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' CHECKLIST OF AMERICAN PHYCITINAE 


- Pages 317-329 from 


United States National Museum Bulletin 207 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


By Carl Heinrich 


Checklist of American Phycitinae 


(Synonyms in italics) 


1. CrypToBLaBEs Zeller 


1. 


gnidiella (Milliére): Europe, Africa, Asia, 
Bermuda, Venezuela, Brazil 


2. Acrosasis Zeller 


Mineola Hulst 
Seneca Hulst 
Acrocaula Hulst 


. indigenella (Zeller): Eastern U.S. and Can- 


ada, California 
nebulo (Walsh) 
nebulella (Riley) 
zelatella (Hulst) 


3. grossbecki (Barnes and McDunnough), new 


Doe 


comb.: Florida 


. vaccinii Riley: U.S. 
. amplexella Ragonot: Eastern U.S. 
. tricolorella Grote: U.S., Canada 


scitulella Hulst 


. comptella Ragonot: Western U.S. 
. minimella Ragonot: LHastern U. S. 


nigrosignella Hulst 


. feltella Dyar: Eastern U. S., Canada 
. palliolella Ragonot: Eastern U. S., Canada 


albocapitella Hulst 


. caryalbella Ely: U.S. (Connecticut) 

. juglandis (LeBaron): Eastern U.S. 

. sylviella Ely: Eastern U. S., Canada 

. kearfottella Dyar: Eastern U. S. 

. caryae Grote: Eastern U. S., Canada 

. evanescentella Dyar: U.S. (Florida) 

. stigmella Dyar: Eastern U.S. 

. aurorella Ely: Eastern U. S. 

. peplifera Dyar: Eastern U.S. 

. exsulella (Zeller), new comb.: Eastern U.S. 


septentrionella Dyar 


. angusella Grote: Eastern U. S., Canada 


eliella Dyar 


. demotella Grote: Eastern U. S. 

. latifasciella Dyar: Eastern U.S. 

. irrubriella Ely: Eastern U. S. 

. normella Dyar: Eastern U.S. (Connecticut) 
. malipennella Dyar: Eastern U. S. (Con- 


necticut) 


. dyarella Ely: Eastern U. S. (Connecticut) 

. ostryella Ely: Eastern U. S., Canada 

. secundella Ely: Eastern U. S., Canada 

. coryliella Dyar: Eastern U. S. 

. hebescella Hulst: Eastern U. S. (New Jer- 


sey) 


. cirroferella Hulst: Eastern U. S. (Texas) 
. cunulae Dyar and Heinrich: Eastern U. S. 
. caryivorella Ragonot: Eastern and South- 


western U.S. 


35. comacornella (Hulst), new comb.: Eastern 


U.S. (Texas) 

36. betulella Hulst: Eastern and Western U.S., 
Canada 

37. rubrifasciella Packard: Eastern U. S., 
Canada 


alnella McDunnough 
38. comptoniella Hulst: Eastern U. S., Canada 
39. myricella Barnes and McDunnough: U. S. 
(Florida) 
40. tumidulella (Ragonot), new comb.: U. S. 
(Florida) 


3. RHopDOPHAEA Guénée 


41. caliginella (Hulst), new comb.: U.S. (Cali- 
fornia, Arizona) 
caliginoidella (Dyar) 
42. supposita (Heinrich), new comb.: Canada 
(British Columbia) 


. TRACHYCERA Ragonot 


43. pallicornella (Ragonot): U.S. (Texas) 


. Anazasis Heinrich, new genus 


44. ochrodesma (Zeller), new comb.: U. S., 
(Florida), México, Guatemala, Panamé 
Colombia, West Indies 

crassisquamella (Hampson) 


. Miuprrxra Dyar 


45. constitutionella Dyar: México, Guatemala 


. SEMATONEURA Ragonot 


46. atrovenosella Ragonot: México, Costa Rica, 
Colombia, Ecuador, Pert, Argentina 
47. abitus Heinrich, new species: Ecuador 


. Hypsipyta Ragonot 


48. grandella (Zeller): U. S. (Florida), West 
Indies and Tropical America to Argen- 
tina 

cnabella Dyar 

49. ferrealis (Hampson), new comb.: ‘Tropical 
America (Costa Rica to Brazil) 

50. dorsimacula (Schaus), new comb.: Costa 
Rica 

51. fluviatella Schaus: Costa Rica 


. Hemrpritocera Ragonot 


52. chinographella Ragonot: French Guiana, 
Brazil, Pert 

53. bigrana (Zeller): México, Colombia 

54. plumigerella (Ragonot), new comb.: ‘‘Amer. 


Merid.”’ 

55. letharda (Schaus), new comb.: Panama, 
México 

56. jocarella (Schaus): Costa Rica, Panama, 
Brazil 


57. exoleta (Zeller): Colombia 
317 


‘ 
4 
? 
. 
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: i 
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318 


10. 


11. 


12. 


13. 


14. 


15. 


16. 


17. 


18. 


19: 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Crocipomera Zeller 
58. turbidella Zeller: 
U.S. (Texas) 
59. fissuralis (Walker): 
Puerto Rico 
adonea (Felder and Rogenhofer) 
60. stenopteryx (Dyar), new comb.: México 
Cuniperta Heinrich, new genus 
61. subtinctella (Ragonot), new comb.: 
ern U.S. and Canada 
Horas Heinrich, new genus 
62. disjunctus Heinrich, new species: 
Apanarsa Heinrich, new genus 
63. intransitella (Dyar), new comb.: 
(Arizona, New Mexico) 
Brrinus Heinrich, new genus 
64. russeolus Heinrich, new species: 
Guiana 
Buertetia Barnes and McDunnough 
65. grisella Barnes and McDunnough: U. S. 
(Arizona) 
Hypareynria Ragonot 


Cuba, Jamaica, México, 


Dominican Republic, 


West- 


Colombia 


U.S. 


British 


66. definitella (Zeller): Puerto Rico, Virgin 
Islands, Colombia, Brazil 
67. slossonella (Hulst), new comb.: U. S. 


(Florida), México 
tenuella (Barnes and McDunnough) 
Craranrica Heinrich, new genus 


68. annuliferella (Dyar), new comb.: U. S. 
(New Mexico, Arizona) 
69. hystriculella (Hulst), new comb.: U. S. 


(Texas, Florida) 

70. bicolorella (Barnes and McDunnough), new 
comb.: U.S. (Arizona, Nevada, Cali- 
fornia) 

Mye topsis Heinrich, new genus 

71. coniella (Ragonot), new comb.: 
ada, México 

nefas (Dyar) 

72. immundella (Hulst), 
(Texas) 

73. subtetricella (Ragonot), new comb.: 
Canada 

zonulella (Ragonot) 
obnupsella (Hulst) 

74. minutularia (Hulst), 
(Texas) 

75. alatella (Hulst), new comb.: 
rectistrigella (Ragonot) 
fragilella (Dyar) 
piazzella (Dyar) 

AnypsipyLa Dyar 

76. univitella Dyar: Cuba, México, Guatemala, 
Panam4, Venezuela, Brazil, Peri, Ecua- 
dor, Jamaica 


U.S., Can- 


new comb.: U. S. 


5s, 


new comb.: 


Western U.S. 


20. ApomyEtots Heinrich, new genus 


77. bistriatella (Hulst), new comb.: Eastern 
U.S., Canada 


bilineatella (Ragonot) 


Uy Ss - 


21. 


22. 


23. 


24. 


25. 


26. 


Ecromyetots Heinrich, new genus. 


78. decolor (Zeller), new comb.: Tropical 
America 
ephestiella (Hampson) 
79. ceratoniae (Zeller), new comb.: Europe, 


U. S. (Florida), Puerto Rico, Jamaica, 


Argentina 
oporedestella (Dyar) 
80. muriscis (Dyar), new comb.: Tropical 
America 
palpalis (Dyar) 
81. furvidorsella (Ragonot), new comb.: Puerto 
Rico 
82. zeteki Heinrich, new species: Panam&é 
ParAMYELO!Is Heinrich, new genus. 
83. transitella (Walker), new comb.: U. &S., 


tropical America 
notatalis (Walker) 
solitella (Zeller) 
duplipunctella (Ragonot) 
venipars (Dyar) 
cassiae (Dyar) 
Psrupopivona Dyar 
84. commensella Dyar: México 
85. cispha Dyar: Guatemala, Costa Rica, Brit- 
ish Honduras 
86. santa-maria Dyar: Guatemala 
87. carabayella Dyar: Pert, Bolivia, Colombia 
ProToMOERBES Heinrich, new genus 
88. aberrans Heinrich, new species: 
89. separabilis Heinrich, new species: 
Diatomocera Ragonot 
Cabima Dyar 
90. tenebricosa (Zeller): 
ana, Costa Rica 
91. dosia (Dyar), new comb.: 


Colombia 
Colombia 


Colombia, French Gui- 


Panama 


92. excisalis (Hampson), new comb.: French 
Guiana, Bolivia (?) 
93. decurrens (Dyar), new comb.: Panam& 
94. majuscula Heinrich, new species: Brazil 
95. albosigno Heinrich, new species: Brazil 
96. hoplidice (Dyar), new comb.: Panama 
97. extracta Heinrich, new species: Costa Rica 
98. mochlophleps (Dyar), new comb.: México 
Psrupocasima Heinrich, new genus 
99. castronalis Heinrich, new species: Brazil 
100. fearnella (Schaus), new comb.: Costa Rica, 
Guatemala 
101. guianalis Heinrich, new species: French 
Guiana, British Guiana 
102. euzopherella (Dyar), new comb.: Panam& 
103. pombra (Dyar), new comb.: Panama 
104. nigristrigella (Ragonot), new comb.: Brazil 
105. arizonensis Heinrich, new species: U. S. 
(Arizona) 
106. expunctrix (Dyar and Heinrich), new comb.: 
Brazil 
107. perrensiella (Ragonot, new comb.: Argen- 
tina 


Lae A 


: + tie 


27. 


28. 


29. 


30. 


31. 


32. 


AMERICAN MOTHS OF THE 


108. rubrizonalis (Hampson), new comb.: French 
Guiana, Brazil 
Hyatosrita Ragonot 


109. stictoneurella Ragonot: México, Guatemala, 
Brazil 

110. celiella Schaus: Costa Rica 

111. insequens Heinrich, new species: Bolivia, 
Colombia 

112. majorina Heinrich, new species: México 

113. fulgidula Heinrich, new species: Cuba 

114. egenella (Ragonot), new comb.: Brazil 

115. xanthoudemia (Dyar), new comb.: Panama, 
Costa Rica 

116. angulinella (Schaus), new comb.: Costa 
Rica 

117. clevelandella (Dyar): Panam4 


118. semibrunneella Ragonot: Colombia 

Funpewya Zeller 

119. pellucens Zeller: U.S. (Florida), West In- 
dies, Brazil, Bolivia 

cistipennis (Dyar) 

120. argentina Dyar: U. S. (Florida, Texas), 
West Indies, Venezuela, Brazil, Argen- 
tina 

eucasis Dyar 

121. agapella Schaus: Gal&pagos Islands 

122. ignobilis Heinrich: México, Guatemala, 
Costa Rica, Cuba, Puerto Rico, Haiti 

123. ahemora Dyar: México, Guatemala, Costa 
Rica 

DirunpeEtua Dyar 

124. corynophora Dyar: 
French Guiana 

125. subsutella (Schaus), new comb.: 


Guatemala, Panamé, 


Costa Rica 


126. distractor Heinrich, new species: Puerto 
Rico 
127. tolerata Heinrich, new species: Bolivia 


Coprarruria Ragonot 

128. dasypyga (Zeller): 

Promy.eEa Ragonot 

129. lunigerella Ragonot: Western U. S. and 

Canada. 
130. lunigerella glendella (Dyar): 
131. dyari Heinrich, new name: 
zimmermani (Druce) 
drucet (Dyar) 

132. druceii (Ragonot), new comb.: 

133. mindosis Dyar: México 

134. dasystigma Dyar: México 

ANADELOSEMIA Dyar 

135. senesciella (Schaus): Costa Rica 

136. tecmessella (Schaus): Costa Rica 

137. fifria Dyar: Guatemala 

138. base Dyar: Guatemala 

139. obstitella (Schaus), new comb.: Costa Rica 

140. texanella (Hulst), new comb.: U.S. (Texas, 
Florida), Puerto Rico, Cuba 

dulciella (Hulst) 

condigna Heinrich, new species: 

zona) 


Colombia, Guatemala 


Colorado 
México 


Guatemala 


141. U.S. (Ari- 


SUBFAMILY PHYCITINAE 


33. 


34. 


35. 


36. 


37. 


38. 


39. 


319 


Dasypyea Ragonot 

142. alternosquamella Ragonot: 
Canada 

stictophorella Ragonot 

Rampyitua Dyar 

143. orio Dyar: México 

144. polydectella (Schaus): 

145. subcaudata (Dyar), new comb.: 
Costa Rica, Brazil 


Western U. S., 


Costa Rica 
Guatemala, 


146. lophotalis Heinrich, new species: México, 
Guatemala 

Fuurapa Heinrich, new genus 

147. querna (Dyar), new comb.: Panama 

148. carpasella (Schaus), new comb.: Galipagos 
Islands 

Scoryuus Heinrich, new genus 

149. cubensis Heinrich, new species: Cuba 


Davara Walker 
Homalopalpia Dyar 
Eucardinia Dyar 
caricae (Dyar), new comb.: 
Tropical America 
dalera (Dyar) 


150. U. S. (Florida), 


151. columnella (Zeller), new comb.: Colombia 
152. nerthella (Schaus), new comb.: México, 
Guatemala, Costa Rica 
euthales (Dyar) 
153. paranensis (Dyar), new comb.: Brazil 
154. azonaxsalis Walker: Brazil 
155. interjecta Heinrich, new species: Puerto 


Rico, Dominican Republic 
156. rufulella (Ragonot), new comb.: Puerto Rico 
Sarasota Hulst 
Cuba Dyar 
157. plumigerella Hulst: U.S. (Florida) 
158. furculella (Dyar), new comb.: Cuba, Puerto 
Rico, Dominica, Virgin Islands 
159. ptyonopoda (Hampson), new comb.: 
ward Islands 
Presmopopa Zeller 
Discopalpia Ragonot 
Amphycitopsis Dyar 
160. rubicundella Zeller: Brazil 
161. xanthomera Dyar: Guatemala, 
Costa Rica, French Guiana 
zanthozona Dyar 
trichomata (Zeller): Colombia 
flavicans (Zeller): Colombia, French Guiana 
fratella Dyar 
ragonoti (Dyar), new comb.: 
temala, Costa Rica 
isabella (Dyar), new comb.: 
xanthopolys Dyar: Panama 
parva Heinrich, new species: Panama 
semirufella (Zeller): Colombia 
apocerastes Dyar: México, Costa Rica, 
French Guiana, Brazil, Dominica 
montella Schaus: Costa Rica 


Wind- 


Panama 


162. 
163. 


164. México, Gua- 


165. Costa Rica 
166. 
167. 
168. 
169. 


170. 


+ Hi be 


wf 
ah 


shiv 
SDA, 


320 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


40. AtHrtoca Heinrich, new genus 


41. 


42. 


43. 


44. 


45. 


46. 


47. 


48. 


171. subrufella (Hulst), new comb.: 

ida), Cuba, Virgin Islands 

filiolella (Aulst). Virgin Islands 
ptychis (Dyar) 


U.S. (Flor- 


172. bondari Heinrich, new species: Brazil 
PRAEDONULA Heinrich, new genus 
173. almonella (Dyar), new comb.: Panama 


Peavus Heinrich, new genus 


174. burdettellus (Schaus), new comb.: Costa 
Rica, Guatemala 
semproniella (Schaus) 
175. dissitus Heinrich, new species: Brazil 


176. subaquilellus (Ragonot), new comb.: Guate- 
mala 
Gasinius Heinrich, new genus 


177. paulsoni (Ragonot), new comb.: Chile 
CrracanTuHiA Ragonot 

Procandiopa Dyar 
178. mamella (Dyar), new comb.: Panama, 

Guatemala 

179. vepreculella Ragonot: Ecuador 
Mecarruria Ragonot 
180. peterseni (Zeller): Guatemala, Colombia, 


Brazil, Pert 


181. squamifera Heinrich, new species: Costa 
Rica 

182. frustrator Heinrich, new species: Costa Rica 

183. schausi Heinrich, new species: Costa Rica 

184. cervicalis Dyar: Cuba 

185. alpha Heinrich, new species: Guatemala, 
México, Costa Rica, Panamé, Bolivia, 
Brazil 

186. beta Heinrich, new species: México, Guate- 


mala, Costa Rica, Trinidad 
Drescoma Dyar 
187. cyrdipsa Dyar: México, Guatemala, Pan- 
ama, French Guiana 
188. cinilixa Dyar: Guatemala, Panamé 
Mowoptitora Hulst 


189. pergratialis (Hulst): U.S. 
grotella (Ragonot) 
nubilella Hulst 
Zamaetria Dyar 
190. dixolophella Dyar: Panam& 
191. pogerythrus Dyar: México, Guatemala 
192. hospitabilis Dyar: Cuba 
193. masculinus Dyar: Guatemala 
194. australella (Hulst), new comb.: Was: 
(Texas, Florida) 
bumeliella (Barnes and McDunnough): 
U.S. (Texas, Florida) 
195. fraterna Heinrich, new species: Cuba 
196. laidion (Zeller): U.S. (Florida), Tropical 
America 
deia Dyar 


striella Dyar 
197. ipsetona Dyar: 


399671—56——2 


Costa Rica 


49. 


ANEGCEPHALESIS Dyar 


198. arctella (Ragonot), new comb.: U. S. 
(Florida), Bahamas, Cuba 
cathaeretes Dyar 
50. Maatriopsts Heinrich, new genus 
199. denticosella (Dyar), new comb.: ‘Tropical 


51. 


52. 


53. 


54. 


55. 


America 
cristalis (Hampson) 
AncyLostomi1a Ragonot 
200. stercorea (Zeller): 
America 
ignobilis (Butler) 
diffissella (Zeller) 
201. sauciella (Zeller): Colombia 
202. argyrophleps Dyar: México, Guatemala 
203. euchroma Dyar: Brazil 
Carisranius Heinrich, new genus 
204. pellucidellus (Ragonot), new comb.: Puerto 
Rico, St. Vincent, Jamaica, Surinam, 
Brazil 
melanoplaga (Hampson) 
205. decoloralis (Walker), new comb.: 
Us: 
metagrammalis (Walker) 
Surfurellus (Hulst) 
floridellus (Hulst) 
206. guatemalellus (Ragonot), 
Guatemala 
Ertexua Zeller 
207. zinckenella (Treitschke): 
U. S., Tropical America 
etiella (Treitschke) 
schisticolor Zeller 
villosella Hulst 
rubribasella Hulst 
Guyptocera Ragonot 
208. consobrinella (Zeller): 


U.S. (Florida), Tropical 


Southern 


new comb.: 


Europe, Asia, 


Eastern U. S., Can- 


ada, 
busckella (Dyar) 
Pima Hulst 
209. boisduvaliella (Guénée), new comb.: 
rope, Canada 
albiplagiatella (Packard), new comb.: East- 
ern U. S., Canada 


Eu- 


210. 


211. albiplagiatella occidentalis Heinrich, new 
race: Western U.S. 

212. fosterella Hulst: Western U. S., Canada 

213. vividella (McDunnough), new comb.: Can- 
ada 

214. albocostalialis (Hulst), new comb.: Western 
U.S., Canada 

215. albocostalialis subcostella (Ragonot), new 


comb.: Southwestern U. S. 
fulvirugella (Ragonot,) Newcomb.: Western 
U.S. (California) 
granitella (Ragonot), new comb.: 


216. 
PANE Western 


piperella (Dyar) 


( 
; 
; : 
Pa Ph, 
bY, Win 
\ i 
, 
- ’ 
, 
y 
< 
j " 
“ , . 
I i f 
d B ret 
mer 
; = y 
4 
a 
‘ i 
fy jl 
mer f moval 
ay a aan 


56. 


57. 


58. 


59. 
60. 


61. 


62. 


63. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


218. parkerella (Schaus), new comb.: Western 
U.S. (Montana) 

InterseEctIo Heinrich, new genus 

219. denticulella (Ragonot), new comb.: North- 


western U. S., Canada 
220. columbiella (McDunnough), new 
Northwestern U. S., Canada 


comb.: 


221. ruderella (Ragnot), new comb.: ‘‘N. 
Amer.” (California?) 
222. niviella (Hulst) new comb.: U.S., Canada 


AmpBeEsa Grote 

223. laetella Grote: Western U. S., Canada 

224, walsinghami (Ragonot): Western U. S. 

monodon Dyar 

walsinghami mirabella Dyar, new status: 
U.S. (Southern California) 

lallatalis (Hulst): . Western U. S. (Nevada, 
Utah) 

Carast1a Hiibner 


225. 


226. 


227. bistriatella (Hulst), new comb.: Western 
U.S. (California) 

228. incorruscella (Hulst), new comb.: Western 
U.S. (California) 

229. actualis (Hulst), new comb.: Western U.S., 


Canada 
Immyrua Dyar 
230. nigrovittella Dyar: 
Oreana Hulst 
231. unicolorella (Hulst): Eastern U. S., Canada 
leucophaeella (Hulst) 
Oxysria Heinrich 


Eastern U. S. 


232. aliculella (Hulst), new comb.: Southwestern 
U.S. 
oberthuriella (Ragonot) 
233. furciferella (Dyar) new comb.: Southwest- 


ern U.S. (Arizona) 
SaLesriacus Heinrich, new genus 
234. odiosellus (Hulst), new comb.: Western 
Weise 
bakerella (Dyar) 
yumaella (Dyar) 
SaLEBrRiaRi4 Heinrich, new genus 
235. turpidella (Ragonot), new comb.: Southern 
WES: 
ademptandella (Dyar) 
nubiferella (Ragonot), new comb.: 
engeli (Dyar) U.S. 
annulosella (Ragonot), new comb.: 
(Texas, North Carolina) 
robustella (Dyar) 
tenebrosella (Hulst), new comb.: 
quercicolella (Ragonot) 
heinrichalis (Dyar) 
pumilella (Ragonot) new comb.: 
ern U.S. 
georgiella (Hulst) 
fructetella (Hulst) new comb.: 
rectistrigella (Dyar) 


236. 
237. 
238. 


U.S. 


UPS: 
239. Ves: 
240. Southeast- 


241. Uos; 


64. 


65. 


66. 


67. 


68. 


321 


QUASISALEBRIA Henrich, new genus 
242. admixta Heinrich, new species: 
Uss: 
OrtTHourPis Ragonot 
243. jugosella Ragonot: Eastern U. S., Canada 
244. pasadamia (Dyar), new comb.: U.S., Can- 
ada 
Potoprustis Ragonot 
245. arctiella (Gibson): 
Meroptera Grote 
Emmerita Hampson 
246. mirandella Ragonot: 
247. cviatella Dyar: 
248. pravella (Grote): 


Western 


Alaska, Canada 


Western U. 8. 
U.S. (illinois, Mississippi) 
U.S., Canada 


249. abditiva Heinrich, new species: Canada 
NepHorrreryx Hiibner 
250. subfuscella (Ragonot), new comb.: Eastern 


U.S., Canada 
semiobscurella (Hulst) 
251. delassalis Hulst: Western U. S. 
purpurella (Hulst) 
pudibundella (Ragonot) 


252. delassalis fraudifera Heinrich, new race: 
Canada (British Columbia), U. S.(Wash- 
ington) 

253. rubescentella (Hulst): U.S. 

254. fernaldi (Ragonot), new comb.: U. §&., 
Canada 

255. dammersi Heinrich, new species: Western 


U.S. (California, Arizona) 

dammersi floridensis Heinrich, new race: 
U.S. (Florida) 

vetustella (Dyar), new comb.: 
ada 


256. 


257. U. S., Can- 


258. inconditella (Ragonot), new comb.: West- 
ern U. S. (Arizona, Colorado) 
259. subcaesiella (Clemens), new comb.: U. S., 
Canada 
contatella (Grote) 
260. virgatella (Clemens), new comb.: U. S., 
Canada ; 
quinquepunctella (Grote) 
261. carneella Hulst: U.S., Canada 
inquilinella (Ragonot) 
262. basilaris Zeller: U.S., Canada 
263. termitalis (Hulst), new comb.: Western 
U.S., Canada 
levigatella (Hulst) 
264. termitalis yuconella Dyar, new status: 
Alaska 
265. bifasciella Hulst: U.S. (Arizona) 
nogalesella (Dyar) 
266. uvinella (Ragonot), new comb.: Eastern 


afflictella (Aulst) 
liquidambarella (Dyar) 
celtidella (Hulst), new comb.: 


267. Uns: 


ee REAR 


' { 
¥ 
et ake 
: winner | 
nia 
i Maik ins 2) 
vy 7 4 : Nai! | 


LAA Re 


‘ 
I Ere ity 


322 


69. 


70. 


dike 


72. 


73. 


74, 


75. 


76. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


268. rubrisparsella (Ragonot): U.S. 
rufibasella (Ragonot) 
croceella (Hulst) 
texanella (Hulst) 
269. gilvibasella Hulst: 
lacteella (Hulst) 
270. crassifasciella Ragonot: 
decipientella Dyar 
crataegella B. and McD. 
271. bisra Dyar: México 
Trascata Hulst 
272. reductella (Walker): Eastern U.S. 
gleditschiella (Fernald) 
Tusa Heinrich, new genus 
273. finitella (Walker), new comb.: 
Canada 
melanellus (Hulst) 
274. umbripennis (Hulst), new comb.: 
(Colorado) 
gillettella (Dyar) 
275. oregonella. (Barnes and McDunnough), new 
comb.: U.S. (Oregon) 
276. infinitella (Dyar), new comb.: 
HomoroerapHa Ragonot 
277. lanceolella Ragonot: Pert 
TELETHusIA Heinrich, new genus 
278. ovalis (Packard), new comb.: 
latifasciatella (Packard) 
geminipunctella (Ragonot) 
modestella (Hulst) 
279. rhypodella (Hulst), new comb.: 
gon’’) 
Puosus Heinrich, new genus 
280. brucei (Hulst), new comb.: 
281. funerellus (Dyar), new comb.: 
Canada 


U.S. (Texas) 
Eastern U.S. 


Eastern U.S., 


Uy: S: 


México 


U.S., Canada 


U.S. (“Ore- 


Western U. S. 
Western U.S., 


282. curvatellus (Ragonot), new comb.: Western 

283. incertus ‘Heinrich, new species: Western 
U.S. (California) 

Actrix Heinrich, new genus 

284. nyssaecolella (Dyar), new comb.: Eastern 
U.S. 

285. dissimulatrix Heinrich, new species: Eastern 
U.S. (Virginia) 

Styitopautpr1a Hampson 


West Indies, México 
U.S. (Texas, 


286. lunigerella Hampson: 
287. scobiella (Grote), new comb.: 
Colorado) 
decimerella (Hulst) 


288. argentinensis Heinrich, new species: Ar- 
gentina 

Pyza Grote 

289. fasciolalis (Hulst), new comb.: Canada 
(British Columbia) 

290. impostor Heinrich, new species: Western 
U.S., Canada 

291. aequivoca Heinrich, new species: Western 
Canada 


399671—56—_3 


ites 


292. insinuatrix Heinrich, new species: Canada 
(Manitoba) 
aenigmatica Heinrich, new species: U. S., 
Canada 
criddlella Dyar: Canada (Manitoba) 
fusca (Haworth), new comb.: Holarctic 
moestella (Walker) 
frigidella (Packard) 
cacabella (Hulst) 
triplagiatella (Dyar) 
hypochalciella (Ragonot), new comb.: North- 
western U. S., Canada. 
blackmorella (Dyar) 
hanhamella Dyar: Canada (Manitoba) 
scintillans (Grote): Western U. S. (Califor- 
nia) 
feella Dyar 


293. 


294. 


295. 


296. 


297. 
298. 


299. sylphiella Dyar: Northwestern U.S., Canada 

300. rainierella Dyar: Northwestern U.S. (Wash- 
ington) 

301. aeneella Hulst: Western U. S. (Colorado, 
Utah) 

302. aeneoviridella Ragonot: Western U. S., 
Canada 

303. metalicella Hulst: Western U.S. (Colorado, 
Utah) 


304. fasciella Barnes and McDunnough: North- 
western U. S. (California) 
nigricula Heinrich, new species: 

U.S. (Nevada) 
viridisuffusella Barnes and McDunnough: 
Western U.S. (California) 

Diorycrria Zeller 
Pinipestis Grote 
307. abietella (Denis and Schiffermiiller): 
ern Hemisphere 
decuriella (Hisbner) 
abietivorella (Grote) 
elegantella (Hulst) 
sysstratiotes Dyar: 
reniculella (Grote): 
ponderosae Dyar: 
California) 
majorella Dyar: México 
muellerana Dyar 
disclusa Heinrich: Eastern U.S. 
auranticella (Grote): Western U.S., Canada 
miniatella Ragonot 
zanthaenobares Dyar 
erythropasa (Dyar): 
(Arizona) 
315. horneana (Dyar): 


305. Western 


306. 


North- 


Guatemala 
Northern U.S., Canada 
Western U.S. (Montana, 


308. 
309. 
310. 
dll. 
312. 
313. 


314. Southwestern U, S. 


Cuba 


316. pygmaeella Ragonot: Eastern U.S. 
317. zimmermani (Grote): U.S., Canada 
delectella (Hulst) 
austriana (Cosens) 
318. cambiicola (Dyar): Western U. S. 
319. amatella (Hulst): Eastern U. S. 


320. albovittella (Hulst): Western U.S. 


Lf ARMS e 


Leia tee Ny cael 


Reem asap ye wit ie 
if) et Bil are | Peak avs Beak Ai 
a ae AA bay 33 


POG Tea at 


miler hn: 


fr; 


: ‘asta af 


78. 


a9. 


80. 


81. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


321. gulosella (Hulst), new comb.: Western U.S. 


(Colorado, New Mexico) 


322. baumhoferi Heinrich, new species: South- 
western U.S. (Arizona) 
323. subtracta Heinrich, new species: South- 


western U. S. (New Mexico) 
324. clarioralis (Walker): Eastern U. S. 
brunneella (Dyar) 
OrycToMEToPiIA Ragonot 
325. fossulatella Ragonot: 
ical America 
moeschleri (Ragonot) 
SaraTa Ragonot 


U. S. (Texas), Trop- 


326. edwardsialis (Hulst), new comb.: Western 
Wass 
polyphemella (Ragonot) 
327. Western 


pullatella (Ragonot), new comb.: 
USS 


328. punctella (Dyar), new comb.: México 

329. punctella septentrionaria Heinrich, new race: 
Western U.S. 

330. incanella (Hulst), new comb.: Western U.S. 

aridella (Dyar) 

331. atrella (Hulst), new comb.: Western U. S. 
(Colorado) 

332. caudellella (Dyar), new comb.: Western 
U.S., Canada 

333. dnopherella Ragonot: Western U.S. (Cali- 
fornia) 

334. nigrifasciella Ragonot: Western U.S., Can- 
ada 

335. cinereella Hulst: Western U. S. (Colorado) 


336. rubrithoracella (Barnes and McDunnough), 


new comb.: Western U.S. 

337. tephrella Ragonot: Western U. S. (Wash- 
ington) 

338. alpha Heinrich, new species: Canada (Sas- 
katchewan) 

339. beta Heinrich, new species: Western U. S., 
Canada 

340. gamma Heinrich, new species: Western 
U.S. (California) 

341. iota Heinrich, new species: Western U. S. 
(California) 

342. perfuscalis (Hulst): Western U. S. 

excantalis (Hulst) 

343. epsilon Heinrich, new species: Western U.S. 

344. phi Heinrich, new species: Western U.S. 

345. kappa Heinrich, new species: Western U.S. 
(Arizona) 

346. delta Heinrich, new species: Western U. S. 

PuitopEMa Heinrich, new genus 

347. rhoiella (Dyar), new comb.: Western U. S. 


Lipograpuis Ragonot 
348. fenestrella (Packard): 
fornia) 

humilis Ragonot 
349. leoninella (Packard): 


Western U.S. (Cali- 


Western U. S., Can- 


ada 
pallidella (Dyar) 


82. 


83. 


84. 


85. 


86. 


87. 


88. 


323 
350. truncatella (Wright), new comb.: South- 
western U. S. (California) 
351. umbrella (Dyar), new comb.: 
U.S. (California) 
352. subosseella Hulst: Bahamas 
Apvr.puta Heinrich, new genus 
353. petrella (Zeller), new comb.: 
rubiginella (Walker) 
rufinalis (Walker) 
hapsella (Hulst) 
354. ochripunctella (Dyar), new comb.: 
U.S. (California) 
Tora Heinrich, new genus 
355. galdinella (Schaus), new comb.: 
Islands 
Ura Walker 
356. lithosella (Ragonot), new comb.: 
western U. S., México 
luteella Hulst) 
roseitinctella (Dyar), new comb.: 
western U. S., México 
senta Heinrich, new species: 
U.S. (Texas, Arizona) 
rubedinella (Zeller), new comb.: 
ida), Tropical America 
translucida (Walker) 
rufescentalis (Walker) 
minualis (Walker) 
deprivalis (Walker) 
venezuelalis Walker 
pyrrhochrellus (Ragonot) 
Easmopatpus Blanchard 
360. lignosellus (Zeller): U.S., Tropical America 
angustellus Blanchard 
tartarella (Zeller) 
incautella (Zeller) 
major (Zeller) 
anthracellus Ragonot 
carbonella (Hulst) 
puer Dyar 
Acroncosa Barnes and McDunnough 
361. albiflavella Barnes and McDunnough: West- 
ern U. S. (California) 
362. albiflavella castrella Barnes and McDun- 
nough: Western U.S. (New Mexico) 
363. similella Barnes and McDunnough: West- 
ern U.S. (Nevada, Utah) 
PassaDENA Hulst 
364. flavidorsella (Ragonot): 
México 
canescentella (Hulst) 
constantella Hulst 
cinctella (Hulst) 
UxorHora Ragonot 
Acromeseres Dyar 
365. groteii Ragonot: Eastern U. S. 
tephrosiella Dyar 
366. guarinella (Zeller): 
dialithus (Dyar) 


Southwestern 


Wes: 


Western 


Galipagos 


South- 


BY lc South- 


358. Southwestern 


359. U.S. (Flor- 


Western U. S., 


Cuba, Colombia 


aly 
ee 
5 
' 
Dia be 
1: 
4 
Nia, Oy wig k 
4 ee “ 
‘ 


324 
89. 


90. 


91. 


92. 


93. 


94. 


95. 


96. 


oT. 


98. 


os 


UNITED STATES NATIONAL MUSEUM 


Cuorrera Dyar 

367. idiotes Dyar: Panamé4 

368. extrincica (Dyar), new comb.: 

369. postica (Zeller), new comb.: 

Tacoma Hulst 

370. feriella Hulst: Southwestern U.S. 
submedianella Dyar 

ApELPERGA Heinrich, new genus 

371. cordubensiella (Ragonot), new comb.: 

gentina 

Evumysia Dyar 

372. mysiella (Dyar): Western U. S. 

373. maidella (Dyar): Western U. S., Canada 
374. pallidipennella (Hulst), new comb.: West- 
ern U.S 
fuscatella (Hulst): 

fornia) 
376. semicana Heinrich, new species: 
U.S. (Washington) 
Drvitraca Barnes and McDunnough 
377. ochrella Barnes and McDunnough: South- 
ern U.S. (Florida) 
378. simulella Barnes and McDunnough: South- 
ern U. S. (Florida) 
379. parvulella Barnes and McDunnough: South- 
ern U.S. (Florida) 
380. parvulella consociata Heinrich, new race: 
Colombia 
Macrorrainia Ragonot 
Dolichorrhinia Ragonot 
381. aureofasciella Ragonot: Southwestern U.S., 
México 
382. placidella (Zeller): 
Ocata Hulst 
383. dryadella Hulst: Southern U.S. (Florida) 
platanella (Grossbeck) 
Vaupivia Ragonot 
Maricopa Hulst 
384. coquimbella Ragonot: 
385. lativittella (Ragonot): 
México 
aureomaculella (Dyar) 
386. walkerella (Ragonot), new comb.: 
ProrasiA Heinrich, new genus 
387. mirabilicornella (Dyar), new comb.: 
ern U.S. (California) 
Hererocrapuis Ragonot 
Mona Hulst 
388. morrisonella Ragonot: 
coloradensis Ragonot 
olbiella (Hulst) 
ignistrigella Ragonot 
palloricostella (Walter) 
STaupincerIA Ragonot 
389. albipenella (Hulst): 
olivacella Dyar 
perluteella Dyar 


Cuba 
Colombia 


Ar- 


Sioe Western U. S. (Cali- 


Western 


Brazil 


Chile 
Southwestern U. S., 


Chile 


West- 


U.S., México 


Western U.S. 


100. 


101. 


102. 


103. 


104. 


105. 


106. 


107. 


108. 


109. 


110. 


Lae 


112. 


BULLETIN 207 


Houustia Ragonot 
390. undulatella (Clemens): 
rubiginalis (Walker) 
obsipella (Hulst) 
fumosella (Hulst) 
Honora Grote 
391. mellinella Grote: U.S. 
ochrimaculella Ragonot 
subsciurella Ragonot: Western U. §, 
sciurella Ragonot: Western U. S. 
fornia) 
dotella Dyar: Western U. S. (California) 
montinatatella (Hulst): Western U. S. 
canicostella Ragonot 
perdubiella (Dyar), new comb.: Western 
Was: 


U.S., Canada 


392. 
393. (Cali- 
394, 
395. 


396. 


Hownorinus Heinrich, new genus 
397. fuliginosus Heinrich, new species: 
Oncoxasis Zeller 
Endommasis Hampson 
398. anticella Zeller: Tropical America 
nigritella (Hampson) 
Casotia Ragonot 
Encystia Hampson 
semidiscella Ragonot: Argentina 
schini (Berg): Argentina 
rhythmatica Dyar: Panamé 
cundajensis ‘(Zeller): Colombia 
impeditella (Zeller) 
bonhoti (Hampson), new comb.: 
Jamaica 
Canarsta Hulst 
404. ulmiarrosorella (Clemens): 
pneumatella (Hulst) 
ulmella (Ragonot) 
fuscatella (Hulst) 
gracilella Hulst 
feliculella Dyar 
Harnocua Dyar 
405. velessa Dyar: 
EKurytamasis Dyar 
406. ignifatua Dyar: 
Cuba 
Evurytamipia Ragonot 
407. ignidorsella (Ragonot): 
México, Panamé 
Wonperra Grossbeck 
408. neaeriatella Grossbeck: U.S. (Florida) 
Oxrpotumia Hampson 
Synothmia Hampson 
409. endopyrella Hampson: México, Bahamas 
bahamasella (Hampson) 
Sryntosasts Hampson 
410. rubripurpurea Hampson: 
Drv1ana Ragonot 
Dannemora Hulst 
411. eudoreella Ragonot: 
edentella (Hulst) 


Pert 


399. 
400. 
401. 
402. 


403. Bahamas, 


U.S., Canada 


Panamaé 


Panamdé, Puerto Rico, 


U. S. (Arizona), 


México, Brazil 


Eastern U. 8S. 


A 
rf 


PROP hay | 


113. 


114, 


115. 


116. 


ave 


118. 


119. 


120. 


121. 


122. 


123. 


124. 


125. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


PaxuatKa Hulst 

412. nymphaeella (Hulst): Eastern U. S. 
verecuntella (Grossbeck) 

CacozopHEra Dyar 


413. venosa Dyar: Guatemala 
Psorosina Dyar 
414. hammondi (Riley): Eastern and Central 


U.S., Canada 
angulella Dyar 
Parriciota Heinrich, new genus 
415. semicana Heinrich, new species: 
Pacontus Heinrich, new genus 
416. corniculatus Heinrich, new species: 
Rico 
AptunGa Heinrich, new genus 
417. macropasa (Dyar), new comb.: 
México 
418. imperfecta (Dyar), new comb.: 
ANDERIDA Heinrich, new genus 
419. sonorella (Ragonot), new comb.: 
U.S. (Arizona) 
placidella (Dyar) 
Cassrana Heinrich, new genus 
420. malacella (Dyar), new comb.: 
Puerto Rico, Virgin Islands 
Mescrinia Ragonot 
421. triloses Dyar: 
mosces Dyar 
pandessa Dyar: Guatemala 
bacerella Dyar: Cuba 
estrella Barnes and McDunnough: U. S. 
(Florida) 
moorei Heinrich, new species: British Guiana 
parvula (Zeller): Colombia 
commatella (Zeller): Colombia 
berosa Dyar: Panam4, Puerto Rico 
peruella Schaus: Peri 
430. discella Hampson: México, Guatemala 
431. indecora Dyar: México 
Nonta Ragonot 
Hypermescinia Dyar 
432. exiguella (Ragonot): 
lambella (Dyar) 
Puestinia Hampson 
433. costella Hampson: 
Comotia Dyar 
434. torsicornis Dyar: Panamé 
435. convergens (Dyar), new comb.: 
Bema Dyar 
Relmis Dyar 
436. neuricella (Zeller), new comb.: 
America 
myja Dyar 
437. fritilla Dyar: Guatemala 
438. ydda (Dyar), new comb.: Panamé, French 
Guiana 
439. yddiopsis (Dyar), new comb.: Cuba 
440. fifaca (Dyar), new comb.: Panama 


Utah 


Puerto 


Guatemala, 
Guatemala 


México, 


México, 


Panamé 
422. 
423. 
424, 
425. 
426. 
427. 


428. 
429. 


- 


Tropical America 


Jamaica, Puerto Rico 


Guatemala 


Tropical 


126. 


127. 


128. 


129. 


130. 


325 


Homorosoma Curtis 
Phycidea Zeller 
441. electellum (Hulst): U.S., México, Guate- 
mala, Cuba, British West Indies 
opalescellum (Hulst) 
texanellum Ragonot 
tenuipunctella Ragonot 
differtella Barnes and McDunnough 


442. stypticellum Grote: U.S., Canada 
uncanale Hulst 
443. striatellum Dyar: Southwestern U. S. 
444, oslarellum Dyar: Western U.S. 
445. oslarellum breviplicitum Heinrich, new race: 
Southwestern U.S. (California) 
446. illuviellum Ragonot: U.S. (Arizona, Colo- 
rado), México 
candidella Hulst 
447, illuviellum emendator Heinrich, new race: 
Western U.S. 
448. imitator Heinrich, new species: Southwestern 
U.S. (California) 
449. longiventrellum Ragonot: Chile 
noctividella Ragonot 
450. albescentellum Ragonot: Western U.S. 
elongellum Dyar 
451. impressale Hulst: Western U. S., Canada 
452. inornatellum (Hulst): Eastern U.S. 
453. deceptorium Heinrich, new species: U. S. 
(Pennsylvania), Canada 
454. discrebile Heinrich, new species, Brazil 
455. peregrinum Heinrich, new species: U. S. 
(California), Costa Rica 
456. vepallidum Heinrich, new species: Argentina 
457. ditaeniatellum Ragonot: Chile 
458. oconequensis (Dyar), new comb.: Pert 
459. assitum Heinrich, new species: Pert 
460. acmaeopterum Ragonot: Chile 
461. nimbosellum Ragonot: Chile 
462. unionellum Ragonot: México 
Paraconia Ragonot 
463. magellanella (Ragonot): Chile 


Rorrupa Heinrich, new genus 
464. mucidella mucidella (Ragonot), new comb.: 
Western U.S., Canada 
465. mucidella reliquella (Dyar), new comb.: 
Eastern U. S., Canada 
466. mucidella olivaceela (Ragonot), new comb.: 
Tropical America 
musiosum (Dyar) 
cubella (Dyar) 
467. mucidella affusella (Ragonot), new comb.: 
Argentina 
STREPHOMESCINIA Dyar 
468. schausella Dyar: 
Unapiuia Hulst 
Strymaz Dyar 


Cuba 


Siti ress 


Bias ts 
te 


ahs ) 


¢ 


Cf a aes estan 


326 


131. 


132. 


133. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


469. erronella (Zeller): Tropical America. 
ubacensis (Zeller) 
bipunctella (Hampson) 
dorae (Dyar) 
pyllis (Dyar) 
470. maturella (Zeller): Colombia, Guatemala, 
Cuba 
471. albidiorella (Richards and Thomason); new 
comb.: Pert 
472. floridensis Heinrich, new species: U.S. (Flor- 
ida) 
473. nasutella Hulst: U.S. (New Mexico) 


Laetitia Ragonot 
Laosticha Hulst 

474. coccidivora (Comstock): 
pallida (Comstock) 
dilatifasciella (Ragonot) 
hulstiit Cockerell 


U_S. 


475. coccidivora quadricolorella (Dyar), new 
comb.: Southwestern U. S. 

476. coccidivora cardini Dyar: Cuba, U. S. 
(Florida) 

477. obscura Dyar: Cuba 

478. portoricensis Dyar: Puerto Rico 

479. melanostathma (Meyrick), new comb.: Ar- 
gentina 

480. amphimetra (Meyrick), new comb.: Argen- 
tina 

481. zamacrella Dyar: Western U.S. (California) 

482. myersella Dyar: Hastern U.S. 

483. ephestiella (Ragonot): Southwestern U. S. 
(Arizona) 

lustrella (Dyar) 
484. fiskella Dyar: Eastern U. S. (North Caro- 


lina) 


485. glomis (Dyar), new comb.: Panamé 
Bapnata Heinrich, new genus 
486. basimaculatella (Ragonot), mew comb.: 


Western U. S. 
eremiella (Dyar) 
487. goyensis (Ragonot), new comb.: 
Uruguay, Argentina 
488. goyensis olivacea Heinrich, new race: Argen- 
tina 
489. homoeosomella (Zeller), new comb.: 
ical America. 
bodkini (Dyar) 
rusto (Dyar) 
taboga (Dyar) 
saissetiae (Dyar) 
490. haywardi Heinrich, new species: Argentina 
491. glabrella (Dyar), new comb.: Guatemala 
492. squalida (Walker), new comb.: Brazil 
Raacea Heinrich, new genus 
493. packardella (Ragonot), new comb.: 
ern U.S. 
orobanchella (Dyar) 
494. stigmella (Dyar), new comb.: 
U.S. (California), México 
maculicula (Dyar) 


Brazil, 


Trop- 


West- 


Southwestern 


134. 


135. 


136. 


137. 


138. 


139. 


140. 


141. 


142. 


143. 


Zopuopia Hiibner 
Dakruma Grote 
495. convolutella (Hiibner): 
ada 
grossulariella (Hiibner) 
turbatella (Grote) 
grossulariae (Riley) 
franconiella (Hulst) 
bella Hulst 
ihouna Dyar 
dilativitta Dyar 
magnificans Dyar 
Metitara Walker 


Europe, U.S., Can- 


496. prodenialis Walker: U.S. 
bolliz (Zeller) 
497. dentata (Grote): U.S. 


doddalis Dyar 

Otycetia Dyar 

498. junctolineella (Hulst): Southern U.S. (Texas) 

499. junctolineella pectinatella (Hampson): Méx- 
ico 

500. nephelepasa (Dyar:) México 

501. subumbrella Dyar: Western U.S. 

Otyca Walker 

502. phryganoides Walker: 
lic, Haiti 

ALBERADA Heinrich 

503. parabates (Dyar): 

504. bidentella (Dyar): 
(Texas, Arizona) 

505. holochlora (Dyar): 
(Texas) 

Nanaia Heinrich 

506. substituta Heinrich: Pert 

Cactostastis Ragonot 

Neopyralis Bréthes 


Dominican Repub- 


U.S., México 
Southwestern U. S. 


Southwestern U. S. 


507. cactorum (Berg): Argentina, Uruguay, Aus- 
tralia 

508. ronnai (Bréthes): Brazil 

509. doddi Heinrich: Argentina 

510. mundelli Heinrich: Pert 


511. bucyrus Dyar: 
CauEta Heinrich 
512. ponderosella (Barnes and McDunnough): 
Western U.S., México 
purgatoria (Dyar) 
interstitialis (Dyar) 
phoenicis (Dyar) 
Rumatua Heinrich 
513. glaucatella (Hulst): Southern U. S. 
514. bihinda (Dyar): Western U.S. 
515. polingella (Dyar): Southwestern U.S. (Ari- 
zona, Texas) 
Yosremit1a Ragonot 
516. graciella (Hulst): Western U.S. 
517. longipennella (Hulst): Southwestern U. S. 
(Texas) 
518. fieldiella (Dyar): 
Arizona) 
519. didactica Dyar: 


Argentina 


Western U.S. (California, 


México 


144. 


145. 


146. 
147. 


148. 


149. 


150. 


151. 


154. 


155. 


156. 


157. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Tucumania Dyar 
520. tapiacola Dyar: 
521. porrecta Dyar: 
EREMBERGA Heinrich 
522. leuconips (Dyar): 
523. creabates (Dyar): 
fornia) 
524. insignis Heinrich: México 
SALAMBONA Heinrich 
525. analamprella (Dyar): 
Parotyca Dyar 
526. asthenosoma (Dyar): 
Sieeueaira Heinrich 
527. chilensis Heinrich: Chile 
528. huanucensis Heinrich: Pert 
529. transilis Heinrich: Pert 
AMALAFRIDA Heinrich 
530. leithella (Dyar): 
Colombia 
Ozamia Ragonot 
531. lucidalis (Walker): West Indies 
532. fuscomaculella (Wright): Southwestern 
U.S. (California) 
heliophila Dyar 
fuscomaculella clarefacta Dyar: Uz. S. 
(Texas), México 
thalassophila Dyar: 
immorella (Dyar), new comb.: 
stigmaferella Dyar: Argentina 
537. hemilutella Dyar: Argentina 
538. punicans Heinrich: Argentina 
Cactosrosis Dyar 
539. fernaldialis (Hulst): Southwestern U. S. 
gigantella (Ragonot) 
cinerella (Hulst) 
longipennella (Hampson): 
elongatella (Hampson): 
maculifera Dyar: México © 
insignatella Dyar: México 
strigalis (Barnes and McDunnough): West- 
ern U. S., México 


Argentina 
Uruguay 


Western U. S. (Arizona) 
Western U. S. (Cali- 


Argentina 


French Guiana 


West Indies, Venezuela, 


533. 


534. 
535. 
536. 


U.S. (California) 
México 


540. México 
541. 
542. 
543. 


. Drescomopsis Dyar 


544. soraella (Druce): 
drucella (Dyar) 
subelisa Dyar 


Tropical America 


. Intatiza Dyar 


545. gurbyris Dyar: Panamé 

Lascetina Heinrich, new genus 

546. canens Heinrich, new species: 
U.S. (Texas), México 

Metepuestia Ragonot 

547. simplicula (Zeller): U.S. (Florida), Puerto 
Rico, Colombia, British West Indies 

Sretaa Heinrich, new genus 

548. arizonella (Hulst), new comb.: 
ern U.S. (Arizona) 

Entmemacornis Dyar 

549. proselytes Dyar: Guatemala 

550. pulla Heinrich, new species: Brazil 


Southern 


Southwest- 


158. 
159. 


160. 


161. 


162. 


163. 


164. 


165. 


166. 


167. 


168. 


169. 


170. 


327 


Cayrnni1a Hampson 
551. rufitinctalis Hampson: 
Riosa, Heinrich, new genus 


French Guiana 


552. nexa Heinrich, new species: Argentina 
Moerrszgs Dyar 
553. dryopella (Schaus): Costa Rica 


554. alveolella (Ragonot), new comb.: Brazil 
555. emendata Heinrich, new species: Panamé, 
French Guiana 
Moopnopsis Dyar 
Campyloplesis Dyar 
556. decipiens Dyar: México 
557. perangusta (Dyar), new comb.: Trinidad 
558. inornatella (Ragonot), new comb.: Costa 
Rica, Brazil 
559. parallela Heinrich, new species: Brazil, Pera 
560. inveterella (Dyar), new comb.: Guatemala 
561. portoricensis Heinrich, new species: Puerto 


Rico 
Eputica Ragonot 
562. compedella (Zeller): 
KuzorHera Zeller 


Tropical America 


563. semifuneralis (Walker): U. S., Canada, 
México 
aglaeella Ragonot 
pallulella (Hulst) 
564. ostricolorella Hulst: Eastern U. 8S. 


565. nigricantella Ragonot: Southwestern U. S., 
México 
griselda Dyar 
Exvurprrius Heinrich, new genus 
566. negator Heinrich, new species: 


Evtoetra Heinrich, new genus 


Pert 


567. ochrifrontella (Zeller), new comb.: U. S., 

Canada 
ferruginella (Ragonot) 

ProsonuzopHeErRA Heinrich, new genus 

568. impletella (Zeller), new comb.: Colombia, 
Jamaica, Puerto Rico 

Farnosia Heinrich, new genus 

569. quadripuncta (Zeller), new comb.: Costa 


Rica, Panam4, French Guiana, Colombia 
Gennavius Heinrich, new genus 
570. junctor Heinrich, new species: 

ana 
Micromescinia Dyar 
571. pygmaea Dyar: 
Epuestiopgs Ragonot 
572. gilvescentella Ragonot: 

ada, México 
nigrella Hulst 
infimella Ragonot: 
erythrella Ragonot: 
coloradella (Hulst) 
benjaminella Dyar 
mignonella Dyar: U.S. (Texas) 
erasa Heinrich, new species: U.S. (Florida) 
lucidibasella Ragonot: Chile 
productella Ragonot: Colombia (?) 
indentella Dyar: Bermuda 


French Gui- 


Panamaé 


Western U.S., Can- 


Eastern U. S. 
Western U.S., Canada 


573. 
574. 


575. 
576. 
577. 
578. 
579. 


Hea 


PRS 


ants 


TD ine 


i De apdbritsy ~ 


FO hae a 
ra A 


328 


171. 


172. 


173. 


174. 


175. 


176. 


177. 


178. 


179. 


180. 


UNITED STATES NATIONAL MUSEUM BULLETIN 


580. plorella Dyar: Panam& 
vestilla (Dyar) 
581. stictella (Hampson), new comb.: 
West Indies 
uniformella Hampson 
granulella Hampson 
582. noniella Dyar: Panama 
Azazura Schaus 
Calamophleps Dyar 
583. muciella Schaus: Costa Rica, Guatemala, 
Panama 
squalidella (Dyar) 
584. nodoses (Dyar), new comb.: 


Bahamas, 


Panama 


585. lophophera (Dyar), new comb.: Panama 
Moopna Hulst 
586. ostrinella (Clemens): U.S., Canada 


obtusangulella (Ragonot) 
pelviculella Hulst 
587. bisinuella Hampson: 
Vituta Ragonot 
588. edmandsae (Packard): Eastern U. S., Can- 
ada 
dentosella Ragonot 


México, U.S. (Texas) 


589. edmandsae serratilineella Ragonot, new sta- 
tus: Western U.S., Canada 
590. lugubrella Ragonot, new comb.: Western 
U.S. (California) 
591. pinei Heinrich, new species: Western U. S. 
(Utah, Nevada) 
592. inanimella (Dyar), new comb.: México, 
Guatemala 
ticitoa (Dyar) 
593. laura (Dyar), new comb.: Guatemala 


Manuatta Hulst 
Hornigia Ragonot 
594. setonella (McDunnough), new comb.: U.S. 
(Utah), Canada (British Columbia) 
595. broweri Heinrich, new species: Eastern U.S. 
(Maine) 
Verna Heinrich, new genus 


596. supplicella (Dyar), new comb.: México, 
Guatemala, Panam4, Brazil 

Vacozpanta Heinrich, new genus 

597. divergens (Butler), new comb.: Chile 

Moopnetua Heinrich, new genus 

598. paula Heinrich, new species: Guatemala, 


Brazil, Argentina 

Vouatica Heinrich, new genus 

599. pachytaeniella (Ragonot), new comb.: Bra- 
zil 


600. trinitatis Heinrich, new species: Trinidad 


Vezina Heinrich, new genus 
601. parasitaria Heinrich, new species: 
tina, Brazil 

Caupgtuia Dyar 

602. apyrella Dyar: Eastern U. S. (Maryland) 

603. albovittella Dyar: Eastern U. S. 

604. nigrella (Hulst), new comb.: Western U.S. 
arizonella (Walter) 


Argen- 


181. 


182. 


183. 


184. 


185. 


186. 


187. 


207 
605. declivella (Zeller), new comb.: Panama, 
Colombia 
animosella (Dyar) 
606. colorella (Dyar), new comb.: Panama 


607. clara Heinrich, new species: Puerto Rico 

Micropuestia Dyar 

608. animalcula Dyar: Panam& 

Sosrpatra Heinrich, new genus 

609. rileyella (Ragonot), new comb.: Western 
U.S., México 

micaceella (Hampson): México 

anthophila (Dyar), new comb.: 
U.S. (Texas) 


610. 


611. Western 


612. thurberiae (Dyar), new comb.: Western U.S. 

613. nonparilella (Dyar), new comb.: Western 
U.S. (Arizona) 

614. majorella (Dyar), new comb.: México 

615. divergens (Dyar): Panamé 

Berruuria Ragonot 

616. championella Ragonot: Guatemala 


Risva Heinrich 

617. innoxia Heinrich: Cuba 

618. contigua Heinrich, new species: 

619. patriciella (Dyar), new comb.: 

Propra Guénée 

620. interpunctella (Hiibner): 

interpunctalis (Hiibner) 
zeae (Fitch) 

latercula (Hampson) 
glycinwora (Matsumura) 

621. dolorosa Dyar: Guatemala 

AnaGastA Heinrich, new genus 

622. kiihniella (Zeller): Cosmopolitan 

tuscofasciella (Ragonot) 
gitonella Druce 

Epuestia Guénée 

Hyphantidium Scott 
623. elutella (Hiibner): Cosmopolitan 
elutea (Haworth) 
semirufa (Haworth) 
rufa (Haworth) 
sericarium (Scott) 
roxburghit Gregson 
unicolorella Staudinger 
amarella Dyar 

624. cautella (Walker): 

defectella (Walker) 
desuetella (Walker) 

cahiritella Zeller 

passulella Barrett 

formosella (Wileman and South) 

625. figulilella Gregson: Europe, Asia, Africa, 
Hawaii, Australia, North America (U.S., 
California), South America 

~ ficulella Barrett 
milleri Zeller 
figuliella Forbes 
Jigulella Curran 
venosella 'Turati 
ernestinella Turati 


Puerto Rico 
Cuba 


Cosmopolitan 


Cosmopolitan 


ne Leer 
Res Se, 
Rey Pts 
eer inna 
PDE EU 
i an 


aD AAC on 


188. 


189. 


190. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Nicrtioprs Schaus 
626. apianella Schaus: 


VaRNERIA Dyar 


Galapagos Islands 


627. postremella Dyar: Eastern U.S. 
628. nannodes Dyar: Panama 
629. atrifasciella Barnes and McDunnough: 


Southern U. S. (Florida) 
630. dubia Heinrich, new species: 
Evrytumia Ragonot 
631. hospitella (Zeller): 
U.S 


Puerto Rico 


Southern and Western 


spaldingella Dyar 
632. hospitella yavapaella Dyar, new status: 
Western U. S. 
633. angulella Ely: Eastern U. S., Canada 
diffusella Ely 
634. fumella Ely: Eastern U.S. (Connecticut) 


191. 


192. 
193. 


194. 


329 


Ere.ieva Heinrich, new genus 
635. quantulella (Hulst), new comb.: 
U.S. (Texas), West Indies 
santiagella (Dyar) 
636. coca (Dyar), new comb.: 
coquilla (Dyar) 
mossa (Dyar) 
uncta (Dyar) 
637. parvulella (Ely), new comb.: 
(Connecticut) 
Casnia Dyar 
638. myronella Dyar: 
Microprycita Dyar 
639. titillella Dyar: Panamé 
Rasrria Heinrich, new genus 
640. conops (Dyar), new comb.: 


Southern 


Panama 


Eastern U. S. 


Eastern U. S. 


Panama 


Species unplaced or unrecognized 


brevistrigella Ragonot [Zophodia] 
came Dyar [EHuzophera] 
cervinistrigalis Walker [Hypochalcia] 
clitellatella Ragonot [Hornigia] 
corrientellus Ragonot [Elasmopalpus] 
daedalella Ragonot [Luzophera] 
disticta Zeller [Psorosa] 

dulciella Hulst [Honora] 

famula Zeller [Myelois} 

flavicornella Ragonot [Phycitopsis] 
formulella Schaus [Moodna] 
fuscifrontella Zeller [Nephopteryz] 


gais Dyar [Euzophera] 


grossipunctella Ragonot [Myelois] 
hulstiella Ragonot [Hypochalcia] 
infusella Zeller [Myelois] 
intextella Zeller [Huzophera] 
trichampa Dyar [Anthropteryz] 
megalopalis Hampson [Huzopherodes] 
nigricans Hulst [Salebria] 

olivella Hampson [Moodna] 
postflavida Dyar [Euzophera] 
putidella Schaus [Hucampyla] 
rinmea Dyar [Huzophera] 
subcanella Zeller [Zophodia] 


U, S, GOVERNMENT PRINTING OFFICE; 1956 


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UNITED STATES NATIONAL MUSEUM BULLETIN 207 


AMERICAN MOTHS 
OF THE SUBFAMILY 
PHYCITINAE 


By CARL HEINRICH 


SMITHSONIAN INSTITUTION +¢ WASHINGTON, D.C., 1956 


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The scientific publications of the National Museum include two series, known, respec- 
tively, as Proceedings and Bulletin. 

The Proceedings series, begun in 1878, is intended primarily as a medium for the pub- 
lication of original papers, based on the collections of the National Museum, that set forth 
newly acquired facts in biology, anthropology, and geology, with descriptions of new forms 
and revisions of limited groups. Copies of each paper, in pamphlet form, are distributed 
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in the different subjects. The dates at which these separate papers are published are 
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The present work forms No. 207 of the Bulletin series. 


RemMIncron KELLoae, 
Director, United States National Museum. 


UNITED STATES 
GOVERNMENT PRINTING OFFICE 
sk SHINGTON : 1956 


41 8 1956 


For sale by the Superintendent of Documents, U. S. Government Printing Office 
Washington 25, D. C. Price $5.50 


II 


Contents 


Introuuctionr st sot) iene eo AES AL he UD NRE YA ERD g 


ianuly Phymtidae see SOS SPNCMUSR |, FL, BO WOOT, BON ea 
sl Pea a ONE ER cuter ten tienes ks ss aise 1s 


Genera and species unplaced, unrecognized, or referred from the Phycitinae : . . . 


@hecklishorAmericansehycitinae: ss tes SSS US PS A I Ss 


Page 


oy , om ah aaa “aw Mathonnd Bec. 
ea A iN aes denne nen f 

DR age a 

a witey 


bigs i he PAP is at hi 

pei eRe es is AN: ye a a pais 
AES, tis ata al et {anit ovine ination 
‘hy bef TTA as ha 
rs cartiolk ia. fas wi on 


Sanh se nr 


tigi coi mse a " nate aowhpohey ! ice 
Aa Pucionhaidy oh Wetaral.” relyoitaehy: Entiat’ etalon’ geek ha oft real aN, 
pane egtint vodele) qolitouona, ak obher tan) ogi of sigan mpetui, 
nalitnemrae water “useing, dyad qoaaarbe ase ie Vj liye 


Dv i pet a, aa pale. a Heidi be iy sia 


Introduction 


This paper completes a 25-year study of the New 
World moths of the subfamily Phycitinae. It is based 
chiefly on the collections in the United States National 
Museum and the Hulst collection, formerly at Rutgers 
University, supplemented by material from the Cornell 
and Canadian national collections and specimens— 
mostly tropical American—from the British Museum, 
the Janse collection, and the collections of several South 
American lepidopterists. 

Recognized and included in the classification are 194 
genera, 619 species, and 21 subspecies (local races). Of 
these, 60 genera, 81 species, and 8 races are described 
as new. The new species and races represent only a 
fraction of the undescribed material examined. The 
remainder consists mostly of females, chiefly from trop- 
ical America and without authentically associated males 
or host plants. Their description would have added 
nothing to our scientific knowledge and the additional 
names would have been only a nuisance to other workers. 
Already too many names have been given such material. 


Acknowledgments 


A work of this kind could not be carried through 
without generous assistance from other entomologists. 
To each of them I owe a debt of gratitude: To Carl 
Muesebeck, Chief of the Division of Insect Investi- 
gation of the U. S. Bureau of Entomology and Plant 
Quarantine, for his support and encouragement at all 
stages of the project; toB. B. Pepper, State Entomologist 
of New Jersey, and John B. Schmitt for permission to 
examine the genitalia of the Hulst types and for their 
courtesies to me at Rutgers University; to J. Bourgogne 
of the Muséum d’Histoire Naturelle, Paris, for the 
privilege of studying the genitalia of the Ragonot 
types of American species; to N. D. Riley and to 
W. H. T. Tams for the loan of unidentified tropical 
American Phycitinae from the British Museum (Natural 
History) and to Tams especially for photographs of 
many types and their genitalia; to Martin Herring of the 
Zoologisches Museum der Universitat, Berlin, for the 
loan of Ragonot types; to W. T. M. Forbes, Department 
of Entomology, Cornell University, for the loan of his 
extensive collections from Puerto Rico, the Virgin Is- 
lands, and Surinam; to J. McDunnough and T. N. Free- 
man, Canadian National Museum, Ottawa, for the loan 
of Canadian specimens; to A.J.T.Janse of the Transvaal 
Museum, Pretoria, South Africa, for the loan of South 
American Phycitinae from his collection and for much 
valuable information; to A. da Costa-Lima, Escola 
Nacional de Agronomia, Universidad Rural, Distrito 
Federal, Brazil, for the loan of Brazilian specimens; to 


Frank Morton Jones for a gift of Phycitinae collected at 
Martha’s Vineyard, Mass.; to John A. Comstock for a 
loan of southern California specimens; to my colleagues 
at the U. S. National Museum—to J. F. Gates Clarke, 
for extensive notes on the phycitid types in the Museums 
of Paris, London, Oxford, and Berlin, and to Hahn 
Capps, for assistance in the tedious business of slide 
preparations. 

My greatest debt is to the artists of the Bureau of 
Entomology and Plant Quarantine for the drawings ac- 
companying this paper. Where genitalia are used in in- 
sect classification verbal descriptions are not enough. 
Figures must accompany and supplement them to give 
the reader a true picture of structural characters. The 
drawings in this paper were begun in 1930 by Eleanor 
A. Carlin and continued by her until October 30, 1940, 
when she retired from the Bureau. From that time the 
drawings were made by Sara H. De Bord, who has made 
the majority of the drawings here published. Her con- 
tribution was of especial value because she was not only 
a capable artist but a trained entomologist as well, and 
her interest in the paper and her devotion to her share 
in it was so complete that she worked well on into her 
last illness (she was retired on disability August 12, 
1948, and died March 12, 1950). Since her death some 
drawings were made by Arthur Cushman and Addie 
Egbert, and the former did most of the assembling of 
the plates. The drawings were all made under my 
supervision and for any inaccuracies in them I am alone 
responsible. 

The indices were prepared by Mrs. Marguerite W. 
Poole. 


Abbreviation of references 


To conserve space and eliminate useless repetition, 
titles to certain publications frequently cited are here 
abbreviated as follows: 

The Ragonot ‘‘Monographie des Phycitidae et des 
Galleriidae,” published as vol. 7 (1893) and vol. 8 
(1901, completed by Hampson) of the Romanoff 
“Mémoires sur les Lépidoptéres,”’ is cited as ‘‘Mono- 
graph, pt. 1,” or “Monograph, pt. 2.” 

Ragonot’s ‘“‘Diagnoses of North American Phycitidae 
and Galleriidae,” 1887, is cited as ‘‘N. Amer. Phyciti- 
dae,” and his ‘‘Nouveau Genera et Espéces de Phyci- 
tidae et Galleriidae,” 1888, as ‘‘Nouv. Gen.” 

Walker’s ‘List of the Specimens of Lepidopterous 
Insects in the Collections of the British Museum,” 
1854-66, is cited as “‘List.” 

Hulst’s ‘‘The Phycitidae of North America,’”’ pub- 
lished in the Transactions of the American Entomologi- 

v 


VI INTRODUCTION 


cal Society, vol. 17, pp. 93-228, pls. 6-8, March—July 
1890, is cited as “‘Phycitidae of N. Amer.” 

Barnes and McDunnough, “Contributions to the 
Natural History of the Lepidoptera of North America,” 
vols. 2 (1913-1914) and 3 (1916-1917), is cited as 
“Contributions.” 

McDunnough’s “Check List of the Lepidoptera of 
Canada and the United States of America, Part II, 
Microlepidoptera,” published in the Memoirs of the 
Southern California Academy of Science, vol. 2, No. 1, 
1939, is cited as ‘Check list.” 

Forbes’s ‘‘Lepidoptera of New York and Neighboring 
States,” Cornell University Agricultural Experiment 
Station Memoir 68, 1923, is cited as ‘Cornell Mem. 68.” 

The distributional records for species in this paper are 
obviously incomplete. They are based (with a few 
exceptions noted in the text) solely on specimens I 
have examined. This was the only safe procedure. 
So many misidentifications have been made in the past, 
even by lepidopterists of repute, that the records in 
literature can not be accepted merely on the authority 
of an author. Unless the specimens upon which his 
statements were based can be examined and the state- 
ments themselves verified, it is best to ignore them. 
By taking them simply on faith and repeating them we 
not only run the risk of perpetuating error, but do an 
injustice to past workers who did not have or could not 
use the evidence available to us. 

This caution applies with even greater force to 
“accepted” generic and specific synonymy. J have been 
very fortunate in being able to examine the genitalia 
of so many holotypes and in having authentic specimens 
of most of the type species of described genera occurring 
in the New World. All unqualified synonymy in this 
paper is based upon genitalic examination of such 
material. From the synonymy of some species common 
to both the New and Old Worlds I have omitted some 
names—chiefly of Old World synonyms—because I 
could not examine their types and had no certainty as 
to correctness of their synonymizing. Such omissions 
are discussed in the text. 


Classification and arrangement 


A general revision has a twofold purpose, a taxonomic 
and a practical one: To define accurately, to delineate 
as nicely as possible, and to name categories which, 
as far as our knowledge permits, represent objective 
realities in nature; and to arrange these categories in 
an order that permits their ready identification. Both 
purposes must be served if the revision is to have any 
value as a contribution to knowledge or to be of 
practical use to other workers. 

To satisfy both requirements I have adopted in this 
paper a dual classification: a definition and division 
into named categories of races, species, genera and 
subfamilies; and an artificial, unnamed division, 
between genus and subfamily, into groups of genera 
or, in a few instances within a single genus, into groups 
of species. 


The named categories themselves are more or less 
tentative. They are not adequate expressions of the 
truth. They are only approximations to it. As we 
learn more we shall have to amend or replace our defini- 
tions and the categories will come a little closer to the 
realities they represent. The names (except for 
homonyms) will always be available; but the concepts 
will change. There are several indications that taxo- 
nomic groupings between genus and subfamily may 
eventually be possible; and that when we have a clearer 
picture of host relations and larval characters, and more 
extensive collections from unexplored regions, we may 
be able to establish tribes on a legitimate taxonomic 
basis; but at present this is impossible. What few 
definite derivations we can trace from genus to genus 
show that tribal groupings would cut across the lines 
of any artificial system we might be able to use. 

The artificial system here adopted (based on vena- 
tional characters) is proposed merely for key purposes. 
The keys themselves, except for the one separating the 
subfamilies, are in no true sense a part of the taxonomic 
system. They are keys, and keys only. They are 
intended merely to open a ready way to the descriptions 
of the genera and have been constructed on the assump- 
tion that they must work for all normal specimens. 
I hope so, for a key that will unlock a door only 75 
(or even 90) percent of the time is a tool of little worth. 
Here, a word of caution. No possible key will work 
for abnormal specimens. The worker in Phycitidae 
must be always on the alert for them, for the family 
contains an unusual number of freaks (chiefly vena- 
tional abnormalities). Any one wishing to identify 
phycitids must resign himself to the tedium of dissection 
and slide making. Here, as in all the serious business of 
science, there is no easy way, no short cut to knowledge. 

The groupings of genera and species, prefaced by 
brief summaries of their common characters, which I 
have interposed within the text, are intended only to 
assist the reader and are not.to be understood, in any 
sense, as definitions of taxonomic (tribal or subgeneric) 
groups. In a few instances they may be; but they are, 
in intent, only divisions of convenience. 

The only portion of the keys offered as a description 
of taxonomic units is that separating the subfamilies 
Anerastiinae and Phycitinae. This long-established 
division of the family Phycitidae seems to be a sound 
one, and in the main the subfamilies themselves appear 
to be natural entities, although their definition leaves 
much to be desired. Probably when the Anerastiinae 
are thoroughly studied we may find other features more 
constant than the reduced and concealed tongue. 
There may even be some shifting of genera across the 
subfamily lines. However, this is only hopeful antici- 
pation for the future and will remain so until the Old 
World genera and species of the family are thoroughly 
revised. For the present we shall have to content 
ourselves with an imperfect definition. 

The chart opposite this page shows my interpretation 
of the genera in their relation to each other and to the 
system based on venational characters. 


INTRODUCTION 


GENERA OF AMERICAN PHYCITINAE GROUPED ACCORDING TO GENITALIA AND VENATION 


Relationships on genitalic characters shown in horizontal arrangement. Venational groupings are vertical 


Group 11 - H. W. vein 4s absent | Group 11 


Acrobasis 
Rhodophaea 


Wanita cere Prosoeuzophera 


Promylea 
Anadelosemia 
Dasypyga 
(Rampylta 2 ) 


Monopt! lota 


Caristanius 


Patriciola 
Tulsa 
Telethusia Psorosina 


Staudingeria 
Heterographis| 


Adelperga Stylobasis 
Protasia Oedothmia 
Valdivia 
}Ocala 

Macrorrhinia 

Eumysia Divitiaca 


VIII INTRODUCTION 


The sequence of genera and of species within the 
genus here offered isan attempt to bring together in linear 
arrangement the forms showing affinities in structure 
and development. It is only an attempt and I shall 
quarrel with no one who objects to it. 

When I began this study I had hoped to write a 
monographic treatise and explore the phylogeny of the 
family, but I now find that I know so much less than I 
thought I did and that the accumulated knowledge of 


others is so meager that any attempt along these lines 
would be & vain and futile performance. 

We don’t know what a primitive phycitid was like. 
We don’t know which forms evolved from which, or 
how. We weren’t there. We may surmise; but the 
guess of one ignoramus is as good as that of another, and 
there is nothing to be gained from either. I have had 
to be content with a mere revision. Would that it were 
more worthy. 

Cari HEINRICH 


Carl Heinrich (1880-1955) 


This monograph was written by Carl Heinrich in the months following his retirement from 
Government service in 1949. Upon it he focus>! the extensive knowledge gained during his 36 
years as entomologist with the U. S. Department of Agriculture. Its publication, toward which 
the Department of Agriculture has contributed substantially, was undertaken in 1954, and the 
author had completed his review of the galley proofs at the time of his death, age 75, on May 31, 


1955. 


A biographical memoir of Car] Heinrich and a bibliography of his scientific writings appeared 
in the Proceedings of the Entomological Society of Washington for October 1955 (vol. 57, No. 5, 
pp- 249-255). In addition to the present bulletin, the U. S. National Museum has published a 
number of his papers, of which several, as noted, are now out of print: 


1921. On some forest Lepidoptera with descrip- 
tions of new species, larvae, and pupae. 
Proc. U. S. Nat. Mus. No. 2305, vol. 57, 
pp- 53-96, 13 pls., June 17, 1920. (Out 
of print.) 

1923. Revision of the North American moths of 
the subfamily Eucosminae of the family 
Olethreutidae. U. S. Nat. Mus. Bull. 
123, iv-+298 pp., 1 fig., 59 pls., Apr. 12, 
1923. (Out of print.) 

1926. Revision of the North American moths of 
the subfamilies Laspeyresiinae and Ole- 
threutinae. U.S. Nat. Mus. Bull. 132, 
iv-+216 pp., 2 figs., 76 pls., Feb. 2, 1926. 
(Out of print.) 

1927. The American moths of the genus Diatraea 
allies. Proc. U. S. Nat. Mus. No. 2691, 
vol. 71, Art. 19, 48, pp., 20 pls., Aug. 23, 
1927. Joint authorship with H. G. Dyar. 
(Out of print.) 


1929. Notes on some North American moths of the 
subfamily Eucosminae. Proc. U.S. Nat. 
Mus. No. 2779, vol. 75, Art. 8, 23 pp., 5 
pls., Apr. 5, 1929. 

1932. Notes on and descriptions of some American 
moths. Proc. U.S. Nat. Mus. No. 2879, 
vol. 79, Art. 13, 16 pp., 1 fig., 7 pls., Aug. 
10, 1931. 

1938. Moths of the genus Rupela (Pyralididae. 


Schoenobiinae). Proc. U. S. Nat. Mus. 
No. 3019, vol. 84, pp. 355-388. 12 pls., 
July 3, 1937. 


1940. The cactus-feeding Phycitinae: A contribu- 
tion toward a revision of the American 
pyralidoid moths of the family Phycitidae. 
Proc. U. S. Nat. Mus. No. 3053, vol. 86, 
pp- 331-413, 29 pls., Mar. 16, 1939. 

1945. The genus Fundella Zeller: A contribution 
toward a revision of the American Pyrali- 
deid moths of the family Phycitidae. 
Proc. U. S. Nat. Mus. No. 3190, vol. 96, 
pp. 105-114. 3 pls., May 18, 1945. 


American Moths of the Subfamily Phycitinae 


Family PayciripaE 


Mors: Labial palpus well developed. Maxillary 
palpus present, variously developed, rarely vestigial. 
Tongue well developed or reduced, rarely absent; when 
distinguishable, basal portion scaled. Forewing entire 
(not divided); 11 veins or less; vein 7 always absent; 8 
and 9 stalked or united; 1c absent (represented only by a 
fold or crease in the wing membrane); no areole. Hind 
wing with 8 veins or less; vein 8 closely approximate or 
contiguous to or anastomosing or completely fused with 
7 beyond cell; 1c always present; a fringe of pecten on 
lower median vein at base; frenulum of female simple 
(a single strong spine). 

Larva: With primary setae only; two setae on 
prespiracular shield of prothorax; setae IV and V ap- 
proximate and under the spiracle on abdominal seg- 
ments 1 to 8; normally a sclerotized, pigmented ring 


1In Cactoblastis the aborted tongue is completely concealed by 
the broadly scaled basal segments of the labial palpi. However, 
the genitalia, habitus and larval affinities show that Cactoblastis 
is ajtrue phycitine and must be placed with the other closely 
related genera of the cactus-feeding Phycitinae. 


encircling or partially encircling the tubercle of seta 
IIb on mesothorax and a smilar ring encircling the 
tubercle of seta IIT on eighth abdominal segment (this 
character absent from the following genera of the 
American Phycitinae; Htiella, Oryctometopia, Ulophora, 
Rotruda, Rhagea, and Unadilia). Prolegs normal; 
crochets in a complete circle. 

The subfamilies of Phycitidae are separated by the 
following key: 


Tongue normally well developed; if sometimes reduced, 
not concealed between the labial palpi (except in 
Cactoblastis ); ocelli always present . . Phycitinae 


Tongue reduced or vestigial; when merely reduced, 
concealed between the labial palpi; if sometimes 
exposed between the palpi (Bandera), then ocelli 
absent Anerastiinae 


Subfamily Phycitinae 


The larger groups of the subfamily Phycitinae are 
separated by the following key: 


Key to the larger groups of Phycitinae 


Hind wing with veins 3 and 4 both present . 
Hind wing with vein 3 present, 4 absent . . 
Hind wing with veins 3 and 4 both absent . 


Group I 


[Hind wing with veins 3 and 4 both present] 


Keys To THE VENATIONAL Divisions AND GENERA OF Group I 


Hind wing with vein 3 appreciably before the outer angle of cell; cell less than one-half the 


wing length 


BEVUTR UNTHRL HIS Venational division A 


Cryptoblabes (p. 10) 


Hind wing with vein 3 closely approximate to or from the angle of the cell (rarely shortly 
stalked with 4-5); 7 and 8 approximate, contiguous, or shortly and weakly anastomosed 
beyond cell; cell at lower angle nearly half as long as wing; if shorter, then vein 3 of 
moderate length and the free (divergent) part of 3 decidedly shorter than vein 2. If 
vein 3 sometimes appreciably before outer angle of cell (Acrobasis), then cell one-half 


threrwanpylencthy.) (eye ey A ee sss 


Big re neha Venational division B (key, p. 2) 


Hind wing with vein 3 closely approximate to or from the angle of cell; veins 7-8 solidly 
anastomosed beyond cell for at least three-fourths of their lengths. 


300329—56——2 


Venational division C (key, p. 7) 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Hind wing with vein 3 closely approximate to or from the angle of cell; veins 7-8 approxi- 


12. 


13. 


tion. 


mate or very weakly and shortly anastomosed beyond cell; 2 and 3 both long, the 
divergent element of 3 nearly as long as 2; cell distinctly ie than half the length of 
WI cr Peas atau ry inl: Ararat Sea rea eee te (tse ee a . Venational division D (key, p. 8) 


Venational division B 


. Hind wing with discocellular vein oblique. ...... Sa Coc taben net: Re Wie WLI Sik Yer 2 


Hind wing with discocellular vein curved . . 3 


. Forewing with subbasal ridge of raised scales; ed ae sii cell someinat ils dhe 


one-third the length of the wing; eighth abdominal segment of male with midven- 

tral hair tuftyy) «essde satomredo. - dehsReey vita. bedokyv: Mildrixia (p. 26) 

Forewing smooth; hind wing with cell one-fifth the length of the wing; eighth abdominal 
segment of male with sternite developed as a sclerotized, digitate pocket. 

Drescoma (p. 88) 


. Hind wing with vein 3 from before, but near, lower outer angle of cell. . . . . . . 4 


Hind wing with vein 3 from the angle or the stalk of veins 4-5. . ........ 8 


. Hind wing with veins 4-5 connate; vein 6 of forewing always straight. ...... 5 


Hind wing with veins 4-5 approximate, contiguous, anastomosed or stalked for a short 
distance from cell, if sometimes connate (on individuals of Hypsipyla) vein 6 of fore- 
wing slightly bent towards base ........-..2.... Satis) ih ee eee eR 


. Basal segment of male antenna triangulate. ........... Acrobasis (p. 11) 


Basal segment of male antenna simple (cylindrical): ...... Rhodophaea? (p. 24) 


. Forewing with subbasal ridge of raised scales; antenna of male pubescent (cilia distinctly 


shorter than width of shaft) .............4... Anabasis (p. 25) 
Forewing smooth; antenna of male ciliate (cilia distinctly longer than width of shaft) . 7 


. Forewing with vein 6 straight, remote from veins 8-9 at base . . Sematoneura (p. 27) 


Forewing with vein 6 bent, shortly separated from 8-9 at base . . . Hypsipyla (p. 27) 


. Vestiture of head, thorax, labial palpi, and femora a mixture of scales and hairs; male with 


harpe short (stubby), clasper absent, apical process of gnathos an inverted heart- 
shaped lobe with short, slender spine; female with ventral surface of genitalia sclero- 
tized throughout its length, ductus seminalis from near junction of bursa and ductus 
DUESHED te tects eee ron yn icc ie Aa an |, Nena Beep On ane . Polopeustis (p. 120) 
Vestiture entirely of scales; or; if occasionally mixed with hair (Saraéa airella), male with 
harpe elongate, clasper present, knoblike, apical process an elongate, stout hook; 
female with ductus bursae unsclerotized, ductus seminalis from bursa remote from 
junction of bursa and ductusbursae .........2.2.+.-.-++-+++-+-+.-. 9 


. Forewing with subbasal ridge ofraisedscales . ........-.4...- spay yl 


Foréwimgismootll iis) 2 be ey GN Eee ala a INT URS Saleh ee a 16 


. Hind wing with vein 3 from the stalk of vems4—5 =... ..... Passadena (p. 175) 


Hind wing with vein 3 approximate to but not from the stalk of vems 4-5 ..... 11 


. Labial palpus porrect, beaklike . . ..........++e+-- . KEtiella (p. 98) 


Labial palpus obliqueorupturned . . 2. 2... ee ee ee 12 
Forewing with vein 6 from the upper angle of the cell, bent towardsbase. ........ 
Hypargyria (p. ee 
Forewing with vein 6 from well below the angle of cell, straight. . . .. . ay scree 
Forewing with veins 4 and 5 slightly separated at base and approximate for a short ste 
tance beyond 2 te cnndw aa wool ae-tad vitor eface eel deeiias o Paar beeoms 14 
Forewing with veins 4 and 5 connate or stalked; if sometimes shortly separated at base 
notapproximate beyond. ...... 2. ee eee ee es 15 


2 The genus Trachycera is omitted from our key as the male is unknown except from Ragonot’s descrip- 
The female before me has the venation of Rhodophaea. It is distinguished from other females of 


Division B in haying two scobinate, cuplike signa similar to those of Davara in division D. (See p. 25.)]+ 


14, 


15. 


16. 


17. 


18. 


19. 


20. 


21. 


22. 


23. 


24, 


25. 


26. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Hind wing with veins 7-8 anastomosed beyond cell; maxillary palpus of male in the form 
oftaneierette...... OR CLIk Were eReIUG aioth dir olan Immyrla (p. 111) 
Hind wing with veins 7-8 approximate beyond cell; maxillary palpus of male squamous. 
Ulophora (p. 176) 
Labial palpus oblique, second segment on male grooved; male with eighth abdominal 
segment simple; female with genitalopeningsimple .... . Ortholepis (p. 119) 
Labial palpus upturned, second segment on male not grooved; eighth abdominal segment 
of male with compound tufts; female with strongly sclerotized plate at genital open- 


ing attached to supplemental eighth-segment collar ...... Tlascala (p. 133) 
Labial palpus porrect, beaklike; male with aedeagus flanged and strongly spined; female 
with ductus seminalis-from bursa .....-.+...-.» Polliyom phonyage ditore |) ily 


Labial palpus oblique or upturned; if sometimes appearing porrect (Stylopalpia) due to 
long, deflected third segment, male with aedeagus simple; female with ductus semi- 


nahstirom ductus burske)) .Joge 2 Jol Seyieqg oy enigy aliwy yogis |. 21 
Forewing with veins 4-5 approximate for a short distance from cell. . . . 2... 18 
Forewing with veins4—5 stalked. . 2. 2 ek ee, 19 
Labial palpi extending at least twice the length of head beyond it; harpe of male elongate; 

female with ductus bursae much longer than bursa ........ Pima (p. 101) 
Labial palpi extending little more than the length of head beyond it; harpe of male short; 

female with ductus bursae little, if any, longer than bursa . . Interjectio (p. 106) 


Male with second segment of labial palpus grooved on inner side; female with ductus 
seminalis from bursa remote from junction of bursa and ductus bursae . Sarata (p. 159) 
Male with second segment of labial palpus not grooved; female with ductus seminalis 
from bursa near junction of bursaand ductus bursae . ............ 20 
Female with a strongly sclerotized plate behind genital opening . . Philodemia (p. 165) 
Female without sclerotization at genital opening ........ Lipographis (p. 166) 
Hind wing with vein 3 from the stalk of 4-5 or closely approximate to it for some distance 
from lower outer angle of cell; vein 2 alwaysratherneartheangle. ....... 22 
Hind wing with vein 3 connate with the stalk of veins 4—5 or connected with it at base by 
a very short spur; if sometimes approximate to the stalk of 4-5 (Megarthria, Acron- 
cosa), vein 2 always from well before lower outer angle of cell; or, if vein 3 some- 
times shortly fused with the stalk of veins 4-5 (Actriz), male with apical process of 
gnathos developed as a square or inverted heart-shaped plate and female with 
caudal half of bursa copulatrix densely spinose. . . ..... 2... ee 28 
Hind wing of male with anal angle folded into a pocket; female with strong sclerotizations 
behind or surrounding genital opening of genitalia... .........4.. 23 
Hind wing of male without folded pocket at anal angle; genital opening of female 
simple. ico, soley Sodan bestjowloe goods Miw has gerd er papi: 24 
Male genitalia with uncus hammer-clawed (long, curved, constricted at middle and 
broadly divided at apex); female with bursa containing strongly sclerotized folds 
or stoutly spmedibandstsimer lagunohds sigan, .outd boozad, Fundella (p. 59) 
Male genitalia with uncus otherwise (sometimes broadly divided at apex but the divided 
elements small and spinelike and the middle of uncus not appreciably constricted) ; 
bursa sometimes with a small granulate patch but otherwise membranous and 


RUDAPITOG 5 hiss econ Shwe Ni Ay cisync socse Ae Mates es © OMQUNse SOONIOOR: Difundella (p. 62) 
Hind wing with vein 3 approximate to the stalk of veins 4—5 for some distance from outer 
pngle‘obealliisls et SvSiibogl SURRY Gy ugnooyiay Starts Wig ¢ Scorylus (p. 72) 
Hind wing with vein 3 from the stalk of veins 4-5... 2... ee ee 25 


Male genitalia with transtilla a sinuate, sclerotized scobinate band involved with gnathos 
and with a long free spine involved with anellus; female with ductus seminalis from 


ductusbirsaesioielor . .sialqmigs alivaayny. ine egdyq leds tig? gauge ig Bia, 26 
Male genitalia with transtilla incomplete or absent; female with ductus seminalis from 
bursa l! ...Ueo brayed aanntelh taods ww busompienin 2S ete? tipw wii DM 27 
Maxillary palpus of male filiform. . ........2.2.200.. Coptarthria (p. 64) 


Maxillary palpus of malesquamous. ............. Anadelosemia (p. 67) 


27. 


28. 


29. 


30. 


31. 


32. 


33. 


34, 


35. 


36. 


37. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Male genitalia with divided element of incomplete transtilla strongly sclerotized, gnathos 
absent; female with ductus bursae partially sclerotized .... . Gabinius (p. 84) 
Male genitalia with transtilla absent, gnathos well developed and with apical process 
a strong hook; female with ductus bursae membranous throughout. 
Ceracanthia (p. 85) 
Male genitalia with transtilla a sinuate sclerotized band involved with gnathos; female 
with bursa small, membranous and ductus bursae much longer than bursa, signum 
(if present) a small granulate patch or small plate with single minute thorn, genital 
opening narrow (the ductus bursae never expanded into a widened opening) ; if bursa 
sometimes large and ductus bursae proportionally shorter (Rampylla), collar of 
eighth :sepment modified. \y.%5)..\)5))9 hea orleans Bee eben ae 29 
Male genitalia without transtilla or, if present, otherwise; female genitalia never as 
aboyeuin-all details} ents sanehaaulie shee heal: Duke betes Bas pees 31 
Forewing with veims 4—5 approximate for a short distance from cell; hind wing with 
vein 2 from well before lower outer angle of cell. . . . .......2...-. 30 
Forewing with veins 4-5 stalked; hind wing with vein 2 from near outer angle of cell. 
Dasypyga (p. 69) 
Female with ductus seminalis from ductus bursae. ........ Promylea (p. 65) 
Female with ductus seminalis from bursa copulatrix ........ Rampylla (p. 70) 
(males: venation group D, couplet 8) 
Male with transtilla of genitalia complete and strongly sclerotized; or, if incomplete, the 
elements enlarged, strongly sclerotized and modified; when complete not in the 
form of a squarish plate. Female with a single signum developed as a small, scobin- 
ate or granulate cup or patch or (Adanarsa) as a single short, stout thorn; genital 
openingvelways broady sty.) sd}isoss heilidoe alata aaatieise vimaciie et Ss cae 32 
Male genitalia with transtilla incomplete or absent. Female with signa or signum, if 
present, otherwise developed. If transtilla complete then weakly sclerotized or 
developed as a square plate; and if signum of female a small scobinate patch (Zegar- 


thria alpha) genital opening narrow... .......2..+.+-+.+.4+2-s 39 
Forewing with vein 6 bent towards base. . ......-....-.+-++-+--- 33 
Forewing with vein 6 straight. 2 2.0.0. 2 ee ee 36 
Transtilla of male complete, strongly arched and with median areaforked ..... . 34 
Transtilla of male complete, but not arched nor with median area forked. .... . 35 


Antenna of male with shaft unipectinate. Ductus bursae of female very short, less than 
one-half as long as bursa, scobinate-granulate and more or less sclerotized but not 
transversely wrinkled ............2..26: Hemiptilocera (p. 30) 

Antenna of male with shaft pubescent. Ductus bursae of female longer, about half as 
long as bursa and with strong, sclerotized wrinkling before genital opening. 

Crocidomera (p.32) 

Maxillary palpus squamous. Forewing of male with costal fold and a fovea in cell 
slightly beyond base. Highth abdominal segment with sternite developed as a 
sclerotized pocket. . ...........24.4- ONG tits mt Heras (p.34) 

(male only, female unknown) 

Maxillary palpus filiform. Forewing of male without costal fold or fovea. Eighth 

abdominal segment simple... ........ 52 ee eee Birinus (p. 36) 
(male only, female unknown) 

Male antenna with shaft unipectinate; transtilla incomplete, its elements long, stout with 
their apices broadly and irregularly developed and hooked. Female with a large 
semicircular sclerotized and scobinate plate on membrane behind genital opening. 

Bertelia (p. 36) 

Male antenna with shaft pubescent; transtilla complete. Sclerotized plate on membrane 
behind genital opening of female, if present, not semicircular. . ....... 37 

Hind wing with veins 7-8 anastomosed for a short distance beyond cell. Male with trans- 
tilla strongly arced but with its median area bearing a smooth narrow crosspiece, not 
forked. Female with signum a single, short, stout, hooked thorn. Adanarsa (p. 35) 


38. 


39. 


40. 


41. 


42, 


43. 


44, 


45. 


46. 


47. 


48. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Hind wing with veins 7-8 approximate for a short distance from cell. Male transtilla 
otherwise. Female with signum a single, small, scobinate or granulate, cup-shaped 


patcliys, heels ier a sued ET “Qpdzly “Pl, Si 38 
Male with apical process of gnathos a stout houk. Fennle with ductus burdie appreci- 
ably longer than bursa; ductus seminalis from ductus bursae . . Cuniberta (p. 34) 


Male without projecting apical process, the lateral arms supporting a thinly sclerotized 
subanal plate. Female with ductus bursae much shorter than bursa; ductus semi- 
nalis from lobe of bursa near junction of bursa and ductus bursae. Chararica (p. 38) 

Male with sternite of eighth abdominal segment developed as a sclerotized pocket. 
Female with signum a small, depressed, granulate-scobinate patch; or, if signum 
absent, ductus bursae narrow throughout and sclerotized for about one-third of its 
length from shortly beyond its junction with bursa, the sclerotized portion sharply 
bentigveriteok.. - .. mous Croom Ha, viedioie, rut, Megarthria (p. 86) 

Male with eighth abdominal ebeniaite ib developed as a sclerotized pocket. Female 
with signum (or signa) developed as sclerotized and strongly spined plates or bands, 
or entirely absent. If signa absent, ductus bursae not as above. If sometimes 
(Olybria) a single more or less strongly sclerotized band in bursa at junction of bursa 
and ductus bursae, the band finely serrate or edged with short spines . . . . .40 

Antenna of male with shaft unipectinate. Female bursa with a single strongly spined 
sipnammosele jon elagiquos asiouh Bogatvig adalat pagoteq Monoptilota (p. 89) 

Antenna of male with shaft pubescent. Female bursa with two signa or none . . .41 

Male genitalia with a pair of long, strong, sclerotized arms from the ventrolateral angles 
of uncus; gnathos absent. Female with two signa consisting of stoutly and coarsely 
spined bands: ductus bursae strongly sclerotized, fattened (ribbonlike). 

Caristanius (p. 97) 

Male genitalia with uncus otherwise; gnathos present and well developed. Female with- 
out signa; or, when present, consisting of two strongly spined bands or plates; when 
signa are present, ductus bursae not as above, if partially flattened and ribbonlike 
thenhyerymarrowe “Ute wtiibe, viitigdel Dodlay®, oc, T-h elev AUPE Qniwe . .42 

Male genitalia with aedeagus expanding to lateral, flanged projections before apex, the 
flanges each bearing a cluster of strong spines. Female with ductus seminalis from 


ducinsibursde: sTaheel Ha Base soit inuiels Vibe ais Stylopalpia (p. 140) 
Male genitalia with aedeagus otherwise, if sometimes spined (Pyla), not flanged before 
apex. Female with ductus seminalis from bursa copulatrix ......... 43 
Masallany palpus ‘vestigial .. °°.0". Gaertn ond) oho. PG Wee Ophah URE tyne 44 


Maxillary palpus squamous, or filiform, or (on some males) in the form of an aigrette . 45 
Penis of male unarmed except for a eran cluster of very weak, short, slender spines. 


Female with ovipositor strongly sclerotized. . . ..... ‘ Télethiusia (p. 186) 
Penis of male armed with a single, strong cornutus. Female with ovipositor normal (not 
sirosply ‘scleretized)d ier pets Sew Se Saskies sured al. Phobus (p. 138) 
Forewing with some rough (raised) scaling in median area beyond outer border of ante- 
median line, but without subbasal ridge of raised scales . . . . . Tulsa (p. 134) 
Forewing Sfadath Yr eet om oee ated CUCSTLONIO COTY Ah pis Ae, TAG 
Fore tibia with a long inner and a short outer claw ....... Nleei cask (p. 174) 
Honatibin atherwasexisoiu, res eit, TRADE SEY Ailes Ws, WAT 


Hind wing with veins 7-8 distinctly anastomosed for about half hein ddngths (more than 
the free length of vein 8). Male with apical process of gnathos a broad shield 
without central terminal spine .........2.2.. oy iter PAGtrix, (ps 139) 

Hind wing with veins 7 and 8 approximate; if sometimes contiguous or anastomosed be- 
yond cell, then weakly and very shortly so. Male with apical process of gnathos 
otherwise al Sjad eased holla adchanibesritos <ihisciphagg CIN. LECENER CIOL, OFT... 48 

Male genitalia with a pair of straight, strongly sclerotized arms [not to be confused with 
similar projections from vinculum in Nephopteryz crassifasciella] projecting back- 
ward from lower, posterior angles of tegumen. Female with ductus bursae of gen- 
italia flattened (cibboniike), waved (twice bent) and sclerotized throughout, the 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


sclerotization developed at genital opening into a stout, squarish ventral 


platejalycog-to.ntaniloon Sluis heals ge Beenie eM okepeas, ae Olybria (p. 113) 
Male genitalia without projecting arms from tegumen. Ductus bursae of female 
otherwisejs:'. dois’ elasne tl. alood tote 3 ROR, PY Ree OSG. Wield olely 49 


49. Harpe of male genitalia with a long hair brush from inner surface, along lower edge of 
basal half of sclerotized costa. Bursa copulatrix of female with the lobe giving off 
ductus seminalis strongly sclerotized; or most of dorsal area of bursa strongly and 
smoothly sclerotized; or bursa with conspicuous, round or oval, strongly pigmented 
and sclerotized, densely granulate patches [absent in Nephopteryx subcaesiella (fic. 
826)], the bursa otherwise spinose overitsmembranous areas. ........ 50 
Harpe of male without such hair brush. Bursa of female not as above ..... 53 
50. Male with sacculus of harpe strongly produced at apex. Female with most of dorsal 
surface of bursa strongly and smoothly sclerotized..... . Glyptocera (p. 100) 
Male with sacculus of harpe simple (not produced at apex). Female with only the lobe 
giving off ductus seminalis sclerotized: or bursa with granulate sclerotized 
patchesgvicais aa sou oneal aiogh Soesitn eee Es aisacte glares Ge 51 

51, Penis of male armed with numerous strong, slender spines.. Female with lobe of bursa 

giving off ductus seminalis strongly sclerotized. ......... Oreana (p. 112) 
Penis of male armed with two stout spines or (very rarely) one spine. Bursa of female 
with granulate patches, the lobe giving off ductus seminalis not sclerotized. . . 52 

52. Harpe of male genitalia with an appressed, stout, thorny or serrate clasper. Female 
with ductus bursae unsclerotized adjacent to bursa, but with strong sclerotization 
dti-cenital openings; izcas via cat iy esi. takede pong Meroptera (p. 121) 

Harpe of male with clasper digitate, slender, simple (without spining). Female with 
ductus bursae sclerotized along ventral surface from junction with bursa, the sclero- 
tization terminating before genital opening, the latter simple (unsclerotized). 

Nephopteryx (p. 123) 


53. Forewing with veins 4—5 stalked for nearly half their lengths. . . . . Tacoma (p. 178) 
Forewing with veins 4-5 not stalked (slightly separated or closely approximate at 
cell) tax neciios ier berated olemiol of palomneme eoasbes Aiiiy sileisivs.. . . 54 


54. Male genitalia with transtilla complete but very weakly sclerotized (a narrow angulate 
band); aedeagus slender, elongate, not spined or divided. Female genitalia with 
bursa entirely membranous and smooth; ductus bursae rather narrow, tubular and 
sclerotized. throughout, nowhere appreciably widened; genital opening narrow (no 
wider than narrowest part of ductus bursae)........... Tota (p. 170) 

Male genitalia with transtilla incomplete or absent; aedeagus moderately broad to stout, 
if sometimes slender and elongate then spined or partially divided or (Chorrera) 
vinculum with produced lateral lobes from terminal margin. Female with or with- 
out signa in bursa; if without signa, bursa spinose or with same sclerotizations 
adjacent to ductus bursae; if bursa entirely membranous and smooth, genital 
opening decidedly broadened . . . .. 2... 2... ee ee ee ee ts 55 

55. Male with aedeagus of genitalia spined or partially divided; if sometimes simple, then a 
strong knoblike and spinose projection from harpe at base of costa. Female without 
signa in bursa, the latter usually simple (unsclerotized and smooth), but sometimes 
with sclerotized, convolute, longitudinal bands near junction of bursa and extending 
into the ductus, the bursa never spinose . .......-..... Pyla (p. 142) 

Male with aedeagus neither spined nor divided; if harpe sometimes with projection from 
base of costa, the latter neither knoblike nor spinose. Female with or without signa 
in bursa; if without signa, the bursa spinose (at lease partially so) or sclerotized at 
junction of bursa and ductus bursae, but such sclerotization not in the form of 
econvolute, Joneitudinal bands, 2. 2. ee ee ww ee ee OU , 38 

56. Male with maxillary palpus squamous or filiform. Female without signa; a serrate or 
minutely spined sclerotization at junction of bursa and ductus bursae, the latter 
broad and strongly sclerotized or with a pair of elongate sclerotized plates behind 


57. 


58. 


59. 


60. 


61 


62. 


63. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


genital opening; if signa present, the ductus bursae very narrow and expanded 


into a membranous globe shortly before genital opening. . ......... 57 
Male with maxillary palpus in the form of an aigrette. Female with or without signa; 
bursa and ductus bursae not as above. . . 2... 6 ee ee ee 59 
Male genitalia with aedeagus stout; penis armed with a single, long stout spine. Bursa 
of female geriitaliawwithoutisignan oli i! soot lS ance sNodle lag spelled 58 


Male genitalia with aedeagus very slender; penis unarmed. Bursa of female containing 
two signa developed as opposed, longitudinal bands bearing a row of short stout 


EPINES..;\. (sg cs le elt, aetdete bis (busolo).roits Chorrera (p. 177) 
Maxillary palpus of male minute, filiform. Female without sclerotized plate or plates 
behind «genitaliopaning ay. js cobindin, newt aecaud tical elenies Ambesa (p. 108) 


Maxillary palpus of male squamous (broadly scaled). Female with a pair of elongate 
sclerotized plates on inner dorsal surface of ductus bursae behind genital 
openinewamayyis eesdst. leokje (sal .oiolayanimi-cam 20 obs Catastia (p. 110) 

Male with sacculus of harpe slightly produced at apex. Female with ductus bursae of 
genitalia cylindrical; narrow except at genital opening; appreciably longer than 
length of bursa; sclerotized for half its length from junction with bursa, the sclero- 
tization longitudinally ribbed ............. Elasmopalpus (p. 172) 

Male with sacculus of harpe not produced at apex. Ductus bursae of female other- 
Wiseweneti | .wanowiie idente! divtaiiniivads yore doisdiiocule laches sti tis . 60 

Hind wing with vein 2 from before but rather near outer angle of cell Atiw qwatcucee 61 

Hind wing with vein 2 from well before outer angle ofcell . . . 1... .... 62 

Male with penis armed with a single stout spine. Female with ductus brusae of genitalia 
sclerotized (at least towards genital opening). Forewing with veins 8-9 stalked for 
less than two-thirds of their lengths. Hind wing with veins 4-5 stalked for approxi- 
mately one-half their lengths. ... 2... 2... ee ae Salebriaria (p. 115) 

Male with penis unarmed. Female with ductus bursae cartilagenous, except at its 
junction with bursa. Forewing with veins 8-9 stalked for over two-thirds of their 
lengths. Hind wing with veins 4-5 stalked for over three-fourths of their 
lengths: Sein Gets. below orneth Insdusdie¥ a ctiie se - Quasisalebria (p. 118) 

Male genitalia with a strong, straight or curved arm from base of costa of harpe; penis 
unarmed. Female genitalia with ductus seminalis from bursa adjacent to junction 
of bursa and ductus bursae (signa present). ......... Adelphia (p. 168) 

Male genitalia without projecting arm from base of costa of harpe; penis armed with a 
single strong spine. Female with ductus seminalis from bursa remote from junction 
of bursa and ductus bursae (signa present or absent). .... Le ay S463 

Male with cornutus on penis a long straight spine, over half as long as wed Garuke Female 
with bursa densely spinose, signa absent; ductus seminalis from near anterior end of 
bursa “sah: diate weeds Cine easier i sieeve leeeapetan eit Salebriacus (p. 114) 

Male with cornutus on penis a short, curved spine, somewhat less than one-third as long 
as aedeagus. Female with bursa smooth except for strongly spined signa (and in 
one species a strongly spined collar at middle of bursa) ; ductus seminalis from bursa 
Sa juceninboronevOh tie Siena Skee roe ele ene alee tantel esis tess Ufa (p. 170) 


Venational division C 


. Hind wing with discocellular vein straight, vertical. . . . . . . Homoeographa (p. 135) 
Hind wing with discocellular vein straight, oblique. ......... Atheloca (p. 81) 
Hind wing with discocellular vein fainted O.Otw. stand . Srad otalngar ori. ... 2 

. Male with uncus of genitalia more or less spoon-shaped (the lateral margins deeply 


concave at middle). Female with signa present, consisting of 2 or more sclerotized 
disks or series of contiguous, bluntthorns. . . . 1... 2... 1 ee ee eee 3 
Male with uncus triangulate or subtriangulate. Female with or without signa; if present 
MOT ASIADOVOSs : RRR renee is cS AT, GI Nye boos se opie caer catteb ett 3 4 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


. Male with costal fold on forewing; shaft of antenna notched at base . Diatomocera (p. 50) 
Male without costal fold on forewing; antennal shaft not notched . Pseudocabima (p. 53) 

. Hind wing with vein 3 from the stalk of veins 4-5. ........ Anypsipyla (p. 42) 
Hind wing with vein 3 from the angle of the cell; if sometimes approximate to the stalk 
of veins 4—5 for a short distance, never actually fromit. ........ 2a NE XS 

. Maxillary palpi of both sexes filiform. Male with complete transtilla ....... 6 
Maxillary palpi squamous. Transtilla of maleimcomplete ........2.... 8 

. Male with antennal shaft pubescent. Female with ductus seminalis from near middle or 
towards anterior (closed) end of bursa .......2.2.2.2.2.2.4.22... : 7 


Male with antennal shaft shortly ciliate (cilia as long as width of shaft). Female with 
ductus seminalis from bursa near junction of bursa and ductus bursae. 

Kcetomyelois (p. 43) 

. Forewing with vein 2 from near outer angle of cell. Male with transtilla weakly sclerotized 

(a thin band or sub-triangulate plate); apical process of gnathos broadly U- 

shapedih sabivaiaen ts .. aeeiey tithe Devs Decrees enter shah: etd Tay: ret ge Myelopsis (p. 40) 

Forewing with vein 2 from well before the angle. Male with transtilla strongly sclero- 

tized and arched; apical process of gnathos a simple, stout hook . . Apomyelois (p. 42) 

. Hind wing with cell moderately long (from a little over to slightly less than one-half the 

length ciitheiswing)iingh) .. 2a. dk Roa bela aaa do. eoluad tin, eels. 9 

Hind wing with cell short (not over one-third the length of wing). . Protomoerbes (p. 49) 

. Forewing with vein 2 from near lower outer angle of cell; vein 10 from the cell. Male 

with eighth abdominal segment simple. Female with signum in bursa (a cluster of 

coarse scobinations); a sclerotized plate behind genital opening (on inner dorsal 

surface of ductus bursae). ............2... Paramyelois (p. 46) 

Forewing with vein 2 from well before the angle; vein 10 from the stalk of veins 8-9. 

Male with a pair of ventrolateral hair tufts on eighth abdominal segment. Female 

with bursa and ductus bursae simple (membranous throughout, with neither signum 

in bursa nor plate behind genital opening) ......... Pseudodivona (p. 48) 


Venational division D 


1. Normal dark discal spots on forewing at end of cell replaced by a conspicuous white spot 
or line (obscured only on clarioralis) on discocellular vein. [Male genitalia with 
eostal area of harpe broadly sclerotized and produced at apex; clasper present, 
erect; penis armed with numerous, straight spines. Female with signa developed as 
2 or 3 clusters of strong, slender spines; ductus bursae flattened, strongly sclerotized 
over most of its length, the sclerotization terminating just before simple genital 


roy OLSSON kes el eal lA be SiN all A, ian) adh Seam ati eh on Saat 78 Dioryctria (p. 149) 
No such white spot on discocellular vein. . .......2.2.+..4.+..4.-. sae 
2. Forewing with ridge of raised scales preceding antemedial line .......... 3 


Forewing smooth; if sometimes with a few roughened scales, no such subbasal ridge. . 4 

3. Forewing with veins 4-5 closely approximate for a short distance from cell. Hind 
wing with cell less than one-fourth the length of wing. Maxillary palpus of male 

in) the form\ofjansierette)) 278. fine soe 4 6 jue ee es Zamagiria (p. 90) 
Forewing with veins 4-5 connate or very shortly stalked. Hind wing with cell slightly 
less than one-third the length of wing. Maxillary palpus of male subsquamous. 
Anegcephalesis (p. 93) 

4. Male genitalia with uncus and tegumen greatly reduced; uncus a narrow, weakly sclero- 
tized, angulate band. Female with a single signum in bursa, consisting of a small, 
sclerotized plate supporting a very short thornlike spine. . ... . Peadus (p. 83) 

Male genitalia with uncus and tegumen well developed. Female with signum or signa 
Gf present) otherwise’. 6... “eesti! corace ot) Ween tal BS 5 


10. 


11. 


12. 


13. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


. Male genitalia with transtilla complete, developed as a narrow, slightly arched band, 


attached to harpes only by membrane. Female with signum a cluster of bluntly 
pointed thorns, more or less surrounded by fine scobinations or strongly pigmented 


Eraniuations s °S SOOM RN, STR Aibe artiunind Hyalospila (p. 56) 
Male genitalia incomplete or absent; if complete (Magiriopsis) not a narrow band. 
Female with signum or signa (if present) otherwise. .......2.2.... 6 

. Hind wing with discocellular vein incomplete ........2... Fulrada (p. 71) 


(based on male; female unknown) 
Hind wing with discocellular vein complete. . . . 2... .... 2 ee ee ee 7 


. Hind wing with cell very short (about one-fifth the length of wing). Male genitalia 


with lateral arms of gnathos broad, expanded and curled at their extremities. 
Praedonula (p. 82) 
Hind wing with cell longer (from one-fourth to one-third the length of wing). Gnathos 
oftmale! otlierwiselt! Hrw. OL 20 8 clay ve heal cn ny alts 8 


. Hind wing of male with anal area (involving vein la) thicker? aad folded, Genie 


a produced pocket; underside of wing with roughened scale or hair tufts on some 


OLGHELVeMIS hr tii TS 4 ca pica, oc. « ACORN EheaILD, Abewes Rampylla (p. 70) 
(females: Venation Group B, couplet 30) 
Hind wing of male without such modification. . ......... IHS BOG 


. Male with uncus of genitalia bifid. Female with one or two small signa developed 


as granulate depressions in bursa; if signa sometimes absent (some species of 
Piegnoppeo) puts: Melnbrapousy . . . 0... yes my ee ee ee 10 
Uncus of male undivided (triangulate or benanonen iaene of rorinle developed as 
strongly, spined bands or plates; if sometimes absent (Ancylostomia), bursa weakly 


but, extensively sclenotizedy). QING? .. me actran-» ‘agasne ee taut repmatend oslanns 12 
Gnathos complete, a thin, weakly sclerotized, transverse band. Female with two 
SISTEM DUIS ee. ek ye te et a DONG AMAe sian orld jeeps MPa ded Ye bn Spy. elih 
Gnathos incomplete, the lateral arms strong, broad; between their separated apices a 
well sclerotized anal plate. Female with one signum or none. . Piesmopoda (p. 77) 


Forewing with vein 6 from below upper angle of cell, separated at base from the stalk 
of veins 8-9. Male genitalia with a stout free spine associated with anellus. 
Davara (p. 73) 
Forewing with vein 6 from upper angle of cell, connate with the stalk of veins 8-9. 
Male genitalia without free spine associated with anellus . . . . Sarasota (p. 76) 
Male antenna unipectinate for basal half of shaft, shortly ciliate beyond. Female 
with two signa developed as strongly spined plates . ..... Magiriopsis (p. 94) 
Male antenna with shaft pubescent. Female with one signum ornone ..... . 13 
Male with maxillary palpus in the form of an aigrette; eighth abdominal segment with 
compound ventral scale and hair tufts. Female genitalia without signum; the 
bursa copulatrix weakly sclerotized throughout. . ..... Ancylostomia (p. 95) 
Male with maxillary palpus squamous; eighth abdominal segment with paired ventro- 
lateral hair tufts. Female with signum, consisting of a single round, curved plate, 
densely armed with long stiff spines; bursa otherwise membranous. 
Oryctometopia (p. 158) 


10 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Genus 1: Cryptoblabes 


[Venational division A. Hind wing with vein 3 distinctly before 
lower outer angle of cell; 7 and 8 approximate, or weakly and 
shortly anastomosed beyond cell. Forewing with vein 6 bent 
towards base; 10 from the cell. Male genitalia with transtilla 
complete; uncus bilobed.] 


1. Genus Cryptoblabes Zeller 


Cryptoblabes Zeller, Isis von Oken, 1848, p. 644.—Ragonot, 
Monograph, pt. 1, pp. xliv, 12, 1893.—Staudinger and Rebel, 
Catalog der Lepidopteren des palaearctischen Faunenge- 
bietes, vol. 2, p. 42, 1901.—Mayrick, Revised handbook 
of British Lepidoptera, p. 397, 1928.—Bisset, zn Pierce and 
Metcalfe, Genitalia of the British Pyrales, p. 57, 1938.— 
Janse, Journ. Ent. Soc. South Africa, vol. 14, p. 143, 1941. 
(Type of genus: Cryptoblabes rutilella Zeller, a synonym of 
bistriga (Haworth); figs. 2, 131, 638.) 

Tongue well developed. Antenna pubescent; shaft 
of male notched at base and with curved, horny hook 
protruding from the notch. Labial palpus upturned, 
slender, reaching a little above vertex; third segment 
about two-thirds the length of second, acuminate. 
Maxillary palpus squamous. Forewing smooth; 11 
veins; vein 2 from well before the lower outer angle of 
cell (from lower median vein of cell at about three- 
fourths); vein 3 also before the angle (from lower 
median at about five-sixths); 4 and 5 closely approxi- 
mate at base, rarely (in individual specimens) connate; 
6 bent towards base, from upper angle or from very 
close to upper angle of cell; 10 from the cell, separated 
from stalk of 8-9 at base; male without costal fold. 
Hind wing with vein 2 from middle or just beyond mid- 
dle of lower median vein of cell; 3 from before and 
more or less removed from the outer angle of cell; 4 
and 5 from the angle, closely approximate at base, 
thence diverging, 7 and 8 closely approximate, con- 
tiguous or weakly anastomosing for a short distance 
beyond cell; cell less than one-half the length of wing, 
but not ‘very short” as stated by Ragonot in his 
generic key (Monograph, p. xliv); discocellular vein 
curved. Highth abdominal segment of male simple. 

Male genitalia with apical process of gnathos a small, 
simple hook. Transtilla complete; developed as a 
narrowly banded bridge with more or less elongate 
central projection. Uncus broad, with apical margin 
broadly rounded and invaginate, giving the uncus a 
bilobed appearance. Harpe (in European species) with 
strong hair tufts arising from articulated plates in 
intersegmental area between base of sacculus and ter- 
minal margin of eighth abdominal segment, or with 
long hair tuft from sacculus near its base (rutvlelia). 
Anellus with elongate, narrow, lateral lobes. Aedeagus 
simple; penis with or without cornutus, latter, when 
present, a single, long, spine. Vinculum broad and 
with broad terminal margin more or less concave. 

Female genitalia with ductus bursae membranous, 
finely scobinate towards junction with bursa; genital 
Opening simple except for a narrow, sclerotized band 
behind and above the opening; bursa membranous 
more or less finely scobinate; signum present, developed 
as a stout, blunt, flattened thorn (rutilella) or a patch 


of dense granulations (gnidiella); ductus seminalis 
from bursa. 

The foregoing description was drawn from European 
species which are obviously congeneric. Numerous 
other species have been described in the genus from 
India, Formosa, Japan, Australia and the islands of 
the Pacific. Whether these are all congeneric I do not 
know. Two unidentified species before me from the 
Philippines have genitalia similar in habitus to those 
of the genotype (rutilella) except that the dorsal, inter- 
segmental tufts at base of harpe are lacking. Their 
unci have the same characteristic bilobed appearance. 
The venational character which has been generally 
accepted as defining the genus (i. e., the position of vein 
3 of hind wing in relation to the lower outer angle of 
cell) is variable and illusive, being closer to the angle 
in gnidiella (fig. 1) than in rutilella (ig. 2) and still 
closer in the Philippine species. Indeed, in American 
examples of Acrobasis (=Mineola) vein 3 is often as far 
from the angle as it is in gnidiella. The shorter cell of 
Cryptoblabes, coupled with the position of vein 3, will, 
however, suffice to maintain the group separation made 
between the two genera in our key. 

Cryptoblabes is a distinctly Old World genus with no 
indigenous New World species. Itis represented in the 
Western Hemisphere by only one introduced European 
species (gnidiella). 


1. Cryptoblabes gnidiella (Milliére) 
Fieures 1, 132, 639 
Ephesiia gnidiella Mulliére, Iconographie et description de 
chenilles et Lépidoptéres inedits, vol. 2, p. 308, 1864. 
Cryptoblabes gniediella (Millitre) Ragonot, Monograph, pt. 1, 
p. 16, 1893.—Dyar, Ins. Inse. Menstr., vol. 3, p. 88, 1915. 

Forewing pale brownish fuscous with a faint rosy 
overcast due to more or less diffused longitudinal streaks 
of reddish scales (in fresh and well marked specimens 
especially along the fold, on the veins in outer area, and 
below costa from base); some whitish dusting along 
costa and in the cell, most pronounced as a pale shade 
between the dark discal spots at end of cell; transverse 
lines obscure and not sharply outlined, but distinguish- 
able, whitish ocherous; the antemedial line oblique and 
curved, set well out towards middle of wing; subter- 
minal nearly straight, parallel with termen; discal dots 
separate, blackish fuscous. Hind wing whitish, trans- 
lucent, the veins darkened, a narrow dark shade along 
costa and a narrower dark line along terminal margin. 
Alar expanse, 11-16 mm. 

Male genitalia with heavy hair tufts from plates 
articulating with base of sacculus of harpe; penis armed 
with spinelike cornutus about two-thirds as long as 
aedeagus. Female genitalia with signum developed as 
a dense scobinate-granulate patch; ductus seminalis 
from bursa near attachment of ductus bursae. 

TYPE LOCALITY: France (type in Paris Mus.). 

Foop pLants: Fruits of Chaenomeles japonica, pome- 
eranates, oranges, citron, grapes, raisins, etc. (often 
fallen and desiccated fruit), onion seeds, leaves and 
flowers of Daphne gnidium, flowers of Ricinus communis, 
green corn stalks (reared moth, in USNM, from Hawaii), 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 11 


young stems of Tamarix and the stems, leaves, and 
flowers of Lythrum. Apparently has a various larval 
habit and a wide variety of hosts. A moth Gn USNM) 
from Muar Johore, Malaya (Clausen), was reported as 
reared from a larval predator on Aleurocanthus. 

Distrisution: Mediterranean countries of Europe, 
Africa, and Asia. Presumably widely distributed in 
the east and among the Pacific islands; but some of the 
published records may apply to other species. New 
World distribution: Brermupa (Jan., Feb., Apr., May). 
VeneEzvuELA: El Valle (Aug.). Braz: Séio Paulo 
(Feb.). 


Genera 2-17: Acrobasis to Chararica 


[Venational division B. Hind wing with vein 3 from the lower 
outer angle of the cell or (if from before the angle) close to it; 
7 and 8 approximate beyond cell, rarely shortly and weakly 
anastomosed. Male genitalia with transtilla complete or, 
where incomplete (Bertelia, Hypargyria), the elements strongly 
developed and with expanded apices. Uncus triangulate, or 
hoodlike with rounded apical margin, or spatulate (Birinus).] 


2. Genus Acrobasis Zeller 


Acrobasis Zeller, Isis von Oken, 1839, p. 176; 1848, p. 606.— 
Herrich-Schaffer, Systematische Bearbeitung der Schmet- 
terlinge von Europa, vol. 4, p. 99, 1849.—Heinemann, Die 
Schmetterlinge Deutschlands und der Schweiz, vol. 1, 
pt. 2, p. 175, 1865.—Hulst, Phycitidae of N. Amer., p. 120, 
1890; U. S. Nat. Mus. Bull. 52, p. 418, 1902.—Ragonot, 
Monograph, pt. 1, p. 85, 1893.—Dyar, Proc. Ent. Soc. 
Washington, vol. 10, p. 41, 1908.—Spuler, Die Schmetter- 
linge Europas, vol. 2, p. 214, 1910.—Barnes and McDun- 
nough, Contributions, vol. 2, p. 221, 1914.—Forbes, Cornell 
Mem. 68, p. 614, 1923.—Meyrick, Revised handbook of 
British Lepidoptera, p. 397, 1928.—Pierce and Metcalfe, 
Genitalia of the British Pyrales, p. 10, 1938.—Bisset, in 
Pierce and Metcalfe, op. cit., p. 55, 1938.—Janse, Journ. 
Ent. Soc. South Africa, vol. 4, p. 148, 1941. (Type of 
genus: Phycis tumidella Zincken (=Acrobasis  zelleri 
Ragonot); figs. 133, 640.) 

Mineola Hulst, Phycitidae of N. Amer., p. 126, 1890; U. S. Nat. 
Mus. Bull. 52, p. 419, 1902.—Forbes, Cornell Mem. 68, 
p. 618, 1923. (Type of genus: Myelois indigenella Zeller.) 

Seneca Hulst, Phycitidae of N. Amer., p. 177, 1890. (Type of 
genus: Cateremna tumidulella Ragonot. New synonymy.) 

Acrocaula Hulst, Canadian Ent., vol. 32, p. 170, 1900. (Type 
of genus: Acrocaula comacornella Hulst. New synonymy.) 


Tongue well developed. Antenna pubescent; on 
male, basal segment enlarged and angulate, the shaft 
with a slight sinus at base. Labial palpus upturned, 
reaching to or a trifle above vertex; third segment 
slightly more than half the length of second, acuminate. 
Maxillary palpus filiform. Forewing smooth or with 
transverse antemedian ridge of raised scales; 11 veins; 
vein 2 from well before the angle of the cell; 3 rather 
well separated from 4, but somewhat nearer to 4 than 
to 2; 4 and 5 closely approximate at base or connate 
(rarely, in individual specimens, shortly stalked); 
6 from below upper angle of cell, straight; 10 from the 
cell, usually (except in individual specimens) separated 
from stalk of 8-9 at base; male without costal fold. 
Hind wing with vein 2 from well before lower outer 
angle of cell (from outer two-thirds of lower median); 


3 from before but near the angle; 4 and 5 from the 
angle, connate; 7 and 8 shortly anastomosed beyond 
cell; cell about half the wing length; discocellular vein 
curved. Eighth abdominal segment of male with 
midventral hair tuft. 


Male genitalia with apical process of gnathos a simple, 
elongate hook, or an elongate trifurcate hook. Trans- 
tilla complete, sharply angulate and reaching as far 
back as base of apical process of gnathos; terminal 
margin narrow and indented. Uncus broadly triangu- 
late. Harpe simple.’ Anellus a narrowly sclerotized 
U- or V-shaped plate. Aedeagus simple, rather stout; 
penis with numerous sclerotized wrinklings, otherwise 
unarmed. Vinculum stout, a trifle longer than broad, 
slightly tapering; terminal margin truncate and more 
or less concave. 

Female genitalia with ductus bursae and bursa popu- 
latrix membranous except for a dorsal sclerotized plate 
in genital opening; ductus and bursa more or less scobi- 
nate; signum, if present, consisting of a granulate cup 
or & minute central spine surrounded by a dense cluster 
of scobinations; ductus seminalis from a lobe of bursa 
near junction of bursa and ductus bursae. 


Acrobasis as here defined is something of a composite 
genus, dividing into two distinct groups on the develop- 
ment of the apical process of gnathos. Typical Acro- 
basis, comprising the European species with both 
smooth-winged forms (including the type, twmidella) 
and those with the raised-scale ridge on forewing, and 
all smooth-winged American species (formerly under 
Mineola), have the apical process of gnathos produced 
as a simple, elongate hook (fig. 133b). All our Ameri- 
can species with the raised-scale ridge on forewing 
(except minimella Ragonot) have the apical process of 
gnathos trifurcate, that is, produced as an elongate 
hook with a lateral projection from each side before 
apex. This latter group is strictly North American, 
limited in distribution to the United States, and Canada 
east of the Rocky Mountains (except for A. betulella 
Hulst). As far as I know there are no Old World species 
with a similar gnathos. I had hoped to distinguish 
this distinctly American group as a separate genus under 
one of Hulst’s available names (Seneca or Acrocauwla) on 
the basis of the trifurcate projection of gnathos and the 
raised-scale ridge on the forewing; but minimella Rago- 
not prevents this, as it falls between the two groups, 
having the raised-scale ridge and the simple projection 
of gnathos. The females offer no characters that will 
serve to differentiate the groups. Their genitalia are 
so similar that they cannot be used, in many cases, 
even for specific separation, much less for group division. 

The males of the typical American group with raised- 
scale ridge are somewhat more variable than the females, 
exhibiting slight differences in the shape of the transtilla 
and apical projection of gnathos. Figures of these 
structures are given, for what they are worth, for all the 
species represented by authentic males. The differ- 
ences are comparative only, and I suspect that, when 
extended series of the several species are available, they 


12 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


will prove to be more individual than specific in 
character. 

The reasons for sinking Seneca and Acrocaula into the 
synonymy of Acrobasis are given in the discussion of 
their types (pp. 22 and 24). 


Genus Acrobasis, Species 2-7: A. indigenella to 
A. comptella 


[Male with apical process of gnathos a simple hook; forewing 
smooth.] 


2. Acrobasis indigenella (Zeller) 
Ficures 3, 134, 641 


Myelois indigenella Zeller, Isis von Oken, 1848, p. 867.—Riley, 
Fourth annual report on the noxious, beneficial and other 
insects, of the State of Missouri, p. 38, 1872. 

Phycita nebulo Walsh, Prairie Farmer, p. 308, 1860; Proc. 
Boston Soc. Nat. Hist., vol. 9, p. 312, 1863. 

Phycis indiginella (Zeller) Weed, in Forbes, Fifteenth report of 
the State Entomologist on the noxious and beneficial insects 
of the State of Illinois, p. 65, 1889. 

Acrobasis indiginella (Zeller) Riley, Canadian Ent., vol. 16, 
p. 237, 1884.—Ragonot, Monograph, pt. 2, p. 118, 1893.— 
Forbes, Cornell Mem. 68, p. 619, 1923. 

Phycita (Acrobasis) nebulo (Walsh) Riley, Fourth annual report 
on the noxious, beneficial and other insects, of the State of 
Missouri, p. 38, 1872. 

Phycita (Acrobasis) nebulo nebulella Riley, Fourth annual report 
on the noxious, beneficial and other insects, of the State of 
Missouri, p. 42, 1872. 

Myelois zelatella Hulst, Ent. Amer., vol. 3, p. 136, 1887. 

Mineola indiginella (Zeller) Hulst, Phycitidae of N. Amer., p. 130, 
1890.—Quaintance and Siegler, U. S. Dep. Agr. Farmers’ 
Bull. 1270, p. 49, 1922.—Essig and Keifer, Monthly Bull. 
California Dep. Agr., vol. 22; p. 155, 1933.—McDonough, 
Check list, No. 6115, 1939. 

Mineola indigenella nebulella (Riley) Hulst, Phycitidae of 
N. Amer., p. 131, 1890. 

Forewing gray-brown densely dusted with white; the 
white dusting concentrated on the upper half of wing 
and somewhat between subterminal line and outer 
margin, forming two strongly contrasted white patches, 
one from costa in subbasal area, the other more or less 
triangular and extending from costa, between the 
transverse dark lines, into cell and including the blackish 
discal spots; the whitish terminal area is less sharply 
defined and on many specimens somewhat faint; trans- 
verse antemedial line curving obliquely from basal third 
of costa to middle of inner margin, indicated chiefly by 
its outer dark margin, which begins as a conspicuous 
black triangle on costa; from inner margin at one-third 
a blackish line curves upward to meet the antemedial 
line near costa; the area enclosed between them reddish 
ocherous; a similar, smaller spot of the same color on 
base of inner margin; extreme base of costa blackish; 
subterminal line sinuate, bordered inwardly and out- 
wardly by dark lines which begin as blackish spots on 
costa; from the outer of these a dark band extends 
transversely across to the base of antemedial line at 
inner margin, somewhat obscured in the dark ground 
color on all but the palest and most contrastingly 
marked specimens; discal spots at end of cell black, 


more or less confluent, usually a black bar along dis- 
cocellular vein; a more or less broken, black line along 
terminal margin. Hind wing subpellucid, pale smoky 
fuscous; the veins very faintly, if at all, darkened; 
a narrow, obscure, dark line along termen. Alar ex- 
panse, 15-20mm. Male genitalia as given for the genus. 
Female genitalia without signum. 

Type Locatitizs: ‘‘Carolina”’ (indigenella, in BM); 
Illinois? (nebulo, lost); Missouri? (nebulelia, lost); “New 
York and Canada,” (zelatella; the supposed type, co’, 
in AMNH, ex Rutgers, is labeled “Blanco County, 
Texas’’). 

Foop piants: Apple, crabapple, plum, prune, cherry, 
quince, Crataegus, Cotoneaster, Pyrocantha coccinea. 
Larva feeding on leaves and forming serpentine resting 
and hibernating case of silk and frass. 

Disrrisution: Unirep Starrs: Maine, Sebec Lake 
(uly); New Hampshire, Hampton (July); Vermont, 
Clarendon; Massachusetts, Newton Highlands; Connec- 
ticut, Hast River (July); New York, Catskill Mts., Ilion; 
New Jersey, Rutherford; Pennsylvania, Chambersburg 
(June, July), Germantown (July), New Brighton (July), 
Pittsburgh (June); Virginia, Colonial Beach (July), 
Norfolk (May); North Carolina, Tryon (Aug.); Georgia, 
Savannah (Apr.); Illinois, Chicago, Decatur (May, 
June, July, Aug.); Mssourt, Mossele (June, July), 
Norborne (Apr., June, July), St. Louis (Aug.); Nebraska, 
Wahoo (May); Kansas, Onaga, Wichita (June); Missis- 
sippi, ‘Agr. College” (May, June, July); Arkansas, 
Siloam Springs (June), Washington County (July); 
Texas, Abilene, Blanco County (Sept.), Fort Worth 
(Sept.), Houston (May, June, Aug.), Kerrville, Victoria 
(June); California, Lomita, Los Angeles County (Mar.) 
Orange County (June). (The California records all 
from plum.) Canapa: Ontario, Trenton (June, July); 
Quebec, Meach Lake (July). 

Apparently generally distributed east of the Rocky 
Mountains and rather recently introduced into Cali- 
fornia. 

This species has been considered of economic impor- 
tance as a defoliator of fruit trees in the Middle States, 
and in the official list of common names approved by 
the American Association of Economic Entomologists 
is designated as the “leaf crumpler.”” However, it does 
not seem to be more than a minor pest of local and 
occasional concern. Several references are made to it 
in the economic literature but none of these adds any- 
thing of significance to our knowledge of the insect 
beyond what is given in the early papers by Walsh and 
Riley. 

Riley’s nebulella was described by him as a variety 
of nebulo and distinguished from the latter by the more 
diffused dark shading and the separation of the discal 
spots on forewing. In the series before me there is 
considerable variation in the extent and intensity of the 
dark coloring and the discal mark varies even more, 
being sometimes divided into two distinct spots or fused 
into a single bar on different sides of the same specimen, 
so the varietal designation is hardly worth maintaining. 
In 1908 (Proc. Ent. Soc. Washington, vol. 10, p. 45) 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 13 


Dyar, on the evidence of a supposed type of nebulella 
in the National Collection, resurrected the name and 
applied it to the ‘‘pecan leaf casebearer.’’ Barnes and 
McDunnough (Contributions, vol. 2, p. 222, 1914; 
vol. 3, p. 221, 1917) called attention to the spuriousness 
of the alleged “‘type” and gave the pecan casebearer its 
proper reference (juglandis LeBaron); but on the 
strength of Dyar’s identification the name nebulella 
had already appeared, and continued to be used for some 
years in economic publications for the pecan leaf case- 
bearer. In his 1939 Check List McDunnough applies 
the name in an entirely new sense, transferring it to 
Meroptera with the well-known unicolorella Hulst as a 
syhonym. This was most unfortunate and altogether 
unnecessary. We know what unicolorella Hulst is, and 
its type is at hand for reference. The type of nebulella 
is nonexistent and McDunnough’s new reference has 
nothing to back it but an entomologist’s interpretation 
of Riley’s description and very poor and over-inked 
figure of the forewing. I see nothing in either to rule 
out the original interpretation, so shall let the name 
sleep in synonymy. 


3. Acrobasis grossbecki (Barnes and McDunnough), new combi- 
nation 


Mineola indigenella nebulella Grossbeck (not Riley), Bull. Amer. 
Mus. Nat. Hist., vol. 37, p. 129, 1917.—Barnes and Mc- 
Dunnough, Contributions, vol. 3, p. 220, 1917. 

Mineola grossbecki Barnes and McDunnough, Contributions, 
vol. 3, p. 221, 1917——McDunnough, Check list, No. 6116, 
1939. 

Forewing purplish brown, the dark ground color 
more extended than in indigenella, obscuring the pale 
antemedial line and completely obliterating the reddish 
ocherous subbasal patch on inner margin usually present 
in the genus; triangular black spot on costa, beginning 
the outer dark border of antemedial line, distinct and 
sharply contrasted as in indigenella; white areas 
restricted more than in indigenella, the subbasal one 
narrowly triangulate with its point on inner margin, 
midcostal one extending to and including the discal 
spots in its lower angle; whitish dusting in terminal 
area very faint; subterminal line obscure; discal spots 
at end of cell black, separated. Hind wing shiny, 
smoky fuscous. Alar expanse, 15-16 mm. 

Genitalia as in indigenella. 


Tyre tocauity: Lakeland, Fla. (type in USNM). 

Foop puant: Crataegus (larva feeding on the leaves). 

Distrisution: Known only from the type locality. 
May be a Florida race of indigenella, but appears to 
be a distinct species despite the likeness of its genitalia 
to those of indigenella. 


4. Acrobasis vaccinii Riley 
Fieure 642 


Acrobasis vaccinii Riley, Canadian Ent., vol. 16, p. 237, 1884; in 
Rep. [U. 8.] Comm. Agr. for 1884, p. 355, 1885.—Smith, 
in Rep. [U. S.] Comm. Agr. for 1884, p. 394, 1885.—Saunders, 
Insects injurious to fruits, p. 375, 1883 (as ‘“The Cranberry 
Fruit-worm”).—Ragonot, Monograph, pt. 1, p. 121, 1893.— 
Forbes, Cornell Mem. 68, p. 618, 1923. 


Mineola vaccinii (Riley) Hulst, Phycitidae of N. Amer., p. 128, 
1890.—Brown, Oregon Agr. Exp. Station Bull. 225, p. 19, 
1927.—Crowley, Washington Agr, Exp. Station Bull. 230, 
p- 24, 1929——McDunnough, Check list, No. 6114, 1939.— 
Beckwith, Journ. Econ. Ent., vol. 34, p. 169, 1941. 

Averaging smaller than indigenella; dark ground 
color similar but more extended and without the con- 
trasting black costal triangle; pale antemedial line 
obliterated by a transverse extension of the ground 
color, bordered inwardly by an almost vertical, rather 
narrow white band which expands narrowly on costa 
towards base (the remains of the much-reduced sub- 
basal white area); midcostal white patch also much 
restricted, barely including at its lower angle the 
separated black discal spots; on fresh specimens some 
sprinkling of rufous scaling is distinguished under high 
magnification, but no reddish or other contrastingly 
colored, angulate, subbasal patch on inner margin (as 
in indigenella and tricolorella). Hind wing pale smoky 
fuscous. Alar expanse 14-18 mm. 

Male genitalia differing in no significant detail from 
those of indigenella. Female genitalia with bursa 
more or less heart-shaped (less elongate than that of 
indigenella or grossbecki); signum present as a minute 
granulate cuplike patch. 


Typr Locauity: Massachusetts (type in USNM). 

Foop puant: Cranberry, blueberry (larva in the 
fruit). 

Distripution: Massachusetts (type series, no exact 
locality, June), Wareham (June, July); Connecticut, East 
River (July); New Jersey, Pemberton (May), Whitesbog 
(June); Wisconsin; Michigan; Georgia; Mississippi, 
Biloxi, Poplarville; Washington, Long Beach (June), 
Seaview (July). 

Presumably generally distributed on the range of its 
food plants in the United States and Canada. The 
foregoing records are from reared and typical examples 
in the National Collection. 

This species, popularly known as the ‘cranberry 
fruitworm,”’ is of some importance, especially to cran- 
berry growers, and has a rather extensive economic 
literature, mostly in annual reports, bulletins, and other 
publications of state entomologists and experiment 
stations. None of these adds anything of biological or 
taxonomic significance to the earlier records of Riley 
and Smith. 


5. Acrobasis amplexella Ragonot 


Acrobasis amplexella Ragonot, N. Amer. Phycitidae, p. 3, 1887; 
Monograph, pt. 1, p. 97, 1893.—Forbes, Cornell, Mem. 68, 
p. 618, 1923. 
Mineola amplexella (Ragonot) Hulst, Phycitidae of N. Amer., 
p. 127, 1890.—McDunnough, Check list, No. 6112, 1939. 
This is probably nothing but a color form of vaccinii. 
I can find no difference from the latter except in the 
greater extension of the basal dark area of forewing and 
the consequent further restriction of the subbasal white 
area which is a narrow band throughout, not expanding 
along costa towards base. 
Rearing will have to settle the status of amplezella. 
In the material before me there are only collected 


14 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


specimens. All reared examples we have from either 
cranberry or blueberry are typical vaccinii. 

Alar expanse, 12-18 mm. 

Typz xocauity: North Carolina (type in Paris 
Mus.). 


Foop prant: Unknown. 

Distrizution: Maine, Monmouth (June), Sebec 
Lake (July); New Hampshire (June, July); Massachu- 
setts, Cohasset (July), Framingham (June), Winchendon 
(July), Worcester (July) ; Connecticut, Hast River (July); 
New York, Liberty (June), Sullivan County (July); 
Pennsylvania, Hazleton (May); North Carolina, Tryon 
(July). 

In the Barnes and old U. S. National Museum Col- 
lections these examples were about equally divided 
under the two names, amplexella and vaccinii. 


6. Acrobasis tricolorella Grote 
Figures 135, 643 


Acrobasis iricolorella Grote, Bull. U. S. Geol. Geogr. Surv. Terr., 
vol. 4, p. 694, 1878.—Ragonot, Monograph, pt. 1, p. 93, 
1893.—Forbes, Cornell Mem. 68, p. 618, 1923. 

Mineola tricolorella (Grote) Hulst, Phycitidae of N. Amer., p. 
127, 1890.—McDunnough, Check list, No. 6111, 1939. 
Mineola scitulella Hulst, Canadian Ent., vol. 32, p. 169, 1900.— 

Hungerford, Idaho Agr. Exp. Station Bull. 149, p. 29, 
1927; Bull. 164, p. 29, 1929.—Pack and Dowdle, Journ. 
Econ. Ent., vol. 23, p. 321, 1930.—Haegele, Journ. Heon. 
Ent., vol. 25, p. 1078, 1982.—Essig and Kiefer, Monthly 
Bull. California Dep. Agr., vol. 22, p. 153, 1983.—McDun- 
nough, Check list, No. 6110, 1939. (New synonymy.) 
Forewing grayish fuscous more or less dusted with 
whitish scales, in some specimens well diffused over the 
middle of wing, giving it a pale slate ground color, but 
normally concentrated into a pale patch from costa 
before subterminal line, including the discal mark, and 
a pale terminal suffusion below apex; usual subbasal 
white area constricted into a narrow, sharply defined 
antemedial band, outwardly oblique from costa to top 
of cell, thence vertical to inner margin and bordered 
outwardly on its vertical portion by a more or less 
triangular, contrasted orange or reddish orange patch; 
bordering the white line and the orange patch out- 
wardly, a black oblique angulate line extending to 
near middle of inner margin and beginning on costa in 

a more or less angulate and diffused blackish patch 

(similar to but not so sharply defined nor contrasted as 

the black costal patch on indigenella) ; subterminal line 

distinct, narrow, white, angled inwardly at vein 6 and 
lower fold and curved outwardly between, bordered 

inwardly by a narrow black line and outwardly by a 

black costal spot and a more or less pronounced orange 

or reddish orange band (well marked in many eastern 
and western specimens, but sometimes obscured by 
dark scaling); discal spots usually fused into a slightly 
curved, black bar along discocellular vein, rarely 
separated. Hind wing smoky white to pale smoky 

fuscous. Alar expanse, 18-22 mm. 

Male genitalia with cucullus of harpe of more even 
width throughout and apex more evenly rounded than 
in other species of the genus. Eighth abdominal seg- 


ment of male with a single central, ventral hair tuft 
(supplementary ventral hair tufts on the other species). 
Female genitalia with signum present as a small, 
granulate, cup-shaped patch. 

TYPE LOCALITIES: Oldtown, Maine (éricolorella, in 
BM); Colorado (scitulella, in AMNH, ex Rutgers). 

Foop Puants: Apple, apricot, catalina cherry, plum, 
prune (larvae feeding in buds and fruits, probably also 


on leaves), also recorded from galls on chokecherry 
(Park and Dowdle). 


Distrrsution: Unitep Status: Maine, Oldtown, 
Bar Harbor (July), Orono; New Hampshire, Hampton 
(July); Massachusetts, Amherst (Hatch Exper. Station, 
July); New York, Catskill Mts., Ilion (July); New 
Mexico, Albuquerque (July); Colorado, Denver, Glen- 
wood Springs (May, July, Aug., Sept.), Grand Junction 
(Aug.), Gunnison County (July); Utah, Bellevue (May), 
Dividend (Aug.), Eureka (Aug.), Logan (July), Park 
City (July), Provo (July); Zdaho, Boise (Sept.), Emmett 
(July); California, Loma Linda (Aug.), Mount Lowe 
(May, July), San Diego (June, July), Santa Barbara 
(Aug.), Warner Mts. (Modoc County, July); Oregon 
Lake View (Aug), The Dalles (June); Washington, 
Prosser (June), Pullman (July), Walla Walla (Aug.), 
Wenatchee (Aug.). Canapa: Ontario, Ottawa (July); 
Manitoba, Cartwright; British Columbia, Arrowhead 
Lake (June). 

The species seems to be abundant in our western 
states and relatively scarce im the east, to judge by 
examples in collections, and has attracted some atten- 
tion as a fruit pest in Utah, Idaho, and California. 
There is nothing to distinguish western from eastern 
specimens and the one detail that Hulst relied upon 
for the separation of his scitulella (the presence of 
an orange outer border to the subterminal line) does not 
hold. It is present in eastern and western examples 
and equally variable in both. I am therefore sinking 
the name in the synonymy of tricolorella. 


7. Acrobasis comptella Ragonot 
Figure 646 


Acrobasis compitella Ragonot, N. Amer. Phycitidae, p. 4, 1887.— 
Hulst, Ent. Amer., vol. 5, p. 156, 1889 (makes synonym of 
caliginella Hulst). 


Forewing dark gray dusted with white, the white 
dusting concentrated on basal area, on costal median 
half of wing (forming a pale angulate patch which 
includes the discal spots), and in terminal area beyond 
subterminal line; in some specimens the white dusting 
is more extended, making most of the basal, median, 
and terminal areas pale ashy gray; outwardly bordering 
basal pale area a black line (narrowing from a shallow 
triangulate patch on costa) extends obliquely outward 
to top of cell thence vertically to inner margin, bordered 
outwardly on vertical part by a triangulate, tawny or 
reddish brown patch; subterminal line white, sinuate, 
bordered inwardly by a narrow, blackish line and out- 
wardly, at costa, by a black smudge; discal spots at end 
of cell distinct, separate, black. Hind wing pellucid, 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 15 


whitish or pale smoky fuscous. Alar expanse, 14-21 
mm. 

Male genitalia exhibiting no distinctive specific 
characters. Female genitalia with several short, paral- 
lel lines of fine scobinations in bursa; signum present as 
a small, granulate, cup-shaped patch. 

Typr Locatity: California (type in Paris Mus.). 

Foop puant: Unknown. 

Distrisution: California, Cisco (July), Colfax (July), 
Pasadena, San Diego (June), Santa Catalina Isl. (May), 
Warners (San Diego County, Aug.); Arizona, Gila 
County, Redington; New Mezico, Albuquerque (July), 
Las Vegas; Utah, Provo (July). 

Hulst in 1889 made comptella a synonym of his 
caliginella and it has remained as such in our lists. 
However, the two are generically as well as specifically 
distinct, caliginella having the basal segment of the 
male antenna cylindrical (not triangularly expanded at 
apex as in Acrobasis). It is superficially similar in 
color and markings to comptella; but the black line 
bordering the whitish basal patch of forewing is 
distinctly broken, its vertical portion not reaching to 
inner margin. I am removing both caliginella and 
Mineola supposita Heinrich to Rhodophaea. 


Genus Acrobasis: Species 8. A. minimella 


[Male with apical process of gnathos a simple hook; forewing 
with raised-scale ridge.] 


8. Acrobasis minimella Ragonot 
Fieure 140 


Acrobasis minimella Ragonot, Ent. Amer., vol. 5, p. 113, 1889; 
Monograph, pt. 1, p. 105, 1898.—MeDunnough, Check list, 
No. 6088, 1939. 

Acrobasis nigrosignella Hulst, Phycitidae of N. Amer., p. 123, 
Map ee Proc. Ent. Soc. Washington, vol. 10, p. 43, 

Forewing grayish fuscous with a purplish suffusion; 
some whitish dusting on basal area and, very faintly, 
from costa before subterminal line to discal spots; 
antemedial line obscure except towards inner margin, 
where it is a narrow whitish line; a blackish triangular 
costal patch following the antemedial line and continued 
as thin black line on its outer border to inner margin; 
vertical scale ridge black; area between scale ridge and 
antemedial line ocherous or reddish; discal spots at 
end of cell small, obscure, separated. Hind wing smoky 
fuscous. Alar expanse, 13-16 mm. 

Female genitalia exhibiting no specific difference to 
distinguish them from those of other species having the 
raised-scale ridge on forewing. 

TypE LocaLity: Texas (minimella, 9, in Paris Mus., 
and nigrosignella, in AMNH, ex Rutgers). 

Foop prant: Oak (this food plant record from speci- 
men, in USNM, reared at Falls Church, Va., under 
Hopkins No. 9847, C. F. Johansen). 

Distrizution: Teras (Apr.); Mississippi, Starkville 
(July); Louisiana, Winfield (June); North Carolina, 
Southern Pines (Apr., May, June, Aug.), Tryon (May); 


Virginia, Falls Church; District of Columbia, Washing- 
ton (June); New Jersey, Lakehurst (July). 

The species is easily recognized by its size, color, and 
male characters. The sex-scaling is present and con- 
sists of long broad black costal streaks on underside of 
fore and hind wings. 


Genus Acrobasis, Species 9-22: A. feltella to A. 
demotella 


[Male: Apical process of gnathos trifurcate; forewing with raised- 
scale ridge; black sex-scaling beneath.] 


9. Acrobasis feltella Dyar 
Ficure 141 
Acrobasis feltella Dyar, Proc. Ent. Soc. Washington, vol. 11, 
p. 214, 1910.—Ely, Ins. Inse. Menstr., vol. 1, p. 51, 1913.— 
McDunnough, Check list, No. 6080, 1939. 

Head, basal segment of antenna, thorax, and basal 
area of forewing white; a faint rosy tint on the posterior 
of thorax and a more obvious rosy shading on lower 
half of pale basal area of forewing (more intense and 
extended on the female than on the male); remainder 
of wing dark gray-brown (in fresh specimens blackish 
brown) with a faint, pale grayish shading in terminal 
area and a whitish spot on inner margin near tornus 
(the lower end, and contrasted portion of the otherwise 
obscure subterminal line); black discal dots at end of 
cell distinguishable but somewhat obscured in the dark 
ground color, usually separate, but occasionally fused; 
antemedial scale ridge blackish. Hind wing of male 
white at base, shading to smoky fuscous outwardly; 
the veins in both sexes faintly darkened. Alar expanse, 
14-18 mm. 

Black sex-scaling consisting of a short patch at base 
of costa on forewing. 

Type tocatity: Warner, N. Y. (type in USNM). 

Foop prant: Hickory (larva boring in petioles). 

DisrrisuTion: Unirep States: New York, Warner 
(July) ; Connecticut, Hast River (July) ; Zilinois, Putnam 
County (June). Canapva: Ontario, Merivale (June). 

This species, palliolella Ragonot, and caryalbella Ely 
are identical in color, maculation, and all superficial 
characters. They exhibit trifling differences in their 
male genitalia, especially in the shapes of their trans- 
tillae and the apical processes of their gnathi. These 
differences are probably no more than individual in 
character. We figure them for what they are worth. 
Ely (1913) noted differences in the larval cases of 
feltella and caryalbella which should be significant. He 
also saw, or thought he saw, a difference in the sex- 
scaling of Dyar’s type and the type of caryalbella. In 
this he was in error; for the sex-scaling is identical in 
both types and on the males of palliolella and juglandis 
as well. I suspect that the three names (feltella, 
palliolella, and caryalbella) apply to a single species; but 
this cannot be determined until the biologies and larvae 
of the various hickory-feeding forms of the genus are 
more thoroughly studied. Until that is done it seems 
best to keep the names separated. 


16 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


10. Acrobasis palliolella Ragonot 
Ficgure 142 
Acrobasis palliolella Ragonot, N. Amer. Phycitidae, p. 4, 1887; 
Monograph pt. 1, p. 92, 1893.—Hulst, Phycitidae of N. 
Amer., p. 121, 1890. 
Acrobasis albocapitella Hulst, Ent. Amer., vol. 4, p. 116, 1888. 

Not distinguishable superficially from feltella. Tri- 
fling differences in the male genitalia are shown in the 
figure. They are probably not significant. 

Type tocauities: North America (palliolella, in 
Paris Mus.); Canada (albocapitella, in AMNH, ex 
Rutgers). 

Foop pLant: Presumably hickory. Life history not 
known. 

Distrisution: Unirep States: Illinois, Chicago 
(July); Pennsylvania, New Brighton (July); North 
Carolina, Plymouth (May); Connecticut, East River 
(July). Canapa: Ontario, Ottawa (July). 

The name palliolella has been variously misapplied 
and has appeared frequently in economic literature for 
the ‘‘pecan leaf casebearer”’ (juglandis LeBaron). In 
our latest checklist (McDunnough, 19389) it appears as 
a synonym of juglandis but I do not think this is correct. 

A long series of juglandis before me shows consider- 
able variation in color but at the same time consistent 
differences from palliolella, whose closest affinities are 
feliella Dyar and caryalbella Ely. 


11. Acrobasis caryalbella Ely 
Fiaure 143 
Acrobasis caryalbella Ely, Ins. Insc. Menstr., vol. 1, p. 52, 
1913.—McDunnough, Check list, No. 6081, 1939. 


Acrobasis angusella Dyar (not Grote), Proc. Ent. Soc. Washing- 
ton, vol. 10, p. 42, 1908. 


Ely distinguishes his species from Dyar’s feltella 
chiefly on the differences in their larval cases (‘‘co- 
coons”). There is nothing else to separate them except 
some slight and probably not significant differences in 
their genitalia. These are shown in the figure. 

Typxz Locauity: East River, Conn. (type in USNM). 

Foop puant: Hickory. 

Known only from reared examples from the type 
locality and the female from hickory (June) without 
locality label, bearing Riley’s No. 376 and referred by 
Dyar to angusella Grote. The sex-scaling on the male 
is the same as that on feltella and palliolella. 


12. Acrobasis juglandis (LeBaron) 
Fieurrs 138, 644 


Phycita juglandis LeBaron, Second annual report on the noxious 
insects of the State of Illinois, p. 23, 1872. 

Acrobasis juglandis (LeBaron) Riley, Fourth annual report on 
the noxious, beneficial and other insects of the State of 
Missouri, p. 42.—Barnes and McDunnough, Contributions, 
vol. 3, p. 221, 1917—McDunnough, Check list, No. 6082, 
1939.—Moznette (and others), U. S. Dep. Agr. Farmers’ 
Bull. 1829, p. 16, 1940.—Craighead, U. S. Dep. Agr. Misc. 
Publ. 657, p. 449, 1950. 

Acrobasis nebulella Dyar (not Riley), Proc. Ent. Soc. Washing- 
ton, vol. 10, p. 45, 1908. 

Aerobasis palliolella Dyar (not Ragonot), Proc. Ent. Soc. Wash- 
ington, vol. 10, p. 44, 1908.—Forbes (in part), Cornell 
Mem. 68, p. 617, 1923. 


Similar to the three preceding species except: White 
basal area more or less shaded with ashy gray; without 
rosy tints on lower basal and outer areas; generally 
paler in outer area, mouse gray, with the white dusting 
from midcosta somewhat more intense; a distinct 
blackish costal triangle following the antemedial line. 
Hind wing smoky fuscous shading to rather dull white 
towards base on the male, darker and more uniformly 
colored on the female. Alar expanse, 14-17 mm. 

TypE Locairy: Illinois (type lost). 

Foop puiants: Hickory, pecan, walnut, butternut 
(larvae feeding on leaves, buds, and flowers). 

Distrisution: Iilinois, Chicago (July); Missouri; 
Mississippi, Wiggins (May); Texas, Black Springs, 
Brownsville (May), Cuero (June), Kerrville (May, 
June), Victoria (May, June); Georgia, Albany (July), 
Atlanta, Blackshear (May, June), Cairo (May, June); 
Florida, Monticello (May, June), Orlando (May), 
Palatka (May), Tallahassee (May); South Carolina, 
Mt. Pleasant (July); North Carolina, Edgecombe 
County (May), Plymouth (May); District of Columbia, 
Washington (June). 

A large reared series in the National Museum is 
mostly from pecan. Also before me a series reared 
from walnut and butternut that appears to be a suffused, 
dark form of juglandis. Two specimens of the latter 
series are from Ontario, Canada. 

In our Gulf States the species is of some importance 
as a defoliator of pecan, and is popularly known as 
the ‘‘pecan leaf casebearer.”” It has numerous refer- 
ences in economic literature. I have retained only one 
of these (Moznette, 1940), for it gives all the biological 
information available on the species under its correct 
specific name. Dyar’s unfortunate identifications have 
greatly confused the nomenclature, with the result that 
most economic references previous to 1939 are under 
nebulella or palliolella. Hulst (Phycitidae of N. Amer., 
p. 131, 1890) and Ragonot (Monograph, pt. 1, p. 120, 
1893) are also at fault in applying the name juglandis. 
Their descriptions apply to examples of indigenella and 
not to the “pecan leaf casebearer.”’ 

The sex-scaling of juglandis is like that on feltella. 


13. Acrobasis sylviella Ely 
Fieure 144 
Acrobasis sylviella Ely, Proc. Ent. Soc. Washington, vol. 10, 
p. 161, 1908.—McDunnough, Check list, No. 6094, 1939. 

Forewing pale ashy gray; the basal area, thorax, and 
head but slightly paler, not contrastingly whitish; no 
triangular black spot on costa outside the antemedial 
line, the dark outer border of the antemedial line a 
narrow band or weak, diffused shade from costa. Hind 
wing pale smoky fuscous on male, slightly darker on 
female. Alar expanse, 19-21 mm. 

Typr Locauity: East River, Conn. (type in USNM). 

Foop PiLant: Ostrya. 

Distrisution: Unirep Srares: Connecticut, East 
River (July); Pennsylvania, New Brighton (May). 
Canapa: Ontario, South March (June). 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 17 


The Pennsylvania specimen was in the Barnes 
Collection as cirroferella Hulst. This is incorrect, 
however, as the male type of cirroferella is without 
sex-scaling. The Canadian specimens (one male and 
one female) were reared from Osirya and are responsible 
for the food plant record. They were tentatively 
identified by McDunnough as sylviella and I think 
correctly. They are considerably darker than the type 
series of the Pennsylvania specimen; but the fact that 
they were reared, and probably under excess moisture, 
would easily account for the difference. 

The black sex-scaling of sylviella is similar to that of 
feltella but slightly more extended, reaching slightly 
beyond basal fourth of costa on the underside of fore- 
wing. 

14, Acrobasis kearfottella Dyar 
Figure 145 


Acrobasis kearfottella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 34, 1905.—Ely, Ins. Inse. Menstr., vol. 1, p. 53, 1913.— 
McDunnough, Check list, No, 6079, 1939. 

p- ‘The most distinct and strikingly marked of the Ameri- 

can Acrobasis species; costal half or third of basal area 

of forewing snow white, this white area extending out 
along costa to subterminal line and broadening to in- 
clude the blackish discal spots; midcostal margin nar- 
rowly black-edged; white area uncrossed at any place 
by dark lines; subterminal line faint, but distinct, dull 
white; remainder of forewing dark gray-brown. Hind 
wing of male white shading to fuscous at apex and 
terminal margin; hind wing of female pale glossy brown 
throughout. Thorax and head of male show white; of 
female concolorous with dark area of forewing. Alar 
expanse, 18-21 mm. 


TYPE Locatity: Cleveland, Ohio (type in USNM). 

Foop piant: Hickory (larvae feeding on the leaves). 

Distrisution: Ohio, Cleveland (June); New York, 
lion (July); Connecticut, East River (July); Pennsyl- 
vania, New Brighton (July); North Carolina, Black 
Mountain; Jilinois, Oconee (Aug.). 

Very little is known of the life history. Ely’s paper 
describes the cocoon. The black sex-scaling on the 
male is similar to that of juglandis and the preceding 
hickory-feeding species. 


15. Acrobasis caryae Grote 
Fiaures 1387, 146 


Acrobasis caryae Grote, Papilio, vol. 1, p. 18, 1881; Bull. U. S. 
Geol. Geog. Surv. Terr., vol. 6, p. 591, 1882.—Hulst, 
Phycitidae of N. Amer., p. 122, 1890.—Ragonot, Mono- 
graph, pt. 1, p. 105, 1893——Barnes and McDunnough, 
Contributions, vol. 2, p. 222, 1914.—Forbes, Cornell Mem. 
68, p. 617, 1923—McDunnough, Check list, No. 6100, 
1939.—Moznette and others, U. S. Dep. Agr. Farmers’ 
Bull. 1829, p. 2, 1940.—Craighead, U. S. Dep. Agr. Misc. 
Publ. 657, p. 449, 1950. 

Acrobasis caryaevorella Dyar (not Ragonot), Proc. Ent. Soc. 
Washington, vol. 10, p. 44, 1908. 

Acrobasis hebescella Dyar (not Hulst), Proc. Ent. Soc. Washing- 
ton, vol. 10, p. 44, 1908. 


Forewing glossy gray (in southern specimens from 
pecans pale and with little darker shading except 


narrowly along antemedial line); basal area concolorous 
with median area except in some of the darker speci- 
mens; northern specimens from hickory normally dark 
grayish fuscous; antemedial line whitish towards inner 
margin; raised-scale ridge black, preceded by some white 
scaling and followed by a narrow, more or less obscured, 
flesh-colored patch; subterminal line pale gray, obscure; 
discal dots distinct and separate but not strongly 
contrasted against ground color. Hind wings smoky 
fuscous. Alar expanse, 18-20 mm. 

Genitalia exhibiting no distinguishing specific char- 
acters; figured from southern male reared from pecan 
nut. The scale tufting on the eighth abdominal 
segment of the male consists of a single, rather long, 
central ventral tuft like that shown in figure 137 and 
similar to that of the European type of the genus 
(tumidella). 

TypeE Locatity: Illinois (type in BM). 

Foop piants: Hickory, pecan (overwintering larvae 
feeding in early spring upon opening leaves and in the 
stems of new growth; later generations in the nuts. 
Larva does not make a case during feeding period). 

Disrrisurion: Unrrep States: Florida, Monticello 
(June, July, Aug.), Tallahassee (May) ; Georgia, Albany 
(July); Mississippt, Goodman (July), Ocean Springs 
(May, Sept.), Pascagoula (June), Wiggins (June, July) ; 
Texas, Boerne (June), Bosque (May), Brownwood 
(Apr., May, June, July, Aug., Sept.), Colorado River 
(Apr., May), Cuero (Aug., Sept.), Dallas (May), Fort 
McKevett (June), Pecan Bayou (July), Pioneer (Aug.), 
San Saba (May), Texas A. and M. College Station 
(June, July), Victoria (June, July, Sept.); Illinois, 
Chicago, (July), Decatur (June); Pennsylvania, New 
Brighton (July, Aug., Sept.); North Carolina, Mill 
Brook; District of Columbia, Washington (May, June) ; 
Connecticut, East River (July, Aug.). Canapa: On- 
tario, Merivale (June). 

This is the “pecan nut casebearer’”’ of economic 
literature. It has a rather extended literature but is 
of importance only as a pecan pest in the Gulf States. 
Most of the economic references before 1929 are to 
hebescella and caryaevorella as a result of Dyar’s mis- 
identification of those species. I cite only one economic 
reference here, as the Moznette (1940) paper gives all 
the biological information available on the species as 
@ pecan insect. Its biology as a hickory insect in the 
north is imperfectly known. 

The sex-scaling of the male is characteristic, consist- 
ing of a short black patch on base of costa of forewing 
(as in feltella) and a long black streak along the top of 
cell on the underside of forewing. This combination 
is peculiar to caryae and evanescentella. 


16. Acrobasis evanescentella Dyar 
Fieure 147 


Acrobasis evanescentella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 44, 1908.—McDunnough, Check list, No. 6086, 1939. 
Doubtfully distinct from caryae. The dark areas of 
forewing beyond base have a purplish luster, and the 
pale (whitish dusting) is more distinct, forming a pale 


18 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


grayish white spot on costa before subterminal line 
(and extended to include the discal spots); sex-scaling 
on underside of wing as in caryae, except that the 
streak along top of cell is somewhat obscured by an 
overlay of pale scales. Under these it is black (not 
“pale gray”as stated by Dyar) and no narrower than 
that of caryae. 

The only authentic specimens I have seen are those 
of the original type series. They are in excellent con- 
dition. The other specimens which Dyar later asso- 
ciated with them are all typical caryae. The note 
with the type lot (‘Chittenden 250”) tells nothing 
about the larval habits, so we do not know whether 
there are any biological characters to distinguish 
evanescentella from caryae. We shall have to hold the 
name until the biology is thoroughly studied. 

Type wocauity: Orlando, Fla. (May) (type in 
USNM). 

Foop PLant: Pecan. 


17. Acrobasis stigmella Dyar 
Fiaure 148 
Acrobasis stigmella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 43, 1908.—Forbes, Cornell Mem. 68, p. 616, 1923.— 
McDunnough, Check list, No. 6089, 1939. 

Forewing purplish gray; the basal area very dark 
(darker than remainder of wing); scale ridge scarcely 
darker, its outer margin edged with dull red; discal dots 
obscure; subterminal line very faint. Hind wing of 
male pale smoky fuscous with a slight ocherous tint; 
of female darker and without the ocherous tint on 
upper surface. 

On the male the upper surface of the head is yellowish 
white, the thorax, purplish gray. On the female the 
head and thorax are concolorous, purplish gray. 
Alar expanse, 17-21 mm. 

Typ tocauity: Fort Lee, N. J. (type lost). 

Foon piant: Hickory. 

Distrisution: New Jersey; Connecticut, Hast River 
(Aug.); District of Columbia, Washington (May, June), 
Virginia, Falls Church (June); Illinois, Decatur (May), 
Putnam County (July). 

In his description Dyar states that it is based upon 
two males and one female from ‘‘Fort Lee, N. J., May 
1896 (H. G. Dyar)” and one female from ‘‘Hast River, 
Conn., Aug. 20, 1906 (Chas. R. Hly).” The female 
from Hast River is in the National Collection but there 
are no specimens from Fort Lee and none dated 1896. 
The specimen (a male in good condition) bearing Dyar’s 
type label is one reared by him at Washington, D. C., 
June 19, 1900. This probably was overlooked by him 
at the time he prepared his description and not identified 
or labeled until later. It is unquestionably stigmella 
but, in the light of his published declaration, cannot be 
accepted as the type. 

The species is quite distinct and easily identified— 
especially the males by their contrasting yellow-white 
heads against their dark thoraces. The sex-scaling on 
underside of the male and the contrasting dark basal 
patch of forewing distinguish it from everything else 


except aurorella Kly. The sex-scaling consists of a nar- 
row black streak on forewing extending for about one- 
fourth of costa from base, a strongly contrasted and 
rather broad black midcostal streak on hind wing and 
some black scaling on the extreme base of the veins of 
cell of the hind wing. The underside of hind wing is 
otherwise a uniform ocherous white. 

The life history is also characteristic. In early spring 
(Mar.) the young overwintering larva is found within 
the unopened leaf-bud, its presence indicated by a 
small round frass lid over the entrance hole. For a 
short time the larva feed within the bud, chiefly upon 
the bud sheath. When it opens the young leaves are 
partially eaten and then the larva enters the new shoot. 
Thereafter the entire feeding life and pupal period is 
spent within the new growth. The larva makes a 
larval case during this generation. The life history of 
later summer generations is not known. 


18. Acrebasis aurorella Ely 
Fieure 149 


Acrobasis aurorella Ely, Proc. Ent. Soc. Washington, vol. 12, 
p. 67, 1910.—Forbes, Cornell Mem. 68, p. 616, 1923.— 
McDunnough, Check list, No. 6090, 1939. 

Close to and similar to stigmella but differing markedly 
in the ground color of the forewing. The sex-scaling is 
the same. ‘The upper surface of the head on the male 
is also ocherous white but much duller and less con- 
trasted against the dark gray thorax. Forewing with 
basal area blackish gray sharply contrasted against re- 
mainder of wing; median and outer areas of wing pale 
pinkish ocherous or pale gray with a pinkish overcast; 
subterminal line extremely faint; discal dots distinct 
but not strongly contrasted. Hind wings as in stigmella. 
Alar expanse, 19-22 mm. 

Typ Locatiry: Washington, D.C. (typein USNM). 

Foop prant: Unknown. 

Distrisution: Connecticut, Hast River (Aug.); New 
York, Ilion (Aug.); New Jersey, Montclair (June); 
Pennsylvania, New Brighton (Sept.); District of Colum- 
bia, Washington (June). Also two specimens from the 
Fernald Collection, without locality and labeled ‘‘demo- 
tella Grt.” 

19. Acrobasis peplifera Dyar 
Fieure 150 

Acrobasis peplifera Dyar, Ins. Inse. Menstr., vol. 13, p. 13, 
1925.—McDunnough, Check list, No. 6106, 1939 

Forewing dark gray with a vinous tint; basal area a 
dark wine red; scale ridge black, followed by a red line, 
then by a whitish line or triangle on inner margin, 
shading into black towards costa; discal dots obscured 
in the dark ground color, more or less confluent. Hind 
wings pale smoky fuscous, darker in the female than 
male; veins faintly outlined by dark scaling. Alar ex- 
panse, 13-17 mm. 

The male sex-scaling on underside is as follows: On 
forewing a rather wide black stripe on basal fifth of 
costa; on hind wing a black streak from base along top 
of cell, widening out to costa at its middle and putting 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 19 


out a thin black branch along lower vein of cell (this 
lower streak extending out as far as base of vein 2). 


Tyre tocauity: Millbrook, N. C. (type in USNM). 

Foop piants: Hickory, pecan. 

Distrisution: North Carolina, Elizabeth City (Aug.), 
Millbrook (Aug.), New Bern (Aug.), Tryon (Aug.); 
Illinois, Putnam County (July); Arkansas, Washington 
County (Aug.); Texas, Cuero (July); Georgia, Albany 
(June); Florida, Monticello. 

Doubtfully distinct from exsulella, but the names had 
better be kept separate until more is known about the 
biologies of the two color forms. 


20. Acrobasis exsulella (Zeller), new combination 
Figure 151 


Myelois exsulella Zeller, Isis von Oken, 1848, p. 868. 

Rhodophaea exsulella (Zeller) Ragonot, Ent. Amer., vol. 5, p. 114, 
1889; Monograph, pt. 1, p. 80, 1893.—Hulst, Phycitidae of 
N. Amer., p. 120, 1890.—McDunnough, Check list, No. 
6076, 1939. 

Acrobasis septentrionella Dyar, Ins. Insc. Menstr., vol. 13, p. 13, 
1925.—McDunnough, Check list, No. 6107, 1939. (New 
synonymy.) 

Similar to peplifera except: Averaging larger; basal 
area of forewing more decidedly and evenly reddish; 
outer area somewhat paler; scale ridge not blackish, 
red and concolorous with remainder of basal area, very 
weak. Alar expanse, 14-20 mm. 

Type LocauitiEs: North America [Georgia?] (ersulel- 
la, in Zool. Mus. Univ. Berlin) ; Florida (septentrionella, 
in USNM). 

Foop piants: Hickory, pecan. 

Distrisution: Florida (type, no other locality, 
Apr.), Orlando (Apr.); Texas, Brownwood (Mar.); 
North Carolina, Raleigh (May); Maryland, Plummers 
Isl. (June). 

The types of both ezsulella and septentrionella are 
females. The former is supposed to be in Berlin. 
The figure of it given by Ragonot (Monograph, pl. 5, 
fig. 19) is a very good match for Dyar’s species and I 
have no hesitation in synonymizing the latter. The 
scale ridge is present on forewing but could be easily 
overlooked, especially ona female that had been spread. 
Even on the unspread and unrubbed females in the 
National Collection it is not discernible except under 
considerable magnification. The structure is more 
prominent on the male. 

Our Texas specimens were reared from larvae feeding 
on the expanding buds of pecan. Dyar’s paratype from 
North Carolina was reared from hickory. We have no 
further information on the biology. The sex-scaling 
of the male is like that of peplifera, which will probably 
prove to be nothing more than a variety or color form 
of exsulella. 

21. Acrobasis angusella Grote 
Figure 152 


Acrobasis angusella Grote, North Amer. Ent., vol. 1, p. 51, 1880; 
Papilio, vol. 1, p. 14, 1880; Bull. U. S. Geol. Geogr. Surv. 
Terr., vol. 6, p. 590, 1882.—Ragonot, Monograph, pt. 1, 
p. 104, 1893.—Barnes and McDunnough, Contributions, 


vol. 2, p. 221, 1914.—Forbes, Cornell Mem. 68, p. 615, 
1923—McDunnough, Check list, No. 6091, 1939. 

Acrobasis eliella Dyar, Proc. Ent. Soc. Washington, vol. 10, p. 
43, 1908. 

Head and thorax whitish clay color, more or less 
shaded with pinkish or reddish suffusion (darker on 
females than on males). Forewing with basal area 
reddish and with some dusting of black scales on darker 
specimens; scale ridge black or black with an inter- 
mixture of red scales, followed outwardly by a reddish 
or reddish ocherous patch narrowing towards costa; 
pale antemedial line chiefly indicated on lower half of 
wing, obscure, followed on costa by a dark fuscous 
triangulate shade; median area gray with a diffused 
pale shade surrounding discal spots and extending to 
costa; discal dots separate, distinct, but not strongly 
contrasted against ground color; subterminal line 
distinct, denticulate, preceded by a narrow dark border 
and followed in terminal area by a broad reddish or 
reddish ocherous suffusion. Hind wing pale smoky 
fuscous. Alar expanse, 17-22 mm. 

Type tocauities: West Farms, N. Y. (angusella, 
in BM); East River, Conn. (eliella, in USNM). 

Foop puant: Hickory (larvae boring in the leaf 
stems). 

DistrisuTion: Unitep States: Massachusetts, North 
Adams (Aug.); New York, West Farms, Ilion (July, 
Aug.) ; New Jersey (June) ; Pennsylvania, New Brighton 
(May, Aug.); Maryland, Beltsville (May) ; Connecticut, 
East River (July, Aug., Sept.). Canapa: Ontario, 
Ottawa (July). 

Barnes and McDunnough (1914) were correct in their 
criticism of Dyar’s identification of angusella and in 
their reference of eliella to synonomy, but their descrip- 
tion of the male sex-scaling is at fault. It is more 
correctly described by Ragonot in his monograph. It 
consists of a very short black patch on costa of forewing; 
a long black streak along the upper vein of cell, expand- 
ing almost to the costal edge at middle and terminating 
well beyond the end of the cell, and from the base of 
this streak a second short black streak along lower line 
of cell for about half its length. On some males there 
are also a few black scales on vein 1c shortly beyond its 
base. 


22. Acrobasis demotella Grote 
Figure 153 


Acrobasis demotella Grote, Papilio, vol. 1, p. 14, 1881; Bull. U. S. 
Geol. Geogr. Surv. Terr., vol. 6, p. 590, 1882.—Hulst, 
Phycitidae of N. Amer., p. 122, 1890.—Ragonot, Mono- 
graph, pt. 1, p. 103, 1893.—Dyar, Proc. Ent. Soc. Washing- 
ton, vol. 10, p. 42, 1908.—Barnes and McDunnough, Con- 
tributions, vol. 2, p. 221, 1914.—Forbes, Cornell Mem. 68, 
p. 616, 1923.—McDunnough, Check list, No. 6092, 1939. 


Color and markings in general similar to those of 
angusella except: Central area of forewing a uniformly 
suffused dark grayish fuscous; basal area a paler red- 
dish shade without dark dusting except for some 
fuscous smudging of the scale ridge on inner margin; 
antemedial line more distinct, dull white and on most 
specimens completely indicated to costa; discal dots 


20 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


and subterminal line much obscured, the latter not 
denticulate, and followed by 2@ rather faint reddish 
shading, distinct only on dark unrubbed or unfaded 
specimens. Hind wing pale smoky fuscous. Alar 
expanse, 20-24 mm. 

Typ Locauity: West Farms, N. Y. (typein BM). 

Foop piant: Black walnut (larvae feeding in buds 
and stems). 

Distrisution: New Hampshire, Durham; New York, 
West Farms, Long Island; Pennsylvania, New Brighton 
(June); North Carolina, Black Mountain (June), Tryon 
(May); Jliinois, Chicago, Decatur (May), Putnam 
County (June); Missouri, St. Louis (June). 

The sex-scaling of the male is of the same type as 
that of angusella, differing as follows: On forewing the 
black costal streak is longer, extending to basal fifth of 
costa; the upper streak on hind wing is somewhat 
shorter, narrow at base, swelling to an oblong patch at 
middle; the streak along lower vein of cell is broader 
and longer, extending to the lower outer angle of cell. 

Hulst gives a description of the larva and life history 
as supplied him by Fernald. Nothing substantial has 
been added since then to our knowledge of the biology 
of the species. 


Genus Acrobasis, Species 23-40: A. latifasciella 
to A. tumidulella 


[Male: Apical process of gnathos trifurcate; forewing with 
Taised-scale ridge; without sex scaling.] 


23. Acrobasis latifasciella Dyar 
Figure 139 


Acrobasis latifasciella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 45, 1908.—Forbes, Cornell Mem. 68, p. 617, 1923.— 
MeDunnough, Check list, No. 6096, 1939. 

In color and maculation similar to angusella Grote; 
but without black sex-scaling on underside of male 
wings. Alar expanse, 18-21 mm. 

Type tocauity: New Brighton, Pa. (type in USNM). 

Foop puants: Hickory, walnut. 

DistripuTion: New York, Ilion (Sept.); Pennsyl- 
vania, New Brighton (Aug.); District of Columbia, 
Washington; Maryland, Plummers Isl. (Aug.); Illinois, 
Putnam County (June). 

Nothing is known of the biology except the food 
plants. 

24, Acrobasis irrubriella Hly 
Fiaure 154 


Acrobasis irrubriella Ely, Proc. Ent. Soc. Washington, vol. 10; 
p. 161, 1908.—Forbes, Cornell Mem. 68, p. 618, 1923.— 
MecDunnough, Check list, No. 6095, 1939. 

Color and maculation similar to those of wngusella 
and latifasciella except: Thorax and basal area of fore- 
wing showing little or no trace of reddish scaling; 
reddish ocherous band on outer border of scale ridge 
narrower and fainter; no reddish shading in terminal 
area beyond subterminal line. Alar expanse, 19-21 
mm. 

Male genitalia with apex of uncus more rounded than 


that of latifasciella—at best, a character of doubtful 
specific value. 

TypELocatity: Hast River, Conn. (typein USNM). 

Foop puant: Unknown, probably Carya (hickory or 
walnut). 

Distrisution: Connecticut, East River (July); Indi- 
ana, Mineral Springs (Aug.). 


25. Acrobasis normella Dyar 
Fieure 155 


Acrobasis normella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 46, 1908.—Forbes, Cornell Mem. 68, p. 617, 1923.— 
McDunnough, Check list, No. 6097, 1989. 

Similar to wrrubriella except: Averages smaller; fore- 
wing of a more even glossy texture; more white dusting 
on basal area, giving it a more decidedly pale gray ap- 
pearance; the white dusting from outer half of costa to 
and surrounding the discal dots also a trifle stronger. 
Alar expanse, 17-19 mm. 

Typ Locatity: Kast River, Conn. (type in USNM). 

Foop pirant: Presumably hickory. 

Represented in the National Collection by 15 speci- 
mens from the type locality. In addition, there are 
before me two smaller (15 mm.), darker, more suffused 
specimens (male and female) from the Barnes collection 
that McDunnough had identified as irrubriella. They 
were reared from hickory (June) and are, I believe, only 
a color form of normella. The male genitalia (fig. 1552) 
show a striking departure in the assymetrical and greatly 
reduced lateral elements of the trifurcate apical projec- 
tion of gnathos, but apparently this is the result of a 
deformation of the organ in this particular specimen. 


26. Acrobasis malipennella Dyar 
Ficure 156 


Acrobasis malipennella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 47, 1908.—Forbes, Cornell Mem. 68, p. 618, 1923.— 
McDunnough, Check list, No. 6093, 1939. 

Similar to the species following (dyarella) in color and 
markings except that the darker areas of forewing are 
dark fuscous and lack the intense red suffusion charac- 
teristic of the latter. The structural differences noted 
by Dyar (the broader, shorter forewing, its broader 
cell, the close approximation of veins 2 and 3, different 
on each forewing, and the connate condition of 4 and 5 
and their remoteness from 3) are due to deformation. 
The male type never matured properly and the fore- 
wings are not fully developed. I believe the specimen 
is nothing but a color form of dyarella and a freak at 
that. The male genitalia are very close, differing only 
in the somewhat more broadly rounded apex of gnathos, 
a, difference of no specific significance in this group. It 
is a pity Dyar ever described it and a still greater pity 
that we cannot ignore his name; for it will probably 
have to replace dyarella Ely which was based on normal 
specimens and is represented by types in good condition. 

Typr Locauity: East River, Conn. (Aug.; type in 
USNM). 

Foop prant: Unknown. 

Known only from the unique type. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE a1 


27. Acrobasis dyarella Ely 
Fieurs 157 
Acrobasis dyarella Ely, Proc. Ent. Soc. Washington, vol. 12, p. 67, 
1910.—McDunnough, Check list, No. 6099, 1939. 

Head yellow gray. Thorax much suffused with red 
scaling. Basal area of forewing red dusted with whitish 
towards costa; scale ridge black, followed outwardly by 
a broad red band which extends and diffuses outwardly 
towards costa, obscuring and partially obliterating the 
blackish costal triangle; red scaling generally scattered 
over lower median area of wing; some obscure whitish 
dusting on median costal area and about the small, 
separated discal spots; subterminal line faint, bordered 
inwardly and outwardly, except on costa, by reddish 
lines; terminal area and dark markings otherwise, dark 
gray. Hind wing glossy, pale smoky fuscous. Alar 
expanse, 19-20 mm. 

Type Locauity: East River, Conn. (type in USNM). 

Foop pirant: Unknown. 

Represented only by the male type (Aug.) and female 
paratype (Sept.). 

28. Acrobasis ostryella Ely 
Fiaure 158 
Acrobasis ostryella Ely, Ins. Insc. Menstr., vol. 1, p. 54, 1913.— 
McDunnough, Check list, No. 6084, 1939. 

Similar to dyarella except: Reddish color more gener- 
ally diffused over outer areas of forewing, not forming 
a strongly accented band following the scale ridge, and 
of a purplish red shade; the coastal triangle and scale 
ridge contrastingly black. Hind wing dark smoky 
fuscous. Alar expanse, 15-18 mm. 

Type Locauity: East River, Conn. (type in USNM). 

Foop Piants: Ostrya virginiana, Carpinus. 

Distrisution: Unirep States: Connecticut, East 
River (July). Canapa: Ontario, South March (June). 


29. Acrobasis secundella Ely 
Figure 159 
Acrobasis secundella Ely, Ins. Inse. Menstr., vol. 1, p. 55, 1913.— 
McDunnough, Check list, No. 6083, 1939. 

Doubtfully distinct from ostryella. The holotype 
and one other reared male from the type locality and 
two Canadian specimens before me are darker and a 
nearly uniformly suffused purplish, with a pale dusting 
on basal area of forewing, and about the discal spots a 
rather faint and pale gray rather than white. However, 
other reared specimens from hazel are a perfect match 
for the type of ostryella. Alar expanse, 15-18 mm. 

The male genitalia show a trifling difference in the 
length of the lateral projections of the apical projection 
of gnathos. Ely states that the larval case of secundella 
is longer and more slender than that of ostryella; but 
the life history needs further investigation before any 
such difference can be evaluated. 

Type tocauity: East River, Conn. (type in USNM). 

Foop piant: Corylus. 

Distrisution: Unitep States: Connecticut, East 
River (July). 

Canapa: Ontario, Merivale (June). 


30. Acrobasis coryliella Dyar 
Ficure 160 


Acrobasis coryliella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 47, 1908.—Ely, Ins. Inse. Menstr., vol. 1, p. 53, 1913.— 
Forbes, Cornell Mem. 68, p. 618, 1923.—MeDunnough, 
Check list, No. 6098, 1939. 

Forewing a dull, ratber lusterless gray, paler in the 
central area and without any reddish or purplish 
shading; following scale ridge on inner margin a faintly 
ocherous patch obscured by gray scaling; the scale ridge 
and other dark markings blackish, but the usual black 
triangle on costa following antemedial line replaced by 
a narrow line; the usual transverse dark shade from 
inner upper edge of subterminal line to middle of lower 
margin; subterminal line bordered inwardly by a narrow 
blackish line; discal dots normally confluent, forming a 
curved line along discocellular vein of cell. Hind wing 
pale smoky fuscous; darker on female. Alar expanse, 
17-20 mm. 

Type Locauity: Unspecified [New York?] (type in 
USN). 

Foop piant: Corylus. 

Distrisution: New York; Connecticut, East River 
(July, Aug.); Massachusetts, Newton Highlands; Jili- 
bac Decatur (June, July), Putnam County (June, 
July). 

In addition to a long series reared from hazel at East 
River, Conn., there are three specimens from the 
Fernald and Brooklyn Museum Collections (one male 
and two females from [llinois) labeled ‘‘Acrobasis 
hebescella,” the two females labeled ‘type’; and one 
male from Decatur, Ill., which McDunnough had 
tentatively identified as A. sylviella Ely. 

The species is easy to recognize from its rather uni- 
form gray shade and strongly contrasted, short, black 
scale ridge. 

31. Acrobasis hebescella Hulst 


Acrobasis hebescella Hulst, Phycitidae of N. Amer., p. 126, 1890.— 
Ragonot, Monograph, pt. 1, p. 109, 1893.—Barnes and 
MecDunnough, Contributions, vol. 3, p. 194, 1916.— 
MeDunnough, Check list, No. 6085, 1939. 

The only authentic representation of this species is the 
female type, which is in very poor condition but ap- 
parently does not, or did not originally, differ in any 
significant detail from coryliella Dyar except as to its 
host. It was reared from a cocoon found onoak. This 
may or may not be significant. The name is just 
another of those that must wait for clarification until 
someone shall make a careful and more thorough 
study of the life histories of the various Acrobasis 
species. Alar expanse, 16.5 mm. 

TypE tLocauiry: ‘Jersey pines, June” 
AMNH, ex Rutgers). 

Foop pLant: Oak. 


(type in 


32. Acrobasis cirroferella Hulst 


Acrobasis cirroferella Hulst, Canadian Ent., vol. 24, p. 60, 1892.— 
MeDunnough, Check list, No. 6109, 1939. 


The type is a male without abdomen. There is no 
sex-scaling. Close to coryliella, but with dark areas of 


22 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


forewing more brownish gray and the whitish areas 
more strongly contrasted; central costal area dis- 
tinctly white; dark outer margin of antemedial line 
brown, narrow on costa; costa before it white, entire 
basal area having some white dusting; discal dots 
distinct, separate. Hind wing pale fuscous. Alar ex- 
panse, 18 mm. 

TypE Locantiry: Austin, Tex. (type in AMNH, ex 
Rutgers). 

Foop prant: Unknown. 

The above description is drawn from the type, which 
is worn and faded. I have seen nothing that exactly 
matches it. 


33. Acrobasis cunulae Dyar and Heinrich 
Fieure 162 


Acrobasis cunulae Dyar and Heinrich, Proc. Ent. Soc. Washing- 
ton, vol. 31, p. 37, 1929.—McDunnough, Check list, No. 
6102, 1939. 

Forewing pale slate gray; basal area paler; midcostal 
area with some faint pale dusting, especially about the 
discal spots; scale ridge weak and little or not at all 
darker than the ground color of the wing; subterminal 
line distinct, with a narrow, dentate, dark inner border, 
neither the pale line itself nor its dark border strongly 
contrasted; discal dots blackish, separated and rather 
conspicuous. Hind wing smoky fuscous. Alar ex- 
panse, 20-24 mm. 

The male genitalia have what appears to be a dis- 
tinguishing specific character in the decidedly broad- 
ened lateral elements of the apical projection of gnathos. 

TyvE tocatiry: Mobile, Ala. (type in USNM). 

Foop piant: Pecan. 

Distrizution: Florida, Monticello (May); Georgia, 
Cairo (May), DeWitt (May); Alabama, Auburn (May), 
Mobile (May); Mississippi, Wiggins (May). 

Close to but apparently distinct from caryivorella. 


34. Acrobasis caryivorella Ragonot 
Fiaure 161 


Acrobasis caryivorella Ragonot, N. Amer. Phycitidae, p. 4, 1887; 
Monograph, pt. 1, p. 108, 1893.—Hulst, U. S. Nat. Mus, 
Bull. 52, p. 419, 1903.—Hill, Florida Ent., vol. 21, p. 12, 
1938.—McDunnough, Check list, No. 6087, 1989.—Craig- 
head, U. S. Dep. Agr. Misc. Publ. 657, p. 450, 1950. 

Acrobasis caryaevorella Hulst, Phycitidae of N. Amer., p. 121, 
1890 (misspelling). 

Acrobasis conjivorella Hulst, in J. B. Smith, List of the Lepi- 
doptera of Boreal America, No. 4262, 1891 (misspelling). 

Acrobasis caryae Dyar (not Grote), Proc. Ent. Soc. Washington, 
vol. 10, p. 46, 1908. 


Forewing dark bluish gray, nearly black; basal area 
towards costa (above the scale ridge) and a small tri- 
angular area on costa adjacent to subterminal line 
powdered with grayish white; scale ridge black, on 
some specimens bordered outwardly by a faint ocherous 
red patch (especially on specimens reared from hickory) ; 
subterminal line pale gray, faint; discal dots distinct, 
separate or confluent. Hind wing smoky white to 
smoky fuscous, darker on female than on male. Alar 
expanse, 19-24 mm. 


TypE Locauiry: Missouri (type in Paris Mus.). 

Foop puants: Hickory, walnut, pecan. Larvae 
boring in the buds and new growth of the stems. 

Distrisution: Massachusetts, Melrose (June, July); 
Maryland, Beltsville (May, July, Aug.), Hyattsville 
(June), Prince Georges County, (June); North Carolina, 
“N. Car. Dept. Agr.”; South Carolina, Summerton 
(May); Georgia, Albany (May), Barnesville (May, 
June) ; Florida, Monticello (May, July), Orlando (Apr.); 
Mississippi, Biloxi (Aug.), State College; Missouri; 
Texas, Austin (Aug.), Brownwood (May, June), 
Menard (June), Victoria, Waco (Apr., May, June); 
New Mexico, Carlsbad (Aug.). 

The species is of some importance in the Gulf States 
as an enemy of pecan and the name caryworella has 
appeared several times in economic publications but 
nearly always wrongly applied to specimens of caryae 
Grote. Specimens of the true caryivorella have also 
been identified as caryae on the basis of Dyar’s (1908) 
misapplication of the two names. 


35. Acrobasis comacornella (Hulst), new combination 
Figure 136 


Acrocaula comacornella Hulst, Canadian Ent., vol. 32, p. 170, 
1900.—McDunnough, Check list, No. 6078, 1939. 

Forewing with dark areas glossy, purplish brown; 
extreme base of wing dark, followed by a rather narrow 
subbasal whitish area; median costal area narrowly 
whitish; outer area uniformly dark; subterminal line 
obscure, not bordered by darker lines; discal dots 
distinct, dark brown, the lower dot twice the size of 
the upper; a little red on the antemedial line towards 
costa. Hind wing whitish with a faint ocherous 
fuscous tint; rather glossy; veins very faintly darkened; 
a narrow dark line along termen. Alar expanse, 17 mm. 

Detail of male genitalia figured from type. 

Type tocatiry: Blanco County, Tex. (type in 
AMNH, ex Rutgers). 

Foop puant: Unknown. 

The male type is obviously an Acrobasis without sex- 
scaling. It resembles very much a specimen of caryi- 
vorella (in USNM) from Victoria, Tex., except that the 
hind wing is paler, the subterminal line less distinct, 
and the discal dots larger and more contrasted. I sus- 
pect that it is nothing more than a variety of caryi- 
vorella. 

36. Acrobasis betulella Hulst 


Ficurs 164 


Acrobasis betulella Hulst, Phycitidae of N. Amer., p. 125, 1890.— 
Ragonot, Monograph, pt. 1, p. 107, 1893.—Dyar, Proc. 
Ent. Soc. Washington, vol. 10, p. 47, 1908.—Forbes, Cornell 
Mem. 68, p. 618, 1923.—McDunnough, Check list, No. 
6101, 1939. 

Forewing dark gray faintly tinted with reddish violet 
and with white dusting on basal area and forming a 
triangulate patch from costa before subterminal line, 
the white dusting faint (less contrasted than on caryi- 
vorella); scale ridge black, without any red bordering 
patch or bar; discal dots at end of cell distinct, separate; 
antemedial line obscure, almost obsolete; subterminal 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 23 


line faint. 
20-24 mm. 

Male genitalia figured from specimen from the origi- 
nal Hulst series in the National Collection (bearing a 
Hulst ‘‘type”’ label and presumably a paratype). 

Typr Locauity: New York (type, 2, in AMNH, ex 
Rutgers). 

Foop prant: Betula. 

Distrisution: Unirep States: Maine (no further 
locality, July), Sebec Lake (July); New Hampshire, 
Center Harbor (July), Hampton (June); Massachusetts, 
Amherst (June, July), Melrose; Connecticut, Hast River 
(July); New York (no further locality, July); Colorado, 
Platt Canon (July); California, Siskiyou County. 
Canapva: Ontario, Trenton. 

There is little or nothing to separate collected speci- 
mens of betulella from comtoniella or rubrifasciella except 
the complete absence of any reddish outer border to the 
scale ridge on forewing (a distinction that does not hold 
for all specimens of rubrifasciella) and a more glossy 
sheen on the specimens of comptoniella and rubrifasciella 
(a comparative difference that is displayed only when 
series of the three species are seen side by side). The 
thing that really distinguishes betulella is its host plant, 
Betula. It also differs from the other species of Ameri- 
can Acrobasis with the scale ridge on forewing in that it 
has been found in the Rocky Mountain region and in 
areas west thereof. The Colorado and California 
records are from the specimens in the National Collec- 
tion mentioned by Dyar. I have seen no later collec- 
tions from any area west of the Rockies. 


Hind wing smoky fuscous. Alar expanse, 


37. Acrobasis rubrifasciella Packard 
Figure 165 


Acrobasis rubrifasciella Packard, Ann. Lyc. Nat. Hist. New 
York, vol. 10, p. 267, 1873.—Grote, Bull. U. S. Geol. Geog. 
Surv. Terr., vol. 4, p. 693, 1878.—Hulst, Phycitidae of N. 
Amer., p. 124, 1890.—Ragonot, Monograph, pt. 1, p. 106, 
1893.—Forbes, Cornell Mem. 68, p. 617, 1923.—McDun- 
nough, Canadian Ent., vol. 65, p. 206, 1933; Check list, 
No. 6103, 1939. 

Phycis rubifasciella (Packard) Beutenmuller, Canadian Ent., 
vol. 22, p. 16, 1890 (spelling! and larva). 

Acrobasis alnella McDunnough, Canadian Ent., vol. 54, p. 36, 
1922. 


Similar to betulella except that normally there is a 
faint, but distinguishable, wine-red bar outwardly 
bordering the scale ridge on forewing and that, in 
series, the forewing surface has a more glossy appear- 
ance. Neither of these differences holds for all speci- 
mens; nor are the genitalic differences noted by Mc- 
Dunnough (1933), the width of the lateral flanges of 
the apical projection of gnathos, reliable as a specific 
character. Alar expanse, 20-24 mm. 

TypE Locatities: Orono, Maine (rubrifasciella, in 
MCZ); Ottawa, Canada (alnella, Canadian Nat. Coll.). 

Foop piant: Alnus. 

Distrievution: Unitep States: Maine, Orono (June); 
New Hampshire, Durham; Massachusetts, Amherst 
(June) ; Connecticut, Hast River (July); New York, Cats- 
kill Mts.; North Carolina, Black Mts. Canapa: On- 


tario, Ottawa (July); Quebec, Meach Lake (July); Nova 
Scotia, Truro (July, Aug.). 


38. Acrobasis comptoniella Hulst 


Acrobasis comptoniella Hulst, Phycitidae of N. Amer., p. 125, 
1890.—Ragonot, Monograph, pt. 1, p. 108, 1893.—Dyar, 
Proc. Ent. Soc. Wash., vol. 10, p. 46, 1908.—Forbes, Cornell 
Mem, 68, p. 618, 1928.—MecDunnough, Check list, No. 6104, 
1939.—Craighead, U. S. Dept. Agr. Misc. Pub. 657, p. 
450, 1950. 

Superficially like rubrifasciella except wine-red bar 
bordering scale ridge of forewing always present and 
of a more intense and darker shade. Hind wing gen- 
erally darker. Alar expanse, 21-25 mm. 


Type Locatity: Long Island, N. Y. (type, 9, in 
AMNH, ex Rutgers; paratype, 9, from type locality 
in USNM). 

Foop piants: Comptonia and Myrica. 

Disrripution: Unirep Srarss: Maine, Bar Harbor 
(July), Kennebunk (July), Mount Desert (July); New 
Hampshire, Center Harbor, Durham; Massachusetts, 
Melrose (June), North Saugus; Connecticut, East River 
(July); New York, Long Island; New Jersey, Bergenfield, 
New Lisbon (June); Michigan, Dickinson County. 
Canapa: Ontario, Trenton. 

The best way to separate comptoniella from the two 
preceding species is by rearing from their respective 
food plants. 


39. Acrobasis myricella Barnes and McDunnough 
Fiaurs 163 


Acrobasis comptoniella Grossbeck (not Hulst), Bull. Amer. Mus. 
Nat. Hist., vol. 37, p. 129, 1917. 
Acrobasis myricella Barnes and McDunnough, Contributions, 
pee p. 221, 1917.—McDunnough, Check list, No. 6105, 
Close to comptoniella Hulst, but superficially quite 
different; smaller, white dusting in pale (basal and 
central-costal) areas much denser and more contrasted; 
dark areas blackish gray with very faint purplish over- 
tint ; reddish bar bordering black scale ridge narrow and 
inconspicuous; subterminal line distinct, white. Alar 
expanse, 17-19 mm. 
Typr Locatity: Fort Myers, Fla. (Apr., May; type 
in USNM). 
Foop puant: Myrica. 


Distrisution: Florida. 

Except for the type series from the type locality I 
have seen only one specimen (a female from Royal Palm 
State Park, Fla., Apr. 5, 1929, F. M. Jones, collector) 
that can be definitely assigned to the name myricella. 
We have, however, in the National Collection a series 
of males and females reared by Chas. R. Ely at East 
River, Conn., from Myrica cerifera, that are inter- 
mediate between myricella and typical comptoniella 
(reared from Comptonia), like the former in size and in 
the intensity and extent of the white dusting on fore- 
wing, but with the subterminal line obscure as in 
comptoniella. I think they are only a variety of 
comptoniella. Indeed, myricella may prove to be only 


94 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


a race of Hulst’s species; but with our present knowl- 
edge we must retain it as a distinct species. © 


40. Aecrobasis tumidulella (Ragonot), new combination 
Fieure 645 


cai tumidulella Ragonot, N. Amer. Phycitidae, p. 13, 

1887. 

Seneca tumidulella (Ragonot) Hulst, Phycitidae of N. Amer., p. 
178, 1890. 

Hyphantidium tumidulellum (Ragonot) Hampson, in Ragonot, 
Monograph, pt. 2, p. 74, 1901.—McDunnough, Check list, 
No. 6321, 1939. 

This species was based on a single female with the 
habitus and raised-scale ridge of an Acrobasis but with 
vein 4 absent from hind wing. Bourgogne informs me 
that the venation is alike on both hind wings. On the 
strength of this venation the species was referred to 
Group II of the Phycitinae and made the type of Hulst’s 
Seneca. However, I am firmly convinced that the speci- 
men is nothing but an Acrobasis with abnormal venation, 
another of those freaks that turn up all too frequently in 
the Phycitidae. Ihave examined the female genitalia of 
the type (figured here) and can find nothing to distin- 
guish them from those of caryivorella. I suspect that 
tumidulella is nothing more than an abnormal specimen 
of caryivorella; but we shall have to await final disposi- 
tion of the name until a similar freak male is recovered 
from the type locality. 


Typr Locatity: Florida (type in Paris Mus.). 
Foop prant: Unknown. 


3. Genus Rhodophaea Guénée 


Rhodophaea Guénée, Ann. Soc. Ent. France, ser. 2, vol, 3, p. 312, 
1845; Europaeorum Microlepidopterorum index methodi- 
cus... , p. 74, 1845.—Ragonot, Ent. Monthly Mag., vol. 
22, p. 19, 1885; Monograph, pt. 1, p. 63, 68, 1898. (Type 
of genus; Phycis advenella Zincken; figs. 166, 649.) 

Characters of Acrobasis except: Male antenna simple, 
basal segment cylindrical, no sinus in base of shaft; 
forewing always smooth; vein 2 of forewing from cell 
before lower outer angle, but somewhat nearer the 
angle than in Acrobasis; male genitalia with apical 
process of gnathos an elongate hook (partially divided 
on advenella); eighth abdominal segment of male simple 
or (on advenella) with midventral hair tuft. 

This genus is distinguished from the smooth-winged 
species of Acrobasis only by its simple male antenna. 
Our two American species do not go any too well with 
advenella; the European type of the genus, differing in 
having an undivided apical projection from gnathos 
and simple eighth abdominal segment. However, in 
these characters they agree with otber obviously con- 
generic European species, marmorea (Haworth), lega- 
tella (Hiibner), suavella (Zincken). R. advenella has a 
somewhat differently shaped transtilla from caliginella, 
supposita, and the three aforementioned Huropean 
species. In all of these the terminal margin of trans- 
tilla is more or less indented (as in Acrobasis) while in 
advenella it is rather deeply U-shaped. 

None of the American species that hitherto have been 


listed under Rhodophaea belongs there. 


They have 
entirely different genitalia. . 


41. Rhodophaea caliginella (Hulst), new combination 
Fieure 647 


Nephopteryx caliginella Hulst, Ent. Amer., vol. 3, p. 131, 1887; 
vol. 5, p. 156, 1889. 

Mineola caliginella (Hulst), Phycitidae of N. Amer., p, 128, 
1890; U. S. Nat. Mus. Bull. 52, p. 419, 1902.— Barnes and 
McDunnough, Contributions, vol. 4, p. 174, 1918.— 
McDunnough, Check list, No. 6113, 1939. 

Acrobasis caliginella (Hulst) Ragonot, Monograph, pt. 1, p. 115, 
1893. 


Myelois caliginoidella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 33, 1905.—McDunnough, Check list, No. 6072, 1939. 
(New synonymy.) 

Similar in color and markings to Acrobasis comptella 
Ragonot except that the narrow black line outwardly 
bordering the whitish basal area of forewing does not 
extend all the way to inner margin. This slight dif- 
ference in maculation seems to be constant and will 
distinguish the females of the two species which, other- 
wise, are difficult to tell apart. Alar expanse, 18-25 
mm. 

TypE LocaLities: Arizona (caliginella, in AMNH, 
ex Rutgers); Santa Clara, Calif. (caliginoidella, in 
USNM). 

Foop puLant: Scrub oak. This host record from 
reared specimens in National Collection received from 
Commander Dammers, Riverside, Calif., June 1938. 

Distripurion: California, Alma (Aug.), Atascadero 
(July), Los Angeles County (July), Riverside (June), 
San Diego (May, June, July, Aug.), Santa Clara; 
Arizona. 

In addition to the female type in the Rutgers Col- 
lection there is also a female from Arizona (‘‘7810”) in 
the National Collection bearing Hulst’s “‘type’’ label. 
This specimen was originally in the Fernald Collection. 
A female from California donated by Hulst. to the 
Brooklyn Museum Collection and twice labeled ‘‘Acro- 
basis comptelia” in his and Ragonot’s handwriting is 
also in the National Museum. This specimen, except 
that it lacks an abdomen, is in good condition. It is 
certainly caliginella and presumably was responsible 
for Hulst’s synonymizing of caliginella and compiella. 
The males of caliginella have hitherto been known as 
caliginoidella Dyar. Hulst evidently never saw a male 
of his species. 


42. Rhodophaea supposita (Heinrich), new combination 
Fiaures 167, 648 


Mineola supposita Heinrich, Proc. Ent. Soe. Washington, vol. 
42, p. 33, 1940. 

Forewing very dark grayish fuscous with a powder- 
ing of white scales on basal and midcostal areas and 
very faintly in the area bordering terman; antemedial 
line narrow, slanting from inner third of costa to just 
before middle of inner margin, slightly notched at 
vein 1b, pale ashy gray bordered inwardly from top of 
cell to inner margin by a dull red triangular patch which 
has an obscure, straight, blackish line along its inner 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 25 


edge; inner margin from base to antemedial line nar- 
rowly bordered by reddish scales; subterminal line 
narrow, slightly outcurved between vein 6 and lower 
fold, pale gray, inwardly bordered by a narrow black 
line; a blackish fuscous patch outwardly bordering the 
antemedial line from costa to cell; a similar dark shade 
on costa near apex; these blackish patches shading into 
the dark central area of wing; black discal dots at end 
of cell distinct and separate; some obscure dull red 
shading in terminal area toward tornus; along termen 
a narrow discontinuous black line. Hind wing pale 
smoky fuscous with veins, terminal margin, and apical 
area darker. Alar expanse, 16-20 mm. 

Male genitalia similar to those of caliginella except 
transtilla broader at apex and arms of anellus stouter. 
Female genitalia differing from those of caliginella 
chiefly in that there are no patches of small scobina- 
tions in bursa near its junction with ductus bursae. 

TypE .Locauity.: Vancouver, British Columbia 
(type in Canadian Nat. Coll.). 

Foop puant: Cotoneaster. 

Known so far only from the type series from Van- 
couver. It is distinguished from caliginella chiefly by 
its generally darker color. 


4, Genus Trachycera Ragonot 


Trachycera Ragonot, Monograph, pt. 1, p. 2, 1893. (Type of 


genus, Rhodophaea pallicornella Ragonot.) 

This genus is close to Rhodophaea, being distinguished 
from it chiefly by male characters. The male of 
pallicornella from which these were drawn is apparently 
lost. Clarke was unable to locate it at Paris, where it 
should have been; and, as no other males agreeing with 
Ragonot’s description or figure (Monograph, pl. 5, 
fig. 20) are available, we are unable to check his charac- 
ters. Ragonot separates Trachycera from Rhodophaea 
widely in his generic key (Monograph, pt. 1, pp. xliii 
and xliv) on the basis of the trifid or bifid condition of 
the median vein of hind wing. This is an error, how- 
ever, for the true Rhodophaea species are no more bifid 
than is Trachycera. The female of pallicornella has 
essentially the same venation as the type of Rhodophaea 
(advenella). 

The distinguishing male characters given by Ragonot 
are: Serratiform male antenna; very short labial palpus 
(scarcely reaching to middle of face); and minute maxil- 
lary palpus. 

The female has a pair of small signa in the bursa 
copulatrix, developed as granulate cups (as in Davara). 


43. Trachycera pallicornella (Ragonot) 
Fiaure 650 
Rhodophaea pallicornella Ragonot, N. Amer. Phycitidae, p. ,3 
1887.—Hulst, Phycitidae of N. Amer., p. 119, 1890. 
Trachycera pallicornella (Ragonot), Monograph, pt. 1, p. 2, 
1893.—McDunnough, Check list, No. 6057, 1939. 

The holotype of pallicornella is a female labeled in 
Ragonot’s handwriting ‘‘ty. or. Pl. V, fig. 20.’ Accord- 
ing to Clarke the Ragonot figure represents it accu- 

300329—56——3 


rately. Unfortunately it is a mended specimen and 
the glued-on abdomen is spurious. Its genitalia are 
pyraustine rather then phycitid. 


I have before me a female which is an exact match for 
Ragonot’s figure. It is somewhat smaller (15 mm.) 
than the type (“19 mm.”) but this difference is easily 
within the normal range for many species of medium- 
sized Phycitinae. It was collected at Devils River, 
Tex. (May). The genitalia are figured from this 
specimen. 

Forewing pale gray with some blackish dusting on 
base, especially on base of costa; a faint purplish gray 
shade on lower part of postmedian area; antemedial 
band rather broad, red narrowly lined with black on 
inner and outer sides, the back outer margin somewhat 
widened at costa; subterminal line narrow, nearly 
vertical, with an outward bulge between vein 6 and 
lower fold, whitish, bordered inwardly by a narrow 
black line and by a black outer patch at costa near apex; 
discal and terminal dots obsolete. Hind wing dull 
whitish with a faint yellow tint and shading to pale 
fuscous towards apex. Alar expanse, 15-19 mm. 


TypPE Locatity: Texas (type in Paris Mus.). 
Foop piant: Unknown. 


5. Anabasis, new genus 


Typx or Genus: Myelois ochrodesma Zellar. 


Tongue well developed. Antenna pubescent; on 
male, basal segment enlarged and angulate (as in 
Acrobasis), shaft simple. Labial palpus upturned 
reaching to vertex (slightly longer on female than on 
male). Maxillary palpus rather broadly scaled. Fore- 
wing with a transverse, antemedian ridge of raised 
scales; 11 veins; vein 2 from before but near lower 
outer angle of cell; 3 from the angle, approximate to 2 
at base, nearer to 2 than to 4; 4 and 5 closely approxi- 
mate for some distance from base; 6 from below upper 
angle of cell, slightly bent towards base; 10 from the 
cell; male without costal fold. Hind wing with vein 2 
from well before lower outer angle of cell; 3 from before 
but very near the angle; 4 and 5 from the angle, closely 
contiguous (or more or less anastomosed) for about 
half their lengths; 7 and 8 contiguous or weakly anasto- 
mosed for some distance beyond cell; cell one-third 
the length of wing; discocellular vein curved. Eighth 
abdominal segment of male with ventral hair tuft and 
a pair of modified, ventrolateral tufts. 

Genitalic characters as in Acrobasis except: Harpe 
with a transverse, sclerotized ridge from base of costa 
to lower outer angle of sacculus; a cluster of modified 
scales on outer edge of inner margin in the angle be- 
tween sacculus and cucullus; terminal margin of 
vinculum more rounded. (These may be only specific 
characters.) 

A development from and quite close to Acrobasis, 
which it replaces in tropical America; distinguished 
from that genus chiefly by shorter cell and the con- 
tiguous position of veins 4 and 5 of hind wing. Except 


26 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


on denuded wings under strong magnification they 
appear to be stalked for half their lengths. Contains 
one tropical American species. 


44. Anabasis ochrodesma (Zeller), new combination 
Ficures 168, 652 


Myelois ochrodesma Zeller, Horae Soc. Ent. Rossicae, vol. 16, 
p. 209, 1881. 

Piesmopoda ochrodesma (Zeller) Ragonot, Monograph, pt. 1, 
p. 165, 1893. 

Acrobasis crassisquamella Hampson, in Ragonot, Monograph, 
pt. 2, p. 520, 1901 (mew synonymy). 

Forewing grayish brown finely powdered with 
blackish scales and with a faint rosy suffusion; ante- 
medial line oblique, narrow, obscured by a heavy whit- 
ish ocherous ridge of raised scales on its inner margin 
and bordered outwardly by a narrow black line; sub- 
terminal line obscure, narrow, when distinguishable, 
sinuate, ocherous white, bordered inwardly by an 
obscure, broken, black line; discal and terminal dots 
obsolete. Hind wing smoky white; veins and a narrow 
shade along termen, fuscous. Alar expanse, 13-16 mm. 

Genitalia as given for the genus; bursa of female with 
small signum. 

TypE LocaLities: Honda, Colombia (ochrodesma, in 
BM); Teapa, Tabasco, México (crassisquamella, in BM). 

Foop piants: Cassia alata, Cassia nodosa, Cassia tora 
(U. S. Dep. Agr. Florida rearings; larva a leaf-folder), 
Sciacassia siamea. 

Distrisution: Unitep Stratis: Florida, Coconut 
Grove (May), Miami (May), St. Petersburg (June). 
Puxrrto Rico: Bayamén (Sept.), Coamo Springs (Apr.), 
Mayagtiez (Jan.), Puerto Real (Vieques Isl., Apr.), Rio 
Piedras (Feb.). Virein Istanps: Kingshill (St. Croix, 
Oct., Nov., Dec.). Cusa: Santiago de las Vegas 
(Havana, Dec.). Grenada. JAMAICA. TRINIDAD: 
Fyzabad (Feb.), Tunapuna (Apr.). México: Tabasco, 
Teapa. Guatemana: Quirigué (May). Panam: 
Corozal (July), Porto Bello (Apr.). Conomspra: Honda. 

An easily recognized tropical American species whose 
range has been extended into southern Florida. 


6. Genus Mildrixia Dyar 


Mildrizia Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 405, 1914. 
(Type of genus; Mildrixia constitutionella Dyar.) 

Tongue well developed. Antenna of male strongly 
ciliate, the cilia over three times longer than width of 
segments; basal segment elongate, subtubular, flat- 
tened and broadening towards apex; first segment of 
shaft swollen and with a short spine and scale tuft from 
inner upper angle (this with the rough scaling of basal 
segment gives the latter when fully scaled the appear- 
ance of the triangulate first segment of Acrobasis) ; 
antenna of female simple and very weakly pubescent. 
Labial palpus obliquely ascending, reaching to slightly 
above vertex; moderately rough scaled beneath; third 
segment acuminate, about two-thirds the length of 
second. Maxillary palpus moderately large, squamous. 
Forewing narrowly elongate, with transverse, ante- 


medial ridge of raised scales; 11 veins; vein 2 from 
before, but near lower outer angle of cell; 3 from the 
angle, approximate to 2 at base and for a short distance 
beyond; 4 and 5 connate or very shortly stalked; 6 
from very close to upper angle of cell, closely approxi- 
mate to 8 at base, nearly straight (very slightly bent 
towards base); 10 from the cell, closely approximate to 
the stalk of 8-9 for some distance from cell; male with- 
out costal fold. Hind wing with vein 2 from before 
but close to lower outer angle of cell; 3 contiguous with 
the stalk of 4-5 for some distance from angle, on 
undenuded wings appears stalked with 4-5; 4 and 5 
stalked for more than half their lengths; 7 and 8 
contiguous or weakly anastomosed for a short distance 
beyond cell; cell one-third the length of wing; disco- 
cellular vein oblique. Eighth abdominal segment of 
male with broad ventral hair tuft. 

Male genitalia of the old world Acrobasis type 
except: Uncus subtriangulate, its apex rather broadly 
rounded; transtilla terminating posteriorly in a U- 
shaped projection with elongate, slender, widely spaced 
and divergent arms; vinculum longer than broad, evenly 
tapering to bluntly pointed terminal margin; anellus 
an elongate, semitubular plate with short lateral lobes 
near base; penis armed with a short, sclerotized plate 
and numerous sclerotized wrinklings. 

Female genitalia of the Acrobasis type but without 
any sclerotized plate or plates at genital opening; a 
single signum in bursa, developed as a small, cupped, 
granulate plate. 

A distinct genus, distinguished at once by its male 
antenna, venation, and transtilla. Contains one 
tropical American species. 


45. Mildrixia constitutionella Dyar 
Figures 169, 651 


Mildrizia constitutionella Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 405, 1914. 

Forewing grayish fuscous with some whitish dusting, 
especially in median area about the discal spots; ante- 
medial scale ridge blackish preceded by a narrow white 
line and followed by an indistinct dark shade; discal 
dots at end of cell distinct, black; just beyond the lower 
diseal dot, an outwardly angled mark from the upper 
edge of which a narrow dark shade extends to the 
inner costal edge of the subterminal line (distinct only 
on well-marked and unrubbed specimens) ; subterminal 
line narrow, denticulate, pale, bordered inwardly and 
outwardly by somewhat broader dark lines; terminal 
dots blackish, more or less confluent. Hind wing trans- 
lucent, opalescent white, the veins faintly darkened 
toward their outer extremities, especially on the females; 
a dark shade along costa and a narrow one along termen. 
Alar expanse, 19-22 mm. 

Genitalia as given for the genus. 

Tyrer Locanitry: Jalapa, México (type in USNM). 

Foop puant: Unknown. 


Distrizution: Méxtco: Jalapa. 
cin Santa Maria (June, July, Oct.). 


GUATEMALA: Vol- 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 27 


7. Genus Sematoneura Ragonot 


Sematoneura Ragonot, Nouv. Gen., p. 9, 1888; Monograph, pt. 
1, p. 186, 1893. (Type of genus: Sematoneura atrovenosella 
Ragonot.) 

Tongue well developed. Antenna of male shortly 
ciliate (cilia a trifle longer than width of shaft), of 
female weakly pubescent. Labial palpus upturned, 
reaching vertex, cylindrical, slender; third segment 
about two-thirds length of second, acuminate. Max- 
illary palpus filiform. Forewing smooth; 11 veins; 
vein 2 from well before lower outer angle of cell; 
3 from the angle, much nearer to 4 than to 2; 4 and 5 
closely approximate for a short distance from cell; 
6 from well below upper angle of cell, straight; 8 and 9 
long stalked; 10 shortly stalked with 8-9; male without 
costal fold. Hind wing with vein 2 from well before 
outer angle of cell; 3 from before, but near the angle; 
4 and 5 closely approximate for a short distance from 
the angle; 7 and 8 closely approximate or contiguous 
beyond cell; cell about half the length of wing; dis- 
cocellular vein curved. Eighth abdominal segment of 
male with a single, broad ventral hair tuft. 

Male genitalia with apical process of gnathos a 
simple, elongate hook, slightly notched at apex. Uncus 
broadly triangulate. Transtilla complete, stout, arched, 
its central area developed as a flat, broad lobe with 
slightly concave terminal margin. Harpe with costa 
sclerotized and produced at apex into a short project- 
ing digitus. Anellus a slightly curved plate with 
moderately long lateral arms. Aedeagus simple; penis 
armed with a single elongate, moderately stout cornutus, 
about one-third as long as aedeagus. Vinculum stout, 
as broad as or a trifle broader than long, tapering to 
broad, truncate terminal margin. 

Female genitalia with bursa and ductus bursae 
simple, without signum, smooth except for minute 
granulations in bursa; ductus bursae shorter than bursa; 
genital opening simple; ductus seminalis from bursa 
near its junction with ductus bursae. 

The foregoing description is drawn from the type 
species (atrovenosella). In male genitalia the new 
species (abitus), tentatively included in the genus, 
departs in some apparently essential details of structure, 
having a different type of transtilla and gnathos and 
lacking the apical projection from costa of barpe; but 
in all other structural characters it agrees with atroveno- 
sella. When its female is discovered a new generic 
placement may be necessary. 


46. Sematoneura atrovenosella Ragonot 
Fieures 4, 171, 653 
Sematoneura atrovenosella Ragonot, Nouv. Gen., p. 10, 1888; 
Monograph, pt. 1, p. 136, 1898. 

Forewing gray with more or less ochraceous dusting 
above inner margin and in outer area between the veins; 
the veins conspicuously outlined by blackish scaling; a 
similar narrow, dark line along the lower fold; these 
dark lines expanded and intensified at basal third 
indicating the remains of an antemedial band, and 
broken in outer area by a rather broad, faint, pale 


subterminal band; lower discal dot at end of cell 
faintly indicated; a line of blackish dots along termen 
between the vein ends. Hind wings dusky white, 
translucent; the veins darkened and a narrow dark 
line along termen. Alar expanse, 26-35 mm. 

Genitalia as given for the genus. 

Typn Locatity: Chanchamayo, Pert (type in Zool. 
Mus. Univ. Berlin). 

Foop prant: Unknown. 

DistrisuTion: México: Coatepec. Costa Rica: 
Huan Vifas (Jan., Feb., Nov.), Tuis (May). Conom- 
BIA: La Selva (San Juan Chaco, Sept.), Juntas (San 
Juan Chaco, Feb.). Ecuapor: Alpayacu (Rio Pas- 
taza), Quito. Prrt: Chanchamayo, Santo Domingo 
(Nov.). Argentina: Tucuman. 

One example before me (a female from Santo Domin- 
go, Peri, 6,000 ft.) differs in coloration from normal 
specimens in having a dark suffusion over the basal area 
to the antemedial line and a distinct antemedial pale 
line with continuous, black outer border. It also lacks 
any trace of ocherous dusting on the forewing. In 
genitalia and otherwise in color and maculation it is 
normal. I believe that it is only a color form. The 
specimen was from the unplaced material in the British 
Museum. 

47, Sematoneura abitus, new species 
Fiaurs 172 


Similar in color and markings to atrovenosella except a 
short strongly contrasted black streak just below costa 
at base and a broad black streak along median fold 
extending from base to end of cell. The ciliations of the 
male antenna are also a trifle shorter than those of 
atrovenosella. Alar expanse, 31 mm. 

The male genitalia differ markedly from those of 
atrovenosella in several details. Apical projection of 
gnathos is a rather short triangulate, pointed hook. 
The costa of harpe is broadly sclerotized but lacks the 
projecting digitus at apex. The gnathos is developed 
into a strongly sclerotized, hairpinlike, backwardly 
projecting loop with dense scobinations along its inner 
margin. 

Typs Locatity: Alpayacu, Rio Pastaza, East Ecua- 
dor (type in BM). 

Foop prant: Unknown. 

Described from unique male type collected by M. G. 
Palmer at 6,000 ft. The specimen is not in good con- 
dition but the essential features of the pattern are 
distinguishable and the male genitalia are so distinctive 
that description seems justified. In the absence of a 
female the generic placement cannot be made with 
absolute certainty. I expect, however, that the female 
genitalia will exhibit no radical difference from those of 
the type of the genus. 


8. Genus Hypsipyla Ragonot 


Aypsipyla Ragonot, Nouv. Gen., p. 10, 1888; Monograph, pt. 1, 
p. 187, 1898. (Type of genus: Hypsipyla pagodella Ragonot, 
synonym of Magiria robusta Moore; India; figs. 173, 656.) 


Tongue well developed. Antenna of male shortly 


28 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


ciliate (cilia but slightly longer than width of shaft, 
except on male of dorsimacula where they are about 
twice as long as width of shaft). Labial palpus of male 
upturned, reaching vertex, slender; third segment about 
half as long as second, acuminate; of female obliquely 
ascending. Maxillary palpus filiform (moderately large 
in pagodella and grandella, minute in other species). 
Forewing smooth; 11 veins; vein 2 from well before 
lower outer angle of cell; 3 from the angle, much closer 
to 4 than to 2; 4 and 5 approximate for a short distance 
from cell, occasionally connate; rarely short stalked; 6 
more or less bent towards base and more or less approxi- 
mate to upper angle of cell; 10 normally from the cell, 
rarely connate or shortly stalked with 8-9; male without 
costal fold. Hind wing with vein 2 from well before 
outer angle of cell; 3 from before, but near the angle; 4 
and 5 normally shortly stalked, occasionally connate, 
partially anastomosed or (in some large females) closely 
approximate for a short distance from cell; 7 and 8 
closely approximate beyond cell; cell half or (males of 
grandella) somewhat less than half the length of wing; 
discocellular vein curved. Highth abdominal segment 
of male simple or with two or three pairs of ventral hair 
tufts. 


Male genitalia with apical process of gnathos a hook, 
forked at apex. Uncus more or less triangulate; apex 
rounded. ‘Transtilla complete, stout, arched, its cen- 
tral area produced into two widely spaced horns; the 
latter stout in all species except pagodella. Harpe with 
costa, strongly sclerotized but not produced at apex. 
Anellus V- or U-shaped with long lateral arms. Aedea- 
gus simple; penis armed with a single, more or less 
twisted, fattened bladelike cornutus (except dorsima- 
cula). Vinculum stout, short or but slightly longer 
than broad, with truncate, broad terminal margin. 


Female genitalia with bursa and ductus bursae 
simple, unsclerotized except for a narrow band along 
ventral margin of genital opening; with or without 
signum; when present, the latter developed as a small, 
scobinate, cup-shaped plate; ductus bursae shorter 
than bursa; ductus seminalis from bursa at its junction 
with ductus bursae. 


The.genus is very close to and difficult to distinguish 
from Sematoneura. It is characterized chiefly by the 
bent condition of vein 6 of forewing. In Sematoneura 
this vein is always perfectly straight and remote from 
8-9 at base. In Hypsipyla 10 is also normally from the 
cell and there is frequent stalking of 4 and 5 of hind 
wing, neither of these conditions occurs in Sematoneura; 
but the venation is so individually variable in Hypsipyla 
that it cannot be trusted. 


48, Hypsipyla grandella (Zeller) 
Fieures 5, 174, 655 


Nephoteryx grandella Zeller, Isis von Oken, 1848, p. 881. 

Hypsipyla grandella (Zeller) Ragonot, Monograph, pt. 1, p. 189, 
1893.—Dyar, Ins. Inse. Menstr., vol. 7, p. 41, 1919.—Monte, 
Rev. de Ent., Brazil, vol. 3, p. 281, 1933. 

Hypsipyla cnabella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 405, 
1914. 


Forewing grayish fuscous shaded (especiaily on lower 
half of wing) with dull rust-red; veins outlined in black; 
antemedial pale line narrow, incomplete, rounded out- 
ward at middle and indented at vein 1b, bordered out- 
wardly by a narrow, discontinuous, black line; beyond 
this in median area between top of cell and vein 1b, an 
expanded faint whitish patch; some faint whitish dust- 
ing also in the middle-outer area between the veins; 
subterminal line faint, indicated chiefly by the intensi- 
fied black streaks bordering it inwardly on the veins, 
sinuate, deeply notched at lower fold; discal spots 
obsolete; terminal black dots between the vein ends 
distinct. Hind wing hyaline white with a fuscous 
shade along costa, some fuscous shading on the vein 
ends and a narrow fuscous line along termen. Alar 
expanse, 23-45 mm. 

Male genitalia with uncus rather abruptly narrowed 
beyond its broad base, the apex narrowly rounded; 
apical hook of gnathos narrow, short; paired horns of 
transtilla curving outward (away from each other); 
vinculum distinctly broader than long, its terminal 
margin very broad and but slightly convex, nearly 
straight. Highth abdominal segment of male simple. 
Female genitalia with signum. 

TYPE LocaLitins: Brazil (grandella, location of type 
unknown to me) ; Cérdoba, México (cnabella, in USNM). 

Foop piants: Cedrela and Swietenia (larva bores it 
fruits and branches). 

Distripution: Unirep States: Florida, Miami 
(Nov.). Mexico: Cérdoba (Feb., Sept.), Jalapa. 
GuaTreMaLa: Cayuga (Apr., May, Oct.), Chejel (June), 
Quirigué (June). Honpuras: La Cambra (Feb.). 
Costa Rica: Avangarez (July), Juan Vifias (Jan., 
Feb., Nov.), San José (Jan.), San de Montes de Oca 
(Oct.), Tuis (May, June). Panam: Almirante (Aug.), 
Summit (C. Z., Mar.). Purrtro Rico: Cayey (May). 
Cusa: Santiago de las Vegas (Apr.). Hartt: Pétion- 
ville (June, Dec.). Jamaica. TRINIDAD (Dec.). Con- 
ompia: ‘Above Rio Negro.” VEnEzuELA: El Valle 
(July), Maracay, Trompillo (July). British GUIANA: 
Georgetown (July). Brazit: Aragatuba (Sao Paulo, 
Apr.), Baia (Feb.), Campo Bello, Castro (Paranda), 
Espirito Santo, Nova Teutonia (May), Santa Catarina 
(Aug., Sept.). Paraguay: Sapucay (Oct.), Villarrica 
(Sept., Oct.). HEcuapor: Loja. ArGcentina: Tucuman. 
Pert: Lima. 

Generally distributed throughout tropical America 
wherever its food plants occur. 

A native American species close to and superficially 
similar to the Indina H. robusta; but with different male 
and female genitalia. It is apparently of some eco- 
nomic importance in the West Indies and South America 
as a pest of mahogany and the Cedrela species. Like 
many borers it varies greatly in size, and the venation 
is more than ordinarily unstable even for a phycitid. 
Vein 10 of forewing may be from the cell, separated 
from, closely approximate or connate with 8-9 or some- 
times shortly stalked with them, Veins 4 and 5 may 
be anything from approximate towards base to shortly 
stalked. Vein 6 is always slightly bent towards base 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 29 


but less so and more remote from the base of 8-9 on 
large females than on the smaller females and average- 
size males. On the hind wing 4 and 5 are usually 
shortly stalked or connate but on some large specimens 
are closely approximate for nearly half their lengths 
beyond the lower angle of the cell. 


49. Hypsipyla ferrealis (Hampson), new combination 
Fiaures 176, 657 
Crocidomera ferrealis Hampson, Ann. Mag. Nat. Hist., ser. 10, 
vol. 4, p. 352, 1929. 

Maculation of forewing similar to that of grandella 
except: Brownish fuscous, the general color decidedly 
more brown than gray, the dull reddish dusting giving 
the wing a somewhat rosy brown tint; dark lining on the 
veins less conspicuous and more discontinuous; ante- 
medial pale line very faint, obsolete on many specimens, 
indicated chiefly by its broken black outer margin; the 
white spot beyond this black margin (conspicuous in 
grandella) absent or indicated only by a faint ocherous 
white shade; subterminal line very faint, indicated by a 
black shading on the veins along its inner margin, sinu- 
ate, rather deeply indentate at vein 6 and lower fold. 
Hind wing smoky fuscous with a faint brownish or 
ocherous tint, more or less smoky white towards base; 
veins darkly outlined. Eighth abdominal segment of 
male with three pairs of ventral hair tufts. Alar 
expanse, 20-43 mm. 

Male genitalia with uncus of the same shape, but 
wider than that of grandella; vinculum narrower, 
longer than broad; transtilla with horns of central area 
divergent, forming a round-bottomed \V, the central 
connecting part of the transtilla slender. Female geni- 
talia without signum. 

TypE Locatity: Sixaola River, Costa Rica (type in 
BM). 

Foop PLANT: 
the seeds). 

Distrisution: Costa Rica: Cachi, Cain, Juan 
Vifias (May), Puerto Limén, Sixaola River (May, 
Sept.), Tuis (May). Conompra: San Antonio (Dec.). 
VENEZUELA: Maturaca (Sept.). FRencH GUIANA: 
Cayenne, St. Jean Maroni, St. Laurent Maroni. 
Trinipap: Caparo. Braziu: Pard. 

A distinct species easily distinguished from grandella 
by its dark hind wings. The foregoing food plant and 
Venezuelan records are from a series of small reared 
specimens (20-24 mm.) submitted by Dr. Ballou in 
1942. These are not only considerably smaller than 
average from the other localities listed; but are some- 
what grayer in color. The genitalia, however, are 
like those of typical Costa Rican examples. In 
venation ferrealis appears somewhat less variable than 
grandella. Vein 10 of forewing is always from the 
cell and 4 and 5 usually shortly stalked, rarely con- 
tiguous for a short distance beyond the cell. 


Carapa guianensis (larvae feeding on 


50. Hypsipyla dorsimacula (Schaus), new combination 
Figures 175, 654 


Myelois dorsimacula Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p. 245, 1913. 


The female type is badly rubbed and the markings 
consequently obscured; ground color bronzy brown; 
faint indications of a pale antemedial line rather far 
out on wing; fainter indications of a subterminal line; 
at end of cell a dark brown spot on discocellular vein 
and shortly separated from it a similar spot in cell, 
between them a pale spot (this marking at end of cell 
seems the characteristic pattern character of the 
species) ; below discocellular vein, on lower fold, a short 
blackish streak. Hind wing semihyaline, lilacine, 
darker towards apex; veins not appreciably darker. 
Alar expanse, 40 mm. 

Female genitalia without signum. 

TypEe Locauity: Sixaola River, Costa Rica (Sept.) 
(type in USNM). 

Foop prant: Unknown. 

In addition to the female type I have before me 
what I believe to be a male of the same species from 
the Janse Collection, taken at La Selva, San Juan, on 
the Chaco slopes of Colombia (4,600 ft., Sept.). It is 
as badly rubbed as the type but shows the same charac- 
teristic markings at end of cell and on the fold beneath; 
ground color of forewing rust-red; hind wing hyaline 
white with a very faint ocherous tint; antennal ciliations 
longer than on other species of the genus; at least 
twice as long as width of shaft. Male genitalia with 
stouter gnathos and quite differently shaped transtilla 
from previous species, the prongs of transtilla rather 
narrowly separated; penis without cornutus. The 
venation is alike on both specimens except for vein 
10 of forewing, which is from the cell on the female 
and short-stalked with 8-9 on the male; 4 and 5 of 
fore and hind wings are short-stalked; 6 of forewing 
is sharply bent towards base and connate with 8-9. 
Eighth abdominal segment of male simple. 


51. Hypsipyla fluviatella Schaus 
Fiaure 177 


Hypsipyla fluviatella Schaus, Ann. Mag. Nat. Hist., ser. 8, 
vol. 11, p. 246, 1913. 

Forewing long, and narrower in proportion than other 
species of the genus; reddish brown, darker brown in 
basal area; antemedial line well out towards middle 
of wing, indicated by its brown outer border, the 
latter out-bent from costa, thence nearly vertical to 
inner margin, forming three lunules, defined by narrow, 
pale buff inner shadings (the remains of the pale 
antemedial line), also preceded on inner margin by some 
silvery gray dusting; a narrow, elongate, pale buff 
patch on inner margin at tornus; a similar, wider, pale 
streak from cell to outer margin, occupying the space 
between veins 5 and 8 and bisected longitudinally by 
a narrow red-brown streak along vein 6. Hind wing 
hyaline white, inner margin rather broadly tinted with 
ocherous, and a narrow ocherous line along outer 
margin. Alar expanse, 45-46 mm. 

Male genitalia with uncus broad throughout, its 
terminal margin broadly rounded. Transtilla com- 
plete but the central fusion weak; the horns widely 
spaced, forming a broad, shallow U. Apical process 


30 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


of gnathos rather broad, oval, flattened. Highth 
abdominal segment of male simple. Female unknown. 

TypE LocaLity: Sixaola River, Costa Rica (type in 
USNM). 

Foop prant: Unknown. 

A distinct species easily distinguished by its narrow, 
peculiarly marked, red-brown forewings and the oval, 
flattened shape of the apical process of gnathos. The 
male hind wings are distinctly triangulate, but this is 
probably only a sex character. 

The forewing venation is fairly stable; vein 10 from 
the cell; 6 bent towards base and narrowly separated 
from 8-9 at base; 4 and 5 closely approximate for a 
short distance from cell. Hind wing with veins 4 and 5 
shortly stalked or closely approximate for some distance 
from the cell. 

The species is represented only by the type series 
of four males from the type locality. 


9. Genus Hemiptilocera Ragonot 


Hemiptilocera Ragonot, Nouv. Gen., p. 9, 1888; Monograph, pt. 
1, p. 144, 1898. (Type of genus: Hemiptilocera chino- 
graphella Ragonot.) 


Tongue well developed. Antenna of male with basa! 
segment elongated, cylindrical; a tuft of scales on base 
of shaft (weak on chinographella); shaft unipectinate 
for two-thirds, crenulate and pubescent beyond. An- 
tenna of female like that of the male type (chino- 
graphella) except for lack of scale tuft on shaft and for 
shorter basal segment; on other species of the genus 
shaft simple and pubescent. Labial palpus ascending; 
reaching to or nearly to vertex (shorter on chinographella 
than on other species); slender. Maxillary palpus 
small, squamous. Forewing smooth; 11 veins; vein 
2 from well before lower outer angle of cell; 3 from the 
angle, much closer to 4 than to 2; 4 and 5 connate 
(chinographella) or closely approximate at base and 
for a very short distance from cell (other species of 
genus); 6 bent towards base, close to or connate with 
8 at base; 10 from the cell, closely approximate to the 
stalk of 8-9 for some distance from cell; male without 
costal fold. Hind wing with vein 2 from well before 
outer angle of cell; 3 from the angle, connate with or 
closely approximate to 4-5; 4 and 5 stalked for about 
half their lengths; 7 and 8 closely approximate beyond 
cell; cell one-half or slightly less than one-half the 
length of wing; discocellular vein curved. Eighth 
abdominal segment of male with one or more paired 
hair tufts and sternal plates modified, one strongly 
sclerotized element in the form of an open loop. 

Male genitalia with apical process of gnathos a hook 
with notched apex. Uncus triangulate, apex bluntly 
pointed. Transtilla complete, stout, arched, its central 
area produced into widely spaced horns. Harpe with 
costa strongly sclerotized (produced at apex on bigrana 
and plumigerella, not produced on chinographella). 
Anellus with short, stout, lateral arms. Penis armed 
with strongly sclerotized cornutus and numerous 


sclerotized wrinklings. Vinculum stout, longer than 
broad, terminal margin broad and more or less indented. 

Female genitalia with signum developed as a small, 
strongly scobinate pocket; bursa large; ductus bursae 
short, more or less sclerotized and strongly scobinate- 
granulate, the scobinations and granulations extending 
into bursa for a short distance from place of junction 
with ductus; genital opening with sclerotized and more 
or less wrinkled plate on lower margin (except on 
exoleta); ductus seminalis from bursa near junction of 
bursa and ductus bursae. 


This genus is distinguished by its pectinate male 
antenna, the strong, stalking of veins 4—5 of hind wing, 
the long vinculum with broad terminal margin, the 
short arms of anellus and its short, granulate-scobinate 
ductus bursae. Eventually it may have to be restricted 
to its type species (chinographella) and a new generic 
placement found for the other species now included. 
All of these have simple pubescent female antennae; 
while those of chinographella are pectinate in both sexes. 
There are also several differences between males of 
chinographella and those of plumigerella and bigrana 
(notably in the shape of the transtilla, the costal 
development of harpe, and the size of the antennal 
tuft); but unfortunately we do not know the males of 
three other species (letharda, jocarella, exoleta) and until 
they are known it seems the wiser course not to attempt 
further generic separation. All the species have similar 
habitus and wing maculation and (except for exoleta) 
female genitalia showing only specific differences. 


52. Hemiptilocera chinographella Ragonot 
Figures 178, 658 


Hemiptilocera chinographella Ragonot, Nouv. Gen., p. 9, 1888; 
Monograph, pt. 1, p. 144, 1893. 

Male antenna with scale tuft on base of shaft weak, 
clay colored. Antenna of female pectinate, the pecti- 
nations a trifle shorter than those of the male. Thorax 
and basal segment of antenna clay-yellow. Forewing 
clay yellow (‘‘olivaceous ocherous” according to 
Ragonot) dusted and shaded with dull reddish brown 
and dark grayish fuscous, the ground color predomi- 
nating in the basal area, along the costa and (more 
faintly) bordering the termen and as a narrow longi- 
tudinal streak between the transverse line and including 
at its middle the lower discal spot; antemedial line 
faint, indicated chiefly by the broken elements of its 
outer dark border (a short notched blackish streak 
slanting outwardly from costa, a blackish spot on top 
of cell, another on lower vein of cell and a third on lower 
fold, these three blackish spots in a vertical line out 
near middle of wing, on a few well-marked specimens 
connected by a very faint, twice-outcurved dark line); 
subterminal line somewhat stronger, inwardly margined 
by a black spot, outwardly margined by a duller dark 
shade, more or less accented at costa and on the veins; 
discal dots at end of cell separated, distinct, especially 
the lower one, black; along termen a row of distinct, 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 31 


well contrasted, black dots (these a rather characteristic 
feature of most of the species of the genus); on the 
female a conspicuous whitish patch on inner margin at 
inner edge of the subterminal line (this whitish patch 
not present on the males before me). Hind wing 
semihyaline, shaded with smoky fuscous towards apex, 
on the veins and narrowly along termen. Eighth 
abdominal segment of male with a single moderately 
long pair of ventrolateral hair tufts. Alar expanse, 
22-26 mm. 

Male genitalia with central part of transtilla quadri- 
form (the horns flattened and with flattened lobes from 
their bases, the space between the horns even through- 
out); harpe with costa not produced at apex; apex of 
cornutus enlarged, sharply bent and bearing a row of 
thornlike spines. Female genitalia with ventral plate 
at genital opening smooth or nearly so; granulations of 
ductus bursae dense and forming a continuous sclero- 
tized mass. 

Typn Locatity: Chanchamayo, Peri (type in Zool. 
Mus. Univ. Berlin). 


Foop puant: Unknown. 


Distripution: Frencu Guiana: Cayenne. Braziu: 
Paré. Prrt: Chanchamayo. 


53. Hemiptilocera bigrana (Zeller) 
Fieures 180, 660 


Myelois bigrana Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 200, 


Hemiptilocera bigrana (Zeller) Ragonot, Monograph, pt. 1, p. 145, 
1893. 


Male antenna with hair tuft strong, black. Antenna 
of female pubescent. Thorax and base of antenna dull 
whitish dusted with fuscous. Forewing pale gray dusted 
with dull rosy and dark grayish fuscous; the dark dust- 
ing more evenly distributed than in chinographella; 
lower discal spot at end of cell large, strongly contrasted; 
upper spot weak or not distinguishable; antemedial line 
not defined; subterminal defined by its dark borders, 
which consist of blackish streaks on the veins. Hind 
wing hyaline white with a faint smoky fuscous shade at 
apex, on the outer half of the veins and narrowly along 
termen. Eighth abdominal segment of male with a long, 
strong pair of ventrolateral hair tufts and two other 
pairs of modified scale tufts. Alar expanse, 25-29 mm. 

Male genitalia with central part of gnathos a stout 
crescent-shaped projection with the horns widely 
spaced; harpe with apex of costa produced into a short 
spine; cornutus a spatulate ribbed blade. Female with 
ventral plate at genital opening deeply wrinkled; ductus 
bursae with a central, elongate, irregular patch of scob- 
inations, extending into adjacent area of bursa. 


Typs Locauity: Honda, Colombia (type in BM). 
Foop prant: Unknown. 
DistripuTion: México: Guerrero (Aug.), Iguala 


(Guerrero, June), Popocatépetl Parks (Distrito Federal, 
June). Conomera: Honda. 


The Mexican records are from specimens in the U. S. 
National Museum. 


54. Hemiptilocera plumigerella (Ragonot), new combination 
Ficure 179 


Nephopteryx plumigerella Ragonot, Nouy. Gen., p. 14, 1888; 
Monograph, pt. 1, p. 261, 1893. 

This species is known only from the male type. From 
Ragonot’s description the forewing color and markings 
must be similar to those of chinographella except much 
more heavily overshaded with vinous brown. The lower 
discal spot is conspicuous as in bigrana and the antenna] 
tuft is stout and ochraceous in color. Hind wing iri- 
descent, semitransparent, grayish brown. Eighth 
abdominal segment of male with a single pair of long, 
stout ‘ventrolateral hair tufts and another pair of 
much shorter, central tufts. Alar expanse, 21 mm. 

Male genitalia with transtilla as in bigrana, except 
that the crescent-shaped central projection is much more 
slender; cornutus a short bluntly pointed plate with a 
row of short, blunt spines near apex. 

Type tocatity: ‘America Meridionalis’” (type in 
Paris Mus.). 

Foop pirant: Unknown. 


55. Hemiptilocera letharda (Schaus), new combination 
Ficure 662 


Chloropaschia letharda Schaus, Proc. Ent. Soc. Washington, 
vol. 24, p. 237, 1922; Ann. Carnegie Mus., Pittsburgh, vol. 
16, p. 112, 1925. 

Forewing olive buff shaded with vinaceous fawn on 
subcoastal, median, and basal areas; the wing markings 
black, and some faint scatterings of black scales on fore- 
wing and thorax; antemedial line indicated only by 
broken fragments of its narrow outer border; discal 
black spots at end of cell both conspicuous, the lower 
one large; black dots on veins forming the inner margin 
of the pale subterminal line and black dots along termen 
also conspicuous. Hind wing semitransparent, smoky 
white, the veins slightly darkened; a dark shade towards 
apex and a narrow dark line along termen. Alar 
expanse, 29 mm. 

Female genitalia similar to those of bigrana but with 
a heavier concentration of scobinations in ductus bursae 
and a much stronger signum. 

Type Locauiry: Cabima, Panamé (May; type in 
USN). 

Foop piant: Unknown. 

In addition to the female type I have before me a 
British Museum female from Presidio, México, origi- 
nally identified as H. bigrana. It is paler than the 
Schaus type, but this apparently is due to its faded and 
slightly rubbed condition. Schaus recognized his 
original misplacement of the species in the Epipaschiidae 
and in his 1925 paper referred it to the Phycitinae, trans- 
ferring the specimen to Hemiptilocera in the National 
Collection; but I am unable to find any published 
reference of his or Dyar’s giving of the generic reference. 


32 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


56. Hemiptilocera jocarella (Schaus) 


Figure 659 
Acrobasis jocarella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p. 245, 1913. 
Hemiptilocera jocarella (Schaus) Dyar, Ins. Inse. Menstr, vol. 7, 
p. 42, 1919. 


Forewing reddish brown, the reddish (vinous) shade 
predominant in cell and, broadly, along lower fold; a 
dull olivaceous shade along costa and in terminal area; 
antemedial line obsolete, indicated only by fragments of 
its outer border (a black narrow streak from costa to 
cell, a black spot on lower margin of cell, and a thin, 
in-bent black streak from vein 1b to inner margin); 
subterminal line indicated by a pale black margined 
spot on costa and an outward series of short whitish 
streaks on veins 6 to 1b, these spots inwardly and out- 
wardly margined by black dots; a series of black dots 
along termen (less distinct than in the other species of 
the genus) ; the usual black discal spots, only the lower 
one pronounced, and it but slightly so. Hind wing 
glossy purplish or smoky brown; the veins but faintly 
darkened; a fine dark line along termen. Alar expanse, 
21-26 mm. 

Female genitalia exhibit (in the amount of scobina- 
tion of ductus bursae and the smaller size of signum) 
but trifling differences from those of letharda. The male 
of jocarella is unknown. 

Type Ltocatity: Avangarez, Costa Rica (type in 
USNM). 


Foop PLANT: Unknown. 


Distripution: Costa Rica: Avengarez (July). 
Panami: Porto Bello (Dec.). Brazm: Nova Teutonia 
(May). 


These records from four females in the U. S. National 
Museum. When males can be associated it is quite 
ikely that jocarella will prove to be only the female 
form of plumigerella. 


57. Hemiptilocera exoleta (Zeller) 
Fieure 661 


Myelois exoleta Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 201, 


Hemiptilocera exoleta (Zeller) Ragonot, Monograph, pt. 1, p. 146, 
1893. 


This species is represented only by the female type. 
T have seen nothing that matches Ragonot’s description 
or Zeller’s rather crude figure; but from both and from 
details of the female genitalia the reference is doubtful. 
Ragonot himself questions the correctness of his generic 
placement; but in the absence of a male no better 
placement could or can be made. 

The forewing shows the usual distinctively contrasted 
row of terminal dots and the maculation otherwise is 
that of a Hemiptilocera except that the usual discal dots 
are replaced by areddishlunule. According to Ragonot 
the cell of hind wing is also short for a Hemiptilocera. 
Alar expanse, 25 mm. 


The genitalia show a peculiar development of the 


eighth segment collar, a central-dorsal, invaginated, 
sclerotized pocket flanked by a pair of irregular, elon- 
gate, flattened lobes and on dorsum of ovipositor a pair 
of shallow sclerotized pockets (fig. 661a). The genital 
opening also is unsclerotized, lacking the usual ventral 
shield. 

Typr tocatity: Honda, Colombia (type in BM). 

Foop prant: Unknown. 

A male will be needed for certain generic placement. 


10. Genus Crocidomera Zeller 


Crocidomera Zeller, Isis von Oken, 1848, p. 865.—Ragonot, Mono- 
graph, pt. 1, p. 132, 1893. (Type of genus: Crocidomera 
turbidella Zeller.) 

Tongue well developed. Antenna pubescent; male 
with a short, blunt, spine from upper outer angle of 
basal segment of shaft; basal segment of male antenna 
swollen and broadly scaled. Labial palpus upcurved, 
reaching to vertex or slightly above it; third segment 
over half as long as second, acuminate. Maxillary 
palpus small, squamous. Forewing smooth; 11 veins; 
vein 2 from well before lower outer angle of cell; 3 from 
the angle; 4 and 5 approximate for a short distance from 
cell; 6 from below upper angle of cell, very slightly bent 
towards base; 8-9 stalked for about half their lengths; 
10 from the cell approximate to the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from well 
before lower outer angle of cell; 3 from the angle; 4—5 
contiguous for about one-third their lengths beyond cell; 
7 and 8 closely approximate beyond cell; cell slightly 
less than half the length of wing; discocellular vein 
curved. Eighth abdominal segment of male with com- 
pound ventral and ventrolateral tufts. 

Male genitalia with apical process of gnathos an 
elongate hook, blunt and rounded or slightly notched at 
apex. Uncus broad, hoodlike, apical margin broadly 
rounded or broad and truncate. 'Transtilla complete, 
stout, arched, with a strongly forked central projection. 
Harpe with costa rather broadly sclerotized, but not 
produced at apex, clasper more or less developed, simple, 
erect. Anellus with broad, dorsoventrally flattened 
lateral arms. Aedeagus with longitudinal rows of 
thornlike spines towards apex; penis with sclerotized 
wrinklings, but otherwise unarmed. Vinculum stout, 
slightly constricted from middle to moderately broad 
terminal margin; slightly longer than broad. 

Female genitalia with signum developed as a small 
scobinate cup-shaped patch; ductus bursae moderately 
long (shorter than bursa), expanding gradually to the 
wide genital opening and with some strong sclerotized 
wrinklings before genital opening; lower margin of 
genital opening sclerotized, wrinkled and more or less 
finely scobinate; dense, fine scobinations on the dorsal 
membrane behind genital opening; ductus seminalis 
from bursa near junction of bursa and ductus bursae. 

A tropical American genus ranging as far north as our 
Texas border and probably into southern Florida; easily 
distinguished by its genitalia and male antenna. 


e 
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 35 


58. Crocidomera turbidella Zeller 
Ficures 182, 664 


Crocidomera turbidella Zeller, Isis von Oken, 1848, p. 865.— 
Ragonot, Monograph, pt. 1, p. 132, 1893. 

Ground color of forewing variable, pale ocherous gray 
or white shaded with faintly reddish or reddish brown on 
outer area and in a narrowing shade towards base along 
inner margin; an indistinct blackish spot on costa 
beyond base; two other blackish dots marking the place 
of the obsolete antemedial line, one on costa, the other 
at top of cell; a dark patch on inner margin at what 
would be the inner margin of the antemedial line, on 
well marked specimens containing one or two minute 
black dots or dashes; subterminal pale line faint but 
distinguishable, indicated chiefly by an inner border of 
black spots on the veins and similar, fainter dark streaks 
(or a confluent dark shade) bordering it outwardly; a 
row of black dots along termen, quite marked on fresh 
specimens; discal dots at end of cell small, black, more 
or less confluent, when separated the lower not appreci- 
ably enlarged; below these, on the fold of some speci- 
mens, a larger spot of reddish or brown scales. Hind 
wing transparent, opalescent white with a faint fuscous 
shading at apex and at the vein ends and a fine dark 
line along termen. Alar expanse, 20-25 mm. 

Male genitalia with apical process of gnathos bluntly 
rounded at apex; uncus subtriangulate, its apical mar- 
gin evenly rounded; central projection of transtilla 
V-shaped, the prongs divergent and rather slender; 
harpe with outer margin of cucullus evenly rounded. 
Female genitalia with sclerotization along lower margin 
of genital opening narrow and but slightly wrinkled. 

Tyrer Locaity: ‘South America” (type in BM). 

Foop piant: Unknown. 

DistrRipution: Cusa: Baracoa, Santiago (June, Oct.). 
Jamaica. M*sxico: Jalapa. Unirep Statszs: Terzas, 
San Benito (May, Oct.). These localities from examples 
(in USNM) from which the foregoing description was 
drawn. 

I have omitted the Moschler reference cited by 
Ragonot for I suspect that the Puerto Rican specimens 
which he and Ragonot had and from which the Ragonot 
description was partly drawn are not turbidella but 
Jissuralis without the peculiar longitudinal black streak 
on forewing characteristic of the type of fissuralis. I 
have seen no specimens of Crocidomera from any South 
American locality except the Bolivian example men- 
tioned in the following discussion of fissuralis. I doubt 
that this could be Zeller’s species. 


59. Crocidomera fissuralis (Walker) 
Figures 183, 665 


Nephopteryx fissuralis Walker, List, vol. 27, p. 58, 1863. 

Myelois (?) adonea Felder and Rogenhofer, Reise... Novara..., 
Lepidoptera, pl. 137, fig. 8, 1874. 

Crocidomera fissuralis (Walker) Méschler, Die Lepidopteren- 
Fauna von Portorico, p. 327, 1890.—Ragonot, Monograph, 
pt. 1, p. 138, 1893. 


A photograph of the female type of fissuralis shows a 
specimen with a strongly contrasted, black longitudinal 


300329—56——4 


streak on forewing extending along lower median vein 
from base of wing to subterminal line and giving off two 
short forks, one along vein 2, the other along vein 3. 
The figure of adonea, presumably also a female, shows a 
similar marking. I have seen nothing to match this 
peculiar pattern except one female from the Janse col- 
lection from the Provincia del Sara (Department of 
Santa Cruz), Bolivia. The genitalia of this specimen 
match those of fissuralis fairly well except for the 
sclerotization of the ductus bursae, which is more like 
that of stenopteryx. The specimen cannot be placed with 
certainty until a male from the type locality is associated 
with it. I doubt very much that the peculiar longi- 
tudinal streaking represents anything more than an 
aberrational or varietal character; for I have before me 
a series of three males and three females from Puerto 
Rico which lack the longitudinal streak, but are obvi- 
ously distinct specifically from what I have recognized 
as turbidella. Their female genitalia are identical in all 
details with those of the type of fissuralis. On super- 
ficial characters they differ from turbedilla chiefly in 
having the lower discal spot at end of cell more pro- 
nounced and distinctly enlarged in comparison to the 
upper discal spot. Alar expanse, 25-30 mm. 

Male genitalia with terminal margin of uncus notched ; 
apical projection of gnathos slightly notched at apex; 
central projection of transtilla with its prongs converging 
towards their apices; harpe with cucullus triangulate, 
its apex narrowly rounded; aedeagus much stouter and 
its thornlike spines stronger and more numerous than 
those of turbidella or stenopteryx; terminal margin of 
vinculum nearly straight, terminal part of vinculum pro- 
portionally about twice as wide as that of either turbi- 
della or stenopteryx. Female genitalia with ventral 
plate at genital opening deeply wrinkled. 

TYPE LOCALITIES: Santo Domingo [Dominican Re- 
public] (jissuralis, adonea, both in BM). 

Foop prant: Unknown. 

DistrisuTion: Dominican REPUBLIC. PuERTO 
Rico: Aguirre Central (Apr.), Coamo Springs (Apr.), 
Culebra Isl. (Feb.). 


60. Crocidomera stenopteryx (Dyar), new combination 
Ficures 181, 663 


Dioryctria stenopteryx Dyar, Ins. Insc. Menstr., vol. 10, p. 16, 
Q 

Forewing ocherous gray with blackish markings; a 
black dot at base of cell; three black streaks beyond on 
lower median vein; a couple of black dots on costa near 
base; a black discal spot at lower, outer angle of clel; 
antemedial pale line indicated below median vein, pre- 
ceded by a dark patch on inner margin (somewhat 
tinted with reddish brown on female), margined out- 
wardly by an obscure, blackish line, beyond which 
on inner margin a somewhat diffused dark shade; vein 
1b more or less outlined in black scaling; subterminal 
line faint, bordered inwardly and outwardly by some 
blackish dots or streaklets on the veins; a fine black line 
along termen (formed by the confluent terminal dots). 
Hind wing transparent, hyaline white with a faint 


34 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


grayish shade along costa and a fine fuscous line along 
termen. Alar expanse, 25-27 mm. 


Male genitalia with terminal margin of uncus broad, 
very slightly convex; apical process of gnathos slightly 
notched at apex; prongs of central projection of trans- 
tilla slightly convergent toward their apices; harpe with 
terminal margin of cucullus oblique, apex bluntly 
pointed. Female genitalia with ductus bursae smoothly 
sclerotized between its sclerotized, wrinkled part and 
the schrotized and wrinkled margin of genital opening. 


TYPE LocALiITy: Tehuacan, Mexico (type in USNM). 


Foon piant: Unknown. 
Known only from the type locality. 


11. Cuniberta, new genus 


TypE or GENUS: Nephopteryx subtinctella Ragonot. 

Tongue well developed. Antenna of male with basal 
segment elongate, cylindrical; shaft weakly serrate and 
pubescent (the cilia about as long as width of segments), 
basal segments swollen and incurved, forming a sinus 
containing a, row of minute thornlike spines and overlaid 
with a spread of appressed scales; antenna of female 
simple and very weakly pubescent. Labial palpus up- 
turned, scarcely reaching vertex; third segment shorter 
than second, acuminate. Maxillary palpus small, squa- 
mous. Forewing smooth; 11 veins; vein 2 from before 
(but rather near) lower outer angle of cell; 3 from the 
angle, but little further from 2 at base than from 4; 4 
and 5 short stalked; 6 from below upper angle of cell, 
straight; 8 and 9 stalked for half their lengths; 10 from 
the cell, closely approximate to basal half of the stalk 
of 8-9; male without costal fold. Hind wing with vein 2 
from well before lower outer angle of cell; 3 from the 
angle, connate with 4-5; 4 and 5 stalked for approxi- 
mately half their lengths; 7 and 8 closely approximate 
beyond cell; cell slightly less than half the length of 
wing; discocellular vein curved. Eighth abdominal seg- 
ment with 2 pairs of ventrolateral hair tufts. 


Male genitalia with apical process of gnathos an 
elongate, stout, rather broad hook with blunt, notched 
apex. Uncus triangulate. Transtilla complete, stout, 
arched, produced at middle into a broad U-shaped pro- 
jection. Harpe simple. Anellus with rather broad, 
dorsoventrally flattened lateral arms. Aedeagus mod- 
erately slender with a single row of very minute serra- 
tions along one lateral margin towards apex, otherwise 
simple; penis with fine sclerotized wrinklings, otherwise 
unarmed. Vinculum stout, somewhat longer than 
broad. 


Female genitalia with signum developed as a small 
granulate cup-shaped patch; bursa small; ductus bursae 
considerably longer than bursa, unsclerotized except for 
a narrow sclerotization along lower margin of genital 
opening; ductus seminalis from ductus bursae near 
genital opening. 

The genus is close to both Hemiptilocera and Crocido- 
mera and shares some of the characters of each but is 
distinct from both in the definite stalking of veins 4-5 


of forewing and in the attachment of the ductus semi- 
nalis of the female genitalia. 
Contains one North American species. 


61. Cuniberta subtinctella (Ragonot), new combination 
FieurEes 170, 666 
Nephopteryx subtinctella Ragonot, N. Amer. Phycitidae, p. 7, 
1887; Monograph, pt. 1, p. 302, 1893.—Hulst, Phycitidae of 
N. Amer., p. 146, 1890.—McDunnough, Check list, No. 
6175, 1939. 

