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Full text of "Bulletin - United States National Museum"

CHECKLIST OF AMERICAN PHYCITINAE 



Pages 317-329 from 

United States National Museum Bulletin 207 

AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 

By Carl Heinrich 



Checklist of American Phycitinae 



(Synonyms in italics) 



1. Cryptoblabes Zeller 

1. gnidiella (Milliere): Europe, Africa, Asia, 

Bermuda, Venezuela, Brazil 

2. AcROBASis Zeller 

Mineola Hulst 
Seneca Hulst 
Acrocaula Hulst 

2. indigenella (Zeller): Eastern U. S. and Can- 

ada, California 
nebulo (Walsh) 
nebulella (Riley) 
zelatella (Hulst) 

3. grossbecki (Barnes and McDunnough), new 

comb.: Florida 

4. vaccinii Riley: U. S. 

5. amplexella Ragonot: Eastern U. S. 

6. tricolorella Grote: U. S., Canada 

scitvlella Hulst 

7. comptella Ragonot: Western U. S. 

8. minimella Ragonot: Eastern U. S. 

nigrosignella Hulst 

9. feltella Dyar: Eastern U. S., Canada 

10. palliolella Ragonot: Eastern U. S., Canada 

albocapitella Hulst 

11. caryalbella Ely: U. S. (Connecticut) 

12. juglandis (LeBaron): Eastern U. S. 

13. sylviella Ely: Eastern U. S., Canada 

14. kearfottella Dyar: Eastern U. S. 

15. caryae Grote: Eastern U. S., Canada 

16. evanescentella Dyar: U. S. (Florida) 

17. stigmella Dyar: Eastern U. S. 

18. aurorella Ely: Eastern U. S. 

19. peplifera Dyar: Eastern U. S. 

20. exsulella (Zeller), new comb.: Eastern U. S. 

septentrionella Dyar 

21. angusella Grote: Eastern U. S., Canada 

eliella Dyar 

22. demotella Grote: Eastern U. S. 

23. latifasciella Dyar: Eastern U. S. 

24. irrubriella Ely: Eastern U. S. 

25. normella Dyar: Eastern U. S. (Connecticut) 

26. malipennella Dyar: Eastern U. S. (Con- 

necticut) 

27. dyarella Ely: Eastern U. S. (Connecticut) 

28. ostryella Ely: Eastern U. S., Canada 

29. secundella Ely: Eastern U. S., Canada 

30. coryliella Dyar: Eastern U. S. 

31. hebescella Hulst: Eastern U. S. (New Jer- 

sey) 

32. cirroferella Hulst: Eastern U. S. (Texas) 

33. cuniilae Dyar and Hernrich: Eastern U. S. 

34. carjavorella Ragonot: Eastern and South- 

western U. S. 



35. comacornella (Hulst), new comb.: Eastern 

U. S. (Texas) 

36. betulella Hulst: Eastern and Western U. S., 

Canada 

37. rubrifasciella Packard: Eastern U. S., 

Canada 
alnella McDunnough 

38. comptoniella Hulst: Eastern U. S., Canada 

39. myiicella Barnes and McDunnough: U. S. 

(Florida) 

40. tumidulella (Ragonot), new comb.: U. S. 

(Florida) 

3. Rhodophaea Gu^n^e 

41. caliginella (Hulst), new comb.: U. S. (Cali- 

fornia, Arizona) 
caliginoidella (Dyar) 

42. supposita (Heinrich), new comb.: Canada 

(British Columbia) 

4. Trachycera Ragonot 

43. paUicornella (Ragonot): U. S. (Texas) 

5. Anabasis Heinrich, new genus 

44. ochrodesma (Zeller), new comb.: U. S., 

(Florida), Mexico, Guatemala, Panamd 
Colombia, West Indies 
crassisquamella (Hampson) 

6. MiLDRixiA Dyar 

45. constitutionella Dyar: Mexico, Guatemala 

7. Sematoneura Ragonot 

46. atrovenosella Ragonot: Mexico, Costa Rica, 

Colombia, Ecuador, Peru, Argentina 

47. abitus Heinrich, new species: Ecuador 

8. Hypsipyla Ragonot 

48. grandeUa (Zeller): U. S. (Florida), West 

Indies and Tropical America to Argen- 
tina 
cnahella Dyar 

49. ferreahs (Hampson), new comb.: Tropical 

America (Costa Rica to Brazil) 

50. dorsimacula (Schaus), new comb.: Costa 

Rica 

51. fluviatella Schaus: Costa Rica 

9. Hemiptilocera Ragonot 

52. chinographella Ragonot: French Guiana, 

Brazil, Peril 

53. bigrana (Zeller) : Mexico, Colombia 

54. plumigerella (Ragonot), new comb.: "Amer. 

Merid." 

55. letharda (Schaus), new comb.: Panamd, 

Mexico 

56. jocarella (Schaus) : Costa Rica, Panamd, 

Brazil 

57. exoleta (Zeller) : Colombia 

317 



318 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



10. Crocidomera Zeller 

58. turbidella Zeller: Cuba, Jamaica, Mexico, 

U. S. (Texas) 

59. fissuralis (Walker) : Dominican Republic, 

Puerto Rico 
adonea (Felder and Rogenhofer) 

60. stenopteryx (Dyar), new comb.: Mexico 

11. CuNiBERTA Heinrich, new genus 

61. subtinctella (Ragonot), new comb.: West- 

ern U. S. and Canada 

12. Heras Heinrich, new genus 

62. disjunctus Heinrich, new species: Colombia 

13. Adanarsa Heinrich, new genus 

63. intransitella (Dyar), new comb.: U. S. 

(Arizona, New Mexico) 

14. Birinus Heinrich, new genus 

64. russeolus Heinrich, new species: British 

Guiana 

15. Bertelia Barnes and McDunnough 

65. gi-isella Barnes and McDunnough: U. S. 

(Arizona) 

16. Hypargtria Ragonot 

66. definitella (Zeller) : Puerto Rico, Virgin 

Islands, Colombia, Brazil 

67. slossonella (Hulst), new comb.: U. S. 

(Florida), Mexico 
tenuella (Barnes and McDminough) 

17. Chararica Heinrich, new genus 

68. annuliferella (Dyar), new comb.: U. S. 

(New Mexico, Arizona) 

69. hystriculella (Hulst), new comb.: U. S. 

(Texas, Florida) 

70. bicolorella (Barnes and McDunnough), new 

comb.: U. S. (Ai-izona, Nevada, Cali- 
fornia) 

18. Myelopsis Heinrich, new genus 

71. coniella (Ragonot), new comb.: U. S., Can- 

ada, Mexico 
nefas (Dyar) 

72. immundella (Hulst), new comb.: U. S. 

(Texas) 

73. subtetricella (Ragonot), new comb.: U. S., 

Canada 
zonulella (Ragonot) 
obnupsella (Hulst) 

74. minutularia (Hulst), new comb.: U. S. 

(Texas) 

75. alatella (Hulst), new comb.: Western U. S. 

rectistrigella (Ragonot) 
jragilella (Dyar) 
piazzella (Dyar) 

19. Anypsipyla Dyar 

76. univitella Dyar: Cuba, Mexico, Guatemala, 

Panamd, Venezuela, Brazil, Perii, Ecua- 
dor, Jamaica 

20. Apomyelois Heinrich, new genus 

77. bistriatella (Hidst), new comb.: Eastern 

U. S., Canada 
bilineatella (Ragonot) 



21. EcTOMYELOis Heiurich, new genus. 

78. decolor (Zeller), new comb.: Tropical 

America 
ephestiella (Hampson) 

79. ceratoniae (Zeller), new comb.: Europe, 

U. S. (Florida), Puerto Rico, Jamaica, 
Argentina 
oporedesteUa (Dyar) 

80. muriscis (Dyar), new comb.: Tropical 

America 
palpalis (Dyar) 

81. furvidorsella (Ragonot), new comb.: Puerto 

Rico 

82. zcteki Heinrich, new species: Panama 

22. Paramyelois Heimich, new genus. 

83. transitella (Walker), new comb.: U. S., 

tropical America 
notatalis (Walker) 
soliteUa (Zeller) 
duplipunctella (Ragonot) 
venipars (Dyar) 
cassiae (Dyar) 

23. PsEUDODivoNA Dyar 

84. commensella Dyar: Al^xico 

85. cispha Dyar: Guatemala, Costa Rica, Brit- 

ish Honduras 

86. santa-maria Dyar: Guatemala 

87. carabayella Dyar: Peru, Bohvia, Colombia 

24. Protomoerbes Heinrich, new genus 

88. aberrans Heinrich, new species: Colombia 

89. separabilis Heinrich, new species: Colombia 

25. DiATOMOCERA Ragonot 

Cabima Dyar 

90. tenebricosa (Zeller): Colombia, French Gui- 

ana, Costa Rica 

91. dosia (Dyar), new comb.: Panamd 

92. excisalis (Hampson), new comb.: French 

Guiana, Bolivia (?) 

93. decurrens (Dyar), new comb.: PanamS, 

94. majuscula Heinrich, new species: Brazil 

95. albosigno Heinrich, new species: Brazil 

96. hoplidice (Dyar), new comb.: Panama 

97. extracta Heinrich, new species: Costa Rica 

98. mochlophleps (Dyar), new comb.: Mexico 

26. PsEUDOCABiMA Hcinrich, new genus 

99. castronalis Heinrich, new species: Brazil 

100. fearnella (Schaus), new comb.: Costa Rica, 

Guatemala 

101. guianalis Heinrich, new species: French 

Guiana, British Guiana 

102. euzopherella (Dyar), new comb.: PanamS, 

103. pombra (Dyar), new comb.: Panama 

104. nigristrigella (Ragonot), new comb.: Brazil 

105. arizonensis Heinrich, new species: U. S. 

(Arizona) 

106. expunctrix (Dyar and Heinrich), new comb.: 

Brazil 

107. perrensiella (Ragonot, new comb.: Argen- 

tina 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



319 



108. rubrizonalis (Hampson), new comb.: French 

Guiana, Brazil 

27. Hyalospila Ragonot 

109. stictoneurella Ragonot: Mexico, Guatemala, 

Brazil 

110. celiella Schaus: Costa Rica 

111. insequens Heim-ich, new species: Bolivia, 

Colombia 

112. majorina Heim-ich, new species: Mexico 

113. fulgidula Heinrich, new species: Cuba 

114. egeneUa (Ragonot), new comb.: Brazil 

115. xanthoudemia (Dyar), new comb.: Panama, 

Costa Rica 

116. anguIineUa (Schaus), new comb.: Costa 

Rica 

117. clevelandella (Dyar): Panama 

118. semibrunneella Ragonot: Colombia 

28. FuNDELLA Zeller 

119. pellucens Zeller: U. S. (Florida), West In- 

dies, Brazil, Bolivia 
cistipennis (Dyar) 

120. argentina Dyar: U. S. (Florida, Texas), 

West Indies, Venezuela, Brazil, Argen- 
tina 
eucasis Djar 

121. agapella Schaus: Galapagos Islands 

122. ignobilis Heinrich: Mexico, Guatemala, 

Costa Rica, Cuba, Puerto Rico, Haiti 

123. ahemora Dyar: Mexico, Guatemala, Costa 

Rica 

29. DiFUNDELLA Dyar 

124. corynophora Dyar: Guatemala, Panamfi, 

French Guiana 

125. subsutella (Schaus) , new comb. : Costa Rica 

126. dis tractor Heinrich, new species: Puerto 

Rico 

127. tolerata Heinrich, new species: Bolivia 

30. CoPTARTHRiA Ragouot 

128. dasypyga (ZeUer): Colombia, Guatemala 

31. Prom YLEA Ragonot 

129. limigerella Ragonot: Western U. S. and 

Canada. 

130. lunigerella glendella (Dyar) : Colorado 

131. dyari Heinrich, new name: Mexico 

zimmermani (Druce) 
drucei (Dyar) 

132. druceii (Ragonot), new comb.: Guatemala 

133. mindosis Dyar: Mexico 

134. dasystigma Dyar: Mexico 

32. Anadelosemia Dyar 

135. senesciella (Schaus): Costa Rica 

136. tecmessella (Schaus): Costa Rica 

137. fifria Dyar: Guatemala 

138. base Dyar: Guatemala 

139. obstiteUa (Schaus), new comb.: Costa Rica 

140. texanella (Hulst), new comb.: U. S. (Texas, 

Florida), Puerto Rico, Cuba 
dulciella (Hulst) 

141. condigna Heinrich, new species: U. S. (Ari- 

zona) 



33. Dasypyga Ragonot 

142. alternosquameUa Ragonot: Western U. S., 

Canada 
stictophorella Ragonot 

34. Rampylla Dyar 

143. orio Dyar: Mexico 

144. polydectella (Schaus) : Costa Rica 

145. subcaudata (D^^ar), new comb.: Guatemala, 

Costa Rica, Brazil 

146. lophotalis Heimich, new species: Mexico, 

Guatemala 

35. FuLRADA Heinrich, new genus 

147. querna (Dyar), new comb.: Panamd, 

148. carpasella (Schaus), new comb.: Gahlpagos 

Islands 

36. ScoRYLUS Heinrich, new genus 

149. cubensis Heinrich, new species: Cuba 

37. Davara Walker 

Homalopalpia Dyar 
Eucardinia Dyar 

150. caricae (Dyar), new comb.: U. S. (Florida), 

Tropical America 
dalera (Dyar) 

151. columneUa (Zeller), new comb.: Colombia 

152. nerthella (Schaus), new comb.: Mexico, 

Guatemala, Costa Rica 
euthales (Dyar) 

153. paranensis (D^^ar), new comb.: Brazil 

154. azonaxsalis Walker: Brazil 

155. interjecta Heinrich, new species: Puerto 

Rico, Dominican Republic 

156. rufuleUa (Ragonot), new comb.: Puerto Rico 

38. Sarasota Hulst 

Cuba Dyar 

157. plumigerella Hulst: U. S. (Florida) 

158. furculella (Dyar), new comb.: Cuba, Puerto 

Rico, Dominica, Virgin Islands 

159. ptyonopoda (Hampson), new comb.: Wind- 

ward Islands 

39. PiESMOPODA Zeller 

Discopalpia Ragonot 
Amphycltopsis Dyar 

160. rubicundella Zeller: Brazil 

161. xanthomera Dyar: Guatemala, Panamd 

Costa Rica, French Guiana 
xanthozona Dyar 

162. trichoma ta (Zeller): Colombia 

163. flavicans (Zeller): Colombia, French Guiana 

jratella Dyar 

164. ragonoti (Dyar), new comb.: Mexico, Gua- 

temala, Costa Rica 

165. Isabella (Dyar), new comb.: Costa Rica 

166. xanthopolys Dyar: Panamd 

167. parva Heinrich, new species: Panami 

168. semirufella (Zeller): Colombia 

169. apocerastes Dyar: Mexico, Costa Rica, 

French Guiana, Brazil, Dominica 

170. montella Schaus: Costa Rica 



320 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



40. Atheloca Heinrich, new genus 

171. subrufella (Hulst), new comb.: U. S. (Flor- 

ida), Cuba, Virgin Islands 
filiolella (Hulst). Virgin Islands 
piychis (Dyar) 

172. bondari Heinrich, new species: BrazU 

41. Praedonula Heinrich, new genus 

173. almonella (Dyar), new comb.: Panama 

42. Peadus Heinrich, new genus 

174. burdettellus (Schaus), new comb.: Costa 

Rica, Guatemala 
semproniella (Schaus) 

175. dissitus Heinrich, new species: Brazil 

176. subaquilellus (Ragonot), new comb.: Guate- 

mala 

43. Gabinius Heinrich, new genus 

177. paulsoni (Ragonot), new comb.: Chile 

44. Ceracanthia Ragonot 

Procandiopa Dyar 

178. mamella (Dyar), new comb.: PanamS,, 

Guatemala 

179. vepreculella Ragonot: Ecuador 

45. Megarthria Ragonot 

180. peterseni (Zeller): Guatemala, Colombia, 

Brazil, Peru 

181. squamifera Heinrich, new species: Costa 

Rica 

1 82. frustrator Heinrich, new species: Costa Rica 

183. schausi Heinrich, new species: Costa Rica 

184. cervicalis Dyar: Cuba 

185. alpha Heinrich, new species: Guatemala, 

Mexico, Costa Rica, Panama, Bolivia, 
Brazil 

186. beta Heinrich, new species: Mexico, Guate- 

mala, Costa Rica, Trinidad 

46. Drescoma Dyar 

187. cyidipsa Dyar: Mexico, Guatemala, Pan- 

ama, French Guiana 

188. cinilixa Dyar: Guatemala, PanamS, 

47. MoNOPTiLOTA Hulst 

189. pergratialis (Hulst): U. S. 

grotella (Ragonot) 
nubilella Hulst 

48. Zamagiria Dyar 

190. dixolophella Dyar: Panama 

191. pogerythrus Dyar: Mexico, Guatemala 

192. hospitabihs Dyar: Cuba 

193. masculinus Dyar: Guatemala 

194. australella (Hulst), new comb.: U. S. 

(Texas, Florida) 
humeliella (Barnes and McDunnough): 
U. S. (Texas, Florida) 

195. fraterna Heinrich, new species: Cuba 

196. laidion (Zeller): U. S. (Florida), Tropical 

America 
deia Dyar 
striella Dyar 

197. ipsetona Dyar: Costa Rica 



49. Anegcephalesis Dyar 

198. arcteUa (Ragonot), new comb.: U. S. 

(Florida), Bahamas, Cuba 
cathaeretes Dyar 

50. Magiriopsis Heinrich, new genus 

199. denticosella (Dyar), new comb.: Tropical 

America 
crisfalis (Hampson) 

51. Ancylostomia Ragonot 

200. stercorea (Zeller): U. S. (Florida), Tropical 

America 
ignobilis (Butler) 
diffissella CZeller) 

201. sauciella (Zeller): Colombia 

202. argyrophleps Dyar: Mexico, Guatemala 

203. euchi'oma Dyar: Brazil 

52. Caristanius Heimich, new genus 

204. pellucidellus (Ragonot), new comb.: Puerto 

Rico, St. Vincent, Jamaica, Surinam, 
Brazil 
melanoplaga (Hampson) 

205. decoloralis (Walker), new comb.: Southern 

U.S. 
metagrammalis (Walker) 
Jurjurellus (Hulst) 
floridellus (Hulst) 

206. guatemalellus (Ragonot), new comb.: 

Guatemala 

53. Etiella Zeller 

207. zinckenella (Treitschke): Europe. Asia, 

U. S., Tropical America 
etiella (Treitschke) 
schisticolor Zeller 
mllosella Hulst 
rubribasella Hulst 

54. Glyptocera Ragonot 

208. consobrinella (Zeller): Eastern U. S., Can- 

ada 
busckella (Dyar) 

55. Pima Hulst 

209. boisduvaliella (Gu^n^e), new comb.: Eu- 

rope, Canada 

210. albiplagiatella (Packard), new comb.: East- 

ern U. S., Canada 

211. albiplagiatella occidentalis Heinrich, new 

race: Western U. S. 

212. fostereUa Hulst: Western U. S., Canada 

213. vivideUa (McDunnough), new comb.: Can- 

ada 

214. albocostalialis (Hulst), new comb.: Western 

U. S., Canada 

215. albocostaliaUs subcosteUa (Ragonot), new 

comb.: Southwestern U. S. 

216. fulvirugella (Ragonot,) New comb.: Western 

U. S. (California) 

217. gi-aniteUa (Ragonot), new comb.: Western 

U.S. 
piperella (Dyar) 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITESTAE 



321 



218. parkerella (Schaus), new comb.: Western 

U. S. (Montana) 

56. Interjectio Heinrich, new genus 

219. denticulella (Ragonot), new comb.: North- 

western U. S., Canada 

220. columbiella (McDunnough), new comb.: 

Northwestern U. S., Canada 

221. ruderella (Ragnot), new comb.: "N. 

Amer." (Cahfornia?) 

222. niviella (Hulst) new comb.: U. S., Canada 

57. Ambesa Grote 

223. laetella Grote: Westerxi U. S., Canada 

224. walsinghami (Ragonot): Western U. S. 

monodon Dyar 

225. walsinghami mirabella Dyar, new status: 

U. S. (Southern Cahfornia) 

226. lallatahs (Hulst): Western U. S. (Nevada, 

Utah) 

58. Catastia Hiibner 

227. bistriateUa (Hulst), new comb.: Western 

U. S. (California) 

228. incorrusceUa (Hulst), new comb.: Western 

U. S. (California) 

229. actualis (Hulst), new comb.: Western U. S., 

Canada 

59. Immyrla Dyar 

230. nigrovittella Dyar: Eastern U. S. 

60. Oreana Hulst 

231. unicolorella (Hulst): Eastern U. S., Canada 

leucophaeella (Hulst) 

61. Olybria Heinrich 

232. ahculella (Hulst), new comb.: Southwestern 

U.S. 
oherthuriella (Ragonot) 

233. furciferella (Dyar) new comb.: Southwest- 

ern U. S. (Arizona) 

62. Salebriacus Heinrich, new genus 

234. odiosellus (Hulst), new comb.: Western 

U. S. 
hakerella (Dyar) 
yumaella (Dyar) 

63. Salebriaria Heinrich, new genus 

235. turpidella (Ragonot), new comb.: Southern 

U.S. 
ademptandella (Dyar) 

236. nubiferella (Ragonot), new comb.: U. S. 

237. engeh (Dyar) U. S. 

238. annulosella (Ragonot), new comb.: U. S. 

(Texas, North Carolina) 
rohustella (Dyar) 

239. tenebrosella (Hulst), new comb.: U. S. 

quercicolella (Ragonot) 
heinrichalis (Dyar) 

240. pumilella (Ragonot) new comb.: Southeast- 

ern U. S. 
georgiella (Hulst) 

241. fructetella (Hulst) new comb.: U. S. 

rectistrigella (Dyar) 



64. Quasisalebria Henrich, new genus 

242. admixta Heinrich, new species: Western 

U. S. 

65. Ortholepis Ragonot 

243. jugosella Ragonot: Eastern U. S., Canada 

244. pasadamia (Dyar), new comb.: U. S., Can- 

ada 

66. Polopeustis Ragonot 

245. arctiella (Gibson): Alaska, Canada 

67. Meroptera Grote 

Emmerita Hampson 

246. mirandella Ragonot: Western U. S. 

247. cviatella Dyar: U. S. (Illinois, Mississippi) 

248. pravella (Grote): U. S., Canada 

249. abditiva Heinrich, new species: Canada 

68. Nephopteryx Hiibner 

250. subfuscella (Ragonot), new comb.: Eastern 

U. S., Canada 
semiobscurella (Hulst) 

251. delassahs Hulst: Western U. S. 

purpurella (Hulst) 
pudibundella (Ragonot) 

252. delassalis fraudifera Heinrich, new race: 

Canada (British Columbia), U. S. (Wash- 
ington) 

253. rubescentella (Hulst): U. S. 

254. fernaldi (Ragonot), new comb.: U. S., 

Canada 

255. dammersi Heinrich, new species: Western 

U. S. (California, Arizona) 

256. dammersi floridensis Heinrich, new race: 

U. S. (Florida) 

257. vetustella (Dyar), new comb.: U. S., Can- 

ada 

258. inconditella (Ragonot), new comb.: West- 

ern U. S. (Arizona, Colorado) 

259. subcaesiella (Clemens), new comb.: U. S., 

Canada 
contatella (Grote) 

260. virgatella (Clemens), new comb.: U. S., 

Canada 
quinquepundella (Grote) 

261. carneella Hulst: U. S., Canada 

inquilinella (Ragonot) 

262. basilaris Zeller: U. S., Canada 

263. termitalis (Hulst), new comb.: Western 

U. S., Canada 
letngatella (Hulst) 

264. termitahs yuconella Dyar, new status: 

Alaska 

265. bifasciella Hulst: U. S. (Arizona) 

nogalesella (Dyar) 

266. uvinella (Ragonot), new comb.: Eastern 

U.S. 
afflidella (Hidst) 
liguidambarella (Dyar) 

267. celtidella (Hulst), new comb.: U. S. 



322 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



268. rubrisparsella (Ragonot): U. S. 

rufihasella (Ragonot) 
croceella (Hulst) 
texanella (Hulst) 

269. gilvibasella Hulst: U. S. (Texas) 

lacteella (Hulst) 

270. crassifasciella Ragonot: Eastern U. S. 

decipientella Dyar 
crataegella B. and McD. 

271. bisra Dyar: Mexico 

69. Tlascala Hulst 

272. reductella (Walker): Eastern U. S. 

gleditschiella (Fernald) 

70. Tulsa Heinrich, new genus 

273. finitella (Walker), new comb.: Eastern U.S., 

Canada 
melanellus (Hulst) 

274. umbripennis (Hulst), new comb.: U. S. 

(Colorado) 
gillettella (Dyar) 

275. oregonella. (Barnes and McDunnough), new 

comb.: U. S. (Oregon) 

276. infinitella (Dyar), new comb.: Mexico 

71. Homoeographa Ragonot 

277. lanceolella Ragonot: Peru 

72. Telethusia Heinrich, new genus 

278. ovalis (Packard), new comb.: U. S., Canada 

latifasciatella (Packard) 
geminipunctella (Ragonot) 
modestella (Hulst) 

279. rhypodella (Hulst), new comb.: U. S. ("Ore- 

gon") 

73. Phobus Heinrich, new genus 

280. brucei (Hulst), new comb.: Western U. S. 

281. funerellus (Dyar), new comb.: WesternU.S., 

Canada 

282. cm-vateUus (Ragonot), new comb.: Western 

U.S. 

283. incertus Heinrich, new species: Western 

U. S. (CaHfornia) 

74. AcTRix Heinrich, new genus 

284. nyssaecolella (Dyar), new comb.: Eastern 

U.S. 

285. dissimulatrix Heinrich, new species: Eastern 

U. S. (Virginia) 

75. Stylopalpia Hampson 

286. lunigerella Hampson: West Indies, Mexico 

287. scobiella (Grote), new comb.: U. S. (Texas, 

Colorado) 
decimerella (Hulst) 

288. argentinensis Heinrich, new species: Ar- 

gentina 

76. Pyla Grote 

289. fasciolalis (Hulst), new comb.: Canada 

(British Columbia) 

290. impostor Heinrich, new species: Western 

U. S., Canada 

291. aequivoca Heinrich, new species: Western 

Canada 



292. insinuatrix Heinrich, new species: Canada 

(Manitoba) 

293. aenigmatica Heinrich, new species: U. S., 

Canada 

294. criddlella Dyar: Canada (Manitoba) 

295. fusca (Haworth), new comb.: Holarctic 

moesteUa (Walker) 
frigidella (Packard) 
cacabella (Hulst) 
triplagiatella (Dyar) 

296. hypochalciella (Ragonot), new comb.: North- 

western U. S., Canada. 
blackmorella (Dyar) 

297. hanhamella Dyar: Canada (Manitoba) 

298. scintillans (Grote): Western U. S. (Califor- 

nia) 
Jeella Dyar 

299. sylphiella Dyar: Northwestern U. S., Canada 

300. rainierella Dyar: Northwestern U. S. (Wash- 

ington) 

301. aeneella Hulst: Western U. S. (Colorado, 

Utah) 

302. aeneoviridella Ragonot: Western U. S., 

Canada 

303. metahcella Hulst: Western U. S. (Colorado, 

Utah) 

304. fasciella Barnes and McDunnough: North- 

western U. S. (California) 

305. nigricula Heinrich, new species: Western 

U. S. (Nevada) 

306. viridisufifusella Barnes and McDunnough: 

Western U. S. (California) 
77. DioRYCTRiA Zeller 

Pinipestis Grote 

307. abietella (Denis and Schiffermiiller): North- 

ern Hemisphere 
decuriella (Htibner) 
abietivorella (Grote) 
elegantella (Hulst) 

308. sysstratiotes Dyar: Guatemala 

309. reniculella (Grote): Northern U. S., Canada 

310. ponderosae Dyar: Western U. S. (Montana, 

California) 

311. majorella Dyar: Mexico 

muellerana Dyar 

312. disclusa Heinrich: Eastern U. S. 

313. auranticella (Grote): Western U. S., Canada 

miniatella Ragonot 
xanthaenobares Dyar 
314. erythropasa (Dyar): Southwestern U. S. 
(Arizona) 

315. horneana (Dyar): Cuba 

316. pygmaeeUa Ragonot: Eastern U. S. 

317. zimmermani (Grote): U. S., Canada 

delectella (Hulst) 
austriana (Cosens) 

318. cambiicola (Dyar): Western U. S. 

319. amatella (Hulst): Eastern U. S. 

320. albovittella (Hulst): Western U. S. 



399671—56 3 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



323 



321. gulosella (Hulst), new comb.: Western U. S. 

(Colorado, New Mexico) 

322. baumhoferi Heinrich, new species: South- 

western U. S. (Arizona) 

323. subtracta Heinrich, new species: South- 

western U. S. (New Mexico) 

324. clarioraUs (Walker): Eastern U. S. 

brunneella (Dyar) 

78. Oeyctometopia Ragonot 

325. fossulatella Ragonot: U. S. (Texas), Trop- 

ical America 
moeschleri (Ragonot) 

79. Sarata Ragonot 

326. edwardsialis (Hulst), new comb.: Western 

U.S. 
polyphemella (Ragonot) 

327. pullatella (Ragonot), new comb.: Western 

U.S. 

328. punctella (Dyar), new comb.: Mexico 

329. punctella septentrionaria Heinrich, new race: 

Western U. S. 

330. incanella (Hulst), new comb.: Western U.S. 

aridella (Dyar) 

331. atrella (Hulst), new comb.: Western U. S. 

(Colorado) 

332. caudellella (Dyar), new comb.: Western 

U. S., Canada 

333. dnopherella Ragonot: Western U. S. (Cah- 

fornia) 

334. nigi'ifasciella Ragonot: Western U. S., Can- 

ada 

335. cinereella Hulst: Western U. S. (Colorado) 

336. rubrithoracella (Barnes and McDunnough), 

new comb.: Western U. S. 

337. tephrella Ragonot: Western U. S. (Wash- 

ington) 

338. alpha Heinrich, new species: Canada (Sas- 

katchewan) 

339. beta Heinrich, new species: Western U. S., 

Canada 

340. gamma Heinrich, new species: Western 

U. S. (California) 

341. iota Heinrich, new species: Western U. S. 

(California) 

342. perfuscalis (Hulst): Western U. S. 

ezcantalis (Hulst) 

343. epsilon Heinrich, new species: Western U.S. 

344. phi Heim-ich, new species: Western U. S. 

345. kappa Heinrich, new species: Western U. S. 

(Arizona) 

346. delta Heinrich, new species: Western U. S. 

80. Philodema Heinrich, new genus 

347. rhoiella (Dyar), new comb.: Western U. S. 

81. Lipographis Ragonot 

348. fenestrella (Packard): Western U. S. (Cali- 

fornia) 
humilis Ragonot 

349. leoninella (Packard): Western U. S., Can- 

ada 
pallidella (Dyar) 



350. truncatella (Wright), new comb.: South- 

western U. S. (California) 

351. umbrella (Dyar), new comb.: Southwestern 

U. S. (California) 

352. subosseella Hulst: Bahamas 

82. Adelphia Heinrich, new genus 

353. petrella (Zeller), new comb.: U. S. 

rubiginella (Walker) 
rvfinalis (Walker) 
hapsella (Hulst) 

354. ochripunctella (Dyar), new comb.: Western 

U. S. (California) 

83. Tota Heinrich, new genus 

355. galdinella (Schaus), new comb.: Galdpagos 

Islands 

84. Ufa Walker 

356. lithosella (Ragonot), new comb.: South- 

western U. S., Mexico 
luteella Hulst) 

357. roseitinctella (Dyar), new comb.: South- 

western U. S., Mexico 

358. senta Heinrich, new species: Southwestern 

U. S. (Texas, Arizona) 

359. rubedinella (Zeller), new comb.: U. S. (Flor- 

ida), Tropical America 
translucida (Walker) 
rufescentalis (Walker) 
minualis (Walker) 
deprivalis (Walker) 
Venezuelans Walker 
pyrrhochrellus (Ragonot) 

85. Elasmopalpus Blanchard 

360. lignosellus (Zeller) : U. S., Tropical America 

angustellus Blanchard 
tartarella (Zeller) 
incautella (Zeller) 
major (Zeller) 
anthracellus Ragonot 
carbonella (Hulst) 
puer Dyar 

86. AcRONCosA Barnes and McDunnough 

361. albiflavella Barnes and McDunnough: West- 

ern U. S. (California) 

362. albiflavella castrella Barnes and McDun- 

nough: Western U. S. (New Mexico) 

363. similella Barnes and McDunnough: West- 

ern U. S. (Nevada, Utah) 

87. Passadena Hulst 

364. flavidorsella (Ragonot): Western U. S., 

Mexico 
canescentella (Hulst) 
constantella Hulst 
cinctella (Hulst) 

88. Ulophoea Ragonot 

Acromeseres Dyar 

365. groteii Ragonot: Eastern U. S. 

tephrosiella Dyar 

366. guarinella (Zeller): Cuba, Colombia 

dialithus (Dyar) 



324 



XINITED STATES NATIONAL MUSEUM BULLETUSf 20 7 



89. Chorrera Dyar 

367. idiotes Dyar: Panamd 

368. extrincica (Dyar), new comb.: Cuba 

369. postica (Zeller), new comb.: Colombia 

90. Tacoma Hulst 

370. feriella Hulst: Southwestern U. S. 

submedlanella Dyar 

91. Adelperga Heinrich, new genus 

371. cordubensiella (Ragonot), new comb.: Ar- 

gentina 

92. EuMYSiA Dyar 

372. mysiella (Dyar) : Western U. S. 

373. maidella (Dyar): Western U. S., Canada 

374. pallidipennella (Hulst), new comb.: West- 

ern U. S. 

375. fuscatella (Hulst): Western U. S. (Cali- 

fornia) 

376. semicana Heinrich, new species: Western 

U. S. (Washington) 

93. DiviTiACA Barnes and McDunnough 

377. ochreUa Barnes and McDunnough: South- 

ern U. S. (Florida) 

378. simulella Barnes and McDunnough: South- 

ern U. S. (Florida) 

379. parvulella Barnes and McDunnough: South- 

ern U. S. (Florida) 

380. parvulella consociata Heinrich, new race: 

Colombia 

94. Macrorrhinia Ragonot 

Dolichorrhmia Ragonot 

381. aureofasciella Ragonot: Southwestern U. S., 

Mexico 

382. placideUa (Zeller): Brazil 

95. OcALA Hulst 

383. dryadella Hulst: Southern U.S. (Florida) 

platanella (Grossbeck) 

96. Valdivia Ragonot 

Maricopa Hulst 

384. coquimbella Ragonot: Chile 

385. lativittella (Ragonot): Southwestern U. S., 

Mexico 
aureomaculella (Dyar) 

386. walkerella (Ragonot), new comb.: Chile 

97. Protasia Heinrich, new genus 

387. mirabilicornella (Dyar), new comb.: West- 

ern U. S. (California) 

98. Heterographis Ragonot 

Mona Hulst 

388. morrisonella Ragonot: U. S., Mexico 

coloradensis Ragonot 
olbiella (Hulst) 
ignistrigella Ragonot 
palloricostella (Walter) 

99. Staudingeria Ragonot 

389. albipenella (Hulst): Western U. S. 

olivacella Dyar 
perluteella Dyar 



100. HuLSTiA Ragonot 

390. undulatella (Clemens): U. S., Canada 

rubiginalis (Walker) 
obsipella (Hulst) 
fumosella (Hulst) 

101. Honora Grote 

391. mellinella Grote: U. S. 

ochrimaculella Ragonot 

392. subsciurella Ragonot: Western U. 8. 

393. sciurella Ragonot: Western U. S. (Cali- 

fornia) 

394. dotella Dyar: Western U. S. (California) 

395. montinatatolla (Hulst): Western U. S. 

canicostella Ragonot 

396. perdubiolla (Dyar), new comb.: Western 

U. S. 

102. HoNORiNus Heinrich, new genus 

397. fuliginosus Heinrich, new species: Perii 

103. Omcolabis Zeller 

Endommasis Hampson 

398. anticella Zeller: Tropical America 

nipritella (Hampson) 

104. Cabotia Ragonot 

Encystia Hampson 

399. semidiscella Ragonot: Argentina 

400. schini (Berg): Argentina 

401. rh^'thmatica Dyar: PanamA 

402. cundajensis '(Zeller): Colombia 

impeditella (Zeller) 

403. bonhoti (Hampson), new comb.: Bahamas, 

Jamaica 

105. Canarsia Hidst 

404. ulmiarrosorella (Clemens): U. S., Canada 

pneumatella (Hulst) 
ulmella (Ragonot) 
fuscatella (Hulst) 
gracilella Hulst 
fellcalella Dyar 

106. Harnocha Dyar 

405. velessa Dyar: Panamd 

107. EuRYTHMASis Dyar 

406. ignifatua Dyar: Panamd, Puerto Rico, 

Cuba 

108. EuRYTHMiDiA Ragonot 

407. ignidorsella (Ragonot): U. S. (Arizona), 

Mexico, Panamd 

109. WuNDERiA Grossbeck 

408. neaeriatella Grossbeck: U. S. (Florida) 

110. Oedothmia Hampson 

Synotlimia Hampson 

409. endopyrella Hampson: Mexico, Bahamas 

bahamasella (Hampson) 

111. Stylobasis Hampson 

410. rubripurpurea Hampson: Mexico, Brazil 

112. DiviANA Ragonot 

Dannemora Hulst 

411. eudoreella Ragonot: Eastern U. S. 

edentella (Hulst) 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



325 



113. Palatka Hulst 

412. nymphaeella (Hulst): Eastern U. S. 

verecuntella (Grossbeck) 

114. Cacozophera Dyar 

413. venosa Dyar: Guatemala 

115. PsoROSiNA Dyar 

414. hammondi (Riley): Eastern and Central 

U. S., Canada 
angulella Dyar 

116. Patriciola Heiru'ich, new genus 

415. semicana Heinrich, new species: Utah 

117. Paconius Heinrich, new genus 

416. corniculatus Heinrich, new species: Puerto 

Rico 

118. Aptunga Heinrich, new genus 

417. macropasa (Dyar), new comb.: Guatemala, 

Mexico 

418. imperfecta (Dyar), new comb.: Guatemala 

119. Anderida Heinrich, new genus 

419. sonorella (Ragonot), new comb.: Mexico, 

U. S. (Arizona) 
placidella (Dyar) 

120. Cassiana Heinrich, new genus 

420. malacella (Dyar), new comb.: Mexico, 

Puerto Rico, Virgin Islands 

121. Mescinia Ragonot 

421. triloses Dyar: Panamd 

mosces Dyar 

422. pandessa Dyar: Guatemala 

423. bacerella Dyar: Cuba 

424. estrella Barnes and McDunnough: U. S. 

(Florida) 

425. mooreiHeiru-ich, new species: British Guiana 

426. parvula (Zeller): Colombia 

427. commatella (Zeller): Colombia 

428. berosa Dyar: Panamd, Puerto Rico 

429. peruella Schaus: Perd 

430. discella Hampson: Mexico, Guatemala 

431. indecora Dyar: Mexico 

122. NoNiA Ragonot 

Hypermescinia Dyar 

432. exiguella (Ragonot): Tropical America 

lambella (Dyar) 

123. Phestinia Hampson 

433. costella Hampson: Jamaica, Puerto Rico 

124. CoMOTiA Dyar 

434. torsicornis Dyar: Panamd 

435. convergens (Dyar), new comb.: Guatemala 

125. Bema Dyar 

Relmis Dyar 

436. neuriceUa (Zeller), new comb.: Tropical 

America 
myja Dyar 

437. fritilla Dyar: Guatemala 

438. ydda (Dyar), new comb.: Panamd, French 

Guiana 

439. yddiopsis (Dyar), new comb.: Cuba 

440. fifaca (Dyar), new comb.: Panama 



126. HoMOEOsoMA Curtis 

Phycidea Zeller 

441. electellum (Hulst): U. S., Mexico, Guate- 

mala, Cuba, British West Indies 
opalescellum (Hulst) 
texanellum Ragonot 
tenuipunctella Ragonot 
dvffertella Barnes and McDunnough 

442. stypticellum Grote: U. S., Canada 

uncanale Hulst 

443. striatellum Dyar: Southwestern U. S. 

444. oslarellum Dyar: Western U. S. 

445. oslarellum breviplicitum Heinrich, new race: 

Southwestern U. S. (California) 

446. illuviellum Ragonot: U. S. (Arizona, Colo- 

rado), Mexico 
candidella Hulst 

447. illuviellum emendator Heinrich, new race: 

Western U. S. 

448. imitator Heim-ich, new species: Southwestern 

U. S. (California) 

449. longiventrellum Ragonot: ChUe 

noctivideUa Ragonot 

450. albescentellum Ragonot: Western U. S. 

elongellum Dyar 

451. impressale Hulst: Western U. S., Canada 

452. inornatellum (Hulst): Eastern U. S. 

453. deceptorium Heinrich, new species: U. S. 

(Pennsylvania), Canada 

454. discrebile Heinrich, new species, Brazil 

455. peregrinum Heinrich, new species: U. S. 

(California), Costa Rica 

456. vepaliidum Heim-ich, new species: Argentina 

457. ditaeniateUum Ragonot: Chile 

458. oconequensis (Dyar), new comb.: Perd 

459. assitum Heim-ich, new species: Peru 

460. acmaeopterum Ragonot: Chile 

461. nimbosellum Ragonot: Chile 

462. unionellum Ragonot: Mdxico 

127. Patagonia Ragonot 

463. magellanella (Ragonot): Chile 

128. RoTRUDA Heiru-ich, new genus 

464. mucidella mucidella (Ragonot), new comb.: 

Western U. S., Canada 

465. mucidella reliquella (Dyar), new comb.: 

Eastern U. S., Canada 

466. mucidella ohvaceela (Ragonot), new comb.: 

Tropical America 
musiosum (Dyar) 
cubella (Dyar) 

467. mucidella affusella (Ragonot), new comb.: 

Argentina 

129. Strephomescinia Dyar 

468. schausella Dyar: Cuba 

130. Unadilla Hulst 

Strymax Dyar 



326 



tnsriTED STATES NATIONAL MUSEUM BULLETIN 207 



469. erronella (Zeller): Tropical America. 

ubacensis (Zeller) 
bipundella (Hampson) 
dorae (Dyar) 
pyllis (Dyar) 

470. maturella (Zeller): Colombia, Guatemala, 

Cuba 

471. albidiorella (Richards and Thomason); new 

comb.: Peril 

472. floridensis Heinrich, new species: U. S. (Flor- 

ida) 

473. nasutella Hulst: U. S. (New Mexico) 

131. Laetilia Ragonot 

Laosticha Hulst 

474. coccidivora (Comstock): U. S. 

pallida (Comstock) 
dilatifasciella (Ragonot) 
hulstii Cockerell 

475. coccidivora quadricolorella (Dyar), new 

comb.: Southwestern U. S. 

476. coccidivora cardini Dyar: Cuba, U. S. 

(Florida) 

477. obscura Dyar: Cuba 

478. portoricensis Dyar: Puerto Rico 

479. melanostathma (Meyrick), new comb.: Ar- 

gentina 

480. amphimetra (Meyrick), new comb.: Argen- 

tina 

481. zamacrella Dyar: Western U. S. (California) 

482. myersella Dyar: Eastern U. S. 

483. ephestiella (Ragonot): Southwestern U. S. 

(Arizona) 
lustrella (Dyar) 

484. fiskella Dyar: Eastern U. S. (North Caro- 

lina) 
485. glomis (Dyar), new comb.: Panamd 

132. Baphala Heinrich, new genus 

486. basimaculatella (Ragonot), new comb.: 

Western U. S. 
eremiella (Dyar) 

487. goyensis (Ragonot), new comb.: Brazil, 

Uruguay, Argentina 

488. goyensis oUvacea Heinrich, new race: Argen- 

tina 

489. homoeosomella (Zeller), new comb.: Trop- 

ical America. 
bodkini (Dyar) 
riisto (Dyar) 
taboga (Dyar) 
saissetiae (Dyar) 

490. haywardi Heinrich, new species: Argentina 

491. glabrella (Dyar), new comb.: Guatemala 

492. squalida (Walker), new comb.: Brazil 

133. Rhagea Heinrich, new genus 

493. packardella (Ragonot), new comb.: West- 

ern U. S. 
orobanchella (Dyar) 

494. stigmella (Dyar), new comb.: Southwestern 

U. S. (California), Mexico 
maculicula (Dyar) 



134. ZopHODiA Hiibner 

Dakruma Grote 

495. convolutella (Hiibner): Europe, U. S., Can- 

ada 
grossulariella (Hiibner) 
turbatella (Grote) 
grossulariae (Riley) 
Jranconiella (Hulst) 
bella Hulst 
ihouna Dyar 
dilativitta Dyar 
magnificans Dyar 

135. Melitara Walker 

496. prodenialis Walker: U. S. 

bollii (Zeller) 

497. dentata (Grote): U. S. 

doddalis Dyar 

136. Olycella Dyar 

498. junctolineella (Hulst): Southern U.S. (Texas) 

499. junctolineella pectinatella (Hampson): Mex- 

ico 

500. nephelepasa (Dyar:) Mexico 

501. subumbrella Dyar: Western U.S. 

137. Olyca Walker 

502. phryganoides Walker: Dominican Repub- 

lic, Haiti 

138. Alberada Heinrich 

503. parabates (Dyar): U. S., Mexico 

504. bidentella (Dyar): Southwestern U. S. 

(Texas, Arizona) 

505. holochlora (Dyar): Southwestern U. S. 

(Texas) 

139. Nanaia Heinrich 

506. substituta Heinrich: Peru 

140. Cactoblastis Ragonot 

Neopyralis Brethes 

507. cactorum (Berg): Argentina, Uruguay, Aus- 

tralia 

508. ronnai (Brethes): BrazU 

509. doddi Heim-ich: Argentina 

510. mundelli Heinrich: Peni 

511. bucyrus Dyar: Ai-gentina 

141. Cahela Heinrich 

512. ponderosella (Barnes and McDunnough): 

Western U. S., Mexico 
purgatoria (Dyar) 
interstitialis (Dyar) 
phoenicis (Dyar) 

142. RuMATHA Heinrich 

513. glaucatella (Hulst): Southern U. S. 

514. bihinda (Dyar): Western U. S. 

515. polingeUa (Dyar): Southwestern U. S. (Ari- 

zona, Texas) 

143. YosEMiTiA Ragonot 

516. graciella (Hulst): Western U. S. 

517. longipennella (HuJst): Southwestern U. S. 

(Texas) 

518. fieldiella (Dyar): Western U. S. (California, 

Arizona) 

519. didactica Dyar: Mexico 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



327 



144. TucuMANiA Dyar 

520. tapiacola Dyar: Argentina 

521. porrecta Dyar: Uruguay 

145. Eremberga Heinrich 

522. leuconips (Dyar): Western U. S. (Arizona) 

523. creabates (Dyar): Western U. S. (Cali- 

fornia) 

524. insignis Heinrich: Mexico 

146. Salambona Heiru-ich 

525. analamprella (Dyar): Argentina 

147. Parolyca Dyar 

526. asthenosoma (Dyar): French Guiana 

148. SiGELGAiTA Heinrich 

527. chilensis Heinrich: Chile 

528. huanucensis Heini'ich: Per6 

529. transilis Heim-ich: Peril 

149. Amalafrida Heinrich 

530. leithella (Dyar): West Indies, Venezuela, 

Colombia 

150. OzAMiA Kagonot 

531. lucidalis (Walker): West Indies 

532. fuscomaculella (Wright): Southwestern 

U. S. (California) 
heliophila Dyar 

533. fuscomaculella clarefacta Dyar: U. S. 

(Texas), Mexico 

534. thalassophila Dyar: U. S. (California) 

535. immorella (Dyar), new comb.: Mexico 

536. stigmaferella Dyar: Argentina 

537. hemilutella Dyar: Argentina 

538. punicans Heinrich: Argentina 

151. Cactobrosis Dyar 

539. fernaldialis (Hulst): Southwestern U. S. 

gigantella (Ragonot) 
cinerella (Hulst) 

540. longipennella (Hampson): Mexico 

elongatella (Hampson): 

541. macuhfera Dyar: Mexico 

542. insignatella Dyar: Mexico 

543. strigalis (Barnes and McDunnough): West- 

ern U. S., Mexico 

152. Drescomopsis Dyar 

544. soraella (Druce): Tropical America 

drucella (Dyar) 
subelisa Dyar 

153. Illatila Dyar 

545. gurbyris Dyar: Panamd 

154. Lascelina Heinrich, new genus 

546. canens Heim-ich, new species: Southern 

U. S. (Texas), Mexico 

155. Metephestia Ragonot 

547. simphcula (Zeller): U. S. (Florida), Puerto 

Rico, Colombia, British West Indies 

156. Selga Heinrich, new genus 

548. arizonella (Hulst), new comb.: Southwest- 

ern U. S. (Arizona) 

157. Entmemacornis Dyar 

549. proselytes Dyar: Guatemala 

550. pulla Heinrich, new species: Brazil 



158. Catennia Hampson 

551. rufitinctalis Hampson: French Guiana 

159. RiojA, Heinrich, new genus 

552. nexa Heimich, new species: Argentina 

160. Moerbes Dyar 

553. dryopella (Schaus): Costa Rica 

554. alveolella (Ragonot), new comb.: Brazil 

555. emendata Heinrich, new species: Panamfi, 

French Guiana 

161. MooDNOPSis Dyar 

Campyloplesis Dyar 

556. decipiens Dyar: Mexico 

557. perangusta (Dyar), new comb.: Trinidad 

558. inornatella (Ragonot), new comb.: Costa 

Rica, Brazil 

559. parallela Heinrich, new species: Brazil, Peril 

560. inveterella (Dyar), new comb.: Guatemala 

561. portoricensis Heinrich, new species: Puerto 

Rico 

162. Edulica Ragonot 

562. compedella (Zeller): Tropical America 

163. EuzopHERA Zeller 

563. semifuneralis (Walker): U. S., Canada, 

Mexico 
aglaeella Ragonot 
pallulella (Hulst) 

564. ostricolorella Hulst: Eastern U. S. 

565. nigi'icantella Ragonot: Southwestern U. S., 

M6xico 
griselda Dyar 

164. ExuPERius Heim-ich, new genus 

566. negator Heinrich, new species: Peril 

165. EuLOGiA Heinrich, new genus 

567. ochi-ifrontella (Zeller), new comb.: U. S., 

Canada 
ferruginella (Ragonot) 

166. Prosoeuzophera Heinrich, new genus 

568. impletella (Zeller), new comb.: Colombia, 

Jamaica, Puerto Rico 

167. Farnobia Heinrich, new genus 

569. quadripuncta (Zeller), new comb.: Costa 

Rica, Panama, French Guiana, Colombia 

168. Gennadius Heinrich, new genus 

570. jimctor Heinrich, new species: French Gui- 

ana 

169. MiCROMESciNiA Dyar 

571. pygmaea Dyar: Panama 

170. Ephestiodes Ragonot 

572. gilvescentella Ragonot: Western U. S., Can- 

ada, Mexico 
nigrella Hulst 

573. infimella Ragonot: Eastern U. S. 

574. erythrella Ragonot: Western U. S., Canada 

coloradella (Hulst) 
benjaminella Dyar 

575. mignonella Dyar: U. S. (Texas) 

576. erasa Heim-ich, new species: U. S. (Florida) 

577. lucidibasella Ragonot: Chile 

578. productella Ragonot: Colombia (?) 

579. indentella Dyar: Bermuda 



328 



XnsriTED STATES NATIONAL MUSEUM BULLETIN 207 



580. plorella Dyar: PanamS, 

vestilla (Dyar) 

581. stictella (Hampson), new comb.: Bahamas, 

West Indies 
uniformella Hampson 
granulella Hampson 

582. noniella Dyar: Panama 

171. AzAERA Schaus 

Calamophleps Dyar 

583. muciella Schaus: Costa Rica, Guatemala, 

Panama 
squalidella (Dyar) 

584. nodoses (Dyar), new comb.: Panama 

585. lophophera (Dyar), new comb.: Panama 

172. MooDNA Hulst 

586. ostrinella (Clemens): U. S., Canada 

obtusangulella (Ragonot) 
pelmculella Hulst 
587. bisinuella Hampson: Mexico, U. S. (Texas) 

173. ViTULA Ragonot 

588. edmandsae (Packard): Eastern U. S., Can- 

ada 
dentosella Ragonot 

589. edmandsae serratilineeUa Ragonot, new sta- 

tus: Western U. S., Canada 

590. lugubrella Ragonot, new comb.: Western 

U. S. (California) 

591. pinei Heinrich, new species: Western U. S. 

(Utah, Nevada) 

592. inanimella (Dyar), new comb.: Mexico, 

Guatemala 
ticitoa (Dyar) 

593. laura (Dyar), new comb.: Guatemala 

174. Manhatta Hulst 

Homigia Ragonot 

594. setonella (McDunnough), new comb.: U. S. 

(Utah), Canada (British Columbia) 

595. broweri Heinrich, new species: Eastern U.S. 

(Maine) 

175. Verina Heimich, new genus 

596. supplicella (Dyar), new comb.: Mexico, 

Guatemala, Panama, Brazil 

176. Vagobanta Heinrich, new genus 

597. divergens (Butler), new comb.: Chile 

177. MooDNELLA Heiurich, new genus 

598. paula Heinrich, new species: Guatemala, 

Brazil, Ajgentina 

178. VoLATicA Heim-ich, new genus 

599. pachytaeniella (Ragonot), new comb.: Bra- 

zil 

600. trinitatis Heinrich, new species: Trinidad 

179. Vezina Heinrich, new genus 

601. parasitaria Heinrich, new species: Argen- 

tina, Brazil 

180. Caudellia Dyar 

602. apyrella Dyar: Eastern U. S. (Maryland) 

603. albovittella Dyar: Eastern U. S. 

604. nigrella (Hulst), new comb.: Western U. S. 

arizonella (Walter) 



605. declivella (ZeUer), new comb.: PanamS,, 

Colombia 
animosella (Dyar) 

606. colorella (Dyar), new comb.: Panam& 

607. clara Heimich, new species: Puerto Rico 

181. MicROPHESTiA Dyar 

608. animalcula Dyar: Panama 

182. SosiPATRA Heinrich, new genus 

609. rileyella (Ragonot), new comb.: Western 

U. S., Mexico 

610. micaceella (Hampson): Mfeico 

611. anthophila (Dyar), new comb.: Western 

U. S. (Texas) 

612. thurberiae (Dyar), new comb.: Western U. S. 

613. nonparilella (Dyar), new comb.: Western 

U. S. (Arizona) 

614. majorella (Dyar), new comb.: M6xico 

615. divergens (Dyar): Panamd 

183. Bethulia Ragonot 

616. championella Ragonot: Guatemala 

184. RiBUA Heinrich 

617. innoxia Heinrich: Cuba 

618. contigua Heinrich, new species: Puerto Rico 

619. patriciella (Dyar), new comb.: Cuba 

185. Plodia Gu6n6e 

620. interpuncteUa (Hiibner): Cosmopolitan 

interpunctalis (Hiibner) 
zeae (Fitch) 
latercula (Hampson) 
glycinivora (Matsumura) 

621. dolorosa Dyar: Guatemala 

186. Anagasta Heim-ich, new genus 

622. kiihniella (Zeller): Cosmopolitan 

iuscojasciella (Ragonot) 
gitonella Druce 

187. Ephestia Gu^n^e 

Hyphantidium Scott 

623. elutella (Hiibner): Cosmopolitan 

elutea (Haworth) 
semirufa (Haworth) 
rufa (Haworth) 
sericarium (Scott) 
roxburghii Gregson 
unicolorella Staudinger 
amarella Dyar 

624. cautella (Walker): Cosmopolitan 

defectella (Walker) 

desuetella (Walker) 

cahiritella ZeUer 

passvlella Barrett 

jormosella (Wileman and South) 

625. figulileUa Gregson: Europe, Asia, Africa, 

Hawaii, Australia, North America (U. S., 
California), South America 

ficulella Barrett 

milleri Zeller 

figuliella Forbes 

Jigulella Curran 

venosella Turati 

emestinella Turati 



AMERICAN MOTHS OP THE SUBFAMILY PHYCITINAE 329 

188. NiCETiODES Schaus 191. Erelieva Heinrich, new genus 

626. apianella Schaus: Galapagos Islands 635. quantulella (Hulst), new comb.: Southern 

189. Varneria Dyar U. S. (Texas), West Indies 

627. postremella Dyar: Eastern U. S. santiagella (Dyar) 

628. nannodes Dyar: Panama 636. coca (Dyar), new comb.: Panama 

629. atrifasciella Barnes and McDunnough: .^^^ ^^ ^^^ 

Southern U. S. (Florida) ^^^^^ ^j^^^^^ 

630. dubia Heinrich, new species: Puerto Rico uncta (Dyar) 

190. EuRTTHMiA Ragonot 637. parvulella (Ely), new comb.: Eastern U. S. 

631. hospiteUa (Zeller): Southern and Western (Connecticut) 

U. S. 192. Cabnia Dyar 

spaldingella Dyar 638. myronella Dyar: Eastern U. S. 

632. hospiteUa yavapaella Dyar, new status: 193. Microphycita Dyar 

Western U. S. 639. titillella Dyar: Panama 

633. anguleUa Ely: Eastern U. S., Canada ^94. Rabiria Heinrich, new genus 

diffusella Ely 640. conops (Dyar), new comb.: Panama 

634. fumella Ely: Eastern U. S. (Connecticut) 

Species unplaced or unrecognized 

hrevistrigella Ragonot [Zophodia] 
came Dyar [Euzophera] 
cervinistrigalis Walker [Hypochalcia] 
clitellatella Ragonot [Hornigia] 
corrientellus Ragonot [Elasmopalpus] 
daedalella Ragonot [Euzophera] 
disticta Zeller [Psorosa] 
dulciella Hulst [Honora] 
Jamula Zeller [Myelois] 
flavicornella Ragonot [Phycitopsis] 
jormulella Schaus [Moodna] 
fuscifrontdla Zeller [Nephopteryx] 
gais Dyar [Euzophera] 
grossipunctella Ragonot [Myelois] 
hulstieUa Ragonot [Hypochalcia] 
injusella Zeller [Myelois] 
intextella Zeller [Euzophera] 
irichampa Dyar [Anthropteryx] 
megalopolis Hampson [Euzopherodes] 
nigricans Hulst [Salebria] 
olivella Hampson [Moodna] 
postflavida Dyar [Euzophera] 
putidella Schaus [Eucampyla] 
rinmea Dyar [Euzophera] 
subcanella Zeller [Zophodia] 



^ 
^ 



m^ 

UNITED STATES NATIONAL MUSEUM BULLETIN 207 




AMERICAN MOTHS 



OF THE SUBFAMILY 



PHYCITINAE 



By CARL HEINRICH 



iC 



SMITHSONIAN INSTITUTION • WASHINGTON, D. C, 1956 



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The scientific publications of the National Museum include two series, known, respec- 
tively, as Proceedings and Bulletin. 

The Proceedings series, begun in 1878, is intended primaiily as a medimn for the pub- 
lication of original papers, based on the collections of the National Museum, that set forth 
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The present work forms No. 207 of the Bulletin series. 

Remington Kellogg, 
Director, United States National Museum. 



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Contents 

Fsge 

Introduction v 

Family Phycitidae 1 

Subfamily Phycitinae 1 

Group I 1 

Group II 180 

Group III 310 

Genera and species unplaced, unrecognized, or referred from the Phycitinae ; . . . 312 

Checklist of American Phycitinae : 317 

Figures 331 

Indexes 567 



Introduction 



This paper completes a 25-year study of the New 
World moths of the subfamily Phycitinae. It is based 
chiefly on the collections in the United States National 
Museum and the Hulst collection, formerly at Rutgers 
University, supplemented by material from the Cornell 
and Canadian national collections and specimens — 
mostly tropical American — from the British Museum, 
the Janse collection, and the collections of several South 
American lepidopterists. 

Recognized and included in the classification are 194 
genera, 619 species, and 21 subspecies (local races). Of 
these, 60 genera, 81 species, and 8 races are described 
as new. The new species and races represent only a 
fraction of the undescribed material examined. The 
remainder consists mostly of females, chiefly from trop- 
ical America and without authentically associated males 
or host plants. Their description would have added 
nothing to om- scientific knowledge and the additional 
names would have been only a nuisance to other workers. 
Already too many names have been given such material. 

Acknowledgments 

A work of this kind could not be carried through 
without generous assistance from other entomologists. 
To each of them I owe a debt of gratitude: To Carl 
Muesebeck, Chief of the Division of Insect Investi- 
gation of the U. S. Bureau of Entomology and Plant 
Quarantine, for his support and encouragement at all 
stages of the project; toB.B.Pepper,State Entomologist 
of New Jersey, and John B. Schmitt for permission to 
examine the genitaUa of the Hulst types and for their 
courtesies to me at Rutgers University; to J. Bourgogne 
of the Muse.imi d'Histoire Naturelle, Paris, for the 
privilege of studying the genitalia of the Ragonot 
types of American species; to N. D. Riley and to 
W. H. T. Tams for the loan of unidentified tropical 
American Phycitinae from the British Museum (Natural 
History) and to Tams especially for photographs of 
many types and their genitalia ; to Martin Herring of the 
Zoologisches Aluseum der Universitat, Berlin, for the 
loan of Ragonot types; to W. T. M. Forbes, Department 
of Entomology, Cornell University, for the loan of his 
extensive collections from Puerto Rico, the Virgin Is- 
lands, and Surinam; to J. McDunnough and T. N. Free- 
man, Canadian National Museum, Ottawa, for the loan 
of Canadian specimens ; to A. J. T. Janse of the Transvaal 
Museum, Pretoria, South Africa, for the loan of South 
American Phycitinae from his collection and for much 
valuable information; to A. da Costa-Lima, Escola 
Nacional de Agronomia, Universidad Rural, Distrito 
Federal, Brazil, for the loan of Brazilian specimens; to 



Frank Morton Jones for a gift of Phycitinae collected at 
Martha's Vineyard, Mass. ; to John A. Comstock for a 
loan of southern California specimens ; to my colleagues 
at the U. S. National Museum — to J. F. Gates Clarke, 
for extensive notes on the phycitid types in the Museums 
of Paris, London, Oxford, and Berlin, and to Hahn 
Capps, for assistance in the tedious business of slide 
preparations. 

My greatest debt is to the artists of the Bureau of 
Entomology and Plant Quarantine for the drawings ac- 
companying this paper. Where genitalia are used in in- 
sect classification verbal descriptions are not enough. 
Figures must accompany and supplement them to give 
the reader a true picture of structural characters. The 
drawings in this paper were begun in 1930 by Eleanor 
A. Carlin and continued by her untU October 30, 1940, 
when she retired from the Bureau. From that time the 
drawings were made by Sara H. De Bord, who has made 
the majority of the drawings here published. Her con- 
tribution was of especial value because she was not only 
a capable artist but a trained entomologist as well, and 
her interest in the paper and her devotion to her share 
in it was so complete that she worked well on into her 
last illness (she was retired on disability August 12, 
1948, and died March 12, 1950). Since her death some 
drawings were made by Arthur Cushman and Addie 
Egbert, and the former did most of the assembling of 
the plates. The drawings were aU made imder my 
supervision and for any inaccuracies in them I am alone 
responsible. 

The indices were prepared by Mrs. Marguerite W. 
Poole. 

Abbreviation of references 

To conserve space and eliminate useless repetition, 
titles to certain publications frequently cited are here 
abbreviated as follows: 

The Ragonot "Monographic des Phycitidae et des 
GaUeriidae," published as vol. 7 (1893) and vol. 8 
(1901, completed by Hampson) of the Romanofif 
"Memoires sur les Lepidop teres," is cited as "Mono- 
graph, pt. 1," or "Monograph, pt. 2." 

Ragonot's "Diagnoses of North American Phycitidae 
and GaUeriidae," 1887, is cited as "N. Amer. Phyciti- 
dae," and his "Nouveau Genera et Especes de Phyci- 
tidae et GaUeriidae," 1888, as "Nouv. Gen." 

Walker's "List of the Specimens of Lepidopterous 
Insects in the Collections of the British Museiun," 
1854-66, is cited as "List." 

Hulst's "The Phycitidae of North America," pub- 
lished in the Transactions of the American Entomologi- 



VI 



INTRODUCTION 



cal Society, vol. 17, pp. 93-228, pis. 6-8, March-July 
1890, is cited as "Phycitidae of N. Amer." 

Barnes and McDunnough, "Contributions to the 
Natural History of the Lepidoptera of North America," 
vols. 2 (1913-1914) and 3 (1916-1917), is cited as 
"Contributions." 

McDunnough's "Check List of the Lepidoptera of 
Canada and the United States of America, Part II, 
Microlepidoptera," published in the Memoirs of the 
Southern California Academy of Science, vol. 2, No. 1, 
1939, is cited as "Check list." 

Forbes's "Lepidoptera of New York and Neighboring 
States," Cornell University Agricultural Experiment 
Station Memoir 68, 1923, is cited as "Cornell Mem. 68." 

The distributional records for species in this paper are 
obviously incomplete. They are based (with a few 
exceptions noted in the text) solely on specimens I 
have examined. This was the only safe procedure. 
So many misidentifications have been made in the past, 
even by lepidopterists of repute, that the records in 
literature can not be accepted merely on the authority 
of an author. Unless the specimens upon which his 
statements were based can be examined and the state- 
ments themselves verified, it is best to ignore them. 
By taking them simply on faith and repeating them we 
not only run the risk of perpetuating error, but do an 
injustice to past workers who did not have or could not 
use the evidence available to us. 

This caution applies with even greater force to 
"accepted" generic and specific synonymy. I have been 
very fortunate in being able to examine the genitaha 
of so many holo types and in having authentic specimens 
of most of the type species of described genera occurring 
in the New World. All unqualified synonymy in this 
paper is based upon genitaHc examination of such 
material. From the synonymy of some species common 
to both the New and Old Worlds I have omitted some 
names — chiefly of Old World synonyms — ^because I 
coxild not examine their types and had no certainty as 
to correctness of their synonymizing. Such omissions 
are discussed in the text. 

Classification and arrangement 

A general revision has a twofold purpose, a taxonomic 
and a practical one: To define accurately, to delineate 
as nicely as possible, and to name categories which, 
as far as our knowledge permits, represent objective 
realities in nature; and to arrange these categories in 
an order that permits their ready identification. Both 
purposes must be served if the revision is to have any 
value as a contribution to knowledge or to be of 
practical use to other workers. 

To satisfy both requirements I have adopted in this 
paper a dual classification: a definition and division 
into named categories of races, species, genera and 
subfamilies; and an artificial, unnamed division, 
between genus and subfamily, into groups of genera 
or, in a few instances within a single genus, into groups 
of species. 



The named categories themselves are more or less 
tentative. They are not adequate expressions of the 
truth. They are only approximations to it. As we 
learn more we shall have to amend or replace our defini- 
tions and the categories will come a Httle closer to the 
realities they represent. The names (except for 
homonyms) will always be available; but the concepts 
will change. There are several indications that taxo- 
nomic groupings between genus and subfamily may 
eventually be possible; and that when we have a clearer 
picture of host relations and larval characters, and more 
extensive collections from imexplored regions, we may 
be able to estabhsh tribes on a legitimate taxonomic 
basis; but at present this is impossible. What few 
definite derivations we can trace from genus to genus 
show that tribal groupings would cut across the lines 
of any artificial system we might be able to use. 

The artificial system here adopted (based on vena- 
tional characters) is proposed merely for key purposes. 
The keys themselves, except for the one separating the 
subfamilies, are in no true sense a part of the taxonomic 
system. They are keys, and keys only. They are 
intended merely to open a ready way to the descriptions 
of the genera and have been constructed on the assump- 
tion that they must work for all normal specimens. 
I hope so, for a key that wiU unlock a door only 75 
(or even 90) percent of the time is a tool of little worth. 
Here, a word of caution. No possible key wUl work 
for abnormal specimens. The worker in Phycitidae 
must be always on the alert for them, for the family 
contains an imusual number of freaks (chiefiy vena- 
tional abnormalities). Any one wishing to identify 
phycitids must resign himself to the tedium of dissection 
and slide making. Here, as in all the serious business of 
science, there is no easy way, no short cut to knowledge. 

The groupings of genera and species, prefaced by 
brief summaries of their common characters, which I 
have interposed within the text, are intended only to 
assist the reader and are not to be imderstood, in any 
sense, as definitions of taxonomic (tribal or subgeneric) 
groups. In a few instances they may be; but they are, 
in intent, only divisions of convenience. 

The only portion of the keys offered as a description 
of taxonomic imits is that separating the subfamilies 
Anerastiinae and Phycitinae. This long-established 
division of the family Phycitidae seems to be a sound 
one, and in the main the subfamilies themselves appear 
to be natural entities, although their definition leaves 
much to be desired. Probably when the Anerastiinae 
are thoroughly studied we may find other featin-es more 
constant than the reduced and concealed tongue. 
There may even be some shifting of genera across the 
subfamily lines. However, this is only hopeful antici- 
pation for the future and will remain so until the Old 
World genera and species of the family are thoroughly 
revised. For the present we shall have to content 
ourselves with an imperfect definition. 

The chart opposite this page shows my interpretation 
of the genera in their relation to each other and to the 
system based on venational characters. 



INTRODXrCTION 



ittL 



GENERA OF AMERICAN PHYCITINAE GROUPED ACCORDING TO GENITALIA AND VENATION 

Relationships on genitalic characters shown in horizontal arranfiement. Venational groupings are vertical 




Acrobasts 

Rhodofihaea 

Trachycera 

Anabasis 

Mildrlxla 

Sematoneun 

Hyslpyla 

Kemlptiloc< 

Crocldotnen 

Cunlberta 

Heras 

Adanarsa 

BIrlnus 

Berts I la 

Hypargyritt 

Chararica 



Protomoerbes 
Pseudod I vona 
Paramyelols 



Ectonyetols 
Apomyelols 
Anypstpyla 
Myelopsis 



Fundel la 

DIfundella 

Coplarthria 

Promylea 

AJiadelosamla 

Oasypyga 

(Rampylla 9 } 



Interjectio 
Catastia 



Ltpooraphi: 
^ Adelphia 
Tota 



Cborrer: 
TaccAa 



/ 
/ 

/ 



Euzopher 
Exuperlu 
Eulogla 



Da vara 

Sara'sota 

Plesmopoda 



Zamagirla 
Anegcephales 
-topsis 
Ancylostofflla 



Oioryctria 
Oryctometopii 



• Dresconopsts 
Cactobrosfs 

Atralafrida 

Sio«loa>ta 

Parol yea 

Salanibona 

Ereniberga 

Tucumanla 

Yosemltla 

Rumatha 

Cahela 

Cactob I ast i s 

Nana! a 

!rada 
01 yea 
Olycella 
Mel Kara 
Zophodia 
Rhagea 
Baphala 
Laetilia 



Cassiar>a 

Anderida 
Aptunga 



Canarsia 
Cabot i a 
Oncol at) is 

Hulstia 
Staudtnaei 



Eumys 



Phestlnii 

Nonia 

Mescinia 



Eurythmasi' 
Eurythmidii 
Wunderia 



Bethjila 
Soslpatra - 
Mlcrophestli 



vni 



INTRODUCnON 



The sequence of genera and of species within the 
genus here offered is an attempt to bring together in linear 
arrangement the forms showing affinities in structure 
and development. It is only an attempt and I shall 
quarrel with no one who objects to it. 

When I began this study I had hoped to write a 
monographic treatise and explore the phylogeny of the 
family, but I now find that I know so much less than I 
thought I did and that the accumulated knowledge of 



others is so meager that any attempt along these lines 
would be a vain and futile performance. 

We don't know what a primitive phycitid was like. 
We don't know which forms evolved from which, or 
how. We weren't there. We may surmise; but the 
guess of one ignoramus is as good as that of another, and 
there is nothing to be gained from either. I have had 
to be content with a mere revision. Would that it were 
more worthy. 

Carl Heineich 



Carl Heinrich (1880-1955) 

This monograph was written by Carl Heinrich in the months following his retirement from 
Government service in 1949. Upon it he focus: I the extensive knowledge gained during his 36 
years as entomologist with the U. S. Department of Agriculture. Its publication, toward which 
the Department of Agriculture has contributed substantially, was undertaken in 1954, and the 
author had completed his review of the galley proofs at the time of his death, age 75, on May 31, 
1955. 

A biographical memoir of Carl Heinrich and a bibhography of his scientific writings appeared 
in the Proceedings of the Entomological Society of Washington for October 1955 (vol. 57, No. 5, 
pp. 249-255). In addition to the present bulletin, the U. S. National Museum has published a 
number of his papers, of which several, as noted, are now out of priat: 



1921. On some forest Lepidoptera with descrip- 
tions of new species, larvae, and pupae. 
Proc. U. S. Nat. Mus. No. 2305, vol. 57, 
pp. 53-96, 13 pis., June 17, 1920. (Out 
of print.) 

1923. Revision of the North American moths of 
the subfamily Eucosminae of the family 
Olethreutidae. U. S. Nat. Mus. Bull. 
123, iv+298 pp., 1 fig., 59 pis., Apr. 12, 
1923. (Out of print.) 

1926. Revision of the North American moths of 

the subfamihes Laspeyresiinae and Ole- 
threutinae. U. S. Nat. Mus. Bull. 132, 
iv+216 pp., 2 figs., 76 pis., Feb. 2, 1926. 
(Out of print.) 

1927. The American moths of the genus Diatraea 

allies. Proc. U. S. Nat. Mus. No. 2691, 
vol. 71, Art. 19, 48, pp., 20 pis., Aug. 23, 
1927. Joint authorship with H. G. Dyar. 
(Out of print.) 



1929. Notes on some North American moths of the 
subfamily Eucosminae. Proc. U. S. Nat. 
Mus. No. 2779, vol. 75, Art. 8, 23 pp., 5 
pis., Apr. 5, 1929. 

1932. Notes on and descriptions of some American 
moths. Proc. U. S. Nat. Mus. No. 2879, 
vol. 79, Art. 13, 16 pp., 1 fig., 7 pis., Aug. 
10, 1931. 

1938. Moths of the genus Rupela (Pyralididae. 
Schoenohiinae). Proc. TJ. S. Nat. Mus. 
No. 3019, vol. 84, pp. 355-388. 12 pis., 
July 3, 1937. 

1940. The cactus-feeding Phycitinae: A contribu- 
tion toward a revision of the American 
pyralidoid moths of the family Phycitidae. 
Proc. U. S. Nat. Mus. No. 3053, vol. 86, 
pp. 331^13, 29 pis.. Mar. 16, 1939. 

1945. The genus Fundella Zeller: A contribution 
toward a revision of the American Pyrali- 
doid moths of the family Phycitidae. 
Proc. U. S. Nat. Mus. No. 3190, vol. 96, 
pp. 105-114. 3 pis.. May 18, 1945. 



American Moths of the Subfamily Phycitinae 



Family Phycitidae encircling or partially encircling the tubercle of seta 

lib on mesothorax and a smilar ring encircling the 

Moth: Labial palpus well developed. Maxillary tubercle of seta III on eighth abdominal segment (this 

palpus present, variously developed, rarely vestigial. character absent from the following genera of the 

Tongue well developed or reduced, rarely absent; when American Phycitinae; Etiella, Oryctometopia, Ulophora, 

distinguishable, basal portion scaled. Forewing entire Rotruda, Rhagea, and Unadilla). Prologs normal; 

(not divided); 11 veins or less; vein 7 always absent; 8 crochets in a complete circle. 

and 9 stalked or united; Ic absent (represented only by a ^he subfamilies of Phycitidae are separated by the 

fold or crease in the wing membrane) ; no areole. Hind following key: 
wing with 8 veins or less ; vein 8 closely approximate or 

contiguous to or anastomosing or completely fused with Tongue normally well developed ; if sometimes reduced, 

7 beyond cell; Ic always present; a fringe of pecten on not concealed between the labial palpi (except in 

lower median vein at base ; frenulum of female simple Cactoblastis ') ; ocelli always present . . Phycitinae 

^*Stl:'witr primary setae only; two setae on tongue reduced or vestigial; when merely reduced, 

prespiracular shield of prothorax; setae IV and V ap- concealed between the labial palpi; if sometimes 

pro^dmate and under the spiracle on abdominal seg- «f «s«d between the palpi (Bandera) then oceUi 

ments 1 to 8; normally a sclerotized, pigmented ring absent Anerastimae 

' In Cactoblastis the aborted tongue is completely concealed by Subfamilv Phvcitinae 

the broadly scaled basal segments of the labial palpi. However, J * 

the cenitalia, habitus and larval afiSnities show that Cactoblastis ™, , * ii_ i_j •^ t>u 

is aftrue phycitine and must be placed with the other closely The larger groups of the subfamily Phycitinae are 

related genera of the cactus-feeding Phycitinae. separated by the following key: 

Key to the larger groups of Phycitinae 

Hind wing with veins 3 and 4 both present Group I 

Hind wing with vein 3 present, 4 absent Group II 

Hind wing with veins 3 and 4 both absent Group III 



Group I 

[Hind wing with veins 3 and 4 both present] 
Keys to the Venational Divisions and Genera of Group I 

Hind wing with vein 3 appreciably before the outer angle of cell ; cell less than one-half the 

wing length Venational division A 

Cryptoblabes (p. 10) 

Hind wing with vein 3 closely approxinaate to or from the angle of the cell (rarely shortly 
stalked with 4-5) ; 7 and 8 approximate, contiguous, or shortly and weakly anastomosed 
beyond cell; cell at lower angle nearly half as long as wing; if shorter, then vein 3 of 
moderate length and the free (divergent) part of 3 decidedly shorter than vein 2. If 
vein 3 sometimes appreciably before outer angle of cell (Acrobasis), then cell one-half 
the wing length Venational division B (key, p. 2) 

Hind wing with vein 3 closely approximate to or from the angle of cell; veins 7-8 solidly 
anastomosed beyond cell for at least three-fourths of their lengths. 

Venational division C (key, p. 7) 



■UNITED STATES NATIONAL MUSEUM BULLETIN 207 

Hind wing with vein 3 closely approximate to or from the angle of cell; veins 7-8 approxi- 
mate or very weakly and shortly anastomosed beyond cell; 2 and 3 both long, the 
divergent element of 3 nearly as long as 2; cell distinctly less than halt the length of 
wing Venational division D (key, p. 8) 

Venational division B 

1. Hind wing with discoceUular vein oblique 2 

Hind wing with discoceUular vein ciirved 3 

2. Forewing with subbasal ridge of raised scales; hind wing with cell somewhat less than 

one-third the length of the wing; eighth abdominal segment of male with midven- 

tral hair tuft Mildrixia (p. 26) 

Forewing smooth; hind wing with cell one-fifth the length of the wing; eighth abdominal 
segment of male with sternite developed as a sclerotized, digitate pocket. 

Drescoma (p. 88) 

3. Hind wing with vein 3 from before, but near, lower outer angle of cell 4 

Hind wing with vein 3 from the angle or the stalk of veins 4-5 8 

4. Hind wing with veins 4-5 connate; vein 6 of forewing always straight 5 

Hind wing with veins 4-5 approximate, contiguous, anastomosed or stalked for a short 

distance from ceU, if sometimes connate (on individuals of Hypsipyla) vein 6 of fore- 
wing slightly bent towards base 6 

5. Basal segment of male antenna triangulate Acrobasis (p. 11) 

Basal segment of male antenna simple (cyHndrical) Rhodophaea " (p. 24) 

6. Forewing with subbasal ridge of raised scales; antenna of male pubescent (cUia distinctly 

shorter than width of shaft) Anabasis (p. 25) 

Forewing smooth; antenna of male ciHate (cUia distinctly longer than width of shaft) . 7 

7. Forewing with vein 6 straight, remote from veins 8-9 at base . . Sematoneura (p. 27) 
Forewing with vein 6 bent, shortly separated from 8-9 at base . . . Hypsipyla (p. 27) 

8. Vestiture of head, thorax, labial palpi, and femora a mixture of scales and hairs; male with 

harpe short (stubby), clasper absent, apical process of gnathos an inverted heart- 
shaped lobe with short, slender spine; female with ventral surface of genitalia sclero- 
tized throughout its length, ductus seminalis from near junction of bursa and ductus 

bursae Polopeustis (p. 120) 

Vestiture entirely of scales; or, if occasionally mixed with hair (Sarata atrella), male with 
harpe elongate, clasper present, knoblike, apical process an elongate, stout hook; 
female with ductus bursae unsclerotized, ductus seminaHs from bursa remote from 
junction of bursa and ductus bursae 9 

9. Forewing with subbasal ridge of raised scales 10 

Forewing smooth 16 

10. Hind wing with vein 3 from the stalk of veins 4-5 Passadena (p. 175) 

Hind wing with vein 3 approximate to but not from the stalk of veins 4-5 11 

11. Labial palpus porrect, beaklike Etiella (p. 98) 

Labial palpus oblique or upturned 12 

12. Forewingwith vein 6 from the upper angle of the cell, bent towards base 

Hypargyria (p. 37) 
Forewing with vein 6 from well below the angle of cell, straight 13 

13. Forewing with veins 4 and 5 sHghtly separated at base and approximate for a short dis- 

tance beyond ■,-,, 14 

Forewing with vein^ 4 and 5 connate or stalked ; if sometimes shortly separated at base 
not approximate beyond 15 



' The genus Trachycera is omitted from our key as the male is unknown except from Ragonot's descrip- 
tion. The female before me has the venation of Rhodophaea. It is distinguished from other females of 
Division B in having two scobinate, cuplike signa similar to those of Davara in division D. (See p. 25.)]"; 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITENAE 

14. Hind wing with veins 7-8 anastomosed beyond cell; maxillary palpus of male in the form 

of an aigrette Immyrla (p. Ill) 

Hind wing with veins 7-8 approximate beyond cell; maxillary palpus of male squamous. 

Ulophora (p. 176) 

15. Labial palpus oblique, second segment on male grooved; male with eighth abdominal 

segment simple; female with genital opening simple Ortholepis (p. 119) 

Labial palpus upturned, second segment on male not grooved; eighth abdominal segment 
of male with compound tufts ; female with strongly sclerotized plate at genital open- 
ing attached to supplemental eighth-segment coUar Tlascala (p. 133) 

16. Labial palpus porrect, beaklike; male with aedeagus flanged and strongly spined; female 

with ductus seminaUs from bursa 17 

Labial palpus obhque or upturned ; if sometimes appearing porrect {Stylopalpia) due to 
long, deflected third segment, male with aedeagus simple; female with ductus semi- 
nalis from ductus bursae 21 

17. Forewing with veins 4-5 approximate for a short distance from cell 18 

Forewing with veins 4-5 stalked 19 

18. Labial palpi extending at least twice the length of head beyond it; harpe of male elongate; 

female with ductus bursae much longer than bursa Pima (p. 101) 

Labial palpi extending little more than the length of head beyond it; harpe of male short; 
female with ductus bursae little, if any, longer than bursa . . Interjectio (p. 106) 

19. Male with second segment of labial palpus grooved on inner side; female with ductus 

seminaUs from bursa remote from junction of bursa and ductus bursae . Sarata (p. 159) 

Male with second segment of labial palpus not grooved; female with ductus seminalis 

from bursa near junction of bursa and ductus bursae 20 

20. Female with a strongly sclerotized plate behind genital opening . . Philodemia (p. 165) 
Female without sclerotization at genital opening Lipographis (p. 166) 

21 . Hind wing with vein 3 from the stalk of 4-5 or closely approximate to it for some distance 

fromlowerouterangleof cell; vein 2 always rather near the angle 22 

Hind wing with vein 3 connate with the stalk of veins 4-5 or connected with it at base by 
a very short spur; if sometimes approximate to the stalk of 4-5 (Megarthria, Acron- 
cosa), vein 2 always from well before lower outer angle of cell; or, if vein 3 some- 
times shortly fused with the stalk of veins 4-5 {Actrix), male with apical process of 
gnathos developed as a square or inverted heart-shaped plate and female with 
caudal half of bursa copulatrix densely spinose 28 

22. Hind wing of male with anal angle folded into a pocket ; female with strong sclerotizations 

behind or surrounding genital opening of genitaha 23 

Hind wing of male without folded pocket at anal angle; genital opening of female 
simple 24 

23. Male genitaha with uncus hammer-clawed (long, curved, constricted at middle and 

broadly divided at apex); female with bursa containing strongly sclerotized folds 

or stoutly spined bands Fundella (p. 59) 

Male genitaha with uncus otherwise (sometimes broadly divided at apex but the divided 
elements small and spinelike and the middle of uncus not appreciably constricted) ; 
bm-sa sometimes with a small granulate patch but otherwise membranous and 
unarmed Difundella (p. 62) 

24. Hind wing with vein 3 approximate to the stalk of veins 4-5 for some distance from outer 

angle of cell Scorylus (p. 72) 

Hind wing with vein 3 from the stalk of veins 4-5 25 

25. Male genitalia with transtUla a sinuate, sclerotized scobinate band involved with gnathos 

and with a long free spine involved with aneUus; female with ductus seminahs from 

ductus bursae 26 

Male genitalia with transtilla incomplete or absent; female with ductus seminalis from 
bm-sa 27 

26. Maxillary palpus of male filiform Coptarthria (p. 64) 

Maxillary palpus of male squamous Anadelosemia (p. 67) 



UNITED STATES NATIONAL MUSEXnH BULLETIN 207 

27. Male genitalia with divided element of incomplete transtilla strongly sclerotized, gnathos 

absent; female with ductus bursae partially sclerotized Gabinius (p. 84) 

Male genitalia with transtilla absent, gnathos well developed and with apical process 
a strong hook; female with ductus bursae membranous throughout. 

Ceracanthia (p. 85) 

28. Male genitaha with transtilla a sinuate sclerotized band involved with gnathos; female 

with bursa small, membranous and ductus bursae much longer than bursa, signum 
(if present) a small granulate patch or small plate with single miaute thorn, genital 
opening narrow (the ductus bursae never expanded into a widened opening) ; if bursa 
sometimes large and ductus bursae proportionally shorter {Bampylla), collar of 

eighth segment modified 29 

Male genitalia without transtUla or, if present, otherwise; female genitalia never as 
above in all details 31 

29. Forewing with veins 4-5 approximate for a short distance from cell; hind wing with 

vein 2 from well before lower outer angle of cell 30 

Forewing with veins 4-5 stalked; hind wing with vein 2 from near outer angle of cell. 

Dasypyga (p. 69) 

30. Female with ductus seminahs from ductus bursae Promylea (p. 65) 

Female with ductus seminalis from bursa copulatrix Rampylla (p. 70) 

(males: venation group D, couplet 8) 

31. Male with transtilla of genitalia complete and strongly sclerotized; or, if incomplete, the 

elements enlarged, strongly sclerotized and modified; when complete not in the 
form of a squarish plate. Female with a single signum developed as a small, scobin- 
ate or grantdate cup or patch or (Adanarsa) as a single short, stout thorn; genital 

opening always broad 32 

Male genitalia with transtiUa incomplete or absent. Female with signa or signum, if 
present, otherwise developed. If transtilla complete then weakly sclerotized or 
developed as a square plate; and if signum of female a small scobiaate patch {Megar- 
thria alpha) genital opening narrow 39 

32. Forewing with vein 6 bent towards base 33 

Forewing with vein 6 straight 36 

33. Transtilla ofmale complete, strongly arched and with median area forked 34 

TranstiUa of male complete, but not arched nor with median area forked 35 

34. Antenna of male with shaft unipectinate. Ductus bursae of female very short, less than 

one-half as long as bursa, scobinate-granulate and more or less sclerotized but not 

transversely wrinkled Hemiptilocera (p. 30) 

Antenna of male with shaft pubescent. Ductus bursae of female longer, about half as 
long as bursa and with strong, sclerotized wrinkling before genital opening. 

Crocidomera (p.32) 

35. Maxillary palpus squamous. Forewing of male with costal fold and a fovea in cell 

slightly beyond base. Eighth abdominal segment with stemite developed as a 
sclerotized pocket Heras (p.34) 

(male only, female unknown) 

Maxillary palpus filiform. Forewing of male without costal fold or fovea. Eighth 

abdominal segment simple Birinus (p. 36) 

(male only, female unknown) 

36. Male antenna with shaft unipectinate; transtilla incomplete, its elements long, stout with 

their apices broadly and irregularly developed and hooked. Female with a large 
semicircular sclerotized and scobinate plate on membrane behind genital opening. 

Bertelia (p. 36) 

Male antenna with shaft pubescent; transtilla complete. Sclerotized plate on membrane 

behind genital opening of female, if present, not semicircular 37 

37. Hind wing with veins 7-8 anastomosed for a short distance beyond cell. Male with trans- 

tilla strongly arced but with its median area bearing a smooth narrow crosspiece, not 
forked. Female with signum a single, short, stout, hooked thorn . Adanarsa (p. 35) 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 

Hind wing with veins 7-8 approximate for a short distance from cell. Male transtilla 
otherwise. Female with signum a single, small, scobinate or granulate, cup-shaped 
patch 38 

38. Male with apical process of gnathos a stout hook. Female with ductus bursae appreci- 

ably longer than bursa; ductus seminalis from ductus bursae . . Cuniberta (p. 34) 
Male without projecting apical process, the lateral arms supporting a thinly sclerotized 
subanal plate. Female with ductus bursae much shorter than bursa; ductus semi- 
nalis from lobe of bursa near junction of bursa and ductus bursae . Chararica (p. 38) 

39. Male with sternite of eighth abdominal segment developed as a sclerotized pocket. 

Female with signum a small, depressed, granulate-scobinate patch; or, if signum 
absent, ductus bursae narrow throughout and sclerotized for about one-third of its 
length from shortly beyond its junction with bursa, the sclerotized portion sharply 

bent over itself Megarthria (p. 86) 

Male with eighth abdominal sternite not developed as a sclerotized pocket. Female 
with signum (or signa) developed as sclerotized and strongly spined plates or bands, 
or entirely absent. If signa absent, ductus bursae not as above. If sometimes 
(Olybria) a single more or less strongly sclerotized band in bursa at junction of bursa 
and ductus bursae, the band finely serrate or edged with short spines 40 

40. Antenna of male with shaft unipectinate. Female bursa \vith a single strongly spined 

signum Monoptilota (p. 89) 

Antenna of male with shaft pubescent. Female bursa with two signa or none ... 41 

41. Male genitalia with a pair of long, strong, sclerotized arms from the ventrolateral angles 

of uncus; gnathos absent. Female with two signa consisting of stoutly and coarsely 
spined bands; ductus bursae strongly sclerotized, fattened (ribbonlike). 

Caristanius (p. 97) 
Male genitalia with uncus otherwise; gnathos present and well developed. Female with- 
out signa; or, when present, consisting of two strongly spined bands or plates; when 
signa are present, ductus bursae not as above, if partially flattened and ribbonlike 
then very narrow 42 

42. Male genitalia with aedeagus expanding to lateral, flanged projections before apex, the 

flanges each bearing a cluster of strong spines. Female with ductus seminalis from 

ductus bursae Stylopalpia (p. 140) 

Male genitalia with aedeagus otherwise, if sometimes spined (Pyla), not flanged before 
apex. Female with ductus seminalis from bursa copulatrLx 43 

43. Maxillary palpus vestigial 44 

Maxillary palpus squamous, or filiform, or (on some males) in the form of an aigrette . 45 

44. Penis of male unarmed except for a small cluster of very weak, short, slender spines. 

Female with ovipositor strongly sclerotized Telethusia (p. 136) 

Penis of male armed with a single, strong comutus. Female with ovipositor normal (not 
strongly sclerotized) Phobus (p. 138) 

45. Forewing with some rough (raised) scaling in median area beyond outer border of ante- 

median line, but without subbasal ridge of raised scales Tulsa (p. 134) 

Forewing smooth 45 

46. Fore tibia with a long inner and a short outer claw Acroncosa (p. 174) 

Fore tibia otherwise 47 

47. Hind wing with veins 7-8 distinctly anastomosed for about half their lengths (more than 

the free length of vein 8). Male with apical process of gnathos a broad shield 

without central terminal spine Actrix (p. 139) 

Hind wing with veins 7 and 8 approximate ; if sometimes contiguous or anastomosed be- 
yond cell, then weakly and very shortly so. Male with apical process of gnathos 
otherwise 48 

48. Male genitalia with a pair of straight, strongly sclerotized arms [not to be confused with 

similar projections from vinculum in Nephopteryx crassifasciella] projecting back- 
ward from lower, posterior angles of tegumen. Female with ductus bursae of gen- 
italia flattened (ribbonlike), waved (twice bent) and sclerotized throughout, the 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 

sclerotization developed at genital opening into a stout, squarish ventral 

plate Olybria (p. 113) 

Male genitalia without projecting arms from tegumen. Ductus bursae of female 
otherwise 49 

49. Harpe of male genitalia with a long hair brush from inner surface, along lower edge of 
basal half of sclerotized costa. Bursa copulatrix of female with the lobe giving ofi' 
ductus seminalis strongly sclerotized; or most of dorsal area of bursa strongly and 
smoothly sclerotized; or bursa with conspicuous, round or oval, strongly pigmented 
and sclerotized, densely granulate patches [absent in Nephopteryx suhcaesiella (fig. 

826)], the bursa otherwise spinose over its membranous areas 50 

Harpe of male without such hair brush. Bursa of female not as above 53 

60. Male with sacculus of harpe strongly produced at apex. Female with most of dorsal 

sm-face of bursa strongly and smoothly sclerotized Glyptocera (p. 100) 

Male with sacculus of harpe simple (not produced at apex) . Female with only the lobe 
giving off ductus seminaUs sclerotized: or biu-sa with granulate sclerotized 
patches 51 

51. Penis of male armed with numerous strong, slender spines. Female with lobe of bursa 

giving off ductus seminalis strongly sclerotized Oreana (p. 112) 

Penis of male armed with two stout spines or (very rarely) one spine. Bursa of female 
with granulate patches, the lobe giving off ductus seminalis not sclerotized ... 52 

52. Harpe of male genitalia with an appressed, stout, thorny or serrate clasper. Female 

with ductus bursae unsclerotized adjacent to bm-sa, but with strong sclerotization 

at genital opening Meroptera (p. 121) 

Harpe of male with clasper digitate, slender, simple (without spining). Female with 
ductus bursae sclerotized along ventral surface from junction with bursa, the sclero- 
tization terminating before genital opening, the latter simple (unsclerotized) . 

Nephopteryx (p. 123) 

53. Forewing with veins 4-5 stalked for nearly half their lengths Tacoma (p. 178) 

Forewing with veins 4-5 not stalked (slightly separated or closely approximate at 

cell) 54 

54. Male genitalia with transtiUa complete but very weakly sclerotized (a narrow angulate 

band); aedeagus slender, elongate, not spined or divided. Female genitalia with 
bursa entirely membranous and smooth; ductus bursae rather narrow, tubular and 
sclerotized throughout, nowhere appreciably widened; genital opening narrow (no 

wider than narrowest part of ductus bursae) Tota (p. 170) 

Male genitalia with transtUla incomplete or absent; aedeagus moderately broad to stout, 
if sometimes slender and elongate then spined or partially divided or {Chorrera) 
vinculum with produced lateral lobes from terminal margin. Female with or with- 
out signa in bursa; if without signa, bursa spinose or with same sclerotizations 
adjacent to ductus bursae; if bursa entirely membranous and smooth, genital 
opening decidedly broadened 55 

55. Male with aedeagus of genitalia spined or partially divided; if sometimes simple, then a 

strong knoblike and spinose projection from harpe at base of costa. Female without 
signa in bursa, the latter usually simple (unsclerotized and smooth), but sometimes 
with sclerotized, convolute, longitudinal bands near jimction of biu-sa and extending 

into the ductus, the bursa never spinose Pyla (p. 142) 

Male with aedeagus neither spined nor divided; if harpe sometimes with projection from 
base of costa, the latter neither knoblike nor spinose. Female with or without signa 
in bursa; if without signa, the bursa spinose (at lease partially so) or sclerotized at 
junction of bursa and ductus bursae, but such sclerotization not in the form of 
convolute, longitudinal bands 56 

56. Male with maxillary palpus squamous or filiform. Female without signa; a serrate or 

minutely spined sclerotization at junction of bursa and ductus bursae, the latter 
broad and strongly sclerotized or with a pair of elongate sclerotized plates behind 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAB 

genital opening; if signa present, the ductus bursae very narrow and expanded 
into a membranous globe shortly before genital opening 57 

Male with maxillary palpus in the form of an aigrette. Female with or without signa; 

bursa and ductus bursae not as above 59 

67. Male genitalia with aedeagus stout; penis armed with a single, long stout spine. Bursa 
of female genitaUa without signa 58 

Male genitaha with aedeagus very slender; penis unarmed. Bui-sa of female containing 
two signa developed as opposed, longitudinal bands bearing a row of short stout 
spines Chorrera (p. 177) 

58. MaxiUary palpus of male minute, filiform. Female without sclerotized plate or plates 

behind genital opening Ambesa (p. 108) 

Maxillary palpus of male squamous (broadly scaled). Female with a pair of elongate 
sclerotized plates on inner dorsal surface of ductus bursae behind genital 
opening Catastia (p. 110) 

59. Male with sacculus of harpe slightly produced at apex. Female with ductus bursae of 

genitaha cylindrical; narrow except at genital opening; appreciably longer than 
length of bursa; sclerotized for haK its length from junction with bursa, the sclero- 

tization longitudinally ribbed Elasmopalpus (p. 172) 

Male with sacculus of harpe not produced at apex. Ductus bursae of female other- 
wise 60 

60. Hind wing with vein 2 from before but rather near outer angle of cell 61 

Hind wing with vein 2 from well before outer angle of cell 62 

61. Male with penis armed with a single stout spine. Female with ductus brusae of genitaUa 

sclerotized (at least towards genital opening). Forewing with veins 8-9 stalked for 
less than two-thirds of their lengths. Hind wing with veins 4-5 stalked for approxi- 
mately one-haK their lengths Salebriaria (p. 115) 

Male with penis unarmed. Female with ductus bm-sae cartilagenous, except at its 
junction with bursa. Forewing with veins 8-9 stalked for over two-thirds of their 
lengths. Hind wing with veins 4-5 stalked for over three-fourths of their 
lengths Quasisalebria (p. 118) 

62. Male genitaha with a strong, straight or curved arm from base of costa of harpe; penis 

imarmed. Female genitaUa with ductus seminaUs from bursa adjacent to jimction 

of bursa and ductus biu^ae (signa present) Adelphia (p. 168) 

Male genitaUa without projecting arm from base of costa of harpe; penis armed with a 
single strong spine. Female with ductus seminaUs from bursa remote from junction 
of bursa and ductus bursae (signa present or absent) 63 

63. Male with cornutus on penis a long straight spine, over half as long as aedeagus. Female 

with bursa densely spinose, signa absent; ductus seminaUs from near anterior end of 

bursa Salebriacus (p. 114) 

Male with cornutus on penis a short, curved spine, somewhat less than one-third as long 
as aedeagus. Female with bursa smooth except for strongly spined signa (and in 
one species a strongly spined coUar at middle of bursa) ; ductus seminalis from bursa 
adjacent to one of the signa Ufa (p. 170) 

Venational division C 

1. Hind wing with discocellular vein straight, vertical Homoeographa (p. 135) 

Hind wing with discocellular vein straight, obUque Atheloca (p. 81) 

Hind wing with discocellular vein curved 2 

2. Male with uncus of genitalia more or less spoon-shaped (the lateral margins deeply 

concave at middle). Female with signa present, consisting of 2 or more sclerotized 

disks or series of contiguous, blunt thorns 3 

Male with uncus triangulate or subtriangulate. Female with or without signa; if present 
not as above 4 



tOSriTED STATES NATIONAL MUSEUM BULLETIN 207 

3. Male with costal fold on forewing; shaft of antenna notched at base . Diatomocera (p. 50) 
Male without costal fold on forewing; antennal shaft not notched . Pseudocabima (p. 53) 

4. Hind wing with vein 3 from the stalk of veins 4-5 Anypsipyla (p. 42) 

Hind wing with vein 3 from the angle of the cell; if sometimes approximate to the stalk 

of veins 4-5 for a short distance, never actually from it 5 

5. Maxillary palpi of both sexes filiform. Male with complete transtilla 6 

Maxillary palpi squamous. Transtilla of male incomplete 8 

6. Male with antennal shaft pubescent. Female with ductus seminalis from near middle or 

towards anterior (closed) end of bursa 7 

Male with antennal shaft shortly ciliate (ciha as long as width of shaft). Female with 
ductus seminalis from bursa near junction of bursa and ductus bursae. 

EctomyeloiB (p. 43) 

7. Forewing with vein 2 from near outer angle of cell. Male with transtilla weakly sclerotized 

(a thin band or sub-triangulate plate); apical process of gnathos broadly U- 

shaped Myelopsis (p. 40) 

Forewing with vein 2 from well before the angle. Male with transtUla strongly sclero- 
tized and arched ; apical process of gnathos a simple, stout hook . . Apomyelois (p. 42) 

8. Hind wing with cell moderately long (from a little over to slightly less than one-half the 

length of the wing) 9 

Hind wing with cell short (not over one-third the length of wing). . Protomoerbes (p. 49) 

9. Forewing with vein 2 from near lower outer angle of cell; vein 10 from the cell. Male 

with eighth abdominal segment simple. Female with signum in bursa (a cluster of 
coarse scobinations) ; a sclerotized plate behind genital opening (on inner dorsal 

surface of ductus bursae) Paramyelois (p. 46) 

Forewing with vein 2 from well before the angle; vein 10 from the stalk of veins 8-9. 
Male with a pair of ventrolateral hair tufts on eighth abdominal segment. Female 
with bm^a and ductus bursae simple (membranous throughout, with neither signxmi 
in bursa nor plate behind genital opening) Pseudodivona (p. 48) 

Venational division D 

1. Normal dark discal spots on forewing at end of cell replaced by a conspicuous white spot 

or Une (obscured only on clarioralis) on discocellular vein. [Male genitalia with 
costal area of harpe broadly sclerotized and produced at apex; clasper present, 
erect ; penis armed with numerous, straight spines. Female with signa developed as 
2 or 3 clusters of strong, slender spines; ductus bursae flattened, strongly sclerotized 
over most of its length, the sclerotization terminating just before simple genital 

opening] Dioryctria (p. 149) 

No such white spot on discocellular vein 2 

2. Forewing with ridge of raised scales preceding antemedial line 3 

Forewing smooth ; if sometimes with a few roughened scales, no such subbasal ridge. . 4 

3. Forewing with veins 4-5 closely approximate for a short distance from cell. Hind 

wing with cell less than one-fourth the length of wing. Maxillary palpus of male 

in the form of an aigrette Zamagiria (p. 90) 

Forewing with veins 4-5 connate or very shortly stalked. Hind wing with cell slightly 
less than one-third the length of wing. Maxillary palpus of male subsquamous. 

Anegcephalesis (p. 93) 

4. Male genitalia with uncus and tegumen greatly reduced; uncus a narrow, weakly sclero- 

tized, angulate band. Female with a single signum in bursa, consisting of a small, 

sclerotized plate supporting a very short thomlike spine Peadus (p. 83) 

Male genitalia with uncus and tegumen well developed. Female with signum or signa 
(if present) otherwise 5 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 

5. Male genitalia with transtilla complete, developed as a narrow, slightly arched band, 

attached to harpes only by membrane. Female with signum a cluster of bluntly 
pointed thorns, more or less surromided by fine scobinations or strongly pigmented 

granulations Hyalospila (p. 56) 

Male genitalia incomplete or absent; if complete (Magiriopsis) not a narrow band. 
Female with signum or signa (if present) otherwise 6 

6. Hind wing with discoceUular vein mcomplete Fulrada (p. 71) 

(based on male ; female unknown) 
HLod wing with discoceUular vein complete 7 

7. Hind wing with cell very short (about one-fifth the length of wing). Male genitalia 

with lateral arms of gnathos broad, expanded and curled at their extremities. 

Praedonula (p. 82) 

Hind wing with cell longer (from one-fourth to one-third the length of wing). Gnathos 

of male otherwise 8 

8. Hind wing of male with anal area (involving vein la) thickened and folded, forming 

a produced pocket; underside of wing with roughened scale or hair tufts on some 

of the veins Rampylla (p. 70) 

(females: Venation Group B, couplet 30) 
Hind wing of male without such modification 9 

9. Male with uncus of genitalia bifid. Female with one or two small signa developed 

as granulate depressions in bursa; if signa sometimes absent (some species of 

Piesmopoda) , bursa membranous 10 

Uncus of male undivided (triangulate or pentagonal). Signa of female developed as 
strongly, spined bands or plates; if sometimes absent (Ancylostomia) , bursa weakly 
but extensively sclerotized 12 

10. Gnathos complete, a thin, weakly sclerotized, transverse band. Female with two 

signa in bursa 11 

Gnathos incomplete, the lateral arms strong, broad; between their separated apices a 
well sclerotized anal plate. Female with one signum or none . . Piesmopoda (p. 77) 

11. Forewing with vein 6 from below upper angle of cell, separated at base from the stalk 

of veins 8-9. Male genitalia with a stout free spine associated with anellus. 

Davara (p. 73) 

Fore^ving with vein 6 from upper angle of cell, connate with the stalk of veins 8-9. 

Male genitalia without free spine associated with anellus .... Sarasota (p. 76) 

12. Male antenna unipectinate for basal half of shaft, shortly ciliate beyond. Female 

with two signa developed as strongly spined plates Magiriopsis (p. 94) 

Male antenna with shaft pubescent. Female with one signum or none 13 

13. Male with maxillary palpus in the form of an aigrette; eighth abdominal segment with 

compound ventral scale and hair tufts. Female genitalia without signum; the 

bursa copulatris weakly sclerotized throughout Ancylostomia (p. 95) 

Male with maxillary palpus squamous; eighth abdominal segment with paired ventro- 
lateral hair tufts. Female with signum, consisting of a single round, curved plate, 
densely armed with long stiff spines ; bursa otherwise membranous. 

Oryctometopia (p. 158) 



10 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



Genus 1: Cryptoblabes 

[Venational division A. Hind wing with vein 3 distinctly before 
lower outer angle of cell; 7 and 8 approximate, or weakly and 
shortly anastomosed beyond cell. Forewing with vein 6 bent 
towards base; 10 from the cell. Male genitalia with transtilla 
complete; uncus bilobed.] 

1. GenuB Cryptoblabes Zeller 

Cryptoblabes ZeUer, Isis von Oken, 1848, p. 644. — Ragonot, 
Monograph, pt. 1, pp. xliv, 12, 1893. — Staudinger and Rebel, 
Catalog der Lepidopteren des palaearctischen Faunenge- 
bietes, vol. 2, p. 42, 1901. — Mayrick, Revised handbook 
of British Lepidoptera, p. 397, 1928. — Bisset, in Pierce and 
Metcalfe, Genitalia of the British Pyrales, p. 57, 1938. — 
Janse, Journ. Ent. Soc. South Africa, vol. 14, p. 143, 1941. 
(Type of genus: Cryptoblabes rutilella Zeller, a synonym of 
Ustriga (Haworth); figs. 2, 131, 638.) 

Tongue well developed. Antenna pubescent; shaft 
of male notched at base and with curved, homy hook 
protruding from the notch. Labial palpus upturned, 
slender, reaching a little above vertex; third segment 
about two-thirds the length of second, acuminate. 
Maxillary palpus squamous. Forewing smooth; 11 
veins; vein 2 from well before the lower outer angle of 
cell (from lower median vein of cell at about three- 
fourths); vein 3 also before the angle (from lower 
median at about five-sixths) ; 4 and 5 closely approxi- 
mate at base, rarely (in individual specimens) connate; 
6 bent towards base, from upper angle or from very 
close to upper angle of cell; 10 from the cell, separated 
from stalk of 8-9 at base; male without costal fold. 
Hind wing with vein 2 from middle or just beyond mid- 
dle of lower median vein of cell; 3 from before and 
more or less removed from the outer angle of cell; 4 
and 5 from the angle, closely approximate at base, 
thence diverging, 7 and 8 closely approximate, con- 
tiguous or weakly anastomosing for a short distance 
beyond cell; cell less than one-half the length of wing, 
but not "very short" as stated by Kagonot in his 
generic key (Monograph, p. xUv); discoceUular vein 
curved. Eighth abdominal segment of male simple. 

Male genitalia with apical process of gnathos a small, 
simple hook. Transtilla complete; developed as a 
narrowly banded bridge with more or less elongate 
central projection. Uncus broad, with apical margin 
broadly rounded and invaginate, giving the uncus a 
bilobed appearance. Harpe (in European species) with 
strong hair tufts arising from articulated plates in 
intersegmental area between base of sacculus and ter- 
minal margin of eighth abdominal segment, or with 
long hair tuft from sacculus near its base (rutilella). 
AneUus with elongate, narrow, lateral lobes. Aedeagus 
simple; penis with or without cornutus, latter, when 
present, a single, long, spine. Vinculum broad and 
with broad terminal margin more or less concave. 

Female genitalia with ductus bursae membranous, 
finely scobinate towards junction with bursa; genital 
opening simple except for a narrow, sclerotized band 
behind and above the opening; bursa membranous 
more or less finely scobinate; signum present, developed 
as a stout, blunt, flattened thorn (rutilella) or a patch 



of dense granulations (gnidielia) ; ductus seminahs 
from bursa. 

The foregoing description was drawn from European 
species which are obviously congeneric. Numerous 
other species have been described in the genus from 
India, Formosa, Japan, Australia and the islands of 
the Pacific. Whether these are aU congeneric 1 do not 
know. Two unidentified species before me from the 
Philippines have genitaha similar in habitus to those 
of the genotype (rutilella) except that the dorsal, inter- 
segmental tufts at base of harpe are lacking. Their 
unci have the same characteristic bilobed appearance. 
The venational character which has been generally 
accepted as defining the genus (i. e., the position of vein 
3 of hind wing in relation to the lower outer angle of 
cell) is variable and illusive, being closer to the angle 
in gnidielia (fig. 1) than in rutilella (fig. 2) and still 
closer in the Philippine species. Indeed, in American 
examples of Acrobasis (=Mineola) vein 3 is often as far 
from the angle as it is in gnidielia. The shorter cell of 
Cryptoblabes, coupled with the position of vein 3, wiU, 
however, suffice to maintain the group separation made 
between the two genera in our key. 

Cryptoblabes is a distinctly Old World genus with no 
indigenous New World species. It is represented in the 
Western Hemisphere by only one introduced European 
species (gnidielia). 

1. Cryptoblabes gnidielia (Millifere) 

FiGUEES 1, 132, 639 

Ephestia gnidielia Millihre, Iconographie et description de 

chenilles et L^pidoptferes inedits, vol. 2, p. 308, 1864. 
Cryptoblabes gniediella (Millifere) Ragonot, Monograph, pt. 1, 
p. 16, 1893. — Dyar, Ins. Insc. Menstr., vol. 3, p. 88, 1915. 

Forewing pale brownish fuscous with a faint rosy 
overcast due to more or less diffused longitudinal streaks 
of reddish scales (in fresh and weU marked specimens 
especially along the fold, on the veins in outer area, and 
below costa from base); some whitish dusting along 
costa and in the cell, most pronounced as a pale shade 
between the dark discal spots at end of cell; transverse 
lines obscure and not sharply outlined, but distinguish- 
able, whitish ocherous; the antemedial Une obhque and 
curved, set weU out towards middle of wing; subter- 
minal nearly straight, parallel with termen; discal dots 
separate, blackish fuscous. Hind wing whitish, trans- 
lucent, the veins darkened, a narrow dark shade along 
costa and a narrower dark line along terminal margin. 
Alar expanse, 11-16 mm. 

Male genitalia with heavy hair tufts from plates 
articulating with base of sacculus of harpe; penis armed 
with spinelike cornutus about two-thirds as long as 
aedeagus. Female genitalia with signum developed as 
a dense scobinate-granulate patch; ductus seminalis 
from bursa near attachment of ductus bursae. 

Type localitt: France (type in Paris Mus.) . 

Food plants: Fruits of Chaenomeles japonica, pome- 
granates, oranges, citron, grapes, raisins, etc. (often 
fallen and desiccated fruit), onion seeds, leaves and 
flowers of Daphne gnidium, flowers of Ricinus communis, 
green corn stalks (reared moth, in USNM, from Hawaii), 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



11 



young stems of Tamarix and the stems, leaves, and 
flowers of Ly thrum. Apparently has a various larval 
habit and a wide variety of hosts. A moth (in USNM) 
from Muar Johore, Malaya (Clausen), was reported as 
reared from a larval predator on Aleurocanthus. 

Distribution: Mediterranean countries of Europe, 
Africa, and Asia. Presumably widely distributed in 
the east and among the Pacific islands; but some of the 
published records may apply to other species. New 
World distribution: Bermuda (Jan., Feb., Apr., May). 
Venezuela: El Valle (Aug.). Brazil: Sao Paulo 
(Feb.). 

Genera 2-17: Acrohasis to Chararica 

[Venational division B. Hind wing with vein 3 from the lower 
outer angle of the cell or (if from before the angle) close to it; 
7 and 8 approximate beyond cell, rarely shortly and weakly 
anastomosed. Male genitalia with transtilla complete or, 
where incomplete (Bertelia, Hypargyria) , the elements strongly 
developed and with expanded apices. Uncus triangulate, or 
hoodlike with rounded apical margin, or spatulate {Birinus).] 

2. Genus Acrohasis Zeller 

Acrobasis Zeller, Isis von Oken, 1839, p. 176; 1848, p. 606. — 
Herrich-Schaffer, Systematische Bearbeitung der Schmet- 
terlinge von Europa, vol. 4, p. 99, 1849. — Heinemann, Die 
Schmetterlinge Deutschlands und der Schweiz, vol. 1, 
pt. 2, p. 175, 1865.— Hulst, Phycitidae of N. Amer., p. 120, 
1890; U. S. Nat. Mus. Bull. 52, p. 418, 1902.— Ragonot, 
Monograph, pt. 1, p. 85, 1893. — Dyar, Proc. Ent. Soc. 
Washington, vol. 10, p. 41, 1908. — Spuler, Die Schmetter- 
linge Europas, vol. 2, p. 214, 1910. — Barnes and McDun- 
nough, Contributions, vol. 2, p. 221, 1914. — Forbes, Cornell 
Mem. 68, p. 614, 1923. — Meyrick, Revised handbook of 
British Lepidoptera, p. 397, 1928. — Pierce and Metcalfe, 
Genitalia of the British Pyrales, p. 10, 1938. — Bisset, in 
Pierce and Metcalfe, op. cit., p. 55, 1938. — Janse, Journ. 
Ent. Soc. South Africa, vol. 4, p. 148, 1941. (Type of 
genus; Phycis tumidella Zincken (= Acrohasis zelleri 
Ragonot) ; figs. 133, 640.) 

Mineola Hulst, Phycitidae of N. Amer., p. 126, 1890; U. S. Nat. 
Mus. Bull. 52, p. 419, 1902.— Forbes, Cornell Mem. 68, 
p. 618, 1923. (Type of genus: Myelois indigenella Zeller.) 

Seneca Hulst, Phycitidae of N. Amer., p. 177, 1890. (Type of 
genus: Cateremna tumidulella Ragonot. New synonymy.) 

Acrocaula Hulst, Canadian Ent., vol. 32, p. 170, 1900. (Type 
of genus: Acrocaula comacornella Hulst. New synonymy.) 

Tongue weU developed. Antenna pubescent; on 
male, basal segment enlarged and angulate, the shaft 
with a slight sinus at base. Labial palpus upturned, 
reaching to or a trifle above vertex; third segment 
slightly more than half the length of second, acuminate. 
Maxillary palpus filiform. Forewing smooth or with 
transverse antemedian ridge of raised scales; 11 veins; 
vein 2 from M^ell before the angle of the cell; 3 rather 
well separated from 4, but somewhat nearer to 4 than 
to 2; 4 and 5 closely approximate at base or connate 
(rarely, in individual specimens, shortly stalked); 
6 from below upper angle of cell, straight; 10 from the 
cell, usually (except in individual specimens) separated 
from stalk of 8-9 at base; male without costal fold. 
Hind wing with vein 2 from well before lower outer 
angle of cell (from outer two-thirds of lower median) ; 



3 from before but near the angle; 4 and 5 from the 
angle, connate; 7 and 8 shortly anastomosed beyond 
ceU; cell about half the wing length; discocellular vein 
curved. Eighth abdominal segment of male with 
midventral hair tuft. 

Male genitaUa with apical process of gnathos a simple, 
elongate hook, or an elongate trifurcate hook. Trans- 
tilla complete, sharply angulate and reaching as far 
back as base of apical process of gnathos; terminal 
margin narrow and indented. Uncus broadly triangu- 
late. Harpe simple. Anellus a narrowly sclerotized 
U- or V-shaped plate. Aedeagus simple, rather stout; 
penis with numerous sclerotized wrinklings, otherwise 
unarmed. Vinculum stout, a trifle longer than broad, 
slightly tapering; terminal margin truncate and more 
or less concave. 

Female genitalia with ductus bursae and biu-sa popu- 
latrix membranous except for a dorsal sclerotized plate 
in genital opening; ductus and bursa more or less scobi- 
nate; signum, if present, consisting of a granulate cup 
or a minute central spine surrounded by a dense cluster 
of scobinations ; ductus seminalis from a lobe of bursa 
near junction of bursa and ductus bursae. 

Acrobasis as here defined is something of a composite 
genus, dividing into two distinct groups on the develop- 
ment of the apical process of gnathos. Typical Acro- 
basis, comprising the European species with both 
smooth-winged forms (including the type, tumidella) 
and those with the raised-scale ridge on forewing, and 
all smooth-winged American species (formerly under 
Mineola), have the apical process of gnathos produced 
as a simple, elongate hook (fig. 133b). All our Ameri- 
can species with the raised-scale ridge on forewing 
(except minimella Ragonot) have the apical process of 
gnathos trifurcate, that is, produced as an elongate 
hook with a lateral projection from each side before 
apex. This latter group is strictly North American, 
limited in distribution to the United States, and Canada 
east of the Rocky Mountains (except for A. betulella 
Hulst) . As far as I know there are no Old World species 
with a similar gnathos. I had hoped to distinguish 
this distinctly American group as a separate genus under 
one of Hulst's available names (Seneca or Acrocaula) on 
the basis of the trifurcate projection of gnathos and the 
raised-scale ridge on the forewing; but minimella Rago- 
not prevents this, as it falls between the two groups, 
having the raised-scale ridge and the simple projection 
of gnathos. The females offer no characters that will 
serve to differentiate the groups. Their genitaha are 
so similar that they cannot be used, in many cases, 
even for specific separation, much less for group division. 

The males of the typical American group with raised- 
scale ridge are somewhat more variable than the females, 
exhibiting slight differences in the shape of the transtilla 
and apical projection of gnathos. Figures of these 
structures are given, for what they are worth, for all the 
species represented by authentic males. The differ- 
ences are comparative only, and I suspect that, when 
extended series of the several species are available, they 



12 



TTNITED STATES NATIONAL MUSEUM BULLETIN 207 



will prove to be more individual than specific in 
character. 

The reasons for sinking Seneca and Acrocaula into the 
synonjTtny of Acrobasis are given in the discussion of 
their types (pp. 22 and 24). 



Genus Acrohasis, Species 2-7: A. indigenella to 
A. comptella 

[Male with apical process of gnathos a simple hook; forewing 
smooth.] 

2. Acrobasis indigenella (Zeller) 
FiGUBES 3, 134, 641 

lots indigenella Zeller, Isis von Oken, 1848, p. 867. — Riley, 

Fourth annual report on the noxious, beneficial and other 

insects, of the State of Missouri, p. 38, 1872. 
Phycita nebulo Walsh, Prairie Farmer, p. 308, 1860; Proc. 

Boston Soc. Nat. Hist., vol. 9, p. 312, 1863. 
Phycis indiginella (Zeller) Weed, in Forbes, Fifteenth report of 

the State Entomologist on the noxious and beneficial insects 

of the state of Illinois, p. 65, 1889. 
Acrobasis indiginella (Zeller) Riley, Canadian Ent., vol. 16, 

p. 237, 1884.— Ragonot, Monograph, pt. 2, p. 118, 1893.— 

Forbes, Cornell Mem. 68, p. 619, 1923. 
Phycita (Acrobasis) nebulo (Walsh) Riley, Fourth annual report 

on the noxious, beneficial and other insects, of the State of 

Missouri, p. 38, 1872. 
Phycita (Acrobasis) nebulo nebulella Riley, Fourth annual report 

on the noxious, beneficial and other insects, of the State of 

Missouri, p. 42, 1872. 
Myelois zelatella Hulst, Ent. Amer., vol. 3, p. 136, 1887. 
Mineola indiginella (Zeller) Hulst, Phycitidae of N. Amer., p. 130, 

1890. — Quaintance and Siegler, U. S. Dep. Agr. Farmers' 

Bull. 1270, p. 49, 1922.— Essig and Keifer, Monthly Bull. 

California Dep. Agr., vol. 22, p. 155, 1933. — McDonough, 

Check list. No. 6115, 1939. 
Mineola indigenella nebulella (Riley) Hulst, Phycitidae of 

N. Amer., p. 131, 1890. 

Forewing gray-brown densely dusted with white ; the 
white dusting concentrated on the upper half of wing 
and somewhat between subterminal line and outer 
margin, forming two strongly contrasted white patches, 
one from costa in subbasal area, the other more or less 
triangular and extending from costa, between the 
transverse dark lines, into cell and including the blackish 
discal spots; the whitish terminal area is less sharply 
defined and on many specimens somewhat faint; trans- 
verse antemedial line curving obliquely from basal third 
of costa to middle of inner margin, indicated chiefly by 
its outer dark margin, which begins as a conspicuous 
black triangle on costa ; from inner margin at one-third 
a blackish line curves upward to meet the antemedial 
line near costa ; the area enclosed between them reddish 
ocherous; a similar, smaller spot of the same color on 
base of inner margin; extreme base of costa blackish; 
subterminal line sinuate, bordered inwardly and out- 
wardly by dark lines which begin as blackish spots on 
costa; from the outer of these a dark band extends 
transversely across to the base of antemedial line at 
inner margin, somewhat obscured in the dark ground 
color on all but the palest and most contrastingly 
marked specimens; discal spots at end of cell black, 



more or less confluent, usually a black bar along dis- 
cocellular vein; a more or less broken, black line along 
terminal margin. Hind wing subpeUucid, pale smoky 
fuscous; the veins very faintly, if at aU, darkened; 
a narrow, obscure, dark line along termen. Alar ex- 
panse, 15-20 mm. Male genitalia as given for the genus. 
Female genitalia without signum. 

Type localities: "Carolina" (indigenella, in BM); 
Illinois? (nebulo, lost) ; Missouri? (nebulella, lost); "New 
York and Canada," (zelatella; the supposed type, cf , 
in AMNH, ex Kutgers, is labeled "Blanco County, 
Texas"). 

Food plants: Apple, crabapple, plum, prune, cherry, 
quince, Crataegus, Cotoneaster, Pyrocantha coccinea. 
Larva feeding on leaves and forming serpentine resting 
and hibernating case of silk and frass. 

Disteieution: United States: Maine, Sebec Lake 
(July); New Hampshire, Hampton (July); Vermont, 
Clenendon; Massachusetts, Newton Highlands; Connec- 
ticut, East River (July) ; New York, Catskill Mts., Hion; 
New Jersey, Rutherford; Pennsylvania, Chambersburg 
(June, July), Germantown (July), New Brighton (July), 
Pittsburgh (June); Virginia, Colonial Beach (July), 
Norfolk (M&j); North Carolina, Tryon (Aug.); Georgia, 
Savannah (Apr.); Illinois, Chicago, Decatur (May, 
June, July, Aug.); Missouri, Mossele (June, July), 
Norborne (Apr., June, July), St. Louis (Aug.) ; Nebraska, 
Wahoo (May) ; Kansas, Onaga, Wichita (June) ; Missis- 
sippi, "Agr. CoUege" (May, June, July); Arkansas, 
Siloam Springs (June), Washington County (July); ^ 
Texas, Abilene, Blanco County (Sept.), Fort Worth 
(Sept.), Houston (May, June, Aug.), Kerrville, Victoria 
(June); California, Lomita, Los Angeles County (Mar.) 
Orange County (June). (The California records all 
from plum.) Canada: Ontario, Trenton (June, July); 
Quebec, Meach Lake (July). 

Apparently generally distributed east of the Rocky 
Mountains and rather recently introduced into Cali- 
fornia. 

This species has been considered of economic impor- 
tance as a defoliator of fruit trees in the Middle States, 
and in the official list of common names approved by 
the American Association of Economic Entomologists 
is designated as the "leaf crumpler." However, it does 
not seem to be more than a minor pest of local and 
occasional concern. Several references are made to it 
in the economic literature but none of these adds any- 
thing of significance to our knowledge of the insect 
beyond what is given in the early papers by Walsh and 
Riley. 

Riley's nebulella was described by him as a variety 
of nebulo and distinguished from the latter by the more 
diffused dark shading and the separation of the discal 
spots on forewing. In the series before me there is 
considerable variation in the extent and intensity of the 
dark coloring and the discal mark varies even more, 
being sometimes divided into two distinct spots or fused 
into a single bar on different sides of the same specimen, 
so the varietal designation is hardly worth maintaining. 
In 1908 (Proc. Ent. Soc. Washington, vol. 10, p. 45) 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



13 



Dyar, on the evidence of a supposed t5rpe of nebulella 
in the National Collection, resurrected the name and 
applied it to the "pecan leaf casebearer." Barnes and 
McDunnough (Contributions, vol. 2, p. 222, 1914; 
vol. 3, p. 221, 1917) called attention to the spuriousness 
of the alleged "type" and gave the pecan casebearer its 
proper reference (juglandis LeBaron); but on the 
strength of Dyar's identification the name nebulella 
had already appeared, and continued to be used for some 
years in economic publications for the pecan leaf case- 
bearer. In his 1939 Check List McDunnough applies 
the name in an entirely new sense, transferring it to 
Meroptera with the well-known unicolorella Hulst as a 
synonym. This was most unfortunate and altogether 
unnecessary. We know what unicolorella Hulst is, and 
its type is at hand for reference. The type of nebulella 
is nonexistent and McDunnough's new reference has 
nothing to back it but an entomologist's interpretation 
of Riley's description and very poor and over-inked 
figure of the forewing. I see nothing in either to rule 
out the original interpretation, so shall let the name 
sleep in synonymy. 

3. Acrobasis grossbecki (Barnes and McDunnough), new combi- 
nation 

Mineola indigenella nebulella Grossbeck (not Riley), Bull. Amer. 
Mus. Nat. Hist., vol. 37, p. 129, 1917.— Barnes and Mc- 
Dunnough, Contributions, vol. 3, p. 220, 1917. 

Mineola grossbecki Barnes and McDunnough, Contributions, 
vol. 3, p. 221, 1917.— McDunnough, Check list. No. 6116, 
1939. 

Forewing purplish brown, the dark ground color 
more extended than in indigenella, obscuring the pale 
antemedial line and completely obliterating the reddish 
ocherous subbasal patch on inner margin usually present 
in the genus; triangular black spot on costa, beginning 
the outer dark border of antemedial line, distinct and 
sharply contrasted as in indigenella; white areas 
restricted more than in indigenella, the subbasal one 
narrowly triangulate with its point on inner margin, 
midcostal one extending to and including the discal 
spots in its lower angle; whitish dusting in terminal 
area very faint; sub terminal line obscure; discal spots 
at end of cell black, separated. Hind wing shiny, 
smoky fuscous. Alar expanse, 15-16 mm. 

Genitalia as in indigenella. 

Type locality: Lakeland, Fla. (type in USNM). 

Food plant: Crataegus (larva feeding on the leaves). 

Distribution: Known only from the type locality. 
May be a Florida race of indigenella, but appears to 
be a distinct species despite the likeness of its genitalia 
to ithose of indigenella. 

4. Acrobasis vaccinii Riley 
FiQ0BE 642 

Acrobasis vaccinii Riley, Canadian Ent., vol. 16, p. 237, 1884; in 
Rep. [U. S.] Comm. Agr. for 1884, p. 355, 1885.— Smith, 
in Rep. [U. S.] Comm. Agr. for 1884, p. 394, 1885. — Saunders, 
Insects injurious to fruits, p. 375, 1883 (as "The Cranberry 
Fruit-worm"). — Ragonot, Monograph, pt. 1, p. 121, 1893. — 
Forbes, Cornell Mem. 68, p. 618, 1923. 