Forewing gray, more or less dusted with whitish on 
basal and median areas; antemedial line nearly vertical, 
out-angled at upper and lower margins of cell, bordered 
outwardly by a black line which is expanded and 
strongly accented on costa, bordered inwardly on lower 
margin by a reddish or reddish olivaceous patch; on 
some specimens a similar shade in fold beyond the ante- 
medial line; subterminal line sinuous, bordered in- 
wardly by a fine black line which, in most specimens, 
expands on costa into a conspicuous black spot or 
streak; on costa following the subterminal line a similar 
more or less expanded black spot; discal dots at end of 
cell usually confluent and forming a thin black lunule 
along the discocellular vein; a thin black streaklet on 
vein 2. Hind wing pale smoky fuscous; veins scarcely 
darker; a faintly darkened line along terminal margin. 
Alar expanse, 22-26 mm. 

Genitalia as given for the genus; male with apex of 
uncus narrowly rounded; vinculum evenly tapering to 
rather broad terminal margin. 


TypE LocaLity: California (type in Paris Mus.). 


Foop Prant: Unknown. 

Distripution: Unirep Srarus: Utah, Park City 
(June), Provo (uly, Aug.); California, Shasta Retreat 
(Siskiyou County, Aug.). Canapa: British Columiia, 
Kaslo (June). 


12. Heras, new genus 


Types or GENus: Heras disjyunctus, new species. 

Tongue well developed. Antenna of male with first 
segment rather long, cylindrical; shaft with a sinus and 
heavy scale tuft at base, otherwise weakly pubescent. 
Labial palpus upceurved, reaching above vertex; dorso- 
ventrally flattened; third segment somewhat shorter 
than second. Maxillary palpus squamous. Forewing 
smooth; 11 veins; vein 2 from well before lower outer 
angle of cell; 3 from the angle; 4 and 5 connate or very 
shortly stalked, shortly separated from 3 at base; 6 
from slightly below upper angle of cell, very slightly 
bent towards base; 10 from the cell, closely approximate 
to the stalk of 8-9; on male, a long narrow costal fold 
and, on upper surface of wing, a fovea (depressed 
pocket) in cell slightly beyond base. Hind wing with 
vein 2 from well before lower outer angle of cell; 3 from 
the angle, connate with 4-5; 4 and 5 stalked for half 
their lengths; 7 and 8 closely approximate beyond cell; 
cell about half the length of wing; discocellular vein 
curved. Eighth abdominal segment of male with ster- 
nal plate developed as a narrow sclerotized pocket at 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 35 


its center; in the intersegmental area two pairs of mem- 
branous eversable lobes (not haired), one long ventro- 
lateral pair and one shorter dorsolateral pair. 

Male genitalia with apical process of gnathos a nar- 
row, somewhat flattened hook with slightly forked apex. 
Uncus triangulate. Transtilla complete, stout; a rather 
short bridge with long widely spaced lateral arms pro- 
jecting backward; and a similarly spaced, shorter pair 
of arms projecting forward and articulating with the 
anellus. Harpe with a strong, long hair tuft from outer 
surface of base of sacculus, otherwise simple. Anellus 
a narrow plate with very long, slender, strongly sclero- 
tized, lateral arms. Aedeagus rather long and slender, 
sclerotized only on dorsal half; penis with some weak 
wrinklings and minute scobinations near apex, other- 
wise unarmed. Vinculum approximately triangulate, 
but slightly longer than its greatest width; its central, 
ventral area unsclerotized. 

I very much dislike to erect & new genus on a single 
male; but the genitalic and secondary male characters 
of this example are so striking and its distinctness from 
any known genus is so obvious it seems best to give it 
a name and separate designation. The genus is ap- 
parently closest to Hemiptilocera. 


62. Heras disjunctus, new species 
Figure 184 


Forewing rosy fuscous with costal area beyond ante- 
medial line broadly clay colored (pale ocherous); the 
rose shade predominant on upper part of wing, the fus- 
cous shade more accented in lower fold and along inner 
margin; antemedial line weak, indicated chiefly by a 
distinct but small whitish ocherous spot near inner mar- 
gin; subterminal more distinct, whitish ocherous, ter- 
minating at inner margin in another pale spot similar 
to the one on antemedial line, inner dark margin of 
subterminal line narrow and very faint; discal dots at 
end of cell confluent, blackish; terminal dots confluent, 
some faint blackish streaking on the veins before and 
beyond the subterminal line. Hind wing pale smoky 
fuscous; darker along the veins and towards outer mar- 
gin. Alar expanse, 22 mm. 

Male genitalia with the long posteriorily projecting 
arms of transtilla terminating in flattened lobes; apex 
of uncus narrowly rounded. Vinculum tapering to 
evenly rounded terminal margin. Female unknown. 

TyPE tocatity. Don Amo, Colombia (200 ft., July) 
(type in Janse Coll.). 

Foop prant. Unknown. 

Described from unique male type. Superficially (in 
maculation and color) it strongly resembles Hyalospila 
stictoneurella Ragonot. 


13. Adanarsa, new genus 


Typs or Genus: Rhodophaea intransitella Dyar. 

Tongue well developed. Antenna simple and pubes- 
cent in both sexes. Labial palpus upturned, reaching 
to vertex; slightly flattened laterally; third segment 
about half the length of second, blunted and slightly 


broadened (ventrally) by scales at apex. Maxillary 
palpus small, squamous. Forewing smooth; 11 veins; 
vein 2 from well before lower outer angle of cell; 3 from 
the angle; 4 and 5 shortly stalked, separated at base 
from 3; 6 from below upper angle of cell, straight; 8 and 
9 stalked for half their lengths; 10 from the cell, closely 
approximate to the stalk of 8-9 for a short distance be- 
yond cell; male without costal fold. Hind wing with 
vein 2 from well before lower outer angle of cell; 3 from 
the angle, approximate to 4-5 at base; 4 and 5 stalked 
for half or nearly half their lengths; 7 and 8 weakly 
anastomsed for a short distance beyond cell; cell half 
the length of wing; discocellular vein curved. Eighth 
abdominal segment of male with a small pair of ventro- 
lateral hair tufts. 


Male genitalia with apical process of gnathos a slen- 
der, rather long hook with slightly forked apex. Uncus 
semitriangulate. Transtilla complete, strongly sclero- 
tized and arched, supporting at its center a rather nar- 
row, smooth, curved crosspiece. Harpe with strongly 
sclerotized, erect clasper, otherwise simple. Anellus 
with short, broad, dorsoventrally flattened lateral arms. 
Aedeagus with a row of very fine serrations along one 
lateral edge towards apex; penis armed with a single, 
slender, sinuate, cornutus. Vinculum stout, about as 
long as greatest width; terminal margin broad. 

Female genitalia with cornutus developed as a single, 
short, stout, hooked thorn; ductus bursae much shorter 
than bursa, broad, flattened and with a broad transverse 
sclerotized band across it at junction of ductus and 
bursa; genital opening weakly and narrowly sclerotized 
along its lower margin and with a narrow, transverse 
sclerotized band in the membrane just behind the open- 
ing; ductus seminalis from bursa near its junction with 
ductus bursae. 

A distinct genus distinguished from related genera 
with complete transtilla by the strongly sclerotized 
clasper and the slight but definite anastomoning of 
veins 7-8 of hind wing. The amount of anastomosis 
varies in different specimens of the type species but is 
always present and always for somewhat less than half 
the length of the veins. 


63. Adanarsa intransitella (Dyar), new combination 
Ficures 185, 667 


Rhodophaea intransitella Dyar, Proc. Ent. Soc. Washington, vol. 
7, p. 33, 1905.—MeDunnough, Check list, No. 6075, 1939. 
Forewing pale ash gray with a fine scattered dusting 
of black scales and a very faint clouding of ocherous 
fuscous above inner margin between the transverse 
lines; antemedial line obsolete, indicated only by small 
black spot on costa and a larger black spot on inner 
margin at what would be the inner margin of the trans- 
verse line; outer line faint, indicated chiefly by border- 
ing black dashes on costa, a faint blackish line along 
its outer border and a few inwardly bordering black 
dots; lower discal spot black, followed outwardly by an 
obscure dark streak; a row of black dots along termen. 
Hind wing whitish, subpellucid; more or less shaded 


36 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


with fuscous towards apex and with a dark line along 
termen. Alar expanse, 16-19 mm. 

Male genitalia with apex of uncus evenly but rather 
narrowly rounded; clasper broadly flaring at apex; vin- 
culum no longer than broad; terminal margin abruptly 
truncate, nearly straight (very slightly concave at 
middle); aedeagus stout. Female genitalia as given for 
the genus. 

Type tocauity: Albuquerque, N. Mex. (type in 
USNM). 

Foop prant: Unknown. 

Distrisution: New Mexico, Albuquerque (July); 
Arizona, Christmas, Kingman (Oct.), Phoenix (Mar.). 


14. Birinus, new genus 


TYPE OF GENUS: Birinus russeolus, new species. 

Tongue well developed. Antenna simple and pubes- 
cent. Labial palpus upturned, slender, barely reaching 
to vertex; third segment shorter than second, acuminate. 
Maxillary palpus minute, filiform. Forewing smooth; 
11 veins; vein 2 from before (but moderately near) 
lower outer angle of cell; 3 from the angle, closer to 4-5 
than to 2; 4 and 5 very shortly stalked; 6 from very near 
to upper angle of cell, bent towards base, approximate 
at base to stalk of 8-9; 8 and 9 long stalked (for more 
than two-thirds of their lengths) ; 10 from the cell, closely 
approximate to stalk of 8-9 for a considerable distance 
beyond cell. Male without costal fold. Hind wing with 
vein 2 from well before lower outer angle of cell; 3 from 
the angle, connate with the stalk of 4—5; 4 and 5 stalked 
for a little over half their lengths; 7 and 8 closely approx- 
imate for a short distance beyond cell; cell about half 
the length of wing; discocellular vein curved. Highth 
abdominal segment of male simple. 

Male genitalia with apical projection of gnathos long, 
nearly straight, needlelike. Uncus spatulate, broadest 
at apical margin. Transtilla complete, a very slightly 
curved band (nearly straight on posterior margin), 
broad at its bases, narrow at middle. Harpe with sac- 
culus strongly sclerotized throughout its length and with 
apex produced as a short spur at lower outer angle of 
harpe; costa very short, sclerotized; bent upward at a 
sharp right angle a short distance from base, not pro- 
duced; cucullus forming more than half of the harpe 
area. Sclerotized part of anellus greatly reduced. Aede- 
agus long, moderately stout, smooth; penis armed with 
two narrow, blade like cornuti and numerous fine granu- 
lations. Vinculum triangulate, sclerotized only along 
margins. 

The foregoing description is incomplete, as the female 
is unknown; but the new genus seems to be justified by 
the male genitalia, which are unlike anything else in the 
American fauna. 


64. Birinus russeolus, new species 
Ficures 8, 186 


Forewing reddish brown, the rust-red shading a little 
more pronounced in outer costal and marginal areas and 
along lower vein of cell; a faint pale, clay-colored blotch 


in outer median area between vein 8 and the lower fold, 
enclosing in its center a small patch of blackish brown 
scales and at its inner margin bordered by a similar 
blackish smudge formed by the confluent discal spots; 
on the fold below and just before lower outer angle of 
cell a somewhat larger blackish brown patch preceded 
by a small clay-colored spot; a thin line of dark scales 
along the remainder of the fold to base of wing; the usual 
antemedial and subterminal lines obsolete; terminal dots 
very faint. Hind wing pale smoky fuscous, the veins 
brown and the cell filled with brown scaling. On the 
underside of fore and hind wings dark brown sex-scaling 
(a male character) covers the upper wing area (above 
lower margin of cell) from base to somewhat beyond the 
outer margin of the cell. Forefemora of male with a 
strong, clay-colored, hair tuft from upper basal angle 
(also a male character). Alar expanse, 22 mm. 

Male genitalia as given for the genus. Female un- 
known. 


TYPE LocaLiry: Tumatumari, Potaro River, British 
Guiana (type in Cornell). 

Foop prant: Unknown. 

Described from unique male type collected by W. T. 
M. Forbes, June 20, 1927 (Cornell lot 760 sub. 114). 


15. Genus Bertelia Barnes and McDunnough 


Bertelia Barnes and McDunnough, Contributions, vol. 2, p. 140, 
1913. (Type of genus: Bertelia grisella Barnes and McDun- 
nough.) 

Tongue well developed. Antenna of male with a 
strong posterior scale tuft on basal segment; shaft with 
a deep incurvation (sinus) at base, the sinus containing 
a few minute spinelike thorns but no scale tuft, shaft 
beyond sinus strongly unipectinate; antenna of female 
simple and pubescent. Labial palpus upturned on male, 
reaching a trifle higher than vertex; oblique on female; 
third segment on male slender, acuminate, about half 
the length of second, on female shorter and somewhat 
expanded with scales at apex. Maxillary palpus squa- 
mous. Forewing smooth; 11 veins; vein 2 from well 
before lower outer angle of cell, 3 from the angle; 4 and 
5 shortly stalked; 6 from below upper angle of cell, 
straight; 8 and 9 stalked for a trifle more than half their 
lengths; 10 from the cell, closely approximate to the 
stalk of 8-9; male without costal fold. Hind wing with 
vein 2 from well before lower outer angle of cell; 3 from 
the angle; 4 and 5 contiguous, shortly anastomosed or 
stalked, usually stalked for less than half their lengths; 
7 and 8 closely approximate for half their lengths beyond 
cell; cell about half the length of wing; discocellular vein 
curved. Highth abdominal segment of male with a pair 
of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos de- 
veloped as a stout hook, slightly notched at apex. 
Uncus triangulate. Transtilla incomplete, but with 
the elements long and stout, their apices broadly and 
irregularly developed and hooked. Harpe simple. 
Anellus U-shaped, its lateral arms dorsolaterally flat- 
tened. Aedeagus with a short row of minute scobina- 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 37 


tions along one lateral edge towards apex, moderately 
stout; penis with a few sclerotized wrinklings, otherwise 
unarmed. Vinculum stout, slightly longer than great- 
est width; terminal margin broad. 

Female genitalia with bursa large and elongate, 
signum present, developed as a small, scobinate, cup- 
shaped depression, ductus bursae very short; genital 
opening with a narrow, short, sclerotized plate on its 
lower margin and a large semicircular, sclerotized and 
scobinate dorsal plate in the membrane behind the 
opening; © pair of ventral scobinate plates in the inter- 
segmental area between eighth segment collar and 
Ovipositor; ductus seminalis from lobe of bursa near 
itsjunction with ductus bursae. 

A distinct genus containing one described North 
American species. The venation of the hind wing is 
individually variable in the amount of stalking or 
anastomosis of veins 4 and 5. 


65. Bertelia grisella Barnes and McDunnough 
Fieures 187, 669 


Bertelia grisella Barnes and McDunnough, Contributions, vol. 2, 
p. 140, 19183 —McDunnough, Check List, No. 6140, 1939. 

Forewing pale ashy gray dusted with fuscous; faint 
interrupted black streaking on upper and lower veins 
of cell, the lower fold and some of the veins beyond 
cell; antemedial line obscure, indicated chiefly by its 
narrow blackish outer border (out-angled from costa) 
and by a whitish incurved line between cell and inner 
margin, preceding which is an obscure dark shading; 
subterminal line nearly obsolete, followed on costa by 
a blackish shade; discal dots obsolete on many speci- 
mens, occasionally indicated by a small blackish dot 
at lower outer angle of cell. Hind wing semihyaline 
white with a faint ocherous tint; veins not appreciably 
darkened; a faint fuscous line along outer margin. Alar 
expanse, 24-30 mm. 

Male genitalia with apex of uncus bluntly and nar- 
rowly rounded; terminal margin of vinculum slightly 
angled; lateral arms of anellus moderately long and 
broad. Female genitalia as given for the genus. 

Type Locatity: Redington, Ariz. (type in USNM). 

Foop prant: Unknown. 

DistrisutTion: Arizona, Redington, Santa Catalina 
Mts. (Sept.). 


16. Genus Hypargyria Ragonot 


Hypargyria Ragonot, Nouv. Gen., p. 9, 1888, Monograph, pt. 1, 
p. 122, 1893.—Janse, Journ. Ent. Soc. South Africa, vol. 4, 
p. 149, 1941. (Type of genus: Hypargyria metalliferella 
Ragonot; India.) 

Tongue well developed. Antenna pubescent, basal 
segment on male elongate with a short spur of scales 
from its upper inner angle (giving the base of antenna 
much the same appearance as that of the undenuded 
two first segments of the male antenna of Mildrizia, 
fig. 169f); male shaft with a deep sinus towards base 
containing a longitudinal row of very minute teeth but 
no scale tuft; antenna of female simple. Labial palpus 


upturned, reaching a little above vertex; third segment 
nearly as long as second, acuminate. Maxillary palpus 
squamous. Forewing with transverse, antemedial ridge 
of raised scales; 11 veins; vein 2 from before but near 
lower outer angle of cell; 3 from the angle, at base about 
equidistant from 4-5; 4 and 5 connate or very shortly 
stalked; 6 from upper angle of cell, connate with stalk 
of 8-9, straight or but slightly bent towards base; 10 
from the cell, approximate to the stalk of 8-9; male 
with a notch in costa very close to base and on under- 
side at base of costa a small knot of modified scales and 
(projecting into the costal notch) a very short brush of 
stiff hairs. Hind wing with vein 2 from well before 
lower outer angle of cell; 3 from the angle, connate with 
stalk of 4-5; 4 and 5 stalked for slightly less than half 
their lengths; 7 and 8 closely approximate for less than 
half their lengths beyond cell; cell slightly more than 
one-third the length of wing; discocellular vein curved. 
Eighth abdominal segment of male with compound 
ventral scale tufts. 

Male genitalia with gnathos weak, lacking a central 
projection, the lateral arms meeting at the base of a 
rudimentary subanal plate. Uncustriangulate. Trans- 
tilla incomplete, but with the elements long and stout, 
their apices broadly developed. Harpe with costa 
broadly and very strongly sclerotized and stoutly pro- 
jecting at apex; a fine moderately long hair tuft from a 
sclerotized disk attached to base of sacculus. Anellus a 
broad, deep plate (bearing short, knoblike, stoutly 
spined, lateral projections on the American species). 
Aedeagus smooth; penis armed with two or more short, 
rather stout, straight spines, a deeply wrinkled, sclero- 
tized band, and a cluster of fine moderately long spines. 
Vinculum very stout, considerably larger than uncus 
and tegumen combined, longer than broad. 

Female genitalia with signum developed as a small, 
round, scobinate, cup-shaped plate; ductus bursae 
shorter than bursa, a broad, strongly sclerotized band 
at the junction of ductus and bursa and a narrower 
sclerotized band at genital opening; behind genital 
opening a conspicuous pair of strongly sclerotized, gran- 
ulate, pocket like lobes; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

Presumably an Old World genus of tropical and prob- 
ably African origin; possessing some structural char- 
acters of Acrobasis, Mildrixia, and Bertelia but amply 
distinct from any of them. It contains two American 
species. 


66. Hypargyria definitella (Zeller) 
Figures 188, 668 
Myelois definitella Zeller. 


p. 205, 1881. 
Hypargyria definitella (Zeller) Ragonot, Monograph, pt. 1, p. 124, 
1893. 


Horae Soc. Ent. Rossicae, vol. 16, 


Forewing purplish ocherous to purplish brown, most 
of basal area and costal half of median area white 
sparsely dusted with red scales, the red dusting most 
abundant along midcosta; a small ocherous patch on 
inner margin near base; antemedial line evenly curved, 


38 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


ocherous, bordered outwardly by a red or purplish line 
continued from a rather pronounced costal dash, and 
inwardly by the vertical scale ridge, the latter red, 
reddish ocherous or purple with some admixture of 
blackish scaling; subterminal line very faint with faint 
purplish borders; discal spots at end of cell separated, 
blackish; black terminal dots faint. Hind wing hyaline 
white with a faint smoky tint on some specimens; the 
veins darkened (brown) and a narrow brown line along 
termen. Undersides of male fore and hind wings in 
the area between vein 2 and costa and from near end of 
cell outward covered with shining silvery scales; also 
on forewing a short black median streak from base, 
more or less extended into cell along lower edge of upper 
vein of cell and on hind wing a similar black streak 
on upper vein of cell; these black sex-scalings not con- 
stant and altogether absent from occasional males. 
Alar expanse, 16-20 mm. 

Genitalia as given for the genus. 

TypE Locatity: Honda, Colombia (type in BM). 

Foop prant: Unknown. 

Distrisution: Purrto Rico: Puerto Real (Vieques 
Isl., Apr.), San Germdén. Virern Istanps: Kingshill 
(St. Croix, June, Oct.). CouomBra: Honda, Valparaiso. 
Braziu: Castro, Santa Catarina. 

The males of this species can be distinguished at once 
from any other American phycitid by the shining 
silvery scaling on the undersides of the wings; a char- 
acter, however, shared by the Old World type of the 
genus. The Old World metalliferliae exhibits 2 number 
of slight but consistent male genitalic differences: The 
heavier and more abundant spining on the penis, a 
different shape to the apical projection of costa of harpe, 
a different shape to the apices of the elements of trans- 
tilla (not developed into paired hooks as in the Ameri- 
can species), and an anellus without spined lateral pro- 
jections. Such differences are certainly specific but no 
more. Through the courtesy of the British Museum I 
have been able to examine males of metalliferella from 
Pusa in India and Nyasaland in Africa. There were 
no differences of any kind between them. 


67. Hypargyria slossonella (Hulst), new combination 


Salabria slossonella Hulst, Canadian Ent., vol. 32, p. 170, 1900. 

Acrobasis tenuella Barnes and McDunnough, Contributions, vol. 
2, p. 181, 1913. 

Acrobasis slossonella (Hulst) Barnes and McDunnough, Contri- 
butions, vol. 3, p. 195, 1916—McDunnough, Check list, 
No. 6108, 1939. 

Not distinguishable from definitella except that the 
males lack entirely the silvery scaling on the under- 
sides of the fore and hind wings. 

I suspect that it is only a variety or race of definitella; 
but until more material is available and something is 
known of their life histories the two forms will have to 
be kept as separate species. The genitalia of slossonella 
exhibit no differences of any specific significance from 
those of definitella. 

TypsE LocaLities: Miami, Fla. (slossonella, in AMNH, 
ex Rutgers); Everglades, Fla. (tenuella, in USNM). 


Foop pLrant: Unknown. 

DistrisutTion: Unrrep States: Florida, Everglades 
(Apr.), Fort Myers (Apr.), Miami (‘‘February-March’’). 
México: Oaxaca. 


17. Chararica, new genus 


Typ or Genus: Myelois annuliferella Dyar. 

Tongue well developed. Antenna simple and pubes- 
cent on both sexes. Labial palpus upturned, reaching 
to vertex, slender; third segment about as long as sec- 
ond, acuminate. Maxillary palpus squamous. Fore- 
wing smooth; 11 veins; vein 2 from before, but near, 
lower outer angle of cell; 3 from the angle; 4 and 5 
separated at base, distance separating them slightly 
less than that separating 3 and 4; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for slightly less 
than half their lengths; 10 from the cell, approximate 
to the stalk of 8-9 for a short distance; male without 
costal fold. Hind wing with vein 2 from well before 
lower outer angle of cell; 3 from the angle, connate 
with or very closely approximate to 4 at base; 4 and 5 
shortly stalked; 7 and 8 approximate for a short dis- 
tance beyond cell; cell about half the length of wing; 
discocellular vein curved. Eighth abdominal segment 
of male with a pair of ventrolateral hair tufts. 

Male with gnathos weak, lacking a central projec- 
tion, the lateral arms articulating with a thinly sclero- 
tized subanal plate. Uncus triangulate, sharply taper- 
ing to a blunt point. Transtilla complete, with a cen- 
tral sclerotized apron connecting transtilla and uncus, 
and with a pair of widely spaced lateral arms each 
bearing at its apex a clutter of slender spinelike setae. 
Harpe with costa slightly concaved, strongly sclero- 
tized, not produced at apex; cucullus large, forming 
about two-thirds of the harpe, outer margin broadly 
rounded. Anellus with stubby, stout, convergently 
directed, lateral projections. Aedeagus with lateral 
margins serrated toward apex; penis with a few weakly 
sclerotized wrinklings or granulations, otherwise un- 
armed. Vinculum stout, nearly square in outline. 

Female genitalia with signum developed as a small, 
finely granulate-scobinate, cup-shaped disk; bursa large; 
ductus bursae, broad, short, less than half as long 
as bursa, weakly sclerotized and transversely wrinkled 
towards genital opening; genital opening broad, with 
strongly sclerotized transverse plate along lower margin 
and some weak scobinations on the membrane above 
and behind the opening; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

This genus is erected for three North American spe- 
cies now listed under Rhodophaea, but differing from 
that genus in both venational and genitalic characters. 
It is easily recognized by its male genitalia. The spe- 
cies also have a pattern character which aids in iden- 
tification: the usual discal dots on forewing at end of 
cell are replaced by a small obicular marking. This is 
weak on bicolorella but present and distinguishable on 
most specimens. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 39 


68. Chararica annuliferella (Dyar), new combination 
Ficures 189, 670 


Myelois annuliferella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 33, 1905. 

Rhodophaea annuliferella (Dyar) Barnes and McDunnough, 
Check list of the Lepidoptera of Boreal America, No. 5516, 
1917.—McDunnough, Check list, No. 6074, 1939. 

Forewing dark gray with a faint, pale ocherous shade 
along inner margin; antemedial line well out on wing, 
outwardly arched in cell to slightly beyond middle of 
wing, inbent from lower fold to inner margin, white, 
bordered outwardly by a narrow black line; basal area 
with veins black and faint intervenous whitish dusting; 
subterming line rather near terminal margin, slightly 
sinuous, fine, white with a narrow, inner, black border; 
some faint white dusting in median area, especially 
along costa; usual discal dots at end of cell replaced 
by small black obicular mark with a whitish center. 

Hind wing hyaline white; veins not appreciably dark- 

ened; a faint, small fuscous shade at apex and a very 

faint, dark line along termen. Alar expanse, 19-22 mm. 

Genitalia of male with lateral arms of transtilla much 
reduced, widely spaced, their spinelike hair tufts long. 

Vinculum with terminal margin decidedly incurved at 

middle. Female genitalia with transverse sclerotized 

band on lower margin of genital opening narrow. 


Typxz Locatity: Gallinas Canyon, N. Mex. (type in 
USNM). 

Foop prant: Unknown. 

Distrisution: New Merico, Gallinas Canyon (July) ; 
Arizona, Colorado Desert (Yuma County), ‘‘So. Ariz.,”’ 
Kingman (Oct.), Yavapai County. 


69. Chararica hystriculella (Hulst), new combination 
Figures 190, 671 


Acrobasis hystriculella Hulst, Ent. Amer., vol. 3, p. 135, 1887. 

Rhodophaea hystriculella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 
114, 1889; Monograph, pt. 1, p. 78, 1893.—McDunnough, 
Check list, No. 6073, 1939. 

Myelois hystriculella (Hulst) Hulst, Phycitidae of N. Amer., p. 
119, 1890. 

Forewing whitish gray with extreme base dark fuscous 
and a similar fuscous shade over the outer area from 
subterminal line on costa obliquely to inner margin near 
antemedial line, and outward to termen; some black 
streaking on upper and lower veins of cell and vein 1b 
before the antemedial line; antemedial line well out 
towards middle of wing, twice angled outwardly, indi- 
cated chiefly by a fine black outwardly bordering line; 
subterminal line faint, with narrow dark inner and outer 
borders, beginning as blackish dashes on costa; obicular 
spot at end of cell conspicuous, black with a narrow 
whitish center; terminal dots black, more or less con- 
fluent. Hind wing hyaline white with a faint fuscous 
shade at apex and a narrow dark line along termen, 
these dark shadings very slight on the males, more ex- 


tended and stronger on some females; veins occasionally 
darkened on females, not darkened on males. Alar 
expanse, 17-23 mm. 

Male genitalia with uncus narrowly triangulate; lat- 
eral arms of transtilla rather long and their terminal 
hair tufts correspondingly shortened, not so widely 
spaced as those of annuliferella; terminal margin of 
vinculum very slightly concave, nearly straight. Fe- 
male genitalia with transverse sclerotized band on lower 
margin of genital opening broad (at least twice as wide 
as that of annuliferella). 

Typ Locauity: Texas (type, 2, in AMNH, ex Rut- 
gers). 

Foop prant: Unknown. 

Distrisution: Texas, Blanco County (Sept.), Browns- 
ville (May, June), Chisos Mts. (June), Cotula (Mar., 
Apr., May), Devils River (May), Kenedy (May), Kerr- 
ville (Aug.), Nueces River (Zavalla County, Apr.), 
Sabinal (Mar.), San Antonio (June, July), San Benito 
(Mar.), San Diego (Apr., May, June); Florida, Coconut 
Grove, Miami. 


70. Chararica bicolorella (Barnes and McDunnough), new 
combination 


Rhodophaea bicolorella Barnes and McDunnough, Canadian Ent., 
vol. 49, p. 404, 1917.—MceDunnough, Check list, No. 6077, 
1939. 

Forewing with costal half of basal area black, streaked 
and peppered with white, giving this area of the wing 
a slate-colored appearance to the naked eye; outer area 
of wing from subterminal line to outer margin and costa 
to lower fold a similarly dark shade; remainder of wing 
ocherous, shading outwardly to tawney or ruddy ocher- 
ous; antemedial line obsolete except along outer margin 
of blackish basal patch; subterminal line faint, narrowly 
and weakly bordered inwardly and outwardly by black- 
ish lines; obicular mark at end of cell very faint but 
distinguishable on most specimens; terminal dots con- 
fluent.. Hind wing hyaline white with a very faint 
ocherous line on outer margin for a short distance from 
apex. Alar expanse, 20-24 mm. 

Genitalia essentially like those of hystriculella. 

Typr Locauity: Christmas, Gila County, Ariz. (type 
in USNM). 

Foop prant: Unknown. 

Distrisution: Arizona, Christmas, Mohave County 
(May, June, July, Aug., Sept.), Redington; Nevada, 
Clark County (Apr.), ‘“‘So. Nevada” (July); California, 
San Bernadino County (Apr.). 

A striking species easily distinguished by its color pat- 
tern, but not structurally different from hystriculella. 
The original type series consists of two males and two 
females, not four males as stated by the authors. 

Both bicolorella and hystriculella have a strong hair 
tuft on the metathorax of the male adjacent to the base 
of the leg. This character is lacking in annuliferella. 


40 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Genera 18-21: Myelopsis to Ectomyelois 


[Venational division C. Forewing with 11 veins; 4and 5 stalked 
for half their lengths or less. Hind wing with veins 7 and 8 anas- 
tomosed for over half their lengths beyond cell (the free element 
of 8 shorter than the anastomosed stalk of 7-8). Transtilla of 
male genitalia complete.] 


18. Myelopsis, new genus 


Typr or GENus: Myelois coniella Ragonot. 

Tongue well developed. Antenna simple, pubescent. 
Labial palpus upturned, reaching slightly above ver- 
tex; second segment somewhat roughly scaled in front; 
third segment slightly shorter than second, acuminate. 
Maxillary palpus filiform. Forewing smooth; 11 veins, 
vein 2 from before but near lower outer angle of cell; 
3 from the angle; 4 and 5 shortly stalked, the stalk at 
base separated from 3 for a distance but slightly less 
than that between 3 and 2; 6 from below upper angle 
of cell, straight; 10 from the cell separated from 8-9 
at base, and more or less divergent beyond; male with 
out costal fold. Hind wing with vein 2 from well be- 
fore outer angle of cell; 3 from the angle, connate with 
the stalk of 4-5; 4 and 5 stalked for slightly less than 
half their lengths; 7 and 8 strongly anatomosed beyond 
cell, the free element of 8 short; cell half the length of 
wing; discocellular vein curved. Eighth abdominal 
segment of male simple. 

Male genitalia with apical process of gnathos U- 
shaped (consisting of a pair of widely spaced, short 
arms). Uncus stout, more or less triangulate, apex 
rather narrowly rounded. Transtilla complete, but 
weakly sclerotized. Harpe simple; costa strongly scler- 
otized and projecting at apex (except in subtetricella). 
Anellus U-shaped, narrowly sclerotized throughout. 
Aedeagus smooth; penis with sclerotized wrinklings. 
Vinculum triangulate, tapering, longer than greatest 
width. 

Female genitalia with or without signum, latter when 
present weak. Ductus bursae membranous. Genital 
opening simple. Ductus seminalis from middle or 
towards terminal end of bursa. 

This genus comprises several North American species 
that have been referred to Myelois Hibner. ‘The latter 
a, heterogenous assemblage of species, very few of which 
are actually congeneric with the type of genus (medu- 
lallis Hiibner synonym of cribrella Hiibner). The lat- 
ter has veins 7 and 8 of hind wing very shortly and 
weakly anastomosed beyond the cell, the free element 
of vein 8 correspondingly long and the cell itself over 
half the length of the wing. It belongs properly in 
our venational division B. None of the American and 
very few of the Old World species that have been as- 
signed to it are properly referrable to Myelois. The 
European éetricella Schiffermueller belongs in Myelopsis. 


71. Myelopsis coniella (Ragonot), new combination 
Fieures 191, 673 


Myelois coniella Ragonot, N. Amer. Phycitidae, p. 3, 1887; 
Monograph, p. 1, p. 53, 1893.—Hulst, Phycitidae of N. 


Amer., p. 118, 1890.—McDunnough, Check list, No. 6071, 
1939. 


Rampylla nefas Dyar, Ins. Insc. Menstr., col. 10, p. 172, 1922 
(new synonymy). 

Forewing pale ash gray (on Utah and Nevada spec- 
imens) to blackish gray; on darker specimens the basal 
and terminal areas are contrastingly paler than the 
area between the transverse lines; at extreme base on 
inner margin an obscure ocherous patch (not distin- 
guishable on worn or faded specimens) ; antemedial line 
oblique, white with a broad, black outer border; sub- 
terminal line sinuate, more or less contrastingly pale 
and inwardly bordered by a blackish line or varying 
intensity; discal dots at end of cell usually distinct, 
separated, black; terminal dots obscure, when distin- 
guishable, more or less confluent. Hind wing hyaline 
white to pale smoky fuscous with a fine dark line along 
termen. Alar expanse, 16-22 mm. 

Male genitalia with transtilla slender, arched and 
very weakly sclerotized at the central attachment of 
its elements. Harpe with costa narrowly sclerotized 
and projecting a trifle beyond the apex of the cucullus. 
Female genitalia without signum; bursa membranous; 
ductus bursae with some minute scobinations near its 
junction with bursa, otherwise smooth. 

TYPE Locatities: Nevada (coniella, in Paris Mus.); 
Mexico City, Mexico (nefas, in USNM). 

Foop puant: Unknown. 

DistriputTion: Unitep states: Nevada, Montgom- 
ery Pass (Mineral County, Sept.), Utah, Dividend 
(Aug.), Eureka (June, Aug.), Provo (July, Aug., 
Sept.), Stockton (Sept.), Trout Creek (Ibapah Mts., 
Sept.); Colorado, Glenwood Springs (Aug.); Arizona, 
Pinal Mts. (July), no definite locality (Aug., Sept.); 
New Mexico, Gallinas Canyon; Texas, Burnet County, 
(Mar.), Kerrville (Mar.); Nebraska, Sioux County, 
(July); Michigan, Dickinson County; Maine, Bar Har- 
bor (Aug.), Mount Desert (Aug.). Cawnapa: British 
Columbia, Kaslo (July, Aug.) ; Manitoba, Aweme (Aug.); 
Ontario, Ottawa (July, Aug.). Mexico: Mexico City 
(Sept.), Tehuacdn (Sept.). 

A variable species in color but with remarkably con- 
stant genitalia. Dyar’s nefas has much darker fore- 
wings than specimens from Utah or Nevada but no 
darker than some specimens from Arizona and New 
Mexico. 


72. Myelopsis immundella (Hulst), new combination 


Myelois immundella Hulst, Phycitidae of N. Amer., p. 117, 
1890.—Ragonot, Monograph, pt. 1, p. 49, 1893.—MceDun- 
nough, Check list, No. 6068, 1939. 

The type is without abdomen. In size, wing shape, 
pattern, and general coloration it is like the following 
species (subtetricella) except that the antemedial and 
subterminal lines of forewing are more whitish and dis- 
tinct and their dark borders (especially the outer border 
of the antemedial line) blackish and more strongly con- 
trasted against the dark ground color of the wing. The 
discal dots are also more strongly contrasted. 

The name may represent only a color form of subtetri- 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 41 


cella, but until other specimens matching the type are 
found and their genitalia studied this cannot be deter- 
mined one way or the other. 
TyprELocaLity: Texas (typein AMNH, ex Rutgers). 
Foop puant: Unknown. 


73. Myelopsis subtetricella (Ragonot), new combination 
Fiaures 192, 672 

Myelois subtetricella Ragonot, Ent. Amer., vol. 5, p. 113, 1889; 
Monograph, pt. 1, p. 47, 1893.—Hulst, Phycitidae of N. 
Amer., p. 117, 1890.—McDunnough, Check list, No. 6062, 
1939. 

Myelois zonulella Ragonot, Ent. Amer., vol. 5, p. 113, 1889; 
Monograph, pt. 1, p. 49, 1893.—Forbes, Cornell Mem. 68, p. 
613, 1923.—McDunnough, Check list, No. 6066, 1939. (New 
synonymy.) 

Myelois obnupsella Hulst, Phycitidae of N. Amer., p. 118, 1890.— 
Ragonot, Monograph, pt. 1, p. 48, 1893.—Barnes and 
McDunnough, Contributions, vol. 3, p. 193, 1916.—Forbes, 
Cornell Mem. 68, p. 613, 1923.—McDunnough, Check list, 
No. 6063, 1939. (New synonymy.) 

Forewing brownish gray with some faint whitish dust- 
ing on basal and median costal areas; antemedial line 
but slightly oblique, rather faint, dull whitish with a 
more or less obscured dark outer border; subterminal 
line obsolete or very faintly indicated; discal dark dots 
at end of cell separated, only the lower one distinct and 
always distinguishable. Hind wings dull smoky white 
to pale fuscous; veins darkened slightly in several speci- 
mens; a narrow dark line along termen. Alar expanse, 
20-24 mm. 


Male genitalia similar to those of coniella except scle- 
rotized costal margin of harpe abruptly terminated before 
apex of cucullus and not projecting as a free spur at apex. 
Female genitalia with a small weak signum; bursa 
weakly sclerotized, finely scobinate, and with a longi- 
tudinal sclex otized groove in area near ductus bursae; 
ductus bursae very short; ductus seminalis from bursa 
well towards its terminal end, 


Typs Locauities: ‘North America” (subtetricella, in 
Paris Mus.); north Illinois (zonulella, in BM); Canada 
(obnupsella, in AMNH, ex Rutgers). 


Foop piant: Unknown. 


DisrriputTion: Unrrep States: New Hampshire, 
Hampton; Massachusetts, Cohasset (July), Forest Hills 
(May, June), Framington (May), Winchendon (May); 
Pennsylvania, Beaver County (May), New Brighton 
(May, June), Pittsburgh (May); Jilmois, Arlington 
Heights (May), Chicago (May), Quincey (June); Ohio, 
Calla; Florida (no specific locality, Mar.). Canapa: 
Alberta, Bilby (June), Edmonton (May); Manitoba, 
Aweme (May, June). 

The species is quite distinct and easily identified by 
male and female genitalic characters. The Florida rec- 
ord cited above is from a spurious “type (male)’’ of 
Myelois immundella Hulst, originally in the Fernald Col- 
lection and now in the U. S. National Museum. It is 
not immundella, and in genitalia, color and markings 
agrees perfectly with other males of subtetricella. Rag- 
onot’s zonulella was described from four females in the 
British Museum labeled ‘‘N, Ill.” and bearing the num- 


ber “82-54.” I have examined the genitalia of two of 
these and they agree in all details with those of the type 
of subtetricella. As Ragonot designated no holotype I am 
selecting as lectotype one of the specimens J examined. 


74, Myelopsis minutularia (Hulst), new combination 
FiauRE 675 
Dioryctria minutularia Hulst, Ent. Amer., vol. 3, p. 135, 1887. 
Myelois minutulella Hulst, Phycitidae of N. Amer., p, 118, 1890.— 
Ragonot, Monograph, pt 1, p. 48, 1893. 
Myelois minutularia (Hulst) McDunnough, Check list, No. 6064, 
1939. 

The status of this species is in doubt. It is known 
only from females which look like small dark examples 
of coniella, of which it may be only a race or variety. 
However, the ductus bursae of minutularia is longer than 
that of typical coniella and the bursa shows considerably 


more scobination. Hind wing semihyaline, smoky 
white. Alar expanse, 11-13 mm. 
Type xnocauity: Blanco County, Tex. (type in 


AMNG, ex Rutgers). 
Foop pLant: Unknown. 


The only known distribution is Texas; examples be- 
fore me are from Blanco and Burnet Counties. The 
statement by Hulst in his original description that his 
types are males is an error. The male is unknown. 


75. Myelopsis alatella (Hulst), new combination 
Ficures 193, 194, 195, 674 


Acrobasis alatella Hulst, Ent. Amer., vol. 3, p. 135, 1887. 

Myelois rectistrigella Ragonot, N. Amer. Phycitidae, p. 3, 1887. 

Myelois alatella (Hulst) Hulst, Phycitidae of N. Amer., p. 118, 
1890.—Ragonot, Monograph, pt. 1, p. 52, 1893—McDun- 
nough, Check list, No. 6070, 1939. 

Myelois fragilella Dyar, Proc. Ent. Soc. Washington, vol. 6, p 
114, 1904.—McDunnough, Check list, No. 6060, 1939. 
(New synonymy.) 

Myelois piazzella Dyar, Ins. Inse. Menstr., vol. 13, p. 11, 1925.— 
McDunnough, Check list, No. 6061, 1939. (New 
synonymy.) 

Forewing ash gray more or less dusted with fuscous, 
general color varying from pale ash gray to grayish 
fuscous (but not so dark as some specimens of coniella) ; 
antemedial line oblique, indicated by its narrow, black 
outer border which is shortly and sharply out-angled 
at middle; subterminal line rather close and paralled 
to outer margin, sinuate, sharply indented between 
costa and vein 6, very slightly so at lower fold, often 
obscure, sometimes with a distinct inwardly bordering 
black line; discal dots separated, black, lower one (at 
least) always distinct. Hind wing semihyaline smoky 
white, somewhat darkened towards apex and with more 
or less darkening of the veins; a fine dark line along 
termen. Alar expanse, 20-26 mm. 

Male genitalia with transtilla a thin, weakly 
sclerotized subtriangulate plate. Harpe with costa 
broadly sclerotized, produced at apex, but not extending 
to apex of cucullus. Female genitalia with a small 
signum; a rather large round area of dorsal surface of 
bursa thinly sclerotized. 


42 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


TyprE Locauities: Napa, Calif. (alatella, in AMNH, 
ex Rutgers); California (rectistrigella, in Paris Mus.); 
Pecos, N. Mex. (fragilella, in USNM); San Diego, 
Calif. (piazzella, in USNM). 

Foop puant: Unknown. 

Distrisution: California, Clarkville (June), Monache 
Meadows (July), Napa, Placerville (May), San Diego 
(Mar., Aug.), San Francisco (Apr.); Utah, Bellevue 
(Apr.); Colorado, Gunnison County (near Almont, 
July); New Mexico, Fort Wingate (June, July), Jemez 
Springs (June), Pecos (June). 

An individually variable species in color and to some 
extent in male genitalia. The actual holotypes of 
alatella, fragilella, and piazzella seem different enough; 
but there are all intergrades among them in a series 
from any given locality. Indeed the two cotypes of 
alatella from Napa, Calif. (alike in color and markings) 
show considerable variation in details of male genitalia 
(width of the sclerotized costa of harpe, shape of 
transtilla, and spacing of the apical prongs of gnathos). 
The genitalia of the cotype from Napa (in USNM, fig. 
193) shows an extreme of variation. The other cotype 
(in AMNH, ex Rutgers, the actual holotype) has 
genitalia identical with those of piazzella shown in 
figure 195. At most, the Dyar names represent forms 
or varieties, but not species or local races. 


19. Genus Anypsipyla Dyar 


Anypsipyla Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 327, 1914. 
(Type of genus: Anypsipyla univitella Dyar.) 

Tongue well developed. Antenna of male shortly 
ciliate, the cilia no longer than width of shaft; of female 
pubescent. Labial palpus obliquely upturned, reach- 
ing slightly above vertex; third segment about as long 
as second, acuminate. Maxillary palpus very slightly 
dilated with scales at apex (subsquamous). Forewing 
smooth; 11 veins; vein 2 from before but close to lower 
outer angle of cell; 3 from the angle, approximate to 
2; 4 and 5 stalked for approximately half their lengths, 
approximate (rarely connate) to 3 at base; 6 from below 
upper angle of cell, straight; 8 and 9 stalked for about 
half their lengths; 10 from the cell, at base closely 
approximate to or connate with stalk of 8-9, thence 
divergent; male with short narrow costal fold. Hind 
wing with vein 2 from well before lower outer angle of 
cell; vein 3 from the stalk of 4-5; 4 and 5 long stalked; 
7 and 8 anastomosed beyond cell for appreciably more 
than half their lengths; cell about half the length of 
wing; discocellular vein curved. Hight abdominal 

OME of male with a single pair of ventrolateral hair 
tuits. 

Male genitalia with apical projection of gnathos an 
elongate hook with slightly notched apex. Uncus sub- 
triangulate (hoodlike). Transtilla complete, strongly 
arched. Harpe with costa strongly sclerotized through- 
out and projecting at apex beyond apex of cucullus; 
otherwise simple. Anellus a narrow band with slender 
lateral arms. Aedeagus simple; penis with some 
weakly sclerotized wrinklings, otherwise unarmed. 


Female genitalia with or without signa, if present, in 
the form of a row of very small, weak, thornlike spines; 
bursa very finely scobinate, ductus bursae considerably 
longer than bursa, simple; genital opening simple; 
ductus seminalis from anterior (terminal) end of bursa. 

A distinct genus with one tropical American species. 


76. Anypsipyla univitella Dyar 
Ficures 39, 196, 679 


Anypsipyla univitella Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 327, 
1914. 

Forewing fuscous gray with a broad white subcostal 
streak extending from near base to apex and touching 
costa near base and at apex; a black streak along mid- 
costal edge and on fresh specimens a fine black line along 
lower vein of cell and some faint black streaking on the 
outer veins; a fine powdering of reddish scales on the 
white subcostal stripe; discal dots faint or absent; trans- 
verse lines obsolete. Hind wing hyaline white with a 
smoky tint along costa and at apex and a fine dark line 
along termen. ‘Alar expanse, 20-32 mm. 

Male genitalia with terminal margin of uncus rather 
broadly round; apical projection of costa of harpe blunt; 
transtilla truncately arched. Female genitalia as given 
for the genus. The signa are usually absent and when 
present consist of from 2 to 10 very weak spines. 

TypE Locatity: Corozal, Canal Zone, Panama (type 
in USNM). 

Foop piants: Cassia brasiliensis, Samanea samdn 
(larva feeding in pods), Pacae (larva in fruit). 

Distrisution: Cusa: Victoria de las Tunas, San 
Blas (Trinidad Mts., May). Mexico: Colima (May, 
Nov.). Guatmmata: Cayuga (Apr.), Quirigué (Mar.). 
PanamA: Corozal (Apr., Noy.), Las Sabanas (Apr.), 
Porto Bello (May). VeEnezurza: El Valle (Apr.). 
Braziu: “S. E. Brazil,” Tapera (Pernambuco). PERU: 
Lima (Feb.). Ecuapor. Jamaica: Kingston (Dec.). 

Probably generally distributed in tropical America, 
where its host plants occur. 


20. Apomyelois, new genus 


Types oF Genus: Dioryctria bistriatella Hulst. 

Tongue well developed. Antenna simple and pubes- 
cent on both sexes. Labial palpus upturned, slender, 
reaching to slightly above vertex; third segment slightly 
shorter than second, acuminate. Maxillary palpus fili- 
form. Forewing smooth; 11 veins; vein 2 from well 
before lower outer angle of cell; 3 from the angle; 4 and 
5 stalked for slightly less than half their lengths, the 
stalk separated from 3 at base; 6 from below upper 
angle of cell, straight; 8 and 9 long stalked (for over 
two-thirds their lengths); 10 from the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from well 
before lower outer angle of cell; vein 3 from the angle; 
4 and 5 stalked for two-thirds of their length, the stalk 
connate with 3; 7 and 8 anastomosed beyond cell for 
about half their lengths (the anastomoses slightly longer 
than the free part of vein 8); cell a trifle more than half 
the length of the wing; discocellular vein curved. Kighth 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 43 


abdominal segment with a pair of weak, short, ventro- 
lateral hair tufts. 

Male genitalia with apical projection of gnathos an 
elongate, slender, simple hook. Uncus subtriangulate, 
apex rounded. Transtilla complete, well sclerotized 
and strongly arched; produced at middle into a flat, 
broadly and bluntly forked projection. Harpe with 
costa sclerotized throughout and projection slightly at 
apex; otherwise simple. Anellus U-shaped. Aedeagus 
smooth, slightly flaring at apex; penis with a single, 
straight, short, weakly sclerotized, spikelike cornutus 
and a few minute and weak scobinations; otherwise 
unarmed. 

Female genitalia with signa present as an oval cluster 
of thornlike scobinations; bursa otherwise smooth, large; 
ductus bursae considerably shorter than bursa, simple; 
genital opening simple; ductus seminalis from anterior 
(terminal) end of bursa. 

This genus is another subtraction from the composite 
genus Myelois of Authors. Of all the American species 
that have been referred to that genus it is the nearest 
to the type of Myelois (medullalis Hiibner, a synonym 
of cribreila Hiibner) of any American species, agreeing 
with cribrelia in forewing venation, except that the stalk- 
ing of vein 10 with 8-9 is less consistent in cribrella than 
in bistriatella. In eribrella 10 is often short stalked (as 
in fig. 38); but it is as often from the cell, connate with 
or approximate to or distinctly separated from the stalk 
of 8-9. In bistriatella it is from the stalk of 8-9 on all 
specimens that I have seen. However, this difference 
has no more than specific significance and would not of 
itself justify any separation of bistriatella from Myelois. 
There are some other differences that, in my judgment, 
are of generic character and justify such separation. 
The hind wing venation and length of cell are similar in 
cribrella and bistriatella except for the anastomosis of 
veins 7 and 8; in cribrella this anastomosis is very weak 
and shorter than it is in bistriatella and would place 
typical Myelois in our venational division B, while 
Apomyelois would go definitely into division C. In male 
genitalia cribrella (fig. 203) differs in having apical pro- 
jection of gnathos developed as a deeply, strongly, and 
narrowly forked process and the transtilla developed as 
a simple, strongly arched, narrow band. The female 
genitalia of cribrella (fig. 684) differs in more striking 
fashion: the ductus bursae being much longer than 
bursa and strongly granulate and partially sclerotized 
throughout most of its length; and ductus seminalis is 
from the bursa between the signum and the junction of 
bursa and ductus bursae. 

The new genus contains one North American species. 


77. Apomyelois bistriatella (Hulst), new combination 
Ficures 40, 197, 676 


Dioryctria bistriatella Hulst, Ent. Amer., vol. 3, p. 136, 1887. 

Myelois bilineatella Ragonot, N. Amer. Phycitidae, p. 3, 1887; 
Monograph, pt. 1, p. 48, 1893.—Hulst, Phycitidae of 
N. Amer., p. 117, 1890; U. S. Nat. Mus. Bull. 62, p. 418, 
1902. 

Myelois bistriatella (Hulst) Hulst, Phycitidae of N. Amer., p. 117, 
1890.—Ragonot, Monograph, pt. 1, p. 51, 1893.—Barnes 


and McDunnough, Contributions, vol. 3, p. 194, 1916.— 
Forbes, Cornell Mem. 68, p. 613, 1923.—McDunnough, 
Check list, No. 6067, 1939. 

Forewing gray-brown faintly dusted with white on 
costal half of basal area and in central area from mid- 
costa to lower margin of cell; transverse lines white, 
rather sharply contrasted, especially towards inner 
margin and without appreciably contrasted blackish 
borders; antemedial line transverse, from costa dis- 
tinctly before middle, straight, except for an occasional 
slight notch in cell; subterminal line somewhat narrower 
and less distinct, sinuate; dark discal dots at end of 
cell often fused into a single spot or line along discocel- 
lular vein, usually set off by the surrounding white 
dusting of the central area; terminal dots very faint, 
more or less confluent. Hind wing dull smoky white, 
the veins slightly darkened and a narrow dark line 
along termen. Alar expanse, 19-22 mm. 

Genitalia as given for the genus. 


Type Locauities: Washington, D. C. (bistriatella, 
type lost?); “America septentrionale” (bilineatella, in 
Paris Mus.). 

Foop prant: Unknown. 

Distrisution: Unirep States: Florida (Mar.); Dis- 
trict of Columbia, Washington; New York; Massachusetts, 
Framingham (May); Jilinois, Edgebrook (May); Wis- 
consin; Iowa, Ames (May). Canapa: Ontario, Ottawa 
(June, July), Trenton (May, June). 

Probably much more widely distributed throughout 
eastern and central United States and Canada, nowhere 
apparently a very abundant species. 

The supposed type of bistriatella is labeled “Iowa, 
H. S. Saunders, June 6, 1886.” It is definitely that 
species but, unless it is mislabeled or the type locality 
given by Hulst in his original description (“‘Washington, 
D. C.”’) is wrong, it could not be the actual holotype. 
I have seen no specimens anywhere labeled “‘Washing- 
ton, D. C.”’ There is a female in the National Museum 
from the Fernald Collection, bearing a Hulst type label 
but no locality. This might be the true type. It is a 
perfect match for the Iowa specimen in the Rutgers 
Collection. Since there can be no reasonable doubt as 
to what the name stands for we may as well consider 
the holotype lost and forget it. 


21. Ectomyelois, new genus 


Tyrer or genus: Myelois decolor Zeller. 

Tongue well developed. Antenna of male shortly 
ciliate (cilia about the length of width of shaft or 
slightly less), otherwise simple; of female simple and 
pubescent. Labial palpus upturned, reaching to or 
nearly to apex (not above it); second segment some- 
what broadened with scales; third segment short, 
distinctly shorter than second, acuminate. Maxillary 
palpus filiform. Forewing smooth; 11 veins; vein 2 
from well before angle; 3 from the angle, shortly 
separated from the stalk of 4-5 at base; 4-5 shortly 
stalked (very shortly stalked in most specimens and 
never for more than half the length of the veins); 


44 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


6 from below upper angle, straight; 8 and 9 long stalked, 
for over two-thirds of their lengths; 10 from the cell, 
closely approximate to the stalk of 8-9 for some distance 
from cell; male without costal fold. Hind wing with 
vein 2 from well before lower outer angle of cell; 3 from 
the angle; 4-5 stalked for not over half their lengths 
(usually for less), the stalk connate with or very closely 
approximate to 3 at base; 7 and 8 strongly anastomosed 
for most of their lengths beyond cell (free element of 8 
very short); cell half the length of wing; discocellular 
vein curved. Highth abdominal segment of male 
simple or with a weak, short pair of ventrolateral hair 
tufts. 

Male genitalia similar to those of the foregoing 
genus (Apomyelois) except: Apical process of gnathos 
slightly notched at apex; costa of harpe not produced 
at apex (except slightly in muriscis and zeteki); penis 
without cornutus; vinculum more truncate and less 
tapering. 

Female genitalia with signum sometimes absent, 
when present consisting of an elongate patch of scobina- 
tions; ductus bursae normally longer than bursa, in- 
dividually variable, simple, except for a weak scleroti- 
zation at genital opening; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

In male genitalia there is little or nothing of a generic 
character to separate Hctomyelois from Apomyelois. 
The two genera are distinguished by the shorter cell 
and much more extended anastomosis of veins 7 and 8 
of hind wing and the different place of departure of 
ductus seminalis from the bursa of the female. 

The genus is erected for another group of species 
(American and European) removed from Myelois of 
Authors. These species all appear to be of tropical or 
semitropical origin. They are distinguished from 
typical Myelois by the much stronger anastomosis of 
veins 7 and 8 of hind wing and the consequent reduction 
of the free element of vein 8. 


78. Ectomyelois decolor (Zeller), new combination 
Ficures 198, 677 


Myelois decolor Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 222, 
1881.—Ragonot, Monograph, pt. 1, p. 58, 1893.—Dyar, 
Proc. U. 8. Nat. Mus., vol. 47, p. 326, 1914.—Wolcoitt, 
Journ. Agr. Univ. Puerto Rico, vol. 20, No. 1, p. 476, 1936. 

Nephopieryx ephestiella Hampson, Ann. Mag. Nat. Hist., ser. 7, 
vol. 7, p. 257, 1901 (new synonymy). 


Forewing dark grayish fuscous with some white 
powdering in basal area and considerably more in the 
median area from slightly above inner margin and in 
outer area between subterminal line and termen; trans- 
verse lines white, well contrasted, especially the ante- 
medial line which is rather wide, sharply oblique, 
slightly indented at lower fold and (in some specimens) 
in the cell, outwardly bordered by a more or less ob- 
scure dark shade; subterminal line fainter, narrow, 
sinuate, obscurely and narrowly dark margined; some 
faint blackish streaking on the veins; discal dots at end 
of cell distinct, separated; terminal dots normally well 
contrasted and separate, blackish. Hind wings smoky 


white to pale smoky fuscous; the veins darkened and a 
narrow dark line along termen. Eighth abdominal 
segment of male with hair tufts. Alar expanse, 19-30 
mm, 

Male genitalia with outer margin of uncus rather 
evenly rounded; central area of transtilla produced into 
a moderately broad plate with notched terminal margin; 
anellus a broad plate with wide, flattened, incurved, 
stubby arms; vinculum nearly square in outline, its 
terminal margin very slightly concave. 

Female genitalia exhibiting considerable individual 
variation in the size of bursa and corresponding length 
of ductus bursae which is usually considerably longer 
than bursa; signum patch of variable shape but usually 
elongate. The female genitalia exhibit no distinctively 
specific characters. 

Type LocaLities: Honda, Colombia (decolor, in 
BM); Nassau, Bahamas (ephestiella, in BM). 

Foop PLANTs: Annona squamosa, Ceratonia siliqua, 
Hymenaea courbil; these records from reared specimens 
in the U.S. National Museum. Presumably the species 
has much the same hosts and habits as the closely re- 
lated Ectomyelois ceratoniae. The larvae feed in the 
fruits and are very difficult to separate from those of 
ceratoniae. 

Distripution: Cusa: Baracoa (Aug., Oct., Nov.), 
Havana, Santiago de las Vegas (Mar.), “Santiago Prov- 
ince” (Sept., Oct., Dec.). Purrro Rico: Arecibo, 
San Germdn (Apr.). Jamaica. Banamas: Nassau. 
GuaTEMALA: Cayuga (Mar., June, Aug.). PanamA: 
Porto Bello (Mar., Dec.). Conomspra: Honda, ‘West 
Slopes” (4,400 ft., Feb.). Venrzurxa: Aroa. BritisH 
Gu1ana: Tumatumari (June). FREncH Guiana: Cay- 
enne, St. Jean Maroni. Surtnam: Surinam River. 
Braziu: Paré (June), Ponte Nova (Rio Xingu, Amazon- 
as), Santa Catarina (July). 

Ragonot considered decolor as a probable variety of 
ceratoniae; but there is a consistent difference in the 
shape of the transtilla between the two which indicates 
more than varietal or racial difference; and in unrubbed 
and unfaded specimens the color difference is obvious 
and consistent. E. decolor seems to be confined to the 
New World while ceratoniae occurs in both the New and 
Old World. Hampson’s ephestiella is nothing more than 
a rather large example of decolor. Like other species in 
this genus, decolor varies greatly in size. 


79. Ectomyelois ceratoniae (Zeller), new combination 


Figures 199, 678 


Myelois ceratoniae Zeller, Isis von Oken, 1839, p. 176; 1848, p. 
675.—Ragonot, Monograph, pt. 1, p. 57, 1893.—Staudinger 
and Rebel, Catalog der Lepidopteran des palaearctischen 
Faunengebietes, vol. 2, No. 787, 1901.—Spuler, Die Schmet- 
terlinge Europas, vol. 2, p. 216, 1910.—Forbes, Cornell 
Mem. 68, p. 614, 1923.—Meyrick, Revised handbook of 
British Lepidoptera, p. 395, 1928.—Wolcott, Journ. Agr. 
Univ. Puerto Rico, vol. 20, No. 1, p. 475, 1936.—Corbet 
and Tams, Proc. Zool. Soc. London, ser. B, vol. 113, p. 68, 
1943. 

Myelois oporedestella Dyar, Proc. Ent. Soc. Wash., vol. 18, p. 30, 
1911.—McDunnough, Check list, No. 6065, 1939. (New 
synonymy.) 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 45 


Color and markings similar to decolor except: Fore- 
wing more uniformly gray, with less of the white dust- 
ing, especially on median area; antemedial line narrower 
and usually more distinctly notched. The chief char- 
acter, however, is in the transtilla of the male genitalia. 
The central projection of this organ is more constricted 
and decidedly narrower on ceratoniae than on decolor, 
and this difference seems to be consistent. In several 
preparations of each species from different rearings and 
localities I have found no intergrading examples. The 
female genitalia offer no satisfactory distinguishing 
characters. As in decolor, individual differences (even 
in the proportional length of the ductus bursae) are 
greater than any difference between the two species. 
Alar expanse, 16-24 mm. 


Type Locaities: Laibach, Austria (ceratoniae, in 
BM); Miami, Fla. (oporedestella, in USNM). 


Foop piants: Carissa grandiflora, Cassia bicapsularis, 
Ceratonia siliqua, Erisbotyra japonica (chiefly in mum- 
mied fruits), Livistona chinensis, Robinia, Tamarindus 
indica, Vachellia insularis. Also on dried figs, dates, 
raisins, and nuts in storage. Primarily a leguminous 
feeder. The favored host seems to be the pods and 
seeds of the corob (Ceratonia siliqua). 


Distrisution: Unitep Srares: Florida, Homestead 
(May), Key West (Apr.), Miami (May, July, Aug., 
Noyv.). Purrto Rico: Arecibo (May), Trujillo Alto 
(Mar., July). Jamaica (July). Ancentina: Buenos 
Aires (Feb.), Catamarca (May, June). Also in the Old 
World in the Mediterranean areas of Europe, Africa 
and Asia and (by introduction in dried fruits) extending 
into Central Europe and England. 


Apparently of Mediterranean origin, introduced by 
commerce and established in some tropical and semi- 
tropical areas of the New World. Probably much more 
widely distributed than indicated by the above records 
from specimens before me. The species is of minor 
importance as a feeder on the seeds of the corob. It 
has been found rather frequently at our port quarantine 
stations in shipments of English walnuts from Italy. 

I have omitted all European synonymy as I have not 
been able to verify its correctness. This, with further 
references to European literature will be found in 
Ragonot’s monograph and the Staudinger and Rebel 
catalog. Myelois phoenicis Durrant may be only a 
color variety or race of ceratoniae; a small series before 
me reared from dates from Algeria has the ground color 
of forewing white, but the male genitalic characters 
of ceratoniae. Corbet and Tams list phoenicis as a 
synonym. 

80. Ectomyelois muriscis (Dyar), new combination 
Figures 200, 680 


Myelois transitella Dyar (not Walker), Proc. U. S. Nat. Mus., 
vol. 47, p. 326, 1914. 

Hypsipyla muriscis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330, 
1914, 


Myelois palpalis Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 1919 
(new synonymy). 


Forewing dull rusty brownish ocherous to reddish 
brown; costal third to half of wing strongly dusted with 
white, the white area rather well contrasted against 
dark ground color; antemedial line angulate, obscure, 
indicated chiefly by a brown or blackish spot on its 
outer margin at or just below costa; subterminal line 
better defined, sinuate, margined inwardly and out- 
wardly by narrow dark lines, the latter especially 
emphasized at costa; discal spots usually distinct and 
separated, sometimes one or the other obscured by the 
white dusting or by an extension of the ground color, 
rarely fused into a line along discocellular vein, blackish 
brown; terminal dots more or less distinct. Hind wing 
dull, translucent white to smoky fuscous (as a rule 
darker on female than on male); a dark shade toward 
apex, some dark shading on the veins and a fine dark 
line along termen. Eighth abdominal segment of male 
simple. Alar expanse, 16-26 mm. 

Male genitalia with apical projection of gnathos slen- 
der, very long, extending at least as far backward as 
apex of uncus (when genitalia are in natural position it 
extends well beyond the uncus); transtilla a rather nar- 
row, sclerotized band, looped backward in a rounded 
arch; sclerotized costa of harpe very slightly and 
bluntly produced at apex. 

Female genitalia with or without signa, when present 
a patch of coarse scobinations, the patch varying in size 
in different specimens; ductus bursae simple or very 
weakly sclerotized on ventral surface at genital opening. 

Type Ltocauities: Cabima, Panamé (muriscis, in 
USNM); Cayuga, Guatemala (palpalis, in USNM). 

Foop piants: Mammea americana (larvae feeding in 
the fruit), Theobroma cacao (larvae in the pods). 

Distrisution: Hartt. Puerto Rico, Mayagiiez 
(July). Britis West Invres: Trinidad, several exam- 
ples with no more specific locality, St. Clair (Mar.); 
Grenada, several examples with no more specific locality; 
Tobago (Apr.). GuatTemaa: Cayuga (Jan., Feb., Apr., 
May, June), Quirigué (Sept.). Costa Rica: Esperanza 
(May, Aug.). PanamA: Alhajuelo (Apr.), Cabima 
(May), Porto Bello (Apr., Oct.), Rio Trinidad (Mar., 
June). Cotomera: La Esperanza (Dec.), no specific 
locality (June). Bonfivia: “East Bolivia’ (Oct.). 
British Guiana: ‘“‘Mazaruni Clearing’? (Aug., Oct.). 
Frenca Guiana: Cayenne, St. Jean Maroni, St. 
Laurent Maroni. Braziu: Rio de Janeiro (June). 

This species is primarily a feeder in the pods of the 
cacao and is well distributed in tropical America wher- 
ever its host occurs. All specimens in the National 
Museum (except the holotypes of muriscis and palpalis) 
had been identified by Dyar as ‘‘Myelois transitella 
Walker.” The two species are easily confused on super- 
ficial characters, especially among faded and stained 
tropical specimens; but their genitalia are quite distinct. 

Dyar’s types of muriscis and palpalis are males and 
alike in genitalic and all other characters. It is very 
probable that muriscis eventually will prove to be no 
more than a variety (or synonym) of furvidorsella 
Ragonot. 


46 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


81. Ectomyelois furvidorsella (Ragonot), new combination 
Figure 681 


Myelois furvidorsella Ragonot, Nouv. Gen., p. 8, 1888; Mono- 
graph, pt. 1, p. 56, 1893. 

This is probably the same as muriscis. 'The genitalia 
of the female type (fig. 681) are somewhat unusual in 
that bursa and ductus bursae are perfectly smooth, with 
no trace of signum or sclerotization of the ductus at 
genital opening. However, I have seen similar geni- 
talia in typical examples of muriscis from Central 
America; but I have seen so few examples of muriscis 
(only males) from Puerto Rico that I prefer to keep the 
names apart till more material is available. 

Alar expanse, 22 mm. 


TypE Locauity: Puerto Rico (type in Paris Mus.). 
Foop PLant: Unknown. 


82. Ectomyelois zeteki, new species 
FiaureEs 201, 682 


Forewing pale brownish gray faintly dusted with 
blackish fuscous; antemedial line obscure, angulate, 
indicated chiefly by its blackish fuscous outer border, 
the latter incomplete on many specimens; subterminal 
line sinuate, rather close to termen, on well-marked 
specimens consisting chiefly of white spots on the veins, 
preceded and followed by dark streaks, obscure on 
many specimens; also on well-marked specimens a me- 
dian, longitudinal blackish fuscous streak from base of 
wing to antemedial line; discal dots at end of cell very 
faint or completely obliterated, when distinguishable 
more or less confluent; terminal dark dots faint, con- 
fluent. Hind wing translucent, white with a faint 
smoky tint towards apex; a dark line along outer margin 
and some darkening of the veins. Highth abdominal 
segment of male simple. Alar expanse, 17-22 mm. 

Male genitalia similar to those of muriscis except: 
Uncus narrower; apical projection of gnathos shorter, 
not reaching so far backward as apex of uncus; transtilla 
a narrow band forming a truncated arch with slightly 
concaved posterior margin; sclerotized costa of harpe 
projecting somewhat further at its apex. Female geni- 
talia with scobinations of signum patch stouter and 
sclerotization of ductus bursae at genital opening fore- 
wing a larger and more strongly pigmented shield than 
those of any other species of the genus. 

Type tocatity: Near Capira, Panama (type in 
USNM, 61316; paratypes in USNM, Cornell Univ., 
Transvaal Mus. (Janse Coll.), Paris Mus., BM). 

Foop puant: Cassia moschata. 


Described from male type, and 16 male and 19 
female paratypes from the type locality; all reared (May 
1941, Zetek No. 4807) by James Zetek, who has con- 
tributed much valuable material to the National Collec- 
tion, and for whom the species is named. 

It is easily distinguished from any other American 
phycitid by its male genitalia. 


Genera 22-24: Paramyelois to Protomoerbes 


[Venational division C. Forewing with 11 veins; 4 and 5 stalked 
for less than half their lengths. Hind wing with veins 7 and 8 
anastomosed for most of their lengths (free element of 8 short). 
Transtilla incomplete; but (except in Paramyelois) its free 
elements well developed.] 


22. Paramyelois, new genus 


Type or Genus: Myelois solitella Zeller. 

Tongue well developed. Antenna of male shorily 
ciliate (cilia shorter than width of shaft), simple; of 
female pubescent. Labial palpus oblique, laterally flat- 
tened (broad and flat from lateral view) ; second segment 
roughly scaled beneath; third segment shorter than 
second, somewhat roughly scaled. Maxillary palpus 
squamous (rather heavily and broadly scaled). Fore- 
wing smooth; 11 veins; vein 2 from before but rather 
near lower outer angle of cell; 3 from the angle, well 
separated from the stalk of 4-5 at base, but nearer to 
4—5 than to 2; 4 and 5 shortly stalked; 6 from below 
upper angle of cell, straight; 8 and 9 long stalked; 10 
from the cell, approximate to the stalk of 8-9 for a 
short distance from base; male without costal fold. 
Hind wing with vein 2 from well before lower outer 
angle of cell; 3 from the angle, closely approximate to 
or connate with the stalk of 4—5 at base; 4 and 5 nor- 
mally stalked for half or slightly less than half their 
lengths, rarely (in some small specimens) stalked for 
over half their lengths: 7 and 8 strongly anastomosed 
for most of their lengths, free element of 8 short; cell 
slightly over half the length of wing in male (as in 
Kuropean Myelois), half the length of wing in female; 
discocellular vein curved. Highth abdominal segment 
of male simple. 

Male genitalia with apical process of gnathos a short, 
rather broad, blunt hook, slightly notched at apex. 
Uncus with broad base; narrowed and triangulate just 
beyond; apex acutely rounded. Tegumen short and 
broad. Transtilla incomplete. Harpe very broad at 
base; costa strongly and broadly sclerotized and forming 
a broad, pointed projection before middle, not appreci- 
ably sclerotized beyond; sacculus large and strongly 
sclerotized; cucullus greatly reduced. Anellus a curved 
shield with long, strongly sclerotized, smooth, slender, 
tapering and pointed lateral arms. Aedeagus scobinate 
on one lateral edge at apex; penis with a few sclerotized 
wrinklings, otherwise unarmed. Vinculum stout, slightly 
longer than broad, truncate, scarcely tapering to broad 
terminal margin. 

Female genitalia with weak signum consisting of a 
cluster of rather coarse scobinations; finer scobinations 
scattered over the caudal half of bursa. Ductus bursae 
shorter than bursa, broadened and sclerotized on inner 
ventral and lateral surfaces towards genital opening; on 
inner dorsal surface behind the opening a pair of small 
sclerotized plates. Ductus seminalis from bursa near 
its junction with ductus bursae. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 47 


This genus is easily recognized and is distinguished 
from other sections of the ‘‘Myelois’’ complex by its 
labial palpi and male genitalia. 

In their normal position the palpi are directed in a 
straight line obliquely from the face; but many speci- 
mens show the third segment more or less deflected for- 
ward, and some with both the second and third seg- 
ments more or less porrected, results of the death con- 
tortions of the moths. Several European species listed 
under Myelois have oblique palpi but they are all more 
or less cylindrical and do not have the broadly scaled 
and flattened lateral aspect of those of Paramyelois. 
The male genitalia with their incomplete transtilla (its 
elements reduced and well separated) are unique among 
the groups nearly related to Myelois or any of the Ameri- 
can species that previously have been referred to that 
genus. 

I have chosen a synonym as type of the new genus 
advisedly, as the type specimen of the oldest name 
(transitella) is a female, and there may be some question 
of my application of the name to the species here treated. 
There can be no such doubt in regard to solitella. 


83. Paramyelois transitella (Walker), new combination 
Figures 202, 683 


Nephopteryx transitella Walker, List, pt. 27, p. 54, 1863. 

Nephopteryx notatalis Walker, List, pt. 27, p. 57, 1863. 

Myelois solitella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 217, 
1881.—Ragonot, Monograph, pt. 1, p. 55, 1893. (New 
synonymy.) 

Myelois duplipunctella Ragonot, N. Amer. Phycitidae, p. 3, 1887; 
Monograph, pt. 1, p. 56, 1893.—MceDunnough, Check list, 
No. 6059, 1939. (New synonymy.) 

Myelois transitella (Walker) Ragonot, Monograph, pt. 1, p. 42, 
1893 (in part)—McDunnough, Check list, No. 6058, 1939 
(in part). 

Myelois venipars Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 404, 
1914.—Mote, Monthly Bull. California Dep. Agr., vol. 11, 
p. 628, 1922.—Glick, Arizona Comm. Agr. and Hort., Four- 
teenth Ann. Rep., p. 78, 1922.—Essig, Insects of western 
North America, p. 708, 1929.—Hixon, Journ. Econ. Ent., 
vol. 27, p. 547, 1934. (New synonymy.) 

Emporia cassiae Dyar, Ins. Inse. Menstr., vol. 5, 91, 1917 
(new synonymy). 

Similar in color and maculation to Ectomyelois muris- 
cis except: Ground color on lower half of wing darker; 
the dark borders of the transverse lines and the discal 
dots decidedly darker, blackish; the white areas more 
strongly contrasted; the dark outer border of antemedial 
complete in most examples and enlarged below costa 
into a conspicuous blackish spot. Hind wings a clearer 
white on the males; more or less smoky on the females. 
Alar expanse, 15-28 mm. 

Genitalia as given for the genus. The male genitalia 
show little or no individual variation. Among the fe- 
males, however, there is considerable variability in mi- 
nor details, namely, the amount of sclerotization about 
genital opening and the amount of scobination in the 
bursa, but these are differences of no specific significance. 

TYPE Locauitigs: ‘United States,” probably Florida 
(transitella, in BM); Santo Domingo (notatalis, in BM); 
Colombia (solitella, in BM); Florida (duplipunctella, in 


Paris Mus.); Hermosillo, México (venipars, in USNM); 
Georgetown, British Guiana (cassiae, in USNM). 

Foop piants: Orange, grapefruit, peach, apple, dates, 
figs, Acacia farnesiana, Aesculus glabra, Cassia grandis, 
Genipa americana, Gleditsia triacanthos, Pithecolobium 
flexicaule, Robinia, Sapindus drummondii, Yucca, English 
walnut. These records from reared specimens in the 
U.S. National Museum. 

Distrisution: Unirep Srarzs: Arizona, Maricopa 
County (Dec.), Mesa (Nov.), Phoenix (Aug., Sept., 
Nov., Dec.), Tempe, Yuma; Texas, Anahuae (March), 
Brownsville (Dec.), Dallas (May), Fort Davis (Oct.), 
Harlingen (July), Hidalgo County (Apr.), Kerrville, 
Louise (Feb.), Mercedes (Feb.), Mission, San Antonio 
(May), San Benito (Sept.) ; Oklahoma, Stillwater (June) ; 
Louisiana, Forbing (Oct.); Alabama, Mobile (Apr.); 
Georgia, St. Simons; Florida, Orlando (Oct., Nov.), Vero 
Beach (Apr., May); North Carolina, Durham. CuBa. 
Dominican Repusiic. Mexico: Hermosillo, Oaxaca. 
GuaTEMAta: Cayuga (Mar., Apr.), Chejel (June). Pan- 
amMA: El Cermeno (Apr., June). Contompia. Braziu: 
Tapera (Pernambuco). Prrt: Lima, Rio Pacaya (June, 
July, Aug.). 

This species has attracted some attention in the south- 
west as a minor orchard pest and is known to economic 
entomologists as the “‘nayel-orange worm.” The larvae 
feed on the nuts, in the seed pods, or on the fruits of 
numerous trees but they seem to prefer the fallen and 
mummied fruits or the dry seed pods or injured or 
diseased fruits. Rarely do they attack sound fruit on 
the trees. They have been reported as infesting sound 
oranges, but such behavior is probably an exceptional 
departure from normal habit. 

The foregoing synonymy requires some comment. I 
have not seen the types of transitella or duplipunctella 
(both females), but from the original. descriptions and 
the Ragonot figures they cannot apply to anything else 
than the species we have hitherto known in the United 
States as venipars Dyar. Of the synonymy of venipars 
and solitella there is no possible doubt. The type of the 
latter is a male (not a female as stated by Zeller) and 
figures of its genitalia, supplied by Tams and Clarke, 
show agreement in every detail with those of venipars. 
Clarke has also furnished excellent photographs of the 
female types of transitella and notatalis and of the geni- 
talia of transitella. The latter show only trifling indi- 
vidual differences from the genitalia of Dyar’s female 
type of venipars. Unfortunately, the type of notatalis 
lacks an abdomen; but photographs of the moths and 
their palpi show no essential differences between the two 
types; so Ragonot’s reference of notatalis to synonymy 
must be accepted. In his specific key to the species of 
“Myelois” Ragonot (Monograph, pt. 1, p. 27) places 
transitella in a group with veins 7 and 8 of hind wing 
approximate. This characterization was obviously 
based upon a freak specimen. Dyar found one such 
freak (a female from Grenada) among the examples of 
muriscis which he misidentified as “‘transitella”’; but in 
over a hundred examples of the true transitella before 
me veins 7 and 8 are strongly anastomosed, and this is 


48 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


a good character of much more than specific value 
despite its lapse in individual specimens. Freaks of all 
kinds can and do turn up anywhere in the Phycitinae. 
Dyar’s cassiae was described from stained and faded 
females; but, even so, it is strange that he did not see 
their resemblance to his venipars, especially in their 
palpi, and still more strange that he should refer them 
to the Old World anerastiid genus H#mporia. They have 
normal phycitine tongues and their genitalia agree with 
those of the female type of venipars. 

Bondar’s (Instituto de Cacau da Bahia Boletim 5, 
p- 72, 1939) identification of a lepidoperon in cacao pods 
as duplipunctella Ragonot (the genus given as “Myel- 
osis’”’) is probably incorrect. What he had was pre- 
sumably Hetomyelois muriscis. 


23. Genus Pseudodivona Dyar 


Pseudodivona Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 405, 1914. 
(Type of genus: Pseudodivona commensella Dyar.) 

Tongue well developed. Antenna shortly ciliate on 
male, cilia about as long as width of shaft (longer on 
carabayella). Labial palpus oblique, broadly scaled and 
laterally flattened; third segment short, acuminate. 
Maxillary palpus squamous. Forewing smooth; 11 
veins; vein 2 from before lower outer angle of cell; 3 
from the angle; 4 and 5 stalked for slightly less than 
half their lengths, the shaft separated at base from 3; 
6 from below upper angle of cell, slightly curved towards 
base; 8 and 9 stalked for two-thirds their lengths; 10 
from the stalk of 8-9; 11 from cell rather near outer 
angle and running close to the stalk of 8-9-10; male 
without costal fold. Hind wing from well before lower 
outer angle of cell; 2 from very close to the angle, or 
from the angle (cispha), closely approximate to or con- 
nate with stalk of 4—5 at base; 4 and 5 stalked for about 
half their lengths; 7 and 8 anastomosed for most of their 
lengths (free element of 8 very short); cell less than one- 
half (but more than a third) the length of wing; disco- 
cellular vein curved. Highth abdominal segment of 
male with a strong pair of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos de- 
veloped as an elongate, stout, flattened hook with 
forked or notched apex. Uncus subtriangulate, with 
rounded terminal margin. Transtilla incomplete; its 
elements well developed, elongate and knobbed at their 
apices. Harpe simple with outer margin evenly rounded; 
costa sclerotized for about four-fifths its length; but not 
produced. Anellus a narrow curved band with slender 
lateral lobes. Aedeagus moderately slender, nearly 
straight; penis unarmed. Vinculum stout, decidedly 
longer than broad, tapering, expanded towards angulate 
terminal margin. 

Female genitalia without signum; bursa, ductus bur- 
sae, and genital opening simple; ductus seminalis from 
bursa near junction of bursa and ductus bursae. 

In genitalic and many other characters as well as 
wing pattern and color, this genus resembles Moerbes, 
to which it is apparently closely related. It differs 
chiefly in having vein 4 present and well developed in 
hind wing, a different development of the elements of 


transtilla, and strong hair tufts on the eighth abdominal 
segment of the male. Four tropical American species 
are recognized. How many of these are really distinct 
species it is impossible to determine from the scanty ma- 
terial available. Nothing is known of their biology or 
habits. 


84. Pseudodivona commensella Dyar 
Fieures 41, 204 


Pseudodivona commensella Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 406, 1914. 

Forewing dull white on area above lower margin of 
cell; the area from cell to inner margin a glossy vinous 
brown; a blackish brown smudge on costa at base; 
antemedial line obscure except where it cuts the brown 
shade and forms a contrasting white spot at inner mar- 
gin, bordered outwardly by a blackish brown, out- 
wardly angled line, obscure on all but fresh specimens 
and frequently broken into two blackish spots, one on 
costa, the other in the cell; subterminal line indicated 
by a pair of narrow and narrowly spaced blackish dashes 
from costa near apex and some inwardly bordering 
blackish spots or streaks on the veins; the veins other- 
wise more or less darkly streaked; lower discal spot at 
end of cell enlarged, blackish, more or less confluent 
with a smaller, much fainter upper spot; a row of black- 
ish dots along termen. Hind wing pale, glossy, semi- 
translucent grayish white; veins darkened and clearly 
outlined; a narrow dark line along termen. Alar 
expanse, 20-22 mm. 

Male genitalia with trifling differences in the shape of 
the apical projection of gnathos between this and the 
following species of the genus and some differences in 
the curve of the outer margin of the harpe, but I suspect 
that these differences are individual rather than specific 
in character. 

TypEr Locatity: Jalapa, México (type in USNM). 

Foop Prant: Unknown. 

Distrisution: Mexico: Jalapa, Orizaba. 

Known only from the four males of Dyar’s type 
series. 

85. Pseudodivona cispha Dyar 


Fieure 205 
Pseudodivona cispha Dyar, Ins. Insc. Menstr., vol. 7, p. 53, 1919. 


A smaller, less distinctly marked species than the 
preceding one (commensella); the brownish area of 
forewing narrower and paler (not “reddish” as stated 
by Dyar); the dark markings fewer, fainter and paler 
and, except for the spot on base of costa and a short 
wedge at apex, not blackish; discal dots inconspicuous, 
light brown, the lower dot much smaller than on com- 
mensella. Alar expanse, 16-18 mm. 

Female genitalia like those of P. santa-maria. 

TyprE LocaLity: Volcin Santa Maria, Guatemala 
(type in USNM). 

Foop PLaAnt: Unknown. 

DistrisutTion: GUATEMALA: Cayuga (Aug.), Voleén 
Santa Maria (July, Oct.). Costa Rica: Tuis (May). 
British Honpuras: Rio Grande (Sept.), Punto Gorda 
(July). 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 49 


86. Pseudodivona santa-maria Dyar 
Ficure 690 


bea cee santa-maria Dyar, Ins. Insc. Menstr., vol. 7, p. 54, 
1919 

Known only from the two females of the type series. 
The coloration and markings are more like those of 
commensella except that the dark striping of the veins is 
fainter and the discal dots smaller, paler and less con- 
spicuous, like those of cispha. It is quite possible that 
these specimens are only larger, darker, better marked, 
female examples of cispha and equally possible that 
both cispha and santa-maria are only varieties of com- 
mensella. Alar expanse, 21 mm. 


Type tocatity: Volcfn Santa Maria, Guatemala 
(July; type in USNM). Paratype from Cayuga, 
Guatemala (May). 

Foop pLant: Unknown. 


87. Pseudodivona carabayella Dyar 
Ficurss 206, 691 


Pseudodivona carabayella Dyar, Ins. Insc. Menstr. vol. 7, p. 54, 
1919 

Larger and more strikingly marked than any of the 
preceding species. Forewing with pale areas pure 
white; lower area of wing (between cell and inner mar- 
gin) vinous brown (not “purplish red’’ as in Dyar’s 
original description) except for an extension of the 
white behind the antemedial line where it reaches 
almost to inner margin; an elongate black patch on 
costa at base; antemedial line indicated by the usual 
white spot on inner margin and its black outer border, 
the latter is rather broad from costa, strongly angled 
and extends from costa to the lower margin of the cell; 
a subbasal black spot in the cell; subterminal line indi- 
cated above cell only by its widely spaced black inner 
and outer borders; the inner black border a strong slant- 
ing black dash extending from costa almost to the lower 
discal dot at end of cell (indicating a deep angulation of 
the subterming line); the outer black border a much 
shorter, slanting, dash from apex to vein 6; from about 
vein 5 the subterminal line is indicated by a faint white 
line through the brownish ground color and is bordered 
inwardly by a few rather faint blackish spots; discal 
spots, distinct, black and somewhat enlarged, the lower 
one particularly. Hind wing white, more or less tinted 
with grayish on some specimens; veins darkly outlined; 
a narrow dark line along termen. Alar expanse, 23-28 
mm. 
Apical process of gnathos of male genitalia figured 
from type. Another male from Incachaca, Bolivia (in 
BM), exhibits some variation from the type in the shape 
of the apical process of gnathos (fig. 206a). It is a 
small specimen (23 mm.) and seems to have the dark 
areas and markings of forewing paler, but it is a rubbed 
and faded example; other specimens from the Schaus 
Collection in the National Museum and from the same 
Bolivian locality are typical in all details. The British 
Museum specimen is probably nothing more than an 
individual variant. 


TYPE LocALITy: Oconeque, Carabaya, Peri (type in 
USNM). 

Foop pLrant: Unknown. 

DistrisutTion: Pert: Carabaya, Oconeque, Tinguri. 
Bourvia: Cochabamba,¥Incachaca. Conompra: San 
Antonio (Dec.). 

All Peruvian examples in the U.S. National Museum, 
British Museum, and Janse Collection are males. The 
only female of the species that I have seen is the 
specimen from Incachaca, Bolivia, from which the 
genitalia are figured. 

The species is undoubtedly a distinct one. The 
widely spaced, strong, black dashes bordering the sub- 
terminal line indicate this as well as the ciliations of the 
male antenna, which are longer than those of any of 
the preceding species, being somewhat longer than the 
width of the antennal shaft. 


24, Protomoerbes, new genus 


Typr or genus: Protomoerbes aberrans, new species. 

Characters of Pseudodivona except: Labial palpus 
upturned (but otherwise as in Pseudodivona) ; forewing 
with vein 3 closely approximate to the stalk of 4-5 at 
base, male with narrow costal fold; hind wing with veins 
4 and 5 stalked for at least three-fourths of their length, 
cell one-third the length of wing; eighth abdominal 
segment of male without hair tufts; transtilla of male 
genitalia incomplete, its elements elongate-angulate, 
their apices not knobbed or expanded. 

In many details this genus is more like Moerbes of 
Group II than Pseudodivona. It differs from both 
genera in its upturned rather than oblique palpi. The 
cilia of the male antenna are also shorter (slightly less 
than the width of the shaft), but this is hardly a generic 
character. Wing pattern, color, and general habitus 
are like those of both Pseudodivona and Moerbes. All 
three have the contrasting white spot on inner margin 
of forewing indicating the base of the antemedial line. 
Protomoerbes in every way seems to be an intermediate 
and connecting link between Pseudodivona and Moerbes. 

It is represented by only two species from Colombia. 
Their females are unknown. 


88. Protomoerbes aberrans, new species 
Fiaure 208 


Forewing white; basal area, median area below cell 
and outer area below apex shaded with pale brown; a 
yellow longitudinal median streak from base to end of 
cell cutting the antemedial line; antemedial line a con- 
spicuous white spot on inner margin and a fainter white 
spot on costa, bordered outwardly below costa and on 
inner margin by blackish scaling and inwardly by a 
subcostal black streak reaching nearly to base of wing 
and by scattered black dusting at inner margin; veins 
and lower fold beyond antemedial more or less streaked 
or dusted with black, the black streaks especially marked 
and angulate at inner margin of subterminal line; lower 
discal dot expanded and extended along lower vein of 
cell, black; upper discal dot not distinguishable; sub- 


50 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


terminal white line narrow, faint, sinuate and sharply 
dentate below costa, bordered at costa by a pair of 
narrow, short, blackish dashes; a row of separate black 
dots along termen. Hind wing white, translucent. 
The veins faintly darkened and a narrow dark shade 
along termen. Alar expanse, 27-30 mm. 

Male genitalia with aedeagus smooth and evenly 
tapering from base. 

TypE Locauity: Colombia (type in USNM, 61317; 
paratype in BM). 

Foop prant: Unknown. 

Described from male type and two male paratypes 
labeled ‘‘Colombia, Fassel.’? The specimens are only 
in fair condition. Fresher examples would probably 
show the black longitudinal streakings somewhat more 
contrasted and extended. ‘The species has the general 
habitus of Pseudodivona commensella but is larger. It 
is easily separated from commensella and all the other 
similarly colored and marked species of Pseudodivona 
and Moerbes by the yellow longitudinal median streak 
on forewing. ‘This is easily distinguished under slight 
magnification, even on slightly rubbed specimens. 


89. Protomoerbes separabilis, new species 
Fieure 207 


Similar to aberrans except: Less distinctly and exten- 
sively streaked with black; median yellow longitudinal 
streak missing from forewing, replaced by a narrow 
extension of the white ground color, extended to the 
base of the wing; aedeagus with a thornlike projection 
from underside near apex (similar to the projection 
from the aedeagus of Moerbes emendata but larger and 
more blunt). The male genitalia also differ from those 
of M. emendata in having a proportionally longer and 
more evenly tapering vinculum. Hind wing smoky 
white, semitranslucent; the veins distinctly darkened. 
Alar expanse, 29 mm. 

TYPE Locatity: San Antonio, Colombia (type in BM; 
paratype in USNM, 61318). 

Foop prant: Unknown. 

Described from male type and one male paratype 
from the type locality, labeled ‘San Antonio, W. 
Colombia, Dec. 07, 5800 feet, M. G. Palmer.’ The 
paratype lacks an abdomen. 


Genera 25 and 26: Diatomocera and Pseudocabima 


[Venational division C. Forewing with 11 veins; 4 and 5 stalked 
for less than half their lengths. Hind wing with veins 7 and 8 
anastomosed for most of their lengths (free element of 8 short). 
Transtilla complete but weakly sclerotized. Uncus spoon- or 
semispoon-shaped.] 


25. Genus Diatomocera Ragonot 


Diatomocera Ragonot, Monograph, pt. 1, p. 250, 1893. 
genus; Homoeosoma tenebricosa Zeller.) 

Cabima Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 329, 1914. 
(Type of genus: Cabima dosia Dyar. New synonymy.) 


Tongue well developed. Antenna of male shortly 
ciliate (the cilia about as long as width of shaft); the 


(Type of 


shaft with notch at base; of female simple and pubes- 
cent. Labial palpus of male upturned, reaching to or 
almost to vertex, slender; third segment nearly as long 
as second, acuminate; palpus of female oblique and 
slightly longer than that of male. Maxillary palpus 
filiform. Forewing smooth; 11 veins; vein 2 from well 
before lower outer angle of cell; 3 from the angle; 4 and 
5 shortly stalked (for half or less than half their lengths), 
the stalk separated from 3 at base; 6 from below upper 
angle of cell, very slightly bent towards base; 8 and 9 
long stalked; 10 from the cell, but approximate to the 
stalk of 8-9 for some distance; male with an elongate, 
narrow costal fold. Hind wing with vein 2 from well 
before lower outer angle of cell; 3 from the angle, connate 
with the stalk of 4-5; 4 and 5 long stalked; 7 and 8 
anastomosed for most of their lengths beyond cell, free 
element of 8 short; cell about half the length of the cell 
on male, on female longer; discocellular vein curved. 
Highth abdominal segment of male with a pair of 
ventrolateral hair tufts. 

Male genitalia with apical process of gnathos U- 
shaped, consisting of a pair of short, blunt arms. Uncus 
spoon- or semispoon-shaped. ‘Transtilla complete but 
weakly sclerotized (except along its lateral edges), con- 
sisting of a broad, more or less finely scobinate plate; 
weakly attached to harpes. Harpe simple; costa sclero- 
tized for most of its length, but not produced. Anellus 
a curved plate with short lateral lobes. Aedeagus stout, 
straight (or but slightly bent near middle), more or less 
tapering to apex, moderately long; penis with a few 
weakly sclerotized wrinklings and more or less finely 
spined, otherwise unarmed. Vinculum stout, elongate, 
constricted towards angulate (or narrowly rounded) 
terminal margin. 

Female genitalia with bursa more or less finely 
scobinate; signa present, consisting of a cluster of two 
or more sclerotized disks; ductus bursae unsclerotized; 
genital opening simple; ductus seminalis from bursa in 
the neighborhood of the signa (sometimes between them 
and the junction of bursa.and ductus bursae, but not 
near the junction). 

The genus is readily recognized by its combination of 
male characters, the most striking feature of which is 
the spoon= or semispoon-shaped uncus which is found 
in only two other American genera—Pseudocabima, 
which lacks the antennal notch and long costal fold of 
forewing, and Entmemacornis, which has veins 4 and 5 
of hind wing completely united. Diatomocera is ap- 
parently confined to tropical America. Nine species 
are here recognized. They are represented by scanty 
and scattered material and nothing is known of their 
life histories or habits. From the greasy condition of 
some of the specimens it may be assumed that the larvae 
are borers. 

90. Diatomocera tenebricosa (Zeller) 
Fiaures 42, 209, 565, 686 


Homoeosoma tenebricosa Zeller, Horae Soc, Ent. Rossicae, vol. 16, 
p. 242, 1881. 

Diatomocera tenebricosa (Zeller) Ragonot, Monograph, pt. 1, p. 
250, 1893. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 51 


I have not seen any specimens from the type locality 
but have before me a male and female from French 
Guiana and a somewhat larger pair (20-21 mm.) from 
Costa Rica compared by Schaus with the type. They 
are either this species or varieties thereof. The follow- 
ing description is drawn from them. 

Ground color of forewing gray shaded with reddish 
brown between the veins; the veins themselves outlined 
with black, the blackish streaks broken by a very faint 
dull whitish antemedial line and by a more distinct 
subterminal line and more or less interrupted between; 
at extreme base the vein markings fused into a blackish 
patch; subterminal line close to outer margin, out- 
wardly angled between the fork of veins 4 and 5; discal 
dots small, separated, set obliquely, blackish; terminal 
dots faint. Hind wings grayish brown, paler and semi- 
translucent on the male; the veins faintly darkened, 
brown; a narrow brown line along termen. Alar ex- 
panse, 16-21 mm. 

Male genitalia with uncus but slightly constricted 
near its middle (wider in this area than in any of the 
following species); penis finely spined. Female geni- 
talia with signum a small cluster of bluntly rounded, 
closely appressed disks. 

Typ tocatity: Honda, Colombia (type in BM). 

Foop prant: Unknown. 

Distrisution: Cotomsr1a: Honda. Frencu Guiana: 
Cayenne. Costa Rica: Juan Vifias (May), Sixaola 
River (Mar.). 

The species is easily distinguished from anything else 
in the genus by its smaller size and the shape of its 
uncus. There are several minor differences between the 
genitalia of specimens from French Guiana and Costa 
Rica; in the spacing between the apical prongs of gna- 
thos, in the shape of the terminal margin of vinculum, 
in the amount of spining on the median area of tran- 
stilla, and in the number of disks forming the female 
signum. These are shown in our figures. Such differ- 
ences, however, do not seem to be of anything more 
than individual or (at most) varietal significance. The 
foldings of the median (membranous) area of the tran- 
stilla shown in figure 209 are superficial and result from 
the manner in which the slide preparation was made. 


91. Diatomocera dosia (Dyar), new combination 
Fiaures 48, 210, 685 


Cabima dosia Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330, 1914. 


Forewing dull white; the veins streaked with black 
broken into short dashes and dots; antemedial line not 
defined except by the black vein streakings on outer 
margin; subterminal line faint, angulate, the apex of 
angle between the fork of veins 4 and 5; discal and ter- 
minal dots distinct, separate, black; a faint shading of 
ocherous fuscous scales above and below vein 1b at 
middle; costa at base black for most of the length of 
fold on male. Hind wing dull semitransparent white, 
slightly darker on female; the veins more or less out- 
lined in pale brown; a smoky shade along costa and a 


fine, brown line along termen. Alar expanse, 24-31 
mm. 
Male genitalia with penis finely granulate-scobinate. 
Female genitalia with signum a chain of bluntly pointed 
disks. 

Typ tocatity: Cabima, Panama (type in USNM). 

Foop piant: Unknown. 

Known only from the type series (nine specimens) 
from the type locality (May). 


92. Diatomocera excisalis (Hampson), new combination 
Ficurzs 211, 687 


Crocidomera excisalis Hampson, Ann. Mag. Nat. Hist., ser. 10, 
vol. 4, p. 353, 1929. 

Similar to dosia Dyar and probably no more than a 
variety of that species. Distinguished by its generally 
smaller size, some trifling differences in male genitalia, 
and a shorter chain of disks forming the signum of the 
female. 

Alar expanse, 23-25 mm. 

Male genitalia considerably smaller than those of 
dosia but otherwise similar. 

Typ LocaLity: Cayenne, French Guiana (type in 
BM). 

Foop piant: Unknown. 

DistrisutTion: FrencH Guiana: Cayenne, St. Lau- 
rent Maroni. 

Also before me is a female from the unidentified ma- 
terial of the British Museum from eastern Bolivia 
(“‘Aug.—Oct., 1920, T. Steinbach’’) which is superficially 
a very good match for the female paratype of excisalis 
and may be a variety of that species. Unfortunately 
it lacks an abdomen, so positive identification cannot 
be made. 


93. Diatomocera decurrens (Dyar), new combination 
Ficure 212 


Cabima decurrens Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 330, 
1914, 

Forewing ‘‘luteous gray’ (the ground color of a dis- 
tinctly yellowish tint); black markings on veins as in 
excisalis but especially strong along vein 1b; the sub- 
terminal line somewhat more distinct. Hind wing semi- 
transparent white with a very faint ocherous tint; veins 
very faintly darkened; a narrow, pale brown line along 
termen. Alar expanse, 21-28 mm. 

Male genitalia distinguished from those of dosia and 
excisalis by its much narrower (more constricted) 
vinculum; penis with a few weak, minute scobinations. 
The sternite of eighth abdominal segment is also differ- 
ently shaped from that of dosia or excisalis. 

Type Locauiry: Rio Trinidad, Panamé (type in 
USNM). 

Foop prant: Unknown. 

DistriputTion: Panami: Corazal (Mar.), La Chorrera 
(May), Rio Trinidad (Mar.). 

A distinct species easily distinguished by its male gen- 
italia and ocherous forewing. Known only from males 
of the original type series. I fail to see the difference 


52 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


in the antennal notch from that of dosia which Dyar 
mentions. His examination of the material before him 
was obviously cursory for he refers to three of the La 
Chorrera specimens as ‘“‘females.’’ 


94. Diatomocera majuscula, new species 
Fiaure. 213 


Forewing pale, dull, brownish gray; the veins outlined 
by dark brown; entire basal area to antemedial line suf- 
fused with blackish brown; antemedial line indicated by 
three detached dull-white marks, a spot on costa, a 
smaller one in cell, and a third, somewhat diffused, white 
smudge on inner margin; subterminal line indicated by 
a white spot on costa, a white spot on inner margin and 
a much fainter, pale, outwardly curved line cutting the 
darkened veins from vein 1 to vein 5; discal spots dis- 
tinct, separated, black. Hind wing white, transparent; 
the veins very slightly darkened; a gray brown shade 
along costa, and a narrow pale-brown line along termen. 
Alar expanse, 32 mm. 

Male genitalia distinguished chiefly by their large 
size. Highth segment tufts of abdomen also more ro- 
bust than those of any other species of the genus. 

Typz Locatiry: Ponta Nova, Rio Xingu, Amazonas, 
Brazil (type in USNM, 61319). 

Foop prant: Unknown. 

Described from male type, from the Dognin Collec- 
tion in the U. S. National Museum, that had been iden- 
tified as Sematoneura atrovenosella. The species can be 
easily identified by its large size and the contrasted, 
blackish basal area of forewing. 


95. Diatomocera albosigno, new species 
Figure 214 


Forewing dull ocherous brown; the veins outlined in 
black; broken outwardly by a faint subterminal line 
which is indicated by a small white spot on costa, a 
smaller, much weaker spot on inner margin and a very 
faint, pale outcurved line between veins 1 and 6; ante- 
medial line replaced by a large white blotch, as broad as 
long and extending from inner margin to or almost to 
costa, bordered outwardly by a narrow blackish line; 
discal spots replaced by a narrow blackish line along 
the discocellular vein, Hind wing dull, smoky white, 
semitranslucent; veins faintly darkened; a thin fuscous 
line along termen. Alar expanse, 21-23 mm. 

Typr tocanity: “S. E. Brazil” [probably Paran4] 
(type in BM; paratype in USNM, 61320). 

Foop prant: Unknown. 

Described from male type and one male paratype 
from the type locality labeled ‘‘S. E. Brazil, E. D. Jones, 
1920-303.”’ Hasily recognized by the large white spot 
on the subbasal area of forewing; the only species of 
Diatomocera so marked. 


96. Diatomocera hoplidice (Dyar), new combination 
Fieure 215 


a 
Cabima hoplidice Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330” 
1914. 


Forewing dark gray; unicolorous except for faint 
darker (blackish) shading on some of the veins and a 
dark edging to the costa; antemedial and subterminal 
lines obsolete; discal dots replaced by a very faint dark 
line along the discocellular vein. Hind wings very dark, 
smoky gray (almost black). Alar expanse, 26 mm. 

Male genitalia with aedeagus broad and abruptly 
tapered toward apex. 

TypE LocALity: Puerto Bello, Panamé (Mar.; type 
in USNM). 

Foop prant: Unknown. 

Known only from the male type. Hasily recognized 
because of its nearly uniform dark coloration and lack 
of any transverse markings on forewing. 


97. Diatomocera extracta, new species 


FI@urREsS 217, 688 


Ground color of forewing gray with a faint ocherous 
tint, especially in basal area; veins darkened; antemedial 
line well out, near middle of wing, vertical, bordered 
outwardly by a narrow reddish brown line and in- 
wardly by a fainter line of the same color; subtermina 
line indicated only by breaks in the dark lining of some 
of the veins; discal spots replaced by an oblique brown 
line along discocellular vein. Hind wing semitranslu- 
cent, white with a very faint grayish ocherous tint, 
slightly darker on female; veins very faintly darkened; 
a fine dark (brownish) line along termen. Alar expanse, 
21-25 mm. 

Male genitalia with harpe narrow, not expanded to- 
ward outer margin; aedeagus slightly bent; penis armed 
with an elongate, dense cluster of slender dark spines 
and a scattering of fine scobinations. Female genitalia 
with bursa transversely elongate, weakly scloerotized in 
the area about the signa and junction of ductus semin- 
alis, also finely scobinate in this area, the scobinations 
extending into the ductus bursae; signa consisting of 
two or three very narrow, thin, elongate disks. 

TypE Locatity: Tuis, Costa Rica (type in USNM, 
61321). 

Foop pirant: Unknown. 

Described from male type from the type locality, col- 
lected by W. Schaus (“May 28—June 4”), and one male 
and one female paratypes, collected by Schaus and 
Barnes (Nov.). 

The species is closest to but quite distinct from moch- 
lophleps Dyar. Most nearly resembles pale or faded 
examples of Pseudocabima rubrizonalis (Hampson), with 
which it was confused in the National Collection. 


98. Diatomocera mochlophleps (Dyar), new combination 
Fieurss 216, 689 
Cabima mochlophleps Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 404, 1914 
Forewing purplish gray with faint pale reddish brown 
shading between the veins; the area from base to near 
middle slightly paler, its outer margin faintly indicating 
an antemedial line, somewhat curved and inwardly 
oblique from costa; subterminal line well inward from 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 53 


termen, angulate with apex of the angle within the fork 
of veins 4—5, appears denticulate due to pale interrup- 
tions on the veins, preceded by short black dashes on 
the veins; veins otherwise faintly blackish; a distinct 
and characteristic discal mark beginning as a black 
streak or dot at lower outer angle of cell and continued 
as a curved line along discocellular vein and for a short 
distance inward along upper vein of cell; a row of black 
dots along terminal margin. Hind wing translucent, 
white; the veins darkened, a smoky shade along costa 
and a narrow blackish line along termen. Alar expanse, 
31 mm. 

Male genitalia similar to those of extracta, except api- 
cal part of uncus broader and vinculum stouter. Fe- 
male genitalia with bursa globular; the signa consider- 
ably larger and more triangulate than those of extracta. 


Typr Locauity: Zacualpan, México (type in USNM). 

Foop prant: Unknown. 

In addition to the male type and a female from the 
type locality there is before me a smaller (27 mm.) male 
from the Janse Collection labeled simply ‘‘Mexico.” It 
is more suffused, lacks the pale basal shade on forewing, 
and shows scarcely any trace of asubterminal line. The 
black discal mark, however, is present and strongly con- 
trasted. This is the characteristic feature of the species. 
It resembles an inverted comma, with the tail pointed 
towards the base of the wing. 


26. Pseudocabima, new genus 


Typs or GENUS: Myelois euzopherella Dyar. 

Characters of Diatomocera except: No notch in shaft 
of male antenna; forewing of male without costal fold; 
hind wing with vein 3 frequently stalked with the stalk 
of veins 4-5 (apparently not a constant specific char- 
acter). In one species (rubrizonalis) the apical process 
of gnathos differs from that of any other species of either 
Pseudocabima or Diatomocera in that it is developed as a 
flattened hook with cleft apex and not as a U- or V- 
shaped pair of prongs. Such a departure from type is 
unusual within generic limits, but of no more than spe- 
cific significance, because the species otherwise is per- 
fectly normal. 

The new genus is proposed with great reluctance; for 
its species are very closely related to the bulk of those in 
Diatomocera, though none has been previously associated 
with them; but some separation must be made if we are 
to define our superspecific groups with any exactness. 
Pseudocabima, Diatomocera, and Entmemacornis are all 
obviously closely related, but they are separable on con- 
sistent, if slight, structural differences. 

Ten species are here recognized as belonging to the 
genus. One of these is North American (arizonensis). 
The remainder are tropical and probably only a fraction 
of the species inhabiting Central and South America. 

Before me are five single examples of what appear to 
be as many new species. It seems advisable to leave 
them undescribed until more material is available, their 
sexes can be associated, and more is known about the 
individual and local ranges of variability within species 


of the genus. There seems to be some variability, both 
in color and in minor details of genitalic structure. 


99. Pseudocabima castronalis, new species 
Figures 218, 696 


Forewing gray, the ground color lightened by white 
dusting over much of the median area; antemedial line 
distinguishable throughout, whitish, nearly vertical, 
slightly out-bent from before middle of costa to middle 
of cell, thence slanting inwardly very slightly to vein 1b 
and thence outwardly to the inner margin, followed on 
inner margin by a dark blotch; subterminal line from 
outer fourth of costa, bluntly angulate at middle (the 
line more curved than angled between vein 6 and lower 
fold) ; discal dots replaced by a pale ocherous brown spot 
covering the discocellular vein; fainter extension of this 
brownish shade between some of the veins in outer area, 
especially near costa beyond and before the subterminal 
line; some blackish streaking on the veins, conspicuous 
as three short black streaks following the brownish dis- 
cal spot; hind wing smoky white, semitranslucent on 
the male; darker, more brownish on female; the veins 
but faintly darkened. Alar expanse, 23-27 mm. 

Male genitalia with aedeagus tapering to narrow 
apical end; penis without spining or scobinations. Fe- 
male genitalia with signum a curved chain of more or 
less bluntly pointed disks. 

Type Locauity: Castro, Parand, Brazil (type in 
USNM, 61322; paratypes in BM). 

Foop prant: Unknown. 

Described from male type from the type locality, and 
one male paratype and one female paratype labeled 
“S.E. Brazil, E. D. Jones, 1920-303,” the latter two from 
unplaced material in the British Museum. They are 
somewhat discolored and consequently appear more 
yellowish than the type, which is in better condition, 
showing no trace of grease. This specimen was de- 
posited in the National Museum in 1905 by Schaus as 
representative of a Hampson manuscript species. It 
bears the name label (in Hampson’s handwriting): 
“Coptarthria castroalis Hampson, Type o.’’ Apparently 
Hampson never published a description. 

The best character for recognition of the species is 
the ocherous-brown discal spot with its outwardly bor- 
dering contrasted black streaks. 


100. Pseudocabima fearnella (Schaus), new combination 
Figure 219 


Myelois fearnella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p. 245, 1913. 


Forewing gray, a reddish brown shade along lower 
fold and some dusting of the same color along inner mar- 
gin; antemedial line obsolete; subterminal line faint but 
distinguishable, bent as in castronalis, pale gray; veins 
discontinuously lined with black; discal spots black, 
separated or (at most) partially fused. Hind wing 
white, semitransparent; the veins slightly darkened; 
a darker more distinct line along termen. Alar expanse, 
19-23 mm. 


54 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Male genitalia differing only in trifling details from those 
of castronalis, apical part of uncus being slightly broad- 
er and a trace of fine scobination appearing on the 
penis. The female is unknown. 

TypE LocaLity: Avengarez, Costa Rica (July; type 
in USNM). 

Foop prant: Unknown. 

In addition to the male type there are before me two 
males from Cayuga, Guatemala (Feb., June). They 
are smaller than the type and somewhat rubbed and 
faded and, as a result, considerably paler. However, 
they agree in all other details. The species is evidently 
very close to castronalis. 


101. Pseudocabima guianalis, new species 
Fieurrs 220, 697 


Forewing pale brownish gray; some darker dusting 
for a wide area along inner margin; veins very faintly 
and irregularly outlined by dark scaling thickened at 
lower angle of cell, the curvation of the line inward; 
antemedial line obsolete, very faintly indicated on one 
or two specimens; subterminal line obscure, more 
sharply angulate than on preceding species. Hind 
wing translucent white; veins partially and faintly 
darkened; a pale smoky brown line along termen. Alar 
expanse, 25-33 mm. 

Male genitalia with uncus narrower in middle than 
that of castronalis or fearnella; penis armed with a 
cluster of very fine weak spines and scobinations. Sig- 
num of female genitalia a short cluster of blunt, thorn- 
like disks. 

TyPE Locatity: St. Jean Maroni, French Guiana 
(type in USNM, 61323; paratypes in BM and Cornell 
Univ.). 

Foop prant: Unknown. 

Described from male type and one male and two fe- 
male paratypes from the type locality, collected by 
W. Schaus; one female paratype from Rockstone, Hs- 
sequebo, British Guiana (Schaus); one male paratype 
from Tumatumari, Potaro River, British Guiana, June 
26, 1927 (Cornell Lot 760, sub. 114); one male and one 
female paratypes from Mackenzie, Demerara River, 
British Guiana, June 21, 22, 1927 (Cornell lot 760, sub. 
102, 104). One of the female paratypes from St. Jean 
Maroni bears a Hampson “cotype” label inscribed 
“Coptarthria guianalis.”’ Evidently another example 
of an undescribed Hampson species. 


102. Pseudocabima euzopherella (Dyar), new combination 
Figures 228, 693 


Myelois euzopherella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 326, 
1914. 


Forewing gray more or less dusted with reddish brown 
(the specimens inclined to be greasy, giving a distinctly 
brownish shade to the wing); transverse lines distinct, 
whitish; antemedial line nearly vertical, a slight bend 
in cell, a narrow dark, outer, bordering line; subterminal 
line with a similar dark inner border, slightly and 
bluntly angled or rounded at vein 5; the distinctive 


mark a round blackish brown smudge, touching outer 
margin of cell, consisting of a black discal dot surround- 
ed by blackish or brown smudges; some broken black 
streaks faintly indicated on the veins. Hind wing pale 
smoky fuscous, subtranslucent; veins darkened; a fine 
dark line along termen. Alar expanse, 19-24 mm. 

Male genitalia with central area of uncus very narrow; 
penis armed with numerous fine spines and scobinations. 
Female genitalia exhibiting no distinctive specific char- 
acters; signum a row of rather stout, thornlike disks. 

TypE tocaLity: Rio Trinidad, Panamé (type in 
USNM). 

Foop piant: Unknown. 

Distrisution: PanamMA; Cabima (May), Corozal 
(May), Tabernilla, Rio Trinidad (June). 

Vein 3 of hind wing seems to be consistently from the 
stalk of veins 4 and 5, though the amount of stalking 
varies in different specimens. In the forewing, 10 is 
from the cell, approximate to the stalk of 8-9 but not 
from it as stated by Dyar. 


103. Pseudocabima pombra (Dyar), new combination 
Fiaure 221 


Myelois pombra Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 326, 
1914. 

Forewing pale ocherous gray; transverse lines faint, 
whitish, the antemedial near middle of wing, vertical, 
the subterminal rather well back from termen, slightly 
angled at middle; no discal markings; some scattered 
dark (brownish) dusting on the veins. Hind wing con- 
colorous with forewing, semitranslucent; the vems not 
appreciably darkened. Alar expanse, 17 mm. 

Male genitalia similar to those of ewzopherella except 
smaller and stem of uncus more slender, differences of 
doubtful specific value. 

Typr Locatiry: Cabima, Panamdé (May; type in 
USNM). 

Foop prant: Unknown. 

Known only from the male type. Evidently close 
to and possibly only a pale suffused variety of ewzophe- 
rella. The type is somewhat rubbed. On a fresh 
specimen in better condition the dark shading on the 
veins of forewing presumably would be more apparent. 


104. Pseudocabima nigristrigella (Ragonot), new combination 
FIGures 222, 699 


Myelois nigristrigella Ragonot, Nouv. Gen., p. 7, 1888; Mono- 
graph, pt. 1, p. 41, 1893. 

Forewing white dusted with blackish, giving it an 
ashy gray color to the naked eye; under magnification 
some rust colored scaling between the veins especially 
above and along inner margin; antemedial line near 
middle of wing, nearly vertical (very slightly convex), 
thin, white, bordered outwardly by a narrow, strongly 
contrasted, blackish brown line; subterminal line cury- 
ing outward from costa to between veins 4-5, thence 
vertical to inner margin, white, bordered inwardly by a 
narrow blackish brown line; a black line along dis- 
cocellular vein and continued for a short distance along 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 55 


vein 3; a faint concentration of the black dusting on the 
other veins. Hind wing semitransparent, more or less 
smoky white; darker on female than on male; the veins 
darkened, a broad dark shade along costa and a narrow 
dark line along termen. Alar expanse, 22-24 mm. 

Male genitalia figured from type. On another male 
in the British Museum from the type locality (June) the 
stem of uncus is broader, the terminal part of vinculum 
less constricted and its terminal margin less sharply 
angled. (Another example of the amount of individual 
variation that may be expected within specific limits in 
this genus.) Female genitalia figured from specimen in 
British Museum. The signum consists of three closely 
grouped, moderately large, bluntly rounded disks. 

TypE Locauity: Rio Negro, Amazonas, Brazil (May; 
type in Paris Mus.). 

Foop puant: Unknown. 

Known only from the type locality. A distinct 
species easily identified by the contrasted black trans- 
verse lines on the ash gray forewing. 


105. Pseudocabima arizonensis, new species 
Friaure 698 


Similar in color and markings to the preceeding 
species (nigristrigella) but darker, the black dusting 
heavier (especially on basal area), giving the wing a 
coarser, more pepper-and-salt appearance; no rust- 
colored scaling between the veins; the antemedial and 
subterminal lines blackish bordered on both sides but 
the borders (especially of antemedial line) much less 
contrasted than in nigristrigella; subterminal line acute- 
ly angled at middle; somewhat expanded black streak- 
ing on veins 2, 3, the stalk of 4, 5, and sometimes 6 for 
a short distance from cell; terminal dots fused into a 
continuous black line along termen. Hind wing shining 
white on male, pale smoky white on female; veins very 
faintly darkened; a fine dark line along termen. Alar 
expanse, 24-27 mm. 

Male genitalia exhibiting no specific characters. 
Female genitalia with signum a compact cluster of 
numerous, closely appressed disks. 

Typr Locauity: Redington, Ariz. (type in USNM, 
61324; paratypes in Paris Mus., Cornell Univ., Trans- 
vaal Mus. (Janse Coll.), and BM). 

Foop prant: Unknown. 

Described from male type and 2 male and 6 female 
paratypes from the type locality; 8 male and 11 female 
paratypes from the Baboquivari Mts., Ariz. (June, 
July, Aug., Sept.); and 4 male and 1 female paratypes 
from Mohave County, Ariz. (Aug., Sept.). 

So far, this is the only species of the genus recovered 
from the United States. In general habitus it resembles 
Euzophera nigricantella Ragonot, also from Arizona. 


106. Pseudocabima expunctrix (Dyar and Heinrich), new combina- 
tion 


Figures 224, 692 


Myelois expunctrix Dyar and Heinrich, Proc. Ent. Soc. Washing- 
ton, vol. 31, p. 116, 1929. 


Forewing slate gray; some black scaling on the veins 
and (under magnification) a faint scattering of white 
scales over outer area; antemedial and subterminal 
lines and discal spots lacking; a row of black dots at 
the vein ends along termen. Hind wing semitrans- 
lucent white, a smoky shade at apex, along costa, and 
on the veins (especially of the female, the veins of the 
male wing not appreciably darkened); a fine dark line 
along termen. Alar expanse, 22-30 mm. 

Female genitalia with signum a long chain of pointed, 
thornlike disks. 

TypE LocALiTy: Baia, Brazil (type in USNM). 

Foop piant: “Stems of leguminous tree.’ 

Known only from the type series. The only reared 
species of the genus and the only one without any 
trace of transverse markings. One of the male para- 
types proves to be a specimen of Fundella argentina 
Dyar. In the forewing veins 4 and 5 are somewhat 
longer stalked than in other species of the genus except 
perrensiella, the stalking being for a half to slightly 
more than half their lengths. 


107. Pseudocabima perrensiella (Ragonot), new combination 
Fiecure 695 


Myelois perrensiella Ragonot, Nouv. Gen., p. 8, 1888; Mono- 
graph, pt. 1, p. 52, 1893. 

From Ragonot’s description and figure the species 
must be similar to erpunctriz except for the presence 
of a distinct pale subterminal line, a trace of an ante- 
medial line, and a thin dark line along discocellular vein. 
Veins 4-5 are ‘long-stalked,” as in exrpunctriz. Alar 
expanse, 28 mm. 

The female genitalia differ from those of all other 
described species of the genus in having a sharp, par- 
tially sclerotized, deeply wrinkled, and densely scobi- 
nate bend in ductus bursae near its junction with bursa 
copulatrix; the signum consists of a curved band of 
bluntly rounded, closely impacted, thornlike disks. 

Typr Locauity: Goya, Argentina (type in Paris 
Mus.). 

Foop prant: Unknown. 

Known only from the female type. 


108. Pseudocabima rubrizonalis (Hampson), new combination 
Fieurus 225, 694 


Crocidomera rubrizonalis Hampson, Ann. Mag. Nat. Hist., ser. 
10, vol. 4, 353, 1929. 

Forewing pale gray; the costal edge and the veins 
beyond cell purplish brown; the basal area suffused 
with some faint dark dusting on the male, considerably 
darker on the female; antemedial line at middle of wing, 
vertical or nearly so, dull white, bordered on inner and 
outer sides by reddish brown bands, the outer one the 
wider and somewhat broadened in cell; subterminal 
line faint, when distinguishable, grayish white, set well 
back from outer margin (the space between it and 
antemedial line correspondingly reduced), sharply out- 
angled, the apex of the angle in the fork of veins 4-5; a 
black line, slightly curved, along discocellular vein; 


56 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


terminal dots faint, more or less confluent. Hind wing 
(of male) translucent white, the veins pale brown in 
outer area; a fine brown line along termen. Alar 
expanse, 26-30 mm. 

Male genitalia with apical process of gnathos a 
single, narrow, flattened hook with notched apex. 

TypE Locatity: St. Jean Maroni, French Guiana 
(type in BM). 

Foop prant: Unknown. 


DistripuTion: French Gurana: Cayenne, St. Jean 


Maroni. Brazi: Parand, Taperinha. 

The foregoing description based on three males in 
the National Museum. What I take to be a female from 
Cayenne in the Janse Collection is also before me. It 
differs from the males in haying broader forewing, the 
antemedial line outwardly oblique from costa, the 
subterminal line more distinct, and the hind wing a 
glossy brown. ‘The bursa is large, as long as ductus 
bursae; the signum a single straight line of rather sharp, 
spinelike thorns. 


Genus 27: Hyalospila 


[Venational division D. Forewing with 11 veins; 4 and 5 closely 
approximate for some distance from cell. Hind wing with all 
veins very long; cell less than one-third the length of wing. 
Transtilla complete, developed as a narrow, slightly arched band.] 


27. Genus Hyalospila Ragonot 


Hyalospila Ragonot, Nouv. Gen. p. 11, 1888; Monograph, pt. 1, 
p. 168, 1898. (Type of genus: Hyalospila stictoneurella 
Ragonot.) 

Tongue well developed. Antenna of male simple; 
shaft very weakly pubescent. Labial palpus upcurved, 
reaching above vertex; slender; segment 3 as long as or a 
trifle longer than 2, acuminate. Maxillary palpus sub- 
squamous (scaling more or less dilated). Forewing 
smooth; 11 veins; vein 2 from before lower outer angle 
of cell; 3 from the angle, shortly separated from 4 at 
base; 4 and 5 closely approximate for some distance 
from cell (not stalked as stated by Ragonot); 6 from 
below upper angle of cell, straight; 8 and 9 stalked for 
not more (usually less) than half the length of 8; 10 
from the cell, approximate to the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from before 
lower outer angle of cell; 8 from the angle, closely 
approximate to the stalk of 4-5; 4 and 5 stalked for 
about half their lengths; 7 and 8 contiguous or very 
weakly anastomosed for not over half their lengths 
beyond cell; all veins very long; cell less than one-third 
the length of wing; discocellular vein curved. Highth 
abdominal segment of male with a pair of ventrolateral 
hair tufts. 

Male genitalia with apical process of gnathos an 
elongate, slender hook with slightly notched apex. 
Uncus triangulate. Transtilla complete, a simple, nar- 
row, slightly arched band. Harpe narrow; costa 
strongly sclerotized throughout and projecting slightly 


at apex; otherwise simple. Anellus a small shield with 
long, slender lateral arms. Aedeagus simple; penis 
armed with numerous fine scobinations or one or more 
clusters of slender spines. ’ 

Female genitalia with signa present, consisting of a 
single cluster of bluntly pointed thorns, frequently 
surrounded by fine scobinations or strongly pigmented 
granulations; bursa various, strongly granulate over 
mich of one side, weakly sclerotized towards or at,junc- 
tion with bursa, more or less finely scobinate or (except 
for signum patch)! smooth; ductus bursae weakly 
sclerotized towards genital opening or with genital 
opening simple; ductus seminalis from bursa near 
junction of bursa and ductus bursae. 

A distinct, easily recognized genus apparently 
limited to tropical America. The species also are easily 
identified by their genitalia. It is difficult to, place 
Hyalospila satisfactorily in any linear arrangement for 
it partakes of the characters of two distinct groups. On 
male genitalia, especially its complete bridgelike trans- 
tilla, it should go with the genera of the main Acrobasis- 
Myelois stem having this organ well developed, while 
on other characters, general habitus, and the long veins 
and short cell of hind wing it seems more closely related 
to genera of the Piesmopoda group. ; 


109. Hyalospila stictoneurella Ragonot 
Fiaures 44, 226, 703 


Hyalospila stictoneurella Ragonot, Nouv. Gen., p. 11, 1888; 
Monograph, pt.1, p. 169, 1893. 

Forewing purplish brown, a narrow ocherous-white 
band along costa, extending from costa to upper vein of 
cell and showing under magnification a scattered pow- 
dering of reddish scales; the extreme costal edge at its 
middle, darkened; the veins more or less streaked with 
black scaling; antemedial line indistinct, indicated by 
an oblique blackish streak from costa, a small whitish 
dot in cell and another on vein 1b, each followed by a 
black dot; subterminal line faint but distinguishable, 
close to and parallel with termen, not dentate; discal 
dots at end of cell confluent, blackish brown. Hind 
wing semitransparent whitish; the veins brown, a brown 
border along costa and a fine brown line along termen. 
Alar expanse, 19-22 mm. 

Male genitalia with transtilla a short, slightly arched 
bridge, somewhat wider than that of any other known 
species and with a minute, pointed projection at middle. 
Penis armed with three small clusters of fine, short 
spines. Vinculum but slightly longer than greatest 
width, tapering to its rather broad, rounded terminal 
margin. 

Female genitalia with bursa unsclerotized, weakly 
scobinate in the area about the signum; the latter a 
small patch of small, thornlike spines (fig. 703a) ; ductus 
bursae considerably longer than bursa, weakly sclero- 
tized for a short distance from genital opening and with 
a patch of scobinations at its middle, otherwise mem- 
branous and simple. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 57 


Type Locatity: Las Mercedes, ‘‘Amer. centr. mer.’ 
[Guatemala?] (type in Paris Mus.). 

Foop prant: Unknown. 

Distrisution: México: Jalapa, Misantla (May), 
Orizaba. Guatemata: Las Mercedes [?], Volcin Santa 
Maria (Aug., Oct., Nov.). Costa Rica: Juan Vifias 
(Nov.). Braz: Campo Bello, Santa Catarina (Oct.), 
“S. E. Brazil” [Paran4?]. 


110. Hyalospila celiella Schaus 
Fiaures 227, 700 


Hyalospila celiella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p. 248, 1913. 

Ground color in outer area between the veins and in 
entire area between lower margin of cell and inner mar- 
gin (except for vein 1b) pale ocherous brown; a narrow 
pale band along costa (fainter than that on stictoneurella, 
less whitish and, under magnification, showing a fine 
irroration of purplish brown scales, especially on costal 
edge) ; upper vein of cell white for a short distance from 
subbasal area; lower vein of cell white from base to end 
of cell except for a black spot near middle; a blackish 
brown line bordering the lower edge of cell from base to 
middle; a short white streak on vein 1b before middle, 
preceded and followed by blackish dots (these and the 
black spot on lower vein of cell all that remain to indi- 
cate an obsolete antemedial line); vein 1b otherwise 
more or less outlined by brown scaling; outer veins 
shortly streaked with blackish brown; subterminal line 
indicated only by an interruption of the blackish vein 
streaks near outer margin; discal spots a pair of short 
black streaklets at end of cell. Hind wing semihyaline, 
white with a very faint smoky tint in outer area; veins 
pale brown; a darker brown line along outer margin. 
Alar expanse, 19-21 mm. 

Male genitalia easily distinguished by the stout, 
greatly elongated vinculum. The abdominal tuft and 
sclerotized sternite of eighth segment similar to those 
of stictoneurella. 

Female genitalia with a weak sclerotization of bursa 
in the area surrounding the ductus seminalis and ex- 
tending for a short distance into the ductus bursae; no 
spining at middle of ductus bursae; otherwise as in 
stictoneurella. The sclerotization of the ductus bursae 
at genital opening varies individually in extent and 
amount as it does in stictoneurella. 

TyPE Locatity: Juan Vifias, Costa Rica (type in 
USNM). 

Foop piant: Unknown. 

Represented in the National Collection by the male 
holotype (Jan.) and two females (June, Nov.), all from 
the type locality. 


111. Hyalospila insequens, new species 


Similar to celiella except: Larger; ground color of fore- 
wing darker, some rosy scaling overlaying most of the 
pale area between cell and inner margin; dark streaking 


300329—56——_5 


on veins fainter; hind wing a clearer white, the veins 
not appreciably darkened. Alar expanse, 23-24 mm. 


Male genitalia similar to those of celiella except vin- 
culum not greatly elongated, slightly less than twice as 
long as its greatest width. Female unknown. 

Type tocauity: Incachaca, Cochabamba, Bolivia 
(type in USNM, 61325; paratype in BM). 

Foop puant: Unknown. 

Described from male type, collected by J. Steinbach, 
and one male paratype from San Antonio, western 
Colombia, 5,800 feet, Nov. 1907, M. G. Palmer, 
collector. 

The species has the same dark streak from base of 
forewing along the under edge of the cell as celiella, but 
not so strongly accented. The discal spots are slightly 
more pronounced and more or less confluent. The 
shorter vinculum at once distinguishes it. 


112. Hyalospila majorina, new species 
Fieure 701 


Forewing pale gray-brown; costal area to and includ- 
ing the cell dull white, the white shade narrowing 
gradually beyond cell to apex of costa; lower discal dot 
at end of cell enlarged, dark brown, completely en- 
circled by white; antemedial line obsolete; subterminal 
line very faint, close to and parallel with costa, indicated 
chiefly by short, whitish streaklets on a few of the upper 
veins and a slight paling of the ground color from vein 
4 to inner margin. Hind wing semitranslucent, whitish 
with a faint brown tint; the veins darker; a fine brown 
line along termen. Alar expanse, 27 mm, 

Female genitalia with ductus bursae considerably 
broadened for most of its length, wrinkled and weakly 
sclerotized on one side at junction with bursa; bursa 
finely scobinate over part of one side, the scobinations 
extending into ductus; genital opening simple. 

TypE Locatity: Misantla, México (type in USNM, 
61326). 

Foop prant: Unknown. 

Described from female type, collected by Robert 
Miiller, Sept. 1914 (“4362”), and one female paratype, 
the latter from Jalapa, México. The male is unknown. 
I should not have described a new species from females 
alone, but in this case a male from México should be 
easily matched. The females are readily distinguished 
by their large size and their genitalia. 


113. Hyalospila fulgidula, new species 
Fiaures 228, 702 


Ground color of forewing a clear bright white, clouded 
by a faint, pale drab shade along inner margin, this 
shade beyond lower outer angle of cell extending ob- 
liquely upward to apex; costa at base reddish; a minute 
red dot on costal edge at one-third and below it a con- 
spicuous, broad, oblique, black dash crosses the cell; 
below this one or two black dots on vein 1b; a short, 
black dash along lower margin of cell at base; some few 
scattered black scales on edge of inner margin and on 


58 UNITED STATES! NATIONAL MUSEUM BULLETIN 207 


some of the veins and a very sparse and scattered dust- 
ing of red scales in the outer white area; discal dots at 
end of cell separate, black, the lower conspicuous, the 
upper minute; subterminal line clearly indicated by a 
row of black dots along its inner border and an out- 
wardly bordering, short, black dash from costa; a row 
of detached black dots from vein 6 to lower fold along 
edge of termen. Hind wing glossy, smoky white, dark- 
ening outwardly and with a brown shade along outer 
margin. Abdominal tufts and eighth segment sternite 
of male as in stictoneurella. Alar expanse, 12-13 mm. 
Male genitalia with transtilla a very narrow, squarely 
arched band; penis very finely scobinate for about the 
length of aedeagus, otherwise unarmed. Female geni- 
talia with ductus bursae somewhat swollen and densely 
but minutely scobinate towards its junction with bursa, 
the scobinations extending on one side into bursa; 
ductus bursae also sclerotized for a short distance from 
genital opening, on its dorsal surface the sclerotization 
forming a shield projecting caudally beyond the opening. 


TYPE LOCALITY: Santiago Province, Cuba (type in 
USNM, 613827). 


Foop ruant: Unknown. 


Described from’ male type (June) and three female 
paratypes (June, Sept., Nov.) from the type locality, 
collected by W. Schaus. A distinct species, easily iden- 
tified by its small size, the squarely arched transtilla, 
and the bright white ground color and contrasted black 
spotting of its forewing. 


114. Hyalospila egenella (Ragonot), new combination 
Fieure 704 


Piesmopoda egenella Ragonot, Nouv. Gen., p. 11, 1888; Mono- 
graph, pt. 1, p. 165, 1893. 

Forewing grayish brown finely powdered with reddish 
scales; the costal area white with scattered red scaling; 
antemedial line not distinguishable; subterminal line in- 
dicated chiefly by its slightly darkened inner and outer 
borders, close to and parallel with termen, not sinuate. 
Hind wing smoky white, semitranslucent, the veins 
darkened and a dark line along termen; the smoky tint 
accented somewhat towards apical area. Alar expanse, 
15-20 mm. 

Female genitalia distinguished chiefly by the en- 
larged, wrinkled and scobinate ductus bursae and the 
enlarged blunt thorns forming the signum (fig. 704a); 
ductus bursae weakly and narrowly sclerotized at gen- 
ital opening. The male is unknown. 

Typ tocatity: Rio Negro, Brazil (type in Paris 
Mus.). 


Foop PLiant: Unknown. 


A small (15 mm.), somewhat rubbed female in the 
U. S. National Museum from Santa Catarina, Brazil 
(July), seems to be this species. It has genitalia similar 
to those of Ragonot’s type, differing only in minor indi- 
vidual details, a somewhat more extended scobination 
of ductus bursae, and faint traces of pale yellowish 
sclerotization in some of the folds at junction of bursa 


and ductus bursae. It also shows traces of a dark discal 
spot on forewing at lower outer angle of cell which Rag- 
onot states is absent from his type. None of these 
differences is significant. 


115. Hyalospila xanthoudemia (Dyar), new combination 
Figures 229, 709 
Piesmopoda xanthoudemia Dyar, Proc. U. 8. Nat. Mus., vol. 47, 
p. 333, 1914. 

Forewing olivaceous ocherous; the costal area beyond 
extreme base and including the cell and the outer area 
above vein 6, white with a scattered powdering of red- 
dish scales; extreme costal edge red, this shade espe- 
cially noticeable at base; antemedial line, discal and 
terminal dots obsolete; subterminal line faint but visible, 
whitish, very close to termen, not sinuate. Hind wing: 
whitish with a faint ocherous fuscous tint, especially in 
outer area, somewhat darker on female than on male; 
veins faintly darkened; a fine, pale brown line along 
termen. Alar expanse, 16-19 mm. 

Male genitalia with penis armed with a band (about 
one-third as long as aedeagus) of fine scobinations. Fe- 
male genitalia with bursa simple except for the signum 
patch; ductus bursae much longer than bursa, slender 
for most of its length, without scobinations and unsclero= 
tized except very weakly at genital opening. 

Typr Locaniry: Rio Trinidad, Panamé (type in 
USNM). 

Foop Pirant: Unknown. 

Distrisution: PanamAd: Chiriqui (April), Paraiso 
(Jan.), Rio Trinidad (Mar.). Costa Rica: Experenza 
(May), Juan Vifias (Oct.). 

Doubttully distinct from angulineella (Schaus). 


116. Hyalospila angulineella (Schaus), new combmation 
Fieure 707 


Piesmopoda angulineella Schaus, Ann. Mag. Nat. Hist., ser. 8, 
vol. 11, p. 246, 1913. 

Known only from the female type. Color and mark- 
ings as on zanthoudemia except for faint traces of an 
angulate dark antemedial line on forewing, and darker 
(pale smoky brown) hind wing. 

Ductus bursae of female genitalia longer than that of 
zanthoudemia. Otherwise the ductus shows but trifling 
differences which are somewhat exaggerated in the figure. 

TYPE LOCALITY: Juan Viiias, Costa Rica (June; type 
in USNM). 

Foop Puant: Unknown. 


117. Hyalospila clevelandella (Dyar) 
Fiaures 230, 705, 706 
Oryctometopia clevelandella Dyar, Proc. U. §. Nat, Mus., vol. 47, 
p. 331, 1914. ee 
Hyalospila clevelandella (Dyar) Dyar, Ins. Insc. Menstr., vol. 7, 
p. 48, 1919. : 
Forewing gray-brown from lower margin of cell and 
(in outer area) below vein 5, costal area white sparsely. 
irrovated with red scales; extreme base of costal edge 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 59 


brown; costal edge beyond more or less reddish; ante- 
medial line indicated only by fragments of its outer 
border, a couple of red dots or dashes in the white area 
and a faint, dark, gray-brown dot on lower fold of vein 
1b; discal dots at end of cell weak, lower brown, upper 
red; subterminal line faint but distinguishable, not sinu- 
ate, close to and parallel with termen, dull white. Hind 
wing smoky white with the veins and lower fold dis- 
tinctly darkened and a dark smoky shade along termen. 
Alar expanse, 13-16 mm. 

Male genitalia with vinculum sharply constricted into 
a digitate projection slightly beyond base; anellus an 
elongate, irregularly shaped, curved plate with elongate, 
very slender (almost threadlike) lateral lobes; penis 
armed with a single, dense cluster of dark brown, slender 
spines, the cluster as long or nearly as long as aedeagus. 

Female genitalia with ventral surface of half of bursa 
and ductus bursae covered with a mat of closely placed, 
pigmented granulations, the granulations extending 
around partly to dorsal surface; signum patch (on dor- 
sum of bursa) surrounded by a teardrop-shaped mass of 
granulations (fig. 705a); genital opening simple. 

Typr nocauity: Porto Bello, Panamé (type in 
USNM). 

Foop pLant: Unknown. 

Distrisution: PanamA: Porto Bello (Dec.), Taboga 
Isl. (Feb.), Tabogilla Isl. (Feb.). 

The species can be at once identified by its peculiar 
genitalia. The female paratype has genitalia similar in 
all but the most trifling details to those of semibrunneella 
Ragonot, and if it is actually conspecific with the males 
of clevelandella the latter name will fall as a variety or 
synonym of semibrunneella. However, there is some 
doubt that this is the case; for we have in the National 
Collection a series of four females from Cayuga, Guate- 
mala, and Jalapa, México, of the same size (16-17 mm.) 
and identical color and maculation as the female para- 
type of clevelandella, but with quite different genitalia 
(fig. 706). Also in the collection are four other females 
with the same color and markings and the same size as 
the males of clevelandella (13-14 mm.) but with differ- 
ent female genitalia. Either of these two groups of 
specimens could be the females of Dyar’s species so, for 
obvious reasons, I am not attempting to name them or 


to propose any synonymy. 


118. Hyalospila semibrunneella Ragonot 
Ficure 708 
Hyalospila semibrunneella Ragonot, Nouv. Gen., p. 12, 1888; 
Monograph, pt. 1, p. 169, 1893. 

Color and maculation similar to those of clevelandella 
except antemedial line more distinct. The female gen- 
italia agree substantially with those of the female para- 
type of clevelandella. Weshall have to wait discovery of 
a male of semibrunneella from the type locality before 
the status of the two supposed species can be determined. 

Type tocatity: “New Granada” [Colombia] (Mar.; 
type in Paris Mus.). 

Foop prant: Unknown. 


Genus 28: Fundella 


[Venational division B. Forewing with veins 4-5 connate or 
approximate at base. Hindwing with vein 3 from the stalk of 
4-5 or closely approximate to it for some distance; cell short; 
on male anal area folded into a pocket. Male genitalia with uncus 
hammer-clawed (long, curved, constricted at middle and broadly 
divided at apex); transtilla absent; sacculus of harpe not pro- 
duced; cornutus present, a single, strong spine. Eighth abdom- 
inal segment of male with pair of hair tufts.] 


28. Genus Fundella Zeller 


Fundella Zeller, Isis von Oken, 1848, p. 866.—Ragonot, Mono- 
graph, pt. 1, p. 210, 1893.—Janse, Journ. Ent. Soc. South 
Africa, vol. 4, p. 163, 1941.—Heinrich, Proc. U.S. Nat. Mus., 
vol. 96, p. 105, 1945. (Type of genus: Fundeila pellucens 
Zeller.) 

Ballovia Dyar, Proc. U. S. Nat. Mus., vol. 44, p. 8328, 1913; Ins. 
Inse. Menstr., vol. 7, p. 40, 1919. (Type of genus: Ballovia 
cistipennis Dyar.) 

Tongue well developed. Antenna of male weakly 
pubescent, basal segment somewhat enlarged, shaft 
laterally flattened and very slightly excavate at base 
(fig. 231e) (except on ignobilis and ahemora) and with a 
very small blackish scale tuft in the excavation (except 
on ignobilis); of female, slender, simple. Front of male 
head deeply grooved to hold labial palpi; of female, 
rounded. Labial palpus upcurved, reaching to vertex, 
clothed with broad appressed scales; on male closely 
appressed to face, with second segment over three times 
as long as first and with third segment very short (about 
one-sixth the length of second); on female with second 
segment shorter and third about one-third the length of 
second. Maxillary palpus minute, filiform. Forewing 
smooth; 11 veins; vein 2 from before lower outer angle 
of cell; 3 from the angle, approximately equidistant 
from 2 and 4; 4 and 5 connate or approximate at base; 
6 from below upper angle of cell, straight; 8 and 9 long 
stalked, 9 short; 10 from the cell, parallel for some dis- 
tance but not closely approximate to the stalk of 8-9. 
Hind wing with vein 2 from close to lower outer angle 
of cell; 3 from the stalk of 4—5 or closely approximate to 
it for some distance; 4 and 5 stalked for over half (about 
two-thirds) their lengths; 7 and 8 closely approximate 
beyond cell for less than half their lengths; cell short, 
about one-third the length of wing; discocellular vein 
curved; on male, anal area (involving veins la and 1b) 
thickened and folded under to form a pocket enclosing 
enlarged scales and hair tufts. Eighth abdominal seg- 
ment of male bearing a thin, short pair of ventrolateral 
hair tufts. 

Male genitalia with uncus long, curved, strongly scler- 
otized, constricted at middle and broadly divided at 
apex (hammer-clawed) ; gnathos terminating in a short, 
stout hook or a short, broad plate (achemora); transtilla 
absent. Harpe rather short, with clasper. Vinculum 
narrow, short. Aedeagus stout with long, stout, pro- 
jecting, curved spine or spines at apex (except in 
argentina); cornutus a single, strong spine. 

Female genitalia without signum (pellucens) or with 
signum well developed and consisting of a large oval or 
pear-shaped cluster of thornlike spines (argentina, aga- 


60 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


pella), or curved sclerotized bands armed with stout, 
thornlike spines (ahemora, ignobilis) ; bursa large; ductus 
bursae short, broad (narrowest in agapella); area sur- 
rounding genital opening strongly sclerotized, the dorsal 
sclerotization in the form of a band connected with the 
supporting rods of eighth segment collar, and armed 
with two or four spinelike projections (except in ignobilis 
and some examples of argentina); ductus seminalis from 
caudal area of bursa. 

This genus is easily distinguished by its striking male 
characters; the strongly sclerotized, long-stemmed, bi- 
furcate (hammer-clawed) uncus; the large pocket on 
anal area of hind wing; the long, embedded labial palpus 
with very short third segment; and minute maxillary 
palpus. A similar bifurcate uncus is not found in any 
other American genus except Difundella Dyar. In the 
type species of the latter (corynophora Dyar) the uncus 
is somewhat produced and exhibits a slight bifurcation 
at apex; but other species, which must also be referred 
to Difundella, lack this character. Difundella separates 
readily on other male structures—its strongly hooked, 
partially free sacculus of harpe, its rounded frons, and 
the narrow, strongly sclerotized, deeply invaginated 
pocket of the sternite of the eighth abdominal segment. 

In Fundella the wing pattern varies so much within 
any given species that it affords no reliable character 
for specific identification, and the several species can be 
separated with certainty only by their genitalia. 


119. Fundella pellucens Zeller 
Figures 6, 231, 713 


Fundella pellucens Zeller, Isis von Oken, vol. 41, p. 866, 1848; 
Horae Soc. Ent. Rossicae, vol. 15, p. 236, 1881.—Ragonot, 
Monograph, pt. 1, p. 210, 1893.—Heinrich, Proc. U. 8. Nat. 
Mus., vol. 96, p. 107, 1945. 

Ballovia cistipennis Dyar, Proc. U. 8. Nat. Mus., vol. 44, p. 323, 
1913. 

Fundella cistipennis (Dyar) Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 
1919.—Wolcott, Journ. Agr. Uniy. Puerto Rico, vol. 17, 

pp. 241-255, 1933; Journ. Agr. Univ. Puerto Rico, vol. 18, 
p. 432, 1934; vol. 20, p. 477, 1936.—Scott, Journ. Agr. Univ. 
Puerto Rico, vol. 24, pp. 35-47, 1940. 

Male antennal shaft with very small black basal tuft 
(fig. 231e). Forewing grayish fuscous more or less dusted 
with whitish and with interspersed reddish brown scales 
(in ‘many specimens the ground color is reddish brown), 
giving the moth a distinctly gray or gray-brown appear- 
ance to the naked eye; a conspicuous, round, darker 
brown or fuscous spot in the center of the area usually 
occupied by the antemedian line, this dark spot more or 
less obscured in some specimens, but in typical examples 
outlined by whitish areas inwardly and outwardly and 
not reaching to inner margin or costa of the wing; discal 
mark at end of cell obscure, often absent; subterminal 
line (when distinguishable) faint, white, indented at 
vein 6 and at submedian fold; a row of dark spots along 
termen (present only in specimens having an appreciable 
dusting of white scales). Hind wing white, translucent, 
a faint fuscous border along costa and (in some speci- 


mens) a fuscous line on termen for a short distance from 
apex; cilia white; anal pocket yellowish white. Mid- 
tibia with a fringe of pale hairlike scales along dorsum. 
Hind tibia with a rather long and slender tuft of pale 
(whitish ocherous), hairlike scales from the knee joint 
(fig. 231f). 

Female essentially like the male in color and markings 
except that the dark spot near the base of the forewing 
is more diffused, sometimes reaching to the costa. Hind 
wing usually with a dark shade along termen. 

Alar expanse, 19-24 mm. 


Male genitalia with a large, strongly sclerotized 
subanal plate, constricted before and beyond its middle. 
Harpe with apex notched below costa; clasper short, 
curved, situated near middle of harpe, and armed with 
several setae at its knobbed apex. Aedeagus with a 
cluster of several long, curved spines from apex; cor- 
nutus long, straight, stout. 


Female genitalia with bursa copulatrix finely scobi- 
nate but without signum; ductus bursae flattened, 
broad, twisted and constricted near genital opening, 
sclerotized throughout, the sclerotization involving 
bursa adjacent to ductus bursae and ductus seminalis; 
sclerotized band behind genital opening armed with 
four long, stout, projecting spines; collar of eighth seg- 
ment invaginated at dorsal margin to form a sclerotized 
pocket (fig. 713a). 

TYPE Locaitius: St. Thomas, British West Indies 
(pellucens, in BM); Barbados (cistupennis, in USNM). 


Foop piants: Vigna unguiculata (cowpeas, black- 
eyed peas, and garden peas), Bauhinia variagata, Can- 
avalia ensiformis (swordbeans), Canavalia maritima 
(black bean), Cajan cajan (pigeon pea), Phaseolus luna- 
tus (cultivated and wild limabeans), Phaseolus sp. 
(Brazilian specimens), Cassia occidentalia (one reared 
specimen from McCubbins Mills, Puerto Rico, before 
me; most records from this last plant are doubtful and 
probably the result of a misidentification of Fundella 
argentina as cistipennis). 

According to Scott the favored host in Puerto Rico is 
the cowpea (Vigna unguiculata), and the species, while 
frequent in limabeans, seldom does serious damage. 
Potentially it is an insect of economic importance. The 
larvae are primarily pod borers, but also bore into the 
stems and feed on the flowers of their hosts. They 
attack, as far as known, only Leguminosae. 


DistrisuTion: Unirep States: Florida, Hobe Sound 
(May), Miami, (Apr., May), Jupiter (Apr.), Coconut 
Grove, Marco Isl., Tampa (Mar.); Walton, Jensen 
(U. S. Dep. Agr. rearings from limabeans, Feb. 
1944), Riviera Beach, Vero Beach (J. R. Malloch, Dec. 
1941), Barsapos. Hartz: Damien (Dec., Feb.), 
Port-au-Prince. MontTSERRAT (Jan.). CuBA: Santiago, 
Matanzas. Vircin Isuanps: St. Croix (Mar., Oct., 
Nov.). Purrto Rico: San Juan, Rio Piedras (Mar.— 
May), Isabella, Catano (July), Vieques Isl. (Apr.). 
Braziu: Bafa (May), Cearé. Botrvra. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 61 


120. Fundella argentina Dyar 
Figures 234, 711 


Fundella pellucens Zeller (in part, ‘‘var. b’’), Isis von Oken, vol. 
41, p. 867, 1848; Horae Soc. Ent. Rossicae, vol. 16, p. 237, 
fig. 41b, 1881. 

Fundella argentina Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 1919.— 
Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. 109, 1945. 

Fundella eucasis Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 1919. 

Male antennal shaft with even smaller black basal 
scale tuft than that of pellucens. Forewing gray with- 
out the reddish brown, interspersed scaling character- 
istic of typical examples of pellucens; entire basal area 
to antemedian line dark fuscous gray (with but very 
slight dusting of whitish scales toward base in some 
specimens); this dark basal patch contrasted against 
the paler gray color of the remainder of the wing, ex- 
tending from costa to inner margin and bordered out- 
wardly by a narrow whitish line. Otherwise not dis- 
tinguishable, superficially, from pellucens. 

Female essentially like the male in color and markings 
except that the basal area of forewing is concolorous 
with or contrastingly paler than the remainder of the 
wing. A narrow dark line or a diffused dark shading 
outwardly bordering the obscure antemedian line. 

Alar expanse, 15-23 mm. 

Male genitalia without sclerotized subanal plate. 
Terminal projection of gnathos varying from round to 
pointed (fig. 234b) at apex. Harpe tapering to bluntly 
pointed apex; clasper a single, straight, slightly rough- 
ened, appressed spine, situated beyond middle of harpe. 
Aedeagus simple; cornutus a single, straight spine. 

Female genitalia with signum well developed and 
consisting of a large pear-shaped cluster of thornlike 
spines; sclerotized band behind genital opening, divided 
in the middle, simple (fig. 711a) in Argentinian and 
Brazilian specimens, or armed with a pair of median, 
spinelike projections (fig. 711), rather long in West In- 
dian specimens or short and disappearing in Mexican 
and Venezuelan specimens. 

TYPE Locaitigs: Tucumén, Argentina (argentina, in 
USNM); Caracas, Venezuela (eucasis, in USNM). 

Foop puant: Cassia spp. (reared examples in Na- 
tional Collection from Cassia bicapsularis and C. corym- 
bosa), Poinciana gilliesi. 

Distrisution: Unirep Srares: Florida, Biscayne 
Bay (May), Coconut Grove (Apr.), Stock Island (Apr.); 
Texas, Brownsville (Nov.). Mxico: Several examples 
reared from pods and blossoms of Cassia bicapsularis 
at Brownsville, Tex., quarantine station. CuBa: Bara- 
gua (Mar.), Habana, Matanzas, Santiago Province. 
Puerto Rico: Bayamén (Mar., Sept.), Vieques Isl. 
(Apr., July), Coamo Springs (Apr.), Aguirre Central 
(Aug.), San German (Aug.), San Juan (Noy.). Hartt: 
Pétionville (June). Virein Isuanps: St. Croix (Oct.- 
Nov.). Jamatca. VENEZUELA: El Valle (June). 
Braziu: Bafa (May). Argentina: Tucumén (Mar.). 

In collections this species has appeared most fre- 
quently under the name pellucens. Both argentina and 
pellucens have about the same distribution and are 
abundant in the West Indies, though, from material 


at hand, pellucens seems to be rarer on the mainland. 
Throughout its range argentina shows considerable var- 
iation in female genitalia. West Indian specimens have 
rather conspicuous spinelike extensions of the sclero- 
tized band behind the genital opening. These are en- 
tirely lacking in Brazilian specimens, and if one had 
only these extremes he would be justified in assuming 
that they were at least racially distinct. However, 
Venezuelan and Mexican examples show an intermedi- 
ate form with very short projections, and Central Amer- 
ican specimens, when recovered in sufficient numbers, 
will probably show all intergradations. The male geni- 
talia are remarkably uniform throughout the range of 
the species, exhibiting only minor individual variations 
in the shape of the terminal projection of the gnathos. 


121. Fundella agapella Schaus 
Figure 710 


Fundella agapella Schaus, Zoologica, vol. 5, No. 2, p. 47, 1923.— 
Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. 111, 1945. 


Female palpi, head, thorax, and forewing whitish 
gray; dark markings drab gray; transverse antemedian 
line of forewing white, defined chiefly by its narrow, 
dark outer border, sharply sinuate, indented a trifle 
just below costa, more deeply at top of cell and still 
more deeply at fold below cell; discal dot at end of cell 
obscure; white subterminal line indented at vein 6 and 
at submedian fold, bordered inwardly by a distinct dark 
shade .as broad as the white line itself and outwardly 
by a similar, fainter, dark shading, the latter conspic- 
uous only at apex. Hind wing as in the other species 
of Fundelia. Alar expanse, 12 mm; 

Genitalia like those of intermediate examples of ar- 
gentina except that the signum is considerably smaller 
in proportion to the size of the bursa. 

Typr Locarity: Tagus Cove, Albemarle, Galipagos 
Islands (type in USNM). 

Foop rrant: Unknown. 

Known only from the female type. Superficially a 
distinct species. The female genitalia, however, would 
indicate that agapella is only a race of argentina. A 
male will be needed for exact placement, and until it is 
available we shall have to treat agapella as a species. 


122. Fundella ignobilis Heinrich 
Ficures 232, 712 
Fundella ignobilis Heinrich, Proc. U. 8. Nat. Mus., vol. 96, p. 
112, 1945. 

Male antennal shaft without any trace of black basal 
scale tuft. Otherwise partaking of the pattern mark- 
ings of both pellucens and argentina; in some specimens 
dark basal patch of forewing round and reaching neither 
costa, nor inner margin (as in typical pellucens), in ma- 
jority of specimens, however, basal patch occupying 
whole basal area (as in typical argentina); median and 
outer areas of wing averaging a trifle paler than in 
argentina and without the reddish brown scaling of 
pellucens. 


62 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Female superficially similar to argentina except a 
trifle paler, on the average. 

Alar expanse, 13-22 mm, 

Male genitalia with gnathos terminating in a short, 
stout hook. Harpe with apex truncate; clasper mod- 
erately long, curved, and weakly haired at apex. Ae- 
deagus with a single, long, strong, curved spine from 
below apex; cornutus a short, stout, curved thorn. 

Female genitalia without spines adjacent to genital 
opening. Bursa copulatrix with signa consisting of a 
pair of partially fused bands, each armed with a row 
of short, stout, thornlike spines; ductus bursae short 
and broad, with median area unsclerotized; eighth seg- 
ment collar completely sclerotized except for a small, 
round, transparent spot on midventer, sclerotization 
extending to and over area behind genital opening. 

TypE LOcALity: Oaxaca, México (type in USNM). 

Foop prant: Unknown. 


Distrisution: México: Cérdoba (May), Guadala- 
jara, Jalapa, Oaxaca, Orizaba, Tehuacén (May, June, 
July). Guatemata: Cayuga. Costa Rica. Cusa: 
Santiago (June), Sierra Miestra (May). Purrto Rico: 
Aguirre Central (Aug.). Harri: Pétionville (June). 


123. Fundella ahemora Dyar 
Figures 233, 714 


Fundella ahemora Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 403, 
1914.—Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. 113, 1945. 
Antenna of male with small black scale tuft at base 
of shaft. Forewing with no or a very faint dark basal 
patch (when present covering basal area to antemedian 
line); antemedian line whitish, very faint; subterminal 
line white, faint but less obscure than antemedian, 
without dark borders except for an inner and an outer 
dark spot at inner margin of wing; veins from cell rather 
strongly outlined by dark scaling (the most conspicuous 
superficial character of the species). A thick, dark 
(brownish) hair tuft covering outer surface of male 
foretibia, a character not found in other species of the 
genus. Alar expanse, 18-23 mm. 


Male genitalia with gnathos terminating in a broad 
tongue-like plate. Harpe somewhat tapering but with 
apex truncate; a strong tuft of long scales from costa; 
clasper long, curved, slender, with a few hairs at apex. 
Aedeagus with a pair of long, curved, flattened spines 
from apex; cornutus a long, straight, slender spine. 

Female genitalia with a pair of long, widely spaced, 
basally curved spines from sclerotized area immediately 
behind genital opening. Bursa copulatrix with signa 
consisting of two rather short bands, each armed with 
a row of long spines. Ductus bursae bulged in the 
middle and with a strongly sclerotized median collar. 
Collar of eighth segment partially sclerotized and fused 
ventrally. 

TYPE LocaLity.—Orizaba, México (type in USNM). 

Foop pLant.— Unknown. 


DistRIBUTION: México: Orizaba, Jalapa, Teapa 
(Dec.), Cérdoba (Apr., Dec.), Cuernavaca (July). 


GUATEMALA: Quirigué (Mar.), Cayuga (Jan., May), 
Parulhé (July). Costa Rica: Juan Vifas (Nov.). 

Superficially the most easily distinguished species in 
the genus. The large foretibial tuft at once identifies 
the male, and both sexes can be separated by the rather 
conspicuous dark outlining of the veins. The veins are 
similarly dark scaled in the other species, but the con- 
trast of the dark veins against the pale intervenular area 
is more marked in ahemora. 


Genus 29: Difundella 


[Venational division B. Forewing with veins 4-5 connate or 
closely approximate at base. Hind wing with vein 3 from stalk 
of 4-5 or closely approximate to it for a short distance; on male, 
anal area folded into a pocket. Male genitalia without transtilla; 
sacculus of harpe strongly sclerotized and produced (free or 
partially free); penis without cornutus or other armature. 
Abdomen of male with lateral pockets and hair tufts between 
segments 2 and 3; eighth abdominal sternite developed as a 
narrow pocket; no hair tufts.] 


This genus shows affinities to Fundella but in many 
characters resembles more closely Rampylla and Cop- 
tarthria. In general habitus (wing pattern, color, and 
maculation) the moths of Difundella, Coptarthria, and 
Rampylla are strikingly similar, but the three genera 
are different on structural characters. Rampylla differs 
from the other two in the free length of vein 3 of 
male hind wing; Coptarthria in its notched male antenna; 
and Difundelia in its anellus (a simple plate without the 
long free spine of Rampylla and Coptarthria) and in the 
possession of scaled pockets between the second and 
third segments of the male abdomen. 

The species of Difundella differ considerably from 
each other on structural details, falling into two distinct 
groups which divide as follows: 

—Labial palpus reaching above vertex in both sexes. Hind 
wing with cell less than one-third the length of wing; 
vein 1b of male bent before middle and with a tuft of 
yellow hairlike scales on its under side (within the bend). 
Gnathos greatly reduced, its apical projection fine, 
needlelike. Costa of harpe without projections. 

—Labial palpus not reaching vertex on males, barely reaching 
vertex on females. Hind wing with cell more than 
one-third (but less than half) the length of wing; vein 
1b of male not bent; rough sex-scaling bordering 1c on 
under side of wing beyond base. Gnathos with apical 
process enlarged and strongly sclerotized. Costa of 
harpe with strongly sclerotized projection or projections. 

The second group probably deserves a separate generic 
designation; but the material before me representing its 
two species is too scanty and not in good enough con- 
dition, and the association of the females with their 
proper males too uncertain, to permit proper evalua- 
tion of generic characters for separation at this time. 


29. Genus Difundella Dyar 


Difundella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 327, 1914, 
(Type of genus: Difundella corynophora Dyar.) 
Tongue well developed. Antenna of male weakly 
pubescent. Labial palpus ascending, recurved, slender, 
smooth scaled; third segment acuminate. Maxillary 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 63 


palpus with second segment slightly thickened with 
scales. Forewing smooth; 11 veins; vein 2 from before 
lower outer angle of cell; 3 from the angle, nearly 
equidistant from 2. and 4; 4 and 5 connate or closely 
approximate at base and approximate for a short dis- 
tance beyond base; 6 from below upper angle of cell, 
slightly bent towards base; 10 from the cell, more or less 
approximate to the stalk of 8-9. Hind wing with vein 
2 from close to lower outer angle of cell; 3 from the 
stalk of 4-5 or closely approximate with it for a short 
distance from angle of cell; 4 and 5 long stalked (for over 
one-half their lengths); 7 and 8 closely approximate 
beyond cell, or shortly and weakly anastomosed; cell 
short, less than one-half the wing length; discocellular 
vein curved, partially obsolescent; on male, anal angle 
folded under to form a pocket enclosing a long hair- 
pencil. Eighth abdominal segment of male with sternite 
developed as a narrow, sclerotized pocket; on each side 
between abdominal segments 2 and 3 a shallow pocket 
containing a modified scale tuft. 

Male genitalia with uncus stout, but variously shaped. 
Transtilla absent. Harpe with sacculus very strongly 
sclerotized, free or partially free and curved. Aedeagus 
with strongly sclerotized and pointed apex; penis 
unarmed. Vinculum stout. A long hair tuft from 
intersegmental area adjacent to base of sacculus of 
harpe. 

Female genitalia with bursa copulatrix membranous; 
signum, if present, a small patch of weak scobinations; 
ductus bursae membranous (unsclerotized) except about 
genital opening; ductus seminalis from bursa near 
junction of bursa and ductus bursae. 

The characteristic pattern features of the species of 
Difundella (as of Coptarthria and most species of 
Rampylia) are: The strongly contrasted, fine, blackish, 
transverse lines forming the outer border of the ante- 
medial and the inner border of subterminal lines of 
forewing; the almost straight, oblique or vertical ante- 
medial line, set well out from base of wing; the oval, 
pale discal spot covering the discocellular vein; and the 
black streaks on veins 2 to 6 just beyond it. 


Genus Difundella, Species 124 and 125: D. cory- 
nophora and D. subsutella 


[Labial palpus reaching above vertex in both sexes. Hind wing 
with cell less than one-third the length of wing; vein 1b of male 
bent before middle and with a tuft of yellow hairlike scales on its 
under side (within the bend). Gnathose greatly reduced, its 
apical projection fine, needlelike. Costa of harpe without 
projections.] 
124. Difundella corynophora Dyar 
Fiaures 7, 235, 715 
Difundella corynophora Dyar, Proc. U. 8. Nat. Mus., vol. 47’ 
p. 327, 1914. 


Forewing with basal area (to antemedial line), and 
upper median area beyond antemedial line and from 
lower margin of cell to costa, blackish fuscous; some 


extension of this dark shade extends narrowly to inner 
margin along the outer dark border of the antemedial 
line; ground color of remainder of wing a ruddy ocherous; 
some extension of this ocherous shade invades the dark 
basal area along the lower fold and forms the centers of 
the transverse lines; beyond the cell the ocherous shade 
is broken by black streaks on veins 2 to 6 and beyond 
subterminal line it is more or less clouded by blackish 
fuscous; discal spot ruddy ocherous, covering disco- 
cellular vein; antemedial line oblique; its narrow dark 
borders black shading to reddish; the dark borders of 
subterminal line also more reddish brown than black; 
all the dark transverse lines less contrasted and con- 
spicuous than those of other species in the genus. Hind 
wing dark smoky fuscous; the veins and terminal edge 
black and (on the female) a blackish shade at apex. 
Alar expanse, 15-16 mm. 

Male genitalia with uncus produced and slightly 
bifurcate at apex. Free projection of sacculus curved 
back towards lower margin of harpe. Aedeagus forked; 
the longer element of the fork spined at apex. Female 
genitalia with signum a small round patch; genital 
opening surrounded by an oblong, strongly sclerotized 
plate; in intersegmental area behind this plate a pair of 
ventrolateral pockets (fig. 715b). 

Typr tocauiry: La Chorrera, Panama (type in 
USNM). 

Foop piant: Unknown. 

DistrisuTion: GuATEMALA: Cayuga (Aug.), Chejel 
(June). Panam&: La Chorrera. Franco GUurIANA: 
Cayenne. 

Easily identified by its genitalia. 


125. Difundella subsutella (Schaus), new combination 
FicursE 236 


Ulophora subsutella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p. 248, 1913. 

Rampylla subsutella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7, 
p. 84, 1919. 

The only representative of this species is the male 
type which is somewhat rubbed and the markings con- 
sequently obscured. It differs from corynophora in 
having the pale ground color of the outer areas of fore- 
wing more reddish than ocherous; the black outer 
margin of antemedial line more sharply defined, black 
throughout, outwardly oblique to lower fold and thence 
slightly curved inward to lower margin. Hind wing 
semihyaline white with a faint brownish ocherous tint; 
veins not appreciably darkened; a fine brown line along 
termen. Alar expanse, 17.5 mm. 

Male genitalia with uncus triangulate. Gnathos 
reduced even more than that of corynophora, the lateral 
arms represented by mere stubs. Projecting part of 
sacculus curving away from harpe. Aedeagus sickle 
shaped; its apical half sharply curved, very strongly 
sclerotized, and tapering to a sharp point. 

Typr LOCALITY: Juan Vifias, Costa Rica (Jan.; type 
in USNM). 

Foop puant: Unknown. 


64 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Genus Difundella, Species 126 and 127: D. dis- 
tractor and D. tolerata 
[Labial palpus not reaching vertex on males, barely reaching 
vertex on females. Hind wing with cell more than one-third 
(but less than half) the length of wing; vein 1b of male not bent; 
rough sex-scaling bordering Ic on underside of wing beyond base. 
Gnathos with apical process enlarged and strongly sclerotized. 
Costa of harpe with strongly sclerotized projection or projections.] 


126. Difundella distractor, new species 
Figures 237, 716 


Dark areas of forewing dark brownish gray; the pale 
outer areas dull whitish; antemedial line well out 
towards middle of wing, nearly vertical, slightly notched 
at vein 1b, red-brown with a fine black line along its 
outer edge and preceded by a rather broad whitish 
blotch, extending from just below costa, nearly to inner 
margin; a short black streak along lower fold for a short 
distance from base of wing; the discal spot whitish; 
blackish lining on the veins beyond cell very weak; 
subterminal line very slightly bent between veins 4 
and 6 otherwise nearly vertical, outwardly bordered by 
a narrow shade of the dark ground color and inwardly 
by a fine black line, Hind wing semihyaline white; a 
blackish brown line along termen; the veins not dark- 
ened; the male wing on the undersurface rather loosely 
and coarsely covered with yellowish scales, especially 
along the veins. Alar expanse, 14.5-15.5 mm. 

Male genitalia with uncus semispoon-shaped. Apical 
projection of gnathos large, triangulate, strongly sclero- 
tized. Harpe with subtriangulate cucullus; a single, 
long, stout, flat, curved, tapering projection from mid- 
costa. Aedeagus long, slender, smooth, tapering to a 
point. Vinculum’ about twice as long as greatest 
width. Female genitalia without signum; genital 
plate large, medially notched and supported by strongly 
sclerotized, involuted seventh abdominal segment. 

TyprE Locatity: Palmas Abajas, Puerto Rico (type 
in Cornell Univ., paratype in USNM, 61328). 

Foop PLant: Unknown. 

Described from male type labeled ‘‘Palmas Abajas 
(near Guayama), P. R., 1900 ft., [date unreadable], 
W. A. Hoffman’; and one female paratype from Aguirre 
Central, Puerto Rico, ‘‘Apr. 2-3, 31,’’ M. D. Leonard, 
collector. The female is in good condition and served 
for the foregoing color and pattern description. The 
male is badly rubbed and has the palpi and antennae 
broken off; but enough of the markings remain to show 
that they were the same as those of the female. 


127. Difundella tolerata, new species 
Fieures 238, 717 


Similar to the foregoing species (distractor) except 
that pale areas of forewing’ are much more restricted, 
limited to an irregular area bordering inner margin of 
subterminal line ‘(extending back to cell above and 
nearly to antemedial line below); some diffused pale 
shading along the lower fold in basal area; the pale 
areas very dull ocherous white, not as well contrasted 
as in distractor; discal spot ocherous. Hind wing trans- 


lucent, white with a very faint ocherous tint on male; 
a faint dark line along termen. Alar expanse, 19 mm. 

Male genitalia with uncus strongly sclerotized and 
stout (longer than basal width), broadened and rounded 
towards apex. Apical projection of gnathos a long, 
broad, strongly sclerotized, obliquely bent band. 
Harpe with narrow, spatulate cucullus; two projections 
from costa, the first a double thornlike projection from 
near middle, the second a rather slender spine from 
outer third. Aedeagus with a couple of short spines 
at apex. Vinculum no longer than greatest width. 

Female genitalia with signum a narrow, elongate 
patch of scobinations; genital opening simple; ductus 
bursae about three times as long as bursa. From 
the dorsocaudal margin of seventh segment an in- 
vaginated, sclerotized shield supports at each lateral 
margin a short, blunt, weakly pigmented hornlike 
process. 

Typr Locatity: Kast Bolivia (type in BM). 

Foop puant: Unknown. 

Described from two specimens from the British Mu- 
seum Collection labeled “Ost Bolivia, Aug.—Oct., 1920, 
T. Steinbach,’’ the male type and a female matching the 
male in size, color, and markings. J do not make’ this 
female a paratype (although the foregoing description 
of the female genitalia is made from it) because there is 
a slight doubt that it is the true female of the species. 
Its genitalia are similar in all but the most trifling details 
to those of a series of females from Cayuga, Guatemala, 
associated in our collection with males of Coptarthria 
dasypyga. One or the other of our identifications of 
females (or both of them) may be in error. We shall 
not know until more material is available. 


Genera 30-33: Coptarthria to Dasypyga 


[Venational division B. Male genitalia with transtilla a sinuate, 
sclerotized, more or less scobinate band involved with gnathos; 
a long, free spine associated with anellus. Highth abdominal 
segment of male with sternite developed as a narrow sclerotized 
pocket.] 4 


30. Genus Coptarthria Ragonot 


Coptarthria Ragonot, Monograph, pt. 1, p. 251, 18938. (Type of 


genus: Myelois dasypyga Zeller.) 

Tongue well developed. Antenna‘of male with basal 
segment of shaft considerably elongated, flattened and 
deeply notched at outer extremity; shaft pubescent. 
Labial palpus upturned, slender, barely reaching vertex; 
third segment about half the length of second, bluntly 
pointed. Maxillary palpus filiform. Forewing smooth; 
11 veins; vein 2 from before lower outer angle of cell; 3 
from the angle, closer to 4 at base than to 2; 4 and 5 
short stalked or connate and closely approximate for a 
short distance from base; 6 from below upper angle of 
cell, bent towards base; 10 from the cell; male without 
costal fold. Hind wing with vein 2 from close to lower 
outer angle of cell; 3 from middle of stalk of 4-5, or 
closely approximate to it; 4. and 5 stalked for slightly 
more than half their lengths; 7 and 8 closely approxi- 
mate beyond cell; cell short, about one-third the length 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 65 


of wing; discocellular vein curved. Highth abdominal 
segment of male with sternite developed as a narrow, 
sclerotized pocket. 

Male genitalia with uncus moderately stout; deeply 
concaved apically (probably only a specific character). 
Transtilla a sinuate, sclerotized band involved with and 
fusing into gnathos. Gnathos proper identifiable only 
by its rather weak lateral arms. Harpe with apex of 
cucullus slightly hooked; sacculus simple, not produced. 
Anellus a small plate with greatly reduced lateral lobes, 
the latter indicated chiefly by their short hair tufts; 
dependent from near base of anellus plate and associated 
with it a long, slender U-shaped band supporting from 
the bottom of the U a long, strongly sclerotized, free 
spine, the latter lying dorsad of the aedeagus. Aedeagus 
small, simple; penis with a few weak scobinations, other- 
wise unarmed. 

The genus is distinguished from its nearest allies by 
the notched shaft of its male antenna. Female char- 
acters could not be included in the foregoing descrip- 
tion, as examples of this sex have not been satisfactorily 
associated with males of the type species, the only 
known representative of the genus. 


128. Coptarthria dasypyga (Zeller) 
Ficures 10, 239, (?) 718 


Myelois dasypyga Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 215, 
1881 


Coptarthria dasypyga (Zeller) Ragonot, Monograph, pt. 1, p. 251, 
1893. 


I have seen no Colombian examples, but there are six 
males in the National Museum from Guatemala which 
Dyar identified as dasypyga. There is no reason to 
question his identification; for the specimens have the 
typical Coptarthria antenna, are the right size, and agree 
in color and maculation with Ragonot’s description of 
the Zeller type. 


Forewing gray-brown with a very slight intermixture 
of ocherous shading above inner margin; the basal area 
a trifle paler than remainder of wing; branches of median 
vein (veins 2 to 5) faintly streaked with brown or black- 
ish brown; transverse lines pale with dark borders and 
faintly tinged with reddish scaling at middle and near 
costa; antemedial line well out near middle of wing, 
nearly vertical, straight except for a slight inward angu- 
lation at vein 1b, its inner border a weak brown line, its 
outer bordering line black; subterminal line well back 
from termen, vertical to vein 4, thence oblique to inner 
margin, its inner border a black line; a rather large, oval, 
ochraceous spot on discocellular vein, margined by faint 
black scaling; along termen a row of conspicuous, more 
or less confluent black spots. Hindwing semitranslu- 
cent white; the veins faintly darkened (pale, ocherous 
brown) and a faint dark line along termen. Alar ex- 
panse, 14-17 mm. 

Male genitalia as given for the genus. 

TYPE Locatity: Honda, Colombia (type, o, in BM). 

Foop piant: Unknown. 

300329—56——6 


Distrisution: Coomera: Honda, GuaTeMata: Cay- 

= om , May), Quirigué (Mar.), Volcén Santa Maria 
y). 

Associated with the males in the National Collection 
are five females from Cayuga (Apr., May) identical with 
the males in all superficial characters. Their genitalia 
are like those of the Bolivian female I have associated 
tentatively with the type of Difundella tolerata (fig. 717). 
However, there is also a female with the same color and 
markings from Juan Vifas, Costa Rica (Feb.), which 
has different genitalia (fig. 718), similar to those of 
Anadelosemia. From the limited material available 
and the few and scattered distributional records it is im- 
possible to determine which females go with which males, 

31. Genus Promylea Ragonot 


Promylea Ragonot, N. Amer. Phycitidae, p. 5, 1887; Monograph, 
pt. 1, p. 207, 1893.—Hulst, Phycitidae of N. Amer., p. 139, 
1890. (Type of genus: Promylea lunigerella Ragonot). 


Tongue well developed. Antenna pubescent; shaft 
of male without notch or other modifications. Labial 
palpus upturned, slender, reaching vertex; third seg- 
ment about half the length of second, acuminate. Max- 
illary palpus squamous. Forewing smooth; 11 veins; 
vein 2 from before, but rather near lower outer angle of 
cell; 3 from the angle, closer to 4 than to 2 at base; 4 
and 5 closely approximate for a short distance from 
base; 6 from below upper angle of cell, straight; 8 and 9 
long stalked, the free element of 9 short; 10 from the 
stalk of 8-9, or from the cell, connate with or closely 
approximate with it for a short distance beyond base 
(definitely stalked with 8-9 in most of the specimens of 
lunigerella); male without costal fold. Hind wing with 
vein 2 from before lower outer angle of cell; 3 from the 
angle, connate with 4; 4 and 5 closely approximate or 
anastomosed for half their lengths beyond cell; 7 and 8 
approximate or partially anastomosed for less than half 
their lengths beyond cell; cell nearly half the length of 
wing; discocellular vein curved. Eighth abdominal seg- 
ment of male with sternite developed as a narrow sclero- 
tized pocket. 

Male genitalia asin Coptarthria except: Apical margin 
of uncus rounded; lateral arms of gnathos more strongly 
developed; penis sometimes with a weak cornutus. 


Female genitalia with bursa and ductus bursae mem- 
branous, bursa small, considerably shorter than ductus; 
signum present but weak, a patch of scobinations or a 
small plate supporting a very small thorn; genital open- 
ing simple; ductus seminalis from ductus bursae. Col- 
lar of eighth abdominal segment with a broad, flaring, 
sclerotized apron projecting from center of anterior 
dorsal margin; in the intersegmental area between collar 
and seventh segment a sclerotized and coarsely granu- 
late pocket (fig. 721a). 

The genus is close to both Coptarthria and Anadelo- 
semia but distinct, differing from the former in its simple 
male antenna and from the latter and all the genera of 
this immediate group having the sinuate, involved 
transtilla and the free spine associated with anellus by 
its peculiarly developed, female, eighth-segment collar. 


66 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


In habitus the moths of Promylea differ strikingly from 
those of Coptarthria; the transverse lines of forewing 


being more widely separated and the antemedial line 
decidedly oblique. 


129. Promylea lunigerella Ragonot 
Figures 9, 240, 721 


Promylea lunigerella Ragonot, N. Amer. Phycitidae, p. 5, 1887; 
Monograph, pt. 1, p. 20, 1893.—Hulst, Phycitidae of N. 
Amer., p. 139, 1890.—McDunnough, Check list, No. 6146, 
1939. 

Forewing gray or brownish gray, pale in median and 
most of basal areas and (in many specimens from Van- 
couver and Washington State) with a faint rosy tint; 
antemedial line indicated chiefly by its blackish outer 
border, strongest towards costa and oblique from costa 
before one-third to inner margin, preceded by reddish 
brown or ocherous brown patch, broad and inwardly 
dark-margined on inner margin, attenuated and paling 
out towards costa; subterminal line pale gray, bordered 
inwardly by a blackish brown line and outwardly by a 
much fainter dark line (grayish or reddish brown), out- 
wardly curved between veins 8 and 1b; discal dots rarely 
separated, normally fused into a thin blackish lunule 
on the discocellular vein; terminal dots obscure, when 
distinguishable more or less fused into a line along ter- 
minal margin. Hind wings pale smoky fuscous; the veins 
little if any darkened; a very faint brownish line along 
termen. Alar expanse, 20-24 mm. 

TYPE Locatiry: Vancouver Isl., British Columbia 
(type in Paris Mus.). 

Foop puant: Unknown. : 

Distripution: Canapa; British Columbia, Fitzgerald 
(June), Duncans (Vancouver Isl.), Victoria (June, July, 
Aug.), Unirep Statss: Washington, Bellingham (Aug.), 
Friday Harbor (July, Aug.), Mt. Constitution (July); 
California, Glen Alpine (Lake Tahoe, July). 


130. Promylea lunigerella glendella (Dyar) 
Figures 241, 720 


Myelois glendella Dyar, Journ. New York Ent. Soc., vol. 14, p. 30, 
1906. 


Promylea glendella (Dyar) Barnes and McDunnough, Check list 
of the Lepidoptera of Boreal America, No. 5584, 1917.— 
McDunnough, Check list, No. 6147, 1939. 

There is nothing to distinguish this from many of our 
specimens of lunigerella from Washington and British 
Columbia except some slight differences in genitalia 
of very doubtful significance, and the name should prob- 
ably go into synonymy; but until material is available 
from intervening areas and something is known of the 
life history, glendella may be retained as a possible local 
race. It certainly is nothing more than that. 

TypPE tocaLity: Glenwood Springs, Colo. (type in 
USNM). 

Foop prant: Unknown. 

Represented in the National Museum by two males 
and three females from the type locality (Aug., Sept., 
Oct.). In addition to the foregoing there are before me 
five examples of another variety of lunigerella from Fal- 


len Leaf Lake, Calif., Aug. 8 and 12, 1932, H. H. Keifer, 
collector. They are quite different in color, having 
paler gray forewings with much fainter transverse dark 
markings and more whitish hind wings than our other 
examples of lunigerella. They are probably nothing 
but a color form and for obvious reasons are going with- 
out a name at present. 


131. Promylea dyari, new name 
FIcuRE 243 


Dioryctria zimmermani Druce (not Grote), Biologia Centrali 
Americana, Lepidoptera Heterocera, vol. 2, p. 564, 1899. 
Dioryctria drucet Dyar, Ins. Inse. Menstr., vol. 7, p. 44, 1919 

(preoccupied). 

Similar to lunigerella except: Larger and darker; the 
paler areas of forewing a dark ashy gray; outer black 
border of antemedial line broader, forming a small tri- 
angle on costa; inner border of subterminal line also 
slightly stronger, black; the subterminal line itself is 
more sharply angled below costa and from about vein 6 
proceeds to inner angle in a slanting almost straight line 
(a difference that strikes the eye but is hardly signifi- 
cant; for on some examples of Junigerella the subterminal 
line is similarly shaped); terminal dots confluent, form- 
ing a conspicuous black line along termen. Alar ex- 
panse, 30 mm. 

Male genitalia with tezumen and vinculum somewhat 
stouter (broader in proportion to their width) than those 
of lunigerella; penis armed with a weak cornutus. 


TYPE Locatity: Rinconada, Vera Cruz, México (type 
in USNM). 

Foop prant: Unknown. sods 

The species is known only from the male type which 
both Druce and Dyar mistook for a female. ‘The refer- 
ence to Dioryctria is difficult to understand in Dyar’s 
case; for vein 3 of hind wing is appreciably too short for 
that genus. The transfer of “Dioryctria drucer Dyar” 
and “‘Nephopteryx druceit Ragonot”’ to the genus Promy- 
lea makes the former a secondary homynym and necessi- 
tates the newname. It is possible that the two “drucei” 
represent only different sexes of one species; but this 
cannot be determined without more material of each, so 
for the present they must be treated as separate species. 


132. Promylea druceii (Ragonot), new combination 
FIcuRE 722 


Nephopteryx drucezi Ragonot, Nouv. Gen., p. 15, 1888; Mono- 
graph, pt. 1, p. 801, 1893. 

Lhave not seen any specimens matching Ragonot’s de- 
scription or figure (Monograph, pl. 9, fig. 17) ; but Ihave 
before me an excellent photograph of the type supplied 
by Tams. This shows a moth similar to dyari but with 
the dark markings (especially the dark borders of the 
transverse lines) much more|expanded, the outer border 
of the antemedial line forming a large triangle on costa. 
According to Ragonot these dark areas have a decided 
purple tint, rather than the dull black or blackish brown 
of dyart. The spot on the inner margin before the ante- 
median line is also purplish black rather than orange (as 


AMERICAN’ MOTHS OF THE SUBFAMILY PHYCITINAE 67 


in dyari) and there is considerable purplish dusting on 
the paler areas of the wing. The moth is also smaller 
than Dyar’s type. Alar expanse, 24 mm. 

Typz Locauity: Totonicapin, Guatemala (type in 
BM). 

Foop prant: Unknown. 

Represented only by the type. 


133. Promylea mindosis Dyar 
Figures 242, 723 


Promylea mindosis Dyar, Ins. Insc. Menstr., vol. 10, p. 172, 1922. 


A narrow-winged, suffused, dark species. Ground 
color of forewing very dark gray-brown; the blackish 
transverse lines narrow and faint; antemedial line ob- 
solete except for its faint outer border; subterminal line 
distinguishable but faint; discal spots at end of cell 
weak but apparently separated. Hind wing very pale 
smoky fuscous, translucent. Alar expanse, 25-26 mm. 

Male genitalia with penis with weak cornutus. 
Female genitalia with siznum developed as a small 
plate bearing a minute, knoblike projection. 

Typr Locauity: Mexico City, Mexico (Aug.; type in 
USN). 

Foop pirant: Unknown. 

Known only from the type series of two males and 
one female from the type locality. 


134. Promylea dasystigma Dyar 
Fiaure 724 


Promylea dasystigma Dyar, Ins. Insc. Menstr., vol. 10, p. 172, 
1922. 


Similar to mindosis, differing only in slight details: 
The forewing is a trifle darker, more blackish than 
brownish gray; the subterminal line ends on inner 
margin in a small but distinct white spot; discal spots 
fused into a thin, blackish lunule on discocellular vein. 
The apron from the eighth-segment collar is differently 
shaped and larger and the intersegmental pocket be- 
tween seventh segment and collar proportionally wider 
than those of mindosis (compare figs. 723a and 724a). 
Alar expanse, 25-26 mm. 

Female genitalia with signum a very weak patch of 
scobinations. 

Typ Locatity: Mexico City, Mexico (Aug.; type in 
USNM). 

Foop puant: Unknown. 

Known only from the two females of the type series. 


32. Genus Anadelosemia Dyar 


Anadelosemia Dyar, Ins. Insc. Menstr., vol. 7, p. 51,1919. (Type 


of genus: Nephopteryx senesciella Schaus). 

Characters of Promylea except: Forewing with vein 
10 always from the cell and not closely approximate to 
the stalk of 8-9. Hind wing with vein 2 from close to 
lower outer angle of cell; 3 from the stalk of 4-5 or at 
least anastomosed with it for a short distance; 4 and 5 
stalked for over half their lengths beyond cell; cell about 
one-third the length of wing. Female abdomen with 


eighth-segment collar simple; no dorsal pocket between 
seventh and eighth segments. On male a hair-pencil 
from lower, outer side of metathorax near base of leg. 


Very close to Promylea but apparently distinct 
enough. The general habitus of the moths is similar 
except that the species of Anadelosemia (except for 
obstitella) are decidedly smaller. 


135. Anadelosemia senesciella (Schaus) 
Figures 11, 244 


Nephopteryx senesciella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 
11, p. 251, 1913. 


Anadelosemia senesciella (Schaus) Dyar, Ins. Insc. Menstr., vol. 
7, p. 52, 1919, 


Forewing ashy white, the basal area to antemedial 
line stained with pale brown; costal edge at base black; 
antemedial line white, broad and oblique from costa to 
lower fold, thence crescentiform to inner margin, on 
upper half bordered by a broad blackish spot, diffused 
on costa and continued below fold as a narrow line to 
inner margin, a weaker dark inner border on lower half; 
a dark (brownish) shade extending obliquely across the 
wing from costa just before subterminal line to near 
middle of inner margin, irregular and more or less 
diffused over remainder of outer area; discal spots dis- 
tinct, blackish (on this and most other species of the 
genus different on opposing forewings, separated or 
fused into a line); subterminal line sinuate-dentate, 
rather close to termen, bordered inwardly by a few 
blackish dots and outwardly by a black line which be- 
gins as a strong dash on costa; a row of blackish dots 
along termen. Hind wing semihyaline tinted with 
brown; the veins darkened and a narrow dark shade 
along termen. Alar expanse, 15 mm. 

Male genitalia distinguished chiefly by the shapes of 
uncus, harpe, and vinculum (fig. 244); penis armed with 
an elongate, narrow, flattened, bladelike cornutus. 

TYPE LOCALITY: Juan Vifias, Costa Rica (Jan.; type 
in USNM). 

Foop prant: Unknown. 

Known only from the male type. 


136. Anadelosemia tecmessella (Schaus) 
Ceracanthia tecmessella Schaus, Ann, Mag. Nat. Hist., ser. 8, 
vol. 11, p. 251, 1913. 
Anadelosemia tecmessella (Schaus) Dyar, Ins. Insc. Menstr., vol. 
7, p. 52, 1919. 

Forewing dull ashy white; an oblique blackish shade 
at base; antemedial line narrow, white, a broken narrow 
inner black border indicated; bordering the antemedian 
line on costal half, a rather large triangular bronzy 
brown spot; this color diffused outwardly along costal 
edge; the oblique dark shade across wing beyond cell 
extended to include the remainder of the outer area; 
subterminal line but slightly paler than the brownish 
color of outer area, otherwise as in senesciella; discal dots 
distinct, blackish brown, well separated. Hind wing 
pale smoky brown, veins and terminal margin darker. 
Alar expanse, 14 mm. 


68 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Typ LocaLity: Avangarez, Costa Rica (uly; type 
in USNM). 

Foop puant: Unknown. 

Known only from the female type. It lacks an 
abdomen so genitalia could not be studied. However, 
the coloration of forewing suggests a species distinct 
from anything else in the genus. 


137. Anadelosemia fifria Dyar 
FiauRE 726 


Anadelosemia fifria Dyar, Ins. Insc. Menstr., vol. 7, p. 52, 1919. 


Forewing similar to that of senesciella except: Whitish 
ground color more extended filling most of outer area; a 
dark brown shading at extreme base; dark shadings 
fuscous brown rather than blackish; antemedial white 
line narrow, its'dark borders also narrower; the post 
media, transverse dark shade also narrower; terminal 
dots more or less confluent. Alar expanse, 15 mm. 

Female genitalia with signum, developed as a narrow, 
small, shallow, granulate cup. 

TYPE LocaLity: Cayuga, Guatemala (type in USNM). 

Foop prant: Unknown. 

Represented only by the female type and one other 
female from the type locality (May). 


138. Anadelosemia base Dyar 
FIGURE 727 


Anadelosemia base Dyar, Ins. Insc. Menstr., vol. 7, p. 52, 1919. 


Similar to the foregoing species (jifria) except: Dark 
markings a paler brown; the outer dark border of costal 
half of antemedian line a small triangulate spot with 
some extension outward on costal edge; terminal dots 
rather weak but not confluent. Alar expanse, 15 mm. 

Female genitalia without signum; ductus bursae pro- 
portionally much longer than that of jifria; ductus 
seminalis from near middle of ductus bursae. (In the 
other species it branches off from the ductus bursae 
very near its Junction with the bursa copulatrix.) 

TYPE LocaLity: Cayuga, Guatemala (type in USNM). 

Foop prant: Unknown. 

Known only from the female type and one other 
female from the type locality. Superficially hardly dis- 
tinguishable from jifria. However, the differences in 
their female genitalia suggest two distinct species. 


139. Anadelosemia obstitella (Schaus), new combination 
FicurE 728 
Nephopteryzx obstitella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 
11, p. 251, 1913. 

The largest and most strikingly marked species in the 
genus; the antemedial and subterminal lines shining 
white; the inner, black bordering line of the former 
continuous from costa to inner margin and strongly con- 
trasted against the ashy gray ground color of the basal 
area of the forewing; outer black border on costal half 
of antemedial line and the black dashes preceding and 
following the subterminal line enlarged and well con- 
trasted. Hind wing semihyaline white with a very faint 


brownish tint; veins faintly darkened; a narrow, pale 
fuscous shading along termen, especially towards apex. 
Alar expanse, 22 mm. 

Female genitalia with signum present as a narrow, 
weak, elongate, irregular scobinate patch (see enlarge- 
ment). The shape and development of signum is a 
character of very doubtful value in this genus and 
probably subject to considerable individual variation. 

Type LocaLity: Mount Pods, Costa Rica (type in 
USNM). 

Foop prant: Unknown. 

Known only from the female type. 


140. Anadelosemia texanella (Hulst), new combination 
Figures 246, 729 


Myelois texanella Hulst, Canadian Ent., vol. 24, p. 60, 1892. 

Myelois dulciella Hulst, Canadian Ent., vol. 32, p. 176, 1900. 

Tacoma texanella (Hulst) Barnes and McDunnough, Contribu- 
tions, vol. 3, p. 193, 1916; vol. 4, p. 174, 1918—MceDun- 
nough, Check list, No. 6143, 1939. 

Forewing ashy white (due to a fine, sparse peppering 
of fuscous scales on the white ground color); costal edge 
at extreme base black; antemedial line not differenti- 
ated except by the narrow, curved, black line forming 
its outer border from costa to inner margin and by @ 
preceding brown spot on inner margin; subterminal line 
sinuate, narrow, bordered inwardly by a continuous 
black line and outwardly by a faint, narrow, brownish 
shade continued from a blackish dash on costa; discal 
spots black, separated. Hind wing smoky white, dark- 
ening outwardly; a fine brown line along termen. Alar 
expanse, 13-16 mm. 

Male genitalia figured from type of dulciella. They 
exhibit several specific characters: a long, slender, 
strongly sclerotized, spinelike, apical projection from 
enathos (the other species whose males are known show 
no such structure, the only elements attached to the 
lateral arms of gnathos at their junction being the 
transtilla and the base of the more or less sclerotized 
subanal plate); tezumen considerably elongated in pro- 
portion to the vinculum; harpe short and broad; penis 
finely spined at apex. Female genitalia without signum; 
ductus seminalis from ductus bursae near junction of 
bursa copulatrix. The distinctive female structural 
character is the shape of the eighth-segment collar (fig. 
729). 

TypPE LocaLitixs: Blanco County, Tex. (texanella, 
in AMNH, ex Rutgers); Hastings, Fla. (dulciella, in 
AMNG, ex Rutgers). 

Foop piant: Unknown. 

DistrisuTion: UnitEpD States: Texas, Blanco 
County, San Benito (Apr., Sept.); Florida, Hastings 
(Oct.). Purrto Rico: San Germén (Apr.); Cusa: Santa 
Clara, Central Soledad (‘“‘E. EH. A. Cuba, Ento. no. 
10234,” May). 

The Puerto Rican and Cuban examples are males 
which agree in every detail with the type of dulciella. 
The type of tezanella lacks an abdomen, so its genitalia 
could not be checked (the other Texas examples are 
females) ; but careful study of the two types discovered 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 69 


no difference that would justify any doubt of the syn- 
onymy proposed by Barnes and McDunnough (1918). 


141. Anadelosemia condigna, new species 
Fieures 245, 730 


Forewing similar to that of teranella except: Outer 
black border of antemedial line nearer middle of wing, 
nearly vertical and more denticulate; subterminal line 
with a discontinuous, blackish outer border; some red- 
dish spotting in the postmedial area near inner margin; 
costa at base not black, but a transverse black marking 
at extreme base; discal spots confluent, forming a black- 
ish or reddish brown line along discocellular vein. Hind 
wing semihyaline white with a narrow fuscous shade 
along termen. Alar expanse, 15-18 mm. 

Male genitalia similar to those of senesciella, but dif- 
fering in the shapes of uncus, harpe, and vinculum; penis 
armed with a narrow, flat, bladelike cornutus similar 
to that of senesciella. Female genitalia without signum, 
similar to that of texana except for the shape of the 
eighth-segment collar (fig. 730). 

TyprrE Locauity: Prescott, Ariz. (type in USNM, 
61329; paratypes in Cornell Univ. and BM). 

Foop prant: Unknown. 

Described from male type from the type locality 
(July) and seven male and four female paratypes from 
the Baboquivari Mts., Pima County, Ariz., collected by 
O. C. Poling, May 1-15, 1924. 


33. Genus Dasypyga Ragonot 


Dasypyga Ragonot, N. Amer. Phycitidae, p. 5, 1887; Monograph, 
pt. 1, p. 206, 1893.—Hulst, Phycitidae of N. Amer., p. 138, 
1890. 


Tongue well developed. Antenna weakly pubescent. 
Labial palpus upcurved, reaching to vertex (female) or 
nearly to it (male) ; third segment half as long as second, 
acuminate. Maxillary palpus small, squamous. Fore- 
wing with some raised scaling on basal area (probably 
only a specific character); 11 veins; vein 2 from before 
but near lower outer angle of cell; 3 from the angle, 
closer to 4-5 than to 2; 4 and 5 shortly stalked; 6 from 
below upper angle of cell, straight; 10 from the cell, 
closely approximate for some distance with the stalk of 
8-9; male without costal fold. Hind wing with vein 2 
from near lower outer angle of cell; 3 from the angle, 
connate with the stalk of 4-5; 4 and 5 long stalked (for 
about two-thirds of their lengths); 7 and 8 closely ap- 
proximate or contiguous for a short distance beyond 
cell; cell one-third the length of wing; on male with anal 
angle folded into a thickened pocket containing a hair 
tuft. Eighth abdominal segment of male with sternite 
developed as a narrow, sclerotized pocket. 

Male genitalia similar to those of Anadelosemia ex- 
cept: Sacculus of harpe strongly sclerotized and for half 
its length developed as a free arm extending across 
harpe; cornutus well developed; a pair of hair tufts from 
intersegmental area adjacent to outer surfaces of the 
sacculi at their bases. 

Female with signum a small patch of scobinations; 
ductus bursae considerably longer than bursa, tubular 


and strongly sclerotized for about one-fifth its length 
from genital opening, and for over half its remaining 
length sclerotized, slightly flattened and bent (sinuate) ; 
ductus seminalis from bursa near its junction with 
ductus bursae. 

A distinct genus distinguished from the other genera 
with similar transtillae and venation by its stout, greatly 
produced sacculus and sclerotized ductus bursae. Con- 
tains one North American species. 


142. Dasypyga alternosquamella Ragonot 
Ficures 12, 247, 719 


Dasypyga alternosquamella Ragonot, N. Amer. Phycitidae, p. 5, 
1887; Monograph, pt. 1, p. 206, 1893.—Hulst, Phycitidae 
of N. Amer., p. 138, 1890.—Heinrich, Proc. U. S. Nat. Mus., 
vol. 57, p. 84, 1920 (larva, pupa, life history).—Essig, 
Insects of western North America, p. 709, 1926.—McDun- 
nough, Check list, No. 6145, 1939. 

Dasypyga alternosquamella stictophorella. Ragonot, N. Amer. 
Phycitidae, p. 5, 1887; Monograph, pt. 1, p. 206, 1893. 

Forewing pale salmon pink; basal area black dusted 
with white, this black area (from the upper vein of cell 
to inner margin) extending almost to middle of wing; 
the black and white scaling coarse and more or less 
raised in base and along outer margin of antemedial 
line which cuts the black areas as a narrow, pale (och- 
raceous red), obliquely curved line; subterminal line 
absent, but along termen a rather broad border of coarse 
black and whitish scales; on most specimens a clear 
white line extends outward from lower angle of cell 
along vein 5 towards and sometimes to the dark terminal 
border and usually enclosing a detached patch of black 
scales forming a discal spot at end of cell; above this 
(below vein 6) a fine red line runs to near outer dark 
margin and thence angles sharply to apex; on well 
marked and fresh specimens traces of a similar red line 
on the Jower fold in outer area. Hind wing very pale, 
shining, smoky fuscous with a faint dark line along 
termen. Alar expanse, 19-24 mm. 

Genitalia as given for the genus; figured from reared 
examples; cornutus of male penis about half as long as 
aedeagus, somewhat flattened, twisted, and bluntly 
pointed; penis also minutely scobinate towards apex. 

TyprE LocaLity: California (type in Paris Mus.). 

Foop puant: Razamofskya cryptopoda. 

DistrisutTion: Unirep Status: California, Lake 
Arrowhead (May); Arizona, Mohave County (Sept.), 
Williams (June); Colorado, Glenwood Springs, Monu- 
ment (May, June, Aug.); Washington, Bellingham 
(June), Seattle. Canapva: British Columbia, Kaslo 
(June). 

A species that can be at once recognized by its pecu- 
liar markings and coloration. The white longitudinal 
line on forewing is of varying length on different speci- 
mens but is present on all that I have seen, though 
sometimes weak. Its presence is the characteristic 
feature given by Ragonot for his variety stictophorella. 
It is presumably absent from the type of alternosqua- 
mella. Isuspect that the latter is an individual variant. 
Notes on the life history and descriptions of larva and 
pupa are given in my paper. 


70 UNITED STATES ‘NATIONAL MUSEUM BULLETIN 207 


Genus 34: Rampylla 


[Male: Venational division D. Transtilla present but variously 
modified. Harpe with apex of sacculus produced, strongly 
sclerotized and pointed. Hind wing triangulate; anal angle 
folded and produced; sex tufts and scalings on lower surface. 
Female: Venational division B. Bursa and ductus bursae 
simple; ductus seminalis from bursa. Hind wing with cell one- 
third the length of wing.] 


34. Genus Rampylla Dyar 


Rampylla Dyar, Ins. Inse. Menstr., vol. 7, p. 84, 1919. 
of genus: Rampzylla orto Dyar). 

Tongue well developed. Antenna pubescent; shaft 
of male slightly thickened. Labial palpus upturned, 
not reaching vertex in male, a trifle longer in female; 
second segment slightly rough scaled beneath; third 
segment bluntly acuminate, about half the length of 
second (shorter on male than female). Maxillary 
palpus squamous, small. Forewing smooth except for 
a slight, projecting scale tuft from inner margin near 
base on male; 11 veins; vein 2 from before lower outer 
angle of cell; 3 from the angle, well separated from 2; 
4 and 5 approximate at base and for a very short dis- 
tance beyond; 6 from below upper angle of cell, straight; 
10 from the cell; male without costal fold. Hind wing 
with vein 2 from before the lower outer angle of cell; 
3 from the angle and in the male almost as long as vein 
2; in the female considerably shorter; 4 and 5 very 
shortly stalked or contiguous for a short distance from 
cell, connate with 3; 7 and 8 contiguous or closely ap- 
proximate for a short distance from cell; cell in male 
one-fourth the length of wing, in female approximately 
one-third; discocellular vein curved; on male anal area 
(involving vein 1a) thickened and folded, forming a 
produced pocket, enclosing a long hair pencil; under- 
side of male wing with roughened scale or hair tufts on 
some of the veins. Highth abdominal segment of male 
with sternite developed as a narrow, sclerotized pocket. 
Metathorax of male with a stout pencil of spatulate 
scales from just above base of leg. 

Male genitalia with transtilla present, variously modi- 
fied (greatly reduced in lophotalis). Harpe with sac- 
culus produced into a strong free hook at apex. A long 
free spine associated with anellus (as in the four preced- 
ing genera). 

Female genitalia with bursa more or less finely scobin- 
ate and with some concentration of these fine scobina- 
tions but no. definable signum; ductus bursae simple, 
short (shorter than bursa except in lophotalis). An 
invaginated, sclerotized, dorsal pocket at apical end of 
seventh abdominal segment or (in lophotalis) a sclero- 
tized, granulate, dorsal pocket between ovipositor and 
eighth-segment collar. 

The length of vein 3 in proportion to 2 of hind wing 
places the males in our venational division D and, in 
conjunction with their decidedly triangulate hind wings 
and the sex-scalings and tuftings on their under surfaces, 
readily distinguishes the genus. The females on hind 
wing venation fall into division B. The four species 
here recognized are all tropical American. They ex- 


(Type 


hibit distinct specific differences in genitalia, color, and 
maculation. The transverse lines of forewing, in their 
rather close approximation and narrow black borders, 
are similar to those of Coptarthria to which Rampylla 
seems most nearly related. 


143. Rampylla orio Dyar 
Fieures 45, 248 


Rampylla orio Dyar, Ins. Insc. Menstr., vol. 7, p. 84, 1919. 


Forewing violaceous gray; under magnification ex- 
treme base and a rather broad area along inner margin 
to beyond middle tinted with purplish red; antemedial 
line obsolete; subterminal line narrow, blackish, rather 
close to and nearly parallel with termen, vertical from 
costa to vein 8, inwardly angled between veins 8 and 5, 
thence oblique to inner margin, bordered inwardly by a 
faint ocherous shade and outwardly by a narrow ocher- 
ous line; discal dots at end of cell separate, ocherous; a 
thin dark line along terminal margin. Hind wing trans- 
lucent white with a fuscous shade at apex and anal area 
yellow; veins not appreciably darkened; terminal mar- 
gin darkened only towards apex; on underside of hind 
wing (male) a yellow hair tuft at origin of veins 4 and 5 
from cell; a white fringe continuing outwardly on these 
veins and a similar white fringe on vein 7 above the 
yellow hair tuft. Alar expanse, 22 mm. 

Male genitalia with lateral portion of uncus on each 
side produced into a broad, strongly sclerotized, projec- 
tion armed at apex with a cluster of long, slender, strong, 
black spines. ‘Transtilla produced caudally into a bi- 
lobed, scobinate process fusing with reduced arms of 
gnathos. Tegumen with a long, stout, curved, free arm 
arising from base at each ventrolateral angle. Harpe 
with produced sacculus developed as a long, stout, taper- 
ing hook, curved across face of harpe. Anellus heart- 
shaped; associated spine straight. Aedeagus partially 
sclerotized; penis with some faint, sclerotized wrinklings 
and a few microscopic scobinations, otherwise unarmed. 

Tyrr Locatity;: Zacualpaén, México (type in USNM). 

Foop prant: Unknown. 

Known only from the male type. 


144, Rampylla polydectella (Schaus) 
Figure 732 
Salebria polydectella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 


p. 250, 1913. 
Rampylla polydectella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7, 
1919. 


Forewing brownish gray with a faint purplish tint; a 
narrow border along inner margin between the trans- 
verse lines and a somewhat broader area along termen 
dusted with dull, grayish white; antemedial line narrow, 
faint, pale gray, indicated chiefly by its narrow, black 
outer-bordering line, the line well out towards middle of 
wing and slightly sinnous, nearly vertical; subterminal 
line equally thin and pale gray with a similar black inner 
border, angled outward slightly at middle and nearly 
parallel with termen; a black line along terminal margin; 
discal spots small, confluent, ocherous; faint indication 


AMERICAN MOTHS OF THE. SUBFAMILY PHYCITINAE vel 


of black streaklets on veins 2, 3, and 4 just beyond cell. 
Hind wing glossy brown-gray; the veins faintly dark- 
ened and a fine dark line along termen. Alar expanse, 
20 mm. 

Female genitalia distinguished chiefly by the shape 
of eighth-segment collar and the invaginated, sclero- 
tized pocket from seventh segment. 

TYPE Locatity: Juan Vifias, Costa Rica (June; type 
in USNM). 

Foop puant: Unknown. 

Known only from the female type. 


145. Rampylla subcaudata (Dyar), new combination 
Ficures 249, 733 


Cerocanthia subcaudata Dyar, Ins. Insc. Menstr., vol. 7, p. 42, 


Forewing ocherous gray, the basal area to antemedial 
line purplish tinted; a similar purplish shade along outer 
border of the subterminal line; a large, somewhat 
darker (fuscous) patch surrounding the discal spot and 
extending from antemedial almost to subterminal line 
and from vein 2 to costa, darkest on veins 2 to 4; ante- 
medial line faint, narrow, ocherous, indicated by a thin 
blackish brown line forming its outer border, the latter 
near middle of wing, slightly sinuate and nearly vertical; 
subterminal line with a narrow, blackish brown, inner 
border, slightly outbent or angled at middle; discal 
mark a narrow ocherous spot along discocellular vein; 
a narrow blackish line along terminal margin. Hind 
wing of male subpellucid white with a faint yellowish 
tint, decidedly ocherous along inner margin; of female 
with pale smoky tint; underside of male hind wing with 
a yellowish scale tuft covering the bases of veins 2 to 
5. Alar expanse, 16 mm. 

Male genitalia with margins of uncus evenly rounded, 
lacking any sclerotized projections. Harpe with 
apical projection of sacculus a slender, upcurved 
hook; a long, stout, hair and scale tuft from a pad ad- 
jacent to base of sacculus. Penis armed with a narrow, 
weakly sclerotized, bladelike cornutus. Female geni- 
talia similar to those of polydectella, differing only in 
slight details in the shape of the eighth-segment collar 
and the sclerotized pocket from seventh segment (fig. 
733a). 

TypE Locauity: Cayuga, Guatemala (May; type in 
USNM). 


Foop PLant: Unknown. 

In addition to the male type there is one other speci- 
men in the National Collection from Quirigué, Guate- 
mala (May), a female with slightly darker hind wings 
whose genitalia are here figured. In addition there are 
before me two somewhat larger specimens (19 mm.), a 
male in the Janse Collection from San José, Costa Rica 
(“H. Schmidt, 8-11”), and a female from the British 
Museum labeled ‘‘S. E. Brazil, E. D. Jones, 1920-303.” 
The genitalia of the latter are almost identical with 
those of polydectella. The genitalia of the male are like 
those of the type of subcaudata. The two specimens 
are identical in size, color, and markings. Except for 


size (which is not significant) they agree superficially 
with the type of subcaudata. Despite the differences in 
fore and hind wing coloration I suspect that subcaudata 
may not be specifically distinct from polydectella. 


146. Rampylla lophotalis, new species 
Figures 250, 731 


Similar to subcaudata except: Ground color of fore- 
wing more yellowish; the dark areas, especially just 
preceding the antemedial line and in the dark area’ sur- 
rounding the discal spot, blackish; dark lines bordering 
the transverse lines blackish rather than blackish brown; 
antemedial line farther out on wing, extending from 
midcosta to inner margin beyond middle, narrowing the 
area between antemedial and subterminal lines. Hind 
wing with the scale tuft on underside blackish rather 
than yellow, the blackish shade extending to the costa 
and visible through the wing from above. Alar ex- 
panse, 17.5-18 mm. 

Male genitalia with two pairs of strongly sclerotized, 
curved, lateral projections from triangulate uncus. 
Gnathos entirely absent (unless the lower pair of pro- 
jecting arms from uncus can be interpreted as lateral 
arms of a gnathos, which is very doubtful, as there is 
no separation whatever between them and the uncus). 
Transtilla reduced to a shortened and slender, trans- 
verse, centrally bent band. Tegumen with a short, 
two-pronged projection from each ventrolateral angle. 
Anellus and harpe as in subcaudata. Female genitalia 
with a thick, sclerotized roll on the back of ovipositor; 
a broad, coarsely granulate pocket between ovipositor 
and collar; eighth-segment collar narrow; ductus bursae 
slightly longer than bursa. 

Typr Locatity: Jalapa, México (type in USNM, 
61330). 

Foop piant: Unknown. 

Described from male type and one female paratype 
from the type locality and one male paratype from 
Volcén Santa Maria, Guatemala (Nov., Schaus and 
Barnes, collectors). The male holotype bears a label 
in Hampson’s handwriting ‘‘Cerocanthia lophotalis 
Hampson, o type.”’ As far as I know Hampson never 
published a description of the species. 


Genus 35: Fulrada 


[Venational division D. Hind wing with veins 4 and 5 contiguous 
or closely approximate beyond cell (not stalked) ; cell one-fourth 
the length of wing; discocellular vein incomplete. Eighth ab- 
dominal segment of male with broad ventral tuft. Transtilla 
vestigial. Harpe with apex of sacculus not produced.] 


35. Fulrada, new genus 


Typr or Genus: Dasypyga querna Dyar. 

Tongue well developed. Antenna weakly pubescent; 
shaft of male simple. Labial palpus upturned, slender, 
reaching to vertex; third segment slightly shorter than 
second, acuminate. Maxillary palpus squamous, small, 
appressed to face. Forewing smooth; 11 veins; vein 2 


72 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


from before but near lower outer angle of cell; 3 from 
the angle, nearly equidistant from bases of 2 and 4; 4 
and 5 closely approximate or connate and contiguous, 
for a short distance beyond base; 6 from below upper 
angle, straight or very slightly bent towards base; 10 
from the cell, approximate to the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from before 
lower outer angle of cell; 3 from the angle, long (its 
free; length slightly shorter than 2); 4 and 5 contiguous 
and. closely approximate for slightly less than half their 
distances from angle of cell (not stalked); 7 and 8 closely 
approximate beyond cell; cell one-fourth the length of 
wing; discocellular vein incomplete. Highth abdominal 
segment of male with a broad, ventral hair or scale tuft 
and sternite developed as a narrow sclerotized pocket. 

Male genitalia with uncus subtriangulate, its apical 
margin truncate. Gnathos represented only by its 
lateral arms, to which (at their junction) is attached 
the base of subanal plate. Transtilla represented only 
by a bilobed central vestige behind (above) the anellus. 
Harpe with sacculus not produced. Anellus semi- 
tubular (querna) or a broad slightly curved plate with 
a greatly reduced, associated, free spine (carpasella). 
Vinculum stout, as broad or nearly as broad as long; 
terminal margin broad and slightly rounded. 

The foregoing description is incomplete, as females 
are unknown. ‘The genus is apparently close to both 
Anadelosemia and Rampylla, differing from the former 
chiefly on hind wing venation and from the latter on 
secondary male characters. 


147. Fulrada querna (Dyar), new combination 
Fieure 251 


Dasypyga querna Dyar, Proc. U. S: Nat. Mus., vol. 47, p. 331, 
1914. 


Forewing ocherous gray with a faint dusting of 
reddish scales on basal area and a very faint reddish 
tint over the area below the discal spots; a scattered 
blackish powdering in costal areas; antemedial line 
oblique, indicated only by a row of (4 or 5) well sepa- 
rated black dots forming its inner border and an outer 
black spot on costa; discal dots at end of cell, small, 
separated, black; a row of small but distinct black dots 
along termen. Hind wing translucent, smoky white, 
darker towards apex; veins faintly darkened and a 
narrow dark line along termen. Alar expanse, 11 mm. 

Male genitalia with anellus semitubular, oblong, 
bottle shaped, more or less involved with the vestigial 
hairy-lobed transtilla. Harpe with a strongly haired, 
transverse, sclerotized ridge extending from near base 
of costa to lower outer angle of cucullus. Aedeagus 
short; penis armed with a very weak, flattened cornutus. 
Tuft on eighth abdominal segment a row of very fine, 
hairlike scales. 


Typn Locauity: La Chorrera, Panamé (May; type 
in USNM). 

Foop prant: Unknown. 

Represented only by the male type. 


148. Fulrada carpasella (Schaus), new combination 
FigurE 252 


Piesmopoda carpasella Schaus, Zoologica, vol. 5, No. 2, p. 47, 
1923. 


Forewing white finely irrorated with black and 
brown; some pale tawny shading on the white ante- 
medial line along the inner margin of its black outer 
border, also postmedially below vein 2 and on discocel- 
lular vein; base black, this shade expanding’ obliquely 
to inner margin and outwardly edged by some pale 
tawny scaling; antemedial line oblique, indicated 
chiefly by its black outer border, the latter strongly 
contrasted from costa to lower margin of cell, very faint 
from cell to inner margin, slightly outcurved from costa; 
subterminal line parallel with termen, slightly sinuous 
and with well-contrasted dark outer and inner borders, 
the inner one a narrow black line, the outer a black, 
angulate costal dash continued as a rather broad 
brownish shade to tornus; discal dots separated, small 
black dots on the outer angles of cell; a row of well- 
contrasted black dots along termen. Hind wing 
whitish towards base, shading to fuscous outwardly, a 
rather broad brownish shade along termen. Alar 
expanse, 12 mm. 

Male genitalia with a vestigial free spine associated 
with the broad, large, slightly curved plate’ of anellus. 
Harpe with rather long, slender, erect clasper: Aedea- 
gus rather long, slender, straight. Highth abdominal 
tuft consisting of long, spatulate scales. 

Typr LocaLity: Conway Bay, Indefatigable, Galé- 
pagos Islands (Apr.; type in USNM). . 

Foop pirant: Unknown. 

Known only from the male type. 


Genus 36: Scorylus 


[Venational division B. Male antenna with shallow sinus in 
shaft at base. Hind wing with veins 4 and 5 stalked for about 
two-thirds; cell about one-third the length of wing; discocellular 
vein complete, curved. Highth abdominal segment of male with 
broad, ventral scale tuft. Metathorax with stout scale and 
hair tuft near base of leg. Transtilla vestigial. Anellus with- 
out trace of associated free spine. Harpe with apex of sacculus 
produced.] 


36. Scorylus, new genus 


TYPE OF GENUS: Scorylus cubensis, new species. 

Tongue well developed. Antenna finely pubescent; 
the male shaft with a shallow sinus at base (a slight ex- 
cavation covering several basal segments); within the 
sinus and extending slightly past it a thin layer of modi- 
fied appressed scales. Labial palpus upturned, reaching 
vertex, second segment slightly rough scaled; third seg- 
ment slightly shorter than second, acuminate. Maxil- 
lary palpus squamous. Forewing smooth; 11 veins; 
vein 2 from before lower outer angle of cell; 3 from the 
angle, nearer to 4 than to 2;4 and 5 closely approximate 
for a short distance from cell; 6 from below upper angle 
of cell, slightly bent towards base; 8 and 9 stalked for 
slightly more than half their lengths; 10 from the cell, 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 73 


approximate to the stalk of 8-9; male without costal 
fold. Hind wing with vein 2 from before, but close to 
lower outer angle of cell; 3 from the angle, contiguous to 
the stalk of 4-5 for nearly half the length of the stalk, 
the free length of 3 decidedly shorter than vein 2; 4 and 
5 stalked for about two-thirds their lengths; 7 and 8 
closely approximate for some distance beyond cell; cell 
about one-third the length of wing; discocellular vein 
curved. Highth abdominal segment with broad, stout 
ventral scale tuft and sternite developed as a narrow 
sclerotized pocket. A stout scale and hair tuft from 
metathorax at base of leg. 


Male genitalia with uncus hoodlike. Gnathos well 
developed, its apical projection a long, tapering spike. 
Transtilla represented by a modified central vestige. 
Harpe with sacculus produced at apex into a sclerotized 
hook; a strong hair tuft from lobe near base of sacculus; 
anellus a triangulate plate with short, blunt, broad lat- 
eral lobes; aedeagus moderately stout, slightly swollen 
and bent towards base; penis armed with an elongate, 
narrow cluster of short, thin spines. Vinculum stout, 
slightly tapering and somewhat longer than basal width. 

Female genitalia without signum but with a concen- 
tration of fine granulations in bursa near its junction 
with ductus bursae; ductus bursae and its genital open- 
ing simple; ductus seminalis from ductus bursae. 

This genus shows affinities to both Fulrada and 
Anadelosemia but is distinct from both. Its venation 
is similar to that of Anadelosemia, to which it appears to 
be most closely allied. 


149. Scorylus cubensis, new species 
Figures 253, 725 


Forewing white with blackish fuscous and reddish 
markings; basal area strongly irrorated with red and 
some scattered reddish scaling in the median white area; 
antemedial line indicated only by a blackish outer line 
from costa, expanding into a black spot in cell, continued 
thence as a very thin blackish fuscous line to inner mar- 
gin, bordered outwardly (from cell to inner margin) by 
an olivaceous patch; a faint triangulate olivaceous- 
fuscous shade over outer area from just beyond middle 
of inner margin to subterminal line at costa; a small 
black spot on midcosta; subterminal line sinuate, 
bordered inwardly by a strong, rather broad, blackish 
line and outwardly by a fainter reddish line continued 
from a strong black spot on costa; discal dots black, 
more or less confluent along discocellular vein; below 
them on the olivaceous fuscous shade a patch of reddish 
scaling; a row of black dots along termen. Hind wing 
translucent white; veins very faintly darkened near 
outer margin; a narrow dark shade along termen. Alar 
expanse, 12-13 mm. 

Male genitalia with outer margin of uncus evenly 
rounded. ‘Transtilla fragment in the shape of two short, 
oblong, pointed plates weakly joined at their bases. 
Apical projection of sacculus sharply upturned, mod- 
erately long, thornlike. Terminal margin of vinculum 


truncate. Female genitalia with bursa large, as long as 
ductus bursae. 

TYPE Locauity: Santiago Province, Cuba (type in 
USNM, 61331). 

Foop piant: Unknown. 

Described from male type and one male and one fe- 
male paratypes from the type locality (June, Dec., 
Schaus and Barnes, collectors). 


Genera 37-39: Davara to Piesmopoda 


[Venational division D. Forewing with veins 4 and 5 approxi- 
mate for a short distance from cell. Hind wing with cell less 
than one-third the length of wing. Male genitalia with uncus 
bifid (divided to base); harpe with strong, hooked clasper from 
near apex of sacculus.] 


37. Genus Davara Walker 


Davara Walker, List, pt. 19, p. 1020, 1859.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 530, 1901. (Type of genus: 
Davara azonazsalis Walker.) 

Homalopalpia Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 332, 1914. 
(Type of genus: Homalopalpia dalera Dyar. New synonymy.) 

Eucardinia Dyar, Ins. Insc. Menstr., vol. 6, p. 188, 1918; vol. 7, 
p- 50, 1919. (Type of genus: Ulophora caricae Dyar.) 

Tongue well developed. Antenna of male (except in 
rufulella) with basal segment enlarged, deeply notched 
and with a brush of short fine bristles in the notch 
(fig. 254d); male shaft simple or flattened and dilated 
towards base, pubescent. Labial palpus upcurved; on 
male (except in rufulella) reaching well above vertex; 
broadly scaled; third segment considerably shorter than 
second. Maxillary palpus squamous. Forewing more 
or less rough scaled at base on male; 11 veins; vein 2 
from before, but near lower outer angle of cell;3 from the 
angle, nearer to 4 at base than to 2; 4 and 5 approximate 
for a short distance from cell; 6 from below upper angle 
of cell, straight or very slightly curved towards base; 8 
and 9 long stalked; 10 from the cell, approximate to the 
stall of 8-9; male without costal fold. Hind wing with 
vein 2 from before the angle of the cell; 3 from the 
angle, long (its free length slightly shorter than 2); 4 and 
5 anastomosed for slightly less than half their lengths 
beyond angle of cell; 7 and 8 contiguous or shortly 
anastomosed beyond cell; cell less than one-third the 
length of wing; discocellular vein curved. Eighth 
abdominal segment of male with sternite developed as 
a narrow, sclerotized pocket. 

Male genitalia with uncus bifid (divided to base). 
Gnathos weak, a thin narrow band. Transtilla absent 
(except in interjecta). Harpe with strong, hooked 
clasper from near apex of sacculus; a long hair tuft from 
lobe near base of sacculus. Anellus a narrow curved 
plate with long, lateral arms; a stout, free spine associ- 
ated with anellus. Penis armed with a thin, narrow, 
curved, flattened, bladelike cornutus. 

Female genitalia with two signa, developed as small, 
granulate depressions; ductus bursae shorter than bursa, 
with paired cuplike plates behind genital opening or 
with genital opening simple (interjecta) ; ductus seminalis 


74 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


from bursa near its junction with ductus bursae. Collar 
of eighth segment complete. 

Davara and. the two following genera (Sarasota and 
Piesmopoda) form a compact group distinguished from 
all other American phycitid genera by the peculiar bifid 
development of their unci. Davara was described by 
Walker on the basis of a single female which he mistook 
for a male; it was referred as a synonym of the Old 
World Phycita by Hampson (1903). Dyar did not 
recognize it. His Homalopalpia was erected on male 
antennal and palpal structures which normally should 
be of generic value, but in this particular instance do 
not seem to hold as separating Davara from Piesmopoda. 
They fall down in the case of rufulella, which on genitalic 
characters of both male and female must be referred to 
Davara. In my opinion Davara and Piesmopoda should 
be retained as separate genera. Their species differ in 
habitus as well as in genitalic structure. In Davara the 
male transtilla is absent and the female bursa always 
has two signa. In Piesmopoda the transtilla is present 
and developed as two long, slender, curved free arms 
and the female’ bursa has a single signum or none. 
Unfortunately an anomalous species (interjecta) seems 
to upset the division. It has the female and all the 
secondary male characters of typical Davara but male 
genitalia of the Piesmopoda type. However, there are 
some minor differences in its male and female genitalia 
which may eventually allow its separation from both 
Davara and Piesmopoda under a separate generic desig- 
nation. For the present I am referring it tentatively to 
Davara. Its distinctive characters are discussed more 
fully under the specific description. 


150. Davara caricae (Dyar), new combination 
Fiaures 46, 254, 735 

Ulophora caricae Dyar, Proc. Ent. Soc. Washington, vol. 14, p. 
218, 1913. 

Homalopalpia dalera Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 332, 
1914. (New synonymy.) 

Eucardinia caricae (Dyar) Dyar, Ins. Insc. Menstr., vol. 6, p. 
139, 1918; Ins. Insc. Menstr., vol. 7, p. 50, 1919.—McDun- 
nough, Check list, No. 6119, 1939. 


Antenna of male with the brush in notch of basal 
segment black; the basal segment itself a very pale buff; 
basal segments of shaft decidedly flattened and broad- 
ened and with black serrations on upper edge. Labial 
palpus of male very broadly scaled, the second segment 
reaching well above vertex; less broadly scaled and 
somewhat shorter on female; reddish brown with a 
peppering of pale buff scaling on outer side and on male 
more or less shaded with blackish brown on inner side. 
Forewing tan-gray shaded with reddish brown; basal 
area, (on male) blackish brown ‘and rough scaled; a 
whitish or pale buff shade precedes the antemedial line 
and a similar transverse shade crosses the disk and 
includes the discocellular mark; the space between this 
transverse pale shade‘and antemedial line suffused with 
reddish brown (on some female examples almost purplish 
fuscous); a similar reddish brown suffusion over outer 
area; antemedial line oblique, slightly angled between 


cell and inner margin, faint, ocherous, bordered in- 
wardly and outwardly by narrow dark lines; subterminal 
line indistinct except for its brown inner and outer 
borders, sinuate; discal spots fused into a narrow lunu- 
late line on the discocellular vein. Hind wing soiled 
white, with a narrow fuscous shade along termen. Alar 
expanse, 14-18 mm. 
Male genitalia with a cluster of fine, long, hairlik 
spines surrounding the strong, free, forked spine asso- 
ciated with anellus; lateral arms of anellus considerably 
shorter than in other species of the genus. Female 
genitalia with a pair of strong ventrolateral ridges on the 
invaginated portion of the eighth-segment collar. 


Tyre tocauities: Miami, Fla. (caricae, in USNM); 
LaChorrera, Panama (dalera, in USNM),. 


Food pLant: Carica papayae (larvae in the fruit). 


Distrizution: Unitep States: Florida, Florida City 
(Apr., May), Fort Pierce (May), Miami (Apr., Dec.), 
Royal Palm State Park. Purrtro Rico: Bayamén 
(June, Dec.), El] Yunque (Apr.), Jajoma Alta (June), 
Lares (June, Sept., Nov.). Cusa: Baracoa (July, Aug.) 
Santiago de las Vegas (Feb., Mé4r.), Santiago Province 
(@Jan., June, Oct.). Harri: Damien (Aug.). Domini- 
can ReEpusiic: San Francisco Mts. (Aug., Sept.). 
Trinipap: Mt. Harris. GuatEemata: Cayuga (Apr., 
May), Chejel (June, Aug.), Purulhi (June, July). 
Costa Rica: Guapiles (May), Juan Vinias (Feb., May, 
June), Silio (May). Ecuapor: Quevedo (‘‘Nov.- 
Dec.”). ; 

The types of both caricae and dalera are males. Dyar 
(1919) recognized the generic synonymy of Hucardinia 
and Homalopalpia but never admitted the specific 
identity of their types, although he had every reason 
to suspect it. His designation of new Cuban types for 
caricae in 1918 is obviously invalid, for he had previ- 
ously (1913) designated Florida types for what he ad- 
mitted was the same species under the same name. I 
believe he was correct in his surmise that caricae is the 
same as columnella Zeller, but as I have never examined 
any Colombian examples of any Davara species I 
hesitate to propose the synonymy. 


151. Davara columnella (Zeller), new combination 
Figure 736 


Myelois columnella Zeller, Horae Soc. Ent. Rossicae, vol. 16, 
pp. 209, 210, 1881. 
Piesmopoda columnella (Zeller) Ragonot, Monograph, pt. 1, p. 
161, 1893. 
Homalopalpia columnella (Zeller) Dyar, Ins. Insc. Menstr., vol. 
7, p. 49, 1919. 
I have seen no specimens from the type locality but 
have before me a photograph of the female type which 
agrees with the females of a series from Costa Rica 
identified by Schaus as columnella and correctly referred 
by Dyar to his dalera, and I have little doubt that the 
names columnella and caricae (=dalera) stand for the 
same species. The female genitalia of Zeller’s type 
(here figured) show some trifling differences in the size 
of the plates behind genital opening and in the eighth- 
segment collar from those of typical caricae; but these 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 75 


are probably only individual differences. Variations 
as great are exhibited among reared examples of caricae 
from different localities. Alar expanse, 15 mm. 
Typxr Locatity: Honda, Colombia (type in BM). 
Foop puant: Unknown (presumably papaya). 


152. Davara nerthella (Schaus), new combination 


Ficure 738 
Piesmopoda nerthella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 
11, p. 247, 1913. 
Homalopalpia euthales Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 403, 1914. 
Homalopalpia nerthella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7, 
p. 49, 1919. 

Similar to caricae except averaging somewhat larger; 
brush in notch of basal segment of male antenna 
ocherous, the segment itself smaller; the subbasal pale 
shade before the antemedial line more extended on the 
male and much more so on the female, reducing con- 
siderably the blackish brown shading of the basal area 
and forming with the slightly paler antemedial line a 
broad pale pinkish ocherous band; antemedial line 
straight, not angled below cell, its inner and outer 
bordering lines very faint and narrow, reddish brown; 
discal dots separate (never fused) and often only the 
lower one distinguished; on females more or less of an 
ocherous tint over the brownish median shade following 
the antemedial line, especially towards costa. Alar 
expanse, 19-22 mm. 

Male genitalia like that of the following species 
(paranensis) except tuft from near base of sacculus pale 
yellow. Highth-segment collar without ridges on the 
invaginated portion; narrower on venter than in other 
species; posterior ventral margin without notch or but 
slightly notched. The extent of this notching is indi- 
vidually variable in all the species and is not a reliable 
character for specific separation. 

TYPE LOCALITIES: Juan Vifias, Costa Rica (nerthella, 
in USNM); Orizaba, México (euthales; in USNM). 

Foop prant: Unknown. 

DistrisutTion: Mfxico: Jalapa, Orizaba. Guatr- 
MALA: Volcdén Santa Maria (June, July). Costa Rica: 
Juan Vinas (Jan.); other Costa Rican females without 
further locality designation in Janse Collection. 

Doubtfully distinct from paranensis. The chief dif- 
ferences between the males are in coloration and be- 
tween the females in the width and notching of the 
eighth-segment collar. There is the same amount of 
individual variation in wing color and maculation as in 
other species, some specimens generally paler than 
others, some with the subterminal line distinct, others 
with it almost obsolete. 

Dyar (1919) recognized the synonymy of his euthales 
with nerthella. The type of the former is a male, of 
the latter a female. 


153. Davara paranensis (Dyar), new combination 
Fieure 255 


Homalopalpia paranensis Dyar, Ins. Insc. Menstr., vol. 7, p. 49, 
1919, 


Similar to nerthella except brush in notch of basal seg- 
ment of male antenna brownish; subbasal pale shade 
before antemedial line of forewing much narrower, re- 
stricted (especially on females) by a greater extension 
of the black basal scaling; median area along costa 
rather strongly tinted with ocherous drab or reddish 
ocherous (on the type and one female). Alar expanse, 
18-19 mm. 

Male genitalia with no fine spine cluster surrounding 
the free, forked spine associated with anellus. Tuft 
from near base of sacculus black. Female genitalia 
similar to those of azonazsalis. 

Type LocaLity: Castro, Parané, Brazil (type in 
USN). 

Foop prant: Unknown. 

At first glance Dyar’s type seems quite distinct from 
males of nerthella, its palpi and antennae being much 
darker and the pale transverse antemedial shade less 
contrasted against the ground color. However the 
specimen is stained and none of the other examples of 
the species before me is in very good condition. Be- 
sides the type, I have before me a female from the type 
locality, another female from Santa Catarina, Brazil, 
and a male from the British Museum collection from 
Sao Paulo, Brazil, that is without abdomen but a good 
match for Dyar’s type. Both it and the type had been 
originally identified by Hampson as columnella Zeller. 

I suspect that when additional South American ma- 
terial is available paranensis will prove to be nothing 
more than a variety of nerthella, and that eventually 
both nerthella and paranensis will fall to azonarsalis of 
Walker. 

154. Davara azonaxsalis Walker 
FIaure 737 
Davara azonazsalis Walker, List, pt. 19, p. 1020, 1859. 
Phycita azonazsalis (Walker) Hampson, in Ragonot, Monograph, 
pt. 2, p. 531, 1901. 

I have seen no specimens of Davara from the type 
locality and none from anywhere of the size of Walker’s 
type (30 mm.). A photograph of the type and its geni- 
talia supplied by Tams are before me. The antemedial 
line of forewing shows an angulation between cell and 
inner margin similar to that on typical caricae. The 
subterminal line is rather distinctly marked and the 
basal area much like that of females of nerthella, but 
not so strongly contrasted. None of these features, 
however, is enough for specific separation. 

The female genitalia show an appreciably wider 
eighth-segment collar than that of nerthella and a dis- 
tinct notch in its ventroposterior margins. The much 
smaller female of paranensis from Castro has similar 
but somewhat smaller genitalia. 

TypE Locauity: Rio de Janiero, Brazil (type in Ox- 
ford Univ. Mus.). 

Foop piant: Unknown. 


155. Davara (?) interjecta, new species 
Fiaurus 256, 734 


Male antenna with basal segment enlarged and 
notched as in caricae, but the usual brush of fine bristles 


76 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


in the notch here replaced by smooth, appressed, silvery 
and ocherous scales; first segment of shaft broadly 
flattened, the shaft shortly ciliate (the cilia about as 
long as width of shaft). Labial palpus broadly scaled, 
the second segment reaching well above vertex, deep 
red-brown to blackish brown on outer side, the third 
segment black scaled, at least on inner surface. Thorax 
deep brown mixed with blackish and buff scaling. Fore- 
wing reddish brown more or less shaded with black in 
median area, the raised scaling at base mixed black and 
dark red-brown; antemedial line obsolete or very faintly 
indi¢ated on the paler specimens, when distinguishable 
oblique, straight, narrow, pale buff; lower discal spot 
at end of cell usually distinct, blackish; subterminal 
line obscure or obsolete; an interrupted row of partially 
confluent black dots along termen. Hind wing trans- 
lucent, white, the veins more or less darkened, a narrow 
fuscous shade along termen. 

Female generally paler than the male. Palpi, head, 
thorax, and forewing light reddish brown with a faint 
sprinkling of whitish scales, especially bordering ter- 
men; ground color darkened in median area along costa; 
usually a broad, olivaceous shade along inner margin 
at base; no defined transverse markings. 

Alar expanse, 17-19 mm. 

Male genitalia of the Piesmopoda type with incom- 
plete transtilla developed as a pair of opposed, long, 
slender, curved, strongly sclerotized arms, their apices 
curved towards each other; anellus a narrow, broadly 
V-shaped band, with long, curved, slender, haired lat- 
eral arms, their apices bulbed. Female genitalia with 
genital opening simple, no sclerotized plates behind the 
opening. 

Type tocauity: El Yunque, Luquillo Mts., Puerto 
Rico (type in Cornell Univ.; paratypes in Cornell and 
USNM, 61332). 

Foop prant: Unknown. 

Described from male type and two male and five fe- 
male paratypes from the type locality (1,500-2,000 ft., 
Apr. 22, 23, and Mar. 29,1930, Cornell lot 795, sub. 38, 
40, and 9, 'W. T. M. Forbes, ‘collector) and two male 
paratypes from San Francisco Mts., Santo Domingo 
(Sept. 1905, A. Busck, collector). The males, with the 
exception of the holotype and one paratype, are badly 
rubbed. The females are in better condition. 

This species is referred with reservations to Davara. 
In its structural characters it straddles both Davara and 
Piesmopoda and fits comfortably in neither genus. Its 
female genitalia are those of Davara except that the 
usual sclerotized plates behind genital opening are lack- 
ing. Its male genitalia are those of Piesmopoda except 
that the apices of the elements of transtilla pomt toward 
(rather than away from) each other; and the apices of 
the lateral arms of anellus are swollen (bulbed). In all 
known species of Piesmopoda the apices of the elements 
of transtilla point away from each other and the apices 
of the lateral arms of transtilla are pointed. The male 
antenna of interjecta also is abnormal for Davara in that 
there is no brush of fine spines in the notch of the basal 
segment. In my opinion these differences could permit 


generic separation of interjecta from both Davara and 
Piesmopoda; but they are so slight that, without further 
evidence from biology or the early stages, a new generic 
designation does not seem justified at this time. 


156. Davara rufulella (Ragonot), new combination 
FicureEs 257, 739 


Piesmopoda rufulella Ragonot, Bull. Soc. Ent. France, 1888, 
p. cxxxix; Monograph, pt. 1, p. 165, 1893. 

Male antenna simple. Labial palpus upturned, not 
broadly scaled, cylindrical; reaching to slightly above 
vertex on male, nearly to vertex onfemale; terminal 
segment acuminate. Forewing pale red-brown; the 
basal area a trifle paler with some faint olivaceous shad- 
ing in inner margin; antemedial line oblique, straight, 
obscure, indicated chiefly by its outer dark margin, 
which begins as a blackish smudge on costa and contin- 
ues to inner margin as a slight darkening of the ground 
color; more or less blackish dusting in the cell beyond 
antemedial line; on paler specimens some peppering of 
white scales in median and outer areas, especially on 
midcosta; subterminal line obsolete or very faintly indi- 
cated; discal spots obscure, confluent along discocellular 
vein.. Hind wing translucent white; the veins dark- 
ened; a faint, narrow fuscous shade along termen. Alar 
expanse, 13-16 mm. 

Male genitalia without cluster of fine spines surround- 
ing free spine associated with anellus, the free spine 
itself short, stout, broadly forked. Lateral arms of 
anellus straight. Harpe with apex of cucullus pointed; 
clasper long, stout, strongly curved. Female genitalia 
with two pairs of contiguous sclerotized plates behind 
genital opening; ductus bursae sclerotized at genital 
opening. 

Type Locauity: Puerto Rico (type in Zool. Mus. 
Univ. Berlin). 

Foop piant: Unknown. 

Distrizution: Purrro Rico: Bayamén (Apr., Sept. ), 
Comerio (Nov.), Jajoma Alto (June), Lares (Dec. ), La 
Sardinera (Dorado, June), Palmas Abajas (June), San 
German (Apr.). 

I have seen no specimens from any but Puerto Rican 
localities. As mentioned in the discussion of the genus, 
rufulella is aberrant in that it lacks the modified basal 
segment of male antenna and the characteristic broad 
scaling of the male labial palpi. However, the genitalia, 
both male and female, are characteristic of the genus, 
showing only specific differences from. other species of 
Davara. 

38. Genus Sarasota Hulst 


Sarasota Hulst, Journ. New York Ent. Soc., vol. 8, p. 222, 1900. 
(Type of genus: Sarasota plumigerella Hulst). 
Cuba Dyar, Ins. Inse: Menstr., vol. 7, p. 50, 1919. 
genus: Cuba furculella Dyar. New synonymy.) 
Tongue well developed. Antenna simple in both 
sexes, shaft weakly pubescent. Labial palpus up- 
turned, reaching to slightly above vertex; third segment 
slightly over half the length of second, acuminate. 
Maxillary palpus small, squamous. Forewing smooth; 
11 veins; vein 2 from before but near lower outer angle 


(Type of 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE TE 


of cell; 3 from the angle, approximately equidistant at 
base from 2 and 4, 4 and 5 approximate for a short 
distance from cell; 6 from upper angle of cell, slightly 
bent towards base, connate with the stalk of 8-9; 10 
from the cell, well separated from the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from before 
lower outer angle of cell; 3 from the angle, nearly as 
long as 2, connate with 4; all veins long; 4 and 5 con- 
tiguous or weakly anastomosed for about half their 
lengths from cell; 7 and 8 weakly anastomosed beyond 
cell, their free elements long; cell slightly less than one- 
fourth the length of wing; discocellular vein curved for 
a short distance from lower angle, thence vertical. 
Kighth abdominal segment of male with sternite de- 
veloped as a narrow, sclerotized pocket. 

Male genitalia similar to those of typical Davara 
except: Free spine associated with anellus entirely 
lacking; harpe with two-pronged clasper and apical end 
of sacculus produced; penis with a pair of curved, flat- 
tened, bladelike cornuti. Female genitalia with ductus 
bursae sclerotized for a short distance from genitalia 
opening; a single, angulate, projecting plate behind 
genital opening; otherwise as in Davara. 


157. Sarasota plumigerella Hulst 
FIGURE 258 


Sarasota plumigerella Hulst, Journ. New York Ent. Soc., vol. .8, 
p. 222, 1900.—Grossbeck, Bull. Amer. Mus. Nat. Hist., 
vol. 37, p. 128, 1917.—McDunnough, Check list, No. 6120, 
1939. 

Basal segment of antenna red spotted with black. 
Prothorax wine red; mesothorax and metathorax black. 
Forewing black at base followed by a whitish ocherous 
shade; antemedial line oblique, straight, ocherous 
white, preceded by more blackish scaling, followed (es- 
pecially on costa) by a strong black shade; remainder 
of wing blackish fuscous stained with wine red, the 
costa on outer half distinctly reddish; subterminal line 
very faint, sinuate; discal spots obscure or absent; 
terminal dots more or less confluent, blackish; cilia red. 
Hind wing dull, translucent white; a dark shade at 
apex and a narrow dark line along termen. Midtibia 
with a strong yellow hair tuft. Alar expanse, 11-14 
mm. 

Male genitalia with terminal margin of vinculum not 

produced at the edges but exhibiting no other specific 
characters. Female genitalia slightly smaller but other- 
wise not distinct from those of furculella. 

Typrr Locauity: Palm Beach, Fla. (type in USNM). 

Foop puiants: Laguncularia racemosa, Coccolobis 
uvifera (larvae feeding under a light silk webbing on 
the leaves and flower buds). 

Distrrisution: Florida, Palm Beach, Ramrod Key 
(Apr.), Stock Isl. (Apr.), Sugar Loaf Key (Apr.). 


158. Sarasota furculella (Dyar), new combination 
Ficures 48, 259, 740 
Cuba furculella Dyar, Ins. Insc. Menstr., vol. 7, p. 50, 1919. 


Larger and paler than plumigerella. Thorax and 
forewing violaceous gray more or less tinted with red- 


dish; the blackish shade of plumigerella replaced in 
Surculella by red; antemedial line dull white with a 
strong ocherous tint, especially towards inner margin, 
its dark outer border red; some white dusting in median 
costal area; discal dots more distinct, separate, red or 
fuscous; subterminal line more distinct, narrow, sinuate, 
dull white; terminal dots few, but distinct and separated, 
blackish.’ Hind wings white with a faint smoky tint; 
the veins very slightly darkened. Midtibial hair tuft 
as in plumigerella. Alar expanse, 14-16 mm. 

Male genitalia with terminal margin of vinculum 
slightly produced at the sides; otherwise as in plumi- 
gerella. Female genitalia a trifle larger than those of 
plumigerella but showing no specific characters. 

TyprE LocaLity: Santiago, Cuba (type in USNM). 

Foop puant: Unknown. 

Distrisution: Cusa: Baracoa, Matanzas, Santiago 
(June). Purrtro Rico: Dorado (May), Puerto Real 
(Vieques Isl., Apr.). Dominica (Dec.). Viren Is- 
LANDS: Kingshill (St. Croix, June, Dec.). 

Very close to plumigerella but apparently a distinct 
species, separable chiefly on color. 


159. Sarasota ptyonopoda (Hampson), new combination 


Phycita ptyonopoda Hampson, Ann. Mag. Nat. Hist., ser. 6, vol. 
16, p. 347, 1895. 

Hyalospila ptyonopoda (Hampson) Hampson, in Ragonot, Mono- 
graph, pt. 2, p. 530, pl. 56, fig. 9, 1901. 

I have seen no examples of this species. From the 
description and the figure in the Ragonot Monograph 
the coloration and markings must be similar to those of 
jurculella. According to Hampson the male has a 
strong, long tuft of scales from the hind femur. Clarke 
has reexamined the type and tells me that there is a 
large expanded dorsal tuft from the base of the hind 
femur and that there is no tuft on the midtibia. These 
differences in tufting distinguish the species easily from 
either plumigerella or furculella. A fine photograph of 
the male genitalia, taken by Clarke, is before me. 
They are like those of furculella Dyar. Unfortunately 
the abdomen had been glued on the type so there will 
remain some question of placement for the species until 
another male from the type locality and with tufted 
hind femora is recorded. Alar expanse, 16 mm. 

Typr nocauity: St. Vincent, Windward Islands, 
British West Indies (type in BM). 

Foop puant: Unknown. 

In the U. S. National Museum is a rubbed male from 
Jalapa, México, labeled in Hampson’s handwriting 
“Phycita ptyonopoda Hampson,” but it cannot be that 
species. What remain of the legs show a yellow hair 
tuft on midtibia and no trace of tuft on the hind femur 
(the hind tibiae are missing). This specimen is an 
Atheloca sp., close to bondari. 


39. Genus Piesmopoda Zeller 


Piesmopoda Zeller, Isis von Oken, 1848, p. 863.—Hulst, Phy- 
citidae of N. Amer., p. 132, 1890.—Ragonot, Monograph, 
pt. 1, p. 158, 18938. (Type of genus: Pzesmopoda rubicun- 
della Zeller.) 


78 UNITED STATES NATIONAL MUSEUM, BULLETIN. 207 


Discopalpia Ragonot, Monograph, pt. 1, p. 167, 1893. 
genus: Myelots flavicans Zeller. New synonymy.) 

Amphycitopsis Dyar, Ins. Inse.’ Menstr., vol. 7, p. 45, 1919. 
(Type of genus: Amphycitopsis isabella Dyar. New syn- 
onymy.) 

Tongue well developed. Antenna pubescent or 
shortly ciliate (the cilia no longer than width of shaft) ; 
on male, shaft simple, notched at base or with curved 
excavation (a long sinus) towards base; basal segment 
of male sometimes swollen but not notched or otherwise 
modified. Labial palpus upturned, of varying length, 
not reaching vertex (isabella, fratella) or extending to or 
above vertex; third segment acuminate or (ragonott) 
broadly dilated with scales. Maxillary palpus minute 
(folded over tongue), filiform or subsquamous (third 
segment slightly dilated with scales). Forewing smooth; 
venation as in Sarasota except vein 8 normally consider- 
ably closer to 4 at base than to 2; male without costal 
fold. Hind wing as in Sarasota. Highth abdominal 
segment ‘with sternite developed as a narrow, sclero- 
tized pocket. 

Male genitalia with uncus bifid. Gnathos incom- 
plete, represented by its broad lateral arms, separated 
at their apices, between which lies a rather well sclero- 
tized subanal’ plate. Transtilla well developed but 
incomplete, consisting of a pair of long, very slender, 
curved arms whose sharply pointed apices are directed 
away from each other. Harpe with @ strong, hooked 
clasper from near apex of sacculus; a strong hair tuft 
from lobe near base of sacculus. Anellus a curved plate 
(U- or V-shaped) with long, slender, haired and bluntly 
pointed lateral arms. Penis armed with a narrow, 
flattened and more or less curved, bladelike cornutus; 
sometimes with two such cornuti. 

Female genitalia with single signum or none, signum 
when present developed as a small granulate depres- 
sion; ductus bursae with genital opening simple or more 
or less sclerotized, but without the paired cuplike plates 
found in Davara; ductus seminalis from bursa near its 
junction with ductus bursae. EHighth-segment collar 
more or less invaginate (except in apocerastes and 
montella) . 

A distinct genus defined and easily recognized by its 
male and female genitalia. As used by Ragonot and 
subsequent authors the name covered a composite of 
disparate elements. The antennal and palpal char- 
acters upon which Presmopoda, Discopalpia, and Am- 
phycitopsis were distinguished are in this instance of no 
more ‘than specific significance. They are discussed 
more fully under their type species. 

The genus is apparently limited to tropical America. 
Nothing is known of the life history or host association 
of any of the species. 


(Type of 


160. Piesmopoda rubicundella Zeller 
Figure 260 


Piesmopoda rubicundella Zeller, Isis von Oken, 1848, p. 864.— 
Ragonot,; Monograph, pt. 1, p. 160, 1893. 


Antenna of male with basal segment swollen, triangu- 
late; shaft notched at base. Labial palpus slender, 


third segment acuminate, reaching well above vertex. 

Forewing pale (olivaceous ocherous) at base and for 
some distance along inner margin; the median costal 
area broadly white, heavily dusted with red scaling; 
more or less of this red dusting also on costal half at 
base; antemedial line not distinguishable; subterminal 
line distinct, straight, oblique, pale reddish or ocherous 
gray bordered by dark purplish fuscous lines; the entire 
outer fourth of wing more or less suffused with purplish 
fuscous; discal dots more or less distinct, separated, red 
or, reddish fuscous. Hind wing pale smoky fuscous, 
lighter towards base and semitransparent; veins dis- 
tinctly darkened. Alar expanse, 13 mm. 

Male genitalia having harpe with cucullus narrowly 
elongate, evenly tapering. Anellus a broadly sclero- 
tized, V-shaped band. 

TyPE Locanity: Brazil (type in Mus. Univ. Berlin). 

Foop piranr: Unknown. 

I have examined the male type but have seen no 
other examples from Brazil. The type is not distin- 
guishable from males of the following species, (canthom- 
era) except for minor differences in male. genitalia, 
which may or may not be significant. 


161. Piesmopoda xanthomera Dyar 
Ficures 266, 745 
Piesmopoda zanthomera Dyar, Proc. U. S. Nat. Mus., vol. 47; 
p. 332, 1914. 
Piesmopoda xanthozona Dyar, Ins. Insc. Menstr., vol. 7, p. 45, 
1919 (new synonymy). 

With the same male characters as rubicundella and 
superficially not distinguishable from it. I suspect that 
zanthomera is nothing more than a synonym or variety 
of rubicundella, but this cannot be proven until females 
of the latter from Brazil are available. Dyar’s two 
names apply only to the sexes (canthomera to the females 
and zanthozona to the males). Alar expanse, 13-17 mm. 

Male genitalia, figured from type of zanthozona, differ 
from those of rubicundella chiefly in the stronger tufting 
from the base of the harpe, a character of very doubtful 
specific value.’ Female genitalia, figured from type of 
zanthomera and checked with those of females from all 
localities here cited, distinguished chiefly by the shape 
of the eighth-segment collar and its sclerotized invagi- 
nate portion and the pair of corrugate patches on the 
ventral surface of the latter. Bursa with signum: 
Genital opening simple. MIO 

TYPE LocAuitTiEes: La Chorrera, Panama (anthomera, 
in USNM); St. Jean Maroni, French Guiana (xantho- 
zona, in USNM). 

Foop prant: Unknown. 

DistRiBuTION: GUATEMALA: Cayuga (Apr., May). 
Costa Rica: San José (July). Panam: La Chorrera 
(May), Porto Bello (Oct.).. France Guiana: Cayenne, 
St. Jean Maroni. 


162. Piesmopoda trichomata (Zeller) 
Figure 744 


Myelois trichomata Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 
194, 1881. : 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 79 


Ragonot (Monograph, pt. 1, p. 160) makes this a 
synonym of rubicundella, considering it merely the fe- 
male of the latter. This is a very dubious placement. 
The Zeller type material in the British Museum con- 
sists of two female cotypes (photographs of which are 
before me) alike in all details and similar to rubicundella 
except that the antemedial line is indicated by broken 
remnants of its outer border, a dark dash from costa, 
and a more or less diffused dark spot on lower margin of 
cell. The lower half of wing is somewhat suffused into 
dark shading and contrasted against the whitish mid- 
costal area, as in rubicundella. Zeller’s figure is mis- 
leading in that it shows much of the wing bright yellow, 
asin floricans. Alar expanse, 13 mm. 

The genitalia of the female cotype here figured, ac- 
cording to Tams and Clarke, agree in all details with 
those of the other cotype. The latter should be con- 
sidered the holotype, as it is the better preserved speci- 
men. Bursa with signum. Genital opening simple. 
The narrow eighth-segment collar easily identifies the 
species. It is unlike any other in the genus that I have 
seen. 

Typsr Locautity: Honda, Colombia (type in BM). 

Foop piant: Unknown. 


Known only from the type specimens. 


163. Piesmopoda flavicans (Zeller) 
Fiaures 262, 746 


Myelois flavicans Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 193, 


Discopalpia flavicans (Zeller) Ragonot, Monograph, pt. 1, p. 167, 
1893 (in part, 2). 
Piesmopoda flavicans (Zeller) Dyar, Proc. U. 8. Nat. Mus., vol. 
47, p. 333, 1914. 
Piesmopoda fratella Dyar, Ins. Insc. Menstr., vol. 7, p. 45, 1919 
(new synonymy). 
Antenna of male simple. Labial palpus slender; on 
male not reaching vertex; on female extending slightly 
above vertex (as in isabella). 


Forewing bright yellow; outer third purple dusted 
with blackish towards apex; the yellow ground color ex- 
tended further outward on costa and inner margin than 
at middle; antemedial line obsolete; subterminal line 
faint, narrow, whitish, straight, oblique and close to 
outer margin. Hind wing whitish, stained with smoky 
fuscous towards apex and on the outer parts of the veins. 
Alar expanse, 14-15 mm. 

Male genitalia like those of zanthopolys except that 
elements of transtilla are stouter, lateral arms of anellus 
are bent sharply away from each other, and terminal 
margin of vinculum is acutely rounded (rather than 
straight). Female genitalia similar to those of rantho- 
polys except for slight differences in the structure of 
eighth-segment collar. 

Type tocauities: Honda, Colombia (flavicans, in 
BM); St. Jean Maroni, French Guiana (fratella, in 
USNM). 


Foop prant: Unknown. 


DistrisuTion: Frencu Guiana: Cayenne, St. Jean 
Maroni (Mar.). Cotomspra: Honda. 

Specimens of five different species in the U. S. Na- 
tional Collection had been identified by Hampson, 
Dyar, and Schaus as flavicans. Among them was one 
female from French Guiana which Hampson identified 
(correctly) as flavicans and which Dyar later included in 
his series of fratella. Clarke and Tams have checked 
our genitalic figures of the several Piesmopoda species 
with the female type of flavicans in the British Museum 
and inform me that the genitalia of fratella agree in all 
details with those of favicans. Ragonot also misidenti- 
fied Zeller’s species (at least as far as males are con- 
cerned) and on the strength of their peculiar male palpi 
erected the genus Discopalpia, with flavicans as type. 
Dyar (1914) noted the misidentification and renamed 
the males of flavicans Ragonot (not Zeller) as Discopal- 
pia ragonoti. Later evidence from genitalia justifies 
the new specific name; but nomenclatorily the type of 
Discopalpia must remain flavicans Zeller, and Dyar’s 
citation of a new type (flavicans Ragonot=ragonoti 
Dyar) is not justified, despite Ragonot’s misidentifica- 
tion and the characters derived therefrom for his genus. 
Fortunately these characters (as far as Piesmopoda are 
concerned) are of specific significance only, so Disco- 
palpia would fall, however we interpreted its type; 
but I for one hold that when a specific name is cited as 
type of a new genus, that species remains the type of 
the genus regardless of what specimens were before the 
author of the genus or how he described his generic 
concept. The decision is one of nomenclature and 
not of zoology. 


164. Piesmopoda ragonoti (Dyar), new combination 


Figures 265, 747 


Discopalpia ragonoti Dyar, Ins. Insc. Menstr., vol. 7, p. 44, 1919. 
Discopalpia flavicans Ragonot (in part, co’; not Zeller), Mono- 
graph, pt. 1, p. 167, 1893. 

Antenna of male simple; basal segment cylindrical; 
shaft without notch or other modification. Labial 
palpus very long, the second segment reaching well 
above vertex in both sexes; on male the third segment 
broadly expanded with long scales, fan shaped; a strong 
admixture of black scaling on outer sides of the palpi of 
both sexes. 

Forewing as in flavicans. Alar expanse, 14-16 mm. 

Male genitalia distinguished chiefly by the very 
heavy, black, broad-haired tuft from base of sacculus of 
harpe. Female genitalia with signum; genital opening 
simple; distinguished from those of other yellow-winged 
species by minor differences in the configuration of the 
eighth-segment collar; closest to those of zanthopolis 
Dyar. 

TYPE LOCALITY: 
USNM). 

Foop prant: Unknown. 

Disrrisution: México: Distrito Federal. Guatr- 
mata: Cayuga (Feb., Apr., May). Costa Rica: Juan 
Vifias (Jan.). 


Cayuga, Guatemala (type in 


80 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


165. Piesmopoda isabella (Dyar), new combination 
Figures 264, 749 
Amphycitopsis isabella Dyar, Ins. Inse. Menstr., vol. 7, p. 45, 
1919 

Male antenna simple. Labial palpus short, hardly 
reaching vertex on male and but a trifle beyond vertex 
on female. 

Forewing as in flavicans. Alar expanse, 18-20 mm. 

Male genitalia with cucullus of harpe subtriangulate, 
harpe wider in proportion to its length and less evenly 
tapering than in other species except apocerastes which 
has similar male genitalia. Female genitalia with 
signum; genital opening simple; distinguished by the 
conficuration of the eighth-segment collar. 

TYPE LocALITy: Juan Vifias, Costa Rica (type in 
USNM). 

Foop prant: Unknown. 

Represented by the male type and a slightly larger 
female from the type locality (Jan.). The latter had 
been identified by Schaus as “Piesmopoda flavicans 
Zeller.”’ The species served as type for Dyar’s genus 
Amphycitopsis, erected solely on the basis of the short 
labial palpi of the male. 


166. Piesmopoda xanthopolys Dyar 
Figures 261, 748 


Piesmopoda xanthopolys Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 
332, 1914 

Male antenna simple. Labial palpi slender, extend- 
ing above vertex in both sexes, somewhat longer on 
female than on male. 

Forewing as in flavicans. On the female from the 
type locality the purplish shading on the outer area of 
the forewing is somewhat more extended and the yellow 
area of the wing proportionally more restricted than in 
flavicans; but this difference seems to be individual 
rather than specific. In zanthopolis as well as the other 
species with coloration similar to flavicans the extent of 
purple shading is variable between the sexes and even 
among individuals of one sex. Alar expanse, 13-16 mm. 

Male genitalia figured from specimen from La Chor- 
rera, Panamd. Their most obvious feature seems to 
be the rather short arms of the bifid uncus (proportion- 
ally shorter than those of any other Piesmopoda except 
flavicans which has stouter transtilla and differently 
shaped anellus and vinculum). Female genitalia with 
signum present. Genital opening simple. Very close 
to those of flavicans, with incurvation of posterior- 
ventral margin of eighth-segment collar similar and ex- 
hibiting only minor differences in the collar otherwise. 

TyPE Locauity: Porto Bello, Panam& (type in 
USNM). 

Foop prant: Unknown. 

Distrisution: PanamA: Corozal (July), La ae 
rera (May), Porto Bello (Sept., Dec.). 

Known only from the original type series. Dyar in 
1919 (Ins. Insc. Menstr., vol. 7, p. 44) placed zantho- 
polys in the synonymy of flavicans. The two species 
are very close, but apparently distinct. 


167. Piesmopoda parva, new species 
Figures 263, 750 


Male antenna simple except for a very slight incur- 
vation of the shaft towards base. Labial palpus slender, 
reaching vertex; terminal segment acuminate. 


Forewing yellow with a slight olivaceous tint; costa 
tather broadly margined from base to near apex with 
white faintly peppered with red scaling; a few red and 
black scales at extreme base and for a short distance 
from base along inner margin; no antemedial line; sub- 
terminal line straight, oblique, close to termen, with 
narrow purplish fuscous borders and preceded by a fus- 
cous shade which extends, triangularly, almost to the 
cell. Hind wings translucent white with a faint smoky 
tint, darkening slightly towards apex. Alar expanse, 
10-11 mm. 

Male genitalia distinguished by the slender arms of 
the divided uncus and the slender, naillike spine asso- 
ciated with anellus. Female genitalia with signum; 
genital opening simple. Distinguished by the broad 
and deep excurvation in posteroventral margin of the 
eighth-segment collar. 

Typr tocauity: La Chorrera, Panamé (type in 
USNM, 61333). 

Foop prant: Unknown. 

Described from male type from the type locality 
(May), one male paratype from Cabima, Panam4 (May 
1911), and one female paratype from Taboga Isl., 
Panamé (Feb. 1912), all collected by A. Busck. These 
specimens had been included by Dyar among his para- 
types of Piesmopoda xanthomera. They are somewhat 
rubbed but otherwise in good condition. The species 
is the smallest of the Presmopoda. 


168. Piesmopoda semirufella (Zeller) 
Figure 752 


Myelois semirufella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 
196, 1881. 

Piesmopoda semirufella (Zeller) Ragonot, Monograph, pt. 1, p. 
160, 1893.—Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 332, 
1914; Ins. Insc. Menstr., vol. 7, p. 46, 1919. 

Several different species have been identified as semi- 
rufella.. I have seen but one example that can be defi- 
nitely placed to Zeller’s name, a female from Cayuga, 
Guatemala, collected by Schaus and Barnes (Jan.). 
The genitalia of this specimen have been checked by 
Clarke with the genitalia of the type of semirufella and 
he finds them identical. They have the signum present 
and a broad, strongly sclerotized plate at genital open- 
ing. The latter structure at once identifies the female. 
Males of the species have not been properly associated. 

Superficially, semirufella is not distinguishable from 
females of apocerastes Dyar. Indeed, several females of 
the latter in both the British Museum and U. S. Na- 
tional Collections had been identified as Zeller’s species. 
Alar expanse, 16 mm. 

TypEr Locatiry: Colombia (type, 2, in BM). 

Foop piant: Unknown. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAEB 81 


169. Piesmopoda apocerastes Dyar 
Fieure 751 


Piesmopoda apocerastes Dyar, Ins. Insc. Menstr., vol. 7, p. 45, 


Male antenna with basal segment cylindrical, slender; 
shaft with a long sinus (involving about eight of the 
basal segments) lined by flattened blackish scales which 
terminate in a slight tuft at the outer extremity of the 
sinus. Labial palpus cylindrical, slender, reaching to 
slightly above vertex. 

Forewing, except for a whitish border along costa, 
suffused reddish brown to the naked eye, very slightly 
darkened towards outer margin (under magnification 
the ground color shows a strong under tinting of oliva- 
ceous ocherous); whitish costal border peppered with 
scattered red scaling, a concentration of these along ex- 
treme costal margin; discal dots separate, red; ante- 
medial line obsolete or, at most, faintly indicated on 
some specimens by an obscure, narrow, dark, trans- 
verse shade; subterminal line faint, weakly bordered by 
narrow dark (reddish fuscous) lines. Hind wings smoky 
white to pale smoky fuscous, darkening towards apex 
and outer margin; the veins darkened. Alar expanse, 
15-16 mm. 

Male genitalia similar to those of isabella Dyar. Fe- 
male genitalia without signum; a narrow, strongly scle- 
rotized plate at genital opening with weakly sclerotized, 
anterior, lobelike projection. 

TYPE LOCALITY: Juan Vifas, Costa Rica (type in 
USNM). 

Foop puant: Unknown. 

Distrisution: México: Jalapa. Costa Rica: Juan 
Vifias (May, Nov.). Dominica (British West Indies, 
Feb.). Frencno Guiana: St. Jean Maroni. Brazit: 
Parané, Castro. 

A distinct species easily identified by its female geni- 
talia. Several of the females before me in the National 
Collection had been identified by Hampson as Piesmo- 
poda semirufella. The genitalia of female specimens 
from all the above-mentioned localities have been 
checked. 

170. Piesmopoda montella Schaus 


Fiaurs 743 


Piesmopoda montella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 
11, p. 247, 1913. 

Labial palpus of female slender, reaching slightly 
higher than vertex. 

Forewing light olivaceous brown; costal margin to 
subterminal line broadly white irrorated with red- 
brown, the extreme costal edge black at base, reddish 
brown at middle; no trace of any antemedial line; sub- 
terminal line slightly outcurved below vein 6, bordered 
inwardly and outwardly from costa to vein 2 by blackish 
bands into which some reddish scales are intermixed; 
discal dots separated, reddish brown; a few blackish 
dots on terminal margin. Hind wing pale, semihyaline 
brown, darkening towards outer margin; the veins 
darkly outlined. Alar expanse, 24 mm. 

Female genitalia without signum; bursa copulatrix 


small, oblong; ductus bursae very short and broad, al- 
most as broad at middle as the bursa, weakly sclerotized 
at genital opening, finely sclerotized otherwise. 
Kighth-segment collar simple, not fused ventrally. 

Typp tocauity: Mount Pods, Costa Rica (May; 
type in USNM). 

Foop prant: Unknown. 

A distinct species easily identified by its large size 
and distinctive genitalia; represented only by the fe- 
male type. 


Genus 40: Atheloca 


[Venational division C. Forewing with veins 4 and 5 connate 
and contiguous or partially anastomosed for one-third their 
lengths from cell; 6 from upper angle of cell, connate with the 
stalk of 8-9, bent towards base. Hind wing with cell less than 
one-fifth the length of wing; discocellular vein oblique, straight. 
Male genitalia with uncus hoodlike; lateral arms of gnathos fus- 
ing into anellus; harpe broadly angled at base of cucullus.] 


40. Atheloca, new genus 


Typr or Genus: Nephopteryz subrufella Hulst. 

Tongue well developed. Antenna of male simple, 
shaft pubescent. Labial palpus slender, upturned, 
reaching to or a trifle above vertex; third segment 
shorter than second, acuminate. Maxillary palpus fili- 
form. Forewing smooth; 11 veins, vein 2 from before 
lower outer angle of cell; 3 from the angle, much nearer 
to 4 at base than to 2; 4 and 5 connate, contiguous or 
partially anastomosed beyond base for one-third of 
their lengths; 6 from upper angle of cell, curved towards 
base and connate with the stalk of 8-9; 10 from the cell, 
approximate to the stalk of 8-9; male without costal 
fold. Hind wing with vein 2 from before lower outer 
angle of cell; 3 from the angle, nearly as long as vein 2, 
closely approximate to the stalk of 4-5 at base; 4 and 
5 stalked for half their lengths; 7 and 8 anastomosed 
for most of their lengths beyond cell, the free element 
of vein 8 very short; cell less than one-fifth the length 
of wing; discocellular vein oblique, straight. Abdomen 
of male with a pair of invaginated hair tufts at base; 
sternite of eighth segment developed as a sclerotized 
digitate pocket. 

Male genitalia with uncus hoodlike, triangulate, 
densely haired on outer surface. Gnathos represented 
only by its lateral arms which fuse into anellus at their 
apices. Transtilla absent. Harpe broadly angled at 
base of cucullus. Anellus a stout, triangulate plate 
with strongly sclerotized, sharply out-curved, smooth 
lateral arms (these latter may possibly represent ele- 
ments of a divided transtilla fused with the anellus, but 
this is extremely doubtful). Aedeagus and penis simple. 
Vinculum longer than greatest width, but slightly taper- 
ing to truncate terminal margin. 

Female genitalia with signa present in the form of 
two small scobinate patches; ductus bursae with a small, 
weakly sclerotized collar near the junction with bursa 
copulatrix; genital opening simple, ductus seminalis 
from bursa near the junction of bursa and ductus bur- 
sae; eighth-segment collar completely fused ventrally. 


82 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


171. Atheloca subrufella (Hulst), new combination 
Figures 267, 741 


Nephopteryx subrufella Hulst, Ent. Amer., vol. 3, p. 182, 1887. 

Nephopteryz filiolella Hulst, Ent. Amer., vol. 4, p. 117, 1888 (new 
synonymy). 

Piesmopoda subrufella (Hulst) Hulst, Phycitidae of N. Amer., 
p. 133, 1890.—Ragonot, Monograph, pt. 1, p. 166, 1893.— 
Grossbeck, Bull. Amer. Mus. Nat: Hist., vol. 37, p. 129, 1917. 

Piesmopoda filiolella (Hulst) Hulst, Phycitidae of N. Amer., p. 
133, 1890,—Ragonot, Monograph, pt. 1, p. 166, 1893. 

Sarasota subrufella (Hulst) Barnes and McDunnough, Check list 
of the Lepidoptera of Boreal America, No. 5558, 1917.— 
McDunnough, Check list, No. 6121, 1939. 

Sarasota filiolella (Hulst) Barnes and McDunnough, Check List 
of the Lepidoptera of Boreal America, No. 5559, 1917.— 
McDunnough, Check list, No. 6122, 1939." 

Hyalospila ptychis Dyar, Ins. Insc. Menstr., vol. 7, p. 49, 1919 
(new synonymy). 

Forewing ocherous fuscous shaded with reddish or 
purplish red except along costa; costa at extreme base 
edged with black, between the transverse lines rather 
broadly bordered by dull white sprinkled with red 
scales; antemedial line sometimes obsolete, when pres- 
ent indicated by a transverse black band interfused with 
reddish and preceded by a narrow dusting of white 
scales; subterminal line faint, pale, bordered inwardly 
and outwardly by narrow blackish or purplish red 
bands; discal spots usually distinct (at least the lower 
one), well separated, blackish (rarely with a touch of 
red); a more or less distinct row of blackish dots along 
termen. Hind wing smoky white, translucent; the 
veins darkened and a distinct dark shade along termen. 
Midtibia of male with strong hair tuft from base on 
inner side. Hind tibia of male without appreciable 
hair tuft. Alar expanse, 12-19 mm. 

Male genitalia as given for the genus. Female geni- 
talia with sclerotized portion of ductus bursae very nar- 
row. 

TypE Locauities: Florida (subrufella, nm AMNH, ex 
Rutgers); “Texas” (filiolella, in AMNH, ex Rutgers); 
Cuba. (piychis, in USNM). 

Foop prant: Unknown. 

Distrisution: Unrrep States: Florida, Chokolos- 
kee, Christmas Harbor (Mar.), Duardia (May), Fort 
Drum, Hastings (Aug., Sept.), Lake Alfred (May), 
Paradise Key (Mar.), Royal Palm State Park (June, 
Sept.), Stemper (May, July, Aug.), Vero Beach (Apr.), 
Winter Park (July, Aug.). Cusa: Pinar del Rio, no 
specific locality (type of ptychis). VIRGIN ISLANDS: 
Kingshill (St. Croix, ““Nov.—Dec.’’). 

The types of both subrufella and jfiliolella are females, 
neither of which bears a locality label; subrufella bears 
a label containing only the number ‘‘61”’; filzolella only 
a date label “March.” The latter is without abdomen 
and in very poor condition; but exhibits no difference 
from the type of subrufella except its somewhat larger 
size. In the National Museum there is a female from 
Christmas Harbor (‘‘March’’) labeled by Hulst ‘“Ne- 
phopteryx filiolella, type.”” It is an exact match for the 
type in the Rutgers Collection, as are three other large 
females from Florida in the National Collection, obvi- 
ously the same as subrufella. We have a large series of 


the species but no Texas examples, and I doubt very 
much the correctness of Hulst’s citation as the type 
locality of his filiolella. The species is obviously a 
tropical one which has extended its range to Florida. 
Dyar’s ptychis is merely a Cuban example, differing in 
no wise from typical subrufella. 


172. Atheloca bondari, new species 


Hyalospila ptychis Bondar (not Dyar), Rev. de Ent., Brazil, vol. 
11, p. 199, 1940.—Lepesme, Les insectes des Palmier’s, p. 
343, 1947 (Paris). 

Similar to subrufella except that the pale costal area 
of forewing is less contrasted and conspicuous. The 
genitalia male and female exhibit no essential differences 
from those of subrufella. The one distinguishing char- 
acter, seems to be a strong, dorsal, yellow hair tuft from 
the base of the male hind tibia. This is lacking from 
all specimens of subrufella and is an addition to the 
similar tuft on male midtibia, present on both subrufella 
and bondari. Alar expanse, 15-16 mm. 

TypE Locauity: Baia, Brazil (type in USNM, 61335). 

Foop PLant: Cocos nucifera (Lepesme also records 
C. coronata, C. vagans, Attalea funifera, and A. piassa- 
bossu). 

Described from male type and three male and five 
female paratypes all from the type locality and reared 
(June and July 1939, under Bondar Nos. 2521 and 2561) 
from larvae feeding in the seeds and at the base of the 
fruits of Cocos nucifera. These were received from Dr. 
Gregorio Bondar, for whom the species is named. He 
gives a good account of the habits of the species in the 
above-cited paper. J am responsible for the misidenti- 
fication to ptychis, for at the time I overlooked the differ- 
ence in leg tuftings between ptychis and the Brazilian 
specimens. 


Genera 41 and 42: Praedonula and Peadus 


[Venational division D. Forewing with veins 4 and 5 closely 
approximate for a short distance from cell; vein 6 straight; 10 
from the cell. Hind wing with cell one-fourth to one-fifth the 
length of wing; discocellular vein more or less curved. Male 
antenna with a shallow sinus in shaft involving the first half 
dozen segments. Male genitalia with a stout hair tuft from near 
base of sacculus.] 


41. Praedonula, new genus 


Type or cenus: Phycita almonella Dyar. 

Tongue well developed. Male antenna with a shallow 
sinus in shaft at base (involving the first six segments), 
the sinus containing a scattering of minute papillalike 
setae and a narrow ridge of scales along its outer edges, 
otherwise pubescent. Labial palpus very slender, 
upturned, reaching to or slightly above vertex; third 
segment slightly shorter than second, acuminate. 
Maxillary palpus subsquamous (the scales on second 
segment expanded, on third rather long and drawing to 
® point). Forewing smooth; 11 veins; vein 2 from 
before but near lower outer angle of cell (nearer the 
angle in male than in female); vein 3 from the angle; 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 83 


4 and 5 closely approximate at base and for a short 
distance beyond; 6 straight, from slightly below upper 
angle of cell (male) or from the angle (female); 10 from 
the cell, closely approximate to the stalk of 8-9 (male) 
or slightly separated from it (female); male without 
costal fold. Hind wing with vein 2 from before lower 
outer angle of cell; 3 from the angle, nearly as long as 
2, approximate at base to the stalk of 4-5; 4-5 stalked 
for at least half their lengths (female), somewhat longer 
stalked on male; 7 and 8 contiguous or weakly anas- 
tomosed for a short distance beyond cell, the free 
element of 8 long; cell short, about one-fifth the length 
of wing; discocellular vein oblique, very slightly curved. 
Eighth abdominal segment with sternite developed as 
a narrow (digitate) sclerotized pocket. 


Male genitalia with uncus hoodlike, rounded. Gna- 
thos strongly developed, the lateral arms broad, ex- 
panded and curled at their extremities and supporting 
a sclerotized subanal plate with a short thornlike spur 
at its base. Transtilla absent. Harpe stout, simple, 
slightly broadened at middle; at base of cucullus a 
stout hair tuft. Anellus a heart-shaped plate with 
stubby lateral lobes. Penis unarmed. Vinculum stout 
(but sclerotized narrowly along its margins); slightly 
longer than broad; scarcely tapering. 


Female genitalia with signa present in the form of two 
small granulate patches; ductus bursae with genital 
opening surrounded by a sclerotized plate; ductus 
seminalis from bursa near junction of bursa and ductus 
bursae. Eighth-segment collar completely fused 
ventrally. 

This genus is obviously closely related to the preced- 
ing genus (Aftheloca) and that which follows (Peadus). 
From the former it differs in the male antennal character 
and the weak anastomosis of veins 7-8 of hind wing. 
From Peadus it differs chiefly in the simple (undivided) 
harpe and the much more strongly developed uncus and 
tegumen of its male genitalia. It has no close relation- 
ship to the Old World Phycita to which Dyar referred 
its type species. 


173. Praedonula almonella (Dyar), new combination 
Fiaures 47, 268, 742 


Phycita almonella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 333, 
1914. 


The type series (a male and two females) are some- 
what rubbed and the coloration and markings of fore- 
wing consequently obscured. Superficially the species 
resembles Atheloca subrufella; the ground color of fore- 
wing a gray brown, darkening in outer area and shading 
to ocherous brown along inner margin at base; the 
costal margin broadly margined with white rather 
heavily dusted with red scales; antemedial line not dis- 
tinguishable; subterminal line faint, oblique and close 
to outer margin; discal dots very faint, separated. 
Hind wing translucent white shaded with fuscous at 
apex; the veins slightly darkened; on underside of male 
& coarse yellow sex-scaling between costa and cell, along 
lower margin of cell, extending for a short distance along 


veins 2 and 3, and along’ vein lb from base for nearly 
half its length. Alar expanse, 12-14 mm. 

Male genitalia with terminal margin of vinculum 
evenly rounded; aedeagus slightly bulged at middle, a 
row of short teeth: along lateral edge towards apex. 
Female genitalia simple and membranous except for a 
fine sprinkling of minute scobinations and the two 
small signa. 

Type nocauity: Porto Bello, Panam& (type in 
USNM). 

Foop piant: Unknown. 

Distrisution: PanamA: La Chorrera (May), Porto 
Bello (May). 

Known only from Dyar’s original type series. 


42, Peadus, new genus 


Type or genus: Piesmopoda burdettella Schaus. 

Tongue well developed. Male antenna with a shal- 
low sinus in shaft at base (as in Praedonula except here 
the hollow of the sinus is overlaid with rather coarse, 
appressed scales and without any indication of a tuft 
or lateral scale ridge), pubescent. Labial palpus slen- 
der, upturned, reaching vertex; third segment shorter 
than second, acuminate. Maxillary palpus subsqua- 
mous (as in Praedonula). Forewing smooth; venation 
as in Praedonula except vein 2 further from lower outer 
angle of cell, and 6 from below upper angle. Hind wing 
with cell one-fourth the length of wing; discocellular 
slightly but evenly curved. Eighth abdominal seg- 
ment with sclerotized pocket of sternite long and 
needlelike and with a large, flattened, fanlike tuft of 
long slender scales. 

Male genitalia with uncus and tegumen greatly re- 
duced; the uncus a narrow, weakly sclerotized angulate 
band. Gnathos indistinguishable (burdetellus) or rep- 
resented only by a very weakly sclerotized, transverse 
band (dissitus). Transtilla absent. Harpe short, stout; 
sacculus broad and broadly produced at apex; giving 
the harpe a partially divided appearance; the free costal 
half of harpe strongly recurved and bearing two very 
stout spines, one on outer lower margin near the angle 
produced by the projection of sacculus and another at 
lower angle of cucullus; the cucullus itself narrow and 
greatly reduced; a strong hair tuft from intersegment 
adjacent to base of sacculus. Anellus a shallow, curved 
plate with stubby lateral arms. Aedeagus spined at 
apex; penis armed with a couple of more or less curved 
and crinkled sclerotized bands and a cluster of short, 
stout spines. Vinculum long, stout and evenly taper- 
ing; considerably longer than greatest width. 

Female genitalia with signum developed as a short, 
slender thorn; area of bursa immediately surrounding 
ductus seminalis scobinate and weakly sclerotized; 
ductus bursae much shorter than bursa, broad and 
more or less sclerotized; behind genital opening a pair 
of narrow elongate plates extending backward and fus- 
ing into the ventrally divided eighth-segment collar 
(except in subaquilellus); ductus seminalis from bursa 
near junction of bursa and ductus bursae. 


84 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


The genus is quite distinct from any other on male 
genitalia. Its closest relatives seem to be Praedonula 
and Hyalospila. On many genitalic characters it re- 
sembles the following genus (Gabinius), especially in 
the development of uncus and harpe; but separates 
from it on hind wing venation, especially the length 
and position of vein 2. 


174. Peadus burdettellus (Schaus), new combination 
Figures 269, 754 


Piesmopoda burdetiella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 
11, p. 247, 1913. 

Discopalpia semproniella Schaus, Ann. Mag. Nat. Hist., ser. 8, 
vol. 11, p. 249, 1913. 

Hyalospila burdettella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7, 
p. 48, 1919. 

Forewing pale brown shaded with red and blackish 
scales; the costal border white sparsely dusted with 
red scales and with medial costal edge reddish; ante- 
medial line obscure except on lower half, far out, oblique 
from costa to cell and below cell inwardly concave, 
white, bordered outwardly on costa by a faint reddish 
streak; some black scaling along basal half of vein 1b; 
a distinct black dot on lower vein of cell at middle, and 
on the vein 1b on the outer edge of the antemedial line; 
subterminal line sinuous, bordered inwardly by a nar- 
row, dark brownish shade and followed in outer area 
(especially towards apex) by some white dusting; discal 
dots distinct, separated, black, the lower one somewhat 
elongated; an irregular black line along terminal mar- 
gin reaching almost to apex. Hind wings translucent, 
smoky white, darkening outwardly; the veins slightly 
darkened and a fine dark line along termen.. Alar ex- 
panse, 19.5—20 mm. 

Male genitalia with no trace of sclerotized gnathos; 
aedeagus with apex bluntly pointed, bearing a line of 
short, coarse spines along its edge. Female genitalia 
with ductus bursae weakly sclerotized near genital open- 
ing; sclerotized plates behind genital opening narrow, 
bladelike; sclerotization of bursa near ductus seminalis 
slight. 

TYPE LocaLitizus: Mount Pods (Juan Vinas), Costa 
Rica (burdettella, in USNM); Juan Vifias (semproniella, 
in USNM). 

Foop prant: Unknown. 

Distrisution: Costa Rica: Juan Vifias (Jan.), 
Mount Pods (May). GuatEemaua: Volcén Santa Maria 
(July). 

Dyar established the above synonymy of Schaus’ 
species. The genitalia of their male types are identical. 


175. Peadus dissitus, new species 
Figures 270, 755 


Similar to burdetiellus in color and markings except 
for a distinct whitish longitudinal shade through the 
cell of forewing and a stronger accentuation of the black 
scaling; a thin black streak from base along half the 
lower fold and a similar, shorter black streak on it just 
before the subterminal line; outer margin of antemedian 
line indicated by strong black dots on upper and lower 


veins of cell and on vein 1b; inner dark margin of sub- 
terminal line broadened by black streak. Alar expanse, 
20-22 mm. 

Male genitalia with gnathos a weakly sclerotized 
transverse band; aedeagus with an expanded, flangelike, 
densely and finely spined apex. Other minor differ- 
ences from bwrdetiella (especially in the shapes of 
cucullus and the projecting part of sacculus of the harpe) 
are shown in the figures. Female genitalia with ductus 
bursae much shorter than in burdettella, strongly sclero- 
tized; sclerotized plates behind genital opening broad- 
ened at their bases; sclerotization of bursa near 
ductus seminalis appreciably stronger. 

Typn tocauiry: “S. E. Brazil’? [Parané?] (type in 
BM;; paratypes in BM and USNM, 61334). 

Foop PLANT: Unknown. 

Described from male type and three male and three 
female paratypes from the type locality, “EH. D. Jones, 
1920—803”’. 


176. Peadus subaquilellus (Ragonot), new combination 
Figure 753 


Hyalospila subaquilella Ragonot, Nouv. Gen., p. 11, 1888; susie 
graph pt. 1, p. 170, 1893; pt. 2, p. 38, fig. Ds, 1901. 

This species is known only from the female'type. “A 
drawing of its genitalia made by Clarke is figured: I 
have seen nothing to match Ragonot’s figure and de- 
scription which indicate a form with dark reddish brown 
forewing shaded somewhat with black at base and on 
the costa’ but without other markings; hind wing 
“transparent,” smoky, the veins and terminal margin 
appreciably darkened. Alar expanse, 20 mm. 

The generic placement is tentative, pending discovery 
ofamale. The female genitalia are not typical, lacking 
the sclerotized plates behind genital opening, but seem 
to indicate a closer relationship to, Peadus than to any 
other genus. 

Type LocaLiry: “Cerro Zunil, ” Guatemala (type i in 
BM). 

Foop purant: Unknown. 


Genus 43 : Gabinius 


[Venational division B. Forewing with veins 4 and 5 connate; 
2 from near lower outer angle of cell. Hind wing with 2 from 
close to lower outer angle of cell; 3 from the stalk of 4—5; cell less 
than one-third the length of wing. Uncus reduced, weakly 
sclerotized, triangulate. Tegumen greatly reduced but with 
strong, projecting lateral arms. Transtilla incomplete. Harpe 
reduced; apex of costa spined. Vinculum short, stout.] 


43. Gabinius, new genus 


Typr or Genus: Promylea paulsoni Ragonot. 

Tongue well developed. Antenna of male simple, 
pubescent. Labial: palpus upturned, reaching above 
vertex; second segment somewhat broadly scaled; third 
segment shorter than second, bluntly acuminate. Max- 
illary palpus squamous. Forewing smooth; 11 veins; 
vein 2 from before but near lower outer angle of cell, 


nearly as close to 3:at base as 3 is to 4;3 from the angle; 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 85 


4 and 5 connate; 6 from below upper angle of cell, 
straight; 8 and 9 long stalked, the free element of 9 very 
short; 10 from the cell, approximate to the stalk of 8-9 
for a short distance; male without costal fold. Hind 
wing with vein 2 from close to lower angle of cell; 3 
from the stalk of 4-5; 4 and5 very long stalked; 7 and 8 
closely approximate beyond cell; cell less than one-third 
the length of wing; discocellular vein curved. Highth 
abdominal segment with sternite developed as a short 
(stubby) sclerotized pocket with thin, curved lateral 
arms extending into a slender U-shaped plate, fringed 
with moderately long scales. 


Male genitalia with uncus reduced, broader than long, 
triangulate and weakly sclerotized. Tegumen support- 
ing a pair of long, broad, pointed, strongly sclerotized, 
backwardly projecting arms; otherwise greatly reduced. 
Gnathos absent. ‘Transtilla incomplete; its elements 
pointed towards each other, their apices expanded and 
nearly touching. Harpe short, nearly as broad as long; 
costa broadly sclerotized, but shorter than remainder of 
harpe, bearing a stout, rather long, projecting spine at 
apex; cucullus much reduced, bearing a similar, stout 
but shorter spine on costal edge at apex; sacculus 
broadly sclerotized, but not produced. Anellus a broad 
crescentiform plate; its lateral lobes reduced to weakly 
haired knobs. Aedeagus smooth, straight; penis armed 
with a pair of curved, more or less wrinkled, sclerotized 
plates. Vinculum stout, as broad as long; its terminal 
margin evenly rounded. 


Female genitalia with signum developed as a thin, 
keellike blade on a narrow, elongate plate; ductus bursae 
shorter than bursa and with a sclerotized collar near 
simple genital opening; ductus seminalis from’ bursa 
near junction of bursa and ductus bursae. 


A distinct genus, apparently most closely related to 
Peadus but falling into a different venational division 
(B). The shape and structure of the uncus, the re- 
duced tegumen, and the short harpe with its stout 
projecting spines suggest the relationship to Peadus. 
The very short cell of hind wing separates it from most 
genera of division B. It is only remotely related to 
Promylea, to which Ragonot referred its type species. 


177. Gabinius paulsoni (Ragonot), new combination 


Ficures 271, 756 


Promylea paulsoni Ragonot, Nouv. Gen., p. 12, 1888; Mono- 
graph, pt. 1, p. 208, 18938. 

Ground color of forewing olivaceous gray strongly 
tinted with vinous brown in the dorsal area, this latter 
shade extending obliquely almost to apex; costal area 
from base to subterminal line, including the cell and 
tapering to costa beyond it, white with a scattered 
peppering of red-brown scales; costal edge from ante- 
medial line to above end of: cell edged with blackish 
brown; antemedial line distinct on lower half of wing, 
vertical with a slight inward concavity, followed out- 
wardly by a narrow blackish brown band and inwardly 
by a blotch of the same shade; the antemedial line 
indistinct and oblique on upper area of wing; subtermi- 


nal line, narrow, faint, sinuate, whitish, followed and 
preceded for a short distance from costa by blackish 
brown shadings; a scattering of white scaling in the 
terminal area below apex; lower discal spot blackish 
brown, more or less distinct, the upper very small and 
faint; a blackish irregular line along terminal margin, 
not reaching to apex. Hind wing translucent, yellowish 
white with a smoky tint towards apex and along ter- 
minal margin. Alar expanse, 20-23 mm. 

Male genitalia with characters as given for the 
genus. 

TYPE Locauity: Quillota, Chile (type in Paris Mus.). 

Foop puant: Unknown. 

The female in the National Collection matches in 
every detail Ragonot’s description and figure of the 
male type (Monograph, pl. 10, fig. 6). It is labeled 
“Chile, Silva.’ Superficially, the maculation and color 
resemble those of the Honora species. 


Genera 44-46: Ceracanthia to Drescoma 


[Venational division B. Forewing with veins 4 and 5 slightly 
separated at cell. Hind wing with cell less than half the length 
of wing (sometimes very short); 4 and 5 strongly stalked. Eighth 
sternite of male developed as a digitate pocket, sometimes the 
latter flattened and distorted. Male genitalia with vinculum 
very long, sclerotized only along its margins, arched dorsally 
(like a bent hairpin), its terminal margin broad; harpe with 
tufts on a projecting arm from base of sacculus.] 


44, Genus Ceracanthia Ragonot 


Ceracanthia Ragonot, Monograph, pt. 1, p. 230, 1893. (Type of 
genus: Ceracanthia vepreculella Ragonot.) 
Procandiopa Dyar, Ins. Inse. Menstr., vol. 7, p. 50, 1919. (Type 
of genus: Procandiopa mamella Dyar. . New synonymy.) 

Tongue well developed. Antenna pubescent; basal 
segment elongate, cylindrical (longer and more heavily 
scaled in male than female); shaft of male with an 
elongate, shallow sinus at base, from middle of sinus a 
short, sharp, sclerotized spine, a similar, shorter spine 
at apical end of sinus. Labial palpus upturned, reach- 
ing above vertex, cylindrical; third segment nearly as 
long as second, acuminate. Maxillary palpus sub- 
squamous (scaling of second segment somewhat ex- 
panded); folded across base of tongue. Forewing 
smooth; 11 veins; vein 2 from near lower outer angle 
of cell; 3 from the angle, about equidistant from 2 and 
4; 4 and 5 slightly separated at base, thence divergent; 
6 from slightly below upper angle of cell, straight; 10 
from the cell, approximate'to the stalk of 8-9 for a very 
short distance from cell; male without costal fold 
Hind wing with vein 2 from before, but near lower outer 
angle of cell; 8 from the stalk of 4-5; 4 and 5 stalked for 
half their lengths; 7 and 8 closely approximate for a 
short distance from cell; cell slightly over one-third the 
length of wing; discocellular vein curved. Eighth ab- 
dominal segment with sternite developed as a strongly 
sclerotized, digitate pocket. 

Male genitalia with uncus hoodlike (not tapering). 
Apical process of gnathos a simple, elongate, slender 
hook. Transtilla absent.’ Harpe broad, stout, sacculus 


86 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


partially divided towards apex and with a projecting 
arm from its base supporting a heavy hair tuft. Anellus 
a rather narrow band with short, stubby, lateral arms. 
Aedeagus somewhat curved and with a slightly more 
sclerotized bulge from outer third; penis with a few 
minute scobinations or some fine sclerotized wrinkles, 
otherwise unarmed. Vinculum elongate (considerably 
longer than greatest width), U-shaped, sclerotized only 
(and narrowly) along its margins, arched dorsally (like 
a bent hairpin). 

Female genitalia with signum; ductus bursae and 
genital opening simple; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

This genus is uncomfortably close to the genus: fol- 
lowing (Megarthria), from which it differs chiefly in 
the closer approximation of vein 2 of hind wing to the 
lower outer angle of cell, the decided stalking of 3 with 
4-5, and the slightly longer cell. The male antennal 
character on which Ragonot and Dyar erected their 
genera is probably here (as in Megarthria) of specific 
rather than generic significance. 


178. Ceracanthia mamella (Dyar), new combination 
Fieaures 13, 272, 278, 757 


Procandiopa mamella Dyar, Ins. Insc. Menstr., vol. 7, p. 51, 1919. 


Forewing yellowish white along costa; remainder of 
wing a purplish shade with a scattered peppering of 
blackish scales especially along base of median fold; 
antemedial line obscure, far out on wing and outwardly 
angled in cell, indicated chiefly by a discontinuous, nar- 
tow, blackish outer bordering line; subterminal line 
somewhat more distinct, smuate, yellowish white, fol- 
lowed on costa by a short blackish dash and bordered 
inwardly by a black line which expands into patch at 
middle; discal dots: blackish; terminal dots confluent 
forming a blackish line along the outer margin. Hind 
wing pale smoky brown; the veins faintly darkened and 
a distinctly smoky shade along termen. Alar expanse, 
15-16 mm. 

Male genitalia with ontee margin of harpe evenly 
rounded. Female genitalia with signum a flat, some- 
what granulate plate, bearing a short thorn near its 
center. 

Tyre tocauity: Rio Trinidad, Panama (type in 
USNM). 

Foop piuAnt: Unknown. 

Distrisution: PanaMA: Rio Trinidad (Mar., May). 
GuatEmaLa: Cayuga (May). 

Represented only by the original type series, a male 
and female from the type locality and a male and female 
from Guatemala. 

The sinus in the male antenna (fig. 278) appears 
smoothly scaled to the naked eye but under magnifica- 
tion shows several minute, erect, papillalike scales, simi- 
lar to those on Megarthria peterseni but less conspicuous. 


179. Ceracanthia vepreculella Ragonot 
Figure 273 
Ceracanthia vepreculella Ragonot, Monograph, pt. 1, p. 230, 1893. 
_I have seen no examples of this species but from 


Ragonot’s description and figure it appears to be similar 
to mamella except that the general color is a more uni- 
form yellowish gray without the contrastingly paler 
costa and the purplish shade on the remainder of the 
wing, characteristic of mamella. It is also a larger 
species. Alar expanse, 23 mm: 

Male genitalia with outer margin of harpe angulate. 

Typrr Locatity: Loja, Ecuador (type in Paris Mus.). 

Foop prant: Unknown. 


45. Genus Megarthria Ragonot 


Megarthria Ragonot, Monograph, pt. 1, p. 156, 1893. 
genus: Myelois petersenz Zeller.) 

Characters similar to those of Ceracanthia except: 
Vein 2 of hind wing further removed from lower outer 
angle of cell; the cell itself shorter, lessithan one-third 
the length of wing; vein 3 connate with the stalk of 4-5 
or contiguous with it for a short distance from the angle 
of cell, but not from the stalk; eighth abdominal seg- 
ment of male with sternite developed usually as a 
laterally flattened, more or less bent, digitate pocket, 
the supporting lateral arms of the sternite bearing a 
pair of scale tufts. 

Male genitalia with anellus an elongate shield; trans- 
tilla absent (except in schaust). 

Female genitalia: with or without signum; ductus 
seminalis from bursa adjacent to junction of bursa and 
ductus bursae. 

The shaft of the male antenna has a rather broad 
sinus at base (as in Ceracanthia) but is specifically vari- 
able. The lower outer angle of cell in the hind wing is 
as far out as in Ceracanthia, but the discocellular vein 
curves inward more deeply, making the cell itself BPEREs 
ciably shorter than that of Ceracanthia. 

In the National Collection all the specimens aoae 
the male type of cervicalis Dyar had been identified as 
peterseni Zeller. They are strikingly similar in color 
and maculation, but their structures show that four 
species are present among the males and at least two 
among the females, With our. present knowledge the 
sexes cannot be associated with any certainty; so until 
something 1 is known of their host association and the 
species are reared it seems a safer procedure to anticipate 
later synonymy and give separate names to the males 
and females rather than to link the females nomencla- 
torily to any of the male types. 


(Type of 


180. Megarthria peterseni (Zeller) 
Fieures 14, 274, 279 


Myelois petersenit Zeller, Horae' Soc. Ent. Rossicae, vol. 16, P. 
198, 1881. 

Megarthria petersent (Zeller) Ragonot, Monograph, pt. 1, p. 187, } 
1893 


Giéountl color of forewing white on costal half of wing, 
olivaceous brown on lower half, this brown shade 
extending upward to costa at extreme base and obliquely 
upward from lower outer angle of cell to costa at apex; 
an oblique blackish brown band: from: costa at about 
one-third extends to the paler brown ground color on 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 87 


lower half of wing, dividing the white costal area into 
two strongly contrasted white patches; dark basal area 
also blackish towards costa; a similar blackish shade 
from apex extends obliquely inward towards lower outer 
angle of cell; some blackish shading along the outer 
veins and a narrowly elongate, blackish brown spot on 
midcosta; subterminal line very faint, except at costa, 
dull white, some faint whitish dusting in the outer 
brown area just below apex; a faint peppering of red 
scales on the white areas and more or less over the 
blackish brown markings on costal half of wing; discal 
dots blackish brown, separated and usually distinct; a 
row of black dots along termen. Hind wing semitrans- 
parent, whitish with a smoky tint towards apex and 
along costa and termen; the veins faintly darkened and 
a blackish line along terminal margin. Eighth abdomi- 
nal tufts more or less swollen hairlike scales. Alar 
expanse, 22-26 mm. 

Male genitalia with transtilla absent; harpe with 
sacculus partially divided (towards its apex); hair tufts 
from extended basal arm from sacculus, yellow; penis 
bearing a patch of fine scobinations; vinculum con- 
stricted near middle. 

Antenna of male (fig. 279) with a broad sinus in base 
of shaft occupying a half-dozen fused segments; the 
sinus with many minute, papillalike setae (or scales) on 
its inner surface, but without hair or scale tuft; a small 
but strongly sclerotized spine from lateral edge of smus 
beyond its base and a similar small spine from apex 
of the sinus. 


Typn LocaLiry: Honda, Colombia (type, @, in BM). 

Foop prant: Unknown. 

DistRisuTIoN: GuatemMaLa: Voleén Santa Maria 
(June, July). Cotomp1a: Honda (Apr.). Braziu: 
Santa Catarina (July). Pert: Oconeque (Carabaya). 

I have seen no specimens from Colombia; but the 
male examples before me from Guatemala, Brazil, and 
Peri agree in antennal characters with the type of 
petersenti as described by Ragonot. Evidently the 
species has a wide distribution in Central and South 
America. 


181. Megarthria squamifera, new species 
Figures 275, 280 


Color and markings as in petersent. Male genitalia 
also similar except lateral arms of gnathos stouter and 
vinculum less constricted. Digitate pocket of eighth 
abdominal sternite not appreciably flattened. Male 
antenna (fig. 280) with a broad, elongate sinus; from 
one lateral edge of sinus a flat brush of long stiff hairs; 
the opposing edge concave for most of its length, the 
concavity ending in a sharp projecting point at each 
end; inner surface of sinus smooth. 

Tyrr tocatity: Mount Pods, Costa Rica (type in 
USNM, 61336). 


Foop pirant: Unknown. 
Described from male type, collected by Schaus and 
Barnes (May). 


182. Megarthria frustrator, new species 


Color markings and male genitalia similar to those of 
petersent. Male antenna like that of squamifera. Dif- 
fers from other males of the genus in having a narrow 
ridge of rough, protruding scales along the costa of 
forewing for more than half its length from base. Alar 
expanse, 20 mm. 

TyPE LocaLiry: Juan Viiias, Costa Rica (type in 
USNM, 61337). 

Foop piant: Unknown. 

Described from male type, collected by Schaus and 
Barnes (Feb.). 


183. Megarthria schausi, new species 
FicurE 276 


Color and markings as in peterseni. Male genitalia 
with vinculum not constricted ; apical process of gnathos 
terminating in a weak, short spine. Transtilla present 
developed as a square, sclerotized plate, pendant from 
protruding lobes from the costobasal area of the harpes 
and with thin short projecting arms from its lower 
(anterior) corners. Hair tufts from projecting basal 
arm of sacculus short, yellow. Male antenna like that 
of squamifera. Alar expanse, 18 mm. 

TypE Locauity: Juan Vifias, Costa Rica (type in 
USNM, 61338). 

Foop pirant: Unknown. 

Described from male type, collected by Schaus (Jan.) 
and named in memory of him. The species is easily 
identified by its platelike transtilla. I have seen nothing 
resembling this structure in any other male of the genus. 


184, Megarthria cervicalis Dyar 
Ficures 277, 281 


Megarthria cervicalis Dyar, Ins. Insc. Menstr., vol. 7, p. 42, 1919. 


Forewing as on petersent except that dark areas are 
paler; less blackish brown and with more red scaling 
on the dark antemedial band and midcostal spot, the 
latter almost entirely reddish. Eighth abdominal tuft 
small, consisting of broadly flattened and contorted 
scales; digitate pocket from sternite of eighth segment 
strongly bent and decidedly flattened. Alar expanse, 
20 mm. 

Male genitalia with vinculum not constricted, of even 
width throughout and with terminal margin evenly 
rounded; penis armed with a small, flat, bladelike cor- 
nutus; sacculus of harpe not divided at apex; hair tuft 
from projecting basal arm of sacculus, black. 

Male antenna (fig. 281) with a short, shallow sinus 
at base of shaft and with a very small scale tuft from 
the base of the sinus (under the lower magnification 
looks like a small triangulate spine). 

Tyrr tocaLity: Taénamo, Cuba (Aug.; type in 
USNM). 

Foop piant: Unknown. 

Known only from its male type. <A distinct species 
easily identified by its male antennae and genitalia. 


88 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


185. Megarthria alpha, new species 
Ficure 760 


Female. Color and markings as in peterseni males. 
Alar expanse, 17-26 mm. 

Genitalia with signum developed as a teardrop 
shaped, finely granulate-scobinate, depressed patch: 
ductus bursae shorter than bursa with a narrow scle- 
rotized band near genital opening. LHighth-segment 
collar with a thin apron projecting anteriorly (the 
shape and size of this individually variable; identical 
in no two specimens; compare figures 760a and 760b). 


Typr LocaLity: Volcén Santa Maria, Guatemala 
(type in USNM, 61339). 

Foop prant: Unknown. 

Described from type (Sept.) and three paratypes 
from the type locality (Schaus and Barnes, collectors, 
July, Oct.) and one paratype from each of the following 
localities: Purulhé, Guatemala (Schaus and Barnes, 
July); Quirigué, Guatemala (Schaus and Barnes, Apr.), 
Jalapa, México (Schaus); Orizaba, México (R. Miiller, 
Mar. 13); Mount Pods, Costa Rica (Schaus, May); 
Juan Vinas, Costa Rica (Schaus and Barnes, Jan.); Rio 
Trinidad, Panamé (A. Busck, Mar. 1912); Incachaca, 
Cochabamba, Bolivia (J. Steinbach); Santa Catarina, 
Brazil (F. Hoffmann, July 12, 1935). All females; 
genitalic preparations made of all specimens. 

I believe that alpha will eventually prove to be the 
female of petersent, but have no convincing evidence at 
this time that it is so. 


186. Megarthria beta, new species 
Figure 759 


Superficially indistinguishable from alpha but with 
quite different genitalia. Alar expanse, 17-23 mm. 

Bursa without trace of signum; ductus bursae much 
longer than bursa, unsclerotized at genital opening ex- 
cept for a narrow, very weakly sclerotized band on 
lower margin ; anterodorsal projection of eighth-segment 
collar slight. 

TYPE LOCALITY: Orizaba, México (type in USNM, 
61340). 

Foop pirant: Unknown. 

Described from type (Schaus Collector, no data); one 
paratype from Jalapa, México (Schaus, no data); three 
paratypes from Cayuga, Guatemala (Schaus and 
Barnes, Feb., May, Oct.); one paratype from San José, 
Costa Rica (H. Schmidt); two paratypes from Porto 
Bello, Panamé (A. Busck, May 1912); and one para- 
type from Caparo, west-central Trinidad (F. Birch, no 
data). All females; the paratypes from Trinidad and 
Costa Rica in the Js anse Collection. 


46. Genus Drescoma Dyar 


Drescoma Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 328, 1914. 
(Type of genus: Drescoma cyrdipsa Dyar.) 
Tongue well developed.. Antenna pubescent; basal 
segment normal (ot swollen or elongated); shaft of 
male simple. Labial palpus upturned, reaching vertex; 


moderately slender, the second segment somewhat 
broadly scaled; third segment nearly as long as second, 
acuminate. Maxillary palpus subsquamous (scaling 
of second segment somewhat expanded); folded across 
base of tongue. Forewing smooth; 11 veins; vein 2 
from near lower, outer angle of cell; 3 from the angle 
about equidistant from 2 and 4; 4 and 5 slightly sepa- 
rated at base, thence divergent; 6 from below (but near) 
upper angle of cell, straight; 8 and 9 long stalked; 10 
from the cell approximate to the stalk of 8-9 for some 
distance; male without costal fold but with a distinct 
notch in costa beyond base. Hind wing with vein 2 
from lower outer angle of cell; 3 apparently from the 
stalk of 4-5, but actually contiguous or weakly anas- 
tomosed with it for half its length; 4 and 5 stalked for 
over half their lengths; 7-8 contiguous or partially 
anastomosed beyond cell, the free parts of the veins 
very long; cell very short, one-fifth the length of wing 
or less; discocellular vein oblique; on male upper vein 
of cell notched just beyond base and with some modi- 
fied sex-scaling above and beyond the notch; also on 
male a short fold on anal margin enclosing a hair pencil. 
Kighth abdominal segment without tuft; sternite de- 
veloped as a short sclerotized, digitate pocket. 

Male genitalia as in Megarthria except: Transtilla 
present and in the form of a wide, very shallow U; 
harpe with an appressed, clasperlike projection near 
apex of sacculus; vinculum terminating in a strongly 
sclerotized, sinuate, transverse bar. 

Female genitalia with signum developed as a long, 
strong, curved hook; ductus bursae shorter than bursa; 
genital opening simple; ductus seminalis from bursa 
near junction of bursa and ductus bursae. 

A distinct genus closely related to Ceracanthia and . 
Megarthria and to Drescomopsis in group II; but easily 
distinguished by the bent (notched) upper vein of cell 
in the male hind wing, the very short hind wing cell 
in both sexes, the notched forewing of the male, the 
peculiarly developed transtilla and terminal margin of 


vinculum. 
187. Drescoma cyrdipsa Dyar 


Figures 15, 282, 758 
Drescoma cyrdipsa Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 328, 
1914. 


Color and markings as in Megarthria peterseni except 
for darker hind wings, especially those of the females 
which are dark smoky fuscous. The average size is 
also consistently smaller. 

Hind wing of male with notched projection of upper 
vein of cell into the cell deep and wide; on underside of 
wing a patch of semimetallic scales above the notch; 
a black patch preceding it, and following it a line of 
yellow and black scales along the vein. 

Male genitalia with transtilla triangularly broadened 
toward harpes; clasperlike projection from sacculus 
rather broadly triangulate. Female genitalia with 
bursa minutely granulate; coarser granulations in ductus 
bursae at its junction with bursa. ‘The long, strong, 
thornlike cornutus may be a specific character also, 
but is more probably generic. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 89 


Type tocauity: La Chorrera, Panamaé (type in 
USNM). 

Foop piant: Unknown. 

Disrrisution: México: Chiapas (May). GuatE- 
MALA: Cayuga (Jan., May, June, Aug.). Panam: 
Cabima (May), Corozal (Feb., Nov.), La Chorrera 
(Apr., May), Rio Trinidad (Mar., June), Tabernilla. 
Frenca Guiana: St. Jean Maroni. 


188. Drescoma cinilixa Dyar 
Figure 283 


Drescoma cinilixa Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 329, 
1914. 


Markings and color of forewing as in cyrdipsa except 
dark ground color and dark costal markings paler, 
more ocherous fuscous than brown. Underside of male 
hind wing with a patch of black sex-scaling above the 
notch in cell, orange-yellow sex-scaling on the upper 
vein of cell preceding the notch, on several veins follow- 
ing the notch, and in the median fold at base of wing; 
the indentation of upper vein of cell also much shallower 
than on cyrdipsa, a concavity rather than a strongly 
triangular notch. Hind wing paler; dull smoky white, 
darkening towards termen and apex. Alar expanse, 
15-16 mm. 

Male genitalia with transtilla narrowed towards 
harpes; clasperlike projection from sacculus slender, 
sharply pointed. Female unknown. 

Typn tocauiry: La Chorrera, Panamé (type in 
USNM). 

Foop pirant: Unknown. 

DistrisuTion: GuaTEMALA: Cayuga (May). Pa- 
naMA: La Chorrera (May). 

Distinguishable from cyrdipsa by its paler color, the 
secondary sexual characters of the male hind wing, and 
the differently shaped transtilla and clasper. 


Genus 47. Monoptilota 


[Venational division B. Forewing with veins 4 and 5 closely ap- 
proximate for a short distance from cell. Hind wing with vein 2 
from well before angle of cell; 3 from the angle; cell one-third the 
length of wing. Male genitalia with uncus bifid, its divided ele- 
ments widely separated; transtilla incomplete; apical process of 
gnathos a broadly triangulate hook. Male antenna unipectinate 
with sinus and scale tuft in base of shaft.] 


47. Genus Monoptilota Hulst 


Monoptilota Hulst, Canadian Ent., vol. 32, p. 13, 1900.—Forbes, 
Cornell Mem. 68, p. 621, 1923. (Type of genus: Monoptilota 
nubilella Hulst.) 

Tongue well developed. Antenna of male (figs. 
285g—-h) with basal segment elongate, shaft unipectinate 
and with a long shallow sinus at base containing a stout, 
appressed scale tuft; of female simple, smooth. Labial 
palpus obliquely upturned, reaching to vertex; second 
segment broadly scaled; third segment small, acumi- 
nate. Maxillary palpussquamous. Forewing smooth; 
11 veins; vein 2 from well before lower outer angle of 
cell; 3 from the angle, much closer at base to 4 than to 

3003293—56——7 


2; 4-5 closely approximate for a short distance from 
cell; 6 from below upper angle of cell, straight; 8 and 9 
stalked for less than half their lengths; 10 from the cell; 
male without costal fold. Hind wing with vein 2 from 
well before lower outer angle of cell; 3 from the angle, 
connate with or very closely approximate at base to 
stalk of 4-5; 4 and 5 stalked for about half their lengths; 
7 and 8 closely approximate for a short distance from 
cell; cell one-third the length of wing; discocellular vein 
curved. Eighth abdominal segment of male with a 
pair of fine, weak, hair tufts. 

Male genitalia with uncus bifid, the divided parts 
widely separated and hooked at their apices. Gnathos 
terminating in a triangulate, sharply hooked central 
process, its supporting lateral arms strongly arched and 
arising well down from ventrolateral projections of 
tegumen; an elaborate well-sclerotized subanal plate 
attached to alimentary tube. Transtilla incomplete, 
consisting of two, widely spaced, slender, weakly sclero- 
tized plates. Anellus a simple, moderately broad, par- 
tially curved band. Aedeagus short, stout; penis un- 
armed. Vinculum short (shorter than broad) ; terminal 
margin truncate. 

Female genitalia with signum developed as a spined 
plate (individually variable); bursa otherwise smooth; 
ductus bursae rather stout, expanding gradually into 
the bursa, smooth except for a broad sclerotized banding 
near genital opening; ductus seminalis from bursa near 
junction of bursa and ductus bursae. 

A unique genus of doubtful affinities. 
the one North American species. 


Contains but 


189. Monoptilota pergratialis (Hulst) 
Ficurss 27, 285, 763, 764 


Nephopteryx pergratialis Hulst, Trans. Amer. Ent. Soc., vol. 13, 
p. 162, 1886; Phycitidae of N. Amer., p. 143, 1890.—Rago- 
not, Monograph, pt. 1, p. 267, 1893. 

Nephopteryx grotella Ragonot, N. Amer. Phycitidae, p. 6, 1887. 

Monoptilota nubilella Hulst, Canadian Ent., vol. 32, p. 14, 1900.— 
Chittenden, U. S. Dep. Agr. Div. Ent. Bull. 23, pp. 9-17, 
1900.—Welden, Jour. Econ. Ent., vol. 1, p. 148, 1908.— 
Forbes, Cornell Mem. 68, p. 621, 1923. 

Monoptilota pergratialis (Hulst) Barnes and McDunnough, Con- 
tributions, vol. 3, p. 195, 1916.—Brannon, Journ. Econ. 
Ent., vol. 27, p. 719, 1984.—Brimley, Insects of North Caro- 
lina, p. 300, 1988.—McDunnough, Check list, No. 6141, 
1939. 

Forewing dark fuscous (gray-brown) with blackish 
shading on many of the veins and on costal half of wing; 
along terminal margin a heavy dusting of white between 
the veins, giving much of wing an ashy appearance; 
antemedial line obscure, indicated chiefly by a diffused 
blackish brown outer border, broadest and strongest 
from costa to lower margin of cell, frequently inter- 
rupted or obscured in the ground color towards inner 
margin; subterminal line sinuate, grayish white with 
dentate blackish brown inner and outer borders; discal 
dots distinct, separated, blackish; a black line along 
terminal margin more or less interrupted by white 
streaklets at the vein ends. Hind wing of male semi- 
hyaline white with veins slightly darkened and a black- 
ish brown line along terminal margin; of female much 


90 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


darker, dull smoky white to brown, with veins and ter- 
minal edge correspondingly darker. Alar expanse, 
17-30 mm. 

Male genitalia with characters as given for the genus. 
Female genitalia with signum extremely variable, rang- 
ing from a small, weakly spined plate, like that of the 
paratype (fig. 764), to a large plate with rather long 
slender spines (fig. 763). In one large female of a reared 
series from Virginia the signum is completely absent. 
This variability in female structure is not matched by 
anything in the male, where the genitalia are remark- 
ably uniform for large and small specimens alike. 

Typr Locatity: Florida (pergratialis, 9, m AMNH, 
ex Rutgers; grotella, 9, in Paris Mus.); Auburn, Ala. 
(nubilella, &', in USNM). 

Foop PLANT: Limabean (larva a borer in the stems). 

Distrisution: Maryland, Cabin John Bridge (Aug., 
Sept.), Salisbury (Sept.); Virginia, Norfolk (May), St. 
Elmo (Jan., Feb., Mar.); North Carolina, Vance County 
(Aug., Oct.); South Carolina, Florence (June, July), 
Lyna Plantation (June); Georgia, Savannah (June); 
Florida, Coconut Grove (Apr., May), Miami (Apr.); 
Alabama, Auburn (July), Montgomery (July); Arizona, 
Baboquivari Mts. (June, Aug., Sept.), Huachuca Mts. 
(uly, Aug.), Nogales (uly), Palmerlee, Washington 
Mts., White Mts. (June). 

The species is of some importance as an enemy of 
limabeans and is known in economic literature as the 
‘dimabeam vine borer.”’ The Chittenden (1900) paper 
cited gives what is known of the life history. There are 
several later references in publications devoted to eco- 
nomic entomology but they add nothing to our knowl- 
edge of the insect. It is not known outside of the 
United States; at least no specimens have been received 
or identified from any of the tropical American regions 
where the species might be expected to occur; and in 
the United States its distribution seems to be limited 
to the eastern area from the District of Columbia south 
to Florida and adjacent Gulf States and to southern 
Arizona. The only known host is the limabean. The 
southern Arizona distribution raises a question as to 
another possible host (probably a wild legume), for the 
Arizona, localities are mostly out of the range of lima- 
bean cultivation. 


Genera 48-50: Zamagiria to Magariopsis 


[Venational division D. Forewing with veins 4 and 5 connate, 
shortly stalked or closely approximate for a short distance from 
base; 8 close to 4-5 at base; 6 bent towards base connate with or 
shortly separated from the stalk of 8-9 at base. Hind wing with 
4 and 5 anastomosed from just beyond angle of cell for about 
half their lengths; cell short. Eighth abdominal segment of 
male with compound, ventral tufts. Labial palpi of male up- 
curved; appressed to face or to each other; third segment greatly 
reduced, acuminate. Gnathos with apical process broadly pro- 
duced and lateral arms elongate and arising from ventrolateral 
projection from tegumen. Female with ductus bursae short and 
5 sclerotized towards genital opening and junction with 
bursa. 


48. Genus Zamagiria Dyar 


Zamagiria Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 329, 1914. 
(Type of genus: Zamagiria dixolophella Dyar.) 

Tongue well developed. Antenna of male shortly 
ciliate, a deep sinus containing heavy scale tuft at base 
of shaft; of female simple and weakly pubescent. Male 
head between the antennae deeply grooved to hold an 
appressed, matted tuft of long scales arising from the 
upper edge of frons. Labial palpus recurved-ascending; 
second segment very long; broadly dilated and hol- 
lowed within to hold the maxillary palpus; third seg- 
ment short, acute. In repose the labial palpi are 
appressed to each other and fit into the groove on head 
covering both the maxillary palpi and the scale tuft 
from frons. Maxillary palpus of male in the form of a 
large aigrette; of the female minute, filiform. Forewing 
with a ridge of roughened scales preceding the ante- 
medial line but not reaching to costa; 11 veins; vein 2 
from before the lower outer angle of cell; 3 from the 
angle, close to 4—5 at base; 4 and 5 very closely approxi- 
mate for a short distance from cell; 6 from upper angle 
of cell, bent towards base, connate with 8-9; 8 and 9 
long stalked; 10 from the cell, approximate to the stalk 
of 8-9 for a short distance from cell; male without costal 
fold but rough scaled on underside of costa at base. 
Hind wing with vein 2 from well before the angle of the 
cell; 3 from the angle; 4 and 5 anastomosed from just 
beyond the angle for about half their lengths; 7 and 8 
contiguous or weakly anastomosed for a short distance 
beyond cell; cell less than one-fourth the length of wing; 
discocellular vein curved. Highth abdominal segment 
of male with compound ventral scale tufts. 

Male genitalia with uncus triangulate. Gnathos with 
apical process broadly produced and variously modified ; 
lateral arms elongate and arising well down from ventro- 
lateral projections of tegumen. Transtilla absent. 
Harpe with sacculus more or less produced at apex; 
clasper in some form usually present; cucullus narrow 
and reduced. Aedeagus stout; penis armed with strong 
spine or spines (except in hospitabilis). 

Female genitalia with bursa armed with strong spine 
cluster or clusters and usually partially sclerotized; 
ductus bursae short, strongly sclerotized towards genital 
opening and junction of bursa and ductus bursa (the 
sclerotizations more or less contorted) ; genital opening 
broad; ductus seminalis from bursa towards junction 
of bursa and ductus bursae. 

An easily recognized, compact genus exhibiting strik- 
ing structural specific differences in genitalia. 


190. Zamagiria dixolophella Dyar 
Figure 286 


Zamagiria dixolophella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 
329, 1914. 


Aigrette of male maxillary palpus reddish. 
Forewing dark smoky gray, the basal area below costa 
a trifle paler; antemedial line faint, narrow, oblique and 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 91 


sinuate, well out on wing, bordered outwardly by a 
narrow blackish line, inwardly (from top of cell to inner 
margin by a broad salmon-ocherous patch filling the 
space between antemedian line and the vertical scale 
ridge; the latter thin, blackish, bordered inwardly by a 
white line; subterminal line near outer margin, indis- 
tinct, pale, bordered inwardly by some obscure blackish 
streaklets on the veins; discal dots separated; all the 
markings obscure except the white inner margin of the 
subbasal scale ridge. Hind wing translucent, pale 
smoky fuscous; the veins and terminal margin faintly 
darkened. Alar expanse, 21 mm. 

Male genitalia with flaring apical process of gnathos 
terminating in a shortly forked hook. Harpe with a 
broadly triangulate projection (clasper) from near apex 
of sacculus, the sacculus otherwise not appreciably 
produced at apex. Penis armed with three clusters of 
straight, strong spines. 

TypE Locauity: Corozal, Canal Zone, Panamd (Nov.; 
type in USNM). 

Foop prant: Unknown. 

Represented only by the male type. 


191. Zamagiria pogerythrus Dyar 
Fiaures 49, 287, 765 
Zamagiria pogerythrus Dyar, Ins. Insc. Menstr., vol. 7, p. 47, 
1919. 


Aigrette of male maxillary palpus red. 

Forewing pale brownish gray dusted and blotched 
with white, slightly darker along costa; the white scaling 
most conspicuous about the lower discal spot, as a more 
or less diffused blotch in median area following the 
antemedial line, as an interrupted white line preceding 
the subbasal scale ridge, and as a short, narrow shade 
from apex in terminal area; antemedial pale line faint, 
its outer blackish border interrupted on the males, 
continuous on females; the subbasal scale ridge broken 
into two or three patches of black intermixed with some 
red raised scales; the patch between these and the ante- 
medial line more restricted and fainter than on dixo- 
lophella, ocherous fuscous; subterminal line well con- 
trasted, indented at vein 6, white, margined inwardly by 
blackish streakings on the veins and inwardly and out- 
wardly by dark spots on costa; discal dots separate, 
black, the lower one elongately enlarged and the most 
conspicuous black marking on the wing; a row of 4 or 
5 black dots along terminal margin. Hind wing whit- 
ish, translucent; a dark shade along costa and a narrow 
dark line along termen; the veins very faintly darkened. 
Alar expanse, 20-22 mm. 

Male genitalia with apical process of gnathos con- 
torted. Harpe with apex of sacculus shortly and 
bluntly produced at apex. Penis armed with two 
clusters of straight, stout, moderately long spines. 

Female genitalia with signum developed as a narrow, 
elongate, strongly spined plate extending the length of 
the bursa; bursa otherwise unsclerotized and minutely 
spinose; ductus bursae very broad (as broad as long and 


broader than bursa), its membrane thickened and 
bearing on its inner dorsal surface, near genital opening, 
a large pair of conjoined sclerotized plates, and on its 
inner ventral surface, near junction of bursa and ductus, 
a large, thickened, corrugate, triangulate, sclerotized 
plate bearing minute spines over its inner surface. 
Eighth-segment collar ventrally fused. 

Type tocauity: Chejel, Guatemala (type @, in 
USNM). 

Foop prant: Unknown. 

Distrisution: Méx1co: Campeche (July). 
MALA: Chejel (June, Aug.), Purulhé (July). 


GUATE- 


192. Zamagiria hospitabilis Dyar 
Ficure 288 
Zamagiria hospitabilis Dyar, Ins. Insc. Menstr., vol. 7, p. 48, 
1919. 


Aigrette of maxillary palpus pale red. 

Forewing ocherous brown heavily overlaid with black, 
giving the wing a dark, blackish brown ground color; 
two strongly contrasted white markings, the inner 
border of the broken, black, subbasal scale ridge and a 
large, irregular blotch on lower median area just beyond 
the antemedial line; antemedial line not strongly con- 
trasted but distinguishable throughout, sinuate, whit- 
ish, bordered outwardly by a thin black line; subtermi- 
nal line more contrasted, dull white, indented at vein 6 
and at lower fold; discal dots confluent, forming a black 
lunate mark on discocellular vein; outer area beyond 
subterminal line very faintly dusted with white; ter- 
minal dots confluent forming a narrow black line along 
outer margin of wing. Hind wing semihyaline white 
with a dark shade along costa; the veins and terminal 
margin faintly darkened. Alar expanse, 21 mm. 

Male genitalia with apical process of gnathos broadly 
flaring and serrate. Harpe with sacculus shortly pro- 
duced at apex, very long and with broadly triangulate, 
median, clasperlike projection (similar to that of 
dizolophella). Lateral margins of aedeagus armed with 
short, stout spines; penis unarmed except for a small, 
flat, weakly sclerotized plate. 

Typx Locauity: Ténamo, Cuba (type in USNM). 

Foop pirant: Unknown. 

Represented only by the male type. 


193. Zamagiria masculinus Dyar 
FicurE 289 
Zamagiria masculinus Dyar, Ins. Inse. Menstr., vol. 7, p. 46, 
1919. 
Aigrette of male maxillary palpus whitish ocherous. 
Forewing very pale ocherous gray thinly dusted with 
white in median area; the ocherous tint strongest in the 
patch between scale ridge and antemedial line, along 
the costal edge, and broadly along lower fold; ante- 
medial and subterminal lines distinguishable, but not 
strongly accented, whitish; black scaling of subbasal 
scale ridge limited to one or two dots; blackish outer 


92 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


border of antemedial line interrupted; inner blackish 
border of subterminal line broken into short black 
streaks on the veins; discal dots separated, blackish; a 
row of small blackish dots (5 or 6) along termen between 
the vein ends; on inner margin at lower outer edge of 
antemedial line a small white spot; none of the white 
markings strongly contrasted; the usual black markings 
broken into dots and very short dashes; overall shade a 
pale clay color. Hind wing translucent, white with a 
faint ocherous tint; a narrow pale brownish line along 
termen; the veins very faintly darkened. Alar expanse, 
25 mm. 

Male genitalia with gnathos terminating in a broad, 
inverted, heart-shaped apical projection. Harpe with 
sacculus short, sharply pointed and shortly projecting 
at apex. Penis armed with a single stout spine about 
one-third as long as aedeagus. 


TypEr Locality: Cayuga, Guatemala (Apr.; type in 
USNM). 


Foop pLrant: Unknown. 


The largest known species of Zamagiria. 
only from the male type. 


Known 


194. Zamagiria australella (Hulst), new combination 
Figures 292, 766 


Selagia australella Hulst, Canadian Ent., vol. 32, p. 174, 1900.— 
McDunnough, Check list, No. 6232, 1939. 

Immyrla bumeliella Barnes and McDunnough, Contributions, 
vol. 2, p. 182, 1913.—McDunnough, Check list, No. 6188, 
1939. (New synonymy.) 

Aigrette of male maxillary palpus ocherous white. 

Forewing white dusted with black scales, giving the 
wing a pale gray color, lightest in basal and through the 
median areas, slightly darker along costa; subbasal scale 
ridge conspicuous and normally unbroken, black with a 
fine white inner border; the patch between scale ridge 
and antemedial line olivaceous ocherous; antemedial 
line oblique, dentate-sinuate, faint, indicated chiefly by 
its black outer bordering line; subterminal line dentate- 
sinuate, bordered inwardly and outwardly at costa by 
blackish dashes, outwardly below costa by a narrow 
brownish shade and inwardly by a fine black line; discal 
dots distinct, separate, black; a small blackish or 
brownish spot on inner margin a slight distance beyond 
antemedial line and diffused black smudges on veins 2, 
3 and 4 for a short distance from cell. Hind wing 
semihyaline white; a darker line along termen and the 
veins very slightly darkened. Alar expanse, 15-18 mm. 

Male genitalia with apical process of gnathos a nearly 
square plate. Harpe. with costa narrowly sclerotized 
and shortly produced at’ apex; sacculus shortly and 
bluntly produced. Penis armed with a single stout 
cornutus about half as long as aedeagus. 

Female genitalia with bursa sclerotized and densely 
spined at posterior end, the sclerotization extending 
into ductus bursae; occupying most of the remainder of 
ductus bursae a funnel-shaped, convoluted, sclerotized 
plate. 


Typr LocaALitizs: Blanco County, Tex. (australella, 


in AMNH, ex Rutgers); Fort Myers, Fla. (6umeliella 
in USNM). 

Foop piant: Bumelia microcarpa. 

Distrisution: Texas, Blanco County; Florida, Fort 
Myers (May), Miami. 

The genitalia of the female type of australella are 
identical with those of a reared female paratype of 
bumeliella, and the two moths otherwise agree, so there 
is DO question of the synonymy. However, I doubt 
somewhat the correctness of the locality label on Hulst’s 
type and suspect that it may be a Florida specimen. 


195. Zamagiria fraterna, new species 
Figure 291 


Aigrette of male maxillary palpus white. 

Forewing white on dorsal half and along inner margin 
beyond antemedial line to tornal angle; extreme base of 
costa rough scaled, black (similar black sex-scaling on 
underside of wing at base) ; antemedial line at middle of 
wing incomplete, indicated only by an angulate white 
line on lower half, bordered inwardly and outwardly by 
small black smudges; along lower margin to antemedial 
line the ground color is pale brown; a faint dusting of 
brown or purplish brown scales on the white ground for 
a Darrow margin along costa; scale ridge interrupted, a 
series of black dots with the inner white bordering line 
very faintly indicated; the usual patch between scale 
ridge and antemedial line pale brown, a trifle paler than 
the ground color on lower basal area of wing; some simi- 
larly colored scales in the lower postmedian area be- 
tween the antemedian line and the end of cell and in the 
interspaces of veins 3 to 5; subterminal line incomplete, 
only its middle portion distinguishable; discal dots dis- 
tinct, separated, black, the lower one enlarged; black 
scaling along veins 3, 4 and 5. Hind wing hyaline 
white, a pale brownish gray line along terminal margin 
and a similar shade along costa; at extreme base of wing 
a few black sex-scales on the veins. Alar expanse, 
21.5 mm. 

Male genitalia with apical process of gnathos roundly 
spatulate. Harpe considerably broadened towards 
middle, thence sharply tapered to the narrow cucullus; 
costa strongly humped at middle; sacculus produced at 
apex into a long, strongly sclerotized, curved, free, 
hooklike arm. Anellus a U-shaped plate. Penis armed 
with a single stout spine (about one-fourth as long as 
aedeagus), a strongly sclerotized, corrugated plate and 
a concentration of rather coarse granulations. 

TYPE LOCALITY: Santiago de las Vegas, Cuba (type 
in USNM, 61841). 

Foop piant: ‘Caimitillo.” 

Described from male type reared by A. Otero, Aug. 
17, 1932, and labeled: ‘“‘Leaf tier on Caimitillo, E. E. A. 
Cuba, Ento. No. 10006.” 


196. Zamagiria laidion (Zeller) 
Figures 290, 767 


Myelois laidion Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 211, 


Piesmopoda laidion (Zeller) Ragonot, Monograph, pt. 1, p. 162, 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 93 


Zamagiria laidion (Zeller) Dyar, Ins. Inse. Menstr., vol. 7, p. 46, 
1 


Zamagiria deia Dyar, Ins. Insc. Menstr., vol. 7, p. 46, 1919 (new 
synonymy). 

Zamagiria striella Dyar, Ins. Insc. Menstr., vol. 7, p. 47, 1919 
(new synonymy). 

Aigrette of male maxillary palpus dull white. 

Forewing pale gray, the overall tint shading from 
grayish white to very pale bluish gray (in fresh reared 
examples) ; costal border slightly darker; the patch be- 
tween scale ridge and antemedial line reddish brown; 
similar reddish brown scaling spread over basal area 
bordering inner margin and frequently blotching the 
median area over vein 1b and the lower fold (especially 
on the males); also some scattering of reddish brown 
streaking on veins 8 and 4 for a very short distance from 
cell; black scale ridge more or less complete; on several 
males a black or black and red-brown streak along vein 
1b from scale ridge to base of wing; transverse lines 
faint, the outer blackish border of the antemedial and 
the inner dark border of subterminal more or less inter- 
rupted (more so on males than females); discal dots 
separated, black, the lower one slightly enlarged; a 
row of 5 or 6 small black dots along termen. Hind 
wing semihyaline white; a dark line along terminal mar- 
gin and some faint darkening of the veins. Alar ex- 
panse, 15-22 mm. 

Male genitalia similar to those of fraterna except: 
Costa of harpe convex but not decidedly humped; an- 
nellus inverted;* penis with cornutus a single, very 
stout, long spine (over two-thirds as long as aedeagus) 
surrounded by a cluster of small granulations. 

Female genitalia with a finely spined plate (signum) 
near junction of bursa and ductus bursae, and a strongly 
sclerotized lateral patch in bursa near its anterior end; 
ductus bursae with a sclerotized plate occupying most 
of its length, the posterior end of the plate folded over 
into triangulate ventral lips. 

Types LocaLitres: Honda, Colombia (/aidion, in BM); 
Chejel, Guatemala (deia and striella, in USNM). 

Foop piants: Achras sapota, Mimusops emarginata, 
Eriobotyra japonica (larvae feeding on leaves and 
flowers). 

DisrrisuTion: Unirep States: Florida, Key West 
(Apr., May), Miami (Jan., Dec.). Guatemaua: Chejel 
(June). Panamd: Porto Bello (Feb.). Cotompta: 
Honda. Braziu: Castro (Parand) Obidos (Amazon re- 
gion, Sept.), Vigosa (Minas Geraes, Sept.). Boxrvia: 
“Hast Bolivia” (“Aug.—Oct., T. Steinbach”). 

Hitherto laidion has been recognized only from female 
examples. A reared series in the National Museum 
from Florida has enabled us to associate the sexes and 
has established the synonymy of deia and striella, both 
described from males. Dyar’s type of striella is merely 
an extreme example of a common color variant with 


3 This structure seems more like a transtilla than an anellus; 
for its straight posterior margin lies between the costal bases of 
the harpes, and it could be interpreted as a transtilla or com- 
bination of transtilla and anellus, except that in other species of 
the genus there is no trace of even the vestiges of a true transtilla. 


more or less black streaking on the base of vein 1b of 
forewing. The reared Florida specimens have also 
given us the host records cited above. 


197. Zamagiria ipsetona Dyar 
Fiaure 768 


Zamagiria ipsetona Dyar, Ins. Insc. Menstr., vol. 7, p. 47, 1919. 


Forewing gray, more heavily marked with black than 
in preceding species; the blackish streakings on the 
veins (2 to 6 in this species) longer and stronger; lower 
discal spot elongated into a black dash; dark dashes 
(reddish brown) on the veins in outer area following the 
faint subterminal line; the black scale ridge not con- 
tinuous; antemedial line sinuate and nearly vertical; 
the whitish areas limited to a pale oval patch surround- 
ing the black-streaked veins and discal dots and a faint 
shade preceding the black scale ridge; the usual red- 
brown scaling limited to the patch between scale ridge 
and antemedial line and weak shadings between some 
of the veins in postmedian area. Hind wing translu- 
cent white; a narrow dark shade along costa and a 
blackish line along termen; veins appreciably darkened. 
Alar expanse, 23-24 mm. 

Female genitalia with two elongate spined plates in 
bursa; another more weakly spined plate at junction of 
bursa and ductus bursae; from the junction a scleroti- 
zation extends along one side of bursa for about four- 
fifths of its length; ductus bursae very short, strongly 
sclerotized towards genital opening, the plate folded 
over at the opening into ventrolateral lips. Eighth- 
segment collar completely fused ventrally. 


TYPE LOCALITY: Juan Vifias, Costa Rica (type in 
USNM). 


Foop pLant: Unknown. 


Represented by three females from the type locality 
(Feb., June). The male is unknown. 


49. Genus Anegcephalesis Dyar 


Anegcephalesis Dyar, Ins. Insc. Menstr., vol. 5, p. 46, 1917.— 
Heinrich, Ins. Insc. Menstr., vol. 5, p. 48, 1917 (larva). 
(Type of genus: Anegcephalesis cathaeretes Dyar.) 


Characters of Zamagiria except: Maxillary palpus 
minute, subsquamous in both sexes. Forewing with 
veins 4 and 5 connate or very shortly stalked. Hind 
wing with cell longer (slightly less than one-third the 
length of wing). 

The genus is very close to Zamagiria but its separa- 
tion seems to be justified by the differences in the male 
maxillary palpi and the longer cell of hind wing. The 
venational differences noted above (between veins 4 and 
4 of forewing) may be only a specific character. Dyar 
distinguished Anegcephalesis from Zamagiria on the 
difference in maxillary palpi, but mistook the appressed 
scale tuft from frons for that organ. This scale tuft is 
present and equally well developed in Zamagiria and 
Anegcephalesis. 


94. UNITED STATES NATIONAL MUSEUM BULLETIN 207 


198. Anegcephalesis arctella (Ragonot), new combination 
Figures 50, 294, 770 


Phycita arctella Ragonot, N. Amer. Phycitidae, p. 4, 1887; 
Monograph, pt. 1, p. 180, 1893. 

Anegcephalesis cathaeretes Dyar, Ins. Insc. Menstr., vol. 5, p. 46, 
1917 (new synonymy). 

Anegcephalesis catheretes (Dyar), Ins. Insc. Menstr., vol. 7, p. 
48, 1919 (food plant and altered spelling)—MceDunnough, 
Check list, No. 6176, 1939.—Bruner, Scaramuzza, and 
Otero, Bull. 63, Estaci6n Exp. Agron., Cuba, p. 69, 1945. 

Forewing brownish gray dusted with white; the 
white shading more pronounced (on the males) in the 
median area above the base of the cell, as a small spot 
on inner margin below the dark markings on veins 2 
to 5, and in outer area beyond the dark outer border of 
the subterminal line, on the female the white dusting 
is more generally distributed over median and outer 
areas and, on some specimens, on the basal area, giving 
the wing an ashy gray appearance; subbasal ridge black 
bordered inwardly by a white line and followed by a 
dull olivaceous ocherous patch; antemedial line well 
out towards middle of wing, oblique, notched at vein 1b 
and evenly curved above it, outwardly bordered by a 
continuous black line; subterminal line distinct; in- 
dented just below costa and very slightly so at 1b; 
discal spots fused into a lunate black line along dis- 
cocellular vein; a dark brownish smudge over veins 1 
to 5 adjacent to cell. Hind wings translucent white; 
the veins very faintly darkened. A narrow brown line 
along termen. Alar expanse, 20-23 mm. 

Male genitalia with uncus narrowly triangulate. 
Apical process of gnathos a thin, broadly ovate plate. 
Harpe with appressed, elongate clasper; sacculus not 
produced at apex. Aedeagus sharply bent; penis armed 
with an elongate, flattened, partially bent, platelike 
cornutus, and some sclerotized wrinklings and granu- 
lations. 

Female genitalia with signum developed as an elon- 
gate, strongly spined plate (about half as long as bursa) ; 
opposite the signum a narrower, much more weakly 
sclerotized plate beginning at junction of bursa and 
ductus bursae (this plate about half the length of 
signum) ; bursa, otherwise smooth; ductus bursae tubu- 
lar, sclerotized. Highth-segment collar fused ven- 
trally. 


Typ Locauity: Nassau, Bahamas (arctella, in Paris 
Mus.; cathaeretes, mn USNM). 

Foop piant: Dipholis salicifolia. 

Distrisution: Unitep States: Florida, Miami, 
Paradise Key (Mar.), Royal Palm State Park (Feb.). 
Bawamas: Nassau (New Providence Isl., Feb., Mar., 
April, May, Sept.). Cusa: Baracoa (Oct.), Santiago 
Province (Feb., June, Sept., Oct., Dec.). 

Ragonot’s arctella was described from a single female. 
His description and figure agree in every detail with 
Dyar’s cathaeretes represented by a large series in the 
U. S. National Museum. Both were described from 
the same type locality. There is no doubt that both 
names apply to the same species. 


50. Magiriopsis, new genus 


TyPE oF GENUS: Sematoneura denticosella Dyar. 

Characters of Zamagiria except: Antenna of male 
unipectinate for basal half of shaft, serrate and shortly 
ciliate beyond; shaft with very shallow sinus near base, 
containing a few roughened scales and a short row of 
minute spines but without scale tuft. Labial palpi of 
male broad, dorsoventrally flattened and appressed to 
face (not to each other as in Zamagiria and Anegcephale- 
sis); hollowed inner surface; third segment greatly re- 
duced and completely hidden by scaling of second 
segment. Male head without scale tuft from upper 
edge of frons, not deeply grooved between antennae but 
with an enlarged scale tuft behind antennae. Maxillary 
palpus of female squamous (of male, as in Zamagiria, 
in the form of an aigrette). Forewing smooth; vein 2 
from well before outer angle of cell; 6 slightly bent 
towards base but from below upper angle of cell, sepa- 
rate from stalk of 8-9 at base. Hind wing with cell 
less than one-third the length of wing. 

Male genitalia with complete transtilla. 


199. Magiriopsis denticosella (Dyar), new combination 
Figures 293, 769 


Sematoneura denticosella Dyar, Proc. U. S. Nat. Mus., vol. 42, 
p. 105, 1912. 

Hypsipyla denticosella (Dyar), Ins. Insc. Menstr., vol. 7, p. 41, 
1919. 


Crocidomera cristalis Hampson, Ann. Mag. Nat. Hist., ser. 10, 
vol. 4, p. 352, 1929 (new synonymy). 

Aigrette of male maxillary palpus whitish ocherous. 

Forewing reddish (rust) brown, the costal third lightly 
dusted with white, especially along the costa and be- 
tween the veins, the dark color somewhat accented on 
the veins and, on occasional female specimens, some 
black scaling on the veins before the usual location of 
subterminal line; the latter rarely indicated ; antemedial 
line and discal spots obsolete. Hind wing translucent, 
opalescent with a smoky shade at apex; a dark line along 
termen and the outer veins faintly darkened. Alar 
expanse, 32-40 mm. 

Male genitalia with uncus narrowly triangulate. 
Apical process of gnathos a long, strong spine, sharply 
hooked at apex. Transtilla a weakly sclerotized, shield- 
like bridge attached to costal bases of harpes. Harpe 
with costa strongly sclerotized for basal third of harpe 
and terminating in a broad projection; sacculus with a 
broad projection from base and a broadly projecting 
clasper from apex. Aedeagus long and stout; penis 
armed with three clusters of strong spines (about as 
long as width of aedeagus). Vinculum elongate (two 
and one-half times as long as greatest width), stout, not 
appreciably tapering; posterior margin reinforced and 
squarely excised. 

Female genitalia with two irregularly shaped and 
strongly spined plates in bursa, and bursa at junction 
with ductus bursa strongly sclerotized, the wrinkled 
sclerotization extending part way into the ductus; 
ductus bursae otherwise only sclerotized (weakly) at 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 95 


genital opening. Highth-segment collar not fused 
ventrally. 

TypPE LocaLitiEs: Orizaba, México (denticosella, in 
USNM); Juan Vifias, Costa Rica (cristalis, in BM). 

Foop piant: Unknown. 

Distrisution: México: Orizaba (Oct.), Misantla 


(Aug.). Guatemata: Cayuga (Jan., Aug.), Chejel 
(June). Costa Rica: Juan Vifias (Jan., Nov.). Co- 
LomBiA: Upper Rio Negro. VeEnnzuEua: Aroa, San 


Esteban Valley. British Guiana: Omai. Braziu: 
Ponte Nova (Rio Xingu, Amazonas), Santos (Mar.). 
Both denticosella and cristalis were described from 
females. The synonymy is obvious. Both Dyar and 
Hampson had seen males many years before but evi- 
dently overlooked them when writing their descriptions. 


Genus 51: Ancylostomia 


Venational division D. Forewing with veins 4 and 5 contiguous 
or stalked for about one-third their lengths from cell; 3 connate 
with 4 at base; 6 bent towards base, connate with the stalk of 
8-9. Hind wing with veins 4 and 5 stalked to middle; 3 closely 
approximate to the stalk of 4-5 at base; cell short. Labial palpus 
obliquely ascending with third segment long and porrect. Uncus 
pentagonal. Gnathos produced at apex into a pair of long, 
flattened, pointed, contorted, bandlike projections. Female 
with bursa copulatrix weakly sclerotized throughout; eighth- 
segment collar modified ventrally. Eighth abdominal segment 
of male with compound ventral tufts. 


51. Genus Ancylostomia Ragonot 


Ancylostomia Ragonot, Monograph, pt. 1, p. 567, 1893. (Type 


of genus: Myelois stercorea Zeller.) 

Tongue well developed. Antenna of male with a 
sinus and strong scale tuft in shaft at base, shaft other- 
wise flattened, weakly serrate and pubescent; of female, 
simple and pubescent. Labial palpus obliquely ascend- 
ing, with third segment porrect; second segment long, 
reaching well above vertex, on male hollowed to receive 
maxillary palpus; third segment slender, acuminate, on 
male half as long as second, on female about the same 
length as second segment. Maxillary palpus of male 
in the form of an aigrette; of female, squamous. Fore- 
wing smooth; 11 veins; vein 2 from before but near 
lower outer angle of cell; 3 from the angle, connate with 
4; 4 and 5 contiguous or stalked for about one-third 
their lengths from cell; 6 from upper angle of cell, con- 
nate with the stalk of 8-9, bent towards base; 8 and 9 
stalked for slightly more than half their lengths; 10 from 
the cell, approximate for a short distance to the stalk of 
8-9; male without costal fold. Hind wing with vein 
2 from before lower outer angle of cell; 3 from the angle, 
closely approximate to 4 at base; veins 2 and 3 very 
long; 4 and 5 stalked to middle; 7 and 8 anastomosed or 
contiguous for a short distance from cell (for less than 
half their lengths); cell less than one-third (about one- 
fourth) the length of wing; discocellular vein curved. 
Eighth abdominal segment of male with compound, 
ventral scale tufts. 

Male genitalia with uncus pentagonal (the sides 


parallel, the terminal margin angled). Gnathos pro- 
duced at apex into a pair of long, flattened, sharply 
pointed and slightly contorted, bandlike projections; 
the lateral arms short and articulated to base of uncus. 
Transtilla absent. Harpe elongate, slender, cucullus 
reduced, its apex bluntly pointed; sacculus produced 
at apex into a slender, free, curved hook. Penis armed 
with a pair of thin, somewhat flattened and twisted 
spines as long (or nearly as long) as aedeagus. Vincu- 
lum slightly longer than greatest width; terminal margin 
broadly rounded. 

Female genitalia without signum; bursa weakly 
sclerotized throughout; ductus bursa strongly sclero- 
tized except for a narrow space beyond junction of 
bursa, at junction with bursa finely ridged and spined, 
the ridges and spines extending for a short distance into 
bursa, at genital opening the margins of ductus fuse 
into eighth-segment collar. Eighth-segment collar very 
strongly sclerotized; enlarged but not fused ventrally; 
its ventrocaudal angles produced and pointed; from its 
ventrolateral angles a pair of invaginated sclerotized 
pockets. 

The genus is easily identified by its peculiarly de- 
veloped gnathos and the eighth-segment collar of the 
female. Its species are tropical American in distribu- 
tion with (in the case of stercorea) a slight extension of 
range into southern Florida. 


200. Ancylostomia stercorea (Zeller) 
Figures 295, 771 


Myelois stercorea Zeller, Isis von Oken, 1848, p. 873. 

Anerastia ignobilis Butler, Proc. Zool. Soc. London, 1878, p. 494. 

Pempelia diffissella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 
178, 1881. 

Ancylostomia stercorea (Zeller) Ragonot, Monograph, pt. 1, p. 
568, 1893. 


Forewing pale ocherous along the costa, through the 
median fold and over most of the basal area; some 
pinkish brown shading between the veins in the pale 
areas giving the ocherous ground color a rosy tint 
(especially on reared and fresh specimens); the lower 
outer half of wing heavily shaded with brown more or 
less dusted with black (on reared examples this area is 
decidedly blackish brown, its inner margin oblique 
from lower outer angle of cell to basal third of inner 
margin); a distinct brown or black streak under vein 8 
from apex about half-way to cell and a similar longer 
dark streak along vein 6 from terminal margin to the 
cell; a thin whitish line along the lower margin of cell 
from beyond base and continuing into vein 4 for a short 
distance; a similar shorter white line on the subbasal 
half of vein 1b; a small black dot on the middle of the 
white streak on vein 1b and a similar black dot above 
it on the white streak on lower margin of cell; a single, 
large, conspicuous black discal spot at lower outer angle 
of cell; antemedial and subterminal lines obsolete; a 
few black dots or streaklets in outer areas about the 
middle of veins 2, 3, and 4 and on dark specimens some 
blackish or dark brown shading on upper and lower 


96 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


veins of cell at base of wing. Hind wing translucent 
white with a smoky shade along costa, towards apex 
and along termen; the veins slightly darkened. Alar 
expanse 16-28 mm. 

Genitalic characters as given for the genus. 
segment collar of female laterally corrugated. 

TYPE LOCALITIES: Brazil (stercorea, in BM); Jamaica 
(ignobilis, in BM); Honda, Colombia (diffissella, in 
BM). 

Foop pPLant: Cajanus cajan (larvae feeding in the 
pods). 

DistRipution: Unirep States: Florida, Cocanut 
Grove (Apr., May), Goulds (May), Homestead (Apr.), 
Jupiter (Apr.), Miami (May). Cua: Santiago Prov- 
ince (May, June, Oct., Dec.), Santiago de las Vegas 
(May). Harrr (June). Dominican Repusiic (Aug.). 
Pusrto Rico: Isabela (Apr.), Puerto Real (Vieques 
Isl., Apr.). Viremn Istanps: Kingshill (St. Croix; 
June, “Noyv.—Dec.”). Jamatca (Mar.). BAHAMAS: 
Nassau (May). Grenapa: Nevis (Jan.); St. Kitts 
(June). Trinmpap (Mar., May). México: Cuerna- 
vaca (July), Jalapa, Orizaba, Zacualpin (July). Gua- 
TEMALA: Chejel (July, Aug.), Volcin Santa Maria 
(June, July). Costa Rica: Juan Vifias (June.) Pan- 
amA: La Chorrera (May), Rio Trinidad (Mar.). Co- 
Lompia: Honda. Frencw Guiana: Cayenne. Bra- 
ziL: Castro. 

The food plant and Florida records are from a large 
series reared by the Special Survey of the U.S. Bureau 
of Entomology and Plant Quarantine in 1944. Wealso 
have a couple of reared adults and several collected 
larvae from chickpea (Cicer) and black-eyed pea 
(Dolichos). The favored host, however, seems to be 
the pigeonpea (Cajanus). In the pods of that plant 
the larvae are abundant throughout the West Indies 
and in southern Florida. 


HKighth- 


201. Ancylostomia sauciella (Zeller) 
Figure 296 


Pempelia sauciella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 
183, 1881. 

Ancylostomia sauciella (Zeller) Ragonot, Monograph, pt. 1, p. 
569, 1893. 

Forewing cinnamon red mixed with rose; costa paler; 
a narrow subcostal reddish brown shade extending from 
base to apex; the veins somewhat accented by reddish 
scaling; a small blackish discal dot at lower outer angle 
of cell; a similar dot on basal third of vein 1b in a white 
streaklet on the vein; from apex a short oblique blackish 
shade. Hind wing yellowish white, translucent; a thin 
dark line along termen and the veins slightly darkened. 
Alar expanse, 20 mm. 

Male genitalia as in stercorea except uncus narrower 
and apical projecting arms of gnathos shorter and 
decidedly broader. 

TypPE Locauity: Maraquita, Colombia (type in BM). 

Foop prant: Unknown. 

Known only from the type. 


202. Ancylostomia argyrophleps Dyar 


Ancylostomia argyrophleps Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 406, 1914. 

Similar to sauciella except: Costa and lower half of 
basal area carneous white; a broad subcostal band 
extending from base to apex, red-brown shaded with 
black especially towards apex; lower outer area of the 
same color with a smoky fuscous shade along outer half 
of inner margin from near tornus; a white line along 
lower margin of cell and vein 5 from basal third, enclos- 
ing a black dot at lower outer angle of cell; a similar 
black dot on basal third of vein 1b. Hind wing trans- 
lucent, semi-irridescent white with a faint smoky tint, 
the latter more pronounced towards apex; veins dark- 
ened in outer area on females, not appreciably so on 
males. Alar expanse, 20-25 mm. 

Male genitalia as in stercorea except apical processes 
of gnathos a trifle broader (but not so broad as on 
sauciella). Highth-segment collar of female smooth. 
Female genitalis otherwise essentially as in stercorea. 

TyprE Locatiry: Orizaba, México (type, co’, m 
USNM). 

Foop pranr: Unknown. 

Distrisution: México: Orizaba (Aug.), Cuernavaca 
(July). Guatemata: Chejel (July). 


203. Ancylostomia euchroma Dyar 


Ancylostomia euchroma Dyar, Ins. Inse. Menstr., vol. 7, p. 53, 


Forewing below the cell from base to tornus red- 
brown; the upper area white-lined between the veins 
and along the median fold in cell; the veins red-brown; 
a black dot at lower outer angle of cell, one on lower 
margin of cell before its middle and another on basal 
third of vein 1b; a diffused pale shade surrounds this 
last dot and extends obliquely backward to inner 
margin, an oblique line of black dots on veins 2, 3, and 
4; on the female a smoky brown shade from apex 
extending narrowly along costa to base; on male the 
shade from apex is short and cinnamon red, and the 
dark area on lower half of wing is a bright cinnamon 
red. Hind wing in the male translucent white with a 
faint smoky shade at apex; on the female the smoky 
shade extends further back from apex and outer margin 
and the veins in outer area are appreciably darkened. 
Alar expanse, 24-25 mm. 

Male genitalia with the projecting apical bands from 
gnathos as broad as those of sauciella but longer (at 
least as long as those of stercorea). Female genitalia 
with the sclerotized ribbing and spining at junction of 
bursa and ductus bursae slightly stronger than those of 
either stercorea or argyrophleps. Highth-segment collar 
very weakly corrugated. 

Type Locatiry: Castro, Parand, Brazil (type in 
USNM). 

Foop prant: Unknown. 

Represented only by the female type and male 
paratype from the type locality. May be only a race 
of sauciella. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 97 


Genus 52: Caristanius 


{Venational division B. Forewing with veins 4-5 separated at 
base and divergent shortly beyond, smooth; hind wing with vein 
3 approximate to the stalk of 4-5 at base. Highth abdominal 
segment of male with compound tufts. Antenna of male with 
sinus and scale tufts on base of shaft. Labial palpus obliquely 
ascending; second segment of male grooved. Maxillary palpus 
of male in the form of an aigrette. Male genitalia with a pair of 
long, strongly sclerotized arms from ventrolateral margins of 
uncus; transtilla absent; gnathos absent; a single strong cornutus 
on penis. Female genitalia with signa present, developed as 
coarsely spined plates; ductus bursae ribbonlike, sclerotized.] 


52. Caristanius, new genus 


Type or GENUs: Oligochroa pellucidella Ragonot. 


Tongue well developed. Antenna of male with sinus 
and stout scale tuft on base of shaft, finely pubescent; 
of female simple. Labial palpus obliquely ascending; 
on male stout and smooth scaled (the palpi appressed 
to each other), second segment long, reaching well above 
vertex, grooved to hold maxillary palpus, third segment 
very short, hidden in scaling of second; on female slen- 
der, shorter, reaching slightly above vertex, more 
roughly scaled, third segment over half as long as sec- 
ond, acuminate. Maxillary palpus of male in the form 
of an aigrette; of female squamous. Forewing smooth; 
11 veins; vein 2 from before lower outer angle of cell; 
3 from the angle; 4 and 5 shortly separated at base and 
divergent very shortly beyond; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for half or less 
than half their lengths; 10 from the cell; male without 
costal fold. Hind wing with vein 2 from before lower 
outer angle of cell; 3 from the angle, approximate to 
stalk of 4-5 at base; veins 2 and 3 of moderate length; 
4 and 5 stalked for half their lengths; 7 and 8 approxi- 
mate beyond cell (for less than half their lengths) ; cell 
less than half the length of wing (but more than one- 
third); discocellular vein curved. Eighth abdominal 
segment of male with compound ventral scale tuft. 


Male genitalia with uncus short, decidedly broader 
than long; a pair of long, strong, sclerotized arms from 
its ventrolateral angles. Gnathos absent (unless the 
fused arms from uncus can be interpreted as parts of 
this organ, which is extremely doubtful). Transtilla 
absent. Harpe witb costa strongly sclerotized and with 
a strong projection from its base or a strong clasperlike 
projection from below it near base; sacculus short, nar- 
row, weak, bluntly produced at apex; cucullus very nar- 
row, weak and reduced. Anellus a narrow band with a 
central, bifurcate, bandlike projection, flanked by elon- 
gate, lateral lobes. Aedeagus elongate, moderately 
slender, straight; penis armed with a single strong cornu- 
tus and some fine spines and granulations. Vinculum 
stout, as long as or longer than greatest width. 

Female genitalia with signa present, developed as 
elongate, curved, sclerotized, strongly and coarsely 
spined plates; ductus bursae flattened, ribbonlike, 
strongly sclerotized except for a narrow space near 
middle, also granulate towards bursa, the granulations 
extending into bursa; at genital opening ductus bursae 

300329—56——8 


very strongly and broadly sclerotized; ductus seminalis 
from bursa near junction of bursa and ductus bursae. 

The species referred here have been placed in Elasmo- 
palpus. ‘They agree with the type of that genus (ligno- 
sellus) on antennal, palpal, and venational characters; 
but differ strikingly on male and female genitalic struc- 
ture. The characteristic uncus with its long, produced, 
basal arms, the reduced, weak sacculus, the lack of any 
distinguishable gnathos, and the flattened, ribbonlike 
ductus bursae with its strongly sclerotized development 
at genital opening at once distinguish Caristanius from 
Elasmopalpus. 


204. Caristanius pellucidellus (Ragonot), new combination 
Ficures 297, 775 


Oligochroa pellucidella Ragonot, Bull. Soc. Ent. France, 1888, 
p. exl.—Mbéschler, Die Lepidopteren—Fauna von Portorico, 
p. 329, 1890. 

Elasmopalpus pellucidellus (Ragonot), Monograph, pt. 1, p. 429, 
1893. 

Rhodophaea melanoplaga Hampson, in Ragonot, Monograph, pt. 
2, p. 519, 1901 (mew synonymy). 


neo SSS 


oe a 
Ground color of forewing variable, pale brownish gray 
to dark gray with a faint purplish or reddish brown tint; 
transverse markings obsolete on most specimens; on 
some the antemedial line indicated by a narrow, very 
faint, pale line between vein 1b and inner margin, pre- 
ceded by a pale reddish or brownish patch more or less 
overlaid by black scaling (on most specimens before me 
this spot entirely absent); subterminal line distinguish- 
able only on a few specimens, very faint, indicated 
chiefly by some short blackish streaklets on the veins; 
discal spots usually distinct but faint, blackish, sepa- 
rated; a row of black dots along termen. Hind wing 
transparent white with a dark shade along costa and at 
apex and extending as a dark line downward along ter- 
men; on the males this dark line extends only to about 
middle of termen, on females to or nearly to anal angle 
of wing. Alar expanse, 19-25 mm. 


Male genitalia with costa of harpe broadly and stoutly 
sclerotized, a large oval projection from base and its 
apical end folded backward and contorted. Cornutus 
about one-third as long as aedeagus. Vinculum about 
as long as greatest width. 

Female genitalia with granulations of ductus bursae 
extending for a very short distance into bursa. Eighth- 
segment collar not extended to ventral surface; at- 
tached laterally to the extended sclerotization of ductus 
bursae at genital opening; stoutly sclerotized dorsally 
and with an inwardly projecting curved sclerotized 
apron; between this apron and ductus bursae a gland 
of unknown function extends into abdomen, terminat- 
ing ina bulb with thickened membrane. This structure 
not noted on other species of the genus. 

Typx Locatities: Puerto Rico (pellucidellus, in Paris 
Mus.); Sio Paulo, Brazil (melanoplaga, in BM). 

Foop prant: Unknown. 

DistriputTion: Puerto Rico: Puerto Real (Vieques 
Isl., Apr., July), Rio Piedras (Aug., Sept.), San German 
(Apr., Aug.). Sv. Vincent. Jamaica. Surinam: Zan- 


98 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


derij I (Apr.). Braziz: Sio Paulo, “S. H. Brazil” 
[probably Castro]. 


The species is somewhat variable in color and mark- 
ings, but is easily identified by its genitalia. Hampson’s 
melanoplaga was based on large Brazilian females (25 
mm.). All specimens from the West Indies and Suri- 
nam that J have seen are smaller (19 to 21 mm.). How- 
ever, there does not seem to be any reason to keep 
melanoplaga as a racial designation on a mere difference 
in size, as there is nothing else to distinguish the Bra- 
zilian examples. 


205. Caristanius decoloralis (Walker), new combination 


Figures 298, 773 


Trachonitis decoloralis Walker, List, vol. 27, p. 42, 1863. 

Nephopteryx metagrammalis Walker, List, vol. 27, p. 42, 1863. 

Nephopteryx furfurella Hulst, Ent. Amer., vol. 3, p. 131, 1887; 
Phycitidae of N. Amer., p. 143, 1890 (mew synonymy). 

Elasmopalpus decoloralis (Walker) Ragonot, Ent. Amer., vol. 5, 
p. 115, 1889; Monograph, pt. 1, p. 420, 1893.—MceDun- 
nough, Check list, No. 6228, 1939. 

Elasmopalpus floridellus Hulst, Phycitidae of N. Amer., p. 158, 
1890.—Ragonot, Monograph, pt. 1, p. 423, 1893. 

Elasmopalpus decoralis Hulst, Phycitidae of N. Amer., p. 158, 
1890 (misspelling for decoloralis). 

Elasmopalpus decorellus Hulst, in J. B. Smith, List of the Lepi- 
doptera of Boreal America, No. 4341, 1891 (misspelling for 
decoloralis). % 

Elasmopalpus furfurellus (Hulst), Barnes and McDunnough, 
Contributions, vol. 3, p. 196, 1916.—McDunnough, Check 
list, No. 6230, 1939. 

Forewing bluish gray or pale fawn gray with more or 
less white dusting over median area; transverse lines 
indistinct; antemedial line indicated chiefly by its 
blackish outer border broken into dots on vein 1b, lower 
vein of cell and a subcostal spot or short dark streak 
from costa to top of cell; preceding antemedial line a 
reddish patch on inner margin extending to or into cell 
and more or less shaded by black scaling; subterminal 
line sinuate, pale and very faint; lower discal spot at 
end of cell distinct, the upper discal dot much smaller, 
sometimes distinct but frequently obscured; a row of 
black dots along termen. Hind wing translucent, whit- 
ish with a pronounced smoky tint, especially over outer 
half of wing; the veins more or less darkened and a fine 
dark line along termen. Alar expanse, 21-25 mm. 

Male genitalia with projecting arms from uncus 
somewhat curved. Costa of harpe sclerotized for the 
length of the harpe, narrowly sclerotized beyond base; 
a strong, curved, clasperlike projection from below costa 
near base, projected beyond costal edge. Cornutus as 
long as aedeagus. Vinculum considerably longer than 
greatest width. 

Female genitalia with granulations of ductus bursae 
extending deeply into bursa; ductus bursae scobinate 
on lower surface at genital opening. Eighth-segment col- 
lar narrowed dorsally, complete but not fusing ven- 
trally, without sclerotized dorsal apron. 

TYPE LocALItTIEs: United States (decoloralis and me- 
tagrammalis, in BM); Florida (furfurellus and floridellus, 
in AMNH, ex Rutgers). 


Foop piant: Chamaecrista spp. (brachiata, fascicu- 
lata, robusta) larvae feeding on the leaves. These rec- 
ords from rearings by the Special Survey (1944) of the 
Division of Foreign Plant Quarantine of the U. S. 
Bureau of Entomology and Plant Quarantine. 

Distrisution: Florida, Orlando (Feb.), no specific 
locality (Mar.), Fort Myers (May), St. Petersburg 
(June), Stuart (May), Tampa, Vero Beach (Apr., May, 
June, Oct., Dec.); Texas, Brownsville; South Carolina, 
Florence (June), Hiltonhead Isl. (Aug.), Pawleys Beach 
(Sept.). 

This species seems to be confined to the southeast- 
ern United States. In his description of decoloralis 
Walker mentions the protruding arms of the uncus. 
Hulst noted similar structures in his floridellus and asso- 
ciated them with the genitalia, suspecting the synonymy 
of his and Walker’s species. I have before me a photo- 
graph of the genitalia of the type of decoloralis supplied 
by Clarke. They are identical with those of the male 
type of floridellus. The synonymy of floridellus and 
JSurfurellus was established by Barnes and McDunnough 
(1916) and that of metagrammalis with decoloralis by 
Ragonot (1889). 


206. Caristanius guatemalellus (Ragonot), new combination 
FIGURE 774 


Salebria guatemalella Ragonot, Nouv. Gen., p. 18, 1888. 
Laodamia guatemalella (Ragonot), Monograph, pt. 1, p. 414, 
1893. 

This species is apparently known only from the female 
type. Ragonot’s description and figure suggest a large, 
pale brownish, suffused form of pellucidellus; the fore- 
wing pale ocherous brown tinted with reddish, trans- 
verse lines absent, the lower discal spot distinct, and the 
veins sparsely powdered with blackish scales; hind wing 
semitransparent, white faintly tinted with ocherous. 
Alar expanse, 26 mm. 

The genitalia determined the present generic refer- 
ence. The granulations of ductus bursae extend deeply 
into the bursa as in decoloralis; the eighth-segment collar 
has a dorsal, sclerotized, invaginated apron smaller 
than and differently shaped from that of pellucidellus, 
and the collar itself is completely sclerotized ventrally. 


TypE LocaLity: San Geranimo, Guatemala (type in 
BM). 
Foop prant: Unknown. 


Genus 53: Etiella 


[Venational division B. Forewing with veins 4-5 separated and 
divergent from base, a raised-scale ridge beyond base; hind wing 
with vein 3 approximate to the stalk of 4-5 at base. Eighth 
abdominal segment of male with paired tufts. Labial palpus 
porrect, long; second segment of male grooved. Maxillary pal- 
pus in the form of an aigrette. Male genitalia with apical process 
of gnathos a simple, short hook; transtilla incomplete and 
vestigial; harpe with a strong curved arm projecting the length 
of the harpe from base of costa, harpe otherwise weakly sclero- 
tized; two strong cornuti on penis. Female genitalia with signa 
developed as curved, sclerotized bands armed with slender 
spines; ductus bursae short, tubular, sclerotized.] 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 99 


53. Genus Etiella Zeller 


Etiella Zeller, Isis von Oken, 1839, p. 179; 1846, p. 733.—Heine- 
mann, Die Schmetterlinge Deutschlands und der Schweiz, 
Abt. 2, vol. 1, pt. 2, p. 154, 1865.—Meyrick, Proc. Linn. 
Soc. New South Wales, vol. 3, p. 629, 1882.—Hulst, Phy- 
citidae of N. Amer., p. 169, 1890; U. S. Nat. Mus. Bull. 52, 
p. 428, 1902.—Ragonot, Monograph, pt. 1, p. 569, 1893.— 
Spuler, Die Schmetterlinge Europas, vol. 2, p. 208, 1910.— 
Forbes, Cornell Mem. 68, p. 629, 1923.—Janse, Journ. Ent. 
Soc. South Africa, vol. 7, p. 15, 1944. (Type of genus: 
Phycis zinckenella Treitschke.) 

Ramphodes Guénée, Europaeorum Microlepidopterorum index 
methodicus ..., p. 81, 1845. (Type of genus: Phycis 
zinckenella Treitschke.) 


Tongue well developed. Antenna of male with basal 
segment enlarged and bearing a short, bluntly pointed 
projection on inner side near base; shaft with sinus and 
large hair and scale tuft at base, pubescent; antenna of 
female simple. Labial palpus porrect; very long (the 
length of head and thorax); smoothly scaled; second 
segment about five times the length of third on male 
and grooved to hold maxillary palpus; third segment 
short on male, longer on female, acuminate, frequently 
bent downward, especially on female. Maxillary palpus 
of male in the form of an aigrette; of female small, 
squamous. Forewing with ridge of raised scales beyond 
base; 11 veins; vein 2 from before but near lower outer 
angle of cell; 3 from the angle, approximately equi- 
distant from 2 and 4; 4 and 5 shortly separated at base, 
diverging from cell; 6 from below upper angle of cell, 
straight; 8 and 9 stalked for half their lengths; 10 from 
the cell, separated and divergent from the stalk of 8-9; 
male without costal fold. Hind wing with vein 2 from 
well before lower outer angle of cell; 3 from the angle, 
appreciably shorter than 2, closely approximate to the 
stalk of 4-5 at base (separated by a very short vein); 
4 and 5 normally stalked (rarely weakly anastomosed 
or contiguous) to about middle; 7 and 8 closely approxi- 
mate for less than half their lengths from cell; cell about 
one-third the length of wing on male, slightly longer on 
female; discocellular vein curved. Eighth abdominal 
segment of male with a pair of weak ventrolateral hair 
tufts. 

Male genitalia with uncus hoodlike, its apical margin 
evenly rounded. Apical process of gnathos a simple, 
sharp hook. ‘Transtilla incomplete and vestigial (its 
elements rarely distinguishable except under high 
magnification). Harpe with a strongly sclerotized, 
curved, tapering, pointed arm projecting from base of 
costa and as long as costa; remainder of harpe weakly 
sclerotized and abruptly narrowed at middle (the shape 
of harpe probably a specific character). Anellus V- 
shaped; its arms rather broad, blunt and haired. 
Aedeagus moderately long, stout; penis armed with 
two strong cornuti. Vinculum stout; as long as greatest 
width; evenly tapering to a blunt point. 

Female genitalia with bursa elongate, finely scobinate, 
armed with signa consisting of curved, sclerotized bands 
armed with slender spines and situated near junction of 
bursa and ductus bursae;a sclerotized lobe on bursa near 
ductus bursae; ductus bursae short (much shorter than 


bursa), tubular, sclerotized, ribbed and broadened 
towards genital opening; ductus seminalis from bursa 
adjacent to sclerotized lobe. 


A distinct genus, not to be confused with anything 
else and easily identified by its combination of antennal, 
palpal, wing, and genitalic characters. Ragonot, Hulst, 
and Janse list five Walker names in the generic synony- 
my. These supposed genera were based on Old World, 
tropical species which have all been referred as syno- 
nyms of zinckenella. I have no reason to question this 
synonymy but have omitted the references as I have 
not been able to check the genitalia or their types. The 
only species occurring in the New World is zinckenella. 


The larva differs from typical phycitid larvae in that 
it lacks altogether the sclerotized rings about seta IIb 
of mesothorax and seta III of the eighth abdominal 
segment. 


207. Etiella zinckenella (Treitschke) 
Figures 17, 326, 840 


Phycis zinckenella Treitschke, Die Schmetterlinge von Europa, 
vol. 9, pt. 1, p. 201, 1832. 

Phycis etiella Treitschke, Die Schmetterlinge von Europa, vol. 
10, p. 3, p. 174, 1835.—Duponchel, Histoire naturelle des 
lépidoptéres, ou papillons de France, vol. 10, p. 180, 1836.— 
Milliére, Iconographie et description de chenilles et lépidop- 
téres inedits, vol. 1, p. 248, 1861. (Originally proposed as 
new name for zinckenella.) 

Pempelia Etiella zinckenella (Treitschke) Zeller, Isis von Oken, 
1839, p. 179; 1846, p. 755.—Herrich-Schiffer, Systematische 
Bearbeitung der Schmetterlinge von Europa, vol. 4, p.":72, 
1849. 

Ramphodes zinckenella (Treitschke) Guénée, Europaeorum 
Microlepidopterorum index methodicus ..., p. 81, 1845. 

Etiella zinckenella (Treitschke) Heinemann, Die Schmetterlinge 
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 154, 
1865.—Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 177, 
1881.—Hulst, Phycitidae of N. Amer., p. 170, 1890; U. 8S. 
Nat. Mus. Bull. 52, p. 428, 1902.—Ragonot, Monograph, 
pt. 1, p. 572, 1893.—Chittenden, U. 8. Dep. Agr. Bur. Ent. 
Bull. 82 (pt. 3), p. 25, 1909.—Essig, Insects of western 
North America, p. 709, 1926.—Walcott, Journ. Agr. Univ. 
Puerto Rico, vol. 20, no. 1, p. 476, 1936.—McDunnough, 
Check list, No. 6274, 1939. 

Etiella zinckenella schisticolor Zeller, Horae Soc. Ent. Rossicae, 
vol. 16, p. 178, 1881.—Hyslop, U. S. Dep. Agr. Bur. Ent. 
Bull. 95, pt. 6, p. 82, 1912. 

Etiella villosella Hulst, Ent. Amer., vol. 3, p. 133, 1887. 

Etiella schisticolor (Zeller) Ragonot, Ent. Amer., vol. 5, p. 116, 
1889; Monograph, pt. 1, p. 274, 1893.—Hulst, Phycitidae 
of N. Amer., p. 170, 1890. 

Etiella rubribasella Hulst, Phycitidae of N. Amer., p. 170, 1890.— 
Ragonot, Monograph, pt. 1, p. 572, 1893. 


Forewing gray more or less lightened by white 
scaling, especially in median areas; a broad white band 
along costa, extending from base to or nearly to apex; 
extreme costal edge dark gray to red; on occasional 
specimens some red scaling at extreme base of wing and 
in outer median area; transverse lines obsolete; the 
antemedial line replaced by a ridge of raised scales near 
basal third and extending from inner margin to top of 
cell, the raised scales metallic ocherous or orange red 
bordered outwardly by a broad ocherous or orange 
patch; discal spots obsolete. Hind wing whitish with 
a faint smoky tint, to dark smoky fuscous; the veins 


100 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


and terminal margin darker. Alar expanse, 15-28 mm. 

Genitalia as given for the genus. 

TypE LOCALITIES: Sicily (zinckenella, in Hungarian 
Nat. Mus., Budapest; etiella); California (schisticolor, 
in BM); Colorado (villosella, in AMNH, ex Rutgers); 
Florida (rubribasella, in AMNH, ex Rutgers). 

Foop pPiants: Pods and seeds of various Leguminosae 
(Astragalus, Cajanus, Colutea, Crotolaria, Dolichos, 
Glycina, Lupinus, Phaseolus, Pisum, Vicina, Vigna). 

Distrizsution: Throughout the tropical and sub- 
tropical areas of the world and in the warmer temperate 
regions of Hurope, Asia, and North America. The fol- 
lowing American records are from specimens before me. 
Unirep Statss: Florida, Archer (Mar.), Buena Vista 
(May), Crescent City (Mar.), Dade City (Aug.), Eg- 
mont (Apr., June), Everglade (Apr.), Jupiter (Apr.), 
Lake Alfred (June, July, Nov.); Texas, Brownsville 
(June, July, Aug.), Cotulla (May), Dallas (June), 
Gainesville (Nov.), Kerrville, Logan (June); Arizona, 
Baboquivari Mts. (June, July, Aug.), Nogales (July), 
Woodruff (June); California, Alameda County (Sept.), 
Garden City (Jan., Feb.), Loma Linda (June, Aug., 
Sept., Oct.), Palo Alto (Sept.), San Diego (May, June, 
Sept.), San Gabriel Mts. (June), Santa Paula, upper 
Ojai (uly); Washington, Pullman (May, July, Aug.), 
Yakima (May, June), Walla Walla (June, Aug.), 
Wenatchee (July); Idaho, Springfield (June); Nevada, 
Pyramid Lake, Reno; Utah, Provo (July), Vineyard 
(June); Colorado, Boulder Creek Canyon (May), Den- 
ver (May); Oklahoma, Stillwater (Aug.); Rhode Island, 
Weekapaugh (Aug.) ; New Hampshire, Hampton (Sept.). 
Canava: Saskatchewan, Oxbow (June). Cusa: San- 
tiago (Feb., June, Oct., Nov.). Purrtro Rico: Dorado 
(May), Isabella (Jan.), Mayagiiez (Jan.), Palmas Aba- 
jas (near Guayaman), Puerto Real (Vieques Isl., Apr.), 
Rio Piedras (Apr.), San Germén (Apr.). GRENADA. 
Jamaica: Newport (Feb.). México: Eldorado (Mar.), 
Mexico City (Nov.), Oaxaca, Crizaba, Tehuacén (Apr.). 
Frence Guiana: St. Laurent Maroni. Brazin: Santa 
Catarina (July, Dec.), Sao Paulo (May). Unucuay: 
Montevideo. Paraguay: Villarrica (Jan., Apr., July, 
Sept., Nov.). Prrt: Angasmarca. GanApacos: Con- 
way Bay (Apr.). 

Presumably to be found in every Central and South 
American country. 

This species is of economic importance as an occa- 
sional pest of cultivated beans and to American ento- 
mologists is known as the “limabean pod borer.” It 
has an extensive literature and has been described under 
many names, having at least 13 Old World syn- 
onyms. I have omitted these (for reasons given under 
discussion of the genus) and listed only the American 
synonyms. Some authors have treated schisticolor and 
rubribasella as distinct species, others as races of 
zinckenella. ‘They appear as the latter in our latest list 
(McDunnough, 1939). However, they are no more 
than color forms intergrading with the typical zincke- 
nella and deserve no separate designation. The 
accepted Old World synonymy is given in the Ragonot 
Monograph (1893). The best and most complete 


accounts of the life history and immature stages will be 
found in the U. S. Department of Agriculture bulletins 
cited here (Chittenden, 1909, and Hyslop, 1912). For 
additional references the “Review of Applied Hnto- 
mology” should be consulted. 


Genus 54: Glyptocera 


[Venational division B. Forewing with veins 4-5 connate, 
smooth; hind wing with veins 4-5 distinctly stalked for over half 
their lengths; 3 connate with the stalk of 4-5. Eighth abdominal 
segment of male with compound tufts. Antenna of male with 
shallow, spined sinus at base of shaft. Labial palpus upturned, 
rough scaled beneath. Maxillary palpus squamous. Male 
genitalia with sacculus of harpe produced at apex as a long, free 
spine; apex of gnathos a short, stout hook; transtilla complete 
but weakly sclerotized; a single strong cornutus on penis. 
Female genitalia with bursa partly sclerotized, otherwise finely 
spined but without definable signa; ductus bursae flattened and 
partially sclerotized.] 


54. Genus Glyptocera Ragonot 


Glyptocera Ragonot, Ent. Amer., vol. 5, p. 114, 1889; Bull. Soc. 
Ent. France, 1890, p. vii; Monograph, pt. 1, p. 209, 18938.— 
Hulst, Phycitidae of N. Amer., p. 140, 1890.—Forbes, 
Cornell Mem. 68, p. 621, 1923. (Type of genus: Nepho- 
pteryz consobrinella Zeller.) 

Tongue well developed. Antenna of male pubescent, 
shaft with shallow sinus towards base containing a row 
of short, toothlike spines; antenna of female simple. 
Labial palpus upturned; second segment rough scaled 
beneath; third segment about one-third as long as 
second, acuminate. Maxillary palpus rather large, 
squamous. Forewing smooth; 11 veins; vein 2 from 
before but rather near lower outer angle of cell; 3 from 
the angle, separated from 4—5 at base, but about half 
as far from them as from 2; 4 and 5 connate; 6 from 
below upper angle of cell, straight; 8 and 9 long stalked 
(for slightly over two-thirds their lengths) ; 10 from the 
cell, closely approximate to the stalk of 8-9 for nearly 
half its length; male without costal fold. Hind wing 
with vein 2 from before but near lower outer angle of 
cell; 3 from the angle, connate with the stalk of 4-5; 
4 and 5 distinctly stalked for over half their lengths; 7 
and 8 closely approximate for a short distance from 
cell; cell about half the length of wing; discocellular 
vein curved. Highth abdominal segment of male with 
compound ventral scale tuft. 

Male genitalia with apical process of gnathos a short, 
stout, simple hook. ‘Transtilla complete but very 
weakly sclerotized, a simple, arched band. Harpe with 
a row of fine, erect hairs from below costa near base; a 
slender, short, clasperlike projection from just below 
base of costa; sacculus produced at apex as a long, free, 
spinelike projection. Vinculum a small V-shaped plate 
with elongate, bandlike, central projection attaching 
to apex of aedeagus. Penis armed with a single, strong 
cornutus. Vinculum stout, longer than greatest width. 

Female genitalia with bursa sclerotized over nearly 
half of one surface, densely and finely spined over most 
of remaining area; ductus bursae flattened, a broad 
sclerotized band extending its length on ventral surface; 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 101 


ductus seminalis from bursa near junction of bursa and 
ductus bursae; genital opening simple. 

The combination of male characters and the rather 
broadly squamous maxillary palpi distinguish the 
genus. The long, straight, free sacculus at once identi- 
fies it and distinguishes it from the genera which follow, 
and which are related to Nephopteryx and Salebria. The 
weak transtilla also occurs in Meroptera and some species 
of Nephopteryz. The female genitalia are similar to 
those of Nephopteryr. 

Glyptocera contains but the one North American 
species. 


208. Glyptocera consobrinella (Zeller) 
Fieurss 327, 811 


Nephopteryx consobrinella Zeller, Verh. zool.-bot. Ges. Wien, vol. 
22, p. 548, 1872. 

Glyptocera consobrinella (Zeller) Ragonot, Ent. Amer., vol. 5, p. 
114, 1889; Monograph, pt. 1, p. 210, 1893.—Hulst, Phyciti- 
dae of N. Amer., p. 140, 1893.—Forbes, Cornell Mem. 68, 
p- 621, 1923.—McDunnough, Check list, No. 6148, 1939. 

Ambesa busckella Dyar, Proc. Ent. Soe. Washington, vol. 6, p. 
108, 1904.—Barnes and McDunnough, Contributions, vol. 
3, p. 196, 1916. 

Forewing ashy gray shaded and marked with black, 

the blackish shade most conspicuous broadly bordering 
the antemedial line on inner side, expanded narrowly 
along inner margin to base and also broadening the 
black outer border of the line at costa; basal area other- 
wise pale clay color more or less shaded with pale sal- 
mon especially in lower fold; on many specimens a 
blotch of the same salmon shade over the middle of 
inner margin; antemedial line well out beyond basal 
third, slightly oblique, sharply sinuate, whitish gray 
with black inner and outer bordering lines; subterminal 
line outwardly rounded at middle and angled above 
and below the bulge, margined inwardly and outwardly 
by distinct black lines; discal spots confluent, forming 
a black line along discocellular vein which expands be- 
low into short black streaklets or smudges on the lower 
veins; terminal dots confluent, forming a black line 
along termen. Hind wing smoky white with a faint 
ocherous tint; the veins not appreciably darkened; a 
narrow dark line along termen. Alar expanse, 20-25 
mm. 
Genitalia as given for the genus. On some males in 
addition to the strong cornutus there is a second much 
smaller and weaker spine but this is variable and ap- 
parently not a constant structure. 

TYPE LOCALITIES: Texas (consobrinella, in MCZ); 
Plummers Island, Md. (busckella, in USNM). 

Foop piant: Viburnum, maple (Dyar and Ely 
rearings). 

Disrrisution: Unitep Statss: Maine, Lincolnville 
(May, reared by Dyar on Viburnum), Orono, Sebec 
Lake (July), Weld (July); New Hampshire, Hampton 
(July); Vermont, Clarendon; Massachusetts, Framing- 
ham (June), New York, Plattsburg (July), Valcour Isl. 
(July); Connecticut, East River (June, July); New Jer- 
sey, Basking Ridge; Pennsylvania, New Brighton (June, 
July, Aug.), Pittsburgh (July); Maryland, Plummers 


Isl. (May, Aug.); District of Columbia, Washington 
(July, reared by Dyar from larva on maple); Tezas, 
Kerrville (Mar.); Jilinois, Chicago (July). Canapa: 
Ontario Trenton (July); Quebec, St. Johns (June); Nova 
Scotia, Cape Breton Isl. (June); Newfoundland, Hum- 
ber Mouth (Bay of Fundy, Aug.). 

Presumably generally distributed over the eastern 
section of the continent from Canada to Texas. 


Genera 55-58: Pima to Catastia 


[Venational division B. Forewing with veins 4-5 separate or 
connate at base, smooth; hind wing with veins 4 and 5 anas- 
tomosed for about half their lengths, discocellular vein consider- 
ably extended at lower angle. Eighth abdominal segment of 
male with paired tufts. Antenna of male with shallow, spined 
sinus at base of shaft (also on Catastia a weak scale tuft). Labial 
palpus porrect or oblique, not grooved. Maxillary palpus of 
male squamous or subsquamous. Male genitalia with costa of 
harpe strongly sclerotized throughout its length and slightly 
produced at apex; gnathos terminating in a short, stout hook; 
transtilla incomplete or absent; penis armed with two stout 
cornuti (Pima, Interjectio) or a single strong cornutus (Ambesa, 
Catastia). Female genitalia with bursa partially sclerotized and 
sometimes (Pima, Interjectio) granulate-scobinate but without 
definable signa; ductus bursae more or less sclerotized and con- 
siderably broadened at genital opening.] 


55. Genus Pima Hulst 


Pima Hulst, Ent. Amer., vol. 4, p. 114, 1888; Phycitidae of N. 
Amer., p. 164, 1890. (Type of genus: Pima fosterella Hulst.) 

Epischnia Authors (not Hiibner) Ragonot, Ent. Amer., vol. 5, 
p. 115, 1889; Monograph (in part) pt. 1, p. 493, 1893.— 
Forbes, Cornell Mem. 68, p. 629, 1923.—McDunnough, 
Canadian Ent., vol. 67, p. 176, 1935. 


Tongue well developed. Antenna finely pubescent; 
on male with a very slight incurvation in base of shaft 
containing a row of minute, black, toothlike spines (6 
to 8). Labial palpus porrect (the second segment 
oblique, the third projected forward); extending at 
least twice the length of the head beyond it; second 
segment broadly (triangularly) scaled; third segment 
as long as second. Maxillary palpus minute but rather 
broadly scaled. Forewing smooth; 11 veins; vein 2 
from before lower outer angle of cell; 3 from the angle, 
closer to 4 than to 2; 4 and 5 separated at base; 6 from 
well below upper angle of cell, straight; 8 and 9 stalked 
for one-half of less than half their lengths; 10 from the 
cell, more or less approximate to the stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from before 
lower outer angle of cell; vein 3 from 4 well beyond 2 
and considerably shorter than 2; 4 and 5 anastomosed 
just beyond 3 for nearly half their lengths; 7 and 8 
contiguous or closely approximate for a short distance 
from cell; cell slightly less than half the length of wing; 
discocellular vein curved, greatly extended at lower 
angle (running into 4 just beyond base of vein 8). 
Eighth abdominal segment of male with a pair of ven- 
trolateral hair tufts. 

Male genitalia with uncus hoodlike; its terminal mar- 
gin broadly rounded. Apical process of gnathos a short, 
stout hook. ‘Transtillaabsent. Harpe elongate, taper- 
ing to bluntly pointed apex; costa broadly and strongly 


102 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


sclerotized for the entire length of the harpe, its apex 
blunt, slightly produced and usually forked; sacculus 
short and narrow, its inner surface covered with dense, 
spinelike hairs (probably the “‘strong spines’”’ mentioned 
by Hulst in his original description of the genus). Vin- 
culum a broad plate with weak lateral lobes. Aedeagus 
long, moderately stout, smooth, slightly curved towards 
base; penis armed with two stout cornuti. Vinculum 
stout, tapering, considerably longer than greatest width. 

Female genitalia with bursa stout, scobinate-granu- 
late and usually with sclerotized patches or folds; ductus 
bursae long (much longer than bursa), flat (ribbonlike), 
sclerotized and granulate for its entire length, broadest 
towards genital opening; ductus seminalis from lobe of 
bursa near junction of bursa and ductus bursae. Eighth- 
segment collar with middle of dorsal anterior margin 
slightly produced. 

Hulst erected the genus Pima for one species (foster- 
ella) which he later (1890) synonymized with albiplagia- 
tella. Ragonot (1889) referred fosterella and the other 
congeneric American species to Hpischnia Hiibner. The 
latter as defined by Ragonot (1893) is a composite of 
several disparate elements and none of the species occur- 
ring in the New World agrees with the type species of 
Epischnia (prodromella, Hiibner). The latter has quite 
different genitalia (figs. 426, 884). Its male antenna 
has a deep sinus but the latter is without the row of 
toothlike spines characteristic of Pima; and the third 
segment of the labial palpus is much shorter (less than 
half the length of the second segment). There is also 
a heavy scale-and-hair tuft on the underside of the pro- 
thorax. This may be only a specific character (as simi- 
lar tuftings are in some other phycitid genera); but the 
structure is entirely lacking on the New World species 
and on the Huropean boisduvaliella Guénée, which is a 
typical Pima on all characters. The venation of Pima 
is like that of Epischinia (fig. 26). 

The species here referred to Pima (except granitella 
and parkerella) have a strikingly similar habitus; but are 
individually variable in size and color, especially among 
examples of western species, several of which exhibit 
both pale and dark forms. The extent of the white 
costal streak on forewing is also individually variable 
and reliance on this and other color features has resulted 
in considerable confusion in the application of names. 
The most reliable specific characters are in the genitalia, 
especially those of the females. The chief male differ- 
ences are in the shape and size of the cornuti, the shape 
of the anellus, and the configuration and width of the 
apex of the sclerotized costa of harpe. These differ- 
ences are trifling and also subject to some variation, es- 
pecially the notching at the apex of costa of harpe. 

The larvae feed in the flowers and seed pods of vari- 
ous Leguminosae. From scattered reared examples in 
the National Collection the species do not seem to be 
confined to specific plants; but there have been no ex- 
tensive and systematic rearings, and these will be needed 
to clear up host relationships and to differentiate any 
possible food-plant races. 


Genus Pima, Species 209-216: P. boisduvaliella 
to P. fulvirugella 


[Forewing with strongly contrasted white costal stripe.] 


209. Pima boisduvaliella (Guénée), new combination 
Figures 299, 776 

Epischnia boisduvaliella Guénée, Europaeorum Microlepidopter- 
orum index methodicus .. . , p. 81, 1845.—Ragonot, Ent. 
Monthly Mag., vol. 22, p. 23, 1885; Monograph, pt. 1, p. 
518, 1893 (part).—Lafaury, Ann. Soc. Ent. France, ser. 6, 
vol. 5, p. 398, 1885.—Spuler, Die Schmetterlinge Europas, 
vol. 2, p. 209, 1910.—Meyrick, Revised handbook of British 
Lepidoptera, p. 382, 1928.—McDunnough, Canadian Ent., 
vol. 67, p. 176, 1935 (part); Check list, No. 6251, 1939. 

Forewing pale fawn on lower half shading into muddy 
fawn along the lower border of the white costal stripe 
where it forms a more or less distinct dark band through 
the center of the wing from base to termen; no indica- 
tion of transverse lines; extreme costal edge blackish 
gray weakly peppered with white, this dark shade en- 
croaching on the white costal stripe beyond middle; 
white costal stripe extending to apex, its lower margin 
edged by a fine gray-black line; discal dots at end of cell 
minute, blackish, the upper one lying within the black- 
ish edge of the white stripe and frequently indistinguish- 
able, the lower one distinct but not conspicuous; some 
faint scattered black dotting and dusting on and below 
vein 1b beyond base. Hind wing pale ocherous brown, 
on some specimens with a faint ocherous tint. Alar 
expanse, 19-25 mm. 

Male genitalia with apex of costa of harpe slightly 
expended, concave or weakly notched, the upper angle 
of the notch rounded, the apical margin oblique. 
Cornuti both broadened for more than half their 
lengths; the broad part of the shorter thorn longitudi- 
nally grooved (fluted) ; the longer thorn but slightly less 
than half the length of the aedeagus. 

Female genitalia with bursa narrowly heart shaped; a 
small rounded or oval sclerotized plate in bursa near its 
middle; bursa longitudinally wrinkled, the wrinklings 
weakly sclerotized; also a couple of broader, more 
tortuous, sclerotized folds, the one curving about junc- 
tion of bursa and ductus bursae serrate along its edge; 
bursal granulations weak, scobinations in lobed area 
adjacent to junction of ductus bursae fine but rather 
dense; lower margin of ductus bursae at genital opening 
straight, not produced. 

TypE Locatity: Switzerland (location of type un- 
known). 

Foop pLants: Ononis, Anthyllis, Lotus, Astragalus 
(European records). 

DistRIBUTION: Europr. Canapa: Manitoba, Aweme 
(May), Beulah (June, Aug.), Winnipeg; Saskatchewan, 
Regina (June, Aug.); Alberta, Lethbridge (June). 

This European species is definitely established in 
North America, but most of the American references to 
boisduvaliella apply to other native species. The only 
American examples of the true boisduvaliella I have 
seen are some ten specimens from the Canadian localities 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 103 


cited above. Old World synonymns and doubtful 
American references are omitted from the foregoing 
literature citations. There can be no reasonable doubt 
of the correctness of the European synonymy of farrella 
(Curtis) or lafauriella (Constant) (their references will 
be found in the Ragonot Monograph); but the status 
of the supposed Asiatic variety tabulella Staudinger is 
doubtful. Ragonot’s reference of albocostalialis Hulst 
as a variety of boisduvaliella is obviously incorrect. 

In size boisduvaliella averages appreciably smaller 
than any other species of Pima. Occasional examples 
have an expanse equal to that of small specimens of some 
of our American species (24-25 mm.); but most speci- 
mens expand 24 mm. or less, while in the other species 
the average expanse is well over 25 mm. 

The male genitalia are similar in all essential charac- 
ters to those of albiplagiatella; but the female genitalia, 
while of the same general habitus, differ in marked and 
apparently consistent details—the folds of the bursa are 
more weakly sclerotized, the bursa itself decidedly 
smaller, and the granulations and scobinations in bursa 
weaker. The contrasts are somewhat greater than 
shown in the figures. 


210. Pima albiplagiatella (Packard), new combination 
Figures 305, 777 


Myelois albiplagiatella Packard, Ann. Lyc. Nat. Hist. New York, 

feud! vol. 10, p. 269, 1874. 

Epischnia boisduvaliella albiplagiatella (Packard) McDunnough, 
Canadian Ent., vol. 67, p. 70 (larva), p. 176 (part), 1935; 
Check list, No. 6251, 1939. 

Larger than boisduvaliella. The ground color on lower 
half of forewing very pale fawn, paler than that of 
boisduvaliella and containing little of no dark scaling, 
the latter when present confined to a couple of faint 
gray dots on basal third of vein 1b and some very 
sparse gray scaling along inner margin near tornus; the 
contrasted dark band bordering lower margin of the 
white costal stripe brown rather than ruddy; a similar, 
weaker, narrower, brown shade along lower fold for 
most of its length; extreme costal edgepale brownish 
gray, this color encroaching on the white stripe beyond 
middle as in other species; lower discal dot distinguish- 
able but very faint. Hind wing whitish ocherous. Alar 
expanse, 26-31 mm. 

Male genitalia like those of boisduvaliella. Female 
genitalia similar to those of boisduvaliella but con- 
sistently different in minor details. These differences 
were noted in the discussion of boisdwvaliella and are 
shown in the figures. 

Typr Locatiry: New Hampshire (type in MCZ). 

Foop puiant: Lathyrus maritima and presumably 
other Leguminosae. 

Distrisution: Unirep Stares: New Hampshire, 
Hampton (June, July). Canapa: Nova Scotia, White 
Point Beach (Queens County, July). 

Typical albiplagiatella is probably generally distrib- 
uted in eastern Canada and northeastern United States. 
The foregoing records are from specimens whose 
genitalia I have been able to examine, the Nova Scotia 


record from a specimen supplied by McDunnough. He 
has referred albiplagiatella as a race of boisduvaliella. It 
is indeed very close to the European species, but I 
believe the differences in female genitalia justify more 
than racial separation. 


211. Pima albiplagiatella occidentalis, new race 
Figure 303 


A variety occurring in the Rocky Mountain and 
Pacific Coast States. Extremely variable in color and 
the amount of blackish dusting on forewing. Average 
specimens in Washington, Colorado, and New Mexico 
with ground color very pale fawn (cream white in some 
specimens) ; the dark border of the white costal stripe 
ranging from pale ocherous brown to dark gray-brown; 
two distinct black dots on basal third of vein 1b, enclos- 
ing a white spot; the remainder of 1b more or less 
streaked with black outwardly and a fine peppering of 
black scales and white scales along inner margin near 
tornus; costal edge brownish or blackish gray, encroach- 
ing on and attenuating the white costal streak beyond 
middle, and sometimes blotting it out before apex. In 
southern California the paler specimens show more 
blackish dusting expecially along the outer veins, and 
sometimes a faint white subterminal line can be dis- 
tinguished; the darkest specimens are almost a uniform 
dark gray with the blackish gray dusting obliterating 
all markings except the white spot on vein 1b and the 
contrasted white costal streak; between these two ex- 
tremes there is every intergrade in series from any given 
locality; lower discal spot small, but usually distinct, 
blackish. Hind wings ocherous white to dark smoky 
gray. Alar expanse, 24-31 mm. 

Male genitalia like those of typical albiplagiatella 
except that apex of sclerotized costa of harpe is some- 
what more swollen; in average specimens the outer 
edge of apex is slightly concaved; one example from 
Alamogordo, N. Mex., has the edge almost straight, 
but there is a gradual intergradation from this to forms 
with the apex as in our figure of albiplagiatella (fig. 
305c). The cornuti are like those of typical albiplagia- 
tella. The female genitalia are like those of eastern 
albiplagiatella. A series of 24 preparations from the 
various western localities exhibits no significant varia- 
tion from type and nothing approaching the bursa of 
boisduvaliella. 

Type LocaLity: Pullman, Wash. (type in USNM, 
61342). 

Foop piants: Astragalus, Lathyrus. 

Described from male type (C. V. Piper, collector, 
May 1905) and one female paratype (J. F. G. Clarke, 
May 30, 1924) from the type locality, and paratypes as 
follows: One male from above Golden, Colo. (H. G. 
Dyar No. 17468, May 29, 1901); one female, Beulah, 
Colo. (June 21, 1900, W. D. Kearfott Collection) ; 2 male 
and one female, Alamogordo, N. Mex. (May 15, 17, 
1929, reared by M. W. Talbot from Astragalus wootont) ; 
one female, Phoenix, Ariz. (Apr., Kunze, collector); one 
male, Palm Springs, Calif.; one male and one female, 


104 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Loma Linda, Calif. (May, June); one female, Clare- 
mont, Calif. (Baker, no date); and one female, San 
Diego, Calif. (H. G. Dyar, May 22, 1924). These are 
from a series of 85 specimens in the U. S. National 
Collection from the following localities: United States: 
Colorado, Beulah (June), Denver, Golden (May); New 
Mexico, Alamogordo, (May), Jemez Springs (Apr., 
May), Pecos; Arizona, Phoenix (Apr.). ‘Southern 
Arizona”; California, Claremont, Loma Linda (Mar., 
Apr., May, June), ‘Los Angeles County” (May), Mir- 
age Lake (San Bernardino County, Apr.), Olancha (Apr., 
May), Palm Springs (Mar.), San Diego (Mar., May), 
“Shasta County,” ‘Sierra Nevada’; Oregon, Baker 
(June); Washington, Copalis (a gray specimen with very 
dark hind wings, reared under Special Survey No. 
26286, Mar. 27, 1945, from Lathyrus sp.), Palouse Falls 
(May), Pullman (May, June, July, Aug.), Walla Walla 
(June, July), Yakima (May). 


212. Pima fosterella Hulst 
Fieures 300, 783 


Pima fosterella Hulst, Ent. Amer., vol. 4, p. 114, 1888. 

Pima albiplagiaiella Hulst (not Packard), Phycitidae of N. Amer., 
p. 164, 1890. 

Epischnia albiplagiatella Ragonot (not Packard), Monograph, 
pt. 1, p. 518, 1893. 

Epischnia fulvirugella McDunnough (not Ragonot), Canadian 
Ent., vol. 70, p. 178, 19385. 

Epischnia fosterella (Hulst) MecDunnough, Check list, No. 6252, 
1939. 


A large, pale species similar in general appearance to 
typical eastern albiplagiatella, but without any trace of 
a dark shade in lower fold; the dark shade along lower 
border of the white stripe also paler, a light drab brown; 
a single small black spot on basal third of vein 1b, but 
no white spot and seldom any further dark shading on 
1b, but more or less gray dusting along outer two-thirds 
of inner margin; white costal streak usually obliterated 
before apex, rarely reaching apex; lower discal dot 
usually well contrasted, but minute, blackish. Hind 
wing whitish ocherous or pale smoky fuscous. Alar 
expanse, 27-35 mm. 

Male genitalia with harpe somewhat longer in pro- 
portion to tegumen and uncas than in preceding species; 
sclerotized costa broadened and forked at apex, the 
prongs of the fork poimted. Cornuti spaced apart; 
neither one appreciably flattened or ribbed towards 
base; the longer slightly less than one-third the length 
of aedeagus. 

Female genitalia with little or no sclerotization of 
bursa except immediately about junction of bursa and 
ductus bursae. The extreme of sclerotization is shown 
in figure 783, from an Arizona female; the female type 
shows none except about the junction with ductus. 
Bursa finely scobinate over entire inner surface. Duc- 
tus bursae produced at apex into a projecting shield, 
its apical margin variable, pointed to evenly rounded. 

Typr wtocauity: Colorado (type in AMNH, ex 
Rutgers). 

Foop pirant: Unknown. 

Distrrsution: Unirep States: Colorado, Baileys 


(July), Denver (June), Durango (June), Golden (June), 
Gunnison County (near Altmont, July); Arizona, 
Williams; Utah, Hureka, (May), Park City (July); Mon- 
tana, Miles City. Canapa: Nordegg (June). 

A distinct species easily recognized by its female 
genitalia. The type () at Rutgers bears only a number 
label (‘43’) but is obviously a Colorado specimen and 
an authentic type. A male paratype matching it is 
in the National Museum. It and one other male and 
two females labeled only ‘‘Colo.”? All Colorado exam- 
ples are larger specimens. The Nordegg specimen (¢) 
had been received from Dr. McDunnough and formed 
part of the series he had treated as fulvirugella in his 
1935 paper. It and a male from EHureka, Utah, show 
some black scaling on the outer veins. They are super- 
ficially very much like some specimens of the western 
race of albiplagiatella and except for their genitalia 
could easily be confused with them. 


213. Pima vividella (McDunnough), new combination 
Figures 302, 780 


Epischnia vividella McDunnough, Canadian Ent., vol. 67, p. 
179, 1935; Check list, No. 6256, 1939. 

Forewing salmon pink below costal white stripe, 
shading below and towards tornus into pinkish ocherous; 
some gray dusting along outer two-thirds of inner 
margin; a black dot, followed by an obscure white one, 
on vein 1b at basal third; white costal stripe attenuated 
by smoky costal scaling on its outer half and obliterated 
before apex. Hind wing pale smoky with a faint 
ocherous tint. Alar expanse, 27-30 mm. 

Male genitalia with apex of costa of harpe forked as 
in fosterella; but upper prong somewhat longer and 
sharper than the lower one. Cornuti closely approxi- 
mate; the longer one with flattened but not ribbed 
basal part, a trifle longer than one-third of the aedeagus. 
Female genitalia resemble those of fosterella except: 
Bursa, proportionally smaller, with two pitted and 
sclerotized patches; sclerotization at junction of bursa 
and ductus bursae serrate along one edge. 

Tyrer tocatity: Lethbridge, Alberta (type in Ca- 
nadian Nat. Coll.). 

Foop piant: Unknown. 

Distrisution: Manitoba, Aweme (June); Saskatche- 
wan, Saskatoon (June); Alberta, Lethbridge (June). 
Also recorded by McDunnough from Beulah, Manitoba 
(June) and Indian Head, Saskatchewan (July). 

A good species, close to but distinct from fosterella; 
easily identified by its male cornuti, female genitalia, 
and salmon-colored forewings. 


214. Pima albocostalialis (Hulst), new combination 
Fiaures 301, 778, 779 


Ephestia albocostalialis Hulst, Trans. Amer. Ent. Soc., vol. 13, 
p. 64, 1886. 

Epischnia albocostalis (Hulst), Phycitidae of N. Amer., p. 163, 
1890. (Emended spelling). 

Epischnia boisduvaliella albocostalis (Hulst) Ragonot, Mono- 
graph, pt. 1, p. 520, 1893. 

Epischnia albocostalialis (Hulst) McDunnough, Canadian Ent., 
vol. 67, p. 178, 1935; Check list, No. 6254, 1939. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 105 


Forewing dark gray shading into very dark grayish 
or blackish brown towards the white costal streak; the 
latter ending just before apex; no spottings or other 
markings on the wing, even the lower discal dot lost in 
the dark suffusion of the wing. Hind wing whitish 
at base, shading into a smoky outer area. Alar expanse, 
26-34 mm. 

Male genitalia with apex of costa of harpe narrow, 
very slightly notched, reaching only to end of cucullus. 
Harpe itself shorter in proportion to length of tegumen 
and uncus than that of any preceding species except 
boisduvaliella. Cornuti spaced apart as in fosterella, 
the longer one about one-third the length of aedeagus. 

Female genitalia with two rather large, irregularly 
shaped, pitted and sclerotized patches in bursa, one at 
the terminal end, the other on the left side (viewed 
ventrally); bursa, otherwise membranous except for a 
sclerotization about junction with ductus bursae; 
ductus bursae not produced at genital opening, its 
apical margin straight. 

Type tocauity: California (type in AMNH, ex 
Rutgers). 

Foop prant: Unknown. 

Distrisution: California, Claremont, ‘En route Im- 
perial to Bishop” (May), Loma Linda (Mar.), River- 
side (Apr.), Shasta Retreat (Siskiyou County, July); 
Washington, Olympia (June), Rochester (June), Seattle, 
Wenatchee (May). McDunnough also records the spe- 
cies from Lethbridge, Alberta, Canada. His specimen 
is undoubtedly this species, but may represent one of 
the varieties that follow. 

The type in the Rutgers Collection lacks antennae 
and abdomen but matches otherwise the examples in 
the National Collection, so there can be no question of 
the application of Hulst’s name. He emended its spell- 
ing to albocostalis in 1890, but gave no reason for doing 
so, hence we shall probably have to perpetuate the 
original barbarous spelling. 


215. Pima albocostalialis subcostella (Ragonot), new combination 


Epischnia subcostella Ragonot, N. Amer. Phycitidae, p. 10, 1887; 
Monograph, pt. 1, p. 520, 1893.—MeDunnough, Check list, 
No. 6255, 1939. 

Forewing with white costal streak as in typical albo- 
costalialis; below it a bordering band of blackish brown 
with a lighter brown shade below it as far as lower fold; 
these dark shades terminating beyond cell in what would 
be the position of the antemedial line if one were pres- 
ent, the outer margin of the dark shade outwardly 
angled at middle; remainder of wing pale, ashy gray 
with a dusting of blackish scales along outer two-thirds 
of inner margin and (in some specimens) blackish 
streaklets on the veins before termen; on basal third of 
vein 1b a white dot preceded by a black shade. Hind 
wing white; smoky shading limited to a narrow line 
along termen and a very small area at apex; on the fe- 
males the smoky tints slightly more extended. Alar 
expanse, 25-27 mm. 

Genitalia, male and female, as in typical albocostal- 
talis. 


Type Locaity: Utah (type in Paris Mus.). 

Foop piant: Unknown. 

A male and three females from Eureka, Utah (May, 
June), a female from Bellevue, Washington County, 
Utah (May) and a male from Pyramid Lake, Nev., are 
before me. They match Ragonot’s description and 
figure of subcostella in every detail. The name may 
represent nothing more than a color variety of albo- 
costalialis and if there were not an old name available 
for it, and one that until now was supposed to repre- 
sent a distinct species, I should not have named it. I 
am holding subcostella as a trinomial against the possi- 
bility that it may represent a valid local race. 

There are also before me what appear to be two 
other varieties with male genitalia identical to those of 
albocostalialis: 

Variety a: A large form with a reddish brown band 
bordering the white costal stripe; the red-brown color 
shading into ocherous fawn on lower and outer areas 
of the wing; lower discal dot present, black; no white 
spot or other appreciable marking on vein 1b. Hind 
wing ocherous white with a pale broken line along ter- 
men. Alar expanse, 34-35. Represented by three 
males from Manitou, Colorado (H. G. Dyar, coll. No. 
6062-6065, May 5, 1891). 

Variety b: A variable variety ranging from dark gray, 
suffused examples to a couple with the entire median 
area of the forewing a ruddy fawn color; the white spot 
is present on vein 1b, but very faint on the darker 
specimens. Hind wings white to pale smoky fuscous. 
Alar expanse, 26-27 mm. 


The female genitalia differ rather markedly from 
those of subcostella or typical albocostalialis in that the 
sclerotized patches in bursa are situated on opposite 
sides of that organ. This arrangement is consistent for 
the females from both New Mexico and Arizona. 


Distrisution: Colorado, Glenwood Springs (June); 
New Mexico, Fort Wingate (June), Pecos (June); Ari- 
zona, Huachuca Mts., White Mts. (Aug.). 

Superficially this form looks like nothing but a color 
variant of subcostella. However, if the single female 
genitalic difference should hold through extended series 
it will need further designation. 


216. Pima fulvirugella (Ragonot), new combination 
Ficure 304 
Epischnia fulvirugella Ragonot, N. Amer. Phycitidae, p. 10, 
1887; Monograph pt. 1, p. 521, 1893—MceDunnough, Check 
list, No. 6253, 1939. 

I have seen nothing that exactly matches Ragonot’s 
description or figure (Monograph, pl. 16, fig. 43). A 
male before me from San Francisco has a similar, at- 
tenuated white subcostal streak and rather pronounced 
black lining on the veins from cell. Its genitalia, how- 
ever, do not match those of the type, and the moth 
itself is smaller (27 mm.). Ragonot gives the alar 
expanse of his type as 30 mm. 

Type Locauity: California (type in Paris Mus.). 

Foop pranr: Unknown. 


106 


Genus Pima, Species 217 and 218: P. granitella 
and P. parkerella 


[Forewing with white costal streak obscure or absent.] 


217. Pima granitella (Ragonot), new combination 
Fieures 307, 781 

Epischnia granitella Ragonot, N. Amer. Phycitidae, p. 9, 1887; 
Monograph, pt. 1, p. 523, 1893.—Hulst, Phycitidae of N. 
Amer., p. 162, 1890.—McDunnough, Check list, No. 6258, 
1939. 

Megasis piperella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 
110, 1904. 

Forewing gray finely powdered with white, the white 
dusting concentrated between the veins; the latter finely 
lined with blackish brown giving the wing (to the naked 
eye) a longitudinally dark-lined, over-all pale bluish 
gray appearance; on some specimens a trace of the usual 
costal white strip as a narrow white line along top of 
cell for a short distance from base; the lower blackish 
discal spot usually distinct but occasionally obliter- 
ated; on a few specimens the subterminal line slightly 
indicated by interruptions in the blackish streaks on the 
veins, but, otherwise, transverse lines absent. Hind 
wing pale brown, unicolorous, except for a narrow, very 
slightly darker line along termen. Alar expanse, 22- 
23 mm. 

Male genitalia with sclerotized costa of harpe broad- 
ened, slightly notched and produced at apex. Cornuti 
lying one before the other, of nearly equal length, not 
(or but very slightly) broadened and not flattened at 
base; the longer one slightly less than one-third the 
length of aedeagus. Female genitalia similar to those 
of albocostalialis except sclerotized patches in bursa, 
narrower, more elongate. 

TypE Locatity: California (granitella, in Paris Mus.); 
Pullman, Wash. (piperella, in USNM). 

Foon piant: Crotolaria. 

Distrisution: Colorado, Glenwood Springs (Apr., 
May); Utah, Eureka (May, June), Richfield (May), 
Stockton (May); New Mexico, Jemez Springs (Apr., 
May); Arizona, Dewey (Apr.); Nevada, Baker (May); 
California, Argus Mts. (May), Crows Landing (May), 
Los Angeles County (May); Washington, Pullman, 
Walla Walla (May). 

A distinct species, easily recognized by its wing pat- 
tern. 

218. Pima parkerella (Schaus), new combination 
Figures 306, 782 


Epischnia parkerella Schaus, Proc. Ent. Soc. Washington, vol. 26, 
p. 196, 1924.—MecDunnough, Check list, No. 6250, 1939. 

Forewing olive buff, costal and inner margins whitish 
more or less overlaid with a fine peppering of blackish 
scales; on some specimens a scattering of similar dark 
scales in the central area and near outer margin along 
some of the veins; no contrasted white costal strip; 
transverse lines well marked on most specimens, white, 
rather broad (absent on one example before me); the 
antimedial line outwardly oblique from costa to inner 
margin, nearly straight, at most with a slight bend at 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


vein 1b; subterminal line inwardly oblique from costa 
to inner margin, with a decided notch at lower fold, in- 
wardly margined by a faint dark shade; discal spots usu- 
ally obsolete, when distinguishable very faint. Hind 
wing very pale fuscous with a narrow dark line along 
termen; on the paler examples the hind wing has a slight 
ocherous-gray tint. Alar expanse, 30-35 mm. 

Male genitalia somewhat stouter than those of pre- 
ceding species. Sclerotized costa of harpe not appre- 
ciably broadened at apex nor projecting beyond apex 
of cucullus; very slightly if any notched at apex. Cor- 
nuti lying close to each other, the apex of one projecting 
slightly beyond that of the other; about one-third as 
long as aedeagus. 

Female genitalia with bursa minutely scobinate, at 
left posterior angle developed as a protruding lobe with 
thickened membrane; ductus bursae developed into a 
projecting shield at genital opening. 

Typ Locatity: Bozeman, Mont. (type in USNM). 

Foop priant: “Locoweed”’ (Astragalus). 

Represented in the National Collection by nine ex- 
amples from the type locality (July). The species is 
easily recognized by its forewing markings. It is the 
only Pima with anything approaching distinct trans- 
verse lines. 


56. Interjectio, new genus 


Tyrer oF GENus: Ambesa columbiella McDunnough. 

Characters as in Pima except: Labial palpus extend- 
ing little more than the length of the head beyond it, 
third segment less than half as long as second; apical 
process of gnathos a broad, flanged plate terminating in 
a short slender hook; harpe short, hardly extending be- 
yond apical margin of uncus; vinculum little longer than 
greatest width; genitalia generally broader andmore 
chunky than those of Pima; ductus bursae of female 
short, little, if any, longer than bursa. 

This genus is intermediate between Pima and Ambesa. 
From the latter it differs chiefly in having at least two 
cornuti on the penis and the granulations of the ductus 
bursae uninterrupted for its entire length. 


219. Interjectio denticulella (Ragonot), new combination 
Fieure 308 


Pristophora denticulella Ragonot, N. Amer. Physicidae, p. 6, 1887. 
Ambesa lallatalis Authors not Hulst (in part) Ragonot, Ent, 
Amer., vol. 5, p. 115, 1889; Monograph, pt. 1, p. 241, 1893. 
Ambesa denticulella (Ragonot) McDunnough, Canadian Ent., 
vol. 67, p. 174, 1985; Check list, No. 6158, 1939. 
Forewing white dusted and marked with black, 
making the ground color (of fresh specimens) white 
with a very faint bluish tint; the dark markings strongly 
contrasted, consisting of streaks and spots chiefly indi- 
cating broken margins of the antemedial and postmedial 
lines; the antemedial line itself obscure except between 
lower fold and inner margin when it is an inwardly 
curved white line preceded by a black spot and followed 
by a thin black marginal line; above, indicated only by 
its outwardly oblique, broken outer margin, consisting 
of a short black line from costa and black streaklets on 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 107 


upper and lower veins of cell; subterminal line indicated 
by an oblique, deeply dentate, broken, black inner 
border; black discal dots at end of cell distinct, the 
upper minute, the lower a short but conspicuous streak; 
a row of black dots along termen; the blackish shading 
otherwise consists of very fine dusting between the 
veins, supplemented by some faint brownish shading 
in the folds. Hind wing whitish brown, the veins very 
slightly dakened; a pale brown line along termen. Alar 
expanse, 32-34 mm. 

Male genitalia with apical process of gnathos with 
the lateral angles of the flanged plate incurved and 
broadly rounded. Apex of cucullus not projecting 
beyond sclerotized costa of harpe. Cornuti two stout 
thorns, less than half as long as aedeagus; the latter 
short, stout, straight. 

Type tocauity: North America, but otherwise 
unspecified (type in Paris Mus.). 

Foop pLant: Unknown. 

Distrisution: Unirep States: Washington, Wenat- 
chee (May). Canapa: British Columbia, Keremeos 
(June). 

This species was removed by McDunnough (1935) 
from the synonymy of Ambesa lallatalis (Hulst) where 
it was placed originally by Ragonot (1889), presumably 
on the basis of specimens of supposed Jallatalis sent him 
by Hulst. The latter species was misidentified by 
Hulst himself as well as by later authors. 


220. Interjectio columbiella (McDunnough), new combination 
Fiaure 784 


Ambesa columbiella McDunnough, Canadian Ent., vol. 67, p. 
175, 1935; Check list, No. 6159, 1939. 

Similar to denticulella but without the well-contrasted 
dark markings of that species; the black spot on inner 
margin, preceding the antemedial line, entirely lacking 
and the dark markings themselves more brown than 
blackish, except for the small, narrow black lower discal 
dot, an occasional black dot on vein 1b near base, and 
some blackish dots along termen; the brown markings 
limited to streakings on the veins and (where the trans- 
verse lines are distinguishable) to a narrow dark shade 
along inner margin of the subterminal line. Hind 
wings as in denticulella. Alar expanse, 23-34 mm. 

The species averages smaller than denticulella, espe- 
cially the females, which are even smaller than any of 
the males before me. 

Male genitalia similar to those of denticulella except 
cornuti distinctly more slender. Female genitalia 
(figured from a Pullman, Wash., specimen from a series 
associated with males of the same locality) with 
sclerotized ductus bursae projecting as a short shield 
at genital opening. 

Type Loca.ity: Oliver, British Columbia (type in 
Canadian Nat. Coll.) 

Foop prant: Unknown. 

Distrizution: Unirep States: Utah, Stockton 
(June); Washington, Pullman (May, June), Yakima 
(June). Canava: British Columbia, Oliver (June); 
Alberta, Lethbridge (July). 


221. Interjectio ruderella (Ragonot), new combination 
Fieurn 785 
Epischnia ruderella Ragonot, N. Amer. Phycitidae, p. 9, 1887; 
Monograph, pt. 1, p. 514, 1893.—Hulst, Phycitidae of N. 
or p. 162, 1890.—McDunnough, Check list, No. 6249, 

This species is known only from the female type, 
which, from Ragonot’s description and figure, must be 
very close to some of the small females of columbiella, 
especially paler examples from Pullman, Wash., except 
that the lower discal spot is a more contrasted black 
streaklet. Alar expanse, 24 mm. 

The genitalia show a broader and stronger sclerotiza- 
tion of the ductus bursae at its junction with bursa than 
typical columbiella; but this character is approached in 
Utah examples of the latter species. 

Type Locauity: “North America” [given as “without 
doubt from California” in the Ragonot Monograph] 
(type in Paris Mus.). 

Foop pLant: Unknown. 


222. Interjectio niviella (Hulst), new combination 


Figure 309 


Iipographis niviella Hulst, Ent. Amer., vol. 4, p. 117, 1888. 

Ambesa niviella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 241, 1893.—Hulst, Phycitidae 
of N. Amer., p. 141, 1890.—McDunnough, Check list, No. 
6157, 1939. 

Forewing chalk white; a large black patch bordering 
inner margin of antemedial line and extending from 
inner margin to top of cell; antemedial line faint but 
complete, oblique from costa to lower vein of cell, thence 
concave to inner margin, bordered outwardly by a black 
line, which is more or less broken into spots on upper 
half; black streaks and wedges on the outer veins indi- 
cating the deeply notched, otherwise obscure sub- 
terminal line; both discal spots conspicuous, black; a 
row of short black streaklets along outer margin; on 
some specimens smears of a faint, pale, ocherous brown 
tint towards apex; a fine brown line along termen. 
Alar expanse, 25-28 mm. 

Male genitalia with apical process of gnathos a broad- 
ly crescentiform plate with the lateral angles slightly 
produced; the terminal hook short and slender. Harpe 
with apex of cucullus projecting beyond the sclerotized 
costa. Cornuti a cluster of several short, stubby 
thorns. Aedeagus rather slender, sinuously curved. 

Type nocauity: Colorado (type in AMNH, ex- 
Rutgers). 

Foop prant: Unknown. 

Distrisution: Unirep States: Colorado; Iowa, 
Sioux City. Canapa: Manitoba, Aweme (July), Cart- 
wright, Winnipeg. 

The type is a female without abdomen. I have seen 
but one other female and it too was without abdomen, 
so the female genitalia could not be studied. These 
specimens matched the males in every detail of color 
and maculation. 


108 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


57. Genus Ambesa Grote 


Ambesa Grote, N. Amer. Ent., vol. 1, p. 98, 1880.—Hulst, Phy- 
citidae of N. Amer., p. 141, 1890.—Ragonot, Monograph, 
pt. 1, p. 237, 1893. (Type of genus: Ambesa laetella Grote.) 

Tongue well developed. Antenna as in Pima and 
Interjectio. Labial palpus oblique, extending above 
vertex; second segment roughly and rather broadly 
scaled; third segment not defected forward, about one- 
third as long as second, acuminate. Maxillary palpus 
minute, filiform. Forewing smooth; venation as in 
Pima. Hind wing with vein 3 from the angle of the 
cell (but separated from discocellular vein by a short 
spur), longer in proportion to 2 than in Pima, 4 and 5 
anastomosed for nearly half their lengths, 7 and 8 closely 
approximate for a short distance from cell; cell one- 
third the length of wing; discocellular vein curved, ex- 
tended at lower angle but not so far asin Pima. Highth 
abdominal segment of male simple or (/aetella) with a 
pair of ventrolateral hair tufts. 

Male genitalia similar to those of Interjectio except: 
Harpe longer in proportion to combined tegumen and 
uncus; its sclerotized costa with upper angle at apex 
produced into a sharp point; sacculus finely haired, not 
with coarse spinelike hairs of Pima or Interjectio. 
Anellus a simple plate without lateral projections. 
Penis armed with a single, long, stout cornutus; over 
half as long as aedeagus. Vinculum as long as greatest 
width. 


Female genitalia with bursa unsclerotized except at 
junction of bursa and ductus bursae; ductus bursae 
sclerotized except for narrow space at middle, broad- 
ened at genital opening. 

A North American genus close to both Pima and 
Interjectio, distinguished from both by its palpi, weakly 
haired sacculus, single strong cornutus, and the inter- 
rupted sclerotization of ductus bursae. The known 
species occur only in the western parts of the United 
States and Canada. 


223. Ambesa laetella Grote 
Fieurus 18, 310, 790 


Ambesa laetella Grote, N. Amer. Ent., vol. 1, p. 98, 1880.—Hulst, 
Phycitidae of N. Amer., p. 141, 1890.—Ragonot, Mono- 
graph, pt. 1, p. 240, 1893.—McDunnough, Check list, No. 
6156, 1939. 

A brilliant, brightly colored species with clearly 
marked pattern; forewing markings a rich red-brown 
against a surrounding suffusion of ashy white; on mid- 
costa a broad, elongate red-brown patch extending from 
outer margin of antemedial line to end of cell and from 
costal edge well into cell, shading into a blackish line on 
its lower margin, and bordered on its outer edge by an 
angled black discal mark formed of the fused discal 
spots; upper half of basal area, median area from the 
brown midcostal patch to vein 1b, and most of the area 
beyond as far as subterminal line, ash white; a brownish 
suffusion, paler than the costal patch, along inner mar- 
gin below vein 1b, on some specimens intensified into a 
constrastingly darkened patch above middle of inner 


margin; the transverse lines well separated, clearly 
marked ; antemedial line narrow, angled, white, bordered 
outwardly by a narrow blackish brown line; subterminal 
line sinuate, narrow, white, preceded and followed by 
blackish brown costal spots, the inner one continued as 
a brown inner bordering line, the outer expanding below 
into a pale brownish suffusion filling tornal area; sub- 
apical area dusted with white; terminal dots fused into 
a black line along terminal edge. Hind wing pale 
smoky fuscous with a very faint yellowish tint. Alar 
expanse, 28-30 mm. 

Male genitalia with apical process of gnathos con- 
tinued below into a pair of narrow, converging, sclero- 
tized lobes. ‘Transtilla absent. Harpe with cucullus 
narrow; sclerotized costa terminating im a short, sharp 
point at apex; an erect clasper arising from below costa 
near base. ‘Terminal margin of vinculum narrowly 
rounded. Highth abdominal segment of male with 
paired tufts. 

Female genitalia with terminal, sclerotized portion 
of ductus bursae abruptly widened and transversely 
wrinkled toward genital opening. Highth-segment col- 
lar narrowly sclerotized. 

Typs Locatity: Colorado (type in BM). 

_ Foop prant: Unknown. 

Disrrrpurion: Unirep States: Arizona, White Mts. 
(July, Aug.), Williams; Colorado, Fort Collins (Aug.), 
Glenwood Springs (Aug.); Montana, Bozeman (Aug.), 
Cut Bank (July); Utah, Provo, ‘‘South Utah” (July); 
Nevada, Verdi (June); California, Inyo County (June), 
San Bernardino Mts. (July, Aug.), Sierra Nevada Mts.; 
Washington, Pullman (June, July, Aug.). CANADA: 
Manitoba, Aweme (Aug.); Alberta, Calgary (July). 

An easily recognized species and one of the most 
beautiful of the American Phycitidae. 


224, Ambesa walsinghami (Ragonot) 
Figures 312, 791, 792 


Pristophora walsingham: Ragonot, N. Amer. Phycitidae, p. 6, 
1887. 

Ambesa walsinghami (Ragonot) Hulst, Phycitidae of N. Amer., 
p. 142, 1890.—Ragonot, Monograph, pt. 1, p. 239, 1893.— 
McDunnough, Check list, No. 6153, 1939. 

Ambesa monodon Dyar, Ins. Insc. Menstr., vol. 1, p. 34, 1913.— 
MeDunnough, Check list, No. 6154, 1939. (New synonymy.) 

Forewing blackish gray faintly tinted with purplish 
fuscous over the lower half of the wing; on costal half 
from base to antemedial line, white faintly streaked 
along the veins with black, the outer margin of the 
whitish area oblique from lower angle of cell to costa. 
near beginning of subterminal line and, on some fresh 
specimens, bordered outwardly by a transverse blackish 
darkening of the ground color; some further ashy white 
dusting in the subapical area beyond the subterminal 
line; transverse lines well contrasted; the white ante- 
medial line distinct only from inner margin to cell, 
concave to lower fold thence inwardly oblique to the 
cell, beyond which it is lost in the white dusting above, 
its outer black margin begins as a black, oblique streak 
from costa and continues outwardly as a fine black line 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 109 


along upper edge of cell as far as discocellular vein, 
whence it loops backward along the lower vein of cell 
and thence along outer edge of the white line to inner 
margin; subterminal line, parallel with termen, sinuate, 
white, bordered inwardly by a black line and outwardly 
by a narrow dark shade, both borders enlarged at 
costa into contrasted black spots; discal dots not dis- 
tinguishable; along termen a row of more or less con- 
fluent black dots. Hind wings varying from smoky 
white to pale brownish, the veins faintly darkened. 
Alar expanse, 19-28 mm. 

Male genitalia with uncas broadly elongate, the side 
margins parallel. Apical process of gnathos without 
pendant lobes. Transtilla present, incomplete. Harpe 
with cucullus moderately broad; apex of sclerotized 
costa produced into a long point at upper (outer) angle; 
no erect clasper. Eighth abdominal segment of male 
without tufts. 

Female genitalia with terminal sclerotized portion of 
ductus bursae gradually broadened to genital opening, 
its terminal margin deeply concaved. LHighth-segment 
collar broadly sclerotized, ventrally fused. 

Tyre Locauiries: California (walsinghami, in Paris 
Mus.); Stockton, Utah (monodon, in USNM). 

Foop piant: Prunus virginiana melanocarpa. 

Disrrisution: California, Cloverdale (June), Deer 
Park Spring (Lake Tahoe), Plumas County (July, Aug.) ; 
Utah, Bellevue (May), Eureka (June, July), Provo 
(July), Stockton (July); Washington, Kamiack Butte 
(May). 

Dyar’s type of monodon is a small female whose 
genitalia (fig. 792) show minor differences from typical 
California specimens, but a series from Utah shows all 
intergradations between the extremes displayed in the 
figures. The food plant record is from a series reared 
by J. F. G. Clarke in 1934 at Kamiack Butte, Wash., 
which I then identified as mirabella Dyar. Their 
abdomens are distinctly gray and their hind wings (es- 
pecially those of the females) are brownish. In exam- 
ples of typical walsinghami from the coastal region of 
California (Cloverdale) the hind wings are whitish and 
the abdomens gray or grayish ocherous. 


225. Ambesa walsinghami mirabella Dyar, new status 
Fieurts 313, 793 


Ambesa mirabella Dyar, Proce. Ent. Soc. Washington, vol. 10, 
p. 59, 1908.—Essig, Insects of western North America, p. 
709, 1826.—McDunnough, Check list, No. 6155, 1939. 

Not distinguishable from typical walsinghami in color 
or maculation of forewing. The hind wing of a dis- 
tinctly ocherous tint and the abdomen ocherous over 

the entire upper surface. Alar expanse, 25-28 mm. 

The male genitalia show only a trifling difference 
from those of typical walsinghami in forking of apex of 
costa (fig. 313). Female genitalia with a shallower con- 
cavity in terminal margin of the projecting ductus bur- 
sae at genital opening. Figure 793 shows the extreme of 
reduction in the concavity. Other examples of mira- 
bella show intergrading approaches to the deep con- 
cavity of walsinghami. 


TYPE Locauity: San Diego, Calif. (type in USNM). 

Foon piant: “Plum.” 

Distrisution: California, Atascadero (San Luis 
Obispo County, July), Camp Baldy (San Bernardino 
Mts., July), Cloudburst Canyon (Los Angeles County, 
July), Mount Lowe (July), Pasadena, Pine Valley (San 
Diego County), San Diego (July). According to Essig 
(1926) the larvae were taken in large numbers on prune 
trees at Hopeland, Calif. 


The name mirabella represents, at most, only a south- 
ern California race of walsinghami. 


226. Ambesa lallatalis (Hulst) 
Fiaures 311, 789 


Neophopteryz lallatalis Hulst, Trans. Amer. Ent. Soc., vol. 13, 
p. 161, 1886. 

Ambesa lallatalis (Hulst) Phycitidae of N. Amer., p. 142, 1890 
(part). 

Forewing white heavily dusted with blackish giving 
the entire wing a pale ashy gray appearance; markings 
as in walsinghami but much fainter; the transverse lines 
obscure; antemedial line indicated by its broken outer 
blackish border which has the same outer loop over the 
cell so characteristic of walsinghami but much fainter 
and often interrupted; on better marked examples a 
whitish crescent on inner margin indicates the base of 
the normal antemedial line; beyond this, dark lines 
extend along vein 1b and the edge of inner margin as 
far as base of subterminal line, defining a narrow oval 
patch along inner margin; subterminal line sinuate, de- 
fined by its black, dentate inner border, the latter inter- 
rupted by a rather broad pale shade extending along 
outer half of lower fold; the outer area (beyond sub- 
terminal line) and the costal area at base also paler than 
remainder of wing; a thin blackish line along outer mar- 
gin. Hind wing white to pale smoky brown. Eighth 
abdominal segment of male without tufts. Alar ex- 
panse, 26-30 mm. 

Male genitalia with apical process of gnathos a simple, 
narrow, moderately long hook. Transtilla absent. 
Harpe without clasper; cucullus moderately broad; 
sclerotized costa terminating in a very shortly project- 
ing point at apex. Anellus a narrow, elongate, plate. 
Cornutus almost as long as aedeagus. Vinculum with 
terminal margin broad. 


Female genitalia with terminal margin of ductus 
bursae at genital margin scobinate, convex, slightly 
notched in the middle (giving the projecting lower sur- 
face of the ductus a bilobed appearance). Highth-seg- 
ment collar broadly sclerotized, wrinkled at lateral mar- 
gins, not ventrally fused. 

Typr tocauity: Nevada (type in AMNH, ex Rut- 
gers). 

Foop piant: Unknown. 

In addition to the female type in the Rutgers Collec- 
tion, I have seen only four other authentic specimens, a 
series of three males and one female from Bellevue, 
Washington County, Utah, in the National Collection, 
collected by G. P. Engelhardt, June 21, 1917. The 


110 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


genitalia of the Utah female agree in every detail with 
those of the type. 

The Ragonot and Dyar references to Jallatalis have 
been omitted from the above synomy as they apply to 
other species. For comments on the misapplication of 
Hulst’s name see under Interjectio denticulella (p. 106) 
and Phobus brucei (p. 138). 


58. Genus Catastia Hitibner 


Catastia Hiibner, Verzeichniss bekannter Schmetterlinge, p. 372, 
1825.—Heinemann, Die Schmetterlinge Deutschlands und 
der Schweiz, Abt. 2, vol. 1, pt. 2, p. 164, 1865.—Ragonot, 
Monograph, pt. 1, p. 479, 1893.—Spuler, Die Schmetterlinge 
Europas, vol. 2, p. 210, 1910.—Hemming, Hubner, vol. 2, p. 
168, 1937.—Janse, Journ. Ent. Soc. South Africa, vol 7, p. 
13, 1944 (description and figures). (Type of genus: Noctua 
marginea Schiffermiiller; figs. 314, 796.) 

Tongue well developed. Antenna finely pubescent; 
on male with a shallow sinus in base of shaft containing 
a row of minute black teeth and a short, weak scale 
tuft. Labial palpus oblique, not extending above 
vertex; broadly scaled, the scales tightly appressed 
(except on ineorruscella and actualis); third segment 
over one-third the length of second,‘ usually projected 
forward and partially concealed in the scaling of second 
segment, acuminate. Maxillary palpus squamous 
(broadly scaled). Forewing smooth; 11 veins; vein 2 
from before lower outer angle of cell; 3 from the angle, 
closer to 4 than to 2; 4 and 5 separated at base; 6 from 
below upper angle of cell, straight; 8 and 9 stalked for 
nearly half their lengths; 10 from the cell; male without 
costal fold. Hind wing as in Ambesa (except that in 
the European marginea vein 3 is slightly longer in 
proportion to 2; this species barely coming within our 
venational group B). Eighth abdominal segment of 
male with a pair of ventrolateral hair tufts. 

Male genitalic characters as in Ambesa except: 
Harpe not so long in proportion to combined length of 
tegumen and uncus; cucullus narrow on all species. 
Transtilla incomplete, but its elements more strongly 
sclerotized. Vinculum as long or a trifle longer (mar- 
ginea) than greatest width, stout. 

Female genitalia with lower surface of ductus bursae 
towards genital opening unsclerotized or very weakly 
sclerotized (except in marginea); a pair of narrow, 
elongate plates on inner dorsal surface of ductus bursae 
at genital opening; otherwise as in Ambesa. 

This genus is very close to Ambesa, differing chiefly 
in its shorter labial palpus with deflected third segment; 
its squamous maxillary palpus; stronger sclerotization 
of the elements of transtilla; the presence of the two 
elongate sclerotized plates in the ductus bursae towards 
genital opening; and the slight scale tuft in the sinus 
of the male antennal shaft. 

The European type of the genus (figs. 314, 796) dif- 
fers from our American species in having the lower 
surface of the ductus bursae sclerotized and produced 


“4 Denuded example of palps of the type species (marginea) 
show the third segment half again as long as the figure (52b) in 
Janse’s 1944 paper. 


at genital opening, the pointed projection from apex 
of costa a trifle longer, the vinculum narrowly rounded 
at its extremity, and vein 3 of hind wing a trifle longer 
in relation to 2. These differences, however, are more 
specific than generic in character and do not seem to 
justify a separate generic designation for our American 
species, despite the obvious likenesses in structure. 
The life history of none of the species is known. 


227. Catastia bistriatella (Hulst), new combination 
Figures 316, 797 


Pyla bistriatella Hulst, Canadian Ent., vol. 27, p. 54, 1895.— 
McDunnough, Check list, No. 6238, 1939. 

Head, thorax, and forewing black with a deep greenish 
blue iridescence; the forewing crossed by two rather 
broad white lines; the antemedial line slightly oblique, 
nearly straight; the subterminal line set well back from 
termen, somewhat wavy. Hind wing a uniform, glossy 
black-brown. Labial palpus not reaching to vertex 
(shorter than the palpi of the other species of the 
genus) ; second segment broadly scaled, the scales flatly 
appressed ; third segment shorter than that of any other 
species of the genus, almost completely hidden in the 
scaling of second segment. Alar expanse, 23-25 mm. 

Male genitalia with projecting spine at apex of 
sclerotized costa very short. Terminal margin of 
vinculum moderately broad, slightly produced at the 
lateral edges, very slightly convex. Female genitalia 
with ventral surface of ductus bursae at genital opening 
not sclerotized; bursa copulatrix with an irregular, 
lined, weakly sclerotized patch towards anterior end 
(probably an individual rather than a specific charac- 
ter). 

TYPE LOcALITy: Yosemite, Calif. (type in AMNH, 
ex Rutgers). 

Foop prant: Unknown. 

Distrizution: California, Humphreys Basin (Fresno 
County, Aug.), Yosemite. 

A striking, easily recognized species. Hulst is in 
error in stating that the maxillary palpi are not scale 
tufted. They are squamous like those of the other 
cogeneric species but are difficult to see behind the 
heavily scaled labial palpi. The genitalia of the male 
type agree in every detail with those of the Humphreys 
Basin male figured. 


228. Catastia incorruscella (Hulst), new combination 
Figures 315, 795 
Pyla incorruscella Hulst, Canadian Ent., vol. 27, p. 55, 1895.— 
McDunnough, Check list, No. 6240, 1939. 

Fore and hind wings a deep, blackish brown, some- 
what lustrous but without metallic iridescence; trans- 
verse lines similar to those of bistriatella, but thinner and 
a dull ocherous white. On thorax and palpi a scatter- 
ing of whitish scales; the scaling on second segment of 
labial palpus slightly roughened; third segment about 
the same proportional length as on marginea and actu- 
alis, longer than that of bistriatella. Alar expanse, 20- 
22 mm. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE Gla! 


Male genitalia differ from those of bistriatella only in 
insignificant details. Female genitalia having bursa 
without sclerotized patch. However, a sclerotized 
patch similar to that shown for bistriatella (fig. 797) is 
present in the bursa of a female in the National Collec- 
tion from Slate Peak, Wash. On this specimen there 
is also a weak sclerotization of the ventral surface of the 
ductus bursae at genital opening. 


Type Locauity: Colorado (type in AMNH, ex Rut- 
gers). 

Foop prant: Unknown. 

Distrisution: Colorado; Arizona, White Mountains 
(June); Washington, Slate Peak (Okanogan County, 
Aug.). 

I have seen no Colorado examples except the male 
type, but the Arizona locality is represented in the 
National Collection by a series of males and females in 
excellent condition (collected by Grace M. and John L. 
Sperry at Colter’s Ranch in the White Mountains, June 
17-18, 1937). Their male genitalia are like those of the 
type. 

229. Catastia actualis (Hulst), new combination 
Ficures 317, 794 


Nephopteryx actualis Hulst, Trans. Amer. Ent. Soc., vol. 13, p. 
161, 1886. 

Dioryctria actualis (Hulst), Phycitidae of N. Amer., p. 135, 
1890.—Ragonot, Monograph, pt. 1, p. 203, 1893.—MeDun- 
nough, Check list, No. 6132, 1939. 

Monoptilota actualis (Hulst), U. S. Nat. Mus. Bull. 52, p. 421, 
1902 (this combination the result of an accidental misplace- 
ment). 

Forewing with basal and outer areas ocherous, the 
basal area more or less dusted with ashy fuscous, espe- 
cially towards inner margin, the outer area with some 
blackish streaks on the veins; median area (between 
the transverse lines) ashy fuscous, the whitish dusting 
concentrated into a pale suffusion transversely across 
the wing from costa before subterminal line to or almost 
to inner margin at base of antemedial line; antemedial 
line white, oblique, notched below cell and more or less 
dentate above; bordered outwardly by a diffused black- 
ish smudge at costa and below cell by a blackish line, 
and preceded on inner margin by a black patch (except 
on California examples); a blackish spot precedes and 
one usually follows the sinuate subterminal white line 
on costa, the inner costal spot continuing as a blackish 
bordering line to inner margin; discal dots distinct, 
separated, black; a row of blackish dots along termen. 
Hind wing ocherous brown, darker brown on most fe- 
males; a dark line along termen and some darkening of 
the veins. Alar expanse, 24-28 mm. 

Male genitalia differing only in insignificant details 
from those of bistriatella and incorruscella. 

Female genitalia with sclerotized wrinklings of bursa 
more extended than in other species of the genus; duc- 
tus bursae weakly granulate towards genital opening 
(differences of little or no significance). 

Type tocauity: Colorado (type in AMNH, ex Rut- 
gers). 

Foop piant: Unknown. 


DistrisuTion: Unirep Strats: Colorado, Denver 
(June), Florissant (July), Gunnison County (near Al- 
mont, June, July), Platt Canyon (July), no specific lo- 
cality (June) ; Nevada; Secret Pass (Elko County, June); 
California, Deer Park Springs (Lake Tahoe, July), Tuo- 
lumne Meadows (July); Washington, Easton, Walla 
Walla (June). Canapva: Manitoba, Aweme (July); 
British Columbia, Blue Lake (west of Lytton, Aug.). 

The foregoing description was drawn from typical 
Colorado examples. The British Columbia and Wash- 
ington specimens before me are much darker, the ocher- 
ous coloration of forewing replaced by deep brown and 
the general color of the wing a suffused blackish fuscous 
with only the white transverse lines and some whitish 
dusting in the median area contrasted; the hind wings 
deep brown to blackish brown with no ocherous tinting. 
This is merely a color form, not a race, and probably 
represents nothing more than individual response to a 
moist condition. The Nevada record, cited above, is 
from a single male in the Canadian National Collection 
(Grace H. and John L. Sperry, collectors). It also rep- 
resents a divergent color form with very dark brown 
hind wing, a dark ground color on forewing and strongly 
marked, white, transverse lines. The type in the Rut- 
gers Collection is a male. Its genitalia agree in every 
detail with those of our pale and dark forms. 


Genera 59-64: Immyrla to Quasisalebria 


[Venational division B. Veins 4 and 5 of forewing slightly 
separated at base, in Oreana connate or, rarely, very shortly 
stalked; vein 6 straight. Hind wing with veins 7-8 approximate 
or very shortly anastomosed beyond cell. Antenna of male with 
sinus and scale tuft at base of shaft. Labial palpus erect or 
oblique, smooth scaled; on male second segment grooved to hold 
maxillary palpus. Maxillary palpus of male in the form of an 
aigrette. Male genitalia with transtilla absent, or incomplete 
and with its elements rudimentary; penis normally armed with 
a single strong cornutus (except Oreana which has several cornuti 
and Quasisalebria which has none). Female genitalia with bursa 
more or less finely and densely spined or scobinate.] 


59. Genus Immyrla Dyar 


Immyrla Dyar, Journ. New York Ent. Soc., vol. 14, p. 108, 
1906.—Forbes, Cornell Mem. 68, p. 627, 1923. (Type of 
genus: Immyrla nigrovittella Dyar.) 

Tongue well developed. Antenna pubescent; a sinus 
and large-scale tuft on base of shaft of male. Labial 
palpus erect; reaching above vertex; smoothly scaled; 
second segment on male grooved to hold the maxillary 
palpus, appressed to face; third segment minute (less 
than one-fifth the length of second and hidden in the 
scaling of the latter on male, a trifle longer and partially 
exposed on female), acuminate. Maxillary palpus of 
male in the form of an aigrette; of female squamous. 
Forewing with subbasal scale ridge; 11 veins; vein 2 
from before lower outer angle of cell; 3 from the angle, 
closer to 4 than to 2; 4 and 5 approximate for a short 
distance from their bases; 6 from below upper angle of 
cell, straight; 8 and 9 stalked for over half their lengths; 
10 from the cell, approximate for a short distance to the 


112 


stalk of 8-9; male without costal fold. Hind wing with 
vein 2 from before lower outer angle of cell; 3 from 4 well 
beyond 2, in some specimens (fig. 19) connected with 
discocellular by a short spur before its separation from 
4; 4 and 5 anastomosed for less than half their lengths; 
7 and 8 anastomosed for a short distance beyond cell; 
cell less than half the length of wing; discocellular vein 
curved and considerably extended at lower angle. 
Highth abdominal segment of male with compound 
ventral scale tufts. 

Male genitalia as in Catastia except transtilla com- 
pletely absent, a fine brush of long hairs arismg from 
inner surface of harpe along lower edge of basal half of 
the sclerotized costa (as in Meroptera). 

Female genitalia with bursa copulatrix finely and 
densely spined; ductus bursae flattened, granulated, 
inbent at middle; not longer than bursa, its lateral 
margins strongly and broadly sclerotized towards 
genital opening, opening deeply concave; ductus semi- 
nalis from a thickened (but not sclerotized) lobe of 
bursa, near junction of bursa and ductus bursae. 

This and the five genera following form a group 
closely allied to Salebria Hiibner. All have grooved 
and erect or oblique male labial palpi, the male maxil- 
lary palpus in the form of an aigrette, a scale tuft in 
sinus on base of shaft of male antenna, no transtilla or 
only the greatly reduced rudiments of one, and (except 
for Oreana and Quasisalebria) a single, long, strong 
cornutus on penis. Jmmyrla is distinguished from all 
nearly related genera by the raised scale ridge on fore- 
wing. Ortholepis also has this character, but the costal 
sclerotization of its harpe is much weaker, and it has a 
complete transtilla. 


230. Immyrla nigrovittella Dyar 
Figures 19, 318, 798 


Immyrla nigrovitiella Dyar, Journ. New York Ent. Soc., vol. 14, 
p. 109, 1906.—Forbes, Cornell Mem. 68, p. 627, 1923.— 
MecDunnough, Check list, No. 6187, 1989. 

Forewing dark gray, the basal area darker than re- 
mainder of wing; median and outer areas a paler 
brownish gray with a faintly darker shade preceding 
the subterminal line, a very sparse dusting of whitish 
scales on the paler areas; antemedial line faint, narrow, 
oblique and more or less curved, dull white, followed 
by a narrow, blackish brown border and preceded by a 
conspicuous black ridge of raised scales extending from 
inner margin to top of cell; subterminal line obscure, 
sinuate, pale, without dark bordering lines; discal spots 
faint, blackish and confluent, forming a curved line 
along discocellular vein. Hind wing pale fuscous, the 
veins very slightly darkened. Both fore and hind 
wings have a rather slick, glossy finish. Alar expanse, 
20-21 mm. 

Genitauia: As given for the genus. 

Type Locatity: Pittsburgh, Pa. (type in USNM). 

Foop prant: Hickory. 

Disrrizution: Pennsylvania, New Brighton (June), 
Pittsburgh (May); New York, Ilion (June). 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


The only species so far discovered referable to the 
genus. The food plant record is from a female without 
a locality label in the National Collection, reared from a 
pupa. ‘The label reads simply ‘102, Hickory, pupated 
ViI—17.” The specimen also bears an identification 
label in Dyar’s handwriting. 


60. Genus Oreana Hulst 


Oreana Hulst, Ent. Amer., vol. 4, p. 115, 1888. 
Dioryctria unicolorella Hulst.) 


(Type of genus: 


Characters as in Immyrla except: Forewing smooth; 
veins 4 and 5 connate (rarely very shortly stalked) ; vein 
10 from the stalk of 8-9 a short distance from cell. 
Hind wing with 4 and 5 stalked for more than half their 
lengths. 

Male genitalia with numerous strong slender cornuti 
on penis. 

Female genitalia with bursa copulatrix strongly 
sclerotized in the lobed area giving off the ductus semi- 
nalis; sclerotized lateral margins of ductus bursae not 
produced at genital opening and ventral margin of the 
opening not appreciably concave (these differences in 
the ductus bursae probably only of specific significance). 

Oreana sank into the synonymy of Meroptera when 
Ragonot (1889) referred its type species (wnicolorella) to 
the latter genus, but it must be restored, as wnicolorella 
is not a Meroptera on genitalic characters. Its chain of 
numerous, strong cornuti distinguish it from any species 
in Meroptera or the American genera, closely allied to 
Salebria. Oreana, on most characters, seems nearest to 
Immyrla, from which it is at once distinguished by its 
smooth forewing. It contains but the one described 
American species. 


231. Oreana unicolorella (Hulst) 
Fieures 319, 788 


Dioryciria unicolorella Hulst, Ent. Amer., vol. 3, p. 136, 1887. 

Oreana unicolorella (Hulst), Ent. Amer., vol. 4, p. 115, 1888. 

Meroptera unicolorella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 
115, 1889; Monograph, pt. 1, p. 315, 1893.—Hulst, Phy- 
citidae of N. Amer., p. 149, 1890.—Forbes, Cornell Mem. 
68, p. 624, 1923. 

Myelois leucophaeella Hulst, Canadian Ent., vol. 24, p. 60, 1892. 

Meroptera leucophaeella (Hulst), Barnes and McDunnough, Con- 
tributions, vol. 3, p. 194, 1916. 

Meroptera nebulella McDunnough (not Riley), Check list, No. 
6185, 1939. 


Forewing mouse gray, the basal area slightly paler 
than remainder of wing; transverse lines grayish white, 
obscure; antemedial line oblique, notched below cell, 
bordered outwardly from costa for a short distance by an 
obscure blackish shade; subterminal line sinuate, with- 
out appreciable dark borders; discal dots faint, brown, 
usually separated but sometimes confluent; on underside 
of male forewing a streak of blackish sex scaling along 
basal third of costa. Hind wing pale smoky brown, the 
veins slightly darkened. Alar expanse, 18-22 mm. 

Male genitalia with harpe, tegumen, and vinculum 
similar to those of Immyrla nigrovittella. Uncus some- 
what smaller in proportion; in natural position inclined 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


downward, but when flattened out it shows an appre- 
ciable constriction at middle. Anellus a simple shield- 
shaped plate. Female genitalia as given for the genus; 
spining of bursa not so dense or continuous as in 
nigrovittella. 

Type tocauities: Washington, D. C. [sic] (unicol- 
orella, in AMNH, ex Rutgers); Iowa (leucophaeella, in 
AMNH, ex Rutgers). 

Foop piant: Apple (record from female in National 
Collection labeled “bred from apple, emerged 25—V— 
1904, Ottawa, J. Fletcher’). 

Distrisution: Unirep Sratres: Maine, Augusta 
(June); New Hampshire, Dublin; Connecticut, Hast 
River (July); New Jersey, Montclair (July); Pennsyl- 
vania, New Brighton (May, June, July, Aug.), Oak 
Station (June), Pittsburgh (June, July) ; Iowa; Colorado; 
Oregon, Milton (July); Washington, Pullman (June). 
Canapa: Ontario, Ottawa (May), Trenton (July); Que- 
bec, Montreal (July), St. Hilaire (July); Nova Scotia, 
Cape Breton Isl. (July). 

There is some mislabeling of the types or, what is more 
likely, errors in the citation of type localities by Hulst. 
In his original description of unicolorella he cites ““Wash- 
ington, D. C.” and in his 1890 paper gives ‘‘Canada.” 
His type is a male (without abdomen) from Iowa, 
labeled: “HH. S. Sanders, June 13, 1886.” The type of 
leucophaeella is a female with typical genitalia, labeled 
“Colo., Gillette.”” The type locality citation of “Iowa” 
in the original description can be written off as another 
Hulst lapsus, although the species occurs there and is 
represented by several examples in the National Collec- 
tion. Barnes and McDunnough (1916) first put leuco- 
phaeella into synonymy with unicolorella, and there does 
not seem to be any reason to doubt the correctness of 
that procedure. The two specimens at Rutgers labeled 
“type” by Hulst certainly represent one and the same 
species. 


61. Olybria, new genus 


Type or Genus: Myelois aliculella Hulst. 


Tongue well developed. Antenna finely and densely 
pubescent; on male a sinus and scale tuft in shaft at 
base. Labial palpus oblique, reaching well above vertex; 
second segment of male grooved to hold the maxillary 
palpus; third segment short, about one-third the length 
of second, acuminate, partially concealed by scaling of 
second segment. Maxillary palpus of male in the form 
of an aigrette; of female small, squamous. Forewing 
smooth; 11 veins; vein 2 from before but near lower 
outer angle of cell; 2, 3, and 4 equidistant at base; 4 and 
5 shortly separated at base and thence approximate 
(parallel) for a very short distance; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for about half 
their lengths; 10 from the cell, shortly separated from 
the stalk of 8-9 at base; male without costal fold. Hind 
wing with vein 2 from well before lower outer angle of 
cell; 3 from the angle, connate with the stalk of 4-5; 
4 and 5 stalked for half their lengths; 7 and 8 closely 
approximate for half their lengths beyond cell; cell 


113 


slightly less than half the length of wing; discocellular 
vein curved, considerably extended at lower angle. 
Eighth abdominal segment of male with a pair of thin, 
weak, ventrolateral hair tufts. 


Male genitalia with uncus small, hoodlike, projected 
downward (at right angle to tegumen). Apical process 
of gnathos a short, stout, curved hook. Tegumen with 
a pair of straight, strongly sclerotized arms projecting 
backward from its lower, posterior angles. Transtilla 
absent. Harpe with costa strongly sclerotized through- 
out its length and produced at apex into a sharp spine; 
cucullus narrowly elongate, tapering to pointed apex. 
Anellus a broadly U-shaped band, supplemented (in 
JSurciferella) by some sclerotization of the membranous 
tube surrounding the aedeagus. Aedeagus rather long 
and stout; penis armed with a single long, strongly 
sclerotized, rather slender cornutus. Vinculum stout, 
longer than greatest width, narrowly truncate at 
terminal margin; the latter very slightly concave. 


Female genitalia with bursa smoothly sclerotized over 
most of dorsal surface, its ventral surface finely sco- 
binate with contorted, wrinkled and spined bands more 
or less encircling the bursa at junction of bursa and 
ductus bursae; ductus bursae at least as long as bursa, 
rather broad, flattened (ribbonlike) and waved (twice 
bent), sclerotized throughout, at genital opening the 
sclerotization forming a stout, squarish ventral plate; 
ductus seminalis from a lobe of bursa near junction of 
bursa and ductus bursae. 

This genus is distinguished from its nearest allies of 
the Salebria complex by the projecting arms from 
tegumen of the male genitalia, the ribbonlike, sclerotized 
ductus bursae and heavy, squarish genital plate of the 
female, and the simple, paired tufts of the eighth abdom- 
inal segment of the male. This last character is 
shared by the genus Salebriacus, which separates from 
Olybria on other differences of genitalia and venation. 

Two North American species, referred from Salebria 
(of authors), represent the only known components of 
the genus. 


232. Olybria aliculella (Hulst), new combination 
Figures 320, 786 


Myelois aliculella Hulst, Ent. Amer., vol. 3, p. 135, 1887. 

Salebria oberthuriella Ragonot, N. Amer. Phycitidae, p. 9, 1887. 

Salebria aliculella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 367, 1893.—Hulst, Phycitidae 
of N. Amer., p. 154, 1890.—Barnes and McDunnough, 
Contributions, vol. 3, p. 197, 1916—McDunnough, Check 
list, No. 6217, 1939. 

Forewing white dusted with black, making the general 
color ashy gray, the black dusting concentrated on 
extreme base and in short streaklets on the lower veins 
at termen; antemedial line narrow, oblique, slightly 
notched at vein 1b, white, bordered outwardly by a 
black line which begins on costa as a conspicuous, 
triangular, black spot; on inner margin, preceding the 
antemedial line, a large orange spot; subterminal line 
sinuate, narrow, white, bordered inwardly by a narrow 
black line and outwardly by a broad orange band which 


114 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


ends in a black spot at costa; along lower fold between 
the transverse lines fresh specimens show a faint shading 
of olivaceous ocherous; a similar shade often along 
discocellular vein; completely surrounding the dis- 
cocellular vein a large black ring (obicular). Hind 
wing translucent white with a faint ocherous tint 
especially towards outer margin and anal angle; the 
veins very faintly darkened and a dark line along 
termen. Alar expense, 19-22 mm. 

Male genitalia showing only comparative differences 
to distinguish them from those of furciferella. ‘These 
are shown in the figures. Female genitalia with a row 
of stiff, flattened setae along lower, posterior margin of 
the eighth-segment collar. 

Type LocaLity: Arizona (aliculella, in AMNH, ex 
Rutgers; oberthuriella, in Paris Mus.). 

Foop piuant: Ceanothus ? (Hulst, 1890). 

Distrrpution: Arizona, Kingman (Oct.), White 
Mts. (Aug.), Wilgus, Williams, state locality only 
(June); New Mexico, Albuquerque, Fort Wingate 
(June), Jemez Springs (June, July); Texas, Big Bend 
region (May). 

Easily identified by the orange spot preceding the 
antemedial line and the large black obicular mark on 
the disc of the forewing. 


233. Olybria furciferella (Dyar), new combination 
Ficurts 321, 787 
Salebria furciferella Dyar, Journ. New York Ent. Soc., vol. 12, 
p. 106, 1903.—Barnes and McDunnough, Contributions, 
vol. 3, p. 197, 1916 —McDunnough, Check list, No. 6216, 
1939. 

Sunilar to aliculella except: Forewing more strongly 
dusted with black, making ground color a decided ash 
gray; the obicular spot on disk absent, replaced by a 
couple of black streaks on upper and lower veins at end 
of cell, forking from a black streak extending along the 
upper vein of cell from the black costal dash bordering 
the antemedial white line; the subbasal orange patch 
on inner margin preceding the antemedial line and the 
orange shade following the subterminal line reduced 
and, on some specimens, obscured by black scaling. 
Hind wing as in aliculella. Alar expanse, 21-23 mm. 

Male genitalia similar to those of aliculella except 
projecting arms of tegumen stouter; aedeagus and 
cornutus longer. Female genitalia with a fringe of 
fine hairlike setae along the lower posterior margin of 
eighth-segment collar. 

TypE tocaLity: Ashfork, Ariz. (type in USNM). 

Foop prant: Unknown. 


Distrisution: Arizona, Ashfork and Prescott (June, 
type, o’, and 2 paratypes, 9), also 2 females with only 
the state locality, from the Fernald and Brooklyn 
Museum Collections and originally made cotypes of 
aliculella Hulst. 

The species is easily recognized by the black streak 
along the upper vein of cell and is distinct from alicu- 
lella though obviously very close to it. 


62. Salebriacus, new genus 


Typr or Genus: Nephopteryx odiosella Hulst. 

Characters of Olybria except: Male antenna with 
scale tuft in shallow sinus on base of shaft but greatly 
reduced, a mere vestige. Forewing with vein 2 slightly 
further from 3 at base than 3 is from 4; 8 and 9 stalked 
for considerably more than half their lengths. Paired 
tufts on eighth abdominal segment of male very weak, 
mere vestiges. 

Male genitalia with uncus hoodlike, elongate (longer 
than wide) and not projected downward. Apical 
hooked process of gnathos stout, long, reaching nearly 
as far as apex of uncus. Tegumen simple. Transtilla 
present but incomplete and its elements much reduced. 
Harpe with apex of sclerotized costa produced as a 
stout, free spine before apex of cucullus. Anellus a 
simple shield. Entire genitalia more robust and 
proportionally shorter than those of Olybria or the genus 
Salebriaria, which follows. 

Female genitalia with bursa very broad in proportion 
to its length, not sclerotized, nearly half the inner sur- 
face covered with a dense, spinose mat; ductus bursae 
semitubular, broad and short, partially sclerotized (the 
sclerotization interrupted near middle of the ductus); 
ductus seminalis from near anterior end of bursa. 
Highth-segment collar narrow ventrally and laterally, 
dorsally produced as a spatulate apron. 

This genus is another restriction from Salebria of 
authors. It is distinguished chiefly by the weak tuft 
of the male antenna, the squat, stout, male genitalia; 
the robust, hooked projection of gnathos, the eighth- 
segment collar of the female, and the place of departure 
from bursa copulatrix of the ductus seminalis. 

It contains but one North American species. 


234. Salebriacus odiosellus (Hulst), new combination 
Figures 322, 799 


Nephopteryz odiosella Hulst, Ent. Amer., vol. 3, p. 132, 1887. 

Salebria odiosella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 366, 1893.—Hulst, Phycitidae 
of N. Amer., p. 155, 1890.—Barnes and McDunnough, 
Contributions, vol. 3, p. 197, 1916—MceDunnough, Check 
list, No. 6219, 1939. 

Salebria bakerella Dyar, Journ. New York Ent. Soc., vol. 12, p. 
105, 1904.—McDunnough, Check list, No. 6220, 1939. 
(New synonymy.) 

Salebria yumaella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 35, 1905—McDunnough, Check list No. 6218, 1939. 
(New synonymy.) 

Forewing white dusted with black, making the 
ground color ashy gray, palest over the median area; 
antemedial line narrow, oblique from costa to lower 
margin of cell, thence slightly incurved to lower margin, 
white, margined outwardly on upper half by a rather 
broad black band and inwardly on lower margin by a 
more or less expanded black patch; subterminal line 
sinuate, narrow, white, bordered inwardly and, to a 
lesser extent, outwardly by blackish lines which expand 
into distinct black dashes at costa; discal dots at end 
of cell, separated, usually distinct, blackish; on most 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


specimens a diffuse oblique shade from upper part of 
subterminal line obliquely across wing toward inner 
margin, and on a few of the more contrastingly marked 
specimens a faint ocherous shading along the lower 
fold; terminal dots along outer margin more or less 
confluent, obscure on many specimens. Hind wing 
white, translucent; faintly shaded with pale brown at 
apex and along outer margin for a short distance from 
apex. Alar expanse, 19-26 mm. 

Male genitalia with vinculum rather abruptly nar- 
rowed from middle to a truncate terminal margin. 
Female genitalia with sclerotized portion of ductus 
bursae at genital opening produced and broadened, its 
terminal margin and lateral angles concave. 

Type Locauities: “Colorado” [sic] (odiosellus, in 
AMNH, ex Rutgers). Ormsby County, Nevada (baker- 
ella, in USNM); Yuma County, Ariz. (ywmaella, in 
USNM). 

Foop prant: Unknown. 


Disrrisution: Teras, Blanco County [?]; Arizona, 
Mohave County (July, Sept.), Yavapai County, Yuma 
County; Colorado; Utah, Bellevue (May, June, July); 
Nevada, Ormsby County (July), Pyramid Lake, Reno; 
California, Jacumba (May), Mexican Wells (Clarke 
Mts., Sept.), Morongo Valley (May). 

The types of odiosellus, bakerella, and yumaella are all 
males with identical genitalia. That of yumaella is a 
small, somewhat suffused specimen with the black 
markings less strongly contrasted than those of typical 
Nevada examples. In our series there is a complete 
intergradation between the extreme forms. 

Barnes and McDunnough (1916) called attention to 
the confused citations of the type locality of odiosellus. 
Hulst gives Colorado in his original description, but in 
his 1893 paper cites ‘“‘Texas.”” The type at Rutgers is 
labeled ‘‘Blanco Co., Texas.’’ I suspect that this is a 
mislabeling, for Colorado seems a more likely locality 
than central Texas. In my paper on the cactus-feeding 
Phycitinae (Proc. U.S. Nat. Mus., vol. 86, p. 389, 1939) 
I misapplied the Hulst name to a species described by 
Dyar as Ozamia clarefacta. This error is discussed 
further under the treatment of clarefacta (p. 258). 


63. Salebriaria, new genus 


Typsz or Genus: Salebria ademptandella Dyar. 

Tongue well developed. Maleantenna pubescent with 
strong scale tuft in sinus at base of shaft. Labial palpus 
obliquely upturned, reaching above vertex, smoothly 
scaled; second segment on male grooved to hold the 
maxillary palpus, appressed to face; third segment mi- 
nute and hidden in scaling of second on male, somewhat 
longer and partially exposed on female, acuminate. 
Maxillary palpus of male in the form of an aigrette; of 
female subsquamous. Forewing smooth; 11 veins; vein 
2 from before but near lower outer angle of cell; 3 
usually somewhat nearer to 4 than to 2, sometimes equi- 
distant from them; 4 and 5 shortly separated at base, 
rarely (in smaller specimens) closely approximate; 6 
from below upper angle of cell, straight; 8 and 9 stalked 


115 


for well over half their lengths (except in fructetella) ; 
male without costal fold; on underside of male wing a 
streak of appressed, black sex-scaling along basal third 
of costa. Hind wing with vein 2 from before, but 
rather near lower outer angle of cell; 3 from the angle, 
connate with the stalk of 4—5; 4 and 5 stalked for approx- 
imately half their lengths (for slightly over half in 
Jructetella); 7 and 8 approximate beyond cell (except in 
pumilella and fructetella where they are contiguous or 
weakly anastomosed for a short distance); cell slightly 
less than half the length of wing. Eighth abdominal 
segment of male with compound ventral scale tufts. 


Male genitalia with uncus subtriangulate (apex rather 
broadly rounded). Apical process of gnathos a short 
hook, slender (except on fructetella). Transtilla absent. 
Harpe with apex of sclerotized costa produced as a short 
free spine before apex of cucullus. Anellus a semitubu- 
lar shield. Penis armed with a single, strongly sclero- 
tized, moderately long cornutus. Vinculum stout, sub- 
triangulate, slightly longer than greatest width. 


Female genitalia with much of inner surface of bursa 
covered with a dense matting of fine spines (especially 
towards distal end); bursa also sometimes partially 
sclerotized (pumilella, fructetella) ; ductus bursae shorter 
than bursa, strongly sclerotized (at least towards genital 
opening), partially flattened (ribbonlike in pumuilella); 
ductus seminalis from lobe of bursa adjacent to the 
junction of bursa and ductus bursae. 

This genus is a further restriction from Salebria. It 
is a compact group of very closely related species; in its 
smooth forewings and male genitalia markedly distinct 
from typical Salebria; closest to Salebriacus from which 
it is at once distinguished by the compound scale tufts 
on eighth abdominal segment of the male and the place 
of departure of the ductus seminalis from bursa in the 
female. How many valid species are represented by the 
several names in our North American lists and definitely 
referable to the genus cannot be exactly determined 
until larger series of reared specimens are available. 
The genitalia (except for those of pumilella and fruc- 
tetella) offer little or nothing in the nature of trust- 
worthy specific characters. The maculation and color 
differences on forewings that have been used by previous 
authors seem to be equally unreliable. 


235. Salebriaria turpidella (Ragonot), new combination 
Fiaures 323, 802 


Salebria turpidella Ragonot, Nouv. Gen., p. 19, 1888; Monograph, 
pt. 1, p. 346, 1893.—Forbes, Cornell Mem. 68, p. 625, 
1923.—McDunnough, Check list, No. 6196, 1939. 

Salebria ademptandella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 114, 1908.—McDunnough, Check list, No. 6198, 
1939. (New synonymy.) 

Forewing powdery gray, the ground color variable, 
ranging from pale ash gray (ademptandella) to a more 
suffused pale brownish gray; basal area usually some- 
what paler than median area; the transverse lines but 
little paler than the ground color, indicated chiefly by 
their dark margins; the latter narrow, blackish, well 
contrasted, especially on the paler examples; antemedial 


116 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


line sinuate-angulate, bordered outwardly by a black 
line and inwardly, on lower half, by a similar narrow 
black line; on many specimens a small, faint, whitish 
patch just beyond the antemedial line on inner margin; 
subterminal line sinuate, bordered inwardly by a black- 
ish line, the latter fainter than that bordering the ante- 
medial line; discal spots black, confluent, normally 
forming a black line along the discocellular vein, but on 
individual specimens tending to separation on one or 
the other forewing. Hind wing smoky white to brown, 
variable in both sexes. Alar expanse, 17-18 mm. 

Male genitalia show no distinguishable differences 
from those of typical turpidella, nubiferella, or annulo- 
sella. Female genitalia with the spining on anterior half 
of bursa of a nearly uniform fineness (no dense concen- 
tration of darker spines at the closed end); bursa with- 
out appreciable sclerotization at middle. 


TYPE LOCALITIES: Unitep Srares: (turpidella, 3, 
in Paris Mus.); Kerrville, Tex. (ademptandella, in 
USNM). 

Foop prant: Unknown. 

Disrrisution: Texas, Burnet County (Oct.), Kerr- 
ville (Apr., May, June), Sabinal (Sept.), also one Texas 
male from the Boll Collection (No. 558) labeled 
‘“Kurope”’; Florida, Gotha (F. Rauterberg, collector, 
2 specimens, of and Q). 

The foregoing from typical examples. Also before 
me are transitional examples between turpidella and 
nubiferella from Texas, Blanco County (May, July) 
and Shovel Mountain, and North Carolina, Southern 
Pines (Apr., July, Aug.) and Tryon (May, Sept.). 
These specimens are dark, with the size and markings 
of turpidella but with female genitalia more like those of 
anmulosella. Some of them had been identified as 
annulosella. Others were under turpidella. I doubt 
very much if turpidella is anything more than a variety 
of nubiferella. Dyar’s ademptandella is nothing but a 
paler form of the typical turpidella. 


236. Salebriaria nubiferella (Ragonot), new combination 


Salebria nubiferella Ragonot, N. Amer. Phycitidae, p. 8, 1887; 
Monograph, pt. 1, p. 344, 1893.—Hulst, Phycitidae of N. 
ane p. 150, 1890—McDunnough, Check list, No. 6191, 

Salebria annulosella nubiferella (Ragonot) Forbes, Cornell Mem. 
68, p. 625, 1923. 

If this and the preceding “species” (turpidella) are 
anywise distinct, the name nubiferella will apply to a 
larger form (21 mm.) with the basal area of forewing 
dark and concolorous with the darker shading in median 
and outer areas and with a more strongly contrasted 
white patch on the inner margin just beyond the ante- 
medial line; but neither of these differences, nor those 
used by Ragonot (Monograph, pp. 329, 345) to separate 
turpidella and nubiferella are constant. According to 
Clarke’s notes the species is represented in the Paris 
Museum only by the unique type, a male labeled “‘type 
orig., pl. XIV fig. 23, Amer. Sept. don. C. V. Riley.” I 


have examined its genitalia and they show nothing to 
distinguish them from those of turpidella, annulosella, 
or engelt. 

What appears to be a typical female in the National 
Museum from Putnam County, Ill. (July) has the well- 
contrasted white spot on inner margin, dark hind wings 
and dark gray forewings, the latter with discal dots 
fused and the dark (blackish) borders of antemedial and 
subterminal lines as in turpidella, but somewhat weaker. 
Its alar expanse is 20mm. The female genitalia have a 
rather dense and dark concentration of fine spines at 
the anterior end of bursa and a thickening and slight 
sclerotization of the membrane at middle of bursa, the 
sclerotized part showing what seems to be a fixed longi- 
tudinal fold. In these particulars the genitalia are like 
those of annulosella. 

Type Locauity: “Amer. Sept.’’ [Texas] (type in Paris 
Mus.). 

Foop pranr: Unknown. 

Besides the aforementioned female there are before 
me a male (19 mm.) from Putnam County, Ill. July), 
and a male (18 mm.) and female (19 mm.) from near 
St. Louis, Mo. (June) which appear to be conspecific 
with the Illinois female. They have the white spot on 
inner margin less strongly marked, but the strength of 
this marking does not seem to be of any significance. 
Presumably several of the Texas examples from the 
intermediate specimens mentioned under turpidella 
should be referred here if a specific distinction can be 
maintained between turpidela and nubiferella. We 
shall have to have host plant association and good 
reared series before such a distinction can be made with 
any certainty. 


237. Salebriaria engeli (Dyar), new combination 


Salebria engeli Dyar, Journ. New York Ent. Soc., vol 14, p. 107, 
1906.—Forbes, Cornell Mem. 68, p. 625, 1923.—McDun- 
nough, Check list, No. 6195, 1939. 

Forewing dark brownish gray, the transverse lines 
obscured; antemedial line indicated only by its very 
faint, narrow, angulate, outer black border, followed 
outwardly on inner margin by a strongly contrasted 
white patch; subterminal line faint, but slightly paler 
than the ground color and with an obscure, narrow, dark 
inner border, sinuate; discal dots more or less confluent, 
blackish with some pale scaling on their outer margins; 
a row of separated blackish dots along outer margin. 
Hind wings smoky fuscous, on darker specimens with 
a pale brownish tint; the veins darkened. Alar expanse, 
18-20 mm. 

Genitalia as in annulosella. 

TypE Locauity: Oak Station, Pa. (type in USNM). 

Foop prant: Unknown. 

DistrisutTion: Pennsylvania, Hazleton, New 
Brighton (July), Oak Station (July); Maryland, Plum- 
mers Isl. (July); Illinois, Decatur (May); Tezas, 
Kerrville (May, June, July). 

An intermediate form between typical nubiferella and 
annulosella, probably only a variety of the former. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 117 


238. Salebriaria annulosella (Ragonot), new combination 
Fiaure 800 


Salebria annulosella Ragonot, N. Amer. Phycitidae, p. 7, 1887; 
Monograph, pt. 1. p. 346, 1893.—Forbes, Cornell Mem. 68, 
p. 625, 1923.—MeDunnough, Check list, No. 6197, 1939. 

Salebria robustella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 114, 1908.—McDunnough, Check list, No. 6211, 1939 
(New synonymy.) 

Forewing gray; antemedial line but slightly paler than 
the ground color and not sharply defined, margined on 
outer side at costa by a more or less triangulate blackish 
patch and on inner side at inner margin by a somewhat 
larger blackish patch which extends to or nearly to base 
of wing; discal dots distinctly separated, black, sur- 
rounded by pale dusting; subterminal line obscure; a 
row of distinct blackish dots along terminal margin. 
Alar expanse, 18-20 mm. 

Female genitalia figured from specimen from Burnet 
County, Tex. They are like those of the type in Paris 
and differ in no essential details from those of females 
of nubilella. 

Type LocauitiEs: Texas (annulosella, in Paris Mus.) ; 
Burnet County (robustella, in USNM). 

Foop prant: Unknown. 

Distrisution: Texas, Blanco County (Apr.); Burnet 
County (Apr.); North Carolina, Raleigh (June). 

According to Clarke’s notes, two examples, forming 
the basis of Ragonot’s diagnosis of the species in his 
Monograph, are in the Paris Museum. One is a female 
from Texas, obviously the type but not so labeled. It 
was the specimen figured in the Monograph (pl. 9, fig. 
6). The other specimen is a male labeled: ‘Dallas, 
Texas, Boll, 24-V-689.’’ A female in the U. S. Na- 
tional Museum, also a Boll specimen from Texas (No. 
559) and probably a mate to the Paris male, bears a 
name label (Salebria annulosella) in Ragonot’s hand- 
writing. I doubt that either of these Boll specimens is 
a typical annulosella. Our female in markings is inter- 
mediate between annulosella and tenebrosella with female 
genitalia like the latter species. Dyar’s type of robus- 
tella (a male) matches Ragonot’s figure and description 
of annilosella in all details except that the discal spots 
are obscured on one forewing. A female, obviously the 
other sex of Dyar’s type, had been identified by him as 
annulosella. ‘The remaining Texas examples before me 
(Blanco County) had been identified by Hulst as either 
nubiferella or pumilella. 


239. Salebriaria tenebrosella (Hulst), new combination 
Fieurss 801, 803 


Nephopteryz tenebrosella Hulst, Ent. Amer., vol. 3, p. 131, 1887. 

Nephopteryx quercicolella Ragonot, N. Amer. Phycitidae, p. 7, 
1887. 

Salebria tenebrosella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 347, 1893.—Hulst, Phycitidae 
of N. Amer., p. 151, 1890.—McDunnough, Check list, No. 
6199, 1939. 

Salebria heinrichalis Dyar, Ins. Insc. Menstr., vol. 5, p. 45, 
1917.—McDunnough, Check list, No. 6192, 1989. (New 
synonymy.) 

Forewing dark gray more or less shaded with black, 
especially in basal area; antemedial line whitish, dif- 


fused, oblique, sometimes interrupted at middle, pre- 
ceded by a dark red patch on lower half; cutting this 
red patch an oblique black line which fuses into the 
black basal shade on costal half of basal area; subter- 
minal line obscure, sinuate, very slightly paler than 
the ground color; discal dots coalesced into a black 
lunulate line along discocellular vein, partially obscured 
in the dark ground color over middle of wing but set 
off by some pale (whitish) dusting along its outer edge; 
separated blackish dots along terminal margin. Hind 
wings brownish gray; the veins slightly darkened. 
Alar expanse, 17-18 mm. 

Male genitalia showing no distinguishing characters 
from those of preceding species. 


Type Locatities: Texas (tenebrosella, in AMNH, ex 
Rutgers); ‘“‘America Septentrionalis” (querciocolella, in 
Paris Mus.) ; Falls Church, Va. (heinrichalis, in USNM). 

Foop piant: Oak (larva a leaf tier). 

Distripution: Texas; Missouri (Aug.); Virginia, 
Falls Church (Apr.). 

In addition to typical examples from the above lo- 
calities there is before me a series of males and females 
of a slightly larger average size, 18-20 mm., with a 
somewhat more diffused and contrasted whitish ante- 
medial line and little or no trace of the subbasal red 
patch on forewing and no red on the thorax. On 
typical fenebrosella the tips of the patagia are shaded 
with red. The females of these variant examples also 
have larger genitalia (fig. 803). They may represent a 
food plant race or a color form of tenebrosella but hardly 
anything more. In view of the already obscure specific 
limits of the described species I do not feel justified in 
adding a further name. 

The variety is represented in the National Collection 
from the following localities: Massachusetts, Cohasset 
(July), Martha’s Vineyard (July); New York, Utica 
(Aug.); Maryland, Plummers Isl. (Aug.); District of 
Columbia, Washington (June); North Carolina, Tryon 
(May); Georgia, Atlanta (June); Jilinois, Decatur 
(May), Lacon (June); Missouri, near St. Louis, and one 
small example from the Murtfeldt Collection labeled 
“130 M. apple, 5-8-89.”’ Several of these had been 
previously misidentified as Oreana leucophaeella (Hulst). 


240. Salebriaria pumilella (Ragonot), new combination 
Fiaures 325, 804 


Salebria pumilella Ragonot, N. Amer. Phycitidae, p. 8, 1887; 
Monograph, pt. 1, p. 344, 1893—MceDunnough, Check list, 
No. 6190, 1939. 

Salebria georgiella Hulst, Canadian Ent., vol. 27, p. 57, 1895.— 
McDunnough, Check list, No. 6210, 1939. (New synony- 
my.) 

Forewing dark gray shaded with brown; a rather pale 
chocolate brown, triangulate patch on inner margin on 
inner side of antemedial line; the latter narrow, white, 
its blackish bordering lines broken, and obscure except 
the inner one on lower half of wing; a similar black 
border on the inner edge of the chocolate brown patch; 
on some specimens (especially faded examples) a slight 
ocherous shading at extreme base of wing; on most fresh 


118 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


specimens a narrowly elongate, grayish fuscous patch 
on midcosta; some white dusting on inner margin 
beyond antemedial line and in cell towards its outer 
margin; subterminal line fine, white, its dark borders 
obscure; discal spots confluent, forming a narrow, 
blackish line along discocellular vein; the blackish dots 
along terminal margin weak, more or less confluent. 
Hind wing pale smoky fuscous. Alar expanse, 15-17 
mm. 

Male genitalia with spine from apex of sclerotized 
costa of harpe projecting straight out beyond apex of 
cucullus (not curved dorsally as in the other species). 

Female genitalia with ductus bursae flattened, ribbon- 
like, sclerotized throughout and dorsoventrally folded 
towards bursa copulatrix, its lower margin produced at 
genital opening into a subtriangulate projecting shield 
with somewhat rounded terminal margin; bursa with 
nearly half of one side strongly and smoothly sclero- 
tized. 


TypE LOCALITIES: Texas (pumilella, in Paris Mus.); 
Charlotte Harbor, Fla. (georgiella, mn AMNH, ex Rut- 
gers). 

Foop pLrant: Unknown. 

Distrisution: Florida, Charlotte Harbor (Mar.); 
Texas, Burnet County (Apr.), also 3 examples (co and 
99) with only the state locality and without dates; North 
Carolina, Southern Pines (June, Aug.). 

Hulst’s references to pumilella in his Phycitidae of 
N. Amer. (p. 150) are omitted from the above synonymy, 
since he had misidentified Ragonot’s species. The 
technical description he gives was simply copied from 
Ragonot’s original description. There is no doubt 
about the synonymy of georgiella. 

Tn all examples I have seen, veins 4 and 5 of forewing 
are rather closely approximate. 


241. Salebriaria fructetella (Hulst), new combination 
Ficures 324, 805 


Myelois fructetella Hulst, Canadian Ent., vol. 24, p. 59, 1892. 

Salebria rectistrigella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 115, 1908. 

Salebria fructetella (Hulst) Barnes and McDunnough, Contribu- 
tions, vol. 3, p. 194, 1916—McDunnough, Check list, No. 
6203, 1939. 

Forewing glossy brownish gray shaded with blackish 
brown and dusted with white; the black shading form- 
ing a blotch on costa following, and a similar blotch on 
inner margin preceding the antemedial line, and an 
obscure dark shade extending from costa near apex 
obliquely inward across the subterminal line; white 
dusting limited to a more or less triangulate cloud from 
midcosta surrounding the upper discal spot and a 
smaller cloud on costa preceding the antemedial line; 
antemedial line narrow, slightly oblique, straight or (on 
some examples) with a slight notch at middle, white and 
distinctly contrasted against the ground color but with- 
out appreciable black bordering lines; discal dots sep- 
arated, blackish; subterminal line obscure, whitish gray, 
vertical except for a median outward bulge; terminal 
dots obscure, brown. Hind wing smoky white shading 


to fuscous along termen; veins very slightly darkened. 
Alar expanse, 15-18 mm. 

Male genitalia distinguished from those of other 
species in the genus by the spoon-shaped apical projec- 
tion of gnathos, the V-shape of the sclerotized part of 
anellus, and the longer cornutus. 

Female genitalia with ductus bursae very short, its 
median area membranous, a moderately broad sclero- 
tized and granulate band at genital opening, strongly 
sclerotized and longitudinally ridged towards junction 
with bursa; this peculiar sclerotization extending into 
and occupying nearly half of the bursa, anterior (closed) 
end of bursa fused into a thickened (but not sclerotized) 
membrane; remainder of bursa covered by a mat of 
fine spines. 

TyPE LocALitius: Blanco County, Tex. (fructetella, 
in AMNH, ex Rutgers); Kerrville, Tex. (rectistrigella, 
in USNM). 

Foop Piant: Oak (larva a leaf feeder). 

Distrisution: Florida, Key West, Orlando (Mar.); 
Louisiana, Natchitoches Parish (Aug.); Texas, Blanco 
County (June), Kerrville (May, June), Sabinal (Sept.), 
Shovel Mountain (@June, July); Arizona, Williams 
(Sept.), Missourt (June, reared); District of Columbia 
(May, Aug., reared); New York, Bellport (June, Sept., 
reared). 

Superficially fructetella strikingly resembles Acrobasis 
amplexella Ragonot (especially the females). The reared 
examples before me had been identified to the latter 
name. The male antennal character and the genitalia 
of both sexes, however, easily separate the two species. 


64. Quasisalebria, new genus 


TYPE OF GENUS: Quasisalebria admixta, new species. 

Characters of Salebriaria except: Labial palpus erect, 
appressed to flattened face. Forewing with vein 8 and 
9 stalked for slightly more than two-thirds of their 
lengths. Hind wing with veins 4 and 5 stalked for over 
three-fourths of their lengths; 7 and 8 shortly anas- 
tomosed beyond cell. 

Male genitalia with costa of harpe sclerotized for its 
entire length, not appreciably produced at apex, but 
with a strongly sclerotized, free, articulating arm from 
base; shieldlike part of anellus with a pair of long, 
divergent, lateral horns; penis without cornutus. 

Female genitalis with ductus bursae thickened 
(cartilaginous in texture) except at junction with bursa 
copulatrix; bursa finely spined only in area adjacent to 
junction with ductus bursae. 

The type of this genus is, in many respects, close to 
Salebriaria fructetella and appears to be an aberrant 
offshoot of Salebriaria. I propose the new generic 
designation reluctantly; but no other procedure is pos- 
sible if we are to have any exact definition of genera for 
the species allied to Salebria. Nothing in our American 
fauna is properly referable to the latter genus, which is 
characterized by a forewing with partial scale ridge in 
the subbasal area; harpe (fig. 332) with erect clasper 
from near middle and costal margin weakly sclerotized ; 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 119 


bursa copulatrix of female without spining or granula- 
tions (membranous and smooth in the type, palumbella). 


242. Quasisalebria admixta, new species 
Figures 328, 806 


Forewing ashy white shaded with olivaceous brown 
or grayish fuscous in outer area and on lower half of 
basal area; the whitish ground color strongly contrasted 
on costal half of basal area and in a more or less tri- 
angulate area extending from median half of costa into 
cell and including the discal spots; antemedial line 
distinct, narrow, slightly curved, white, bordered out- 
wardly on costa by a strongly contrasted, black, tri- 
angulate patch and preceded on inner margin by a 
similar quadrate spot; subterminal line narrow, white, 
close to termen, and outwardly bulged at middle, 
bordered by fine blackish lines which begin as strong 
black smudges at costa; discal dots separated, the lower 
one always distinct, the upper sometimes absent. Hind 
wing semihyaline, white with a brownish shade at apex 
and a narrow brown line along upper half of termen; 
the veins not appreciably darkened. Alar expanse, 
19-21 mm. 

Male genitalia. Characters as given for the genus. 
The peculiar development of the anellus may be only 
of specific significance. 

TypE Locautity: Provo, Utah (type in USNM, 
61343). 

Foop prant: Unknown. 

Described from male type and two male and four 
female paratypes from the type locality (July, Aug.); 
one male paratype from Redington, Ariz.; two female 
paratypes from Bellevue, Washington County, Utah 
(May); and one female paratype from Glenwood 
Springs, Colo. (Aug.). 

The species is superficially similar to Salebriaria 
fructetella Hulst, but with the white areas and blackish 
markings of forewing more strongly contrasted. 


Genus 65: Ortholepis 


[Venational division B. Veins 4 and 5 of forewing connate 
(rarely in individual specimens, slightly separated at base). 
Hind wing with vein 2 from rather near lower outer angle of 
cell. Male genitalia with transtilla complete but its median 
area weakly sclerotized and granulate; costa of harpe strongly 
sclerotized throughout but not produced at apex; penis armed 
with a single, long, strong cornutus. Female genitalia without 
signum or scobinations in bursa.] 


65. Genus Ortholepis Ragonot 


Ortholepis Ragonot, N. Amer. Phycitidae, p. 6, 1887; Monograph, 
pt. 1, p. 214, 1893.—Hulst, Phycitidae of N. Amer., p. 140, 
1890. (Type of genus: Ortholepis jugosella Ragonot.) 

Tongue well developed. Antenna of male pubescent, 
shaft with sinus towards base containing a row of black 
toothlike spines more or less concealed under a weak 
seale tuft; antenna of female simple. Labial palpus 
obligue; second segment roughly and rather broadly 
scaled, on male slightly grooved on inner side; third 
segment very short, acuminate, reaching to height of 


vertex. Maxillary palpus of male squamous (jugosella) 
or in the form of an aigrette (pasadamia); of female 
minute and filiform. Forewing with ridge of raised 
scales on inner side of antemedial line, not reaching 
costa or inner margin; 11 veins; 2 from before lower 
outer angle of cell; 3 from the angle, separated at base 
from 4-5; 4 and 5 connate, rarely (in individual speci- 
mens) slightly separated at base; 6 from below upper 
angle of cell, straight, 8 and 9 stalked for slightly more 
than half their lengths; 10 from the cell, shortly sepa- 
rated from 8-9 at base and thence divergent; male 
without costal fold. Hind wing with vein 2 from before 
but rather near lower outer angle of cell; 3 from the 
angle, connate with the stalk of 4-5, short as compared 
with 2; 4 and 5 stalked for half or a trifle over half their 
lengths; 7 and 8 contiguous or anastomosed for a very 
short distance beyond cell; cell less than half the length 
of wing; discocellular vein curved, considerably extended 
at lower angle. Eighth abdominal segment of male 
simple. 

Male genitalia with uncus subtriangulate; slightly 
produced (lobed) near its lower, lateral angles. Apical 
process of gnathos a short, simple hook. Transtilla 
complete but its median area weakly sclerotized and 
granulate; its lateral elements broadly sclerotized. 
Harpe narrow; costal margin sclerotized throughout but 
not produced. Anellus U-shaped, narrowly sclerotized. 
Penis armed with a single stout cornutus nearly as long 
as aedeagus. Vinculum stout, triangulate; about as 
long as greatest width. 

Female genitalia without signum; bursa elongate, 
large, longitudinally wrinkled, partially sclerotized in 
the lobed area bearing the ductus seminalis; ductus 
bursae considerably shorter than bursa; flattened, 
strongly sclerotized, at least near and at its junction 
with bursa copulatrix; ductus seminalis from lobe of 
bursa near junction of bursa and ductus bursae. 
Eighth-segment collar with sclerotized part on dorsum 
reduced to a U-shaped band. 

An American genus showing affinities to Polopeustis 
and the various genera of the Salebria complex; but 
easily distinguished by its genitalia. Ragonot’s de- 
scription is somewhat misleading. Veins 4 and 5 of 
forewing are normally connate and not “nearly paral- 
lel”? except well beyond base; from base to near middle 
they are divergent. 


243. Ortholepis jugosella Ragonot 
Figures 23, 329, 808 


Ortholepis jugosella Ragonot, N. Amer. Phycitidae, p. 6, 1887; 
Monograph, pt. 1, p. 214, 1893.—Hulst, Phycitidae of N. 
Amer., p. 140, 1890.—McDunnough, Check list, No. 6149, 
1939. 

Maxillary palpus of male squamous. 

Forewing gray, rather shiny; palest (on some speci- 
mens ash gray) on costal half of median area and on the 
anterior costal half of basal area; remainder of basal 
and median areas and the area beyond the subterminal 
line brownish gray with a faint purplish suffusion; 


120 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


antemedial line oblique from costa to lower margin of 
cell, thence vertical to inner margin, its upper half 
obscure, indicated chiefly by a blackish outer border, 
lower half distinct, whitish; the antemedial line pre- 
ceded by a raised tuft of brown and blackish scales 
which are bordered inwardly by a more or less con- 
trasted white line; subterminal line faint, narrow, 
slightly bulged at middle; discal dots black, separated, 
distinct and rather large, especially the upper one; 
terminal dots reduced, obscure and more or less conflu- 
ent. Hind wing light brown; a thin blackish line along 
termen; the veins not appreciably darkened. Alar 
expanse, 19-20 mm. 

Male genitalia as given for the genus. Female geni- 
talia with posterior half of ductus bursa very weakly 
sclerotized. 

Type Locaity: “‘America septentrionalis” (type in 
Paris Mus.). 

Foop pLiant: Hickory (Carya alba) and wild azalea 
(Ragonot records). Hickory and walnut are the more 
probable hosts. 

Distripution: Unirep StratEs: Connecticut, Hast 
River (July). Canapa: Nova Scotia, White Point 
Beach (Queens County, July). 


244. Ortholepis pasadamia (Dyar), new combination 
Fiaure 807 
Immyrla pasadamia Dyar, Ins. Insc. Menstr., vol. 5, p. 45, 
1917.—McDunnough, Check list, No. 6189, 1939. 
Maxillary palpus of male in the form of a short aig- 
rette. 


Forewing uniformly dark glossy gray with a purplish 
tint; the transverse lines well marked, narrow, whitish; a 
contrasted white inner margin to the subbasal tuft; dis- 
cal dots usually confluent, distinguishable but not 
strongly contrasted against the dark ground color. 
Hind wing smoky fuscous; the veins faintly darkened. 
Alar expanse, 17-20 mm. 

Male genitalia as in jugosella. Female genitalia as in 
jugosella except ductus bursae sclerotized along ventral 
surface to genital opening. 

TypPE LOcALITy: St. Johns, Quebec (type in USNM). 

Foop Puant: Betula. 

Disrrizution: Unirep States: Pennsylvania, Ha- 
zleton (June); New Hampshire, Dublm, Hampton 
(July) ; Maine, Mount Desert Island (July), Sebec Lake 
(July); Washington, Meadow Creek (Grant County, 
Apr.), reared specimen, Walla Walla (June). Canapba: 
Ontario, Blacotasing (July), Ottawa (July), Waubamia 
(Perry Sound, July); Quebec, St. Johns (June). 

The foregoing description was drawn from typical 
examples represented in the National Collection by a 
series of 16 males and females. There are also before 
me four specimens from Maine, New Hampshire, and 
Quebec of what appears to be a color form in which the 
transverse lines (except for the white inner border of the 
raised-scale patch) are almost completely obliterated; 
the ground color of the forewing is darker (more pur- 
plish) and the hind wing is brown (as in jugosella) ; there 


is also a faint narrow dusting of whitish scales in the 
median area of forewing. The genitalia of these speci- 
mens are identical with those of typical pasadamia. 
Examples of both forms have been reared from Betula. 


Genus 66: Polopeustis 


[Venational division B. Vestiture of head, thorax, labial palpi, 
and femora a mixture of scales and hairs.] 


66. Genus Polopeustis Ragonot 


Polopeustis Ragonot, Monograph, pt. 1, p. 233, 1893.—Staudin- 
ger and Rebel, Catalog der Lepidopteren des palaearctichen 
Faunengebietes, vol. 2, p. 30, 1901.—Spuler, Die Schmetter- 
linge Europas, vol. 2, p. 212, 1910.—Forbes, Cornell Mem. 
68, p. 622, 1923.—Janse, Journ. Ent. Soc. South Africa, vol. 
5, p. 30, 1942. (Type of genus: Phycis annulatella Zetter- 
stedt; figs. 24, 330, 809.) 

Tongue well developed. Antenna of male ciliate- 
pubescent (the cilia a trifle shorter than width of seg- 
ments) the shaft curved towards base and containing 
two or three short teeth in the incurvation; of female 
simple. Labial palpus obliquely ascending, not reach- 
ing height of vertex; third segment short, less than one- 
third of second. Maxillary palpus minute, filiform. 
Forewing smooth; 11 veins; vein 2 from before but near 
lower outer angle of cell; 3 from the angle; 4 and 5 
separated at base; 6 from below upper angle of cell, 
straight; 8 and 9 stalked for over half their lengths; 10 
from the cell, shortly separated from the stalk of 8-9 at 
base; male without costal fold. Hind wing with vein 2 
from well before outer angle of cell; 3 from the angle; 4 
and 5 anastomosed for about half their lengths beyond 
cell; 7 and 8 closely approximate for a short distance 
beyond cell; cell less than half the length of wing; dis- 
cocellular vein curved, produced atlowerangle. Highth 
abdominal segment of male with pair of ventrolateral 
hair tufts. 

Vestiture of head, thorax, labial palpi, femora, and 
and foretibiae a mixture of scales and hairs. 

Male genitalia with uncus about as broad as long; at 
apical margin broadly rounded. Apical process of 
enathos (from ventral view) an inverted heart-shaped 
lobe terminating in a short, slender, hooked spine. 
Transtilla absent. Harpe short, stubby; its apex bluntly 
rounded; costa broadly and strongly sclerotized through- 
out, but not produced at apex; otherwise simple. Anel- 
lus a U-shaped plate terminating in short lateral lobes. 
Aedeagus slender; penis armed with moderately stout, 
elongate cornutus or two similar cornuti. Vinculum 
stout, slightly longer than greatest width; tapermg to 
blunt and rather broad terminal margin. 

Female genitalia without signum; bursa copulatrix 
small and with a rather broad, fused cartilagenouslike 
thickening around its lateral and anterior margins, 
otherwise minutely granulate and containing a weak 
elongate chitinized strip; ductus bursae twice as long as 
bursa, flattened, its ventral surface sclerotized and gran- 
ulate through its length, the sclerotization extending 
into bursa; genital opening simple. Highth-segment 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 121 


collar with sclerotized area reduced to a narrow, more 
or less U-shaped dorsolateral band (similar to that of 
Ortholepis). 

An Old World genus of Holarctic distribution with 
one European and one North American species; easily 
identified by its genitalia and hairy vestiture. 


245. Polopeustis arctiella (Gibson) 
Fiaures 331, 810 


Pyla arctiella Gibson, Report of the Canadian Arctic Expedition 
(1913-18), vol. 3, pt. 1, p. 46, 1920. 

Polopeustis annulatella arctiella (Gibson) McDunnough, Ca- 
nadian Ent., vol 67, p. 174, 1935; Check list, No. 6150, 1939. 

Forewing slate gray with a fine scattered sprinkling 
of white, the whitish dusting more concentrated in basa] 
area and the area beyond the subterminal line; trans- 
verse lines rather broad, well contrasted, white; ante- 
medial line oblique and slightly angulate; subterminal 
line sinuate; a narrow blackish shade bordering the 
antemedial outwardly and the subterminal inwardly; 
discal dots obscure, more or less confluent. Hind wings 
smoky white; the veins darkened; a narrow dark shade 
along termen. Alar expanse, 21-25 mm. 

Male genitalia with two cornuti on penis; otherwise as 
in annulatella. Female genitalia differ from those of 
annulatella chiefly in the shape of the sclerotized area of 
eighth-segment collar (compare figs. 809a and 810a). 
The differences in shape and extent of sclerotized area 
of bursa are probably individual in character. 

Type Locauity: Collinson Point, Alaska (type in 
Canadian Nat. Coll.). 

Foop piant: Unknown. 

Distrisution: AuasKa: Collinson Point (July). 
Canava: Labrador, Hopedale (July), Nain; Manitoba, 
Fort Churchill (June, July). 

Gibson’s name may represent no more than a New 
World race of annulatella; but arctiella is at least that. 
Good series of both are before me and their genitalic 
differences appear to be constant; the male of annulat- 
ella has but one cornutus, that of arctiella has consist- 
ently two cornuti. If and when intergrading examples 
of Polopeustis are recovered from northern Siberia the 
name arctiella may be reduced to subspecific status or 
referred as a synonym to annulatella. Meanwhile a 
specific separation seems the safer procedure. 


Genera 67-70: Meroptera to Tulsa 


[Venational division B. Veins 4 and 5 of forewing very shortly 
stalked, connate or closely approximate at base, in Tulsa approxi- 
mate at base and for a short distance beyond; § and 9 long stalked; 
10 frequently connate or shortly stalked with 8-9, if from the cell 
approximate to the stalk of 8-9 for some distance from its base. 
Antenna of male with sinus and scale tuft in base of shaft. Labial 
palpus erect or obliquely upturned. Maxillary palpus of male 
in the form of an aigrette or squamous. Male genitalia with 
transtilla frequently complete, but if so, weakly sclerotized; 
harpe with clasper always present and well sclerotized, digitate 
or enlarged and spined; harpe with long hair brush from inner 
surface along lower edge of basal half of sclerotized costa; penis 
armed with two stout, rather short cornuti. Female genitalia 


300329—56——_9 


with bursa finely and densely spined, usually with one or more 
sclerotized, granulate patches.] 


67. Genus Meroptera Grote 


Meroptera Grote, Canadian Ent., vol. 14, p. 29, 1882.—Hulst, 
Phycitidae of N. Amer., p. 148, 1890.—Ragonot, Mono- 
graph, pt. 1, p. 312, 1893.—Forbes, Cornell Mem. 68, 
p. 624, 1923. (Type of genus: Pempelia pravella Grote.) 

Emmerita Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 76, 
1930. (Type of genus: Meroptera mirandella Ragonot. New 
synonymy.) 


Tongue well developed. Antenna weakly pubescent; 
on male with sinus and scale tuft in base of shaft. La- 
bial palpus upturned, closely appressed to face; smooth 
scaled; reaching above vertex; second segment long, on 
male hollowed to receive maxillary palpus; third seg- 
ment short (about one-fourth of second), acuminate. 
Maxillary palpus of male in the form an of aigrette; 
of female squamous. Forewing smooth; 11 veins; vein 
2 from before but rather near lower outer angle of cell; 
3 from the angle, at base slightly nearer to 4-5 than to 
2; 4 and 5 very shortly stalked, connate or closely ap- 
proximate at base; 6 from below upper angle of cell, 
straight or (rarely) very slightly bent towards base; 8 
and 9 stalked for over two-thirds their lengths; 10 
shortly stalked or connate with the stalk of 8-9 (indi- 
vidually variable); male without costal fold. Hind 
wing with vein 2 from before but near lower outer angle 
of cell; 3 from the angle, connate with 4; 4 and 5 stalked 
for slightly more than half their lengths; 7 and 8 closely 
approximate for a short distance beyond cell; cell less 
than one-half the length of wing; discocellular vein 
curved, outwardly produced at lower angle of cell. 
Highth abdominal segment of male with compound 
scale tufts. 

Male genitalia with uncus triangulate or subtriangu- 
late, its apex bluntly rounded. Apical process of 
gnathos a short, stout hook. Transtilla complete but 
weakly sclerotized; a narrow, flatly arched band. 
Harpe with costa sclerotized throughout but not pro- 
duced at apex; a fine brush of long hairs arising from 
inner surface along lower edge of basal half of sclero- 
tized costa; cucullus simple, narrow, tapering slightly 
to rounded apex; from near base of harpe an appressed, 
stout, thorny or serrate clasper. Penis armed with 
two, stout, rather short cornuti less than one-half the 
length of aedeagus. Vinculum stout, as long as or 
somewhat longer than greatest width, its terminal 
margin bluntly rounded or narrowly truncate. 

Female genitalia without signum; bursa with a couple 
of conspicuous round or oval, strongly pigmented and 
sclerotized, densely granulate patches, otherwise bursa 
finely spinose over its membranous areas; ductus bursae 
shorter than bursa, not sclerotized adjacent to bursa, 
but with strong sclerotization at broadened genital 
opening; ductus seminalis from a lobe of bursa near 
junction of bursa and ductus bursae. 

This genus, while distinct from, is very close to 
Nephopteryz, from which it is distinguished only by the 


122 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


peculiar development of the clasper on harpe of the 
male genitalia and the strong sclerotization about the 
genital opening of the female. Hampson’s Hmmerita 
has not a single character to separate it from Meroptera. 
The stalking of vein 10 of forewing with 8-9, upon 
which Hampson evidently relied, is shared by the types 
of both Meroptera and Nephopteryr. This stalking is 
very short at most and is not even specifically constant 
in either genus. 

The genus as here defined contains but four North 
American species, and (to the best of my knowledge) 
no Old World representatives. 


246. Meroptera mirandella Ragonot 
Figures 21, 333, 816 


Meroptera mirandella Ragonot, Monograph, pt. 1, p. 313, 1893. 

Eimmerita mirandella (Ragonot) Hampson, Ann. Mag. Nat. Hist., 
ser. 10, vol. 5, p. 76, 1930.—McDunnough, Check list, 
No. 6181, 1939. 

Forewing ocherous white with a very faint and 
scattered dusting of blackish scales in median area; 
antemedial line narrow, oblique, indented at lower fold, 
bordered outwardly by a black line which begins as an 
enlarged, more or less triangular dash on costa, in- 
wardly by a straight black line from top of cell to inner 
margin; subterminal line obscure, dentate-sinuate, 
margined inwardly and outwardly for a short distance 
from costa by black lines, on well marked specimens the 
inner line continued as a fine dark border to tornus; 
also on well marked examples a faint blackish or 
fuscous shade extending obliquely across the wing from 
the inner costal edge of the antemedial line; discal dots 
faint, but usually distinguishable, blackish, separated, 
obliquely placed; terminal black dots more or less con- 
fluent. Hind wing white with a faint creamy or smoky 
tint; a very faint brownish line along termen; veins not 
or very slightly darkened. Alar expanse, 23-26 mm. 

Male genitalia with broad, irregularly fan-shaped 
clasper. Female genitalia with sclerotization of genital 
opening a narrow, corrugate, sclerotized band with 
short lateral, inward projections; bursa with two 
opposed, lateral, granulate patches. 

TypE Locatity: Colorado (type in Paris Mus.). 

Foop piant: Populus (this record from an Arizona 
specimen in the National Collections reared from a 
cocoon on a cottonwood leaf). 

Distrisution: Colorado, Denver; Arizona, Douglas 
(Aug.), Nogales (July), Phoenix (Apr.), Pinal Mts. 
(May), Readington, Tucson (Apr.), Yuma (June). 

One of the females before me from Phoenix is de- 
cidedly abnormal in venation, the forewing having 
vein 10 from the cell and closely approximate to the 
stalk of 8-9, 6 bent at base and from very near the upper 
angle of the cell and 4 and 5 closely approximate at base 
and for some little distance from the cell. Its genitalia 
are normal and it is clearly an individual aberration, 
but an example of what occurs all too often in the 
Phycitidae and which should caution us to use generic 
keys with discretion and to place unusual specimens 
only after examination of their genitalia. 


247. Meroptera cviatella Dyar 
Fieure 817 


Meroptera cviatella Dyar, Proc. Ent. Soc. Washington, vol. 7, p.34, 
1805.—MecDunnough, Check list, No. 6182, 1939. 
Salebria cviatella (Dyar) Forbes, Cornell Mem. 68, p. 627, 1923. 

Forewing bright reddish brown with a more or less 
distinct, transverse purplish gray shading in outer area 
and bordering outwardly the subterminal line; this 
latter area sometimes dusted with blackish scales; the 
central area near antemedial line more or less dusted 
with whitish scales; antemedial line oblique, narrow, 
strongly indented at lower fold, its upper half obscured; 
the white line bordered outwardly by a conspicuous 
black band, broad on costa and very gradually narrow- 
ing towards inner margin; inwardly bordered on lower 
half by a similar broad, vertical black band; subterminal 
line diffused, whitish; discal dots confluent, black, the 
lower one sometimes obscured; a row of more or less 
confluent black dots along termen. Hind wing pale 
smoky brown, darkened slightly towards termen. Alar 
expanse, 23-25 mm. 

Male genitalia similar to those of mirandella. Fe- 
male genitalia with sclerotization of genital opening a 
narrow, dark, granulate band, broken and with two 
short, divergent extensions at middle, these extensions 
fusing into a small, thin, triangulate, sclerotized patch 
on the lower median surface of the ductus bursae. 

Typx Locatity: Chicago, Ill. (type in USNM). 

Foop pianr: Populus (cottonwood). Larva a borer 
in buds and new shoots. 

Distrizution: Illinois, Chicago (June, July), Lacon 
(Aug.), Putnam County (July, Aug.); Mississippi, 
Starkville (July). 

The above food-plant record is from Putnam County 
specimens reared by Mr. Murry O. Glenn. One of 
his series is labeled ‘bred from larva on Amorpha 
canescens.”’ IJ doubt if Amorpha is a true food plant or 
the presence of the larva on this plant anything more 
than an accidental last-stage migration from cotton- 
wood. 


248. Meroptera pravella (Grote) 
Figures 22, 334, 812 


Pempelia pravella Grote, Bull. U. S. Geol. Geogr. Surv. Terr., 
vol. 4, No. 3, p. 694, 1878. 

Meroptera pravella (Grote), Canadian Ent., vol. 14, p. 30, 1882.— 
Packard, U. S. Dep. Agr. Div. Ent. Bull. 13, p. 23, 1887; 
U. S. Dep. Agr. Fifth Rep., Ent. Comm., p. 574, 1890.— 
Ragonot (in part), Monograph, pt. 1, p. 314, 1893.—Mc- 
Dunnough, Check list, No. 6184, 1939. 

Forewing dark gray; the basal area, except for a 
narrow dark gray shading at extreme base, pale ashy 
gray and contrastingly paler; antemedial line obscure 
and often partially obliterated, when distinguishable 
it is oblique, narrow, dull white, distinguished chiefly 
by its black borders, which tend to coalesce into a 
broad, diffused, blackish band; subterminal line faint 
but distinguishable, grayish white, sinuate, in fresh 
specimens bordered inwardly by an obscure, narrow, 
blackish line; discal dots separated, blackish, not con- 
spicuous; a faint blackish line along termen. Hind 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 123 


wing very pale smoky fuscous. 
mm. 

Male genitalia with an elongate, stout, tapering 
clasper extending about half the length of harpe; 
figured from male from Edmonton, Alberta, and com- 
pared by Clarke with the genitalia of the type of 
pravella in the British Museum. Female genitalia with 
sclerotization at genital opening in the form of a broad, 
stout, curved, granulate and wrinkled, lunate plate 
with strong lateral arms projecting into the ductus 
bursae. 

TypPE Locatiry: Oldtown, Maine (type in BM). 

Foop piant: Populus, Salix [?]. 

Disrrisution: Unitep States: Maine; New Hamp- 
shire, Hampton (July); New York, Ilion (June); Colo- 
rado, Chimney Gulch (Golden, June). Canapa: Que- 
bec, Knowlton (Feb., reared in laboratory from larva 
on aspen), Norway Bay (June); Ontario, Constance 
Bay (Feb., from poplar), Grand Bend (July), Hymers 
(June), Mer Bleue (June), Ottawa (Mar., from poplar), 
Smoky Falls (Mattagami River, June), Trenton (June); 
Manitoba, Aweme (May, June, July), Winnipeg; Sa- 
skatchewan, Indian Head (June, July); Alberta, Bilby 
(June, July), Edmonton (May, July), Nordegg (July); 
British Columbia, Canim Lake (June), ‘100 Mile 
House’”’ (June). 

These records (except for the type locality) are from 
specimens before me from the U. S. and Canadian 
National Collections. The Salix plant record is from 
Packard. I have not seen the reared Brunswick, 
Maine, specimens upon which it was based but have 
little doubt that the name pravella was correctly ap- 
plied. Most of the specimens in our collections that 
have been identified as pravella as well as many of the 
references in literature are referable to Nephopteryx 
subfuscella (Ragonot) (=semiobscurella (Hulst)). This 
confusion is discussed under subfuscella. The two 
species are easily confused, especially with worn exam- 
ples, if their genitalia are not examined; but in unrubbed 
specimens pravella is easily separated from subfuscella 
by the lack of any reddish or reddish ocherous scaling 
adjacent to inner margin at the base of forewing. Also 
included under pravella in the U. S. and Canadian 
National Collections were 21 Canadian examples of a 
new species (hereinafter described as abditiva) similar 
in all superficial characters to pravella but with dis- 
tinctly different male and female genitalia. 


Alar expanse, 20-22 


249. Meroptera abditiva, new species 


Ficures 335, 813, 814 


In color and maculation like pravella but with con- 
sistently different genitalia. Alar expanse, 19-22 mm. 

Male genitalia with harpe having a stout but short, 
stubby clasper, less than one-fourth the length of harpe. 
Female genitalia with the sclerotization at genital open- 
ing in the form of a rather narrow granulate, curved, 
transverse band without inwardly projecting arms. 
The pigmented, granulated area of the bursa is individ- 
ually variable in extent and sometimes divided into two 
distinct patches by a slight break at the anterior 


(closed) end of the bursa. Extremes of variation are 
shown in figures 813 and 814. However, there are no 
intergradations whatever between abditiva and pravella 
in the structure of the female genital plate nor in the 
male clasper. 

Typp Locatity: Knowlton, Quebec (type in Cana- 
dian Nat. Coll.; paratypes in Canadian Nat. Coll. and 
USNM, 61344). 


Foop piant: Populus tremuloides. 


Described from male type and 3 female paratypes 
from the type locality reared (in laboratory) Feb. 12, 
1 and 2, 1930, from larvae feeding on leaves of P. tre- 
muloides (“aspen’’) by J. McDunnough; and 17 para- 
types from the following Canadian localities: Quebec, 
Mount St. Hilaire, June 30, 1908, G. Chagnon (9); 
Norway Bay, June 4, 1938, E. G. Lester (@). New 
Brunswick, Chamcook, June 23, 1938, T. N. Freeman 
(9); Eel River, June 21, 1941, T. N. Freeman (<*). 
Nova Scotia, Beddeck, June 23, 1936 and June 27, 1938, 
T. N. Freeman (29); White Point Beach, Queens, Feb. 
12 and 20, 1936, J. McDunnough (2 9, reared). 
Ontario, Trenton, May 29 and June 25, 1908, Evans 
(2 9); Vineland Station, June 15, 1936, W. L. Putnam 
(2, reared). Saskatchewan, Christopher Lake, June 
19, 1939, A. R. Brooks (). British Columbia, 
Canim Lake, June 25, 1938, J. K. Jacob (9); Jesmond, 
July 13, 1937, J. K. Jacob (2 9); Kaniloops, June 14, 
1937, J. K. Jacob (@); Shingle Creek, Penticton, 
June 25, 1935, A. K. Cartrell (9). 

In as much as pravella and abditiva have the same 
hosts and larval habits, an overlapping distribution, 
and similar habitus, it is necessary to examine their 
genitalia to distinguish them apart. 


68. Genus Nephopteryx Hiibner 


Nephopieryx Hiibner, Verzeichniss bekannter Schmett[er]linge, 
p. 370, 1825.—Zeller, Isis von Oken, 1846, p. 731.—Grote, 
Bull. U. 8. Geol. Geogr. Surv. Terr., vol. 4, p. 695, 1878; 
North Amer. Ent., vol. 1, p. 11, 1879.—Ragonot, Ent. 
Monthly Mag., vol. 22, p. 19, 1885 (citation of type); Mono- 
graph, pt. 1, p. 254, 1893.—Hulst, Phycitidae of N. Amer., 
p. 142, 1890.—Forbes, Cornell Mem. 68, p. 622, 1923.— 
Hemming, Hubner, vol. 2, p. 229, 1937.—Bisset, in Pierce 
and Metcalfe, Genitalia of the British Pyrales, p. 61, 1938.— 
Janse, Journ. Ent. Soc. South Africa, vol. 5, p. 34, 1942. 
(Type of genus: Phycita rhenella Zincken; Europe; figs. 25, 
336, 815.) 

Sciota Hulst, Ent. Amer., vol. 4, p. 115, 1888. 
Sciota croceella Hulst.) 


Characters of Meroptera except: Labial palpus erect 
or obliquely upturned. Maxillary palpus of male in 
the form of an aigrette or squamous. Forewing with 
10 usually connate with the stalk of 8-9 or closely 
approximate to it, rarely stalked. Transtilla frequently 
incomplete or absent. Clasper of harpe digitate, slen- 
der, simple (without spining). One cornutus on penis in 
uvinella, other species have two cornuti as in Meroptera. 
Female genitalia with ductus bursae sclerotized along 
ventral surface for most of its length from junction with 
bursa, the sclerotization terminating before genital 
opening, the latter simple (unsclerotized). 


(Type of genus: 


124 


As here defined the genus includes a number of 
species with two types of maxillary palpi, several of 
which have been hitherto referred to either Salebria or 
Myrlaea. The reference of some of our North American 
species to the latter genus on the strength of a slight 
obliqueness of their labial palpi is not warranted as 
none of them agrees with the Old World type of Myrlaea 
(albistrigella Staudinger) on either male or female 
genitalic characters. The ductus bursae and bursa of 
albistrigella are perfectly smooth, without granulations, 
scobinations, or sclerotizations of any kind; and its 
harpe lacks the hair brush characteristic of Meroptera 
and Nephopterysx. 

The two types of male maxillary palpi (aigrettelike 
and squamous) do not justify generic separation of the 
species here any more than they do in Dioryctria or 
Ortholepis although such a difference should be, and in 
the Phycitidae usually is, of generic significance. On 
basilaris, the palpus is midway between the two types, 
though somewhat more aigrettelike than squamous. 
The uniform type of their male and female genitalia and 
the similar habitus of the included species indicate a 
distinct and natural group. Nephopteryz is very close 
to Meroptera but is distinguished by its simple, slender 
clasper, simple female genital opening, and differently 
sclerotized ductus bursae. 


Genus Nephopteryxz, Species 250-267: N. sub- 
fuscella to N. celtidella 


[Males with aigrettelike palpi.] 


250. Nephopteryx subfuscella (Ragonot), new combination 
Fieures 337, 821 


Salebria subfuscella Ragonot, N. Amer. Phycitidae, p. 8, 1887; 
Monograph, pt. 1, pp. 329, 350, 1893.—Hulst, Phycitidae 
of N. Amer., p. 151, 1890.—McDunnough, Check list, No. 
6123, 19389. 

Salebria semiobscurella Hulst, Phycitidae of N. Amer., p. 151, 
1890.—Ragonot, Monograph, pt. 1, p. 352, 1893.—Barnes 
and McDunnough, Contributions, vol. 3, p. 197, 1916.— 
Forbes, Cornell Mem. 68, p. 626, 1923.—McDunnough, 
Check list, No. 6212, 1939.—Craighead, U. S. Dep. Agr. 
Misc. Publ. 657, p. 454, 1950. (New synonymy.) 

Meroptera pravella (authors not Grote) Hulst (in part), Phyci- 
tidae of N. Amer., p. 148, 1890.—Ragonot (in part), Mono- 
graph, pt. 1, p. 314, 1893.—Grossbeck, Bull. Amer. Mus. 
Nat. Hist., vol. 37, p. 180, 1917.—Forbes, Cornell Mem. 68, 
p. 624, 1923. 


Color and maculation resembling those of Meroptera 
pravella. Forewing gray; the basal area contrastingly 
paler with some dark shading at extreme base and more 
or less reddish or orange scaling on base of inner margin 
(at least a trace on all except badly rubbed specimens); 
antemedial line obscure, indicated chiefly by its fused 
dark inner and outer borders which form a rather broad, 
oblique, blackish band, the antemedial line itself distin- 
guishable on most specimens only as an incomplete, me- 
dian, pale (whitish) streak in the dark band ;subterminal 
line obscure but complete, narrow, sinuate, pale gray with 
fine dark bordering lines; some whitish dusting over 
central area of wing, especially on pale examples; discal 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


dots blackish, occasionally separated, more often fused 
into a curved line; dots along terminal line fine, weak, 
blackish, usually separated, on some specimens con- 
fluent. Hind wing pale brown to smoky fuscous; veins 
darkened, especially on the darker females; a narrow 
dark shade along termen. Alar expanse, 18-22 mm. 

Male genitalia with apical process of gnathos triangu- 
late. Clasper bent downward, parallel to surface of 
harpe. Female genitalia with granulate sclerotized 
patch on bursa a continuous band across posterior 
ventral surface and forward on right side of bursa. 

Type tocauity: Not given (subfuscella, in Paris 
Mus.); Blanco County, Tex. (semiobscurella, in AMNH, 
ex Rutgers). 

Foop piants: Rhus, locust [7]. Larva a leaf-folder. 

Distripution: Unitrp statrs: Maine, Augusta 
(June); Massachusetts, Martha’s Vineyard (May, July); 
Connecticut, East River (June); Rhode Island, Weeka- 
paugh (Aug.); New York, Ilion (June), Rossville (Long 
Island, Feb., Mar.), Shore of Lake Ontario (near 
Rochester, May, June); Pennsylvania, Oak Station 
(Aug.); District of Columbia, Washington; North Caro- 
lina, Tryon (May); Florida, Fort Myers (May), Lake 
Alfred (July); Texas, Burnet County (Apr.), Kerrville 
(May, Aug.); Missouri, Kirkwood (Mar., Apr., May); 
Iitinois, Decatur (July, Aug.); Washington, Almota 
(July), Bellingham (Nov.), Pullman (Feb., May, June, 
July, Aug., Nov.), Snake River (Jan., Feb., July), 
Walla Walla (uly), Wawawai (May). Canapa: Que- 
bec, Levis, Meach Lake (July), Mount St. Hilaire (July); 
Ontario, Merivale (Mar.), Ottawa (Mar.), Trenton 
(May, June, July). 