Mineola vaccinii (Riley) Hulst, Phycitidae of N. Amer., p. 128, 
1890.— Brown, Oregon Agr. Exp. Station Bull. 225, p. 19, 
1927.— Crowley, Washington Agr. Exp. Station Bull. 230, 
p. 24, 1929.— McDunnough, Check list. No. 6114, 1939.— 
Beckwith, Journ. Econ. Ent., vol. 34, p. 169, 1941. 

Averaging smaller than indigenella; dark ground 
color similar but more extended and without the con- 
trasting black costal triangle; pale antemedial line 
obliterated by a transverse extension of the ground 
color, bordered inwardly by an almost vertical, rather 
narrow white band which expands narrowly on costa 
towards base (the remains of the much-reduced sub- 
basal white area); midcostal white patch also much 
restricted, barely including at its lower angle the 
separated black discal spots; on fresh specimens some 
sprinkling of rufous scaling is distinguished under high 
magnification, but no reddish or other contrastingly 
colored, angulate, subbasal patch on inner margin (as 
in indigenella and tricolorella) . Hind wing pale smoky 
fuscous. Alar expanse 14-18 mm. 

Male genitalia differing in no significant detail from 
those of indigenella. Female genitalia with bursa 
more or less heart-shaped (less elongate than that of 
indigenella or grossbecki); signum present as a minute 
granulate cuplike patch. 

Type locality: Massachusetts (type in USNM). 

Food plant: Cranberry, blueberry (larva in the 
fruit). 

Distribution: Massachusetts (type series, no exact 
locahty, June), Wareham (June, July); Connecticut,Ea,st 
River (July);iVew Jersey, Pemberton (May), Whitesbog 
(June); Wisconsin; Michigan; Georgia; Mississippi, 
Biloxi, Poplarville; Washington, Long Beach (June), 
Sea view (July). 

Presumably generally distributed on the range of its 
food plants in the United States and Canada. The 
foregoing records are from reared and typical examples 
in the National Collection. 

This species, popularly known as the "cranberry 
fruitworm," is of some importance, especially to cran- 
berry growers, and has a rather extensive economic 
literature, mostly in annual reports, bulletins, and other 
pubHcations of state entomologists and experiment 
stations. None of these adds anything of biological or 
taxonomic significance to the earlier records of Riley 
and Smith. 

5. Acrobasis amplexella Ragonot 

Acrobasis amplexella Ragonot, N. Amer. Phycitidae, p. 3, 1887; 

Monograph, pt. 1, p. 97, 1893.— Forbes, Cornell, Mem. 68, 

p. 618, 1923. 
Mineola amplexella (Ragonot) Hulst, Phycitidae of N. Amer., 

p. 127, 1890.— McDunnough, Check list. No. 6112, 1939. 

This is probably nothing but a color form of vaccinii. 
I can find no difference from the latter except in the 
greater extension of the basal dark area of forewing and 
the consequent further restriction of the subbasal white 
area which is a narrow band throughout, not expanding 
along costa towards base. 

Rearing will have to settle the status of amplexella. 
In the material before me there are only collected 



14 



tnsriTED STATES NATIONAL MUSEUM BULLETIN 207 



specimens. All reared examples we have from either 
cranberry or blueberry are typical vaccinii. 

Alar expanse, 12-18 mm. 

Type locality: North Carolina (type in Paris 
Mus.). 

Food plant: Unknown. 

Distribution: Maine, Monmouth (June), Sebec 
Lake (July); New Hampshire (June, July); Massachu- 
setts, Cohasset (July), Framingham (Jime), Winchendon 
(July), Worcester (July) ; Connecticut, East River (July) ; 
New York, Liberty (June), Sullivan County (July); 
Pennsylvania, Hazleton (May); North Carolina, Tryon 

(July). 

In the Barnes and old U. S. National Museum Col- 
lections these examples were about equally divided 
under the two names, amplexella and vaccinii. 

6. Acrobasis tricolorella Grote 
Figures 135, 643 

Acrobasis tricolorella Grote, Bull. U. S. Geol. Geogr. Surv. Terr., 
vol. 4, p. 694, 1878. — Ragonot, Monograph, pt. 1, p. 93, 
1893.— Forbes, Cornell Mem. 68, p. 618, 1923. 

Mineola tricolorella (Grote) Hulst, Phycitidae of N. Amer., p. 
127, 1890.— McDunnough, Check list, No. 6111, 1939. 

Mineola scitulella Hulst, Canadian Ent., vol. 32, p. 169, 1900. — 
Hungerford, Idaho Agr. Exp. Station Bull. 149, p. 29, 
1927; Bull. 164, p. 29, 1929.— Pack and Dowdle, Journ. 
Econ. Ent., vol. 23, p. 321, 1930. — Haegele, Journ. Econ. 
Ent., vol. 25, p. 1073, 1932.— Essig and Kiefer, Monthly 
Bull. California Dep. Agr., vol. 22, p. 153, 1933.— McDun- 
nough, Check list, No. 6110, 1939. (New synonymy.) 

Forewing grayish fuscous more or less dusted with 
whitish scales, in some specimens well diffused over the 
middle of wing, giving it a pale slate ground color, but 
normally concentrated into a pale patch from costa 
before subterminal line, including the discal mark, and 
a pale terminal suffusion below apex; usual subbasal 
white area constricted into a narrow, sharply defined 
antemedial band, outwardly oblique from costa to top 
of cell, thence vertical to iimer margin and bordered 
outwardly on its vertical portion by a more or less 
triangular, contrasted orange or reddish orange patch; 
bordering the white line and the orange patch out- 
wardly, a black oblique angulate line extending to 
near middle of inner margin and beginning on costa in 
a more or less angulate and diffused blackish patch 
(similar to but not so sharply defined nor contrasted as 
the black costal patch on indigenella) ; subterminal line 
distinct, narrow, white, angled inwardly at vein 6 and 
lower fold and curved outwardly between, bordered 
inwardly by a narrow black line and outwardly by a 
black costal spot and a more or less pronounced orange 
or reddish orange band (well marked in many eastern 
and western specimens, but sometimes obscured by 
dark scaling) ; discal spots usually fused into a slightly 
curved, black bar along discocellular vein, rarely 
separated. Hind wing smoky white to pale smoky 
fuscous. Alar expanse, 18-22 mm. 

Male genitalia with cucullus of harpe of more even 
width throughout and apex more evenly rounded than 
in other species of the genus. Eighth abdominal seg- 



ment of male with a single central, ventral hair tuft 
(supplementary ventral hair tufts on the other species). 
Female genitalia with signum present as a small, 
granulate, cup-shaped patch. 

Type localities: Oldtown, Maine (tricolorella, in 
BM); Colorado (scitulella, in AMNH, ex Rutgers). 

Food Plants: Apple, apricot, catalina cherry, plum, 
prune (larvae feeding in buds and fruits, probably also 
on leaves), also recorded from galls on chokecherry 
(Park and Dowdle). 

Distribution: United States: Maine, Oldtown, 
Bar Harbor (July), Orono; New Hampshire, Hampton 
(July) ; Massachusetts, Amherst (Hatch Exper. Station, 
July); New York, Catskill Mts., Illion (July); New 
Mexico, Albuquerque (July); Colorado, Denver, Glen- 
wood Springs (May, July, Aug., Sept.), Grand Junction 
(Aug.), Gunnison County (July); Utah, BeUevue (May), 
Dividend (Aug.), Eureka (Aug.), Logan (July), Park 
City (July), Provo (July); Idaho, Boise (Sept.), Emmett 
(July); California, Loma Linda (Aug.), Mount Lowe 
(May, July), San Diego (June, July), Santa Barbara 
(Aug.), Warner Mts. (Modoc County, July); Oregon 
Lake View (Aug), The Dalles (June); Washington, 
Prosser (Jtme), PuUman (July), Walla WaUa (Aug.), 
Wenatchee (Aug.). Canada: Ontario, Ottawa (July); 
Manitoba, Cartwright; British Columbia, Arrowhead 
Lake (June). 

The species seems to be abundant in our western 
states and relatively scarce in the east, to judge by 
examples in collections, and has attracted some atten- 
tion as a fruit pest in Utah, Idaho, and California. 
There is nothing to distinguish western from eastern 
specimens and the one detail that Hulst relied upon 
for the separation of his scitulella (the presence of 
an orange outer border to the subterminal line) does not 
hold. It is present in eastern and western examples 
and equally variable in both. I am therefore sinking 
the name in the synon3Tny of tricolorella. 

7. Acrobasis comptella Ragonot 

Figure 646 

Acrobasis comptella Ragonot, N. Amer. Phycitidae, p. 4, 1887. — 
Hulst, Ent. Amer., vol. 5, p. 156, 1889 (makes synonym of 
caliginella Hulst). 

Forewing dark gray dusted with white, the white 
dusting concentrated on basal area, on costal median 
half of wing (forming a pale angulate patch which 
includes the discal spots), and in terminal area beyond 
subterminal line; in some specimens the white dusting 
is more extended, making most of the basal, median, 
and terminal areas pale ashy gray; outwardly bordering 
basal pale area a black line (narrowing from a shallow 
triangulate patch on costa) extends obliquely outward 
to top of cell thence vertically to iimer margin, bordered 
outwardly on vertical part by a triangulate, tawny or 
reddish brown patch; subterminal line white, sinuate, 
bordered inwardly by a narrow, blackish line and out- 
wardly, at costa, by a black smudge; discal spots at end 
of cell distinct, separate, black. Hind wing pellucid. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITDSrAE 



15 



whitish or pale smoky fuscous. Alar expanse, 14-21 
mm. 

Male genitalia exhibiting no distinctive specific 
characters. Female genitalia with several short, paral- 
lel lines of fine scobinations in bui-sa ; signum present as 
a small, granulate, cup-shaped patch. 

Type locality: California (type in Paris Mus.). 

Food plant: Unknown. 

Distribution: California, Cisco (July), Colfax (July), 
Pasadena, San Diego (June), Santa Catalina Isl. (May), 
Warners (San Diego County, Aug.); Arizona, Gila 
County, Eedington; New Mexico, Albuquerque (July), 
Las Vegas; Utah, Provo (July). 

Hulst in 1889 made comptella a synonym of his 
caliginella and it has remained as such in our lists. 
However, the two are genericaUy as well as specifically 
distinct, caliginella having the basal segment of the 
male antenna cylindrical (not triangularly expanded at 
apex as in Acrobasis). It is superficially similar in 
color and markings to comptella; but the black line 
bordering the whitish basal patch of forewing is 
distinctly broken, its vertical portion not reaching to 
inner margin. I am removing both caliginella and 
Mineola supposita Heinrich to Rhodophaea. 

Genus Acrobasis: Species 8. A. minimella 

[Male with apical process of gnathos a simple hook; forewing 
with raised-scale ridge.] 

8. Acrobasis minimella Ragonot 
Figure 140 

Acrobasis minimella Ragonot, Ent. Amer., vol. 5, p. 113, 1889; 

Monograph, pt. 1, p. 105, 1893. — McDunnough, Check list, 

No. 6088, 1939. 
Acrobasis nigrosignella Hulst, Phycitidae of N. Amer., p. 123, 

1890.— Dyar, Proc. Ent. Soc. Washington, vol. 10, p. 43, 

1908. 

Forewing grayish fuscous with a purplish sirffusion! 
some whitish dusting on basal area and, very faintly, 
from costa before sub terminal line to discal spots; 
antemedial line obscm-e except towards inner margin, 
where it is a narrow whitish line; a blackish triangular 
costal patch following the antemedial line and continued 
as thin black line on its outer border to inner margin; 
vertical scale ridge black; area between scale ridge and 
antemedial line ocherous or reddish; discal spots at 
end of cell small, obscure, separated. Hind wing smoky 
fuscous. Alar expanse, 13-16 mm. 

Female genitalia exhibiting no specific difference to 
distinguish them from those of other species having the 
raised-scale ridge on forewing. 

Type locality: Texas (minimella, 9, in Paris Mus., 
and nigrosignella, in AMNH, ex Rutgers) . 

Food plant: Oak (this food plant record from speci- 
men, in USNM, reared at Falls Church, Va., under 
Hopkins No. 9847, C. F. Johansen). 

Distribution: Texas (Apr.) ; Mississippi, Starkville 
(July); Louisiana, Winfield (June); North Carolina, 
Southern Pines (Apr., May, June, Aug.), Tryon (May); 



Virginia, Falls Church ; District oj Columbia, Washing- 
ton (June) ; New Jersey, Lakehurst (July) . 

The species is easily recognized by its size, color, and 
male characters. The sex-scaling is present and con- 
sists of long broad black costal streaks on underside of 
fore and hind wings. 

Genus Acrobasis, Species 9-22: A.feltella to A. 
demotella 

(Male: Apical process of gnathos trifurcate; forewing with raised- 
scale ridge; black sex-scaling beneath.] 

9. Acrobasis feltella Dyar 

Figure 141 

Acrobasis feltella Dyar, Proc. Ent. Soc. Washington, vol. 11, 
p. 214, 1910.— Ely, Ins. Inso. Menstr., vol. 1, p. 51, 1913.— 
McDunnough, Check list, No. 6080, 1939. 

Head, basal segment of antenna, thorax, and basal 
area of forewing white ; a faint rosy tint on the posterior 
of thorax and a more obvious rosy shading on lower 
half of pale basal area of forewing (more intense and 
extended on the female than on the male) ; remainder 
of wing dark gray-brown (in fresh specimens blackish 
brown) with a faint, pale grayish shading in terminal 
area and a whitish spot on inner margin near tornus 
(the lower end, and contrasted portion of the otherwise 
obscure subterminal line) ; black discal dots at end of 
cell distinguishable but somewhat obscured in the dark 
ground color, usually separate, but occasionally fused; 
antemedial scale ridge blackish. Hind wing of male 
white at base, shading to smoky fuscous outwardly; 
the veins in both sexes faintly darkened. Alar expanse, 
14-18 mm. 

Black sex-scaling consisting of a short patch at base 
of costa on forewing. 

Type locality: Warner, N. Y. (type in USNM). 

Food plant: Hickory (larva boring in petioles). 

Distribution: United States: New York, Warner 
(July) ; Connecticut, East River (Jidy) ; Illinois, Putnam 
County (June). Canada: Ontario, Merivale (June). 

This species, palliolella Ragonot, and caryalbella Ely 
are identical in color, maculation, and all superficial 
characters. They exhibit trifling differences in their 
male genitalia, especially in the shapes of their trans- 
tillae and the apical processes of their gnathi. These 
differences are probably no more than individual in 
character. We figure them for what they are worth. 
Ely (1913) noted differences in the larval cases of 
feltella and caryalbella which should be significant. He 
also saw, or thought he saw, a difference in the sex- 
scaling of Dyar's type and the type of caryalbella. In 
this he was in error; for the sex-scaUng is identical in 
both types and on the males of palliolella and juglandis 
as well. I suspect that the three names (feltella, 
palliolella, and caryalbella) apply to a single species; but 
this cannot be determined until the biologies and larvae 
of the various hickory-feeding forms of the genus are 
more thoroughly studied. Until that is done it seems 
best to keep the names separated. 



16 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



10. Acrobasis palliolella Ragonot 
Figure 142 

Acrobasis palliolella Ragonot, N. Amer. Phycitidae, p. 4, 1887; 

Mcnograph pt. 1, p. 92, 1893. — Hulst, Phycitidae of N. 

Amer., p. 121, 1890. 
Acrobasis albocapitella Hulst, Ent. Amer., vol. 4, p. 116, 1888. 

Not distinguishable superficially from feltella. Tri- 
fling differences in the male genitalia are shown in the 
figure. They are probably not significant. 

Type localities: North America (palliolella, in 
Paris Mus.); Canada (albocapitella, in AMNH, ex 
Rutgers). 

Food plant: Presumably hickory. Life history not 
known. 

Distribution: United States: Illinois, Chicago 
(July); Pennsylvania, New Brighton (July); North 
Carolina, Plymouth (May); Connecticut, East River 
(July). Canada: Ontario, Ottawa (July). 

The name palliolella has been variously misapplied 
and has appeared frequently in economic literature for 
the "pecan leaf casebearer" (juglandis LeBaron). In 
our latest checklist (McDunnough, 1939) it appears as 
a synonym oi juglandis but I do not think this is correct. 

A long series of juglandis before me shows consider- 
able variation in color but at the same time consistent 
differences from palliolella, whose closest aflinities are 
feltella Dyar and caryalbella Ely. 

11. Acrobasis caryalbella Ely 
Figure 143 

Acrobasis caryalbella Ely, Ins. Insc. Menstr., vol. 1, p. 52, 
1913.— McDunnough, Check list. No. 6081, 1939. 

Acrobasis angusella Dyar (not Grote), Proc. Ent. Soc. Washing- 
ton, vol. 10, p. 42, 1908. 

Ely distinguishes his species from Dyar's feltella 
chiefly on the differences in their larval cases ("co- 
coons"). There is nothing else to separate them except 
some slight and probably not significant differences in 
their genitalia. These are shown in the figure. 

Type locality: East River, Conn, (type in USNM). 

Food plant: Hickory. 

Known only from reared examples from the type 
locality and the female from hickory (June) without 
locality label, bearing Riley's No. 376 and referred by 
Dyar to angusella Grote. The sex-scaling on the male 
is the same as that on feltella and palliolella. 

12. Acrobasis juglemdis (LeBaron) 
Figures 138, 644 

Phycita juglandis LeBaron, Second annual report on the noxious 
insects of the State of Illinois, p. 23, 1872. 

Acrobasis juglandis (LeBaron) Riley, Fourth annual report on 
the noxious, beneficial and other insects of the State of 
Missouri, p. 42. — Barnes and McDunnough, Contributions, 
vol. 3, p. 221, 1917.— McDunnough, Check list. No. 6082, 
1939. — Moznette (and others), U. S. Dap. Agr. Farmers' 
Bull. 1829, p. 16, 1940.— Craighead, U. S. Dep. Agr. Misc. 
Publ. 657, p. 449, 1950. 

Acrobasis nebulella Dyar (not Riley), Proc. Ent. Soc. Washing- 
ton, vol. 10, p. 45, 1908. 

Acrobasis palliolella Dyar (not Ragonot), Proc. Ent. Soc. Wash- 
ington, vol. 10, p. 44, 1908. — Forbes (in part), Cornell 
Mem. 68, p. 617, 1923. 



Similar to the three preceding species except: White 
basal area more or less shaded with ashy gray; without 
rosy tints on lower basal and outer areas; generally 
paler in outer area, mouse gray, with the white dusting 
from midcosta somewhat more intense; a distinct 
blackish costal triangle following the antemedial line. 
Hind wing smoky fuscous shading to rather dull white 
towards base on the male, darker and more uniformly 
colored on the female. Alar expanse, 14-17 mm. 

Type locality: Illinois (tji^pe lost) . 

Food plants: Hickory, pecan, walnut, butternut 
(larvae feeding on leaves, buds, and flowers) . 

Disteibution: Illinois, Chicago (July); Missouri; 
Mississippi, Wiggins (May); Texas, Black Springs, 
Brownsville (May), Cuero (June), KerrvUle (May, 
June), Victoria (May, June); Georgia, Albany (July), 
Atlanta, Blackshear (May, June), Cairo (May, June); 
Florida, Monticello (May, Jime), Orlando (May), 
Palatka (May), Tallahassee (May); South Carolina, 
Mt. Pleasant (July); North Carolina, Edgecombe 
County (May), Plymouth (May); District of Columbia, 
Washington (June). 

A large reared series in the National Museum is 
mostly from pecan. Also before me a series reared 
from walnut and butternut that appears to be a suffused, 
dark form of juglandis. Two specimens of the latter 
series are from Ontario, Canada. 

In our Gulf States the species is of some importance 
as a defoliator of pecan, and is popularly known as 
the "pecan leaf casebearer." It has numerous refer- 
ences in economic literature. I have retained only one 
of these (Moznette, 1940), for it gives all the biological 
raformation available on the species under its correct 
specific name. Dyar's unfortunate identifications have 
greatly confused the nomenclature, with the result that 
most economic references previous to 1939 are under 
nebulella or paUiolella. Hulst (Phycitidae of N. Amer., 
p. 131, 1890) and Ragonot (Monograph, pt. 1, p. 120, 
1893) are also at fault in applying the name juglandis. 
Their descriptions apply to examples of indigeneUa and 
not to the "pecan leaf casebearer." 

The sex-scaling of juglandis is like that on feltella. 

13. Acrobasis sylviella Ely 

FlQUHB 144 

Acrobasis sylviella Ely, Proc. Ent. Soc. Washington, vol. 10, 
p. 161, 1908.— McDunnough, Check list. No. 6094, 1939. 

Forewing pale ashy gray; the basal area, thorax, and 
head but slightly paler, not contrastingly whitish; no 
triangular black spot on costa outside the antemedial 
line, the dark outer border of the antemedial line a 
narrow band or weak, diffused shade from costa. Hind 
wing pale smoky fuscous on male, slightly darker on 
female. Alar expanse, 19-21 mm. 

Type locality: East River, Conn, (type in USNM). 

Food plant: Ostrya. 

Distribution: United States: Connecticut, East 
River (July); Pennsylvania, New Brighton (May). 
Canada: Ontario, South March (June). 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



17 



The Pennsylvania specimen was in the Barnes 
Collection as cirrojerella Hulst. This is incorrect, 
however, as the male type of cirrofcrella is without 
sex-sc£,ling. The Canadian specimens (one male and 
one female) were reared from Ostrya and are responsible 
for the food plant record. They were tentatively 
identified by McDunnough as sylviella and I think 
correctly. They are considerably darker than the type 
series of the Pennsylvania specimen; but the fact that 
they were reared, and probably under excess moisture, 
would easily account for the difference. 

The black sex-scaling of syltriella is similar to that of 
feltella but slightly more extended, reaching slightly 
beyond basal foiu-th of costa on the underside of fore- 
wing. 

14. Acrobasis kearfottella Dyar 
FlGUBE 145 

Acrohasis kearfottella Dyar, Proc. Ent. Soe. Washington, vol. 7, 
p. 34, 1905.— Ely, Ins. Insc. Menstr., vol. 1, p. 53, 1913.— 
McDunnough, Check list, No. 6079, 1939. 

[•'■ The most distinct and strikingly marked of the Ameri- 
can Acrobasis species ; costal half or third of basal area 
of forewing snow white, this white area extending out 
along costa to subterminal line and broadening to in- 
clude the blackish discal spots; midcostal margin nar- 
rowly black-edged; white area uncrossed at any place 
by dark lines; subterminal line faint, but distinct, duU 
white; remainder of forewing dark gray-brown. Hind 
wing of male white shading to fuscous at apex and 
terminal margin ; hind wing of female pale glossy brown 
throughout. Thorax and head of male show white; of 
female concolorous with dark area of forewing. Alar 
expanse, 18-21 mm. 

Type locality: Cleveland, Ohio (type in USNM). 

Food plant: Hickory (larvae feeding on the leaves). 

Distribution: Ohio, Cleveland (June); New York, 
Ilion (July); Connecticut, East River (July); Pennsyl- 
vania, New Brighton (July); North Carolina, Black 
Mountain; Illinois, Oconee (Aug.). 

Very little is known of the life history. Ely's paper 
describes the cocoon. The black sex-scaling on the 
male is similar to that of juglandis and the preceding 
hickory-feeding species. 

15. Acrobasis caryae Grote 
Figures 137, 146 

Acrobasis caryae Grote, Papilio, vol. 1, p. 13, 1881; Bull. U. S. 
Geol. Geog. Surv. Terr., vol. 6, p. 591, 1882.— Hulst, 
Phycitidae of N. Amer., p. 122, 1890.— Ragonot, Mono- 
graph, pt. 1, p. 105, 1893. — Barnes and McDunnough, 
Contributions, vol. 2, p. 222. 1914.— Forbes, Cornell Mem. 
68, p. 617, 1923.— McDunnough, Check list. No. 6100, 
1939. — Moznette and others, U. S. Dep. Agr. Farmers' 
Bull. 1829, p. 2, 1940.— Craighead, U. S. Dep. Agr. Misc. 
Publ. 657, p. 449, 1950. 

Acrobasis caryaevorella Dyar (not Ragonot), Proc. Ent. Soc. 
Washington, vol. 10, p. 44, 1908. 

Acrobasis hebescella Dyar (not Hulst), Proc. Ent. Soc. Washing- 
ton, vol. 10, p. 44, 1908. 

Forewing glossy gray (in southern specimens from 
pecans pale and with little darker shading except 



nanowly along antemedial line) ; basal area concolorous 
with median area except in some of the darker speci- 
mens; northern specimens from hickory normally dark 
grayish fuscous; antemedial line whitish towards inner 
margin ; raised-scale ridge black, preceded by some white 
scaling and followed by a narrow, more or less obscured, 
flesh-colored patch; subterminal line pale gray, obscure; 
discal dots distinct and separate but not strongly 
contrasted against ground color. Hind wings smoky 
fuscous. Alar expanse, 18-20 mm. 

GenitaHa exhibiting no distinguishing specific char- 
acters; figured from southern male reared from pecan 
nut. The scale tufting on the eighth abdominal 
segment of the male consists of a single, rather long, 
central ventral tuft like that shown in figure 137 and 
similar to that of the European tji^e of the genus 
(tumidella). 

Type locality: Illinois (type in BM). 

Food plants: Hickory, pecan (overwintering larvae 
feeding in early spring upon opening leaves and in the 
stems of new growth; later generations in the nuts. 
Larva does not make a case during feeding period). 

Distribution: United States: Florida, Monticello 
(June, July, Aug.), Tallahassee (May) ; Georgia, Albany 
(July) ; Mississippi, Goodman (July) , Ocean Springs 
(May, Sept.), Pascagoula (June), Wiggins (June, July); 
Texas, Boerne (June), Bosque (May), Brownwood 
(Apr., May, June, July, Aug., Sept.), Colorado River 
(Apr., May), Cuero (Aug., Sept.), Dallas (May), Fort 
McKevett (June), Pecan Bayou (July), Pioneer (Aug.), 
San Saba (May), Texas A. and M. College Station 
(Jime, July), Victoria (June, July, Sept.); Illinois, 
Chicago, (July), Decatur (June); Pennsylvania, New 
Brighton (July, Aug., Sept.) ; North Carolina, Mill 
Brook; District of Columbia, Washington (May, June); 
Connecticut, East River (July, Aug.). Canada: On- 
tario, Merivale (June). 

This is the "pecan nut casebearer" of economic 
literature. It has a rather extended literature but is 
of importance only as a pecan pest in the Gulf States. 
Most of the economic references before 1929 are to 
hebescella and caryaevorella as a result of Dyar's mis- 
identification of those species. I cite only one economic 
reference here, as the Moznette (1940) paper gives all 
the biological information available on the species as 
a pecan insect. Its biology as a hickory insect in the 
north is imperfectly known. 

The sex-scaling of the male is characteristic, consist- 
ing of a short black patch on base of costa of forewing 
(as in feltella) and a long black streak along the top of 
cell on the underside of forewing. This combination 
is peculiar to caryae and evanescentella. 

16. Acrobasis evanescentella Dyar 

Figure 147 

Acrobasis evanescentella Dvar, Proc. Ent. Soc. Washington, vol. 
10, p. 44, 1908.— McDunnough, Check list, No. 6086, 1939. 

Doubtfully distinct from caryae. The dark areas of 
forewing beyond base have a purplish luster, and the 
pale (whitish dusting) is more distinct, forming a pale 



18 



UNITED STATES NATIONAL MUSEtTM BULLETIN 207 



grayish white spot on costa before subterminal line 
(and extended to include the discal spots) ; sex-scaling 
on underside of wing as in caryae, except that the 
streak along top of cell is somewhat obscured by an 
overlay of pale scales. Under these it is black (not 
"pale gray"as stated by Dyar) and no narrower than 
that of caryae. 

The only authentic specimens I have seen are those 
of the original type series. They are in excellent con- 
dition. The other specimens which Dyar later asso- 
ciated with them are all typical caryae. The note 
with the type lot ("Chittenden 250") tells nothing 
about the larval habits, so we do not know whether 
there are any biological characters to distinguish 
evanescentella from caryae. We shall have to hold the 
name until tbe biology is thoroughly studied. 

Type locality: Orlando, Fla. (May) (type in 
USNM). 

Food plant: Pecan. 

17. Acrobasis stigmella Dyar 

FlGtTKB 148 

Acrobasis stigmella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 43, 1908.— Forbes, Cornell Mem. 68, p. 616, 1923.— 
McDunnough, Check list. No. 6089, 1939. 

Fore wing purplish gray; the basal area very dark 
(darker than remainder of wing) ; scale ridge scarcely 
darker, its outer margin edged wdth dull rod; discal dots 
obscure; subterminal hne very faint. Hind wing of 
male pale smoky fuscous with a sUght ocherous tint; 
of female darker and without the ocherous tint on 
upper surface. 

On the male the upper surface of the head is yellowish 
white, the thorax, purplish gray. On the female the 
head and thorax are concolorous, purphsh gray. 
Alar expanse, 17-21 mm. 

Type locality: Fort Lee, N. J. (type lost). 

Food plant: Hickory. 

Distribution: New Jersey; Connecticut, East River 
(Aug.) ; District of Columbia, Washington (May, June) , 
Virginia, Falls Church (Jime) ; Blinois, Decatur (May), 
Putnam County (July). 

In his description Dyar states that it is based upon 
two males and one female from "Fort Lee, N. J., May 
1896 (H. G. Dyar)" and one female from "East River, 
Conn., Aug. 20, 1906 (Chas. R. Ely)." The female 
from East River is in the National Collection but there 
are no specimens from Fort Lee and none dated 1896. 
The specimen (a male in good condition) bearing Dyar's 
type label is one reared by him at Washington, D. C, 
Jime 19, 1900. This probably was overlooked by him 
at the time he prepared his description and not identified 
or labeled untU later. It is unquestionably stigmella 
but, in the light of his published declaration, cannot be 
accepted as the type. 

The species is quite distinct and easily identified — 
especially the males by their contrasting yeUow-white 
heads against their dark thoraces. The sex-scaling on 
underside of the male and the contrasting dark basal 
patch of forewing distinguish it from everything else 



except aurorella Ely. The sex-scaling consists of a nar- 
row black streak on forewing extending for about one- 
fourth of costa from base, a strongly contrasted and 
rather broad black midcostal streak on hind wing and 
some black scahng on the extreme base of the veins of 
cell of the hind wing. The underside of hind wing is 
otherwise a uniform ocherous white. 

The hfe history is also characteristic. In early spring 
(Mar.) the young overwintering larva is found within 
the unopened leaf-bud, its presence indicated by a 
small roimd frass Ud over the entrance hole. For a 
short time the larva feed within the bud, chiefly upon 
the bud sheath. When it opens the young leaves are 
partially eaten and then the larva enters the new shoot. 
Thereafter the entire feeding life and pupal period is 
spent within the new growth. The larva makes a 
larval case during this generation. The life history of 
later summer generations is not known. 

18. Acrobasis auroreUa Ely 
FlQUEE 149 

Acrobasis aurorella Ely, Proc. Ent. Soc. Washington, vol. 12, 
p. 67, 1910.— Forbes, CorneU Mem. 68, p. 616, 1923.— 
McDunnough, Check list, No. 6090, 1939. 

Close to and similar to stigmella but differing markedly 
in the ground color of the forewing. The sex-scaling is 
the same. The upper surface of the head on the male 
is also ocherous white but much duller and less con- 
trasted against the dark gray thorax. Forewing with 
basal area blackish gray sharply contrasted against re- 
mainder of wing; median and outer areas of wing pale 
pinkish ocherous or pale gray with a pinkish overcast; 
subterminal line extremely faint; discal dots distinct 
but not strongly contrasted. Hind wings as in stigmella. 
Alar expanse, 19-22 mm. 

Type locality: Washington, D. C. (type in USNM). 

Food plant: Unknown. 

Distribution: Connecticut, East River (Aug.); New 
York, Ilion (Aug.); New Jersey, Montclair (June); 
Pennsylvania, New Brighton (Sept.); District of Colum- 
bia, Washington (June). Also two specimens from the 
Fernald Collection, without locality and labeled "demo- 
tella Grt." 

19. Acrobasis peplifera Dyar 
FiGTJBE 150 

Acrobasis peplifera Dyar, Ins. Insc. Menstr., vol. 13, p. 13, 
1925.— McDunnough, Check list. No. 6106, 1939 

Forewing dark gray with a vinous tint; basal area a 
dark wine red ; scale ridge black, followed by a red line, 
then by a whitish line or triangle on inner margin, 
shading into black towards costa; discal dots obscured 
in the dark ground color, more or less confluent. Hind 
wings pale smoky fuscous, darker in the female than 
male; veins faintly outlined by dark scaling. Alar ex- 
panse, 13-17 mm. 

The male sex-scaUng on underside is as follows: On 
forewing a rather wide black stripe on basal fifth of 
costa ; on hind wing a black streak from base along top 
of cell, widening out to costa at its middle and putting 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



19 



out a thin black branch along lower vein of cell (this 
lower streak extending out as far as base of vein 2). 

Type locality: Millbrook, N. C. (type in USNM). 

Food plants: Hickory, pecan. 

Distribution: North Carolina, Elizabeth City (Aug.), 
MiUbrook (Aug.), New Bern (Aug.), Tryon (Aug.); 
Illinois, Putnam County (July) ; Arkansas, Washington 
County (Aug.); Texas, Cuero (July); Georgia, Albany 
(June) ; Florida, Monticello. 

Doubtfully distinct from exsulella, but the names had 
better be kept separate untU more is known about the 
biologies of the two color forms. 

20. Acrobasis exsulella (Zeller), new combination 

Figure 151 

Myelois exsulella Zeller, Isis von Oken, 1848, p. 868. 
Rhodophaea exsulella (Zeller) Ragonot, Ent. Amer., vol. 5, p. 114, 

1889; Monograph, pt. 1, p. 80, 1893. — Hulst, Phycitidae of 

N. Amer., p. 120, 1890.— McDunnough, Check list. No. 

6076, 1939. 
Acrobasis seplentrionella Dyar, Ins. Insc. Menstr., vol. 13, p. 13, 

1925.— McDunnough, Check list. No. 6107, 1939. (New 

synonymy.) 

Similar to peplifera except: Averaging larger; basal 
area of forewing more decidedly and evenly reddish; 
outer area somewhat paler; scale ridge not blackish, 
red and concolorous with remainder of basal area, very 
weak. Alar expanse, 14-20 mm. 

Type localities: North America [Georgia?] {exsulel- 
la, in Zool. Mus. Univ. Berlin) ; Florida (septentrioneUa, 
in USNM). 

Food plants: Hickory, pecan. 

Distribution: Florida (type, no other locality, 
Apr.), Orlando (Apr.); Texas, Brownwood (Mar.); 
North Carolina, Kaleigh (May); Maryland, Plummers 
Isl. (June). 

The types of both exsulella and seplentrionella are 
females. The former is supposed to be in Berlin. 
The figure of it given by Kagonot (Monograph, pi. 5, 
fig. 19) is a very good match for Dyar's species and I 
have no hesitation in synonymizing the latter. The 
scale ridge is present on forewing but could be easily 
overlooked, especially on a female that had been spread. 
Even on the unspread and unrubbed females in the 
National Collection it is not discernible except under 
considerable magnification. The structure is more 
prominent on the male. 

Our Texas specimens were reared from larvae feeding 
on the expanding buds of pecan. Dyar's paratype from 
North Carolina was reared from hickory. We have no 
further information on the biology. The sex-scaling 
of the male is like that of peplifera, which will probably 
prove to be nothing more than a variety or color form 
of exsulella. 

21. Acrobasis anguselia Grote 
Figure 152 

Acrobasis angusella Grote, North Amer. Ent., vol. 1, p. 51, 1880; 
Papilio, vol. 1, p. 14, 1880; BuU. U. S. Geol. Geogr. Surv. 
Terr., vol. 6, p. 590, 1882.— Ragonot, Monograph, pt. 1, 
p. 104, 1893. — Barnes and McDunnough, Contributions, 



vol. 2, p. 221, 1914.— Forbes, Cornell Mem. 68, p. 615, 
1923— McDunnough, Check list. No. 6091, 1939. 
Acrobasis eliella Dyar, Proc. Ent. Soc. Washington, vol. 10, p. 
43, 1908. 

Head and thorax whitish clay color, more or less 
shaded with pinkish or reddish suffusion (darker on 
females than on males). Forewing with basal area 
reddish and with some dusting of black scales on darker 
specimens; scale ridge black or black with an inter- 
mixture of red scales, followed outwardly by a reddish 
or reddish ocherous patch narrowing towards costa; 
pale antemedial line chiefly indicated on lower half of 
wing, obscure, followed on costa by a dark fuscous 
triangulate shade; median area gray with a diffused 
pale shade surrounding discal spots and extending to 
costa; discal dots separate, distinct, but not strongly 
contrasted against groimd color; sub terminal line 
distinct, denticulate, preceded by a narrow dark border 
and followed in terminal area by a broad reddish or 
reddish ocherous suffusion. Hind wing pale smoky 
fuscous. Alar expanse, 17-22 mm. 

Type localities: West Farms, N. Y. {anguselUi, 
in BM); East River, Conn, {eliella, in USNM). 

Food plant: Hickory (larvae boring in the leaf 
stems) . 

Distribution: United States: Massachusetts, North 
Adams (Aug.); New York, West Farms, Ilion (July, 
Aug.) ; New Jersey (June) ; Pennsylvania, New Brighton 
(May, Aug.) ; Maryland, Beltsvihe (May) ; Connecticut, 
East River (July, Aug., Sept.). Canada: Ontario, 
Ottawa (July). 

Barnes and McDunnough (1914) were correct in their 
criticism of Dyar's identification of angusella and in 
their reference of eliella to synonomy, but their descrip- 
tion of the male sex-scaUng is at fault. It is more 
correctly described by Ragonot in his monograph. It 
consists of a very short black patch on costa of forewing ; 
a long black streak along the upper vein of cell, expand- 
ing almost to the costal edge at middle and terminating 
well beyond the end of the cell, and from the base of 
this streak a second short black streak along lower fine 
of cell for about half its length. On some males there 
are also a few black scales on vein Ic shortly beyond its 



22. Acrobasis demotella Grote 
Figure 153 

Acrobasis demotella Grote, Papilio, vol. 1, p. 14, 1881; Bull. U. S. 
Geol. Geogr. Surv. Terr., vol. 6, p. 590, 1882.— Hulst, 
Phycitidae of N. Amer., p. 122, 1890.— Ragonot, Mono- 
graph, pt. 1, p. 103, 1893. — Dyar, Proc. Ent. Soc. Washing- 
ton, vol. 10, p. 42, 1908. — Barnes and McDunnough, Con- 
tributions, vol. 2, p. 221, 1914.— Forbes, Cornell Mem. 68, 
p. 616, 1923.— McDunnough, Check list. No. 6092, 1939. 

Color and markings in general similar to those of 
angusella except: Central area of forewing a uniformly 
suffused dark grayish fuscous; basal area a paler red- 
dish shade without dark dusting except for some 
fuscous smudging of the scale ridge on inner margin; 
antemedial line more distinct, dull white and on most 
specimens completely indicated to costa; discal dots 



20 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



and subterminal line much obscured, the latter not 
denticulate, and followed by a rather faint reddish 
shading, distinct only on dark unrubbed or unfaded 
specimens. Hind wing pale smoky fuscous. Alar 
expanse, 20-24 mm . 

Type locality: West Farms, N. Y. (type in BM). 

Food plant: Black walnut (larvae feeding in buds 
and stems). 

Distkibution: New Hampshire, Durham; iVew York, 
West Farms, Long Island ; Pennsylvania, New Brighton 
(June); North Carolina, Black Mountain (June), Tryon 
(May); Illinois, Chicago, Decatur (May), Putnam 
County (June); Missouri, St. Louis (June). 

The sex-scaling of the male is of the same type as 
that of angusella, differing as foUows: On forewing the 
black costal streak is longer, extending to basal fifth of 
costa; the upper streak on hind wing is somewhat 
shorter, narrow at base, swelling to an oblong patch at 
middle; the streak along lower vein of cell is broader 
and longer, extending to the lower outer angle of cell. 

Hulst gives a description of the larva and life history 
as supplied him by Fernald. Nothing substantial has 
been added since then to our Imowledge of the biology 
of the species. 

Genus Acrohasis, Species 23-40: A. latifasciella 
to A. tumidulella 

[Male: Apical process of gnathos trifurcate; forewing with 
raised-scale ridge; without sex scaling.] 

23. Acrobasis latifasciella Dyar 
Figure 139 

Acrobasis latifasciella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 45, 1908.— Forbes, Cornell Mem. 68, p. 617, 1923.— 
McDunnough, Check list, No. 6096, 1939. 

In color and macula tion similar to angusella Grote; 
but without black sex-scaling on underside of male 
wings. Alar expanse, 18-21 mm. 

Type locality: New Brighton, Pa. (type in USNM). 

Food plants: Hickory, walnut. 

Distribution: New York, Ilion (Sept.); Pennsyl- 
vania, New Brighton (Aug.); District oj Columbia, 
Washington; Maryland, Plummers Isl. (Aug.); Illinois, 
Putnam County (June). 

Nothing is known of the biology except the food 
plants. 

24. Acrobasis irrubriella Ely 
FiGTJHB 154 

Acrobasis irrubriella Ely, Proc. Ent. Soc. Washington, vol. 10> 
p. 161, 1908.— Forbes, Cornell Mem. 68, p. 618, 1923.— 
McDunnough, Check list, No. 6095, 1939. 

Color and maculation similar to those of angusella 
and latifasciella except: Thorax and basal area of fore- 
wing showing little or no trace of reddish scaling; 
reddish ocherous band on outer border of scale ridge 
narrower and fainter; no reddish shading in terminal 
area beyond subterminal line. Alar expanse, 19-21 
mm. 

Male genitalia with apex of uncus more rounded than 



that of latifasciella — at best, a character of doubtful 
specific value. 

Type locality: East River, Conn, (type in USNM). 

Food plant: Unknown, probably Carya (hickory or 
walnut). 

Distribution: Connecticut, East River (July); Indi- 
ana, Mineral Springs (Aug.). 

25. Acrobasis normella Dyar 

FlGTJRB 155 

Acrobasis normella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 46, 1908.— Forbes, Cornell Mem. 68, p. 617, 1923.— 
McDunnough, Check list, No. 6097, 1939. 

Similar to irrubriella except: Averages smaller; fore- 
wing of a more even glossy texture ; more white dusting 
on basal area, giving it a more decidedly pale gray ap- 
pearance; the white dusting from outer half of costa to 
and surroimding the discal dots also a trifle stronger. 
Alar expanse, 17-19 mm. 

Type locality: East River, Conn, (type in USNM). 

Food plant: Presumably hickory. 

Represented in the National Collection by 15 speci- 
mens from the type locaHty. In addition, there are 
before me two smaller (15 mm.), darker, more suffused 
specimens (male and female) from the Barnes collection 
that McDunnough had identified as irrubriella. They 
were reared from hickory (June) and are, I believe, only 
a color form of normella. The male genitalia (fig. 155a) 
show a stiiking departure in the assymetrical and greatly 
reduced lateral elements of the trifurcate apical projec- 
tion of gnathos, but apparently this is the result of a 
deformation of the organ in this particular specimen. 

26. Acrobasis malipennella Dyar 
Figure 156 

Acrobasis malipennella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 47, 1908.— Forbes, Cornell Mem. 68, p. 618, 1923.— 
McDunnough, Check list, No. 6093, 1939. 

Similar to the species following (dyarella) in color and 
markings except that the darker areas of forewing are 
dark fuscous and lack the intense red suffusion charac- 
teiistic of the latter. The structural differences noted 
by Dyar (the broader, shorter forewing, its broader 
cell, the close approximation of veins 2 and 3, different 
on each forewing, and the connate condition of 4 and 5 
and their remoteness from 3) are due to deformation. 
The male type never matured properly and the fore- 
wings are not fully developed. I beheve the specimen 
is nothing but a color form of dyarella and a freak at 
that. The male genitalia are very close, differing only 
in the somewhat more broadly rounded apex of gnathos, 
a difference of no specific significance in this group. It 
is a pity Dyar ever described it and a still greater pity 
that we cannot ignore bis name; for it will probably 
have to replace dyarella Ely which was based on normal 
specimens and is represented hj types in good condition. 

Type locality: East River, Conn. (Aug.; type in 
USNM). 

Food plant: Unknown. 

Known only from the unique type. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



21 



27. Acrobasis dyarella Ely 

Figure 157 

Acrobasis dyarella Ely, Proc. Ent. Soc. Washington, vol. 12, p. 67, 
1910.— McDunnough, Check list, No. 6099, 1939. 

Head yellow gray. Thorax much suflFused with red 
scaling. Basal area of forewing red dusted with whitish 
towards costa; scale ridge black, followed outwardly by 
a broad red band which extends and diffuses outwardly 
towards costa, obscuring and partially obliterating the 
blackish costal triangle; red scaling generally scattered 
over lower median area of wing; some obscure whitish 
dusting on median costal area and about the small, 
separated discal spots; sub terminal line faint, bordered 
inwardly and outwardly, except on costa, by reddish 
lines; terminal area and dark markings otherwise, dark 
gray. Hind wing glossy, pale smoky fuscous. Alar 
expanse, 19-20 mm. 

Type locality: East River, Conn, (type in USNM) . 

Food plant: Unknown. 

Represented only by the male type (Aug.) and female 
paratype (Sept.). 

28. Acrobasis ostryeUa Ely 

Figure 158 

Acrobasis ostryella Ely, Ins. Insc. Menstr., vol. 1, p. 54, 1913. — 
McDunnough, Check Ust, No. 6084, 1939. 

Similar to dyarella except: Reddish color more gener- 
ally diffused over outer areas of forewing, not forming 
a strongly accented band following the scale ridge, and 
of a purplish red shade; the coastal triangle and scale 
ridge contrastingly black. Hind wing dark smoky 
fuscous. Alar expanse, 15-18 mm. 

Type locality: East River, Conn, (type in USNM). 

Food plants: Ostrya virginiana, Carpinus. 

Distribution: United States: Connecticut, East 
River (July) . Canada: Ontario, South March (June) . 

29. Acrobasis secundella Ely 
Figure 159 

Acrobasis secundella Ely, Ins. Insc. Menstr., vol. 1, p. 55, 1913. — 
McDunnough, Check list, No. 6083, 1939. 

Doubtfully distinct from ostryella. The holotype 
and one other reared male from the type locality and 
two Canadian specimens before me are darker and a 
nearly uniformly suffused pm-plish, with a pale dusting 
on basal area of forewing, and about the discal spots a 
rather faint and pale gray rather than white. However, 
other reared specimens from hazel are a perfect match 
for the type of ostryella. Alar expanse, 15-18 mm. 

The male genitalia show a trifling difference in the 
length of the lateral projections of the apical projection 
of gnathos. Ely states that the larval case of secundella 
is longer and more slender than that of ostryella; but 
the life history needs further investigation before any 
such difference can be evaluated. 

Type locality: East River, Conn, (type in USNM). 

Food plant: Corylus. 

Distribution: United States: Connecticut, East 
River (July). 

Canada: Ontario, Merivale (June). 



30. Acrobasis coryliella Dyar 
Figure 160 

Acrobasis coryliella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 47, 1908.— Ely, Ins. Insc. Menstr., vol. 1, p. 53, 1913. — 
Forbes, Cornell Mem. 68, p. 618, 1923. — McDunnough, 
Check list, No. 6098, 1939. 

Forewing a dull, rather lusterless gray, paler in the 
central area and without any reddish or purpUsh 
shading; following scale ridge on inner margin a faintly 
ocherous patch obscured by gray scaling; the scale ridge 
and other dark markings blackish, but the usual black 
triangle on costa following antemedial line replaced by 
a narrow line; the usual transverse dark shade from 
inner upper edge of subterminal line to middle of lower 
margin; subterminal line bordered inwardly by a narrow 
blackish line ; discal dots normally confluent, forming a 
curved line along discocellular vein of cell. Hind wing 
pale smoky fuscous; darker on female. Alar expanse, 
17-20 mm. 

Type locality: Unspecified [New York?] (type in 
USNM). 

Food plant: Corylus. 

Distribution: New York; Connecticut, East River 
(July, Aug.); Massachusetts, Newton Highlands; Illi- 
nois, Decatur (June, July), Putnam County (June, 
July). 

In addition to a long series reared from hazel at East 
River, Conn., there are three specimens from the 
Fernald and Brooklyn Museum Collections (one male 
and two females from Illinois) labeled "Acrobasis 
hebescella," the two females labeled "type"; and one 
male from Decatur, 111., which McDunnough had 
tentatively identified as A. sylviella Ely. 

The species is easy to recognize from its rather uni- 
form gray shade and strongly contrasted, short, black 
scale ridge. 

31. Acrobasis hebescella Hulst 

Acrobasis hebescella Hulst, Phycitidae of N. Amer., p. 126, 1890. — 
Ragonot, Monograph, pt. 1, p. 109, 1893. — Barnes and 
McDunnough, Contributions, vol. 3, p. 194, 1916. — 
McDunnough, Check list. No. 6085, 1939. 

The only authentic representation of this species is the 
female type, which is in very poor condition but ap- 
parently does not, or did not originally, differ in any 
significant detail from coryliella Dyar except as to its 
host. It was reared from a cocoon found on oak. This 
may or may not be significant. The name is just 
another of those that must wait for clarification until 
someone shall make a careful and more thorough 
study of the life histories of the various Acrobasis 
species. Alar expanse, 16.5 mm. 

Type locality: "Jersey pines, June" (type in 
AMNH, ex Rutgers). 

Food plant: Oak. 

32. Acrobasis cirroferella Hulst 

Acrobasis cirroferella Hulst, Canadian Ent., vol. 24, p. 60, 1892. — 
McDunnough, Check list. No. 6109, 1939. 

The type is a male without abdomen. There is no 
sex-scaling. Close to coryliella, but with dark areas of 



22 



XHSriTED STATES NATIONAL MUSEUM BULLETIN 207 



forewing more brownish gray and the whitish areas 
more strongly contrasted; central costal area dis- 
tinctly white; dark outer margin of antemedial line 
brown, narrow on costa; costa before it white, entire 
basal area having some white dusting; discal dots 
distinct, separate. Hind wing pale fuscous. Alar ex- 
panse, 18 mm. 

Type locality: Austin, Tex. (type in AMNH, ex 
Rutgers). 

Food plant: Unknown. 

The above description is drawn from the type, which 
is worn and faded. I have seen nothing that exactly 
matches it. 

33. Acrobasis cvmnlae Dyar and Heinrich 

FiGUEE 162 

Acrobasis cunulae Dyar and Heinrich, Proc. Ent. Soc. Washing- 
ton, vol. 31, p. 37, 1929. — McDunnough, Check list, No. 
6102, 1939. 

Forewing pale slate gray; basal area paler; midcostal 
area with some faint pale dusting, especially about the 
discal spots; scale ridge weak and little or not at all 
darker than the ground color of the wing; sub terminal 
line distinct, with a narrow, dentate, dark inner border, 
neither the pale line itself nor its dark border strongly 
contrasted; discal dots blackish, separated and rather 
conspicuous. Hind wing smoky fuscous. Alar ex- 
panse, 20-24 mm. 

The male genitaha have what appears to be a dis- 
tinguishing specific character in the decidedly broad- 
ened lateral elements of the apical projection of gnathos. 

Type locality: Mobile, Ala. (type in USNM). 

Food plant: Pecan. 

Disteibution: Florida, Monticello (May); Georgia, 
Cairo (May), DeWitt (M.&y); Alabama, Auburn (May), 
Mobile (May); Mississippi, Wiggins (May). 

Close to but apparently distinct from caryivoreUa. 

34. Acrobasis caryivorella Ragonot 
FiGUBB 161 

Acrobasis caryivorella Ragonot, N. Amer. Phycitidae, p. 4, 1887; 
Monograph, pt. 1, p. 108, 1893.— Hulst, U. S. Nat. Mus. 
Bull. 52, p. 419, 1903.— Hill, Florida Ent., vol. 21, p. 12, 
1938.— McDunnough, Check list, No. 6087, 1939.— Craig- 
head, U. S. Dep. Agr. Misc. Publ. 657, p. 450, 1950. 

Acrobasis caryaevorella Hulst, Phycitidae of N. Amer., p. 121, 
1890 (misspelling). 

Acrobasis conjivorella Hulst, in J. B. Smith, List of the Lepi- 
doptera of Boreal America, No. 4262, 1891 (misspelling). 

Acrobasis caryae Dyar (not Grote), Proc. Ent. Soc. Washington, 
vol. 10, p. 46, 1908. 

Forewing dark bluish gray, nearly black; basal area 
towards costa (above the scale ridge) and a small tri- 
angular area on costa adjacent to subterminal line 
powdered with grayish white; scale ridge black, on 
some specimens bordered outwardly by a faint ocherous 
red patch (especially on specimens reared from hickory) ; 
subterminal line pale gray, faint; discal dots distinct, 
separate or confluent. Hind wing smoky white to 
smoky fuscous, darker on female than on male. Alar 
expanse, 19-24 mm. 



Type locality: Missoiu-i (type in Paris Mus.). 

Food plants: Hickory, walnut, pecan. Larvae 
boring in the buds and new growth of the stems. 

Distribution: Massachusetts, Melrose (June, July); 
Maryland, Beltsville (May, July, Aug.), HyattsviUe 
(June), Prince Georges County, (June) ; North Carolina, 
"N. Car. Dept. Agr."; South Carolina, Summerton 
(May); Georgia, Albany (May), Barnesville (May, 
3wie>) ; Florida, MonticeUo (May, July), Orlando (Apr.); 
Mississippi, Biloxi (Aug.), State College; Missouri; 
Texas, Austin (Aug.), Brovrawood (May, June), 
Menard (June), Victoria, Waco (Apr., May, June); 
New Mexico, Carlsbad (Aug.). 

The species is of some importance in the Gulf States 
as an enemy of pecan and the name caryivorella has 
appeared several times in economic pubhcations but 
nearly always wrongly applied to specimens of caryae 
Grote. Specimens of the true caryivorella have also 
been identified as caryae on the basis of Dyar's (1908) 
misapplication of the two names. 

35. Acrobasis comacornella (Hulst), new combination 
Figure 136 

Acrocaula comacornella Hulst, Canadian Ent., vol. 32, p. 170, 
1900.— McDunnough, Check list. No. 6078, 1939. 

Forewing with dark areas glossy, purplish brown; 
extreme base of wing dark, followed by a rather narrow 
subbasal whitish area; median costal area narrowly 
whitish; outer area uniformly dark; subterminal line 
obscure, not bordered by darker lines; discal dots 
distinct, dark brovra, the lower dot tvrice the size of 
the upper; a little red on the antemedial line towards 
costa. Hind wing whitish with a faint ocherous 
fuscous tint; rather glossy; veins very faintly darkened; 
a narrow dark line along termen. Alar expanse, 17 mm. 

Detail of male genitalia figured from type. 

Type locality: Blanco Coimty, Tex. (type in 
AMNH, ex Rutgers). 

Food plant: Unknown. 

The male type is obviously an Acrobasis without sex- 
scaling. It resembles very much a specimen of caryi- 
vorella (in USNM) from Victoria, Tex., except that the 
hind wing is paler, the subterminal line less distinct, 
and the discal dots larger and more contrasted. I sus- 
pect that it is nothing more than a variety of caryi- 
voreUa. 

36. Acrobasis betuleUa Hulst 
FiGUBB 164 

Acrobasis betulella Hulst, Phycitidae of N. Amer., p. 125, 1890. — 
Ragonot, Monograph, pt. 1, p. 107, 1893. — Dyar, Proc. 
Ent. Soc. Washington, vol. 10, p. 47, 1908.— Forbes, Cornell 
Mem. 68, p. 618, 1923.— McDunnough, Check list. No. 
6101, 1939. 

Forewing dark gray faintly tinted with reddish violet 
and with white dusting on basal area and forming a 
triangulate patch from costa before subterminal line, 
the white dusting faint (less contrasted than on caryi- 
vorella); scale ridge black, without any red bordering 
patch or bar; discal dots at end of cell distinct, separate; 
antemedial line obscure, almost obsolete; subterminal 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



23 



line faint. Hind wing smoky fuscous. Alar expanse, 
20-24 mm. 

Male genitalia figured from specimen from the origi- 
nal Hulst series in the National Collection (bearing a 
Hulst "type" label and presumably a paratype). 

Type locality: New York (type, 9, in AMNH, ex 
Rutgers). 

Food plant: Betula. 

Distribution: United States: Maine (no further 
locality, July), Sebec Lake (July); New Hampshire, 
Center Harbor (July), Hampton (June); Massachusetts, 
Amherst (June, July), Melrose; Connecticut, East River 
(July); New York (no further locality, July); Colorado, 
Piatt Canon (July); California, Siskiyou County. 
Canada: Ontario, Trenton. 

There is little or nothing to separate collected speci- 
mens of hetulella from comtoniella or rubrifasciella except 
the complete absence of any reddish outer border to the 
scale ridge on forewing (a distinction that does not hold 
for all specimens of rubrifasciella) and a more glossy 
sheen on the specimens of comptoniella and rubrifasciella 
(a comparative difference that is displayed only when 
series of the three species are seen side by side). The 
thing that really distinguishes betulella is its host plant, 
Betula. It also differs from the other species of Ameri- 
can Acrobasis with the scale ridge on forewing in that it 
has been found in the Rocky Mountain region and in 
areas west thereof. The Colorado and California 
records are from the specimens in the National Collec- 
tion mentioned by Dyar. I have seen no later collec- 
tions from any area west of the Rockies. 

37. Acrobasis rubrifasciella Packard 
Figure 165 

Acrobasis rubrifasciella Packard, Ann. Lye. Nat. Hist. New 
York, vol. 10, p. 267, 1873.— Grote, Bull. U. S. Geol. Geog. 
Surv. Terr., vol. 4, p. 693, 1878.— Hulst, Phycitidae of N. 
Amer., p. 124, 1890. — Ragonot, Monograph, pt. 1, p. 106, 
1893.— Forbes, Cornell Mem. 68, p. 617, 1923.— McDun- 
nough, Canadian Ent., vol. 65, p. 206, 1933; Check list. 
No. 6103, 1939. 

Phycis rubifasciella (Packard) Beutenmuller, Canadian Ent., 
vol. 22, p. 16, 1890 (spellingl and larva). 

Acrobasis alnella McDunnough, Canadian Ent., vol. 64, p. 36, 
1922. 

Similar to betulella except that normally there is a 
faint, but distinguishable, wine-red bar outwardly 
bordering the scale ridge on forewing and that, in 
series, the forewing surface has a more glossy appear- 
ance. Neither of these differences holds for aU speci- 
mens; nor are the genitalic differences noted by Mc- 
Dunnough (1933), the width of the lateral flanges of 
the apical projection of gnathos, reliable as a specific 
character. Alar expanse, 20-24 mm. 

Type localities: Orono, Maine (rubrifasciella, in 
MCZ); Ottawa, Canada (alnella, Canadian Nat. CoU.). 

Food plant: Alnus. 

Distribution: United States: Maine, Orono (June); 
New Hampshire, Durham; Massachusetts, Amherst 
(June) ; Connecticut, East River (July) ; New York, Cats- 
kill Mts.; NoHh Carolina, Black Mts. Canada: On- 



tario, Ottawa (July) ; Quebec, Meach Lake (July) ; Nova 
Scotia, Truro (July, Aug.). 

38. Acrobasis comptoniella Hulst 

Acrobasis comptoniella Hulst, Phycitidae of N. Amer., p. 125 
1890. — Ragonot, Monograph, pt. 1, p. 108, 1893. — Dyar' 
Proc. Ent. Soc. Wash., vol. 10, p. 46, 1908.— Forbes, Cornell 
Mem. 68, p. 618, 1923.— McDunnough, Check hst, No. 6104, 
1939.— Craighead, U. S. Dept. Agr. Misc. Pub. 657, p 
450, 1950. 

Superficially like rubrifasciella except wine-red bar 
bordering scale ridge of forewing always present and 
of a more intense and darker shade. Hind wing gen- 
erally darker. Alar expanse, 21-25 mm. 

Type locality: Long Island, N. Y. (type, 9, in 
AMNH, ex Rutgers; paratjrpe, 9, from type locality 
in USNM). 

Food plants: Comptonia and Myrica. 

Distribution: United States: Maine, Bar Harbor 
(July), Kennebunk (July), Mount Desert (July); New 
Hampshire, Center Harbor, Durham; Massachusetts, 
Melrose (June), North Saugus; Connecticut, East River 
(July) ; New York, Long Island ; New Jersey, Bergenfield, 
New Lisbon (June); Michigan, Dickinson County. 
Canada: Ontario, Trenton. 

The best way to separate comptoniella from the two 
preceding species is by rearing from their respective 
food plants. 

39. Acrobasis myricella Barnes and McDunnough 
FiouBE 163 

Acrobasis comptoniella Grossbeck (not Hulst), Bull. Amer. Mus. 

Nat. Hist., vol. 37, p. 129, 1917. 
Acrobasis myricella Barnes and McDunnough, Contributions, 

vol. 3, p. 221, 1917. — McDunnough, Check list. No. 6105, 

1939. 

Close to comptoniella Hulst, but superficially quite 
different; smaller, white dusting in pale (basal and 
central-costal) areas much denser and more contrasted ; 
dark areas blackish gray with very faint purplish over- 
tint; reddish bar bordering black scale ridge narrow and 
inconspicuous; sub terminal line distinct, white. Alar 
expanse, 17-19 mm. 

Type locality: Fort Myers, Fla. (Apr., May; type 
in USNM). 

Food plant: Myrica. 

Distribution: Florida. 

Except for the type series from the type locality I 
have seen only one specimen (a female from Royal Palm 
State Park, Fla., Apr. 5, 1929, F. M. Jones, collector) 
that can be definitely assigned to the name myricella. 
We have, however, in the National Collection a series 
of males and females reared by Chas. R. Ely at East 
River, Conn., from Myrica cerifera, that are inter- 
mediate between myricella and typical comptoniella 
(reared from Comptonia), like the former in size and in 
the intensity and extent of the white dusting on fore- 
wing, but with the subterminal line obscure as in 
comptoniella. I think they are only a variety of 
comptoniella. Indeed, myricella may prove to be only 



24 



■UNITED STATES NATIONAL MUSEUM BULLETIN 207 



a race of Hulst's species; but with our present knowl- 
edge we must retain it as a distinct species. 

40. Acrobasis tumidulella (Ragonot), new combination 
FiGTJBE 645 

Cateremna tumidulella Ragonot, N. Amer. Phycitidae, p. 13, 

1887. 
Seneca tumidulella (Ragonot) Hulst, Phycitidae of N. Amer., p. 

178, 1890. 
Hyphantidium tumidulellum (Ragonot) Hampson, in Ragonot, 

Monograph, pt. 2, p. 74, 1901. — McDunnough, Check list, 

No. 6321, 1939. 

This species was based on a single female with the 
habitus and raised-scale ridge of an Acrobasis but with 
vein 4 absent from hind wing. Bourgogne informs me 
that the venation is alike on both hind wings. On the 
strength of this venation the species was referred to 
Group II of the Phycitinae and made the type of Hulst's 
Seneea. However, I am firmly convinced that the speci- 
men is nothing but an Acrobasis with abnormal venation, 
another of those freaks that turn up all too frequently in 
the Phycitidae. I have examined the female genitaUa of 
the type (figured here) and can find nothing to distin- 
guish them from those of caryimrella. I suspect that 
tumidvlella is nothing more than an abnormal specimen 
of caryivorella; but we shall have to await final disposi- 
tion of the name imtil a similar freak male is recovered 
from the type locahty. 

Type locality: Florida (type in Paris Mus.). 

Food plant: Unknown. 

3. Genus Rhodophaea Guenee 

Rhodophaea Gu§n6e, Ann. Soc. Ent. France, ser. 2, vol. 3, p. 312, 
1845; Europaeorum Microlepidopterorum index methodi- 
cus . . . , p. 74, 1845. — Ragonot, Ent. Monthly Mag., vol. 
22, p. 19, 1885; Monograph, pt. 1, p. 63, 68, 1893. (Type 
of genus; Phycis advenella Zincken; figs. 166, 649.) 

Characters of Acrobasis except: Male antenna simple, 
basal segment cylindrical, no sinus in base of shaft; 
forewing always smooth; vein 2 of forewing from ceU 
before lower outer angle, but somewhat nearer the 
angle than in Acrobasis; male genitaha with apical 
process of gnathos an elongate hook (partially divided 
on advenella) ; eighth abdominal segment of male simple 
or (on advenella) with midventral hair tuft. 

This genus is distinguished from the smooth-winged 
species of Acrobasis only by its simple male antenna. 
Our two American species do not go any too well with 
advenella, the European type of the genus, differing in 
having an undivided apical projection from gnathos 
and simple eighth abdominal segment. However, in 
these characters they agree with other obviously con- 
generic European species, marmorea (Haworth), lega- 
teUa (Hiibner), suavella (Zincken). B. advenella has a 
somewhat differently shaped transtilla from caliginella, 
supposita, and the three aforementioned European 
species. In all of these the terminal margin of trans- 
tiUa is more or less indented (as in Acrobasis) while in 
advenella it is rather deeply U-shaped. 

None of the American species that hitherto have been 



listed imder Rhodophaea belongs there. They have 
entirely different genitaha. 

41. Rhodophaea caliginella (Hulet), new combination 

Figure 647 

Nephopteryx caliginella Hulst, Ent. Amer., vol. 3, p. 131, 1887; 

vol. 5, p. 156, 1889. 
Mineola caliginella (Hulst), Phycitidae of N. Amer., p. 128, 

1890; U. S. Nat. Mus. Bull. 52, p. 419, 1902.— Barnes and 

McDunnough, Contributions, vol. 4, p. 174, 1918. — 

McDunnough, Check list. No. 6113, 1939. 
Acrobasis caliginella (Hulst) Ragonot, Monograph, pt. 1, p. 115, 

1893. 
Myelois caliginoidella Dyar, Proc. Ent. Soc. Washington, vol. 7, 

p. 33, 1905.— McDunnough, Check hst. No. 6072, 1939. 

(New synonymy.) 

Similar in color and markings to Acrobasis comptella 
Kagonot except that the narrow black line outwardly 
bordering the whitish basal area of forewing does not 
extend all the way to inner margin. This slight dif- 
ference in macidation seems to be constant and will 
distinguish the females of the two species which, other- 
wise, are difficult to tell apart. Alar expanse, 18-25 
mm. 

Type localities: Arizona (caliginella, in AMNH, 
ex Rutgers); Santa Clara, Calif, (caliginoidella, in 
USNM). 

Food plant: Scrub oak. This host record from 
reared specimens in National Collection received from 
Commander Dammers, Riverside, Calif., June 1938. 

Distribution: California, Alma (Aug.), Atascadero 
(July), Los Angeles County (July), Riverside (June), 
San Diego (May, June, July, Aug.), Santa Clara; 
Arizona. 

In addition to the female type in the Rutgers Col- 
lection there is also a female from Arizona ("7810") in 
the National Collection bearing Hulst's "type" label. 
This specimen was originally in the Fernald Collection. 
A female from Cahfornia donated by Hulst to the 
Brookl3Ti Museum Collection and twice labeled "Acro- 
basis comptella" in his and Ragonot's handwriting is 
also in the National Museum. This specimen, except 
that it lacks an abdomen, is in good condition. It is 
certainly caliginella and presumably was responsible 
for Hulst's sjTionymizing of caliginella and comptella. 
The males of caliginella have hitherto been known as 
caliginoidella Dyar. Hulst evidently never saw a male 
of his species. 

42. Rhodophaea supposita (Heinrich), new combination 

FiQUEEs 167, 648 

Mineola supposita Heinrich, Proc. Ent. Soc. Washington, vol. 
42, p. 33, 1940. 

Foreiving very dark grayish fuscous with a powder- 
ing of white scales on basal and midcostal areas and 
very faintly in the area bordering terman; antemedial 
line narrow, slanting from inner third of costa to just 
before middle of inner margin, sUghtly notched at 
vein lb, pale ashy gray bordered inwardly from top of 
cell to inner margin by a dull red triangular patch which 
has an obscure, straight, blackish hue along its inner 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



25 



edge; inner margin from base to antemedial line nar- 
rowly bordered by reddish scales; subterminal line 
narrow, slightly outcurved between vein 6 and lower 
fold, pale gray, inwardly bordered by a narrow black 
line; a blackish fuscous patch outwardly bordering the 
antemedial line from costa to cell ; a similar dark shade 
on costa near apex; these blackish patches shading into 
the dark central area of wing; black discal dots at end 
of cell distinct and separate; some obsciu-e dull red 
shading in terminal area toward tornus; along termen 
a narrow discontinuous black line. Hind wing pale 
smoky fuscous with veins, terminal margin, and apical 
area darker. Alar expanse, 16-20 mm. 

Male genitalia similar to those of caliginella except 
transtilla broader at apex and arms of anellus stouter. 
Female genitalia differing from those of caliginella 
chiefly in that there are no patches of small scobina- 
tions in bursa near its junction with ductus bursae. 

Type locality.: Vancouver, British Columbia 
(type in Canadian Nat. Coll.). 

Food plant: Cotoneaster. 

Known so far only from the type series from Van- 
couver. It is distinguished from caliginella chiefly by 
its generally darker color. 

4. Genus Trachycera Ragonot 

Trachycera Ragonot, Monograph, pt. 1, p. 2, 1893. (Type of 
genus, Rhodophaea pallicornella Ragonot.) 

This genus is close to Rhodophaea, being distinguished 
from it chiefly by male characters. The male of 
pallicornella from which these were drawn is apparently 
lost. Clarke was unable to locate it at Paris, where it 
should have been; and, as no other males agreeing with 
Ragonot's description or figure (Monograph, pi. 5, 
fig. 20) are available, we are unable to check his charac- 
ters. Ragonot separates Trachycera from Rhodophaea 
widely in his generic key (Monograph, pt. 1, pp. xliii 
and xliv) on the basis of the trifid or bifid condition of 
the median vein of hind wing. This is an error, how- 
ever, for the true Rhodophaea species are no more bifid 
than is Trachycera. The female of pallicornella has 
essentially the same venation as the type of Rhodophaea 
(advenella). 

The distinguishing male characters given by Ragonot 
are: Serratiform male antenna; very short labial palpus 
(scarcely reaching to middle of face); and minute maxil- 
lary palpus. 

The female has a pair of small signa in the bursa 
copulatrix, developed as granulate cups (as in Davara). 

43. Trachycera pallicornella (Ragonot) 
Figure 650 

Rhodophaea pallicornella Ragonot, N. Amer. Phycitidae, p. ,3 
1887.— Hulst, Phycitidae of N. Amer., p. 119, 1890. 

Trachycera pallicornella (Ragonot), Monograph, pt. 1, p. 2, 
1893.— McDunnough, Check list, No. 6057, 1939. 

The holotype of pallicornella is a female labeled in 
Ragonot's handwriting "ty. or. PI. V, fig. 20." Accord- 
ing to Clarke the Ragonot figure represents it accu- 

300329 — 56 3 



rately. Unfortunately it is a mended specimen and 
the glued-on abdomen is spurious. Its genitalia are 
pyraustine rather then phycitid. 

I have before me a female which is an exact match for 
Ragonot's figure. It is somewhat smaller (15 mm.) 
than the type ("19 mm.") but this difference is easily 
within the normal range for many species of medium- 
sized Phycitinae. It was collected at Devils River, 
Tex. (May). The genitalia are figured from this 
specimen. 

Forewing pale gray with some blackish dusting on 
base, especially on base of costa; a faint purplish gray 
shade on lower part of postmedian area; antemedial 
band rather broad, red narrowly lined with black on 
inner and outer sides, the back outer margin somewhat 
widened at costa; subterminal line narrow, nearly 
vertical, with an outward bulge between vein 6 and 
lower fold, whitish, bordered inwardly by a narrow 
black line and by a black outer patch at costa near apex ; 
discal and terminal dots obsolete. Hind wing dull 
whitish with a faint yellow tint and shading to pale 
fuscous towards apex. Alar expanse, 15-19 mm. 

Type locality: Texas (type in Paris Mus.), 

Food plant: Unknown. 

5. Anabasis, new genus 

Type of genus: Myelois ochrodesma ZeUar. 

Tongue well developed. Antenna pubescent; on 
male, basal segment enlarged and angulate (as in 
Acrohasis), shaft simple. Labial palpus upturned 
reaching to vertex (slightly longer on female than on 
male). Maxillary palpus rather broadly scaled. Fore- 
wing with a transverse, antemedian ridge of raised 
scales; 11 veins; vein 2 from before but near lower 
outer angle of cell; 3 from the angle, approximate to 2 
at base, nearer to 2 than to 4 ; 4 and 5 closely approxi- 
mate for some distance from base; 6 from below upper 
angle of cell, slightly bent towards base; 10 from the 
cell; male without costal fold. Hind wing with vein 2 
from well before lower outer angle of cell ; 3 from before 
but very near the angle ; 4 and 5 from the angle, closely 
contiguous (or more or less anastomosed) for about 
half their lengths ; 7 and 8 contiguous or weakly anasto- 
mosed for some distance beyond cell; cell one-third 
the length of wing; discocellvdar vein curved. Eighth 
abdominal segment of male with ventral hair tuft and 
a pair of modified, ventrolateral tufts. 

Genitalic characters as in Acrohasis except: Harpe 
with a transverse, sclerotized ridge from base of costa 
to lower outer angle of sacculus; a cluster of modified 
scales on outer edge of inner margin in the angle be- 
tween sacculus and cucuUus; terminal margin of 
vinculum more rounded. (These may be only specific 
characters.) 

A development from and quite close to Acrohasis, 
which it replaces in tropical America; distinguished 
from that genus chiefly by shorter cell and the con- 
tiguous position of veins 4 and 5 of hind wing. Except 



26 



TXNITED STATES NATIONAL MUSEUM BULLETIN 207 



on denuded wings under strong magnification they 
appear to be stalked for half their lengths. Contains 
one tropical American species. 

44. Anabasis ocfarodesma (Zeller), new combination 
Figures 168, 652 

Myelois ochrodesma Zeller, Horae Soc. Ent. Rossicae, vol. 16, 

p. 209, 1881. 
Piesmopoda ochrodesma (Zeller) Ragonot, Monograph, pt. 1, 

p. 165, 1893. 
Acrobasis crassisquamella Hampson, in Ragonot, Monograph, 

pt. 2, p. 520, 1901 (new synonymy). 

Forewing grayish brown finely powdered with 
blackish scales and with a faint rosy suffusion; ante- 
medial line oblique, narrow, obscured by a heavy whit- 
ish ocherous ridge of raised scales on its inner margin 
and bordered outwardly by a narrow black fine; sub- 
terminal line obsciu-e, narrow, when distinguishable, 
sinuate, ocherous white, bordered inwardly by an 
obscm-e, broken, black line; discal and terminal dots 
obsolete. Hind wing smoky white; veins and a narrow 
shade along termen, fuscous. Alar expanse, 13-16 mm. 

Genitalia as given for the genus; bursa of female with 
small signum. 

Type localities: Honda, Colombia (ochrodesma, in 
BM) ; Teapa, Tabasco, Mexico (crassisquamella, in BM) . 

Food plants: Cassia alata. Cassia nodosa. Cassia tora 
(U. S. Dep. Agr. Florida rearings; larva a leaf-folder), 
Sciacassia siamea. 

Distribution: United States: Florida, Coconut 
Grove (May), Miami (May), St. Petersburg (June). 
Puerto Kico: Bayam6n (Sept.), Coamo Springs (Apr.), 
Mayagiiez (Jan.), Puerto Eeal (Vieques Isl., Apr.), Rio 
Piedras (Feb.). Virgin Islands: Kingshill (St. Croix, 
Oct., Nov., Dec). Cuba: Santiago de las Vegas 
(Havana, Dec). Grenada. Jamaica. Trinidad: 
Fyzabad (Feb.), Tunapuna (Apr.). Mexico: Tabasco, 
Teapa. Guatemala: Quirigu^ (May). PanamX: 
Corozal (July), Porto BeUo (Apr.). Colombia: Honda. 

An easUy recognized tropical American species whose 
range has been extended into southern Florida. 



6. Genus Mildrixia Dyar 

Mildrixia Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 405, 1914. 
(Type of genus; Mildrixia consiitutionella Dyar.) 

Tongue well developed. Antenna of male strongly 
ciliate, the cUia over three times longer than width of 
segments; basal segment elongate, subtubular, flat- 
tened and broadening towards apex; first segment of 
shaft swoUen and with a short spine and scale tuft from 
inner upper angle (this with the rough scahng of basal 
segment gives the latter when fully scaled the appear- 
ance of the triangulate first segment of Acrobasis); 
antenna of female simple and very weakly pubescent. 
Labial palpus obliquely ascending, reaching to slightly 
above vertex; moderately rough scaled beneath; third 
segment acuminate, about two-thirds the length of 
second. Maxillary palpus moderately large, squamous. 
Forewing narrowly elongate, with transverse, ante- 



medial ridge of raised scales; 11 veins; vein 2 from 
before, but near lower outer angle of cell; 3 from the 
angle, approximate to 2 at base and for a short distance 
beyond; 4 and 5 connate or very shortly stalked; 6 
from very close to upper angle of cell, closely approxi- 
mate to 8 at base, nearly straight (very slightly bent 
towards base); 10 from the cell, closely approximate to 
the stalk of 8-9 for some distance from cell; male with- 
out costal fold. Hind wing with vein 2 from before 
but close to lower outer angle of cell; 3 contiguous with 
the stalk of 4-5 for some distance from angle, on 
undenuded wings appears stalked with 4-5; 4 and 5 
stalked for more than half their lengths; 7 and 8 
contiguous or weakly anastomosed for a short distance 
beyond cell; cell one-third the length of wing; disco- 
cellular vein oblique. Eighth abdominal segment of 
male with broad ventral hair tuft. 

Male genitalia of the old world Acrobasis type 
except: Uncus subtriangulate, its apex rather broadly 
rounded; transtiUa terminating posteriorly in a U- 
shaped projection with elongate, slender, widely spaced 
and divergent arms; vinculum longer than broad, evenly 
tapering to bluntly pointed terminal margin; anellus 
an elongate, semitubular plate with short lateral lobes 
near base; penis armed with a short, sclerotized plate 
and numerous sclerotized wrinklings. 

Female genitalia of the Acrobasis type but without 
any sclerotized plate or plates at genital opening; a 
single signum in bursa, developed as a small, cupped, 
granulate plate. 

A distinct genus, distinguished at once by its male 
antenna, venation, and transtiUa. Contains one 
tropical American species. 

45. Mildrixia coustitutionella Dyar 

FiGUHES 169, 651 

Mildrixia constitutionella Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 405, 1914. 

Forewing grayish fuscous with some whitish dusting, 
especially in median area about the discal spots; ante- 
medial scale ridge blackish preceded by a narrow white 
line and followed by an indistinct dark shade; discal 
dots at end of cell distinct, black; just beyond the lower 
discal dot, an outwardly angled mark from the upper 
edge of which a narrow dark shade extends to the 
inner costal edge of the subterminal line (distinct only 
on weU-marked and unrubbed specimens) ; subterminal 
fine narrow, denticulate, pale, bordered inwardly and 
outwardly by somewhat broader dark lines; terminal 
dots blackish, more or less confluent. Hind wing trans- 
lucent, opalescent white, the veins faintly darkened 
toward their outer extremities, especially on the females; 
a dark shade along costa and a narrow one along termen. 
Alar expanse, 19-22 mm. 

Genitaha as given for the genus. 

Type locality: Jalapa, Mexico (type in USNM). 

Food plant: Unknown. 

Distribution: Mf;xico: Jalapa. Guatemala: Vol- 
cdn Santa Maria (June, July, Oct.). 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



27 



7. Genus Sematoneura Ragonot 

Sematoneura Ragonot, Nouv. Gen., p. 9, 1888; Monograph, pt. 
1, p. 136, 1893. (Type of genus: Sematoneura atrovenosella 
Ragonot.) 

Tongue well developed. Antenna of male shortly 
ciliate (cilia a trifle longer than width of shaft), of 
female weakly pubescent. Labial palpus upturned, 
reaching vertex, cylindrical, slender; third segment 
about two-thirds length of second, acuminate. Max- 
illary palpus filiform. Forewing smooth; 11 veins; 
vein 2 from well before lower outer angle of cell; 

3 from the angle, much nearer to 4 than to 2; 4 and 5 
closely approximate for a short distance from cell; 
6 from well below upper angle of cell, straight; 8 and 9 
long stalked; 10 shortly stalked with 8-9; male without 
costal fold. Hind wing with vein 2 from well before 
outer angle of cell; 3 from before, but near the angle; 

4 and 5 closely approximate for a short distance from 
the angle; 7 and 8 closely approximate or contiguous 
beyond ceU; cell about half the length of wing; dis- 
cocellular vein curved. Eighth abdominal segment of 
male with a single, broad ventral hair tuft. 

Male genitalia with apical process of gnathos a 
simple, elongate hook, slightly notched at apex. Uncus 
broadly triangulate. TranstiUa complete, stout, arched, 
its central area developed as a flat, broad lobe with 
slightly concave terminal margin. Harpe with costa 
sclerotized and produced at apex into a short project- 
ing digitus. Anellus a slightly curved plate with 
moderately long lateral arms. Aedeagus simple; penis 
armed with a single elongate, moderately stout cornutus, 
about one-third as long as aedeagus. Vinculum stout, 
as broad as or a trifle broader than long, tapering to 
broad, truncate terminal margin. 

Female genitalia with biu-sa and ductus bursae 
simple, without signum, smooth except for minute 
granulations in bursa ; ductus biu-sae shorter than bm-sa ; 
genital opening simple; ductus seminalis from bursa 
near its junction with ductus bursae. 

The foregoing description is drawn from the type 
species {atrovenosella). In male genitalia the new 
species (abitus), tentatively included in the genus, 
departs in some apparently essential details of structure, 
having a different type of transtilla and gnathos and 
lacking the apical projection from costa of harpe; but 
in all other structural characters it agrees with atroveno- 
sella. When its female is discovered a new generic 
placement may be necessary. 

46. Sematoneura atrovenosella Ragonot 
Figures 4, 171, 653 

Sematoneura atrovenosella Ragonot, Nouv. Gen., p. 10, 1888; 
Monograph, pt. 1, p. 136, 1893. 

Forewing gray with more or less ochraceous dusting 
above inner margin and in outer area between the veins; 
the veins conspicuously outUned by blackish scaling; a 
similar narrow, dark line along the lower fold; these 
dark lines expanded and intensified at basal third 
indicating the remains of an antemedial band, and 
broken in outer area by a rather broad, faint, pale 



subterminal band; lower discal dot at end of cell 
faintly indicated; a line of blackish dots along term en 
between the vein ends. Hind wings dusky white, 
translucent; the veins darkened and a narrow dark 
line along termen. Alar expanse, 26-35 mm. 

Genitaha as given for the genus. 

Type locality: Chancbamayo, Per6 (type in Zool. 
Mus. Univ. Berlin). 

Food plant: Unknown. 

Distribution: Mexico: Coatepec. Costa Rica: 
Huan Vinas (Jan., Feb., Nov.), Tuis (May). Colom- 
bia: La Selva (San Juan Chaco, Sept.), Juntas (San 
Juan Chaco, Feb.). Ecuador: Alpayacu (Rfo Pas- 
taza), Quito. Peri6: Chancbamayo, Santo Domingo 
(Nov.). Argentina: Tucuman. 

One example before me (a female from Santo Domin- 
go, Peru, 6,000 ft.) differs in coloration from normal 
specimens in having a dark suffusion over the basal area 
to the antemedial line and a distinct antemedial pale 
line with continuous, black outer border. It also lacks 
any trace of ocherous dusting on the forewing. In 
genitaha and otherwise in color and maculation it is 
normal. I believe that it is only a color form. The 
specimen was from the unplaced material in the British 
Museum. 

47. Sematoneura abituB, new species 

Figure 172 

Similar in color and markings to atrovenosella except a 
short strongly contrasted black streak just below costa 
at base and a broad black streak along median fold 
extending from base to end of cell. The ciliations of the 
male antenna are also a trifle shorter than those of 
atrovenosella. Alar expanse, 31 mm. 

The male genitalia differ markedly from those of 
atrovenosella in several details. Apical projection of 
gnathos is a rather short triangulate, pointed hook. 
The costa of harpe is broadly sclerotized but lacks the 
projecting digitus at apex. The gnathos is developed 
into a strongly sclerotized, hairpinlike, backwardly 
projecting loop with dense scobinations along its inner 
margin. 

Type locality: Alpayacu, Rio Pastaza, East Ecua- 
dor (type in BM). 

Food plant: Unknown. 

Described from unique male type collected by M. G. 
Palmer at 6,000 ft. The specimen is not in good con- 
dition but the essential features of the pattern are 
distinguishable and the male genitalia are so distinctive 
that description seems justified. In the absence of a 
female the generic placement cannot be made with 
absolute certainty. I expect, however, that the female 
genitalia will exhibit no radical difference from those of 
the type of the genus. 

8. Genus Hypsipyla Ragonot 

Hypsipyla Ragonot, Nouv. Gen., p. 10, 1888; Monograph, pt. 1, 
p. 137, 1893. (Type of genus: Hypsipyla pagodclla Ragonot, 
synonym of Magiria robusta Moore; India; figs. 173, 656.) 

Tongue well developed. Antenna of male shortly 



28 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



ciliate (cilia but slightly longer than width of shaft, 
except on male of dorsimacula where they are about 
twice as long as width of shaft). Labial palpus of male 
upturned, reaching vertex, slender ; third segment about 
half as long as second, acuminate; of female obhquely 
ascending. Maxillary palpus filiform (moderately large 
in pagodella and grandella, minute in other species). 
Forewing smooth; 11 veins; vein 2 from well before 
lower outer angle of cell; 3 from the angle, much closer 
to 4 than to 2 ; 4 and 5 approximate for a short distance 
from cell, occasionally connate; rarely short stalked; 6 
more or less bent towards base and more or less approxi- 
mate to upper angle of cell; 10 normally from the cell, 
rarely connate or shortly stalked with 8-9 ; male without 
costal fold. Hind wing with vein 2 from well before 
outer angle of cell; 3 from before, but near the angle; 4 
and 5 normally shortly stalked, occasionally connate, 
partially anastomosed or (in some large females) closely 
approximate for a short distance from cell; 7 and 8 
closely approximate beyond cell; cell half or (males of 
grandella) somewhat less than half the length of wing; 
discocellular vein curved. Eighth abdominal segment 
of male simple or with two or three pairs of ventral hair 
tufts. 

Male genitalia with apical process of gnathos a hook, 
forked at apex. Uncus more or less triangulate; apex 
rounded. TranstiUa complete, stout, arched, its cen- 
tral area produced into two widely spaced horns; the 
latter stout in aU species except pagodella. Harpe with 
costa strongly sclerotized but not produced at apex. 
Anellus V- or U-shaped with long lateral arms. Aedea- 
gus simple; penis armed with a single, more or less 
twisted, fattened bladelike cornutus (except dorsima- 
cula). Vinculum stout, short or but slightly longer 
than broad, with truncate, broad terminal margin. 

Female genitaUa with bursa and ductus bursae 
simple, unsclerotized except for a narrow band along 
ventral margin of genital opening; with or without 
signum; when present, the latter developed as a small, 
scobinate, cup-shaped plate; ductus bursae shorter 
than bm-sa; ductus seminalis from bursa at its junction 
with ductus bursae. 

The. genus is very close to and difficult to distinguish 
from Sematoneura. It is characterized chiefly by the 
bent condition of vein 6 of forewing. In Sematoneura 
this vein is always perfectly straight and remote from 
8-9 at base. In Hypsipyla 10 is also normally from the 
cell and there is frequent stalking of 4 and 5 of hind 
wing, neither of these conditions occurs in Sematoneura; 
but the venation is so individually variable in Hypsipyla 
that it cannot be trusted. 

48. Hypsipyla grandella (Zeller) 
Figures 5, 174, 655 

Nephoteryx grandella Zeller, Isis von Oken, 1848, p. 881. 
Hypsipyla grandella (Zeller) Ragonot, Monograph, pt. 1, p. 139, 

1893.— Dyar, Ins. Insc. Menstr., vol. 7, p. 41, 1919. — Monte, 

Rev. de Ent., Brazil, vol. 3, p. 281, 1933. 
Hypsipyla cnabella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 405, 

1914. 



Forewing grayish fuscous shaded (especially on lower 
half of wing) with dull rust-red; veins outlined in black; 
antemedial pale line narrow, incomplete, roimded out- 
ward at middle and indented at vein lb, bordered out- 
wardly by a narrow, discontinuous, black line; beyond 
this in median area Isetween top of cell and vein lb, an 
expanded faint whitish patch; some faint whitish dust- 
ing also in the middle-outer area between the veins; 
subterminal line faint, indicated chiefly by the intensi- 
fied black streaks bordering it inwardly on the veins, 
sinuate, deeply notched at lower fold; discal spots 
obsolete; terminal black dots between the vein ends 
distinct. Hind wing hyaline white with a fuscous 
shade along costa, some fuscous shading on the vein 
ends and a narrow fuscous line along termen. Alar 
expanse, 23^5 mm. 

Male genitalia with uncus rather abruptly narrowed 
beyond its broad base, the apex narrowly roimded; 
apical hook of gnathos narrow, short; paired horns of 
transtilla curving outward (away from each other); 
vinculum distinctly broader than long, its terminal 
margin very broad and but slightly convex, nearly 
straight. Eighth abdominal segment of male simple. 
Female genitalia with signum. 

Type localities: Brazil (grandella, location of type 
unknown to me) ; C6rdoba, Mexico (cnabella, ia.\JSNM). 

Food plants: Cedrela and Svnetenia (larva bores iri 
fruits and branches). 

Distribution: United States: Florida, Miami 
(Nov.). Mexico: C6rdoba (Feb., Sept.), Jalapa. 
Guatemala: Cayuga (Apr., May, Oct.), Chejel (June), 
Quirigud (June). Honduhas: La Cambra (Feb.). 
Costa Eica: Avangarez (July), Juan Vinas (Jan., 
Feb., Nov.), San Jos6 (Jan.), San de Montes de Oca 
(Oct.), Tins (May, June). PanamA: Almirante (Aug.), 
Summit (C. Z., Mar.). Puerto Rico: Cayey (May). 
Cuba: Santiago de las Vegas (Apr.) . Haiti: Petion- 
ville (June, Dec). Jamaica. Trinidad (Dec). Col- 
ombia: "Above Eio Negro." Venezuela: El VaUe 
(July), Maracay, Trompillo (July). British Guiana: 
Georgetown (July). Brazil: Aragatuba (Sao Paulo, 
Apr.), Bala (Feb.), Campo Bello, Castro (Parana), 
Espirito Santo, Nova Teutonia (May), Santa Catarina 
(Aug., Sept.). Paraguay: Sapucay (Oct.), Villarrica 
(Sept., Oct.). Ecuador: Loj a. Argentina: Tucumdn. 
PerIj: Lima. 

Generally distributed throughout tropical America 
wherever its food plants occvu-. 

A native American species close to and superficially 
similar to the Indina H. robusta; but with different male 
and female genitalia. It is apparently of some eco- 
nomic importance in the West Indies and South America 
as a pest of mahogany and the Cedrela species. Like 
many borers it varies greatly in size, and the venation 
is more than ordinarily unstable even for a phycitid. 
Vein 10 of forewing may be from the cell, separated 
from, closely approximate or connate with 8-9 or some- 
times shortly stalked with them. Veins 4 and 5 may 
be anything from approximate towards base to shortly 
stalked. Vein 6 is always slightly bent towards base 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



29 



but less so and more remote from the base of 8-9 on 
large females than on the smaller females and average- 
size males. On the hind wing 4 and 5 are usually 
shortly stalked or connate but on some large specimens 
are closely approximate for nearly half their lengths 
beyond the lower angle of the cell. 

49. Hypsipyla ferrealis (Hampson), new combination 

FiGUBES 176, 657 
Crocidomera ferrealis Hampson, Ann. Mag. Nat. Hist., ser. 10, 
vol. 4, p. 352, 1929. 

Maculation of forewing similar to that of grandella 
except: Brownish fuscous, the general color decidedly 
more brown than gray, the dull reddish dusting giving 
the wing a somewhat rosy brown tint; dark lining on the 
veins less conspicuous and more discontinuous; ante- 
medial pale line very faint, obsolete on many specimens, 
indicated chiefly by its broken black outer margin ; the 
white spot beyond this black margin (conspicuous in 
grandella) absent or indicated only by a faint ocherous 
white shade ; subterminal line very faint, indicated by a 
black shading on the veins along its inner margin, sinu- 
ate, rather deeply indentate at vein 6 and lower fold. 
Hind wing smoky fuscous with a faint brownish or 
ocherous tint, more or less smoky white towards base ; 
veins darkly outlined. Eighth abdominal segment of 
male with three pairs of ventral hair tufts. Alar 
expanse, 20-43 mm. 

Male genitalia with uncus of the same shape, but 
wider than that of grandella; vinculum narrower, 
longer than broad; transtilla with horns of central area 
divergent, forming a roimd-bottomed V, the central 
connecting part of the transtilla slender. Female geni- 
talia without signum. 

Type locality: SLxaola Kiver, Costa Rica (type in 
BM). 

Food plant: Carapa guianensis (larvae feeding on 
the seeds). 

Distribution: Costa Rica: Cachl, Cain, Juan 
Vinas (May), Puerto Lim6n, Sixaola River (May, 
Sept.), Tuis (May). Colombia: San Antonio (Dec). 
Venezuela: Maturaca (Sept.). French Guiana: 
Cayenne, St. Jean Maroni, St. Laurent Maroni. 
Trinidad: Caparo. Brazil: Parfi. 

A distinct species easily distinguished from grandella 
by its dark hind wings. The foregoing food plant and 
Venezuelan records are from a series of small reared 
specimens (20-24 mm.) submitted by Dr. Ballon in 
1942. These are not only considerably smaller than 
average from the other localities hsted; but are some- 
what grayer in color. The genitalia, however, are 
like those of typical Costa Rican examples. In 
veneLtion ferrealis appears somewhat less variable than 
grandella. Vein 10 of forewing is always from the 
cell and 4 and 5 usually shortly stalked, rarely con- 
tiguous for a short distance beyond the cell. 

50. Hypsipyla doreimacula (Schaus), new combination 

Figures 175, 654 

Myelois dorsimacula Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. lli 
p. 245, 1913. 



The female type is badly rubbed and the markings 
consequently obscured; ground color bronzy brown; 
faint indications of a pale antemedial hne rather far 
out on wing; fainter indications of a subterminal line; 
at end of ceU a dark brown spot on discocellular vein 
and shortly separated from it a similar spot in cell, 
between them a pale spot (this marking at end of cell 
seems the characteristic pattern character of the 
species) ; below discocellular vein, on lower fold, a short 
blackish streak. Hind wing semihyaline, lilacine, 
darker towards apex; veins not appreciably darker. 
Alar expanse, 40 mm. 

Female genitalia without signum. 

Type locality: Sixaola River, Costa Rica (Sept.) 
(type in USNM). 

Food plant: Unknown. 

In addition to the female type I have before me 
what I believe to be a male of the same species from 
the Janse Collection, taken at La Selva, San Juan, on 
the Chaco slopes of Colombia (4,600 ft., Sept.). It ia 
as badly rubbed as the type but shows the same charac- 
teristic markings at end of cell and on the fold beneath; 
ground color of forewing rust-red; hind wing hyaline 
white with a very faint ocherous tint; antennal ciliations 
longer than on other species of the genus; at least 
twice as long as width of shaft. Male genitalia with 
stouter gnathos and quite differently shaped transtUla 
from previous species, the prongs of transtilla rather 
narrowly separated; penis without cornutus. The 
venation is alike on both specimens except for vein 
10 of forewing, which is from the cell on the female 
and short-stalked with 8-9 on the male; 4 and 5 of 
fore and hind wings are short-stalked; 6 of forewing 
is sharply bent towards base and connate with 8-9. 
Eighth abdominal segment of male simple. 

51. Hypsipyla fluviatella Schaus 
Figure 177 

Hypsipyla fluviatella Schaus, Ann. Mag. Nat. Hist., ser. 8, 
vol. 11, p. 246, 1913. 

Forewing long, and narrower in proportion than other 
species of the genus; reddish brown, darker brown in 
basal area; antemedial line w*^ll out towards middle 
of wing, indicated by its brown outer border, the 
latter out-bent from costa, thence nearly vertical to 
inner margin, forming three lunules, defined by narrow, 
pale buff inner shadings (the remains of the pale 
antemedial line) , also preceded on inner margin by some 
silvery gray dusting; a narrow, elongate, pale buff 
patch on inner margin at tomus; a similar, wider, pale 
streak from ceU to outer margin, occupying the space 
between veins 5 and 8 and bisected longitudinally by 
a narrow red-brown streak along vein 6. Hind wing 
hyaline white, inner margin rather broadly tinted with 
ocherous, and a narrow ocherous line along outer 
margin. Alar expanse, 45-46 mm. 

Male genitalia with uncus broad throughout, its 
terminal margin broadly rounded. Transtilla com- 
plete but the central fusion weak; the horns widely 
spaced, forming a broad, shallow U. Apical process 



30 



UNITED STATES NATIONAL MUSEUM BULLETIN 2 07 



of gnathos rather broad, oval, flattened. Eighth 
abdominal segment of male simple. Female unknown. 

Type locality: Sixaola Elver, Costa Eica Ctype in 
USNM). 

Food plant: Unknown. 

A distinct species easily distinguished by its narrow, 
peculiarly marked, red-brown forewings and the oval, 
flattened shape of the apical process of gnathos. The 
male hind wings are distinctly triangulate, but this is 
probably only a sex character. 

The forewing venation is fairly stable; vein 10 from 
the cell; 6 bent towards base and narrowly separated 
from 8-9 at base; 4 and 5 closely approximate for a 
short distance from ceU. Hind wing with veins 4 and 5 
shortly stalked or closely approximate for some distance 
from the cell. 

The species is represented only by the type series 
of four males from the type locality. 

9. Genus Hemiptilocera Bagonot 

Hemiptilocera Ragonot, Nouv. Gen., p. 9, 1888; Monograph, pt. 
1, p. 144, 1893. (Type of genus: Hemiptilocera chino- 
graphella Ragonot.) 

Tongue well developed. Antenna of male with basa^ 
segment elongated, cylindrical; a tuft of scales on base 
of shaft (weak on chinographella) ; shaft unipectinate 
for two-thirds, crenulate and pubescent beyond. An- 
tenna of female like that of the male type (chino- 
graphella) except for lack of scale tuft on shaft and for 
shorter basal segment; on other species of the genus 
shaft simple and pubescent. Labial palpus ascending; 
reaching to or nearly to vertex (shorter on chinographella 
than on other species); slender. Maxillary palpus 
small, squamous. Forewing smooth; 11 veins; vein 
2 from well before lower outer angle of cell; 3 from the 
angle, much closer to 4 than to 2; 4 and 5 connate 
(chinographella) or closely approximate at base and 
for a very short distance from cell (other species of 
genus); 6 bent towards base, close to or connate with 
8 at base; 10 from the ceU, closely approximate to the 
stalk of 8-9 for some distance from cell; male without 
costal fold. Hind wing with vein 2 from well before 
outer angle of ceU; 3 from the angle, connate with or 
closely approximate to 4-5; 4 and 5 stalked for about 
half their lengths; 7 and 8 closely approximate beyond 
cell; cell one-half or slightly less than one-half the 
length of wing; discocellular vein curved. Eighth 
abdominal segment of male with one or more paired 
hair tufts and sternal plates modified, one strongly 
Bclerotized element in the form of an open loop. 

Male genitaha with apical process of gnathos a hook 
with notched apex. Uncus triangulate, apex bluntly 
pointed. Transtilla complete, stout, arched, its central 
area produced into widely spaced horns. Harpe with 
costa strongly sclerotized (produced at apex on bigrana 
and plumigerella, not produced on chinographella). 
Anellus with short, stout, lateral arms. Penis armed 
with strongly sclerotized cornutus and numerous 



sclerotized wrinklings. Vinculum stout, longer than 
broad, terminal margin broad and more or less indented. 

Female genitalia with signum developed as a small, 
strongly scobinate pocket; bursa large; ductus bursas 
short, more or less sclerotized and strongly scobinate- 
granulate, the scobinations and granulations extending 
into bursa for a short distance from place of junction 
with ductus; genital opening with sclerotized and more 
or less wrinkled plate on lower margin (except on 
exoleta); ductus seminalis from bursa near junction of 
bursa and ductus bursae. 

This genus is distinguished by its pectinate male 
antenna, the strong, stalking of veins 4-5 of hind wing, 
the long vinculum with broad terminal margin, the 
short arms of aneUus and its short, granulate-scobinate 
ductus biursae. Eventually it may have to be restricted 
to its type species (chinographella) and a new generic 
placement found for the other species now included. 
All of these have simple pubescent female antennae; 
while those of chinographella are pectinate in both sexes. 
There are also several differences between males of 
chinographella and those of plumigerella and bigrana 
(notably in the shape of the transtilla, the costal 
development of harpe, and the size of the antennal 
tuft) ; but unfortunately we do not know the males of 
three other species (letharda, jocarella, exoleta) and until 
they are known it seems the wiser com-se not to attempt 
further generic separation. All the species have similar 
habitus and wing maculation and (except for exoleta) 
female genitalia showing only specific differences. 

52. Hemiptilocera chinographella Ragonot 

Figures 178, 658 

Hemiptilocera chinographella Ragonot, Nouv. Gen., p. 9, 1888; 
Monograph, pt. 1, p. 144, 1893. 

Male antenna with scale tuft on base of shaft weak, 
clay colored. Antenna of female pectinate, the pecti- 
nations a trifle shorter than those of the male. Thorax 
and basal segment of antenna clay-yellow. Forewing 
clay yellow ("ohvaceous ocherous" according to 
Eagonot) dusted and shaded with dull reddish brown 
and dark grayish fuscous, the ground color predomi- 
nating in the basal area, along the costa and (more 
faintly) bordering the termen and as a narrow longi- 
tudinal streak between the transverse line and including 
at its middle the lower discal spot; antemedial line 
faint, indicated chiefly by the broken elements of its 
outer dark border (a short notched blackish streak 
slanting outwardly from costa, a blackish spot on top 
of cell, another on lower vein of cell and a third on lower 
fold, these three blackish spots in a vertical line out 
near middle of wing, on a few well-marked specimens 
connected by a very faint, twice-outciu-ved dark line) ; 
subterminal line somewhat stronger, inwardly margined 
by a black spot, outwardly margined by a duller dark 
shade, more or less accented at costa and on the veins; 
discal dots at end of cell separated, distinct, especially 
the lower one, black; along termen a row of distinct, 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



31 



well contrasted, black dots (these a rather characteristic 
feature of most of the species of the genus); on the 
female a conspicuous whitish patch on inner margin at 
inner edge of the subterminal line (this whitish patch 
not present on the males before me). Hind wing 
semihyaline, shaded with smoky fuscous towards apex, 
on the veins and nan-owly along termen. Eighth 
abdominal segment of male with a single moderately 
long pair of ventrolateral hair tufts. Alar expanse, 
22-26 mm. 

Male genitalia with central part of transtilla quadri- 
form (the horns flattened and with flattened lobes from 
their bases, the space between the horns even through- 
out); harpe with costa not produced at apex; apex of 
cornutus enlarged, sharply bent and bearing a row of 
thornlike spines. Female genitalia with ventral plate 
at genital opening smooth or nearly so ; granulations of 
ductus bursae dense and forming a continuous sclero- 
tized mass. 

Type locality: Chanchamayo, Peru (type in Zool. 
Mus. Univ. Berlin). 

Food plant: Unknown. 

Distribution: Feench Guiana: Cayenne. Brazil: 
Par^. Perij: Chanchamayo. 

53. Hemiptilocera bigrana (Zeller) 
Figures 180, 660 

Myelois bigrana Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 200, 

1881. 
Hemiptilocera bigrana (Zeller) Ragonot, Monograph, pt. 1, p. 145, 

1893. 

Male antenna with hair tuft strong, black. Antenna 
of female pubescent. Thorax and base of antenna dull 
whitish dusted with fuscous. Forewing pale gray dusted 
with dull rosy and dark grayish fuscous ; the dark dust- 
ing more evenly distributed than in chinographella; 
lower discal spot at end of cell large, strongly contrasted ; 
upper spot weak or not distinguishable ; antemedial line 
not defined; subterminal defined by its dark borders, 
which consist of blackish streaks on the veins. Hind 
wing hyaline white with a faint smoky fuscous shade at 
apex, on the outer half of the veins and narrowly along 
termen. Eighth abdominal segment of male with a long, 
strong pair of ventrolateral hair tufts and two other 
pairs of modified scale tufts. Alar expanse, 25-29 mm. 

Male genitalia with central part of gnathos a stout 
crescent-shaped projection with the horns widely 
spaced ; harpe with apex of costa produced into a short 
spine ; cornutus a spatulate ribbed blade. Female with 
ventral plate at genital opening deeply wrinkled ; ductus 
bursae with a central, elongate, irregular patch of scob- 
inations, extending into adjacent area of bursa. 

Type locality: Honda, Colombia (type in BM). 

Food plant: Unknown. 

Distribution: Mexico: Guerrero (Aug.), Iguala 
{Guerrero, June), Popocatepetl Parks (Distrito Federal, 
June). Colombia: Honda. 



The Mexican records are from specimens in the U. S. 
National Museum. 

54. Hemiptilocera plumigerella (Ragonot), new combination 

Figure 179 

Nephopteryx plumigerella Ragonot, Nouv. Gen., p. 14, 1888; 
Monograph, pt. 1, p. 261, 1893. 

This species is known only from the male t3rpe. From 
Ragonot's description the forewing color and markings 
must be similar to those of chinographella except much 
more heavily overshaded with vinous brown. The lower 
discal spot is conspicuous as in bigrana and the antennal 
tuft is stout and ochraceous in color. Hind wing iri- 
descent, semitransparent, grayish brown. Eighth 
abdominal segment of male with a single pair of long, 
stout [ventrolateral hair tufts and another pair of 
much shorter, central tufts. Alar expanse, 21 mm. 

Male genitalia with transtilla as in bigrana, except 
that the crescent-shaped central projection is much more 
slender ; cornutus a short bluntly pointed plate with a 
row of short, blunt spines near apex. 

Type locality: "America Meridionalis" (type in 
Paris Mus.). 

Food plant: Unknown. 

55. Hemiptilocera letharda (Schaus), new combination 
Figure 662 

Chloropaschia letharda Schaus, Proc. Ent. Soc. Washington, 
vol. 24, p. 237, 1922; Ann. Carnegie Mus., Pittsburgh, vol. 
16, p. 112, 1925. 

Forewing olive buff shaded with vinaceous fawn on 
subcoastal, median, and basal areas ; the wing markings 
black, and some faint scatterings of black scales on fore- 
wing and thorax; antemedial line indicated only by 
broken fragments of its narrow outer border; discal 
black spots at end of cell both conspicuous, the lower 
one large ; black dots on veins forming the inner margin 
of the pale subterminal line and black dots along termen 
also conspicuous. Hind wing semitransparent, smoky 
white, the veins slightly darkened ; a dark shade towards 
apex and a narrow dark line along termen. Alar 
expanse, 29 mm. 

Female genitalia similar to those of bigrana but with 
a heavier concentration of scobinations in ductus bursae 
and a much stronger signum. 

Type locality: Cabima, Panamd (May; tj^e in 
USNM). 

Food plant: Unknown. 

In addition to the female type I have before me a 
British Museum female from Presidio, Mexico, origi- 
nally identified as H. bigrana. It is paler than the 
Schaus type, but this apparently is due to its faded and 
slightly rubbed condition. Schaus recognized his 
original misplacement of the species in the Epipaschiidae 
and in his 1925 paper referred it to the Phycitinae, trans- 
ferring the specimen to Hemiptilocera in the National 
Collection; but I am unable to find any published 
reference of his or Dyar's giving of the generic reference. 



32 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



56. Eemlptilocera jocarella (Schaus) 

FlQITBE 659 

Acrdbasis jocarella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p. 245, 1913. 

Hemiptilocera jocarella (Schaus) Dyar, Ins. Insc. Menstr, vol. 7, 
p. 42, 1919. 

Forewing reddish brown, the reddish (vinous) shade 
predominant in cell and, broadly, along lower fold; a 
dull olivaceous shade along costa and in terminal area; 
antemedial line obsolete, indicated only by fragments of 
its outer border (a black narrow streak from costa to 
cell, a black spot on lower margin of cell, and a thin, 
in-bent black streak from vein lb to inner margin); 
subterminal line indicated by a pale black margined 
spot on costa and an outward series of short whitish 
streaks on veins 6 to lb, these spots inwardly and out- 
wardly margined by black dots; a series of black dots 
along termen (less distinct than in the other species of 
the genus) ; the usual black discal spots, only the lower 
one pronounced, and it but slightly so. Hind wing 
glossy purplish or smoky brown; the veins but faintly 
darkened ; a fine dark line along termen. Alar expanse, 
21-26 mm. 

Female genitalia exhibit (in the amount of scobina- 
tion of ductus bursae and the smaller size of signmn) 
but trifling differences from those of letharda. The male 
of jocarella is unknown. 

Ttpe locality: Avangarez, Costa Rica (type in 
USNM). 

Food plant: Unknown. 

Distribution: Costa Rica: Avengarez (July). 
PanamA: Porto Bello (Dec). Brazil: Nova Teutonia 
(May). 

These records from four females in the U. S. National 
Museum. When males can be associated it is quite 
ikely that jocarella will prove to be only the female 
form of plumigerella. 

57. Hemiptilocera exoleta (Zeller) 
Figure 661 

Myelois exoleta Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 201, 

1881. 
Hemiptilocera exoleta (Zeller) Ragonot, Monograph, pt. 1, p. 146, 

1893. 

This species is represented only by the female type. 
I have seen nothing that matches Ragonot's description 
or Zeller's rather crude figure; but from both and from 
details of the female genitalia the reference is doubtful. 
Ragonot himseK questions the correctness of his generic 
placement; but in the absence of a male no better 
placement could or can be made. 

The forewing shows the usual distinctively contrasted 
row of terminal dots and the maculation otherwise is 
that of a Hemiptilocera except that the usual discal dots 
are replaced by a reddish lunule. According to Ragonot 
the cell of hind wing is also short for a Hemiptilocera. 
Alar expanse, 25 mm. 

The genitalia show a peculiar development of the 



eighth segment collar, a central-dorsal, invaginated, 
sclerotized pocket flanked by a pair of irregular, elon- 
gate, flattened lobes and on dorsum of ovipositor a pair 
of shallow sclerotized pockets (fig. 661a). The genital 
opening also is unsclerotized, lacking the usual ventral 
shield. 

Type locality: Honda, Colombia (type in BM) . 

Food plant: Unknown. 

A male will be needed for certain generic placement. 

10. Genus Crocidomera Zeller 

Crocidomera Zeller, Isis von Oken, 1848, p. 865. — Ragonot, Mono- 
graph, pt. 1, p. 132, 1893. (Type of genus: Crocidomera 
turbidella Zeller.) 

Tongue well developed. Antenna pubescent; male 
with a short, blimt, spine from upper outer angle of 
basal segment of shaft; basal segment of male antenna 
swollen and broadly scaled. Labial palpus upcurved, 
reaching to vertex or slightly above it; third segment 
over half as long as second, acuminate. Maxillary 
palpus small, squamous. Forewing smooth; 11 veins; 
vein 2 from well before lower outer angle of cell; 3 from 
the angle ; 4 and 5 approximate for a short distance from 
cell; 6 from below upper angle of cell, very slightly bent 
towards base; 8-9 stalked for about half their lengths; 
10 from the cell approximate to the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from well 
before lower outer angle of cell; 3 from the angle; 4-5 
contiguous for about one-third their lengths beyond cell; 
7 and 8 closely approximate beyond cell; cell slightly 
less than half the length of wing; discocellular vein 
curved. Eighth abdominal segment of male with com- 
pound ventral and ventrolateral tufts. 

Male genitalia with apical process of gnathos an 
elongate hook, blunt and rounded or slightly notched at 
apex. Uncus broad, hoodlike, apical margin broadly 
rounded or broad and truncate. TranstUla complete, 
stout, arched, with a strongly forked central projection. 
Harpe with costa rather broadly sclerotized, but not 
produced at apex, clasper more or less developed, simple, 
erect. AneUus with broad, dorsoventrally flattened 
lateral arms. Aedeagus with longitudinal rows of 
thornlike spines towards apex; penis with sclerotized 
wrinklings, but otherwise unarmed. Vinculum stout, 
slightly constricted from middle to moderately broad 
terminal margin; sh'ghtly longer than broad. 

Female genitalia with signum developed as a small 
scobinate cup-shaped patch; ductus bursae moderately 
long (shorter than bursa), expanding gradually to the 
wide genital opening and with some strong sclerotized 
wrinklings before genital opening; lower margin of 
genital opening sclerotized, wrinkled and more or less 
finely scobinate; dense, fine scobinations on the dorsal 
membrane behind genital opening; ductus seminalis 
from bursa near junction of bursa and ductus bursae. 

A tropical American genus ranging as far north as our 
Texas border and probably into southern Florida ; easily 
distinguished by its genitalia and male antenna. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITtNAE 



33 



58. Crocidomera turbidella Zeller 
Figures 182, 664 

Crocidomera turbidella Zeller, Isis von Oken, 1848, p. 865. — 
Ragonot, Monograph, pt. 1, p. 132, 1893. 

Ground color of forewing variable, pale ocherous gray 
or white shaded with faintly reddish or reddish brown on 
outer area and in a narrowing shade towards base along 
inner margin; an indistinct blackish spot on costa 
beyond base; two other blackish dots marking the place 
of the obsolete antemedial line, one on costa, the other 
at top of cell; a dark patch on inner margin at what 
would be the inner margin of the antemedial line, on 
well marked specimens containing one or two minute 
black dots or dashes; sub terminal pale line faint but 
distinguishable, indicated chiefly by an inner border of 
black spots on the veins and similar, fainter dark streaks 
(or a confluent dark shade) bordering it outwardly; a 
row of black dots along termen, quite marked on fresh 
specimens; discal dots at end of cell small, black, more 
or less confluent, when separated the lower not appreci- 
ably enlarged; below these, on the fold of some speci- 
mens, a larger spot of reddish or brown scales. Hind 
wing transparent, opalescent white with a faint fuscous 
shading at apex and at the vein ends and a fine dark 
line along termen. Alar expanse, 20-25 mm. 

Male genitalia with apical process of gnathos bluntly 
rounded at apex; uncus sub triangulate, its apical mar- 
gin evenly rounded; central projection of transtilla 
V-shaped, the prongs divergent and rather slender; 
harpe with outer margin of cucullus evenly rounded. 
Female genitalia with sclerotization along lower margin 
of genital opening narrow and but slightly wrinkled. 

Type locality: "South America" (type in BM). 

Food plant: Unknown. 

Distribution: Cuba: Baracoa, Santiago (June, Oct.). 
Jamaica. Mexico: Jalapa. United States: Texas, 
San Benito (May, Oct.) . These localities from examples 
(in USNM) from which the foregoing description was 
drawn. 

I have omitted the Moschler reference cited by 
Ragonot for I suspect that the Puerto Rican specimens 
which he and Ragonot had and from which the Ragonot 
description was partly drawn are not turbidella but 
fissuralis without the peculiar longitudinal black streak 
on forewing characteristic of the type oi fissuralis. I 
have seen no specimens of Crocidomera from any South 
American locality except the Bolivian example men- 
tioned in the following discussion of fissuralis. I doubt 
that this could be Zeller's species. 

59. Crocidomera fissuralis (Walker) 
Figures 183, 665 

Nephopteryx fissuralis Walker, List, vol. 27, p. 58, 1863. 
Myelois (?) adonea Felder and Rogenhofer, Reise . . . Novara . . ., 

Lepidoptera, pi. 137, fig. 8, 1874. 
Crocidomera fissuralis (Walker) Moschler, Die Lepidopteren- 

Fauna von Portorico, p. 327, 1890. — Ragonot, Monograph, 

pt. 1, p. 133, 1893. 

A photograph of the female type oi fissuralis shows a 
specimen with a strongly contrasted, black longitudinal 



streak on forewing extending along lower median vein 
from base of wing to sub terminal line and giving off two 
short forks, one along vein 2, the other along vein 3. 
The figure of adonea, presumably also a female, shows a 
similar marking. I have seen nothing to match this 
peculiar pattern except one female from the Janse col- 
lection from the Provincia del Sara (Department of 
Santa Cruz), BoHvia. The genitalia of this specimen 
match those of fissuralis fairly well except for the 
sclerotization of the ductus bursae, which is more like 
that of stenopteryx. The specimen cannot be placed with 
certainty until a male from the type locality is associated 
with it. I doubt very much that the peculiar longi- 
tudinal streaking represents anything more than an 
aberrational or varietal character; for I have before me 
a series of three males and three females from Puerto 
Rico which lack the longitudinal streak, but are obvi- 
ously distinct specifically from what I have recognized 
as turbidella. Their female genitalia are identical in all 
details with those of the tj^pe oi fissuralis. On super- 
ficial characters they differ from turbedilla chiefly in 
having the lower discal spot at end of cell more pro- 
nounced and distinctly enlarged in comparison to the 
upper discal spot. Alar expanse, 25-30 mm. 

Male genitalia with terminal margin of uncus notched ; 
apical projection of gnathos slightly notched at apex; 
central projection of transtUla with its prongs converging 
towards their apices; harpe with cucullus trianguJate, 
its apex narrowly rounded ; aedeagus much stouter and 
its thornlike spines stronger and more numerous than 
those of turbidella or stenopteryx; terminal margin of 
vinculum nearly straight, terminal part of vinculum pro- 
portionally about twice as wide as that of either turbi- 
della or stenopteryx. Female genitalia with ventral 
plate at genital opening deeply wrinkled. 

Type localities: Santo Domingo [Dominican Re- 
public] (fissuralis, adonea, both in BM). 

Food plant: Unknown. 

Distribution: Dominican Republic. Puerto 
Rico: Aguirre Central (Apr.), Coamo Springs (Apr.), 
Culebralsl. (Feb.). 

60. Crocidomera stenopteryx (Dyar), new combination 

Figures 181, 663 

Dioryctria stenopteryx Dyar, Ins. Insc. Menstr., vol. 10, p. 16, 
1922. 

Forewing ocherous gray with blacldsh markings; a 
black dot at base of cell ; three black streaks beyond on 
lower median vein ; a couple of black dots on costa near 
base; a black discal spot at lower, outer angle of del; 
antemedial pale line indicated below median vein, pre- 
ceded by a dark patch on inner margin (somewhat 
tinted with reddish brown on female), margined out- 
wardly by an obscure, blackish line, beyond which 
on inner margin a somewhat diffused dark shade; vein 
lb more or less outlined in black scaling; subterminal 
line faint, bordered inwardly and outwardly by some 
blackish dots or streaklets on the veins ; a fine black line 
along termen (formed by the confluent terminal dots). 
Hind wing transparent, hyaline white with a faint 



34 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



grayish shade along costa and a fine fuscous line along 
termen. Alar expanse, 25-27 mm. 

Male genitaUa with terminal margin of uncus broad, 
very slightly convex; apical process of gnathos slightly 
notched at apex; prongs of central projection of trans- 
tiUa slightly convergent toward their apices ; harpe with 
terminal margin of cucullus oblique, apex bluntly 
poiated. Female genitalia with ductus bursae smoothly 
sclerotized between its sclerotized, wrinkled part and 
the schrotized and wrinkled margin of genital opening. 

Type locality: Tehuacan, Mexico (type in USNM). 

Food plant: Unknown. 

Ejiown only from the type locality. 

11. CuniLerta, new genus 

Type of genus: Nephopteryx subtinctella Kagonot. 

Tongue well developed. Antenna of male with basal 
segment elongate, cylindrical; shaft weakly serrate and 
pubescent (the cilia about as long as width of segments), 
basal segments swollen and incurved, forming a sinus 
containing a row of minute thornlike spines and overlaid 
with a spread of appressed scales; antenna of female 
simple and very weakly pubescent. Labial palpus up- 
turned, scarcely reaching vertex; third segment shorter 
than second, acuminate. MaxiUary palpus small, squa- 
mous. Forewing smooth; 11 veins; veiu 2 from before 
(but rather near) lower outer angle of cell; 3 from the 
angle, but little further from 2 at base than from 4; 4 
and 5 short stalked; 6 from below upper angle of cell, 
straight; 8 and 9 stalked for half their lengths; 10 from 
the cell, closely approximate to basal half of the stalk 
of 8-9 ; male without costal fold. Hind wing with vein 2 
from well before lower outer angle of cell; 3 from the 
angle, connate with 4-5; 4 and 5 stalked for approxi- 
mately half their lengths; 7 and 8 closely approximate 
beyond cell; cell slightly less than half the length of 
wing; discoceUular vein curved. Eighth abdominal seg- 
ment with 2 pairs of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos an 
elongate, stout, rather broad hook with blunt, notched 
apex. Uncus triangulate. Transtilla complete, stout, 
arched, produced at middle into a broad U-shaped pro- 
jection. Harpe simple. AneUus with rather broad, 
dorsoventrally flattened lateral arms. Aedeagus mod- 
erately slender with a single row of very minute serra- 
tions along one lateral margin towards apex, otherwise 
simple; penis with fine sclerotized wrinklings, otherwise 
unarmed. Viuculum stout, somewhat longer than 
broad. 

Female genitaha with signum developed as a small 
granulate cup-shaped patch; bursa small; ductus bursae 
considerably longer than bursa, unsclerotized except for 
a narrow sclerotization along lower margin of genital 
opening; ductus seminalis from ductus bursae near 
genital opening. 

The genus is close to both Hemiptilocera and Crocido- 
mera and shares some of the characters of each but is 
distinct from both in the definite stalking of veins 4-5 



of forewing and iu the attachment of the ductus semi- 
nalis of the female genitalia. 

Contains one North American species. 

61. Cuniberta subtinctella (Ragonot), new combination 
FiGUBES 170, 666 

Nephopieryx subtinctella Ragonot, N. Amer. Phycitidae, p. 7, 
1887; Monograph, pt. 1, p. 302, 1893. — Hulst, Phycitidae of 
N. Amer., p. 146, 1890.— McDunnough, Check list. No. 
6175, 1939. 

Forewing gray, more or less dusted with whitish on 
basal and median areas; antemedial line nearly vertical, 
out-angled at upper and lower margins of cell, bordered 
outwardly by a black line which is expanded and 
strongly accented on costa, bordered inwardly on lower 
margin by a reddish or reddish olivaceous patch; on 
some specimens a similar shade in fold beyond the ante- 
medial line; subterminal line sinuous, bordered in- 
wardly by a fine black line which, in most specimens, 
expands on costa into a conspicuous black spot or 
streak; on costa following the subterminal line a similar 
more or less expanded black spot; discal dots at end of 
cell usually confluent and forming a thin black lunide 
along the discoceUular vein; a thin black streaklet on 
vein 2. Hind wing pale smoky fuscous; veins scarcely 
darker; a faintly darkened line along terminal margin. 
Alar expanse, 22-26 mm. 

Genitalia as given for the genus; male with apex of 
uncus narrowly rounded; vinculum evenly tapering to 
rather broad terminal margin. 

Type locality: California (type in Paris Mus.). 

Food plant: Unknown. 

Distribution: United States: Utah, Park City 
(June), Provo (July, Aug.); California, Shasta Ketreat 
(Siskiyou County, Aug.). Canada: British Columbia, 
Kaslo (June). 

12. Heras, new genus 

Type op genus: Heras disjunctus, new species. 

Tongue well developed. Antenna of male with first 
segment rather long, cylindrical; shaft with a sinus and 
heavy scale tuft at base, otherwise weakly pubescent. 
Labial palpus upcurved, reaching above vertex; dorso- 
ventraUy flattened; third segment somewhat shorter 
than second. MaxUlary palpus squamous. Forewing 
smooth; 11 veins; vein 2 from weU before lower outer 
angle of cell; 3 from the angle; 4 and 5 connate or very 
shortly stalked, shortly separated from 3 at base; 6 
from slightly below upper angle of ceU, very slightly 
bent towards base; 10 from the cell, closely approximate 
to the stalk of 8-9; on male, a long narrow costal fold 
and, on upper surface of wing, a fovea (depressed 
pocket) in cell slightly beyond base. Hind wing with 
vein 2 from well before lower outer angle of ceU ; 3 from 
the angle, connate with 4-5; 4 and 5 stalked for half 
their lengths; 7 and 8 closely approximate beyond cell; 
cell about half the length of wing; discoceUular vein 
curved. Eighth abdominal segment of male with ster- 
nal plate developed as a narrow sclerotized pocket at 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITESTAE 



35 



its center; in the intersegmental area two pairs of mem- 
branous eversable lobes (not haired), one long ventro- 
lateral pair and one shorter dorsolateral pair. 

Male genitalia with apical process of gnathos a nar- 
row, somewhat flattened hook with slightly forked apex. 
Uncus triangulate. Transtilla complete, stout; a rather 
short bridge with long widely spaced lateral arms pro- 
jecting backward; and a similarly spaced, shorter pair 
of arms projecting forward and articulating with the 
anellus. Harpe with a strong, long hair tuft from outer 
surface of base of sacculus, otherwise simple. Anellus 
a narrow plate with very long, slender, strongly sclero- 
tized, lateral arms. Aedeagus rather long and slender, 
sclerotized only on dorsal half; penis with some weak 
wrinklings and minute scobinations near apex, other- 
wise unarmed. Vinculum approximately triangulate, 
but slightly longer than its greatest width; its central, 
ventral area unsclerotized. 

I very much dislike to erect a new genus on a single 
male; but the genitalic and secondary male characters 
of this example are so striking and its distinctness from 
any known genus is so obvious it seems best to give it 
a name and separate designation. The genus is ap- 
parently closest to Hemiptilocera. 

62. Heras disjunctus, new species 

Figure 184 

Forewing rosy fuscous with costal area beyond ante- 
medial line broadly clay colored (pale ocherous); the 
rose shade predominant on upper part of wing, the fus- 
cous shade more accented in lower fold and along inner 
margin; antemedial hne weak, indicated chiefly by a 
distinct but small whitish ocherous spot near inner mar- 
gin; subterminal more distinct, whitish ocherous, ter- 
minating at inner margin in another pale spot similar 
to the one on antemedial Hne, inner dark margin of 
subterminal line narrow and very faint; discal dots at 
end of cell confluent, blackish; terminal dots confluent, 
some faint blackish streaking on the veins before and 
beyond the subterminal line. Hind wing pale smoky 
fuscous ; darker along the veins and towards outer mar- 
gin. Alar expanse, 22 mm. 

Male genitalia with the long posteriorily projecting 
arms of transtilla terminating in flattened lobes; apex 
of uncus narrowly rounded. Vinculum tapering to 
evenly rounded terminal margin. Female unknown. 

Type locality. Don Amo, Colombia (200 ft., July) 
(type in Janse Coll.). 

Food plant. Unknown. 

Described from unique male type. Superficially (in 
maculation and color) it strongly resembles Hyalospila 
stictoneurella Ragonot. 

13. Adanarsa, new genus 

Type of genus: Rhodophaea intransitella Dyar. 

Tongue weU developed. Antenna simple and pubes- 
cent in both sexes. Labial palpus upturned, reaching 
to vertex; slightly flattened laterally; thu-d segment 
about half the length of second, blunted and slightly 



broadened (ventrally) by scales at apex. Maxillary 
palpus small, squamous. Forewing smooth; 11 veins; 
vein 2 from well before lower outer angle of cell; 3 from 
the angle; 4 and 5 shortly stalked, separated at base 
from 3 ; 6 from below upper angle of cell, straight; 8 and 
9 stalked for half their lengths; 10 from the cell, closely 
approximate to the stalk of 8-9 for a short distance be- 
yond cell; male without costal fold. Hind wing with 
vein 2 from well before lower outer angle of cell ; 3 from 
the angle, approximate to 4-5 at base; 4 and 5 stalked 
for half or nearly half their lengths; 7 and 8 weakly 
anastomsed for a short distance beyond cell; cell half 
the length of wing; discocellular vein cm-ved. Eighth 
abdominal segment of male with a small pair of ventro- 
lateral hair tufts. 