Many of the above records are from reared examples 
from sumac, and all such had been identified by Riley 
and Dyar as Meroptera pravella. Barnes and McDun- 
nough (1916) were the first to question and correct this 
identification, assigning the name Salebria semiobscurella 
Hulst to the sumac feeder. At that time subfuscella 
had not been recognized by American lepidopterists. 
It resembles pravella but has the reddish scaling on base 
of inner margin of forewing. Unquestionably it is the 
same as what Hulst later described as semiobscurella. 
No other American species that could have been referred 
to Meroptera or Salebria has this red-scale character 
with the habitus, otherwise, of pravella. The food 
plant record of dried peaches (‘‘peches desséchées’’) given 
for subfuscella by Ragonot (Monograph, pt. 1, p. 352) 
on the basis of two imperfect specimens sent him so 
labeled by Riley is obviously incorrect and can be 
ignored. I question also “locust” as a probable or even 
occasional food plant. That record, given above, is 
based on four males from the Fernald collection that 
had been identified by Dyar as pravella. ‘They had been 
reared (Apr. 1889, May 1893) from larvae collected by 
Miss Murtfeldt at Kirkwood, Mo., onlocust. Her note 
(“‘324M’’) states that most of the larvae in the lot were 
“‘Salebria contatella Grote” but that a few seemed to be 
a different species. I suspect that the latter (the sub- 
fuscella larvae) had merely migrated to the locust after 
feeding on nearby sumac. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 125 


251. Nephopteryx delassalis Hulst 
Fieure 818 


Nephopteryx delassalis Hulst, Trans. Amer. Ent. Soc., vol. 13, 
p. 161, 1886. 


Salebria purpurella Hulst, Canadian Ent., vol. 24, p. 61, 1892.— 
Forbes, Cornell Mem. 68, p. 627, 1923. 


Salebria pudibundella Ragonot, Monograph, pt. 1, p. 350, 1893.— 
McDunnough, Check list, No. 6208, 1939. (New synonymy.) 

Myelaea delassalis (Hulst) Barnes and McDunnough, Contribu- 
tions, vol. 3, p. 198, 1916.—McDunnough, Check list, No. 
6225, 1939. 


Thorax vinous red with a scattered dusting of white 
and black scales. Forewing vinous red with a more or 
less diffused bluish white shading in basal area immedi- 
ately preceding inner border of the antemedial line; 
this border a broad black band (the most conspicuous 
marking on the wing), vertical, and fusing at costa with 
the outer black border; the antemedial line itself faint, 
but on most specimens its lower half distinct, narrow, 
white, slightly oblique and inwardly angulate at lower 
fold, bordered outwardly by a rather narrow, inter- 
rupted black band; some blackish dusting on the whitish 
subbasal area, black scaling along lower margin of cell, 
on some of the lower veins from cell and on costa, espe- 
cially towards apex; discal dots (when distinguishable) 
separated, red, more or less shaded with black; dots 
along termen very faint, blackish, more or less confluent. 
Hind wing pale ocherous fuscous. Alar expanse, 22- 
26 mm. 


Male genitalia similar to those of fernaldi; transtilla 
absent; apical process of gnathos narrow (not triangu- 
late); clasper short, slightly curved. Female genitalia 
with two granulate patches on bursa copulatrix, a small 
one on posterior dorsal surface, near the left side of 
bursa and a larger on ventral surface at the antero- 
lateral margin (closed end) of bursa. 


Typr Locatities: Nevada (delassalis, in AMNH, ex 
Rutgers); New Mexico (purpurella, in AMNH, ex Rut- 
gers); Colorado (pudibundelia, in Paris Mus.). 

Foop piant: Unknown. 


Distrisution: Unirep Sratrs: Arizona, White 
Mts; Colorado, Beulah (June), Fort Collins; Utah, 
Vineyard (June, July) ; Nevada; California, Inyo County 
(May, June, July). 

The types of delassalis and purpurella are both fe- 
males. Their genitalia are alike. Barnes and McDun- 
nough (1916) first noted this synonymy and the mis- 
identification of his species by Hulst (1900) and the 
consequent misapplication of the name delassalis to 
specimens of fernaldi Ragonot. The descriptions under 
delassalis in Hulst’s 1900 revision and in Ragonot’s 
monograph apply to fernaldi and not delassalis. On 
the other hand, Ragonot’s description of pudibundella 
applies in detail to the true delassalis. 

The venation of forewing is individually variable, as 
with many species of Nephopteryz, vein 10 being either 
connate or closely approximate at base to the stalk 8-9 
(rarely short stalked with it) and veins 4 and 5 either 
connate or closely approximate at base. 


252. Nephopteryx delassalis fraudifera, new race 


Superficially appears quite distinct from delassalis; 
the entire median area and much of the basal area of 
forewing being heavily dusted with whitish, giving the 
general ground color a whitish blue-gray shade similar 
to that of inconditella rather than the vinous red of 
typical delassalis; the vinous red limited in fraudifera 
to the thorax, extreme base of forewing, a broad band 
outwardly bordering the subterminal line, and a faint 
diffused shading just preceding it. The red shade 
somewhat darker than in typical delassalis; the black 
borders of antemedial line also somewhat broader, es- 
pecially at costa. Alar expanse, 24-26 mm. 

Male and female genitalia agreeing in all details with 
those of delassalis. 

Typr Locauiry: Oliver, British Columbia, (type in 
Canadian Nat. Coll.; paratypes in USNM, 61345, and 
Canadian Nat. Coll.). 

Foop piant: Unknown. 

Described from male type and one female paratype 
from the type locality (June 8 and 9, 1923, G. B. Gar- 
rett, collector) ; and paratypes from the following locali- 
ties: “Shingle Cr. Road,” Keremeos, British Columbia 
June 22, 1935, A. N. Cartrell (9); Salmon Arm, British 
Columbia, June 20, 1916, ‘“W. R. B.”’ (9); Kaslo, Brit- 
ish Columbia, June 13, 1903, H. G. Dyar, “19366” (9); 
Alberni, British Columbia, June 20, 1922, ““‘W. R. B.” 
(@), and Bellingham, Wash., May 30, 1922, J. F. G. 
Clarke (). 

Presumably a distinct food plant as well as local race. 


253. Nephopteryx rubescentella (Hulst) 

Mineola rubescentella Hulst, Canadian Ent., vol. 32, p. 169, 1900. 
Nephopteryx rubescentella (Hulst), U. S. Nat. Mus. Bull. 52, p. 
419, 1903.—MecDunnough, Check list, No. 6173, 1939. 

Ground color of forewing slightly paler than that of 
typical delassalis. Thorax also paler, pale purplish 
gray or grayish ocherous. The dark bands bordering 
antemedial line on forewing dull red or reddish orange, 
containing no black except for occasional scattered 
scales. Maculation otherwise as in typical delassalis. 
Alar expanse, 26 mm. 

Male genitalia similar to those of fernaldi. 

Typm LOCALITY: Tennessee (type in USNM). 

Foop prant: Unknown. 

In addition to the male type, the National Collection 
contains a male from Denver, Colo. (Aug.). I have 
seen no other specimens. A female from the type 
locality will be needed before the exact status of rubes- 
centella can be determined. It may be no more than 
a race or variety of fernaldi. 


254. Nephopteryx fernaldi (Ragonot), new combination 
Friaures 340, 819 


Salebria fernaldi Ragonot, N. Amer. Phycitidae, p. 9, 1887. 

Salebria delassalis Hulst (not Hulst), Phycitidae of N. Amer., 
p. 154, 1890. 

Myrlaea delassalis (Hulst not Hulst) Ragonot, Monograph, pt. 1, 
p. 402, 1893. 

Myrlaea fernaldi (Ragonot) McDunnough, Check list, No. 6226, 
1939. 


126 


Forewing whitish ocherous with basal half of costa 
faintly washed with reddish ocherous, also some traces 
of this shading on the pale ocherous thorax and along 
inner margin of forewing from base to antemedial line, 
strongest on costa just above the dark borders of the 
antemedial line; some blackish dusting along costa near 
apex and occasionally on a few of the veins; antemedial 
line obscure, broken, frequently obscured entirely by 
its black borders; the latter forming a broad, strongly 
contrasted black blotch which extends from inner 
margin to top of cell (not reaching costa); discal dots 
much reduced or absent, if present more or less con- 
fluent. Hind wing pale whitish ocherous; the veins 
very slightly darkened. Alar expanse, 22-25 mm. 


Male genitalia figured from Colorado specimen identi- 
fied by Hulst as delassalis (one of his spurious “‘types’’). 
There are no structural differences to distinguish the 
male genitalia of the two species. 

Female genitalia with a single, rather large, granulate 
patch in bursa, on left side and extending onto ventral 
surface near posterior end of bursa; the bursa copula- 
trix itself as broad as long, approximately round, the 
membrane at anterior end thickened (cartilaginous). 

Type Locauity: Arizona (type in Paris Mus.). 

Foop prant: Unknown (probably Amorpha sp.). 

Distripution: Unirep States: Arizona; Colorado, 
Denver, Glenwood Springs (July), also two males and 
a female with only the state locality, identified by Hulst 
as ‘delassalis’’; Kansas, Thomas County; Jowa, Ames. 
Canava: Manitoba, Cartwright (June, July), Winnipeg. 

The female genitalia and ground color of thorax and 
forewing easily separate this species from delassalis, 
with which it has been confused due to Hulst’s later 
(1890) misidentification of his own species. 


255. Nephopteryx dammersi, new species 
Figures 338, 822 


Ground color pale ashy gray strongly shaded with 
pale rust red on thorax, at extreme base, on basal half 
of costal edge and on basal third of inner margin of 
forewing; more or less of this red shading on the black 
inner border of the antemedial line and forming its 
outer border near costa; an obscure, ill-defined, rusty 
blotch on middle of lower fold; antemedial line distin- 
guishable on most specimens, narrow, whitish, slightly 
oblique and indented between cell and inner margin 
(the antemedial line, where it can be distinguished for 
any appreciable distance, has a similar configuration 
and slant on most of our American Nephopteryz), 
bordered outwardly on lower half by a narrow, inter- 
rupted black line and inwardly by a broad black band, 
the latter extending only from inner margin to middle 
of cell and (as noted above) more or less shaded with 
rust red; subterminal line obsolete or nearly so; discal 
dots usually distinct, especially the lower one, sepa- 
rated, blackish; terminal dots minute, very faint, not 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


confluent. Hind wing dull white with a faint ocherous 
tint on male; pale smoky fuscous on female; the veins 
faintly darkened and a faint, narrow, dark shade along 
termen. Alar expanse, 25-26 mm. 

Male genitalia similar to those of fernaldi. There is 
some difference in the armature of the penis between the 
two species; the cornuti are somewhat shorter and 
stouter and there is a darker pigmentation of some of 
the scobinations on penis in dammersi (fig. 338); but 
there is so much individual variation of these structures 
within any given species of Nephopteryx that they can 
not be safely used to separate species. Female genitalia 
with two small and one large granulate patch in bursa. 

Type tocauity: Cajon Valley, San Bernardino 
Valley, Calif. (type in USNM, 61346). 

Foop pLant: Amorpha californica. 

Described from male type and three male and six 
female paratypes from the type locality (reared by 
Commander J. Dammers Apr. 15, 1933, and July 20, 
30, 1932), and one female from the Huachuca Mts., 
Ariz. (“July 8-15’). In addition I have before me a 
male from Douglas County, Ariz. (July 20, 1940, 
collected by Fritz Forbes), which appears to be con- 
specific. It is a trifle smaller (24 mm.) than the speci- 
mens of the type series. Without a matching female 
from the same locality it appears unwise to include it 
among the paratypes. 

The species is named in honor of Commander 
Dammers, who has given many fine reared and collected 
specimens to our National Collection. It is close to 
fernaldi, but on female genitalic characters seems to be 
a distinct species and not a color form or local race. 


256. Nephopteryx dammersi floridensis, new race 


Smaller and considerably darker than typical dam- 
mersi; the ground color dark gray finely peppered with 
white especially in median and subbasal areas making 
these areas a trifle paler than remainder of wing; the 
rust red markings of typical dammersi replaced by 
lavender-red in floridensis and this color more extended, 
forming a faint suffusion over much of the median and 
outer areas in addition to the stronger markings on 
costa, inner margin and extreme base; subterminal line 
distinct and with narrow dark borders. Hind wing 
smoky fuscous; the veins darkened; a narrow blackish 
line along termen. Alar expanse, 21-22 mm. 

Genrrauia: Male and female as in typical dammersi. 

TypE Locauity: Williamsburg, Fla. (type in USNM, 
61347). 

Foop PpLant: Amorpha herbacea. 

Described from male type from the type locality, 
reared under S. S. No. 16970A, June 25, 1944; and one 
female paratype from Tampa, Fla., reared under S. S. 
No. 16859, June 29, 1944. Larva collected and both 
specimens reared by members of the Special Survey of 
the Division of Foreign Plant Quarantine of the U. S. 
Bureau of Entomology and Plant Quarantine. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 127 


257. Nephopteryx vetustella (Dyar), new combination 
FicurE 820 


Salebria vetustella Dyar, Journ. New York Ent. Soc., vol. 12, 
p. 106, 1904.—Forbes, Cornell Mem. 68, p. 626, 1923. 
Myrlaea vetustella (Dyar) Barnes and McDunnough, Check list 
of the Lepidoptera of Boreal America, No. 5650, 1916.— 
Forbes, Cornell Mem. 68, p. 627, 1923.—McDunnough, 

Check list, No. 6224, 1939. 

Forewing pale ash gray with a slight purplish shading 
over outer half; basal area almost to inner dark border 
of antemedial line, Indian red or reddish orange, this 
reddish shade also on collar and top of head; antemedial 
line distinct from upper vein of cell to inner margin, 
narrow, oblique, inwardly notched between cell and 
inner margin, bordered inwardly by a broad black band 
and outwardly by a narrow black line, the black borders 
continuous to and fused at costa; subterminal line 
usually distinct but faint and with a weak dark inner 
bordering line, sinuate; discal dots separated, small, 
blackish; a weak row of blackish dots along termen. 
Hind wing pale smoky brown; the veins and terminal 
margin slightly darkened. Alar expanse, 22-25 mm. 

Male genitalia similar to those of fernaldi. Female 
genitalia with a single long granulate patch in bursa 
extending nearly the length of the bursa on its left side 
(seen from below) and curving onto ventral surface near 
junction of bursa and ductus bursae. 


Typr Locauity: Plummers Isl., Md. (typein USNM). 


Foop puant: Unknown (Amorpha sp. ?). 

Disrrisution: Unirep Sratses: Maryland, Plum- 
mers Isl. (Apr., May); Pennsylvania, Oak Station 
(June), Pittsburgh (May, June) ; New York, Ilion (May) ; 
Illinois, Edgebrook (June), Elkhart (Aug.), Palos Park 
(June); Iowa, Iowa City (June), Sioux City (July); 
Missouri, St. Louis; Florida, St. Petersburg (June). 
Canapa: Ontario, Ottawa (June, July); Quebec, Meach 
Lake (June). 

This species forms one of a group of closely related 
species (delassalis to vetustella) that feed upon Amorpha 
and have similar male genitalia but differ markedly in 
the number, arrangement, and relative sizes of the gran- 
ulate patches of the female bursa. These differences 
appear to be constant and the specific character holds 
even in series of individuals from widely different 
localities. The male genitalia exhibit some minor dif- 
ferences, especially in the relative size and position of the 
two cornuti; but here individual variation makes the 
apparent differences untrustworthy for specific differ- 
entiation. During dissection of the male organs a 
simple transtilla can be faintly seen in all the species of 
the group, but it is so weakly sclerotized that it is not 
visible or but partially distinguishable in balsam mounts. 
The structure is obsolescent and cannot be considered 
“present”? in the sense in which it is in Meroptera for 
example or even in a few other species of Nephopteryz. 


258. Nephopteryx inconditella (Ragonot), new combination 
Figure 825 


Salebria contatella inconditella Ragonot, Monograph, pt. 1, 
348, 350, 1893.—McDunnough, Check list, No. 6205b, 1939, 


Thorax and forewing pale ash gray with a faint bluish 
tint; some reddish scaling at extreme base of wing and a 
small spot of the same shade at middle of lower fold; 
pale antemedial line distinguishable from inner margin 
to cell, deeply notched at vein 1b, its black borders 
complete from inner margin to costa, the inner border 
broad and fusing with the narrow outer one near costa; 
subterminal line distinct, sinuate-serrate, narrowly bor- 
dered by obscure blackish lines; discal spots obscure but 
usually distinguishable, separated; terminal dots weak, 
confluent; on most specimens a narrow, dark transverse 
shade extending from costa at inner edge of subterminal 
line to middle of inner margin. Hind wings translucent, 
whitish with a faint ocherous tint, the veins slightly 
darkened; a narrow brownish shade along terminal 
margin. Alar expanse, 25-30 mm. 

Male genitalia of the fernaldi type but clasper some- 
what more strongly sclerotized than in the seven preced- 
ing species. Female genitalia with a single, small, 
round, granulate patch on ventral surface of bursa very 
near junction of bursa and ductus bursae. 

Type Locauity: Colorado (type in Paris Mus.). 

Foop prant: Unknown. 

Distrisution: Colorado; Arizona, Huachuca Mts., 
Palmerlee, and four Arizona specimens with only the 
state locality and without dates, one of these a pseudo- 
type of ‘Pinipestis albovittella Hulst.”’ 

The species was described as a pale western variety of 
contatella (Grote), to which it is apparently most nearly 
related and which, except for its paler ground color, it 
resembles. Its female genitalia however clearly indi- 
cate a distinct species. The ground color of forewing 
is intermediate between that of subcaesiella (=conta- 
tella) and that of dammersi. 


259. Nephopteryx subcaesiella (Clemens), new combination 
Fiaurus 339, 826 


Pempelia subcaesiella Clemens, Proc. Acad. Nat. Sci. Philadel- 
phia, p. 206, 1860. 

Pempelia contatella Grote, North Amer. Ent., vol. 1, p. 49, 1880.— 
Comstock, in Rep. [U. S.] Comm. Agr. for 1880, p. 261, 1881 
(in part). 

Salebria contatella (Grote), Bull. U. S. Geol. Geogr. Surv. Terr., 
vol. 6, no. 3, p. 590, 1882.—Beutenmiiller, Canadian Ent. 
vol 22, p. 16, 1890 (larva).—Hulst, Phycitidae of N. Amer., 
p. 152, 1890; U. S. Nat. Mus. Bull. 52, p. 424, 1903.— 
Packard, U. S. Dep. Agr. Fifth Rep. Ent. Comm., p. 361, 
1890.—Ragonot, Monograph, pt. 1, p. 348, 1893.—Forbes, 
Cornell Mem. 68, p. 626, 1923.—McDunnough, Canadian 
Ent., vol. 78, p. 109, 1946. 

Salebria virgatella subcaesiella (Clemens) Barnes and McDun- 
nough, Check list of the Lepidoptera of Boreal America, No. 
5631a, 1916—McDunnough, Check list, No. 6205a, 1939.— 
Craighead, U. 8. Dep. Agr. Mise. Publ. 657, p. 453, 1950. 


Color and maculation similar to those of inconditella 
but the ground color of forewing a much darker gray, 
basal area contrastingly paler gray; the reddish scaling 
on base of wing ranging from wine to rusty ocherous, 
always present but sometimes reduced to a few scales, 
rarely extended for a short distance onto the thorax; 
inner black border of antemedial line somewhat broader 
and more diffused than on inconditella, complete to costa 


128 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


(unbroken at any part of its length) ; following the ante- 
medial line 2 pale grayish spot (sometimes very faint but 
usually more contrasted than in inconditella); discal 
spots distinct, black, separated. Hind wing smoky 
white to smoky fuscous or brown, darker on females 
than on males, a distinct dark shade along termen. 
Alar expanse, 21-28 mmi 

Male genitalia as in inconditella and virgatella except 
for the armature of the penis. In both inconditella and 
virgatella there are two moderately stout cornuti situ- 
ated on penis, one to the side and slightly behind the 
other. In subcaesiella the second cornutus is greatly 
reduced. McDunnough (1946) states that there is only 
one cornutus in subcaesiella (contatella). That could 
easily be the case on individual specimens, but the 
normal condition is two cornuti on penis. Every prep- 
aration J have seen shows at least a vestige of the 
second cornutus. In all three species the lateral ele- 
ments of transtilla are indicated, but very weakly 
sclerotized. Female genitalia without granulate patch 
or patches on bursa. 

Typr Locanitius: Not given, presumably Pennsyl- 
vania (subcaesiella, in Acad. Nat. Sci. Philadelphia); 
“New England” (contatella, i BM). 

Foop piant: Robinia pseudoacacia. Wisteria also 
recorded as food plant. 

DistrisutTion: Unirep States: Maine, Augusta 
(May, June), Orono; New Hampshire, Hampton (July); 
Massachusetts, Amherst (June), Framingham (May), 
Martha’s Vineyard (July, Aug.), Newton Highlands; 
New Jersey, Essex County Park (June), New Lisbon 
(Aug.) ; Pennsylvania, New Brighton (May, July, Aug.), 
Oak Station (June), Pittsburgh (May, June, July); 
Maryland, Plummers Isl. (June, Aug.), Somerset Heights 
(Aug.); District of Columbia, Washington (Apr., May, 
July); Virginia, Falls Church (Aug.), Snickers Gap 
(July); North Carolina, Black Mountain (July), Tryon 
(May); Tennessee, no specific locality (May); Iihnois, 
Decatur (July), Elkhart, Oconee (July); Jowa, Iowa 
City (Aug.), Sioux City (June) ; Missouri, “Cent. Mo.”’ 
(Aug.), Kirkwood (Apr., May), St. Louis (June); Ar- 
kansas, Washington County (‘“‘July—Aug.”). CanNapa: 
Nova Scotia, Smith’s Cove (recorded by McDunnough, 
1946). 

I have not seen any Canadian examples but Mc- 
Dunnough’s description leaves no doubt of what he had. 
He notes the differences in the cornuti and larval 
characters between subcaestella (contatella) and virgatella 
quinquepunctella) and treats them as a distinct species. 
The difference in their female genitalia and the con- 
sistent difference in maculation of forewing are added 
evidence that they are not merely races of one variable 
species. 


260. Nephopteryx virgatella (Clemens), new combination 
Figure 827 


Pempelia virgatella Clemens, Proc. Acad. Nat. Sci. 
p. 205, 1860. 

Pempelia contatella quinquepunctella Grote, North Amer, Ent., 
vol. 1, p. 50, 1880.—Comstock, in Rep. [U. 8.] Comm. Agr. 
for 1880, pp. 261-262, 1881 (part; larva). 


Philadelphia, 


Salebria contatella quinquepunctella (Grote) Hulst, Phycitidae of 
North Amer., p. 152, 1890; U.S. Nat. Mus. Bull. 52, p. 424, 
1903.—Ragonot, Monograph, pt. 1, p. 348, 1893.—Forbes, 
Cornell Mem. 68, p. 626, 1923. 

Salebria virgatella (Clemens) Barnes and MceDunnough, Check 
list of the Lepidoptera of Boreal America, No. 5631, 1916.— 
McDunnough, Check list, No. 6205, 1939. 

Salebria quinquepunctella (Grote) McDunnough, Canadian Ent., 
vol. 78, p. 109, 1946 (larva). 

Ground color of forewing (on most specimens) 2 
trifle paler than that of subcaesiella, gray with a faint 
brownish or purplish tint; a reddish (or pale purplish) 
brown shade along lower fold, cutting the antemedial 
line and its black borders; a similar, narrower, shorter 
streak on median fold; outer black border of ante- 
medial line between top of cell and inner margin reduced 
to two black dots, one on the lower margin of cell, the 
other on vein 1b and enclosed within the pale patch 
following the antemedial line; both these dots and the 
black discal dots at end of cell well contrasted; sub- 
terminal line faint, not appreciably darkly bordered, 
Interrupted at the folds. Hind wing smoky white to 
brown; the veins more or less darkened and a smoky 
brown shade along termen. Alar expanse, 22-26 mm. 

Male genitalia similar to those of inconditella; both 
cornuti moderately stout and situated one to the side 
of and slightly behind the other. Female genitalia 
with two strong granulate patches on bursa, a rather 
large one on middle of dorsal surface curving around 
left side onto ventral surface, and a smaller ventral 
patch near junction of bursa and ductus bursae. 

TypE Locatiries: Not given, presumably Pennsyl- 
vania (virgatella, in Acad. Nat. Sci. Philadelphia); New 
York (quinquepunctella, in BM). 

Foop piant: Robinia pseudoacacia. 

Distrisution: UnrTrep States: Maine; Massachusetts, 
Amherst (June), Martha’s Vineyard (June, July, Aug.); 
New York, Orient (Long Island, Aug., Sept.) and one 
specimen with only state locality; New Jersey, Essex 
County Park (Aug.); Pennsylvania, Buena Vista (Aug.); 
New Brighton (May, July); West Virginia, Jefferson 
County (Aug.); District of Oolumbia, Washington 
(June); Virginia, Berryville (May); North Caroline, 
Tryon; ilinois, Elkhart, Putnam County (May); Mis- 
souri, St. Louis; Arkansas, Washington County (‘“‘July— 
Aug.”). Canapa: Ontario, London, Trenton (June); 
Nova Scotia, Bridgetown (July), Smith’s Cove. 

The differences separating virgatella from subcaesiella, 
of which it was long considered only a variety, are dis- 
cussed under the latter species. 


261. Nephopteryx carneella Hulst 
° Figure 823 


Nephopteryx carneella Hulst, Ent. Amer., vol. 3, p. 131, 1887. 

Nephopteryx inquilinella Ragonot, N. Amer. Phycitidae, p. 8, 
1887; Monograph, pt. 1, p. 290, 1893.—Hulst, Ent. Amer., 
vol. 5, p. 156, 1889; Phycitidae of N. Amer., p. 145, 1890.— 
McDunnough, Check list, No. 6171, 1939. 

Salebria carneella (Hulst), Phycitidae of N. Amer., p. 153, 1890.— 
Ragonot, Monograph, pt. 1, p. 367, 1893.—Barnes and Mc- 
Dunnough, Contributions, vol. 3, p. 197, 1916.—Forbes, 
Cornell Mem. 68, p. 627, 1923.—McDunnough, Check list, 
No. 6204, 1939. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 129 


Ground color of forewing bluish gray, the gray shade 
most obvious in median area and as a narrow band 
preceding the inner border of the antemedial line; base 
maroon red, this shade more or less suffusing the gray 
thorax; antemedial line obscure, bordered inwardly by 
a broad madder-red band which extends to costa; a 
similar red shade in outer area bordering the faint 
sinuate subterminal line and some red suffusion over 
median part of the lower fold; discal spots distinct, 
separated, black; terminal dots very faint more or less 
confluent. Hind wing smoky white with a faint yellow- 
ish tint; somewhat darker on females; a fine brown line 
along termen. Alar expanse, 20-23 mm. 

Male genitalia of the fernaldi type; the two cornuti 
lie side by side, one slightly shorter and more slender 
than the other. Female genitalia with bursa and re- 
mainder of genitalia considerably smaller than in pre- 
ceding species; two granulate patches, one large, one 
considerably smaller, placed opposite each other on 
lateral margins of the bursa. 

Typr LocALitiEs: Maine [?] (carneella, in AMNH, 
ex Rutgers; the male type bears no locality label, but 
in his original description Hulst gives New Mexico as 
the type locality; this, as pointed out by Barnes and 
McDunnough, is probably an error or pure guesswork 
on Hulst’s part); Wisconsin (inquilinella, in Paris Mus.). 

Foop piant: Saliz (carneella); galls of sawfly (Huura 
Salicisnodum) on willow. 


Distrisution: Unirep Sratrs: Maine, Monmouth 
(July), and two examples with only state locality; Mas- 
sachusetts, Amherst (June), Martha’s Vineyard (Apr.); 
Indiana, Hessville (June); Wisconsin; Michigan, Dick- 
inson County. Canapa: Ontario, Ottawa (June), 
Trenton (July); Manitoba, Aweme (May). 

Hulst was correct the first time (1889) in making 
inquilinella 2 synonym of his carneella. The genitalia 
of their male types are identical. 


262. Nephopteryx basilaris Zeller 
Figures 341, 829 


Nephopteryx basilaris Zeller, Verh. zool.-bot Ges. Wein, vol. 22, 
p. 548, 1872.—Grote, N. Amer. Ent., vol. 1, p. 51, 1880.— 
Hulst, N. Amer. Phycitidae, p. 145, 1890. 

Salebria basilaris (Zeller) Ragonot, Monograph, pt. 1. p. 353, 
1893.—Forbes, Cornell Mem. 68, p. 627, 1923.—McDun- 
nough, Check list, No. 6209, 1939. 

Forewing pale ash gray to dark gray with a faint 
bluish tint; basal area pale wood brown or pale orange, 
a black shade along its inner margin expanding upward 
at antemedial line to middle of cell; the lower fold in 
basal area streaked with red or reddish brown with 
some scattering of similarly colored scales on the pale 
area above; antemedial line distinct to top of cell, well 
contrasted, grayish white, nearly vertical, inwardly 
dentate between cell and inner margin; margined out- 
wardly by a narrow black line; subterminal line faint 
but distinguishable, sinuate-dentate, followed in outer 
area (on some specimens) by a broad reddish shade; 
discal dots obscured; a row of small blackish dots along 
terminal margin. Hind wings white with a faint ocher- 

300329—56——10 


ous or smoky tint; the veins very faintly darkened; a 
thin brownish line along termen. Alar expanse, 24—27 
mm. 


Male genitalia resembling those of subcaesiella; the 
clasper and lateral elements of transtilla somewhat 
stronger; the smaller cornutus on penis longer and 
stouter but also on some specimens reduced to a mere 
vestige. Female genitalia with two granulate patches 
on ventral surface of bursa, a weaker granulation of the 
surface connecting them. 

TypE Locauity: Massachusetts (type in BM), 

Foop prant: Unknown. 

Distrisution: Unirep Sratres: Maine, Augusta 
(June), Orono; New Hampshire, Hampton (June); Mas- 
sachusetts, Martha’s Vineyard (July), Wilmington 
(June); New York, Catskill Mts., Ilion (June); New 
Jersey, Newark; Illinois, Lacon (June); Indiana, Hess- 
ville (May, July); Michigan, one example, state locality 
only; Colorado, Fort Collins (July), and one example 
state locality only; Utah, Provo Canyon (July). Can- 
ava: Ontario, Budbury, Hymers (July), Trenton (July); 
Manitoba, Aweme (June), McCreary, Winnipeg. 

A strikingly marked species, the most easily identified 
in the genus. The maxillary palpus of the male is, as 
noted by Ragonot (Monograph, p. 354), not strictly in 
the form of an aigrette. The scales are moderately 
long and slender, but not hairlike, intermediate between 
those of a squamous and a typical aigrettelike palpus. 
The labial palpus is grooved on inner surface of the 
second segment as in most species having maxillary 
palpi of the aigrette type. 


263. Nephopteryx termitalis (Hulst), new combination 
Fiaures 342, 828 
Pempelia termitalis Hulst, Trans. Amer. Ent. Soc., vol 13, p. 162, 


Salebria termitalis (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889,—Hulst, Phycitidae of N. Amer., p. 153, 1890. 

Salebria levigatella Hulst, Canadian Ent., vol. 24, p. 61, 1892.— 
Forbes, Cornell Mem. 68, p. 626, 1923.—McDunnough, 
Check list, No. 6207, 1939. (New synonymy.) 

Myrlaea termitalis (Hulst) Ragonot, Monograph, pt. 1, p. 401, 
1893.—McDunnough, Check list, No. 6223, 1939. 

Forewing dark suffused gray (darker than on basilaris) ; 
basal area dull reddish orange with little black on most 
specimens except along costal edge; on others some 
black scaling at extreme base and, narrowly, along 
inner margin; antemedial line nearly obliterated by its 
black borders which are more or less fused and from a 
broad, nearly vertical band from inner margin to costa, 
not strongly contrasted against the dark ground color 
of the wing; subterminal line, discal and terminal dots 
obscure. 

Thorax dark gray, on some specimens more or less 
suffused by the orange color of the basal area of fore- 
wing. One specimen before me (a male from Inyo 
County, Calif.) has the entire thorax and base of fore- 
wing to the black inner border of the antemedial line a 
dull madder-red, and the outer third of wing faintly 
suffused with the same reddish shade. Hind wing dull 


130 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


ocherous white to smoky white. 
mm 


Alar expanse, 23-27 mm. 

Male genitalia with apical process of gnathos tri- 
angulate. Transtilla weak but distinguishable. Clasper 
erect and slightly curved, slender, digitate. One 
moderately sized and one much smaller cornutus on 
penis. Female genitalia with a single moderately large 
granulate patch on posterior lateral corner of bursa and 
extending in weaker granulation transversely across the 
middle of the lower surface. 


Typ Locatitizs: Colorado (fermitalis, in AMNH, 
ex Rutgers); “Amherst, Massachusetts” [sie] (leviga- 
tella, in AMNH, ex Rutgers). 

Foop prant: Unknown. 


DistRipuTion: UNITED States: Colorado, Glenwood 
Springs, Gunnison County near Altmont, and two 
examples (c', 2) with only the state locality; Utah, 
Spanish Fork (July); Arizona, Prescott (June); Cah- 
fornia, Inyo County (June), Placer County (June). 
Canava: Ontario, Trenton (July), Manitoba, Winni- 
peg; Alberta, Bilby (June); British Columbia, Clinton 
(June). 

Hulst also gives Massachusetts and Wisconsin as 
localities for his levigatella. His type of the latter how- 
ever has no locality label. A female of levigatella from 
the Fernald Collection is in the National Collection. 
It also bears a Hulst “type” label, but no locality or 
date. The type of termitalis is a male, not a female as 
given in Hulst’s original description. Genitalia of 
conspecific females from western localities agree in every 
detail with those of the type of levigatella so there can 
be no question of the synonymy of the two Hulst names. 

The labial palpi appear a trifle oblique (less tightly 
appressed to the face than on most of the preceding 
species) which may account for Ragonot’s reference of 
the species to Myrlaea. 


Alar expanse, 23-27 


264. Nephopteryx termitalis yuconella (Dyar), new status 


Salebria yuconella Dyar, Ins. Insc. Menstr., vol. 13, p. 12, 1925.— 
McDunnough, Check list, No. 6194, 1939. 

A slightly darker more suffused local race of termitalis; 
the basal area of forewing almost entirely suffused with 
blackish except for a narrow grayish white shade along 
inner margin of the black inner border of the antemedial 
line; no appreciable red or orange shading anywhere on 
forewing or thorax. Alar expanse, 25-27 mm. 

Genitatta: Male and female as in typical termitalis. 


Typz tocatity: Near Fort Yukon, Alaska (type in 
USNM). 


Foop rirant: Unknown. 


Distrisution: ALasKa: Dawson (June), Fort Yukon. 

Specimens of yuconella as of typical termitalis show 
a strong tendency to become greasy with age, indicating 
that their larvae are borers; but nothing is known about 
the biology of either form. 


265. Nephopteryx bifasciella Hulst 
Figures 343, 831 


Nephopteryx bifasciella Hulst, Ent. Amer., vol. 3, p. 132, 1887. 

Salebria bifasciella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 366, 1893.—Hulst, Phycitidae 
of N. Amer., p. 154, 1890.—Barnes and McDunnough, 
Contributions, vol. 3, p. 197, 1916.—McDunnough, Check 
list, No. 6214, 1939. 

Salebria nogalesella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 35, 1905.—MecDunnough, Check list, No. 6215, 1939. 
(New synonymy.) 

Forewing ash gray with a fine powdering of black 
scales, giving the wing a faint pale bluish tint; the black 
borders of the transverse lines strongly contrasted; 
antemedial line distinct, at least from below upper vein 
of cell, narrow, slightly oblique and somewhat angulate 
at middle, its outer black border complete, slightly 
broadened at costa, its inner black border narrow, 
extending from inner margin only to top of cell; sub- 
terminal line sinuate, bordered inwardly by a narrow 
black line, and outwardly by a much fainter, paler 
dark line; discal dots usually distinguishable but faint, 
separate or confluent (sometimes both ways on the 
same specimen); dots along terminal margin very 
faint, more or less confluent. Hind wing white with a 
very faint ocherous or smoky tint, slightly darker on 
female than on male. Alar expanse, 20-22 mm. 

Male genitalia with apical process of gnathos tri- 
angulate. Clasper sharply curved and running close 
and parallel to surface of harpe, simple and strongly 
sclerotized. Cornuti rather short, but individually 
variable in size. There is also some slight variability 
in the size and shape of the apical process of gnathos. 
The genitalia of the type of bifasciella and those of its 
synonym nogalesella are more nearly alike than those 
of any other two males before me. Female genitalia 
with bursa nearly round, armed with two large granulate 
patches, arranged as in figure 831 but with the position 
of the anterior patch (at closed end of bursa) somewhat 
variable. In one specimen from Palmerlee there is a 
third patch on the right side of bursa and the bursa 
itself is narrower and considerably elongated. ‘These 
differences probably represent nothing more than 
individual aberrations. Males from the same locality 
are normal. 

Type Locauities: Arizona (bifasciella, in AMNH, 
ex Rutgers); Nogales, Ariz. (nogalesella, in USNM). 

Foop prant: Unknown. 

DistRiBuTIon: Arizona, Baboquivari Mts. (July), 
Huachuca Mts. (July, Aug.), Nogales (July), Palmerlee, 
Redington, ‘‘Southern Arizona” (Aug.), and two ex- 
amples with only the state locality. 


266. Nephopteryx uvinella (Ragonot), new combination 
Fieures 344, 824 


Meroptera uvinella Ragonot, N. Amer. Phycitidae, p. 8, 1887; 
Monograph, pt. 1, p. 315, 1893.—Hulst, Phycitidae of N. 
Amer., p. 148, 1890.—McDunnough, Check list, No. 6183, 
1939. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 131 


Salebria afflictella Hulst, Canadian Ent., vol. 32, p. 170, 1900.— 
Forbes, Cornell Mem. 68, p. 625, 1923.—McDunnough, 
Check list, No. 6200, 1939. (New synonymy.) 

Meroptera liquidambarella Dyar, Proc. Ent. Soc. Washington, 
vol. 6, p. 108, 1904. 

Meroptera afflictella (Hulst) Dyar, Proc. Ent. Soc. Washington, 
vol. 7, p. 34, 1905. 

Forewing fuscous (gray-brown), the median area 
heavily dusted with white giving it a pale ash gray 
color; basal area of the ground color, rarely with some 
red scaling in the median fold; dark base followed by 
an oblique, straight, whitish band; this latter bordered 
outwardly by a broad, somewhat diffused, dark brown 
or blackish band through which may be distinguished 
faint white traces of the true antemedian line; subter- 
minal line with a very slight central bulge, rarely some- 
what crenulate, whitish gray, bordered inwardly by a 
diffused brown shade and outwardly by a narrow brown 
line; discal dots more or less fused; blackish terminal 
dots confluent. Hind wings pale to dark smoky fus- 
cous. Alar expanse, 14-18 mm. 


Male genitalia distinguished chiefly by the armature 
of the penis. On the paratype of afflictella from Mont- 
clair, N. J., there appears to be two cornuti, one closely 
appressed to the other; but in other preparations before 
me (including the types of wvinella and liquidambarella) 
the two cornuti are completely fused into a single rather 
short and stout, longitudinally ribbed cornutus. The 
clasper is a slender, curved, sharply pointed, smooth, 
sclerotized hook. Vestiges of a divided transtilla 
distinguishable on most preparations. 


Female genitalia small (approximately the size and 
form of those of carneella); bursa with two moderately 
sized granulate patches, the bursal lobe giving off the 
ductus seminalis also granulate and partially (smoothly) 
sclerotized; this lobe as usual arises from the dorsum of 
bursa but is deflected to the left. 


TYPE LOCALITIES: United States (wvinella, in USNM) 
Elizabeth, N. J. (affictella, in AMNH, ex Rutgers); 
Washington, D. C., (liquidambarella, in USNM). 

Foop piant: Liquidambar styraciflua (sweetgum). 
Larva a leaf-tier. 


DistrisutTion: Connecticut, East River (May), Stam- 
ford (Aug.); New Jersey, Elizabeth (Aug.), Lakeland 
(May), Montclair (June, Aug.), New Lisbon (June); 
District of Columbia, Washington (May, Aug.) ; Virginia, 
Skyland (July); North Carolina, Greensboro (Aug.); 
Georgia, Savannah (Sept.); Florida, Lakeland (May, 
June). 


Ragonot’s wvinella has been an unknown entity ever 
since its description and was suspected of being only a 
variety of Meroptera pravella. Fortunately the type 
was secured by Dr. Barnes when the Oberthiir Collec- 
tion was sold. It lacks one forewing but is otherwise 
intact and its habitus and genitalia leave no doubt that 
it is the same as the sweet-gum feeder described by 
Hulst and Dyar. The type of affictella is a female. 
The types of uvinella and liquidambarella are both males. 
The genitalia of all of them are before me. 


267. Nephopteryx celtidella (Hulst), new combination 
Figures 345, 830 


Salebria celtidella Hulst, Phycitidae of N. Amer., p. 155, 1890.— 
Beutenmiller, Canadian Ent., vol. 22, p. 17, 1890 (larva).— 
Ragonot, Monograph, pt. 1, p. 354, 1893.—Forbes, Cornell 
Mem. 68, p. 626, 1923.—McDunnough, Check list, No. 
6206, 1939. 

Ground color ocherous (clay color) more or less shaded 
with fuscous gray over submedian and (on especially 
dark females) outer basal areas; the ground color espe- 
cially well contrasted at base and in the central area 
about the discal spots, also on thorax, at extreme base 
of wing and on thorax sometimes of a tawny shade; ante- 
medial line far out towards middle of wing, oblique, sinu- 
ate, faint, indicated chiefly by its narrow black borders 
which are more or less broken into dots on the veins; 
subterminal line sinuate-serrate, bordered inwardly by 
a row of black (somewhat confluent) dots and outwardly 
by a row of small black wedges on the veins; discal dots 
distinct, well separated, black; a row of distinct black 
dots along termen. Hind wing pale to rather dark 
smoky fuscous. Alar expanse, 18-22 mm. 

Male genitalia with apical process of gnathos smaller 
than that of preceding species (not triangulate). Clasper 
rather short, bent across surface of harpe, blunt. Sclero- 
tized lateral elements of transtilla distinguishable, 
rather long, slender. 

Female genitalia with a round, moderately large 
granulate patch on ventral surface of bursa and a smaller 
patch near junction of bursa and ductus bursae and the 
base of the lobe giving off the ductus seminalis. 

Typr Locatity: New York (in AMNH, ex Rutgers). 

Foop puant: Celtis. Larva a leaf-tier. 

Distripution: Unitep Starrs: New York, Long 
Island; Maryland, Plummers Isl. (May, July); Flor- 
ida, Palm Beach (Feb.); Texas, Brownsville (July), Vic- 
toria (May), Zavalla County (Apr.); Mississippi, “Agr. 
College” (Apr.), Starkville (July); Missouri, St. Louis 
(Aug.); Zilinois, Oconee (Aug.). 

A distinct species distinguished from any of the spe- 
cies with aigrettelike maxillar palpi on the male by its 
clay-colored, black-mottled forewings. Its habitus is 
nearest to that of rubrisparsella in the group with 
squamous, male maxillary palpi. 


Genus Nephopteryx, Species 268-271: N. rubri- 
sparsella to N. bisra 


[Males with squamousmaxillary palpi.] 


268. Nephopteryx rubrisparsella (Ragonot) 
Ficures 346, 832 


Pristrophora rubrisparsella Ragonot, N. Amer. Phycitidae, p. 6, 
87 


Pristophora rufibasella Ragonot, N. Amer. Phycitidae, p. 7, 1887. 

Sciota croceella Hulst, Ent. Amer., vol. 4, p. 115, 1888. 

Nephopteryx rubrisparsella (Ragonot), Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 284, 1893.—Hulst, Phycitidae 
of N. Amer., p. 145, 1890.—Forbes, Cornell Mem. 68, p. 623, 
1923.—McDunnough, Check list, No. 6169, 1939. 


132 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Psorosa texanella Hulst, Canadian Ent., vol. 32, p. 174, 1900. 
Hulstea texanella (Hulst), U.S. Nat. Mus. Bull. 52, p. 432, 1903.— 
MeDunnough, Check list, No. 6342, 1939. (Newsynonymy.) 
Similar to celtidella in color and maculation except: 
Ocherous basal area of forewing more or less shaded with 
reddish purple and similar suffusions over much of the 
median and outer areas; also a broad blackish suffusion 
over the area of the antemedial line and extending some- 
what beyond it; the inner dark border of subterminal 
line more nearly continuous and the outer bordering 
dashes much fainter except in dark and very well 
marked examples. Alar expanse, 17-20 mm. 


Male genitalia figured from type of texanella Hulst, 
which agree in every detail with those of typical rubri- 
sparsella and of the male type of croceella. ‘They differ 
from those of celtidella in having a longer, more strongly 
sclerotized clasper, and different cornuti (shown in lat- 
eral view in our figures). The cornuti lie side by side 
in both celtidella and rubrisparsella, but in the former 
species they are of equal length while in rubrisparsella, 
one cornutus is much shorter than the other—not too 
reliable a character, but apparently consistent here. 
Transtilla, on dissection, distinguishable as a complete 
band, but central area very weak and in balsam prepa- 
ration only the well sclerotized lateral elements easily 
seen. 


Female genitalia with a broad granulate band par- 
tially encircling middle of bursa, similar to that in sub- 
fuscella except that in subfuscella it is chiefly over the 
ventral surface of bursa while in rubrisparsella it is on 
the dorsal. 


TYPE LOCALITIES: United States (rubrisparsella, in 
Paris Mus.); Texas (rufibasella, in Paris Mus.); Blanco 
County, Tex. (croceella and texanella, in AMNH, ex 
Rutgers). 

Foop pnant: Celtis. 


Distrisution: Texas, Black Jack Springs, Blanco 
County, Kerrville (June); Oklahoma; Missouri, St. 
Louis (June); Jilinois, Lacon (July), Putnam County 
(uly); West Virginia, Jefferson County (Aug.); Mary- 
land, Plummers Isl.; District of Columbia, Washington 
(May, June). 

Very close to celfidella and distinguished from it 
chiefly by its genitalia and squamous male maxillary 
palpi. Hulst’s texanella was described from an ab- 
normal specimen with vein 4 lacking in the hind wing. 
I have before me two similar abnormal females (from 
Oklahoma and Illinois) and in the following species 
(gilvibasella) a couple of examples, out of a long series 
of normal specimens, that also have vein 4 absent. 

The type of croceella in addition to the Hulst name 
label also bears the following label in Ragonot’s hand- 
writing: ‘‘Nephopteryx rubrisparsella Rag.—rufibasella 
Rag.” 

269. Nephopteryx gilvibasella Hulst 
Figure 836 
Nephopteryx gilvibasella Hulst, Phycitidae of N. Amer., p. 145, 


1890.—Ragonot, Monograph, pt. 1, p. 289, 1893.—McDun- 
nough, Check list, No. 6172, 1939. 


Salebria lacteella Hulst, Canadian Ent., vol. 32, p. 71, 1900.— 
cia and McDunnough, Contributions, vol. 3, p. 197, 
1916. 

Much paler than the preceding species (rubrisparsella) 
and without its blackish shadings. Ground color pale 
gray with a very faint bluish tint; extreme base of wing 
ocherous orange, this shade extended somewhat on 
inner margin and indicated on many specimens in 
median area over the lower fold; a broad band of the 
same color immediately preceding the antemedial line 
and extending from inner margin to middle of cell; 
above it a blackish shade extending to costa; lower half 
of antemedial line clearly indicated, narrow, whitish, 
bordered outwardly by a narrow black line which fuses 
towards costa into the blackish shade above the ocher- 
ous patch; subterminal line faintly bordered inwardly 
by a faint blackish line; discal dots small, separated, 
sometimes obscure, but usually distinct; a row of weak 
but discernible blackish dots along termen. Hind wing 
subpellucid; whitish with a very faint ocherous tint; a 
weak, pale brown line along termen; the veins not 
appreciably darkened. Alar expanse, 17-20 mm. 

Male genitalia similar to those of rubrisparsella. The 
eighth-segment tufts much simpler, reduced to two 
lateral pairs with scales of a uniform shape and size. 

Female genitalia with a broad granulate band on 
bursa similar to that in rubrisparsella; bursa bulged into 
a small, strongly granulate lobe at its left posterior 
angle (this lobe opposite to that giving off ductus 
seminalis) ; also a narrow row of moderately long spines 
across upper side of bursa at junction of bursa and 
ductus bursae. 


TyprE Locatitizs: Not given (gilvibasella, in AMNH, 
ex Rutgers); Blanco County, Tex. (lacteella, in AMNH, 
ex Rutgers). 


Foop prant: Unknown. 


Distrisution: Texas, Blanco County (Sept.), 
Brownsville (Mar., July), San Benito (Mar., Apr., 
May, July, Aug., Sept.). 

The species is close to but distinct from rubrisparsella. 
The types of gilvibasella and lacteella are both females 
with identical genitalia. A long series from Browns- 
ville and San Benito are before me. 


270. Nephopteryx crassifasciella Ragonot 
Fieurts 347, 835 


Nephopteryx crassifasciella Ragonot, N. Amer. Phycitidae, p. 8, 
1887; Monograph, pt. 1, p. 285, 1893—Hulst, Phycitidae 
of N. Amer., p. 146, 1890.—MceDunnough, Check list, No. 
6170, 1939. 

Nephopteryx decipientella Dyar, Proc. Ent. Soc. Washington, vol. 
7, p. 34, 1905.—McDunnough, Check list, No. 6162, 1939. 
(New synonymy.) 

Nephopteryx crataegella Barnes and McDunnough, Contributions, 
vol. 3, p. 222, 1917—McDunnough, Check list, No. 6164, 
1939. (New synonymy.) 


Forewing ash gray more or less suffused by somber 
purplish or gray brown, shading into blackish brown on 
the borders of the transverse lines; the pale color limited 
to a rather narrow band preceding the inner border of 
the antemedial line, part of the median area following 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 133 


the antemedial line and surrounding the discal dots, 
and some faint pale dusting immediately preceding 
terminal margin; extreme base of wing purplish brown 
with a shading (in some specimens) of reddish scales, 
especially towards inner margin; antemedial line pale 
gray, distinguishable only from inner margin to middle 
of cell, bordered inwardly by a broad, vertical, blackish 
brown band which extends to costa, and outwardly by 
a narrow black line which fuses into the inner dark 
border towards costa; subterminal line sinuate, pale 
gray with narrow, black-brown borders; discal dots 
distinct, sometimes partially fused. Hind wing light 
to dark smoky brown; the veins slightly darkened and 
a dark line along termen. Alar expanse, 16-17 mm. 

Male genitalia with clasper short, blunt, curved. 
Cornuti moderately long, approximately the same size 
and lying side by side. A long slender arm projecting 
from each dorsolateral angle of vinculum. Female 
genitalia, with two large, opposing, roundly oval, gran- 
ulate patches on bursa, and some rather strong, darkly 
pigmented spining on the lobe giving off the ductus 
seminalis. 

Type LocauitiEs: California [?] (crassifasciella, lost?) ; 
unknown (decipientella, in USNM); Lakeland, Fla. 
(crataegella, in USNM). 


Foop piants: Vaccinium, Crataegus. 


Distrisution: Maryland, Plummers Isl. (May); 
District of Columbia, Washington (May) ; Georgia, Sape- 
loe (Sept.); Florida, Lakeland (May). 

A male from the District of Columbia (reared from 
Vaccinium by Chambliss, May 10, 1895) and a similar 
female from Plummers Isl., Md., in the National Col- 
lection agree in every detail with Ragonot’s figure and 
description of crassifasciella and were so identified by 
Dyar. His decipientella was described from a single 
specimen without locality label. It is merely a dark- 
suffused male with the pale coloration limited to a more 
restricted area about the discal spots and some light 
dusting between the outer veins. The Georgia example 
(reared from Vaccinium) is a female and matches the 
type of decipientella except for venation. It is another 
of the all too frequent phycitid freaks with vein 4 miss- 
ing from hind wing and 7-8 long stalked. The Barnes 
and MeDunnough type of crataegella (also a male and 
reared from Crataegus) differs from typical crassifasciella 
only in a somewhat more extended and lighter colora- 
tion of the pale areas of forewing. All three males 
before me agree in genitalic structure. 

The species is easily identified by the peculiar develop- 
ment of the vinculum, not found in any other American 
species in the genus. 

Dr. Bourgogne informs me that the type of inquilinella 
could not be found at Paris. It is probably lost, and I 
suspect that the California citation in Ragonot’s Mono- 
graph was probably a guess. I have seen nothing from 
the Pacific Coast States that could possibly be his 
species. 


271. Nephopteryx bisra Dyar 
Fievurse 833 
Nephopteryx bisra Dyar, Ins. Inse. Menstr., vol 7, p. 51, 1919. 


Forewing pale gray-brown; base shaded with black- 
ish; a straight, narrow, oblique whitish line along inner 
margin of the inner border of the antemedial line, the 
latter a thin, oblique, notched, whitish line, obscured 
towards costa; its inner border a wide blackish band 
reaching to costa; its outer border a narrow black line; 
a pale patch on inner margin following the antemedial 
line; subterminal line distinct, whitish gray, sinuate, 
margined by narrow blackish brown inner and outer 
lines; discal spots fused into a lunate mark with a pale 
surrounding shade; a blackish line along termen. Hind 
wing pale gray-brown, paler than ground color of fore- 
wing; veins not appreciably darkened; a narrow, dark 
(brownish) line along termen. Alar expanse, 21 mm. 

Female genitalia with a large granulate patch cover- 
ing most of dorsal surface of bursa, a smaller latero- 
ventral patch and some scattered granulations at left, 
posterior angle on ventral side; bursa otherwise rather 
densely and finely spinose. 

Typs Locauity: Orizaba, México (type in USNM). 

Foop prant: Unknown. 

Known only from the female type. The placement 
in the second group of Nephopteryz is provisional, pend- 
ing discovery of a male. In the type, vein 10 of fore- 
wing is shortly stalked with the stem of 8-9. 


69. Genus Tlascala Hulst 


Tlascala Hulst, Phycitidae of N. Amer., p. 146, 1890.—Forbes, 
Cornell Mem. 68, p. 623, 1923. (Type of genus: Nepho- 
pleryx reductella Walker.) 


Tongue well developed. Antenna weakly pubescent; 
on male, with sinus and enlarged scale tuft in base of 
shaft. Labial palpus upturned, not appressed to face, 
cylindrical, smooth scaled, reaching above vertex; 
second segment of male not hollowed; third segment 
moderately long, acuminate. Maxillary palpi of both 
sexes squamous, appressed to face. Forewing with 
most of inner border of antemedial line consisting of 
raised scales; 11 veins; vein 2 from near lower outer 
angle of cell; 3 from the angle, about equidistant from 
2 and 4; 4 and 5 very shortly stalked or connate, rarely 
(in individual specimens) closely approximate at base; 
6 from below upper angle of cell, straight; 8-9 long 
stalked (for three-fourths of their lengths) ; 10 from the 
cell approximate to or connate with the stalk of 8-9 at 
base, and approximate to it for some distance beyond 
base; male without costal fold. Hind wing venation 
similar to that of Meroptera and Nephopteryx. Highth 
abdominal segment of male with compound scale tufts. 

Male genitalia with strongly sclerotized, broadly tri- 
angulate clasper with a row of irregular teeth along its 
outer margin. Penis armed with two, equally sized, 
stout, sharply curved, thornlike cornuti. Otherwise 
as in Nephopteryz. 


134 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Female genitalia without signum; bursa finely and 
densely spined over most of inner surface and with a 
cluster of longer and stronger spines near middle of 
lateral margin; ductus bursae short (less than half the 
length of bursa); armed on ventral surface by an elon- 
gate pair of granulate plates; at genital opening a 
strongly sclerotized, centrally interrupted genital plate, 
attached to a narrow, sclerotized collar (incomplete 
dorsally and ventrally) and supplemental to the regular 
eighth-segment collar; ductus seminalis from bursa near 
junction of bursa and ductus bursae. 

The genus is close to and has several characters in 
common with both Meroptera and Nephopteryz, resem- 
bling the former in the well developed genital plate of 
the female genitalia and the strong, serrate clasper of 
the male, differing from both genera in the rough scaling 
on forewing, the ungrooved labial palpi and much 
stouter antennal tuft of the male, the sharply curved 
cornuti on penis, and the supplemental collar attached 
to the female genital plate. As here defined it contains 
only its American type species. 


272. Tlascala reductella (Walker) 
FIGURES 28, 348, 834 


Nephopteryx reductella Walker, List, vol. 27, p. 638, 1863.— 
Ragonot, Ent. Amer., vol. 5, p. 115, 1889; Monograph, pt. 1, 
p. 283, 1893. 

Pempelia gleditschiella Fernald, in Comstock, in Rep. [U. §.] 
Comm. Dep. Agr. for 1880, p. 262, 1881.—Packard, U. S. 
Dep. Agr. Fifth Rep. Ent. Com., p. 652, 1890. 

Tlascala reductella (Walker) Hulst, Phycitidae of N. Amer., p. 
146, 1890.—Forbes, Cornell Mem. 68, p. 624, 1923.— 
McDunnough, Check list, No. 6180, 1939. 

In color and maculation similar to the Kuropean 
Nephopteryz rhenella except for the raised scaling of the 
subbasal black bar. 

Forewing dull ash gray with purplish brown shading 
at extreme base and paler brownish shading on outer 
median and terminal areas; antemedial line complete 
but faint, narrow, oblique, slightly indented on lower 
half, preceded by a broad black band of rough scales 
and followed by a narrow black band; subterminal line 
obscure, sinuate-dentate, faintly bordered by dark inner 
and outer lines; discal dots distinct, blackish, separated; 
a row of faint blackish dots along termen. Hind wing 
pale smoky fuscous; the veins very slightly darkened 
and a narrow dark line along termen. Alar expanse, 
17-23 mm. 

Male genitalia with cornuti set side by side near outer 
end of penis, their apices turned away from each other. 
Female genitalia as given for the genus. 

TypE LOCALITIES: Honduras [sic] (reductella, in BM); 
District of Columbia (gleditschiella, in USNM). 

Foop Piant: Gleditsia. Larva a leaf-tier. 

DistrisutTion: District of Columbia (May, July); 
Maryland, Plummers Isl. (June); North Carolina, Hil- 
tonhead Isl. (Aug.); Pennsylvania; Illinois, Decatur 
(Mar., Apr., May, July), Quincy (May); Jowa, Ames; 
Kansas, Lawrence (May); Missouri, St. Louis (May); 
Texas, Kenedy (Apr.), Paris (May), Victoria (July), 


Zavalla County (Apr.); Louisiana, Crown Point (June, 
larva), New Orleans (larva, June). 

Walker gives Honduras as the type locality. I 
rather doubt the correctness of this citation, for I have 
never seen anything from Central America that even 
remotely resembled the species. I have not seen his 
types, but have no reason to question the correctness of 
Ragonot’s reference of gleditschiella to synonymy. 


70. Tulsa, new genus 


Type of genus: Nephopteryx finitella Walker. 

Characters of Tlascala except: 

Forewing with some rough scaling in the median 
area beyond the outer margin of the antemedian line, 
sometimes a small tuft on lower fold, always a few 
roughened scales in the discal spots; veins 4—5 approxi- 
mate at base and for a short distance beyond. Male 
genitalia with sacculus of harpe considerably enlarged, 
strongly sclerotized, densely and finely spined along 
entire lower margin, and produced at apex; clasper a 
thin, dished plate produced into an elongate, curved, 
sharply pointed claw at each lower angle. Uncus 
broadened at apex. Transtilla complete, very weakly 
but evenly sclerotized throughout. Cornuti straight, 
set one before the other. Female genitalia with several 
lines of fine spines running from bursa into ductus 
bursae almost to genital opening; genital plate and its 
attached supplemental collar strongly wrinkled (more 
so than shown in the figures) ; no granulations in ductus 
bursae. 

The genus is very close to Tlascala and I propose the 
new name with some misgiving; but the habitus of the 
moths and their genitalia, male and female, differ so 
much from those of the type of Tiascala that something 
more than a species-group difference is indicated. When 
larvae and host relations of the Tulsa species are known 
we shall probably find additional supporting characters 
for the genus. Specifically the genitalia are remarkably 
similar, offering little or nothing to distinguish the 
species. Four are here recognized. 


273. Tulsa finitella (Walker), new combination 
Figure 349 


Nephopteryx finitella Walker, List, vol. 27, p. 53, 1863.—Ragonot, 
Monograph, pt. 1, p. 282, 1893. 

Tlascala finitella (Walker) Hulst, Phycitidae of N. Amer., p. 147, 
1830.—Forbes, Cornell Mem. 68, p. 624, 1923.—McDun- 
nough, Check list, No. 6177, 1939. 

Elasmopalpus melanellus Hulst, Phycitidae of N. Amer., p. 157, 
1890.—Barnes and McDunnough, Contributions, vol. 3, 
p. 199, 1916 (makes synonym of finitella). 


Forewing very dark gray with the blackish borders of 
the antemedial and postmedial lines but slightly con- 
trasted; some slight dusting of white on subbasal, me- 
dial, and terminal areas; antemedial line somewhat 
stronger, its inner blackish bordering line more or less 
interrupted on the veins, its outer border continuous 
but faint; discal dots tending to coalesce; an obscure 
row of blackish dots along termen; raised scales con- 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 135 


spicuous on lower half of inner border of antemedial 
line and as a patch on middle of lower fold; a raised 
scale or two in the discal spots. Hind wing pale smoky 
fuscous with a darker shading towards termen; the 
veins slightly darkened; the entire wing darker on 
northern examples. Alar expanse, 21-25 mm. 

Male genitalia with no distinguishing specific features. 
Female genitalia similar to those of wmbripennis but 
smaller (about the size of infinitella) and with more 
decided wrinkling of the genital plate and its supple- 
mental collar. 

Typr LocatitiEs: United States (jinitella, in BM); 
Florida (melanellus, in AMNH, ex Rutgers). 

Foop prant.—Blueberry. Thisrecord isfromareared 
female (Brunswick, Ga., Quaintance No. 31501) in the 
National Collection. The larva, however, may have 
been accidentally on that plant. 

DistrisutTion: Unirep States: Florida, Charlotte 
Harbor (Mar.), Fort Myers (Apr.), Miami (Mar.), St. 
Petersburg (Apr.), Tampa, also examples with only 
state locality; Georgia, Brunswick (June); North Caro- 
lina, Raleigh (June); Virginia, Richmond (May); Dis- 
trict of Columbia, Washington (May, June); New Jersey, 
Essex County (May), Newark (May); Massachusetts, 
Martha’s Vineyard (Aug.); Jndiana, Hessville (June). 
Canava: Ontario, Trenton; Quebec, Kazubazua (June). 
Walker also reports the species from Nova Scotia. 

The type of melanellus in the Rutgers Collection is a 
female without abdomen. Matching cotypes (o and 9) 
from Mrs. Slosson’s material are in the National Collec- 
tion. There can be no question of the synonymical 
reference by Barnes and McDunnough. 


274. Tulsa umbripennis (Hulst), new combination 
Fieures 350, 842 


Pinipestis umbripennis Hulst, Canadian Ent., vol. 27, p. 57, 1895. 

Ortholepis gillettella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 107, 1904. 

Tlascala umbripennis (Hulst) Barnes and McDunnough, Contri- 
butions, vol. 3, p. 195, 1916.—McDunnough, Check list, 
No. 6178, 1939. 

Fore and hind wings dark brown with a somewhat 
glossy sheen not possessed by the other species of the 
genus; discal dots on forewing confluent, forming a bar 
on discocellular vein; raised scales and maculation as in 
Jjinitella. Alar expanse, 25-26 mm. 

Male genitalia, figured from type of gillettella, agree 
in every detail with those of the male type of umbri- 
pennis. Female genitalia with bursa copulatrix 
slightly larger than that of any of the other Tulsa 
species except oregonella. 

TypEtocauitizs: Colorado (umbripennis, in AMNH, 
ex Rutgers; gillettella, in USNM). 

Foop prant: Unknown. 

Known only from the type locality. A series before 
me is from Chimney Gulch, Colo. (June and July). 
The types of umbripennis and gillettella have only the 
state locality, but the latter was probably from the 
neighborhood of Fort Collins. 


275. Tulsa oregonella (Barnes and McDunnough), new combination 
FicurE 351 


Tlascala oregonella Barnes and McDunnough, Contributions, 
ve p. 175, 1918.—McDunnough, Check list, No. 6179, 

Forewing a dull, powdery, slate gray; basal area a 
trifle paler; the transverse lines somewhat more distinct 
and better defined than on wmbripennis; inner black 
border of antemedial line slightly narrower than in 
preceding species; dark borders of subterminal line very 
faint; discal dots separated. Hind wing smoky gray- 
brown, paler than that of wmbripennis and not glossy. 
Alar expanse, 26-28 mm. 

Female genitalia similar to those of wmbripennis. 

Type Locauity: Crater Lake, Oreg. (July; type in 
USNM). 

Foop puant: Unknown. 

Known only from the type series. 


276. Tulsa infinitella (Dyar), new combination 
Figure 841 


Tlascala infinitella Dyar, Ins. Insc. Menstr., vol. 7, p. 52, 1919. 


Similar to oregonella except: Forewing slightly darker; 
antemedial line obscure; the discal dots confluent, form- 
ing a blackish line of slightly roughened scales. Hind 
wing as in southern specimens of finitella; pale smoky 
fuscous at base with a darker shading towards termen. 
Alar expanse, 27 mm. 

Female genitalia like those of finitella except lateral 
elements of genital plate smoother. 

Tyrer Locauity: Orizaba, México (type in USNM). 

Foop piant: Unknown. 

Known only from the female type. 


Genus 71: Homoeographa 


[Venational division C. veins 4 and 5 shortly stalked or weakly 
anastomosed for a short distance beyond cell. Hind wing with 
discocellular vein of cell vertical, straight. Antenna of male 
with sinus and scale tuft in base of shaft. Labial palpus obliquely 
upturned, second segment laterally flattened and broadly scaled, 
on male grooved to hold maxillary palpus. Maxillary palpus of 
male in the form of an aigrette. Highth abdominal segment of 
male with compound scale tufts. Male genitalia without trans- 
tilla; clasper present, digitate; penis armed with two moderately 
stout cornuti. Female genitalia without signum; bursa with 
deep, convoluted, sclerotized folds; genital opening simple.] 


71. Genus Homoeographa Ragonot 


Homoeographa Ragonot, Nouv. Gen., p. 24, 1888; Monograph, 
pt. 1, pp. xlvi, 432, 1893. (Type of genus: Homoeographa 
lanceolella Ragonot.) 

Tongue well developed. Antenna pubescent; male 
with sinus and strong tuft in base of shaft. Labial 
palpus obliquely upturned, reaching well above vertex; 
second segment flattened and broadly scaled, on male 
grooved to hold the maxillary palpus; third segment 
short, partially hidden in scaling of second. Maxillary 
palpus of male in the form of an aigrette; of female 


136 


squamous. Forewing smooth; 11 veins; vem 2 from 
before the lower outer angle of cell; 3 from the angle, 
separated from 4-5; 4 and 5 shortly stalked or weakly 
anastomosed for less than half their lengths from cell; 
6 from below upper angle of cell, slightly bent at base; 
8 and 9 stalked for over half their lengths; 10 from the 
cell, connate or closely anastomosed at base with the 
stalk of 8-9. Hind wing with vein 2 from well before 
lower outer angle of cell; 3 from the angle, long; 4 and 5 
stalked for about four-fifths of their lengths; 7 and 8 
anastomosed for three-fourths of their lengths; cell 
short, one-third the length of wing; discocellular vein 
vertical, straight. Highth abdominal segment of male 
with compound ventral scale tufts. 

Male genitalia with uncus subtriangulate and with 
blunt, moderately broad, notched apical margin. Apical 
process of gnathos a moderately stout hook with a 
slender, digitate basal projection. Transtilla absent. 
Harpe with costa sclerotized throughout, but not 
produced at apex; cucullus simple, curved, apex 
bluntly pointed; clasper present, simple, erect, digitate. 
Penis armed with two moderately stout cornuti, about 
one-third as long as aedeagus. Vinculum about twice 
as long as broad, evenly tapering to its truncate, 
strongly sclerotized, anterior margin. Vinculum U- 
shaped with somewhat enlarged base. 

Female genitalia with bursa elongate, narrow, signum 
absent, two or three deep, convolute sclerotized folds 
at posterior half, the sclerotization extending for a 
short distance into ductus bursae; genital opening 
simple. Ductus seminalis from bursa, near junction of 
bursa and ductus bursae. 

The genus is easily distinguished by its hind wing 
venation and genitalia. The male genitalia indicate 
a close relationship to the Nephopteryx group of genera, 
However the characteristic hair brush on the harpe of 
the latter are absent from Homoeographa. It contains 
only one known tropical American species. 


277. Homoeographa lanceolella Ragonot 
Ficures 352, 839 


Homoeographa lanceolella Ragonot, Nouv. Gen., p. 25, 1888; 
Monograph, pt. 1, p. 483, 1893. 

Forewing gray heavily dusted with white on costal 
half; antemedial line indicated by a narrow, white, in- 
wardly notched line between cell and inner margin and 
above that by its incompleted outer border, a black 
line oblique from costa to cell thence inwardly angled to 
lower vein of cell; subterminal line faint, sinuate, in- 
dicated chiefly by blackish gray bordering streaks from 
costa, the inner one the longer and continued as a weak 
blackish shading to inner margin; discal dots small, 
separated, blackish gray; in outer area black streaklets 
bordering vein 1b above and veins 3 and 6 below. 
Hind wing semitranslucent smoky white; the veins 
slightly darkened and a narrow dark line along termen. 
Head ashy white. Alar expanse, 21 mm. 

Female genitalic characters as given for the genus. 

TypE LocaLity: Callao, Peri (type in Paris Mus.). 

Foop piant: Unknown. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Known only from the type series in the Muséum 
National d’Histoire Naturelle, Paris, and the British 
Museum. 


Genera 72-76: Telethusia to Pyla 


[Venational division B. Veins 4 and 5 of forewing usually 
separated at base (shortly stalked in Actriz); 10 from the cell, 
separated at base from stalk of 8-9. Hind wing with cell less 
than half the length of wing (about one-thid in Pyla). Antenna 
of male with sinus and scale tuft in base of shaft. Labial palpus 
oblique or upturned. Maxillary palpus various (minute, 
squamous or aigrettelike). Male genitalia with transtilla 
usually absent, if present (Phobus, Stylopalpia) incomplete or 
its median area very weakly sclerotized; harpe with sclerotized 
costa sometimes produced at base, never at apex; clasper absent 
or more or less developed (strongly so in many species of Pyla); 
aedeagus frequently divided (bifid) or spined; penis unarmed or 
finely scobinate or finely and weakly spined, rarely (Phobus) 
with a single cornutus. Female genitalia without signum; bursa 
frequently smooth or weakly spined, occasionally with some 
sclerotized folds continued from ductus bursae; the latter more 
or less sclerotized in part, in many Pyla species broadly expanded 
towards genital opening.] 


72. Telethusia, new genus 


Typr or Genus: Pempelia ovalis Packard. 

Tongue well developed. Antenna pubescent; on male 
with sinus and enlarged scale tuft in base of shaft. 
Labial palpus obliquely upturned, reaching to vertex 
on male, above vertex on female; laterally flattened and 
broadly scaled; second segment of male grooved on 
inner side to hold the tongue; third segment consider- 
ably shorter than second, bluntly pointed, more or less 
deflected forward and partially hidden in scaling of 
second segment. Maxillary palpus minute (a mere 
vestige). Forewing smooth; 11 veins; vem 2 from 
before lower outer angle of cell; 3 from the angle, 
nearer to 4 than to 2; 4 and 5 separated at base, parallel 
for a short distance beyond; 6 from below upper angle 
of cell, straight; 8 and 9 stalked for half or nearly half 
their lengths; 10 from the cell separated from the stalk 
of 8-9 at base and divergent from it shortly beyond; 
male without costal fold. Hind wing with vein 2 from 
before lower outer angle of cell; 3 from the angle, 
connate at base with the stalk of 4-5; 4 and 5 stalked 
for about half their lengths; 7 and 8 contiguous or 
weakly anastomosed for a short distance beyond cell; 
cell less than half the length of wing; discocellular vein 
curved, outwardly produced at lower angle of cell. 
Eighth abdominal segment of male with compound 
scale tufts. 

Male genitalia with uncus subtriangulate; apex 
bluntly rounded. Apical process of gnathos a short, 
stout hook. Transtilla absent. Harpe simple; clasper 
rudimentary. Aedeagus simple, straight, not taper- 
ing; penis unarmed except for a small comb of very 
weak, short, slender spines. Vinculum stout, longer 
than greatest width, tapering slightly to truncated 
terminal margin. 

Female genitalia with ovipositor strongly sclerotized; 
apophyses (supporting rods) of ovipositor and eighth 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 137 


segment collar stout; bursa elongate, finely scobinate 
over two-thirds of its inner surface but without signum 
or other sclerotization; ductus bursae unsclerotized 
except for a narrow, weak band near genital opening; 
ductus seminalis from junction of bursa and ductus 
bursae. At least half of membrane between collar and 
ovipositor finely and densely spinose. 

The genus is erected for a species hitherto referred to 
Nephopteryz. It differs from the latter in its vestigial 
maxillary palpi, the lack of cornutus or cornuti of penis 
or the hair brush on harpe of its male genitalia, the 
hardened ovipositor of female, the shorter stalking of 
veins 8 and 9, and the somewhat more separated condi- 
tion at base of veins 4 and 5 of forewing. 


278. Telethusia ovalis (Packard), new combination 
Ficures 353, 843 


Pempelia ovalis Packard, Ann. Lyc. Nat. Hist. New York, vol. 10, 
p. 269, 1873. 

Nephopteryz latifasciatella Packard, Ann. Lyc. Nat. Hist., New 
York, vol. 10, p. 269, 1873. 

Nephopteryz ovalis (Packard) Grote, Bull., U. 8. Geol. Geogr. 
Surv. Terr., vol. 4, p. 696, 1878; North Amer. Ent., vol. 1, 
p. 11, 1879.—Hulst, Phycitidae of N. Amer., p. 144, 1890.— 
Ragonot, Monograph, pt. 1, p. 269, 1893.—Forbes, Cornell 
Mem. 68, p. 623, 1923——McDunnough, Check list, No. 
6163, 1939. 

Nephopteryz ovalis geminipunctella Ragonot, Monograph, pt. 1, 
p. 270, 1893. 

Nephopteryx modestella Hulst, Canadian Ent., vol. 34, p. 170 
1900.—Barnes and McDunnough, Contributions, vol. 3, 
p. 196, 1916. 

Forewing ashy gray-white with a distinct powdery 
appearance, the white dusting conspicuous on median 
and basal areas; antemedial line oblique, zigzag (twice 
notched), narrow, white, bordered inwardly by a broad 
dark fuscous (blackish gray) band which is interrupted 
by a streak of dull ocherous orange at lower fold (this 
ocherous shade also continued along fold in median 
area) ; antemedial line bordered outwardly by a blackish 
gray bar at costa and similarly colored dots on cell and 
just above inner margin; subterminal line sinuate 
bordered inwardly by a more or less broken blackish 
gray line and outwardly by a broader blackish gray, 
brownish, or brownish ocherous shade (the latter when 
present interrupted by blackish streaklets on the veins), 
both borders strongly accented at costa; on most speci- 
mens a narrow, oblique, dark shading across median 
area from costal inner margin of subterminal line; discal 
dots distinct, separated, blackish, the lower sometimes 
expanded into a black smudge; a row of black dots along 
termen. Hind wing pale smoky fuscous, more whitish 
in some specimens, darker and somewhat brownish gray 
in others. Alar expanse, 20-28 mm. 

Genitalia as given for the genus. 

Type Locauities: Maine (ovalis and latifasciatella, 
in MCZ); Washington State (geminipunctella, in Paris 
Mus.); Massachusetts (modestella, in AMNH, ex Rut- 
gers). 

Foop puants: Antennaria, Eriophyllum ignotum. 
These records from Washington specimens reared by 
J. F. G. Clarke. Presumably on other Compositae. 


Distrisution: Unitep Srates: Maine, Orono (July), 
Wales (July); New Hampshire, Durham, Hampton 
(Aug.), Fort Washington (July); Vermont, Clarenton; 
Massachusetts; Connecticut, East River (July); New 
York, Catskill Mts., Ilion (June, July); Colorado, Gun- 
nison County (near Altmont, July); Utah, Stockton 
(June, July); Montana, Missoula (Aug.); Washington, 
Bellingham (June), Chuckanut Bay (Whatcom County, 
June), Godman Springs (Blue Mts., July), Kamiack 
Butte (May), Pullman (June, July); California, Placer 
County (June), San Jacinto Mts. (July), Tuolumne 
Meadows (July, Aug.). Canapa: Ontario, Trenton 
(July); Alberta, Banff (June, July); British Columbia, 
Wellington. 


The species is variable in color, especially in the 
Western areas of the United States and Canada. Most 
of the specimens from Washington and British Colum- 
bia have the white dusting on forewing more conspicu- 
ous and the dark markings more strongly contrasted 
than on eastern examples. However, there are inter- 
grades, and no sharp line can be drawn on color between 
the two areas. In the Tuolumne Meadows of Cali- 
fornia there is a larger (26-28 mm.) and somewhat paler 
form. Specimens from Colorado, Alberta, and occa- 
sionally from Washington form another darker and 
duller variety, the forewing showing little or no brown 
shading, the pattern markings a dull black, and the 
pale areas more gray than whitish and less strongly 
contrasted against the dark markings than in other 
Washington or eastern specimens. ‘Three specimens 
before me from California have the transverse dark 
lines much weaker and the over-all color an ashy gray 
with a slight bluish tint. The Utah examples are the 
most distinctive of all the forms, their hind wings de- 
cidedly paler, the forewing a very pale ashy gray and 
all the darker pattern markings more or less obscured 
or obliterated. I do not think that these varieties 
represent anything but color forms or that any one of 
them is entitled to a subspecific designation. More 
and wider collecting throughout the Middle and Far 
West will probably turn up still other color variants. 
The species itself, despite its variability, is easily identi- 
fied by its genitalia. 


279. Telethusia rhypodella (Hulst), new combination 


Glyptoteles rhypodella Hulst, Ent. Amer., vol. 3, p. 137, 1887. 

Nephopteryx rhypodella (Hulst), Phycitidae of N. Amer., p. 144, 
1890.—Ragonot, Monograph, p. 1, p. 270, 1893.—Barnes 
and McDunnough, Contributions, vol. 3, p. 196, 1916.— 
McDunnough, Check list, No. 6165, 1939. 

There are no specimens in the Rutgers Collection or 
elsewhere that I have seen matching Hulst’s description, 
nor any available Oregon examples that could be re- 
ferred to rhypodella. The alleged type at Rutgers, a 
female without locality label and bearing only the num- 
ber 42, is a typical representative of Phobus curvatella 
(Ragonot). Unfortunately the Hulst ‘“types’’ are fre- 
quently as unreliable as his type designations and this 
particular type is probably spurious. Hulst’s descrip- 
tions, on the other hand, are usually more reliable and I 


138 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


suspect that when sufficient Oregon material is available 
we shall find that rhypodella is merely one of the nu- 
merous color forms of ovalis. 

Typr Locauity: ‘Oregon’ (type lost?). 

Foop prant: Unknown. 


73. Phobus, new genus 


Type or GENus: Dioryctria brucei Hulst. 

Characters of Telethusia except: Eighth abdominal 
segment of male with a pair of ventrolateral hair tufts. 
Penis of male genitalia armed with a single cornutus; 
usually also a cluster of very fine, minute, slender spines 
at apex of aedeagus (but not on membranous penis). 
Transtilla represented at least by its lateral elements, 
sometimes the median area is recognizable but is very 
weakly sclerotized and the completed band not a con- 
stant character. Female genitalia with ovipositor nor- 
mal (not strongly sclerotized) ; apophyses of ovipositor 
and eighth-segment collar slender; bursa copulatrix 
simple (smooth); membrane between collar and ovi- 
positor smooth. 

The foregoing differences seem to justify separation 
from Telethusia, with which the genus is very closely 
related. 


280. Phobus brucei (Hulst), new combination 
Figures 354, 844 


Dioryctria brucei Hulst, Canadian Ent., vol. 27, p. 55, 1895. 

Ambesa lallatalis Hulst (not Hulst), U. S. Nat. Mus. Bull. 52, p. 
422, 1903. 

Tacoma lallatalis Dyar (not Hulst), Proc. Ent. Soc. Washington, 
vol. 6, p. 227, 1904. 

Nephopteryz lallatalis brucei (Hulst) McDunnough, Check list, 
No. 6160, 1939. 

Forewing whitish more or less dusted and shaded with 
blackish or fuscous scales, making the general color 
cream white (with some pale brownish shading on paler, 
weakly marked examples) to ashy gray with a faint 
bluish gray tint (on well-marked specimens) ; the trans- 
verse lines irregular and more or less interrupted and 
not strongly contrasted; antemedial line oblique, ser- 
rate, interrupted at lower fold by a pale olivaceous- 
ocherous shade which extends rather broadly the length 
of the fold and also cuts the subterminal line; a similar 
but somewhat weaker shade fills the cell; outer margin 
of antemedial line consisting of a thin blackish line 
curving outwardly from costa to top of cell, a small 
blackish dot or dash on lower vein of cell and a similar 
blackish marking on vein 1b; subterminal line markedly 
serrate, deeply indented (almost to cell) below costa, on 
well-marked (darker) specimens bordered inwardly by a 
black line from costa at least to cell, this line frequently 
continued along top of cell to the black outer marking 
of antemedial line, forming a continuous, long, narrow 
hook along the median and postmedian subcostal area; 
below the lower fold the blackish outer border of the 
subterminal line is also continued back, as a black line 
under vein 1b to and fusing with a narrow blackish line 
on the outer edge of the antemedial line forming a nar- 
row, oval marking on lower margin between the trans- 


verse lines; on dark examples a more or less conspicuous, 
blackish, quadrate patch inwardly bordering the ante- 
medial line at inner margin; on pale specimens this 
patch pale brown, more or less obscured; discal dots 
obscured, rarely distinguishable; a row of small narrow 
black or brownish dots along termen. Hind wings 
subpellucid with a faint ocherous tint; the veins not 
appreciably darkened; a faint narrow dark shade along 
termen. Alar expanse, 26-29 mm. 

Male genitalia with cornutus small, slender. Female 
genitalia, with bursa very short, not much longer than 
ductus bursae. 

Type tocaLiry: Colorado (type in AMNH, ex Rut- 
gers). 

Foop prant: Unknown. 

Distrisution: Colorado; Utah, Eureka (June, July), 
Stockton (June, July, Sept.); Nevada, Ormsby, and one 
female with only the state locality (a pseudotype of 
lallatalis Hulst); California, San Luis Obispo; Washing- 
ton, Pullman (July). 

All specimens of this species in the National Collec- 
tion had been identified as lallatalis Hulst on Dyar’s 
misidentification of the latter species. As a result 
Dyar referred brucei as a synonym of lallatalis. Hulst, 
however, was primarily to blame for the confusion; for 
he had identified and sent out as “types” of Jallatalis 
specimens of both brucei and Interjectio denticulella. 
The true brucei resembles superficially both lallatalis 
and denticulella in some of its more striking details of 
maculation, but is easily distinguished from both by 
its male and female genitalia. 


281. Phobus funerellus (Dyar), new combination 


Salebria funerella Dyar, Ins. Inse. Menstr., vol. 13, p. 12, 1925.— 
McDunnough, Check list, No. 6193, 1939. 

Forewing blackish to dark brownish gray more or less 
dusted with white on basal, median, and terminal areas; 
the white dusting very faint and scattered on the type 
series from Southern California, which have a uniform 
blackish gray ground color, much stronger on specimens 
from Washington, British Columbia, and New Mexico; 
transverse lines and thin blackish borders complete (not 
interrupted as in brucei), sinuate; the antemedial line 
oblique, sharply serrate, narrow, whitish gray, bordered 
outwardly by a narrow black line and inwardly by a 
broad, unbroken blackish or (on paler examples) dark 
gray-brown band, this band distinguishable and con- 
trasted even on the darkest, most suffused examples; 
subterminal line serrate but not deeply indented below 
costa, bordered inwardly by a narrow, continuous black 
line, the latter not continued inwardly below costa or 
on vein 1b as it is on brucei; subterminal line bordered 
outwardly by a rather broad dark band; discal dots 
usually distinct, black, more or less confluent; a row of 
blackish dots along termen. Hind wing brown; the 
veins darkened and a narrow blackish line along termen. 
Alar expanse, 24.5-30 mm. 

Genitalia similar to those of bruce. 

Typz Locauity: Southern California (type in USNM). 

Foop puant: Unknown. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 139 


Distrisution: Unirep States; California (Southern 
California without more definite locality, the type 
series), Clarksville (El Dorado County, June); Washing- 
ton, Pullman; New Mexico, Fort Wingate (July). 
Canava: British Columbia, Departure Bay (Aug.), 
Duncans (Vancouver Isl., July), Nicola (July), Welling- 
ton (June). Also one male without state locality 
labeled ‘‘Larima Co. [sic], Aug., 1901, Schaus collector.”’ 

The species is distinct and easily distinguished from 
brucei on the color and pattern of forewing, especially 
by the broad black band extending from inner margin 
to costa before the antemedial line and by the shallow 
indentation of the subterminal line below costa. 


282. Phobus curvatellus (Ragonot), new combination 
Ficures 355, 845 


Nephopteryx curvatella Ragonot, N. Amer. Phycitidae, p. 7, 
1887.—Barnes and McDunnough, Contributions, vol. 3, 
p. 196, 1916.—McDunnough, Check list, No. 6166, 1939. 

Nephopteryx rhypodella Ragonot (not Hulst), Monograph, pt. 1, 
p. 270, 1893. 

Forewing ashy bluish gray; the transverse lines com- 
plete, narrow, white, obscure except on the well-marked 
darker examples, indicated chiefly by the fine, black 
outer border of the antemedial line and similar black 
inner border of the subterminal] line; preceding the ante- 
medial line a quadrate blackish spot on inner margin; 
limited above by a weak, smaller, pale, somewhat oilva- 
ceous shade in the lower fold, this pale shade not 
interrupting the antemedial line itself; indentations of 
subterminal line as in funerellus; discal dots obscure, 
the lower one sometimes distinct (under magnification) 
and frequently with a dark shade below it which forms 
a round dark spot, to the naked eye one of the more 
conspicuous markings on the wing; a row of more or 
less confluent black dots along termen. Hind wing 
translucent, whitish with a smoky shade towards apex; 
the veins darkened; a fine brown line along termen. 
Alar expanse, 26-30 mm. 


Male genitalia with cornutus of the same length as 
that of brucei and funerellus but somewhat stouter. 
Genitalia otherwise like those of the species following 
(incertus). Female genitalia with bursa very long, and 
narrow throughout its length, but little wider than the 
ductus bursae. 


TypE Locatity: America Septentrionalis (type im 
Paris Mus.). 

Foop puant: Unknown. 

Distrisution: California, Loma Linda (June, July), 
Los Angeles County (1,060 ft., June), Monachee 
Meadows (Tulare County, July), Mount Lowe (July), 
San Gabriel Mts. (1,700 ft., July); Arizona, Nogales 
(May), Santa Rita Mts. (May); Utah, Bellevue 
(Washington County, May, June); Colorado, Silverton 
(July). 

The spurious type of Telethusia rhypodella (Hulst) 
in the Rutgers Collection belongs here. It is discussed 
under the treatment of rhypodella (p. 137). 


283. Phobus incertus, new species 
Ficures 356, 846 


Color and markings of forewing as in curvatellus 
except duller, lacking the bluish tint of the latter 
species; the quadrate dark spot preceding the ante- 
medial line also continued as a broad band to costa, 
though frequently interrupted by a pale shading at 
lower fold. 

Male genitalia with cornutus appreciably stouter and 
longer than that of any of the other species of the genus. 
Bursa copulatrix of the female genitalia less than half 
the length of that of curvatellus but twice the length of 
that of brucei or funerellus. 


Type Locauity: Strawberry Valley (6,000 ft.), San 
Jacinto Mts., Calif. (type in USNM, 61348). 

Described from male type and five male and five 
female paratypes from the type locality, collected by F. 
Grinnell, Jr., June 16, 17, and 18, 1908. 

Except for the genitalic differences this might easily 
be a higher altitude race of curvatellus, but the differ- 
ences in size of cornutus and length of bursa seem to 
be constant characters and greater than to be expected 
in variants of one species. 


74, Actrix, new genus 


TypzE or Genus: Tacoma nyssaecolella Dyar. 


Tongue well developed. Antenna weakly pubescent; 
on male with sinus and scale tuft in base of shaft. 
Labial palpus upcurved, slender, reaching above vertex; 
second segment somewhat flattened and very slightly 
rough scaled, not grooved on male; third segment about 
two-thirds the length of second, acuminate. Maxillary 
palpus squamous. Forewing smooth; 11 veins; vein 2 
from near lower outer angle of cell; 3 from the angle, 
approximately equidistant from 2 and 4 at base; 4 and 
5 shortly stalked; 6 from below upper angle of cell, 
straight; 8 and 9 very long stalked; 10 from the cell, 
approximate to the stalk of 8-9 for some distance; male 
without costal fold. Hind wing with vein 2 from before 
but near lower outer angle of cell; 3 from the angle, 
connate or (on occasional specimens) shortly fused 
with the stalk of 4-5; 4 and 5 stalked for two-thirds of 
their lengths; 7 and 8 strongly anastomosed beyond 
cell for half or a trifle more than half their lengths; 
cell slightly less than half the length of wing; dis- 
cocellular vein curved, extended outwardly at lower 
angle of cell. Eighth abdominal segment of male with 
pair of ventrolateral hair tufts. 


Male genitalia with uncus broader than long, its 
outer lateral angles slightly lobed. Apical process of 
gnathos broad, shieldlike, its lateral arms greatly 
reduced. ‘Transtilla absent. Harpe without clasper. 
Aedeagus straight, divided towards apex, the projecting 
divided elements strongly sclerotized; penis with some 
very weak scobinations, but otherwise unarmed. 
Vinculum short, stout, about as long as greatest width, 
evenly rounded to blunt terminal margin. 


140 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Female genitalia with bursa small, covered with 
dense slender spines on posterior half, otherwise very 
finely spinose, without signum; ductus bursae granu- 
late, short, expanded into a broad, strongly sclerotized, 
contorted plate towards genital opening; ductus semi- 
nalis from lobe of bursa near junction of bursa and 
ductus bursae. 

The genus is a further restriction from Nephopteryz 
of authors. Its nearest relationship seems to be Pyla, 
which it resembles in the peculiar modification of the 
ductus bursae of the female and the aedeagus of the 
male. Its venation, however, is quite distinct and, 
except for the strong anastomosis of veins 7 and 8 of 
hind wing, closer to that of Tacoma. 


284. Actrix nyssaecolella (Dyar), new combination 


Ficures 357, 838 
Tacoma nyssaecolella Dyar, Proc. Ent. Soc. Washington vol. 6, 
p. 112, 1904.—Forbes, Cornell Mem. 68, p. 621, 1923.— 
Craighead, U. S. Dep. Agr. Misc. Publ. 657, p. 453, 1950. 
Nephopteryx nyssaecolella (Dyar) Barnes and McDunnough, 
Contributions, vol. 3, p. 196, 1916—McDunnough, Check 
list, No. 6174, 1939. 

Forewing brownish gray (“lilaceous gray”) paler in 
basal area beyond extreme base, in the half of median 
area just beyond antemedial line, and to a lesser extent 
in outer area (beyond the subterminal line), these pale 
areas of an ashy hue, due to a faint peppering of white 
scales; antemedial line narrow, oblique and very slightly 
angled at middle, dull white, preceded on inner margin 
by a subquadrate blackish brown patch and bordered 
outwardly at costa by a triangulate black patch which 
(on most specimens) continues as a more or less inter- 
rupted black line to inner margin; subterminal line 
sinuate (bulged at middle) and weakly serrate, dull 
white, bordered inwardly by a dark shade and outwardly 
by a narrower dark line, these dark borders especially 
marked and blackish at costa; discal dots confluent, 
blackish; a row of more or less confluent black dots 
along termen. Hind wing pale smoky brown, with a 
glossy sheen; veins very faintly darkened; a narrow 
brown line along termen. Alar expanse, 15-18 mm. 

Male genitalia with terminal margin of uncus con- 
cave, its lateral lobes turned laterally outward. Apical 
process of gnathos a convex shield, slightly longer than 
broad and with apical, lateral angles produced backward 
into bluntly pointed spines. Divided elements of aedea- 
gus produced as short, sharply and oppositely curved 
hooks. Cucullus of harpe sharply curved towards its 
apex. Female genitalia distinguished at once by the 
shape of the ventral sclerotized plate of ductus bursae 
and the eighth-segment collar, which is complete and 
strongly sclerotized ventrally. 

Typr Locatity: Washington, D.C. (typein USNM). 

Foop prant: Nyssa sylvatica (larva a leaf-folder). 

Distrisution: Massachusetts, Nantucket (July); 
Connecticut, Kast River (July, Aug.); Rhode Island, 
Weekapaugh (July) ; New Jersey, Anglesea (May, June); 
Pennsylvania, New Brighton (July, Aug.), Oak Station 
(Aug.) ; District of Columbia, Washington (Aug.); North 
Carolina, Southern Pines (Aug.), Tryon (Aug.). 


285. Actrix dissimulatrix, new species 
Fieures 358, 837 


Superficially not distinguishable from nyssaecolella 
except (on the specimens before me) a slightly stronger 
white dusting on median area of forewing especially 
over inner margin immediately following the antemedial 
line. This difference can hardly be expected to hold 
for the species. 

The genitalia of both sexes are very different from 
those of nyssaecolella. Male genitalia with terminal 
margin of uncus evenly rounded, its lateral lobes turned 
inward and downward, partially encircling the anal 
tube. Apical process of gnathos heart shaped, without 
produced angles. Divided elements of aedeagus pro- 
duced as long, straight, stout thornlike projections, one 
of which is coarsely scobinate. Female genitalia with 
a stout pair of strongly sclerotized, curved, lateral arms 
projecting forward from the ventral sclerotized plate 
of the ductus bursae. 


Typ Locatity: Cape Henry, Va. (type in USNM, 
61349). 

Foop prant: Nyssa sylvatica. 

Described from male type and two male and one 
female paratypes from the type locality, reared by A. 
Busck, Aug. 10, 16 and 18, 1927, from larvae feeding on 
the leaves of Nyssa sylvatica. Four larvae were pre- 
served from the collection. Three of these are blackish 
brown in color and are undoubtedly nyssaecolella. One 
larva is yellow with a pale yellowish head and thoracic 
shield. Itis probably dissimulatriz. No difference was 
noted in larval habits between the two forms. 


75. Genus Stylopalpia Hampson 


Stylopalpia Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7, 
p. 257, 1901. (Lype of genus: Stylopalpia lunigerella 
Hampson.) 

Tongue well developed. Antenna very shortly pubes- 
cent, shaft of male somewhat flattened and with a 
shallow sinus at base containing a row of short spines 
more or less concealed by a small, weak scale tuft. 
Labial palpus obliquely upturned; third sesment very 
long (nearly twice the length of second), slender and 
porrect in lunigerella, much shorter (about half the 
length of second) and oblique in the other two species, 
in these reaching a little above vertex, acuminate in all 
species. Maxillary palpus minute, its scaling slightly 
expanded. Forewing smooth; 11 veins; vein 2 from 
before the lower outer angle of cell; 3 from the angle; 
4 and 5 separated at base and divergent beyond, 4 but 
slightly nearer to 5 at base than to 3; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for slightly more 
than half their lengths; 10 from the cell, well separated 
from the stalk of 8-9 at base but just beyond approach- 
ing it for a very short distance; male without costal 
fold. Hind wing with vein 2 from well before lower 
outer angle of cell; 3 from the angle; 4 and 5 stalked 
for about half their lengths; 7 and 8 approximate or 
contiguous for a very short distance beyond cell; cell 
less than half the length of wing; discocellular vein 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 141 


curved, produced at lower angle, a short spur connecting 
it and vein 8. Highth abdominal segment with weak 
compound tufts in lwnigerella, paired ventral tufts in 
the other two species. 

Male genitalia with uncus subtriangulate, its terminal 
margin more or less broadly rounded. Apical process 
of gnathos a simple, rather short, stout hook. Trans- 
tilla incomplete, sclerotized only in its reduced lateral 
elements (the central portion, as shown in the figures, 
distinguishable but not sclerotized). Harpe with large, 
strongly sclerotized, erect, scoop-shaped clasper, situ- 
ated towards base, below costa; cucullus narrow, 
elongate, very slightly tapering to rounded apex. 
Anellus shield-shaped, with small lateral lobes. Aedea- 
gus expanding to lateral, flanged projections before 
apex, the latter flanges, each bearing a cluster of strong 
spines; penis unarmed. Vinculum stout; slightly longer 
than broad; tapering slightly to broadly rounded ter- 
minal margin. 

Female genitalia without signum. Bursa mem- 
branous with only a little fine spining at junction of 
bursa and ductus bursae; ductus bursae weakly scle- 
rotized at, and just before, genital opening, the latter 
broad; ductus seminalis from middle of ductus bursae. 

The genus was originally erected on the peculiar 
palps of its type species. This character, however, 
proves to be specific rather than generic. The two 
species here included do not have it; but agree with 
the type species on every other detail of venation, 
antennal structure, and genitalia. The genus can 
easily be maintained on its combination of male and 
female genitalic characters. 


286. Stylopalpia lunigerella Hampson 


Ficures 30, 359, 848 
Stylopalpia lunigerella Hampson, Ann. Mag. Nat. Hist., ser. 7, 
vol. 7, p. 258, 1901. 
Third segment of labial palpus very long, slender 
and porrect. Eighth abdominal segment of male with 
compound tufts. 


Forewing ocherous (clay color) dusted with blackish, 
making the general shade dark gray, the ocherous color 
forming a contrasting band along costa and more or 
less lightening the lower median area and the base of 
inner margin; antemedial line indicated by a pale 
lunulate line between cell and inner margin and, on 
its upper half, by faint traces of its narrow, blackish, 
outer border; subterminal line very close to outer 
margin, slightly bulged at middle, not serrate, pre- 
ceded by some black streaklets on the veins and whitish 
or pale ocherous streaklets between them; discal dots 
separated, blackish; a few of black dots along termen. 
On female a somewhat broader brownish shade at apex 
and along termen and some darkening of the outer 
parts of the veins. Alar expanse, 18-24 mm. 

Male genitalia with clasper of harpe considerably 
longer than deep, serrate along lower and inner margins. 
Female genitalia with genital opening very broad. 

Type Locauity: Jamaica (type in BM). 


Foop pirant: Unknown. 

DistriputTion: Banamas, Nassau. Purrto Rico: 
Aguirre Central (Apr., June, Aug.), Camuy (Apr.), 
Coamo Springs (Apr.), Ponce (Sept.), San Germén 
(June). Cua: Santiago Province (June, July, Oct.). 
Jamaica. M*&xtco: Colima (Jan., June, July). 

The species is easily identified by its peculiar palpi, 
alike in both sexes. 


287. Stylopalpia scobiella (Grote), new combination 
Ficure 360, 847 


Nephopteryx scobiella Grote, N. Amer. Ent., vol. 1, p. 5, 1890.— 
Hulst, Phycitidae of N. Amer., p. 148, 1890.—Ragonot, Ent. 
Amer., vol. 5, p. 115, 1889; Monograph, pt. 1, p. 266, 1893.— 
McDunnough, Check list, No. 6161, 1939. 

Lipographis decimerella Hulst, Ent. Amer., vol. 4, p. 117, 1888. 

Third segment of labial palpus short, oblique. Except 
on a few of the grayer specimens, head and thorax 
distinctly ocherous. 

Forewing pale gray to grayish ocherous, extreme base 
of wing ocherous and a similar pale ocherous shade 
rather broadly bordering the costa; transverse markings 
nearly obliterated; antemedial line indicated only by a 
small Junate white spot on vein 1b, preceded and fol- 
lowed by black dots, a similar blackish dot or streaklet 
at lower margin of cell (representing a median fragment 
of the usual black outer border of the antemedial line) ; 
subterminal line obscure, a very faint pale line weakly 
bordered inwardly by a darker shade; lower discal spot 
a blackish streaklet, upper discal dot usually absent, if 
present very faint; a row of fine blackish dots along 
termen. Hind wings whitish with a faint ocherous or 
smoky tint; the veins little if any darkened; a thin 
brownish line along termen. Alar expanse, 24-26 mm. 

Male genitalia with clasper of harpe broadly oval, 
not serrate. Terminal margins of uncus and vinculum 
very broadly rounded, the vinculum not appreciably 
tapering. Eighth abdominal segment of male with 
paired ventral tufts. Female with genital opening more 
constricted than that of lunigerella. 

Typ LocaLitiEs: Bosque County, Tex. (scobiella, in 
BM); Blanco County, Tex. (decimerella, in AMNH, 
ex Rutgers). 

Foop pirant: Unknown. 

Distrisution: Texas, Barber (Apr.), Beeville (May), 
Blanco County (May), Bosque County, Burnet County 
(Apr., Sept., Oct.), Kerrville, Sabinal (Mar., Apr.), 
Sapulpa (May), San Diego (Apr.), Victoria (Apr., Sept.), 
Zavalla County (Apr.); Colorado, Glenwood Springs. 
Also four specimens from Texas with only the state 
locality. The species probably also occurs in northern 
México. 


288. Stylopalpia argentinensis, new species 
Ficure 361 


Labial palpus and eighth abdominal tufts as in scobi- 
ella. 'The head and thorax brown. 

Forewing pale brown with a strong dusting of white 
scales faintly peppered with black in median area; basal 


142 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


area brownish ocherous, shading to brown at extreme 
base and with a clouding of white in midbasal area; ante- 
medial line complete, well out on wing, oblique, inwardly 
notched at vein 1b, white, bordered outwardly by a 
narrow blackish line; subterminal line also complete, 
further back from termen than in the two preceding 
species, oblique (parallel to termen), whitish ocherous 
with a narrow blackish inner border; discal dots distinct, 
separated, black. Hind wing dark brown; the fringe 
whitish ocherous with a dark median band. Alar ex- 
panse, 24 mm. 

Male genitalia similar to those of scobiella except: 
Uncus narrower and tapering to more narrowly rounded 
terminal margin; vinculum longer in proportion to its 
width; spines on aedeagus fewer and coarser. 

TyrEeocauity: “Villa Anna, F.S.C. Fe., Argentina”’ 
(type in BM). 

Foop prant: Unknown. 

Described from unique male, collected by K. J. 
Hayward, Dec. 1925. 


76. Genus Pyla Grote 


Pyla Grote, New check list of North American moths, p. 55, 
1882.—Ragonot, N. Amer. Phycitidae, p. 9, 1887; Mono- 
graph, pt. 1, p. 481, 1893.—Hulst, Phycitidae of N. Amer., 
p. 161,1890. (Typeof genus: Nephopterysx scintillans Grote.) 

Tongue well developed. Antenna finely pubescent; 
on male with sinus and scale tuft in base of shaft. 
Labial palpus oblique; second segment broadly scaled, 
somewhat flattened laterally, reaching above vertex; on 
male with a slight groove to hold the maxillary palpus; 
third segment short, less than one-third the length of 
second, porrect. Maxillary palpus of male in the form 
of an aigrette, semiaigrette (i. e., the scales hairlike 
but short), or more or less squamous. Forewing smooth; 
11 veins; vein 2 from near lower outer angle of cell; 3 
from the angle, slightly nearer to 4 at base than to 2; 
4 and 5 slightly separated at base, parallel for a short 
distance beyond cell; 6 from below upper angle of cell 
straight; 8 and 9 stalked from one-half to a third of 
their lengths; 10 from the cell, slightly separated at 
base from the stalk of 8-9; male without costal fold. 
Hind wing with vein 2 from before but rather near 
lower outer angle of cell; 3 from the angle, connate with 
4; 4 and 5 stalked or anastomosed for half or slightly 
less than half of their lengths; 7 and 8 contiguous or 
very weakly anastomosed for a short distance beyond 
cell; cell less than half the length of wing; discocellular 
vem curved, outwardly produced at lower angle and 
connected with vein 3 by ashort spur. Eighth abdom- 
inal segment of male with a pair of ventrolateral hair 
tufts (fig. 372b), or two or three pairs containing some 
modified scales (fig. 367c). 

Male genitalia with uncus broad, more or less trian- 
gulate. Apical process of gnathos a short, stout hook. 
Transtilla absent. Harpe usually with base of costa 
produced into a strong projecting hook, or spine, or 
spined lobe; frequently a strong hooked or spined clasper 
from median basal area; sacculus simple; costa strongly 
sclerotized but sclerotization rather abruptly terminated 


before apex of harpe. Aedeagus usually partially divided 
(bifurcate) or armed with projecting spine or spines, 
rarely simple; penis unarmed (except for a very weak 
cornutus in fusca). Vinculum stout, slightly tapered to 
truncate or more or less broadly rounded terminal 
margin. 

Female genitalia without signum; bursa copulatrix 
usually simple, sometimes with strongly sclerotized, 
convolute, longitudinal bands near junction of bursa 
and ductus bursae and extending a short distance into 
the ductus; ductus bursae short, widening to broad gen- 
ital opening, usually strongly and elaborately sclero- 
tized towards genital opening; genital opening rarely 
simple (fasciolalis, mridisuffusella); ductus seminalis 
from bursa near junction of bursa and ductus bursae. 

The genus as here defined includes what superficially 
appears to be two distinct entities, one group of species 
with gray forewings and another with shiny brown 
wings, the latter the typical Pyla of authors. For con- 
venience of identification I am designating them as 
species groups. The division is not supported by any 
consistent structural character or combination of char- 
acters. The differences exhibited by the several species 
in male and female genitalia, male maxillary palpi, and 
male abdominal tufts are striking, but apparently only 
of specific significance. 

Nothing is known of the food habits or early stages of 
any of the species except fusca, which is recorded from 
Erica in the Old World. I suspect that the Hricaceae 
will prove to be the chief hosts of the genus. 


Genus Pyla, Species 289-297: P. fasciolalis to 
P. hanhamella 


[Ground color of forewing gray.] 


289. Pyla fasciolalis (Hulst), new combination 
Figures 362, 3638, 849 


Pinipestis fasciolalis Hulst, Trans. Amer. Ent. Soce., vol. 13, p. 
162, 1886. 

Nephopteryx fasciolalis (Hulst) Ragonot, Ent. Amer., vol. 5, p. 
115, 1889; Monograph, pt. 1, p. 271, 1893 (?).—Hulst, 
Phycitidae of N. Amer., p. 144, 1890.—McDunnough, Check 
list, No. 6167, 1890. 

Maxillary palpus of male squamous. 

Forewing gray finely powdered with white, giving the 
wing an ashy gray appearance; antemedial line distinct 
throughout, whitish, oblique, notched at top of cell and 
on lower fold, bordered outwardly by a black line begin- 
ning as a black smudge on costa, inwardly by a mod- 
erately broad black line extending from inner margin to 
cell; subterminal line well marked, bulged at middle and 
more or less dentate, bordered inwardly by a blackish 
band and outwardly by a somewhat broader, fainter 
dark band, these dark borders strongest near costa; 
discal dots distinct, small, normally separated, occa- 
sionally partially coalesced; a row of small black dots 
along termen. Hind wing smoky white with a pale 
brownish tint; veins very faintly darkened ; a narrow dark 
shade along termen. Alar expanse, 27-30 mm. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 143 


Male genitalia with harpe simple; clasper vestigial. 
Aedeagus deeply divided; one of the divided elements 
slightly forked at apex. Female genitalia with strongly 
sclerotized, convulute bands in bursa; genital opening 
simple. 

TYPE Locauity: British Columbia (type in AMNH, 
ex Rutgers). 

Foop prant: Unknown. 

Distrizution: British Columbia, Goldstream (Aug.), 
Necola (July), Saunichton (July). 

The only specimen in the Rutgers Collection is a large 
male (30 mm.) from British Columbia labeled “Dioryc- 
tria fasciolalis Hulst, Type.” I think we may safely 
assume this to be the actual type although in his original 
description Hulst gives ‘‘Nevada”’ as the type locality, 
probably one of his characteristic lapses. A perfect 
match for the type in color, markings, and genitalia, is 
found in a specimen from Necola, in the Canadian Na- 
tional Collection. The other records cited above (Gold- 
stream and Saanichton) are from specimens in the U. 8. 
National Collection. 

Also before me are four examples of what I take to be 
a variety of fasciolalis—two males from Gunnison 
County, Colo. (July), and 2 females from Wallace, 
Idaho (Aug.). Their blackish markings on forewing 
are a trifle stronger, the vinculum of male genitalia (fig. 
363) is somewhat shorter than in typical fasciolalis. The 
convolute bands in the bursa of the female are also a 
trifle longer. I doubt very much if these differences 
ndicate anything more than a possible local race. 


290. Pyla impostor, new species 
Fieures 364, 850 


Maxillary palpus of male in the form of an aigrette. 

Forewing color and markings similar to those of 
fasciolalis except: Somewhat duller and darker; white 
dusting sparser; the transverse pale lines fainter, in 
some specimens much obscured; their blackish borders 
less strongly contrasted against the ground color of the 
wing. Hind wing pale smoky fuscous, the brownish 
tint of fasciolalis very faint or altogether lacking. Alar 
expanse, 23-30 mm. 

Male genitalia with a long, slender, somewhat flat- 
tened, outwardly curved clasper on harpe; base of costa 
not modified. Aedeagus divided for about half its 
length, moderately slender, the divided elements rigid, 
pointed and unforked at their apices. Female with 
convolute, sclerotized folds extending from bursa shortly 
into ductus bursae; ductus bursae itself partially sclero- 
tized, the sclerotization forming broad ventrolateral 
bands extending from just beyond the convolute folds 
of the bursa to genital opening. 

TYPE Loca.ity: Slate Peak, Whatcom County, Wash. 
(6,000-7,000 ft.; type in USNM, 61350; paratypes in 
USNM and Canadian Nat. Coll.). 

Foop prant: Unknown. 

Described from male type and one female paratype 
from the type locality, collected by J. F. G. Clarke, Aug. 


2, 1940; and paratypes from the following localities: 
Bogachiel Peak, Olympic Mts., Wash., Aug. 9, 1936, 
Dr. A. F. Braun (?); Chimney Gulch, Golden, Colo., 
Oslar (oc); Colorado with only the state locality (a 
pseudotype, o, of fascialis Hulst, from the Fernald 
Collection); Big Belt Mts., Mont., July 18, 1928, J. 
McDunnough (co); upper Gallatin Canyon, Mont., 
7,000 ft., July 4, 1928, J. McDunnough (c’); Banff, 
Alberta, July 20, 1925, Owen Bryant, (co); Lethbridge, 
Alberta, July 3, 1922, H. L. Seamans (c’); Moraine 
Lake, Alberta, July 3, 4, 7, 1923, J. McDunnough (4 &* 
and 2 °); Waterton Lakes, Alberta, July 23, 28, 1923, 
J. McDunnough (co and 2); Hope Mts., British Co- 
lumbia, July 22, 1932, A. N. Gartrell (o&’); Mount Rey- 
elstoke, British Columbia, 6,000 ft., July 12, 1923, E. 
E. Buckell (0). 

Most of the foregoing were in our collections as 
fasciolalis on the basis of the false Hulst type in the 
National Museum. It is superficially like fasciolalis, 
but a distinct mountain-top species easily identified by 
its genitalia. 


291. Pyla aequivoca, new species 
Fiaures 366, 855 


Maxillary palpus of male in the form of an aigrette. 

Superficially like ampostor; the ground color of the 
male forewing a trifle more brownish gray, and the dark 
outer border of antemedial line somewhat broader and 
diffused into the ground color at costa, differences that 
could not be expected to hold in any extended series. 
Distinguished by its genitalia. Alar expanse, 26-29 mm. 

Male genitalia with clasper arising from midbasal 
area of harpe as in impostor, but much shorter and angu- 
late. Anellus U-shaped, its lateral arms long and 
slender. Aedeagus much shorter and somewhat stouter; 
its divided elements broader, decidedly flattened, termi- 
nating in sharp spines and more or less laterally spined 
towards apices. Eighth abdominal segment of male 
with two pairs of hair tufts. 

Typr Ltocauity: Banff, Alberta, Canada (type in 
Canadian Nat. Coll., paratype in USNM, 61351). 

Foop puant: Unknown. 

Described from male type and three male paratypes 
from the type locality, collected by C. B. D. Garrett, 
June 21 and 30 and July 6, 1922; and one male paratype 
from Hymers, Ontario June 8, 1915. In addition to the 
type series I have before me a female from the Canadian 
National Collection collected at Aweme, Manitoba, 
Aug. 26, 1921, by Norman Criddle. Its genitalia differ 
markedly from those of impostor. The membrane of 
bursa is thickened near junction with ductus bursae but 
lacks the sclerotizations of impostor; the ventrolateral 
bands of the ductus bursae are differently shaped and 
fuse into the ventrolateral sclerotizations of the inter- 
segmental area before the eighth-segment collar. While 
I have little doubt that this female is conspecific with 
the males, I am not designating it as a paratype. 


144 


292. Pyla insinuatrix, new species 


Fiaures 365, 856 


Maxillary palpus of male in the form of an aigrette. 

Forewing paler than in the preceding species, due to 
stronger white dusting, giving the paler areas a faint 
bluish tint; transverse lines and dark markings more 
strongly contrasted; the white antemedial line especially 
well marked on its lower half, its inner black border 
below cell expanded into a moderately wider blackish 
band or patch; blackish inner and outer borders of sub- 
terminal line well marked, especially the former; discal 
dots conspicuous, tending to coalesce. Hind wing 
subpellucid smoky white, darkening towards apex and 
termen; the veins very faintly darkened and a narrow 
dark line along termen. Alar expanse, 24-26 mm. 

Male genitalia with uncus hoodlike, constricted to- 
wards base. Harpe with a strong, outwardly produced, 
spined, knob from base of costa; no appreciable clasper. 
Aedeagus slender, with a very slight bifurcation at 
apex; the bifurcate projections straight. ‘Two pairs of 
ventrolateral tufts on eighth abdominal segment; some 
of the hairs broadly expanded at their apices (as in 
aenigmatica, fig. 367¢c). 

Female genitalia with bursa copulatrix membranous 
except for a very faint sclerotization of the lobe giving 
off the ductus seminalis; ductus bursae flattened, 
weakly sclerotized, expanding at genital opening into 
sclerotized, scobinate ventrolateral lobes. 

Type Locatiry: Aweme, Manitoba, Canada (type in 
Canadian Nat. Coll.; paratypes in USNM, 61352). 

Foop pranr: Unknown. 

Described from male type and four male and two 
female paratypes from the type locality, collected by 
Norman Criddle July 13 and Aug. 10, 1925; Aug. 10, 
1921; Aug. 10, 1925; Aug. 19, 1915). 


293. Pyla aenigmatica, new species 
Figures 367, 853 


Mazxillary palpus of male in the form of a semiaigrette 
(the hairs short). 

Forewing as on insinuatriz except darker, the ground 
color like that of impostor; the transverse lines distinct; 
lower half of antemedial line bordered inwardly by a 
subquadrate blackish patch, costal half of the outer 
dark border rather broad and well contrasted, blackish; 
the dark borders of subterminal line well contrasted 
towards costa; discal dots confluent; dots along termen 
minute, weak. Alar expanse, 25-28 mm. 

Male genitalia with tegumen having two, strongly 
spined, protruding lobes on each ventrolateral margin. 
Harpe with base of costa enlarged and coarsely scobi- 
nate; clasper small, semicircular, erect. Aedeagus 
slender; shortly bifurcate; bifurcate elements at apex 
spinelike, bent sharply at right angles to the aedeagus. 
Anellus a semitubular shield with rather long, strongly 
sclerotized lateral arms. Eighth abdominal segment 
with two pairs of ventrolateral hair tufts; one pair with 
the hairs expanded at their apices (forming knoblike 
clusters). 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Female genitalia with bursa membranous. Ductus 
bursae short, strongly sclerotized along lateral margins, 
the sclerotizations expanding laterally and at right 
angles at genital opening into a pair of convolute, finely 
scobinate lobes. 

Typrz LocaLity: Wellington, British Columbia (type 
in USNM, 61353; paratype in Canadian Nat. Coll.). 

Foop prant: Unknown. 

Described from male type and one male paratype 
from the type locality, “‘21-VI-04,” G. W. Taylor, and 
paratypes from the following localities: Goldstream, 
British Columbia, “30-VIII-20” (9); Salmon Arm, 
British Columbia, ‘‘22-6-20, W. R. B.” (co); Pine 
Grove, Colo., July 8, 1901, H. G. Dyar, “17310” (&); 
East River, Conn., Aug. 21 and Sept. 3, 1908, C. R. 
Ely (f and 9); Oak Station, Pa., Aug. 20, 1911, Fred 
Marloff (#); Watchung Mts., N. J., “64-99,” W. D. 
Kearfott (). 

The hind wings are a trifle darker on the eastern 
examples, which were, in our collection, identified as 
Susca. 

The species is evidently closely related to insinuatriz, 
but is distinct and easily distinguished by its genitalia. 


294. Pyla criddlella Dyar 
FieurEe 368 


Pyla criddlella Dyar, Journ. New York Ent. Soc., vol. 15, p. 110, 
1907.—McDunnough, Check list, No. 6241. 


Maxillary palpus of male squamous. 

Forewing blackish gray, semilustrous, unicolorous 
except for a slight darkening of the ground color border- 
ing the transverse lines; the latter very faint, but slightly 
lighter than the ground color; discal and terminal dots 
obscured. Hind wing brownish gray. Alar expanse, 
18 mm. 

Male genitalia with harpe simple except for a greatly 
reduced, upcurving, triangulate clasper. Aedeagus 
deeply bifurcate; the right divided element (in ventral 
view) produced into a sharp abruptly curved hook. A 
single pair of ventrolateral hair tufts on eighth abdom- 
inal segment. 


Typ Locatity: Aweme, Manitoba, Canada (June; 
type in USNM). 

Foop ruant: Unknown. 

Known only from the male type. 


295. Pyla fusca (Haworth), new combination 
Ficures 369, 852 


Phycis fusca Haworth, Lepidoptera Brittanica, pt. 3, p. 493, 1828. 

Phycita fusca (Haworth) Stephens, Illustrations of British ento- 
mology, Haustellata, vol. 4, p. 310, 1834. 

Pempelia fusca (Haworth) Stainton, Manual of British butterflies 
and moths, vol. 2, p. 176, 1859.—Packard, Ann. Lye. Nat. 
Hist. New York, vol. 10, p. 271, 1873. 

Nephopteryx moestella Walker, List, vol. 27, p. 53, 1863. 

Eudorea (?) frigidella Packard, Proc. Boston Soc. Nat. Hist., 
vol. 11, p. 53, 1866. 

Salebria fusca (Haworth) Heinemann, Die Schmetterlinge 
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 156, 
1865.—Grote, Bull. U. S. Geol. Geogr. Surv. Terr., vol. 4, 
p. 695, 1878; North Amer. Ent., vol. 1, p. 11, 1879.— 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 145 


Staudinger and Rebel, Catalog der Lepidopteren des palae- 
arctischen Faunengebietes, vol. 2, p. 34, 1901.—Spuler, Die 
Schmetterlinge Europas, vol. 2, p. 211, 1910.—Meyrick, 
Revised handbook of British Lepidoptera, p. 380, 1928.— 
Ford, Guide to the smaller British Lepidoptera, p. 10, 1949. 

Pinipestis cacabella Hulst, Ent. Amer., vol. 3, p. 133, 1887. 

Laodamia fusca (Haworth) Ragonot, Ent. Amer., vol. 5, p. 115, 
1889; Monograph, pt. 1, p. 408, 1893.—Hulst, Phycitidae 
of N. Amer., p. 156, 1890; U. 8. Nat. Mus. Bull. 52, p. 425, 
1902.—Forbes, Cornell Mem. 68, p. 628, 1923.—MceDun- 
nough, Check list, No. 6227, 1939. 

Salebria triplagiatella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 109, 1904.—Barnes and McDunnough, Contributions, 
vol. 3, p. 196, 1916. 

Dioryctria fusca (Haworth) Pierce and Metcalfe, The genitalia of 
the British Pyrales, p. 3, pl. 2, 1938. 


Maxillary palpus of male in the form of an aigrette. 


Forewing blackish gray, generally of a duskier hue 
than that of any of the preceding species; transverse 
lines usually faint and a dull whitish gray, rarely con- 
trasted against the ground color and when so, chiefly 
the lower half of antemedial line; the latter bordered 
outwardly at costa and inwardly at inner margin by 
blackish patches more or less contrasted against the 
ground color of the wing; a similar dark shade inwardly 
bordering the subterminal line; discal and terminal dots 
tending to coalesce, black. Hind wing dusky white 
between the veins; the latter appreciably darkened; a 
smoky shade along termen. Alar expanse, 25-30 mm. 


Male genitalia with harpe simple except for a thin, 
saucer-shaped, erect clasper with a toothlike projection 
from its upper inner angle. Aedeagus slender with its 
anterior end abruptly expanded and anterior margin 
straight (as in hypochalctella); one side produced into 
an extended, strongly sclerotized arm, sharply bent and 
pointed at apex (as in criddlella)*; penis armed with a 
single, moderately long, hairlike cornutus. Three pairs 
of ventrolateral hair tufts on eighth abdominal segment 
of male, some of the hairs broadly expanded at their 
apices. 

Female genitalia with bursa membranous; ductus 
bursae sclerotized for most of its length, the sclerotiza- 
tions extending for a short distance into the bursa; 
genital opening simple except for some weak granula- 
tions on and behind the ductus bursae. 

TYPE LOCALITIES: England (fusca, in BM); eastern 
Canada (moestella, in BM); Caribou Is., Labrador 
(frigidella, in MCZ); “New York’ ® (cacabella, in 
AMNH, ex Rutgers); Winnipeg, Manitoba (triplagia- 
tella, in USNM). 

Foon piants: Erica and probably some other Hrica- 
ceae. The only authentic Old World record is Erica 
(Meyrick 1938, Ford 1949). Ragonot (Monograph, 
p. 408) records Vaccinium myrtellus and Salix caprea as 
probabilities; but these plants only on the basis of food 
accepted in the laboratory by larvae hatched from eggs 
from gravid females by Porritt (Ent. Monthly Mag., 


5 This extension of aedeagus was misidentified by Pierce and 
Metcalfe (1938) as a cornutus. They overlooked the true 
cornutus attached to the vesica. 

6 So given in Hulst’s original description. 
ever, bears no locality label. 


The male type, how- 


vol. 19, p. 11, 1882). A female in the U. S. National 
Museum from Ottawa, Canada, was reared by James 
Fletcher (Aug. 1889) from a “black larva’’ found on 


Betula. Isuspect, however, that the larva had migrated 
to that plant. We have no other New World rearing 
records. 


Distrrisution: Holarctic. In the Old World from 
Great Britain to Japan. The American records from 
specimens are: Unirep Srarus: Maine, Orono; New 
Hampshire, Hampton (June), Mount Washington (July) ; 
Massachusetts, Framingham (July), Martha’s Vineyard 
(Aug.); New York, Rochester (June), Waterville (Aug.); 
Colorado, Glenwood Springs (Aug.); Washington, Pull- 
man. Canapa: Newfoundland, Port aux Basque (Aug.), 
St. George Bay (Harry’s River and Stephenville, Aug.), 
Spruce Brook (Aug.); Labrador, Caribou Isl., Hopedale, 
Nain; Nova Scotia, Baddeck (Cape Breton Isl., Aug.); 
Quebec, Chelesea (May); Ontario, Albany River (St. 
Martin’s Falls), Hymers (Aug.), Ottawa (June, Aug.); 
Manitoba, Aweme (June, July, Aug.), Winnpieg; Alberta, 
Banff (July), Calgary (Aug.), Edmonton (May); British 
Columbia, Fraser Mills (June), Kaslo (July, Aug.), 
Shawnigan Lake (Aug.), Victoria (July). AwasKa: 
Cordova, Fort Yukon, Juneau (July), Rampart (July). 

The species can be readily distinguished by its aede- 
agus, threadlike cornutus, and the peculiar sclerotization 
of its ductus bursae. On habitus and all its structural 
characters it is closely related to the gray-winged 
species of Pyla. Superficially it could easily be con- 
fused with impostor, equivoca, or aenigmatica. It is not 
congeneric with faecella (Zeller), the type of Laodama, 
to which genus Ragonot referred it. The latter differs 
markedly in male and female genitalia (figs. 427 and 
885), and on venation falls into our venational group D. 
Both Pyla and Laodamia have the cell of hind wing 
short; but in Laodamia vein 3 is appreciably longer in 
relation to vein 2 (fig. 52). 

Packard’s frigidella was retained by Ragonot as a 
separable variety from fusca, but it is at most only one 
of its color variants and is not entitled to any trinomial 
designation as a race. 

Several Old World references and synonyms have 
been omitted from the above synonymy. I do not 
question them, but have not been able to verify them. 
Anyone interested will find the names and references in 
Hulst (Phycitidae of N. Amer., 1890) and Ragonot 
(Monograph, 1893). 


296. Pyla hypochalciella (Ragonot), new combination 
Fieures 370, 854 


Nephopteryx ovalis hypochalciella Ragonot, N. Amer. Phycitidae, 
p. 7, 1887.—Hulst, Phycitidae of N. Amer., p. 144, 1890. 

Nephopteryx hypochalciella Ragonot, Monograph, pt. 1, p. 272, 
1893.—McDunnough, Check list, No. 6168, 1939. 

Pyla blackmorella Dyar, Ins. Inse. Menstr., vol. 9, p. 68, 1921.— 
McDunnough, Check list, No. 6248, 1939. (New syn- 
onymy.) 

Maxillary palpus of male in the form of an aigrette. 
Forewing very dark gray-brown, the dark areas of 
some of the darkest specimens almost black;a very faint 


146 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


powdering of white on costal half of median area and, 
narrowly, along terminal margin; the transverse lines 
powdery, grayish white, obscure on some specimens; 
antemedial line oblique, expanded slightly towards costa, 
without distinct inner dark border and with but faint 
indication of a blackish brown outer bordering shade 
towards costa; subterminal line more or less obscured, on 
well-marked examples preceded by a thin blackish line 
and followed by a broad band of the darkest prevailing 
ground color; discal dots black, well separated; a row of 
small black dots along termen, tending to fuse and on a 
few specimens forming a fine black line. Hind wing a 
uniform very dark satiny brown; the cilia paler, shading 
from pale brown to white at their tips. Alar expanse, 
22-26 mm. 

Male genitalia having harpe with an erect clasper 
armed along its outer margin with a row of stout spines 
(in the figure this looks like an enlargement of the base 
of costa, but it arises below costa and the base of the 
costa itself is simple). Anellus bearing two pairs of 
comblike, heavy spines, one pair ventral, one dorsal, the 
latter situated behind the former. Aedeagus slender; 
its anterior end abruptly expanded and the anterior 
margin straight; apical fourth bifid, the divided elements 
terminating in laterally curved horns. A single pair 
of simple hair tufts on eighth abdominal segment of 
male. 

Female genitalia with bursa membranous; ductus 
bursae partially flattened, sclerotized throughout, the 
sclerotization expanding abruptly into a wide funnel at 
genital opening. 

Type LocaLitiEs: ‘Washington Territory” (hypo- 
chalciella, in Paris Mus.); Mount Tzouhalem, southern 
Vancouver Isl., British Columbia (6lackmorella, in 
USNM). 

Foop prant: Unknown. 


Distrisution: Unitep Starrs: Washington, Friday 
Harbor (June, July). Canava: British Columbia (south- 
ern Vancouver Isl.), Cowichan District (June), Duncans 
(June), Mount Malahat (June), Mount Tzouhalem 
(June). 

The species is easily identified by its peculiarly armed 
anellus. In ground color of forewing it and the species 
following (hanhamella) are intermediate between typical 
gray- and brown-winged members of the two Pyla 
species groups. However, except for a very faint trace 
of it in hanhamella, they both lack the bronzy luster on 
forewing so characteristic of the typical brown group. 


297. Pyla hanhamella Dyar 
Figures 371, 860 
Pyla hanhamella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 109, 
1904.—McDunnough, Check list, No. 6239, 1939. 

Maxillary palpus of male subsquamous (a short brush 
of hairs mixed with flattened scales). 

Forewing color and markings as in hypochalciella 
except for traces of a metallic sheen at base. Hind 
wing pale brown; cilia white with a fine dark subbasal 
line. Alar expanse, 20-24 mm. 


Male genitalia having clasper of harpe a moderately 
long, erect spike; costa at base simple (not produced). 
Aedeagus slightly bent towards middle; shortly bifid at 
apex, the divided elements coarsely scobinate. A single 
pair of simple hair tufts on eighth abdominal segment. 

Female genitalia with the lobe of bursa giving off the 
ductus seminalis partially sclerotized, otherwise mem- 
branous; ductus bursae flattened, sclerotized through- 
out, concavely bent at middle, the sclerotization termi- 
nating in a sinuate, thickened, narrow, liplike band 
along the lower margin of the genital opening. 

TYPE LOCALITY: Winnipeg, Manitoba (type in 
USNM). 

Foop prant: Unknown. 

Distrisution: Manitoba, Aweme (June, July), Win- 
nipeg (June). 


Genus Pyla, Species 298-306: P. scintillans to 
P. viridisuffusella 
[Ground color of forewing bronzy brown.] 


298. Pyla scintillans (Grote) 
Figures 29, 372, 373, 857 

Nephopteryzr scintillans Grote, Papilio, vol. 1, p. 18, 1881. 

Pyla scintillans (Grote), New check list of North American 
moths, p. 55, 1882.—Hulst, Phycitidae of N. Amer., p. 161, 
1890.—Ragonot, Monograph, pt. 1, p. 482, 1893.—MeDun- 
nough, Check list, No. 6235, 1939. 

Pyla feella Dyar, Ins. Insc. Menstr., vol. 9, p. 68, 1921.—Mc- 
Dunnough, Check list, No. 6247, 1939. (New synonymy.) 

Maxillary palpus of male subsquamous (the scales 
short, flattened, forming a small expanded brush). 

Forewing dark bronzy brown; the scaling shiny, 
metallic; transverse lines absent, indicated only on well 
marked specimens by very faint, moderately broad, 
blackish brown bands (vestiges of their dark borders) ; 
discal and terminal dots obsolete. Alar expanse, 
20-26 mm. 

Male genitalia having harpe with strong clasper, 
developed as a stout outwardly curved hook with an 
extended, elongate, bladelike base, the latter more or 
less serrate. Considerable individual variation is shown 
in the clasper and the shape ofits base. Inone example, 
from Inyo County (presumably a variety of scintillans 
but possibly a distinct species), the clasper hook is 
markedly longer than in the examples figured, and the 
bladelike base narrower. Costa of harpe produced at 
base into a pointed, stout, very coarsely spined projec- 
tion. Aedeagus bifid for less than half its length; one 
of the divided elements with a short, thornlike spine 
projecting from lateral margin before apex; the other 
with 2 or 3 similar spines from lateral margin near apex 
(usually 3, rarely 2, a single specimen from El Dorado 
County, exhibiting only one). A single pair of ventro- 
lateral abdominal hair tufts on eighth segment. 

Female genitalia with bursa small, membranous 
throughout; ductus bursae very short, expanded abruptly 
into a sclerotized cup, its lower surface developed as a 
pair of flattened, pointed, elongate-oval blades which 
project beyond genital opening. Only trifling individual 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


differences can be distinguished between Dyar’s feella 
and females from other California localities. 

TYPE LOCALITIES: Summit, Sierra Nevada Mts., 
Calif. (scintillans, in BM); Bullfrog Lake (10,634 ft.), 
Sierra Nevada Mts., Calif. (feella, in USNM). 

Foop prant: Unknown. 

Disrrisution: California, Cisco (Placer County, 
July), Deer Park Springs (Lake Tahoe, July), El 
Dorado County (July), Inyo County (July), Mineral- 
king (Tulare County, July, Aug.), Sierra Nevada Mts. 
(Bullfrog Lake and Summit, Aug.), Tuolumne Meadows 
(July). 

299. Pyla sylphiella Dyar 
Fieures 375, 858 


Pyla sylphiella Dyar, Ins. Insc. Menstr., vol. 9, p. 68, 1921.— 
McDunnough, Check list, No. 6246, 1939. 

Maxillary palpus of male subsquamous. 

Superficially like scintillans, averaging a trifle darker; 
but distinguished only by its genitalia. Alar expanse, 
19-25 mm. 

Male genitalia with clasper of harpe similar to that 
of scintillans; produced enlargement of base of costa 
considerably stouter and more coarsely spined. Aede- 
agus with only a pair of lateral spines from adedeagus 
near its apex (one spine from each of the divided 
elements opposite and pointed away from each other). 
These differences are slight but appear to be consistent 
through long series. Female genitalia with ventral 
surface of the cup-shaped portion of ductus bursae 
bent into broad, deep, strongly sclerotized folds. 

Type tocatity: Mount Rainier, Wash. (type in 
USNM). 

Foop piant: Unknown. 

Distripution: Unirep States: Washington, Mount 
Rainier (Aug.), Paradise Valley (Mount Rainier, 
Aug.), Sheep Lake (Yakima County, Aug.), Skyline 
Ridge (Mount Baker District, Aug.), Slate Peak 
(Whatcom County, Aug.). Canapa: British Columbia, 
Mount Cheam (Aug.), Mount McLean (Aug.). 

The species is very close to scintillans but apparently 
distinct. The male genitalia differ only in minor details 
and the color and maculation offer little if anything to 
separate the two; but the female genitalia are markedly 
different and, from the specimens available, sylphiella 
appears to have a more northerly distribution. 


300. Pyla rainierella Dyar 
Fiaures 374, 859 


Pyla rainierella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 109, 
1904.—McDunnough, Check list, No. 6243, 1939. 

Maxillary palpus of male subsquamous. 

Moths averaging a trifle smaller than the preceding 
species (sylphiella), but certainly distinguished from it 
only by genitalia. Alar expanse, 16-20 mm. 

Male genitalia with production from base of costa 
of harpe considerably smaller and less coarsely spined 
than that of either sylphiella or scintillans. Aedeagus 
short, the apices of its divided elements bent abruptly 
downward as sharp, parallel, spinelike hooks. Female 


147 


genitalia with the sclerotized portion of the cup-shaped 
area of ductus bursae developed laterally as triangulate 
plates. 

Type tocatiry: Mount Rainier, Wash. (type in 
USN). 

Foop pirant: Unknown. 

Distrisution: Washington, Mount Rainier (Aug.), 
Paradise Valley (Mount Rainier, July), Sheep Lake 
(Yakima County, Aug.), Slate Peak (Whatcom 
County, Aug.), Table Mountain (Aug.). 

Like sylphiella, this species is chiefly distinguished by 
its female genitalia, the sclerotized area of ductus 
bursae at genital opening resembles somewhat that of 
Jasciella but differs in shape and is like that of no other 
species in the genus. The aedeagus easily separates 
the male of fasciella from either scintillans or sylphiella 


301. Pyla aeneella Hulst 
Ficures 376, 864 


Pyla aeneella Hulst, Canadian Ent., vol. 27, p. 55, 1895.—Mc- 
Dunnough, Check list, No. 6242, 1939. 

Maxillary palpus of male in the form of a semiaigrette 
(the hairs short). 

Forewing unicolorous, without any trace of dark 
transverse shadings or discal spots; brown with a bronzy 
green irridescence. Hind wing concolorous with fore- 
wing. Alar expanse 23-25 mm. 

Male genitalia with costa of harpe at base produced 
into a thin, rounded lobe with finely serrate edge; 
clasper erect, short, stout, thornlike. Aedeagus short, 
broadest at middle, divided to middle; the divided ele- 
ments rather broadly flattened and abruptly, asym- 
metrically bent at their apices. Female genitalia with 
ductus bursae broadly cup-shaped from shortly beyond 
its junction with bursa; the ventral surface weakly 
sclerotized, granulate, and with slight infoldings on the 
lower median area. 

Typr Locaity: Colorado (in AMNH, ex Rutgers). 

Distrisution: Colorado; Utah, Silver Falls (July), 
Stockton (May, June). 

A good series of the Utah specimens is in the National 
Collection. The genitalia of the females agree in every 
detail with those of the Colorado type in the Rutgers 
Collection. The metallic iridescence of the forewings 
is conspicuous but its greenish tint is very faint. 


302. Pyla aeneoviridella Ragonot 
Ficures 378, 862 


Pyla aeneoviridella Ragonot, N. Amer. Phycitidae, p. 9, 1887; 
Monograph, pt. 1, p. 482, 1893.—Hulst, Phycitidae of N. 
Amer., p. 161, 1890.—Barnes and McDunnough, Contribu- 
tions, vol. 2, p. 222, 1914—McDunnough, Check list, No. 
6237, 1939. 


Maxillary palpus in the form of a semiaigrette (the 
scales somewhat flattened, not so decidedly hairlike as 
in aeneella). 

Forewing unicolorous bronzy brown, without dark 
markings of any kind; the iridescent scaling with a 
faint greenish tint and somewhat more strongly con- 
centrated at the base of the wing than in the median 


148 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


and outer areas. The color of both the fore and hind 
wings is similar to that of aeneella and the two species 
can only be safely distinguished by their genitalia. 
Alar expanse, 23-29 mm. 

Male genitalia with an angulate, serrate and coarsely 
spined projection from costal base of harpe; clasper out- 
bent, strongly spined and more or less serrate. Aedea- 
gus short, slightly bent at middle and with a single stout, 
stubby spine projecting from one side (extent of indi- 
vidual variation shown in figs. 378a, b). Female geni- 
talia with a weak sclerotization of the lobe of bursa 
giving off the ductus seminalis; ductus bursae with 
ventral surface of cup-shaped area more strongly 
sclerotized than that of aeneella and differently sculp- 
tured. 


TYPE LocaLity: Evanston, Wyo. (type in Paris Mus.)- 

Foop prant: Unknown. 

Distrisution: Unitep States: Colorado, Tennessee 
Pass (July); Wyoming, Big Horn Mts. (July), 
Evanston, Yellowstone Park (July, Aug.); Montana, 
Bozeman (July), Glacier Park (June); Oregon, Wallowa 
Mts. (Arnold Lake, July); Washington, Olympic Mts. 
(Hurricane Ridge, June, July). Canapa: Alberta, 
Laggan (July). 

The Washington and Oregon specimens are consider- 
ably darker than those from the other localities, the 
specimens from Oregon having almost black hind wings 
and blackish brown forewings. 

In his original description and in his Monograph 
Ragonot gives “N. Y.’’ as the type locality. This was 
a misreading of the label of his type. ‘The correction 
was made by Barnes and McDunnough in the reference 
cited above. 


303. Pyla metalicella Hulst 
Figures 377, 863 


Pyla metalicella Hulst, Canadian Ent., vol. 27, p. 54, 1895.— 
McDunnough, Check list, No. 6236, 1939. 

Maxillary palpus of male in the form of a semiaigrette 
(the hairs rather short). 

Appreciably lighter than aeneoviridella. The fore- 
wing a unicolorous light bronzy brown with greenish 
yellow iridescence; discal dots more or less distinct, 
separated, blackish; no other markings. Average size 
larger than that of aeneoviridella. Alar expanse, 25-32 
mm. 

Male genitalia with a long, somewhat flattened, out- 
wardly curved clasper on harpe (similar to that of 
wmpostor, but proportionally longer); costa of harpe at 
base simple. Aedeagus very shortly divided at apex; 
a pair of very short, sharp, straight spines on ventral 
surface near apex. 

Female genitalia with lobe of bursa giving off ductus 
seminalis weakly sclerotized; cuplike area of ductus 
bursae, funnel shaped (triangulate), strongly sclerotized 
over its entire ventral surface and containing a broad, 
centrally located, funnel-shaped fold. Individual vari- 
ations in this fold are shown in figs. 863 and 863a. 

Type Locauity: Colorado (type in AMNH, ex 
Rutgers). 


Foop prant: Unknown. 

DistriBuTion: Colorado, Silverton (July, Aug.), and 
two specimens with only the state locality; Utah, Silver 
Lake (July). The Silverton locality is represented in 
the National Collection by 17 specimens. Also in the 
National Collection is a female from Colorado (Bruce), 
labeled ‘“Pyla aeneella Hulst, Type,” another of Hulst’s 
pseudotypes and possibly part of his original ‘“‘type” 
series of aeneella. ‘The actual type of metalicella is a 
male with only the state locality. Its genitalia agree 
in every detail with those from Silverton specimens. 


304, Pyla fasciella Barnes and McDunnough 
Figures 379, 861 


Pyla fasciella Barnes and McDunnough, Canadian Ent., vol. 49, 
p. 405, 1917.—McDunnough, Check list, No. 6244, 1939. 

Maxillary palpus of male minute, squamous. 

Forewing blue-black, some paler bluish iridescence 
over basal area and (in strong light) a faint, brownish 
iridescence in outer area; antemedial line represented 
by a narrow oblique black band near, but before middle; 
subterminal line a similar curved band well back from 
and parallel with termen; the area between the two 
black bands darker than remainder of wing, forming a 
faint, broad, median, black fascia; discal dots obsolete. 
Hind wings very dark brown, shiny. Alar expanse, 
21-24 mm. 

Male genitalia without clasper on harpe; base of 
costa of harpe produced into a knoblike projection, 
finely spinose along margin. Aedeagus simple. A 
single pair of ventrolateral hair tufts on eighth segment. 
Female genitalia similar to those of rainierella except 
that the paired plates of ductus bursae at genital open- 
ing are narrower and differently shaped. 

Typrk tocatity: Mount Shasta, Calif. (type in 
USNM). 

Foop prant: Unknown. 

Known only from northern California. Represented 
in the National Collection by the type series from 
Mount Shasta, 7,000 ft., July—three males and two 
females (not four males and two females as given in the 
original description) ; and one male from Bartle, Calif. 
(June 14, 1939, Grace H. and John L. Sperry). In 
their original description the authors give the expanse 
as “24-31 mm.” This is probably a printer’s error, 
for the largest specimen before me is a scant 24 mm. 


305. Pyla nigricula, new species 
Fieure 380 


Maxillary palpus of male minute, squamous. 

Superficially like fasciella except: Transverse dark 
lines of forewing obsolete, only the antemedial black 
band very faintly indicated; no contrasted dark median 
fascia; the entire median and outer areas a dark 
purplish brown. Alar expanse, 26 mm. 

Male genitalia with uncus broader and squattier than 
that of fasciella. Projection from costal base of harpe 
differently shaped, bluntly poimted; clasper developed 
as a stout, smooth, curved, pointed hook. Aedeagus 
simple. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 149 


Typrxocauity: Verdi, Nev. (type in USNM, 61354). 

Foop piant: Unknown. 

Described from unique male collected by A. H. 
Vachell, “June 1 to 10.”” This specimen had been in 
our collection under scintillans Grote. 


306. Pyla viridisuffusella Barnes and McDunnough 
Figures 381, 851 


Pyla viridisuffusella Barnes and McDunnough, Canadian Ent., 

Mayan’ p. 406, 1917—McDunnough, Check list, No. 6245, 

Maxillary palpus of male in the form of a short 
aigrette. 

Forewing heavily suffused with light, bronzy green, 
irridescent scaling, especially strong over basal area and 
in a line indicating the subterminal line; two transverse 
blackish bands, an oblique, antemedial one and another 
forming an inner border to the subterminal line; discal 
dots, when distinguishable, confluent, forming a line 
along discocellular vein. Hind wing very dark brown. 
Alar expanse, 17-20 mm. 

Male genitalia with harpe simple. Aedeagus bifur- 
cate to middle, the divided elements asymmetrical (one 
longer than the other). Female genitalia with a pair 
of strongly sclerotized, convolute bands extending from 
posterior end of bursa well into ductus bursae; genital 
opening simple. 

Tyrer ocauity: Tuolumne Meadows, 
County, Calif. (type in USNM). 

Foop prant: Unknown. 

Distrisution: California, Humphreys Basin (Fresno 
County, Aug.), Johnsons Park (Sierra Nevada Mts.), 
Kernick Meadows (9,250 ft., July), Mineralking (Tulare 
County, July, Aug.), Tuolumne Meadows (July, Aug.). 

The most brilliant of the Pyla species. Its genitalia, 
both male and female most resemble those of fasciolalis 
Hulst. 


Tuolumne 


Genera 77 and 78: Dioryctria and Oryctometopia 


[Venational division D. Forewing with veins 4 and 5 closely 
approximate for a short distance from cell (Dioryctria), or connate 
or very shortly stalked (Oryctometopia); vein 6 straight or bent 
towards base; 10 from the cell. Hind wing with cell less than 
one-third the length of wing; discocellular vein curved; veins 4 
and 5 stalked for at least half their lengths. Male antenna with 
@ shallow sinus or slight incurvation in base of shaft, containing 
& row of fine spines or a weak scale tuft. Male genitalia with 
transtilla incomplete or absent; harpe with costa strongly scle- 
rotized and produced at apex (Dioryctria) or with one or more 
short, stout, thornlike spines from lower margin of sacculus.] 


77. Genus Dioryctria Zeller 


Dioryctria Zeller, Isis von Oken, 1846, p. 632.—Heinemann, Die 
Schmetterlinge Deutschlands und der Schweiz, Abt. 2, vol. 
1, pt. 2, p. 148, 1865.—Ragonot, Ent. Monthly Mag., vol. 
22, pp. 52, 56, 1885; Monograph, pt. 1, p. 187, 1893.—Hulst, 
Phycitidae of N, Amer., p. 134, 1890.—Spuler, Die Schmet- 
terlinge Europas, vol. 2, p. 213, 1910.—Forbes, Cornell Univ. 
Agr. Exp. Station, Mem. 68, p. 619, 1923.—Meyrick, Re- 
vised Handbook of British Lepidoptera, p. 383, 1928.— 
Bisset, in Pierce and Metcalfe, Genitalia of the British 


Pyrales, p. 57, 1938 (notes Ragonot’s fixation (1885) of 
type of genus).—Janse, Journ. Ent. Soc. South Africa, vol. 
4, p. 161, 1941. (Type of genus: Tinea abietella Denis and 
Schiffermiiller.) 

Pinipestis Grote, Canadian Ent. vol. 10, p. 19, 1878; Bull. U. 8S. 
Geol. Geogr. Surv. Terr., vol. 4, p. 699, 1878.—Hulst, 
Phycitidae of N. Amer., p. 136, 1890. (Type of genus: 
Nephopteryx zimmermani Grote.) 

Tongue well developed. Antenna finely pubescent 
or very shortly ciliate, rarely unipectinate (in males of 
some Old World species, pineae Staudinger, mendacella 
Standinger); on all males a shallow sinus in base of 
shaft, containing a short row of minute black, thornlike 
spines, more or less concealed by rough scaling. Labial 
palpus upturned, reaching to or a trifle above vertex; 
second segment grooved on inner side; third segment 
short (less than one-third the length of second), acumi- 
nate. Maxillary palpus of male small and squamous or 
(rarely) in the form of an aigrette. Forewing smooth 
or with two or more tufts or raised (ruffed) scales; 11 
veins; vein 2 from before lower outer angle of cell; 3 from 
the angle, slightly nearer to 4 than to 2 at base; 4 and 5 
from the cell, approximate at and for about one-fourth 
their distance from cell; 6 bent towards base and from 
close to upper angle of cell (in type species, straight and 
from well below the angle on some specimens); 10 from 
the cell, approximate to the stalk of 8-9 for some dis- 
tance beyond cell; 8 and 9 long stalked; male without 
costal fold. Hind wing with vein 2 from before lower 
outer angle of cell; 3 from the angle, long (almost as long 
as 2); 4 and 5 stalked for half or slightly more than half 
their lengths; 7 and 8 closely approximate for less than 
half their lengths beyond cell; rarely shortly anasto- 
mosed; cell Jess than one-third the length of wing; dis- 
cocellular vein curved, outwardly extended at lower 
angle. Eighth abdominal segment of male with com- 
pound ventral scale tufts. 

Male genitalia decidedly elongated (least so in gulo- 
sella). Uncus broad, stout, its terminal margin more or 
less broadly rounded. Apical process of gnathos a 
short, stout hook. ‘Transtilla incomplete or absent; its 
lateral elements, when distinguishable, slender and 
usually attached to costal base of harpe. Harpe with 
costal area broadly sclerotized and produced at apex; 
cucullus narrowly elongate, bluntly pointed or very 
narrowly rounded at apex; sacculus short, simple; 
clasper present, erect, usually finely haired at or near 
apex, but not bearing strong spines, thorns, or serra- 
tions. Anellus with well-developed lateral lobes. Ae- 
deagus long, moderately stout; penis with strong cor- 
nuti, consisting of numerous straight, slender spines (as 
long or nearly as long as width of aedeagus) and usually 
one or more longer, stouter spines placed back of them 
on the penis. Vinculum stout; longer than broad 
(frequently considerably elongated), gradually tapering 
to a moderately broad, rounded, or abruptly angled 
terminal margin. 

Female genitalia with well-developed signa, consisting 
of two or three clusters of strong, slender spines, their 
bases in each of the clusters more or less fused into 
sclerotized plates; the clusters in end of bursa near 


150 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


ductus bursae and one of them frequently in or extend- 
ing into the ductus; ductus bursae flattened, strongly 
sclerotized over most of its length, the sclerotization 
more or less longitudinally wrinkled and terminating just 
before the simple genital opening; ductus seminalis from 
bursa near the junction of bursa and ductus bursae. 

Dioryctria is one of the most, if not the most, distinct 
and sharply defined genus in the Phycitidae; and is so 
despite the variations in structure exhibited by its male 
antennae, male maxillary palpi, and its wing scaling. 
Its male and female genitalia have a characteristic 
habitus, difficult to describe, but easily observed in slides 
or figures. There is also a maculation character of the 
forewing common to nearly all the species; the usual 
black dots at end of cell are absent, being replaced by a 
white spot or line on the discocellular vem. The only 
North American species without such a marking is 
clarioralis, where a considerable area about the end of 
cell is clouded with a dark suffusion. The raised-scale 
character upon which Grote distinguished his genus 
Pinipestis from Dioryciria consists of a mere rufiing of 
the scales on two or three spots on the forewing. It is 
at best an elusive character, subject to opinion as much 
as to observation, and in some of the forms(zimmermani, 
cambiicola) the tufts may be either raised or flattened on 
unspread specimens. Naturally, on spread specimens 
they are usually flattened, whatever was their condition 
in nature. Ragonot was perfectly justified (1893) in 
relegating Pinipestis to synonymy. 

As here defined the genus is of world-wide distribution 
in the Northern Hemisphere and all the species whose 
larval habits are known are borers in conifers. Most of 
the American species are represented in the National 
Collection by numerous specimens reared in connection 
with the forest-insect investigations of the U.S. Bureau 
of Entomology and Plant Quarantine. 


307. Dioryctria abietella (Denis and Schiffermiiller) 
Fieures 51, 382, 865 


Tinea abietella Denis and Schiffermiiller, Systematische Ver- 
zeichniss der Schmetterlinge der Wienergegend . . . , p. 138, 
1776.—Fabricius, Mantissa insectorum . . . , vol. 2, p. 245, 
1787; Entomologica systematica . . . , vol. 3, pt. 2, p. 302, 
1793. 

Tinea decuriella Hibner, Sammlung europaischer Schmetterlinge, 
p- 35 and Lepidoptera 8, Tineae 2, pl. 11, fig. 74, 1796. 
Phycis abietella (Denis and Schiffermiiller) Zincken, in Germar 
and Zincken, Mag. der Ent., vol. 3, p. 160, 1818.—Treitschke, 
Die Schmetterlinge von Europa, vol. 9, p. 177, 1832.— 
Ratzeburg, Die Forst-Insecten . . . , vol. 2, p. 244, pl. 15, 

fig. 2, 1840. 

Dioryctria abietella (Denis and Schiffermiiller) Zeller, Isis von 
Oken, p. 736, 1846——Heinemann, Die Schmetterlinge 
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 148, 
1865.—Ragonot, Ent. Monthly Mag., vol. 22, p. 52, 1885; 
Monograph, pt. 1, p. 198, 1893.—Hulst, Phycitidae of N. 
Amer., p. 135, 1890.—Spuler, Die Schmetterlinge Europas, 
vol. 2, p. 213, 1910.—Joannis, Ann. Soc. Ent. France, vol. 
85, p. 259, 1916; Bull. de la Station de Recherches forestiérs 
du Nord de 1’ Afrique, vol. 1, p. 192, 1921.—Forbes, Cornell 
Mem. 68, p. 621, 1923.—Meyrick, Revised handbook of 
British Lepidoptera, p. 384, 1928.—Pierce and Metcalfe, 
Genitalia of the British Pyrales, p. 2, pl. 1, 1938.—Keen, 
U.S. Dep. Agr. Mise. Publ. 273, p. 39, 19838.—McDunnough, 


Check list, No. 6129, 1939.—Janse, Journ. Ent. Soc. South 
Africa, vol. 4, p. 161, 1941—MacKay, Canadian Ent., vol. 
75, p. 94, 1943.—Craighead, U. S. Dep. Agr. Misc. Publ. 
657, p. 451, 1950. 

Pinipestis abietivorella Grote, Bull. U. S. Geogr. Geol. Surv. Terr., 
vol. 4, p. 701, 1878. 

Pinipestis reniculella Packard (not Grote), U. 8S. Dep. Agr. Ent. 
Bull. 13, p. 21, 1887; U.S. Dep. Agr. Fifth Rep. Ent. Comm., 
p. 854, 1890. 

Myelois elegantella Hulst, Canadian Ent., vol. 24, p. 59, 1892. 


Maxillary palpus of male squamous. 


Forewing smooth; bluish gray, dusted with white, the 
white dusting of variable intensity, when pronounced, 
more or less concentrated in basal area, along the outer 
border of antemedial line, the inner border of subter- 
minal line and, weakly, along terminal margin; trans- 
verse lines white, distinct, narrow; antemedial line 
oblique, inwardly angled at cell and vein 1b, preceded 
on costa by a blackish shade and bordered outwardly 
by a black line, and usually preceded by a pale patch 
on inner margin, this often no more than a smear of 
olivaceous white scales and never so conspicuous or 
well contrasted as in reniculella; subterminal line 
sinuate-angulate, preceded and followed by blackish 
bordering lines; discal mark a white, lunate spot; a fine 
black line along terminal margin. Hind wing dusky 
white, darkened slightly towards outer margin and on 
the veins. Alar expanse, 20-30 mm. 

Male genitalia with uncus subtriangulate and rather 
narrow in normal position; a short, slight incurvation 
of the lateral margins near base, and the lateral margins 
themselves infolded. (When uncus is flattened in prep- 
arations and the lateral folds pushed out, the uncus 
appears as in fig. 382, but never takes the form of the 
flattened unci of the species which have a longer incurva- 
tion of the lateral margins, such as zimmermani). Harpe 
with one or more spines projecting from the terminal 
margin of the sclerotized costal area below its apex 
(there is considerable individual variation in this feature, 
a few examples of which, from small American speci- 
mens, are shown in figs. 882c-e). Penis armed with a 
single stout spine behind anterior spine cluster. 

Female genitalia chiefly distinguished by a longitu- 
dinal fissure on the ventral surface of the sclerotized 
portion of ductus bursae, variations of which are shown 
in figures 865 and 865a. The females of sysstratiotes 
from Guatemala also show traces of such a fissure but 
this species is only doubtfully distinct from abietella. 

TYPE LOCALITIES: Austria (abietella, location un- 
known); Germany (decuriella, type lost); Amherst, 
Mass. (abietivorella, in BM); Seattle, Wash. (elegantella, 
in AMNH, ex Rutgers). ; 

Foop PLants: Pine, spruce, fir; in the spruces and firs 
(Abies, Pseudotsugha, etc.) chiefly in the cones. The 
favored host seems to be Pinus of which it attacks all 
species. The larvae exhibit a variety of habits. They 
are both primary and secondary. ‘They bore into new 
and otherwise uninfested terminals and into terminals 
that have been attacked by Rhyacionia buoliana or the 
white pine weevil (Pissodes strobi.) They attack both 
healthy and diseased cones. They bore into and feed 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 151 


on the cambium of smooth bark on the trunks and 
branches; and also feed in the galls on any part of the 
tree. 

Disrrisution: Apparently throughout the range of 
the genus Pinus in the Northern Hemisphere. Ameri- 
can records from specimens (moths) before me as 
follows: Unirep States: Maine, Orono (Aug.); Massa- 
chusetts, Amherst, Framingham (Oct.), Martha’s Vine- 
yard (Sept.), Pepperell (Aug.); Connecticut, Bradford, 
Lyme (Aug.), New Haven (June); New York, Long 
Island (Garden City, Oct., Great Neck, July), Warrens- 
burg (Sept.); New Jersey, Lakehurst (May); District 
of Columbia, Washington (July, Aug.); Florida, Alton 
(June), Eustis (June, July), Gainesville (June), Orlando 
(June), ‘So. Florida” (June, July, Aug.); Jdlinois, Dun- 
dee; Nebraska, Halsey (Apr., June, Aug.); Montana, 
Dillon (July), Elliston, Evaro (Mar.); Colorado, Glen- 
wood Springs; Arizona, Prescott (July); California, 
Berkeley, Patrick’s Creek (Sept.), Sacramento, Shasta 
Retreat (July); Oregon, Ashland (July, Aug., Sept., 
Oct.), Salem (Aug.), Colestin (June), Silver Lake (Aug.), 
Sprague River (July); Washington, Hoquiam, Pullman, 
Rock Lake (Whitman County, Sept.); Seattle. Can- 
apa: Labrador, Dublin Shore (Lunenburg County) ; 
Quebec, Montreal (June); Saskatchewan, Lutherland 
(June, Aug., Sept.); British Columbia, Kaslo (June). 
GuateMaLa: A series of males and females in the U. S. 
National Museum, reared from pine cones, May 1927, 
by J. G. Salas and labeled simply “Guatemala, C. A.’ 

The species is of considerable economic importance, 
especially to young pine trees in our Western States, 
and particularly in reforestation areas. It has an 
extensive literature in the Old World. I have listed 
here only the more important references and have 
omitted purely European synonyms. For additional 
references the reader is referred to Ragonot (Mono- 
graph, p. 198), Hulst (Phycitidae of N. Amer., 1890), 
the Journal of Economic Entomology, and the Review 
of Applied Entomology. The most satisfactory infor- 
mation on life history and larval habits will be found in 
the MacKay (1943) and Craighead (1950) papers. 


308. Dioryctria sysstratiotes Dyar 
Ficure 866 


Dioryctria sysstratiotes Dyar, Ins. Insc. Menstr., vol. 7, p. 48, 


Forewing smooth, similar to that of abietella except: 
A pale brownish suffusion in median area forming a 
rather large patch below discal spot; a similar brownish 
shade outwardly bordering subterminal line; the patch 
preceding antemedial line on inner margin, more dis- 
tinct, larger, pale olivaceous brown. Hind wing some- 
what darker, translucent smoky white with a very faint 
brownish tint towards outer margin. 

Alar expanse, 23-28 mm. 

Female genitalia as in abietella except no (or only a 
faint trace of) median ventral cleft in ductus bursae. 

TyPE Locauity: Cayuga, Guatemala (type in 
USNM). 

Foop PLant: presumably Pinus. No rearing records. 


Disrripution: GuaTEMALA: Cayuga (June), Chejel 
(June), Purulhé (July). 

Known only from females. Doubtfully distinct from 
abietella except as arace or colorform. Its exact status 
will have to await discovery of a male. 


309. Dioryctria reniculella (Grote) 
Figures 383, 867 


Pinipestis reniculella Grote, North Amer. Ent., vol. 1, p. 67, 


Dioryctria reniculella (Grote), Ragonot, Monograph, pt. 1, p. 200 
1893.—Forbes, Cornell Mem. 68, p. 620, 1923.—McDun™ 
nough, Check list, No. 6131, 1939.—Brown, Canada Dep- 
Agr. Publ. 712, Techn. Bull. 31, p. 13, 1941.—MacKay, Cana- 
dian Ent., vol. 75, p. 94, 1943.—Craighead, U. 8. Dep. 
Agr. Misc. Publ. 657, p. 451, 1950. 

Maxillary palpus of male squamous. 

Forewing smooth; in color and maculation similar to 
that of abietella except: Ground color pale brownish 
gray; the transverse lines and discal spot more sharply 
contrasted, more distinctly white; usually a rather large 
olivaceous patch on inner margin preceding the ante- 
medial line; hind wing darker, pale smoky fuscous. 
Alar expanse, 22-26 mm. 

Male genitalia with uncus tonguelike; more elongate 
and narrower than that of any other American species; 
its terminal margin very narrowly rounded; no incur- 
vation of the lateral margins (its shape not appreciably 
altered by flattening in slide preparations). A very 
slight spur from the lower outer angle of the sclerotized 
costal area of harpe, but no other spines from below 
apex of costa. Penis armed only with anterior spine 
cluster; no single stout spine on penis behind the an- 
terior cluster. 

Female genitalia with only one strong spine cluster, 
that at junction of bursa and ductus bursae; girdle of 
spines in bursa before junction with ductus, weak, 
broken, and the spines themselves greatly reduced. 

Type Locauity: New York (type in BM). 

Foop Puants: Various spruces; rarely in balsam fir 
and tamarack. Reported as occasional in jack pine, 
but such records are doubtful. The larvae feed in 
terminals and cones and to a lesser extent upon the 
foliage of the terminals. 

Distrisution: Unirep Srates: Maine, Blue Hill 
(July), Sebee Lake (July); Connecticut, East River 
(July), New Haven (June, July); New York; Illinois, 
Putnam County (July) ; Michigan, East Lansing (Aug.) ; 
Colorado, Estes Park (July); California, Fallen Leaf 
Lake (Aug.), Mount Lowe (July); Washington, Belling- 
ham (Aug.), Hoquiam, Kamiack Butte (Aug.), Pullman 
(July, Aug.). Canapa: Nova Scotia, Cape Breton 
(Aug.); Quebec, St. Therese Isl. (St. John’s County, 
July); Ontario, Westree; Saskatchewan (June); British 
Columbia, Seton Lake (June, July), Victoria (July). 

The species is easily distinguished by its genitalia. 
In the past it has been frequently confused with 
abietella and until 1893 was treated by Hulst and 
Ragonot as a synonym of the latter. The larvae of the 
two species often occur together in spruce cones, so 
there is no safe way to separate them on their larval 


152 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


habits. However, reniculella is primarily a spruce 
feeder, while abietella shows a marked preference for 
the pines. Both species are borers, as are all the 
Dioryciria species. It is unfortunate that Miss Mac- 
Kay in her otherwise excellent paper (1943) should refer 
to reniculella as ““The spruce foliage worm.” It does 
feed to some extent upon the foliage of terminal shoots; 
but likewise, and more frequently within the terminals 
themselves and in the cones, and in economic literature 
is quite properly referred to as ‘“‘the spruce cone moth.” 
It is a strictly American species limited in distribution, 
apparently, to the northern United States and Canada. 


310. Dioryctria ponderosae Dyar 
Figures 384, 868 


Dioryciria ponderosae Dyar, Ins. Insc. Menstr., vol. 2, p. 2, 
1914.—McDunnough, Check list, No. 6130, 1939. 


Maxillary palpus of male squamous. 

Forewing smooth; like that of abietella except: Black- 
ish markings more strongly contrasted; a broad black 
band inwardly bordering the antemedial line, a similar 
band on some specimens of abietella but not so broad 
nor so well contrasted; subterminal line outwardly 
angled at middle; the white transverse lines and discal 
spot well contrasted. Hind wing white, clouded with 
smoky fuscous towards outer margin; the veins slightly 
darkened. Alar expanse, 27-30 mm. 

Male genitalia with uncus considerably shorter and 
broader than that of either abietella or reniculella; its 
terminal margin broadly rounded; lateral margins in- 
curved near its base. Harpe with apex of sclerotized 
costa produced into a blunt, curved hook, no spine from 
lower outer angle of the sclerotized costal area; clasper 
reduced. Female genitalia distinguished by its short 
ductus bursae: 

TypE Locatity: Lamedeer, Mont. (type in USNM). 

Foop prant: Pinus ponderosa. Larva a borer in the 
cambium of the bark. 

Distrisution: Montana, Lamedeer (June), Colorado, 
Glenwood Springs (June, July, Aug.); California, 
American River (May). 

A distinct species known only from a half-dozen 
specimens from the above localities. It belongs defi- 
nitely with the smooth-winged Dioryctria, although a 
few roughened scales can be distinguished in the black 
inner borders of the antemedial line on a couple of the 
specimens. Its genitalia separate it easily from all 
other smooth-winged species. The name has been 
misapplied to a color variety of zimmermani. This 
misidentification is discussed under the latter name. 
The Missoula, Mont., female (Hopkins U.S. No. 11508) 
mentioned by Dyar in his description of ponderosae 
belongs to this variety of zimmermani. 


311. Dioryctria majorella Dyar 
Fiaures 385, 871 


Dioryctria majorella Dyar, Ins. Insc. Menstr., vol, 7, p. 43, 1919. 
Dioryctria muellerana Dyar, Ins. Insc. Menstr., vol. 7, p. 85, 1919 
(new synonymy). 


Maxillary palpus of male squamous. 


Forewing smooth, similar in color and markings to 
that of sysstratiotes except: Somewhat more suffused; 
the whitish markings fainter and a duller, more sordid 
white; the pale discal spot obscure; the blackish inner 
border of the subterminal line and the blackish band 
preceding the antemedial line broader. Hind wing 
smoky white, the veins faintly darkened. Alar expanse, 
28-33 mm. 

Male genitalia distinguished by greatly increased 
width of the sclerotized costal area of harpe; a short 
spur projects from the outer margin of this sclerotized 
area just below its apex. D. erythropasa has a similar 
harpe; but differs markedly in other details of the geni- 
talia—differently shaped uncus, narrower clasper, and 
different spining of penis. 

Female genitalia differ from those of sysstratiotes only 
in insignificant details. The female genitalia of both 
species differ from those of abietella chiefly in the greater 
broadening and thickening of the membrane of ductus 
bursae near its junction with bursa copulatrix. 

TYPE Locauities: Jalapa, México (majorella, 9, in 
USNM); Zacualpén, México (muellerana, o, in USNM). 

Foop pPLant: Pine. 

Known only from Dyar’s two types and a pair (o' and 
9) from British Honduras. The Honduran specimens 
are smaller than the Mexican types and in wretched 
condition, but their genitalia are a perfect match in all 


details. In his description of muellerana Dyar sug- 
gested that it might be the male of majorella. This 
synonymy is proven by the Honduran examples. They 


were reared from cones of Pinus caribaea. 


312. Dioryctria disclusa Heinrich 
Fiaure 372 


Dioryctria disclusa Heinrich, im Farrier and Tauber, Iowa State 
Coll. Journ. Sci, vol. 27, p. 495, 1953. 


Maxillary palpus of male squamous. 

Forewing smooth; basal area to antemedial line 
orange yellow; area beyond brownish red, more or less 
shaded or suffused with yellowish orange (on some 
specimens the ground color of the entire wing yellowish 
orange), usually the red shade most conspicuous in the 
area between subterminal line and termen; transverse 
lines narrow, white; a white streak along lower margin 
of cell between the transverse lines; antemedial line 
faint, oblique, nearly straight ; subterminal line stronger, 
set well out, rather near terminal margin, outwardly 
angulate between veins 6 and 1b; discal mark (when dis- 
tinguishable) a white line along discocellular vein; some 
very short white dashes on terminal margin; cilia smoky 
white. Hind wing smoky white to pale smoky gray, the 
paler examples showing a very faint ocherous tint; 
veins slightly darkened; cilia whitish. Alar expanse, 
24-29 mm. 

Male genitalia like those of auranticella. Female 
genitalia essentially like those of auranticella. The 
differences shown in the figures for the two species 
are the extremes and represent, at most, individual 
variations. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 153 


Typrr Locauity: Tryon, N.C. (type in USNM). 

Foop piants: Pinus spp. Larvae feeding in the 
cones. 

DistrisuTion: Massachusetts, Martha’s Vineyard 
(June) ; New Jersey, Lakehurst (July) ; District of Colum- 
bia; West Virginia, Roosevelt (June), North Carolina, 
Raleigh (June), Tryon (June); Jowa, Ames (June). 

Examples of this species have been in the National 
Collection as auranticella and it was on the basis of this 
misapplication of Grote’s name that Dyar described his 
zanthaenobares. The true auranticella is strictly a 
western species, while the distribution of disclusa, as 
far as I know, is limited to the eastern and central areas 
of the United States. 

The paper by Farrier and Tauber gives all the infor- 
mation known on the life history, food habits, and 
behavior of the insect. 


313. Dioryctria auranticella (Grote) 
Ficures 386, 873 


Nephopteryz auranticella Grote, Ann. Mag. Nat. Hist., ser. 5, vol. 
11, p. 57, 1883; Trans. Kansas Acad. Sci., vol. 8, p. 57, 1883. 

Dioryctria miniatella Ragonot, N. Amer. Phycitidae, p. 4, 1887.— 
Hulst, Ent. Amer., vol. 5, p. 156, 1889. 

Dioryctria auranticella (Grote) Hulst, Phycitidae of N. Amer., p. 
134, 1890.—Ragonot, Monograph, pt. 1, p. 194, 1893. 
McDunnough, Check list, No. 6126, 1939. 

Dioryctria zanthaenobares Dyar, Proc. Ent. Soc. Washington, vol. 
13, p. 81, 1911.—Keen, U. S. Dep. Agr. Misc. Publ. 273, p. 
38, 1938.—McDunnough, Check list, No. 6127, 1939.— 
Heinrich, in Farrier and Tauber, Iowa State Coll. Journ. 
Sci., vol. 27, p. 495, 1953. 

Maxillary palpus of the male in the form of an 
aigrette. 

Forewing smooth; color and markings similar to those 
of disclusa, but on the average somewhat paler, the 
ground color ranging from yellowish orange to brownish 
red; on the darker suffused, reddish examples the yel- 
lowish color of the basal area is less contrasted than in 
ordinary disclusa. Very pale examples of the two 
species cannot be distinguished superficially. In size 
auranticella averages larger. Alar expanse, 27-33 mm. 

Male genitalia with apex of cucullus of harpe extend- 
ing beyond apex of the sclerotized costa. Female 
genitalia exhibiting only slight and comparative differ- 
ences from those of dasclusa. 

Tyrer Locaities: New Mexico (auranticella, in Univ. 
Kansas); Arizona (miniatella, in Paris Mus.); Kaslo, 
British Columbia (zanthaenobares, in USNM). 

Foop piants: Pinus spp. Larvae feed chiefly in the 
cones, sometimes in the twigs. The favored host seems 
to be Pinus ponderosa. 

DistrisutTion: Unitsep Strats: New Merico, state 
locality only; Arizona, Mohave County (July); Colo- 
rado, Glenwood Springs (Aug.), Rocky Mountain Na- 
tional Park (July); Utah, Eureka (July, Aug.); Zdaho, 
Coeur d’Alene (July); Montana, Bitterroot River 
(July), Camas (July); California, Gasquets (May), 
Pasadena, Warner Mts. (July); Oregon, Monumental 
Pass (Aug.), Silver Lake (Aug.); Washington, Pullman 
(July), Rock Lake (June, July), Seattle; Nebraska, 

300329—56——11 


Halsey (Aug.), Valentine (July, Aug.). CaNnapa: 
British Columbia; Kaslo (Aug.), Trout Creek (Ibapah 
Mts.). The Nebraska records, our most easterly, are 
from examples reared from Pinus ponderosa in refor- 
ested areas. The insect was probably introduced there 
on western nursery stock. 

The species is represented by a large series of speci- 
mens in the National Collection, a majority of them 
reared from cones in connection with the forest insect 
investigations of the U. S. Bureau of Entomology and 
Plant Quarantine. The type of auranticella in the Snow 
Collection at the University of Kansas is a pale female 
in rather poor condition. Three other similar females, 
in better condition are in the Rutger’s College Collec- 
tion. All of these are labeled simply “(New Mexico, 
Snow, Coll.” Females of auranticella are not separable 
from those of disclusa, especially the pale, yellowish 
examples, except by their locality labels. On average 
specimens the white streak along the lower margin of 
the cell of forewing is shorter in auranticella, not reach- 
ing beyond the outer angle of the cell; but this feature is 
not constant in either species. Between males there 
is never any need for confusion; for auranticella is the 
only known American species with aigrettelike maxillary 
palpi. 

A similar maxillary palpus occurs in Dioryctria laurata 
(Heinrich) from Japan (described as a Salebria in Proc. 
Ent. Soc. Washington, vol. 30, p. 61, 1928). The unique 
male type has the labial palpi erect and appressed 
close to the face and a strong scale tuft in the sinus of 
the antennal shaft. Hence the original reference to 
Salebria. It is probably a snyonym of Dvioryctria 
pryert Ragonot (Monograph, pt. 1, p. 194, 1923), 
described from a single female from the Holland Col- 
lection, now in the Carnegie Museum, Pittsburgh. I 
have never seen this specimen. 


314. Dioryctria erythropasa (Dyar) 
Figures 387, 869 
Pinipestis erythropasa Dyar, Ins. Inse. Menstr., vol. 2, p. 112, 
1914. 


Dioryciria erythropasa (Dyar) Barnes and McDunnough, Check 
list of the Lepidoptera of Boreal America, No. 5564, 1917.— 
McDunnough, Check list, No. 6128, 1939. 


Maxillary palpus of male squamous. 

Forewing with some roughened (raised) scale tufts in 
median area (one in lower fold and another in cell just 
beyond antemedial line and a slight roughening of the 
white scales of the discal spot), otherwise smooth; 
ground color red-brown of a somewhat darker, more 
rosy shade and lacking any of the orange suffusion com- 
mon to auranticella and disclusa; more or less dusted 
with white in median area, the white concentrated into 
a broad patch extending from inner margin to top of cell 
just beyond antemedial line; the transverse lines thin, 
white; antemedial line oblique, irregularly and very 
weakly dentate; subterminal line nearly vertical, slightly 
denticulate, bordered inwardly by a dark red-brown 
line; a similar dark line forms an outer border to the 
antemedial line; discal spot a slightly enlarged, lunate, 


154 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


white line on discocellular vein; outer area beyond sub- 
terminal line red-brown, terminal dots confluent, form- 
ing a more or less continuous blackish line along termen; 
cilia reddish brown. Hind wing smoky white; the veins 
darkened; a very narrow dark shade along termen; cilia 
whitish, cut by a dark subbasal line. Alar expanse, 
23-28 mm. 

Male genitalia of the majorella type but with uncus 
short, broad, its terminal margin angulate. Harpe with 
slender, digitate clasper. Vinculum narrower, more 
gradually tapered. Penis with numerous anterior 
spines, but without the usual enlarged posterior cornu- 
tus. Female genitalia with bursa copulatrix greatly 
reduced, much shorter than ductus bursae. 

Tyrr LocaLity: Chiricahua National Forest, Ariz. 
(type in USNM). 

Foop puant; Pinus chihuahuana. Larvae feeding 
in the cones. 

Distripution: Arizona, Chiricahua National For- 
est (May); Redington (Aug.). 

This species, with pygmaeella, forms a connecting link 
between the smooth-winged Dioryctria species and those 
with distinctly roughened scales formerly referred to 
Pinipestis. 

315. Dioryctria horneana (Dyar) 
Fieure 874 


Pinipestis horneana Dyar, Ins. Inse. Menstr., vol. 7, p. 43, 1919. 


Maxillary palpus of male squamous. 

Forewing bright red-brown shaded with white, the 
ground color brighter and more on the red shade than 
that of any of the foregoing species; a rather broad, 
diffused, white shade preceding the antemedial line 
and two rather large, confluent patches following it, one 
in the cell and another in lower fold; a strong broad 
oblique white shade extending from inner margin near 
subterminal line to costal beginning of that line and 
fusing with and more or less obscuring it; the scales of 
these white areas as well as those of the white discal 
spot decidedly roughened; a narrow band of appressed 
white scales along terminal margin; antemedial line 
narrow, white, nearly vertical, notched above and below 
its middle, followed on basal half by a faint, narrow, 
gray outer bordering line; subterminal white line rather 
close to outer margin, vertical with a slight bulge at 
middle; terminal black dots narrow weak, confluent. 
Hind wing yellowish white with a very faint smoky tint; 
veins but slightly darkened; a fine, pale brown line along 
termen. ‘Top of head and collar of thorax red-brown; 
remainder of thorax whitish. Alar expanse, 25-28 mm. 

Genitalia similar to those of zimmermant. 

Type tocauiry: Herradura, Pinar del Rio, Cuba 
(type in USNM). 

Foop piant: Pinus sp. 

Known only from the female type and a male from 
thesame rearing. Dyar in his original description gives 
Santiago de las Vegas as the type locality, but was evi- 
dently in error. Dr. S. C. Bruner, Chief of the De- 
partmenta Agronémica of Cuba, has given us the cor- 


rect locality. The species is easily distinguished from 
others of the zimmermani complex by the bright reddish 
ground color of its forewings and from the other red- 
winged American species by its strongly tufted fore- 
wings. 

The labial palpi of the male are more closely appressed. 
to the face than those of most Dioryctria species. 


316. Dioryctria pygmaeella Ragonot 
Ficures 388, 877 


Dioryciria pygmaeella Ragonot, N. Amer. Phycitidae, p. 5, 1887; 
Monograph, pt. 1, p. 192, 1893. 

Pinipestis pygmaeella (Ragonot) Hulst, Phycitidae of N. Amer., 
p. 136, 1890.—Forbes, Cornell Mem. 68, p. 620, 1923.— 
McDunnough, Check list, No. 6136, 1939. 

Maxillary palpus of male squamous. 

Forewing smooth except for a slight roughening of 
the white scales of the transverse lines and discal spot; 
purplish gray with irregular patches of dull dark red 
and a more or less extended white dusting; the red 
patches consisting of a rather broad band preceding the 
antemedial line, a similar band from middle of inner 
margin to cell and a rather broad shade outwardly 
bordering the subterminal line; the white dusting con- 
centrated over the median areas not occupied by the 
red median band, in subbasal area just behind the red 
band, and along termen following the red subterminal 
shade; transverse lines thin, whitish, rather faint (under 
magnification, the scales silvery and somewhat rough- 
ened); antemedial line oblique, notched at vein 1b, 
bordered outwardly by a black line which expands at 
costa into a black patch; subterminal line sinuate- 
angulate with a broad, black, inner, bordering line; 
terminal dots fused into a fine black line along termen; 
white discal spot an oblique lunule; a fine black line 
along the lower half of the inner margin of the red band 
preceding antemedial line (but not a raised-scale ridge 
as stated by Ragonot); also some dusting of black scales 
on the red of extreme basal area. Hind wing pale smoky 
gray; the veins darkened and a narrow dark shade 
along termen. Alar expanse, 15-21 mm. 

Male genitalia with a cluster of long strong posterior 
spines and one or two weak anterior spines on penis. 
Female genitalia with the spine clusters in bursa con- 
sisting of narrow, straight bands of slender spines; 
ductus bursae sclerotized from junction with bursa 
almost to genital opening. 

Type Locatity: Florida (type in Paris Mus.). 

Foop pLant: Taxodium distichum. Larvae in the 
cones. ‘This record from reared series from Maryland 
(Heinrich, 1920) and Virginia (Busck, 1927) in the U.S. 
National Museum. . 

Distrisution: Florida, Winter Park (July); Virginia, 
Cape Henry (Aug.); Maryland, Pokomoke (Aug., Sept.). 
Probably over the range of its host plant. 

‘A distinct, easily recognized species, intermediate be- 
tween the smooth-winged species and. mbGEe with definite 
meee Seale TCE ! 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 155 


317. Dioryctria zimmermani (Grote) 
Ficures 389, 875, 878 


Nephopteryx (Dioryctria) zimmermani Grote, Canadian Ent., 
vol. 9, p. 163, 1877. 

Nephopteryx (Pinipestis) zimmermani (Grote), Canadian Ent., 
vol. 10, p. 19, 1878.—Packard, U. S. Dep. Agr. Fifth Rep. 
Ent. Comm., p. 73, 1890. 

Pinipestis zimmermani (Grote), Bull. U. S. Geol. Geogr. Surv. 
Terr., vol. 4, p. 699, 1878; op. cit., vol. 6, p. 589, 1882; N. 
Amer. Ent., vol. 1, p. 11, pl. 2, fig. 10, 1879. 

Nephopteryx zimmermani (Grote) Kellicott, Canadian Ent., vol. 
11, p. 114, 1879; Ent. Amer., vol. 1, p. 173, 1885. 

Dioryctria zimmermanni (Grote) Ragonot, Ent. Amer., vol. 5, 
p. 114, 1889; Monograph, pt. 1, p. 190, 1893 (emended 
spelling of specific name).—Forbes, Cornell Mem. 68, p. 620, 
1923. 


Pinipestis zimmermanni (Grote) Hulst, Phycitidae of N. Amer., 
p. 137, 1890.—McDunnough, Check list, No. 6133, 1939. 

Salebria delectella Hulst, Canadian Ent., vol. 27, p. 57, 1895.— 
Ragonot, Monograph, pt. 2, p. 550, 1901. (New synonymy.) 

Dioryctria delectella (Hulst) Dyar, Proc. Ent. Soc. Washington, 
vol. 6, p. 227, 1904. 

Retinia austriana Cosens, Canadian Ent., vol. 38, p. 362, 1906.— 
Busck, Journ. New York Ent. Soc., vol. 15, p. 236, 1907. 

Pinipestis delectella (Hulst) Barnes and McDunnough, Check 
list of the Lepidoptera of Boreal America, No. 5570, 1916.— 
MeDunnough, Check list, No. 6134, 1939. 

Dioryctria ponderosae Heinrich (not Dyar), in Keen, U. S. Dep. 
Agr. Misc. Pub. 273, p. 38, 1938. 

Dioryctria zimmermani (Grote) Craighead, U. 8S. Dep. Agr. 
Misc. Publ. 657, p. 452, 1950. 


Maxillary palpus of male squamous. 


Forewing with a ridge of raised (roughened) scales 
preceding and one following the antemedial line, some 
rough scaling of the discal spots and on some specimens 
in the outer median area above inner margin and im- 
mediately before subterminal line, the raised scaling 
somewhat variable and nowhere reaching to costa, 
easily and frequently flattened in the spreading and 
setting of specimens; color variable, blackish gray with 
a rather broad, but faint, whitish dusting before the 
subterminal line and, on some specimens but to a lesser 
extent, immediately following the antemedial line and 
narrowly and faintly along the inner margin of the 
subbasal scale ridge; basal and terminal areas normally 
shaded with red, the extent and tint of the shading 
extremely variable and on some specimens almost 
obliterated or confined to dull patches of an oblivace- 
ous hue; when strongly accented, extended over base 
and onto the collar of the prothorax, usually most dis- 
tinct between subbasal scale ridge and antemedial line; 
the raised-scale ridges themselves, black; transverse 
lines dull white, always distinguishable but sometimes 
faint, bordered inwardly and outwardly by black lines 
which broaden into dark wedges on costa; white discal 
spot usually distinct; a black line along terminal margin. 
Hind wing white more or less shaded with smoky fuscous 
along costa and termen, less so on male than female; a 
fine dark line along termen; veins only faintly darkened. 
Alar expanse, 25-33 mm. ' 

Male genitalia with uncus but slightly longer than 
broad, the lateral margins slightly concave; terminal 
margin rounded; when flattened in preparation; as in 
figure 390a of cambiicola. .Harpe with costa broadly 


sclerotized and terminating at apex in a long curved 
pointed hook, a short spine from its lower outer angle; 
clasper digitate; cucullus narrow, pointed at apex. 
Penis with posterior spine, long, strong, straight, evenly 
tapering to a sharp point. Vinculum stout, consider- 
ably longer than broad, evenly tapering to roundly 
angulate terminal margin. 

In the female genitalia the variation in the spining 
of bursa shown in the figures is merely individual and 
is equaled or exceeded in any series of eastern or western 
specimens. Ductus bursae much longer than bursa, 
sclerotized for its entire length except for a short dis- 
tance from genital opening, the sclerotization ribbon- 
like, broadening and bent towards bursa, longitudinally 
ribbed on caudal half and terminating caudally in a 
produced, bluntly pointed or acutely rounded central 
projection. Bursa proportionally small; the spine 
clusters closely grouped at its posterior half; the en- 
larged lobe giving off the ductus seminalis appreciably 
thickened. 

Typ Locauities: Buffalo, N. Y. (zimmermani, in 
BM; paratype, o, in USNM); Colorado (delectella, in 
AMNH, ex Rutgers); Toronto, Ontario, Canada (aus- 
triana, in Royal Ontario Mus.). 


Foop prant: Pinus spp. Most if not all species of 
pine in this country are attacked. The spruce records 
given by Packard (1895) have never been verified. 
They were probably based upon misidentified larvae. 
I doubt very much that zimmermanz feeds on anything 
but pine. The larvae bore into the cambium of the 
trunk, branches, and twigs, causing considerable dam- 
age to the new growth of older trees and sometimes even 
killing younger trees (8 inches or less in diameter) by 
completely girdling their boles. The place of attack is 
usually indicated by a resinous mass of exuded pitch 
mixed with frass and larval exuviae. 

Distripution: Unirep States: New York, Buffalo, 
Coram (Long Island, Aug.), Warrensburg (Aug.); Con- 
necticut, Woodstock (Aug.); Rhode Island, Washington 
County (June); Massachusetts, Dover (Aug.), Martha’s 
Vineyard (Aug.); New Hampshire, Hampton (Sept.); 
North Carolina, Tryon (Aug.); Ohio, Akron (July, Aug.), 
Lake County (July, Aug.), Mentor (June), Scioto 
County (July) ; Ziinois, Oregon (July, Aug.) ; Nebraska, 
Ainsworth (Aug.), Halsey (May, June, July, Aug.), 
Meadville (Aug.), Norden (Aug.), Wyoming, Wyoming 
National Forest (June, July) ; Montana, Banner (July), 
Missoula (July); Colorado, only the state locality (fe- 
male cotypes of delectella, Bruce, collector) ; New Mezico, 
Taos Junction (July); Arizona, Santa Catalina Mts. 
(Bear Canyon, July), White Mts. (Aug.); California, 
Placerville, San Mateo (June), Ventura; Oregon, Butte 
Falls, Coletin; Washington, Friday Harbor (Aug.), 
Rock Lake (July). Canapa: Ontario, Toronto. 

The foregoing records are from specimens before me, 
most of them reared. The range of the species probably 
extends over the entire northern areas of the United 
States wherever its hosts occur and presumably over 
a considerable area in southern Canada. 

Hulst’s delectella was described from, Colorado females 


156 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


which, except for somewhat larger expanse (82 mm.) 
than average eastern specimens, are typical zimmermani 
in coloration and genitalia. The unfortunate reference 
to ponderosae in the Keen paper (1938) was due to my 
misidentification of reared specimens from the reforesta- 
tion areas of Nebraska. These came to us in good 
series but were spread and the raised scales on the fore- 
wings had been flattened, and on most of the examples 
(as also on some eastern specimens) the characteristic 
red shading on basal area of forewing was lacking. 
Superficially they looked like ponderosae; but had the 
typical zimmermani genitalia, and unrubbed examples 
clearly showed the raised scaling. 

Economically zimmermani is our most important 
Dioryctria. In this country it does more serious dam- 
age, especially to young trees, in both the East and the 
West, than abietella. As far as I know it is strictly a 
bark borer and does not attack the cones. The most 
complete and accurate account of the life history is that 
in the Craighead (1950) paper. The life history of the 
insect in Nebraska is also treated in the Keen (1938) 
paper under “‘ponderosae.” 


318. Dioryctria cambiicola (Dyar) 
Figures 390, 876 
Pinipestis cambticola Dyar, Ins. Insc. Menstr., vol. 2, p. 2, 
1914.—MecDunnough, Check list, No. 6137, 1939. 

Maxillary palpus of male squamous. 

Forewing as in zimmermani except: Ground color of 
basal, submedial and terminal areas a dark, dull, reddish 
brown (somewhat paler on Arizona specimens); the 
black scaling greatly reduced; the white scaling follow- 
ing antemedial and preceding subterminal lines and 
bordering terminal margin; the transverse lines and 
the discal mark, dull silvery. Hind wing smoky fuscous; 
the veins darkened and a dark line along termen. Alar 
expanse, 28-32 mm. 

Genitalia, male and female, show no specific differ- 
ences from those of zimmermant. The extent of the 
bend in the female ductus bursae shown by the figures 
is purely individual and can be easily accented or 
diminished in preparing the slides. 

TypE Locatity: Flathead Reservation, Mont. (type 
in USNM). 

Foop pLants: Pinus ponderosae, P. scopulorum, P. 
coulteri. The larvae are cambium borers in new growth, 
causing pitch exudations like those of zimmermant. 

DistriBution: Montana, Flathead Reservation (July, 
Aug.); Missoula (July); Colorado, Boulder (Sept.), 
Palmer Park (July, Aug.); New Mexico, Las Vegas; 
Arizona, Flagstaff, White Mts. (Aug.); California, 
Julian (Sept.). 

The species is doubtfully distinct from zimmermani 
except as a possible race. It differs chiefly in its dark 
hind wings. It apparently has a limited distribution 
in our Western States. In his original description Dyar 
mentions an eastern specimen (presumably from Wash- 
ington, D. C.) reared from a cone of Pinus taeda, Aug. 
14, 1882. I have also before me a similar female from 
Cape Henry, reared June 9, 1927. I suspect that both 


these examples may be hybrids of zimmermani and 
amatella. 
319. Dioryctria amatella (Hulst) 
Fiaure 879 


Nephopteryz amatella Hulst, Ent. Amer., vol. 3, p. 131, 1887. 

Dioryctria amatella (Hulst), Ragonot, Ent. Amer., vol. 5, p. 114, 
1889; Monograph, pt. 1, p. 191, 1893.—Craighead, U. S. 
Dep. Agr. Mise. Publ. 657, 1950. 

Pinipestis amatella (Hulst), Phycitidae of N. Amer., p. 136, 
1890.—McDunnough, Check list, No. 6135, 1939. 

Maxillary palpus of male squamous. 

Forewing with raised scales as in zimmermani but the 
scale ridges somewhat narrower; ground color a dark 
wood brown; transverse lines, discal spot, a thin trans- 
verse band preceding the subbasal ridge, a blotch fol- 
lowing the antemedial line, a similar shade near inner 
margin of subterminal line on lower half of wing, and 
a thin zigzag pale shade just within terminal margin, 
silvery white and strongly contrasted against the 
ground color; black scaling limited to the subbasal and 
submedian scale ridges, the thin borders of the trans- 
verse lines, a spot at extreme base, and the thin terminal 
line. Hind wing smoky with a somewhat glossy pale 
brownish tint; the veins slightly darkened and a thin 
dark line along terminal margin. Alar expanse, 27— 
32 mm. 

Genitalia essentially like those of zimmermani. 

TypE Locauity: Florida (type in AMNH, ex Rutgers). 

Foop pLants: Pinus spp. Larvae feeding in cones 
and terminals, more often in the cones than terminals and 
apparently favoring diseased cones. 

Disrrisution: Florida, Alton (June), Camp Pinchot 
(June), Fort Mead (May), Lake City (May, June), 
Monticello (Sept.), Orlando (June), ‘Southern Florida’’ 
(June, July), Starke (May); ZLowiszana, New Orleans 
(Sept., Nov.), Woodworth (June) ; Texas, Conroe (May); 
Mississippi, Hattiesburg (June), Picayune (May); 
Maryland, Baltimore (Aug.); District of Columbia, 
Washington (July). 

Close to but apparently distinct from zimmermani 
and cambiicola, distinguished from both by the strong 
contrast of its white markings. Generally distributed 
in the Gulf States where its abundance, especially in 
Florida and Louisiana, makes it something of a pest. A 
brief account of what is known of its life history is given 
in the Craighead (1950) paper. 


320. Dioryctria albovittella (Hulst) 
Fiaure 880 
Pinipestis albovittella Hulst, Phycitidae of N. Amer. p. 138, 
1900.—Barnes and McDunnough, Contributions, vol. 4, p. 
174, 1918.—MeDunnough, Check list, No. 6138, 1939. 
Dioryciria albovittella Ragonot, Monograph, pt. 1, p. 193, 1893. 

Maxillary palpus of male squamous. 

Forewing with a narrow black subbasal scale ridge, 
little if any raised scaling otherwise; forewing gray 
densely dusted with white, making the extreme basal, 
median, and terminal areas a pale ash color; transverse 
lines and discal mark well contrasted, white; antemedial 
line nearly vertical, slightly notched above and below 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 157 


middle, bordered outwardly by a thin black line and 
inwardly by a broad dark shade which includes the 
black raised-scale ridge; a white blotch on lower half of 
wing just beyond the antemedial line; subterminal line 
well defined, median section broadly triangulate, bor- 
dered inwardly by a blackish line and outwardly by a 
narrow dark shade; discal marking a white spot covering 
discocellular vein; a row of confluent black dots along 
termen. Hind wing white with a faint ocherous tint; a 
fuscous shade at apex and, narrowly, along termen. 
Alar expanse, 23-30 mm. 

Male genitalia like those of zimmermani. Female 
genitalia similar to those of cambiicola, differing only in 
trifling details of the spining in the bursa, not exhibiting 
any consistent differences of a specific character. 

Typr tocatiry: Colorado (type in AMNH, ex 
Rutgers). 

Foop piant: Cones of Pinus monophylla. This rec- 
ord from reared Nevada specimens received from the 
Forest Insect Division of the U. S. Bureau of Entomol- 
ogy and Plant Quarantine under Hopkins U. S. No. 
32009. 

Distripution: New Mexico, Jemez Mts. (Aug.), 
Jemez Springs (Aug.); Arizona, Mohave County (July, 
Aug.) ; Colorado; Utah, Dividend (Aug.), Eureka (Sept.), 
Ibapah Mts. (Trout Creek, July); Nevada, Topaz Lake 
(Aug.). 

Another close relative of zimmermani and cambiicola, 
distinguished chiefly by its much paler forewings. In 
his original description Hulst gives Hot Springs, N. 
Mex., as the type locality; but this, as pointed out by 
Barnes and McDunnough (1918) is evidentally a lapsus. 
The type is a male, labeled ‘‘Colo. Bruce.” 


321. Dioryctria gulosella (Hulst), new combination 
Figure 392 


Acrobasis gulosella Hulst, Phycitidae of N. Amer., p. 126, 1890.— 
Ragonot, Monograph, pt. 1, p. 109, 1893; pt. 2, p. 520, pl. 
50, fig. 11, 1901. 

Pinipestis gulosella (Hulst) Barnes and McDunnough, Check 
list of the Lepidoptera of Boreal America, No. 5575, 1916.— 
MceDunnough, Check list, No. 6139, 1939. 

Maxillary palpus of male squamous. 

Forewing with a subbasal scale ridge and small patch 
of raised scales in median area; dark gray with a fine, 
sparse, white dusting, making the general color a dark 
ash gray; transverse lines whitish gray, distinct; ante- 
medial line twice notched, edged outwardly by a thin 
black line, preceded by an obscure pale patch on lower 
half of wing, this followed on its inner margin by a black 
scale ridge continued as a thin black line to costa; sub- 
terminal line outwardly angled at middle, bordered 
inwardly by a black line and outwardly by a narrow 
dark shade; discal spot white; a narrow black line along 
termen. Hind wing white, smoky at apex and some- 
what along termen (especially on females); the veins 
more or less darkened; a fine brown line along terminal 
margin. Alar expanse, 21-27 mm. 

Genitalia similar to those of bawmhoferi except for the 
shape of the uncus. The latter has a more broadly 


rounded terminal margin on gulosella (compare figs. 
391 and 392). 

Type tocauity: Hot Springs, N. Mex. (type in 
AMNH, ex Rutgers). 

Foop prant: Not definitely known, but undoubtedly 
pine. 

Disrrisution: New Mexico, Hot Springs (Aug.); 
Colorado, Glenwood Springs (July, Aug.), also one 
female with only the state locality (Bruce, collector) 
and bearing a Hulst “type” label. The type from New 
Mexico is also a female. The Colorado specimens (2 
of and 4 9) are all in the National Collection. They 
are a perfect match for Ragonot’s figure. 


322. Dioryctria baumhoferi, new species 
Figures 391, 881 


Maxillary palpus of male squamous. 

Forewing with raised scaling as on typical zimmer- 
mani; blackish gray with the usual black markings indi- 
cated by a more intense darkening of the ground color; 
the transverse lines faint but distinguishable, gray; 
antemedial line bordered inwardly by a broad black 
band including the subbasal scale ridge; a similar 
blackish, transverse shade across the middle of the 
wing; subterminal line outwardly angulate at middle, 
bordered inwardly by a black line and outwardly by a 
rather broad black shade extended into streaks on some 
of the veins; a black line along terminal margin; discal 
spot whitish gray, sometimes very faint. Hind wing 
smoky gray; the veins darkened and the smoky shade 
intensified along termen. Alar expanse, 25-28 mm. 

Male genitalia having uncus triangulate with nar- 
rowly rounded apical margin. Female genitalia of the 
zimmermani type but with ductus bursae shorter in 
proportion to length of bursa and somewhat broader. 

Typr LocaLity: Prescott, Ariz. (type in USNM, 
61355). 

Foop piant: Pinus ponderosa. 
new growth. 

Described from male type and one male and nine 
female paratypes from the type locality, reared under 
Hopkins Nos. 9932C and 18506, June 5, 6, and 11, 
1928, by the late L. G. Baumhofer of the Forest Insect 
Division of the U. S. Bureau of Entomology and Plant 
Quarantine. Baumhofer’s extensive rearings and field 
studies, especially in the Nebraska National Forest, 
have contributed what knowledge we have of the 
biology of the Dioryctria species in that area. 

The new species is close to gulosella Hulst, from which 
it is distinguished by the much darker color of its fore 
and hind wings, differently shaped male uncus, and the 
somewhat stouter spining of the male penis. It may 
prove to be a local race of guloselia but is at least as 
distinct from it as cambiicola is from zimmermant. 


Larvae feeding in 


323. Dioryctria subtracta, new species 
FiaureEs 393, 882 


Maxillary palpus of male squamous. 
Forewing with a subbasal ridge of raised scales and 


158 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


a small tuft in lower fold just beyond antemedial line; 
dark gray finely peppered with white making the ground 
color a dark ash gray; the subbasal scale ridge, outer 
border of antemedial and inner border of subterminal 
lines, and the small raised patch following antemedial 
line, black strongly contrasted against the ground 
color, the outer border of antemedial line somewhat 
fainter than the other black markings; subbasal scale 
ridge narrow, reaching almost to costa; between it and 
antemedial line a broad pale band; antemedial line 
obscure, indicated chiefly by its blackish outer bordering 
line which is almost vertical and parallel with the scale 
ridge; subterminal line also faint, pale gray, outwardly 
angled at middle, bordered inwardly by a narrow black 
line; discal spot obscure, a narrow grayish white line on 
discocellular vein; a fine, strongly contrasted, black line 
along terminal margin. Hind wing white, on female a 
faint smoky tint towards apex; the outer parts of the 
veins faintly darkened and a narrow brown line along 
terminal margin. Alar expanse, 23-25 mm. 

Male genitalia with uncus rather short in proportion 
to width; its terminal margin broadly rounded. Harpe 
narrow and rather short. Penis armed with two 
groups of subterminal spines and a single, straight, 
slender, rather short posterior spine; one of the anterior 
groups consisting of a line of short spines along lateral 
margin of penis near its apex. 

TypH Locatity: Fort Wingate, N. Mex. (type in 
USNM, 61356). 

Foop pirant: Unknown. 

Described from male type and one female paratype 
from the type locality (July), and one female from Glen- 
wood Springs, Colo. (Sept.) which McDunnough had 
identified as gulosella Hulst. They are superficially 
similar to the type of that species except for the stronger 
contrast of the black markings on forewing; but are 
easily separated by their radically different genitalia. 
The expanded (bulbous) shape of the caudal end of the 
female ductus bursae is found in only one other Ameri- 
can species of Dioryctria (clarioralis) and there in a 
lesser degree. 


324, Dioryctria clarioralis (Walker) 
Fiaures 394, 883 


Nephopteryz clarioralis Walker, List, vol. 27, p. 54, 1863. 

Dioryctria clarioralis (Walker) Ragonot, Ent. Amer., vol. 5, p. 
114, 1889; Monograph, pt. 1, p. 193, 1893.—Hulst, Phy- 
citidae of N. Amer., p. 136, 1890—Forbes, Cornell Mem. 
68, p. 620, 1923.—MeDunnough, Check list, No. 6125, 1939. 

Ulophora brunneella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 106, 1904.—Barnes and McDunnough, Contributions, 
vol. 3, p. 195, 1916 (make synonym of clarioralis). 

Maxillary palpus of male squamous. 

Forewing with a weak subbasal ridge of raised scales 
(the species has always been included in the smooth- 
winged Dioryctria group, but unrubbed and unpressed 
examples always show some traces of a raised subbasal 
ridge); ground color grayish brown with black patches 
and more or less white shading in the median and ter- 
minal areas; a broad blackish band preceding the ante- 
medial line, paling towards inner margin, bordered in- 


wardly by the black raised-scale ridge and outwardly by 
the thin, black inner border of the antemedial line; the 
latter thin, oblique, sometimes weakly notched below 
costa and more rarely at lower fold, white without (or 
with only an occasional trace, near inner margin) the 
normal black outer bordering line; a more or less ex- 
tended black smudge in cell, sometimes extended as 
far as the black inner border of subterminal line and 
usually completely obliterating any trace of a white 
discal spot; some white streaking on lower vein of cell; 
subterminal line distinct, sharply indented between 
costa and vein 5, thence vertical and straight to inner 
margin, whitish with a thin black inner border; a fine 


_line of confluent black dots along terminal margin. 


Hind wing smoky gray or brownish; the veins more or 
less darkened; a fine dark line along termen. Alar 
expanse, 22-29 mm. 

Male genitalia with uncus triangulate, appreciably 
longer than greatest width, evenly tapering to very 
narrowly rounded apex. Female with ductus bursae 
broadened near genital opening but less so than in 
subtracta; bursa much larger and more heavily spined 
than that of subtracta, at least as long as ductus bursae. 

TYPE LOCALITIES: “United States” (clarioralis, in 
BM); Tryon, N. C. (brunneella, in USNM). 

Foop piant: Pinus palustris. This record from a 
specimen received from L. A. Hetrick reared from larva 
feeding in the cone. 

Distrisution: Florida, Dunedin (Mar.), Fort Myers 
(Apr.), Miami; North Carolina, Tryon (May, June); 
Massachusetts, Martha’s Vineyard (June, July). 

Another intermediate between the smooth-winged 
and rough-scaled species, easily distinguished by its 
genitalia and wing pattern. 


78. Genus Oryctometopia Ragonot 


Oryctometopia Ragonot, Nouv. Gen., p. 11, 1888; Monograph, 
pt. 1, p. 153, 1893.—Janse, Journ. Ent. Soc. South Africa, 
vol. 4, p. 156, 1941. (Type of genus: Oryctometopia fossula- 
tella Ragonot.) 

Tongue well developed. Antenna of male with a 
slight sinus and scale tuft in base of shaft; pubescent. 
Labial palpus obliquely upturned, reaching to or a trifle 
above vertex; third segment about two-thirds the length 
of second, bluntly pointed, more or less deflected for- 
ward. Maxillary palpi of both sexes broadly squamous, 
the scales forming a flat cover over the face. Forewing 
smooth; 11 veins; vein 2 from before but near lower 
outer angle of cell; 3 from the angle, equidistant at base 
from 2 and 4, parallel with 2 from just beyond base for 
its remaining length; 4 and 5 connate or very shortly 
stalked; 6 from below upper angle of cell, straight; 8 
and 9 stalked for about two-thirds the length of 8; 10 
from the cell, approximate to stalk of 8-9 at base; male 
with costal fold containing a row of coarse scales. Hind 
wing with vein 2 from before lower outer angle of cell; 
3 from the angle; 4 and 5 stalked for about half their 
lengths; 7 and 8 closely approximate beyond cell at least 
for half their lengths; all veins long; cell about one- 
fourth the length of wing; discocellular vein slightly 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 159 


curved, not extended at lower angle. Highth abdom- 
inal segment of male with a pair of long, slender, ventro- 
lateral hair tufts. 

Male genitalia with uncus subtriangulate. Apical 
process of gnathos a short, stout, hook. Transtilla ab- 
sent. Harpe with one or more short stout thornlike 
spines projecting from lower margin of sacculus. Aedea- 
gus slender, rather long, sinuate. Penis armed with a 
single thornlike cornutus. Vinculum stout, longer 
than broad, subtriangulate, narrowed from middle to 
terminal margin. 


Female genitalia with signum consisting of a single 
round curved plate, densely armed with long, stiff 
spines and covering the ventral and lateral caudal half 
of the bursa copulatrix; bursa otherwise membranous; 
a narrow sclerotized collar about ductus bursae at its 
junction with bursa, the ductus bursae otherwise un- 
sclerotized; genital opening simple; ductus seminalis 
from bursa near junction of bursa and ductus bursae. 

The genus, so far as we know, is confined to the New 
World and contains but one tropical American species. 


325. Oryctometopia fossulatella Ragonot 
Ficures 53, 395, 900 


Oryctomeiopia fossulatella Ragonot, Nouv. Gen. p. 11, 1888; Mon- 
ograph, pt. 1, p. 153, 1893. 

Phycita moeschleri Ragonot, Nouv. Gen., p. 12, 1888; Monograph, 
pt. 1, p. 182, 1893—Méschler, Die Lepidopteren-Fauna 
von Portorico, p. 328, 1890. (New synonymy.) 

Forewing gray to pale brownish gray; the transverse 
lines obscure but usually distinguishable; antemedial 
line nearly vertical, far out from base, on well marked 
specimens irregularly serrate and bordered outwardly 
by a thin dark line which is more or less broken, pre- 
ceded by an obscure reddish olivaceous (or reddish 
brown) patch on inner margin, the latter bordered 
inwardly by same blackish scaling; a similar obscure 
reddish brown patch over lower fold just before the sub- 
terminal line; subterminal line more distinct than ante- 
medial, sinuate (outwardly bulged at middle), dull 
whitish gray, bordered inwardly and outwardly by 
narrow dark lines; discal dots, when distinguishable, 
small, separated, blackish; a row of obscure blackish 
dots along termen; on most specimens a dull whitish 
patch over middle of inner margin. Hind wing trans- 
lucent white; the veins not appreciably darkened; a con- 
trasting dark shade along costa and narrowly along ter- 
men; on males, a fine dark subbasal line through the 
white cilia. Alar expanse, 15-20 mm. 


Genitalia as given for the genus. The thornlike 
spines on sacculus of the male harpe are individually 
variable, consisting of one stout, hooked spine and two 
or more slenderer spines, their number and size varying 
not only in individuals but on opposite harpes of the 
same specimen. Cornutus of penis with a flattened 
platelike base. 

Type LocauitiEs: “Irazu’’? [Mount Irazi], Costa 


Rica (fossulatella, in BM); Puerto Rico (moeschleri, in 
Paris Mus.). 


Foop piant: Bauhinia mexicana. Larvae in the 
pods. This record from Brownsville, Tex., specimens 
reared by the Division of Foreign Plant Quarantine of 
the U. S. Bureau of Entomology and Plant Quarantine. 

Distrisution: Unitep States: Texas, Brownsville 
(June). M*sxico: Chiapas (May), Jalapa, Oaxaca, Te- 
huacin (Apr.). GuatTeMaua: Cayuga (May, June, 
Sept.), Chejel (June, July, Aug.), Purulhé (July), Vol- 
cin Santa Maria (May, June, July). Costa Rica: 
Esperanza (May), Mount Irazi. Panami: Cabima 
(May), Corazal (Apr.), La Chorrera (May), Porto 
Bello (Oct.). VenrEzunLA: Aragus (Rancho Grande, 
May). Braz: Rio de Janeiro (‘10-1-31”’). Puzrtro 
Rico: Coamo Springs (Apr.), Covado (May), Puerto 
Real (Vieques Isl., Apr. ), San German (Apr.). Vira@in 
Isuanps: Kingshill (St. Croix, June). 

The species is easily distinguished by its genitalia; 
but in color and maculation is variable, as the foregoing 
description indicates. Such variability is purely indi- 
vidual and has no racial or local significance whatsoever. 
Ragonot described fossulatella from a large (20 mm.) 
male and his moeschleri from a small (17 mm.) female 
color variant, which accounts for his two names and 
their placement in different genera. 


Genera 79-81: Sarata to Lipographis 


[Venational division B. Forewing smooth; veins 4 and 5 sepa- 
rated at base. Hind wing with veins 4 and 5 stalked; cell usually 
short, about one-third the length of wing (longer in Lipographis, 
about one-half). Labial palpus porrect, broadly scaled, beak- 
like. Male genitalia with transtilla incomplete or absent; harpe 
with costa partially sclerotized, not produced, clasper reduced or 
absent, otherwise simple. Female genitalia with bursa smooth 
or scobinate, without signum; ductus seminalis from bursa.] 


79. Genus Sarata Ragonot 


Sarata Ragonot, N. Amer. Phycitidae, p. 11, 1887; Monograph, 
pt. 1, p. 614, 1893.—Hulst, Phycitidae of N. Amer. p. 168, 
1890. (Type of genus: Sarata dnopherella Ragonot.) 

Tongue well developed. Antenna pubescent; shaft 
of male cylindrical, slightly swollen at base and smooth- 
ly scaled or with a very slight ridge of roughened scales 
along a few of the basal segments. Labial palpus 
porrect, beaklike; second segment oblique, laterally 
flattened, broadly scaled; third segment deflected for- 
ward, about the length of second (sometimes a trifle 
shorter or longer), bluntly acuminate. Maxillary pal- 
pus minute, filiform. Forewing smooth; 11 veins; 
vein 2 from well before lower outer angle of cell; 3 from 
the angle; 4 and 5 separated at base; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for from one-half 
to two-thirds their lengths; 10 from the cell, approxi- 
mate to the stalk of 8-9 for a considerable distance from 

base; male without costal fold. Hind wing with vein 2 

from well before lower outer angle of cell; 3 from the 

angle or separated from it by a very short spur; 4 and 

5 stalked for two-thirds their lengths; 7 and 8 closely 

approximate beyond cell for nearly half their lengths; 

cell about one-third the length of wing; discocellular 


160 


vein curved, outwardly produced at lower angle of cell. 
Eighth abdominal segment of male with a pair of weak, 
ventrolateral hair tufts. 

Male genitalia with uncus triangulate, its apex 
rounded. Apical process of gnathos terminating in an 
elongate hook, moderately long and bladelike (except in 
incanella where it is short, and digitate from an enlarged 
base); lateral arms of gnathos broad and stout. Trans- 
tilla incomplete; its lateral elements well sclerotized and 
short or moderately long and slender, their apices 
pointed. Harpe, elongate, slender; apex of cucullus 
rounded; clasper present as a very short, blunt, wartlike 
projection from near middle of basal margin of cucullus 
(except in incanella where it is broader and more scoop- 
like), the size and shape of the wartlike clasper indi- 
vidually variable; costa not produced, strongly sclero- 
tized only on basal half. Anellus a shallow, broadly 
U-shaped shield. Aedeagus long, straight, not appre- 
ciably tapering or expanded towards apex, stout to 
moderately slender (incanella); penis (except in incan- 
ella) armed with a single, long, stout, spikelike cornutus, 
rarely a second elongate slenderer spine (on penis of 
incanella the single cornutus is @ short, stout thorn sit- 
uated near apex). Vinculum stout, as long as or but a 
trifle longer than broad; terminal margin broad. 

Female genitalia with bursa strongly scobinate over 
much of inner surface and more or less thickened 
(cartilaginous) at or near anterior end; ductus bursae 
simple (unsclerotized and unspined throughout), nor- 
mally distinctly shorter than bursa; ductus seminalis 
from bursa near its middle well forward of junction of 
bursa and ductus bursae). 


The species here referred to Sarata form a homogene- 
ous group. Some of them on the basis of a smooth 
male antennal shaft have hitherto been listed under the 
Old World genus Megasis; but none agrees with the 
type of the latter (rippertella (Zeller), fig. 428) on 
genitalic characters or the stalking of veins 4—5 of hind 
wing, which is always shorter (and frequently incom- 
plete, a mere approximation or anastomosis of the basal 
half of the veins in rippertella). The difference between 
a smooth antennal shaft and one with some roughened 
scales towards base is very slight, and should have no 
weight against the uniformity of the genitalic and vena- 
tional characters of Sarata otherwise. Indeed the 
species of Sarata are much closer to those of Lipographis 
than to the type of Megasis. 

None of our species has been reared and nothing is 
known of the food plants of any of them, so that any 
association of females with males is purely speculative. 
Such associations as have been made are open to grave 
suspicion. The females differ from the males not only 
in size but also in pattern and color, and within any 
given species of females the color varies more between 
individuals than it does between the species themselves. 
For this reason I have treated the males and females 
separately, giving to the latter new temporary names 
which can go into synonymy when the sexes are properly 
associated. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


The following three species hitherto listed in Megasis 
or Sarata must be referred elsewhere: 

Sarata rhoiella Dyar becomes the type of a new genus 
(Philodema). 

Sarata umbrella Dyar goes to Lipographis. 

Megasis indianella Dyar is an anerastiine and a 
synonym (see p. 315) of Ragonotia olivella (Hulst). 


Genus Sarata (males), Species 326-330: 
S. edwardsialis to S. incanella 


[Antennal shaft at base smooth scaled.] 


326. Sarata edwardsialis (Hulst), new combination 


Figure 396 


Megaphysis edwardsialis Hulst, Trans. Amer. Ent. Soc., vol. 13, 
p. 163, 1886. 

Megasis polyphemella Ragonot, N. Amer. Phycitidae, p. 10, 1887; 
Monograph, pt. 1, p. 545, 1893. 

Megasis edwardsialis (Hulst), Ent. Amer., vol. 5, p. 156, 1889; 
Phycitidae of N. Amer., p. 165, 1890.—McDunnough, Check 
list, No. 6259, 1939. 

Forewing pale grayish brown more or less smudged 
with darker gray; blackish streaks on several of the 
veins, especially marked on vein 1b, the lower vein of 
cell, and the veins immediately preceding and following 
the subterminal line; the latter faintly indicated; ante- 
medial line obsolete; discal spots poorly defined, often 
obliterated, where distinguishable, separated; a row of 
narrow black dots along termen, more or less accented. 
Hind wing a little lighter grayish brown than ground 
color of forewing, the veins not appreciably darkened, 
a slightly darker line along termen; cilia paler, a dull 
white with a faint fuscous tint. Alar expanse, 35-46 
mm. 

Male genitalia with uncus evenly tapering to narrowly 
rounded apex. Apical process of gnathos about half as 
long as uncus. Elements of transtilla very short. 
Aedeagus long, stout; penis armed with a single, straight 
cornutus, nearly as long as aedeagus, also a small sup- 
plemental sclerotized patch. Terminal margin of vin- 
culum concave. 

TyprE LocaLitiEs: Nevada (edwardsialis, in AMNH, 
ex Rutgers) ; California (polyphemella, in Paris Mus.). 

Foop prant: Unknown. 

Distrisution: Colorado, Boulder (Mar.), Glenwood 
Springs (Mar., Apr.), Salida; Utah, Dividend (Mar.), 
Eureka (Apr., May); Nevada; California, ““Middle Cali- 
fornia’”’; Washington, Grand Coulee (Apr.), Pullman 
(Apr.). 

Average specimens (40 mm. or more) make this the 
largest species in the genus. The Ragonot figure of 
polyphemella (Monograph, pl. 19, fig. 8a) is a very good 
likeness of normal examples, except that hind wing is a 
trifle too dark. 


327. Sarata pullatella (Ragonot), new combination 
FIGurE 397 


Megasis pullatella Ragonot, N. Amer. Phycitidae, p. 10, 1887; 
Monograph, pt. 1, p. 547, 1893. 


Smaller on the average than edwardsialis and darker. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 161 


Forewing almost uniformly"suffused dark grayish fus- 
cous (blackish gray on some examples); on some speci- 
mens the basal and terminal areas paler by contrast 
and the antemedial line indicated, but very faint, nearly 
vertical ; on occasional specimens a paler brownish shade 
in the cell; faint blackish streakings on the veins in 
terminal area (especially on specimens with pale outer 
area) and more or less of a black streak on lower vein 
of cell; subterminal line very faint, often completely 
obscured, when distinguishable indicated chiefly by a 
broken black shading forming its inner border; discal 
dots obscured. Hind wing pale to dark gray-brown; 
the cilia whitish. Alar expanse, 29-36 mm. 

Male genitalia figured from type and a typical speci- 
men from Dividend, Utah, to show extent of individual 
variation. Uncus not so evenly tapering as in edward- 
sialis; its lateral margins slightly angled at middle. 
Aedeagus less stout and the single long cornutus on penis 
more slender than those of edwardsialis; no supplemental 
patch on penis. 

Type Locatity: California (type in Paris Mus.). 

Foop piant: Unknown. 

Distrrisution: Utah, Dividend (Apr.); California, 
San Diego (Jan.); Jdaho, Malta; Washington, Kamiack 
Butte (Feb.), Pullman (Apr.). 

Easily confused with smaller specimens of edwardsialis, 
and all specimens in the National Collection had been so 
identified; but otherwise distinguished by its genitalia. 

Hulst in his Phycitidae of North America (1890) 
made pullatella a synonym of his excantalis and it has 
since appeared as such in our lists. Since the type of 
excantalis is a female, the synonymy is doubtful, to say 
the least. 


328. Sarata punctella (Dyar), new combination 
Figure 398 
Megasis punctella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 404, 
1915. 


Forewing pale brownish gray with a slight rufous tint 
through the cell; antemedial line indicated by an outer 
border of three black dots, one below costa, a second on 
lower vein of cell, and a third on vein 1b; subterminal 
line obscure but usually distinguishable, a whitish spot 
on costa, preceded and followed by blackish dots and, 
below, bordered inwardly by short blackish streaklet on 
veins; on typical specimens some faint, blackish streak- 
lets on the veins of outer area (following the subterminal 
line); a row of very faint blackish dots along termen; 
discal dots obsolete or very faint. Some specimens 
show little or no trace of the blackish markings. Hind 
wings pale brownish gray (“mouse gray”’); the cilia but 
slightly paler, a narrow dark line along termen. Alar 
expanse, 25-30 mm. 

Male genitalia with uncus elongate, slightly and 
evenly tapering to rounded apex. Apical process of 
gnathos appreciably shorter than in preceding species, 
slender. Elements of transtilla long, slender. Penis 
armed with two cornuti—one stout, slightly bent or 

300329—56—12 


sinuate, about half as long as aedeagus; the other a 
slender, flattened spine, as long as aedeagus. 

TypE Locauity: Tehuacin, México (type in USNM). 

Foop prant: Unknown. 

Distrisution: M&xico: ‘Las Vigas’’ [probably Las 
Viagas, Vera Cruz], Tehuacin (Oazaca, Sept.), Uruapan 
(Michoacdn, Mar.). Also three specimens labeled sim- 
ply “V.5.” One of the latter was before Dyar when he 
described hisspecies. It had been identified by Druceas 
“Zophodia inornatella Rag.” 

Dyar’s short original description is thoroughly ade- 
quate and accurate for the type but takes no account 
of suffused examples which lack the characteristic 
blackish markings. 


329. Sarata punctella septentrionaria, new race 


Like typical punctella but larger; the Arizona, New 
Mexico, and Colorado examples a suffused pale gray- 
brown with the usual dark markings obsolete or nearly 
so; the Wyoming example more grayish with most of the 
veins faintly streaked with blackish scaling. The 
genitalia agree with those of the type of punctella. Alar 
expanse, 32-35 mm. 

Typr Locauity: Palmerlee, Ariz. (type in USNM, 
61357). 

Foop prant: Unknown. 

Described from male type from the type locality, 
“Apr. 1-7,” and one male paratype from each of the 
following localities: Fort Wingate, N. Mex. (Apr. 14, 
1908); Golden, Colo., Dyar and Caudell, No. 16259; 
Medicine Bow, Wyo. July 4, 1936, I. H. Blake, No. 
321-11.” 

Possibly a distinct species but probably only a larger 
northern variety of punctella. When collections are 
made in the poorly explored areas of northern México 
the species should show a continuous distribution. 


330. Sarata incanella (Hulst), new combination 
Ficure 399 


Epischnia incanella Hulst, Canadian Ent., vol. 27, p. 56, 1895.— 
McDunnough, Check list, No. 6257, 1939. 

Megasis aridella Dyar, Proc. Ent. Soc., Washington, vol. 7, p. 
35, 1905.—McDunnough, Check list, No. 6263, 1939. (New 
synonymy.) 

Forewing gray heavily dusted with white especially 
in outer area and along costa at base and middle; the 
median area between the transverse lines darker than 
remainder of wing; dark shading also on basal area below 
costa; black streaking on the veins, especially pro- 
nounced in outer area, on vein 1b and upper and lower 
veins of cell; transverse lines white, distinctly outlined ; 
antemedial line zigzag, nearly vertical, with some black- 
ish shading along outer margin; subterminal line sharply 
indented at veins 6 and 1b; discal dots separated, black- 
ish; terminal dots faint but distinguishable, separated. 
Hind wing whitish with a faint smoky tint; a very faint 
dark line along termen; cilia concolorous with wing. 
Alar expanse, 31-36 mm. 

Male genitalia with uncus as long as greatest width; 
its apical margin broadly rounded. Apical process of 


162 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


gnathos a short, slender hook arising from a thickened 
base. Clasper of harpe broad, squarish. Aedeagus 
rather short; penis armed with a single small thornlike 
cornutus. Vinculum with terminal margin broadly and 
evenly rounded; in all other species of the genus the 
terminal margin distinctly concave. 

TypE LocaLities: Colorado (incanella, in USNM); 
Stockton, Utah (aridella, in USNM). 

Foop prant: Unknown. 

Distrizution: Colorado; Utah, Stockton (May); 
California, Inyo County (May). 

The palest of the Sarata species, except tephrella Rag- 
onot. The genitalia of the types of incanella and ari- 
della are identical. 


Genus Sarata (males) Species 331-337: S. atrella 
to S. tephrella 


[Antennal shaft with a ridge of roughened scales at base.] 


331. Sarata atrella (Hulst), new combination 
Ficure 400 


Megasis atrella Hulst, Phycitidae of N. Amer., p. 166, 1890.— 
Ragonot, Monograph, pt. 1, p. 548, 1893.—McDunnough, 
Check list, No. 6268, 1939. 

Vestiture of labial palpi, head, and prothorax a mix- 
ture of scales and long hairs. 

Forewing blackish gray; transverse lines whitish gray, 
dull; the antemedial line obscure; subterminal line 
stronger, always distinguishable, bordered inwardly and 
outwardly by black streaklets on the veins; discal spots 
obsolete, obscured in the dark ground color; blackish 
dots along termen confluent. Hind wing pale to rather 
dark smoky fuscous, a thin dark line along termen. Alar 
expanse, 30-33 mm. 

Male genitalia with uncus elongate, evenly tapering 
to narrowly rounded apex. Apical process of gnathos 
about half as long as uncus, ventrally flattened. Ele- 
ments of transtilla short. Penis armed with a single 
strong cornutus, as long as aedeagus, no supplemental 
sclerotized plate. — 

Type Locauiry: West Cliff, Colo. (type in AMNH, 
ex Rutgers). 

Foop prant: Unknown. 

Distrizution: Colorado, Custer County, West Cliff, 
and two specimens with only the state locality. 

The species is easily distinguished by the hairy vesti- 
ture not possessed by any other known species in the 
genus. 


332. Sarata caudellella (Dyar), new combination 
Ficure 401 
Megasis caudellella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p- 110, 1904.—McDunnough, Check list, No. 6261, 1939. 
Similar to the foregoing species (atrella) except: With- 
out hairy vestiture and with some fine white powdering 
on the blackish gray ground color of forewing; the 
transverse lines distinct, oblique and nearly straight; 
antemedial line followed on costa and preceded at inner 


margin by obscure dark blotches; subterminal line bor- 
dered inwardly by a continuous irregular blackish 
shade, outwardly by a short, faint, dark streaking of 
the veins; the veins otherwise not appreciably streaked; 
discal dots faint, but usually distinguishable, more or 
less confluent; dots along terminal margin weak, fused 
into a faint blackish line. Hind wing pale brownish 
gray, semilustrous; a dark line along termen. Alar ex- 
panse, 28-32 mm. 

Male genitalia differ from those of atrella only in 
trifling details. 


Typz Locatitry: Golden, Colo. (type in USNM). 
Foop piant: Unknown. 


Disrrisution: Unirep Starrs: Colorado, Golden 
(May); Washington, Grand Coulee (Apr.). CANADA: 
Saskatchewan, Oxbow (June) ; Manitoba, Aweme (Apr.), 
Miniota. 

Close to but distinct from atrella, from which it is 
distinguished chiefly by its smooth-scaled vestiture 
and the strong contrast of the whitish transverse lines 
of forewing, especially the well-marked antemedial line; 


333. Sarata dnopherella Ragonot 
Fieure 402 
Sarata dnopherella Ragonot, N. Amer. Phycitidae, p. 11, 1887; 
Monograph, pt. 1, p. 616, 1893. 

This and the two species following (nigrifasciella and 
cinereelia) are very close, separable only by minor color 
differences and trifling variations in the genitalia of their 
types. Such variations are more than covered in the 
series of nigrifasciella and cinereella before me. I sus- 
pect that the names represent nothing but color varie- 
ties of one variable species; but am keeping them sepa- 
rate until life-history information and more extensive 
collections are available and more exact definitions of 
species and possible races can be made. 

Ragonot’s dnopherella is authentically represented 
only by its type. His description and figure suggest a 
grayish brown form suffused with blackish brown and 
with the transverse lines very weakly contrasted and 
poorly defined against the ground color. Alar expanse, 
32 mm. 

Male genitalia with uncus broadly and bluntly tri- 
angulate, evenly tapering to narrowly rounded apex. 
Elements of transtilla moderately long, slender (about 
half the length of those of atrella). Penis armed with a 
single stout cornutus, slightly more than half as long 
as aedeagus and preceded basally by a small, weakly 
sclerotized patch. 

TyprE Locaity: California (type in Paris Mus.). 

Foop piant: Unknown. 

Hulst (Phycitidae of N. Amer., p. 168, 1890) treated 
dnopherella as a synonym of his perfuscalis and it so 
appears in our lists. As perfuscalis was described from 
a female the synonymising of the two names was 
arbitrary and, under the circumstances, unwarranted. 
Ragonot (Monograph, p. 616, 1893) very rightly ques- 
tioned it. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 163 


334. Sarata nigrifasciella Ragonot 
Fiaure 403 
Sarata nigrifasciella Ragonot, N. Amer. Phycitidae, p. 11, 1887; 
Monograph, pt. 1, p. 615, 1893.—Hulst, Phycitidae of N. 
eae p. 169, 1890.—McDunnough, Check list, No. 6266, 
1939. 

Forewing pale to dark ashy gray, some specimens 
tinted with a faint brownish shade (one specimen before 
me from Mineralking, Calif., a very pale, sordid 
brownish gray). The transverse lines distinct, but 
indicated chiefly by the blackish outer border of the 
antemedial line and a similar inner border to the sub- 
terminal line. This character, however, is variable and 
on the Idaho specimen before me is almost obsolete. 
Hind wing very pale brown, with a fine dark terminal 
line. Alar expanse, 29-32 mm. 

Male genitalia similar to those of dnopherella except 
for the narrower apical process of gnathos (viewed 
ventrally). This character, however, is not reliable; 
for in a typical series of cinereella all intergrades are 
found between the gnathos of typical nigrifasciella and 
that of dnopherella. 

Type Locatity: America Septentrionalis (type in 
Paris Mus.). 

Foop puant: Unknown. 

Disrrisution: Unirep Srares: New Mexico, Fort 
Wingate (Mar.); Colorado, Chimney Gulch (June), 
Platte Canyon (June); Jdaho, Wallace (Apr.); Cali- 
fornia, Mineralking (July), Yosemite (this specimen 
labeled nigrifasciella in Ragonot’s handwriting). Can- 
ADA: Manitoba, Aweme (Apr.). 

This species, if such it be, is distinguished from the 
preceding and following species chiefly by its more 
marked transverse lines in forewing and the narrower 
apical process of its gnathos. Ragonot’s description 
of the females (also from “Amer. Sept.’’) associated 
with his male type can be ignored, for they represent 
two distinct species. 


335. Sarata cinereella Hulst 


Sarata cinereella Hulst, Canadian Ent., vol. 32, p. 172, 1900.— 
McDunnough, Check list, No. 6271, 1939. 

Forewing dark ash gray, nearly concolorous; the 
transverse lines obsolete or nearly so; the subterminal 
line faintly indicated on most specimens and, on one or 
two, very faint traces of the antemedial line; as on 
dnopherella and nigrifasciella there is more or less 
blackish dusting on the veins. Hind wing pale brown- 
ish gray. Alar expanse, 29-33 mm. 

Male genitalia of type show a slightly longer cornutus 
than that of nigrifasciella, but other examples inter- 
grade in all characters between the two types. 

Type Locauity: Salida, Colo. (type in AMNH, ex 
Rutgers). 

Foop piant: Unknown. 

Disrrisution: Colorado, Denver (Mar.), Glenwood 
Springs, Salida, also three examples with only state 
locality, two of them bearing Hulst ‘‘type’’ labels 


(Bruce, collector), and the other a pseudotype of 
‘“Anerastia excantalis Hulst.”’ 
Probably only a suffused form of nigrifasciella. 


336. Sarata rubrithoracella (Barnes and McDunnough), new 
combination 


Fieurs 404 
Megasis rubrithoracella Barnes and McDunnough, Contributions, 
veer p. 140, 1913 —McDunnough, Check list, No. 6268, 

Forewing pale brownish gray rather evenly dusted 
with white intermixed with a sparse peppering of 
blackish scales; the transverse lines distinct, but faint, 
whitish; antemedial line bordered outwardly by a thin, 
broken, blackish shade; subterminal line bordered in- 
wardly by a continuous narrow blackish band; no ap- 
preciable streaking on the veins; discal spots obsolete; 
terminal dots confluent, forming a faint, dark line. 
Hind wing pale grayish brown; a thin dark line along 
termen. Alar expanse, 22-29 mm. 

Male genitalia show little to distinguish them from 
those of the preceding three species except for the very 
short lateral elements of transtilla and a slightly longer 
cornutus (our drawing of the aedeagus is in reversed 
position from that of other species; if drawn as were 
the other aedeagi the apex of cornutus would point to 
the right). 

Tyrer Locatity: White Mts., Ariz. (type in USNM). 

Foop pLant: Unknown. 

Distrisution: Arizona, White Mts.; New Mezico, 
Fort Wingate (Apr., July). 

Apparently a distinct species. Differs from the 
others by a distinct rufous-ocherous shading on the 
thorax and, in genitalia, from the species of the dnoph- 
erella—cinereella group by the short transtilla elements 
of its genitalia. The rufous ocherous thorax occurs 
also in two species of females (kappa and phi). 


337. Sarata tephrella Ragonot 
Fiaure 405 
Sarata tephrella Ragonot, Monograph, pt. 2, p. 616, 1893.— 
McDunnough, Check list, No. 6268, 1939. 

Known to me only from Ragonot’s description and 
figure, and the genitalia of its type. Evidently a much 
paler species than any other of those with a rough-scaled 
antennal shaft, and having much the general habitus of 
a Lipographis. Forewing heavily dusted with white; 
some ocherous brown shading in median and outer 
areas; transverse lines whitish, distinguishable but 
poorly defined. Hind wing dark gray. Alar expanse, 
28 mm. 

Male genitalia similar to those of the dnopherella- 
cinereella group except elements of transtilla larger and 
cornutus more slender and more evenly tapering. 

Type tocauity: ‘Washington Territory” (type in 
Paris Mus.). 

Foop prant: Unknown. 

Described and so far known only from its male type. 


164 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Genus Sarata (females), Species 338-346: 
S. alpha to S. delta 


[The females are of a uniform pattern and similar coloration; the 
ground color gray, sometimes tinted with pale brown and more 
or less dusted with white, variations in color more individual 
than specific; the transverse lines strongly contrasted, white with 
strongly accented blackish or brownish borders on outer margin 
of antemedial and inner margin of subterminal lines; the lines 
oblique and straight or notched, the notching also more individual 
than specific in character. The only reliable specific characters 
are in the genitalia, and for certain identification it is necessary 
to dissect nearly all females.] 


338. Sarata alpha, new species 
FIGURE 888 


A bright species with the white lines and their black 
borders sharply contrasted. Forewing blackish gray 
with a strong dusting of white (however, in one speci- 
men from the type locality, the median area distinctly 
darker than basal or outer areas); antemedial line 
slightly curved ; subterminal line straight or with a very 
slight bend at lower fold; discal dots distinct, more or 
less confluent. Hind wing whitish with a faint brown 
tint; the veins slightly darkened; a broadened blackish 
brown line along termen; cilia white. Alar expanse, 
21-24 mm. 

Female genitalia with bursa large and greatly elon- 
gated; densely and finely spined over most of interior 
surface, the denser spining in longitudinal rows, partial- 
ly divided by lines of the clear membrane; anterior end 
thickened (cartilaginous), the amount of thickening 
individually variable; ductus bursae very short. 

TYPE LOCALITY: Oxbow, Saskatchewan, Canada (type 
in USNM, 61358). 

Foop pirant: Unknown. 

Described from female type and two female paratypes 
from the type locality, May 14, 1907, Frederick Knab; 
and additional female paratypes as follows: One from 
Aweme, Manitoba, Apr. 12, 1908. N. Criddle; one from 
Regina, Saskatchewan, June 5, 1907; and two from 
Chimney Gulch, Golden, Colo., July, Oslar. A female 
in the Rutgers Collection (C. H. slide No. 2186) from 
Colorado identified as atrella also goes here. Needless 
to say there is no trace on any of these females of the 
hairy vestiture of the male of atredla. 


339. Sarata beta, new species 
Figure 889 


Similar to alpha except less glossy. Forewing duller; 
less white dusting; transverse lines and their black 
borders less strongly contrasted; discal dots obscured. 
Hind wing dark smoky gray; the veins not darkened. 
Alar expanse, 23-26 mm. 

Female genitalia like those of alpha in shape and 
proportions except that spining covers appreciably less 
of the bursa surface, leaving half or more than half of 
the latter membranous and unspined. 

Type Locauity: Colorado (type in USNM, 61359). 

Foop prant: Unknown. 

Described from female type with only a state locality 


label; one female paratype from Custer County, Colo.; 
and one female paratype from Chilcotin, British 
Columbia, May 2, 1920, E. K. Buckell No. 137. The 
two Colorado examples were in the Barnes and National 
Museum Collections as females of atrella Hulst. 


340. Sarata gamma, new species 


Figure 890 


This is the female figured in the Ragonot Monograph 
(pt. 1, pl. 23, fig. 2b.) as a paratype of his dnopher- 
ella. Its genitalia, here figured, are similar to those of 
alpha and beta except for slight differences in the spiming 
of the bursa, as shown in the figure. 

Type Locatity: California (type in Paris Mus.). 

Foop prant: Unknown. 


341. Sarata iota, new species 
FIGURE 894 


This name is proposed for the female paratype of 
pullatella Ragonot, described and figured by him 
(Monograph, pt. 1, p. 547, pl. 19, fig. 9b, 1893). 

Forewing blackish gray with very little pale dusting; 
the antemedial line broader and more strongly con- 
trasted than the subterminal. Hind wing dark grayish 
brown. Alar expanse, 24 mm. 

Genitalia (C. H. slide No. 3113) with bursa, consider- 
ably smaller and more sparsely spined than that of any 
of the preceding species; ductus bursze about half as 
long as bursa. 

Tyrer Locauity: California (type in Paris Mus.). 

Foop puant: Unknown. 


It is very likely that this is the female of pullatella; 
but at the present time there is no certainty about any 
of the sex associations in the genus. 


342. Sarata perfuscalis (Hulst) 
Figure 893 


Nephopteryx perfuscalis Hulst, Trans. Amer. Ent. Soc., vol. 13, 
p. 161, 1886 

Anerastia excantalis Hulst, Trans, Amer. Ent. Soc., vol. 13, 
p. 163, 1886 (mew synonymy). 

Megasis excantalis (Hulst), Ent. Amer., vol. 5, p. 156, 1889; 
Phycitidae of N. Amer., p. 165, 1890.—McDunnough, 
Check list, No. 6260, 1939. 

Sarata perfuscalis (Hulst), Phycitidae of N. Amer., p. 168, 1890.— 
MceDunnough, Check list, No. 6269, 1939. 


Forewing dull, dark gray more or less dusted with 
white; the terminal and (usually) the median areas the 
paler, the basal area the darker; transverse lines dis- 
tinct, sordid white, their dark borders well contrasted, 
especially on specimens with considerable white dusting. 
Hind wing smoky grayish brown. Alar expanse, 25— 
29 mm. 

Female genitalia distinguished by the spining of the 
large bursa. These spines are arranged in an elongate, 
ribbed band which extends most of the length of the 
inner dorsolateral surface, curving onto ventral surface 
at anterior end; the area of bursa under the spines more 
or less sclerotized. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 165 


TyprE Locauity: California (perfuscalis, excantalis, in 
AMNGH, ex Rutgers). 

Foop prant: Unknown. 

Distrisution: California (state locality only); 
Washington, Seattle; Utah, Eureka (Mar., Apr.), 
Stockton (Apr.). 

Hulst associated his perfuscalis with dnopherella 
Ragonot; and excantalis with puillatella Ragonot. So 
much for superficial sex associations. Their genitalia 
show the two females to be obviously conspecific. 


343. Sarata epsilon, new species 
Figure 892 


The smallest of the female species. Similar in colora- 
tion to perfuscalis except that the dark borders of the 
transverse whitish lines are somewhat broader and more 
strongly contrasted. Alar expanse, 19-21 mm. 

Female genitalia with bursa moderately large, evenly 
and finely spined on anterolateral half and extreme 
anterior end. 


TypE Locauity: Yosemite, Calif. (type in USNM, 
61360). 

Foop prant: Unknown. 

Described from female type from the type locality 
identified by Ragonot as a female of nigrifasciella and 
bearing that name label in his handwriting and female 
paratypes as follows: One from Colorado, Cockerell, 
collector, identified by Hulst as atrella; two from Golden, 
Colo., May, Dyar and Caudell Nos. 16252 and 16253, 
and identified by Dyar as paratypes of caudellella; 
seven from Chimney Gulch, Golden, Colo., June, 
Oslar; and one from Fort Wingate, N. Mex., March. 
Here also is referrable one of the two female paratypes of 
nigrifasciella Ragonot in the Paris Museum (C. H. slide 
No. 2891). 

Most of the foregoing examples show at least one of 
the black discal spots. On each of the seven specimens 
from Chimney Gulch, Colo., is 8 minute white spot on 
discocellular vein between the black dots and on these 
also there is some very dark brown shading on the 
otherwise blackish borders of the white transverse lines. 


344. Sarata phi, new species 
Fiaure 891 


Thorax shaded with rufus ocherous. 

Forewing more evenly dusted with white; dark 
borders of the transverse lines distinctly brownish; dis- 
cal spots obsolete, replaced by a faint, white line or spot 
on the discocellular vein. Alar expanse, 23-27 mm. 

Female genitalia essentially like those of epsilon. The 
figure shows the bursa twisted into a reverse position 
from that of epsilon to show the somewhat greater 
development of the thickened (cartilagenous) lateral 
margin (a variable and probably only an individual 
character). 

TypE LocaLity: White Mts., Ariz. 
61361). 

Foop piant: Unknown. 

Described from female type from the type locality 


(type in USNM, 


(the female paratype of rubithoracella Barnes and 
McDunnough); and five female paratypes from Fort 
Wingate, N. Mex. March, June, July. Also before 
me, but not included among the paratypes, is a large 
female (26.5 mm.) from Denver, Colo., Apr. 1, 1904, 
Oslar. On this specimen the patagia are more putty 
white than rufus. Its genitalia, however, agree in 
detail with those of typical phi from Arizona and New 
Mexico. The species is uncomfortably close to epsilon. 
It probably does represent the female of rubithoracella 
but the verification of that relation will have to wait. 
upon rearing evidence. 


345. Sarata kappa, new species 
FIGURE 887 


Forewing dull, as in beta, but with considerable white 
dusting, rather evenly distributed; the transverse lines 
more irregular and their black borders more strongly 
contrasted; subterminal line with slight notches at vein 
6 and lower fold; lower discal dot faint, but distinguish- 
able. The thorax of the type is strongly shaded with 
rufus-ocherous. Alar expanse, 23 mm. 

Female genitalia with bursa copulatrix very small 
(the smallest of any of the Sarata species); the greater 
part of its inner dorsal surface covered with a dense mat 
of very fine spines. Ductus bursae as long as bursa. 

Tyrer Locauity: Arizona (type in USNM, 61362). 

Foop piant: Unknown. 

Described from a pseudotype of perfuscalis Hulst 
from the Fernald Collection, bearing only a state 
locality and numbered ‘‘7820.” 

In coloration similar to phi except for the blackish 
borders of the transverse lines of forewing. Dis- 
tinguished from that and other species of the genus by 
its genitalia. 

346. Sarata delta, new species 
Ficure 886 


This name is proposed for the second of the female 
paratypes of nigrifasciella Ragonot (in Paris Mus., C. 
H. slide No. 3111) whose genitalia are here figured. 


80. Philodema, new genus 


Tyre or Genus: Sarata rhoiella Dyar. 

Tongue well developed. Antenna pubescent; shaft 
of male slightly flattened and with a shallow sinus near 
base, the latter containing some slightly roughened 
scales and a few, minute serrations. Labial palpus 
porrect (as in Sarata but shorter). Maxillary palpus 
vestigial. Venation as in Sarata except veins 4 and 5 
of hind wing stalked for half or less than half their 
lengths and cell a short one-third the length of wing. 
Eighth abdominal segment of male with ventrolateral 
hair tufts. 

Male genitalia as in Sarata except: More squat, 
broader in proportion to their length; harpe short in 
proportion to its width; no erect clasper; anellus 
strongly sclerotized, its central area developed into a 
pair of produced, pointed, bladelike arms, the usual 


166 


lateral lobes absent; aedeagus slender, its basal end 
broadened and flattened; penis without cornutus or 
other armature except for a few weak scobinations at 
apex; vinculum stout, shorter than its greatest width. 

Female genitalia with bursa small, simple, membran- 
ous; ductus bursae short with a strongly sclerotized, 
curved, wide, centrally notched, dorsal plate behind 
genital opening; ductus seminalis from a small lobe of 
bursa near junction of bursa and ductus bursae. 

The genus falls between Sarata and Hypochalcia and 
has several features of each of these genera but can go 
into neither of them on the sum of its characters. It 
appears to be a New World analogue of the Old World 
Hypochalcia, agreeing with the latter on most genitalic 
characters except for its short vinculum and peculiarly 
developed anellus. The type of Hypochalcia (ahenella 
(Zeller), fig. 54) has an elongate vinculum. It also 
differs from Philodema in having much longer, smoother 
and slenderer labial palpi, rather broad, squamous 
maxillary palpi and smoother more glossy wing vesti- 
ture. 

Philodema differs markedly from Sarata in that there 
is no sexual dimorphism, the males and females being 
alike in color and markings. 


347, Philodema rhoiella (Dyar), new combination 
Fiaures 406, 895 


Sarata rhoiella Dyar, Journ. New York Ent. Soc., vol. 12, p. 105, 
1903.—McDunnough, Check list, No. 6267, 1939. 

Forewing pale, sordid, brownish gray; extreme basal 
area dark smoky gray; the transverse lines indicated 
chiefly by their dark borders, the latter dark smoky 
gray; the outer border of the antemedial line more or 
less broken and diffused; subterminal line bordered 
inwardly by an irregular (zigzag) border, somewhat ac- 
cented on the veins, and outwardly by a more obscure 
dark shade; discal dots distinct, separated. Hind wing 
smoky gray; the veins slightly darkened; a narrow dark 
shade along termen. Alar expanse, 23-33 mm. 

Genitalia as given for the genus; figured from para- 
types from the type locality. The male holotype was 
without an abdomen. 

Tyre tocauity: Platte Canyon, Colo. (type in 
USNM). 

Foop puant: Rhus toxicodendron. 

Distripution: Colorado, Platte Canyon (July) ; Utah, 
“So. Utah” (July). 

The Utah specimens (2 <) are larger (82-33 mm.) 
than any of the Colorado examples; but have identical 
genitalia and wing maculation; nothing is known of the 
life history except Dyar’s statement that two specimens 
of the type series were reared from larvae on poison-ivy. 


81. Genus Lipographis Ragonot 


Lipographis Ragonot, N. Amer. Phycitidae, p. 10, 1887; Mono- 
graph, pt. 1, p. 562, 1893.—Hulst, Phycitidae of N. Amer., 

p. 166, 1890. (Type of genus: Pempelia fenestrella Packard.) 
Tongue well developed. Antenna weakly pubescent; 
on male, shaft flattened, serrate, and with sinus and 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


strong scale tuft at base (except in umbrella and subos- 
seella where the shallow sinus has a weak tuft of rough- 
ened scales). Labial palpus porrect, second segment 
oblique, laterally flattened, broadly scaled; third seg- 
ment deflected forward, decidedly shorter than second, 
its proportions obscured by its long scaling and the 
extended scaling of second segment. Maxillary palpus 
subsquamous (small but broadly scaled, vestigial in 
umbrella). Forewing smooth; venation as in Sarata 
except for a somewhat shorter stalking of veins 8 and 9 
of forewing and a longer cell in hind wing (nearly one- 
half the length of the wing); 4 and 5 are also shorter 
stalked, about one-half their lengths.’ Highth abdomi- 
nal segment of male with a pair of ventrolateral hair or 
scale tufts (absent in subosseella). 


Male genitalia with apical projection of gnathos a 
short stout hook (except in subosseella). Transtilla 
absent except in fruncatella and subosseella where it is 
represented by its short, weak, divided elements. Anel- 
lus with short, weak, lateral lobes (except in subosseella). 
Aedeagus broadly expanded towards apex (except in 
truncatella); penis armed with one or more strongly 
sclerotized, curved, spinelike cornuti (the latter always 
decidedly less than half as long as the aedeagus). Geni- 
talia otherwise as in Sarata. 


Female genitalia with bursa membranous and greatly 
reduced, if sometimes elongate (truncatella) narrow; 
ductus bursae scobinate and partially sclerotized near 
its junction with bursa copulatrix, greatly broadened in 
proportion to width of bursa (except in éruncatella); 
genital opening simple, unsclerotized; ductus seminalis 
from bursa near junction of bursa and ductus bursae. 


Lipographis agrees with Philodema and differs from 
Sarata in that the males and females are alike in color 
and markings. It is distinguished from both Sarata 
and Philodema chiefly by its female genitalia. The 
latter resemble those of the type of the Old World 
Divona Ragonot (ilignella (Zeller)) except that the 
bursa of 2lignella is strongly scobinate, partially sclero- 
tized, and proportionally much larger. 

In his original description of Lipographis, Ragonot 
designated fenestrella as type of the genus. Later 
(Monograph, 1898) he cites humilis as its type. ‘This 
substituted designation is invalid, regardless of the fact 
that humilis was an originally included species and may 
have served as the basis for the original generic descrip- 
tion. That humilis now proves to be a synonym of 
fenestrella is also immaterial and irrelevant. 

One species (swbosseella) originally described in Iipo- 
graphis is here provisionally retained in the genus. It 
may eventually have to have a new generic placement 
as its only representation (the male type) is aberrant 
in several genitalic details. 


7 The venation of fenestrella and leoninella exhibit considerable 
individual variation; veins 4 and 5 of forewing are normally ap- 
proximate for a short distance from cell but sometimes divergent 
and (rarely) even shortly stalked. In one freak specimen before 
me vein 4 is also absent from hind wing, another example which 
advises caution against relying too much upon one structure for 
the identification of phycitids. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 167 


348. Lipographis fenestrella (Packard) 
Ficurrs 31, 407, 896 


Pempelia fenestrella Packard, Ann. New York Lyc. Nat. Hist., 
vol. 10, p. 259, 1873. 

Nephopteryx fenestrella (Packard) Grote, Bull. U. 8S. Geol. Geogr. 
Surv. Terr., p. 697, 1878. 

Lipographis humilis Ragonot, N. Amer. Phycitidae, p. 11, 1887; 
Monograph, pt. 1, p. 563, 1893.—Hulst, Phycitidae of N. 
Amer., p. 167, 1890. (New synonymy.) 

Lipographis fenestrella (Packard) Ragonot, N. Amer. Phycitidae, 
p- 10, 1887; Monograph, pt. 1, p. 564, 1893.—Hulst, Phy- 
citidae of N. Amer. p. 166, 1890.—McDunnough, Check 
list, No. 6272, 1939. 

Forewing ash gray, dusted with white and shaded 
with brownish ocherous; the transverse lines narrow, 
white; antemedial line oblique, nearly straight, bordered 
inwardly by a broad brownish ocherous band marked 
by black dots or streaklets on vein 1b and upper and 
lower veins of cell, followed outwardly by two or three 
similar black dots; white dusting along lower vein of 
cell, median part of vein 1b and along some of the veins 
preceding the subterminal line; subterminal line parallel 
to termen, very slightly indented at veins 6 and 1b, 
bordered outwardly by a broad brownish ocherous band 
(the latter interrupted by blackish streaklets on the 
veins) and from costa by short, faint, narrow, inner and 
outer, blackish bordering lines; along termen a narrow 
dusting of white; terminal dots more or less confluent, 
individually variable, forming sometimes a straight, 
sometimes a scalloped, black line; discal dots separated, 
small, blackish; usually a brownish ocherous shade 
along median area of lower fold. Hind wing dull white 
with a faint, smoky tint towards apex and termen;a fine 
blackish line along terminal margin; the veins not 
appreciably darkened. Alar expanse, 19.5-24 mm. 

Male genitalia with aedeagus decidedly bulged from 
shortly beyond base; penis armed with a comb of 5 
stout, curved spines of a graduating length. Female 
genitalia with bursa greatly reduced and but slightly 
longer than ductus bursae; the latter appreciably 
broader than the bursa. 

Type tocauity: California (fenestrella, in MCZ; hu- 
milis, in Paris Mus.). 

Foop prant: Unknown. 

Distrizution: California, ‘Middle California,’ Palo 
Alto (May), San Diego (Apr., May, June, July, Aug., 
Oct.), San Francisco. 

The type of Ragonot’s humilis is merely a small, 
rather dark male of fenestrella with identical genitalia. 
In any considerable series of fenestrella the palpal differ- 
ences cited by Ragonot can be observed. They are 
indeed more apparent than real and more due to pro- 
portionate differences in the sizes of the individual 
specimens and to differing positions of the palps. In 
our latest checklists humilis is listed as a subspecies or 
variety of fenestrella. It is not even that. 


349. Lipographis leoninella (Packard) 


Pempelia leoninella Packard, Ann. New York Lyc. Nat. Hist., 
vol. 10, p. 259, 1873. 

Nephopteryzx leoninella (Packard) Grote, Bull. U. S. Geol. Geogr. 
Surv. Terr., vol. 4, p. 697, 1878. 


Lipographis leoninella (Packard) Ragonot, Ent. Amer., vol. 5, 
p. 115, 1889; Monograph, pt. 1, p. 565, 1893.—MceDunnough 
Check list, No. 6273, 1939. 

Lipographis fenestrella leoninella (Packard) Hulst, Phycitidae of 
N. Amer., p. 167, 1890. 

Pyla pallidella Dyar, Journ. New York Ent. Soc., vol. 12, p, 107, 
1903.—Barnes and McDunnough, Contributions, vol. 3, p. 
199, 1916 (make synonym of leoninella). 

Forewing similar in maculation to that of fenestrella 
except: General color more ocherous than gray, the gray 
shading limited to the median area between the trans- 
verse line; basal area of wing pale ocherous; the 
inner border of the antemedial and outer border of the 
subterminal lines yellow; lower fold between the trans- 
verse lines pale ocherous; no appreciable black streaking 
on the veins of outer area; discal spot at lower outer 
angle of cell larger, more conspicuous. Hind wing paler, 
with a faint ocherous tint towards apex and termen. 
Alar expanse, 21-24 mm. 

Male and female genitalia like those of fenestrella. 

Type tocaities: California (leoninella, in MCZ); 
Salt Lake, Utah (pallidella, in USNM). 

Foop piant: Unknown. 

Distrisution: Unitep States: California, ‘“Mid- 
dle Calif.,”” Olancha (Inyo County, June), Palo Alto 
(May), Sonoma County (May); Utah, Richfield (Aug.), 
Salt Lake, Stockton (July), Vineyard (June, July). 
Canava: Manitoba, Cartwright (Aug.). 

Hulst treated leoninella as a variety of fenestrella, and 
probably correctly. There is nothing to separate the 
two except coloration. However, as nothing isknown 
about their biology, it seems the better part of wisdom 
to keep the two names apart. Larval characters and 
habits and hosts may indicate separate species or at 
least distinct races. 


350. Lipographis truncatella (Wright), new combination 
Figures 408, 898 


Hypochalcia truncatella Wright, Ent. News, vol. 27, p. 25, 1916.— 
McDunnough, Check list, No. 6276, 1939. 

Forewing a dull, pale, brownish ocherous, dusted with 
white and a fine peppering of black scales, making the 
general color an ashy gray with a strong suffusion of the 
ground color, the latter most pronounced in basal area 
and in lower half of median area; antemedial line faint, 
without any appreciable inner border, its outer border 
indicated by black dots on costa, upper and lower veins 
of cell, and on vein 1b; subterminal line distinct, with a 
narrow, faint, but distinguishable and continuous inner, 
black, bordering line. Hind wing pale gray, very faintly 
tinted with ocherous toward base and shading into a 
smoky hue towards apex and termen; a strong narrow 
dark shade along termen. Alar expanse, 22-25 mm. 

Male genitalia distinguished chiefly by its much 
slenderer aedeagus, narrowing at apex, and the single, 
very short, thornlike, curved cornutus on penis. Dif- 
ferences also in the shape of the sclerotization of the 
eighth segment tergite of abdomen are shown in the 
figure. Female genitalia figured from a San Diego 
specimen in the National Collection (W. S. Wright, 
June 23, 1911). Bursa narrowly elongate; ductus bur- 


168 UNITED STATES NATIONAL 
sae much shorter than bursa, and narrow (no wider than 
bursa, except at genital opening). 

TypPE LocaLity: San Diego, Calif. (type probably 
lost). 

Foop prant: Unknown. 

Distrisution: California, Chula Vista (June), San 
Diego (June), 

Despite its striking specific differences in genitalia 
and more broadly scaled labial palpi this species fits 
well into Lipographis. It is certainly not a Hypochal- 
cia. The latter, an Old World genus, as far as I know 
is not represented in our fauna. 


351. Lipographis umbrella (Dyar), new combination 
Ficures 410, 897 


Sarata umbrella Dyar, Proc. Ent. Soc. Washington, vol. 10, p. 59, 
1908.—McDunnough, Check list, No. 6270, 1939. 

Male antenna with a weak ridge of roughened scales 
in shallow sinus at base of shaft. 

Forewing orange yellow; transverse lines narrow, 
white; antemedial line oblique, somewhat curved, set 
well out on wing and with only the faintest indication 
of a dark outer border, the latter sometimes containing 
a few black scales; subterminal line nearly straight, with 
only a slight median bulge, inwardly more or less bor- 
dered with black, the latter color varying from a thin, 
weak line to large smudges extending well into the 
median area of the wing; on some specimens a narrow 
oblique blackish shade just beyond basal attachment of 
wing; discal dots obscure, often obliterated by streaks 
of white scaling or extensions of the black border of the 
antemedial line. Hind wing semilustrous, ocherous 
with a smoky suffusion, the latter most pronounced on 
dark specimens; veins not appreciably darkened. Alar 
expanse, 26.5-31 mm. 

Male genitalia similar to those of fenestrella and 
leoninella; differing from them chiefly in the armature 
of the penis; the latter consists of a comb of 6 or 7 curved 
spines and another straight spine, near but distinctly 
separated from the comb. Female genitalia differing 
only in minor details from those of fenestrella. 

TYPE LOCALITY: San Diego, Calif. (type in USNM). 

Foop prant: Unknown. 

Distrisution: California, Laguna, Long Beach 
(Sept.), Los Angeles (Sept.), Petaluma (Sept.), San 
Diego (Aug., Sept.). 

Dyar placed the species in Sarata on the basis of its 
male antenna character; but its genitalia as well as the 
lack of any sexual dimorphism in wing maculation or 
color show that it belongs in Lipographis. 


352. Lipographis (?) subosseella Hulst 
Fieure 409 
Iipographis subosseella Hulst, Canadian Ent., vol. 24, p. 62, 
1893.—Ragonot, Monograph, pt. 1, p. 565, 1893. 

Male antenna with a very weak scale ridge in shallow 
sinus at base of shaft. 

Thorax and forewing sordid white overshaded with 
dull ocherous; the whitish ground color most noticeable 


MUSEUM BULLETIN 207 

along costa, the ocherous shade strongest along lower 
fold and in outer area, making the general color of the 
wing (to the naked eye) a pale brownish yellow; trans- 
verse lines nearly obsolete; antemedial line distinguish- 
able only as an oblique whitish streak from lower vein 
of cell to inner margin, preceded on inner margin by a 
blackish brown smudge; subterminal line indicated only 
by its dark borders, a pale brownish, rather broad outer 
band and a fainter, narrower, broken inner line; discal 
dots separated, weak, blackish brown; a half-dozen 
narrow blackish dots along termen. Hind wing white 
with some fuscous shading at apex, on the outer veins, 
and, narrowly, along termen; cilia shining white. Alar 
expanse, 16 mm. 

Male genitalia uncus triangulate, its apex narrowly 
rounded. Apical process of gnathos an elongate, rather 
narrow, veutrally flattened hook. Harpe elongate, 
slender, with very small thornlike clasper. Anellus 
without lateral lobes. Aedeagus broadly expanded at 
apex; penis armed with a single, stout, curved cornutus 
and a small supplemental detached sclerotized plate. 
Kighth abdominal segment simple (without tufts). 

TypE LocaLity: Bahama Islands (type in AMNH, 
ex Rutgers). 

Foop prant: Unknown. 

An anomalous species, differing in its tropical habitat, 
wing pattern, and many details of male genitalia from 
other species of Lipographis, in which genus it is tenta- 
tively retained. Eventually, when more material is 
available, especially some female examples, a new ge- 
neric reference may be needed. At present the species 
is known only from its unique male type. 


Genera 82-86: Adelphia to Acroncosa 


[Venational division B. Forewing smooth; vein 2 from near lower 
outer angle of cell; 4 and 5 separated at base. Hind wing with 
7 and 8 closely approximate for a short distance from cell, rarely 
(in individual specimens) weakly anastomosed; cell short, less 
than one-half the length of cell. Female genitalia with signa 
developed as opposed, strongly spined plates (except in Tota, 
where bursa is smooth); ductus seminalis from bursa.] 


82. Adelphia, new genus 


Typr or Genus: Pempelia petrella Zeller. 

Tongue well developed. Antenna pubescent; on male 
a short, shallow sinus with small scale tuft in base of 
shaft. Labial palpus oblique on male, upturned on 
female, reaching to, but not appreciably above vertex; 
second segment long, on male grooved to hold the maxil- 
lary palpus; third segment minute, acuminate. Maxil- 
lary palpus of male in the form of an aigrette; of female 
squamous. Forewing smooth; 11 veins; vein 2 from 
before but rather near lower outer angle of cell; 3 from 
the angle, but little further at base from 2 than from 4; 
4 and 5 separated at cell and divergent very shortly be- 
yond it; 6 from below upper angle of cell, straight; 8 
and 9 stalked for slightly less than half their lengths; 
10 from the cell, approximate to the stalk of 8-9 for a 
short distance from cell; male without costal fold. Hind 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 169 


wing with vein 2 from well before lower outer angle of 
cell; 3 from the angle connected with 4 by a short 
spur; 4 and 5 contiguous or anastomosed for about 
half their lengths; 7 and 8 closely approximate for less 
than half their lengths; cell less than half the length of 
wing; discocellular vein curved, produced outwardly at 
lower angle. Eighth abdominal segment of male with 
two pairs of ventrolateral hair tufts (the tufts stouter 
and the hairs broadened and flattened in ochripunctella). 

Male genitalia with uncus broad, hoodlike. Apical 
process of gnathos a simple, strongly sclerotized hook. 
Transtilla absent. Harpe elongate, slender; costa 
strongly sclerotized on basal half; a strongly sclerotized 
arm or hook arising from the lower edge of the sclero- 
tized costa at base; cucullus narrow, tapering to a blunt 
point. Anellus with short, weak lateral lobes. Penis 
without cornuti or other appreciable armature. Vin- 
culum stout. 

Female genitalia with signa strongly developed, con- 
sisting of two opposed, densely spined plates (in petrella 
an additional collar of similar spines about the posterior 
third of bursa); ductus bursae short, funnel-shaped, 
sclerotized except at junction with bursa where it is 
more or less finely scobinate; genital opening simple; 
ductus seminalis from bursa at junction of the bursa and 
ductus bursae. 

This and the two genera following (Tota and Ufa) are 
closely related to Elasmopalpus, and each contains a 
species withdrawn from the latter genus. Such with- 
drawal was necessary if Hlasmopalpus was to be ac- 
curately defined. With its previous inclusions such 
definition was impossible. 


3538. Adelphia petrella (Zeller), new combination 
Fieurss 411, 904 


Pempelia petrella Zeller, Isis von Oken, 1846, p. 771; 1848, p. 886; 
Verh. zool.—bot. Ges. Wien, vol. 22, p. 545, 1872. 

Nephopteryx rubiginella Walker, List, vol. 27, p. 55, 1863. 

Nephopteryz rufinalis Walker, List, vol. 27, p. 56, 1863. 

Nephopteryx hapsella Hulst, Ent. Amer., vol. 3, p. 132, 1887. 

Elasmopalpus petrellus (Zeller) Hulst, Phycitidae of N. Amer., 
p. 158, 1890.—Ragonot, Monograph, pt. 1, p. 421, 1893.— 
Forbes, Cornell Mem. 68, p. 628, 1923.—McDunnough, 
Check list, No. 6229, 1939. 

Head and thorax reddish brown or brownish ocherous 
with some gray shading on the mesothorax. 

Forewing brownish gray with a fine white dusting in 
median area especially toward costa; antemedial line 
very faint, an irregular, interrupted, fine white line well 
out on the wing, bordered outwardly by two or three 
small black dots and preceded by a broad brownish 
ocherous or reddish brown band; the inner edge of the 
latter more or less shaded with black heavily dusted 
with white; extreme base of wing shaded with blackish 
gray; subterminal line usually distinct, narrow, and 
finely dentate, white with a very weak dark inner border 
except at costa but, on all well-marked specimens, 
followed outwardly by some black streaklets on the 
veins; discal dots separated, black, the lower one always 
distinct, the upper weaker and sometimes obscured; a 
row of small black dots along terminal margin set off 


by a narrow dusting of white. Hind wing pale grayish 
brown; the veins slightly darkened; on most specimens 
a narrow smoky shade along outer margin. Alar ex- 
panse, 20-27 mm. 

Male genitalia with terminal margin of uncus rounded 
and with a very slight notch at apex. Apical process 
of gnathos a very stout hook. Projecting hook from 
subcostal base of harpe a long, stout, nearly straight 
arm, nearly as long as costa of harpe. Aedeagus not 
appreciably widened towards apex.’ Female genitalia 
with bursa considerably elongated, slender for over half 
its length and with a collar of strong spines near its 
middle. 

Type tocauities: North America (petrella, in Mus. 
Univ. Berlin); United States (rubiginella and rufinalis, 
in BM); Florida (hapsella, in AMNH, ex Rutgers). 

Foop prant: Unknown. 

Distrisution: Florida, Coconut Grove, Glenwood, 
Lake Alfred (July), Orlando (Feb., Mar., Apr.), St. 
Petersburg (Feb.), Tampa, Vero Beach (Apr., June, 
Sept., Oct., Dec.); Georgia (Feb., Mar.); Texas, Browns- 
ville, San Benito, Victoria (Mar.); North Carolina, 
Raleigh (Apr.), Southern Pines (Aug.), Tryon (May, 
Aug.); Virginia, Virginia Beach (Aug.); District of 
Columbia (May, July); New Jersey, Anglesea (May), 
Woodbury (May); Jowa, Iowa City (July). 

The Walker species (rubiginella and rujinalis) are 
included in the synonymy on the strength of Ragonot’s 
reference which is probably correct, for he presumably 
examined their types. I have not. Hulst’s hapsella 
agrees in genitalia and all other characters with typical 
petrella. Its type is a female, not a male as stated by 
Hulst. Honora obsipella Hulst is also listed as a 
synonym of petrella in our lists, but incorrectly. It is 
a synonym of Hulstia undulatella (see p. 196). 


354. Adelphia ochripunctella (Dyar), new combination 
Figures 412, 901 


Salebria ochripunctella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 59, 1908 —MeDunnough, Check list, No. 6202, 1939. 
Forewing mouse gray; the transverse lines obsolete 
except for a faint indication of the antemedial line which 
is oblique, nearly straight, and a dull pale ocherous gray; 
a similarly faint ocherous discal spot at end of cell. Ex- 
cept for these markings the wing is unicolorous. Hind 
wing subpellucid, whitish with a faint smoky tint at 
apex and narrowly along terminal margin; the veins 
not appreciably darkened. Alar expanse, 17-21 mm. 
Male genitalia with uncus slightly constricted towards 
terminal margin, which is slightly concave; its lateral, 
apical angles produced into short, ventrally projecting 
lobes. Apical process of gnathos a small hook. Pro- 
jecting hook from subcostal base of harpe, curved, about 
half the length of costa of harpe. Aedeagus enlarged 
(bulging) towards apex, and with some minute scobina- 


8 The lateral projection from near apex of aedeagus shown in 
our drawing (fig. 411a) is merely a partially sclerotized fragment 
of the membrane connecting aedeagus and anellus and not a 
projecting thorn or spine such as occurs in some species of Pyla. 


170 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


tions on its outer surface (the latter are visible only 
under high magnification and are somewhat exaggerated 
in fig. 412a). Female genitalia with bursa much re- 
duced as compared to that of petrella; armed only with 
two opposed, spined plates. 

TypE Locauity: San Diego, Calif. (type in USNM). 

Foop piant: Eremocarpus setigerus. ‘This record 
from an El Segundo specimen reared by W. D. Pierce. 

Distrizution: California, El Segundo (Oct.), San 
Diego (Oct., Nov.). 

A distinct species easily identified by its peculiar 
wing markings and male genitalia. 


83. Tota, new genus 


Typr or Genus: Elasmopalpus galdinella Schaus. 

Characters of Adelphia except: Labial palpus con- 
siderably longer, reaching well above vertex, especially 
on the male. Hind wing with veins 4 and 5 stalked for 
two-thirds of their lengths. Eighth abdominal segment 
of male with compound ventral tufts. Male genitalia 
with complete transtilla (a narrow, angulate, band); 
aedeagus slender, elongate; penis armed with an elon- 
gate, sclerotized plate bearing a row of six minute, 
thornlike spines. Female genitalia with bursa simple, 
without signa, spines, or other sclerotization; ductus 
bursae narrow, tubular, sclerotized throughout its 
length, not expanded or funnel shaped; genital opening 
narrow. 

The genus is distinguished from the others in this im- 
mediate group by its genitalia and the rather long stalk- 
ing of veins 4 and 5 of hind wing. Like the genus pre- 
ceding (Adelphia) and that following (Ufa), it appears 
to be closely related to Elasmopalpus, in which its type 
and only known species was placed by Schaus. 


355. Tota galdinella (Schaus), new combination 
Fiaures 413, 899 


Elasmopalpus galdinella Schaus, Zoologica, vol. 5, no. 2, p. 46, 


Forewing blackish brown with the transverse lines 
strongly contrasted, narrow, white; some scattered 
white dusting in the subbasal and outer areas and, on 
the male, appreciable white dusting over the medial 
area; antemedial line oblique, irregularly dentate; sub- 
terminal line zigzag, nearly vertical; no appreciable dis- 
cal spots; a row of faint, separated, black dots along 
terminal margin. Hind wing pale brownish (more 
whitish on the male); the veins faintly darkened and a 
smoky shade along outer margin. Alar expanse, 18- 
24 mm. 

Male genitalia with tegumen longer than greatest 
width, its terminal margin straight and slightly pro- 
duced at the lateral angles. Female genitalia with char- 
acters as given for the genus. 

Type Locatiry: Conway Bay, Indefatigable, Galé- 
pagos Islands (type in USNM). 

Foop puant: Unknown. 

Distrisution: GauApagos Isuanps: “Camp Beta” 
(Jan.), Conway Bay (Apr.), South Seymore (Apr.). 


84. Genus Ufa Walker 


Ufa Walker, List, vol. 27, p. 79, 1863. 
venezuelalis Walker.) 

Tongue well developed. Antenna pubescent; on 
male with a short, shallow sinus with small scale tuft 
in base of shaft. Labial palpus oblique in both sexes 
(except in the female of rubedinella where it is some- 
what upcurved); third segment nearly half the length 
of second; extending well above vertex. Maxillary 
palpi as in Adelphia. Venation as in Adelphia except: 
Vein 3 of forewing on the average, closer to 4 than to 2; 
hind wings with vein 4 and 5 anastomosed for about 
half their lengths (or in some specimens of rubedinellus 
and senta stalked for a trifle over half), cell one-third 
the length of wing. Highth abdominal segment of male 
with compound, ventral hair and scale tufts (except on 
lithosella which has simple hair tufts). 


Male genitalia without any sclerotized arm or hook 
arising from base of costa of harpe; the costa strongly 
sclerotized (in lithosella this sclerotized part of costa 
terminating well before the apex of the harpe); clasper 
present as a digitate or weak, rounded or triangulate, 
erect projection from upper edge of sacculus. Anellus 
a broad plate with short, lateral lobes. Aedeagus stout. 
Penis armed with a single, stout, rather short, curved 
spines. 

Female genitalia similar to those of Adelphia; ductus 
bursae short and broad, more or less funnel shaped, 
with broad genital opening; ductus seminalis from 
bursa well before junction of bursa and ductus bursae. 

This genus, while sharing many of the characters of 
Adelphia is closest to Elasmopalpus, from which it is 
distinguished by its more oblique female labial palpi, 
the strongly sclerotized costa of harpe, the shape of its 
anellus, the stouter, much shorter apical projection of 
gnathos, and the broader female ductus brusae. 


(Type of genus: Ufa 


356. Ufa lithosella (Ragonot), new combination 
Fiaures 414, 903 
Selagia lithosella Ragonot, N. Amer. Phycitidae, p. 9, 1887; 
Monograph, pt. 1, p. 474, 1893.—Hulst, Phycitidae of N. 
Amer., p. 160, 1890.—McDunnough, Check list, No. 6233, 
1939. 
Honora luteella Hulst, Journ. New York Ent. Soc., vol. 8, p. 223, 
1900 p 


Ancylostomia lithosella (Ragonot) Dyar, Ins. Insc. Menstr., vol. 
7, p. 53, 1919. 

Forewing pale ocherous; the ground color interrupted 
by a narrow, paler, cream-colored shade along costal 
edge, along lower margin of the cell, sometimes along 
lower fold and (in a few examples) between the veins 
in outer area; transverse lines obsolete; on some speci- 
mens a dark grayish spot on vein 1b indicates what 
remains of a dark outer margin to the antemedial line; 
on occasional examples a dark grayish shade on middle 
of inner margin; discal dots sometimes absent but the 
lower one usually distinguishable, minute, blackish; 
the usual dark dots along outer margin rarely dis- 
tinguishable and when so only as a slight darkening of 
the ground color. Hind wing semitransparent white 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 171 


with a more or less pale smoky tint; the veins and 
terminal margin not appreciably darkened, except on 
the darkest specimens. Alar expanse, 26.5-32 mm. 

Male genitalia with harpe very short, its costa 
broadly and strongly sclerotized, the sclerotized costal 
area terminating in an abrupt projection, well before 
apex of cucullus. Lateral lobes of anellus knoblike. 
Cornutus a short, stout, slightly curved, bluntly 
pointed, hornlike thorn with a few minute serrate 
projections on one side. Eighth abdominal segment 
of male with a single pair of ventrolateral hair tufts. 

Female genitalia with bursa armed with one large and 
one small, round, strongly spined plate; the ductus 
seminalis from bursa near the caudal margin of the 
smaller plate; ductus bursae smooth except for a 
slight, irregularly shaped sclerotization near genital 
opening. 

TypeE Locatities: Arizona (lithosella, in Paris Mus.); 
Santa Rita Mts., Ariz. (luteella, in USNM). 

Foop prant: Unknown. 

Distrisution: Unirep States: Arizona, Baboqui- 
vari Mts. (July, Oct.), Chiricahua Mts., Huachuca 
Mts., Nogales (May), Palmerlee, Paradise (Cochise 
County, Mar.), Santa Rita Mts. (June), Wilgus (Co- 
chise County); New Mexico, Albuquerque. Mfxtco: 
Cuernavaca (June), Durango (city), México (city, Oct.), 
Venadio (Sinaloa), Zacualpin (June). 

A distinct species, easily recognized by its genitalia 
and obviously not closely related to Selagia where it 
was placed by Ragonot, nor to Ancylostomia where it 
was placed in the National Collection by Dyar. 


357. Ufa roseitinctella (Dyar), new combination 


Ancylostomia roseitinctella Dyar, Proc. U. S. Nat. Mus., vol. 42, 
p. 105, 1912. 

Forewing pale ocherous, the ground color suffused 
with pale rose red broadly along inner margin and costa 
and over most of median and outer areas; the ocherous 
color on most specimens limited to the basal area and 
(on a few examples) to a narrow, pale, longitudal shade 
through the middle of the wing; transverse lines obso- 
lete; discal dots minute, but usually distinguishable 
(at least the lower one), blackish, separated; terminal 
dots absent. Hind wing pale smoky fuscous, with a 
very faint ocherous tint; the veins very slightly dark- 
ened and a faint, dark line along termen. Alar expanse, 
26-32 mm. 

Female genitalia similar to those of lithosella. 

Type tocauiry: Cuernavaca, México (type in 
USNM). 

Foop prant: Unknown. 

Distrrisution: Unrrep Status: Arizona, Huachuca 
Mts. (Sept.), Paradise (Cochise County, July, Oct.), 
Pea Mts., Wilgus Mts. México: Cuernavaca 

y). 

This species is known only from females. It is very 
close to and may only be a color form of lithosella, but 
this cannot be determined until a male is recovered. 
The Arizona specimens were in our collection under 
(Selagia) Zamagiria australella (Hulst). 


358. Ufa senta, new species 
Fiecurss 415, 905 


Forewing pale canary yellow with a narrow brownish 
red shade along inner margin; costa pale on outer half, 
whitish with a fine, sparse peppering of fuscous scales; 
from apex inward along vein 6 a reddish fuscous streak 
continued, on most specimens, as a faint, more or less 
interrupted, dark shade along top of cell to base of wing; 
lower discal spot minute, but usually distinct, dull red; 
cilia peppered, ashy gray white. Hind wing translucent 
white with a smoky shade towards outer margin; veins 
in outer area of wing faintly darkened; a narrow fuscous 
shade along termen. Alar expanse, 22-25 mm. 

Male genitalia having harpe with costa sclerotized to 
apex; clasper moderately long, slender, digitate. Anellus 
with lateral lobes moderately long, strongly sclerotized, 
tapering to blunt points. Cornutus a sharply curved, 
strong, rather slender spine with a broad base. 

Female genitalia with bursa rather narrowly elongate 
with a lateral lobe near junction of bursa and ductus 
bursa; signa consisting of two greatly elongated oval 
lobes densely armed with long, slender spines; a second 
cluster of minute spines adjacent to the posterior, 
lateral lobe; ductus bursa short, triangulate, sclerotized 
almost to genital opening. 


Type tocauity: Big Bend, Tex. (type in USNM, 
61363; paratype, co’, in BM). 

Foop prant: Unknown. 

Described from male type from the type locality 
(Apr. 15, 1926); four male and one female paratypes 
from Palmerlee, Ariz.; and one female paratype from 
the Huachuca Mts., Ariz. 

A distinct, easily recognized species, apparently near 
to rubedinella Zeller. 


359. Ufa rubedinella (Zeller), new combination 
Ficurss 416, 902 


Pempelia rubedinella Zeller, Isis von Oken, 1848, p. 885; Verh. 
zool.-bot. Ges. Wien., vol. 24, p. 480, 1874; Horae Soc. Ent. 
Rossicae, vol. 16, p. 181, 1881. 

Acrobasis translucida Walker, List, vol. 27, p. 29, 1863. 

Nephopteryx rufescentalis Walker, List, vol. 27, p. 58, 1863. 

Nephopteryx minualis Walker, List, vol. 27, p. 60, 1863. 

Nephopteryx deprivalis Walker, List, vol. 27, p. 60, 1863. 

Ufa venezuelalis Walker, List, vol. 27, p. 60, 1863. 

Elasmopalpus pyrrhochrellus Ragonot, Nouv. Gen., p. 23, 1888; 
Monograph, pt. 1, p. 429, 1893 (new synonymy). 

Elasmopalpus rubedinellus (Zeller) Ragonot, Nouv. Gen., p. 23, 
1888; Monograph, pt. 1, p. 340, 1893.—Wolcott, Journ. 
Agr. Univ. Puerto Rico, vol. 25, no. 2, p. 134, 1941. 


Forewing a pale, glossy ocherous, more or less shaded 
and marked with red or reddish brown; the females 
usually darker and showing more of the reddish shading 
than the males, some examples entirely suffused with 
red except for a contrasted, whitish or very pale ocher- 
ous shade along costa; three small but conspicuous 
blackish or dark reddish dots in median area, two well 
out from base, obliquely placed on lower vein of cell 
and vein 1b, and one at lower outer angle of cell (on 
some specimens indications of another dot at costa 
opposite the one on vein ib and, very rarely, faint 


172 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


traces of an upper discal dot at end of cell); from apex 
a short oblique, reddish shade more or less peppered 
with fuscous; faint tracings of red on the outer extrem- 
ities of the veins and, on darker examples, a reddish 
blush over the entire outer area; subterminal line in- 
dicated by a narrow, oblique, twice-indented, red or 
reddish fuscous line rather close to terminal margin. 
Hind wings shiny, translucent white with a fine brown 
line along termen; the veins not appreciably darkened. 
Alar expanse, 16-22 mm. 

Male genitalia with erect clasper of harpe short, 
rounded, knoblike. Anellus with lateral lobes short, 
stubby. Cornutus shaped like that of senta but appre- 
ciably stouter. Terminal margin of vinculum slightly 
produced at lateral angles. Female genitalia with 
rather small narrow bursa containing a collar of fine 
spines at junction of bursa and ductus bursae in addi- 
tion to the normal] pair of spined signa; the broad, short 
ductus bursae sclerotized throughout, its lower terminal 
margin at genital opening sharply concave. 


TYPE LOCALITIES: Brazil (rubedinella, in BM); Santo 
Domingo (translucida and rufescentalis, in BM); Hon- 
duras (minualis, in BM); Venezuela (deprivalis and 
venezuelalis, in BM); Corrientes, Argentina (pyrrhochrel- 
lus, in Paris Mus.). 

Foop piants: Limabeans, black-eyed peas. These 
records from Florida specimens reared by the Special 
Survey of the Division of Foreign Plant Quarantine of 
the U. S. Bureau of Entomology and Plant Quarantine. 
Larvae feeding on the leaves. 

DistRIBUTION.—UniTED States: Florida, Egmont 
(Apr.), Hobe Sound (May), Hypoluxo, Lake Beach 
(Feb.), Palm Beach, Palmetto, Vero Beach (Sept., 
Oct., Dec.), Winter Park (July). Cusa: Havana, 
Matanzas (June), Santiago Province (Oct.). Domrni- 
can Repusiic. Puerto Rico: Catafio July), Coamo 
Springs (Apr.), Dovado (May), Isabela Substation 
(Apr.), Palmas Abajas (Apr.), Puerto Real (Vieques Isl., 
Apr.), San German (Apr.). Viren Isuanps: Kingshill 
(St. Croix, Mar., Apr., June). Jamaica. TRINIDAD: 
Quare River Valley (Jan.). M*#x1co: Chiapas (May), 
Guadalajara, Guerrero (Dec.), Oaxaca, Sierra de Gue- 
rrero (Nov.), ‘Mexican Sub-region.” GUATEMALA: 
Cayuga (Feb., Mar., Apr., May, June, Dec.), Chejel 
(Aug.), Guatemala (city, Mar.), Quirigué (Mar.). 
Honpuras. Costa Rica: Juan Vifias (Jan., June, 
Dec.). Panamd: Alhajuela (Mar., Apr.), Corazal 
(May, June, July), La Chorrera (Apr.), Obispo, Paraiso 
(Apr., May), Porto Bello (Apr., May, Oct.), Taboga 
Isl. (Jan., July), Tabernilla. Venrzunta. FRENcH 
Gurana: St. Jean Maroni, St. Laurent Maroni. Bo- 
Livia: Esperanza, Prov. del Saré (Dept. Santa Cruz, 
May). Braz: Rio de Janiero (Nov.). Paraguay: 
Villarrica (Jan., Mar., Oct., Nov., Dec.). AnrgEn- 
TINA: Corrientes, Metin (Prov. del Salta, Feb.). 
Pert: Chanchamayo. Generally distributed through- 
out the tropical and subtropical regions of the New 
World. 

In many respects (its habitus, upturned female labial 
palpi, sexual dimorphism, and similarity of larval 


habit) this species seems to go with Elasmopalpus 
lignosellus, with which it has been associated; but on 
male and female genitalic characters it belongs definitely 
with the species here assigned to Ufa, and on all larval 
characters it is radically different from lignosellus. In 
structural and pattern characters its larva resembles 
that of Caristanius decoloralis. 


85. Genus Elasmopalpus Blanchard 


Elasmopalpus Blanchard, in Gay, Historia fisca y politica de 
Chile. Zoologia, vol. 7, p. 104, 1852.—Hulst, Phycitidae of 
N. Amer., p. 157, 1890.—Ragonot, Monograph, pt. 1, p. 418, 
1893.—Forbes, Cornell Mem. 68, p. 628, 1923.—Janse, 
Journ. Ent. Soc. South Africa, vol. 7, p. 4, 1944. (Type of 
genus: Hlasmopalpus angustellus Blanchard.) 

Tongue well developed. Antenna weakly pubescent; 
on male a sinus and a short scale tuft in base of shaft. 
Labial palpus of male erect and appressed to face, reach- 
ing well above vertex; third segment minute, hidden in 
scaling of second segment; of female upcurved, reaching 
slightly above vertex; third segment but slightly shorter 
than second, acuminate. Maxillary palpus of male in 
the form of an aigrette; of female, squamous. Forewing 
smooth; 11 veins; vein 2 from close to lower outer angle 
of cell; 3 from the angle; 2, 3 and 4 approximately 
equidistant at base (the position of 2 individually vari- 
able, on some specimens very close and occasionally 
connate with 3); 4 and 5 separated at base, approximate 
for a short distance from cell; 6 from below upper angle 
of cell, straight; 8 and 9 stalked for less than half their 
lengths; 10 from the cell; male without costal fold. 
Hind wing with vein 2 from well before lower outer angle 
of cell; 3 from the angle, separated from 4 by a very 
short spur; 4 and 5 stalked for at least two thirds of their 
lengths; 7 and 8 closely approximate, contiguous or 
weakly anastomosed fora very short distance from cell; 
cell about one-third the length of wing; discocellular 
vein curved, produced at lower angle. Highth abdom- 
inal segment of male with compound ventral scale and 
hair tufts. 


Male genitalia with uncus rather narrowly triangu- 
late, its apex narrowly rounded. Transtilla absent. 
Apical process of gnathos a slender, elongate hook. 
Harpe with costa not appreciably sclerotized; sacculus 
with upper margin an erect, irregularly serrate ridge, 
slightly produced at apex. Anellus U-shaped, with 
lateral arms strongly sclerotized and produced as 
curved pointed horns. Aedeagus slender; penis armed 
with a single, slightly curved, slender, strongly sclero- 
tized cornutus, from slightly more than one-third to 
one-half as long as aedeagus. Vinculum stout, longer 
than greatest width, tapering to narrowly rounded or 
bluntly pointed terminal margin. 

Female genitalia with signa consisting of two opposed, 
strongly spined plates; ductus bursae cylindrical, longer 
than bursa, sclerotized for half its length from junction 
with bursa (the sclerotized part longitudinally ribbed), 
membranous on posterior half, expanding to wide 
genital opening, weakly sclerotized on inner dorsal 
surface behind genital opening; ductus seminalis from a 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 173 


lobe of bursa near junction of bursa and ductus bursae. 
As here defined the genus is limited to its type 
species. None of the other species that have hitherto 
been assigned to Hlasmopalpus fits comfortably into the 
genus. 
360. Elasmopalpus lignosellus (Zeller) 
Fiaures 33, 417, 906 


Pempelia lignosella Zeller, Isis von Oken, 1848, p. 885; Verh. zool.- 
bot. Ges. Wien, vol. 22; p. 544, 1872; vol. 24, p. 430, 1874; 
Horae Soc. Ent. Rossicae, vol. 16, p. 180, 1881.—Riley, in 
Rep. [U. 8.] Comm. Agr. for 1881, pp. 142-145, 1882. 

Elasmopalpus angustellus Blanchard, in Gay, Historia fisca y 
politica de Chile. Zoologica, vol. 7, p. 105, 1852.—Berg, Bull. 
Soc. Imp. Nat., Moscou, p. 228, 1875; Anales Soc. Cient. 
Argentina, vol. 4, pt. 4, p. 209, 1877 (makes synonym of 
Pempelia lignosella). 

Pempelia lignosella tartarella Zeller, Verh. zool.-bot. Ges. Wien, 
vol. 22, p. 544, 1872; Horae Soc. Ent. Rossicae, vol. 16, p. 
180, 1881. 

Pempelia lignosella incautella Zeller, Verh. zool.-bot. Ges. Wien, 
vol. 22, p. 544, 1872; Horae Soc. Ent. Rossicae, vol. 16, p. 
180, 1881. 

Pempelia lignosella major Zeller, Verh. zool.-bot. Ges. Wien, vol. 
24, p. 430, 1874. 

Elasmopalpus anthracellus Ragonot, Nouv. Gen., p. 23, 1888; 
Monograph, pt. 1, p. 428, 1893 (new synonymy). 

Dasypyga carbonella Hulst, Ent. Amer., vol. 4, p. 114, 1888. 

Elasmopalpus lignosellus (Zeller) Ragonot, Ent. Amer., vol. 5, p. 
115, 1889; Monograph, pt. 1, p. 425, 1893.—Hulst, Phyci- 
tidae of N. Amer., p. 159, 1890.—Chittenden, U. 8S. Dep. 
Agr. Div. Ent. Bull. 23, pp. 17-22, 1900.—Luginbill and 
Ainslie, U. S. Dep. Agr. Bull. 539, 27 pp., 1917.—Forbes, 
Cornell Mem. 68, p. 628, 1923.—Metcalf and Flint, Destruc- 
tive and useful insects, ed 1, pp. 338-339, 1928; ed 2, pp. 367— 
368, 1939.—McDunnough, Check list, No. 6231, 1939.— 
Costa-Lima, Insetos do Brazil, p. 2, p. 93, 1950.—Craighead, 
U.S. Dep. Agr. Misc. Publ. 657, p. 454, 1950. 

Elasmopalpus puer Dyar, Ins. Insc. Menstr., vol. 7, p. 53, 1919 
(new synonymy), 

Ground color of forewing (on male) dull ocherous 
fawn, transverse lines obsolete; on paler specimens a 
subbasal black spot on vein 1b, one on lower vein of cell 
beyond middle and another at lower outer angle of cell; 
a row of faint, more or less confluent blackish dots along 
terminal margin; a blackish shade of varying width 
along costa and inner and outer margins; on females the 
dark shading more extended, reducing the ocherous 
ground color to a narrow, longitudinal shade, or com- 
pletely suffusing the wing; many female examples en- 
tirely black, with some occasional sparse reddish scaling 
at extreme base of wing. Hind wing translucent white, 
with a faint darkening of the outer veins and a narrow 
brownish shade along terminal margin. Alar expanse, 
16-24 mm. 

Male and female genitalia: as given for the genus. 

TYPE Locauities: Brazil (lignosellus, in BM); Rio 
Negro, Brazil (anthracellus, in Paris Mus.); Concepcién, 
Chile (angustellus, in Paris Mus.); Valparaiso, Chile 
(major, in BM); Santiago Province, Cuba (pwer, in 
USNM); Texas (tartarella and incautella, in BM; car- 
bonella, in AMNH, ex Rutgers). 

Foop piants: Beans, corn, cowpeas, chufa (Cyperus 
exculentus), crabgrass, sudangrass, Johnsongrass, Japa- 


nese cane, milo maze, sugar cane, sorghum, peanuts, 
turnips, wheat, strawberry plants, flax, cotton, black 
locust. Larvae boring into the stems of growing plants 
and, to a lesser extent, feeding on the leaves. 


Disrrizution.—Unitep States: Florida, Everglade 
(Apr.), Fort Myers (Apr.), Hastings (Sept., Oct.), Lake 
Alfred (July), Lakeland (Oct.), Miami (Oct.); Georgia, 
Savannah (July); Alabama, Auburn (Aug.), Eufaula; 
Lowsiana, Baton Rouge (July, Aug.), New Orleans 
(June); Texas, Blanco County (Mar.), Brownsville 
(Mar., June, Oct.), Burnet County (Mar.), College 
Station, Dallas (Oct.), Dickinson (Oct.), Gainesville 
(Aug.); Kerrville (Aug.), San Benito (Apr., June, July), 
Smith Point (Sept.), Victoria (Apr., July); Arizona, 
Baboquivari Mts. (Oct.), Mesa, (Aug.), Phoenix (Aug.), 
Tempe (Oct.); California, San Diego (Aug.); South 
Carolina, Columbia; Virginia, Cape Henry (June), Nor- 
folk (Sept.); District of Columbia, Washington (Aug., 
Oct., Nov., Dec.); New Jersey, Montclair (Sept.); 
Massachusetts, Cohasset (July, Aug.), Newton High- 
lands. Brrmupa (Mar., Apr., May). Cusa: Havana, 
Matanzas, Santiago de las Vegas (Apr.), Santiago 
Province (Oct.). Purrro Rico: Anasco, Coamo Springs 
(Apr.), Puerto Real (Vieques Isl., Apr.), Rio Piedras 
(Aug.), San German (Apr.), Villalba (July). Viren 
Isuanps: St. Croix (Apr.). Topsaco (June). JAMAIca. 
México: Barmos (Sinaloa, Mar.), Chiapas, Colima 
(Mar.), Guadalajara, Izuala (June), Mexico City (Nov.), 
Orizaba, Tehuacin (May, June, Aug.). GUATEMALA: 
Cayuga (Apr., June, July), Chejel (June, July, Aug.), 
Guatemala City (Mar.), Perulhi (July), Quirigud 
(Mar.), Volecén Santa Maria (Nov.), PanamA: Alha- 
juela (Apr.), Corozal (May, July), La Chorrera (Apr., 
May), Paraiso (May), Porto Bello (Mar., Apr., May, 
Sept., Oct.), Rio Trinidad (Mar.). VEnEzuEva: Las 
Cruces, Colén (Dec.). Frencu Guiana: Cayenne, St. 
Jean Maroni. Braziu: Castro (Parand), Rio Negro, 
Santa Catarina (Oct.), Sdo Paulo (Sept.). Paraguay: 
Paraguayan Chaco (Makthlawaiya, Nov., Nanahua, 
Mar.), Villarrica (Feb., Sept., Oct., Nov.). ARGENTINA: 
Tucumin (Nov.), Villa Cina (Feb., Mar., Nov.). 
Curie: Valparaiso (Apr.). Prrt: Cafiete (Oct.). Gen- 
erally distributed throughout the tropical and temperate 
regions of the New World. 


An insect of some economic importance in our 
Southern States and known in economic literature as 
the “lesser cornstalk borer.’”’ It is individually variable 
in color and the sexes are markedly dimorphic. As a 
result it has received several names. The latter have 
no taxonomic value for they represent at most only 
color forms. As far as I know the species is confined 
to the New World, where it is abundant and widely 
distributed. 

The Luginbill and Ainslie paper (1917) gives all avail- 
able information on the life history of the species. 
There has been no contribution of any importance since 
its publication. 


174 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


86. Genus Acroncosa Barnes and McDunnough 


Acroncosa Barnes and McDunnough, Canadian Ent., vol. 49, 
p. 404, 1917. (Type of genus: Acroncosa albiflavella Barnes 
and McDunnough.) 

Tongue well developed. Antenna simple except for 

a few minute thornlike projections on base of shaft of 

the male (no appreciable sinus or scale tuft); very 

shortly and weakly pubescent in both sexes. Labial 
palpus short, slender, oblique, third segment shorter 
than second, blunt, projected slightly forward; not 
reaching to height of vertex. Maxillary palpus 
squamous. Forewing smooth; 11 veins; vein 2 from 
before but near lower outer angle of cell; 3 from the 
angle; 4 and 5 separated at base; 6 from below upper 
angle of cell, straight; 8 and 9 stalked for half or a 
trifle over half their lengths; 10 from the cell, separated 
at base from 8-9 and not approximate to its stalk; 
male without costal fold. Hind wing with vein 2 from 
well before lower outer angle of cell; 3 from the angle 
and approximate to 4 for a short distance beyond the 
angle; 4 and 5 contiguous or anastomosed beyond the 
angle for about half their lengths (superficially, on 
undenuded specimens, they appear long stalked); 7 and 

8 closely approximate for a short distance from cell; 

cell less than half the length of wing; discocellular vein 

curved, considerable produced outwardly at lower angle. 

Eighth abdominal segment of male with a pair of ventro- 

lateral hair tufts (the hairs somewhat flattened). 


Male genitalia with uncus broadly triangulate, its 
apex bluntly pointed. Transtilla incomplete, its ele- 
ments moderately sized, irregularly triangulate plates. 
Harpe simple; costa sclerotized nearly to apex, not 
produced. Anellus a shallow U-shaped plate with small 
lateral lobes. Aedeagus long, stout; penis armed with 
masses of spines varying from long (one-third the length 
of aedeagus) to minute. (Figs. 418c-e show the shape 
and character of the longer spines, greatly enlarged.) 
Vinculum stout, longer than greatest width, tapering 
to truncate terminal margin. 


Female genitalia with signa consisting of two small, 
opposed, strongly spined, round plates; bursa otherwise 
membranous except for a thickening of the lobe giving 
off the ductus seminalis; ductus bursae long, slender, 
tubular, unsclerotized except for a couple of weak spine 
clusters near junction with bursa; genital opening 
simple; ductus seminalis from bursa near its junction 
with ductus bursae. 

Foretibia with a long inner and short outer claw. I 
should suspect this character to be merely specific 
except that it is equally developed in both sexes of both 
species of the genus. 


361. Acroncosa albiflavella Barnes and McDunnough 
Ficurss 35, 418, 907 
Acroncosa albiflavella Barnes and McDunnough, Canadian Ent., 
vol. 49, p. 405, 1917.—McDunnough, Check list, No. 6151, 
1939. 
Forewing white; the transverse white lines lost in the 
ground color; antemedial line indicated only by a broad 


oblique inner orange band extending from inner margin 
to costa; subterminal line indicated by a similar, 
narrower, sinuate, outer orange band, the latter termi- 
nating at costa in a small blackish spot; a well-con- 
trasted black discal dot at lower, outer angle of cell and 
a smaller black dot on inner margin of the subbasal 
orange band at vein 1b; on same specimens a few 
widely scattered black scales on the white ground color 
of median area. Hind wings whitish with a very faint 
ocherous or smoky tint; the veins not appreciably 
darkened. Alar expanse, 18-21 mm. 

Genitalia as given for the genus. 

Type Locatiry: Loma Linda, San Bernardino Coun- 
ty, Calif. (type in USNM). 

Foop piant: Unknown. 

Distrisution: California, Loma Linda (July, Aug.), 
Mona Lake (July). 


362. Acroncosa albiflavella castrella Barnes and McDunnough 


Acroncosa albiflavella castrella Barnes and McDunnough, 
Canadian Ent., vol. 49, p. 405, 1917. 

A local race of albiflavella differing only in having a 
stronger peppering of black scales in the median area of 
forewing. Alar expanse, 20-22 mm. 

Genitalia like those of typical albiflavella. 


Type tocatity: Fort Wingate, N. Mex. (type in 
USNM). 

Foop prant: Unknown. 
pe usenuaeNs New Mexico, Fort Wingate (July, 

ug.). 

Apparently a distinguishable local race, but can be 
identified only by locality label and a somewhat 
stronger dusting of black scales on forewing. 


363. Acroncosa similella Barnes and McDunnough 


Acroncosa similella Barnes and McDunnough, Canadian Ent., 
vol. 49, p. 405, 1917——MceDunnough, Check list No. 6152, 
1939. 

Forewing similar to albiflavella except: A much 
heavier peppering of black scales on forewing; orange 
inner border of antemedial line extending only from 
inner margin to top of cell; a concentration of black 
scaling forming a more or less broken line along outer 
margin of the antemedial line; a similar black line 
inwardly bordering the subterminal line; basal, median, 
and outer areas dusted with black scales. Alar expanse, 
21-26 mm. 

Genitalia like those of albiflavella. 

Type LocaLity: Pyramid Lake, Nev. 
USNM). 

Foop Prant: Unknown. 

Distrisution: Nevada, Pyramid Lake; Utah, Eureka 
(June). 

Not structurally distinct from and possibly only a 
local race of albiflavella. The incomplete subbasal 
orange band on forewing, however, suggests a distinct 
species. 


(type in 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 175 


Genus 87: Passadena 


[Venational division B. Forewing with weak subbasal scale 
ridge; veins 4 and 5 shortly stalked; 10 from the cell. Hind 
wing with vein 2 near angle of cell; 3 from the stalk of 4-5; 4 and 
5 long stalked. Labial palpus short, porrect. Male genitalia 
with a strongly developed clasper on harpe. Female genitalia 
with signa developed as opposed, strongly spined plates; ductus 
seminalis from bursa.] 


87. Genus Passadena Hulst 


Passadena Hulst, Canadian Ent., vol. 32, p. 171, 1900. 
genus: Passadena constantella Hulst). 

Tongue well developed. Antenna weakly pubescent; 
on male with a shallow sinus and scale tuft in base of 
shaft. Labial palpus short, porrect; second segment 
oblique, the third deflected forward, minute, more or 
less hidden in the thick scaling of second segment; the 
latter grooved on male to hold the maxillary palpus. 
Maxillary palpus of male in the form of an aigrette; of 
female, squamous. Forewing with weak subbasal 
ridge of raised scales; 11 veins; vein 2 from before but 
near lower outer angle of cell; 3 from the angle; 4 and 5 
shortly stalked; 6 from below upper angle of cell, 
straight; 8 and 9 stalked for about half their lengths; 
10 from the cell; male without costal fold. Hind wing 
with vein 2 from near lower outer angle of cell; 3 from 
the stalk of 4-5; 4 and 5 long stalked (for over two- 
thirds their lengths; 7 and 8 approximate or contiguous 
for a very short distance from cell; cell about half the 
length of wing; discocellular vein curved, outwardly 
produced at lower angle. Eighth abdominal segment 
of male with a weak pair of ventrolateral hair tufts. 

Male genitalia with uncus triangulate; apical process 
of gnathos a strong, narrow, elongate hook. Transtilla 
absent. Harpe with costa narrowly sclerotized for its 
entire length; a strong, projecting clasper from upper 
edge of sacculus near base (the clasper a striking feature; 
its peculiar shape probably a specific character). Penis 
armed with several short rows of weak spines with a 
mat of fine scobinations between them. Vinculum 
stout, stubby, slightly broader than long, terminal mar- 
gin broadly truncate. 

Female genitalia with signa developed as a pair of 
opposed, strongly spined plates, one (ventral) rather 
large, elongately oval, the other (dorsal) very small and 
bearing only two or three spines; ductus bursae short, 
broad, sclerotized for most of its length, and bearing on 
midventral surface a row of minute spines; genital open- 
ing simple; ductus seminalis from bursa near junction 
of bursa and ductus bursae. 

A distinct, monotypical, New World genus with one 
western North American species. 


(Type of 


364. Passadena flavidorsella (Ragonot) 
Figures 36, 419, 908 


Anoristia flavidorsella Ragonot, N. Amer. Phycitidae, p. 9, 
1887.—Hulst, Phycitidae of N. Amer., p. 160, 1890. 

Meroptera canescentella Hulst, Phycitidae of N. Amer., p. 149, 
1890.—Ragonot, Monograph, pt. 1, p. 319, 1893.—MceDun- 
nough, Check list, No. 6186, 1939. (New synonymy.) 

Getulia flavidorsella (Ragonot), Monograph, pt. 1, p. 528, 1893. 


Passadena constantella Hulst, Canadian Ent., vol. 32, p. 171, 1900. 

Megasis cinctella Hulst, Canadian Ent., vol. 32, p. 172, 1900 (new 
synonymy). 

Passadena cinctella (Hulst) Barnes and McDunnough, Contribu- 
tions, vol. 3, p. 198, 1916.—MeDunnough, Check list, No. 
6222, 1939. 

Passadena flavidorsella Barnes and McDunnough, Check list of 
the Lepidoptera of Boreal America, No. 5647, 1917.— 
McDunnough, Check list, No. 6221, 1939. 

Forewing whitish gray, more or less finely dusted with 
black scales; transverse lines narrow, whitish; ante- 
medial line somewhat oblique, twice indented, bordered 
outwardly by a narrow black line, inwardly by a broader 
black band of roughened scales, the black borders form- 
ing a conspicuous black band divided by a narrow pale 
line; subterminal line sinuate-angulate, bordered in- 
wardly by a black line (most strongly accented at 
costa) and outwardly at costa by a short black streak; 
discal dots faint, only distinct on specimens with a pale 
(whitish) ground color; a row of inconspicuous black 
dots along termen. Hind wing white to pale smoky 
fuscous, frequently with a slightly smoky shade towards 
apex; veins not appreciably darkened; a fine brownish 
line along outer margin. Alar expanse, 15-20 mm. 

Genitalia as given for the genus. The male can be 
readily identified by the peculiar shape of the clasper 
of harpe (fig. 419e). 

Type Locauitigs: Arizona (flavidorsella, in Paris 
Mus.); Texas (canescentella, in AMNH, ex Rutgers); 
California (constantella, in AMNH, ex Rutgers); Argus 
Mts., Calif. (cinctella, in USNM). 

Foop puant: Unknown. 

Disrrisution: Texas, Brownsville (May), “Central 
Texas” (the types of canescentella, 2 and o’, without 
date); New Mexico, state locality only (July, Cockerell, 
collector); Arizona, Ajo (Pima County, Mar.), Babo- 
quivari Mts. (Apr., May, July, Aug., Sept.), Catalina 
Springs (Apr.), Oracle (July), Phoenix (Apr.), Reding- 
ton, Sells Post Office (Pima County, Apr.), “Salt River 
Mts.” (Sept.), “Southern Arizona” (Aug.), Tempe 
(Feb., May), Tucson (May), Wenden (Yuma County, 
Aug.), Yuma County (‘Colorado Desert,’ Mar.); 
California, Argus Mts. (Apr., May), Inyo County 
(June, July), Jacumba (June), La Puerta (July), Los 
Angeles (May), Mason Valley (San Diego County, 
Apr.); Utah, Richfield (May). Also reported by Rago- 
not from Sonora, México. 

The species is somewhat variable in the ground color 
and amount of blackish dusting on forewing, some 
specimens being considerably paler than others; but 
the pattern markings are constant, the most conspicu- 
ous of which is the broad black subbasal band with 
roughened scales, formed by the borders of the ante- 
medial line. 

The female type of canescentella Hulst in the Rut- 
gers Collection lacks an abdomen, as does the matching 
male paratype in the National Museum (originally from 
the Fernald Collection). Both specimens are rubbed; 
but the pattern characters are plainly distinguishable 
and their palps, antennae, and venation are unmistak- 
ably those of Passadena. 


176 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Genus 88: Ulophora 


[Venational division B. Forewing with subbasal scale ridge; 
veins 4 and 5 approximate for a short distance from cell. Hind 
wing with vein 2 from well before angle; 4 and 5 stalked for about 
half their lengths; cell short (less than one-third the length of 
wing). Male antenna simple. Labial palpus uncurved, long, 
slender. Maxillary palpus squamous. Male genitalia with 
transtilla incomplete or absent; penis armed with a longitudinal 
row of short, slender spines.] 


88. Genus Ulophora Ragonot 


Ulophora Ragonot, Ann. Soc. Ent. France, ser. 6, vol. 10, Buil., 
p. vii, 1890; Monograph, pt. 1, p. 155, 1893.—Forbes, 
Cornell Mem. 68, p. 619, 1923. (Type of genus: Ulophora 
grotezz Ragonot.) 

Acromeseres Dyar, Ins. Insc. Menstr., vol. 7, p. 41, 1919. (Type 
of genus: Acromeseres dialithus Dyar. New synonymy.) 

Tongue well developed. Antenna simple; shaft 
somewhat broadly flattened towards base on male but 
without sinus or scale tuft or thornlike spines, on female 
threadlike, weakly pubescent on both sexes. Labial 
palpus upcurved, reaching well above vertex, slender, 
smoothly scaled; third segment about one-third the 
length of second, acuminate. Maxillary palpus squa- 
mous, more broadly so on male than on female. Fore- 
wing with subbasal ridge of raised scales; 11 veins; vein 
2 from before but rather near lower outer angle of cell; 
3 from the angle, at base nearer to 4 than to 2; 4 and 5 
approximate at base and for a very short distance 
beyond; 6 from below upper angle of cell, straight; 8 
and 9 stalked for slightly more than half their lengths; 
10 from the cell, nowhere approximate to the stalk of 
8-9; male without costal fold. Hind wing with vein 2 
from well before lower outer angle of cell; 3 from the 
angle connate with or very closely approximate to the 
stalk of 4-5 at base; 4 and 5 stalked for about half their 
lengths; 7 and 8 closely approximate for a short dis- 
tance from cell; cell slightly less than one-third the 
length of wing; discocellular vein curved, not appreci- 
ably produced at lower angle. Highth abdominal 
segment of male with a weak pair of ventrolateral hair 
tufts. 

Male genitalia with uncus triangulate, its apex 
narrowly rounded. Apical process of gnathos a stout, 
curved hook. ‘Transtill2 incomplete or absent (its 
elements, when distinguishable, very small and weakly 
sclerotized). Harpe simple; costa broadly but weakly 
sclerotized; clasper vestigial. Aedeagus straight; penis 
armed with a longitudinal row of short, slender, 
straight spines (16 to 20, the number individually vari- 
able). Vinculum stout, considerably longer than great- 
est width, but slightly tapered to rounded terminal 
margin. 

Female genitalia without signa; bursa with a mat of 
fine scobinations at and near junction of bursa and 
ductus bursae, otherwise smooth and membranous; 
ductus bursae longer than bursa, slender, tubular, 
unsclerotized, genital opening simple, narrow; ductus 
seminalis from middle of bursa. 


A distinct genus of uncertain affinities; contains one 
North American and one very closely related neo- 
tropical species. 


365. Ulophora groteti Ragonot 
Ficure 420 


Ulophora groteit Ragonot, Ann. Soc. Ent. France, ser. 6, vol. 10, 
Bull., p. vii, 1890; Monograph, pt. 1, p. 156, 1893.—Hulst, 
Phycitidae of N. Amer., p. 222, 1890.—Forbes, Cornell 
Mem. 68, p. 619, 1923.—McDunnough, Check list, No. 
6117, 1939. 


Ulophora tephrosiella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 107, 1904.—Forbes, Cornell Mem. 68, p. 619, 1923.— 
MeDunnough, Check list, No. 6118, 1989. (New synonymy.) 

Forewing brown dusted with white on basal and 
median areas and narrowly along terminal margin, 
giving the paler areas an ashy gray appearance and ac- 
centing the brown shade along costa and bordering the 
transverse lines; the latter narrow, faint, silvery scaled ; 
antemedial line at middle of wing, oblique, somewhat 
sinuate-dentate, bordered outwardly by a narrow brown 
line and inwardly by a broad brown band, the inner 
edge of which forms a stout raised-scale ridge extending 
from inner margin to top of cell, the inner margin rather 
strongly dusted with white; subterminal line sinuate, 
with rather broad inner and outer brown borders; discal 
dots obscure, more or less confluent, brown; the terminal 
dots confluent, forming a fine brown line along outer 
margin. Hind wing pale to dark smoky brown; the 
veins somewhat darker; a fine dark brown line along 
termen. Alar expanse, 11-19 mm. 

Male genitalia distinguished from those of the species 
following (guarinella) chiefly by the broader harpes and 
more narrowly rounded apex of uncus. The number 
and length of the spines on penis is an individual charac- 
ter. Our figure 420a shows the minimum in number 
and the maximum in length. Female genitalia like 
those of guarinella. 

Typr LocaLities: North Carolina (grote, in Paris 
Mus.); Washington, D. C. (ephrosiella, in USNM). 

Foon eiant: Tephrosia spp. Larvae feeding on pods 
and seeds. 

DistrisutTion: New Jersey, Woodbine (Sept.); Dis- 
trict of Columbia (July, Aug.); Virginia, Pendleton 
(Aug.), Skyland (July); North Carolina, Southern Pines 
(June, July, Aug., Sept.), Tryon (July, Aug.); South 
Carolina, Clemson College (July); Georgia, Atlanta 
(June), Spalding County (Nov.); Florida, Glenwood, 
Lake Alfred (May, July), Lakeland (Sept.), Polk 
County (Aug.), Tampa (June), Winter Haven (July); 
Alabama, Auburn; Louisiana, Vernon Parish (Aug.); 
Texas, Herne, Sandflat. 

Dyar’s tephrosiella was described from small speci- 
mens. ‘There are no structural or color differences to 
distinguish these from typical grotew. The larva lacks 
the sclerotized rings around seta IIb of mesothorax and 
seta III of the eighth abdominal segment, normally 
characteristic of phycitid larvae. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE Al¥pye 


366. Ulophora guarinella (Zeller) 
Ficures 421, 909 


Myelois guarinella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 
208, 1881. 

Ulophora guarinelia (Zeller) Ragonot, Monograph, pt. 1, p. 155, 
1893. 


Acromeseres dialithus Dyar, Ins. Insc. Menstr., vol. 7, p. 42, 1919. 
(New synonymy.) 

Forewing decidedly paler than that of grotei and, 
except for the band preceding the antemedial line, an 
almost uniform pale gray; the transverse lines obscure; 
antemedial line bordered inwardly by an orange-red 
band with more or less black shading on its lower half 
and a black scale ridge along its inner edge; subterminal 
line nearly obsolete, indicated chiefly by a very faint, 
narrow, dark gray inner border; discal dots more or less 
obscured, when distinguishable, blackish and either 
separated or coalesced; terminal dots faint, separated. 
Hind wing whitish with a faint brownish or smoky tint 
towards apex and along outer margin; the veins very 
slightly darkened. Alar expanse, 13-18 mm. 

Male genitalia are similar to those of groteii except 
for narrower harpes and a more bluntly pointed uncus. 
The female genitalia show no distinguishing characters. 


TypPE Locatitizes: Honda, Colombia (guarinella, in 
BM); Santiago, Cuba (dialithus, in USNM). 

Foop piant: Unknown. 

Distrisution: Cusa: Santiago (Jan., Oct.), Baracoa. 
Cotompia: Honda. 

In structure, color, and maculation Dyar’s dialithus 
is a perfect match to the male type of guarinella. The 
species is very close to groteii but apparently distinct. 
Nothing is known of its life history. 


Genus 89: Chorrera 


[Venational division B. Forewing smooth; veins 3, 4 and 5 equi- 
distant and approximate towards base. Hind wing with vein 2 
well before angle; 4 and 5 stalked for over half their lengths. 
Male genitalia with transtilla incomplete; aedeagus needlelike; 
penis unarmed; vinculum with pair of anteriorally produced 
lateral lobes from terminal margin. Female genitalia developed 
as two elongate, narrow, opposed plates, armed with short, longi- 
tudinally arranged spines; ductus bursae slender, globularly ex- 
panded near genital opening.] 


89. Genus Chorrera Dyar 


Chorrera Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330, 1914. 
(Type of genus: Chorrera idiotes Dyar.) 

Tongue well developed. Antenna weakly pubescent; 
on male a slight scale tuft on shaft shortly beyond basal 
segment, no appreciable sinus. Labial palpus obliquely 
ascending, reaching to slightly above vertex, slender; 
second segment roughly scaled; third less than one-half 
the length of second, acuminate. Maxillary palpus 
squamous. Forewing smooth; 11 veins; vein 2 from 
before but rather near lower outer angle of cell; 3 from 
the angle; 3, 4 and 5 equidistant and approximate 
towards base; 6 from below upper angle of cell, straight ; 


8 and 9 stalked for half or less than half their length; 
10 from the cell; male without costal fold. Hind wing 
with vein 2 from well before outer angle of cell; 3 from 
the angle; 4 and 5 stalked for over half their lengths 
(very long stalked in extrincica) ; 7 and 8 closely approxi- 
mate for a short distance beyond cell; cell less than one- 
half the length of wing; discocellular vein curved, con- 
siderably produced at lower angle. Eighth abdominal 
segment of male with a pair of very weak ventrolateral 
hair tufts. 


Male genitalia with uncus rather narrowly triangu- 
late, tapering abruptly at apex. Apical process of 
gnathos a short, stout hook. Transtilla incomplete, the 
divided elements small. Harpe short, with strongly 
sclerotized, thornlike, more or less appressed clasper, 
about half as long as harpe. Anellus a shallow U- or 
V-shaped plate with somewhat produced lateral lobes. 
Aedeagus, long, slender (needlelike), slightly curved; 
penis unarmed. Vinculum stout, about twice as long 
as greatest width; slightly tapering to a moderately 
broad terminal margin with a pair of anteriorly pro- 
duced lateral lobes. 

Female genitalia with signa developed as two 
elongate, narrow, opposed plates armed with short, 
stout, thornlike spines, longitudinally arranged; bursa 
otherwise membranous, large, more or less pear shaped; 
ductus bursae for half its length from bursa, very 
narrow, ribbonlike, sclerotized and bent at middle, 
globular and membranous beyond, and thence triangu- 
larly expanding into the simple genital opening; ductus 
seminalis from bursa near its junction with ductus 
bursae. 

The genus is certainly distinct and easily distin- 
guished from other phycetine genera by its peculiar 
genitalia. It contains what appear to be three tropical 
American species. These may eventually prove to be 
no more than races of a single variable species, but at 
present we are not justified in such a grouping. We 
know nothing of their life history and our knowledge of 
their distribution is too fragmentary to permit more 
than speculation as to their status. 


367. Chorrera idiotes Dyar 
Ficures 34, 422, 914 


Chorrera idiotes Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 331, 
1914. 

Forewing gray with a fine dusting of white scales, 
making the ground color a dark ashy gray; transverse 
lines narrow, faint, whitish; antemedial line slightly 
oblique, sinuate-angulate, with a narrow, obscured, 
outer edging of black scales; subterminal line somewhat 
more distinct, sinuate, with narrow inner and outer 
dark borders, pronounced and black at costa; discal 
dots very faint, when distinguishable, black and sepa- 
rated; a row of faint, confluent, blackish dots along 
terminal margin. Hind wing translucent white; a dark 
smoky shade along costa and a narrower smoky shade 
along terminal margin; the veins not darkened except 


178 


UNITED STATES NATIONAL 
(on some females) at their marginal extremities. Alar 
expanse, 15.5-19 mm. 

Male genitalia with a short knoblike projection from 
base of clasper of harpe. Lateral lobes of anellus 
short, stubby. Female genitalia as given for the genus. 

Typ tocauity: La Chorrera, Panama (May; type 
in USNM). 

Koop prant: Unknown. 

Known only from the type locality. 


368. Chorrera extrincica (Dyar), new combination 
Figures 423, 913 


Rhodophaea extrincica Dyar, Ins. Insc. Menstr. vol. 7, p. 41, 1919. 


Similar to idiotes except: Forewing with less whitish 
dusting; the transverse lines and their dark borders more 
obscure; ground color a nearly uniform brownish gray 
(fuscous). Hind wing with narrower smoky borders 
along costa and terminal margin; veins 4 and 5 longer 
stalked (4 very short and on some specimens vestigial or 
altogether absent). Alar expanse, 13-15 mm. 

Male genitalia with clasper of harpe a simple hook (as 
in postica); lateral lobes of anellus longer and more 
slender. Female genitalia with appreciably smaller 
bursa and entire genitalia shorter. 

TYPE LOCALITY: Santiago, Cuba (type in USNM). 

Foop prant: Unknown. 

Known only from the type locality (May, June, Oct.). 
Represented in the National Collection by 12 males and 
2 females. 


369. Chorrera postica (Zeller), new combination 
Fieure 424 


Myelois postica Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 213, 
1881. 


Nephopteryx postica (Zeller) Ragonot, Monograph, pt. 1, p. 292, 
1893. 


Similar to idiotes except: Thorax and basal two- 
thirds of forewing heavily dusted with white, making 
ground color an ashy white as far on forewing as the 
dark gray transverse shade extending from costa at 
beginning of subterminal line to outer third of inner 
margin. Hind wing without dark shade along costa 
and with only a faint, narrow, dark shade on inner 
Margin at apex. Alar expanse, 18 mm. 

Type LocaLity: Honda, Colombia (type in BM). 

Foop prant: Unknown. 

Known only from the type locality. 


Genus 90: Tacoma 


[Venational division B. Forewing with veins 4 and 5 stalked for 
nearly half their lengths. Hind wing with vein 2 near lower angle 
of cell; 4 and 5 long stalked; cell about one-third the length of 
wing. Antenna of male simple. Labial palpus upturned, slen- 
der. Maxillary palpus squamous. Male genitalia with apical 
process of gnathos a pair of converging bladelike projections; 
transtilla absent; harpe simple; penis unarmed. Female genitalia 
with signa developed as two plates armed with long, curved, 
clawlike spines; ductus bursae from middle of bursa, long, slender, 
membranous; ductus seminalis from bursa remote from junction 
of ductus bursae and bursa.] 


MUSEUM BULLETIN 207 


90. Genus Tacoma Hulst 


Tacoma Hulst, Ent. Amer., vol. 4, p. 115, 1888; Phycitidae of 
N. Amer., p. 139, 1890.—Ragonot, Monograph, pt. 1, p. 205, 
1893. (Type of genus: Tacoma feriella Hulst.) 

Tongue well developed. Antenna simple. Labial 
palpus upturned, cylindrical, slender, reaching to vertex; 
third segment one-third the length of second, acuminate. 
Maxillary palpus squamous. Forewing smooth; 11 
veins; vein 2 from before but rather near lower outer 
angle of cell; 3 from the angle; 4 and 5 stalked for 
slightly less than half their lengths; 6 from below upper 
angle, straight; 8 and 9 stalked for a half or a trifle more 
than half their lengths; 10 from the cell; male without 
costal fold. Hind wing with vein 2 from before but near 
lower outer angle of cell; 3 from the angle, connate with 
the stalk of 4-5; 4 and 5 long stalked (for over two-thirds 
their lengths); 7 and 8 closely approximate for a short 
distance from cell; cell about one-third the length of 
wing; discocellular vein curved and outwardly produced 
at lower angle of cell. Eighth abdominal segment of 
male simple. 


Male genitalia with uncus triangulate, its apex bluntly 
rounded. Apical process of gnathos a pair of converg- 
ing, flattened, pointed, bladelike projections. Trans- 
tilla absent. Harpe simple, narrowly elongate. Anel- 
lus a simple shield. Aedeagus simple, straight; penis 
unarmed. Vinculum stout, subtriangulate with nar- 
rowly rounded terminal margin; about as long as 
greatest width. 


Female genitalia with bursa shaped like an elongate 
potato, with ductus bursae arising from its middle; 
signa developed as two strongly spined plates, one of 
irregular shape armed with numerous, slender, long, 
curved spines and situated at junction of bursa and 
ductus bursae, the other a narrow curved band with a 
row of well spaced, strong, curved, clawlike spines 
along one edge; also in bursa, several (6 or more) small, 
detached, curved, thornlike spines; ductus bursae much 
longer than bursa, slender, membranous throughout; 
genital opening simple, small; ductus seminalis from 
bursa remote from junction of bursa and ductus bursae. 

A monotypical genus of unmistakable distinctness, 
easily identified by its genitalia. Except for their 
genitalia the sexes are not distinguishable on any 
external characters. The spining of the female abdo- 
men mentioned by Hulst (1890) is nonexistent. It is 
impossible to conceive what he saw or thought he saw, 
for there is no such structure on either the female or the 
male. 

370. Tacoma feriella Hulst 


Ficures 37, 425, 912 


Tacoma feriella Hulst, Ent. Amer., vol. 4, p. 115, 1888; Phycitidae 
of N. Amer., p. 189, 1890.—Ragonot, Monograph, p. 1, p. 
205, 1893.—McDunnough, Check list, No. 6142, 1939. 


Tacoma submedianella Dyar, Ins. Insc. Menstr., vol. 1, p. 34, 
1913.—McDunnough, Check list, No. 6144, 1939. (New 
synonymy.) 

Forewing gray, with an irregularly intermixed pow- 
dering of black and white scales, the black predomi- 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 179 


nating, making the over-all color dark gray with a faint 
bluish tint; the transverse lines grayish white, faint 
but distinguishable; antemedial line oblique, narrow 
from costa to lower margin of cell, thence (on most 
specimens) more or less expanded into a conspicuous 
white blotch of variable but usually quadrate shape; 
on some examples the white blotch is reduced and on a 
few completely absent; in the latter the antemedial is a 
narrow white line throughout, notched below costa and 
at lower fold and with a complete, narrow, outer, black 
border; on examples with the antemedial line expanded 
into a white blotch, the blackish outer border is broken 
or obliterated below the cell; subterminal line inwardly 
notched at veins 6 and 1b, shaded inwardly by a 
narrow, blackish border; discal dots, when distinguish- 
able, confluent, forming, on well marked specimens, a 
narrow black line along discocellular vein; terminal dots 
more or Jess confluent, usually forming a blackish line 
along outer margin. Hind wing whitish with a more 
or less smoky brown tint; veins faintly darkened and a 
narrow brown shade or line along outer margin. Alar 
expanse, 15-20 mm. 
Genitalia as given for the genus. 


Types.—In Rutgers University (feriella); U. S. 
National Museum (submedianella) 

Type Locauitins: Texas (feriella, in AMNH, ex 
Rutgers) ; La Puerta, Calif. (submedianella, in USNM). 

Foop piant: Mistletoe. This record from speci- 
mens from Riverside, Calif., reared by Commander 
Dammers. 

Distribution —Tezas, Blanco County (Sept.), Burnet 
County (Sept.), Kerrville, Plano (Aug., Sept.), Rio Frio 
(May); New Mexico; Mesilla Park (May); Arizona, 
Fish Creek Station (Sept.), Redington, Santa Rita Mts. 
(June), Wilgus; California, La Puerta (July), Mount 
Lowe (July), Riverside (July, Sept.). 

As indicated by the foregoing description the species 
is individually variable. Dyar’s submedianella was 
described from rather small California examples in 
which the characteristic white patch over the lower 
half of the antemedial line was partially or wholly 
obliterated by dark scaling. In the series before me 
there is every intergrade between the form with a 
conspicuous white patch and that without it; the 
extremes are not peculiar to any locality, nor do they 
exhibit any genitalic differences. 


180 


Group II 
[Hind wing with vein 3 present, 4 absent.] 


K&rys TO THE VENATIONAL DIVISIONS AND GENERA OF GROUP II 


Forewing with 11 veins; vein 2 from the cell . . . . . Venational division A (key, p. 180) 
Forewing with 11, 10, or 9 veins; vein 2 stalked or united with 3. 
Venational division B® (key, p. 184) 
Forewing with 10 veins; veins 8 and 9 ee 4 and 5 stalked; 2 from the cell or [Azaera] 
from the stalk of 4-5 . . ..... =... .  Venational division C (key, p. 184) 
Forewing with 10 veins; 8 and 9 stalked; 4 and 5 united; 2 from the cell. 
Venational division D (key, p. 185) 
Forewing with 9 veins; 8 and 9 united"; 4 and 5 united; 2 and 3 from the cell. 
Venational division E (key, p. 185) 
Forewing with 9 veins; 8 and 9 united; 4 absent; 3 and 5 stalked; 2 from the cell. 
Venational division F (key, p. 186) 
Venational division A 


1. Cell of hind wing partially open; only a fragment of the discocellular vein distinguishable. 
Psorosina (p. 208) 


Cell closed; discocellular vein complete and curved ............... 2 
2. Hind wing with vein 2 from lower outer angle ofcell ........2..2.2...8 
Hind wing with vein 2 from before the angle. .......... 8 
3. Forewing with vein 2 from the lower outer angle of wall: 2 emabnnte Meae Hoses (. 196) 


Forewing with vein 2 from before the angle ava ; 

4, Forewing with vein 2 from well before the angle; ein Albdamizel seamnent a wt 
without hair tufts; transtilla absent (elements not distinguishable); ductus bursae 
sclerotized for most of itslength ... . ... . . Ilatila (p. 263) 

Forewing with vein 2 from near lower angle; sielnian opdbmeadl segment of male with 
hair tufts; transtilla present Gameenied at least by Sh a: elements) ; 


ductus mene membranous. . . 2 sree 

5. Labial palpus upturned; maxillary wala a walle squamous; Sema a feansail a de- 
pressed, small cluster of blunt, stout, thornlike spies . . . Drescomopsis (p. 262) 

Labial palpus oblique; maxillary palpus 2 male in the form of an a signum or 
signa of female otherwise .. . 6 


6. Male genitalia with apical process ae eaminos on annie aie! hooting a sinelll anes 
signa of female consisting of a chain of thornlike spines on bulbous bases and a 


scattering of similarspines . . . . .. . . Cabotia (p. 200) 
Male genitalia with apical process of anailias a gett. ieae hook; signa of female 
consisting of large, round, densely spined plates... .....-...+.+--7 


9 The genus Nonia in this division could easily be confused on forewing venation with the species and 
genera of division F. However, in Nonia it is vein 2 that is absent (united with 3) and 4 and 5 that are 
stalked; while in group F vein 2 is present, 4 absent, and 3 stalked with 5, two radically different developments 
to the same end result. The genitalia of Nonia show its close relationship to forms with veins 2 and 3 stalked 
and vein 2 tending to disappear. 

10Tn occasional specimens of Vitula a vestige of vein 9 may be present on one forewing or the other, 
but the normal condition is for vein 9 to be absent. 


11 Some specimens of Bema show a trace of vein 9 on one side or the other, but this is an abnormal 
condition. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


7. Male antenna with a stout hook from basal segment of shaft; tufts on eighth abdominal 

segment of male compound; transtilla incomplete. . ..... Oncolabis (p. 199) 

Male antenna with shaft simple; eighth abdominal segment of male with paired tufts; 

transtila completes {exam Ae Qo. Ole ons aatels| De Honorinus (p. 199) 

8. Veins 3 and 5 of hind wing approximate at base (or contiguous for a short distance 

IexOViCl) eS * OE eS comes ee REN rte) ap artes were aeccee arn CCRT (25. | 9 

Veins 3 and 5 of hind wing connate (rarely very shortly stalked)... ...... 15 

Veins 3 and 5 of hind wing appreciably stalked. . ............2.2.. 24 

0.2 Maxillary palpus!squamoush Saver a? HERA iO emo inode ww sige: yao 

Midxillary: palpus«filiform™ fein... .Oha 80% enohei ane & gieveliin.geivew. 12 

10. Labial palpus upturned; shaft of male antenna with sinus and scale tuft at base . .11 

Labial palpus porrect; shaft of male antenna simple. ....... Patriciola (p. 209) 

11. Forewing with vein 10 stalked with 8-9; eighth abdominal segment of male with com- 

pound! ventrallituitsy, Pars HO Sl aries. Bilge’? vn. Canarsia (p. 201) 

Forewing with vein 10 from the cell; eighth abdominal segment of male with paired 

ventrolateral ttufts@ TiRG% 10 ort 20). beeccrohaia 308 Teciey. Paconius (p. 210) 

12. Apical process of gnathos bifid or produced as large, partially fused lobes; ductus bursae 

of female sclerotized only at genital opening . ............2... 13 

Apical process of gnathos a short, blunt hook; ductus bursae sclerotized for most of its 

length SO FReRIOVIO Die geen Je loletog. Uso old arost Ani awed’ eo. 14 

13. Antennal shaft of male simple; ductus seminalis of female from middle of bursa; signum 

BbsenG) SH SLND Dc a meh Yr Pet og er BILE BULA CRO, ae Aptunga (p. 211) 

Antennal shaft of male with some rough scaling in a shallow sinus towards base; ductus 

seminalis from anterior end of bursa; signum present .... . Baphala (p. 235) 

14. Labial palpus porrect; eighth abdominal segment of male simple . . Volatica (p. 290) 

Labial palpus oblique; eighth abdominal segment of male with paired dorsal tufts. 

Vezina (p. 291) 

15, Antennae bipectinate (in both sexes) ; veins 7 and 8 of hind wing closely approximate. . 16 

Antennae pubescent; veins 7 and 8 of hind wing anastomosed. . ......... 17 

16. Labial palpus of male porrect; bursa of female without signum . . . Melitara (p. 240) 

Labial palpus of male obliquely ascending; bursa of female with signum . Olycella (p. 241) 

17. Veins 7 and 8 of hind wing anastomosing beyond cell for not more than half the length 

Ofsvenn’ 72511)! Uy Teme ee ae eae ate lo, wees la@eahnn wileliosysolalt. . 18 

Veins 7 and 8 anastomosing for most of their lengths beyond cell. . . . ..... 21 

18. Cell of hind wing not more than half thelength of wing. ............ 19 

Cell of hind wing definitely more than half thelength of wing .......... 20 

19. Maxillary palpus filiform; antennal shaft of male with modified, papillalike setae on basal 

segments; female withsignum .......2...2..2... Zophodia (p. 238) 
Maxillary palpus squamous; antennal shaft of male simple; female without signum. 

Olyca (p. 243) 

20. Maxillary palpus squamous; male with transtilla complete, forewing without costal 

Folds AR NUR eae Mea BOSC, Chl Lees 5 Ona Euzophera (p. 272) 

Maxillary palpus filiform; male with transtilla incomplete, forewing with costal fold. 

Eulogia (p. 275) 

2ishabialipalpus' upturned! OuhaMnt (io Asie), WDE Ames eh ep Geet loekneeliok . 22 

Labial palpus oblique or obliquely ascending. . ...........0.52285 23 

22. Forewing with vein 10 shortly stalked with 8-9; male with transtilla incomplete, but 

elements considerably enlarged. . . ...........4. Moerbes (p. 268) 

Forewing with vein 10 from the cell (usually separated, rarely connate, at base, with 

8-9); male with transtilla complete. .......2.2... Moodnopsis (p. 269) 

23. Cell of hind wing less than half (more than one-third) the length of cell; forewing of male 


with costal fold; eighth abdominal segment of male with paired ventrolateral hair 
tufts, |. . REN RR pow to Ru.BOLOR GAO, Bots Entmemacornis (p. 266) 
Cell of hind wing more than half the length of cell; forewing of male without costal fold; 
eighth abdominal segment of male simple .......... Exuperius (p. 274) 


181 


182 


24. 


25. 


26. 


27. 


28. 


29. 


30. 


31. 


32. 


33. 


34. 


35. 


36. 


37. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Hind wing with cell one-third the length of wg. ......... Edulica (p. 271) 
Hind wing with cell approximately half the length of wmg. ........... 25 
Penis of male armed with two stout thornlike cornute; signa of female consisting of two 


large, opposed plates armed with long stout spines, a scattering of similar spines 
between the plates, and a band of shorter spines at junction of bursa and ductus 


bursae.. 3 SAAD APUG RN 00, oe, SO Ae eve Adelperga (p. 187) 
Penis of male smooth or armed only with weak spines or sclerotized wrinklings; signa of 
female (if present) otherwise; iacthiieny Hiasoenene yaihae de ie ee eae 26 
Forewing with vein 2 from lower outer aele of cell... .scocieupe sued eee 2 27 
Forewing with vein 2 from before the angle ...........2.2.4..... 28 


Forewing with veins 2 and 3 closely approximate at base; males with strong costal fold on 
forewing, eighth segment with paired ventrolateral hair tufts, transtilla complete. 
Cassiana (p. 212) 
Forewing with veins 2 and 3 connate; male without costal fold, eighth abdominal segment 
without hair tufts, transtilla complete. ............ Rioja (p. 267) 
Hind wing with veins 7 and 8 anastomosed for most of their lengths beyond cell. . . . 29 
Hind wing with veins 7 and 8 approximate, contiguous or anastomosed beyond cell; but if 
anastomosed, then for less than two-thirds the length of vein7........ 31 
Forewing with vein 10 stalked with 8-9. ............ Cayennia (p. 267) 
Forewing with vein 10 from the cell, separated at base and divergent from stem of 8-9 . 30 
Male forewing with costal fold; eighth abdominal segment of male simple; female genitalia 
with ductus seminalis from bursa between signum and ductus bursae. Selga (p. 265) 
Male forewing without costal fold; eighth abdominal segment of male with paired ventro- 
lateral hair tufts; female genital with ductus seminalis from anterior end of 
bursa .. .. .okpeu. alu to duensies lininobds Mia sasinoe Anderida (p. 211) 
Male genitalia with apical process of Briathos greatly reduced and fused into subanal 
plate; female genitalia with signum a single, long, stout, curved spine. 
Lascelina (p. 264) 
Male genitalia with apical process of gnathos well developed; female genitalia with sig- 


num (if present) otherwise .*.)) 020.) ee eee we a ale. 32 
Male genitalia with apical process of gnathos developed as a stout, elongate hook; female 
genitalia with ductus seminalis from ductus bursae. ........2.... 33 


Male genitalia with apical process of gnathos usually flanged or lobed, frequently bifid, 
sometimes fused or partially fused, if completely fused (Cahela, Rumatha) consider- 
ably enlarged SESE female pominle with ductus seminalis from bursa copulatrix. 


40 

Hapa palpi jporrect enor eins: oo tele dete ene eee atis/lod ventana & 34 
Labial palpi oblique or upturned . . . . ~~... 2 ee eee eee ee wes 38 
Highth abdominal segment of male with ventrolateral hair tufts and lateral eversible 
HOES ee ee se AES Toh OM a Sey foe) cr se Ae eae mea 35 
Eighth abdominal segment of male without tufts or lobes, ventrolateral hair tufts only 
(no eversible lobes)... .... AU MLE ie ricco Geo MRI SNE cb eS - Cet ex%h'c 36 
Maxillary palpus of male filiform. ............. Macrorrhinia (p. 190) 
Maxillary palpus of male in the form ofanaigrette. ......... Ocala (p. 191) 
Aedeagus of male of nearly even width throughout; female genitalia with ductus bursae 
simple, and ductus seminalis from ductus bursaenearitsmiddle ........ 37 


Aedeagus of male abruptly tapered from base and very slender therefrom; female geni- 
talia very slender for half its length from genital opening, and thence to bursa 
swollen and spiraled, armed throughout the spiraled portion with a line of fine, sharp 
BPINES WOM: «, s. a. s,s tee sy oy op AMOReEO< Roots pene Valdivia © 191) 

Kighth abdominal segment of male with a pair of short ventrolateral hair tufts; harpe 
with clasper present, developed as a strongly sclerotized, nearly straight hook; Gamal 
genitalia with ductus bursae as long as, or longer than, bursa. . . Eumysia (p. 187) 

Eighth abdominal segment. of male without hair tufts; harpe without clasper; ductus 
bursae considerably shorter than bursa. . .......... Protasia (p. 193) 


38. 


39. 


40. 


41. 


42. 


43. 


44. 


45. 


46. 


47. 


48. 


49. 


50. 


51. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Maxillary palpus of male squamous; female genitalia with an acutely bent, sclerotized 


elbow in ductus bursae before genital opening. ......... Hulstia (p. 195) 
Maxillary palpus of male filiform or in the form of an aigrette; female genitalia with 
duchus bursacistraightmyin ye meee. Nem eos Cleese wlio, nilasinon oll . 39 
Maxillary palpus of male filiform; female genitalia with ductus bursae long (as long or 
longer -thantbursa) loge anne, Oly i) SaBotTR) wees Heterographis (p. 193) 
Maxillary palpus of male in the form of an aigrette; female genitalia with ductus bursae 
shorti(much'shorter than’ bursa)iiyili-ieteedesey es ee Staudingeria (p. 194) 
Tongue considerably reduced, exposed or completely hidden by labial palpi . . . .41 
Tongue well developed and exposed.) WS" 20. NON S RpY Ow Raggio. , 43 


Hind wing with veins 7 and 8 anastomosed for at least half their distances from cell; 
female genitalia with ductus seminalis from anterior end of bursa copulatrix. 

Laetilia (p. 230) 

Hind wing with veins 7 and 8 very shortly anastomosed (for less than half their distances 

from cell); female genitalia with ductus seminalis from bursa near its junction with 


ductus*bursae% 24 os = MAY Sho Le Sopte aay Mane pital bo te? , 42 
Male genitalia with apical process of gnathos partially fused; female genitalia with 
SISTA Oe AIRE DILL) OLS 1) Soe A PaaS Cactoblastis (p. 245) 


Male genitalia with apical process of gnathos bifid; female genitalia without signum. 
Eremberga (p. 253) 


Mieixill aneya Al MSPETLOMUN voy co ws iee 5) aso) lay cis) ce! ea ah “Fes Semen heh Fomeepe Subd alemaey ot Boy ot 44 
Maxillary palpus squamous, fan shaped or flamboyant ..........2... 45 
Labial palpi of both sexes porrect; eighth abdominal segment of male simple. 


Rhagea (p. 237) 
Labial palpi of male upturned, of female oblique; eighth abdominal segment of male 


with paired ventrolateral hair tufts. ........... Cactobrosis (p. 260) 
Maxillary palpus flamboyant (extending above front). ...... Sigelgaita (p. 255) 
Maxillary palpus fan shaped and held vertical to the face. . . .. Yosemitia (p. 250) 
Maxillary palpus: squamous « «+ oo: e06 ws SEY YS Oo Roy Ca pyroqon » 46 


Shaft of male antenna pubescent and with a series of modified papillalike setae on several 
basal segments; female with labial palpus obliquely ascending . . Ozamia (p. 257) 
Shaft of male antenna pubescent (but without modified setae on basal segments) or 
pectinate; female with labial palpus porrect or obliquely porrect . ...... 47 
Vinculum of male long (distinctly longer than broad) ; female genitalia without signum. 48 
Vinculum short (little, if any, longer than broad); signum present ........ 50 
Maxillary palpus extending well above middle of face; eighth abdominal segment of male 
SUING Ve ter ir eR RE EN Scr. Wi cat Nanaia (p. 245) 
Maxillary palpus not extending above middle of face; eighth abdominal segment of male 
Withyventrolateral ham itifisgss webie AioelVsuisins: serusswsturn sf ideue a thustienee eater ° 49 
Antenna of male pubescent; ductus seminalis of female from near anterior end of bursa; 
bursaveryismiallis raieeesik See ee LO Ss ee SE EIN Salambona (p. 254) 
Antenna of male bipectinate; ductus seminalis of female from middle of bursa; bursa 
lareeig wrt MRR Ley nee 1s el ce SS AL iy, a, Amalafrida (p. 256) 
Antenna of male unipectinate; (female unknown) ........ . Parolyca (p. 255) 
Antenna of male bipectinate; ductus seminalis of female from middle of bursa. 
Alberada (p. 244) 
Antenna of male pubescent; ductus seminalis from bursa towards (near) junction of 
bursa with ductussbursnewe vis eit) Fe Le to ee dn tatu Hebe tat 51 
Habitat, South America; male genitalia with apical process of gnathos bifid. 
Tucumania (p. 252) 


_ Habitat, southwestern United States and northern México; male genitalia with apical 


process\of gnathos uscd ee AL iin ha paie, chink anny © cios diies entwehait « 52 


183 


184 


52. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Male genitalia with aedeagus moderately long and sclerotized throughout, harpe without 
subbasal sclerotized pocket, anellus with base of plate narrowly sclerotized; female 
with dark markings on forewing longitudinal, no discal spot . . . Cahela (p. 247) 

Male genitalia with aedeagus short and partially sclerotized, harpe with subbasal sclero- 
tized pocket, anellus with base of plate broadly sclerotized; female with dark mark- 
ings of forewing transverse (at least in part), discal spot prominent. 

Rumatha (p. 248) 


Venational division B 


. Forewing with veins 2 and 3 of forewing stalked see 
Forewing with veins 2 and 3 of forewing united. . 6 
. Forewing with veins 4 and 5 stalked. . +8 
Forewing with veins 4 and 5 united . 5 8 
. Forewing with veins 8 and 9 stalked. Lie cceucsie FF fo way Ciara acl ae anes e cy regenera 
Forewing with veins 8andQ9 united. ......... . . . Micromescinia (p. 277) 
. Cell of forewing open (no trace of disease vein). . .. . . . Gennadius (p. 277) 
Cell of forewing closed . ..........2... . . . Mescinia (p. 212) 


. Forewing with veins 8 and 9 stalked; hind wing with 3 and 5 iptolleed: 


Eurythmasis (p. 203) 
Forewing with veins 8 and 9 united; hind wing with 3 and 5 closely approximate at base. 
eirephonyegints © on 


, Hindi wing with yeins)3 and aystalked -)) sq-Uem ane y-li se ieciielsinlii mele 


Hind wing with veins 3 and 5 closely approximate or connate at base. . .... . : 
. Cell of forewing open (no trace of discocellular vein) . . .. . . . Farnobia (p. 276) 
Cell of forewing closed . ...-.. 2... +22 - 2s e Ge Sige test wpa t att gee 
. Cell of hind wing open (no trace of discocellular vein). . . . . . . Wunderia (p. 204) 
Cell of hind wing closed. ............ ... . . .Hurythmidia (p. 204) 
. Forewing with veins 8 and 9 stalked... ....... =... . . Phestinia (p. 216) 
Forewing with veins 8and9 united. ........=...... . Nonia (p. 215) 
Venational division C 
. Hind wing with discocellular vein straight and vertical . . . . . . . BY, Sea, 
Hind wang with’ discocellular vem curved: s.) -ete) -n sien eee cee et rauem 
Horewine with) veins 4 and orstalkcedss 5). ee) sien tere mete nny nts) ee eee 
Forewing with veins 4and 5connate.............. Patagonia (p. 225) 
. Male genitalia with a comb of strong spines along side of aedeagus; female genitalia with 


two large signa composed of opposed, cup-shaped, strongly spined, concave plates; 
ductus seminalis from ductus bursae. .........-... Rotruda (p. 225) 
Male genitalia with aedeagus smooth or with only minute scobinations at apex; female 
genitalia with signum a single, coarsely spined plate, or absent; ductus seminalis 


fromm bursa.’ Oyen rita En hiak aia SSE L are MPA reat sy te goed HgnivedsOmnA (p. 219) 

. Hind wing with veins 3 and 5 approximate at base. .... . Pataca are sory itty scat te 5 
Hind wing with veins 3 and 5 connate orstalked. ..............-.7 

. Cell of hind wing one-third the length of wing Breed iin) TELE ORE SRE tee 6 
Cell of hind wing one-fourth the length of wing. .......... Gvenna (p. 288) 

. Eighth abdominal segment of male with compound diossall tufts; shaft of male antenna 
SIMPLE ce ee eee. nee, Nhe Sty eae Tet Cot Baran ae ce Ree ata Vitula (p. 285) 
Eighth abdominal segment of male simple; shaft of male antenna with shallow smus 
towardstbases tee Tee gee ay SRE Dinteeiee eo, erie haere eee Moodnella (p. 289) 

. Hind wing with vein 2 oe the lower outer angle of cell. . . . . Harnocha (p. 202) 
Hind wing with vein 2 from before theangle. .........-+-+-+-+-+-+-+-s 8 

. Hind wing with vein 2 near lower outer angle of cell. . . 2... . +++ ---- 9 


Hind wing with vein 2 from well before the angle... .........2-.--. 10 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 185 


9. Forewing with vein 3 separate from, connate with, or rarely (in individual specimens) 


very shortly stalked with the stem of 4-5 . . . . . .Ephestiodes (p. 278) 
Forewing with vein 3 from the stalk of 4-5 (well stalked maith it) . . . Azaera (p. 282) 
10. Cell of hind wing approximately one-third the length of wing. . . 2478) Tal 


Cell of hind wing slightly more than one-half the length of wing . . Vagobanta (p. 289) 

11. Male with transtilla incomplete; shaft of antenna with shallow sinus towards base. 
Moodna (p. 283) 

Male with transtilla complete; shaft of antenna simple. . . . . . Manhatta (p. 287) 


Venational division D 


1. Hind wing with discocellular vein straight and vertical; cell very short (about one-fifth 


the length of wing) . . . . Metephestia (p. 264) 

Hind wing with dintdealittanevein Gaeved: call tonger ania slightly less than one-third 
to one-half the length of wing) . SM tee eee ee ae et ee 

2. Hind wing with vein 2 from lower outer ariel of cell Pili re Cee ale at ee ote: 
Hind wing with 2 from before the angle. ... . . 4 


3. Veins 7 and 8 of hind wing contiguous for some chesney hepend eet (but sa: werent 
sing) ; labial palpus upturned; antennal shaft of male simple . Oedothmia (p. 205) 

Veins 7 and 8 anastomosed beyond cell; labial palpus oblique; shaft of male antenna 
with a hook from its basal segment and deeply curved for several segments beyond. 
Stylobasis (p. 205) 


4, Hind wing with vein 2 from near the angle ofcell. . 2. 2. 2... 2 ee 
Hind wing with vein 2 from well before the angle. ....... 6 

5. Labial palpus porrect; male penis without armature. .. . tes cD iaitiasts es 189) 
Labial palpus upturned; penis armed with two stout, straighit cornuti. Diviana (p. 206) 

6. Hind wing with veins 3 and 5 shortly stalked. . . . . . . . Prosoeuzophera (p. 275) 
Hind wing with veins 3 and 5 connate or very closely approximate at base. . . . . 7 

7. Labial palpus oblique, extending above vertex of head; veins 7 and 8 of hind wing closely 
approximate for half their distance beyond cell. . . . . . . . Palatka (p. 207) 

Labial palpus upturned, short, not reaching vertex; veins 7 and 8 anastomosed for most 

of their distance beyond cell. . . . . . . 2... . . . Cacozophera (p. 208) 


Venational division E 


1. Hind wing with discocellular vein straight and vertical. . . . . .. . 2 
Hind wing with discocellular vein straight and oblique. . ...... Beay (pr 217) 
Hind wing with discocellular vein curved. . . Sako 


2. Cell of hind wing less than one-fourth the wing length! Gali ious dude fold. on fore- 
wing, with a weak pair of ventrolateral hair tufts on eighth abdominal segment. 

Comotia (p. 217) 

Cell of hind wing about one-third the wing length; male with costal fold, but with eighth 


segment SIMplO oes wwe news Unaaiitia Grr) 
3. Hind wing with vein 2 from near foere Peer snes ‘of ene . . . .  Nicetiodes (p. 304) 
Hind wing with vein 2 from well before angle of cell. . . . 4 


4, Male with apical process of gnathos enlarged (broadened), ped ted: septic or fonaede 
female with ductus seminalis from bursa near its junction ae ductus bursae. 

Sosipatra (p. 294) 

Male with apical process of gnathos otherwise; female with ductus seminalis from bursa 

well removed from the junction of bursa ate ductus bursae. . . . 5 

5. Labial palpus upturned; ductus bursae of female membranous for a or ‘all of its 

length, not flattened. . . 6 

Labial palpus oblique or parece actus ipa of feats peleronized fOr a eonsidersbio 

Pant Onis lenrthvandanatroneds: 4. 2 secttucetae © easy. eee ee eet 


300329—56——_13 


186 


10. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


. Male with transtilla complete or its elements greatly enlarged and approximate at their 


apices; forewing with a strong costal fold; eighth abdominal segment with com- 
pound dorsal tufts; female with apophyses of ovipositor and intersegmental area 
between ovipositor and eighth-segment collar, short. . . . . . Ephestia (p. 301) 
Male with transtilla incomplete, its elements well separated at their apices, not enlarged; 
forewing without costal fold; eighth segment with paired dorsal hair tufts; female 
with apophyses of ovipositor and intersegmental area between ovipositor and 


eighth-segment collar, verylong. .........2.+.2... Anagasta (p. 299) 

. Male genitalia with apical process of gnathos asec’: ductus seminalis of female 
from very near anterior end of bursa. . ............ Ribua (p. 297) 

Male genitalia with apical process of gnathos svmmetmieal: ductus seminalis of female 
from middle or near middle of bursa. . . .......2-.-..2.+-..4:+-.-- 8 

. Labial palpi of both sexes porrect. .......-..2...-. . . . . Plodia (p. 298) 
Labial palpi of both sexes oblique. . .......2.2.2..4.++++e48020448-8 9 

. Tongue reduced; alarexpanse8mm.. ........... ‘ Caan (p. 294) 
Tongue well devdlaei: alar expanse 10mm.ormore. . . . area 1(0) 
Bursa of female with signum (antenna of male with a sheila: sinus in | ghett jonas 
DESC), koe tin Biyeye tic BeBe 3 Ae ass te iis . . Caudellia (p. 292) 
Bursa of ennel’s without Siac, Gosia imino) - SRY eo . . Bethulia (p. 296) 


Venational division F 


. Male genitalia with a projecting spur from base of costa of harpe; apical process of gna- 


thos U-shaped; female with ductus bursae sclerotized and flattened for most of its 
lenethy ee 2 oe se ye) SRR BEEET FAT OUW. BEOSD AUR cranes Varneria (p. 305) 
Male genitalia with ane! pela from costa of harpe near or -teyand middle; apical process 
of gnathos (if present) not U-shaped; female with ductus bursae membranous and 
tubular throughout... ............. 2 


. Male with eighth abdominal segment simple; gnathos ammina fae | ma very dee cou 


narrowly forked projection; aedeagus with bluntly pointed apex; penis pened sili 
an elongate, spiraled, suleviiiza and spined plate; female genitalia with signa ar- 
ranged as a series of (3 to 6) detached, broadly based, thornlike spines and narrow- 
bladed disks situated near anterior end of bursa and a cluster of several smaller 
spines near junction of bursa and ductus bursae. . . . . . . Eurythmia (p. 306) 
Male with compound dorsolateral tufts on eighth abdominal segment; gnathos without 
apical projection, its arms fusing at their distal ends into the sclerotized subanal 
plate; aedeagus with apex enlarged and flaring; penis unarmed except for a few sclero- 
tized wrinklings; signa of female consisting of a single elongate series of (5 to 14) 
thornlike spines... .....-++-++.-. ....... . Erelieva (p. 308) 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Genera 91 and 92: Adelperga and Eumysia 


[Venational division A. Forewing with 11 veins; 10 from cell, 
8 and 9 stalked, 2 and 3 from the cell, 4 and 5 stalked or connate. 
Hind wing with veins 7 and 8 closely approximate or contiguous; 
2 from close to lower, outer angle of cell; discocellular vein 
curved. Harpe of male genitalia with an elongate, strongly 
sclerotized clasper; gnathos terminating in a stout, short or 
moderately long, hooked process; transtilla incomplete; vinculum 
stout, as broad or broader than long.] 


91. Adelperga, new genus 


Tyrer or Genus: Heterographis cordubensiella Ragonot. 

Tongue well developed. Antenna simple, pubescent. 
Labial palpus obliquely upturned, reaching vertex, 
third segment short, projected forward. Maxillary 
palpus squamous. Forewing with vein 2 from or from 
very near outer angle of cell; 2 and 3 approximate at 
base; 4 and 5 stalked; 6 straight from below upper 
outer angle of cell; 10 from cell, closely approximate to 
8-9; male without costal fold. Hind wimg with vein 2 
from very close to angle of cell; 3 and 5 stalked; 7 and 8 
approximate; cell less than half (more than one-third) 
the length of the wing; discocellular vein curved. 
Eighth abdominal segment with a pair of short ventro- 
lateral hair tufts. 

Male genitalia with apical process of gnathos a rather 
short hook. Uncus broadly and bluntly rounded at 
apex. Harpe with a decided incurvation between sac- 
culus and cucullus; clasper present, developed as a 
long, strongly sclerotized and apically curved and 
swollen arm, projecting into the incurvation between 
sacculus and cucullus. Anellus semitubular. Aedea- 
gus long, stout, cleft and strongly sclerotized towards 
apex; penis armed with a pair of short stout thornlike 
cornuti and numerous granulations. 

Female genitalia with bursa copulatrix thickened and 
sclerotized at junction with ductus bursae; signa 
strongly developed, consisting of two large, opposed 
plates armed with long, stout spines, a scattering of 
similar spines between the plates, and a band of shorter 
spines where ductus and bursa join; ductus bursae 
short, flattened, very broad and sclerotized throughout; 
genital opening very large; ductus seminalis from 
bursa near junction with ductus bursae. 

This genus is easily identified by its genitalia. It 
shows striking affinities to Passadena of group I in shape 
of harpe, with incurvation between sacculus and cucul- 
lus, and in the development of an enlarged, strongly 
sclerotized and projecting clasper. Passadena also has 
opposed signa similarly spined. 


371. Adelperga cordubensiella (Ragonot), new combination 
Ficures 429, 772 

Heterographis cordubensiella Ragonot, Nouv. Gen., p. 30, 1888. 
Hulstia cordubensiella (Ragonot), Monograph, pt. 2, p. 128, 1901. 

Forewing with color and markings of Huzophera semi- 
funeralis (especially the pale color form of its synonym 
aglaeella Ragonot; see p. 273); the antemedial line far 
out (at or very near middle of wing), nearly vertical, 
slightly notched at top of cell and at lower fold, white 


187 


bordered outwardly by a black line; subterminal white 
line sinuate, incurved at vein 6 and the lower fold, 
bordered inwardly by black; the two transverse lines 
rather close (as in typical Huzophera) and the space 
between them dusted with blackish scales; otherwise 
the ground color of the wing is ash gray strongly shaded 
with reddish ocherous, especially in the enlarged basal 
area; a blackish spot on inner margin near base; a 
smaller blackish spot on costa just beyond the sub- 
terminal line and a row of black dots along termen. 
Hind wing whitish, shading to pale smoky fuscous 
towards apex and outer margin. Alar expanse, 15-18 
mm, 
Genitalia with characters as given for the genus. 


Type Locauity: Cérdoba, Argentina (type in Paris 
Mus.). 

Foop prant: Unknown. 

Distrisution: Argentina: Cérdoba, Las Vasquez, 
Tucuman. 

The figure in the Ragonot Monograph (pl. 22, fig. 20) 
is misleading. It shows the hind wing much too dark 
and uniformly colored, and shows none of the strong 
blackish dusting in the area between the antemedial 
and subterminal lines. 


92. Genus Eumysia Dyar 


Eumysia Dyar, Ins. Insc. Menstr., vol. 18, p. 220, 1925. 
of genus: Yosemitia mysiella Dyar.) 

Tongue well developed. Antenna strongly ciliate in 
male (cilia about three times the width of shaft), simple 
in female. Labial palpus porrect, long and beaklike 
(projecting about three times the length of the head 
beyond it). Maxillary palpus minute, filiform. Fore- 
wing smooth; 11 veins; vein 2 from before angle of cell; 3 
from the angle; 4 and 5 very shortly stalked or con- 
nate; 6 from below upper angle of cell, straight; 10 from 
cell, approximate to stalk of 8-9 at base; male without 
costal fold. Hind wing with vein 2 close to lower outer 
angle of cell; 3 and 5 stalked; 7 and 8 closely approxi- 
mate or contiguous for some distance from cell; cell 
slightly less than one-half the length of wing; disco- 
cellular vein curved. Eighth abdominal segment of 
male with a pair of short, weak hair tufts. 

Male genitalia with gnathos terminating in an 
elongate hook. Uncus narrowly and bluntly rounded 
at apex. Harpe with lower margin evenly curved, no 
incurvation between sacculus and cucullus; clasper 
present, developed as a strongly sclerotized, nearly 
straight hook. Anellus a simple plate. Aedeagus 
short, straight, moderately slender; penis unarmed. 
Vinculum broad, short, truncate. 

Female genitalia with bursa and ductus bursae 
simple; ductus seminalis from middle of ductus bursae. 

The genus was originally erected for nine species 
which Dyar removed from Zophodia and Yosemitia. 
As here restricted it includes only three of these (mysi- 
ella, maidella, and fuscella), a fourth species (pallidi- 
pennella), which Dyar had referred from Yosemitia to 
Zophodia, and a new species from Washington State. 


(Type 


188 UNITED STATES NATIONAL MUSEUM BULLETIN 207. 


There are no structural differences in genitalia that can 
be used to distinguish these supposed species. The 
form of the clasper of the harpe distinguishes Humysia 
from any other genus in group II. A similar develop- 
ment of the clasper occurs in Nephopteryx of group I. 
In general habitus Humysia most resembles Ragonotia 
in the Anerastiinae. 


372. Eumysia mysiella (Dyar) 
Fieures 430, 920 


Yosemitia mysiella Dyar, Proc. Ent. Soc. Washington, vol. 7, p. 
36, 1905.—Barnes and McDunnough, Contributions, vol. 3, 
p. 199, 1916. 

Eumysia mysiella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221, 
1925.—McDunnough, Check list No. 6295, 1939. 

Forewing white dusted with blackish fuscous, making 
the general color a light ashen gray; antemedial white 
line sinuate, nearly vertical, bordered inwardly from 
inner margin to top of cell by a broad ocherous bar, 
this bar preceded by a blackish margin, especially 
towards inner margin; similar blackish scaling out- 
wardly bordering the antemedial white line, sometimes 
faint, but usually distinct towards costa; subterminal 
line obscure, more or less shaded inwardly by dark 
scaling; blackish discal dots (especially the one at lower, 
outer angle of cell) usually distinct. Hind wing semi- 
hyaline, white. Alar expanse, 21-27 mm. 

Genitalia with characters as given for the genus. 

TypE Locauity: Stockton, Utah (type in USNM). 

Foop prant: Unknown. 

Distrisution: Utah, Stockton (Sept.), Hureka 
(June); Arizona, Phoenix (Aug.), Tempe (Aug.), Red- 
ington; New Mexico, Las Vegas (July), Jemez Springs 
(July). 

373. Eumysia maidella (Dyar) 
Figure 431 


Yosemitia maidella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 36, 1905. 

Eumysia maidella (Dyar), Ins. Inse. Menstr., vol. 13, p. 221, 
1925.—McDunnough, Check list No. 6297, 1939. 

Similar to mysiella but averaging larger, with dark 
markings more intensified along the veins and bordering 
antemedial and subterminal lines, ocherous subbasal 
bar fainter and in some specimens absent. Alar ex- 
panse, 27-32 mm. 

Genitalia similar to those of mysvella. 

TYPE LOCALITY: Stockton, Utah (in USNM). 

Foop Piant: Unknown. 

Distrisution: Unitep Starrs: Utah, Stockton 
(June, July), Eureka (June), Cedar Mts. (Iron County, 
July); Colorado, Fort Collins; Arizona (Sept.); New 
Mexico, Fort Wingate (May); California, Loma Linda 
(Aug.). Canapa: British Columbia, Kaslo (Aug.). 

Probably not specifically distinct from mysvella. 
There are no structural differences between them. The 
character given by Dyar, costa of male forewing ‘‘con- 
cave” (maidella) or ‘‘not concave’ (mysiella) is purely 
imaginary. Some females as well as some males of 
both ‘‘species” appear to have the costa slightly concave 
at middle but this is an optical illusion due to imperfect 


flattening of the wings in their spreading. The differ- 
ences in size and intensity of markings are slight. How- 
ever, since we know nothing of the biology of any 
Eumysia and since such differences may coincide with 
differences in host plants it seems advisable to keep the 
names separated for the present. 


374. Eumysia pallidipennella (Hulst), new combination 


Volusia pallidipennella Hulst, Canadian Ent., vol. 27, p. 56, 1895. 

Trivolusia pallidipennata Hulst, U.S. Nat. Mus. Bull. 52, p. 438, 
1903 (misspelling). 

Yosemitia pallidipennella (Hulst) Barnes and McDunnough, 
Contributions, vol. 3, p. 199, 1916. 

Zophodia pallidipennella (Hulst) McDunnough, Check list, No. 
3606, 1939. 

Paler and averaging smaller than mysiella with dark 
dusting fainter, giving the wing a more ocherous gray 
than ashy gray tint; ocherous subbasal bar somewhat 
more strongly accented. The genitalia of the female 
type show no appreciable differences from those of 
mysiella or maidella. The name may represent nothing 
more than a race or variety but it should be kept specifi- 
cally distinct until closer relationship is proven. In a 
series of some 40-odd specimens before me the color 
seems fairly constant. Alar expanse, 19-24 mm. 

TypE Locauity: Colorado (type in AMNH, ex Rut- 
gers). 

Foop prant: Unknown. 

Distrisution: Colorado; Utah, Stockton (June, Aug., 
Sept.), Vineyard (June, July); New Mexico, Mesilla 
Park (May); California, San Francisco (Apr.), Loma 
Linda (Apr., Aug., Oct.); Washington, Pullman (July, 
Aug.). . 

375. Kumysia fuscatella (Hulst) 

Zophodia fuscatella Hulst, Canadian Ent., vol. 32, p. 173, 1900. 
Eumysia fuscatella (Hulst) Dyar, Ins. Insc. Menstr., vol. 13, p. 
221, 1925.—McDunnough, Check list, No. 6298, 1939. 

Forewing mouse gray; transverse white lines but 
faintly indicated. Hind wings pale smoky fuscous. 
Alar expanse, 21 mm. 

Typr Locatity: Los Angeles County, Calif. (type in 
USNM). 

Foop puant: Unknown. 

Known only from the male type. 


376. Eumysia semicana, new species 


Antenna white annulated with black. Head, palpi, 
thorax, and forewing white densely dusted with blackish 
fuscous, giving the moth a predominately slate-gray 
color with ashy white streakings on forewing in the 
interspaces between the veins, through the middle of 
the cell and bordering the upper and lower veins of cell; 
antemedial and subterminal lines obsolete, indicated 
only by an obscure, transverse, angulate, dark shading 
at basal third and a couple of dark spots on costa near 
apex, from the inner of which a dark shade extends 
toward cell; base of wing (before transverse shade) more 
or less clouded by dark scaling; discal spots at end of 
cell, small, blackish; some obscure blackish dots along 
termen; cilia fuscous, with a median white band. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 189 


Hind wing smoky fuscous; cilia slightly paler, with a 
whitish median band. Alar expanse, 30 mm. 

TypE Locatity: Yakima, Wash. (type in USNM, 
61364). 

Foop pirant: Unknown. 

Described from female type and one female paratype 
from the type locality, collected by A. Rolfs, May 20, 
1932, and received from J. F. G. Clarke under his Nos. 
4008 and 4009. 

A large, dark species. It may eventually prove to be 
only an extremely dark color form or race of maidella, 
but superficially it is so different in color and so unlike 
any other described species in the genus that a separate 
specific designation seems warranted. 


Genus 93: Divitiaca 


[Venational division D. Forewing with 10 veins; 10 from cell, 
8 and 9 stalked; 4 and 5 united; 2 and 3 from the cell. Hind 
wing with 7 and 8 approximate; 2 from close to lower outer angle 
of cell; discocellular vein curved. Labial palpi porrect. Male 
genitalia with uncus triangulate, tapering evenly to rounded or 
bluntly pointed extremity, large in proportion to reduced 
tegumen; transtilla incomplete; harpe with a transverse sclero- 
tized ridge from base of costa to lower angle of cucullus; penis 
without cornuti or other appreciable sclerotizations; vinculum 
broad, stout, short. Female genitalia with ductus seminalis 
from ductus bursae.] 


93. Genus Divitiaca Barnes and McDunnough 


Divitiaca Barnes and McDunnough, Contributions, vol. 2, p. 183, 
1913. (Type of genus: Divitiaca ochreila Barnes and 
McDunnough.) 

Tongue well developed. Antenna pubescent; in 
male a shallow sinus and slight scale tuft at base of 
shaft; in female simple. Labial palpus porrect; long, 
extending over twice the length of the head beyond it; 
broadly scaled; third segment about half the length of 
second. Maxillary palpus minute, filiform. Forewing 
smooth; 10 veins; vein 2 from slightly before angle of 
cell; 3 from angle; 4 and 5 united; 6 from below upper 
angle of cell, straight; 10 from cell, separated from the 
stalk of 8-9; male without costal fold. Hind wing 
with vein 2 from very close to angle of cell; 3 and 5 
stalked; 7 and 8 closely approximate for some distance 
from cell; cell slightly less than one-half the length of 
the wing; discocellular vein curved. Eighth abdominal 
segment of male with a pair of short ventrolateral hair 
tufts and a lateral pair of eversible lobes with long hair 
tufts 

Male genitalia with apical process of gnathos a stout 
hook. Harpe with broadened cucullus, its outer 
margin straight and vertical or slightly rounded; a 
transverse sclerotized ridge extending from base of costa 
to lower angle of cucullus. Anellus a slightly curved 
plate with short lateral lobes. Aedeagus short, slender, 
needlelike; penis without armature. Vinculum short, 
extremity narrowly rounded. 

Female genitalia with bursa and ductus bursae mem- 
branous; bursa large, sausage shaped or trilobed (twice 


constricted near junction with ductus bursae), without 
signum; ductus bursae long, very slender, with or with- 
out a few fine spines at junction with bursa; ductus 
seminalis from ductus bursae nearer to bursa than to 
genital opening. 

This genus is closely related to Macrorrhinia and 
Ocala, agreeing with them in genitalic and most other 
structural characters except for the absence of vein 4 of 
forewing, a consistent character for all the species and 
specimens of the genus. 


377. Divitiaca ochrella Barnes and McDunnough 


Ficures 101, 433, 922 


Divitiaca ochrella Barnes and McDunnough, Contributions, vol. 
2, p. 188, 1918—McDunnough, Check list, No. 6357, 1939. 


Forewing pale ocherous; veins faintly outlined, 
whitish; antemedial line indicated by an outwardly 
curved and slanting row of blackish spots; between this 
and base of wing, above inner margin, a small, more or 
less diffused, blackish spot; subterminal line faintly 
indicated by a broken series of blackish streaks or spots 
on the veins; a dark discal spot at lower outer angle of 
cell, the discal spot at upper outer angle very faint or 
absent. Hind wing semihyaline, whitish. Alar ex- 
panse, 12-17 mm. 

Male genitalia with apical process of gnathos a 
moderately long, stout, slightly curved and evenly 
tapering hook. Female genitalia with ductus bursae 
weakly spined at junction with bursa copulatrix; bursa 
sausage-shaped, in part minutely scobinate. 

Type tocauity: Everglade, Fla. (Apr., type in 
USNM). 

Foop prant: Unknown. 

Known only from the type series and one other male 
from the type locality, and one male from Marco, Fla., 
all in the National Collection. 


378. Divitiaca simulella Barnes and McDunnough 


Divitiaca simulella Barnes and McDunnough, Contributions, 
vol. 2, p. 183, 1913—McDunnough, Check list, No. 6356, 
1939. 

Similar to ochrella, differing only in its smaller average 
size, and somewhat darker fore and hind wings. The 
hind wing is a pale smoky hyaline hue. Alar expanse, 
12-14 mm. 

The male genitalia of the type are a trifle smaller than 
those of the type of ochrella and the apex of uncus 
seems slightly more pointed; but these are scarcely 
more than individual differences. The female genitalia 
are essentially like those of ochrella. 


Typr tocatity: Everglade, Fla. (Apr., type in 
USNM). 

Foop piant: Unknown. 

Represented in the National Collection by the type 
series and one other male from the Everglades. Very 
doubtfully distinct from ochrella. Ina reared series the 
color differences would probably disappear and speci- 
mens reared under moist conditions would show more 


190 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


intense and diffused markings and a considerably darker 
coloration. 


379. Divitiaca parvulella Barnes and McDunnough 
Fieuries 432, 921 


Divitiaca parvulella Barnes and McDunnough, Contributions, vol. 
2, p. 183, 1913 —McDunnough, Check list, No. 6355, 1939. 
Averages smaller than simulella or ochrella. Ground 
color and dark markings of forewing similar to those of 
simulella but with a rather broad whitish band along 
costa. Hind wings pale to dark smoky fuscous. Alar 
expanse, 9-12 mm. 

Male genitalia with apical process of gnathos a long, 
stout arm, swollen and abruptly hooked at the end. 
Female genitalia with bursa and ductus bursae smooth; 
bursa trilobed (twice constricted near junction with 
ductus bursae). 


Typr Locatity: Marco, Fla. (type m USNM). 
Foop prant: Achyranthus ramosissima. 


Distrisution: Florida, Marco (Apr.), Vero Beach 
(Apr.), Key West (Apr.). 

Easily distinguished from other species in the genus 
by the contrastingly white costa of forewing and the 
peculiar development of gnathos and bursa. The Key 
West specimens (1 ¢' and 3 9) were reared Apr. 7, 1945, 
in connection with the Special Survey of the Division of 
Foreign Plant Quarantine, U. S. Department of Ari- 
culture, from larvae feeding in the flowers of Achyranthus. 
They have given us our only food-plant record for the 
genus. The male is slightly darker and somewhat more 
strongly marked than either of the two males of the type 
series. The reared females and a collected female from 
Vero Beach are even darker, their hind wings being a 
dark smoky fuscous. The larva lacks the sclerotized 
rings about seta Iib of mesothorax and seta III of 
eighth abdominal segment characteristic of most phyci- 
tine larvae. 


380. Divitiaca parvulella consociata, new race 


Similar to typical parvulella except somewhat paler 
and larger. The hind wing of the female is darker than 
that of the male but not so dark as in the reared females 
of parvulella from Key West, a difference of little or no 
significance as between collected and reared specimens. 

The genitalia, male and female, are also slightly larger 
than those of Florida parvulella but not structurally 
different. Alar expanse, 11-13.5 mm. 


TypE Locauity: Valle de Medellin, Colombia (type 
in USNM, 61365). 


Foop pLrant: Unknown. 


Described from male type and one male and one 
female paratype from the type locality, received from 
F. L. Gallego M. under his No. 111, and dated “January 
1942.” 


I name these examples with great reluctance and do 
so only as a precautionary measure, since we have no 
examples of Divitiaca from any intervening area between 
Colombia and the United States. 


Genera 94-97: Macrorrhinia to Protasia 


[Venational division A. Forewing with 11 veins; 10 from cell, 8 
and 9 stalked; 4 and 5 stalked; 2 and 3 from cell. Characters 
otherwise as for previous group (Dzvitiaca).] 


94. Genus Macrorrhinia Ragonot 


Macrorrhinia Ragonot, N. Amer. Phycitidae, p. 13, 1887. 
of genus: Macrorrhinia aureofasciella Ragonot.) 
Dolichorrhinia Ragonot, Nouv. Gen., p. 28, 1888; Monograph, 
pt. 2, pp. xi, 190, 1901—Hulst, Phycitidae of N. Amer., p. 

190, 1890. 

Tongue well developed. Antenna pubescent (cilia 
about as long as width of shaft); in male a shallow sinus 
with a very small tuft at base of shaft; in female simple. 
Labial palpus porrect, downcurved; long, extending at 
least three times length of head beyond it. Maxillary 
palpus minute, filiform. Forewing smooth; 11 veins; 
vein 2 from slightly before angle of cell; 3 from angle; 
4 and 5 short stalked; 6 from below upper angle of cell, 
straight; 10 from cell, at base approximate to stem of 
8-9; male without costal fold. Hind wing with vein 2 
from close to angle of cell; 3 and 5 stalked; 7 and 8 
approximate or weakly anastomosed beyond cell; cell 
slightly less than one-half the length of wing; discocel- 
lular vein curved. Eighth abdominal segment of male 
with a pair of short ventrolateral hair tufts and a lateral 
pair of eversible lobes with long hair tufts (as in Diiti- 
aca). 

Male genitalia with apical process of gnathos a stout 
hook. Harpe with broadened cucullus; a transverse 
sclerotized ridge extending from base of costa to lower 
angle of cucullus. Anellus a slightly curved plate with 
short, thick, lateral lobes. Aedeagus short, slender, 
needlelike; penis without armature. Vinculum short 
(as broad as long); extremity rounded. 

Female genitalia with bursa and ductus bursae mem- 
branous throughout, except for a slight sclerotization of 
the genital opening; bursa large, sausage shaped, with- 
out signum; ductus bursae long, very slender for two- 
thirds of its length, suddenly and considerably expanded 
before junction with bursa; ductus seminalis from ductus 
bursae near genital opening. 

Ragonot (1888) proposed Dolichorrhinia as a new 
name for Macrorrhinia Ragonot (1887) since the latter 
was, in his opinion, a homonym, “resembling too much 
the name Macrorhinus already employed twice.” The 
name is uncomfortably close to and much nearer a 
homonym of Macrorrhina Berthold (1827); but under 
our interpretation of the International Code the name 
Macrorrhinia is homonymous with neither and will have 
to replace Dolichorrhinia which lepidopterists since 
Ragonot have been using for the genus. 


(Type 


381. Macrorrhinia aureofasciella Ragonot 
Fiaures 61, 437, 924 
Macrorrhinia aureofasciella Ragonot, N. Amer. Phycitidae, p. 13, 
1887. 


Dolichorrhinia aureofasciella (Ragonot) Hulst, Phycitidae of N. 
Amer., p. 190, 1890; U.S. Nat. Mus. Bull. 52, p. 433, 1902.— 
Hampson, 7m Ragonot, Monograph, pt. 2, p. 190, 1901.— 
McDunnough, Check list, No. 6351, 1939. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 191 


Forewing ocherous gray; antemedial line broad, 
slightly oblique, orange, preceded by a blackish line 
which extends from middle of cell to inner margin; 
discal dots at end of cell, blackish, the lower one always 
distinct; subterminal line very faint, whitish, indis- 
tinctly shaded inwardly by dark scaling, indented at 
vein 6 and lower fold; terminal dots faint. Hind wing 
translucent, whitish with a pale smoky tint. Alar 
expanse, 13-22 mm. 

Male genitalia with apical hooked process of gnathos 
but slightly curved; aedeagus with a row of 4 or 5 very 
small spines near apex. Female genitalia as given for 
the genus. 

Typm Locauity: Arizona (type in Paris Mus.). 

Foop prant: Unknown. 

Distrisution: Unirep Srates: Arizona, Baboqui- 
vari Mts. (Apr. to Dec.), Greaterville, Palmerlee (Apr.), 
Santa Rita Mts. (May, June, July), Wilgus Mts.; 
New Mexico, Albuquerque (July), Las Vegas; Tezas, 
Alice (June), Blanco County (July, Oct.), Brownsville, 
Burnet County, Kerrville (Apr., July), Shovel Moun- 
tain. México: Sonora. 

Superficially aureofasciella is easily confused with 
Eumysia pallidipennella, especially the female. Their 
genitalia, however, readily separate the two species. 


382. Macrorrhinia placidella (Zeller) 


Myelois placidella Zeller, Isis von Oken, p. 874, 1848. 
Dolichorrinia placidella (Zeller) Ragonot, Monograph, pt. 2, 
p. 190, 1901. 

This species is represented only by the female type in 
Berlin. According to Ragonot the specimen is in poor 
condition. I have seen nothing from tropical America 
that matches Zeller’s description or Ragonot’s figure 
(Monograph, pl. 31, fig. 20). Very likely the species 
does not belong in Macrorrhinia, but without evidence 
to the contrary must be left where Ragonot placed it. 

Type Locauity: Alegrete, Brazil (type in Mus. Univ. 
Berlin). 


Foop pirant: Unknown. 


95. Genus Ocala Hulst 


Ocala Hulst, Canadian Ent., vol. 24, p. 61, 1892. 
genus: Ocala dryadelia Hulst.) 


(Type of 


Characters of Macrorrhinia except: Labial palpus of 
male grooved to hold maxillary palpus; maxillary pal- 
pus of male in the form of an aigrette. There is no 
structural difference between the two genera in the 
tufting of the male abdomen. Hulst made no dissec- 
tions and consequently failed to see the eversible tufts 
in Macrorrhinia. 

The palpal differences may be of no more than specific 
significance here as they are, for example, in Dioryctria; 
but until more material is available, other possible 
species discovered, and something known about their 
life histories it seems advisible to keep the generic 
names separate. 


383. Ocala dryadella Hulst 
Ficures 59, 438, 923 


Ocala dryadella Hulst, Canadian Ent., vol. 24, p. 61, 1892.— 
MeDunnough, Check list, No. 6359, 1939. 

Dolichorrhinia platanella Grossbeck, Bull. Amer. Mus. Nat. 
Hist., vol. 37, art. 1, p. 131, 1917.—Barnes and McDun- 
nough, Contributions, vol. 3, p. 222, 1917. 

Forewing pale fuscous gray; antemedial white line 
obscure, indicated chiefly by a narrow outer bordering 
of blackish scales, angled slightly at middle and slanting, 
bordered inwardly on lower half by a pale orange spot 
behind which, on inner margin, is a rather conspicuous 
blackish dot; discal spots at end of cell distinct, black- 
ish; subterminal line faint, with a blackish inner border, 
inwardly angled at vein 6 and (slightly) at lower fold; a 
row of blackish dots along termen. Hind wings trans- 
lucent, pale smoky white, with some dark streaking on 
the veins, especially on the females. Alar expanse, 
14-17 mm. 

Male genitalia with outer margin of cucullus straight; 
aedeagus smooth. Female genitalia with ductus bursae 
sclerotized for a short distance from genital opening and 
swollen for more than a third of its length from junction 
with bursa copulatrix. 

Tyrer Locauities: Charlotte Harbor, Fla. (dryadella, 
in AMNH, ex Rutgers); Fort Myers (platanella, in 
AMNBE). 

Foop prant: Unknown. 

Distrrisution: Florida, Charlotte Harbor, 
glade (Apr.), Fort Myers (Apr.), Marco (Apr.). 


Ever- 


96. Genus Valdivia Ragonot 


Valdivia Ragonot, Nouv. Gen., p. 27, 1888; Monograph, pt. 2, 
p. xi, 1901.—Hampson, in Ragonot, Monograph, pt. 2, p. 
191, 1901. (Type of genus: Valdivia coquimbella Ragonot.) 

Maricopa Hulst, Phycitidae of N. Amer., p. 205, 1890. (Type of 
genus: Ciris lativitella Ragonot.) 

Tongue well developed. Antenna ciliate in male 
(cilia over twice the width of the shaft); simple in 
female. Labial palpus porrect (downcurved), long 
(projecting about 3 times the length of the head beyond 
it). Maxillary palpus minute, filiform. Forewing 
smooth; 11 veins; vein 2 from before outer angle of 
cell; 3 from the angle; 4 and 5 stalked; 6 from below 
upper angle of cell, straight; 10 from cell, closely ap- 
proximate at base or connate with the stem of 8-9; male 
without costal fold. Hind wing with vein 2 from very 
close to outer angle of cell; 3 and 5 stalked; 7 and 8 
approximate or weakly anastomosed just beyond cell; 
cell slightly less than one-half the length of the wing. 
Eighth abdominal segment of male with a short pair of 
ventrolateral hair tufts. 

Male genitalia with apical process of gnathos a rather 
short, stout hook. Harpe with broadened cucullus, its 
outer margin rounded; a transverse sclerotized ridge 
extending from base of costa to lower angle of cucullus. 
Anellus an elongate, curved plate with lateral lobes. 
Aedeagus short, slender; penis without armature. 
Vinculum short, terminal margin rather broadly 
rounded. 


192 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Female genitalia with bursa simple, membranous; 
ductus bursae very slender for half its length from 
genital opening, thence to bursa swollen, spiraled and 
armed throughout the spiraled portion with a line of 
fine, sharp spines, sclerotized only at genital opening; 
ductus seminalis from ductus bursae near genital 
opening. 

Valdivia is close to Macrorrhinia and Ocala, dis- 
tinguished by a slightly wider spacing of veins 2, 3 and 
the stem of 4-5 in forewing, the longer cilia of the 
male antenna, the slightly stouter aedeagus, the absence 
of eversible tufts on the male abdomen, the smaller, 
oval bursa and the spiraled and spined ductus bursae. 
Three species are recognized as belonging to the genus. 
A fourth now listed in Valdivia (Maricopa albocostella 
Hulst) will have to be referred to the Anerastiinae. Its 
type (co) in the Rutgers Collection has a greatly reduced 
tongue, completely enclosed by the labial palpi, vein 4 
of forewing absent and a slight excavation in the base 
of the antennal shaft. 


384. Valdivia coquimbella Ragonot 
FIgurE 436 


Valdivia coquimbella Ragonot, Nouv. Gen., p. 27, 1888.—Hamp- 
son, 7n Ragonot, Monograph, pt. 2, p. 191, 1901. 

Forewing fuscous gray; transverse pale lines obscure, 
the antemedial oblique line preceded by a diffused dark 
shade, the subterminal line indicated only by a broken, 
weakly indented line of blackish scaling running parallel 
to the outer margin; discal dots obscure, blackish; a 
row of faint black dots along termen. Hind wing very 
pale smoky fuscous. Alar expanse, 22 mm. 

Male genitalia figured from paratype in the British 
Museum from the type locality. The type in the Paris 
Museum is also a male (not a female as stated by Hamp- 
son) but is without an abdomen. ‘These are the only 
two examples of the species known. The anellus is 
much shorter than that in lativiltella. V. coquimbella 
also lacks any trace of the yellowish spot edging the 
antemedial line, characteristic of the other two species. 

TypE LocaLity: Coquimba, Chile (type in Paris 
Mus.). 

Foop prant: Unknown. 


385. Valdivia lativittella (Ragonot) 
Figures 434, 485, 926, 927 


Ciris lativitiella Ragonot, N. Amer. Phycitidae, p. 18, 1887. 

Ragonotia lativitella (Ragonot), Ent. Amer., p. 117, 1889. 

Maricopa lativitella (Ragonot) Hulst, Phycitidae of N. Amer., 
p- 206, 1890. 

Valdivia lativittella (Ragonot) Hampson, zn Ragonot, Mono- 
graph, pt. 2, p. 191, 1901.—McDunnough, Check list, No. 
6352, 1939. 

Zophodia aureomaculella Dyar, Journ. New York Ent. Soc., vol. 
12, p. 107, 1808. (New synonymy.) 

Eumysia aureomaculella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221, 
1925.—McDunnough, Check list, No. 6296, 1939. 


Paler than coquimbella, the forewing very pale ashy 
gray; on inner margin at base a pale ocherous spot; a 
similar somewhat larger spot extending halfway across 


the wing and bordering the antemedial line; on inner 
margin between the two ocherous patches a blackish 
fuscous spot (rather pronounced on the male, less so on 
the female); antemedial line vertical, faint, mdicated 
chiefly by some irregular and broken, outer, blackish 
fuscous shading; subterminal whitish line more distinct, 
indented at vein 6 and lower fold and margined inwardly 
by a narrow dark line; discal and terminal dots blackish 
fuscous, the latter faint. Hind wing very pale smoky 
fuscous with a faint ocherous tint; a narrow dark line 
along termen; veins faintly darkened. Alar expanse, 
16-21 mm. 

Male genitalia differ from those of coqguimbella chiefly 
in their much longer anellus. 

Type Locatitigs: Arizona (lativittella, in Paris Mus.) ; 
Bremond, Tex. (aureomaculella, in USNM). 


Foop purant: Unknown. 


Distrisution: Unirep Statss: Arizona, Baboqui- 
vari Mts. (July), Greaterville (June), Phoenix (Aug.), 
Pinal Mts. (Aug.); Texas, Bremond (Apr.), Brownsville 
(Mar.), San Benito (Mar., May). Mexico: Durango. 

Twenty-five specimens are before me. A male from 
Durango, Dyar’s type of aureomaculella from east- 
central Texas and a series from Arizona exhibit no es- 
sential differences. The Texan specimens from Browns- 
ville and San Benito (1 @ and 3 6) show some trifling 
differences in male genitalia and considerably weaker 
spining of the ductus bursae in the female (fig. 926), 
but nothing, however, that would justify specific or 
even racial separation. Dyar’s supposed species is an 
obvious synonym. He had not recognized either 
Valdivia or lativittella when he described aureomaculella 
or, later, transferred it to Humysia. The key vena- 
tional character given by Hampson for the separation 
of latiittella from coquimbella (vein 8 of hind wing “free” 
from or anastomosed with 7) does not hold. In our 
Pinal Mountains series of lativittella ves 7 and 8 occur 
both ways, closely approximate or partially anasto- 
mosed. 


386. Valdivia walkerella (Ragonot), new combination 
Fieure 439 


Saluria walkerella Ragonot, Nouv. Gen., p. 44, 1888. 
Hypogryphia walkerella (Ragonot) Hampson, in Ragonot, Mono- 
graph, pt. 2, p. 194, 1901. 

I have seen nothing from Chile that exactly matches 
Ragonot’s description or the figure in his Monograph 
(pl. 37, fig. 21), but the genitalia of the male type clearly 
shows it to be a Valdivia and not a member of the Old 
World genus Hypogryphia where Hampson placed it. 
Veins 4 and 5 of forewing are shortly stalked and the 
ocherous spot on inner margin before the antemedial 
line is present, as in Jativitiella which it strongly re- 
sembles in genitalia and general habitus. Alar expanse, 
18 mm. 

Typr Locanity: Valparaiso, Chile (type in Paris 
Mus.). 


Foop prant: Unknown. 
Known only from the type. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 193 


97. Protasia, new genus 


Typ or Genus: Valdivia mirabilicornella Dyar. 

Tongue short (but somewhat exposed between the 
palpi). Antenna unipectinate in male, simple in fe- 
male. Labial palpus porrect, long (projecting more than 
3 times the length of the head beyond it). Maxillary 
palpus minute, filiform. Forewing smooth; 11 veins; 
vein 2 from before outer angle of cell; 3 from the angle; 
4 and 5 very shortly stalked; 6 from below upper angle 
of cell, straight; 10 from cell, approximate to 8-9 at 
base; male without costal fold. Hind wing with vein 
2 from very close to outer angle of cell; 3 and 5 stalked; 
7 and 8 approximate or weakly anastomosed for a short 
distance beyond cell; cell less than one-half the length 
of the wing. Eighth abdominal segment of male 
simple. 

Male genitalia with apical process of gnathos a stout, 
bluntly pointed hook. Harpe with lower margin 
evenly curved; cucullus elongate, tapering slightly to 
somewhat narrowly rounded apex; a transverse, weakly 
sclerotized ridge extending from slightly below base of 
costa to lower angle of cucullus. Anellus a simple, 
nearly flat plate, without lateral lobes. Aedeagus 
short, stout; penis without armature. Vinculum short, 
terminal margin broadly rounded. 

Female genitalia with bursa simple, membranous, 
elongate; ductus bursae short, simple, membranous 
throughout; ductus seminalis from ductus bursae. 

The genus is readily distinguished from the others in 
this immediate group by its unipectinate male antenna, 
short tongue, lack of hair tufts on eighth abdominal 
segment of male, and short and simple ductus bursae. 


387. Protasia mirabilicornella (Dyar), new combination 
Ficures 440, 925 


Valdivia mirabilicornella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 60, 1908.—McDunnough, Check list, No. 6353, 1939. 
Whitish gray with a sprinkling of dark scales and a 
faint ocherous shading in the median and lower folds of 
forewing; antemedial line dentate, outwardly bordered 
by a thin blackish fuscous line, the latter sharply out- 
angled between costa and middle of cell, very slightly 
out-angled at lower margin of cell and at inner margin, 
preceded by a more or less obscure, broad ocherous 
orange shade, bordered basally on inner margin of wing 
by a blackish fuscous patch, the latter encroaching upon 
and more or less obscuring the ocherous shade; subter- 
minal line with a narrow blackish inner border, sharply 
indented to vein 6, thence bulging outward and again 
indented at lower fold; discal dots distinct, blackish; 
terminal dark dots more or less confluent. Hind wing 
pale brassy whitish (pale smoky fuscous in some speci- 
mens), sheeny; a slightly darker line along termen; 
cilia somewhat paler. Alar expanse, 18-26 mm. 

Male genitalia with uncus evenly tapering from broad 
base to rather narrowly rounded apex; elements of 
transtilla small, weak, recurved (fig. 440b). Female 
genitalia with bursa narrow, nearly three times as long 
as ductus bursae. 

300329—56——14 


Type Locatity: San Diego, Calif. (type in USNM). 

Foop pirant: Unknown. 

Represented by 25 specimens in the National Collec- 
tion, all from the type locality (July, Oct., Nov., Dec., 
Feb.). 

In his description Dyar describes the male antenna 
as “bipectinate.” It is distinctly unipectinate, with 
the pectinations and the shaft covered with long cilia. 


Genera 98-100: Heterographis to Hulstia 


[Venational division A. Forewing with 11 veins; 10 from cell; 
2 from near lower outer angle of cell. Hind wing with 7 and 8 
approximate or very weakly anastomosed; 2 from close to lower 
outer angle of cell; discocellular vein curved. Labial palpi up- 
turned. Male genitalia with uncus triangulate (or spoon-shaped), 
large in proportion to reduced tegumen; tegumen constricted 
before uncus, its lateral margins concave; transtilla incomplete 
(except in Heterographis samaritanella); aedeagus straight, 
slender; penis without cornuti or other armature. Female 
genitalia with signa consisting of many straight disconnected 
thornlike spines; ductus seminalis from ductus bursae.] 


98. Genus Heterographis Ragonot 


Heterographis Ragonot, Ent. Monthly Mag., vol. 22, p. 31, 1885; 
Monograph, pt. 2, p. x, 1901.—Hulst, Phycitidae of N. 
Amer., p. 186, 1890.—Hampson, in Ragonot, Monograph, 
pt. 2, p. 140, 1901.—Bisset, in Pierce and Metcalf, Genitalia 
of the British Pyrales, p. 59, 1938 (fixes type of genus).— 
Janse, Journ. Ent. Soc. South Africa, vol. 8, p. 37, 1945. 
(Type of genus: Huzophera samaritanella Zeller; figs. 441, 
928). 

Mona Peat Ent. Amer., vol. 4, p. 115, 1888. 
Mona olbiella Hulst.) 

Tongue well developed. Antenna simple, pubescent; 
in the male the shaft slightly curved towards base. 
Labial palpus upcurved, reaching above vertex. Max- 
illary palpus minute, filiform. Forewing smooth; 11 
veins; vein 2 from near lower outer angle of cell; 3 from 
the angle; 4 and 5 connate or very shortly stalked, 
closely approximate to 3; 6 from below upper angle of 
cell, straight; 10 from cell, separated from the stalk of 
8-9; male without costal fold. Hind wing with 2 from 
close to outer angle of cell; 3 and 5 stalked; 7 and 8 
closely approximate (occasionally contiguous or weakly 
anastomosed for a short distance) beyond cell; cell less 
than one-half (over one-third) the length of the wing. 
Eighth abdominal segment of male with a strong pair 
of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos a stout 
hook. Elements of transtilla long, slender, curved, 
their apices touching and weakly fused (samaritanella) 
or narrowly separated (morrisonella) ; deeply and round- 
ly arched behind aedeagus. Harpe simple, narrowly 
elongate; apex rounded. Anellus a slightly curved 
plate with lateral arms. Aedeagus long, slender; penis 
without armature. Vinculum at least as long as broad; 
truncate; terminal margin straight or slightly notched. 

Female genitalia with bursa copulatrix and ductus 
bursae membranous except for a slight sclerotization of 
the tube of ductus near genital opening; bursa elongate 
with signa consisting of numerous straight, sharp, broadly 


(Type of genus, 


194 


based, disconnected spines, a few of these spines in 
ductus near its junction with bursa; ductus seminalis 
from ductus bursae near genital opening. 

Represented in North America by a single species. 


388. Heterographis morrisonella Ragonot 
Figures 442, 929 


Heterographis morrisonella Ragonot, N. Amer. Phycitidae, p. 11, 
1887.—Hulst, Phycitidae of N. Amer., p. 186, 1890.—Hamp- 
son, zz Ragonot, Monograph, pt. 2, p. 169, 1901.—Dyar, 
Proe. Ent. Soc. Washington, vol. 7, p. 38, 1905—McDun- 
nough, Check list, No. 6339, 1939. 

Heterographis colorandensis Ragonot, N. Amer. Phycitidae, p. 12, 
1887 


Mona olbiella Hulst, Ent. Amer., vol. 4, p. 116, 1888. 

Heterographis morrisonella coloradensis (Ragonot) Hulst, Phy- 
citidae of N. Amer., p. 186, 1890.—Hampson, in Ragonot, 
Monograph, pt. 2, p. 169, 1901. 

Heterographis morrisonella olbiella (Hulst), Phycitidae of N. 
Amer., p. 186, 1890.—Hampson, 7m Ragonot, Monograph, 
pt. 2, p. 169, 1901. 

Heterographis ignistrigella Ragonot, Monograph, pt. 2, p. 166, 
1901.—McDunnough, Check list, No. 6338, 1939. (New 
synonymy.) 

Honora palloricostella Walter, Proc. Ent. Soc. Washington, vol. 
30, p. 141, 1928.—McDunnough, Check list, No. 6344, 1939. 
(New synonymy.) 

Extremely variable in color and intensity of mark- 
ings, ranging from a creamy ocherous form with only 
the faintest dark dusting to forms much suffused with 
reddish or blackish fuscous. Forewing with costa some- 
what paler (from base to outer line) than the remainder 
of the wing in nearly all specimens, contrastingly whit- 
ish in some; transverse pale lines more or less distinct; 
antemedial line outwardly slanting to lower margin of 
cell, indented below at fold; subterminal line nearly 
parallel with outer margin, indented at vein 6 and lower 
fold; in some specimens an ocherous band, a blackish 
fuscous patch, or a diffused reddish shade preceding the 
antemedial line; similar colorations frequently following 
the subterminal line; the area between the transverse 
lines frequently grayish from the heavy dusting of dark 
scales; usually the blackish dusting is more conspicuous 
along some of the veins. Hind wings from very pale to 
dark smoky fuscous; cilia whitish; veins more or less 
outlined by fuscous shading. Alar expanse, 13-23 mm. 

Male genitalia with lateral margins of uncus straight, 
apex bluntly rounded. Harpe with costa and lower 
margin of harpe nearly parallel; cucullus slightly curved 
near apex. Female genitalia with bursa long; spines of 
signa arranged in more or less regular, longitudinal 
series, the number and arrangement of spines differing 
in different specimens, hardly any two examples being 
identical (extremes of variation are shown in the fig- 
ures) ; ductus bursae long, slender throughout its length, 
weakly sclerotized between junction of ductus seminalis 
and genital opening. 

TypE Locatitius: Texas (morrisonella and ignistri- 
gella, in Paris Mus.); Colorado (coloradensis, in Paris 
Mus.); Salida, Colo. (olbiella, in AMNH, ex Rutgers) 
Tempe, Ariz. (palloricostella, in USNM). 

Foop pruant: Franseria bipinnatifida. A single fe- 
male reared by F. N. Pierce, Sept. 6, 1938, at El Se- 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


gundo, Calif., our only food-plant record. The species 
presumably feeds on other composites. 

Distrisution: Unirep States: Texas, Beeville 
(Oct.), Brownsville (June), Burnet County (Apr.), 
Sabinal (Apr., May, June), San Benito (June, July, 
Sept.), San Diego (May), Zavalla County (Apr.); New 
Mexico, Albuquerque (July), Deming (July), Fort Win- 
gate (June, July), Hot Springs (Aug.), Jemez Springs, 
Las Vegas (May), Mesilla Park (Apr., May), Santa Fe; 
Arizona, Baboquivari Mts. (July), Phoenix (Mar., 
Apr., Aug., Oct.), Tempe (Apr., Aug., Sept., Oct.), 
White Mts. (Aug.), Yuma County (Mar.); Colorado, 
Denver, Glenwood Springs (Aug.); Utah, Provo (July), 
Scover Valley (May), Stockton (June, July, Aug., Oct.), 
Vineyard (June); Nevada, Pyramid Lake; California, 
Balboa, (Sept.), Death Valley (Apr.), El Segundo 
(Sept.), Ibanpa Mts. (Oct.), Jacumba (June), Laguna 
(Sept.), La Puerta (July), Loma Linda (Mar., Apr., 
May, Aug., Sept.), Narrows (Mar.), Palm Springs 
(Mar., Apr., June), San Diego (June); Washington, 
Blue Mts. (Bone Springs, July), Pullman (June, July), 
Walla Walla (May, June), Yakima (June). M*xrco: 
Durango, Sonora. 

The color variations are so many, the color forms so 
widely distributed, and the intergradations so gradual 
that none of the synonymical names can be used as a 
racial designation. The species, however, is clearly 
defined and easily identified by its genitalic characters. 


99. Genus Staudingeria Ragonot 


Staudingeria Ragonot, Ann. Soc. Ent. France, ser. 6, vol. 7, p. 
249, 1887; Monograph, pt. 2, p. x, 1901.—Hulst, Phycitidae 
of N. Amer., p. 185, 1890—Hampson, in Ragonot, Mono- 
graph, pt. 2, p. 181, 1901. (Type of genus: Ancylosis morbo- 
sella Staudinger.) 

Characters of Heterographis except: Labial palpus 
obliquely ascending, third segment very short, less than 
one-third the length of second, in male second segment 
grooved to hold the maxillary palpus; maxillary palpus 
of male in the form of an aigrette; female genitalia with 
bursa very narrowly elongate, ductus bursae much 
shorter than bursa. 

The status of this genus in relation to Heterographis 
is similar to that of Ocala in relation to Macrorrhinia. 
If only our North American species were involved I 
should be inclined to consider the differences in male 
maxillary palpi and the female bursae to be merely spe- 
cific; but apparently some of the species under each 
generic name in the Old World are properly assigned 
and the two groups of species there are separable on the 
male palpal character; so, for the present at least, it 
seems advisable to keep the names separate. I have 
not seen the type of Staudingeria, but there is no reason 
to doubt that our American species is congeneric. Many 
specimens of albipenella (especially what Dyar described 
as olivacella) are an exact match for Ragonot’s figure of 
morbosella (Monograph, pl. 27, fig. 1). The most striking 
difference between our American representatives of 
Staudingeria and Heterographis is in the comparative 
length of the ductus bursae of the female. In Hefero- 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 195 


graphis it is at least as long as the bursa. In Staudin- 


geria it is much shorter than the bursa. 


389. Staudingeria albipenella (Hulst) 
Fiagures 443, 930 


Pempelia albipenella Hulst, Ent. Amer., vol. 3, p. 133, 1887. 

Staudingeria albipennella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 
116, 1889; Monograph, pt. 2, p. 186, 1901.—Dyar, Proc. Ent. 
Soc. Washington, vol. 7, p. 38, 1905.—Hulst, Phycitidae of 
N. Amer., p. 186, 1890. 

Staudingeria olivacella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 111, 1904. 

Siaudingeria perluteella Dyar, Proc. 
vol. 6, p. 111, 1904. 

Staudingeria albipenella (Hulst), McDunnough, Check list, No. 
6337, 1939. 

Variable in color and intensity of markings, ranging 
from a form with forewing a uniform luteous ocherous 
without definite markings, though forms more or less 
dusted and clouded with blackish fuscous, to a form 
with a clear red ground color and gradations of this 
with the red more or less overlaid with blackish fuscous 
scaling; costal edge white or whitish from base to just 
before apex, the whitish costal margin sometimes very 
thin and obscure but more often contrasted against the 
ground color; transverse lines obsolete or obscure, in 
better marked specimens chiefly indicated by their dark 
marginal shadings; such shadings on subterminal line 
deeply indented at vein 6 and slightly indented at lower 
fold; cilia pale luteous ocherous, ocherous fuscous, or red, 
concolorous with the ground color. Hind wing from 
very pale to dark smoky fuscous. Fore and hind wings 
have a slick, glossy sheen. Alar expanse, 18-27 mm. 

Male genitalia with uncus tapering to a blunt point, 
its lateral margins curved; anellus with very short 
lateral lobes; aedeagus somewhat stouter than that of 
Heterographis; vinculum very slightly tapered, terminal 
margin broad and with a shallow indentation. Female 
genitalia with bursa copulatrix a long tube terminating 
in a small round bulb; signum consisting of an irregular 
cluster of separate spines in the bulbous part of bursa, 
a scattering of similar spines in the anterior half of the 
tube and a small cluster of much shorter spines in the 
tube near junction of ductus bursae; ductus not appreci- 
ably sclerotized near genital opening. 

Type Locauities: Southern California (albipenella, 
in AMNH, ex Rutgers); Pullman, Wash. (olivacella, in 
USNM); Fort Collins, Colo. (perluteella, in USNM). 

Foon piant: “Loco weed.” 

Distrisution: New Mexico, Fort Wingate (May); 
Arizona, Phoenix (Apr.), Woodruff (June); California, 
San Diego (June); Colorado, Denver (July, Sept.), Fort 
Collins, Glenwood Springs (June, July); Utah, Provo, 
Richfield (June), Stockton (July, Aug., Sept.); Wyo- 
ming, Yellowstone Park (July) ; Idaho, Blackfoot (June), 
Sawtooth (June); Oregon, Bone Springs (Blue Mts., 
July), Crater Lake (July); Washington, Godman 
Spring (Blue Mts., July), Pullman (Aug., Sept.), 
Vantage Ferry (May), Walla Walla (May, June, July), 
Wenatchee (May, June, July, Aug.), Yakima (May). 

Dyar’s two names (olivacella and perluteella) appear 


Ent. Soc. Washington, 


as racial designations in our lists but the differentiation 
is not justified, for there is complete intergradation 
between them and albipenella in any considerable 
series from a given locality. The most outstanding 
color form is exemplified in a series from Crater Lake, 
Oreg. This is a clear, shining red variety, matched 
only in some specimens from Idaho and Washington. 
However there are intergrades in these localities and 
one in the Crater Lake series between the red form and 
the type of albipenella, so I do not feel justified in 
adding another name. As in Heterographis we appear 
to have only one clear-cut American species of the Old 
World genus Staudingeria, and it seems limited in its 
distribution to the Western States. It is easily identi- 
fied by its genitalia. 


100. Genus Hulstia Ragonot 


Hulstia Ragonot, Monograph, pt. 2, p. x, 1901.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 127, 1901. 

Hulstea Hulst, U. S. Nat. Mus. Bull. 52, p. 482, 1903. (Spelling. 
Type of genus: Nephopteryx undulatella Clemens.) 

Tongue well developed. Antenna simple, finely pu- 
bescent. Labial palpus upturned; rather broadly and 
smoothly scaled; third segment short, pointed. Mazxil- 
lary palpus small, squamous, appressed to face. Fore- 
wing smooth; 11 veins; vein 2 from near lower outer 
angle of cell; 3 from the angle; 4 and 5 stalked, the 
stalk closely approximate to 3 at base; 6 from below 
upper angle of cell, straight; 10 from the cell, at base 
connate or very closely approximate to 8-9; male with- 
out costal fold. Hind wing with vein 2 from close to 
outer angle of cell; 3 and 5 stalked; 7 and 8 contiguous 
or partially anastomosed for a short distance beyond 
cell; cell slightly less than one-half the length of wing; 
discocellular vein curved. Eighth abdominal segment 
of male with a pair of short, ventrolateral hair tufts. 


Male genitalia with apical process of gnathos a stout, 
blunt hook. Elements of incomplete transtilla, small 
and weak. Harpe simple, narrowly elongate; apex 
rounded. Anellus a narrowly elongate plate with 
strongly sclerotized margin and short lateral lobes (re- 
sembling an elongate and closed U). Aedeagus long, 
moderately slender. Vinculum as broad as long, squar- 
ish, terminal margin nearly straight. 


Female genitalia with bursa copulatrix long, tapering 
and expanding gradually from ductus bursae; demarca- 
tion of bursa from ductus obscure; signa consisting of 
a dense mass of straight, short, rather slender spines 
with small bases and filling about half the bursa. 
Ductus bursae short, with a strongly sclerotized, acutely 
bent elbow before genital opening; genital opening 
simple. Ductus seminalis from ductus bursae just 
before sclerotized elbow. 

Hulstia is very close to Heterographis and Staudin- 
geria, agreeing with them in most structural characters 
and differing chiefly in the more pronounced stalking 
of veins 4 and 5 of forewing, the closer approximation 
of vein 10 to the stalk of 8-9 at base, its much reduced 
transtilla, the shape of the female bursa, the densely 


196 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


clustered and narrowly based cornuti, and the sclero- 
tized elbow in ductus bursae. It contains but one 
known species. The other American species that have 
been assigned to it by Hampson (texanella and cordu- 
bensiella) are dissimilar in both structure and habitus. 


390. Hulstia undulatella (Clemens) 
Ficures 64, 444, 931 


Nephopteryx undulatella Clemens, Proc. Acad. Nat. Sci. Phila- 
delphia, p. 205, 1860. 

Scoparia rubiginalis Walker, List, vol. 34, p. 1498, 1865. 

Honora obsipella Hulst, Ent. Amer., vol. 4, p. 118, 1888 (new 
synonymy). 

Heterographis oblitella Ragonot (not Zeller), Ent. Amer., vol. 5, 
p. 116, 1889. 

Honora oblitella undulatella (Clemens) Hulst, Phycitidae of N. 
Amer., p. 187, 1890. 

Honora fumosella Hulst, Canadian Ent., vol. 32, p. 174, 1900.— 
Forbes, Cornell Mem. 68, p. 6338, 1920.—McDunnough, 
Check list, No. 6350, 1939. (New synonymy.) 

Hulstia undulatella (Clemens) Hampson, in Ragonot, Mono- 
graph, pt. 2, p. 128, 1901.—Hssig, Insects of western North 
America, p. 711, 1926.—McDunnough, Check list, No. 6341, 
1939.—Darlington, Trans. Amer. Ent. Soc., vol. 73, p. 91, 
1947. 

Hulstea undulatella (Clemens) Hulst, U. S. Nat. Mus. Bull. 52, 
p- 432, 1903. 


Forewing whitish ocherous, pale gray or dark grayish 
fuscous, more or less dusted with white scaling especially 
in median area of wing, in darkest specimens (color form 
jfumosella) pattern mostly obliterated by dark scaling; 
transverse lines normally distinct, narrow, white; ante- 
medial line oblique, notched at cell and lower fold, 
broadly bordered inwardly by fuscous or fuscous and 
ocherous, especially toward inner margin, and outward- 
ly by a narrow dark shade especially from costa to cell; 
subterminal line sinuate, parallel to and rather near 
termen, bordered inwardly by a narrow dark line and 
outwardly by a fuscous or ocherous fuscous shade; discal 
spots, especially the lower one, usually distinct. Hind 
wings dark smoky fuscous. Palpi, underside of body, 
and legs whitish, more or less shaded with fuscous in 
darker specimens. Alar expanse, 16-22 mm. 

Genitalic characters as given for the genus. 


TYPE LOCALITIES: Pennsylvania (undulatella, lost); 
North America (rubiginalis, in BM); ‘Texas’ [sic] 
(obstpella, in AMNH, ex Rutgers); Newark, N. J. 
(fumosella, in AMNH, ex Rutgers). 

Foop prants: Elm [?], sugar beets. 


Distrizution: Unitep States: Maine; Massachu- 
setts, Martha’s Vineyard (July), Vineyard Haven 
(Aug.); Connecticut, East River (July, Sept.); New 
Hampshire, Hampton (June, Aug., Sept., Oct.); New 
Jersey, Anglesea (June), Newark; North Carolina, 
Southern Pines (Oct.); Florida, Glenwood, Hastings 
(Apr.); Wisconsin, Cranmoor; Montana, Bozeman 
(Aug.); Colorado, Clear Creek, Denver (Aug., Sept.), 
Florisant (July), Glenwood Springs (Aug.), Windsor 
(July) ; Utah, Delta (July), Logan (Aug.), Ogden (July), 
Provo (July), Salt Lake, “Southern Utah” (July), 
Stockton (Aug.), Vineyard (June); Nevada, Clark 
County, Reno (July); New Mexico, Fort Wingate 


(June), Hot Springs, Jemez Springs (July), Sapello 
Canyon (July) ; Arizona, Flagstaff, ‘‘Southern Arizona,” 
Williams (July); California, Alameda (Aug., Oct.), 
Chino (Nov.), Folsom (July), Palo Alto, Sacramento 
(Oct.), San Diego (Sept., Oct., Nov.), Siskiyou (Sept.), 
Truckee (Aug.), Woodland; Washington, Bellingham 
(July, Aug.), Friday Harbor (July), Pullman (May, 
July, Aug., Sept.), Snake River (May), Walla Walla 
(June, July), Wenatchee (July, Aug.). CAaNapa: Que- 
bec, Montreal; Alberta, Calgary (July, Aug.); Manitoba, 
Aweme (July, Aug.); British Columbia, Arrowhead 
Lake, Goldstream (July), Kaslo, Victoria (Aug.). 

This species was described from eastern specimens, 
but seems to be much more abundant in our Western 
States. Very little is known of its life history. Clem- 
ens states that he took larvae and pupae of undulatella 
on elm at Niagara Falls, Canada. He did not rear any 
moths from the larvae and his statement is ambiguous 
as to the rearing of moths from the pupae (which were 
found under the bark of the tree), so the association of 
undulatella with elm as a host is by no means estab- 
lished. In the National Museum we have a half-dozen 
reared specimens from Colorado, Utah, and California 
reared from larvae feeding on sugar-beet foliage. ‘This 
is the only authenticated food-plant record with which 
IT am acquainted. 

The name fumosella represents nothing more than a 
much suffused, dark color form. There are four 
examples in the National Collection matching Hulst’s 
type and several specimens from widely scattered 
localities intergrading between it and typical wndula- 
tella. None of these (including the type of fumosella) 
exhibits any genitalic differences. 

The type of obsipella is a freak specimen with vein 4 
present in one hind wing. On the strength of this 
venation it was referred to the synonymy of Hlasmo- 
palpus petrellus by Hulst in 1890 and so appears in our 
lists. It is a ragged female with only one hind wing 
complete enough to show venation and is labeled 
“Colorado” a more probable locality than Texas as 
given in the original description. Its genitalia are 
those of undulatella. 


Genera 101-104: Honora to Cabotia 


[Venational division A. Forewing with 11 veins; 10 from cell; 
2 from or from very near lower outer angle of cell; 3 closely 
approximate to or connate with stalk of 4-5; 4-5 stalked at least 
for one-half their lengths. Hind wing with 7—8 anastomosed or 
contiguous (Oncolabis) beyond cell; 2 from or from very near 
lower outer angle of cell; discocellular vein curved. Labial palpi 
oblique. Male genitalia with uncus triangulate; tegumen not 
appreciably reduced in proportion to uncus; harpe with short 
erect clasper; transtilla incomplete (except in Honorinus) ; aedea- 
gus short, stout; penis armed with sclerotized folds or granula- 
tions or short spines. Female genitalia with signa consisting of 
densely spined round plate, detached thornlike spines or both; 
ductus seminalis from bursa copulatrix.] 


101. Genus Honora Grote 


Honora Grote, Bull U. 8S. Geol. Geogr. Surv. Terr., vol. 4, p. 702, 
1878; North Amer. Ent., vol. 1, p. 11, 1879.—Hulst, Phycitidae 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 197 


of N. Amer., p. 187, 1890.—Ragonot, Monograph, pt. 2, 
p- x, 1901—Hampson, in Ragonot, Monograph, pt. 2, p. 
183, 1901. (Type of genus: Honora mellinella Grote.) 

Tongue well developed. Antenna simple, pubescent. 
Labial palpus oblique, the third segment porrect (de- 
flected forward). Maxillary palpus small, squamous, 
appressed to face. Forewing smooth; 11 veins; veins 
2 and 3 closely approximate and from the lower outer 
angle of cell; 4 and 5 stalked, the stalk shortly sepa- 
rated from 3 at base; 6 from below upper angle of cell, 
straight; 10 from the cell, at base shortly separated 
from 8-9; male without costal fold. Hind wing with 
vein 2 from the lower outer angle of cell, connate with 
3, 3 and 5 stalked; 7 and 8 anastomosed for nearly half 
their lengths beyond cell; cell one-third the length of 
wing; discocellular vein curved. Eighth abdominal 
segment of male with a pair of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos a 
stout hook. Harpe narrowly elongate; apex rounded 
or bluntly pointed; an erect, narrow, short clasper 
arising from sacculus near base of cucullus. Anellus a 
nearly flat plate with a cupped depression at base and 
posterior margin more or less notched. Aedeagus 
short, moderately stout, slightly bent (elbowed) near 
middle; penis armed with a narrow row of fine, spine- 
like cornuti. Vinculum stout, as broad or slightly 
broader than long; terminal margin broad. 

Female genitalia with signa strongly developed, con- 
sisting of a single large, round, curved, densely spined 
plate and a scattering of detached spines (except in 
dotella) opposite the plate; ductus bursae as long as or 
somewhat shorter than length of bursa, finely scobinate 
for two-thirds of its length from bursa; a narrow sclero- 
tized plate in area above genital opening; ductus semi- 
nalis from a lobed projection of bursa near its junction 
with ductus bursae. 

The genus is apparently confined to North America. 
The species perdubiella (Dyar) is referred here from 
Zophodia. Another, dulciella (Hulst), described in 
Honora, must find placement elsewhere. The genitalia 
of the unique female type (fig. 951) show that it is not 
a Honora; but its proper generic placement will have 
to wait upon discovery of a male. It is treated briefly 
at the end of the paper (see p. 313). 

The species are very close and the genitalic differences 
separating them, especially among the males, are 
trifling. The female genitalia show more obvious varia- 
tion in the amount of spining in the bursa, but this is a 
character of doubtful value. Such differences as exist 
are shown in the several drawings. 


391. Honora mellinella Grote 
Figures 57, 445, 937 


Honora mellinella Grote, Bull. U. S. Geol. Geogr. Surv. Terr., 
vol. 4, p. 702, 1890.—Hulst, Phycitidae of N. Amer., p. 188, 
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 185, 
1901.—McDunnough, Check list, No. 6343, 1939. 

Honora ochrimaculella Ragonot, N. Amer. Phycitidae, p. 12, 
1887.—Hulst, Phycitidae of N. Amer., pp. 188, 189, 1890.— 
Hampson, in Ragonot, Monograph, pt. 2, p. 185, 1901. 


Forewing dark grayish fuscous; costal area from 
upper margin of cell whitish, in some specimens this 
pale costal shading somewhat obscure, in most speci- 
mens rather well contrasted against ground color; ante- 
medial line nearly vertical, white, distinct and slightly 
concave towards inner margin, bordered inwardly by 
faint blackish shading and outwardly on inner margin 
by a yellowish patch; a similar yellowish patch at base 
of wing; subterminal white line, narrow, sinuate, ob- 
scure and frequently obliterated; discal dots at end of 
cell blackish, the upper in the pale costal area, the lower 
in the dark area but normally with a faint, narrow, 
whitish border. Hind wing very pale smoky fuscous, 
with a silky sheen. Alar expanse, 16-24mm. Male 
genitalia with apex of harpe rounded; cucullus not 
appreciably broadened toward apex. Female with a 
rather dense cluster of spines in bursa opposite spined 
plate. 

TyprE LOCALITIES: Texas (mellinella, in BM); Cali- 
fornia (ochrimaculella, in Paris Mus.). 

Foop puant: Palafoxia. 

Distrisution: North Carolina, Southern Pines (Aug., 
Sept.); Zouwisiana, Orange, Sabine Parish (June); 
Florida, Lakeland (Jan.), Tampa; Teras, Blanco 
County, Burnet County, Kerrville (Mar., July, Sept., 
Oct.), New Braunfels (May), San Benito (Mar., Apr.) ; 
New Mexico, Hot Springs (Sept.); Arizona, Palmerlee, 
Phoenix (Oct.), Redington, Santa Rita Mts. (June, 
July); California, Indian Wells (Jan.); Washington, 
Pullman (June), Walla Walla (May, June, July), 
Yakima (May). 

The food plant record given above is from a female 
from Indian Wells, Calif., reared by Commander C. M. 
Dammers. 


392. Honora subsciurella Ragonot 
Fieures 447, 941 


Honora subsciurella Ragonot, N. Amer. Phycitidae, p. 12, 1887.— 
Hulst, Phycitidae of N. Amer., p. 189, 1880.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 186, 1901.—McDunnough, 
Check list, No. 6347, 1939. 

Doubtfully distinct from mellinella except, possibly, 
as a local race. Distinguished chiefly by the paler 
ground color of forewing beyond antemedial line, due 
to a fine powdering of white scales; a broader diffusion 
of the pale costal area (usually extended to include the 
lower discal spot); the bluntly pointed apex of harpe 
(fig. 447); and the sparser spining of the female bursa 
(compare figs. 941 and 937). Alar expanse, 22-28 mm. 

Typr Locauity: Colorado (type in Paris Mus.). 

Foop piant: Unknown. 

Distripution: Colorado, Glenwood Springs (Sept.); 
Utah, Eureka (June, July, Aug.), Stockton (July, Aug.), 
Vineyard (July); California, San Bernardino Mts. 
(Sept.). 

The “red” ground color of forewing is exaggerated by 
Ragonot in his original description and in the figure 
(pl. 31, fig. 12) in his Monograph. Hampson’s de- 
scription (in Ragonot) is more accurate. 


198 


393. Honora sciurella Ragonot 
Ficure 446 


Honora sciurella Ragonot, N. Amer. Phycitidae, p. 12, 1887.— 
Hulst, Phycitidae of N. Amer., p. 189, 1890.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 185, 1901—McDunnough, 
Check list, No. 6346, 1939. 

Known to me only from descriptions of Ragonot and 
Hampson, the Ragonot figures (plate 37, figs. 16, 17) 
and the genitalia of the male type. The harpes of the 
latter (fig. 446) are somewhat longer in proportion to 
the remainder of the genitalia than in the other species 
of the genus. In the National Collection is a female 
(24 mm.) from Walla Walla, Wash. (June), that is 
almost an exact match, even to the dark fuscous hind 
wing, to Ragonot’s figure 17 of the female in Paris. A 
larger female (82 mm.) in the National Collection from 
Monachee Meadows, Calif., is a good match for 
Ragonot’s figure 16, of the male type, except that the 
hind wing is whitish towards the base as described by 
Hampson. In these two examples the spining of the 
bursa is similar to that of montinatatella (fig. 939). The 
exact status of sciwrella cannot be determined until it 
is reared and more specimens in substantial series are 
assembled. 

Alar expanse, ‘32 mm.”’ 

Tyrer Locatity: California (type in Paris Mus.). 

Foop pirant: Unknown. 


394. Honora dotella Dyar 
FIGURES 448, 938 


Honora dotella Dyar, Pomona College Journ. Ent., vol. 2, p. 377, 
1910.—McDunnough, Check list, No. 6345, 1939. 

Distinguished from mellinella by the darker, more 
suffused (blackish fuscous) ground color and the more 
strongly contrasted whitish costal area of forewing; 
antemedial line thin, white, not continued to costa; 
ocherous patch beyond antemedial line obsolete or 
represented only by a pale trace; basal pale spot clay 
colored, concolorous with thorax; subterminal line ob- 
solete; upper discal spot obliterated; lower discal spot 
black, elongate, completely enclosed by the dark ground 
color and with no trace of a white border. Hind wing 
semihyaline white with a fuscous shade along termen 
and some fuscous shading on the veins, especially in 
female. Alar expanse, 17-24 mm. 

Male genitalia distinguished chiefly by the somewhat 
widened apical portion of cucullus. Bursa of female 
genitalia without detached spines opposite the large, 
spined plate (signum). 

Typr Locatiry: Claremont, Calif. (type in USNM). 

Foop prant: Unknown. F 

Distrisution: California, Claremont, Loma Linda 
(Mar.), San Diego (Mar., May, June). 

Apparently a distinct species, but close to mellinella. 


395. Honora montinatatella (Hulst) 
FicureE 939 


Spermatophthora montinatatella Hulst, Ent. Amer., vol. 3, p. 134, 
1887. 


Honora canicostella Ragonot, N. Amer. Phycitidae, p. 12, 1887. 


UNITED STATES NATIONAL 


MUSEUM BULLETIN 207 


Honora montinatatella (Hulst), Ent. Amer., vol. 5, p. 156, 1889; 
Phycitidae of N. Amer., p. 189, 1890.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 186, 1901.—McDunnough, 
Check list, No. 6348, 1939. 

Forewing brownish red with costal area white dusted 
with reddish scales, the pale costal color spreading into 
the cell at middle of wing and enclosing the blackish 
discal spots, broken near middle of costa by a narrow 
band of the ground color which slants outwardly into 
cell; antemedial line incomplete, a thin line of whitish 
scales from inner margin to cell, set further out on wing 
than in other species; subterminal line obsolete or nearly 
so, when present well back from terminal margin so 
that space between antemedial and subterminal lines is 
considerably reduced (less than one-third the wing 
area); ON some specimens an obscure ocherous patch 
bordermg the antemedial line outwardly near inner 
margin. Hind wing palesmokyfuscous. Alar expanse, 
24-28 mm. 

Male genitalia similar to those of mellinella except 
vinculum narrower at terminal margin, shape similar to 
that of subsciurella (fig. 941). 

TYPE LOCALITIES: Sierra Nevada, Calif. (montinata- 
tella, i AMNH, ex Rutgers); California (canicostella, 
in Paris Mus.). 

Foop puant: Unknown. 

In addition to the Hulst type I have seen two other 
specimens (co' and 9, in USNM), from a subalpine 
meadow on Bogachiel Peak of the Olympic Mts., Wash. 
Apparently montinatatella is a high-altitude species. 


396. Honora perdubiella (Dyar), new combination 
FicurEs 449, 940 


Zophodia perdubiella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
. 37, 1905. 

Pie perdubiella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221, 
1925.—McDunnough, Check list, No. 6299, 1989. 

Pale ashy gray; costal area sparsely dusted with 
whitish scales; transverse lines obsolete; lower discal 
spot at end of cell faintly indicated, blackish. Hind 
wing slightly paler than forewing, unmarked, shiny. 
Alar expanse, 22-26 mm. 

Male genitalia distinguished chiefly by the shape of 
the apical projection of gnathos. Female genitalia 
figured from paratype from the type locality; similar 
to those of montinatatella, especially the genitalia of the 
Mount Olympus female of Hulst’s species. 

TypE LocaLiry: Stockton, Utah (June, type in 
USNM). 

Foop pLiant: Unknown. 

Apparently a distinct species. Dyar’s types are 
rubbed so the extremely pale color of forewing may not 
be true of fresh examples. A female in the National 
Collection from Baker, Nev. (May), is much darker, 
shows faint traces of the normal transverse lines and 
indications of the blackish discal and terminal dots. 
However, there is scarcely any whitish dusting in the 
costal area. The forewing is dark grayish fuscous and 
the hind wing fuscous, with a faintly darker line along 
termen and some dark shading on the veins. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 199 


102. Honorinus, new genus 


TyprE or genus: Honorinus fuliginosus, new species. 

Characters of Honora except: Labial palpus of male 
grooved to hold the maxillary palpus; maxillary palpus 
of male in the form of an aigrette; forewing with vein 2 
from before but near outer angle of cell; hind wing with 
cell shorter, somewhat less than one-third the length of 
wing; transtilla complete; abdomen of male with two 
pairs of ventrolateral hair tufts. 

Very close to Honora. In the absence of any typical 
species of Honora from tropical America and because of 
the divergence of the Peruvian species on male charac- 
ters it seems better to give it a separate generic desig- 
nation than to expand and loosen our definition of 
Honora. 

397. Honorinus fuliginosus, new species 


Figures 450, 947 


Forewing sooty fuscous; transverse lines obsolete or 
nearly so; antemedial line faintly indicated in the male 
near inner margin; beyond it a faint ocherous shade; 
some pale scaling between the veins beyond cell; a 
narrow, obscure ocherous shade along costa from base 
to middle; discal and terminal dots obscure. Hind 
wing semihyaline, whitish fuscous with a dark line 
along termen and some dark shading on the veins; 
veins 3 and 5 stalked for more than half their lengths. 
Alar expanse, 24-26 mm. 

Male genitalia with elements of transtilla solidly 
fused at their apices to form a high, narrowly rounded 
arch. Female genitalia with the membrane of posterior 
lobe of bursa thickened. 

Typ Locatity: Angasmarca, Pert (type in USNM). 

Foop prant: Unknown. 

Described from male type and three male and one 
female paratypes, all from the type locality. In addi- 
tion I have before me another female from Mima, Pert, 
which I take to be the same species; but as it has veins 
3 and 5 of hind wing somewhat longer stalked and a 
slightly smaller genitalia than the female paratype I do 
not include it among the types. The males are rubbed 
and do not show the dark ground color as plainly as the 
female, which is in good condition. 


103. Genus Oncolabis Zeller 


Oncolabis Zeller, Isis von Oken, p. 877, 1848.—Ragonot, Mono- 
graph, pt. 1, pl. 1, fig. 26, 1893; pt. 2, p. xi, 1901—Hampson, 
in Ragonot, Monograph, pt. 2, p. 187, 1901. (Type of genus: 
Oncolabis anticella Zeller.) 

Endommasis Hampson, in Ragonot, Monograph, pt. 2, pp. ix, 
124, 1901. (Type of genus: Endommasis nigritella Hamp- 
son. New synonymy.) 

Tongue well developed. Antenna pubescent; basal 
segment elongate; on male, shaft with a stout hook from 
its basal segment and strongly curved for a few seg- 
ments beyond (fig. 454d); on female simple. Labial 
palpus obliquely ascending; long, extending half its 
length above vertex; smooth scaled; segment 2 in male 
grooved to hold maxillary palpus; segment 3 very short, 
acuminate, not deflected forward. Maxillary palpus 


of male in the form of an aigrette; of female filiform. 
Forewing smooth; narrowly elongate; 11 veins; vein 2 
from near lower outer angle of cell; 3 from the angle; 
4-5 stalked for half their lengths, their stalk very closely 
approximate or connate with 3 at base; 6 from below 
upper angle of cell, straight; 10 from cell, closely ap- 
proximate to the stalk of 8-9 for a short distance; male 
without costal fold. Hind wing with vein 2 from very 
close to lower outer angle of cell; 3 and 5 stalked for 
half their lengths; 7 and 8 contiguous or closely approxi- 
mate for a short distance beyond cell; cell one-third the 
wing length; discocellular vein curved. Eighth ab- 
dominal segment of male with compound ventral tufts. 

Male genitalia with apical process of gnathos a 
stout, elongate, pointed hook. Uncus triangulate; apex 
pointed. Harpe elongate, slender; apex rounded; 
clasper vertical, pointed. Anellus a small triangulate 
cupped plate without lateral lobes. Aedeagus stout, 
straight, moderately long; penis armed with a few short 
sclerotized folds and numerous granulations. Vin- 
culum stout, a trifle longer than broad, not appreciably 
tapering to broad terminal margin. 

Female genitalia with signa strongly developed, 
consisting of a large, round, curved, densely spined 
plate and a varying number of detached spines opposite 
the plate; ductus bursae about the same length as 
bursa, with a ring of sclerotized spinose ridges at its 
junction with bursa, minutely granulose for a short 
distance beyond; genital opening simple; ductus 
seminalis from bursa near its junction with ductus 
bursae. 

Hampson distinguished his Hndommasis from 
Oncolabis chiefly on the shape of the labial palpus, 
upturned (‘‘redressé”) in Hndommasis and obliquely 
erect with third segment projected forward in Oncolabis. 
On the strength of this supposed difference the genera 
were widely separated in the generic key in Ragonot’s 
Monograph (pp. ix and xi). This difference was solely 
due to the position of the palpi at time of death of the 
particular specimens. The normal position of the labial 
palpus is appressed to face with third segment erect. 

Similar male antennae and female genitalia occur in 
Stylobasis Hampson. The latter, however, is distin- 
guished by its forewing venation, vein 4 being absent. 


398. Oncolabis anticella Zeller 
Figures 56, 454, 942 


Oncolabis anticella Zeller, Isis von Oken, p. 877, 1848.—Hampson, 
in Ragonot, Monograph, pt. 2, p. 187, 1901. 

Endommasis nigritella Hampson, in Ragonot, Monograph, pt. 
2, p. 124, 1901 (new synonymy). 

Forewing dark brownish fuscous with basal area 
darker, blackish; costal border (except in suffused 
specimens) whitish finely dusted (under magnification) 
with scattered reddish scales; antemedial line obsolete, 
indicated only by a white spot extending from lower 
fold to inner margin and having a small, central, black 
dot; in fresh specimens the white spot narrowly 
bordered inwardly and outwardly by a faint ocherous 
shading. Suffused specimens show little or no trace of 


200 


the white spot or the white border of costa. Hind 
wing hyaline white with a narrow fuscous shade along 
termen and a similar shading on some of the veins, 
especially on the females. Alar expanse, 13-21 mm. 

Male genitalia with lateral margins of vinculum 
slightly concave; terminal margin very slightly angled; 
clasper sharply pointed. Female genitalia exhibiting 
considerable variety in size of bursa, size of the large 
spined plate of signa, and the number of scattered spines 
opposite the spined plate. Extremes of variation are 
shown in figure 942. 

TYPE LocauitiEs: South Brazil (anticella, in BM); 
Santos, Brazil (ngritella, in BM). 

Foop pirant: Hlephantopus sp. 

DisTRIBUTION: CuBA: Santiago de las Vegas (Sept.). 
GuatemaLa: Cayuga (Aug.), Chejel (Aug.), Volcdin 
Santa Maria (July). Costa Rica: La Florida, Juan 
Vinas (May, Nov.). Panamé: Corozal (Apr.), La- 
Chorrera (May), Parafso (May), Porto Bello (Apr., 
May, Oct., Dec.), Rio Trinidad (Mar.). FREencu 
Guiana: St. Jean Maroni. Contompra: Minca. 
Bottvia: Santa Cruz, Prov. del Sara (Jan., Nov.); 
Brazit: Sao Paulo, Amparo, Santos, Ypianga (Sept.) ; 
Santa Catarina, Santa Catarina Isl. Ecuapor: 
Quevedo (Nov. Dec.). Paraguay: Villarrica (July, 
Aug., Sept.). Argrnrina: “Villa Ana, F. C. S. F.” 
(Jan.). 

Over 60 specimens before me from the National 
Museum, British Museum, Cornell, and Janse Collec- 
tions exhibiting wide variation in size and color, from 
suffused examples corresponding to the figure of 
nigritella in the Ragonot Monograph (pl. 49, fig. 20) 
to that of typical anticella (pl. 48, fig. 25). The male 
genitalia are remarkably uniform. Female genitalia 
vary in individual specimens as indicated above. Such 
variations bear no relation to locality or pattern. The 
food plant record is from two reared specimens (o" and @) 
in the National Museum from Santiago de las Vegas 
(HE. H. A. de Cuba, No. 9627). This is our only known 
host record. 


104. Genus Cabotia Ragonot 


Cabotia Ragonot, Nouv. Gen., p. 30, 1888; Monograph, pt. 2, 
pp. xi, 187, 19801.—Hampson, in Ragonot, Monograph, 
pt. 2, p. 188, 1901. (Type of genus: Cabotia semidiscella 
Ragonot.) 

Encystia Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7, p. 256, 
1901. (Type of genus: Encystia bonhoti Hampson. New 
synonymy.) 

Tongue well developed. Antenna pubescent; basal 
segment elongate; on male shaft compressed and 
strongly curved at base (fig. 452d); on female shaft 
simple. Labial palpus oblique; segment 2 reaching 
almost to vertex, in male grooved to hold the maxillary 
palpus; segment 3 short, deflected forward or slightly 
downward. Maxillary palpus of male in the form of an 
aigrette; of female filiform, somewhat broadly scaled. 
Forewing smooth; 11 veins; 2 from very close to lower 
outer angle of cell; 3 from the angle; 4-5 stalked for at 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


least half their lengths, stalk connate with 3; 6 from 
below upper angle of cell, straight; 10 from the cell, 
at base shortly separated from 8-9; male without 
costal fold. Hind wing with vein 2 from lower outer 
angle of cell, connate with 3; 3 and 5 long stalked (for 
over half their lengths); 7 and 8 anastomosed for 
slightly more than half their lengths beyond cell; cell 
about one-third the length of wing; discocellular vein 
curved. Highth abdominal segment of male with a pair 
of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos an ovate 
plate bearing a small thornlike spine near its posterior 
end. Uncus triangulate; apex pointed. Harpe elon- 
gate; apex evenly rounded; clasper vertical, knobbed. 
Anellus a triangulate, cupped plate. Aedeagus short, 
stout, sharply angled at apex; penis with a few scattered 
granulations, otherwise unarmed. Vinculum stout, as 
long as or a trifle longer than broad, truncate, and of 
nearly equal width throughout. 

Female genitalia with signa well developed, consisting 
of a chain of short thornlike spines on bulbous bases and 
a varying number of similar, scattered, discontinuous 
spines; bursa, otherwise smooth; ductus bursae mem- 
branous; genital opening simple, unsclerotized; ductus 
seminalis from bursa near its junction with ductus 
bursae. 

The genus is compact and sharply defined, easily 
distinguished from nearly related genera by its male 
antenna, signa, and the shape of the terminal projection 
of gnathos. Hampson’s description of Encystia would 
indicate something entirely different, but the description 
is erroneous in a number of details: vein 9 of forewing is 
not absent, but well developed; vein 2 of forewing is 
close to, but not from the angle, and the discocellular 
vein of hind wing is curved and not oblique. I have 
before me two females of his series of bonhoti from 
Nassau and photographs of his holotype and its male 
genitalia, which clearly show that it is a typical Cabotia. 

Specific limits within the genus are difficult to deter- 
mine from the limited and scattered material available. 
I suspect that most of the names will eventually fall 
into synonymy. The genitalic differences exhibited by 
the supposed species are trifling, consisting chiefly of 
the terminal plate of gnathos and the number of spines 
composing the signa, all very doubtful characters sub- 
ject to individual variation. The coloration and pat- 
tern—likewise somewhat individually variable in speci- 
mens from a given locality—are much the same for all 
the species except Dyar’s rhythmatica: Forewing luteous 
more or less shaded with fuscous; antemedial line very 
faintly indicated, well out towards middle of wing, 
bordered outwardly at or near costa by a fuscous blotch 
and inwardly near inner margin by a similar shade; 
subterminal line obscure, parallel and close to termen; 
costal area whitish; a sparse scattering of reddish scales 
over most of the wing; discal dots faint; terminal dots 
distinguishable except at tornus and near apex. Hind 
wing pale smoky fuscous, proportionally darker in dark 
females. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 201 


399. Cabotia semidiscella Ragonot 
Figure 55, 451, 935 


Cabotia semidiscella Ragonot, Nouv. Gen., p. 30, 1888.—Hamp- 
son, in Ragonot, Monograph, pt. 2, p. 188, 1901. 

The alar expanse given by Ragonot is 16 mm. In 
the National Museum are three smaller specimens (11 
mm.) identified by Hampson—a male and female from 
St. Jean Maroni, French Guiana (the genitalia of which 
are figured), and a male from St. Laurent du Maroni, 
French Guiana. This last is an abnormal specimen, 
having veins 2 and 3 of forewing long stalked. Also in 
the National Collection are six females from Los Vas- 
quez, Argentina (15-16 mm.), agreeing with the fore- 
going, and two somewhat darker (fresher) females that 
I take to be the same species from Villarrica, Paraguay 
(Feb., Oct.). Inthe British Museum there is a male and 
nine females from southeastern Brazil (E. D. Jones, 
1920-303”) and three females from Villa Ana, Ar- 
gentina (Oct.). 

Typr Locauity: Goya, Argentina (type in Paris 
Mus.). 

Foop pirant: Unknown. 


400. Cabotia schini (Berg) 


Spermatophthora schini Berg, Anales Soc. Cient. Argentina, vol. 
19, p. 275, 1885. 

Cabotia schini (Berg) Ragonot, Nouv. Gen., p. 30, 1888.—Hamp- 
son, 7n Ragonot, Monograph, pt. 2, p. 188, 1901. 

I have seen nothing identified as this species. How- 
ever, it should be readily identifiable if Argentinian 
examples of Cabotia are ever reared from the pepper- 
tree. Alar expanse, 20 mm. 

Type Locaity: Buenos Aires, Argentina (type lost). 

Foop piant: Schinus molle Linnaeus. 

Apparently known only from the type specimens 
reared from galls on the peppertree. This is our only 
food-plant record for the genus. 


401. Cabotia rhythmatica Dyar 
Ficures 453, 933 


Cabotia rhythmatica Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 339, 
1914, 

Forewings less distinctly marked with fuscous and 
with a more rosy suffusion than those of other species 
inthe genus. Uncus shorter and male genitalia stockier 
than in our examples identified as semidiscella or than in 
males of bonhoti. Alar expanse, 13-14 mm. 

Typr Locauity: Porto Bello, Panamdé (Mar., Apr.; 
type in USNM). 

Foop prant: Unknown. 

Known only from the type specimens. 


402. Cabotia cundajensis (Zeller) 
Fiaure 932 


Euzophera cundajensis Zeller, Horae Soc. Ent. Rossicae, vol. 16, 
p. 227, 1881. 

Euzophera impeditella Zeller, Horae Soc. Ent. Rossicae, vol. 16, 
p. 229, 1881. 

Cabotia cundajensis (Zeller) Ragonot, Monograph, pt. 2, p. 189, 
1901. 


I have seen no Cabotia from Colombia; but in the 
National Museum are three females (15-20 mm.) from 
Castro, Parand, Brazil, identified by both Hampson 
and Schaus as cundajensis. The genitalia are figured 
from one of these. They exhibit nothing that can be 
definitely identified as a specific character. Alar ex- 
panse, 16-22 mm. 

Types: In British Museum (cundajensis, impeditella). 

TypE LocauitiEes: Cundai, Colombia (cundajensis, in 
BM); Viani, Colombia (impeditella, in BM). 

Foop prant: Unknown. 

I suspect that this as well as semidiscella will eventu- 
ally prove to be the same as schini (Berg). 


403. Cabotia bonhoti (Hampson), new combination 
Figures 452, 934 


Encystia bonhoti Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7, 
p. 256, 1901. 

Not appreciably different from what we have identi- 
fied as semidiscella Ragonot from French Guiana, 
except for trifling differences in genitalia which are 
probably not of more than individual significance. 
Male genitalia figured from specimen from Trelawney 
Parish, Jamaica; female genitalia from Zeller specimen 
in British Museum, from Nassau. Alar expanse, 
13-16 mm. 

Typr Locatity: Nassau, Bahamas (type in BM). 

Foop piant: Unknown. 

Distrisution: Bawamas: Nassau (July). Jamaica: 
Newport (Feb.), St. Andrews Parish, Trelawney Parish. 
The Jamaican material consists of 36 specimens in the 
National Collection. 


Genus 105: Canarsia 


[Venational division A. Forewing with 11 veins; 10 from the 
stalk of 8-9; 2 from well before lower outer angle of cell; stalk 
of 4-5 separate from 3 at base. Hind wing with 7 and 8 anas- 
tomosed for a short distance beyond cell; vein 2 from before 
lower outer angle of cell; discocellular vein curved. Labial 
palpus upecurved. Male genitalia with uncus triangulate; harpe 
with short erect clasper; transtilla absent; aedeagus short, stout; 
penis armed with several short spines. Female with a girdle of 
slender spines in bursa; bursa partially sclerotized; ductus 
seminalis from bursa couplatrix.] 


105. Genus Canarsia Hulst 


Canarsia Hulst, Phycitidae of N. Amer., p. 179, 1890.—Ragonot, 
Monograph, pt. 2, p. ix, 1901—Hampson, in Ragonot, 
Monograph, pt. 2, p. 119, 1901. (Type of genus: Ne- 
phopteryx ulmiarrosorella Clemens.) 

Tongue well developed. Antenna pubescent; on 
male shaft with a deep sinus and strong scale tuft at 
base; on female simple. Labial palpus upcurved, rough 
scaled, reaching to vertex; third segment short (less 
than half the length of second), pointed. Maxillary 
palpus squamous, appressed to face. Forewing smooth; 
11 veins, vein 2 from well before lower outer angle of 
cell; 3 from angle; 4 and 5 stalked, the stalk well sepa- 
rated from 3; 6 from below upper angle of cell, straight; 
10 stalked with 8-9; male without costal fold. Hind 


202 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


wing with vein 2 before lower outer angle of cell; 3 from 
angle, closely approximate for some distance with 5; 
7 and 8 anastomosed for a short distance beyond cell; 
cell about one-half the wing length; discocellular vein 
curved. Highth abdominal segment of male with 
compound ventral hair tufts. 

Male genitalia and apical process of gnathos a stout 
hook. Elements of transtilla not distinguishable (un- 
sclerotized); uncus triangulate; apex narrowly rounded. 
Harpe narrowly elongate, of nearly even width through- 
out; apex rounded; a small erect blunt clasper arising 
from sacculus near middle of harpe. Anellus a rather 
large flattened plate with prominent lateral lobes. 
Aedeagus short, stout, strongly elbowed near middle 
and with a scattering of fine spines toward apex; penis 
armed with a short row of spinelike cornuti. Vinculum 
stout, about as broad as long; tapering slightly to 
broadly rounded terminal margin. 

Female genitalia with signa consisting of a girdle of 
long, slender spines with enlarged bases; bursa copu- 
latrix partially sclerotized at junction with ductus 
bursae and emergence of ductus seminalis; ductus 
bursae short (shorter than bursa), sclerotized just before 
genital opening, otherwise simple; genital opening 
simple; ductus seminalis from bursa near its junction 
with ductus brusae. 

An easily recognized genus containing one North 
American species. 


404. Canarsia ulmiarrosorella (Clemens) 
Figures 80, 455, 936 

Nephopteryx (?) ulmiarrosorella Clemens, Proc. Acad. Nat. Sci. 
Philadelphia, 1860, p. 205. 

Stenoptycha pneumatella Hulst, Ent. Amer., vol. 3, p. 137, 1887. 

Psorosa ulmella Ragonot, N. Amer. Phycitidae, p. 13, 1887. 

Honora fuscatella Hulst, Ent. Amer., vol. 4, p. 118, 1888. 

Canarsia ulmiarrosorella (Clemens) Hulst, Phycitidae of N. 
Amer., p. 180, 1890; U. S. Nat. Mus. Bull. 52, p. 431, 
1902.—Hampson, in Ragonot, Monograph, pt. 2, p: 120, 
1901.—Forbes, Cornell Mem. 68, p. 632, 1923—MeDun- 
nough, Check list No. 6332, 1939.—Darlington, Trans. 
Amer. Ent. Soc., vol. 78, p. 91, 1947.—Craighead, U. S. 
Dep. Agr. Mise. Publ. 657, p. 454, 1950. 

Canarsia gracilella Hulst, Canadian Ent., vol. 32; p. 174, 1900.— 
Forbes, Cornell Mem. 68, p. 632, 1923—McDunnough, 
Check list, No. 6333, 1939. (New synonymy.) 

Canarsia feliculella Dyar, Journ. New York Ent. Soc., vol. 15, p. 
110, 1907.—McDunnough, Check list No. 6334, 1939. (New 
synonymy.) 

Forewing pale to dark gray; antemedial line well out 
towards middle of wing, nearly vertical, slightly in- 
dented at vein 1b and above the lower fold, white bor- 
dered outwardly by a blackish line; subterminal line 
parallel with termen, indented at vein 6 and lower fold 
and outwardly angled between them, white, inwardly 
bordered by a blackish line; discal spots confluent, form- 
ing a blackish line along discocellular vein; terminal 
dots confluent forming a more or less continuous black 
line along termen. Hind wings pale smoky fuscous, 
somewhat darker on dark specimens. Alar expanse, 
15-20 mm. 

Genitalic characters as given for the genus, the thin 
spines of the signa slightly bent. 


TypEz Locauities: None indicated (ulmiarrosorella, 
type lost; pnewmatella, in AMNH, ex Rutgers) ; Wiscon- 
sin (ulmella, in Paris Mus.); Blanco County, Tex. 
(fuscatella, in AMNH, ex Rutgers); Montclair, N. J. 
(gracilella, rn AMNH, ex Rutgers); Brownsville, Tex. 
(feliculella, nm USNM). 

Foop pLant: Elm (larva a leaf-tier). Also recorded 
from hickory (Forbes). In the National Museum there 
is a series reared from larvae taken under bark of hack- 
berry; but as larvae wander about after feeding, records 
other than elm should be discounted. 

Distrisution: Unitep Starrs: Maine; New Hamp- 
shire, Hampton (July); Massachusetts, Amherst (June, 
July), Framingham (May); Connecticut, East River, 
New Haven (Aug.) ; New York, Otto (July) ; New Jersey, 
Essex County (June), Montclair (Aug.), New Bruns- 
wick; Pennsylvania, New Brighton (May), Pittsburgh 
(May); Maryland, Hyattsville, Plummers Isl. (May); 
District of Columbia, Washington (Mar., Apr., May); 
Ohio, Dayton (Aug.); Illinois, Chicago, Decatur (Apr., 
June), Lacon (Aug.), Oconee (Aug.); Wisconsin; Iowa, 
Ames (May), Sioux City (May, July); Kansas, Law- 
rence (Aug.), Manhattan (May), Onaga; Tennessee, 
Knoxville (May); Missouri, St. Louis (June, Aug.) ; 
Texas, Blanco County (May, June), Brownsville (May), 
Burnet County, Kerrville (Apr.), Plano (July), San 
Benito (Aug., Sept.), Victoria (May). Canapa: Ontario, 
Trenton (June, July); Quebec, St. Hilaire (June, July) ; 
Nova Scotia, Cape Breton Isl. (July). 

Presumably generally distributed east of the Rocky 
Mountains wherever the elm occurs. 

The names gracilella and feliculella represent nothing 
more than pale color forms and have no racial signifi- 
cance. The holotype of gracilella is a male without ab- 
domen, but. is obviously conspecific with ulmiarrosorelia. 
The female paratype (also in the Rutgers Collection) is 
an Hiphestia. Dyar’s type is a male agreeing in all de- 
tails of genitalia with typical wlmiarrosorella. 


Genus 106: Harnocha 


[Venational division C. Forewing with 10 veins; 9 absent; male 
without costal fold. Hind wing with vein 2 from angle of cell; 
3 and 5 stalked; discocellular vein curved. Labial palpi porrect. 
Transtilla incomplete.] 


106. Genus Harnocha Dyar 


Harnocha Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 337, 1914. 
(Type of genus: Harnocha velessa Dyar.) 

Tongue well developed. Antenna pubescent, shaft 
very slightly bent at base. Labial palpus porrect, 
downcurved; extending slightly over twice the length 
of the head beyond it. Maxillary palpus minute, fili- 
form. Forewing smooth; 10 veins; vein 2 from very 
close to lower outer angle of cell, approximate to 3; 3 
from the angle; 4 and 5 stalked for half their lengths, 
the stalk connate with 3; 6 from below upper angle of 
cell, straight; 8 and 9 united (9 absent); 10 from the 
cell, shortly separated from 8 at base; male without cos- 
tal fold. Hind wing with vein 2 from lower outer angle 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


of cell, connate with 3; 3 and 5 stalked for at least half 
their lengths; 7 and 8 anastomosed for at least half their 
lengths beyond cell; cell less than one-half (nearer one- 
third) the length of the wing; discocellular vein curved. 
Eighth abdominal segment of male simple. 

Male genitalia with apical process of gnathos an ovate 
plate terminating in a spinelike point. Uncus triangu- 
late; apex bluntly pointed. Harpe narrowly elongate; 
apex rounded; clasper absent. Anellus a triangulate, 
flattened plate. Aedeagus moderately stout, sinuate 
(slightly bent at one-third and again at two-thirds); 
penis with a few weak scobinations, otherwise unarmed. 
Vinculum stout, as broad as long, not tapering; terminal 
margin evenly rounded. 

This genus is distinct from but apparently very close 
to Cabotia, with which it agrees on a great many struc- 
tural details of venation and male genitalia. It differs 
in having the antennal shaft less strongly bent, the 
labial palpus porrect rather than oblique, vein 9 of fore- 
wing absent, eighth abdominal segment of male without 
paired hair tufts, and harpe without clasper. It con- 
tains but one known tropical American species. 


405. Harnocha velessa Dyar 
Ficures 111, 456 


Harnocha velessa Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 338, 
1914. 

Forewing luteous with a very faint rosy tint; costa 
from base to apex rather broadly margined with white 
very sparsely dusted with dark fuscous scales; an 
oblique blackish band near base and extending from 
inner margin to white costal stripe; a rather conspicuous 
patch of blackish scales near middle, between cell and 
lower fold; a similar smaller blackish spot at lower outer 
angle of cell. Hind wing semihyaline, whitish, shaded 
with pale fuscous along costa and at apex. Alar ex- 
panse, 11-13 mm. 

Typp tocaniry: La Chorrera, Panamd (May, type 
in USNM). 

Foop puant: Unknown. 

Represented in the National Collection by the type, 
eight other males from the type locality, and one male 
from Rio Trinidad, Panama (June). The female is 
unknown. 


Genera 107-109: Eurythmasis to Wunderia 


{Venational division B. Forewing with 10 veins; 9 present; 2 
and 3 stalked or united, from lower outer angle of cell. Hind 
‘wing with 2 from lower outer angle of cell; 3 and 5 stalked for at 
least half their lengths; 7 and 8 strongly anastomosed. Uncus 
triangulate. Transtilla incomplete.] 


107. Genus Eurythmasis Dyar 


Eurythmasis Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 338, 1914. 
(Type of genus: Hurythmasis ignifatua Dyar.) 

Tongue well developed. Antenna pubescent; basal 

segment elongate; on male shaft with a hook from its 


203 


basal segment and curved for a few segments beyond; 
on female simple. Labial palpus oblique, long, extend- 
ing more than half its length above vertex; smooth; 
segment 2 on male grooved to hold the maxillary palpus; 
segment 3 very short, acuminate. Maxillary palpus 
of male in the form of an aigrette; of female filiform. 
Forewing smooth; 10 veins; veins 2 and 3 long stalked, 
from lower outer angle of cell; 4 absent; 5 separated 
from 2-3 at base; 6 from below upper angle of cell, 
straight; 8 and 9 stalked; 10 from cell, not approximate 
to 8-9; male without costal fold. Hind wing with vein 
2 from lower outer angle of cell, connate with the stalk 
of 3-5; 3 and 5 stalked for half their lengths; 7 and 8 
anastomosed for half their lengths; cell one-third the 
wing length; discocellular vein curved. Highth ab- 
dominal segment of male with compound ventral tufts. 

Male genitalia with apical process of gnathos an 
elongate, rather narrow hook. Uncus triangulate; apex 
pointed. Harpe elongate, slender; apex rounded; 
clasper vertical, pointed, slightly curved. Anellus a 
small, weakly sclerotized, cupped plate. Aedeagus 
moderately stout, nearly straight; penis with a few 
minute spines and granulations, otherwise unarmed. 
Vinculum stout, a trifle longer than broad, not appre- 
ciably tapering to broad terminal margin. 

Female genitalia with signa strongly developed, con- 
sisting of a large, curved, oval (or round), densely spined 
plate and a few detached spines near or opposite the 
plate; ductus bursae approximately the same length as 
bursa, minutely scobinate for a short distance from 
junction with bursa; genital opening simple; ductus 
seminalis from bursa near its junction with ductus bursae. 

In habitus and all structural characters except vena- 
tion this genus resembles Oncolabis, to which it is ap- 
parently closely related. It contains one tropical 
American species. 


406. Eurythmasis ignifatua Dyar 
Ficures 95, 457, 945 


Eurythmasis ignifatua Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 338, 1914. 

Forewing gray brown; a white border along costa 
sparsely dusted with reddish scales; on middle of inner 
margin a narrowly elongate, dull ocherous patch with a 
knoblike projection at each end extending to the cell; 
a few reddish scales on dark ground color at base of 
wing and bordering inner margin; transverse lines ob- 
solete. Hind wing semihyaline; in female with some 
fuscous shading on the veins. Alar expanse, 13-14 mm. 

Genitalia as given for the genus; male with terminal 
margin of vinculum slightly angled. 

Type tocatiry: La Chorrera, Panamaé (type in 
USNM). 

Foop prant: Unknown. 

DistriputTion: PanamA: La Chorrera (May). Purr- 
to Rico: Adjuntas (June), Bayamén (Mar., Aug.), 
Lavis (Oct.), San German (Apr., Aug.), Utuado. Cusa: 
Oriente, Loma del Gato (Sept.). 


204 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


108. Genus Eurythmidia Ragonot 


Eurythmidia Ragonot, Monograph pt. 2, p. xii, 1901. Hampson, 
in Ragonot, Monograph, pt. 2, p. 208, 1901. Janse (in 
part), Journ. Ent. Soc. South Africa, vol. 7, p. 4, 1944. 
(Type of genus: Hurythmia ignidorsella Ragonot.) 

Tongue well developed. Antenna pubescent; on 
male, a row of 4 or 5 rough scales on shaft beyond basal 
segment; of female simple. Labial palpus, upturned, 
extending well above vertex; third segment about half 
the length of second, pointed. Maxillary palpus 
squamous, appressed to face. Forewing smooth; 10 
veins; veins 2 and 3 united, from lower outer angle of cell, 
connate with the stalk of 4-5; 4 and 5 stalked for at 
least half their lengths; 6 from below upper angle of 
cell, straight; 8 and 9 stalked; 10 from cell approximate 
to 8-9 at base, thence divergent; male without costal 
fold. Hind wing with vein 2 from the lower outer angle 
of cell, connate with the stalk of 3-5; 3 and 5 long 
stalked (for more than half their lengths); 7 and 8 
anastomosed for nearly two-thirds of their lengths; cell 
short, less than one-third the wing length; discocellular 
vein curved. Eighth abdominal segment of male simple. 

Male genitalia with apical process of gnathos an 
elongate, narrow hook. Uncus triangulate; apex nar- 
rowly rounded. Harpe elongate, slender; apex rounded; 
clasper vertical, short, pointed, weakly sclerotized. 
Anellus a flattened plate. Aedeagus moderately slen- 
der, straight; penis unarmed. Vinculum stout, as 
broad as long, not tapering, truncate. 

Female genitalia with bursa copulatrix elongate; 
signa strongly developed, consisting of a curved, round, 
densely spined plate and a few detached spines near the 
plate; ductus bursae somewhat shorter than bursa, 
minutely scobinate near junction with bursa, the sco- 
binations extending into bursa for a short distance; a 
narrow sclerotized plate behind (and above) genital 
Opening; genital opening otherwise simple; ductus 
seminalis from bursa near its junction with ductus 
bursae. 

The genus is obviously closely related to Hurythmasis, 
with similar habitus and genitalia, differing chiefly in 
venation, the simple eighth segment of male, and the 
simple shaft of the male antenna. It contains one 
species common to the southwestern United States and 
Central America. I mterpret the venation of forewing 
differently from Hampson and Ragonot. They con- 
sider that 4 is absent and 3 and 5 stalked. From the 
affinities of the genus I believe that 4 is present and 
stalked with 5 and 2 and 3 united. By either interpre- 
tation the end result would be the same; but Huryth- 
midia seems to belong with the genera where the 
tendency is for 2 and 38 to fuse. 

The description and figures published by Janse apply 
to Hurythmidia only so far as the female characters are 
concerned. The two specimens from Amula, México, 
which he had before him were from the Druce Collection 
and bore Hampson’s identification labels. The female 
is Hurythmidia ignidorsella but the male is not. It is 
a somewhat rubbed example of Elasmopalpus lignosellus 
Zeller. Evidently Hampson did not check the vena- 


tion, for it is normal for Hlasmopalpus, vein 4 being 
present in both hind wings. Dr. Janse kindly sent me 
the specimens and his slides for examination and has 
asked me to make the correction to his description. 


407. Eurythmidia ignidorsella (Ragonot) 
Ficures 91, 459, 943 
Eurythmia ignidorsella Ragonot, N. Amer. Phycitidae, p. 16, 
1887.—Hulst, Phycitidae of N. Amer., p. 196, 1890. 
Eurythmidia ignidorsella Hampson, in Ragonot, Monograph, pt. 
2, p- 208, 1901.—Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 
338, 1914—McDunnough, Check list, No. 6361, 1939. 

Forewing blackish gray; a white border along costa, 
sparsely dusted with reddish scales; on middle of inner 
margin a whitish orange patch, the orange shade ex- 
tending more or less along inner margin toward base of 
wing; antemedial line obsolete; subterminal line very 
faint (frequently absent), when distinguishable, whitish. 
Hind wing semihyaline, more or less shaded with fuscous 
along costa and outer margin. Alar expanse, 12-16 
mm. 

Genitalia as given for the genus; terminal margin of 
vinculum straight; spined plate of signa rather small. 

Typr LocaLiry: Arizona (type in Paris Mus.). 

Foop piant: Unknown. 

Distrisution: Unirep Staves: Arizona, Paradise 
(Sept.), Prescott (Sept.), M#xtco: Orizaba, Guerrero, 
Amula (May), Cayuga (Apr., May, Aug.), Panamé: 
Alhajuelo (Mar., Apr.), Cafio Saddle (Gatin Lake, 
May), Corazal (Mar.), Paraiso (Apr.), Porto Bello 
(Feb., Mar., May). Reported from additional locali- 
ties in México by Ragonot and Hampson. 

Strikingly similar in color and markings to Huryth- 
masis ignifatua but easily separable on genitalia and 
venation. 


109. Genus Wunderia Grossbeck 


Wumnderia Grossbeck, Bull. Amer. Mus. Nat. Hist., vol. 37, art. 1, 
p. 133, 1917. (Type of genus: Wunderia neaeriatella Gross- 
beck.) 

Tongue well developed. Antenna of female simple, 
weakly pubescent. Labial palpus slender, upturned, 
reaching above vertex, third segment acuminate, 
slightly more than half as long as second. Forewing 
smooth; 10 veins; veins 2 and 3 united, from lower 
outer angle of cell, connate with the stalk of 4-5; 4 and 5 
stalked for half their lengths; 6 from below upper angle 
of cell, straight; 8 and 9 stalked; 10 from the cell, 
separate from 8-9 at base. Hind wing with vein 2 
from the stalk of 3-5; 3 and 5 long stalked; 7 and 8 
anastomosed for nearly two-thirds of their lengths; cell 
open (discocellular vein absent). 

Female genitalia with bursa copulatrix without sig- 
num and simple except for some minute scobinations at 
its posterior end, the scobinations extending into the 
ductus bursae for over half its length; ductus bursae 
with a narrow sclerotized plate behind (and above) 
genital opening; genital opening otherwise simple; 
ductus seminalis from bursa near its junction with 
ductus bursae. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


The genus and its type species were described from 
a single female. The male is unknown, so the above 
diagnosis is incomplete. The habitus of neaeriatella is 
that of a pale ignidorsella so the placement of Wunderia 
near Hurythmidia seems safe enough. 

The genus is readily identified by the open cell of hind 
wing, the otherwise Hurythmidia venation, and the 
absence of asignum. Grossbeck’s description is faulty 
in one respect. The front is not “projected forward in 
the form of a cone.”’ The scaling projects in a conical 
tuft, but the front itself is evenly rounded. 


408. Wunderia neaeriatella Grossbeck 
Figures 90, 948, 949 


Wunderia neaeriatella Grossbeck, Bull. Amer. Mus. Nat. Hist., 
vol. 37, art. 1, p. 133, 1917—McDunnough, Check list, 
No. 6361-1, 1939. 

Markings and color of forewing similar to those of 
Eurythmidia ignidorsella except slightly paler. The 
female type is somewhat rubbed, which accounts for 
the traces of a dark transverse antemedial line and the 
dark discal markings mentioned by Grossbeck. Before 
me are three females from the Cornell Collection, col- 
lected at San German, Puerto Rico, Apr. 16 and 17, 
1930. In these the entire area below the whitish costal 
stripe is pale grayish fuscous without transverse lines 
or appreciable discal spots, and there is a very faint 
orange blotch on middle of dorsal margin (as in 2gni- 
dorsella) distinguishable on one of the specimens. The 
whitish costal stripe is sparsely dusted with reddish 
scales and there is a scattering of similar scales on the 
dark area. The Puerto Rican examples are certainly 
congeneric and I believe conspecific with the Florida 
type. 

There are some differences in the female genitalia 
(shown in figs. 948, 949); but these are merely in the 
size and shape of the bursa, differences which are prob- 
ably of no more than individual significance. The 
Puerto Rican specimens are rather small (12-12.5 mm.) 
compared with the type (14.5 mm.). 

Type wocauity: Everglades, Fla. (Apr., type in 
AMNB#). 

Foop piant: Unknown. 

The male is unknown. 


Genera 110-114: Oedothmia to Cacozophera 


[Venational division D. Forewing with 10 veins; 9 present; 4 
absent; vein 2 from before but near lower outer angle of cell. 
Hind wing with discocellular vein curved. Labial palpi up- 
turned or oblique. Transtilla incomplete or altogether absent] 


110. Genus Oedothmia Hampson 


Oedothmia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 60, 
1930. (Type of genus: Oedothmia endopyrella Hampson.) 

Synothmia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 61, 
1930. (Type of genus: Synothmia bahamasella Hampson. 
New synonymy.) 


Clarke and Tams have compared the types of Hamp- 
son’s two supposed genera and inform me that they 


205 


appear to be no more than sexes of one species. Hamp- 
son’s chief character for separating Oedothmia and 
Synothmia was the shape of the frons, conical on 
Oedothmia and round on Synothmia; but Clarke states 
that his description of the former is at variance with 
the type in several details—the male antenna is simple, 
the shaft not excised at base, and the frons isround, not 
pointed. The venation of the types of the two genera 
is similar. 

Tongue well developed. Antenna pubescent; shaft 
simple. Labial palpus upturned; the second segment 
reaching above vertex; third segment “thickly scaled.” 
Maxillary palpus filiform. Forewing smooth; 10 veins; 
vein 2 from very near to lower outer angle of cell; 3 
from the angle, approximate to 2 for some distance from 
cell; 4 and 5 united, shortly separated from 3 at base; 
6 from below upper angle of cell, straight; 8 and 9 
stalked for about half their lengths; 10 from the cell, 
separated from 8-9 at base; male without costal fold. 
Hind wing with vein 2 from lower outer angle of cell; 
3 from the angle, contiguous (but not fused) to 5 for 
half their lengths; 7 and 8 contiguous beyond cell for 
some distance, but not anastomosed; cell short, less 
than one-third the length of wing; discocellular vein 
curved. 

Male genitalia unknown; the type of O. endopyrella, 
and only known male, is without abdomen. 

Female genitalia resembling those of Hurythmidia 
and Oncolabis; with the signa strongly developed, the 
large spined plate situated in bursa like that of Oncolabis 
but the collar of strong spines at junction of bursa and 
ductus is lacking; genital opening simple; ductus semi- 
nalis from bursa near its junction with ductus bursae. 

The genus is easily distinguishable from its nearest 
relatives in venational division D (except Stylobasis) by 
the approximate condition of veins 2 and 3 of forewing. 
From Stylobasis it is distinguished by the contiguous 
rather than anastomosed condition of veins 7 and 8 of 
hind wing, its upturned labial palpi and simple male 
antennae. 

409. Oedothmia endopyrella Hampson 


Figures 102, 944 


Oedothmia endopyrella Hampson, Ann. Mag. Nat. Hist., ser. 10, 
vol. 5, p. 61, 1930. 

Synothmia bahamasella Hampson, Ann. Mag. Nat. Hist., ser. 10, 
vol. 5, p. 61, 1930 (new synonymy). 

Markings and color similar to those of Oncolabis 
anticella Zeller, from which it can be distinguished most 
readily by its venation. Alar expanse, 16-18 mm. 

Typr Locauitizs: Vera Cruz, México (endopyrella, 
in BM); Nassau, Bahamas (bahamasella, in BM). 

Known only from the two types. 


111. Genus Stylobasis Hampson 


Stylobasis Hampson, in Ragonot, Monograph, pt. 2, pp. xii, 198, 
1901. (Type of genus: Stylobasis rubripurpurea Hampson.) 
Tongue well developed. Antenna pubescent; basal 
segment enlarged; on male shaft with a hook from its 
basal segment and deeply curved for several segments 


206 


beyond (as in Oncolabis), otherwise flattened, on female 
simple. Labial palpus oblique; long, second segment 
extending above vertex, on male grooved to hold the 
maxillary palpus; third segment porrect (deflected for- 
ward). Maxillary palpus filiform. Forewing smooth; 
10 veins; vein 2 from near lower outer angle of cell; 3 
from the angle, approximate to 2 in the male, (closely 
approximate or connate with 5 in the female); 4 absent; 
6 from slightly below upper angle of cell, slightly curved 
in male (straight in the female); 8 and 9 stalked; 10 
from the cell, approximate to 8-9 at base; male without 
costal fold. Hind wing with vein 2 from lower outer 
angle of cell connate with the stalk of 3-5; 3 and 5 
stalked for half or (in female) slightly more than half 
their lengths; 7 and 8 anastomosed for about half their 
lengths (more shortly anastomosed in female) beyond 
cell; cell slightly under one-third the wing length; dis- 
cocellular vem curved. The venation except for the 
absence of vein 4 of forewing is strikingly similar to that 
of Oncolabis, especially in the female. Eighth abdom- 
inal segment of male with compound ventral tuft. 


Male genitalia similar to those of Oncolabis and 
Hurythmasis except for slightly longer vinculum (a 
difference of, at most, specific significance). 

Female genitalia similar to those of Oncolabis; signa 
strongly developed, consisting of a large, round, curved, 
densely spined plate and a few similar detached spines 
near the plate; a fused collar of similar spines surround- 
ing the ductus bursae at its junction with bursa copu- 
latrix; ductus bursae as long as bursa, minutely scob- 
inate for about half its length beyond the collar; genital 
opening simple; ductus seminalis from bursa near its 
junction with ductus bursae. 


410. Stylobasis rubripurpurea Hampson 
Ficurss 106, 458, 946 
Stylobasis rubripurpurea Hampson, in Ragonot, Monograph, pt. 
2, p. 198, 1901. 

The moth has the general habitus of Oncolabis 
anticella; forewing purplish gray (or dark grayish 
fuscous) with whitish costal streak sparsely dusted with 
reddish scales; transverse lines obsolete, the antemedial 
indicated below cell only by an outwardly bordering 
pale yellowish patch. Hampson’s description men- 
tions faint blackish streaks from the base of costa and 
on the median vein and one distinct discal spot. The 
figure in Ragonot (pl. 49, fig. 19) is misleading. It 
shows a form with the veins darkly outlined, a distinct 
whitish subterminal line, and no trace of the yellowish 
patch on inner margin, or of the pale costal streak. Our 
female in the National Collection is rubbed so that the 
dark ground color shows plainly only along the veins, 
thus corresponding with Ragonot’s figure. Alar ex- 
panse, 16-19 mm. 

Genitalia as given for the genus. 

TYPE Locauity: “Jrazu, Mexique” (type in Trans- 
vaal Mus.). 

Foop eiant: Unknown. 

In addition to the male type Hampson mentions a 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


male from Santos, Brazil. These, in addition to the 
female from Juan Vinas, Costa Rica (Apr., in USNM), 
are the only specimens of the species known to me. 

Through the courtesy of Dr. Janse I was able to see 
and figure the genitalia of the male type. He also 
submitted a drawing of the wing venation which is re- 
produced here. The femele in the National Museum 
tentatively identified by Schaus as rubripurpurea differs 
in slight details of venation from the male type, as indi- 
cated in the foregoimg generic description, and may or 
may not be conspecific. However, from its labial palpi 
and the sum of its characters I believe it is properly 
placed. 


112. Genus Diviana Ragonot 


Diviana Ragonot, Nouv. Gen., p. 27, 1888; Monograph, pt. 2, 
pp. xii, 201, 1901.—Hulst, Phycitidae of N. Amer., p. 190, 
1890. (Type of genus: Diviana eudoreella Ragonot.) 

Dannemora Hulst, Phycitidae of N. Amer., p. 212, 1890.—Hamp- 
son, 7m Ragonot, Monograph, pt. 2, pp. xiii, 209, 1901. 
ean of genus: Dannemora edentella Hulst. New synony- 
my. 

Tongue short and weak, but exposed. Antenna 
pubescent; on male, shaft curved toward base, some 
rough scaling in the curve (but no teeth or other spine- 
like projections). Labial palpus upturned, short, barely 
reaching to vertex; the second segment tufted in front; 
the third short, acuminate. Maxillary palpus squa- 
mous (not filiform as stated by Ragonot and Hampson). 
Forewing smooth; subtriangular, rather broad towards 
termen; 10 veins; vein 2 from before the angle of the 
cell; 3 and 5 closely approximate or connate from the 
angle; 4 absent; 6 from below upper angle of cell, 
straight; 8 and 9 stalked; 10 normally from the cell and 
approximate to the stalk of 8-9, rarely connate or very 
shortly stalked with 8-9; male without costal fold. 
Hind wing with vein 2 from before but near lower outer 
angle of cell; 3 and 5 shortly stalked, from the angle; 
7 and 8 anastomosed for a short distance beyond cell 
(for half or less than half their lengths); cell less than 
half the length of wing; discocellular vein curved. 
Eighth abdominal segment of male simple. 

Male genitalia with apical process of gnathos a small 
knob terminating in a short spine. Uncus as broad as 
long, lateral margins parallel, terminal margin bluntly 
angled. Transtilla absent (elements not distinguish- 
able). Harpe short, slender, very slightly tapering to 
narrowly rounded apex; costa’ broadly sclerotized 
throughout and terminating in a short, slender spine. 
Anellus a broad, deeply and widely cleft plate. Ae- 
deagus long, moderately stout, strongly sclerotized, 
with a few scobinations toward apex; penis armed with 
two stout, straight cornuti (slightly less than one-third 
as long as aedeagus). Vinculum very stout, longer than 
combined tegumen and uncus and considerably longer 
than broad; very slightly tapering; terminal margin 
broad, straight. 

Both Diviana and Dannemora were described from 
males. As far as I know there are no known females of 
either type species. In the Ragonot Monograph the 
two supposed genera are widely separated in the generic 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 207 


key on the alternative of vein 10 of forewing from the 
cell or from the stalk of 8-9, an unreliable character at 
best and in this instance thoroughly misleading. In 
two males before me (perfect matches for the figure of 
eudoreella in the Monograph, pl. 23, fig. 12) vein 10 is 
both ways, from the cell on three forewings and dis- 
tinctly stalked on one (compare figs. 103 and 1032). 
On the Hulst type of edentella it is stalked. 

Hampson’s characterization of Dannemora in the 
Monograph is false in a number of details. He included 
in the genus (in addition to its type species) Huzophera 
quadripuncta Zeller, of which he had only females, and 
selected ‘‘characters’”’ from both species and added a 
“character” possessed by neither, namely, a circular 
hyaline depression in the cell on the underside of hind 
wing. There is no such structure in edentella or quadri- 
puncta. The male of the latter has a depression in the 
cell of hind wing, but it is not hyaline, is on the upper 
surface of the wing, and is filled with dense, modified 
sex-scaling. Hampson did not know the male, and the 
hind wing of the female is simple, so it is difficult to see 
where he got this ‘“‘character.”’ Zeller’s species is not 
even closely related to edentella. I am treating it else- 
where (p. 276). Ragonot’s characterization of Diviana 
is misleading in one particular. He states that there are 
some teeth (‘‘dents’’) in the sinus of the shaft of the 
male antenna. Such a character is present in Palatka 
nymphaeella (Hulst); but I do not think he could have 
had an example of this species before him unless his 
description of ewdoreella is incorrect and his figure of it 
completely false. 


411. Diviana eudoreella Ragonot 
Figures 103, 463 


Diviana eudoreella Ragonot, Nouv. Gen., p. 27, 1888; Monograph, 
pt. 2, p. 201, 1901.—Hulst, Phycitidae of N. Amer., p. 190, 
1900.—McDunnough, Check list, No. 6358, 1939. 

Dannemora edentella Hulst, Phycitidae of N. Amer., p. 212, 
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 209, 


1901.—McDunnough, Check list, No. 6362, 1939. (New 
synonymy.) 

Diviana eudoriella Hulst, U. 8. Nat. Mus. Bull. 52, p. 4383, 1902 
(misspelling). 


Forewing blackish brown, the median area heavily 
dusted with white, giving it an ashy gray appearance; 
antemedial white line nearly straight, slanting out- 
wardly from costa to inner margin, slightly notched at 
lower fold, bordered outwardly by a blackish line which 
expands to a triangular blotch at costa, a similar, some- 
what smaller blotch bordering the line at inner margin; 
subterminal white line parallel to termen, notched at 
vein 6 and at lower fold, towards costa bordered in- 
wardly by a fuscous streak; discal dots at end of cell 
distinct, blackish; entire outer area between subterminal 
line and termen blackish brown. Hind wing pale smoky 
fuscous, with a darker line along termen and some very 
faint dark shading on the veins. Alar expanse, 16-19 
mm. 

Male genitalia as given for the genus. 
Type Locauities: ‘‘America Septentrionalis’ (eu- 


doreella, in Paris Mus.); Florida (Apr.; edentella, in 
AMNH, ex Rutgers). 


Foop pLant: Unknown. 


Known only from males. The Hulst type is badly 
tubbed and shows little trace of original markings; but 
its genitalia agree in every detail with those of two well- 
marked specimens (in USNM) from Orlando, Fla. 
(Feb.), and Plummers Isl., Md. (July). These two 
specimens are in perfect agreement with Ragonot’s 
figure of eudoreella. 


113. Genus Palatka Hulst 


Palatka Hulst, Canadian Ent., vol. 24, p. 62, 1892; U. S. Nat. 
Mus. Bull. 52, p. 433, 1902. (Type of genus: Divinia nym- 
phaeella Hulst.) 

Tongue reduced (better developed in female than 
male). Antenna pubescent; shaft of male with a 
shallow sinus at base containing a weak scale tuft and 
a couple of short teeth concealed within the tuft (these 
teeth are strongly sclerotized, pointed, dorsal projections 
from two of the segments); shaft of female simple. 
Labial palpus oblique, extending well above vertex; 
broadly scaled; segment 2 long; segment 3 about one- 
third the length of 2, acuminate. Maxillary palpus 
filiform. Forewing smooth; 10 veins; vein 2 from before 
the angle of the cell; 3 from the angle; 4 absent; 5 from 
slightly above the lower angle and separate from 3; 6 
from below upper angle, straight; 8 and 9 stalked; 10 
from the cell separate at base from stalk of 8-9; male 
without costal fold. Hind wing with vein 2 from well 
before lower outer angle of cell; 3 and 5 from the angle, 
very closely approximate towards base; 7 and 8 closely 
approximate for half their distance beyond cell; cell 
about one-half the length of wing; discocellular vein 
curved. Eighth abdominal segment of male simple. 

Male genitalia with apical process of gnathos an 
elongate, narrow hook. Uncus longer than broad, with 
terminal margin broadly rounded. ‘Transtilla incom- 
plete, elements minute. Harpe elongate, narrow, grad- 
ually tapering to bluntly pointed apex; sacculus pro- 
duced at extremity into blunt, free arm directed towards 
costa. Anellus a broad flattened plate. Aedeagus 
long, moderately stout, smooth; penis armed with two 
stout cornuti, about half as long as aedeagus. Vincu- 
lum stout, slightly longer than broad, tapering to 
broadly rounded terminal margin. 

Female genitalia with bursa copulatrix scobinate but 
without signa, thickened (but not sclerotized) at ter- 
minal end; ductus bursae shorter than bursa, strongly 
sclerotized from junction with bursa for a little more 
than half its length, on ventral surface at genital open- 
ing a broad, strongly sclerotized, triangulate plate; 
ductus seminalis from bursa near its junction with 
ductus bursae. 

The genus is quite distinct from others of venational 
group D and easily identified by its male and female 
genitalia. It is closest to Diviana, from which it is 
readily separable by its hind wing venation, veins 3 and 


208 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


5 approximate rather than stalked. It contains only 
one North American species. 


412. Palatka nymphaeella (Hulst) 
Ficures 104, 462, 954 


Diiana nymphaeella Hulst, Canadian Ent., vol. 24, p. 62, 1892. 

Palatka nymphaeella (Hulst), Canadian Ent., vol. 24, p. 62, 1892; 
U. S, Nat. Mus. Bull. 52, p. 433, 1902.—Barnes and Mc- 
Dunnough, Contributions, vol. 3, p. 222, 1917.—McDun- 
nough, Check list, No. 6360, 1939. 

Diviana verecuntella Grossbeck, Bull. Amer. Mus. Nat. Hist., 
vol. 37, art. 1, p. 132, 1917. 

Forewing grayish ocherous with whitish transverse 
line and more or less marked with brownish fuscous; in 
paler specimens the ocherous tint predominates, in 
darker one, the gray; antemedial line outwardly trans- 
verse from costa, indented at median and lower folds, 
in well-marked (pale) specimens bordered inwardly on 
lower half by a large dark patch; subterminal line 
rather near and parallel to termen, very slightly in- 
dented at vein 6 and lower fold and with some dark 
streaks bordering it near costa; discal spots distinct, 
blackish; a row of blackish dots along termen. Hind 
wings pale smoky fuscous. Alar expanse, 16-18 mm. 

Genitalia as given for the genus. 

Typr Locatities: Charlotte Harbor, Fla. (nymphae- 
ella, in AMNH, ex Rutgers); Fort Myers, Fla. (vere- 
cuntella, i AMNH). 


Foop piuant: Unknown. 


Distrisution: Florida, Charlotte Harbor (Apr.), 
Everglades (Apr.), Fort Myers (Apr.); Texas, San Be- 
nito (Apr.); Connecticut, East River (July). 

The Hulst type in the Rutgers Collection is only a 
fragment. One pair of wings, the antennae, and the 
abdomen are missing; but there is no doubt of what it 
represents. The alar expanse given by Hulst (21 mm.) 
is too great. It is nearer 18 mm. The Connecticut 
specimens (one male and six females in the National 
Collection) are darker than those from Florida and 
Texas, more grayish, and with little or no dark shading 
or blotches bordering the transverse lines. They show 
no genitalic differences. 


114. Genus Cacozophera Dyar 


Cacozophera Dyar, Ins. Insc. Menstr., vol. 7, p. 58, 1919. 
(Type of genus: Cacozophera venosa Dyar.) 

Tongue reduced but exposed. Antenna of female 
pubescent, simple. Labial palpus upturned, short, 
not reaching vertex; third segment acuminate, shorter 
than second. Maxillary palpus filiform. Forewing 
smooth; narrowly elongate; 10 veins; vein 2 from well 
before outer angle of cell; 3 and 5 connate from the angle; 
4 absent; 6 from below upper angle of cell, straight; 8 
and 9 stalked; 10 from the cell, approximate to the 
stalk of 8-9 for some distance. Hind wing with vein 
2 from well before lower outer angle of cell; 3 and 5 con- 
nate from the angle;7 and 8 anastomosed for most of their 
lengths (8 very short); cell less than one-half the 
length of the wing; discocellular vein curved. 


Female genitalia with bursa copulatrix large, finely 
scobinate throughout, the scobinations extending part 
way into the ductus bursae; signa present, consisting 
of a cluster of small, slender disks (three in the only 
specimen available); ductus bursaze slightly shorter 
than bursa, unsclerotized; genital opening simple; 
ductus siminalis from bursa in the neighborhood of the 
signa. 

The genus and its type species were erected on a 
single female. Dyar characterized Cacozophera merely 
as having the venation of his genus Anthropteryz, 
differing only in having the “wings long and narrow, 
trigonate, the apex pointed, not short and square.” 
Anthropteryx itself was also erected on a single female 
and unfortunately is a freak with vein 4 absent from 
one forewing. Dyar did not notice the other forewing 
in which vein 4 was present and stalked with 5, the 
normal condition for the specimen. For further dis- 
cussion of Anthropteryx see page 313. The placement of 
Cacozophera is tentative. Its relationship to the other 
genera cannot be determined until a male is discovered. 


413. Cacozophera venosa Dyar 
Fiaures 105, 950 


Cacozophera venosa Dyar, Ins. Insc. Menstr., vol. 7, p. 58, 1919. 


Forewing brownish fuscous, the area between costa 
and cell and vein 6 uniformly dark; below vein 6 and 
from end of cell the veins darkly outlined and the inter- 
spaces between them contrastingly paler; subterminal 
line distinct from vein 6 to inner margin, whitish, 
evenly curved and parallel with termen; no dis- 
tinguishable antemedial line or discal spots; a faint 
yellowish white shade along inner margin and (under 
magnification) a scattered dusting of dull rosy scales on 
the dark areas; terminal dots blackish, large, inter- 
venular. Hind wing smoky fuscous, the veims and 
terminal margin darker. Alar expanse, 19 mm. 

Genitalia as given for the genus. 

TyprE Locatity: Cayuga, Guatemala (May, type in 
USNM). 

Foop piant: Unknown. 

Known only from the female type. 
species but of uncertain affinities. 


Obviously a good 


Genera 115-117: Psorosina to Paconius 


[Venational division A, Forewing with 11 veins; 10 from the 
cell or connate or shortly stalked with stalk of 8-9; 4 and 5 
approximate, connate or very shortly stalked. Hind wing with 
vein 2 from before lower outer angle of cell; discocellular vein 
curved or incomplete. Male genitalia with costa of harpe 
produced; cornuti developed; transtilla incomplete or absent.] 


115. Genus Psorosina Dyar 


Psorosina Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 113, 
1904.—Forbes, Cornell Mem. 68, p. 632, 1923. (Type of 
genus: Psorosina angulella Dyar.) 

Tongue well developed. Antenna pubescent; shaft 
of male with sinus and large scale tuft at base; shaft of 
female simple. Labial palpus upcurved, slender, rough 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 209 


scaled, reaching to vertex in male, to slightly above in 
female; third segment about half the length of second, 
acuminate. Maxillary palpus squamous. Forewing 
smooth; 11 veins; vein 2 from before lower outer angle 
of cell; 3 from the angle; 4 and 5 connate (or very 
shortly stalked), separated from 3 at base; 6 from 
below upper angle of cell, straight; 8 and 9 stalked; 
10 from cell, separated from stalk of 8-9; male without 
costal fold. Hind wing with vein 2 from before (but 
near) lower outer angle of cell; 3 and 5 stalked; 7 and 
8 anastomosed for half their lengths beyond cell; cell 
about half the length of wing, partially open; dis- 
cocellular vein incomplete. Eighth abdominal segment 
of male with two pairs of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos a stout 
hook. Uncus triangulate. Transtilla incomplete, ele- 
ments minute. Harpe elongate, narrow, not appre- 
ciably tapering; apex rounded; strongly sclerotized 
costa projecting at apex into a short, sharp spine; 
clasper erect, short, weakly sclerotized. Anellus a 
curved U-shaped plate with broad base. Aedeagus 
short, stout, nearly straight; penis armed with a single, 
long, moderately slender cornutus (nearly as long as 
aedeagus). Vinculum stout, about as broad as long, 
slightly tapering to truncate terminal margin. 

Female genitalia with bursa copulatrix scobinate 
over most of its inner surface; signum developed as an 
irregularly shaped spined plate near junction of ductus 
bursae; ductus bursae shorter than bursa, with a pair 
of narrow, sclerotized, strongly spined plates along the 
lateral margins, some minute scobinations in the inter- 
vening area, a fingerlike, ventral, sclerotized plate from 
genital opening, and a few granulations on the inner 
wall above genital opening; ductus seminalis from 
bursa near its junction with ductus bursae. 

A distinct genus with one North American species; 
easily distinguished by venation and genitalia. 


414. Psorosina hammondi (Riley) 
Fieurts 62, 461, 952 


Pempelia hammondi Riley, Fourth annual report on the noxious, 
beneficial and other insects, of the State of Missouri, p. 44, 
1872. 

Psorosa hammondi (Riley) Ragonot, Ent. Amer., vol. 5, p. 116, 
1889. 

Canarsia hammondi (Riley) Hulst, Phycitidae of N. Amer., p. 
180, 1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 
120, 1901. 

Psorosina hammondi (Riley) Dyar, Proc. Ent. Soc. Washington, 
vol. 6, p. 113, 1°04.—Forbes, Cornell Mem. 68, p. 682, 
1923.—McDunnough, Check list, No. 6335, 1939. 

Psorosina angulella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 113, 
1904.—Forbes, Cornell Mem. 68, p. 632, 1923.—McDun- 
nough, Check list, No. 6336, 1939. (New synonymy.) 


Forewing glossy brownish fuscous (purplish brown 
in some lights) ; antemedial line grayish white, straight, 
slightly curved or weakly angled at cell, slanting a trifle 
from costa to inner margin, of varying width depending 
on the amount of white scaling and sometimes (but 
rarely) partially divided by a fine median dark line; 
subterminal line obscure or indistinguishable, fine, 
whitish, nearly vertical from costa to tornus, very 


faintly bordered by blackish lines; on costa adjacent to 
inner border of the subterminal line a more or less 
extended and triangulate grayish white patch; blackish 
discal and terminal dots obscure, the discal dots more 
or less confluent. Hind wings smoky fuscous, glossy. 
Alar expanse, 13-14 mm. 

Genitalia as given for the genus; terminal margin of 
vinculum broadly and shallowly notched. 


Type tocauitins: Illinois (hammondi, in USNM); 
Iowa (angulella, in USNM). 

Foop pxiants: Apple, pear, Prunus maritima. 
(Specimens in USNM also labeled ‘from sycamore” 
and ‘‘from acorns,”’ but these are very dubious records.) 
The larva is a leaf skeletonizer. 

DistrrBuTion: Connecticut, East River (Aug.) ; Rhode 
Island, Weekapaug (Aug., Sept.); Ohio, Cincinnati 
(Aug.) ; Indiana, Bedford (July) ; Jilinois, Oconee (July, 
Aug.); Iowa; Kansas, Wathena (July), Wichita (June) ; 
Missouri, several specimens with no further locality, 
St. Louis (Aug.). Rather generally distributed 
throughout the eastern and central United States and 
Canada. 

Worn females of this species are superficially similar 
to small specimens of Moodna ostrinella (Clemens) and 
the two species have been occasionally confused. They 
are quite distinct, however, on both venation and 
genitalia. The character on which Dyar separated his 
angulella (its “strongly angled’ antemedial line) is 
rather imaginary than real. The pale markings on the 
forewing of hammondi are formed by rather thinly 
spread, whitish scales and vary from specimen to 
specimen in size, shape, and intensity. Dyar’s type is 
rubbed, a condition which accounts in part for the 
angulate condition of the antemedial line, a very slight 
angulation at that. Its genitalia agree in every detail 
with those of typical hammondt. 


116. Patriciola, new genus 


Typr or Genus: Patriciola semicana, new species. 

Tongue well developed. Antennae simple in both 
sexes, shortly pubescent. Labial palpus porrect (beak- 
like), long, projecting about four times the length of 
head beyond it; alike in both sexes. Maxillary palpus 
squamous, large; alike in both sexes. Forewing smooth; 
11 veins; vein 2 from well before the lower outer angle 
of the cell; vein 3 remote from 2 but before the angle of 
the cell; 4 from the angle approximate to 5 for some 
distance from cell; 6 from below upper angle of cell, 
straight; 8 and 9 stalked; 10 connate or very shortly 
stalked with 8-9; male without costal fold. Hind wing 
with vein 2 from well before the lower outer angle of the 
cell; 3 from near but before the angle; 5 from the angle; 
7 and 8 contiguous (touching but not completely fused) 
for nearly half their lengths beyond cell; cell about half 
the length of the wing; discocellular vein curved. 
Eighth abdominal segment of male simple. 

Male genitalia with apical process of gnathos a rather 
broadly based, blunt hook produced anteriorly into 
a long, flat, sclerotized apron (the whole process resem- 


210 


bling, in outline, a bulbus-necked bottle). Uncus 
broader than long; sides nearly parallel ; terminal margin 
broadly rounded. Transtilla incomplete; elements mi- 
nute, folded. Harpe short; appressed clasper well out 
towards outer margin of harpe; from base of costa a 
long, stout, curved, strongly sclerotized and pointed 
clasperlike horn projects across face of harpe. Anellus 
a flattened plate. Aedeagus short, stout, slightly sinu- 
ate, and with apex sharply constricted; armed with a few 
short spines near apex; penis with a small patch of fine 
scobinations, otherwise unarmed. Vinculum stout, as 
broad as long, tapering to truncate terminal margin. 

Female genitalia with bursa copulatrix finely scob- 
inate throughout, otherwise simple, without signum; 
ductus bursae sclerotized, broad and very short, broad- 
ening abruptly into the sclerotized and thickened lower 
lip to an exceptionally wide genital opening; ductus 
seminalis from bursa near its junction with ductus 
bursae. 

A striking genus easily identified by venation and 
genitalia. Contains one North American species. 


415. Patriciola semicana, new species 


Fieures 84, 460, 953 


Forewing ashy grayish white on costal half; shaded 
with faun brown rather heavily dusted with blackish 
fuscous on lower half, especially from base to antemedial 
line and slightly beyond; veins partially outlined by 
dark scaling; antemedial line distinct only from cell to 
inner margin, far out towards middle of wing, white, 
narrow, evenly incurved; subterminal line very faint, 
nearly obsolete, parallel to and rather far in from 
termen; lower discal dot at end of cell, large, distinct 
and blackish, a similar smaller black spot in cell over 
the antemedial line. Hind wing semihyaline, smoky 
white; veins not darkened and terminal margin only 
faintly so; cilia concolorous with wing. Alar expanse, 
25-26 mm. 

Genitalia as given for the genus. 

TYPE Locality: Provo, Utah (type in USNM, 
61367). 

Foop prant: Unknown. 


Described from male type and two female paratypes 
from the type locality, collected by Tom Spalding July 
16, 1909 (co), and Aug. 4, 1908 (¢@). In habitus sem- 
icana resembles most some specimens of Honora but is 
easily separable from anything in that genus. 


117. Paconius, new genus 


TypPE oF GENus: Paconius corniculatus, new species. 

Tongue well developed. Antenna pubescent; shaft 
of male with sinus and small scale tuft at base. Labial 
palpus upcurved, rough scaled, reaching to vertex; third 
segment about half the length of second, acuminate. 
Maxillary palpus squamous. Forewing smooth; 11 
veins; vein 2 from before the lower outer angle of cell; 
3 from very close to angle; 4 and 5 approximate for some 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


distance from cell; 6 from below upper angle of cell, 
straight; 8 and 9 stalked; 10 from the cell, shortly 
separated from the stalk of 8-9; male without costal 
fold. Hind wing with vein 2 from before lower outer 
angle of cell; veins 3 and 5 contiguous for a short dis- 
tance from angle (touching but not fused at any point); 
7 and 8 contiguous for less than half their lengths beyond 
cell; cell about half the length of wing; discocellular 
vein curved. Highth abdominal segment of male with 
a weak pair of ventrolateral hair tufts. 

Male genitalia with apical process of gnathos nar- 
rowly triangulate and acutely pointed. Uncus irregu- 
larly triangulate, tapering to narrowly rounded apex. 
Transtilla absent; elements not distinguishable. Harpe 
divided to base with entire costa developed as a strongly 
sclerotized, long, hornlike projection; remainder of 
harpe much reduced, triangulate and tapering to a 
point; clasper absent. Anellus a narrow, slightly 
curved plate with very long, slender, lateral arms. 
Aedeagus long, slender, abruptly bent toward apex; 
armed at apex with two thornlike spurs; penis armed 
with a cluster of short, rather stout spines. Vinculum 
stout, slightly longer than broad, very slightly tapering 
to broad, evenly rounded, terminal margin. 

Female unknown. 

Paconius has much the same venation, palpi, and 
antennae as the European genus Psorosa; but differs 
markedly in genitalia. In Psorosa the harpe is simple 
and the gnathos, vinculum, and anellus entirely different. 
Paconius has several features in common with Patriciola, 
with which it appears to be most closely related, dif- 
fering most from that genus in its upcurved rather than 
porrect palpi. I dislike very much describing a new 
genus without having females for completion of the 
diagnosis on genitalic characters; but it is so obviously 
new and the male genitalia so different from anything 
else in our fauna that it seems advisable to give it some 
designation. 


416. Paconius corniculatus, new species 
FIGURE 464 


Forewing pale ashy gray; transverse lines nearly ob- 
solete; antemedial line faintly outlined, nearly straight, 
slanting outwardly from costa to inner margin, indicated 
chiefly by a small blackish spot on its inner border at 
inner margin and a couple of short blackish streaks on 
its outer border near costa; costal edge at base of wing 
blackish; subterminal line not defined except by a few 
blackish, inwardly bordering streaks near costa. Hind 
wing semihyaline white with a narrow fuscous shade 
along costa and outer margin. Alar expanse, 19 mm. 

Male genitalia as given for the genus. 


Typr Locatity: San Germdn, Puerto Rico (type in 
Cornell Univ.; paratype in USNM, 61368). 

Foop prant: Unknown. 

Described from male type and one male paratype 
from the type locality collected Apr. 16 and 17, 1930, 
under Cornell lot 795, sub. 34 and 36. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 211 


Genera 118-120: Aptunga to Cassiana 


[Venational division A. Forewing with 11 veins; 10 from cell, 
8 and 9 stalked, 6 straight, 4 and 5 stalked, 2 and 3 separate or 
approximate. Hind wing with 7 veins; 7 and 8 strongly anasto- 
mosed, 3 and 5 approximate at base or stalked, discocellular vein 
curved. Male genitalia with uncus subtriangular, tapering 
abruptly, apical half narrow, bluntly pointed; gnathos ter- 
minating in a small, bifid, hooked process; transtilla incomplete, 
represented by a pair of separate, elongate plates; harpe simple, 
elongate, apex obliquely rounded; vinculum stout, decidedly 
longer than broad; aedeagus moderately stout, straight; penis 
without cornutus. Female genitalia with ductus seminalis from 
bursa.] 


118. Aptunga, new genus 


Typs or Genus: Vitula macropasa Dyar. 

Tongue well developed. Antenna pubescent, shaft 
simple in both sexes. Labial palpus obliquely up- 
turned. Maxillary palpus filiform. Forewing nar- 
rowly elongate, smooth; veins 2 and 3 closely approxi- 
mate from lower outer angle of cell; stem of 4-5 approxi- 
mate to 3; vein 10 from the cell, rather well separated 
from stalk of 8-9; male without costal frold or other 
sexual modifications. Hind wing with vein 2 from well 
before angle of cell; 3 and 5 closely approximate at 
lower, outer angle of cell; 7-8 anastomosed for most of 
their lengths beyond cell (free end of vein 8 very short 
and weak); cell less than half the length of wing. 
Abdomen of male with a strong pair of ventrolateral 
hair tufts from eighth segment. 

Male genitalia with apical process of gnathos small, 
bifid; aedeagus simple (without longitudinal ventral 
sclerotized ridge). 

Female genitalia without signum; bursa copulatrix 
more or less finely scobinate in the area about attach- 
ment of ductus seminalis, otherwise smooth; ductus 
bursae smooth except for a weak sclerotization near 
genital opening and some fine scobinations at junction 
with bursa; ductus seminalis from middle or near 
middle of bursa. 

This genus and the two genera following are closely 
related to each other and Mescinia, agreeing in nearly 
all male genitalic characters but differing in venation. 
Aptunga is similar in hind wing venation to Mescinia, 
differing from the latter in having veins 2 and 3 of 
forewing closely approximate at base rather than 
stalked, and lacking the signum in bursa of the female. 


417. Aptunga macropasa (Dyar), new combination 
Ficures 85, 465, 958 


Vitula macropasa Dyar, Ins. Inse. Menstr., vol. 7, p. 61, 1919. 


Forewing grayish fuscous; antemedial line obscure, 
whitish, bordered outwardly by a diffused dark shade; 
subterminal line narrow, whitish, decidedly slanting 
and slightly angled near middle, shaded inwardly and 
outwardly by dark streaks on the veins; a row of small 
blackish dots along termen; discal dots at end of cell 
separate, somewhat elongate; under magnification costa 
shows a peppering of reddish scales. Hind wing 
whitish, semihyaline, the veins outlined with fuscous 


and a fuscous shade bordering costa and along terminal 
margin. Alar expanse, 19-23 mm. 

Male genitalia with weak sclerotized granulations 
and wrinklings on penis. Female genitalia with a 
narrow band of sclerotized granulations on inner 
dorsal surface of ductus bursae at genital opening. 

TypE tocauity: Purulhé, Guatemala (type in 
USNM). 

Foop prant: Unknown. 

Distripution: GuatemMAua: Chejel (Aug.), Purulha 
(July), Voleén Santa Maria (Nov.). M*x1co: Orizaba. 


418. Aptunga imperfecta (Dyar), new combination 
FicurE 957 


Mescinia inperfecta Dyar, Ins. Insc. Menstr., vol. 7, p. 60, 1919. 


Considerably shorter and somewhat paler than 
macropasa, the general color (in the rubbed type) more 
brownish than grayish fuscous; no trace of reddish 
scaling in costal area of forewing; discal dots fused into 
a single round dark spot. Hind wing darker, less 
hyaline than that of macropasa. Alar expanse, 14 mm. 

Female genitalia with the ductus bursae appreciably 
but not strongly sclerotized and granulate for a short 
distance from genital opening. 

Typr Locatity: Cayuga, Guatemala (Apr., type in 
USNM). 

Foop piant: Unknown. 

Known only from the female type; described by 
Dyar in Mescinia but ruled out of that genus by its 
forewing venation and. lack of signum. 


119. Anderida, new genus 


Typr or Genus: Euzophera sonorella Ragonot. 

Tongue well developed. Antenna pubescent, shaft 
simple in both sexes. Labial palpus obliquely up- 
turned. Maxillary palpus filiform. Forewing narrowly 
elongate, smooth; vein 2 from near angle of cell but 
well separated from 3; 10 from cell, separate from 
stalk of 8-9; male without costal fold or other sexual 
modifications. Hind wing with 3 and 5 stalked; 7-8 
anastomosed for at least two-thirds the length of vein 
7; cell approximately half the length of wing. Abdomen 
of male with strong pair of ventrolateral hair tufts 
from eighth segment. 

Male genitalia as in Aptunga except apical process of 
gnathos partially fused. 

Female genitalia with signum; ductus seminalis from 
anterior end of bursa; ductus bursae with a thin 
sclerotized shield on venter at genital opening and more 
or less finely scobinate towards bursa, otherwise smooth. 


419. Anderida sonorella (Ragonot), new combination 
Ficurss 467, 959 


Euzophera senorella Ragonot, N. Amer. Phycitidae, p. 14, 1887; 
Monograph, pt. 2, p. 59, 1901. 

Euzophera sonorella Ragonot, Nouv. Gen., p. 4, 1888 (correction 
of spelling). 

Eyzophera placidella Dyar, Proc. Ent. Soc. Washington, vol. 10, 


212 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


p. 115, 1908.—MeDunnough, Check list, No. 6314, 1939. 
(New synonymy.) 

Forewing ashy gray on costal half shading to white 
on costa; lower half of wing shaded with ocherous gray; 
rather narrow antemedian and subterminal bands 
strongly contrasted, blackish fuscous, antemedian band 
slightly curved, nearly vertical, ending just before costa; 
discal dots small, blackish, the upper one obscure and 
frequently absent. Hind wing whitish with a faint 
gray or ocherous tint. Alar expanse, 16-22 mm. 

Female genitalia with signum very weak, consisting 
of a cluster of 3 to 5 small disks; bursa copulatrix finely 
scobinate. 

TypE Locauitiss: ‘Senora”’ [sic], México (sonorella, 
in Paris Mus.); Yuma County, Ariz. (placidella, in 
USNM). 

Foop piant: Unknown. 

Disrrisution: Unirep Starrs: Arizona, Huachuca 
Mts., Santa Catalina Mts., Redington, Yuma County 
(Colorado Desert). México: Sonora. 

In some of the female specimens before me from 
Redington, Ariz., the transverse dark markings are 
rather faint, almost obsolete in a few examples and 
there is considerable variation in size, but the general 
habitus and genitalia easily distinguish the species. 


120. Cassiana, new genus 


Type or Genus: Vitula malacella Dyar 

Tongue well developed. Antenna pubescent; in male 
with a row of 6 or 7 short slender spines along outer side 
of shaft towards base, shaft somewhat swollen. Labial 
palpus upturned; in female slightly oblique. Maxillary 
palpus filiform. Forewing smooth; venation as in 
Aptunga, veins 2-3 closely approximate from angle of 
cell; male with a strong costal fold terminating at basal 
third in an outwardly projecting brush of scales. Hind 
wing with veins 3-5 stalked; 7-8 anastomosed for most 
of their lengths (free end of vein 8 a short, weak spine); 
cell about half the length of wing. Abdomen of male 
with a strong pair of ventrolateral hair tufts from eighth 
segment. 

Male genitalia with apical process of gnathos small, 
bifid; aedeagus with a sclerotized, longitudinal, ventral 
ridge for over half its length from base. 

Female genitalia with sisnum; ductus seminalis from 
anterior end of bursa. 


420. Cassiana malacella (Dyar), new combination 
Ficures 466, 955, 956 
ieee malacella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 408, 
914. 

Forewing pale grayish fuscous, the area between cell 
and costa white with a faint, scattered dusting of red- 
dish scales; costal edge from base to one-third (the extent 
of the costal fold) blackish; transverse antemedial line 
obsolete; subterminal line but faintly indicated. Hind 
wing whitish, the veins outlined by dark scales, and a 
narrow smoky border along termen and costa. Alar 
expanse, 15-17 mm. 


Male genitalia with lateral margins of vinculum con- 
cave, terminal margin straight. Female genitalia with 
signum variable, consisting of from one small platelike 
projecting disk to a line of similar disks (examples of 
the extreme types in two Puerto Rican specimens are 
shown in figs. 955, 956); bursa and considerable part of 
ductus bursae finely scobinate. 

Typz Locatity: Tehuacén, México (type in USNM). 

Foop pranr: Unknown. 

Disrrisution: Mrxico: Tehuacén (May). PuErro 
Rico: Bayamén (Sept.), Puerto Real (Apr.), Rio 
Piedras (Sept.). Virrern Isuanps: Kingshill (St. Croix, 
Mar., Dec.). 


Genus 121: Mescinia 


[Venational division B. Forewing with 11 veins: 10 from cell, 
8 and 9 stalked, 4 and 5 stalked, 2and 3 stalked. Male genitalia 
with transtilla incomplete.] 


121. Genus Mescinia Ragonot 


Mescinia Ragonot, Monograph, pt. 2, p. viii, 1901—Hampson, 
in Ragonot, Monograph, pt. 2, p. 83, 1901. (Type of genus: 
Ephestia commatella Zeller.) 

Tongue well developed. Antenna pubescent; shaft 
slightly thickened towards base in male, simple in 
female. Labial palpus obliquely upturned, reaching to 
vertex. Maxillary palpus filiform. Forewing smooth, 
narrowly elongate; termen slanting; 11 veins; veins 2 
and 3 from angle of cell, stalked; 4 and 5 stalked; 6 
straight; 8 and 9 stalked; 10 from the cell, separate from 
the stalk of 8-9; male with a short costal fold at base of 
wing. Hind wing with vein 2 from well before lower 
angle of cell; 3 and 5 from angle and closely approximate 
at base; 6 from upper angle of cell; 7 and 8 strongly 
anastomosed beyond cell (in small species completely 
fused or showing the free part of vein 8 as a short spur 
to costa); discocellular vein curved; cell one-third to 
two-fifths of wing length. 

Male genitalia with uncus subtriangular, tapering 
abruptly from broad base, terminal half narrow and 
apex bluntly pointed (except in indecora), outer surface 
covered with bristlelike scales. Gnathos terminating 
in a small, or moderately small, bifid apical process 
(except in indecora). Harpe simple, with costa sclero- 
tized for most of its length. Anellus U-shaped with 
long lateralarms. Aedeagus straight, moderately stout, 
a strong, longitudinal, sclerotized ridge on ventral side. 
Penis more or less scobinate and frequently with sclero- 
tized patch and wrinklings. Vinculum stout, consider- 
ably longer than broad (except in indecora) and with 
terminal margin truncate. 

Female genitalia with bursa copulatrix membranous, 
finely scobinate over part of inner surface (especially 
about signum and near ductus bursae); signum nor- 
mally an oval or round sclerotized plate covered with 
short, blunt or bluntly pointed thorns, or a single stout 
platelike thorn (indecora), or absent (discella). Ductus 
bursae membranous except for a slight sclerotization 
near genital opening, finely scobinate toward bursa. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 213 


Ductus seminalis from anterior end of bursa or from 
bursa near its juncture with ductus bursae. 

The genus Mescinia is closely related to the foregoing 
three genera (Aptunga, Anderida, Cassiana) and to 
Nonia, all of which have similar genitalia, especially in 
the male, but which are readily separable on venation. 
Mescinia is an American genus without, as far as I know, 
any properly included exotic species. Hampson (in 
Ragonot, Monograph, pt. 2, pp. 84, 86) includes one 
Indian species, micans Hampson, but this obviously is 
wrongly referred for it lacks vein 10 of forewing and 
has 4 and 5 from the cell and not stalked as they should 
be for Mescinia. 

Our species divide into three groups on size as follows: 

Alar expanse less than 15 mm. 

Alar expanse more than 15 but less than 20 mm. 

Alar expanse more than 20 mm, 
The first two groups are typical Mescinia on all struc- 
tural characters, but the two large species in the third 
group are somewhat abnormal, especially indecora, 
which is aberrant on both male and female genitalia. 
The other species (discella) has normal male genitalia, 
but the female lacks a signum, and in both the anasto- 
mosis of veins 7 and 8 of hind wing is considerably 
shorter than in normal Mescinia. 

Among the species of the first group the color and 
pattern differences are trifling, and the distinctions 
stressed by Dyar between his species are as much due 
to the condition of the specimens as to differences in 
color or intensity of dark scaling. Genitalic differences 
are also slight, but apparently more reliable. The best 
characters are found in the signa, the apical process of 
gnathos, and the armature of the penis. With the few 
specimens from widely scattered localities and the 
scanty biological information available, it is almost im- 
possible to define specific limits in this group with any 
certainty. 


Genus Mescinia, Species 421-428: M. triloses to 
M. berosa 


[Alar expanse less than 15 mm.] 


421. Mescinia triloses Dyar 
Figures 468, 965 


Mescinia triloses Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 341, 
914. 

Mescinia mosces Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 341, 
1914 (new synonymy). 

Forewing sordid white, the costal area paler and con- 
trasted; dark markings pale brownish fuscous; discal 
dots more or less confluent; antemedian line obscure; 
subterminal dark line rather well marked in unrubbed 
specimens; in fresh examples a dark streak along outer 
half of fold (replacing the absent vein 1c). Hind wing 
whitish, the veins faintly outlined by dark scaling; 
discocellular vein slightly curved. Veins 2 and 3 of 
forewing are stalked for about one-third and veins 4-5 
for approximately one-half their lengths. Alar ex- 
panse, 10-14 mm. 


Tyrer Locauity: Taboga Isl., Panama (Triloses and 
Mosces, in USNM). 

Foop prant: Unknown. 

DistripuTion: PanamA: Alhajuelo (Apr.), Corazal 
(Feb.), Taboga Isl. (Feb.). 

The specimens of Dyar’s mosces are a trifle darker and 
more strongly marked than those of triloses, but they 
represent only the least rubbed of a series collected on 
the same date at the same locality and exhibit no struc- 
tural differences of any significance in genitalia. I am 
unable to distinguish any trace of the ‘‘faintly pinkish” 
color which Dyar mentions in his descriptions. 


422. Mescinia pandessa Dyar 
Figure 469 


Mescinia pandessa Dyar, Ins. Inse. Menstr., vol. 7, p. 60, 1919. 


In color and markings similar to the females of 
mosces. Possibly nothing more than a variety or race 
of triloses, but exhibiting a few slight structural differ- 
ences: Narrower harpes, a somewhat stronger scobina- 
tion on penis, and a wider spacing of the bifid apical 
process of gnathos. Such differences are hardly signifi- 
cant, but in the absence of females it is safer to retain 
the names in specific rank for the present. Veins 2 and 
3 of forewing stalked for two-thirds and 4 and 5 for 
one-half or slightly more than one-half of their lengths. 
Discocellular vein of hind wing as in triloses. Alar 
expanse, 12-13 mm. 

Type tocauity: Cayuga, Guatemala (Apr., type in 
USNM). 

Foop puant: Unknown. 

Known only from the types series from Cayuga. 
The specimens in the National Collection are all males. 


423. Mescinia bacerella Dyar 
Ficures 94, 470, 962 


Mescinia bacerella Dyar, Ins. Inse. Menstr., vol. 7, p. 59, 1919. 


General color somewhat darker than triloses or 
pandessa; costal area of forewing more contrasted, 
whitish with a scattering of reddish brown scales. 
Genitalia showing only minute differences from those of 
preceding species. Veins 2 and 8 of forewing stalked 
for two-thirds and 4 and 5 stalked for one-half their 
lengths. Discocellular vein of hind wing very slightly 
curved. Alar expanse, 12 mm. 

Typr LocaLity: Havana, Cuba (type in USNM). 

Foop piant: Unknown. 

DisrrisuTion: Cusa: Havana, Sierra Maestra (Jan.). 


424, Mescinia estrella Barnes and McDunnough 
Figures 473, 963 
Mescinia estrella Barnes and MeDunnough, Contributions, vol. 2, 
p. 182, 1913.—McDunnough, Check list, No. 6322, 1939. 
Wing pattern similar to that of other species in this 
group, but (in reared examples especially) dark mark- 
ings more intense and whitish costal area of forewing 
more conspicuously dusted with reddish scales. Male 
genitalia similar to those of pandessa except bifid apical 


214 


process of gnathos (fig. 473) somewhat more widely divi- 
ded (as in parvula). 'The spining of the female signum is 
almost identical to that of triloses (compare figs. 468 
and 963). Veins 2 and 3 and 4 and 5 of forewing 
stalked from slightly less than one-half to about one- 
half their lengths, discocellular of hind wing decidedly 
curved. The length of the stalking of the forewing 
veins should not be given too much importance in 
identifying species, for it is a variable character in any 
considerable series of any given species. Alar expanse, 
12-14 mm. 

Typr Locatity: Hverglade, Fla. (type in USNM). 

Foop pruants: Melanthara radiata (rearings of the 
Special Survey of the Div. Foreign Plant Quar., U. S. 
Bureau of Entomology and Plant Quarantine, Nos. 
24851, 24867); Bidens (larva in the flower heads). 

DistrisuTion: Florida, Everglade (Apr.), Key West 
(Apr.), Sugar Loaf Key (Mar.), Sarasota (Jan., Feb., 
Mar.). 

425. Mescinia moorei, new species 
Ficure 472 


In color and superficial appearance most resembling 
bacarella but differing in details of genitalia, a somewhat 
broader extension of the whitish costal margin of forewing 
and less contrasted dark border of the subterminal line. 
The outstanding character seems to be the narrow, 
evenly spaced, parallel, and somewhat elongate prongs of 
the bifid apical process of gnathos (fig.472c). Veins 2 and 
3 of forewing are stalked for about one-third and veins 
4-5 for approximately half their lengths. Alar expanse, 
11 mm. 

TypEr Locauity: Mon Repos, British Guiana (type in 
USNM, 61369). 

Foop puant: Seeds of Wulffia. 

Described from male type and female paratype from 
the type locality reared by H. W. B. Moore, for whom the 
species is named. There are no dates on the specimens 
and they are in poor condition, somewhat broken though 
not badly rubbed. I should not have described them 
had they not been reared and been previously identified 
as parvula Zeller and that name probably been given to 
Mr. Moore. The species should be easily recognized if 
other specimens from the host plant are discovered in 
British Guiana. 


426. Mescinia parvula (Zeller) 
Fieures 471, 960 


Ephestia parvula Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 249, 
1881 


Mescinia parvula (Zeller) Hampson, in Ragonot, Monograph, 
pt. 2, p. 85, 1901. 

Similar to the other species of the group in color and 
markings, except for the much darker, more contrasted 
lining of the veins of hind wing. Differs strikingly from 
all other described species of the genus in the angulate 
projection from the costa of harpe. Alar expanse, 12 
mm. 


Type Locatiry: Honda, Colombia (type in BM). 
Foop PLant: Unknown. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Disrrisution: Cotomsta: Honda. 

There are no examples of parvula in the U.S. National 
Museum and the series in the British Museum is prob- 
ably mixed, as Hampson mentions a specimen from 
Colombia with veins 2 and 3 from the cell. 


427. Mescinia commatella (Zeller) 
Ficure 961 
Ephestia commatella Zeller, Horae Soc. Ent. Rossicae, vol. 16, 


p. 247, 1881. 
Mescinia comatella (Zeller) Ragonot, Monograph, pt. 2, p. 84, 
1901. 


Known to me only from description and figure of the 
female type. Apparently distinguished by the incom- 
plete but strong white antemedian fascia, slanting 
obliquely from inner margin to top of cell and parallel 
with outer margin. Alar expanse, 13 mm. 

The female genitalia of the type shows a single small 
disklike signum. 

Tyrer Locatity: Ceiba, Colombia (type in BM). 

Foop prant: Unknown. 


428. Mescinia berosa Dyar 
Figure 966 


Mescinia berosa Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 341, 1914. 


The female type from which the original description 
was drawn is faded and somewhat rubbed. A fresher 
female example from Puerto Rico shows the dark areas 
of forewing a dark vinous fuscous and the whitish costal 
area somewhat speckled with rufous scales; discal and 
terminal dots blackish. Hind wing smoky, the veins, 
apical area, and terminal margin darker. Female geni- 
talia with bursa copulatrix narrowly elongate; signum a 
large elongate patch of many stout, bluntly pointed, 
thornlike spines; ductus seminalis from bursa near junc- 
ture of bursa and ductus bursae. Veins 2 and 3 of fore- 
wing stalked for about two-thirds and 4 and 5 for 
slightly more than half their lengths. Discocellular 
vein of hind wing decidedly curved. Alar expanse, 12 
mm. 

Typr uocatity: Rio Trinidad, Panamé (type in 


USNM). 
Foop puant: Unknown. 
DistrisutTion: PanamAé: Rio Trinidad (Mar.). 


Purrto Rico: El Semil, near Villalba, elevation 1,700 
feet (May). 

Known to me only from two female examples in the 
National Collection. 


Genus Mescinia, Species 429: M. peruella 


[Alar expanse more than 15, less than 20 mm.] 


429. Mescinia peruella Schaus 
Fieures 474, 964 

Mescinia peruella Schaus, Proc. Ent. Soc. Washington, vol. 29,. 

p. 186, 1927.—Wille, Rey. de Ent., Brazil, vol. 4, p. 455, 
1934. 

Forewing gray with costa whitish nearly to apex; 

under magnification showing a sparse, scattered dusting: 

of rufous scales; transverse markings and discal spots: 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 215 


hardly distinguishable. Hind wing white, translucent, 
extreme apex and terminal margin faintly smoky. 
Male genitalia with apical process of gnathos mod- 
erately large (proportionally about twice the size of 
that organ in the foregoing species group). Female 
genitalia with spines of signum bluntly and broadly 
rounded. Veins 2 and 3 and 4 and 5 of forewing stalked 
for about one-third their lengths (in some specimens 2-3 
stalked for nearly one-half). Discocellular vein of hind 
wing very slightly curved. Alar expanse, 15-18 mm. 

Typr Locauity: Cafiete Valley, Peri (type in 
USNM). 

Foop pxiant: Cotton (larvae feeding in bolls). 

Distrisution: Pert: Cafiete Valley (Mar.), Lima 
(Sept.). 


Genus Mescinia, Species 430 and 431: M. discella 
and M. indecora 
[Alar expanse over 20 mm.] 


430. Mescinia discella Hampson 
Ficures 475, 967 


Mescinia discella Hampson, in Ragonot, Monograph, pt. 2, p. 84, 
1901. 


Forewing a pale purplish fuscous on lower half, 
yellowish (bronzy) above, shading to white on outer 
half of subcostal area; costal edge yellowish brown 
with some blackish scales towards base; a bright 
orange-yellow patch towards end of cell; along median 
fold a fine line of blackish scales; a similar blackish line 
along the fold replacing vein 1c; from end of cell to sub- 
terminal line blackish dashes on veins 2 and 3, 4 and 5 
and 6, more or less fusing to form a large, conspicuous, 
dark blotch; discal spots fused into a blackish line along 
discocellular vein; a row of faint black dots along 
termen; antemedial line very faint, narrow, white, out- 
wardly angled at lower margin of cell and inwardly 
angled at vein 1b; subterminal line sharply angled 
inward at vein 6, thence curved outward around the 
blackish blotch, disappearing near costa and inner 
margin. Hind wing whitish, semihyaline; the veins 
and terminal margins faintly and finely outlined with 
purplish ocherous; veins 7 and 8 anastomosed for little 
more than half the length of vein 7; discocellular vein 
decidedly curved. Alar expanse, 20-21 mm. 