Male genitalia with apical process of gnathos a slen- 
der, rather long hook with slightly forked apex. Uncus 
semitriangulate. Transtilla complete, strongly sclero- 
tized and arched, supporting at its center a rather nar- 
row, smooth, curved crosspiece. Harpe with strongly 
sclerotized, erect clasper, otherwise simple. Anellus 
with short, broad, dorsoventrally flattened lateral arms. 
Aedeagus with a row of very fine serrations along one 
lateral edge towards apex; penis armed with a single, 
slender, sinuate, cornutus. Vincidum stout, about as 
long as greatest width; terminal margin broad. 

Female genitalia with cornutus developed as a single, 
short, stout, hooked thorn ; ductus bursae much shorter 
than bursa, broad, flattened and with a broad transverse 
sclerotized band across it at junction of ductus and 
bursa ; genital opening weakly and narrowly sclerotized 
along its lower margin and with a naiTow, transverse 
sclerotized band in the membrane just behind the open- 
ing; ductus seminalis from bursa near its junction with 
ductus biu-sae. 

A distinct genus distinguished from related genera 
with complete transtilla by the strongly sclerotized 
clasper and the slight but definite anastomoning of 
veins 7-8 of hind wing. The amount of anastomosis 
varies in different specimens of the type species but is 
always present and always for somewhat less than half 
the length of the veins. 

63. Adanarsa intransitella (Dyar), new combination 

Figures 185, 667 

Rhodophaea intransitella Dyar, Proc. Ent. Soc. Washington, vol. 
7, p. 33, 1905.— McDunnough, Check list, No. 6075, 1939. 

Forewing pale ash gray with a fine scattered dusting 
of black scales and a very faint clouding of ocherous 
fuscous above iimer margin between the transverse 
lines ; antemedial line obsolete, indicated only by small 
black spot on costa and a larger black spot on inner 
margin at what would be the inner margin of the trans- 
verse line ; outer line faint, indicated chiefly by border- 
ing black dashes on costa, a faint blackish line along 
its outer border and a few inwardly bordering black 
dots ; lower discal spot black, followed outwardly by an 
obsciu-e dark streak ; a row of black dots along termen. 
Hind wing whitish, subpellucid; more or less shaded 



36 



TJNITED STATES NATIONAL MUSEUM BULLETIN 207 



with fuscous towards apex and with a dark line along 
termen. Alar expanse, 16-19 mm. 

Male genitalia with apex of uncus evenly but rather 
narrowly rounded; clasper broadly flaring at apex; vin- 
culimi no longer than broad ; terminal margin abruptly 
truncate, nearly straight (very slightly concave at 
middle) ; aedeagus stout. Female genitalia as given for 
the genus. 

Type locality: Albuquerque, N. Mex. (type in 
USNM). 

Food plant: Unknown. 

Distribution: New Mexico, Albuquerque (July); 
ArizoTM, Christmas, Kiugman (Oct.), Phoenix (Mar.). 

14. Birinus, new genus 

Type of genus: Birinus russeolus, new species. 

Tongue well developed. Antenna simple and pubes- 
cent. Labial palpus upturned, slender, barely reaching 
to vertex; third segment shorter than second, acuminate. 
Maxillary palpus minute, filiform. Forewing smooth; 
11 veius; vein 2 from before (but moderately near) 
lower outer angle of cell; 3 from the angle, closer to 4-5 
than to 2 ; 4 and 5 very shortly stalked ; 6 from very near 
to upper angle of cell, bent towards base, approximate 
at base to stalk of 8-9 ; 8 and 9 long stalked (for more 
than two-thirds of their lengths) ; 10 from the cell, closely 
approximate to stalk of 8-9 for a considerable distance 
beyond cell. Male without costal fold. Hind wing with 
vein 2 from well before lower outer angle of cell; 3 from 
the angle, connate with the stalk of 4-5 ; 4 and 5 stalked 
for a little over half their lengths; 7 and 8 closely approx- 
imate for a short distance beyond cell; cell about half 
the length of wing; discocellular vein curved. Eighth 
abdominal segment of male simple. 

Male genitaHa with apical projection of gnathos long, 
nearly straight, needlelike. Uncus spatulate, broadest 
at apical margin. TranstUla complete, a very slightly 
curved band (nearly straight on posterior margin), 
broad at its bases, narrow at middle. Harpe with sac- 
culus strongly sclerotized throughout its length and with 
apex produced as a short spur at lower outer angle of 
harpe; costa very short, sclerotized; bent upward at a 
sharp right angle a short distance from base, not pro- 
duced; cucullus forming more than half of the harpe 
area. Sclerotized part of anellus greatly reduced. Aede- 
agus long, moderately stout, smooth; penis armed with 
two narrow, blade like cornuti and numerous fine granu- 
lations. Vinculum triangulate, sclerotized only along 
margins. 

The foregoing description is incomplete, as the female 
is unknown; but the new genus seems to be justified by 
the male genitalia, which are unlike anything else in the 
American fauna. 

64. BirinuB russeolus, new species 

Figures 8, 186 

Forewing reddish brown, the rust-red shading a little 
more pronounced in outer costal and marginal areas and 
along lower vein of cell; a faint pale, clay-colored blotch 



in outer median area between vein 8 and the lower fold, 
enclosing in its center a small patch of blackish brown 
scales and at its inner margin bordered by a similar 
blackish smudge formed by the confluent discal spots; 
on the fold below and just before lower outer angle of 
cell a somewhat larger blackish brown patch preceded 
by a small clay-colored spot; a thin line of dark scales 
along the remainder of the fold to base of wing; the usual 
antemedial and subterminal lines obsolete; terminal dots 
very faint. Hind wing pale smoky fuscous, the veins 
brown and the cell filled with brown scaling. On the 
underside of fore and hind wings dark brown sex-scaling 
(a male character) covers the upper wing area (above 
lower margin of cell) from base to somewhat beyond the 
outer margin of the cell. Forefemora of male with a 
strong, clay-colored, hair tuft from upper basal angle 
(also a male character). Alar expanse, 22 mm. 

Male genitalia as given for the genus. Female un- 
known. 

Type locality: Tumatumari, Potaro River, British 
Guiana (t3^e in Cornell). 

Food plant: Unknown. 

Described from unique male type collected by W. T. 
M. Forbes, June 20, 1927 (Cornell lot 760 sub. 114). 

15. Genus Bertelia Barnes and McDunnough 

Bertelia Barnes and McDunnough, Contributions, vol. 2, p. 140, 
1913. (Type of genus: BerieZia ^nseHa Barnes and McDun- 
nough.) 

Tongue well developed. Antenna of male with a 
strong posterior scale tuft on basal segment; shaft with 
a deep incurvation (sinus) at base, the sinus containing 
a few minute spinelike thorns but no scale tuft, shaft 
beyond sinus strongly unipectinate; antenna of female 
simple and pubescent. Labial palpus upturned on male, 
reaching a trifle higher than vertex; obUque on female; 
third segment on male slender, acuminate, about half 
the length of second, on female shorter and somewhat 
expanded with scales at apex. Maxillary palpus squa- 
mous. Forewing smooth; 11 veins; vein 2 from well 
before lower outer angle of cell, 3 from the angle; 4 and 
5 shortly stalked; 6 from below upper angle of cell, 
straight; 8 and 9 stalked for a trifle more than half their 
lengths; 10 from the cell, closely approximate to the 
stalk of 8-9 ; male without costal fold. Hind wing with 
vein 2 from well before lower outer angle of cell; 3 from 
the angle; 4 and 5 contiguous, shortly anastomosed or 
stalked, usually stalked for less than half their lengths ; 
7 and 8 closely approxunate for haK their lengths beyond 
cell; cell about half the length of wing; discocellular vein 
curved. Eighth abdominal segment of male with a pair 
of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos de- 
veloped as a stout hook, slightly notched at apex. 
Uncus triangulate. TranstUla incomplete, but with 
the elements long and stout, their apices broadly and 
irregularly developed and hooked. Harpe simple. 
Anellus U-shaped, its lateral arms dorsolaterally flat- 
tened. Aedeagus with a short row of minute scobina- 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITDSrAE 



37 



tions along one lateral edge towards apex, moderately 
stout ; penis with a few sclerotized wrinklings, otherwise 
unarmed. Vinculum stout, shghtly longer than great- 
est width; terminal margin broad. 

Female genitalia with bursa large and elongate, 
signum present, developed as a small, scobinate, cup- 
shaped depression, ductus bursae very short; genital 
opening with a narrow, short, sclerotized plate on its 
lower margin and a large semicircidar, sclerotized and 
scobinate dorsal plate in the membrane behind the 
opening; a pair of ventral scobinate plates in the inter- 
segmental area between eighth segment coUar and 
ovipositor; ductus seminalis from lobe of bursa near 
ilB junction with ductus bursae. 

A distinct genus containing one described North 
American species. The venation of the hind wing is 
individually variable in the amount of stalking or 
anastomosis of veins 4 and 5. 

65. Bertelia grieella Barnes and McDunnough 

Figures 187, 669 

Bertelia grisella Barnes and McDunnough, Contributions, vol. 2, 
p. 140, 1913.— McDunnough, Check List, No. 6140, 1939. 

Forewing pale ashy gray dusted with fuscous; faint 
interrupted black streaking on upper and lower veins 
of cell, the lower fold and some of the veins beyond 
cell; antemedial line obscure, indicated chiefly by its 
narrow blackish outer border (out-angled from costa) 
and by a whitish incurved line between cell and inner 
margin, preceding which is an obscure dark shading; 
subterminal line nearly obsolete, followed on costa by 
a blackish shade; discal dots obsolete on many speci- 
mens, occasionally indicated by a small blackish dot 
at lower outer angle of cell. Hind wing semihyaline 
white with a faint ocherous tint; veins not appreciably 
darkened ; a faint fuscous line along outer margin. Alar 
expanse, 24-30 mm. 

Male genitalia with apex of uncus bluntly and nar- 
rowly rounded; terminal margin of vinculum slightly 
angled; lateral arms of anellus moderately long and 
broad. Female genitalia as given for the genus. 

Type locality: Redington, Ariz, (type in USNM). 

Food plant: Unknown. 

Disthibution: Arizona, Redington, Santa Catalina 
Mts. (Sept.). 

16. Genus Hypargyria Ragonot 

Hypargtjria Ragonot, Nouv. Gen., p. 9, 1888, Monograph, pt. 1, 
p. 122, 1893. — Janse, Journ. Ent. Soc. South Africa, vol. 4, 
p. 149, 1941. (Type of genus: Hypargyria metalliferella 
Ragonot; India.) 

Tongue weU developed. Antenna pubescent, basal 
segment on male elongate with a short spur of scales 
from its upper inner angle (giving the base of antenna 
much the same appearance as that of the undenuded 
two first segments of the male antenna of Mildrixia, 
fig. 169f); male shaft with a deep sinus towards base 
containing a longitudinal row of very minute teeth but 
no scale tuft; antenna of female simple. Labial palpus 



uptm^ned, reaching a little above vertex; third segment 
nearly as long as second, acuminate. Maxillary palpus 
squamous. Forewing with transverse, antemedial ridge 
of raised scales; 11 veins; vein 2 from before but near 
lower outer angle of cell ; 3 from the angle, at base about 
equidistant from 4-5; 4 and 5 connate or very shortly 
stalked; 6 from upper angle of cell, connate with stalk 
of 8-9, straight or but slightly bent towards base; 10 
from the cell, approximate to the stalk of 8-9; male 
with a notch in costa very close to base and on under- 
side at base of costa a small knot of modified scales and 
(projecting into the costal notch) a very short brush of 
stiff hairs. Hind wing with vein 2 from well before 
lower outer angle of cell ; 3 from the angle, connate with 
stalk of 4-5 ; 4 and 5 stalked for slightly less than half 
their lengths ; 7 and 8 closely approximate for less than 
half their lengths beyond cell; cell slightly more than 
one-third the length of wing; discocellular vein curved. 
Eighth abdominal segment of male with compound 
ventral scale tufts. 

Male genitalia with gnathos weak, lacking a central 
projection, the lateral arms meeting at the base of a 
rudimentary subanal plate. Uncus triangulate. Trans- 
tilla incomplete, but with the elements long and stout, 
their apices broadly developed. Harpe with costa 
broadly and very strongly sclerotized and stoutly pro- 
jecting at apex; a fine moderately long hair tuft from a 
sclerotized disk attached to base of sacculus. Anellus a 
broad, deep plate (bearing short, knoblike, stoutly 
spined, lateral projections on the American species). 
Aedeagus smooth; penis armed with two or more short, 
rather stout, straight spines, a deeply wrinkled, sclero- 
tized band, and a cluster of fine moderately long spines. 
Vinculum very stout, considerably larger than uncus 
and tegumen combined, longer than broad. 

Female genitalia with signum developed as a small, 
round, scobinate, cup-shaped plate; ductus bursae 
shorter than bm-sa, a broad, strongly sclerotized band 
at the junction of ductus and bursa and a narrower 
sclerotized band at genital opening; behind genital 
opening a conspicuous pair of strongly sclerotized, gran- 
ulate, pocket like lobes; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

Presumably an Old World genus of tropical and prob- 
ably African origin; possessing some structural char- 
acters of Acrobasis, Mildrixia, and Bertelia but amply 
distinct from any of them. It contains two American 
species. 

66. Hypargyria definitella (Zeller) 
Figures 188, 668 

Myelois definitella Zeller. Horae Soc. Ent. Rossicae, vol. 16, 

p. 205, 1881. 
Hypargyria definitella (Zeller) Ragonot, Monograph, pt. 1, p. 124, 

1893. 

Forewing purplish ocherous to purplish brown, most 
of basal area and costal half of median area white 
sparsely dusted with red scales, the red dusting most 
abundant along midcosta; a small ocherous patch on 
inner margin near base; antemedial line evenly curved. 



38 



UNITED STATES NATIONAL MUSEUM BULLETIN" 207 



ocherous, bordered outwardly by a red or purplish line 
continued from a rather pronounced costal dash, and 
inwardly by the vertical scale ridge, the latter red, 
reddish ocherous or purple with some admixtm-e of 
blackish scaling; subterminal line very faint with faint 
purplish borders; discal spots at end of cell separated, 
blackish ; black terminal dots faint. Hind wing hyaline 
white with a faint smoky tint on some specimens; the 
veins darkened (brown) and a narrow brown line along 
termen. Undersides of male fore and hind wings in 
the area between vein 2 and costa and from near end of 
cell outward covered with shining sUvery scales; also 
on forewing a short black median streak from base, 
more or less extended into ceU along lower edge of upper 
vein of cell and on hind wing a similar black streak 
on upper vein of cell; these black sex-scalings not con- 
stant and altogether absent from occasional males. 
Alar expanse, 16-20 mm. 

Genitalia as given for the genus. 

Type locality: Honda, Colombia (type in BM). 

Food plant: Unknown. 

Distkibution: Puerto Eigo: Puerto Real (Vieques 
Isl., Apr.), San Germdn. Virgin Islands: KingshiQ 
(St. Croix, June, Oct.). Colombia: Honda, Valparaiso. 
Brazil: Castro, Santa Catarina. 

The males of this species can be distinguished at once 
from any other American phycitid by the shining 
silvery scaling on the imdersides of the wings; a char- 
acter, however, shared by the Old World type of the 
genus. The Old World metalliferllae exhibits a number 
of slight but consistent male genitalic diflferences: The 
heavier and more abundant spining on the penis, a 
different shape to the apical projection of costa of harpe, 
a different shape to the apices of the elements of trans- 
tiUa (not developed into paired hooks as in the Ameri- 
can species), and an anellus without spined lateral pro- 
jections. Such differences are certainly specific but no 
more. Through the com-tesy of the British Museum I 
have been able to examine males of metallifereUa from 
Pusa in India and Nyasaland in Africa. There were 
no differences of any kind between them. 

67. Hypargyria slossonella (Hiilst), new combination 

Salabria slossonella Hulst, Canadian Ent., vol. 32, p. 170, 1900. 

Acrobasis tenuella Barnes and McDunnough, Contributions, vol. 
2, p. 181, 1913. 

Acrobasis slossonella (Hulst) Barnes and McDunnough, Contri- 
butions, vol. 3, p. 195, 1916. — McDunnough, Check list, 
No. 6108, 1939. 

Not distinguishable from dejinitella except that the 
males lack entirely the silvery scaling on the under- 
sides of the fore and hind wings. 

I suspect that it is only a variety or race of dejinitella; 
but until more material is available and something is 
known of their life histories the two forms will have to 
be kept as separate species. The genitalia of slossonella 
exhibit no differences of any specific significance from 
those of dejinitella. 

Type localities: Miami, Fla. (sZossoneZto, in AMNH, 
ex Rutgers) ; Everglades, Fla. {tenuella, in tJSNM) . 



Food plant: Unknown. 

Distribution: United States: Florida, Everglades 
(Apr.), Fort Myers (Apr.), Miami ("February-March"). 
Mexico: Oaxaca. 

17. Chararica, new genus 

Type of genus: Myelois annuliferella Dyar. 

Tongue well developed. Antenna simple and pubes- 
cent on both sexes. Labial palpus upturned, reaching 
to vertex, slender; third segment about as long as sec- 
ond, acuminate. Maxillary palpus squamous. Fore- 
wing smooth; 11 veins; vein 2 from before, but near, 
lower outer angle of cell; 3 from the angle; 4 and 5 
separated at base, distance separating them slightly 
less than that separating 3 and 4; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for slightly less 
than half their lengths; 10 from the cell, approximate 
to the stalk of 8-9 for a short distance; male without 
costal fold. Hind wing with vein 2 from well before 
lower outer angle of cell; 3 from the angle, connate 
with or very closely approximate to 4 at base; 4 and 5 
shortly stalked; 7 and 8 approximate for a short dis- 
tance beyond cell; cell about half the length of wing; 
discoceUular vein curved. Eighth abdominal segment 
of male with a pair of ventrolateral hair tufts. 

Male with gnathos weak, lacking a central projec- 
tion, the lateral arms articulating with a thinly sclero- 
tized subanal plate. Uncus triangulate, sharply taper- 
ing to a blunt point. Transtilla complete, with a cen- 
tral sclerotized apron connecting transtUla and uncus, 
and with a pair of widely spaced lateral arms each 
bearing at its apex a clutter of slender spinelike setae. 
Harpe with costa slightly concaved, strongly sclero- 
tized, not produced at apex; cucuUus large, forming 
about two-thirds of the harpe, outer margin broadly 
rounded. Anellus with stubby, stout, convergently 
directed, lateral projections. Aedeagus with lateral 
margins serrated toward apex; penis with a few weakly 
sclerotized wrinMings or granulations, otherwise un- 
armed. Vinculxim stout, nearly square in outhne. 

Female genitalia with signum developed as a small, 
finely granulate-scobinate, cup-shaped disk; bursa large; 
ductus bursae, broad, short, less than half as long 
as bursa, weakly sclerotized and transversely wrinkled 
towards genital opening; genital opening broad, with 
strongly sclerotized transverse plate along lower margin 
and some weak scbbinations on the membrane above 
and behind the opening; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

This genus is erected for three North American spe- 
cies now listed under Rhodophaea, but differing from 
that genus in both venational and genitalic characters. 
It is easily recognized by its male genitalia. The spe- 
cies also have a pattern character which aids in iden- 
tification: the usual discal dots on forewing at end of 
cell are replaced by a small obicular marking. This is 
weak on bicolorella but present and distinguishable on 
most specimens. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



39 



68. Chararica annuliferella (Dyar), new combination 
Figures 189, 670 

Myelois annuliferella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 33, 1905. 

Rhodophaea annuliferella (Dyar) Barnes and McDunnough, 
Check list of the Lepidoptera of Boreal America, No. 5516, 
1917.— McDunnough, Check list, No. 6074, 1939. 

Forewing dark gray with a faint, pale ocherous shade 
along inner margin; antemedial line well out on wing, 
outwardly arched in cell to slightly beyond middle of 
wing, intent from lower fold to inner margin, white, 
bordered outwardly by a narrow black line; basal area 
with veins black and faint intervenous whitish dusting; 
subterming Hne rather near terminal margin, slightly 
sinuous, fifne, white with a narrow, inner, black border; 
some faint white dusting in median area, especially 
along costa; usual discal dots at end of cell replaced 
by small black obicular mark with a whitish center. 
Hind wing hyaline white; veins not appreciably dark- 
ened; a faint, small fuscous shade at apex and a very 
faint, dark line along termen. Alar expanse, 19-22 mm. 

Genitalia of male with lateral arms of transtilla much 
reduced, widely spaced, their spinelike hair tufts long. 
Vinculum with terminal margin decidedly incurved at 
middle. Female genitalia with transverse sclerotized 
band on lower margin of genital opening narrow. 

Type locality: Gallinas Canyon, N. Mex. (type in 
USNM). 

Food plant: Unknown. 

Distribution: New Mexico, Gallinas Canyon (July) ; 
Arizona, Colorado Desert (Yuma County), "So. Ariz.," 
Kingman (Oct.), Yavapai County. 

69. Chararica hystriculella (Hulst), new combination 
FiGUBEs 190, 671 

Acrobasis hystriculella Hulst, Ent. Amer., vol. 3, p. 135, 1887. 
Rhodophaea hystriculella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 

114, 1889; Monograph, pt. 1, p. 73, 1893.— McDunnough, 

Check list, No. 6073, 1939. 
Myelois hystriculella (Hulst) Hulst, Phycitidae of N. Amer., p. 

119, 1890. 

Forewing whitish gray with extreme base dark fuscous 
and a similar fuscous shade over the outer area from 
subterminal line on costa obliquely to inner margin near 
antemedial line, and outward to termen; some black 
streaking on upper and lower veins of cell and vein lb 
before the antemedial line; antemedial line well out 
towards middle of wing, twice angled outwardly, indi- 
cated chiefly by a fine black outwardly bordering line; 
subterminal line faint, with narrow dark inner and outer 
borders, beginning as blackish dashes on costa; obicular 
spot at end of cell conspicuous, black with a narrow 
whitish center; terminal dots black, more or less con- 
fluent. Hind wing hyaline white with a faint fuscous 
shade at apex and a narrow dark line along termen, 
these dark shadings very slight on the males, more ex- 



tended and stronger on some females ; veins occasionally 
darkened on females, not darkened on males. Alar 
expanse, 17-23 mm. 

Male genitalia with uncus narrowly triangulate; lat- 
eral arms of transtilla rather long and their terminal 
hair tufts correspondingly shortened, not so widely 
spaced as those of annuliferella; terminal margin of 
vinculum very slightly concave, nearly straight. Fe- 
male genitalia with transverse sclerotized band on lower 
margin of genital opening broad (at least twice as wide 
as that of annuliferella) . 

Type locality: Texas (type, 9, in AMNH, ex Rut- 
gers). 

Food plant: Unknown. 

Distribution: Texas, Blanco County (Sept.), Browns- 
ville (May, June), Chisos Mts. (June), Cotula (Mar., 
Apr., May), Devils River (May), Kenedy (May), Kerr- 
ville (Aug.), Nueces River (Za valla County, Apr.), 
Sabinal (Mar.), San Antonio (June, July), San Benito 
(Mar.), San Diego (Apr., May, June); Florida, Coconut 
Grove, Miami. 

70. Chararica bicolorella (Barnes and McDunnough), new 
combination 

Rhodophaea bicolorella Barnes and McDunnough, Canadian Ent., 
vol. 49, p. 404, 1917.— McDunnough, Check hst, No. 607(. 
1939. 

Forewing with costal half of basal area black, streaked 
and peppered with white, giving this area of the wing 
a slate-colored appearance to the naked eye; outer area 
of wing from subterminal line to outer margin and costa 
to lower fold a similarly dark shade; remainder of wing 
ocherous, shading outwardly to tawney or ruddy ocher- 
ous; antemedial line obsolete except along outer margin 
of blackish basal patch; subterminal line faint, narrowly 
and weakly bordered inwardly and outwardly by black- 
ish lines; obicular mark at end of cell very faint but 
distinguishable on most specimens; terminal dots con- 
fluent. Hind wing hyaline white with a very faint 
ocherous line on outer margin for a short distance from 
apex. Alar expanse, 20-24 mm. 

Genitalia essentially like those of hystriculella. 

Type locality: Christmas, Gila County, Ariz, (type 
in USNM). 

Food plant: Unknown. 

Distribution: Arizona, Christmas, Mohave County 
(May, June, July, Aug., Sept.), Redington; Nevada, 
Clark County (Apr.), "So. Nevada" (July); California, 
San Bernadino County (Apr.) . 

A striking species easily distinguished by its color pat- 
tern, but not structurally different from hystriculella. 
The original type series consists of two males and two 
females, not four males as stated by the authors. 

Both bicolorella and hystriculella have a strong hair 
tuft on the metathorax of the male adjacent to the base 
of the leg. This character is lacking in annuliferella. 



40 



■UNITED STATES NATIONAL MUSEUM BULLETIN 207 



Genera 18-21: Myelopsis to Ectomyelois 

[Venational division C. Forewing with 11 veins; 4 and 5 stalked 
for half their lengths or less. Hind wing with veins 7 and 8 anas- 
tomosed for over half their lengths beyond cell (the free element 
of 8 shorter than the anastomosed stalk of 7-8). TranstiUa of 
male genitalia complete.] 

18. Myelopsis, new genus 

Type of genus: Myelois coniella Ragonot. 

Tongue well developed. Antenna simple, pubescent. 
Labial palpus upturned, reaching slightly above ver- 
tex; second segment somewhat roughly scaled in front; 
third segment slightly shorter than second, acuminate. 
Maxillary palpus filiform. Forewing smooth; 11 veins, 
vein 2 from before but near lower outer angle of cell; 
3 from the angle; 4 and 5 shortly stalked, the stalk at 
base separated from 3 for a distance but slightly less 
than that between 3 and 2 ; 6 from below upper angle 
of ceU, straight; 10 from the cell separated from 8-9 
at base, and more or less divergent beyond; male with 
out costal fold. Hind wing with vein 2 from well be- 
fore outer angle of cell; 3 from the angle, connate with 
the stalk of 4-5 ; 4 and 5 stalked for slightly less than 
half their lengths; 7 and 8 strongly anatomosed beyond 
cell, the free element of 8 short; cell half the length of 
wing; discocellular vein curved. Eighth abdominal 
segment of male simple. 

Male genitalia with apical process of gnathos U- 
shaped (consisting of a pair of widely spaced, short 
arms). Uncus stout, more or less triangulate, apex 
rather narrowly rounded. TranstiUa complete, but 
weakly sclerotized. Harpe simple; costa strongly scler- 
otized and projecting at apex (except in subtetriceUa) . 
Anellus U-shaped, narrowly sclerotized throughout. 
Aedeagus smooth; penis with sclerotized wrinklings. 
Vinculmn triangulate, tapering, longer than greatest 
width. 

Female genitalia with or without signum, latter when 
present weak. Ductus bursae membranous. Genital 
opening simple. Ductus seminalis from middle or 
towards terminal end of bursa. 

This genus comprises several North American species 
that have been referred to Myelois Hiibner. The latter 
a heterogenous assemblage of species, very few of which 
are actually congeneric with the type of genus (medii- 
lallis Hiibner synonym of crihrella Hiibner). The lat- 
ter has veins 7 and 8 of hind wing very shortly and 
weakly anastomosed beyond the cell, the free element 
of vein 8 correspondingly long and the cell itself over 
half the length of the wing. It belongs properly in 
om: venational division B. None of the American and 
very few of the Old World species that have been as- 
signed to it are properly referrable to Myelois. The 
European tetricella SchiffermueUer belongs in Myelopsis. 

71. Myelopsis coniella (Ragonot), new combination 
FiGTJBES 191, 673 

Myelois coniella Ragonot, N. Amer. Phycitidae, p. 3, 1887; 
Monograph, p. 1, p. 53, 1893. — Hulst, Phycitidae of N. 



Amer., p. 118, 1890.— McDunnough, Check list, No. 6071, 
1939. 
Rampylla nefas Dyar, Ins. Insc. Menstr., col. 10, p. 172, 1922 
(new synonymy). 

Forewing pale ash gray (on Utah and Nevada spec- 
imens) to blackish gray; on darker specimens the basal 
and terminal areas are contrastingly paler than the 
area between the transverse lines; at extreme base on 
inner margin an obscure ocherous patch (not distin- 
guishable on worn or faded specimens) ; antemedial line 
oblique, white with a broad, black outer border; sub- 
terminal line sinuate, more or less contrastingly pale 
and inwardly bordered by a blackish line or varying 
intensity; discal dots at end of cell usually distinct, 
separated, black; terminal dots obscure, when distin- 
guishable, more or less confluent. Hind wing hyaline 
white to pale smoky fuscous with a fine dark line along 
termen. Alar expanse, 16-22 mm. 

Male genitaUa with transtilla slender, arched and 
very weakly sclerotized at the central attachment of 
its elements. Harpe with costa narrowly sclerotized 
and projecting a trifle beyond the apex of the cucuUus. 
Female genitalia without signum; bursa membranous; 
ductus bursae with some minute scobinations near its 
junction with bursa, otherwise smooth. 

Type localities: Nevada {coniella, in Paris Mus.) ; 
Mexico City, Mexico {nejas, in USNM). 

Food plant: Unknown. 

Distribution: United states: Nevada, Montgom- 
ery Pass (Mineral County, Sept.), Utah, Dividend 
(Aug.), Eureka (June, Aug.), Provo (July, Aug., 
Sept.), Stockton (Sept.), Trout Creek (Ibapah Mts., 
Sept.); Colorado, Glenwood Springs (Aug.); Arizona, 
Pinal Mts. (July), no definite locality (Aug., Sept.); 
New Mexico, Gallinas Canyon; Texas, Burnet County, 
(Mar.), Kerrville (Mar.); Nebraska, Sioux County, 
(July) ; Michigan, Dickinson County ; Maine, Bar Har- 
bor (Aug.), Moimt Desert (Aug.). Canada: British 
Columbia, Kaslo (July, Aug.) ; Manitoba, Aweme (Aug.) ; 
Ontario, Ottawa (July, Aug.). Mexico: Mexico City 
(Sept.), Tehuacdn (Sept.). 

A variable species in color but with remarkably con- 
stant genitalia. Dyar's nefas has much darker fore- 
wings than specimens from Utah or Nevada but no 
darker than some specimens from Arizona and New 
Mexico. 

72. Myelopsis immundella (Hulst), new conxbination 

Myelois immundella Hulst, Phycitidae of N. Amer., p. 117, 
1890.— Ragonot, Monograph, pt. 1, p. 49, 1893.— McDun- 
nough, Check list, No. 6068, 1939. 

The type is without abdomen. In size, wing shape, 
pattern, and general coloration it is like the following 
species (subtetriceUa) except that the antemedial and 
subterminal lines of forewing are more whitish and dis- 
tinct and their dark borders (especially the outer border 
of the antemedial line) blackish and more strongly con- 
trasted against the dark groimd color of the wing. The 
discal dots are also more strongly contrasted. 

The name may represent only a color form of subtetri- 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



41 



cella, but until other specimens matching the type are 
found and their genitalia studied this cannot be deter- 
mined one way or the other. 

Type locality: Texas (type in AMNH, ex Rutgers). 

Food plant: Unknown. 

73. Myelopsis subtetricella (Ragonot), new combination 
Figures 192, 672 

Myelois subtetricella Ragonot, Ent. Amer., vol. 5, p. 113, 1889; 

Monograph, pt. 1, p. 47, 1893.— Hulst, Phycitidae of N. 

Amer., p. 117, 1890.— McDunnough, Check list, No. 6062, 

1939. 
Myelois zonulella Ragonot, Ent. Amer., vol. 5, p. 113, 1889; 

Monograph, pt. 1, p. 49, 1893.— Forbes, Cornell Mem. 68, p. 

613, 1923.— McDunnough, Check list. No. 6066, 1939. (New 

synonymy.) 
Myelois obnupsella Hulst, Phycitidae of N. Amer., p. 118, 1890. — 

Ragonot, Monograph, pt. 1, p. 48, 1893. — Barnes and 

McDunnough, Contributions, vol. 3, p. 193, 1916. — Forbes, 

Cornell Mem. 68, p. 613, 1923. — McDunnough, Check list. 

No. 6063, 1939. (New synonymy.) 

Forewing brownish gray with some faint whitish dust- 
ing on basal and median costal areas; antemedial line 
but slightly oblique, rather faint, dull whitish with a 
more or less obscured dark outer border; subterminal 
line obsolete or very faintly indicated; discal dark dots 
at end of cell separated, only the lower one distinct and 
always distinguishable. Hind wings dull smoky white 
to pale fuscous ; veins darkened slightly in several speci- 
mens; a narrow dark line along termen. Alar expanse, 
20-24 mm. 

Male genitalia similar to those of coniella except scle- 
rotized costal margin of harpe abruptly terminated before 
apex of cucullus and not projecting as a free spur at apex. 
Female genitalia with a small weak signum; bursa 
weakly sclerotized, finely scobinate, and with a longi- 
tudinal sclerotized groove in area near ductus bursae; 
ductus bursae very short; ductus seminalis from biursa 
well towards its terminal end. 

Type localities: "North America" {subtetricella, in 
Paris Mus.); north Illinois {zonulella, in BM); Canada 
{obnupsella, in AMNH, ex Rutgers). 

Food plant: Unlaiown. 

Distribution: United States: New Hampshire, 
Hampton; Massachusetts, Cohasset (July), Forest Hills 
(May, June), Framington (May), Winchendon (May); 
Pennsylvania, Beaver County (May), New Brighton 
(May, June), Pittsburgh (May); Illinois, Arlington 
Heights (May), Chicago (May), (^uincy (June); Ohio, 
Calla; Florida (no specific locality, Mar.). Canada: 
Alberta, Bilby (June), Edmonton (May); Manitoba, 
Aweme (May, June). 

The species is quite distinct and easUy identified by 
male and female genitalic characters. The Florida rec- 
ord cited above is from a spurious "tjrpe (male)" of 
Myelois immundella Hulst, originally in the Fernald Col- 
lection and now in the U. S. National Museum. It is 
not immundella, and in genitaha, color and markings 
agrees perfectly with other males of subtetricella. Rag- 
onot's zonulella was described from four females in the 
British Museum labeled "N. lU." and bearing the num- 



ber "82-54." I have examined the genitalia of two of 
these and they agree in all details with those of the type 
of subtetricella. As Ragonot designated no holotype I am 
selecting as lectotype one of the specimens I examined. 

74. Myelopsis minutularia (Hulst)) new combination 

Figure 675 

Dioryctria minutularia Hulst, Ent. Amer., vol. 3, p. 135, 1887. 
Myelois minutulella Hulst, Phycitidae of N. Amer., p. 118, 1890. — 

Ragonot, Monograph, pt 1, p. 48, 1893. 
Myelois minutularia (Hulst) McDunnough, Check list. No. 6064, 

1939. 

The status of this species is in doubt. It is known 
only from females which look like small dark examples 
of coniella, of which it may be only a race or variety. 
However, the ductus bursae of minutularia is longer than 
that of typical coniella and the bursa shows considerably 
more scobination. Hind wing semihyaline, smoky 
white. Alar expanse, 11-13 mm. 

Type locality: Blanco County, Tex. (type in 
AMNH, ex Rutgers). 

Food plant: Unknown. 

The only knoAvn distribution is Texas; examples be- 
fore me are from Blanco and Burnet Counties. The 
statement by Hulst in his original description that his 
types are males is an error. The male is unknown. 

75. Myelopsis alatella (Hulst), new combination 
Figures 193, 194, 195, 674 

Acrobasis alatella Hulst, Ent. Amer., vol. 3, p. 135, 1887. 

Myelois rectistrigella Ragonot, N. Amer. Phycitidae, p. 3, 1887. 

Myelois alatella (Hulst) Hulst, Phycitidae of N. Amer., p. 118, 
1890.— Ragonot, Monograph, pt. 1, p. 52, 1893. — McDun- 
nough, Check list, No. 6070, 1939. 

Myelois fragilella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 
114, 1904.— McDunnough, Check list. No. 6060, 1939. 
(New synonymy.) 

Myelois piazzella Dyar, Ins. Insc. Menstr., vol. 13, p. 11, 1925. — 
McDunnough, Check list. No. 6061, 1939. (New 
synonymy.) 

Forewing ash gray more or less dusted with fuscous, 
general color varying from pale ash gray to grayish 
fuscous (but not so dark as some specimens of coniella) ; 
antemedial line oblique, indicated by its narrow, black 
outer border which is shortly and sharply out-angled 
at middle; subterminal line rather close and paralled 
to outer margin, sinuate, sharply indented between 
costa and vein 6, very slightly so at lower fold, often 
obscure, sometimes with a distinct inwardly bordering 
black line; discal dots separated, black, lower one (at 
least) always distinct. Hind wing semihyaline smoky 
white, somewhat darkened towards apex and with more 
or less darkening of the veins; a fine dark line along 
termen. Alar expanse, 20-26 mm. 

Male genitalia with transtilla a thin, weakly 
sclerotized sub triangulate plate. Harpe with costa 
broadly sclerotized, produced at apex, but not extending 
to apex of cucullus. Female genitalia with a small 
signum; a rather large round area of dorsal surface of 
bursa thinly sclerotized. 



42 



UNITED STATES NATIONAL MUSEUM BULLETIKr 2 07 



Type localities: Napa, Calif, {alatella, in AMNH, 
ex Rutgers); California (rectistrigella, in Paris Mus.); 
Pecos, N. Mex. (Jragilella, in USNM); San Diego, 
Calif, (piazzella, in USNM). 

Food plant: Unknown. 

Distribution: California, Clarkville ( June) , Monache 
Meadows (July), Napa, Placerville (May), San Diego 
(Mar., Aug.), San Francisco (Apr.); Utah, Bellevue 
(Apr.); Colorado, Gunnison County (near Almont, 
July) ; New Mexico, Fort Wingate (June, July) , Jemez 
Springs (June), Pecos (June). 

An individually variable species in color and to some 
extent in male genitalia. The actual holotypes of 
alatella, fragilella, and piazzella seem different enough; 
but there are all intergrades among them in a series 
from any given locality. Indeed the two cotypes of 
alatella from Napa, Calif. (aUke in color and markings) 
show considerable variation in details of male genitalia 
(width of the sclerotized costa of harpe, shape of 
transtilla, and spacing of the apical prongs of gnathos) . 
The genitalia of the cotype from Napa (in USNM, fig. 
193) shows an extreme of variation. The other cotype 
(in AMNH, ex Rutgers, the actual holotype) has 
genitalia identical with those of piazzella shown in 
figure 195. At most, the Dyar names represent forms 
or varieties, but not species or local races. 

19. Genus Anypsipyla Dyar 

Anypsipyla Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 327, 1914. 
(Type of genus: Anypsipyla univitella Dyar.) 

Tongue well developed. Antenna of male shortly 
cihate, the cUia no longer than width of shaft; of female 
pubescent. Labial palpus obliquely uptm-ned, reach- 
ing slightly above vertex; third segment about as long 
as second, acuminate. MaxiUary palpus very slightly 
dilated with scales at apex (subsquamous) . Forewing 
smooth; 11 veins; vein 2 from before but close to lower 
outer angle of cell; 3 from the angle, approximate to 
2 ; 4 and 5 stalked for approximately half their lengths, 
approximate (rarely connate) to 3 at base; 6 from below 
upper angle of cell, straight; 8 and 9 stalked for about 
half their lengths; 10 from the cell, at base closely 
approximate to or connate with stalk of 8-9, thence 
divergent; male with short narrow costal fold. Hind 
wing with vein 2 from well before lower outer angle of 
cell; vein 3 from the stalk of 4-5; 4 and 5 long stalked; 
7 and 8 anastomosed beyond cell for appreciably more 
than half their lengths; cell about half the length of 
wing; discocellular vein curved. Eight abdominal 
segment of male with a single pair of ventrolateral hair 
tufts. 

Male genitalia with apical projection of gnathos an 
elongate hook with slightly notched apex. Uncus sub- 
triangulate (hoodlike). Transtilla complete, strongly 
arched. Harpe with costa strongly sclerotized through- 
out and projecting at apex beyond apex of cucullus; 
otherwise simple. Anellus a narrow band with slender 
lateral arms. Aedeagus simple; penis with some 
weakly sclerotized wrinklings, otherwise unarmed. 



Female genitalia with or without signa, if present, in 
the form of a row of very small, weak, thornlike spines ; 
bursa very finely scobinate, ductus bursae considerably 
longer than bursa, simple; genital opening simple; 
ductus seminalis from anterior (terminal) end of bursa. 

A distinct genus with one tropical American species. 

76. Anypsipyla univitella Dyar 
FiGTiRES 39, 196, 679 

Anypsipyla univitella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 327, 
1914. 

Forewing fuscous gray with a broad white subcostal 
streak extending from near base to apex and touching 
costa near base and at apex; a black streak along mid- 
costal edge and on fresh specimens a fine black line along 
lower vein of ceU and some faint black streaking on the 
outer veins; a fine powdering of reddish scales on the 
white subcostal stripe; discal dots faint or absent; trans- 
verse lines obsolete. Hind wing hyaline white with a 
smoky tint along costa and at apex and a fine dark line 
along termen. Alar expanse, 20-32 mm. 

Male genitalia with terminal margin of uncus rather 
broadly round; apical projection of costa of harpe blunt; 
transtilla truncately arched. Female genitalia as given 
for the genus. The signa are usually absent and when 
present consist of from 2 to 10 very weak spines. 

Type locality: Corozal, Canal Zone, Panamd (type 
in USNM). 

Food plants: Cassia brasiliensis, Samanea samdn 
(larva feeding in pods), Pacae (larva in fruit). 

Distribution: Cuba: Victoria de las Tunas, San 
Bias (Trinidad Mts., May). Mexico: Colima (May, 
Nov.). Guatemala: Cayuga (Apr.), Quirigud (Mar.). 
PanamA: Corozal (Apr., Nov.), Las Sabanas (Apr.), 
Porto Bello (May). Venezuela: El Valle (Apr.). 
Brazil: "S. E. Brazil," Tapera {Pemambuco). Peru: 
Lima (Feb.). Ecuador. Jamaica: Kingston (Dec). 

Probably generally distributed in tropical America, 
where its host plants occiu*. 

20. Apomyelois, new genus 

Type of genus: Dioryctria bistriatella Hulst. 

Tongue well developed. Antenna simple and pubes- 
cent on both sexes. Labial palpus upturned, slender, 
reaching to slightly above vertex; third segment slightly 
shorter than second, acuminate. Maxillary palpus fili- 
form. Forewing smooth; 11 veins; vein 2 from well 
before lower outer angle of cell; 3 from the angle; 4 and 
5 stalked for slightly less than half their lengths, the 
stalk separated from 3 at base; 6 from below upper 
angle of cell, straight; 8 and 9 long stalked (for over 
two-thirds their lengths) ; 10 from the stalk of 8-9 ; male 
without costal fold. Hind wing with vein 2 from weU 
before lower outer angle of ceU; veia 3 from the angle; 
4 and 5 stalked for two-thirds of their length, the stalk 
connate with 3; 7 and 8 anastomosed beyond cell for 
about half their lengths (the anastomoses slightly longer 
than the free part of vein 8) ; cell a trifle more than half 
the length of the wing; discocellular vein curved. Eighth 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



43 



abdominal segment with a pair of weak, short, ventro- 
lateral hair tufts. 

Male genitalia with apical projection of gnathos an 
elongate, slender, simple hook. Uncus subtriangulate, 
apex roxmded. Transtilla complete, weU sclerotized 
and strongly arched; produced at middle into a flat, 
broadly and bluntly forked projection. Harpe with 
costa sclerotized throughout and projection slightly at 
apex; otherwise simple. Anellus U-shaped. Aedeagus 
smooth, slightly flaring at apex; penis with a single, 
straight, short, weakly sclerotized, spikelike cornutus 
and a few minute and weak scobinations; otherwise 
unarmed. 

Female genitalia with signa present as an oval cluster 
of thornlike scobinations ; bursa otherwise smooth, large; 
ductus bursae considerably shorter than bursa, simple; 
genital opening simple; ductus seminalis from anterior 
(terminal) end of bursa. 

This genus is another subtraction from the composite 
genus Myelois of Authors. Of all the American species 
that have been referred to that genus it is the nearest 
to the type of Myelois {medullalis Hiibner, a synonym 
of cribrella Hiibner) of any American species, agreeing 
with cribrella in forewing venation, except that the stalk- 
ing of vein 10 with 8-9 is less consistent in cribrella than 
in bistriatella. In cribrella 10 is often short stalked (as 
in fig. 38) ; but it is as often from the cell, connate with 
or approximate to or distinctly separated from the stalk 
of 8-9. In bistriatella it is from the stalk of 8-9 on aU 
specimens that I have seen. However, this difference 
has no more than specific significance and would not of 
itself justifj' any separation of bistriatella from Myelois. 
There are some other differences that, in my judgment, 
are of generic character and justify such separation. 
The hind wing venation and length of cell are similar in 
cribrella and bistriatella except for the anastomosis of 
veins 7 and 8 ; in cribrella this anastomosis is very weak 
and shorter than it is in bistriatella and would place 
typical Myelois in our venational division B, while 
Apomyelois would go definitely into division C. In male 
genitaUa cribrella (fig. 203) differs in having apical pro- 
jection of gnathos developed as a deeply, strongly, and 
narrowly forked process and the transtilla developed as 
a simple, strongly arched, narrow band. The female 
genitalia of cribrella (fig. 684) differs in more striking 
fashion: the ductus biusae being much longer than 
bursa and strongly granulate and partially sclerotized 
throughout most of its length; and ductus seminalis is 
from the bursa between the signum and the junction of 
bursa and ductus bm-sae. 

The new genus contains one North American species. 

77. Apomyelois bistriatella (Hulst), new combination 
Figures 40, 197, 676 

Dioryctria bistriatella Hulst, Ent. Amer., vol. 3, p. 136, 1887. 
Myelois bilineatella Ragonot, N. Amer. Phycitidae, p. 3, 1887; 

Monograph, pt. 1, p. 48, 1893. — Hulst, Phycitidae of 

N. Amer., p. 117, 1890; U. S. Nat. Mus. BuU. 62, p. 418, 

1902. 
Myelois bistriatella (Hulst) Hulst, Phycitidae of N. Amer., p. 117, 

1890. — Ragonot, Monograph, pt. 1, p. 51, 1893. — Barnes 



and McDunnough, Contributions, vol. 3, p. 194, 1916. — 
Forbes, Cornell Mem. 68, p. 613, 1923.— McDunnough, 
Check list, No. 6067, 1939. 

Forewing gray-brown faintly dusted with white on 
costal half of basal area and in central area from mid- 
costa to lower margin of cell; transverse lines white, 
rather sharply contrasted, especially towards inner 
margin and without appreciably contrasted blackish 
borders; antemedial line transverse, from costa dis- 
tinctly before middle, straight, except for an occasional 
sUght notch in ceU ; subterminal line somewhat narrower 
and less distinct, sinuate; dark discal dots at end of 
cell often fused into a single spot or line along discocel- 
lular vein, usually set off by the surrounding white 
dusting of the central area; terminal dots very faint, 
more or less confluent. Hind wing dull smoky white, 
the veins slightly darkened and a narrow dark line 
along termen. Alar expanse, 19-22 mm. 

Genitaha as given for the genus. 

Type localities: Washington, D. C. (bistriatella, 
type lost?) ; "America septentrionale" (bilineatella, in 
Paris Mus.). 

Food plant: Unknown. 

Distribution: United States: Florida (Msir.); Dis- 
trict of Columbia, Washington; A^ew York; Massachusetts, 
Framingham (May) ; Illinois, Edgebrook (May) ; Wis- 
consin; loiva, Ames (May). Canada: Ontario, Ottawa 
(June, July), Trenton (May, June). 

Probably much more widely distributed thi-oughout 
eastern and central United States and Canada, nowhere 
apparently a very abundant species. 

The supposed type of bistriatella is labeled "Iowa, 
H. S. Saunders, June 6, 1886." It is definitely that 
species but, unless it is mislabeled or the type locality 
given by Hulst in his original description ("Washington, 
D. C") is wrong, it could not be the actual holotype. 
I have seen no specimens anywhere labeled "Washing- 
ton, D. C." There is a female in the National Museum 
from the Fernald Collection, bearing a Hulst type label 
but no locality. This might be the true type. It is a 
perfect match for the Iowa specimen in the Rutgers 
Collection. Since there can be no reasonable doubt as 
to what the name stands for we may as well consider 
the holotype lost and forget it. 

21. Ectomyelois, new genus 

Type of genus: Myelois decolor ZeUer. 

Tongue well developed. Antenna of male shortly 
ciliate (cilia about the length of width of shaft or 
shghtly less), otherwise simple; of female simple and 
pubescent. Labial palpus upturned, reaching to or 
nearly to apex (not above it); second segment some- 
what broadened with scales; third segment short, 
distinctly shorter than second, acuminate. Maxillary 
palpus filiform. Forewing smooth; 11 veins; vein 2 
from well before angle; 3 from the angle, shortly 
separated from the stalk of 4-5 at base; 4-5 shortly 
stalked (very shortly staUied in most specimens and 
never for more than half the length of the veins); 



44 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



6 from below upper angle, straight; 8 and 9 long stalked, 
for over two-thirds of their lengths; 10 from the cell, 
closely approximate to the stalk of 8-9 for some distance 
from cell; male without costal fold. Hind wing with 
vein 2 from well before lower outer angle of cell; 3 from 
the angle; 4-5 stalked for not over half their lengths 
(usually for less), the stalk connate with or very closely 
approximate to 3 at base; 7 and 8 strongly anastomosed 
for most of their lengths beyond cell (free element of 8 
very short); cell half the length of wing; discocellular 
vein ciu'ved. Eighth abdominal segment of male 
simple or with a weak, short pair of ventrolateral hair 
tufts. 

Male genitalia similar to those of the foregoing 
genus (Apomyelois) except: Apical process ofgnathos 
slightly notched at apex; costa of harpe not produced 
at apex (except slightly in muriscis and zeteki); penis 
without cornutus; vinculum more truncate and less 
tapering. 

Female genitalia with signum sometimes absent, 
when present consisting of an elongate patch of scobina- 
tions; ductus bursae normally longer than bursa, in- 
dividually variable, simple, except for a weak scleroti- 
zation at genital opening; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

In male genitalia there is little or nothing of a generic 
character to separate Ectomyelois from Apomyelois. 
The two genera are distinguished by the shorter cell 
and much more extended anastomosis of veins 7 and 8 
of hind wing and the different place of departure of 
ductus seminalis from the biu'sa of the female. 

The genus is erected for another group of species 
(American and European) removed from Myelois of 
Authors. These species all appear to be of tropical or 
semitropical origin. They are distinguished from 
typical Myelois by the much stronger anastomosis of 
veins 7 and 8 of hind wing and the consequent reduction 
of the free element of vein 8. 

78. Ectomyelois decolor (Zeller), new combination 
Figures 198, 677 

Myelois decolor Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 222, 
1881. — Ragonot, Monograph, pt. 1, p. 58, 1893. — Dyar, 
Proc. U. S. Nat. Mus., vol. 47, p. 326, 1914.— Wolcott, 
Journ. Agr. Univ. Puerto Rico, vol. 20, No. 1, p. 476, 1936. 

Nephopteryx ephestiella Hampson, Ann. Mag. Nat. Hist., ser. 7, 
vol. 7, p. 257, 1901 (new synonymy). 

Forewing dark grayish fuscous with some white 
powdering in basal area and considerably more in the 
median area from slightly above inner margin and in 
outer area between subterminal line and termen; trans- 
verse lines white, well contrasted, especially the ante- 
medial line which is rather wide, sharply oblique, 
slightly indented at lower fold and (in some specimens) 
in the cell, outwardly bordered by a more or less ob- 
scure dark shade; subterminal line fainter, narrow, 
sinuate, obscurely and narrowly dark margined; some 
faint blackish streaking on the veins; discal dots at end 
of cell distinct, separated; terminal dots normally well 
contrasted and separate, blackish. Hind wings smoky 



white to pale smoky fuscous ; the veins darkened and a 
narrow dark line along termen. Eighth abdominal 
segment of male with hair tufts. Alar expanse, 19-30 
mm. 

Male genitalia with outer margin of uncus rather 
evenly rounded; central area of transtilla produced into 
a moderately broad plate with notched terminal margin; 
anellus a broad plate with wide, flattened, incurved, 
stubby arms; vinculum nearly square in outline, its 
terminal margin very slightly concave. 

Female genitalia exhibiting considerable individual 
variation in the size of bursa and corresponding length 
of ductus bm^ae which is usually considerably longer 
than bursa; signum patch of variable shape but usually 
elongate. The female genitalia exhibit no distinctively 
specific characters. 

Type localities: Honda, Colombia {decolor, in 
BM); Nassau, Bahamas (ephestiella, in BM). 

Food plants: Annona squamosa, Ceratonia sUiqua, 
Hymenaea courbil; these records from reared specimens 
in the U. S. National Museum. Presumably the species 
has much the same hosts and habits as the closely re- 
lated Ectomyelois ceratoniae. The larvae feed in the 
fruits and are very difficult to separate from those of 
ceratoniae. 

Distribution: Cuba: Baracoa (Aug., Oct., Nov.), 
Havana, Santiago de las Vegas (Mar.), "Santiago Prov- 
ince" (Sept., Oct., Dec). Puerto Kico: Arecibo, 
San German (Apr.). Jamaica. Bahamas: Nassau. 
Guatemala: Caynga (Mar., June, Aug.). PanamX: 
Porto Bello (Mar., Dec). Colombia: Honda, "West 
Slopes" (4,400 ft., Feb.). Venezuela: Aroa. British 
Guiana: Tumatumari (June). French Guiana: CAy- 
enne, St. Jean Maroni. Surinam: Surinam Kiver. 
Brazil: Pard (June), Ponte Nova (Rio Xingu, Amazon- 
as), Santa Catarina (July). 

Ragonot considered decolor as a probable variety of 
ceratoniae; but there is a consistent difference in the 
shape of the transtilla between the two which indicates 
more than varietal or racial difference; and in unrubbed 
and unfaded specimens the color difference is obvious 
and consistent. E. decolor seems to be confined to the 
New World while ceratoniae occurs in both the New and 
Old World. Hampson's ephestiella is nothing more than 
a rather large example of decolor. Like other species in 
this genus, decolor varies greatly in size. 

79. Ectomyelois ceratoniae (Zeller), new combination 
Figures 199, 678 

Myelois ceratoniae Zeller, Isis von Oken, 1839, p. 176; 1848, p. 
676. — Ragonot, Monograph, pt. 1, p. 57, 1893. — Staudinger 
and Rebel, Catalog der Lepidopteran des palaearctischen 
Faunengebietes, vol. 2, No. 787, 1901. — Spuler, Die Schmet- 
terlinge Europas, vol. 2, p. 216, 1910. — Forbes, Cornell 
Mem. 68, p. 614, 1923. — Meyrick, Revised handbook of 
British Lepidoptera, p. 395, 1928. — Wolcott, Journ. Agr. 
Univ. Puerto Rico, vol. 20, No. 1, p. 476, 1936.— Corbet 
and Tarns, Proc. Zool. Soc. London, ser. B, vol. 113, p. 68, 
1943. 

Myelois oporedestella Dyar, Proc. Ent. Soc. Wash., vol. 13, p. 30, 
1911.— MoDunnough, Check list. No. 6065, 1939. (New 
synonymy.) 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



45 



Color and markings similar to decolor except: Fore- 
wing more imiformly gray, with less of the white dust- 
ing, especially on median area; antemedial line narrower 
and usually more distinctly notched. The chief char- 
acter, however, is in the transtilla of the male genitaUa. 
The central projection of this organ is more constricted 
and decidedly narrower on ceratoniae than on decolor, 
and this difference seems to be consistent. In several 
preparations of each species from different rearings and 
localities I have found no intergrading examples. The 
female genitaha offer no satisfactory distinguishing 
characters. As in decolor, individual differences (even 
in the proportional length of the ductus bursae) are 
greater than any difference between the two species. 
Alar expanse, 16-24 mm. 

Type localities: Laibach, Austria (ceratoniae, in 
BM); Miami, Fla. (oporedestella, in USNM). 

Food plants: Carissa grandiHora, Cassia bicapsularis, 
Ceratonia siliqua, Erishotyra japonica (chiefly in mum- 
mied fruits), Livistona chinensis, Rohinia, Tamarindus 
indica, Vachellia instdaris. Also on dried figs, dates, 
raisins, and nuts in storage. Primarily a leguminous 
feeder. The favored host seems to be the pods and 
seeds of the corob (Ceratonia siliqua) . 

Distribution: United States: Florida, Homestead 
(May), Key West (Apr.), Miami (May, Jidy, Aug., 
Nov.). Puerto Rico: Arecibo (May), TrujOlo Alto 
(Mar., July). Jamaica (July). Argentina: Buenos 
Aires (Feb.), Catamarca (May, June). Also in the Old 
World in the Mediterranean areas of Europe, Africa 
and Asia and (by introduction in dried fruits) extending 
into Central Europe and England. 

Apparently of Mediterranean origin, introduced by 
commerce and estabhshed in some tropical and semi- 
tropical areas of the New World. Probably much more 
widely distributed than indicated by the above records 
from specimens before me. The species is of minor 
importance as a feeder on the seeds of the corob. It 
has been foimd rather frequently at our port quarantine 
stations in shipments of English walnuts from Italy. 

I have omitted all European synonymy as I have not 
been able to verify its correctness. This, with further 
references to European literature will be foimd in 
Ragonot's monograph and the Staudinger and Rebel 
catalog. Myelois phoenicis Durrant may be only a 
color variety or race of ceratoniae; a small series before 
me reared from dates from Algeria has the ground color 
of forewing white, but the male genitalic characters 
of ceratoniae. Corbet and Tams list phoenicis as a 
synonym. 

80. Ectomyelois muriscis (Dyar), new combination 
Figures 200, 680 

Myelois Iransitella Dyar (not Walker), Proc. U. S. Nat. Mus., 

vol. 47, p. 326, 1914. 
Hypsipyla muriscis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330, 

1914. 
Myelois palpalis Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 1919 

(new synonymy). 



Forewing duU rusty brownish ocherous to reddish 
brown; costal third to half of wing strongly dusted with 
white, the white area rather weU contrasted against 
dark ground color; antemedial line angidate, obscure, 
indicated chiefly by a brown or blackish spot on its 
outer margin at or just below costa; subterminal line 
better defined, sinuate, margined inwardly and out- 
wardly by narrow dark lines, the latter especially 
emphasized at costa; discal spots usually distinct and 
separated, sometimes one or the other obscured by the 
white dusting or by an extension of the ground color, 
rarely fused into a line along discocellular vein, blackish 
brown; terminal dots more or less distinct. Hind wing 
dull, translucent white to smoky fuscous (as a rule 
darker on female than on male) ; a dark shade toward 
apex, some dark shading on the veins and a fine dark 
line along termen. Eighth abdominal segment of male 
simple. Alar expanse, 16-26 mm. 

Male genitaha with apical projection of gnathos slen- 
der, very long, extending at least as far backward as 
apex of uncus (when genitalia are in natural position it 
extends well beyond the uncus) ; transtUla a rather nar- 
row, sclerotized band, looped backward in a rounded 
arch; sclerotized costa of harpe very slightly and 
bluntly produced at apex. 

Female genitaUa mth or without signa, when present 
a patch of coarse scobinations, the patch varying in size 
in different specimens; ductus bursae simple or very 
weakly sclerotized on ventral surface at genital opening. 

Type localities: Cabima, Panamd (muriscis, in 
USNM); Cayuga, Guatemala (palpalis, in USNM). 

Food plants: Mammea americana (larvae feeding in 
the fruit), Theobroma cacao (larvae in the pods). 

Distribution: Haiti. Puerto Rico, Mayaguez 
(July). British West Indies: Trinidad, several exam- 
ples with no more specific locality, St. Clair (Mar.); 
Grenada, several examples with no more specific locality; 
Tobago (Apr.). Guatemala: Cayuga (Jan., Feb., Apr., 
May, June), Quirigu^ (Sept.). Costa Rica: Esperanza 
(May, Aug.). PanamX: Alhajuelo (Apr.), Cabima 
(May), Porto Bello (Apr., Oct.), Rio Trmidad (Mar., 
June). Colombia: La Esperanza (Dec), no specific 
locality (June). BolIvia: "East Bolivia" (Oct.). 
British Guiana: "Mazaruni Clearing" (Aug., Oct.). 
French Guiana: Cayenne, St. Jean Maroni, St. 
Laurent Maroni. Brazil: Rio de Janeiro (June). 

This species is primarily a feeder in the pods of the 
cacao and is weU distributed in tropical America wher- 
ever its host occurs. All specimens in the National 
Museum (except the holotypes of muriscis and palpalis) 
had been identified by Dyar as "Myelois transitella 
Walker." The two species are easily confused on super- 
ficial characters, especially among faded and stained 
tropical specimens; but their genitalia are quite distinct. 

Dyar's types of muriscis and palpalis are males and 
aUke in genitalic and all other characters. It is very 
probable that muriscis eventually will prove to be no 
more than a variety (or synonym) of jurvidorsella 
Ragonot. 



46 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



81. Ectomyelois furvidorsella (Ragonot), new combination 

Figure 681 

Myelois furvidorsella Ragonot, Nouv. Gen., p. 8, 1888; Mono- 
graph, pt. 1, p. 56, 1893. 

This is probably the same as muriscis. The genitalia 
of the female type (fig. 681) are somewhat unusual in 
that bursa and ductus biirsae are perfectly smooth, with 
no trace of signum or sclerotization of the ductus at 
genital opening. However, I have seen similar geni- 
talia in typical examples of muriscis from Central 
America; but I have seen so few examples of muriscis 
(only males) from Puerto Rico that I prefer to keep the 
names apart tUl more material is available. 

Alar expanse, 22 mm. 

Type locality: Puerto Rico (type in Paris Mus.). 

Food plant: Unknown. 

82. Ectomyelois zeteki, new species 
Figures 201, 682 

Forewing pale brownish gray faintly dusted with 
blackish fuscous; antemedial Une obscure, angxilate, 
indicated chiefly by its blackish fuscous outer border, 
the latter incomplete on many specimens; sub terminal 
line sinuate, rather close to termen, on weU-marked 
specimens consisting chiefly of white spots on the veins, 
preceded and followed by dark streaks, obscure on 
many specimens; also on well-marked specimens a me- 
dian, longitudinal blackish fuscous streak from base of 
wing to antemedial line; discal dots at end of ceU very 
faint or completely obliterated, when distinguishable 
more or less confluent; terminal dark dots faint, con- 
fluent. Hind wing translucent, white with a faint 
smoky tint towards apex; a dark line along outer margin 
and some darkening of the veins. Eighth abdominal 
segment of male simple. Alar expanse, 17-22 mm. 

Male genitalia similar to those of muriscis except: 
Uncus narrower; apical projection of gnathos shorter, 
not reaching so far backward as apex of uncus ; transtiQa 
a narrow band forming a truncated arch with slightly 
concaved posterior margin; sclerotized costa of harpe 
projecting somewhat further at its apex. Female geni- 
talia with scobinations of signum patch stouter and 
sclerotization of ductus bursae at genital opening fore- 
wing a larger and more strongly pigmented shield than 
those of any other species of the genus. 

Type locality: Near Capira, Panamd (type in 
USNM, 61316; paratypes in USNM, Cornell Univ., 
Transvaal Mus. (Janse Coll.), Paris Mus., BM). 

Food plant: Cassia moschata. 

Described from male type, and 16 male and 19 
female paratypes from the type locality; all reared (May 
1941, Zetek No. 4807) by James Zetek, who has con- 
tributed much valuable material to the National Collec- 
tion, and for whom the species is named. 

It is easily distinguished from any other American 
phycitid by its male genitaha. 



Genera 22-24: Paramyelois to Protomoerhes 

[Venational division C. Forewing with 11 veins; 4 and 5 stalked 
for less than haU their lengths. Hind wing with veins 7 and 8 
anastomosed for most of their lengths (free element of 8 short) . 
Transtilla incomplete; but (except in Paramyelois) its free 
elements well developed.] 



22. Paramyelois, new genus 

Type of genus: Myelois solitella Zeller. 

Tongue well developed. Antenna of male shortly 
cilia te (cilia shorter than width of shaft), simple; of 
female pubescent. Labial palpus oblique, laterally flat- 
tened (broad and flat from lateral view) ; second segment 
roughly scaled beneath; third segment shorter than 
second, somewhat roughly scaled. Maxillary palpus 
squamous (rather heavily and broadly scaled). Fore- 
wing smooth; 11 veins; vein 2 from before but rather 
near lower outer angle of cell; 3 from the angle, well 
separated from the stalk of 4-5 at base, but nearer to 
4-5 than to 2; 4 and 5 shortly stalked; 6 from below 
upper angle of cell, straight; 8 and 9 long stalked; 10 
from the cell, approximate to the stalk of 8-9 for a 
short distance from base; male without costal fold. 
Hind wing with vein 2 from well before lower outer 
angle of cell; 3 from the angle, closely approximate to 
or connate with the stalk of 4-5 at base; 4 and 5 nor- 
mally stalked for half or slightly less than half their 
lengths, rarely (in some small specimens) stalked for 
over half their lengths: 7 and 8 strongly anastomosed 
for most of their lengths, free element of 8 short; cell 
slightly over half the length of wing in male (as in 
European Mj/eZois), half the length of wing in female; 
discoceUular vein curved. Eighth abdominal segment 
of male simple. 

Male genitalia with apical process of gnathos a short, 
rather broad, blunt hook, slightly notched at apex. 
Uncus with broad base; narrowed and triangulate just 
beyond; apex ateutely rounded. Tegumen short and 
broad. Transtilla incomplete. Harpe very broad at 
base; costa strongly and broadly sclerotized and forming 
a broad, pointed projection before middle, not appreci- 
ably sclerotized beyond; sacculus large and strongly 
sclerotized ; cucullus greatly reduced. Anellus a curved 
shield with long, strongly sclerotized, smooth, slender, 
tapering and pointed lateral arms. Aedeagus scobinate 
on one lateral edge at apex; penis with a few sclerotized 
wrinklings, otherwise unarmed. Vinculum stout, slightly 
longer than broad, truncate, scarcely tapering to broad 
terminal margin. 

Female genitalia with weak signum consisting of a 
cluster of rather coarse scobinations; finer scobinations 
scattered over the caudal half of bursa. Ductus biu:sae 
shorter than bursa, broadened and sclerotized on inner 
ventral and lateral surfaces towards genital opening; on 
inner dorsal surface behind the opening a pair of small 
sclerotized plates. Ductus seminalis from bursa near 
its junction with ductus biu-sae. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



47 



This genus is easily recognized and is distinguished 
from other sections of the "Alyelois" complex by its 
labial palpi and male genitalia. 

In their normal position the palpi are directed in a 
straight line obliquely from the face; but many speci- 
mens show the third segment more or less deflefcted for- 
ward, and some with both the second and third seg- 
ments more or less porrected, results of the death con- 
tortions of the moths. Several European species listed 
under Myelois have oblique palpi but they are all more 
or less cylindrical and do not have the broadly scaled 
and flattened lateral aspect of those of Paramyelois. 
The male genitalia with their incomplete transtilla (its 
elements reduced and well separated) are unique among 
the groups nearly related to Myelois or any of the Ameri- 
can species that previously have been referred to that 
genus. 

I have chosen a sjmonym as type of the new genus 
advisedly, as the type specimen of the oldest name 
{transiteUa) is a female, and there may be some question 
of my application of the name to the species here treated. 
There can be no such doubt in regard to solitella. 

83. Paramyelois transit ella (Walker), new combination 
Figures 202, 683 

Nephopteryx transiteUa Walker, List, pt. 27, p. 54, 1863. 

Nephopteryx notatalis Walker, List, pt. 27, p. 57, 1863. 

Myelois solitella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 217, 
1881.— Ragonot, Monograph, pt. 1, p. 55, 1893. (New 
synonymy.) 

Myelois duplipunctella Ragonot, N. Amer. Phycitidae, p. 3, 1887; 
Monograph, pt. 1, p. 56, 1893. — McDunnough, Check list, 
No. 6059, 1939. (New synonymy.) 

Myelois transiteUa (Walker) Ragonot, Monograph, pt. 1, p. 42, 
1893 (in part).— McDunnough, Check list. No. 6058, 1939 
(in part). 

Myelois venipars Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 404, 
1914. — Mote, Monthly Bull. California Dep. Agr., vol. 11, 
p. 628, 1922. — Glick, Arizona Comm. Agr. and Hort., Four- 
teenth Ann. Rep., p. 78, 1922. — Essig, Insects of western 
North America, p. 708, 1929. — Hixon, Journ. Econ. Ent., 
vol. 27, p. 547, 1934. (New synonymy.) 

Emporia cassiae Dyar, Ins. Insc. Menstr., vol. 5, 91, 1917 
(new synonymy). 

Similar in color and maculation to Ectomyelois muris- 
cis except: Ground color on lower half of wing darker; 
the dark borders of the transverse lines and the discal 
dots decidedly darker, blackish; the white areas more 
strongly contrasted ; the dark outer border of antemedial 
complete in most examples and enlarged below costa 
into a conspicuous blackish spot. Hind wings a clearer 
white on the males ; more or less smoky on the females. 
Alar expanse, 15-28 mm. 

Genitalia as given for the genus. The male genitalia 
show little or no individual variation. Among the fe- 
males, however, there is considerable variability in mi- 
nor details, namely, the amount of sclerotization about 
genital opening and the amount of scobination in the 
bursa, but these are differences of no specific significance. 

Type localities: "United States," probably Florida 
(transiteUa, in BM) ; Santo Domingo (^notatalis, in BM) ; 
Colombia (solitella, in BM) ; Florida (duplipunctella, in 



Paris Mus.); Hermosillo, Mexico (venipars, in USNM); 
Georgetown, British Guiana (cassiae, in USNM). 

Food plants: Orange, grapefruit, peach, apple, dates, 
figs, Acacia farnesiana, Aesculus glabra. Cassia grandis, 
Genipa americana, Gleditsia tnacanthos, Pithecolobium 
flexicaule, Robinia, Sapindus drummondii, Yucca, English 
walnut. These records from reared specimens in the 
U. S. National Museum. 

Disteibution: United States: Arizona, Maricopa 
County (Dec), Mesa (Nov.), Phoenix (Aug., Sept., 
Nov., Dec), Tempe, Yuma; Texas, Anahuac (March), 
Brownsville (Dec), Dallas (May), Fort Davis (Oct.), 
Harlingen (July), Hidalgo County (Apr.), KerrviUe, 
Louise (Feb.), Mercedes (Feb.), Mission, San Antonio 
(May) , San Benito (Sept.) ; Oklahoma, Stillwater (June) ; 
Louisiana, Forbing (Oct.); Alabama, Mobile (Apr.); 
Georgia, St. Simons; Florida, Orlando (Oct., Nov.), Vero 
Beach (Apr., May) ; North Carolina, Durham. Cuba. 
Dominican Eepublic. Mexico: Hermosillo, Oaxaca. 
Guatemala: Cayuga (Mar., Apr.), Chejel (June). Pan- 
ama: El Cermeno (Apr., June). Colombia. Brazil: 
Tapera (Pernambuco) . PerIj: Lima, Rio Pacaya (June, 
July, Aug.). 

This species has attracted some attention in the south- 
west as a minor orchard pest and is known to economic 
entomologists as the "navel-orange worm." The larvae 
feed on the nuts, in the seed pods, or on the fruits of 
numerous trees but they seem to prefer the fallen and 
mummied fruits or the dry seed pods or injured or 
diseased fruits. Rarely do they attack sound fruit on 
the trees. They have been reported as infesting sound 
oranges, but such behavior is probably an exceptional 
departure from normal habit. 

The foregoing sjTionymy requu'es some comment. I 
have not seen the types of transiteUa or duplipunctella 
(both females), but from the original descriptions and 
the Ragonot figm-es they cannot apply to anything else 
than the species we have hitherto known in the United 
States as venipars Dyar. Of the synonymy of venipars 
and solitella there is no possible doubt. The type of the 
latter is a male (not a female as stated by Zeller) and 
figures of its genitalia, supplied by Tams and Clarke, 
show agreement in every detaU with those of venipars. 
Clarke has also furnished excellent photographs of the 
female types of transiteUa and notatalis and of the geni- 
talia of transiteUa. The latter show only trifling indi- 
vidual differences from the genitalia of Dyar's female 
type of venipars. Unfortunately^, the type of notatalis 
lacks an abdomen; but photogi-aphs of the moths and 
their palpi show no essential differences between the two 
types; so Ragonot's reference of notatalis to synonymy 
must be accepted. In his specific key to the species of 
"Myelois" Ragonot (Monograph, pt. 1, p. 27) places 
transiteUa in a group with veins 7 and 8 of hind wing 
approximate. This characterization was obviously 
based upon a freak specimen. Dyar found one such 
freak (a female from Grenada) among the examples of 
muriscis which he misidentified as "transiteUa"; but in 
over a hundred examples of the true transiteUa before 
me veins 7 and 8 are strongly anastomosed, and this is 



48 



UNITED STATES NATIONAL MUSEUM BULLETIN" 207 



a good character of much more than specific value 
despite its lapse ia individual specimens. Freaks of all 
kinds can and do turn up anywhere in the Phycitinae. 
Dyar's cassiae was described from stained and faded 
females; but, even so, it is strange that he did not see 
their resemblance to his venipars, especially in their 
palpi, and still more strange that he should refer them 
to the Old World anerastiid genus Emporia. They have 
normal phycitine tongues and their genitalia agree with 
those of the female type of venipars. 

Bondar's (Instituto de Cacau da Bahia Boletim 5, 
p. 72, 1939) identification of a lepidoperon in cacao pods 
as duplipunctella Ragonot (the genus given as "Myel- 
osis") is probably incorrect. What he had was pre- 
sumably Ectomyelois muriscis. 

23. Genus Pseudodivona Dyar 

Pseudodivona Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 405, 1914. 
(Type of genus: Pseudodivona commensella Dyar.) 

Tongue well developed. Antenna shortly cUiate on 
male, cUia about as long as width of shaft (longer on 
carabayella). Labial palpus oblique, broadly scaled and 
laterally flattened; third segment short, acuminate. 
Maxillary palpus squamous. Forewing smooth; 11 
veins; vein 2 from before lower outer angle of cell; 3 
from the angle; 4 and 5 stalked for slightly less than 
half their lengths, the shaft separated at base from 3; 
6 from below upper angle of cell, slightly curved towards 
base; 8 and 9 stalked for two-thirds their lengths; 10 
from the stalk of 8-9; 11 from ceU rather near outer 
angle and running close to the stalk of 8-9-10; male 
without costal fold. Hind wing from well before lower 
outer angle of cell; 2 from very close to the angle, or 
from the angle (cispha), closely approximate to or con- 
nate with stalk of 4-5 at base; 4 and 5 stalked for about 
half their lengths ; 7 and 8 anastomosed for most of their 
lengths (free element of 8 very short) ; cell less than one- 
half (but more than a third) the length of wing; disco- 
ceUular vein curved. Eighth abdominal segment of 
male with a strong pair of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos de- 
veloped as an elongate, stout, flattened hook with 
forked or notched apex. Uncus subtriangulate, with 
rounded terminal margin. TranstUla incomplete; its 
elements well developed, elongate and knobbed at their 
apices. Harpe simple with outer margin evenly rounded ; 
costa sclerotized for about four-fifths its length; but not 
produced. AneUus a narrow curved band with slender 
lateral lobes. Aedeagus moderately slender, nearly 
straight; penis unarmed. Vinculum stout, decidedly 
longer than broad, tapering, expanded towards angulate 
terminal margin. 

Female genitalia without signum; bursa, ductus bur- 
sae, and genital opening simple; ductus seminalis from 
bm-sa near junction of bursa and ductus bursae. 

In genitalic and many other characters as well as 
wing pattern and color, this genus resembles Moerbes, 
to which it is apparently closely related. It differs 
chiefly in having vein 4 present and weU developed in 
hind wing, a different development of the elements of 



transtUla, and strong hair tufts on the eighth abdominal 
segment of the male. Four tropical American species 
are recognized. How many of these are really distinct 
species it is impossible to determine from the scanty ma- 
terial available. Nothing is known of their biology or 
habits. 

84. Pseudodivona commenaella Dyar 
FiaxjRES 41, 204 

Pseudodivona commensella Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 406, 1914. 

Forewing dull white on area above lower margin of 
cell; the area from cell to inner margin a glossy vinous 
brown; a blackish brown smudge on costa at base; 
antemedial line obscure except where it cuts the brown 
shade and forms a contrasting white spot at inner mar- 
gin, bordered outwardly by a blackish brown, out- 
wardly angled line, obscure on all but fresh specimens 
and frequently broken into two blackish spots, one on 
costa, the other in the cell; subterminal line indicated 
by a pair of narrow and narrowly spaced blackish dashes 
from costa near apex and some inwardly bordering 
blackish spots or streaks on the veins; the veins other- 
wise more or less darkly streaked; lower discal spot at 
end of cell enlarged, blackish, more or less confluent 
with a smaller, much fainter upper spot; a row of black- 
ish dots along termen. Hind wing pale, glossy, semi- 
translucent grayish white; veins darkened and clearly 
outlined; a narrow dark line along termen. Alar 
expanse, 20-22 mm. 

Male genitalia with trifling differences in the shape of 
the apical projection of gnathos between this and the 
following species of the genus and some differences in 
the ciurve of the outer margin of the harpe, but I suspect 
that these differences are individual rather than specific 
in character. 

Type locality: Jalapa, Mexico (type in USNM) . 

Food plant: Unknown. 

Distribution: Mexico: Jalapa, Orizaba. 

Known only from the four males of Dyar's type 
series. 

85. Pseudodivona cispba Dyar 

Figure 205 

Pseudodivona cispha Dyar, Ins. Insc. Menstr., vol. 7, p. 53, 1919. 

A smaller, less distinctly marked species than the 
preceding one (commensella); the brownish area of 
forewing narrower and paler (not "reddish" as stated 
by Dyar); the dark markings fewer, fainter and paler 
and, except for the spot on base of costa and a short 
wedge at apex, not blackish; discal dots inconspicuous, 
light brown, the lower dot much smaller than on com- 
mensella. Alar expanse, 16-18 mm. 

Female genitalia like those of P. santa-maria. 

Type locality: Volcdn Santa Maria, Guatemala 
(type in USNM). 

Food plant: Unknown. 

Distribution: Guatemala: Cayuga (Aug.), Volcdn 
Santa Maria (July, Oct.). Costa Eica: Tuis (May). 
British Honduras: Rio Grande (Sept.), Pun to Gorda 
(July). 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



49 



86. Pseudodivona eanta-maria Dyar 

Figure 690 

Pseudodivona sanla-maria Dyar, Ins. Insc. Menstr., vol. 7, p. 54, 
1919. 

Known only from the two females of the type series. 
The coloration and markings are more like those of 
commensella except that the dark striping of the veins is 
fainter and the discal dots smaller, paler and less con- 
spicuous, like those of cispha. It is quite possible that 
these specimens are only larger, darker, better marked, 
female examples of cispha and equally possible that 
both cispha and santa-maria are only varieties of com- 
mensella. Alar expanse, 21 mm. 

Type locality: Volcdn Santa Maria, Guatemala 
(July; type in USNM). Paratype from Cayuga, 
Guatemala (May). 

Food plant: Unknown. 

87. Pseudodivona carabayella Dyar 

Figures 206, 691 

Pseudodivona carabayella Dyar, Ins. Insc. Menstr. vol. 7, p. 54, 
1919. 

Larger and more strikingly marked than any of the 
preceding species. Forewing with pale areas pure 
white; lower area of wing (between cell and inner mar- 
gin) vinous brown (not "purplish red" as in Dyar's 
original description) except for an extension of the 
white behind the antemedial line where it reaches 
almost to inner margin; an elongate black patch on 
costa at base; antemedial line indicated by the usual 
white spot on inner margin and its black outer border, 
the latter is rather broad from costa, strongly angled 
and extends from costa to the lower margin of the cell ; 
a subbasal black spot in the cell; subterminal line indi- 
cated above cell only by its widely spaced black inner 
and outer borders; the inner black border a strong slant- 
ing black dash extending from costa almost to the lower 
discal dot at end of cell (indicating a deep angulation of 
the subterming line); the outer black border a much 
shorter, slanting, dash from apex to vein 6 ; from about 
vein 5 the subterminal line is indicated by a faint white 
line through the brownish ground color and is bordered 
inwardly by a few rather faint blackish spots; discal 
spots, distinct, black and somewhat enlarged, the lower 
one particularly. Hind wing white, more or less tinted 
with grayish on some specimens; veins darkly outlined; 
a narrow dark line along termen. Alar expanse, 23-28 
mm. 

Apical process of gnathos of male genitalia figured 
from type. Another male from Incachaca, Bolivia (in 
BM) , exhibits some variation from the type in the shape 
of the apical process of gnathos (fig. 206a). It is a 
small specimen (23 mm.) and seems to have the dark 
areas and markings of forewing paler, but it is a rubbed 
and faded example; other specimens from the Schaus 
Collection in the National Museum and from the same 
Bolivian locality are typical in all details. The British 
Museum specimen is probably nothing more than an 
individual variant. 



Type locality: Oconeque, Carabaya, Peini (tjrpe in 
USNM). 

Food plant: Unknown. 

Distribution: PertJ: Carabaya, Oconeque, Tinguri. 
Bolivia: CocAa6am6a,1(. Incachaca. Colombia: San 
Antonio (Dec). 

All Peruvian examples in the U. S. National Museum, 
British Museum, and Janse Collection are males. The 
only female of the species that I have seen is the 
specimen from Incachaca, Bolivia, from which the 
genitalia are figured. 

The species is undoubtedly a distinct one. The 
widely spaced, strong, black dashes bordering the sub- 
terminal line indicate this as well as the ciliations of the 
male antenna, which are longer than those of any of 
the preceding species, being somewhat longer than the 
width of the antennal shaft. 

24. Protomoerbes, new genus 

Type of genus: Protomoerbes aberrans, new species. 

Characters of Pseudodivona except: Labial palpus 
upturned (but otherwise as in Pseudodivona) ; forewing 
with vein 3 closely approximate to the stalk of 4-5 at 
base, male with narrow costal fold; hind wing with veins 
4 and 5 stalked for at least three-fourths of their length, 
cell one-third the length of wing; eighth abdominal 
segment of male without hair tufts; transtilla of male 
genitalia incomplete, its elements elongate-angulate, 
their apices not knobbed or expanded. 

In many details this genus is more like Moerbes of 
Group II than Pseudodivona. It differs from both 
genera in its upturned rather than oblique palpi. The 
cilia of the male antenna are also shorter (slightly less 
than the width of the shaft), but this is hardly a generic 
character. Wing pattern, color, and general habitus 
are like those of both Pseudodivona and Moerbes. All 
three have the contrasting white spot on inner margin 
of forewing indicating the base of the antemedial line. 
Protomoerbes in every way seems to be an intermediate 
and connecting link between Pseudodivona and Moerbes. 

It is represented by only two species from Colombia. 
Their females are unknown. 

88. Protomoerbes aberrans, new species 
Figure 208 

Forewing white; basal area, median area below cell 
and outer area below apex shaded with pale brown; a 
yellow longitudinal median streak from base to end of 
cell cutting the antemedial line; antemedial line a con- 
spicuous white spot on inner margin and a fainter white 
spot on costa, bordered outwardly below costa and on 
inner margin by blackish scaling and inwardly by a 
subcostal black streak reaching nearly to base of wing 
and by scattered black dusting at inner margin; veins 
and lower fold beyond antemedial more or less streaked 
or dusted with black, the black streaks especially marked 
and angulate at inner margin of subterminal line; lower 
discal dot expanded and extended along lower vein of 
cell, black; upper discal dot not distinguishable; sub- 



50 



UNITED STATES NATIONAL MUSEUM BXTLLETIN 207 



terminal white line narrow, faint, sinuate and sharply 
dentate below costa, bordered at costa by a pair of 
narrow, short, blackish dashes; a row of separate black 
dots along termen. Hind wing white, translucent. 
The veins faintly darkened and a narrow dark shade 
along termen. Alar expanse, 27-30 mm. 

Male genitalia with aedeagus smooth and evenly 
tapering from base. 

Type locality: Colombia (type in USNM, 61317; 
paratype in BM) . 

Food plant: Unknown. 

Described from male type and two male paratypes 
labeled "Colombia, Fassel." The specimens are only 
in fair condition. Fresher examples would probably 
show the black longitudinal streakings somewhat more 
contrasted and extended. The species has the general 
habitus of Pseudodivona commensella but is larger. It 
is easily separated from commensella and aU the other 
similarly colored and marked species of Pseudodivona 
and Moerbes by the yellow longitudinal median streak 
on forewing. This is easily distinguished under slight 
magnification, even on slightly rubbed specimens. 

89. Protomoerbes separabiKs, new species 
FiGUEB 207 

SimUar to aberrans except: Less distinctly and exten- 
sively streaked with black; median yellow longitudinal 
streak missing from forewing, replaced by a narrow 
extension of the white ground color, extended to the 
base of the wing; aedeagus with a thornlike projection 
from imderside near apex (similar to the projection 
from the aedeagus of Moerbes emendata but larger and 
more blunt) . The male genitalia also differ from those 
of M. emendata in having a proportionally longer and 
more evenly tapering vinculum. Hind wing smoky 
white, semi translucent; the veins distinctly darkened. 
Alar expanse, 29 mm. 

Type locality: San Antonio, Colombia (type inBM; 
paratype in USNM, 61318). 

Food plant: Unknown. 

Described from male type and one male paratype 
from the type locality, labeled "San Antonio, W. 
Colombia, Dec. 07, 5800 feet, M. G. Palmer." The 
paratype lacks an abdomen. 

Genera 25 and 26: Diatomocera and Pseudocabima 

[Venational division C. Forewing with 11 veins; 4 and 5 stalked 
for less than half their lengths. Hind wing with veins 7 and 8 
anastomosed for most of their lengths (free element of 8 short) . 
Transtilla complete but weakly sclerotized. Uncus spoon- or 
semispoon-shaped.] 

25. Genus Diatomocera Ragonot 

Diatomocera Ragonot, Monograph, pt. 1, p. 250, 1893. (Type of 

genus; Homoeosoma tenebricosa Zeller.) 
Cahima Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 329, 1914. 

(Type of genus: Cabima dosia Dyar. New synonymy.) 

Tongue well developed. Antenna of male shortly 
ciliate (the cilia about as long as width of shaft) ; the 



shaft with notch at base; of female simple and pubes- 
cent. Labial palpus of male upturned, reaching to or 
almost to vertex, slender; third segment nearly as long 
as second, acuminate; palpus of female oblique and 
slightly longer than that of male. Maxillary palpus 
filiform. Forewing smooth; 11 veins; vein 2 from weU 
before lower outer angle of cell; 3 from the angle; 4 and 
5 shortly stallved (for half or less than half their lengths) , 
the stalk separated from 3 at base; 6 from below upper 
angle of cell, very slightly bent towards base; 8 and 9 
long stalked; 10 from the cell, but approximate to the 
stalk of 8-9 for some distance; male with an elongate, 
narrow costal fold. Hind wing with vein 2 from well 
before lower outer angle of cell; 3 from the angle, connate 
with the stalk of 4-5; 4 and 5 long stalked; 7 and 8 
anastomosed for most of their lengths beyond cell, free 
element of 8 short ; cell about half the length of the cell 
on male, on female longer; discocellular vein curved. 
Eighth abdominal segment of male with a pair of 
ventrolateral hair tufts. 

Male genitalia with apical process of gnathos U- 
shaped, consisting of a pair of short, blunt arms. Uncus 
spoon- or semispoon-shaped. Transtilla complete but 
weakly sclerotized (except along its lateral edges), con- 
sisting of a broad, more or less finely scobinate plate; 
weakly attached to harpes. Harpe simple; costa sclero- 
tized for most of its length, but not produced. Anellus 
a curved plate with short lateral lobes. Aedeagus stout, 
straight (or but slightly bent near middle), more or less 
tapering to apex, moderately long; penis with a few 
weakly sclerotized wrinklings and more or less finely 
spined, otherwise imarmed. Vinculum stout, elongate, 
constricted towards angulate (or narrowly rounded) 
terminal margin. 

Female genitaHa with bursa more or less finely 
scobinate; signa present, consisting of a cluster of two 
or more sclerotized disks; ductus bursae unsclerotized; 
genital opening simple; ductus seminalis from bursa in 
the neighborhood of the signa (sometimes between them 
and the junction of bursa and ductus bursae, but not 
near the junction). 

The genus is readily recognized by its combination of 
male characters, the most striking feature of which is 
the spoon- or semispoon-shaped uncus which is found 
in only two other American genera — Pseudocabima, 
which lacks the antennal notch and long costal fold of 
forewing, and Entmemacornis, which has veins 4 and 5 
of hind wing completely united. Diatomocera is ap- 
parently confined to tropical America. Nine species 
are here recognized. They are represented by scanty 
and scattered material and nothing is known of their 
life histories or habits. From the greasy condition of 
some of the specimens it may be assumed that the larvae 
are borers. 

90. Diatomocera tenebricosa (Zeller) 
Figures 42, 209, 565, 686 

Homoeosoma tenebricosa Zeller, Horae Soc. Ent. Rossicae, vol. 16, 

p. 242, 1881. 
Diatomocera tenebricosa (Zeller) Ragonot, Monograph, pt. 1, p. 

250, 1893. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



u 



I have not seen any specimens from the type locaHty 
but have before me a male and female from French 
Guiana and a somewhat larger pair (20-21 mm.) from 
Costa Rica compared by Schaus with the type. They 
are either this species or varieties thereof. The follow- 
ing description is drawn from them. 

Ground color of forewing gray shaded with reddish 
brown between the veins; the veins themselves outlined 
with black, the blackish streaks broken by a very faint 
dull whitish antemedial line and by a more distinct 
subterminal line and more or less interrupted between; 
at extreme base the vein markings fused into a blackish 
patch; subterminal line close to outer margin, out- 
wardly angled between the fork of veins 4 and 5 ; discal 
dots small, separated, set obliquely, blackish; terminal 
dots faint. Hind wings grayish brown, paler and semi- 
translucent on the male; the veins faintly darkened, 
brown; a narrow brown line along termen. Alar ex- 
panse, 16-21 mm. 

Male genitalia with uncus but slightly constricted 
near its middle (wider in this area than in any of the 
following species); penis finely spined. Female geni- 
talia with signum a small cluster of bluntly rounded, 
closely appressed disks. 

Type locality: Honda, Colombia (type in BM). 

Food plant: Unknown. 

Distribution: Colombia: Honda. Fbench Guiana: 
Cayenne. Costa Rica: Juan Vinas (May), Sixaola 
River (Mar.). 

The species is easily distinguished from anything else 
in the genus by its smaller size and the shape of its 
imcus. There are several minor differences between the 
genitalia of specimens from French Guiana and Costa 
Rica; in the spacing between the apical prongs of gna- 
thos, in the shape of the terminal margin of vinculum, 
in the amount of spining on the median area of tran- 
stiUa, and in the number of disks forming the female 
signum. These are shown in our figures. Such differ- 
ences, however, do not seem to be of anything more 
than individual or (at most) varietal significance. The 
foldings of the median (membranous) area of the tran- 
stilla shown in figure 209 are superficial and result from 
the manner in which the slide preparation was made. 

91. Diatomocera dosia (Dyar), new combination 

Figures 43, 210, 685 
Cabima dosia Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330, 1914. 
Forewing dull white; the veins streaked with black 
broken into short dashes and dots; antemedial line not 
defined except by the black vein streakings on outer 
margin; subterminal line faint, angulate, the apex of 
angle between the fork of veins 4 and 5 ; discal and ter- 
minal dots distinct, separate, black; a faint shading of 
ocherous fuscous scales above and below vein lb at 
middle; costa at base black for most of the length of 
fold on male. Hind wing duU semitransparent white, 
slightly darker on female; the veins more or less out- 
lined in pale brown; a smoky shade along costa and a 



fine, brown line along termen. Alar expanse, 24-31 
mm, 

Male genitalia mth penis finely granulate-scobinate. 
Female genitalia with signum a chain of blimtly pointed 
disks. 

Type locality: Cabima, Panama (type in USNM). 

Food plant: Unknown. 

Known only from the type series (nine specimens) 
from the type locality (May). 

92. Diatomocera excisalis (Hampson), new combination 
Figures 211, 687 

Crocidomera excisalis Hampson, Ann. Mag. Nat. Hist., ser. 10, 
vol. 4, p. 353, 1929. 

Similar to dosia Dyar and probably no more than a 
variety of that species. Distinguished by its generally 
smaller size, some trifling differences in male genitalia, 
and a shorter chain of disks forming the signum of the 
female. 

Alar expanse, 23-25 mm. 

Male genitalia considerably smaller than those of 
dosia but otherwise similar. 

Type locality: Cayenne, French Gudana (type in 
BM). 

Food plant: Unknown. 

Distribution: French Guiana: Cayenne, St. Lau- 
rent Maroni. 

Also before me is a female from the unidentified ma- 
terial of the British Museum from eastern Bolivia 
("Aug.-Oct., 1920, T. Steinbach") which is superficially 
a very good match for the female paratype of excisalis 
and may be a variety of that species. Unfortunately 
it lacks an abdomen, so positive identification cannot 
be made. 

93. Diatomocera decurrens (Dyar), new combination 
Figure 212 

Cabima decurrens Dyar, Free. U. S. Nat. Mus., vol. 47, p. 330, 
1914. 

Forewing "luteous gray" (the ground color of a dis- 
tinctly yellowish tint); black markings on veins as in 
excisalis but especially strong along vein lb; the sub- 
terminal line somewhat more distinct. Hind wing semi- 
transparent white with a very faint ocherous tint ; veins 
very faintly darkened ; a narrow, pale brown line along 
termen. Alar expanse, 21-28 mm. 

Male genitalia distinguished from those of dosia and 
excisalis by its much narrower (more constricted) 
vinculum; penis with a few weak, minute scobinations. 
The sternite of eighth abdominal segment is also differ- 
ently shaped from that of dosia or excisalis. 

Type locality: Rio Trinidad, Panami (type in 
USNM). 

Food plant: Unknown. 

Distribution: PanamX: Corazal (Mar.), La Chorrera 
(May), Rio Trinidad (Mar.). 

A distinct species easily distinguished by its male gen- 
italia and ocherous forewing. Known only from males 
of the original type series. I fail to see the difference 



52 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



in the antennal notch from that of dosia which Dyar 
mentions. His examination of the material before him 
was obviously cursory for he refers to three of the La 
Chorrera specimens as "females." 

94. Diatomocera majuacula, new species 

FlQUEB 213 

Forewing pale, dull, brownish gray; the veins outlined 
by dark brown; entire basal area to antemedial line suf- 
fused with blackish brown; antemedial line indicated by 
three detached dull-white marks, a spot on costa, a 
smaller one in cell, and a third, somewhat diffused, white 
smudge on inner margin; sub terminal line indicated by 
a white spot on costa, a white spot on inner margin and 
a much fainter, pale, outwardly curved line cutting the 
darkened veins from vein 1 to vein 5; discal spots dis- 
tinct, separated, black. Hind wing white, transparent; 
the veins very slightly darkened; a gray brown shade 
along costa, and a narrow pale-brown line along termen. 
Alar expanse, 32 mm. 

Male genitalia distinguished chiefly by their large 
size. Eighth segment tufts of abdomen also more ro- 
bust than those of any other species of the genus. 

Type locality: Ponta Nova, Eio Xingu, Amazonas, 
Brazil (type in USNM, 61319). 

Food plant: Unknown. 

Described from male type, from the Dognin Collec- 
tion in the U. S. National Museum, that had been iden- 
tified as Sematoneura atrovenosella. The species can be 
easily identified by its large size and the contrasted, 
blackish basal area of forewing. 

95. Diatomocera albosigno, new species 

Figure 214 

Forewing dull ocherous brown; the veins outlined in 
black; broken outwardly by a faint sub terminal line 
which is indicated by a small white spot on costa, a 
smaller, much weaker spot on inner margin and a very 
faint, pale outcurved line between veins 1 and 6; ante- 
medial line replaced by a large white blotch, as broad as 
long and extending from inner margin to or almost to 
costa, bordered outwardly by a narrow blackish line; 
discal spots replaced by a narrow blackish line along 
the discocellular vein. Hind wing dull, smoky white, 
semi translucent; veins faintly darkened; a thin fuscous 
line along termen. Alar expanse, 21-23 mm. 

Type locality: "S. E. Brazil" [probably Parand] 
(type in BM; paratype in USNM, 61320). 

Food plant: Unknown. 

Described from male type and one male paratype 
from the type locality labeled "S. E. Brazil, E. D. Jones, 
1920-303." Easily recognized by the large white spot 
on the subbasal area of forewing; the only species of 
Diatomocera so marked. 

96. Diatomocera hoplidice (Dyar), new combination 

Figure 215 

n 
Cabima hoplidice Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330* 
1914. 



Forewing dark gray; unicolorous except for faint 
darker (blackish) shading on some of the veins and a 
dark edging to the costa; antemedial and sub terminal 
lines obsolete; discal dots replaced by a very faint dark 
line along the discocellular vein. Hind wings very dark, 
smoky gray (almost black) . Alar expanse, 26 mm. 

Male genitalia with aedeagus broad and abruptly 
tapered toward apex. 

Type locality: Puerto Bello, Panamd (Mar.; type 
in USNM). 

Food plant: Unknown. 

Known only from the male type. Easily recognized 
because of its nearly uniform dark coloration and lack 
of any transverse markings on forewing. 

97. Diatomocera extracta, new species 
Figures 217, 688 

Ground color of forewing gray with a faint ocherous 
tint, especially in basal area; veins darkened; antemedial 
line well out, near middle of wing, vertical, bordered 
outwardly by a narrow reddish brown line and in- 
wardly by a fainter line of the same color; subtermina 
line indicated only by breaks in the dark lining of some 
of the veins ; discal spots replaced by an oblique brown 
line along discocellular vein. Hind wing semitranslu- 
cent, white with a very faint grayish ocherous tint, 
slightly darker on female; veins very faintly darkened; 
a fine dark (brownish) line along termen. AJar expanse, 
21-25 mm. 

Male genitalia with harpe narrow, not expanded to- 
ward outer margin; aedeagus slightly bent; penis armed 
with an elongate, dense cluster of slender dark spines 
and a scattering of fine scobiaations. Female genitalia 
with bursa transversely elongate, weakly scloerotized in 
the area about the signa and junction of ductus semin- 
alis, also finely scobinate in this area, the scobinations 
extending into the ductus bursae; signa consisting of 
two or three very narrow, thin, elongate disks. 

Type locality: Tuis, Costa Kica (type in USNM, 
61321). 

Food plant: Unknown. 

Described from male type from the type locality, col- 
lected by W. Schaus ("May 28-June 4"), and one male 
and one female paratypes, collected by Schaus and 
Barnes (Nov.). 

The species is closest to but quite distinct from moch- 
lophleps Dyar. Most nearly resembles pale or faded 
examples of Pseudocahima rubrizonalis (Hampson), with 
which it was confused in the National Collection. 

98. Diatomocera mochlophleps (Dyar), new combination 
Figures 216, 689 

Cabima mochlophleps Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 404, 1914 

Forewing purpHsh gray with faint pale reddish brown 
shading between the veins; the area from base to near 
middle slightly paler, its outer margin faintly indicating 
an antemedial line, somewhat curved and inwardly 
oblique from costa; sub terminal line well inward from 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



53 



termen, angulate with apex of the angle within the fork 
of veins 4-5, appears denticiJate due to pale interrup- 
tions on the veins, preceded by short black dashes on 
the veins; veins otherwise faintly blackish; a distinct 
and characteristic discal mark beginning as a black 
streak or dot at lower outer angle of cell and continued 
as a curved line along discocellular vein and for a short 
distance inward along upper vein of cell ; a row of black 
dots along terminal margin. Hind wing translucent, 
white; the veins darkened, a smoky shade along costa 
and a narrow blackish line along termen. Alar expanse, 
31 mm. 

Male genitalia similar to those of extracta, except api- 
cal part of uncus broader and vinculum stouter. Fe- 
male genitalia with bursa globular; the signa consider- 
ably larger and more triangulate than those of extracta. 

Type locality: Zacualpan, Mexico (tjrpe in USNM). 

Food plant: Unknown. 

In addition to the male type and a female from the 
type locality there is before me a smaller (27 mm.) male 
from the Janse Collection labeled simply "Mexico." It 
is more suffused, lacks the pale basal shade on forewing, 
and shows scarcely any trace of a sub terminal line. The 
black discal mark, however, is present and strongly con- 
trasted. This is the characteristic feature of the species. 
It resembles an inverted comma with the tail pointed 
towards the base of the wing. 

26. Pseudocabima, new genus 

Type op genus: Myelois euzopherella Dyar. 

Characters of Diatomocera except: No notch in shaft 
of male antenna; forewing of male without costal fold; 
hind wing with vein 3 frequently stalked with the stalk 
of veins 4-5 (apparently not a constant specific char- 
acter) . In one species (rubrizonalis) the apical process 
of gnathos differs from that of any other species of either 
Pseudocabima or Diatomocera in that it is developed as a 
flattened hook with cleft apex and not as a U- or V- 
shaped pair of prongs. Such a departure from type is 
unusual within generic limits, but of no more than spe- 
cific significance, because the species otherwise is per- 
fectly normal. 

The new genus is proposed with great reluctance ; for 
its species are very closely related to the bulk of those in 
Diatomocera, though none has been previously associated 
with them; but some separation must be made if we are 
to define our superspecific groups with any exactness. 
Psevdocabima, Diatomocera, and Entmemacornis are all 
obviously closely related, but they are separable on con- 
sistent, if slight, structural differences. 

Ten species are here recognized as belonging to the 
genus. One of these is North American (arizonensis) . 
The remainder are tropical and probably only a fraction 
of the species inhabiting Central and South America. 

Before me are five single examples of what appear to 
be as many new species. It seems advisable to leave 
them undescribed until more material is available, their 
sexes can be associated, and more is known about the 
individual and local ranges of variability within species 



of the genus. There seems to be some variability, both 
in color and in minor details of genitalic structure. 

99. Pseudocabima castronalis, new species 
Figures 218, 696 

Forewing gray, the ground color lightened by white 
dusting over much of the median area; antemedial line 
distinguishable throughout, whitish, nearly vertical, 
slightly out-bent from before middle of costa to middle 
of cell, thence slanting inwardly very slightly to vein lb 
and thence outwardly to the inner margin, followed on 
inner margin by a dark blotch; subterminal line from 
outer fourth of costa, bluntly angulate at middle (the 
line more curved than angled between vein 6 and lower 
fold) ; discal dots replaced by a pale ocherous brown spot 
covering the discocellular vein ; fainter extension of this 
brownish shade between some of the veins in outer area, 
especially near costa beyond and before the subterminal 
line; some blackish streaking on the veins, conspicuous 
as three short black streaks following the brownish dis- 
cal spot; hind wing smoky white, semitranslucent on 
the male; darker, more brownish on female; the veins 
but faintly darkened. Alar expanse, 23-27 mm. 

Male genitalia with aedeagus tapering to narrow 
apical end; penis without spining or scobinations. Fe- 
male genitalia with signum a curved chain of more or 
less bluntly pointed disks. 

Type locality: Castro, Parand, Brazil (type in 
USNM, 61322; paratypes in BM). 

Food plant: Unknown. 

Described from male type from the type locality, and 
one male paratype and one female paratype labeled 
"S. E. Brazil, E. D. Jones, 1920-303," the latter two from 
unplaced material in the British Museum. They are 
somewhat discolored and consequently appear more 
yellowish than the type, which is in better condition, 
showing no trace of grease. This specimen was de- 
posited in the National Museum in 1905 by Schaus as 
representative of a Hampson manuscript species. It 
bears the name label (in Hampson's handwriting): 
"Coptarthria castroalis Hampson, Type cT ." Apparently 
Hampson never published a description. 

The best character for recognition of the species is 
the ocherous-brown discal spot with its outwardly bor- 
dering contrasted black streaks. 

100. Pseudocabima fearnella (Schaus), new combination 

Figure 219 

Myelois fearnella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p. 245, 1913. 

Forewing gray, a reddish brown shade along lower 
fold and some dusting of the same color along inner mar- 
gin; antemedial line obsolete; subterminal line faint but 
distinguishable, bent as in castronalis, pale gray; veins 
discontinuously lined with black; discal spots black, 
separated or (at most) partially fused. Hind wing 
white, semi transparent; the veins slightly darkened; 
a darker more distinct line along termen. Alar expanse, 
19-23 mm. 



54 



tnsriTED STATES NATIONAL MUSEUM BULLETIN 207 



Male genitalia differing only in trifling details from those 
of castronalis, apical part of uncus being slightly broad- 
er and a trace of fine scobination appearing on the 
penis. The female is unknown. 

Type locality: Avengarez, Costa Rica (July; type 
in USNM). 

Food plant: Unknown. 

In addition to the male type there are before me two 
males from Cayuga, Guatemala (Feb., June). They 
are smaller than the type and somewhat rubbed and 
faded and, as a result, considerably paler. However, 
they agree in all other details. The species is evidently 
very close to castronalis. 

101. Pseudocabima guianalis, new species 
FiGUBES 220, 697 

Forewing pale brownish gray; some darker dusting 
for a wide area along inner margin; veins very faintly 
and irregularly outlined by dark scaling thickened at 
lower angle of cell, the curvation of the line inward; 
antemedial line obsolete, very faintly indicated on one 
or two specimens; sub terminal hne obscure, more 
sharply angulate than on preceding species. Hind 
wing translucent white; veins partially and faintly 
darkened ; a pale smoky brown line along termen. Alar 
expanse, 25-33 mm. 

Male genitalia with uncus narrower in middle than 
that of castronalis or fearnella; penis armed with a 
cluster of very fine weak spines and scobiaations. Sig- 
nmn of female genitalia a short cluster of blunt, thorn- 
like disks. 

Type locality: St. Jean Maroni, French Guiana 
(type in USNM, 61323; para types in BM and Cornell 
Univ.). 

Food plant: Unknown. 

Described from male type and one male and two fe- 
male paratypes from the type locality, collected by 
W. Schaus; one female paratype from Eockstone, Es- 
sequebo, British Guiana (Schaus); one male paratype 
from Tumatumari, Potaro River, British Guiana, June 
26, 1927 (Cornell Lot 760, sub. 114); one male and one 
female paratypes from Mackenzie, Demerara River, 
British Guiana, June 21, 22, 1927 (Cornell lot 760, sub. 
102, 104). One of the female paratypes from St. Jean 
Maroni bears a Hampson "cotype" label inscribed 
"Coptarthria guianalis." Evidently another example 
of an undescribed Hampson species. 

102. Pseudocabima euzopherella (Dyar), new combination 
Figures 223, 693 

Myelois euzopherella Dyar, Proo. U. S. Nat. Mus., vol. 47, p. 326, 
1914. 

Forewing gray more or less dusted with reddish brown 
(the specimens inclined to be greasy, giving a distinctly 
brownish shade to the wing) ; transverse lines distinct, 
whitish; antemedial line nearly vertical, a slight bend 
in cell, a narrow dark, outer, bordering line; subterminal 
line with a similar dark inner border, slightly and 
blimtly angled or rounded at vein 5; the distinctive 



mark a round blackish brown smudge, touching outer 
margia of cell, consisting of a black discal dot surround- 
ed by blackish or brown smudges; some broken black 
streaks faintly indicated on the veins. Hind wing pale 
smoky fuscous, sub translucent; veins darkened; a fine 
dark line along termen. Alar expanse, 19-24 mm. 

Male genitalia with central area of uncus very narrow; 
penis armed with numerous fine spines and scobinations. 
Female genitalia exhibiting no distinctive specific char- 
acters ; signum a row of rather stout, thornlike disks. 

Type locality: Rio Trinidad, Panamd (type in 
USNM). 

Food plant: Unknown. 

Distribution: Panama; Cabima (May), Corozal 
(May), Tabermlla, Rio Trinidad (June). 

Vein 3 of hind wing seems to be consistently from the 
stalk of veins 4 and 5, though the amount of stalking 
varies in different specimens. In the forewing, 10 is 
from the cell, approximate to the stalk of 8-9 but not 
from it as stated by Dyar. 

103. Pseudocabima pombra (Dyar), new combination 
Figure 221 

Myelois pombra Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 326, 
1914. 

Forewing pale ocherous gray; transverse lines faint, 
whitish, the antemedial near middle of wing, vertical, 
the subterminal rather well back from termen, slightly 
angled at middle; no discal markings; some scattered 
dark (brownish) dusting on the veins. Hind wing con- 
colorous with forewing, semitranslucent; the veins not 
appreciably darkened. Alar expanse, 17 mm. 

Male genitaha similar to those of euzopherella except 
smaller and stem of uncus more slender, differences of 
doubtful specific value. 

Type locality: Cabima, Panama (May; type in 
USNM). 

Food plant: Unknown. 

Known only from the male type. Evidently close 
to and possibly only a pale suffused variety of euzophe- 
rella. The type is somewhat rubbed. On a fresh 
specimen in better condition the dark shading on the 
veins of forewing presumably would be more apparent. 

104. Pseudocabima nigristrigeDa (Ragonot), new combination 
Figures 222, 699 

Myelois nigristrigella Ragonot, Nouv. Gen., p. 7, 1888; Mono- 
graph, pt. 1, p. 41, 1893. 

Forewing white dusted with blackish, giving it an 
ashy gray color to the naked eye; under magnification 
some rust colored scaling between the veins especially 
above and along inner margin; antemedial line near 
middle of wing, nearly vertical (very shghtly convex), 
thin, white, bordered outwardly by a narrow, strongly 
contrasted, blackish brown line; subterminal line curv- 
ing outward from costa to between veins 4-5, thence 
vertical to inner margin, white, bordered inwardly by a 
narrow blackish brown line; a black line along dis- 
cocellular vein and continued for a short distance along 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



55 



vein 3 ; a faint concentration of the black dusting on the 
other veins. Hind wing semitransparent, more or less 
smoky white ; darker on female than on male ; the veins 
darkened, a broad dark shade along costa and a narrow 
dark line along termen. Alar expanse, 22-24 mm. 

Male genitalia figured from type. On another male 
in the British Museum from the type locality (June) the 
stem of uncus is broader, the terminal part of vinculum 
less constricted and its terminal margin less sharply 
angled. (Another example of the amount of individual 
variation that may be expected within specific limits in 
this genus.) Female genitalia figured from specimen in 
British Museum. The signum consists of thi-ee closely 
grouped, moderately large, bluntly rounded disks. 

Type locality: Rio Negro, Amazonas, Brazil (May; 
type in Paris Mus.). 

Food plant: UnknowTi. 

Known only from the type locality. A distinct 
species easily identified by the contrasted black trans- 
verse lines on the ash gi'ay forewing. 

105. PBeudocabima arizonensis, new species 
Figure 698 

Similar in color and markings to the preceeding 
species (nigristrigella) but darker, the black dusting 
heavier (especially on basal area), giving the wing a 
coarser, more pepper-and-salt appearance; no rust- 
colored scahng between the veins; the antemedial and 
subterminal lines blackish bordered on both sides but 
the borders (especially of antemedial line) much less 
contrasted than in nigristrigella; subterminal line acute- 
ly angled at middle; somewhat expanded black streak- 
ing on veins 2, 3, the stalk of 4, 5, and sometimes 6 for 
a short distance from ceU; terminal dots fused into a 
continuous black line along termen. Hind wing shining 
white on male, pale smoky white on female; veins very 
faintly darkened; a fine dark line along termen. Alar 
expanse, 24-27 mm. 

Male genitalia exhibiting no specific characters. 
Female genitalia with signum a compact cluster of 
numerous, closely appressed disks. 

Type locality: Redington, Ariz, (type in USNM, 
61324; paratypes in Paris Mus., Cornell Univ., Trans- 
vaal Mus. (Janse Coll.), and BM). 

Food plant: Unknown. 

Described from male type and 2 male and 6 female 
paratypes from the type locality; 8 male and 11 female 
paratypes from the Baboquivari Mts., Ariz. (June, 
July, Aug., Sept.) ; and 4 male and 1 female paratypes 
from Mohave County, Ariz. (Aug., Sept.). 

So far, this is the only species of the genus recovered 
from the United States. In general habitus it resembles 
Euzophera nigricantella Ragonot, also from Arizona. 

106. Pseudocabima expunctrix (Dyar and Heinrich), new combina- 
tion 

Figures 224, 692 

Myelois expunctrix Dyar and Heinrich, Proo. Ent. Soc. Washing- 
ton, vol. 31, p. 116, 1929. 



Forewing slate gray; some black scaling on the veins 
and (under magnification) a faint scattering of white 
scales over outer area; antemedial and subterminal 
lines and discal spots lacking; a row of black dots at 
the vein ends along termen. Hind wing semitrans- 
lucent white, a smoky shade at apex, along costa, and 
on the veins (especially of the female, the veins of the 
male wing not appreciably darkened) ; a fine dark line 
along termen. Alar expanse, 22-30 mm. 

Female genitalia with signum a long chain of pointed, 
thornlike disks. 

Type locality: Baia, Brazil (type in USNM). 

Food plant: "Stems of leguminous tree." 

Known only from the type series. The only reared 
species of the genus and the only one without any 
trace of transverse marldngs. One of the male para- 
types proves to be a specimen of Fundella argentina 
Dyar. In the forewing veins 4 and 5 are somewhat 
longer stalked than in other species of the genus except 
perrensiella, the stalking being for a half to slightly 
more than half their lengths. 

107. Pseudocabima perrensiella (Ragonot), new combination 
Figure 695 

Myelois perrensiella Ragonot, Nouv. Gen., p. 8, 1888; Mono- 
graph, pt. 1, p. 52, 1893. 

From Ragonot's description and figure the species 
must be similar to expunctrix except for the presence 
of a distinct pale subterminal line, a trace of an ante- 
medial line, and a thin dark line along discocellular vein. 
Veins 4-5 are "long-stalked," as in expunctrix. Alar 
expanse, 28 mm. 

The female genitalia differ from those of all other 
described species of the genus in having a sharp, par- 
tially sclerotized, deeply wrinkled, and densely scobi- 
nate bend in ductus bursae near its junction with bursa 
copidatrix; the signum consists of a curved band of 
bluntly rounded, closely impacted, thornlike disks. 

Type locality: Goya, Argentina (type in Paris 
Mus.). 

Food plant: Unknown. 

Known only from the female type. 

108. Pseudocabima rubrizonalis (Ilainpson), new combination 
Figures 225, 694 

Crocidomera rubrizonalis Hampson, Ann. Mag. Nat. Hist., ser. 
10, vol. 4, 353, 1929. 

Forewing pale gray; the costal edge and the veins 
beyond cell purplish brown; the basal area suffused 
with some faint dark dusting on the male, considerably 
darker on the female; antemedial line at middle of wing, 
vertical or nearly so, dull white, bordered on inner and 
outer sides by reddish brown bands, the outer one the 
wider and somewhat broadened in cell; subterminal 
line faint, when distinguishable, grayish white, set well 
back from outer margin (the space between it and 
antemedial line correspondingly reduced), sharply out- 
angled, the apex of the angle in the fork of veins 4-5; a 
black line, slightly curved, along discocellular vein; 



56 



XnsriTED STATES NATIONAL MTJSEtTM BULLETIN 207 



terminal dots faint, more or less confluent. Hind wing 
(of male) translucent white, the veins pale brown in 
outer area; a fine brown line along termen. Alar 
expanse, 26-30 mm. 

Male genitalia with apical process of gnathos a 
single, narrow, flattened hook with notched apex. 

Type locality: St. Jean Maroni, French Guiana 
(type in BM). 

Food plant: Unknown. 

Distribution: French Guiana: Cayenne, St. Jean 
Maroni. Brazil: Parand, Taperinha. 

The foregoing description based on three males in 
the National Museum. What I take to be a female from 
Cayenne in the Janse Collection is also before me. It 
differs from the males in having broader forewing, the 
antemedial line outwardly oblique from costa, the 
subterminal line more distinct, and the hind wing a 
glossy brown. The bursa is large, as long as ductus 
bursae; the signuin a single straight line of rather sharp, 
spinelike thorns. 



Genus 27: Hyalospila 

[Venational division D. Forewing with 11 veins; 4 and 5 closely 
approximate for some distance from cell. Hind wing with all 
veins very long; cell less than one-third the length of wing. 
TranstUla complete, developed as a narrow, slightly arched band.] 

27. Genus Hyalospila Ragonot 

Hyalospila Ragonot, Nouv. Gen. p. 11, 1888; Monograph, pt. 1, 
p. 168, 1893. (Type of genus: Hyalospila stictoneurella 
Ragonot.) 

Tongue well developed. Antenna of male simple; 
shaft very weakly pubescent. Labial palpus upcurved, 
reaching above vertex; slender; segment 3 as long as or a 
trifle longer than 2, acuminate. Maxillary palpus sub- 
squamous (scaling more or less dilated). Forewing 
smooth; 11 veins; vein 2 from before lower outer angle 
of cell; 3 from the angle, shortly separated from 4 at 
base; 4 and 5 closely approximate for some distance 
from cell (not stalked as stated by Ragonot); 6 from 
below upper angle of cell, straight; 8 and 9 stalked for 
not more (usually less) than half the length of 8; 10 
from the cell, approximate to the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from before 
lower outer angle of cell; 3 from the angle, closely 
approximate to the stalk of 4-5; 4 and 5 stalked for 
about half their lengths; 7 and 8 contiguous or very 
weakly anastomosed for not over half their lengths 
beyond cell; all veins very long; cell less than one-third 
the length of wing; discocellular vein curved. Eighth 
abdominal segment of male with a pair of ventrolateral 
hair tufts. 

Male genitalia with apical process of gnathos an 
elongate, slender hook with slightly notched apex. 
Uncus triangulate. Transtilla complete, a simple, nar- 
row, slightly arched band. Harpe narrow; costa 
stA-ongly sclerotized throughout and projecting slightly 



at apex; otherwise simple. Anellus a small shield with 
long, slender lateral arms. Aedeagus simple; penis 
armed with numerous fine scobinations or one or more 
clusters of slender spines. 

Female genitalia with signa present, consisting of a 
single cluster of bluntly pointed thorns, frequently 
surrounded by fine scobinations or strongly pigmented 
granulations; bursa various, strongly granulate over 
much of one side, weakly sclerotized towards or at junc- 
tion with bursa, more or less finely scobinate or (except 
for signum patch) smooth; ductus bursae weakly 
sclerotized towards genital opening or with genital 
opening simple; ductus seminalis from bursa near 
junction of bursa and ductus bursae. 

A distinct, easily recognized genus apparently 
limited to tropical America. The species also are easily 
identified by their genitalia. It is difficult to place 
Hyalospila satisfactorily in any linear arrangement for 
it partakes of the characters of two distinct groups. On 
male genitalia, especially its complete bridgelike trans- 
tilla, it should go with the genera of the main Acrobasis- 
Myelois stem having this organ well developed, while 
on other characters, general habitus, and the long veins 
and short cell of hind wing it seems more closely related 
to genera of the Piesmopoda group. 

109. Hyalospila stictoneurella Ragonot 
Figures 44, 226, 703 

Hyalospila stictoneurella Ragonot, Nouv. Gen., p. 11, 1888; 
Monograph, pt.l, p. 169, 1893. 

Forewing purplish brown, a narrow ocherous-white 
band along costa, extending from costa to upper vein of 
cell and showing under magnification a scattered pow- 
dering of reddish scales; the extreme costal edge at its 
middle, darkened; the veins more or less streaked with 
black scaling; antemedial line indistinct, indicated by 
an oblique blackish streak from costa, a small whitish 
dot in ceD and another on vein lb, each followed by a 
black dot; subterminal line faint but distinguishable, 
close to and parallel with termen, not dentate; discal 
dots at end of cell confluent, blackish brown. Hind 
wing semitransparent whitish; the veins brown, a brown 
border along costa and a fine brown line along termen. 
Alar expanse, 19-22 mm. 

Male genitalia with transtilla a short, slightly arched 
bridge, somewhat wider than that of any other known 
species and with a minute, pointed projection at middle. 
Penis armed with three small clusters of fine, short 
spines. Vinculum but slightly longer than greatest 
width, tapering to its rather broad, rounded terminal 
margin. 

Female genitalia with bursa unsclerotized, weakly 
scobinate in the area about the signum; the latter a 
small patch of small, thornlike spines (fig. 703a) ; ductus 
bursae considerably longer than bursa, weakly sclero- 
tized for a short distance from genital opening and with 
a patch of scobinations at its middle, otherwise mem- 
branous and simple. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



Type locality: Las Mercedes, "Amer. centr. mer." 
[Guatemala?] (type in Paris Mus.). 

Food plant: Unknown. 

Distribution: Mexico: Jalapa, Misantla (May), 
Orizaba. Guatemala: Las Mercedes [?], Volcdn Santa 
Maria (Aug., Oct., Nov.). Costa Rica: Juan Vinas 
(Nov.). Brazil: Campo Bello, Santa Catarina (Oct.), 
"S. E. Brazil" [Parand?]. 

110. Hyalospila celiella Schaus 
Figures 227, 700 

Hyalospila celiella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p. 248, 1913. 

Ground color in outer area between the veins and in 
entire area between lower margin of cell and inner mar- 
gin (except for vein lb) pale ocherous brown; a narrow 
pale band along costa (fainter than that on stictoneurella, 
less whitish and, imder magnification, showing a fine 
irroration of purplish brown scales, especially on costal 
edge) ; upper vein of cell white for a short distance from 
subbasal area; lower vein of cell white from base to end 
of cell except for a black spot near middle; a blackish 
brown line bordering the lower edge of cell from base to 
middle; a short white streak on vein lb before middle, 
preceded and followed by blackish dots (these and the 
black spot on lower vein of cell all that remain to indi- 
cate an obsolete antemedial line); vein lb otherwise 
more or less outlined by brown scaling; outer veins 
shortly streaked with blackish brown; sub terminal line 
indicated only by an interruption of the blackish vein 
streaks near outer margin; discal spots a pair of short 
black streaklets at end of cell. Hind wing semihyaline, 
white with a very faint smoky tint in outer area ; veins 
pale brown; a darker brown line along outer margin. 
Alar expanse, 19-21 mm. 

Male genitalia easily distinguished by the stout, 
greatly elongated vinculum. The abdominal tuft and 
sclerotized sternite of eighth segment similar to those 
of stictoneurella. 

Female genitalia with a weak sclerotization of bursa 
in the area surrounding the ductus seminalis and ex- 
tending for a short distance into the ductus bursae; no 
spining at middle of ductus bursae; otherwise as in 
stictoneurella. The sclerotization of the ductus bursae 
at genital opening varies individually in extent and 
amount as it does in stictoneurella. 

Type locality: Juan Vinas, Costa Rica (type in 
USNM). 

Food plant: Unknown. 

Represented in the National Collection by the male 
holotype (Jan.) and two females (June, Nov.), all from 
the type locality. 

111. Hyalospila inseqaens, new species 

Similar to celiella except: Larger; ground color of fore- 
wing darker, some rosy scaling overlaying most of the 
pale area between cell and inner margin; dark streaking 



on veins fainter; hind wing a clearer white, the veins 
not appreciably darkened. Alar expanse, 23-24 mm. 

Male genitalia similar to those of celiella except vin- 
culum not greatly elongated, slightly less than twice as 
long as its greatest width. Female unknown. 

Type locality: Incachaca, Cochabamba, Bolivia 
(type in USNM, 61325; paratype in BM). 

Food plant: Unknown. 

Described from male type, collected by J. Steinbach, 
and one male paratype from San Antonio, western 
Colombia, 5,800 feet, Nov. 1907, M. G. Pabner, 
collector. 

The species has the same dark streak from base of 
forewing along the under edge of the cell as celiella, but 
not so strongly accented. The discal spots are slightly 
more pronounced and more or less confluent. The 
shorter vinculum at once distinguishes it. 

112. Hyalospila majorina, new species 
Figure 701 

Forewing pale gray-brown; costal area to and includ- 
ing the cell dull white, the white shade narrowing 
gradually beyond cell to apex of costa; lower discal dot 
at end of cell enlarged, dark brown, completely en- 
circled by white; antemedial line obsolete; subterminal 
line very faint, close to and parallel with costa, indicated 
chiefly by short, whitish streaklets on a few of the upper 
veins and a slight paling of the ground color from vein 
4 to inner margin. Hind wing semitranslucent, whitish 
with a faint brown tint; the veins darker; a fine brown 
line along termen. Alar expanse, 27 mm. 

Female genitalia with ductus bursae considerably 
broadened for most of its length, wrinkled and weakly 
sclerotized on one side at junction with bursa; bursa 
finely scobinate over part of one side, the scobinations 
extending into ductus ; genital opening simple. 

Type locality: Misantla, Me-xico (type in USNM, 
61326). 

Food plant: Unknown. 

Described from female type, collected by Robert 
Miiller, Sept. 1914 ("4362"), and one female paratype, 
the latter from Jalapa, Mexico. The male is unknown. 
I should not have described a new species from females 
alone, but in this case a male from Mexico should be 
easily matched. The females are readily distinguished 
by their large size and their genitalia. 

113. Hyalospila fulgidula, new species 
Figures 228, 702 

Ground color of forewing a clear bright white, clouded 
by a faint, pale drab shade along inner margin, this 
shade beyond lower outer angle of cell extending ob- 
Hquely upward to apex ; costa at base reddish ; a minute 
red dot on costal edge at one-third and below it a con- 
spicuous, broad, oblique, black dash crosses the cell; 
below this one or two black dots on vein lb; a short, 
black dash along lower margin of cell at base; some few 
scattered black scales on edge of inner margin and on 



800329 — B6 



58 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



some of the veins and a very sparse and scattered dust- 
ing of red scales in the outer white area; discal dots at 
end of cell separate, black, the lower conspicuous, the 
upper minute; subterminal line clearly indicated by a 
row of black dots along its inner border and an out- 
wardly bordering, short, black dash from costa; a row 
of detached black dots from vein 6 to lower fold along 
edge of termen. Hind wing glossy, smoky white, dark- 
ening outwardly and with a brown shade along outer 
margin. Abdominal tufts and eighth segment sternite 
of male as in stietoneurella. Alar expanse, 12-13 mm. 

Male genitaha with transtiUa a very narrow, squarely 
arched band; penis very finely scobinate for about the 
length of aedeagus, otherwise unarmed. Female geni- 
talia with ductus bursae somewhat swoUen and densely 
but minutely scobinate towards its junction with bursa, 
the scobinations extending on one side into bursa; 
ductus binsae also sclerotized for a short distance from 
genital opening, on its dorsal surface the sclerotization 
for min g a shield projecting caudally beyond the opening. 

Type locality: Santiago Province, Cuba (type in 
USNM, 61327). 

Food plant: Unknown. 

Described from male type (June) and three female 
paratypes (June, Sept., Nov.) from the type locality, 
collected by W. Schaus. A distinct species, easily iden- 
tified by its small size, the squarely arched transtiUa, 
and the bright white ground color and contrasted black 
spotting of its forewing. 

114. Hyalospila egenella (Ragonot), new combination 

Figure 704 

Piesmopoda egenella Ragonot, Nouv. Gen., p. 11, 1888; Mono- 
graph, pt. 1, p. 165, 1893. 

Forewing grayish brown finely powdered with reddish 
scales; the costal area white with scattered red scaling; 
antemedial line not distinguishable; subterminal line in- 
dicated chiefly by its slightly darkened inner and outer 
borders, close to and parallel with termen, not sinuate. 
Hind wing smoky white, semitranslucent, the veins 
darkened and a dark line along termen; the smoky tint 
accented somewhat towards apical area. Alar expanse, 
15-20 mm. 

Female genitalia distinguished chiefly by the en- 
larged, wrinkled and scobinate ductus bursae and the 
enlarged blimt thorns forming the signum (fig. 704a); 
ductus bursae weakly and narrowly sclerotized at gen- 
ital opening. The male is unknown. 

Type locality: Rio Negro, Brazil (type in Paris 
Mus.), 

Food plant: Unknown. 

A small (15 mm.), somewhat rubbed female in the 
U. S. National Museum from Santa Catarina, Brazil 
(July), seems to be this species. It has genitalia similar 
to those of Ragonot's type, differing only in minor indi- 
vidual details, a somewhat more extended scobination 
of ductus bursae, and faint traces of pale yellowish 
sclerotization in some of the folds at junction of bursa 



and ductus bursae. It also shows traces of a dark discal 
spot on forewing at lower outer angle of cell which Rag- 
onot states is absent from his type. None of these 
differences is significant. 

115. Hyalospila xanthoudemia (Dyar), new combination 
FiGTJEES 229, 709 

Piesmopoda xanthoudemia Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 333, 1914. 

Forewing olivaceous ocherous; the costal area beyond 
extreme base and including the cell and the outer area 
above vein 6, white with a scattered powdering of red- 
dish scales; extreme costal edge red, this shade espe- 
cially noticeable at base; antemedial line, discal and 
terminal dots obsolete; subterminal line faint but visible, 
whitish, very close to termen, not sinuate. Hind wing 
whitish with a faint ocherous fuscous tint especially in 
outer area, somewhat darker on female than on male; 
veins faintly darkened; a fine, pale brown line along 
termen. Alar expanse, 16-19 mm. 

Male genitalia with penis armed with a band (about 
one-third as long as aedeagus) of fine scobinations. Fe- 
male genitalia with bursa simple except for the signiun 
patch; ductus bursae much longer than bursa, slender 
for most of its length, without scobinations and unsclero- 
tized except very weakly at genital opening. 

Type locality: Rio Trinidad, Panamd (type in 
USNM). 

Food plant: Unknown. 

Distribution: PanamI: ChiriquI (AprU), Paraiso 
(Jan.), Rio Trinidad (Mar.). Costa Rica: Experenza 
(May), Juan Vinas (Oct.). 

Doubtfully distinct from angulineeUa (Schaus). 

116. Hyalospila angulineeUa (Scbaus), new combination 
Figure 707 

Piesmopoda angulineeUa Schaus, Ann. Mag. Nat. Hist., ser. Sj-, 
vol. 11, p. 246, 1913. 

Known only from the female type. Color and mark- 
ings as on xanthoudemia except for faint traces of an 
angulate dark antemedial line on forewing, and darker 
(pale smoky brown) hind wing. 

Ductus bursae of female genitalia longer than that of 
xanthoudemia. Otherwise the ductus shows but trifling 
differences which are somewhat exaggerated in the figiu-e. 

Type locality: Juan Vinas, Costa Rica (June; type 
in USNM). 

Food Plant: Unknown. 

117. Hyalospila clevelandella (Dyar) 
FiGUBES 230, 705, 706 

Oryctometopia clevelandella Dyar, Proc. U. S. Nat, Mus., vol. 47, 

p. 331, 1914. 
Hyalospila clevelandella (Dyar) Dyar, Ins. Insc. Menstr., vol. 7, 

p. 48, 1919. 

Forewing gray-brown from lower margin of cell and 
(in outer area) below vein 5, costal area white sparsely 
irrovated with red scales; extreme base of costal edge 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



^ 



brown; costal edge beyond more or less reddish; ante- 
medial line indicated only by fragments of its outer 
border, a couple of red dots or dashes in the white area 
and a faint, dark, gray-brown dot on lower fold of vein 
lb; discal dots at end of cell weak, lower brown, upper 
red ; subterminal line faint but distinguishable, not sinu- 
ate, close to and parallel with termen, dull white. Hind 
wing smoky white with the veins and lower fold dis- 
tinctly darkened and a dark smoky shade along termen. 
Alar expanse, 13-16 mm. 

Male genitalia with vinculum sharply constricted into 
a digitate projection slightly beyond base; anellus an 
elongate, irregularly shaped, ciu-ved plate with elongate, 
very slender (almost threadlike) lateral lobes; penis 
armed with a single, dense cluster of dark brown, slender 
spines, the cluster as long or nearly as long as aedeagus. 

Female genitalia with ventral sm-face of half of bursa 
and ductus bursae covered with a mat of closely placed, 
pigmented granulations, the granulations extending 
around partly to dorsal surface; signum patch (on dor- 
sum of bursa) surrounded by a teardrop-shaped mass of 
granulations (fig. 705a) ; genital opening simple. 

Type locality: Porto Bello, Panamd (type in 
USNM). 

Food plant: Unknown. 

Distribution: PanamA: Porto BeUo (Dec), Taboga 
Isl. (Feb.), TabogiUa Isl. (Feb.). 

The species can be at once identified by its peculiar 
genitalia. The female paratype has genitalia similar in 
all but the most trifling details to those of semibrunneella 
Ragonot, and if it is actually conspecific with the males 
of clevelandella the latter name will fall as a variety or 
synonym of semibrunneella. However, there is some 
doubt that this is the case ; for we have in the National 
Collection a series of four females from Cayuga, Guate- 
mala, and Jalapa, Mexico, of the same size (16-17 mm.) 
and identical color and maculation as the female para- 
type of clevelandella, but with quite different genitaha 
(fig. 706). Also in the collection are four other females 
with the same color and markings and the same size as 
the males of clevelandella (13-14 mm.) but with differ- 
ent female genitalia. Either of these two groups of 
specimens could be the females of Dyar's species so, for 
obvious reasons, I am not attempting to name them or 
to propose any synonymy. 

118. Hyalospila Bemibnmneella Ragonot 
FiGUBE 708 

Hyalospila semibrunneella Ragonot, Nouv. Gen., p. 12, 1888; 
Monograph, pt. 1, p. 169, 1893. 

Color and maculation similar to those of clevelandella 
except antemedial line more distinct. The female gen- 
itaha agree substantially with those of the female para- 
type of clevelandella. We shall have to wait discovery of 
a male of semibrunneella from the type locality before 
the status of the two supposed species can be determined. 

Type locality: "New Granada" [Colombia] (Mar. ; 
type in Paris Mus.). 

Food plant: Unknown. 



Genus 28: Fundella 

[Venational division B. Forewing with veins 4-5 connate or 
approximate at base. Hindwing with vein 3 from the stalk of 
4-5 or closely approximate to it for some distance; cell short; 
on male anal area folded into a pocket. Male genitalia with uncus 
hammer-clawed (long, curved, constricted at middle and broadly 
divided at apex); transtilla absent; sacculus of harpe not pro- 
duced; cornutus present, a single, strong spine. Eighth abdom- 
inal segment of male with pair of hair tufts.] 

28. Genus Fundella Zeller 

Fundella Zeller, Isis von Oken, 1848, p. 866. — Ragonot, Mono- 
graph, pt. 1, p. 210, 1893. — Janse, Journ. Ent. Soc. South 
Africa, vol. 4, p. 163, 1941.— Heinrich, Proc. U. S. Nat. Mus., 
vol. 96, p. 105, 1945. (Type of genus: Fundella pellucena 
Zeller.) 

Ballovia Dyar, Proc. U. S. Nat. Mus., vol. 44, p. 323, 1913; Ins. 
Insc. Menstr., vol. 7, p. 40, 1919. (Type of genus: Ballovia 
cislipennis Dyar.) 

Tongue well developed. Antenna of male weakly 
pubescent, basal segment somewhat enlarged, shaft 
laterally flattened and very slightly excavate at base 
(fig. 23 le) (except on ignobilis and ahemora) and with a 
very small blackish scale tuft in the excavation (except 
on ignobilis); of female, slender, simple. Front of male 
head deeply grooved to hold labial palpi; of female, 
rounded. Labial palpus upcurved, reaching to vertex, 
clothed with broad appressed scales; on male closely 
appressed to face, with second segment over three times 
as long as first and with third segment very short (about 
one-sixth the length of second); on female with second 
segment shorter and third about one-third the length of 
second. Maxillary palpus minute, filiform. Forewing 
smooth; 11 veins; vein 2 from before lower outer angle 
of cell; 3 from the angle, approximately equidistant 
from 2 and 4; 4 and 5 connate or approximate at base; 
6 from below upper angle of cell, straight; 8 and 9 long 
stalked, 9 short; 10 from the cell, parallel for some dis- 
tance but not closely approximate to the stalk of 8-9. 
Hind wing with vein 2 from close to lower outer angle 
of ceU ; 3 from the stalk of 4-5 or closely approximate to 
it for some distance; 4 and 5 stalked for over half (about 
two-thirds) their lengths; 7 and 8 closely approximate 
beyond ceU for less than half their lengths; cell short, 
about one-third the length of wing; discocellular vein 
curved; on male, anal area (involving veins la and lb) 
thickened and folded under to form a pocket enclosing 
enlarged scales and hair tufts. Eighth abdominal seg- 
ment of male bearing a thin, short pair of ventrolateral 
hair tufts. 

Male genitalia with uncus long, curved, strongly scler- 
otized, constricted at middle and broadly divided at 
apex (hammer-clawed) ;gnathos terminating in a short, 
stout hook or a short, broad plate {ahemora) ; transtilla 
absent. Harpe rather short, with clasper. Vinculum 
narrow, short. Aedeagus stout with long, stout, pro- 
jecting, ciu-ved spine or spines at apex (except in 
argentina) ; cornutus a single, strong spine. 

Female genitalia without signum (pellu^ens) or with 
signum well developed and consisting of a large oval or 
pear-shaped cluster of thomlike spines (argentina, aga- 



60 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



pella), or ciu-ved sclerotized bands armed with stout, 
thornlike spines (ahemora, ignohilis) ; bursa large; ductus 
bursae short, broad (narrowest in agapella); area sur- 
roimding genital opening strongly sclerotized, the dorsal 
sclerotization in the form of a band connected with the 
supporting rods of eighth segment collar, and armed 
with two or four spinelike projections (except in ignohilis 
and some examples of argentina) ; ductus seminalis from 
caudal area of bursa. 

This genus is easily distinguished by its striking male 
characters; the strongly sclerotized, long-stemmed, bi- 
furcate (hammer-clawed) uncus; the large pocket on 
anal area of hind wing; the long, embedded labial palpus 
with very short third segment; and minute maxillary 
palpus. A similar bifurcate uncus is not found in any 
other American genus except Dijundella Dyar. In the 
type species of the latter {corynophora Dyar) the uncus 
is somewhat produced and exhibits a slight bifurcation 
at apex; but other species, which must also be referred 
to Dijundella, lack this character. Dijundella separates 
readily on other male structures — its strongly hooked, 
partially free sacculus of harpe, its rounded irons, and 
the narrow, strongly sclerotized, deeply invaginated 
pocket of the sternite of the eighth abdominal segment. 

In Fundella the wing pattern varies so much within 
any given species that it affords no reliable character 
for specific identification, and the several species can be 
separated with certainty only by their genitalia. 

119. Fundella pellucens Zeller 
Figures 6, 231, 713 

Fundella pellucens Zeller, Isis von Oken, vol. 41, p. 866, 1848; 
Horae Soc. Ent. Rossicae, vol. 15, p. 236, 1881. — Ragonot, 
Monograph, pt. 1, p. 210, 1893.— Heinrich, Proc. U. S. Nat. 
Mus., vol. 96, p. 107, 1945. 

Ballovia cistipennis Dyar, Proc. U. S. Nat. Mus., vol. 44, p. 323, 
1913. 

Fundella cistipennis (Dyar) Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 
1919. — Wolcott, Journ. Agr. Univ. Puerto Rico, vol. 17, 
pp. 241-255, 1933; Journ. Agr. Univ. Puerto Rico, vol. 18, 
p. 432, 1934; vol. 20, p. 477, 1936.— Scott, Journ. Agr. Univ. 
Puerto Rico, vol. 24, pp. 35-47, 1940. 

Male antennal shaft with very small black basal tuft 
(fig. 23 le). Forewing grayish fuscous more or less dusted 
with whitish and with interspersed reddish brown scales 
(in many specimens the ground color is reddish brown), 
giving the moth a distinctly gray or gray-brown appear- 
ance to the naked eye; a conspicuous, round, darker 
brown or fuscous spot in the center of the area usually 
occupied by the antemedian line, this dark spot more or 
less obscured in some specimens, but in typical examples 
outlined by whitish areas inwardly and outwardly and 
not reaching to inner margin or costa of the wing; discal 
mark at end of cell obscure, often absent; sub terminal 
line (when distinguishable) faint, white, indented at 
vein 6 and at submedian fold; a row of dark spots along 
termen (present only in specimens having an appreciable 
dusting of white scales). Hind wing white, translucent, 
a faint fuscous border along costa and (in some speci- 



mens) a fuscous line on termen for a short distance from 
apex; cUia white; anal pocket yellowish white. Mid- 
tibia with a fringe of pale hairlike scales along dorsum. 
Hind tibia with a rather long and slender tuft of pale 
(whitish ocherous), hairlike scales from the knee joint 
(fig. 231f). 

Female essentially like the male in color and markings 
except that the dark spot near the base of the forewing 
is more diffused, sometimes reaching to the costa. Hind 
wing usually with a dark shade along termen. 

Alar expanse, 19-24 mm. 

Male genitalia with a large, strongly sclerotized 
subanal plate, constricted before and beyond its middle. 
Harpe with apex notched below costa; clasper short, 
curved, situated near middle of harpe, and armed with 
several setae at its knobbed apex. Aedeagus with a 
cluster of several long, curved spines from apex; cor- 
nutus long, straight, stout. 

Female genitalia with bm-sa copulatrix finely scobi- 
nate but without signum; ductus bursae flattened, 
broad, twisted and constricted near genital opening, 
sclerotized throughout, the sclerotization involving 
bursa adjacent to ductus bursae and ductus seminalis; 
sclerotized band behind genital opening armed with 
four long, stout, projecting spines; coUar of eighth seg- 
ment invaginated at dorsal margin to form a sclerotized 
pocket (fig. 713a). 

Type localities: St. Thomas, British West Indies 
{pellucens, in BM); Barbados {cistipennis, in USNM). 

Food plants: Vigna unguiculata (cowpeas, black- 
eyed peas, and garden peas), Bauhinia variagata, Can- 
avalia ensiformis (swordbeans), Canavalia maritima 
(black bean), Cajan eajan (pigeon pea), Phaseolus luna- 
tus (cultivated and wild limabeans), Phaseolus sp. 
(Brazilian specimens), Cassia ocddentalia (one reared 
specimen from McCubbins MiUs, Puerto Rico, before 
me; most records from this last plant are doubtful and 
probably the result of a misidentification of Fundella 
argentina as cistipennis). 

According to Scott the favored host in Puerto Rico is 
the cowpea {Vigna unguiculata), and the species, while 
frequent in limabeans, seldom does serious damage. 
Potentially it is an insect of economic importance. The 
larvae are primarily pod borers, but also bore into the 
stems and feed on the flowers of their hosts. They 
attack, as far as known, only Leguminosae. 

Distribution: United States: Florida, Hobe Sound 
(May), Miami, (Apr., May), Jupiter (Apr.), Coconut 
Grove, Marco Isl., Tampa (Mar.), Walton, Jensen 
(U. S. Dep. Agr. rearings from limabeans, Feb. 
1944), Riviera Beach, Vero Beach (J. R. Malloch, Dec. 
1941). Barbados. Haiti: Damien (Dec, Feb.), 
Port-au-Prince. Montserrat (Jan.). Cuba: Santiago, 
Matanzas. Virgin Islands: St. Croix (Mar., Oct., 
Nov.). Puerto Rico: San Juan, Rio Piedras (Mar.- 
May), Isabella, Catano (July), Vieques Isl. (Apr.). 
Brazil: Bala (May), Ceard. Bolivia. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



61 



120. Fundella argentina Dyar 
Figures 234, 711 

Fundella pellucens Zeller (in part, "var. b"), Isis von Oken, vol. 

41, p. 867, 1848; Horae Soc. Ent. Rossicae, vol. 16, p. 237, 

fig. 41b, 1881. 
Fundella argentina Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 1919. — 

Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. 109, 1945. 
Fundella eucasis Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 1919. 

Male antennal shaft with even smaller black basal 
scale tuft than that of pellucens. Forewing gray with- 
out the reddish brown, interspersed scaling character- 
istic of typical examples of pellucens; entire basal area 
to antemedian line dark fuscous gray (with but very 
slight dusting of whitish scales toward base in some 
specimens) ; this dark basal patch contrasted against 
the paler gray color of the remainder of the wing, ex- 
tending from costa to inner margin and bordered out- 
wardly by a narrow whitish line. Otherwise not dis- 
tinguishable, superficially, from pellucens. 

Female essentially like the male in color and markings 
except that the basal area of forewing is concolorous 
with or contrastingly paler than the remainder of the 
wing. A narrow dark line or a diffused dark shading 
outwardly bordering the obscure antemedian line. 

Alar expanse, 15-23 mm. 

Male genitalia without sclerotized subanal plate. 
Terminal projection of gnathos varying from round to 
pointed (fig. 234b) at apex. Harpe tapering to bluntly 
pointed apex; clasper a single, straight, sUghtly rough- 
ened, appressed spine, situated beyond middle of harpe. 
Aedeagus simple; cornutus a single, straight spine. 

Female genitalia with signima well developed and 
consisting of a large pear-shaped cluster of thornlike 
spines; sclerotized band behind genital opening, divided 
in the middle, simple (fig. 711a) in Argentinian and 
Brazilian specimens, or armed with a pair of median, 
spinelike projections (fig. 711), rather long in West In- 
dian specimens or short and disappearing in Mexican 
and Venezuelan specimens. 

Type localities: Tucumdn, Argentina (argentina, in 
USNM); Caracas, Venezuela {eucasis, in USNM). 

Food plant: Cassia spp. (reared examples in Na- 
tional Collection from Cassia bicapsularis and C. corym- 
hosa), Poinciana gilliesi. 

Distribution: United States: Florida, Biscayne 
Bay (May), Coconut Grove (Apr.), Stock Island (Apr.); 
Texas, Brownsville (Nov.). Mexico: Several examples 
reared from pods and blossoms of Cassia bicapsularis 
at Brownsville, Tex., quarantine station. Cuba: Bara- 
guA (Mar.), Habana, Matanzas, Santiago Province. 
Puerto Eico: Bayam6n (Mar., Sept.), Vieques Isl. 
(Apr., July), Coamo Springs (Apr.), Aguirre Central 
(Aug.), San Germdn (Aug.), San Juan (Nov.). Haiti: 
Potion viUe (Jime). Virgin Islands: St. Croix (Oct.- 
Nov.). Jamaica. Venezuela: El Valle (June). 
Brazil: Bafa (May). Argentina: Tucumdn (Mar.). 

In collections this species has appeared most fre- 
quently under the name pellucens. Both argentina and 
pellucens have about the same distribution and are 
abundant in the West Indies, though, from material 



at hand, pellucens seems to be rarer on the mainland. 
Throughout its range argentina shows considerable var- 
iation in female genitalia. West Indian specimens have 
rather conspicuous spinelike extensions of the sclero- 
tized band behind the genital opening. These are en- 
tirely lacking in Brazilian specimens, and if one had 
only these extremes he would be justified in assuming 
that they were at least racially distinct. However, 
Venezuelan and Mexican examples show an intermedi- 
ate form with very short projections, and Central Amer- 
ican specimens, when recovered in sufficient numbers, 
will probably show all intergradations. The male geni- 
talia are remarkably uniform throughout the range of 
the species, exhibiting only minor individual variations 
in the shape of the terminal projection of the gnathos. 

121. Fundella agapella Schaus 

Figure 710 

Fundella agapella Schaus, Zoologica, vol. 5, No. 2, p. 47, 1923. — 
Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. Ill, 1945. 

Female palpi, head, thorax, and forewing whitish 
gray; dark markings drab gray; transverse antemedian 
line of forewing white, defined chiefly bjj^ its narrow, 
dark outer border, sharply sinuate, indented a trifle 
just below costa, more deeply at top of cell and still 
more deeply at fold below cell; discal dot at end of cell 
obscure; white sub terminal line indented at vein 6 and 
at submedian fold, bordered inwardly by a distinct dark 
shade as broad as the white line itself and outwardly 
by a similar, fainter, dark shading, the latter conspic- 
uous only at apex. Hind wing as in the other species 
of Fundella. Alar expanse, 12 mm; 

Genitalia like those of intermediate examples of ar- 
gentina except that the signum is considerably smaller 
in proportion to the size of the bursa. 

Type locality: Tagus Cove, Albemarle, Galdpagos 
Islands (type in USNM). 

Food plant: Unknown. 

Known only from the female type. Superficially a 
distinct species. The female genitalia, however, would 
indicate that agapella is only a race of argentina. A 
male will be needed for exact placement, and until it is 
available we shall have to treat agapella as a species. 

122. Fundella ignobilis Heinrich 
Figures 232, 712 

Fundella ignobilis Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. 
112, 1946. 

Male antennal shaft without any trace of black basal 
scale tuft. Otherwise partaking of the pattern mark- 
ings of both pellucens and argentina; in some specimens 
dark basal patch of forewing roimd and reaching neither 
costa nor inner margin (as in typical pellucens) , in ma- 
jority of specimens, however, basal patch occupying 
whole basal area (as in typical argentina) ; median and 
outer areas of wing averaging a trifle paler than in 
argentina and without the reddish brown scaling of 
pellucens. 



62 



XHSriTED STATES NATIONAL MUSEUM BULLETIN 207 



Female superficially similar to argentina except a 
trifle paler, on the average. 

Alar expanse, 13-22 mm. 

Male genitalia with gnathos terminating in a short, 
stout hook. Harpe with apex truncate; clasper mod- 
erately long, curved, and weakly haired at apex. Ae- 
deagus with a single, long, strong, curved spine from 
below apex; cornutus a short, stout, curved thorn. 

Female genitalia without spines adjacent to genital 
opening. Bursa copulatrix with signa consisting of a 
pair of partially fused bands, each armed with a row 
of short, stout, thornlike spines; ductus bursae short 
and broad, with median area unsclerotized; eighth seg- 
ment collar completely sclerotized except for a small, 
round, transparent spot on midventer, sclerotLzation 
extending to and over area behind genital opening. 

Type locality: Oaxaca, Mexico (type in USNM) . 

Food plant: Unknown. 

Distribution: M:fixico: Cdrdoba (May), Guadala- 
jara, Jalapa, Oaxaca, Orizaba, Tehuacdn (May, June, 
July). Guatemala: Cayuga. Costa Rica. Cuba: 
Santiago (June), Sierra Miestra (May). Puerto Rico: 
Aguirre Central (Aug.). Haiti: P^tionville (June). 

123. Fundella ahemora Djar 
Figures 233, 714 

Fundella ahemora Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 403, 
1914.— Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. 113, 1945. 

Antenna of male with small black scale tuft at base 
of shaft. Forewing with no or a very faint dark basal 
patch (when present covering basal area to antemedian 
line); antemedian line whitish, very faint; sub terminal 
line white, faint but less obscure than antemedian, 
without dark borders except for an inner and an outer 
dark spot at inner margin of wing; veins from cell rather 
strongly outlined by dark scaling (the most conspicuous 
superficial character of the species). A thick, dark 
(brownish) hair tuft covering outer surface of male 
foretibia, a character not found in other species of the 
genus. Alar expanse, 18-23 mm. 

Male genitalia with gnathos terminating in a broad 
tongue-like plate. Harpe somewhat tapering but with 
apex truncate; a strong tuft of long scales from costa; 
clasper long, curved, slender, with a few hairs at apex. 
Aedeagus with a pair of long, curved, flattened spines 
from apex; cornutus a long, straight, slender spine. 

Female genitalia with a pair of long, widely spaced, 
basally curved spines from sclerotized area immediately 
behind genital opening. Bursa copulatrix with signa 
consisting of two rather short bands, each armed with 
a row of long spines. Ductus bursae bulged in the 
middle and with a strongly sclerotized median collar. 
CoUar of eighth segment partially sclerotized and fused 
ventrally. 

Type locality. — Orizaba, Mexico (type in USNM). 

Food plant. — Unknown. 

Distribution: M:6xico: Orizaba, Jalapa, Teapa 
(Dec), C6rdoba (Apr., Dec), Cuernavaca (July). 



Guatemala: Quirigud (Mar.), Cayuga (Jan., May), 
Parulhd (July). Costa Rica: Juan Vinas (Nov.). 

Superficially the most easily distinguished species in 
the genus. The large foretibial tuft at once identifies 
the male, and both sexes can be separated by the rather 
conspicuous dark outlining of the veins. The veins are 
similarly dark scaled in the other species, but the con- 
trast of the dark veins against the pale intervenular area 
is more marked in ahemora. 

Genus 29: Difundella 

[Venational division B. Forewing with veins 4-5 connate or 
closely approximate at base. Hind wing with vein 3 from stalk 
of 4-5 or closely approximate to it for a short distance; on male, 
anal area folded into a pocket. Male genitalia without transtiUa; 
sacculus of harpe strongly sclerotized and produced (free or 
partially free); penis without cornutus or other armature. 
Abdomen of male with lateral pockets and hair tufts between 
segments 2 and 3; eighth abdominal sternite developed as a 
narrow pocket; no hair tufts.] 

This genus shows affinities to Fundella but in many 
characters resembles more closely Bampylla and Cop- 
tarthria. In general habitus (wing pattern, color, and 
maculation) the moths of Difundella, Coptarthria, and 
Bampylla are strikingly similar, but the three genera 
are different on structural characters. Bampylla differs 
from the other two in the free length of vein 3 of 
male hind wing; Coptarthria in its notched male antenna; 
and Difundella in its anellus (a simple plate without the 
long free spine of Bampylla and Coptarthria) and in the 
possession of scaled pockets between the second and 
third segments of the male abdomen. 

The species of Difundella differ considerably from 
each other on structural details, falling into two distinct 
groups which divide as follows: 

— Labial palpus reaching above vertex in both sexes. Hind 
wing with cell less than one-third the length of wing; 
vein lb of male bent before middle and with a tuft of 
yellow hairlike scales on its under side (within the bend) . 
Gnathos greatly reduced, its apical projection fine, 
needlelike. Costa of harpe without projections. 

— Labial palpus not reaching vertex on males, barely reaching 
vertex on females. Hind wing with cell more than 
one-third (but less than half) the length of wing; vein 
lb of male not bent; rough sex-scaling bordering Ic on 
under side of wing beyond base. Gnathos with apical 
process enlarged and strongly sclerotized. Costa of 
harpe with strongly sclerotized projection or projections. 

The second group probably deserves a separate generic 
designation; but the material before me representing its 
two species is too scanty and not in good enough con- 
dition, and the association of the females with their 
proper males too uncertain, to permit proper evalua- 
tion of generic characters for separation at this time. 

29. Genus Difundella Dyar 

Difundella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 327, 1914. 
(Type of genus: Difundella corynophora Dyar.) 

Tongue well developed. Antenna of male weakly 
pubescent. Labial palpus ascending, recurved, slender, 
smooth scaled; third segment acuminate. Maxillary 



AMERICAN MOTHS OF THE STJBFAMILT PHTCITESTAE 



63 



palpus with second segment slightly thickened with 
scales. Forewing smooth ; 1 1 veins ; vein 2 from before 
lower outer angle of cell; 3 from the angle, nearly 
equidistant from 2 and 4; 4 and 5 connate or closely 
approximate at base and approximate for a short dis- 
tance beyond base; 6 from below upper angle of cell, 
sUghtly bent towards base; 10 from the cell, more or less 
approximate to the stalk of 8-9. Hind wing with vein 
2 from close to lower outer angle of cell; 3 from the 
stalk of 4-5 or closely approximate with it for a short 
distance from angle of cell ; 4 and 5 long stalked (for over 
one-half their lengths); 7 and 8 closely approximate 
beyond cell, or shortly and weakly anastomosed; cell 
short, less than one-half the wing length ; discocellular 
vein curved, partially obsolescent; on male, anal angle 
folded under to form a pocket enclosing a long hair- 
pencil. Eighth abdominal segment of male with sternite 
developed as a narrow, sclerotized pocket ; on each side 
between abdominal segments 2 and 3 a shallow pocket 
containing a modified scale tuft. 

Male genitalia with uncus stout, but variously shaped. 
Transtilla absent. Harpe with sacculus very strongly 
sclerotized, free or partially free and curved. Aedeagus 
with strongly sclerotized and pointed apex; penis 
imarmed. Vinculum stout. A long hair tuft from 
intersegmental area adjacent to base of sacculus of 
harpe. 

Female genitalia with bursa copulatrix membranous; 
sigmmi, if present, a small patch of weak scobinations; 
ductus bursae membranous (unsclerotized) except about 
genital opening; ductus seminalis from bursa near 
junction of biu-sa and ductus bursae. 

The characteristic pattern features of the species of 
Difundella (as of Coptarthria and most species of 
Rampylla) are: The strongly contrasted, fine, blackish, 
transverse lines forming the outer border of the ante- 
medial and the inner border of subterminal lines of 
forewing; the almost straight, oblique or vertical ante- 
medial line, set well out from base of wing; the oval, 
pale discal spot covering the discocellular vein; and the 
black streaks on veins 2 to 6 just beyond it. 

Genus Difundella, Species 124 and 125: D. cory- 
nophora and D. subsutella 

[Labial palpus reaching above vertex in both sexes. Hind wing 
with cell less than one-third the length of wing; vein lb of male 
bent before middle and with a tuft of yellow hairlike scales on its 
under side (within the bend). Gnathose greatly reduced, its 
apical projection fine, needlelike. Costa of harpe without 
projections.] 

124. Difundella corynophora Dyar 

Figures 7, 235, 715 

Difundella corynophora Dyar, Proc. U. S. Nat. Mus., vol. 47' 
p. 327, 1914. 

Forewing with basal area (to antemedial line), and 
upper median area beyond antemedial line and from 
lower margin of cell to costa, blackish fuscous; some 



extension of this dark shade extends narrowly to inner 
margin along the outer dark border of the antemedial 
line ; ground color of remainder of wing a ruddy ocherous ; 
some extension of this ocherous shade invades the dark 
basal area along the lower fold and forms the centers of 
the transverse lines ; beyond the cell the ocherous shade 
is broken by black streaks on veins 2 to 6 and beyond 
subterminal line it is more or less clouded by blackish 
fuscous; discal spot ruddy ocherous, covering disco- 
cellular vein; antemedial line oblique; its narrow dark 
borders black shading to reddish; the dark borders of 
subterminal line also more reddish brown than black; 
all the dark transverse lines less contrasted and con- 
spicuous than those of other species in the genus. Hind 
wing dark smoky fuscous; the veins and terminal edge 
black and (on the female) a blackish shade at apex. 
Alar expanse, 15-16 mm. 

Male genitalia with uncus produced and slightly 
bifurcate at apex. Free projection of sacculus curved 
back towards lower margin of harpe. Aedeagus forked ; 
the longer element of the fork spined at apex. Female 
genitaha with signum a small round patch; genital 
opening smrounded by an oblong, strongly sclerotized 
plate ; in intersegmental area behind this plate a pair of 
ventrolateral pockets (fig. 715b). 

Type locality: La Chorrera, Panamfi (tjrpe in 
USNM). 

Food plant: Unknown. 

Distribution: Guatemala: Cayuga (Aug.), Chejel 
(June). PanamA: La Chorrera. French Guiana: 
Cayenne. 

EasUy identified by its genitalia. 

125. Difundella subsutella (Schaus), new combination 
Figure 236 

Ulophora subsutella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 

p. 248, 1913. 
Rampylla subsutella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7, 

p. 84, 1919. 

The only representative of this species is the male 
type which is somewhat rubbed and the markings con- 
sequently obscured. It differs from corynophora in 
having the pale ground color of the outer areas of fore- 
wing more reddish than ocherous; the black outer 
margin of antemedial Une more sharply defined, black 
throughout, outwardly obUque to lower fold and thence 
slightly curved inward to lower margin. Hind wing 
semihyaline wliite with a faint brownish ocherous tint; 
veins not appreciably darkened; a fine brown line along 
termen. Alar expanse, 17.5 mm. 

Male genitalia with imcus triangulate. Gnathos 
reduced even more than that of corynophora, the lateral 
arms represented by mere stubs. Projecting part of 
sacculus curving away from harpe. Aedeagus sickle 
shaped; its apical haK sharply curved, very strongly 
sclerotized, and tapering to a sharp point. 

Type locality: Juan Vinas, Costa Rica (Jan. ; type 
in USNM). 

Food plant: Unknown. 



64 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



Genus Difundella, Species 126 and 127: D. dis- 
tractor and D. tolerata 

[Labial palpus not reaching vertex on males, barely reaching 
vertex on females. Hind wing with cell more than one-third 
(but less than half) the length of wing; vein lb of male not bent; 
rough sex-scaUng bordering Ic on underside of wing beyond base. 
Gnathos with apical process enlarged and strongly sclerotized. 
Costa of harpe with strongly sclerotized projection or projections.] 

126. Difundella distractor, new species 

Figures 237, 716 

Dark areas of forewing dark brownish gray; the pale 
outer areas dull whitish; antemedial line well out 
towards middle of wing, nearly vertical, slightly notched 
at vein lb, red-brown with a fine black line along its 
outer edge and preceded by a rather broad whitish 
blotch, extending from just below costa, nearly to inner 
margin; a short black streak along lower fold for a short 
distance from base of wing; the discal spot whitish; 
blackish lining on the veins beyond cell very weak; 
subterminal line very sHghtly bent between veins 4 
and 6 otherwise nearly vertical, outwardly bordered by 
a naiTOW shade of the dark ground color and inwardly 
by a fine black line. Hind wing semihyaline white; a 
blackish brown line along termen; the veins not dark- 
ened; the male wing on the imdersurface rather loosely 
and coarsely covered with yellowish scales, especially 
along the veins. Alar expanse, 14.5-15.5 mm. 

Male genitalia with imcus semispoon-shaped. Apical 
projection of gnathos large, triangulate, strongly sclero- 
tized. Harpe with sub triangulate cucuUus; a single, 
long, stout, flat, cm-ved, tapering projection from mid- 
costa. Aedeagus long, slender, smooth, tapering to a 
point. Vinculum about twice as long as greatest 
width. Female genitalia without signum; genital 
plate large, medially notched and supported by strongly 
sclerotized, involuted seventh abdominal segment. 

Type locality: Palmas Abajas, Puerto Rico (type 
in Cornell Univ., paratype in USNM, 61328). 

Food plant: Unknown. 

Described from male type labeled "Palmas Abajas 
(near Guayama), P. R., 1900 ft., [date unreadable], 
W. A. Hofl^man" ; and one female paratype from Aguirre 
Central, Puerto Rico, "Apr. 2-3, 31," M. D. Leonard, 
collector. The female is in good condition and served 
for the foregoing color and pattern description. The 
male is badly rubbed and has the palpi and antennae 
broken off; but enough of the markings remain to show 
that they were the same as those of the female. 

127. Difundella tolerata, new species 

FiGTJBBS 238, 717 

Similar to the foregoing species (distractor) except 
that pale areas of forewing are much more restricted, 
limited to an irregular area bordering inner margin of 
subterminal line (extending back to cell above and 
nearly to antemedial line below); some diffused pale 
shading along the lower fold in basal area; the pale 
areas very duU ocherous white, not as well contrasted 
as in distractor; discal spot ocherous. Hind wing trans- 



lucent, white with a very faint ocherous tint on male; 
a faint dark line along termen. Alar expanse, 19 mm. 

Male genitalia with uncus strongly sclerotized and 
stout (longer than basal width), broadened and rounded 
towards apex. Apical projection of gnathos a long, 
broad, strongly sclerotized, obliquely bent band. 
Harpe with narrow, spatulate cucullus; two projections 
from costa, the first a double thornlike projection from 
near middle, the second a rather slender spine from 
outer third. Aedeagus with a couple of short spines 
at apex. Vinculum no longer than greatest width. 

Female genitalia with signum a narrow, elongate 
patch of scobinations; genital opening simple; ductus 
bursae about three times as long as bursa. From 
the dorsocaudal margin of seventh segment an in- 
vaginated, sclerotized shield supports at each lateral 
margin a short, blunt, weakly pigmented hornlike 
process. 

Type locality: East Bolivia (type in BM). 

Food plant: Unknown. 

Described from two specimens from the British Mu- 
seum Collection labeled "Ost Bohvia, Aug.-Oct., 1920, 
T. Steinbach," the male t3rpe and a female matching the 
male in size, color, and markings. I do not make this 
female a paratype (although the foregoing description 
of the female genitalia is made from it) because there is 
a slight doubt that it is the true female of the species. 
Its genitalia are similar in all but the most trifling details 
to those of a series of females from Cajniga, Guatemala, 
associated in our collection with males of Coptarthria 
dasypyga. One or the other of our identifications of 
females (or both of them) may be in error. We shall 
not know imtil more material is available. 

Genera 30-33: Coptarthria to Dasypyga 

[Venational division B. Male genitalia with transtilla a sinuate, 
sclerotized, more or less scobinate band involved with gnathos; 
a long, free spine associated with anellus. Eighth abdominal 
segment of male with sternite developed as a narrow sclerotized 
pocket.] 

30. Genus Coptarthria Ragonot 

Coptarthria Ragonot, Monograph, pt. 1, p. 251, 1893. (Type of 
genus: Myelois dasypyga Zeller.) 

Tongue well developed. Antenna of male with basal 
segment of shaft considerably elongated, flattened and 
deeply notched at outer extremity; shaft pubescent. 
Labial palpus upturned, slender, barely reaching vertex; 
third segment about half the length of second, bluntly 
pointed. MaxiUary palpus filiform. Forewing smooth; 
11 veins; vein 2 from before lower outer angle of cell; 3 
from the angle, closer to 4 at base than to 2; 4 and 5 
short stalked or connate and closely approximate for a 
short distance from base; 6 from below upper angle of 
ceU, bent towards base; 10 from the cell; male without 
costal fold. Hind wing with vein 2 from close to lower 
outer angle of cell; 3 from middle of stalk of 4-5, or 
closely approximate to it; 4 and 5 stalked for slightly 
more than half their lengths; 7 and 8 closely approxi- 
mate beyond cell; ceU short, about one-third the length 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



of wing; discocellular vein curved. Eighth abdominal 
segment of male with sternite developed as a narrow, 
sclerotized pocket. 

Male genitalia with uncus moderately stout; deeply 
concaved apically (probably only a specific character). 
Transtilla a sinuate, sclerotized band involved with and 
fusing into gnathos. Gnathos proper identifiable only 
by its rather weak lateral arms. Harpe with apex of 
cucullus slightly hooked; sacculus simple, not produced. 
Anellus a small plate with greatly reduced lateral lobes, 
the latter indicated chiefly by their short hair tufts; 
dependent from near base of anellus plate and associated 
with it a long, slender U-shaped band supporting from 
the bottom of the U a long, strongly sclerotized, free 
spine, the latter lying dorsad of the aedeagus. Aedeagus 
small, simple ; penis with a few weak scobinations, other- 
wise unarmed. 

The genus is distinguished from its nearest allies by 
the notched shaft of its male antenna. Female char- 
acters could not be included in the foregoing descrip- 
tion, as examples of this sex have not been satisfactorily 
associated with males of the type species, the only 
known representative of the genus. 

128. Coptarthria dagypyga (Zeller) 
Figures 10, 239, (?) 718 

Myelois dasypyga Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 215, 

1881. 
Coptarthria dasypyga (Zeller) Ragonot, Monograph, pt. 1, p. 251, 

1893. 

I have seen no Colombian examples, but there are six 
males in the National Museum from Guatemala which 
Dyar identified as dasypyga. There is no reason to 
question his identification; for the specimens have the 
typical Coptarthria antenna, are the right size, and agree 
in color and maculation with Ragonot's description of 
the Zeller type. 

Forewing gray-brown with a very slight intermixture 
of ocherous shading above inner margin ; the basal area 
a trifle paler than remainder of wing; branches of median 
vein (veins 2 to 5) faintly streaked with brown or black- 
ish brown; transverse lines pale with dark borders and 
faintly tinged with reddish scaling at middle and near 
costa; antemedial line well out near middle of wing, 
nearly vertical, straight except for a slight inward angu- 
lation at vein lb, its inner border a weak brown line, its 
outer bordering line black; subterminal line well back 
from termen, vertical to vein 4, thence oblique to inner 
margin, its inner border a black line; a rather large, oval, 
ochraceous spot on discocellular vein, margined by faint 
black scaling; along termen a row of conspicuous, more 
or less confluent black spots. Hindwing semi translu- 
cent white; the veins faintly darkened (pale, ocherous 
brown) and a faint dark line along termen. Alar ex- 
panse, 14-17 mm. 

Male genitalia as given for the genus. 

Type locality: Honda, Colombia (type, cT, in BM). 

Food plant: Unknown. 



Distribution: Colombia: Honda, Guatemala: Cay- 
uga (Apr., May), Quirigu^ (Mar.), VolcAn Santa Maria 
(July). 

Associated with the males in the National Collection 
are five females from Cayuga (Apr., May) identical with 
the males in all superficial characters. Their genitalia 
are like those of the Bolivian female I have associated 
tentatively with the type otDifundella tolerata (fig. 717). 
However, there is also a female with the same color and 
markings from Juan Vinas, Costa Rica (Feb.), which 
has different genitalia (fig. 718), similar to those of 
Anadelosemia. From the limited material available 
and the few and scattered distributional records it is im- 
possible to determine which females go with which males. 
31. Genus Promylea Ragonot 

Promylea Ragonot, N. Amer. Phycitidae, p. 5, 1887; Monograph, 
pt. 1, p. 207, 1893.— Hulst, Phycitidae of N. Amer., p. 139, 
1890. (Type of genus: Promylea lunigerella Ragonot). 

Tongue well developed. Antenna pubescent; shaft 
of male without notch or other modifications. Labial 
palpus upturned, slender, reaching vertex; third seg- 
ment about half the length of second, acuminate. Max- 
illary palpus squamous. Forewing smooth; 11 veins; 
vein 2 from before, but rather near lower outer angle of 
cell; 3 from the angle, closer to 4 than to 2 at base; 4 
and 5 closely approximate for a short distance from 
base; 6 from below upper angle of cell, straight; 8 and 9 
long stalked, the free element of 9 short; 10 from the 
stalk of 8-9, or from the ceU, connate with or closely 
approximate with it for a short distance beyond base 
(definitely stalked with 8-9 in most of the specimens of 
lunigerella); male without costal fold. Hind wing with 
vein 2 from before lower outer angle of cell ; 3 from the 
angle, connate with 4; 4 and 5 closely approximate or 
anastomosed for half their lengths beyond cell ; 7 and 8 
approximate or partially anastomosed for less than half 
their lengths beyond ceU ; ceU nearly half the length of 
wing; discocellular vein curved. Eighth abdominal seg- 
ment of male with sternite developed as a narrow sclero- 
tized pocket. 

Male genitalia as in Coptarthria except: Apical margin 
of uncus rounded ; lateral arms of gnathos more strongly 
developed; penis sometimes with a weak cornutus. 

Female genitalia with bursa and ductus bursae mem- 
branouSj bursa small, considerably shorter than ductus; 
signum present but weak, a patch of scobinations or a 
small plate supporting a very small thorn ; genital open- 
ing simple; ductus seminalis from ductus bursae. Col- 
lar of eighth abdominal segment with a broad, flaring, 
sclerotized apron projecting from center of anterior 
dorsal margin ; in the intersegmental area between collar 
and seventh segment a sclerotized and coarsely granu- 
late pocket (fig. 721a). 

The genus is close to both Coptarthria and Anadelo- 
semia but distinct, differing from the former in its simple 
male antenna and from the latter and all the genera of 
this immediate gi-oup having the sinuate, involved 
transtilla and the free spine associated with aneUus by 
its peculiarly developed, female, eighth-segment collar. 



66 



TTNITED STATES NATIONAL MUSEUM BULLETIN 207 



In habitus the moths of Promylea differ strikingly from 
those of Coptarthria; the transverse lines of forewing 
being more widely separated and the antemedial line 
decidedly obUque. 

129. Promylea lunigerella Ragonot 

Figures 9, 240, 721 

Promylea lunigerella Ragonot, N. Amer. Phycitidae, p. 5, 1887; 
Monograph, pt. 1, p. 20, 1893.— Hulst, Phycitidae of N. 
Amer., p. 139, 1890.— McDunnough, Check list, No. 6146, 
1939, 

Forewing gray or brownish gray, pale in median and 
most of basal areas and (in many specimens from Van- 
couver and Washington State) with a faint rosy tint; 
antemedial line indicated chiefly by its blackish outer 
border, strongest towards costa and oblique from costa 
before one-third to inner margin, preceded by reddish 
brown or ocherous brown patch, broad and inwardly 
dark-margined on inner margin, attenuated and paling 
out towards costa; subterminal line pale gray, bordered 
inwardly by a blackish brown line and outwardly by a 
much fainter dark line (grayish or reddish brown), out- 
wardly ciurved between veins 8 and lb; discal dots rarely 
separated, normally fused into a thin blackish lunule 
on the discoceUular vein; terminal dots obscure, when 
distinguishable more or less fused into a line along ter- 
minal margin. Hind wings pale smoky fuscous ; the veins 
little if any darkened; a very faint brownish line along 
termen. Alar expanse, 20-24 mm. 

Type localitt: Vancouver IsL, British Colimibia 
(type in Paris Mus.). 
n Food plant: Unknown. 

Distkibtjtion: Canada: British Columbia, Fitzgerald 
(June), Dimcans (Vancouver Isl.), Victoria (June, July, 
Aug.), United States: Washington, BelMngham (Aug.), 
Friday Harbor (July, Aug.), Mt. Constitution (July); 
California, Glen Alpine (Lake Tahoe, July). 

130. Promylea lunigerella glendella (Dyar) 
Figures 241, 720 

Myelois glendella Dyar, Journ. New York Ent. Soc, vol. 14, p. 30, 

1906. 
Promylea glendella (Dyar) Barnes and McDunnough, Check list 

of the Lepidoptera of Boreal America, No. 6584, 1917. — 

McDunnough, Check list. No. 6147, 1939. 

There is nothing to distinguish this from many of our 
specimens of lunigerella from Washington and British 
Columbia except some slight differences in genitalia 
of very doubtful significance, and the name should prob- 
ably go into synonymy; but untU material is available 
from intervening areas and something is known of the 
life history, glendella may be retained as a possible local 
race. It certainly is nothing more than that. 

Type locality: Glenwood Springs, Colo, (tjrpe in 
USNM). 

Food plant: Unknown. 

Represented in the National Museum by two males 
and three females from the type locality (Aug., Sept., 
Oct.). In addition to the foregoing there are before me 
five examples of another variety of lunigerella from Fal- 



len Leaf Lake, Calif., Aug. 8 and 12, 1932, H. H. Keifer, 
collector. They are quite different in color, having 
paler gray forewings with much fainter transverse dark 
markings and more whitish hind wings than our other 
examples of lunigerella. They are probably nothing 
but a color form and for obvious reasons are going with- 
out a name at present. 

131. Promylea dyari, new name 
Figure 243 

Dioryctria eimmermani Druce (not Grote), Biologia Centrali 
Americana, Lepidoptera Heterocera, vol. 2, p. 564, 1899. 

Dioryctria drucei Dyar, Ins. Insc. Menstr., vol. 7, p. 44, 1919 
(preoccupied). 

Similar to lunigerella except: Larger and darker; the 
paler areas of forewing a dark ashy gray; outer black 
border of antemedial fine broader, forming a small tri- 
angle on costa; inner border of subterminal fine also 
slightly stronger, black; the subterminal line itself is 
more sharply angled below costa and from about vein 6 
proceeds to inner angle in a slanting almost straight line 
(a difference that strikes the eye but is hardly signifi- 
cant; for on some examples of lunigerella the subterminal 
line is similarly shaped) ; terminal dots confluent, form- 
ing a conspicuous black line along termen. Alar ex- 
panse, 30 mm. 

Male genitalia with tegumen and vinculum somewhat 
stouter (broader in proportion to their width) than those 
of lunigerella; penis armed with a weak cornutus. 

Type locality: Rinconada, Vera Cruz, Mexico (type 
in USNM). 

Food plant: Unknown. 

The species is known only from the male type which 
both Druce and Dyar mistook for a female. The refer- 
ence to Dioryctria is difficult to understand in Dyar's 
case; for vein 3 of hind wing is appreciably too short for 
that genus. The transfer of "Dioryctria drucei Dyar" 
and "Nephopteryx druceii Ragonot" to the genus Promy- 
lea makes the former a secondary homsnaym and necessi- 
tates the new name. It is possible that the two "drucei" 
represent only different sexes of one species; but this 
caiOnot be determined without more material of each, so 
for the present they must be treated as separate species. 

132. Promylea druceii (Ragonot), new combination 
Figure 722 

Nephopteryx druceii Ragonot, Nouv. Gen., p. 15, 1888; Mono- 
graph, pt. 1, p. 301, 1893. 

I have not seen any specimens matching Ragonot's de- 
scription or figure (Monograph, pi. 9, fig. 17) ; but I have 
before me an excellent photograph of the type supplied 
by Tarns. This shows a moth similar to dyari but with 
the dark markings (especially the dark borders of the 
transverse lines) much more expanded, the outer border 
of the antemedial line forming a large triangle on costa. 
According to Ragonot these dark areas have a decided 
purple tint, rather than the duU black or blackish brown 
of dyari. The spot on the inner margin before the ante- 
median line is also piirplish black rather than orange (as 



AMERICAN MOTHS OP THE SXJBFAMIliT PHTCITINAE 



in dyari) and there is considerable purplish dusting on 
the paler areas of the wing. The moth is also smaller 
than Dyar's type. Alar expanse, 24 mm. 

Type locality: Totonicapdn, Guatemala (type in 
BM). 

Food plant: Unknown. 

Represented only by the type. 

133. Promy^Iea mindosis T)yar 

FiGTTREs 242, 723 

Promylea mindosis Dyar, Ins. Insc. Menstr., vol. 10, p. 172, 1922. 

A narrow-winged, suffused, dark species. Ground 
color of forewing very dark gray-brown; the blackish 
transverse Hues narrow and faint; antemedial line ob- 
solete except for its faint outer border; subterminal line 
distinguishable but faint; discal spots at end of cell 
weak but apparently separated. Hind wing very pale 
smoky fuscous, translucent. Alar expanse, 25-26 mm. 

Male genitaUa with penis with weak cornutus. 
Female genitalia with signum developed as a small 
plate bearing a minute, knoblLke projection. 

Type locality: Mexico City, Mexico (Aug. ; type in 
USNM). 

Food plant: Unknown. 

Ejiown only from the type series of two males and 
one female from the type locaUty. 

134. Promylea dasystigma Dyar 

Figure 724 

Promylea dasystigma Dyar, Ins. Insc. Menstr., vol. 10, p. 172, 
1922. 

Similar to mindosis, differing only in slight details: 
The forewing is a trifle darker, more blackish than 
brownish gray; the subterminal line ends on inner 
margin in a small but distinct white spot; discal spots 
fused into a thin, blackish lunule on discocellular vein. 
The apron from the eighth-segment collar is differently 
shaped and larger and the intersegmental pocket be- 
tween seventh segment and collar proportionally wider 
than those of mindosis (compare figs. 723a and 724a). 
Alar expanse, 25-26 mm. 

Female genitalia with signum a very weak patch of 
scobinations. 

Type locality: Mexico City, Mexico (Aug. ; type in 
USNM). 

Food plant: Unknown. 

Known only from the two females of the type series. 

32. Genus Anadelosemia Dyar 

Anadelosemia Dyar, Ins. Insc. Menstr., vol. 7, p. 51, 1919. (Type 
of genus: Nephopteryx senesciella Schaus). 

Characters of Promylea except: Forewing with vein 
10 always from the cell and not closely approximate to 
the stalk of 8-9. Hind wing with vein 2 from close to 
lower outer angle of ceU; 3 from the stalk of 4-5 or at 
least anastomosed with it for a short distance; 4 and 5 
stalked for over half their lengths beyond ceU; cell about 
one-third the length of wing. Female abdomen with 



eighth-segment collar simple ; no dorsal pocket between 
seventh and eighth segments. On male a hair-pencil 
from lower, outer side of metathorax near base of leg. 

Very close to Promylea but apparently distinct 
enough. The general habitus of the moths is similar 
except that the species of Anadelosemia (except for 
obstitella) are decidedly smaller. 

135. Anadelosemia senesciella (Schaus) 
Figures 11, 244 

Nephopteryx senesciella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 

11, p. 251, 1913. 
Anadelosemia senesciella (Schaus) Dyar, Ins. lusc. Menstr., vol. 

7, p. 52, 1919. 

Forewing ashy white, the basal area to antemedial 
line stained with pale brown; costal edge at base black; 
antemedial Une white, broad and oblique from costa to 
lower fold, thence crescentiform to inner margin, on 
upper half bordered by a broad blackish spot, diffused 
on costa and continued below fold as a narrow line to 
inner margin, a weaker dark inner border on lower half; 
a dark (brownish) shade extending obliquely across the 
wing from costa just before subterminal line to near 
middle of inner margin, irregular and more or less 
diffused over remainder of outer area; discal spots dis- 
tinct, blackish (on this and most other species of the 
genus different on opposing forewings, separated or 
fused into a line); subterminal line sinuate-dentate, 
rather close to termen, bordered inwardly by a few 
blackish dots and outwardly by a black hne which be- 
gins as a strong dash on costa; a row of blackish dots 
along termen. Hind wing semihyaline tinted with 
brown; the veins darkened and a narrow dark shade 
along termen. Alar expanse, 15 mm. 

Male genitalia distinguished chiefly by the shapes of 
uncus, harpe, and vincidum (fig. 244) ; penis armed with 
an elongate, narrow, flattened, bladeUke cornutus. 

Type locality: Juan Vinas, Costa Rica (Jan.; type 
in USNM). 

Food plant: Unknown. 

Kjiown only from the male type. 

136. Anadelosemia tecmeesella (Schaus) 

Ceracanthia tecmessella Schaus, Ann. Mag. Nat. Hist., ser. 8, 

vol. 11, p. 251, 1913. 
Anadelosemia tecmessella (Schaus) Dyar, Ins. Insc. Menstr., vol. 

7, p. 52, 1919. 

Forewing duU ashy white; an oblique blackish shade 
at base ; antemedial Une narrow, white, a broken narrow 
inner black border indicated ; bordering the antemedian 
line on costal half, a rather large triangular bronzy 
brown spot; this color diffused outwardly along costal 
edge; the oblique dark shade across wing beyond cell 
extended to include the remainder of the outer area; 
subterminal line but slightly paler than the brownish 
color of outer area, otherwise as in senesciella; discal dots 
distinct, blackish brown, well separated. Hind wing 
pale smoky brown, veins and terminal margin darker. 
Alar expanse, 14 mm. 



68 



UNITED STATES NATIONAL MUSEUM BULLETDT 207 



Type locality: Avangarez, Costa Rica (July; type 
in USNM). 

Food plant: Unknown. 

Known only from the female type. It lacks an 
abdomen so genitalia could not be studied. However, 
the coloration of forewing suggests a species distinct 
from anything else in the genus. 

137. Anadelosemia fifria Dyar 

Figure 726 
Anadelosemia fifria Dyar, Ins. Insc. Menstr., vol. 7, p. 52, 1919. 

Forewing similar to that oi senesciella except: Whitish 
ground color more extended filling most of outer area ; a 
dark brown shading at extreme base; dark shadings 
fuscous brown rather than blackish; antemedial white 
line narrow, its dark borders also narrower; the post 
media, transverse dark shade also narrower; terminal 
dots more or less confluent. Alar expanse, 15 mm. 

Female genitalia with signum, developed as a narrow, 
smaU, shallow, granulate cup. 

Type locality: Cayuga, Guatemala (type in USNM). 

Food plant: Unknown. 

Represented only by the female type and one other 
female from the type locality (May). 

138. Anadelosemia base Dyar 

FiGUBE 727 

Anadelosemia base Dyar, Ins. Insc. Menstr., vol. 7, p. 52, 1919. 

Similar to the foregoing species (Jvfria) except: Dark 
markings a paler brown; the outer dark border of costal 
haK of antemedian line a small triangulate spot with 
some extension outward on costal edge; terminal dots 
rather weak but not confluent. Alar expanse, 15 mm. 

Female genitalia without signum; ductus bursae pro- 
portionally much longer than that of fijria; ductus 
seminalis from near middle of ductus bmsae. (In the 
other species it branches off from the ductus bursae 
very near its junction with the bursa copula trix.) 

Type locality: Cayuga, Guatemala (type in USNM). 

Food plant: Unknown. 

Known only from the female type and one other 
female from the type locahty. Superficially hardly dis- 
tinguishable from jljria. However, the differences in 
their female genitaha suggest two distinct species. 

139. Anadelosemia obstitella (Schaus), new combination 

Figure 728 

Nephopteryx obstitella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 
11, p. 251, 1913. 

The largest and most strikingly marked species in the 
genus; the antemedial and subterminal lines shining 
white; the inner, black bordering line of the former 
continuous from costa to inner margin and strongly con- 
trasted against the ashy gray ground color of the basal 
area of the forewing; outer black border on costal half 
of antemedial line and the black dashes preceding and 
following the subterminal line enlarged and well con- 
trasted. Hind wing semihyaline white with a very faint 



brownish tint; veins faintly darkened; a narrow, pale 
fuscous shading along termen, especially towards apex. 
Alar expanse, 22 mm. 

Female genitalia with signum present as a narrow, 
weak, elongate, irregular scobinate patch (see enlarge- 
ment). The shape and development of signum is a 
character of very doubtful value in this genus and 
probably subject to considerable individual variation. 

Type locality: Mount Pods, Costa Rica (type in 
USNM). 

Food plant: Unknown. 

Known only from the female type. 

140. Anadelosemia texanella (Hulst), new combination 
Figures 246, 729 

Myelois texanella Hulst, Canadian Ent., vol. 24, p. 60, 1892. 

Myelois dulciella Hulst, Canadian Ent., vol. 32, p. 176, 1900. 

Tacoma texanella (Hulst) Barnes and McDunnough, Contribu- 
tions, vol. 3, p. 193, 1916; vol. 4, p. 174, 1918.— McDun- 
nough, Check list, No. 6143, 1939. 

Forewing ashy white (due to a fine, sparse peppering 
of fuscous scales on the white ground color) ; costal edge 
at extreme base black; antemedial line not differenti- 
ated except by the narrow, curved, black line forming 
its outer border from costa to inner margin and by a 
preceding brown spot on inner margin; subterminal line 
sinuate, nan-ow, bordered inwardly by a continuous 
black line and outwardly by a faint, narrow, brownish 
shade continued from a blackish dash on costa; discal 
spots black, separated. Hind wing smoky white, dark- 
ening outwardly; a fine brown Kne along termen. Alar 
expanse, 13-16 mm. 

Male genitalia figured from type of dulciella. They 
exhibit several specific characters: a long, slender, 
strongly sclerotized, spinelike, apical projection from 
gnathos (the other species whose males are known show 
no such structure, the only elements attached to the 
lateral arms of gnathos at their junction being the 
transtilla and the base of the more or less sclerotized 
subanal plate) ; tegumen considerably elongated in pro- 
portion to the vinculum; harpe short and broad; penis 
finely spined at apex. Female genitalia without signum; 
ductus seminalis from ductus bursae near junction of 
bursa copulatrix. The distinctive female structural 
character is the shape of the eighth-segment collar (fig. 
729). 

Type localities: Blanco County, Tex. (texanella, 
in AMNH, ex Rutgers); Hastings, Fla. {duldella, in 
AMNH, ex Rutgers). 

Food plant: Unknown. 

Distribution: United States: Texas, Blanco 
County, San Benito (Apr., Sept.); Florida, Hastings 
(Oct.). Puerto Rico: San German (Apr.) ; Cuba: Santa 
Clara, Central Soledad ("E. E. A. Cuba, Ento. no. 
10234," May). 

The Puerto Rican and Cuban examples are males 
which agree in every detail with the type of duldella. 
The type of texanella lacks an abdomen, so its genitalia 
could not be checked (the other Texas examples are 
females) ; but careful study of the two types discovered 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



69 



no diflference that woiild justify any doubt of the syn- 
onymy proposed by Barnes and McDunnough (1918). 

141, Anadelosemia condigna, new species 
Figures 245, 730 

Forewing similar to that of texanella except: Outer 
black border of antemedial line nearer middle of wing, 
nearly vertical and more denticulate; sub terminal line 
with a discontinuous, blackish outer border; some red- 
dish spotting in the postmedial area near inner margin; 
costa at base not black, but a transverse black marking 
at extreme base; discal spots confluent, forming a black- 
ish or reddish brown line along discocellxilar vein. Hind 
wing semihyaline white with a narrow fuscous shade 
along termen. Alar expanse, 15-18 mm. 

Male genitalia similar to those of senesciella, but dif- 
fering in the shapes of uncus, harpe, and vinculum; penis 
armed with a narrow, flat, bladelilie cornutus similar 
to that of senesciella. Female genitalia without signum, 
similar to that of texana except for the shape of the 
eighth-segment collar (fig. 730). 

Type locality: Prescott, Ariz, (type in USNM, 
61329; paratypes in Cornell Univ. and BM). 

Food plant: Unknown. 

Described from male type from the type locality 
(July) and seven male and four female paratypes from 
the Baboquivari Mts., Pima County, Ariz., collected by 
O. C. Pohng, May 1-15, 1924. 

33. Genus Dasypyga Ragonot 

Dasypyga Ragonot, N. Amer. Phycitidae, p. 5, 1887; Monograph, 
pt. 1, p. 206, 1893.— Hulst, Phycitidae of N. Amer., p. 138, 
1890. 

Tongue well developed. Antenna weakly pubescent. 
Labial palpus upcurved, reaching to vertex (female) or 
nearly to it (male) ; third segment half as long as second, 
acuminate. Maxillary palpus small, squamous. Fore- 
wing with some raised scaling on basal area (probably 
only a specific character); 11 veins; vein 2 from before 
but near lower outer angle of cell; 3 from the angle, 
closer to 4-5 than to 2 ; 4 and 5 shortly stalked ; 6 from 
below upper angle of cell, straight; 10 from the cell, 
closely approximate for some distance with the stalk of 
8-9; male without costal fold. Hind wing with vein 2 
from near lower outer angle of cell; 3 from the angle, 
connate with the stalk of 4-5; 4 and 5 long stalked (for 
about two-thirds of their lengths) ; 7 and 8 closely ap- 
proximate or contiguous for a short distance beyond 
cell; cell one-third the length of wing; on male with anal 
angle folded into a thickened pocket containing a hair 
tuft. Eighth abdominal segment of male with sternite 
developed as a narrow, sclerotized pocket. 

Male genitalia similar to those of Anadelosemia ex- 
cept: Sacculus of harpe strongly sclerotized and for half 
its length developed as a free arm extending across 
harpe; cornutus well developed; a pair of hair tufts from 
intersegmental area adjacent to outer surfaces of the 
sacculi at their bases. 

Female with signum a small patch of scobinations ; 
ductus bursae considerably longer than bursa, tubular 



and strongly sclerotized for about one-fifth its length 
from genital opening, and for over half its remaining 
length sclerotized, sUghtly flattened and bent (sinuate) ; 
ductus seminalis from biu^a near its junction with 
ductus bursae. 

A distinct genus distinguished from the other genera 
with similar transtillae and venation by its stout, greatly 
produced sacculus and sclerotized ductus bursae. Con- 
tains one North American species. 

142. Dasypyga alternosquamella Ragonot 
Figures 12, 247, 719 

Dasypyga allernosquamella Ragonot, N. Amer. Phycitidae, p. 5, 
1887; Monograph, pt. 1, p. 206, 1893.— Hulst, Phycitidae 
of N. Amer., p. 138, 1800.— Heinrich, Proc. U. S. Nat. Mus., 
vol. 57, p. 84, 1920 (larva, pupa, life history). — Essig, 
Insects of western North America, p. 709, 1926. — McDun- 
nough, Check list, No. 6145, 1939. 

Dasypyga alternosquamella stictophorella Ragonot, N. Amer. 
Phycitidae, p. 5, 1887; Monograph, pt. 1, p. 206, 1893. 

Forewing pale salmon pinls; basal area black dusted 
with white, this black area (from the upper vein of cell 
to inner margin) extending almost to middle of wing; 
the black and white scahng coarse and more or less 
raised in base and along outer margin of antemedial 
line which cuts the black areas as a narrow, pale (och- 
raceous red), obliquely curved line; subterminal line 
absent, but along termen a rather broad border of coarse 
black and whitish scales; on most specimens a clear 
white line extends outward from lower angle of cell 
along vein 5 towards and sometimes to the dark terminal 
border and usually enclosing a detached patch of black 
scales forming a discal spot at end of cell; above this 
(below vein 6) a fine red line rims to near outer dark 
margin and thence angles sharply to ape.x; on well 
marked and fresh specimens traces of a similar red line 
on the lower fold in outer area. Hind wing very pale, 
shining, smoky fuscous with a faint dark line along 
termen. Alar expanse, 19-24 mm. 

Genitalia as given for the genus ; figured from reared 
examples ; cornutus of male penis about half as long as 
aedeagus, somewhat flattened, twisted, and bluntly 
pointed ; penis also minutely scobinate towards apex. 

Type locality: California (type in Paris Mus.). 

Food plant: Razamqfskya cryptopoda. 

Distribution: United States: California, Lake 
AiTowhead (May); Arizona, Mohave County (Sept.), 
Williams (June); Colorado, Glenwood Springs, Monu- 
ment (May, June, Aug.); Washington, Bellingham 
(June), Seattle. Canada: British Columbia, Kaslo 
(June). 

A species that can be at once recognized by its pecu- 
liar markings and coloration. The white longitudinal 
line on forewing is of varying length on different speci- 
mens but is present on all that I have seen, though 
sometimes weak. Its presence is the characteristic 
feature given by Ragonot for his variety stictophorella. 
It is presumably absent from the type of alternosqua- 
mella. I suspect that the latter is an individual variant. 
Notes on the life history and descriptions of larva and 
pupa are given in my paper. 



1Q 



UNITED STATES NATIONAL MUSEUM: BULLETIN 207 



Genus 34: Rampylla 

[Male: Venational division D. Transtilla present but variously 
modified. Harpe with apex of sacculus produced, strongly 
sclerotized and pointed. Hind wing triangulate; anal angle 
folded and produced; sex tufts and scalings on lower surface. 
Female: Venational division B. Bursa and ductus bursae 
simple; ductus seminalis from bursa. Hind wing with cell one- 
third the length of wing.] 

34. Genus Rampylla Dyar 

Rampylla Dyar, Ins. Insc. Menstr., vol. 7, p. 84, 1919. (Type 
of genus: Rampylla orio Dyar). 

Tongue well developed. Antenna pubescent; shaft 
of male slightly thickened. Labial palpus upturned, 
not reaching vertex in male, a trifle longer in female; 
second segment slightly rough scaled beneath; third 
segment bluntly acuminate, about half the length of 
second (shorter on male than female). MaxiQary 
palpus squamous, small. Forewing smooth except for 
a sHght, projecting scale tuft from inner margin near 
base on male; 11 veins; vein 2 from before lower outer 
angle of cell; 3 from the angle, well separated from 2; 
4 and 5 approximate at base and for a very short dis- 
tance beyond; 6 from below upper angle of cell, straight; 
10 from the cell; male without costal fold. Hind wing 
with vein 2 from before the lower outer angle of cell; 
3 from the angle and in the male almost as long as vein 
2; in the female considerably shorter; 4 and 5 very 
shortly stalked or contiguous for a short distance from 
cell, connate with 3 ; 7 and 8 contiguous or closely ap- 
proximate for a short distance from cell; cell in male 
one-fourth the length of wing, in female approximately 
one-third; discocellular vein curved; on male anal area 
(involving vein la) thickened and folded, forming a 
produced pocket, enclosing a long hair pencil; under- 
side of male wing with roughened scale or hair tufts on 
some of the veins. Eighth abdominal segment of male 
with sternite developed as a narrow, sclerotized pocket. 
Metathorax of male with a stout pencil of spatulate 
scales from just above base of leg. 

Male genitalia with transtilla present, variously modi- 
fied (greatly reduced in lophotalis). Harpe with sac- 
culus produced into a strong free hook at apex. A long 
free spine associated with anellus (as in the four preced- 
ing genera). 

Female genitaUa with bm-sa more or less finely scobin- 
ate and with some concentration of these fine scobina- 
tions but no definable sigmmi; ductus bursae simple, 
short (shorter than bursa except in lophotalis). An 
invaginated, sclerotized, dorsal pocket at apical end of 
seventh abdominal segment or (in lophotalis) a sclero- 
tized, granulate, dorsal pocket between ovipositor and 
eighth-segment coUar. 

The length of vein 3 in proportion to 2 of hind wing 
places the males in our venational division D and, in 
conjunction with their decidedly triangulate hind wings 
and the sex-scalings and tuf tiogs on their imder surfaces, 
readily distinguishes the genus. The females on hind 
wing venation fall into division B. The four species 
here recognized are all tropical American. They ex- 



hibit distinct specific differences in genitaha, color, and 
maculation. The transverse lines of forewing, in their 
rather close approximation and narrow black borders, 
are similar to those of Coptarthria to which Rampylla 
seems most nearly related. 

143. Rampylla orio Dyar 

Figures 45, 248 
Rampylla orio Dyar, Ins. Insc. Menstr., vol. 7, p. 84, 1919. 

Forewing violaceous gray; under magnification ex- 
treme base and a rather broad area along inner margin 
to beyond middle tinted with purplish red; antemedial 
line obsolete; sub terminal line narrow, blackish, rather 
close to and nearly parallel with termen, vertical from 
costa to vein 8, inwardly angled between veins 8 and 5, 
thence obhque to ioner margin, bordered iawardly by a 
faint ocherous shade and outwardly by a narrow ocher- 
ous line; discal dots at end of cell separate, ocherous; a 
thin dark line along terminal margin. Hind wing trans- 
lucent white with a fuscous shade at apex and anal area 
yellow; veins not appreciably darkened; terminal mar- 
gin darkened only towards apex; on underside of hind 
wing (male) a yeUow hair tuft at origin of veins 4 and 5 
from cell; a white fringe continuing outwardly on these 
veins and a similar white fringe on vein 7 above the 
yellow hair tuft. Alar expanse, 22 mm. 

Male genitalia with lateral portion of uncus on each 
side produced iato a broad, strongly sclerotized, projec- 
tion armed at apex with a cluster of long, slender, strong, 
black spines. TranstiUa produced caudally iato a bi- 
lobed, scobinate process fusing with reduced arms of 
gnathos. Tegumen with a long, stout, curved, free arm 
arising from base at each ventrolateral angle. Harpe 
with produced sacculus developed as a long, stout, taper- 
ing hook, curved across face of harpe. Anellus heart- 
shaped; associated spine straight. Aedeagus partially 
sclerotized; penis with some farat, sclerotized wrinklings 
and a few microscopic scobinations, otherwise unarmed. 

Type localitt: Zacualp^n, M6xico (type in USNM). 

Food plant: Unknown. 

Known only from the male type. 

144. Rampylla polydectella (Schans) 
Figure 732 

Salebria polydectella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 

p. 250, 1913. 
Rampylla polydectella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7, 

1919. 

Forewing brownish gray with a faint purplish tint; a 
narrow border along inner margin between the trans- 
verse lines and a somewhat broader area along termen 
dusted with duU, grayish white; antemedial line narrow, 
faint, pale gray, indicated chiefly by its narrow, black 
outer-bordering hne, the line well out towards middle of 
wing and shghtly sinnous, nearly vertical; subterminal 
line equally thin and pale gray with a similar black inner 
border, angled outward slightly at middle and nearly 
parallel with termen; a black line along terminal margin; 
discal spots small, confluent, ocherous; faint indication 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



of black stroaklets on veins 2, 3, and 4 just beyond cell. 
Hind wing glossy brown-gray; the veins faintly dark- 
ened and a fine dark line along termen. Alar expanse, 
20 nun. 

Female genitalia distinguished chiefly by the shape 
of eighth-segment collar and the invaginated, sclero- 
tized pocket from seventh segment. 

Type locality: Juan Vinas, Costa Rica (June ; type 
in USNM). 

Food plant: Unknown. 

Known only from the female type. 

145. Rampylla subcaudata (Dyar), new combination 

Figures 249, 733 

Cerocanthia subcaudata Dyar, Ins. Insc. Menstr., vol. 7, p. 42, 
1919. 

Forewing ocherous gray, the basal area to antemedial 
line purplish tinted ; a similar purplish shade along outer 
border of the subterminal line; a large, somewhat 
darker (fuscous) patch surrounding the discal spot and 
extending from antemedial almost to subterminal line 
and from vein 2 to costa, darkest on veins 2 to 4 ; ante- 
medial line faint, narrow, ocherous, indicated by a thin 
blackish brown line forming its outer border, the latter 
near middle of wing, shghtly sinuate and nearly vertical ; 
subterminal line with a narrow, blackish brown, inner 
border, slightly outbent or angled at middle; discal 
mark a narrow ocherous spot along discocellular vein; 
a narrow blackish line along terminal margin. Hind 
wing of male subpellucid white with a faint yellowish 
tint, decidedly ocherous along inner margin; of female 
with pale smoky tint; underside of male hind wing with 
a yellowish scale tuft covering the bases of veins 2 to 
5. Alar expanse, 16 mm. 

Male genitalia with margins of uncus evenly rounded, 
lacking any sclerotized projections. Harpe with 
apical projection of sacculus a slender, upciurved 
hook; a long, stout, hair and scale tuft from a pad ad- 
jacent to base of sacculus. Penis armed with a narrow, 
weakly sclerotized, bladehke cornutus. Female geni- 
talia similar to those of polydectella, differing only in 
slight details in the shape of the eighth-segment coUar 
and the sclerotized pocket from seventh segment (fig. 
733a). 

Type locality: Cayuga, Guatemala (May; type in 
USNM). 

Food plant: Unknown. 

In addition to the male type there is one other speci- 
men in the National Collection from Quirigud, Guate- 
mala (May), a female with slightly darker hind wings 
whose genitalia are here figured. In addition there are 
before me two somewhat larger specimens (19 mm.), a 
male in the Janse Collection from San Jos6, Costa Rica 
("H. Schmidt, 8-11"), and a female from the British 
Museum labeled "S. E. Brazil, E. D. Jones, 1920-303." 
The genitalia of the latter are almost identical with 
those of polydectella. The genitalia of the male are like 
those of the type of subcaudata. The two specimens 
are identical in size, color, and markings. Except for 



1h 

size (which is not significant) they agree superficially 
with the type of subcaudata. Despite the differences in 
fore and hind wing coloration I suspect that subcaudata 
may not be specifically distinct from polydectella. 

146. Rampylla lophotalis, new species 
Figures 250, 731 

Similar to subcaudata except: Ground color of fore- 
wing more yellowish; the dark areas, especially just 
preceding the antemedial line and in the dark area sur- 
rounding the discal spot, blackish ; dark lines bordering 
the transverse lines blackish rather than blackish brown; 
antemedial line farther out on wing, extending from 
midcosta to inner margin beyond middle, narrowing the 
area between antemedial and subterminal lines. Hind 
wing with the scale tuft on underside blackish rather 
than yellow, the blackish shade extending to the costa 
and visible through the wing from above. Alar ex- 
panse, 17.5-18 mm. 

Male genitalia with two pairs of strongly sclerotized, 
curved, lateral projections from triangulate uncus. 
Gnathos entirely absent (unless the lower pair of pro- 
jecting arms from uncus can be interpreted as lateral 
arms of a gnathos, which is very doubtful, as there is 
no separation whatever between them and the uncus). 
Transtilla reduced to a shortened and slender, trans- 
verse, centrally bent band. Tegumen with a short, 
two-pronged projection from each ventrolateral angle. 
Anellus and harpe as in subcaudata. Female genitalia 
with a thick, sclerotized roll on the back of ovipositor; 
a broad, coarsely granulate pocket between ovipositor 
and collar; eighth-segment collar narrow; ductus bursas 
shghtly longer than bursa. 

Type locality: Jalapa, Mexico (type in USNM, 
61330). 

Food plant: Unknown. 

Described from male type and one female paratype 
from the type locality and one male paratype from 
Volcdn Santa Maria, Guatemala (Nov., Schaus and 
Barnes, collectors). The male holotype bears a label 
in Hampson's handwriting "Cerocanthia lophotalis 
Hampson, c? type." As far as I know Hampson never 
published a description of the species. 

Genus 35: Fulrada 

[ Venational division D. Hind wing with veins 4 and 5 contiguous 
or closely approximate beyond cell (not stalked) ; cell one-fourth 
the length of wing; discocellular vein incomplete. Eighth ab- 
dominal segment of male with broad ventral tuft. Transtilla 
vestigial. Harpe with apex of sacculus not produced.] 

35. Fulrada, new genus 

Type of genus: Dasypyga guerna Dyar. 

Tongue well developed. Antenna weakly pubescent; 
shaft of male simple. Labial palpus upturned, slender, 
reaching to vertex; third segment slightly shorter than 
second, acuminate. Maxillary palpus squamous, small, 
appressed to face. Forewing smooth; 11 veins; vein 2 



72 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



from before but near lower outer angle of cell; 3 from 
the angle, nearly equidistant from bases of 2 and 4; 4 
and 5 closely approximate or connate and contiguous, 
for a short distance beyond base; 6 from below upper 
angle, straight or very shghtly bent towards base; 10 
from the cell, approximate to the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from before 
lower outer angle of cell; 3 from the angle, long (its 
free length slightly shorter than 2); 4 and 5 contiguous 
and closely approximate for slightly less than half their 
distances from angle of cell (not stalked) ; 7 and 8 closely 
approximate beyond cell; cell one-fourth the length of 
wing; discocellular vein incomplete. Eighth abdominal 
segment of male with a broad, ventral hair or scale tuft 
and sternite developed as a narrow sclerotized pocket. 

Male genitalia with uncus subtriangulate, its apical 
margin truncate. Gna'thos represented only by its 
lateral arms, to which (at their junction) is attached 
the base of subanal plate. Transtilla represented only 
by a bilobed central vestige behind (above) the anellus. 
Harpe with sacculus not produced. Anellus semi- 
tubular (guerna) or a broad slightly curved plate with 
a greatly reduced, associated, free spiae (carpasella). 
Vinculum stout, as broad or nearly as broad as long; 
terminal margin broad and slightly rounded. 

The foregoing description is incomplete, as females 
are unknown. The genus is apparently close to both 
Anadelosemia and Eampylla, differing from the former 
chiefly on hiud wing venation and from the latter on 
secondary naale characters. 

147. Fulrada qiiema (Dyar), new combination 
FlQUEE 251 

Dasypyga querna Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 331, 
1914. 

Forewing ocherous gray with a faint dusting of 
reddish scales on basal area and a very faint reddish 
tint over the area below the discal spots; a scattered 
blackish powdering ia costal areas; antemedial line 
obUque, indicated only by a row of (4 or 5) well sepa- 
rated black dots forming its inner border and an outer 
black spot on costa; discal dots at end of cell, small, 
separated, black; a row of smaU but distinct black dots 
along termen. Hind wing translucent, smoky white, 
darker towards apex; veins faintly darkened and a 
narrow dark line along termen. Alar expanse, 1 1 mm. 

Male genitalia with anellus semitubular, oblong, 
bottle shaped, more or less involved with the vestigial 
hairy-lobed transtilla. Harpe with a strongly haired, 
transverse, sclerotized ridge extending from near base 
of costa to lower outer angle of cucullus. Aedeagus 
short; penis armed with a very weak, flattened cornutus. 
Tuft on eighth abdominal segment a row of very fine, 
hairlike scales. 

Type locality: La Chorrera, Panamd (May; type 
in USNM). 

Food plant: Unknown. 

Represented only by the male type. 



148. Fulrada carpasella (Schaus), new combination 

FiGUBE 252 

Piesmopoda carpasella Schaus, Zoologica, vol. 5, No. 2, p. 47, 
1923. 

Forewing white finely irrorated with black and 
brown; some pale tawny shading on the white ante- 
medial line along the inner margin of its black outer 
border, also postmedially below vein 2 and on discocel- 
lular vein; base black, this shade expanding obliquely 
to inner margin and outwardly edged by some pale 
tawny scaling; antemedial line oblique, indicated 
chiefly by its black outer border, the latter strongly 
contrasted from costa to lower margin of cell, very faint 
from cell to inner margin, slightly outcurved from costa; 
subterminal line parallel with termen, slightly sinuous 
and with well-contrasted dark outer and inner borders, 
the inner one a narrow black line, the outer a black, 
angulate costal dash continued as a rather broad 
brownish shade to tornus; discal dots separated, small 
black dots on the outer angles of cell; a row of well- 
contrasted black dots along termen. Hind wing 
whitish towards base, shading to fuscous outwardly, a 
rather broad brownish shade along termen. Alar 
expanse, 12 mm. 

Male genitalia with a vestigial free spine associated 
with the broad, large, slightly curved plate of anellus. 
Harpe with rather long, slender, erect clasper. Aedea- 
gus rather long, slender, straight. Eighth abdominal 
tuft consisting of long, spatulate scales. 

Type locality: Conway Bay, Indefatigable, Gald- 
pagos Islands (Apr.; type in USNM). 

Food plant: Unknown. 

Known only from the male type. 



Genus 36: Scorylus 

[Venational division B. Male antenna with shallow sinus in 
shaft at base. Hind wing with veins 4 and 5 stalked for about 
two-thirds; cell about one-third the length of wing; discocellular 
vein complete, curved. Eighth abdominal segment of male with 
broad, ventral scale tuft. Metathorax with stout scale and 
hair tuft near base of leg. Transtilla vestigial. Anellus with- 
out trace of associated free spine. Harpe with apex of sacculus 
produced.] 

36. Scorylus, new genus 

Type of gentts: Scorylus cuhensis, new species. 

Tongue well developed. Antenna finely pubescent; 
the male shaft with a shallow sinus at base (a shght ex- 
cavation covering several basal segments); within the 
sinus and extending slightly past it a thin layer of modi- 
fied appressed scales. Labial palpus upturned, reaching 
vertex, second segment slightly rough scaled ; third seg- 
ment slightly shorter than second, acuminate. Maxil- 
lary palpus squamous. Forewing smooth; 11 veins; 
vein 2 from before lower outer angle of ceU; 3 from the 
angle, nearer to 4 than to 2 ; 4 and 5 closely approximate 
for a short distance from cell ; 6 from below upper angle 
of cell, slightly bent towards base; 8 and 9 stalked for 
slightly more than half their lengths; 10 from the cell. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



73 



approximate to the stalk of 8-9; male without costal 
fold. Hind wing with vein 2 from before, but close to 
lower outer angle of cell ; 3 from the angle, contiguous to 
the stalk of 4-5 for nearly half the length of the stalk, 
the free length of 3 decidedly shorter than vein 2 ; 4 and 
5 stalked for about two-thirds their lengths; 7 and 8 
closely approximate for some distance beyond cell; cell 
about one-third the length of wing; discocellular vein 
curved. Eighth abdominal segment with broad, stout 
ventral scale tuft and sternite developed as a narrow 
sclerotized pocket. A stout scale and hair tuft from 
metathorax at base of leg. 

Male genitalia with uncus hoodlike. Gnathos well 
developed, its apical projection a long, tapering spike. 
Transtiila represented by a modified central vestige. 
Harpe with sacculus produced at apex into a sclerotized 
hook; a strong hair tuft from lobe near base of sacculus; 
anellus a triangulate plate with short, blunt, broad lat- 
eral lobes; aedeagus moderately stout, slightly swollen 
and bent towards base; penis armed with an elongate, 
narrow cluster of short, thin spines. Vinculum stout, 
slightly tapering and somewhat longer than basal width. 

Female genitalia without signum but with a concen- 
tration of fine granulations in bursa near its junction 
with ductus bursae ; ductus bursae and its genital open- 
ing simple; ductus seminalis from ductus bursae. 

This genus shows affinities to both Fulrada and 
Anadelosemia but is distinct from both. Its venation 
is similar to that of Anadelosemia, to which it appears to 
be most closely allied. 

149. Scorylus cubensis, new species 
Figures 253, 725 

Forewing white with blackish fuscous and reddish 
markings; basal area strongly ii-rorated with red and 
some scattered reddish scaling in the median white area; 
antemedial line indicated only by a blackish outer line 
from costa, expanding into a black spot in cell, continued 
thence as a very thin blackish fuscous line to inner mar- 
gin, bordered outwardly (from cell to ioner margin) by 
an olivaceous patch; a faint triangulate olivaceous- 
fuscous shade over outer area from just beyond middle 
of inner margin to subterminal line at costa; a small 
black spot on midcosta; subterminal line sinuate, 
bordered inwardly by a strong, rather broad, blackish 
line and outwardly by a fainter reddish line continued 
from a strong black spot on costa; discal dots black, 
more or less confluent along discocellular vein; below 
them on the olivaceous fuscous shade a patch of reddish 
scaling; a row of black dots along termen. Hind wing 
translucent white; veins very faintly darkened near 
outer margin; a narrow dark shade along termen. Alar 
expanse, 12-13 mm. 

Male genitalia with outer margin of uncus evenly 
rounded. Transtiila fragment in the shape of two short, 
oblong, pointed plates weakly joined at their bases. 
Apical projection of sacculus sharply upturned, mod- 
erately long, thornlike. Terminal margin of vinculum 



truncate. Female genitalia with bursa large, as long as 
ductus bursae. 

Type locality: Santiago Province, Cuba (type in 
USNM, 61331). 

Food plant: Unknown. 

Described from male type and one male and one fe- 
male paratypes from the type locality (June, Dec, 
Schaus and Barnes, collectors). 

Genera 37-39: Davara to Piesmopoda 

[Venational division D. Forewing with veins 4 and 5 approxi- 
mate for a short distance from cell. Hind wing with cell less 
than one-third the length of wing. Male genitalia with uncus 
bifid (divided to base) ; harpe with strong, hooked clasper from 
near apex of sacculus.] 

37. Genus Davara Walker 

Davara Walker, List, pt. 19, p. 1020, 1859. — Hampson, in 

Ragonot, Monograph, pt. 2, p. 530, 1901. (Type of genus: 

Davara azonaxsalis Walker.) 
Homalopalpia Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 332, 1914. 

(Type of genus: Homalopalpia daleraDy&T. New synonymy.) 
Eucardinia Dyar, Ins. Insc. Menstr., vol. 6, p. 138, 1918; vol. 7, 

p. 50, 1919. (Type of genus: Ulophora caricae Dyar.) 

Tongue well developed. Antenna of male (except in 
rufulella) with basal segment enlarged, deeply notched 
and with a brush of short fine bristles in the notch 
(fig. 254d) ; male shaft simple or flattened and dilated 
towards base, pubescent. Labial palpus upcurved; on 
male (except in rujulella) reaching well above vertex; 
broadly scaled ; third segment considerably shorter than 
second. Maxillary palpus squamous. Forewing more 
or less rough scaled at base on male ; 1 1 veins ; vein 2 
from before, but near lower outer angle of cell ; 3 from the 
angle, nearer to 4 at base than to 2 ; 4 and 5 approximate 
for a short distance from cell; 6 from below upper angle 
of cell, straight or very slightly cm-ved towards base; 8 
and 9 long stalked; 10 from the cell, approximate to the 
stalk of 8-9; male without costal fold. Hind wing with 
vein 2 from before the angle of the cell; 3 from the 
angle, long (its free length slightly shorter than 2) ; 4 and 
5 anastomosed for slightly less than half their lengths 
beyond angle of cell; 7 and 8 contiguous or shortly 
anastomosed beyond cell; cell less than one-third the 
length of wing; discocellular vein ciu-ved. Eighth 
abdominal segment of male with sternite developed as 
a narrow, sclerotized pocket. 

Male genitalia with uncus bifid (divided to base). 
Gnathos weak, a thin narrow band. Transtiila absent 
(except in inter jecta). Harpe with strong, hooked 
clasper from near apex of sacculus ; a long hair tuft from 
lobe near base of sacculus. Anellus a narrow curved 
plate with long, lateral arms; a stout, free spine associ- 
ated with anellus. Penis armed with a thin, narrow, 
curved, flattened, bladelike cornutus. 

Female genitalia with two signa, developed as small, 
granulate depressions ; ductus bursae shorter than bursa, 
with paired cupUke plates behind genital opening or 
with genital opening simple (interjecta) ; ductus seminalis 



74 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



from bursa near its junction with ductus bursae. Collar 
of eighth segment complete. 

Davara and the two following genera (Sarasota and 
Piesmopoda) form a compact group distinguished from 
all other American phycitid genera by the peculiar bifid 
development of their unci. Davara was described by 
Walker on the basis of a single female which he mistook 
for a male; it was referred as a synonym of the Old 
World Phydta by Hampson (1903). Dyar did not 
recognize it. His Homalopalpia was erected on male 
antennal and palpal structures which normally should 
be of generic value, but ia this particidar instance do 
not seem to hold as separating Davara from Piesmopoda. 
They fall down in the case of rwfulella, which on genitalic 
characters of both male and female must be referred to 
Davara. In my opinion Davara and Piesmopoda should 
be retained as separate genera. Their species differ in 
habitus as well as in genitalic structure. In Davara the 
male transtiUa is absent and the female bursa always 
has two signa. In Piesmopoda the transtiUa is present 
and developed as two long, slender, curved free arms 
and the female bursa has a single signum or none. 
Unfortunately an anomalous species {interjecta) seems 
to upset the division. It has the female and all the 
secondary male characters of typical Davara but male 
genitalia of the Piesmopoda type. However, there are 
some minor differences in its male and female genitalia 
which may eventually allow its separation from both 
Davara and Piesmopoda imder a separate generic desig- 
nation. For the present I am referring it tentatively to 
Davara. Its distinctive characters are discussed more 
fuUy imder the specific description. 

150. Davara caricae (Dyar), new combination 

PiGTJHBs 46, 254, 735 

Ulophora caricae Dyar, Proc. Ent. See. Washington, vol. 14, p. 

218, 1913. 
Homalopalpia dalera Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 332, 

1914. (New synonymy.) 
Eucardinia caricae (Dyar) Dyar, Ins. Insc. Menstr., vol. 6, p. 

139, 1918; Ins. Insc. Menstr., vol. 7, p. 50, 1919.— McDun- 

nough. Check list. No. 6119, 1939. 

Antenna of male with the brush in notch of basal 
segment black; the basal segment itself a very pale buff; 
basal segments of shaft decidedly flattened and broad- 
ened and with black serrations on upper edge. Labial 
palpus of male very broadly scaled, the second segment 
reaching well above vertex; less broadly scaled and 
somewhat shorter on female; reddish brown with a 
peppering of pale buff scaUng on outer side and on male 
more or less shaded with blackish brown on inner side. 
Forewing tan-gray shaded with reddish brown; basal 
area (on male) blacldsh brown and rough scaled; a 
whitish or pale buff shade precedes the antemedial line 
and a similar transverse shade crosses the disk and 
includes the discocellular mark; the space between this 
transverse pale shade and antemedial Une suffused with 
reddish brown (on some female examples almost purplish 
fuscous) ; a similar reddish brown suffusion over outer 
area; antemedial hne obUque, sUghtly angled between 



cell and inner margin, faint, ocherous, bordered in- 
wardly and outwardly by narrow dark lines; sub terminal 
Une indistinct except for its brown umer and outer 
borders, sinuate; discal spots fused into a narrow lunu- 
late line on the discocellular vein. Hind wing soiled 
white, with a narrow fuscous shade along termen. Alar 
expanse, 14-18 mm. 

Male genitaha with a cluster of fine, long, hairlike 
spines surrounding the strong, free, forked spine asso- 
ciated with anellus; lateral arms of anellus considerably 
shorter than in other species of the genus. Female 
genitalia with a pair of strong ventrolateral ridges on the 
invaginated portion of the eighth-segment collar. 

Type localities: Miami, Fla. {caricae, in USNM); 
LaChorrera, Panamd {dalera, in USNM). 

Food plant: Carica papayae (larvae in the fruit). 

Distribution: United States: Florida, Florida City 
(Apr., May), Fort Pierce (May), Miami (Apr., Dec), 
Royal Palm State Park. Puerto Rico: Bayam6n 
(June, Dec), El Yunque (Apr.), Jajoma Alta (June), 
Lares (June, Sept., Nov.). Cuba: Baracoa (July, Aug.) 
Santiago de las Vegas (Feb., Mdr.), Santiago Province 
(Jan., June, Oct.). Haiti: Damien (Aug.). Domini- 
can Republic: San Francisco Mts. (Aug., Sept.). 
Trinidad: Mt. Harris. Guatemala: Cayuga (Apr., 
May), Chejel (June, Aug.), Purulhd (June, July). 
CosTA Rica: Guapiles (May), Juan Vinas (Feb., May, 
June), SUio (May). Ecuador: Quevedo ("Nov.- 
Dec"). 

The types of both caricae and dalera are males, Dyar 
(1919) recognized the generic synonymy of Eucardinia 
and Homalopalpia but never admitted the specific 
identity of their types, although he had every reason 
to suspect it. His designation of new Cuban types for 
caricae in 1918 is obviously invalid, for he had previ- 
ously (1913) designated Florida types for what he ad- 
mitted was the same species under the same name. I 
believe he was correct in his surmise that caricae is the 
same as columnella Zeller, but as I have never examined 
any Colombian examples of any Davara species I 
hesitate to propose the synonymy. 

151. Davara columnella (Zeller), new combination 
Figure 736 

Myelois columnella Zeller, Horae Soc. Ent. Eossicae, vol. 16, 

pp. 209, 210, 1881. 
Piesmopoda columnella (Zeller) Ragonot, Monograph, pt. 1, p. 

161, 1893. 
Homalopalpia columnella (Zeller) Dyar, Ins. Insc. Menstr., vol. 

7, p. 49, 1919. 

I have seen no specimens from the type locality but 
have before me a photograph of the female type which 
agrees with the females of a series from Costa Rica 
identified by Schaus as columnella and coixectly referred 
by Dyar to his dalera, and I have little doubt that the 
names columnella and caricae {= dalera) stand for the 
same species. The female genitalia of Zeller's type 
(here figured) show some trifling differences in the size 
of the plates behind genital opening and in the eighth- 
segment coUar from those of typical caricae; but these 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



75 



are probably only individual differences. Variations 
as great are exhibited among reared examples of caricae 
from different localities. Alar expanse, 15 nun. 

Type locality: Honda, Colombia (type in BM). 

Food plant: Unknown (presumably papaya). 

152. Daveira nerthella (Schaus), new combination 

Figure 738 

Piesmopoda nerthella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 

11, p. 247, 1913. 
Homalopalpia eulhales Dyar, Proc. U. S. Nat. Mus., vol. 47, 

p. 403, 1914. 
Homalopalpia nerthella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7, 

p. 49, 1919. 

Similar to caricae except averaging somewhat larger; 
brush in notch of basal segment of male antenna 
ocherous, the segment itself smaller; the subbasal pale 
shade before the antemedial line more extended on the 
male and much more so on the female, reducing con- 
siderably the blackish brown shading of the basal area 
and forming with the slightly paler antemedial line a 
broad pale pinkish ocherous band; antemedial line 
straight, not angled below cell, its inner and outer 
bordering lines very faint and narrow, reddish brown; 
discal dots separate (never fused) and often only the 
lower one distinguished; on females more or less of an 
ocherous tint over the brownish median shade following 
the antemedial line, especially towards costa. Alar 
expanse, 19-22 mm. 

Male genitalia like that of the following species 
{paranensis) except tuft from near base of sacculus pale 
yellow. Eighth-segment collar without ridges on the 
invaginated portion; narrower on venter than in other 
species; posterior ventral margin without notch or but 
slightly notched. The extent of this notching is indi- 
vidually variable in all the species and is not a reliable 
character for specific separation. 

Type localities: Juan Villas, Costa Rica {nerthella, 
in USNM); Orizaba, M6xico (euthales; in USNM). 

Food plant: Unknown. 

Distribution: M:6xico: Jalapa, Orizaba. Guate- 
mala: Volcfin Santa Maria (June, July). Costa Rica: 
Juan Vinas (Jan.) ; other Costa Rican females without 
further locality designation in Janse Collection. 

Doubtfully distinct from paranensis. The chief dif- 
ferences between the males are in coloration and be- 
tween the females in the width and notching of the 
eighth-segment collar. There is the same amount of 
individual variation in wing color and maculation as in 
other species, some specimens generally paler than 
others, some with the subterminal line distinct, others 
with it almost obsolete. 

Dyar (1919) recognized the synonymy of his euthales 
with nerthella. The type of the former is a male, of 
the latter a female. 

153. Dayara paranensis (Dyar), new combination 
FiGUBB 255 

Homalopalpia paranensis Dyar, Ins. Insc. Menstr., vol. 7, p. 49, 
1919. 



Similar to nerthella except brush in notch of basal seg- 
ment of male antenna brownish; subbasal pale shade 
before antemedial line of forewing much narrower, re- 
stricted (especially on females) by a greater extension 
of the black basal scaling; median area along costa 
rather strongly tinted with ocherous drab or reddish 
ocherous (on the type and one female). Alar expanse, 
18-19 mm. 

Male genitaha with no fine spine cluster surroimding 
the free, forked spine associated with anellus. Tuft 
from near base of sacculus black. Female genitalia 
similar to those of azonaxsalis. 

Type locality: Castro, Parang, Brazil ''type in 
USNM). 

Food plant: Unknown. 

At first glance Dyar's type seems quite distinct from 
males of nerthella, its palpi and antennae being much 
darker and the pale transverse antemedial shade less 
contrasted against the ground color. However the 
specimen is stained and none of the other examples of 
the species before me is in very good condition. Be- 
sides the type, I have before me a female from the type 
locahty, another female from Santa Catarina, Brazil, 
and a male from the British Museum collection from 
Sao Paulo, Brazil, that is without abdomen but a good 
match for Dyar's type. Both it and the type had been 
originally identified by Hampson as columnella Zeller. 

I suspect that when additional South American ma- 
terial is available paranensis will prove to be nothing 
more than a variety of nerthella, and that eventually 
both nerthella and paranensis will fall to azonaxsalis of 
Walker. 

154. Dayara azonaxsalis Walker 
Figure 737 

Davara azonaxsalis Walker, List, pt. 19, p. 1020, 1859. 
Phycita azonaxsalis (Walker) Hampson, in Ragonot, Monograph, 
pt. 2, p. 531, 1901. 

I have seen no specimens of Davara from the type 
locality and none from anywhere of the size of Walker's 
type (30 mm.). A photograph of the type and its geni- 
talia supplied by Tams are before me. The antemedial 
line of forewing shows an angulation between ceU and 
inner margin similar to that on tj^iical caricae. The 
subterminal line is rather distinctly marked and the 
basal area much like that of females of nerthella, but 
not so strongly contrasted. None of these features, 
however, is enough for specific separation. 

The female genitaha show an appreciably wider 
eighth-segment collar than that of nerthella and a dis- 
tinct notch in its ventroposterior margins. The much 
smaller female of paranensis from Castro has similar 
but somewhat smaller genitalia. 

Type locality: Rio de Janiero, Brazil (type in Ox- 
ford Univ. Mus.). 

Food plant: Unknown. 

155. Davara (?) interjecta, new species 
FiouKBS 256, 734 

Male antenna with basal segment enlarged and 
notched as in caricae, but the usual brush of fine bristles 



76 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



in the notch here replaced by smooth, appressed, silvery 
and ocherous scales; first segment of shaft broadly 
flattened, the shaft shortly ciliate (the cilia about as 
long as width of shaft). Labial palpus broadly scaled, 
the second segment reaching well above vertex, deep 
red-brown to blackish brown on outer side, the third 
segment black scaled, at least on inner smiace. Thorax 
deep brown mixed with blackish and buflf scaling. Fore- 
wing reddish brown more or less shaded with black in 
median area, the raised scaling at base mixed black and 
dark red-brown; antemedial line obsolete or very faintly 
indicated on the paler specimens, when distiuguishable 
oblique, straight, narrow, pale buff; lower discal spot 
at end of cell usually distinct, blackish; sub terminal 
line obscure or obsolete; an interrupted row of partially 
confluent black dots along termen. Hind wing trans- 
lucent, white, the veins more or less darkened, a narrow 
fuscous shade along termen. 

Female generally paler than the male. Palpi, head, 
thorax, and forewiug hght reddish brown with a faint 
sprinkling of whitish scales, especially bordering ter- 
men; ground color darkened in median area along costa; 
usually a broad, olivaceous shade along inner margin 
at base; no defined transverse markings. 

Alar expanse, 17-19 mm. 

Male genitalia of the Piesmopoda type with incom- 
plete transtilla developed as a pair of opposed, long, 
slender, cxurved, strongly sclerotized arms, their apices 
curved towards each other; anellus a narrow, broadly 
V-shaped band, with long, curved, slender, haired lat- 
eral arms, their apices bulbed. Female genitaha with 
genital opening simple, no sclerotized plates behind the 
opening. 

Type locality: El Yunque, Luqmllo Mts., Puerto 
Kico (type in Cornell Univ.; paratypes in Cornell and 
USNM, 61332). 

Food plant: Unknown. 

Described from male type and two male and five fe- 
male paratypes from the type locality (1,500-2,000 ft., 
Apr. 22, 23, and Mar. 29, 1930, Cornell lot 795, sub. 38, 
40, and 9, W. T. M. Forbes, collector) and two male 
paratypes from San Francisco Mts., Santo Domingo 
(Sept. 1905, A. Busck, collector). The males, with the 
exception of the holotype and one paratjT)e, are badly 
rubbed. The females are in better condition. 

This species is referred with reservations to Davara. 
In its structural characters it straddles both Davara and 
Piesmopoda and fits comfortably in neither genus. Its 
female genitalia are those of Davara except that the 
usual sclerotized plates behind genital opening are lack- 
ing. Its male genitalia are those of Piesmopoda except 
that the apices of the elements of transtilla point toward 
(rather than away from) each other; and the apices of 
the lateral arms of aneUus are swollen (bulbed) . In all 
known species of Piesmopoda the apices of the elements 
of transtilla point away from each other and the apices 
of the lateral arms of transtilla are pointed. The male 
antenna of interjecta also is abnormal for Davara in that 
there is no brush of fine spines in the notch of the basal 
segment. In my opinion these differences could permit 



generic separation of interjecta from both Davara and 
Piesmopoda; but they are so shght that, without further 
evidence from biology or the early stages, a new generic 
designation does not seem justified at this time. 

156. Davara rufulella (Ragonot), new combination 

FiGiTRBS 257, 739 

Piesmopoda rufulella Ragonot, Bull. Soc. Ent. France, 1888, 
p. cxxxix; Monograph, pt. 1, p. 165, 1893. 

Male antenna simple. Labial palpus upturned, not 
broadly scaled, cylindrical; reaching to slightly above 
vertex on male, nearly to vertex on female; terminal 
segment acuminate. Forewing pale red-brown; the 
basal area a trifle paler with some faint olivaceous shad- 
ing in inner margin; antemedial line oblique, straight, 
obscure, indicated chiefly by its outer dark margin, 
which begins as a blackish smudge on costa and contin- 
ues to inner margin as a slight darkening of the ground 
color; more or less blackish dusting in the cell beyond 
antemedial line; on paler specimens some peppering of 
white scales in median and outer areas, especially on 
midcosta; sub terminal line obsolete or very faintly indi- 
cated; discal spots obscure, confluent along discocellular 
vein. Hind wing translucent white; the veins dark- 
ened; a faint, narrow fuscous shade along termen. Alar 
expanse, 13-16 mm. 

Male genitalia without cluster of fine spines surround- 
ing free spine associated with anellus, the free spine 
itself short, stout, broadly forked. Lateral arms of 
anellus straight. Harpe with apex of cuculius pointed; 
clasper long, stout, strongly curved. Female genitalia 
with two pairs of contiguous sclerotized plates behind 
genital opening; ductus bm-sae sclerotized at genital 
opening. 

Type locality: Puerto Rico (t3^e in Zool. Mus. 
Univ. Berlin). 

Food plant: Unknown. 

Disthibution: Pueeto Eico: Bayam^n (Apr., Sept.), 
Comerio (Nov.), Jajoma Alto (June), Lares (Dec), La 
Sardinera (Dorado, June), Palmas Abajas (June), San 
German (Apr.). 

I have seen no specimens from any but Puerto Eican 
localities. As mentioned in the discussion of the genus, 
rufulella is aberrant in that it lacks the modified basal 
segment of male antenna and the characteristic broad 
scaling of the male labial palpi. However, the genitalia, 
both male and female, are characteristic of the genus, 
showing only specific differences from other species of 
Davara. 

38. Genus Sarasota Hulst 

Sarasota Hulst, Journ. New York Ent. Soc, vol. 8, p. 222, 1900, 
(Type of genus: Sarasota plumigerella Hulst). 

Cuba Dyar, Ins. Insc. Menstr., vol. 7, p. 50, 1919. (Type of 
genus: Cuba furculella Dya,!. New synonymy.) 

Tongue well developed. Antenna simple in both 
sexes, shaft weakly pubescent. Labial palpus up- 
turned, reaching to slightly above vertex; third segment 
slightly over half the length of second, acuminate. 
MaxUlary palpus small, squamous. Forewing smooth; 
11 veins; vein 2 from before but near lower outer angle 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



77 



of cell; 3 from the angle, approximately equidistant at 
base from 2 and 4, 4 and 5 approximate for a short 
distance from cell; 6 from upper angle of cell, slightly 
bent towards base, connate with the stalk of 8-9; 10 
from the cell, well separated from the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from before 
lower outer angle of cell; 3 from the angle, nearly as 
long as 2, connate with 4; all veins long; 4 and 5 con- 
tiguous or weakly anastomosed for about half their 
lengths from cell; 7 and 8 wealdy anastomosed beyond 
cell, their free elements long ; cell slightly less than one- 
fourth the length of wing; discocellular vein curved for 
a short distance from lower angle, thence vertical. 
Eighth abdominal segment of male with sternite de- 
veloped as a narrow, sclerotized pocket. 

Male genitalia similar to those of typical Davara 
except: Free spine associated with anellus entirely 
lacking; harpe with two-pronged clasper and apical end 
of sacculus produced; penis with a pair of curved, flat- 
tened, bladelike cornuti. Female genitalia with ductus 
bursae sclerotized for a short distance from genitalia 
opening; a single, angulate, projecting plate behind 
genital opening; otherwise as in Davara. 

157. Sarasota plumigerella Hulst 
Figure 258 

Sarasota plumigerella Hulst, Journ. New York Ent. Soc, vol. 8, 
p. 222, 1900.— Grossbeck, Bull. Amer. Mus. Nat. Hist., 
vol. 37, p. 128, 1917.— McDunnough, Check list. No. 6120, 
1939. 

Basal segment of antenna red spotted with black. 
Prothorax wine red ; mesothorax and metathorax black. 
Forewing black at base followed by a whitish ocherous 
shade; antemedial line oblique, straight, ocherous 
white, preceded by more blackish scaling, followed (es- 
pecially on costa) by a strong black shade; remainder 
of wing blackish fuscous stained mth wine red, the 
costa on outer half distinctly reddish; sub terminal line 
very faint, sinuate; discal spots obscm-e or absent; 
terminal dots more or less confluent, blackish; cilia red. 
Hind wing dull, translucent white; a dark shade at 
apex and a narrow dark line along termen. Mid tibia 
with a strong yellow hair tuft. Alar expanse, 11-14 
mm. 

Male genitalia with terminal margin of vinculum not 
produced at the edges but exhibiting no other specific 
characters. Female genitalia slightly smaller but other- 
wise not distinct from those oi furculella. 

Type locality: Palm Beach, Fla. (type in USNM). 

Food plants: Laguncularia racemosa, Coccolobis 
uvvfera Garvae feetUng under a light silk webbing on 
the leaves and flower buds). 

Distribution: Florida, Pahn Beach, Ramrod Key 
(Apr.), Stock Isl. (Apr.), Sugar Loaf Key (Apr.). 

158. Sarasota furculella (Dyar), new combination 
Figures 48, 259, 740 
Cuba furculella Dyar, Ins. Insc. Menstr., vol. 7, p. 50, 1919. 

Larger and paler than plumigerella. Thorax and 
forewing violaceous gray more or less tinted with red- 



dish; the blackish shade of plumigerella replaced in 
furculella by red; antemedial line dull white with a 
strong ocherous tint, especially towards inner margin, 
its dark outer border red; some white dusting in median 
costal area; discal dots more distinct, separate, red or 
fuscous ; sub terminal line more distinct, narrow, sinuate, 
dull white; terminal dots few, but distinct and separated, 
blackish. Hind wings white with a faint smoky tint; 
the veins very shghtly darkened. Midtibial hair tuft 
as in plumigerella. Alar expanse, 14-16 mm. 

Male genitalia with terminal margin of vinculum 
slightly produced at the sides; otherwise as in plumi- 
gerella. Female genitalia a trifle larger than those of 
plumigerella but showing no specific characters. 

Type locality: Santiago, Cuba (type in USNM). 

Food plant: Unknown. 

Distribution: Cuba: Baracoa, Matanzas, Santiago 
(June). Puerto Rico: Dorado (May), Puerto Real 
(Vieques Isl., Apr.). Dominica (Dec). Virgin Is- 
lands: Kingshill (St. Croix, June, Dec). 

Very close to plumigerella but apparently a distinct 
species, separable chiefly on color. 

159. Sarasota ptyonopoda (Hampson), new combination 

Phycita ptyonopoda Hampson, Ann. Mag. Nat. Hist., ser. 6, vol. 
16, p. 347, 1895. 

Hyalospila ptyonopoda (Hampson) Hampson, in Ragonot, Mono- 
graph, pt. 2, p. 530, pi. 56, fig. 9, 1901. 

I have seen no examples of this species. From the 
description and the figure in the Ragonot Monograph 
the coloration and markings must be similar to those of 
furculella. According to Hampson the male has a 
strong, long tuft of scales from the hind femur. Clarke 
has reexamined the type and teUs me that there is a 
large expanded dorsal tuft from the base of the hind 
femur and that there is no tuft on the midtibia. These 
differences in tufting distinguish the species easily from 
either plumigerella or furculella. A fine photograph of 
the male genitalia, taken by Clarke, is before me. 
They are like those oi furculella Dyar. Unfortunately 
the abdomen had been glued on the type so there will 
remain some question of placement for the species until 
another male from the type locality and with tufted 
hind femora is recorded. Alar expanse, 16 mm. 

Type locality: St. Vincent, Windward Islands, 
British West Indies (type in BM). 

Food plant: Unloiown. 

In the U. S. National Museum is a rubbed male from 
Jalapa, M6xico, labeled in Hampson's handwriting 
"Phycita ptyonopoda Hampson," but it cannot be that 
species. What remain of the legs show a yellow hair 
tuft on midtibia and no trace of tuft on the hind femur 
(the hind tibiae are missing). This specimen is an 
Atheloca sp., close to bondari. 

39. Genus Piesmopoda Zeller 

Piesmopoda Zeller, Isis von Oken, 1848, p. 863. — Hulst, Phy- 
citidae of N. Amer., p. 132, 1890. — Ragonot, Monograph, 
pt. 1, p. 158, 1893. (Type of genus: Piesmopoda rubicun- 
deUa Zeller.) 



78 



XnsriTED STATES NATIONAL MUSEUM BULLETIN 207 



Discopalpia Ragonot, Monograph, pt. 1, p. 167, 1893. (Tjrpe of 
genus: My elois flavicans Zeller. New synonymy.) 

Amphycitopsis Dyar, Ins. Insc. Menstr., vol. 7, p. 45, 1919. 
(Type of genus: Amphycitopsis Isabella Dyar. New syn- 
onymy.) 

Tongue well developed. Antenna pubescent or 
shortly ciliate (the cilia no longer than width of shaft) ; 
on male, shaft simple, notched at base or with curved 
excavation (a long sinus) towards base; basal segment 
of male sometimes swollen but not notched or otherwise 
modified. Labial palpus upturned, of varying length, 
not reaching vertex (isabella, fratella) or extending to or 
above vertex; third segment acuminate or {ragonoti) 
broadly dilated with scales. MaxiUary palpus minute 
(folded over tongue), filiform or subsquamous (third 
segment sUghtly dilated with scales) . Forewing smooth ; 
venation as in Sarasota except vein 3 normally consider- 
ably closer to 4 at base than to 2 ; male without costal 
fold. Hind wing as in Sarasota. Eighth abdominal 
segment with stemite developed as a narrow, sclero- 
tized pocket. 

Male genitalia with uncus bifid. Gnathos incom- 
plete, represented by its broad lateral arms, separated 
at their apices, between which lies a rather well sclero- 
tized subanal plate. Transtilla well developed but 
incomplete, consisting of a pair of long, very slender, 
curved arms whose sharply pointed apices are directed 
away from each other. Harpe with a strong, hooked 
clasper from near apex of sacculus; a strong hair tuft 
from lobe near base of sacculus. AneUus a curved plate 
(U- or V-shaped) with long, slender, haired and bluntly 
pointed lateral arms. Penis armed with a narrow, 
flattened and more or less curved, bladelike cornutus; 
sometimes with two such comuti. 

Female genitalia with single signum or none, signum 
when present developed as a small granulate depres- 
sion; ductus bursae with genital opening simple or more 
or less sclerotized, but without the paired cupHke plates 
found in Davara; ductus seminalis from bursa near its 
junction with ductus bursae. Eighth-segment collar 
more or less invaginate (except in apocerastes and 
montella) . 

A distinct genus defined and easUy recognized by its 
male and female genitaUa. As used by Ragonot and 
subsequent authors the name covered a composite of 
disparate elements. The antennal and palpal char- 
acters upon which Piesmopoda, Discopalpia, and Am- 
phycitopsis were distinguished are in this instance of no 
more than specific significance. They are discussed 
more fully under their type species. 

The genus is apparently limited to tropical America. 
Nothing is known of the life history or host association 
of any of the species. 

160. Piesmopoda rubicundella Zeller 
FiGUBE 260 

Piesmopoda rubicundella Zeller, Isis von Oken, 1848, p. 864. — 
Ragonot, Monograph, pt. 1, p. 160, 1893. 

Antenna of male with basal segment swollen, triangu- 
late; shaft notched at base. Labial palpus slender, 



third segment acuminate, reaching well above vertex. 

Forewing pale (olivaceous ocherous) at base and for 
some distance along inner margin; the median costal 
area broadly white, heavily dusted with red scaling; 
more or less of this red dusting also on costal half at 
base; antemedial line not distinguishable; sub terminal 
Une distinct, straight, oblique, pale reddish or ocherous 
gray bordered by dark purplish fuscous lines; the entire 
outer fourth of wing more or less suffused with purplish 
fuscous; discai dots more or less distinct, separated, red 
or reddish fuscous. Hind wing pale smoky fuscous, 
lighter towards base and semitransparent; veins dis- 
tinctly darkened. Alar expanse, 13 mm. 

Male genitaUa having harpe with cucuUus narrowly 
elongate, evenly tapering. Anellus a broadly sclero- 
tized, V-shaped band. 

Type locality: BrazU (type in Mus. Univ. BerUn) . 

Food plant: Unknown. 

I have examined the male type but have seen no 
other examples from Brazil. The type is not distin- 
guishable from males of the following species (xanthom- 
era) except for minor differences in male genitalia, 
which may or may not be significant. 

161. Piesmopoda xanthomera Dyar 

Figures 266, 745 

Piesmopoda xanthomera Dyar, Proe. U. S. Nat. Mus., vol. 47, 

p. 332, 1914. 
Piesmopoda xanthozona Dyar, Ins. Insc. Menstr., vol. 7, p. 45, 

1919 (new synonymy). 

With the same male characters as rubicundella and 
superficially not distinguishable from it. I suspect that 
xanthomera is nothing more than a synonym or variety 
of rubicundella, but this cannot be proven imtil females 
of the latter from Brazil are available. Dyar's two 
names apply only to the sexes {xanthomera to the females 
and xanthozona to the males). Alar expanse, 13-17 mm. 

Male genitalia, figured from type of xanthozona, differ 
from those of rubicundella chiefly ia the stronger tuf tiug 
from the base of the harpe, a character of very doubtful 
specific value. Female genitalia, figured from type of 
xanthomera and checked with those of females from all 
localities here cited, distinguished chiefly by the shape 
of the eighth-segment coUar and its sclerotized invagi- 
nate portion and the pair of corrugate patches on the 
ventral siu-face of the latter. Bursa with signum. 
Genital opening simple. 

Type localities: La Chorrera, Panamd {xanthomera, 
in USNM); St. Jean Maroni, French Guiana {xantho- 
zona, in USNM). 

Food plant: Unknown. 

Distribution: Guatemala: Cayuga (Apr., May). 
Costa Rica: San Jos^ (July). PanamI: La Chorrera 
(May), Porto Bello (Oct.). French Guiana: Cayenne, 
St. Jean Maroni. 

162. PieBmopoda trichomata (Zeller) 

Figure 744 

Myelois trichomata Zeller, Horae Soo. Ent. Rossicae, vol. 16, p. 
194, 1881. 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



Ragonot (Monograph, pt. 1, p. 160) makes this a 
synonym of rubicundella, considering it merely the fe- 
male of the latter. This is a very dubious placement. 
The Zeller type material in the British Museimi con- 
sists of two female cotypes (photographs of which are 
before me) alike in all details and similar to rubicundella 
except that the antemedial line is indicated by broken 
remnants of its outer border, a dark dash from costa, 
and a more or less diffused dark spot on lower margin of 
cell. The lower half of wing is somewhat suffused into 
dark shading and contrasted against the whitish mid- 
costal area, as in rubicundella. Zeller's figure is mis- 
leading in that it shows much of the wing bright yellow, 
as mfloricans. Alar expanse, 13 mm. 

The genitaha of the female cotype here figured, ac- 
cording to Tams and Clarke, agree in all details with 
those of the other cotype. The latter should be con- 
sidered the holotype, as it is the better preserved speci- 
men. Bm-sa with signum. Genital opening simple. 
The narrow eighth-segment collar easily identifies the 
species. It is imlike any other in the genus that I have 
seen. 

Type locality: Honda, Colombia (type in BM). 

Food plant: Unknown. 

Known only from the type specimens. 

163. Piesmopoda flavicans (Zeller) 
Figures 262, 746 

Myelois flavicans Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 193, 

1881. 
Discopalpia flavicans (Zeller) Ragonot, Monograph, pt. 1, p. 167, 

1893 (in part, ?). 
Piesmopoda flavicans (Zeller) Dyar, Proc. U. S. Nat. Mus., vol. 

47, p. 333, 1914. 
Piesmopoda fratella Dyar, Ins. Insc. Menstr., vol. 7, p. 45, 1919 

(new synonymy). 

Antenna of male simple. Labial palpus slender; on 
male not reaching vertex; on female extending sUghtly 
above vertex (as in Isabella). 

Forewing bright yellow; outer third purple dusted 
with blackish towards apex; the yellow ground color ex- 
tended further outward on costa and inner margin than 
at middle; antemedial line obsolete; subterminal hne 
faint, narrow, whitish, straight, oblique and close to 
outer margin. Hind wing whitish, stained with smoky 
fuscous towards apex and on the outer parts of the veins. 
Alar expanse, 14-15 mm. 

Male genitalia like those of xanthopolys except that 
elements of transtUla are stouter, lateral arms of anellus 
are bent sharply away from each other, and terminal 
margin of vinculum is acutely rounded (rather than 
straight). Female genitalia similar to those of xantho- 
polys except for slight differences in the structure of 
eighth-segment collar. 

Type localities: Honda, Colombia (flamcans, in 
BM); St. Jean Maroni, French Guiana (fratella, in 
USNM). 

Food plant: Unknown. 



Distribution: French Guiana: Cayenne, St. Jean 
Maroni (Mar.). Colombia: Honda, 

Specimens of five different species in the U. S. Na- 
tional Collection had been identified by Hampson, 
Dyar, and Schaus as flavicans. Among them was one 
female from French Guiana which Hampson identified 
(correctly) as flavicans and which Dyar later included in 
his series oi fratella. Clarke and Tams have checked 
our genitalic figures of the several Piesmopoda species 
with the female type of flavicans in the British Museum 
and inform me that the genitalia oi fratella agree in all 
details with those oi flavicans. Ragonot also misidenti- 
fied Zeller's species (at least as far as males are con- 
cerned) and on the strength of their pecuUar male palpi 
erected the genus Discopalpia, with flavicans as type. 
Dyar (1914) noted the misidentification and renamed 
the males of flavicans Ragonot (not Zeller) as Discopal- 
pia ragonoti. Later evidence from genitalia justifies 
the new specific name; but nomenclatorily the type of 
Discopalpia must remain flavicans Zeller, and Dyar's 
citation of a new type {flavicans ^&gonot= ragonoti 
Dyar) is not justified, despite Ragonot's misidentifica- 
tion and the characters derived therefrom for his genus. 
Fortimately these characters (as far as Piesmopoda are 
concerned) are of specific significance only, so Disco- 
palpia would fall, however we interpreted its type; 
but I for one hold that when a specific name is cited as 
type of a new genus, that species remains the type of 
the genus regardless of what specimens were before the 
author of the genus or how he described his generic 
concept. The decision is one of nomenclature and 
not of zoology. 

164. Piesmopoda ragonoti (Dyar), new combination 
Figures 265, 747 

Discopalpia ragonoti Dyar, Ins. Insc. Menstr., vol. 7, p. 44, 1919. 
Discopalpia flamcans Ragonot (in part, cf; not Zeller), Mono- 
graph, pt. 1, p. 167, 1893. 

Antenna of male simple; basal segment cylindrical; 
shaft without notch or other modification. Labial 
palpus very long, the second segment reaching well 
above vertex in both sexes; on male the third segment 
broadly expanded with long scales, fan shaped ; a strong 
admixture of black scaling on outer sides of the palpi of 
both sexes. 

Forewing as in flavicans. Alar expanse, 14-16 mm. 

Male genitalia distinguished chiefly by the very 
heavy, black, broad-haired tuft from base of sacculus of 
harpe. Female genitalia with signimi; genital opening 
simple; distinguished from those of other yellow-winged 
species by minor differences in the configuration of the 
eighth-segment collar; closest to those of xanthopolis 
Dyar. 

Type locality: Cayuga, Guatemala (type in 
USNM). 

Food plant: Unknown. 

Distribution: Mexico: Distrito Federal. Guate- 
mala: Cayuga (Feb., Apr., May). Costa Rica: Juan 
Vinas (Jan.). 



80 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



165. Piesmopoda Isabella (Dyar), new combination 

FiGUHEs 264, 749 

Amphyciiopsis Isabella Dyar, Ins. Insc. Menstr., vol. 7, p. 45, 
1919. 

Male antenna simple. Labial palpus short, hardly 
reaching vertex on male and but a trifle beyond vertex 
on female. 

Forewing as ia flavicans. Alar expanse, 18-20 mm. 

Male genitalia with cucullus of harpe subtriangulate, 
harpe wider in proportion to its length and less evenly 
tapering than in other species except apocerastes which 
has similar male genitalia. Female genitalia with 
signum; genital opening simple; distinguished by the 
configuration of the eighth-segment collar. 

Type localitt: Juan Vinas, Costa Kica (type in 
USNM). 

Food plant: Unknown. 

Represented by the male type and a shghtly larger 
female from the type locahty (Jan.). The latter had 
been identified by Schaus as "Piesmopoda flavicans 
Zeller." The species served as type for Dyar's genus 
Amphyciiopsis, erected solely on the basis of the short 
labial palpi of the male. 

166. Piesmopoda xanthopolys Dyar 
Figures 261, 748 

Piesmopoda xanthopolys Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 
332, 1914. 

' Male antenna simple. Labial palpi slender, extend- 
ing above vertex in both sexes, somewhat longer on 
female than on male. 

Forewing as in flavicans. On the female from the 
type locality the purplish shading on the outer area of 
the forewing is somewhat more extended and the yellow 
area of the wing proportionally more restricted than in 
Havicans; but this difference seems to be individual 
rather than specific. In xanthopolis as well as the other 
species with coloration similar to flavicans the extent of 
piu^jle shading is variable between the sexes and even 
among individuals of one sex. Alar expanse, 13-16 mm. 

Male genitalia figured from specimen from La Chor- 
rera, Panamd. Their most obvious feature seems to 
be the rather short arms of the bifid uncus (proportion- 
ally shorter than those of any other Piesmopoda except 
flavicans which has stouter transtilla and differently 
shaped anellus and vinculum). Female genitalia with 
signum present. Genital opening simple. Very close 
to those of flavicans, with incurvation of posterior- 
ventral margin of eighth-segment collar similar and ex- 
hibiting only minor differences in the collar otherwise. 

Type locality: Porto Bello, Panamd (type in 
USNM). 

Food plant: Unknown. 

Disteibution: Panama: Corozal (July), La Chor- 
rera (May), Porto Bello (Sept., Dec). 

Known only from the original type series. Dyar in 
1919 (Ins. Insc. Menstr., vol. 7, p. 44) placed xantho- 
polys in the sjmonymy oi flavicans. The two species 
are very close, but apparently distinct. 



167. Piesmopoda parva, new species 
FiGTJBES 263, 750 

Male antenna simple except for a very shght incm-- 
vation of the shaft towards base. Labial palpus slender, 
reaching vertex; terminal segment acuminate. 

Forewing yellow with a slight ohvaceous tint; costa 
rather broadly margined from base to near apex with 
white faintly peppered with red scaling; a few red and 
black scales at extreme base and for a short distance 
from base along inner margin; no antemedial line; sub- 
terminal line straight, oblique, close to termen, with 
narrow purphsh fuscous borders and preceded by a fus- 
cous shade which extends, triangularly, almost to the 
cell. Hind wings translucent white with a faint smoky 
tint, darkening slightly towards apex. Alar expanse, 
10-11 mm. 

Male genitalia distinguished by the slender arms of 
the divided uncus and the slender, naillike spine asso- 
ciated with anellus. Female genitalia with signum; 
genital opening simple. Distinguished by the broad 
and deep excurvation in posteroventral margin of the 
eighth-segment collar. 

Type locality: La Chorrera, Panamd (type in 
USNM, 61333). 

Food plant: Unknown. 

Described from male type from the type locahty 
(May), one male paratype from Cabima, Panamd (May 
1911), and one female paratype from Taboga Isl., 
Panamd (Feb. 1912), all collected by A. Busck. These 
specimens had been included by Dyar among his para- 
types of Piesm,opoda xanthomera. They are somewhat 
rubbed but otherwise in good condition. The species 
is the smallest of the Piesmopoda. 

168. Piesmopoda semirufella (Zeller) 
Figure 752 

Myelois semirufella ZeUer, Horae Soc. Ent. Rossicae, vol. 16, p. 
196, 1881. 

Piesmopoda semirufella (Zeller) Ragonot, Monograph, pt. 1, p. 
160, 1893.— Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 332, 
1914; Ins. Insc. Menstr., vol. 7, p. 46, 1919. 

Several different species have been identified as semi- 
rufella. I have seen but one example that can be defi- 
nitely placed to Zeller's name, a female from Cayuga, 
Guatemala, collected by Schaus and Barnes (Jan.). 
The genitaha of this specimen have been checked by 
Clarke with the genitalia of the type of semirufella and 
he finds them identical. They have the signum present 
and a broad, strongly sclerotized plate at genital open- 
ing. The latter structure at once identifies the female. 
Males of the species have not been properly associated. 

Superficially, semirufella is not distinguishable from 
females of apocerastes Dyar. Indeed, several females of 
the latter in both the British Museum and U. S. Na- 
tional Collections had been identified as Zeller's species. 
Alar expanse, 16 mm. 

Type locality: Colombia (type, ?, in BM). 

Food plant: Unknown. 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



81 



169. Piesmopoda apocerastes Dyar 
Figure 751 

Piesmopoda apocerastes Dyar, Ins. Insc. Menstr., vol. 7, p. 45, 
1919. 

Male antenna with basal segment cylindrical, slender ; 
shaft with a long sinus (involving about eight of the 
basal segments) lined by flattened blackish scales which 
terminate in a slight tuft at the outer extremity of the 
sinus. Labial palpus cylindrical, slender, reaching to 
slightly above vertex. 

Forewing, except for a whitish border along costa, 
suffused reddish brown to the naked eye, very slightly 
darkened towards outer margin (under magnification 
the groimd color shows a strong under tinting of oliva- 
ceous ocherous) ; whitish costal border peppered with 
scattered red scaling, a concentration of these along ex- 
treme costal margin; discal dots separate, red; ante- 
medial line obsolete or, at most, faintly indicated on 
some specimens by an obscure, narrow, dark, trans- 
verse shade ; subterminal line faint, weakly bordered by 
narrow dark (reddish fuscous) lines. Hind wings smoky 
white to pale smoky fuscous, darkening towards apex 
and outer margin; the veins darkened. Alar expanse, 
15-16 mm. 

Male genitalia similar to those of Isabella Dyar. Fe- 
male genitalia without signum; a naiTow, strongly scle- 
rotized plate at genital opening with weakly sclerotized, 
anterior, lobelike projection. 

Type locality: Juan Vinas, Costa Kica (type in 
USNM). 

Food plant: Unknown. 

Distribution: Mexico: Jalapa. Costa Rica: Juan 
Vinas (May, Nov.). Dominica (British West Indies, 
Feb.). French Guiana: St. Jean Maroni. Brazil: 
Parand, Castro. 

A distinct species easily identified by its female geni- 
talia. Several of the females before me in the National 
Collection had been identified by Hampson as Piesmo- 
poda semirufella. The genitalia of female specimens 
from all the above-mentioned localities have been 
checked. 