Male genitalia with bifid apical process of gnathos 
elongate, rather large; anellus with enlarged base and 
curved lateral arms; penis with a strong cluster of 
sclerotized folds. Female genitalia with signum re- 
placed by a concentration of scobinations; ductus 
bursae weakly sclerotized towards genital opening; 
ductus seminalis from bursa near its junction with 
ductus bursae. 

TYPE LOcALITY: Jalapa, México (type in USNM). 

Foop pirant: Unknown. 

Disrrisution: Méxtco: Jalapa, Orizaba. 
MALA: Volcfén Santa Maria (Nov.). 

The species is easily recognized by its bronzy color 
and the conspicuous blackish markings beyond the cell. 


GUATE- 


431. Mescinia indecora Dyar 
Ficures 476, 968 


Mescinia indecora Dyar, Ins. Insc. Menstr., vol. 8, p. 35, 1920. 


Forewing grayish fuscous with a faint purplish tint; 
the median area between faint, dark, antemedian and 
subterminal lines weakly shaded with blackish fuscous; 
costal area narrowly sordid whitish; discal dots blackish. 
Hind wings semihyaline, the veins faintly outlined in 
ocherous fuscous, a similar narrow shade along termen; 
veins 7 and 8 anastomosed for little more than half the 
length of vein 7; discocellular vein curved. Alar 
expanse, 22—23 mm. 

Male genitalia with uncus broadly triangulate, 
slightly tapering and with apex broadly rounded; apical 
process of gnathos completely fused, large, roughly 
triangular and with angles more or less rounded; anellus 
with lateral arms broad and narrowly divided; vin- 
culum stout and about as long as broad. Female 
genitalia of type damaged, so that little remains of 
bursa except the signum, which is a rather large, 
flattened, platelike thorn. 

Typr Locatity: Zacualpin, México (type in USNM). 

Foop piant: Unknown. 

Disrrisution: México: Zacualpdén (July), Techua- 
cin (July). 

The species fits badly in Mescinia on genitalic char- 
acters. Uncus, anellus, gnathos, the short vinculum, 
and the female signum are unlike any of these structures 
in the other species of the genus and suggest a separate 
generic placement, which may be necessary when more 
material can be studied and something is known of the 
earlier stages and biology. At present indecora is 
represented by only two specimens in the National 
Museum. 


Genus 122: Nonia 


[Venational division B. Forewing with 9 veins: 10 from the 
cell, 8 and 9 united, 4 and 5 stalked, 2 and 3 united. Male 
genitalia with transtilla incomplete.] 


122. Genus Nonia Ragonot 


Nonia Ragonot, Monograph, pt. 2, p. xiii, 1901.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 260, 1901. (Type of genus: 
Homoeosoma exiguella Ragonot.) 

Hypermescinia Dyar, Proc. Ent. Soc. Washington, vol. 47, p. 
341, 1914. (Type of genus: Hypermescinia lambella Dyar. 
New synonymy.) 

Tongue well developed. Antenna pubescent; shaft 
simple in both sexes. Labial palpus obliquely up- 
turned. Maxillary palpus filiform. Forewing smooth; 
9 veins; veins 2 and 8 united from very near angle of 
cell; 4 and 5 stalked; 8 and 9 united; 10 from the cell, 
separate from 8-9; male with fringe of hair or scales 
from underside of costa towards base, but without 
costal fold. Hind wing with vein 2 from well before 
lower angle of cell; 3 and 5 from angle and closely ap- 
proximate at base; 6 from upper angle of cell; 7 and 8 
completely fused beyond cell; discocellular vein curved; 
cell one-third of wing length. Abdomen of male with 


216 


a strong pair of ventrolateral tufts from eighth segment. 

Male genitalia of the Mescinia type but with all the 
parts more slender. Female genitalia with signa a 
cluster of several very small disks; ductus seminalis 
from bursa near its juncture with ductus bursae. 

The male genitalia show the close affinity of Nonia to 
Mescinia and show that Dyar’s interpretation of the 
forewing venation rather than that of Ragonot and 
Hampson is the correct one; that is, 2 and 3 united and 
4 and 5 stalked rather than 4 and 5 united and 3 stalked 
with 5. Ragonot and Hampson are also in error in 
regard to the discocellular vein of hind wing. I have 
examined the type of exiguella, and it has the disco- 
cellular vein decidedly curved, as have all the Central 
and South American and Puerto Rican specimens in 
the National Museum and Cornell University Collec- 
tions. The genus contains only one described species. 


432. Nonia exiguella (Ragonot) 
Figures 89, 477, 969 


Homocosoma exiguella Ragonot, Nouv. Gen., p. 35, 1888. 

Nonia exiguella (Ragonot), Monograph, pt. 2, p. 260, 1901. 

Hypermescinia lambella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 
341, 1914 (new synonymy). 

Forewing whitish gray overshaded (especially in fresh 
specimens) with vinaceous fawn, whitish along costa; 
antemedial line from costa beyond one-third, broken, 
represented (except in the darkest, most strongly 
marked specimens) by two or three blackish spots; a 
dark fuscous shade on inner margin at base; subterminal 
line, narrow, blackish, slanting, from costa to outer 
fourth of inner margin; discal spots conspicuous and 
confluent, blackish. Hind wings whitish in male, 
somewhat fuscous in female, veins and termen slightly 
darker. Alar expanse, 10-12 mm. 

Male genitalia figured from types of exiguella and 
lambella, which are identical in all details. 

TYPE LocaALities: Colombia (eriguella, in Mus. Univ. 
Berlin); Tabernilla, Panam4 (lambella, m USNM). 

Foop pirant: Unknown. 

Distripution: Pusrro Rico: Dorado (Dorado, 
May), La Sardinera (June); San German (Mayagiiez, 
Apr.); Isabela and Puerto Real (Vieques Isl., Apr.). 
Virein Istanps: Kingshill (St. Croix, Nov., Dec.). 
Jamaica. GuatEMALA: Cayuga (Feb., May). Pan- 
AMA: La Chorrera (May), Porto Bello (Feb.), Taber- 
nilla, Rio Trinidad (June). Cotompi1a. BRaziu: 
Vicosa (Minas Gerais, Oct.). Paraguay: Villarrica 
(July). 


Genus 123: Phestinia 


[Venational division B. Forewing with 10 veins; 10 from the 
cell, 8 and 9 long stalked, 2 and 3 united.] 


123. Genus Phestinia Hampson 


Phestinia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 57, 
1930. (Type of genus: Phestinia costella Hampson.) 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Tongue well developed. Antenna of female simple, 
a few projecting scales at each of the joints. Labial 
palpus upturned, slender, somewhat rough scaled; 
second segment reaching to vertex; third segment nearly 
as long as second, bluntly pointed. Maxillary palpus 
small but rather broadly and flatly scaled (squamous) 
and appressed to face. Forewing smooth; 10 veins; 
veins 2 and 3 united, from just before lower outer angle 
of cell; 4 and 5 from the angle, stalked for nearly half 
their lengths, the stalk separated at base from 2-3; 6 
from below upper angle of cell, straight; 8 and 9 long 
stalked; 10 from the cell, separate at base from stalk of 
8-9. Hind wing with vein 2 from well before lower 
outer angle of cell; 3 and 5 from the angle, connate (or 
very closely approximate) at base; 7 and 8 anastomosed 
for most of their lengths beyond cell; cell about one- 
third the length of wing; discocellular vein curved. 

Female genitalia with bursa copulatrix smooth except 
for signum; signum a small disk with short projecting 
spine (or spines) from lower margin; ductus bursae con- 
siderably longer than bursa, simple except for a mod- 
erately broad sclerotization of the tube at genital open- 
ing; ductus seminalis from bursa near its junction with 
ductus bursae. 

The genus was erected for a single species described 
from a single female specimen (not a male as stated by 
Hampson); so the foregoing diagnosis is incomplete. 
However, the venation and female genitalia are sufii- 
cient to show its distinctness from and close relationship 
to Nonia. It differs from the latter chiefly in having 
veins 9 of forewing and 8 of hind wing both present. 


433. Phestinia costella Hampson 
Fieures 92, $70 


Phestinia costella Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, 
p. 57, 1930. 

In general habitus similar to Nonia exiguella, but 
with darker areas more suffused and dark antemedial 
and discal markings less strongly contrasted. Hamp- 
son’s description of the dark suffusion as “red-brown” 
is somewhat exaggerated. There is a faint vinaceous 
tint to the brown, but it is hardly describable as red. 
Clarke has examined the genitalia of the female type 
and furnished me with a sketch of the signum (fig. 970b) 
and venation (fig. 92). I have seen no examples from 
the type locality, but have before me a rather worn 
female from the Cornell Collection, taken at San Ger- 
man, Puerto Rico, Apr. 17, 1930, which is certainly 
congeneric, and probably conspecific, with costella. It 
differs in that the signum (fig. 970a) has only one spine 
from the lower margin of signum. There are two spines 
in the type of costella. It is also smaller. Otherwise 
the Puerto Rican example agrees with the type. The 
venational details of the two specimens are identical. 
Alar expanse, 16-20 mm. 

Tyrr Locatity: Constant Springs, Jamaica (type in 
BM). 

Foop prant: Unknown. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 217 


Genera 124 and 125: Comotia and Bema 


[Venational division E. Forewing with veins 9 and 4 absent, 
rarely (in some specimens of Bema) with a vestigial indication of 
9 from 8 near apex. Hind wing with discocellular vein straight, 
vertical or slanting. Male genitalia with hooked apical process 
of gnathos small, bifid or partially fused; transtilla incomplete; 
uncus slender.] 


124. Genus Comotia Dyar 


Comotia Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 343, 1914. 
(Type of genus: Comotia torsicornis Dyar.) 

Antenna of male (fig. 478d) pubescent; basal segment 
enlarged and elongate, flattened on inner side (not tri- 
angular or armed with a spine as stated by Dyar); shaft 
flattened, the first half-dozen segments enlarged and 
excavated into a pocket (sinus) enclosing modified 
scales and followed by a ridge of crests on the following 
three or four segments; female antenna simple. Tongue 
well developed. Labial palpus upturned. Maxillary 
palpus filiform. Forewing narrowly elongate, smooth; 
nine veins; vein 2 from before angle; 3 and 5 shortly 
stalked; 10 from cell, close to 8; male without costal 
fold. Hind wing with vein 2 from well before lower 
angle of cell; 3 and 5 from the angle and approximate 
at base; 7 and 8 completely fused beyond cell (rarely 
a vestige of 8 discernible as a short spur near apex); 
discocellular vein straight and vertical; cell very short, 
less than one-fourth of wing length. Abdomen of male 
with a pair of weak ventrolateral tufts from eighth 
segment. 

Male genitalia of the Mescinia type, but with vincu- 
lum shorter (but slightly longer than broad), apical 
process of gnathos bifid, aedeagus constricted towards 
apex. Female with signum a single projecting disk, 
ductus bursae unsclerotized at genital opening, ductus 
seminalis from bursa near its junction with ductus 
bursae. 

434, Comotia torsicornis Dyar 
Ficures 117, 478 


Comotia torsicornis Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 343, 
1914, 


Forewing sordid white, lightly dusted with fuscous; 
costal area not appreciably paler; discal dots at end of 
cell separate, dark fuscous; some dark fuscous dots 
along termen. Hind wing translucent, smoky white 
with veins and termen faintly darkened. Alar expanse, 
13 mm. 

Male genitalia with some fine scobinations on penis. 

TypE Loca.ity: Porto Bello, Panam& (Mar., type in 
USNM). 

Foop prant: Unknown. 

Known only from the male type which is somewhat 
rubbed. Fresh specimens would probably exhibit pat- 
tern markings of a more distinctive type and contrasting 
color. The female from Taboga Isl. which Dyar asso- 
ciated with his type does not belong in Comotia. It is 
& specimen of Bema myja. 


300329—56—_15 


435. Comotia conyergens (Dyar), new combination 
Bema convergens Dyar, Ins. Insc. Menstr., vol. 7, p. 59, 1919. 


A trifle larger than torsicornis and with slightly longer 
labial palpi (they reach to vertex in torsicornis and 
slightly beyond in convergens), but otherwise similar. 
The species is known only from the male type and one 
other male from the type locality (not a female, as 
stated by Dyar). These specimens are somewhat 
rubbed, so no distinct wing pattern is discernible. The 
exact status of convergens will have to wait upon more 
and better material of both it and torsicornis. I doubt 
that it is specifically distinct. Dyar’s placement of it 
in Bema was obviously a blunder. Alar expanse, 15 
mm. 


Type Locatity: Cayuga, Guatemala (May, type in 
USNM). 
Foop piant: Unknown. 


Comotia sp. 


Fiaure 971 


The only female of the genus that I have seen is a 
rubbed specimen in the Cornell Collection from Jajoma 
Alta, Puerto Rico (collected by W. A. Hoffman, June 18, 
1930). Enough of the scaling remains to indicate that 
fresh specimens of Comotia will exhibit a rather well- 
marked pattern with a sprinkling of rufous scales among 
the fuscous scaling of the darker areas. The venation 
of fore and hind wings is identical with that of the male. 
I suspect that this specimen is a female of torsicornis, 
but exact specific identification must wait upon receipt 
of more material from Puerto Rico and Central America. 
Alar expanse, 15 mm. 


125. Genus Bema Dyar 


Bema Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 356, 1914. 
of genus: Bema myja Dyar.) 

Relmis Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 336, 1914. (Type 
of genus: Relmis ydda Dyar. New synonymy.) 

Antenna of male pubescent, with basal segment en- 
larged, triangular, and with shaft projecting from one 
corner (fig. 479e); shaft slightly swollen towards base, 
otherwise simple. Antenna of female simple. Male 
head (fig. 479d) hollowed between antennae and with a 
strongly sclerotized, hollowed tubercle projecting up- 
ward from front and bearing a ring of modified scales; 
these structures absent from female. Labial palpus 
upturned. Maxillary palpus filiform. Forewing nar- 
rowly elongate, smooth; normally with nine veins (in 
some specimens, see figs. 119 and 120, a vestige of vein 
9 present); vein 10 from the cell closely approximate to 
8; 2 from before lower angle of cell; 3 and 5 separate; 
male with costal fold. Hind wing with 2 from well be- 
fore lower angle of cell; 3 and 5 from the angle and ap- 
proximate at base; 7 and 8 completely fused beyond cell 
(rarely with a vestige of vein 8 near terminal end of 7); 
discocellular vein straight and slanting inward from 
lower angle; cell short, less than one-third the wing 


(Type 


218 


length. In male a pair of hair tufts on underside of 
prothorax and lying within grooved forecoxae. Ab- 
domen of male with two pairs of dorsolateral hair tufts 
from eighth segment; ventral membranous area of eighth 
segment covered with enlarged scales, these and the hair 
tufts set in elongate sclerotized tubercles (fig. 479c). 

Male genitalia similar to those of Nona except harpe 
broadly oval, sclerotized costa strongly arched and ter- 
minating in a short, free spur at apex; apical process of 
gnathos partially fused; vinculum long and narrow, with 
lateral margins subparallel (very slightly concave), and 
anterior margin indented. 

Female genitalia with signum a single projecting 
digitate disk; ductus bursae unsclerotized at genital 
opening; ductus seminalis from anterior end of bursa. 

The genus is easily identified by its characteristic gen- 
italia. It is difficult to see how Dyar justified his 
separation of species into two genera (Bema and Relmis) 
since he had a female of myja and females of ydda and 
jfifaca before him and not a single structural character 
to separate them except a trifling difference in the length 
of the labial palpi, at best specific, and here no greater 
between females of myja and ydda than between the 
females and males of myja. 


436. Bema neuricella (Zeller), new combination 
Figures 118, 119, 120, 479, 972 
Ephestia neuricella Zeller, Isis von Oken, p. 862, 1848.—Ragonot, 
Monograph, pt. 2, p. 288, 1901. 
Bema myja Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 336, 1914 
(new synonymy). 

Forewing grayish fuscous, basal area paler; ante- 
medial white line straight and slanting outwardly from 
costa to inner margin and with a blackish shading along 
its outer margin; subterminal line obscure, whitish, in- 
dented below costa (at vein 8) and, very slightly, at 
submedian fold; discal dots obscure, blackish; veins 
more or less outlined by dark scaling. In older speci- 
mens the dark markings are a pale brownish fuscous. 
Hind wings semitransparent, smoky white with veins 
darker and a dark shade at apex and along termen. 
Alar expanse, 15-19 mm. 

Specific differences in male genitalia are slight in the 
genus and comparative only, mostly in the size of the 
aedeagi, the harpes, and the width of the vinculum. 
Female genitalia with signum very small and of nearly 
uniform size but variable in shape and number of digi- 
tate projections. Variations in signa shown in figures 
972a-d. Equal variability exhibited in a series from 
one locality. 

Type tocauities: “St. Thomas,’”’ [Virgin Islands] 
(neuricella, 9, in Mus. Univ. Berlin); Taboga Isl., 
Panamé (myja, o', in USNM). 

Foop PLant: Inga sp. 

Distrizution: Cusa: Habana, Santiago (8 moths 
reared under E. E. A. de Cuba No. 9600, from larvae 
boring in seeds of Inga sp.; A. Otero, collector; only 
host record available). Banamas: Cay Santo Domingo 
(Sept.). Purrto Rico: Lares (July, Dec.). Viren 
Isuanps: St. Thomas, Kingshill (St. Croix, Oct., Nov., 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Dec.). Guaremata: Cayuga (Apr., May., June). 
PanamA: Taboga Isl. (Feb.). Trinipap. FRENCH 
Gurana: St. Laurent du Maroni. 

This species is represented by 65 specimens (11 & 
and 54 9) in the National and Cornell University Col- 
lections. All the males are more or less rubbed and 
show very little of the original color or pattern. The 
females are in better condition, especially those from 
Guatemala and St. Croix, and once the sexes are prop- 
erly associated it is not difficult to identify the species. 


437. Bema fritilla Dyar 
Bema fritilla Dyar, Ins. Insc. Menstr., vol. 7, p. 59, 1919. 


The only representative of this species is the male 
type. It is badly rubbed and shows no trace of the 
original pattern. I suspect that it is nothing but a 
runted specimen of neuricella (=myja). There is 
nothing to distinguish it from the type of myja except 
its smaller size (11 mm.) and proportionally smaller 
genitalia, which are otherwise identical. The difference 
in anal tufts which Dyar thought he saw was purely 
imaginary. The white scaling he mentions is that on 
the membranous area of the eighth segment and is 
identical on the two types. I am retaining the name 
for the present, pending receipt of similar small males 
from the type locality. The female from Cayuga 
(May) which Dyar associated with his type is not con- 
specific or even congeneric. Itis a Unadilla and prob- 
ably a small specimen of maturella Zeller, if what we 
have under that name is correctly identified. 

TypE Locatiry: Cayuga, Guatemala (June, type in 
USNM). 

Foop prant: Unknown. 


438. Bema ydda (Dyar), new combination 
Figure 974 


Relmis ydda Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 337, 1914. 


Forewing dark gray (in fresh specimens with a faint 
violaceous tint); base (and thorax) darker; antemedial 
pale line faint, bordered outwardly by a broad dark 
band; subterminal line pale, indented as in myja, 
bordered inwardly by a broad dark shade reaching to 
cell; outer area beyond subterminal line of a similar 
dark shade; discal dots obscured, more or less fused. 
Hind wing smoky fuscous. Abdominal tufts of eighth 
segment appreciably stouter than those of other species 
of Bema. Alar expanse, 19-20 mm. 

Male genitalia with vinculum broader in proportion 
to its length than that of myja or the other species of 
the genus. The entire genitalia are larger than those 
of myja, otherwise similar. Female genitalia as in myja 
except for enlarged signum. 

TypE Locatity: Rio Trinidad, Panamsé (May, June; 
type in USNM). 

Foop pirant: Unknown. 

Represented in the National Collection only by the 
female type and one other female from the type locality 
and a male from St. Jean Maroni, French Guiana. 
The outer border of the antemedial and the inner border 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 219 


of the subterminal lines have the appearance, to the 
naked eye, of broad, dark, weakly contrasted, trans- 
verse fasciae. 

439. Bema yddiopsis (Dyar) 
Relmis yddiopsis Dyar, Ins. Insc. Menstr., vol. 7, p. 58, 1919. 


Similar to ydda except ground color paler. Thorax 
and base of forewing pale clay color; terminal area of 
forewing also paler than the dark shade inwardly border- 
ing the subterminal line. Alar expanse, 20-21 mm. 

Male genitalia similar to those of myja except some- 
what larger; vinculum as narrow in proportion as that 
of myja. 

Type Locauity: Ténamo, Cuba (Mar., type in 
USNM). 

Foop piant: Unknown. 

Known only from the type and one other male from 
the type locality. The type (in fair condition) shows 
a few reddish scales in the dark area bordering the sub- 
terminal line, but I am unable to find any trace of the 
“red streak along submedian fold” mentioned by Dyar. 


440. Bema fifaca (Dyar), new combination 
Figure 973 


Relmis fifaca Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 337, 1914. 


Known only from the unique female type. This 
specimen similar in color and markings to fresh speci- 
mens of myja, of which it may be only a variety. The 
labial palpi seem a, trifle longer than those of myja and 
the signum is somewhat larger but not different enough 
to rule it out of that species. The outward dentations 
of the antemedial whitish line, mentioned by Dyar, 
also appear in some specimens of myja, depending upon 
how much the specimen is rubbed or how much the 
dark outer shading encroaches upon the white line. 
They are not constant, and the line properly should be 
described as slanting and straight. Alar expanse, 
18 mm. 

Type tocauity: Porto Bello, Panama (Dec., type 
in USNM). 

Foop piant: Unknown. 


Genera 126-128: Homoeosoma to Rotruda 


[Venational division C. Forewing with 10 veins: 10 from cell, 
8 and 9 united, 4 and 5 stalked or connate, 2 and 3 from cell. 
Hind wing with veins 7 and 8 strongly anastomosed; 3 and 5 
from cell; discocellular straight, vertical. Male genitalia with 
uncus broad, subtriangular; apical process of gnathos fused; 
vinculum stout, broad. Female genitalia with or without 
Signum; bursa, except for signum and some weak scobinations, 
not sclerotized; ductus seminalis from bursa or ductus bursae.] 


126. Genus Homoeosoma Curtis 


Homoeosoma Curtis, Ent. Mag., vol. 1, p. 190, 1883.—Hulst, 
Phycitidae of N. Amer., p. 191, 1890.—Ragonot, Mono- 
graph, pt. 2, p. xiii, 1901—Hampson in Ragonot, Mono- 
graph, pt. 2, p. 227, 1901.—Janse, Journ. Ent. Soc. South 
Africa, vol. 8, p. 26, 1945. (Type of genus: Phycis gemina 
Haworth, synonym of Tinea sinuella Fabricius; figs. 112, 
480, 978.) 


Phycidea Zeller, Isis von Oken, p. 178, 1839. 
Tinea sinuella Fabricius.) 

Tongue well developed. Antenna of male pubescent 
and with a slight notch at base of shaft; of female, 
simple. Ocelli present but small and lenses flattened 
in some specimens. Frons rounded, labial palpus up- 
curved; somewhat rough scaled; reaching to or slightly 
above vertex; third segment slightly shorter than 
second. Maxillary palpus filiform. Forewing smooth; 
narrowly elongate; termen slanting; 10 veins: veins 2 
and 3 from near lower outer angle of cell, separate; 4 and 
5 stalked for at least half the length of vein 5; 6 from 
below upper angle of cell, straight; 9 absent; 10 from 
cell; male with a slight costal fold at base of wing. 
Hind wing with seven veins; vein 2 from before lower 
angle of cell; 3 and 5 from angle of cell and closely 
approximate at base; 6 from upper angle of cell; 7 and 
8 fused beyond cell almost to costa; discocellular vein 
straight, nearly vertical and obsolescent; cell one-third 
to two-fifths of wing length. Eighth abdominal seg- 
ment of male simple (fig. 482e) with a single stout, 
central thorn associated with sternite (figs. 483f-g), or 
with a pair of weak ventrolateral hair tufts (fig. 489e). 

Male genitalia with uncus broad, subtriangular, 
outer surface densely covered with bristlelike scales. 
Gnathos terminating in a broadly triangulate or tear- 
shaped fused apical process with a very small spine at 
caudal end. Harpe simple; costa strongly sclerotized 
for most or all of its length; apex broadly rounded. 
Anellus U-shaped with narrow basal plate and long 
slender lateral arms, or semitubular. Aedeagus more 
or less sinuate, smooth or with a few minute scobina- 
tions at apex. Penis with a patch or elongate roll of 
sclerotized wrinklings, otherwise unarmed. Vinculum 
stout and broad, short to long and with terminal 
margin broadly rounded. 

Female genitalia with bursa copulatrix membranous, 
finely scobinate over most of its inner surface; signum 
a coarsely spined plate or altogether absent. Ductus 
bursae membranous throughout, finely scobinate only 
at or near junction with bursa copulatrix. Genital 
opening simple (unsclerotized and without adjacent 
sclerotized plates or other armature). Ductus semi- 
nalis from bursa near its junction with ductus bursae. 
Ovipositor lobes membranous or sclerotized. 

The genus, as here defined, is composed of two quite 
dissimilar groups which eventually will require generic 
separation. However, until the exact status of Pata- 
goma is established they may remain together in 
Homoeosoma, for I believe that the name Patagonia can 
be used eventually to include the whole of one species 
group. From Homoeosoma I am removing mucidel- 
lum Ragonot, reliqueella Dyar, olivaceella Ragonot, 
musiosum Dyar, cubellum Dyar, affusellum Ragonot, and 
the European nimbellum to a new genus (Rotruda), since 
they differ strikingly on both male and female genitalic 
characters from other species generally referred to 
Homoeosoma. Further generic division of Homoeosoma 
may be necessary when the genitalia of Old World 


(Type of genus: 


220 


species can be thoroughly studied. Our American 
species divide as follows: 


Male with apical process of gnathos broadly triangulate 
(figs. 481, 975), anellus U-shaped; female with signum, 
ovipositor lobes unsclerotized. 

Male with apical process of gnathos considerably smaller 
and tear-shaped, anellus semitubular; female without 
signum, ovipositor lobes sclerotized. 


The species of the first group are further divisible into 
subgroups on the following male characters: 


Highth abdominal segment simple. 

Eighth abdominal segment with a strong thornlike process 
associated with sternite. 

Highth abdominal segment with a pair of ventrolateral scale 
tufts. 


Genus Homoeosoma, Species 441-447: H. electel- 
lum to H. illuviellum emendator 


[Male with apical process of gnathos broadly triangulate, anellus 
U-shaped, eighth abdominal segment simple; female with signum, 
ovipostor lobes unsclerotized.] 


44). Homoeosoma electellum (Hulst) 
Figures 481, 975 


Anerastia electella Hulst, Ent. Amer., vol. 3, p. 137, 1887. 

Ephestia opalescella Hulst, Ent. Amer., vol. 3, p. 138, 1887 (new 
synonymy). 

Homoeosoma texanella Ragonot, N. Amer. Phycitidae, p. 15, 1887. 

Homoeosoma tenuipunctella Ragonot, N. Amer. Phycitidae, p. 15, 
1887. 

Homoeosoma olectella (Hulst), Phycitidae of N. Amer., p. 198, 
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 243, 
1901. 

Homoeosoma epalescella (Hulst), Phycitidae of N. Amer., p. 192, 
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 243, 
1901, 

Homoeosoma electellum (Hulst), U. 8. Nat. Mus. Bull. 52, p. 434, 
1903.—Kearfott, Canadian Ent., vol. 37, p. 123, 1905 (de- 
scription of larva).—Barnes and McDunnough, Contribu- 
tions, vol. 8, p. 200, 1916.—Forbes, Cornell Mem. 68, p. 634, 
1920.—McDunnough, Check list, No. 6374, 1939.—Satter- 
thwait and Swain, Journ. Econ. Ent., vol. 39, p. 575, 1946. 

Homoeosoma opalescellum (Hulst), U. S. Nat. Mus. Bull. 52, 
p. 434, 1903—McDunnough, Check list, No. 6375, 1939. 

Homoeosoma differtella Barnes and McDunnough, Contributions, 
vol. 2, p. 184, pl. 4, fig. 9, 1913. 


This and the three species following have similar 
genitalia and cannot be separated by any consistent 
characters in these organs. In all, the vinculum is pro- 
duced dorsolaterally into a pair of free arms (fig. 481a), 
the lateral margins of the apical process of gnathos are 
deeply concave (fig. 481c), and the signum is situated 
near the cephalic end of the bursa. There is consider- 
able variation in the gnathi and signa, but it is more 
individual than specific in character. H. electellum is 
readily separable from the other three species on its 
forewing pattern. The contrasting white subcostal 
streak (when distinguishable) is diagnostic. It is more 
or less obsolescent in the paler specimens. Wing color 
varies from pale powdery gray to whitish ocherous. 
The name opalescellum applies to the more yellowish 
specimens but represents nothing more than a color 
form. H. electellum is close to the Kuropean nebulellum 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


which has similar genitalia and larval habits, nebulellum 
being an important enemy of sunflower in Russia (see 
V. Schzegoleff, Journ. Oil Industry, Moscow, p. 44, 
November 1928). However, there appear to be enough 
differences in habitus to distinguish the two as distinct 
species. Alar expanse, 15-25 mm. 

TyeE Locatities: Blanco County, Tex. (electellum, 
in AMNH, ex Rutgers); Texas (texanellum, in Paris 
Mus.); California (opalescellum, in USNM; tenuipunc- 
tellum, in Paris Mus.) ; Everglades, Fla. (differtellum, in 
USNM). 

Foop piants: Flower heads of various asteraceous 
plants (Anthemis, Aster, Bidens, Brauneria, Chry- 
santhemum, Coreopsis, Dahlia, Helianthus, Heliopsis, 
Rudbeckia, Tagetes, Viguiera, Ximenesia); also from 
flowers of Opuntia and cotton and several U. S. Dep. 
Agr. rearings (at Mission, Hidalgo, and Brownsville, 
Tex.) from fruit of orange (on the trees). Sunflower 
seems to be the most favored food plant. 

Distrisution: Unitep States: Florida, Miami 
(Apr.), Orlando (June); Lowisiana, Baton Rouge 
(July), Vernon Parish (July); Texas, Blanco County, 
Brownsville (Mar., Apr., Aug., Sept.), College Station 
(June), Hidalgo, Liberty (July), Mercedes (Feb.), 
Mission (Jan.), New Braunfels (May), Presidio (Sept.), 
Sabinal (Mar.), San Benito (July, Aug., Sept.), San 
Diego (May), Smith Point (Sept.), Victoria (Mar., 
May, Sept.), Zavilla (Apr.); Mississippi, Natchez 
(May); Missouri, New Madrid (Aug.), Scott County 
(Oct.); Jowa, Ames (Aug.), Sioux City (June); Kansas, 
Onaga; Nebraska, Fort Calhoun; South Dakota, Elk 
Point (Aug.); Colorado, Boulder (Sept.), Denver 
(June), Pueblo (Sept.); New Mexico, Fort Wingate 
(May, June), Jemez Springs (Apr.), Las Vegas, Pecos 
(June); Arizona, South Liberty (Sept.), Santa Rita 
Mts. (Aug.); California, Sacramento (Aug.); Wash- 
ington, Bonneville (July), Pullman (June, July, Aug., 
Sept.), Walla Walla (June, July), Wenatchee (Aug.). 
México: Cuernavaca (July), Orizaba (May), Tehuacin 
(May, June). GuatTemata: Guatemala City. CuBsa: 
Havana (Sept.). Bermupa (Jan., Mar., Apr., May, 
July). 

The foregoing localities are for specimens before me. 
The species is generally distributed throughout the 
United States. 


442. Homoeosoma stypticellum Grote 
Figure 976 


Homoeosoma stypticella Grote, Bull. U. S. Geol. Geogr. Surv. 
Terr. vol. 4, p. 703, 1878.—Hulst, Phycitidae of N. Amer., 
p- 193, 1890.—Hampson, tn Ragonot, Monograph, pt. 2, 
p. 252, 1901. 

Homoeosoma uncanale Hulst, Trans. Amer. Ent. Soc., vol. 3, 
p. 162, 1886; U. S. Nat. Mus. Bull. 52, p. 434, 1903.— 
Barnes and McDunnough, Contributions, vol. 4, p. 175, 


1918.—McDunnough, Check list, No. 6324, 1939. (New 
synonymy.) 

Homoeosoma uncanalis Hulst, Phycitidae of N. Amer., p. 192, 
1890. 


Homoeosoma stypicellum Hulst, U. 8S. Nat. Mus. Bull. 52, p. 434, 
1903 (misspelling). 

Homoeosoma stypticellum (Grote), Forbes, Cornell Mem. 68, 
p. 634, 1920.—McDunnough, Check list, No. 6371, 1939. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 221 


Distinguished from other species with the electellum 
type of genitalia by the broad, transverse, antemedial 
dark brown band and the more or less extended dark 
shading on inner margin of subterminal line of forewing, 
this latter dark shading sometimes extending to the 
dark discal spots at outer margin of cell. Hind wings 
smoky fuscous. Alar expanse, 18-25 mm. 

I agree with McDunnough that the female from 
Colorado in the Hulst Collection which McDunnough 
has so labeled is the probable type of uncanale. The 
differences shown in figure 976 between it and the 
signum of typical eastern siypticellum are only individual 
in character and no more than those exhibited by other 
eastern examples of stypticellum. I find nothing on 
which to distinguish uncanale, even as a western race, 
from stypticellum. 

Typrr tocauities: Maine (stypticellum, in BM [?]); 
Custer County, Colo. (uncanale, in AMNH, ex Rutgers). 

Foop piant: Cirsium, Rhus. 

Distrisution: Unrrep States: Maine, Orono, Bar 
Harbor (July), Wales (July); Massachusetts, Martha’s 
Vineyard (July, Aug.); Connecticut, East River (July, 
Aug.); Rhode Island, Weekapaug (July, Aug.); New 
York, Mlion (June); Pennsylvania, Oak Station (Mar.), 
New Brighton (July); Jilinois, Palos Park (Aug.); 
Arkansas, Washington County (July); Texas; Colorado, 
Custer County, Glenwood Springs (Aug.), Logan’s 
Peak (July). Canapa: Quebec, St. Hilaire (July); 
Ontario, Ottawa (June), Trenton (June); Manitoba, 
Aweme (June, July), Rounthwaite (Aug.); Saskatch- 
ewan, Regina (July). 

Among the foregoing, one example (from Palos Park, 
Til., Aug. 6, 1939) was reared by A. K. Wyatt from 
larvae feeding in the flower heads of ‘‘swamp thistle.” 


443. Homoeosoma striatellum Dyar 
Fiaurt 977 
Homoeosoma striatellum Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 38, 1905.—McDunnough, Check list, No. 6367, 1939. 

Pale slate gray; the veins of forewing outlined by 
blackish scaling, transverse lines nearly obsolete, in 
some specimens indicated. by a narrow, dark, angulate, 
antemedial band and some dark shading towards costa, 
on inner side of subterminal oblique line; discal spots 
faintly indicated or absent. Hind wings whitish to very 
pale smoky fuscous. Signum of female somewhat vari- 
able, the extent of variation shown in the figure. Alar 
expanse, 18-23 mm. 

Typr Locauity: Phoenix, Ariz. (type in USNM). 

Foop puant: Unknown. 

Distripution: Arizona, Baboquivari Mts., Ajo, 
Pima County (Mar.), Phoenix (Mar., Apr.), Tempe 
(Feb.); California, Death Valley (Apr.). 


444, Homoeosoma oslarellum Dyar 
Figure 979 


Homoeosoma oslarellum Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 38, 1905.—McDunnough, Check list, No. 6373, 1939. 


Dark grayish fuscous with a scattering of whitish 
scales along costa of forewing; transverse marking obso- 


lete. Hind wing pale smoky fuscous. Alar expanse, 
17-23 mm. 

Female genitalia with signum similar to that of 
striatellum, considerably larger than that of either 
stypticellum or electellum, situated somewhat nearer the 
middle of the bursa. 

Type tocauiry: Chimney Gulch, Golden, Colo. 
(type in USNM). 

Foop piant: Unknown. 

Distrisution: Colorado, Golden; New Mexico, Beulah 
(July); California, San Francisco (Apr.). 


445. Homoeosoma oslarellum breviplicitum, new race 


A Southern California race differing from typical 
oslarellum by the much shorter roll of sclerotized wrink- 
lings of penis (but half the length of that in oslarellum 
or the other preceding species), its paler wing color and 
forewing markings. Color of forewing slate gray (inter- 
mediate between that of oslarellum and striatellum); a 
faint but distinguishable and sharply angulate, narrow, 
dark, antemedial band; veins more or less outlined by 
blackish scaling, especially in outer area of wing; hind 
wing more whitish and with the veins more darkly con- 
trasted than in typical oslarellum. Alar expanse, 20-25 
mm. 

Type tocauity: San Diego, Calif. (type in USNM, 
61370). 

Foop prant: Unknown. 

Described from male type and male paratype from 
the type locality (May 3, 1924, H. G. Dyar); one female 
paratype from San Diego (June 14, 1924, J. M. Dam- 
mers) and three female paratypes from Riverside, 
Calif. (Apr. 14 and 17, 1937, Grace B. and John L. 
Sperry). 


446. Homoeosoma illuviellum Ragonot 


Figure 482 


Homoeosoma illuviella Ragonot, Nouv. Gen., p. 33, 1888; Mono- 
graph, pt. 2, p. 245, 1901—Hulst, Phycitidae of N. Amer., 
p. 192, 1890. 

Homoeosoma candidella Hulst, Ent. Amer., vol. 4, p. 118, 1888. 

Homoeosoma illuviellum Ragonot, Hulst, U. 8. Nat. Mus. Bull. 
52, p. 484, 1903.—McDunnough, Check list, No. 6366, 1929. 

White with a faint scattering of dark scales on fore- 
wing; transverse lines obsolete, their usual position 
indicated only by a couple of dark dots in the ante- 
medial and a very faint transverse shading of dark dust- 
ing in the subterminal areas; costal edge dark beyond 
middle; a small but rather conspicuous dark dot at 
lower outer angle of cell and a much smaller, more 
obscure dot at upper outer angle. Hind wings pale 
smoky fuscous; cilia white. Alar expanse, 22-28 mm. 

Male genitalia with apical process of gnathos stouter 
and with less deeply concave lateral margins than in 
the preceding species; vinculum not produced dorso- 
laterally. 

Typr tocaities: Sonora, México (iluviellum, in 
Paris Mus.); Arizona (candidellum, in AMNH, ex 
Rutgers). 

Foop puant: Unknown. 


222 


DistRipution: Unirep Sratres: Arizona (no further 
locality), Prescott (July); Colorado. M*#xtco: Sonora. 

The type and only specimen in Paris is from “Senora” 
[sic], México, Morrison, 1883, labeled in Ragonot’s 
handwriting “Homoeosoma illunella Rag. pl. xxxii, 
fig. 11, type original.” It is a female and not a male as 
stated in the original description. I have examined the 
genitalia. The type of candidellum has been examined 
by Ragonot as the label indicates. It also is a female 
with a glued-on abdomen. However, there is no doubt 
that it is conspecific with dlunellum. 


447. Homoeosoma illuviellum emendator, new race 
Fiaure 980 


Differs from typical dluviellum only in color and in- 
tensity of markings; head, thorax, and forewing a pale 
slate gray rather than white; dark spots as in Wluviellum 
but more conspicuous and the dark shading along outer 
line more pronounced; hind wings darker. Alar ex- 
panse; 25-29 mm. 

Genitalia as in dluviellum. 

The new name is given with considerable hesitation 
as this may prove to be only a colorform. However, as 
several distinct species in Homoeosoma cannot be sepa- 
rated by genitalia and the specimens before me are so 
different in color from true dluviellum, I suspect that 
they represent something more than a mere color form, 
possibly a food-plant race. 

Type Locauity: Chimney Gulch, Golden, Colo. 
(May; type in USNM, 61371). 

Foop piant: Unknown. 

Described from male type, one male paratype from 
Breckenridge, Colo. (June), one female paratype from 
Eureka, Utah (June 2, 1917, Tom Spalding), and two 
female paratypes from Richfield, Utah (May 20, 22, 
1930). 


Genus Homoeosoma, Species 448-453: H. imitator 
to H. deceptorium 


[Male with apical process of gnathos broadly triangulate, anellus 
U-shaped, eighth abdominal segment with a strong thornlike 
process associated with sternite; female with signum, ovipositor 
lobes unsclerotized.] 


448. Homoeosoma imitator, new species 
FIGURES 483, 984 


In color and markings similar to striatellum Dyar but 
strikingly different in structure. The presence of the 
thornlike process on eighth abdominal segment at once 
distinguishes it from all the species of the foregoing sub- 
group, and the produced lateral arms of vinculum from 
all the species of this group except longiventrellum. The 
eighth segment process varies in length in different 
specimens. Extremes of variation are shown in figures 
483f and 483g. 

Alar expanse, 19-24 mm. 

Type Locatiry: Palm Springs, Riverside County, 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Calif. (type in USNM, 61372; paratypes in Los An- 
geles County Mus., Calif.). 

Foop prant: Unknown. 

Described from male type and three male and three 
female paratypes from the type locality (Mar.), one 
male paratype from West Riverside, Calif. (Feb. 16, 
1906), and two female paratypes from Palo Verde, 
Imperial County, Calif. (Feb. 10, 1940). 


449. Homoeosoma longiventrellum Ragonot 
Figure 484 


Homoeosoma longiventrella Ragonot, Nouv. Gen., p. 34, 1888; 
Monograph, pt. 2, p. 253, 1901. 

Homoeosoma longiventrella noctividella Ragonot, Monograph, pt. 
2, p. 253, 1901. 

There are no specimens from tropical America agree- 
ing with Ragonot’s description or figure of longiventrella 
in the National Collection. The type is in the British 
Museum but lacks an abdomen. In Paris, Clarke lo- 
cated but one specimen in the Ragonot Collection, a 
male from Chiriqui, Panamé, labeled Gn Ragonot’s 
handwriting) “var. noctividella Rag.” If the genitalia 
(figured) of this specimen represent, as they presumably 
do, the true longiventrella the species should be easily 
recognized, for it is the only one of the tropical species 
with projecting arms from vinculum and a thornlike 
projection from the eighth abdominal segment. The 
projecting arms of vinculum are exceptionally long. 

TypE Locatities: Chile (?) (longiventrellum, in BM); 
locality not known to me (noctividellum, in Mus. Univ. 
Berlin). 

Foop prant: Unknown. 


450. Homoeosoma albescentellum Ragonot 
Fiaures 487, 983 


Homoeosoma albescentella Ragonot, N. Amer. Phycitidae, p. 15, 
1887.—Hulst, Phycitidae of N. Amer., p. 192, 1890.— 
Hampson, in Ragonot, Monograph, pt. 2, p. 250, 1901. 

Homoeosoma elongellum Dyar, Proc. Ent. Soc. Washington, vol. 
5, p. 227, 1903.—McDunnough, Check list, No. 6365, 1929. 
(New synonymy.) 

Homoeosoma albescentellum (Ragonot) Hulst, U. S. Nat. Mus. 
Bull. 52, p. 434, 1903.—McDunnough, Check list, No. 6368, 
1939. 

Rather pale brownish fuscous, dusted with white 
scales and with darker markings accented. The white 
scaling concentrated on the costal half of forewing, 
giving the lower half a contrasting brownish shade; 
transverse antemedial band blackish brown, normally 
angulate but outwardly rounded in occasional speci- 
mens; subterminal line inwardly angled at vein 6 and 
inwardly shaded with blackish brown, especially towards 
costa; discal spots at end of cell small, but distinct. 
Hind wing smoky fuscous. 

From others in the subgroup having a ventral, thorn- 
like process from the eighth abdominal segment of the 
male, this and the three species following are distin- 
guished by their simple vinculi (without produced lat- 
eral arms). Alar expanse, 21-30 mm. 

Male genitalia figured from type of elongellum. Fe- 
male genitalia figured from the type of albescentellum 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


in the Paris Museum. This latter is the only specimen 

of the species in the Ragonot Collection. There are no 

characters of structure or pattern by which elongellum 
can be separated from albescentellum. 

Type Locauitiss: California (albescentellum, in Paris 
Mus.); Williams, Ariz. (elongellum, in USNM). 

Foop pLant: Unknown. 

Distrisution: California; Arizona, Williams (July), 
Redington, Santa Catalina Mts.; Colorado, Silverton 
(July). 

451. Homoeosoma impressale Hulst 
Ficures 486, 981 

Homoeosoma impressalis Hulst, Trans. Amer. Ent. Soc., vol. 13, 
p- 163, 1886; Phycitidae of N. Amer., p. 191, 1890.— 
Hampson, in Ragonot, Monograph, pt. 2, p. 239, 1901. 

Homoeosoma uncanalis Ragonot (not Hulst), Monograph, pt. 2, 
p. 253, 1901. 

Homoeosoma impressale Hulst, U. S. Nat. Mus. Bull. 52, p. 434, 
1903.—Barnes and McDunnough, Contributions, vol. 3, p. 
200, 1916; vol. 4, p. 174, 1918 —McDunnough, Check list, 
No. 6363, 1939. 

McDunnough’s identification (Contributions, vol. 4, 
pt. 2, p. 174, 1918) of the true type of this species in the 
Rutgers Collection is without any doubt the correct 
one. The specimen is a male without abdomen, 
labeled Nevada, ‘3838.’ It also bears a Hulst name 
label ““Homoeosoma uncanale,’”’ which is obviously in- 
correct and can be ignored as an error in labeling. The 
Nevada specimen is one of the well-marked examples 
of the species and agrees with the original description. 

The markings are variable in the extreme, ranging 
from pure white examples with only a couple of small 
blackish discal dots at end of cell and a row of faint 
blackish dots along termen to white examples with a 
broad, slanting, blackish, antemedial band and a narrow 
blackish subterminal band. On the right side of one 
specimen before me from Manitoba the blackish bands 
are fused, filling the entire center of the wing. The 
specimens without transverse markings are superficially 
like those of tlluviellum, usually showing the discal spot 
at upper outer angle of cell a trifle stronger but other- 
wise only separable on genitalia. The two figures in 
the Ragonot Monograph (pl. 33, figs. 7, 18) are good 
illustrations of the normal variation in pattern. Alar 
expanse, 23-34 mm. 

TYPE LOCALITY: 
Rutgers). 

Foop prant: Unknown. 

Distrizution: Unirep States: Arizona; Nevada; 
Colorado; Utah, Stockton (June, July); California, 
Davis Creek, Modoc County (June); Washington, 
Pullman (Aug.), Walla Walla (June). Canapa: Mani- 
toba, Cartwright, Rounthwaite (Aug.). 


Nevada (type in AMNH, ex 


452. Homoeosoma inornatellum (Hulst) 
Figures 485, 982 
Euzophera inornatella Hulst, Canadian Ent., vol. 32, p. 173, 


1900.—Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 228, 
1904. 


223 


Homoeosoma inornatellum (Hulst), McDunnough, Check list, 
No. 6369, 1939, 

A very faintly marked species, especially in the male; 
forewing pale gray or grayish fuscous with a faint, white 
subcostal streak (much as in electellum); antemedial 
markings consisting of one or two obscure dark dots; 
discal dots at end of cell faint. In the female the 
markings are somewhat stronger. Hind wings pale 
smoky fuscous. Superficially the males look much like 
some examples of electellum, but the two species are 
easily separated on structure. In addition to the 
differences in male genitalia and the eighth abdominal 
segment, inornatellum has veins 2 and 3 of forewing 
approximate at origin while in electellum these veins are 
well separated, about as far apart as 3 and 4. Alar 
expanse 23-25 mm. 

From other species with simple vinculi (albescentellum, 
impressale, deceptorium), inornatellum is readily dis- 
tinguished by the shape of the harpe and the apical 
process of gnathos. The signum of the female is 
situated very near the anterior end of the bursa, as in 
electellum. Dyar’s identification of the type of inorna- 
tellum with stypticellum is an obvious error. The two 
species are not even superficially similar. 

Tyrr Locauity: Anglesea, N. J. (type in AMNH, ex 
Rutgers). 

Foop piant: Unknown. 

Distrisution: New Jersey, Anglesea (May, June); 
Pennsylvania, Mount Airy. 

A series of males from the type locality is in the 
National Collection. 


453. Homoeosoma deceptorium, new species 


Figures 488, 985 


Color and markings as in the larger, most strongly 
marked specimens of stypticellum, except that the white 
dusting on forewing is more restricted to the costal half 
of the wing; transverse antemedial band broad, slanting, 
not appreciably angulate, blackish brown; subterminal 
dark band narrower but distinct; discal spots at outer 
angles of cell distinct. Hind wings pale smoky fuscous 
with veins outlined by darker scaling. Alar expanse 
23-27 mm. 

Male genitalia chiefly distinguished from those of 
other associated species in this subgroup by the shape 
of the apical process of gnathos. Signum of female 
located near middle of bursa. 

TyprE Locauity: New Brighton, Pa. (type in USNM, 
61373; paratypes in Canadian Nat. Coll.). 

Foop piant: Unknown. 

Described from male type and four male and one 
female paratypes from the type locality (July, Aug.); 
one male and one female paratype from Alcove, Quebec 
(July 7, 1936, F. A. Urquart). 


224 


Genus Homoeosoma, Species 454: H. discrebile 


[Male with apical process of gnathos broadly triangulate, anellus 
U-shaped, eighth abdominal segment with a pair of ventrolateral 
scale tufts; female with signum, ovipositor lobes unsclerotized.] 


454, Homoeosoma discrebile, new species 
Figures 489, 986 


| White with a very faint cream tint and with three 


trongly contrasted brown markings on forewing, a 
transverse slanting antemedial band, a somewhat 
narrower but equally distinct subterminal band and a 
large discal spot formed by the fusion of the two dots 
at outer angles of cell. Hind wings white; veins not 
outlined by dark shading. Alar expanse, 15-17 mm. 

Male genitalia similar to those of electellum; vinculum 
with produced dorsolateral arms. Signum of female 
large, situated well back of the anterior end of bursa. 

Typr tocauity: “S. KH. Brazil” (F. D. Jones, ‘‘1920- 
303”; type in BM; paratypes in BM and USNM, 
61374). 

Foop PLiant: Unknown. 

Described from male type and three female paratypes 
from the type locality, and one female paratype from 
Castro, Parand, Brazil. 

An easily recognized species. 


Genus Homoeosoma, Species 455-461: H. pere- 
grinum to H. nimbosellum 
{Male with apical process of gnathos considerably smaller and 


tear-shaped, anellus semitubular; female without signum, ovi- 
postor lobes sclerotized.] 


455. Homoeosoma peregrinum, new species 


Fieurrs 490, 987 


Forewing very narrow in the male (8 by 1/4 mm.); in 
female somewhat broader. Color brown, evenly pep- 
pered with whitish scales, making the general color 
grayish fuscous; markings obscure, only the faintest 
indication of a narrow antemedial dark line and the 
discal spot at end of cell. Hind wing of a very pale 
smoky hue with darkened veins and a fuscous line along 
termen. Alar expanse, 15-17 mm. 

Male genitalia chiefly distinguished from those of 
other species in the group (except assitum) by the very 
small, semicircular sclerotized area of penis (assitum is 
distinguished by its smooth aedeagus and slenderer, 
more evenly tapering harpes); aedeagus in peregrinum 
finely scobinate at apex. The structural differences 
among the species of this group are slight and hardly of 
specific value. In the female the amount and extent 
of scobination of the bursa varies somewhat from species 
to species, but almost as much from specimen to speci- 
men. These scobinations (except in oconequensis) are 
more concentrated in a single circular area where they 
have the appearance of a weak signum, which they are 
not. 

TYPE Locaity: Carmel, Calif. (typein USNM, 61375; 
paratypes in USNM and Lange Coll.). 

Foop PLant: Anaphalis margaritosa. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Described from male type and three male and two 
female paratypes from the type locality, reared by 
W. H. Lange (Mar. 4, 1938, and Mar. 25, 1939); and 
one male paratype from Juan Vinas, Costa Rica (May). 

H. peregrinum is the only species of the group: known 
from the United States, and since it also occurs in Costa 
Rica, the natural assumption is that it is a wanderer 
from tropical America. It may be the species that 
Hampson misidentified with his Old World ephestidiella 
(in Ragonot, Monograph, pt. 2, p. 256, 1901). ‘The 
latter belongs to quite a different species group which is 
not found in the New World. 


456. Homoeosoma vepallidum, new species 
Figures 491, 991, 992 


Pale brownish fuscous dusted with white, giving the 
insect a sordid whitish tint; antemedial markings of 
forewing reduced to a dark spot extending from imner 
margin to vein 1b; discal dots at outer angles of cell 
present but ebscure; subterminal line obscure, slanting, 
narrow, white, and margined imwardly by a few very 
faint, dark spots; a faint but distinct white subcostal 
streak. Hind wing smoky white, veins faintly dark- 
lined. Alar expanse, 18-20 mm. 

Genitalia without outstanding specific characters. 

Typz Locauity: “Villa Ana, F. C. S. F.,”’ Argentina 
(type and paratype in BM). 

Foop prant: Unknown. 

Described from male type and female paratype from 
the type locality, Nov. 29, 1923, R. J. Hayward. A 
female from the Cornell University Collection, taken in 
the Sierra de Cérdoba, Argentina, Mar. 9, 1920, is not 
included among the types because it is paler in color 
and has a larger bursa, copulatrix (fig. 992) than the 
paratype from Villa Ana, but it is undoubtedly the 
same species. 


457. Homoeosoma ditaeniatellum Ragonot 
Ficurs 990 
Homoeosoma ditaeniatella Ragonot, Nouv. Gen., p. 33, 1888; 
Monograph, pt. 2, p. 236, 1901. 

According to Clarke’s notes this specimen is badly 
rubbed and stained and the figure and description drawn 
from it consequently somewhat misleading. The species 
is probably considerably more whitish than the type 
would seem to indicate. Its identification will have to 
wait upon more material from the type locality. Alar 
expanse, 18 mm. 

TYPE LOCALITY: Quillota, Chile (Paulson, 1887; type 
in Paris Mus.). 

Known only from the female type. 


458. Homoeosoma oconequensis (Dyar), new combination 
FigurE 989 


Eucampyla oconequensis Dyar, Ins. Insc. Menstr., vol. 7, p. 60, 


A pale fuscous gray species almost unmarked; the 
faintest indication of a narrow, transverse, dark ante- 
medial line and a dark discal dot at lower outer angle 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


of cell. Hind wing pale smoky white; veins faintly 
outlined. Alar expanse, 22 mm. 

Genitalia distinguished by the very sparse scobina- 
tion of bursa copulatrix without any trace of a concen- 
tration of these scobinations into a conspicuous spot. 

Tyrer Locauity: Oconeque, Carabaya, Peri (Schaus, 
collector; type in USNM). 

Foop prant: Unknown. 

Known only from the female type. 


459. Homoeosoma assitum, new species 
Fiaures 492, 988 


A small pale gray species, very faintly but appreciably 
marked; forewing with a narrow, slanting, dark ante- 
medial line; a very faint, straight, slanting, white, sub- 
terminal line; two small, obscure discal dots at outer 
angles of cell. Hind wing smoky white; veins darkly 
outlined; a narrow dark line along termen; cilia white. 
The species resembles peregrinum but is smaller and 
paler. Alar expanse, 13-17 mm. 

Male genitalia distinguished by their narrow, taper- 
ing harpes and smooth aedeagus. 

TYPE LocaLity: Cafiete, Pera (type in USNM, 61376; 
paratypes in USNM, BM, Paris Mus.). 

Foop pirant: Unknown. 

Described from male type and seven male and four 
female paratypes from the type locality, taken Mar. 2, 
1943, on the foliage of a composite by E. J. Hambleton. 


460. Homoeosoma acmaeopterum Ragonot 
Ficure 493 
Homoeosoma acmaeoptera Ragonot, Nov. Gen., p. 34, 1888; 
Monograph, pt. 2, p. 255, 1901. 

According to Clarke’s notes there are five specimens 
under this name in the Ragonot Collection at Paris 
representing what appears to be an assortment of spe- 
cies. The male type, so labeled by Ragonot and from 
which the figure in his Monograph (pl. 33, fig. 21) was 
made, is the only specimen that can definitely be called 
acmaeopterum. According to Clarke, the Ragonot 
figure is a fair representation except that the forewing 
is too short in proportion to its width and its median 
dark area too strongly contrasted. Alar expanse, 22 
mm. 

Type Locauity: Coquimbo, Chile (type in Paris 
Mus.). 
Foop pirant: Unknown. 


461. Homoeosoma nimbosellum Ragonot 
Fiaure 494 
Homoeosoma nimbosella Ragonot, Nouv. Gen., p. 34, 1888; 
Monograph, pt. 2, p. 255, 1901. 

The type is a male, and not a female, as stated by 
Ragonot. According to Clarke, the Ragonot figure 
(Monograph, pl. 30, fig. 22) is somewhat inaccurate, 
being too dark and with the subterminal line too well 
defined. Alar expanse, 18 mm. 

Type xocatity: Valparaiso, Chile (type in Paris 
Mus.). 

Foop prant: Unknown. 

Known only from the type in Paris. 

8003293—bé6——16 


225 


Genus Homoeosoma, Species 462 (unplaced): 
H. unionellum 


462. Homoeosoma unionellum Ragonot 


Homoeosoma unionella Ragonot, Nouv. Gen., p. 33, 1888.— 
Hampson, in Ragonot, Monograph, pt. 2, p. 235, 1901. 

A photograph of the type shows a large, narrow- 
winged, pale specimen (yellowish white, according to 
description) with costal edge of forewing beyond middle, 
blackish. A sketch of the female genitalia shows that 
the species belongs in the first group (Homoeosoma 
Species 441-454), but more exact placement is impos- 
sible without a male. Alar expanse, 28 mm. 

TyrE Locatity: Milpas, México (type in BM). 

Foop prant: Unknown. 

Known only from the female type. 


127. Genus Patagonia Ragonot 


Patagonia Ragonot, Monograph, pt. 2, p. xiii, 1901—Hampson, 
in Ragonot, Monograph, pt. 2, p. 236, 1901. (Type of 
genus: Homoeosoma magellanella Ragonot.) 

Characters of the second Homoeosoma species group, 
except veins 4 and 5 of forewing connate rather than 
stalked. 

So far represented only by the single female example 
of the type species. J suspect that this is an individual, 
venational aberrant and that when further examples 
are taken they will show the normal Homoeosoma vena- 
tion of forewing, 4 and 5 appreciably stalked. If, 
however, the connate condition of 4-5 were to prove 
constant and the genitalia of the male show no marked 
variation from those of the second Homoeosoma group, 
the venational difference would have a purely specific 
significance and Patagonia could be retained as a valid 
genus on the genitalic characters of the group; and the 
species of the group could be referred to it. 


463. Patagonia magellanella (Ragonot) 
Figure 993 
Homoeosoma magellanella Ragonot, Nouv. Gen., p. 35, 1888. 
Patagonia magellanella (Ragonot), Monograph, pt. 2, p. 226, 
1901. 

According to Clarke’s notes, the figure of the type in 
Ragonot’s Monograph (pl. 33, fig. 25) is faulty in that 
the forewing is too broad, the whitish costal streak not 
extensive or white enough, the subterminal striations 
too much accented, and the wing generally too dark. 
However, the pattern should make specimens from the 
type locality easily identifiable. Alar expanse, 19 mm. 

Tyrer Locatity: Punta Arenas, Straits of Magellan, 
Chile (type in Paris Mus.). 

Foop pirant: Unknown. 


128. Rotruda, new genus 


Tyrer oF GeNus: Homocosoma mucidellum Ragonot. 
Tongue, antenna, palpi, venation, and habitus as in 
Homoeosoma. Male genitalia with a comb of strong 
spines along side of aedeagus. Female genitalia with 


226 


two large signa composed of opposed, cup-shaped, 
strongly spined concave plates; ductus seminalis from 
ductus bursae. 

In addition to mucidella and its American allies the 
European Homoeosoma nimbella Zeller should be re- 
ferred here (fig. 496). 


464. Rotruda mucidella mucidella (Ragonot), new combination 
Figures 495, 994 


Homoeosoma mucidella Ragonot, N. Amer. Phycitidae, p. 15, 
1887; Monograph, pt. 2, p. 258, 1901.—Hulst, Phycitidae 
of N. Amer., p. 193, 1890. 

Homoeosoma mucidellum Ragonot, Hulst, U. S. Nat. Mus. Bull. 
52, p. 434, 1903.—Dyar, Pomona College Journ. Ent., vol. 2, 
p- 378, 1910.—Barnes and McDunnough, Contributions, 
vol. 2, p. 223, 1914.—McDunnough, Check list, No. 6370, 
1939. 

There is nothing except color differences to separate 
typical mucidella from the following supposed species 
which I am treating as races or synonyms, and these 
differences are not clear-cut between tropical examples 
and dark specimens from eastern United States. What 
color differences there are depend upon the amount of 
white dusting and the intensity of the dark markings. 
Typical mucidellum is the most heavily dusted with 
white and has the dark markings of forewing most 
sharply contrasted (frequently blackish), and to the 
naked eye seems a paler form than any of the others, 
although there are intergrading examples between it 
and religuella. Alar expanse, 15-22 mm. 

There is considerable variation in the number of 
spines in the comb along the side of aedeagus (10 to 15), 
but this variation is as great between examples of 
typical mucidella as it is between it and any of its races. 


Tyrer Locatity: California (type in Paris Mus.). 

Foop prants: Aster (and presumably other com- 
posites). 

Distrisution: Unirep Strarss: California, Clare- 
mont, Colfax (July), Deer Park Springs (Lake Tahoe), 
La Puerta Valley (July), Loma Linda (July), Los 
Angeles County (Aug.), Polente (Sept.), Riverside 
(Nov.), Sacramento (Sept.), San Bernardino Mts. 
(July), San Diego (July), Shasta Retreat (Siskiyou 
County, July, Aug., Sept.), Warner Mts. (Modoc 
County, July); New Mexico, Jemez Springs, Las Vegas, 
Hot Springs (June, Sept.); Arizona, Palmerlee, Tempe 
(June); Colorado, Denver (July), Glenwood Springs 
(Aug., Oct.); Utah, Provo (July), Salt Lake, Stockton 
(June, Aug.); Wyoming, Jackson Hole (July); Oregon, 
Crater Lake (July); Washington, Bellingham (June), 
Pullman (May, July, Aug., Oct.), Snake River (July), 
Walla Walla (June, July), Wenatchee (Aug.). Can- 
ADA: British Columbia, Kaslo (June). 

The range of distribution is probably the entire 
United States, southern Canada, and northern México 
from the Rocky Mountains westward to the Pacific 
Ocean. 

The European nimbella (fig. 496) is very close to 
mucidella, differing only in the greater number of spines 
on aedeagus (19 to 23). Were it not for this apparently 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


consistent difference I should be inclined to treat 
mucidella as an American race of the Old World species. 


465. Rotruda mucidella reliquella (Dyar), new combination 


Homoeosoma reliquellum Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p- 112, 1904.—Forbes, Cornell Mem. 68, p. 634, 1920.— 
MeDunnough, Check list, No. 6372, 1939. 

Darker than typical mucidella with forewing some- 
what suffused, the dark markings less sharply con- 
trasted and the whitish dusting confined mostly to the 
costal half of the wing. Apparently confined to the 
areas of Canada and the United States east of the 
Rocky Mountains. 

Alar expanse, 14-19 mm. 

TypE Locauity: Center Harbor, N. H. (type in 
USNM). 

Foop piants: Aster, Cirsium spinosisimum, Sitihas 
caroliniana, Sonchus asper, lettuce. 

Distrisution: Unirep Srares: New Hampshire, 
Center Harbor (July), Hampton (June, July); Massa- 
chusetts, Amherst (June), Cohasset, Framingham (May), 
Winchendon (June); Connecticut, East River (July, 
Aug.); New Jersey, Montclair (Aug.), Newark (Aug.); 
Pennsylvania, New Brighton (May, June, July), Oak 
Station (June), Pittsburgh (May, June, Aug.); Illinois, 
Edgebrook (Oct.), Chicago (July) ; District of Columbia, 
Washington (May); Maryland, Plummer’s Isl. (May); 
Virginia, Great Falls (July); North Carolina, Tryon 
(May); Georgia, Savannah (May); Florida, Big Pine 
Key (Apr.), Chokoloskee, Everglades (Apr.), Ft. 
Myers (Apr.), Lake Alfred (July), Miami (Apr.), Pom- 
pano (Mar.). Cawnapa: Ontario, Trenton (May, June, 
July, Aug.). 


466. Rotruda mucidella olivaceella (Ragonot), new combination 


Homoeosoma olivaceella Ragonot, Nouv. Gen., p. 33, 1888; Mono- 
graph, pt. 2, p. 244, 1901. 

Homoeosoma musiosum Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 342, 1915 (new synonymy). 

Homoeosoma mucidella Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 342, 1915. 

Homoeosoma cubella Dyar, Ins. Insc. Menstr., vol. 7, p. 62, 1919 
(mew synonymy). 

There is nothing, structural or otherwise, to separate 
the female types of musiosum and cubella from each 
other or from what Dyar correctly identified as muci- 
della from Central America. I have examined the type 
of olivaceella (&, in Mus. Univ. Berlin) as well as its 
genitalia. It is merely a small (13 mm.), somewhat 
rubbed and faded specimen. I am holding olivaceella 
as a tropical American race of mucidella more on geo- 
graphical distribution than anything else. It shows, in 
contrast to typical mucidella, the same color variations 
and obscurity of pattern markings as the eastern United 
States race reliqguella. When examples are recovered 
from the intermediate areas of northeastern México it 
will probably be necessary to sink reliquella into the 
synonymy of olivaceella. The races at best are dubious 
entities, and I doubt that even the distinctness of 
mucidella mucidella will survive extensive collection in 
the central areas of United States. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Alar expanse, 12-18 mm. 

Type Locauitizs: “St. Thomas,’ [Virgin Islands] 
(oliwaceella, in Mus. Univ. Berlin); Porto Bello, Panama 
(musiosum, in. USNM); Baracoa, Cuba (cubella, in 
USNM). 

Foop piants: Unknown. 

Distrisution: Cusa: Baracoa. VircGin IsLanps: St. 
Thomas. GuateMaLa: Cayuga (Apr., May), Quirigué 
(Feb.). Costa Rica: Juan Vifias (June). PanamA: 
Cabima (May), Corozal (Nov.), Porto Bello (Feb., 
Mar.), Rio Trinidad (Mar., May). Brazi: Sania Cata- 
rina, New Bremen; Minas Gerais, Vicosa (Oct.). Para- 
Guay: Villarrica (Sept., Oct.). 


467. Rotruda mucidella affusella (Ragonot), new combination 


Homoeosoma affusella Ragonot, Nouv. Gen., p. 34, 1888; Mono- 
graph, pt. 2, p. 260, 1901. 

The name affusella is doubtfully placed here in associ- 
ation with mucidella, for Ragonot’s description and 
figure suggest something quite different, possibly similar 
to what I have described as Homocosoma vepallidum. 
However, Clarke’s notes on the type discount the figure 
(Monograph, pl. 33, fig. 24) and, to a considerable 
degree, the original description. He states that “the 
figure is poor. The dorsal spot at basal third is dis- 
tinct but the excurved line costad is very faint. The 
s. t. line of dashes, which is so prominent in the figure, 
is scarcely discernible in the specimen. On the fore- 
wing there is a heavier dusting of white than is indi- 
cated in the drawing.’’ This suggests a rather pale, 
not too well marked example of mucidella. We shall 
have to await recovery of examples matching the type 
from Argentina before the status of the name affusella 
can be definitely settled. The abdomen of the type 
was no help. It has been glued on and is obviously 
spurious. I examined the genitalia (?). They are 
those of E’phestia elutella. 

Alar expanse, 18 mm. 

Typr Locauity: Cérdoba, Argentina (type in Paris 
Mus.). 

Foop prant: Unknown. 


Genus 129: Stirephomescinia 


[Venational division B. Forewing with 9 veins: 10 from cell; 
8 and 9 united; 4 and 5 united; 2 and 3 stalked. Male genitalia 
with transtilla incomplete.] 


129, Genus Strephomescinia Dyar 


Strephomescinia Dyar, Ins. Insc. Menstr., vol. 7, pl 60, 1919. 
(Type of genus: Strephomescinia schausella Dyar.) 

Tongue well developed. Antenna pubescent; shaft 
simple. Labial palpus upturned. Maxillary palpus 
filiform. Forewing smooth; 9 veins; veins 2 and 3 long 
stalked from angle of cell; 4 and 5 united; 6 straight; 8 
and 9 united; 10 from the cell but closely approximate 
to 8-9 for some distance; male with weak costal fold. 
Hind wing with vein 2 from well before angle of cell, 
straight and parallel with 3; 3 and 5 from angle and 


227 


closely approximate at base; 6 from upper angle of cell; 
7 and 8 completely fused beyond cell; discocellular vein 
curved; cell about one-fourth the wing length. Abdo- 
men of male with a pair of ventrolateral hair tufts. 

Male genitalia with uncus triangulate. Gnathos 
terminating in a small bifid apical process. Harpe 
simple; costa broadly sclerotized for most of its length. 
Aedeagus straight, simple, moderately slender. Penis 
very weakly scobinate, otherwise unarmed. Vinculum 
stout, short, terminal margin truncate. 

This genus is known only from the male of the type 
species but is easily identified by its venation. The 
genitalia show the close affinity of Strephomescinia to 
both Homoeosoma and Mescinia. 


468. Strephomescinia Schausella Dyar 


Figures 93, 497 


Strephomescinia schausella Dyar, Ins. Insc. Menstr., vol. 7, p. 60, 
1919. 

Forewing whitish gray, irregularly and faintly 
marked with fuscous; a dark streak along submedian 
fold and some dark streaking on the veins toward 
termen; discal spots elongate, separate; no transverse 
lines distinguishable. Hind wing translucent, with the 
veins outlined by dark scaling; terminal and costal 
margins narrowly dark-margined. The one available 
specimen is badly rubbed. Fresh specimens would 
probably exhibit a more definable pattern. Alar ex- 
panse, 9.5 mm. 

Male genitalia with apex of uncus bluntly pointed; 
cucculus of harpe oval, apex evenly rounded; elements 
of divided transtilla long and slender. 

Type tocauiry: Santiago, Cuba (June; type in 
USNM). 

Foop prant: Unknown. 

So far represented only by the unique male type. 


Genus 130: Unadilla 


[Venational division E. Forewing with 9 veins: 10 from cell; 
9 absent (8 and 9 united): 4 absent (4 and 5 united); 2 and 3 
from cell. Hind wing with discocellular vein straight and verti- 
cal. Male genitalia with transtilla incomplete; uncus broad 
throughout. Female genitalia with genital opening simple 
(unsclerotized and without adjacent armature).] 


130. Genus Unadilla Hulst 


Unadilla Hulst, Phycitidae of N. Amer., p. 197, 1890.—Ragonot, 
Monograph, pt. 2, p. xiii, 1901—Hampson, in Ragonot, 
Monograph, pt. 2, p. 261, 1901.—Janse, Journ. Ent. Soc. 
South Africa, vol. 8, p. 25,1945. (Type of genus: Unadilla 
nasutella Hulst.) 

Strymax Dyar, Proc. Ent. Soc. Washington, vol. 47, p. 344, 
1914.—Richards and Thomson, Trans. Ent. Soc. London, 
vol. 80, p. 201, 1932. (Type of genus: Sirymax dorae Dyar. 
New synonymy.) 

Tongue well developed. Antenna pubescent; shaft 
simple. Labial palpus upturned, somewhat obliquely 
so in the female. Maxillary palpus filiform. Forewing 
smooth; 9 veins; veins 2 and 3 separate, 2 near and 3 
from the angle of cell; 3 and 5 closely approximate at 
base; 4 absent; 6 straight; 8 and 9 united; 10 from cell, 


228 


well separated from 8-9; male with a short costal fold. 
Hind wing with vein 2 from well before angle of cell; 3 
from before but near angle, approximate to 5; 6 from 
near upper angle of cell; 7 and 8 completely fused be- 
yond cell (in occasional specimens a rudiment of 8 visible 
as a short spur from outer fifth of vein 7) ; discocellular 
vein straight and vertical; cell approximately one-third 
the length of the wing. Eighth abdominal segment of 
male simple. 

Male genitalia with uncus broad, subtriangular, 
outer surface densely covered with bristlelike scales. 
Gnathos terminating in a moderately large, fused or 
partially fused, hooked apical process. Harpe simple; 
costa, strongly sclerotized for most of its length; apex 
broadly rounded. Anellus U-shaped; lateral arms 
curved and partially encircling aedeagus. Aedeagus 
straight; somewhat constricted at apex; smooth or 
with a patch of minute scobinations at apex. Penis 
with some scobinate wrinklings, otherwise unarmed. 
Vinculum stout and broad, long and tapering slightly 
to more or less narrowly rounded terminal margin. 

Female genitalia with bursa copulatrix membranous, 
finely scobinate over caudal half of inner surface; 
signum consisting of a girdle of fine, narrow, serrate, 
ridgelike disks about middle of bursa, or completely 
absent. Ductus bursae membranous throughout, finely 
scobinate towards junction with bursa. Genital open- 
ing unsclerotized. Ductus seminalis from bursa near 
its junction with ductus bursae. 

Richards and Thomson (1932) referred Strymax 
Dyar to Ephesiia as a subgenus. They did not treat 
Unadilla. Hulst erected the latter for a single species 
(nasutella Hulst) which Hampson placed with two other 
Zeller species (ubacensis and maturella) in the synonymy 
of erronella Zeller. Upon the basis of this synonymy, 
which is doubtful, to say the least, Hampson cites 
erronella as the type of Unadilla. Janse follows him 
in this. Nomenclatorily nasutella must be the type of 
Unadilla whether nasutella proves to be a synonym of 
erronella or not. 

Strymazx is an obvious synonym of Unadilla. It has 
no close affinity to Ephestia even in the broad sense in 
which that genus is interpreted by Richards and 
Thomson. Hind wing venation and genitalia seem to 
indicate much closer relation to Homoeosoma. 

On structures of the male and female genitalia the 
species divide into two distinct groups, as follows: 

Male with apical process of gnathos partially fused; female 
with an encircling band of signa. 


Male with apical process of gnathos completely fused; 
female without signa. 


Genus Unadiila, Species 469: U. erronella 


[Male with apical process of gnathos partially fused; female 
with an encircling band of signa.] 
469. Unadilla erronella (Zeller) 
Fiaures 121, 498, 995 


Homoeosoma erronella Zeller, Horae Soc. Ent. Rossicae, vol. 16, 
p. 238, 1881. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Homoeosoma ubacensis Zeller, Horae Soc. Ent. Rossicae, vol. 16, 
p. 239, 1881. 

Unadilla erronella (Zeller) Hampson, in Ragonot, Monograph, 
pt. 2, p. 262, 1901. 

Ephestia bipunctella Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 
7, p. 255, 1901 (new synonymy). 

Strymazx dorae Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 344, 1914 
(new synonymy). 

Strymaz pyllis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 344, 1914 
(new synonymy). 

Ephestia (Strymazx) bipunctella (Hampson) Richards and Thom- 
son, Trans. Ent. Soc. London, vol. 80, p. 201, 1932. 


Ephestia (Strymaz) pyllis (Dyar) Richards and Thomson, 
Trans. Ent. Soc. London, vol. 80, p. 202, 1932. 


Hampson referred ubacensis as a synonym of erronella, 
and, I believe, correctly so, judging from photographs 
of the male types and their genitalia. The only differ- 
ence between the two is in the more intensified macu- 
lation of ubacensis. The same difference is exhibited 
by the male types of dorae and pyllis, the latter being a 
pale example with the normal subbasal markings of 
forewing almost obsolete. However, in a series of 
dorae from Panamé there are several intergrading 
examples. U. erronella is also variable in size and to 
some extent in structure, the vinculum being shorter in 
some specimens from Puerto Rico than in those from 
Panamé4 and Colombia, but otherwise the genitalia are 
remarkably constant. In careful preparations of the 
genitalia of Dyar’s supposed two species (dorae and 
pyllis) I am unable to find any of the differences shown 
in the figures published by Richards and Thomson. 
The general color of the forewing is whitish gray; sub- 
basal markings, when present, consisting of from one to 
three pale brownish spots forming a broken antemedial 
transverse shade; discal dots faint; hind wing white in 
the male, more or less shaded with pale smoky fuscous 
in the female. Alar expanse, 9-14 mm. 


TYPE LOCALITIES: Honda, Colombia (erronel/a, in 
BM); Ubaque, Colombia (ubacensis, in BM); Nassau, 
Bahamas (bipunctella, in BM); La Chorrera, Panamd 
(dorae, in USNM); Porto Bello, Panama (pyllis, in 
USNM). 


Foop piant: Unknown. 


DistrisuTion: Conompia; Honda, Mariquita, 
Ubaque. Panamd: Corazal (Mar., Apr., May), La 
Chorrera (Apr., May), Paraiso (Apr.), Porto Bello 
(Feb.), Rio Trinidad (Mar., May), Taboga Isl. (Feb., 
June). Banamas: Nassau. Pusrto Rico: Aguirre 
Central (Apr.), Cataéno (Apr., June), Coamo Springs 
(Apr.), Dorado (May, June), Isabela (Apr.), San 
German (Aug.); Puerto Real (Vieques Isl., Apr., July). 
Virein Isuanps: Kingshill (St. Croix, Mar., Dec.). 

The Puerto Rican specimens (some 40-odd in the 
Cornell University Collection) may possibly represent 
a distinct race from the mainland form, but I am 
unable to find any valid character for their separation. 
The somewhat shorter vinculum of the male is matched 
in some Panamé specimens and is of very doubtful 
significance. U. erronella is easily identified as it is 
the only described species with a partially divided 
apical process of gnathos and a girdle of signa. The 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


remaining species (with the possible exception of 
nasutella) belong in the following group. 


Genus Unadilla, Species 470-472: U. maturella 
to U. floridensis 


[Male with apical process of gnathos completely fused; female 
without signa.] 


470. Unadilla maturella (Zeller) 
Fieures 499, 996 


Homoecosoma maturella Zeller, Horae Ent. Soc. Rossicae, vol. 16, 
p. 240, 1881. 


Hampson made maturella a synonym of erronella, 
but a sketch by Clarke of the male genitalia of the male 
type shows a completely fused apical process of gnathos 
which places maturella in this group of species. A 
photograph of the type also shows a pale grayish white 
form with a strong, dark, transverse, antemedial shade 
extending on a straight, outward slant from below costa 
almost to inner margin. I have before me a series of 
similarly marked specimens from Guatemala and Cuba 
with genitalia similar to those of Zeller’s type, and 
presumably conspecific. Alar expanse, 14-16 mm. 

TypE Ltocatity: Colombia (type in BM). 

Foop prant: Unknown. 

Distrisution: Cotompia. GuaTEMALA: Cayuga 
(Mar., Apr., May). Cusa: Baracoa (May), Havana. 


471. Unadilla albidiorella (Richards and Thomson), new combination 


Ephestia (Strymaz) albidiorella Richards and Thomson, Trans, 
Ent. Soc. London, vol. 80, p. 201, 1932. 


Described from a single male. From description and 
figure published by the authors it must be very close to 
maturella if not a race of that species. I have seen no 
Peruvian examples of Unadilla. Alar expanse, 17 mm. 

Typr tocatity: Rio Ucayale, Contamino, Pert 
(“Oct.—Dec.,”’ type in BM). 

Foop pirant: Unknown. 


472. Unadilla floridensis, new species 


Darker and more decidedly gray than the other 
species of the genus; dark pattern markings faint, in 
some specimens almost obsolete, in others conspicuous 
only as a single antemedial shade or spot and a narrow 
fainter dark inner shade along subterminal line of fore- 
wing; white dusting rather sparse. Alar expanse, 
8-13 mm. 

The genitalia (co and 9) are similar to those of 
maturella, exhibiting no differences of a specific nature. 


Type Locatity: Key West, Fla. (type in USNM, 
61377). 

Foop piants: Pulchea odorata, Melanthera radiata. 

Described from male type and 10 male and 4 female 
paratypes from the type locality, reared Apr. 23 to May 
1, 1945, by the Special Survey of the Division of Foreign 
Plant Quarantine, U. S. Bureau of Entomology and 
Plant Quarantine, from larvae feeding in the blossoms 


229 


and seed pods of Pulchea and Melanthera. These are 
the only food-plant records we have for the genus 
Unadilla. The larvae are similar in markings and 
habitus to those of Rotruda mucidella and like them lack 
the sclerotized rings about seta IIb of mesothorax and 
seta IIT of the eighth abdominal segment. 


Genus Unadilla, Species 473 (unplaced): U. 
nasutella 


473. Unadilla nasutella Hulst 
Unadilla nasutella Hulst, Phycitidae of N. Amer., p. 197, 1890. 


The male type in the Rutgers Collection lacks an 
abdomen, so it will be impossible to place nasutella in 
its proper species group or to determine its synonymy 
until more specimens from the southwestern United 
States are discovered. Except for the Hulst type I 
have seen no example of Unadilla from New Mexico, 
Arizona, or Texas. Alar expense, 13.5 mm. 


Type tocauity: Hot Springs, N. Mex. (type in 
AMNH, ex Rutgers). 

Foop piant: Unknown. 

Hampson referred nasutella as a synonym of erronella 
Zeller and it so stands in our lists. This synonymy is 
probably incorrect and is certainly unwarranted on the 
evidence before us. 


Genera 131-151: Laetilia to Cactobrosis 


{Venational division A. Forewing smooth, oblong, broadest 
toward termen; color alike in both sexes; 11 veins; 10 from cell, 
8 and 9 stalked, 6 straight, 4 and 5 stalked, 2 and 3 separate or 
approximate, 2 from near lower outer angle of cell; no costal 
fold or other secondary sexual modifications. Hind wing with 
7 veins; 3 and 5 connate or stalked; 2 from before lower outer 
angle of cell; discocellular vein curved, complete; no sex-scaling 
or other sexual modifications; cell one-half or somewhat less 
than one-half the length of the wing. Abdomen of male with a 
pair (rarely two pairs) of ventrolateral hair tufts at base of 
eighth segment or with eighth segment simple. Male genitalia 
with uncus broad, subtriangular, never hook-shaped or otherwise 
modified, apex broadly rounded, its outer (dorsal) surface densely 
covered with bristlelike scales; gnathos terminating in a flanged 
(or lobed) and hooked apical process, frequently bifid, sometimes 
fused or partially fused; transtilla incomplete (represented by 2a 
pair of separate, more or less elongate-triangulate plates), never 
forming a bridge or otherwise modified; harpe simple, without 
clasper or extensions from sacculus or costa; anellus U-shaped; 
aedeagus straight or slightly sinuate, usually smooth, but occa- 
sionally with a few very small scobinations at apex; penis smooth, 
finely scobinate or with sclerotized wrinklings but not other- 
wise armed; vinculum stout and broad, short or long, and with 
terminal margin normally broadly rounded. Female genitalia 
with bursa copulatrix membranous, smooth or more or less mi- 
nutely scobinate on inner surface, never strongly sclerotized or 
pigmented; signum frequently absent, when present consisting 
of a small, ribbed, weakly serrate, finely scobinate or cupped 
plate; ductus bursae membranous throughout or scobinate for a 
short distance from genital opening, gradually widening into and 
not sharply differentiated from bursa copulatrix; genital opening 
normally simple, sometimes minutely scobinate, occasionally 
with sclerotized dorsal or ventral plates or a few setae on the inner 
surfaces of the ductus at the opening, otherwise unmodified or 
unarmed; ductus seminalis from bursa, rarely from ductus bursae 
near junction with bursa.] 


230 


131. Genus Laetilia Ragonot 


Laetitia Ragonot, Entomologica Americana, vol. 5, p. 116, 1889; 
Bull. Soc. Ent. France, p. viii, January 1890.—Hulst, 
Phycitidae of N. Amer., p. 182, 1890.—Hampson, in Rago- 
not, Monograph, pt. 2, pp. 116, 560, 1901.—Forbes, Cornell 
Mem. 68, p. 631, 1920. (Type of genus: Dakruma cocci- 
divora Comstock.) 

Laosticha Hulst, U. S. Nat. Mus. Bull. 52, p. 431, 1902.—Dyar, 
Proc. Ent. Soc. Washington, vol. 6, p. 159, 1904. (Type of 
genus: Dakruma ephestiella Ragonot.) 

Tongue short (greatly reduced in melanostathma and 
glomis, but not completely enclosed and hidden by the 
labial palpi). Antenna pubescent, shaft simple in both 
sexes. Labial palpus oblique or obliquely upturned 
(males of melanostathma and zamacrella). Maxillary 
palpus filiform or subsquamous (coccidivora). Hind 
wing with veins 7 and 8 anastomosing for at least half 
their distance beyond cell; 3 and 5 stalked. Highth 
abdominal segment of male simple. 

Male genitalia with apex of uncus normally broadly 
rounded (somewhat narrowly rounded in coccidivora, 
cardini, and myersella) ; apical flanged process of gnathos 
stout, the lobes fusing posteriorly; aedeagus smooth, 
moderately stout; penis without cornuti, rarely with 
a few sclerotized folds (zamacrella, myersella), otherwise 
smooth; vinculum stout, rarely longer than broad 
(jiskella). 

Female genitalia with signum consisting of a narrow, 
flanged or a small rounded, bluntly dentate plate; bursa 
copulatrix, except for signum and occasional minute 
scobinations surrounding the signum, smooth; ductus 
bursae with a sclerotized ventral plate and a pair of 
dorsal sclerotized plates at genital opening, otherwise 
smooth; ductus seminalis from anterior end of bursa 
copulatrix. 

Larvae predaceous on scale insects. 

The genus as here defined is structurally a somewhat 
composite group but is, I think, a natural one, linking 
in one direction with Rhagaea, Zophodia, and the cactus- 
feeding phycitine genera and on another with two or 
three as yet undefined genera of coccid feeders in the 
Anerastiinae. Typical ZLaetilia (coccidivora to portori- 
censis) are as much an erastiine as phycitine on male 
genitalic characters (especially the general habitus of 
the organs as well as the peculiar development of the 
gnathos), but the tongue, while much reduced, is dis- 
tinetly exposed as in most other short-tongued Phyci- 
tinae. Three of the transition species (melanostathma, 
amphimetra, and glomis) have the tongue rudimentary 
and on this structure could go into the Anerastiinae, 
were it not for their genitalia. The remaining transi- 
tion species (zamacrella, myersella, ephestiella and jfis- 
kella), all nearctic species, are definitely phycitine on 
all characters. On the sum of its characters Laetillia 
must be included in the Phycitinae. Upon one or more 
structural differences it can be distinguished from any 
other phycitine or anerastiine genus. 


474. Laetilia coccidivora (Comstock) 
Fiaures 63, 502, 503, 997 


Dakruma coccidivora Comstock, North Amer. Ent., vol. 1, p. 26, 
1879; Rep. Comm. Agr. for 1879, p. 241, 1880.—Packard, 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


U. 8. Ent. Comm. Bull. 7, p. 54, 1881; U.S. Dep. Agr. Fifth 
Rep. Ent. Comm., p. 413, 1890. 

Dakrum pallida Comstock, Rep. Comm. Agr. for 1879, p. 243, 
1880. 


Laetilia coccidivora (Comstock) Ragonot, Ent. Amer., vol. 5, p. 
116, 1889; Bull. Soc. Ent. France, p. viii, 1890.—Hulst, 
Phycitidae of N. Amer., p. 182, 1890; U. S. Nat. Mus. Bull. 
52, p. 431, 1903.—Hampson, in Ragonot, Monograph, pt. 2, 
p. 117, 1901.—Hunter, Pratt, and Mitchell, U. S. Dept. 
Agr. Bur. Ent. Bull. 113, pp. 24, 26, 1912.—Forbes, Cornell 
Mem. 68, p.632, 1920.—McDunnough, Check list, No. 6328, 
1939. 


Zophodia dilatifasciella Ragonot, N. Amer. Phycitidae, p. 13, 
1887.—Hulst, Phycitidae of N. Amer., p. 174, 1890; U. S. 
Nat. Mus. Bull. 52, p. 429, 1903.—Hampson, im Ragonot, 
Monograph, pt. 2, p. 21, 1901.—Hunter, Pratt, and Mitchell, 
U.S. Dep. Agr. Bur. Ent. Bull. 113, pp. 24, 26, 1912.—Mc- 
Dunnough, Check list, No. 6309, 1939. (New synonymy.) 


Laetilia coccidivora hulstit Cockerell, Amer. Nat., vol. 31, p. 588, 
1897.—Hulst, U. S. Nat. Mus. Bull. 52, p. 431, 1903.— 
MeDunnough, Check list, No. 6328a, 19389. 


Laetilia hulstii (Cockerell) Dyar, Proc. Ent. Soc. Washington, 
vol. 6, p. 228, 1904. 

Palpi, face, head, and thorax brownish fuscous dusted 
with white, the white more pronounced on undersurface 
of second segment of labial palpus, on head, and on 
collar. Forewing pale brownish fuscous heavily dusted 
with white and with dark brown bordering the trans- 
verse lines; the white dusting concentrated on the costal 
half of wing, giving this area (especially between cell 
and costa) a distinctly whitish appearance; area between 
cell and inner margin (and, beyond cell, between vein 
3 and tornal margin) more or less suffused with pale 
ocherous fuscous; costa for a short distance from base 
blackish brown; in most specimens a rather obscure 
transverse pale brownish basal band extending from 
just below base of costa to inner margin near antemedial 
line (only observable on unfaded specimens, sometimes 
obsolete); from middle of basal band a short blackish 
streak; antemedial line slanting outwardly from costa, 
slightly angled or bent at fold, white and margined in- 
wardly by a very narrow blackish brown band which 
fades out towards costa, and outwardly margined by a 
broad blackish brown band, especially contrasted to- 
wards costa; subterminal line, faint, smuate (inwardly 
angled opposite cell and also, but very slightly, at fold), 
inwardly margined by a narrow blackish brown band 
and outwardly edged for a short distance from costa 
by a broader, more or less triangular, blackish shading; 
a small blackish dot or line at upper angle of cell and 
another at lower outer angle, these occasionally fused 
but normally well separated; along termen at the vem 
ends a row of obscure brownish dots; cilia ashy gray. 
Hind wing whitish with a somewhat smoky tint, a dark 
shading along costa, a narrow dark line along termen 
and some dark shading on the veins; cilia white with a 
broad, slightly darker, basal band. Alar expanse, 
10-20 mm. 

Male genitalia with gnathos terminating in a flanged 
process with drooping lobes and a pair of short, strongly 
sclerotized, divergent posterior horns; apex of uncus 
truncate; penis simple, without cornuti or sclerotized 
folds. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 231 


Female genitalia with signum rather large, a single, 
elongate, strongly sclerotized, pocketlike projection into 
bursa; bursa copulatrix, except for signum, smooth. 

Type Locauities: Washington, D. C. (coccidivora, in 
USNM); Sanford, Fla. (pallida, in USNM); Las Cruces, 
N. Mex. (hulstti, in AMNH, ex Rutgers); Arizona 
(dilatifacsiella, in Paris Mus.). 

Foop: Various Coccidae.'2 Specific records include 
Pulvinaria vitis (L.) (originally identified as Lecanium 
acericola or Pulvinaria innumerabilis), Pulvinaria amyg- 
dali Ck11., Pulvinaria bigeloviae Ck11., Toumeyella 
numismaticum (Ck11.), Toumeyella spp., Coccus hesperi- 
dum L., Eriococcus quercus (Coms.), Saissetia oleae 
(Bern.), Kermes spp., Lecaniodiaspis sp., Cerococcus 
quercus Coms., Lecanium arizonensis King, Dactylopius 
sp., Dactylopius confusus Ck11., Neolecanium cornu- 
parvum (Tho.), Dactylopius tormentosus (Lam.), Pseu- 
dococcus sp. The larvae will also feed in flowers of 
cactus (Platypuntia spp.) 

Distrisution: District of Columbia (July); Virginia, 
Falls Church (Aug.), Vienna (July); Maryland, Plum- 
mers Isl. (June); Pennsylvania, Longhorne (Oct.), 
Rockville (Apr.); Ohio, Newark (Nov.); Florida, no 
specific locality (May), Everglades (Apr.), Chokoloskee, 
Miami (Feb.); Alabama, Mobile (June); Mississippi, 
Biloxi (June); Louisiana, Baton Rouge (Aug., Nov.), 
New Orleans; Texas, Sabinal (Mar.), Victoria (Feb.), 
Brownsville (Jan., Feb., Apr.), Kerrville (Apr., May), 
San Antonio (Oct.), Uvalde (May, Aug.), Sterling City 
(Oct.), Dallas (Nov.), Corpus Christi (Feb.); New 
Mexico, Mesilla Park (June), Hot Springs; Arizona, 
Tucson (June, Nov.), Paradise (Mar., Sept., Oct.), 
Redington, Palmerlee, Huachuca Mts., Wilgus Mts., 
Baboquivari Mts. (July, Sept., Oct.); California, San 
Bernardino (Nov.), San Diego (Oct.), Palm Springs, 
no locality (Mar.). 

There is considerable variation in size in this species 
and corresponding differences in the shape of the 
flanged processes of the gnathos and the length of the 
lateral arms of anellus, but nothing that would justify 
separation into races; for the extremes of difference in 
size and structure are present in the reared cotype series 
from Washington, D. C. The name dilatifasciella ap- 
plies to Arizona specimens which average larger than 
those from eastern localities and show some occasional 
traces of ocherous shading on the lower half of forewing. 
Here also there is no consistency which would justify 
even a racial separation. The name has escaped 
synonymy so far only because specimens identified to 
it have been placed in the wrong genus. The larvae of 
coccidivora like those of other species of Laetilia are 
predaceous on the Coccidae. For obvious reasons they 
do not attack the smaller hard-scaled species but other- 
wise are not restricted, feeding on any of the larger 
scales and mealy bugs that occur in colonies of sufficient 
number to provide food. They seem to be fond of the 


122 Dr. Harold Morrison was good enough to check the coccid 
records in connection with rearings of L. coccidivora and has 
supplied the names of coccids here given. 


cochineal scale on cactus and on this plant will occa- 
sionally vary their diet by feeding upon the flowers. 
Several moths have been reared from larvae which fed 
altogether upon the flowers of Platypuntia. As a conse- 
quence coccidivora has been listed as a cactus insect; but 
this plant feeding is, I believe, only a departure from 
the normal predaceous habit. Certainly coccidivora 
does not favor the coccids on cactus above those on 
other plants. Its impartiality in diet and lack of host 
restriction are probably accountable for its ineffective- 
ness as an agent of control in scale infestations. 


475. Laetilia coccidivora quadricolorella (Dyar), new combination 
Fiaures 504, 999 


Atascosa quadricolorella Dyar, Proc. Ent. Soc, Washington, vol. 6, 
p. 116, 1904. 

Poujadia quadricolorella (Dyar), Barnes and McDunnough, 
Check list of the Lepidoptera of Boreal America, no. 5819, 
1917.—MecDunnough, Check list, No. 6422, 1939. 

Differs from that of typical coccidivora in that the 
white dusting is more abundant on head, thorax, and 
forewing, and the brownish fuscous shading largely re- 
placed by ocherous; the basal patch ocherous with a 
narrow outer border of blackish scales; band outwardly 
bordering the antemedial line ocherous with a blackish 
brown dash or narrow smudge at top of cell; area 
between cell and inner margin and between antemedial 
and subterminal lines more or less ocherous (in Cali- 
fornia specimens before me completely so) ; subterminal 
line narrowly bordered inwardly and outwardly by 
blackish scales, especially towards costa; black scales 
more or less lining the upper margin of cell; blackish 
discal dots at end of cell and along termen at vein ends 
rather sharply contrasted against the white dusting of 
the ground color. Hind wing clear white with no dark 
scaling on the veins; along termen a narrow brown line. 
Alar expanse, 15-20 mm. 

Male genitalia with apex of gnathos more rounded, 
posterior horns of apical process of gnathos somewhat 
shorter and closer together than those of typical 
coccidivora; but otherwise substantially the same. 
Female genitalia not essentially different from those of 
typical coccidivora. 

Type tocauiry: Las Cruces, N. Mex. (type in 
USNM). 


Foop: Unknown (none of the specimens before me 
has been reared but the larvae presumably feed upon 
more than one species of coccids). 

Distrisution: New Mezico, Las Cruces, Fort Win- 
gate (May); Arizona, Redington; California, Mojave 
(Aug.), Olancha (May, June). 

This local color variety (it is hardly more than that) 
has probably escaped synonymy because it was de- 
scribed in the wrong genus and subfamily. When it is 
reared we may discover that it has no right even to 
racial distinction. In New Mexico and Arizona it and 
typical coccidivora occur in the same localities. The 
California specimens before me, however, show such 
marked contrasts in color (intensity of the white dusting 
and light ocherous tint of the normally brownish areas 


232 


of forewing) that I hesitate to sink the name completely. 
It is possible that hulstii Cockerell and quadricolorella 
stand for the same form (types of both are from Las 
Cruces and have clear white hind wings); but the type 
of the former (9) is unrecognizable and the name may 
very well be left where it is. 

Dyar’s type is a male and not a female as he stated 
in his description. 


476. Laetilia coccidivora cardini Dyar 
Fiaures 505, 998 


Laetilia cardini Dyar, Ins. Inse, Menstr., vol. 6, p. 139, 1918. 


Not distinguishable from small specimens of typical 
coccidivora except that the hind wing is somewhat 
darker. In addition to the female type I have before 
me another female from the type locality and three 
males and eight females from Orlando, Fla. The 
Florida specimens were reared from larvae feeding on 
mealy bugs on grapefruit. 

The female genitalia differ only in their smaller size 
from genitalia of normal coccidivora, but even in this 
detail do not differ from equally small examples of 
coccidwora. The male genitalia show apex of uncus 
rather more pointed and the posterior horns of apical 
process of gnathos more widely spaced than those of 
typical coccidivora. However there is considerable 
variability in these structures in coccidivora and the 
differences exhibited by the male of cardini may not be 
constant. In my opinion they do not justify more than 
racial separaton. 

TypE LocaLity: Santiago de las Vegas, Cuba (type 
in USNM). 

Foop: Mealy bugs. 

Distrizution: Cusa: Havana, Santiago de las 
Vegas. Unrtep Statss: Florida, Orlando (June). 


477. Laetilia obscura Dyar 
Ficures 506, 1000 


Laetilia obscura Dyar, Ins. Insc. Menstr., vol. 6, p. 140, 1918. 


Brownish fuscous with a sparse scattering of whitish 
scales on head, thorax, and subcostal area of forewing. 
Usual transverse markings on forewing nearly obsolete; 
antemedial line very faint, straight, shaded outwardly 
towards costa by dark brown; subterminal line a mere 
trace; discal dot obscure; cilia very little paler than 
ground color. Hind wing pale smoky fuscous, darkened 
outwardly and with blackish brown terminal margin; 
cilia slightly paler with dark basal band. Alar expanse, 
9-11 mm. 

Male genitalia similar to those of coccidivora cardini 
but with posterior horns of gnathos more widely 
spaced. Female genitalia with signum similar to but 
weaker and narrower than that of coccidwora or any of 
its varieties. 

Typ Locauity: Santiago de las Vegas, Cuba (type 
in USNM). 

Foop: Presumably Coccidae (type series apparently 
reared but without labels to indicate food or dates of 
emergence). 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Distrizution: Cusa. 

Represented in the National Collection only by 
specimens from the type locality. It may prove to be 
merely a dark, suffused race of coccidivora; but the 
uniformly dark color, the almost complete obliteration 
of the usual transverse markings, and the much narrower 
signum suggest a distinct species. 


478. Laetilia portoricensis Dyar 
Fiaures 500, 1001 


Laetilia portoricensis Dyar, Ins. Insc. Menstr., vol. 3, p. 62, 
1915.—Wolcott, Journ. Agr. Univ. Puerto Rico, vol. 20, 
No. 1, p. 479, 1936. 

Palpi, face, head, and thorax pale brownish fuscous 

with a few white scales on inner side of palpus and on 
face. Forewing pale brownish fuscous with area 
between costa and upper vein of cell and antemedial 
and subterminal lines white; on this white area a 
rather conspicuous midcostal, blackish brown dash; 
basal area paler than ground color, whitish toward 
costa; transverse pale lines obscure, indicated chiefly 
by their dark outer borders; antemedial line slightly 
angled at cell, bordered outwardly by blackish brown 
which is conspicuous on costa and forms a patch at the 
cell; subterminal line outcurved between veins 2 and 5, 
rather broadly bordered with blackish brown towards 
costa; a blackish brown discal dot at upper outer angle 
of cell; more or less blackish shading along submedian 
fold and upper vein of cell; a few indistinct dark spots 
on pale termen between the veins; cilia concolorous 
with pale terminal margin. Hind wing pale smoky 
brown; terminal margin dark brown; cilia sordid 
whitish with dark basal band. Alar expanse, 10-13 
mm. 
Male genitalia with gnathos terminating in a, pair of 
rather weakly sclerotized, liplike lobes without posterior 
horns; uncus rather broad but laterally folded, apical 
margin rounded; penis without cornuti. Female geni- 
talia with signum a rather small, nearly round, sclero- 
tized, scobinate disk; bursa copulatrix, except for 
signum and area immediately adjoining it, smooth; 
vinculum long. 

Typ Locality: Rio Piedras, Puerto Rico (type in 
USNM). 

Foon: Saissetia oleae, Lecanium sp. 

Distrisution: Purrto Rico: Rio Piedras (Nov.), 
Bayamén (Aug.), Comerio (Aug.). 

The genitalia and the midcostal dark dash on fore- 
wing readily distinguish this species from anything else 
in the genus. Dyar described it as “gray,” but the 
ground color and general effect, except for the contrasted 
whitish subcostal area, is more brown than gray. 


479. Laetilia melanostathma (Meyrick), new combination 
Fieures 501, 507, 1002 


Euzophera melanostathma Meyrick, Exotic Microlepitoptera, vol. 
5, pt. 5, p. 134, 19387, 


Palpi, head, and thorax pale grayish brown. Fore- 
wing light gray with a whitish suffusion in subcostal 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


area and a faint lilac tint due to sparsely scattered, 
reddish brown scales; antemedial line very faintly in- 
dicated, straight, slanting inwardly from costa, and 
bordered outwardly in cell by a blackish blotch, this 
black marking not continued to costa; at outer end of 
cell a similar conspicuous, large, round black spot; a 
faint dark streak on midcosta; subterminal line faint, 
sinuous, bordered outwardly and inwardly by blackish 
bands, the latter rather broad and well defined from 
costa but fading out toward inner margin; a few irregu- 
lar black dots on termen between the veins; cilia 
slightly paler than ground color of the wing. Hind 
wing very pale smoky fuscous, translucent and almost 
white towards base, somewhat darker in the female; 
terminal margins dark; cilia slightly paler than wing. 
Alar expanse, 11-12 mm. 

Male genitalia with gnathos terminating in a broad 
heart-shaped process composed of two partially cleft, 
partially fused lobes ending in a single, slender posterior 
horn; vinculum rather long and broad; penis without 
cornuti. 

Female genitalia with ductus bursae strongly and 
broadly sclerotized on ventral side toward genital open- 
ing; bursa copulatrix minutely scobinate; signum 
similar to that of coccidivora, but smaller. 

Type Locatiry: Concordia, Entre Rios, Argentina 
(type in BM). 

Foon: Tachardiella argentina. 

As yet known only from the type locality. 

The four specimens examined had been reared (Nov. 
1936) and were submitted by K. J. Hayward, and pre- 
sumably formed part of the original series from which 
the type and paratypes were selected. 

This species is a somewhat anomalous Laetilia. The 
very short tongue, short and acuminate third segment 
of labial palpus, and definitely filiform maxillary palpus 
suggest other generic placement; but the genitalia show 
that it is closely allied to typical Laetilia. It certainly 
does not belong in Euzophera where it was originally 
placed. The two conspicuous black spots on the cell 
of forewing will identify it specifically, and distinguish 
it from other described species of the genus with the 
possible exception of amphimetra. 


480. Laetilia amphimetra (Meyrick), new combination 


Euzophera amphimetra Meyrick, Trans. Ent. Soc. London, vol. 
89, pt. 4, p. 47, 1939. 

This species is known to me only from the description. 
In the National Collection there are five badly rubbed 
specimens (one o' and four 9) reared from larvae 
feeding on a Ceroplastes sp. at Valemar, Chile, by 
P. A. Barry (“9-21-40”) which might be amphimetra. 
What is left of the pattern matches well enough with 
Meyrick’s description. The markings are similar to 
those of melanostathma and the male genitalia are 
identical, except that the vinculum is shorter in the 
Chilean specimen. It and its accompanying females, 
however, are appreciably darker than typical examples 
of melanostathma. If the Chilean specimens are 


233 


amphimetra, the latter is probably no more than a color 
variety of melanostathma. 

Typr tocauity: Concordia, Entre Rios, Argentina 
(type in BM). 

Foon: Ceroplastes sp. on guava. 


481. Laetilia zamacrella Dyar 
Ficures 508, 1004 


Laetilia zamacrella Dyar, Ins. Insc. Menstr., vol. 13, p. 12, 
1925.—McDunnough, Check list, No. 6327, 1939. 


Palpi, face, head, thorax, and forewing grayish 

fuscous rather evenly dusted with white (the head 
densely so) making the general color to the naked eye a 
pale slate gray. Forewing blackish at base for a very 
short distance; antemedial line nearly vertical, slightly 
indented at fold, white, bordered within and without by 
blackish brown, outer dark border appreciably broad- 
ened at top of cell; a conspicuous blackish discal spot 
at outer margin of cell; subterminal line sinuate (out- 
wardly angled between veins 1 and 5), white, narrowly 
bordered inwardly and outwardly by blackish fuscous; 
along termen a row of more or less fused blackish spots; 
cilia ashy gray. Hind wing semihyaline with a pale 
fuscous shade bordering costa and a fine fuscous line 
along termen from apex to vein 1b; veins slightly darker 
than wing membrane; cilia white. Alar expanse, 24-25 
mm. 
Male genitalia with apical process of gnathos par- 
tially fused, large, the lobes broadly flaring, the posterior 
projections stubby and close together; penis with 
several long sclerotized folds but without cornuti. 
Female genitalia with a pair of strongly sclerotized 
dorsal plates in ductus bursae at genital opening; 
ventral plate at opening large; signum a small flanged 
plate; bursa copulatrix smooth except for a few minute 
scobinations surrounding signum. 

Typrgz xocauity: Mount Wilson, Calif. 
USNM). 

Foon: Scales on pine. 

Distrisution: California, Mount Wilson (July), 
Mount Hamilton (Apr., May). 

The seven Mount Hamilton specimens were reared 
by H. E. Burke, under Hopkins No. 19013d, from 
larvae feeding on an unidentified scale on Pinus radiata. 
The moth superficially resembles Dioryctria abietella 
and might easily be mistaken for a Dioryctria species 
except for its reduced hind wing venation and the 
black rather than white discal spot on forewing. Its 
genitalia, except for their squat appearance, resemble 
those of Zophodia and Cactobrosis and show the close 
affinity of Laetilia to the cactus-feeding phycitine 
genera, 


(type in 


482, Laetilia myersella Dyar 
Fiaures 509, 1003 
Laetilia myersella Dyar, Proc. Ent. Soc. Washington, vol. 12, 
p. 54, 1910.—Forbes, Cornell Mem. 68, p. 632, 1920.— 
McDunnough, Check list, No. 6330, 1939. 
Palpi, face, head, and thorax brownish fuscous sparse- 
ly dusted with white, the white dusting most conspic- 


234 


uous on the middle third of tegulae, throwing into con- 
trast their dark basal and even darker apical areas. 
Forewing ashy white with brownish areas forming a 
rather strongly contrasted pattern; basal area brownish 
fuscous with a narrow pale subbasal border; antemedial 
line well marked, somewhat sinuate and slanting out- 
ward from costa to immer margin, white, with a narrow 
dark inner border and a broad darker outer border 
which is appreciably broadened towards inner margin; 
discal dots fused into a large, dark spot at end of cell; 
subterminal line slightly sinuate (outwardly angled be- 
tween veins 1 and 5), white, rather broadly bordered 
inwardly and outwardly by dark brownish fuscous; 
terminal area of the pale ground color with terminal 
margin dark brown; cilia pale brownish fuscous. Hind 
wing pale fuscous with a dark line along termen; cilia 
whitish. Alar expanse, 14-16 mm. 

Male genitalia with apical process of gnathos partially 
fused, large, the lobes flaring but not so broadly so as 
those of zamacrella, the posterior projections sharp and 
appreciably separated; penis with several long sclero- 
tized folds but without cornuti (similar to that of 
zamacrella) . 

Female genitalia with a pair of strongly sclerotized 
plates in ductus bursae at genital opening; ventral 
plate at opening large; signum a small plate with from 
two to four bladelike projections; bursa copulatrix 
smooth except for a few minute scobinations surround- 
ing signum. 

Typn LocaLity: Rockville, Pa. (type in USNM). 

Foon: Scales on pine. 

Distrisution: Pennsylvania, Rockville (May) ; North 
Carolina, Tryon, Southern Pines (Sept.). 

Twelve specimens examined. 

This species is apparently confined to the Eastern 
States. It is readily separable on habitus and structure 
from the western zamacrella but is obviously closely 
related to that species. 


483. Laetilia ephestiella (Ragonot) 
FicureE 510 


Dakruma ephestiella Ragonot, N. Amer. Phycitidae, p. 13, 1887. 

Laetilia ephestiella (Ragonot) Hulst, Phycitidae of N. Amer., 
p. 185, 1890.—Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 159, 1904; vol. 7, p. 37, 1905.—McDunnough, Check list, 
No. 6326, 1939. 

Lasiosticha ephestiella (Ragonot), Monograph, pt. 2, p. 109, 1901. 

Laosticha ephestiella (Ragonot) Hulst, U. S. Nat. Mus. Bull. 52, 
p. 431, 1903. 

Maricopa lustrella Dyar, Proc. Ent. Soc. Washington, vol. 5, 
p. 227, 1903. 


Ashy fuscous, resembling myersella very closely in 
color and markings but with the transverse pale lines 
and dark areas of forewing less strongly contrasted; 
antemedial and subterminal lines faint but distinguish- 
able and otherwise as on myersella, their dark borders 
a paler brown, not so broad and more obscured than 
those on myersella. Hind wing translucent, with a very 
pale ocherous tint; termen hardly darker; cilia con- 
colorous. Alar expanse, 17-20 mm. 

Male genitalia with apical process of gnathos partially 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


fused, large, the lobes flaring and similar to those of 
myersella, the posterior projections short, blunt and 
close together as in zamacrella. Uncus narrower than 
that of either myersella or zamacrella. 

TYPE LOCALITIES: Arizona (ephestiella, in Paris Mus.) ; 
Williams, Ariz. (June; lustrella, in USNM). 

Foop: Unknown, presumably scales. 

Distripurion: Arizona. 

This species is known to me only from Dyar’s male 
type. I have no reason to question the synonymy 
which he proposed in 1905. The three species, zama- 
crella, myersella, and ephestiella, can readily be separated 
on color and genitalic characters but they are obviously 
very closely related and form a distinct subgroup within 
the genus. 

484. Laetilia fiskella Dyar 
Figures 511, 1005 


Laetilia fiskella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 221, 
1904.—McDunnough, Check list, No. 6331, 1939. 

Similar in color and markings to myersella except that 
central area of forewing (between antemedial and sub- 
terminal lines) is much darker, blackish brown; the basal 
area is paler than the median area, but the entire wing 
lacks the dusting of white so characteristic of myersella; 
antemedial white line outwardly angulate at middle; 
discal dot as in myersella, but sometimes obscured by 
the general dark suffusion. Hind wing dark smoky 
fuscous; cilia slightly paler. Alar expanse, 16-20 mm. 

Male genitalia of the Baphala type; uncus with lateral 
edges concave; apical process of gnathos consisting of a 
pair of tear-shaped lobes fused at apex; vinculum long; 
penis smooth. Female genitalia with pair of sclerotized 
plates in ductus bursae at genital opening; ventral plate 
at genital opening large; ductus bursae long and much 
more slender than in myersella; siznum a small plate 
armed with several irregularly shaped, thornlike projec- 
tions; bursa copulatrix triangulate, smooth except for a 
few minute scobinations surrounding signum. 

Typr Locatity: Tryon, N. C. (type in USNM). 

Foop: Unknown, presumably scale insects. 

Known to me only from the type series, two males 
and three females from the type locality (not one male 
and four females as stated by Dyar). The species is a 
transition one linking typical Laetilia and Baphala; it 
has Baphala genitalia but lacks the other diagnostic 
characters of that genus, the eighth abdominal segment 
of the male being simple, the male antenna without 
sinus or sex-scaling on the base of shaft, and veins 3 and 
5 of hind wing strongly stalked. 


485. Laetilia glomis (Dyar), new combination 
FieurE 512 


Euzophera glomis Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 335, 
1914. 


Clay colored, with extreme base of forewing blackish; 
antemedial line slanting outwardly to inner margin and 
with a slight notch at vein 1b, distinct and blackish; 
subterminal line faint, narrow, subdenticulate, blackish, 
bordered outwardly by 8 somewhat wider pale band; 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 235 


two faint, dark discal dots at outer angles of cell; along 
termen a row of faint blackish dots. Hind wing glossy 
smoke brown; cilia very slightly paler, nearly concolor- 
ous with wing. Alar expanse, 16 mm. 

Male genitalia with lateral edges of uncus concave 
(but not so deeply as in jiskella); apical process of 
gnathos partially fused, the lobes small, rounded, the 
posterior fused portion rather broad and forked at apex; 
vinculum short (as wide as long); penis with a few 
weakly sclerotized folds, otherwise unarmed. 

Tyrer Locauity: Taboga Isl., Panama (Feb., type in 
USNM). 

Foop: Unknown, presumably scales. 

Represented only by the unique type, a male (not 
female, as stated by Dyar). The tongue is greatly re- 
duced and less exposed by the labial palpi than that of 
melanostathma Meyrick, indicating glomis as another 
direct link between Laetilia and the coccid-feeding 
Anerastiinae. Its genitalia, however, show that glomis 
is better placed in the Phycitinae and in Laetilia than 
with the coccid-feeding complex referred to the Aner- 
astiinae. 


Baphala, new genus 


Type oF Genus: Huzophera homoeosomella Zeller. 

Tongue well developed. Antenna pubescent; in male 
some rough scaling in a very shallow sinus toward base 
of shaft. Labial palpus obliquely upturned, reaching 
to level of or slightly above vertex. Maxillary palpus 
filiform. Hind wing with veins 7 and 8 anastomosing 
for most of their length (free end of 8 very short and 
weak); 3 and 5 closely approximate or connate at lower 
angle of cell; cell less than one-half (little more than 
one-third) the length of the wing. Abdomen of male 
with a pair of ventrolateral hair tufts from eighth 
segment. 

Male genitalia with uncus narrowed at middle, the 
lateral edges deeply concave, apex rounded; apical 
process of gnathos a pair of moderately large lobes 
fusing and hooked posteriorly; aedeagus smooth; penis 
with weakly sclerotized wrinklings, otherwise unarmed; 
vinculum stout, longer than broad. 

Female genitalia with signum a small coarsely dentate 
plate; bursa copulatrix more or less finely scobinate on 
inner surface; ductus bursae with a sclerotized ventral 
plate and internal scobinations at genital opening, 
otherwise smooth; ductus seminalis from anterior end 
of bursa. 

Larvae predaceous on scale insects. 

The genus is close to but distinct from Laetilia, easily 
distinguished by its stronger tongue, the sex-scaling on 
shaft of male antenna, the narrower median area of 
uncus, the abdominal tufts on eighth segment of the male, 
and the shorter cell and the approximate (rarely connate) 
condition of veins 8 and 5of hind wing. Structural differ- 
ences separating the species are slight and chiefly in the 
size and shape of the terminal process of gnathos. 
Differences in the female signa (usually of specific 
value) are unreliable in Baphala, for they are as great 
among individuals of a given species as they are be- 


tween any two of the species themselves. The hind 
wings are similarly colored and exhibit the same sexual 
difference in the several species, whitish and semihyaline 
in the male, pale smoky fuscous in the female. 


486. Baphala basimaculatella (Ragonot), new combination 
Figures 514, 1006 


Vitula basimaculatella Ragonot, N. Amer. Phycitidae, p. 15, 
1887.—Hampson, in Ragonot, Monograph, pt. 2, p. 82, 
1901.—McDunnough, Check list, No. 6325, 1939. 

Vitula basimaculella Hulst, Phycitidae of N. Amer., p. 179, 1890; 
U.S. Nat. Mus. Bull. 52, p. 431, 1902 (misspelling). 

Laetilia eremiella Dyar, Proc. Ent. Soc. Washington, vol. 12, p. 
54, 1910.—McDunnough, Check list, No. 6329, 1939. 
(New synonymy.) 

Forewing pale whitish gray, very little darker along 
inner margin; discal dots and transverse dark markings 
blackish and strongly contrasted especially toward 
costa; the discal dots normally separated but occasion- 
ally partially fused. The palest species in the genus 
and the only one known from the United States. The 
male type in Paris lacks an abdomen but is obviously 
what Dyar later described as eremiella. It was sent to 
Ragonot by C. V. Riley and is labeled ‘‘Am. Sept.” 
In the Stainton Collection in the British Museum are 
two matching specimens from Texas. The male geni- 
talic figure was drawn from one of these. The two 
specimens in Paris from Loja, Ecuador, associated by 
Hampson with basimaculatella are not that species. I 
have examined the genitalia (male) of one of these Loja 
specimens but am unable to match them with those of 
any described species of Phycitinae known to me. 

Alar expanse, 15-19 mm. 

TYPE LOCALITIES: United States (basimaculatella, in 
Paris Mus.); Stockton, Utah (eremiella, in USNM). 

Foop: Unknown. Larvae presumably predaceous 
on coccids. 


Distrisution: Teras; Utah, Stockton (Aug.), Provo 
(July, Aug.). 


487. Baphala goyensis (Ragonot), new combination 
Fieures 513, 1009 


Zophodia goyensis Ragonot, Nouv. Gen., p. 31, 1888; Monograph, 
pt. 2, p. 25, 1901. 


Extreme base of forewing blackish fuscous; contrast- 
ingly whitish beyond to the broad blackish fuscous 
antemedial band; latter somewhat irregular, more or 
less angulate; ground color of wing between lower 
margin of cell and costa and between antemedial and 
subterminal transverse markings, whitish; remainder 
of wing shading to smoky fuscous with only a faint 
dusting of white at apex; dark border of subter- 
minal line well contrasted especially towards costa; 
midcosta contrastingly edged with blackish fuscous; a 
single rather large discal spot at end of cell (formed by 
the complete fusion of the ordinary pair of discal dots). 
Easily identified by the conspicuous discal spot and 
other blackish fuscous markings. Alar expanse 18-20 
mm. 


236 


Typr Locauity: Goya, Argentina (type in Paris 
Mus:). 

Foop: Black and wax scales (Ceroplastes and Saissetia 
spp.). 

Distrisution: Southeastern Brazil, Uruguay, Ar- 
gentina. Six specimens are before me: A male collected 
by Schaus at Castro, Parand, Brazil; another Brazilian 
male without locality label, reared from a larva feeding 
on a Ceroplastes and received from Dr. Costa Lima; and 
four females reared at the South American Parasite 
Laboratory of the U. S. Bureau of Entomology and 
Plant Quarantine, at Montevideo, Uruguay, from 
larvae feeding on black and wax scales. 


488. Baphala goyensis olivacea, new race 


Ficure 1010 


Similar in color and markings to goyensis except that 
the thorax and the dark markings on forewing are 
olivaceous rather than blackish fuscous, and the discal 
dots at end of cell are usually separated and not fused 
into a single conspicuous dark spot. Alar expanse, 
18-20 mm. 

The genitalia show no characters of specific signifi- 
cance to distinguish them from those of typical goyensis. 

Typr Locatity: Posadas, Argentina (type in USNM, 
61378; paratypes in Paris Mus. and BM). 

Foon: Scale insects (not identified). 

Described from male type and two male and six 
female paratypes from type locality, reared (Dec., Jan.) 
under No. 578-7, at the South American Parasite 
Laboratory of the U. 8. Bureau of Entomology and 
Plant Quarantine, at Montevideo, Uruguay, from larvae 
predaceous on scales; and one female paratype from 
Metaén, Argentina (Apr.). This last is from the col- 
lection of the British Museum and had been identified 
as squalida Walker. Judging from a photograph of the 
type of Walker’s species I do not think it could possibly 
be that species. Even in its somewhat faded condition 
the olivaceous color of the thorax and dark forewing 
markings are distinctly discernible. If this color had 
been present in the type of squalida, Walker or Ragonot 
would certainly have mentioned it. 


489. Baphala homoeosomella (Zeller), new combination 
Figures 517, 1011 


Euzophera homocosomella Zeller, Horae Soc. Ent. Rossicae, vol. 
16, p. 231, 1881—Hampson, in Ragonot, Monograph, pt. 2, 
p. 61, 1901. 

Vitula bodkint Dyar, Ins. Insc. Menstr., vol. 1, p. 99, 1913; Proc. 
U.S. Nat. Mus., vol. 47, p. 339, 1914 (mew synonymy). 

Vitula rusto Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 338, 1914 
(new synonymy). 

Vitula taboga Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 339, 1914 
(mew synonymy). 

Vitula saissetiae Dyar, Proc. Ent. Soc. Washington, vol. 31, p. 
16, 1929 (new synonymy). 


A pale brownish gray species, averaging smaller than 
either basimaculatella or goyensis and with the trans- 
verse dark markings on forewing weakly contrasted, 
much obscured in some specimens; discal dots when 
distinguishable small, not fused; the whitish dusting 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


limited chiefly to a shade along costa and some pale 
dusting in basal area. Alar expanse, 11-16 mm. 

There are no male genitalic structural characters to 
separate any of Dyar’s supposed species from typical 
homoeosomella and only trifling and inconsistent color 
differences to distinguish them from each other. The 
females show considerable variation in their signa, but 
as much or more among specimens of any one of the 
forms as between typical females of the forms them- 
selves. The greatest variation is shown among speci- 
mens from a reared series of typical saissetiae (figs. 
1011b-4). 

Tyre LocaLities: Honda, Colombia (homoeosomella, 
in BM); Georgetown, British Guiana (bodkini, in 
USNM); Parafso (rusto, in USNM), Taboga Isl. (taboga, 
in USNM), Barro Colorado Isl. (saissetiae, in USNM), 
all in Canal Zone, Panamé. 

Foon: Saissetia spp. and Ceroplastes spp. (wax and 
black scales), Toumayella sp. 

Distrizution: Cupa. VireGin Istanps: St. Croix 
(July). PanamA Canat Zone: Barro Colorado Isl. 
(May), Paraiso (May), Taboga Isl. (Feb.), Rio Trinidad 


(May). Brivise Guiana: Georgetown (June). Co- 
LomBIA: Honda (Apr., Aug., Sept.). Brazin: Sado 
Paulo (Feb.). 


The most widely distributed and apparently the 
commonest phycitine predaceous on scales in tropical 
America. 

490. Baphala haywardi, new species 
Fieures 516, 1007 


In color and markings similar to the more strongly 
marked color forms of homoeosomella, the female hardly 
distinguishable from the female paratype of saissetiae; 
basal area of forewing, midcostal area above cell, and 
terminal area beyond subterminal line rather strongly 
dusted with whitish scales; dark transverse antemedial 
band pale grayish brown, outwardly angulate and 
diffusing into a similar shade through most of area be- 
tween inner margin and cell; smuate subterminal white 
line bordered inwardly and outwardly by pale grayish 
brown; discal dots at.end of cell similarly colored; a row 
of somewhat darker dots along termen. Alar expanse, 
15-16 mm. 

Male genitalia with vinculum shorter and apical 
process of gnathos stouter than those of homocosomella 
or any of the other described species of the genus. 
Female genitalia with fine scobinations of bursa dis- 
tributed over most of inner surface; sclerotization of 
ductus bursae at genital opening weaker than that of 
homoeosomella (compare figs. 1011h and 1007). 

Typr Locauity: Concordia, Entre Rios, Argentina 
(type in USNM, 61879). 

Foon: Ceroplastes grandis. 

Described from male type and female paratype from 
type locality, reared by K. J. Hayward from larvae 
feeding on the large wax scale (co, ‘9—-4-1935,” 9, 
“14-VIT-1935,” Hayward No. 3185). They were sub- 
mitted as examples of homoeosomella, which they could 
easily be on color and markings; but their genitalia, 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


especially those of the male, seem to rule them out 
from that species. 


491. Baphala glabrella (Dyar), new combination 
Fiaurss 515, 1008 


Euzophera glabrella Dyar, Ins. Insc. Menstr., vol. 7, p. 57, 1919. 


A suffused, grayish brown species with markings ob- 
scure, the pale transverse antemedial and subterminal 
lines and the dark discal dots faintly indicated; general 
color darker (more brownish) than that of homoeo- 
somella. Probably only a variety of that species, but 
a reared series would be needed to determine this. The 
genitalia show some trifling differences from homoco- 
somella, but none that could be classed as specific. 

Alar expanse, 15-16 mm. 

Type Locauity: Cayuga, Guatemala (Apr., May, 
June; type in USNM). 

Foop: Unknown. 

The species is known only from the original series of 
five specimens upon which the name was based. 


492. Baphala squalida (Walker), new combination 


Nephopteryz squalida Walker, List, pt. 27, p. 59, 1863. 
Zophodia squalida (Walker) Ragonot, Monograph, pt. 2, p. 25, 
1901. 


I know this species only from photographs of the male 
type and its genitalia submitted by Tams. The photo- 
graph of the moth shows a specimen almost completely 
suffused with blackish fuscous except for a small area 
from outer part of cell to costa. The subterminal pale 
line of forewing, the two discal dots, and the row of 
terminal dots are distinct, but the antemedial line and 
the usually contrasted dark transverse markings are 
indistinguishable in the general dark suffusion. The 
hind wing shows the veins strongly outlined by dark 
scaling and the apical area of the wing darkly clouded 
(unusual markings for a male of this genus). Alar ex- 
panse, 20 mm. 

Typr Locauity: Rio de Janeiro, Brazil (type in BM). 

Foop: Unknown. 

Certain specific application of Walker’s name will 
have to wait upon recovery of more material in reared 
series from the type locality. If, as I suspect, a series of 
rather dark homoeosomelia in the United States National 
Museum from Sao Paulo (four males and eight females 
reared from larvae on wax scales) should prove to be 
squalida, Walker’s name will take precedence and 
homoeosomella will fall as a synonym. However, none 
of our Brazilian specimens is over 16 mm. in expanse 
and their small size seems to rule them out. 


133. Rhagea, new genus 


Typ or GeNus: Zophodia packardella Ragonot. 

Tongue well developed. Antenna pubescent, shaft 
simple in both sexes. Labial palpus porrect. Maxil- 
lary palpus filiform. Hind wing with veins 7 and 8 
anastomosing for about half their distance beyond cell; 
3 and 5 stalked. Eighth abdominal segment of male 
simple. 


237 


Male genitalia with apex of uncus broad, apical 
process of gnathos bifid, large; aedeagus moderately 
long and stout; penis with some very weakly sclerotized 
scobinations or pubescence but without cornutus. 

Female genitalia without signum; bursa copulatrix 
smooth or (stigmella) minutely scobinate-granulate; 
ductus bursae with a weak bandlike sclerotized ventral 
plate or scobinate patch on inner surface at genital 
opening, otherwise membranous; ductus seminalis from 
anterior end of bursa. 

Larvae feeding in roots of Crassulaceae or flowers of 
Orobanchaceae. 

This genus is close to Zophodia Hiibner. It differs in 
having porrect labial palpi in both sexes, the eighth seg- 
ment of male simple, the antennal shaft of the male 
simple and the female bursa without trace of signum. 


493. Rhagea packardella (Ragonot), new combination 
Ficures 518, 1012 


Zophodia packardella Ragonot, N. Amer. Phycitidae, p. 12, 
1887.—Hulst, Phycitidae of N. Amer., p. 173, 1890; U. 8. 
Nat. Mus. Bull. 52, p. 430, 1903—Hampson, in Ragonot, 
Monograph, pt. 2, p. 22, 1901.—McDunnough, Check list, 
No. 6305, 1939. 

Zophodia orobanchella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 111, 1904—-McDunnough, Check list, No. 6307, 1939. 
(New synonymy.) 

Palpi, head, thorax, and forewings white dusted with 
brown, giving the moth a pale ash gray appearance. 
Forewing with costal area paler than remainder of 
wing; no trace of any transverse antemedial line; dark 
dustings outlining most of the veins and forming rather 
conspicuous longitudinal lines through center of cell, on 
basal third of lower vein of cell, and along the basal 
third of the fold; a large conspicuous discal spot at end 
of cell covering the entire crossvein between veins 3 
and 8; subterminal line indicated by dark shading which 
extends from costa near apex to about vein 2, indented 
between veins 4 and 6 and distinct only towards costa; 
terminal margin dark; cilia ash gray. Hind wing pale 
smoky white, with veins and terminal area slightly 
darker and a fine dark line along termen; cilia sordid 
white with slightly darker basal band. Alar expanse 
18.5-25 mm. 

Male genitalia with vinculum shorter and divided 
apical process of gnathos larger in proportion to re- 
mainder of genitalia than those of stigmella. Female 
genitalia with inner surface of ductus bursae finely 
scobinate-granulate at genital opening; bursa copula- 
trix minutely and sparsely scobinate. 

Typr tocauities: California (packardella, in Paris 
Mus.); Wawawai, Wash. (orobanchella, in USNM). 

Foop piant: Orobanche ludoviciana Nutall (larvae 
feeding in the flowers). 

Distrisution: Washington, Wawawai (Aug.); Utah, 
Provo (July), Eureka (July); California, Dixieland 
(Apr.); Arizona, Phoenix (Mar.); Colorado, Fort Collins 
(Sept.). 

Dyar had his specimens of orobanchella under pack- 
ardella in the National Collection but, to the best of my 
knowledge, the synonymy had not been published. 


238 


The species is easily recognized by the large, dark, 
discal spot on the forewing, the most conspicuous mark- 
ing on the insect. 


494. Rhagea stigmella (Dyar), new combination 
Ficures 519, 520, 521, 1013, 1014 

Zophodia stigmella Dyar, Pomona Coll. Journ. Ent., vol. 2, No. 4, 
p. 378, 1910.—Barnes and McDunnough, Check list of the 
Lepidoptera of Boreal America, No. 5716, 1917. 

Yosemitia maculicula Dyar, Ins. Insc. Menstr., vol. 1, p. 34, 
1918.—Barnes and McDunnough, Contributions, vol. 3, 
p. 200, 1916. 

Zophodia stigmella maculicula (Dyar), Barnes and McDunnough, 
Check list of the Lepidoptera of Boreal America, No. 5716, 
1917. 

Eumysia stigmella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221, 
1925.—McDunnough, Check list, No. 6300, 1939. 

Eumysia maculiella Dyar, Ins. Insc. Menstr., vol. 13, p. 221, 
1925 (misspelling for maculicula). 

Eumysia stigmella maculicula (Dyar), McDunnough, Check list, 
No. 6300, 1939. 

Palpi, face, and head pale brown dusted with white, 
the scales brown with their tips white. Thorax grayish 
white shading to pale argillaceous. Forewing grayish 
(ashy) white with blackish brown markings and a 
rather broad pale argillaceous shading along the fold 
and fainter traces of the same color between the veins 
beyond cell; veins, except on lower half of wing, faintly 
lined with blackish brown; antemedial line incomplete, 
acutely angled, in most specimens indicated only by a 
broad blackish spot on the basal third of inner margin 
extending from inner margin to vein 1b, a rather con- 
spicuous blackish patch on lower vein of cell just beyond 
middle and an obscure blackish shade on basal third of 
costa; subterminal line often obsolete, otherwise indi- 
cated by a narrow blackish streak from costa to vein 8, 
a more or less broken line between veins 5 and 2, and a 
small blackish patch or spot on vein 1b; rarely are the 
transverse markings strongly indicated; a more or less 
pronounced blackish discal spot at lower outer angle of 
cell and a few obscure blackish spots along termen 
between the vein ends; cilia argillaceous with the scale 
tips white. Hind wing semihyaline white with a 
smoky shade toward apex, a fine dark line along termen, 
and the veins faintly outlined by smoky scaling; cilia 
white with a very faint ocherous basal shade. Alar 
expanse, 21-30 mm. 

Male genitalia with vinculum appreciably longer and 
apical process of gnathos proportionately smaller than 
those of packardella. The type of stigmella (fig. 519) is 
abnormal in that the apical process of gnathos is par- 
tially fused. The normal bifid condition isshownin figures 
520 and 521. Female genitalia with a weak, bandlike, 
sclerotized ventral plate on imner surface of ductus 
bursae at genital opening; bursa copulatrix smooth. 

Larva unusual for the family in that the sclerotized 
rings about setal tubercles IIb on mesothorax and III 
of eighth abdominal segment are absent. In this 
respect it is similar to the caterpillar of Hiiella zincken- 
ella which it otherwise resembles in superficial appear- 
ance. 

TYPE LOCALITIES: San Diego, Calif. (stigmella, in 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


USNM); La Puerta, Calif. (maculicula, in USNM). 

Foop puant: (Sedum) Hasseanthus elongatus (larvae 
boring in the roots; probably also in the roots of other 
succulents). 

Distrizution: Unitep Srares: California, San 
Diego (Mar., May, June, July, Aug., Oct.), Loma Linda 
(June), La Puerta (July), Laguna (May), Half Moon 
Bay (Aug.). México: Baja California (June). 

This is a variable species in size, markings, and struc- 
ture. Extremes of difference in genitalia are shown in 
the figures. Barnes and McDunnough (1916) pointed 
out that maculicula was only a color variety of stigmella 
and in their Check List reduced the name to subspecific 
rank. JI do not believe that it deserves even this status 
for it represents only one of many color variants. In 
some specimens the transverse markings on forewing 
are nearly obsolete, in others more or less indicated but 
incomplete, and in one specimen before me (Half Moon 
Bay, Calif., Aug. 11, 1937, W. H. Lange No. 27) the 
discal spot and transverse lines are conspicuous and 
the latter complete except at the fold, the subterminal 
line running from outer fourth of costa transversely to 
outer third of inner margin, broken between veins 2 and 
1b and with a sharp outward angulation near yein 5. 
The genitalic differences shown in our figures at first 
glance seem rather striking; but they are not consistent 
and represent only individual variation. 

This species seems to be native to southern California 
and northwestern México. We have a series of reared 
specimens from Laguna, Calif. A few larvae and pupae 
have been intercepted at quarantine ports in California 
from roots of succulents (‘“Sedum sp.’’) shipped from 
México and from another similar interception at San 
Francisco a large (85 mm.) male was reared. 


134. Genus Zophodia Hiibner 


Zophodia Hiibner, Verzeichniss bekannter Schmettfer]linge, p. 
370 [1825].—Ragonot, Ent. Monthly Mag., vol. 22, p. 19, 
1885.—Hulst, Phycitidae of N. Amer., p. 172, 1890.— 
Hampson, in Ragonot, Monograph, pt. 2, p. 18, 1901.— 
Spuler, Die Schmetterlinge Europas, vol. 2, p. 207, 1910.— 
Dyar, Ins. Insc. Menstr., vol. 13, p. 220, 1925.—Heinrich, 
Proc. U. 8. Nat. Mus., vol. 86, p. 401, 1939.—Janse, Journ. 
Ent. Soc. South Africa, vol. 8, p. 39, 1945. (Type of genus: 
Tinea convolutella Hubner.) 

Dakruma Grote, Bull. U. 8. Geol. Geogr. Surv. Terr., vol. 4, p. 
702, 1878. (Type of genus: Dakruma turbatella Grote.) 


Tongue well developed. Antenna of male pubescent 
and with a series of modified, papillalike setae on the 
inner sides of several basal segments of the shaft; of 
female simple and very shortly pubescent. Labial 
palpus oblique in the male, porrect in the female. 
Maxillary palpus filiform. Hind wing with veins 7 and 
8 anastomosing beyond the cell; 3 and 5 connate (in 
occasional specimens very shortly stalked). Highth 
abdominal segment with a pair of weak ventrolateral 
hair tufts. 

Male genitalia with apical process of gnathos bifid, 
large; apex of harpe evenly rounded; vinculum long; 
anellus with base of plate narrowly sclerotized, arms 
moderately long, slender, slightly twisted; aedeagus 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 239 


moderately long and stout; penis partially ribbed and 
pubescent. 


Female genitalia with a small weak signum developed 
as a plate with an inwardly projecting flange; bursa 
copulatrix small, minutely and very weakly scobinate; 
ductus bursae minutely scobinate, with two rather large, 
sclerotized, dorsal plates at genital opening; ductus 
seminalis from bursa near signum. 


As here defined the genus is limited to its type species. 
Its distribution is central and southern Europe, the 
northern part of the United States and southern Canada. 


495. Zophodia convolutella (Hiibner) 
Ficures 522, 1015 


Tinea convolutella Hiibner, Sammlung europiischer Schmetter- 
linge, Lepidoptera 8, Tineae 2, pl. 5, fig. 34, 1796. 

Tinea grossulariella Hiibner, Geschichte europaischer Schmetter- 
linge, Tinea II, pl. C.a.b., fig. 2.a.b.c., [1807-1809] (larva). 

Phycis grossulariella (Hiibner) Zincken, in German and Zincken, 
Mag. der Ent., vol. 3, p. 144, 1818.—Treitschke, Die Schmet- 
terlinge von Europa, vol. 9, pt. 1, p. 172, 1832; op. cit., vol. 
10, pt. 3, p. 275, 1835.—Duponchel, Histoire naturelle des 
Lépidoptéres, ou papillons de France, vol. 10, p. 206, pl. 
279, fig. 9, 1836. 

Zophodia grossularialis Hiibner, Verzeichniss bekannter Schmet- 
tferllinge p. 370, [1825] (emended spelling for grossulariella 
and to replace convolutella). 

Zophodia convolutella (Hiibner), Verzeichniss bekannter Schmet- 
tferllinge (sic), p. 370, [1825]—Heinemann, Schmetterlinge 
Deutschlands und der Schweiz, Abt. 2, vol. 1, no. 2, p. 190, 
1865.—Ragonot, Ent. Monthly Mag., vol. 22, p. 19, 1865.— 
Hampson, in Ragonot, Monograph, pt. 2, p. 20, 1901.— 
Staudinger and Rebel, Catalog der Lepidopteren des palae- 
arctischen Faunengebietes, vol. 2, p. 25, 1901.—Spuler, 
Die Schmetterlinge Europas, vol. 2, p. 207, 1910.—Heinrich, 
Proc. U. 8. Nat. Mus., vol. 86, p. 402, 1939. 

Myelois (Zophodia) convolutella (Hiibner) Zeller, Isis von Oken, 
1839, p. 178; 1848, p. 679. 

Homoeosoma convolutella (Hiibner) Herrich-Schaffer, Systema- 
tische Bearbeitung der Schmetterlinge von Europa, vol. 4, 
p. 107, 1849. 

Pempelia grossulariae Riley, First annual report on the noxious, 
beneficial and other insects of the State of Missouri, p. 140, 
1869; Papilio, vol. 1, p. 108, 1881 (suggests synonymy with 
convolutella).—Packard, Guide to the study of insects, p. 
331, 1869. 

Dakruma turbatella Grote, Bull. U. 8. Geol. Geogr. Surv. Terr. 
vol. 4, pp. 702, 703, 1878; North Amer. Ent., vol. 1, p. 11, 
1879. 

Myelots convolutella (Hiibner) Packard, Guide to the study of 
insects, ed. 7, p. 331, 1880. 

Dakruma grossulariae (Riley) Grote, North Amer. Ent., vol. 1, 
p. 68, 1880. 

Dakruma convolutella (Hiibner) Grote, New check list of North 
American moths, p. 55, 1882 (gives grossulariae and turba- 
tella as synonyms). 

Zophodia grossulariae (Riley) Hulst, Phycitidae of N. Amer., p. 
173, 1880; U.S. Nat. Mus. Bull. 52, p. 429, 1903.—Hampson, 
in Ragonot, Monograph, pt. 2, p. 21, 1$01.—Dyar, Proc. 
Ent. Soc. Washington, vol. 7, p. 37, 1905; Ins. Insc. Menstr., 
vol. 13, p. 221, 1925.—Pack, Utah Agr. Exp. Station. Bull. 
216, pp. 1-12, 1930.—McDunnough, Check list, No. 6303, 
1939. 

Euzophera franconiella Hulst, Phycitidae of N. Amer., p. 177, 
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 61, 1901. 

Zophodia bella Hulst, Canadian Ent., vol. 24, p. 61, 1892.—Dyar, 
Proc. Ent. Soc. Washington, vol. 6, p. 228, 1904. 

Zophodia franconiella (Hulst) Barnes and MecDunnough, Check 
list of the Lepidoptera of Boreal America, No. 5706, 1917. 


Zophodia grossulariae franconiella (Hulst) Dyar, Ins. Insc. 
Menstr., vol. 13, p. 221, 1925. 

Zophodia grossulariae thouna Dyar, Ins. Insc. Menstr., vol. 13, 
p. 221, 1925. 

Zophodia grossulariae dilativitta Dyar, Ins. Insc. Menstr., vol. 
13, p. 222, 1925. 

Zophodia grossulariae magnificans Dyar, Ins. Inse. Menstr., vol. 
13, p. 222, 1925. 

Forewing fuscous dusted with white, the white color 
strongest in central costal area, the general color gray; 
antemedial line outwardly oblique to lower vein of cell 
and notched between cell and inner margin, white, 
bordered outwardly by a more or less extended black 
shade; subterminal line oblique, slightly dentate and 
sinuate, white, bordered inwardly by a black line and 
outwardly by a narrow black line for a short distance 
from costa; the fold and veins at extreme base of wing 
and in area beyond subterminal line faintly outlined in 
black; discal mark at end of cell black, curved, rarely 
replaced by a pair of dots; a row of black dots along 
termen between the vein ends. Hind wing pale smoky 
white with a narrow dark line along termen. Alar 
expanse, 25-36 mm. 

Male genitalia with lateral margins of vinculum 
broadly and shallowly excavate, its posterior margins 
straight. Female genitalia with signum small and 
weak. The eighth-segment collar is subject to con- 
siderable variation in the size and shape of the unsclero- 
tized dorsal area; but these variations do not conform to 
the varieties that have been named and can be found 
in any series from one locality. 


TYPE LocaLities: Germany (convolutella and grossu- 
lariella, no known existing types); Missouri (grossu- 
lariae, no known existing type); Oldtown, Maine 
(turbatella, in BM); Franconia, N. H. (franconiella, in 
AMNH, ex Rutgers); Massachusetts (bella, in AMNH, 
ex Rutgers); southern Utah (ihowna, in USNM); San 
Diego, Calif. (dilativitta, in USNM) Seattle, Wash. 
(magnificans, in USNM). 

Foon piants: Ribes grossularia Linnaeus and other 
Ribes species (larva feeding in the fruit). 

Distrisution: Evurorr (central and southern). 
Unitep States: Maine, Orono; New Hampshire, 
Hampton (May), Durham; Missouri; Colorado, Mani- 
tou, Denver (Apr.), Fort Collins (Mar., Apr.); Utah, 
Logan (‘June’), Beaver Canyon (‘vii’); Oregon; 
California, San Diego; Washington, Seattle, Bellingham 
(Apr.). Canapa: Quebec, St. Johns County (Apr.), 
Mount St. Hilaire (May); Ontario, Hymers; Alberta, 
Edmonton (May), Bilby (May); British Columbia, 
Kaslo (Apr.), Wellington (Apr.), Alberni (May), 
Goldstream (May), Vancouver Isl. (Apr.). 

The foregoing localities are for the specimens before 
me. The species is generally distributed over the 
northern part of the United States and southern 
Canada. 

The species is more extensively treated in my paper 
on the cactus-feeding Phycitinae (Proc. U.S. Nat. Mus., 
vol. 86, pp. 402-405, 1939). In this country it is popu- 
larly known as the ‘gooseberry fruit-worm.” It has a 
rather extensive economic literature and is the most im- 


240 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


portant lepidopterous pest of the gooseberry here and 
abroad, often doing serious injury. It is also recorded 
as an occasional enemy of currants. 

There is one generation a year, the moths flying from 
mid-April to early June. About 10 months are passed 
in the pupal stage, the insects overwintering as pupae in 
loose cocoons on the ground under fallen leaves and 
other rubbish. 


135. Genus Melitara Walker 


Melitara Walker, List, pt. 27, p. 136, 1863.—Hulst, Phycitidae of 
N. Amer., p. 171, 1890.—Ragonot, Monograph, pt. 2, p. 12, 
1901.—Dyar, Proc. Ent. Soc. Washington, vol. 30, p. 133, 
1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 338, 
1939.—Dodd, Biological campaign against prickly-pear, 
Brisbane, Australia, pp. 38, 39, 58, 1940. (Type of genus: 
Melitara prodenialis Walker.) 

Megaphycis Grote, Canadian Ent., vol. 14, p. 30, 1882. 
of genus: Zophodia bollii Zeller.) 

Tongue developed (stout, but rather short). Antenna 
of male bipectinate, of female shortly bipectinate. La- 
bial palpus porrect. Maxillary palpus squamous. Hind 
wing with veins 7 and 8 approximate beyond cell; 3 and 
5 connate (rarely very shortly stalked). Eighth ab- 
dominal segment of male simple. 

Male genitalia with apical process of gnathos bifid; 
harpe with apex evenly rounded; vinculum short; anel- 
lus with base of plate narrowly sclerotized, arms mod- 
erately long and stout; aedeagus stout, moderately long. 

Female genitalia without signum, the latter replaced 
by a few very fine scobinations (not distinguishable in 
most preparations except under high magnification), 
bursa copulatrix otherwise simple; ductus seminalis from 
bursa near junction of ductus bursae and bursa copula- 
trix. 


Larva bluish, not banded; sclerotized plates sur- 
rounding body setae rather small; 3 setae in group VII 
on abdominal segments 7 and 8. 

The larvae feed gregariously in the joints of various 
species of Platypuntia. 

Eggs laid in chains (‘egg sticks”). 

Melitara and the 16 genera following (Olycella to Cac- 
tobrosis) form what appears to be a natural group of 
genera and species more closely related to each other 
than to other Phycitinae, although they also show dis- 
tinct affinities to Zophodia and the coccid feeders of the 
Laetilia complex. They all have a common host asso- 
ciation, their larvae being internal feeders in the fruits 
and stems of various cacti. A fuller treatment of the 
several species will be found in my paper on the cactus- 
feeding Phycitinae (Proc. U. S. Nat. Mus., vol. 86, 
pp. 331-413, 1939). 

The genus Melitara as here defined is separated from 
the other cactus-feeding Phycitinae by the following 
combination of characters: Antennae bipectinate in 
both sexes; labial palpi porrect in both sexes; veins 7 and 
8 of hind wing approximate; veins 3 and 5 of hind wing 
connate; eighth abdomial segment of male simple; 
larvae not banded or conspicuously spotted, gregarious 
in habit throughout feeding period. 


(Type 


Two species are recognized as belonging to the genus, 
and its distribution is apparently limited to the United 
States and adjacent areas in northern México. 


496. Melitara prodenialis Walker 
Fieures 74, 527, 1016 

Melitara prodenialis Walker, List, vol. 27, p. 137, 1863.—Hulst, 
Phycitidae of N. Amer., vol. 17, p. 171, 1890; U. S. Nat, 
Mus. Bull. 52, p. 429, 1903.—Ragonot, Monograph, pt. 2, 
p- 13, 1901.—Hunter, Pratt, and Mitchell, U. S. Dep. Agr. 
Bur. Ent. Bull. 113, p. 28, 1912.—Dodd, Council for Sci. 
and Ind. Res., Australia, Bull. 34. p. 27, 1927; Biological 
campaign against prickly-pear, Brisbane, Australia, p. 77, 
1940.—Dyar, Proc. Ent. Soc. Washington, vol. 30, p. 133, 
1928.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 339, 
1939.—MeDunnough, Check list, No. 6277, 1939. 

Zophodia bollit Zeller, Verh. zool.-bot. Ges. Wien, p. 550, pl. 3, 
fig. 21, 1872. 

Megaphycis bollit (Zeller) Grote, Canadian Ent., vol. 14, p. 30, 
1882. 


Melitara prodenialis bollii (Zeller) Dyar, Proc. Ent. Soc. Wash- 
ington, vol. 30, p. 133, 1928. 

Palpi, head, and thorax cinereous fuscous sparsely 
dusted with white, especially on basal segments of Jabial 
palpi; posterior margin of thorax blackish. Forewing 
cinereous fuscous with a heavy dusting of white on costal 
half; the fuscous and whitish areas contrasted but not 
sharply defined, the white dusting most pronounced 
between antemedial and subterminal lines and in 
subapical area beyond subterminal line; a few black 
scales scattered over entire wing; antemedial line 
narrow, black, outwardly angled from basal fourth of 
costal margin to 1b, less sharply defined from there to 
inner margin; subterminal line narrow, black out- 
wardly margined by a narrow border of white, beyond 
which is a faintly dark shading which forms an obscure 
line paralleling the subterminal line, the parallel black 
lines most pronounced from costal margin near apex to 
vein 6; subterminal line irregularly dentate and smuate, 
the angulations rather shallow; discal dots fused, form- 
ing a black line or smudge along discocellular vein; a 
row of black dots along termen at the vein ends; cilia 
grayish fuscous; underside of wing grayish fuscous, in 
some specimens with a more ocherous tint. Hind wing 
white, semihyaline with more or less fuscous suffusion 
at apex and along costal and terminal margins, the 
fuscous shading more extended in female than male; 
cilia white with a narrow, dark subbasal line. Alar 
expanse, 31-45 mm. 

Male genitalia with outer margins of vinculum evenly 
curved; elements of transtilla rather broad. Female 
genitalia with ductus bursae rather slender for most of 
its length. 

Type Locatitizs: ‘United States” (prodenialis, in 
BM); Texas (bollvi, Cambridge Mus. Nat. Hist.). 

Foop piants: Opuntia (Platypuntia) spp. 

Distrisution: Teras, Dallas, Freeport, Utopia, Col- 
lege Station (Oct.), Brownsville; Mississippi, Biloxi 
(Sept.); Oklahoma, Wichita National Forest (June); 
Florida, Altamont (Sept., Oct.), Key West, Lakeland 
(Apr.), Crescent City (May), Miami (Oct.), St. Peters- 
burg (Mar., June, Sept., Oct.), Fort Meade (Apr.), Fort 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Myers (Apr.), Venice (May); North Carolina, Southern 
Pines (June); Delaware, Indian River Bay (July); New 
Jersey, Lakehurst (Sept.); New York, Rye (July). 


497. Melitara dentata (Grote) 
Fieures 528, 1017 


Zophodia dentata Grote, Canadian Ent., vol. 8, p. 158, 1876; 
Bull. U. S. Geol. Geogr. Surv. Terr., vol. 3, p. 799, 1877. 
Megaphycis dentata (Grote), Canadian Ent., vol. 14, p. 30, 1882. 
Melitara deniaia (Grote) Hulst, Phycitidae of N. Amer., p. 172, 
1890; U. S. Nat. Mus. Bull. 52, p. 429, 1903.—Kellogg, 
Kansas Univ. Quart., vol. 1, p. 39, 1892.—Ragonot, Mono- 
graph, pt. 2, p. 14, 1901.—Hunter, Pratt, and Mitchell, 
U. S. Dep. Agr. Bur. Ent. Bull. 113, p. 28, 1912.—Dodd, 
Council for Sci. and Ind. Res., Australia, Bull. 34, p. 29, 
1927; Biological campaign against prickly-pear, Brisbane, 
Australia, p. 79, 1940.—Dyar, Proc. Ent. Soc. Washington, 
vol. 30, p. 134, 1928.—Heinrich, Proc. U. 8. Nat. Mus. vol. 
86, p. 341, 1989.—McDunnough, Check list No. 6278, 1939. 

Melitara doddalis Dyar, Ins. Insc. Menstr., vol. 13, p. 13, 1925; 
Proc. Ent. Soc. Washington, vol. 30, p. 184, 1928.—Dodd, 
Council for Sci. and Ind. Res., Australia, Bull. 34, p. 29, 
1927; Biological campaign against prickly-pear, Brisbane, 
Australia, p. 80, 1940. 

Melitara junctolineella Hulst (in part), Canadian Ent., vol. 32, 
p. 173, 1900.—Barnes and McDunnough, Contributions, 
vol. 3, p. 199, 1916. 

Melitara bollit Dodd (not Zeller), Council for Sci. and Ind. Res., 
Australia, Bull. 34, p. 29, 1927. 

General color and pattern as in prodenialis except: 
Blackish shading on posterior margin of thorax less pro- 
nounced and in some specimens not distinguishable. 
Forewing with white dusting rather evenly distributed 
over the entire wing, the whitish and fuscous areas not 
contrasted except (in some specimens) for a rather 
narrow pale suffusion along costal margin and a more 
or less pronounced dark shade from end of cell to middle 
of inner margin; the subterminal line with no black 
shading beyond its white border except for a short dis- 
tance from apex, markedly dentate and sinuate, the 
angulations deep, the angulation between veins 5 and 
6 reaching to the cell. Hind wing semihyaline, in the 
male almost pure white with little or no fuscous shading, 
the latter, when present, confined to a narrow band 
along costal margin and a fine line along termen; in the 
female fuscous shadings nearly always pronounced, 
though limited to a narrow line along costal margin, a 
slight clouding at apex, and a thin line along termen; 
cilia white. Alar expanse, 33-50 mm. 

Male genitalia with outer margins of vinculum slight- 
ly sinuate; elements of transtilla slightly narrower than 
those of prodenialis. Slight differences in the shape of 
the anellus between the two species shown in figures 
527e-f and 528b. Female genitalia essentially like 
those of prodenialis except that ductus bursae is nor- 
mally stouter. 

Typr Locaities: Clear Creek Canyon, Colo. (den- 
tata, in BM); Mesilla Park, N. Mex. (doddalis, in 
USNM). 

Foop puants: Opuntia (Platypuntia) spp. 

DistrisuTion: Unirep States: Wyoming, Evanston 
(July), Medicine Bow (July); Colorado, Glenwood 
Springs (July, Aug., Sept.), Fort Collins, Denver, Rocky 


241 


Ford (Sept.); Utah, Buckskin Valley (Iron County, 
June, July), Eureka (Aug.), Dividend (Aug., Sept.); 
Kansas, Manhattan (Sept.); Arizona, Mormon Lake 
(July), Douglas (Aug., Sept.), Oracle (Sept.), Globe 
(Sept.), Quijotoa (Oct.), Chiricahua Mts.; New Mexico, 
Mesilla Park (Sept.), Silver City (Sept.), Jemez Springs 
(July, Aug., Sept.); Texas, Uvalde (Sept., Oct., Nov.), 
Henrietta (Oct.), Trent (Oct.), Rock Springs, Laredo 
(Sept.), Shafter (Sept.), Albany, Panhandle (Aug.). 
México: Chihuahua (city), Julimes (Sept., Oct.), Mo- 
relia, (Oct.). 

Shortly after the publication of my 1939 paper on 
the cactus-feeding phycitids (in which I discussed the 
synonymy of dentata and doddalis) we received through 
Mr. Leith Hitchcock a series of moths from Uvalde, 
Tex., labeled Melitara bollii. They were evidently the 
specimens that Dodd distinguished from prodenialis in 
treating bolli as a distinct species. Dyar was respon- 
sible for the original identification of bollii with pro- 
denialis as a race or variety. He apparently was con- 
fused by Dodd’s biological notes on the Uvalde moths. 
Some of these moths he placed with other Texas speci- 
mens of true bollii, while others from the same Uvalde 
rearing he included under his doddalis. M. bollit 
Zeller is a synonym of prodenialis while bollii Dodd (not 
Zeller) is merely a southern Texas form of the exceed- 
ingly variable dentata Grote. 


136. Genus Olycella Dyar 


Olycella Dyar, Proc. Ent. Soc. Washington, vol. 30, p. 134, 
1928.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 348, 
1939.—Janse, Journ. Ent. Soc. South Africa, vol. 8, p. 40, 
1945. (Type of genus: Melitara junctolineella Hulst.) 

Tongue developed (as in Melitara). Antenna of male 
bipectinate, of female shortly bipectinate. Labial pal- 
pus obliquely ascending (sometimes in the female the 
third segment is bent forward, which gives the palpus a 
porrect appearance, but the second segment is always 
deflected upward and reaches nearly as high as the top 
of the head). Maxillary palpus squamous. Hind wing 
with veins 7 and 8 approximate beyond the cell; 3 and 
5 connate. Eighth abdominal segment of male simple, 

Male genitalia with apical process of gnathos bifid; 
harpe with apex evenly rounded; vinculum short; 
anellus with base of plate narrowly sclerotized, arms 
moderately long and stout; aedeagus stout, moderately 
long. 

Female genitalia with signum, the latter a small 
ridged plate; bursa copulatrix wrinkled, otherwise 
simple and without scobinations; ductus bursae with a 
pair of sclerotized plates on inner wall at genital open- 
ing; ductus seminalis from center of bursa. 

Larva white with broad blackish or purplish cross 
bands on the caudal margins of the segments; sclero- 
tized plates surrounding setae rather small; three setae 
in group VII on abdominal segments 7 and 8. 

The larvae feed gregariously for a short period after 
hatching (probably during the first instar) but there- 
after are solitary in habit. They feed in the joints of 
various Platypuntias. 


242 


The genus is close to Melitara, distinguished from it 
only by the following characters: Labial palpi obliquely 
ascending; larvae transversely banded and solitary in 
habit during most of the feeding period. 

Three species and one local race are here recognized. 
They are remarkably alike in structure, the genitalia 
exhibiting difference of only an individual character. 
The species, however, can be distinguished by color 
differences which seem to be constant. 


aa 498. Olycella junctolineella (Hulst) 
Fieures 75, 529, 1018 


Melitara junctolineella Hulst, Canadian Ent., vol. 32, p. 173, 1900; 
U.S. Nat. Mus. Bull. 52, p. 429, 1803.—Hunter, Pratt, and 
Mitchell, U. S. Dep. Agr. Bur. Ent. Bull. 113, p. 25, 1912— 
Barnes and McDunnough, Contributions, vol. 3, p. 199, 
1916.—Dodd, Council for Sci. and Ind. Res., Australia, 
Bull. 34, p. 27, 1927. 

Olyca junctolineella (Hulst) Barnes and McDunnough, Check list 
of the Lepidoptera of Boreal America, No. 5695, 1917. 
Olycella junctolineella (Hulst) Dyar, Proc. Ent. Soc. Washington, 
vol. 30, p. 134, 1928.—Heinrich, Proc. U. 8. Nat. Mus., 
vol. 86, p. 344, 1939.—MeDunnough, Check list, No. 6281, 
1939.—Dodd, Biological campaign against prickly-pear, 

Brisbane, Australia, pp. 39, 59, 81, 1940. 

Head, thorax, and forewing ocherous fuscous dusted 
with white and marked with patches and lines of black 
scales. Labial palpus with the apical ends of the seg- 
ments blackish. Maxillary palpus crossbanded with 
black scales. Thorax with some black dusting on 
posterior margin: Forewing with whitish dusting 
slightly mtensified in costal area; veins faintly outlined 
in black; a row of more or less obscure black dots on 
termen between the vein ends; antemedial line inter- 
rupted, sometimes obscure, in fresh, well-marked speci- 
mens its outer dentation much extended and meeting a 
shade from the inner angulation of the subterminal line 
at the fold (which indicates the normal position of the 
absent vein 1c); subterminal line interrupted, strongly 
indicated only between veins 5 and the fold and for a 
short distance from inner margin; black discal dots at 
end of cell more or less fused and pronounced; cilia 
ocherous fuscous. Hind wing pure white in male; in 
female generally suffused with fuscous. Alar expanse, 
38-55 mm. 


The genitalia presenting no outstanding specific 
characters. 

TYPE LOcALITy: Texas (type in AMNH, ex Rutgers). 

Foon piants: Opuntia (Platypuntia) spp. 

Distrisution: Texas, Brownsville (Apr., June, July, 
Aug.), Corpus Christi (Sept., Oct.), Burnet County, 
San Benito (Mar., Aug., Sept.), Shovel Mountain 
(May), Kerrville (Apr.), Victoria (Oct., Nov.), Laredo 
(Sept.). 

This species is most readily distinguished from the 
others in the genus by its ocherous fuscous color, which 
seems to be constant. It is remarkably so in the 
specimens before me. O. nephelepasa and subumbrella 
are decidely gray in appearance. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Rather full notes on the life history and larval habits 
of junctolineella are given by Dodd and by Hunter, 
Mitchell, and Pratt. The latter also give descriptions 
of the larva and pupa. 

The species has two generations a year. 


499. Olycella junctolineella pectinatella (Hampson) 
Fiaure 530 


Olyca pectinatella Hampson, 7n Ragonot, Monograph, pt. 2, p. 35, 
1901. 


Olyca junctolineella (Hulst) Barnes and McDunnough, Check list 
of the Lepidoptera of Boreal America, No. 5695, 1917 (in 
art). 
Olycella cote (Huist) Dyar, Proc. Ent. Soc. Washington, 
vol. 30, p. 1384, 1928 (in part). 
Olycella junctolineella pectinatella (Hampson) Heinrich, Proc. 
U.S. Nat. Mus., vol. 86, p. 345, 1989. 

Known only from two males from the type locality. 

These two specimens are a trifle paler than normal 
junetolineella and are less clearly marked except for the 
pronounced discal spots. The transverse lines on the 
forewing are almost obsolete and the veins very slightly 
indicated by dark shading. 

Inasmuch as we have no larvae or females, or any 
information on the life history of the form from Jalapa 
(which is far south of the known range of typical 
junctolineella), I do not think we are justified in treating 
it as @ Mere synonym; or, on the evidence before us, 
as a distinct species. 

TYPE Locauity: Jalapa, State of Veracruz, México 
(type in USNM). 

Foop puant: Opuntia (Platypuntia) spp. 


500. Olycella nephelepasa (Dyar) 


Olyca nephelepasa Dyar, Ins. Insc. Menstr., vol. 7, p. 55, 1919. 

Olycella nephelepasa (Dyar), Proc. Ent. Soc. Washington, vol. 30, 
p. 134, 1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 
346, 1989. 

Similar in pattern and general appearance to junc- 
tolineella, but darker. The forewing is grayish fuscous 
with a slight brownish tint, but decidedly more grayish 
brown than ocherous fuscous. Also the hind tibiae of 
nephelepasa are heavily dusted with fuscous, while those 
of junctolineella are nearly pure white. Alar expanse, 
42-52 mm. 

The genitalia cannot be distinguished from those of 
junctolineella. 

Type Locatity: Tehuacén, México (type in USNM),. 

Foop piants: Opuntia (Platypuntia) spp. 

Disrrisution: México: Tehuacin (Sept.), México 
(city), Cuernavaca, Aguascalientes, San Luis Potosi 
(June). 

The known distribution of this species is confined to 
the central plateau of México. In the National Col- 
lection there is one female (determined as nephelepasa) 
from Monclova, México. This specimen is colored like 
typical subwmbrella and is, I think, only a southern 
example of that species. 

O. nephelepasa has two generations a year. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 243 


501. Olycella subumbrella (Dyar) 
Ficure 1019 


Olyca subumbrella Dyar, Ins. Insc. Menstr., vol. 13, p. 14, 1925. 

Olycella nephelepasa (Dyar), Proc. Ent. Soc. Washington, vol. 30, 
p. 134, 1928 (in part). 

Olycella subumbrella (Dyar) Heinrich, Proc. U. S Nat. Mus., 
vol. 86, p. 347, 1939.—Dodd, Biological campaign against 
prickly-pear, Brisbane, Australia, p. 39, 1940. 

Similar to nephelepasa except that the forewing is less 
brownish and more grayish. The general color is de- 
cidedly gray rather than brownish or ocherous. Alar 
expanse, 40-55 mm. 

Male genitalia as in nephelepasa and junctolineella. 
Female genitalia like those of junctolineella. The figure 
shows the extreme variation from typical junctolineella; 
but the differences in the shape of the sclerotized areas 
of the collar of the eighth segment and the length of the 
supporting rods of the collar are not specific. Every 
intergrade between this and typical junctolineella may 
be found in each of the species in the genus. 

Type Locauiry: Carlsbad, N. Mex. (type in USNM). 

Foop pLants: Opuntia (Platypuntia) spp. 

Distrisution: Texas, El Paso (Mar.); New Mexico, 
Carlsbad (Sept.); Arizona, Dewey, Redington, Palmer- 
lee, Paradise (Cochise County, Mar., Apr., May, June), 
Douglas (May, Aug.,) Pinal Mts. (Apr.), Haualapai 
Mts. (May); California, Warner (Sept.), Santa Clara 
(Apr.); Utah, Dividend (May, June), Stockton (May), 
Richfield (May); Nebraska, Scotts Bluff (June). 

In addition to the above there are before me two 
specimens from Monclova, Coahuila, México (E. 
Mortensen Collection, Sept. 1926), which probably are 
referable here. One (a male) was in the collection 
under junctolineella, the other (a female) under nephele- 
pasa. The male is in very poor condition but obviously 
belongs with the female. The latter is in fair shape, 
and its color is that of typical swbumbrella. More 
material is needed from northern México before we can 
determine what species inhabits that region. 

In 1928 Dyar sank subumbrella in the synonymy of 
nephelepasa; but Dodd informs me that the larval 
habits of the two are quite different. In nephelepasa 
“the larvae are banded with blue or blue-black and do 
not keep entrance hole in plant open for discharge of 
frass,’”’ while in subumbrella the larvae have ‘“‘rather pale 
purplish bands and maintain the hole open for the dis- 
charge of frass.’”’ These differences in larval habit, 
coupled with the slight but apparently consistent color 
differences in the moths seem to warrant the separation 
of subumbrella from nephelapasa. 

O. subumbrella has one generation a year. 


137. Genus Olyca Walker 


Olyca Walker, List, pt. 11, p. 725, 1857.—Hampson, in Ragonot, 
Monograph, pt. 2, p. 34, 1901.—Dyar, Proc. Ent. Soc. 
Washington, vol. 30, p. 133, 1928.—Heinrich, Proc. U. 8S. 
Nat. Mus., vol. 86, p. 348, 1939. (Type of genus: Olyca 
phryganoides Walker.) 


Tongue reduced (shorter than in Méelitara). An- 
tennae pubescent in both sexes (the pubescence longer 


in the male than in the female), slightly serrate in the 
male. Labial palpus of the male obliquely ascending; 
of the female porrect and downcurved. Maxillary 
palpus squamous. Hind wing with veins 7 and 8 
shortly anastomosed beyond cell; 3 and 5 connate (oc- 
casionally very shortly stalked). Eighth abdominal 
segment of male simple. 


Male genitalia with apical process of gnathos bifid; 
harpe with apex evenly rounded; vinculum short; 
anellus with base of plate narrowly sclerotized, arms 
moderately long and stout; aedeagus stout, moderately 
long, apex armed with many minute, hairlike spines. 


Female genitalia without signum; bursa copulatrix 
simple except for a few microscopic scobinations; ductus 
bursae short; ductus seminalis from bursa somewhat 
caudad of middle. 

Larvae not banded, solitary in habit, feeding in 
Platypuntias (presumably in the stems). 

Eggs unknown. 

Olyca is readily separated from other genera of the 
cactus-feeding Phycitinae having veins 3:and 5 of hind 
wing connate by having the antennae pubescent in both 
sexes. 

The male genitalia are similar to those of Olycella, 
differing only in slight details; the vinculum is slightly 
shorter, the uncus broader in proportion to its length, 
the cleft apical process of gnathos smaller, the elements 
of transtilla longer and straighter. 

The females differ chiefly in that they lack the signum 
in the bursa and the sclerotized plates in the opening 
of the ductus bursae. 

The genus as here defined contains only the type 
species from the West Indies. 


502. Olyca phryganoides Walker 
Figures 71, 531, 1020 

Olyca phryganoides Walker, List, pt. 11, p. 726, 1857—Hampson, 
in Ragonot, Monograph, pt. 2, p. 35, 1901.—Dyar, Proc. 
Ent. Soc. Washington, vol. 30, p. 134, 1928.—Heinrich, 
Proc. U. S. Nat. Mus., vol. 86, p. 348, 1939.—Dodd, Bio- 
logical campaign against prickly-pear, Brisbane, Australia, 
p. 40, 1940. 

General color (except hind wings) pinkish white, more 
or less spotted and suffused with black. Palpi, thorax, 
and underside of body heavily dusted and shaded with 
black. Forewing of male with no distinguishable ante- 
medial line, the latter being replaced by two more or 
less transversely extended black spots; subterminal line 
only partially and faintly indicated, irregularly dentate; 
vein ends marked with blackish dots or dashes; black 
discal spot large, conspicuous; below the discal dot a 
more or less extended black smudge. 

On the female about three-fourths of the forewing is 
suffused with black, the pinkish white color being 
strongly contrasted and limited to a rather narrow area 
along the costa, with a triangular projection at the end 
of the cell; terminal area and a patch on inner margin 
opposite discal spot also pale, but duller and less con- 
trasted than the costal color. Hind wing white with a 
blackish fuscous shade on terminal margin and a some- 


244 


what wider dark shade along costal margin; these dark 
shadings broader in the female than the male; also in 
the female the veins are outlined by fuscous scaling; 
cilia white with a dark subbasal line. Alar expanse, 
35-47 mm. 

Male genitalia figured from specimen from Azuda. 

Female genitalia with bursa copulatrix containing a 
scattering of microscopic scobinations, otherwise simple; 
scobinations in genital opening stronger and more 
dense, also in genital opening a few fine setae (the latter 
probably a generic character). 

Eggs unknown. 

Larva ‘cream or buff colored, with dark spiracular 
markings,” according to Dodd. 

Typ LocaLity: Hispaniola (type in BM). 

Foop piant: Opuntia (Platypuntia) spp. 

Distrinution: Dominican Repusuic: Azuda (Jan.). 
Hartt: Port-au-Prince (Jan.). 


Nothing has been published on the life history of this 
species, and little is known about it. ‘The larvae are 
presumably solitary in habit and confined to the Platy- 
puntias. The distribution of phryganoides is probably 
confined to the West Indies. 


138. Genus Alberada Heinrich 


Alberada Heinrich, Proce. U. S. Nat. Mus., vol. 86, p. 350; 1939. 
(Type of genus: Melitara parabates Dyar.) 

Tongue as in Melitara. Antenna of male> bipecti- 
nate, of female pubescent. Labial palpus porrect and 
downcurved. Maxillary palpussquamous. Hind wing 
with veins 7 and 8 anastomosing beyond cell; 3 and 5 
stalked. Highth abdominal segment of male simple. 

Male genitalia with apical process of gnathos bifid, 
the two prongs rather widely separated; harpe with the 
apex evenly rounded; vinculum short; anellus with base 
of plate narrowly sclerotized, arms moderately long and 
stout; aedeagus stout, weakly sclerotized in middle 
except on midventer. 

Female genitalia with sigsnum developed as a ridged 
plate; bursa copulatrix finely scobinate and wrinkled; 
ductus seminalis from middle of bursa. 

Larvae bluish, not banded; solitary feeders in the 
joints of Cylindropuntias. 

Eggs laid singly or in masses of two or three. 

The genus is close to Melitara, differing in the follow- 
ing characters: Veins 7 and 8 of hind wing anastomosed, 
3 and 5 stalked, aedeagus only partially sclerotized, 
apical process of gnathos with prongs well separated, 
bursa copulatrix with signum. 

The distribution is apparently limited to México and 
the southwestern part of the United States. 


503. Alberada parabates (Dyar) 
Fieures 533, 1021 


Melitara parabates Dyar, Proc. U. S. Nat. Mus., vol. 44, p. 322, 
1913; Proc. Ent. Soc. Washington, vol. 30, p. 184, 1928.— 
Barnes and McDunnough, Contributions, vol. 4, p. 175, 
1918.—Dodd, Council for Sci. and Ind: Res., Australia, 
Bull. 34, p. 27, 1927. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Alberada parabates (Dyar) Heinrich, Proc. U. 8. Nat. Mus., vol. 
86, p. 350, 1939. 

Forewing fuscous with area between lower vein of 
cell and costal margin and from antemedial to sub- 
terminal lines heavily dusted with white; area between 
lower vein of cell and inner margin and from base to 
subterminal line suffused with ocherous fuscous; on the 
middle of this area a more or less extended smudge of 
blackish brown; antemedial line black, bordered inwardly 
by a line of white scales, dentate and sinuate, a sharp 
dentation at vein 11, a longer one in the cell (extending 
nearly to middle of wing), another equally long and 
acute dentation at the fold, and two very slight denta- 
tions between 1b and inner margin; subterminal line 
black with a white outer border, dentate and sinuate, 
the angulations deep, the angulation between 5 and 6 
reaching almost to cell; area beyond subterminal line 
dark fuscous, paler in some specimens; along termen a 
row of black dots at the vein ends; discal black dot at 
end of cell conspicuous in most specimens. Hind wing 
white, semihyaline; costal margin bordered with 
fuscous and a fine dark fuscous line on termen for a 
short distance from apex; in many females a stronger 
fuscous shading in apical area. Alar expanse, 35-48 
mm. 

Male genitalia over twice as large as those of biden- 
tella; aedeagus more extensively sclerotized. Female 
genitalia larger than those of the other species of the 
genus and with scobinations in bursa finer. 

Eggs laid singly or in masses of two or three. 

Typr tocatiry: Cerritos, San Luis Potosi, México 
(type in USNM). 

Foop prants: Opuntia (Cylindropuntia) imbricata 
(Haworth) and probably several other Cylindropuntias. 

DistrizpuTion: Unirep states: California, San 
Diego, Warner (Sept.), Palm Springs (Apr.), Oceanside 
(Aug.), Riverside (Oct.); Arizona, Christmas (Gila 
County), Fort Grant (July), Oracle (July), Redington, 
Santa Catalina Mts. (Sept.), Baboquivari Mts. (Apr., 
June, July, Aug., Sept., Oct.), Sells Post Office (Indian 
Oasis, Apr.), Douglas (June, Sept.), Mohave County 
(Aug.); Texas, Presidio County (July), Brewster County. 
México: San Luis Potosi, Cerritos (Aug.), Tamaulipas, 
Tula (June). 

504. Alberada bidentella (Dyar) 
Ficures 532, 1022 


Zophodia bidentella Dyar, Proc. Ent. Soc. Washington, vol. 10, 
p. 114, 1908. 

Eumysia bidentella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221, 
192 


Alberada didentella (Dyar) Heinrich, Proc. U. S. Nat. Mus., vol. 
86, p. 352, 1939. 


Much smaller and paler than parabates but with simi- 
lar pattern, the ground color more ocherous than fus- 
cous, the white dusting on forewing heavier, the denta- 
tions of antemedial and subterminal lines shorter; discal 
dots distinct and not fused as is frequently the case in 
parabates. Alar expanse, 19-24 mm. 

Male genitalia similar to those of parabates but much 
smaller and with central ventral part of aedeagus more 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


narrowly sclerotized. Female genitalia appreciably 
smaller than those of parabates; bursa wrinkled and 
more coarsely scobinate. 

Typs Locaity: San Antonio, Tex. (type in USNM). 

Foop piant: Unknown. 

Distrisution: TZexras, San Antonio (July), San 
Benito (June, Aug., Sept.), Brownsville (June) ; Arizona, 
Phoenix, ‘‘route between Dewey and Salome.” 

A uniformly marked and colored species, known only 
from collected specimens. 


505. Alberada holochlora (Dyar) 
Figure 1023 


Zophodia holochlora Dyar, Ins. Insc. Menstr., vol. 13, p. 15, 1925. 
Alberada holochlora (Dyar) Heinrich, Proc. U. S, Nat. Mus., vol. 
86, p. 352, 1939. 


This is probably a synonym or, at most, a variety of 
bidentella. 'The three females of the type series are the 
only specimens I have seen. They are a trifle smaller 
than typical bidentella, and there are some slight, though 
hardly significant, differences in the female genitalia 
(shown in figs. 1022 and 1023). However, until males 
of holochlora are discovered and bidentella has been 
reared, it will be wiser to keep the two as separate 
species. 

According to Dodd the larvae are solitary in habit 
and dark blue and the eggs laid singly. 

Alar expanse, 18 mm. 

Types Locauity: Uvalde, Tex. (type in USNM). 

Foop piant: Opuntia (Cylindropuntia) leptocaulis De 
Candolle. 


139. Genus Nanaia Heinrich 


Nanaia Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 353, 1939. 
(Type of genus: Nanaia substituta Heinrich.) 


Tongue well developed. Antenna of male bipec- 
tinate; of female pubescent. Labial palpus obliquely 
porrect (second segment obliquely upturned nearly to 
top of face and third segment bent forward or slightly 
downcurved) ; third segment long (in the female as long 
as second segment), pointed in the male, blunt in the 
female. Maxillary palpus large, developed as a broad, 
curved, somewhat flattened tuft of scales which reaches 
well above middle of face. Hind wing with veins 7 and 
8 anastomosing beyond cell; 3 and 5 stalked. Eighth 
abdominal segment of male simple. 

Male genitalia with apical process of gnathos bifid, 
the two prongs narrowly separated; harpe with the apex 
somewhat tapering but bluntly rounded; vinculum 
moderately long (longer than broad); anellus with base 
of plate narrowly sclerotized, arms long, tapering, and 
slightly twisted; aedeagus moderately stout, strongly 
sclerotized throughout. 

Female genitalia without signum; ductus bursae and 
bursa copulatrix simple except for fine scobinations in 
ductus at genital opening; ductus bursae long; bursa 
copulatrix small; ductus seminalis from middle of bursa. 

Larva bluish, not banded; sclerotized plates sur- 
rounding body setae small; 2 setae in group VII on ab- 
dominal segments 7 and 8. 


245 


The larvae are solitary feeders in the trunks of Cylin- 
dropuntia and Trichocereus. 

Ege and egg-laying habits unknown. 

This genus is close to Alberada but distinguished by 
several characters: The forewings are distinctly nar- 
rower, the vinculum is longer in proportion to its width, 
the aedeagus more evenly sclerotized, the apical process 
of gnathos more narrowly cleft, the anellus more de- 
cidedly curved, the bursa simple, without signum or 
scobinations, the transverse markings on forewing al- 
most obliterated, and the maxillary palpi much larger. 
The maxillary palpi are similar to those of Sigelgaita, 
the moths of which resemble in general habitus those of 
Nanaia. The two genera, however, are easily dis- 
tinguished by their different labial palpi, porrect in 
Nanaia, upturned in the males of Sigelgaita. 

Known only from Pert. 


506. Nanaia substituta Heinrich 
Fieures 534, 1048 


Nanaia substituta Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 354 


General color (except hind wings) ocherous fuscous 
peppered with black and white; the type darker than 
most of the paratypes. Forewing with pale color con- 
fined to costal half of wing; terminal area and the area 
between cell and inner margin darker, with very little 
white dusting; in most specimens a rather pronounced, 
broad, longitudinal, ocherous fuscous shade in the fold; 
transverse and discal markings almost obsolete, in a few 
specimens the antemedial line faintly indicated and in 
the palest of the paratypes the discal black dots dis- 
tinguishable, also some black scaling along the veins. 
Hind wing white (whitish ocherous on female) with a 
smoky tint toward apex and termen; terminal margin 
blackish fuscous; cilia smoky white with a dark sub- 
basal line. Alar expanse, 37-40 mm. 

Female genitalia with no appreciable scobinations or 
granulations in bursa; ductus bursae minutely scobinate 
at genital opening, otherwise smooth. 


TypE Locauiry: Cuzco. Pert (type in USNM). 
Foop piant: Opuntia (Cylindropuntia)  exaltata 
Berger. 


140. Genus Cactoblastis Ragonot 


Cactoblastis Ragonot, Monograph pt. 2, p. 15, 1901.—Dyar, Proc. 
Ent. Soc. Washington, vo. 30, p. 135, 1928.—Heinrich, Proc. 
U. S. Nat. Mus., vol. 86, p. 354, 1939.—Janse, Journ. Ent. 
Soc. South Africa, vol. 8, p. 45, 1945. (Type of genus: 
Zophodia cactorum Berg.) 

Neopyralis Bréthes, in Ronna, Chacaras e Quinaes, vol. 20, No. 1, 
p. 18, 1920. (Type of genus: Neopyralis ronnai Bréthes.) 


Tongue considerably reduced. Antenna of male 
pubescent, of female shortly pubescent. Labial palpus 
of male ascending (upcurved), of female porrect. Hind 
wing with veins 7 and 8 shortly anastomosed beyond 
cell; 3 and 5 shortly stalked. Eighth abdominal seg- 
ment of male simple. 

Male genitalia with apical process of gnathos partially 
fused, the prongs separated only for a short distance; 


246 


harpe with apex evenly rounded; vinculum short; 
anellus with base of plate narrowly sclerotized, arms 
moderately long and rather slender, slightly twisted, 
very finely serrate on outer edges toward apices; 
aedeagus stout, moderately long. 

Female genitalia with sigsnum developed as a series of 
more or less fused plates; bursa copulatrix weakly and 
very finely scobinate; ductus seminalis from bursa at 
junction of ductus bursae and bursa copulatrix. 

Larva bright orange or red, with rows of large black 
spots resembling broken crossbands; two setae in group 
VII on abdominal segments 7 and 8. 

The larvae feed gregariously in the joints of Platy- 
puntia, Oylindropuntia, Trichocereus, Echinopsis, and 
Denmoza. 

Eggs laid in long chains. 

The genus as here defined is distinguished from other 
cactus-feeding phycitids by the following combination 
of characters: Antennae of both sexes pubescent; labial 
palpi upcurved in the male, porrect in the female; veins 
7 and 8 of hind wing shortly anastomosed; veins 3 and 5 
shortly stalked; apical process of gnathos partially 
fused; eighth abdominal segment of male simple; larvae 
bright orange or red, with rows of large black spots 
resembling broken crossbands, gregarious in habit. 

Four (possibly five) species are recognized as belong- 
ing to the genus. Its natural distribution is apparently 
limited to South America south of the Equator, but at 
least one of its species (cactorum) has been introduced 
and become established in Australia. 


507. Cactoblastis cactorum (Berg) 
Ficures 66, 535, 1024 


Zophodia cactorum Berg, Anales Soc. Cient. Argentina, vol. 19, p. 
276, 1885. 

Cactoblastis cactorum (Berg) Ragonot, Monograph, pt. 2, p. 16, 
1901.— Dodd, Council for Sci. and Ind. Res., Australia, Bull. 
34, p. 30, 1927; Bull. Ent. Res., vol. 27, p. 509, 1936; Bio- 
logical campaign against prickly-pear, Brisbane, Australia, 
177 pp., 1940.—Dyar, Proc. Ent. Soc. Washington, vol. 30, 
p. 135, 1928.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 
356, 1939. 

Head sordid whitish ocherous. Palpi pale cinereous, 
the tips of the maxillary palpi and the ends of the seg- 
ments of the labial palpi blackish fuscous. Thorax dull 
ocherous fuscous rather heavily dusted with blackish 
fuscous on posterior half. Forewing ocherous fuscous 
more or less dusted with white on costal half between 
antemedial line and apex; antemedial line black, angu- 
late, the apex of angle at vein 1b, sometimes obscure 
except on costal half; subterminal line black, ‘with a 
narrow whitish outer border and beyond this a faint 
fuscous band, the black line straight from near apex to 
vein 6, thence sinuate and dentate to inner margin, the 
ends of the dentations rounded; a black spot at end of 
cell and a few scattered black scales on disc; along 
termen at vein ends a row of seven distinct black dots. 
Hind wing white, semihyaline, costal margin narrowly 
bordered with fuscous and on termen a fine black line, 
the latter not extending to inner angle, on the female 
the fuscous shading is rather broad at apex and along 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


termen for a short distance from apex. Midtibia pale 
cinereous with a narrow, black, transverse band at 
outer fourth. Alar expanse, 23-40 mm. 

Genitalia not exhibiting any marked specific char- 
acters. 


TYPE LOCALITY: 
known). 

Foop puants: Opuntia (Platypuntia) spp. Appar- 
ently limited to the Platypuntias. 

Distrisution: Arecentina: La Plata, Concordia, 
Tacanitas, Santiago del Estero. Unrucauay: Piriapolis. 
AUSTRALIA (introduced and established). 


According to Dodd “‘cactorum is a native of Uruguay 
and the northern Argentine provinces of Entre Rios, 
Corrientes, Sante Fe, Santiago del Estero, Tucumén, 
Salta, and Chaco.” He also includes Paraguay and 
southern Brazil in its possible range, but we have no 
adult specimens from the latter localities. 

This is the species that has been used with such 
remarkable success in the biological campaign against 
the prickly-pear in Queensland and New South Wales. 
All available information on the species is given in 
Dodd’s 1940 paper. 


Argentina (location of type un- 


508. Cactoblastis ronnai (Bréthes) 


Neopyralis ronnai Bréthes, in Ronna, Chacaras e Quinaes, vol. 
20, No. 1, p. 18, 1920.—Costa-Lima, Terceiro catalogo dos 
insectos que vivem nas plantas do Brazil, p. 268, No. 1031, 
1936. 

Cactoblastis ronnai (Bréthes) Heinrich, Proc. U. 8. Nat. Mus., 
vol. 86, p. 357, 1939. 


Probably a synonym of cactorum. Its identity will 
have to wait upon rearing of Cactoblastis from southern 
Brazil. 

TyPE tocatity: Rio Grande do Sul, Brazil (type 
lost?). 

Foop pian: “Spineless cactus.” 


509. Cactoblastis doddi Heinrich 
Fieures 536, 1025 


Cactoblastis bucyrus Dodd (not Dyar), Council for Sci. and Ind. 
Res., Australia, Bull. 34, p. 30, 1927. 

Cactoblastis doddi Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 
358, 1939.—Dodd, Biological campaign against prickly-pear, 
Brisbane, Australia, pp. 39, 59, 75, 1940, 

Similar to that of cactorum except as follows: White 
dusting on forewing less contrasted, sparser; general 
color darker, decidedly grayish fuscous in specimens 
from Tucumén; dentations of subterminal line of fore- 
wing acute and their ends pointed; black dots along 
termen very faintly imdicated, normally altogether 


absent. Himd wing of male semihyaline white; of 
female dark smoky fuscous throughout. Alar expanse, 
31-41 mm. 


Male genitalia similar to those of cactorum, differing 
chiefly in the shorter cleft between the prongs at apex 
of gnathos. This character, however, is subject to 
some individual variation, and should be used with 
discretion. Female genitalia with scobinations of 
bursa. somewhat more uniformly distributed than in 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


other species of Cactoblastis, not an altogether reliable 
or satisfactory character in this genus. 

Typr Locauity: Tapia, Tucumén, Argentina (type 
in USNM). 

Foop puants: Opuntia (Platypuntia) sulphurea G. 
Don, Opuntia (Platypuntia) ficus-indica (Linnaeus). 

Represented only by the type series from the type 
locality. 

According to Dodd, this species ‘‘is distributed along 
the eastern edge and foothills of the Andes from Men- 
doza right to the northern boundary of the Republic in 
O. sulphurea, and almost certainly into southern Bolivia 
at altitudes to 8,000 fect and probably more. Hence, as 
far as our information goes, No. 49 (doddi) inhabits 
territory lying in between that of cactorum and the 
Peruvian insect (mundelli).”’ 

O. sulphurea seems to be the favored host of doddi. 
Dodd tells me that cactorum does not attack this cactus 
although it is abundant in territory within the range 
of that insect. He also states that there are consistent 
differences in the eggs and egg sticks between the two 
species and that their larvae can be distinguished in the 
field. I am unable to separate alcoholic specimens of 
the larvae with any certainty. The moths can be dis- 
tinguished easily enough by the following combination 
of characters: Forewing without terminal row of dots or 
with but 3 or 4 very faintly indicated; hind wing of 
male semihyaline white, of female dark smoky fuscous. 


510. Cactoblastis mundelli Heinrich 
Figure 537 
Cactoblastis mundelli Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 
359, 1939. 

Head ocherous. Palpi cinereous, dusted with black. 
Thorax ocherous fuscous, heavily dusted with white and 
black scales, especially on tegulae and posterior margin. 
Forewing ocherous fuscous with a fine dusting of white 
scales in costal area from base to apex; some black scal- 
ing on the veins; discal dot at end of cell somewhat 
obscured by a dark smudge which extends beyond the 
cell toward vein 1b; transverse black lines distinct and 
well contrasted against the ground color; dentations of 
subterminal line as in doddi; a row of seven small black 
dots on termen at the vein ends. Hind wing of male 
dull white with a faint smoky tint; veins faintly out- 
lined in fuscous ocherous; some fuscous shading along 
costa and a fine fuscous line on termen from apex to 
about vein 1b; hind wing of female pale smoky fuscous 
throughout. Alar expanse, 38-42 mm. 

Male genitalia with base of apical process of gnathos 
nearly square when viewed from beneath. Female 
genitalia not specifically different from those of cac- 
torum except for a somewhat shorter ductus bursae. 

Type Locatity: Arequipa, Peri (type in USNM). 

Foop piant: Opuntia (Cylindropuntia) exaltata Berger 
(apparently does not attack the Platypuntias). 

The species is known only from the type series from 
the type locality. 

It is easily recognized by the color of the hind wings 
and the ocherous suffusion on the forewings. 


247 


511. Cactoblastis bucyrus Dyar 
Fiaures 538, 1026 


Cactoblastis bucyrus Dyar, Ins. Insc. Menstr., vol. 10, p. 16, 1922; 
Proc. Ent. Soc. Washington, vol. 30, p. 135, 1928.—Heinrich, 
Proce. U. S. Nat. Mus., vol. 86, p. 360, 1939.—Dodd, Bio- 
logical campaign against prickly-pear, Brisbane, Australia, 
p. 39, 1940. 

The male is much darker than the males of other 
species of Cactoblastis. Palpi and thorax heavily dusted 
with blackish scales. Head and collar ocherous. Fore- 
wing brownish fuscous; white dusting inconspicuous, 
the pale scales more ocherous than white in the male, 
in female more whitish than ocherous and a trifle more 
noticeable; black antemedial and subterminal lines 
somewhat obscured by the dark ground color, conspicu- 
ous only toward costa, dentations of subterminal line 
as in doddi; a row of seven black dots along termen at 
vein ends. Hind wing brownish fuscous in both sexes. 
Alar expanse, 30-41 mm. 

Male genitalia with arms of anellus somewhat shorter 
than those of cactorum, doddi, and mundelli. Female 
genitalia with bursa very minutely and sparsely scobin- 
ate. The signum, like that of other species of Cacto- 
blastis, is individually variable. Extremes of variation 
are shown in figures 1026 and 1027. 

Typr tocauity: Mendoza, Argentina 
USNM). 


Foop piants: Trichocereus, Echinopsis, Denmoza. 


Distripution: ARGENTINA: Tucumén, Tapia, Men- 
doza, Catamarca, Andalgalé. 


(type in 


141. Genus Cahela Heinrich 


Cahela Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 361, 1939. 
(Type of genus: Olyca ponderosella Barnes and McDun- 
nough.) 

Tongue short but stout. Antennae of male and fe- 
male pubescent, the pubescence shorter in the female. 
Labial palpus of male obliquely upturned, of female 
porrect, with third segment downcurved and second and 
third segments longer than those of male; male palpus 
not extending above middle of front and with third 
segment short. Maxillary palpus squamous. Hind 
wing with veins 7 and 8 shortly anastomosing beyond 
cell; 3 and 5 stalked. Eighth abdominal segment of 
male simple. 

Male genitalia with apical process of gnathos fused; 
harpe with apex evenly rounded; vinculum short; 
anellus with base of plate narrowly sclerotized, arms 
long and broad, slightly twisted; aedeagus stout, 
sclerotized throughout. 

Female genitalia with signum developed as a ridged 
plate (a hollow, blunt, flattened, more or less thornlike 
projection into the bursa) ; bursa copulatrix large, finely 
scobinate expecially in the neighborhood of the signum; 
ductus bursae scobinate at genital opening; ductus 
seminalis from bursa near signum. 

Larvae whitish, not banded or conspicuously spotted; 
solitary in habit; stem borers in Cylindropuntias. 

Eggs laid singly. 


248 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


This genus and the following (Rumatha) are dis- 
tinguished from all others in the cactus-feeding group 
by the complete fusion of the apical process of the 
gnathos. Several male characters distinguish the two 
genera from each other, but Cahela is most easily recog- 
nized by the black longitudinal lines between the veins 
on the forewing. 


The genus is apparently limited in distribution to the 
southwestern part of the United States and northern 
México. 


512, Cahela ponderosella (Barnes and McDunnough) 
Figures 539, 1027 


Olyca ponderosella Barnes and MecDunnough, Contributions, vol. 
4, No. 2, p. 175, 1918. 

Zophodia purgatoria Dyar, Ins. Insc. Menstr., vol. 13, p. 222, 1925. 

Cactobrosis interstitialis Dyar, Ins. Insc. Menstr., vol. 13, p. 223, 
1925; Proc. Ent. Soc. Washington, vol. 30, p. 1386, 1928. 

Cactobrosis phoenicis Dvyar, Ins. Insc. Menstr., vol. 18, p. 223, 
1925; Proc. Ent. Soc. Washington, vol. 30, p. 136, 1938. 

Cactobrosis (?) ponderosella (Barnes and McDunnough) Dyar, 
Proc. Ent. Soc. Washington, vol. 30, p. 136, 1928.—Mc- 
Dunnough, Check list, No. 6284, 1939. 

Cahela ponderosella (Barnes and McDunnough) Heinrich, Proc. 
U.S. Nat. Mus., vol. 86, p. 361, 1939. 

Head, thorax, forewings, and body dark fuscous-gray 
peppered with white and with pronounced, longitudinal, 
black lines on the forewing; a long black line through 
the cell and extending from near base of wing to termen; 
another long black line from base to tornus running 
parallel and very close to the fold; in outer area, from 
beyond cell to apex and termen, five other shorter black 
lines, the largest and most pronounced above vein 6; 
all the black lines between and not on the veins; ante- 
medial and subterminal lines normally obsolete; in very 
few specimens a faint indication of a partial, black, 
antemedial line and in several specimens a dark shade 
from end of cell to middle of inner margin, but no 
trace of any transverse subterminal line; body some- 
what paler than forewing or thorax. Hind wing white, 
semihyaline, termen for a short distance from costa very 
faintly and narrowly edged with fuscous; a more pro- 
nounced fuscous shading along termen on the female. 
Alar expanse, 30-42 mm. 

Male genitalia show some variation in the shape and 
size of the terminal process of the gnathos in different 
specimens from any given locality. The extremes of 
variation are shown in the figure. In the female geni- 
talia the size of the signum and bursa varies somewhat 
in different specimens but the variations are slight and 
can be found in any series from one locality. 

TYPE LocaLities: Palm Springs, Calif. (ponderosella 
and phoenicis, in USNM); Colorado Desert, Yuma 
County, Ariz. (purgatoria, in USNM); Presidio, Tex. 
(nterstitialis, in USNM). 

Foop pPuant: Opuntia (Cylindropuntia) imbricata 
(Haworth) and probably other Cylindropuntias. 

Distrizution: California, Palm Springs (Apr., Aug.), 
San Bernardino (Apr., May); Utah, St. George (May, 
June); Nevada, Charlestown Mts. (July), Clark County 
(Apr., May, June); Arizona, Yuma County, Mohave 


County (Apr., May), Dewey (June), “‘en route from 
Dewey to Salome” (Apr.), Maricopa County (July), 
Prescott (Apr., June), Redington, Baboquivari Mts. 
(Pima County, May, July, Aug.), Phoenix (May), 
Tucson: (June), Douglas (May), Christmas (Gila 
County), Paradise (Cochise County, June); Texas, 
Brewster County, Alpine (Apr.), Presidio. 

Dodd states that the range of the species includes 
the central plateau of México, which is what we should 
expect. JI have seen no specimens from México. 

The species is remarkably uniform in color and mark- 
ing but varies considerably in size, which accounts for 
some of the synonymy. Superficially ponderosella is 
similar to both Eremberga leuconips (Dyar) and Cacto- 
brosis strigalis (Barnes and McDunnough). They also 
are dark gray with conspicuous black longitudinal lines 
on their forewings, but in leuconips and strigalis the 
black lines are on the veins, while in ponderosella they 
lie between the veins. This character at once dis- 
tinguishes it from other known cactus-feeding phycitids. 


142. Genus Rumatha Heinrich 


Rumatha Heinrich, Proc. U.S. Nat. Mus., vol. 86, p. 363, 1939. 
(Type of genus: Zophodia bihinda Dyar.) 

Tongue more or less developed (very short in glauca- 
tella, but not hidden by palpi). Antenna of male 
shortly serrate and pubescent, of the female simple and 
pubescent, the pubescence shorter in the female than in 
the male. Labial palpi porrect in both sexes; third 
segment of palpus about half as long as second. Max- 
illary palpus squamous. Hind wing with veins 7 and 8 
anastomosing beyond cell; 3 and 5 stalked. Highth 
abdominal segment of male simple. 


Male genitalia with apical process of gnathos fused; 
harpe with apex evenly rounded and with a subbasal 
sclerotized pocket (pkt., fig. 540) between sacculus and 
costa; uncus truncate and short in proportion to its 
breadth; vinculum short, truncate, almost square in 
outline; anellus with base rather broadly sclerotized, 
arms short, broad, slightly twisted; aedeagus very short, 
stout, partially sclerotized (on ventral half only). The 
entire genitalia have a squat appearance that is charac- 
teristic. 

Female genitalia with signum developed as a ridged 
plate with inwardly projecting ridge bluntly serrate 
(except in glaucatella, in which the signum is as in 
Cahela ponderosella); bursa copulatrix large, finely 
scobinate, especially in neighborhood of signum; ductus 
bursae scobinate at genital opening and with a pair of 
more or less defined sclerotized plates on the dorsal 
membrane of the ductus at the opening; ductus semi- 
nalis from bursa near signum (but somewhat farther 
removed than in Cahela). 

Larva of only one species (glaucatella) known; whit- 
ish, not banded or conspicuously spotted; solitary im 
habit; stem borer in Cylindropuntia. 

Eggs laid singly. 

This genus is very close to Cahela. Both genera have 
similar larvae and host associations and a like structure 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


of the gnathos; but there are too many other differences 
in male characters and adult habitus to permit their 
lumping. The partially sclerotized aedeagus, the short 
stout arms of the anellus, the squat appearance of the 
whole male genitalia, the porrect male labial palpi, and 
the serrate male antennae at once distinguish the males 
of Rumatha from those of Cahela; and the wing patterns 
readily separate both sexes. In Rumatha the discal 
dot is prominent and the transverse lines on the fore- 
wing are well defined for at least half their length. In 
Cahela the distinctive wing markings are longitudinal. 

Three species are recognized as belonging to the 
genus. Its distribution is limited apparently to the 
southwestern part of the United States and possibly the 
adjacent regions of northern México, although as yet 
no specimens have been received from México. 


513. Rumatha glaucatella (Hulst) 
Fieures 542, 1028 


Honora glaucatella Hulst, Ent. Amer., vol. 4, p. 117, 1888. 

Zophodia glaucatella (Hulst) Phycitidae of North Amer., p. 174, 
1890; U. S. Nat. Mus. Bull. 52, p. 4380, 1903.—Ragonot, 
Monograph, pt. 2, p. 23, 1901—McDunnough Check list, 
No. 6311, 1939. 

Rumatha glaucatella (Hulst) Heinrich, Proc. U. 8. Nat. Mus., 
vol. 86, p. 365, 1939. 

Palpi, head, and thorax pale fuscous, sparsely 
sprinkled with white; posterior margin of thorax edged 
with blackish fuscous. Forewing dull white, sparsely 
sprinkled with fuscous and with a very pale fuscous 
stain in a broad area bordering inner margin; ante- 
medial line angulate, fuscous, rather faint but complete 
and always distinguishable; subterminal line double, 
consisting of two parallel, faint, pale fuscous lines, 
almost vertical and but very slightly dentate; discal 
spot at end of cell blackish fuscous, prominent; a row of 
small blackish dots along termen between the vein ends. 
Hind wing whitish with a very pale fuscous line edging 
termen. Alar expanse, 15-20 mm. 

Male genitalia much smaller than those of either 
bihinda or polingella; basal portion of aedeagus narrower 
in proportion; harpe with apex more bluntly rounded 
than that of polingella but with width of harpe less in 
proportion to its length than that of bihinda. 

Female genitalia with signum similar to that of 
Cahela ponderosella, the inwardly projecting edge not 
appreciably serrate; sclerotized plates in genital open- 
ing very weak, hardly distinguishable except under 
very high magnification. 

Larvae solitary in habit, white, not banded or conspic- 
uously spotted. 

TypE Locatity: Texas (type in AMNH, ex Rutgers). 

Foop prant: Opuntia (Cylindropuntia) leptocaulis De 
Candolle. 

Distrisution: Texas, San Benito (May, June, July, 
Aug.), Brownsville (June), San Diego (May), Laredo 
(July), San Antonio; Florida (one female, so labeled and 
without other locality, from the Fernald Collection, in 
USNM). 

The labial palpus of the male is somewhat misleading. 

3003293—56——17 


249 


In natural position the third segment is projected 
forward, but in relaxed and badly prepared specimens 
it may be bent upward. 


514. Rumatha bihinda (Dyar) 
Figures 540, 1030 


Zophodia bihinda Dyar, Ins. Insc. Menstr., vol. 10, p. 173, 1922. 

Eumysia bihinda (Dyar), Ins. Insc. Menstr., vol. 13, p. 221, 
1925.—McDunnough, Check list No. 6301, 1939. 

Rumatha bihinda (Dyar) Heinrich, Proc. U. 8S. Nat. Mus., vol. 86, 
p. 366, 1939. 

Palpi, head, thorax, forewing, and abdomen dark 
fuscous, dusted with white, giving a decidedly grayish 
fuscous appearance to the moth; the white dusting 
heavy on costal half of forewing and upper surface of 
abdomen; discal spots and transverse markings on 
forewings blackish fuscous. Forewing with area be- 
tween cell and inner margin brownish, with little or no 
white dusting and with transverse lines obscured; costal 
half (especially above cell) strongly suffused with white; 
transverse antemedial line blackish, distinct only from 
costa to fold; subterminal line markedly dentate and 
sinuate, blackish, oblique, broad and conspicuous from 
costa to vein 8, with a slight dentation between veins 
8 and 6 and a deep angulation between veins 5 and 6 
extending to cell, between vein 5 and the fold straight 
and inwardly slanting, thence obscure to inner margin 
of forewing; discal dots at end of cell normally con- 
spicuous and fused into a single black spot, obscure in a 
few specimens; a row of black dots along termen at the 
vein ends; in some specimens faint traces of a black 
longitudinal line through center of cell and a line of 
blackish scales along the fold. Hind wing white, 
semihyaline, with a fine, faint, fuscous line along 
termen and some fuscous shading on costal margin. 
Undersurface of abdomen decidedly brownish fuscous, 
sparsely dusted with white. Legs with femora whitish, 
with some fuscous spotting; coxae uniformly dark 
brown, with no white dustings or markings, strongly 
contrasted against femora. Alar expanse, 30-36 mm. 

Male genitalia appreciably larger than those of other 
species in the genus; harpe broader in proportion to its 
length and with apex more broadly rounded. Female 
genitalia similar to those of polingella and hardly to be 
distinguished; signum with inner projecting edge irregu- 
larly and bluntly serrate. 

Typr Locauity: Jemez Springs, N. Mex. (type in 
USNM). 

Foop piant: Unknown. 

Distrrisution: Zeras, Alpine (Apr.); New Mezico, 
Jemez Springs (June, July); Arizona, Yuma County 
(Apr.), “en route from Dewey to Salome’ (Apr.), 
Dewey (May), Mohave County (Mar.); Nevada, Clark 
County (Mar., Apr., May), Bellevue (Washington 
County, May). 

This species has never been reared and its larva is 
unknown. From its close relationship to glaucatella we 
may expect that its host will prove to be one of the 
Cylindropuntias. 


250 


515. Rumatha polingella (Dyar) 
Fieures 541, 1029 


Zophodia polingella Dyar, Journ. New York Ent. Soc., vol. 14, 
p- 31, 1806.—McDunnough, Check list, No. 6312, 1939. 

Rumatha polingella (Dyar) Heinrich, Proc. U. 8. Nat. Mus., vol. 
86, p. 367, 1939. 

Similar in appearance to that of bihinda but with 
transverse antemedial and subterminal lines more dis- 
tinctly continued to inner margin of forewing; indenta- 
tions of subterminal line not so deep as in bihinda and 
not extending to cell; a scattering of pinkish scales 
among the white scales on costal area of forewing. Alar 
expanse, 23-35 mm. 

Male genitalia similar to those of bihinda but with 
harpe narrower, apex of harpe more acutely rounded, 
and aedeagus slightly narrower in proportion to its 
length. Female genitalia essentially like those of 67- 
hinda except that the sclerotized plates in genital open- 
ing are not so distinct and the signum is on the average 
smaller. 


TYPE Locatiry: Southern Arizona (type in USNM). 

Foop puant: Opuntia (Cylindropuntia) leptocaulis 
De Candolle. 

Distrisution: Arizona, Douglas (June, Aug.), Red- 
ington, Palmerlee, Paradise (Cochise County, July, 
Sept.), Pinal Mts. (Apr.), Baboquivari Mts. (June, July, 
Aug., Sept.), Santa Catalina Mts. (Aug.), ‘southern 
Arizona”’ (Apr.); Texas, Presidio (Aug.). 

The Texas record is from a small reared male, giving 
us our first food plant record for the species. 


143. Genus Yosemitia Ragonot 


Yosemitia Ragonot, Monograph, pt. 2, p. 17, 1901.—Heinrich, 
Proc. U.S. Nat. Mus., vol. 86, p. 368, 1939. (Type of genus: 
Spermatopthora graciella Hulst.) 

Yosemetia Hulst, U. S. Nat. Mus. Bull. 52, p. 429, 1903.—Dyar, 
Ins. Insc. Menstr., vol. 18, p. 220, 1925. (Misspelling.) 

Tongue well developed. Antenna of male weakly 
serrate and pubescent, of female simple and shortly 
pubescent. Labial palpi obliquely porrect. Maxillary 
palpus fan shaped and held vertically to the face. Hind 
wing with veins 7 and 8 anastomosing beyond cell; 
veins 3 and 5 stalked. Highth abdominal segment of 
male simple. 

Male genitalia with apical process of gnathos bifid; 
harpe with apex evenly rounded; vinculum moderately 
long; anellus with arms broad, short, slightly twisted, 
and base of plate broadly sclerotized; aedeagus short 
and slender, sclerotized throughout. 

Female genitalia with signum developed as a small, 
shortly spined plate; ductus bursae short; bursa copu- 
latrix finely scobinate, especially in area about signum; 
ductus seminalis from bursa near junction of bursa and 
ductus bursae. 

Larvae bluish, dark, not banded, or conspicuously 
spotted. 

The larvae feed gregariously (sometimes singly) in 
Echinocereus, Coryphantha, Homalocephala, and pre- 
sumably also in Echinocactus and Neomamillaria. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Eggs laid singly. 

This genus as here defined is distinguished by the fol- 
lowing combination of characters: Male antenna ser- 
rate and pubescent; labial palpi porrect in both sexes; 
maxillary palpi fan shaped; male genitalia with vincu- 
lum moderately long and rather narrow, apical process 
of gnathos bifid, anellus small and stout with broad 
short arms, aedeagus slender; eighth abdominal sezsment 
of male simple; female genitalia with signum a small, 
shortly spined plate, ductus bursae short and ductus 
seminalis from bursa copulatrix near junction of bursa 
and ductus bursae; larvae not banded and normally 
gregarious. 

The male genitalia have a characteristic habitus 
which makes them easy to place generically; but the 
differences between species are very slight and not alto- 
gether trustworthy, hardly more than might be expected 
within specific limits. 

Four species are recognized as belonging to the genus. 
Its distribution is the southwestern part of the United 
States and México. 


516. Yosemitia graciella (Hulst) 
Figures 68, 547, 1033 


Spermatopthora graciella Hulst, Ent. Amer., vol. 3, p. 184, 1887. 

Zophodia graciella (Hulst), Phycitidae of N. Amer., p. 173, 1890. 

Yosemitia graciella (Hulst) Ragonot, Monograph, pt. 2, p. 13, 
1901.—Barnes and McDunnough, Contributions, vol. 3, p. 
199, 1916.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 369, 
1939.—MeDunnough, Check list, No. 6293, 1939. 

Yosemeiza graciella (Hulst), U. S. Nat. Mus. Bull. 52, p. 429, 
1908.—Dyar, Ins. Insc. Menstr., vol. 13, p. 220, 1925. 

Forewing pale brownish fuscous dusted and streaked 
with black and dusted with white scales; the white 
scaling concentrated on and strongly whitening the 
costal half of the wing; the black scaling thinly dusted 
over the lower half of the wing (between cell and inner 
margin) and outlining the veins; transverse lines incom- 
plete, blackish; the antemedial line indicated only by a 
transverse dash in the cell and a dot or very short streak 
on inner margin; subterminal line prominent from costa 
near apex to vein 8 (Sometimes to vein 6), inwardly 
slanting, interrupted between veins 6 and 5, obscure 
between vein 2 and inner margin; discal dots fused into 
a line of black scales on discocellular vein; a row of 
small black dots along termen between the vein ends. 
Hind wing whitish to very pale smoky fuscous, faintly 
darker at apex, along costa, and narrowly along termen; 
cilia white with a very fine, pale fuscous, subbasal line. 
Alar expanse, 25-30 mm. 

Male genitalia very little different from those of other 
species in the genus; the vinculum is not so broad as 
that of fieldiella or so long as that of didactica; the anelli 
of the several species seem to offer the best characters 
for separating the species on genitalic characters; those 
of graciella and longipennelia are much alike, but in 
didactica the arms appear to be more sharply twisted 
and in fieldiella the basal portion is more narrowly 
sclerotized and the free arms, therefore, correspondingly 
longer. These characters, however, may not be con- 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 251 


stant in long series. Female genitalia with signum a 
trifle smaller than that of any other Yosemitia, other- 
wise not specifically distinguished. 

Larvae dark, dull blue and solitary or gregarious in 
habit, according to Dodd. 

Type tLocauity: Blanco County, Tex. 
AMNHG, ex Rutgers). 

Foop puants: Echinocereus viridiflorus Engelmann, 
E. polyacanthus Engelmann, and Coryphantha aggregata 
(Engelmann). 

Distrrsution: Colorado, Denver (July), and one 
specimen with only the State designation; Nevada, 
Clark County (Apr., May); California, San Bernardino 
County (Apr.), Providence Mts. (May), one specimen 
with only State designation (Apr.); Arizona, Yavapai 
County, Ajo (Pima County, Mar.), Baboquivari Mts. 
(May), White Mts. (June), Pinal Mts. (Apr.), Quijotoa 
Mts, (June), Santa Rita Mts. (June), Sells Post Office 
(Pima County, May), ‘‘en route from Dewey to Salome” 
(Apr.), Mojave County (May), Roosevelt (June), 
Phoenix (Mar., Apr.), Redington, no locality except 
the State (2 specimens reared from Coryphantha aggre- 
gata, June); New Mexico (Mar.); Texas (no specific 
locality, one specimen reared from Echinocereus viridi- 
florus; Apr.). 


(type in 


517. Yosemitia longipennella (Hulst) 
Ficures 546, 1032 


Zophodia longipennella Hulst, Ent. Amer., vol. 4, p. 118, 1888. 

Zophodia graciella (Hulst, in part), Phycitidae of N. Amer., p. 
173, 1890. 

Yosemitia graciella (Hulst, in part) Ragonot, Monograph, pt. 2, 
p. 13, 1901. 

Yosemetia graciella longipennella (Hulst) Barnes and McDun- 
nough, Check list of the Lepidoptera of Boreal America, 
No. 5699, 1917. 

Yosemetia longipennella (Hulst) Dyar, Ins. Insc. Menstr., vol. 13, 
p. 220, 1925. 

Yosemitia longipennella (Hulst) Heinrich, Proc. U. S. Nat. Mus., 
vol. 86, p. 370, 1939.—McDunnough, Check list, No. 6294, 
1939. 

Similar to graciella except black dusting on forewing 
sparse, veins not or but very faintly outlined by black 
scales, transverse subterminal line not interrupted 
between veins 6 and 5, dentate. Alar expanse, 21-26 
mm. 

Genitalia similar to those of graciella but with signum 
of female a trifle larger. 

Larvae dark dull blue and gregarious in habit, accord- 
ing to Dodd. 

Tyrer Locauity: Texas (type in AMNH, ex Rutgers). 

Foop piant: Homalocephala texensis (Hopffer). Ac- 
cording to Dodd the larvae also feed in Neomamillaria. 

Disrrisution: Texas, Uvalde (June), “Big Bend” 
(Apr.), San Antonio (June), Van Horn (June), San 
Diego (Apr.), San Benito (Apr., May). 


518. Yosemitia fieldiella (Dyar) 
Figure 544 


Zophodia fieldiella Dyar, Ins. Inse. Menstr., vol. 1, p. 35, 1913.— 
McDunnough, Check list, No. 6310, 1939. 


Yosemitia fieldiella (Dyar) Heinrich, Proc. U. S. Nat. Mus., 
vol. 86, p. 371, 1939. 

Forewing heavily dusted with white on costal half 
and with some scattered white scales on remainder of 
wing; area between inner margin and cell pale brownish; 
antemedial line incomplete, distinct only from costa to 
lower vein of cell; subterminal line complete but obscure 
except for the blackish costal dash, dentate, the incurva- 
tion between veins 6 and 5 shallow; from antemedial 
line, where it meets the cell, a thin, obscure, curved, 
blackish line extends to upper outer angle of cell; 
discal dot small, obscure; on some specimens a few black 
scales outlining vein 6; on termen a row of obscure 
blackish dots lying between the vein ends. Hind 
wing white, in female faintly tinted with smoky fuscous; 
a very pale fuscous line along termen. Alar expanse, 
22-25 mm. 

Male genitalia with basal portion of anellus more 
narrowly sclerotized and arms correspondingly longer 
than in the other species of the genus; vinculum also 
broader and shorter. Female genitalia similar to those 
of longipennella, but signum somewhat larger. 

Type Locauity: La Puerta Valley, Calif. (type in 
USNM). 


Foop piant: Unknown 


Distrizution: California, La Puerta Valley (July); 
Arizona, Catalina Springs (May). 


519. Yosemitia didactica Dyar 
Figure 545 
Yosemitia didactica Dyar, Proc, U.S. Nat. Mus., vol. 47, p. 
408, 1915.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 
372, 1939. 

Forewing heavily dusted with white on costal half; 
a short blackish line on midcosta; lower half of wing 
concolorous with thorax; antemedial line obscure, 
incomplete; discal dot at outer end of cell distinct, 
blackish; subterminal line complete, dentate, double for 
a short distance from apex and thence outwardly 
margined by a narrow pale shade, obscure toward inner 
margin, parallel to termen; veins 5 to 9 very faintly 
outlined by dark scaling; terminal row of dots almost 
obsolete. Hind wing whitish, faintly smoke tinted, 
somewhat darker toward apex and along termen; cilia 
with a pale smoky subbasal line. Alar expanse, 
22-23 mm. 

Male genitalia with vinculum rather longer than that 
of any other species in the genus; anellus with arms 
bent about aedeagus. Female genitalia similar to 
those of graciella. 

Type Locauity: Tehuacin, México (type in USNM). 

Foop piant: Unknown. 

Distrisution: Mixico: Tehuacén (May, June), 
Orizaba. 

This species resembles graciella but is somewhat paler 
and the male has slightly darker (smoky) hind wings. 
It is at once distinguished by the dark line on the 
midcosta of the forewing. Its life history is unknown. 


252 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


144. Genus Tucumania Dyar 


Tucumania Dyar, Ins. Insc. Menstr., vol. 13, p. 224, 1925.— 
Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 373, 1939. 
(Type of genus: Tucumania tapiacola Dyar.) 

Tongue rather short, but stout (as im Melitara). 
Antenna of male shortly serrate and pubescent, of 
female simple and shortly pubescent. Labial palpus 
of male upturned, reaching almost to level of top of 
eye; of female porrect (the second segment oblique, 
the third slightly downcurved). Maxillary palpus 
squamous. Hind wing with veins 7 and 8 anastomos- 
ing for a short distance beyond cell; 3 and 5 stalked. 
Highth abdominal segment of male simple. 


Male genitalia with apical process of gnathos bifid; 
harpe with apex bluntly pointed or elliptically rounded; 
vinculum moderately long (it is somewhat foreshortened 
in fig. 543), broad; anellus with base of plate moderately 
sclerotized, arms rather broad and long, slightly 
twisted; aedeagus long, slender; penis weakly scobinate 
toward outer extremity. 


Female genitalia with signum a small ridged or 
granulate plate; bursa copulatrix with some fine 
scobinations in the area about signum; ductus seminalis 
from bursa near junction of bursa and ductus bursaae. 


Larva purplish or wine colored with sclerotized areas 
about body tubercles dark brown and large; two setae in 
group VII on abdominal segments 7 and 8. 


The larvae are solitary feeders in the joints of Platy- 
puntias. 

Eggs laid singly. 

This genus is distinguished from others having serrate 
and pubescent male antennae and squamous maxillary 
palpi by its host association, its upturned male palpi, 
slender aedeagus, female genitalia with signum, and 
with ductus seminalis from the bursa. It is nearest to 
Eremberga, but that genus is broad-winged and has a 
flat, more strongly sclerotized anellus, a scobinate 
aedeagus, stouter male genitalia, no signum, and the 
ductus seminalis coming from the ductus bursae. In 
Tucumania the wings are long and rather narrow. 


The known distribution is Argentina and Uruguay. 


520. Tucumania tapiacola Dyar 
Ficures 65, 5438, 1034 


Tucumania tapiacola Dyar, Ins. Insc. Menstr., vol. 13, p. 225, 
1925.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 374, 
1939.—Dodd, Biological campaign against prickly-pear, 
Brisbane, Australia, pp. 39, 59, 83, 1940. 

Palpi, face, head, thorax, and forewing dark grayish 
fuscous with a sparse scattering of obscure whitish 
scales (Dyar states that the coxae and parts of the 
femora and tibiae of the legs are black, but even on 
these parts there is some scattered pale scaling and the 
ground color is fuscous rather than black). Forewing 
almost uniformly dark, sometimes a very faint luteous 
tint in the median area and a slight pale suffusion in 
terminal area; transverse lines black but not strongly 


contrasted against the dark ground color; antemedia 
line bidentate, its apex extending almost to center of 
cell; subterminal line dentate, sinuate, the dentations 
short, bordered outwardly by a pale line and beyond 
this by a rather broad blackish band, from costa well 
before apex; discal spot at end of cell large; veins beyond 
cell faintly outlined by dark scaling; a row of black dots 
along termen at the vein ends. Hind wing whitish, 
semihyaline, strongly shaded with fuscous at apex and 
narrowly along margin of termen almost to anal angle, 
this fuscous shading more extended on the female. Alar 
expanse, 27-30 mm. 


Male genitalia with apex of harpe bluntly pointed; 
anellus with the apices of the arms appreciably broad- 
ened. These are presumably specific characters. I 
have seen no males of any other species of Tucumama. 
Female genitalia with scobinations of bursa very weak 
and distinguishable only in area surrounding signum; 
signum somewhat granulate. 

TyrE Locatiry: Tapia, Tucum4n, Argentina (type in 
USNM). 

Foop piants: Opuntia (Platypuntia) discolor Britton 
and Rose, O. (Platypuntia) aurantiaca Lindley. 


DistRIBUTION: ARGENTINA. 


Only three specimens are before me, the male type 
and a pair (o' and @) reared in Australia from Argen- 
tine stock and sent me by Dodd. 


521. Tucumania porrecta Dyar 
Ficuré 1035 


Tucumania porrecta Dyar, Ins. Insc. Menstr., vol. 13, p. 225, 
1925.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 375, 1939. 
Large and paler than fapiacola. ‘Thorax pale fawn 
color. Forewing pale purplish fuscous with black 
markings diffused; antemedial and subterminal lines 
narrow, black, irregularly dentate, distinguishable 
throughout but somewhat interrupted; discal dots at 
end of cell rather large but not sharply contrasted 
against ground color of the wing because of scattered 
black dusting in the surrounding area; dots along termen 
distinct; a short black streak from base through middle 
of cell to apex of angulate antemedial line. Hind wing 
white, faintly smoke tinted, especially toward apex. 
Legs pale purplish fuscous; femora and tibiae trans- 
versely banded with blackish fuscous on outer sides. 
Alar expanse, 32-35 mm. 

Female genitalia with scobinations of bursa very fine 
but denser than in tapiacola; a small patch of somewhat 
larger scobinations in neck of bursa; signum larger, with 
a thin even keel but no granulations. 

Typr Locauity: Paysandi, Uruguay 
USNM). 

Foop prant: Opuntia (Platypuntia) sp. 

Distrisution: Urueuay. 

Represented in the National Collection only by the 
type and paratype from the type locality (Dodd, Feb. 
1925), both females. The male is unknown. 


(type in 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 253 


145. Genus Eremberga Heinrich 


Eremberga Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 375, 1939. 
(Type of genus: Cactobrosis leuconips Dyar.) 

Tongue reduced (small and weak). Antenna of male 
serrate and pubescent, of female simple and shortly 
pubescent. Labial palpus of male upturned, of female 
obliquely porrect. Maxillary palpus squamous. Hind 
wing with veins 7 and 8 very shortly anastomosed be- 
yond cell: 3 and 5 stalked. Highth abdominal segment 
of male simple. 

Male genitalia with apical process of gnathos bifid; 
harpe with apex evenly rounded; vinculum broad and 
short; anellus with base of plate broadly and strongly 
sclerotized, arms short, broad, not twisted or bent and 
with apices pointed; aedeagus moderately long, rather 
slender, sclerotized throughout and with a minutely 
scobinate flange at apex. 

Female genitalia without signum; bursa copulatrix 
smooth or with a few scattered microscopic scobina- 
tions; ductus bursae short, scobinate at genital open- 
ing; ductus seminalis from ductus bursae. 

Larva white with dark spots forming incomplete 
crossbands; two setae in group VII on abdominal seg- 
ments 7 and 8. 

The larvae are solitary or semigregarious feeders in 
Echinocereus. The larva of only one species (leuconips) 
is known but the characters here given presumably 
apply to the genus. 

Eggs laid singly. 

This genus is close to Tucumania and has many 
characters in common with Olyca. The latter, however, 
has veins 3 and 5 of the hind wing connate, the ductus 
seminalis from the bursa rather than from the ductus 
bursae, the male labial palpus oblique, the aedeagus 
stout, and the basal plate of the anellus narrowly 
sclerotized. The characters separating Hremberga from 
Tucumania have been discussed in connection with the 
latter genus. 

Three species are here recognized as belonging to 
Eremberga. Its distribution is the southwestern part 
of the United States and México. 


522. Eremberga leuconips (Dyar) 
Fiaures 548, 1031 
Cactobrosis leuconips Dyar, Ins. Inse. Menstr., vol. 13, p. 224, 
1925; Proc. Ent. Soc. Washington, vol. 30, p. 136, 1928.— 
McDunnough, Check list, No. 6288, 1939. 
Eremberga leuconips (Dyar) Heinrich, Proc. U. 8. Nat. Mus., 
vol. 86, p. 376, 1939. 

Palpi, head, thorax, and forewing grayish fuscous 
densely sprinkled with white, giving the insect a pale 
slate color. Labial palpus banded with blackish fuscous 
toward the ends of the segments. Forewing with vein 
3 to 10 outlined in black, the black scaling especially 
strong on lower vein of cell; antemedial and subterminal 
lines very fine and faint but usually discernible, black; 
antemedial line acutely angulate and irregularly sinuate 
and dentate, more or less broken and normally obliter- 
ated at costa; subterminal line also irregularly sinuate 


and dentate, decidedly slanting, obscured toward costa; 
no distinct discal marks at end of cell and no dots along 
termen. Hind wing of male glistening white, semi- 
hyaline with a band of fuscous shading along costa and 
a fine, pale fuscous line along termen for a short distance 
from apex; of female, dark smoky fuscous, the fuscous 
shading extending into the cilia and strongly outlining 
most of the veins, and the wing paler towards base. 
Alar expanse, 26-27 mm. 

Male genitalia with lateral edges of anellus finely and 
irregularly serrate; vinculum with terminal margin 
evenly rounded, lateral margins not concave or exca- 
vate. Female genitalia without any trace of signum; 
bursa nearly smooth; ductus seminalis from ductus 
bursae a short distance from genital opening. 

Typr Locauity: Baboquivari Mts., Ariz. (type in 
USNM). 

Foop piant: Echinocereus polyacanthus Engelmann. 

Distrisution: Arizona, Baboquivari Mts. (July, 
Sept.), Roosevelt (July), Oracle (July), Huachuca Mts. 
(Aug.), Chiricahua Mts., Mohave County (Sept.). 

This species bears a strong resemblance to Cactobrosis 
strigalis (Barnes and McDunnough) and might easily 
be confused with it. The latter, however, has filiform 
maxillary palpi while those of leuconips are distinctly 
squamous. Dyar seems to have overlooked this char- 
acter in placing many of his species. Also there is a 
difference in the longitudinal markings. In sétrigalis 
the strongest black longitudinal line is that along the 
top of the cell and vein 6, while in leuconips the strongest 
line is that along the lower vein of the cell. 

In two males and some of the females of lewconips 
there is a faint brownish fuscous suffusion on the lower 
third of the forewing (bordering the inner margin), but 
this is not distinguishable on all specimens and does 
not seem to be a specific character. 


523. Eremberga creabates (Dyar) 
Fiaure 550 


Olyca creabates Dyar, Ins. Inst. Menstr., vol. 11, p. 29, 1923. 

Cactobrosis creabates (Dyar), Proc. Ent. Soc. Washington, vol. 30, 
p. 136, 1928—McDunnough, Check list, No. 6289, 1939. 

Eremberga creabates (Dyar) Heinrich, Proc. U. 8. Nat. Mus., vol. 
86, p. 377, 1939. 

Palpi grayish fuscous. Head grayish fuscous shaded 
with white. Thorax luteous, whitish toward anterior 
margin. Forewing with basal area (to antemedial line), 
and all the area between antemedial and subterminal 
transverse dark shade, and the fold and costa, white; 
area between fold and inner margin luteous; outer area 
(beyond subterminal dark shade) ashy white, shading 
to luteous at tornus; transverse antemedial line well 
contrasted against ground color, thin black, irrorate, 
forming a sharp angle at the fold, the apex of the angle 
extending almost to the middle of the fold; subterminal 
line obsolete, replaced by a dark, transverse shade be- 
low end of cell and some scattered blackish dusting 
toward apex; veins 2 to 10 and upper and lower veins of 
cell more or less outlined in black, the lines very faint 


254 


on all the veins except vein 4; along termen, between the 
vein ends, a row of very faint blackish dots; no discal 
marks at end of cell, Hind wing shiny white, semihya- 
line, with a faint pale fuscous shading along costa, on 
veins 6, 7, and 8, and at extreme apex. Alar expanse, 
34 mm. 

Genitalia with lateral margins of anellus smooth; 
vinculum with terminal margin straight, rather broad, 
lateral margins excavate. 

Typr LocaLiry: San Diego, Calif. (July; type in 
USNM). 

Foop prant: Unknown. 

Known only from the unique male type. It is a 
striking species and should be easily recognized from 
the description and genitalic features. 


524, Eremberga insignis Heinrich 
Figure 549 
Eremberga insignis Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 
378, 1939. 

Palpi, face, head, thorax, and forewing dark grayish 
fuscous. Forewing very faintly dusted with white on 
costal half; lower half of wing faintly shaded with dull 
luteous ocherous; antemedial and subterminal lines as 
in leuconips, except antemedial not obliterated toward 
costa; veins 2 to 9 very faintly outlined in black, the 
black lining most distinct on lower vein of cell; a con- 
spicuous black spot at end of cell; along termen, be- 
tween the vein ends, a row of rather conspicuous black 
dots. Hind wing shiny white, semihyaline, with a 
fuscous shade bordering costa and a pale fuscous line 
on termen for a short distance from apex. Alar expanse, 
35 mm. 

Male genitalia with lateral margins of anellus smooth; 
vinculum with terminal margin straight and narrow, 
lateral margins outwardly angled. 

Typr LocaLity: San Luis Potosi, México (type in 
USNM). 

Foop piant: Unknown. 

This species is known only from the male type. It is 
easily distinguished from the other two species in the 
genus by the conspicuous discal spot on forewing. 


146. Genus Salambona Heinrich 


Salambona Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 379, 1939. 
(Type of genus: Zophodia analamprella Dyar.) 

Tongue well developed. Antenna of male pubescent 
and slightly serrate, of female simple and shortly pubes- 
cent. Labial palpi of both sexes porrect with the third 
segment downcurved, the third segment slightly longer 
in the female than in the male. Maxillary palpus 
squamous. Hind wing with veins 7 and 8 anastomosed 
for more than one-half their lengths beyond the cell; 3 
and 5 stalked. Eighth abdominal segment with a pair 
of strong ventrolateral hair tufts. 

Male genitalia with apical process of gnathos bifid, 
small; uncus constricted toward apex; harpe with apex 
oblique; vinculum long; anellus with base of plate nar- 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


rowly sclerotized, arms long, curved and twisted part 
way around aedeagus; aedeagus long, stout. 

Female genitalia without signum or scobinations in 
bursa; bursa small, smooth; ductus bursae long, slender, 
smooth; ductus seminalis from near end of bursa. 

Larvae grayish green or blackish, according to Dodd; 
not banded or conspicuously spotted; solitary feeders in 
fruits of Platypuntia. 

Eggs laid singly. 

The genus is distinguished from other genera in the 
cactus-feeding group by the following combination of 
characters: Antenna of male serrate and pubescent; 
labial palpi of both sexes porrect and downcurved; 
maxillary palpus squamous; harpe of genitalia with 
apex oblique; vinculum long; eighth abdominal segment 
of male bearing a pair of ventrolateral tufts; bursa 
copulatrix of female small and without signum or 
scobonations (smooth); ductus seminalis from near end 
of bursa; larvae unbanded, dark, fruit feeders in 
Platypuntias. 

Only the type species is recognized as belonging to 
the genus. It is known only from Argentina. 


525. Salambona analamprella (Dyar) 
Fieures 552, 1045 


Zophodia analamprella Dyar, Ins. Insc. Menstr., vol. 10, p. 17, 
1922. 

Salambona analamprella (Dyar) Heinrich, Proc. U. 8. Nat. Mus., 
vol. 86, p. 380, 1939.—Dodd, Biological campaign against 
prickly-pear, Brisbane, Australia, p. 40, 1940. 

Palpi, head, thorax, and forewings dark stone gray; 
the scales under magnification dark grayish fuscous 
tipped with dull white. Forewing with the costa 
broadly margined (to top of cell) with white, the white 
streak diminishing toward base of wing and terminating 
before apex; no transverse lines, or discal or terminal 
dots. Hind wing semihyaline with a smoky shade 
along costa and a narrow smoke-brown line along 
termen; the smoky shade somewhat more extended on 
the female. Alar expanse 25-27 mm. 

Male genitalia characters as given for the genus. 
Female genitalia with bursa very small and ductus 
bursae long and very slender. 

TypE LocaLity: Carmen Patagones, Argentina (type 
in USNM). 

Foop pLant: Opuntia (Platypuntia) sulphurea G. 
Don and probably other species of Platypuntia. 

Distrisution: ARGENTINA: Carmen Patagones 
(Jan.), Andalgalé (Mar.), La Rioja. 

According to Dodd this insect is the chief enemy of 
the cochineal scales in Argentina and is usually preda- 
ceous on cochineal (Dactylopius spp.) but not un- 
commonly the larvae feed in Opuntia fruit and flower 
buds. From the genitalic and other structural charac- 
ters of the moth I am inclined to doubt this. I think 
analamprella will prove to be primarily a cactus feeder 
and only secondarily predaceous on the cochineal scales 
on the cactus. It is the other way around with Laetila 
coccidivora (Comstock). The latter is a true predator 
and follows its coccid hosts no matter to what plant 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


they may go. It also varies its diet somewhat by 
occasional feedings on buds and flowers. According to 
Dodd it sometimes feeds in Opuntia flowers; but this is 
a secondary habit and the association with Opuntia 
accidental. Laetilia is close to but not a part of the 
cactus-feeding group of Phycitinae. Salambona, on the 
other hand, is, in all adult characters, definitely a 
member of the group. 

The species is a striking one, easily recognized by the 
strongly contrasted, white costal stripe on forewing. 


147. Genus Parolyea Dyar 


Parolyca Dyar, Proc. Ent. Soc. Washington, vol. 30, p. 17, 
1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 381, 
1939.—(Type of genus: Olyca asthenosoma Dyar.) 


Tongue well developed. Antenna of male unipecti- 
nate. Labial palpus of male upcurved. Maxillary 
palpus squamous. Hind wing with veins 7 and 8 
shortly anastomosed beyond cell; 8 and 5 shortly 
stalked. Eighth abdominal segment with a strong 
pair of ventrolateral tufts. 

Male genitalia with apical portion of gnathos bifid; 
uncus narrowed well before apex; harpe with apex 
oblique; vinculum long, its terminal margin rounded, 
its lateral margins excavate (probably a specific charac- 
ter only); anellus with base of plate broadly sclerotized, 
arms long, rather broad and slightly twisted; aedeagus 
long, stout; penis weakly scobinate. 

The genus is known only from the male of its type 
species. Its biology is unknown, but from the genitalic 
and other structural characters of the adult its larvae 
are presumed to be cactus feeders. It is easily recog- 
nized, for it is the only genus in the cactus-feeding 
group with unipectinate antenna. The habitat is 
French Guiana. 


526. Parolyca asthenosoma (Dyar) 
Figures 69, 551 


Olyca asthenosoma Dyar, Ins. Insc. Menstr., vol. 7, p. 55, 1919. 

Parolyca asthenosoma (Dyar), Proc. Ent. Soc. Washington, vol. 
30, p. 137, 1929.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, 
p. 381, 1939. 

Palpi, head, and thorax sordid white. Forewing 
white, with a yellowish tint on area between fold and 
inner margin; antemedial band angulate, consisting of 
parallel black lines and a central white line; a black 
oblique dash in median area from inner margin to dashes 
at apex, black dots on the veins, and a black spot on 
inner margin; a black discal dot at end of cell and some 
black scaling on bases of veins 2 to 4; a row of small 
black dots along termen, between the vein ends. Hind 
wing white, semihyaline, with a pale fuscous shade 
along costa, a narrow fuscous line along termen, and 
some pale fuscous scaling on veins 2 to 8. Alar expanse, 
30 mm. 

Male genitalia with lateral margins of vinculum 
excavate. 

Type Locatity: Maroni River, French Guiana (type 
in USNM). 


255 


Foop prant: Unknown. 
Known only from the unique male type. 


148. Genus Sigelgaita Heinrich 


Sigelgaita Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 382, 1939. 
(Type of genus: Sigelgaita chilensis Heinrich.) 

Tongue well developed. Antenna of male bipectinate 
(in transilis with a few flattened setae on the inner row 
of pectinations of the first five or six segments of the 
shaft); antenna of female shortly pubescent. Labial 
palpus of male upcurved, of female porrect (the second 
segment obliquely upturned, the third bent forward). 
Maxillary palpus large, extending above front, flam- 
boyant. Hind wing with veins 7 and 8 anastomosing 
beyond cell; 3 and 5 stalked. Eighth abdominal seg- 
ment with two pairs of thin hair tufts (very slight in 
chilensis). 

Male genitalia with apical process of gnathos small, 
bifid; harpe with apex oblique; vinculum long; anellus 
with base of plate somewhat broadly sclerotized, arms 
moderately long, slightly twisted (in transilis) ; aedeagus 
moderately stout, long; penis weakly scobinate. 

Female genitalia with signum weak or absent; bursa 
small and finely scobinate; ductus bursae moderately 
long, finely scobinate toward bursa and genital opening; 
ductus seminalis from middle of bursa. 

Larva blue or blue-green according to Dodd; not 
banded or conspicuously spotted; two setae in group 
VII on abdominal segments 7 and 8. 

The larvae are solitary feeders in the fruits of Zuly- 
chnia, Trichocereus, and Platypuntia. 

Egg and egg-laying habits unknown. 

This genus is closest to Amalafrida but in many 
characters more nearly resembles Nanaia. The max- 
illary palpi are long in both Sigelgaita and Nanaia but 
are not so closely appressed to the face in the former as 
in the latter. The labial palpi of the males (upcurved 
in Sigelgaita, porrect in Nanaia) readily separate the 
two genera. 

527. Sigelgaita chilensis Heinrich 
Fieures 554, 1046 
Sigelgaita chilensis Heinrich, Proc. U. 8S. Nat. Mus., vol. 86, p. 
383, 1939. 

Palpus, head, and thorax fuscous, strongly irrorated 
with white; head and collar more whitish than fuscous; 
posterior margin of thorax shaded with black. Fore- 
wing fuscous, dusted with white, giving the wing an 
ashy gray (in some specimens a bluish gray) color; a 
white suffusion filling the cell; antemedial line near 
middle of wing, black, outwardly angulate; from upper 
angle of cell to middle of inner margin a more or less 
prominent blackish shade; subterminal band dentate, 
consisting of a thin, black, inner line, a parallel outer 
black line, and a central pale line, the dentations of the 
outer line acute and extended in short dashes onto the 
veins; a row of black dots along termen between the 
vein ends. Hind wing whitish, smoky fuscous toward 
termen, apex, and costa, and on the veins; cilia white 


256 
with a pale fuscous subbasal line. Alar expanse, 31-44 
mm. 

Male genitalia with harpe fairly broad, aedeagus 
somewhat stouter than in ¢ransilis, vinculum shorter. 
Female genitalia with signum present, the latter con- 
sisting of three or four minute, more or less coalesced, 
blunt spines. 

TypPE Locauity: Ovalle, Chile (type in USNM). 

Foop piant: Hulychnia acida Philippi, Trichocereus 
chiloensis (Colla). 

Distrizution: Curie: La Serena, Ovalle, Tofa. 

Superficially this species and huanucensis resembles 
Nanaia substituta. The latter, however, lacks alto- 
gether the dark shade between the outer angle of the 
cell and the inner margin so characteristic of chilensis 
and huanucensis. The forewings of the three species 
are similar—long, and narrow and of about the same size 
and shape. 

528. Sigelgaita huanucensis Heinrich 
Ficure 1047 
Sigelgaita huanucensis Heinrich, Proc. U. S. Nat. Mus., vol. 86, 
p. 383, 1939. 

Similar to chilensis, except as follows: Paler, white 
dusting on head, thorax, and forewing more pro- 
nounced; general color of forewing brownish rather than 
gray; transverse dark shade from outer upper angle of 
cell to inner margin pale brown; antemedial and sub- 
terminal lines interrupted, the latter indicated only by 
blackish scaling on the veins; a pale brownish shade in 
area bordering inner margin. Hind wing pure white, 
with a very faint fuscous shade along costa and a 
thin, pale fuscous line on termen for a short distance 
from apex, this line a trifle broader on female than on 
male and extending nearly to anal angle of wing. Alar 
expanse, 45 mm. 

Female genitalia without signum. 

TypE LocaLity: Hudnuco, Peri (type in USNM). 

Foop prant: Opuntia (Platypuntia) ficus-indica (Lin- 
naeus). 

Known only from the female type and male paratype 
from the type locality. 


529. Sigelgaita transilis Heinrich 
Figure 553 
Sigelgaita transilis Heinrich, Proc. U. S. Nat. Mus., vol. 86, 
p. 384, 1939. 

Palpi, head, thorax, and forewing dark grayish 
fuscous finely peppered with white, giving them a slate- 
gray color. Forewing with antemedial and subterminal 
lines obscured, indicated by faint whitish lines bordered, 
for a short distance from costa, by blackish streaks; 
discal spot at end of cell blackish, rather large; a row of 
black dots along termen between the vein ends. Alar 
expanse, 26 mm. 

Genitalia with harpe narrower than that of chilensis, 
vinculum considerably longer, and aedeagus slenderer 
and appreciably tapering toward apex. 

TYPE Locatity: Santa Eulalia, Pera (type in USNM). 

Foop puant: Trichocereus sp. 

Known only from the male type. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


149. Genus Amalafrida Heinrich 


Amalafrida Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 385, 1939. 
(Type of genus: Cactoblastis leithella Dyar.) 

Tongue well developed. Antenna of male bipec- 
tinate; on each of the inner pectinations of the first five 
segments a row of from three to five flattened, spinelike 
setae; antenna of female simple and finely pubescent. 
Labial palpus of male obliquely ascending, of female 
obliquely porrect. Maxillary palpus squamous. Hind 
wing with veins 7 and 8 anastomosing for over half 
their lengths beyond cell; veins 3 and 5 stalked. Eighth 
abdominal segment of male with two pairs of ventro- 
lateral hair tufts. 

Male genitalia with apical process of gnathos bifid; 
harpe with apex oblique; vinculum long; anellus with 
base of plate rather broadly sclerotized, arms long, 
slightly twisted; aedeagus long, moderately stout; 
Penis scobinate. 

Female genitalia without signum; bursa copulatrix 
large, weakly and scatteringly scobinate; ductus bursae 
long, slender; ductus seminalis from about middle of 
bursa. 

Larva grayish in color with a tendency toward pale 
transverse bands‘after the manner of Olycella larvae, 
according to Dodd; solitary tunnelers in Platypuntia. 

Eggs unknown. 

The genus at present is represented by only the type 
species. It is closest to Sigelgaita, one species of which 
(transits) also has setiferous pectinations on some of 
the basal segments of the male antennal shaft. The 
form of the maxillary palpi, as well as the shape of the 
forewings, distinguish the two genera. In Sigelgaita 
the forewing is much longer in proportion to its width 
and the termen more rounded than is the case in 
Amalafrida. According to Dodd, leithella differs mark- 
edly from the species of Sigelgaita in larval and pupal 
habits. 

530. Amalafrida leithella (Dyar) 
Ficures 555, 1044 

Cactoblastis leithella Dyar, Proc. Ent. Soc. Washington, vol. 30, 
p. 135, 1928. 

Amalafrida leithella (Dyar) Heinrich, Proc. U. S. Nat. Mus., vol. 
86, p. 386, 1939.—Dodd, Biological campaign against 
prickly-pear, Brisbane, Australia, p. 40, 1940. 

Forewing with the areas between cell and costa, be- 
tween vein 1b and inner margin for a short distance, 
and along costal half of termen, white with a scattering 
of black scales; ground color of remaining areas ocherous 
fuscous, very faintly shaded with rufous above inner 
margin; transverse and discal markings black; ante- 
medial line incomplete, indicated by a thin, blackish, 
irregular line from inner margin to cell and a broad 
black streak from costa to about middle of the fold (in 
some specimens this fuses with a black streak, which 
extends from middle of vein 1b to end of cell) ; subter- 
minal line black, faint (obscured below vein 6 in some 
specimens), sinuate and dentate, outwardly bordered 
by a whitish line and beyond this by a second, very 
faint, parallel, pale fuscous line; at end of cell a large, 
irregular, black spot; a line of distinct black dots along 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


termen between the vein ends. Hind wing of male 
white, semihyaline, with a narrow pale fuscous shade 
along costa and termen; of female dark smoky fuscous 
shading to white towards base; cilia white with a 
fuscous basal band. Alar expanse, 30-33 mm. 

Male genitalia as given for the genus. Female geni- 
talia with bursa very large and irregularly shaped, 
minutely scobinate. 


Type LocaLity: Curacao, Dutch West Indies (type 
in USNM). 

Foop piant: Opuntia (Platypuntia) sp. 

Distrisution: Dutcu West Inpres: Curacao (Jan.). 
VENEZUELA. Caracas (Jan.). Coxomsta: Province of 
Colombia (Jan.) 

Superficially leithella resembles Cactoblastis cactorum 
but is easily distinguished on structural characters of 
the male and female genitalia and of the male antennae. 


150. Genus Ozamia Ragonot 


Ozamia Ragonot, Monograph, pt. 2, p. 34, 1901.—Heinrich, Proc. 
U. S. Nat. Mus., vol. 86, p. 386, 1939. (Type of genus: 
Trachonitis lucidalis Walker.) 


Antenna of male serrate (except in hemilutella and 
punicans, where it is simple) and pubescent with a series 
of modified, papillalike setae on the inner side of several 
basal segments of the shaft (fig. 557d); antenna of the 
female simple and pubescent. Labial palpi obliquely 
ascending in both sexes. Maxillary palpus squamous. 
Hind wing with veins 7 and 8 anastomosing beyond the 
cell; 3 and 5 stalked. Eighth abdominal segment bear- 
ing one pair or two (odiosella) pairs of ventrolateral 
hair tufts. 

Male genitalia with apex of gnathos small or moder- 
ately large, bifid; apex of harpe oblique (except in 
punicans); vinculum long; anellus with base of plate 
broadly rather than narrowly sclerotized, arms long, 
slightly twisted and curved; aedeagus rather long and 
moderately stout (except in lucidalis); penis scobinate. 

Female genitalia with signum weak or absent (luci- 
dalis), when present developed as a thin, short, scobi- 
nate or shortly thorned plate or a series or cluster of 
small, weak spines; bursa copulatrix minutely scobinate, 
at least toward ductus bursae (wrinkled in the South 
American species); ductus bursae long or moderately 
long, scobinate toward bursa; ductus seminalis from 
bursa near signum. 

Larva wine colored, olive green, or blackish, not 
banded or conspicuously spotted; with two setae in 
group VII of abdominal segments 7 and 8; solitary 
feeders in fruits and flower buds of Opuntia and Cereus, 
sometimes (some South American species) in the stems 
of Cereus. 

This genus divides into two natural groups: The 
North American species with unwrinkled bursa and 
minutely scobinate ductus bursae, and all fruit or bud 
feeders; and the South American species with wrinkled 
bursa and coarsely scobinate ductus bursae and either 
fruit or stem feeders. The West Indian species 
(lucidalis) is anomalous in some genitalic characters 

300329—56——18 


257 


(small abdominal tufts, rather slender aedeagus, long 
ductus bursae, and no signum), but on habitus and 
other characters it appears closely allied to the North 
American group. When males of all the species are 
known it may be possible to give a separate generic 
designation to the South American forms, but in the 
absence of definitive male characters that does not seem 
justified. 

The papillalike setae on the male antennal shaft of 
Ozamia also occur in Cactobrosis and Zophodia, but 
the last two genera are distinguished by filiform maxil- 
lary palpi. 

Eight species are here recognized as belonging to the 
genus. Its distribution appears to be the southwestern 
part of the United States, Central and South America, 
and the West Indies. 


531. Ozamia lucidalis (Walker) 
Ficures 556, 1042 


Trachonitis lucidalis Walker, List, vol. 27, p. 39, 1863. 

Ozamia lucidalis (Walker) Ragonot, Monograph, pt. 2, p. 34, 
1801.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 388, 
1939.—McDunnough, Check list, No. 6290, 1939. 


Palpi, head, thorax, and forewing sordid white. 
Forewing with ferruginous-fuscous spottings on the area 
bordering inner margin; transverse markings black, 
shading to ferruginous fuscous toward inner margin; 
antemedial line angulate, white, bordered on inner and 
outer sides by black or ferruginous, the outer black 
marking at costa a broad spot; subterminal line dentate, 
slanting from costa near apex to outer fourth of inner 
margin, bordered inwardly and outwardly by dark lines, 
shading from black to ferruginous; discal spot at end of 
cell irregular, frequently extended beyond cell into two 
short dashes, black; a row of black dots along termen 
at the vein ends. Hind wing white, semihyaline, with 
a fine fuscous line along termen; cilia white with a faint, 
dark, subbasal line. Abdominal tufts small. Alar 
expanse, 25-30 mm. 

Male genitalia with apical process of gnathos small; 
end of vinculum bluntly rounded. Female genitalia 
without signum; bursa and part of ductus bursae 
minutely scobinate; ductus bursae long, slender, bent 
at middle. 

TypPE Locatity: Santo Domingo [Dominican Repub- 
lic] (type in BM). 

Foop puant: Opuntia (Platypuntia) sp. 

Distripution: Cusa; JAmMarca, Kingston (Jan.). 

I have seen no specimens from the type locality. 


532. Ozamia fuscomaculella (Wright) 
Fiaures 557, 1043 


Euzophera fuscomaculella Wright, Ent. News, vol. 27, p. 27, 
1916.—MceDunnough, Check list, No. 6320, 1939. 

Ozamia heliophila Dyar, Ins. Inse. Menstr., vol. 18, p. 222, 1925. 

Ozamia odiosella fuscomaculella (Wright) Heinrich, Proc. U. S. 
Nat. Mus., vol. 86, pl 390, 1939. 


Ground color and markings of forewing similar to 
those of lucidalis except that transverse markings are 
blackish throughout, paling somewhat toward inner 


258 


margin but not shading into ferruginous; no ferruginous 
coloring on forewing. Abdominal tufts much stronger 
than in lucidalis and in two distinct pairs. Alar 
expanse, 23-28 mm. 

Male genitalia with apical process of gnathos rather 
large; terminal margin of vinculum bluntly angulate. 
Female genitalia with signum a thin short line of 
minute spines; bursa copulatrix smooth except toward 
ductus bursae, where it is finely scobinate; ductus 
bursae of moderate length, swollen toward and gradually 
expanding into bursa. 

TYPE LOcALITIES: San Diego, Calif. (fuscomaculella, 
in W. S. Wright Collection); Los Angeles, Calif. (helio- 
pila, in USNM). 

Foon pian: Opuntia (Platypuntia) spp. 

Disrripution: California, San Diego (May, June, 
Aug.), Los Angeles (July), Pasadena (Aug.). 


533. Ozamia fuscomaculella clarefacta Dyar 


Fieures 67, 559, 1040 


Ozamia clarefacta, Dyar, Ins. Inse. Menstr., vol. 7, p. 55, 1919— 
McDunnough, Check list, No. 6292, 1939. 

Ozamia odiosella Heinrich (not Hulst), Proc. U. 8. Nat. Mus., 
vol. 86, p. 389, 1939.—Dodd, Biological campaign against 
prickly-pear, Brisbane, Australia, pp. 39, 60, 84, 1940. 

The variety (or race) is apparently limited to south- 
eastern Texas and eastern México. It is distinguished 
by the presence of a greenish tint over the area bordering 
the inner margin of forewing. According to Dodd this 
green shade is quite distinct on live or freshly killed 
moths, but is not observable on most pinned examples. 
There is also a slight difference in the female signum. 
In clarefacta it consists of a narrow, minutely spined 
plate. Otherwise clarefacta has nothing to distinguish 
it from typical fuscomaculella. 

TYPE LOCALITY: Orizaba, México (type in USNM). 

Foop PLants: Opuntia (Platypuntia) spp. 

Distrisution: Unirep Starrs: Texas, Brownsville, 
Victoria (May), Burnet County (Oct.), Uvalde (June, 
July), Kerrville (May, June), San Benito (Aug.). 
Mexico: Orizaba (Apr.), Jalapa. 

In my 1939 paper I misidentified this variety with 
Hulst’s odiosella on the basis of a female from Texas in 
the National Collection which Hulst had labeled ‘‘Saleb- 
ria odiosella Hulst, Type” and which both Dyar and I 
had supposed was the true type of that species. It is 
not the actual type, which I had previously overlooked. 
This is in the Rutgers Collection. It is a male, super- 
ficially similar to the spurious ‘‘type”’ in the National 
Museum but structurally quite distinct; it has 8 veins in 
the hind wing and the maxillary palpus in the form of an 
aigrette (as in Salebria where Hulst placed it). The 
true odiosella is treated on page 114. 


534. Ozamia thalassophila Dyar 


Fiaure 1039 


Ozamia thalassophila Dyar, Ins. Insc. Menstr., vol. 18, p. 15, 
1925.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 391, 
1939.—McDunnough, Check list, No. 6291, 1939. 


Palpi, head, thorax, and forewing dark grayish 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


fuscous, lightly sprinkled with white. Forewing with 
some white dusting on costal half and a slightly more 
brownish shade on inner half; antemedial and subter- 
minal transverse lines whitish, bordered with black or 
blackish fuscous, the pattern as in lucidalis and odio- 
sella; discal spot at end of cell curved, black; between 
this and subterminal lines one or two small, obscure, 
blackish dots; upper and lower veins of cell faintly out- 
lined by white scales; a row of black dots along termen 
at or close to the vein ends; cilia pale ocherous fuscous. 
Hind wing white with a narrow fuscous shade along 
termen; cilia white, with a fuscous subbasal line. Alar 
expanse, 28 mm. 

Female genitalia, with signum a small cluster of two 
or three more or less fused and minute spines; bursa 
copulatrix partially wrinkled (im the region of the sig- 
num); ductus bursae of moderate length, finely sco- 
binate toward bursa. 

TYPE LocaLity: Oceanside, Calif. (type in USNM). 

Foop piant: Opuntia (Cylindropuntia) sp. 

Known only from the unique female type, reared 
(Aug. 1924) from larva in a Cylindropuntia, presumably 
feeding in the fruit. 


535. Ozamia immorella (Dyar), new combination 


Fieure 1036 
Euzophera immorella Dyar, Proc. U. 8. Nat. Mus., vol. 44, p. 322, 
1913. 

Similar in color and markings to thalassophila except 
for a faint purplish red suffusion over the ground color 
of forewing (especially noticeable on freshly reared 
examples) and less white dusting on costal area of wing. 
Alar expanse, 25-31 mm. 

Male genitalia with apical process of gnathos rather 
large (as in fuscomaculella); terminal margin of vin- 
culum bluntly rounded. Female genitalia with signum 
a narrow, bluntly spined plate, the signum longer than 
any other in the genus; bursa copulatrix somewhat 
wrinkled in the neighborhood of signum, finely scobi- 
nate towards ductus bursae, the scobinations extending 
for a short distance into ductus. iq 

Typr Locatity: Tehuacén, México (type in Oe 

Foop PLant: Opuntia (Platypuntia) sp. 

Distripution: México: Caxaxa, Tehuacin (uly), 
Zacatecas (Dec.) 

This species is very close to thalassophila but appar- 
ently distinct. I overlooked it when treating the 
cactus-feeding Phycitinae in 1939. Since that time a 
series (including one male) was reared on Dec. 10, 1946, 
by George Callaghan of the U.S. Bureau of Entomology 
and Plant Quarantine from larvae feeding in the fruits 
of prickly-pear at Zacatecas, México. 


536. Ozamia stigmaferella Dyar 


Fieure 1038 
Ozamia stigmaferella Dyar, Ins. Insc. Menstr., vol. 10, p. 17, 
1922.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 391, 
1939. 
Palpi, head, and thorax dark grayish fuscous sparsely 
sprinkled with white, forewing dark grayish fuscous; 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


extreme base black; remainder of basal area and area 
between cell and costa and to the subterminal line 
white lightly dusted with black; antemedial line obso- 
lete, indicated only by a large black spot on costa 
(corresponding to the black shade outwardly bordering 
the white antemedial line in the North American 
species of Ozamia); subterminal line whitish, dentate, 
bordered inwardly and outwardly by black; a short, 
dentate, black line from vein 8 to vein 2, midway 
between the end of cell and the subterminal line and 
parallel with the latter; a black curved mark at end of 
cell; a row of black dots along termen at or near the 
vein ends; a faint whitish color dusted with black in 
apical area. Hind wing white, semihyaline, with a 
fuscous shade at apex and for a short distance along 
termen; cilia whitish with a very faint fuscous subbasal 
line. Alar expanse, 26 mm. 

Female genitalia with signum a small buttonlike 
thorn; bursa copulatrix wrinkled and finely scobinate; 
ductus bursae long, coarsely scobinate toward bursa. 


Type tocatity: Catamarca, Argentina (type in 
USNM). 

Foop piant: Cereus validus Haworth. 

This species is known only from the female type, 
reared Mar. 7, 1921, by W. B. Alexander from a larva 
that had excavated a hollow in a stem of Cereus validus. 

O. stigmaferelia and the two following species (hemilu- 
tella and punicans) are the South American repre- 
sentatives of the genus and differ from those from the 
United States, México, and the West Indies in having 
the ductus bursae of the female coarsely scobinate and 
the bursa copulatrix decidedly wrinkled. 


537. Ozamia hemilutella Dyar 


Fiaures 560, 1041 
Ozamia hemilutella Dyar, Ins. Insc. Menstr., vol. 10, p. 17, 
Pdi ta Proc. U. S. Nat. Mus., vol. 86, p. 392, 
1939, 

Palpi, head, and thorax brownish fuscous, finely and 
evenly sprinkled with white, making the general color 
(to the naked eye) pale gray; collar of thorax with a 
slight yellowish tint. Forewing with area between 
lower vein of cell, vein 2, and inner margin pale yellow 
without any markings; remainder of wing pale gray, 
concolorous with head and thorax; antemedial line 
obsolete; from costa just before middle to middle of 
lower vein of cell, a rather broad, transverse, brownish 
shade; a brown discal dot at end of cell and some brown 
shading just beyond; subterminal line faint, weakly 
dentate, parallel with termen, bordered inwardly and 
outwardly by thin faintly brownish lines; a row of 
minute black dots along termen at the vein ends; cilia 
pale gray. Hind wing white, semihyaline with a thin 
fuscous line along termen and some fuscous shading 
bordering the costa. Abdomen with one pair of strong 
tufts. Alar expanse, 27-30 mm. 

Male genitalia with apical process of gnathos small; 
end of vinculum bluntly angulate. Female genitalia 
with signum a single, weak, irregular, thornlike patch; 


259 


bursa copulatrix wrinkled and finely scobinate; ductus 
bursae coarsely scobinate toward bursa. 

Type ocauity: La Rioja, Argentina (type in 
USNM). 

Foop piant: Cereus validus Haworth. 

DistRIBUTION: ARGENTINA: La Rioja, Santiago del 
Estro. 


The moth is easily identified by the clear yellow inner 
area of the forewing. The larvae feed in the fruits and 
flower buds and, possibly, to some extent, in the stems 
of Cereus. Dodd states that they also attack fruits of 
Platypuntias. 


538. Ozamia punicans Heinrich 


Fieures 558, 1037 


Ozamia punicans Heinrich, Proc. U. S. Nat, Mus., vol. 86, p. 393, 
939. 