170. Piesmopoda montella Schaus 
Figure 743 

Piesmopoda montella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 
11, p. 247, 1913. 

Labial palpus of female slender, reaching shghtly 
higher than vertex. 

Forewing light olivaceous brown; costal margin to 
subterminal line broadly white irrorated with red- 
brown, the extreme costal edge black at base, reddish 
brown at middle ; no trace of any antemedial line ; sub- 
terminal line slightly outcurved below vein 6, bordered 
inwardly and outwardly from costa to vein 2 by blackish 
bands into which some reddish scales are intermixed; 
discal dots separated, reddish brown; a few blackish 
dots on terminal margin. Hind wing pale, semihyaline 
brown, darkening towards outer margin; the veins 
darkly outlined. Alar e.xpanse, 24 mm. 

Female genitalia without signum; bursa copulatrix 



small, oblong; ductus bursae very short and broad, al- 
most as broad at middle as the bursa, weakly sclerotized 
at genital opening, finely sclerotized otherwise. 
Eighth-segment collar simple, not fused ventrally. 

Type locality: Moimt Pods, Costa Rica (May; 
type in USNM). 

Food plant: Unknown. 

A distinct species easily identified by its large size 
and distinctive genitalia; represented only by the fe- 
male type. 

Genus 40: Atheloca 

[Venational division C. Forewing with veins 4 and 5 connate 
and contiguous or partially anastomosed for one-third their 
lengths from cell; 6 from upper angle of cell, connate with the 
stalk of 8-9, bent towards base. Hind wing with cell less than 
one-fifth the length of wing; discocellular vein oblique, straight. 
Male genitaha with uncus hoodlike; lateral arms of gnathos fus- 
ing into aneUus; harpe broadly angled at base of cucullus.] 

40. Atheloca, new genus 

Type of genus: Nephopteryx subrufella Hulst. 

Tongue well developed. Antenna of male simple, 
shaft pubescent. Labial palpus slender, upturned, 
reaching to or a trifle above vertex; third segment 
shorter than second, acuminate. Maxillary palpus fili- 
form. Forewing smooth; 11 veins, vein 2 from before 
lower outer angle of cell ; 3 from the angle, much nearer 
to 4 at base than to 2; 4 and 5 connate, contiguous or 
partially anastomosed beyond base for one-third of 
their lengths; 6 from upper angle of cell, curved towards 
base and connate with the stalk of 8-9 ; 10 from the cell, 
approximate to the stalk of 8-9; male without costal 
fold. Hind wing with vein 2 from before lower outer 
angle of cell; 3 from the angle, nearly as long as vein 2, 
closely approximate to the stalk of 4-5 at base; 4 and 
5 stalked for half their lengths; 7 and 8 anastomosed 
for most of their lengths beyond cell, the free element 
of vein 8 very short; cell less than one-fifth the length 
of wing; discocellular vein oblique, straight. Abdomen 
of male with a pair of invaginated hair tufts at base; 
sternite of eighth segment developed as a sclerotized 
digitate pocket. 

Male genitalia Avith uncus hoodlike, triangulate, 
densely haired on outer surface. Gnathos represented 
only by its lateral arms which fuse into anellus at their 
apices. Transtilla absent. Harpe broadly angled at 
base of cucullus. AneUus a stout, triangulate plate 
with strongly sclerotized, sharply out-curved, smooth 
lateral arms (these latter may possibly represent ele- 
ments of a divided transtilla fused with the anellus, but 
this is extremely doubtful). Aedeagus and penis simple. 
Vinculum longer than greatest width, but slightly taper- 
ing to truncate terminal margin. 

Female genitalia with signa present in the form of 
two small scobinate patches ; ductus bursae with a small, 
weakly sclerotized collar near the jimction with bursa 
copulatrix; genital opening simple, ductus seminalis 
from bursa near the junction of bursa and ductus bur- 
sae; eighth-segment collar completely fused ventrally. 



82 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



171. Atheloca subrufella (Hulst), new combination 

FiGTJBBs 267, 741 

Nephopteryx subrufella Hulst, Ent. Amer., vol. 3, p. 132, 1887. 
Nephopteryx filiolella Hulst, Ent. Amer., vol. 4, p. 117, 1888 (new 

synonymy). 
Piesmopoda subrufella (Hulst) Hulst, Phycitidae of N. Amer., 

p. 133, 1890.— Ragonot, Monograph, pt. 1, p. 166, 1893.— 

Grossbeek, Bull. Amer. Mus. Nat. Hist., vol. 37, p. 129, 1917. 
Piesmopoda filiolella (Hulst) Hulst, Phycitidae of N. Amer., p. 

133, 1890. — Ragonot, Monograph, pt. 1, p. 166, 1893. 
Sarasota subrufella (Hulst) Barnes and McDunnough, Check list 

of the Lepidoptera of Boreal America, No. 5558, 1917. — 

McDunnough, Check list. No. 6121, 1939. 
Sarasota filiolella (Hulst) Barnes and McDunnough, Check List 

of the Lepidoptera of Boreal America, No. 5559, 1917. — 

McDunnough, Check list. No. 6122, 1939. 
Hyalospila ptychis Dyar, Ins. Inso. Menstr., vol. 7, p. 49, 1919 

(new synonymy). 

Forewing ocherous fuscous shaded with reddish or 
purplish red except along costa; costa at extreme base 
edged with black, between the transverse lines rather 
broadly bordered by dull white sprinkled with red 
scales; antemedial line sometimes obsolete, when pres- 
ent indicated by a transverse black band interfused with 
reddish and preceded by a narrow dusting of white 
scales; sub terminal Hne faint, pale, bordered inwardly 
and outwardly by narrow blackish or purphsh red 
bands; discal spots usually distinct (at least the lower 
one), well separated, blackish (rarely with a touch of 
red) ; a more or less distinct row of blackish dots along 
termen. Hind wing smoky white, translucent; the 
veins darkened and a distinct dark shade along termen. 
Midtibia of male with strong hair tuft from base on 
inner side. Hind tibia of male without appreciable 
hair tuft. Alar expanse, 12-19 mm. 

Male genitaha as given for the genus. Female geni- 
taUa with sclerotized portion of ductus bursae very nar- 
row. 

Type localities: Florida (subrufella, in AMNH, ex 
Rutgers); "Texas" (filiolella, in AMNH, ex Rutgers); 
Cuba (ptychis, in USNM). 

Food plant: Unknown. 

Distribution: United States: Florida, Chokolos- 
kee, Christmas Harbor (Mar.), Duardia (May), Fort 
Drum, Hastings (Aug., Sept.), Lake Alfred (May), 
Paradise Key (Mar.), Royal Palm State Park (June, 
Sept.), Stemper (May, July, Aug.), Vero Beach (Apr.), 
Winter Park (July, Aug.). Cuba: Pinar del Rio, no 
specific locality (type of ptychis). Virgin islands: 
Kingshill (St. Croix, "Nov.-Dec"). 

The types of both subrufella and filiolella are females, 
neither of which bears a locahty label; subrufella bears 
a label containing only the number "Ql" ; filiolella only 
a date label "March." The latter is without abdomen 
and in very poor condition; but exhibits no difference 
from the type of subrufella except its somewhat larger 
size. In the National Museum there is a female from 
Christmas Harbor ("March") labeled by Hulst "Ne- 
phopteryx filioleUa, type." It is an exact match for the 
type in the Rutgers Collection, as are three other large 
females from Florida in the National Collection, obvi- 
ously the same as subrufella. We have a large series of 



the species but no Texas examples, and I doubt very 
much the correctness of Hulst's citation as the type 
locality of his filiolella. The species is obviously a 
tropical one which has extended its range to Florida. 
Dyar's ptychis is merely a Cuban example, differing in 
no wise from typical subrufella. 

172. Atheloca bondari, new species 

Hyalospila ptychis Bondar (not Dyar), Rev. de Ent., Brazil, vol. 
11, p. 199, 1940. — Lepesme, Les insectes des Palmier's, p. 
343, 1947 (Paris). 

Similar to subrufella except that the pale costal area 
of forewing is less contrasted and conspicuous. The 
genitalia male and female exhibit no essential differences 
from those of subrufella. The one distinguishing char- 
acter, seems to be a strong, dorsal, yellow hair tuft from 
the base of the male hind tibia. This is lacking from 
all specimens of subrufella and is an addition to the 
similar tuft on male midtibia, present on both subrufeUa 
and bondari. Alar expanse, 15-16 mm. 

Type locality: Baia, Brazil (type m USNM, 61335). 

Food plant: Cocos nucifera (Lepesme also records 
O. coronata, O. vagans, Attalea funifera, and A. piassa- 
bossu). 

Described from male type and three male and five 
female paratypes aU from the type locahty and reared 
(June and July 1939, imder Bondar Nos. 2521 and 2561) 
from larvae feeding in the seeds and at the base of the 
fruits of Cocos nucifera. These were received from Dr. 
Gregorio Bondar, for whom the species is named. He 
gives a good account of the habits of the species in the 
above-cited paper. I am responsible for the misidenti- 
fication to ptychis, for at the time I overlooked the differ- 
ence in leg tuftings between ptychis and the Brazilian 
specimens. 

Genera 41 and 42: Praedonula and Peadus 

IVenational division D. Forewing with veins 4 and 6 closely 
approximate for a short distance from cell; vein 6 straight; 10 
from the cell. Hind wing with cell one-fourth to one-fifth the 
length of wing; discocellular vein more or less curved. Male 
antenna with a shallow sinus in shaft involving the first half 
dozen segments. Male genitalia with a stout hair tuft from near 
base of sacculus.) 

41. Praedonula, new genus 

Type of genus: Phydta almoneUa Dyar. 

Tongue well developed. Male antenna with a shallow 
sinus in shaft at base (involving the first six segments), 
the sinus containing a scattering of minute papiQalike 
setae and a narrow ridge of scales along its outer edges, 
otherwise pubescent. Labial palpus very slender, 
upturned, reaching to or shghtly above vertex; third 
segment slightly shorter than second, acuminate. 
Maxillary palpus subsquamous (the scales on second 
segment expanded, on third rather long and drawing to 
a point). Forewing smooth; 11 veins; vein 2 from 
before but near lower outer angle of ceU (nearer the 
angle in male than in female); vein 3 from the angle; 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



83 



4 and 5 closely approximate at base and for a short 
distance beyond; 6 straight, from slightly below upper 
angle of cell (male) or from the angle (female) ; 10 from 
the cell, closely approximate to the stalk of 8-9 (male) 
or slightly separated from it (female) ; male without 
costal fold. Hind wing with vein 2 from before lower 
outer angle of cell; 3 from the angle, nearly as long as 
2, approximate at base to the stalk of 4-5; 4-5 stalked 
for at least half their lengths (female), somewhat longer 
stalked on male; 7 and 8 contiguous or weakly anas- 
tomosed for a short distance beyond cell, the free 
element of 8 long; cell short, about one-fifth the length 
of wing; discoceUular vein oblique, very slightly curved. 
Eighth abdominal segment with sternite developed as 
a narrow (digitate) sclerotized pocket. 

Male genitalia with uncus hoodlike, roimded. Gna- 
thos strongly developed, the lateral arms broad, ex- 
panded and curled at their extremities and supporting 
a sclerotized subanal plate with a short thornlike spur 
at its base. Transtilla absent. Harpe stout, simple, 
slightly broadened at middle; at base of cucullus a 
stout hair tuft. AneUus a heart-shaped plate with 
stubby lateral lobes. Penis unarmed. Vinculum stout 
(but sclerotized narrowly along its margins); slightly 
longer than broad; scarcely tapering. 

Female genitalia with signa present in the form of two 
small granulate patches; ductus bursae with genital 
opening surrounded by a sclerotized plate; ductus 
seminahs from btrrsa near junction of bm-sa and ductus 
bursae. Eighth-segment collar completely fused 
ventraUy. 

This genus is obviously closely related to the preced- 
ing genus (Atheloca) and that which follows (Peadus). 
From the former it differs in the male antennal character 
and the weak anastomosis of veins 7-8 of hind wing. 
From Peadus it differs chiefly in the simple (imdivided) 
harpe and the much more strongly developed uncus and 
tegumen of its male genitalia. It has no close relation- 
ship to the Old World Phycita to which Dyar referred 
its type species. 

173. Praedoniila almonella (Dyar), new combination 

Figures 47, 268, 742 

Phycita almonella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 333, 
1914. 

The type series (a male and two females) are some- 
what rubbed and the coloration and markings of fore- 
tving consequently obscured. Superficially the species 
resembles Atheloca subrufella; the ground color of fore- 
wing a gray brown, darkening in outer area and shading 
to ocherous brown along inner margin at base; the 
costal margin broadly margined with white rather 
heavily dusted with red scales; antemedial line not dis- 
tinguishable; subterminal line faint, oblique and close 
to outer margin; discal dots very faint, separated. 
Hind wing translucent white shaded with fuscous at 
apex; the veins slightly darkened; on underside of male 
a coarse yellow sex-scaling between costa and cell, along 
lower margin of cell, extending for a short distance along 



veins 2 and 3, and along vein lb from base for nearly 
half its length. Alar expanse, 12-14 mm. 

Male genitaUa with terminal margin of vinculum 
evenly rounded; aedeagus shghtly bulged at middle, a 
row of short teeth along lateral edge towards apex. 
Female genitalia simple and membranous except for a 
fine sprinkling of minute scobinations and the two 
small signa. 

Type locality: Porto BeUo, Panamd (type in 
USNM). 

Food plant: Unknown. 

Distribution: Panama: La Chorrera (May), Porto 
Bello (May). 

Known only from Dyar's original type series. 

42. Peadus, new genus 

Type of genus: Piesmopoda burdettella Schaus. 

Tongue well developed. Male antenna with a shal- 
low sinus in shaft at base (as in Praedonula except here 
the hollow of the sinus is overlaid with rather coarse, 
appressed scales and without any indication of a tuft 
or lateral scale ridge), pubescent. Labial palpus slen- 
der, upturned, reaching vertex; third segment shorter 
than second, acuminate. Maxillary palpus subsqua- 
mous (as in Praedonula). Forewing smooth; venation 
as in Praedonula except vein 2 further from lower outer 
angle of ceU, and 6 from below upper angle. Hind wing 
with cell one-fourth the length of wing; discoceUular 
slightly but evenly curved. Eighth abdominal seg- 
ment with sclerotized pocket of sternite long and 
needlelike and with a large, flattened, fanlike tuft of 
long slender scales. 

Male genitalia with imcus and tegumen greatly re- 
duced; the uncus a narrow, weakly sclerotized angulate 
band. Gnathos indistinguishable (burdetellus) or rep- 
resented only by a very weakly sclerotized, transverse 
h&nd (dissitus) . Transtilla absent. Harpe short, stout ; 
sacculus broad and broadly produced at apex; giving 
the harpe a partially divided appearance; the free costal 
half of harpe strongly recurved and bearing two very 
stout spines, one on outer lower margin near the angle 
produced by the projection of sacculus and another at 
lower angle of cucuUus; the cucullus itself narrow and 
greatly reduced; a strong hair tuft from intersegment 
adjacent to base of sacculus. AneUus a shaUow, curved 
plate with stubby lateral arms. Aedeagus spined at 
apex; penis armed with a couple of more or less curved 
and crinkled sclerotized bands and a cluster of short, 
stout spines. Vinculum long, stout and evenly taper- 
ing; considerably longer than greatest width. 

Female genitaha with signum developed as a short, 
slender thorn; area of bursa immediately surrounding 
ductus seminalis scobinate and weakly sclerotized; 
ductus bursae much shorter than bursa, broad and 
more or less sclerotized; behind genital opening a pair 
of narrow elongate plates extending backward and fus- 
ing into the ventraUy divided eighth-segment collar 
(except in subaquUellus) ; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 



84 



XnsriTED STATES NATIONAL MUSEUM BULLETIN 207 



The genus is quite distinct from any other on male 
genitalia. Its closest relatives seem to be Praedonula 
and Hyalospila. On many genitalic characters it re- 
sembles the following genus (Gabinius), especially in 
the development of uncus and harpe; but separates 
from it on hind wing venation, especially the length 
and position of vein 2. 

174. Peadus burdettellus (Schaus), new combination 
Figures 269, 754 

Piesmopoda burdettella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 

11, p. 247, 1913. 
Discopalpia semproniella Schaus, Ann. Mag. Nat. Hist., ser. 8, 

vol. 11, p. 249, 1913. 
Hyalospila burdettella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7, 

p. 48, 1919. 

Forewing pale brown shaded with red and blackish 
scales; the costal border white sparsely dusted with 
red scales and with medial costal edge reddish; ante- 
medial line obscure except on lower half, far out, oblique 
from costa to cell and below ceU inwardly concave, 
white, bordered outwardly on costa by a faint reddish 
streak; some black scaling along basal half of vein lb; 
a distinct black dot on lower vein of cell at middle, and 
on the vein lb on the outer edge of the antemedial line; 
subterminal line sinuous, bordered inwardly by a nar- 
row, dark brownish shade and followed in outer area 
(especially towards apex) by some white dusting; discal 
dots distinct, separated, black, the lower one somewhat 
elongated ; an irregular black line along terminal mar- 
gin reaching almost to apex. Hind wings translucent, 
smoky white, darkening outwardly; the veins slightly 
darkened and a fine dark line along termen. Alar ex- 
panse, 19.5-20 mm. 

Male genitalia with no trace of sclerotized gnathos; 
aedeagus with apex bluntly pointed, bearing a line of 
short, coarse spines along its edge. Female genitalia 
with ductus bursae weakly sclerotized near genital open- 
ing; sclerotized plates behind genital opening narrow, 
bladelike; sclerotization of bursa near ductus seminalis 
slight. 

Type localities: Mount Pods (Juan Vinas), Costa 
Rica (burdettella, in USNM) ; Juan Vinas (semproniella, 
in USNM). 

Food plant: Unknown. 

Distribution: Costa Eica: Juan Vinas (Jan.), 
Mount Pods (May). Guatemala: Volcdn Santa Maria 
(July). 

Dyar estabUshed the above synonynay of Schaus' 
species. The genitalia of their male types are identical. 

175. Peadus dissitus, new species 
Figures 270, 755 

Sinular to burdettellus in color and markings except 
for a distinct whitish longitudinal shade through the 
cell of forewing and a stronger accentuation of the black 
scaling; a thin black streak from base along half the 
lower fold and a similar, shorter black streak on it just 
before the subterminal line; outer margin of antemedian 
line indicated by strong black dots on upper and lower 



veins of cell and on vein lb; inner dark margin of sub- 
terminal line broadened by black streak. Alar expanse, 
20-22 mm. 

Male genitalia with gnathos a weakly sclerotized 
transverse band; aedeagus with an expanded, flangelike, 
densely and finely spined apex. Other minor diflfer- 
ences from burdettella (especially in the shapes of 
cucvllus and the projecting part of sacculus of the harpe) 
are shown in the figures. Female genitalia with ductus 
bursae much shorter than in burdettella, strongly sclero- 
tized; sclerotized plates behind genital opening broad- 
ened at their bases; sclerotization of bursa near 
ductus seminalis appreciably stronger. 

Type locality: "S. E. Brazil" [Parand?] (type in 
BM; paratypes in BM and USNM, 61334). 

Food plant: Unknown. 

Described from male type and three male and three 
female paratypes from the type locality, "E. D. Jones, 
1920—303". 

176. Peadus subaquilellus (Ragonot), new combination 
Figure 753 

Hyalospila subaquilella Ragonot, Nouv. Gen., p. 11, 1888; Mono- 
graph pt. 1, p. 170, 1893; pt. 2, p. 38, fig. 24, 1901. 

This species is known only from the female type. A 
drawing of its genitalia made by Clarke is figured. I 
have seen nothing to match Eagonot's figure and de- 
scription which indicate a form with dark reddish brown 
forewing shaded somewhat with black at base and on 
the costa but without other markings; hind wing 
"transparent," smoky, the veins and terminal margin 
appreciably darkened. Alar expanse, 20 mm. 

The generic placement is tentative, pending discovery 
of a male. The female genitaHa are not typical, lacking 
the sclerotized plates behind genital opening, but seem 
to indicate a closer relationship to Peadus than to any 
other genus. 

Type locality: "Cerro Zunil," Guatemala (type in 
BM). 

Food plant: Unknown. 



Genus 43 : Gabinius 

[Venational division B. Forewing with veins 4 and 5 connate; 
2 from near lower outer angle of cell. Hind wing with 2 from 
close to lower outer angle of cell; 3 from the stalk of 4r-5; cell less 
than one-third the length of wing. Uncus reduced, weakly 
sclerotized, triangulate. Tegumen greatly reduced but with 
strong, projecting lateral arms. TranstUla incomplete. Harpe 
reduced; apex of costa spined. Vinculum short, stout.] 

43. Gabinius, new genus 

Type of genus: Promylea paulsoni Eagonot. 

Tongue well developed. Antenna of male simple, 
pubescent. Labial palpus upturned, reaching above 
vertex; second segment somewhat broadly scaled; third 
segment shorter than second, bluntly acuminate. Max- 
illary palpus squamous. Forewing smooth; 11 veins; 
vein 2 from before but near lower outer angle of cell, 
nearly as close to 3 at base as 3 is to 4; 3 from the angle; 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



4 and 5 connate; 6 from below upper angle of cell, 
straight; 8 and 9 long stalked, the free element of 9 very 
short; 10 from the cell, approximate to the stalk of 8-9 
for a short distance; male without costal fold. Hind 
wing with vein 2 from close to lower angle of cell; 3 
from the stalk of 4-5; 4 and 5 very long stalked; 7 and 8 
closely approximate beyond cell; cell less than one-third 
the length of wing; discocellular vein curved. Eighth 
abdominal segment with sternite developed as a short 
(stubby) sclerotized pocket with thin, curved lateral 
arms extending into a slender U-shaped plate, fringed 
with moderately long scales. 

Male genitalia with uncus reduced, broader than long, 
triangulate and weakly sclerotized. Tegumen support- 
ing a pair of long, broad, pointed, strongly sclerotized, 
backwardly projecting arms; otherwise greatly reduced. 
Gnathos absent. Transtilla incomplete; its elements 
pointed towards each other, their apices expanded and 
nearly touching. Harpe short, nearly as broad as long; 
costa broadly sclerotized, but shorter than remainder of 
harpe, bearing a stout, rather long, projecting spine at 
apex; cucuUus much reduced, bearing a similar, stout 
but shorter spine on costal edge at apex; sacculus 
broadly sclerotized, but not produced. Anellus a broad 
crescentiform plate ; its lateral lobes reduced to weakly 
haired knobs. Aedeagus smooth, straight; penis armed 
with a pair of ciu-ved, more or less wrinkled, sclerotized 
plates. Vinculum stout, as broad as long; its terminal 
margin evenly rounded. 

Female genitalia with signum developed as a thin, 
keellike blade on a narrow, elongate plate ; ductus bursae 
shorter than bursa and with a sclerotized collar near 
simple genital opening; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

A distinct genus, apparently most closely related to 
Peadus but falling into a different venational division 
(B). The shape and structure of the uncus, the re- 
duced tegumen, and the short harpe with its stout 
projecting spines suggest the relationship to Peadus. 
The very short cell of hind wing separates it from most 
genera of division B. It is only remotely related to 
Promylea, to which Ragonot referred its type species. 

177. Gabinius paulsoni (Ragonot), new combination 
Figures 271, 756 

Promylea paulsoni Ragonot, Nouv. Gen., p. 12, 1888; Mono- 
graph, pt. 1, p. 208, 1893. 

Ground color of forewing olivaceous gray strongly 
tinted with vinous brown in the dorsal area, this latter 
shade extending obliquely almost to apex; costal area 
from base to subterminal line, including the cell and 
tapering to costa beyond it, white with a scattered 
peppering of red-brown scales; costal edge from ante- 
medial hne to above end of cell edged with blackish 
brown; antemedial hne distinct on lower half of wing, 
vertical with a shght inward concavity, followed out- 
wardly by a narrow blackish brown band and inwardly 
by a blotch of the same shade; the antemedial hne 
indistinct and oblique on upper area of wing; subtermi- 



8& 

nal hne, narrow, faint, sinuate, whitish, followed and 
preceded for a short distance from costa by blackish 
brown shadings; a scattering of white scaUng in the 
terminal area below apex; lower discal spot blackish 
brown, more or less distinct, the upper very small and 
faint; a blackish irregular hne along terminal margin, 
not reaching to apex. Hind wing translucent, yellowish 
white with a smoky tint towards apex and along ter- 
minal margin. Alar expanse, 20-23 mm. 

Male genitaHa with characters as given for the 
genus. 

Type locality: Quillota, Chile (type in Paris Mus.). 

Food plant: Unknown. 

The female in the National Collection matches in 
every detail Ragonot's description and figure of the 
male type (Monograph, pi. 10, fig. 6). It is labeled 
"Chile, Silva." Superficially, the maculation and color 
resemble those of the Honora species. 

Genera 44-46: Ceracanthia to Drescoma 

[Venational division B. Forewing with veins 4 and 5 slightly 
separated at cell. Hind wing with cell less than half the length 
of wing (sometimes very short) ; 4 and 5 strongly stalked. Eighth 
sternite of male developed as a digitate pocket, sometimes the 
latter flattened and distorted. Male genitalia with vinculum 
very long, sclerotized only along its margins, arched dorsally 
(like a bent hairpin), its terminal margin broad; harpe with 
tufts on a projecting arm from base of sacculus.] 

44. Genus Ceracanthia Ragonot 

Ceracanthia Ragonot, Monograph, pt. 1, p. 230, 1893. (Type of 

genus: Ceracanthia vepreculella Ragonot.) 
Procandiopa Dyar, Ins. Inso. Menstr., vol. 7, p. 50, 1919. (Type 

of genus: Procandiopa mamella Dyar. New synonymy.) 

Tongue weU developed. Antenna pubescent; basal 
segment elongate, cyhndrical (longer and more heavily 
scaled in male than female); shaft of male with an 
elongate, shallow sinus at base, from middle of sinus a 
short, sharp, sclerotized spine, a similar, shorter spine 
at apical end of sinus. Labial palpus uptiu-ned, reach- 
ing above vertex, cyhndrical; third segment nearly as 
long as second, acuminate. Maxillary palpus sub- 
squamous (scaling of second segment somewhat ex- 
panded); folded across base of tongue. Forewing 
smooth; 11 veins; vein 2 from near lower outer angle 
of cell; 3 from the angle, about equidistant from 2 and 
4 ; 4 and 5 slightly separated at base, thence divergent ; 
6 from slightly below upper angle of cell, straight; 10 
from the cell, approximate to the stalk of 8-9 for a very 
short distance from cell; male without costal fold 
Hind wing with vein 2 from before, but near lower outer 
angle of cell; 3 from the stalk of 4-5; 4 and 5 stalked for 
half their lengths; 7 and 8 closely approximate for a 
short distance from cell; cell slightly over one-third the 
length of wing; discocellular vein curved. Eighth ab- 
dominal segment with sternite developed as a strongly 
sclerotized, digitate pocket. 

Male genitalia with uncus hoodlike (not tapering). 
Apical process of gnathos a simple, elongate, slender 
hook. Transtilla absent. Harpe broad, stout, sacculus 



86 



TTNITED STATES NATIONAL MUSEUM BULLETIN 207 



partially divided towards apex and with a projecting 
arm from its base supporting a heavy hair tuft. Anellus 
a rather narrow band with short, stubby, lateral arms. 
Aedeagus somewhat curved and with a slightly more 
sclerotized bulge from outer third; penis with a few 
minute scobinations or some fine sclerotized wrinkles, 
otherwise unarmed. Vinculum elongate (considerably 
longer than greatest width), U-shaped, sclerotized only 
(and narrowly) along its margins, arched dorsaUy (like 
a bent hairpin) . 

Female genitalia with signum; ductus bursae and 
genital opening simple; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

This genus is uncomfortably close to the genus fol- 
lowing (Megarthria), from which it differs chiefly in 
the closer approximation of vein 2 of hind wing to the 
lower outer angle of cell, the decided stalking of 3 with 
4-5, and the slightly longer cell. The male antennal 
character on which Eagonot and Dyar erected then- 
genera is probably here (as in Megarthria) of specific 
rather than generic significance. 

178. Ceracanthia mamella (Dyar), new comhination 
Figures 13, 272, 278, 757 
Procandiopa mamella Dyar, Ins. Insc. Menstr., vol. 7, p. 51, 1919, 

Forewing yellowish white along costa; remainder of 
wing a purplish shade with a scattered peppering of 
blackish scales especially along base of median fold; 
antemedial line obscure, far out on wing and outwardly 
angled in cell, indicated chiefly by a discontinuous, nar- 
row, blackish outer bordering line; subterminal line 
somewhat more distinct, sinuate, yellowish white, fol- 
lowed on costa by a short blackish dash and bordered 
inwardly by a black line which expands into patch at 
middle; discal dots blackish; terminal dots confluent 
forming a blackish line along the outer margin. Hind 
wing pale smoky brown; the veins faintly darkened and 
a distinctly smoky shade along termen. Alar expanse, 
15-16 mm. 

Male genitalia with outer margin of harpe evenly 
rounded. Female genitalia with signmn a flat, some- 
what granulate plate, bearing a short thorn near its 
center. 

Ttpe locality: Kfo Trinidad, Panamd (type in 
USNM). 

Food plant: Unknown. 

Distribution: PanamA: Kio Trinidad (Mar., May). 
Guatemala: Cayuga (May). 

Represented only by the original type series, a male 
and female from the type locality and a male and female 
from Guatemala. 

The sinus in the male antenna (fig. 278) appears 
smoothly scaled to the naked eye but xmder magnifica- 
tion shows several minute, erect, papillalike scales, simi- 
lar to those on Megarthria peterseni but less conspicuous. 

179. Ceracanthia vepreculella Ragonot 
FiQUEE 273 
CercxarUhia vepreculella Ragonot, Monograph, pt. 1, p. 230, 1893. 
I have seen no examples of this species but from 



Ragonot's description and figure it appears to be similar 
to mamella except that the general color is a more uni- 
form yellowish gray without the contrastingly paler 
costa and the purplish shade on the remainder of the 
wing, characteristic of mamella. It is also a larger 
species. Alar expanse, 23 mm. 

Male genitalia with outer margin of harpe angulate. 

Type locality: Loja, Ecuador (type in Paris Mus.). 

Food plant: Unknown. 

45. Genus Megarthria Ragonot 

Megarthria Ragonot, Monograph, pt. 1, p. 156, 1893. (Type of 
genus: Myelois peterseni Zeller.) 

Characters similar to those of Ceracanthia except: 
Vein 2 of hind wing fiu:ther removed from lower outer 
angle of cell; the cell itself shorter, less than one-third 
the length of wing; vein 3 connate with the stalk of 4-5 
or contiguous with it for a short distance from the angle 
of cell, but not from the stalk; eighth abdominal seg- 
ment of male with sternite developed usually as a 
laterally flattened, more or less bent, digitate pocket, 
the supporting lateral arms of the sternite bearing a 
pair of scale tufts. 

Male genitalia with anellus an elongate shield; trans- 
tilla absent (except in schausi) . 

Female genitalia with or without signum; ductus 
seminalis from bursa adjacent to junction of bursa and 
ductus bursae. 

The shaft of the male antenna has a rather broad 
sinus at base (as in Ceracanthia) but is specifically vari- 
able. The lower outer angle of cell in the hind wing is 
as far out as in Ceracanthia, but the discocellular vein 
curves inward more deeply, making the cell itself appre- 
ciably shorter than that of Ceracanthia. 

In the National Collection all the specimens except 
the male type of cervicalis Dyar had been identified as 
peterseni Zeller. They are strikingly similar in color 
and maculation, but their structures show that foiur 
species are present among the males and at least two 
among the females. With our present knowledge the 
sexes cannot be associated with any certainty; so untU 
something is known of their host association and the 
species are reared it seems a safer procedure to anticipate 
later synonjony and give separate names to the males 
and females rather than to link the females nomencla- 
torUy to any of the male types. 

180. Megarthria peterseni (Zeller) 
FiGUKES 14, 274, 279 

Myelois peterseni Zeller, Horae' Soc. Ent. Rossi cae, vol. 16, p. 

198, 1881. , 

Megarthria peterseni (Zeller) Ragonot, Monograph, pt. 1, p. 157, 

1893. 

Ground color of forewing white on costal half of wing, 
olivaceous brown on lower half, this brown shade 
extendiog upward to costa at extreme base and obliquely 
upward from lower outer angle of cell to costa at apex; 
an obhque blackish brown band from costa at about 
one-third extends to the paler brown ground color on 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



87' 



lower half of wing, dividing the white costal area into 
two strongly contrasted white patches; dark basal area 
also blackish towards costa; a similar blackish shade 
from apex extends obUquely inward towards lower outer 
angle of ceU; some blackish shading along the outer 
veins and a narrowly elongate, blackish brown spot on 
midcosta; sub terminal line very faint, except at costa, 
dull white, some faint whitish dusting in the outer 
brown area just below apex; a faint peppering of red 
scales on the white areas and more or less over the 
blackish brown markings on costal half of wing; discal 
dots blackish brown, separated and usually distinct; a 
row of black dots along termen. Hind wing semitrans- 
parent, whitish with a smoky tint towards apex and 
along costa and termen; the veins faintly darkened and 
a blackish line along terminal margin. Eighth abdomi- 
nal tufts more or less swollen hairhke scales. Alar 
expanse, 22-26 mm. 

Male genitalia with transtilla absent; harpe with 
sacculus partially divided (towards its apex) ; hair tufts 
from extended basal arm from sacculus, yellow; penis 
bearing a patch of fine scobiaations; vinculum con- 
stricted near middle. 

Antenna of male (fig. 279) with a broad sinus in base 
of shaft occupying a half-dozen fused segments; the 
sinus with many minute, papUlaUke setae (or scales) on 
its inner surface, but without hair or scale tuft; a small 
but strongly sclerotized spine from lateral edge of sinus 
beyond its base and a similar small spine from apex 
of the sinus. 

Type locality: Honda, Colombia (type, cf , inBM). 

Food plant: Unknown. 

Distribution: Guatemala: Volcdn Santa Maria 
(June, July). Colombia: Honda (Apr.). Brazil: 
Santa Catarina (July). PbrIj: Oconeque (Carabaya). 

I have seen no specimens from Colombia; but the 
male examples before me from Guatemala, BrazU, and 
Peni agree in antennal characters with the type of 
peterseni as described by Ragonot. Evidently the 
species has a wide distribution in Central and South 
America. 

181. Megarthria squamifera, new species 
Figures 275, 280 

Color and markings as in peterseni. Male genitalia 
also similar except lateral arms of gnathos stouter and 
vinculum less constricted. Digitate pocket of eighth 
abdominal sternite not appreciably flattened. Male 
antenna (fig. 280) with a broad, elongate sinus; from 
one lateral edge of sinus a flat brush of long stiff hairs; 
the opposing edge concave for most of its length, the 
concavity ending in a sharp projecting point at each 
end; inner surface of sinus smooth. 

Type locality: Mount Pods, Costa Rica (type in 
USNM, 61336). 

Food plant: Unknown. 

Described from male type, collected by Schaus and 
Barnes (May). 



182. Megarthria frustrator, new species 

Color markings and male genitaUa similar to those of 
peterseni. Male antenna like that of squamijera. Dif- 
fers from other males of the genus in having a narrow 
ridge of rough, protruding scales along the costa of 
forewing for more than half its length from base. Alar 
expanse, 20 mm. 

Type locality: Juan Vinas, Costa Rica (type in 
USNM, 61337). 

Food plant: Unknown. 

Described from male type, collected by Schaus and 
Barnes (Feb.). 

183. Megarthria schausi, new species 
Figure 276 

Color and markings as in peterseni. Male genitalia 
with vinculum not constricted; apical process of gnathos 
terminating in a weak, short spine. Transtilla present 
developed as a square, sclerotized plate, pendant from 
protruding lobes from the costobasal area of the harpes 
and with thin short projecting arms from its lower 
(anterior) corners. Hair tufts from projecting basal 
arm of sacculus short, yellow. Male antenna like that 
of squamifera. Alar expanse, 18 mm. 

Type locality: Juan Vinas, Costa Rica (type in 
USNM, 61338). 

Food plant: Unknown. 

Described from male type, collected by Schaus (Jan.) 
and named in memory of him. The species is easUy 
identified by its platelike transtilla. I have seen nothing 
resembling this structure in any other male of the genus. 

184. Megarthria cervicalis Dyar 

Figures 277, 281 

Megarthria cervicalis Dyar, Ins. Insc. Menstr., vol. 7, p. 42, 1919. 

Forewing as on peterseni except that dark areas are 
paler; less blackish brown and with more red scaling 
on the dark antemedial band and midcostal spot, the 
latter almost entirely reddish. Eighth abdominal tuft 
small, consisting of broadly flattened and contorted 
scales; digitate pocket from sternite of eighth segment 
strongly bent and decidedly flattened. Alar expanse, 
20 mm. 

Male genitalia with vinculum not constricted, of even 
width throughout and with terminal margin evenly 
rounded; penis armed with a small, flat, bladelike cor- 
nutus; sacculus of harpe not divided at apex; hair tuft 
from projecting basal arm of sacculus, black. 

Male antenna (fig. 281) with a short, shallow sinus 
at base of shaft and with a very small scale tuft from 
the base of the sinus (under the lower magnification 
looks like a small triangulate spine). 

Type locality: Tdnamo, Cuba (Aug.; type in 
USNM). 

Food plant: Unknown. 

Ejiown only from its male type. A distinct species 
easily identified by its male antennae and genitalia. 



■UNITED STATES NATIONAL MUSEUM BULLETIN 207 



185. Megarthria alpha, new species 

FiGUBE 760 

Female. Color and markings as in peterseni males. 
Alar expanse, 17-26 mm. 

Genitalia with signum developed as a teardrop 
shaped, finely granulate-scobinate, depressed patch: 
ductus bursae shorter than bursa with a narrow scle- 
rotized band near genital opening. Eighth-segment 
collar with a thin apron projecting anteriorly (the 
shape and size of this individually variable; identical 
in no two specimens; compare figures 760a and 760b). 

Type locality: Volcdn Santa Marfa, Guatemala 
(type in USNM, 61339). 

Food plant: Unltnown. 

Described from type (Sept.) and three para types 
from the type locaUty (Schaus and Barnes, collectors, 
July, Oct.) and one paratype from each of the following 
localities: Purulhd, Guatemala (Schaus and Barnes, 
July); Quirigud, Guatemala (Schaus and Barnes, Apr.), 
Jalapa, M6xico (Schaus); Orizaba, M6xico (R. MuHer, 
Mar. 13); Mount Pods, Costa Rica (Schaus, May); 
Juan Vinas, Costa Rica (Schaus and Barnes, Jan.); Rio 
Trinidad, Panama (A. Busck, Mar. 1912); Incachaca, 
Cochabamba, BoHvia (J. Steinbach); Santa Catarina, 
Brazil (F. Hoffmann, July 12, 1935). All females; 
genitalic preparations made of aU specimens. 

I beheve that alpha wiU eventually prove to be the 
female of peterseni but have no convincing evidence at 
this time that it is so. 

186. Megarthria beta, new species 

FiGiJHB 759 

Superficially indistinguishable from alpha but with 
quite different genitaha. Alar expanse, 17-23 mm. 

Bursa without trace of signum; ductus bm«ae much 
longer than bursa, unsclerotized at genital opening ex- 
cept for a narrow, very weakly sclerotized band on 
lower margin; anterodorsal projection of eighth-segment 
coUar shght. 

Type locality: Orizaba, Mexico (type in USNM, 
61340). 

Food plant: Unknown. 

Described from type (Schaus Collector, no data) ; one 
paratype from Jalapa, Mexico (Schaus, no data) ; three 
paratypes from Cayuga, Guatemala (Schaus and 
Barnes, Feb., May, Oct.); one paratype from San Jos6, 
Costa Rica (H. Schmidt); two paratypes from Porto 
Bello, Panama (A. Busck, May 1912); and one para- 
type from Caparo, west-central Trinidad (F. Birch, no 
data). All females; the paratypes from Trinidad and 
Costa Rica in the Ja,nse Collection. 

46. Genus Drescoma Dyar 

Drescoma Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 328, 1914. 
(Type of genus: Drescoma cyrdipsa Dyar.) 

Tongue well developed. Antenna pubescent; basal 
segment normal (not swollen or elongated); shaft of 
male simple. Labial palpus upturned, reaching vertex; 



moderately slender, the second segment somewhat 
broadly scaled ; third segment nearly as long as second, 
acuminate. Maxillary palpus subsquamous (scaling 
of second segment somewhat expanded); folded across 
base of tongue. Forewing smooth; 11 veins; vein 2 
from near lower outer angle of cell; 3 from the angle 
about equidistant from 2 and 4; 4 and 5 slightly sepa- 
rated at base, thence divergent; 6 from below (but near) 
upper angle of ceU, straight; 8 and 9 long stalked; 10 
from the cell approximate to the stalk of 8-9 for some 
distance; male without costal fold but with a distinct 
notch in costa beyond base. Hind wing with vein 2 
from lower outer angle of cell; 3 apparently from the 
stalk of 4-5, but actually contiguous or weakly anas- 
tomosed with it for half its length; 4 and 5 stalked for 
over half their lengths; 7-8 contiguous or partially 
anastomosed beyond cell, the free parts of the veins 
very long; cell very short, one-fifth the length of wing 
or less; discoceUular vein oblique; on male upper vein 
of ceU notched just beyond base and with some modi- 
fied sex-scaling above and beyond the notch; also on 
male a short fold on anal margin enclosing a hair pencil. 
Eighth abdominal segment without tuft; sternite de- 
veloped as a short sclerotized, digitate pocket. 

Male genitalia as in Megarthria except: Transtilla 
present and in the form of a wide, very shallow U; 
harpe with an appressed, clasperlike projection near 
apex of sacculus; vinculum terminating in a strongly 
sclerotized, sinuate, transverse bar. 

Female genitalia with signum developed as a long, 
strong, curved hook; ductus bursae shorter than bursa; 
genital opening simple; ductus seminalis from bm-sa 
near junction of bursa and ductus bursae. 

A distinct genus closely related to Ceracanthia and 
Megarthria and to Drescomopsis in group II; but easily 
distinguished by the bent (notched) upper vein of cell 
in the male hind wing, the very short hind wing cell 
in both sexes, the notched forewing of the male, the 
peculiarly developed transtilla and terminal margin of 
vinculum. 

187. Drescoma cyrdipsa Dyar 
Figures 15, 282, 758 
Drescoma cyrdipsa Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 328, 
1914. 

Color and markings as in Megarthria peterseni except 
for darker hind wings, especially those of the females 
which are dark smoky fuscous. The average size is 
also consistently smaller. 

Hind wing of male with notched projection of upper 
vein of cell into the cell deep and wide; on underside of 
wing a patch of semimetallic scales above the notch; 
a black patch preceding it, and following it a line of 
yeUow and black scales along the vein. 

Male genitalia with transtilla triangularly broadened 
toward harpes; clasperlike projection from sacculus 
rather broadly triangulate. Female genitalia with 
bursa minutely granulate; coarser granulations in ductus 
bursae at its junction with bursa. The long, strong, 
thornlike cornutus may be a specific character also, 
but is more probably generic. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



89 



Type locality: La Chorrera, Panama (type in 
USNM). 

Food plant: Unknown. 

Distribution: Mexico: Chiapas (May). Guate- 
mala: Cayuga (Jan., May, June, Aug.). Panama: 
Cabima (May), Corozal (Feb., Nov.), La Chorrera 
(Apr., May), Eio Trinidad (Mar., June), Tabernilla. 
French Guiana: St. Jean Maroni. 

188. Drescoma cinilixa Dyar 

FiGUEB 283 

Drescoma cinilixa Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 329, 
1914. 

Markings and color of forewing as in cyrdipsa except 
dark ground color and dark costal markings paler, 
more ocherous fuscous than brown. Underside of male 
hind wing with a patch of black sex-scaling above the 
notch in cell, orange-yellow sex-scaling on the upper 
vein of cell preceding the notch, on several veins follow- 
ing the notch, and in the median fold at base of wing; 
the indentation of upper vein of cell also much shallower 
than on cyrdipsa, a concavity rather than a strongly 
triangular notch. Hind wing paler; dull smoky white, 
darkening towards termen and apex. Alar expanse, 
15-16 mm. 

Male genitalia with transtilla narrowed towards 
harpes; clasperUke projection from sacculus slender, 
sharply pointed. Female unknown. 

Type locality: La Chorrera, Panamd (type in 
USNM). 

Food plant: Unknown. 

Distribution: Guatemala: Cayuga (May). Pa- 
namA: La Chorrera (May). 

Distinguishable from cyrdipsa by its paler color, the 
secondary sexual characters of the male hind wing, and 
the differently shaped transtilla and clasper. 

Genus 47. Monoptilota 

[Venational division B. Forewing with veins 4 and 5 closely ap- 
proximate for a short distance from cell. Hind wing with vein 2 
from well before angle of cell ; 3 from the angle ; cell one-third the 
length of wing. Male genitalia with uncus bifid, its divided ele- 
ments widely separated; transtilla incomplete; apical process of 
gnathos a broadly triangulate hook. Male antenna unipectinate 
with sinus and scale tuft in base of shaft.] 

47. Genus Monoptilota Hulst 

Monoptilota Hulst, Canadian Ent., vol. 32, p. 13, 1900. — Forbes, 
Cornell Mem. 68, p. 621, 1923. (Type of genus: Monoptilota 
nubilella Hulst.) 

Tongue well developed. Antenna of male (figs. 
285g-h) with basal segment elongate, shaft unipectinate 
and with a long shallow sinus at base containing a stout, 
appressed scale tuft; of female simple, smooth. Labial 
palpus obliquely upturned, reaching to vertex; second 
segment broadly scaled; third segment small, acumi- 
nate. Maxillary palpus squamous. Forewing smooth ; 
11 veins; vein 2 from well before lower outer angle of 
cell; 3 from the angle, much closer at base to 4 than to 

30032&— 56 7 



2; 4-5 closely approximate for a short distance from 
cell; 6 from below upper angle of ceU, straight; 8 and 9 
stalked for less than half their lengths; 10 from the cell; 
male without costal fold. Hind wing with vein 2 from 
well before lower outer angle of cell; 3 from the angle, 
connate with or very closely approximate at base to 
stalk of 4-5; 4 and 5 stalked for about half their lengths; 
7 and 8 closely approximate for a short distance from 
cell; cell one-third the length of wing; discoceUular vein 
curved. Eighth abdominal segment of male with a 
pair of fine, weak, hair tufts. 

Male genitaUa with uncus bifid, the divided parts 
widely separated and hooked at their apices. Gnathos 
terminating in a triangulate, sharply hooked central 
process, its supporting lateral arms strongly arched and 
arising well down from ventrolateral projections of 
tegumen; an elaborate well-sclerotized subanal plate 
attached to alimentary tube. Transtilla incomplete, 
consisting of two, widely spaced, slender, weakly sclero- 
tized plates. Anellus a simple, moderately broad, par- 
tially cm-ved band. Aedeagus short, stout; penis un- 
armed. Vinculum short (shorter than broad) ; terminal 
margin truncate. 

Female genitalia with signum developed as a spined 
plate (individually variable); bursa otherwise smooth; 
ductus bursae rather stout, expanding gradually into 
the bursa, smooth except for a broad sclerotized banding 
near genital opening; ductus seminaHs from bursa near 
junction of bursa and ductus bursae. 

A unique genus of doubtful affinities. Contains but 
the one North American species. 

189. Monoptilota pergratialig (Hulst) 
Figures 27, 285, 763, 764 

Nephopteryx pergratialis Hulst, Trans. Amer. Ent. Soc, vol. 13, 
p. 162, 1886; Phycitidae of N. Amer., p. 143, 1890. — Rago- 
not. Monograph, pt. 1, p. 267, 1893. 

Nephopteryx grotella Ragonot, N. Amer. Phycitidae, p. 6, 1887. 

Monoptilota nubilella Hulst, Canadian Ent., vol. 32, p. 14, 1900. — 
Chittenden, U. S. Dep. Agr. Div. Ent. Bull. 23, pp. 9-17, 
1900.— Welden, Jour. Econ. Ent., vol. 1, p. 148, 1908.— 
Forbes, Cornell Mem. 68, p. 621, 1923. 

Monoptilota pergratialis (Hulst) Barnes and McDunnough, Con- 
tributions, vol. 3, p. 195, 1916. — Brannon, Journ. Econ. 
Ent., vol. 27, p. 719, 1934.— Brimley, Insects of North Caro- 
lina, p. 300, 1938.— McDunnough, Check list. No. 6141, 
1939. 

Forewing dark fuscous (gray-brown) with blackish 
shading on many of the veins and on costal half of wing; 
along terminal margin a heavy dusting of white between 
the veins, giving much of wing an ashy appearance; 
antemedial line obscure, indicated chiefly by a diffused 
blackish brown outer border, broadest and strongest 
from costa to lower margin of cell, frequently inter- 
rupted or obsciu-ed in the ground color towards inner 
margin; sub terminal line sinuate, grayish white with 
dentate blackish brown inner and outer borders ; discal 
dots distinct, separated, blackish; a black line along 
terminal margin more or less interrupted by white 
streaklets at the vein ends. Hind wing of male semi- 
hyaline white with veins slightly darkened and a black- 
ish brown line along terminal margin; of female much 



90 



UNITED STATES NATIONAL MUSEUM BULLETIN 2 07 



darker, dull smoky white to brown, with veins and ter- 
minal edge correspondingly darker. Alar expanse, 
17-30 mm. 

Male genitalia with characters as given for the genus. 
Female genitaha with signum extremely variable, rang- 
ing from a small, weakly spiaed plate, like that of the 
para type (fig. 764), to a large plate with rather long 
slender spines (fig. 763). In one large female of a reared 
series from Virginia the signum is completely absent. 
This variability in female structure is not matched by 
anything in the male, where the genitalia are remark- 
ably imiform for large and small specimens ahke. 

Type locality: Florida (pergratialis, 9, in AMNH, 
ex Eutgers; grotella, 9, in Paris Mus.); Auburn, Ala. 
(nubilella, cf , in USNM). 

Food plant: Limabean (larva a borer in the stems). 

Distribution: Maryland, Cabin John Bridge (Aug., 
Sept.), Salisbury (Sept.); Virginia, Norfolk (May), St. 
Elmo (Jan., Feb., Mar.) ; North Carolina, Vance Coimty 
(Aug., Oct.); South Carolina, Florence (June, July), 
Lyna Plantation (June); Georgia, Savannah (June); 
Florida, Coconut Grove (Apr., May), Miami (Apr.); 
Alabama, Auburn (July), Montgomery (July); Arizona, 
Baboquivari Mts. (June, Aug., Sept.), Huachuca Mts. 
(July, Aug.), Nogales (July), Palmerlee, Washington 
Mts., White Mts. (June). 

The species is of some importance as an enemy of 
limabeans and is known in economic literature as the 
"limabeam vine borer." The Chittenden (1900) paper 
cited gives what is known of the life history. There are 
several later references in publications devoted to eco- 
nomic entomology but they add nothing to our knowl- 
edge of the insect. It is not known outside of the 
United States; at least no specimens have been received 
or identified from any of the tropical American regions 
where the species might be expected to occur; and in 
the United States its distribution seems to be limited 
to the eastern area from the District of Columbia south 
to Florida and adjacent GnLf States and to southern 
Arizona. The only known host is the limabean. The 
southern Arizona distribution raises a question as to 
another possible host (probably a wild legume), for the 
Arizona localities are mostly out of the range of lima- 
bean cultivation. 

Genera 48-50: Zamagiria to Magariopsis 

[Venational division D. Forewing with veins 4 and 5 connate, 
shortly stalked or closely approximate for a short distance from 
base; 3 close to 4-5 at base; 6 bent towards base connate with or 
shortly separated from the stalk of 8-9 at base. Hind wing with 
4 and 5 anastomosed from just beyond angle of cell for about 
half their lengths; cell short. Eighth abdominal segment of 
male with compound, ventral tufts. Labial palpi of male up- 
curved; appressed to face or to each other; third segment greatly 
reduced, acuminate. Gnathos with apical process broadly pro- 
duced and lateral arms elongate and arising from ventrolateral 
proiection from tegumen. Female with ductus bursae short and 
strongly sclerotized towards genital opening and junction with 
bursa.] 



48. Genus Zamagiria Dyar 

Zamagiria Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 329, 1914. 
(Type of genus: Zamagiria dixolophella Dyar.) 

Tongue well developed. Antenna of male shortly 
ciliate, a deep sinus containing heavy scale tuft at base 
of shaft ; of female simple and weakly pubescent. Male 
head between the antennae deeply grooved to hold an 
appressed, matted tuft of long scales arising from the 
upper edge of frons. Labial palpus recurved-ascending; 
second segment very long; broadly dilated and hol- 
lowed within to hold the maxillary palpus; third seg- 
ment short, acute. In repose the labial palpi are 
appressed to each other and fit into the groove on head 
covering both the maxillary palpi and the scale tuft 
from frons. Maxillary palpus of male in the form of a 
large aigrette; of the female minute, filiform. Forewing 
with a ridge of roughened scales preceding the ante- 
medial line but not reaching to costa; 11 veins; vein 2 
from before the lower outer angle of cell; 3 from the 
angle, close to 4-5 at base; 4 and 5 very closely approxi- 
mate for a short distance from ceU; 6 from upper angle 
of cell, bent towards base, connate with 8-9; 8 and 9 
long stalked; 10 from the cell, approximate to the stalk 
of 8-9 for a short distance from cell; male without costal 
fold but rough scaled on underside of costa at base. 
Hind wing with vein 2 from well before the angle of the 
cell; 3 from the angle; 4 and 5 anastomosed from just 
beyond the angle for about half their lengths; 7 and 8 
contiguous or weakly anastomosed for a short distance 
beyond ceU; cell less than one-fourth the length of wing; 
discoceUular vein curved. Eighth abdominal segment 
of male with compound ventral scale tufts. 

Male genitalia with uncus triangulate. Gnathos with 
apical process broadly produced and variously modified; 
lateral arms elongate and arising well down from ventro- 
lateral projections of tegumen. Transtilla absent. 
Harpe with sacculus more or less produced at apex; 
clasper in some form usually present; cucullus narrow 
and reduced. Aedeagus stout; penis armed with strong 
spine or spines (except in hospitabUis) . 

Female genitalia with bursa armed with strong spine 
cluster or clusters and usually partially sclerotized; 
ductus bursae short, strongly sclerotized towards genital 
opening and junction of biu'sa and ductus bursa (the 
sclerotizations more or less contorted) ; genital opening 
broad; ductus seminalis from bursa towards junction 
of bursa and ductus bursae. 

An easily recognized, compact genus exhibiting strik- 
ing structural specific differences in genitalia. 

190. Zamagiria dixolophella Dyar 

Figure 286 

Zamagiria dixolophella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 
329, 1914. 

Aigrette of male maxillary palpus reddish. 
Forewing dark smoky gray, the basal area below costa 
a trifle paler; antemedial line faint, narrow, oblique and 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



91 



sinuate, well out on wing, bordered outwardly by a 
narrow blackish line, inwardly (from top of cell to inner 
margin by a broad salmon-ocherous patch filling the 
space between antemedian line and the vertical scale 
ridge; the latter thin, blackish, bordered inwardly by a 
white line; subterminal line near outer margin, indis- 
tinct, pale, bordered inwardly by some obscure blackish 
streaklets on the veins; discal dots separated; all the 
markings obscure except the white inner margin of the 
subbasal scale ridge. Hind wing translucent, pale 
smoky fuscous; the veins and terminal margin faintly 
darkened. Alar expanse, 21 mm. 

Male genitalia with flaring apical process of gnathos 
terminating in a shortly forked hook. Harpe with a 
broadly triangulate projection (clasper) from near apex 
of sacculus, the sacculus otherwise not appreciably 
produced at apex. Penis armed with three clusters of 
straight, strong spines. 

Type locality: Corozal, Canal Zone, Panamd (Nov.; 
typeinUSNM). 

Food plant: Unknown. 

Represented only by the male type. 

191. Zamagiria pogerythrus Dyar 

Figures 49, 287, 765 

Zamagiria pogerythrus Dyar, Ins. Insc. Menstr., vol. 7, p. 47, 
1919. 

Aigrette of male maxillary palpus red. 

Forewing pale brownish gray dusted and blotched 
with white, slightly darker along costa ; the white scaling 
most conspicuous about the lower discal spot, as a more 
or less diffused blotch in median area following the 
antemedial line, as an interrupted white line preceding 
the subbasal scale ridge, and as a short, narrow shade 
from apex in terminal area; antemedial pale line faint, 
its outer blackish border interrupted on the males, 
continuous on females; the subbasal scale ridge broken 
into two or three patches of black intermixed with some 
red raised scales; the patch between these and the ante- 
medial line more restricted and fainter than on dixo- 
lophella, ocherous fuscous; subterminal line well con- 
trasted, indented at vein 6, white, margined inwardly by 
blackish streakings on the verus and inwardly and out- 
wardly by dark spots on costa; discal dots separate, 
black, the lower one elongately enlarged and the most 
conspicuous black marking on the wing; a row of 4 or 
5 black dots along terminal margin. Hind wing whit- 
ish, translucent; a dark shade along costa and a narrow 
dark line along termen; the veins very faintly darkened. 
Alar expanse, 20-22 mm. 

Male genitalia with apical process of gnathos con- 
torted. Harpe with apex of sacculus shortly and 
bluntly produced at apex. Penis armed with two 
clusters of straight, stout, moderately long spines. 

Female genitalia with signiun developed as a narrow, 
elongate, strongly spined plate extending the length of 
the bursa; bursa otherwise unsclerotized and minutely 
spinose; ductus bursae very broad (as broad as long and 



broader than bm-sa), its membrane thickened and 
bearing on its inner dorsal surface, near genital opening, 
a large pair of conjoined sclerotized plates, and on its 
inner ventral surface, near junction of bursa and ductus, 
a large, thickened, corrugate, triangulate, sclerotized 
plate bearing minute spines over its inner surface. 
Eighth-segment collar ventrally fused. 

Type locality: Chejel, Guatemala (type d', in 
USNM). 

Food plant: Unknown. 

Distribution: M:^xico: Campeche (July). Guate- 
mala: Chejel (June, Aug.), Purulhd (July). 

192. Zamagiria hogpitabilis Dyar 

FiGUBB 288 

Zamagiria hospitabilis Dyar, Ins. Insc. Menstr., vol. 7, p. 48, 
1919. 

Aigrette of maxillary palpus pale red. 

Forewing ocherous brown heavily overlaid with black, 
giving the wing a dark, blackish brown groimd color; 
two strongly contrasted white markings, the inner 
border of the broken, black, subbasal scale ridge and a 
large, irregular blotch on lower median area just beyond 
the antemedial line; antemedial line not strongly con- 
trasted but distinguishable throughout, sinuate, whit- 
ish, bordered outwardly by a thin black line; subtermi- 
nal line more contrasted, dull white, indented at vein 6 
and at lower fold ; discal dots confluent, forming a black 
lunate mark on discocellular vein; outer area beyond 
subterminal line very faintly dusted with white; ter- 
minal dots confluent forming a narrow black Hne along 
outer margin of wing. Hind wing semihyaline white 
with a dark shade along costa; the veins and terminal 
margin faintly darkened. Alar expanse, 21 mm. 

Male genitalia with apical process of gnathos broadly 
flaring and serrate. Harpe with sacculus shortly pro- 
duced at apex, very long and with broadly triangulate, 
median, clasperlike projection (similar to that of 
dixolophella) . Lateral margins of aedeagus armed with 
short, stout spines; penis unarmed except for a small, 
flat, weakly sclerotized plate. 

Type locality: Tdnamo, Cuba (type in USNM). 

Food plant: Unknown. 

Represented only by the male type. 

193. Zamagiria masciilinus Dyar 

FiauBE 289 

Zamagiria mascuUnus Dyar, Ins. Insc. Menstr., vol. 7, p. 46, 
1919. 

Aigrette of male maxillary palpus whitish ocherous. 

Forewing very pale ocherous gray thinly dusted with 
white in median area; the ocherous tint strongest in the 
patch between scale ridge and antemedial line, along 
the costal edge, and broadly along lower fold; ante- 
medial and subterminal lines distinguishable, but not 
strongly accented, whitish; black scaling of subbasal 
scale ridge limited to one or two dots; blackish outer 



92 



UNITED STATES NATIONAL MUSEUM BULLETm 207 



border of antemedial line interrupted; inner blackish 
border of subterminal line broken into short black 
streaks on the veins; discal dots separated, blackish; a 
row of small blackish dots (5 or 6) along termen between 
the vein ends; on inner margin at lower outer edge of 
antemedial line a small white spot; none of the white 
markings strongly contrasted; the usual black markings 
broken into dots and very short dashes; overall shade a 
pale clay color. Hind wing translucent, white with a 
faint ocherous tint; a narrow pale brownish line along 
termen; the veins very faintly darkened. Alar expanse, 
25 mm. 

Male genitalia with gnathos terminating in a broad, 
inverted, heart-shaped apical projection. Harpe with 
sacculus short, sharply pointed and shortly projecting 
at apex. Penis armed with a single stout spine about 
one-third as long as aedeagus. 

Type locality: Cayuga, Guatemala (Apr.; type in 
USNM). 

Food plant: Unknown. 

The largest known species of Zamagiria. Eaiown 
only from the male type. 

194. Zamagiria australella (Hulst), new combination 

Figures 292, 766 

Selagia australella Hulst, Canadian Ent., vol. 32, p. 174, 1900. — 

McDunnough, Check list, No. 6232, 1939. 
Immyrla humeliella Barnes and McDunnough, Contributions, 

vol. 2, p. 182, 1913.— McDunnough, Check list. No. 6188, 

1939. (New synonymy.) 

Aigrette of male maxUlary palpus ocherous white. 

Forewing white dusted with black scales, giving the 
wing a pale gray color, lightest in basal and through the 
median areas, slightly darker along costa ; subbasal scale 
ridge conspicuous and normally unbroken, black with a 
fine white inner border; the patch between scale ridge 
and antemedial line olivaceous ocherous; antemedial 
line oblique, dentate-sinuate, faint, indicated chiefly by 
its black outer bordering line; subterminal line dentate- 
sinuate, bordered inwardly and outwardly at costa by 
blackish dashes, outwardly below costa by a narrow 
brownish shade and inwardly by a fine black line; discal 
dots distinct, separate, black; a small blackish or 
brownish spot on inner margin a slight distance beyond 
antemedial line and diffused black smudges on veins 2, 
3 and 4 for a short distance from cell. Hind wing 
semihyaline white; a darker line along termen and the 
veins very slightly darkened. Alar expanse, 15-18 mm. 

Male genitalia with apical process of gnathos a nearly 
square plate. Harpe with costa narrowly sclerotized 
and shortly produced at apex; sacculus shortly and 
bluntly produced. Penis armed with a single stout 
cornutus about half as long as aedeagus. 

Female genitalia with bm-sa sclerotized and densely 
spined at posterior end, the sclerotization extending 
into ductus bursae; occupying most of the remainder of 
ductus bursae a funnel-shaped, convoluted, sclerotized 
plate. 

Type localities: Blanco County, Tex. {australella, 



in AMNH, ex Kutgers) ; Fort Myers, Fla. {humeliella 
in USNM). 

Food plant: Bumelia microcarpa. 

Distribution: Texas, Blanco County; Florida, Fort 
Myers (May), Miami. 

The genitalia of the female type of australella are 
identical with those of a reared female paratype of 
humeliella, and the two moths otherwise agree, so there 
is no question of the synonymy. However, I doubt 
somewhat the correctness of the locality label on Hulst's 
type and suspect that it may be a Florida specimen. 

195. Zamagiria fratema, new species 

FlQTJEE 291 

Aigrette of male maxillary palpus white. 

Forewing white on dorsal half and along inner margin 
beyond antemedial line to tornal angle; extreme base of 
costa rough scaled, black (similar black sex-scaling on 
underside of wing at base) ; antemedial Une at middle of 
wing incomplete, indicated only by an angulate white 
line on lower half, bordered inwardly and outwardly by 
small black smudges; along lower margin to antemedial 
line the ground color is pale brown; a faint dusting of 
brown or purplish brown scales on the white groimd for 
a narrow margin along costa; scale ridge interrupted, a 
series of black dots with the inner white bordering line 
very faintly indicated; the usual patch between scale 
ridge and antemedial line pale brown, a trifle paler than 
the ground color on lower basal area of wing; some simi- 
larly colored scales in the lower postmedian area be- 
tween the antemedian line and the end of cell and in the 
interspaces of veins 3 to 5 ; subterminal line incomplete, 
only its middle portion distinguishable; discal dots dis- 
tinct, separated, black, the lower one enlarged; black 
scaling along veins 3, 4 and 5. Hind wing hyaline 
white, a pale brownish gray line along terminal margin 
and a similar shade along costa; at extreme base of wing 
a few black sex-scales on the veins. Alar expanse, 
21.5 mm. 

Male genitalia with apical process of gnathos roundly 
spatulate. Harpe considerably broadened towards 
middle, thence sharply tapered to the narrow cucidlus; 
costa strongly humped at middle; sacculus produced at 
apex into a long, strongly sclerotized, curved, free, 
hooklike arm. Anellus a U-shaped plate. Penis armed 
with a single stout spine (about one-fourth as long as 
aedeagus), a strongly sclerotized, corrugated plate and 
a concentration of rather coarse granulations. 

Type locality: Santiago de las Vegas, Cuba (type 
in USNM, 61341). 

Food plant: "Caimitillo." 

Described from male type reared by A. Otero, Aug. 
17, 1932, and labeled: "Leaf tier on Caimitillo, E. E. A. 
Cuba, Ento. No. 10006." 

196. Zamagiria laidion (Zeller) 
FiGUBES 290, 767 

Myelois laidion Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 211, 

1881. 
Piesmopoda laidion (Zeller) Ragonot, Monograph, pt. 1, p. 162, 

1893. 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



93 



Zamagiria laidion (Zeller) Dyar, Ins. Insc. Menstr., vol. 7, p. 46, 

1919. 
Zamagiria deia Dyar, Ins. Insc. Menstr., vol. 7, p. 46, 1919 (new 

synonymy). 
Zamagiria striella Dyar, Ins. Insc. Menstr., vol. 7, p. 47, 1919 

(new synonymy). 

Aigrette of male maxillary palpus dull white. 

Forewing pale gray, the overall tint shading from 
grayish white to very pale bluish gray (in fresh reared 
examples); costal border slightly darker; the patch be- 
tween scale ridge and antemedial Hne reddish brown; 
similar reddish brown scaling spread over basal area 
bordering inner margin and frequently blotching the 
median area over vein lb and the lower fold (especially 
on the males) ; also some scattering of reddish brown 
streaking on veins 3 and 4 for a very short distance from 
cell; black scale ridge more or less complete; on several 
males a black or black and red-brown streak along vein 
lb from scale ridge to base of wing; transverse lines 
faint, the outer blacldsh border of the antemedial and 
the inner dark border of subterminal more or less inter- 
rupted (more so on males than females); discal dots 
separated, black, the lower one shghtly enlarged; a 
row of 5 or 6 small black dots along termen. Hind 
wing semihyaline white; a dark line along terminal mar- 
gin and some faint darkening of the veins. Alar ex- 
panse, 15-22 mm. 

Male genitalia similar to those of Jraterna except: 
Costa of harpe convex but not decidedly humped; an- 
nellus inverted ; ^ penis with cornutus a single, very 
stout, long spine (over two-thirds as long as aedeagus) 
surrounded by a cluster of small granulations. 

Female genitalia with a finely spined plate (signum) 
near junction of bursa and ductus bursae, and a strongly 
sclerotized lateral patch in bursa near its anterior end; 
ductus bursae with a sclerotized plate occupying most 
of its length, the posterior end of the plate folded over 
into triangulate ventral lips. 

Type localities: Honda, Colombia {laidion, in BM) ; 
Chejel, Guatemala {deia and striella, in USNM). 

Food plants: Achras sapota, Mimusops emarginata, 
Eriobotyra japonica (larvae feeding on leaves and 
flowers) . 

Distribution: United States: Florida, Key West 
(Apr., May), Aliami (Jan., Dec). Guatemala: Chejel 
(June). PanamX: Porto Bello (Feb.). Colombia: 
Honda. Brazil: Castro {Parand) Obidos (Amazon re- 
gion, Sept.), Vi50sa {Minas Geraes, Sept.). Bolivia: 
"East Bolivia" ("Aug.-Oct., T. Steinbach"). 

Hitherto laidion has been recognized only from female 
examples. A reared series in the National Museum 
from Florida has enabled us to associate the sexes and 
has established the synonymy of deia and striella, both 
described from males. Dyar's type of striella is merely 
an extreme example of a common color variant with 

' This structure seems more like a transtilla than an anellus; 
for its straight posterior margin lies between the costal bases of 
the harpes, and it could be interpreted as a transtilla or com- 
bination of transtilla and anellus, except that in other species of 
the genus there is no trace of even the vestiges of a true transtilla. 



more or less black streaking on the base of vein lb of 
forewing. The reared Florida specimens have also 
given us the host records cited above. 

197. Zamagiria ipsetona Dyar 

Figure 768 

Zamagiria ipsetona Dyar, Ins. Insc. Menstr., vol. 7, p. 47, 1919. 

Forewing gray, more heavily marked with black than 
in preceding species; the blackish streakings on the 
veins (2 to 6 in this species) longer and stronger; lower 
discal spot elongated into a black dash; dark dashes 
(reddish brown) on the veins in outer area following the 
faint subterminal line; the black scale ridge not con- 
tinuous; antemedial line sinuate and nearly vertical; 
the whitish areas limited to a pale oval patch surround- 
ing the black-streaked veins and discal dots and a faint 
shade preceding the black scale ridge; the usual red- 
brown scaling limited to the patch between scale ridge 
and antemedial line and weak shadings between some 
of the veins in postmedian area. Hind wing translu- 
cent white; a narrow dark shade along costa and a 
blackish line along termen; veins appreciably darkened. 
Alar expanse, 23-24 mm. 

Female genitalia with two elongate spined plates in 
bursa; another more weakly spined plate at junction of 
bursa and ductus bursae; from the junction a scJeroti- 
zation extends along one side of bursa for about four- 
fifths of its length; ductus bursae very short, strongly 
sclerotized towards genital opening, the plate folded 
over at the opening into ventrolateral lips. Eighth- 
segment collar completely fused ventrally. 

Type locality: Juan Vinas, Costa Rica (type in 
USNM). 

Food plant : Unknown. 

Represented by three females from the type locality 
(Feb., June). The male is unknovra. 

49. Genus Anegcephalesis Dyar 

Anegcephalesis Dyar, Ins. Insc. Menstr., vol. 5, p. 46, 1917. — 
Heinrich, Ins. Insc. Menstr., vol. 5, p. 48, 1917 (larva). 
(Type of genus: Anegcephalesis cathaereles Dyar.) 

Characters of Zamagiria except: Maxillary palpus 
minute, subsquamous in both sexes. Forewing with 
veins 4 and 5 connate or very shortly stalked. Hind 
wing with cell longer (slightly less than one-third the 
length of wing) . 

The genus is very close to Zamagiria but its separa- 
tion seems to be justified by the differences in the male 
maxillary palpi and the longer cell of hind wing. The 
venational differences noted above (between veins 4 and 
4 of forewing) may be only a specific character. Dyar 
distinguished Anegcephalesis from Zamagiria on the 
difference in maxillary palpi, but mistook the appressed 
scale tuft from frons for that organ. This scale tuft is 
present and equally well developed in Zamagiria and 
Anegcephalesis. 



94 



UNITED STATES NATIONAL MUSEUM BULLETESr 207 



198. Anegcephalesis arctella (Ragonot), new combination 

FiGTTRBS 50, 294, 770 

Phycita arctella Ragonot, N. Amer. Phycitidae, p. 4, 1887; 
Monograph, pt. 1, p. 180, 1893. 

Anegcephalesis cathaeretes Dyar, Ins. Insc. Menstr., vol. 5, p. 46, 
1917 (new synonymy) . 

Anegcephalesis catheretes (Dyar), Ins. Insc. Menstr., vol. 7, p. 
48, 1919 (food plant and altered spelling). — McDunnough, 
Check list, No. 6176, 1939. — Bruner, Scaramuzza, and 
Otero, Bull. 63, Estaoi6n Exp. Agron., Cuba, p. 69, 1945. 

Forewing brownish gray dusted with white; the 
white shading more pronounced (on the males) in the 
median area above the base of the cell, as a small spot 
on inner margin below the dark markings on veins 2 
to 5, and in outer area beyond the dark outer border of 
the subterminal hne, on the female the white dusting 
is more generally distributed over median and outer 
areas and, on some specimens, on the basal area, giving 
the wing an ashy gray appearance ; subbasal ridge black 
bordered inwardly by a white line and followed by a 
dull ohvaceous ocherous patch; antemedial line well 
out towards middle of wing, obUque, notched at vein lb 
and evenly curved above it, outwardly bordered by a 
continuous black line; subterminal line distinct; in- 
dented just below costa and very slightly so at lb; 
discal spots fused into a lunate black line along dis- 
cocellular vein; a dark brownish smudge over veins 1 
to 5 adjacent to cell. Hind wings translucent white; 
the veins very faintly darkened. A narrow brown line 
along termen. Alar expanse, 20-23 mm. 

Male genitalia with uncus narrowly triangulate. 
Apical process of gnathos a thin, broadly ovate plate. 
Harpe with appressed, elongate clasper; sacculus not 
produced at apex. Aedeagus sharply bent; penis armed 
with an elongate, flattened, partially bent, platelike 
cornutus, and some sclerotized wrinklings and granu- 
lations. 

Female genitalia with signum developed as an elon- 
gate, strongly spined plate (about half as long as bursa) ; 
opposite the signum a narrower, much more weakly 
sclerotized plate beginning at junction of bursa and 
ductus bursae (this plate about half the length of 
signum); bursa otherwise smooth; ductus bursae tubu- 
lar, sclerotized. Eighth-segment collar fused ven- 
trally. 

Type locality: Nassau, Bahamas (arctella, in Paris 
Mus. ; cathaeretes, in USNM). 

Food plant: Dipholis salicijolia. 

Distribution: United States: Florida, Miami, 
Paradise Key (Mar.), Koyal Palm State Park (Feb.). 
Bahamas: Nassau (New Providence Isl., Feb., Mar., 
April, May, Sept.). Cuba: Baracoa (Oct.), Santiago 
Province (Feb., June, Sept., Oct., Dec). 

Ragonot's arctella was described from a single female. 
His description and figure agree in every detail with 
Dyar's cathaeretes represented by a large series in the 
U. S. National Museum. Both were described from 
the same type locality. There is no doubt that both 
names apply to the same species. 



50. Magiriopsis, new genus 

Type of genus: Sematoneura denticosella Dyar. 

Characters of Zamagiria except: Antenna of male 
unipectinate for basal half of shaft, serrate and shortly 
ciliate beyond; shaft with very shallow sinus near base, 
containing a few roughened scales and a short row of 
minute spines but without scale tuft. Labial palpi of 
male broad, dorsoventraUy flattened and appressed to 
face (not to each other as in Zamagiria and Anegcephale- 
sis); hoUowed umer surface; third segment greatly re- 
duced and completely hidden by scaling of second 
segment. Male head without scale tuft from upper 
edge of frons, not deeply grooved between antennae but 
with an enlarged scale tuft behind antennae. Maxillary 
palpus of female squamous (of male, as in Zamagiria, 
in the form of an aigrette). Forewing smooth; vein 2 
from weU before outer angle of cell; 6 slightly bent 
towards base but from below upper angle of cell, sepa- 
rate from stalk of 8-9 at base. Hind wing with cell 
less than one-third the length of wing. 

Male genitaha with complete transtilla. 

199. MagiriopBis denticosella (Dyar), new combination 

Figures 293, 769 

Sematoneura denticosella Dyar, Proc. U. S. Nat. Mus., vol. 42, 

p. 105, 1912. 
Hypsipyla denticosella (Dyar), Ins. Insc. Menstr., vol. 7, p. 41, 

1919. 
Crocidomera cristalis Hampson, Ann. Mag. Nat. Hist., ser. 10, 

vol. 4, p, 352, 1929 (new synonymy). 

Aigrette of male maxillary palpus whitish ocherous. 

Forewing reddish (rust) brown, the costal third lightly 
dusted with white, especially along the costa and be- 
tween the veins, the dark color somewhat accented on 
the veins and, on occasional female specimens, some 
black scaling on the veins before the usual location of 
subterminal line; the latter rarely indicated; antemedial 
line and discal spots obsolete. Hind wing translucent, 
opalescent with a smoky shade at apex; a dark line along 
termen and the outer veins faintly darkened. Alar 
expanse, 32-40 mm. 

Male genitaha with uncus narrowly triangulate. 
Apical process of gnathos a long, strong spine, sharply 
hooked at apex. Transtilla a weakly sclerotized, shield- 
like bridge attached to costal bases of harpes. Harpe 
with costa strongly sclerotized for basal third of harpe 
and terminating in a broad projection; sacculus with a 
broad projection from base and a broadly projecting 
clasper from apex. Aedeagus long and stout; penis 
armed with three clusters of strong spines (about as 
long as width of aedeagus). Vinculum elongate (two 
and one-half times as long as greatest width), stout, not 
appreciably tapering; posterior margin reinforced and 
squarely excised. 

Female genitalia with two irregularly shaped and 
strongly spined plates in bursa, and bursa at jimction 
with ductus bursa strongly sclerotized, the wrinkled 
sclerotization extending part way into the ductus; 
ductus bursae otherwise only sclerotized (weakly) at 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



95 



genital opening. Eighth-segment collar not fused 
ventrally. 

Type localities: Orizaba, Mexico (denticosella, in 
USNM); Juan Vifias, Costa Rica (cristalis, in BM). 

Food plant: Unknown. 

Disteibution: Mexico: Orizaba (Oct.), Misantla 
(Aug.). Guatemala: Cayuga (Jan., Aug.), Chejel 
(June). Costa Rica: Juan Vinas (Jan., Nov.). Co- 
lombia: Upper Rio Negro. Venezuela: Aroa, San 
Esteban Valley. British Guiana: Omai. Brazil: 
Ponte Nova (Rio Xingu, Amazonas), Santos (Mar.). 

Both denticosella and cristalis were described from 
females. The synonymy is obvious. Both Dyar and 
Hampson had seen males many years before but evi- 
dently overlooked them when writing their descriptions. 

Genus 51: Ancylostomia 

Venational division D. Forewing with veins 4 and 5 contiguous 
or stalked for about one-third their lengths from cell; 3 connate 
with 4 at base; 6 bent towards base, connate with the stalk of 
8-9. Hind wing with veins 4 and 5 stalked to middle; 3 closely 
approximate to the stalk of 4-5 at base; cell short. Labial palpus 
obliquely ascending with third segment long and porrect. Uncus 
pentagonal. Gnathos produced at apex into a pair of long, 
flattened, pointed, contorted, bandlike projections. Female 
with bursa copulatrix weakly sclerotized throughout; eighth- 
segment collar modified ventrally. Eighth abdominal segment 
of male with compound ventral tufts. 

51. Genus Ancylostomia Ragonot 

Ancylostomia Ragonot, Monograph, pt. 1, p. 567, 1893. (Type 
of genus: Myelois stercorea Zeller.) 

Tongue well developed. Antenna of male with a 
sinus and strong scale tuft in shaft at base, shaft other- 
wise flattened, weakly serrate and pubescent; of female, 
simple and pubescent. Labial palpus obliquely ascend- 
ing, with third segment porrect; second segment long, 
reaching well above vertex, on male hollowed to receive 
maxillary palpus; third segment slender, acuminate, on 
male half as long as second, on female about the same 
length as second segment. Maxillary palpus of male 
in the form of an aigrette ; of female, squamous. Fore- 
wing smooth; 11 veins; vein 2 from before but near 
lower outer angle of cell; 3 from the angle, connate with 
4; 4 and 5 contiguous or stalked for about one-third 
their lengths from cell ; 6 from upper angle of cell, con- 
nate with the stalk of 8-9, bent towards base; 8 and 9 
stalked for shghtly more than half their lengths; 10 from 
the cell, approximate for a short distance to the stalk of 
8-9; male without costal fold. Hind wing with vein 
2 from before lower outer angle of cell ; 3 from the angle, 
closely approximate to 4 at base; veins 2 and 3 very 
long; 4 and 5 stalked to middle; 7 and 8 anastomosed or 
contiguous for a short distance from cell (for less than 
half their lengths); cell less than one-third (about one- 
fourth) the length of wing; discocellular vein curved. 
Eighth abdominal segment of male with compound, 
ventral scale tufts. 

Male genitaha with uncus pentagonal (the sides 



parallel, the terminal margin angled). Gnathos pro- 
duced at apex into a pair of long, flattened, sharply 
pointed and slightly contorted, bandlike projections; 
the lateral arms short and articulated to base of uncus. 
Transtilla absent. Harpe elongate, slender, cucullus 
reduced, its apex bluntly pointed; sacculus produced 
at apex into a slender, free, curved hook. Penis armed 
with a pair of thin, somewhat flattened and twisted 
spines as long (or nearly as long) as aedeagus. Vincu- 
him. shghtly longer than greatest width; terminal margin 
broadly rounded. 

Female genitalia without signum; bursa weakly 
sclerotized throughout; ductus bursa strongly sclero- 
tized except for a narrow space beyond junction of 
bursa, at junction with bursa finely ridged and spined, 
the ridges and spines extending for a short distance into 
bursa, at genital opening the margins of ductus fuse 
into eighth-segment collar. Eighth-segment collar very 
strongly sclerotized; enlarged but not fused ventrally; 
its ventrocaudal angles produced and pointed; from its 
ventrolateral angles a pair of invaginated sclerotized 
pockets. 

The genus is easily identified by its pecuharly de- 
veloped gnathos and the eighth-segment coUar of the 
female. Its species are tropical American in distribu- 
tion with (in the case of stercorea) a shght extension of 
range into southern Florida. 

200. Ancylostomia stercorea (Zeller) 

Figures 295, 771 

Myelois stercorea Zeller, Isis von Oken, 1848, p. 873. 

Anerastia ignobilis Butler, Proc. Zool. Soc. London, 1878, p. 494. 

Pempelia diffissella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 

178, 1881. 
Ancylostomia stercorea (Zeller) Ragonot, Monograph, pt. 1, p. 

568, 1893. 

Forewing pale ocherous along the costa, through the 
median fold and over most of the basal area; some 
pinkish brown shading between the veins in the pale 
areas giving the ocherous ground color a rosy tint 
(especially on reared and fresh specimens) ; the lower 
outer half of wing heavily shaded with brown more or 
less dusted with black (on reared examples this area is 
decidedly blackish brown, its inner margin obhque 
from lower outer angle of cell to basal third of inner 
margin) ; a distinct brown or black streak under vein 8 
from apex about half-way to ceU and a similar longer 
dark streak along vein 6 from terminal margin to the 
cell; a thin whitish fine along the lower margin of cell 
from beyond base and continuing into vein 4 for a short 
distance; a similar shorter white fine on the subbasal 
half of vein lb; a small black dot on the middle of the 
white streak on vein lb and a similar black dot above 
it on the white streak on lower margin of cell; a single, 
large, conspicuous black discal spot at lower outer angle 
of cell; antemedial and subterminal lines obsolete; a 
few black dots or streaklets in outer areas about the 
middle of veins 2, 3, and 4 and on dark specimens some 
blackish or dark brown shading on upper and lower 



96 



rnsriTED states national museitm bulletin 207 



veins of cell at base of wing. Hind wing translucent 
white with a smoky shade along costa, towards apex 
and along termen; the veins shghtly darkened. Alar 
expanse 16-28 mm. 

Genitalic characters as given for the genus. Eighth- 
segment collar of female laterally corrugated. 

Type localities : BrazU (stercorea, in BM) ; Jamaica 
(ignobilis, iu BM); Honda, Colombia {diffissella, in 
BM). 

Food plant: Cajanus cajan (larvae feeding in the 
pods). 

Distribution: United States: Florida, Cocanut 
Grove (Apr., May), Goulds (May), Homestead (Apr.), 
Jupiter (Apr.), Miami (May). Cuba: Santiago Prov- 
ince (May, June, Oct., Dec), Santiago de las Vegas 
(May). Haiti (June). Dominican Eepublic (Aug.). 
PuEETO Eico: Isabela (Apr.), Puerto Real (Vieques 
Isl., Apr.). Virgin Islands: Kingshill (St. Croix; 
June, "Nov.-Dec"). Jamaica (Mar.). Bahamas: 
Nassau (May). Grenada: Nevis (Jan.); St. Kitts 
(June). Trinidad (Mar., May). Mexico: Cuerna- 
vaca (July), Jalapa, Orizaba, Zacualpdn (July). Gua- 
temala: Chejel (July, Aug.), Volcdn Santa Maria 
(June, July). Costa Rica: Juan Vinas (June.) Pan- 
amX: La Chorrera (May), Rio Trinidad (Mar.). Co- 
lombia: Honda. French Guiana: Cayenne. Bra- 
zil: Castro. 

The food plant and Florida records are from a large 
series reared by the Special Survey of the U. S. Bureau 
of Entomology and Plant Quarantine in 1944. We also 
have a couple of reared adults and several collected 
larvae from chickpea (Cicer) and black-eyed pea 
(Dolichos). The favored host, however, seems to be 
the pigeonpea {Cajanus). In the pods of that plant 
the larvae are abundant throughout the West Indies 
and in southern Florida. 

201. Ancylostomia sauciella (Zeller) 

Figure 296 

Pempelia sauciella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 

183, 1881. 
Ancylostomia sauciella (Zeller) Ragonot, Monograph, pt. 1, p. 

569, 1893. 

Forewiag cinnamon red mixed with rose; costa paler; 
a narrow subcostal reddish brown shade extending from 
base to apex; the veins somewhat accented by reddish 
scaliug; a small blackish discal dot at lower outer angle 
of cell; a similar dot on basal third of vein lb ih a white 
streaklet on the vein; from apex a short oblique blackish 
shade. Hind wing yellowish white, translucent; a thin 
dark line along termen and the veins slightly darkened. 
Alar expanse, 20 mm. 

Male genitalia as in stercorea except uncus narrower 
and apical projecting arms of gnathos shorter and 
decidedly broader. 

Type locality: Maraquita, Colombia (type in BM). 

Food plant: Unknown. 

Known only from the type. 



202. Ancylostomia argyropMeps Dyar 

Ancylostomia argyrophleps Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 406, 1914. 

Similar to sauciella except: Costa and lower half of 
basal area carneous white; a broad subcostal band 
extending from base to apex, red-brown shaded with 
black especially towards apex; lower outer area of the 
same color with a smoky fuscous shade along outer half 
of inner margin from near tomus; a white line along 
lower margin of cell and vein 5 from basal third, enclos- 
ing a black dot at lower outer angle of cell; a similar 
black dot on basal third of vein lb. Hind wing trans- 
lucent, semi-irridescent white with a faint smoky tint, 
the latter more pronounced towards apex; veins dark- 
ened in outer area on females, not appreciably so on 
males. Alar expanse, 20-25 mm. 

Male genitalia as in stercorea except apical processes 
of gnathos a trifle broader (but not so broad as on 
sauciella). Eighth-segment collar of female smooth. 
Female genitalis otherwise essentially as in stercorea. 

Type locality: Orizaba, Mexico (type, cT, in 
USNM). 

Food plant: Unknown. 

Distribution: Mexico: Orizaba (Aug.), Cuemavaca 
(July). Guatemala: Chejel (July). 

203. Ancylostomia euchroma Dyar 

Ancylostomia euchroma Dyar, Ins. Insc. Menstr., vol. 7, p. 63, 
1919. 

Forewing below the cell from base to tornus red- 
brown; the upper area white-lined between the veins 
and along the median fold in cell; the veins red-brown; 
a black dot at lower outer angle of cell, one on lower 
margin of cell before its middle and another on basal 
third of vein lb; a diffused pale shade surrounds this 
last dot and extends obliquely backward to inner 
margin, an obhque line of black dots on veins 2, 3, and 
4; on the female a smoky brown shade from apex 
extending narrowly along costa to base; on male the 
shade from apex is short and cinnamon red, and the 
dark area on lower half of wing is a bright cinnamon 
red. Hind wing in the male translucent white with a 
faint smoky shade at apex; on the female the smoky 
shade extends further back from apex and outer margin 
and the veins in outer area are appreciably darkened. 
Alar expanse, 24-25 mm. 

Male genitalia with the projecting apical bands from 
gnathos as broad as those of sauciella but longer (at 
least as long as those of stercorea). Female genitaha 
with the sclerotized ribbing and spining at junction of 
bursa and ductus bursae slightly stronger than those of 
either stercorea or argyrophleps. Eighth-segment collar 
very weakly corrugated. 

Type locality: Castro, Parand, Brazil (type in 
USNM). 

Food plant: UnknoMTi. 

Represented only by the female type and male 
paratype from the type locality. May be only a race 
of sauciella. 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



97 



Genus 52: Caristanius 

[Venational division B. Forewing with veins 4-5 separated at 
base and divergent shortly beyond, smooth; hind wing with vein 
3 approximate to the stalli of 4-5 at base. Eighth abdominal 
segment of male with compound tufts. Antenna of male with 
sinus and scale tufts on base of shaft. Labial palpus obliquely 
ascending; second segment of male grooved. Ma.xillary palpus 
of male in the form of an aigrette. Male genitalia with a pair of 
long, strongly sclerotized arms from ventrolateral margins of 
uncus; transtilla absent; gnathos absent; a single strong cornutus 
on penis. Female genitalia with signa present, developed as 
coarsely spined plates; ductus bursae ribbonlike, sclerotized.] 



52. Caristanius, new genus 

Type of genus: Oligochroa pellucidella Ragonot. 

Tongue well developed. Antenna of male with sinus 
and stout scale tuft on base of shaft, finely pubescent; 
of female simple. Labial palpus obliquely ascending; 
on male stout and smooth scaled (the palpi appressed 
to each other), second segment long, reaching well above 
vertex, grooved to hold maxillary palpus, third segment 
very short, hidden in scaling of second; on female slen- 
der, shorter, reacliing slightly above vertex, more 
roughly scaled, third segment over half as long as sec- 
ond, acuminate. Maxillary palpus of male in the form 
of an aigrette; of female squamous. Forewing smooth; 
11 veins; vein 2 from before lower outer angle of cell; 

3 from the angle; 4 and 5 shortly separated at base and 
divergent very shortly beyond; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for half or less 
than half their lengths; 10 from the cell; male without 
costal fold. Hind wing with vein 2 from before lower 
outer angle of cell; 3 from the angle, approximate to 
stalk of 4-5 at base; veins 2 and 3 of moderate length; 

4 and 5 stalked for half their lengths ; 7 and 8 approxi- 
mate beyond cell (for less than half their lengths) ; cell 
less than half the length of wing (but more than one- 
third) ; discocelhdar vein curved. Eighth abdominal 
segment of male with compound ventral scale tuft. 

Male genitalia with uncus short, decidedly broader 
than long; a pair of long, strong, sclerotized arms from 
its ventrolateral angles. Gnathos absent (unless the 
fused arms from uncus can be interpreted as parts of 
this organ, which is extremely doubtful). Transtilla 
absent. Harpe with costa strongly sclerotized and with 
a strong projection from its base or a strong clasperlike 
projection from below it near base; sacculus short, nar- 
row, weak, bluntly produced at apex; cucullus very nar- 
row, weak and reduced. Anellus a narrow band with a 
central, bifurcate, bandlike projection, flanked by elon- 
gate, lateral lobes. Aedeagus elongate, moderately 
slender, straight; penis armed with a single strong cornu- 
tus and some fine spines and granulations. Vinculum 
stout, as long as or longer than greatest width. 

Female genitalia with signa present, developed as 
elongate, curved, sclerotized, strongly and coarsely 
spined plates; ductus bursae flattened, ribbonlike, 
strongly sclerotized except for a narrow space near 
middle, also granulate towards bursa, the granulations 
extending into bursa; at genital opening ductus bursae 

300329 — 56 8 



very strongly and broadly sclerotized ; ductus seminalia 
from bursa near junction of bursa and ductus bursae. 

The species referred here have been placed in Elasmo- 
•palpus. They agree with the type of that genus {ligno- 
sellus) on antennal, palpal, and venational characters; 
but differ strikingly on male and female genitalic struc- 
ture. The characteristic uncus with its long, produced, 
basal arms, the reduced, weak sacculus, the lack of any 
distinguishable gnathos, and the flattened, ribbonlike 
ductus bursae with its strongly sclerotized development 
at genital opening at once distinguish Caristanius from 
Elasmopalpus. 

204. Caristanius pellucidellus (Ragonot), new combination 

Figures 297, 775 

Oligochroa pellucidella Ragonot, Bull. Soc. Ent. France, 1888, 

p. cxl. — Moschler, Die Lepidopteren-Fauna von Portorico, 

p. 329, 1890. 
Elasmopalpus pellucidellus (Ragonot), Monograph, pt. 1, p. 429, 

1893. 
Rhodophaea melanoplaga Hampson, in Ragonot, Monograph, pt. 

2, p. 519, 1901 (new synonj'my). _ 

Ground color of forewing variable, pale brownish gray 
to dark gray %vith a faint purplish or reddish brown tint; 
transverse markings obsolete on most specimens; on 
some the antemedial line indicated by a narrow, very 
faint, pale line between vein lb and inner margin, pre- 
ceded by a pale reddish or brownish patch more or less 
overlaid by black scaling (on most specimens before me 
this spot entirely absent) ; subterminal line distinguish- 
able only on a few specimens, very faint, indicated 
chiefly by some short blackish streaklets on the veins; 
discal spots usually distinct but faint, blackish, sepa- 
rated; a row of black dots along termen. Hind wing 
transparent white with a dark shade along costa and at 
apex and extending as a dark line downward along ter- 
men; on the males this dark line extends only to about 
middle of termen, on females to or nearly to anal angle 
of wing. Alar expanse, 19-25 mm. 

Male genitalia with costa of harpe broadly and stoutly 
sclerotized, a large oval projection from base and its 
apical end folded backward and contorted. Cornutus 
about one-third as long as aedeagus. Vinculum about 
as long as greatest width. 

Female genitalia with granidations of ductus bursae 
extending for a very short distance into bursa. Eighth- 
segment collar not extended to ventral surface; at- 
tached laterally to the extended sclerotization of ductus 
bursae at genital opening; stoutly sclerotized dorsally 
and with an inwardly projecting curved sclerotized 
apron; between this apron and ductus bursae a gland 
of unknown function extends into abdomen, terminat- 
ing in a bulb with thickened membrane. This structure 
not noted on other species of the genus. 

Type localities: Puerto Rico {pellucidellus, in Paris 
Mus.); Sao Paulo, Brazil {melanoplaga, in BM). 

Food plant: Unknown. 

Distribution: Puerto Rico: Puerto Real (Vieques 
Isl., Apr., July), Rio Piedras (Aug., Sept.), San German 
(Apr., Aug.). St. Vincent. Jamaica. Surinam: Zan- 



UNITED STATES NATIONAL MUSEXUVI BULLETIN 207 



derij I (Apr.). Bbazil: Sao Paulo, "S. E. Brazil" 
[probably Castro]. 

The species is somewhat variable in color and mark- 
ings, but is easily identified by its genitalia. Hampson's 
melanoplaga was based on large Brazilian females (25 
mm.). All specimens from the West Indies and Suri- 
nam that I have seen are smaller (19 to 21 nam.). How- 
ever, there does not seem to be any reason to keep 
melanoplaga as a racial designation on a mere difference 
in size, as there is nothing else to distinguish the Bra- 
zilian examples. 

205. Carbtanius decoloralis (Walker), new combination 

Figures 298, 773 

Trachonitis decoloralis Walker, List, vol. 27, p. 42, 1863. 
Nephopteryx meiagrammalis Walker, List, vol. 27, p. 42, 1863. 
Nephopieryx furfurella Hulst, Ent. Amer., vol. 3, p. 131, 1887; 

Phycitidae of N. Amer., p. 143, 1890 (new synonymy). 
BHasmopalpus decoloralis (Walker) Ragonot, Ent. Amer., vol. 5, 

p. 115, 1889; Monograph, pt. 1, p. 420, 1893.— McDun- 

nough. Check list, No. 6228, 1939. 
Elasmopalpus floridellus Hulst, JPhycitidae of N. Amer., p. 158, 

1890. — Ragonot, Monograph, pt. 1, p. 423, 1893. 
Elasmopalpus decoralis Hulst, Phycitidae of N. Amer., p. 158, 

1890 (misspelling for decoloralis). 
Elasmopalpus decorellus Hulst, in J. B. Smith, List of the Lepi- 

doptera of Boreal America, No. 4341, 1891 (misspelling for 

decoloralis) . 
Elasmopalpus furfurellus (Hulst), Barnes and McDunnough, 

Contributions, vol. 3, p. 196, 1916. — McDunnough, Check 

list, No. 6230, 1939. 

Forewing bluish gi'ay or pale fawn gray with more or 
less white dusting over median area; transverse lines 
indistinct; antemedial line indicated chiefly by its 
blackish outer border broken into dots on vein lb, lower 
vein of cell and a subcostal spot or short dark streak 
from costa to top of cell; preceding antemedial line a 
reddish patch on inner margin extending to or into cell 
and more or less shaded by black scaling; sub terminal 
line sinuate, pale and very faint; lower discal spot at 
end of cell distinct, the upper discal dot much smaller, 
sometimes distinct but frequently obscured; a row of 
black dots along termen. Hind wing translucent, whit- 
ish with a pronounced smoky tint, especially over outer 
half of wing; the veins more or less darkened and a fine 
dark line along termen. Alar expanse, 21-25 mm. 

Male genitalia with projecting arms from uncus 
somewhat curved. Costa of harpe sclerotized for the 
length of the harpe, narrowly sclerotized beyond base; 
a strong, curved, clasperlike projection from below costa 
near base, projected beyond costal edge. Cornutus as 
long as aedeagus. Vinculum considerably longer than 
greatest width. 

Female genitalia with granulations of ductus bm-sae 
extending deeply into bursa; ductus bursae scobinate 
on lower surface at genital opening. Eighth-segment col- 
lar narrowed dorsally, complete but not fusing ven- 
traUy, without sclerotized dorsal apron. 

Type localities: United States (decoloralis and me- 
tagrammalis, in BM) ; Florida (JurJureUus and floridellits, 
in AMNH, ex Rutgers). 



Food plant: Chamaecrista spp. (brachiata, fascicu- 
lata, robusta) larvae feeding on the leaves. These rec- 
ords from rearings by the Special Survey (1944) of the 
Division of Foreign Plant Quarantine of the U. S. 
Bureau of Entomology and Plant Quarantine. 

Distribution: Florida, Orlando (Feb.), no specific 
locahty (Mar.), Fort Myers (May), St. Petersburg 
(June), Stuart (May), Tampa, Vero Beach (Apr., May, 
June, Oct., Dec); Texas, Brownsville; South Carolina, 
Florence (Jime), HUtonhead Isl. (Aug.), Pawleys Beach 
(Sept.). 

This species seems to be confined to the southeast- 
ern United States. In his description of decoloralis 
Walker mentions the protruding arms of the imcus. 
Hulst noted similar structures in his floridellus and asso- 
ciated them with the genitalia, suspecting the synonymy 
of his and Walker's species. I have before me a photo- 
graph of the genitalia of the type of decoloralis supplied 
by Clarke. They are identical with those of the male 
type of floridellus. The synonymy of floridellus and 
furfurellus was established by Barnes and McDunnough 
(1916) and that of metagrammalis with decoloralis by 
Ragonot (1889). 

206. Caristanius guatemalellus (Ragonot), new combination 

FlGTTHE 774 

Salehria guatemalella Ragonot, Nouv. Gen., p. 18, 1888. 
Laodamia guatemalella (Ragonot), Monograph, pt. 1, p. 414, 
1893. 

This species is apparently known only from the female 
type. Ragonot's description and figure suggest a large, 
pale brownish, suffused form of peUucidellus; the fore- 
wing pale ocherous brown tinted with reddish, trans- 
verse lines absent, the lower discal spot distinct, and the 
veins sparsely powdered with blackish scales; hind wing 
semitransparent, white faintly tinted with ocherous. 
Alar expanse, 26 mm. 

The genitalia determined the present generic refer- 
ence. The granulations of ductus bursae extend deeply 
into the bursa as in decoloralis; the eighth-segment collar 
has a dorsal, sclerotized, invaginated apron smaller 
than and differently shaped from that of peUucidellus, 
and the collar itself is completely sclerotized ventrally. 

Type locality: San Geranimo, Guatemala (type in 
BM). 

Food plant: Unknown. 



Genus 53: Etiella 

[Venational division B. Forewing with veins 4r-5 separated and 
divergent from base, a raised-scale ridge beyond base; hind wing 
with vein 3 approximate to the stalk of 4r-5 at base. Eighth 
abdominal segment of male with paired tufts. Labial palpus 
porrect, long; second segment of male grooved. Maxillary pal- 
pus in the form of an aigrette. Male genitalia with apical process 
of gnathos a simple, short hook; transtilla incomplete and 
vestigial; harpe with a strong curved arm projecting the length 
of the harpe from base of costa, harpe otherwise weakly sclero- 
tized; two strong cornuti on penis. Female genitalia with signa 
developed as curved, sclerotized bands armed with slender 
spines; ductus bursae short, tubular, sclerotized.] 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



99 



53. Genus Etiella Zeller 

Eiiella Zeller, Isis von Oken, 1839, p. 179; 1846, p. 733.— Heine- 
mann, Die Schmetterlinge Deutschlands und der Schweiz, 
Abt. 2, vol. 1, pt. 2, p. 154, 1865.— Meyrick, Proc. Linn. 
Soc. New South Wales, vol. 3, p. 629, 1882.— Hulst, Phy- 
citidae of N. Amer., p. 169, 1890; U. S. Nat. Mus. Bull. 52, 
p. 428, 1902.- Ragonot, Monograph, pt. 1, p. 569, 1893.— 
Spuler, Die Schmetterlinge Europas, vol. 2, p. 208, 1910. — 
Forbes, Cornell Mem. 68, p. 629, 1923.— Janse, Journ. Ent. 
Soc. South Africa, vol. 7, p. 15, 1944. (Type of genus: 
Phycis zinckenella Treitschke.) 

Ramphodes Gu6n6e, Europaeorum Microlepidopterorum index 
methodicus . . ., p. 81, 1845. (Type of genus: Phycis 
zinckenella Treitschke.) 

Tongue well developed. Antenna of male with basal 
segment enlarged and bearing a short, bluntly pointed 
projection on inner side near base; shaft with sinus and 
large hair and scale tuft at base, pubescent; antenna of 
female simple. Labial palpus porrect; very long (the 
length of head and thorax); smoothly scaled; second 
segment about five times the length of third on male 
and grooved to hold maxillary palpus; third segment 
short on male, longer on female, acuminate, frequently 
bent downward, especially on female. Maxillary palpus 
of male in the form of an aigrette; of female small, 
squamous. Forewing with ridge of raised scales beyond 
base; 11 veins; vein 2 from before but near lower outer 
angle of cell; 3 from the angle, approximately equi- 
distant from 2 and 4 ; 4 and 5 shortly separated at base, 
diverging from cell; 6 from below upper angle of cell, 
straight; 8 and 9 stalked for half their lengths; 10 from 
the ceU, separated and divergent from the stalk of 8-9 ; 
male without costal fold. Hind wing with vein 2 from 
well before lower outer angle of cell; 3 from the angle, 
appreciably shorter than 2, closely approximate to the 
stalk of 4-5 at base (separated by a very short vein) ; 
4 and 5 normally stalked (rarely weakly anastomosed 
or contiguous) to about middle; 7 and 8 closely approxi- 
mate for less than haK their lengths from cell; cell about 
one-third the length of wing on male, slightly longer on 
female; discocellular vein curved. Eighth abdominal 
segment of male with a pair of weak ventrolateral hair 
tufts. 

Male genitalia with uncus hoodlike, its apical margin 
evenly rounded. Apical process of gnathos a simple, 
sharp hook. Transtilla incomplete and vestigial (its 
elements rarely distinguishable except under high 
magnification). Harpe with a strongly sclerotized, 
curved, tapering, pointed arm projecting from base of 
costa and as long as costa; remainder of harpe wealdy 
sclerotized and abruptly narrowed at middle (the shape 
of harpe probably a specific character). Anellus V- 
shaped; its arms rather broad, blunt and haired. 
Aedeagus moderately long, stout; penis armed with 
two strong cornuti. Vinculum stout ; as long as greatest 
width; evenly tapering to a blunt point. 

Female genitaha with bujsa elongate, finely scobinate, 
armed with signa consisting of curved, sclerotized bands 
armed with slender spines and situated near junction of 
bursa and ductus bursae ; a sclerotized lobe on bursa near 
ductus bursae; ductus bursae short (much shorter than 



bursa), tubular, sclerotized, ribbed and broadened 
towards genital opening; ductus seminalis from bursa 
adjacent to sclerotized lobe. 

A distinct genus, not to be confused with anything 
else and easily identified by its combination of antennal, 
palpal, wing, and genitalic characters. Ragonot, Hulst, 
and Janse list five Walker names in the generic synony- 
my. These supposed genera were based on Old World, 
tropical species which have all been referred as s3tio- 
njins of zinckenella. I have no reason to question this 
synonymy but have omitted the references as I have 
not been able to check the genitalia or their types. The 
only species occiu-ring in the New World is zinckenella. 

The larva differs from typical phycitid larvae in that 
it lacks altogether the sclerotized rings about seta lib 
of mesothorax and seta III of the eighth abdominal 
segment. 

207. Etiella zinckenella (Treitschke) 
Figures 17, 326, 840 

Phycis zinckenella Treitschke, Die Schmetterlinge von Europa, 
vol. 9, pt. 1, p. 201, 1832. 

Phycis etiella Treitschke, Die Schmetterlinge von Europa, vol. 
10, p. 3, p. 174, 1835. — Duponchel, Histoire naturelle des 
l^pidoptSres, ou papillons de France, vol. 10, p. 180, 1836. — 
Milliere, Iconographie et description de chenilles et l§pidop- 
tfires inedits, vol. 1, p. 248, 1861. (Originally proposed as 
new name for zinckenella.) 

Pempelia Eiiella zinckenella (Treitschke) Zeller, Isis von Oken, 
1839, p. 179; 1846, p. 755.— Herrioh-Schafifer, Systematische 
Bearbeitung der Schmetterlinge von Europa, vol. 4, p.^'72, 
1849. 

Ramphodes zinckenella (Treitschke) Gu^n^e, Europaeorum 
Microlepidopterorum index methodicus . . . , p. 81, 1845. 

Eiiella zinckenella (Treitschke) Heinemann, Die Schmetterlinge 
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 154, 
1865. — Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 177, 
1881.— Hulst, Phycitidae of N. Amer., p. 170, 1890; U. S. 
Nat. Mus. Bull. 52, p. 428, 1902.— Ragonot, Monograph, 
pt. 1, p. 572, 1893.— Chittenden, U. S. Dep. Agr. Bur. Ent. 
Bull. 82 (pt. 3), p. 25, 1909. — Essig, Insects of western 
North America, p. 709, 1926. — Walcott, Journ. Agr. Univ. 
Puerto Rico, vol. 20, no. 1, p. 476, 1936.— McDunnough, 
Check list. No. 6274, 1939. 

Etiella zinckenella schisticolor Zeller, Horae Soc. Ent. Rossicae, 
vol. 16, p. 178, 1881.— Hyslop, U. S. Dep. Agr. Bur. Ent. 
Bull. 95, pt. 6, p. 82, 1912. 

Etiella villosella Hulst, Ent. Amer., vol. 3, p. 133, 1887. 

Etiella schisticolor (Zeller) Ragonot, Ent. Amer., vol. 5, p. 116, 
1889; Monograph, pt. 1, p. 274, 1893.— Hulst, Phycitidae 
of N. Amer., p. 170, 1890. 

Etiella rubribasella Hulst, Phycitidae of N. Amer., p. 170, 1890. — 
Ragonot, Monograph, pt. 1, p. 572, 1893. 

Forewing gray more or less lightened by white 
scaling, especially in median areas; a broad white band 
along costa, extending from base to or nearly to apex; 
extreme costal edge dark gray to red; on occasional 
specimens some red scaling at extreme base of wing and 
in outer median area; transverse lines obsolete; the 
antemedial line replaced by a ridge of raised scales near 
basal third and extending from inner margin to top of 
cell, the raised scales metallic ocherous or orange red 
bordered outwardly by a broad ocherous or orange 
patch; discal spots obsolete. Hind wing whitish with 
a faint smoky tint, to dark smoky fuscous; the veins 



100 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



and terminal margin darker. Alar expanse, 15-28 mm. 

Genitalia as given for the genus. 

Type localities: S|icily (zinckenella, in Hungarian 
Nat. Mus., Budapest; etiella); California (schisticolor, 
in BM); Colorado (mllosella, in AMNH, ex Eutgers); 
Florida (ruhribasella, in AMNH, ex Rutgers) . 

Food plants : Pods and seeds of various Leguminosae 
(Astragalus, Cajanus, Colutea, Crotolaria, Dolichos, 
Glycina, Lwpinus, Phaseolus, Pisum, Vicina, Vigna). 

Disteibution: Throughout the tropical and sub- 
tropical areas of the world and in the warmer temperate 
regions of Europe, Asia, and North America. The fol- 
lowing American records are from specimens before me. 
United States: Florida, Archer (Mar.), Buena Vista 
(May), Crescent City (Mar.), Dade City (Aug.), Eg- 
mont (Apr., June), Everglade (Apr.), Jupiter (Apr.), 
Lake Alfred (June, July, Nov.) ; Texas, Brownsville 
(June, July, Aug.), CotuUa (May), Dallas (June), 
Gainesville (Nov.), KerrvUIe, Logan (Jime); Arizoria, 
Baboquivari Mts. (June, July, Aug.), Nogales (July), 
Woodruff (June); California, Alameda County (Sept.), 
Garden City (Jan., Feb.), Loma Linda (June, Aug., 
Sept., Oct.), Palo Alto (Sept.), San Diego (May, Jime, 
Sept.), San Gabriel Mts. (June), Santa Paula, upper 
Ojai (July); Washington, Pullman (May, July, Aug.), 
Yakima (May, June), Walla Walla (June, Aug.), 
Wenatchee (July); Idaho, Springfield (June); Nevada, 
Pyramid Lake, Reno; Utah, Provo (July), Vineyard 
(June); Colorado, Boulder Creek Canyon (May), Den- 
ver (May) ; Oklahoma, Stillwater (Aug.) ; Rhode Island, 
Weekapaugh (Aug.) ; New Hampshire, Hampton (Sept.) . 
Canada: Saskatchewan, Oxbow (June). Cuba: San- 
tiago (Feb., June, Oct., Nov.). Puerto Rico: Dorado 
(May), Isabella (Jan.), Mayagiiez (Jan.), Palmas Aba- 
jas (near Guayaman), Puerto Real (Vieques IsL, Apr.), 
Rio Piedras (Apr.), San Germdn (Apr.). Grenada. 
Jamaica: Newport (Feb.). Mexico: Eldorado (Mar.), 
Mexico City (Nov.), Oaxaca, Orizaba, Tehuacdn (Apr.). 
French Guiana: St. Laurent Maroni. Brazil: Santa 
Catarina (July, Dec), Sao Paulo (May). Uruguay: 
Montevideo. Paraguay: ViUarrica (Jan., Apr., July, 
Sept., Nov.). Per^: Angasmarca. GalApagos: Con- 
way Bay (Apr.) . 

Presumably to be found in every Central and South 
American country. 

This species is of economic importance as an occa- 
sional pest of cultivated beans and to American ento- 
mologists is known as the "limabean pod borer." It 
has an extensive literature and has been described under 
many names, having at least 13 Old World syn- 
onyms. I have omitted these (for reasons given under 
discussion of the genus) and listed only the American 
synonyms. Some authors have treated schisticolor and 
ruhribasella as distinct species, others as races of 
zinckenella. They appear as the latter in our latest list 
(McDunnough, 1939). However, they are no more 
than color forms intergrading with the typical zincke- 
nella and deserve no separate designation. The 
accepted Old World synonymy is given in the Ragonot 
Monograph (1893). The best and most complete 



accounts of the life history and immature stages will be 
found in the U. S. Department of Agriculture bulletins 
cited here (Chittenden, 1909, and Hyslop, 1912). For 
additional references the "Review of Applied Ento- 
mology" should be consulted. 

Genus 54: Glyptocera 

[Venational division B. Forewing with veins 4^5 connate, 
smooth; hind wing with veins 4-5 distinctly stallsied for over half 
their lengths; 3 connate with the stalk of 4^5. Eighth abdominal 
segment of male with compound tufts. Antenna of male with 
shallow, spined sinus at base of shaft. Labial palpus upturned, 
rough scaled beneath. Maxillary palpus squamous. Male 
genitalia with sacculus of harpe produced at apex as a long, free 
spine; apex of gnathos a short, stout hook; transtilla complete 
but weakly sclerotized; a single strong cornutus on penis. 
Female genitalia with bursa partly sclerotized, otherwise finely 
spined but without definable signa; ductus bursae flattened and 
partially sclerotized.] 

54. Genus Glyptocera Ragonot 

Glyptocera Ragonot, Ent. Amer., vol. 5, p. 114, 1889; Bull. Soc. 
Ent. France, 1890, p. vii; Monograph, pt. 1, p. 209, 1893. — 
Hulst, Phycitidae of N. Amer., p. 140, 1890. — Forbes, 
Cornell Mem. 68, p. 621, 1923. (Type of genus: Nepho- 
pteryx consobrinella Zeller.) 

Tongue well developed. Antenna of male pubescent, 
shaft with shallow sinus towards base containing a row 
of short, toothlike spines; antenna of female simple. 
Labial palpus upturned; second segment rough scaled 
beneath; third segment about one-third as long as 
second, acuminate. Maxillary palpus rather large, 
squamous. Forewing smooth; 11 veins; vein 2 from 
before but rather near lower outer angle of cell; 3 from 
the angle, separated from 4-5 at base, but about half 
as far from them as from 2; 4 and 5 connate; 6 from 
below upper angle of cell, straight ; 8 and 9 long stalked 
(for slightly over two- thirds their lengths) ; 10 from the 
cell, closely approximate to the stalk of 8-9 for nearly 
half its length; male without costal fold. Hind wing 
with vein 2 from before but near lower outer angle of 
cell; 3 from the angle, connate with the stalk of 4-5; 
4 and 5 distinctly stalked for over half their lengths; 7 
and 8 closely approximate for a short distance from 
cell; cell about half the length of wing; discocellular 
vein curved. Eighth abdominal segment of male with 
compoxmd ventral scale tuft. 

Male genitalia with apical process of gnathos a short, 
stout, simple hook. Transtilla complete but very 
weakly sclerotized, a simple, arched band. Harpe with 
a row of fine, erect hairs from below costa near base; a 
slender, short, clasperlike projection from just below 
base of costa; sacculus produced at apex as a long, free, 
spinelike projection. Vinculum a small V-shaped plate 
with elongate, bandlike, central projection attaching 
to apex of aedeagus. Penis armed with a single, strong 
cornutus. Vinculum stout, longer than greatest width. 

Female genitalia with bursa sclerotized over nearly 
half of one surface, densely and finely spiaed over most 
of remaining area; ductus bursae flattened, a broad 
sclerotized band extending its length on ventral surface; 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



101 



ductus seminalis from bursa near junction of bursa and 
ductus bursae; genital opening simple. 

The combination of male characters and the rather 
broadly squamous maxillary palpi distinguish the 
genus. The long, straight, free sacculus at once identi- 
fies it and distinguishes it from the genera which follow, 
and which are related to Nephopteryz and Salehria. The 
weak transtilla also occurs in Meroptera and some species 
of Nephopteryz. The female genitalia are similar to 
those of Nephopteryr. 

Glyplocera contains but the one North American 
species. 

208. Glyptocera consobrinella (ZeUer) 
Figures 327, 811 

Nephopteryz consobrinella Zeller, Verb, zool.-bot. Ges. Wien, vol. 
22, p. 548, 1872. 

Glyptocera consobrinella (Zeller) Ragonot, Ent. Amer., vol. 5, p. 
114, 1889; Monograph, pt. 1, p. 210, 1893.— Hulst, Phyciti- 
dae of N. Amer., p. 140, 1893.— Forbes, Cornell Mem. 68, 
p. 621, 1923.— McDunnough, Check list. No. 6148, 1939. 

Ambesa busckella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 
108, 1904. — Barnes and McDunnough, Contributions, vol. 
3, p. 196, 1916. 

Forewing ashy gray shaded and marked with black, 
the blackish shade most conspicuous broadly bordering 
the antemedial line on inner side, expanded narrowly 
along inner margin to base and also broadening the 
black outer border of the line at costa; basal area other- 
wise pale clay color more or less shaded with pale sal- 
mon especially in lower fold; on many specimens a 
blotch of the same salmon shade over the middle of 
inner margin; antemedial line well out beyond basal 
third, slightly oblique, sharply sinuate, whitish gray 
with black inner and outer bordering lines; subterminal 
line outwardly rounded at middle and angled above 
and below the bulge, margined inwardly and outwardly 
by distinct black lines; discal spots confluent, forming 
a black line along discocellular vein which expands be- 
low into short black streaklets or smudges on the lower 
veins; terminal dots confluent, forming a black line 
along termen. Hind wing smoky white with a faint 
ocherous tint; the veins not appreciably darkened; a 
narrow dark line along termen. Alar expanse, 20-25 
mm. 

Genitalia as given for the genus. On some males in 
addition to the strong cornutus there is a second much 
smaller and weaker spine but this is variable and ap- 
parently not a constant structure. 

Type localities: Texas (consobrinella, in MCZ); 
Plummers Island, Md. (busckella, in USNM). 

Food plant: Viburnum, maple (Dyar and Ely 
rearings). 

Distribution: United States: Maine, Lincoln ville 
(May, reared by Dyar on Viburnum), Orono, Sebec 
Lake (July), Weld (July); New Hampshire, Hampton 
(July); Vermont, Clarendon; Massachusetts, Framing- 
ham (June), New York, Plattsburg (July), Valcour Isl. 
(July) ; Connecticut, East River (Jime, July) ; New Jer- 
sey, Basldng ^idge; Pennsylvania, New Brighton (June, 
July, Aug.), Pittsburgh (July); Maryland, Plummers 



Isl. (May, Aug.); District oj Columbia, Washington 
(July, reared by Dyar from larva on maple); Texas, 
Kerrville (Mar.); Illinois, Chicago (July). Canada: 
Ontario Trenton (July) ; Quebec, St. Johns (June) ; Nova 
Scotia, Cape Breton Isl. (June); Newfoundland, Hum- 
ber Mouth (Bay of Fundy, Aug.). 

Presumably generally distributed over the eastern 
section of the continent from Canada to Texas. 

Genera 55-58: Pima to Catastia 

[Venational division B. Forewing with veins 4-5 separate or 
connate at base, smooth; hind wing with veins 4 and 5 anas- 
tomosed for about half their lengths, discocellular vein consider- 
ably extended at lower angle. Eighth abdominal segment of 
male with paired tufts. Antenna of male with shallow, spined 
sinus at base of shaft (also on Catastia a weak scale tuft). Labial 
palpus porrect or oblique, not grooved. Maxillary palpus of 
male squamous or subsquamous. Male genitalia with costa of 
harpe strongly sclerotized throughout its length and slightly 
produced at apex; gnathos terminating in a short, stout hook; 
transtilla incomplete or absent; penis armed with two stout 
cornuti {Pima, Inter jectio) or a single strong cornutus [Ambesa, 
Catastia). Female genitalia with bursa partially sclerotized and 
sometimes (Pima, Interjectio) granulate-scobinate but without 
definable signa; ductus bursae more or less sclerotized and con- 
siderably broadened at genital opening.] 

55. Genus Pima Hulst 

Pima Hulst, Ent. Amer., vol. 4, p. 114, 1888; Phycitidae of N. 

Amer., p. 164, 1890. (Type of genus: Pima fosterella Hulst.) 
Epischnia Authors (not Hubner) Ragonot, Ent. Amer., vol. 5, 

p. 115, 1889; Monograph (in part) pt. 1, p. 493, 1893. — 

Forbes, Cornell Mem. 68, p. 629, 1923.— McDunnough, 

Canadian Ent., vol. 67, p. 176, 1935. 

Tongue well developed. Antenna finely pubescent; 
on male with a very slight incurvation in base of shaft 
containing a row of minute, black, toothlike spines (6 
to 8). Labial palpus porrect (the second segment 
oblique, the third projected forward); extending at 
least twice the length of the head beyond it; second 
segment broadly (triangularly) scaled; thnd segment 
as long as second. Maxillary palpus minute but rather 
broadly scaled. Forewing smooth; 11 veins; vein 2 
from before lower outer angle of cell; 3 from the angle, 
closer to 4 than to 2 ; 4 and 5 separated at base ; 6 from 
well below upper angle of cell, straight; 8 and 9 stalked 
for one-half of less than half their lengths; 10 from the 
ceU, more or less approximate to the stalk of 8-9 ; male 
without costal fold. Hind wing with vein 2 from before 
lower outer angle of cell; vein 3 from 4 well beyond 2 
and considerably shorter than 2; 4 and 5 anastomosed 
just beyond 3 for nearly half their lengths; 7 and 8 
contiguous or closely approximate for a short distance 
from cell; cell slightly less than half the length of wing; 
discocellular vein curved, greatly extended at lower 
angle (running into 4 just beyond base of vein 3). 
Eighth abdominal segment of male with a pair of ven- 
trolateral hair tufts. 

Male genitalia with uncus hoodlike; its terminal mar- 
gin broadly rounded. Apical process of gnathos a short, 
stout hook. Transtilla absent. Harpe elongate, taper- 
ing to bluntly pointed apex; costa broadly and strongly 



102 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



sclerotized for the entire length of the harpe, its apex 
blunt, slightly produced and usually forked; sacciilus 
short and narrow, its inner surface covered with dense, 
spinelike hairs (probably the "strong spines" mentioned 
by Hulst in his original description of the genus) . Vin- 
culum a broad plate with weak lateral lobes. Aedeagus 
long, moderately stout, smooth, slightly curved towards 
base; penis armed with two stout cornuti. Vinciilum 
stout, tapering, considerably longer than greatest width. 

Female genitalia with bursa stout, scobinate-granu- 
late and usually with sclerotized patches or folds; ductus 
bin^ae long (much longer than bursa), flat (ribbonlike), 
sclerotized and granulate for its entire length, broadest 
towards genital opening; ductus seminalis from lobe of 
bursa near jimction of bursa an'd ductus bursae. Eighth- 
segment collar with middle of dorsal anterior margin 
slightly produced. 

Hulst erected the genus Pima for one species {foster- 
ella) which he later (1890) synonymized with alhijplagkb- 
teUa. Kagonot (1889) referred fosterella and the other 
congeneric American species to Epischnia Hiibner. The 
latter as defined by Eagonot (1893) is a composite of 
several disparate elements and none of the species occur- 
ring in the New World agrees with the type species of 
Epischnia (prodromella, Hiibner) . The latter has quite 
different genitalia (figs. 426, 884). Its male antenna 
has a deep sinus but the latter is without the row of 
toothlike spines characteristic of Pima; and the third 
segment of the labial palpus is much shorter (less than 
half the length of the second segment). There is also 
a heavy scale-and-hair tuft on the underside of the pro- 
thorax. This may be only a specific character (as simi- 
lar tuf tings are in some other phycitid genera) ; but the 
structm-e is entirely lacking on the New World species 
and on the European boisduvalieUa Gu6n6e, which is a 
tjrpical Pima on all characters. The venation of Pima 
is like that of Epischinia (fig. 26). 

The species here referred to PiwM (except graniteUa 
and parkerella) have a strikingly similar habitus ; but are 
individually variable in size and color, especially among 
examples of western species, several of which exhibit 
both pale and dark forms. The extent of the white 
costal streak on forewing is also individually variable 
and reliance on this and other color features has resulted 
in considerable confusion in the application of names. 
The most reliable specific characters are in the genitalia, 
especially those of the females. The chief male differ- 
ences are in the shape and size of the cornuti, the shape 
of the aneUus, and the configuration and width of the 
apex of the sclerotized costa of harpe. These differ- 
ences are trifling and also subject to some variation, es- 
pecially the notching at the apex of costa of harpe. 

The larvae feed in the flowers and seed pods of vari- 
ous Legumiinosae. From scattered reared examples in 
the National Collection the species do not seem to be 
confined to specific plants; but there have been no ex- 
tensive and systematic rearings, and these will be needed 
to clear up host relationships and to differentiate any 
possible food-plant races. 



Genus Pima, Species 209-216: P. boisduvalieUa 
to P. fulvirugella 

[Forewing with strongly contrasted wMte costal stripe.] 

209. Pima boisduvalieUa (Guenee), new combination 
Figures 299, 776 

Epischnia boisduvalieUa Guinea, Europaeorum Microlepidopter- 
orum index methodicus . . . , p. 81, 1845. — Ragonot, Ent. 
Monthly Mag., vol. 22, p. 23, 1885; Monograph, pt. 1, p. 
518, 1893 (part). — Lafaury, Ann. Soc. Ent. France, ser. 6, 
vol. 5, p. 398, 1885. — Spuler, Die Schmetterlinge Europas, 
vol. 2, p. 209, 1910. — Meyrick, Revised handbook of British 
Lepidoptera, p. 382, 1928. — McDunnough, Canadian Ent., 
vol. 67, p. 176, 1935 (part) ; Check list. No. 6251, 1939. 

Forewing pale fawn on lower half shading into muddy 
fawn along the lower border of the white costal stripe 
where it forms a more or less distinct dark band through 
the center of the wing from base to termen; no indica- 
tion of transverse Hnes; extreme costal edge blackish 
gray weakly peppered with white, this dark shade en- 
croaching on the white costal stripe beyond middle; 
white costal stripe extending to apex, its lower margin 
edged by a fine gray-black line; discal dots at end of cell 
minute, blackish, the upper one lying within the black- 
ish edge of the white stripe and frequently indistinguish- 
able, the lower one distinct but not conspicuous; some 
faint scattered black dotting and dusting on and below 
vein lb beyond base. Hind wing pale ocherous brown, 
on some specimens with a faint ocherous tint. Alar 
expanse, 19-25 mm. 

Male genitaUa with apex of costa of harpe slightly 
expended, concave or weakly notched, the upper angle 
of the notch rounded, the apical margin oblique. 
Cornuti both broadened for more than half their 
lengths; the broad part of the shorter thorn longitudi- 
nally grooved (fluted) ; the longer thorn but shghtly less 
than half the length of the aedeagus. 

Female genitaha with bursa narrowly heart shaped; a 
small rounded or oval sclerotized plate in bursa near its 
middle; bursa longitudinally wrinkled, the wrinklings 
weakly sclerotized; also a couple of broader, more 
tortuous, sclerotized folds, the one curving about junc- 
tion of bursa aind ductus bUrsae serrate along its edge; 
bm-sal granulations weak, scobinations in lobed area 
adjacent to junction of ductus btu-sae fine but rather 
dense; lower margin of ductus biu-sae at genital opening 
straight, not produced. 

Type locality: Switzerland (location of type un- 
known). 

Food plants: Ononis, Anihyllis, Lotus, Astragalus 
(European records). 

Distribution: Eubope. Canada: Manitoba, Aweme 
(May), Beulah (June, Aug.), Winnipeg; Saskatchewan, 
Regina (June, Aug.) ; Alberta, Lethbridge (Jime) . 

This European species is definitely estabUshed in 
North America, but most of the American references to 
boisduvalieUa apply to other native species. The only 
American examples of the true boisduvalieUa I have 
seen are some ten specimens from the Canadian localities 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



103 



cited above. Old World synonymns and doubtful 
American references are omitted from the foregoing 
literature citations. There can be no reasonable doubt 
of the correctness of the European synonymy oifarrella 
(Curtis) or lajauriella (Constant) (their references will 
be found in the Ragonot Monograph); but the status 
of the supposed Asiatic variety tabulella Staudinger is 
doubtful. Ragonot's reference of albocostalialis Hulst 
as a variety of boisduvaliella is obviously incorrect. 

In size boisduvaliella averages appreciably smaller 
than any other species of Pima. Occasional examples 
have an expanse equal to that of small specimens of some 
of our American species (24-25 mm.) ; but most speci- 
mens expand 24 mm. or less, while in the other species 
the average expanse is well over 25 mm. 