Palpi fuscous sprinkled with white. Head and thorax 
fuscous heavily dusted with white and more or less 
shaded with pale rust color, especially on top of head 
and on collar of thorax. Forewing pale gray (fuscous 
heavily dusted with white) marked with darker gray 
and with large blotches of pale rust color; the rust shade 
filling about one-fourth of the basal area and nearly all 
the area between antemedial and subterminal lines, 
lower vein of cell, vein 2, and vein 1b; antemedial line 
obscure, indicated chiefly by a rather broad dark gray 
shade from costa to lower vein of cell and a thin dark 
gray line thence to inner margin; subterminal line faint, 
somewhat sinuate but not dentate, approximately paral- 
lel with termen, bordered inwardly and outwardly by 
obscure dark gray; apical mark at end of cell irregular, 
dark gray; between cell and subterminal line some faint 
rust shading in the interspaces between the veins; a row 
of black dots along termen between the vein ends; cilia 
pale rust red. Hind wing white, semihyaline, with a 
fuscous shade in costal area to top of cell and vein 8, 
some fuscous shading on the vein ends, and a fine fuscous 
line along termen to vein 1b; cilia shiny white. Abdom- 
inal tufts as in hemilutella. Alar expanse, 36-40 mm. 

Male genitalia with apical process of gnathos moder- 
ately large; apex of harpe more rounded than in other 
species of Ozamia; end of vinculum more rounded than 
angulate and lateral margins excavate; penis bearing a 
number of coarse spines. Female genitalia larger than 
those of any other Ozamia; with signum a small granu- 
lose plate containing a stubby central thorn; bursa 
copulatrix wrinkled and finely scobinate; ductus bursae 
long, very coarsely scobinate toward bursa. 

TyprE LocaLity: Tapia, Tucumdn, Argentina (type 
in USNM). 

Foop piant: Cereus validus Haworth. 

According to Dodd, punicans differs from other 
species of Ozamia in that it is a stem borer and appar- 
ently does not attack the fruits or flower buds. It 
differs also in that the apex of the harpe is not definitely 
oblique, and the maxillary palpi are somewhat narrowly 
scaled. However, the latter are of the squamous 
rather than the filiform type, and from its general 
habitus the species is obviously closely related to 


260 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


hemilutella. The moth can be easily identified by the 
rust-red cilia and blotches on the forewing. It, so far, 
is known only from the type series from Tucumdén. 


151. Genus Cactobrosis Dyar 


Cactobrosis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 406, 1915; 
Proc. Ent. Soc. Washington, vol. 30, p. 135, 1928.—Hein- 
rich, Proc. U.S. Nat. Mus., vol. 86, p. 394, 1939. 

Tongue well developed. Antenna of male with a 
series of modified, papillalike setae on the inner sides of 
several basal segments of the shaft, bipectinate (fer- 
naldialis, longipennella) orstrongly serrate and pubescent 
(maculifera, strigahis); antenna of female simple and 
shortly pubescent. Labial palpus upturned in the 
male, oblique in the female. Maxillary palpus filiform. 
Hind wing with veins 7 and 8 anastomosing beyond 
the cell; 3 and 5 shortly stalked. Highth abdominal 
segment bearing a pair of ventrolateral hair tufts (the 
tufts long and dense except in strigalis). 

Male genitalia with apex of gnathos large, bifid; apex 
of harpe evenly rounded; vinculum long (moderately 
long in strigalis); anellus with base of plate narrowly 
sclerotized, arms long, slender, slightly twisted ; aedeagus 
long, stout (shorter and less stout in sfrigalis); penis 
more or less densely pubescent (armed with short, hair- 
like spines). 

Female genitalia without sisnum; ductus bursae long, 
finely scobinate only at genital opening or (in séfrigalis 
only) sparsely so at junction of bursa copulatrix and 
ductus bursae, with two small sclerotized dorsal plates 
and a single ventral plate at genital opening (the ventral 
plate absent in strigalis); bursa copulatrix large, smooth 
(except in strigalis, in which it has a few minute scobina- 
tions) ; ductus seminalis from near end of bursa. 

Larvae bluish, not banded or conspicuously spotted; 
with two setae in group VII on abdominal segments 7 
and 8; gregarious feeders in Ferocactus, Echinocereus, 
Peniocereus, and, probably, Carnegiea. 

Eggs laid singly. 

The genus as here defined is distinguished from all] 
other genera of the cactus-feeding group by its filiform 
maxillary palpi. Zophodia, which it resembles in most 
structural characters, is not a cactus-feeding genus, has 
the male antenna unserrate, the labial palpus of female 
porrect, and a small signum in the bursa, copulatrix. 

Five species are recognized as belonging to the genus. 
They are fairly easy to distinguish but subject to much 
individual variation in wing markings. 

The known distribution is the southwestern part of 
the United States and México. 


539. Cactobrosis fernaldialis (Hulst) 
Fiaures 73, 524, 1049 


Megaphycis fernaldialis Hulst, Trans. Amer. Ent. Soc., vol. 13, 
p. 163, 1886. 

Euzophera gigantella Ragonot, Nouv. Gen., p. 32, 1888; Mono- 
graph, pt. 2, p. 51, 1901.—Dyar, Proc. Ent. Soc. Washing- 
ton, p. 158, 1904. 

Melitara fernaldialis (Hulst), Phycitidae of N. Amer., p. 172, 
1890; U. S. Nat. Mus. Bull. 52, p. 429, 1903.—Schwarz, 


Psyche, vol. 8, p. 15, 1901.—Hunter, Pratt, and Mitchell, 
U.S. Dept. Agr. Bur. Ent. Bull. 113, p. 29, 1912. 

Honora cinerella Hulst, Journ. New York Ent. Soc., vol. 8, 
p. 223, 1901; U. S. Nat. Mus. Bull. 52, p. 433, 1903. 

Melitara fernaldalis Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 36, 1905 (misspelling for fernaldialis Hulst). 

Cactobrosis fernaldalis (Dyar), Proc. U. S. Nat. Mus., vol. 47, 
p- 407, 1914; Ins. Insc. Menstr., vol. 13, p. 223, 1925 (in 
part); Proc. Ent. Soc. Washington, vol. 30, p. 135, 1928 
(in part). 

Cactobrosis fernaldialis (Hulst) Heinrich, Proc. U. S. Nat. Mus., 
vol. 86, p. 396, 1939—McDunnough, Check list, No. 
6283, 1939.—Dodd, Biological campaign against prickly- 
pear, Brisbane, Australia, p. 40, 1940. 

Mats: Antenna bipectinate. Palpi, head, and thorax 
grayish fuscous dusted with white. Forewing grayish 
fuscous dusted with white and more or less blotched 
with black; some specimens with a faint ocherous- 
fuscous tint in the middle of the cell and on the area 
between vein 1b and the cell; normally with antemedial 
and subterminal transverse markings indistinct, but 
indicated by whitish angulate and dentate bands shaded 
inwardly and outwardly by black; a blackish shade at 
end of cell, often extending to costa; below it on inner 
margin a similar dark spot; veins 2 to 8 faintly lined 
with black and in many specimens the fold to a little 
beyond its middle. Hind wing white, semihyaline, 
shaded in costal area above vein 6 and cell with pale 
fuscous, with some fuscous scaling on the veins and a 
fine fuscous line along termen; anal margin and adjoin- 
ing cilia faintly ocherous; cilia otherwise white, with a 
narrow, fuscous subbasal line. Alar expanse, 36—47 
mm 


Male genitalia essentially like those of longipennella 
but somewhat larger, in size and habitus like those of 
maculifera. 


Fematz: In color and markings like the male except 
that there is never any black streak on the fold of the 
forewing; some specimens are heavily dusted with black 
over the entire base of the forewing as far as the ante- 
medial line; others have the transverse lines and con- 
trasted dark spots almost obliterated and the wing of a 
pale slate color with only the faintest remnants of the 
normal markings. Alar expanse, 34-50 mm. 

Female genitalia with the sclerotized ventral plate 
in ductus bursae at opening smaller than those in 
longipennella, insignatella, and maculifera. 

TYPE Locauities: Arizona (fernaldialis, in AMNH, 
ex Rutgers; gigantella, in Paris Mus.); Santa Rita Mts., 
Ariz. (cinerella, in USNM). 

Foop piants: Ferocactus wislizent (Engelmann) and 
probably other species of Ferocactus; Peniocereus greggu 
(Engelmann) ; Carnegiea gigantea (Englemann) ; Homalo- 
cephala texensis (Hopffer). 

Distrisution: Arizona, Catalina Springs (Apr.), 
Oracle (July), Tuscon (June), Baboquivari Mts. (Apr., 
May, June, July, Aug., Oct., Nov.), Christmas, Reding- 
ton, Pinal Mts., Santa Rita Mts. (May, June), Hua- 
chuca Mts. (Aug.), Douglas (Apr., May), Mohave 
County (May), Sells Post Office (Indian Oasis, Apr.), 
Dewey (June), Maricopa County (July), “en route from 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 261 


Dewey to Salome’ (Apr., May); California, San Diego 
(May, Oct.); Texas, Brownsville (Jan., U. S. Dep. Agr. 


rearing). 


540. Cactobrosis longipennella (Hampson) 
Fiaures 523, 1050 


Euzophera longipennella Hampson, in Ragonot, Monograph, pt. 
2, p. 52, 1901. 

Moodna elongatella Hampson, in Ragonot, Monograph, pt. 2, p. 
269, 1901. 

Cactobrosis longipennella (Hampson) Dyar, Proc. U.S. Nat. Mus., 
vol. 47, p. 407, 1914.—Heinrich, Proc. U. 8. Nat. Mus., vol. 
86, p. 397, 1939. 

Cactobrosis elongatella (Hampson) Dyar, Proc. U. 8. Nat. Mus., 
vol. 47, p. 407, 1914. 

Cactobrosis fernaldialis Dyar (part), Ins. Insc. Menstr., vol. 13, 
p. 223, 1925; Proc. Ent. Soc. Washington, vol. 30, p. 135, 
1928. 

Mats: Like that of fernaldialis except that pectina- 
tions of antenna are about half the length of those on 
fernaldialis; transverse pale markings on forewing obso- 
lete in some specimens. Alar expanse, 34-40 mm. 

Male genitalia similar to those of fernaldialis except 
smaller; harpe not so markedly creased. 


Frmate: Similar in color and markings to the female 
of fernaldialis. Alar expanse, 33-43 mm. 

Female genitalia with sclerotized ventral plate in 
ductus bursae at genitalic opening larger and the oppos- 
ing small plates on the dorsal wall of the ductus 
narrower than those of fernaldialis. 


Typr tocauities: Tres Marfas Isl., México (longi- 
pennella, in BM); Orizaba, México (elongatella, in 
USNM). 

Foop puant: Unknown, probably Ferocactus. 

Distrisution: México: Orizaba, Oaxaca, Tehuacén 
(June), Cuernavaca (June, July), Zacualpin (Mar., 
Oct.). 

I have seen no examples from the type locality of 
longipennella. 


541. Cactobrosis maculifera Dyar 
Ficures 525, 1051 


Cactobrosis maculifera Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 
407, 1914; Proc. Ent. Soc. Washington, vol. 30, p. 136, 
1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 398, 1939. 

Mate: Antenna strongly serrate and fasciculate. 

Palpi, head, and thorax pale clay color (‘‘luteous’’). 

Forewing luteous gray shaded and spotted with dark 

grayish fuscous, the luteous tint pronounced on basal 

third of costa and over the submedian area of the wing; 
transverse antemedial and subterminal lines obsolete; 

a fuscous shade from costa before middle to cell, another 

from costa at middle, and, below these, corresponding 

streaks or spots on lower vein of cell and on vein 1b; 

a thin blackish line on the fold from its base to near its 

middle; a similar dark streak on vein 1b at outer third; 

short, broken, dark streaks on the veins at or near the 
cell; a clouded fuscous spot at end of cell; outer half of 
costa shaded with fuscous; a row of dark spots along 
termen at or very close to the vein ends. Hind wing 
white, semihyaline with only the faintest indication of 


a fuscous line on termen toward apex. Alar expanse, 
32-45 mm. 

Male genitalia agreeing in size and nearly all details 
with those of longipennella except that the arms of the 
anellus are a trifle longer in maculifera. 

Frmate: In color and pattern like the male except 
that there is some gray shading on the head and thorax 
and considerably more gray on the forewing (the single 
specimen before me is in much better condition than 
the males, which may account for some of the differ- 
ences); basal third of wing clouded with dark fuscous; 
terminal area more faintly clouded; subterminal line 
faintly indicated, sharply angulate at middle, broken 
below; the luteous shade more contrasted than in the 
male, but restricted to middle of cell and the area be- 
tween veins 1b and the fold. Hind wing white, semi- 
hyaline with a narrow fuscous shade along termen and 
on the veins near their apices. Alar expanse, 37 mm. 

Female genitalia similar to those of insignatella but 
with sclerotized ventral plate in ductus bursae at gen- 
ital opening smaller. 

TypE LocaLity: Oaxaca, México (type in USNM). 

Foop prant: Unknown. 


Distrisution: M*£xico: 
(Sept.). 

This species may be distinguished from other species 
of Cactobrosis by the strong luteous (pale clay) shade 
on the forewing and the serrate-fasciculate male 
antenna. 


Oaxaca, Salina Cruz 


542. Cactobrosis insignatella Dyar 
FicureE 1052 


Cactobrosis insignatella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 
407, 1914; Proc. Ent. Soc. Washington, vol. 30, p. 136, 
1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 399, 1939. 


Mate: Unknown. 


Frema.e: Palpi, head, thorax, and forewing of a soft, 
nearly uniform grayish fuscous (with a more brownish 
than slate-gray tint). Forewing without discal spot or 
transverse dark markings; costa at base very slightly 
paler than ground color of wing, concolorous with collar 
of thorax; an obscure pale shade on midcosta and the 
faintest indication of a pale subterminal line, the latter 
broadly angulate at middle; some faint dark shading 
on the veins from cell to termen and a row of small, 
dark dots along termen near the vein ends. Hind wing 
white, semihyaline, with a pale fuscous line along ter- 
men; cilia white with a narrow, pale fuscous, subbasal 
line. Alar expanse, 37-40 mm. 

Genitalia with the dorsal plates in ductus bursae at 
genital opening strongly sclerotized; ventral plate at 
opening slightly larger than in any of the other species. 


Type Locauity: Oaxaca, México (type in USNM). 
Foop prant: Unknown. 


Known only from the female type and paratype from 
the type locality. These specimens resemble suffused 
specimens of fernaldialis and longipennella except that 
the latter are more slate colored. The slight genitalic 


262 


differences seem to indicate that insignatella is a good 
species and not a mere color form. 


543. Cactobrosis strigalis (Barnes and McDunnough) 
Figures 526, 1053 


Euzophera strigalis Barnes and McDunnough, Canadian Ent., 
vol. 44, p. 127, 1912; Contributions, vol. 1, No. 4, pl. 1, 
fig. 14, 1912. 

Caciobrosis strigalis (Barnes and McDunnough), Check list of 
the Lepidoptera of Boreal America, No. 5697, 1917.—Dyar, 
Ins. Inse. Menstr., vol. 13, p. 224, 1925; Proc. Ent. Soc. 
Washington, vol. 30, p. 136, 1928.—Heinrich, Proc. U. 8. 
Nat. Mus., vol. 86, p. 400, 1989.—McDunnough, Check 
list, No. 6287, 1939. 

Mats: Antenna strongly serrate. Palpi, head, thorax, 
and forewing grayish fuscous sprinkled with whitish 
(the ends of the scales white) making the ground color 
a pale slate-gray. Forewing with the veins outlined in 
black, the strongest black line being that along upper 
vein of cell and vein 6; transverse lines and discal mark 
absent; no dots along termen. Hind wing white, semi- 
hyaline, with a faint fuscous shade bordering costa, and 
a fine fuscous line along termen for a short distance from 
apex. Tufts on eighth abdominal segment weak. Alar 
expanse, 30-43 mm. 


Male genitalia with vinculum moderately long, but 
considerably shorter than in other species of Cactobrosis; 
aedeagus also shorter. 


Frmaue: Similar to the male in color and markings 
except that hind wing is more or less suffused with 
smoky fuscous beyond the base, especially along the 
veins and termen. Alar expanse, 33-44 mm. 

Female genitalia with sclerotized plates on dorsal wall 
of ductus bursae behind the genital opening well de- 
veloped, but with opposing ventral sclerotized plate 
absent, replaced by minute scobinations; ductus bursae 
shorter than in other Cactobrosis species; bursa copula- 
trix not entirely smooth, having a few weak scobinations 
toward ductus. 

TypE LOCALITY: Hureka, Utah (type in USNM). 

Foop piant: Echinocereus rigidissimus (Engelmann), 
E. pectinatus (Scheidweiler), and probably a number of 
other species of H-chinocereus. 


Distripution: Unirep States: Utah, Eureka (Aug., 
Sept.). Dividend (Sept.); Cahfornia, San Gorgonio 
Pass (July); Arizona, Tucson (Apr., July); Texas, Brew- 
ster County (July, Aug.), Alpine (Apr.). Mexico: 
Distrito Federal, México (National University, male 
reared from £. pectinatus, June 3, 1931). 

In a number of respects (its shorter vinculum and 
ductus bursae, its weak abdominal tufts, and its par- 
tially scobinate bursa copulatrix) this species fits badly 
into Cactobrosis. Eventually it may need a separate 
generic designation, but this had better be postponed 
until the life histories of the other species of Cactobrosis 
are more fully known. 

The forewing markings of strigalis resemble those of 
Euremberga leuconips (Dyar). The latter, however, is 
easily distinguished by its squamous maxillary palpi. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Genus 152: Drescomopsis 


[Venational division A. Forewing with 11 veins: 4 and 5 approxi- 
mate or connate. Hind wing with vein 2 from the lower outer 
angle of cell; discocellular vein curved. Male genitalia with 
transtilla complete; harpe with base of sacculus produced and 
bearing stout hair tuft. Labial palpus upturned.] 


152. Genus Drescomopsis Dyar 


Drescomopsis Dyar, Ins. Insc. Menstr., vol. 7, p. 61, 1919. (Type 
of genus: Drescomopsis subelisa Dyar.) 

Tongue well developed. Antenna pubescent; shaft 
of male slightly swollen at base, otherwise simple. La- 
bial palpus upcurved, broadly scaled; reaching a little 
above vertex. Maxillary palpus squamous, appressed 
to face. Forewing smooth except for a projecting tuft 
of rough scales from base of costa in male; 11 veins; vein 
2 from very close to lower outer angle of cell; 3 from the 
angle, shortly separated from 4; 4 and 5 connate or very 
closely approximate at bases; 6 from below upper angle 
of cell, straight; 8 and 9 stalked; 10 from the cell, ap- 
proximate to the stalk of 8-9 for some distance; male 
without costal fold. Hind wing with vein 2 from lower 
outer angle of cell; 3 and 5 shortly stalked; 7 and 8 ap- 
proximate (or contiguous) for a short distance (less than 
half their lengths) beyond cell; cell about one-third the 
length of wing, shorter in male than female; on under 
side of male wing, between cell and vein 1c, an elongate 
pocket enclosing hair tuft and a mass of modified scales; 
discocellular vein curved. Wighth abdominal segment 
of male with a pair of ventrolateral hair tufts and ster- 
nite produced as a sclerotized pocket (projecting into 
abdomen). 

Male genitalia with apical process of gnathos a slen- 
der, elongate, flattened hook. Uncus rounded, hook- 
like. Transtillas complete, thrust anteriorally from 
costal margins of harpes. Harpe broad, apex broadly 
and evenly rounded; clasper appressed to harpe; base of 
sacculus produced and bearing a stout hair tuft. Anel- 
lus a slightly curved, elongate plate. Aedeagus slender, 
smooth, moderately long, sharply bent beyond one- 
third from base; penis without cornuti or other appreci- 
able armature. Vinculum long, narrow, strongly 
arched dorsally, strongly sclerotized only on lateral and 
terminal margins; constricted laterally and flaring to a 
broad lateral margin. 

Female genitalia with signum consisting of a de- 
pressed, tear-shaped patch of short scobinations with a 
sparser scattering of weaker scobinations in surrounding 
area, (signa shown in frontal and lateral views in figs. 
761 and 762); bursa otherwise simple; ductus bursae 
shorter than bursa, simple; genital opening simple; duc- 
tus seminalis from bursa near its junction with ductus 
bursae. 

The genus is very closely related to and apparently 
derived from Drescoma in group I, agreeing with it on 
most important genitalic characters, differing chiefly in 
the absence of vein 4 of hind wing. 

Contains but one tropical American species. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 2 


544. Drescomopsis soraella (Druce) 
Ficures 16, 284, 761, 762 


Homoeosoma soraella Druce, Biologia Centrali-Americana, Lepi- 
doptera Heterocera, vol. 2, p. 565, 1899. 

Drescoma soraella (Druce) Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 328, 1914. 

Drescoma drucella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 328, 
1914 (new synonymy). 

Drescomopsis subelisa Dyar, Ins. Inst. Menstr., vol. 7, p. 62, 1919 
(new synonymy). 

Forewing with costal area above middle of cell and 
from just beyond base to subterminal line white; re- 
mainder of wing a dull purplish fuscous; a rather broad 
glossy dark brown oblique antemedial band dividing 
the white area and extending half way across wing; ex- 
treme base of wing on costal half similarly colored; a 
narrowly lunulate brown patch on midcosta and a simi- 
lar shade extending from apex transversely toward cell, 
broken toward costa by faint indication of a subterminal 
pale line; discal dots small but distinct; under magnifi- 
cation the dark brown markings show an admixture of 
reddish scales. Hind wings dark smoky fuscous, the 
veins and terminal margin slightly darker. Alar ex- 
panse, 14-16 mm. 

Male genitalia with apical process of gnathos sharply 
pointed; clasper a short, sharp spine; transtilla a 
narrow, short, arched band with lateral ends produced, 
pointed, and directed anteriorally. Female genitalia as 
given for the genus. There is considerable individual 
variation in the size of the bursa, but as this is an 
expansible organ (when unsclerotized) its size and shape 
mean little or nothing. 

TypE LocauitiEs: Jalapa, México (soraelia, in BM); 
Porto Bello, Panama (drucella, in USNM); Cayuga, 
Guatemala (swbelisa, in USNM). 

Foop piant: Unknown. 

Distripution: M*f&xico: Cérdoba (May), Jalapa. 
GuatemaLa: Cayuga (Apr., June). Costa Rica: 
Juan Vifias (June, Nov.), Sitio. PanamA: Paraiso 
(May), Porto Bello (Mar., Oct.), Rio Trinidad (Mar.). 
Ecuapor: Zaruma. Braziu: Santa Catarina, Santa 
Catarina Isl. (June). 

Dyar evidently did not look very carefully at his 
male type of drucella or the specimens from Costa Rica 
he had as soraella or he never would have put them into 
a genus with eight veins in hind wing, nor associated 
with his males of drucella some small females of Dres- 
coma cyrdipsa. The true females of his drucella he 
later described as Drescomopsis subelisa. 


Genera 153 and 154: Illatila and Lascelina 


[Venational division A. Forewing with 11 veins; 4 and 5 stalked. 
Hind wing with vein 2 close to or from lower outer angle of cell; 
discocellular vein curved. Male genitalia with transtilla 
absent; apical process of gnathos considerably reduced. Labial 
palpus upturned.] 


153. Genus Illatila Dyar 


Illatila Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 334, 1914. 
of genus: Jllatila gurbyris Dyar.) 


(Type 


Tongue well developed. Antenna pubescent, simple. 
Labial palpus upturned, slender; reaching a little above 
vertex; third segment acuminate, more than half as long 
assecond. Maxillary palpus filiform, appressed to face. 
Forewing smooth; 11 veins; vein 2 from before the lower 
outer angle of cell, well separated from 3; 3 from the 
angle; 4 and 5 short stalked, the stalk separate from 3 
at base; 6 from below upper angle of cell, straight; 8 and 
9 stalked; 10 from the cell, closely approximate to 8-9 
for a short distance from cell; male without costal fold. 
Hind wing with vein 2 from lower outer angle of cell; 
3 and 5 stalked for less than half their lengths; 7 and 8 
anastomosed beyond cell for a trifle less than half their 
lengths; cell short, less than one-third the length of 
wing; discocellular vein curved; on male the inner 
margin thickened and folded with a fringe of hairs 
enclosed within the fold. Eighth abdominal segment 
of male with sternite produced as a sclerotized pocket, 
projecting into abdomen. 

Male genitalia with apical process of gnathos weak, 
straight, very slender (needlelike). Uncus subtriangu- 
late. Transtilla absent (elements not distinguishable). 
Harpe rather narrow, apex evenly rounded; sacculus 
produced at extremity into a short spine; base of 
sacculus hairy, but not produced. Anellus a curved 
plate. Adeagus short, moderately stout; cornuti pres- 
ent, consisting of rather short stout spines in clusters. 
Vinculum stout, triangulate; terminal margin rounded. 

Female genitalia with strong signa, developed as a 
small cluster of blunt, stout, thornlike spines; ductus 
bursae sclerotized throughout, tubular, stout; genital 
opening simple; ductus seminalis from bursa between 
signa and ductus bursae. 

This genus is quite distinct in genitalic characters 
from anything else in group II. Its nearest affinities 
seem to be with Difundella and related genera in group I. 
It contains but one tropical American species. 


545. Illatila gurbyris Dyar 
Ficures 60, 561, 1054 
Illatila gurbyris Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 334, 1914, 


Dyar’s original description is accurate and adequate 
except that the ground color is a rusty brown rather 
than “brownish gray,’’ discocellular vein of forewing 
outlined by blackish scaling with a broader pale outer 
border (Dyar’s “discal spot”). Alar expanse, 12-13 
mm. 

Male genitalia with gnathos weakly attached to 
tegumen at base of uncus, its lateral arms produced 
anteriorly (in relation to head of insect) into widely 
spaced ribbonlike bands. Apex or uncus_ broadly 
rounded. Cornuti consisting of three spine clusters, two 
of them of stout and one (apical) of slender spines. 
Vinculum a trifle longer than broad (foreshortened in 
fig. 561); tapering; terminal margin narrowly rounded. 
Tegumen with lateral margin at base produced into a 
rounded projection with serrate edge. 

Female genitalia with bursa finely scobinate over 
most of its inner surface; ductus bursae bulging and 


264 


heavily rugose near junction with bursa, nearly as long 
as bursa. 

TyPE LocaLity: Taboga Isl., Panamd (Feb.; type in 
USNM). 

Foop prant: Unknown. 

Represented only by the original type series. Hasily 
distinguished from anything else in the family by its 
unique male and female genitalia. 


154. Lascelina, new genus 


Typr or GENus: Lascelina canens, new species. 

Tongue well developed. Antenna pubescent; shaft 
of male with sinus and strong scale tuft at base; of fe- 
male simple. Labial palpus upturned, reaching vertex 
in male, well above vertex in female; slender; rough 
scaled; third segment slightly shorter than second, 
bluntly pointed. Maxillary palpus minute, filiform. 
Forewing smooth; 11 veins; vein 2 from before, but near, 
lower outer angle of cell; 3 from the angle; 4—5 shortly 
stalked, the stalk separate from 3 at base; 6 from below 
upper angle of cell, straight; 8-9 stalked; 10 from the 
cell, approximate to stalk of 8-9 for some distance from 
cell; male without costal fold. Hind wing with vein 2 
from very close to lower outer angle of cell; 3 and 5 
stalked for half their lengths; 7 and 8 anastomosed for 
half their lengths; cell slightly less than half the wing 
length; discocellular vein curved. Highth abdominal 
segment with sternite produced as a sclerotized pocket 
projecting into abdomen. 

Male genitalia with gnathos and its apical process 
greatly reduced, latter fused with subanal plate. Uncus 
triangulate. Transtilla absent (elements not distin- 
guishable). Harpe broad, evenly curved; clasper pres- 
ent as a short erect lobe; a pair of strong, detached hair 
tufts adjacent to bases of sacculi. Anellus a triangu- 
late, slightly curved plate. Aedeagus short, stout; 
penis armed with minute spines and fine scobinations. 
Vinculum long (longer than combined tegumen and 
uncus) ; U-shaped; slightly tapering to rounded terminal 
margin. 

Female genitalia with bursa copulatrix elongate; 
signum present, consisting of a single long stout curved 
spine; ductus bursae very short, sclerotized near junc- 
tion with bursa, the sclerotizations extending into the 
neck of bursa; genital opening simple. 

A distinct genus of uncertain affinities, easily distin- 
guished by its male and female genitalia. Represented 
only by its type species. 


546. Lascelina canens, new species 
Fieures 562, 1055 


Forewing grayish white (the ground color that of 
wood ashes); antemedial and subterminal lines faint, 
indicated chiefly by the dark markings bordering them; 
antemedial line bordered inwardly by a thin broken 
blackish line and terminating at costa in a small reddish 
brown patch; a similar, smaller, reddish brown patch on 
midcosta; subterminal line bordered on both sides 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


towards costa by blackish fuscous; discal spots at end of 
cell, small, distinct, blackish; along termen a confluent 
row of blackish dots. Some reddish brown and blackish 
scaling on the labial palpi and a strong admixture of 
blackish scales in the antennal tuft of the male. Hind 
wing smoky white, with a dark line along termen. Alar 
expanse, 11-16 mm. 

Male genitalia with uncus very narrowly rounded at 
apex; costa of harpe broadly sclerotized; terminal mar- 
gin oblique, straight; anellus with lateral margins 
slightly produced; aedeagus bulged before apex. Fe- 
male genitalia as given for the genus. 

Typr Locatity: Brownsville, Tex. (type m USNM, 
61380; paratypes in BM, Paris Mus., and Cornell. 
Canadian National, and Janse Collections). 

Foop piant: Maytenus phyllanthoides. 

Described from male type and 24 male and 17 female 
paratypes from the type locality (40 of these reared 
Apr. 29, Aug. 4, and Dec. 7, 1938, and June 16, 1943, by 
members of the Foreign Plant Quarantine Division of the 
U. S. Department of Agriculture at Brownsville, Tex., 
from larvae feeding on the leaves of Maytenus) ;1maleand 
41 female paratypes from San Benito, Tex. (Mar., Apr., 
May, Aug., Sept.); 1 female paratype from Redington, 
Ariz., and 1 female paratype from Prescott, Ariz. (July). 
In addition to the foregoing I have also seen a male from 
Los Mochis, Sinaloa, México, intercepted in quarantine 
at Nogales, Ariz. (Feb. 27, 1941). 


Genus 155: Metephestia 


[Venational division D. Forewing with 10 veins; 4 absent; 10 
from the cell; hind wing with 3-5 approximate at base; discocel- 
lular vein straight, vertical. Male genitalia with uncus bifur- 
cate.] 


155. Genus Metephestia Ragonot 


Meiephestia Ragonot, Monograph, pt. 2, p. viii, 1901.—Hamp- 
son, in Ragonot, Monograph, pt. 2, p. 87, 1901. (Type of 
genus: Ephestia simplicula Zeller.) 

Tongue well developed. Antenna very shortly pu- 
bescent;shaft of male with a shallow sinus and small scale 
tuft at base; shaft of female simple. Labial palpus up- 
turned, slender, reaching to vertex; third segment acu- 
minate, about half as long as second. Maxillary palpus 
filiform. Forewing smooth; 10 veins; vein 2 from before 
but near lower outer angle of cell; 3 from the angle; 4 
absent (united with 5); 5 separated slightly from 3 at 
base; 6 from below upper angle of cell, very slightly 
curved (nearly straight); 8 and 9 stalked; 10 from the 
cell, separate from the stalk of 8-9; male without costal 
fold. Hind wing with vein 2 from well before lower 
outer angle of cell; 3 and 5 from the angle, approximate 
at base; 7 and 8 anastomosed for less than half their 
lengths beyond cell; cell very short, about one-fifth the 
length of wing; discocellular vein straight, vertical. 
Abdomen of male with a strong pair of ventrolateral hair 
tufts. 

Male genitalia with apical projection of gnathos pro- 
duced into a long, slender hook with notched apex. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Uncus bifurcate. Transtilla absent. Harpe simple 
except for short, bluntly rounded clasper; slender; sac- 
culus reduced; costa not appreciably sclerotized; apex 
rounded. Anellus a narrow curved band with long 
lateral lobes and long, slender, sclerotized lat- 
eral attachments to tegumen. Aedeagus long, stout, 
partially sclerotized; penis armed with a single, stout 
cornutus. Vinculum elongate, narrowing abruptly be- 
yond base. 

Female genitalia without signum; bursa copulatrix 
with some fine sclerotized wrinklings and a small lobe of 
thickened membrane near ductus bursae; ductus bursae 
much shorter than bursa, expanding abruptly to en- 
larged genital opening; a pair of weakly sclerotized 
plates on membrane behind genital opening; ductus and 
genital opening otherwise simple; ductus seminalis from 
bursa near thickened lobe. 

A distinct genus of uncertain affinities, with male gen- 
italia unlike anything else in group II. Contains but 
one described species. 


547. Metephestia simplicula (Zeller) 
Fiaures 100, 563, 1056 


Ephestia simplicula Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 
246, 1881. 

Metephestia simplicula (Zeller) Hampson, in Ragonot, Mono- 
graph, pt. 2, p. 87, 1901. 

Forewing unicolorous light gray brown to dark gray 
(reared examples much darker than flown specimens); 
transverse lines obsolete; in fresh specimens faint traces 
of dark terminal and discal dots, otherwise unmarked. 
Hind wing semihyaline white with a faint ocherous tint 
towards apex and termen. Alar expanse, 11-16 mm. 

Genitalia as given for the genus; apex of harpe nar- 
rowly rounded; vinculum longitudinally grooved and 
with notched apex. 


Typr Locauity: Honda, Colombia (type in BM). 


Foop puants: Indigofera tinctoria and I. verbasifo- 
lium. 

Distrisution: Unirep Sratss: Florida, Key West 
(Mar., Apr.). Pusrro Rico: Catafio (Apr., July), 
Coamo Springs (Apr.), Dovado (May), Rio Piedras 
(Aug.), San German (Apr.); Puerto Real (Vieques Isl., 
Apr.). Cotomsra: Honda (Apr.). Also reported by 
Hampson from St. Vincent Isl., British West Indies. 


The above food-plant records are from a series of 
moths reared by the Special Survey of the Division of 
Foreign Plant Quarantine of the U. S. Bureau of Ento- 
mology and Plant Quarantine from larvae webbing the 
leaves of Indigofera. This is the first record we have of 
the species from the United States or of its host plant. 


Genera 156-159: Selga to Rioja 


[Venational division A. Forewing with 11 veins; 4 and 5 stalked. 
Hind wing with 2 from before lower outer angle of cell; 3 and 5 
stalked or connate (Entmemacornis); discocellular vein curved. 
Male genitalia with apical process of gnathos bifid; transtilla 
complete (except in Selga). Labial palpi oblique (Selga) or 
upturned.] 


265 
156. Selga, new genus 


Typ or GENUS: Heterographis arizonella Hulst. 

Tongue well developed. Antenna pubescent; shaft 
of male simple. Labial palpus oblique, reaching height 
of vertex; slender; third segment nearly as long as second, 
apex bluntly pointed. Maxillary palpus minute, fili- 
form. Forewing smooth; 11 veins; vein 2 from before 
but near lower, outer angle of cell; 3 from the angle; 4 
and 5 shortly stalked (for less than half their lengths), 
stalk separated at base from 3; 6 from below upper angle 
of cell, straight; 8 and 9 stalked; 10 from the cell, sepa- 
rate and divergent from the stalk of 8-9; male with a 
narrow costal fold, extending for over one-fourth of costa 
from base. Hind wing with vein 2 from before lower 
outer angle of cell; 3 and 5 from the angle, shortly 
stalked (for less than half their lengths); 7 and 8 strongly 
anastomosed for most of their lengths beyond cell; cell 
one-half the Jength of wing; discocellular vein curved. 
Kighth abdominal segment of male simple. 

Male genitalia with apical process of gnathos U- 
shaped (consisting of a pair of widely spaced, short, 
blunt arms). Uncus stout, broadly rounded. Trans- 
tilla incomplete. Harpe simple, apex rounded; costa 
sclerotized throughout and projecting slightly at apex. 
Anellus a broad, curved plate. Aedeagus short, stout; 
penis with sclerotized wrinklings, otherwise unarmed. 
Vinculum stout, rather short (broader than long); 
terminal margin broadly rounded. 

Female genitalia with bursa copulatrix elongate, 
minutely scobinate; signum present, consisting of a 
small coarsely and bluntly spined plate. Ductus bursae 
unsclerotized, shorter than bursa; with a pair of narrow 
elongate dorsal plates and minute scobinations at genital 
opening, otherwise simple. Ductus seminalis from 
bursa about half-way between signum and junction of 
ductus bursae. 

A distinct genus easily distinguished on male genitalia 
from anything else in group IT with 11 veins in forewing. 
Its male genitalia are strikingly similar to those of 
Myelopsis coniella in group I to which it apparently is 
closely related. It contains one North American 
species. 


548. Selga arizonella (Hulst), new combination 
Ficurss 564, 1057 


Heterographis arizonella Hulst, Journ. New York Ent. Soc., vol. 
8, p. 222, 1900.—McDunnough, Check list, No. 6340, 1939. 


Forewing white, finely and evenly peppered with 
blackish scales giving the wing a pale ashy gray color; 
antemedial line white, slightly notched in cell and more 
deeply notched at lower fold, outwardly margined by a 
blackish line and inwardly margined by a broader 
blackish shade; subterminal line parallel with and 
rather near to terminal margin, indented slightly at vein 
6 and lower fold, white bordered inwardly and out- 
wardly by blackish lines; blackish discal dots at end of 
cell small, more or less confluent, in more strongly 
marked specimens a fine blackish line extends from 
them to middle of inner margin; a row of small blackish 


266 


dots along terminal margin. Hind wing glistening 
white with some faint fuscous shading on the veins and 
along terminal and costal margins. Alar expanse, 
14-19 mm. 

Genitalia as given for the genus; aedeagus sharply 
constricted towards apex; apical margin of anellus 
broadly notched; elements of transtilla long, slender. 


Type LocaLity: Catalina Springs, Ariz. (type in 
USNM). 


Foop prant: Unknown. 
Distrisution: Arizona, Catalina Springs (Apr.), 
Redington, ‘‘Southern Arizona” (May), Tempe (June). 


157. Genus Entmemacornis Dyar 


Entmemacornis Dyar, Ins. Insc. Menstr., vol. 7, p. 57, 1919. 
(Type of genus: Entmemacornis proselytes Dyar.) 

Tongue well developed. Antenna pubescent; of 
male (fig. 566f) with basal segment broadened and shaft 
with notch at base; of female simple. Labial palpus 
upturned, reaching to vertex, slender; third segment 
nearly as long as second, acuminate. Maxillary palpus 
minute, filiform. Forewing smooth; 11 veins; vein 2 
from well before lower, outer angle of cell; 3 from the 
angle; 4 and 5 shortly stalked (for less than half their 
lengths), the stalk separated from 3 at base; 6 from 
below upper angle of cell, slightly curved towards base; 
8 and 9 long stalked; 10 from the cell but approximate 
to the stalk of 8-9 for some distance; male with a 
narrow costal fold. Hind wing with vein 2 from well 
before lower outer angle of cell; 3 and 5 connate from 
the angle; 7 and 8 anastomosed almost to costa (free 
element of 8 very short); cell less than half the length 
of the wing; discocellular vein curved. Eighth abdom- 
inal segment of male with a pair of ventrolateral hair 
tufts. 

Male genitalia with apical process of gnathos bifid, 
consisting of a pair of short blunt arms (similar to but 
not so widely separated as those of Selga). Uncus 
semispoon-shaped (of the Diatomocera type). Tran- 
stilla complete but weakly sclerotized, a broad, finely 
scobinate plate, weakly attached to harpes. Harpe 
simple; terminal margin evenly rounded; costa sclero- 
tized throughout but not produced; on outer surface 
bordering lower margin a row of modified, flattened 
setae. Anellus a curved plate with short lateral lobes. 
Aedeagus stout, straight, moderately long; penis with a 
few sclerotized wrinklings and several minute spines 
(the latter distinguishable only under high magnifica- 
tion), otherwise unarmed. Vinculum stout, elongate, 
constricted towards angulate terminal margin. 

Female genitalia with bursa copulatrix long, minutely 
but sparsely scobinate, the scobinations extending into 
ductus bursae; signa present, consisting of small 
sclerotized disks; ductus bursae long, slender, un- 
sclerotized; genital opening simple; ductus seminalis 
from bursa near its anterior end. 

On nearly all characters except hind-wing venation 
this genus agrees with Diatomocera in group I. Were 
it not for the fact that the four specimens of the type 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


series of Hntmemacornis proselytes are consistent in 
venation, I should be inclined to discount the absence 
of vein 4 in hind wing as an individual aberration and 
unite the two genera; but apparently the same relation- 
ship prevails here as that between Drescoma and Dresco- 
mopsis. Entmemacornis is closely related to and 
obviously derived from Diatomocera but is apparently 
distinct. 
549. Entmemacornis proselytes Dyar 
Fieures 81, 566, 1058 


Enimemacornis proselytes Dyar, Ins. Insc. Menstr., vol. 7, p. 58, 


Thorax gray brown; tegulae tips black. Forewing 
eray-brown dusted with dull white and reddish scales 
(the latter only distinguishable under magnification) ; 
antemedial line slanting, notched at lower fold, whitish 
and concolorous with the general color of the basal area, 
defined inwardly by a narrow, faint, dark border and 
outwardly by a diffused dark shade; from base of wing 
a median longitudinal black streak extending to but 
not cutting the antemedial pale line; subterminal line 
faint, parallel and rather close to termen, outcurved 
between veins 6 and lower fold, formed by whitish 
scaling on the veins and bordered by narrow inner and 
outer dark lines more or less interrupted between the 
veins; distal and terminal dots, small, faint, blackish; 
between cell and subterminal line black streaks on veins 
5 and 6; costal fold of male about one-third the length 


of costa. Hind wing pale, glossy, brownish gray; the 
veins and terminal margin darker. Alar expanse, 
15-17 mm. 


Male genitalia with terminal margin of uncus convex; 
apex of harpe narrowly rounded; arms of bifid apical 
process of gnathos slightly convergent toward their 
apices; plate of anellus narrow. Female genitalia with 
bursa bent sharply at middle, its terminal margin 
straight. 

TypELocatiry: Cayuga, Guatemala (typein USNM). 

Foop piant: Unknown. 

Known only from the type series, two males and two 
females from the type locality. 


550. Entmemacornis pulla, new species 
FicurE 567 


Forewing blackish gray; antemedial line slanting, 
notched at lower fold, dull white with a very faint, 
narrow, blackish outer border; subterminal line some- 
what irregular, slanting sharply inwardly from costa to 
vein 8, then outwardly convex to lower fold, thence 
straight to inner margin, dull white with some blackish 
streaks on the veins bordering it on both sides; discal 
and terminal dots small, faint, blackish; basal area of 
wing before antemedial line and between cell and inner 
margin dull reddish ocherous; a dusting of similar color 
along the fold beyond antemedial line; from base of 
wing a median longitudinal black streak extends to 
antemedial line; bordering termen a narrow faint dust- 
ing of whitish scales; costal fold of male less than one- 
fourth the length of costa. Hind wing very pale 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


brownish gray; veins and terminal margin but little 
darker. Alar expanse, 19.5 mm. 

Male genitalia with terminal margin of uncus slightly 
concave; apex of harpe rather broadly rounded; arms 
of bifid apical process of gnathos divergent toward their 
apices; plate of anellus broad. 

Type Locatiry: Santa Catarina, Brazil (type in 
USN, 61381). 

Foop prant: Unknown. 

Described from unique male type collected by Fritz 
Hoffmann Sept. 27, 1934. 

The species is easily separated from proselytes by its 
darker (blackish gray) ground color, more strongly con- 
trasted transverse pale lines, shorter costal fold, and dif- 
ferent genitalia. The tegulae are black tipped like those 
of proselytes, but their blackness is less contrasted 
against the dull reddish ocherous of the remainder of the 
thorax. 


158. Genus Cayennia Hampson 


Cayennia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 62, 
1930. (Type of genus: Cayennia rufitinctalis Hampson.) 

Tongue well developed. Antenna pubescent (cilia- 
tions less than the width of shaft). Labial palpus up- 
turned, reaching to vertex, slender, third segment nearly 
as long as second. Maxillary palpus minute, filiform. 
Forewing smooth; 11 veins; vein 2 from well before lower 
outer angle of cell; 3 from the angle; 4-5 stalked for 
nearly half their lengths, approximate at base to 3; 6 
from below upper angle of cell, very slightly curved; 8 
and 9 long stalked; 10 from the stalk of 8-9; 11 from 
well out on cell, running close to stalk of 8-9 for a short 
distance; male without costal fold. Hind wing with 
vein 2 from before lower outer angle of cell; 3 and 5 from 
the angle, stalked (for approximately one-half) ; 7 and 8 
anastomosed for most of their lengths beyond cell (free 
element of 8 a very short spur); cell nearly one-half the 
length of the wing; discocellular vein curved. Eighth 
abdominal segment of male with a pair of ventrolateral 
hair tufts. 

Male genitalic characters same as those for Entmema- 
cornis except penis with only sclerotized wrinklings. 

A distinct genus close to Hntmemacornis and agreeing 
with it in all male genitalic characters, differing only in 
having simple antennae and different venation (vein 10 
of forewing stalked with 8-9, and 3-5 of hind wing 
stalked) and in lacking costal fold on male forewing. 


551. Cayennia rufitinctalis Hampson 
Ficurss 72, 568 


Cayennia rufitinctalis Hampson, Ann. Mag. Nat. Hist., ser. 10, 
vol. 5, p. 62, 1930. 

Forewing pale ashy gray peppered with fuscous, the 
basal half of inner area and tornal half of terminal area 
tinged with rufous; antemedial line near middle of wing, 
narrow, slanting outwardly from costa to middle of cell 
thence inwardly to lower fold and thence outwardly to 
inner margin, whitish with blackish lines before and 
beyond it on the veins; subterminal line inwardly 
oblique and slightly concave from apex to inner margin 


267 


before tornus, whitish, margined inwardly by short 
blackish dashes on the veins; discal dots small, blackish, 
below them on an inwardly slanting line blackish dots 
on veins 3 and 2b; a row of small blackish dots along 
termen. Hind wing semihyaline white with veins, 
terminal and costal margins shaded with pale brown. 
Alar expanse, 17.5 mm. 


Male genitalia with apical portion of uncus narrow 
(considerably reduced as compared with that of 
Entmemacornis proselytes); transtilla triangulate; vin- 
culum tapering, but slightly constricted before narrow, 
angulate terminal margin. 

Typr LocaLity: Cayenne, French Guiana (type in 
BM). 

Foop piant: Unknown. 


The female is unknown. Hampson mentions only 
the male type in his description; but in the U. S. 
National Museum there is a male “‘cotype’”’ so labeled 
by Hampson from the type locality. The species is 
easily identified by its genitalia and the medial position 
of the antemedial line of forewing. 


159. Rioja, new genus 


TYPE OF GENUS: Rioja nexa, new species. 

Tongue well developed. Antenna pubescent. Labial 
palpus upturned, reaching vertex, slender; third seg- 
ment almost as long as second. Maxillary palpus 
minute, filiform. Forewing smooth, 11 veins; veins 2 
and 3 connate from lower outer angle of cell; 4 and 5 
short stalked (for decidedly less than half their lengths), 
approximate at base to 3; 6 from below upper angle of 
cell, straight; 8 and 9 long stalked; 10 from cell, ap- 
proximate to stalk of 8-9 at base; male without costal 
fold. Hind wing with 2 from before lower outer angle 
of cell; 3 and 5 stalked for less than half their lengths; 
7 and 8 anastomosed beyond cell for half their lengths; 
cell a trifle less than half as long as wing; discocellular 
vein curved. Eighth abdominal segment of male with- 
out hair tufts; sternite developed as a triangulate, 
sclerotized pocket. 


Male genitalia with apical process of gnathos bifid. 
Uncus semispoon-shaped, constricted before apical end. 
Transtilla a complete arched bridge. Harpe simple; 
cucullus broad and with terminal margin evenly 
rounded; costa sclerotized but not produced. Anellus 
a narrow curved band with long lateral arms partially 
encircling the aedeagus. Aedeagus stout, moderately 
long; penis minutely scobinate, otherwise simple. 
Vinculum stout, longer than broad, slightly tapering. 

This genus forms a link between the two preceding 
genera and Moerbes Dyar. I should hesitate to describe 
it upon a single male if it were not obvious that the 
specimen is not a freak, and if it did not have characters 
that prevented its inclusion in Entmemacornis, Cayen- 
nia, or Moerbes. 

Its uncus and gnathos are of the Entmemacornis 
type, its eighth-segment pocket similar to that of 
Moerbes; but its venation is unique. The connate con- 


268 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


dition of veins 2 and 3 (from the angle of the cell) at 
once distinguishes it from all near relations. 


552. Rioja nexa, new species 
Figures 70, 569 


Forewing blackish with basal area to antemedial area 
dusted with whitish scaling and contrastingly paler than 
remainder of wing; also some whitish dusting forming a 
pale transverse shade across wing at end of cell; ante- 
medial line outwardly slanting from costa to inner mar- 
gin with a slight notch at lower fold, white, outwardly 
bordered by a narrow black line; subterminal line faint, 
irregularly dentate, pale (whitish gray), bordered in- 
wardly by a broad blackish suffusion; discal dots con- 
fluent, forming a blackish line along discocellular vein; 
a row of faint blackish dots along termen. Hind wing 
hyaline white with a faint fuscous shade on costa and 
on termen towards apex. Alar expanse, 17 mm. 

Male genitalia with arms of bifid apical process of 
gnathos divergent toward their pointed apices; aedeagus 
bulging and slightly bent before its broad, truncate 
apex. 

Type LOCALITY: La Rioja, Argentina (type in USNM, 
61382). 

Foop piant: Unknown 

Described from a single male collected by Schaus. 
The specimen is undated. 


Genera 160-162: Moerbes to Edulica 


[Venational division A. Forewing with 11 veins; 4 and 5 stalked. 
Hind wing with 2 from before lower outer angle of cell; 3 and 
5 connate or stalked (Edulica); discocellular vein curved. Male 
genitalia with apical process of gnathos a stout hook with forked 
apex; transtilla complete or, if incomplete (Moerbes), elements well 
developed and their apices approximate. Labial palpi oblique or 
upturned (Edulica).] 


160. Genus Moerbes Dyar 


Moerbes Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 337, 1914. (Type 
of genus: Zophodia dryopella Schaus.) 

Tongue well developed. Antenna shortly ciliate on 
male (cilia as long as width of shaft). Labial palpus 
oblique, broadly scaled, reaching to level of vertex; 
third segment short (one-fourth the length of second), 
acuminate. Maxillary palpus moderately large, some- 
what rough scaled. Forewing smooth; 11 veins; vein 2 
from well before lower, outer angle of cell; 3 from the 
angle; 4 and 5 stalked (for a trifle less than half their 
lengths), the stalk shortly separated from 3 at base; 
6 from below upper angle of cell, slightly curved; 8 and 
9 stalked for about two-thirds their lengths; 10 stalked 
with 8-9; male with a short, narrow costal fold. Hind 
wing with vein 2 from well before lower, outer angle of 
cell; 3 and 5 from the angle, connate; 7 and 8 anasto- 
mosed for most of their lengths (free element of 8 very 
short) ; cell approximately one-third the length of wing; 
discocellular vein curved. Eighth abdominal segment 
of male without hair tufts, but with sternite developed 
as a shallow, triangulate, sclerotized pocket. 


Male genitalia with apical process of gnathos devel- 
oped as an elongate, stout, flattened hook with forked 
apex. Uncus subtriangulate with broad terminal mar- 
gin. ‘Transtilla incomplete, its elements elongate-angu- 
late and well developed. Harpe simple with terminal 
margin evenly rounded; costa sclerotized for four-fifths 
of its length, but not produced. Anellus a narrow, 
curved band with minute lateral lobes. Aedeagus mod- 
erately stout, nearly straight; penis simple. Vinculum 
stout, longer than broad, slightly tapering to blunt ter- 
minal margin. 

A distinct genus showing strong affinities in general 
habitus (wing markings and color) and male genitalic 
structure to Pseudodiwona of group I. However, the 
venational difference (the absence or presence of vein 4) 
is consistent in both genera and the association of 
Moerbes with the genus following (Moodnopsis) seems a 
natural and proper one. It contains three tropical 
American species represented in the National Museum 
by eight males. I have seen no females. 


553. Moerbes dryopella (Schaus) 
Ficures 82, 570 


Zophodia dryopella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p. 249, 1913. 


Moerbes dryopella (Schaus) Dyar (part), Proc. U. S. Nat. Mus., 
vol. 47, p. 337, 1914; Ins. Insc. Menstr., vol. 7, p. 54, 1919. 


Forewing white with a broad, dull, ocherous brown 
shade along inner margin, and a sparse peppering of 
black scales in the white areas; antemedial line rather 
broad, defined chiefly by thin inner and outer bordering 
black lines, and conspicuous as a white spot on inner 
margin, outwardly slanting from costa, inwardly angled 
at lower part of cell and thence incurved to inner mar- 
gin; subterminal line faint, defined by two black dashes 
before and beyond it on costa and thin, broken, blackish 
bordering lines, inwardly angled at vein 6, thence paral- 
lel to termen; costa from base to antemedial line black, 
and a fine black transverse subbasal line; lower discal 
spot at end of cell present, black; below it a similar black 
spot or dash merging into the brown shade bordering 
inner margin; discal spot at upper outer angle of cell 
obsolete; terminal blackish dots faint. Hind wing very 
pale smoky fuscous, veins and terminal margin darker. 
Alar expanse, 23 mm. 

Male genitalia with apex of apical projection of 
gnathos considerably broadened and rather deeply bi- 


furcate. Apices of transtilla elements well separated. 
Aedeagus smooth, stout. Terminal margin of vinculum 
angulate. 


TYPE LOCALITY: Juan Vinas, Costa Rica (Apr.; type 
in USNM). 

Foop piant: Unknown. 

Represented only by the unique male type. The 
specimens from Panamé referred by Dyar to Schaus’ 
species (Proc. U.S. Nat. Mus., vol. 47, p. 337, 1914) are 
congeneric but not conspecific. I am describing them 
in this paper as Moerbes emendata. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 269 


554. Moerbes alveolella (Ragonot), new combination 
Fiaure 571 


Zophodia alveolella Ragonot, Nouv. Gen., p. 31, 1888; Mono- 
graph, pt. 2, p. 25, 1901. 

The type in Paris is a female according to Ragonot. 
I have not seen it; but have before me two males from 
Santa Catarina, Brazil (Sept. 2, and Oct. 26, 1934, col- 
lected by Fritz Hoffman), which are a perfect match for 
Ragonot’s excellent figure (Monograph, pl. 25, fig. 7). 
They are identical in color and maculation with the type 
of dryopella but markedly different in structure: 8 and 
9 of forewing longer stalked (over two-thirds); apical 
process of gnathos longer, its apex narrow and shallowly 
bifurcate; apices of transtilla elements touching (but 
not fused) and forming a high arch over aedeagus; 
aedeagus more slender; terminal margin of vinculum 
bluntly rounded (almost straight). Alar expanse, 21-23 
mm. 


TyprE Loca.ity: Rio de Janeiro, Brazil (type in Paris 
Mus.). 


Foop puant: Unknown. 


555. Moerbes emendata, new species 
Figure 572 


Moerbes dryopella Dyar (not Schaus), Proc. U. S. Nat. Mus., vol. 
47, p. 337, 1914. 

Similar in markings and color to the two preceding 
species except that the ground color of forewing is a 
more soiled white (less contrasted, except for the white 
spot on inner margin formed by the base of the ante- 
medial line), and both upper and lower discal spots are 
present and distinct. The male genitalia are also 
markedly different: Uncus appreciably narrower than 
that of alveolella or dryopella; aedeagus with a thornlike 
projection from undersurface before apex; vinculum 
sharply tapering from middle to angulate terminal 
margin. Alar expanse, 16-20 mm. 

Tyrer Locatity: Cabima, Panamé (type in USNM, 
61383). 

Foop piant: Unknown. 

Described from male type from the type locality 
(May); one male paratype from La Chorrera, Panamé; 
one male paratype from Porto Bello, Panama (Mar.); 
one male from Lino, Panamé (from the Janse Collec- 
tion); one male from 6 miles up the Maroni River, 
French Guiana (Schaus, collector), and one male without 
locality label, collected by Schaus and presumably also 
from French Guiana. The Panama specimens, except 
for the one from Lino, were collected by Busck and are 
those referred by Dyar to dryopella Schaus. ‘The Porto 
Bello specimen is abnormal in that veins 8 and 9 are 
united on both forewings; but this is a freak. Other- 
wise the specimen is normal and on the other specimens 
the venation is normal. The species can be distin- 
guished at once by the thornlike projection from its 
aedeagus. 


161. Genus Moodnopsis Dyar 


Moodnopsis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 408, 1914. 
(Type of genus: Moodnopsis decipiens Dyar.) 

Campyloplesis Dyar, Ins. Insc. Menstr., vol. 7, p. 61, 1919. (Type 
of genus: Campyloplesis inveterella Dyar. New synonymy.) 

Tongue well developed. Antenna of male shortly 
ciliate (cilia no longer than width of shaft); of female 
pubescent. Labial palpus of male obliquely ascending, 
cylindrical, reaching nearly to vertex, third segment less 
than half as long as second; of female porrect, laterally 
somewhat flattened, second segment oblique, long, ex- 
tending to the level of vertex, third segment deflected 
forward, about half the length of second. Maxillary 
palpus filiform. Forewing smooth; 11 veins; vein 2 
from well before lower outer angle of cell; 3 from the 
angle; 4 and 5 stalked, the stalk approximate at base 
(and in some specimens for a short distance from base) 
with 3; 6 from below upper angle, slightly curved to- 
wards base; 8 and 9 long stalked; 10 from the cell, more 
or less approximate (rarely connate) to the stalk of 8-9 
at base; male with a long costal fold (extending for half 
or more than half the length of costa). Hind wing with 
vein 2 from well before lower, outer angle of cell; 3 and 5 
connate from the angle; 7 and 8 anastomosed beyond 
cell almost to apex (completely anastomosed in a couple 
of males); cell about one-third the length of the wing; 
discocellular vein curved. Eighth abdominal segment 
of male with a short pair of ventrolateral hair tufts and 
sternite developed as a triangulate sclerotized pocket. 

Male genitalia with apical process of gnathos de- 
veloped as a stout, flattened hook with forked apex. 
Transtilla complete, the apices of its elements enlarged 
and fused into a spreading, bulbous knob. Harpe with 
terminal margin evenly rounded; costa strongly sclero- 
tized for four-fifths of its length, but not produced; on 
outer surface, bordering inner margin, a row of modified, 
flattened setae. Anellus a narrow, curved band with 
short lateral lobes. Aedeagus smooth, straight or 
slightly curved, tapering slightly from base to apex. 
Vinculum stout, longer than broad, tapering or abruptly 
constricted towards angulate terminal margin. 

Female genitalia with bursa copulatrix finely scobi- 
nate; signum present as an elongate narrow projecting 
plate with serrate edge; ductus bursae shorter than 
bursa, simple; genital opening simple; ductus seminalis 
from bursa towards (but not near) its junction with 
ductus bursae. 

Dyar described Moodnopsis from two large dark fe- 
male specimens and Campyloplesis from two small pale 
males. On the evidence of these alone the generic sep- 
aration would seem valid enough; but the evidence of 
associated males and females of species other than the 
type of genus shows no consistent character for such 
separation. The palpal differences are purely sexual 
and the trifling venational differences are either individ- 
ual or, at most, specific in character. Moodnopsis is a 
distinct genus easily identified by its male genitalia, and 
is somewhat more closely related to Moerbes than to 


270 


Moodna, from which Dyar distinguished it. His diag- 
nosis of Campyloplesis is in error in two important par- 
ticulars: The male labial palpi are not ‘“‘porrect” but 
obliquely ascenting, almost upcurved; veins 3, 4, and 5 
of forewing are not “stalked,” 3 being only approximate 
to the stalk of 4-5 at base and for a very short distance 
beyond. 


556. Moodnopsis decipiens Dyar 
Fieures 83, 1062 
Moodnopsis decipiens Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 408, 
1914. 


The two females from which the species was described 
(and the only specimens available) are rubbed so that 
the forewing pattern cannot be clearly distinguished. 
The ground color is a brownish gray (rather dark) with 
a faint dusting of whitish scales along costa and (under 
magnification) a scattered peppering of reddish scales 
over most of the wing; antemedial line indicated only by 
a rather broad dark outer border, this broken; the sub- 
terminal pale line faintly indicated, defined chiefly by 
dark streaks on the veins before and beyond it, the inner 
streaks rather long; discal dots faint but distinguish- 
able, blackish; dots along terminal margin very faint 
and more or less confluent. Hind wing pale fuscous; 
the veins and terminal margin darker. Alar expanse, 
27-28 mm. 

Female genitalia with signum large, the serrations 
along projecting edge bluntly and rather broadly 
rounded. 


TYPE LocaLity: Orizaba, México (type in USNM). 


Foop erant: Unknown. 

In unrubbed specimens the dark streaking along the 
veins would be more emphasized. Such lining of the 
veins is characteristic of all the species in the genus; but 
decipiens stands out because of its greater size and 
darker ground color. 


557. Moodnopsis perangusta (Dyar), new combination 
Ficure 573 


Euzophera perangusta Dyar, Ins. Insc. Menstr., vol. 7, p. 57, 1919. 


Ground color of forewing as in decipiens but trans- 
verse lines more distinct; the antemedial line deeply 
notched at lower fold; the subterminal irregularly den- 
tate and parallel with termen; dark borders of the trans- 
verse pale lines narrower and dark lining of the veins 
much less pronounced than in decipiens; blackish discal 
spots confluent; veins 4 and 5 longer stalked (for more 
than half their lengths); 8 and 9 longer stalked and 10 
closely approximate to the stalk of 8-9 for some distance 
beyond cell; costal fold extending to well beyond middle 
of costa. Hind wings translucent white, gray toward 
apex and along upper half of terminal margin. Alar 
expanse, 25 mm. 

Male genitalia with bifurcation of apical process of 
gnathos deeper, aedeagus smaller and less tapering, 
vinculum broader at, and less constricted before, ter- 
minal margins than those of other species in the genus. 
The genitalic differences among the several species are 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


better illustrated than described and can be readily 
distinguished in the drawings. 

TypE LocaLity: Montserrat, Trinidad, British West 
Indies (type in USNM). 

Foop prant: Unknown. 

Known only from the male type. It is possibly the 
male of decipiens. The size and coloration of forewing 
would suggest this. The white hind wings and vena- 
tion, however, would seem to indicate a distinct species, 
though the differences noted may be only sexual. The 
length of the stalking of veins 4 and 5 of forewing is 
certainly unreliable, varying in individuals of species 
where we have more than two specimens. 


558. Moodnopsis inornatella (Ragonot), new combination 
Ficures 574, 1061 


Zophodia inornatella Ragonot, Nouv. Gen., p. 31, 1888; Mono- 
graph, pt. 2, p. 25, 1901. 

The type of this species is a worn female from Costa 
Rica without abdomen. In his original description 
Ragonot gives the type locality as Brazil; but according 
to Clarke this could easily have been a misreading of a 
small pin label with ‘‘Druce” inscribed on it. The 
female in the British Museum is labeled ‘ty. original’ 
in Ragonot’s handwriting and bears the further infor- 
mation, ‘Irazu, 6-7,000 ft., H. Rogers.” This must be 
the type, for Dr. Burgogne states that it is not in the 
Ragonot Collection at the Paris Museum. 

A photograph of the type agrees well with a series of 
specimens in the National Museum (one male and three 
females) from Juan Vifias, Costa Rica. 

The general color of the forewing is paler than that 
of either decipiens or perangusta; but the markings are 
similar, the dark lining of the veins, if anything, more 
distinct. Worn specimens show no trace of an ante- 
medial line, but in one of the females before me (in 
better condition than the others) it is faintly outlined. 
The male costal fold is shorter than in perangusta, 
extending only to middle of costa. Hind wing of 
female very pale fuscous; of male soiled white; veins 
and edge of terminal margin darker. Alar expanse, 
20-28 mm. 

The distinguishing characters of the male genitalia 
are the shape of terminal projection of gnathos, the 
shape of apical lobe of transtilla, and the constricted 
vinculum. The female genitalia are similar to those of 
decipiens except that the signum is smaller and has 
weaker, less rounded serrations along its projecting edge, 
trifling differences of rather doubtful value. 

TypE Locauity: Irazt, Costa Rica (type in BM). 

Foop piant: Unknown. 

In addition to the Costa Rican specimens I have 
before me a2 male from Santa Catarina, Brazil (July 22, 
1935, Fritz Hoffmann, collector) which I take to be 
anornatella or a variety of it. The fore and hind wings 
are considerably paler (more whitish) and the ante- 
medial and subterminal lines more clearly indicated. 
It is the same size as the male from Juan Vinas (20 
mm.), has a costal fold of the same length, and agrees 
with it in every genitalic detail except that the forking 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 271 


of the apical process of gnathos is a trifle shallower. It 
certainly is not conspecific with the males of the follow- 
ing species from the same locality and collection. 

Upon superficial examination vein 3 of forewing seems 
to be connate with the stalk of 4-5 in inornatella and in 
most specimens of the other species, and is so described 
by Ragonot; but if the wings are partially denuded on 
underside at outer end of cell, more or less of a sepa- 
ration shows between 8 and 4—5 at their bases. 


559. Moodnopsis parallela, new species 
Fiaure 575 


Forewing similar in color and markings to that of the 
females of inornatella; costal fold very long, extending 
for at least two-thirds the length of costa. Male geni- 
talia with apical process of gnathos longer than that of 
other species of Moodnopsis except inveterella and its 
apical bifurcation U-shaped; aedeagus very stout, 
straight and evenly tapering; apical lobe of transtilla 
considerably enlarged; vinculum constricted before 
terminal margin, similar to that of inornatella. Alar 
expanse, 21-23 mm. 

Type Locauity: Santa Catarina, Brazil (type in 
USNM, 61384). 

Foop prant: Unknown. 

Described from male type and four male paratypes 
from the type locality (Oct. 2, 1934, July 7, 8, 12, 1935, 
Fritz Hoffmann, collector). In addition to these I have 
before me a somewhat larger male (26 mm.) with identi- 
cal genitalia from Tinguri, Carabaya, Peri (Schaus, 
collector) 

The female is unknown. 


560. Moodnopsis inveterella (Dyar), new combination 
Ficures 86, 576 


Campyloplesis inveterella Dyar, Ins. Insc. Menstr., vol. 7, p. 61, 


Similar in color and maculation to inornatella Ragonot 
but much smaller and with quite different genitalia. 

The venation of the type is figured (fig. 86) to show 
extent of variation in venation within the genus between 
it and the large female of decipiens (fig. 83). The com- 
plete fusion of veins 7 and 8 of hind wing in the type of 
inveterella is apparently an abnormality, a short spur of 
8 showing in one of the hind wings of the paratype. 
The costal fold extends for half the length of costa. 
Alar expanse, 14.5 mm. 

The distinguishing characters of the genitalia are: 
The stout, flat-faced apical process of gnathos (fig. 576b) 
with shallow V-shaped bifurcation at its apex; the nar- 
row, looped knob at apex of transtilla (fig. 576c); and the 
very abruptly tapered vinculum with its narrow termi- 


nal margin. The aedeagus is moderately stout and 
tapering. 

TypE tLocauity: Cayuga, Guatemala (type in 
USNM). 


Foop prant: Unknown. 
Known only from the male type (June) and a male 
paratype (Apr.) from the type locality. 


561. Moodnopsis portoricensis, new species 
Fiaures 577, 1060 


In size, color, and maculation similar to inveterella but 
with differently shaped uncus, apical process of gnathos, 
and transtilla. Alar expanse, 14.5-16 mm. 

The female genitalia are distinguished only by a very 
narrow, short signum, with sharply serrate edge and set 
far towards the anterior end of the bursa. The position 
of the signum is a character of very doubtful value. 

Type tocauity: Lares, Puerto Rico (type in Cornell 
Univ.; paratypes in USNM, 61385). 

Foop prant: Unknown. 

Described from male type and one male and one 
female paratype from the type locality, collected by 
Francesco Sein, Jr., July, 1931. 


162. Genus Edulica Ragonot 


Edulica Ragonot, Monograph, pt. 2, p. ix, 1901.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 122, 1901. (Type of genus: 
Euzophera compedella Zeller.) 

Tongue well developed. Antenna pubescent; basal 
joint enlarged on male. Labial palpus upturned; third 
segment approximately half as long as second, dorsally 
flattened. Maxillary palpus squamous, appressed to 
face. Forewing smooth; 11 veins; vein 2 from near 
lower outer angle of cell; 4 and 5 shortly stalked, from 
the angle; 3 connate or very shortly stalked with 4-5; 
6 from upper angle of cell, curved; 8 and 9 stalked; 10 
from the stalk of 8-9; male without costal fold. Hind 
wing with vein 2 from very close to lower outer angle of 
cell; 3 and 5 stalked for at least half their lengths; 7 and 
8 anastomosed beyond cell for approximately half their 
lengths; cell one-third the length of wing; discocellular 
vein curved. EKighth abdominal segment of male with 
sternite developed as a shallow, triangulate, sclerotized 
pocket, otherwise simple. 

Male genitalia with apical process of gnathos devel- 
oped as an elongate, stout, flattened hook, with very 
slightly notched apex. Uncus stout with broadly 
rounded terminal margin. Transtilla complete, devel- 
oped as a strongly sclerotized arch with broad, flaring 
apical crossband. Harpe with a strong hair tuft from 
base of sacculus; constricted between sacculus and 
cucullus; terminal margin rounded; costa sclerotized for 
two-thirds its length, not produced; clasper short, ap- 
pressed. Anellus a slightly curved, narrow band with 
well developed lateral lobes. Aedeagus short, stout; 
penis armed with many strongly sclerotized folds and 
a cluster of moderately stout, straight, elongate spines 
(cornuti) about one-fourth as long as aedeagus. Vincu- 
lum stout, as broad as long, triangulate; terminal end 
pointed. 

Female genitalia with bursa copulatrix sclerotized 
towards junction with ductus bursae and with several 
sclerotized folds extending well into the ductus, finely 
scobinate over most of inner surface; signum present as 
a small, cupped, scobinate plate; ductus bursae much 
shorter than bursa; genital opening simple. 

The genus is apparently close to but distinct from 


272 UNITED STATES NATIONAL MUSEUM BULLETIN 2:07 


Euzophera, easily distinguished from that genus by the 
close association of veins 3, 4 and 5 of forewing, and the 
short cell of hind wing. Contains one tropical Ameri- 
can species. Hampson includes a species from Mada- 
gascar but this is probably improperly placed. 


562. Edulica compedella (Zeller) 
Fieures 58, 585, 1071 


Euzophera compedella Zeller, Horae Soc. Ent. Rossicae, vol. 16, 
p. 224, 1881. 

Edulica compedella (Zeller) Ragonot, Monograph, pt. 2, p. ix, 
1901.—Hampson, zn Ragonot, Monograph, pt. 2, p. 122, 
1901. 

Ground color of forewing varying from somber brown 
to gray-brown; costal half of wing dusted with ashy 
grayish white; the discal spots blackish, confluent; some 
scattered blackish dots on several of the veins; ante- 
medial line not defined; the subterminal one weakly so 
by its dark inner and outer bordering lines, irregularly 
dentate; terminal dots distinct, blackish. Hind wing 
of male whitish with dark brown shading on veins and 
along termen; of female smoky fuscous, darkening 
towards apex and termen. Alar expanse, 20-25 mm. 

Genitalia as given for the genus. 

TyPE Locatity: Honda, Colombia (type in BM). 

Koop Pprant: Unknown. 

Distrisution: PanamA: Porto Bello (Apr., May). 
Cotompia: Honda. French Guiana: St. Jean 
Maroni. Braziu: Amazonas, Ponte Nova (Rio 
Xingu). 


Genera 163 and 164: Huzophera and Exuperius 


[Venational division A. Forewing with 11 veins; 4 and 5 stalked. 
Hind wing with vein 2 from well before lower outer angle of cell; 
3 and 5 connate; cell long; discocellular vein curved. Male geni- 
talia with apical process of gnathos a stout, elongate hook with 
pointed apex; transtilla complete. Labial palpi upturned] 


163. Genus Euzophera Zeller 


Stenoptycha Heinemann (not Zeller), Schmetterlinge Deutsch- 
lands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 190, 1866. 
Melia Heinemann, Schmetierlinge Deutschlands und der 
Schweiz, Abt. 2, vol. 1, pt. 2, p. 209, 1866 (mew name, pre- 

occupied, proposed for Stenoptycha Heinemann). 

Euzophera Zeller, Trans. Ent. Soc. London, ser. 3, vol. 5, p. 456, 
1867 (mew name for Stenoptycha Heinemann); Stettiner 
Ent. Zeit., vol. 28, p. 377, 1867.—Hulst, Phycitidae of 
N. Amer., p. 174, 1880 (cites type as Myelozs cinerosella 
Zeller).—Hampson, Moths, vol. 4, p. 72, 1896, in Blanford, 
Fauna of British India (cites type as Ephestia biviella 
Zeller); 7n Ragonot, Monograph, pt. 2, p. 36, 1901.— 
Ragonot, Monograph, pt. 2, p. vii, 1901 (cites type as 
Phycis pinguis Haworth).—Forbes, Cornell Mem. 68, p. 
630, 1923.—Bisset, zn Pierce and Metcalf, Genitalia of the 
British Pyrales, p. 59, 1938 (notes fixation of type by 
Hulst).—Janse, Journ. Ent. Soc. South Africa, vol. 8, p. 31, 
1945. (Type of genus: Myelois cinerosella Zeller; figs. 77, 
578, 1063.) 


Tongue well developed. Antenna simple, shaft 
pubescent (ciliations very short). Labial palpus up- 
turned, reaching to or almost to vertex; second segment 
rough scaled; third segment about half as long as 


second, acuminate. Maxillary palpus subsquamous 
(somewhat rough scaled), appressed to face. Forewing 
smooth; 11 veins; vem 2 from well before lower, outer 
angle of cell; 3 from the angle; 4 and 5 stalked (for less 
than half the lengths) ; 6 from below upper angle of cell, 
straight; 8 and 9 long stalked; 10 normally from the 
cell, separated from or approximate to (rarely connate 
with or from the stalk of) 8-9; male without costal fold. 
Hind wing with vein 2 from well before lower outer 
angle of cell; 3 and 5 from the angle, connate, rarely (in 
individual specimens) very shortly stalked; 7 and 8 
strongly anastomosed beyond cell (for more than half 
their lengths); cell long, slightly more than half the 
length of wing; discocellular vein curved. Wighth 
abdominal segment of male simple except in cinerosella 
which has a pair of short ventrolateral hair tufts. 

Male genitalia with apical process of gnathos devel- 
oped as an elongate, stout, simple hook with pointed 
apex. Uncus stout, broadly triangulate. Transtilla 
complete, developed as a strongly sclerotized arch with 
prominent, caudally projecting, lateral lobes. Harpe of 
nearly equal width throughout; terminal margin 
rounded and with a short rounded projection from apex 
of costa; costa otherwise, for most of its length, strongly 
sclerotized; clasper not developed. Anellus a slightly 
curved U-shaped shield, the lateral lobes well developed 
and strongly sclerotized in the American species. 
Aedeagus moderately long, stout; penis armed with 
numerous, short, sharp spines. Vinculum stout, as long 
as (or very little longer than) broad; terminal margin 
rounded. 

Female genitalia with numerous sclerotized or scob- 
inate folds in bursa copulatrix adjacent to or at junction 
with ductus bursae (except im the European type, 
cinerosella), finely scobinate over most of inner surface; 
signum present as a small, cupped, scobinate plate; 
ductus bursae with genital opening simple; ductus 
seminalis from bursa near signum (in American species) 
or from terminal end of bursa (in cinerosella). Dorsal 
sclerotized area of eighth segment collar a narrow band 
with central tonguelike projection. 

Euzophera, like Zophodia, has served as the receptacle 
for a heterogeneous lot of species, most of which must 
be referred to various other genera. As here defined 
the genus contains only three described American 
species that can be referred to it with any certainty. 
These form a compact group with constant genitalic 
characters and similar forewing pattern: antemedian 
line far out on wing, subterminal line well back from the 
terminal margin, the space between the lines corre- 
spondingly reduced, and a white spot on discocellular 
vein. On venation, palpi, and male genitalia they 
agree with the European type of genus, cinerosella, but 
differ from it in wing pattern and two details of struc- 
ture, cinerosella having hair tufts on the eighth abdom- 
inal segment of the male, and the ductus seminalis from 
the anterior end of the female bursa (figs. 578b, 1063). 
Eventually it may be possible to give our American 
species a separate generic designation; but upon present 
knowledge they must be retained in Huzophera. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Venation in Huzophera is subject to considerable 
individual variation especially as regards the relation 
of vein 10 of forewing to 8-9. In our American species 
and the European cinerosella (fig. 77) vein 10 is from the 
cell and normally distinctly separated from the stalk of 
8-9, though in individual specimens of any American 
species it may be either connate with or approximate, 
for some distance, to the stalk. In the European 
pinguis Haworth, it may be from the middle of the 
stalk, short stalked, or connate with it, or (rarely) from 
the cell and distinctly separate. 


563. Euzophera semifuneralis (Walker) 
Ficures 579, 1064 


Nephopteryx semifuneralis Walker, List, pt. 27, p. 57, 1863. 

Euzophera aglacella Ragonot, N. Amer. Phycitidae, p. 14, 1887; 
Monograph, pt. 2, p. 65, 1901.—Hulst, Phycitidae of N. 
Amer., p. 177, 1890.—Essig, Insects of western North Amer- 
ica, p. 710, 1929.—McDunnough, Check list, No. 6316, 1939. 

Stenoptycha pallulella Hulst, Ent. Amer., vol. 3, p. 137, 1887. 

Euzophera semifuneralis (Walker) Ragonot, Ent. Amer., vol. 5, p. 
116, 1889.—Forbes, 8. A., Psyche, vol. 5, p. 295, 1890.— 
Hulst, Phycitidae of N. Amer., p. 175, 1890.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 66, 1901.—Blakeslee, U. S. 
Dep. Agr. Bull. 261, 13 pp., 1915.—Forbes, W. T. M., 
Cornel Mem. 68, p. 631, 1923.—HEssig, Insects of western 
North America, p. 710, 1929.—Keifer, Monthly Bull. 
Dep. Agr. California, vol. 20, p. 618, 1931.—McDun- 
nough, Check list, No. 6317, 1939. 

Forewing with basal and terminal areas dark to pale 
reddish brown more or less dusted with white along 
costa and (in very pale specimens) in apical-terminal 
area; area between the transverse lines densely dusted 
with blackish scales; antemedial line more or less ver- 
tical to lower margin of cell, inwardly angled at lower 
fold, white bordered outwardly by a black line; sub- 
terminal line somewhat irregular, usually out-angled 
evenly at middle, white, bordered inwardly by a black 
line; discal black dots more or less obscured in the black 
dusting of median area, usually a white mark on dis- 
cocellular vein; a line of black dots along terminal 
margin, confluent in some specimens, distinctly sepa- 
rated in others. Hind wing white to smoky fuscous, 
more or less darkened along terminal margin, at apex, 
and along some of the veins. Alar expanse, 16.5-28 
mm. (Hampson gives extreme expanse as 34 mm.). 

Male genitalia with lateral, projecting lobes of trans- 
tilla and lateral lobes of anellus more slender than those 
of other American species; cucullus of harpe narrower 
and more elongate; vinculum somewhat longer than 
broad; uncus evenly tapering. Female genitalia with 
bursa rather small and wrinkled over most of its surface. 

Type tocauitiss: “North America” (semifuneralis, 
in BM); Sonora, México (aglaeella, in Paris Mus.); 
Colorado (pallulella, in AMNH, ex Rutgers). 

Foop piants: Various deciduous trees (apple, apri- 
cot, pear, peach, plum, persimmon, cherry, mulberry, 
walnut, pecan, olive, linden, poplar, Liquidambar, 
Ginkgo, mountain ash). Larva a bark borer. Also 
stems of cotton and corn. 

Distrisution: Unirep States: New Hampshire, 
Hampton; Connecticut, East River (July, Sept.); New 


273 


York (no exact locality); New Jersey, Hackensack 
(Nov.), Montclair (May, Aug.), Morristown (June); 
Maryland, Plummers Isl. (May); District of Columbia, 
Washington (May, Sept.); Virginia, Cape Henry (July), 
Vienna (May); North Carolina, Southern Pines (Mar., 
Apr.), Tryon (May); South Carolina, Anderson (Oct.); 
Georgia, Fort Valley; Michigan, Monroe; Indiana, Bed- 
ford (Apr.); Illinois, Decatur (Apr.); Kansas, Lawrence 
(May), Onaga; Missouri, St. Louis (June); Mississippi, 
Jackson (Feb.), Tishomingo (Nov.); Texas, Blanco 
County, Brownsville, Brownwood (Oct.), Justine (June), 
Kerrville (Apr.), Paris, Plano (Sept.), San Benito (July), 
San Diego (May), Shovel Mountain, Snyder (Mar., 
Apr.), Victoria (Apr.), Zavalla County (Apr.); New 
Mexico, Mesilla, Roswell (Apr.); Arizona, Baboquivari 
Mts. (Apr., May), Chiricahua Mts., Garces, Huachuca 
Mts., Palmerlee, Scotsdale (May), Yavapai County; 
Colorado, Denver (June); Utah, Bellevue (May); Cali- 
fornia, Corningi (Jan.), Inyo County (June, July), 
Loma Linda (Sept.), Piru (Ventura County, July), Pla- 
cerville (Jan., May), Putah Canyon (Yolo County, 
Nov.). Canapa: British Columbia, Alberni (July), 
Duncans (Vancouver Isl., Aug.). Mzs&xico: Sonora. 

The species is variable in color but is easily distin- 
guished by the reddish basal and terminal areas of fore- 
wing and the narrowed, black-dusted area between the 
transverse lines. The name aglaeella represents a color 
form, with paler red-brown areas and more strongly 
contrasted, blackish median dustings and markings. 
It is commoner in the Western and Southwestern States 
and México than elsewhere but cannot be maintained 
asarace;forit also occurs in the Kast and there are inter- 
grades between it and the typical dark form, common to 
the Eastern and Central States, but occurring through- 
out the range of the insect. The species is of some im- 
portance as an orchard insect, the larvae often domg 
considerable damage as a bark borer in plum, olive, and 
walnut trees. It is known in economic literature as 
the ‘‘American plum borer.” 


564. Euzophera ostricolorella Hulst 
Figures 580, 1065 


Euzophera ostricolorella Hulst, Phycitidae of N. Amer. p. 175, 
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 67, 
1901.—Heinrich, Proc. U. 8. Nat. Mus., vol. 57, p. 87, 1920 
(life history, larva and pupa).—Forbes, Cornell Mem. 68, 
p. 631, 1923.—McDunnough, Check list, No. 6319, 1939. 

Forewing purplish brown, dark grayish fuscous along 
costa and with some grayish dusting in apical area; 
transverse lines whitish, not darkly bordered; anteme- 
dial line vertical to lower margin of cell, deeply angled 
below, at fold; subterminal line sinuate; a whitish line 

along discocellular vein at end of cell; along termen a 

row of obscure, blackish dots; in some specimens a suf- 

fusion of dark red brown in basal area and above inner 
margins beyond base, especially on the folds. Hind 
wing rather pale smoky fuscous with a fine dark line 

along terminal margin. Alar expanse, 29-40 mm. 

Male genitalia with lateral, projecting lobes of trans- 
tilla broader than those of semifuneralis; anellus very 


274 


broad, its lateral lobes wide and widely spaced, their 
apices narrowly rounded; harpe with cucullus broaden- 
ing toward apex; vinculum about as broad as long; 
uncus abruptly narrowed beyond base, thence tapering. 

Female genitalia with bursa large, densely spined and 
wrinkled towards junction with ductus bursae; ductus 
bursae very short, deeply wrinkled. 

Type Locatity: New York (in AMNH, ex Rutgers). 

Foop puant: Liriodendron tulipifera. 

Distrisution: New York, Long Island; District of 
Columbia, Washington (June); Maryland, Oxon Hill, 
Plummers Isl. (Oct.); Virginia, Leesburg; South Caro- 
lina, Greenville (June), Hendersonville (June), Paris 
Mountain (June), Rockhill (June), Yorkville (June); 
Georgia, Atlanta (Oct.). 

Probably distributed throughout the range of its host. 

This distinct species apparently has only one host, the 
‘tulip tree.’”’? The larva bores in the bark on the damp 
side near the base of the tree. 


565. Euzophera nigricantella Ragonot 
Ficures 581, 1066 


Euzophera nigricantella Ragonot, N. Amer. Phycitidae. p. 14, 
1887.—Hulst, Phycitidae of N. Amer., p. 177, 1890.—Hamp- 
son, in Ragonot, Monograph, pt. 2, p. 63, 1901.—McDun- 
nough, Check list, No. 6315, 1939. 

Euzophera griselda Dyar, Proc. U. 8. Nat. Mus. vol. 44, p. 322, 
1913 (new synonymy). 

Forewing gray evenly dusted with blackish; trans- 
verse pale lines paler, but obscure, shaped as in the pre- 
ceding species; antemedial line with an obscure narrow 
blackish outer border; subterminal with faint dark bor- 
ders on inner and outer sides; a whitish spot on disco- 
cellular vein; terminal dots, faint, blackish. Hind wings 
white with some dark shading on the veins and along 
terminal margin. Alar expanse, 21-27 mm. 

Male genitalia with lateral projecting lobes of trans- 
tilla largest of any known American Huzophera; anellus 
longer than broad, the lateral lobes untapering, nar- 
rowly spaced, and with outer margins at apices inwardly 
hooked; harpe with cucullus stubby, its lower margin 
slightly concave; vinculum as broad as long; uncus as in 
ostricolorella. Female genitalia with a wrinkled sclero- 
tized collar about ductus bursae at its junction with 
bursa copulatrix. 

Type tocauities: Arizona (nigricantella; in Paris 
Mus.); Tehuacén, México (griselda, in USNM). 

Foop piant: Unknown. 


Distrisution: Untrep Staves: Arizona, Baboqui- 
vari Mts. (Apr., June, July, Aug., Sept., Oct.), Reding- 
ton, Tempe (Mar.), Yavapai County; New Mexico, Al- 
buquerque (July) ; Texas, San Benito (May). M*xico: 
Sonora; Tehuacén (May, Sept.), San Jose de Guaymas 
(Apr.). 

Dyar’s griselda was described from females which 
match specimens of nigricantella from Arizona in every 
detail of maculation, color, and genitalia. At the time 
he described griselda Dyar had not recognized Ragonot’s 
species. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


164. Exuperius, new genus 


TYPE OF GENUS: Hruperius negator, new species. 

Tongue well developed. Antenna simple, pubescent. 
Labial palpus upturned, barely reaching to vertex; first 
segment broadly scaled; third segment nearly as long as 
second, acuminate. Mazxillary palpus minute, filiform. 
Forewing smooth; 11 veins; vein 2 from before lower 
outer angle of cell; 3 from the angle; 4 and 5 stalked (for 
less than half their lengths), separate from 3 at base: 6 
slightly bent, from below, but near upper angle of cell; 
8 and 9 rather long stalked (for two-thirds their lengths) ; 
10 from the cell, approximate to the stalk of 8-9 for 
some distance; male without costal fold. Hind wing 
with vein 2 from well before lower outer angle of cell; 
3 and 5 connate, from the angle; 7 and 8 anastomosed 
beyond cell for most of their lengths (free segment of 8 
short); cell long, slightly over half the length of wing; 
discocellular vein curved. Highth abdominal segment 
of male simple. 

Male genitalia with apical process of gnathos de- 
veloped as an elongate, stout, simple hook with pointed 
apex. Uncus narrow, constricted towards base, shaped 
like a narrow, bluntly pointed arrowhead. ‘Transtilla 
complete, but median portion a narrow, rather weakly 
sclerotized band. Harpe with broadened, evenly 
rounded cucullus; spoon-shaped; costa broadly sclero- 
tized but not produced; clasper present, short, knobbed 
at apex; from near base of cucullus a strong, long, semi- 
detached hair tuft. Anellus a broadly U-shaped nar- 
row band with long, flattened, lateral projections. 
Aedeagus long, stout; penis armed with a dense mass of 
minute granulations. Vinculum considerably shorter 
than broad, terminal margin broadly and evenly 
rounded. 

Female unknown. 

This genus is close to Huzophera and agrees with it in 
characters of venation, labial palpi, and gnathos; but its 
peculiar genitalia (aside from the apical projection of 
enathos) differ in every detail from those of typical 
Euzophera and suggest a separate generic designation. 
They resemble those of no other American group that 
I know. The partial forking of vein 1b of forewing at 
base (shown in fig. 76) is present on both forewings but 
may be an individual abnormality. It is most unusual 
for a phycetine. 


566. Exuperius negator, new species 


Ficures 76, 584 


Forewing brown, dusted with grayish white over basal 
area and for a short distance beyond antemedial line 
near costa and inner margin; some blackish dusting in 
cell beyond its middle and a black spot on costa on each 
side of the subterminal line; antemedial line indicated 
by a blackish band slanting slightly inward from slightly 
beyond basal third of costa to inner margin, straight; 
subterminal line somewhat wavy, obscure except on 
costal half where it is distinctly white; a small but con- 
spicuous white spot above the stalk of veins 4—5 at end 
of cell; terminal dots blackish, more or less confluent. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 275 


Hind wing hyaline white shaded with blackish fuscous, 
broadly along costa and narrowly along terminal margin; 
veins faintly outlined by dark shading. Alar expanse, 
22 mm. 

Male genitalia as given for the genus; aedeagus 
broadened at base, sharply constricted just beyond. 


Typr Locauity: La Chorerra, Patamayo District, 
Pert (type in Cornell Univ.). 

Foop piant: Unknown. 

Described from unique male type collected on the 
Cornell University Expedition (Apr. 1920) under lot 
No. 607. 

The white discal spot on forewing and the genitalia 
should easily identify the species. The former is also 
common to our American species of Huzophera but in 
that genus the transverse lines are much closer together 
than in negator. 


Genus 165: Eulogia 


[Venational division A. Forewing with 11 veins; 4-5 approx- 
imate, connate or very shortly stalked. Hind wing with 3 and 5 
connate; cell long; discocellular vein curved. Male genitalia 
with apical process of gnathos a short, flat, furcate plate; trans- 
tilla incomplete. Labial palpus upturned.] 


165. Eulogia, new genus 


Typz or Gunus: Ephestia ochrifrontella Zeller. 

Characters of Huzophera except: Labial palpus with 
third segment as long or nearly as long as second. Fore- 
wing of male with a narrow, elongate costal fold; veins 
4 and 5 approximate, connate or very shortly stalked. 
Male genitalia with apical process of gnathos a short, 
flat plate, furcate at apex; transtilla incomplete; aede- 
agus slender; penis armed with a single curved cornutus. 
Female genitalia with bursa copulatrix smooth except 
for a very few scobinations surrounding signum; signum 
a cluster of small, short, thornlike spines (not ona plate) ; 
ductus bursae strongly sclerotized at and just before 
genital opening, the sclerotized part of ductus tubular. 

On venation, wing maculation, and palpal characters 
the type species (ochrifrontella) could go well enough in 
Euzophera, where it has been placed; but the genitalia 
rule it out. All the really congeneric species of Europe 
and North America placed in Huzophera have similarly 
constructed, complete transtillae, the same type of 
hooked projection from gnathos, similar multiple cor- 
nuti on the penis, and similar wide-mouthed, unsclero- 
tized genital openings in the female. On all of these 
characters ochrifrontella is ruled out. 


567. Eulogia ochrifrontella (Zeller), new combination 
Ficures 78, 79, 583, 1069 
Ephestia ochrifrontella Zeller, Verh. zool.-bot. Ges. Wien, vol. 25, 


p. 337, 1876. 
Euzophera ferruginella Ragonot, N. Amer. Phycitidae, p. 14, 


Euzophera ochrifrontella (Zeller) Hulst, Phycitidae of N. Amer., 
p. 177, 1890.—Hampson, in Ragonot, Monograph, pt. 2, 


p- 67, 1901.—Ely, Proc. Ent. Soc. Washington, vol. 12, p. 
203, 1910.—Forbes, Cornell Mem. 68, p. 631, 1920.— 
McDunnough, Check list, No. 6318, 1939. 

Forewing copper colored, with the area between the 
transverse lines heavily dusted with black; antemedial 
line far out (slightly beyond middle) on wing, pale 
ocherous, angulate; subterminal line, parallel with 
termen, slightly indented at vein 6 and lower fold; an 
obscure pale ocherous line along discocellular vein; in 
some specimens the blackish dusting extends beyond 
the transverse lines over most of the basal and terminal 
areas of the wing, but this is not a normal condition. 
Hind wing pale smoky fuscous; veins and terminal 
margin slightly darker. Alar expanse, 11-15 mm. 

Genitalia as given for the genus. In the female the 
ductus bursae has a slightly sclerotized band near its 
junction with bursa. 

Typr tocauities: Texas (ochrifrontella, in MCZ); 
North Carolina (ferruginella, in Paris Mus.). 

Foop puants: Pecan, oak, apple. 

Distrisution: Unitrep Statrs: Massachusetts, Co- 
hasset (July), Framingham (Sept., Oct.), Martha’s 
Vineyard (July, Aug.); Connecticut, East River (July, 
Aug., Sept., Oct.), Stamford (July); New Jersey, Eliza- 
beth (Aug.); Pennsylvania, Adams County (June), 
Arendtsville (Aug.), Hazleton (July), New Brighton 
(June, July); Maryland, Plummers Isl. (May, June, 
July); North Carolina; Florida, Orlando (Mar.); Texas; 
Illinois, Decatur (May, June); Kansas, Onaga. Can- 
ADA: Manitoba, Rounthwaite (July), Winnipeg; British 
Columbia, Duncans (Vancouver Isl., June), Kaslo 


‘@uly). 


The foregoing food-plant records are from specimens 
in the National Museum. Nothing, as far as I know, 
has been published on the life history. 

The venation is variable in different specimens and 
sometimes on opposing wings of the same insect. On 
forewing vein 10 may be separate from, approximate to, 
or connate with the stalk of 8 and 9; 4 and 5 may be 
approximate or connate or, sometimes, very shortly 
stalked. On the hind wing the anastomisis of 7 and 8 
varies, but is normally short. There is no appreciable 
individual variation in genitalia. 


Genus 166: Prosoeuzophera 


[Venational division D. Forewing with 10 veins; 4 absent; 3 and 
5 connate. Hind wing with veins 3 and 5 connate or shortly 
stalked. Male genitalia with apical process of gnathos a stout, 
elongate hook with pointed apex; transtilla complete. Labial 
palpi upturned.] 


166. Prosoeuzophera, new genus 


Type or Genus: Euzophera impletella Zeller. 

Characters of Huzophera except: Forewing with 10 
veins; vein 4 absent; 3 and 5 connate from lower outer 
angle of cell. 

A direct derivative of Huzophera but with advanced 
venation, replacing Luzophera in tropical America. 


276 


568. Prosoeuzophera impletella (Zeller), new combination 


Figures 107, 582, 1067 


Euzophera impletella Zeller, Horae Soc. Ent. Rossicae, Vol. 16, 
p. 234, 1881. 

In color and maculation like small, pale examples of 
Euzophera semifuneralis with which it was synonymized 
by Hampson (Ragonot, Monograph, pt. 2, p. 66, 
1901). He either overlooked or ignored the distinct 
venational difference between impletella and the North 
American species of Huzophera. The genitalia of 
impletella (o' and ) exhibit differences of only a 
specific character from the Huzophera, quite distinct 
specifically from semifuneralis, as our figures show; but 
the absence of vein 4 and the connate condition of 
veins 3 and 5 of forewing are constant. Alar expanse, 
15-16.5 mm. 

TyrE Locality: Honda, Colombia (Gn BM). 

Foop piant: Unknown. 


Distrisution: Cotompra: Honda, Maraquita. Ja- 
marca (2, in USNM). Purrto Rico: Villa Margarita 
(Catano, Apr.) 


Genera 167-169: Farnobia to Micromescinia 


[Venational division B. Forewing with 10 or 11 veins; 2 and 3 
stalked or united. Cell open or incomplete in fore or hind wings. 
Male genitalia with transtilla complete.] 


167. Farnobia, new genus 


TyrE oF Genus: Huzophera quadripuncta Zeller. 

Tongue well developed. Antenna of male with basal 
segment narrowly elongate, bearing a short spur from 
inner side near apex; shaft deeply curved toward base, 
pubescent; of female simple. Labial palpus upturned, 
scarcely reaching vertex; third segment shorter than 
second. Mazxillary palpus minute, rather broadly and 
flatly scaled, appressed to face. Forewing smooth; 10 
veins; veins 2 and 3 united; 4 and 5 short stalked; 6 
curved and connate with the stalk of 8-9-10; 8 and 9 
stalked; 10 from the stalk of 8-9; cell open (discocellular 
vein absent); in male costa enlarged, lobed and sinuate, 
and outer margin concave between veins 7 and 5; an 
elongate discal fovea in male between veins 6 and the 
stalk of 3-5. Sternite of eighth abdominal segment of 
male developed as a broadly triangulate sclerotized 
pocket. 


Male genitalia with apical process of gnathos a short, 
stout, blunt hook. Uncustriangulate. Transtilla com- 
plete, a heavy arched bridge with enlarged lateral lobes. 
Harpe with incurvation between sacculus and cucullus; 
cucullus narrow and narrowly rounded at apex; costa 
sclerotized for most of its length, but not produced. 
Anellus a narrow curved band with broad, curved, 
bandlike lateral projections (clasping the sides of 
aedeagus). Aedeagus stout, somewhat bent; penis 
armed with numerous sclerotized folds and two or three 
stout, very short, thornlike spine clusters. Vinculum 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


stout, as broad as long, bluntly triangulate; terminal 
margin reinforced and broadly rounded. 

Female genitalia with bursa copulatrix large and 
elongate, finely scobinate over surface of anterior half, 
thickened and partially sclerotized at junction with 
ductus bursae; signum present as a sclerotized cup 
evenly covered with short, stout, blunt spines; ductus 
bursae very short, a mere neck between genital opening 
and bursa; genital opening with a curled-over, strongly 
sclerotized and centrally emarginate lower margin; 
ductus seminalis from a lobe of bursa near its junction 
with ductus bursae. 


In genitalia this genus resembles the Hemiptiolocera— 
Crocidomera complex of group I with which it is appar- 
ently related. It is even more nearly related to the 
genus following (Gennadius) which also has the cell of 
forewing open and (except for specific differences) sim- 
ilar male genitalia. However, Farnobia is so different 
in venational characters that it need not be confused 
with anything else in the Phycitidae. The forewing 
venation might be interpreted differently from what 
T indicate (i. e., vein 4 rather than 3 absent and 3 and 
5 short stalked); but in view of the outward position 
of 2 and the close association of Farnobia with 
Gennadius (which has 2 and 3 distinctly stalked) the 
correct interpretation appears to be 2 and 3 united. 


569. Farnobia quadripuncta (Zeller), new combination 
FicuREs 98, 99, 586, 1070 


Euzophera quadripuncta Zeller, Horae Soc. Ent. Rossicae, vol. 16, 

p. 226. 

Dannemora quadripuncta (Zeller) Hampson, in Ragonot, Mono- 
graph, pt. 2, p. 209, 1901. 

Forewing brown with a purplish tint; the costal area 
dusted with ocherous gray, this pale shading extending 
into the cell at middle; antemedial line obscure, blackish, 
dentate and outwardly oblique from costa to inner mar- 
gin; subterminal line better defined, blackish, sharply 
dentate at the veins; three rather conspicuous blackish 
dots forming an angle near extremity of cell and, in some 
specimens, traces of a fourth dot on costa marking be- 
ginning of antemedial line. Hind wings whitish, semi- 
hyaline, with a narrow dark line along termen and dark 
shading on some of the veins. On the underside of the 
male hind wing the costal area is filled with coarse 
ocherous scaling terminating in a blackish fuscous patch 
at the outer angle of thelobe. Alar expanse, 21-24 mm. 

Genitalia as given for the genus. 

TyprE Locatity: Honda, Colombia (type in B. M). 


Foop prant: Unknown. 

Distrizution: Costa Rica: Esperanza (May). 
Pawnam&: La Chorrera (May), Cabima (May). France 
Guiana: St. Laurent du Maroni. Cotompra: Honda. 
Also recorded by Hampson from Brazil. 

When Hampson placed quadripuncta in Dannemora 
he had only females of Zeller’s species and no authentic 
examples of the type of Hulst’s genus. The two species 
have very little in common, structurally or otherwise. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 277 


168. Gennadius, new genus 


Type or Genus: Gennadius junctor, new species. 

Tongue well developed. Antenna of male with basal 
segment narrowly elongate, bearing a short spur from 
middle of inner side; shaft with a short spur from basal 
joint, deeply curved for several segments beyond, pu- 
bescent. Labial palpus oblique, slender, dorsoventrally 
flattened, reaching above vertex; third segment about 
as long as second. Maxillary palpus moderately sized, 
flatly scaled and appressed to face. Forewing with a 
subbasal clump of raised scales (possibly a male or spe- 
cific character) ; 11 veins; veins 2 and 3 stalked; 4 and 5 
stalked (for about half their lengths); 6 curved, and 
connate with the stalk of 8-9-10; 8 and 9 stalked; 10 
from the stalk of 8-9; cell open (discocellular vein ab- 
sent); male without costal fold. Hind wing of male 
with vein 2 from very close to angle of cell; 3 and 5 
stalked; 7 and 8 united; in male costa triangularly pro- 
duced and outer margin concave between veins 7 and 5; 
cell closed, very short (less than one-fourth the length 
of wing in male); discocellular vein curved. Sternite of 
eighth abdominal segment of male developed as a 
triangulate, sclerotized pocket. 

Male genitalia as in Farnobia except: Apical hook of 
gnathos slightly furcate at apex; lateral lobes of trans- 
tilla bluntly pointed rather than rounded; lateral arms 
of anellus shorter, narrower and tapering; harpe not 
incurvate between succulus and cucullus. Most of 
these differences are probably specific rather than 
generic in character. 

Female unknown. 

The genus is very close to Farnobia as indicated by 
the antennal and genitalic structures and the open cell 
of forewing, but is easily separated by the stalking rather 
than fusion of veins 2 and 3 of forewing and by the 
closed cell and the absence of vein 8 in hind wing. 


570. Gennadius junctor, new species 
Figures 97, 587 


Forewing brown with a purplish tint; the costal area 
dusted with ocherous gray; some blackish streaking on 
upper vein of cell and along median fold and a few 
blackish spots indicating the broken margins of the 
transverse lines; a conspicuous white patch along inner 
margin between antemedial and subterminal lines and 
extending from inner margin to cell; transverse lines 
obscure. Hind wing very pale brown; terminal margin 
and veins little if any darker; in the male rather coarsely 
scaled over most of undersurface and with a border of 
coarse, thick, ocherous scaling along costa. Alar 
expanse, 22.5 mm. 

Male genitalia as given for the genus; figure 587b 
shows the penis extruded to display the cornuti. 

Typr Locauity: St. Jean Maroni, French Guiana 
(type in USNM, 61886). 

Foop piant: Unknown. 

Described from male type and one male paratype 
from the type locality (Schaus, collector, no date given). 


The species is easily identified by its structural 
characters and the elongate white patch on inner 
margin of forewing. 


169. Genus Micromescinia Dyar 


Micromescinia Dyar, Proc. U. 8S. Nat. Mus., vol. 47, p. 347, 1914. 
(Type of genus: Micromescinia pygmaea Dyar.) 

Tongue well developed. Antenna simple. Labial 
palpus oblique, reaching to or a trifle above vertex; slen- 
der; third segment nearly as long as second, acuminate. 
Maxillary palpus minute, filiform. Forewing smooth; 
10 veins; veins 2 and 3 stalked; 4 and 5 long stalked; 6 
from below upper angle of cell, very slightly curved; 8 
and 9 united (9 absent); male with a short costal fold 
enclosing a hair tuft. Hind wing with 2 from before 
(but near) lower outer angle of cell; 3 and 5 stalked, 
from the angle; 7 and 8 completely fused beyond cell; 
cell partially open (only a rudiment of discocellular 
vein), long, a trifle more than half as long as wing. 
Eighth abdominal segment of male simple. 

Male genitalia with apical process of gnathos devel- 
oped as a stout, curved, bluntly pointed hook. Uncus 
stout, hoodlike, scarcely tapering and with broadly 
rounded terminalmargin. Transtilla complete, a bridge 
with central furcate projection, not appreciably arched. 
Harpe with costa produced and angulate at middle. 
Anellus a U-shaped plate with rather short lateral arms. 
Aedeagus rather slender, simple, slightly curved; penis 
armed with a thin elongate sclerotized band. Vincu- 
lum moderately stout, about as broad as long, triangu- 
late. 

Female genitalia with bursa copulatrix weakly and 
sparsely scobinate; signa present as a cluster of (3 or 4) 
small, weakly sclerotized discs. Ductus bursae sclero- 
tized, granulate and flattened for two-thirds of its length 
from genital opening and with a produced, angulate, 
sclerotized shield behind genital opening. Ductus sem- 
inalis from bursa near signa (near anterior end of bursa). 

A distinct genus showing affinities, especially in fe- 
male genitalia, to Hphestiodes and several of the follow- 
ing genera. Contains one tropical American species. 


571. Micromescinia pygmaea Dyar 
Ficures 96, 595, 1083 
Micromescinia pygmaea Dyar, Proc. U. 8S. Nat. Mus., vol. 47, 
p. 347, 1914. 

Forewing pale ocherous with a narrow purplish red 
shade along costa and a similarly colored broad suffu- 
sion along termen. Hind wing pale smoky fuscous, paler 
towards base. Alar expanse, 9-10 mm. 

Genitalia as given for the genus; cucullus of harpe 
angulate, apex bluntly pointed. 

Type tocauity: Porto Bello, Panamé (type in 
USNM). 

Foop piant: Unknown. 

Distripution: PanamA: Porto Bello (Mar.), Taber- 
nilla, Taboga Isl. (Feb.). 

Known only from the original type series (two males 
and one female) in the National Collection. 


278 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Genera 170-177: Ephestiodes to Moodnella 


[Venational division C. Forewing with 10 veins; 8 and 9 united 
(except in aberrant individuals); male with costal fold. Labial 
palpi oblique. Male abdomen with compound dorsal tufts on 
eighth segment or simple. Transtilla complete or incomplete; 
if complete, an angulate bridge; if incomplete, its elements well 
developed. Ductus bursae sclerotized for at least part of its 
length from genital opening and with projecting sclerotized shield 
behind genital opening.] 


170. Genus Ephestiodes Ragonot 


Ephestiodes Ragonot, N. Amer. Phycitidae, p. 16, 1887; Mono- 
graph, pt. 2, pp. xiii, 264, 1901.—Hulst, Phycitidae of N. 
Amer., p. 194, 1890.—Forbes, Cornell Mem. 68, p. 635, 1923. 
(Type of genus: Ephestiodes gilvescentella Ragonot.) 

Tongue well developed. Antenna simple, pubescent. 

Labial palpus oblique, reaching to vertex, somewhat 

rough scaled; segment 3 about aslong as 2. Mazxillary 

palpus filiform. Forewing smooth; 10 veins; vein 2 

from very near to lower outer angle of cell; 3 from the 

angle; 4 and 5 stalked, the stalk approximate to or con- 
nate with 3 at base; 6 from below upper angle of cell, 
straight or very slightly curved; 8 and 9 united (9 ab- 

sent); 10 from the cell, separated from 8; male with a 

short costal fold. Hind wing with vein 2 from before 

(but near) lower outer angle of cell; 3 and 5 long stalked; 

7 and 8 anastomosed almost to costa (8 a mere vestige) ; 

cell about half the length of wing; discocellular vein 

curved. Highth abdominal segment of male with com- 
pound dorsal scale tufts (except in noniella). 


Male genitalia with apical projection of gnathos 
forked, U- or V-shaped. Uncus broad and with apical 
margin broadly rounded. ‘Transtilla incomplete (ex- 
cept in stictella and noniella) but with the elements 
strongly developed, their apices approximate and broad- 
ly flared. Harpe normally (except in Jucidibasella, 
stictella, and noniella) with a transverse sclerotized ridge 
across base of cucullus; costa strongly sclerotized but 
not produced. Anellus a narrow, semicircular band 
with flanged base. Aedeagus straight, moderately long; 
penis armed (except in noniella) with a weak, thin, flat 
bladelike cornutus or a pair of similar cornuti. Vincu- 
lum stout, triangulate. 


Female genitalia with signum developed as a short 
comb of from two to six blunt, short, thornlike teeth, or 
as a flat plate with a row of such teeth along one edge 
(plorella); bursa copulatrix otherwise simple except for 
a few weak scobinations surrounding signum or at 
junction of ductus bursae. Ductus bursae longer than 
bursa (except in stictella), sclerotized, granulate and 
flattened for at least half (normally for most) of its 
length from genital opening, with a produced, strongly 
sclerotized shield behind genital opening and usually 
with a sclerotized band or shield below the opening; 
ductus seminalis from bursa close to signum. 

The North American species of the genus form a com- 
pact group agreeing on all structural characters. In 
tropical America, however, there is considerable varia- 
tion from type and two of the species (stictella and 
noniella) are distinctly aberrant on male characters. 


Eventually it may be possible to give them separate 
generic designations, but at present there are no charac- 
ters to be found (apart from male genitalia) to permit 
such separation. The species divide into three groups 
as follows: 


Transtilla incomplete, but strongly developed, with apices 
flared and approximate. Harpe normally with a trans- 
verse sclerotized ridge at base of cucullus. Eighth ab- 
dominal segment of male with tufts. (Comprising all 
the typical E’phestiodes.) 

Transtilla complete, with greatly produced, asymmetrical, 
caudal projections. Harpe with apex of sacculus pro- 
duced into a free arm. Highth abdominal segment of 
male with tufts. (Tropical only.) 

Transtilla complete, a narrow band with central loop. 
Harpe simple. Eighth abdominal segment of male 
simple. (Tropical only.) 


While adult specimens seem to be abundant and are 
often taken in numbers, little if anything is known of 
life histories within the genus and none of the species 
has, so far, proven to be of any economic importance. 
Most of the species are similar in color and maculation, 
but each shows some distinguishing difference in male 
genitalia. These are not easy to describe, but can be 
seen readily enough in the drawings. The female geni- 
talia of the North American species are remarkably 
uniform in general structure, exhibiting more variation 
within than between species. In all of them the dorsal 
sclerotized area of the eighth-segment collar is reduced 
to a narrow band, the sclerotized shield behind genital 
opening has a central angulate projection from its pos- 
terior margin, and the sclerotized band below the open- 
ing is narrow. 


Genus Ephestiodes, Species 572-580: E. gilves- 
centella to E. plorella 


[Transtilla incomplete but strongly developed, with apices flared 
and approximate. Harpe normally with a transverse sclerotized 
ridge at base of cucullus. Eighth abdominal segment of male 
with tufts.] 


572. Ephestiodes gilvescentella Ragonot 
Figures 108, 589, 1072, 1073 


Ephestiodes gilvescentella Ragonot, N. Amer. Phycitidae, p. 16, 
1887; (Ragonot and Hampson), Monograph, pt. 2, p. 264, 
1901.—Hulst, Phycitidae of N. Amer., p. 194, 1890.— 
McDunnough, Check list, No. 6379, 1939. 

Ephestiodes nigrella Hulst, Journ. New York Ent. Soc., vol. 8, 
p. 224, 1800.—Donohoe and Barnes, Journ. Econ. Ent., 
vol. 27, p. 1071, 1984.—McDunnough, Check list, No. 6382, 
1939. (New synonymy.) 

Forewing pale gray, the median area (between the 
transverse lines) dark gray; antemedial line outwardly 
angled at middle and with a very slight notch at lower 
fold, white with a narrow blackish outer border; sub- 
terminal line fainter, slightly sinuate and parallel with 
termen, whitish with a narrow blackish inner border; 
discal dots at end of cell, separate, distinct, blackish; 
terminal dots obscure, more or less confluent. Hind 
wing white to pale smoky fuscous; the veins but faintly 
darkened. Alar expanse, 12-16 mm. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 279 


Male genitalia with apical process of gnathos 
V-shaped, the prongs narrowly triangulate. Harpe 
with strong transverse sclerotized ridge at base of 
cucullus and projecting slightly beyond lower margin 
of harpe; costa smooth; apex angled. ‘Transtilla with 
flaring apices of its elements divergent. Aedeagus 
moderately stout. 

Female genitalia show some individual variation in 
the number of teeth in the signum (4 to 6). The shape 
of the shield behind genital opening is also variable 
(compare figs. 1072 and 1078). 

Type Locaitizs: California (gilvescentella, in Paris 
Mus.); Los Angeles, Calif. (nigrella, in USNM). 

Foop piants: Raisins, cotton, Gaura parviflora. 
These food-plant records from reared specimens in the 
National Collection. Moths have been frequently seen 
about stored unprocessed raisins in California and have 
been reared from larvae feeding on the raisins, but 
apparently doing only minor injury. The life history 
has not been worked out. The habits of the larvae are 
presumably similar to those of Vitula and Ephestia. 

Distrisution: Unitep States: Colorado, Denver; 
Utah, Eureka (July, Aug.), Provo (June, July, Aug., 
Sept.), Stockton (July), Vineyard (June); New Mexico, 
Mesilla (Apr., July); Arizona, Fish Creek Station 
(Sept.), Tempe (Apr.); California, Blythe (Sept.), 
Claremont, Fresno (July), Laguna Beach, Loma Linda 
(July), Los Angeles (Mar., Apr., July), Pasadena 
(June), Riverside (Apr., June, July), Sacramento 
(Mar.), San Diego (Mar., May, June), San Gabriel 
(July) ; Washington, Almota (Apr.), Bellingham (Aug.), 
Pullman (June, July, Aug., Sept.), Snake River (June), 
Wenatchee (Aug.). Canapa: British Columbia, Kere- 
meos (Aug.), Seton Lake (Aug.), Victoria, Wellington. 
México: Baja California, Ensenada (Apr.) 

The name nigrella applies only to a suffused color 
form in which the dark dusting on the central area of 
forewing extends into the terminal area. In series from 
any given locality intergrades occur between it and 
typical gilvescentella. The species appears to be con- 
fined to the North American region from the Rocky 
Mountains westward. 


573. Ephestiodes infimella Ragonot 
Fieure 590 


Ephestiodes infimella Ragonot, N. Amer. Phycitidae, p. 16, 1887; 
(Ragonot and Hampson), Monograph, pt. 2, pp. 264, 265, 
1901.—Hulst, Phycitidae of N. Amer., p. 194, 1900.— 
Forbes, Cornell Mem. 68, p. 635, 1923.—McDunnough, 
Check list, No. 6380, 1939. 

Forewing similar in pattern and color to the more 
suffused specimens of gilvescentella but averaging 
smaller. In many specimens, especially those where the 
basal area is pale and contrasted against the rest of the 
wing, there is more or less shading of reddish luteous; 
color as variable as in gilvescentella. Hind wing pale to 
dark smoky fuscous. It is the common species in 
eastern and central United States. Alar expanse, 
10-15 mm. 


Male genitalia with the prongs of apical] process of 
gnathos slender, cylindrical, and widely spaced at base. 
Harpe with the transverse, sclerotized ridge at base of 
costa not projecting beyond lower margin of harpe; 
costa irregularly serrate at middle (a variable and not 
too reliable character, costa however never entirely 
smooth under high magnification); apex rounded. 
Transtilla with apices of its elements divergent at their 
apices but less flaring than those of gilvescentella. 
Aedeagus moderately stout. Female genitalia not 
essentially different from those of gilvescentella. 

Type Locauity: North Carolina (type in Paris Mus.). 

Foop pxrants: Wild cherry, seeds of Ambrosia 
(Forbes). The wild cherry record is from a specimen 
reared from larvae collected by A. Busck at Cape 
Henry, Va. 

Distripution: Unitep States: Massachusetts, Mar- 
tha’s Vineyard (Aug.); Connecticut, East River; New 
Jersey, Anglesea (June), Montclair (Aug.); Pennsyl- 
vania, Hazleton (July), Oak Station (July) ; Maryland, 
Hyattsville (Aug.), Plummers Isl. (May, July) ; District 
of Columbia, Washington (Aug.); Virginia, Cape Henry 
(July) ; North Carolina, Tryon (Aug.); Kentucky (Aug.) ; 
Illinois, Chicago (June), Putnam County (May, Sept.) ; 
Iowa; Missouri, St. Louis (June); Arkansas, Washing- 
ton County (July); Teras, Burnet County. Also re- 
corded by Hampson from Colombia; but this record is 
undoubtedly based on a misidentification. From all 
available evidence the species is limited in its distribu- 
tion to North America. 


574, Ephestiodes erythrella Ragonot 
Figures 591, 1074 


Ephestiodes erythrella Ragonot, N. Amer. Phycitidae, p. 16, 1887; 
(Ragonot and Hampson), Monograph, pt. 2, pp. 264, 266, 
1901.—Hulst, Phycitidae of N. Amer., p. 195, 1900.— 
McDunnough, Check list, No. 6384, 1939. 

Eurythmia coloradella Hulst, Canadian Ent., vol. 32, p. 175, 
1900.—McDunnough, Check list, No. 6390, 1939. (New 
synonymy.) 

Ephestiodes benjaminella Dyar, Proc. U. 8S. Nat. Mus., vol. 27, 
p. 922, 1905.—McDunnough, Check list, No. 6383, 1939. 
(New synonymy.) 

Forewing vinous red, dusted with whitish gray in the 
basal area and with some faint blackish dusting along 
costa and on the veins; in the dry areas of Colorado and 
Utah the ground color somewhat paler, with consider- 
able whitish gray dusting in the area between the trans- 
verse lines and little or no appreciable blackish dusting 
on costa and veins; transverse lines whitish, somewhat 
more distinct than in gilvescentella and similarly shaped 
and dark margined (in some specimens they appear 
curved, in others straight, due to the extent of the black- 
ish shade bordering them, but in reality slightly angled 
as in the other North American species) ; discal dots ob- 
scure. Hind wings pale to moderately dark smoky 
fuscous. Alar expanse, 13-17 mm. 

Male genitalia similar to those of infimella except 
apex of harpe angulate. Female genitalia showing no 
distinctive specific characters. 


280 


Typr wocatitius: California (erythrella, in Paris 
Mus.); Colorado (Coloradella, in AMNH, ex Rutgers); 
Kaslo, British Columbia (benjaminella, in USNM). 


Foop pLant: Unknown. 


DistrisutTion: Unitep States: Colorado; Wyoming, 
Jackson Hole (June); Utah, Eureka (May), Provo 
(June, July); California, Alfa (Placer County, July), 
Baxter (Placer County, July), Camp Baldy (San Ber- 
nardino Mts., July), Makehlumne Hill (June); Wash- 
ington, Pullman (June, July, Aug.). Canapa: British 
Columbia, Goldstream (Sept.), Kaslo (June, July, Aug.), 
Wellington. A long series in the National Collection 
from Utah. 


Like gilvescentella and infimella, erythrella is variable 
in coloration and intensity of markings and in the shape 
of the terminal margin of the sclerotized shield behind 
genital opening of the female. Extremes of individual 
variation in the shield are shown in figures 1074a, b. 
The same amount of variation occurs in gilvescentella. 
It has much the same range as the latter, but need not 
be confused with it, for in dark or light specimens it can 
be readily identified by its distinctly reddish ground 
color. Occasional specimens of injimella are also red- 
dish but that species does not occur in the same areas 
as erythrella. I am unable to find any character of 
specific value to distinguish either coloradella or ben- 
gaminella from California examples of erythrella,so I am 
sinking the names in synonymy. 


575. Ephestiodes mignonella Dyar 
Fiaures 592, 1075 


Ephestiodes mignonella Dyar, Proc. Ent. Soc. Washington, vol. 
10, p. 113, 1908.—McDunnough, Check list, No. 6381, 
1939. 

Forewing very pale grayish white; a blackish streak 
on costa at base and, in many specimens, a subbasal 
black streak on median vein; transverse lines as in pre- 
ceding species, white, the antemedial with a strongly 
contrasted, outer black band and the subterminal with a 
narrower inner black band, especially pronounced on 
costal half of wing (sometimes the black borders partially 
obscured on lower half of wing by pale putty-colored 
scaling); discal dots fused, black; terminal dots fainter, 
confluent. Hind wings whitish, with a smoky tinge 
at apex, along termen, and faintly on the veins, espe- 
cially on the females. Alar expanse, 12-14 mm. 

Male genitalia similar to those of injfimella except: 
Harpe with apex angled; enlarged apices of elements of 
transtilla parallel and facing each other. Female gen- 
italia showing no distinctive specific characters. 

TypE Locauity: Kerrville, Tex. (type in USNM). 

Foop pLrant: Unknown. 

Represented by 13 specimens from the type locality 
in the National Collection (Apr.). 

The species is easily distinguished by the very pale 
ground color and strongly contrasted black markings of 
forewing. 


UNITED STATES NATIONAL 


MUSEUM BULLETIN 207 


576. Ephestiodes erasa, new species 


Fieures 588, 1077 


Forewing very pale gray with some pale putty- 
colored scaling on lower half, especially towards base; 
transverse lines faint, distinguishable only under low- 
powered magnification; their dark borders nearly 
obliterated; discal dots very faint, obsolescent in some 
specimens. Hind wings smoky white, little paler than 
forewing. Alar expanse, 11-13 mm. 

Male genitalia with transverse ridge projecting 
beyond lower margin of harpe (more so than in gilvescen- 
tella); lower margin of cucullus incurved just beyond 
transverse ridge; cucullus somewhat reduced. Female 
genitalia exhibiting no distinguishing specific characters. 

Tyrer tocatity: Lake Alfred, Fla. (type m USNM, 
61387). 

Foop prant: Unknown. 

Described from male type and six female paratypes 
from the type locality, May 6 and July 2, 1929, L. J. 
Bottimer, collector. In addition to the type series I 
have before me a female from Gainesville, Fla. (Apr.), 
D. M. Bates, and one other female from Winter Park, 
Fla. Guly 23, 1939), H. Fernald. All the foregomg 
were collected at light. 

The species is at once distinguished by its pale, almost 
unmarked forewings. 


577. Ephestiodes lucidibasella Ragonot 
Fieures 593, 1081 


Ephestiodaes lucidibasella Ragonot, Nouv. Gen., p. 35, 1888; 
Monograph, pt. 2, pp. 264, 265, 1901. 

Forewing rosy gray, the median and terminal areas 
heavily and evenly dusted with blackish; the transverse 
lines decidedly oblique, widely spaced on costa. Rago- 
not’s figure (Monograph, pl. 34, fig. 6) shows a species 
with an extension of the pale ground color along the 
lower fold, and according to Clarke’s notes the figure is 
a good representation of the type. Hind wings semi- 
transparent, whitish with a slight smoky tint. Alar 
expanse, 16 mm. 

Gnathos of male genitalia with prongs of apical 
process somewhat longer than those of the North Amer- 
ican species, slender, widely spaced at base and curving 
apart slightly toward their apices. Harpe without 
trace of asclerotized, transverse ridge at base of cucullus; 
edge of costa minutely and irregularly serrate; apex 
evenly rounded. ‘Transtilla with the enlarged apices 
of its elements concave, parallel, and facing each other. 
Female genitalia distinguished by the shape of the 
sclerotized shield from ductus bursae behind genital 
opening. 

TYPE LOCALITY: Quillota, Chile (type in Paris Mus.). 

Foop piant: Unknown. 

All known specimens are from Chile. In the Paris 
Museum, according to Clarke’s notes, are three males 
from the type locality and one male from Valparaiso. 
A female paratype is in the British Museum. There 
are no examples in the U. S. National Museum. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 281 


578. Ephestiodes productella Ragonot 
Ficure 1078 
Ephestiodes productella Ragonot, Nouv. Gen., p. 36, 1888; Mono- 
graph, pt. 2, pp. 264, 265, 1901. 

This species is known only from the female type in 
Paris. According to Clarke’s note it answers very well 
to Ragonot’s description but not to his figure in the 
Monograph (pl. 35, fig. 1). 

The genitalia are characteristic and should easily 
identify the species when other females are discovered. 
The shield behind genital opening has an angulate pro- 
jection from posterior margin as in North American 
species, but is appreciably larger and the sclerotized 
band below genital opening is much stouter and broader. 
Alar expanse, 15 mm. 

TYPE LocaLity: Given by Ragonot as “Am. Mér.” 
in original description, and as Chiriqui, Colombia, in 
Monograph (type in Paris Mus.). Nothing on type to 
indicate the exact locality, but undoubtedly from 
tropical America. 

Foop prant: Unknown. 


579. Ephestiodes indentella Dyar 
Figure 1076 


Ephestiodes indentella Dyar, Ins. Insc. Menstr., vol. 3, p. 89, 1915. 


Forewing pale gray, the median area shading with 
blackish scaling; the transverse lines widely spaced on 
costa, strongly contrasted (especially the antemedial), 
white; antemedial line decidedly oblique, notched at 
lower fold and with a black outer border; subterminal 
indented at vein 6 and slightly at lower fold, and with 
a narrow black inner border; discal dots separate; ter- 
minal dots confluent, not extending to either costa or 
tornus. Hind wing pale smoky fuscous; veins and 
terminal margin but slightly darkened. Alar expanse, 
12.5-17 mm. 

Female genitalia with a wide, deep notch in posterior 
margin of shield behind genital opening; sclerotized 
band below genital opening narrow, sinuous; eighth-seg- 
ment collar broadly and deeply sclerotized on dorsum, 
its anterior margin produced and rounded. 

Male unknown. 

TyprE Locauity: Bermuda (type in USNM). 

Foop prant: Unknown. 

Known only from Bermuda. In addition to the type 
series (Apr.) I have before me 6 females from the 
British Museum (Mar., Apr.). The species is readily 
identified by its genitalia. It and plorella are tenta- 
tively assigned to the first Ephestiodes species group, 
but accurate placement of them will have to wait upon 
discovery of males. 


580. Ephestiodes plorella Dyar 
Ficures 109, 1080 


Ephestiodes plorella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 344, 
1914. 

Eurythmia vestilla Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 339, 
1914 (new synonymy). 


Forewing rather pale brownish gray, uniformly 
300329—56——19 


colored except for a somewhat paler basal area and a 
rather broad brownish outer band along the antemedial 
line; antemedial band but slightly oblique; subterminal 
line obscure, whitish with only faint trace of a dark 
inner border; discal dots obscure, when distinguishable, 
more or less confluent; terminal dots faint, confluent. 
Hind wing whitish, some of the veins and terminal area 
very faintly darkened. Alar expanse, 11-14 mm. 

Female genitalia with signum developed as a flat plate 
with serrate edge, the ductus seminalis arising from a 
hole in the plate; bursa with a few scobinations at junc- 
tion with ductus bursae; ductus bursae sclerotized for 
only half its length; no sclerotized band below genital 
opening; posterior margin of large sclerotized shield 
behind genital opening evenly rounded except for a very 
slight narrow central notch; collar of eighth segment 
natrow but deeply sclerotized on dorsum, its produced 
anterior margin concave and with asmall shallow central 
notch. 

Male unknown. 


Type Locauity: Corazal, Canal Zone, Panama 
(plorella and vestilla, in USNM). 

Foop pirant: Unknown. 

A rather suffused, poorly marked species without much 
to distinguish it superficially from faded examples of our 
North American gilvescentella or infimella. The female 
genitalia, however, are markedly different from those of 
any other species in the genus. The absence of a sclero- 
tized band below genital opening, the shapes of the dor- 
sal genital plate and eighth-segment collar, and the pe- 
culiar signum at once distinguish them. The venation 
also differs in that vein 3 of forewing is very closely ap- 
proximate to, connate, or short stalked with 4-5, and 
the latter are very long stalked. Dyar’s vestilla was 
placed by him in Zurythmia on the belief that veins 4 
and 5 of forewing were united as they appear to be on 
superficial examination; but denuded wings (fig. 109) 
show them long stalked. The female genitalia of his 
type of vestella are identical with those of his plorella. 


Genus Ephestiodes, Species 581: ZH. stictella 


[Transtilla complete, with greatly produced, asymmetrical, 
caudal projections. Harpe with apex of sacculus produced into 
afree arm. Highth abdominal segment of male with tufts.] 


581. Ephestiodes stictella (Hampson), new combination 
Figure 596, 1082 

Unadilla stictella Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7, 
p. 255, 1901. 

Ephestiodes uniformella Hampson, Ann. Mag. Nat. Hist., ser. 7, 
vol. 7, p. 256, 1901 (new synonymy). 

Ephestiodes granulella Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 
7, p. 256, 1901 (new synonymy). 

The type of stictella is afemale. A photograph of its 
genitalia definitely fixes the species to what I here iden- 
tify in long series from the Cornell Collection from Puerto 
Rico and St. Croix. The type of uniformella is a male 
without abdomen and according to Clarke a stained spec- 
imen., A photograph shows it to be a dark specimen but 
no darker than many examples of the St. Croix series. 


282 


The type of granulella is a male and a photograph of its 
genitalia fixed it. I have before me other specimens of 
stictella from the Bahamas and of granulella from Ja- 
maica that agree in all characters. Hampson’s reference 
of stictella to Unadilla ignored the diagnostic venational 
character of that genus, the vertical discocellular vein of 
hind wing. In stictella the discocellular vein is obviously 
curved. ‘The species is variable in color, ranging from 
very dark to whitish gray, the ground color (dark or 
light) being rather uniform over the forewing, the basal 
area no darker or lighter than the median and terminal 
areas. Antemedial band rather broad, whitish, oblique 
and nearly straight, outwardly bordered on costal half 
by a narrow blackish line; subterminal line narrow, paral- 
lel and near to termen, slightly irregular, whitish bor- 
dered inwardly towards costa by a thin, faint, blackish 
line; discocellular spots more or less obsolescent, when 
distinct, separate and blackish. Hind wing whitish to 
pale smoky fuscous, shaded with smoky fuscous towards 
apex and termen. Alar expanse, 10-12 mm. 

Male genitalia with transtilla elements fusing at a 
point and thence developed into two extended, curving, 
asymmetrical arms; harpe with sacculus produced at ex- 
tremity into a clasperlike free arm; costa smooth; apex 
of harpe bluntly pointed; vinculum triangulate, shorter 
than broad; aedeagus rather slender. Female genitalia 
with ductus bursae shorter than bursa, sclerotized 
throughout its length; a narrow flaring sclerotized plate 
below genital opening; eighth-segment collar divided, its 
apophyses separated from the dorsal part and firmly at- 
tached to the broad shield behind genital opening (fig. 
1082b).; dorsal portion of collar (ig. 1082a) produced an- 
teriorly into an elongate tongue and laterally into curved, 
projecting arms. 

Type Locaurry: Nassau, Bahamas (stictella, uniform- 
ella, and granulella, in BM). 

Foop prant: Unknown. 


DistriputTion: Banamas: Nassau. Jamaica, Runa- 
way Bay (Mar.). Purrro Rico: Coamo Springs (Apr.), 
Palmas Abajas (June, July), San German (Apr., July), 
Puerto Real (Vieques Isl., Apr.). Virein IsLanps: 
Kingshill (St. Croix; May, Oct., Nov., Dec.). 

This species on characters of the genitalia seems to de- 
serve a separate generic designation; but until the males 
of the other tropical species (described from females) are 
known it seems best to retain it in H’phestiodes, with 
which it agrees on all other than genitalic characters. 


Genus Hphestiodes, Species 582: EH. noniella 
[Transtilla complete, a narrow band with central loop. Harpe 
simple. Eighth abdominal segment of male simple.] 


582. Ephestiodes noniella Dyar 
Figures 597, 1079 
Ephestiodes noniella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 345, 
1914. 


Forewing (to the naked eye) gray, the basal area with 
an olivaceous tint; under magnification the remainder 
of the wing shows a strong dusting of rosy scaling; ante- 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


medial line faint, oblique, narrow, whitish, and faintly 
bordered outwardly near costa by a narrow dark line; 
subterminal line almost obliterated; discal dots not dis- 
tinguishable. Hind wing pale smoky fuscous; the veins 
and terminal margin but slightly darkened. Wighth 
abdominal segment of male simple. Alar expanse, 
9.5-10 mm. 

Male genitalia with the prongs of apical process of 
gnathos well separated but somewhat reduced. Trans- 
tilla complete, a narrow band with a central loop. 
Harpe simple; costa smooth; apex narrowly rounded. 
Aedeagus short; penis without armature. Female geni- 
talia like those of typical Ephestiodes except: Shield 
behind genital opening smaller; sclerotized band below 
genital opening developed as a half oval, caudally pro- 
jecting shield. 

Typn tocauity: La Chorrera, Panamd (type in 
USNM). 

Foop prant: Unknown. 


Known only from the type series, two females from 
the type locality (May) and one malefrom Rio Trinidad, 
Panam4 (Sept.). 

The species is an abnormal Hphestiodes on male char- 
acters (the complete, thin, looped transtilla, lack of 
cornutus on penis, and absence of tufts on eighth 
segment), but separate generic designation does not 
seem warranted at this time. 


171. Genus Azaera Schaus 


Azaera Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, p. 250, 
1918. (Type of genus: Azaera muciella Schaus.) 

Calamophleps Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 342, 1914. 
(Type of genus: Calamophleps squalidella Dyar.) 

Characters of typical H’phestiodes except vein 3 of 
forewing from the stalk of 4—5. 

This genus is of doubtful status. There are nothing 
but differences of specific value in the genitalia to 
separate it from the type of Hphestiodes and, in the 
latter genus, occasional specimens of gilvescentella have 
vein 3 of forewing connate cr even very shortly stalked 
with 4-5. However, none that I have seen (and | have 
examined long series of H’phestiodes) shows such short 
stalking on both forewings and none exhibits the con- 
siderable stalking of Azaera (see fig. 110). Asa definite 
stalking of 3 with 4 and 5 is a rare character in New 
World Phycitinae, we had better retain the generic 
separation for the present. 

Dyar was in error in assuming that Azaera Schaus 
was a homonym of Azara D’Orbigny, so Schaus’ name 
must take precedence over Calamophleps. 


583. Azaera muciella Schaus 


Figures 110, 594, 1084 


Azaera muciella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, 
p- 250, 1913. 

Calamophleps squalidella Dyar, Proc. U. 8. Nat. Mus., vol. 47, 
p. 342, 1914. 


Forewing color and markings similar to those of 
Ephestiodes plorella Dyar except that subterminal line 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 


is always distinguishable. The species can be identified 
satisfactorily only by its venation and genitalia. Hind 
wing pale smoky fuscous; veins and terminal margin 
darker. Alar expanse, 11-13 mm. 

Male genitalia with prongs of apical process of gnathos 
rather stout and curved towards each other at their 
apices. Apices of elements of transtilla greatly de- 
veloped, widely flaring. Harpe with transverse ridge 
at base of cucullus projecting beyond lower margin of 
harpe; lower margin of cucullus with a slight notch at 
apex. Female genitalia with sclerotized band below 
genital opening forming a rather broad oval shield; 
ductus bursae much longer than bursa, sclerotized for 
no more than half its length; dorsal sclerotized portion 
of eighth-segment collar a rather broad band. 

TYPE LOCALITIES: Juan Vifias, Costa Rica (muciella, 
in USNM), Porto Bello, Panama (squalidella, in USNM). 

Foop prant: Unknown. 

DistrisuTion: Costa Rica: Juan Vifias (Feb.). 
GuatEmata: Cayuga (May). PanamA: Corozal (Mar. 
Apr., May), La Chorrera (May), Porto Bello (May, 
Dec.). 

Dyar acknowledged the synonymy of muciella and 
squalidella in his description of the latter. It is borne 
out by their genitalia. Both types are females. 


584. Azaera nodoses (Dyar), new combination 
Ficure 1085 
mea nodoses Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 348, 
4, 

Smaller and paler than muciella, with the dark borders 
indicating the transverse lines fragmentary and obscure; 
but chiefly distinguished by its genitalia. Alar expanse, 
10 mm. 

Female genitalia with ductus bursae short (no longer 
than bursa) and sclerotized for its entire length; dorsal 
sclerotized portion of eighth-segment collar a narrow 
band. 

Type tocauity: Corozal, Panamé (Apr., type in 
USNM). 

Foop prant: Unknown. 

Known only from the unique female type. 


585. Azaera lophophora (Dyar), new combination 


Calamophleps lophophora Dyar, Proc. U. 8. Nat. Mus., vol. 47, 
p. 343, 1914 

Similar to muciella but darker, smoky gray; the 
transverse lines and their dark borders more distinct. 
May be only a dark form of muciella; but this cannot 
be determined until more specimens are available, as 
the two females of the type series (the only specimens 
known) lack abdomens. Alar expanse, 11-12 mm. 

Typ Locatity: Porto Bello, Panamdé (May, type in 
USNM). 

Foop piant: Unknown. 

The female paratype is from La Chorrera, Panama 


(May). 


283 


172. Genus Moodna Hulst 


Moodna Hulst, Phycitidae of N. Amer., p. 193, 1890.—Ragonot, 
Monograph, pt. 2, p. xiii, 1901—Hampson, in Ragonot, 
Monograph, pt. 2, p. 267, 1901.—Forbes, Cornell Mem. 68, 
p. 636, 1923. (Type of genus: Moodna pelviculella Hulst.) 

Tongue well developed. Antenna pubescent; shaft 
of male with a short, shallow sinus towards base. 
Labial palpus oblique, subcylindrical, reaching vertex, 
somewhat rough scaled; third segment nearly as long 
as second. Maxillary palpus filiform, short. Fore- 
wing smooth; 10 veins; vein 2 from before, but rather 
near, lower outer angle of cell; 3 from the angle; 4 and 5 
stalked (for at least half their lengths), the stalk shortly 
separated from 3 at base; 6 from below upper angle!of 
cell, straight; 8 and 9 united (9 absent); 10 from the 
cell, separated from 8 at base; male with a strong costal 
fold enclosing hair tuft. Hind wing with vein 2 from 
well before lower outer angle of cell; 3 and 5 from the 
angle, connate, rarely (in individual specimens) very 
shortly stalked; 7 and 8 anastomosed to or almost to 
costa (8 absent or a mere vestige at costa); cell less than 
half (but more than a third) the length of the wing; 
discocellular vein curved. Eighth abdominal segment 
of male with compound, dorsal tufts. 

Male genitalia with apical projection of gnathos a 
broad, short, pointed hook. Uncus broad; terminal 
margin broad, straight or but slightly rounded. Trans- 
tilla incomplete. Harpe with apex broadly rounded; 
two strongly sclerotized subbasal projections from 
costa. Anellus a narrow semicircular sclerotized band. 
Aedeagus long, straight, not appreciably tapering, 
simple; penis bearing one or two elongate, thin, weakly 
sclerotized plates and a few sclerotized wrinklings, 
otherwise unarmed. Vinculum long, triagulate. 

Female genitalia with signum a small cluster of 
rounded, short, projecting discs; bursa copulatrix small, 
wrinkled, the wrinklings weakly sclerotized; ductus 
bursae long (much longer than bursa), sclerotized for a 
short distance from genital openings; ductus seminalis 
from bursa close to signum; an extended ventral lobe 
from membranous area between ovipositor and eighth- 
segment collar (figs. 1086a and 1087a). 

The species of the Moodna-Vitula complex offer a 
difficult problem in generic placement. Typical species 
of either genus are obviously different in characters of 
venation, genitalia, and male antennae; but there are 
a number of aberrant species that possess characters of 
both Moodna and Vitula and will fit in neither genus. 
If we should attempt to unite all under one genus, we 
should have a group that could not be defined. I have 
therefore divided the complex into small units, resur- 
recting Hulst’s Manhatta and erecting separate genera 
for some new species and the aberrant tropical species 
that have been referred to Moodna, or misplaced else- 
where. One name that appears under Moodna in our 
lists (Z’phestia nigrella Hulst) has veins 4 and 5 of fore- 
wing united and is treated elsewhere in this paper as 
Caudellia nigrella (p. 293). It has the habitus of 
Moodna, similar male antennae, is similar in color and 
maculation to its type, and is apparently derived from 


284 


Moodna; but is structurally an advanced development. 
As here defined, Moodna is limited to two species from 
the United States and México (ostrinella Clemens and 
bisinuella Hampson). The genus is easily distinguished 
from other genera in venational group C by the follow- 
ing combination of characters: Male antenna with a 
shallow sinus in shaft; eighth abdominal segment of 
male with tuft; hind wing with vein 2 from well before 
angle of cell and veins 3-5 connate or very shortly 
stalked; gnathos terminating in a short, simple hook; 
transtilla incomplete. 


Two species (Moodna olivella Hampson and Hornigia 
clitellatella Ragonot) that Hampson (Ragonot Mono- 
graph, pt. 2, pp. 268, 269) includes in the genus are un- 
known to me, and I am unable to place them generically. 
They are treated briefly at the end of this paper. 

On the strength of the supposed synonymy of Man- 
hatta Hulst (=Hornigia Ragonot, 1887), Hampson 
cites the European biviella Zeller as the type of Moodna. 
This, of course, is inadmissible, for not only was biviella 
not among the species originally included in Moodna, 
but in describing his genus Hulst designated pelviculella 
as its type. 

586. Moodna osirinella (Clemens) 
Fieures 114, 599, 1086 


Ephestia ostrinella Clemens, Proc. Acad. Nat. Sci. Philadelphia, 
1860, p. 206.—Hulst, Phycitidae of N. Amer., p. 220, 1890. 

Horginia obtusangulella Ragonot, N. Amer. Phycitidae, p. 16, 
1887. 

Moodna pelviculella Hulst, Phycitidae of N. Amer., p. 194, 1890. 

Manhatta obtusangulella (Ragonot) Hulst, Phycitidae of N. 
Amer., p. 197, 1890. 

Moodna obtusangulella (Ragonot) Hampson, in Ragonot, Mono- 
graph, pt. 2, p. 269, 1901. 

Manhatta ostrinella (Clemens), Hulst, U. S. Nat. Mus., Bull. 52, 
p. 486, 1903. 


Moodna ostrinella (Clemens) Barnes and McDunnough, Check 
list of the Lepidoptera of Boreal America, No. 5795, 1917.— 
Forbes, Cornell Mem. 68, p. 636, 1923.—McDunnough, 
Check list, No. 6896, 1939.—Darlington, Trans. Amer. Ent. 
Soce., vol. 73, p. 91, 1947. 


Forewing blackish fuscous; basal area strongly shaded 
with a deep violet red; terminal area also more or less 
shaded with the same reddish color; antemedial le 
distinct, rather wide, nearly straight, white; subterminal 
line obscure, parallel to termen, slightly and irregularly 
denticulate; discal spots distinct, more or less confluent 
and usually set off by some surrounding whitish dusting 
which extends, in well marked specimens, to costa. 
Hind wing smoky white to pale fuscous; veins darkly 
outlined and a narrow dark shade along terminal mar- 
gin. Alar expanse, 11-17 mm. 


Male genitalia with inner costal projection from harpe 
long, slender, tapering, vertical from costa; outer pro- 
jection curved outwardly, the two projections pointed 
away from each other. Uncus with sides nearly paral- 
lel. Vinculum evenly tapering. Female genitalia with 
ductus bursae weakly sclerotized toward genital opening. 

Typ LOCALITIES: Pennsylvania [?] (ostrinella, type 
lost); Texas (obtusangulella, in Paris Mus.); Newburgh, 
N. Y. (pelviculella, in AMNH, ex Rutgers). 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Foop puiants: Betula, Rhus, Quercus, rose, pear, 
peach, apple, loquat, iris, cotton, Pinus. The larva is 
more or less a scavenger, feeding on dried seeds and 
mummied fruits, on dry rose buds, and in galls on roses, 
in acorns and old cotton bolls. Its favorite food in the 
neighborhood of Washington, D. C., seems to be the 
racemes of Rhus. It has much the habits of an E’phestia 
except that it is an outdoor insect and does not attack 
stored products to any extent. 

Distrisution: Unirep Srares: Maine; Vermont, 
Highgate (June); Massachusetts, Cohasset (July); Con- 
necticut, Kast River (Aug.); New York, East Aurora 
(Aug.), Ilion (Aug.), Newburgh (July), Rochester (Aug.); 
New Jersey, Dayton (July), Greenwood Lake (June), 
Pine Brook; Pennsylvania, Oak Station (Aug.), West 
Chester (July); Maryland, Hyattsville; District of 
Columbia, Washington (Mar., Apr., July); Virgina, 
Arlington County (Aug.), Blacksburg (June), Cape 
Henry (Aug.), Great Falls (Mar., Apr., May); North 
Carolina, Southern Pines; Florida, Camp Pinchot 
(Dec.), Hastings (Mar.), Lake Alfred (Aug.), Miami 
(July, Aug.), Monticello (Mar.), ‘Southern Florida” 
(June); Texas, Houston (July), Kountze (Apr.), Vic- 
toria (Mar., May); Ohio, New Richmond (Apr.); 
Iitinois, Oconee (Aug.); Jowa, Ames (Aug. Sept.). 
Canapva: Quebec, Chelsea (Apr.), Meach Lake; On- 
tario, Trenton (July). Distribution apparently limited 
to United States and Canada east of the Rockies. 


587. Moodna bisinuella Hampson 
Ficures 600, 1087 


Moodna bisinuella Hampson, in Ragonot, Monograph, pt. 2, 
p. 268, 1901. 

Forewing blackish fuscous with transverse lines 
obscure; on fresh specimens some reddish scaling in 
basal area and along the folds (visible only under 
magnification) ; antemedial line as in ostrinella but very 
faint; discal spots usually distinguishable and more or 
less confluent. Hind wing whitish with apical area 
somewhat smoky; the veins and a line along terminal 
margin considerably darker. Alar expanse, 17-22 mm. 

Male genitalia with inner costal projection from 
harpe curved outwardly; outer projection curved in- 
wardly, the two projections pointed toward each other. 
Uncus with sides strongly convex. Vinculum sharply 
angled at anterior end. Aedeagus considerably stouter 
than that of osirinella. Female genitalia with ductus 
bursae strongly sclerotized and striated towards genital 
opening. 

TYPE LOCALITY: Orizaba, México (type in USNM; 
paratypes in BM). 

Foop pLtant: Corn. 

Distrisution: México: Matamoros (July, Aug.), 
Orizaba, Tehuacén (Sept., Oct.). Unitmsp Sratzs: 
Texas, Crystal Springs (Oct.), Weslaco (June). 

Larvae of bisinuella have been frequently intercepted 
in ears of green corn from México at border ports by 
the Division of Foreign Plant Quarantine of the 
U. S. Bureau of Entomology and Plant Quarantine. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


The species has apparently invaded the United States 
from México and has become established in a few Texas 
localities. A larva indistinguishable from bisinuella 
has also been intercepted in green corn from British 
Honduras but no adults from that locality have been 
reared to verify the identification. 

Hampson’s description of the male antenna is mis- 
leading. He states that it has ‘deux sinus profonds 4 
la base de la tige.’’ The shaft is slightly swollen from 
the second to the seventh segments and beyond the 
seventh segment has a single short shallow sinus. His 
male type and several reared males are before me and 
each of them shows an antennal sinus like that of 
ostrinella. 


173. Genus Vitula Ragonot 


Vitula Ragonot, N. Amer. Phycitidae, p. 14, 1887; Monograph, 
pt. 2, p. 81, 1901.—Hulst, Phycitidae of N. Amer., p. 178, 
1890.—Forbes, Cornell Mem. 68, p. 631, 1923. (Type of 
genus: Vitula dentosella Ragonot). 

Eccopsia Hulst, U. 8. Nat. Mus. Bull. 52, p. 430, 1903.—Dyar, 


Proc. Ent. Soc. Washington, vol. 6, p. 158, 1904. (Type of 
genus: Vitula serratilineella Ragonot.) 
Tongue well developed. Antenna simple. Labial 


palpus upturned in male, more oblique in female, 
reaching a little above vertex; subcylindrical, somewhat 
rough scaled; third segment slightly shorter than 
second. Maxillary palpus filiform, short. Forewing 
smooth; 10 veins; vein 2 from before, but rather near 
lower outer angle of cell; 3 from the angle; 4 and 5 
stalked for half their lengths, the stalk separated from 
3 at base; 6 from below upper angle of cell, straight; 
8 and 9 united (a vestige of 9 present occasionally on 
one side or another of individual specimens); 10 from 
the cell, approximate at base to 8; male with a strong 
costal fold enclosing hair tuft. Hind wing with vein 2 
from well before lower outer angle of cell: 3 and 5 from 
the angle, approximate at base; 7 and 8 anastomosed 
for all or most of their lengths beyond cell (if present, 
8 a mere vestige at costa); cell about one-third the 
length of the wing (with veins 2, 3, and 5 appreciably 
long); discocellular vein curved. Eighth abdominal 
segment of male with compound dorsal tufts. 

Male genitalia with apical process of gnathos a short, 
bluntly pointed hook. Uncus broad; terminal margin 
broadly rounded. ‘Transtilla complete, an angulate 
bridge. Harpe with apex rounded; costa slightly 
humped near middle but without projections. Anellus 
a narrow, semicircular, sclerotized band with very short 
lateral lobes. Aedeagus long, straight, not tapering; 
penis bearing a single elongate, thin, weakly sclerotized 
plate, otherwise unarmed. Vinculum stout, slightly 
longer than broad, not appreciably tapering; terminal 
margin broad. 

Female genitalia with signum a small cuplike dise or 
a cluster of two or more such discs; bursa copulatrix 
small, more or less scobinate, the scobinations (except 
in laura) prominent asa partial girdlenearsignum;ductus 
bursae appreciably longer than bursa, strongly sclero- 
tized for a short distance from genital opening and with 


285 


a projecting, angulate, sclerotized plate behind genital 
opening; ductus seminalis from bursa close to signum; 
ventral membrane between eighth-segment collar and 
ovipositor not extruded. 

As here defined Vitula is distinguished from Moodna 
by its simple male antenna, complete transtilla, simple 
harpe (without costal projection) and the approximate, 
rather than connate, condition of veins 3 and 5 of hind 
wing. On the strength of the synonymy of dentosella 
and edmandsw Hulst cites edmandsii as type of the 
genus and is followed in this by Ragonot in his Mono- 
graph. This is not admissible nomenclatorially, as only 
dentosella was included under the original description 
of the genus and was definitely named at that time as 
type by Ragonot. 


588. Vitula edmandsae (Packard) 
Figures 115, 605 


Nephopteryx edmandsii Packard, Proc. Essex Inst., vol. 4, p. 120, 
1864; Guide to the study of insects, p. 331, 1869. 

Vitula dentosella Ragonot, N. Amer. Phycitidae, p. 14, 1887. 

Vitula edmandsii (Packard) Hulst, Ent. Amer., vol. 5, p. 156, 
1889; Phycitidae of N. Amer., p. 178, 1890.—Ragonot, 
Monograph, p. 2, p. 82, 1901.—Forbes, Cornell Mem. 68, 
p. 631, 1923.—Frison, Ann. Ent. Soc. Amer. vol. 19, p. 226, 
1926.—MceDunnough, Check list, No. 6323, 1939.—Corbet 
and Tams, Proc. Zool. Soc. London, vol. 113 (B), p. 64, 1943. 

Forewing gray more or less dusted with blackish 
fuscous, especially in the median area (between the 
transverse lines) ; on some specimens a reddish ocherous 
shade along lower fold and inner margin; pale ante- 
medial line obscure, defined chiefly by its contrasted, 
blackish outer border, the latter from costa near middle, 
outwardly angled at cell and usually with a slight notch 
at lower fold; subterminal line parallel to and well in 
from terminal margin, rather deeply notched at vein 6, 
and slightly so at lower fold, bordered inwardly by a 
thin black line; discal dots black, rarely separated, 
usually fused into a line along discocellular vein. Hind 
wing pale smoky fuscous, veins darker, a narrow dark 
line along terminal margin. Alar expanse, 15-22 mm. 

Genitalia as given for the genus. The female bursa 
shows more or less minute scaling near to and at junc- 
tion of bursa and ductus bursae. 

TyPE LocauitiEs: Bridport, Vt. (edmandsae, ia 
MCZ); North Carolina (dentosella, in Paris Mus.). 

Foop: Honeycomb of bees (larvae feeding on wax, 
pollen, and comb). 

DistriputTion: Unirep Sratrs: Vermont, Bridport; 
Massachusetts, Boston (May), Framingham (Sept.); 
Connecticut, East River (July); New Jersey, New Lisbon 
(Sept.); Pennsylvania, Oak Station (Sept.), New 
Brighton (June, Aug., Sept., Oct.) ; District of Columbia, 
Washington (June, July, Aug.); Maryland, Plummers 
Isl. (June, July, Aug.) ; North Carolina, Black Mountain 
(June), Tryon (June, Aug., Sept.) ; Kentucky, Lexington 
(Mar.); Missouri, St. Louis (June); Jilinois, Lacon 
(Aug.), Oconee (July); Arkansas, Washington County 
(July); Florida, Archer (Mar.). Canapa: Ontario, 
Trenton (Aug., Sept.) ; Quebec, St. Hilaire (June, Sept.). 


286 


Occasionally examples of edmandsae and its variety 
serratilineella show a vestige of vein 9 on forewing; but 
in normal specimens the fusion of 8 and 9 beyond the 
cell is complete, and the species obviously belongs in the 
group with vein 9 absent. Large pale examples are 
quite similar in habitus to Anagasta kuhniella and have 
been confused with that species in some collections. 
The mistake is very easy to make if one does not exam- 
ine the venation of all specimens before him. In his 
original description Packard stated that he was naming 
the species after Miss A. M. Edmands of Cambridge. I 
am therefore emending his name to give it the feminine 
ending required by the International Code. 


589. Vitula edmandsae serratilineella Ragonot, new status 


Figure 1088 


Vitula serratilineella Ragonot, N. Amer. Phycitidae, p. 15, 
1887.—Hulst, Phycitidae of N. Amer., p. 179, 1890.— 
Hampson, in Ragonot, Monograph, pt. 2, p. 83, 1901.— 
Dyar, Proc. Ent. Soc. Washington, vol. 5, p. 104, 1903 (de- 
scribes egg and larva); Proc. Ent. Soc. Washington, vol. 6, 
p. 158, 1904.—Hamlin and Reed, Journ. Econ. Ent., vol. 20, 
p. 840, 1927.—EHssig, Insects of western North America, 
p. 710, 1929.—Simmons, Reed, and McGregor, U. 8S. Dep. 
Agr. cire. 157, p. 38, 1931—McDunnough, Check list, No. 
6324, 1939. 


Eccopisa serratilineella (Ragonot), Monograph, pt. 2, p. 33, 
1901.—Hampson, in Ragonot, Monograph, pt. 2, p. 560, 
1901. 


Eccopsia serratilineella (Ragonot) Hulst, U. S. Nat. Mus. Bull. 
52, p. 430, 1903. 

Not structurally different from typical edmandsae and 
superficially distinguishable only by its somewhat paler 
(more whitish) hind wings. J am keeping the name to 
designate a western race which is of some importance 
as a minor pest of dried stored fruits in California; but 
it is probably not entitled even to this distinction. It 
also attacks the honeycombs of bees in the West, but, 
as in the Kastern States, does not seem to do any serious 
damage to thriving bee colonies. In the literature of 
economic entomology it is known as the “dried fruit 
moth.” Alar expanse, 14-25 mm. 

_Typr tocauity: North America (probably Southern 
California; type in Paris Mus.). 

Foop: Honeycombs of bees, dried fruits (apples, figs, 
raisins, prunes). 

Distrisution: Unitep States: California, Alameda 
County (Apr.), Claremont, El Segundo (sand dunes, 
Mar.), Fresno (May, June, July, Dec.), Humboldt, Los 
Angeles (Apr.), Mount View (Apr.), San Diego (Apr., 
June, July), Santa Clara County (Apr.), Santa Cruz 
County (Feb., June); Oregon (no definite locality); 
Washington, Pullman (Feb., Mar., May, July), Wen- 
atchee (Aug., Sept.); Wyoming, Cody (July); Nevada, 
Verdi (June); Utah, Stockton (Sept.); Colorado, Fort 
Collins; New Mexico, Pecos (June), Raton (Oct.); Ari- 
zona, Baboquivari Mts. (July). Camnapa: British Co- 
lumbia, Kaslo (Aug., Sept.), Victoria (June, July), 
Wellington (July). 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


590. Vitula lugubrella (Ragonot), new combination 
Fiaure 607 

Hornigia lugubrella Ragonot, N. Amer. Phycitidae, p. 17, 1887. 

Manhatia lugubrella (Ragonot) Hulst, Phycitidae of N. Amer., 
p. 197, 1890. 

Moodna lugubrella (Ragonot) Hampson, 7n Ragonot, Monograph, 
pt. 2, p. 270, 1901.—MecDunnough, Check list, No. 6395, 
1939. 

I have seen no California specimens that match 
Ragonot’s description or figure. Evidently the fore- 
wing shows some appreciable dusting of reddish scales 
and has the antemedial line straight and approximately 
vertical (more or less oblique from costa to inner mar- 
gin). The genitalia of one of Ragonot’s male paratypes 
(from an abdomen loaned by the Paris Museum) are 
similar to those of edmandsae except that the vinculum 
is longer (like that of pinei) and the harpe tapering from 
middle to a more narrowly rounded apex, as shown in 
figure 607. 


TypxE Locatity: California (type in Paris Mus.). 
Foop PLant: Unknown. 


591. Vitula pinei, new species 
Figures 606, 1090 


Forewing white finely dusted with fuscous, giving the 
wing a pale gray ground color to the naked eye; trans- 
verse lines white bordered inwardly and outwardly by 
sharply contrasted blackish lines; antemedial line 
slightly angulate; subterminal line bent. inward slightly 
for ashort distance from costa, shortly and sharply angled 
out at middle and slightly notched at lower fold; discal 
dots fused into a black line along discocellular vein and 
extending (in fresh specimens) to the black inner border 
of subterminal line; terminal dots fused into a black line 
along termen; hair tuft enclosed by costal fold, white. 
Hind wing whitish, semihyaline with a smoky line along 
termen and some smoky shading on the veins. Alar 
expanse, 19-21 mm. 


Genitalia differing in slight details from those of 
edmandsae; vinculum longer, its terminal margin more 
evenly rounded; hump of costa of harpe more angu- 
late; aedeagus stouter; female genitalia considerably 
longer and sclerotization of ductus bursae somewhat 
more extended; comparative characters only. 


Tyrer nLocauity: Hureka, Utah (type m USNM, 
61388). 


Foop pLant: Pine cones. 


Described from male type and one male paratype 
from the type locality, July 14 and 17, 1911, collected 
by Tom Spalding; and one female paratype from Baker, 
Nev., reared by T. O. Thacher, May 28, 1940, from larva 
feeding in cone of Pinus monophylla. The larva was 
presumably feeding on the seeds or bracts. 

The species is easily distinguished from edmandsae by 
the sharply contrasted black double borders of the ante- 
medial and subterminal lines. Superficially it bears a 
rather striking resemblance to Laetilia zamacrella Dyar. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


592. Vitula inanimella (Dyar), new combination 
Fieure 1089 


Moodna inanimella Dyar, Proc. U. S. Nat. Mus., vol. 54, p. 372, 


Euzophera ticitoa Dyar, Ins. Insc. Menstr., vol. 7, p. 56, 1919 
(new synonymy). 

A suffused gray-brown species with transverse lines 
faintly indicated by their very slightly darker borders; 
similar in maculation to edmandsae, but more uniformly 
colored. The genitalia are also similar, those of the 
males differing only in insignificant details, the differ- 
ences no greater than those between individuals of 
edmandsae. Female genitalia have the bursae smooth 
except for the band of scobinations near signum and the 
signa themselves smaller. The type of ticitoa (fig. 
1089a) has one signum, the female paratype of inani- 
mella from Orizaba, México (fig. 1089) has two signa, a 
difference of not specific significance in this genus or the 
allied Ephestia groups. The type of ticitoa shows a 
vestige of vein 9 on one forewing which probably ac- 
counts for Dyar’s placement of it in Huzophera. I am 
unable to find a valid character for separating Dyar’s 
supposed species and am therefore sinking ficitoa into 
the synonymy of inanimella. Alar expanse, 20-21 mm. 

Tyre Locauities: Zacualpin, México (inanimella, in 
USNM); Volcaén Santa Maria, Guatemala (ticitoa, in 
USNM). 

Foop puants: Unknown. 

Distrisution: México: Orizaba, Zacualpin (May). 
Guatema.a: Volcin Santa Maria (May). 

Represented in the National Collection only by the 
original type series of two males and three females. 


593. Vitula laura (Dyar), new combination 
Fieure 1091 


Euzophera laura Dyar, Ins. Insc. Mensir., vol. 7, p. 56, 1919. 


Forewing dark purplish gray, costal area white; 
antemedial line indicated by an oblique blackish band 
extending across the white area and obsolete below; 
subterminal line white, thin, parallel with and rather 
close to costa, evenly curved, bordered before and 
beyond by narrow bands of the dark ground color; some 
faint white dusting bordering termen; discal dots dis- 
tinct, separate, black; a row of faint blackish dots along 
terminal margin. Hind wing a glossy smoky brown; 
veins and terminal margin darker. Alar expanse, 20 
mm. 


Female genitalia with bursa copulatrix finely, evenly, 
and sparsely scobinate; signum a cluster of a half-dozen 
small discs; ductus bursae with a sclerotized collar for- 
ward of the short sclerotized area from genital opening. 

TypE Locatity: Cayuga, Guatemala (July; type in 
USNM). 

Foop prant: Unknown. 

Known only from the unique female type. It is 
obviously not a Euzophera, and in habitus and female 
genitalia is not too good a Vitula, in which genus it is 
placed only provisionally. A male will be necessary 
for definite placement. Vein 9 of forewing is absent 


287 


and the venation otherwise and the female genitalia 
indicate that it belongs somewhere in the Moodna-— 
Vitula complex. 


174. Genus Manhatta Hulst 


Hornigia Ragonot, N. Amer. Phycitidae, p. 16, 1887. (Type of 
genus: Ephestia biviella Zeller; Europe; figs. 113, 601, 1092.) 

Manhatta Hulst, Phycitidae of N. Amer., p. 196, 1890 (mew 
name for Hornigia Ragonot 1887, preoccupied by Hornigia 
Ragonot 1885 in Galleriidae). 

Characters of Vitula except veins 3 and 5 of hind 
wing connate. 

Hampson (in Ragonot, Monograph, pt. 2, p. 267, 
1901) made Manhatta a synonym of Moodna, apparently 
on the connate condition of veins 3 and 5 of the hind 
wing of biviella. This synonymy cannot stand if we 
are to maintain any generic separation between Moodna 
and Vitula; for Manhatta has only the hind wing vena- 
tion and somewhat longer hind wing cell of Moodna. In 
other characters (simple male antenna and complete 
transtilla) it agrees with Vitula, thus occupying an inter- 
mediate position between the two genera, but appar- 
ently more closely related to Vitula than to Moodna. 
In addition to its type (biviella), the only European 
representative of the Moodna—Viiula complex, it contains 
two North American species. 


594. Manhatta setonella (McDunnough), new combination 
Figures 602, 1093 


Moodna setonella McDunnough, Canadian Ent., vol. 59, p. 270, 
1927; Check list, No. 6398, 1939. 

Forewing whitish gray, more or less dusted with 
blackish scales on lower half of wing, giving that area a 
somewhat smoky tint; black discal spots and borders of 
the transverse lines strongly contrasted, the transverse 
lines themselves not distinguishable from the ground 
color of wing and indicated only by their black borders; 
antemedial line bordered outwardly only, the black 
border normally straight and oblique, but in some 
specimens slightly angled; subterminal line bulging 
slightly at middle, bordered inwardly by a narrow black 
line and outwardly by a black dash at costa, continu- 
ing as a paler shade to inner margin; discal dots distinct, 
separate; a few obscure blackish dots on terminal mar- 
gin, beginning below apex and ending above tornus. 
Hind wings semihyaline, whitish with veins and terminal 
margin pale smoky fuscous. Alar expanse, 14-16 mm. 


Male genitalia with costa of harpe produced at apex 
into a short free spur. Female genitalia with ductus 
bursae sclerotized for half its length from genital opening, 
the sclerotized portion constricted at middle; a small 
sclerotized patch at junction of ductus and bursa; spin- 
ing of bursa as in typical Vitula. 

Typr LocaLity: Seton Lake, British Columbia (June; 
type in Canadian Nat. Coll.). 

Foop puant: Unknown. 

In addition to female paratypes from the type locality 
there are in the U.S. National Collection two males from 
Eureka and Provo, Utah (June). The species is easily 


288 


recognized byits genitalia; the apically produced costa of 
harpe and the patch in the neck of the bursa at once 
distinguishing it from anything else in the Moodna— 
Vitula group. Superficially the moths resemble those 
of Vitula pine but are smaller and lack the dark border 
on inner side of the antemedial line. 


595. Manhatta broweri, new species 
Fieure 1094 


Forewing pale brownish fuscous with a white-powdered 
area about the discal dots and extending to costa, and 
some white dusting at tornus; transverse lines white, 
narrow; at base below fold and extending beyond ante- 
medial line along fold a faint reddish ocherous shade; 
dark ground color concentrated and intensified as rather 
diffused broad bands outwardly bordering the anteme- 
dial and inwardly bordering the subterminal lines; ante- 
medial line slightly angulate; subterminal line irregu- 
larly dentate, parallel with termen; discal dots more or 
less confluent, brown. Hind wing pale fuscous, veins 
and terminal margin slightly darker. Alar expanse, 14— 
16 mm. 

Genitalia, (male and female) differing very little from 
those of Vitula edmandsae and exhibiting no distinguish- 
ing specific characters. 

TYPE LocALITY: Bar Harbor, Maine (type in USNM, 
61389). 

Foop prant: Unknown. 

Described from male type and one male and five female 
paratypes from the type locality, July 25-30, 1933, and 
one male paratype from Mount Desert Isl., Maine, July 
17, 1934, all collected by Dr. E. A. Brower for whom the 
species isnamed. Paratypes deposited in Dr. Brower’s 
collection. 

The species is easily identified by its color and macu- 
lation. Its male genitalia (of the edmandsae type) dis- 
tinguish it from the other two species of Manhatia and 
its hind wing venation from any species of Vitula. It 
is the only known species of Manhatta in the eastern 
United States. 


175. Verina, new genus 


TypE or Genus: Moodna supplicellaiDyar 

Tongue well developed. Antenna pubescent (in male 
the cilia less that the width of shaft in length); shaft of 
male with a few rough scales at base, above, and just 
beyond, a slight, very shallow sinus. Labial palpus 
oblique, slender, reaching above vertex; third segment as 
long assecond. Maxillary palpus filiform,short. Fore- 
wing smooth; 10 veins; vein 2 from before, but near, 
lower outer angle of cell; 3 from the angle; 4 and 5 long 
stalked; 6 from below upper angle of cell, straight; 8 
and 9 united; 10 from the cell approximate to 8 at base; 
male with a strong costal fold enclosing hair tuft. Hind 
wing with vein 2 from well before lower outer angle of 
cell; 3 and 5 from the angle, closely approximate at base; 
7 and 8 anastomosed for all or most of their lengths be- 
yond cell; cell very short (about one-fourth the length 
of wing), a trifle longer in female than in male; disco- 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


cellular vein slightly curved (nearly straight in male). 
Kighth abdominal segment of male with compound dor- 
sal tufts. 

Male genitalia with apical process of gnathos forked, 
U-shaped. Uncus broad, terminal margin broadly 
rounded. Transtilla mcomplete. Harpe with a short 
blunt digitate projection from near middle of costa. 
Anellus a U-shaped sclerotized band. Aedeagus long, 
not appreciably tapering; penis bearing a couple of 
elongate, thin, weakly sclerotized plates, otherwise un- 
armed. Vinculum elongate-angulate, tapering. 

Female genitalia with ductus bursae sclerotized for 
most of its length from genital opening; sclerotized por- 
tion slightly flattened; bursa copulatrix sparsely and 
finely scobinate; an extruded lobe from membranous 
area between eighth-segment collar and ovipositor (as 
in Moodna). 

The genus is distinguished from the other genera in 
group C by the following’ combination of characters: 
Male antenna with slight sinus in shaft; eighth abdom- 
inal segment of male with tuft; veins 3 and 5 of hind 
wing approximate at base, cell very short and discocel- 
lular vein very slightly curved; gnathos terminating in a 
forked process; harpe with digitate projection from costa; 
transtilla incomplete. 

It contains one tropical American species. 


596. Verina supplicella (Dyar), new combination 
Fieures 116, 603, 1097 


Moodna supplicella Dyar, Proc. U. 8S. Nat. Mus., vol. 47, p. 342, 
1914. 


Forewing dark vinous, dusted with black especially 
in area between the transverse lines; basal and tornal 
areas slightly paler; transverse lines white, slender, 
powdery and broken, the antemedial line far out and 
oblique, the subterminal parallel with and close to 
termen; discal and terminal dots not distinguishable. 
Hind wing translucent; whitish to pale fuscous; the 
veins and terminal margin darker. Head and thorax 
ocherous, more or less shaded with reddish or blackish 
scaling. Labial palpus ocherous with reddish scaling 
on terminal segment. Alar expanse, 13-16 mm. 

Male genitalia with terminal margin of cucullus of 
harpe oblique and slightly concave, its lower angle 
produced, vinculum evenly tapering to abruptly pointed 
extremity; arms of forked process of gnathos approx- 
imate at their apices. Female genitalia with sclerotized 
portion of ductus bursae strongly striated at genital 
opening, finely granulate otherwise. 

Typr Locatiry: Rio Trinidad, Panamé (type in 
USNM). 

Foop puant: Dried leaves. 

Distrisution: Mexico: No specific locality (Aug., 
reared from leaf trash in banana cargo). GUATEMALA: 
No specific locality (Apr., reared from dried leaves 
intercepted at quarantine), Guatemala City (Mar.). 
Panami: Cabima (May), Porto Bello (Apr., May, 
Oct.), Rio Trinidad (June). Braziu: Santa Catarina 
(July). 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


This species has been intercepted a number of times 
at our quarantine ports in banana trash. The larva is 
apparently a feeder on dry vegetable refuse. The moth 
is easily identified by its male genitalia and (in fresh 
specimens) by the broken powdery transverse lines and 
wine color of its forewings. 


176. Vagobanta, new genus 


Type oF GENus: Cryptoblabes divergens Butler. 

Characters of Verina except: Antenna of male simple; 
eighth abdominal segment of male simple; forewing with 
vein 2 shortly separated from, approximate to, or con- 
nate with 3 (more separated in females than in males); 
hind wing with veins 3 and 5 shortly stalked; cell long 
(one-half or a trifle over one-half the length of wing); 
discocellular vein decidedly curved. 

Contains one tropical American species. 


597. Vagobanta divergens (Butler), new combination 
Fiaures 604, 1100 


Cryptoblabes divergens Butler, Trans. Ent. Soc. London, p. 60, 
1883. 


Moodna divergens (Butler) Hampson, in Ragonot, Monograph, 
pt. 2, p. 269, 1901. 

Forewing powder gray, the basal area distinctly paler; 
in fresh specimens an obscure shading of reddish scales 
on middle of lower fold; antemedial line distinct, de- 
cidedly oblique, white, bordered outwardly by a black- 
ish band, nearly straight but in some specimens with a 
notch at lower fold; subterminal line obscure, sinuate, 
very faintly bordered by dark line on inner side. Hind 
wing translucent, white, with a pale brown line along 
termen and the veins slightly darkened. Alar expanse, 
21-25 mm. 

Male genitalia with arms of U-shaped apical projec- 
tion of gnathos very short and widely spaced. Ele- 
ments of divided transtilla long and stout. Harpe with 
digitate projection from about middle of costa; outer 
margin of cucullus rounded and turned up slightly at 
apex. Anellus V-shaped, with stout base. Aedeagus 
stout. Vinculum stout, but slightly tapering; terminal 
end broadly and abruptly angulate, reinforced by a 
sclerotized, pocketlike fold. 

Female genitalia with bursa copulatrix large, finely 
and sparsely scobinate; signum weak; ductus bursae 
sclerotized for half its length, the sclerotized portion 
flattened and bent. 

TypxE LocaLity: Coquimbo, Chile (type in BM). 

Foop piant: Puya alpestris. 

Distrisution: Cute: Angol, Coquimbo. 

A reared series of eight specimens from Angol, Chile, 
is in the National Collection, received from D. S. 
Bullock but undated. 


177. Moodnella, new genus 


Typr or Genus: Moodnella paula, new species. 
Tongue well developed. Antenna with slight sinus 
in shaft of male near base. Labial palpus oblique, 
slender, reaching a trifle above vertex, third segment 
800329—56——20 


289 


shorter than second. Maxillary palpus filiform, rather 
long (as long as third segment of labial palpus and about 
twice the length of the maxillary palpi of the other 
genera in the Moodna-Vitula group). Wing venation 
asin Vitula. Forewing with strong costal fold enclosing 
scale tuft. Hind wing with cell one-third the length of 
wing; discocellular vein curved. Eighth abdominal 
segment of male simple. 

Male genitalia as in Vitula. 

Female genitalia with ductus bursae very weakly 
sclerotized for a very short distance from genital open- 


Differs from Vitula in having a sinus in the shaft of 
male antenna, no tufts on eighth abdominal segment of 
the male; from Moodna in its complete transtilla, simple 
harpe, simple eighth abdominal segment, approximate 
condition of veins 3 and 5 of hind wing; from Verina 
in its complete transtilla, small hooked, apical process 
of gnathos, simple harpe, simple eighth abdominal seg- 
ment, longer cell and more distinctly curved discocellu- 
lar vein of hind wing; and from all three of these genera 
by its longer maxillary palpi. 

Contains one tropical American species. 


598. Moodnella paula, new species 
Ficures 608, 1096 


Forewing gray suffused with rufous and shaded with 
blackish, the blackish shade extending in a narrow band 
along costa and diffused irregularly in the median area; 
antemedial white line far out on wing, at or a trifle 
beyond middle, distinct on lower half, fading out 
towards costa, bordered outwardly by an irregularly 
diffused black, band; subterminal line narrow, parallel 
with termen, nearly straight, slightly oblique from costa 
to vein 6, shortly out-angled just below, thence straight 
to inner margin, white, bordered inwardly by a narrow 
blackish band; the area between the transverse lines 
greatly restricted. Hind wings white to smoky fuscous; 
the veins faintly darkened and a dark line along termen. 
Head and thorax reddish ocherous with a scattered 
dusting of blackish scales. Alar expanse, 14-15 mm. 

Male genitalia with uncus narrowly rounded at apex; 
vinculum long, stout, scarcely tapering, terminal margin 
broad but very slightly convex (nearly straight). 

TypE Ltocauity: Guatemala City, Guatemala (type 
in USNM, 61390). 

Foop prant: Unknown. 

Described from male type and one male paratype 
from the type locality collected by C. N. Ainslie, Mar. 
1932; one female paratype from Santa Catarina, Brazil, 
collected by Fritz Hoffmann, July 9, 1935; one female 
paratype from Tigre, Argentina, Aug. 1939, from the 
collection of Fernando Bourquin; and one female para- 
type from Vicosa, Minas Gerais, Brazil, E. J. Hamble- 
ton, collector, ‘‘12—-1-34,” from the Cornell Collection. 

A pretty little species easily recognized by its reddish 
fuscous color, the narrow interspace between the trans- 
verse lines of forewing and the distinct whiteness of 
these lines on their lower halves. 


290 
* Genera 178 and 179: Volatica and Vezina 


[Venational division A. ‘Forewing with 11 veins; 8 and 9 long 
stalked (9 rather weak). Hind wing with veins 3 and 5 approxi- 
mate and from lower angle of cell. Labial palpi porrect or 
oblique. Maxillary palpi minute. Transtilla complete and de- 
veloped as an angulate bridge, or incomplete. Ductus bursae 
with projecting sclerotized shield behind genital opening.] 


178. Volatica, new genus 


TyPE oF GENUS: Zophodia pachytaeniella Ragonot. 

Tongue well developed. Antenna pubescent, simple. 
Labial palpus porrect (second segment oblique, third 
deflected forward), extending twice the length of head 
beyond it, broadly scaled, third segment less than half 
the length of second. Maxillary palpus minute, fili- 
form. Forewing smooth; 11 veins; vein 2 from before, 
but near, lower outer angle of cell; 3 from the angle; 4 
and 5 stalked (for at least half their lengths), the stalk 
separated from 3 at base; 6 from below upper angle of 
cell, straight; 8 and 9 long stalked (free element of 9 
sometimes weak, but always present); 10 from the cell, 
separated from the stalk of 8-9 at base; male without 
costal fold. Hind wing with vein 2 from well before 
lower outer angle of cell; 3 and 5 from the angle, approxi- 
mate at base; 7 and 8 anastomosed beyond cell for at 
least half their lengths; cell about half the length of 
wing; discocellular vein curved. Highth abdominal 
segment of male simple. 

Male genitalia of the Vitula type but larger and more 
robust. 

Female genitalia with or without signa; bursa copu- 
latrix more or less finely scobinate; ductus bursae shorter 
Gn pachytaeniella) or very little longer than bursa, 
(trinitatis), sclerotized for over half its length, the 
sclerotized portion flattened except just before genital 
opening and with a more or less rounded, projecting, 
sclerotized shield behind genital opening; ductus 
seminalis from bursa near signum. 

The genitalia, male and female, show the close rela- 
tionship of this genus to the Vitula group despite the 
forewing venation which is definitely that of venational 
division A. Vein 9 while sometimes weak is always 
present. In Vitula on the other hand vein 9 is nor- 
mally absent, being present and weak only in occasional 
specimens. Volatica is distinguished from all the other 
genera of the Moodna—Vitula complex by its porrect, 
broadly scaled labial palpi. 


599. Volatica pachytaeniella (Ragonot), new combination 
Figures 610, 1098 

Zophodia pachytaeniella Ragonot, Nouv. Gen., p. 31, 1888.— 

Hampson, zn Ragonot, Monograph, pt. 2, p. 23, 1901. 
Forewing white lightly sprinkled with brownish 
fuscous scales giving the costal half of the wing a soiled 
ashy white appearance; lower half of wing smeared with 
a pale drab shade; on fresh specimens a pink streak 
along lower margin at base; antemedial line angulate, 
indicated chiefly by a rather broad blackish brown outer 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


border; subterminal line more or less irregularly dentate, 

bordered inwardly by a narrow blackish fuscous ‘line; 

discal spot at lower outer angle of cell distinct, upper’ 
discal spot occasionally distinct but often obscure or 

absent. Hind wing white; a broad fuscous shade along 

costa; some darkening of the veins and a distinct dark 

line along termen, broadening in most specimens. Alar 

expanse, 26-33 mm. 

Male genitalia with vinculum slightly constricted 
before terminal margin; apical end of harpe somewhat 
spatulate. Female genitalia with bursa copulatrix 
elongate, much longer than ductus. bursae; signum 
present, consisting of a short band of partially fused 
discs; ductus bursaesclerotized for its entire length. 

TypE LocaAuity: Rio de Janeiro, Brazil (type in Paris 
Mus.). 


Foop PLANT: Unknown. 


Distrisution: Braziu: Parand, Castro, Ypiranga; 
Rio de Janeiro, Petrépolis; Santa Catarina (no dates on 
any specimens before me). 


A large, distinct species so far only known from 
Brazil, easily identified by maculation, color, and 
genitalic structure. 


600. Volatica trinitatis, new species 
Fieures 609, 1099 


Somewhat narrower winged than pachytaeniella, 
similarly colored, but with whitish area of forewing 
restricted to a narrower border along costa; transverse 
lines obscured, only the antemedial line indicated in 
male by a partial dark outer border, very faint in the 
males before me, absent in the female; the latter shows 
some dark scaling on the veins and has fuscous hind 
wings; hind wing of male white with little or no dark 
shading along costa or termen. Alar expanse, 24-26 
mm. 

Male genitalia with apical process of gnathos longer 
than in pachyiaeniella (over twice as long as broad); 
terminal margin of harpe evenly rounded; vinculum 
not appreciably constricted towards terminal margin. 
Female genitalia with bursa copulatrix somewhat 
shorter than ductus bursae, without signum; ductus 
bursae sclerotized for not more than two-thirds of its 
length, bent slightly near membranous part of the 
ductus. ‘There are also differences in the shape of the 
dorsal projecting shield at genital opening between 
trimiatis and pachytaeniella, but these differences may 
not be significant. 

Typr Locatity: Fyzabad, Trinidad (type in Cornell 
Univ. Coll.). 

Foop prant: Unknown. 

Described from male type and one male and one 
female paratype from the type locality. (Feb. 17, 19, 
21, 1928) from the Cornell Collection, and one male 
paratype (USNM 61391) from Trinidad, without date 
or more exact locality, collected by A. Busck. 

The species is very close to but apparently distinct 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 291 


from pachytaeniella, easily distinguished by its genitalia, 
the more extended drab suffusion and less distinct 
transverse lines on forewing. 


179. Vezina, new genus 


TypE oF Genus: Vezina parasitaria, new species. 

Tongue well developed. Antenna pubescent, simple. 
Labial palpus oblique; reaching as high as vertex; 
rather broadly and roughly scaled; third segment 
shorter than second. Maxillary palpus short, filiform. 
Forewing smooth; 11 veins; vein 2 from before but 
rather near lower outer angle of cell; 3 from the angle; 
4 and 5 stalked for half their lengths, separated from 3 
at base; 6 from below upper angle of cell, straight; 8 
and 9 long stalked; 10 from the cell, separated from 8 
at base; male with a strong costal fold enclosing hair 
tuft. Hind wing with vein 2 from well before lower 
outer angle of cell; 3 and 5 from the angle, approximate 
at base; 7 and 8 anastomosed for most of their lengths 
beyond cell; cell about one-third the length of the wing 
in male, one-half in female; discocellular vein curved. 
Eighth abdominal segment of male with two pairs of 
dorsolateral hair tufts. 

Male genitalia with apical process of gnathos a short, 
bluntly pointed hook. Uncus broad; terminal margin 
broadly rounded. ‘Transtilla incomplete; its elements 
weakly sclerotized. Harpe with a short angulate pro- 
jection from costa near middle and with apex of costa 
produced into a free hook at apex (as in Anagasta 
kithniella). Anellus a stout, broad, semicircular band 
with broad, deep, V-shaped ventral incision. Aedeagus 
long, stout; penis armed with a thin, narrow, sclerotized 
band and a stout, thornlike, broadly based spine. 
Vinculum stout; tapering slightly; terminal margin 
moderately broad. 

Female genitalia with signum a small cuplike disc or 
discs; bursa minutely and sparsely granulate; ductus 
bursae granulate towards junction with bursa, with a 
strongly sclerotized collar surrounding a broad genital 
opening and with a broad, projecting, sclerotized shield 
behind genital opening; ductus seminalis from bursa 
close to signum. 

This genus, like Volatica, is close to and obviously 
related to Vitula despite its forewing venation. It is 
distinguished by the following combination of charac- 
ters: Vein 9 of forewing present; eighth abdominal 
segment of male with paired dorsal tufts; transtilla in- 
complete; penis armed with a stout, thornlike cornutus; 
anellus a broad, semicircular band with V-shaped ven- 
tral incision; ductus bursae with broad sclerotization 
surrounding broad genital opening. 


601. Vezina parasitaria, new species 


Figures 598, 1068 


Forewing dark gray (grayish brown on worn and 
faded specimens); transverse lines white and strongly 
contrasted from inner margin to cell, thence to costa 


pale but more obscure, narrowly bordered inwardly and 
outwardly by black lines; these most obvious from lower 
margin of cell to costa; antemedial line at middle of 
wing and nearly vertical; subterminal line oblique and 
well back from termen; the interspace between the lines 
short; discal spots at end of cell, confluent and forming 
a thin black line along discocellular vein; terminal dots 
obscure. Hind wing shining white, with a fuscous 
shade along costa, some fuscous shading at apex and a 
narrow dark line along termen. Alar expanse, 17-25 
mm. 

Male genitalia with vinculum but slightly longer 
than broad; terminal margin angulate. Female geni- 
talia with a short, thornlike pouch projecting from 
venter of ductus bursae near genital opening. 

Typr tocauity: José C. Paz, Province of Buenos 
Aires, Argentina (type in USNM, 61392; paratypes in 
Mus. Argentino Cienc. Nat., Buenos Aires, and BM). 

Foop: Larvae feeding in larval cases of Oiketicus 
kirbyi Guilding. 

Described from male type and one male and one 
female paratype from the type locality; one male para- 
type from Juan B. Gonnet, Province of Buenos Aires; 
five female paratypes from Tigre, Argentina; and one 
female paratype from southeast Brazil, F. D. Jones, 
1920, this last from the British Museum Collection. 
The Argentinian specimens were received from Dr. 
Everard E. Blanchard, Director and Sefor José A. 
Pastrana, Ingeniero, of the Instituto de Sanidad 
Vegetal of the Ministerio de Agricultura in Argentina. 
They had been reared by Sefior Pastrana and Dr. Pablo 
Kohler from small larvae in the larval cases of the 
psychid Otketicus kirbyi Guilding. Dr. Kébler identi- 
fied them as a new species to which he gave the manu- 
script name ‘‘Canarsia parasitaria” upon the assump- 
tion that the species was parasitic on the Ovketicus. 
With Dr. Koéhler’s permission I am adopting his manu- 
script specific name, but am in doubt as to the parasitic 
nature of the larva. According to Sefior Pastrana the 
actual feeding habits have not been noted, and I am 
inclined to believe (from its close affinities to the 
Moodna-Vitula complex) that parasitaria is a scavenger 
rather than a parasite or predator upon the living 
Otketicus larva or pupa. In a letter of Dec. 29, 1947, 
Dr. Blanchard states that, in the Province of José C. 
Paz, Sefior N. Jauch has observed larvae of parasitaria 
on the remains of bagworms and a few days later all 
traces of bagworms had disappeared, suggesting that 
the larva are feeders upon the bagworms at least to the 
extent of eating the larval and pupal exuviae or the 
remains of the dead females of Oiketicus. 

The species is a striking one easily recognized by its 
bright white transverse lines with narrow black borders 
contrasted against the otherwise dark gray ground color 
of the forewing. The female genitalia are different in 
the armature about genital opening from any other in 
the subfamily. 


292 
Genera 180-185: Caudellia to Plodia 


[Venational division E. Forewing with 9 veins; 10 from the 
cell; 9 absent; 4 absent; 2 and 3 from the cell. Hind wing with 
discocellular vein curved. Male genitalia with transtilla com- 
plete or its elements greatly enlarged. Female genitalia with 
ductus bursae sclerotized for a considerable part of its length 
and usually flattened.] 


180. Genus Caudellia Dyar 


Caudellia Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 116, 
1904.—Forbes, Cornell Mem. 68, p. 636, 1923. (Type of 
genus: Caudellia apyrella Dyar.) 


Tongue well developed. Antenna pubescent; shaft 
of male with a short, shallow sinus near base (as in 
Moodna). Labial palpus oblique, reaching to or slightly 
above vertex, somewhat flattened laterally; third seg- 
ment nearly as long as second (somewhat broadly scaled 
and deflected forward in apyrella and albovittella). Max- 
illary palpus minute, filiform. Forewing smooth; 9 
veins; vein 2 from before but rather near lower outer 
angle of cell; 3 from the angle; 4 absent; 5 more or less 
approximate to 3 at base; 6 from below upper angle of 
cell, straight; 8 and 9 united (9 absent); 10 from the 
cell, separated from 8 at base; male with costal fold 
enclosing a scale tuft. Hind wing with vein 2 from well 
before lower outer angle of cell; 3 and 5 connate, rarely 
very shortly stalked; 7 and 8 anastomosed for most or 
all of their lengths beyond cell (8 when present very 
short) ; cell one-half or slightly less than half the length 
of wing; discocellular vein curved. Wighth abdominal 
segment of male with compound dorsal tufts. 


Male genitalia with transtilla complete (declivella) or 
its elements broadened and often fused with arms of 
gnathos. Aedeagus straight, smooth; penis with cor- 
nutus developed as a weakly sclerotized, narrow, flat- 
tened band, otherwise unarmed. 


Female genitalia with ductus bursae sclerotized for a 
third or more of its length from genital opening; signa 
present as a cluster of small, bluntly rounded, projecting 
discs or spines more or less fused at their bases; ductus 
seminalis from bursa adjacent to signa (approximately 
at middle of bursa). 

This genus and the following genera with sclerotized 
ductus bursae form a compact, strictly American group 
closely related to and evidently derived from the 
Moodna-Vitula complex. The moths of Caudellia also 
resemble Moodna in color and maculation and male 
antennal structures. The species differ markedly from 
each other in male genitalia. On labial palpi they 
divide into two groups: 


Palpus somewhat broadly scaled and third segment deflected 
forward. 
Palpus slender and third segment not deflected. 


The deflection of the third segment in our two species 
from the Eastern United States is rather slight and does 
not give the palpus nearly so pronouncedly porrect an 
appearance as, for example, that of Plodia, and I do not 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


believe justifies any generic separation of the two 
species groups. 


Genus Caudellia, Species 602 and 603: C.. apyrella 
and C. albovittella 


[Labial palpus somewhat broadly scaled and third segment de- 
tected forward.] 
602. Caudellia apyrella Dyar 
Figures 122, 613 
Caudellia apyrella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 116, 1904.—Forbes, Cornell Mem. 68, p. 636, 1923.— 
McDunnough, Check list, No. 6376, 1939. 

Forewing vinous brown overlaid with blackish brown, 
the blackish shading most obvious at apex; a faint pale 
patch on outer third of costa; transverse lines and discal 
spots obscured, the antemedial line very faintly indi- 
cated, oblique, pale vinous brown. Hind wing whitish, 
faintly tinted with fuscous; veins not appreciably 
darkened. Alar expanse, 14-15 mm. 

Male genitalia with gnathos terminating in a stubby, 
angulate projection; harpe simple. 

Tyrer LocaLity: Plummers Island, Md. (type in 
USNM). 

Foop prant: Unknown. 

Known only from the type (co, July) and paratype 
(o@, June) from the type locality. 


603. Caudellia albovittella Dyar 
Ficure 1103 
Caudellia albovitiella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p- 116, 1904.—Forbes, Cornell Mem. 68, p. 636, 1923.— 
MeDunnough, Check list, No. 6377, 1939. 

Forewing vinous brown shaded with blackish brown, 
the blackish shading less diffused than in apyrella, con- 
centrated as a dark median streak at base, a dark blotch 
on outer side of antemedial line and a fainter shade at 
apex; antemedial line oblique, strongly contrasted, 
white, preceded by a short white dash on inner margin; 
subterminal line obscure, distinguishable on fresh speci- 
mens as a thin irregular pale line near termen; discal 
dots faint, blackish, separate; terminal dots confluent. 
Hind wing pale fuscous, veins very slightly darkened. 
Alar expanse, 13-21 mm. 

Male genitalia not distinguishable from those of 
apyrella. Female genitalia with ductus bursae scler- 
otized for slightly less than half its length from genital 
opening; signa a double line of 6 to 8 short discs, close 
together and fusing at their bases. 

Typ Ltocatity: Plummers Island, Md. 

Foop prant: Unknown. 

Distrisution: Maryland, Hyattsville (July), Plum- 
mers Isl. (June, July); Missouri, St. Louis (Aug.). 

Probably not specifically distinct from apyrella which 
may be only a suffused color form or food-plant race; but 
as nothing is known of the biology and no females cor- 
responding to the males of apyrella are available for 
genitalic comparison, the two will have to be kept as 
separate species for the time being. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Genus Caudellia, Species 604-607: C. nigrella to 
C. clara 


[Labial palpus slender and third segment not deflected.] 


604. Caudellia nigrella (Hulst), new combination 
Ficures 611, 1102 

Ephestia nigrella Hulst, Phycitidae of N. Amer., p. 200, 1890. 

Mescinia nigrella (Hulst) Ragonot, Monograph, pt. 2, p. 85, 1901. 

Moodna nigrella (Hulst) Barnes and McDunnough, Check list of 
the Lepidoptera of Boreal America, No. 5796, 1917.— 
McDunnough, Check list, No. 6397, 1939. 

Ephestia arizonella Walter, Proc, Ent. Soc. Washington, vol. 30, 
p. 141, 1928.—McDunnough, Check list, No. 6405, 1939. 
(New synonymy.) 

In color and markings resembles Moodna ostrinella. 
Forewing blackish fuscous; basal area more or less 
shaded with red or reddish ocherous, especially on lower 
half, this reddish shade sometimes extending outward 
for a short distance along lower fold, reappearing in 
tornal area; antemedial line distinct, rather wide, nearly 
straight, and in many specimens apparently vertical but 
actually a trifle oblique; subterminal line obscure, paral- 
lel to termen, slightly and irregularly denticulate; discal 
dots sometimes obscured by the blackish dusting of 
median area but usually distinct, more or less confluent 
and set off by some surrounding pale dusting which 
extends in well marked specimens to costa. Hind wing 
white, smoky white or pale fuscous; veins darkly out- 
lined, some dark shading and a narrow dark line along 
termen. Alar expanse, 14-20 mm. 

Male genitalia with gnathos terminating in a short, 
bluntly pointed, straight spike; elements of transtilla 
thin, broad, lightly sclerotized, fusing with arms of 
gnathos and extending beyond them to subanal plate 
(fig. 6112); harpe with a very short digitate projection 
from basal third of costa, apex broadly rounded; vin- 
culum long, its terminal margin sharply angled; aedea- 
gus long and stout. Female genitalia with ductus 
bursae sclerotized for about half its length, the sclero- 
tized portion longitudinally wrinkled for its entire 
length; signum a longitudinal series of four or five small 
discs. 

Tyrr Locaities: Blanco County, Tex. (nigrella, in 
USNM); Tempe, Ariz. (arizonella, in USNM). 

Foop prant: Unknown. 

Distrrsution: Teras, Blanco County (Aug.), Browns- 
ville (Mar.), San Benito (Mar., June, July, Sept.); 
Arizona, Catalina Springs (May), Mohave County 
(Sept.), Redington, Tempe (Aug., Sept.), Yuma (June) ; 
California, Death Valley (Apr.), La Puerta Valley 
(July), Palm Springs (Mar.). 

Like Moodna ostrinella a variable species in color. 
The name arizonella refers only to a color form with 
pale reddish ocherous shading on the basal area of fore- 
wing. Its genitalia (male and female) agree in every 
detail with those of typical nigrella. The latter name 
has been “kicked around” rather carelessly by later 
authors since its original placement by Hulst. He 
alone seems to have examined the venation, which would 
allow reference to Ephestia but not to Moodna and 
certainly not to Mescinia. 


293 


605. Caudellia declivella (Zeller), new combination 
Ficures 612, 1104 


Ephestia declivella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 
244, 1881.—Hampson, in Ragonot, Monograph, pt. 2, p. 
305, 1901.—Dyar, Proc. Ent. Soc. Washington, vol. 31, p. 
17, 1929.—Richards and Thomson, Trans. Ent. Soc. London, 
vol. 80, p. 175, 1932. 

Ephestia animosella Dyar, Proc. U. 8S. Nat. Mus., vol. 47, p. 345, 
1914. 

Forewing red heavily dusted with black especially in 
the median area, the red shade more obvious in basal 
and terminal areas and along the fold; antemedial line 
strongly contrasted, white, narrow, straight, oblique; 
from antemedial line to base a subcostal streak of white 
scaling (only distinguishable in fresh specimens); sub- 
terminal line faint and very narrow, near to and parallel 
with termen; discal dots obscure, some white dusting in 
the area surrounding them. Hind wing pale trans- 
lucent fuscous, darker on the veins and towards terminal 
margin. Alar expanse, 10-13 mm. 


Male genitalia with transtilla complete, developed as 
a bridge with humped central projection; apical projec- 
tion of gnathos a very short, blunt, knoblike hook; 
harpe simple; aedeagus rather slender. 


Female genitalia with ductus bursae sclerotized for 
most of its length, some sclerotized ridges at the junc- 
tion with bursa copulatrix; signa a cluster of minute, 
blunt discs. 


Typr Locauitigs: Honda, Colombia (declivella, in 
BM); Porto Bello, Panama (animosella, in USNM). 

Foop puant: Unknown. Larva probably a scavenger 
on dried vegetable matter. 


Distrinution: PanamA: La Chorrera (Apr.), Porto 
Bello, (May), Rio Trinidad (Mar., May, June). 
CotomeraA: Honda, Maraquita. 


The species is easily recognized by its male genitalia 
which are similar to those of Manhatia biviella of 
Europe, but unlike anything else from the Americas. 


606. Caudellia colorella (Dyar), new combination 
Ficures 614, 1101 


Ephestia colorella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 345, 
1914.—Richards and Thomson, Trans. Ent. Soc. London, 
vol. 80, p. 192, 1932. 

Superficially similar in every respect to declivella 
except that reddish areas of forewing are paler, ocherous 
red rather than dull red as in declivella. In the hind 
wing veins 3 and 5 are sometimes very shortly stalked. 
They are occasionally so in declivella, but in the latter 
normally are connate (not separate as Dyar states in 
his original description of animosella). Alar expanse, 
11-14 mm. 

Male genitalia with transtilla completely fused with 
gnathos and greatly broadened posteriorly. Harpe 
with short, blunt projections from costa at middle and 
apex; uncus broadly triangulate. Female genitalia 
with ductus bursae sclerotized for two-thirds its length 
from genital opening, not sclerotized at junction with 
bursa, 


‘294 


Typr LocaLity: Taboga Island, Panamé (type in 
USNM). 

Foop pirant: Unknown. 

DistRiBuTIoN: Panamd: Cabima (May), Corozal 
(Mar.), Porto Bello (May), Taboga Isl. (Feb.). 


607. Caudellia clara, new species 
Figure 615 


Pattern and color of forewing much as in the two pre- 
ceding species except the red shading more confined to 
‘streaks along the folds and rather inconspicuous; white 
antemedial line and the white extension from it along 
costa to base more strongly contrasted; general ground 
color purplish fuscous; discal dots rather well marked 
‘and set in a clear white field. Hind wing whitish; the 
veins and a narrow line along termen pale fuscous. 
Alar expanse, 13.5 mm. 

Male genitalia with elements of transtilla very broad 
and completely fused with arms of transtilla; gnathos 
terminating in an elongate, stout, blunt hook; uncus 
broad throughout, its terminal margin straight; harpe 
with an enlarged flattened scooplike projection from 
base of costa; costa broadly sclerotized and slightly and 
bluntly projecting at apex. Female unknown. 

TypE Locality: Hl Yunque, Luquillo Mts., Puerto 
Rico (type in Cornell Univ. Coll.; paratype in USNM, 
61393). 

Foop eLant: Unknown. 

Described from male type and one male paratype 
from the type locality, 1,500 to 2,000 ft., Cornell lot No. 
795, sub. 38, Apr. 22, 1930. 

_ A distinct species, but in color and maculation hardly 
separable from declivella. However, the male genitalia 
are distinctive and easily identify the species. 


181. Genus Microphestia Dyar 


Microphestia Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 346, 1914. 
(Type of genus: Microphestia animalcula Dyar.) 

Tongue short (but not completely enclosed by labial 
palpi). Antenna of female roughly scaled. Labial 
palpus oblique, slender, reaching nearly to vertex; third 
segment slightly shorter than second. Mazxillar palpus 
minute, filiform. Forewing smooth; 9 veins; vein 2 
from very near lower, outer angle of cell; 2, 3, and 5 
slightly separated and approximately equidistant at 
base; 4 absent; 6 from below upper angle of cell, 
straight; 9 absent; 10 from cell, separated from 8 at 
base. Hind wing with 2 from well before lower angle 
of cell; 3 and 5 shortly stalked (not long stalked as 
stated by Dyar); 7 and 8 completely anastomosed 
beyond cell; cell one-half the length of wing; discocellu- 
lar vein curved. 

Female genitalia with ductus bursae sclerotized for 
most of its length from genital opening, the sclerotized 
area flattened; a triangulate, projecting, sclerotized 
shield behind genital opening. 

The above diagnosis is incomplete, as the male is 
unknown. The genus is doubtfully distinct from 


“UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Caudellia. It is distinguished from the latter chiefly 
by its reduced tongue and the small size of its type, 8 
mm. 
608. Microphestia animalcula Dyar 
Figure 1114 


Microphestia animaleula Dyar, Proc. U. S. Nat. Mus., vol. 47, 
p. 346, 1914. 


Forewing unicolorous, dark glossy brown; under 
magnification the faintest indication of a thin, pale, 
oblique antemedial line; no other markings. Hind wing 
pale fuscous. Alar expanse, 8 mm. 

Female genitalia having bursa without signum; 
ductus bursae no longer than bursa; ductus seminalis 
from approximately middle of bursa. 

Typr LocaLity: Rio Trinidad, Panam4é (Mar.; type 
in USNM). 

Foop prant: Unknown. 

Known only from the unique female type. 


182. Sosipatra, new genus 


TyprE oF GENus: Ephestia rileyella Ragonot 

Characters of Caudellia except: Shaft of male antenna 
simple; labial palpus oblique or erect; apical process of 
gnathos enlarged (broadened), undivided, knobbed or 
looped; harpe with apex of costa produced as short 
spine at apex, or apex of costa and cucullus upturned; 
transtilla complete, an angulate bridge entirely free of 
gnathos; ductus seminalis from bursa copulatrix near 
its junction with ductus bursae; eighth abdominal seg- 
ment of male with compound dorsal tufts or simple. In 
the hind wing, veins 3 and 5 are either connate or 
shortly stalked. Signum, when present, a single, blunt, 
thornlike disc, adjacent to ductus seminalis. 


The new genus brings together a group of American 
species having a consistent female character in the 
position of ductus seminalis in relation to the bursa and 
a male character in the broadened apical process of 
gnathos. On the harpe and the eighth abdominal seg- 
ment of the male it divides into two groups as follows: 


Harpe with costa produced at apex into a short spine; eighth 
abdominal segment of male with compound tufts. 
Harpe with apex of costa and cucullus upturned; eighth ab- 

dominal segment of male simple. 


Genus Sosipatra, Species 609-612: S. rileyella to 
S. thurberiae 


[Harpe with costa produced at apex into a short spine; eighth 
abdominal segment of male with compound tufts.] 


609. Sosipatra rileyella (Ragonot), new combination 
Figures 616, 1105 


Ephestia rileyella Ragonot, N. Amer. Phycitidae, p. 17, 1887.— 
Hulst, Phycitidae of N. Amer., p. 198, 1890.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 294, 1901.—Richards and 
Thomson, Trans. Ent. Soc. London, vol. 80, p. 182, 1932.— 
McDunnough, Check list, No. 6400, 1939. 


Forewing cream white very sparsely sprinkled with 
black scales; costal edge for half the distance from base 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


to antemedial line black; antemedial line indicated by 
two black spots, one above the other, on median and 
lower folds, rarely with additional black dots above and 
below them; subterminal line indicated by an oblique 
series of black spots on the veins; discal spot on lower 
outer angle of cell conspicuous, black, the discal spot 
on upper angle of cell more or less obsolescent. Hind 
wings translucent, shining white; a fine brown line 
along termen; veins faintly, if at all, darkened. Alar 
expanse, 15-23 mm. 

Male genitalia distinguished chiefly by the shapes of 
transtilla (fig. 616b) and apical process of gnathos 
(fig. 616a). Female genitalia with signum. 

Type Locauity: Utah (type in Paris Mus.). 

Foop piants: Yucca, Nolina. 

Distrisution: Unirep States: Utah, Hureka (June), 
Penah (Feb.); Colorado, Grand Junction (July); Cali- 
fornia, Loma Linda (Mar.), Los Angeles County (June, 
July, Sept.), Mohave (May), Morongo (Apr.), Phelan 
(May), Pipes Canyon (San Bernardino Mts.; Mar., 
Apr., May), San Pasqual (Apr.); Arizona, Catalina 
Springs (May), Chiricahua Mts. (Apr., May), Mohave 
County (Sept., Oct.), Phoenix (Apr.), Pinal Mts. (May); 
New Mexico, Albuquerque (July), Valencia County; 
Texas, Brownsville. M*xtco: Sonora. 

A species easily recognized by its wing color and 
maculation and its close association with Yucca. The 
larva feeds upon the dry seeds in the pods of several 
species of that genus. There are also in the National 
Collection three specimens reared in 1939 by W. D. 
Pierce from larvae feeding in the seed pods of Nolina 
parryi at Pipes Canyon, San Bernardino Mts., Calif. 
N. parryi is a plant closely related to Yucca and by some 
botanists classified under the latter genus. 


610. Sosipatra micaceella (Hampson), new combination 
Figure 617 


Ephestia micaceella Hampson, in Ragonot, Monograph, pt. 2, 
p. 298, 1901. 

Forewing gray-brown powdered with black, the black 
dusting heaviest on the costal half of wing, under mag- 
nification some white scaling that gives the costal mar- 
gin the bluish gray tint mentioned by Hampson; two 
black spots on costa near base; antemedian line faint 
but distinguishable, pale gray, well out towards middle 
of wing and nearly vertical, bordered outwardly by 
some black dashes, the latter most pronounced on upper 
margin and middle of cell and on lower fold, the two 
upper dashes more or less confluent and extended into 
the cell; subterminal line oblique, parallel to termen, 
inwardly angled at vein 6 and slightly so at lower fold, 
bordered inwardly by a black line (broken more or less 
into short dashes on the veins); discal spots confluent, 
forming a black line along discocellular vein; an obscure 
blackish line along termen. Hind wing translucent, 
white; the veins outlined by pale fuscous, a fuscous 


295 


shade along costa and narrowly bordering the termen. 
Alar expanse, 18-19 mm. 

Male genitalia similar to those of rileyella except for 
slight differences in the shapes of transtilla and apical 
process of gnathos. 

The female is unknown. 

Type Locauity: México (type in the collection of the 
Abbé Joannis). 

Foop prant: Unknown. The type was reared from 
a larva found in a cocoon of (Attacus) Rothschildia ori- 
zaba (Westwood) but the larval habits were not noted. 

Known only from México. There are two males in 
the National Collection from the city of México (Nov.). 
One of these is badly rubbed; but the other is in fair con- 
dition, only the lower half of the forewing being some- 
what rubbed, and it answers well to Hampson’s descrip- 
tion. Richards and Thomson in their paper on Hphestia 
(Trans. Ent. Soc. London, vol. 80, p. 183, 1932) give a 
description of micaceella from a male so determined by 
Hampson in the British Museum. They publish no 
figures, but describe the genitalia; and their description 
raises some doubt as to what they had, for they state 
that the ‘dorsal thickenings” of anellus (our transtilla) 
are “absent, replaced by broad ventral thickenings of 
the tegumen itself.” If they had an example of mica- 
ceella before them, this is obviously a misinterpretation 
of structure from a poor preparation; for micaceella has 
as distinct a transtilla as rileyella and no broad ventral 
thickening of the tegumen. No phycitid in any nearly 
related group has such a tegumen. 


611. Sosipatra anthophila (Dyar), new combination 
Fiaures 618, 1106 
Eurythmia anthophila Dyar, Ins. Insc. Menstr., vol. 13, p. 226, 
1925.—McDunnough, Check list, No. 6394, 1939. 

Forewing bluish gray, rather broadly shaded with 
white along costa and with some faint white dusting in 
terminal area; antemedial line slightly angulate, white 
outwardly bordered by blackish shading, especially be- 
low the white costal suffusion; subterminal line thin, 
white, practically straight, bordered inwardly by a nar- 
row blackish shade and outwardly for a short distance 
from costa by a similar blackish shade; discal dots dis- 
tinct, separate; terminal dots obscure. Hind wing 
white, translucent; a narrow, pale yellowish fuscous line 
along termen. Alar expanse 15-16 mm. 

Male genitalia with apical process of gnathos a short, 
stout, blunt, oval, scooplike hook; transtilla arched, 
flattened at middle; vinculum with terminal margin 
angulate. Female genitalia with the bursa somewhat 
more strongly scobinate than in other species of the 
genus, especially near and at junction of bursa and duc- 
tus bursae; signum present. 

Type Locatity: Uvalde, Tex. (type in USNM). 

Foop piant: Opuntia (larva feeding in the flowers). 

Known only from the type series, a male from Uvalde 
(May) and one male and one female from Devils River, 
Tex. (May). 


296 


612. Sosipatra thurberiae (Dyar), new combination 
Ficures 619, 1107 


Eurythmia thurberiae Dyar, Ins. Inse. Mensir., vol. 5, p. 46, 
1917.—McDunnough, Check list No. 6393, 1939. 

Forewing bluish gray more or less dusted with black, 
in some specimens the entire area between the trans- 
verse lines suffused with black, a fine whitish dusting 
usually sets off the entire basal area; antemedial line 
well out towards middle of wing, narrow and nearly 
vertical, narrowly bordered outwardly by some accentu- 
ation of the black dusting; subterminal line parallel to 
termen, slightly angled inwardly at vein 6 and lower 
fold; discal dots more or less distinct (at least the lower 
one) and from them a broadening wedge of pale scaling, 
extending to costa; terminal dots obscure, more or less 
confluent. Hind wing white to pale smoky fuscous; 
the veins more or less darkened and a narrow dark line 
along termen. Alar expanse, 11-19 mm. 

Male genitalia similar to those of anthophila but 
easily distinguished by the shapes of transtilla and 
apical process of gnathos; terminal margin of vinculum 
evenly rounded. Female genitalia with very slender 
signum. 

TYPE LOCALITY: Bowie, Ariz. (type in USNM). 

Foop puants: Cercis occidentalis (larvae in pods), 
Quercus (larvae in “oak-apple” galls on leaves), Thur- 
beria (larvae in seed pods). 

Distrisution: Arizona, Bowie (June); California, 
Applegate (July, Aug.), Buelton (July), Calpella (July), 
Gasquet (July), Hopeland (July), San Felipe Wash 
(San Diego County, June), Three Rivers (Apr.); 
Oregon, Dundee (Aug.), Eugene (July), Woodburn 
(Aug.). 


Genus Soszpatra, Species 613-615: S. nonparilella 
to S. diwergens 


[Harpe with apex of costa and cucullus upturned; eighth ab- 
dominal segment of male simple.] 


613. Sosipatra nonparilella (Dyar), new combination 
Figure 621 


Ephestia nonparilella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 118, 1904.—McDunnough, Check list No. 6404, 1939. 

Forewing white dusted with black, giving the wing a 
pale ashy gray ground color; discal spots and borders of 
transverse lines black and strongly contrasted; ante- 
medial line indicated chiefly by its outer border, far out 
on wing (about middle), just below costa angled inward 
and running parallel with subterminal line; the latter 
white and straight except for a very slight notch at 
lower fold, parallel with and rather close to termen; 
discal dots confluent, forming a black streak along 
discocellular vein. Hind wing white, translucent; its 
terminal margin but slightly darkened. Alar expanse, 
18 mm. 

Male genitalia distinguished by shape of apical 
process of gnathos (fig. 621a). 

TYPE LOCALITY: Santa Rita Mts., Ariz. (June; type 
in USNM). 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Foop prant: Unknown. 

Known only from the male type. Dyar’s original 
description is in error in one particular. He states that 
veins 3 and 5 of hind wing are “separate, but approxi- 
mate at base.’’ They are distinctly connate. 


614. Sosipatra majorella (Dyar), new combination 
Fieures 620, 1108 


Ephestia majorella Dyar, Ins. Insc. Menstr., vol. 10, p. 173, 1922. 


Forewing similar to that of micaceella but with less 
contrast between the costal and inner areas, and the 
black borders of the transverse lines broader, more 
complete (not broken into dashes); ground color dark 
gray-brown with some lighter rust-brown scaling in the 
folds; antemedial line slightly angled just below costa; 
the black inner border of subterminal line angled out- 
wardly at middle; discal spots confluent, black. Hind 
wing of male light brown, of female, whitish fuscous; 
the veins and terminal margin darker. Alar expanse, 
20 mm. 

Male genitalia similar to those of nonparilella except 
for a somewhat broader transtilla, more slender aedea- 
gus and differently shaped apical projection of gnathos. 
Female genitalia with signum; bursa otherwise smooth. 

Type nocanity: Guadalajara, México (type in 
USNM). 

Foop puant: Unknown. 

Tn addition to the male type there is in the National 
Museum from Mexico City (Aug.) a female which 
Schaus or Dyar had associated with micaceella. Its 
maculation is a much better match for that of majorella, 
with which I associate it. The two species differ radi- 
cally in male genitalia butaresimilarin colorand pattern 
and easily confused. On the other hand, while they 
differ markedly in color, the types of majorella and non- 
parilella exhibit only minor structural differences. 


615. Sosipatra divergens (Dyar) 
Figure 1109 


Ephestia divergens Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 345, 
1914. 


Forewing dark gray-brown; antemedial line vertical, 
at middle of wing, bordered outwardly by black; sub- 
terminal line oblique, slightly curved at fold, margined 
within by a thin black line; discal spots confluent, form- 
ing a blackish line along discocellular vem. Hind wing 
pale fuscous, veins and terminal margin very slightly 
darker. Alar expanse, 16 mm. 

Female genitalia without signum. Male unknown. 

Type tocatity: Taboga Isl., Panama (type in 
USNM). 

Foop prant: Unknown. 

Known only from the unique female type. 


183. Genus Bethulia Ragonot 


Bethulia Ragonot, Nouv. Gen., p. 36, 1888.—Hampson, in Rag- 
onot, Monograph, pt. 2, pp. xiv, 804, 1901. (Type.of genus: 
Bethulia championella Ragonot.) 


The generic descriptions of Ragonot and Hampson are 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 297 


faulty in several details. The type of championella (and 
only known example of the genus) is a female and not a 
male as stated by Ragonot and Hampson. The vena- 
tion is similar to that of Plodia—veins3 and 5 areslightly 
separated at base, not from a point, and 8 and 5 of hind 
wings are connate and not shortly stalked. The female 
genitalia are also similar to those of Plodia except that 
there is no signum. The labial palpi are oblique. 

The genus is obviously very close to Plodia and Ribua 
but in the absence of a male it is impossible to charac- 
terize it properly or determine its status accurately. 


616. Bethulia championella Ragonot 
Ficures 123, 1120 


Bethulia championella Ragonot, Nouv. Gen., p. 37, 1888; Mono- 
graph, pt. 2, p. 304, 1901.—Druce, Biologia Centrali-Amer- 
icana, Lepidoptera, Heterocera, vol. 2, p. 287, 1896. 

The Ragonot figure of the type (Monograph, pl. 35, 
fig. 16) is somewhat misleading. An enlarged photo- 
graph, before me, shows the blackish margins defining 
the antemedial and subterminal lines stronger and some- 
what broader than those in the published figure and a 
distinct scattering of blackish scales over the white areas 
of the forewing; the antemedial line is angulate (not 
“oblique’’) and the black border of the subterminal is 
complete (not broken as in the figure) and shows a 
sharp angulation at vein 6. 

In female genitalia the projecting shield behind geni- 
talia is subtriangulate and broadly flaring to its wide, 
slightly convex terminal margin. 

If other specimens are ever received from Central 
America they should be easily identified; for champion- 
ella is the only described white species, except Socipatra 
rileyella, in any of the genera, of similar venation and 
with the ductus bursae sclerotized and flattened for part 
ofitslength. From rileyella it is easily distinguished by 
its genitalia. 

Type tocauity: San Joaquin, Vera Paz, Guatemala 
(type in BM). 

Foop pLant: Unknown. 

Known only from the type specimen. 


184. Genus Ribua Heinrich 


Ribua Heinrich, Proc. Ent. Soc. Washington, vol. 42, p. 31, 1940. 
(Type of genus; Ribua innozxia Heinrich.) 

Tongue well developed. Antenna simple and pubes- 
cent. Labial palpus oblique in male, porrect in female 
(the third segment downcurved). Maxillary palpus 
minute, filiform. Forewing smooth; 9 veins; vein 2 
from just before lower outer angle of cell; 3 from the 
angle; 4 absent; 5 closely approximate to 3 at base; 6 
from below upper angle of cell, straight; 8 and 9 united; 
10 from the cell, closely approximate to 8 for some dis- 
tance from cell; male with costal fold enclosing a scale 
tuft. Hind wing with vein 2 from well before outer 
angle of cell; 3 and 5 approximate at base; 7 and 8 com- 
pletely anastomosed beyond cell (rarely, in individual 
specimens, with 8 represented as a short spur at costa) ; 
cell slightly less than one-half the length of wing; disco- 


cellular vein curved. Abdomen of male with a pair of 
dorsolateral hair tufts on eighth segment. 

Male genitalia with apical process of gnathos asym- 
metrical, bearing two short hooks. Transtilla a narrow 
band with flattened central process fusing to apical proc- 
ess of gnathos (fig. 622). Harpe with costa slightly pro- 
duced at apex, otherwise simple. Vinculum long and 
broad. 

Female genitalia with ductus bursae flattened and 
sclerotized for most of its length; bursa copulatrix with 
or without signa, latter when present consisting of one 
or two small, bluntly pointed, disclike spines, ductus 
seminalis from near anterior end of bursa. 

The genus is close to but distinct from Plodia, dis- 
tinguished by the following combination of characters: 
Veins 3 and 5, and 8 and 10 closely approximate at base; 
labial palpus of male oblique; tufts on eighth abdominal 
segment of male simple; apical process of gnathos asym- 
metrical and bifurcate; anellus fusing with gnathos; 
ductus seminalis from near anterior end of bursa. 


617. Ribua innoxia Heinrich 
Figures 622, 1115 


Ribua innozia Heinrich, Proc. Ent. Soc. Washington, vol. 42, 
p. 32, 1940, 

Forewing ash gray, obscurely marked and shaded with 
blackish fuscous; an ill-defined dark basal patch; ante- 
medial line faintly indicated, oblique, shaded outwardly 
by a blackish band; subterminal line obscure, pale, 
straight and parallel with termen, narrowly shaded 
inwardly and outwardly by dark scaling; the veins 
faintly dark-shaded; a scattering of dull reddish scales 
on the wing, especially along lower fold, on upper vein 
of cell and bordering subterminal line; all marking 
obscure and dark shading more or less diffused. Hind 
wing dull white; a narrow fuscous shade along costa and 
termen, most conspicuous at apex; veins slightly dark- 
ened. Alar expanse, 12-16 mm. 

Male genitalia with prongs of apical process of 
enathos separated (space between them U-shaped); 
vinculum nearly twice as long as tegumen. Female 
genitalia with two signa. 

TYPE Locatity: Cuba (type in USNM). 

Foop piant: Fungus on pineapple. 

The larvae of this species are frequently intercepted 
at our southern ports on pineapples from Cuba, but 
apparently do no damage to the fruit. A moth was 
recently received from Cuba which had been reared 
from a larva in decaying sugarcane. Apparently the 
species is more of a scavenger than anything else. 


618. Ribua contigua, new species 
Figures 623, 1117 


Forewing reddish brown; black dusting sparser than 
on innoria and largely replaced by red scaling; trans- 
verse lines obliterated. Alar expanse, 12-14 mm. 

Male genitalia with prongs of apical process of 
gnathos close together, the space between them narrowly 
V-shaped and the prongs much shorter than those of 


298 


innoxia; vinculum shorter (notover oneand one-half times 
the length of tegumen) and broader throughout. than 
that.of innoxia. Female genitalia not exhibiting any 
specific characters to distinguish them from ‘those of 
innoxia. 

TypE LocaLity: Dorado, Puerto Rico (type in Cor- 
nell Univ. Coll.; paratype, o, in USNM, 61894). 

Foop prant: Unknown. 

Described from male type and female paratype from 
the type locality, collected May 30, 1930, by W. A. 
Hoffman, and two male paratypes from Catafio, Puerto 
Rico, collected by Leonard and Mills, May 16 and July 
24, 1930. 

Nothing is known of its life history, but its larval 
habits are probably similar to those of innomia. 


619. Ribua patriciella (Dyar), new combination 
Fieure 1116 


Ephestia patriciella Dyar, Ins. Insc. Menstr., vol. 6, p. 140, 1918. 


Forewing coloration similar to that of contigua, but 
the reddish dusting more towards a brown shade and 
less generally distributed; the veins outlined in blackish 
gray which predominates over the red-brown on most of 
the wing; transverse lines and discal spots obsolete. 
Alar expanse, 12 mm. 

Female genitalia without signum. 

TypE wLocauity: Baracoa, Cuba (Oct.; 
USNM). 

Foop piant: Unknown. 

Known only from two specimens in poor condition, 
the type and one female paratype from Santiago, Cuba 
(May). The labial palpi seem more oblique than por- 
rect, but they are twisted so much that their shape can 
not be accurately determined. ‘The male is unknown. 
The three males that Dyar included among his para- 
types are Hphestia cautella. 


type in 


185. Genus Plodia Guénée 


Plodia Guénée, Europaeorum Microlepidopterorum index me- 
thodicus . . . p. 80, 1845.—Hinemann, Die Schmetterlinge 
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 202, 
1865.—Snellen, De Vlinders van Nederland, Microlepidop- 
tera, vol. 1, p. 163, 1882.—Hulst, Phycitidae of N. Amer., p. 
200, 1890.—Ragonot, Monograph, pt. 2, p. xiv, 1901.— 
Hampson, in Ragonot, Monograph, pt. 2, p. 305, 1901.— 
Spuler, Die Schmetterlinge Europas, vol. 2, p. 201, 1910.— 
Forbes, Cornell Mem. 68, p. 636, 1923.—Meyrick, Revised 
handbook of British Lepidoptera, p. 386, 1938.—Richards 
and Thomson, Trans. Ent. Soc. London, vol. 80, p. 203, 
1932.—Pierce and Metcalf, Genitalia of the British Pyrales 
p. 6, 1938.—Janse, Journ. Ent. Soc. South Africa, vol. 8, p. 
26, 1945. (Type of genus: Tinea interpunctella Hubner.) 


Tongue well developed. Antenna simple and pu- 
bescent in both sexes. Labial palpus porrect. Maxil- 
lary palpus minute, filiform. Forewing smooth; 9 
veins; vein 2 from before, but near, lower outer angle 
of cell; 3 from the angle; 4 absent; 5 appreciably sepa- 
rated from 3 at base; 6 from below upper angle of cell, 
straight; 8 and 9 united; 10 from the cell, separated from 
8 at base; male with costal fold enclosing a scale tuft. 
Hind wing with 2 from well before outer angle of cell; 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


3 and 5 connate; 7 and 8 anastomosed from most or all 
of their lengths beyond cell; cell about half the length 
of wing; discocellular vein curved. Abdomen of male 
with two pairs of dorsal hair tufts, or compound dorsal 
tufts on eighth segment. 

Male genitalia with apical projection of gnathos a 
small knob terminating in a very short, simple, blunt 
hook. ‘Transtilla a narrow, curved, sclerotized band, 
not in any way fusing to gnathos. Uncus for most of 
its length rather narrow, its sides straight or but slightly 
tapering to bluntly apical margin. Harpe with a very 
slight, rounded projection from costa at middle and 
a short thornlike projection at apex, otherwise simple. 

Female genitalia with ductus bursae flattened and 
sclerotized for over half its length, shorter than bursa; 
signa, present, developed as a cluster of 3 to 5 contig- 
uous, small, blunt, thornlike projections; ductus semi- 
nalis from about middle of bursa, near signa. 

The genus is distinguished from Ribua by the follow- 
ing combination of characters: Labial palpi of both 
Sexes porrect; veins 3 and 5 of forewing distinctly 
separated at base; 10 from cell and not close to 8 at 
base; 3 and 5 of hind wing connate; eighth abdominal 
segment of male with two or more pairs of dorsal tufts; 
ductus seminalis from middle of bursa copulatrix. In 
the Ragonot key to genera (Monograph, p. xiv) veins 3 
and 5 of hind wing are said to be separate; but this is an 
error. ‘They are definitely connate. 

Plodia is a genus of definitely American origin. Its 
type species has become cosmopolitan through trans- 
portation in ship stores and the commercial distribution 
of dried grains and fruits; but all its nearest relatives 
(Ribua, Caudellia, Bethulia, etc.), and the only other 
known congeneric species (dolorosa), are confined to the 
New World. 


620. Plodia interpunctella (Hiibner) 
Fiegures 124, 624, 1118 


Tinea interpunctella Hiibner, Sammlung europdischer Schmetter- 
linge, Lepidoptera 8, Tineae 5, pl. 45, fig. 310 [1810]-[1813]. 

Elucita interpunctalis Hiibner, Verzeichnis bekannter Schmett- 
[erJlinge, p. 347, 1825 (change in spelling for interpunctella). 

Phycita interpunctella (Hiibner) Treitschke, Die Schmetterlinge 
von Europa, vol. 10, p. 196, 18382.—Duponchel, Histoire 
naturelle des lépidoptéres .. . de France, vol. 10, p. 224, 
1836. 

Myelois interpunctella (Hiibner) Zeller, Isis von Oken, 1839, p. 
176. 

Plodia interpunctella (Hiibner) Guénée, Europaeorum Microlepi- 
dopterorum index methodicus ... , p. 80, 1845.—Heine- 
mann, Die Schmetterlinge Deutschlands und der Schweiz, 
Abt. 2, vol. 1, pt. 2, p. 202, 1865.—Snellen, De Vlinders van 
Nederland, Microlepidoptera, vol. 1, p. 163, 1882.—Ragonot, 
Ent. Monthly Mag., ser. 2, vol. 22, p. 25, 1885.—Hulst, 
Phycitidae of N. Amer., p. 200, 1900; U. S. Nat. Mus. Bull. 
52, p. 487, 1903.— Hampson, in Ragonot, Monograph, pt. 
2, p. 305, 1901.—Spuler, Die Schmetterlinge Europas, vol. 
2, p. 201, 1910.—Forbes, Cornell Mem. 68, p. 636, 1923.— 
Curran, Sci. Agr., vol. 6, p. 386, 1926.—Hudson, Butterflies 
and moths of New Zealand, p. 156, 1928.—Meyrick, Revised 
handbook of British Lepidoptera, p. 386, 1928.—Noyes, 
Bull. Ent. Res., vol. 21, p. 81, 1930.—Richards and Herford, 
Ann, Appl. Biol., vol. 17, p. 380, 1930.—Hamlin, Reed, and 
Phillips, U. S. Dep. Agr. Techn. Bull. 242, 26 pp., 1931.— 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Richards and Thomson, Trans. Ent. Soc. London, vol. 80, 
p. 205, 1932.—Norris, Proc. Zool. Soc. London, pp. 597-611, 
1932.—Dickins, Trans. Ent. Soc. London, vol. 85, p. 338, 
1936.—Barth, Zool. Jarb., Jena, vol. 58, pp. 297-329, 1937; 
Zeitschr. Wiss. Zool., Leipzig, vol. 150, p. 27, 1938.—Leh- 
mensick and Liebers, Zeits. Angew. Ent., Berlin, vol. 24, 
p. 441, 1937; p. 582, 1938.—McDunnough, Check list, No. 
6408, 1939.—Hinton, Bull. Ent. Res., vol. 34, p. 185, 1943.— 
Corbet and Tams, Proc. Zool. Soc. London, p. 64, 1943. 

Ephestia interpunclella (Hiibner) Herrich-Schiffer, Systematische 
Bearbeitung der Schmetterlinge von Europa, vol. 4, p. 110, 
1849.— Westwood and Humphreys, British moths and their 
transformations, p. 274, 1854.—Stainton, Manual of 
British butterflies and moths, vol. 2, p. 169, 1859.—Zeller, 
Verh. zool.-bot. Ges. Wien, vol. 25, p. 336, 1876; vol. 34, p. 
310, 1884.—Riley and Howard, Insect Life, vol. 2, p. 277, 
1890. 

Tinea zeae Fitch, Second report on the noxious, beneficial and 
other insects, of the State of New York, p. 320, 1856. 

Ephestia zeae (Fitch) Clemens, Proc. Acad. Nat. Sci. Philadel- 
phia, p. 206, 1860. 

Ephestia interpunctalis (Hiibner) Butler, Ent. Monthly Mag., 
vol. 15, p. 278, 1879.—Druce, Biologia Centrali-Americana, 
Lepidoptera, Heterocera, vol. 2, p. 286, 1896. 

Unadilla latercula Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7, 
p. 255, 1901 (New synonymy). 

Ephestia glycinivora Matsumura, ‘“‘Oyo-Kinchyugakii”’ (Applied 
entomology), pt. 1, p. 561, 1917; Dai-Nippon Gaichyu 
Zensho (Injurious insects of the Japanese Empire), ed. 2, 
vol. 1, p. 529, 1920 (spelled here as glycinivorella).—Shibuya, 
Konchu Sekai, vol. 36, p. 225, 1982 (makes synonym of 
interpunctella). 

Ephestia (Strymaz) latercula (Hampson) Richards and Thomson, 
Trans. Ent. Soc. London, vol. 80, p. 202, 1932. 


Forewing with basal area ocherous white to pale 
ocherous, in fresh specimens well contrasted against re- 
mainder of wing; area beyond antemedial line rusty 
ocherous to reddish fuscous, the red shading marked 
along costal edge; central area more or less heavily 
dusted with leaden fuscous, the fuscous scaling forming 
an irregular blotch surrounding an ocherous line along 
the discocellular vein; transverse lines sublustrous, 
leaden lined; antemedial line rather broad, irregular, 
set well out towards middle of wing, oblique and 
slightly curved; subterminal line obscure, somewhat 
sinuate, more or less parallel with termen; a faint leaden 
line along termen. Hind wing pale, shining, fuscous; 
veins and terminal margin faintly darkened. The red 
scaling is pronounced on thorax and labial palpi as well 
as on costa of forewing. Alar expanse, 12-19 mm. 

Male genitalia with transtilla a smooth, uninterrupted 
band; uncus not at all expanded at apex. Female geni- 
talia with sclerotized projection of ductus bursae behind 
genital opening rather broad, variously shaped in indi- 
vidual specimens, angulate or rounded. Number of 
spines of signa also variable. 

TYPE LocauitTiEs: Europe (interpunctella, type lost); 
New York (zeae, in USNM); Nassau, Bahamas (later- 
cula, in BM); Japan (glycinivora, in Hokkaido Imperial 
Univ., Sapporo, Japan). 

Foon: All kinds of stored and dried vegetable prod- 

ucts. 
_ Distrrpution: Cosmopolitan. 


This well-known and ubiquitous pest (‘‘the%Indian 
meal moth” of economic literature) requires little com- 


299 


ment. It has an enormous literature. I have cited 
above only the more important systematic references 
and papers dealing with the physiology, morphology 
and biology of the insect. The fullest treatment of 
Plodia interpunctella and the common Ephestia species 
will be found in the paper by Richards and Thomson 
and that by Lehmensick and Liebers (1938). They 
contain an extended bibliography, as does also the paper 
by Hamlin, Reed, and Phillips. For additional refer- 
ences the “Index of American Economic Entomology” 
and the ‘Review of Applied Entomology” should be 
consulted. 

Hampson’s latercula, here placed in synonymy, has 
none of the diagnostic characters of the genus Unadilla 
in which he placed it. Itis obviously a Plodia. Ihave 
seen the moth from Colombia he associated with his 
type and have examined its genitalia; and Clarke and 
Tams have compared the latter with the genitalia of the 
type from the Bahamas. The moths themselves are in 
very poor condition and show the basal area of the wing 
somewhat darker than in typical interpunctella, but the 
female genitalia show variation only of an individual 
nature. 

621. Plodia dolorosa Dyar 


Figures 625, 1119 


Plodia dolorosa Dyar, Ins. Inse. Menstr., vol. 7, p. 63, 1919.— 
Richards and Thomson, Trans. Ent. Soc. London, vol. 80, 
p. 204, 1932. 

Forewing dark gray-brown (on the female type, the 
only unrubbed specimen before me, with a purplish 
sheen), the veins faintly darkened by black scaling; 
transverse lines and discal marks obsolete. Hind wings 
whitish fuscous, darker on the female, with veins clearly 
outlined by dark scaling and a fine, dark fuscous line 
along terminal margin. Alar expanse, 15-18 mm. 

Male genitalia with the elements of transtilla slightly 
curled and weakly united at their apices (not the smooth 
band of interpunctella); apical process of gnathos some- 
what larger; uncus a trifle broader and widened slightly 
at apex. Female genitalia with ductus bursae narrower; 
the dorsal projection at genital opening narrowly tri- 
angulate and pointed; signa more reduced. 

TypE Locauity: Cayuga, Guatemala (typein USNM). 

Foop piant: Unknown. 

Represented in the National Collection by a series of 
12 specimens, both males and females, from the type 
locality (Mar., Apr., May, Sept., Oct.). 


Genera 186-188: Anagasta to Nicetiodes 


[Venational division E. Forewing with 9 veins; 10 from the 
cell; 9 absent; 4 absent; 2 and 3 from the cell. Hind wing with 
discocellular vein curved. Male genitalia with transtilla com- 
plete or its elements greatly enlarged and touching at apices 
(except in Anagasia). Female genitalia with ductus bursae un- 
sclerotized except, occasionally, immediately adjacent to genital 
opening.] 


186. Anagasta, new genus 


Typs or Genus: Ephestia kiihniella Zeller. 
Tongue well developed. Antenna simple and pubes- 


300 


cent in both sexes. Labial palpus upturned, not reach- 
ing vertex in male, extending slightly above in female. 
Maxillary palpus minute, filiform. Forewing smooth; 
9 veins; venation individually variable; vein 2 from 
well before lower outer angle of cell; 4 absent; 3 and 5 
from the lower angle of cell, normally closely approxi- 
mate at base, occasionally connate or shortly stalked; 
6 normally from below upper angle of cell and only 
slightly curved towards base (nearly straight), occa- 
sionally curved upward to the angle and closely approxi- 
mate to 8; 8 and 9 united; 10 from the cell, separated 
from 8 at base; male without costal fold. Hind wing 
with vein 2 from well before lower outer angle of cell; 
3 and 5 normally approximate at base, rarely connate 
or very shortly stalked and, when so, usually on only 
one hind wing of an individual specimen; 7 and § anasto- 
mosed for most or all their lengths beyond cell; cell 
about one-third the length of wing; discocellular vein 
curved. Abdomen of male with two pairs of short, sim- 
ple, rather weak, dorsal hair tufts on eighth segment. 

Male genitalia with apical projection of gnathos a 
very small, shortly bifurcate hook. Transtilla incom- 
plete, its elements long, slender, and widely separated 
at their apices. Uncus long and moderately broad 
(longer than tegumen), beyond its broad base slightly 
tapering to rounded apical margin. Harpe elongate; 
costa produced at apex into a short, free hook. Aedeagus 
enlarged and ventrally cleft at apex. Vinculum stout, 
but slightly longer than greatest width, tapering slightly 
to broad terminal margin. 

Female genitalia with apophyses of ovipositor and 
intersegmental area between ovipositor and eighth seg- 
ment collar very long; ductus bursae membranous 
throughout, strongly spined at junction with bursa; 
genital opening simple and unsclerotized; signa present, 
consisting of from one to four separate, more or less elon- 
gate and angled discs; ductus seminalis from near 
middle of bursa, approximate to signa. 

The generic separation of kuhniella from the other 
stored-product insects of the Ephestia group requires 
some justification in view of the close association of all 
of them in habits, distribution, and indoor association; 
but in any nice definition of E/phestia, kihniella fits 
very badly. On female genitalia it could squeeze into 
Ephestia, even though its long extruded ovipositor sets 
it apart; but on male characters it differs radically. It 
has no trace of a costal fold, while this structure is 
strongly developed in all the true H’phestia. Its trans- 
tilla is not only incomplete but the elements are slender 
and no wise enlarged, connected, or even approximate 
at their apices, quite the opposite of the development of 
that organ in Hphestia. Also its venation is erratic, 
even for a phycitid. Veins 6 and 8 of forewing are 
parallel from a short distance beyond cell as in H’phestia 
as defined in this paper; but the condition of 3 and 5 of 
fore and hind wings is variable to a degree not found in 
the true Ephestias. In addition to its adult characters 
kithniella has the dorsum of the pupal thorax rugose. 
In Ephestia the thorax of the pupa is smooth. Alto- 
gether kiihniella is an aberrant species, probably of a 


/UNITED STATES NATIONAL MUSEUM BULLETIN 207 


different faunal origin from some, at least, of the 
stored-product Ephestia species. Both they and kih- 
mella are obviously of Old World origin. From all 
available evidence the true home of kiihniella seems to 
be the Mediterranean region, probably Asia Minor, as 
suggested in 1930 by both Lebedev and Klemm. 


622. Anagasta kiihniella (Zeller) 
Figures 626, 1124 


Ephestia kihniella Zeller, Stettiner, Ent. Zeit., vol. 40, p. 466, 
1879.—Snellen, Tijds. voor Ent., vol. 28, p. 237, 1885.— 
Barrett, Ent. Monthly Mag., vol. 23, p. 255, 1887.—Klein, 
Proc. Ent. Soc. London, p. lii, 1887.—Omerod, Insect Life, 
vol. 1, p. 314, 1889.—Fletcher, Ent. Soc. Ontario, Twentieth 
Ann. Rept., p. 95, 1889. Insect Life, vol 2, p. 187, 1889; 
Canadian Ent., vol. 22, p. 41, 1890.—Hulst, Phycitidae of 
N. Amer., p. 198, 1890; U. S. Nat. Mus. Bull. 52, p. 435, 
1903.—Riley, Insect Life, vol. 5, p. 276, 1892.—Danysz, 
Mém. de Lab. Parasit. yég. Bourse de Commerce, vol. 1, 
pp. viii-58, 1893.—Howard, Insect Life, vol. 7, p. 416, 
1895.—Johnson, in Forbes, Nineteenth report of the State 
Entomologist on the noxious and beneficial insects of the 
State of Illinois, 65 pp., 1895-1896.—Druce, Biologia-Cen- 
trali Americana, Lepidoptera, Heterocera, vol. 2, p. 286, 
1896.—Fuller, Agr. Gaz. New South Wales, vol. 7, pp. 444—453, 
1896.—Lounsbury, Ent. News, vol. 10, p. 291, 1899.— 
Hampson in Ragonot, Monograph, pt. 2, p. 279, 1901.— 
Carpenter, Econ. Proc. Roy. Dublin Soc., vol. 1, p. 209, 
1903.—Barrett, Lepidoptera of the British Islands, vol. 10, 
p. 54, 1905.—Spuler, Die Schmetterlinge Europas, vol. 2, 
p. 201, 1910.—Mosher, Bull. Illinois State Lab. Nat. Hist., 
vol. 12, p. 24, fig. 74, 1916.—Durant and Beveridge, Journ. 
Roy. Army Med. Corps., vol. 20, pp. 615-634, 1913; reprint 
with notes, Brit. Mus. (Nat. Hist.), London, 1918.—Burk- 
hardt, Zeitschr. Angew. Ent., Berlin, vol. 6, pp. 25-60, 
1919 (biology).—Whiting, Journ. Exp. Zool., vol. 28, pp. 
413-441 (genetics).—Forbes, Cornell Mem. 68, p. 635, 
1923.—Curran, Sci. Agr., vol. 6, p. 385, 1926.—Hering, Anz. 
fiir Schadlingsk., vol. 2, p. 139, 1926.—Richardson, Journ. 
Agr. Res., vol. 32, p. 895, 1926.—White, Proc. Ent. Soe. 
Washington, vol. 29, p. 147, 1927.—Candura, Boll. Lab. 
Zool. Portici, vol. 21, p. 149, 1928.—Hudson, Butterflies and 
moths of New Zealand, p. 156, 1928:—Meyrick, Revised 
handbook of British Lepidoptera, p. 889, 1928.—Kiihn and 
Henke, Ges. Wiss. Géttingen, Math. Phys., Abh., new ser., 
vol. 15, 121 pp., 1929 (physiology).—Brindley, Ent. Soc. 
Amer. Ann., vol. 23, p. 740, 1930.—Klemm, Mitt. Ges. 
Vorratsschutz, vol. 6, p. 26, 1930.—Lebedev, Zeitschr. 
Angew. Ent., Berlin, vol. 6, pp. 597—605, 1930.—Noyes, Bull. 
Ent. Res., vol. 21, p. 77, 1930.—Richards and Herford, Ann. 
Appl. Biol., vol. 17, p. 380, 1980.—Clausen, U. 8. Dep. Agr. 
circ. 168, p. 92, 1931 (Japanese record).—Richards and 
Thomson, Trans. Ent. Soc. London, vol. 80, p. 177, 1932.— 
Norris, Proc. Zool. Soc. London, pp. 597-611, 1932; pp. 
903-934, 1933; pp. 333-360, 1934.—Dickins, Trans. Ent. 
Soc. London, vol. 85, p. 333, 1986.—Busnel, Rev. Path. 
Veg. et Ent. Agr. France, pp. 187-162, Apr. 1937 (arval 
anatomy and physiology).—Lehmensick and Liebers, Zeits. 
angew. Ent., Berlin, vol. 24, p. 442, 1937 (egg).—Pierce and 
Metcalf, Genitalia of the British Pyrales, p. 7, 1938.—Mc- 
Dunnough, Check list, No. 6399, 1939.—Corbet and Tams, 
Entomologist, vol. 78, p. 87, 1945. 

Ephestia fuscofasciella Ragonot, N. Amer. Phycitidae, p. 17, 
1887.—Hulst, Phycitidae of N. Amer., p. 198, 1890.— 
Hampson, in Ragonot, Monograph, pt. 2, p. 278, 1901. 

Ephestia gitonella Druce, Biologia Centrali-Americana, Lepi- 
doptera, Heterocera, vol. 2, p. 286, 1896. 

Ephestia sericaria Authors (not Scott), Ragonot, Bull. Soc. Ent. 
France, vol. 61, p. cclxxiv, 1892.—Richards and Thomson, 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 301 


Trans. Ent. Soc. London, vol. 80, pp. 171, 177, 1932.— 
Corbet and Tams, Proc. Ent. Soc. London (ser. B), vol. 113, 
p. 68, 1943; Entomologist, vol. 76, p. 15, 1943.—Hinton, 
Bull. Ent. Res., vol. 34, p. 195, 1943. 

Forewing gray, more or less dusted with black and 
with blackish markings; antemedial line grayish white 
(this pale shade on some specimens extending over the 
remainder of the basal area), slightly oblique and 
irregularly dentate, bordered outwardly by a black 
shade varying from a narrow line to a broad suffu- 
sion; subterminal line pale gray, obscure on most 
specimens, variable and irregularly dentate, usually 
with a dark inner border; veins in outer area of wing 
somewhat darkly outlined; discal dots at end of cell 
pronounced, sometimes confluent along discocellular 
vein but usually well separated and the lower dot 
elongated slightly along veins 3 and 5; a row of faint 
dark dots along termen. Hind wings translucent, 
whitish, the veins and terminal margin pale brownish 
fuscous. Alar expanse, 18-28 mm. 

Genitalic characters as given for the genus. The 
male genitalia are remarkably uniform and exhibit little 
individual variation, considering the variability of the 
species otherwise. The female genitalia are variable 
(see fig. 1124) in the number and shape of the signa. 

Typr Locauities: Germany (kihniella, in BM); 
Durango, Mexico (gitonella, in BM); Texas (fusco- 
Jfasciella, in Paris Mus.). 

Foon: Flour, grains, nuts, dried fruits, chocolate, 
seeds, biscuits, cakes, jellies, candy, and a host of other 
dried and stored vegetable products. The favored 
foods are wheat and other grain flours. Other records, 
unless based upon reared adults, should be received 
with caution; for the larvae are very difficult to dis- 
tinguish from those of a number of other phycitids 
feeding on dried vegetable matter, and records based 
upon larval identifications are as apt to be incorrect as 
not. The species is of major economic importance only 
as a flour and milled-grain feeder and is often a serious 
pest in flour mills. It is apparently an indoor insect 
and requires a high temperature throughout the year 
for maximum development. 

DistrisutTion: Nearly cosmopolitan. 


“The Mediterranean flour moth” has probably as 
extensive a literature as any other lepidopteron. Only 
the more important systematic, biological, and physio- 
logical references are given above. For further refer- 
ences the papers by Noyes (1931) and Richards and 
Thomson (1932), the ‘Index of American Economic 
Entomology,” “Zoological Record,” and ‘‘Review of 
Applied Entomology’’ should be consulted. 

Considering its variability kiihniella has remarkably 
little synonymy except for numerous misspellings of its 
specific name. A list of these will be found in the 
Richards and Thomson paper (1932). Most modern 
authors use the equivalent spelling kuehniella in place 
of kiihniella; but as the latter was the original ortho- 
graphic form and the species was named in honor of 
Professor Kiihn of Halle we are obliged by the inter- 
national rules to follow it. 


The name fuscofasciella Ragonot applies to a color 
form with broad blackish borders to the transverse 
lines and a contrasted pale area of forewing. Such 
examples occur rather frequently in our Southwestern 
States. Unless venation is noted, it is easy to confuse 
kithniella with large examples of Vitula edmandsae. 
Pale specimens of the two species are strikingly similar 
in habitus. 

The original specimens before Zeller when he described 
kithniella are in the British Museum but, according to 
Clarke and Tams, no one specimen bears a type desig- 
nation so I am here designating a male from the original 
series as lectotype. This specimen bears three labels, 
the first a small white one with the inscription, 
“excl. 4/7/77’; a second label. inscribed in Zeller’s 
handwriting “Zphestia kiihnii Z,’’ “MSS.Z”; and a 
third label with the inscription ‘‘Zell. Coll. 1884.’”’ Tams 
concurs in this selection. 


187 Genus Ephestia Guénée 


Ephestia Guénée, Europaeorum Microlepidopterorum index 
methodicus . . . p. 81, 1845; Ann. Soc. Ent. France, ser. 2, 
vol. 3, p. 319, 1845.—Zeller, Isis von Oken, 1848, pp. 585, 
592.—Herrich-Schaffer, Systematische Bearbeitung der 
Schmetterlinge von Europa, vol. 4, p. 110, 1849.—Heine- 
mann, Schmetterlinge Deutschlands und der Schweiz, Abt. 
2, vol. 1, pt. 2, p. 201, 1865.—Meyrick, Proc. Linn. Soc. 
New South Wales, vol. 3, p. 215, 1878; vol. 4, p. 234, 1879; 
vol. 7, p. 160, 1882; Revised handbook of British Lepidop- 
tera, p. 387, 1928.—Hulst, Phycitidae of N. Amer., p. 197, 
1890; U. S. Nat. Mus. Bull. 52, p. 434, 1903.—Ragonot, 
Monograph, pt. 2, p. xiii, 1901.—Hampson, in Ragonot, 
Monograph, pt. 2, p. 271, 1901.—Forbes, Cornell Mem. 68, 
p. 634, 1923.—Richards and Thomson, Trans. Ent. Soc. 
London, vol. 80, p. 171, 1932.—Bisset in Pierce and Metcalfe, 
Genitalia of the British Pyrales, p. 58, 1938. (Type of 
genus: Tinea elutella Hiibner.) 

Hyphantidium Scott, Proc. Zool. Soc. London, vol. 27, p. 207, 
1859, (Type of genus: Hyphantidium sericariwm Scott.) 

Cadra Walker, List, vol. 30, p. 961, 1864 (Type of genus: Cadra 
defectella Walker.) 


Tongue well developed. Antenna simple and pubes- 
cent in both sexes. Labial palpus upturned, reaching 
slightly above vertex. Maxillary palpus minute, fili- 
form. Forewing smooth; 9 veins; vein 2 from before 
but rather near lower outer angle of cell; 3 from outer 
angle; 4 absent; 5 more or less approximate to 3 at base; 
6 from below upper angle of cell, straight or slightly 
curved towards base (from a slight distance beyond base 
running parallel with 8) ; 9 absent; 10 from the cell; male 
with a strong costal fold enclosing scale tuft. Hind 
wing with vein 2 from well before lower outer angle of 
cell; 3 and 5 approximate at base or short stalked 
(elutella); 7 and 8 anastomosed for most or all of their 
lengths beyond cell; cell one-half or slightly less than 
one-half the length of wing; discocellular vein curved. 
Abdomen of male with compound dorsal tufts. 

Male genitalia with gnathos terminating in a pair of 
well-spaced prongs. Transtilla complete or, if elements 
slightly separated, their apices broadly developed, if 
complete with flaring lateral projections. Uncus short, 
broad; its terminal margin bluntly rounded. Aedeagus 
not appreciably enlarged at apex; penis armed with a 


302 


sclerotized band or a row) of sclerotized wrinklings. 
Vinculum stout, longer than broad, tapering slightly to 
blunt terminal margin. 

Female genitalia with apophyses of ovipositor and 
intersegmental area between ovipositor and eighth seg- 
ment collar short; ductus bursae strongly spined towards 
bursa or bearing a longitudinal row of sclerotized ridges; 
signa present, consisting of a row of narrow sclerotized 
discs; ductus seminalis from bursa approximate to signa. 

Ephestia.as here defined is an Old World genus. Even 
with kihniella and the formerly included American 
species removed, it remains somewhat of a heteroge- 
neous group. I suspect that the type species (elutella) 
has a different faunal origin than cautella, figulilella, 
callideila, and affatella, which, because of their similar 
habits, present distribution and genitalic structures are 
associated with it; but until the Old World species now 
in Ephestia can be thoroughly studied and the incon- 
gruous ones given a better generic placement we shall 
have to keep the more-or less cosmopolitan species 
together. However, elutella has a different hind wing 
venation and a different type of harpe from the others 
and I believe will eventually be generically distin- 
guished.% Three species are represented in the Amer- 
icas, all apparently imports from the Old World. 
They separate into groups as follows: 

Hind wing with veins 3 and 5:stalked; costa of harpe smooth. 

Hind wing with veins 3 and 5 approximate at base; harpe 
with digitate projection from middle or near middle 
of costa. 

Color and markings are of little help in distinguishing 
the species. There is so much individual variation and 
so little specific difference, even between typical speci- 
mens, that superficial habitus can not be trusted. 
Genitalia on the other hand do readily identify and 
distinguish the species. Their specific characters are 
several, obvious, and consistent. These organs should 
be examined in every instance when specific identifica- 
tion is of any importance. 


Genus E’phestia, Species 623: E. elutella 
[Hind wing with veins 3 and 5 stalked; costa of harpe smooth.] 


623. Ephestia elutella (Hiibner) 
Ficures 126, 628, 1122 


Tinea elutella Hubner, Sammlung europaischer Schmetterlinge, 
Lepidoptera 8, Tinea 5, pl. 24, fig. 163, 1796. 

Phycis elutea Haworth, Lepidoptera Britannica, p. 496, 1811 
(amended spelling for elutelia). 

Phycis semirufa Haworth, Lepidoptera Britannica, p. 496, 1811. 

Phycis rufa Haworth, Lepidoptera Britannica, p. 497, 1811. 

Phycis elutella (Hiibner) Zincken, in Germar and Zincken, Mag. 
der Ent., vol. 3, p. 175, 1818.—Treitschke, Die Schmetter- 
linge von Kuropa, vol. 9, p. 194, 1832.—Duponchel, Histoire 
naturelle des lépidoptéres, ou papillons de France, vol. 10, 
p. 204, 1836. 


8 Ephestia vitivora Filipjev (Deuts, Ent. Zeit. “Iris,” Dresden, 
vol. 45, p. 70, 1931) from Transcaucasia is distinct from, but 
obviously very close to, and in the event of any further division 
of Ephestia, congeneric with elutella—suggesting a western Asiatic 
origin for the latter. 


( 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Phycita semirufa (Haworth) Stephens, Illustrations of British 
entomology, Haustellata, vol. 4, p. 305, 1834.— Wood, Index 
entomologicus, p. 210, pl. 46, fig. 1457, 1839.—Westwood 
and Humphreys, British moths and their transformations, 
vol. 2, p. 229, 1845. 

Phycita elutella (Hibner) Stephens, Mlustrations-of British ento- 
mology, Haustellata, vol. 4, p. 804, 1884—Wood, Index 
entomologicus, p. 210, pl. 46, fig. 1454, 1839.—Westwood 
and Humphreys, British moths and their transformations, 
vol. 2, p. 229, 1845. 

Myelois elutella (Hiibner) Zeller, Isis von Oken, 1839, pp. 176, 
343. 

Ephestia elutella (Hiibner) Guénée, Europaeorum Microlepidop- 
terorum index methodicus..., p. 81, 1845.—Herrich- 

' Schaffer, Systematische Bearbeitung der Schmetterlinge von 
Europa, vol. 4, p. 110, 1849.—Stainton, Manual of British 
butterflies and moths, vol. 2, p. 168, 1859.—Heinemann, Die 
Schmetterlinge Deutschlands und der Schweiz, Abt. 2, vol. 1, 
pt. 2, p. 201, 1865.—Zeller, Verh. zool.-bot. Ges. Wien, vol. 
25, p. 338, 1876.—Hulst, Phycitidae of N. Amer., p. 200, 
1900; U.S. Nat. Mus. Bull. 52, p. 435, 1903.—Hampson, in 
Ragonot, Monograph, pt. 2, p. 300, 1901.—Forbes, Cornell 
Mem. 68, p. 635, 1923.—Curran, Sci. Agr.; vol. 6, p. 386, 
1926.—Meyrick, Revised handbook of British Lepidoptera, 
p. 388, 1928.—Munro and Thomson, Empire Marketing 
Board, No. 24, London, H. M. Stationery Office, p. 22, 
1929.—Noyes, Bull. Ent. Res., p. 80, 1930.—Richards and 
Herford, Ann. Appl. Biol., vol. 17, p. 380, 1930.—Back and 
Reed, Journ. Econ. Ent., vol. 23, p. 1004, 1930.—Richards 
and Thomson, Trans. Ent. Soc. London, vol. 80, p. 185, 
1932.—Bovingdon, Empire Marketing Board, No. 67, pp. 1— 
88, 1933.—Norris, Proc. Zool. Soc. London, 1934, pp. 333— 
360, 1934.—Dickins, Trans. Ent. Soc. London, vol. 85, 
p. 348, 1936.—Lehmensick and Liebers, Zeitschr. angew. 
Ent., Berlin, vol. 24, p. 442, 1937.—Pierce and Metcalfe, 
Genitalia of the British Pyrales, p. 6, 1938.—McDunnough, 
Check list, No. 6401, 1939.—Hinton, Bull. Ent. Res., vol. 
34, p. 196, 1943.—Corbet and Tams, Proc. Zool. Soc. Lon- 
don, vol. 113, ser. B, p. 64, 1943. 

Hypantidium sericariwm Scott, Proc. Zool. Soc. London, vol. 27, 
p. 207, 1859.—Corbet and Tams, Entomologist, vol. 78, 
p. 87, 1945 (establish synonymy with elutella). 

Hphestia semirufa (Haworth) Staudinger, Horae Soc. Ent. Ros- 
sicae, vol. 15, p. 229, 1879. 


Ephestia roxburghii Gregson, Entomologist, vol. 6, p. 318, 1873.— 
Ragonot, Monograph, pt. 2, p. 301, 1901.—Richards and 
Thomson, Trans. Ent. Soc. Lond., vol. 80, p. 187, 1932. 


Ephestia elutella unicolorella Staudinger, Horae Soc. Ent. Ros- 
sicae, vol. 15, p. 228, 1879. 


Ephestia unicolorella Staudinger, Horae Soc. Ent. Rossicae, vol. 
16, p. 89, 1881.—Ragonot and Hampson, in Ragonot, Mono- 
graph, pt.2, p. 229, 1901.—Richards and Thomson, Trans. 
Ent. Soc. London, vol. 80, p. 187, 1932. 


Ephestia amarella Dyar, Proc. U. 8. Nat. Mus., vol. 27, p. 921, 
1904. 

Forewing dark brown to pale gray, sometimes with a 
paler shade between lower fold and inner margin; color 
variable; transverse lines usually distinguishable; ante- 
medial line obscure, oblique, bordered outwardly by a 
narrow dark shade, the latter approaching middle of 
wing; subterminal line somewhat sinuate, faintly bor- 
dered on both sides by dark lines. Hind wing smoky 
white to pale fuscous. Alar expanse, 13-18 mm. 

Male genitalia with prongs of apical process of 
gnathos forming a V, more narrowly spaced than in 
other species of the genus. Harpe without projections 
from costa. Female genitalia with ductus bursae rather 
coarsely spined for about half its length towards bursa; 
bursa evenly and minutely spined. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


TypE Locaitizs: Germany (elutella, type lost); Eng- 
land (semirufa, type lost; rufa and roxburghii, in BM); 
Malatia, Asia Minor (unicolorella, in Mus. Univ. Ber- 
lin); Australia (sericarium, in BM); Kalso, Kootenai 
District, British Columbia (amarella), in USNM). 


Foon: Dried fruits, seeds, nuts, chocolate, cocoa, 
tobacco, and a number of other dried vegetable prod- 
ucts. Feeding records based on larval identifications 
should be accepted with extreme caution; for the larvae 
of elutella, are often impossible to distinguish from 
those of cautella, and have been frequently confused 
with them. The so-called characters given in literature 
for separation of the larvae of the two species are not 
constant and should not be depended upon for specific 
identification. In both Europe and the United States 
elutella seems to breed out-of-doors to some extent but 
has been noticed as a pest only in storage houses, in 
England chiefly on stored cocoa and in America on 
tobacco. In our economic literature it is known as 
“the tobacco moth.” Aside from its depredations in 
tobacco warehouses it appears to be only a minor and 
occasional pest on other stored products in the United 
States. 


Distripution: Practically cosmopolitan, but appar- 
ently less numerous and not so generally distributed as 
either Plodia interpunctella or E’phestia cautella. 


The moths are easily separable from the other two 
species found here (cautella and figulilella) and from 
other truly congeneric European species by the short 
stalking of veins 3 and 5 of hind wing. In other true 
Ephestia these veins are always approximate at base. 
They should be examined in every instance where 
specific identification is of any importance. 


The above synonymy gives only the more important 
references. Some Old World references cited by 
Richards and Thomson (1932) and Hulst (1903) have 
been omitted and, except for a few essential articles, 
no attempt has been made to cover the economic 
literature, which is both extensive and repetitious. 


Genus Ephestia, Species 624 and 625: E. cautella 
and #. figulilella 


[Hind wing with veins 2 and 5 approximate at base; harpe with 
digitate projection from middle or near middle of costa.] 


624, Ephestia cautella (Walker) 
Fieurns 125, 629, 1121 


Pempelia cautella Walker, List, vol. 27, p. 73, 1863. 

Cadra defectella Walker, List, vol. 30, p. 962, 1864. 

Nephopteryx desuetella Walker, List, vol. 35, p. 1719, 1866. 