The male genitaha are similar in all essential charac- 
ters to those of albiplagiatella.; but the female genitaha, 
while of the same general habitus, differ in marked and 
apparently consistent details — the folds of the bursa are 
more weakly sclerotized, the bursa itself decidedly 
smaller, and the granulations and scobinations in bursa 
weaker. The contrasts are somewhat greater than 
shown in the figures. 

210. Pima albiplagiatella (Packard), new combination 
Figures 305, 777 

Myelois albiplagiatella Packard, Ann. Lye. Nat. Hist. New York, 

mit: vol. 10, p. 269, 1874. 

Epischnia boisduvaliella albiplagiatella (Packard) McDunnough, 

Canadian Ent., vol. 67, p. 70 (larva), p. 176 (part), 1935; 

Check list, No. 6251, 1939. 

Larger than boisduvaliella. The ground color on lower 
half of foremng very pale fawn, paler than that of 
boisduvaliella and containing httle of no dark scaling, 
the latter when present confined to a couple of faint 
gray dots on basal third of vein lb and some very 
sparse gray scaling along irmer margin near tornus; the 
contrasted dark band bordering lower margin of the 
white costal stripe brown rather than ruddy; a similar, 
weaker, narrower, brown shade along lower fold for 
most of its length; extreme costal edgepale brownish 
gray, this color encroaching on the white stripe beyond 
middle as in other species; lower discal dot distinguish- 
able but very faint. Hind wing whitish ocherou's. Alar 
expanse, 26-31 mm. 

Male genitalia like those of boisduvaliella. Female 
genitaha similar to those of boisduvaliella but con- 
sistently different in minor details. These differences 
were noted in the discussion of boisduvaliella and are 
shown in the figiires. 

Type locality: New Hampshire (type in MCZ). 

Food plant: Laihyrus maritima and presumably 
other Leguminosae. 

Distribution: United States: New Hampshire, 
Hampton (June, July). Canada: Nova Scotia, White 
Point Beach (Queens County, July). 

Typical albiplagiatella is probably generally distrib- 
uted in eastern Canada and northeastern United States. 
The foregoing records are from specimens whose 
genitaha I have been able to examine, the Nova Scotia 



record from a specimen suppUed by McDunnough. He 
has referred albiplagiatella as a race of boisduvaliella. It 
is indeed very close to the European species, but I 
beUeve the differences in female genitaha justify more 
than racial separation. 

211. Pima albiplagiatella occidentalis, new race 
FlGUBE 303 

A variety occurring in the Rocky Mountain and 
Pacific Coast States. Extremely variable in color and 
the amount of blackish dusting on forewing. Average 
specimens in Washington, Colorado, and New Mexico 
with ground color very pale fawn (cream white in some 
specimens) ; the dark border of the white costal stripe 
ranging from pale ocherous brown to dark gray-brown; 
two distinct black dots on basal third of vein lb, enclos- 
ing a white spot; the remainder of lb more or less 
streaked with black outwardly and a fijie peppering of 
black scales and white scales along inner margin near 
tornus; costal edge brownish or blackish gray, encroach- 
ing on and attenuating the white costal streak beyond 
middle, and sometimes blotting it out before apex. In 
southern California the paler specimens show more 
blackish dusting expecially along the outer veins, and 
sometimes a faint white subterminal line can be dis- 
tinguished; the darkest specimens are almost a uniform 
dark gray with the blackish gray dusting obliterating 
all markings except the white spot on vein lb and the 
contrasted white costal streak; between these two ex- 
tremes there is every intergrade in series from any given 
locality; lower discal spot small, but usually distinct, 
blackish. Hind wings ocherous white to dark smoky 
gray. Alar expanse, 24-31 mm. 

Male genitalia like those of typical albiplagiatella 
except that apex of sclerotized costa of harpe is some- 
what more swollen; in average specimens the outer 
edge of apex is slightly concaved; one example from 
Alamogordo, N. Mex., has the edge almost straight, 
but there is a gradual intergradation from this to forms 
with the apex as in our figure of albiplagiatella (fig. 
305c). The cornuti are like those of typical albiplagia- 
tella. The female genitalia are like those of eastern 
albiplagiatella. A series of 24 preparations from the 
various western localities exhibits no significant varia- 
tion from type and nothing approaching the bursa of 
boisduvaliella. 

Type locality: Pullman, Wash, (type in USNM, 
61342). 

Food plants: Astragalus, Lathyrus. 

Described from male type (C. V. Piper, collector, 
May 1905) and one female paratype (J. F. G. Clarke, 
May 30, 1924) from the type locality, and paratypes as 
follows: One male from above Golden, Colo. (H. G. 
Dyar No. 17468, May 29, 1901); one female, Beulah, 
Colo. (June 21, 1900, W. D. Kearfott Collection) ; 2 male 
and one female, Alamogordo, N. Mex. (May 15, 17, 
1929, reared by M. W. Talbot from Astragalus wootoni) ; 
one female, Phoenix, Ariz. (Apr., Kunze, collector); one 
male. Palm Springs, Calif.; one male and one female, 



104 



tnsriTED STATES NATIONAL MUSEUM BULLETUST 207 



Loma Linda, Calif. (May, June); one female, Clare- 
mont, Calif. (Baker, no date); and one female, San 
Diego, Calif. (H. G. Dyar, May 22, 1924). These are 
from a series of 85 specimens in the U. S. National 
Collection from the following localities : United States : 
Colorado, Beulah (June), Denver, Golden (May); New 
Mexico, Alamogordo, (May), Jemez Springs (Apr., 
May), Pecos; Arizona, Phoenix (Apr.). "Southern 
Arizona"; California, Claremont, Loma Linda (Mar., 
Apr., May, Jime), "Los Angeles County" (May), Mir- 
age Lake (San Bernardino County, Apr.) , Olancha (Apr., 
May), Palm Springs (Mar.), San Diego (Mar., May), 
"Shasta County," "Sierra Nevada"; Oregon, Baker 
(June) ; Washington, Copalis (a gray specimen with very 
dark hind wings, reared under Special Survey No. 
26286, Mar. 27, 1945, from Laihyrus sp.), Palouse Falls 
(May), Pullman (May, June, Jiily, Aug.), Walla Walla 
(June, July), Yakima (May). 

212. Pima fosterella Hulat 
Figures 300, 783 

Pima fosterella Hulst, Ent. Amer., vol. 4, p. 114, 1888. 

Pima albiplagiaiella Hulst (not Packard), Phycitidae of N. Anaer., 

p. 164, 1890. 
Epischnia albiplagiatella Ragonot (not Packard), Monograph, 

pt. 1, p. 518, 1893. 
Epischnia fulvirugella McDunnough (not Ragonot), Canadian 

Ent., vol. 70, p. 178, 1935. 
Epischnia fosterella (Hulst) McDunnough, Check list, No. 6252, 

1939. 

A large, pale species similar in general appearance to 
typical eastern albiplagiatella, but without any trace of 
a dark shade in lower fold ; the dark shade along lower 
border of the white stripe also paler, a light drab brown; 
a single small black spot on basal third of vein lb, but 
no white spot and seldom any further dark shading on 
lb, but more or less gray dusting along outer two-thirds 
of inner margin; white costal streak usually obliterated 
before apex, rarely reaching apex; lower discal dot 
usually well contrasted, but minute, blackish. Hind 
wing whitish ocherous or pale smoky fuscous. Alar 
expanse, 27-35 mm. 

Male genitalia with harpe somewhat longer in pro- 
portion to tegumen and uncas than in preceding species; 
sclerotized costa broadened and forked at apex, the 
prongs of the fork pointed. Cornuti spaced apart; 
neither one appreciably flattened or ribbed towards 
base; the longer slightly less than one-third the length 
of aedeagus. 

Female genitalia with little or no sclerotization of 
bursa except immediately about junction of bursa and 
ductus bursae. The extreme of sclerotization is shown 
in figure 783, from an Arizona female; the female type 
shows none except about the junction with ductus. 
Bursa finely scobinate over entire inner surface. Duc- 
tus bmsae produced at apex into a projecting shield, 
its apical margin variable, pointed to evenly rounded. 

Type locality: Colorado (type in AMNH, ex 
Kutgers). 

Food plant: Unknown. 

Disteibution: United States: Colorado, Baileys 



(Jidy), Denver (June), Durango (June), Golden (June), 
Gunnison County (near Altmont, July); Arizona, 
Williams; Utah, Eureka (May), Park City (July); Mon- 
tana. Miles City. Canada: Nordegg (June). 

A distinct species easily recognized by its female 
genitalia. The type (9) at Rutgers bears only a number 
label ("43") but is obviously a Colorado specimen and 
an authentic type. A male paratype matching it is 
in the National Museum. It and one other male and 
two females labeled only "Colo." AU Colorado exam- 
ples are larger specimens. The Nordegg specimen (c?) 
had been received from Dr. McDunnough and formed 
part of the series he had treated as Julvirugella in his 
1935 paper. It and a male from Eureka, Utah, show 
some black scaling on the outer veins. They are super- 
ficially very much like some specimens of the western 
race of albiplagiatella and except for their genitalia 
could easily be confused with them. 

213. Pima vividella (McDunnough), new combination 
FiGTJHES 302, 780 

Epischnia vividella McDunnough, Canadian Ent., vol. 67, p. 
179, 1935; Check list, No. 6256, 1939. 

Forewing salmon pink below costal white stripe, 
shading below and towards tornus into pinkish ocherous; 
some gray dusting along outer two-thirds of inner 
margin; a black dot, followed by an obscure white one, 
on vein lb at basal third ; white costal stripe attenuated 
by smoky costal scaling on its outer half and obliterated 
before apex. Hind wing pale sm.oky with a faint 
ocherous tint. Alar expanse, 27-30 mm. 

Male genitalia with apex of costa of harpe forked as 
in fosterella; but upper prong somewhat longer and 
sharper than the lower one. Cornuti closely approxi- 
mate; the longer one with flattened but not ribbed 
basal part, a trifle longer than one-third of the aedeagus. 
Female genitalia resemble those of fosterella except: 
Bursa proportionally smaller, with two pitted and 
sclerotized patches; sclerotization at junction of bursa 
and ductus bursae serrate along one edge. 

Type locality: Lethbridge, Alberta (type in Ca- 
nadian Nat. Coll.). 

Food plant: Unknown. 

Distribution: Manitoba, Aweme (June); Saskatche- 
wan, Saskatoon (June); Alberta, Lethbridge (June). 
Also recorded by McDunnough from Beulah, Manitoba 
(June) and Indian Head, Saskatchewan (July). 

A good species, close to but distinct trom. Josterella; 
easily identified by its male cornuti, female genitalia, 
and salmon-colored forewings. 

214. Pima albocostalialis (Hulst), new combination 
Figures 301, 778, 779 

Ephestia albocostalialis Hulst, Trans. Amer. Ent. Soc, vol. 13, 
p. 64, 1886. 

Epischnia albocostalis (Hulst), Phycitidae of N. Amer., p. 163, 
1890. (Emended spelling). 

Epischnia boisduvaliella albocostalis (Hulst) Ragonot, Mono- 
graph, pt. 1, p. 520, 1893. 

Epischnia albocostalialis (Hulst) McDunnough, Canadian Ent., 
vol. 67, p. 178, 1935; Check list, No. 6254, 1939. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITDSTAE 



105 



Forewing dark gi'ay shading into very dark gi-ayish 
or blackish brown towards the white costal streak; the 
latter ending just before apex; no spottings or other 
markings on the wing, even the lower discal dot lost in 
the dark suffusion of the wing. Hind wing whitish 
at base, shading into a smoky outer area. Alar expanse, 
26-34 mm. 

Male genitalia with apex of costa of harpe narrow, 
very slightly notched, reaching only to end of cucullus. 
Harpe itself shorter in proportion to length of tegumen 
and uncus than that of any preceding species except 
boisduvaliella. Cornuti spaced apart as in fosterella, 
the longer one about one-third the length of aedeagus. 

Female genitalia with two rather large, irregularly 
shaped, pitted and sclerotized patches in bursa, one at 
the terminal end, the other on the left side (viewed 
ventrally); bursa otherwise membranous except for a 
sclerotization about junction with ductus bursae; 
ductus bursae not produced at genital opening, its 
apical margin straight. 

Type locality: California (type in AMNH, ex 
Rutgers) . 

Food plant: Unknown. 

Distribution: California, Claremont, "En route Im- 
perial to Bishop" (May), Loma Linda (Mar.), River- 
side (Apr.), Shasta Retreat (Siskiyou County, July); 
Washington, Olympia (June), Rochester (June), Seattle, 
Wenatchee (May). McDunnough also records the spe- 
cies from Lethbridge, Alberta, Canada. His specimen 
is undoubtedly this species, but may represent one of 
the varieties that follow. 

The type in the Rutgers Collection lacks antennae 
and abdomen but matches otherwise the examples in 
the National Collection, so there can be no question of 
the application of Hulst's name. He emended its spell- 
ing to albocostalis in 1890, but gave no reason for doing 
so, hence we shall probably have to perpetuate the 
original barbarous spelling. 

215. Pima albocostalialis subcostella (Bagouot), new combination 

Epischnia subcostella Ragonot, N. Amer. Phycitidae, p. 10, 1887; 
Monograph, pt. 1, p. 520, 1893. — McDunnough, Check list, 
No. 6255, 1939. 

Forewing with white costal streak as in tj^jical albo- 
costalialis; below it a bordering band of blackish brown 
with a lighter brown shade below it as far as lower fold; 
these dark shades terminating beyond cell in what would 
be the position of the antemedial line if one were pres- 
ent, the outer margin of the dark shade outwardly 
angled at middle; remainder of wing pale, ashy gray 
with a dusting of blackish scales along outer two-thirds 
of inner margin and (in some specimens) blackish 
streaklets on the veins before termen; on basal third of 
vein lb a white dot preceded by a black shade. Hind 
wing white; smoky shading limited to a narrow line 
along termen and a very small area at apex; on the fe- 
males the smoky tints slightly more extended. Alar 
expanse, 25-27 mm. 

Genitalia, male and female, as in typical albocostal- 
ialis. 



Type locality: Utah (type in Paris Mus.). 

Food plant: Unknown. 

A male and three females from Eureka, Utah (May, 
June), a female from Bellevue, Washington County, 
Utah (May) and a male from Pyramid Lake, Nev., are 
before me. They match Ragonot's description and 
figure of subcostella in every detail. The name may 
represent nothing more than a color variety of albo- 
costalialis and if there were not an old name available 
for it, and one that until now was supposed to repre- 
sent a distinct species, I should not have named it. I 
am holding subcostella as a trinomial against the possi- 
bility that it may represent a valid local race. 

There are also before me what appear to be two 
other varieties with male genitalia identical to those of 
albocostalialis: 

Variety a: A large form with a reddish brown band 
bordering the white costal stripe; the red-brown color 
shading into ocherous fawn on lower and outer areas 
of the wing; lower discal dot present, black; no white 
spot or other appreciable marking on vein lb. Hind 
wing ocherous white with a pale broken Une along ter- 
men. Alar expanse, 34-35. Represented by three 
males from Manitou, Colorado (H. G. Dyar, coll. No. 
6062-6065, May 5, 1891). 

Variety b: A variable variety ranging from dark gray, 
suffused examples to a couple with the entire median 
area of the forewing a ruddy fawn color; the white spot 
is present on vein lb, but very faint on the darker 
specimens. Hind wings white to pale smoky fuscous. 
Alar expanse, 26-27 mm. 

The female genitalia differ rather markedly from 
those of subcostella or typical albocostalialis in that the 
sclerotized patches in bursa are situated on opposite 
sides of that organ. This arrangement is consistent for 
the females from both New Mexico and Arizona. 

Distribution: Colorado, Glenwood Springs (June); 
New Mexico, Fort Wingate (June) , Pecos (June) ; Ari- 
zona, Huachuca Mts., White Mts. (Aug.). 

Superficially this foi'm looks hke nothing but a color 
variant of subcostella. However, if the single female 
genitalic difference should hold through extended series 
it will need further designation. 

216. Pima fulvinigella (Ragonot), new combination 

Figure 304 

Epischnia fulvirugella Ragonot, N. Amer. Phycitidae, p. 10, 
1887; Monograph pt. 1, p. 521, 1893.— McDunnough, Check 
list. No. 6253, 1939. 

I have seen nothing that exactly matches Ragonot's 
description or figure (Monograph, pi. 16, fig. 43). A 
male before me from San Francisco has a similar, at- 
tenuated white subcostal streak and rather pronounced 
black lining on the veins from cell. Its genitalia, how- 
ever, do not match those of the type, and the moth 
itself is smaller (27 mm.). Ragonot gives the alar 
expanse of his type as 30 mm. 

Type locality: California (type in Paris Mus.). 

Food plant: Unkno\vn. 



106 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



Genus Pima, Species 217 and 218: P. granitella 
and P. parkerella 

pPorewing with white costal streak obscure or absent.] 

217. Pima granitella (Ragonot), new combination 
Figures 307, 781 

Epischnia granitella Ragonot, N. Amer. Phyoitidae, p. 9, 1887; 

Monograph, pt. 1, p. 523, 1893. — Hulst, Phycitidae of N. 

Amer., p. 162, 1890. — McDunnough, Check list. No. 6258, 

1939. 
Megasis piperella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 

110, 1904. 

Forewing gray finely powdered with white, the white 
dusting concentrated between the veins; the latter finely 
lined with blackish brown giving the wing (to the naked 
eye) a longitudinally dark-lined, over-all pale bluish 
gray appearance; on some specimens a trace of the usual 
costal white strip as a narrow white line along top of 
cell for a short distance from base; the lower blackish 
discal spot usually distinct but occasionally obliter- 
ated; on a few specimens the subterminal line slightly 
indicated by interruptions in the blackish streaks on the 
veins, but, otherwise, transverse lines absent. Hind 
wing pale brown, unicolorous, except for a narrow, very 
slightly darker line along termen. Alar expanse, 22- 
23 mm. 

Male genitalia with sclerotized costa of harpe broad- 
ened, sUghtly notched and produced at apex. Cornuti 
lying one before the other, of nearly equal length, not 
(or but very slightly) broadened and not flattened at 
base; the longer one slightly less than one-third the 
length of aedeagus. Female genitalia similar to those 
of albocostalialis except sclerotized patches in bursa, 
narrower, more elongate. 

Type locality: California (granitella, in Paris Mus.) ; 
Pullman, Wash, (piperella, in USNM). 

Food plant: Orotolaria. 

Distribution: Colorado, Glenwood Springs (Apr., 
May); Utah, Eureka (May, June), Kichfield (May), 
Stockton (May); New Mexico, Jemez Springs (Apr., 
May); Arizona, Dewey (Apr.); Nevada, Baker (May); 
California, Argus Mts. (May), Crows Landing (May), 
Los Angeles County (May); Washington, Pullman, 
Walla WaUa (May). 

A distinct species, easily recognized by its wing pat- 
tern. 

218. Pima parkerella (Schaue), new combination 
Figures 306, 782 

Epischnia parkerella Schaus, Proc. Ent. Soc. Washington, vol. 26, 
p. 196, 1924.— McDunnough, Check list, No. 6250, 1939. 

Forewing olive buff, costal and inner margins whitish 
more or less overlaid with a fine peppering of blackish 
scales; on some specimens a scattering of similar dark 
scales in the central area and near outer margin along 
some of the veins; no contrasted white costal strip; 
transverse lines well marked on most specimens, white, 
rather broad (absent on one example before me); the 
antimedial line outwardly obHque from costa to inner 
margin, nearly straight, at most with a slight bend at 



vein lb; subterminal line inwardly oblique from costa 
to inner margin, with a decided notch at lower fold, in- 
wardly margined by a faint dark shade; discal spots usu- 
ally obsolete, when distinguishable very faint. Hind 
wing very pale fuscous with a narrow dark Une along 
termen; on the paler examples the hind wing has a slight 
ocherous-gray tint. Alar expanse, 30-35 mm. 

Male genitalia somewhat stouter than those of pre- 
ceding species. Sclerotized costa of harpe not appre- 
ciably broadened at apex nor projecting beyond apex 
of cucullus; very sHghtly if any notched at apex. Cor- 
nuti lying close to each other, the apex of one projecting 
slightly beyond that of the other; about one-third as 
long as aedeagus. 

Female genitalia with bursa minutely scobinate, at 
left posterior angle developed as a protruding lobe with 
thickened membrane; ductus bursae developed into a 
projecting shield at genital opening. 

Type locality: Bozeman, Mont, (type in USNM). 

Food plant: "Locoweed" (Astragalus). 

Represented in the National Collection by nine ex- 
amples from the type locaUty (July). The species is 
easUy recognized by its forewing markings. It is the 
only Pima with anything approaching distinct trans- 
verse lines. 

56. Interjectio, new genus 

Type of genus: Ambesa columbiella McDunnough. 

Characters as in Pima except: Labial palpus extend- 
ing little more than the length of the head beyond it, 
third segment less than half as long as second; apical 
process of gnathos a broad, flanged plate terminating in 
a short slender hook; harpe short, hardly extending be- 
yond apical margin of uncus; vinculum little longer than 
greatest width; genitalia generally broader andmore 
chunky than those of Pima; ductus bursae of female 
short, Httle, if any, longer than bursa. 

This genus is intermediate between Pima and Ambesa. 
From the latter it differs chiefly in having at least two 
cornuti on the penis and the granulations of the ductus 
bursae uninterrupted for its entire length. 

219. Interjectio denticulella (Ragonot), new combination 

Figure 308 

Prisiophora denticulella Ragonot, N. Amer. Physicidae, p. 6, 1887. 
Ambesa lallatalis Authors not Hulst (in part) Ragonot, Ent. 

Amer., vol. 5, p. 115, 1889; Monograph, pt. 1, p. 241, 1893. 
Ambesa denticulella (Ragonot) McDunnough, Canadian Ent., 

vol. 67, p. 174, 1935; Check list, No. 6158, 1939. 

Forewing white dusted and marked with black, 
making the ground color (of fresh specimens) white 
with a very faint bluish tint; the dark markings strongly 
contrasted, consisting of streaks and spots chiefly indi- 
cating broken margins of the antemedial and postmedial 
lines; the antemedial line itself obscure except between 
lower fold and inner margin when it is an inwardly 
curved white line preceded by a black spot and followed 
by a thin black marginal line; above, indicated only by 
its outwardly oblique, broken outer margin, consisting 
of a short black line from costa and black streaklets on 



AMERICAN MOTHS OF THE SUBFAMILY PHyCITINAE 



107 



upper and lower veins of cell; subterminal line indicated 
by an oblique, deeply dentate, broken, black inner 
border; black discal dots at end of cell distinct, the 
upper minute, the lower a short but conspicuous streak; 
a row of black dots along termen; the blackish shading 
otherwise consists of very fine dusting between the 
veins, supplemented by some faint brownish shading 
in the folds. Hihd wing whitish brown, the veins very 
slightly dakened ; a pale brown line along termen. Alar 
expanse, 32-34 mm. 

Male genitalia with apical process of gnathos with 
the lateral angles of the flanged plate incurved and 
broadly rounded. Apex of cucullus not projecting 
beyond sclerotized costa of harpe. Cornuti two stout 
thorns, less than half as long as aedeagus; the latter 
short, stout, straight. 

Type locality: North America, but otherwise 
unspecified (type in Paris Mus.). 

Food plant: Unknown. 

Distribution: United States: Washington, Wenat- 
chee (May). Canada: British Columbia, Keremeos 
(June) . 

This species was removed by McDunnough (1935) 
from the synonymy of Ambesa lallatalis (Hulst) where 
it was placed originally by Ragonot (1889), presumably 
on the basis of specimens of supposed lallatalis sent him 
by Hulst. The latter species was misidentified by 
Hulst himself as weU as by later authors. 

220. Interjectio coltimbiella (McDunoough), new combination 

Figure 784 

Ambesa columbiella McDunnough, Canadian Ent., voL 67, p. 
175, 1935; Check list, No. 6159, 1939. 

Similar to denticulella but without the well-contrasted 
dark markings of that species; the black spot on inner 
margin, preceding the antemedial line, entirely lacking 
and the dark markings themselves more brown than 
blackish, except for the small, narrow black lower discal 
dot, an occasional black dot on vein lb near base, and 
some blackish dots along termen; the brown markings 
Umited to streakings on the veins and (where the trans- 
verse lines are distinguishable) to a narrow dark shade 
along inner margin of the subterminal line. Hind 
wings as in denticulella. Alar expanse, 23-34 mm. 

The species averages smaller than denticulella, espe- 
cially the females, which are even smaller than any of 
the males before me. 

Male genitalia similar to those of denticulella except 
cornuti distinctly more slender. Female genitalia 
(figured from a Pullman, Wash., specimen from a series 
associated with males of the same locaUty) with 
sclerotized ductus bursae projecting as a short shield 
at genital opening. 

Type locality: OUver, British Columbia (type in 
Canadian Nat. Coll.) 

Food plant: Unknown. 

Distribution: United States: Utah, Stockton 
(June); Washington, Pullman (May, June), Yakima 
(June). Canada: British Columbia, Oliver (June); 
Alberta, Lethbridge (July). 



221. Interjectio ruderella (Ragonot), new combination 

Figure 785 

Epischnia ruderella Ragonot, N. Amer. Phycitidae, p. 9, 1887; 
Monograph, pt. 1, p. 514, 1893.— Hulst, Phycitidae of N. 
Amer., p. 162, 1890.— McDunnough, Check list. No. 6249, 
1939. 

This species is known only from the female type, 
which, from Ragonot's description and figure, must be 
very close to some of the small females of columbiella, 
especially paler examples from Pullman, Wash., except 
that the lower discal spot is a more contrasted black 
streaklet. Alar expanse, 24 mm. 

The genitalia show a broader and stronger sclerotiza- 
tion of the ductus bursae at its junction with bursa than 
tjrpical columbiella; but this character is approached in 
Utah examples of the latter species. 

Type locality: "North America" [given as "without 
doubt from California" in the Ragonot Monograph] 
(type in Paris Mus.). 

Food plant: Unknown. 

222. Interjectio niviella (Hulst), new combination 
Figure 309 

Lipographis niviella Hulst, Ent. Amer., vol. 4, p. 117, 1888. 

Ambesa niviella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 241, 1893.— Hulst, Phycitidae 
of N. Amer., p. 141, 1890. — McDunnough, Check list, No. 
6157, 1939. 

Forewing chalk white; a large black patch bordering 
inner margin of antemedial line and extending from 
inner margin to top of cell; antemedial line faint but 
complete, oblique from costa to lower vein of cell, thence 
concave to inner margin, bordered outwardly by a black 
line, which is more or less broken into spots on upper 
half; black streaks and wedges on the outer veins indi- 
cating the deeply notched, otherwise obscure sub- 
terminal line; both discal spots conspicuous, black; a 
row of short black streaklets along outer margin; on 
some specimens smears of a faint, pale, ocherous brown 
tint towards apex; a fine brown line along termen. 
Alar expanse, 25-28 mm. 

Male genitalia with apical process of gnathos a broad- 
ly crescentiform plate with the lateral angles slightly 
produced ; the terminal hook short and slender. Harpe 
with apex of cucuUus projecting beyond the sclerotized 
costa. Cornuti a cluster of several short, stubby 
thorns. Aedeagus rather slender, sinuously curved. 

Type locality: Colorado (type in AMNH, ex- 
Rutgers). 

Food plant: Unknown. 

Distribution: United States: Colorado; Iowa, 
Sioux City. Canada: Manitoba, Aweme (July), Cart- 
wright, Winnipeg. 

The type is a female without abdomen. I have seen 
but one other female and it too was without abdomen, 
so the female genitalia could not be studied. These 
specimens matched the males in every detail of color 
and macidation. 



108 



■UNITED STATES NATIONAL MUSEUM BULLETIN 207 



57. Genus Ambesa Grote 

Anibesa Grote, N. Amer. Ent., vol. 1, p. 98, 1880.— Hulst, Phy- 
citidae of N. Amer., p. 141, 1890. — Ragonot, Monograph, 
pt. 1, p. 237, 1893. (Type of genus: Ambesa laetella Grote.) 

Tongue well developed. Antenna as in Pima and 
Inter jectio. Labial palpus oblique, extending above 
vertex; second segment roughly and rather broadly 
scaled; third segment not defected forward, about one- 
third as long as second, acuminate. Maxillary palpus 
minute, filiform. Forewing smooth; venation as in 
Pima. Hind wing with vein 3 from the angle of the 
cell (but separated from discocellular vein by a short 
spur), longer in proportion to 2 than in Pima, 4 and 5 
anastomosed for nearly half their lengths, 7 and 8 closely 
approximate for a short distance from cell; ceU one- 
third the length of wing; discocellular vein curved, ex- 
tended at lower angle but not so far as in Pima. Eighth 
abdominal segment of male simple or {laetella) with a 
pair of ventrolateral hair tufts. 

Male genitalia simUar to those of Interjectio except: 
Harpe longer in proportion to combined tegumen and 
imcus; its sclerotized costa with upper angle at apex 
produced into a sharp point; sacculus finely haired, not 
with coarse spinelike hairs of Pima or Interjectio. 
Anellus a simple plate without lateral projections. 
Penis armed with a single, long, stout cornutus; over 
half as long as aedeagus. Vinculum as long as greatest 
width. 

Female genitalia with bursa unsclerotized except at 
junction of bursa and ductus bursae; ductus bursae 
sclerotized except for narrow space at middle, broad- 
ened at genital opening. 

A North American genus close to both Pima and 
Interjectio, distinguished from both by its palpi, weakly 
haired sacculus, single strong cornutus, and the inter- 
rupted sclerotization of ductus bursae. The known 
species occur only in the western parts of the United 
States and Canada. 

223. Ambesa laetella Grote 
Figures 18, 310, 790 

Ambesa laetella Grote, N. Amer. Ent., vol. 1, p. 98, 1880. — Hulst, 
Phyoitidae of N. Amer., p. 141, 1890. — Ragonot, Mono- 
graph, pt. 1, p. 240, 1893. — McDunnough, Check list, No. 
6156, 1939. 

A brUliant, brightly colored species with clearly 
marked pattern; forewing markings a rich red-brown 
against a surrounding suffusion of ashy white; on mid- 
costa a broad, elongate red-bro^'m patch extending from 
outer margin of antemedial line to end of cell and from 
costal edge well into cell, shading into a blackish line on 
its lower margm, and bordered on its outer edge by an 
angled black discal mark formed of the fused discal 
spots; upper half of basal area, median area from the 
brown midcostal patch to vein lb, and most of the area 
beyond as far as subterminal line, ash white ; a brownish 
suffusion, paler than the costal patch, along inner mar- 
gin below vein lb, on some specimens intensified into a 
constrastingly darkened patch above middle of inner 



margin; the transverse lines weU separated, clearly 
marked ; antemedial line narrow, angled, white, bordered 
outwardly by a narrow blackish brown line; subterminal 
line sinuate, narrow, white, preceded and followed by 
blackish brown costal spots, the inner one continued as 
a brown inner bordering line, the outer expanding below 
into a pale brownish suffusion filling tornal area; sub- 
apical area dusted with white ; terminal dots fused into 
a black line along terminal edge. Hind wing pale 
smoky fuscous with a very faint yellowish tint. Alar 
expanse, 28-30 mm. 

Male genitalia with apical process of gnathos con- 
tinued below into a pair of narrow, converging, sclero- 
tized lobes. TranstUla absent. Harpe with cucullus 
narrow; sclerotized costa terminating xa a short, sharp 
point at apex; an erect clasper arising from below costa 
near base. Terminal margin of vinculum narrowly 
rounded. Eighth abdominal segment of male with 
paired tufts. 

Female genitalia with terminal, sclerotized portion 
of ductus bursae abruptly widened and transversely 
wrinkled toward genital opening. Eighth-segment col- 
lar narrowly sclerotized. 

Type locality : Colorado (type in BM) . 

Food plant : Unknown. 

Distribution : United States : Arizona, White Mts. 
(July, Aug.), Williams; Colorado, Fort Collins (Aug.), 
Glenwood Springs (Aug.); Montana, Bozeman (Aug.), 
Cut Bank (July); Utah, Provo, "South Utah" (July); 
Nevada, Verdi (June) ; California, Inyo County (June) , 
San Bernardino Mts. (July, Aug.), Sierra Nevada Mts.; 
Washington, Pullman (June, July, Aug.). Canada: 
Manitoba, Aweme (Aug.) ; Alberta, Calgary (July) . 

Aji easily recognized species and one of the most 
beautiful of the American Phycitidae. 

224. Ambesa walsinghami (Ragonot) 
Figures 312, 791, 792 

Pristophora walsinghami Ragonot, N. Amer. Phycitidae, p. 6, 

1887. 
Ambesa walsinghami (Ragonot) Hulst, Phycitidae of N. Amer., 

p. 142, 1890.— Ragonot, Monograph, pt. 1, p. 239, 1893.— 

McDunnough, Check list. No. 6153, 1939. 
Ambesa monodon Dyar, Ins. Insc. Menstr., vol. 1, p. 34, 1913. — 

McDunnough, Check list, No. 6154, 1939. (New synonymy.) 

Forewing blackish gray faintly tinted with purplish 
fuscous over the lower half of the wing; on costal half 
from base to antemedial line, white faintly streaked 
along the veins with black, the outer margin of the 
whitish area oblique from lower angle of cell to costa 
near beginning of subterminal line and, on some fresh 
specimens, bordered outwardly by a transverse blackish 
darkening of the ground color; some further ashy white 
dusting in the subapical area beyond the subterminal 
line; transverse lines well contrasted; the white ante- 
medial line distinct only from inner margin to cell, 
concave to lower fold thence inwardly oblique to the 
cell, beyond which it is lost in the white dusting above, 
its outer black margin begins as a black, oblique streak 
from costa and continues outwardly as a fine black line 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



109 



along upper edge of cell as far as discocellular vein, 
whence it loops backward along the lower vein of cell 
and thence along outer edge of the white line to inner 
margin; subterminal line, parallel with termen, sinuate, 
white, bordered inwardly by a black line and outwardly 
by a narrow dark shade, both borders enlarged at 
costa into contrasted black spots; discal dots not dis- 
tinguishable; along termen a row of more or less con- 
fluent black dots. Hind wings varying from smoky 
white to pale brownish, the veins faintly darkened. 
Alar expanse, 19-28 mm. 

Male genitalia with uncas broadly elongate, the side 
margins parallel. Apical process of gnathos without 
pendant lobes. Transtilla present, incomplete. Harpe 
with cucuUus moderately broad; apex of sclerotized 
costa produced into a long point at upper (outer) angle; 
no erect clasper. Eighth abdominal segment of male 
without tufts. 

Female genitalia with terminal sclerotized portion of 
ductus bursae gradually broadened to genital opening, 
its terminal margin deeply concaved. Eighth-segment 
collar broadly sclerotized, ventrally fused. 

Type localities: California {walsinghami, in Paris 
Mus.); Stockton, Utah (monodon, in USNM). 

Food plant: Prunus virginiana melanocarpa. 

Distribution: California, Cloverdale (June), Deer 
Park Spring (Lake Tahoe) , Plumas Coimty (July, Aug.) ; 
Utah, Bellevue (May), Eureka (June, July), Provo 
(July), Stockton (July); Washington, Kamiack Butte 
(May). 

Dyar's type of monodon is a small female whose 
genitalia (fig. 792) show minor differences from typical 
California specimens, but a series from Utah shows all 
intergradations between the extremes displayed in the 
figures. The food plant record is from a series reared 
by J. F. G. Clarke in 1934 at Kamiack Butte, Wash., 
which I then identified as mirabella Dyar. Their 
abdomens are distinctly gray and then* hind wings (es- 
pecially those of the females) are brownish. In exam- 
ples of typical walsinghami from the coastal region of 
California (Cloverdale) the hind wings are whitish and 
the abdomens gray or grayish ocheroUs. 

225. Ambesa walsinghami mirabella Dyar, new status 
Figures 313, 793 

Ambesa mirabella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 59, 1908. — Essig, Insects of western North America, p. 
709, 1926.— McDunnough, Check list, No. 6155, 1930. 

Not distinguishable from typical walsinghami in color 
or maculation of forewing. The hind wing of a dis- 
tinctly ocherous tint and the abdomen ocherous over 
the entire upper sm-face. Alar expanse, 25-28 mm. 

The male genitalia show only a trifling difference 
from those of tj^pical walsinghami in forking of apex of 
costa (fig. .313). Female genitalia with a shallower con- 
cavity in terminal margin of the projecting ductus bur- 
sae at genital opening. Figure 793 shows the extreme of 
reduction in the concavity. Other examples of mira- 
bella show intergrading approaches to the deep con- 
cavity of walsinghami. 



Type locality: San Diego, Calif, (type in USNM). 

Food plant: "Plum." 

Distribution: Calijomia, Atascadero (San Luis 
Obispo County, July), Camp Baldy (San Bernardino 
Mts., July), Cloudburst Canyon (Los Angeles County, 
July), Mount Lowe (July), Pasadena, Pine Valley (San 
Diego County), San Diego (July). According to Essig 
(1926) the larvae were taken in large numbers on prune 
trees at Hopeland, Calif. 

The name mirabella represents, at most, only a south- 
ern California race of walsinghami. 

226. Ambesa lallatalis (Hulst) 
Figures 311, 789 

Neophopleryx lallatalis Hulst, Trans. Amer. Ent. See, vol. 13, 

p. 161, 1886. 
Ambesa lallatalis (Hulst) Phycitidae of N. Amer., p. 142, 1890 

(part) . 

Forewing white heavily dusted with blackish giving 
the entire wing a pale ashy gray appearance ; markings 
as in walsinghami but much fainter; the transverse hnes 
obscure; antemedial line indicated by its broken outer 
blackish border which has the same outer loop over the 
cell so characteristic of walsinghami but much fainter 
and often interrupted; on better marked examples a 
whitish crescent on inner margin indicates the base of 
the normal antemedial line; beyond this, dark lines 
extend along vein lb and the edge of inner margin as 
far as base of subterminal line, defining a narrow oval 
patch along inner margin; subterminal line sinuate, de- 
fined by its black, dentate inner border, the latter inter- 
rupted by a rather broad pale shade extending along 
outer half of lower fold; the outer area (beyond sub- 
terminal line) and the costal area at base also paler than 
remainder of wing; a thin blackish line along outer mar- 
gin. Hind wing white to pale smoky brown. Eighth 
abdominal segment of male without tufts. Alar ex- 
panse, 26-30 mm. 

Male genitaUa with apical process of gnathos a simple, 
narrow, moderately long hook. Transtilla absent. 
Harpe without clasper; cucuUus moderately broad; 
sclerotized costa terminating in a very shortly project- 
ing point at apex. Anellus a narrow, elongate, plate. 
Cornutus almost as long as aedeagus. Vinculum with 
terminal margin broad. 

Female genitalia with terminal margin of ductus 
bursae at genital margin scobinate, convex, slightly 
notched in the middle (giving the projecting lower sur- 
face of the ductus a bilobed appearance). Eighth-seg- 
ment collar broadly sclerotized, wrinkled at lateral mar- 
gins, not ventrally fused. 

Type locality: Nevada (type in AMNH, ex Rut- 
gers). 

Food plant: Unknown. 

In addition to the female type in the Rutgers Collec- 
tion, I have seen only four other authentic specimens, a 
series of three males and one female from Bellevue, 
Washington County, Utah, in the National Collection, 
collected by G. P. Engelhardt, June 21, 1917. The 



no 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



genitalia of the Utah female agree in every detail with 
those of the type. 

The Ragonot and Dyar references to lallatalis have 
been omitted from the above s3Tiomy as they apply to 
other species. For comments on the misapplication of 
Hulst's name see under Interjectio denticvlella (p. 106) 
and Phobus hrucei (p. 138). 

58. Genus Catastia Hiibner 

Catastia Hiibner, Verzeiehniss bekannter Schmetterlinge, p. 372, 
1825. — Heinemann, Die Schmetterlinge Deutschlands und 
der Schweiz, Abt. 2, vol. 1, pt. 2, p. 164, 1865. — Ragonot, 
Monograph, pt. 1, p. 479, 1893. — Spuler, Die Schmetterlinge 
Europas, vol. 2, p. 210, 1910. — Hemming, Hiibner, vol. 2, p. 
168, 1937. — Janse, Journ. Ent. Soc. South Africa, vol 7, p. 
13, 1944 (description and figures). (Type of genus: Noctua 
marginea Schiffermiiller; figs. 314, 796.) 

Tongue weU developed. Antenna finely pubescent; 
on male with a shallow sinus in base of shaft containing 
a row of minute black teeth and a short, weak scale 
tuft. Labial palpus oblique, not extending above 
vertex; broadly scaled, the scales tightly appressed 
(except on incorruscella and actualis); third segment 
over one-third the length of second,* usually projected 
forward and partially concealed in the scaling of second 
segment, acuminate. Maxillary palpus squamous 
(broadly scaled). Forewing smooth; 11 veins; vein 2 
from before lower outer angle of cell; 3 from the angle, 
closer to 4 than to 2; 4 and 5 separated at base; 6 from 
below upper angle of cell, straight; 8 and 9 stalked for 
nearly half their lengths; 10 from the cell; male without 
costal fold. Hind wing as in Amhesa (except that in 
the European marginea vein 3 is slightly longer in 
proportion to 2; this species barely coming within our 
venational group B). Eighth abdominal segment of 
male with a pair of ventrolateral hair tufts. 

Male genitalic characters as in Amhesa except: 
Harpe not so long in proportion to combined length of 
tegumen and uncus; cucullus narrow on aU species. 
Transtilla incomplete, but its elements more strongly 
sclerotized. Vinculum as long or a trifle longer {mar- 
ginea) than greatest width, stout. 

Female genitaha with lower surface of ductus bm-sae 
towards genital opening unsclerotized or very weakly 
sclerotized (except in marginea); a pair of narrow, 
elongate plates on inner dorsal surface of ductus bursae 
at genital opening; otherwise as in Amhesa. 

This genus is very close to Amhesa, differing chiefly 
in its shorter labial palpus with deflected third segment; 
its squamous maxillary palpus; stronger sclerotization 
of the elements of transtilla; the presence of the two 
elongate sclerotized plates in the ductus bursae towards 
genital opening; and the slight scale tuft in the sinus 
of the male antennal shaft. 

The European type of the genus (figs. 314, 796) dif- 
fers from our American species in having the lower 
surface of the ductus bursae sclerotized and produced 

* Denuded example of palps of the type species {marginea) 
show the third segment half again as long as the figure (52b) in 
Janse's 1944 paper. 



at genital opening, the pointed projection from apex 
of costa a trifle longer, the vinculum narrowly roxmded 
at its extremity, and vein 3 of hind wing a trifle longer 
in relation to 2. These differences, however, are more 
specific than generic in character and do not seem to 
justify a separate generic designation for oiu- American 
species, despite the obvious likenesses in structure. 
The life history of none of the species is known. 

227. Catastia bistriatella (Hulst), new combination 

Figures 316, 797 

Pyla bistriatella Hulst, Canadian Ent., vol. 27, p. 54, 1895. — 
McDunnough, Check list. No. 6238, 1939. 

Head, thorax, and forewing black with a deep greenish 
blue iridescence; the forewing crossed by two rather 
broad white lines ; the antemedial line slightly oblique, 
nearly straight; the subterminal line set well back from 
termen, somewhat wavy. Hind wing a uniform, glossy 
black-brown. Labial palpus not reaching to vertex 
(shorter than the palpi of the other species of the 
genus) ; second segment broadly scaled, the scales flatly 
appressed; third segment shorter than that of any other 
species of the genus, almost completely hidden in the 
scaling of second segment. Alar expanse, 23-25 mm. 

Male genitalia with projecting spine at apex of 
sclerotized costa very short. Terminal margin of 
vinculum moderately broad, sHghtly produced at the 
lateral edges, very slightly convex. Female genitalia 
with ventral surface of ductus bursae at genital opening 
not sclerotized; bursa copulatrix with an irregular, 
lined, weakly sclerotized patch towards anterior end 
(probably an individual rather than a specific charac- 
ter). 

Type localitt: Yosemite, Calif, (type in AMNH, 
ex Rutgers). 

Food plant: Unknown. 

Distribution: California, Humphreys Basin (Fresno 
County, Aug.), Yosemite. 

A striking, easily recognized species. Hulst is in 
error in stating that the maxillary palpi are not scale 
tufted. They are squamous like those of the other 
cogeneric species but are difficult to see behind the 
heavily scaled labial palpi. The genitalia of the male 
type agree in every detail with th'ose of the Humphreys 
Basin male figured. 

228. Catastia incorruscella (Hulst), new combination 
Figures 315, 795 

Pyla incorruscella Hulst, Canadian Ent., vol. 27, p. 55, 1895. — 
McDunnough, Check list. No. 6240, 1939. 

Fore and hind wings a deep, blackish brown, some- 
what lustrous but without metallic iridescence; trans- 
verse lines similar to those of bistriatella, but thinner and 
a dull ocherous white. On thorax and palpi a scatter- 
ing of whitish scales; the scaling on second segment of 
labial palpus slightly roughened; third segment about 
the same proportional length as on marginea and actvr- 
alis, longer than that of bistriatella. Alar expanse, 20- 
22 mm. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITDSTAE 



111 



Male genitalia differ from those of bistriatella only in 
insignificant details. Female genitalia having bursa 
without sclerotized patch. However, a sclerotized 
patch similar to that shown for bistriatella (fig. 797) is 
present in the bursa of a female in the National Collec- 
tion from Slate Peak, Wash. On this specimen there 
is also a weak sclerotization of the ventral surface of the 
ductus bursae at genital opening. 

Type locality: Colorado (type in AMNH, ex Rut- 
gers). 

Food plant: Unknown. 

Distribution: Colorado; Arizona, White Mountains 
(June); Washington, Slate Peak (Okanogan County, 
Aug.). 

I have seen no Colorado examples except the male 
type, but the Arizona locality is represented in the 
National Collection by a series of males and females in 
excellent condition (collected by Grace M. and John L. 
Sperry at Colter's Ranch in the White Moimtains, June 
17-18, 1937). Their male genitalia are like those of the 
type. 

229. Catastia actualis (Hulst), new combination 
Figures 317, 794 

Nephopteryx actualis Hulst, Trans. Amer. Ent. Soc, vol. 13, p. 
161, 1886. 

Dioryclria actualis (Hulst), Phycitidae of N. Amer., p. 135, 
1890.— Ragonot, Monograph, pt. 1, p. 203, 1893.— McDun- 
nough. Check list, No. 6132, 1939. 

Monoptilota actualis (Hulst), TJ. S. Nat. Mus. Bull. 52, p. 421, 
1902 (this combination the result of an accidental misplace- 
ment) . 

Forewing with basal and outer areas ocherous, the 
basal area more or less dusted with ashy fuscous, espe- 
cially towards inner margin, the outer area with some 
blackish streaks on the veins; median area (between 
the transverse lines) ashy fuscous, the whitish dusting 
concentrated into a pale suffusion transversely across 
the wing from costa before subterminal line to or almost 
to inner margin at base of antemedial line; antemedial 
line white, oblique, notched below cell and more or less 
dentate above; bordered outwardly by a diffused black- 
ish smudge at costa and below cell by a blacldsh line, 
and preceded on inner margin by a black patch (except 
on California examples) ; a blackish spot precedes and 
one usually follows the sinuate subterminal white line 
on costa, the inner costal spot continuing as a blackish 
bordering line to inner margin; discal dots distinct, 
separated, black; a row of blacldsh dots along termen. 
Hind wing ocherous brown, darker brown on most fe- 
males ; a dark line along termen and some darkening of 
the veins. Alar expanse, 24-28 mm. 

Male genitalia differing only in insignificant details 
from those of bistriatella and incorruscella. 

Female genitaha with sclerotized wrinklings of bursa 
more extended than in other species of the genus; duc- 
tus bursae weakly granulate towards genital opening 
(differences of little or no significance) . 

Type locality: Colorado (type in AMNH, ex Rut- 
gers). 

Food plant: Unknown. 



Distribution: United States: Colorado, Denver 
(June), Florissant (July), Gunnison County (near Al- 
mont, June, July), Piatt Canyon (July), no specific lo- 
cahty (June) ; Nevada; Secret Pass (Elko County, June) ; 
Calijornia, Deer Park Springs (Lake Tahoe, July), Tuo- 
lumne Meadows (July); Washington, Easton, WaUa 
WaUa (June). Canada: Manitoba, Aweme (July); 
British Columbia, Blue Lake (west of Lytton, Aug.) . 

The foregoing description was drawn from typical 
Colorado examples. The British Columbia and Wash- 
ington specimens before me are much darker, the ocher- 
ous coloration of forewing replaced by deep brown and 
the general color of the wing a suffused blackish fuscous 
with only the white transverse lines and some whitish 
dusting in the median area contrasted; the hind wings 
deep brown to blackish brown with no ocherous tinting. 
This is merely a color form, not a race, and probably 
represents nothing more than individual response to a 
moist condition. The Nevada record, cited above, is 
from a single male in the Canadian National Collection 
(Grace H. and John L. Sperry, collectors). It also rep- 
resents a divergent color form with very dark brown 
hind wing, a dark ground color on forewing and strongly 
marked, white, transverse lines. The tj^e in the Rut- 
gers Collection is a male. Its genitalia agree in every 
detail with those of our pale and dark forms. 



Genera 59-64: Immyrla to Quasisalebria 

[Venational division B. Veins 4 and 5 of forewing slightly 
separated at base, in Oreana connate or, rarely, very shortly 
stalked; vein 6 straight. Hind wing with veins 7-8 approximate 
or very shortly anastomosed beyond cell. Antenna of male with 
sinus and scale tuft at base of shaft. Labial palpus erect or 
oblique, smooth scaled; on male second segment grooved to hold 
maxillary palpus. Maxillary palpus of male in the form of an 
aigrette. Male genitalia with transtilla absent, or incomplete 
and with its elements rudimentary; penis normally armed with 
a single strong cornutus (except Oreana which has several cornuti 
and Quasisalebria which has none). Female genitalia with bursa 
more or less finely and densely spined or scobinate.] 

59. Genus Immyrla Dyar 

Immyrla Dyar, Journ. New York Ent. Soc, vol. 14, p. 108, 
1906.— Forbes, Cornell Mem. 68, p. 627, 1923. (Type of 
genus: Immyrla nigrovittella Dyar.) 

Tongue well developed. Antenna pubescent; a sinus 
and large-scale tuft on base of shaft of male. Labial 
palpus erect; reaching above vertex; smoothly scaled; 
second segment on male grooved to hold the maxillary 
palpus, appressed to face; third segment minute Gess 
than one-fifth the length of second and hidden in the 
scaling of the latter on male, a trifle longer and partially 
exposed on female), acuminate. Maxillary palpus of 
male in the form of an aigrette; of female squamous. 
Forewing with subbasal scale ridge; 11 veins; vein 2 
from before lower outer angle of cell; 3 from the angle, 
closer to 4 than to 2; 4 and 5 approximate for a short 
distance from their bases; 6 from below upper angle of 
cell, straight; 8 and 9 stalked for over half their lengths; 
10 from the cell, approximate for a short distance to the 



112 



tnsriTED STATES NATIONAL MUSEUM BULLETIN 2 07 



stalk of 8-9 ; male without costal fold. Hind wing with 
veiQ 2 from before lower outer angle of cell; 3 from 4 well 
beyond 2, in some specimens (fig. 19) connected with 
discoceUular by a short spur before its separation from 
4; 4 and 5 anastomosed for less than half their lengths; 
7 and 8 anastomosed for a short distance beyond cell; 
cell less than half the length of wing; discoceUular vein 
cin-ved and considerably extended at lower angle. 
Eighth abdominal segment of male with compound 
ventral scale tufts. 

Male genitalia as in Oatastia except transtiUa com- 
pletely absent, a fine brush of long hairs arising from 
inner surface of harpe along lower edge of basal half of 
the sclerotized costa (as in Meroptera). 

Female genitalia with bursa copulatrix finely and 
densely spiued; ductus bursae flattened, gramilated, 
inbent at middle; not longer than bursa, its lateral 
margins strongly and broadly sclerotized towards 
genital opening, opening deeply concave; ductus semi- 
nalis from a thickened (but not sclerotized) lobe of 
bursa, near junction of bursa and ductus bursae. 

This and the five genera following form a group 
closely allied to Salebiia Hiibner. All have grooved 
and erect or oblique male labial palpi, the male maxil- 
lary palpus in the form of an aigrette, a scale tuft in 
sinus on base of shaft of male antenna, no transtilla or 
only the greatly reduced rudiments of one, and (except 
for Oreana and Quasisalehria) a single, long, strong 
cornutus on penis. Immyrla is distinguished from all 
nearly related genera by the raised scale ridge on fore- 
wing. Ortholepis also has this character, but the costal 
sclerotization of its harpe is much weaker, and it has a 
complete transtilla. 

230. ImmyTla nigrovittella D^ar 

FiGTTEBS 19, 318, 798 

Immyrla nigrovittella Dyar, Journ. New York Ent. Soc, vol. 14, 
p. 109, 1906.— Forbes, Cornell Mem. 68, p. 627, 192.3.— 
McDunnough, Check list, No. 6187, 1939. 

Forewing dark gray, the basal area darker than re- 
mainder of wing; median and outer areas a paler 
brownish gi'ay with a faintly darker shade preceding 
the subterminal line, a very sparse dusting of whitish 
scales on the paler areas; antemedial line faiut, narrow, 
oblique and more or less curved, dull white, followed 
by a narrow, blackish brown border and preceded by a 
conspicuous black ridge of raised scales extending from 
inner margin to top of cell; subterminal line obscure, 
sinuate, pale, without dark bordering lines ; discal spots 
faint, blackish and confluent, forming a curved line 
along discoceUular vein. Hind wing pale fuscous, the 
veins very slightly darkened. Both fore and hind 
wings have a rather slick, glossy finish. Alar expanse, 
20-21 mm. 

Genitalia: As given for the genus. 

Type locality: Pittsburgh, Pa. (type in USNM). 

Food plant: Hickory. 

Distribtitign: Pennsylvania, New Brighton (June), 
Pittsburgh (May); New York, lUion (June). 



The only species so far discovered referable to the 
genus. The food plant record is from a female without 
a locality label in the National Collection, reared from a 
pupa. The label reads simply "102, Hickory, pupated 
VII — 17." The specimen also bears an identification 
label in Dyar's handwriting. 

60. Genus Oreana Hulst 

Oreana Hulst, Ent. Amer., vol. 4, p. 115, 1888. (Type of genus: 
Dioryctria unicolorella Hulst.) 

Characters as in Immyrla except: Forewing smooth; 
veins 4 and 5 connate (rarely very shortly stalked) ; vein 
10 from the stalk of 8-9 a short distance from ceU. 
Hind wing with 4 and 5 stalked for more than half their 
lengths. 

Male genitalia with numerous strong slender cornuti 
on penis. 

Female genitalia with bursa copulatrix strongly 
sclerotized in the lobed area giving off the ductus semi- 
nalis; sclerotized lateral margins of ductus bursae not 
produced at genital opening and ventral margin of the 
opening not appreciably concave (these differences in 
the ductus bursae probably only of specific significance). 

Oreana sank into the synonymy of Meroptera when 
Ragonot (1889) referred its type species (unicolorella) to 
the latter genus, but it must be restored, as unicolorella 
is not a Meroptera on genitalic characters. Its chain of 
numerous, strong cornuti distinguish it from any species 
in Meroptera or the American genera closely aUied to 
Salebria. Oreanxi, on most characters, seems nearest to 
Immyrla, from which it is at once distinguished by its 
smooth forewing. It contains but the one described 
American species. 

231. Oreana unicolorella (Hulst) 

FiGTJBEs 319, 788 

Dioryctria unicolorella Hulst, Ent. Amer., vol. 3, p. 136, 1887. 

Oreana unicolorella (Hulst), Ent. Amer., vol. 4, p. 115, 1888. 

Meroptera unicolorella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 
115, 1889; Monograph, pt. 1, p. 315, 1893.— Hulst, Phy- 
citidae of N. Amer., p. 149, 1890. — Forbes, Cornell Mem. 
68, p. 624, 1923. 

Myelois leucophaeella Hulst, Canadian Ent., vol. 24, p. 60, 1892. 

Meroptera leucophaeella (Hulst), Barnes and McDunnough, Con- 
tributions, vol. 3, p. 194, 1916. 

Meroptera nebulella McDunnough (not Riley), Check list, No. 
6185, 1939. 

Forewing mouse gray, the basal area slightly paler 
than remainder of wing; transverse lines grayish white, 
obscure; antemedial line obUque, notched below ceU, 
bordered outwardly from costa for a short distance by an 
obscure blackish shade; subterminal line sinuate, with- 
out appreciable dark borders; discal dots faint, brown, 
usuaUy separated but sometimes confluent; on underside 
of male forewing a streak of blackish sex scaling along 
basal third of costa. Hind wing pale smoky brown, the 
veins slightly darkened. Alar expanse, 18-22 mm. 

Male genitalia with harpe, tegumen, and vinculiun 
similar to those of Immyrla nigrovittella. Uncus some- 
what smaUer in proportion; in natm-al position inclined 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



113 



downward, but when flattened out it shows an appre- 
ciable constriction at middle. Anellus a simple shield- 
shaped plate. Female genitalia as given for the genus ; 
spining of bursa not so dense or continuous as in 
nigrovittella. 

Type localities: Washington, D. C. [sic] {unicol- 
orella, in AMNH, ex Rutgers) ; Iowa (leucophaeella, in 
AMNH, ex Rutgers). 

Food plant: Apple (record from female in National 
Collection labeled "bred from apple, emerged 25 — V — 
1904, Ottawa, J. Fletcher"). 

Distribution: United States: Maine, Augusta 
(Jime); New Hampshire, Dublin; Connecticut, East 
River (July) ; New Jersey, Montclair (July) ; Pennsyl- 
vania, New iSrighton (May, June, July, Aug.), Oak 
Station (June), Pittsburgh (Jime, July) ; Iowa; Colorado; 
Oregon, Milton (July); Washington, Pullman (Jime). 
Canada: Ontario, Ottawa (May), Trenton (July); Que- 
bec, Montreal (July), St. Hilaire (July); Nova Scotia, 
Cape Breton Isl. (July). 

There is some mislabeling of the types or, what is more 
likely, errors in the citation of type localities by Hulst. 
In his original description of unicolorella he cites "Wash- 
ington, D. C." and in his 1890 paper gives "Canada." 
His type is a male (without abdomen) from Iowa, 
labeled: "H. S. Sanders, June 13, 1886." The type of 
leucophaeella is a female with typical genitalia, labeled 
"Colo., Gillette." The type locality citation of "Iowa" 
in the original description can be written off as another 
Hulst lapsus, although the species occurs there and is 
represented by several examples in the National Collec- 
tion. Barnes and McDunnough (1916) first put leuco- 
phaeella into synonymy with unicolorella, and there does 
not seem to be any reason to doubt the correctness of 
that procedure. The two specimens at Rutgers labeled 
"type" by Hulst certainly represent one and the same 
species. 

61. Olybria, new genus 

Type of genus: Myelois aliculella Hulst. 

Tongue well developed. Antenna finely and densely 
pubescent; on male a sinus and scale tuft in shaft at 
base. Labial palpus oblique, reaching well above vertex; 
second segment of male grooved to hold the maxillary 
palpus ; third segment short, about one-third the length 
of second, acuminate, partially concealed by scaling of 
second segment. Maxillary palpus of male in the form 
of an aigrette; of female small, squamous. Forewing 
smooth; 11 veins; vein 2 from before but near lower 
outer angle of cell; 2,3, and 4 equidistant at base; 4 and 
5 shortly separated at base and thence approximate 
(parallel) for a very short distance; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for about half 
their lengths; 10 from the cell, shortly separated from 
the stalk of 8-9 at base; male without costal fold. Hiad 
wing with vein 2 from weU before lower outer angle of 
cell; 3 from the angle, connate with the stalk of 4-5; 
4 and 5 stalked for half their lengths; 7 and 8 closely 
approximate for half their lengths beyond cell; cell 



slightly less than half the length of wing; discocellular 
vein curved, considerably extended at lower angle. 
Eighth abdominal segment of male with a pair of thin, 
weak, ventrolateral hair tufts. 

Male genitalia with uncus small, hoodlike, projected 
downward (at right angle to tegumen) . Apical process 
of gnathos a short, stout, curved hook. Tegumen with 
a pair of straight, strongly sclerotized arms projecting 
backward from its lower, posterior angles. Transtilla 
absent. Harpe with costa strongly sclerotized through- 
out its length and produced at apex into a sharp spine; 
cucullus narrowly elongate, tapering to pointed apex. 
AneUus a broadly U-shaped band, supplemented (in 
furciferella) by some sclerotization of the membranous 
tube surrounding the aedeagus. Aedeagus rather long 
and stout; penis armed with a single long, strongly 
sclerotized, rather slender cornutus. Vinculum stout, 
longer than gi-eatest width, narrowly truncate at 
terminal margin; the latter very slightly concave. 

Female genitalia with bursa smoothly sclerotized over 
most of dorsal surface, its ventral surface finely sco- 
binate with contorted, wrinkled and spined bands more 
or less encircling the bursa at junction of bursa and 
ductus bursae; ductus bursae at least as long as bursa, 
rather broad, flattened (ribbonlike) and waved (twice 
bent), sclerotized throughout, at genital opening the 
sclerotization forming a stout, squarish ventral plate; 
ductus seminalis from a lobe of bursa near junction of 
bursa and ductus bursae. 

This genus is distinguished from its nearest allies of 
the Salebria complex by the projecting arms from 
tegumen of the male genitalia, the ribbonlike, sclerotized 
ductus bursae and heavy, squarish genital plate of the 
female, and the simple, paired tufts of the eighth abdom- 
inal segment of the male. This last character is 
shared by the genus Salebriacus, which separates from 
Olybria on other difi^erences of genitalia and venation. 

Two North American species, referred from Salebria 
(of authors) , represent the only known components of 
the genus. 

232. Olybria aliculella (Hulst), new combination 
Figures 320, 786 

Myelois aliculella Hulst, Ent. Amer., vol. 3, p. 135, 1887. 

Salebria oberthuriella Ragonot, N. Amer. Phycitidae, p. 9, 1887. 

Salebria aliculella (Hulst) Ragonot, Eut. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 367, 1893.— Hulst, Phycitidae 
of N. Amer., p. 154, 1890. — Barnes and McDunnough, 
Contributions, vol. 3, p. 197, 1916. — McDunnough, Check 
list. No. 6217, 1939. 

Forewing white dusted with black, making the general 
color ashy gray, the black dusting concentrated on 
extreme base and in short streaklets on the lower veins 
at termen; antemedial line narrow, oblique, slightly 
notched at vein lb, white, bordered outwardly by a 
black line which begins on costa as a conspicuous, 
triangular, black spot; on inner margin, preceding the 
antemedial line, a large orange spot; subterminal line 
sinuate, narrow, white, bordered inwardly by a narrow 
black line and outwardly by a broad orange band which 



114 



tnsriTED STATES NATIONAL MUSEUM BULLETIN 207 



ends in a black spot at costa; along lower fold between 
the transverse lines fresh specimens show a faint shading 
of olivaceous ocherous; a similar shade often along 
discocellular vein; completely surrounding the dis- 
cocellidar vein a large black ring (obicular). Hind 
wing translucent white with a faint ocherous tint 
especially towards outer margin and anal angle; the 
veins very faintly darkened and a dark line along 
termen. Alar expense, 19-22 mm. 

Male genitalia showing only comparative differences 
to distinguish them from those of Jurcyferella. These 
are shown in the figures. Female genitalia with a row 
of stiff, flattened setae along lower, posterior margin of 
the eighth-segment collar. 

Type locality: Arizona {alicvlella, in AMNH, ex 
Rutgers; oherikuriella, in Paris Mus.). 

Food plant: Ceanothus ? (Hulst, 1890). 

Distribution: Arizona, Kingman (Oct.), White 
Mts. (Aug.), Wilgus, Williams, state locality only 
(June); New Mexico, Albuquerque, Fort Wingate 
(June), Jemez Springs (June, July); Texas, Big Bend 
region (May). 

Easily identified by the orange spot preceding the 
antemedial line and the large black obicular mark on 
the disc of the forewing. 

233. Otybria forciferella (Dyar), new combination 
Figures 321, 787 

Salebria furciferella Dyar, Joum. New York Ent. Soc, vol. 12, 
p. 106, 1903. — Barnes and McDunnough, Contributions, 
vol. 3, p. 197, 1916.— McDunnough, Check list. No. 6216, 
1939. 

Similar to aliculella except: Forewing more strongly 
dusted with black, making ground color a decided ash 
gray; the obicular spot on disk absent, replaced by a 
couple of black streaks on upper and lower veins at end 
of cell, forking from a black streak extending along the 
upper vein of cell from the black costal dash bordering 
the antemedial white line; the subbasal orange patch 
on inner margin preceding the antemedial line and the 
orange shade following the subterminal Une reduced 
and, on some specimens, obscured by black scaling. 
Hind wing as in aliculella. Alar expanse, 21-23 mm. 

Male genitaha similar to those of aliculella except 
projecting arms of tegumen stouter; aedeagus and 
cornutus longer. Female genitalia with a fringe of 
fine hairlike setae along the lower posterior margin of 
eighth-segment collar. 

Type locality: Ashfork, Ariz, (type in USNM). 

Food plant: Unknown. 

Distribution : Arizona, Ashfork and Fresco tt (June, 
type, (f , and 2 paratypes, 9) , also 2 females with only 
the state locality, from the Femald and Brooklyn 
Museum Collections and originally made cotypes of 
aliculella Hulst. 

The species is easily recognized by the black streak 
along the upper vein of cell and is distinct from alicu- 
lella though obviously very close to it. 



62. Salebriacus, new genus 

Type of genus : Nephopteryx odiosella Hulst. 

Characters of Olybria except: Male antenna with 
scale tuft in shallow sinus on base of shaft but greatly 
reduced, a mere vestige. Forewing with vein 2 sUghtly 
further from 3 at base than 3 is from 4; 8 and 9 stalked 
for considerably more than half their lengths. Paired 
tufts on eighth abdominal segment of male very weak, 
mere vestiges. 

Male genitalia with uncus hoodlike, elongate (longer 
than wide) and not projected downward. Apical 
hooked process of gnathos stout, long, reaching nearly 
as far as apex of uncus. Tegumen simple. Transtilla 
present but incomplete and its elements much reduced. 
Harpe with apex of sclerotized costa produced as a 
stout, free spine before apex of cucullus. Anellus a 
simple shield. Entire genitalia more robust and 
proportionally shorter than those of Olybria or the genus 
Salehriaria, which follows. 

Female genitalia with bursa very broad in proportion 
to its length, not sclerotized, nearly half the inner sur- 
face covered with a dense, spinose mat; ductus bursae 
semitubular, broad and short, partially sclerotized (the 
sclerotization interrupted near middle of the ductus); 
ductus seminalis from near anterior end of bursa. 
Eighth-segment collar narrow ventraUy and laterally, 
dorsally produced as a spatulate apron. 

This genus is another restriction from Salebria of 
authors. It is distinguished chiefly by the weak tuft 
of the male antenna, the squat, stout, male genitalia; 
the robust, hooked projection of gnathos, the eighth- 
segment collar of the female, and the place of departure 
from bursa copulatrix of the ductus seminalis. 

It contains but one North American species. 

234. Salebriacus odiosellus (Hulst), new combination 
Figures 322, 799 

Nephopteryx odiosella Hulst, Ent. Amer., vol, 3, p. 132, 1887. 
Salebria odiosella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 

1889; Monograph, pt. 1, p. 366, 1893.— Hulst, Phycitidae 

of N. Amer., p. 155, 1890. — Barnes and McDunnough, 

Contributions, vol. 3, p. 197, 1916. — McDunnough, Check 

list. No. 6219, 1939. 
Salebria bakerella Dyar, Journ. New York Ent. Soc, vol. 12, p. 

105, 1904.— McDunnough, Check list. No. 6220, 1939. 

(New synonymy.) 
Salebria yumaella Dyar, Proc. Ent. Soc. Washington, vol. 7, 

p. 35, 1905.— McDunnough, Check list No. 6218, 1939. 

(New synonymy.) 

Forewing white dusted with black, making the 
ground color ashy gray, palest over the median area; 
antemedial Une narrow, obhque from costa to lower 
margin of cell, thence slightly incurved to lower margin, 
white, margined outwardly on upper half by a rather 
broad black band and inwardly on lower margin by a 
more or less expanded black patch; subterminal line 
sinuate, narrow, white, bordered inwardly and, to a 
lesser extent, outwardly by blackish lines which expand 
into distinct black dashes at costa; discal dots at end 
of cell, separated, usually distinct, blackish; on most 



AMERICAN MOTHS OF THE SXIBFAMILY PHTCITINAE 



115 



specimens a diffuse oblique shade from upper part of 
subterminal line obliquely across wing toward inner 
margin, and on a few of the more contrastingly marked 
specimens a faint ocherous shading along the lower 
fold; terminal dots along outer margin more or less 
confluent, obscure on many specimens. Hind wing 
white, translucent; faintly shaded with pale brown at 
apex and along outer margin for a short distance from 
apex. Alar expanse, 19-26 mm. 

Male genitalia with vinculum rather abruptly nar- 
rowed from middle to a truncate terminal margin. 
Female genitalia with sclerotized portion of ductus 
bursae at genital opening produced and broadened, its 
terminal margin and lateral angles concave. 

Ttpe localities: "Colorado" [sic] (odiosellus, in 
AMNH, ex Rutgers) . Ormsby County, Nevada {haker- 
ella, in USNAI); Yuma County, Ariz, {yumaella, in 
USNM). 

Food plant: Unknown. 

Distribution: Texas, Blanco County [?]; Arizona, 
Mohave County (July, Sept.), Yavapai County, Yuma 
County; Colorado; Utah, Bellevue (May, June, July); 
Nevada, Ormsby County (July), Pyramid Lake, Reno; 
California, Jacumba (May), Mexican Wells (Clarke 
Mts., Sept.), Morongo VaUey (May). 

The types of odiosellus, bakerella, and yumaella are all 
males with identical genitaha. That of yumaella is a 
small, somewhat suiffused specimen with the black 
markings less strongly contrasted than those of typical 
Nevada examples. In our series there is a complete 
intergradation between the extreme forms. 

Barnes and McDunnough (1916) called attention to 
the confused citations of the type locahty of odiosellus. 
Hulst gives Colorado in his original description, but in 
his 1893 paper cites "Texas." The type at Rutgers is 
labeled "Blanco Co., Texas." I suspect that this is a 
mislabeUng, for Colorado seems a more likely locality 
than central Texas. In my paper on the cactus-feeding 
Phycitinae (Proc. U. S. Nat. Mus., vol. 86, p. 389, 1939) 

1 misapplied the Hulst name to a species described by 
Dyar as Ozamia clarejacta. This error is discussed 
further under the treatment of clarejacta (p. 258). 

63. Salebriaria, new genus 

Type of genus : Salebria ademptandella Dyar. 

Tongue well developed. Male antenna pubescent with 
strong scale tuft in sinus at base of shaft. Labial palpus 
obliquely upturned, reaching above vertex, smoothly 
scaled; second segment on male grooved to hold the 
maxillary palpus, appressed to face; third segment mi- 
nute and hidden in scaling of second on male, somewhat 
longer and partially exposed on female, acuminate. 
MaxiUary palpus of male in the form of an aigrette; of 
female subsquamous. Forewing smooth; 11 veins; vein 

2 from before but near lower outer angle of cell; 3 
usually somewhat nearer to 4 than to 2, sttaietimes equi- 
distant from them; 4 and 5 shortly separated at base, 
rarely (in smaller specimens) closely approximate; 6 
from below upper angle of cell, straight; 8 and 9 stalked 



for well over half their lengths (except in jructetella) ; 
male without costal fold; on underside of male wing a 
streak of appressed, black sex-scaling along basal third 
of costa. Hind wing with vein 2 from before, but 
rather near lower outer angle of cell; 3 from the angle, 
connate with the stalk of 4-5 ; 4 and 5 stalked for approx- 
imately half their lengths (for sUghtly over half in 
jructetella) ; 7 and 8 approximate beyond cell (except in 
pumilella and jructetella where they are contiguous or 
weakly anastomosed for a short distance) ; cell slightly 
less than half the length of wing. Eighth abdominal 
segment of male with compound ventral scale tufts. 

Male genitaha with uncus sub triangulate (apex rather 
broadly rounded). Apical process of gnathos a short 
hook, slender (except on jructetella). Transtilla absent. 
Harpe with apex of sclerotized costa produced as a short 
free spine before apex of cucullus. Anellus a semitubu- 
lar shield. Penis armed with a single, strongly sclero- 
tized, moderately long cornutus. Vinculum stout, sub- 
triangulate, slightly longer than greatest width. 

Female genitalia with much of inner surface of bursa 
covered with a dense matting of fine spines (especially 
towards distal end); bursa also sometimes partially 
sclerotized (pumilella, jructetella) ; ductus bursae shorter 
than biu-sa, strongly sclerotized (at least towards genital 
opening), partially flattened (ribbonlike in pumilella); 
ductus seminalis from lobe of bursa adjacent to the 
junction of bursa and ductus bursae. 

This genus is a further restriction from Salebria. It 
is a compact group of very closely related species; in its 
smooth forewings and male genitalia markedly distinct 
from typical Salebria; closest to Salebriacus from which 
it is at once distinguished by the compound scale tufts 
on eighth abdominal segment of the male and the place 
of departure of the ductus seminalis from bursa in the 
female. How many vaUd species are represented by the 
several names in our North American lists and definitely 
referable to the genus cannot be exactly determined 
until larger series of reared specimens are available. 
The genitaha (except for those of pumilella and jruc- 
tetella) offer httle or nothing in the nature of trust- 
worthy specific characters. The maculation and color 
differences on forewings that have been used by previous 
authors seem to be equally unreUable. 

235. Salebriaria turpidella (Ragonot), new comhination 
Figures 323, 802 

Salebria turpidella Ragonot, Nouv. Gen., p. 19, 1888; Monograph, 
pt. 1, p. 346, 1893.— Forbes, Cornell Mem. 68, p. 625, 
1923.— McDunnough, Check list. No. 6196, 1939. 

Salebria ademplandella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 114, 1908.— McDunnough, Check Hat, No. 6198, 
1939. (New synonymy.) 

Forewing powdery gray, the ground color variable, 
ranging from pale ash gray (ademptandella) to a more 
suffused pale brownish gray; basal area usually some- 
what paler than median area; the transverse lines but 
little paler than the ground color, indicated chiefly by 
their dark margins; the latter narrow, blackish, well 
contrasted, especially on the paler examples; antemedial 



116 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



line sinuate-angulate, bordered outwardly by a black 
line and inwardly, on lower half, by a similar narrow 
black liae; on many specimens a small, faint, whitish 
patch just beyond the antemedial line on inner margin; 
subterminal liae sinuate, bordered inwardly by a black- 
ish line, the latter fainter than that bordering the ante- 
medial line; discal spots black, confluent, normally 
forming a black line along the discocellular vein, but on 
individual specimens tending to separation on one or 
the other forewing. Hind wing smoky white to brown, 
variable in both sexes. Alar expanse, 17-18 mm. 

Male genitalia show no distinguishable differences 
from those of typical turpidella, nubiferella, or annulo- 
sella. Female genitaUa with the spining on anterior half 
of bursa of a nearly imiform fineness (no dense concen- 
tration of darker spines at the closed end) ; bursa with- 
out appreciable sclerotization at middle. 

Type localities: United States: {turpidella, (^ , 
in Paris Mus.); Kerrville, Tex. (ademptandella, in 
USNM). 

Food plant: Unknown. 

Distribution: Texas, Burnet County (Oct.), Kerr- 
vUle (Apr., May, June), Sabinal (Sept.), also one Texas 
male from the BoU Collection (No. 558) labeled 
"Europe"; Florida, Gotha (F. Kauterberg, collector, 
2 specimens, cf and 9). 

The foregoing from typical examples. Also before 
me are transitional examples between turpidella and 
nubiferella from Texas, Blanco County (May, July) 
and Shovel Mountain, and North Carolina, Southern 
Pines (Apr., July, Aug.) and Tryon (May, Sept.). 
These specimens are dark, with the size and markings 
of turpidella but with female genitalia more like those of 
annulosella. Some of them had been identified as 
annulosella. Others were under turpidella. I doubt 
very much if turpidella is anything more than a variety 
of nubiferella. Dyar's ademptandella is nothing but a 
paler form of the typical turpidella. 

236. Salebriaria nubiferella (Ragonot), new combination 

Salebria nubiferella Ragonot, N. Amer. Phycitidae, p. 8, 1887; 

Monograph, pt. 1, p. 344, 1893.— Hulst, Phycitidae of N. 

Anaer., p. 150, 1890.— McDunnough, Check list, No. 6191, 

1939. 
Salebria annulosella nubiferella (Ragonot) Forbes, Cornell Mem. 

68, p. 625, 1923. 

If this and the preceding "species" (turpidella) are 
anywise distinct, the name nubiferella will apply to a 
larger form (21 mm.) with the basal area of forewing 
dark and concolorous with the darker shading in median 
and outer areas and with a more strongly contrasted 
white patch on the inner margin just beyond the ante- 
medial line; but neither of these differences, nor those 
used by Ragonot (Monograph, pp. 329, 345) to separate 
turpidella and nubiferella are constant. According to 
Clarke's notes the species is represented in the Paris 
Museum only by the imique type, a male labeled "type 
orig., pi. XIV fig. 23, Amer. Sept. don. C. V. Riley." I 



have examined its genitalia and they show nothing to 
distinguish them from those of turpidella, annulosella, 
or engeli. 

What appears to be a typical female in the National 
Museum from Putnam County, lU. (July) has the well- 
contrasted white spot on inner margin, dark hind wings 
and dark gray forewings, the latter with discal dots 
fused and the dark (blackish) borders of antemedial and 
subterminal lines as in turpidella, but somewhat weaker. 
Its alar expanse is 20 mm. The female genitalia have a 
rather dense and dark concentration of fine spines at 
the anterior end of bursa and a thickening and slight 
sclerotization of the membrane at middle of bursa, the 
sclerotized part showing what seems to be a fixed longi- 
tudinal fold. In these particulars the genitaha are like 
those of annulosella. 

Type locality: "Amer. Sept." [Texas] (type in Paris 
Mus.) . 

Food plant: Unknown. 

Besides the aforementioned female there are before 
me a male (19 mm.) from Putnam County, 111. (July), 
and a male (18 mm.) and female (19 mm.) from near 
St. Louis, Mo. (June) which appear to be conspecific 
with the Illinois female. They have the white spot on 
inner margin less strongly marked, but the strength of 
this marking does not seem to be of any significance. 
Presumably several of the Texas examples from the 
intermediate specimens mentioned under turpidella 
should be referred here if a specific distinction can be 
maintained between turpidella and nubiferella. We 
shall have to have host plant association and good 
reared series before such a distinction can be made with 
any certainty. 

237. Salebriaria engeli (Dyar), new combination 

Salebria engeli Dyar, Journ. New York Ent. Soc, vol 14, p. 107, 
1906.— Forbes, Cornell Mem. 68, p. 625, 1923.— McDun- 
nough, Check list, No. 6195, 1939. 

Forewing dark brownish gray, the transverse lines 
obscured; antemedial line indicated only by its very 
faint, narrow, angulate, outer black border, followed 
outwardly on inner margin by a strongly contrasted 
white patch; subterminal line faint, but slightly paler 
than the ground color and with an obscure, narrow, dark 
inner border, sinuate; discal dots more or less confluent, 
blackish with some pale scaling on their outer margins; 
a row of separated blackish dots along outer margin. 
Hind wings smoky fuscous, on darker specimens with 
a pale brownish tint ; th e veins darkened. Alar expanse, 
18-20 mm. 

Genitaha as in annulosella. 

Type locality: Oak Station, Pa. (type in USNM). 

Food plant: Unknown. 

Distkibution: Pennsylvania, Hazleton, New 
Brighton (July), Oak Station (July); Maryland, Plum- 
mers Isl. (July); Illinois, Decatur (May); Texas, 
KerrvUle (May, June, July). 

An intermediate form between typical nubiferella and 
annulosella, probably only a variety of the former. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



117 



238. Salebriaria annnloBella (Ragonot), new combination 

Figure 800 

Salebria annulosella Ragonot, N. Amer. Phycitidae, p. 7, 1887; 

Monograph, pt. 1. p. 346, 1893.— Forbes, Cornell Mem. 68, 

p. 625, 1923.— McDunnough, Check list. No. 6197, 1939. 
Salebria robustella Dyar, Proc. Ent. Soc. Washington, vol. 10, 

p. 114, 1908.— McDunnough, Check list. No. 6211, 1939 

(New synonymy.) 

Forewing gray ; antemedial line but slightly paler than 
the ground color and not sharply defined, margined on 
outer side at costa by a more or less triangulate blackish 
patch and on inner side at inner margin by a somewhat 
larger blackish patch which extends to or nearly to base 
of wing; discal dots distinctly separated, black, sur- 
rounded by pale dusting; sub terminal line obscure; a 
row of distinct blackish dots along terminal margin. 
Alar expanse, 18-20 mm. 

Female genitalia figured from specimen from Bm-net 
County, Te.x. They are like those of the type in Paris 
and differ in no essential details from those of females 
of nubilella. 

Type localities: Texas (annulosella, in Paris Mus.) ; 
Burnet County (robustella, in USNM). 

Food plant: Unknown. 

Distribution: Texas, Blanco County (Apr.) ; Burnet 
County (Apr.); North Carolina, Raleigh (June). 

According to Clarke's notes, two examples, forming 
the basis of Ragonot's diagnosis of the species in his 
Monograph, are in the Paris Museum. One is a female 
from Texas, obviously the type but not so labeled. It 
was the specimen figured in the Monograph (pi. 9, fig. 
6). The other specimen is a male labeled: "Dallas, 
Texas, Boll, 24-V-689." A female in the U. S. Na- 
tional Museum, also a Boll specimen from Texas (No. 
559) and probably a mate to the Paris male, bears a 
name label (Salebria annulosella) in Ragonot's hand- 
writing. 1 doubt that either of these Boll specimens is 
a typical annulosella. Our female in markings is inter- 
mediate between annulosella and tenebrosella with female 
genitalia like the latter species. Dyar's type of robus- 
tella (a male) matches Ragonot's figure and description 
of annulosella in all details e.xcept that the discal spots 
are obscured on one forewing. A female, obviously the 
other sex of Dyar's type, had been identified by him as 
annulosella. The remaining Texas examples before me 
(Blanco County) had been identified by Hulst as either 
nubijerella or pumilella. 

239. Salebriaria tenebrosella (Hulst), new combination 
Figures 801, 803 

Nephopteryx tenebrosella Hulst, Ent. Amer., vol. 3, p. 131, 1887. 
Nephopteryx quercicolella Ragonot, N. Amer. Phycitidae, p. 7, 

1887. 
Salebria tenebrosella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 

1889; Monograph, pt. 1, p. 347, 1893.— Hulst, Phycitidae 

of N. Amer., p. 151, 1890.— McDunnough, Check list. No. 

6199, 1939. 
Salebria heinrichalis Dyar, Ins. Insc. Menstr., vol. 5, p. 45, 

1917.— McDunnough, Check list, No. 6192, 1939. (New 

synonymy.) 

Forewing dark gray more or less shaded with black, 
especially in basal area; antemedial line whitish, dif- 



fused, oblique, sometimes interrupted at middle, pre- 
ceded by a dark red patch on lower half; cutting this 
red patch an oblique black line which fuses into the 
black basal shade on costal half of basal area; subter- 
minal line obscure, sinuate, very slightly paler than 
the ground color; discal dots coalesced into a black 
lunulate line along discocellular vein, partially obscured 
in the dark ground color over middle of wing but set 
off by some pale (whitish) dusting along its outer edge; 
separated blackish dots along terminal margin. Hind 
wings brownish gray; the veins slightly darkened. 
Alar expanse, 17-18 mm. 

Male genitalia showing no distinguishing characters 
from those of preceding species. 

Type localities: Te.xas (tenebrosella, in AMNH, ex 
Rutgers) ; "America Septentrionalis" (querciocolella, in 
Paris Mus.) ; Falls Church, Va. (heinrichalis, in USNM). 

Food plant: Oak (larva a leaf tier) . 

Disteibution: Texas; Missouri (Aug.); Virginia, 
FaUs Church (Apr.). 

In addition to typical examples from the above lo- 
calities there is before me a series of males and females 
of a slightly larger average size, 18-20 mm., with a 
somewhat more diffused and contrasted whitish ante- 
medial line and little or no trace of the subbasal red 
patch on forewing and no red on the thorax. On 
typical tenebrosella the tips of the patagia are shaded 
with red. The females of these variant examples also 
have larger genitalia (fig. 803) . They may represent a 
food plant race or a color form of tenebrosella but hardly 
anything more. In view of the already obscure specific 
limits of the described species I do not feel justified in 
adding a further name. 

The variety is represented in the National Collection 
from the following localities: Massachusetts, Cohasset 
(July), Martha's Vineyard (July); New York, Utica 
(Aug.) ; Maryland, Plummers Isl. (Aug.) ; District oj 
Columbia, Washington (June) ; North Carolina, Tryon 
(May) ; Georgia, Atlanta (June) ; Illinois, Decatur 
(May), Lacon (Jime) ; Missouri, near St. Louis, and one 
small example from the Miu-tfeldt Collection labeled 
"1.30 M. apple, 5-8-89." Several of these had been 
previously misidentified as Oreana leucophaeella (Hulst) . 

240. Salebriaria pumUella (Ragonot), new combination 
Figures 325, 804 

Salebria pumilella Ragonot, N. Amer. Phycitidae, p. 8, 1887; 

Monograph, pt. 1, p. 344, 1893. — McDunnough, Check list, 

No. 6190, 1939. 
Salebria georgiella Hulst, Canadian Ent., vol. 27, p. 57, 1895. — 

McDunnough, Check list. No. 6210, 1939. (New synonj'- 

my.) 

Forewing dark gray shaded vdth brown; a rather pale 
chocolate brown, triangulate patch on inner margin on 
inner side of antemedial line; the latter narrow, white, 
its blackish bordering lines broken, and obscure except 
the inner one on lower half of wing; a similar black 
border on the inner edge of the chocolate brown patch; 
on some specimens (especially faded examples) a slight 
ocherous shading at extreme base of wing; on most fresh 



118 



tnsriTED STATES NATIONAL MUSEUM BULLETEST 207 



specimens a narrowly elongate, grayish fuscous patch 
on midcosta; some white dusting on inner margin 
beyond antemedial line and in cell towards its outer 
margin; subterminal line fine, white, its dark borders 
obscure; discal spots confluent, forming a narrow, 
blackish line along discocellular vein: the blackish dots 
along terminal margin weak, more or less confluent. 
Hind wing pale smoky fuscous. Alar expanse, 15-17 
mm. 

Male genitalia with spine from apex of sclerotized 
costa of harpe projecting straight out beyond apex of 
cucullus (not curved dorsally as in the other species) . 

Female genitalia with ductus bursae flattened, ribbon- 
like, sclerotized thi'oughout and dorsoventrally folded 
towards bursa copulatrix, its lower margin produced at 
genital opening into a subtriangulate projecting shield 
with somewhat rounded terminal margin; bursa with 
nearly half of one side strongly and smoothly sclero- 
tized. 

Type localities: Texas (pwnilella, in Paris Mus.); 
Charlotte Harbor, Fla. (georgieUa, in AMNH, ex Rut- 
gers). 

Food plant: Unknown. 

Distribution: Florida, Charlotte Harbor (Mar.); 
Texas, Burnet County (Apr.), also 3 examples (c? and 
99) with only the state locality and without dates; North 
Carolina, Southern Pines (June, Aug.) . 

Hulst's references to pwmilella in his Phycitidae of 
N. Amer. (p. 150) are omitted from the above s5Tionymy, 
since he had misidentified Eagonot's species. The 
technical description he gives was simply copied from 
Eagonot's original description. There is no doubt 
about the synonymy of georgieUa. 

In all examples I have seen, veins 4 and 5 of forewing 
are rather closely approximate. 

241. Salebriaria firactetella (Hulst), new combination 
FiGTJEBs 324, 805 

Myelois fructetella Hulst, Canadian Ent., vol. 24, p. 59, 1892. 

Salebria rectistrigella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 115, 1908. 

Salebria fructetella (Hulst) Barnes and McDunnough, Contribu- 
tions, vol. 3, p. 194, 1916. — McDunnough, Check list, No. 
6203, 1939. 

Forewing glossy brownish gray shaded with blackish 
brown and dusted with white; the black shading form- 
ing a blotch on costa following, and a similar blotch on 
inner margin preceding the antemedial line, and an 
obscure dark shade extending from costa near apex 
obliquely inward across the subterminal line; white 
dusting limited to a more or less triangulate cloud from 
midcosta surrounding the upper discal spot and a 
smaller cloud on costa preceding the antemedial line; 
antemedial line narrow, slightly oblique, straight or (on 
some examples) with a slight notch at middle, white and 
distinctly contrasted against the ground color but with- 
out appreciable black bordering lines; discal dots sep- 
arated, blackish; subterminal line obscure, whitish gray, 
vertical except for a median outward bulge; terminal 
dots obscure, brown. Hind wing smoky white shading 



to fuscous along termen; veins very slightly darkened. 
Alar expanse, 15-18 mm. 

Male genitalia distinguished from those of other 
species in the genus by the spoon-shaped apical projec- 
tion of gnathos, the V-shape of the sclerotized part of 
anellus, and the longer cornutus. 

Female genitalia with ductus bursae very short, its 
median area membranous, a moderately broad sclero- 
tized and granulate band at genital opening, strongly 
sclerotized and longitudinally ridged towards junction 
with bursa; this peculiar sclerotization extending into 
and occupying nearly half of the bursa, anterior (closed) 
end of bursa fused into a thickened (but not sclerotized) 
membrane; remainder of bursa covered by a mat of 
fine spines. 

Type localities: Blanco County, Tex. (fructetella, 
in AMNH, ex Eutgers); Kerrville, Tex. (rectistrigella, 
in USNM). 

Food plant : Oak (larva a leaf feeder) . 

Distribution: Florida, Key West, Orlando (Mar.); 
Louisiana, Natchitoches Parish (Aug.); Texas, Blanco 
County (June), Kerrville (May, Jime), Sabinal (Sept.), 
Shovel Mountain (June, July); Arizona, Williams 
(Sept.), Missouri (June, reared); District of Columbia 
(May, Aug., reared); New York, Bellport (June, Sept., 
reared) . 

Superficially /ritcfefeiZa strikingly resembles Acrobasis 
amplexella Eagonot (especially the females) . The reared 
examples before me had been identified to the latter 
name. The male antennal character and the genitalia 
of both sexes, however, easily separate the two species. 

64. Quasisalebria, new genus 

Type of genus: Quasisalebria admixta, new species. 

Characters of Salebriaria except: Labial palpus erect, 
appressed to flattened face. Forewing with vein 8 and 
9 stalked for slightly more than two-thirds of their 
lengths. Hind wing with veins 4 and 5 stalked for over 
three-fourths of their lengths; 7 and 8 shortly anas- 
tomosed beyond cell. 

Male genitalia with costa of harpe sclerotized for its 
entire length, not appreciably produced at apex, but 
with a strongly sclerotized, free, articulating arm from 
base; shieldlike part of anellus with a pair of long, 
divergent, lateral horns; penis without cornutus. 

Female genitalis with ductus bursae thickened 
(cartilaginous in texture) except at junction with bursa 
copulatrix; bursa finely spined only in area adjacent to 
junction with ductus bursae. 

The type of this genus is, in many respects, close to 
Salebriaria fructetella and appears to be an aberrant 
offshoot of Salebriaria. I propose the new generic 
designation reluctantly; but no other procedure is pos- 
sible if we are to have any exact definition of genera for 
the species allied to Salebria. Nothing in our American 
fauna is properly referable to the latter genus, which is 
characterized by a forewing with partial scale ridge in 
the subbasal area; harpe (fig. 332) with erect clasper 
from near middle and costal margin weakly sclerotized; 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



119 



bursa copulatrix of female without spining or granula- 
tions (membranous and smooth in the type, palumbella). 

242. Quasisalebria admlxta, new species 
Figures 328, 806 

Forewing ashy white shaded with olivaceous brown 
or grayish fuscous in outer area and on lower half of 
basal area; the whitish ground color strongly contrasted 
on costal half of basal area and in a more or less tri- 
angulate area extending from median half of costa into 
cell and including the discal spots; antemedial line 
distinct, narrow, slightly curved, white, bordered out- 
wardly on costa by a strongly contrasted, black, tri- 
angulate patch and preceded on inner margin by a 
similar quadrate spot; subterminal line narrow, white, 
close to termen, and outwardly bulged at middle, 
bordered by fine blackish lines which begin as strong 
black smudges at costa; discal dots separated, the lower 
one always distinct, the upper sometimes absent. Hind 
wing semihyaline, white with a brownish shade at apex 
and a narrow brown line along upper half of termen; 
the veins not appreciably darkened. Alar expanse, 
19-21 mm. 

Male genitalia. Characters as given for the genus. 
The pecuhar development of the aneUus may be only 
of specific significance. 

Type locality: Provo, Utah (type in USNM, 
61343). 

Food plant: Unknown. 

Described from male type and two male and four 
female paratypes from the type locality (July, Aug.); 
one male paratype from Kedington, Ariz.; two female 
paratypes from BeUevue, Washington County, Utah 
(May); and one female paratype from Glenwood 
Springs, Colo. (Aug.). 

The species is superficially similar to Salebriaria 
fructetella Hulst, but with the white areas and blackish 
markings of forewing more strongly contrasted. 



Genus 65: Ortholepis 

[Venational division B. Veins 4 and 5 of forewing connate 
(rarely in individual specimens, slightly separated at base). 
Hind wing with vein 2 from rather near lower outer angle of 
cell. Male genitalia with transtilla complete but its median 
area weakly sclerotized and granulate; costa of harpe strongly 
sclerotized throughout but not produced at apex; penis armed 
with a single, long, strong cornutus. Female genitalia without 
signum or scobinations in bursa.] 

65. Genus Ortholepis Ragonot 

Ortholepis Ragonot, N. Amer. Phycitidae, p. 6, 1887; Monograph, 
pt. 1, p. 214, 1893.— Hulst, Phycitidae of N. Amer., p. 140, 
1890. (Type of genus: Ortholepis jugosella 'Ra.gonot.) 

Tongue well developed. Antenna of male pubescent, 
shaft with sinus towards base contarning a row of black 
toothlike spines more or less concealed imder a weak 
scale tuft; anteima of female simple. Labial palpus 
obligue; second segment roughly and rather broadly 
scaled, on male slightly grooved on inner side; third 
segment very short, acuminate, reaching to height of 



vertex. Maxillary palpus of male squamous (jugosella) 
or in the form of an aigrette (pasadamia) ; of female 
minute and filiform. Forewing with ridge of raised 
scales on inner side of antemedial line, not reaching 
costa or inner margin; 11 veins; 2 from before lower 
outer angle of cell ; 3 from the angle, separated at base 
from 4-5; 4 and 5 cormate, rarely (in individual speci- 
mens) slightly separated at base; 6 from below upper 
angle of ceU, straight, 8 and 9 stalked for slightly more 
than half their lengths; 10 from the cell, shortly sepa- 
rated from 8-9 at base and thence divergent; male 
without costal fold. Hind wing with vein 2 from before 
but rather near lower outer angle of cell; 3 from the 
angle, connate with the stalk of 4-5, short as compared 
with 2 ; 4 and 5 stalked for half or a trifle over half their 
lengths; 7 and 8 contiguous or anastomosed for a very 
short distance beyond cell ; cell less than half the length 
of wing; discocellular vein curved, considerably extended 
at lower angle. Eighth abdominal segment of male 
simple. 

Male genitalia with uncus sub triangulate; shghtly 
produced (lobed) near its lower, lateral angles. Apical 
process of gnathos a short, simple hook. Transtilla 
complete but its median area weakly sclerotized and 
granulate; its lateral elements broadly sclerotized. 
Harpe narrow ; costal margin sclerotized throughout but 
not produced. Anellus U-shaped, narrowly sclerotized. 
Penis armed with a single stout cornutus nearly as long 
as aedeagus. Vinculum stout, triangulate; about as 
long as greatest width. 

Female genitalia without signum; bursa elongate, 
large, longitudinally wrinkled, partially sclerotized in 
the lobed area bearing the ductus seminalis; ductus 
bursae considerably shorter than bursa; flattened, 
strongly sclerotized, at least near and at its jimction 
with bursa copulatrix; ductus seminahs from lobe of 
bursa near junction of bursa and ductus bursae. 
Eighth-segment collar with sclerotized part on dorsum 
reduced to a U-shaped band. 

An American genus showing afl[inities to Polopeustis 
and the various genera of the Salehria complex; but 
easily distinguished by its genitalia. Kagonot's de- 
scription is somewhat misleading. Veins 4 and 5 of 
forewing are normally connate and not "nearly paral- 
lel" except well beyond base; from base to near middle 
they are divergent. 

243. Ortholepis jugosella Ragonot 

Figures 23, 329, 808 

Ortholepis jugosella Ragonot, N. Amer. Phycitidae, p. 6, 1887; 
Monograph, pt. 1, p. 214, 1893.— Hulst, Phycitidae of N. 
Amer., p. 140, 1890. — McDunnough, Check list. No. 6149, 
1939. 

Maxillary palpus of male squamous. 

Forewing gray, rather shiny; palest (on some speci- 
mens ash gray) on costal half of median area and on the 
anterior costal half of basal area; remainder of basal 
and median areas and the area beyond the subterminal 
line brownish gray with a faint pm-plish suffusion; 



120 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



antemedial line oblique from costa to lower margin of 
cell, thence vertical to inner margin, its upper half 
obscure, indicated chiefly by a blackish outer border, 
lower half distinct, whitish; the antemedial line pre- 
ceded by a raised tuft of brown and blackish scales 
which are bordered inwardly by a more or less con- 
trasted white line; subterminal line faint, narrow, 
slightly bulged at middle; discal dots black, separated, 
distinct and rather large, especially the upper one; 
terminal dots reduced, obscure and more or less conflu- 
ent. Hiad wing light brown ; a thin blackish line along 
termen; the veins not appreciably darkened. Alar 
expanse, 19-20 mm. 

Male genitalia as given for the genus. Female geni- 
talia with posterior half of ductus bursa very weakly 
sclerotized. 

Type locality: "America septentrionalis" (type in 
Paris Mus.). 

Food plant: Hickory {Carya alba) and wild azalea 
(Ragonot records). Hickory and wahiut are the more 
probable hosts. 

Distribution: United States: Connecticut, East 
River (July). Canada: Nova Scotia, White Point 
Beach (Queens County, July). 

244. Ortholepis pasadamia (Dyar), new combination 

FionBE 807 

Immyrla pasadamia Dyar, Ins. Insc. Menstr., vol. 5, p. 45, 
1917.— McDunnough, Check liat, No. 6189, 1939. 

Maxillary palpus of male in the form of a short aig- 
rette. 

Forewing uniformly dark glossy gray with a purplish 
tint; the transverse lines well marked, narrow, whitish; a 
contrasted white inner margin to the subbasal tuft; dis- 
cal dots usually confluent, distinguishable but not 
strongly contrasted against the dark ground color. 
Hind wing smoky fuscous; the veins faintly darkened. 
Alar expanse, 17-20 mm. 

Male genitaUa as in jugosella. Female genitalia as in 
jugosella except ductus bursae sclerotized along ventral 
surface to genital opening. 

Type locality: St. Johns, Quebec (type in USNM). 

Food plant: Betula. 

Distribution: United States: Pennsylvania, Ha- 
zleton (June); New Hampshire, Dublin, Hampton 
(July) ; Maine, Mount Desert Island (July) , Sebec Lake 
(July); Washington, Meadow Creek (Grant County, 
Apr.), reared specimen, Walla Walla (June). Canada: 
Ontario, Blacotasing (July), Ottawa (July), Waubamia 
(Perry Sound, July) ; Quebec, St. Johns (June) . 

The foregoing description was drawn from typical 
examples represented in the National Collection by a 
series of 16 males and females. There are also before 
me four specimens from Maine, New Hampshire, and 
Quebec of what appears to be a color form in which the 
transverse lines (except for the white inner border of the 
raised-scale patch) are almost completely obliterated; 
the ground color of the forewing is darker (more pur- 
plish) and the hind wing is brown (as in jugosella) ; there 



is also a faint narrow dusting of whitish scales in the 
median area of forewing. The genitalia of these speci- 
mens are identical with those of typical pasadamia. 
Examples of both forms have been reared from Betvla. 



Genus 66: Polopeustis 

[Venational division B. Vestiture of head, thorax, labial palpi, 
and femora a mixture of scales and hairs.] 

66. Genus Polopeustis Ragonot 

Pohpeustis Ragonot, Monograph, pt. 1, p. 233, 1893. — Staudin- 
ger and Rebel, Catalog der Lepidopteren des palaearctichen 
Faunengebietes, vol. 2, p. 30, 1901. — Spuler, Die Schmetter- 
Unge Europaa, vol. 2, p. 212, 1910. — Forbes, Cornell Mem. 
68, p. 622, 1923.— Janse, Journ. Ent. Soc. South Africa, vol. 
5, p. 30, 1942. (Type of genus: Phycis annulatella Zetter- 
stedt; figs. 24, 330, 809.) 

Tongue weU developed. Antenna of male ciliate- 
pubescent (the cilia a trifle shorter than width of seg- 
ments) the shaft curved towards base and containing 
two or three short teeth in the incurvation; of female 
simple. Labial palpus obliquely ascending, not reach- 
ing height of vertex; third segment short, less than one- 
third of second. MaxUlary palpus minute, filiform. 
Forewing smooth; 11 veins; vein 2 from before but near 
lower outer angle of cell; 3 from the angle; 4 and 5 
separated at base; 6 from below upper angle of cell, 
straight; 8 and 9 stalked for over half their lengths; 10 
from the cell, shortly separated from the stalk of 8-9 at 
base; male without costal fold. Hind wing with vein 2 
from well before outer angle of cell; 3 from the angle; 4 
and 5 anastomosed for about half their lengths beyond 
cell; 7 and 8 closely approximate for a short distance 
beyond ceU; cell less than half the length of wing; dis- 
cocellular vein curved, produced at lower angle. Eighth 
abdominal segment of male with pair of ventrolateral 
hair tufts. 

Vestiture of head, thorax, labial palpi, femora, and 
and foretibiae a mixture of scales and hairs. 

Male genitalia with uncus about as broad as long; at 
apical margin broadly rounded. Apical process of 
gnathos (from ventral view) an inverted heart-shaped 
lobe terminating in a short, slender, hooked spine. 
Transtifla absent. Harpe short, stubby; its apex bluntly 
rounded; costa broadly and strongly sclerotized through- 
out, but not produced at apex; otherwise simple. Anel- 
lus a U-shaped plate terminating in short lateral lobes. 
Aedeagus slender; penis armed with moderately stout, 
elongate cornutus or two similar cornuti. Vinculum 
stout, slightly longer than greatest width; tapering to 
blunt and rather broad terminal margin. 

Female genitalia without signum; bursa copulatrix 
small and with a rather broad, fused cartilagenouslike 
thickening around its lateral and anterior margins, 
otherwise minutely granulate and containing a weak 
elongate chitinized strip ; ductus bursae twice as long as 
bursa, flattened, its ventral surface sclerotized and gran- 
ulate through its length, the sclerotization extending 
into bursa; genital opening simple. Eighth-segment 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



121 



collar with sclerotized area reduced to a narrow, more 
or less U-shaped dorsolateral band (similar to that of 
Ortholepis) . 

An Old World genus of Holarctic distribution with 
one European and one North American species; easily 
identified by its genitalia and hairy vestiture. 

245. Polopeustis arctiella (Gibson) 
Figures 331, 810 

Pyla arctiella Gibson, Report of the Canadian Arctic Expedition 
(1913-18), vol. 3, pt. 1, p. 46, 1920. 

Polopeuslis annulatella arctiella (Gibson) McDunnough, Ca- 
nadian Ent., vol 67, p. 174, 1935; Check list. No. 6150, 1939. 

ForeAsdng slate gray with a fine scattered sprinkling 
of white, the whitish dusting more concentrated in basal 
area and the area beyond the sub terminal line; trans- 
verse lines rather broad, well contrasted, white; ante- 
medial line oblique and slightly angulate; sub terminal 
line sinuate; a narrow blackish shade bordering the 
antemedial outwardly and the subterminal inwardly; 
discal dots obscure, more or less confluent. Hind wings 
smoky white; the veins darkened; a narrow dark shade 
along termen. Alar expanse, 21-25 mm. 

Male genitalia with two cornuti on penis ; otherwise as 
in annulatella. Female genitalia differ from those of 
annulatella chiefly in the shape of the sclerotized area of 
eighth-segment collar (compare figs. 809a and 810a). 
The differences in shape and extent of sclerotized area 
of bursa are probably individual in character. 

Type locality: Collinson Point, Alaska (type in 
Canadian Nat. Coll.). 

Food plant: Unknown. 

Distribution: Alaska: Collinson Point (July). 
Canada: Labrador, Hopedale (July), Nain; Manitoba, 
Fort Churchill (June, July). 

Gibson's name may represent no more than a New 
World race of annulatella; but arctiella is at least that. 
Good series of both are before me and their genitalic 
differences appear to be constant; the male of annulat- 
ella has but one cornutus, that of arctiella has consist- 
ently two comuti. If and when intergrading examples 
of Polopeustis are recovered from northern Siberia the 
name arctiella may be reduced to subspecific status or 
referred as a synonym to annulatella. Meanwhile a 
specific separation seems the safer procedure. 

Genera 67-70: Meroptera to Tulsa 

[Venational division B. Veins 4 and 5 of forewing very shortly 
stalked, connate or closely approximate at base, in Tulsa approxi- 
mate at base and for a short distance beyond; 8 and 9 long stalked; 
10 frequently connate or shortly stalked with 8-9, if from the cell 
approximate to the stalk of 8-9 for some distance from its base. 
Antenna of male with sinus and scale tuft in base of shaft. Labial 
palpus erect or obUquely upturned. Maxillary palpus of male 
in the form of an aigrette or squamous. Male genitalia with 
transtilla frequently complete, but if so, weakly sclerotized; 
harpe with clasper always present and well sclerotized, digitate 
or enlarged and spined; harpe with long hair brush from inner 
surface along lower edge of basal half of sclerotized costa; penis 
armed with two stout, rather short cornuti. Female genitalia 



with bursa finely and densely spined, usually with one or more 
sclerotized, granulate patches.] 

67. Genus Meroptera Grote 

Meroptera Grote, Canadian Ent., vol. 14, p. 29, 1882. — Hulst, 
Phycitidae of N. Amer., p. 148, 1890. — Ragonot, Mono- 
graph, pt. 1, p. 312, 1893.— Forbes, Cornell Mem. 68, 
p. 624, 1923. (Type of genus: Pempelia pravella Grote.) 

Emmeriia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 76, 
1930. (Type of genus: Meroptera mirandella Ragonot. New 
synonymy.) 

Tongue well developed. Antenna weakly pubescent; 
on male with sinus and scale tuft in base of shaft. La- 
bial palpus upturned, closely appressed to face; smooth 
scaled; reaching above vertex; second segment long, on 
male hoUowed to receive maxillary palpus; third seg- 
ment short (about one-fourth of second), acuminate. 
Maxillary palpus of male in the form an of aigrette; 
of female squamous. Forewing smooth; 11 veins; vein 

2 from before but rather near lower outer angle of cell; 

3 from the angle, at base slightly nearer to 4-5 than to 
2; 4 and 5 very shortly stalked, connate or closely ap- 
proximate at base; 6 from below upper angle of cell, 
straight or (rarely) very slightly bent towards base; 8 
and 9 stalked for over two-thirds their lengths; 10 
shortly stalked or connate with the stalk of 8-9 (indi- 
vidually variable); male without costal fold. Hind 
wing with vein 2 from before but near lower outer angle 
of cell; 3 from the angle, connate with 4; 4 and 5 stalked 
for slightly more than half their lengths; 7 and 8 closely 
approximate for a short distance beyond cell; cell less 
than one-half the length of wing; discocellular vein 
curved, outwardly produced at lower angle of cell. 
Eighth abdominal segment of male with compound 
scale tufts. 

Male genitalia with uncus triangulate or subtriangu- 
late, its apex bluntly rounded. Apical process of 
gnathos a short, stout hook. Transtilla complete but 
weakly sclerotized; a narrow, flatly arched band. 
Harpe with costa sclerotized throughout but not pro- 
duced at apex; a fine brush of long hairs arising from 
inner surface along lower edge of basal half of sclero- 
tized costa; cucullus simple, narrow, tapering sHghtly 
to rounded apex; from near base of harpe an appressed, 
stout, thorny or serrate clasper. Penis armed with 
two, stout, rather short cornuti less than one-half the 
length of aedeagus. Vinculum stout, as long as or 
somewhat longer than greatest width, its terminal 
margin bluntly rounded or narrowly truncate. 

Female genitalia without signum ; bursa with a couple 
of conspicuous round or oval, strongly pigmented and 
sclerotized, densely granulate patches, otherwise bursa 
finely spinose over its membranous areas; ductus bursas 
shorter than bursa, not sclerotized adjacent to bursa, 
but with strong sclerotization at broadened genital 
opening; ductus seminaUs from a lobe of bursa near 
junction of bursa and ductus bursae. 

This genus, while distinct from, is very close to 
Nephopteryx, from which it is distinguished only by the 



122 



insriTED STATES NATIONAL MUSEUM BULLETIN 207 



peciiliar development of the clasper on harpe of the 
male genitalia and the strong sclerotization about the 
genital opening of the female. Hampson's Emmerita 
has not a single character to separate it from Meroptera. 
The stalking of vein 10 of forewing with 8-9, upon 
which Hampson evidently relied, is shared by the types 
of both Meroptera and Nephopteryx. This stalking is 
very short at most and is not even specifically constant 
in either genus. 

The genus as here defined contains but four North 
American species, and (to the best of my knowledge) 
no Old World representatives. 

246. Meroptera mirandella Ragonot 
Figures 21, 333, 816 

Meroptera mirandella Ragonot, Monograph, pt. 1, p. 313, 1893. 
Emmerita mirandella (Ragonot) Hampson, Ann. Mag. Nat. Hist., 

ser. 10, vol. 5, p. 76, 1930.— McDunnough, Check list, 

No. 6181, 1939. 

Forewing ocherous white with a very faint and 
scattered dusting of blackish scales in median area; 
antemedial line narrow, oblique, indented at lower fold, 
bordered outwardly by a black hne which begins as an 
enlarged, more or less triangular dash on costa, in- 
wardly by a straight black line from top of ceU to inner 
margin; sub terminal line obscure, dentate-sinuate, 
margined inwardly and outwardly for a short distance 
from costa by black lines, on well marked specimens the 
inner line continued as a fine dark border to tornus; 
also on well marked examples a faint blackish or 
fuscous shade extending obliquely across the wing from 
the inner costal edge of the antemedial line; discal dots 
faint, but usually distinguishable, blackish, separated, 
obliquely placed; terminal black dots more or less con- 
fluent. Hind wing white with a faint creamy or smoky 
tint; a very faint brownish line along termen; veins not 
or very sHghtly darkened. Alar expanse, 23-26 mm. 

Male genitalia with broad, irregularly fan-shaped 
clasper. Female genitalia with sclerotization of genital 
opening a narrow, corrugate, sclerotized band with 
short lateral, inward projections; bursa with two 
opposed, lateral, granulate patches. 

Type locality: Colorado (type in Paris Mus.). 

Food plant: Popidus (this record from an Arizona 
specimen in the National Collections reared from a 
cocoon on a cottonwood leaf) . 

Distkibution: Colorado, Denver; Arizona, Douglas 
(Aug.), Nogales (July), Phoenix (Apr.), Pinal Mts. 
(May), Keadington, Tucson (Apr.), Ymna (June). 

One of the females before me from Phoenix is de- 
cidedly abnormal in venation, the forewing having 
vein 10 from the cell and closely approximate to the 
stalk of 8-9, 6 bent at base and from very near the upper 
angle of the cell and 4 and 5 closely approximate at base 
and for some little distance from the cell. Its genitalia 
are normal and it is clearly an individual aberration, 
but an example of what occurs all too often in the 
Phycitidae and which should caution us to use generic 
keys with discretion and to place unusual specimens 
only after examination of their genitalia. 



247. Meroptera cviatella Dyar 
Figure 817 

Meroptera cviatella Dyar, Proc. Ent. Soo. Washington, vol. 7, p. 34, 

1905.— McDunnough, Check list. No. 6182, 1939. 
Salehria cviatella (Dyar) Forbes, Cornell Mem. 68, p. 627, 1923. 

Forewing bright reddish brown with a more or less 
distinct, transverse purplish gray shading in outer area 
and bordering outwardly the sub terminal line; this 
latter area sometimes dusted with blackish scales; the 
central area near antemedial line more or less dusted 
with whitish scales; antemedial line oblique, narrow, 
strongly indented at lower fold, its upper half obscured; 
the white line bordered outwardly by a conspicuous 
black band, broad on costa and very gradually narrow- 
ing towards inner margin; inwai'dly bordered on lower 
half by a similar broad, vertical black band; sub terminal 
line diffused, whitish; discal dots confluent, black, the 
lower one sometimes obscm-ed; a row of more or less 
confluent black dots along termen. Hind wing pale 
smoky brown, darkened slightly towards termen. Alar 
expanse, 23-25 mm. 

Male genitalia similar to those of mirandella. Fe- 
male genitaUa with sclerotization of genital opening a 
narrow, dark, granulate band, broken and with two 
short, divergent extensions at middle, these extensions 
fusing into a small, thin, triangulate, sclerotized patch 
on the lower median surface of the ductus bursae. 

Type locality: Chicago, lU. (type in USNM). 

Food plant: Pojmlus (cottonwood). Larva a borer 
in buds and new shoots. 

Disteibution: Illinois, Chicago (June, July), Lacon 
(Aug.), Putnam County (July, Aug.); Mississippi, 
Starkville (July). 

The above food-plant record is from Putnam Coimty 
specimens reared by Mr. Murry O. Glenn. One of 
his series is labeled "bred from larva on Amorpha 
canescen,s." I doubt if Amorpha is a true food plant or 
the presence of the larva on this plant anything more 
than an accidental last-stage migration from cotton- 
wood. 

248. Meroptera pravella (Grote) 
Figures 22, 334, 812 

Pempelia pravella Grote, Bull. U. S. Geol. Geogr. Surv. Terr., 
vol. 4, No. 3, p. 694, 1878. 

Meroptera pravella (Grote), Canadian Ent., vol. 14, p. 30, 1882. — 
Packard, U. S. Dep. Agr. Div. Ent. Bull. 13, p. 23, 1887; 
U. S. Dep. Agr. Fifth Rep., Ent. Comm., p. 574, 1890.— 
Ragonot (in part). Monograph, pt. 1, p. 314, 1893. — Mc- 
Dunnough, Check list, No. 6184, 1939. 

Forewing dark gray; the basal area, except for a 
narrow dark gray shading at extreme base, pale ashy 
gray and contrastingly paler; antemedial Hne obscure 
and often partially obliterated, when distinguishable 
it is oblique, narrow, duU white, distinguished chiefly 
by its black borders, which tend to coalesce into a 
broad, diffused, blackish band; subterminal line faint 
but distinguishable, grayish white, sinuate, in fresh 
specimens bordered inwardly by an obscure, narrow, 
blackish line; discal dots separated, blackish, not con- 
spicuous; a faint blackish line along termen. Hind 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



123 



wing very pale smoky fuscous. Alar expanse, 20-22 
mm. 

Male genitalia with an elongate, stout, tapering 
clasper extending about half the length of harpe; 
figiu-ed from male from Edmonton, Alberta, and com- 
pared by Clarke with the genitalia of the type of 
pravella in the British Museum. Female genitalia with 
sclerotization at genital opening in the form of a broad, 
stout, curved, granulate and wrinkled, lunate plate 
with strong lateral arms projecting into the ductus 
bursae. 

Type locality: Old town, Maine (type in BM). 

Food plant: Populus, Salix [?]. 

Distribution: United States: Maine; New Hamp- 
shire, Hampton (July); New York, Ilion (June); Colo- 
rado, Chimney Gulch (Golden, June). Canada: Que- 
bec, Knowlton (Feb., reared in laboratory from larva 
on aspen), Norway Bay (June); Ontario, Constance 
Bay (Feb., from poplar). Grand Bend (July), Hymers 
(June), Mer Bleue (June), Ottawa (Mar., from poplar). 
Smoky Falls (Mattagami River, June), Trenton (June) ; 
Manitoba, Aweme (May, June, July), Winnipeg; Sa- 
skatchewan, Indian Head (June, July); Alberta, Bilby 
(June, July) , Edmonton (May, July) , Nordegg (July) ; 
British Columbia, Canim Lake (June), "100 Mile 
House" (June). 

These records (except for the type locahty) are from 
specimens before me from the U. S. and Canadian 
National Collections. The Salix plant record is from 
Packard. I have not seen the reared Brunswick, 
Maine, specimens upon which it was based but have 
httle doubt that the name pravella was correctly ap- 
plied. Most of the specimens in our collections that 
have been identified as pravella as well as many of the 
references in literatiu-e are referable to Nephopteryx 
subfuscella (Ragonot) {=semiobscurella (Hulst)). This 
confusion is discussed under subjuscella. The two 
species are easily confused, especially with worn exam- 
ples, if their genitalia are not examined ; but in unrubbed 
specimens pravella is easily separated from subjuscella 
by the lack of any reddish or reddish ocherous scaling 
adjacent to inner margin at the base of forewing. Also 
included under pravella in the U. S. and Canadian 
National Collections were 21 Canadian examples of a 
new species (hereinafter described as abditiva) similar 
in all superficial characters to pravella but with dis- 
tinctly dijfferent male and female genitalia. 

249. Meroptera abditiva, new species 
FiGTTKES 335, 813, 814 

In color and maculation like pravella but with con- 
sistently different genitalia. Alar expanse, 19-22 mm. 

Male genitalia with harpe having a stout but short, 
stubby clasper, less than one-fourth the length of harpe. 
Female genitalia with the sclerotization at genital open- 
ing in the form of a rather narrow granulate, curved, 
transverse band without inwardly projecting arms. 
The pigmented, granulated area of the bursa is individ- 
ually variable in extent and sometimes divided into two 
distinct patches by a shght break at the anterior 



(closed) end of the bursa. Extremes of variation are 
shown in figures 813 and 814. However, there are no 
intergradations whatever between abditiva and pravella 
in the structure of the female genital plate nor in the 
male clasper. 

Type locality: Knowlton, Quebec (type in Cana- 
dian Nat. Coll. ; paratypes in Canadian Nat. Coll. and 
USNM, 61344). 

Food plant: Populus tremuloides. 

Described from male type and 3 female paratypes 
from the type locahty reared (in laboratory) Feb. 12, 
1 and 2, 1930, from larvae feeding on leaves of P. tre- 
muloides ("aspen") by J. McDunnough; and 17 para- 
types from the following Canadian localities: Quebec, 
Mount St. Hilaire, June 30, 1908, G. Chagnon (9); 
Norway Bay, June 4, 1938, E. G. Lester (cf). New 
Brunswick, Chamcook, June 23, 1938, T. N. Freeman 
(9); Eel River, June 21, 1941, T. N. Freeman (cf). 
Nova Scotia, Beddeck, June 23, 1936 and June 27, 1938, 
T. N. Freeman (29) ; White Point Beach, Queens, Feb. 
12 and 20, 1936, J. McDunnough (2 9, reared). 
Ontario, Trenton, May 29 and June 25, 1908, Evans 
(2 9) ; Vineland Station, June 15, 1936, W. L. Putnam 
(9, reared). Saskatchewan, Christopher Lake, June 
19, 1939, A. R. Brooks (cf). British Columbia, 
Canim Lake, June 25, 1938, J. K. Jacob (9) ; Jesmond, 
July 13, 1937, J. K. Jacob (2 9); Kaniloops, June 14, 
1937, J. K. Jacob {d'); Shingle Creek, Penticton, 
June 25, 1935, A. K. Cartrell (9). 

In as much as pravella and abditiva have the same 
hosts and larval habits, an overlapping distribution, 
and similar habitus, it is necessary to examine their 
genitalia to distinguish them apart. 

68. Genus Nephopteryx Hiibner 

Nephopteryx Hiibner, Verzeichniss bekannter Schmett[er]linge, 
p. 370, 1825.— Zeller, Isis von Oken, 1846, p. 731.— Grote, 
Bull. U. S. Geol. Geogr. Surv. Terr., vol. 4, p. 695, 1878; 
North Amer. Ent., vol. 1, p. 11, 1879.— Ragonot, Ent. 
Monthly Mag., vol. 22, p. 19, 1885 (citation of type) ; Mono- 
graph, pt. 1, p. 254, 1893. — Hulst, Phycitidae of N. Amer., 
p. 142, 1890.— Forbes, Cornell Mem. 68, p. 622, 1923.— 
Hemming, Hubner, vol. 2, p. 229, 1937.— Bisset, in Pierce 
and Metcalfe, Genitalia of the British Pyrales, p. 61, 1938. — 
Janse, Journ. Ent. Soc. South Africa, vol. 5, p. 34, 1942. 
(Type of genus: Phycita rhenella Zincken; Europe; figs. 25, 
336, 815.) 

Sciota Hulst, Ent. Amer., vol. 4, p. 115, 1888. (Type of genus: 
Sciota croceella Hulst.) 

Characters of Meroptera except: Labial palpus erect 
or obhquely upturned. Maxillary palpus of male in 
the form of an aigrette or squamous. Forewing with 
10 usually connate with the stalk of 8-9 or closely 
approximate to it, rarely stalked. Transtilla frequently 
incomplete or absent. Clasper of harpe digitate, slen- 
der, simple (without spining). One cornutus on penis in 
uvinella, other species have two cornuti as in Meroptera. 
Female genitalia with ductus bursae sclerotized along 
ventral surface for most of its length from junction with 
bursa, the sclerotization terminating before genital 
opening, the latter simple (unsclerotized). 



124 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



As here defined the genus includes a number of 
species with two types of maxillary palpi, several of 
which have been hitherto referred to either Salebria or 
Myrlaea. The reference of some of our North American 
species to the latter genus on the strength of a slight 
obliqueness of their labial palpi is not warranted as 
none of them agrees with the Old World type of Myrlaea 
(albistrigella Staudinger) on either male or female 
genitalic characters. The ductus bursae and bursa of 
albistrigella are perfectly smooth, without granulations, 
scobinations, or sclerotizations of any kind; and its 
harpe lacks the hair brush characteristic of Meroptera 
and Nephopteryx. 

The two types of male maxillary palpi (aigrettelike 
and squamous) do not justify generic separation of the 
species here any more than they do in Dioryctria or 
Ortholepis although such a difference should be, and in 
the Phycitidae usually is, of generic significance. On 
basilaris, the palpus is midway between the two types, 
though somewhat more aigrettelike than squamous. 
The uniform type of their male and female genitalia and 
the similar habitus of the included species indicate a 
distinct and natural group. Nephopteryx is very close 
to Meroptera but is distinguished by its simple, slender 
clasper, simple female genital opening, and differently 
sclerotized ductus bursae. 

Genus Nephopteryx, Species 250-267: N. sub- 
fuscella to N. celtidella 

[Males with aigrettelike palpi.] 

250. Nephopteryx suhfuscella (Ragonot), new combination 
FiauHES 337, 821 

Salebria suhfuscella Ragonot, N. Amer. Phycitidae, p. 8, 1887; 
Monograph, pt. 1, pp. 329, 360, 1893.— Hulst, Phycitidae 
of N. Amer., p. 151, 1890.— McDunnough, Check list. No. 
6123, 1939. 

Salebria semiobscurella Hulst, Phycitidae of N. Amer., p. 151, 
1890. — Ragonot, Monograph, pt. 1, p. 352, 1893. — Barnes 
and McDunnough, Contributions, vol. 3, p. 197, 1916. — 
Forbes, Cornell Mem. 68, p. 626, 1923.— McDunnough, 
Check list. No. 6212, 1939.— Craighead, U. S. Dep. Agr. 
Misc. Publ. 657, p. 454, 1950. (New synonymy.) 

Meroptera pravella (authors not Grote) Hulst (in part), Phyci- 
tidae of N. Amer., p. 148, 1890. — Ragonot (in part). Mono- 
graph, pt. 1, p. 314, 1893. — Grossbeck, Bull. Amer. Mus. 
Nat. Hist., vol. 37, p. 130, 1917.— Forbes, Cornell Mem. 68, 
p. 624, 1923. 

Color and maculation resembling those of Meroptera 
pravella. Forewing gray; the basal area contrastingly 
paler with some dark shading at extreme base and more 
or less reddish or orange scaling on base of inner margin 
(at least a trace on all except badly rubbed specimens) ; 
antemedial line obscure, indicated chiefly by its fused 
dark inner and outer borders which form a rather broad, 
oblique, blackish band, the antemedial line itself distin- 
guishable on most specimens only as an incomplete, me- 
dian, pale (whitish) streak in the dark band;subterminal 
line obscure but complete, narrow, sinuate, pale gray with 
fine dark bordering lines; some whitish dusting over 
central area of wing, especially on pale examples; discal 



dots blackish, occasionally separated, more often fused 
into a curved line; dots along terminal line fine, weak, 
blackish, usually separated, on some specimens con- 
fluent. Hind wing pale brown to smoky fuscous; veins 
darkened, especially on the darker females; a narrow 
dark shade along termen. Alar expanse, 18-22 mm. 

Male genitalia with apical process of gnathos triangu- 
late. Clasper bent downward, parallel to sm-face of 
harpe. Female genitalia with granulate sclerotized 
patch on bursa a continuous band across posterior 
ventral surface and forward on right side of bursa. 

Type locality: Not given (subfuscella, in Paris 
Mus.) ; Blanco County, Tex. (semiobscurella, in AMNH, 
ex Rutgers). 

Food plants: Rhus, locust [?]. Larva a leaf-folder. 

Distribution: United states: Maine, Augusta 
(June) ; Massachusetts, Martha's Vineyard (May, July) ; 
Connecticut, East River (June); Rhode Island, Weeka- 
paugh (Aug.); New York, Ilion (June), Rossville (Long 
Island, Feb., Mar.), Shore of Lake Ontario (near 
Rochester, May, June); Pennsylvania, Oak Station 
(Aug.); District of Columbia, Washington; North Caro- 
lina, Tryon (May); Florida, Fort Myers (May), Lake 
Alfred (July); Texas, Burnet County (Apr.), Kerrville 
(May, Aug.); Missouri, Kirkwood (Mar., Apr., May); 
Illinois, Decatur (July, Aug.); Washington, Almota 
(July), BeUingham (Nov.), PuUman (Feb., May, June, 
July, Aug., Nov.), Snake River (Jan., Feb., July), 
Walla Walla (July), Wawawai (May). Canada: Que- 
bec, Levis, Meach Lake (July), Mount St. Hilaire (July) ; 
Ontario, Merivale (Mar.), Ottawa (Mar.), Trenton 
(May, June, July). 

Many of the above records are from reared examples 
from sumac, and all such had been identified by Riley 
and Dyar as Meroptera pravella. Barnes and McDun- 
nough (1916) were the first to question and correct this 
identification, assigning the name Salebria semiobscurella 
Hulst to the sumac feeder. At that time subfuscella 
had not been recognized by American lepidopterists. 
It resembles pravella but has the reddish scaling on base 
of inner margin of forewing. Unquestionably it is the 
same as what Hulst later described as semiobscurella. 
No other American species that could have been referred 
to Meroptera or Salebria has this red-scale character 
with the habitus, otherwise, of pravella. The food 
plant record of dried peaches ("peches dess^ch^es") given 
for subfuscella by Ragonot (Monograph, pt. 1, p. 352) 
on the basis of two imperfect specimens sent him so 
labeled by Riley is obviously incorrect and can be 
ignored. I question also "locust" as a probable or even 
occasional food plant. That record, given above, is 
based on four males from the Fernald collection that 
had been identified by Dyar as pravella. They had been 
reared (Apr. 1889, May 1893) from larvae collected by 
Miss Murtfeldt at Kirkwood, Mo., on locust. Her note 
("324M") states that most of the larvae in the lot were 
"Salebria contatella Grote" but that a few seemed to be 
a different species. I suspect that the latter (the sub- 
fuscella larvae) had merely migrated to the locust after 
feeding on nearby sumac. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



125 



25]. Nephopteryx delasealU HuJst 

Figure 818 

Nephopteryx delassalis Hulst, Trans. Amer. Ent. Soc, vol. 13, 
p. 161, 1886. 

Salebria purpurella Hulst, Canadian Ent., vol. 24, p. 61, 1892. — 
Forbes, Cornell Mem. 68, p. 627, 1923. 

Salebria pudibundella Ragonot, Monograph, pt. 1, p. 350, 1893. — 
McDunnough, Check list. No. 6208, 1939. (New synonymy.) 

Myelaea delassalis (Hulst) Barnes and McDunnough, Contribu- 
tions, vol. 3, p. 198, 1916. — McDunnough, Cheek list. No. 
6225, 1939. 

Thorax vinous red with a scattered dusting of white 
and black scales. Forewing vinous red with a more or 
less diffused bluish white shading in basal area immedi- 
ately preceding inner border of the antemedial line; 
this border a broad black band (the most conspicuous 
marking on the wing), vertical, and fusing at costa with 
the outer black border; the antemedial line itself faint, 
but on most specimens its lower half distinct, narrow, 
white, slightly oblique and inwardly angulate at lower 
fold, bordered outwardly by a rather narrow, inter- 
rupted black band; some blackish dusting on the whitish 
subbasal area, black scaling along lower margin of cell, 
on some of the lower veins from cell and on costa, espe- 
cially towards apex; discal dots (when distinguishable) 
separated, red, more or less shaded with black; dots 
along termen very faint, blackish, more or less confluent. 
Hind wing pale ocherous fuscous. Alar expanse, 22- 
26 mm. 

Male genitalia similar to those oi femaldi; transtUla 
absent; apical process of gnathos narrow (not triangu- 
late); clasper short, slightly curved. Female genitalia 
with two granulate patches on bursa copulatrix, a small 
one on posterior dorsal surface, near the left side of 
bursa and a larger on ventral surface at the antero- 
lateral margin (closed end) of bursa. 

Type localities: Nevada {delassalis, in AMNH, ex 
Rutgers) ; New Mexico (purpurella, in AMNH, ex Eut- 
gers) ; Colorado (pudibundella, in Paris Mus.) . 

Food plant: Unknown. 

Distribution: United States: Arizona, White 
Mts; Colorado, Beulah (June), Fort Collins; Utah, 
Vineyard (June, July) ; Nevada; California, Inyo County 
(May, June, July). 

The types of delassalis and purpurella are both fe- 
males. Their genitalia are alike. Barnes and McDun- 
nough (1916) first noted this synonymy and the mis- 
identification of his species by Hulst (1900) and the 
consequent misapplication of the name delassalis to 
specimens oi femaldi Ragonot. The descriptions under 
delassalis in Hulst's 1900 revision and in Ragonot's 
monograph apply to femaldi and not delassalis. On 
the other hand, Ragonot's description of pudibundella 
applies in detail to the true delassalis. 

The venation of forewing is individually variable, as 
with many species of Nephopteryx, vein 10 being either 
connate or closely approximate at base to the stalk 8-9 
(rarely short stalked with it) and veins 4 and 5 either 
connate or closely approximate at base. 



252. Nephopteryx delassalis fraudlfera, new race 

Superficially appears quite distinct from delassalis; 
the entire median area and much of the basal area of 
forewing being heavUy dusted with whitish, giving the 
general ground color a whitish blue-gray shade similar 
to that of inconditella rather than the vinous red of 
typical delassalis; the vinous red limited in fraudifera 
to the thorax, extreme base of forewing, a broad band 
outwardly bordering the subterminal line, and a faint 
diffused shading just preceding it. The red shade 
somewhat darker than in typical delassalis; the black 
borders of antemedial line also somewhat broader, es- 
pecially at costa. Alar expanse, 24-26 mm. 

Male and female genitalia agreeing in all details with 
those of delassalis. 

Type locality: Oliver, British Columbia, (type in 
Canadian Nat. Coll.; paratypes in USNM, 61345, and 
Canadian Nat. Coll.). 

Food plant: Unknown. 

Described from male type and one female paratype 
from the type locality (June 8 and 9, 1923, G. B. Gar- 
rett, collector) ; and paratypes from the following locali- 
ties: "Shingle Cr. Road," Keremeos, British Columbia 
June 22, 1935, A. N. Cartrell (9); Salmon Arm, British 
Columbia, June 20, 1916, "W. R. B." (9); Kaslo, Brit- 
ish Columbia, June 13, 1903, H. G. Dyar, "19366" (9); 
Alberni, British Columbia, June 20, 1922, "W. R. B." 
(d^), and Bellingham, Wash., May 30, 1922, J. F. G. 
Clarke (d'). 

Presumably a distinct food plant as well as local race. 

253. Nephopteryx rubescentella (Hulst) 

Mineola rubescenklla Hulst, Canadian Ent., vol. 32, p. 169, 1900. 

Nephopteryx rubescentella (Hulst), U. S. Nat. Mus. Bull. 52, p. 

419, 1903.— McDunnough, Check list, No. 6173, 1939. 

Ground color of forewing slightly paler than that of 
t3T)ical delassalis. Thorax also paler, pale purplish 
gray or grayish ocherous. The dark bands bordering 
antemedial line on forewing dull red or reddish orange, 
containing no black except for occasional scattered 
scales. Maculation otherwise as in typical delassalis. 
Alar expanse, 26 mm. 

Male genitalia similar to those oi fernaldi. 

Type locality : Tennessee (type in USNM) . 

Food plant: Unknown. 

In addition to the male type, the National Collection 
contains a male from Denver, Colo. (Aug.). I have 
seen no other specimens. A female from the type 
locality will be needed before the exact status of rubes- 
centella can be determined. It may be no more than 
a race or variety oi femaldi. 

254. Nephopteryx femaldi (Ragonot), new combination 
Figures 340, 819 

Salebria fernaldi Ragonot, N. Amer. Phycitidae, p. 9, 1887. 
Salebria delassalis Hulst (not Hulst), Phycitidae of N. Amer., 

p. 154, 1890. 
Myrlaea delassalis (Hulst not Hulst) Ragonot, Monograph, pt. 1, 

p. 402, 1893. 
Myrlaea fernaldi (Ragonot) McDunnough, Check list. No. 6226, 

1939. 



126 



■UNITED STATES NATIONAL MUSEUM BULLETIH 207 



Forewing whitish ocherous with basal half of costa 
faintly washed with reddish ocherous, also some traces 
of this shading on the pale ocherous thorax and along 
inner margin of forewing from base to antemedial line, 
strongest on costa just above the dark borders of the 
antemedial line; some blackish dusting along costa near 
apex and occasionally on a few of the veins; antemedial 
line obscm"e, broken, frequently obscured entirely by 
its black borders; the latter forming a broad, strongly 
contrasted black blotch which extends from inner 
margin to top of cell (not reaching costa); discal dots 
much reduced or absent, if present more or less con- 
fluent. Hind wing pale whitish ocherous; the veins 
very slightly darkened. Alar expanse, 22-25 mm. 

Male genitalia figured from Colorado specimen identi- 
fied by Hulst as delassalis (one of his spurious "types"). 
There are no structural differences to distinguish the 
male genitalia of the two species. 

Female genitalia with a single, rather lairge, granulate 
patch in bursa, on left side and extending onto ventral 
surface near posterior end of bursa; the bursa copula- 
trix itself as broad as long, approximately round, the 
membrane at anterior end thickened (cartilaginous). 

Type locality: Arizona (type in Paris Mus.). 

Food plant: Unknown (prohsiblj Amorpha sp.). 

Distribution: United States: Arizona; Colorado, 
Denver, Glen wood Springs (July), also two males and 
a female with only the state locality, identified by Hulst 
as "delassalis" ; Kansas, Thomas County; Iowa, Ames. 
Canada: Manitoba, Cartwright (June, July), Winnipeg. 

The female genitaha and ground color of thorax and 
forewing easily separate this species from delassalis, 
with which it has been confused due to Hulst's later 
(1890) misidentification of his own species. 

255. Nephopteryx dammersi, new species 
Figures 338, 822 

Ground color pale ashy gray strongly shaded with 
pale rust red on thorax, at extreme base, on basal half 
of costal edge and on basal third of inner margin of 
forewing; more or less of this red shading on the black 
inner border of the antemedial line and forming its 
outer border near costa; an obscure, ill-defined, rusty 
blotch on middle of lower fold; antemedial line distin- 
guishable on most specimens, narrow, whitish, slightly 
oblique and indented between cell and inner margin 
(the antemedial line, where it can be distinguished for 
any appreciable distance, has a similar configuration 
and slant on most of our American Nephopteryx), 
bordered outwardly on lower half by a narrow, inter- 
rupted black line and inwardly by a broad black band, 
the latter extending only from inner margin to middle 
of cell and (as noted above) more or less shaded with 
rust red; sub terminal line obsolete or nearly so; discal 
dots usually distinct, especially the lower one, sepa- 
rated, blackish; terminal dots minute, very faint, not 



confluent. Hind wing duU white with a faint ocherous 
tint on male; pale smoky fuscous on female; the veins 
faintly darkened and a faint, narrow, dark shade along 
termen. Alar expanse, 25-26 mm. 

Male genitalia similar to those oifernaldi. There is 
some difference in the armature of the penis between the 
two species; the cornuti are somewhat shorter and 
stouter and there is a darker pigmentation of some of 
the scobinations on penis in dammersi (fig. 338); but 
there is so much individual variation of these structures 
within any given species of Nephopteryx that they can 
not be safely used to separate species. Female genitalia 
with two small and one large granulate patch in bursa. 

Type locality: Cajon Valley, San Bernardino 
Valley, Calif, (type in USNM, 61346). 

Food plant: Amorpha calijornica. 

Described from male type and three male and six 
female paratypes from the type locality (reared by 
Commander J. Dammers Apr. 15, 1933, and July 20, 
30, 1932), and one female from the Huachuca Mts., 
Ariz. ("July 8-15"). In addition I have before me a 
male from Douglas County, Ariz. (July 20, 1940, 
collected by Fritz Forbes), which appears to be con- 
specific. It is a trifle smaller (24 mm.) than the speci- 
mens of the type series. Without a matching female 
from the same locahty it appears unwise to include it 
among the paratypes. 

The species is named in honor of Commander 
Dammers, who has given many fine reared and collected 
specimens to our National Collection. It is close to 
jemaldi, but on female genitahc characters seems to be 
a distinct species and not a color form or local race. 

256. Nephopteryx dammersi floridensis, new race 

Smaller and considerably darker than typical dam- 
mersi; the ground color dark gray finely peppered with 
white especially in median and subbasal areas making 
these areas a trifle paler than remainder of wing; the 
rust red markings of typical dammersi replaced by 
lavender-red infloridensis and this color more extended, 
forming a faint suffusion over much of the median and 
outer areas in addition to the stronger markings on 
costa, inner margin and extreme base; subterminal line 
distinct and with narrow dark borders. Hind wing 
smoky fuscous; the veins darkened; a narrow blackish 
line along termen. Alar expanse, 21-22 mm. 

Genitalia: Male and female as in typical dammersi. 

Type locality: Williamsburg, Fla. (t3T)e in USNM, 
61347). 

Food plant: Amorpha herbacea. 

Described from male type from the type locahty, 
reared under S. S. No. 16970A, June 25, 1944; and one 
female paratype from Tampa, Fla., reared imder S. S. 
No. 16859, June 29, 1944. Larva collected and both 
specimens reared by members of the Special Survey of 
the Division of Foreign Plant Quarantine of the U. S. 
Bureau of Entomology and Plant Quarantine. 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



127 



257. Nephopteryr vetustella (Dyar), new combination 

Figure 820 

Salehria vetustella Dyar, Journ. New York Ent. Soc, vol. 12, 
p. 106, 1904.— Forbes, Cornell Mem. 68, p. 626, 1923. 

Myrlaea vetustella (Dyar) Barnes and McDunnough, Check list 
of the Lepidoptera of Boreal America, No. 5650, 1916. — 
Forbes, Cornell Mem. 68, p. 627, 1923.— McDunnough, 
Check list, No. 6224, 1939. 

Forewing pale ash gray with a slight purplish shading 
over outer half; basal area almost to inner dark border 
of antemedial line, Indian red or reddish orange, this 
reddish shade also on collar and top of head; antemedial 
line distinct from upper vein of cell to inner margin, 
narrow, oblique, inwardly notched between cell and 
inner margin, bordered inwardly by a broad black band 
and outwardly by a narrow black line, the black borders 
continuous to and fused at costa; sub terminal line 
usually distinct but faint and with a weak dark inner 
bordering line, sinuate; discal dots separated, small, 
blackish; a weak row of blackish dots along termen. 
Hind wing pale smoky brown; the veins and terminal 
margin slightly darkened. Alar expanse, 22-25 mm. 

Male genitalia similar to those o{ Jemaldi. Female 
genitalia with a single long granidate patch in bursa 
extending nearly the length of the bursa on its left side 
(seen from below) and curving onto ventral surface near 
junction of bursa and ductus bursae. 

Type locality: Plummers Isl., Md. (type in USNM) . 

Food plant: Unknown {Amorpha sp. ?). 

Distribution: United States: Maryland, Hum- 
mers Isl. (Apr., May); Pennsylvania, Oak Station 
(June), Pittsburgh (May, June) ; New York, Dion (May) ; 
j/^inois, Edgebrook (June), Elkhart (Aug.), Palos Park 
(June); Iowa, Iowa City (June), Sioux City (July); 
Missouri, St. Louis; Florida, St. Petersburg (June). 
Canada: Ontario, Ottawa (June, July) ; Qiiebec, Meach 
Lake (June). 

This species forms one of a group of closely related 
species (delassalis to vetustella) that feed upon Amorpha 
and have similar male genitalia but differ markedly in 
the number, arrangement, and relative sizes of the gran- 
ulate patches of the female bm-sa. These differences 
appear to be constant and the specific character holds 
even in series of individuals from widely different 
localities. The male genitalia exhibit some minor dif- 
ferences, especially in the relative size and position of the 
two cornuti ; but here individual variation makes the 
apparent differences untrustworthy for specific differ- 
entiation. During dissection of the male organs a 
simple transtilla can be faintly seen in all the species of 
the group, but it is so weakly sclerotized that it is not 
visible or but partially distinguishable in balsam mounts. 
The structure is obsolescent and cannot be considered 
"present" in the sense in which it is in Meroptera for 
example or even in a few other species of Nephopteryx. 

258. Nephopteryx inconditella (Ragonot), new combination 

Figure 825 

Salebria contatella inconditella Ragonot, Monograph, pt. 1, pp. 
348, 350, 1893.— McDunnough, Check Uat, No. 6205b, 1939. 



Thorax and forewing pale ash gray with a faint bluish 
tint; some reddish scaling at extreme base of wing and a 
small spot of the same shade at middle of lower fold; 
pale antemedial line distinguishable from inner margin 
to cell, deeply notched at vein lb, its black borders 
complete from inner margin to costa, the inner border 
broad and fusing with the narrow outer one near costa; 
subterminal line distinct, sinuate-serrate, narrowly bor- 
dered by obscure blackish lines; discal spots obscure but 
usually distinguishable, separated; terminal dots weak, 
confluent; on most specimens a narrow, dark transverse 
shade extending from costa at inner edge of subterminal 
line to middle of inner margin. Hind wings translucent, 
whitish with a faint ocherous tint, the veins slightly 
darkened; a narrow brownish shade along terminal 
margin. Alar expanse, 25-30 mm. 

Male genitalia of the jemaldi type but clasper some- 
what more strongly sclerotized than in the seven preced- 
ing species. Female genitalia with a single, small, 
round, granulate patch on ventral surface of bursa very 
near junction of bursa and ductus bursae. 

Type locality: Colorado (type in Paris Mus.) . 

Food plant: Unknown. 

Distribution: Colorado; Arizona, Huachuca Mts., 
Palmerlee, and four Arizona specimens with only the 
state locahty and without dates, one of these a pseudo- 
type of "Pinipestis albovittella Hulst." 

The species was described as a pale western variety of 
contatella (Grote), to which it is apparently most nearly 
related and which, except for its paler ground color, it 
resembles. Its female genitalia however clearly indi- 
cate a distinct species. The ground color of forewiug 
is intermediate between that of suhcaesiella {= conta- 
tella) and that of dammersi. 

259. Nephopteryx suhcaesiella (Clemens), new combination 
FiQUBBS 339, 826 

Pempelia suhcaesiella Clemens, Proc. Acad. Nat. Sci. Philadel- 
phia, p. 206, 1860. 

Pempelia contatella Grote, North Amer. Ent., vol. 1, p. 49, 1880. — 
Comstock, in Rep. [U. S.] Comm. Agr. for 1880, p. 261, 1881 
(in part) . 

Salebria contatella (Grote), Bull. U. S. Geol. Geogr. Surv. Terr., 
vol. 6, no. 3, p. 590, 1882. — BeutenmOller, Canadian Ent. 
vol 22, p. 16, 1890 (larva). — Hulst, Phycitidae of N. Amer., 
p. 152, 1890; U. S. Nat. Mus. Bull. 52, p. 424, 1903.— 
Packard, U. S. Dep. Agr. Fifth Rep. Ent. Comm., p. 361, 
1890.— Ragonot, Monograph, pt. 1, p. 348, 1893.— Forbes, 
Cornell Mem. 68, p. 626, 1923. — McDunnough, Canadian 
Ent., vol. 78, p. 109, 1946. 

Salebria virgatella suhcaesiella (Clemens) Barnes and McDun- 
nough, Check list of the Lepidoptera of Boreal America, No. 
5631a, 1916— McDunnough, Check list, No. 6205a, 1939.— 
Craighead, U. S. Dep. Agr. Misc. Publ. 657, p. 453, 1950. 

Color and maculation similar to those of inconditella 
but the ground color of forewing a much darker gray, 
basal area contrastingly paler gray; the reddish scaling 
on base of wing ranging from wine to rusty ocherous, 
always present but sometimes reduced to a few scales, 
rarely extended for a short distance onto the thorax; 
inner black border of antemedial line somewhat broader 
and more diffused than on inconditella, complete to costa 



128 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



(unbroken at any part of its length) ; following the ante- 
medial line a pale grayish spot (sometimes very faint but 
usually more contrasted than in inconditeUa) ; discal 
spots distinct, black, separated. Hind wing smoky 
white to smoky fuscous or brown, darker on females 
than on males, a distinct dark shade along termen. 
Alar expanse, 21-28 mmi 

Male genitalia as in inconditeUa and virgatella except 
for the armature of the penis. In both inconditeUa and 
virgatella there are two moderately stout cornuti situ- 
ated on penis, one to the side and slightly behind the 
other. In suhcaesieUa the second cornutus is greatly 
reduced. McDunnough (1946) states that there is only 
one cornutus in suhcaesieUa (contaiella) . That could 
easUy be the case on individual specimens, but the 
normal condition is two comuti on penis. Every prep- 
aration I have seen shows at least a vestige of the 
second cornutus. In all three species the lateral ele- 
ments of transtiUa are indicated, but very weakly 
sclerotized. Female genitalia without granulate patch 
or patches on bursa. 

Type localities: Not given, presumably Pennsyl- 
vania {suhcaesieUa, in Acad. Nat. Sci. Philadelphia); 
"New England" {contateUa, in BM). 

Food plant: Robinia pseudoaca^ia. Wisteria also 
recorded as food plant. 

Distribution: United States: Maine, Augusta 
(May, June), Orono; iVew Hampshire, Hampton (July) ; 
Massachusetts, Amherst (June), Framingham (May), 
Martha's Vineyard (July, Aug.), Newton Highlands; 
New Jersey, Essex County Park (June), New Lisbon 
(Aug.) ; Pennsylvania, New Brighton (May, July, Aug.), 
Oak Station (June), Pittsburgh (May, June, July); 
Maryland, Plummers Isl. (June, Aug.), Somerset Heights 
(Aug.) ; District of Columbia, Washington (Apr., May, 
July); Virginia, Falls Church (Aug.), Snickers Gap 
(July); North Carolina, Black Mountain (July), Tryon 
(May) ; Tennessee, no specific locality (May) ; Illinois, 
Decatur (July), Elkhart, Oconee (July); Iowa, Iowa 
City (Aug.), Sioux City (June); Missouri, "Cent. Mo." 
(Aug.), Kirkwood (Apr., May), St. Louis (June); Ar- 
kansas, Washington County ("July-Aug."). Canada: 
Nova Scotia, Smith's Cove (recorded by McDunnough, 



I have not seen any Canadian examples but Mc- 
Dunnough's description leaves no doubt of what he had. 
He notes the differences in the cornuti and larval 
characters between suhcaesieUa (contateUa) and virgatella 
quinquepunctella) and treats them as a distinct species. 
The difference in their female genitalia and the con- 
sistent difference in maculation of forewing are added 
evidence that they are not merely races of one variable 
species. 

260. Nephopter3rx virgatella (Clemens), new combination 

Figure 827 

Pempelia virgatella Clemens, Proc. Acad. Nat. Sci. Philadelphia, 

p. 205, 1860. 
Pempelia contateUa quinquepunctella Grote, North Amer. Ent., 

vol. 1, p. 50, 1880.— Comstock, in Rep. [U. S.] Comm. Agr. 

for 1880, pp. 261-262, 1881 (part; larva). 



Salebria contateUa quinquepunctella (Grote) Hulst, Phycitidae of 
North Amer., p. 152, 1890; U. S. Nat. Mus. Bull. 52, p. 424, 
1903. — Ragonot, Monograph, pt. 1, p. 348, 1893. — Forbes, 
Cornell Mem. 68, p. 626, 1923. 

Salebria virgatella (Clemens) Barnes and McDunnough, Check 
list of the Lepidoptera of Boreal America, No. 5631, 1916. — 
McDunnough, Check list. No. 6205, 1939. 

Salebria quinquepunctella (Grote) McDunnough, Canadian Ent., 
vol. 78, p. 109, 1946 (larva). 

Ground color of forewing (on most specimens) a 
trifle paler than that of suhcaesieUa, gray with a faint 
brownish or purplish tint; a reddish (or pale purplish) 
brown shade along lower fold, cutting the antemedial 
line and its black borders; a similar, narrower, shorter 
streak on median fold; outer black border of ante- 
medial line between top of cell and inner margin reduced 
to two black dots, one on the lower margin of cell, the 
other on vein lb and enclosed within the pale patch 
following the antemedial line; both these dots and the 
black discal dots at end of cell well contrasted; sub- 
terminal line faint, not appreciably darkly bordered, 
interrupted at the folds. Hind wing smoky white to 
brown; the veins more or less darkened and a smoky 
brown shade along termen. Alar expanse, 22-26 mm. 

Male genitalia similar to those of inconditeUa; both 
cornuti moderately stout and situated one to the side 
of and slightly behind the other. Female genitalia 
with two strong granulate patches on bursa, a rather 
large one on middle of dorsal surface cm'ving around 
left side onto ventral surface, and a smaller ventral 
patch near junction of bursa and ductus bursae. 

Type localities: Not given, presumably Pennsyl- 
vania (virgatella, in Acad. Nat. Sci. Philadelphia) ; New 
York (quinquepunctella, in BM). 

Food plant: Bohinia pseudoacacia. 

Distribution: UnitedStates: Maine; Massachusetts, 
Amherst (June), Martha's Vineyard (June, July, Aug.); 
New York, Orient (Long Island, Aug., Sept.) and one 
specimen with only state locality; New Jersey, Essex 
County Park (Aug.) ; Pennsylvania, Buena Vista (Aug.) ; 
New Brighton (May, July); West Virginia, Jefferson 
County (Aug.); District of Columbia, Washington 
(June); Virginia, BerryviUe (May); North Carolina, 
Tryon; Illinois, Elkhart, Putnam Coimty (May); Mis- 
souri, St. Louis; Arkansas, Washington County ("July- 
Aug."). Canada: Ontario, London, Trenton (June); 
Nova Scotia, Bridgetown (July), Smith's Cove. 

The differences separating virgatella from suhcaesieUa, 
of which it was long considered only a variety, are dis- 
cussed under the latter species. 

261. Nephopteryx cameella Hulst 
Figure 823 

Nephopteryx cameella Hulst, Ent. Amer., vol. 3, p. 131, 1887. 

Nephopteryx inquilinella Ragonot, N. Amer. Phycitidae, p. 8, 
1887; Monograph, pt. 1, p. 290, 1893.— Hulst, Ent. Amer., 
vol. 5, p. 156, 1889; Phycitidae of N. Amer., p. 145, 1890.— 
McDunnough, Check list, No. 6171, 1939. 

Salebria cameella (Hulst), Phycitidae of N. Amer., p. 153, 1890. — 
Ragonot, Monograph, pt. 1, p. 367, 1893. — Barnes and Mc- 
Dunnough, Contributions, vol. 3, p. 197, 1916. — Forbes, 
Cornell Mem. 68, p. 627, 1923. — McDunnough, Check list, 
No. 6204, 1939. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 



129 



Ground color of forewing bluish gray, the gray shade 
most obvious in median area and as a narrow band 
preceding the inner border of the antemedial line; base 
maroon red, this shade more or less suffusing the gray 
thorax; antemedial line obscure, bordered inwardly by 
a broad madder-red band which extends to costa; a 
similar red shade in outer area bordering the faint 
sinuate sub terminal line and some red suffusion over 
median part of the lower fold; discal spots distinct, 
separated, black; terminal dots very faint more or less 
confluent. Hind wing smoky white with a faint yellow- 
ish tint; somewhat darker on females; a fine brown line 
along termen. Alar expanse, 20-23 mm. 

Male genitalia of the femaldi type; the two cornuti 
lie side by side, one slightly shorter and more slender 
than the other. Female genitalia with bursa and re- 
mainder of genitalia considerably smaller than in pre- 
ceding species; two granulate patches, one large, one 
considerably smaller, placed opposite each other on 
lateral margins of the bursa. 

Type localities: Maine [?] {carneella, ia AMNH, 
ex Rutgers; the male type bears no locality label, but 
in his original description Hulst gives New Mexico as 
the type locality; this, as pointed out by Barnes and 
McDunnough, is probably an error or pure guesswork 
on Hulst's part) ; Wisconsin {inguilinella, in Paris Mus.). 

Food plant: Salix (carneella) ; galls of sawfly (Euura 
Salicisnodum) on willow. 

Distribution: United States: Maine, Monmouth 
(July), and two examples with only state locality; Mas- 
sachusetts, Amherst (June), Martha's Vineyard (Apr.); 
Indiana, Hessville (June); Wisconsin; Michigan, Dick- 
inson County. Canada: Ontario, Ottawa (June), 
Trenton (July); Manitoba, Aweme (May). 

Hulst was correct the first time (1889) in making 
inquilinella a synonym of his carneella. The genitalia 
of their male types are identical. 

262. Nephopteryx basilaris Zeller 
Figures 341, 829 

Nephopteryx basilaris Zeller, Verb, zool.-bot Ges. Wein, vol. 22, 
p. 548, 1872.— Grote, N. Amer. Ent., vol. 1, p. 51, 1880.— 
Hulst, N. Amer. Phycitidae, p. 145, 1890. 

Salebria basilaris (Zeller) Ragonot, Monograph, pt. 1. p. 353, 
1893.— Forbes, Cornell Mem. 68, p. 627, 1923.— McDun- 
nough, Check list, No. 6209, 1939. 

Forewing pale ash gray to dark gray with a faint 
bluish tint; basal area pale wood brown or pale orange, 
a black shade along its inner margin expanding upward 
at antemedial line to middle of cell; the lower fold in 
basal area streaked with red or reddish brown with 
some scattering of similarly colored scales on the pale 
area above; antemedial line distinct to top of cell, well 
contrasted, grayish white, nearly vertical, inwardly 
dentate between cell and inner margin; margined out- 
wardly by a narrow black line; subterminal line faint 
but distinguishable, sinuate-dentate, followed in outer 
area (on some specimens) by a broad reddish shade; 
discal dots obscured ; a row of small blackish dots along 
terminal margin. Hind wings white with a faint ocher- 

300329 — 56 10 



ous or smoky tint; the veins very faintly darkened; a 
thin brownish line along termen. Alar expanse, 24-27 
mm. 

Male genitalia resembling those of subcaesiella; the 
clasper and lateral elements of transtilla somewhat 
stronger; the smaller cornutus on penis longer and 
stouter but also on some specimens reduced to a mere 
vestige. Female genitalia with two granulate patches 
on ventral surface of bursa, a weaker granulation of the 
surface connecting them. 

Type locality: Massachusetts (type in BM). 

Food plant: Unknown. 

Distribution: United States: Maine, Augusta 
(June), Orono; New Hampshire, Hampton (June); Mas- 
sachusetts, Martha's Vineyard (July), Wilmington 
(June); New York, CatskiU Mts., Ilion (June); New 
Jersey, Newark; Illinois, Lacon (June); Indiana, Hess- 
ville (May, July) ; Michigan, one example, state locality 
only; Colorado, Fort Collins (July), and one example 
state locality only; Utah, Provo Canyon (July). Can- 
ada: Ontario, Budbury, Hymers (July), Trenton (July); 
Manitoba, Aweme (June), McCreary, Winnipeg. 

A strikingly marked species, the most easily identified 
in the genus. The maxiUary palpus of the male is, as 
noted by Ragonot (Monograph, p. 354), not strictly in 
the form of an aigrette. The scales are moderately 
long and slender, but not hairlike, intermediate between 
those of a squamous and a typical aigrettelike palpus. 
The labial palpus is grooved on inner surface of the 
second segment as in most species having maxillary 
palpi of the aigrette type. 

263. Nephopteryx termi talis (Hulst), new combination 
Figures 342, 828 

Pempelia termitalis Hulst, Trans. Amer. Ent. Soc, vol 13, p. 162, 

1886. 
Salebria termitalis (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 

1889.— Hulst, Phycitidae of N. Amer., p. 153, 1890. 
Salebria levigatella Hulst, Canadian Ent., vol. 24, p. 61, 1892. — 

Forbes, Cornell Mem. 68, p. 626, 1923. — McDunnough, 

Check list, No. 6207, 1939. (New synonymy.) 
Myrlaea termitalis (Hulst) Ragonot, Monograph, pt. 1, p. 401, 

1893.— McDunnough, Check list. No. 6223, 1939. 

Forewing dark suffused gray (darker than on basilaris) ; 
basal area duU reddish orange with little black on most 
specimens except along costal edge; on others some 
black scaling at extreme base and, narrowly, along 
inner margin; antemedial line nearly obliterated by its 
black borders which are more or less fused and from a 
broad, nearly vertical band from inner margin to costa, 
not strongly contrasted against the dark ground color 
of the wing; subterminal line, discal and terminal dots 
obscure. 

Thorax dark gray, on some specimens more or less 
suffused by the orange color of the basal area of fore- 
wing. One specimen before me (a male from Inyo 
Coimty, Calif.) has the entire thorax and base of fore- 
wing to the black inner border of the antemedial line a 
dull madder-red, and the outer third of wing faintly 
suffused with the same reddish shade. Hind wing dull 



130 



ITNITED STATES NATIONAL MUSEUM BULLETIN 207 



ocherous white to smoky white. Alar expanse, 23-27 
mm. 

Alar expanse, 23-27 mm. 

Male genitalia with apical process of gnathos tri- 
angulate. Transtilla weak but distinguishable. Clasper 
erect and slightly curved, slender, digitate. One 
moderately sized and one much smaller cornutus on 
penis. Female genitaha with a single moderately large 
granulate patch on posterior lateral corner of bursa and 
extending in weaker granulation transversely across the 
middle of the lower surface. 

Type localities: Colorado {termitalis, in AMNH, 
ex Rutgers); "Amherst, Massachusetts" [sic] (leviga- 
tella, in AMNH, ex Rutgers). 

Food plant: Unknown. 

Distribution: United States: Colorado, Glenwood 
Springs, Gunnison County near Altmont, and two 
examples (cf , 9) with only the state locality; Utah, 
Spanish Fork (July); Arizona, Prescott (June); Cali- 
Jornia, Inyo Coimty (Jime), Placer County (June). 
Canada: Ontario, Trenton (July), Manitoba, Winni- 
peg; Alberta, Bilby (June); British Columbia, Clinton 
(June). 

Hulst also gives Massachusetts and Wisconsin as 
localities for his levigatella. His type of the latter how- 
ever has no locality label. A female of levigatella from 
the Fernald Collection is in the National Collection. 
It also bears a Hulst "t3rpe" label, but no locality or 
date. The type of termitalis is a male, not a female as 
given in Hulst's original description. Genitalia of 
conspecific females from western locaUties agree in every 
detail with those of the type of levigatella so there can 
be no question of the synonymy of the two Hulst names. 

The labial palpi appear a trifle oblique (less tightly 
appressed to the face than on most of the preceding 
species) which may account for Ragonot's reference of 
the species to Myrlaea. 

264. Nephopteryx termitalis yuconella (Dyar), new status 

Salehria yuconella Dyar, Ins. Insc. Menstr., vol. 13, p. 12, 1925. — 
McDunnough, Check list, No. 6194, 1939. 

A slightly darker more suffused local race of termitalis; 
the basal area of forewing almost entirely suffused with 
blackish except for a narrow grayish white shade along 
inner margin of the black inner border of the antemedial 
line; no appreciable red or orange shading anjrwhere on 
forewing or thorax. Alar expanse, 25-27 mm. 

Genitalia: Male and female as in typical termitalis. 

Type locality: Near Fort Yukon, Alaska (type in 
USNM). 

Food plant: Unknown. 

Distribution: Alaska: Dawson (June), Fort Yukon. 

Specimens of yuconella as of typical termitalis show 
a strong tendency to become greasy with age, indicating 
that their larvae are borers; but nothing is known about 
the biology of either form. 



265. Nephopteryx bifasciella Hulst 
FiGtTBEs 343, 831 

Nephopteryx bifasciella Hulst, Ent. Amer., vol. 3, p. 132, 1887. 

Salehria bifasciella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 366, 1893. — Hulst, Phycitidae 
of N. Amer., p. 154, 1890. — Barnes and McDunnough, 
Contributions, vol. 3, p. 197, 1916. — McDunnough, Check 
list, No. 6214, 1939. 

Salehria nogalesella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 35, 1905.— McDunnough, Check list, No. 6215, 1939. 
(New synonymy.) 

Forewing ash gray with a fine powdering of black 
scales, giving the wing a faint pale bluish tint; the black 
borders of the transverse lines strongly contrasted; 
antemedial line distinct, at least from below upper vein 
of cell, narrow, slightly oblique and somewhat angulate 
at middle, its outer black border complete, slightly 
broadened at costa, its inner black border narrow, 
extending from inner margin only to top of cell; sub- 
terminal line sinuate, bordered iawardly by a narrow 
black line, and outwardly by a much fainter, paler 
dark line; discal dots usually distinguishable but faint, 
separate or confluent (sometimes both ways on the 
same specimen); dots along terminal margin very 
faint, more or less confluent. Hind wing white with a 
very faint ocherous or smoky tint, slightly darker on 
female than on male. Alar expanse, 20-22 mm. 

Male genitalia with apical process of gnathos tri- 
angulate. Clasper sharply curved and running close 
and parallel to surface of harpe, simple and strongly 
sclerotized. Cornuti rather short, but individually 
variable in size. There is also some slight variability 
in the size and shape of the apical process of gnathos. 
The genitalia of the type of bija^ciella and those of its 
synonym nogalesella are more nearly ahke than those 
of any other two males before me. Female genitaha 
with bursa nearly round, armed with two large granulate 
patches, arranged as in figure 831 but with the position 
of the anterior patch (at closed end of bursa) somewhat 
variable. In one specimen from Palmerlee there is a 
third patch on the right side of bursa and the bursa 
itself is narrower and considerably elongated. These 
differences probably represent nothkig more than 
individual aberrations. Males from the same locality 
are normal. 

Type localities: Arizona (bijasciella, in AMNH, 
ex Rutgers); Nogales, Ariz, (nogalesella, in USNM). 

Food plant: Unknown. 

Distribution: Arizona, Baboquivari Mts. (July), 
Huachuca Mts. (July, Aug.), Nogales (July), Palmerlee, 
Redington, "Southern Arizona" (Aug.), and two ex- 
amples with only the state locahty. 

266. Nephopteryx uviiiella (Ragonot), new combination 

FiGTJBEs 344, 824 

Meroptera uvinella Ragonot, N. Amer. Phycitidae, p. 8, 1887; 
Monograph, pt. 1, p. 315, 1893. — Hulst, Phycitidae of N. 
Amer., p. 148, 1890. — McDunnough, Check list. No. 6183, 
1939. 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITmAE 



131 



Salebria afflictella Hulst, Canadian Ent., vol. 32, p. 170, 1900. — 
Forbes, Cornell Mem. 68, p. 625, 1923.— McDunnough, 
Check list. No. 6200, 1939. (New synonymy.) 

Meroptera liquidambarella Dyar, Proc. Ent. Soc. Washington, 
vol. 6, p. 108, 1904. 

Meroptera afflictella (Hulst) Dyar, Proc. Ent. Soc. Washington, 
vol. 7, p. 34, 1905. 

Forewing fuscous (gray-brown), the median area 
heavily dusted with white giving it a pale ash gray 
color; basal area of the ground color, rarely with some 
red scaling in the median fold; dark base followed by 
an obHque, straight, whitish band ; this latter bordered 
outwardly by a broad, somewhat diffused, dark brown 
or blackish band through which may be distinguished 
faint white traces of the true antemedian line; sub ter- 
minal line with a very slight central bulge, rarely some- 
what crenulate, whitish gray, bordered inwardly by a 
diffused brown shade and outwardly by a narrow brown 
line; discal dots more or less fused; blackish terminal 
dots confluent. Hind wings pale to dark smoky fus- 
cous. Alar expanse, 14-18 mm. 

Male genitalia distinguished chiefly by the armature 
of the penis. On the paratype of afflictella from Mont- 
clair, N. J., there appears to be two cornuti, one closely 
appressed to the other; but in other preparations before 
me (including the types of uvinella and liquidambarella) 
the two cornuti are completely fused into a single rather 
short and stout, longitudinally ribbed cornutus. The 
clasper is a slender, curved, sharply pointed, smooth, 
sclerotized hook. Vestiges of a divided transtiUa 
distinguishable on most preparations. 

Female genitalia small (approximately the size and 
form of those of carneella) ; bursa with two moderately 
sized granulate patches, the bursal lobe giving off the 
ductus seminalis also granulate and partially (smoothly) 
sclerotized ; this lobe as usual arises from the dorsum of 
bursa but is deflected to the left. 

Type localities: United States {uvinella, in USNM) 
Elizabeth, N. J. {afflictella, in AMNH, ex Rutgers); 
Washington, D. C, {liquidambarella, in USNM). 

Food plant: Liquidambar styraciUua (sweetgum). 
Larva a leaf-tier. 

Distribution: Connecticut, East River (May), Stam- 
ford (Aug.); New Jersey, Elizabeth (Aug.), Lakeland 
(May), Montclair (June, Aug.), New Lisbon (June); 
District oj Columbia, Washington (May, Aug.) ; Virginia, 
Skyland (July) ; North Carolina, Greensboro (Aug.) ; 
Georgia, Savannah (Sept.); Florida, Lakeland (May, 
June) . 

Ragonot's uvinella has been an unknown entity ever 
since its description and was suspected of being only a 
variety of Meroptera pravella. Fortunately the type 
was secured by Dr. Barnes when the Oberthiir Collec- 
tion was sold. It lacks one forewing but is otherwise 
intact and its habitus and genitalia leave no doubt that 
it is the same as the sweet-gum feeder described by 
Hulst and Dyar. The type of afflictella is a female. 
The types of uvinella and liquidambarella are both males. 
The genitalia of all of them are before me. 



267. Nephopteryx celtidella (Hulst), new combination 
FiGUBEs 345, 830 

Salebria celtidella Hulst, Phycitidae of N. Amer., p. 155, 1890. — 
Beutenmiiller, Canadian Ent., vol. 22, p. 17, 1890 (larva). — 
Ragonot, Monograph, pt. 1, p. 354, 1893. — Forbes, Cornell 
Mem. 68, p. 626, 1923.— McDunnough, Check list, No. 
6206, 1939. 

Ground color ocherous (clay color) more or less shaded 
with fuscous gray over submedian and (on especially 
dark females) outer basal areas; the ground color espe- 
cially well contrasted at base and in the central area 
about the discal spots, also on thorax, at extreme base 
of wing and on thorax sometimes of a tawny shade ; ante- 
medial line far out towards middle of wing, oblique, sinu- 
ate, faint, indicated chiefly by its narrow black borders 
which are more or less broken into dots on the veins; 
subterminal line sinuate-serrate, bordered inwardly by 
a row of black (somewhat confluent) dots and outwardly 
by a row of small black wedges on the veins ; discal dots 
distinct, well separated, black; a row of distinct black 
dots along termen. Hind wing pale to rather dark 
smoky fuscous. Alar expanse, 18-22 mm. 

Male genitalia with apical process of gnathos smaller 
than that of preceding species (not triangulate). Clasper 
rather short, bent across surface of harpe, blunt. Sclero- 
tized lateral elements of transtiUa distinguishable, 
rather long, slender. 

Female genitalia with a round, moderately large 
granulate patch on ventral surface of bursa and a smaller 
patch near junction of bursa and ductus bursae and the 
base of the lobe giving off the ductus seminahs. 

Type locality: New York (in AMNH, ex Rutgers). 

Food plant: Celtis. Larva a leaf-tier. 

Distribution: United States: New York, Long 
Island; Maryland, Plummers Isl. (May, July); Flor- 
ida, Palm Beach (Feb.); Texas, Brownsville (July), Vic- 
toria (May), ZavaUa County {kpv.); Mississippi, "Agr. 
CoUege" (Apr.), Starkville (Jifly); Missouri, St. Louis 
(Aug.); Illinois, Oconee (Aug.). 

A distinct species distinguished from any of the spe- 
cies with aigrettelike maxillar palpi on the male by its 
clay-colored, black-mottled forewings. Its habitus is 
nearest to that of rubrisparsella in the group with 
squamous, male maxiUary palpi. 

Genus Nephopteryx, Species 268-271: A'', rubri- 
sparsella to N. bisra 

[Males with squamousmaxillary palpi.] 

268. Nephopteryx rubrisparsella (Ragonot) 
Figures 346, 832 

Pristrophora rubrisparsella Ragonot, N. Amer. Phycitidae, p. 6, 
1887. 

Pristophora rufibasella Ragonot, N. Amer. Phycitidae, p. 7, 1887. 

Sciota croceella Hulst, Ent. Amer., vol. 4, p. 115, 1888. 

Nephopteryx rubrisparsella (Ragonot), Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 284, 1893.— Hulst, Phycitidae 
of N. Amer., p. 145, 1890.— Forbes, Cornell Mem. 68, p. 623, 
1923.— McDunnough, Check list. No. 6169, 1939. 



132 



msriTED STATES NATIONAL MUSEUM BULLETIN 2 07 



Psorosa texanella Hulst, Canadian Ent., vol. 32, p. 174, 1900. 
Hulstea texanella (Hulst), U, S. Nat. Mus. BuU. 52, p. 432, 1903.— 
McDunnough, Check list, No. 6342, 1939. (New synonymy.) 

Similar to celtidella in color and maculation except: 
Ocherous basal area of forewing more or less shaded with 
reddish purple and similar suffusions over much of the 
median and outer areas; also a broad blackish suffusion 
over the area of the antemedial line and extending some- 
what beyond it; the inner dark border of subtermiaal 
line more nearly continuous and the outer bordering 
dashes much fainter except in dark and very well 
marked examples. Alar expanse, 17-20 mm. 

Male genitalia figured from type of texanella Hulst, 
which agree in every detail with those of typical rubri- 
sfarsella and of the male type of croceella. They differ 
from those of celtidella in having a longer, more strongly 
sclerotized clasper, and different cornuti (shown in lat- 
eral view in our figxu-es). The cornuti lie side by side 
in both celtidella and rubrisparsella, but in the former 
species they are of equal length while in rubrisparsella, 
one cornutus is much shorter than the other — ^not too 
reliable a character, but apparently consistent here. 
Transtilla, on dissection, distinguishable as a complete 
band, but central area very weak and in balsam prepa- 
ration only the well sclerotized lateral elements easily 
seen. 

Female genitalia with a broad granulate band par- 
tially encircling middle of bursa, similar to that in sub- 
fv^cella except that in subjuscella it is chiefly over the 
ventral surface of bursa while m. rubrisparsella it is on 
the dorsal. 

Type localities: United States {rubrisparsella, in 
Paris Mus.); Texas (rufibasella, in Paris Mus.); Blanco 
Coimty, Tex. {croceella and texanella, in AMNH, ex 
Eutgers). 

Food plant: Celtis. 

Distribution: Texas, Black Jack Springs, Blanco 
County, KerrvHle (June); Oklahoma; Missouri, St. 
Louis (June); Illinois, Lacon (July), Putnam County 
(July); West Virginia, Jefferson County (Aug.); Mary- 
land, Plummers Isl.; District of Columbia, Washington 
(May, June). 

Very close to celtidella and distinguished from it 
chiefly by its genitaUa and squamous male maxillary 
palpi. Hulst's texanella was described from an ab- 
normal specimen with vein 4 lacking in the hind wing. 
I have before me two similar abnormal females (from 
Oklahoma and Illinois) and in the following species 
{gilvibasella) a couple of examples, out of a long series 
of normal specimens, that also have vein 4 absent. 

The type of croceella in addition to the Hulst name 
label also bears the following label in Ragonot's hand- 
writing: "Nephopteryx rubrisparsella Ti&g.=rwfibasella 
Rag." 

269. Nephopteryx gilvibasella Hulst 
Figure 836 

Nephopteryx gilvibasella Hulst, Phycitidae of N. Amer., p. 145, 
1890. — Ragonot, Monograph, pt. 1, p. 289, 1893.— McDun- 
nough, Check list, No. 6172, 1939. 



Salebria lacteella Hulst, Canadian Ent., vol. 32, p. 71, 1900. — 
Barnes and McDunnough, Contributions, vol. 3, p. 197, 
1916. 

Much paler than the preceding species {rubrisparsella) 
and without its blackish shadings. Ground color pale 
gray with a very faint bluish tint; extreme base of wing 
ocherous orange, this shade extended somewhat on 
inner margin and indicated on many specimens in 
median area over the lower fold; a broad band of the 
same color immediately preceding the antemedial line 
and extending from inner margin to middle of cell; 
above it a blackish shade extending to costa; lower half 
of antemedial line clearly indicated, narrow, whitish, 
bordered outwardly by a narrow black line which fuses 
towards costa into the blackish shade above the ocher- 
ous patch; subterminal line faintly bordered inwardly 
by a faint blackish line; discal dots small, separated, 
sometimes obscure, but usually distinct; a row of weak 
but discernible blackish dots along termen. Hind wing 
subpeUucid; whitish with a very faint ocherous tint; a 
weak, pale brown Une along tei-men; the veins not 
appreciably darkened. Alar expanse, 17-20 mm. 

Male genitalia similar to those of rubrisparsella. The 
eighth-segment tufts much simpler, reduced to two 
lateral pairs with scales of a uniform shape and size. 

Female genitalia with a broad granulate band on 
biu-sa similar to that in rubrisparsella; bursa bulged into 
a small, strongly granulate lobe at its left posterior 
angle (this lobe opposite to that giving off ductus 
seminalis) ; also a narrow row of moderately long spines 
across upper side of bursa at junction of bursa and 
ductus bursae. 

Ttpe localities: Not given {gUvibaseUa, in AMNH, 
ex Rutgers) ; Blanco Coimty, Tex. {lacteella, in AMNH, 
ex Rutgers). 

Food plant: Unknown. 

Distbibution: Texas, Blanco County (Sept.), 
Brownsville (Mar., July), San Benito (Mar., Apr., 
May, July, Aug., Sept.). 

The species is close to but distinct from rubrisparsella. 
The types of gilvibasella and lacteella are both females 
with identical genitalia. A long series from Browns- 
ville and San Benito are before me. 

270. Nephopteryx erassifasciella Ragonot 
Figures 347, 835 

Nephopteryx erassifasciella Ragonot, N. Amer. Phycitidae, p. 8, 

1887; Monograph, pt. 1, p. 285, 1893. — Hulst, Phycitidae 

of N. Amer., p. 146, 1890. — McDunnough, Check list, No. 

6170, 1939. 
Nephopteryx decipientella Dyar, Proc. Ent. Soc. Washington, vol. 

7, p. 34, 1905.— McDunnough, Check list. No. 6162, 1939. 

(New synonymy.) 
Nephopteryx crataegella Barnes and McDunnough, Contributions, 

vol. 3, p. 222, 1917.— McDunnough, Check list. No. 6164, 

1939. (New synonymy.) 

Forewing ash gray more or less suffused by somber 
purplish or gray brown, shading into blackish brown on 
the borders of the transverse liaes; the pale color limited 
to a rather narrow band preceding the inner border of 
the antemedial line, part of the median area following 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



133 



the antemedial line and siurounding the discal dots, 
and some faint pale dusting immediately preceding 
terminal margin; extreme base of wing pm-plish brown 
with a shading (in some specimens) of reddish scales, 
especially towards inner margin; antemedial line pale 
gray, distinguishable only from inner margin to middle 
of cell, bordered inwardly by a broad, vertical, blackish 
brown band which extends to costa, and outwardly by 
a narrow black line which fuses into the inner dark 
border towards costa; subterminal line sinuate, paJe 
gray with narrow, black-brown borders; discal dots 
distinct, sometimes partially fused. Hind wing light 
to dark smoky brown; the veins slightly darkened and 
a dark line along termen. Alar expanse, 16-17 mm. 

Male genitaUa with clasper short, blimt, curved. 
Cornuti moderately long, approximately the same size 
and lying side by side. A long slender arm projecting 
from each dorsolateral angle of vinculum. Female 
genitalia with two large, opposing, roundly oval, gran- 
ulate patches on bursa, and some rather strong, darkly 
pigmented spining on the lobe giving off the ductus 
seminalis. 

Type localities: California [?] (crassifasciella, lost?) ; 
unknown (decipientella, in USNM); Lakeland, Fla. 
(crataegella, in USNM) . 

Food plants: Vaccinium, Crataegus. 

Distribution: Maryland, Plummers Isl. (May); 
District of Columbia, Washington (May) ; Georgia, Sape- 
loe (Sept.); Florida, Lakeland (May). 

A male from the District of Columbia (reared from 
Vaccinium by Chambliss, May 10, 1895) and a similar 
female from Plummers Isl., Md., in the National Col- 
lection agree in every detail with Ragonot's figure and 
description of crassifasciella and were so identified by 
Dyar. His decipientella was described from a single 
specimen without locality label. It is merely a dark- 
suffused male with the pale coloration limited to a more 
restricted area about the discal spots and some light 
dusting between the outer veins. The Georgia example 
(reared from Vaccinium) is a female and matches the 
type of decipientella except for venation. It is another 
of the all too frequent phycitid freaks with vein 4 miss- 
ing from hind wing and 7-8 long stalked. The Barnes 
and McDunnough tj'pe of crataegella (also a male and 
reared from Crataegus) differs from typical crassifasciella 
only in a somewhat more extended and lighter colora- 
tion of the pale areas of forewing. All three males 
before me agree in genitalic structure. 

The species is easUy identified by the peculiar develop- 
ment of the vinculum, not found in any other American 
species in the genus. 

Dr. Bourgogne informs me that the type oiinquilinella 
could not be found at Paris. It is probably lost, and I 
suspect that the California citation in Ragonot's Mono- 
graph was probably a guess. I have seen nothing from 
the Pacific Coast States that could possibly be his 
species. 



271. Nephopteryx bisra Dyar 
Figure 833 
Nephopteryx bisra Dyar, Ins. Insc. Menstr., vol 7, p. 51, 1919. 

Fore\ving pale gray-brown; base shaded with black- 
ish; a straight, narrow, oblique whitish line along inner 
margin of the inner border of the antemedial line, the 
latter a thin, oblique, notched, whitish line, obscured 
towards costa; its inner border a wide blackish band 
reaching to costa; its outer border a narrow black line; 
a pale patch on inner margin following the antemedial 
line; subterminal line distinct, whitish gray, sinuate, 
margined by narrow blackish brown inner and outer 
lines; discal spots fused into a lunate mark with a pale 
surrounding shade; a blackish line along termen. Hind 
wing pale gray-brown, paler thajn ground color of fore- 
wing; veins not appreciably darkened; a narrow, dark 
(brownish) line along termen. Alar expanse, 21 mm. 

Female genitalia with a large granulate patch cover- 
ing most of dorsal surface of bursa, a smaller latero- 
ventral patch and some scattered granulations at left, 
posterior angle on ventral side; bursa otherwise rather 
densely and finely spinose. 

Type locality: Orizaba, Mexico (type in USNM). 

Food plant: Unknown. 

Known only from the female type. The placement 
in the second group of Nephopteryx is provisional, pend- 
ing discovery of a male. In the type, vein 10 of fore- 
wing is shortly stalked with the stem of 8-9. 

69. Genus Tlascala Hulst 

Tlascala Hulst, Phycitidae of N. Amer., p. 146, 1890.— Forbes, 
Cornell Mem. 68, p. 623, 1923. (Type of genus: Nepho- 
pteryx reductella Walker.) 

Tongue well developed. Antenna weakly pubescent; 
on male, with sinus and enlarged scale tuft in base of 
shaft. Labial palpus upturned, not appressed to face, 
cylindrical, smooth scaled, reaching above vertex; 
second segment of male not hoUowed; third segment 
moderately long, acuminate. Maxillary palpi of both 
sexes squamous, appressed to face. Forewing with 
most of inner border of antemedial line consisting of 
raised scales; 11 veins; vein 2 from near lower outer 
angle of cell; 3 from the angle, about equidistant from 
2 and 4 ; 4 and 5 very shortly stalked or connate, rarely 
(in individual specimens) closely approximate at base; 
6 from below upper angle of cell, straight; 8-9 long 
stalked (for three-fourths of their lengths); 10 from the 
cell approximate to or connate with the stalk of 8-9 at 
base, and approximate to it for some distance beyond 
base; male without costal fold. Hind wing venation 
similar to that of Meroptera and Nephopteryx. Eighth 
abdominal segment of male with compound scale tufts. 

Male genitalia with strongly sclerotized, broadly tri- 
angulate clasper with a row of irregular teeth along its 
outer margin. Penis armed with two, equally sized, 
stout, sharply curved, thornlike cornuti. Otherwise 
as in Nephopteryx. 



134 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



Female genitalia without signum; bursa finely and 
densely spined over most of inner surface and with a 
cluster of longer and stronger spines near middle of 
lateral margin; ductus bursae short (less than half the 
length of bursa) ; armed on ventral surface by an elon- 
gate pair of granulate plates; at genital opening a 
strongly sclerotized, centrally interrupted genital plate, 
attached to a narrow, sclerotized collar (incomplete 
dorsaUy and ventraUy) and supplemental to the regular 
eighth-segment collar; ductus seminalis from bursa near 
junction of bursa and ductus bursae. 

The genus is close to and has several characters in 
common with both Meroptera and Nephopteryx, resem- 
bling the former in the well developed genital plate of 
the female genitalia and the strong, serrate clasper of 
the male, differing from both genera in the rough scaling 
on forewing, the ungrooved labial palpi and much 
stouter antennal tuft of the male, the sharply curved 
cornuti on penis, and the supplemental collar attached 
to the female genital plate. As here defined it contains 
only its American type species. 

272. Tlascala reductella (Walker) 
Figures 28, 348, 834 

Nephopteryx reductella Walker, List, vol. 27, p. 63, 1863. — 

Eagonot, Ent. Amer., vol. 5, p. 115, 1889; Monograph, pt. 1, 

p. 283, 1893. 
Pempelia gleditschiella Fernald, in Comstock, in Rep. [U. S.] 

Comm. Dep. Agr. for 1880, p. 262, 1881.— Packard, U. S. 

Dep. Agr. Fifth Rep. Ent. Com., p. 652, 1890. 
T^lascala rediMtella (Walker) Hulst, Phycitidae of N. Amer., p. 

146, 1890.— Forbes, Cornell Mem. 68, p. 624, 1923.— 

McDunnough, Check list. No. 6180, 1939. 

In color and maculation similar to the European 
Nephopteryx rhenella except for the raised scaling of the 
subbasal black bar. 

Forewing duU ash gray with pm-phsh brown shading 
at extreme base and paler brownish shading on outer 
median and terminal areas; antemedial line complete 
but faint, narrow, oblique, slightly indented on lower 
haK, preceded by a broad black band of rough scales 
and followed by a narrow black band; subterminal line 
obscure, sinuate-dentate, faintly bordered by dark inner 
and outer lines; discal dots distinct, blackish, separated; 
a row of faint Islackish dots along termen. Hind wing 
pale smoky fuscous; the veins very slightly darkened 
and a narrow dark line along termen. Alar expanse, 
17-23 mm. 

Male genitalia with cornuti set side by side near outer 
end of penis, their apices turned away from each other. 
Female genitaha as given for the genus. 

Type localities: Honduras [sic] {reductella, in BM); 
District of Columbia (glediischiella, in USNM) . 

Food plant : Gleditsia. Larva a leaf-tier. 

Distribution: District oj Columbia (May, July); 
Maryland, Plummers Isl. (June); North Carolina, Hil- 
tonhead Isl. (Aug.); Pennsylvania; Illinois, Decatur 
(Mar., Apr., May, July), Quincy (May); Iowa, Ames; 
Kansas, Lawrence (May); Missouri, St. Louis (May); 
Texas, Kenedy (Apr.), Paris (May), Victoria (July), 



ZavaUa County (Apr.) ; Louisiaria, Crown Point (June, 
larva). New Orleans (larva, June). 

Walker gives Honduras as the type locality. I 
rather doubt the correctness of this citation, for I have 
never seen anything from Central America that even 
remotely resembled the species. I have not seen his 
types, but have no reason to question the correctness of 
Ragonot's reference of gleditschiella to synonymy. 

70. Tulsa, new genus 

Type of genus: Nephopteryx finiiella Walker. 

Characters of Tlascala except: 

Forewing with some rough scaling in the median 
area beyond the outer margin of the antemedian line, 
sometimes a small tuft on lower fold, always a few 
roughened scales in the discal spots; veins 4-5 approxi- 
mate at base and for a short distance beyond. Male 
genitalia with sacculus of harpe considerably enlarged, 
strongly sclerotized, densely and finely spined along 
entire lower margin, and produced at apex; clasper a 
thin, dished plate produced into an elongate, curved, 
sharply pointed claw at each lower angle. Uncus 
broadened at apex. TranstiUa complete, very weakly 
but evenly sclerotized throughout. Cornuti straight, 
set one before the other. Female genitalia with several 
lines of fine spines running from bursa into ductus 
bursae almost to genital opening; genital plate and its 
attached supplemental coUar strongly wrinkled (more 
so than shown in the figures) ; no granulations in ductus 
bursae. 

The genus is very close to Tlascala and I propose the 
new name with some misgiving; but the habitus of the 
moths and their genitalia, male and female, differ so 
much from those of the type of Tlascala that something 
more than a species-group difference is indicated. When 
larvae and host relations of the Tulsa species are known 
we shall probably find additional supporting characters 
for the genus. Specifically the genitalia are remarkably 
similar, offering little or nothing to distinguish the 
species. Four are here recognized. 

273. Tulsa fiaitella (Walker), new combination 

Figure 349 

Nephopteryx finitella Walker, List, vol. 27, p. 53, 1863. — Ragonot, 
Monograph, pt. 1, p. 282, 1893. 

Tlascala finitella (Walker) Hulst, Phycitidae of N. Amer., p. 147, 
1890.— Forbes, Cornell Mem. 68, p. 624, 1923.— McDun- 
nough, Check list. No. 6177, 1939. 

Elasmopalpus melanellus Hulst, Phycitidae of N. Amer., p. 157, 
1890. — Barnes and McDunnough, Contributions, vol. 3, 
p. 199, 1916 (makes synonym ot finitella). 

Forewing very dark gray with the blackish borders of 
the antemedial and postmedial lines but slightly con- 
trasted; some slight dusting of white on subbasal, me- 
dial, and terminal areas; antemedial line somewhat 
stronger, its inner blackish bordering line more or less 
interrupted on the veins, its outer border continuous 
but faint; discal dots tending to coalesce; an obscure 
row of blackish dots along termen; raised scales con- 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



135 



spicuous on lower half of inner border of antemedial 
line and as a patch on middle of lower fold; a raised 
scale or two in the discal spots. Hind wing pale smoky 
fuscous with a darker shading towards termen; the 
veins slightly darkened; the entire wing darker on 
northern examples. Alar expanse, 21-25 mm. 

Male genitaha with no distinguishing specific features. 
Female genitaUa similar to those of umbripennis but 
smaller (about the size of infinitella) and with more 
decided wrinkling of the genital plate and its supple- 
mental collar. 

Type localities: United States (finitella, in BM); 
Florida {melanellus, in AMNH, ex Kutgers). 

Food plant. — Blueberry. This record is from a reared 
female (Brunswick, Ga., Quaintance No. 31501) in the 
National Collection. The larva, however, may have 
been accidentally on that plant. 

Distribution: United States: Florida, Charlotte 
Harbor (Mar.), Fort Myers (Apr.), Miami (Mar.), St. 
Petersburg (Apr.), Tampa, also examples with only 
state locality ; Georgia, Brunswick (June) ; North Caro- 
lina, Ealeigh (June); Virginia, Kichmond (May); Dis- 
trict oj Columbia, Washington (May, June) ; New Jersey, 
Essex County (May), Newark (May); MassachxLsetts, 
Martha's Vineyard (Aug.); Indiana, HessviUe (June). 
Canada: Ontario, Trenton; Quebec, Kazubazua (June). 
Walker also reports the species from Nova Scotia. 

The type of melanellus in the Rutgers Collection is a 
female without abdomen. Matching cotypes ( cT and 9) 
from Mrs. Slosson's material are in the National Collec- 
tion. There can be no question of the synonymical 
reference by Barnes and McDunnough. 

274. Tulsa umbripennis (Hulst), new combination 
Figures 350, 842 

Pinipestis umbripennis Hulst, Canadian Ent., vol. 27, p. 57, 1895. 

Ortholepis gilletteUa Dvar, Proc. Ent. Soe. Washington, vol. 6, 
p. 107, 1904. 

Tlascala umbripennis (Hulst) Barnes and McDunnough, Contri- 
butions, vol. 3, p. 195, 1916. — McDunnough, Check list, 
No. 6178, 1939. 

Fore and hind wings dark brown with a somewhat 
glossy sheen not possessed by the other species of the 
genus ; discal dots on forewing confluent, forming a bar 
on discocellular vein ; raised scales and maculation as in 
finitella. Alar expanse, 25-26 mm. 

Male genitalia, figured from type of gilletteUa, agree 
in every detail with those of the male type of umbri- 
pennis. Female genitalia with bursa copulatrix 
slightly larger than that of any of the other Tulsa 
species except oregonella. 

Type localities: Colorado {umbripennis, in AMNH, 
ex Rutgers; gilletteUa, in USNM). 

Food plant: Unknown. 

Known only from the type locality. A series before 
me is from Chimney GuJch, Colo. (June and July). 
The types of umbripennis and gilletteUa have only the 
state locaUty, but the latter was probably from the 
neighborhood of Fort Collins. 



275. Tulsa oregonella (Bamea and McDunnough), new combination 

Figure 351 

Tlascala oregonella Barnes and McDunnough, Contributions, 
vol. 4, p. 175, 1918.— McDunnough, Check list. No. 6179, 
1939. 

Forewing a dull, powdery, slate gray; basal area a 
trifle paler; the transverse lines somewhat more distinct 
and better defined than on umbripennis; inner black 
border of antemedial line slightly narrower than in 
preceding species ; dark borders of subterminal line very 
faint; discal dots separated. Hind wing smoky gray- 
brown, paler than that of umbripennis and not glossy. 
Alar expanse, 26-28 mm. 

Female genitalia similar to those of umbripennis. 

Type locality: Crater Lake, Oreg. (July; type in 
USNM). 

Food plant: Unknown. 

Known only from the type series. 

276. Tulsa infinitella (Dyar), new combination 

Figure 841 
Tlascala infinitella Dyar, Ins. Insc. Menstr., vol. 7, p. 52, 1919. 

Similar to oregonella except: Forewing slightly darker; 
antemedial line obscure; the discal dots confluent, form- 
ing a blackish Une of sUghtly roughened scales. Hind 
wing as in southern specimens oi finitella; pale smoky 
fuscous at base with a darker shading towards termen. 
Alar expanse, 27 mm. 

Female genitalia like those oi finitella except lateral 
elements of genital plate smoother. 

Type locality: Orizaba, Mexico (type in USNM). 

Food plant: Unknown. 

Known only from the female type. 



Genus 71: Homoeographa 

[Venational division C. veins 4 and 5 shortly stalked or weakly 
anastomosed for a short distance beyond cell. Hind wing with 
discocellular vein of cell vertical, straight. Antenna of male 
with sinus and scale tuft in base of shaft. Labial palpus obliquely 
upturned, second segment laterally flattened and broadly scaled, 
on male grooved to hold maxillary palpus. MaxiUary palpus of 
male in the form of an aigrette. Eighth abdominal segment of 
male with compound scale tufts. Male genitalia without trans- 
tilla; clasper present, digitate; penis armed with two moderately 
stout cornuti. Female genitalia without signum; bursa with 
deep, convoluted, sclerotized folds; genital opening simple.] 



71. Genus Homoeographa Ragonot 

Homoeographa Ragonot, Nouv. Gen., p. 24, 1888; Monograph, 
pt. 1, pp. xlvi, 432, 1893. (Type of genus: Homoeographa 
lanceolella Ragonot.) 

Tongue well developed. Antenna pubescent; male 
with sinus and strong tuft in base of shaft. Labial 
palpus obliquely uptiu-ned, reaching well above vertex; 
second segment flattened and broadly scaled, on male 
grooved to hold the maxiUary palpus; third segment 
short, partially hidden in scaling of second. Maxillary 
palpus of male in the form of an aigrette; of female 



136 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



squamous. Forewing smooth; 11 veins; vein 2 from 
before the lower outer angle of cell; 3 from the angle, 
separated from 4-5; 4 and 5 shortly stalked or weakly 
anastomosed for less than half their lengths from cell; 
6 from below upper angle of cell, slightly bent at base; 
8 and 9 stalked for over half their lengths; 10 from the 
cell, connate or closely anastomosed at base with the 
stalk of 8-9. Hind wing with vein 2 from well before 
lower outer angle of cell; 3 from the angle, long; 4 and 5 
stalked for about four-fifths of their lengths; 7 and 8 
anastomosed for three-fourths of their lengths; cell 
short, one-third the length of wing; discoceUular vein 
vertical, straight. Eighth abdominal segment of male 
with compound ventral scale tufts. 

Male genitaUa with uncus subtriangulate and with 
blunt, moderately broad, notched apical margin. Apical 
process of gnathos a moderately stout hook with a 
slender, digitate basal projection. TranstUIa absent. 
Harpe with costa sclerotized throughout, but not 
produced at apex; cucullus simple, curved, apex 
bluntly pointed; clasper present, simple, erect, digitate. 
Penis armed with two moderately stout cornuti, about 
one-third as long as aedeagus. Vinculiun about twice 
as long as broad, evenly tapering to its truncate, 
strongly sclerotized, anterior margin. Vinculum U- 
shaped with somewhat enlarged base. 

Female genitalia with bursa elongate, narrow, signum 
absent, two or three deep, convolute sclerotized folds 
at posterior half, the sclerotization extending for a 
short distance into ductus bursae; genital opening 
simple. Ductus seminalis from bursa, near junction of 
bursa and ductus bursae. 

The genus is easily distinguished by its hind wing 
venation and genitalia. The male genitalia indicate 
a close relationship to the Nephopteryx group of genera. 
However the characteristic hair brush on the harpe of 
the latter are absent from Homoeographa. It contains 
only one known tropical American species, 

277. Homoeographa lanceoleUa Ragonot 
Figures 352, 839 

Homoeographa lanceoleUa Ragonot, Nouv. Gen., p. 25, 1888; 
Monograph, pt. 1, p. 433, 1893. 

Forewing gray heavUy dusted with white on costal 
half; antemedial line indicated by a narrow, white, in- 
wardly notched liae between cell and inner margin and 
above that by its incompleted outer border, a black 
line obhque from costa to cell thence inwardly angled to 
lower vein of cell; sub terminal line faint, sinuate, in- 
dicated chiefly by blackish gray bordering streaks from 
costa, the inner one the longer and continued as a weak 
blaclash shading to inner margin; discal dots small, 
separated, blackish gray; in outer area black streaklets 
bordering vein lb above and veins 3 and 6 below. 
Hind wing semi translucent smoky white; the veins 
sUghtly darkened and a narrow dark line along termen. 
Head ashy white. Alar expanse, 21 mm. 

Female genitalic characters as given for the genus. 

Type locality: CaUao, Perd (type in Paris Mus.). 

Food plant: Unknown. 



Known only from the type series in the Museum 
National d'Histoire NatureUe, Paris, and the British 
Museum. 



Genera 72-76: Telethusia to Pyla 

[Venational division B. Veins 4 and 5 of forewing usually 
separated at base (shortly stalked in Actrix); 10 from the cell, 
separated at base from stalk of 8-9. Hind wing with cell less 
than half the length of wing (about one-thid in Pyla). Antenna 
of male with sinus and scale tuft in base of shaft. Labial palpus 
oblique or upturned. Maxillary palpus various (minute, 
squamous or aigrettelike). Male genitalia with transtilla 
usually absent, if present (Phobus, Stylopalpia) incomplete or 
its median area very weakly sclerotized; harpe with sclerotized 
costa sometimes produced at base, never at apex; clasper absent 
or more or less developed (strongly so in many species of Pyla) ; 
aedeagus frequently divided (bifid) or spined; penis unarmed or 
finely scobinate or finely and weakly spined, rarely (Phobus) 
with a single cornutus. Female genitalia without signum; bursa 
frequently smooth or weakly spined, occasionally with some 
sclerotized folds continued from ductus bursae; the latter more 
or less sclerotized in part, in many Pyla species broadly expanded 
towards genital opening.] 

72. Telethusia, new genus 

Type of genus: Pempelia ovalis Packard. 

Tongue well developed. Antenna pubescent; on male 
with sinus and enlarged scale tuft in base of shaft. 
Labial palpus obliquely upturned, reaching to vertex 
on male, above vertex on female; laterally flattened and 
broadly scaled; second segment of male grooved on 
inner side to hold the tongue; third segmelit consider- 
ably shorter than second, bluntly pointed, more or less 
deflected forward and partially hidden in scaling of 
second segment. Maxillary palpus minute (a mere 
vestige). Forewing smooth; 11 veins; vein 2 from 
before lower outer angle of cell; 3 from the angle, 
nearer to 4 than to 2 ; 4 and 5 separated at base, parallel 
for a short distance beyond; 6 from below upper angle 
of cell, straight; 8 and 9 stalked for half or nearly half 
their lengths; 10 from the cell separated from the stalk 
of 8-9 at base and divergent from it shortly beyond; 
male without costal fold. Hind wing with vein 2 from 
before lower outer angle of cell; 3 from the angle, 
connate at base with the stalk of 4-5; 4 and 5 stalked 
for about half their lengths; 7 and 8 contiguous or 
weakly anastomosed for a short distance beyond cell; 
cell less than half the length of wing; discoceUular vein 
curved, outwardly produced at lower angle of cell. 
Eighth abdominal segment of male with compound 
scale tufts. 

Male genitalia with uncus subtriangulate; apex 
bluntly rounded. Apical process of gnathos a short, 
stout hook. TranstiUa absent. Harpe simple; clasper 
rudimentary. Aedeagus simple, straight, not taper- 
ing; penis unarmed except for a small comb of very 
weak, short, slender spines. Vinculum stout, longer 
than greatest width, tapering slightly to trimcated 
terminal margin. 

Female genitalia with ovipositor strongly sclerotized; 
apophyses (supporting rods) of ovipositor and eighth 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



137 



segment collar stout; bursa elongate, finely scobinate 
over two-thirds of its inner surface but without signum 
or other sclerotization; ductus bursae unsclerotized 
except for a narrow, weak band near genital opening; 
ductus seminalis from junction of bursa and ductus 
bursae. At least half of membrane between coUar and 
ovipositor finely and densely spinose. 

The genus is erected for a species hitherto referred to 
Nephopteryx. It differs from the latter in its vestigial 
maxillary palpi, the lack of cornutus or cornuti of penis 
or the hair brush on harpe of its male genitaUa, the 
hardened ovipositor of female, the shorter stalking of 
veins 8 and 9, and the somewhat more separated condi- 
tion at base of veins 4 and 5 of forewing. 

278. Telethusia ovalis (Packard), new combination 
Figures 353, 843 

Pempelia ovalis Packard, Ann. Lye. Nat. Hist. New York, vol. 10, 

p. 269, 1873. 
Nephopteryx lalifasciatella Packard, Ann. Lye. Nat. Hist., New 

York, vol. 10, p. 269, 1873. 
Nephopteryx ovalis (Packard) Grote, BuU., U. S. Geol. Geogr. 

Surv. Terr., vol. 4, p. 696, 1878; North Amer. Ent., vol. 1, 

p. 11, 1879.— Hulst, Phycitidae of N. Amer., p. 144, 1890.— 

Ragonot, Monograph, pt. 1, p. 269, 1893. — Forbes, Cornell 

Mem. 68, p. 623, 1923. — McDunnough, Check list, No. 

6163, 1939. 
Nephopteryx ovalis geminipunclella Ragonot, Monograph, pt. 1, 

p. 270, 1893. 
Nephopteryx modestella Hulst, Canadian Ent., vol. 34, p. 170 

1900. — Barnes and McDunnough, Contributions, vol. 3, 

p. 196, 1916. 

Forewing ashy gray-white with a distinct powdery 
appearance, the white dusting conspicuous on median 
and basal areas; antemedial line oblique, zigzag (twice 
notched), narrow, white, bordered inwardly by a broad 
dark fuscous (blacldsh gray) band which is interrupted 
by a streak of dull ocherous orange at lower fold (this 
ocherous shade also continued along fold in median 
area) ; antemedial line bordered outwardly by a blackish 
gray bar at costa and similarly colored dots on cell and 
just above inner margin; subterminal line sinuate 
bordered inwardly by a more or less broken blackish 
gray line and outwardly by a broader blackish gray, 
brownish, or brownish ocherous shade (the latter when 
present interrupted by blacldsh streaklets on the veins), 
both borders strongly accented at costa; on most speci- 
mens a narrow, oblique, dark shading across median 
area from costal inner margin of subterminal line; discal 
dots distinct, separated, blackish, the lower sometimes 
expanded into a black smudge; a row of black dots along 
termen. Hind wing pale smoky fuscous, more whitish 
in some specimens, darker and somewhat brownish gray 
in others. Alar expanse, 20-28 mm. 

Genitaha as given for the genus. 

Type localities: Maine (ovalis and latifasciateUa, 
in MCZ) ; Washington State (geminipunctella, in Paris 
Mus.) ; Massachusetts (modestella, in AMNH, ex Rut- 
gers). 

Food plants: Antennaria, Eriophyllum ignotum. 
These records from Washington specimens reared by 
J. F. G. Clarke. Presumably on other Compositae. 



Distribution: United States: Maine, Orono (July), 
Wales (July); New Hampshire, Durham, Hampton 
(Aug.), Fort Washington (July); Vermont, Clarenton; 
Massachusetts; Connecticut, East River (July); New 
York, CatskUl Mts., Ilion (June, July) ; Colorado, Gun- 
nison Coimty (near Altmont, July); Utah, Stockton 
(June, July); Montana, Missoula (Aug.); Washington, 
Bellingham (June), Chuckanut Bay (Whatcom County, 
June), Godman Springs (Blue Mts., July), Kamiack 
Butte (May) , Pullman (June, July) ; California, Placer 
County (June), San Jacinto Mts. (July), Tuolumne 
Meadows (July, Aug.). Canada: Ontario, Trenton 
(July); Alberta, Banff (June, July); British Columbia, 
Wellington. 

The species is variable in color, especially in the 
Western areas of the United States and Canada. Most 
of the specimens from Washington and British Colum- 
bia have the white dusting on forewing more conspicu- 
ous and the dark markings more strongly contrasted 
than on eastern examples. However, there are inter- 
grades, and no sharp line can be drawn on color between 
the two areas. In the Tuolumne Meadows of Cali- 
fornia there is a larger (26-28 mm.) and somewhat paler 
form. Specimens from Colorado, Alberta, and occa- 
sionally from Washington form another darker and 
duller variety, the forewing showing little or no brown 
shading, the pattern markings a dull black, and the 
pale areas more gray than whitish and less strongly 
contrasted against the dark markings than in other 
Washington or eastern specimens. Three specimens 
before me from California have the transverse dark 
lines much weaker and the over-all color an ashy gray 
with a slight bluish tint. The Utah examples are the 
most distinctive of all the forms, their hind wings de- 
cidedly paler, the forewing a very pale ashy gray and 
all the darker pattern markings more or less obscured 
or obliterated. I do not think that these varieties 
represent anything but color forms or that any one of 
them is entitled to a subspecific designation. More 
and wider collecting throughout the Middle and Far 
West will probably turn up still other color variants. 
The species itself, despite its variability, is easily identi- 
fied by its genitalia. 

279. Telethusia rhypodella (Hulst), new combination 

Glyptoteles rhypodella Hulst, Ent. Amer., vol. 3, p. 137, 1887. 

Nephopteryx rhypodella (Hulst), Phycitidae of N. Amer., p. 144, 
1890.— Ragonot, Monograph, p. 1, p. 270, 1893.— Barnes 
and McDunnough, Contributions, vol. 3, p. 196, 1916.— 
McDunnough, Check list. No. 6165, 1939. 

There are no specimens in the Rutgers Collection or 
elsewhere that I have seen matching Hulst's description, 
nor any available Oregon examples that could be re- 
ferred to rhypodella. The alleged type at Rutgers, a 
female without locality label and bearing only the num- 
ber 42, is a typical representative of Phobus curvatella 
(Ragonot). Unfortunately the Hulst "types" are fre- 
quently as unreliable as his type designations and this 
particular type is probably spurious. Hulst's descrip- 
tions, on the other hand, are usually more reliable and I 



138 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



suspect that when sufficient Oregon material is available 
we shall find that rhypodella is merely one of the nu- 
merous color forms of ovalis. 

Type locality: "Oregon" (type lost?). 

Food plant: Unknown. 

73. Phobus, new genus 

Type of genus: Dioryctria brucei Hulst. 

Characters of Telethusia except: Eighth abdominal 
segment of male with a pair of ventrolateral hair tufts. 
Penis of male genitalia armed with a single cornutus; 
usually also a cluster of very fine, minute, slender spines 
at apex of aedeagus (but not on membranous penis). 
TranstUla represented at least by its lateral elements, 
sometimes the median area is recognizable but is very 
weakly sclerotized and the completed band not a con- 
stant character. Female genitalia with ovipositor nor- 
mal (not strongly sclerotized) ; apophyses of ovipositor 
and eighth-segment collar slender; bursa copula trix 
simple (smooth); membrane between collar and ovi- 
positor smooth. 

The foregoing differences seem to justify separation 
from Telethusia, with which the genus is very closely 
related. 

280. Phobus brucei (Hulst), new combination 
FiGTJBBS 354, 844 

Dioryctria brucei Hulst, Canadian Ent., vol. 27, p. 55, 1895. 
Ambesa lallatalis Hulst (not Hulst), U. S. Nat. Mus. Bull. 52, p. 

422, 1903. 
Tacoma lallatalis Dyar (not Hulst), Proc. Ent. Soc. Washington, 

vol. 6, p. 227, 1904. 
Nephopteryx lallatalis brucei (Hulst) McDunnough, Check list, 

No. 6160, 1939. 

Forewing whitish more or less dusted and shaded with 
blackish or fuscous scales, making the general color 
cream white (with some pale brownish shading on paler, 
weakly marked examples) to ashy gray with a faint 
bluish gray tint (on well-marked specimens) ; the trans- 
verse lines irregular and more or less interrupted and 
not strongly contrasted; antemedial line oblique, ser- 
rate, interrupted at lower fold by a pale olivaceous- 
ocherous shade which extends rather broadly the length 
of the fold and also cuts the subterminal line; a similar 
but somewhat weaker shade fills the cell; outer margin 
of antemedial line consisting of a thin blackish line 
curving outwardly from costa to top of cell, a small 
blackish dot or dash on lower vein of cell and a similar 
blackish marking on vein lb; subterminal line markedly 
serrate, deeply indented (almost to cell) below costa, on 
weU-marked (darker) specimens bordered inwardly by a 
black line from costa at least to cell, this line frequently 
continued along top of cell to the black outer marking 
of antemedial line, forming a continuous, long, narrow 
hook along the median and postmedian subcostal area; 
below the lower fold the blackish outer border of the 
subterminal line is also continued back, as a black line 
under vein lb to and fusing with a narrow blackish line 
on the outer edge of the antemedial line forming a nar- 
row, oval marking on lower margin between the trans- 



verse lines; on dark examples a more or less conspicuous, 
blackish, quadrate patch inwardly bordering the ante- 
medial line at inner margin; on pale specimens this 
patch pale brown, more or less obscured; discal dots 
obscured, rarely distinguishable; a row of small narrow 
black or brownish dots along termen. Hind wings 
subpellucid with a faint ocherous tint; the veins not 
appreciably darkened; a faint narrow dark shade along 
termen. Alar expanse, 26-29 mm. 

Male genitalia with cornutus small, slender. Female 
genitalia with bursa very short, not much longer than 
ductus bursae. 

Type locality: Colorado (type in AMNH, ex Rut- 
gers). 

Food plant: Unknown. 

Distribution: Colorado; Utah, Eureka (June, July), 
Stockton (June, July, Sept.) ; Nevada, Ormsby, and one 
female with only the state locality (a pseudotype of 
lallatalis Hulst); California, San Luis Obispo; Washing- 
ton, Pulbnan (July). 

All specimens of this species in the National Collec- 
tion had been identified as lallatalis Hulst on Dyar's 
misidentification of the latter species. As a result 
Dyar referred hrucei as a synonym of lallatalis. Hulst, 
however, was primarily to blame for the confusion; for 
he had identified and sent out as "types" of lallatalis 
specimens of both brucei and Interjectio denticulella. 
The true brucei resembles superficially both lallatalis 
and denticulella in some of its more strildng details of 
maculation, but is easily distinguished from both by 
its male and female genitalia. 

281. Phobus funerellus (Dyar), new combination 

Salebria funerella Dyar, Ins. Insc. Menstr., vol. 13, p. 12, 1925. — 
McDunnough, Check list, No. 6193, 1939. 

Forewing blackish to dark brownish gray more or less 
dusted with white on basal, median, and terminal areas ; 
the white dusting very faint and scattered on the type 
series from Southern California, which have a uniform 
blackish gray ground color, much stronger on specimens 
from Washington, British Columbia, and New Mexico ; 
transverse lines and thin blackish borders complete (not 
interrupted as in brucei), sinuate; the antemedial line 
oblique, sharply serrate, narrow, whitish gray, bordered 
outwardly by a narrow black line and inwardly by a 
broad, unbroken blackish or (on paler examples) dark 
gray-brown band, this band distinguishable and con- 
trasted even on the darkest, most suffused examples; 
subterminal line seri'ate but not deeply indented below 
costa, bordered inwardly by a narrow, continuous black 
line, the latter not continued inwardly below costa or 
on vein lb as it is on brucei; subterminal line bordered 
outwardly by a rather broad dark band; discal dots 
usually distinct, black, more or less confluent; a row of 
blackish dots along termen. Hind wing brown; the 
veins darkened and a narrow blackish line along termen. 
Alar expanse, 24.5-30 mm. 

Genitalia similar to those of brucei. 

Type locality: Southern California (type in USNM). 

Food plant: Unknown. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



139 



Distribution: United States; California (Southern 
California without more definite locaUty, the tjrpe 
series), Clarksville (El Dorado County, June); Washing- 
ton, Pullman; New Mexico, Fort Wingate (July). 
Canada: British Columbia, Departure Bay (Aug.), 
Duncans (Vancouver Isl., July), Nicola (July), Welling- 
ton (June). Also one male without state locality 
labeled "Larima Co. [sic], Aug., 1901, Schaus collector." 

The species is distinct and easily distinguished from 
brv^ei on the color and pattern of forewing, especially 
by the broad black band extending from inner margin 
to costa before the antemedial line and by the shallow 
indentation of the subterminal line below costa. 

282. Phobus curvalellns (Ragonot), new combination 
Figures 355, 845 

Nephopteryx curvatella Ragonot, N. Amer. Phycitidae, p. 7, 
1887. — Barnes and McDunnough, Contributions, vol. 3, 
p. 196, 1916.— McDunnough, Check list. No. 6166, 1939. 

Nephopteryx rhypodella Ragonot (not Hulst), Monograph, pt. 1, 
p. 270, 1893. 

Forewing ashy bluish gray; the transverse lines com- 
plete, narrow, white, obscure except on the well-marked 
darker examples, indicated chiefly by the fine, black 
outer border of the antemedial line and similar black 
inner border of the subterminal line ; preceding the ante- 
medial line a quadrate blackish spot on inner margin; 
limited above by a weak, smaller, pale, somewhat oliva- 
ceous shade in the lower fold, this pale shade not 
interrupting the antemedial line itself; indentations of 
subterminal line as in junerellus; discal dots obscure, 
the lower one sometimes distinct (under magnification) 
and frequently with a dark shade below it which forms 
a round dark spot, to the naked eye one of the more 
conspicuous markings on the wing; a row of more or 
less confluent black dots along termen. Hind wing 
translucent, whitish with a smoky shade towards apex; 
the veins darkened; a fine brown line along termen. 
Alar expanse, 26-30 mm. 

Male genitalia with cornutus of the same length as 
that of brucei and funerellus but somewhat stouter. 
GenitaUa otherwise like those of the species following 
(incertus). Female genitalia with bursa very long, and 
narrow throughout its length, but little wider than the 
ductus bursae. 

Type locality: America Septentrionalis (type in 
Paris Mus.). 

Food plant: Unknown. 

Distribution: California, Loma Linda (June, July), 
Los Angeles County (1,060 ft., Jime), Monachee 
Meadows (Tulare County, July), Mount Lowe (July), 
San Gabriel Mts. (1,700 ft., July); Arizona, Nogales 
(May), Santa Rita Mts. (May); Utah, Bellevue 
(Washington County, May, June); Colorado, Silverton 
(July). 

The spurious type of Telethusia rhypodella (Hulst) 
in the Rutgers Collection belongs here. It is discussed 
under the treatment of rhypodella (p. 137). 



283. Phobu9 incertus, new species 
Figures 356, 846 

Color and markings of forewing as in curvatellus 
except duller, lacking the bluish tint of the latter 
species; the quadrate dark spot preceding the ante- 
medial line also continued as a broad band to costa, 
though frequently interrupted by a pale shading at 
lower fold. 

Male genitalia with cornutus appreciably stouter and 
longer than that of any of the other species of the genus. 
Bursa copulatrix of the female genitalia less than half 
the length of that of curvatellus but twice the length of 
that of brucei or funerellus. 

Type locality: Strawberry Valley (6,000 ft.), San 
Jacinto Mts., Calif, (type in USNM, 61348). 

Described from male type and five male and five 
female paratypes from the type locality, collected by F. 
Grinnell, Jr., June 16, 17, and 18, 1908. 

Except for the genitalic differences this might easily 
be a higher altitude race of curvatellus, but the differ- 
ences in size of cornutus and length of bursa seem to 
be constant characters and greater than to be expected 
in variants of one species. 

74. Actrix, new genus 

Type of genus: Tacoma nyssaecolella Dyar. 

Tongue well developed. Antenna weakly pubescent; 
on male with sinus and scale tuft in base of shaft. 
Labial palpus upcurved, slender, reaching above vertex; 
second segment somewhat flattened and very slightly 
rough scaled, not grooved on male ; third segment about 
two-thirds the length of second, acuminate. Maxillary 
palpus squamous. Forewing smooth; 11 veins; vein 2 
from near lower outer angle of cell; 3 from the angle, 
approximately equidistant from 2 and 4 at base; 4 and 
5 shortly stalked; 6 from below upper angle of cell, 
straight; 8 and 9 very long stalked; 10 from the cell, 
approximate to the stalk of 8-9 for some distance; male 
without costal fold. Hind wing with vein 2 from before 
but near lower outer angle of cell; 3 from the angle, 
connate or (on occasional specimens) shortly fused 
with the stalk of 4-5 ; 4 and 5 stalked for two-thirds of 
their lengths; 7 and 8 strongly anastomosed beyond 
cell for half or a trifle more than half their lengths; 
cell slightly less than half the length of wing; dis- 
cocellular vein curved, extended outwardly at lower 
angle of cell. Eighth abdominal segment of male with 
pair of ventrolateral hair tufts. 

Male genitalia with uncus broader than long, its 
outer lateral angles slightly lobed. Apical process of 
gnathos broad, shieldhke, its lateral arms greatly 
reduced. Transtilla absent. Harpe without clasper. 
Aedeagus straight, divided towards apex, the projecting 
divided elements strongly sclerotized; penis with some 
very weak scobinations, but otherwise imarmed. 
Vinculum short, stout, about as long as greatest width, 
evenly rounded to blunt terminal margin. 



140 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



Female genitalia with bursa small, covered with 
dense slender spines on posterior half, otherwise very 
finely spinose, without signum; ductus bursae granu- 
late, short, expanded into a broad, strongly sclerotized, 
contorted plate towards genital opening; ductus semi- 
nalis from lobe of bursa near junction of bursa and 
ductus bursae. 

The genus is a further restriction from Nephopteryx 
of authors. Its nearest relationship seems to be Pyla, 
which it resembles in the peculiar modification of the 
ductus bursae of the female and the aedeagus of the 
male. Its venation, however, is quite distinct and, 
except for the strong anastomosis of veins 7 and 8 of 
hind :wing, closer to that of Tacoma. 

284. Actrix nyssaecolella (Dyar), new combination 
FiGUEES 357, 838 
Tacoma nyssaecolella Dyar, Proc. Ent. Soc. Washington vol. 6, 
p. 112, 1904.— Forbes, Cornell Mem. 68, p. 621, 1923.— 
Craighead, U. S. Dep. Agr. Misc. Publ. 657, p. 453, 1950. 
Nephopteryx nyssaecolella (Dyar) Barnes and McDunnough, 
Contributions, vol. 3, p. 196, 1916. — McDunnough, Check 
list, No. 6174, 1939. 

Forewing brownish gray ("lUaceous gray") paler in 
basal area beyond extreme base, in the half of median 
area just beyond antemedial line, and to a lesser extent 
in outer area (beyond the subterminal line), these pale 
areas of an ashy hue, due to a faint peppering of white 
scales; antemedial line narrow, obKque and very slightly 
angled at middle, dull white, preceded on inner margin 
by a subquadrate blackish brown patch and bordered 
outwardly at costa by a triangulate black patch which 
(on most specimens) continues as a more or less inter- 
rupted black line to inner margin; subterminal line 
sinuate (bulged at middle) and weakly serrate, dull 
white, bordered inwardly by a dark shade and outwardly 
by a narrower dark line, these dark borders especially 
marked and blackish at costa; discal dots confluent, 
blackish; a row of more or less confluent black dots 
along termen. Hind wing pale smoky brown, with a 
glossy sheen; veins very faintly darkened; a narrow 
brown line along termen. Alar expanse, 15-18 mm. 

Male genitalia with terminal margin of uncus con- 
cave, its lateral lobes tiu-ned laterally outward. Apical 
process of gnathos a convex shield, slightly longer than 
broad and with apical, lateral angles produced backward 
into bluntly pointed spines. Divided elements of aedea- 
gus produced as short, sharply and oppositely curved 
hooks. CucuUus of harpe sharply curved towards its 
apex. Female genitalia distinguished at once by the 
shape of the ventral sclerotized plate of ductus bursae 
and the eighth-segment collar, which is complete and 
strongly sclerotized ventrally. 

Type locality: Washington, D. C. (type in USNM). 

Food plant: Nyssa sylvatica (larva a leaf-folder). 

Distkibution: Massachusetts, Nantucket (July); 
Connecticut, East River (July, Aug.); Rhode Island, 
Weekapaugh (July) ; New Jersey, Anglesea (May, June) ; 
Pennsylvania, New Brighton (July, Aug.), Oak Station 
(Aug.) ; District of Columbia, Washington (Aug.) ; North 
Carolina, Southern Pines (Aug.), Tryon (Aug.). 



285. Actrix diBsimuIatrix, new species 
Figures 358, 837 

Superficially not distinguishable from nyssaecolella 
except (on the specimens before me) a slightly stronger 
white dusting on median area of forewing especially 
over iimer margin immediately following the antemedial 
line. This difference can hardly be expected to hold 
for the species. 

The genitalia of both sexes are very different from 
those of nyssaecolella. Male genitalia with terminal 
margin of uncus evenly roimded, its lateral lobes turned 
inward and downward, partially encircling the anal 
tube. Apical process of gnathos heart shaped, without 
produced angles. Divided elements of aedeagus pro- 
duced as long, straight, stout thornlike projections, one 
of which is coarsely scobinate. Female genitalia with 
a stout pair of strongly sclerotized, curved, lateral arms 
projecting forward from the ventral sclerotized plate 
of the ductus bursae. 

Type locality: Cape Henry, Va. (type in USNM, 
61349). 

Food plant: Nyssa sylvatica. 

Described from male type and two male and one 
female parat3T)es from the type locality, reared by A. 
Busck, Aug. 10, 16 and 18, 1927, from larvae feeding on 
the leaves of Nyssa sylvatica. Fom- larvae were pre- 
served from the collection. Three of these are blackish 
brown in color and are undoubtedly nyssaecolella. One 
larva is yellow with a pale yellowish head and thoracic 
shield. It is probably dissimvlatrix. No difference was 
noted in larval habits between the two forms. 

75. Genus Stylopalpia Hampson 

Stylopalpia Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7, 
p. 257, 1901. (Type of genus: Stylopalpia lunigerella 
Hampson.) 

Tongue well developed. Antenna very shortly pubes- 
cent, shaft of male somewhat flattened and with a 
shallow sinus at base containing a row of short spines 
more or less concealed by a small, weak scale tuft. 
Labial palpus obliquely upturned; third segment very 
long (nearly twice the length of second), slender and 
porrect in lunigerella, much shorter (about half the 
length of second) and oblique in the other two species, 
in these reaching a little above vertex, acuminate in all 
species. Maxillary palpus minute, its scaling slightly 
expanded. Forewing smooth; 11 veins; vein 2 from 
before the lower outer angle of cell; 3 from the angle; 
4 and 5 separated at base and divergent beyond, 4 but 
slightly nearer to 5 at base than to 3 ; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for slightly more 
than half their lengths; 10 from the cell, well separated 
from the stalk of 8-9 at base but just beyond approach- 
ing it for a very short distance; male without costal 
fold. Hind wing with vein 2 from well before lower 
outer angle of cell; 3 from the angle; 4 and 5 stalked 
for about half their lengths; 7 and 8 approximate or 
contiguous for a very short distance beyond cell; cell 
less than half the length of wing; discocellular vein 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



141 



curved, produced at lower angle, a short spur coimecting 
it and vein 3. Eighth abdominal segment with weak 
compound tufts in lunigerella, paired ventral tufts in 
the other two species. 

Male genitalia with uncus subtriangulate, its terminal 
margin more or less broadly rounded. Apical process 
of gnathos a simple, rather short, stout hook. Trans- 
tilla incomplete, sclerotized only in its reduced lateral 
elements (the central portion, as shown in the figures, 
distinguishable but not sclerotized). Harpe with large, 
strongly sclerotized, erect, scoop-shaped clasper, situ- 
ated towards base, below costa; cucullus narrow, 
elongate, very slightly tapering to rounded apex. 
Anellus shield-shaped, with small lateral lobes. Aedea- 
gus expanding to lateral, flanged projections before 
apex, the latter flanges, each bearing a cluster of strong 
spines; penis unarmed. Vinculum stout; slightly longer 
than broad; tapering slightly to broadly rounded ter- 
minal margin. 

Female genitalia without signum. Bursa mem- 
branous with only a little fine spining at junction of 
bursa and ductus biu^ae; ductus bursae weakly scle- 
rotized at, and just before, genital opening, the latter 
broad; ductus seminaUs from middle of ductus bursae. 

The genus was originally erected on the peculiar 
palps of its type species. This character, however, 
proves to be specific rather than generic. The two 
species here included do not have it; but agree with 
the type species on every other detail of venation, 
antennal structure, and genitalia. The genus can 
easily be maintained on its combination of male and 
female genitalic characters. 

286. Stylopalpia lunigerella Hampson 
Figures 30, 359, 848 

Stylopalpia lunigerella Hampson, Ann. Mag. Nat. Hist., ser. 7, 
vol. 7, p. 258, 1901. 

Third segment of labial palpus very long, slender 
and porrect. Eighth abdominal segment of male with 
compound tufts. 

Forewing ocherous (clay color) dusted with blackish, 
making the general shade dark gray, the ocherous color 
forming a contrasting band along costa and more or 
less lightening the lower median area and the base of 
inner margin; antemedial line indicated by a pale 
lunulate line between cell and inner margin and, on 
its upper half, by faint traces of its narrow, blackish, 
outer border; subterminal line very close to outer 
margin, slightly bulged at middle, not serrate, pre- 
ceded by some black streaklets on the veins and whitish 
or pale ocherous streaklets between them; discal dots 
separated, blackish; a few of black dots along termen. 
On female a somewhat broader brownish shade at apex 
and along termen and some darkening of the outer 
parts of the veins. Alar expanse, 18-24 mm. 

Male genitalia with clasper of harpe considerably 
longer than deep, serrate along lower and inner margins. 
Female genitalia with genital opening very broad. 

Type locality: Jamaica (type in BM). 



Food plant: Unknown. 

Distribution: Bahamas, Nassau. Puerto Rico: 
Aguirre Central (Apr., June, Aug.), Camuy (Apr.), 
Coamo Springs (Apr.), Ponce (Sept.), San German 
(June). Cuba: Santiago Province (June, July, Oct.). 
Jamaica. Mexico: Colima (Jan., June, July). 

The species is easily identified by its peculiar palpi, 
alike in both sexes. 

287. Stylopalpia scobiella (Grote), new combination 
Figure 360, 847 

Nephopteryx scobiella Grote, N. Amer. Ent., vol. 1, p. 5, 1890. — 
Hulst, Phycitidae of N. Amer., p. 143, 1890.— Ragonot, Ent. 
Amer., vol. 5, p. 115, 1889; Monograph, pt. 1, p. 266, 1893.— 
McDunnough, Check list. No. 6161, 1939. 

Lipographis decimerella Hulst, Ent. Amer., vol. 4, p. 117, 1888. 

Third segment of labial palpus short, oblique. Except 
on a few of the grayer specimens, head and thorax 
distinctly ocherous. 

Forewing pale gray to grayish ocherous, extreme base 
of wing ocherous and a similar pale ocherous shade 
rather broadly bordering the costa ; transverse markings 
nearly obliterated; antemedial line indicated only by a 
small lunate white spot on vein lb, preceded and fol- 
lowed by black dots, a similar blackish dot or streaklet 
at lower margin of cell (representing a median fragment 
of the usual black outer border of the antemedial line) ; 
subterminal line obscure, a very faint pale line weakly 
bordered inwardly by a darker shade ; lower discal spot 
a blackish streaklet, upper discal dot usually absent, if 
present very faint; a row of fine blackish dots along 
termen. Hind wings whitish with a faint ocherous or 
smoky tint; the veins little if any darkened; a thin 
brownish line along termen. Alar expanse, 24-26 mm. 

Male genitalia with clasper of harpe broadly oval, 
not serrate. Terminal margins of uncus and vinculum 
very broadly roimded, the vinculum not appreciably 
tapering. Eighth abdominal segment of male with 
paired ventral tufts. Female with genital opening more 
constricted than that of lunigerella. 

Type localities: Bosque County, Tex. (scobiella, in 
BM); Blanco County, Tex. (decimerella, in AMNH, 
ex Rutgers). 

Food plant: Unknown. 

Distribution: Texas, Barber (Apr.), BeeviUe (May), 
Blanco County (May), Bosque Coimty, Burnet County 
(Apr., Sept., Oct.), KerrvUle, Sabinal (Mar., Apr.), 
Sapulpa (May), San Diego (Apr.), Victoria (Apr., Sept.), 
Zavalla County (Apr.); Colorado, Glenwood Springs. 
Also four specimens from Texas with only the state 
locality. The species probably also occurs in northern 
Mexico. 

288. Stylopalpia argentinensis, new species 
Figure 361 

Labial palpus and eighth abdominal tufts as in scobi- 
ella. The head and thorax brown. 

Forewing pale brown with a strong dusting of white 
scales faintly peppered with black in median area; basal 



142 



XnsriTED STATES NATIONAL MUSEUM BULLETIN 207 



area brownish ocherous, shading to brown at extreme 
base and with a clouding of white in midbasal area ; ante- 
medial line complete, weU out on wing, oblique, inwardly 
notched at vein lb, white, bordered outwardly by a 
narrow blackish line; subterminal line also complete, 
further back from termen than in the two preceding 
species, oblique (parallel to termen), whitish ocherous 
with a narrow blackish inner border; discal dots distinct, 
separated, black. Hind wing dark brown; the fringe 
whitish ocherous with a dark median band. Alar ex- 
panse, 24 mm. 

Male genitalia similar to those of scobiella except: 
Uncus narrower and tapering to more narrowly rounded 
terminal margin; vinculum longer in proportion to its 
width; spines on aedeagus fewer and coarser. 

Type locality: "VUla Anna, F. S. C. Fe., Argentina" 
(type in BM). 

Food plant: Unknown. 

Described from imique male, collected by K. J. 
Hayward, Dec. 1925. 

76. Genus Pyla Grote 

Pyla Grote, New check list of North American moths, p. 55, 
1882. — Ragonot, N. Amer. Phycitidae, p. 9, 1887; Mono- 
graph, pt. 1, p. 481, 1893. — Hulst, Phycitidae of N. Amer., 
p. 161,1890. (Type of genua: Nephopieryx scintillans Giote.) 

Tongue well developed. Antenna finely pubescent; 
on male with sinus and scale tuft in base of shaft. 
Labial palpus oblique; second segment broadly scaled, 
somewhat flattened laterally, reaching above vertex; on 
male with a slight groove to hold the maxillary palpus; 
third segment short, less than one-third the length of 
second, porrect. MaxiUary palpus of male in the form 
of an aigrette, semiaigrette (i. e., the scales hairlike 
but short), or more or less squamous. Forewing smooth; 
11 veins; vein 2 from near lower outer angle of ceU; 3 
from the angle, slightly nearer to 4 at base than to 2 ; 
4 and 5 slightly separated at base, parallel for a short 
distance beyond ceU; 6 from below upper angle of cell 
straight; 8 and 9 stalked from one-half to a third of 
their lengths; 10 from the cell, slightly separated at 
base from the stalk of 8-9; male without costal fold. 
Hind wing with vein 2 from before but rather near 
lower outer angle of cell; 3 from the angle, connate with 
4; 4 and 5 stalked or anastomosed for half or slightly 
less than half of their lengths; 7 and 8 contiguous or 
very weakly anastomosed for a short distance beyond 
cell; cell less than half the length of wing; discocellular 
vein curved, outwardly produced at lower angle and 
connected with vein 3 by a short spur. Eighth abdom- 
inal segment of male with a pair of ventrolateral hair 
tufts (fig. 372b), or two or three pairs containing some 
modified scales (fig. 367c). 

Male genitalia with uncus broad, more or less trian- 
gulate. Apical process of gnathos a short, stout hook. 
Transtilla absent. Harpe usually with base of costa 
produced into a strong projecting hook, or spine, or 
spined lobe; frequently a strong hooked or spined clasper 
from median basal area; sacculus simple; costa strongly 
sclerotized but sclerotization rather abruptly terminated 



before apex of harpe. Aedeagus usually partially divided 
(bifurcate) or armed with projecting spine or spines, 
rarely simple; penis unarmed (except for a very weak 
cornutus in/ifscci). Vinculum stout, slightly tapered to 
truncate or more or less broadly roimded terminal 
margin. 

Female genitalia without signum; bursa copulatrix 
usually simple, sometimes with strongly sclerotized, 
convolute, longitudinal bands near junction of bursa 
and ductus bursae and extending a short distance into 
the ductus; ductus bursae short, widening to broad gen- 
ital opening, usually strongly and elaborately sclero- 
tized towards genital opening; genital opening rarely 
simple (fasciolalis, mridisujffusella)] ductus seminalia 
from biu-sa near junction of bursa and ductus bursae. 

The genus as here defined includes what superficially 
appears to be two distinct entities, one group of species 
with gray forewings and another with shiny brown 
wings, the latter the typical Pyla of authors. For con- 
venience of identification I am designating them as 
species groups. The division is not supported by any 
consistent structural character or combination of char- 
acters. The differences exhibited by the several species 
in male and female genitalia, male maxUlary palpi, and 
male abdominal tufts are striking, but apparently only 
of specific significance. 

Nothing is known of the food habits or early stages of 
any of the species except fusca, which is recorded from 
Erica in the Old World. I suspect that the Ericaceae 
will prove to be the chief hosts of the genus. 

Genus Pyla, Species 289-297: P. fasciolalis to 
P. hanhamella 

[Ground color of forewing gray.] 

289. Pyla fasciolalis (Hulst), new combination 
FiGTTEBs 362, 363, 849 

Pinipesiis fasciolalis Hulst, Trans. Amer. Ent. Soc, vol. 13, p. 

162, 1886. 
Nephopteryx fasciolalis (Hulst) Ragonot, Ent. Amer., vol. 5, p. 

115, 1889; Monograph, pt. 1, p. 271, 1893 (?).— Hulst, 

Phycitidae of N. Amer., p. 144, 1890. — McDunnough, Check 

list. No. 6167, 1890. 

Maxillary palpus of male squamous. 

Forewing gray finely powdered with white, giving the 
wing an ashy gray appearance; antemedial line distinct 
throughout, whitish, oblique, notched at top of ceU and 
on lower fold, bordered outwardly by a black line begin- 
ning as a black smudge on costa, inwardly by a mod- 
erately broad black line extending from inner margin to 
ceU ; subterminal line well marked, bulged at middle and 
more or less dentate, bordered inwardly by a blackish 
band and outwardly by a somewhat broader, fainter 
dark band, these dark borders strongest near costa ; 
discal dots distinct, small, normally separated, occa- 
sionally partially coalesced; a row of small black dots 
along termen. Hind wing smoky white with a pale 
brownish tint ; veins very faintly darkened ; a narrow dark 
shade along termen. Alar expanse, 27-30 mm. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



143 



Male genitalia with harpe simple; clasper vestigial. 
Aedeagus deeply divided; one of the divided elements 
slightly forked at apex. Female genitalia with strongly 
sclerotized, convulute bands in bursa; genital opening 
simple. 

Type locality: British Columbia (type in AMNH, 
ex Rutgers). 

Food plant: Unknown. 

Distribution: British Columbia, Goldstream (Aug.), 
Necola (July), Saunichton (July). 

The only specimen in the Rutgers Collection is a large 
male (30 mm.) from British Columbia labeled "Dioryc- 
tria jasciolalis Hulst, Type." I think we may safely 
assume this to be the actual type although in his original 
description Hulst gives "Nevada" as the type locality, 
probably one of his characteristic lapses. A perfect 
match for the typ^ in color, markings, and genitalia, is 
foimd in a specimen from Necola, in the Canadian Na- 
tional Collection. The other records cited above (Gold- 
stream and Saanichton) are from specimens in the U. S. 
National Collection. 

Also before me are four examples of what I take to be 
a variety of Jasciolalis — two males from Gunnison 
County, Colo. (July), and 2 females from Wallace, 
Idaho (Aug.). Their blackish markings on forewing 
are a trifle stronger, the vinculum of male genitalia (fig. 
363) is somewhat shorter than in iypxc&l Jasciolalis. The 
convolute bands in the bursa of the female are also a 
trifle longer. I doubt very much if these differences 
ndicate anything more than a possible local race. 

290. Pyla impostor, new species 
FiQUHES 364, 850 

Maxillary palpus of male in the form of an aigrette. 

Forewing color and markings similar to those of 
Jasciolalis except: Somewhat duller and darker; white 
dusting sparser; the transverse pale lines fainter, in 
some specimens much obscured; their blackish borders 
less strongly contrasted against the ground color of the 
wing. Hind wing pale smoky fuscous, the brownish 
tint oi Jasciolalis very faint or altogether lacking. Alar 
expanse, 23-30 mm. 

Male genitalia with a long, slender, somewhat flat- 
tened, outwardly curved clasper on harpe; base of costa 
not modified. Aedeagus divided for about half its 
length, moderately slender, the divided elements rigid, 
pointed and unforked at their apices. Female with 
convolute, sclerotized folds extending from bursa shortly 
into ductus bursae; ductus bursae itself partially sclero- 
tized, the sclerotization forming broad ventrolateral 
bands extending from just beyond the convolute folds 
of the bursa to genital opening. 

Type locality: Slate Peak, Whatcom County, Wash. 
(6,000-7,000 ft.; type in USNM, 61350; paratypes in 
USNM and Canadian Nat. CoU.). 

Food plant: Unknown. 

Described from male type and one female paratype 
from the type locality, collected by J. F. G. Clarke, Aug. 



2, 1940; and paratypes from the following localities: 
Bogachiel Peak, Olympic Mts., Wash., Aug. 9, 1936, 
Dr. A. F. Braun (9); Chimney Gulch, Golden, Colo., 
Oslar (cf); Colorado with only the state locality (a 
pseudotype, cf , of Jascialis Hulst, from the Fernald 
Collection); Big Belt Mts., Mont., July 18, 1928, J. 
McDunnough (d'); upper GaUatin Canyon, Mont., 
7,000 ft., July 4, 1928, J. McDunnough (c?); Banff, 
Alberta, July 20, 1925, Owen Bryant, (cf ); Lethbridge, 
Alberta, July 3, 1922, H. L. Seamans (cT); Moraine 
Lake, Alberta, July 3, 4, 7, 1923, J. McDunnough (4 cf 
and 2 9); Waterton Lakes, Alberta, July 23, 28, 1923, 
J. McDunnough (cf and 9); Hope Mts., British Co- 
lumbia, July 22, 1932, A. N. Gartrell (cf ); Mount Rev- 
elstoke, British Columbia, 6,000 ft., July 12, 1923, E. 
E. BuckeU (cf ). 

Most of the foregoing were in our collections as 
Jasciolalis on the basis of the false Hulst type in the 
National Museum. It is superficially like Jasciolalis, 
but a distinct mountain-top species easily identified by 
its genitalia. 

291. Pyla aequivoca, new species 
Figures 366, 855 

Maxillary palpus of male in the form of an aigrette. 

Superficially like impostor; the ground color of the 
male forewing a trifle more brownish gray, and the dark 
outer border of antemedial line somewhat broader and 
diffused into the ground color at costa, differences that 
could not be expected to hold in any extended series. 
Distinguished by its genitalia. Alar expanse, 26-29 mm. 

Male genitaha with clasper arising from midbasal 
area of harpe as in impostor, but much shorter and angu- 
late. Anellus U-shaped, its lateral arms long and 
slender. Aedeagus much shorter and somewhat stouter; 
its divided elements broader, decidedly flattened, termi- 
nating in sharp spines and more or less laterally spined 
towards apices. Eighth abdominal segment of male 
with two pairs of hair tufts. 

Type locality: Banff, Alberta, Canada (type in 
Canadian Nat. Coll., paratype in USNM, 61351). 

Food plant: Unknown. 

Described from male type and three male paratypes 
from the type locality, collected by C. B. D. Garrett, 
June 21 and 30 and Jidy 6, 1922; and one male parat3rpe 
from Hymers, Ontario June 8, 1915. In addition to the 
type series I have before me a female from the Canadian 
National CoUection collected at Aweme, Manitoba, 
Aug. 26, 1921, by Norman Criddle. Its genitalia differ 
markedly from those of impostor. The membrane of 
bursa is thickened near jimction with ductus bursae but 
lacks the sclerotizations of impostor; the ventrolateral 
bands of the ductus bursae are differently shaped and 
fuse into the ventrolateral sclerotizations of the inter- 
segmental area before the eighth-segment collar. While 
I have little doubt that this female is conspecific with 
the males, I am not designating it as a paratype. 



144 



tnsriTED STATES NATIONAL MUSEUM BULLETIN 207 



292. Pyla insinuatrix, new species 
Figures 365, 856 

Maxillary palpus of male in the form of an aigrette. 

Forewing paler than in the preceding species, due to 
stronger white dusting, giving the paler areas a faint 
bluish tint; transverse lines and dark markings more 
strongly contrasted; the white antemedial line especially 
well marked on its lower half, its inner black border 
below cell expanded into a moderately wider blackish 
band or patch; blackish inner and outer borders of sub- 
terminal hne well marked, especially the former; discal 
dots conspicuous, tending to coalesce. Hind wing 
subpellucid smoky white, darkening towards apex and 
termen; the veins very faintly darkened and a narrow 
dark line along termen. Alar expanse, 24-26 mm. 

Male genitalia with uncus hoodlike, constricted to- 
wards base. Harpe with a strong, outwardly produced, 
spined, knob from base of costa; no appreciable clasper. 
Aedeagus slender, with a very shght bifiu-cation at 
apex; the bifurcate projections straight. Two pairs of 
ventrolateral tufts on eighth abdominal segment; some 
of the hairs broadly expanded at their apices (as in 
aenigmatica, fig. 367c). 

Female genitalia with bursa copulatrix membranous 
except for a very faint sclerotization of the lobe giving 
oflF the ductus seminalis; ductus bursae flattened, 
weakly sclerotized, expanding at genital opening into 
sclerotized, scobinate ventrolateral lobes. 

Type locality: Aweme, Manitoba, Canada (type in 
Canadian Nat. Coll.; paratypes in USNM, 61352). 

Food plant: Unknown. 

Described from male type and four male and two 
female paratypes from the type locality, collected by 
Norman Criddle July 13 and Aug. 10, 1925; Aug. 10, 
1921; Aug. 10, 1925; Aug. 19, 1915). 

293. Pyla aenigmatica, new species 
Figures 367, 853 

Maxillary palpus of male in the form of a semiaigrette 
(the hairs short). 

Forewing as on insinuatrix except darker, the ground 
color Hke that of impostor; the transverse lines distinct; 
lower half of antemedial line bordered inwardly by a 
subquadrate blackish patch, costal half of the outer 
dark border rather broad and well contrasted, blackish; 
the dark borders of subterminal line well contrasted 
towards costa; discal dots confluent; dots along termen 
minute, weak. Alar expanse, 25-28 mm. 

Male genitalia with tegumen having two, strongly 
spined, protruding lobes on each ventrolateral margin. 
Harpe mth base of costa enlarged and coarsely scobi- 
nate; clasper smaU, semicircular, erect. Aedeagus 
slender; shortly bifurcate; bifurcate elements at apex 
spinehke, bent sharply at right angles to the aedeagus. 
Anellus a semitubular shield with rather long, strongly 
sclerotized lateral arms. Eighth abdominal segment 
with two pairs of ventrolateral hair tufts; one pair with 
the hairs expanded at their apices (forming knobhke 
clusters). 



Female genitalia with bursa membranous. Ductus 
bursae short, strongly sclerotized along lateral margins, 
the sclerotizations expanding laterally and at right 
angles at genital opening into a pair of convolute, finely 
scobinate lobes. 

Type locality: Wellington, British Columbia (type 
in USNM, 61353; paratype in Canadian Nat. Coll.). 

Food plant: Unknown. 

Described from male type and one male paratype 
from the type locaUty, "21-VI-04," G. W. Taylor, and 
paratypes from the following localities: Goldstream, 
British Columbia, "30-VIII-20" (9); Salmon Arm, 
British Columbia, "22-6-20, W. R. B." (cf); Pine 
Grove, Colo., July 8, 1901, H. G. Dyar, "17310" (cf); 
East River, Conn., Aug. 21 and Sept. 3, 1908, C. R. 
Ely (cf and 9 ); Oak Station, Pa., Aug. 20, 1911, Fred 
Marloff (cf); Watchung Mts., N. J., "6^-99," W. D. 
Kearfott (cf). 

The hind wings are a trifle darker on the eastern 
examples, which were, in our collection, identified as 
Jusca. 

The species is evidently closely related to insinuatrix, 
but is distinct and easily distinguished by its genitalia. 

294. Pyla criddlella Dyar 
Figure 368 

Pyla criddlella Dyar, Journ. New York Ent. Soc, vol. 15, p. 110, 
1907.— McDunnough, Check list. No. 6241. 

Maxillary palpus of male squamous. 

Forewing blackish gray, semilustrous, unicolorous 
except for a shght darkening of the ground color border- 
ing the transverse lines; the latter very faint, but slightly 
lighter than the groimd color; discal and terminal dots 
obscured. Hind wing brownish gray. Alar expanse, 
18 mm. 

Male genitalia with harpe simple except for a greatly 
reduced, upcurving, triangulate clasper. Aedeagus 
deeply bifurcate; the right divided element (in ventral 
view) produced into a sharp abruptly curved hook. A 
single pair of ventrolateral hair tufts on eighth abdom- 
inal segment. 

Type locality: Aweme, Manitoba, Canada (June ; 
type in USNM). 

Food Plant: Unknown. 

Known only from the male type. 

295. Pyla fusca (Haworth), new combination 
Figures 369, 852 

Phycis fusca Haworth, Lepidoptera Brittaniea, pt. 3, p. 493, 1828. 

Phycita fusca (Haworth) Stephens, Illustrations of British ento- 
mology, Haustellata, vol. 4, p. 310, 1834. 

Pempelia fusca (Haworth) Stainton, Manual of British butterflies 
and moths, vol. 2, p. 176, 1859. — Packard, Ann. Lyo. Nat. 
Hist. New York, vol. 10, p. 271, 1873. 

Nephopteryx moestella Walker, List, vol. 27, p. 53, 1863. 

Eudorea (?) frigidella Packard, Proo. Boston Soc. Nat, Hist., 
vol. 11, p. 53, 1866. 

Salehria fusca (Haworth) Heinemann, Die Schmetterlinge 
Deutschlanda und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 156, 
1865.— Grote, BuU. U. S. Geol. Geogr. Surv. Terr., vol. 4, 
p. 695, 1878; North Amer. Ent., vol. 1, p. 11, 1879.— 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



145 



Staudinger and Rebel, Catalog der Lepidopteren dea palae- 
arctischen Faunengebietes, vol. 2, p. 34, 1901. — Spuler, Die 
Schmetterlinge Europas, vol. 2, p. 211, 1910. — Meyrick, 
Revised handbook of British Lepidoptera, p. 380, 1928. — 
Ford, Guide to the smaller British Lepidoptera, p. 10, 1949. 

Pinipestis cacabella Hulst, Ent. Amer., vol. 3, p. 133, 1887. 

Laodamia fusca (Haworth) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 408, 1893.— Hulst, Phycitidae 
of N. Amer., p. 156, 1890; U. S. Nat. Mus. Bull. 52, p. 425, 
1902.— Forbes, Cornell Mem. 68, p. 628, 1923.— McDun- 
nough. Check list. No. 6227, 1939. 

Salebria triplagiatella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 109, 1904. — Barnes and McDunnough, Contributions, 
vol. 3, p. 196, 1916. 

Dioryclria fusca (Haworth) Pierce and Metcalfe, The genitalia of 
the British Pyrales, p. 3, pi. 2, 1938. 

Maxillary palpus of male in the form of an aigrette. 

Forewing blackish gray, generally of a duskier hue 
than that of any of the preceding species; transverse 
lines usually faint and a dull whitish gray, rarely con- 
trasted against the ground color and when so, chiefly 
the lower half of antemedial line; the latter bordered 
outwardly at costa and inwardly at inner margin by 
blackish patches more or less contrasted against the 
ground color of the wing; a similar dark shade inwardly 
bordering the subterminal line ; discal and terminal dots 
tending to coalesce, black. Hind wing dusky white 
between the veins; the latter appreciably darkened; a 
smoky shade along termen. Alar expanse, 25-30 mm. 

Male genitalia with harpe simple except for a thin, 
saucer-shaped, erect clasper with a toothlike projection 
from its upper inner angle. Aedeagus slender with its 
anterior end abruptly expanded and anterior margin 
straight (as in hypochalciella) ; one side produced into 
an extended, strongly sclerotized arm, sharply bent and 
pointed at apex (as in criddlella) ^ ; penis armed with a 
single, moderately long, hairlike cornutus. Three pairs 
of ventrolateral hair tufts on eighth abdominal segment 
of male, some of the hairs broadly expanded at their 
apices. 

Female genitalia with bursa membranous; ductus 
bursae sclerotized for most of its length, the sclerotiza- 
tions extending for a short distance into the bursa; 
genital opening simple except for some weak granula- 
tions on and behind the ductus bursae. 

Type locahties : England (Jusca, in BM) ; eastern 
Canada {moestella, in BM) ; Caribou Is., Labrador 
(frigidella, in MCZ); "New York"' (cacabella, in 
AMNH, ex Rutgers); Winnipeg, Manitoba {triplagia- 
tella, m USNM). 

Food plants: Erica and probably some other Erica- 
ceae. The only authentic Old World record is Erica 
(Meyrick 1938, Ford 1949). Ragonot (Monograph, 
p. 408) records Vaccinium myrtellus and Salix caprea as 
probabilities; but these plants only on the basis of food 
accepted in the laboratory by larvae hatched from eggs 
from gravid females by Porritt (Ent. Monthly Mag., 

' This extension of aedeagus was misidentified by Pierce and 
Metcalfe (1938) as a cornutus. They overlooked the true 
cornutus attached to the vesica. 

• So given in Hulst's original description. The male type, how- 
ever, bears no locality label. 



vol. 19, p. 11, 1882). A female in the U. S. National 
Museum from Ottawa, Canada, was reared by James 
Fletcher (Aug. 1889) from a "black larva" found on 
Betula. I suspect, however, that the larva had migrated 
to that plant. We have no other New World rearing 
records. 

Distribution: Holarctic. In the Old World from 
Great Britain to Japan. The American records from 
specimens are: United States: Maine, Orono; New 
Hampshire, Hampton (June) , Mount Washington (July) ; 
Massachusetts, Framingham (July) , Martha's Vineyard 
(Aug.) ; New York, Rochester (June), Waterville (Aug.) ; 
Colorado, Glenwood Springs (Aug.); Washington, Pull- 
man. Canada : Newfoundland, Port aux Basque (Aug.) , 
St. George Bay (Harry's River and Stephenville, Aug.), 
Spruce Brook {Km^.) ; Labrador , Caribou Isl., Hopedale, 
Nain; Nova Scotia, Baddeck (Cape Breton Isl., Aug.); 
Quebec, Chelesea (May); Ontario, Albany River (St. 
Martin's Falls), Hymers (Aug.), Ottawa (June, Aug.); 
Manitoba, Aweme (June, July, Aug.), Winnpieg;.4Z5erta, 
Banff (July), Calgary (Aug.), Edmonton (May) ; British 
Columbia, Eraser Mills (June), Kaslo (July, Aug.), 
Shawnigan Lake (Aug.), Victoria (July). Alaska: 
Cordova, Fort Yukon, Juneau (July), Rampart (July). 

The species can be readily distinguished by its aede- 
agus, threadlike cornutus, and the peculiar sclerotization 
of its ductus bursae. On habitus and aU its structural 
characters it is closely related to the gray-winged 
species of Pyla. Superficially it could easily be con- 
fused with impostor, equivoca, or aenigmatica. It is not 
congeneric with. Jaecella (ZeUer), the type of Laodamia, 
to which genus Ragonot referred it. The latter differs 
markedly in male and female genitalia (figs. 427 and 
885), and on venation falls into our venational group D. 
Both Pyla and Laodamia have the cell of hind wing 
short; but in Laodamia vein 3 is appreciably longer in 
relation to vein 2 (fig. 52) . 

Packard's Jrigidella was retained by Ragonot as a 
separable variety from /wsca, but it is at most only one 
of its color variants and is not entitled to any trinomial 
designation as a race. 

Several Old World references and synonyms have 
been omitted from the above synonymy. I do not 
question them, but have not been able to verify them. 
Anyone interested will find the names and references in 
Hulst (Phycitidae of N. Amer., 1890) and Ragonot 
(Monograph, 1893). 

296. Pyla hypochalciella (Ragonot), new combination 
Figures 370, 854 

Nephopteryx ovalis hypochalciella Ragonot, N. Amer. Phycitidae, 
p. 7, 1887.— Hulst, Phycitidae of N. Amer., p. 144, 1890. 

Nephopteryx hypochalciella Ragonot, Monograph, pt. 1, p. 272, 
1893.— McDunnough, Check list. No. 6168, 1939. 

Pyla blackmorella Dyar, Ins. Insc. Menstr., vol. 9, p. 68, 1921. — 
McDunnough, Check list. No. 6248, 1939. (New syn- 
onymy.) 

Maxillary palpus of male in the form of an aigrette. 

Forewing very dark gray-brown, the dark areas of 

some of the darkest specimens almost black;a veiyfaint 



146 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



powdering of white on costal half of median area and, 
narrowly, along terminal margin; the transverse Hnes 
powdery, grayish white, obscm-e on some specimens; 
antemedial Hne obhque, expanded slightly towards costa, 
without distinct inner dark border and with but faint 
indication of a blackish brown outer bordering shade 
towards costa; subterminal hne more or less obscured, on 
weU-marked examples preceded by a thin blackish line 
and followed by a broad band of the darkest prevailing 
ground color; discal dots black, well separated; a row of 
small black dots along termen, tending to fuse and on a 
few specimens forming a fine black line. Hind wing a 
uniform very dark satiny brown; the cilia paler, shading 
from pale brown to white at their tips. Alar expanse, 
22-26 nam. 

Male genitalia ha\dng harpe with an erect clasper 
armed along its outer margin with a row of stout spines 
(in the figure this looks like an enlargement of the base 
of costa, but it arises below costa and the base of the 
costa itseK is simple). Anellus bearing two pairs of 
combUke, heavy spines, one pair ventral, one dorsal, the 
latter situated behind the former. Aedeagus slender; 
its anterior end abruptly expanded and the anterior 
margin straight; apical fourth bifid, the divided elements 
terminating in laterally curved horns. A single pair 
of simple hair tufts on eighth abdominal segment of 
male. 

Female genitalia with bursa membranous; ductus 
bursae partially flattened, sclerotized throughout, the 
sclerotization expanding abruptly into a wide funnel at 
genital opening. 

Type localities: "Washington Territory" Qiypo- 
chaldella, in Paris Mus.) ; Mount Tzouhalem, southern 
Vancouver Isl., British Columbia {blackmorella, in 
USNM). 

Food plant: Unknown. 

Disteibution: United States: Washington, Friday 
Harbor (Jime, July). Canada: British Columbia (south- 
em Vancouver Isl.), Cowichan District (June), Dimcans 
(June), Mount Malahat (June), Moimt Tzouhalem 
(June). 

The species is easily identified by its peculiarly armed 
anellus. In ground color of forewing it and the species 
following (hanhamella) are intermediate between typical 
gray- and brown-winged members of the two Pyla 
species groups. However, except for a very faint trace 
of it in hanhamella, they both lack the bronzy luster on 
forewing so characteristic of the typical brown group. 

297. Pyla hanhamella Dyar 
FiGUBBs 371, 860 

Pyla hanhamella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 109, 
1904.— McDunnough, Check list, No. 6239, 1939. 

Maxillary palpus of male subsquamous (a short brush 
of hairs mixed with flattened scales) . 

Forewing color and markings as in Kypochaldella 
except for traces of a metallic sheen at base. Hind 
wing pale brown; ciha white with a fine dark subbasal 
line. Alar expanse, 20-24 mm. 



Male genitalia having clasper of harpe a moderately 
long, erect spike; costa at base simple (not produced). 
Aedeagus slightly bent towards middle; shortly bifid at 
apex, the divided elements coarsely scobinate. A single 
pair of simple hair tufts on eighth abdominal segment. 

Female genitalia with the lobe of bursa giving off the 
ductus seminaUs partially sclerotized, otherwise mem- 
branous; ductus bursae flattened, sclerotized through- 
out, concavely bent at middle, the sclerotization termi- 
nating in a sinuate, thickened, narrow, hplike band 
along the lower margin of the genital opening. 

Type locality: Winnipeg, Manitoba (type in 
USNM). 

Food plant: Unknown. 

Distribution: Manitoba, Aweme (Jime, July), Win- 
nipeg (June). 

Genus Pyla, Species 298-306: P. sdntillans to 
P. viridisuffusella 

[Ground color of forewing bronzy brown.] 

298. Pyla scintUlans (Grote) 
FiGXJBBS 29, 372, 373, 857 

Nephopteryx sdntillans Grote, Papilio, vol. 1, p. 18, 1881. 

Pyla sdntillans (Grote), New check list of North American 
moths, p. 56, 1882. — Hulst, Phycitidae of N. Amer., p. 161, 
1890. — Ragonot, Monograph, pt. 1, p. 482, 1893. — McDun- 
nough, Check list, No. 6235, 1939. 

Pyla feella Dyar, Ins. Insc. Menstr., vol. 9, p. 68, 1921. — Mc- 
Dunnough, Check list, No. 6247, 1939. (New synonymy.) 

MaxiUary palpus of male subsquamous (the scales 
short, flattened, forming a small expanded brush). 

Forewing dark bronzy brown; the scaling shiny, 
metallic; transverse lines absent, indicated only on well 
marked specimens by very faint, moderately broad, 
blackish brown bands (vestiges of their dark borders) ; 
discal and terminal dots obsolete. Alar expanse, 
20-26 mm. 

Male genitalia having harpe with strong clasper, 
developed as a stout outwardly ciu-ved hook with an 
extended, elongate, bladelike base, the latter more or 
less sen-ate. Considerable individual variation is shown 
in the clasper and the shape of its base. In one example, 
from Inyo County (presximably a variety of sdntillans 
but possibly a distinct species), the clasper hook is 
markedly longer than in the examples figured, and the 
bladelike base narrower. Costa of harpe produced at 
base into a pointed, stout, very coarsely spined projec- 
tion. Aedeagus bifid for less than half its length; one 
of the divided elements with a short, thornhke spine 
projecting from lateral margin before apex; the other 
with 2 or 3 similar spines from lateral margin near apex 
(usually 3, rarely 2, a single specimen from El Dorado 
County, exhibiting only one) . A single pair of ventro- 
lateral abdominal hair tufts on eighth segment. 

Female genitalia with bursa small, membranous 
throughout; ductus bursae very short, expanded abruptly 
into a sclerotized cup, its lower surface developed as a 
pair of flattened, pointed, elongate-oval blades which 
pro j ect beyond genital opening. Only trifling individual 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



147 



differences cau be distinguished between Dyar's jeella 
and females from other CaHfornia localities. 

Type localities: Summit, Sierra Nevada Mts., 
Calif, {scintillans, in BM); Bullfrog Lake (10,634 ft.), 
Sierra Nevada Mts., Calif, (feella, in USNM). 

Food plant: Unknown. 

Distribution: California, Cisco (Placer County, 
July), Deer Park Springs (Lake Tahoe, July), El 
Dorado County (July), Inyo County (July), Mineral- 
king (Tulare County, July, Aug.), Sierra Nevada Mts. 
(Bullfrog Lake and Summit, Aug.), Tuolumne Meadows 

(July). 

299. Pyla sylphiella Dyar 
Figures 375, 858 

Pyla sylphiella Dyar, Ins. Insc. Menstr., vol. 9, p. 68, 1921. — 
McDunnough, Check list. No. 6246, 1939. 

Maxillary palpus of male subsquamous. 

Superficially like scintillans, averaging a trifle darker; 
but distinguished only by its genitalia. Alar expanse, 
19-25 mm. 

Male genitalia with clasper of harpe similar to that 
of scintillans; produced enlargement of base of costa 
considerably stouter and more coarsely spined. Aede- 
agus with only a pair of lateral spines from adedeagus 
near its apex (one spine from each of the divided 
elements opposite and pointed away from each other). 
These differences are slight but appear to be consistent 
through long series. Female genitalia with ventral 
surface of the cup-shaped portion of ductus bursae 
bent into broad, deep, strongly sclerotized folds. 

Type locality: Mount Kainier, Wash, (type in 
USNM). 

Food plant: Unknown. 

Distribution: United States: Washington, Movmt 
Rainier (Aug.), Paradise Valley (Moimt Rainier, 
Aug.), Sheep Lake (Yakima County, Aug.), Skyline 
Ridge (Moimt Baker District, Aug.), Slate Peak 
(Whatcom Comity, Aug.). Canada: British Columbia, 
Mount Cheam (Aug.), Mount McLean (Aug.). 

The species is very close to scintillans but apparently 
distinct. The male genitalia differ only in minor details 
and the color and maculation offer little if anything to 
separate the two ; but the female genitalia are markedly 
different and, from the specimens available, sylphiella 
appears to have a more northerly distribution. 

300. Pyla rainierella Dyar 

FiQUBEs 374, 859 

Pyla rainierella Dyar, Proc. Ent. See. Washington, vol. 6, p. 109, 
1904.— McDunnough, Check list, No. 6243, 1939. 

Maxillary palpus of male subsquamous. 

Moths averaging a trifle smaller than the preceding 
species (sylphiella), but certainly distinguished from it 
only by genitalia. Alar expanse, 16-20 mm. 

Male genitalia with production from base of costa 
of harpe considerably smaller and less coarsely spined 
than that of either sylphiella or scintillans. Aedeagus 
short, the apices of its divided elements bent abruptly 
downward as sharp, parallel, spinelike hooks. Female 



genitalia with the sclerotized portion of the cup-shaped 
area of ductus bursae developed laterally as triangulate 
plates. 

Type locality: Mount Rainier, Wash, (type in 
USNM). 

Food plant: Unknown. 

Distribution: Washington, Mount Rainier (Aug.), 
Paradise Valley (Mount Rainier, July), Sheep Lake 
(Yakima County, Aug.), Slate Peak (Whatcom 
County, Aug.), Table Moimtain (Aug.). 

Like sylphiella, this species is chiefly distinguished by 
its female genitalia, the sclerotized area of ductus 
bursae at genital opening resembles somewhat that of 
Jasciella but differs in shape and is like that of no other 
species in the genus. The aedeagus easily separates 
the male oi Jasciella from either scintillans or sylphiella 

301. Pyla aeneella Hulst 
Figures 376, 864 

Pyla aeneella Hulst, Canadian Ent., vol. 27, p. 55, 1895. — Mc- 
Dunnough, Check list, No. 6242, 1939. 

Maxillary palpus of male in the form of a semiaigrette 
(the hairs short) . 

Forewing unicolorous, without any trace of dark 
transverse shadings or discal spots; brown with a bronzy 
green irridescence. Hind wing concolorous with fore- 
wing. Alar expanse 23-25 mm. 

Male genitalia with costa of harpe at base produced 
into a thin, rounded lobe with finely serrate edge; 
clasper erect, short, stout, thornhke. Aedeagus short, 
broadest at middle, divided to middle; the divided ele- 
ments rather broadly flattened and abruptly, asym- 
metrically bent at their apices. Female genitaUa with 
ductus bursae broadly cup-shaped from shortly beyond 
its junction with bursa; the ventral surface weakly 
sclerotized, granulate, and with slight infoldings on the 
lower median area. 

Type locality: Colorado (in AMNH, ex Rutgers). 

Distribution: Colorado; Utah, Silver Falls (July), 
Stockton (May, June). 

A good series of the Utah specimens is in the National 
Collection. The genitalia of the females agree in every 
detail with those of the Colorado type in the Rutgers 
Collection. The metaUic iridescence of the forewings 
is conspicuous but its greenish tint is very faint. 

302. Pyla aeneoviridella Ragonot 

Figures 378, 862 

Pyla aeneoviridella Ragonot, N. Amer. Phycitidae, p. 9, 1887; 
Monograph, pt. 1, p. 482, 1893.— Hulst, Phycitidae of N. 
Amer., p. 161, 1890. — Barnes and McDunnough, Contribu- 
tions, vol. 2, p. 222, 1914. — McDunnough, Check list. No. 
6237, 1939. 

Maxillaiy palpus in the form of a semiaigrette (the 
scales somewhat flattened, not so decidedly hairlike as 
in aeneella) . 

Forewing unicolorous bronzy brown, without dark 
markings of any kind; the iridescent scaling with a 
faint greenish tint and somewhat more strongly con- 
centrated at the base of the wing than in the median 



148 



TXNITED STATES NATIONAL MUSEUM BULLETIN 207 



and outer areas. The color of both the fore and hind 
wings is similar to that of aeneella and the two species 
can only be safely distinguished by their genitalia. 
Alar expanse, 23-29 mm. 

Male genitalia with an angulate, serrate and coarsely 
spined projection from costal base of harpe; clasper out- 
bent, strongly spined and more or less serrate. Aedea- 
gus short, slightly bent at middle and with a single stout, 
stubby spine projecting from one side (extent of indi- 
vidual variation shown in figs. 378a, b). Female geni- 
taUa with a weak sclerotization of the lobe of bursa 
giving off the ductus seminaHs; ductus bursae with 
ventral surface of cup-shaped area more strongly 
sclerotized than that of aeneella and differently sculp- 
tured. 

Type locality : Evanston, Wyo. (type in Paris Mus.) • 

Food plant: Unknown. 

Distbibution: United States: Colorado, Tennessee 
Pass (July); Wyoming, Big Horn Mts. (July), 
Evanston, Yellowstone Park (July, Aug.); Montana, 
Bozeman (July), Glacier Park (June); Oregon, Wallowa 
Mts. (Arnold Lake, July); Washington, Olympic Mts. 
(Hurricane Ridge, June, July). Canada: Alberta, 
Laggan (July). 

The Washington and Oregon specimens are consider- 
ably darker than those from the other locaUties, the 
specimens from Oregon having almost black hind wings 
and blackish brown forewings. 

In his original description and in his Monograph 
Ragonot gives "N. Y." as the type locaUty. This was 
a misreading of the label of his type. The correction 
was made by Barnes and McDunnough in the reference 
cited above. 

303. Pyla metalicella HiJst 

Figures 377, 863 

Pyla metalicella Hulst, Canadian Ent., vol. 27, p. 64, 1895. — 
McDunnough, Check list, No. 6236, 1939. 

Maxillary palpus of male in the form of a semiaigrette 
(the hairs rather short). 

Appreciably lighter than aeneoviridella. The fore- 
wing a unicolorous light bronzy brown with greenish 
yellow iridescence; discal dots more or less distinct, 
separated, blackish; no other markings. Average size 
larger than that of aeneoviridella. Alar expanse, 25-32 
mm. 

Male genitalia with a long, somewhat flattened, out- 
wardly curved clasper on harpe (similar to that of 
impostor, but proportionally longer) ; costa of harpe at 
base simple. Aedeagus very shortly divided at apex; 
a pair of very short, sharp, straight spines on ventral 
surface near apex. 

Female genitalia with lobe of bittsa giving off ductus 
seminalis weakly sclerotized; cuplike area of ductus 
bursae, funnel shaped (triangulate) , strongly sclerotized 
over its entire ventral surface and containing a broad, 
centrally located, funnel-shaped fold. Individual vari- 
ations in this fold are shown in figs. 863 and 863a. 

Type locality: Colorado (type in AMNH, ex 
Rutgers) . 



Food plant: Unknown. 

Distribution: Colorado, Silverton (July, Aug.), and 
two specimens with only the state locality; Utah, SUver 
Lake (July). The Silverton locahty is represented in 
the National Collection by 17 specimens. Also in the 
National Collection is a female from Colorado (Bruce), 
labeled "Pyla aeneella Hulst, Type," another of Hulst's 
pseudotypes and possibly part of his original "type" 
series of aeneella. The actual type of metalicella is a 
male with only the state locality. Its genitalia agree 
in every detail with those from Silverton specimens. 

304, Pyla fasciella Barnes and McDunnough 

FiGTTBES 379, 861 

Pyla fasciella Barnes and McDunnough, Canadian Ent., vol. 49, 
p. 405, 1917.— McDunnough, Check list. No. 6244, 1939. 

Maxillary palpus of male minute, squamous. 

Forewing blue-black, some paler bluish iridescence 
over basal area and (in strong light) a faint, brownish 
iridescence in outer area; antemedial line represented 
by a narrow oblique black band near, but before middle; 
subterminal line a similar curved band well back from 
and parallel with termen; the area between the two 
black bands darker than remainder of wing, forming a 
faint, broad, median, black fascia; discal dots obsolete. 
Hind wings very dark brown, shiny. Alar expanse, 
21-24 mm. 

Male genitalia without clasper on harpe; base of 
costa of harpe produced into a knoblike projection, 
finely spinose along margin. Aedeagus simple. A 
single pair of ventrolateral hair tufts on eighth segment. 
Female genitalia similar to those of rainierella except 
that the paired plates of ductus bm-sae at genital open- 
ing are narrower and differently shaped. 

Type locality: Mount Shasta, Calif, (type in 
USNM). 

Food plant: Unknown. 

Known only from northern California. Represented 
in the National Collection by the type series from 
Mount Shasta, 7,000 ft., July — three males and two 
females (not four males and two females as given in the 
original description) ; and one male from Bartle, Cahf . 
(June 14, 1939, Grace H. and John L. Sperry). In 
their original description the authors give the expanse 
as "24-31 mm." This is probably a printer's error, 
for the largest specimen before me is a scant 24 mm. 

305. Pyla nigricula, new species 
FlGTJBB 380 

Maxillary palpus of male minute, squamous. 

Superficially like fascieUa except: Transverse dark 
lines of forewing obsolete, only the antemedial black 
band very faintly indicated; no contrasted dark median 
fascia; the entire median and outer areas a dark 
purplish brown. Alar expanse, 26 mm. 

Male genitalia with uncus broader and squattier than 
that oi jasciella. Projection from costal base of harpe 
differently shaped, bluntly pointed; clasper developed 
as a stout, smooth, curved, pointed hook. Aedeagus 
simple. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



149 



Type locality: Verdi, Nev. (type in USNM, 61354). 

Food plant: Unknown. 

Described from unique male collected by A. H. 
VachelJ, "June 1 to 10." This specimen had been in 
our collection under scintillans Grote. 

306. Pyla TiridigufTusella Barnes and McDunnough 

Figures 381, 851 

Pyla viridisuffuiella Barnes and McDunnough, Canadian Ent., 
vol. 49, p. 406, 1917.— McDunnough, Check list. No. 6245, 
1939. 

Maxillary palpus of male in the form of a short 
aigrette. 

Forewing heavily suffused with light, bronzy green, 
irridescent scaling, especially strong over basal area and 
in a line indicating the subterminal line; two transverse 
blackish bands, an oblique, antemedial one and another 
forming an inner border to the subterminal line; discal 
dots, when distinguishable, confluent, forming a line 
along discocelMar vein. Hind wing very dark brown. 
Alar expanse, 17-20 mm. 

Male genitalia with harpe simple. Aedeagus bifur- 
cate to middle, the divided elements asymmetrical (one 
longer than the other). Female genitalia with a pair 
of strongly sclerotized, convolute bands extending from 
posterior end of bursa well into ductus bursae; genital 
opening simple. 

Type locality: Tuolumne Meadows, Tuolumne 
County, Calif, (type in USNM). 

Food plant: Unknown. 

Distribution: California, Humphreys Basin (Fresno 
County, Aug.), Johnsons Park (Sierra Nevada Mts.), 
Kernick Meadows (9,250 ft., July), Mineralking (Tulare 
County, July, Aug.), Tuolumne Meadows (July, Aug.). 

The most brilliant of the Pyla species. Its genitalia, 
both male and female most resemble those of fasciolalis 
Hulst. 



Genera 77 and 78: Diorydria and Orydometopia 

IVenational division D. Forewing with veins 4 and 6 closely 
approximate for a short distance from cell {Dioryctria) , or connate 
or very shortly stalked {Oryctometopia) ; vein 6 straight or bent 
towards base; 10 from the cell. Hind wing with cell less than 
one-third the length of wing; discocellular vein curved; veins 4 
and 5 stalked for at least half their lengths. Male antenna with 
a shallow sinus or slight incurvation in base of shaft, containing 
a row of fine spines or a weak scale tuft. Male genitaha with 
transtiUa incomplete or absent; harpe with costa strongly scle- 
rotized and produced at apex (Dioryctria) or with one or more 
short, stout, thornlike spines from lower margin of sacculus.] 

77. Genus Dioryctria Zeller 

Dioryctria Zeller, Isis von Oken, 1846, p. 632. — Heinemann, Die 
Schmetterlinge Deutschlands und der Schweiz, Abt. 2, vol. 
1, pt. 2, p. 148, 1865.— Ragonot, Ent. Monthly Mag., vol. 
22, pp. 52, 56, 1885; Monograph, pt. 1, p. 187, 1893.— Hulst, 
Phycitidae of N. Amer., p. 134, 1890.— Spuler, Die Schmet- 
terlinge Europas, vol. 2, p. 213, 1910. — Forbes, Cornell Univ. 
Agr. Exp. Station, Mem. 68, p. 619, 1923. — Meyrick, Re- 
vised Handbook of British Lepidoptera, p. 383, 1928. — 
Bisset, in Pierce and Metcalfe, Genitalia of the British 



Pyrales, p. 57, 1938 (notes Ragonot's fixation (1885) of 
type of genus) .—Janse, Journ. Ent. Soc. South Africa, vol. 
4, p. 161, 1941. (Type of genus: Tinea abietella Denis and 
SchifFermtiller.) 
Pinipestis Grote, Canadian Ent. vol. 10, p. 19, 1878; Bull. U. 8. 
Geol. Geogr. Surv. Terr., vol. 4, p. 699, 1878.— Hulst, 
Phycitidae of N. Amer., p. 136, 1890. (Type of genus: 
Nephopteryx zimmermani Grote.) 

Tongue well developed. Antenna finely pubescent 
or very shortly cihate, rarely unipectinate (in males of 
some Old World species, pineae Staudinger, mendacella 
Standinger); on aU males a shallow sinus in base of 
shaft, containing a short row of minute black, thornlike 
spines, more or less concealed by rough scaling. Labial 
palpus upturned, reaching to or a trifle above vertex; 
second segment grooved on inner side; third segment 
short (less than one-third the length of second), acumi- 
nate. Maxillary palpus of male small and squamous or 
(rarely) in the form of an aigrette. Forewing smooth 
or with two or more tufts or raised (ruffed) scales; 11 
veins ; vein 2 from before lower outer angle of cell ; 3 from 
the angle, slightly nearer to 4 than to 2 at base ; 4 and 5 
from the cell, approximate at and for about one-fourth 
their distance from cell; 6 bent towards base and from 
close to upper angle of cell (in type species, straight and 
from well below the angle on some specimens); 10 from 
the cell, approximate to the stalk of 8-9 for some dis- 
tance beyond ceU; 8 and 9 long stalked; male without 
costal fold. Hind wing with vein 2 from before lower 
outer angle of cell ; 3 from the angle, long (almost as long 
as 2) ; 4 and 5 stalked for half or slightly more than half 
their lengths; 7 and 8 closely approximate for less than 
half their lengths beyond ceU; rarely shortly anasto- 
mosed; cell less than one-third the length of wing; dis- 
cocellular vein curved, outwardly extended at lower 
angle. Eighth abdominal segment of male with com- 
pound ventral scale tufts. 

Male genitalia decidedly elongated (least so in gido- 
sella). Uncus broad, stout, its terminal margin more or 
less broadly rounded. Apical process of gnathos a 
short, stout hook. TranstiUa incomplete or absent; its 
lateral elements, when distinguishable, slender and 
usually attached to costal base of harpe. Harpe with 
costal area broadly sclerotized and produced at apex; 
cucullus narrowly elongate, bluntly pointed or very 
narrowly rounded at apex; sacculus short, simple; 
clasper present, erect, usuaUy finely haired at or near 
apex, but not bearing strong spines, thorns, or serra- 
tions. Anellus with well-developed lateral lobes. Ae- 
deagus long, moderately stout; penis with strong cor- 
nuti, consisting of numerous straight, slender spines (as 
long or nearly as long as width of aedeagus) and usually 
one or more longer, stouter spines placed back of them 
on the penis. Vinculum stout; longer than broad 
(frequently considerably elongated), gradually tapering 
to a moderately broad, rounded, or abruptly angled 
terminal margin. 

Female genitalia with well-developed signa, consisting 
of two or three clusters of strong, slender spines, their 
bases in each of the clusters more or less fused into 
sclerotized plates; the clusters in end of bursa near 



150 



UNITED STATES NATIONAL MUSEUM BUliLETIN 207 



ductus bursae and one of them frequently in or extend- 
ing into the ductus; ductus bursae flattened, strongly 
sclerotized over most of its length, the sclerotization 
more or less longitudinally wriukled and terminating just 
before the simple genital opening; ductus semiualis from 
bursa near the jimction of bursa and ductus bursae. 

Dioryctria is one of the most, if not the most, distinct 
and sharply defined genus in the Phycitidae; and is so 
despite the variations in structure exhibited by its male 
antennae, male maxillary palpi, and its wing scaling. 
Its male and female genitalia have a characteristic 
habitus, difficult to describe, but easUy observed in slides 
or figures. There is also a maculation character of the 
forewing common to nearly all the species; the usual 
black dots at end of cell are absent, being replaced by a 
white spot or line on the discocellular vein. The only 
North American species without such a marking is 
clarioralis, where a considerable area about the end of 
cell is clouded with a dark suffusion. The raised-scale 
character upon which Grote distinguished his genus 
Pinipestis fi'om Dioryctria consists of a mere ruffing of 
the scales on two or three spots on the forewing. It is 
at best an elusive character, subject to opinion as much 
as to observation, and in some of the forms (simmermam, 
cambiicola) the tufts may be either raised or flattened on 
unspread specimens. Naturally, on spread specimens 
they are usually flattened, whatever was their condition 
in nature. Kagonot was perfectly justified (1893) in 
relegating Pinipestis to synonymy. 

As here defined the genus is of world-wide distribution 
in the Northern Hemisphere and all the species whose 
larval habits are Icnown are borers in conifers. Most of 
the American species are represented in the National 
Collection by numerous specimens reared in connection 
with the forest-insect investigations of the U. S. Biu'eau 
of Entomology and Plant Quarantine. 

307. Dioryctria abietella (Denis and Schiffermuller) 
Figures 61, 382, 865 

Tinea abietella Denis and Schiffermuller, Systematische Ver- 
zeichniss der Schmetterlinge der Wienergegend . . . , p. 138, 
1776. — Fabriclus, Mantissa insectorum . . . , vol. 2, p. 246, 
1787; Entomologica systematica . . . , vol. 3, pt. 2, p. 302, 
1793. 

Tinea decuriella Hiibner, Sammlung europaischer Schmetterlinge, 
p. 35 and Lepidoptera 8, Tineae 2, pi. 11, fig. 74, 1796. 

Phycis abietella (Denis and Schiffermiiller) Zincken, in Germar 
and Zincken, Mag. der Ent., vol. 3, p. 160, 1818. — Treitschke, 
Die Schmetterlinge von Europa, vol. 9, p. 177, 1832. — 
Ratzeburg, Die Forst-Insecten . . . , vol. 2, p. 244, pi. 15, 
fig. 2, 1840. 

Dioryctria abietella (Denis and Schiffermuller) ZeUer, Isis von 
Oken, p. 736, 1846. — Heinemann, Die Schmetterlinge 
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 148, 
1865.— Ragonot, Ent. Monthly Mag., vol. 22, p. 62, 1885; 
Monograph, pt. 1, p. 198, 1893. — Hulst, Phycitidae of N. 
Amer., p. 136, 1890. — Spiiler, Die Schmetterlinge Europas, 
vol. 2, p. 213, 1910. — Joannis, Ann. Soc. Ent. France, vol. 
85, p. 259, 1916; Bull, de la Station de Recherches forestiSrs 
du Nord de I'Afrique, vol. 1, p. 192, 1921. — Forbes, Cornell 
Mem. 68, p. 621, 1923. — Meyrick, Revised handbook of 
British Lepidoptera, p. 384, 1928. — Pierce and Metcalfe, 
Genitalia of the British Pyrales, p. 2, pi. 1, 1938. — Keen, 
U. S. Dep. Agr. Misc. Publ. 273, p. 39, 1938.— McDunnough, 



Check list, No. 6129, 1939.— Janse, Journ. Ent. Soc. South 

Africa, vol. 4, p. 161, 1941. — MacKay, Canadian Ent., vol. 

75, p. 94, 1943.— Craighead, U. S. Dep. Agr. Misc. Publ. 

657, p. 451, 1950. 
Pinipestis abieiivorella Grote, Bull. U. S. Geogr. Geol. Surv. Terr., 

vol. 4, p. 701, 1878. 
Pinipestis reniculella Packard (not Grote), U. S. Dep. Agr. Ent. 

Bull. 13, p. 21, 1887; U. S. Dep. Agr. Fifth Rep. Ent. Comm., 

p. 854, 1890. 
Myelois elegantella Hulst, Canadian Ent., vol. 24, p. 59, 1892. 

MaxiUary palpus of male squamous. 

Forewing smooth; bluish gray, dusted with white, the 
white dusting of variable intensity, when pronounced, 
more or less concentrated in basal area, along the outer 
border of antemedial line, the inner border of subter- 
minal line and, weakly, along terminal margin; trans- 
verse lines white, distinct, narrow; antemedial line 
oblique, inwardly angled at cell and vein lb, preceded 
on costa by a blackish shade and bordered outwardly 
by a black line, and usually preceded by a pale patch 
on inner margin, this often no more than a smear of 
olivaceous white scales and never so conspicuous or 
well contrasted as in renicvlella; subterminal line 
sinuate-angulate, preceded and followed by blackish 
bordering lines; discal mark a white, lunate spot; a fine 
black line along terminal margin. Hind wing dusky 
white, darkened slightly towards outer margin and on 
the veins. Alar expanse, 20-30 mm. 

Male genitalia with uncus subtriangulate and rather 
narrow in normal position; a short, slight incurvation 
of the lateral margins near base, and the lateral margins 
themselves infolded. (When uncus is flattened in prep- 
arations and the lateral folds pushed out, the uncus 
appears as in fig. 382, but never takes the form of the 
flattened unci of the species which have a longer incurva- 
tion of the lateral margins, such as zimmermani). Harpe 
with one or more spines projecting from the terminal 
margin of the sclerotized costal area below its apex 
(there is considerable individual variation in this featiu:e, 
a few examples of which, from small American speci- 
mens, are shown in figs. 382c-e). Penis armed with a 
single stout spine behind anterior spine cluster. 

Female genitalia chiefly distinguished by a longitu- 
dinal flssure on the ventral surface of the sclerotized 
portion of ductus bursae, variations of which are shown 
in figures 865 and 865a. The females of sysstratiotes 
from Guatemala also show traces of such a fissure but 
this species is only doubtfully distinct from abietella. 

Type localities: Austria (abietella, location un- 
known); Germany (decuriella, type lost); Amherst, 
Mass. (abietiwrella, in BM) ; Seattle, Wash, (elegantella, 
in AMNH, ex Rutgers). 

Food plants: Pine, spruce, fir; in the spruces and firs 
(Abies, Psevdotsugha, etc.) chiefly in the cones. The 
favored host seems to be Pinus of which it attacks all 
species. The larvae exhibit a variety of habits. They 
are both primary and secondary. They bore into new 
and otherwise uninfested terminals and into terminals 
that have been attacked by Ehyacionia buoliana or the 
white pine weevil (Pissodes strobi.) They attack both 
healthy and diseased cones. They bore into and feed 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



151 



on the cambium of smooth bark on the trunks and 
branches; and also feed in the galls on any part of the 
tree. 

Distribution: Apparently throughout the range of 
the genus Pinus in the Northern Hemisphere. Ameri- 
can records from specimens (moths) before me as 
follows: United States: Maine, Orono (Aug.); Massa- 
chusetts, Amherst, Framingham (Oct.), Martha's Vine- 
yard (Sept.), Pepperell (Aug.); Connecticut, Bradford, 
Lyme (Aug.), New Haven (June); New York, Long 
Island (Garden City, Oct., Great Neck, July), Warrens- 
burg (Sept.); New Jersey, Lakehurst (May); District 
oj Columbia, Washington (July, Aug.); Florida, Alton 
(June), Eustis (June, July), Gainesville (June), Orlando 
(June), "So. Florida" (June, July, Aug.); Illinois, Dim- 
dee; Nebraska, Halsey (Apr., June, Aug.); Montana, 
Dillon (July), EUiston, Evaro (Mar.); Colorado, Glen- 
wood Springs; Arizona, Prescott (July) ; California, 
Berkeley, Patrick's Creek (Sept.), Sacramento, Shasta 
Retreat (July); Oregon, Ashland (July, Aug., Sept., 
Oct.), Salem (Aug.), Colestin (June), Silver Lake (Aug.), 
Sprague River (Jidy) ; Washington, Hoquiam, Pullman, 
Rock Lake (Whitman County, Sept.); Seattle. Can- 
ada: Labrador, Dublin Shore (Jjunenburg County); 
Quebec, Montreal (June); Saskatchewan, Lutherland 
(June, Aug., Sept.); British Columbia, Kaslo (June). 
Guatemala: A series of males and females in the U. S. 
National Museum, reared from pine cones. May 1927, 
by J. G. Salas and labeled simply "Guatemala, C. A." 

The species is of considerable economic importance, 
especially to young pine trees in our Western States, 
and particularly in reforestation areas. It has an 
extensive literature in the Old World. I have listed 
here only the more important references and have 
omitted purely European synonyms. For additional 
references the reader is referred to Ragonot (Mono- 
graph, p. 198), Hulst (Phycitidae of N. Amer., 1890), 
the Journal of Economic Entomology, and the Review 
of Applied Entomology. The most satisfactory infor- 
mation on life history and larval habits will be found in 
the MacKay (1943) and Craighead (1950) papers. 

308. Dioryctria sysstratiotes Dyar 
Figure 866 

Dioryctria sysstratiotes Dyar, Ins. Inso. Menstr., vol. 7, p. 43, 
1919. 

Forewing smooth, similar to that of abietella except: 
A pale brownish suflFusion in median area forming a 
rather large patch below discal spot; a similar brownish 
shade outwardly bordering subterminal line; the patch 
preceding antemedial line on inner margin, more dis- 
tinct, larger, pale olivaceous brown. Hind wing some- 
what darker, translucent smoky white with a very faint 
brownish tint towards outer margin. 

Alar expanse, 23-28 mm. 

Female genitaha as in abietella except no (or only a 
faint trace of) median ventral cleft in ductus bm-sae. 

Type locality: Cayuga, Guatemala (type in 
USNM). 

Food plant : presumably Pinus. No rearing records. 



Distribution: Guatemala: Cayuga (June), Chejel 
(June), Purulhd (July). 

Known only from females. Doubtfully distinct from 
abietella except as a race or color form. Its exact status 
will have to await discovery of a male. 

309. Dioryctria reniculella (Grote) 
Figures 383, 867 

Pinipestis reniculella Grote, North Amer. Ent., vol. 1, p. 67, 
1880. 

Dioryctria reniculella (Grote), Ragonot, Monograph, pt. 1, p. 200 
1893.— Forbes, Cornell Mem. 68, p. 620, 1923.— McDun" 
nough, Check list. No. 6131, 1939.— Brown, Canada Dep- 
Agr. Publ. 712. Techn. Bull. 31, p. 13, 1941.— MacKay, Cana- 
dian Ent., vol. 75, p. 94, 1943.— Craighead, U. S. Dep. 
Agr. Misc. Publ. 657, p. 451, 1950. 

Maxillary palpus of male squamous. 

Forewing smooth; in color and maculation similar to 
that of abietella except: Ground color pale brownish 
gray; the transverse lines and discal spot more sharply 
contrasted, more distinctly white; usually a rather large 
ohvaceous patch on inner margin preceding the ante- 
medial line; hind wing darker, pale smoky fuscous. 
Alar expanse, 22-26 mm. 

Male genitalia with uncus tonguelike; more elongate 
and narrower than that of any other American species; 
its terminal margin very narrowly rounded; no incur- 
vation of the lateral margins (its shape not appreciably 
altered by flattening in slide preparations). A very 
slight spur from the lower outer angle of the sclerotized 
costal area of harpe, but no other spines from below 
apex of costa. Penis armed only with anterior spine 
cluster; no single stout spine on penis behind the an- 
terior cluster. 

Female genitalia with only one strong spine cluster, 
that at junction of bursa and ductus bursae; girdle of 
spines in biu"sa before junction with ductus, weak, 
broken, and the spines themselves greatly reduced. 

Type locality: New York (type in BM). 

Food plants: Various spruces; rarely in balsam fir 
and tamarack. Reported as occasional in jack pine, 
but such records are doubtful. The larvae feed in 
terminals and cones and to a lesser extent upon the 
foliage of the terminals. 

Distribution: United States: Maine, Blue HUl 
(July), Sebec Lake (July); Connecticut, East River 
(July), New Haven (June, Jidy); New York; Illinois, 
Putnam County (July) ; Michigan, East Lansing (Aug.) ; 
Colorado, Estes Park (July); California, Fallen Leaf 
Lake (Aug.), Moimt Lowe (July) ; Washington, Belling- 
ham (Aug.), Hoquiam, Kamiack Butte (Aug.), Pullman 
(July, Aug.). Canada: Nova Scotia, Cape Breton 
(Aug.); Quebec, St. Therese Isl. (St. John's County, 
July) ; Ontario, Westree ; Saskatchewan (June) ; British 
Columbia, Seton Lake (June, July), Victoria (July)- 

The species is easily distinguished by its genitalia. 
In the past it has been frequently confused with 
abietella and until 1893 was treated by Hulst and 
Ragonot as a synonym of the latter. The larvae of the 
two species often occur together in spruce cones, so 
there is no safe way to separate them on their larval 



152 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



habits. However, renicvlella is primarily a spruce 
feeder, while abietella shows a marked preference for 
the pines. Both species are borers, as are all the 
Dioryctria species. It is unfortunate that Miss Mac- 
Kay in her otherwise excellent paper (1943) should refer 
to renicvlella as "The spruce foliage worm." It does 
feed to some extent upon the foliage of terminal shoots; 
but likewise, and more frequently within the terminals 
themselves and in the cones, and in economic literature 
is quite properly referred to as "the spruce cone moth." 
It is a strictly American species limited in distribution, 
apparently, to the northern United States and Canada. 

310. Dioryctria ponderosae Dyar 
FiGtJBEs 384, 868 

Dioryctria ponderosae Dyar, Ins. Insc. Menstr., vol. 2, p. 2, 
1914.— McDunnough, Check list, No. 6130, 1939. 

Maxillary palpus of male squamous. 

Forewing smooth; like that of abietella except: Black- 
ish markings more strongly contrasted; a broad black 
band inwardly bordering the antemedial line, a similar 
band on some specimens of abietella but not so broad 
nor so well contrasted; subterminal line outwardly 
angled at middle; the white transverse lines and discal 
spot well contrasted. Hind wing white, clouded with 
smoky fuscous towards outer margin; the veins slightly 
darkened. Alar expanse, 27-30 mm. 

Male genitalia with uncus considerably shorter and 
broader than that of either abietella or renicvlella; its 
terminal margin broadly rounded; lateral margins in- 
ciu-ved near its base. Harpe with apex of sclerotized 
costa produced into a blunt, curved hook, no spine from 
lower outer angle of the sclerotized costal area ; clasper 
reduced. Female genitalia distinguished by its short 
ductus bursae. 

Type localitt: Lamedeer, Mont, (type in USNM). 

Food plant: Pinus ponderosa. Larva a borer in the 
cambium of the bark. 

Distribution: Montana, Lamedeer (Jvme), Colorado, 
Glenwood Springs (June, July, Aug.); California, 
American River (May) . 

A distinct species known only from a half-dozen 
specimens from the above localities. It belongs defi- 
nitely with the smooth-winged Dioryctria, although a 
few roughened scales can be distinguished in the black 
inner borders of the antemedial line on a couple of the 
specimens. Its genitalia separate it easily from all 
other smooth-winged species. The name has been 
misappKed to a color variety of zimmermani. This 
misidentification is discussed under the latter name. 
The Missoula, Mont., female (Hopkins U. S. No. 11508) 
mentioned by Dyar in his description of ponderosae 
belongs to this variety of zimmermani. 

311. Dioryctria majorella Dyar 
FiQUBES 385, 871 

Dioryctria majorella Dyar, Ins. Insc. Menstr., vol. 7, p. 43, 1919. 
Dioryctria muellerana Dyar, Ins. Insc. Menstr., vol. 7, p. 85, 1919 
(new synonymy). 

Maxillary palpus of male squamous. 



Forewing smooth, similar in color and markings to 
that of sysstratiotes except: Somewhat more suffused; 
the whitish markings fainter and a duller, more sordid 
white; the pale discal spot obscure; the blackish inner 
border of the subterminal line and the blackish band 
preceding the antemedial line broader. Hind wing 
smoky white, the veins faintly darkened. Alar expanse, 
28-33 mm. 

Male genitalia distinguished by greatly increased 
width of the sclerotized costal area of harpe; a short 
spur projects from the outer margin of this sclerotized 
area just below its apex. D. erythropasa has a similar 
harpe; but differs markedly in other details of the geni- 
talia — differently shaped uncus, narrower clasper, and 
different spining of penis. 

Female genitalia differ from those of sysstratiotes only 
in insignificant details. The female genitalia of both 
species differ from those of abietella chiefly in the greater 
broadening and thickening of the membrane of ductus 
bursae near its junction with bursa copulatrix. 

Type localities: Jalapa, Mexico (majorella, 9, in 
USNM) ; Zacualpdn, Mexico {muellerana, &, in USNM). 

Food plant: Pine. 

Ejiown only from Dyar's two types and a pair (cf and 
9) from British Honduras. The Honduran specimens 
are smaller than the Mexican types and in wretched 
condition, but their genitalia are a perfect match in all 
details. In his description of muellerana Dyar sug- 
gested that it might be the male of majorella. This 
synonymy is proven by the Honduran examples. They 
were reared from cones of Pinus caribaea. 

312. Dioryctria disclusa Heinrich 
FiGUBB 872 

Dioryctria disclusa Heinrich, in Farrier and Tauber, Iowa State 
CoU. Journ. Sci, vol. 27, p. 495, 1953. 

MaxiUary palpus of male squamous. 

Forewing smooth; basal area to antemedial line 
orange yellow; area beyond brownish red, more or less 
shaded or suffused with yellowish orange (on some 
specimens the groimd color of the entire wing yellowish 
orange), usually the red shade most conspicuous in the 
area between subterminal line and termen; transverse 
lines narrow, white; a white streak along lower margin 
of cell between the transverse lines; antemedial line 
faint, oblique, nearly straight; subterminal line stronger, 
set well out, rather near terminal margin, outwardly 
angulate between veins 6 and lb; discal mark (when dis- 
tinguishable) a white line along discoceUular vein; some 
very short white dashes on terminal margin; cUia smoky 
white. Hind wing smoky white to pale smoky gray, the 
paler examples showing a very faint ocherous tint; 
veins slightly darkened; cilia whitish. Alar expanse, 
24-29 mm. 

Male genitalia like those of auranticella. Female 
genitalia essentially like those of auranticella. The 
differences shown in the figures for the two species 
are the extremes and represent, at most, individual 
variations. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



153 



Type locality: Tryon, N. C. (type in USNM). 

Food plants: Pinus spp. Larvae feeding in the 
cones. 

Distribution: Massachusetts, Martha's Vineyard 
(June) ; New Jersey, Lakehurst (July) ; District of Colum- 
bia; West Virginia, Roosevelt (June), North Carolina, 
Raleigh (June), Tryon (June); Iowa, Ames (June). 

Examples of this species have been in the National 
Collection as auranticella and it was on the basis of this 
misapplication of Grote's name that Dyar described his 
xanthaenobares. The true auranticella is strictly a 
western species, while the distribution of disclusa, as 
far as I know, is limited to the eastern and central areas 
of the United States. 

The paper by Farrier and Tauber gives aU the infor- 
mation known on the life history, food habits, and 
behavior of the insect. 

313. Dioryctria auranticella (Grote) 
Figures 386, 873 

Nephopteryx auranticella Grote, Ann. Mag. Nat. Hist., ser. 5, vol. 

11, p. 57, 1883; Trana. Kansas Acad. Sci., vol. 8, p. 57, 1883. 
Dioryctria miniatella Ragonot, N. Amer. Phycitidae, p. 4, 1887. — 

Hulst, Ent. Amer., vol. 5, p. 156, 1889. 
Dioryctria auranticella (Grote) Hulst, Phycitidae of N. Amer., p. 

134, 1890.— Ragonot, Monograph, pt. 1, p. 194, 1893. 

McDunnough, Check list. No. 6126, 1939. 
Dioryctria xanthaenobares Dyar, Proc. Ent. Soc. Washington, vol. 

13, p. 81, 1911.— Keen, U. S. Dep. Agr. Misc. Publ. 273, p. 

38, 1938.— McDunnough, Check list, No. 6127, 1939.— 

Heinrich, in Farrier and Tauber, Iowa State Coll. Journ. 

Sci., vol. 27, p. 495, 1953. 

MaxiUary palpus of the male in the form of an 
aigrette. 

Forewing smooth; color and markings similar to those 
of disclusa, but on the average somewhat paler, the 
ground color ranging from yellowish orange to brownish 
red; on the darker suffused, reddish examples the yel- 
lowish color of the basal area is less contrasted than in 
ordinary disclusa. Very pale examples of the two 
species cannot be distinguished superficially. In size 
auranticella averages larger. Alar expanse, 27-33 mm. 

Male genitalia with apex of cucullus of harpe extend- 
ing beyond apex of the sclerotized costa. Female 
genitaha exhibiting only slight and comparative differ- 
ences from those of disclusa. 

Type localities: New Mexico {auranticella, in Univ. 
Kansas); Arizona (miniatella, in Paris Mus.); Kaslo, 
British Columbia {xanthaenobares, in USNM) . 

Food plants: Pinus spp. Larvae feed chiefly in the 
cones, sometimes in the twigs. The favored host seems 
to be Pinus ponderosa. 

Distribution: United States: New Mexico, state 
locality only; Arizona, Mohave County (July); Colo- 
rado, Glenwood Springs (Aug.), Rocky Mountain Na- 
tional Park (July); Utah, Eureka (July, Aug.); Idaho, 
Coeur d'Alene (July); Montana, Bitterroot River 
(July), Camas (July); California, Gasquets (May), 
Pasadena, Warner Mts. (July); Oregon, Monumental 
Pass (Aug.), Silver Lake (Aug.); Washington, Pullman 
(July), Rock Lake (June, July), Seattle; Nebraska, 

30032ft— 56 11 



Halsey (Aug.), Valentine (July, Aug.). Canada: 
British Columbia; Kaslo (Aug.), Trout Creek (Ibapah 
Mts.). The Nebraska records, our most easterly, are 
from examples reared from Pinus ponderosa in refor- 
ested areas. The insect was probably introduced there 
on western nursery stock. 

The species is represented by a large series of speci- 
mens in the National Collection, a majority of them 
reared from cones in connection with the forest insect 
investigations of the U. S. Bureau of Entomology and 
Plant Quarantine. The type of auranticella in the Snow 
Collection at the University of Kansas is a pale femalr 
in rather poor condition. Three other similar females, 
in better condition are in the Rutger's College Collec- 
tion. All of these are labeled simply "New Mexico, 
Snow, Coll." Females of auranticella are not separable 
from those of disclusa, especially the pale, yellowish 
examples, except by their locality labels. On average 
specimens the white streak along the lower margin of 
the cell of forewing is shorter in auranticella, not reach- 
ing beyond the outer angle of the cell; but this feature is 
not constant in either species. Between males there 
is never any need for confusion; for auranticella is the 
only known American species with aigrettelike maxillary 
palpi. 

A similar maxillary palpus occurs in Dioryctria laurata 
(Heinrich) from Japan (described as a Salebria in Proc. 
Ent. Soc. Washington, vol. 30, p. 61, 1928). The unique 
male type has the labial palpi erect and appressed 
close to the face and a strong scale tuft in the sinus of 
the antennal shaft. Hence the original reference to 
Salebria. It is probably a snyonym of Dioryctria 
pryeri Ragonot (Monograph, pt. 1, p. 194, 1923), 
described from a single female from the Holland Col- 
lection, now in the Carnegie Museum, Pittsburgh. I 
have never seen this specimen. 

314. Dioryctria erythropasa (Dyar) 
FiGUHEs 387, 869 

Pinipestis erythropasa Dyar, Ins. Insc. Menstr., vol. 2, p. 112, 

1914. 
Dioryctria erythropasa (Dyar) Barnes and McDunnough, Check 

list of the Lepidoptera of Boreal America, No. 5564, 1917. — 

McDunnough, Check list. No. 6128, 1939. 

Maxillary palpus of male squamous. 

Forewing with some roughened (raised) scale tufts in 
median area (one in lower fold and another in cell just 
beyond antemedial line and a slight roughening of the 
white scales of the discal spot), otherwise smooth; 
ground color red-brown of a somewhat darker, more 
rosy shade and lacking any of the orange suffusion com- 
mon to auranticella and disclusa; more or less dusted 
with white in median area, the white concentrated into 
a broad patch extending from inner margin to top of cell 
just beyond antemedial hne; the transverse lines thin, 
white; antemedial line obHque, irregularly and very 
weakly dentate; sub terminal line nearly vertical, slightly 
denticulate, bordered inwardly by a dark red-brown 
line; a similar dark line forms an outer border to the 
antemedial line; discal spot a sUghtly enlarged, lunate, 



154 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



white line on discocellular vein; outer area beyond sub- 
terminal line red-brown, terminal dots confluent, form- 
ing a more or less continuous blackish line along termen; 
cilia reddish brown. Hind wing smoky white; the veins 
darkened; a very narrow dark shade along termen; ciha 
whitish, cut by a dark subbasal liae. Alar expanse, 
23-28 mm. 

Male genitalia of the majorella type but with uncus 
short, broad, its terminal margia angulate. Harpe with 
slender, digitate clasper. Viuculum narrower, more 
gradually tapered. Penis with numerous anterior 
spines, but without the usual enlarged posterior cornu- 
tus. Female genitaha with bm-sa copulatrbc greatly 
reduced, much shorter than ductus bursae. 

Type locality: Chiricahua National Forest, Ariz. 
(typeinUSNM). 

Food plant: Pinus chihuahvxma. Larvae feeding 
in the cones. 

Disteibution: Arizona, Chiricahua National For- 
est (May); Redington (Aug.). 

This species, with pygmaeella, forms a connecting link 
between the smooth-winged Dioryctria species and those 
with distinctly roughened scales formerly referred to 
Pinipestis. 

315. Dioryctria horneana (Dyar) 

FiGUEB 874 

Pinipestis horneana Dyar, Ins. Insc. Menstr., vol. 7, p. 43, 1919. 

MaxUlary palpus of male squamous. 

Forewing bright red-brown shaded with white, the 
ground color brighter and more on the red shade than 
that of any of the foregoing species; a rather broad, 
diffused, white shade preceding the antemedial line 
and two rather large, confluent patches following it, one 
in the cell and another in lower fold; a strong broad 
oblique white shade extending from inner margin near 
subterminal line to costal beginning of that line and 
fusing with and more or less obscuring it; the scales of 
these white areas as well as those of the white discal 
spot decidedly roughened; a narrow band of appressed 
white scales along terminal margin; antemedial line 
narrow, white, nearly vertical, notched above and below 
its middle, followed on basal half by a faint, nari'ow, 
gray outer bordering line; subterminal white line rather 
close to outer margin, vertical with a slight bulge at 
middle; terminal black dots narrow weak, confluent. 
Hind wing yellowish white with a very faint smoky tint; 
veins but shghtly darkened; a fine, pale brown line along 
termen. Top of head and collar of thorax red-brown; 
remainder of thorax whitish. Alar expanse, 25-28 mm. 

Genitaha similar to those of zimmermani. 

Type locality: Herradura, Pinar del Rio, Cuba 
(typeinUSNM). 

Food plant: Pinus sp. 

Ejiown only from the female type and a male from 
the same rearing. Dyar in his original description gives 
Santiago de las Vegas as the type locaUty, but was evi- 
dently in error. Dr. S. C. Bruner, Chief of the De- 
partmenta Agron6mica of Cuba, has given us the cor- 



rect locahty. The species is easily distinguished from 
others of the zimmermani complex by the bright reddish 
ground color of its forewings and from the other red- 
winged American species by its strongly tufted fore- 
wings. 

The labial palpi of the male are more closely appressed 
to the face than those of most Dioryctria species. 

316. Dioryctria pygmaeella Ragonot 
FiGUEEs 388, 877 

Dioryctria pygmaeella Ragonot, N. Amer. Phycitidae, p. 5, 1887; 

Monograph, pt. 1, p. 192, 1893. 
Pinipestis pygmaeella (Ragonot) Hulst, Phycitidae of N. Amer., 

p. 136, 1890.— Forbes, Cornell Mem. 68, p. 620, 1923.— 

McDunnough, Check list, No. 6136, 1939. 

Maxillary palpus of male squamous. 

Forewing smooth except for a slight roughening of 
the white scales of the transverse lines and discal spot; 
purplish gray with irregular patches of dull dark red 
and a more or less extended white dusting; the red 
patches consisting of a rather broad band preceding the 
antemedial line, a similar band from middle of iimer 
margin to cell and a rather broad shade outwardly 
bordering the subterminal line; the white dusting con- 
centrated over the median areas not occupied by the 
red median band, in subbasal area just behind the red 
band, and along termen following the red subterminal 
shade; transverse lines thin, whitish, rather faint (under 
magnification, the scales silvery and somewhat rough- 
ened); antemedial line oblique, notched at vein lb, 
bordered outwardly by a black line which expands at 
costa into a black patch; subterminal line sinuate- 
angulate with a broad, black, inner, bordering line; 
terminal dots fused into a fime black line along termen; 
white discal spot an oblique lunule; a fine black line 
along the lower half of the inner margin of the red band 
preceding antemedial line (but not a raised-scale ridge 
as stated by Ragonot) ; also some dusting of black scales 
on the red of extreme basal area. Hind wing pale smoky 
gray; the veins darkened and a narrow dark shade 
along termen. Alar expanse, 15-21 mm. 

Male genitalia with a cluster of long strong posterior 
spines and one or two weak anterior spines on penis. 
Female genitalia with the spine clusters in bursa con- 
sisting of narrow, straight bands of slender spines; 
ductus bursae sclerotized from junction with bursa 
almost to genital opening. 

Type locality: Florida (type in Paris Mus.). 

Food plant: Taxodium distichum. Larvae in the 
cones. This record from reared series from Maryland 
(Heinrich, 1920) and Virginia (Busck, 1927) in the U. S. 
National Museum. 

Distribution: Florida, Winter Park (July) ; Virginia, 
Cape Henry (Aug.) ; Maryland, Pokomoke (Aug., Sept.). 
Probably over the range of its host plant. 

A distinct, easily recognized species, intermediate be- 
tween the smooth-winged species and those with definite 
raised-scale ridges. 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



155 



317. Dioryctria zimmermani (Grote) 
FiGUKEs 389, 875, 878 

Nephopieryx (Dioryctria) zimmermani Grote, Canadian Ent., 

vol. 9, p. 163, 1877. 
Nephopieryx (Pinipestis) zimmermani (Grote), Canadian Ent., 

vol. 10, p. 19, 1878.— Packard, U. S. Dep. Agr. Fifth Rep. 

Ent. Comm., p. 73, 1890. 
Pinipestis zimmermani (Grote), Bull. U. S. Geol. Geogr. Surv. 

Terr., vol. 4, p. 699, 1878; op. cit., vol. 6, p. 589, 1882; N. 

Amer. Ent., vol. 1, p. 11, pi. 2, fig. 10, 1879. 
Nephopieryx zimmermani (Grote) Kellicott, Canadian Ent., vol. 

11, p. 114, 1879; Ent. Amer., vol. 1, p. 173, 1885. 
Dioryctria zimmermanni (Grote) Ragonot, Ent. Amer., vol. 6, 

p. 114, 1889; Monograph, pt. 1, p. 190, 1893 (emended 

spelling of specific name). — Forbes, Cornell Mem. 68, p. 620, 

1923. 
Pinipestis zimmermanni (Grote) Hulst, Phycitidae of N. Amer., 

p. 137, 1890.— McDunnough, Check list, No. 6133, 1939. 
Salebria delectella Hulst, Canadian Ent., vol. 27, p. 57, 1895. — 

Ragonot, Monograph, pt. 2, p. 550, 1901. (New synonymy.) 
Dioryctria delectella (Hulst) Dyar, Proc. Ent. Soc. Washington, 

vol. 6, p. 227, 1904. 
Retinia austriana Cosens, Canadian Ent., vol. 38, p. 362, 1906. — 

Busck, Journ. New York Ent. Soc, vol. 15, p. 236, 1907. 
Pinipestis delectella (Hulst) Barnes and McDunnough, Check 

list of the Lepidoptera of Boreal America, No. 5570, 1916. — 

McDunnough, Check list. No. 6134, 1939. 
Dioryctria ponderosae Heinrich (not Dyar), in Keen, U. S. Dep. 

Agr. Misc. Pub. 273, p. 38, 1938. 
Dioryctria zimmermani (Grote) Craighead, U. S. Dep. Agr. 

Misc. Publ. 657, p. 452, 1950. 

Maxillary palpus of male squamous. 

Forewing with a ridge of raised (roughened) scales 
preceding and one following the antemedial line, some 
rough scaling of the disca] spots and on some specimens 
in the outer median area above inner margin and im- 
mediately before subterminal line, the raised scaling 
somewhat variable and nowhere reaching to costa, 
easily and frequently flattened in the spreading and 
setting of specimens; color variable, blacMsh gray with 
a rather broad, but faint, whitish dusting before the 
subterminal line and, on some specimens but to a lesser 
extent, immediately following the antemedial line and 
narrowly and faintly along the inner margin of the 
subbasal scale ridge; basal and terminal areas normally 
shaded with red, the extent and tint of the shading 
extremely variable and on some specimens almost 
obliterated or confined to dull patches of an oblivace- 
ous hue; when strongly accented, extended over base 
and onto the coUar of the prothorax, usually most dis- 
tinct between subbasal scale ridge and antemedial line; 
the raised-scale ridges themselves, black; transverse 
lines dull white, always distinguishable but sometimes 
faint, bordered inwardly and outwardly by black lines 
which broaden into dark wedges on costa; white discal 
spot usually distinct; a black Une along terminal margin. 
Hind wing white more or less shaded with smoky fuscous 
along costa and termen, less so on male than female; a 
fine dark line along termen; veins only faintly darkened. 
Alar expanse, 25-33 mm. 

Male genitaUa with uncus but slightly longer than 
broad, the lateral margins slightly concave; terminal 
margin rounded; when flattened in preparation, as in 
figure 390a of cambiicola. Harpe with costa broadly 



sclerotized and terminating at apex in a long curved 
pointed hook, a short spine from its lower outer angle; 
clasper digitate; cucullus narrow, pointed at apex. 
Penis with posterior spine, long, strong, straight, evenly 
tapering to a sharp point. Vinculum stout, consider- 
ably longer than broad, evenly tapering to roundly 
angulate terminal margin. 

In the female genitalia the variation in the spining 
of bursa shown in the figures is merely individual and 
is equaled or exceeded in any series of eastern or western 
specimens. Ductus biursae much longer than bursa, 
sclerotized for its entire length except for a short dis- 
tance from genital opening, the sclerotization ribbon- 
like, broadening and bent towards bursa, longitudinally 
ribbed on caudal half and terminating caudally in a 
produced, bluntly pointed or acutely roimded central 
projection. Bursa proportionally small; the spine 
clusters closely grouped at its posterior half; the en- 
larged lobe giving off the ductus seminaUs appreciably 
thickened. 

Type localities: Buffalo, N. Y. (zimmermani, in 
BM; paratype, cf , in USNM); Colorado (delectella, in 
AMNH, ex Kutgers) ; Toronto, Ontario, Canada (aus- 
triana, in Royal Ontario Mus.). 

Food plant: Pinus spp. Most if not all species of 
pine in this country are attacked. The spruce records 
given by Packard (1895) have never been verified. 
They were probably based upon misidentified larvae. 
I doubt very much that zimmermani feeds on anything 
but pine. The larvae bore into the cambium of the 
trunk, branches, and twigs, causing considerable dam- 
age to the new growth of older trees and sometimes even 
killing younger trees (8 inches or less in diameter) by 
completely girdling their boles. The place of attack is 
usually indicated by a resinous mass of exuded pitch 
mixed with frass and larval exuviae. 

Distribution: United States: New York, Buffalo, 
Coram (Long Island, Aug.), Warrensburg (Aug.); Con- 
necticut, Woodstock (Aug.); Rhode Island, Washington 
County (June); Massachusetts, Dover (Aug.), Martha's 
Vineyard (Aug.); New Hampshire, Hampton (Sept.); 
North Carolina, Tryon (Aug.) ; Ohio, Akron (July, Aug.), 
Lake County (July, Aug.), Mentor (June), Scioto 
County (July) ; Illinois, Oregon (July, Aug.) ; Nebraska, 
Ainsworth (Aug.), Halsey (May, June, July, Aug.), 
Meadville (Aug.), Norden (Aug.), Wyoming, Wyoming 
National Forest (June, July) ; Montana, Banner (July) , 
Missoula (July); Colorado, only the state locality (fe- 
male cotypes of delectella, Bruce, collector) ; New Mexico, 
Taos Junction (July); Arizona, Santa Catalina Mts. 
(Bear Canyon, July), White Mts. (Aug.); Calijomia, 
Placerville, San Mateo (June), Ventura; Oregon, Butte 
Falls, Coletin; Washington, Friday Harbor (Aug.), 
Rock Lake (July). Canada: Ontario, Toronto. 

The foregoing records are from specimens before me, 
most of them reared. The range of the species probably 
extends over the entire northern areas of the United 
States wherever its hosts occur and presumably over 
a considerable area in southern Canada. 

Hulst's delectella was described from Colorado females 



156 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



which, except for somewhat larger expanse (32 mm.) 
than average eastern specimens, are typical zimmermani 
in coloration and genitalia. The unfortunate reference 
to ponderosae in the Keen paper (1938) was due to my 
misidentification of reared specimens from the reforesta- 
tion areas of Nebraska. These came to us in good 
series but were spread and the raised scales on the fore- 
wings had been flattened, and on most of the examples 
(as also on some eastern specimens) the characteristic 
red shading on basal area of forewing was lacking. 
Superficially they looked like ponderosae; but had the 
typical zimmermani genitalia, and unrubbed examples 
clearly showed the raised scaling. 

Economically zimmermani is our most important 
Dioryctria. In this country it does more serious dam- 
age, especially to young trees, in both the East and the 
West, than abietella. As far as I know it is strictly a 
bark borer and does not attack the cones. The most 
complete and accurate account of the life history is that 
in the Craighead (1950) paper. The life history of the 
insect in Nebraska is also treated in the Keen (1938) 
paper under "ponderosae." 

318. Dioryctria cambiicola (Dyar) 
Figures 390, 876 

Pinipestis cambiicola Dyar, Ins. Insc. Menstr., vol. 2, p. 2, 
1914.— McDunnough, Check list, No. 6137, 1939. 

Maxillary palpus of male squamous. 

Forewing as in zimmermani except: Ground color of 
basal, submedial and terminal areas a dark, dull, reddish 
brown (somewhat paler on Arizona specimens); the 
black scaling greatly reduced; the white scaling follow- 
ing antemedial and preceding subterminal lines and 
bordering terminal margin; the transverse lines and 
the discal mark, dull silvery. Hind wing smoky fuscous ; 
the veins darkened and a dark line along termen. Alar 
expanse, 28-32 mm. 

Genitalia, male and female, show no specific differ- 
ences from those of zimmermani. The extent of the 
bend in the female ductus bursae shown by the figures 
is purely individual and can be easily accented or 
diminished in preparing the slides. 

Type localitt: Flathead Reservation, Mont, (type 
inUSNM). 

Food plants: Finns ponderosae, P. scopulorum, P. 
covlteri. The larvae are cambium borers in new growth, 
causing pitch exudations like those of zimmermani. 

Distbibution: Montana, Flathead Reservation (July, 
Aug.); Missoula (July); Colorado, Boulder (Sept.), 
Palmer Park (July, Aug.); New Mexico, Las Vegas; 
Arizona, Flagstaff, White Mts. (Aug.) ; California, 
Julian (Sept.). 

The species is doubtfully distinct from zimmermani 
except as a possible race. It differs chiefly in its dark 
hind wings. It apparently has a limited distribution 
in our Western States. In his original description Dyar 
mentions an eastern specimen (presumably from Wash- 
ington, D. C.) reared from a cone of Pinus taeda, Aug. 
14, 1882. I have also before me a similar female from 
Cape Henry, reared June 9, 1927. I suspect that both 



these examples may be hybrids of zimmermani and 
amatella. 

319. Dioryctria amatella (Hulst) 
Figure 879 

Nephopteryx amatella Hulst, Ent. Amer., vol. 3, p. 131, 1887. 
Dioryctria amatella (Hulst), Ragonot, Ent. Amer., vol. 5, p. 114, 

1889; Monograph, pt. 1, p. 191, 1893.— Craighead, U. S. 

Dep. Agr. Misc. Publ. 657, 1950. 
Pinipestis amatella (Hulst), Phycitidae of N. Amer., p. 136, 

1890.— McDunnough, Check list. No. 6135, 1939. 

Maxillary palpus of male squamous. 

Forewing with raised scales as in zimmermani but the 
scale ridges somewhat narrower; ground color a dark 
wood brown; transverse lines, discal spot, a thin trans- 
verse band preceding the subbasal ridge, a blotch fol- 
lowing the antemedial line, a similar shade near inner 
margin of subterminal line on lower half of wing, and 
a thin zigzag pale shade just within terminal margin, 
silvery white and strongly contrasted against the 
ground color; black scaling limited to the subbasal and 
submedian scale ridges, the thin borders of the trans- 
verse lines, a spot at extreme base, and the thin terminal 
line. Hind wing smoky with a somewhat glossy pale 
brownish tint; the veins slightly darkened and a thin 
dark line along terminal margin. Alar expanse, 27- 
32 mm. 

Genitalia essentially like those of zimmermani. 

Type localitt: Florida (type in AMNH, ex Rutgers). 

Food plants: Pinus spp. Larvae feeding in cones 
and terminals, more often in the cones than terminals and 
apparently favoring diseased cones. 

Distribution: Florida, Alton (June), Camp Pinchot 
(June), Fort Mead (May), Lake City (May, June), 
Monticello (Sept.), Orlando (June), "Southern Florida" 
(June, July), Starke (May); Louisiana, New Orleans 
(Sept., Nov.), Woodworth (June) ; Texas, Conroe (May) ; 
Mississippi, Hattiesburg (June), Picayune (May); 
Maryland, Baltimore (Aug.); District oj Columbia, 
Washington (July). 

Close to but apparently distinct from zimmermani 
and cambiicola, distinguished from both by the strong 
contrast of its white markings. Generally distributed 
in the Gulf States where its abundance, especially in 
Florida and Louisiana, makes it something of a pest. A 
brief accoimt of what is known of its fife history is given 
in the Craighead (1950) paper. 

320. Dioryctria albovittella (Hulst) 
Figure 880 

Pinipestis albovittella Hulst, Phycitidae of N. Amer. p. 138, 
1900. — Barnes and MclDunnough, Contributions, vol. 4, p. 
174, 1918.— McDunnough, Check list, No. 6138, 1939. 

Dioryctria albovittella Ragonot, Monograph, pt. 1, p. 193, 1893. 

Maxillary palpus of male squamous. 

Forewing with a narrow black subbasal scale ridge, 
little if any raised scaling otherwise; forewing gray 
densely dusted with white, making the extreme basal, 
median, and terminal areas a pale ash color; transverse 
lines and discal mark well contrasted, white; antemedial 
line nearly vertical, slightly notched above and below 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



157 



middle, bordered outwardly by a thin black line and 
inwardly by a broad dark shade which includes the 
black raised-scale ridge; a white blotch on lower half of 
wing just beyond the antemedial line; subterminal line 
well defined, median section broadly triangulate, bor- 
dered inwardly by a blackish line and outwardly by a 
narrow dark shade; discal marking a white spot covering 
discocellular vein; a row of confluent black dots along 
termen. Hind wing white with a faint ocherous tint; a 
fuscous shade at apex and, narrowly, along termen. 
Alar expanse, 23-30 mm. 

Male genitalia like those of zimmermani. Female 
genitalia similar to those of cambiicola, differing only in 
trifling details of the spining in the bursa, not exhibiting 
any consistent differences of a specific character. 

Type locality: Colorado (tj^ie in AMNH, ex 
Rutgers). 

Food plant : Cones of Pinus monophylla. This rec- 
ord from reared Nevada specimens received from the 
Forest Insect Division of the U. S. Bureau of Entomol- 
ogy and Plant Quarantine under Hopkins U. S. No. 
32009. 

Distribution: New Mexico, Jemez Mts. (Aug.), 
Jemez Springs (Aug.) ; Arizona, Mohave County (July, 
Aug.) ; Colorado; Utah, Dividend (Aug.), Eureka (Sept.), 
Ibapah Mts. (Trout Creek, July) ; Nevada, Topaz Lake 

Another close relative of zimmermani and cambiicola, 
distinguished chiefly by its much paler forewings. In 
his original description Hulst gives Hot Springs, N. 
Mex., as the type locality; but this, as pointed out by 
Barnes and McDunnough (1918) is evidentally a lapsus. 
The type is a male, labeled "Colo. Bruce." 

321. Dioryctria gulosella (Hulst), new combination 
Figure 392 

Acrobasis gulosella Hulst, Phycitidae of N. Amer., p. 126, 1890.— 

Ragonot, Monograph, pt. 1, p. 109, 1893; pt. 2, p. 520, pi. 

50, fig. 11, 1901. 
Pinipestis gulosella (Hulst) Barnes and McDunnough, Check 

list of the Lepidoptera of Boreal America, No. 5575, 1916. — 

McDunnough, Check list, No. 6139, 1939. 

Maxillary palpus of male squamous. 

Forewing with a subbasal scale ridge and small patch 
of raised scales in median area; dark gray with a fine, 
sparse, white dusting, making the general color a dark 
ash gray; transverse lines whitish gray, distinct; ante- 
medial line twice notched, edged outwardly by a thin 
black line, preceded by an obscure pale patch on lower 
half of wing, this followed on its inner margin by a black 
scale ridge continued as a thin black line to costa; sub- 
terminal lino outwardly angled at middle, bordered 
inwardly by a black line and outwardly by a narrow 
dark shade; discal spot white; a narrow black line along 
termen. Hind wing white, smoky at apex and some- 
what along termen (especially on females) ; the veins 
more or less darkened ; a fine brown line along terminal 
margin. Alar expanse, 21-27 mm. 

Genitalia similar to those of baumhojeri except for the 
shape of the uncus. The latter has a more broadlj'^ 



roimded terminal margin on gulosella (compare figs. 
391 and 392). 

Type locality: Hot Springs, N. Mex. (type in 
AMNH, ex Rutgers). 

Food plant: Not definitely known, but undoubtedly 
pine. 

Distribution: New Mexico, Hot Springs (Aug.); 
Colorado, Glenwood Springs (July, Aug.), also one 
female with only the state locality (Bruce, collector) 
and bearing a Hulst "type" label. The type from New 
Mexico is also a female. The Colorado specimens (2 
c? and 4 9) are all in the National Collection. Thej'^ 
are a perfect match for Ragonot's figure. 

322. Dioryctria baiimhoferi, new species 
Figures 391, 881 

Maxillary palpus of male squamous. 

Forewing with raised scaling as on typical zimmer- 
mani; blackish gray with the usual black marldngs indi- 
cated by a more intense darkening of the ground color; 
the transverse lines faint but distinguishable, gray; 
antemedial line bordered inwardly by a broad black 
band including the subbasal scale ridge; a similar 
blackish, transverse shade across the middle of the 
wing; subterminal line outwardly angulate at middle, 
bordered inwardly by a black line and outwardly by a 
rather broad black shade extended into streaks on some 
of the veins; a black line along terminal margin; discal 
spot whitish gray, sometimes very faint. Hind wing 
smoky graj"^; the veins darkened and the smoky shade 
intensified along termen. Alar expanse, 25-28 mm. 

Male genitalia having uncus triangulate with nar- 
row^ly rounded apical margin. Female genitalia of the 
zimmermani type but with ductus bursae shorter in 
proportion to length of bursa and somewhat broader. 

Type locality: Prescott, Ariz, (type in USNM, 
61355). 

Food plant: Pinus ponderosa. Larvae feeding in 
new growth. 

Described from male type and one male and nine 
female paratypes from the type locality, reared under 
Hopkins Nos. 9932C and 18506, June 5, 6, and 11, 
1928, by the late L. G. Baumhofer of the Forest Insect 
Division of the U. S. Bureau of Entomology and Plant 
Quarantine. Baumhofer's extensive rearings and field 
studies, especially in the Nebraska National Forest, 
have contributed what knowledge we have of the 
biology of the Dioryctria species in that area. 

The new species is close to gulosella Hulst, from which 
it is distinguished by the much darker color of its fore 
and hind wings, differently shaped male uncus, and the 
somewhat stouter spining of the male penis. It may 
prove to be a local race of gulosella but is at least as 
distinct from it as cambiicola is from zimmermani. 

323. Dioryctria subtracta, new species 

Figures 393, 882 

Maxillary palpus of male squamous. 

Forewing with a subbasal ridge of raised scales and 



158 



UNITED STATES NATIONAL MUSEUM BULLETIN" 207 



a small tuft in lower fold just beyond antemedial line; 
dark gray finely peppered with white making the ground 
color a dark ash gray; the subbasal scale ridge, outer 
border of antemedial and inner border of subterminal 
lines, and the small raised patch following antemedial 
line, black strongly contrasted against the ground 
color, the outer border of antemedial line somewhat 
fainter than the other black markings; subbasal scale 
ridge narrow, reaching almost to costa; between it and 
antemedial line a broad pale band; antemedial line 
obscure, indicated chiefly by its blackish outer bordering 
Une which is almost vertical and parallel with the scale 
ridge; subterminal line also faint, pale gray, outwardly 
angled at middle, bordered inwardly by a narrow black 
line; discal spot obscure, a narrow grayish white line on 
discocellular vein; a fine, strongly contrasted, black line 
along terminal margin. Hind wing white, on female a 
faint smoky tint towards apex; the outer parts of the 
veins faintly darkened and a narrow brown line along 
terminal margin. Alar expanse, 23-25 mm. 

Male genitalia with uncus rather short in proportion 
to width; its terminal margin broadly rounded. Harpe 
narrow and rather short. Penis armed with two 
groups of subterminal spines and a single, straight, 
slender, rather short posterior spine; one of the anterior 
groups consisting of a line of short spines along lateral 
margin of penis near its apex. 

Type localitt: Fort Wingate, N. Mex, (type in 
USNM, 61356). 

Food plant: Unknown. 

Described from male type and one female paratype 
from the type locality (July), and one female from Glen- 
wood Springs, Colo. (Sept.) which McDmmough had 
identified as gulosella Hulst. They are superficially 
similar to the type of that species except for the stronger 
contrast of the black markings on fore wing; but are 
easily separated by their radically different genitalia. 
The expanded (bulbous) shape of the caudal end of the 
female ductus bursae is found in only one other Ameri- 
can species of Dioryctria {clarioralis) and there in a 
lesser degree. 

324. Dioryctria clarioralis (Walker) 
Figures 394, 883 

Nephopteryx clarioraUa Walker, List, vol. 27, p. 54, 1863. 

Dioryctria clarioralis (Walker) Ragonot, Ent. Amer., vol. 5, p. 
114, 1889; Monograph, pt. 1, p. 193, 1893.— Hulst, Phy- 
citidae of N. Amer., p. 136, 1890.— Forbes, Cornell Mem. 
68, p. 620, 1923.— McDunnough, Check list, No. 6125, 1939. 

Ulophora brunneella Dyar, Proe. Ent. Soc. Washington, vol. 6, 
p. 106, 1904. — Barnes and McDunnough, Contributions, 
vol. 3, p. 195, 1916 (make synonym of clarioralis). 

MaxiUary palpus of male squamous. 

Forewing with a weak subbasal ridge of raised scales 
(the species has always been included in the smooth- 
winged Dioryctria group, but unrubbed and unpressed 
examples always show some traces of a raised subbasal 
ridge) ; ground color grayish brown with black patches 
and more or less white shading in the median and ter- 
minal areas; a broad blackish band preceding the ante- 
medial line, paling towards inner margin, bordered in- 



wardly by the black raised-scale ridge and outwardly by 
the thin, black inner border of the antemedial line; the 
latter thin, obhque, sometimes weakly notched below 
costa and more rarely at lower fold, white without (or 
with only an occasional trace, near inner margin) the 
normal black outer bordering hne; a more or less ex- 
tended black smudge in cell, sometimes extended as 
far as the black inner border of subterminal line and 
usually completely obhterating any trace of a white 
discal spot; some white streaking on lower vein of cell; 
subtermraal hne distinct, sharply indented between 
costa and vein 5, thence vertical and straight to inner 
margin, whitish with a thin black inner border; a fine 
line of confluent black dots along terminal margin. 
Hind wing smoky gray or brownish; the veins more or 
less darkened; a fine dark line along termen. Alar 
expanse, 22-29 mm. 

Male genitalia with uncus triangulate, appreciably 
longer than greatest width, evenly tapering to very 
narrowly rounded apex. Female with ductus biu:sae 
broadened near genital opening but less so than in 
subtracta; bursa much larger and more heavily spined 
than that of subtracta, at least as long as ductus bursae. 

Type localities: "United States" (clarioralis, in 
BM); Tryon, N. C. (brunneella, in USNM). 

Food plant: Pinus palustris. This record from a 
specimen received from L. A. Hetrick reared from larva 
feeding in the cone. 

Distribution: Florida, Dunediu (Mar.), Fort Myers 
(Apr.), Miami; North Carolina, Tryon (May, June); 
Massachusetts, Martha's Vineyard (June, July). 

Another intermediate between the smooth-winged 
and rough-scaled species, easily distinguished by its 
genitaUa and wtog pattern. 

78. Genus Oryctometopia Ragonot 

Oryctometopia Ragonot, Nouv. Gen., p. 11, 1888; Monograph, 
pt. 1, p. 153, 1893. — Janse, Joum. Ent. Soc. South Africa, 
vol. 4, p. 156, 1941. (Type of genus: Oryctometopia fossula- 
tella Ragonot.) 

Tongue well developed. Antenna of male with a 
slight sinus and scale tuft in base of shaft; pubescent. 
Labial palpus obliquely upturned, reaching to or a trifle 
above vertex; third segment about two-thirds the length 
of second, bluntly pointed, more or less deflected for- 
ward. Maxillary palpi of both sexes broadly squamous, 
the scales forming a flat cover over the face. Forewing 
smooth; 11 veins; vein 2 from before but near lower 
outer angle of cell; 3 from the angle, equidistant at base 
from 2 and 4, parallel with 2 from just beyond base for 
its remaining length; 4 and 5 connate or very shortly 
stalked; 6 from below upper angle of cell, straight; 8 
and 9 stalked for about two-thirds the length of 8; 10 
from the cell, approximate to stalk of 8-9 at base; male 
with costal fold containing a row of coarse scales. Hind 
wing with vein 2 from before lower outer angle of cell; 
3 from the angle; 4 and 5 stalked for about half their 
lengths; 7 and 8 closely approxim;ate beyond cell at least 
for half their lengths; all veins long; cell about one- 
fourth the length of wing; discocelliilar vein slightly 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITrNAE 



169 



curved, not extended at lower angle. Eighth abdom- 
inal segment of male with a pair of long, slender, ventro- 
lateral hair tufts. 

Male genitalia with uncus subtriangulate. Apical 
process of gnathos a short, stout, hook. TranstiUa ab- 
sent. Harpe with one or more short stout thornhke 
spines projecting from lower margin of sacculus. Aedea- 
gus slender, rather long, sinuate. Penis armed with a 
single thornlike cornutus. Vinculum stout, longer 
than broad, subtriangulate, narrowed from middle to 
terminal margin. 

Female genitalia with signum consisting of a single 
round curved plate, densely armed with long, stiff 
spines and covering the ventral and lateral caudal half 
of the bm-sa copulatrix; bursa otherwise membranous; 
a narrow sclerotized coUar about ductus bursae at its 
junction with bursa, the ductus biu^ae otherwise un- 
sclerotized; genital opening simple; ductus seminalis 
from bursa near junction of bursa and ductus bursae. 

The genus, so far as we know, is confined to the New 
World and contains but one tropical American species. 

325. Oryctometopia fossulatella Ragonot 
Figures 53, 395, 900 

Oryctometopia fossulatella Ra,gonot, Nouv. Gen. p. 11, 1888; Mon- 
ograph, pt. 1, p. 153, 1893. 

Phycita moeschleri Ragonot, Nouv. Gen., p. 12, 1888; Monograph, 
pt. 1, p. 182, 1893. — Moschler, Die Lepidopteren-Fauna 
von Portorico, p. 328, 1890. (New synonymy.) 

Forewing gray to pale brownish gray; the transverse 
lines obscure but usually distinguishable; antemedial 
line nearly vertical, far out from base, on well marked 
specimens irregularly serrate and bordered outwardly 
by a thin dark line which is more or less broken, pre- 
ceded by an obscm-e reddish olivaceous (or reddish 
brown) patch on inner margin, the latter bordered 
inwardly by same blackish scaling; a similar obscure 
reddish brown patch over lower fold just before the sub- 
terminal line ; subterminal line more distinct than ante- 
medial, sinuate (outwardly bulged at middle), dull 
whitish gray, bordered inwardly and outwardly by 
narrow dark lines; discal dots, when distinguishable, 
small, separated, blackish; a row of obscure blackish 
dots along termen; on most specimens a dull whitish 
patch over middle of inner margin. Hind wing trans- 
lucent white; the veins not appreciably darkened; a con- 
trasting dark shade along costa and narrowly along ter- 
men; on males, a fine dark subbasal line through the 
white cilia. Alar expanse, 15-20 mm. 

Genitalia as given for the genus. The thornlike 
spines on sacculus of the male harpe are individually 
variable, consisting of one stout, hooked spine and two 
or more slenderer spines, their number and size varying 
not only in individuals but on opposite harpes of the 
same specimen. Cornutus of penis with a flattened 
platelike base. 

Type localities: "Irazu" [Moimt Irazu], Costa 
Rica {Jossvlatella, in BM); Puerto Rico {moeschleri, in 
Paris Mus.). 



Food plant: Bauhinia mexicana. Larvae in the 
pods. This record from Brownsville, Tex., specimens 
reared by the Division of Foreign Plant Quarantine of 
the U. S. Bureau of Entomology and Plant Quarantine. 

Distribution: United States: Texas, Brownsville 
(June). M:fexico: Chiapas (May), Jalapa, Oaxaca, Te- 
huacdn (Apr.). Guatemala: Cayuga (May, June, 
Sept.), Chejel (June, July, Aug.), PurulhA (July), Vol- 
cdn Santa Maria (May, June, July). Costa Rica: 
Esperanza (May), Mount Irazil. PanamA: Cabima 
(May), Corazal (Apr.), La Chorrera (May), Porto 
BeUo (Oct.). Venezuela: Aragus (Rancho Grande, 
May). Brazil: Rio de Janeiro ("10-1-31")- Puerto 
Rico: Coamo Springs (Apr.), Covado (May), Puerto 
Real (Vieques Isl., Apr. ), San German (Apr.). Virgin 
Islands: Kingshill (St. Croix, June). 

The species is easUy distinguished by its genitalia; 
but in color and maculation is variable, as the foregoing 
description indicates. Such variability is purely indi- 
vidual and has no racial or local significance whatsoever. 
Ragonot described fossulatella from a large (20 mm.) 
male and his moeschleri from a small (17 mm.) female 
color variant, which accoimts for his two names and 
their placement in different genera. 

Genera 79-81: Sarata to Lipographis 

[Venational division B. Forewing smooth; veins 4 and 5 sepa- 
rated at base. Hind wing with veins 4 and 5 stallced; ceU usually 
short, about one-third the length of wing (longer in Lipographis, 
about one-half). Labial palpus porrect, broadly scaled, beak- 
like. Male genitalia with transtilla incomplete or absent; harpe 
with costa partially sclerotized, not produced, clasper reduced or 
absent, otherwise simple. Female genitalia with bursa smooth 
or scobinate, without signum; ductus seminalis from bursa.] 

79. Genus Sarata Ragonot 

Sarata Ragonot, N. Amer. Phycitidae, p. 11, 1887; Monograph, 
pt. 1, p. 614, 1893.— Hulst, Phycitidae of N. Amer. p. 168, 
1890. (Type of genus: Sarata dnopherella Ragonot.) 

Tongue well developed. Antenna pubescent; shaft 
of male cylindrical, slightly swollen at base and smooth- 
ly scaled or with a very slight ridge of roughened scales 
along a few of the basal segments. Labial palpus 
porrect, beaklike; second segment obhque, laterally 
flattened, broadly scaled; third segment deflected for- 
ward, about the length of second (sometimes a trifle 
shorter or longer), bluntly acuminate. Ma.xillary pal- 
pus minute, filiform. Forewing smooth; 11 veins; 
vein 2 from well before lower outer angle of cell ; 3 from 
the angle; 4 and 5 separated at base ; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for from one-half 
to two-thirds their lengths; 10 from the cell, approxi- 
mate to the stalk of 8-9 for a considerable distance from 
base; male without costal fold. Hind wing with vein 2 
from well before lower outer angle of cell; 3 from the 
angle or separated from it by a very short spur; 4 and 
5 stalked for two-thirds their lengths; 7 and 8 closely 
approximate beyond cell for nearly half their lengths; 
ceU about one-third the length of wing; discocellular 



160 

vein curved, outwardly produced at lower angle of cell. 
Eighth abdominal segment of male with a pair of weak, 
ventrolateral hair tufts. 

Male genitaUa with uncus triangulate, its apex 
rounded. Apical process of gnathos terminating in an 
elongate hook, moderately long and bladelike (except in 
incanella where it is short, and digitate from an enlarged 
base) ; lateral arms of gnathos broad and stout. Trans- 
tilla incomplete; its lateral elements well sclerotized and 
short or moderately long and slender, their apices 
pointed. Harpe, elongate, slender; apex of cucuUus 
rounded; clasper present as a very short, blunt, wartlike 
projection from near middle of basal margin of cucuUus 
(except in incanella where it is broader and more scoop- 
like), the size and shape of the wartlike clasper indi- 
vidually variable; costa not produced, strongly sclero- 
tized only on basal half. Anellus a shallow, broadly 
U-shaped shield. Aedeagus long, straight, not appre- 
ciably tapering or expanded towards apex, stout to 
moderately slender (incanella) ; penis (except in incan- 
ella) armed with a single, long, stout, spikelike cornutus, 
rarely a second elongate slenderer spine (on penis of 
incanella the single cornutus is a short, stout thorn sit- 
uated near apex). Vinculum stout, as long as or but a 
trifle longer than broad; terminal margin broad. 

Female genitalia with bursa strongly scobinate over 
much of inner surface and more or less thickened 
(cartilaginous) at or near anterior end; ductus bursae 
simple (unsclerotized and imspined throughout), nor- 
mally distinctly shorter than bursa; ductus seminalis 
from bursa near its middle well forward of junction of 
bursa and ductus bursae). 

The species here referred to Sarata form a homogene- 
ous group. Some of them on the basis of a smooth 
male antennal shaft have hitherto been listed imder the 
Old World genus Megasis; but none agrees with the 
type of the latter (rippertella (Zeller), fig. 428) on 
genitalic characters or the stalking of veins 4-5 of hind 
wing, which is always shorter (and frequently incom- 
plete, a mere approximation or anastomosis of the basal 
half of the veins in rippertella) . The difference between 
a smooth antennal shaft and one with some roughened 
scales towards base is very slight, and should have no 
weight against the uniformity of the genitalic and vena- 
tional characters of Sarata otherwise. Indeed the 
species of Sarata are much closer to those of Lipographis 
than to the type of Megasis. 

None of om- species has been reared and nothing is 
known of the food plants of any of them, so that any 
association of females with males is purely speculative. 
Such associations as have been made are open to grave 
suspicion. The females differ from the males not only 
in size but also in pattern and color, and within any 
given species of females the color varies more between 
individuals than it does between the species themselves. 
For this reason I have treated the males and females 
separately, giving to the latter new temporary names 
which can go into synonymy when the sexes are properly 
associated. 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



The following three species hitherto listed in Megasis 
or Sarata must be referred elsewhere: 

Sarata rhoieUa Dyar becomes the type of a new genus 
(Philodema) . 

Sarata umbrella Dyar goes to Lipographis. 

Megasis indianella Dyar is an anerastiine and a 
synonym (see p. 315) of Bagonotia oliveUa (Hulst). 

Genus Sarata (males), Species 326-330: 
S. edwardsialis to S. incanella 

[Antennal shaft at base smooth scaled.] 

326. Sarata edwardsialis (Hulst), new combination 

Figure 396 

Megaphysis edwardsialis Hulst, Trans. Amer. Ent. Soc, vol. 13, 

p. 163, 1886. 
Megasis polyphemella Ragonot, N. Amer. Phycitidae, p. 10, 1887; 

Monograph, pt. 1, p. 545, 1893. 
Megasis edwardsialis (Hulst), Ent. Amer., vol. 5, p. 156, 1889; 

Phycitidae of N. Amer., p. 166, 1890. — McDunnough, Check 

list, No. 6259, 1939. 

Forewing pale grayish brown more or less smudged 
with darker gray; blackish streaks on several of the 
veins, especially marked on vein lb, the lower vein of 
cell, and the veins immediately preceding and following 
the sub terminal line; the latter faintly indicated; ante- 
medial line obsolete; discal spots poorly defined, often 
obliterated, where distinguishable, separated; a row of 
narrow black dots along termen, more or less accented. 
Hind wing a little lighter grayish brown than ground 
color of forewing, the veins not appreciably darkened, 
a slightly darker line along termen; cilia paler, a dull 
white with a faint fuscous tint. Alar expanse, 35-46 
mm. 

Male genitalia with uncus evenly tapering to narrowly 
rounded apex. Apical process of gnathos about half as 
long as uncus. Elements of transtilla very short. 
Aedeagus long, stout; penis armed with a single, straight 
cornutus, nearly as long as aedeagus, also a small sup- 
plemental sclerotized patch. Terminal margin of vin- 
culum concave. 

Type localities: Nevada (edwardsialis, in AMNH, 
ex Rutgers); California (polyphemella, in Paris Mus.). 

Food plant: Unknown. 

Distribution: Colorado, Boulder (Mar.), Glenwood 
Springs (Mar., Apr.), Salida; Utah, Dividend (Mar.), 
Eureka (Apr., May) ; Nevada; California, "Middle Cali- 
fornia"; Washington, Grand Coulee (Apr.), Pullman 
(Apr.). 

Average specimens (40 mm. or more) make this the 
largest species in the genus. The Ragonot figure of 
polyphemella (Monograph, pi. 19, fig. 8a) is a very good 
likeness of normal examples, except that hind wing is a 
trifle too dark. 

327. Sarata pullatella (Ragonot), new combination 

Figure 397 

Megasis pullatella Ragonot, N. Amer. Phycitidae, p. 10, 1887; 
Monograph, pt. 1, p. 547, 1893. 

Smaller on the average than edwardsialis and darker. 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



161 



Forewing almost uniformly "suffused dark grayish fus- 
cous (blackish gray on some examples) ; on some speci- 
mens the basal and terminal areas paler by contrast 
and the antemedial line indicated, but very faint, nearly 
vertical ; on occasional specimens a paler brownish shade 
in the cell; faint blackish streakings on the veins in 
terminal ai'ea (especially on specimens with pale outer 
area) and more or less of a black streak on lower vein 
of cell; subterminal line very faint, often completely 
obscured, when distinguishable indicated chiefly by a 
broken black shading forming its inner border; discal 
dots obscured. Hind wing pale to dark gray-brown; 
the cilia whitish. Alar expanse, 29-36 mm. 

Male genitalia figured from type and a typical speci- 
men from Dividend, Utah, to show extent of individual 
variation. Uncus not so evenly tapering as in edward- 
sialis; its lateral margins slightly angled at middle. 
Aedeagus less stout and the single long cornutus on penis 
more slender than those of edwardsialis; no supplemental 
patch on penis. 

Type locality: California (type in Paris Mus.). 

Food plant: Unknown. 

Distribution: Utah, Dividend (Apr.) ; California, 
San Diego (Jan.); Idaho, Malta; Washington, Kamiack 
Butte (Feb.), Pullman (Apr.). 

Easily confused with smaller specimens of edwardsialis, 
and all specimens in the National Collection had been so 
identified; but otherwise distinguished by its genitalia. 

Hulst in his Phycitidae of North America (1890) 
made pullatella a synonym of his excantalis and it has 
since appeared as such in our lists. Since the type of 
excantalis is a female, the synonjTny is doubtful, to say 
the least. 

328. Sarata punctella (Dyar), new combination 

Figure 398 

Megasis punctella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 404, 
1915. 

Forewing pale brownish gray with a slight rufous tint 
through the cell; antemedial line indicated by an outer 
border of three black dots, one below costa, a second on 
lower vein of cell, and a third on vein lb; subterminal 
line obscure but usually distinguishable, a whitish spot 
on costa, preceded and followed by blackish dots and, 
below, bordered inwardly by short blackish streaklet on 
veins; on typical specimens some faint, blackish streak- 
lets on the veins of outer area (following the subterminal 
line) ; a row of very faint blackish dots along termen ; 
discal dots obsolete or very faint. Some specimens 
show little or no trace of the blackish markings. Hind 
wings pale brownish gray ("mouse gray") ; the cilia but 
slightly paler, a narrow dark line along termen. Alar 
expanse, 25-30 mm. 

Male genitalia with uncus elongate, slightly and 
evenly tapering to rounded apex. Apical process of 
gnathos appreciably shorter than in preceding species, 
slender. Elements of transtilla long, slender. Penis 
armed with two cornuti — one stout, slightly bent or 

300329 — 56 12 



sinuate, about half as long as aedeagus; the other a 
slender, flattened spine, as long as aedeagus. 

Type locality: Tehuacdn, Mexico (type in USNM). 

Food plant: Unknown. 

Distribution: M:fixico: "Las Vigas" [probably Las 
Viagas, Vera Cruz], Tehuacan (Oaxaca, Sept.), Uruapdn 
(Michoacdn, Mar.). Also three specimens labeled sim- 
ply "V.5." One of the latter was before Dyar when he 
described his species. It had been identified by Druce as 
"Zophodia inomatella Rag." 

Dyar's short original description is thoroughly ade- 
quate and accurate for the type but takes no account 
of suffused examples which lack the characteristic 
blackish markings. 

329. Sarata punctella septentrionaria, new race 

Like typical punctella but larger; the Arizona, New 
Mexico, and Colorado examples a suffused pale gray- 
brown with the usual dark markings obsolete or nearly 
so ; the Wyoming example more gi-ayish with most of the 
veins faintly streaked with blackish scaling. The 
genitalia agree with those of the type of punctella. Alar 
expanse, 32-35 mm. 

Type locality: Palmerlee, Ariz, (type in USNM, 
61357). 

Food plant: Unknown. 

Described from male type from the type locality, 
"Apr. 1-7," and one male paratype from each of the 
following localities: Fort Wingate, N. Mex. (Apr. 14, 
1908); Golden, Colo., Dyar and Caudell, No. 16259; 
Medicine Bow, Wyo. July 4, 1936, I. H. Blake, No. 
"321-11." 

Possibly a distinct species but probably only a larger 
northern variety of punctella. When collections are 
made in the poorly explored areas of northern Mexico 
the species should show a continuous distribution. 

330. Sarata incanella (Hulst), new combination 

Figure 399 

Epischnia incanella Hulst, Canadian Ent., vol. 27, p. 56, 1895.^ 

McDunnough, Check list, No. 6257, 1939. 
Megasis aridella Dyar, Proc. Ent. Soc, Washington, vol. 7, p. 

35, 1905.— McDunnough, Check list. No. 6263, 1939. (New 

synonymy.) 

Forewing gray heavily dusted with white especially 
in outer area and along costa at base and middle; the 
median area between the transverse lines darker than 
remainder of wing; dark shading also on basal area below 
costa; black strealdng on the veins, especially pro- 
nounced in outer area, on vein lb and upper and lower 
veins of cell; transverse lines white, distinctly outlined ; 
antemedial line zigzag, nearly vertical, with some black- 
ish shading along outer margin; subterminal line sharply 
indented at veins 6 and lb; discal dots separated, black- 
ish; terminal dots faint but distinguishable, separated. 
Hind wing whitish with a faint smoky tint ; a very faint 
dark line along termen; cilia concolorous with wing. 
Alar expanse, 31-36 mm. 

Male genitalia with uncus as long as greatest width; 
its apical margin broadly rounded. Apical process of 



162 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



gnathos a short, slender hook arising from a thickened 
base. Clasper of harpe broad, squarish. Aedeagus 
rather short; penis armed with a single small thornlike 
comutus. Vinculmn with terminal margin broadly and 
evenly romided; in aU other species of the genus the 
terminal margin distinctly concave. 

Type localities: Colorado (incandla, in USNM); 
Stockton, Utah {aridella, in USNM). 

Food plant: Unknown. 

Distribution: Colorado; Utah, Stockton (May); 
California, Inyo Coimty (May). 

The palest of the Sarata species, except tephrella Rag- 
onot. The genitalia of the t3rpes of incanella and ari- 
della are identical. 

Genus Sarata (males),' Species 331-337: S. atrella 
to *S. tephrella 

[Antennal shaft with a ridge of roughened scales at base.] 

331. Sarata atrella (Hulst), new combination 
Figure 400 

Megasis atrella Hulst, Phycitidae of N. Amer., p. 166, 1890. — 
Ragonot, Monograph, pt. 1, p. 548, 1893. — McDunnough, 
Check list, No. 6263, 1939. 

Vestiture of labial palpi, head, and prothorax a mix- 
ture of scales and long hairs. 

Forewing blackish gray; transverse lines whitish gray, 
dull; the antemedial line obsciffe; sub terminal line 
stronger, always distinguishable, bordered inwardly and 
outwardly by black streaklets on the veins; discal spots 
obsolete, obscured in the dark ground color; blackish 
dots along termen confluent. Hind wing pale to rather 
dark smoky fuscous, a thin dark line along termen. Alar 
expanse, 30-33 mm. 

Male genitalia with uncus elongate, evenly tapering 
to narrowly rounded apex. Apical process of gnathos 
about half as long as uncus, ventrally flattened. Ele- 
ments of transtiUa short. Penis armed with a single 
strong comutus, as long as aedeagus, no supplemental 
sclerotized plate. 

Type locality: "West Cliff, Colo, (type in AMNH, 
ex Rutgers). 

Food plant: Unknown. 

Distribution: Colorado, Custer County, West Cliff, 
and two specimens with only the state locahty. 

The species is easily distinguished by the hairy vesti- 
ture not possessed by any other known species in the 
genus. 

332. Sarata caudellella (Dyar), new combination 
Figure 401 

Megasis caudellella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 110, 1904.— McDunnough, Check list. No. 6261, 1939. 

Similar to the foregoing species (atrella) except : With- 
out hairy vestiture and with some flne white powdering 
on the blackish gray ground color of forewing; the 
transverse lines distinct, obhque and nearly straight; 
antemedial line followed on costa and preceded at inner 



margin by obscure dark blotches; sub terminal hne bor- 
dered inwardly by a continuous irregular blackish 
shade, outwardly by a short, faint, dark streaking of 
the veins; the veins otherwise not appreciably streaked; 
discal dots faint, but usually distinguishable, more or 
less confluent; dots along terminal margui weak, fused 
into a faint blackish line. Hind wing pale brownish 
gray, semilustrous; a dark line along termen. Alar ex- 
panse, 28-32 mm. 

Male genitalia differ from those of atrella only in 
trifling details. 

Type locality: Golden, Colo, (tj^pe in USNM). 

Food plant: Unknown. 

Distribution: United States: Colorado, Golden 
(May); Washington, Grand Coulee (Apr.). Canada: 
Saskatchewan, Oxbow (June); Manitoba, Aweme (Apr.), 
Miniota. 

Close to but distinct from atrella, from which it is 
distinguished chiefly by its smooth-scaled vestiture 
and the strong contrast of the whitish transverse lines 
of forewing, especiaUy the well-marked antemedial line; 

333. Sarata dnopherella Ragonot 
Figure 402 

Sarata dnopherella Ragonot, N. Amer. Phycitidae, p. 11, 1887; 
Monograph, pt. 1, p. 616, 1893. 

This and the two species following (nigrifasciella and 
cinereella) are very close, separable only by minor color 
differences and trifling variations in the genitalia of their 
types. Such variations are more than covered in the 
series of nigrifasciella and cinereella before me. I sus- 
pect that the names represent nothing but color varie- 
ties of one variable species; but am keeping them sepa- 
rate until life-history information and more extensive 
collections are available and more exact definitions of 
species and possible races can be made. 

Ragonot's dnopherella is authentically represented 
only by its type. His description and figure suggest a 
grayish brown form suffused with blackish brown and 
with the transverse lines very weakly contrasted and 
poorly defined against the ground color. Alar expanse, 
32 mm. 

Male genitalia with uncus broadly and blimtly tri- 
angulate, evenly tapering to narrowly rounded apex. 
Elements of transtiUa moderately long, slender (about 
half the length of those of atrella) . Penis armed with a 
single stout comutus, shghtly more than half as long 
as aedeagus and preceded basally by a small, weakly 
sclerotized patch. 

Type locality: California (type in Paris Mus.). 

Food plant: Unknown. 

Hulst (Phycitidae of N. Amer., p. 168, 1890) treated 
dnopherella as a synonym of his perfuscalis and it so 
appears in our fists. As perfuscalis was described from 
a female the synonymising of the two names was 
arbitrary and, under the circumstances, unwarranted. 
Ragonot (Monograph, p. 616, 1893) very rightly ques- 
tioned it. 



AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



163 



334. Sarata nigrifasciella Ragonot 
Figure 403 

Sarata nigrifasciella Ragonot, N. Amer. Phycitidae, p. 11, 1887; 
Monograph, pt. 1, p. 615, 1893.— Hulst, Phycitidae of N. 
Amer., p. 169, 1890.— McDunnough, Check list. No. 6266, 
1939. 

Forewing pale to dark ashy gray, some specimens 
tinted with a faint brownish shade (one specimen before 
me from Mineralking, CaHf., a very pale, sordid 
brownish gray). The transverse lines distinct, but 
indicated chiefly by the blackish outer border of the 
antemedial line and a similar inner border to the sub- 
terminal hne. This character, however, is variable and 
on the Idaho specimen before me is almost obsolete. 
Hind wing very pale brown, with a fine dark terminal 
line. Alar expanse, 29-32 mm. 

Alale genitalia similar to those of dnopherella except 
for the narrower apical process of gnathos (viewed 
ventrally). This character, however, is not reliable; 
for in a typical series of cinereella all intergrades are 
found between the gnathos of typical nigrifasciella and 
that of dnopherella. 

Type locality: America Septentrionalis (type in 
Paris Mus.). 

Food plant: Unknown. 

Distribution: United States: New Mexico, Fort 
Wingate (Mar.); Colorado, Chimney Gulch (June), 
Platte Canyon (June); Idaho, Wallace (Apr.); Cali- 
fornia, Mineralking (July), Yosemite (this specimen 
labeled nigrifasciella in Eagonot's handwriting). Can- 
ada : Manitoba, Aweme (Apr.) . 

This species, if such it be, is distinguished from the 
preceding and following species chiefly by its more 
marked transverse lines in forewing and the narrower 
apical process of its gnathos. Ragonot's description 
of the females (also from "Amer. Sept.") associated 
with his male type can be ignored, for they represent 
two distinct species. 

335. Sarata cinereella Hulst 

Sarata cinereella Hulst, Canadian Ent., vol. 32, p. 172, 1900. — 
McDunnough, Check Hst, No. 6271, 1939. 

Forewing dark ash gray, nearly concolorous; the 
transverse lines obsolete or nearly so; the subterminal 
line faintly indicated on most specimens and, on one or 
two, very faint traces of the antemedial line; as on 
dnopherella and nigrifasciella there is more or less 
blackish dusting on the veins. Hind wing pale brown- 
ish gray. Alar expanse, 29-33 mm. 

Male genitalia of type show a slightly longer cornutus 
than that of nigrifasciella, but other examples inter- 
grade in all characters between the two types. 

Type locality: Salida, Colo, (type in AMNH, ex 
Rutgers) . 

Food plant: Unknown. 

Distribution: Colorado, Denver (Mar.), Glenwood 
Springs, Salida, also three examples with only state 
locality, two of them bearing Hulst "type" labels 



(Bruce, collector), and the other a pseudotype of 
"Anerastia excantalis Hulst." 
Probably only a suffused form of nigrifasciella. 

336. Sarata rubrithoracella (Barnes and McDunnough), new 
combination 

FiGUBB 404 

Megasis rubrithoracella Barnes and McDunnough, Contributions, 
vol. 2, p. 140, 1913.— McDunnough, Check list, No. 6268, 
1939. 

Forewing pale brownish gray rather evenly dusted 
with white intermixed with a sparse peppering of 
blackish scales; the transverse lines distinct, but faint, 
whitish; antemedial line bordered outwardly by a thin, 
broken, blackish shade; subterminal line bordered in- 
wardly by a continuous narrow blackish band; no ap- 
preciable strealdng on the veins; discal spots obsolete; 
terminal dots confluent, forming a faint, dark line. 
Hind wing pale grayish brown; a thin dark Hne along 
termen. Alar expanse, 22-29 mm. 

Male genitalia show little to distinguish them from 
those of the preceding three species except for the very 
short lateral elements of transtilla and a slightly longer 
cornutus (our drawing of the aedeagus is in reversed 
position from that of other species; if drawn as were 
the other aedeagi the apex of cornutus would point to 
the right) . 

Type locality: White Mts., Ariz, (type in USNM). 

Food plant: Unknown. 

Distribution: Arizona, White Mts.; New Mexico, 
Fort Wingate (Apr., July). 

Apparently a distinct species. Differs from the 
others by a distinct rufous-ocherous shading on the 
thorax and, in genitalia, from the species of the dnoph- 
erella-cinereella group by the short transtilla elements 
of its genitalia. The rufous ocherous thorax occurs 
also in two species of females (kappa and phi). 

337. Sarata tephrella Ragonot 

Figure 405 

Sarata tephrella Ragonot, Monograph, pt. 2, p. 616, 1893. — 
McDunnough, Check list. No. 6268, 1939. 

Known to me only from Ragonot's description and 
figure, and the genitalia of its type. Evidently a much 
paler species than any other of those with a rough-scaled 
antennal shaft, and having much the general habitus of 
a Lipographis. Forewing heavily dusted with white; 
some ocherous brown shading in median and outer 
areas; transverse lines whitish, distinguishable but 
poorly defined. Hind wing dark gray. Alar expanse, 
28 mm. 

Male genitalia similar to those of the dnopherella- 
cinereella group except elements of transtiUa larger and 
cornutus more slender and more evenly tapering. 

Type locality: "Washington Territory" (type in 
Paris Mus.). 

Food plant: Unknown. 

Described and so far known only from its male type. 



164 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



Genus Sarata (females), Species 338-346: 
S. alpJia to S. delta 

[The females are of a uniform pattern and similar coloration; the 
ground color gray, sometimes tinted with pale brown and more 
or less dusted with white, variations in color more individual 
than specific; the transverse lines strongly contrasted, white with 
strongly accented blackish or brownish borders on outer margin 
of antemedial and inner margin of subterminal lines; the lines 
oblique and straight or notched, the notching also more individual 
than specific in character. The only reliable specific characters 
are in the genitalia, and for certain identification it is necessary 
to dissect nearly all females.] 

338. Sarata alpha, new species 
Figure 888 

A bright species with the white lines and their black 
borders sharply contrasted. Forewing blackish gray 
with a strong dusting of white (however, in one speci- 
men from the type locality, the median area distinctly 
darker than basal or outer areas); antemedial line 
slightly curved; subterminal line straight or with a very 
slight bend at lower fold; discal dots distinct, more or 
less confluent. Hind wing whitish with a faint brown 
tint; the veins slightly darkened; a broadened blackish 
brown line along termen; cilia white. Alar expanse, 
21-24 mm. 

Female genitalia with bursa large and greatly elon- 
gated; densely and finely spined over most of interior 
surface, the denser spining in longitudinal rows, partial- 
ly divided by lines of the clear membrane; anterior end 
thickened (cartilaginous), the amount of thickening 
individually variable; ductus bursae very short. 

Type locality: Oxbow, Saskatchewan, Canada (type 
in USNM, 61358). 

Food plant: Unknown. 

Described from female type and two female paratypes 
from the type locality, May 14, 1907, Frederick Knab; 
and additional female paratypes as follows: One from 
Aweme, Manitoba, Apr. 12, 1903. N. Griddle; one from 
Kegina, Saskatchewan, June 5, 1907; and two from 
Chimney Gulch, Golden, Colo., July, Oslar. A female 
in the Rutgers Collection (C. H. slide No. 2186) from 
Colorado identified as atrella also goes here. Needless 
to say there is no trace on any of these females of the 
hairy vestiture of the male of atrella. 

339. Sarata beta, new species 

Figure 889 

Similar to alpha except less glossy. Forewing duller; 
less white dusting; transverse lines and their black 
borders less strongly contrasted; discal dots obscured. 
Hind wing dark smoky gray; the veins not darkened. 
Alar expanse, 23-26 mm. 

Female genitalia like those of alpha in shape and 
proportions except that spining covers appreciably less 
of the bursa sinface, leaving half or more than half of 
the latter membranous and unspined. 

Type locality: Colorado (type in USNM, 61359). 

Food plant: Unknown. 

Described from female type with only a state locality 



label; one female paratype from Custer County, Colo.; 
and one female paratype from Chilcotin, British 
Columbia, May 2, 1920, E. K. Buckell No. 137. The 
two Colorado examples were in the Barnes and National 
Museum Collections as females of atrella Hulst. 

340. Sarata gamma, new species 
Figure 890 

This is the female figured in the Ragonot Monograph 
(pt. 1, pi. 23, fig. 2b.) as a paratype of his dnopher- 
ella. Its genitalia, here figured, are similar to those of 
alpha and beta except for slight differences in the spining 
of the bursa, as shown in the figure. 

Type locality: California (type in Paris Mus.). 

Food plant: Unknown. 

341. Sarata iota, new species 
Figure 894 

This name is proposed for the female paratype of 
pullatella Ragonot, described and figured by him 
(Monograph, pt. 1, p. 547, pi. 19, fig. 9b, 1893). 

Forewing blackish gray with very little pale dusting; 
the antemedial line broader and more strongly con- 
trasted than the subterminal. Hind wing dark grayish 
brown. Alar expanse, 24 mm. 

Genitalia (C. H. slide No. 3113) with bursa consider- 
ably smaller and more sparsely spined than that of any 
of the preceding species; ductus bursae about half as 
long as bursa. 

Type locality: California (type in Paris Mus.). 

Food plant: Unknown. 

It is very likely that this is the female of pullatella; 
but at the present time there is no certainty about any 
of the sex associations in the genus. 

342. Sarata perfuscalis (Hulst) 

Figure 893 

Nephopteryx perfuscalis Hulst, Trans. Amer. Ent. Soc, vol. 13, 

p. 161, 1886. 
Anerastia excantalis Hulst, Trans, Amer. Ent. Soc, vol. 13, 

p. 163, 1886 (new synonymy). 
Megasis excantalis (Hulst), Ent. Amer., vol. 5, p. 156, 1889; 

Phycitidae of N. Amer., p. 165, 1890. — McDunnough, 

Check list, No. 6260, 1939. 
Sarata perfuscalis (Hulst), Phycitidae of N. Amer., p. 168, 1890. — 

McDunnough, Check list, No. 6269, 1939. 

Forewing dull, dark gray more or less dusted with 
white ; the terminal and (usually) the median areas the 
paler, the basal area the darker; transverse lines dis- 
tinct, sordid white, their dark borders well contrasted, 
especially on specimens with considerable white dusting. 
Hind wing smoky grayish brown. Alar expanse, 25- 
29 mm. 

Female genitalia distinguished by the spining of the 
large bursa. These spines are arranged in an elongate, 
ribbed band which extends most of the length of the 
inner dorsolateral surface, curving onto ventral surface 
at anterior end; the area of bursa under the spines more 
or less sclerotized. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



165 



Type locality: California {jperfuscalis, excantalis, in 
AMNH, ex Rutgers). 

Food plant: Unknown. 

Distribution: California (state locality only); 
Washington, Seattle; Utah, Eureka (Mar., Apr.), 
Stockton (Apr.). 

Hulst associated his perfuscalis with dnopherella 
Ragonot; and excantalis with pullatella Ragonot. So 
much for superficial sex associations. Their genitalia 
show the two females to be obviously conspecific. 

343. Sarata epsilon, new species 
Figure 892 

The smallest of the female species. Similar in colora- 
tion to perfuscalis except that the dark borders of the 
transverse whitish lines are somewhat broader and more 
strongly contrasted. Alar expanse, 19-21 mm. 

Female genitalia with bursa moderately large, evenly 
and finely spined on anterolateral half and extreme 
anterior end. 

Type locality: Yosemite, Calif, (type in USNM, 
61360). 

Food plant: Unknown. 

Described from female type from the type locality 
identified by Ragonot as a female of nigrifasciella and 
bearing that name label in his handwriting and female 
paratypes as follows: One from Colorado, Cockerell, 
collector, identified by Hulst as atrella; two from Golden, 
Colo., May, Dyar and Caudell Nos. 16252 and 16253, 
and identified by Dyar as paratypes of caudellella; 
seven from Chimney Gulch, Golden, Colo., June, 
Oslar; and one from Fort Wingate, N. Mex., March. 
Here also is referrable one of the two female paratypes of 
nigrifasciella Ragonot in the Paris Museum (C. H, slide 
No. 2891). 

Most of the foregoing examples show at least one of 
the black discal spots. On each of the seven specimens 
from Chimney Gulch, Colo., is a minute white spot on 
discocellular vein between the black dots and on these 
also there is some very dark brown shading on the 
otherwise blackish borders of the white transverse lines. 



(the female paratype of ruhithoracella Barnes and 
McDunnough); and five female paratypes from Fort 
Wingate, N. Mex. March, Jime, July. Also before 
me, but not included among the paratypes, is a large 
female (26.5 mm.) from Denver, Colo., Apr. 1, 1904, 
Oslar. On this specimen the patagia are more putty 
white than rufus. Its genitalia, however, agree in 
detail with those of typical phi from Arizona and New 
Mexico. The species is uncomfortably close to epsilon. 
It probably does represent the female of ruhithoracella 
but the verification of that relation will have to wait 
upon rearing evidence. 

345. Sarata kappa, new species 

Figure 887 

Forewing dull, as in beta, but with considerable white 
dusting, rather evenly distributed ; the transverse lines 
more irregular and their black borders more strongly 
contrasted ; subterminal line with slight notches at vein 
6 and lower fold; lower discal dot faint, but distinguish- 
able. The thorax of the type is strongly shaded with 
rufus-ocherous. Alar expanse, 23 mm. 

Female genitalia with bursa copulatrix very small 
(the smallest of any of the Sarata species) ; the greater 
part of its inner dorsal surface covered with a dense mat 
of very fine spines. Ductus bursae as long as bursa. 

Type locality: Arizona (type in USNM, 61362). 

Food plant: Unknown. 

Described from a pseudotype of perjuscalis Hulst 
from the Fernald Collection, bearing only a state 
locality and numbered "7820." 

In coloration similar to phi except for the blackish 
borders of the transverse lines of forewing. Dis- 
tinguished from that and other species of the genus by 
its genitalia. 

346. Sarata delta, new species 

Figure 886 

This name is proposed for the second of the female 
paratypes of nigrifasciella Ragonot (in Paris Mus., C. 
H. slide No. 3111) whose genitalia are here figured. 



344. Sarata phi, new species 
Figure 891 

Thorax shaded with rufus ocherous. 

Forewing more evenly dusted with white; dark 
borders of the transverse lines distinctly brownish ; dis- 
cal spots obsolete, replaced by a faint, white line or spot 
on the discocellular vein. Alar expanse, 23-27 mm. 

Female genitalia essentially like those of epsilon. The 
figure shows the bursa twisted into a reverse position 
from that of epsilon to show the somewhat greater 
development of the thickened (cartilagenous) lateral 
margin (a variable and probably only an individual 
character). 

Type locality: White Mts., Ariz, (type in USNM, 
61361). 

Food plant: Unknown. 

Described from female type from the type locality 



80. Philodema, new genus 

Type of genus: Sarata rhoiella Dyar. 

Tongue well developed. Antenna pubescent; shaft 
of male slightly flattened and with a shallow sinus near 
base, the latter containing some shghtly roughened 
scales and a few, minute serrations. Labial palpus 
porrect (as in Sarata but shorter). Maxillary palpus 
vestigial. Venation as in Sarata except vems 4 and 5 
of hind wing stalked for half or less than half their 
lengths and cell a short one-third the length of wing. 
Eighth abdominal segment of male with ventrolateral 
hair tufts. 

Male genitalia as in Sarata except: More squat, 
broader in proportion to their length; harpe short in 
proportion to its width; no erect clasper; aneUus 
strongly sclerotized, its central area developed into a 
pair of produced, pointed, bladelike arms, the usual 



166 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



lateral lobes absent; aedeagus slender, its basal end 
broadened and flattened; penis without cornutus or 
other armature except for a few weak scobinations at 
apex; vinculum stout, shorter than its greatest width. 

Female genitalia with bursa small, simple, membran- 
ous; ductus bursae short with a strongly sclerotized, 
curved, wide, centrally notched, dorsal plate behind 
genital opening; ductus seminalis from a smalt lobe of 
bursa near junction of bursa and ductus bursae. 

The genus falls between Sarata and Hypochalcia and 
has several features of each of these genera but can go 
into neither of them on the sum of its characters. It 
appears to be a New World analogue of the Old World 
Hypochalcia, agreeing with the latter on most genitalic 
characters except for its short vinculum and peculiarly 
developed aneUus. The type of Hypochalcia (ahenella 
(Zeller), fig. 54) has an elongate vinculum. It also 
differs from Philodema in having much longer, smoother 
and slenderer labial palpi, rather broad, squamous 
maxillary palpi and smoother more glossy wing vesti- 
ture. 

Philodema differs markedly from Sarata in that there 
is no sexual dimorphism, the males and females being 
alike in color and markings. 

347. Philodema rhoiella (Dyar), new combination 

FiGTTBES 406, 895 

Sarata rhoiella Dyar, Journ. New York Ent. Soc, vol. 12, p. 105, 
1903.— McDunnough, Check list, No. 6267, 1939. 

Forewing pale, sordid, brownish gray; extreme basal 
area dark smoky gray; the transverse lines indicated 
chiefly by their dark borders, the latter dark smoky 
gray; the outer border of the antemedial line more or 
less broken and diffused; subterminal line bordered 
inwardly by an irregular (zigzag) border, somewhat ac- 
cented on the veins, and outwardly by a more obscure 
dark shade; discal dots distinct, separated. Hind wing 
smoky gray; the veins slightly darkened; a narrow dark 
shade along termen. Alar expanse, 23-33 mm. 

Genitalia as given for the genus; figured from para- 
types from the type locality. The male holotype was 
without an abdomen. 

Type locality: Platte Canyon, Colo, (type in 
USNM). 

Food plant: Rhus toxicodendron. 

Distkibution: Colorado, Platte Canyon (July) ; Utah, 
"So. Utah" (July). 

The Utah specimens (2 cf) are larger (32-33 mm.) 
than any of the Colorado examples; but have identical 
genitalia and wing maculation; nothing is known of the 
life history except Dyar's statement that two specimens 
of the type series were reared from larvae on poison-ivy. 

81. Genus Lipographis Ragonot 

lApographis Ragonot, N. Amer. Phycitidae, p. 10, 1887; Mono- 
graph, pt. 1, p. 562, 1893. — Hulst, Phycitidae of N. Amer., 
p. 166, 1890. (Type of genus: Pempelia fenestrella Packard.) 

Tongue well developed. Antenna weakly pubescent; 
on male, shaft flattened, serrate, and with sinus and 



strong scale tuft at base (except in umbrella and subos- 
seella where the shallow sinus has a weak tuft of rough- 
ened scales). Labial palpus porrect, second segment 
oblique, laterally flattened, broadly scaled; third seg- 
ment deflected forward, decidedly shorter than second, 
its proportions obscm-ed by its long scaling and the 
extended scaling of second segment. Maxillary palpus 
subsquamous (small but broadly scaled, vestigial in 
umbrella). Forewing smooth; venation as ia Sarata 
except for a somewhat shorter stalking of veins 8 and 9 
of forewing and a longer cell ia hind wing (nearly one- 
half the length of the wing); 4 and 5 are also shorter 
stalked, about one-half their lengths.' Eighth abdomi- 
nal segment of male with a pair of ventrolateral hair or 
scale tufts (absent in subosseella) . 

Male genitalia with apical projection of gnathos a 
short stout hook (except in subosseella). Transtilla 
absent except in truncatella and subosseella where it is 
represented by its short, weak, divided elements. Anel- 
lus with short, weak, lateral lobes (except in subosseella). 
Aedeagus broadly expanded towards apex (except in 
truncatella); penis armed with one or more strongly 
sclerotized, ciu-ved, spinelike cornuti (the latter always 
decidedly less than half as long as the aedeagus) . Geni- 
talia otherwise as in Sarata. 

Female genitalia with bursa membranous and greatly 
reduced, if sometimes elongate (truncatella) narrow; 
ductus bursae scobinate and partially sclerotized near 
its junction with bursa copulatrix, greatly broadened in 
proportion to width of bursa (except in truncatella); 
genital opening simple, unsclerotized; ductus seminalis 
from bursa near junction of bursa and ductus bursae. 

Lipographis agrees with Philodema and differs from 
Sarata in that the males and females are alike in color 
and markings. It is distinguished from both Sarata 
and Philodema chiefly by its female genitaUa. The 
latter resemble those of the type of the Old World 
Divona Ragonot (ilignella (Zeller)) except that the 
bursa of ilignella is strongly scobinate, partially sclero- 
tized, and proportionally much larger. 

In his original description of Lipographis, Ragonot 
designated fenestrella as type of the genus. Later 
(Monograph, 1893) he cites humUis as its type. This 
substituted designation is invalid, regardless of the fact 
that humilis was an originally included species and may 
have served as the basis for the original generic descrip- 
tion. That humilis now proves to be a synonym of 
fenestrella is also immaterial and irrelevant. 

One species (subosseella) originally described in lApo- 
graphis is here provisionally retained in the genus. It 
may eventually have to have a new generic placement 
as its only representation (the male type) is aberrant 
in several genitalic details. 

' The venation of fenestrella and leoninella exhibit considerable 
individual variation; veins 4 and 5 of forewing are normally ap- 
proximate for a short distance from cell but sometimes divergent 
and (rarely) even shortly stalked. In one freak specimen before 
me vein 4 is also absent from hind wing, another example which 
advises caution against relying too much upon one structure for 
the identification of phycitids. 



AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 



167 



348. Lipographis fenestrella (Packard) 

Figures 31, 407, 896 

Pempelia fenestrella Packard, Ann. New York Lye. Nat. Hiat., 
vol. 10, p. 259, 1873. 

Nephopteryx fenestrella (Packard) Grote, Bull. U. S. Geol. Geogr. 
Surv. Terr., p. 697, 1878. 

Lipographis humilis Ragonot, N. Amer. Phycitidae, p. 11, 1887: 
Monograph, pt. 1, p. 563, 1893.— Hulst, Phycitidae of N. 
Amer., p. 167, 1890. (New synonymy.) 

Lipographis fenestrella (Packard) Ragonot, N. Amer. Phycitidae, 
p. 10, 1887; Monograph, pt. 1, p. 564, 1893.— Hulst, Phy- 
citidae of N. Amer. p. 166, 1890. — McDunnough, Check 
list, No. 6272, 1939. 

Forewing ash gray, dusted with white and shaded 
with brownish ocherous; the transverse lines narrow, 
white; antemedial line oblique, nearly straight, bordered 
inwardly by a broad brownish ocherous band marked 
by black dots or streaklets on vein lb and upper and 
lower veins of cell, followed outwardly by two or three 
similar black dots; white dusting along lower vein of 
cell, median part of vein lb and along some of the veins 
preceding the subterminal line; subterminal line parallel 
to termen, very slightly indented at veins 6 and lb, 
bordered outwardly by a broad brownish ocherous band 
(the latter interrupted by blackish streaklets on the 
veins) and from costa by short, faint, narrow, inner and 
outer, blackish bordering lines; along termen a narrow 
dusting of white; terminal dots more or less confluent, 
individually variable, forming sometimes a straight, 
sometimes a scalloped, black line; discal dots separated, 
small, blackish; usually a brownish ocherous shade 
along median area of lower fold. Hind wing dull white 
with a faint, smoky tint towards apex and termen; a fine 
blackish line along terminal margin; the veins not 
appreciably darkened. Alar expanse, 19.5-24 mm. 

Male genitalia with aedeagus decidedly bulged from 
shortly beyond base; penis armed with a comb of 5 
stout, curved spines of a graduating length. Female 
genitalia with bm-sa greatly reduced and but slightly 
longer than ductus bursae; the latter appreciably 
broader than the bursa. 

Type locality: California (Jenestrella, in MCZ; hii- 
tnilis, in Paris Mus.) . 

Food plant: Unknown. 

Distribution: California, "Middle California," Palo 
Alto (May), San Diego (Apr., May, June, July, Aug., 
Oct.), San Francisco. 

The type of Ragonot's humilis is merely a small, 
rather dark male of fenestrella with identical genitalia. 
In any considerable series oi Jenestrella the palpal differ- 
ences cited by Ragonot can be observed. They are 
indeed more apparent than real and more due to pro- 
portionate differences in the sizes of the individual 
specimens and to differing positions of the palps. In 
our latest checklists humilis is listed as a subspecies or 
variety of fenestrella. It is not even that. 

349. Lipographis leoninella (Packard) 

Pempelia leoninella Packard, Ann. New York Lye. Nat. Hist., 

vol. 10, p. 259, 1873. 
Nephopteryx leoninella (Packard) Grote, Bull. U. S. Geol. Geogr. 

Surv. Terr., vol. 4, p. 697, 1878. 



Lipographis leoninella (Packard) Ragonot, Ent. Amer., vol. 5, 

p. 115, 1889; Monograph, pt. 1, p. 565, 1893.— McDunnough 

Check list. No. 6273, 1939. 
Lipographis fenestrella leoninella (Packard) Hulst, Phycitidae of 

N. Amer., p. 167, 1890. 
Pyla pallidella Dyar, Journ. New York Ent. Soc, vol. 12, p. 107, 

1903. — Barnes and McDunnough, Contributions, vol. 3, p. 

199, 1916 (make synonym of leoninella). 

Forewing similar in maculation to that of fenestrella 
except: General color more ocherous than gray, the gray 
shading limited to the median area between the trans- 
verse lino; basal area of wing pale ocherous; the 
inner border of the antemedial and outer border of the 
subterminal lines yellow; lower fold between the trans- 
verse lines pale ocherous ; no appreciable black strealdng 
on the veins of outer area; discal spot at lower outer 
angle of ceU larger, more conspicuous. Hind wing paler, 
with a faint ocherous tint towards apex and termen. 
Alar expanse, 21-24 mm. 

Male and female genitalia like those of fenestrella. 

Type localities: California (leoninella, in MCZ); 
Salt Lake, Utah (pallidella, in USNM). 

Food plant: Unknown. 

Distribution: United States: California, "Mid- 
dle Calif.," Olancha (Inyo County, June), Palo Alto 
(May), Sonoma County (May); Utah, Richfield (Aug.), 
Salt Lake, Stockton (July), Vineyard (June, July). 
Canada: Manitoba, Cartwright (Aug.). 

Hulst treated leoninella as a variety of fenestrella, and 
probably correctly. There is nothing to separate the 
two except coloration. However, as nothing is known 
about their biology, it seems the better part of wisdom 
to keep the two names apart. Larval characters and 
habits and hosts may indicate separate species or at 
least distinct races. 

350. Lipographis truncatella (Wright), new combination 

FiQOEES 408, 898 

Hypochalcia truncatella Wright, Ent. News, vol. 27, p. 25, 1916. — 
McDunnough, Check list, No. 6276, 1939. 

Forewing a duU, pale, brownish ocherous, dusted with 
white and a fine peppering of black scales, making the 
general color an ashy gi"ay with a strong suffusion of the 
ground color, the latter most pronounced in basal area 
and in lower half of median area; antemedial line faint, 
without any appreciable inner border, its outer border 
indicated by black dots on costa, upper and lower veins 
of cell, and on vein lb; subterminal line distinct, with a 
narrow, faint, but distinguishable and continuous inner, 
black, bordering line. Hind wing pale gray, very faintly 
tinted with ocherous toward base and shading into a 
smoky hue towards apex and termen; a strong narrow 
dark shade along termen. Alar expanse, 22-25 mm. 

Male genitaha distinguished chiefly by its much 
slenderer aedeagus, narrowing at apex, and the single, 
very short, thornlike, curved cornutus on penis. Dif- 
ferences also in the shape of the sclerotization of the 
eighth segment tergite of abdomen are shown in the 
figuire. Female genitalia figured from a San Diego 
specimen in the National Collection (W. S. Wright, 
June 23, 1911). Bursa narrowly elongate; ductus bur- 



168 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



sae much shorter than bursa, and narrow (no wider than 
bursa, except at genital opening). 

Type locality: San Diego, Calif, (type probably 
lost). 

Food plant: Unknown. 

Disthibution: California, Chula Vista (June), San 
Diego (June). 

Despite its striking specific differences in genitalia 
and more broadly scaled labial palpi this species fits 
well into Lipographis. It is certainly not a Hypochal- 
cia. The latter, an Old World genus, as far as I know 
is not represented in our fauna. 

351. Lipographis umbreUa (Dyar), new combioation 
Figures 410, 897 

Sarata umbrella Dyar, Proc. Ent. Soc. Washington, vol. 10, p. 59, 
1908.— McDunnough, Check list, No. 6270, 1939. 

Male antenna with a weak ridge of roughened scales 
in shallow sinus at base of shaft. 

Forewing orange yellow; transverse lines narrow, 
white; antemedial line obhque, somewhat curved, set 
well out on wing and with only the faintest indication 
of a dark outer border, the latter sometimes containing 
a few black scales; sub terminal line nearly straight, with 
only a shght median bulge, inwardly more or less bor- 
dered with black, the latter color varying from a thin, 
weak line to large smudges extending well into the 
median area of the wing; on some specimens a narrow 
obhque blackish shade just beyond basal attachment of 
wing; discal dots obscure, often obhterated by streaks 
of white scaling or extensions of the black border of the 
antemedial hne. Hind wing semilustrous, ocherous 
with a smoky suffusion, the latter most pronounced on 
dark specimens; veins not appreciably darkened. Alar 
expanse, 26.5-31 mm. 

Male genitalia similar to those of fenestrella and 
leonineUa; differing from them chiefly in the armatm-e 
of the penis; the latter consists of a comb of 6 or 7 curved 
spines and another straight spiue, near but distinctly 
separated from the comb. Female genitaha differing 
only in minor details from those oiJenestreUa. 

Type locality : San Diego, Calif, (type in USNM) . 

Food plant: Unknown. 

Disthibution: Calijornia, Laguna, Long Beach 
(Sept.), Los Angeles (Sept.), Petaluma (Sept.), San 
Diego (Aug., Sept.). 

Dyar placed the species in Sarata on the basis of its 
male antenna character; but its genitalia as well as the 
lack of any sexual dimorphism in wing maculation or 
color show that it belongs in Lipographis. 

352. Lipographis (?) subosseella Hiilst 
Figure 409 

Lipographis subosseella Hulst, Canadian Ent., vol. 24, p. 62, 
1893.— Ragonot, Monograph, pt. 1, p. 565, 1893. 

Male antenna with a very weak scale ridge in shallow 
sinus at base of shaft. 

Thorax and forewing sordid white overshaded with 
dull ocherous; the whitish ground color most noticeable 



along costa, the ocherous shade strongest along lower 
fold and in outer area, making the general color of the 
wing (to the naked eye) a pale brownish yellow; trans- 
verse lines nearly obsolete; antemedial line distinguish- 
able only as an obhque whitish streak from lower vein 
of cell to inner margin, preceded on inner margin by a 
blackish brown smudge; sub terminal line indicated only 
by its dark borders, a pale brownish, rather broad outer 
band and a faint