Ephestia cahiritella Zeller, Stettiner, Ent. Zeit., vol. 28, p. 384, 
1867.—South, Entomologist, vol. 23, p. 304, 1890.—Van 
Deventer, Tijdschr. voor Ent., vol. 46, p. 80, 1904.— 
Barrett, Lepidoptera of the British Islands, vol. 10, p. 56, 
1905. 

Ephestia passulella Barrett, Ent. Monthly Mag., vol. 11, p. 271, 
1875.—Ragonot, Ent. Monthly Mag., vol. 22, p. 24, 1885. 

Salebria cautella (Walker) Cotes and Swinhoe, Catalogue of the 
moths of India, pt. 6, p. 675, 1889. 


303 


Ephestia cautella (Walker) Hampson, Moths, vol. 4, p."66, 1896, 
in Blanford, Fauna of British India.—Ragonot, Monograph, 
pt. 2, p. 292, 1901.—Hulst, U. S. Nat. Mus. Bull. 52, p. 434, 
1903.—Spuler, Die Schmetterlinge Europas, vol. 2, p. 202, 
1910.—Chittenden, U. 8. Dep. Agr. Bur. Ent. Bull. 104, 
40 pp., 1911.—Dyar, Proc. U. S. Nat. Mus., vol. 47, p.'345, 
1914.— Forbes, Cornell Mem. 68, p. 635, 1923.—Curran, 
Sci. Agr., vol. 6, p. 386, 1926.—Meyrick, Revised Handbook 
of British Lepidoptera, p. 388, 1928.—Shibuya, Journ. 
Faculty Agric. Hokkaido Imperial Univ., Sapporo, vol. 22, 
p. 77, 1298.—Richards and Herford, Ann. Appl. Biol., vol. 
17, p. 380, 1930.—Noyes, Bull. Ent. Res., p. 80, 1930.— 
Simmons, Reed, and McGregor, U. 8. Dep. Agr. Cir. 157, 
p. 36, 1931.—Keifer, Monthly Bull. Dep. Agr. California, 
vol. 20, p. 619, 1931.—Richards and Thomson, Trans. Ent. 
Soe. London, vol. 80, p. 197, 1932.—Bovingdon, Empire 
Marketing Board, No. 67, pp. 1-88, 1933.—Norris, Proc. 
Zool. Soc. London, 1934, pp. 333-360, 1934.—Dickins, 
Trans. Ent. Soc. London, vol. 85, p. 342, 1936.—Lehmensick 
and Liebers, Zeitschr. angew. Ent., Berlin, vol 24, p. 443, 
1937.—Pierce and Metcalfe, Genitalia of the British Pyrales, 
p. 7, 1938.—McDunnough, Check list, No. 6403, 1939.— 
Hinton, Bull. Ent. Res., vol. 34, p. 193, 1943.—Corbet 
and Tams, Proc. Zool. Soc. London, vol. 113, ser. B, p. 64, 
1943. 

Cryptoblabes formosella Wileman and South (<), Entomologist, 
vol. 51, p. 219, 1918.—Shibuya, Journ. Faculty Agric. 
Hokkaido Imperial Univ., Sapporo, vol. 22, pp. 17, 88, 1928. 


Color and markings similar to those of elutella and as 
variable, expecially on the females. On the average 
the antemedial line is less oblique in both cautella and 
Jigulilella than in elutella, but this difference does not 
hold for all specimens. £. cavutella is distinguished by 
its genitalia, both male and female. They are quite 
distinct from those of any other phycitid. 

Alar expanse, 11-18 mm. 

Male genitalia with prongs of apical process of 
gnathos forming a broad U, widely spaced. Harpe 
with a short, digitate projection from near middle of 
costa. Transtilla with a broad fusion at middle and 
prominent, round, flattened, lateral lobes. 

Female genitalia with ductus bursae bearing a longi- 
tudinal row of strongly sclerotized and pigmented 
ridges, the latter forming 9, narrow, fluted band some- 
what over half as long as the ductus. Bursa copulatrix 
rather coarsely spined over about half its surface. 

Typr tocauitiges: Ceylon, India (cautella and de- 
fectella, in BM); Moreton Bay, Queensland, Australia 
(desuetella, in BM); Cairo, Egypt (cahiritelia, in BM); 
England (passulella, in BM); Takow, Formosa (for- 
mosella, in BM). 

Foon: All kinds of dried, stored vegetable products. 

Distrizution: Cosmopolitan. 

This species is as abundant, and probably as destruc- 
tive to dry fruits, grains, nuts, and other stored vege- 
table products, as Plodia interpunctella. Surprisingly 
enough it has fewer economic references than either 
elutella or figulella though it appears to be more widely 
distributed and more of a general nuisance than either 
of them. In the list of common names approved by 
the American Association of Economic Entomologists 
it is known as the “almond moth.” In economic litera- 
ture generally it has also been referred to as the ‘‘dried 
currant moth” and the “fig moth.” 


304 


625. Ephestia figulilella Gregson 
Fieures 630, 1123 


Ephestia figulilella Gregson, Entomologist, vol. 5, p. 385, 1871.— 
Ragonot, Ent. Monthly Mag., vol. 22, p. 25, 1885.—Hampson 
and Ragonot, in Ragonot, Monograph, pt. 2, p. 282, 1901.— 
Hulst, U. S. Nat. Mus. Bull. 52, p. 435, 1903.—Caradja, 
Deutsche Ent. Zeitschr., ‘‘Iris,’’ vol. 24, p. 210, 1910.—Spuler, 
Die Schmetterlinge Europas, vol. 2, p. 202, 1910.—Turati and 
Zanon, Atti Soc. Italiana Sci. nat., vol. 61, p. 164, 1922.— 
Keifer, Monthly Bull. Dep. Agr. California, vol. 20, p. 619, 
1931.—Richards and Thomson, Trans. Ent. Soc. London, 
vol. 80, p. 194, 1982.—Donohoe and Barnes, Journ. Econ. 
Ent., vol. 27, pp. 1070, 1075, 1934.—Pierce and Metcalfe, 
Genitalia of the British Pyrales, p. 6, 19838.—McDunnough, 
Check list, No. 6402, 1939.—Hinton, Bull. Ent. Res., vol. 
34, p. 194, 1943.—Corbet and Tams, Proc. Zool. Soc. Lon- 
don, vol. 113, ser. B, p. 68, 1948.—Donohoe et al., U. S. 
Dep. Agr. Techn. Bull. 994, pp. 1-23, 1949. 

Ephestia ficulella Barrett, Ent. Monthly Mag., vol. 11, p. 271, 
1875 (emended spelling for jigulilella)—Meyrick, Proc. 
Linn. Soe. New South Wales, vol. 4, p. 244, 1880.—Chittenden, 
U. S. Dep. Agr. Div. Ent. Bull. 8 (new ser.), p. 10, 1897.— 
Meyrick, Revised hand book of British Lepidoptera, p. 388, 
1928. 

Ephestia millerit Zeller, Verh. zool.-bot. Ges. Wien., vol. 25, p. 
339, 1876.—Druce, Biologia Centrali Americana, Lepidop- 
tera, Heterocera, vol. 2, p. 287, 1890. 

Ephestia figuliella Forbes, Cornell Mem. 68, p. 635, 1923 (mis- 
spelling). 

Ephestia figulella Curran, Sci. Agr., vol. 6, p. 386, 1926 (mis- 
spelling). 

Ephestia venosella Turati, Atti Soc. Italiana Sci. Nat., vol. 65, 
p. 58, 1926. 

Ephestia ernestinella Turati, Atti Soc. Italiana Sci. Nat., vol. 66, 
p. 330, 1927. 


The genitalia are distinctive but there are no con- 
sistent superficial characters for the separation of figu- 
hilella from cautella. 

Alar expanse, 12-17 mm. 

Male genitalia with prongs of apical process of 
gnathos U-shaped with the base of the U flattened and 
the prongs well separated. Harpe with a long digitate 
projection from midcosta. Transtilla with its elements 
broadened and touching (but not fusing) at their apices. 

Female genitalia with ductus bursae armed with a 
spiral of short, broadly based spines; a cluster of similar 
spines at terminal end of bursa copulatrix; a pair of 
fan-shaped scale tufts from intersegmental area ad- 
jacent to genital opening; signa consisting of a row of 
a half dozen elongate, narrow discs. 


Typ tocatitizs: Liverpool, England (jfigulilella, in 
BM); Central America (milleri, in BM); Bengasi, 
Cyrenaica (venosella, [?] Turati Coll.); Giarabub, 
Cyrenaica (ernestinella, [?] Turati Coll.). 

Foop: Dried fruits, nuts, seeds, meal, beans, etc. 

DistrrBution: Europe, Asia, North and West Africa, 
Hawaii, Australia, North and South America. In the 
United States it seems to have established itself only in 
California where it is a minor pest of dried raisins, 
though it has been intercepted at various ports of entry 
in stored products. It is apparently less widely dis- 
tributed and of considerably less economic importance 
than either cautella or elutella. In our economic litera- 
ture it is known as the “‘raisin moth,” and has a growing 
list of economic references, for which the Review of 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Applied Entomology should be consulted. Some Old 
World references cited by Richards and Thomson (1932) 
have been omitted from the foregoing synonymy. 


188. Genus Nicetiodes Schaus 


Nicetiodes Schaus, Zoologica (Contr. New York Zool. Soc.), 
vol. 5, No. 2, p. 48, 1923 (Type of genus: Nicetiodes 
apianella Schaus). 

Tongue well developed. Antenna pubescent; shaft 
of male dilated and with a shallow, ventral sinus 
towards base; shaft of female simple. Labial palpus 
slender, upturned, reaching slightly above vertex; third 
segment acuminate, about half the length of second. 
Maxillary palpus rather broadly dilated with scales on’ 
female, less so on male. Forewing of male with some 
raised scales alone upper margin of cell towards base 
(possibly a specific character), of female smooth; 9 
veins; vein 2 from before but near lower outer angle of 
cell; 3 and 5 from the angle, closely approximate or 
connate at base; 4 absent; 6 from below upper angle, 
straight; 8 and 9 united; 10 from the cell, separate from 
8 at base; male with a strong costal fold enclosing hair 
tuft. Hind wing with vein 2 from very close to lower 
outer angle of cell; 3 and 5 stalked for less than half 
their lengths; 7 and 8 anastomosed for most of their 
lengths beyond cell; cell about half the length of wing; 
discocellular vein curved. Abdomen of male with 
compound dorsal tufts. 

Male genitalia with gnathos terminating in an en- 
larged, angulate knob. ‘Transtilla complete, a narrow, 
low-arched bridge with slight central projection. Uncus 
narrow stemmed, abruptly expanding and flatly bilobed 
at extremity; a short, broad scaphium projecting from 
its base. Harpe with sacculus slightly produced at 
extremity, otherwise simple. Anellus tubular with at- 
tached ventral shield bearing short lateral lobes. 
Aedeagus long, rather stout, flaring and sharply angled 
at apex; penis with a few weakly sclerotized wrinklings, 
otherwise unarmed. Vinculum triangulate, longer than 
broad. 

Female genitalia with ductus bursae scobinate 
towards bursa and narrowly sclerotized at genital open- 
ing; signa present, consisting of a row of small sclero- 
tized discs; ductus seminalis from bursa approximate to 
signa. 

A distinct genus at once distinguished by its peculiar 
uncus and gnathos and projecting sacculus. Contains 
but one described species. 


626. Nicetiodes apianella Schaus 
Ficures 127, 637, 1125 


Nicetiodes apianella Schaus, Zoologica, vol. 5, No. 2, p. 48, 1923. 


Forewing blackish fuscous with a few dull whitish 
scales at base and a shading of them surrounding dis- 
cocellular vein and in area beyond subterminal line; 
transverse lines white, well separated, the antemedial 
line narrow, straight, and nearly vertical; subterminal 
line faint, irregularly and shortly dentate, rather near 
termen; discal dots obscure. Hind wing smoky white; 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


the veins slightly darkened and a similar narrow dark 
shade along terminal margin. Alar expanse, 13 mm. 

Genitalic characters as given for the genus; the 
tubular part of anellus partially scobinate. 

Type Locatity: Conway Bay, Indefatigable, South 
Seymore, Tower Island, Galipagos (type in USNM). 

Foop prant: Unknown. 

Known only from the male type and female paratype 
from the type locality. A rather striking species con- 
spicuous because of its blackish forewings with con- 
trasted white antemedial line. 


Genera 189-191: Varneria to Erelieva 


[Venational division F. Forewing with 9 veins; 2 from the cell; 
3 and 5 stalked; 4 absent; 9 absent; 10 from the cell.] 


189. Genus Varneria Dyar 


Varneria Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 114, 
1904.—Forbes, Cornell Mem. 68, p. 639, 1923. (Type of 
genus: Varneria postremella Dyar.) 


Tongue well developed. Antenna weakly pubescent; 
shaft simple, sometimes showing a slight bend at base 
(but no sinus and the bend as obvious in female as 
male). Labial palpus oblique, slender, laterally flat- 
tened, reaching as high as vertex; third segment slightly 
less than half as long as second. Maxillary palpus 
filiform. Forewing smooth; 9 veins; vein 2 from lower 
outer angle of cell or from very close to the angle, con- 
nate with or approximate to the stalk of 3-5 at base; 
4 absent; 3 and 5 stalked; 6 from below upper angle of 
cell, straight; 8 and 9 united (9 absent) ; 10 from the cell, 
separated from 8 at base; male with strong costal fold 
containing projecting hair tuft. Hind wing with vein 
2 from close to lower outer angle of cell; 3 and 5 long 
stalked; 7 and 8 completely anastomosed beyond cell; 
cell one-half the length of wing; discocellular vein 
curved. Abdomen of male with rather weak, com- 
pound, dorsolateral scale tufts. 


Male genitalia with apical process of gnathos U- 
shaped, the prongs rather short and moderately stout. 
Transtilla complete, a narrow curved band. Harpe 
with projecting spur from base of costa, otherwise 
simple. Anellus a simple shield. Aedeagus straight, 
simple, not appreciably tapering; penis bearing a flat, 
weakly sclerotized plate, otherwise unarmed. Vincu- 
lum subtriangulate, stout, no longer or but slightly 
longer than greatest width. 


Female genitalia with ductus bursae sclerotized and 
flattened for more than half its length from genital 
opening and with sclerotized, projecting, dorsal shield 
behind genital opening; bursa copulatrix with or with- 
out signum, latter when present a small flat plate with 
a few toothlike projections along one edge; ductus 
seminalis from near middle of bursa. 

Four species are here recognized as belonging to the 
genus; but two of these are of doubtful specific status. 
The absence of males in two of the species and the 
scarcity of specimens in those known only from females 


305 


makes the evaluation of characters for specific separa- 
tion very difficult. All of the species have a superficial 
character in common in the vinous red ground color of 
the forewings. 


627. Varneria postremella Dyar 
Ficures 88, 631, 1110 
Varneria postremella Dyar, Proc. Ent. Soc. Washington, vol. 6, 
p. 115, 1904.—Ely, Proc. Ent. Soc. Washington, vol. 12, 
p. 203, 1910.—Forbes, Cornell Mem. 68, p. 639, 1923.— 
McDunnough, Check list, No. 6406, 1939. 

Forewing vinous red; basal half of costa overlaid with 
blackish scales; a line of blackish scales along discal 
fold, spreading somewhat between veins 3 and 6, this 
blackish dusting variable and sparse in some specimens; 
no transverse lines or discal spots distinguishable. 
Hind wing smoky fuscous, costal and terminal margins 
darker. Alar expanse, 9-10.5 mm. 

Male genitalia with costal process from harpe directed 
forward, hooked at apex; lower margin of harpe incurved 
near apex. Terminal margin of vinculum angulate. 
Female genitalia with signum present; projecting shield 
behind genital opening subtriangulate, sides incurved, 
apex truncate. 

Type LocaLity: Kentucky (type in USNM). 

Foop piant: Unknown. 

Disrripution: Connecticut, East River (July) ; Mary- 
land, Hyattsville (Aug.), Plummers Isl. (July); Vir- 
ginia, Mountain Lake (July); Kentucky (July). 

In postremella vein 2 of forewing is connate with the 
stalk of 3-5. In the remaining species of Varneria it is 
closely approximate. This character, however, might 
be expected to vary in individual specimens if more 
were available. 


628. Varneria nannodes Dyar 
Figure 1111 
Varneria nannodes Dyar, Proc. Ent. Soc. Washington, vol. 47, 
p. 346, 1914. 

This species is represented only by the female type, 
which in color, markings, and genitalia exhibits nothing 
to distinguish it specifically from females of postremella. 
It is probably a synonym of the latter; but in the ab- 
sence of a male of nannodes or any matching examples 
between Virginia and Panamé it were wiser to hold any 
synonymy in abeyance. Alar expanse, 8.5 mm. 

Typr Locatity: Cabima, Panamé (type in USNM). 

Foop piant: Unknown. 


629. Varneria atrifasciella Barnes and McDunnough 
Figure 1112 


Varneria atrifasciella Barnes and McDunnough, Contributions, 
vol. 2, p. 184, 19138.—McDunnough, Check list, No. 6407, 
1939. 

Forewing dark vinous red rather heavily dusted with 
blackish scales; in the paratype faint indications of pale 
(yellowish) antemedial and postmedial lines, these, how- 
ever, not distinguishable in the type and their position 
chiefly indicated by the concentration of blackish scaling 


306 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


between them. Hind wing smoky white, the veins 
slightly darker. Alar expanse, 9-9.5 mm. 

Female genitalia without signum; projecting shield 
behind genital opening large and approximately square; 
an eversible lobe in ventral membrane between oviposi- 
tor and eighth-segment collar. 

TyYpE Locautity: Everglades, Fla. (type in USNM). 

Foop prant: Unknown. 

Represented only by the type and one paratype from 
the type locality (Apr.), both females. 


630. Varneria dubia, new species 
Fiaures 632, 1113 


Similar to atrifasciella except that forewing is less 
abundantly shaded with black and female genitalia show 
slight differences in the shape of the shield at genital 
opening and the eighth-segment collar (compare figs. 
1112 and 1118). Alar expanse, 8.5-11 mm. 

Male genitalia with costal process from harpe directed 
backward, curved, pointed at apex; lower margin of 
harpe evenly convex toward apex. Female genitalia 
with projecting shield behind genital opening triangu- 
late; eighth-segment collar appreciably narrower to- 
wards base on dorsum; otherwise as in atrifasciella. 

Typz Locatity: El Yunque, Luquillo Mts., Puerto 
Rico (type in Cornell Univ. Coll.; paratypes in Cornell 
Univ. and USNM, 61395). 

Foon. prant: Unknown. 

Described from male type and one male and four fe- 
male paratypes from the type locality, Mar. 29 to Apr. 
23, 1930, Cornell University lot 795 sub. 9.and 40. In 
addition to the types there are fragments of a broken 
specimen in the Cornell Collection from Jajome Alta, 
Puerto Rico, June 18, 1930, J. A. Hoffman, collector. 

This species is uncomfortably close to atrifasciella and 
may be nothing more than a race or variety of it, but 
in all the females before me the genitalic differences, 
slight though they be, are consistent, and in the absence 
of any authentic males of aétrifasciella or sufficient fe- 
male examples of any Varneria species to determine the 
extent of individual variation in genitalia it seems safest 
to define dubia as a distinct species. 


190. Genus Eurythmia Ragonot 


Hurythmia Ragonot, N. Amer. Phycitidae, p. 16, 1887.—Hulst, 
Phycitidae of N. Amer., p. 195, 1900.—Hampson, zn Rago- 
not, Monograph, pt. 2, pp. xiii, 266, 1901. (Type of genus: 
Ephestia hospitella Zeller.) 

Tongue well developed. Antenna weakly pubescent, 
simple. Labial palpus upturned, reaching vertex, lat- 
erally flattened and somewhat rough scaled; third seg- 
ment slightly more than half as long as second. Maxil- 
lary palpus filiform. Forewing smooth; 9 veins; vein 2 
from very close to lower outer angle of cell; 3 from the 
angle, shortly stalked with 5; vein 4 absent; 6 from be- 
low but near upper angle, very slightly bent; 8 and 9 
united; 10 from the cell, separated from 8 at base; male 
with short costal fold. Hind wing with 2 from close to 


lower outer angle of cell; 3 and 5 stalked for at least half 


their lengths; 7 and 8 anastomosed for most of their 
lengths beyond cell (free element of 8 very short); cell 
less than half the length of wing; discocellular vein 
curved. Highth abdominal segment of male simple. 

Male genitalia with gnathos terminating in a very 
short, stout, narrowly forked projection. Transtilla 
complete, arched and produced at middle into a narrowly 
forked projection with shorter central spur. Harpe with 
costa sclerotized for half its length, the apex of the 
sclerotized margin projecting as a free spur. Anellus a 
curved shield with short lateral lobes. Aedeagus taper- 
ing sharply from enlarged base to bluntly pointed apex; 
penis armed with an elongate sclerotized spiral band 
covered with short saw-toothed spines. Vineulum tri- 
angulate, longer than greatest width and tapering to a 
blunt point. 

Female genitalia with ductus bursae membranous 
throughout; genital opening simple; bursa copulatrix 
with signa a series of (8 to 6) detached, broad-based, 
short, thornlike spines and narrow-bladed discs situated 
near anterior end of the bursa and a cluster of several 
similar, much smaller spines near junction of bursa and 
ductus bursae; ductus seminalis from bursa in the neigh- 
borhood of the larger signa. 

Eurythmia is distinguished chiefly by the armature of 
the penis, the arrangement of the signa in bursa, and the 
simple eighth abdominal segment of the male. The 
venation is similar to that of Varneria except for the 
shorter stalking of veins 3 and 5 of forewing, normally a 
character of, at most, specific significance. Ragonot 
and Hampson interpret the forewing venation differ- 
ently from that given here; namely, 2-3 united and 4-5 
stalked rather than 4 absent and 3-5 stalked. The end 
result would be the same, but the male genitalia do not 
indicate any affinities to the genera having 2 and 3 
stalked and tending to fuse and do show affinities to 
E’phestiodes (especially the aberrant species noniella and 
stictella) where veins 4 and 5 are stalked and 4 shows & 
tendency to disappear (compare venation of EL. plorella, 
fig. 109). 

Apparently the genus as here defined is confined to 
North America; at least, no tropical species have as 
yet been discovered. Several have been described from 
the United States on trifling differences of color and 
maculation. They exhibit no differences in male geni- 
talia of any significance and no consistent characters in 
the female genitalia. There is great variability in the 
number, arrangement, and shape of the spines and discs 
forming the signa; but this is individual and as great 
between females of a given series as between it and any 
of the other named forms. Pending some information 
on the biology, and more material from intervening 
areas of the distribution, we shall have to keep some of 
the names; but I suspect that they all represent but one 
variable species. Figures 633c-d show extremes of 
variation in the forked central projection from transtilla 
between a small Texas example of hospitella and the type 
of diffusella Ely. Such difference is not of more than 
individual significance as we have both extremes and 
all intergrades in male examples of hospitella. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


631. Eurythmia hospitella (Zeller) 
Frioursrs 87, 633, 1130 


Ephestia hospitella Zeller, Verh. zool.-bot. Ges. Wien, vol. 25, 
p. 338, 1875. 

Eurythmia hospitella (Zeller) Ragonot, N. Amer. Phycitidae, p. 
16, 1887.—Hulst, Phycitidae of N. Amer., p. 196, 1900; 
U. S. Nat. Mus. Bull. 52, p. 486, 1903—Hampson, in 
Ragonot, Monograph, pt. 2, p. 266, 1901.—McDunnough, 
Check list, No. 6385, 1939 (in part). 

Eurythmia spaldingella Dyar, Proc. Ent. Soc. Washington, vol. 7, 
p. 39, 1905.—McDunnough, Check list, No. 6391, 1939, 
(New synonymy.) 

Forewing white more or less dusted with fuscous, the 
concentrated brown dusting appearing more or less 
blackish to the naked eye; antemedial line transverse, 
indicated by a dark outer border which is frequently 
produced into a sharp, outward angle at top of cell; 
subterminal line near and parallel to termen, bordered 
inwardly by a dark border and outwardly by some dark 
dusting; discal dots at end of cell small, blackish, 
separate. Hind wing whitish, subpellucid; the veins 
but faintly darkened; a fuscous line along termen. 
Alar expanse, 10-15 mm. 


Male genitalia as given for the genus: Female geni- 
talia with the larger spines and discs, comprising the 
signa, varying from 4 to 6. The type of hospitella has 
5. One of the paratypes of spaldingella has 4, as in 
yavapaella (fig. 1131). The other female paratype of 
spaldingella has 5, as has a specimen from Winter Park, 
Fla. A female of hospitella in the National Museum 
from Texas (fig. 1180a) shows 6. There is no corre- 
spondence between locality or pattern variation and the 
number of the larger signa. 


TYPE LOCALITIES: Texas (hospitella, in BM); Stock- 
ton, Utah (spaldingella, in USNM). 

Foop prant: Unknown. 

Distrisution: Florida, Winter Park (July); Tezas, 
Bosque County (May, July); Brownsville (May), Kerr- 
ville (July); Arizona, Maricopa County, Fish Creek 
(Sept.), Phoenix (Sept.), Liberty (Sept.); Utah, Stock- 
ton (Aug., Sept.). 

Our series (14 specimens). under hospitella in the 
National Collection. contained two species, not dis- 
tinguishable on superficial characters but radically dif- 
ferent in male and female genitalia. Eight of these 
specimens were the true hospitella. The remaining six 
are quantulella Hulst which Ragonot (1889), Hulst 
(1900), Hampson (1901), and later authors have treated 
as a synonym of hospitella, but which, along with par- 
vulella Ely and the tropical forms Dyar described under 
Eurythmia, I am removing to form the new genus 
Erelieva. ‘From Clarke’s notes I suspect that the orig- 
inal type series of hospitella in the British Museum is 
also a mixture. It consists of four females and two 
males. As no one of the cotypes was designated by 
Zeller or any later worker as the actual holotype I am 
here designating the female of his type series dated 
“17-V” as lectotype. It bears the following labels: A 
small white label with the numerals “17/5” to indicate 


307 


date; another white, rectangular label with the inscrip- 
tion ‘‘Bosque Co. Texas’’; the usual Zeller rectangular 
green label inscribed “‘hospitella Z. Texas Stt. 68”; and 
another white label with ‘‘Zell. Coll. 1884.” A sketch 
of the genitalia of this lectotype shows five of the larger 
signa in bursa (2 thorns and 8 discs) similar to the 
arrangement shown in figure 1130. 


632. Eurythmia hospitella yavapaella Dyar, new status 
Figure 1131 


Eurythmia yavapaella Dyar, Journ. New York Ent. Soc., vol. 14, 
p. 108, 1906.—McDunnough, Check list, No. 6392, 1939. 

Forewing more heavily dusted with dark scaling than 
in typical spaldingella; general color, to the naked eye, 
more brownish fuscous than ash gray and with the dark 
borders of the transverse lines more or less obscured in 
the dark overscaling of the wing; hind wing with a faint 
smoky tint. Alar expanse, 13.5-15.5 mm, 

Female genitalia of specimen from Glenwood Springs, 
Colo. (fig. 1131) show four large signa. Another female 
from the same locality and collection shows six of the 
larger signa. A San Diego, Calif., female (fig. 1131a) 
shows five. 


Typr nocauity: Yavapai County, Ariz. (type in 
USN). 

Foop prant: Unknown. 

DistrisutTion: Arizona, Yavapai County; Colorado, 
Glenwood Springs; California, San Diego (May, July), 
San Felipe Wash, San Diego County (June). 

This is at most a race and probably no more than a 
color variety of hospitella. I am holding the name 
merely as a precaution against possible future separa- 
tion. 

633. Eurythmia angulella Ely 
FieurE 1132 


Eurythmia angulella Ely, Proc. Ent. Soc. Washington, vol. 12, 
p- 202, 1910.—McDunnough, Check list, No. 6387, 1939. 

Eurythmia diffusella Ely, Proc. Ent. Soc. Washington, vol. 12, 
p. 202, 1910.—McDunnough, Check list, No. 6388, 1939. 
(New synonymy.) 

Similar to the dark western form, yavapaella, except 
dark margins of transverse lines of forewing somewhat 
more contrasted and hind wings darker. Bursa copu- 
latrix of female also larger than that of any western 
specimens. The name angulella may apply only to a 
northeastern race of hospitella but in the absence of any 
intergrading examples from any central United States 
localities we must keep angulella specifically separated. 
Ely’s diffusella, represented only by his male type, is 
simply a slightly more brownish specimen of angulella 
with slightly wider dark margin to the antemedial line. 
Alar expanse, 12-14 mm. 

Typm Ltocauity: East River, Conn. (angulella and 
diffusella, in USNM). 

Foop piant: Unknown. 

Represented in the National Museum by five males 
and nine females from the type locality (July) and one 
male from Trenton, Ontario, Canada (July). 


308 


634. Hurythmia fumella Ely 
Fieurn 1133 


EHurythmia fumella Ely, Proc. Ent. Soc. Washington, vol. 12, p. 
202, 1910.—McDunnough, Check list, No. 6389, 1939. 


Probably nothing more than an aberrant form of 
angulella; but known only from the unique female type, 
and differing from angulella in the wider diffusion of 
the whitish transverse lines of forewing, especially the 
antemedial line and in the blackish suffusion over the 
area between the lines. Bursa of female with but three 
of the larger signa. Alar expanse, 14 mm. 


Tyr Locauity: Hast River, Conn. (July, im USNM). 
Foop PLant: Unknown. 


191. Erelieva, new genus 


Typs or Genus: Pempelia quantulella Hulst. 

Characters of Hurythmia except: Male genitalia 
without an apical process to gnathos, the arms of 
gnathos fusing at their distal ends into the sclerotized 
subanal plate (subscaphium); aedeagus not tapering, 
expanding into a wide mouth at apex; penis unarmed 
except for some sclerotized wrinklings; transtilla pro- 
duced at middle into a stout, widely forked projection. 
Female genitalia with bursa copulatrix evenly and finely 
scobinate over entire inner surface; signa a single 
elongate series of (5 to 14) thornlike spines; no cluster 
of smaller spines at junction of bursa and ductus 
bursae, except in parvulella where such spines are 
arranged in a spiral extending into the ductus. Highth 
abdominal segment of male with compound: dorso- 
lateral scale tufts. 


The species included in this new genus are removed 
from Hurythmia because ‘of their radically different 
genitalia. They are a tropical group with a couple of 
extensions into the United States, while Hurythmia, as 
now restricted, seems to be confined to the continental 
United States. I suspect that, as in Hurythmia, the 
several specific names (with the exception of parvulella) 
represent no more than varieties of one widely dis- 
tributed, superficially variable species; but this cannot 
be determined from the specimens now available. 


635. Erelieva quantulella (Hulet), new combination 
Fieures 634, 1134 


Pempelia quantulella Hulst, Ent. Amer., vol. 8, p. 134, 1887. 


Eurythmia quantulella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 
116, 1889 (referred as synonym of E. hospitella). 


Eurythmia santiagella Dyar, Ins. Insc. Menstr., vol. 7, p. 62, 
1919. (New synonymy.) 

Eurythmia hospitella Dyar (not Zeller), Ins. Insc. Menstr., vol. 
13, p. 226, 1925. 

Forewing fuscous more or less dusted with white, 
giving the wing a pale gray cloor, usually paler and 
more contrasting on the basal area; antemedial line 
slightly oblique, bordered outwardly by blackish fus- 
cous; subterminal line faint with a narrow, blackish, 
inner border; discal dots faint, but distinct and separate. 
Hind wing dusky white, with veins perceptibly dark- 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


ened and dark fuscous borders along costa and 
outer margin. Alar’ expanse, 11-15 mm. 

Male genitalia as given for the genus. 
talia with from 10 to 14 signa. 

TYPE LOCALITIES: Blanco County, Tex. (quantulella, 
in AMNH, ex Rutgers); Santiago, Cuba (santiagella, in 
USNM). 

Foop puants: Opuntia (flowers), bell pepper, sor- 
ghum. The larvae are evidently flower or seed feeders 
in the plants attacked. 

Disrrizution: Unirep States: Texas, Blanco 
County (Aug.), Carrizo Springs (Oct.), Uvalde (June). 
Purrto Rico: Aguirre Central (Apr.), Coamo Springs 
(Apr.), Palmas Abajas (near Guayama, July), Puerto 
Real (Vieques Isl., Apr., July), San, Germdn (Apr., 
Aug.). Virein Istanps: Kingshill. (St. Croix; Mar., 
Oct., Nov., Dec.), no specific locality (Apr.): Cua: 
Santiago. Harr: Port au Prince (May). 

The type of quantulella in the Rutgers Collection is 
indeed a “‘very frail” specimen as Hulst said. It con- 
sists of a head, thorax, and a fragment of one forewing 
and is a female (not a male as stated by Hulst). A 
similar female from Blanco County, Tex. (nm USNM), is 
also labeled ‘‘type”’ by Hulst. The two specimens are 
obviously conspecific. Dyar’s santiagella has nothing 
to distinguish it from Texas specimens except the 
slightly larger size of his female type (14 mm.). The 
mele associated with it and also from Santiago is con- 
siderably smaller (11.5 mm.). A long series of Cornell 
specimens from Puerto Rico and the Virgin Islands 
exhibits considerable variation in size and the distinct- 
ness of the pale basal area of forewing and the extremes 
of variation in the number of spines (signa) in the bursa. 
There are no consistent genitalic or pattern differences 
to distinguish even local races within the species, which 
is apparently a tropical one that has extended its range 
or been introduced into Texas. 

The only Texas specimens I have seen are those in the 
National Museum, three males and five females. Three 
of the females came originally from Hulst through the 
Brooklyn and Fernald Collections. One female, from 
Uvalde, was reared June 1925 by A. P. Dodd from a 
larva feeding in the flowers of an Opuntia. ‘The remain- 
ing female and the three males were reared at Carrizo 
Springs, Oct. 28, 1944, by members of the Special 
Survey of the Division of Plant Quarantine of the U.S. 
Bureau of Entomology and Plant Quarantine from lar- 
vae feeding in the ripened pods of bell pepper. The 
sorghum record is from reared examples in the National 
Collection from St. Croix. 


Female geni- 


636. Erelieva coca (Dyar), new combination 


Eurythmia coca Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 339, 1914. 

Eurythmia coquilla Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 339, 
1914 (mew synonymy). 

Eurythmia mossa Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 340, 
1914 (new synonymy). 

Eurythmia uncta Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 340, 
1914 (new synonymy). 


Differs from quantulella only in its somewhat smaller 
average size (10-12 mm.) and the more suffused brown- 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 309 


ish color of forewing, the basal area being but slightly if 
any paler than the remainder of the wing. The color 
differences noted by Dyar among his supposed Panamé 
species are not so great as the variation exhibited by the 
Puerto Rican series of guantulella and the palpal differ- 
ences he stresses are purely imaginary. The heads of his 
specimens were so badly battered and the palpi so dis- 
torted it would be impossible to determine their exact 
lengths, shapes, or scaling even if there were such differ- 
ences, which there are not. I have grave doubt that 
coca is anything but a local form of quantulella; but in the 
absence of any examples of Frelieva from the mainland 
between Panama and Texas it seems best to retain coca 
as a specific designation until more material is available. 

The genitalia are not significantly different from those 
of guantulella. 

Typr Locaitius: Taboga Island, Panama (coca and 
uncta, in USNM); La Chorrera, Panama (coguilla, in 
USNM); Taboguilla Island, Panama (mossa, in USNM). 

Foop piant: Unknown. 

The only specimens IJ have seen are the 13 specimens 
(a male and 12 females dated Feb. and May) originally 
associated by Dyar with his several names and one 
female from Corazal, Panama (Mar.), which he had 
erroneously associated with his female type of “Hu- 
rythmia vestilla,” the latter, itself, a synonymy of 
Ephestiodes plorelia. 


637. Erelieva parvulella (Ely), new combination 
Ficure 1135 
Eurythmia parvulella Ely, Proc. Ent. Soc. Washington, vol. 12, 
p- 202, 1910.—McDunnough, Check list, No. 6386, 1939. 

Forewing brown with a very sparse dusting of whitish 
scales and a glossy sheen in contrast to the duller ap- 
pearance of quantulella and coca; transverse lines 
straight, narrow, the antemedial line more distinct than 
in the other two species; veins 3 and 5 very shortly 
stalked. Hind wing glossy fuscous; veins not appreci- 
ably darker and terminal margin but faintly so. Alar 
expanse, 11—-12.5 mm. 

Male genitalia not appreciably different from those 
of quantulella. Female genitalia with from 5 to 7 signa 
and a spiral of small spines at junction of bursa and 
ductus bursae and extending into the ductus. 

Typ LocaLiry: East River, Conn. (type in USNM). 

Foop prant: Unknown. 

A distinct species, distinguished by its glossy brown 
forewing, its darker hind wing, fewer signa, and the spiral 
spining at junction of bursa and ductus bursae. It is 
represented in the National Museum by a good series 
(37 specimens) collected by Ely in July 1909 and 1910 
at the type locality; but, so far as I know, has not turned 
up since from any other locality. It may be indigenous 
to the northeastern United States or an invader from 
the Tropics. I suspect that it is the latter. 


Group II 


[Hind wing with veins 3 and 4 both absent.]} 


Key to the genera of group III 


1. Forewing with veins 2 and 3 united; 4 and 5stalked. ....... OSES be DUE ES 


Forewing with veins 2 and 3stalked;4 and 5 united. . . 
2. Hind wing with discocellular vein absent; 6 greatly Petuced 
Hind wing with discocellular vein present; 6 normal . 


192. Genus Cabnia Dyar 


Cabnia Dyar, Journ. New York Ent. Soe., vol. 12, p. 108, 1904.— 
Forbes, Cornell Mem. 68, p. 639, 1923. (Lype of genus: 
Cabnia myronella Dyar.) 

Tongue very short; but exposed between the well- 
spaced labial palpi. Antenna pubescent; on male with a 
broadly scaled basal sezsment anda short, deep incurva- 
tion in base of shaft. Labial palpus upturned, short, 
not reaching to vertex; second segment triangularly 
scaled; third segment very short and broadly scaled. 
Maxillary palpus minute, hardly distinguishable. Fore- 
wing smooth; 9 veins; veins 2 and 3 stalked; 4 and 5 
united, separated from the stalk of 2-3 at base; 6 from 
below upper angle of cell, straight; 8 and 9 united; 10 
from the cell, well separated from 8 at base; male with 
a strong costal fold. Hind wing with vein 2 from well 
before lower outer angle of cell; 5 from the angle; 7 and 
8 completely anastomosed beyond cell; discocellular 
vein very weak, curved. Highth abdominal segment 
of male simple. 

Male genitalia with apical process of gnathos a very 
short, blunt hook. ‘Transtilla complete. Uncus long 
(but not longer than tegumen), moderately broad, 
beyond its broad base but slightly tapering to rounded 
apical margin. Harpe elongate; costa produced at 
apex into a short, free hook. Aedeagus simple, not 
tapering, and not appreciably enlarged at apex. Penis 
unarmed. 

Female genitalia with ductus bursae membranous, 
short (much shorter than bursa); bursa copulatrix 
elongate, membranous, weakly scobinate over most of 
its inner surface; signa present but very weak, consist- 
ing of a pair of minute disks; genital opening simple; 
ductus seminalis from anterior end of bursa. 

On its abbreviated tongue Cabnia could go in the 
Anerastiinae where it is placed in our recent list of 
North American Lepidoptera. However, on male 
genitalia and habitus it seems to fit better ito the 
Phycitinae. The ocelli are present and the tongue not 

310 


Seas Cabnia (p. 310) 
. Microphycita (p. 310) 
Rabiria (p. 311) 


entirely concealed by the palps. The genus contains 


one North American species. 


638. Cabnia myronella Dyar 
Fiaures 128, 627, 1138 


Cabnia myronella Dyar, Journ. New York Ent. Soc., vol.'12, p. 
108, 1904.—Forbes, Cornell Mem. 68, p. 689, 1923.— 
McDunnough, Check list, No. 6427, 1939. 


Forewing dark cinereus, uniformly colored, many of 
the scales tipped with dull white, giving the wing a 
slightly frosted appearance; transverse lines obsolete, 
or nearly so. Hind wing whitish, subpellucid; a very 
faint dark line along termen. Alar expanse 10-11 mm. 

Genitalic characters as given for the genus. The 
male genitalia are remarkably like those of Anagasta 
kiihniella, differing chiefly in the proportionally shorter 
uncus and the presence of a complete transtilla. 

TypE Locauity: Washington, D. C. (type in USNM). 

Foop piant: Unknown. 

Distripution: Massachusetts, Vinyard Haven (Aug.); 
District of Columbia (June) ; Florida, Lake Placid (Mar., 
Apr.). 


193. Genus Microphycita Dyar 


Microphycita Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 346, 1914. 
(Type of genus: Microphycita titillella Dyar.) 

Tongue well developed. Antenna pubescent, simple. 
Labial palpus upturned, slender; reaching to vertex; 
third segment as long as second, acuminate. Maxillary 
palpus minute, filiform. Forewing smooth; 9 veins; 
veins 2 and 3 united; 4 and 5 stalked, the stalk separated 
from 2 and 3 at base; 6 from below upper angle of cell, 
straight; 9 absent; 10 stalked with 8; male with costal 
fold. Hind wing with veins 2 and 5 rather short; 6 
greatly reduced and obsolescent at base; 7 absent; cell 
open (discocellular vein absent). Highth abdominal 
segment of male simple. 

Male genitalia without apical projection from gnathos, 
the slender lateral arms fusing into sclerotized subanal 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 311 


plate. Transtilla complete and greatly developed. Un- 
cusabsent. Harpe with cucullus projecting into a blunt 
point at apex. Anellus semitubular with elongate, 
slender, lateral lobes. Aedeagus curved, slightly ta- 
pered. Penis without armature. Vinculum as broad 
as long. 

Female genitalia with ductus bursae membranous, 
slender, longer than bursa; bursa copulatrix round, mem- 
branous; signum a finely, densely spined plate; genital 
opening simple; ductus seminalis from middle of bursa. 

A tropical genus of uncertain affinities. Dyar’s inter- 
pretation of the forewing venation differs from mine. 
He considered vein 4 absent and veins 3 and 5 stalked. 
The end result would be the same, whichever interpreta- 
tion is accepted. 


639. Microphycita titillella Dyar 


Figures 130, 636, 1137 
Microphycita titillella Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 346, 
1914. 


Forewing dull ocherous with a narrow dark (fuscous) 
shade along costa and termen, expanding somewhat at 
tornal area; transverse lines obscure; the antemedial line 
near middle of wing, vertical and straight, indicated 
chiefly by a narrow, fuscous, outer border; subterminal 
line well back from terminal margin, also indicated by a 
narrow, inner border, subparallel to antemedial line; no 
distinguishable discal dots. Hind wing pale smoky fus- 
cous. Alar expanse, 6-8 mm. 

Genital characters as given for the genus. 

Type tocauiry: Rio Trinidad, Panamé (type in 
USNM). 

Foop prant: Unknown. 

Distrisution: PanamA: Cabima (May), Rio Trinidad 
(Mar., May, June). 


194, Rabiria, new genus 


Typs or Genus: Microphycita conops Dyar 
Characters of Microphycita except: Forewing with 


vein 10 from the cell, separated from 8 at base. Hind 
wing with veins of normal length; vein 2 from before 
lower outer angle of cell; cell closed, discocellular vein 
curved. Male genitalia with apical process of gnathos 
well developed, U-shaped; uncus strongly developed 
(similar to that of Cabnia); harpe simple, elongate, api- 
cal margin evenly rounded; anellus a simple curved band, 
without lateral lobes; aedeagus straight, not tapering; 
transtilla a narrow, arched bridge. Female genitalia 
with signum a small, smooth disk. 

This genus is erected to receive conops originally in- 
cluded in Microphycita by Dyar along with the type 
species (titilella) of the latter genus. Why Dyar ever 
included the two species under one generic heading— 
even without examination of their genitalia—is difficult 
to understand; for the hind wing venation of the two 
is radically different. 


640. Rabiria conops (Dyar), new combination 
Figures 129, 635, 1136 
me pag conops Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 346, 
4, 

Forewing dark fuscous with a faint bronzy sheen; 
without transverse lines or discal spots. Hind wing 
pale translucent fuscous. Alar expanse, 8 mm. 

Genitalic characters as given for the genus. 

Type tocaurry: La Chorrera, Panamd (type in 
USNM). 

Foop prant: Unknown. 

DistRipution: PanaMA: Cabima (May), La Chorrera 
(Apr., May) Rio Trinidad (May). 

Represented so far only by the original type series, 
three males and one female. The type is a male (not a 
female as stated by Dyar). The genitalia are figured 
from this specimen and from a female (May) from the 
type locality. 


Genera and species unplaced, unrecognized, or referred 
from the Phycitinae 


Unplaced 


[Myelois] grossipunctella Ragonot 
Figures 915, 916 
Myelois grossipunctella Ragonot, Nouv. Gen., p. 8, 1888; Ent. 
Amer., vol. 6, p. 64, 1890; Monograph, pt. 1, p. 51, 1893.— 
McDunnough, Check list, No. 6069, 1939. 

This species was described from a male in the Ober- 
thiir Collection without locality label, but presumably 
from Australia. There is a series in the U.S. National 
Museum from California reared from larvae feeding on 
the “cottony cushion scale’ and undoubtedly intro- 
duced with that insect. This series was reared under 
No. 869-09 by Coquillet, April 1889, and the species 
has not been recovered since in California, so appar- 
ently it did not become established there. It should 
be dropped from our lists. 

It is not a Myelois as the genitalia clearly indicate; 
but its proper placement could only be made by one 
thoroughly acquainted with the Australian Phycitidae. 

Tyrer tocatiry: Australia [?] (location of type 
unknown to me). 

Foon: Icerya purchasi. Larva a predator. 
[Myelois] famula Zeller 

Figure 910 


Myelois famula Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 218, 
1881. 
Myelois restrictella Zeller, Horae Soc. Ent. Rossicae, vol. 16, 
p. 220, 1881. 
Salebria famula (Zeller) Ragonot, Monograph, pt. 1, p. 365, 1893. 
This species and its synonym (restrictella) are known 
only from their female types. Their genitalia are 
identical and suggest a placement much closer to 
Fundella than to either Myelois or Salebria. It will 
have to wait upon discovery of a male. 
TYPE LOCALITIES: Barranquila, Colombia (famula, 
in BM); Honda, Colombia (restrictella, in BM). 


[Nephopteryx] fuscifrontella Zeller 
Figure 911 
Nephopteryz fuscifrontella Zeller, Horae Soc. Ent. Rossicae, vol. 
16, p. 183, 1881.—Ragonot, Monograph, pt. 1, p, 282, 1893. 
In the U.S. National Museum are two females from 
Colombia identified by Ragonot as fuscifrontella. The 
genitalia of one of these are figured. A male will be 
needed for generic placement. Whatever it is, the 
species is not a Nephopteryz. 
Typ Locatiry: Honda, Colombia (type in BM). 
312 


{Salebria] nigricans Hulst 
FicurE 918 
Salebria nigricans Hulst, Canadian Ent., vol. 32, p. 171, 1900.— 
MeDunnough, Check list, No. 6201, 1989. 

I have seen nothing resembling the female type, 
whose genitalia are figured. A male will be needed 
for proper generic placement. 

Typr Locauity: Phoenix, Ariz. (type n AMNH, ex 
Rutgers). 

[Hypochalcia] cervinistrigalis Walker 
Ficure 919 
Hypochalcia cervinistrigalis Walker, List, pt. 27, p. 45, 1863.— 
Ragonot, Monograph, pt. 1, p. 585, 1893. 

Photographs of the female type and its genitalia are 
before me. The latter are figured. I have seen 
nothing like them. A male will be needed for generic 
placement. 

TyYpE LocaLiry: Santo Domingo [Dominican Repub- 
lic] (type in BM). 


Hypochalcia hulstiella Ragonot 


Hypochalcia hulstiella Ragonot, N. Amer. Phycitidae, p. 11, 1887; 
Monograph, pt. 1, p. 603, 1893.—Hulst, Phycitidae of N. 
nae p: 168, 1890.—MecDunnough, Check list, No. 6275, 
1939. 

According to Clarke the type lacks an abdomen and 
is otherwise in very poor condition. It is very likely 
not an American example. Ragonot’s figure (Mono- 
graph, pl. 21, fig. 5) strongly resembles one of the vari- 
ants of the European Hypochalcia ahenella with well- 
marked lines. 

TyprE Locanitry: ‘Texas’? (type in Paris Mus.). 


[Elasmopalpus] corrientellus Ragonot 
Fieure 917 


Elasmopalpus corrientellus Ragonot, Nouv. Gen., p. 23, 1888; 
Monograph, pt. 1, p. 423, 1893. 

This is not an Hlasmopalpus, but generic placement 
cannot be made without an authentic male. The type 
is a female. Its genitalia are figured. 

Typr LocaLity: Corrientes, Argentina (type in Paris 
Mus.). 

[Euzophera] postflavida Dyar 
Fiaure 1128 


Euzophera postflavida Dyar, Ins. Inse. Menstr., vol. 11, p. 29, 
1928. 

This species is represented by the female type and a 

female paratype (in USNM) from St. Laurent Maroni, 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


French Guiana, and a female (in Cornell Collection) 
from Tumatumari, Potaro River, British Guiana (June). 
The male is unknown. The distinctive genetalia and 
coloration readily identify the species. The basal half 
of hind wing and the abdomen (except for the last two 
segments) are ochre yellow; apical half of hind wing and 
the two caudal segments of abdomen blackish. A male, 
however, will be needed before the species can be 
properly placed generically. It is not a Huzophera. 

Typr tocatity: Nouveau Chantier, French Guiana 
(Sept.; type in USNM). 

Foop prant: Unknown. 


[Euzophera] rinmea Dyar 
Figure 1129 
Euzophera rinmea Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 335, 
1914. 

Known only from the female type. 
place without male. 

Typr Locauity: Rio Trinidad, Panama (Mar.; type in 
USNM). 

Foop piant: Unknown. 


Impossible to 


[Euzophera] came Dyar 
Figure 1126 


Euzophera came Dyar, Ins. Inse. Menstr., vol. 7, p. 56, 1919. 


Represented in the National Collection by the female 
type and five other females from the type locality. 
Not a Euzophera but cannot be placed without male. 

TyYpx Locauity: Cayuga, Guatemala (type in USNM). 

Foop prant: Unknown. 


[Euzophera] gais Dyar 
Euzophera gais Dyar, Ins. Insc. Menstr., vol. 7, p. 57, 1919. 


Represented only by the unique female type, which is 
probably a synonym of Anthropteryx irichampa Dyar. 
Both species have similar genitalia. Synonymy and 
proper placement will have to wait upon recovery of a 
male. 

TypE LocaLity: Cayuga, Guatemala (June; type in 
USNM). 

Foop prant: Unknown. 


Genus Anthropteryx Dyar 
FiaurE 1059 


Anthropteryx Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 335, 1914. 
Nels of genus: Anthropteryx irichampa Dyar, loc. cit., p. 
336. 

This genus is probably a good one, but was erected 
upon a freak specimen and the venational ‘‘characters”’ 
as given are incorrect. Vein 4 is absent from the left 
forewing of irichampa but present on the right wing and 
long stalked with 5, and vein 3 is connate with the stalk 
of 4-5, not stalked as stated by Dyar. The species 
belongs, therefore, in venational group A (11 veins in 
forewing and vein 2 from the cell). Its genitalia indi- 
cate close relationship to Moodnopsis. A male will be 
needed before the status of Anthropteryx can be deter- 

300329—56——21 


313 


mined. The type of genus is represented only by its 
unique female type. 

TyrpELocauity: Taboga Island, Panama (Feb.; type in 
USNM). 

Foop pirant: Unknown. 


[Moodna] formulella Schaus 
Figure 1127 


Moodna formuleila Schaus, Ann. Mag. Nat. Hist. ser. 8, vol. 11, 
p. 252, 1913. 

A large (28 mm.), strikingly marked species which 
should be easy to recognize if more specimens are dis- 
covered. So far known only from the unique female 
type. A male will be needed for proper placement. It 
is obviously not a Moodna, having 11 veins in the fore- 
wing, with 4 and 5 stalked for a little more than half 
their lengths; 2 from before the lower outer angle of cell; 
3 from the angle and slightly separated from the stalk 
of 4-5; 6 straight from below upper angle of cell; 8 and 9 
long stalked; 10 from the cell but rather close to the 
stalk of 8-9. Hind wing with 2 from well before the 
angle of the cell; 3 and 5 from the angle, approximate at 
base; 7 and 8 anastomiosed from most of their lengths 
beyond cell; cell one-third the length of wing; discocellu- 
lar vein curved. The transverse lines of forewing are 
rather close together, the antemedial line being near the 
middle of the wing. The dark areas and markings of 
the wing are a reddish brown rather than the usual 
fuscous shades. 

Typ Locauity: Juan Vifias, Costa Rica (Feb.; type 
in USNM). 

Foop prant: Unknown. 


[Eucampyla] putidella Schaus 
Fieure 1095 


Eucampyla putidella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 
11, p. 250, 1913. 

Known only from the female type. A male is needed 
for proper placement. Certainly does not belong in 
Eucampyla, which is an Australian genus not represented 
in the American faunas. May be an aberrant exam- 
ple of Vitula. The venation and female genitalia are 
those of Vitula except for the presence of vein 9 in both 
forewings (8 and 9 being long stalked) and the absence 
of scobinations in the bursa. 

TypE Locatiry: Mount Poas, Costa Rica (May; type 
in USNM). 

Foop puant: Unknown. 


[Honora] dulciella Hulst 
Figure 951 
Honora dulciella Hulst, Journ. New York Ent. Soc.. vol. 8, p. 223, 
1900.—McDunnough, Check list, No. 6349, 1939. 

This species has the venation of Honora but the fe- 
male genitalia are altogether wrong for that genus. A 
male will be needed for proper placement. At present 
dulciella is represented only by the female type. 

Typx Locatity: Palm Beach, Fla. (type in USNM). 

Foop prant: Unknown. 


314 


Unrecognized 


Myelois infusella Zeller 


Myelois infusella Zeller, Isis von Oken, 1848, p. 869. 
Salebria infusella (Zeller) Ragonot, Monograph, pt. 1, p. 352, 
1893. 
Unknown to me. The type is supposed to be in the 
Berlin Museum, but Dr. Hering was unable to locate it. 
TYPE LocaLiry: Port-au-Prince, Haiti. 


Genus Phycitopsis Ragonot 


Phycitopsis Ragonot, N. Amer. Phycitidae, p. 4, 1887; Mono- 
graph, pt. 1, p. 185, 1893.—Hulst, Phycitidae of N. Amer., 
p. 1388, 1890. (Type of genus: Phycitopsis flavicornella 
Ragonot, loc. cit.—Hulst, op. cit., p. 134—MeDunnough, 
Check list, No. 6124, 1939.) 

T have seen nothing from America that could be this 
genus or species. According to Clarke’s notes the fe- 
male type is not in either the Paris or British Museum 
Collections. From the Ragonot descriptions the genus 
should go into venational division D and the species 
must strongly resemble the Old World Phycita spissi- 
cella (Fabricus). I suspect that flavicornella is a Kuro- 
pean specimen and the ‘“Texas’’ locality is 2 mislabeling. 
This is quite likely if the type was a Boll specimen. 

There is a discrepancy between Ragonot’s description 
and his figure of the labial palpus (Monograph, pl. 8, 
fig. 10). The figure shows an oblique, moderately long 
(not @ very short) ascending palp. 

Tyerr Locauity: ‘“Texas’’ (type lost?). 


Euzophera intextella (Zeller) 


Myelois intextella Zeller, Isis von Oken, 1848, p. 871. 
Euzophera intextella (Zeller) Ragonot, Monograph, pt. 2, p. 53, 
1901. 

Unknown to me. The type is supposed to be in the 
Berlin Museum but Dr. Hering has been unable to 
locate it. 

Typrr Locatity: St. Thomas, British West Indies. 


Euzophera daedalella Ragonot 


Euzophera daedalella Ragonot, Nouv. Gen. p. 32, 1888; Mono- 
graph, pt. 2, p. 56, 1910. 

Unknown to me. This type is also supposed to be in 
the Berlin Museum. Dr. Hering was unable to locate 
it. : 
Typr Locatity: Chanchamayo, Peri. 


Zophodia brevistrigella Ragonot 


Zophodia brevistrigella Ragonot, Nouv. Gen., p. 31, 1888; Mono- 
graph, pt. 2, p. 24, 1901. 

According to Ragonot the type (2) was deposited in 
the Staudinger Collection in Berlin. It has not been 
located and so far I have seen nothing that agrees with 
the description and figure published in the Ragonot 
monograph. 

TypxE Locauity: Nova Friburgo, Brazil. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Zophodia subcanella (Zeller) 


Myelois subcanella Zeller, Isis von Oken, 1848, p. 873. 
Zophodia subcanella (Zeller) Ragonot, Monograph, pt. 2, p. 22, 
1901. 
Unknown to me. Type supposed to be in Berlin 
Museum but not located. 
TYPE LocaLity: Brazil. 


Psorosa disticta (Zeller) 
Euzophera disticta Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 


232, 1881. 
Psorosa disticta (Zeller) Ragonot, Monograph, pt. 2, p. 107, 1901. 


A photograph of the genitalia of the female type 
shows two large opposed signa composed of cup- 
shaped, strongly spined concave plates (similar to those 
of Rotruda mucidella). I have seen nothing that 
matches the type exactly either in structure or super- 
ficially. It is obviously not a Psorosa or a Huzophera; 
but generic placement cannot be made without a male. 

Tyrr Locauity: Maraquita, Colombia (type in BM). 


Hornigia clitellatella Ragonot 
Hornigia clitellatella Ragonot, Nouv. Gen., p. 36, 1888. 


In the Ragonot monograph (pt. 2, p. 269, 1901) 
Hampson places clitellatella in Moodna. It apparently 
belongs somewhere near Moodna, in venational group 
C, but accurate placement must wait upon discovery of 
a male from Chile. The female type is in the Paris 
Museum. I have seen nothing agreeing with Ragonot’s 
description or figure (Monograph, pl. 34, fig. 12). 

Typr Locatity: Callao, Chile (type in Paris Mus.). 


Moodna olivella Hampson 


Moodna olivella Hampson, in Ragonot, Monograph, pt. 2, p. 268, 
1901. 

I have seen nothing agreeing with either the descrip- 
tion or figure (pl. 49, fig. 12) in the Monograph. Clarke’s 
notes on the genitalia of the unique male type in the 
British Museum show that olivella cannot be a Moodna. 
The gnathos apparently is more like that of Moodnopsis. 
Placement will have to wait upon recovery of additional 
specimens. 

TypsE LocaLiry: Petrépolis, Brazil. 


Euzopherodes megalopalis Hampson 


Euzopherodes megalopalis Hampson, Ann. Mag. Nat. Hist., ser. 7, 
vol. 14, p. 181, 1904. 

According to Clarke’s notes the type im the Bzitish 
Museum is a female (not a male as stated by Hampson) 
and has a glued-on abdomen, which may or may not 
belong with the remainder of the specimen. <A photo- 
graph of the type shows a moth superficially similar to 
Nicetiodes apianella Schaus in markings. Its venation 
(if correctly given) places it in venational group D 
where we have nothing resembling it. We shall have 
to wait upon additional specimens (male and female) 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 815 


for proper placement. Hampson’s reference to the 
Old World Euzopherodes is obviously wrong. 
Type Locauity: Nassau, Bahamas. 


Genera and species referred from Phycitinae 


Genus Eutrichocera Hampson 


Eutrichocera Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 14, p. 
182, 1904. (Type of genus: Eutrichocera paurolepidalis 
Hampson, loc. cit.) 

This is an Epipaschiid. The genus is a synonym of 

Pococera and its type species a synonym of P. insularella 

Ragonot. 


Tyrr Locatity: Abaco, Bahamas (type, o’, in BM). 


Myelois atristrigella Ragonot 


Myelois atristrigella Ragonot, Monograph, pt. 1, p. 40, pl. 18, 
fig. 3, 1893. 

This species goes to the Hpipaschiidae. I have 
examined the type. It has the normal epipaschiid 
venation, but will require a new genus, for it does not 
fit comfortably in any of our described epipaschiid 
genera. Its peculiar forewing pattern should make it 
easy to identify if more specimens are recovered. At 
present it is known only from the unique male type. 

Typr Locatity: Puerto Rico (type in Zool. Mus. 
Univ. Berlin). 


Genus Psammia Hampson 


Psammia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 71, 
1930. (Type of genus: Psammia flavipicta Hampson, loc. 
cit—McDunnough, Check list, No. 6234, 1939.) 


Genus and species go to Anerastiinae. 
TypE Locauity: Florida (type in BM). 


Megasis indianella Dyar 
Megasis indianella Dyar, Ins. Inse. Menstr., vol. 11, p. 28, 1923.— 
MeDunnough, Check list, No. 6265, 1939. 


Goes to Anerastiinae. Dyar’s species is a synonym 
of Ragonotia olivella Hulst. 
TypE Locauity: Indian Wells, Calif. (type in USNM). 


Euzophera tintilla Dyar 


Euzophera tintilla Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 334, 
1914. 


Goes to Anerastiinae. Described from single male. 
Generic placement uncertain. 

Typr tocauiry: Porto Bello, Panama (Apr.; type in 
USNM). 

Foop: Unknown. Larva probably a coccid feeder. 


Euzophera conquistador Dyar 
Euzophera conquistador Dyar, Proc. U. 8. Nat. Mus., vol. 47, 
p. 335, 1914. 
Goes to Anerastiinae. Described from single female. 
Generic placement uncertain. 


TyprrE LocaLity: Cabima, Panamé& (May; type in 
USNM). 
Foop: Unknown. 


Euzophera mabes Dyar 


Euzophera mabes Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 335, 
1914. 
Goes to Anerastiinae. Described from female. Ge- 
neric placement uncertain. 


Types Locauity: Taboga Island, Panam (Feb.; type 
in USNM). 
Foop: Unknown. 


Euzophera climosa Dyar 


Euzophera climosa Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 335, 
1914, 


The original series before Dyar consisted of three 
males and four females (not one male and five females as 
stated) and only the male holotype and one male from 
the type locality can be definitely assigned to the name. 
The other male from the type locality is congeneric but 
a distinct, undescribed species. The four females repre- 
sent two species, each of which consists of one example 
from Rio Trinidad and one from Porto Bello. Which 
of these female species, if either, represents the other 
sex of climosa is not possible to determine. All these 
specimens belong in the Anerastiinae and will require 
reference to a new genus when that subfamily is revised. 

Typs Locauity: Rio Trinidad, Panamé' (Mar.; type 
in USNM). 

Foopv: Unknown. Probably a coccid feeder. 


Zophodia epischnioides Hulst 
Zophodia epischnioides Hulst, Canadian Ent. vol. 32, p. 173, 1900. 


I have been unable to locate the type of this species in 
the Rutgers Collection or elsewhere. From the descrip- 
tion I suspect that it is an Anerastiine. It is obviously 
not a Zophodia. Disposition of the name can wait upon 
a revision of the Anerastiinae. 

Typr Locatity: Not given (type lost?). 

Foop prant: Unknown. 


Genus Harnochina Dyar 


Harnochina Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 340, 1914. 
(Type of genus: Harnochina rectilinea Dyar, loc. cit.) 


Goes to the Anerastiinae. The tongue is absent. 

Typr Locauity: Corozal, Canal Zone, Panamé (Mar.; 
type in USNM). 

Foop piant: Unknown. 


Genus Nasutes Hampson 
Nasutes Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 53, 
1930. (Type of genus: Nasutes venata Hampson, op. cit. p. 
54.) 


This genus is a synonym of Bandera Ragonot and its 
type species (venata) a synonym of B. cupidinella 
(Hulst). 


316 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


I have before me an enlarged photograph of the geni- 
talia of the male type of venata and a slide of the genitalia 
of the male type of cupidinella; the two, both from Colo- 
rado, are identical in all details. The genus Bandera is 
a true anerastiid on habitus and all characters except 
the tongue. This organ is reduced, but not concealed 
between the labial palpi as in typical Anerastiinae. 
However, Bandera lacks any trace of an ocellus, as do 
several other Anerastiinae. This organ is present in all 
genera of Phycitinae in the Americas. 

Typr Locauity: Colorado (type in BM). 

Foop prant: Unknown. 


Maricopa albocosiella Hulst 


Maricopa albocostella Hulst, Canadian Ent., vol. 32, p. 176, 1900. 

Valdiria albocostella (Hulst) Barnes and McDunnough, Check 
list of the Lepidoptera of Boreal America, No. 5755.— 
McecDunnough, Check list, No. 6354, 1939. 


Not a Valdivia. Goes to Anerastiinae. Has no 
tongue visible. The type is a male with veins 4 and 5 
united in forewing and with a slight excavation in the 
base of the antennal shaft. 

TypE LtocaLity: Anglesea, N. J. (type in AMNH, ex 
Rutgers). 

Foop piant: Unknown. 


Checklist of American Phycitinae 


1. CryprosLaBEs Zeller 


ile 


gnidiella (Milliére): Europe, Africa, Asia, 
Bermuda, Venezuela, Brazil 


2. Acrosasis Zeller 


ore 


. Indigenella (Zeller): 


. feltella Dyar: 
. palliolella Ragonot: 


. caryalbella Ely: 
. juglandis (LeBaron): Eastern U. S. 

. sylviella Ely: Eastern U. S., Canada 
. kearfottella Dyar: 
. caryae Grote: 
. evanescentella Dyar: 
. stigmella Dyar: 
. aurorella Ely: Eastern U.S. 
. peplifera Dyar: 
. exsulella (Zeller), new comb.: 


. angusella Grote: 


. demotella Grote: 

. latifasciella Dyar: 
. irrubriella Ely: Eastern U.S. 
. normella Dyar: 
. Malipennella Dyar: Eastern U. S. (Con- 


. cirroferella, Hulst: 
. cunulae Dyar and Heinrich: Eastern U. S. 
. caryivorella Ragonot: 


Mineola Hulst 

Seneca Hulst 

Acrocaula Hulst 

Eastern U.S. and Can- 
ada, California 

nebulo (Walsh) 

nebulella (Riley) 

zelatella (Hulst) 


. grossbecki (Barnes and McDunnough), new 


comb.: Florida 
. vaccinii Riley: U.S. 
. amplexella Ragonot: Eastern U.S. 
. tricolorella Grote: U.S., Canada 
scitulella Hulst 
. comptella Ragonot: Western U.S. 
. minimella Ragonot: Eastern U.S. 


nigrosignella Hulst 

Eastern U. S., Canada 
Eastern U. S., Canada 
albocapitella Hulst 

U.S. (Connecticut) 


Eastern U. S. 
Eastern U. S., Canada 
U.S. (Florida) 
Eastern U.S. 


Eastern U. S. 
Eastern U. 8S. 
septentrionella Dyar 
Eastern U. S., Canada 
eliella Dyar 
Eastern U. S. 

Eastern U.S. 


Eastern U.S. (Connecticut) 


necticut) 


. dyarella Ely: Eastern U.S. (Connecticut) 
. ostryella Ely: Eastern U.S., Canada 

. secundella Ely: Eastern U.S., Canada 

. coryliella Dyar: Eastern U.S. 

. hebescella Hulst: 


Eastern U. S. (New Jer- 


sey) 
Eastern U.S. (Texas) 


Eastern and South- 
western U.S. 


“I 


35. comacornella (Hulst), new comb.: Eastern 


U.S. (Texas) 

36. betulella Hulst: Eastern and Western U.S., 
Canada 

37. rubrifasciella Packard: Eastern U. S., 
Canada 

alnella McDunnough 

38. comptoniella Hulst: Eastern U. S., Canada 

39. myricella Barnes and McDunnough: U. S. 
(Florida) 

40. tumidulella (Ragonot), new comb.: 
(Florida) 


USS: 


. ReopopHakEa Guénée 


41. caliginella (Hulst), new comb.: 
fornia, Arizona) 
caliginoidella (Dyar) 
42. supposita (Heinrich), new comb.: 
(British Columbia) 


U.S. (Cali- 


Canada 


. TracHycrra Ragonot 


43. pallicornella (Ragonot): U.S. (Texas) 


. ANABASIS Heinrich, new genus 


44. ochrodesma (Zeller), new comb.: U. S., 
(Florida), México, Guatemala, Panama 
Colombia, West Indies 

crassisquamella (Hampson) 


. Mitpreixra Dyar 


45. constitutionella Dyar: México, Guatemala 


. SEMATONEURA Ragonot 


46. atrovenosella Ragonot: México, Costa Rica, 
Colombia, Ecuador, Pert, Argentina 
47. abitus Heinrich, new species: Ecuador 


. HypsrpyLa Ragonot 


48. grandella (Zeller): U. S. (Florida), West 
Indies and Tropical America to Argen- 
tina 

cnabella Dyar 


49. ferrealis (Hampson), new comb.: ‘Tropical 
America (Costa Rica to Brazil) 
50. dorsimacula (Schaus), new comb.: Costa 


Rica 


51. fluviatella Schaus: Costa Rica 


. Hemipritocera Ragonot 


52. chinographella Ragonot: French Guiana, 
Brazil, Pera 
53. bigrana (Zeller): México, Colombia 


54, plumigerella (Ragonot), new comb.: ‘Amer. 


Merid.” 

55. letharda (Schaus), new comb.: Panamé, 
México 

56. jocarella (Schaus): Costa Rica, Panamé, 
Brazil 


57. exoleta (Zeller): Colombia 


317 


318 


10. 


11. 


12. 


13. 


14. 


15. 


16. 


17. 


18. 


i). 


20. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Crocipomera Zeller 
58. turbidella Zeller: 
U.S. (Texas) 
59. fissuralis (Walker): 
Puerto Rico 
adonea (Felder and Rogenhofer) 
60. stenopteryx (Dyar), new comb.: México 
CunisEertTa Heinrich, new genus 
61. subtinctella (Ragonot), new comb.: 
ern U.S. and Canada 
Heras Heinrich, new genus 
62. disjunctus Heinrich, new species: 
Apanarsa Heinrich, new genus 
63. intransitella (Dyar), new comb.: 
(Arizona, New Mexico) 
Birinus Heinrich, new genus 
64. russeolus Heinrich, new species: 
Guiana 
BeErreLia Barnes and McDunnough 
65. grisella Barnes and McDunnough: U. 5. 
(Arizona) 
Hypareyria Ragonot 


Cuba, Jamaica, México, 


Dominican Republic, 


West- 


Colombia 


WES: 


British 


66. definitella (Zeller): Puerto Rico, Virgin 
Islands, Colombia, Brazil 
67. slossonella (Hulst), new comb.: U. S. 


(Florida), México 
tenuella (Barnes and McDunnough) 
CuaAraAnrica Heinrich, new genus 


68. annuliferella (Dyar), new comb.: U. S. 
(New Mexico, Arizona) 
69. hystriculella (Hulst), new comb.: U. S. 


(Texas, Florida) 

70. bicolorella (Barnes and McDunnough), new 
comb.: U.S. (Arizona, Nevada, Cali- 
fornia) 

Myetopstis Heinrich, new genus 

71. coniella (Ragonot), new comb.: 
ada, México 

nefas (Dyar) 

72. immundella (Hulst), 
(Texas) 

73. subtetricella (Ragonot), new comb.: 
Canada 

zonulella (Ragonot) 
obnupsella (Hulst) 


U.S., Can- 


new comb.: U. S. 


wese 


74. minutularia (Hulst), new comb.: U. S. 
(Texas) 
75. alatella (Hulst), new comb.: Western U.S. 


rectistrigella (Ragonot) 
fragilella (Dyar) 
piazzella (Dyar) 
AnypsIpyLa Dyar 
76. univitella Dyar: Cuba, México, Guatemala, 
Panamé, Venezuela, Brazil, Peri, Heua- 
dor, Jamaica 
ApomyYrEtois Heinrich, new genus 
77. bistriatella (Hulst), new comb.: 
U.S., Canada 
bilineatella (Ragonot) 


Eastern 


21. Ecromyxrtots Heinrich, new genus. 


22. 


23. 


24. 


25. 


78. decolor (Zeller), new comb.: ‘Tropical 
America, 
ephestiella (Hampson) 
79. ceratoniae (Zeller), new comb.: Hurope, 


U. S. (@lorida), Puerto Rico, Jamaica, 


Argentina 
oporedestella (Dyar) 
80. muriscis (Dyar), new comb.: ‘Tropical 
America 
palpalis (Dyar) 
81. furvidorsella (Ragonot), new comb.: Puerto 
Rico 
82. zeteki Heinrich, new species: Panama 
PaRAMYELot!s Heinrich, new genus. 
83. transitella (Walker), new comb.: U. S., 


tropical America 
notatalis (Walker) 
solitella (Zeller) 
duplipunctella (Ragonot) 
venipars (Dyar) 
cassiae (Dyar) 
Psrupopivona Dyar 
84. commensella Dyar: México 
85. cispha Dyar: Guatemala, Costa Rica, Brit- 
ish Honduras 
86. santa-maria Dyar: Guatemala 
87. carabayella Dyar: Pert, Bolivia, Colombia 
PROTOMOERBES Heinrich, new genus 
88. aberrans Heinrich, new species: 
89. separabilis Heinrich, new species: 
Dratomocrera Ragonot 
Cabima Dyar 
90. tenebricosa (Zeller): 
ana, Costa Rica 
91. dosia (Dyar), new comb.: Panamé 
92. excisalis (Hampson), new comb.: 
Guiana, Bolivia (?) 
93. decurrens (Dyar), new comb.: 
94. majuscula Heinrich, new species: 
95. albosigno Heinrich, new species: 
96. hoplidice (Dyar), new comb.: 
97. extracta Heinrich, new species: 
98. mochlophleps (Dyar), new comb.: 


Colombia 
Colombia 


Colombia, French Gui- 


French 


Panamaé 
Brazil 
Brazil 
Panama 
Costa Rica 
México 


. Psrupocasima Heinrich, new genus 


99. castronalis Heinrich, new species: Brazil 
100. fearnella (Schaus), new comb.: Costa Rica, 
Guatemala 
101. guianalis Heinrich, new species: 
Guiana, British Guiana 
euzopherella (Dyar), new comb.: Panama 
Panama 


French 


102. 
103. pombra (Dyar), new comb.: 


104. nigristrigella (Ragonot), new comb.: Brazil 

105. arizonensis Heinrich, new species: U. S. 
(Arizona) 

106. expunctrix (Dyar and Heinrich), new comb.: 
Brazil 

107. perrensiella (Ragonot, new comb.: Argen- 
tina 


27. 


28. 


29. 


30. 


31. 


32. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


108. rubrizonalis (Hampson), new comb.: French 
Guiana, Brazil 
Hyatospita Ragonot 


109. stictoneurella Ragonot: México, Guatemala, 
Brazil 

110. celiella Schaus: Costa Rica 

111. insequens Heinrich, new species: Bolivia, 
Colombia 

112. majorina Heinrich, new species: México 

113. fulgidula Heinrich, new species: Cuba 

114. egenella (Ragonot), new comb.: Brazil 

115. xanthoudemia (Dyar), new comb.: Panama, 
Costa Rica 

116. angulinella (Schaus), new comb.: Costa 
Rica 

117. clevelandella (Dyar): Panam& 

118. semibrunneella Ragonot: Colombia 


Funpeuia Zeller 
119. pellucens Zeller: U.S. (Florida), West In- 
dies, Brazil, Bolivia 
cistipennis (Dyar) 
argentina Dyar: U. S. (Florida, Texas), 
West Indies, Venezuela, Brazil, Argen- 
tina 
eucasis Dyar 
121. agapella Schaus: Galapagos Islands 
122. ignobilis Heinrich: México, Guatemala, 
Costa Rica, Cuba, Puerto Rico, Haiti 
ahemora Dyar: México, Guatemala, Costa 
Rica 
DirunpDELia Dyar 
124. corynophora Dyar: 
French Guiana 
125. subsutella (Schaus), new comb.: 


120. 


123. 


Guatemala, Panamé, 


Costa Rica 


126. distractor Heinrich, new species: Puerto 
Rico 
127. tolerata Heinrich, new species: Bolivia 


CopTarTHRIA Ragonot 

128. dasypyga (Zeller): 

PromMyYLEA Ragonot 

129. lunigerella Ragonot: 

Canada. 
130. lunigerella glendella (Dyar): 
131. dyari Heinrich, new name: 
zimmermani (Druce) 
drucei (Dyar) 

132. druceii (Ragonot), new comb.: 

133. mindosis Dyar: México 

134. dasystigma Dyar: México 

ANADELOSEMIA Dyar 

135. senesciella (Schaus): Costa Rica 

136. tecmessella (Schaus): Costa Rica 

137. fifria Dyar: Guatemala 

138. base Dyar: Guatemala 

139. obstitella (Schaus), new comb.: Costa Rica 

140. texanella (Hulst), new comb.: U.S. (Texas, 
Florida), Puerto Rico, Cuba 

dulciella (Hulst) 

condigna Heinrich, new species: 

zona) 


Colombia, Guatemala 
Western U. S. and 
Colorado 
México 


Guatemala 


141. U.S. (Ari- 


33. 


34. 


35. 


36. 


37. 


38. 


39. 


319 


Dasypyea Ragonot 
142. alternosquamella Ragonot: 
Canada 
stictophorella Ragonot 
Rampyuua Dyar 
143. orio Dyar: México 
144. polydectella (Schaus): Costa Rica 
145. subcaudata (Dyar), newcomb.: Guatemala, 
Costa Rica, Brazil 
146. lophotalis Heinrich, new species: 
Guatemala 
Fuurapa Heinrich, new genus 
147. querna (Dyar), new comb.: Panamé 
148. carpasella (Schaus), new comb.: Galdpagos 
Islands 
Scoryuus Heinrich, new genus 
149. cubensis Heinrich, new species: 
Davara Walker 
Homalopalpia Dyar 
Eucardinia Dyar 
150. caricae (Dyar), new comb.: 
Tropical America 
dalera (Dyar) 


Western U. S., 


México, 


Cuba 


U.S. (Florida), 


151. columnella (Zeller), new comb.: Colombia 

152. nerthella (Schaus), new comb.: México, 
Guatemala, Costa Rica 

euthales (Dyar) 

153. paranensis (Dyar), new comb.: Brazil 

154. azonaxsalis Walker: Brazil 

155. interjecta Heinrich, new species: Puerto 
Rico, Dominican Republic 

156. rufulella (Ragonot), newcomb.: Puerto Rico 


Sarasora Hulst 
Cuba Dyar 
157. plumigerella Hulst: U.S. (Florida) 
158. furculella (Dyar), new comb.: Cuba, Puerto 
Rico, Dominica, Virgin Islands 
159. ptyonopoda (Hampson), new comb.: 
ward Islands 
Presmopopa Zeller 
Discopalpia Ragonot 
Amphycitopsis Dyar 
160. rubicundella Zeller: Brazil 
161. xanthomera Dyar: Guatemala, 
Costa Rica, French Guiana 
zanthozona Dyar 
trichomata (Zeller): Colombia 
flavicans (Zeller): Colombia, French Guiana 
fratella Dyar 
ragonoti (Dyar), new comb.: 
temala, Costa Rica 
isabella (Dyar), new comb.: 
xanthopolys Dyar: Panama 
parva Heinrich, new species: Panama 
semirufella (Zeller): Colombia 
apocerastes Dyar: México, Costa Rica, 
French Guiana, Brazil, Dominica 
montella Schaus: Costa Rica 


Wind- 


Panam 4 


162. 
163. 


164. México, Gua- 


165. Costa Rica 
166. 
167. 
168. 
169. 


170. 


320 


40. 


41. 


42. 


43. 


44, 


45. 


46. 


47. 


48. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Atuertoca Heinrich, new genus 
171. subrufella (Hulst), new comb.: 
ida), Cuba, Virgin Islands 
jiliolella (Aulst). Virgin Islands 
ptychis (Dyar) 


U.S. (Flor- 


172. bondari Heinrich, new species: Brazil 
PRAEDONULA Heinrich, new genus 
173. almonella (Dyar), new comb.: Panama 


Prapus Heinrich, new genus 
174. burdettellus (Schaus), new comb.: 
Rica, Guatemala 
semproniella (Schaus) 
175. dissitus Heinrich, new species: Brazil 
176. subaquilellus (Ragonot), new comb.: Guate- 
mala, 
Gasinivs Heinrich, new genus 
177. paulsoni (Ragonot), new comb.: 
Crracantuia Ragonot 
Procandiopa Dyar 
178. mamella (Dyar), 
Guatemala 
179. vepreculella Ragonot: 
Merearraria Ragonot 
180. peterseni (Zeller): 
Brazil, Pert 


Costa 


Chile 


new comb.: Panama, 
Ecuador 


Guatemala, Colombia, 


181. squamifera Heinrich, new species: Costa 
Rica 

182. frustrator Heinrich, new species: Costa Rica 

183. schausi Heinrich, new species: Costa Rica 

184. cervicalis Dyar: Cuba 

185. alpha Heimrich, new species: Guatemala, 


México, Costa Rica, Panam4, Bolivia, 
Brazil 

beta Heinrich, new species: México, Guate- 
mala, Costa Rica, Trinidad 

Drescoma Dyar 

187. cyrdipsa Dyar: México, Guatemala, Pan- 

ama, French Guiana 
188. cinilixa Dyar: Guatemala, Panamé 


186. 


Mownopritora Hulst 
189. pergratialis (Hulst): U.S. 
grotella (Ragonot) 
nubilella Hulst 
ZAMAGIRIA Dyar 
190. dixolophella Dyar: Panama 
191. pogerythrus Dyar: México, Guatemala 
192. hospitabilis Dyar: Cuba 
193. masculinus Dyar: Guatemala 


194. australella (Hulst), new comb.: Weis: 
(Texas, Florida) 
bumeliella (Barnes and McDunnough): 
U.S. (Texas, Florida) 
fraterna Heinrich, new species: 
laidion (Zeller): 
America 
deva Dyar 
striella Dyar 
ipsetona Dyar: 


195. 
196. 


Cuba 
U. S. (Florida), Tropical 


197. Costa Rica 


49. ANEGCEPHALESIS Dyar 


198. arctella (Ragonot), new comb.: U. S. 
(Florida), Bahamas, Cuba 
cathaeretes Dyar 
50. Maeiriopsis Heinrich, new genus 
199. denticosella (Dyar), new comb.: Tropical 


51. 


52. 


53. 


54. 


55. 


America 
cristalis (Hampson) 
ANcYLOSToMIA Ragonot 
200. stercorea (Zeller): 
America 
agnobilis (Butler) 
diffissella (Zeller) 
201. sauciella (Zeller): Colombia 
202. argyrophleps Dyar: México, Guatemala 
203. euchroma Dyar: Brazil 
Caristanius Heinrich, new genus 
204. pellucidellus (Ragonot), new comb.: Puerto 
Rico, St. Vincent, Jamaica, Surinam, 
Brazil 
melanoplaga (Hampson) 
205. decoloralis (Walker), new comb.: 
U.S. 
metagrammalis (Walker) 
furfurellus (Aulst) 
floridellus (Hulst) 
206. guatemalellus (Ragonot), 
Guatemala 
Erreia Zeller 
207. zinckenella (Treitschke): 
U.S., Tropical America 
etiella (Treitschke) 
schisticolor Zeller 
villosella Hulst 
rubribasella Hulst 
GuyprTocera Ragonot 
208. consobrinella (Zeller): 


U.S. (florida), Tropical 


Southern 


new comb.: 


Europe, Asia, 


Eastern U. S., Can- 


ada 
busckella (Dyar) 

Pima Hulst 

209. boisduvaliella (Guénée), new comb.: EKu- 
rope, Canada 

210. albiplagiatella (Packard), new comb.: EHast- 
ern U. S., Canada 

211. albiplagiatella occidentalis Heinrich, new 
race: Western U.S. 

212. fosterella Hulst: Western U.S., Canada 

213. vividella (McDunnough), new comb.: Can- 
ada 

214. albocostalialis (Hulst), new comb.: Western 
U.S., Canada 

215. albocostalialis subcostella (Ragonot), new 
comb.: Southwestern U.S. 

216. fulvirugella (Ragonot,) Newcomb.: Western 
U.S. (California) 

217. granitella (Ragonot), new comb.: Western 


piperella (Dyar) 


56. 


57. 


58. 


59. 
60. 


61. 


62. 


63. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


218. parkerella (Schaus), new comb.: Western 
U.S. (Montana) 

InterRsEctiIo Heinrich, new genus 

219. denticulella (Ragonot), new comb.: North- 


western U.S., Canada 

220. columbiella (McDunnough), new comb.: 
Northwestern U.S., Canada 

221. ruderella (Ragnot), new comb.: ‘‘N. 
Amer.”’ (California?) 

222. niviella (Hulst) new comb.: 

Ambesa Grote 

223. laetella Grote: Western U. S., Canada 

224. walsinghami (Ragonot): Western U. S. 

monodon Dyar 

225. walsinghami mirabella Dyar, new status: 
U.S. (Southern California) 

226. lallatalis (Hulst): Western U. S. (Nevada, 
Utah) 


U.S., Canada 


Carastia Hiibner 

227. bistriatella (Hulst), new comb.: Western 
U.S. (California) 

228. incorruscella (Hulst), new comb.: Western 


U.S. (California) 
229. actualis (Hulst), new comb.: 
Canada 
Immyrua Dyar 
230. nigrovittella Dyar: 
Oreana Hulst 
231. unicolorella (Hulst): Eastern U. S., Canada 
leucophaeella (Hulst) 
Otysrta Heinrich 
232. aliculella (Hulst), new comb.: 
U.S 


Western U.S., 


Eastern U. S. 


Southwestern 


oberthuriella (Ragonot) 
233. furciferella (Dyar) new comb.: 
ern U.S. (Arizona) 
SALEBRIACUS Heinrich, new genus 
234. odiosellus (Hulst), new comb.: 
U.S. 
bakerella (Dyar) 
yumaella (Dyar) 
SaLesBriaria Heinrich, new genus 


Southwest- 


Western 


235. turpidella (Ragonot), new comb.: Southern 
U.S. 
ademptandella (Dyar) 
236. nubiferella (Ragonot), new comb.: U.S. 
237. engeli (Dyar) U.S. 
238. annulosella (Ragonot), new comb.: U. S. 
(Texas, North Carolina) 
robustella (Dyar) 
239. tenebrosella (Hulst), new comb.: U.S. 
quercicolella (Ragonot) 
heinrichalis (Dyar) 
240. pumilella (Ragonot) new comb.: Southeast- 


ern U.S. 
georgiella (Hulst) 
241. fructetella (Hulst) new comb.: 
rectistrigella (Dyar) 


U.S. 


300329—56——22 


64. 


65. 


66. 


67. 


68. 


321 


QUASISALEBRIA Henrich, new genus 

242. admixta Heinrich, new species: 
LOTS 

OrtTHOLEPIS Ragonot 

243. jugosella Ragonot: Eastern U. S., Canada 

244, pasadamia (Dyar), new comb.: U. S., Can- 
ada 

Poxoprustis Ragonot 

245. arctiella (Gibson): 

Merropterra Grote 

Emmerita Hampson 

246. mirandella Ragonot: Western U.S. 

247. cviatella Dyar: U.S. (Illinois, Mississippi) 

248. pravella (Grote): U.S., Canada 


Western 


Alaska, Canada 


249. abditiva Heinrich, new species: Canada 
Nepuorteryx Hiibner 
250. subfuscella (Ragonot), new comb.: Eastern 


U.S., Canada 
semiobscurella (Hulst) 
251. delassalis Hulst: Western U. S. 
purpurella (Hulst) 
pudibundella (Ragonot) 


252. delassalis fraudifera Heinrich, new race: 
Canada (British Columbia), U. S.(Wash- 
ington) 

253. rubescentella (Hulst): U.S. 

254. fernaldi (Ragonot), new comb.: U. S., 
Canada 

255. dammersi Heinrich, new species: Western 


U.S. (California, Arizona) 
dammersi floridensis Heinrich, new race: 
U.S. (Florida) 
vetustella (Dyar), new comb.: 
ada 
inconditella (Ragonot), new comb.: 
ern U.S. (Arizona, Colorado) 
subcaesiella (Clemens), new comb.: 
Canada 
contatella (Grote) 
virgatella (Clemens), new comb.: 
Canada 
quinquepunctella (Grote) 
carneella Hulst: U.S., Canada 
inquilinella (Ragonot) 
basilaris Zeller: U.S., Canada 
termitalis (Hulst), new comb.: 
U.S., Canada 
levigatella (Hulst) 
termitalis yuconella 
Alaska 
bifasciella Hulst: U.S. (Arizona) 
nogalesella (Dyar) 


256. 
257. U. S., Can- 
258. West- 


259. U.S, 


260. UESS:, 


261. 


262. 


263. Western 


264. Dyar, new status: 


265. 


266. uvinella (Ragonot), new comb.: Eastern 
afflictella (Hulst) 
liquidambarella (Dyar) 

267. celtidella (Hulst), new comb.: U.S. 


322 


69. 


70. 


72. 


73. 


74. 


75. 


76. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


268. rubrisparsella (Ragonot): U.S. 
rufibasella (Ragonot) 
croceella (Hulst) 
texanella (Hulst) 
269. gilvibasella Hulst: 
lacteella (Hulst) 
270. crassifasciella Ragonot: 
decupientella Dyar 
crataegella B. and McD. 
271. bisra Dyar: México 
TiascaLa Hulst 
272. reductella (Walker): Eastern U. S. 
gleditschiella (Fernald) 
Tusa Heinrich, new genus 
273. finitella (Walker), new comb.: 
Canada 
melanellus (Hulst) 
274. umbripennis (Hulst), new comb:: 
(Colorado) 
gillettella (Dyar) 
275. oregonella. (Barnes and McDunnough), new 
comb.: U.S. (Oregon) 
infinitella (Dyar), new comb.: 


U.S. (Texas) 
Eastern U.S. 


Eastern U.S., 


UW. ©: 


276. México 


. Homorocraprua Ragonot 


277. lanceolella Ragonot: Pert 
TELETHuUsIA Heinrich, new genus 
278. ovalis (Packard), new comb.: 
latifasciatella (Packard) 
geminipunctella (Ragonot) 
modestella (Hulst) 
279. rhypodella (Hulst), new comb.: 
gon’’) 
Pxosus Heinrich, new genus 
280. brucei (Hulst), new comb.: 
281. funerellus (Dyar), new comb.: 
Canada 
curvatellus (Ragonot), new comb.: 
U.S. 
incertus Heinrich, new species: 
U.S. (California) 
Actrix Heinrich, new genus 
284. nyssaecolella (Dyar), new comb.: 


U.S., Canada 


U.S. (“Ore- 


Western U. S. 
Western U.S., 


282. Western 


283. Western 


Hastern 
285. dissimulatrix Heinrich, new species: Eastern 
U.S. (Virginia) 
SryLopaLpia Hampson 
286. lunigerella Hampson: West Indies, México 
287. scobiella (Grote), new comb.: U.S. (Texas, 
Colorado) 
decimerella (Hulst) 


288. argentinensis Heinrich, new species: Ar- 
gentina 

Pyta Grote 

289. fasciolalis (Hulst), new comb.: Canada 
(British Columbia) 

290. impostor Heinrich, new species: Western 
U.S., Canada 

291. aequivoca Heinrich, new species: Western 
Canada 


77. 


292. insinuatrix Heinrich, new species: Canada 
(Manitoba) 

293. aenigmatica Heinrich, new species: U. S., 
Canada 


294. 
295. 


criddlella Dyar: Canada (Manitoba) 
fusca (Haworth), new comb.: Holarctic 
moestella (Walker) 
frigidella (Packard) 
cacabella (Hulst) 
triplagiatella (Dyar) 
hypochalciella (Ragonot), new comb.: North- 
western U. S., Canada. 
blackmorella (Dyar) 


296. 


297. hanhamella Dyar: Canada (Manitoba) 

298. scintillans (Grote): Western U. S. (Califor- 
nia) 

feella Dyar 

299. sylphiella Dyar: Northwestern U. S., Canada 

300. rainierella Dyar: Northwestern U.S. (Wash- 
ington) 

301. aeneella Hulst: Western U. S. (Colorado, 
Utah) 

302. aeneoviridella Ragonot: Western U. S., 
Canada 

303. metalicella Hulst: Western U.S. (Colorado, 
Utah) 

304. fasciella Barnes and McDunnough: North- 


western U.S. (California) 
nigricula Heinrich, new species: 
U.S. (Nevada) 
viridisuffusella Barnes and McDunnough: 
Western U.S. (California) 
Diorycrria Zeller 
Pinipestis Grote 
307. abietella (Denis and Schiffermiiller): 
ern Hemisphere 
decuriella (Hisbner) 
abietivorella (Grote) 
elegantella (Hulst) 
sysstratiotes Dyar: 
reniculella (Grote): 
ponderosae Dyar: 
California) 
majorella Dyar: México 
muellerana Dyar 
disclusa Heinrich: Hastern U. S. 
auranticella (Grote): Western U.S., Canada 
miniatella Ragonot 
zanthaenobares Dyar 
erythropasa (Dyar): 
(Arizona) 
315. horneana (Dyar): 


305. Western 


306. 


North- 


308. 
309. 
310. 


Guatemala 

Northern U.S., Canada 
Western U.S. (Montana, 
311. 
312. 
313. 


314. Southwestern U. S. 


Cuba 


316. pygmaeella Ragonot: Hastern U.S. 
317. zimmermani (Grote): U.S., Canada 
delectella (Hulst) 
austriana (Cosens) 
318. cambiicola (Dyar): Western U.S. 
319. amatella (Hulst): Eastern U.S. 
320. albovittella (Hulst): Western U.S. 


78. 


80. 


81. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


321. gulosella (Hulst), new comb.: Western U.S. 


(Colorado, New Mexico) 


322. baumhoferi Heinrich, new species: South- 
western U.S. (Arizona) 
323. subtracta Heinrich, new species: South- 


western U. S. (New Mexico) 
clarioralis (Walker): Eastern U.S. 
brunneella (Dyar) 
Orycrometoria Ragonot 
325. fossulatella Ragonot: 
ical America 
moeschleri (Ragonot) 


324. 


U. S. (Texas), Trop- 


. Saravta Ragonot 


326. edwardsialis (Hulst), new comb.: Western 
AISE 
polyphemella (Ragonot) 
327. pullatella (Ragonot), new comb.: Western 
U.S. 
punctella (Dyar), new comb.: México 
punctella septentrionaria Heinrich, new race: 
_ Western U. S. 
incanella (Hulst), new comb.: 
aridella (Dyar) 


328. 
329. 


330. Western U.S. 


331. atrella (Hulst), new comb.: Western U. S. 
(Colorado) 

332. caudellella (Dyar), new comb.: Western 
U.S., Canada 

333. dnopherella Ragonot: Western U. S. (Cali- 
fornia) 

334. nigrifasciella Ragonot: Western U.S., Can- 
ada 

335. cinereella Hulst: Western U. S. (Colorado) 


336. rubrithoracella (Barnes and McDunnough), 


new comb.: Western U.S. 

337. tephrella Ragonot: Western U. S. (Wash- 
ington) 

338. alpha Heinrich, new species: Canada (Sas- 
katchewan) 

339. beta Heinrich, new species: Western U. S., 
Canada 

340. gamma Heinrich, new species: Western 
U.S. (California) 

341. iota Heinrich, new species: Western U. S. 


(California) 
perfuscalis (Hulst): 
excantalis (Hulst) 
epsilon Heinrich, new species: Western U.S. 
phi Heinrich, new species: Western U. S. 


342. Western U. S. 


343. 
344, 


345. kappa Heinrich, new species: Western U.S. 
(Arizona) 

346. delta Heinrich, new species: Western U. S. 

PuitopEMa Heinrich, new genus 

347. rhoiella (Dyar), new comb.: Western U. S. 


Lipoerapuis Ragonot 
348. fenestrella (Packard): 
fornia) 

humilis Ragonot 
349. leoninella (Packard): 
ada 
pallidella (Dyar) 


Western U. S. (Cali- 


Western U. S., Can- 


82. 


83. 


84. 


85. 


86. 


87. 


88. 


323 
350. truncatella (Wright), new comb.: South- 
western U.S. (California) 
351. umbrella (Dyar), new comb.: 
U.S. (California) 
352. subosseella Hulst: Bahamas 
AveE.puiA Heinrich, new genus 
353. petrella (Zeller), new comb.: 
rubiginella (Walker) 
rufinalis (Walker) 
hapsella (Hulst) 
354. ochripunctella (Dyar), new comb.: 
U.S. (California) 
Tora Heinrich, new genus 
355. galdinella (Schaus), new comb.: 
Islands 
Ura Walker 
356. lithosella (Ragonot), new comb.: 
western U. S., México 
luteella Hulst) 
roseitinctella (Dyar), new comb.: 
western U. S., México 
senta Heinrich, new species: 
U.S. (Texas, Arizona) 
rubedinella (Zeller), new comb.: 
ida), Tropical America 
translucida (Walker) 
rufescentalis (Walker) 
minualis (Walker) 
deprivalis (Walker) 
venezuelalis Walker 
pyrrhochrellus (Ragonot) 
Exasmopaupus Blanchard 
360. lignosellus (Zeller): U.S., Tropical America 
angustellus Blanchard 
tartarella (Zeller) 
incautella (Zeller) 
major (Zeller) 
anthracellus Ragonot 
carbonella (Hulst) 
puer Dyar 
Acroncosa Barnes and McDunnough 
361. albiflavella Barnes and McDunnough: West- 
ern U. S. (California) 
362. albiflavella castrella Barnes and McDun- 
nough: Western U.S. (New Mexico) 
363. similella Barnes and McDunnough: West- 
ern U.S. (Nevada, Utah) 
PassapDENA Hulst 
364. flavidorsella (Ragonot): 
México 
canescentella (Hulst) 
constantella Hulst 
cinctella (Hulst) 
Utopnora Ragonot 
Acromeseres Dyar 
365. groteii Ragonot: Eastern U.S. 
tephrosiella Dyar 
366. guarinella (Zeller): 
dialithus (Dyar) 


Southwestern 


U.S. 


Western 


Galdpagos 


South- 


357. South- 


358. Southwestern 


359. U.S. (Flor- 


Western U. S., 


Cuba, Colombia 


324 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


89. CHorrEeRA Dyar 
367. idiotes Dyar: Panama 
368. extrincica (Dyar), new comb.: 
369. postica (Zeller), new comb.: 
90. Tacoma Hulst 
370. feriella Hulst: Southwestern U.S. 
submedianella Dyar 
91. ApELPERGA Heinrich, new genus 
371. cordubensiella (Ragonot), new comb.: Ar- 
gentina 
92. Humysta Dyar 
372. mysiella (Dyar): 
373. maidella (Dyar): 


Cuba 
Colombia 


Western U. S. 
Western U. S., Canada 


374. pallidipennella (Hulst), new comb.: West- 
ern U.S. 

375. fuscatella (Hulst): Western U. S. (Cali- 
fornia) 


376. semicana Heinrich, new species: Western 
U. S. (Washington) 
93. Drvrt1aca Barnes and McDunnough 
377. ochrella Barnes and McDunnough: South- 
ern U.S. (Florida) 
378. simulella Barnes and McDunnough: South- 
ern U.S. (Florida) 
379. parvulella Barnes and McDunnough: South- 
ern U.S. (Florida) 
380. parvulella consociata Heinrich, new race: 
Colombia 
94. MacrorrHInia Ragonot 
Dolichorrhinia Ragonot 
381. aureofasciella Ragonot: Southwestern U.S., 
México 
382. placidella (Zeller): 
95. Ocata Hulst 
383. dryadella Hulst: Southern U.S. (Florida) 
platanella (Grossbeck) 


96. Vauprvia Ragonot 
Maricopa Hulst 
384. coquimbella Ragonot: 
385. lativittella (Ragonot): 
México 
aureomaculella (Dyar) 
386. walkerella (Ragonot), new comb.: 


97. Prorasta Heinrich, new genus 
387. mirabilicornella (Dyar), new comb.: 
ern U.S. (California) 
98. HreTrRoGRAPHis Ragonot 
Mona Hulst 
388. morrisonella Ragonot: 
coloradensis Ragonot 
olbiella (Hulst) 
ignistrigella Ragonot 
palloricostella (Walter) 
99. StauDINGERIA Ragonot 
389. albipenella (Hulst): 
olivacella Dyar 
perluteella Dyar 


Brazil 


Chile 
Southwestern U. S., 


Chile 


West- 


U.S., México 


Western U. S. 


100. Huustia Ragonot 
390. undulatella (Clemens): 
rubiginalis (Walker) 
obsipella (Hulst) 
fumosella (Aulst) 
101. Honora Grote 
391. mellinella Grote: U.S. 
ochrimaculella Ragonot 
392. subsciurella Ragonot: Western U.S. 
393. sciurella Ragonot: Western U. S. (Cali- 
fornia) 
394. dotella Dyar: Western U.S. (California) 
395. montinatatella (Hulst): Western U. S. 
canicostella Ragonot 
396. perdubiella (Dyar), new comb.: Western 
U.S. 


U.S., Canada 


102. Honorinus Heinrich, new genus 
397. fuliginosus Heinrich, new species: 
103. Oncoxasis Zeller 
Endommasis Hampson 
398. anticella Zeller: Tropical America 
migritella (Hampson) 
104. Casotia Ragonot 
Encystia Hampson 
399. semidiscella Ragonot: Argentina 
400. schini (Berg): Argentina 
401. rhythmatica Dyar: Panamé 
402. cundajensis (Zeller): Colombia 
impeditella (Zeller) 
403. bonhoti (Hampson), new comb.: 
Jamaica 
105. Canarsta Hulst 
404. ulmiarrosorella (Clemens): 
pneumatella (Hulst) 
ulmella (Ragonot) 
fuscatella (Hulst) 
gracilella Hulst 
feliculella Dyar 
106. Harnocua Dyar 
405. velessa Dyar: 
107. Eurytamasis Dyar 
406. ignifatua Dyar: 
Cuba 
108. Euryramipia Ragonot 
407. ignidorsella (Ragonot): 
México, Panamé 
109. Wunper1A Grossbeck 
408. neaeriatella Grossbeck: U.S. (Florida) 
110. OzpotTHm1a Hampson 
Synothmia Hampson 
409. endopyrella Hampson: México, Bahamas 
bahamasella (Hampson) 
111. StynoBasis Hampson 
410. rubripurpurea Hampson: México, Brazil 
112. Diviana Ragonot 
Dannemora Hulst 
411. eudoreella Ragonot: 
edentella (Hulst) 


Peri 


Bahamas, 


U.S., Canada 


Panamdé 


Panamé, Puerto Rico, 


U. S. (Arizona), 


Eastern U. S. 


113. 


114. 


115. 


116. 


117. 


118. 


119. 


120. 


121. 


122. 


123. 


124. 


125. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


PanatKa Hulst 

412. nymphaeella (Hulst): Eastern U. S. 
verecuntella (Grossbeck) 

CacozopHEera Dyar 


413. venosa Dyar: Guatemala 
Psorosina Dyar 
414. hammondi (Riley): Eastern and Central 


U.S., Canada 
angulella Dyar 
Patricroua Heinrich, new genus 
415. semicana Heinrich, new species: Utah 
Pacontus Heinrich, new genus 
416. corniculatus Heinrich, new species: 
Rico 
Aptunea Heinrich, new genus 
417. macropasa (Dyar), new comb.: 
México 
418. imperfecta (Dyar), new comb.: 
ANDERIDA Heinrich, new genus 
419. sonorella (Ragonot), new comb.: 
U.S. (Arizona) 
placidella (Dyar) 
Casstana Heinrich, new genus 
420. malacella (Dyar), new comb.: 
Puerto Rico, Virgin Islands 
Mescinra Ragonot 
421. triloses Dyar: 
mosces Dyar 
pandessa Dyar: Guatemala 
bacerella Dyar: Cuba 
estrella Barnes and McDunnough: U. S. 
(Florida) 
moorei Heinrich, new species: British Guiana 
parvula (Zeller): Colombia 
commatella (Zeller): Colombia 
berosa Dyar: Panam4, Puerto Rico 
peruella Schaus: Pert 
430. discella Hampson: México, Guatemala 
431. indecora Dyar: México 
Nona Ragonot 
Hypermescinia Dyar 
432. exiguella (Ragonot): 
lambella (Dyar) 
Puestinia Hampson 
433. costella Hampson: 
Comotta Dyar 
434. torsicornis Dyar: Panamé 
435. convergens (Dyar), new comb.: 
Bema Dyar 
Relmis Dyar 
neuricella (Zeller), new comb.: 
America 
myja Dyar 
437. fritilla Dyar: Guatemala 
438. ydda (Dyar), new comb.: 
Guiana 
439. yddiopsis (Dyar), new comb.: Cuba 
440. fifaca (Dyar), new comb.: Panama 


Puerto 


Guatemala, 
Guatemala 


México, 


México, 


Panam& 
422. 
423. 
424. 
425, 
426. 
427. 


428. 
429. 


Tropical America 


Jamaica, Puerto Rico 


Guatemala 


436. Tropical 


Panamd, French 


126. 


127. 


128. 


129. 


130. 


325 


Homogosoma Curtis 
Phycidea Zeller 
electellum (Hulst): U. S., México, Guate- 
mala, Cuba, British West Indies 
opalescellum (Hulst) 
texanellum Ragonot 
tenuipunctella Ragonot 
differtella Barnes and McDunnough 
. stypticellum Grote: U.S., Canada 
uncanale Hulst 
. striatellum Dyar: Southwestern U. S. 
. oslarellum Dyar: Western U. S. 
. oslarellum breviplicitum Heinrich, new race: 
Southwestern U. S. (California) 
. illuviellum Ragonot: U.S. (Arizona, Colo- 
rado), México 
candidella Hulst 
. iluviellum emendator Heinrich, new race: 
Western U.S. 
. Imitator Heinrich, new species: Southwestern 
U.S. (California) 
. longiventrellum Ragonot: 
noctividella Ragonot 
. albescentellum Ragonot: Western U.S. 
elongellum Dyar 
. impressale Hulst: Western U. S., Canada 
. inornatellum (Hulst): Eastern U.S. 
. deceptorium Heinrich, new species: U. S. 
(Pennsylvania), Canada 
. discrebile Heinrich, new species, Brazil 
. peregrinum Heinrich, new species: 
(California), Costa Rica 
. vepallidum Heinrich, new species: Argentina 
. ditaeniatellum Ragonot: Chile 
. oconequensis (Dyar), new comb.: Pert 
. assitum Heinrich, new species: Pert 
. acmacopterum Ragonot: Chile 
. nimbosellum Ragonot: Chile 
462. unionellum Ragonot: México 
Pataconia Ragonot 
463. magellanella (Ragonot): 
Rorrupa Heinrich, new genus 
464. mucidella mucidella (Ragonot), new comb.: 
Western U. S., Canada 
465. mucidella reliquella (Dyar), new comb.: 
Eastern U. S., Canada 
466. mucidella olivaceela (Ragonot), new comb.: 
Tropical America 
musiosum (Dyar) 
cubella (Dyar) 
467. mucidella affusella (Ragonot), new comb.: 
Argentina 
STREPHOMESCINIA Dyar 
468. schausella Dyar: 
UnapiLua Hulst 
Strymaz Dyar 


441. 


Chile 


De Ss 


Chile 


Cuba 


326 


131. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


469. erronella (Zeller): Tropical America. 
ubacensis (Zeller) 
bipunctella (Hampson) 
dorae (Dyar) 
pyllis (Dyar) 
470. maturella (Zeller): Colombia, Guatemala, 
Cuba 
471. albidiorella (Richards and Thomason); new 
comb.: Pert 
472. floridensis Heinrich, new species: U.S. (Flor- 
ida) 
473. nasutella Hulst: U.S. (New Mexico) 


Laetitia Ragonot 
Laosticha Hulst 

474. coccidivora, (Comstock): 
pallida (Comstock) 
dilatifasciella (Ragonot) 
hulstii Cockerell 


WS. 


475. coccidivora quadricolorella (Dyar), new 
comb.: Southwestern U. S. 

476. coccidivora cardini Dyar: Cuba, U. S. 
(Florida) 

477. obscura Dyar: Cuba 

478. portoricensis Dyar: Puerto Rico 

479. melanostathma (Meyrick), new comb.: Ar- 


dl gentina 


480. amphimetra (Meyrick), new comb.: Argen- 
tina 
481. zamacrella Dyar: Western U.S. (California) 
482. myersella Dyar: Eastern U.S. 
483. ephestiella (Ragonot): Southwestern U. S. 
(Arizona) 
lustrella (Dyar) 
484. fiskella Dyar: Eastern U. S. (North Caro- 


lina) 


485. glomis (Dyar), new comb.: Panamé 
132.3BarHaa Heinrich, new genus 
486. basimaculatella (Ragonot), new comb.: 


133. 


Western U.S. 
eremella (Dyar) 
487. goyensis (Ragonot), new comb.: 
Uruguay, Argentina 
488. goyensis olivacea Heinrich, new race: Argen- 
tina 
489. homoeosomella, (Zeller), new comb.: 
ical America, 
bodkini (Dyar) 
rusto (Dyar) 
taboga (Dyar) 
saissetiae (Dyar) 
490. haywardi Heinrich, new species: 
491. glabrella (Dyar), new comb.: 
492. squalida (Walker), new comb.: 
Ruacea Heinrich, new genus 
493. packardella (Ragonot), new comb.: 
ern U.S. 
orobanchella (Dyar) 
494, stigmella (Dyar), new comb.: 
U.S. (California), México 
maculicula (Dyar) 


Brazil, 


Trop- 


Argentina 
Guatemala 
Brazil 


West- 


Southwestern 


134. ZorHopia Hubner 


136. 


137. 


138. 


139. 


140. 


141. 


143. 


Dakruma Grote 

495. convolutella (Hiibner): 
ada 

grossulariella (Hiibner) 

turbatella (Grote) 

grossularvae (Riley) 

franconiella (Hulst) 

bella Hulst 

thouna Dyar 

dilativitta Dyar 

magmyicans Dyar 


Europe, U.S., Can- 


. Menitara Walker 


496. prodenialis Walker: U.S. 
bolliz (Zeller) 
497. dentata (Grote): U.S. 


doddalis Dyar 
OtycEeLLa Dyar 
498. junctolineella (Hulst): Southern U.S. (Texas) 
499. junctolineella pectinatella (Hampson): Méx- 
ico 
500. nephelepasa (Dyar:) México 


501. subumbrella Dyar: Western U.S. 

Ouyca Walker 

502. phryganoides Walker: Dominican Repub- 
lic, Haiti 


ALBERADA Heinrich 
503. parabates (Dyar): 
504. bidentella (Dyar): 
(Texas, Arizona) 
505. holochlora (Dyar): 
(Texas) 
Nawata Heinrich 
506. substituta Heinrich: Pert 
Cactosiastis Ragonot 
Neopyralis Bréthes 
cactorum (Berg): Argentina, Uruguay, Aus- 
tralia 
. ronnai (Bréthes): Brazil 
. doddi Heinrich: Argentina 
. mundelli Heinrich: Pert 
511. bucyrus Dyar: Argentina 
CaHELa Heinrich 
512. ponderosella (Barnes and McDunnough): 
Western U. S., México 
purgatoria (Dyar) 
interstitialis (Dyar) 
phoenicis (Dyar) 


U.S., México 
Southwestern U. S. 


Southwestern U. S. 


507. 


. Rumatua Heinrich 


Southern U. S. 
Western U.S. 
Southwestern U.S. (Ari- 


513. glaucatella (Hulst): 
514. bihinda (Dyar): 
515. polingella (Dyar): 
zona, Texas) 
Yosrmitia Ragonot 
516. graciella (Hulst): Western U.S. 
517. longipennella (Hulst): Southwestern U. S. 
(Texas) 
518. fieldiella (Dyar): 
Arizona) 
519. didactica Dyar: 


Western U.S. (California, 


México 


144, 


145. 


146. 


147. 


148. 


149. 


150. 


151. 


152. 


156. 


157. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Tucumania Dyar 
520. tapiacola Dyar: 
521. porrecta Dyar: 
EremsBerca Heinrich 
522. leuconips (Dyar): 
523. creabates (Dyar): 
fornia) 
524. insignis Heinrich: México 
SatamBona Heinrich 
525. analamprella (Dyar): 
Parotyca Dyar 
526. asthenosoma (Dyar): 
Sigeveaira Heinrich 
527. chilensis Heinrich: Chile 
528. huanucensis Heinrich: Pert 
529. transilis Heinrich: Pert 
AMALAFRIDA Heinrich 
530. leithella (Dyar): 
Colombia 
Ozamia Ragonot 
531. lucidalis (Walker): West Indies 
532. fuscomaculella (Wright): Southwestern 
U.S. (California) 
heliophila Dyar 
fuscomaculella clarefacta Dyar:  U. S. 
(Texas), México 
thalassophila Dyar: U.S. (California) 
immorella (Dyar), new comb.: México 
stigmaferella Dyar: Argentina 
537. hemilutella Dyar: Argentina 
538. punicans Heinrich: Argentina 
Cacrosrosis Dyar 
539. fernaldialis (Hulst): Southwestern U. S. 
gigantella (Ragonot) 
cinerella (Hulst) 
longipennella (Hampson): 
elongatella (Hampson): 
maculifera Dyar: México 
insignatella Dyar: México 
strigalis (Barnes and McDunnough): West- 
ern U. S., México 
Drescomopsis Dyar 
544. soraella (Druce): 
drucella (Dyar) 
subelisa Dyar 


Argentina 
Uruguay 


Western U.S. (Arizona) 
Western U. S. (Cali- 


Argentina 
French Guiana 


West Indies, Venezuela, 


533. 


534, 
535. 
536. 


540. México 


541. 


542. 
543. 


Tropical America 


. Intatiza Dyar 


545. gurbyris Dyar: Panamé 


. Lascrtina Heinrich, new genus 


546. canens Heinrich, new species: Southern 


U.S. (Texas), México 


. MerepuHestia Ragonot 


547. simplicula (Zeller): U.S. (Florida), Puerto 
Rico, Colombia, British West Indies 

Srxea Heinrich, new genus 

548. arizonella (Hulst), new comb.: 
ern U.S. (Arizona) 

Entmemacornis Dyar 

549. proselytes Dyar: Guatemala 

550. pulla Heinrich, new species: Brazil 


Southwest- 


158. 


159. 


160. 


161. 


162. 


163. 


164. 


165. 


166. 


167. 


168. 


169. 


170. 


327 


Cayrennia Hampson 


551. rufitinctalis Hampson: French Guiana 


Rioga, Heinrich, new genus 

552. nexa Heinrich, new species: Argentina 
Moerrses Dyar 

553. dryopella (Schaus): Costa Rica 


554. alveolella (Ragonot), new comb.: Brazil 
555. emendata Heinrich, new species: Panama, 
French Guiana 
Moopnopsis Dyar 
Campyloplesis Dyar 
556. decipiens Dyar: México 
557. perangusta (Dyar), new comb.: Trinidad 
558. inornatella (Ragonot), new comb.: Costa 
Rica, Brazil 
559. parallela Heinrich, new species: Brazil, Pera 
560. inveterella (Dyar), new comb.: Guatemala 
561. portoricensis Heinrich, new species: Puerto 


Rico 
Eputica Ragonot 
562. compedella (Zeller): 
KuzopHera Zeller 


Tropical America 


563. semifuneralis (Walker): U. S., Canada, 
México 
aglaeella Ragonot 
pallulella (Hulst) 
564. ostricolorella Hulst: Eastern U. S. 


565. nigricantella Ragonot: 
México 
griselda Dyar 
Exvurertivus Heinrich, new genus 
566. negator Heinrich, new species: 
Evutoera Heinrich, new genus 
567. ochrifrontella (Zeller), new comb.: 
Canada 
Serruginella (Ragonot) 
PROSOEUZOPHERA Heinrich, new genus 
568. impletella (Zeller), new comb.: 
Jamaica, Puerto Rico 
Farnosia Heinrich, new genus 
569. quadripuncta (Zeller), new comb.: Costa 
Rica, Panama, French Guiana, Colombia 
Gennapius Heinrich, new genus 
570. junctor Heinrich, new species: 
ana 
Micromescinia Dyar 
571. pygmaea Dyar: 
Epuestioprs Ragonot 
572. gilvescentella Ragonot: 
ada, México 
nigrella Hulst 
573. infimella Ragonot: 
574. erythrella Ragonot: 
coloradella (Hulst) 
benjaminella Dyar 
mignonella Dyar: U.S. (Texas) 
erasa Heinrich, new species: U.S. (Florida) 
lucidibasella Ragonot: Chile 
productella Ragonot: Colombia (?) 
indentella Dyar: Bermuda 


Southwestern U. S., 


Pert 


U. S., 


Colombia, 


French Gui- 


Panama 


Western U.S., Can- 


Eastern U.S. 
Western U.S., Canada 


575. 
576. 
577. 
578. 
579. 


328 


171. 


172. 


173. 


174. 


175. 


176. 


177. 


178. 


179. 


180. 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


580. plorella Dyar: 
vestilla (Dyar) 
581. stictella (Hampson), new comb.: 
West Indies 
uniformella Hampson 
granulella Hampson 
582. noniella Dyar: Panam4 
AZAERA Schaus 
Calamophleps Dyar 
583. muciella Schaus: Costa Rica, Guatemala, 
Panama 
squalidella (Dyar) 
584. nodoses (Dyar), new comb.: Panam’ 
585. lophophera (Dyar), new comb.: Panamé 
Moopna Hulst 
586. ostrinella (Clemens): U.S., Canada 
obtusangulella (Ragonot) 
pelviculella Hulst 
587. bisinuella Hampson: 
Viruta Ragonot 
588. edmandsae (Packard): 
ada 
dentosella Ragonot 


Panama 


Bahamas, 


México, U.S. (Texas) 
Eastern U. S., Can- 


589. edmandsae serratilineella Ragonot, new sta- 
tus: Western U.S., Canada 
590. lugubrella Ragonot, new comb.: Western 
U.S. (California) 
591. pinei Heinrich, new species: Western U. S. 
(Utah, Nevada) 
592. inanimella (Dyar), new comb.: México, 
Guatemala 
ticitoa (Dyar) 
593. laura (Dyar), new comb.: Guatemala 


Manuatta Hulst 
Hornigia Ragonot 
594. setonella (McDunnough), new comb.: U.S. 
(Utah), Canada (British Columbia) 
595. broweri Heinrich, new species: Hastern U.S. 
(Maine) 
Verina Heinrich, new genus 


596. supplicella (Dyar), new comb.: México, 
Guatemala, Panam4, Brazil 

Vacozsanta Heinrich, new genus 

597. divergens (Butler), new comb.: Chile 

Moopnetua Heinrich, new genus 

598. paula Heinrich, new species: Guatemala, 


Brazil, Argentina 

Vouatica Heinrich, new genus 

599. pachytaeniella (Ragonot), new comb.: Bra- 
zil 

600. trinitatis Heinrich, new species: Trinidad 

Vezina Heinrich, new genus 

601. parasitaria Heinrich, new species: 
tina, Brazil 

CaubELui1a Dyar 

602. apyrella Dyar: Eastern U. S. (Maryland) 

603. albovittella Dyar: Eastern U. S. 

604. nigrella (Hulst), new comb.: Western U.S. 

arizonella (Walter) 


Argen- 


181. 


182. 


183. 


184. 


185. 


186. 


187. 


605. declivella (Zeller), new comb.: Panama, 
Colombia 
animosella (Dyar) 
606. colorella (Dyar), new comb.: Panam& 


607. clara Heinrich, new species: Puerto Rico 

MicropHestia Dyar 

608. animalcula Dyar: Panama 

SosipaTRa Heinrich, new genus 

609. rileyella (Ragonot), new comb.: Western 

U.S., México 
micaceella (Hampson): México 
anthophila (Dyar), new comb.: 
U.S. (Texas) 
thurberiae (Dyar), new comb.: Western U. S. 
nonparilella (Dyar), new comb.: Western 
U.S. (Arizona) 

614. majorella (Dyar), new comb.: 

615. divergens (Dyar): Panamé 

Beruutia Ragonot 

616. championella Ragonot: 

Risua Heinrich 

617. mnoxia Heinrich: Cuba 

618. contigua Heinrich, new species: 

619. patriciella (Dyar), new comb.: 

Prop1a Guénée 

620. interpunctella (Hiibner): 

interpunctalis (Hiibner) 
zeae (Fitch) 

latercula (Hampson) 
glycinwwora (Matsumura) 

621. dolorosa Dyar: Guatemala, 

Awacasta Heinrich, new genus 

622. kiihniella (Zeller): Cosmopolitan 

fuscofasciella (Ragonot) 
gitonella Druce 

Epuestia Guénée 

Hyphantidium Scott 
623. elutella (Hitibner): Cosmopolitan 
elutea (Haworth) 
semirufa (Haworth) 
rufa (Haworth) 
sericarium (Scott) 
roxburghi Gregson 
unicolorella Staudinger 
amarella Dyar 

624. cautella (Walker): 

defectella (Walker) 
desuetella (Walker) 

cahiritella Zeller 

passulella Barrett 

formosella (Wileman and South) 

625. figulilella Gregson: Europe, Asia, Africa, 
Hawaii, Australia, North America (U.S., 
California), South America 

jficulella Barrett 
milleri Zeller 
jiguliella Forbes 
jigulella Curran 
venosella Turati 
ernestinella 'Turati 


610. 


611. Western 


612. 
613. 


México 
Guatemala 
Puerto Rico 


Cuba 


Cosmopolitan 


Cosmopolitan 


188. 


189. 


190. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


NIcETIODES Schaus 
626. apianella Schaus: 


Varneria Dyar 


Galfpagos Islands 


627. postremella Dyar: Eastern U. S. 
628. nannodes Dyar: Panam& 
629. atrifasciella Barnes and McDunnough: 


Southern U. S. (Florida) 
630. dubia Heinrich, new species: 
Evrytumia Ragonot 
631. hospitella (Zeller): Southern and Western 
U.S. 


Puerto Rico 


spaldingella Dyar 
632. hospitella yavapaella Dyar, new status: 
Western U.S. 
633. angulella Ely: Eastern U. S., Canada 
diffusella Ely 


634. fumella Ely: Eastern U.S. (Connecticut) 


191. 


192. 
193. 


194. 


329 
Ereieva Heinrich, new genus 
635. quantulella (Hulst), new comb.: Southern 
U.S. (Texas), West Indies 

santiagella (Dyar) 
636. coca (Dyar), new comb.:~ Panam4 

coquilla (Dyar) 

mossa (Dyar) 

uncta (Dyar) 
637. parvulella (Ely), new comb.: Eastern U. S. 


(Connecticut) 
Casnia Dyar 


638. myronella Dyar: Eastern U. S. 
Micropuycira Dyar 
639. titillella Dyar: Panama 


Rasreia Heinrich, new genus 


640. conops (Dyar), new comb.: Panam&é 


Species unplaced or unrecognized 


brevistrigella Ragonot [Zophodia] 
came Dyar [Huzophera] 
cervinistrigalis Walker [Hypochalcia] 
clitellatella Ragonot [Hornigia] 
corrientellus Ragonot [Elasmopalpus] 
daedalella Ragonot [Euzophera] 
disticta Zeller [Psorosa] 

dulciella Hulst [Honora] 

Jamula Zeller [Myelois] 

flavicornella Ragonot [Phycitopsis] 
formulella Schaus [Moodna] 
Suscifrontella Zeller [Nephopteryz] 


gais Dyar [Huzophera] 


grossipunctella Ragonot [Myelois] 
hulstiella Ragonot [Hypochalcia] 
infusella Zeller [Myelois] 
intextella Zeller [Huzophera] 
trichampa Dyar [Anthropteryz] 
megalopalis Hampson [Huzopherodes] 
migricans Hulst [Salebria] 

olwella Hampson [Moodna] 
postflavida Dyar [Huzophera] 
putidella Schaus [Hucampyla] 
rinmea Dyar [Huzophera] 
subcanella Zeller [Zophodia] 


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Figures 


The figures on the following pages are not drawn throughout to any fixed scale, the 
smaller genitalia being shown in greater enlargement than those of the larger species and the 
female genitalia on a uniformly smaller scale than the male genitalia. However, for the 
species of any given genus a uniform scale within the sexes has been attempted. 


The illustrations of venation in figures 1-53 and 55-130 are of males unless otherwise 
‘stated. The illustrations of male genitalia in figures 54, 131-637, and 915 show first under 
each figure, unless otherwise stated, a ventral view of the genitalia with one harpe and the 
aedeagus omitted, and this view is usually accompanied by a drawing of the aedeagus and 
other associated parts. Illustrations of female genitalia are shown in figures 638-914 and 
‘915-1138. For a few species, details of antennae and eyes have been added. 


331 


332 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 1-8.—VENATION (MALE UNLESS 


Oarwnr 


OOF 


. Cryptoblabes gnidiella (Milliére). 
. Cryptoblabes rutilella Zeller. 
. Acrobasis indigenella (Zeller). 


. Hypsipyla grandella (Zeller). 
. Fundella pellucens Zeller; 6a, underside of 


OTHERWISE NOTED). 


Sematoneura atrovenosella Ragonot. 


anal angle of hind wing with pocket un- 1.Cryptoblabes 

folded to show sex-scaling. Ie 2. Cryptoblabes 
Difundella corynophora Dyar. 
Birinus russeolus Heinrich, new species. 


4. Sematoneura 


6. Fundella 


5. Hypsipyla 


8. Birinus 


7. Difundella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 333 


Fieures 9—-16.—VENATION (MALE UNLESS 
OTHERWISE NOTED). 


9. Promylea lunigerella lunigerella Ragonot. 
10. Coptarthria dasypyga (Zeller). 
11. Anadelosemia senesciella (Schaus). 

12. Dasypyga alternosquamella Ragonot. 

13. Ceracanthia mamella (Dyar). 

14, Megarthria peterseni (Zeller). 
Coptarthria 15. Drescoma cyrdipsa Dyar. 


16. Drescomopsis soraella (Druce). 
Se 


11. Anadelosemia 


9. Promylea 


12. Dasypyga 


13. Ceracanthia 


16, Drescomopsis 


334 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 17-24.—VENATION (MALE UNLESS 
OTHERWISE NOTED). 


17. Etiella zinckenella (Treitschke). 

18. Ambesa laetella Grote. 

19. Immyrla nigrovittella Dyar. 

20. Salebria plumbella (Schiffermiller), type 
of the genus Salebria Zeller, not repre- 
sented in the American fauna. 

21. Emmerita mirandella (Ragonot). 

22. Meropiera pravella (Grote). 

23. Ortholepis jugosella Ragonot. 

24. Polopeustis annulatella (Zetterstedt). 


18.Ambesa 
17. Etiella 


20. 


Salebria 


Ortholepis 


22Meroptera Polopeustis 


AMERICAN MOTHS OF THE SUBFAMILY 


25: 
Nephopteryx 


26. 
Epischnia 


28. 


Tlascala 


27. Monoptilota 


30.Stylopalpia 


Lipographis 


PHYCITINAE 335 


Figures 25-32.—VENATION (MALE UNLESS 


25. 
26. 
27. 
28. 
29. 
30. 
31, 
32. 


OTHERWISE NOTED). 


Nephopteryz rhenella (Zincken). 

Epischnia prodromella (Hiibner). 

Monoptilota pergratialis (Hulst). 

Tlascala reductella (Walker). 

Pyla scintillans (Grote). 

Stylopalpia lunigerella Hampson. 

Lipographis fenestrella (Packard). 

Selagia argyrella (Schiffermiller). This 
genus occurs in our lists, but is not rep- 
resented in the American fauna. 


336 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 33-40.—VENATION (MALE UNLESS < 
OTHERWISE NOTED). 


33. Elasmopalpus lignosellus (Zeller). 

34. Chorrera idiotes Dyar. 

35. Acroncosa albiflavella Barnes and McDun- 
nough. 

36. Passadena flavidorsella (Ragonot). 

37. Tacoma feriella Hulst. 

38. Myelois cribrella (Hubner), an Old World 33. Elasmopal pus 34. Chorrera 
genus, not represented in the American 
fauna. 

39. Anypsipyla univitella Dyar. 

40. Apomyelois bistriatella (Hulst). 


36.Passadena 
35. Acroncosa 


4-0. Apomyelois 


AMERICAN MOTHS 


41. Pseudodivona 


SS  —— — 


S 


42. Diatomocera 


45. Rampylla 


47. Praedonula 


43. Cabima 


44. Hyalospila 


46. Davara 


48. Sarasota 


OF THE SUBFAMILY PHYCITINAE 337 


Figures 41-48.—VENATION (MALE UNLESS 
OTHERWISE NOTED) AND MALE GENITALIA. 


41. Pseudodivona commensella Dyar. Vena- 
tion. 

42. Diatomocera tenebricosa (Zeller). Vena- 
tion. 

43. Cabima dosia Dyar, venation, type of 
Cabima Dyar, asynonym of Diatomocera 
Ragonot. 

44, Hyalospila stictoneurella Ragonot. Vena- 
tion. 

45. Rampylla orio Dyar. Venation. 

46. Davara caricae (Dyar). Venation. 

47. Praedonula almonella (Dyar). Venation. 

48. Sarasota furculella (Dyar). Venation. 


338 


Figures 49-54.—VENATION (MALE UNLESS 


49. 


50. 


51. 


52. 
53. 


54. 


OTHERWISE NOTED). 


Zamagiria pogerythrus Dyar. Venation. 

Anegcephalesis arctella (Ragonot), figured 
from its synonym, cathaeretes Dyar. 
Venation. 

Dioryctria abietella (Denis and Schiffer- 
miller). Venation. 

Laodamia faecella (Zeller). Venation. 

Oryctometopia fossulatella Ragonot. Vena- 
tion. 

Hypochalcia ahenella (Schiffermiiller), male 
genitalia; male genitalia 54a, detached 
harpe in ventral view; 54b, dorsal view 
of tegumen and uncus; 54c, aedeagus; 
54d, gnathos; 54e, anellus; 54f, sternite 
and tergite of eighth abdominal seg- 
ment. (Type of an Old World genus 
not represented in the New World.) 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


49. 


Ee SS ete Anagcephalesis 


51. Dioryctria 


52. 
Laodamia 


53. 
Oryctometopia 


ahenella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 339 


Fieures 55-62.—VENATION (MALE UNLESS 
OTHERWISE NOTED). 


55. Cabotia semidiscella Ragonot. 

56. Oncolabis anticella Zeller. 

57. Honora mellinella Grote. 

58. Edulica compedella (Zeller). 

56. Oncolabis 59. Ocala dryadella Hulst. 

60. [llatila gurbyris Dyar. 

61. Macrorrhinia aureofasciella Ragonot. 
62. Psorosina hammondi (Riley). 


~$8. Edulica 


NS = Y 
= 
Cre Sf 


se 
a 


. : 
a Sean Cacia — 60. Illatila 


61. Macrorrhinia 62. Psorosina 


340 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figtres 63-70.—VENATION (MALE UNLESS 
OTHERWISE NOTED). 


63. Laetilia coccidivora (Comstock). 

64. Hulstia undulatella (Clemens). 

65. Tucumania tapiacola Dyar. 

66. Cactoblastis cactorum (Berg), female. 
67. Ozamia fuscomaculella clarefacta Dyar. 
68. Yosemitia graciella (Hulst). 

69. Parolyca asthenosoma (Dyar). 

70. Rioja nera Heinrich, new species. 


63. Laetilia 64. Hulstia 


66. Cactoblastis 


es 


67. Ozamia 68.Yosemitia 


69. Parolyca 70 Rioja 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 341 


Ficures 71-79.—VENATION (MALE UNLESS 
OTHERWISE NOTED). 


71. Olyca phryganoides Walker. 

72. Cayennia rufitinctalis Hampson. 

73. Cactobrosis fernaldialis (Hulst). 

74. Melitara prodenialis Walker. 

75. Olycella junctolineella (Hulst). 

76. Hxuperius negator Heinrich. 

77. Euzophera cinerosella (Zeller). 

78. Hulogia ochrifrontella (Zeller), female. 

79. Hulogia ochrifrontella (Zeller), hind wing of 
female, showing variation in venation. 


73. Cactobrosis 


74. Melitara 
6. Exuperius 
%S on 
75. Olycella 


Euzophera 79. Eulogia 


342 


Figures 80-89.—VENATION (MALE UNLESS 


80. 
81. 
82. 
83. 
84. 
85. 
86. 
87. 
88. 
89. 


OTHERWISE NOTED). 


Canarsia ulmiarrosorella (Clemens). 
Entmemacornis proselytes Dyar. 

Moerbes dryopella (Schaus). 

Moodnopsis decipiens Dyar, female. 
Patriciola semicana Heinrich, new species. 
Aptunga macropasa (Dyar), female. + ys 
Moodnopsis inveterella (Dyar). 

Eurythmia hospitella (Zeller). 

Varneria postremella Dyar. 

Nonia exiguella (Ragonot). 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


6]. Entmemacornis. 


82. Moerbes 


83.Moodnopsis 2 


—~ 
—=- 


85. Aptunga 


= 


87. Eurythmia 86.Moodnopsis 


88 .Vorneria 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 343 


SF 


Eurythmidia 


Phestinia 


Strephomescinia 


Micromescinia 


ma é 


id SS 


awh 2; 97. Gennadius P 99. SS 46 Farnobia? 


98. Farnobiaw” 
AN SS 


Figures 90-99.—VENATION (MALE UNLESS 
OTHERWISE NOTED). 


90. Wunderia neaeriatella Grossbeck, female. 

91. Eurythmidia ignidorsella (Ragonot). 

92. Phestinia costella Hampson. 

93. Strephomescinia schausella Dyar. 

94, Mescinia bacerella Dyar. 

95. Eurythmasis ignifatua Dyar. 

96. Micromescinia pygmaea Dyar. 

97. Gennadius junctor Heinrich, new species. 

98. Farnobia quadripuncta (Zeller), male. 

99. Farnobia quadripuncta (Zeller), hind wing 
of female. 


344 UNITED 


Fieures 100-110.—VENATION (MALE UNLESS 
OTHERWISH NOTED). 


100. Metephestia simplicula (Zeller). 

101. Divitiaca ochrella Barnes and McDun- 
nough. 

102. Oedothmia endopyrella Hampson, figured 
from a sketch, by J. F. G. Clarke, of 
the type of its synonym Synothomia 
bahamasella Hampson. 

103. Diviana eudoreella Ragonot; 103a, fore- 
wing showing variation in venation. 

104. Palatka nymphaeella (Hulst). 

105. Cacozophera venosa Dyar. 

106. Stylobasis rubripurpurea Hampson. 

107. Prosoeuzophera impletella (Zeller). 

108. Ephestiodes gilvescentella Ragonot. 

109. Ephestiodes plorella Dyar, Forewing of 
its synonym Eurythmia vestilla Dyar. 

110. Azaera muciella Schaus. 


STATES NATIONAL MUSEUM BULLETIN 207 


100. Metephestia 


1 10 9 


2 


103. Diviana 


107. 


Prosoeuzophera, 


Ol. Divitiaca 


Se 


102. Oedothmia 


110. Azaera 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 345 


= 


112. Homoeosoma 


SS 


f 
15.Vitula 116. Verina 


117. Comotia 
————— 


119. Bema 2 


~ 
™ 


120. Bema 2 


300329—56——23 


Figures 111-120.—VENATION (MALE UNLESS 
OTHERWISE NOTED). 


111. Harnocha velessa Dyar. 

112. Homoeosoma sinuellum (Fabricius). 

113. Manhatta biviella (Zeller), female. 

114. Moodna ostrinella (Clemens). 

115. Vitula edmandsae (Packard). 

116. Verina supplicella (Dyar). 

117. Comotia torsicornis Dyar. 

118. Bema neuricella (Zeller), male. 

119, 120. Bema neuricella (Zeller), females, 
showing vestiges of vein 9 in forewings. 


346 UNITED STATES NATIONAL MUSEUM BULLETIN 207. 


Figures 121—-130.—VENATION (MALE UNLESS 
OTHERWISE NOTED). 


121. Unadilla erronella (Zeller). 

122. Caudellia apyrella Dyar. 

123. Bethulia championella Ragonot, female. 
124. Plodia interpunctella (Hiibner). 

125. Ephestia cautella (Walker). 

126. Ephestia elutella (Hiibner). 

127. Nicetiodes apianella Schaus. 

126. Cabnia myronella Dyar. 

129. Ribiria conops (Dyar). 

130. Microphycita titillella Dyar. 


121. Unadilla 122. Caudellia 


ee 


SS 


123. Bethulia 24. Plodia 


125. Z 
Ephestia 126. Ephestia 


Saaaaa— 


127. 
Nicetiodes 
128. Cabnia 


N. 


130. Microphycita 


2 


129. Ribiria 


1340. ew 
134. indigenella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 347 


‘134d, 


Figures 131-134.—Mates. 


131. Cryptoblabes rutilella Zeller, right harpe 
denuded and aedeagus omitted; 131a, 
vinculum, tegumen, and uncus; 131b, 
harpe, denuded except for ventral tuft; 
131c, aedeagus; 131d, gnathos; 13le, 
transtilla and anellus; 131f, two views 
(scaled and denuded) of basal segments 
of antenna. 

132. Cryptoblabes gnidiella (Millitre), left 
harpe detached and aedeagus omitted; 
132a, anellus; 132b, transtilla; 132c, 
gnathos; 132d, aedeagus. 

133. Acrobasis tumidella (Zincken); 133a, 
gnathos; 133b, transtilla; 133c, anellus; 
133d aedeagus; 133e, ventral tuft of 
eighth abdominal segment. 

134. Acrobasis indigenella (Zeller) ; 134a, aede- 
agus; 134b, transtilla; 134c,d, basal 
segments of antenna, denuded and 
scaled. 


348 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieurres 135-139.—Ma tess. 


135. Acrobasis tricolorella Grote, 135a, aedea- 
gus; 135b, gnathos; 135c, transtilla; 
135d, ventral tuft of eighth abdominal 
segment. 

136. Acrobasis comacornella (Hulst), type; 
136a, aedeagus; 136b, gnathos; 136c, 
transtilla; 136d, anellus; 136e, ventral 
tuft of eighth abdominal segment. 

137. Acrobasis caryae Grote, specimen reared 
from pecan nut, tuft of eighth ab- 
dominal segment. 

138. Acrobasis juglandis (LeBaron), specimen 
Treared from pecan; 138a, aedeagus; 
138b, gnathos; 138c, transtilla; 138d, 
anellus. 

139. Acrobasis latifasciella Dyar, type; 139a, 
aedeagus. 


iss encolarella 135c. 


136.comacornella 


136d. 


138. juglandis 139, latifasciella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 349 


142. 
palliolella 


lake, 142a. 


144, 
sylviella 


14.6. 
evanescentella, 
149 a. 147a. 
149. \ 
aurorella 


151. 


exsulella 


152a. 


Fiaures 140-154.—mates or AcROBASIS 
SPECIES: UNCUS AND GNATHOS AND (a) 
TRANSTILLA: 


140. 


141. 
142, 


143. 
144, 
145. 
146. 


147. 
148. 


149. 
150. 
151. 
152. 


153. 


154. 


Acrobasis minimella Ragonot, figured 
from type of its synonym A. nigro- 
signella Hulst. 

Acrobasis feltella Dyar, type. 

Acrobasis palliolella Ragonot, male from 
Chicago, Ill. 

Acrobasis caryalbella Ely, type. 

Acrobasis sylviella Ely. 

Acrobasis kearfottella Dyar, type. 

Acrobasis caryae Grote, southern male, 
reared from pecan nut. 

Acrobasis evanescentella Dyar. 

Acrobasis stigmella Dyar, reared specimen 
from Falls Church, Va. (Hopk. U. S. 
No. 12121d). 

Acrobasis aurorella Ely, type. 

Acrobasis peplifera Dyar, type. 

Acrobasis exsulella (Zeller). 

Acrobasis angusella Grote, figured from 
type of its synonym A. eliella, Dyar. 
Acrobasis demotella Grote, specimen 

from New Brighton, Pa. 

Acrobasis irrubriella Ely, type. 


350 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 155-166.—Matzs. 


155. Acrobasis normella Dyar, type, uncus 
and gnathos; 155a, another male, 
showing variation in apical process of 
gnathos; 155b, type, transtilla. 

156. Acrobasis malipennella Dyar, type, uncus 
and gnathos; 156a, transtilla. 

157. Acrobasis dyarella Ely, type, uncus and 
gnathos; 157a, transtilla. 

158. Acrobasis ostryella Ely, type, uncus and 
gnathos; 158a, transtilla. 

159. Acrobasis secundella Ely, type, uncus and 
gnathos; 159a, transtilla. 

160. Acrobasis coryliella Dyar, type, uncus 
and gnathos; 160a, transtilla. 

161. Acrobasis caryivorella Ragonot, specimen 
reared on pecan at Austin, Tex. 
uncus and gnathos; 161a, transtilla. 

162. Acrobasis cunulae Dyar and Heinrich: 
162-162a, Ventral and lateral views of 
uncus and gnathos; 162b, apical proc- 
ess of gnathos from a male showing 
the broadest development in this 
structure; 162c, transtilla. 

163. Acrobasis myricella Barnes and McDun- 
nough, type, uncus and gnathos; 163a, 
transtilla. 

164. Acrobasis betulella Hulst, uncus and 
gnathos; 164a, transtilla. 

165. Acrobasis rubrifasciella Packard, reared 
male from New York, uncus and 
enathos; 165a, transtilla. 

166. Rhodophaea advenella (Zincken), aedea- 
gus omitted; 166a, aedeagus; 166b, 
transtilla; 166c, ventral tuft on eighth 
abdominal segment. 


ostryella 


162. 


162. 


cunulae 


163. 


We, aN Ally 
Wz ate C/I AS. art Wa 
Ny Al ( a wi sy i SS S/ 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 351 


168.ochrodesma tN 


169. 


Ficures 167-170.— Mates. 


167. Rhodophaea supposita (Heinrich), aedeag- 
us omitted; 167a; aedeagus; 167b, 
gnathos; 167c, transtilla. 

168. Anabasis ochrodesma (Zeller), aedeagus 
omitted; 168a, aedeagus; 168b, trans- 
tilla; 168c, sternite and tergite of 
eighth abdominal segment, showing 
ventral tufts. 

169. Mildrizia constitutionella Dyar; 169a, 
aedeagus; 169b, anellus; 169c, gnathos; 
169d, transtilla; 169e, sternite and 
tergite of eighth abdominal segment, 
showing ventral tuft; 169f, basal 
segments of antenna, partially de- 
nuded. 

170. Cuniberta subtinctella (Ragonot); 170a, 
aedeagus; 170b, gnathos; 170c, trans- 
tilla; 170d, anellus. 


352 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 171, 172.—Mates. 


171. Sematoneura atrovenosella Ragonot, ae- 
deagus omitted; 17la, aedeagus: 171b, 
vinculum, tegumen, gnathos and uncus, 
ventral view; 171c, transtilla; 171d, 
anellus; 17le, ventral hair tuft of 
eighth abdominal segment. 

172. Sematoneura abitus Heinrich, new species, 
type; 172a, aedeagus; 172b, gnathos; 
172c, transtilla. 


171. atrovenosello 


172b. 


172.abitus 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 353 


173. a : | V y 
\ 


300329—56——24 


Ficurss 173, 174.—Matss. 


173. Hypsipyla pagodella Ragonot; 173a, 
aedeagus; 173b, gnathos; 173c, trans- 
tilla; 173d, anellus; 178e, sternite, 
tergite and ventrolateral tufts of 
eighth abdominal segment. 

174. Hypsipyla grandella (Zeller), aedeagus 
omitted; 174a, aedeagus; 174b, gna- 
thos; 174c, transtilla; 174d, sternite 
and tergite of eighth abdominal seg- 
ment. 


354 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 175-177. MALEs. 


175. Hypsipyla dorsimacula (Schaus); 175a, 
aedeagus; 175b, gnathos; 175c, trans- 
tilla. 

176. Hypsipyla ferrealis (Hampson); 176a, 
aedeagus; 176b, gnathos; 176c, trans- 
tilla; 176d, anellus; 176e, eighth 
abdominal segment, showing tufts and 
modifications of sternite and tergite. 


177. Hypsipyla fluviatella Schaus; 177a, ae- 175 x 
deagus; 177b, sternite and tergite of e: Ss 
eighth abdominal segment; 177c, gna- 175. dorsimacula 


thos; 177d, transtilla; 177e, anellus. 


“x \T1b. 
MI 
Me 


WS 


YY iy, 
yy 
NY 


177. fluviatella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 355 


178 
chinographella 


SZ 


br 


1798 


A plumigerella 


bigrana 


Figures 178-180.—Matzs, 


178. Hemiptilocera chinographella Ragonot; 
178a, aedeagus; 178b, apex of aedea- 
gus, showing bent and spined cornutus 
in another view; 178c, transtilla; 178d, 
modified sternite and hair tufts of 
eighth abdominal segment; 178e, basal 
segments of antenna. 

179. Hemiptilocera plumigerella (Ragonot), 
type; 179a, aedeagus; 179b, transtilla; 
179c, sternite and tufts of eighth 
abdominal segment. 

180. Hemiptilocera bigrana (Zeller); 180a, 
aedeagus; 180b, transtilla; 180c, tufts 
and modified sternal plates of eighth 
abdominal segment. 


356 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 181-183.— Mats. 


181. Crocidomera stenopteryx (Dyar), type; 
18la, aedeagus. 

182. Crocidomera turbidella Zeller, specimen 
from Cuba, aedeagus omitted; 182a, 
aedeagus; 182b, transtilla; 182c, an- 
ellus; 182d, scale and hair tufts of 
eighth abdominal segment. 

183. Crocidomera fissuralis (Walker), speci- 
men from Puerto Rico (in Cornell); 
183a, aedeagus; 183b, transtilla; 183c, 
anellus; 183d, scale and hair tufts of 
eighth abdominal segment. 


NIN 
PE FAW 


ey 


183. fissuralis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 357 


Figures 184—187.—Matss. 


184. Heras disjunctus Heinrich, new species, 
type; 184a, aedeagus; 184b, gnathos; 
184c, transtilla; 184d, anellus. 

185. Adanarsa intransitella (Dyar), aedeagus 
omitted; 185a, aedeagus; 185b, anellus; 
185c, gnathos; 185d, transtilla; 185e, 
tufts on eighth abdominal segment. 

186. Birinus russeolus Heinrich, new species, 
type; 186a, aedeagus; 186b, gnathos; 
186c, transtilla and anellus. 

187. Bertelia grisella Barnes and McDun- 
nough, type, aedeagus omitted; 187a, 
aedeagus; 187b, transtilla; 187c, tufts 
on eighth abdominal segment. 


186. 


russeolus 


358 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 188-190.— Mates. 


188. Hypargyria definitella (Zeller), aedeagus 
omitted; 188a, aedeagus; 188b, gna- 
thos; 188c, transtilla; 188d, anellus; 
188e, eighth abdominal segment, show- 
ing ventral scale tufts. 

189. Chararica annuliferella (Dyar), aedeagus 
omitted; 189a, aedeagus; 189b, par- 
tially fused gnathos and transtilla; 
189c, anellus. 

190. Chararica hystriculella (Hulst), aedeagus 
omitted; 190a, aedeagus; 190b, par- 
tially fused gnathos and transtilla; ‘ 188. definitella 
190c, anellus; 190d, hair tufts on 
eighth abdominal segment. 


189. annuliferella 


190. hystriculella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


The 


192. subtetricella 


193. clatella 


g 


/195. 
piazzella 
195a. 


359 


Fiagures 191—195.—Mat.zs. 


191, Myelopsis coniella (Ragonot); 19la, 
aedeagus; 191b, gnathos; 191c, trans- 
tilla; 191d, anellus. 

192. Myelopsis subtetricella (Ragonot); 192a, 
aedeagus; 192b, gnathos; 192c, trans- 
tilla. 

193. Myelopsis alatella (Hulst); 193a, aedea- 
gus; 193b, gnathos; 193c, transtilla. 

194. Myelopsis fragilella (Dyar), type, 
synonym of M., alatella (Hulst); 194a, 
transtilla. 

195. Myelopsis piazzella (Dyar), type, @ syno- 
nym of M. alatella (Hulst); 195a, 
transtilla, 


360 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 196-199.—Matss. 


196. Anypsipyla wnivitella Dyar, aedeagus 
omitted; 196a, aedeagus, 196b; trans- 
tilla; 196c, ventrolateral tufts on 
eighth abdominal segment. 

197. Apomyelois bistriatella (Hulst); 197a, 
aedeagus; 197b, gnathos; 197c, trans- 
tilla. 

198. Ectomyelois decolor (Zeller), aedeagus 
omitted; 198a, aedeagus; 198b, trans- 
tilla. 

199. Ectomyelois ceratoniae (Zeller), aedeagus 
omitted; 199a, aedeagus; 199b, trans- 
tilla. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 361 


Ficurrs 200-203.—Matszs. 


200. Hctomyelois muriscis (Dyar), aedeagus 
omitted; 200a, aedeagus; 200b, trans- 
tilla. 

201. Ectomyelois zeteki Heinrich, new species, 
type; 201a, aedeagus. 

202. Paramyelois transitella (Walker); 202a, 
aedeagus; 202b, gnathos; 202c, ele- 
ments of divided transtilla; 202d, 
anellus. 

203. Myelois cribrella (Hiibner), aedeagus 
omitted; 203a, aedeagus; 203b, trans- 
tilla. 


SS ee 
wy PP 


203b. 


362 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 204-209.— Mates. 


204, Pseudodivona commensella Dyar, type, 
aedeagus omitted; 204a, aedeagus; 
204b, elements of transtilla; 204c, ter- 
gite, sternite, and ventrolateral tufts 
of eighth abdominal segment. 

205. Pseudodivona cispha Dyar, type, harpe; 

, . 205a, uncus; 205b, gnathos. 

206. Pseudodivona carabayella Dyar, type, 
uncus; 206a, specimen from Incachaca, 
Bolivia, gnathos. 

207. Protomoerbes separabilis Heinrich, new 
species, type; 207a, aedeagus; 207b, 
tergite and sternite of eighth abdom- 
inal segment. 

208. Protomoerbes aberrans Heinrich, new 
species, type; 208a, aedeagus; 208b, 
elements of transtilla. 

209. Diatomocera tenebricosa (Zeller); 209a, 
transtilla. (See also fig. 565.) 


206a. 
carabayella 


ww 


209.tenebricosa 


AMERICAN MOTHS OF THE SUBFAMILY 


210.dosia 


Diz dectecens 


213. 


majuscula 


PHYCITINAE 363 


Ficurs 210-213.—Ma tes. 


210. Diatomocera dosia (Dyar), type; 210a, 
aedeagus; 210b, hair tufts of eighth 
abdominal segment. 

211. Diatomocera excisalis (Hampson), speci- 
men from St. Laurent Maroni, French 
Guiana (in USNM); 211a, aedeagus. 

212. Diatomocera decurrens (Dyar), type; 
212a, aedeagus. 

213. Diatomocera majuscula Heinrich, new 
species, type; 213a, aedeagus; 213b, 
transtilla; 213c, tergite, sternite, and 
tufts of eighth abdominal segment. 


364 UNITED STATES NATIONAL: MUSEUM BULLETIN 207 


Figures 214-219.—Matss. 


214. Diatomocera albosigno Heinrich, new 
species, type, one harpe omitted; 214a, 
transtilla; 214b, tergite, sternite, and 
hair tufts of eighth abdominal seg- 


ment. 
215. Diatomocera hoplidice (Dyar), type; 215a, 1 UNS 
aedeagus. Wy, 


216. Diatomocera mochlophleps (Dyar), type; 
216a, aedeagus. 

217. Diatomocera extracta Heinrich, new spe- 
cies, type; 217a, aedeagus. 

218. Pseudocabima castronalis Heinrich, new 
species, type; 218a, aedeagus. 

219. Pseudocabima fearnella (Schaus), type; 
219a, aedeagus. 


214. albosigno 


211. 


extracta 


216.” 
mochlophleps 


castronalis 


219. fearnella 


224% 


expunctrix 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 365 


225. 


rubrizonalis 


220. 
221, 


222. 


223. 
224, 


225. 


Ficures 220-225.— Mates. 


Pseudocabima guianalis Heinrich, new 
species, type; 220a, aedeagus. 

Pseudocabima pombra (Dyar), type, un- 
cus. 

Pseudocabima nigristrigella (Ragonot), 
type; 222a, aedeagus; 222b, tergite, 
sternite, and hair tufts of eighth ab- 
dominal segment. 

Pseudocabima euzopherella (Dyar); 223a, 
aedeagus. 

Pseudocabima expunctrix Dyar and Hein- 
rich, type; 224a, aedeagus. 

Pseudocabima rubrizonalis (Hampson), 
“cotype” (in USNM) from Cayenne, 
French Guiana; 225a, aedeagus. 


366 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 226-230—MatgEs. 


226. Hyalospila stictoneurella Ragonot; 226a, 
aedeagus; 226b, transtilla; 226c, an- 
ellus; 226d, tergite, sternite, and hair 
tufts of eighth abdominal segment. 

227. Hyalospila celiella Schaus, type; 227a, 
aedeagus; 227b, transtilla; 227c, an- 


ellus. 

228. Hyalospila fulgidula Heinrich, new spe- ig 
cies, type; 228a, aedeagus; 228b, trans- 226. 
tilla; 228c, anellus. 2260. reves Sorelle 


229. Hyalospila xsanthoudemia (Dyar), type; 
229a, aedeagus; 229b, transtilla; 229c, 
anellus; 229d, tergite, sternite, and hair 
tufts of eighth abdominal segment. 

230. Hyalospila clevelandella (Dyar), type; 
230a, aedeagus; 230b, transtilla; 230c, 
anellus; 230d, tergite, sternite, and hair 
tufts of eighth abdominal segment. 


ee, 
228. 
fulgidula 


229. xanthoudemia 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 367 


Fiaurres 231-233.—Ma tgs. 


231. Fundella pellucens Zeller, aedeagus omit- 
ted; 231a, aedeagus; 231b, lateral view 
of uncus, gnathos, and tegumen; 231c, 
tergite, sternite, and hair tufts of 
eighth abdominal segment; 231d, basal 
segments of antenna, scaled; 23le, 
basal segments of male antenna, de- 
nuded; 231f, hind tibia, showing hair 
tuft. 

232. Fundella ignobilis Heinrich, new species, 
aedeagus omitted; 232a, aedeagus; 
232b, anellus; 232c, lateral view of 
uncus, gnathos, and tegumen. 

233. Fundella ahemora Dyar, aedeagus omit- 
ted; 233a, aedeagus; 233b, lateral view 
of uncus, gnathos, and tegumen; 233c, 
foretibia, showing hair tuft. 


HAO ue 


" 
< ig 


ahemora 


368 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficgures 234-238.—Matus. 


234. Fundella argentina Dyar, aedeagus) omit- 
ted; 234a, aedeagus; 234b, apical 
projection of gnathos, showing the 
extreme of variability in this structure; 
234c, hair tufts on eighth abdominal 
segment. 

235. Difundella corynophora Dyar, aedeagus 
omitted; 235a, aedeagus; 235b, eighth 
abdominal segment, showing sclero- 
tized pocket developed from sternite. 

236. Difundella subsutella (Schaus), type, 
aedeagus omitted; 236a, aedeagus. 

237. Difundella distractor Heinrich, new spe- 
cies, both harpes and aedeagus omit- 
ted; 237a, aedeagus; 237b, harpe, 237c, 
sclerotized parts of eighth abdominal 
segment, showing sclerotized pocket of 
sternite. 

238. Difundella tolerata Heinrich, new species, 
type, both harpes, anellus, and aedea- 
gus omitted; 238a, aedeagus; 238b, 
anellus; 238c, harpe; 238d, scaled 
pocket between second and third 
segments of abdomen. 


237. distractor 


238 .tolerata 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 369 


24a. 


243a. 


Figures 239-243.—Ma tgs. 


239. Coplarthria dasypyga (Zeller); 239a, 


aedeagus; 239b, anellus; 239c, sclero- 
tizations of eighth abdominal segment, 
showing development from sternite. 


240. Promylea lunigerella Ragonot, specimen 


from Duncans, Vancouver Island, 
aedeagus omitted; 240a, aedeagus; 
240b, anellus; 240c, sclerotizations of 
eighth abdominal segment. 


241. Promylea lunigerella glendella (Dyar); 


241a, aedeagus; 241b, sclerotizations 
of eighth abdominal segment. 


242. Promylea mindosis Dyar, type; 242a, 


aedeagus; 242b, sclerotizations of 
eighth abdominal segment. 


243. Promylea dyari Heinrich, type; 248a, 


aedeagus. 


370 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 244-247.—Matsgs. 


244. Anadelosemia senesciella (Schaus), type, 
one harpe and aedeagus omitted and \ 
one harpe detached; 244a, aedeagus; 

244), anellus; 244c, sclerotization of 


eighth abdominal segment. 
245. Anadelosemia condigna Heinrich, new 


species, type; 245a, aedeagus; 245b, ue 
sclerotization of eighth abdominal seg- Gi. 
ment. 


246. Anadelosemia dulciella (Hulst); a syno- 244, senesciella 


nym of A. texanella; 246a, aedeagus; 
246b, anellus; 246c, sclerotization of 
eighth abdominal segment. 

247: Dasypyga alternosquamella Ragonot, 
aedeagus omitted; 247a, aedeagus; 
247b, combined gnathos and anellus; 
247c, sclerotization of eighth abdomi- 
nal segment. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 371 


Ficurrs 248-250.—Matzs. 


248. Rampylla orio Dyar, type; 248a, dorsal 
view of uncus, gnathos and tegumen; 
248b, aedeagus; 248c, anellus; 248d, 
harpe; 248e, sclerotization of eighth 
abdominal segment. 

249. Rampylla subcaudata (Dyar), type; 249a, 
aedeagus; 249b, sclerotization of eighth 
abdominal segment. 

250. Rampylla lophotalis Heinrich, new spe- 
cies; 250a, aedeagus; 250b, harpe; 
250c, transtilla; 250d, anellus; 250e, 
sclerotization of eighth abdominal seg- 
ment; 250f, uncus; 250g, tegumen; 
250h, vinculum. 


250.5. 
lophotalis 


372 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 251-254.—Mates. 


251. Fulrada querna (Dyar), type; 25la, 
aedeagus; 251b, sclerotization and tuft 
of eighth abdominal segment. 

252. Fulrada carpasella (Schaus), type; 252a, 
aedeagus; 252b, anellus; 252c, scale 
tufts and sclerotization of sternite of 
eighth abdominal segment. 

253. Scorylus cubensis Heinrich, new species; 
253a, aedeagus; 253b, sternite and 
tufts of eighth abdominal segment. 

254. Davara caricase (Dyar), aedeagus omit- 
ted; 2548, aedeagus; 254b, anellus; 254c, 
sternite of eighth abdominal segment; 
254d, basal segments of antenna. 


eS 


253. cubensis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 373 


Figures 255-259.—Matzs. 


255. Davara paranensis (Dyar), type; 255a, 
aedeagus. 

256. Davara interjecta Heinrich, new species; 
256a, aedeagus. 

257. Davara rufulella (Ragonot), aedeagus 
omitted; 257a, aedeagus; 257b, forked 
spine associated with anellus; 257e, 
anellus; 257d, sternite and tufts of 
eighth abdominal segment. 

258. Sarasota plumigerella Hulst, type, aede- 
agus omitted; 258a, aedeagus; 258b, 
sternite of eighth abdominal segment. 

259. Sarasota furculella (Dyar), aedeagus 
omitted; 259a, aedeagus; 259b, anellus; 
259c, sternite of eighth abdominal 
segment. 


259.furculella 


374 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 260—264.— Mates. \ ; i 


260. Piesmopoda rubicundella Zeller, type; 
260a, aedeagus. 

261. Piesmopoda xanthopolys Dyar, specimen 
from La Chorrera, Panamd; 261a, 
aedeagus; 261b, dorsal view of uncus 
and tegumen. 

262. Piesmopoda flavicans (Zeller), figured 
from type of its synonym P. fratella; 
262a, aedeagus. 

263. Piesmopoda parva Heinrich, new species, 


type; 263a, aedeagus. 7. ss Bees = 
264. Piesmopoda isabella (Dyar), type; 264a, SS ae 26 ail 
ae 260. rubicundella \ aa 261.xanthopolys 


aedeagus; 264b, anellus; 264c, sternite 
of eighth abdominal segment. 


264.isabella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 875 


—————— 


268.almonella 


sf \ SS ; 
266. — ey, 


Figures 265—268.— Mauss. 


265. Piesmopoda ragonoti (Dyar), type; 265a, 
aedeagus. 

266. Piesmopoda sxanthomera Dyar, figured 
from the type of its synonym P., 
zanthozona Dyar; 266a, aedeagus. 

267. Atheloca subrufella (Hulst), figured from 
type of its synonym Hyalospila ptychis 
Dyar, aedeagus omitted; 267a, aede- 
agus; 267b, anellus and fusing arms of 
gnathos; 267c, harpe; 267d, sternite of 
eighth abdominal segment. 

268. Praedonula almonella (Dyar), type, aede- 
agus omitted; 268a, aedeagus; 268b, 
anellus; 268c, transtilla and gnathos; 
268d, sternite of eighth abdominal 
segment, 


376 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 269-271. Matss. 


269. Peadus burdettellus (Schaus), type; 269a, 
aedeagus; 269b, anellus; 269c, ventral 
view of (detached) left harpe; 269d, 
ventral view of left harpe, denuded; 
269e, sternite and tuft of eighth ab- 
dominal segment. 

270. Peadus dissitus Heinrich, new species, 
type; 270a, aedeagus. 

271. Gabinius paulsoni (Ragonot), type; 271a, 
aedeagus; 271b, dorsal view of uncus, 
tegumen, and vinculum; 271c, trans- 
tilla; 271d, anellus; 27le, sternite of 
eighth abdominal segment. 


270a. 


270.dissitus 


271. pauilsoni 


27\b. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 377 


Ficures 272-274.—Matss. 


272. Ceracanthia mamella (Dyar), specimen 
from type locality, aedeagus omitted; 
272a, aedeagus; 272b, sternite of 
eighth abdominal segment. 

273. Ceracanthia vepreculella Ragonot, type; 
273a, aedeagus; 273b, gnathos; 273c, 
anellus; 273d, sternite of eighth ab- 
dominal segment. 

274. Megarthria peterseni (Zeller), specimen 
from Guatemala; 274a, aedeagus; 
274b, anellus; 274c, ventrolateral tufts 
and sternite of eighth abdominal seg- 

. ment, showing two views of the bent, 
272.mamella : digitate pocket of sternite. 


=) 274. peterseni 


800329—56——25 


378 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


YieuREs 275—-281.— Mates. 


275. Megarthria squamifera Heinrich, new 
- species; 275a, aedeagus; 275b, gnathos. 

276. Megarthria schaust Heinrich, new species; 
276a, aedeagus; 276b, gnathos; 276c, 
transtilla; 276d, anellus. 

277. Megarthria cervicalis Dyar, type; 277a, 
aedeagus; 277b, anellus; 277c, sternite 
and ventral tuft of eighth abdominal 
segment. 

278. Ceracanthia mamella (Dyar), basal seg- 
ments of antenna, scaled and denuded. 

279. Megarthria peterseni (Zeller), basal seg- 
ments of antenna, scaled and denuded. 

280. Megarthria squamifera Heinrich, new 
species, basal segments of antenna, 
scaled and denuded. 

281. Megarthria cervicalis Dyar, type, basal 
segments of antenna, scaled. 


275. Squamifera 


cervicalis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 379 


284.drucella 


Figures 282—284.—MA.gEs. 


282. Drescoma cyrdipsa Dyar, type, aedeagus 
omitted; 282a, aedeagus; 282b, trans- 
tilla; 282c, sternite of eighth abdom- 
inal segment, 

283. Drescoma ciniliza Dyar, type, aedeagus 
omitted; 283a, aedeagus; 283b, trans- 
tilla; 283c, sternite of eighth abdominal 

a segment. 

284. Drescoma drucella Dyar, a synonym of 
Drescomopsis soraella (Druce), aedea- 
gus omitted; 284a, aedeagus; 284b, 
transtilla; 284c, sternite and scale 
tufts of eighth abdominal segment. 


380 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fi@ures 285-287.—Matss. 


285. Monoptilota pergratialis (Hulst), geni- 
talia figured from type of its synonym 
M. nubilella Hulst; 285a, aedeagus; 
285b, elements of transtilla; 285c, 
anellus; 285d, lateral view of gnathos; 
285e, ventral view of gnathos; 285f, 
sclerotization and tuftings of eighth 
abdominal segment of male; 285g, 
basal segments of male antenna, 
denuded; 285h, basal segments of male 
antenna, scaled. 

286. Zamagiria dixolophella Dyar, type, aedea- 
gus omitted and tegumen, uncus, and 
gnathos slightly turned to show them 
in ventrolateral view; 286a, aedeagus; 
286b, male genitalia shown in full 
ventral view with both harpes and 
aedeagus omitted; 286c, compound 
tufts of eighth abdominal segment of 
abdomen. 

287. Zamagiria pogerythrus Dyar; 287a, aedea- 
gus. 


287. pogerythrus 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 381 


Ficurzis 288-290.—Matgs. 


288. Zamagiria hospitabilis Dyar, type, aedea- 
gus omitted; 288a, aedeagus; 288b, 
anellus. 

289. Zamagiria masculinus Dyar, type, aedea- 
gus omitted; 289a, aedeagus. 

. 290. Zamagiria laidion (Zeller), aedeagus 
omitted ; 290a, aedeagus; 290b, anellus. 


290. laidion 290b. 


382 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 291-294.—Matszs. 


291. Zamagiria fraterna Heinrich, new species; 
291a, aedeagus. 

292. Zamagiria australella (Hulst); 292a, 
aedeagus. 

293. Magiriopsis denticosella (Dyar), one harpe 
and aedeagus omitted and uncus and 
gnathos bent to show lateral view; 
293a, aedeagus. 

294. Anegcephalesis arctella (Ragonot); 294, 
aedeagus; 294b, harpe, ventral view; 
294c, compound tufts of eighth abdom- 
inal segment. 


293. denticosel 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 383 


Ficures 295-298.—Matss. 


295. Ancylostomia stercorea (Zeller), aedeagus 
omitted; 295a, aedeagus; 295b, com- 
pound tufts of eighth abdominal seg- 
ment. 

296. Ancylostomia sauciella (Zeller), type; 296a, 
aedeagus; 296b, dorsal view of uncus. 

297. Caristanius pellucidellus (Ragonot), aede- 
agus omitted and right harpe detached; 
297a, aedeagus; 297b, anellus; 297c, 
compound tufts of eighth abdominal 
segment. 

298. Caristanius decoloralis (Walker), aede- 
agus omitted; 298a, aedeagus; 298b. 
anellus, 


384 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 299-305.—Matss. 


299. Pima boisduvaliella (Guénée), anellus; 299. boisduvaliella 
299a, apical portion of harpe; 299b, fi 
cornuti. 302.vividella 

300. Pima fosterella Hulst, anellus; 300a, api- 
cal portion of harpe; 300b, cornuti. 

301. Pima albocostalialis (Hulst), apical por- 
tion of harpe; 301a, cornuti. 

302. Pima vividella (McDunnough), paratype 
from Saskatoon, Saskatchewan (in 
USNM); anellus; 302a, apical portion 
of harpe; 302b, cornuti. 

303. Pima albiplagiatella occidentalis Heinrich: 
New race, anellus. 

304. Pima fulvirugella (Ragonot), type; 304a, 
apical portion of harpe; 304b, cornuti. 

305. Pima albiplagiatella (Packard) ; 305a, ae- 
deagus; 305b, anellus; 305c, ventral 
view of right harpe; 305d, tufts of 
eighth abdominal segment. 


301. 


albocostalialis 


a 
7 


305. | 
albiplagiatella 


AMERICAN MOTHS OF THE SUBFAMILY 


308. denticulella 


307. granitella 


300329—36——26 


PHYCITINAE 385 


Ficures 306-308.—Ma.zs. 


306. Pima parkerella (Schaus), type; 306a, ae- 
deagus; 306b, ventral view of harpe. 

307. Pima granitella (Ragonot), figured from 
type of its synonym Megasis piperella 
Dyar; 307a, aedeagus; 307b, anellus; 
307c, ventral view of right harpe. 

308. Interjectio denticulella (Ragonot), speci- 
men from British Columbia, aedeagus 
omitted and right harpe detached and 
shown in ventral view; 308a, aedeagus; 
308b, anellus. 


386 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieurres 309-311.—Mates. 


309. Interjectio niviella (Hulst), aedeagus 
omitted and right harpe detached and 
shown in ventral view; 309a, aedeagus. 

310. Ambesa laetella Grote, aedeagus omitted; 
310a, aedeagus; 310b, tufts of eighth 
abdominal segment. 

311. Ambesa lallatalis (Hulst); 311a, aedeagus; 
311b, ventral view of right harpe; 311c, 
anellus. 


<—S 


‘ 


By FLT 


A WR + Ah MY i: 
ANZ SAU 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 387 


312. 


313. 


314. 


315. 


316. 


Ficures 312-316.— Mates. 


Ambesa walsinghami (Ragonot), speci- 
men from Plumas County, Calif.; 
312a, aedeagus; 312b, elements of 
transtilla; 312¢, apical portion of harpe. 

Ambesa walsinghami mirabella Dyar, 
apical portion of harpe. 

Catastia marginea (Schiffermiiller): aede- 
agus omitted; 3l4a, aedeagus; 314b, 
transtilla. 

Catastia incorruscella (Hulst), type; 315a, 
aedeagus; 315b, gnathos; 315c, ele- 
ments of transtilla; 315d, anellus; 315e, 
tufts and sclerotization of eighth ab- 
dominal segment. 

Catastia bistriatella (Hulst), specimen 
from Humphreys Basin, Calif.; 316a, 
aedeagus. 


388 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 317-321.—Matzs. 


317. Catastia actualis (Hulst) ; 317a, aedeagus; 
317b, elements of transtilla. 

318. Immyrla nigrovitiella Dyar, aedeagus 
omitted; 318a, aedeagus; 318b, com- 
pound tufts of eighth abdominal seg- 
ment. 

319. Oreana unicolorella (Hulst), two views of 
uncus and gnathos; 319a, aedeagus; 
319b, anellus. 

320. Olybria aliculella (Hulst) ; 320a, aedeagus. 

321. Olybria furctferella (Dyar), type; 321a, 
aedeagus. 


320. 


aliculella < furciferella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 389 


Figures 322-325.—Matzs, 


322. Salebriacus odiosellus (Hulst); 322a, 
aedeagus. 

323. Salebriaria turpidella (Ragonot), figured 
from type of its synonym Salebria 
ademptandella Dyar; 323a, aedeagus; 
323b, anellus. 

324. Salebriaria fructetella (Hulst), figured 
from type of its synonym Salebria 
rectistrigella Dyar; 3242, aedeagus; 
324b, anellus. 

325. Salebriaria pumilella (Ragonot), figured 
from type of its synonym Salebria 
georgiella Hulst; 325a, aedeagus; 325b, 
gnathos; 325c, elements of transtilla; 
325d, anellus; 325e, compound tufts of 
eighth abdominal segment. 


fructetella: 


S 2) 325, 
\ if pumilella 


325d. 


390 


326. 


327. 


328. 
329. 
330. 


331. 
332, 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 326-332.—Matss. 


Etiella zinckenella (Treitschke); 326a, 
ventral view of harpe; 326b, aedeagus. 

Glyplocera consobrinella (Zeller), aedeagus 
omitted; 327a, aedeagus; 327b, trans- 
tilla. 

Quasisalebria admixta Heinrich, type; 
328a, aedeagus. 

Oriholepis jugosella Ragonot; 329a, aede- 
agus. 

Polopeustis annulatella (Zetterstedt) ; 
330a, aedeagus. 

Polopeustis arctiella (Gibson), aedeagus. 

Salebria palumbella (Schiffermiller); 
332a, aedeagus. 


331. 
arctiella 


. ~ ZB 
~ 330.annulatella 


327b. 


332.palu 


i. 327. consobrinella 


mbella. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 391 


Freurss 333-336.— Matzs. 


333. Meroptera mirandella Ragonot, aedeagus 
omitted; 333a, aedeagus; 333b, trans- 
tilla. 

334. Meroptera pravella (Grote), specimen 
from Edmonton, Alberta; 334a, aede- 
agus. 

335. Meroptera abditiva Heinrich, new species, 
type; 335a, aedeagus, 

336. Nephopteryz rhenella (Zincken), aedeagus 
omitted; 336a, aedeagus; 336b, basal 
segments of antenna, denuded. 


392 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 337-342.—MaA.zs. 


337. Nephopteryz subfuscella (Ragonot), ae- 
deagus omitted; 337a, aedeagus; 337b, 
transtilla. 

338. Nephopteryx dammersi Heinrich, new 
species, aedeagus. 

339. Nephopteryz subcaesiella (Clemens), ae- 
deagus. 

340. Nephopteryz fernaldi (Ragonot), speci- 
men from Colorado, aedeagus omitted; 
340a, aedeagus. 

341. Nephopteryz basilaris Zeller; 34la, ae- 
deagus. 

342. Nephopteryx termitalis (Hulst), type; 
3428, aedeagus. 


339. \ 
subcaesiella 340. fernaldi 


320 teematalis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 393 


Molt 


344d- 


346. 
rubrisparsella 


343. 


344, 


345. 


346. 


347. 


348. 


Fiaures 343-348.—Matgzs. 


Nephopteryzx bifasciella Hulst, type; 348a, 
aedeagus. 

Nephopteryx uvinella (Ragonot); 344a, 
aedeagus; 344b, elements of trans- 
tilla; 344c, gnathos; 344d, anellus; 
344e, compound tufts of eighth ab- 
dominal segment. 

Nephopteryx celtidella (Hulst); 345a, 
aedeagus. 

Nephopteryz rubrisparsella (Ragonot), 
figured from type of its synonym 
texanella Hulst, 346a, aedeagus; 346b, 
gnathos; 346c, transtilla; 346d, anellus. 

Nephopteryz crassifasciella Ragonot, one 
harpe omitted, one detached, and 
aedeagus omitted; 347a, aedeagus. 

Tlascala reductella (Walker), figured from 
type of its synonym Pempelia gledit- 
schiella Fernald, aedeagus omitted; 
348a, aedeagus; 348b, compound ven- 
tral tufts of eighth abdominal segment. 


394 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 349-352.—Matzs. 


349. Tulsa finitella (Walker), specimen from 
Florida, harpe; 349a, aedeagus. 

350. Tulsa umbripennis (Hulst), figured from 
type of its synonym Ortholepis gillet- 
tella Dyar; 350a, aedeagus; 350b, 
harpe; 350c, compound tufts of eighth 
abdominal segment. 

351. Tulsa oregonella (Barnes and MeDun- 
nough), type; harpe; 351a, aedeagus. 

352. Homeographa lanceolella Ragonot, type, 
one harpe detached and aedeagus 
omitted; 352a, aedeagus; 352b, gna- 
thos; 352c, anellus; 352d, compound 
tufts of eighth abdominal segment. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 395 


354 


355 
356 


357 


i 355. 
, curvatellus 
4d. 


Bhi bcucei 


356b. 


3538. 


Fieurns 353-357.— Ma tzs. 


Telethusia ovalis (Packard), aedeagus 

omitted; 353a, aedeagus. 

. Phobus brucei (Hulst), type; 354a, 
aedeagus; 354b, elements of trans- 
tilla; 354c, anellus; 354d, gnathos; 
354e, sclerotizations aud tufts of 
eighth abdominal segment. 

. Phobus curvatellus (Ragonot), anellus. 

. Phobus incertus Heinrich, new species; 
356a, aedeagus; 356b, gnathos; 356c, 
anellus. 

. Actriz nyssaecolella (Dyar), type; 357a, 

aedeagus; 357b, anellus; 357c, gnathos; 

357d, uncus. 


396 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 358-361.—Matzs. 


358. Aciriz dissimulatriz Heinrich, type; 358a, 
aedeagus, 358b, gnathos; 358c, anellus. 

359. Stylopalpia lunigerella Hampson, aedea- 
gus omitted; 359a, aedeagus; 359b, 
transtilla; 359c, tufts of eighth abdomi- 
nal segment. 

360. Stylopalpia scobiella (Grote), 360a, aedea- 
gus; 360b, vestiges of transtilla; 360c, 
anellus. 

361. Siylopalpia argentinensis Heinrich, type; 
361a, aedeagus. 


358. dissimulatrix 


361. argentinensis 


364. impostor 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 397 


366.aequivoca 


362. 
363. 
364. 
365. 


366. 


Figures 362—366.—Ma tgs. 


Pyla fasciolalis (Hulst), type; 362a, 
aedeagus; 362b, anellus. 

Pyla fasciolalis (Hulst), variety with 
shorter harpe and vinculum. 

Pyla impostor Heinrich, new species; 
364a, aedeagus. 

Pyla insinuatriz Heinrich, new species, 
type; 365a, aedeagus. 

Pyla aequivoca Heinrich, new species, 
type; 366a, aedeagus. 


398 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 367—370.—Ma.ss. 


367. Pyla aenigmatica Heinrich, new species; 
genitalia dissected and one harpe, 
aedeagus, and anellus omitted; 367a, 
anellus; 367b, aedeagus; 367c, tufts of 
eighth abdominal segment. 

368. Pyla criddlella Dyar, type, aedeagus 
omitted; 368a, aedeagus. 

369. Pyla fusca (Haworth), aedeagus omitted; 
369a, aedeagus; 369b, tufts of eighth 
abdominal segment. 

370. Pyla hypochalciella (Ragonot), figured 
from type of its synonym P. blackmor- 
ella Dyar, aedeagus omitted; 370a, 
aedeagus; 370b, dorsal view of uncus, 
tegumen, and vinculum; 370c, anellus. 


7 


7 (43100. 


hypochalciclla 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 399 


Fieures 371-375.— Ma.gs. 


371. Pyla hanhamella Dyar, type, aedeagus 
omitted; 371la, aedeagus. 

372. Pyla scintillans (Grote), specimen from 
Tuolumne Meadows, Calif., aedeagus 
omitted; 372a, aedeagus; 372b, hair 
tufts of eighth abdominal segment. 

373. Pyla scintillans (Grote), figured from 
type of its synonym, P. feella Dyar, 
aedeagus omitted; 373a, aedeagus. 

374. Pyla rainierella Dyar, type, aedeagus 

puma ae Ky" i omitted; 374a, aedeagus. 

; 375. Pyla sylphiella Dyar, type, aedeagus 
pared bY omitted; 375a, aedeagus. 


\ 


Py sen Dr : 
rene ERT 


id 


375. sylphiella 


374. rainierella 


400 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 376-381.— Ma tes. 


376. Pyla aeneella Hulst, left harpe deformed, 
tight harpe detached, and aedeagus 
omitted; 376a, aedeagus. 

377. Pyla metalicella Hulst: Ventral view of 
unecus, gnathos, tegumen, and vincu- 
lum; 377a, anellus and harpe; 377b, 
aedeagus. 

378. Pyla aeneoviridella Ragonot, aedeagus 
omitted; 278a-b, two variations of the 
aedeagus. 

379. Pyla fasciella Barnes and McDunnough, 
aedeagus omitted; 379a, aedeagus. 

380. Pyla nigricula Heinrich, new species, 
aedeagus omitted; 380a, aedeagus. 

381. Pyla viridisuffusella Barnes and Mc- 
Dunnough, one harpe detached and 
one harpe and aedeagus omitted; 381a, 
aedeagus. 


381. 
viridisuffusella 


i 


380.nigricula 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 401 


tks j , i 


iia 
He 


epee: 
Oe 


( 


or 
Ne 


ca 


Fieures 382—385.—Mauus 


382. Dioryctria abietella (Denis and Schiffer- 
miiller); 382a, aedeagus; 382b, uncus, 
gnathos, and tegumen of a European 
example; 383c-e, variations in the 
costo-terminal part of harpe in Amer- 
can specimens; 383f, compound tufts 
on eighth abdominal segment. 

383. Dioryctria reniculella Grote; 383a, aedea- 
gus. 

384. Dioryctria ponderosae Dyar, type; 384a, 
aedeagus. 

385. Dioryctria majorella Dyar; figured from 
type of its synonym D. muellerana 
Dyar; 385a, aedeagus. 


402 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 386-389.—MaAteEs. 


386. Dioryctria auranticella (Grote) ; 386a, ae- 
deagus. 

387. Dioryctria erythropasa (Dyar), type; 3872, 
aedeagus. 

388. Dioryctria pygmaeella Ragonot; 388a, ae- 
deagus. 

389. Dioryctria zimmermani (Grote), typical 
eastern example; 389a, aedeagus. 


=) 386. auranticella 


388.pygmacella 


\ 


NN J 389. zimmermani 


393. subtracta 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 403 


Figures 390-394.—Matgs. 


. Dioryctria cambiicola (Dyar), type; 390a, 


uncus, with lateral margins flattened; 
390b, aedeagus. 


. Dioryctria baumhoferi Heinrich, new spe- 


cies, type; 391a, aedeagus. 


. Dioryctria gulosella (Hulst), uncus. 
. Dioryctria subtracta Heinrich, new spe- 


cies, type; 393a, aedeagus. 


. Dioryetria clarioralis (Walker), typical 


Florida specimen; 394a, aedeagus. 


404 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieurts 395-398.—MALgEs. 


395. Oryctometopia fossulatella Ragonot, one 
harpe detached and aedeagus omitted; 
395a, aedeagus; 395b, gnathos; 395c, 
eighth abdominal segment, showing 
hair tufts. 

396. Sarata edwardsialis (Hulst), type, one 
harpe detached and aedeagus omitted; 
396a, aedeagus; 396b, clasper of harpe 
(small example), elements of transtilla, 
and anellus. 

397. Sarata pullatella (Ragonot), uncus, 
gnathos, and tegumen of type; 397a, 
aedeagus of type; 397b, uncus, gnathos, 
and tegumen of example from Utah; 
397c, aedeagus of example from Utah; 
397d, elements of transtilla and anellus. 

398. Sarata punctella (Dyar), uncus, gnathos, 
and tegumen; 398a, elements of trans- 
tilla; 398b-d, variations in clasper of 
harpe; 398e, anellus; 398f, aedeagus. 


396b, 


a 


396. edwacdsialis: 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


4.02.dnopherella 


402a. 


x A 
AWAY 
eT i vy 


BAS 404. rubrithoracella 


}405.tephrella 


399. 


400. 


401. 


402. 


403. 


404. 


405. 


405 


Fiaures 399-405.— Mates. 


Saraia incanella (Hulst), aedeagus 
omitted; 399a, aedeagus; 399b, anellus. 

Sarata atrella (Hulst), uncus, gnathos, 
and tegumen; 400a, clasper of harpe; 
400b, aedeagus; 400c, elements of 
transtilla and anellus. 

Sarata caudellella (Dyar), type, uncus, 
gnathos, and tegumen; 401a, clasper of 
harpe, elements of transtilla and 
anellus; 401b, aedeagus. 

Sarata dnopherella Ragonot, type, uncus, 
gnathos, and tegumen; 402a, elements 
of transtilla and anellus; 402b, clasper 
of harpe; 402c, aedeagus. 

Sarata nigrifasciella Ragonot, type, 
uncus, gnathos, and tegumen; 403a, 
aedeagus; 403b, clasper of harpe; 
403c, anellus and elements of transtilla. 

Sarata rubrithoracella (Barnes and 
MecDunnough), type; 404a, aedeagus. 

Saraia tephrella Ragonot, type; 405a, ae- 
deagus; 405b, eighth abdominal seg- 
ment, showing ventrolateral tufts. 


406 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficurrs 406—410.—Mauzs. 


406. Philodema rhoiella (Dyar), paratype from 
type locality; 406a, ventral view of 
right harpe; 406b, anellus; 406c, aede- 
agus. 

407. Lipographis fenestrella (Packard); 407a, 
aedeagus; 407b, tergite and sternite of 
eighth abdominal segment. 

408. Lipographis truncatella (Wright), speci- 
men from Chula Vista, Calif., 408a, 
aedeagus; 408b, elements of transtilla 
and anellus; 408¢, tergite and sternite 
of eighth abdominal segment. 

409. Lipographis subosseella Hulst, type; 409a, 
gnathos; 409b, aedeagus; 409c, anellus; 
409d, elements of transtilla. 

410. Lipographis wmbrella (Dyar), type; 410a, 
aedeagus. 


406. rhoiella 


ag /\N 
410. umbrella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 407 


Figures 411-415.—Matrs. 


411. Adelphia petrella (Zeller), aedeagus 
omitted; 411la, aedeagus; 411b, anellus; 
41le, tufts of eighth abdominal seg- 
ment. 

412. Adelphia ochripunctella (Dyar), type; 
412a, aedeagus; 412b, ventral view of 
harpe; 412c, anellus; 412d, gnathos. 

413. Tota galdinella (Schaus), aedeagus omit- 
ted; 413a, aedeagus; 413b, transtilla; 
413c, anellus; 413d, tufts of eighth 
abdominal segment. 

414. Ufa lithosella (Ragonot) ; 414a, aedeagus. 

415. Ufa senia Heinrich, new species, type; 
415a, aedeagus. 


ZA 
411. petrella \ 


408 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 416—418.— Mass. 


416. Ufa rubedinella (Zeller); 4162, aedeagus; 
416b, harpe; 416c, anellus; 416d, com- 
pound tufts of eighth abdominal seg- 
ment. 

417. Elasmopalpus lignosellus (Zeller), aede- 
agus omitted; 417a, aedeagus; 417b, 
tufts of eighth abdominal segment, 
ventral view; 417c, lateral view of 
tufts of eighth abdominal segment. 

418. Acroncosa albiflavella Barnes and Mc- 
Dunnough; 4182, aedeagus, dorsal 
view; 418b, aedeagus, ventral view; 
418c-e, cornuti of penis, much en- 
larged; 418f, ventrolateral tufts of 
eighth abdominal segment; 418g, an- 
ellus. 


418b. 4.18. albiflavella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 409 


Figures 419-425.— Matgs. 


419. Passadena flavidorsella (Ragonot); 419a, 
aedeagus; 419b, sclerotizations and 
tufts of eighth abdominal segment; 
419c, elements of transtilla and anellus; 
419d, gnathos; 419e, clasper of harpe. 

420. Ulophora groteit Ragonot, aedeagus omit- 
ted; 420a, aedeagus; 420b, elements of 
transtilla; 420c, hair tufts of eighth 
abdominal segment. 

421. Ulophora guarinella (Zeller), specimen 
from Cuba, aedeagus omitted; 421a, 
aedeagus. 

422. Chorrera idiotes Dyar, one harpe detached 
and aedeagus omitted; 422a, elements 
of transtilla and anellus; 422b, aedea- 
gus; 422c, hair tufts of eighth abdom- 
inal segment. 

423. Chorrera extrincica (Dyar), harpe; 423a, 
anellus. 

424. Chorrera postica (Zeller), type, one harpe 
detached and aedeagus omitted; 424a, 
gnathos, elements of transtilla and 
anellus; 424b, tufts of eighth abdom- 
inal segment; 424c, aedeagus. 

425. Tacoma feriella Hulst; 425a, aedeagus. 


oN 
420b. 
421.guarinella 


Lh 


\Y.. 


423. extrincica 


= 


ZZ, | 


1 425.teriella 


% 
424. postica 


300328—5é6——27 


410 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 426—-428.—Mates. 


426. Epischnia prodromella (Hiibner), aedea- 
gus omitted; 426a, aedeagus; 426b, 
compound tufts of eighth abdominal 
segment. 

427. Laodamia faecella (Zeller), one harpe 
detached and aedeagus omitted; 427a, 
aedeagus; 427b, anellus; 427c, com- 
pound tufts of eighth abdominal seg- 
ment. 

428. Megasis rippertella (Zeller) ; 428a, aedea- 
gus; 428b, elements of transtilla; 428c, 
harpe; 428d, eighth abdominal seg- 
ment. 


F: E880. \; 
4 GC 427. 
2 4 aecella 
‘ 


42 Ta. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 41] 


Figures 429-433.—Mates. 


429. Adelperga cordubensiella _ (Ragonot), 
specimen from Tucumén (in USNM); 
429a, aedeagus; 429b, elements of 
transtilla and anellus; 429c, tufts of 

/ Sy eighth abdominal segment. 

429c. 430. Eumysia mysiella (Dyar), type; 430a, 
aedeagus. 

431. Eumysia maidella (Dyar), sclerotization 
and tufts of eighth abdominal segment, 
431la, elements of transtilla and anel- 
lus. 

432. Divitiaca parvulella Barnes and McDun- 
nough; 432a, aedeagus; 432b, anellus; 
432c, eighth abdominal segment, show- 
ing sclerotizations, haired lobes, and 
scale tufts. 

433. Divitiaca ochrella Barnes and McDun- 
nough, type; 433a, dorsal view of 
uncus and tegumen; 433b, aedeagus; 
433c, anellus and elements of trans- 
tilla; 433d, eighth abdominal segments, 
showing sclerotization, haired lobes, 
and scale tufts. 


> 


cordubensiella 


parvulella 


432b. 


SSS 


> SS 


~ ochrella 


412 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 434—438.—Matess. 


434. Valdivia lativittella (Ragonot), figured 
from the type of its synonym Zophodia 
aureomaculella Dyar; 434a, anellus; 
434b, aedeagus; 434c, sclerotization 
and tufts of eighth abdominal seg- 
ment. 

435. Valdivia lativittella (Ragonot), Browns- 
ville, Tex., example; 435a, anellus; 
435b, sclerotization and tufts of 
eighth abdominal segment. 

436. Valdivia coquimbella Ragonot, paratype 
from type locality (in BM); 436a, 
aedeagus; 436b, anellus; 436c, sclero- 
tization and tufts of eighth abdominal 
segment. 

437. Macrorrhinia aureofasciella Ragonot; 
437a, anellus; 437b, aedeagus; 437c, 
sclerotization, lobes and tufts of 
eighth abdominal segment. 

438. Ocala dryadella Hulst, type; 438a, anellus 
and elements of transtilla; 438b, aedea- 
gus; 438c, sclerotization, lobes and 
tufts of eighth abdominal segment. 


wv 
435. \ativittella 


y 


“35a. 


434 lativittella 
/ 4 


43 


aureofasciella 


anyadalian 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 413 


440 


mirabilicornella 


eS 441. 
samaritanella 


morrisonella 


Fiaurses 439-443.—Matzs. 


439. Valdivia walkerella (Ragonot), type; 
439a, aedeagus. 

440, Protasia mirabilicornella (Dyar), type; 
440a, aedeagus; 440b, anellus and 
elements of transtilla; 440c, scleroti- 
zations of eighth abdominal segment. 

441. Helerographis samaritanella (Zeller) ; 
44la, aedeagus; 441b, transtilla and 
anellus; 441c, tufts of eighth abdomi- 
nal segment. 

442. Heterographis morrisonella Ragonot; 
442a, anellus; 442b, aedeagus. 

443. Staudingeria olivacella Dyar, a synonym 
of S. albipenella (Hulst), type; 443a, 
transtilla and anellus; 443b, aedeagus. 


414 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 444-448 —MaAtes. 


444, Hulstia undulatella (Clemens), figured 
from the type of its synonym Honora 
fumosella Hulst; 4442, aedeagus; 444b, 
anellus and elements of transtilla; 
444c, sclerotization and tufts of eighth 
abdominal segment. 

445. Honora mellinella Grote, specimen from 
Texas; 445a, aedeagus; 445b, anellus 
and elements of transtilla; 445c, 
sclerotization and tufts of eighth 
abdominal segment. 

446. Honora sciurella Ragonot, type; 446a, 

: aedeagus; 446b, anellus and elements 
of transtilla; 446c, sclerotization and 
tufts of eighth abdominal segment. 

447. Honora subsciurella Ragonot; 447a, 
aedeagus; 447b, anellus and elements 
of transtilla; 447c, sclerotization and 
tufts of eighth abdominal segment. 

448. Honora dotella Dyar, type; 448a, aedea- 
gus; 448b, anellus and elements of 
transtilla; 448c, sclerotization and 
tufts of eighth abdominal segment. 


7168, 
dotella. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 415 


Fiaures 449-453.—Matgs. 


449. Honora perdubiella (Dyar) type; 449a, 
aedeagus; 449b, anellus and elements 
of transtilla; 449¢, sclerotization and 
tufts of eighth abdominal segment. 

450. Honorinus fuliginosus Heinrich, new 
species, type; 450a, aedeagus; 450b, 
anellus; 450c, transtilla; 450d, sclero- 
tization and tufts of eighth abdominal 


segment. 
Sms My 451. Cabotia semidiscella Ragonot; 45la, ele- 
= ; 450.fuliginosus a ments of transtilla; 451b, aedeagus; 
449, ey } \ i { Ay: 451c, eighth abdominal segment, show- 
perdubiella a) \ ‘gs ing tufts. 


452. Cabotia bonhoti (Hampson), specimen 
from Trelawney Parish, Jamaica; 
452a, anellus and elements of transtilla; 
452b, aedeagus; 452c, tufts of eighth 
abdominal segment; 452d, basal seg- 
ments of male antenna. 

453. Cabotia rhythmatica Dyar, type; 453a, 
anellus and elements of transtilla; 
453b, aedeagus; 453c, tufts of eighth 
abdominal segment. 


453. “Ss 
rhythmatica 


416 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 454—459.— Matss. 


454. Oncolabis anticella Zeller; 454a, aedeagus; 
454b, tufts of eighth abdominal seg- 
ment; 454c, elements of transtilla and 
anellus; 454d, basal segments of an- 
tenna. 

455. Canarsia ulmiarrosorella (Clemens) ; 455a, 
aedeagus; 455b, anellus; 455c, tufts of 
eighth abdominal segment. 

456. Harnocha velessa Dyar, type, aedeagus 
omitted; 456a, aedeagus; 456b, ele- 
ments of transtilla. 

457. Eurythmasis ignifatua Dyar, type, aedea- 
gus omitted; 457a, dorsal view of uncus 
and tegumen; 467b, gnathos and ele- 
ments of transtilla; 457c, aedeagus; 
457d, tufts of eighth abdominal seg- 
ment. 

458. Stylobasis rubripurpurea Hampson, type, 
three-quarter view; 458a, aedeagus. 

459. Eurythmidia ignidorsella (Ragonot), 


aedeagus omitted; 459a, aedeagus; Ve 4 455. 

459b, elements of transtilla; 459c, ter- iy = ulmiarrosorella 
gite and sternite of eighth abdominal i 456. velessa 

segment. 


ry 
ignifatua 


be 


cornic: 


300329—56——28 


AMERICAN MOTHS OF THE 


SUBFAMILY 


PHYCITINAE 417 


460. 


461. 


462. 


463. 


464. 


Figures 460—464.— Ma.gs. 


Patriciola semicana Heinrich, new spe- 
cies; 460a, aedeagus; 460b, ventral 
view of right harpe; 460c, gnathos; 
460d, elements of transtilla; 460e, 
anellus. 

Psorosina hammondi (Riley); 46la, 
aedeagus; 461b, anellus and elements 
of transtilla; 461c, tufts of eighth 
abdominal segment of abdomen. 

Palatka nymphaeella (Hulst); 462a, 
aedeagus; 462b, sclerotization of eighth 
abdominal segment. 

Diviana eudoreella Ragonot, figured from 
type of itssynonym, Dannemora eden- 
tella Hulst; 463a, aedeagus; 463b, 
gnathos; 463c, anellus. 

Paconius corniculatus Heinrich, new 
species, type; 464a, aedeagus; 464b, 
anellus; 464c, tufts of eighth abdom- 
inal segment. 


418 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 465-467.—Ma tgs. 


465. Aptunga macropasa (Dyar); 465a, ele- 
ments of transtilla; 465b, gnathos; 
465e, anellus; 465d, aedeagus; 465e, 
eighth abdominal segment, showing 
tufts and sclerotizations. 

466. Cassiana malacella (Dyar), type; 466a, 
anellus; 466b, aedeagus; 466c, gnathos; 
466d, elements of transtilla; 466e, 
eighth abdominal segment, showing 
tufts and sclerotizations. 

467. Anderida sonorella (Ragonot), figured 
from type of its synonym Hyzophera 
placidella Dyar; 467a, anellus; 467b, 
elements of transtilla; 467c, gnathos; 
467d, aedeagus; 467e, eighth abdom- 
inal segment, showing tufts and scle- 
rotizations. 


(23 b | i 
malacella hy ay F sepa 


66c. 


sonorella 


470 


bacerella 


471. parvula 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 419 


ine 


moorei 


Figurrs 468-473.—Mates. 


468. Mescinia triloses Dyar, type; 468a, aede- 


agus; 468b, gnathos; 468c, elements of 
transtilla; 468d, anellus; 468e, eighth 
abdominal segment, showing tufts and 
sclerotizations. 


469. Mescinia pandessa Dyar, type; 469a, 


gnathos; 469b, elements of transtilla; 
469c, anellus; 469d, aedeagus. 


470. Mescinia bacerella Dyar, type; 470a, ae- 


deagus; 470b, gnathos; 470c, elements 
of transtilla; 470d, anellus. 


471. Mescinia parvula (Zeller), paratype (in 


BM); 471a, aedeagus; 471b, gnathos; 
471c, elements of transtilla; 471d, 
anellus. 


472. Mescinia moorei Heinrich, new species; 


472a, aedeagus; 472b, elements of 
transtilla and anellus; 472c, gnathos. 


473. Mescinia estrella Barnes and McDun- 


nough, anellus and gnathos. 


420 UNITED STATES NATIONAL MUSEUM BULLETIN 207 \ 


Figures 474—476.— Mates. 


474. Mescinia peruella Schaus; 474a, aede- 
agus; 474b, gnathos; 474c, elements of 
transtilla; 474d, anellus. 

475. Mescinia discella Hampson; 475a, aede- 
agus; 475b, gnathos; 475c, elements of 
transtilla; 475d, anellus. 

476. Mescinia indecora Dyar; 476a, aedeagus; 
476b, gnathos; 476c, elements of trans- 
tilla; 476d, anellus; 476e, eighth ab- 
dominal segment, showing tufts and 
sclerotizations. 


74 


peruella 


476. indecora 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 421 


Fiagures 477-479.—Ma tess. 


477. Nonia exiguella (Ragonot), figured from 
type and from type of its synonym 
Hypermescinia lambella Dyar, aedeagus 
omitted; 477a, aedeagus; 477b, dorsal 
view of uncus and tegumen; 477c, 
gnathos; 477d, elements of transtilla; 
477e, anellus; 477f, eighth abdominal 
segment, showing tufts and sclerotiza- 
tions. 

478. Comotia torsicornis Dyar, type, aedeagus 
omitted; 478a, aedeagus; 478b, ele- 
ments of transtilla; 478c, tufts and 
sclerotizations of eighth abdominal 
segment; 478d, basal segments of an- 
tenna. 

479. Bema neuricella (Zeller), figured from 
type of its synonym B. myja Dyar, ae- 
deagus omitted; 479a, elements of 
transtilla; 479b, aedeagus; 479c, eighth 
abdominal segment, showing tuftings 
and sclerotizations; 479d, front view of 
head; 479e, front view of head, de- 
nuded. 


478. 


torsicornis 


479 \ 
Nneuricella 


‘422 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 480—-483.— Mates. 


. 480. Homoeosoma  sinuellum (Fabricius) ; 
480a, gnathos; 480b, elements of trans- 
tilla; 480c, anellus; 480d, aedeagus; 
480e, sclerotizations of eighth abdom- 
inal segment. 

481. Homoeosoma electellum (Hulst); 481a, 
vinculum; 481b, aedeagus; 481c, gna- 
thos; 481d, elements of transtilla; 
481e, anellus. 

- 482. Homoeosoma illuviellum Ragonot; 482a, 

- aedeagus; 482b, gnathos; 482c, anellus; 
482d, elements of transtilla; 482e, 
sclerotizations of eighth abdominal 
segment. 

483. Homoeosomaimitator Heinrich, new species; 
488a, lateral view of uncus, gnathos, and 
tegumen; 483b, aedeagus; 483c, gna- 
thos; 483d, elements of transtilla; 483e, 
anellus; 483f, sclerotizations of eighth 
abdominal segment, showing shortest 
form of ventral process; 483g, longest 
form of ventral process from eighth 
abdominal segment. 


480. sinue\lum 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 423 


485. inornatellum 


impressale 


Fiaures 484-486.—Matxgs. 


484. Homoeosoma longiventrellum Ragonot, 
3 specimens from Chiriqui, Panamé4; 
484a, aedeagus; 484b, gnathos; 484c, 
anellus; 484d, elements of transtilla; 
484e, sclerotizations of eighth abdom- 
inal segment. 

485. Homoeosoma inornatellum (Hulst) ; 485a, 
aedeagus. 

486. Homoeosoma impressale Hulst; 486a, 
aedeagus; 486b, gnathos; 486c, ele- 
ments of transtilla; 486d, anellus, 
486e, sclerotizations of eighth abdom- 
inal segment. 


424 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 487—490.— Ma tgs. 


487. Homoeosoma albescentellum Ragonot, fig- 
ured from type of its synonym H. 
elongellum Dyar; 487a, aedeagus. 

488. Homoeosoma deceptorium Heinrich, new 
species, paratype from Alcove, Quebec; 
488a, gnathos with apical projection 
shown in ventral and ventrolateral 
views; 488b, elements of transtilla; 
A88c, anellus; 488d, aedeagus; 488e, 
sclerotizations of eighth abdominal 
segment. 

489. Homoeosoma discrebile Heinrich, new 
species; 489a, aedeagus; 489b, gnathos; 
489c, anellus; 489d, elements of trans- 
tilla; 489e, tufts and sclerotizations of 
eighth abdominal segment. 

490. Homocosoma peregrinum Heinrich, new 
species; 490a-b, two views of aedeagus. 


discrebile 


( 489d,\ 


peregrinum 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 425 


Fieures 491-496.—Matgs. 


491. Homoeosoma vepallidum Heinrich, new 
species, type; 491a, aedeagus. 

492. Homoeosoma assitum Heinrich, new spe- 
cies, type; 491a, aedeagus. 

493. Homoeosoma acmaeopterum Ragonot, 
type; 498a, gnathos; 493b, elements of 
transtilla; 493c, anellus; 493d, aedea- 
gus; 498e, sclerotizations of eighth ab- 
dominal segment. 

494, Homoeosoma nimbosellum Ragonot, type; 
494a, aedeagus; 494b, gnathos and 
elements of transtilla; 494c, anellus. 

495. Rotruda mucidella (Ragonot); 495a, gna- 
thos; 495b, anellus; 495c, elements of 
transtilla; 495d, aedeagus; 495e, lat- 
eral view of uncus, gnathos, and tegu- 
men; 495f, tufts and sclerotizations of 
eighth abdominal segment. 

496. Rotruda nimbella (Zeller), aedeagus. 


491. vepallidum 


493e. 


695. ee 


mucidella 


426 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 497—502.—Matss. 


497. Strephomescinia schausella Dyar, type, 
aedeagus omitted; 497a, aedeagus; 
497b, anellus. 

498. Unadilla erronella (Zeller); 498a, aedea- 
gus; 498b, gnathos, elements of trans- 
tilla, and anellus; 498c-f, variations in 
vinculum. 

499. Unadilla maturella (Zeller), specimen 
from Guatemala; 499a, aedeagus; 
499b, gnathos; 499c, elements of 
transtilla; 499d, anellus. 

500. Laetilia portoricensis Dyar; 500a, aedea- 
gus; 500b, anellus; 500c, gnathos; 
500d, lateral view of female head. 

501. Laetilia melanostathma (Meyrick), lateral 
view of male head. 

502. Laetilia coccidivora (Comstock), lateral 
view of male head. 


498. erronella 


500. Sz 


ortoricensis 
P melanostathma 


coccidivora 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 427 


Figures 503-507.—Matgss. 


503. Laetilia coccidivora (Comstock); 508a, 
aedeagus; 503b, elements of trans- 
tilla; 503c, gnathos; 508d, anellus. 

504. Laetilia coccidivora quadricolorella 
(Dyar); 504a, aedeagus; 504b, ele- 
ments of transtilla; 504c, gnathos; 
504d, anellus. 

505. Laetilia coccidivora cardini Dyar, speci- 
men from Florida; 505a, aedeagus; 
505b, elements of transtilla; 505c, 
gnathos; 505d, anellus. 

506. Laetilia obscura Dyar; 506a, aedeagus; 
506b, elements of transtilla; 506c, 
gnathos; 506d, anellus. 

507. Laetilia melanostathma (Meyrick); 507a, 
aedeagus; 507b, elements of trans- 
tilla; 507c, gnathos; 507d, anellus. 


504. 


quadricolorella 


506 


obscura 


melanostathma 


428 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 508-511.— Mates. 


508. Laetilia zamacrella Dyar; 508a, anellus; 
508b, gnathos; 508c, elements of 
transtilla; 508d, aedeagus. 

509. Laetilia myersella Dyar; 509a, aedeagus; 
509b, gnathos; 509c, elements of 
transtilla; 509d, anellus. 

510. Laetilia ephestiella (Ragonot); 510a, 
gnathos; 510b, elements of trans- 
tilla; 510c, anellus. 

511. Laetilia fiskella Dyar; 511a, aedeagus; 
511b, gnathos; 511c, elements of trans- 
tilla; 511d, anellus. 


ephestiella 


511. 


Fiskella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 429 


Ja 


J se. ¢ 513c. > 


513d. 


My! 


maculatellae 


Fiaurses 512-514.—Ma.ss. 


512. Laetilia glomis (Dyar), type; 512a, 
aedeagus; 512b, gnathos; 512c, ele- 
ments of transtilla; 512d, anellus. 

518. Baphala goyensis (Ragonot), specimen 
from Parand, Brazil; 513a, aedeagus; 
513b, gnathos; 513c, elements of trans- 
tilla; 513d, anellus. 

514. Baphala basimaculatella (Ragonot), speci- 
men from Texas (in BM); 514a, 
aedeagus; 514b, gnathos; 514c, ele- 
ments of transtilla; 514d, anellus; 514e, 
sclerotizations and tufts of eighth 
abdominal segment. 


430 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 515-517.—Matzs. 


515. Baphala glabrella (Dyar), specimen from 
type locality; 515a, aedeagus; 515b, 
gnathos; 515c, elements of transtilla; 
515d, anellus. 

516. Baphala haywardi Heinrich, new species; 
516a, aedeagus; 516b, gnathos; 516c, 
elements of transtilla; 516d, anellus. 

517. Baphala homoeosomella (Zeller), specimen 
from type locality (in BM, Zeller, 
collection); 517a, aedeagus; 517b-c, 
gnathos with apical projection in 
ventrolateral and ventral views; 517d, 
elements of transtilla; 517e, anellus; 
517£, tufts and sclerotizations of eighth 
abdominal segment; 517g, basal seg- 
ments of antenna. 


homoeosomella 


519.stigmella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 431 


519e. 


Ficurss 518, 519.—Matss. 


518. Rhagea packardella (Ragonot), figured 
from type of its synonym Zophodia 
orobanchella Dyar; 518a, aedeagus; 
518b, gnathos; 518c, elements of trans- 
tilla; 518d, anellus. 

519. Rhagea stigmella (Dyar), type; 519a, 
aedeagus; 519b, gnathos; 519c, ele- 
ments of transtilla; 519d, anellus; 
519e, sclerotizations of eighth abdom- 
inal segment. 


432 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 520, 521.—Mates. 


NT TT ah Iie I mV ossiad r oS Sai] 

520. Rhagea stigmella (Dyar), figured from its 
synonym Yosemitia maculicula Dyar; 
520a, aedeagus; 520b, gnathos; 520c, 
elements of transtilla; 520d, anellus. 

521. Rhagea stigmella (Dyar), variety from 
Half Moon Bay, Calif.; 521a, aedeagus; 
521b, gnathos; 521c, elements of trans- 
tilla; 521d, anellus. 


Ss 
SR 
= 


521. stigmella i) 


52ld. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 433 


Figures 522-524.—Matzs. 


522. Zophodia convolutella (Hiibner), aedeagus 
omitted; 522a, aedeagus; 522b, anellus; 
522c, elements of transtilla; 522d, api- 
cal process of gnathos; 522e, scleroti- 
zations and tufts of eighth abdominal 
segment. 

523. Cactobrosis longipennella (Hampson), fig- 
ured from type of its synonym 
Moodna elongatella Hampson, aedegus 
omitted; 523a, dorsal view of uncus 
and tegumen; 523b, elements of trans- 
tilla; 523c, anellus; 523d, aedeagus; 
523e, tufts of eighth abdominal seg- 
ment. 

524. Cactobrosis fernaldialis (Hulst), part of 
shaft of antenna, ventral view. 


523. longipennella 


J 


434 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 525, 526.—MA.gzs. 


525. Cactobrosis maculifera Dyar; 5252, part of 
shaft of antenna, ventral view; 525b, 
basal segments of antenna, lateral 
view; 525c, elements of transtilla; 525d, 
anellus; 525e, aedeagus. 

526. Cactobrosis strigalis (Barnes and Mc- 
Dunnough); 526a, elements of trans- 
tilla; 526b, anellus; 526c, aedeagus. 


525. maculifera 


526. strigalis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 435 


Ficures 527, 528.—Matss. 


527. Melitara prodenialis Walker; 527a, dorsal 
view of uncus and tegumen; 527b, 
gnathos; 527c, elements of transtilla; 
527d, anellus; 527e-f, lateral and ven- 
tral views of aedeagus. 

528. Melitara dentata (Grote); 528a, ele- 
ments of transtilla; 528b, anellus; 
528c, aedeagus. 


528. dentata 


436 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 529-531.—Matss. 


529. Olycella junctolineella (Hulst); 529a, 
anellus; 529b, elements of transtilla; 
529¢c, aedeagus. 

530. Olycella junctolineella pectinatella (Hamp- 
son), gnathos. 

531. Olyca phryganoides Walker, specimen 
from Azuda, Santo Domingo; 531a, 
elements of transtilla; 531b, aedeagus; 
531c, anellus. 


SWS 
SS QQ SS -\ 
SSS ANY 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 437 


534. substituta 


Ficures 532-534.—Matezs. 


532. Alberada bidentella (Dyar), aedeagus 
omitted; 532a, elements of transtilla; 
532b, anellus; 532c, aedeagus. 

533. Alberada parabates (Dyar); 533a, ele- 
ments of transtilla; 523b, anellus; 
533c, aedeagus. 

534. Nanaia substituta Heinrich, type; 534a, 
elements of transtilla; 534b, anellus; 
534c, aedeagus; 534d, sternite and 
tergite of eighth abdominal segment. 


438 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 535-539.— Mates. 


535. Cactoblastis cactorwm (Berg); 535a, ele- 
ments of transtilla; 535b, anellus; 
535c, aedeagus. 

536. Cactoblastis doddi Heinrich, gnathos. 

537. Cactoblastis mundelli Heinrich, gnathos. 

538. Cactoblastis bucyrus Dyar: 538a, elements 
of transtilla; 538b, anellus; 538c, 
aedeagus. 

539. Cahela ponderosella (Barnes and Mc- 
Dunnough), type; 539a-c, various 
modifications of apical process of 
gnathos; 539d, elements of transtilla; 
539e anellus; 539f, aedeagus. 


535. cactorum Rick 536. doddi 


537. mundelli 


538. bucyrus 


539 ponderosella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 439 


1, polingella 


543. tapiacola 


Figures 540-543.—Matgs. 


540. Rumatha bihinda (Dyar), showing sub- 
basal sclerotized pocket (pkt.); 540a, 
elements of transtilla; 540b, anellus; 
540c, aedeagus. 

541. Rumatha polingella (Dyar); 541a, ele- 
ments of transtilla; 541b, anellus; 541c¢, 
aedeagus. 

542. Rumatha glaucatella (Hulst), aedeagus 
omitted; 542a, elements of transtilla; 
542b, anellus; 542c, aedeagus. 

543. Tucumania tapiacola Dyar, aedeagus 
omitted, vinculum somewhat fore- 
shortened; 543a, elements of transtilla; 
543b, anellus; 548c, aedeagus; 543d, 
sternite and tergite of eighth abdom- 
inal segment. 


440 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 544-547.—Mates. 


544, Yosemitia fieldiella (Dyar), aedeagus 
omitted; 544a, elements of transtilla; 
544b, anellus; 544c, aedeagus. 

545. Yosemitia didactica Dyar, aedeagus omit- 
ted; 545a, elements of transtilla; 545b, 
anellus; 545c, aedeagus. 

546. Yosemitia longipennella (Hulst), aedeagus 
omitted; 546a, elements of transtilla; 
546b, anellus; 546c, aedeagus. 

547. Yosemitia graciella (Hulst), aedeagus 
omitted; 5472, elements of transtilla; 
547b, anellus; 547c, aedeagus; 547d, 
sternite and tergite of eighth abdom- 
inal segment. 


547. graciella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 44] 


Fiaures 548-550.—Ma.gs. 


548, Eremberga leuconips (Dyar); 548a, ele- 
ments of transtilla; 548b, anellus; 548e, 
aedeagus. 

549. Eremberga insignis Heinrich, type; 549a, 
elements of transtilla; 549b, anellus; 
549c, aedeagus. 

550. Hremberga creabates (Dyar), type; 550a, 
elements of transtilla; 550b, anellus; 
550c, aedeagus, 


548. leuconips 


300329—56——29 


442 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 551-553.—Ma tes. 


551. Parolyca asthenosoma (Dyar), type; 551a, 
elements of transtilla; 551b, anellus; 
551c, aedeagus; 551d, part of shaft of 
antenna. 

552. Salambona analamprella (Dyar); 552a, 
elements of transtilla; 552b, anellus; 
552c, aedeagus. 

558. Sigelgaita transilis Heinrich, type; 553a, 
elements of transtilla; 553b, anellus; 
558c, aedeagus; 553d, eighth abdom- 
inal segment, showing hair tufts. 


553. transilis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 443 


Ficures 554, 555.—Matgzs. 


554. Sigelgaita chilensis Heinrich; 554a, ele- 
ments of transtilla; 554b, anellus; 554, 
aedeagus. 

555. Amalafrida leithella (Dyar); 555a, ele- 
ments of transtilla; 555b, anellus; 
555c, aedeagus; 555d, eighth abdominal 
segment, showing hair tufts; 555e, 
basal segments of antenna; 555f, inner 
pectination from one of basal segments 
of antennal shaft, showing attachment 
of modified setae (greatly enlarged). 


555. leithella 


444 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 556—-558.—Matss. 


556. Ozamia lucidalis (Walker) ; 556a, gnathos, 
ventral view; 556b, elements of trans- 
tilla; 556c, anellus; 556d, aedeagus; 
556e, eighth abdominal segment, show- 
ing hair tufts. 

557. Ozamia fuscomaculella (Wright); 5957a, 
elements of transtilla; 557b, anellus; 
557c, aedeagus; 557d, basal segments 
of antenna. 

558. Ozamia punicans Heinrich: 558a, ele- 
ments of transtilla; 558b, anellus; 
558c, aedeagus. 


556. lucidalis 


558. punicans 


557. fuscomaculella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 445 


Fieurss 559, 560.—Matzs. 


559. Ozamia fuscomaculella clarefacta Dyar; 
559a, elements of transtilla; 559b, 
anellus; 559c, aedeagus; 559d, eighth 
abdominal segment, showing hair tufts; 
559e, lateral view of genitalia with one 
harpe omitted. 

560. Ozamia hemilutella Dyar; 560a, elements 
of transtilla; 560b, anellus; 560c, aede- 
agus; 560d, eighth abdominal segment, 
showing hair tufts. 


559. clarefacta 


560. hemilutella 


446 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 561—564.— Mats. 


561. Illatila gurbyris Dyar, aedeagus omitted, 
vinculum foreshortened; 561a, aede- 
agus; 561b, lateral view of tegumen 
showing serrate production from base; 
561c, gnathos; 561d, sternite of eighth 
abdominal segment. 

562. Lascelina canens Heinrich, new species, 
type; 562a, aedeagus; 562b, sternite of 
eighth abdominal segment. 

563. Meitlphestia simplicula (Zeller); 563a, 
uncus; 563b, aedeagus; 563c, gnathos 
and anellus; 563d, tufts of eighth ab- 
dominal segment. 

564. Selga arizonella (Hulst); 564a, aedeagus; 
564b, gnathos; 564c, elements of trans- 
tilla; 564d, anellus. 


~ 563. 


simplicula 


564.arizonella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 447 


Fieures 565-567.—Matgs. 


565. Diatomocera tenebricosa (Zeller), example 
from French Guiana, placed here to 
show affinities in genitalic structure of 
Diatomocera and Entmemacornis; 565a, 
aedeagus; 565b, gnathos; 565c, trans- 
tilla; 565d, anellus; 565e, tufts of 
eighth abdominal segment. 

566. Entmemacornis proselytes Dyar; 566a, 
aedeagus; 566b, gnathos; 566c, trans- 
tilla; 566d, anellus; 566e, tufts of 
eighth abdominal segment; 566f, basal 
segments of antenna, denuded. 

567. Entmemacornis pulla Heinrich, new spe- 

Sky : Z| iy ‘ cies, type; 567a, aedeagus; 567b, 

nebricosa =, | ANG gnathos; 567c, transtilla; 567d, anellus; 

iS 567e, tufts of eighth abdominal seg- 
ment. 


i 


567. pulla 


448 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 568-570.— Mates. 


568. Cayennia rufitinctalis Hampson; 568a, 
aedeagus; 568b, gnathos; 568c, trans- 
tilla; 568d, anellus. 

569. Rioja neza Heinrich, new species, type; 
569a, aedeagus; 569b, tergite and ster- 
nite of eighth abdominal segment; 569c, 
gnathos; 569d, transtilla; 569e, anellus. 

570. Moerbes dryopella (Schaus), type; 570a, 
aedeagus; 570b, gnathos; 570c, ele- 
ments of transtilla; 570d, anellus; 
570e, tergite and sternite of eighth 
abdominal segment. 


569. nexa 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 449 


572b. 


To 


d 


573. paenaista 


300329—56——30 


Fieures 571-573.—Matzs. 


571. Moerbes alveolella (Ragonot); 571a, aede- 
agus; 571b, gnathos; 571c, transtilla; 
571d, anellus; 57le, tergite and ster- 
nite of eighth abdominal segment. 

572. Moerbes emendata Heinrich, new species; 
572a, aedeagus; 572b, gnathos; 572c, 
transtilla; 572d, anellus; 572e, tergite 
and sternite of eighth abdominal seg- 
ment. 

573. Moodnopsis perangusta (Dyar), type; 
573a, aedeagus; 573b, gnathos; 5738c, 
transtilla; 573d, anellus; 573e, tergite, 
sternite and hair tufts of eighth ab- 
dominal segment. 


450 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 574-577.—MA.LEs. 


574. Moodnopsis inornatella (Ragonot); 574a, 
aedeagus; 574b, gnathos; 574c, trans- 
tilla; 574d, anellus; 574e, tergite, ster- 
nite, and tufts of eighth abdominal 
segment. 

575. Moodnopsis parallela Heinrich, new spe- 
cies; 575a, aedeagus; 575b, gnathos; 
575c, transtilla; 575d, anellus; 575e, 
tergite, sternite, and tufts of eighth 
abdominal segment. 

576. Moodnopsis <inveterella (Dyar); 576a, 
aedeagus; 576b, gnathos; 576c, trans- 
tilla; 576d, anellus; 576e, tergite, 
sternite, and tufts of eighth abdominal 
segment. 

577. Moodnopsis portoricensis Heinrich, new 
species, uncus, gnathos, and tegumen 
of male genitalia; 577a, transtilla. 


inornatella 


575.parallela 
575c. 


577. 
portoricensis 


2 
TAS i 


BAe 


VW 576d. 
576.inveterella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 451 


Figures 578-580.— Mats. 


578. Euzophera cinerosella (Zeller), aedeagus 
omitted; 578a, aedeagus; 578b, tufts of 
eighth abdominal segment; 578c, 
gnathos; 578d, transtilla; 578e, anellus. 

579. Euzophera semifuneralis (Walker); 579a, 
gnathos; 579b, transtilla; 579c, anellus; 
579d, aedeagus, 

580. Euzophera osiricolorella Hulst; 580a, 
aedeagus; 580b, gnathos; 580c, anellus; 
580d, transtilla. 


its 
579. semifuneralis 


580. ostricolorella 


452 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 581-584.— Mates. 


581. Euzophera nigricantella Ragonot; 581a, 
gnathos; 581b, transtilla, 581c, anellus; 
581d, aedeagus. 

582. Prosoeuzophera impletella (Zeller), para- 
type (in BM); 582a, aedeagus; 582b, 
gnathos; 582c, transtilla; 582d, anellus. 

583. Eulogia ochrifrontella (Zeller); 583a, 
aedeagus; 583b, tufts of eighth abdom- 
inal segment of male; 583c, gnathos; 
583d, elements of transtilla; 583e, 
anellus. 

584. Exuperius negator Heinrich, type; 584a, 
aedeagus. 


581. nigricantella. 


SS 


5830. 583.ochrifrontella 


negator 


584. 5 583e. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 4538 


Figures 585-587.— Mates. 


585. Edulica compedella (Zeller); 585a, gna- 
thos; 585b, transtilla; 585c, aedeagus, 

586. Farnobia quadripuncta (Zeller); 586a, 
anellus; 586b, gnathos; 586c; transtilla ' 
586d, aedeagus; 586e, sternite of eighth 
abdominal segment. 

587. Gennadius junctor Heinrich, new species; 
587a, sternite of eighth abdominal seg- 
ment; 587b, aedeagus with penis ex- 
truded; 587c, aedeagus with penis in 
normal position; 587d, gnathos; 587e, 
transtilla; 587f, anellus, 


586. quadripunc a 


587. junctor 


454 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 588-591.—MaA.ss. 


588. Ephestiodes erasa Heinrich, new species, 
type; 588a, anellus; 588b, gnathos; 
588c, elements of transtilla; 588d, 
aedeagus. 

589. Ephestiodes gilvescentella Ragonot; 589a, 
aedeagus; 589b, gnathos; 589c, ele- 
ments of transtilla; 589d, anellus; 
589e, tufts of eighth abdominal 
segment. 

590. Ephestiodes infimella Ragonot; 590a, 
gnathos; 590b, elements of transtilla; 
590c, anellus; 590d, aedeagus. 

591. Eurythmia coloradella Hulst, a synonym 
of Ephestiodes erythrella Ragonot; 
591a, gnathos; 591b, elements of trans- 
tilla; 591c, anellus; 591d, aedeagus. 


591. coloradella 


592. mignonella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 455 


592a. 


Ficures 592-594.—Matzs. 


592. Ephestiodes mignonella Dyar; 592a, gna- 
thos, 592b, elements of transtilla; 
592c, anellus; 592d, aedeagus. 

593. Ephestiodes lucidibasella Ragonot, type; 
593a, gnathos; 593b, elements of 
transtilla; 593c, anellus; 593d, aedea- 
gus. 

594, Azaera muciella Schaus; 594a, gnathos; 
594b, elements of transtilla; 594c, 
anellus; 594d, aedeagus. 


456 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 595-598.—Mates. 


595. Micromescinia pygmaea Dyar, aedeagus 
omitted; 595a, anellus; 595b, gnathos; 
595c, transtilla; 595d, aedeagus; 595e, 
tergite and sternite of eighth abdomi- 
nal segment. 

596. Ephestiodes stictella (Hampson), one 
harpe detached and aedeagus omitted; 
596a, aedeagus; 596b, gnathos; 596c, 
transtilla; 596d, tufts of eighth abdomi- 
nal segment. 

597. Ephestiodes noniella Dyar, 597a, aedea- 
gus; 597b, gnathos; 597c, transtilla; 
597d, anellus; 597e, tergite and sternite 
of eighth abdominal segment. 

598. Vezina parasitaria Heinrich, new species, 
type; 598a, anellus; 598b, gnathos; 
598c, elements of transtilla; 598d, ae- 
deagus. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 457 


599. 


ostrinella 


600. 


bisinuella 


ae, 


601. 


biviella 


Fieurres 599-601.—Matgzs. 


599. Moodna ostrinella (Clemens); 599a, gna- 
thos; 599b, elements of transtilla; 
599c, aedeagus; 599d, tufts of eighth 
abdominal segment; 599e, basal seg- 
ment of antenna. 

600. Moodna bisinuella Hampson; 600a, ae- 
deagus; 600b, gnathos; 600c, elements 
of transtilla; 600d, anellus. 

601. Manhatta biviella (Zeller); 601a, aedea- 
gus; 601b, gnathos; 601c, transtilla; 
601d, anellus; 60le, tufts on eighth 
abdominal segment. 


458 UNITED STATES NATIONAL MUSEUM! BULLETIN 207 


Figures 602-604.—Matss. 


602. Manhaita setonella (McDunnough), speci- 1. His ON 
: : Leen 


men from Utah; 602a, gnathos; 602b, 
transtilla; 602c, anellus; 602d, aedea- 
gus. 


603. Verina supplicella (Dyar); 603a, tergite, “602R.S 
sternite, and tufts of eighth abdominal Lorall i 
segment; 603b, aedeagus; 608c, gna- Sexonene css 
thos; 603d, elements of transtilla; 602c. 


603e, anellus. 

604. Vagobanta divergens (Butler); 604a, ter- 
gite and sternite of eighth abdominal 
segment; 604b, aedeagus; 604c, gna- 
thos; 604d, elements of transtilla; 
604e, anellus. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 459 


Ficures 605-608.—Matgzs. 


605. Vitula edmandsae (Packard) ; 605a, aedea- 
gus; 605b, gnathos; 605c, transtilla; 
605d, anellus; 605e, tergite, sternite, 
and tufts of eighth abdominal segment. 

606. Vitula pinei Heinrich, new species; 606a, 
gnathos; 606b, transtilla; 606c, anel- 
lus; 606d, aedeagus. 

607. Vitula lugubrella (Ragonot), harpe. 

608. Moodnella paula Heinrich, type; 608a, 
aedeagus; 608b, gnathos; 608c, trans- 
tilla; 608d, anellus. 


6050. 605. edmandsae 


608. paula 


460 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 609, 610.—Mates. 


609. Volatica trinitatis Heinrich, new species; 
609a, aedeagus; 609b, gnathos, 609c, 
transtilla; 609d, anellus; 609e, tergite 
and sternite of eighth abdominal seg- 
ment. 

610. Volatica pachytaeniella (Ragonot); 610a, 
gnathos; 610b, transtilla; 610c, anellus; 
610d, aedeagus. 


610. pachytaeniella 


610c. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 461 


Fiaures 611-614.—Matzs. 


611. Caudellia nigrella (Hulst), type; 611a, 
gnathos plus transtilla; 611b, anellus; 
61lc, aedeagus. 

612. Caudellia declivella (Zeller); 612a, aedea- 
gus; 612b, gnathos; 612c, transtilla. 

613. Caudellia apyrella Dyar, type; 613, ae- 
deagus; 613b, elements of transtilla. 

614. Caudellia colorella (Dyar), one harpe de- 
tached and aedeagus omitted; 614a, 
aedeagus; 614b, transtilla. 


612.declivella 


613. apyrella 


Bow 


élécolbralla 


462 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 615-619.— Mates. 


615. Caudellia clara Heinrich, type; 615a, 
gnathos; 615b, eighth abdominal seg- 
ment, showing dorsal tufts. 

616. Sosipatra rileyella (Ragonot) ; 616a, gna- 
thos; 616b, transtilla; 616c, anellus; 
616d, aedeagus. 

617. Sosipatra micaceella (Hampson); 6172, 
aedeagus; 617b, gnathos; 617c, trans- 
tilla; 617d, eighth abdominal segment, 
showing dorsal tufts. 

618. Sosipatra anthophila (Dyar), type; 618a, 
aedeagus; 618b, gnathos with side 
sketch showing apical projection in 
lateral view; 618c, transtilla; 618d, 
anellus. 

619. Sosipatra thurberiae (Dyar), type; 6198, 
aedeagus; 619b, gnathos; 619c, trans- 
tilla; 619d, anellus. 


l 


ye RES) 


He 
Sa 


617. micaceella 


618. 
anthophila 


AMERICAN 


620827 


majorella 


ed 622.innoxia 


622c. 


MOTHS OF THE SUBFAMILY PHYCITINAE 463 


Figures 620-623.—Ma.gs. 


620. Sosipaira majorella (Dyar), aedeagus 
omitted; 620a, aedeagus; 620b, gna- 
thos; 620c, transtilla; 620d, tergite and 
sternite of eighth abdominal segment. 

621. Sostpatra nonparilella (Dyar), type, ae- 
deagus omitted; 621a, gnathos; 621b, 
transtilla; 621c, anellus; 621d, aedea- 
gus. 

622. Ribua innoxia Heinrich, aedeagus omit- 
ted; 622a, aedeagus; 622b, apical pro- 
jection of gnathos; 622c, anellus; 622d, 
tergite, sternite, and dorsal tufts of 
eighth abdominal segment. 

623. Ribua contigua Heinrich, new species, 
aedeagus omitted; 623a, aedeagus; 
623b, fused gnathos and transtilla. 


464 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fia@ures 624-627.— Matss. 


624. Plodia interpunctella (Hiibner); 624a, 
aedeagus; 624b, transtilla. 

625. Plodia dolorosa Dyar; 625a, aedeagus; 
625b, transtilla; 625c, tergite, sternite, 
and dorsal tufts of eighth abdominal 
segment. 

626. Anagasta kiuhniella (Zeller), aedeagus 
omitted; 626a, aedeagus; 626b, gna- 
thos; 626c, elements of transtilla; 626d, 
anellus; 626e, tergite, sternite, and 
dorsal tufts of eighth abdominal seg- 
ment. 

627.—Cabnia myronella Dyar, aedeagus omit- 
ted; 627a, aedeagus; 627b, gnathos; 
627c, transtilla; 627d, anellus; 627e, 
lateral view of head. 


624.2" 
interpunctella 


626. kuhniella 


627. myronella 


AMERICAN MOTHS OF 


629a. Ss 
629.cautella 


630. figulilella 


THE SUBFAMILY .PHYCITINAE 465 


Figures 628-630.—Matzs. 


628. Ephestia elutella (Hiibner); 628a, aedea- 
gus; 628b, gnathos; 628c, transtilla; 
628d, eighth abdominal segment, show- 
ing dorsal tufts. 

629. Ephestia cautella (Walker); 629a, aedea- 
gus; 629b, gnathos; 629c, transtilla; 
629d, anellus; 629e, dorsal view of 
uncus and tegumen; 629f, eighth ab- 
dominal segment, showing dorsal tufts. 

630. Ephestia figulilella Gregson; 630a, aedea- 
gus; 630b, gnathos; 630c, transtilla: 
630d, anellus. 


466 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 631-634.—Mat.es 


631. Varneria postremella Dyar, one harpe de- 
tached ‘and aedeagus omitted; 631a, 
transtilla; 631b, aedeagus; 631c, eighth 
abdominal segment, showing dorsal 
tufts. 

632. Varneria dubia Heinrich, new species, 

aedeagus omitted; 632a, aedeagus; 
632b, gnathos; 632c, transtilla; 632d, 
eighth abdominal segment, showing 
dorsal tufts. 

633. Eurythmia hospitella (Zeller); 633a, ae- 
deagus; 633b, gnathos; 633c, transtilla; 
633d, a slight modification of the 
central projection of transtilla drawn 
from an example of angulella Ely 
(-diffusella Ely); 633e, anellus; 633f, 
sclerotizations of eighth abdominal 
segment. 

634. Hrelieva quantulella (Hulst); 6342, aede- 
agus; 634b, gnathos; 634c, transtilla; 
634d, eighth abdominal segment, show- 
ing dorsal tufts. 


Sat 


63lc. 


RE 


VJ % 
guantutella 


634b. 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 467 


635. conops Z\ 
~ 635c. 


637. ~ 


apianella 


Figures 635-637.—MA.zs 


635. Rabiria conops (Dyar), type, aedeagus 
omitted; 635a, gnathos; 635b, eighth 
abdominal segment; 635c, transtilla; 
635d, aedeagus. 

636. Microphycita titillella Dyar, aedeagus 
omitted; 636a, gnathos; 636b, trans- 
tilla; 636c, aedeagus and anellus. 

637. Nicetiodes apianella Schaus, type; 637a, 
gnathos; 637b, transtilla; 637c, anellus; 
637d, aedeagus; 637e, eighth abdom- 
inal segment showing tufts. 


468 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 638-644.—FEMALE GENITALIA. 


638. Cryptoblades rutilella Zeller. 

639. Cryptoblades gnidiella (Milliére). 

640. Acrobasis tumidella (Zincken). 

641. Acrobasis indigenella (Zeller). 

642. Acrobasis vaccinii Riley. 

643. Acrobasis tricolorella Grote. 

644. Acrobasis juglandis (LeBaron), specimen 
reared from pecan. 


642. : 644, 


indiginella vaccinii tricolorella juglandis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 469 


Fieures 645-650.—FEMALE GENITALIA. 


645. Acrobasis tumidulella (Ragonot), type. 
646. Acrobasis comptella Ragonot. 

647. Rhodophaea caliginella (Hulst), type. 
648. Rhodophaea supposita (Heinrich). 

649. Rhodophaea advenella (Zincken). 

650. Trachycera pallicornella (Ragonot). 


16 a otella 


tumidulella 


649. 


supposita .- _—_ advenella pallicornella 


470 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 651-654.— FEMALE GENITALIA. 


651. Mildrizia constitutionella Dyar; 651a, sig- 
num of bursa, enlarged. 

652. Anabasis ochrodesma (Zeller). 

653. Sematoneura atrovenosella (Ragonot). 

654. Hypsipyla dorsimacula (Schaus). 


653. atrovenosella 


654 


. dorsimacula 


652.ochrodesma 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 471 


Figures 655-657.—FEMALE GHNITALIA. 


655. Hypsipyla grandella (Zeller), 
656. Hypsipyla pagodella Ragonot. 
657. Hypsipyla ferrealis (Hampson). 


655.grandella 


656. pagodella 657. ferrealis 


472 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 658-660.— FEMALE GENITALIA. 


658. Hemiptilocera chinographella Ragonot. 
659. Hemiptilocera jocarella (Schaus), type. 
660. Hemiptilocera bigrana (Zeller). 


658,chinographella 


659. 3 
jecarella 660. bigrana 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 473 


Fieures 661-663.—FEMALE GBNITALIA. 


661. Hemiptilocera exoleta (Zeller), type; 66la, 
dorsal view of eighth abdominal seg- 
ment. 

662. Hemiptilocera letharda (Schaus), showing 
to the side of bursa a greatly enlarged 
figure of the spines of signum. 

663. Crocidomera stenopteryx (Dyar), specimen 
from type locality. 


661. exoleta 


662. 
letharda 


663. stenopteryx 


300329—56——31 stots Suiibaihicgs = =" 


474 UNITED STATES NATIONAL MUSEUM BULLETIN 207. 


Ficures 664—-667.— FEMALE GENITALIA. 


664. Crocidomera turbidella Zeller. 

665. Crocidomera fissuralis (Walker), type. 

666. Cuniberia subtinctella (Ragonot). 

667. Adanarsa iniransitella (Dyar), showing a 
great enlargement of signum to one side 
of bursa. 


666. 
subtinctella 


"ences anorenens 


664. turbidella 


667. intransitella 665. fissuralis 


670. 


annuliferella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 475 


1 668. 


definitella 


671. hystriculella 


669.grisella 


Fieures 668-671.—FEMALH GENITALIA. 


668. Hypargyria definitella (Zeller). 

669. Bertelia grisella Barnes and McDun- 
nough. 

670. Chararica annuliferella (Dyar). 

671. Chararica hystriculella (Hulst). 


476 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 672-675.— FEMALE GENITALIA. 


672. Myelopsis subtetricella (Ragonot), type. 

673. Myelopsis coniella (Ragonot). 

674. Myelopsis fragilella (Dyar), a2 synonym 
of M. alatella (Hulst). 

675. Myelopsis minutularia (Hulst), type. 


673. coniella 675. minutularia 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 477 


676. 
bistriatella 


677. decolor 


678, ceratoniae 679. univitella 


Ficurres 676-679.—FEMALE GENITALIA. 


676. Apomyelois bistriatella (Hulst). 
677. Ectomyelois decolor (Zeller). 
678. Pctomyelois ceratoniae (Zeller). 
679. Anypsipyla univitella Dyar. 


478 UNITED STATES NATIONAL MUSEUM BULLETIN. 207 


Fraures 680-684.—FSMALE GENITALIA. 


680. Ectomyelois muriscis (Dyar). 

681. Ectomyelois furvidorsella (Ragonot), type. 
682. Ectomyelois zeteki Heinrich, new species. 
683. Paramyelois transitella (Walker). 

684. Myelois cribrella (Hiibner). 


680. muriscis 


furvidorsella 
684. cribrella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 479 


Fiaurses 685-691.—FEMALE GENITALIA. 


685. Diatomocera dosia (Dyar), paratype; 
685a, enlargement of signa. 

686. Diatomocera tenebricosa (Zeller); 686a, 
enlargement of signa. 

687. Diatomocera excisalis (Hampson), para- 
type from Cayenne, French Guiana 
(in USNM), signa in bursa copulatrix. 

688. Diatomocera extracta Heinrich, new spe- 
cies, bursa copulatrix. 

689. Diatomocera mochlophleps (Dyar), bursa 
copulatrix; 689a, enlargement of sig- 
num in bursa. 

690. Pseudodivona santa-maria Dyar, type, 
bursa copulatrix. 

691. Pseudodivona carabayella Dyar, specimen 
from Incachaca, Bolivia. 


690. 


santa-maria 


687. 
excisalis 


691. 
carabayella 


689. mochlophleps 


’ 

1 

At 
ah 


Nay 


688. extracta 


480 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 692-699.— FEMALE GENITALIA. $ : 
rubrizonalis 


693. 


692. Pseudocabima expunctriz (Dyar and Hein- he 
euzopherella 694. 


rich), signa of female genitalia. 

693. Pseudocabima euzopherella (Dyar), signa 
of female genitalia. 

694. Pseudocabima rubrizonalis (Hampson), 
signa of female genitalia. 

695. Pseudocabima perrensiella (Ragonot), 
type; 695a, enlargement of signa. 

696. Pseudocabima castronalis Heinrich, new 
species, paratype; 696a, enlargement 
of signa. 

697. Pseudocabima guianalis Heinrich, new 
species, signa of female genitalia. 

698. Pseudocabima arizonensis Heinrich, new 
species, signa of female genitalia. 

699. Pseudocabima nigristrigella (Ragonot), 
specimen in BM; 699a, enlargement 
of signa. 


arizonensis 


698. 


guianalis 


695, castronalis 


perrensiella sie nee 
nigristrigella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 481 


701. 


majorina 


700.celiella 


702. 703. 
fulgidula stictoneurella egenella 


300329—56——32 


Ficures 700-704. FEMALE GENITALIA AND 
(a) ENLARGEMENT OF SIGNA. 


700. Hyalospila celiella Schaus. 

701. Hyalospila majorina Heinrich, new spe- 
cies. 

702. Hyalospila fulgidula Heinrich, new spe- 
cies. 

703. Hyalospila stictoneurella Ragonot. 

704. Hyalospila egenella (Ragonot), type. 


482 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieaures 705-709. FaMALE GENITALIA. 


705. Hyalospila clevelandella (Dyar), paratype 
from Porto Bello, Panamdé; 705a, 
enlargement of signum patch. 

706. Hyalospila clevelandella (Dyar), speci- 
mens from México and Guatemala; 
706a, enlargement of signum. 

707. Hyalospila angulineella (Schaus), type; 
707a, enlargement of signa. 

708. Hyalospila semibrunneella (Ragonot), 
type. 

709. Hyalospila xanthoudemia (Dyar); 709a, 
enlargement of signum. 


106a. 


clevelandella ? 


<= 


709a. 


709, 


708. semibrunncella Je Roudsmra 


ignobilis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 483 


pellucens 


ahemora 


Figures 710-714.— FEMALE GENITALIA. 


710. Fundella agapella Schaus, type. 

711. Fundella argentina Dyar; 711a, armature 
of genital opening (in South American, 
Argentine, and Brazilian specimens). 

712. Fundella ignobilis Heinrich, new species. 

713. Fundella pellucens Zeller; 713a, dorsal 
view of eighth segment collar. 

714. Fundella ahemora Dyar. 


484. UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiqures 715-719.— FEMALE GENITALIA. 


715. Difundella corynophora Dyar; 715a, dor- 
sal view of collar of eighth abdominal 
segment; 715b, ventrolateral pockets in 
intersegmental area between seventh 
and eighth abdominal segments, shown 
from dorsal view. 

716. Difundella distractor Heinrich, new spe- 
cies. 

717. Difundella tolerata Heinrich, new species; 
717a, invaginated, sclerotized, dorsal 
shield of seventh segment of abdomen. 

718. ?Coptarthria dasypyga (Zeller), example 
from Costa Rica; 718a, enlargement of 
signum. 

719. Dasypyga alternosquamella Ragonot. 


distractor 


718. dasypyga ? 719, alternosqaamella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 485 


Figurts 720-725.—FEMALE GENITALIA. 


720. Promylea lunigerella glendella (Dyar), 
showing enlargement of signum beside 
bursa; 721a, dorsal view of sclerotized 
apron and pocket attached to collar. 

721, Promylea lunigerella lunigerella Ragonot, 
specimen from Duncans, Vancouver 
Isl., showing enlargement of signum 
beside bursa; 721la, dorsal view of 
sclerotized apron and pocket attached 
to collar; 721b, lateral view of same, 
also showing genital opening of ductus 
bursae. 

722. Promylea druceii (Ragonot), type; 722a, 
dorsal view of sclerotized apron and 
pocket attached to collar. 

723. Promylea mindosis Dyar; 728a, dorsal 
view of sclerotized apron and pocket 
attached to collar. 

724. Promylea dasystigma Dyar; 724a, dorsal 
view of sclerotized apron and pocket 
attached to collar. 

725. Scorylus cubensis Heinrich, new species. 


721. 


glendella lunigerella 


722. 
druceii 723. mindosis 724. deasystigma 


725. cubensis 


486 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 726-733.— FEMALE GENITALIA. 


726. Anadelosemia fifria Dyar, showing en- 
largement of signum to side of bursa. 

727. Anadelosemia base Dyar. 

728. Anadelosemia obstitella (Schaus), type, 
showing enlargement of signum beside 
bursa; 728a, dorsal view of eighth- 
segment collar. 

729. Anadelosemia texanella (Hulst), dorsal 
view of eighth-segment collar. 

730. Anadelosemia condigna Heinrich, new spe- 
cies, dorsal view of eighth-segment 
collar. 

731. Rampylla lophotalis Heinrich, new spe- 
cies; 731a, dorsal views of ovipositor 
and eighth-segment collar. 

732, Rampylla polydectella  (Schaus), type; 
732a, dorsal view of eighth-segment 
collar; 732b, invaginated, sclerotized 
pocket from seventh abdominal seg- 
ment. 

733. Rampylla subcaudata (Dyar), specimen 
from Quirigus, Guatemala; 733a, dor- 
sal views of eighth-segment collar and 
sclerotized pocket of seventh segment. 


726. fifria 
729. 


texanella 


728. obstitella 


727. base 


733. 


subcaudata 


732. 
731. lophotalis polydectella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 487 


735. caricae 


736. 


columnella 


737, azonaxsalis 


Ficures 734-738.—FEMALE GENITALIA, 


734. 
735. 
736. 
737. 
738. 


Davara interjecta Heinrich, new species. 
Davara caricae (Dyar). 

Davara columnella (Zeller), type. 
Davara azonazsalis (Walker), type. 
Davara nerthella (Schaus), type. 


488 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 739-744.— FEMALE GENITALIA. 


739. Davara rufulella (Ragonot). 

740. Sarasota furculella (Dyar), specimen 
from Cuba. 

741. Atheloca subrufella (Hulst), type. 

742. Praedonula almonella (Dyar); 742a, dor- 
salview of eighth-segment collar, 
showing genital opening and its 
attachments. 

743. Piesmopoda montella Schaus, type. 

744. Piesmopoda trichomata (Zeller), cotype. 


740.furculella 


742. almonella 743. montella trichomata 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 489 


745. xanthomera 746.flavicans 748. xanthopol ys 


750. parva 
751 apocerastes 


752. semirufella 


745 


746. 
747. 
748. 
749. 
750. 
751. 
752. 


Ficures 745-752.— FEMALE GENITALIA. 


. Piesmopoda sxanthomera Dyar, type, 
detail to the side of collar shows its 
invagination; similar details shown in 
figures 746 to 750. 

Piesmopoda flavicans (Zeller). 

Piesmopoda ragonoti (Dyar). 

Piesmopoda xanthopolys Dyar, type. 

Piesmopoda isabella (Dyar). 

Piesmopoda parva Heinrich. 

Piesmopoda apocerastes Dyar. 

Piesmopoda semirufella (Zeller). 


490 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


. Fieures 753-758.— FEMALE GENITALIA. 


. 753. Peadus subaquilellus (Ragonot), type, 
with enlargement of signum shown to 
the side of bursa. 

754. Peadus burdettellus (Schaus). 

755. Peadus dissitus Heinrich, new species. 

756. Gabinius paulsoni (Ragonot), specimen 
from ‘“‘Chile, Silva.’ 

757. Ceracanthia mamella (Dyar), type. 

758. Drescoma cyrdipsa Dyar. 


755. dissitus 


756. paul soni 
TST. momella 758. cyrdipsa 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 491 


761. drucella 


760.alpha 


762: subelisa 764..nubilella 765. pogerythrus 


761. 


762. 


763. 
764. 


765. 


Fiaures 759-765.— FEMALE GENITALIA. 


759. 
760. 


Megarthria beta Heinrich, type. 
Megarthria alpha Heinrich, type; 760a, b, 
collar in dorsal view showing variations 
in the apron. 
Drescoma drucella Dyar, a synonym of 
Drescomopsis soraella (Druce). 
Drescomopsis subelisa Dyar, a synonym 
of Drescomopsis soraella (Druce), with 
signum shown in full lateral view. 
Monoptilota pergratialis (Hulst), bursa 
copulatrix of female genitalia. 
Monoptilota nubilella Hulst, paratype, a 
synonym of M. pergratialis (Hulst). 
Zamagiria pogerythrus Dyar. 


492 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 766—770.— FEMALE GENITALIA. 


766. Zamagiria australella (Hulst). 
767. Zamagiria laidion (Zeller). 

768. Zamagiria ipsetona Dyar. 

769. Magiriopsis denticosella (Dyar). 
770. Anegcephaleszs arctella (Ragonot). 


australella 


768. ipsetona - 769. denticosella 770. arctella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 493 


Ficures 771-775.— FEMALE GENITALIA. 


771. Ancylostomia stercorea (Zeller); 771a, 
ventral view of ductus bursae and 
eighth-segment collar with membrane 
of seventh segment removed. 

772. Adelperga cordubensiella (Ragonot), type. 

773. Caristanius decoloralis (Walker); 773a, 
ventral view of eighth-segment collar 
and apical portion of ductus bursae. 

774. Caristanius guatemalellus (Ragonot), 
type. 

775. Caristanius pellucidellus (Ragonot) ; 775a, 
ventral view of apical part of ductus 
bursae and eighth-segment collar; 
775b, dorsal view of eighth-segment 
collar and its attachments to ductus 
bursae. 


“y j 4 74 
ff af 
i ie 


I # 


774. 


775. pellucidellus 


temalellus 


494 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 766—780.— FEMALE GENITALIA. 


776. Pima boisduvaliella (Guénée), bursa cop- 
ulatrix, apical part of ductus bursae, 
and dorsal outline of eighth-segment 
collar of female genitalia. 

777. Pima albiplagiatella (Packard). 

778. Pima albocostalialis (Hulst), bursa copu- 
latrix, apical part of ductus bursae, and 
dorsal outline of eighth-segment collar 
of female genitalia. 

779. Pima albocostalialis (Hulst), a specimen 
showing extent of variation in patches 
of bursa and shape of eighth-segment 
collar. 

780. Pima vwidella (McDunnough), specimen 
from Aweme, Manitoba, bursa copula- 
trix, apical part of ductus bursae, and 
dorsal outline of eighth-segment collar 
of female genitalia. 


780. vividella 779. albocostalialis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 495 


Ficures 781-783.— FEMALE GENITALIA 


781. Pima granitella (Ragonot). 

782. Pima parkerella (Schaus). 

783. Pima fosterella Hulst, specimen from 
Arizona, bursa copulatrix, apical part 
of ductus bursae, and dorsal outline of 
eighth-segment collar of female geni- 
talia. 


7; 781. 


granitella 


782. parkerella 783. fosterella 


496 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 784—788.—FEMALE GENITALIA. 


784. Interjectio columbiella (McDunnough), 
specimen from Pullman, Wash, 

785. Interjectio ruderella (Ragonot), type. 

786. Olybria aliculella (Hulst). 

787. Olybria furciferella (Dyar). 

788. Oreana unicolorella (Hulst). 


786. aliculella 


= 788. unicolorella 


787. furciferella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 497 


Figures 789-790.—FEMALE GENITALIA. 


789. Ambesa lallatalis (Hulst); 789a, dorsal 
view of eighth-segment collar. 
790. Ambesa laetella Grote. 


790. laetella 


789. lallatalis 


498 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 791-793.—FEMALE GENITALIA. 


791. Ambesa walsinghami (Ragonot). 

792. Ambesa walsinghami (Ragonot), figured 
from type of its synonym, A. monodon 
Dyar. 

793. Ambesa walsinghami mirabella Dyar. 


792.walsinghami 


791. walsinghami 793. mirabella 


796. marginea 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 499 


794. actualis 


797. bistriatella 


798. nigrovittella 


Ficures 794-798.—FEMALE GENITALIA. 


794. Catastia actualis (Hulst). 

795. Catastia incorruscella (Hulst), specimen 
from Arizona. 

796. Catastia marginea (Schiffermiiller). 

797. Catastia bistriatella (Hulst), specimen 
from type locality. 

798. Immyrla nigrovittella Dyar. 


500 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 799-803.—FEMALE GENITALIA. 


799. Salebriacus odiosellus (Hulst); 799a, 
dorsal view of eighth-segment collar. 

800. Salebriaria annulosella (Ragonot), speci- 
men from Burnet County, Tex. 

801. Salebriaria tenebrosella (Hulst), type. 

802. Salebriaria turpidella (Ragonot). 

803. Salebriaria tenebrosella (Hulst), figured 
from large example. 


tenebrosella 


NG 


799. odiosellus 803. tencbrosella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 501 


fruc 


807. pasadamia 


808. jugosella 


Figures 804-808.—FEMALE GENITALIA. 


804. Salebriaria pumilella (Ragonot). 

805. Salebriaria fructetella (Hulst), type. 

806. Quasisalebria admizta Heinrich, paratype 
from type locality. 

807. Ortholepis pasadamia (Dyar); 807a, dor- 
sal view of eighth-segment collar; 807b, 
dorsal view of junction of bursa and 
ductus bursae. 

808. Ortholepis jugosella Ragonot; 808a, dorsal 
view of eighth-segment collar. 


502 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaurts 809-812.— FEMALE GENITALIA. 


809. Polopeustis annulatella (Zetterstedt); 
809a, dorsal view of eighth-segment 
collar. 

810. Polopeustis arctiella (Gibson), specimen 
from Labrador; 810a, dorsal view of 

; eighth-segment collar. 

811. Glyptocera consobrinella (Zeller). 

812. Meroptera pravella (Grote). 


611, consobrinella 612. pravella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 503 


Figures 813-817.— FEMALE GENITALIA. 


813. Meroptera abditiva Heinrich, new species, 
paratype from type locality, with 
eighth-segment collar and ovipositor 
omitted. 

814. Meroptera abditiva Heinrich, new species, 
paratype from type locality. 

815. Nephopteryx rhenella (Zincken). 

816. Meroptera mirandella Ragonot. 

817. Meroptera cviatella Dyar. 


815. rhenella 


816. mirandella 


504 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 818-822.—FEMALE GENITALIA. 


818. Nephopteryx delassalis Hulst, type. 

819. Nephopteryx fernaldi (Ragonot). 

820. Nephopteryx vetustella (Dyar). 

$21. Nephopteryx subfuscella (Ragonot), fig- 
ured from type of its synonym Salebria 
semiobscurella Hulst. 

822. Nephopteryz dammerst Heinrich. new 
species. 


818. delassalis 


820. vetustella 


821. subfuscella 822 . dammersi 


sts 


826. subcaesiella 


800329—56——-33 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 505 


827. virgatella 


Figures 823-827.—FEMALE GENITALIA. 


823. Nephopteryx carneella Hulst, reared speci- 
men from Maine. 

824. Nephopteryz uvinella (Ragonot). 

825. Nephopteryx inconditella (Ragonot). 

826. Nephopteryx subcaesiella (Clemens), 

827. Nephopteryx virgatella (Clemens). 


506 UNITED STATES NATIONAL MUSEUM BULLETIN’ 207 


Figures 828-831.—FEMALE GENITALIA. 


828. Nephopteryx termitalis (Hulst), figured 
from type of its synonym Salabria 
levigatella Hulst. 

829. Nephopteryx basilaris Zeller. 

830. Nephopteryx celtidella (Hulst). 

831. Nephopieryx bifasciella Hulst. 831a, ab- 
normal specimen from Yuma, Ariz. 


830. celtidella 


<B% 


831. bifasciella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 507 


Figures 832-836.—FEMALE GENITALIA. 


832. Nephopteryx rubrisparsella (Ragonot). 
833. Nephopteryx bisra Dyar, type. 

834. Tlascala reductella (Walker). 

835. Nephopteryx crassifasciella Ragonot. 
836. Nephopteryx gilvibasella Hulst. 


834. reductella 


835. crassifasciella 836. gilvibasella 


508 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 837—-842.— FEMALE GENITALIA. 


837. Actriz dissimulairix Heinrich. 

838. Actriz nyssaecolella (Dyar). 

839. Homoeographa lanceolella Ragonot, para- 
type (in BM, “Peru, Coll. Walker, 
84-72”). 

840. Etiella zinckenella (Treitschke). 

841. Tulsa infinitella (Dyar), type. 

842. Tulsa umbripennis (Hulst), specimen 
from Chimney Gulch, Colo. 


841. infinitella 


PI 
g 
z 
= 
@ 


840. 3inckenella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 509 


Ficures 843-848.— FEMALE GENITALIA, 


843. Telethusia ovalis (Packard). 

844. Phobus brucet (Hulst). 

845. Phobus curvatellus (Ragonot). 

846. Phobus incertus Heinrich, new species. 
847. Stylopalpia scobiella (Grote). 

848. Stylopalpia lunigerella Hampson. 


843. ovalis 


curvatellus 


84.7. scobiella 


UNITED STATES NATIONAL ‘MUSEUM BULLETIN 207 


510 


xe 


Figures 849-852.—FEMALE GENITALIA. 


849, Pyla fasciolalis (Hulst), with ovipositor 
omitted; 849a, female genitalia of a 
variety from Gunnison County, Colo. 

850, Pyla impostor Heinrich, new species. 

851. Pyla viridisuffusella Barnes and McDun- 
nough. 

852. Pyla fusca (Haworth). 


mr sae 


ROOT OAV ANE Cv nyse ted sem uta pan IVETE 


849a. fasciolalis 


851. viridisuffusella 


852 fusca 


AMERICAN MOTHS OF THE > SUBFAMILY PHYCITINAE 511 


Figures 853-856.—FBMALE GENITALIA. 


853. Pyla aenigmatica Heinrich, new species. 
854. Pyla hypochalciella (Ragonot). 

855. Pyla aequivoca Heinrich, new species. 
856. Pyla insinuatriz Heinrich, new species, 


MOLINA BEA NE RMweserets etry, 


a 


~Stam 


econ 


— 


855. aequivoca 856. insinuatrix 


512 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 857—860.—FEMALE GENITALIA. 


857. Pyla scintillans (Grote), figured from 
type of its synonym P. feella Dyar; 
857a, female genitalia of a specimen 
from Tuolumne Meadows, Calif. 

858. Pyla sylphiella Dyar. 

859. Pyla rainierella Dyar. 

860. Pyla hanhamella Dyar. 


ean e7es 


ESenimenn 


858.sylphiella 


Yi S; y 
Ke By A 
= WA NM NZ 
Hee ie Jt 
PS scintillans iS; 
es : Hates 
yg z 
bog 4 5 
a-i \ 2d _ 857a, 


bey 


cent 


bell 


NZ 
ays 


859. rainierella 860. hanhamella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 513 


Ficures 861-864.—FEMALE GENITALIA. 


861. Pyla fasciella Barnes and McDunnough. 

862. Pyla aeneoviridella Ragonot. 

863. Pyla metalicella Hulst; 863a, specimen 
from Silverton, Colo. 

864. Pyla aeneella Hulst, type. 


861. fascicella 


< 


f 


PREERTO Cocet aE 


at seseue LOMAATEA NES OI RPC COMATOSE 


8630. 863. metalicella 864.aeneella 


300328—56——34 


UNITED: STATES NATIONAL MUSEUM BULLETIN ‘207 


514) 


Fieures 865—867.— FEMALE GENITALIA.’ 


865. Dioryctria abietella (Denis and Schiffer- 
miiller) ; 865a, part of female genitalia 
of a specimen showing variation in the 
suture of the sclerotization of ductus 


bursae. 
866. Dioryctria sysstratiotes Dyar. 
867. Dioryctria reniculella Grote. 


ALORA NAOII OS BLAS) pn ERNEL 


Be otiey 


f 


866. sysstratiotes 


1 
/ 


865. abictella 867. reniculella 


868. ponderosae 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 552 


nemoniaa 


suits 


862 erythropasa 


870. Dioryctria 


Sp. 


871. majorella 


Ficures 868-871.—FEMALE GENITALIA: 


868. Dioryctria ponderosae Dyar, paratype 
from California. 

869. Dioryctria erythropasa (Dyar), paratype: 

870. Dioryctria sp., a probable hybrid of au- 
ranticella and erythropasa. 

871. Dioryctria majorella Dyar, type. 


516 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 872—874.—F2EMALE GENITALIA. 


872.. Dioryctria disclusa Heinrich, paratype 
from New Jersey. 

873. Dioryctria auranticella (Grote). 

874. Dioryctria horneana (Dyar), type. 


| 873. auranticella 


872. disclusa 874. horneana 


517 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


Figures 875-878.—FEMALE GENITALIA. 


ir 
o 
s 
Q 
ive) 
_ 
S 
_ 
o 
- 
S} 
iS) 
'- 
i= 
3 
= 
n 
*) 
RS 
= 
= 
3 
= 
~~ 
= 
o 
> 
i] 
‘= 
Q 
1S 
iw 
eo 


® 
2, 
lal 
~ 
zg 
3 
a 
3 
cal 
Q 
~— 
38 
= 
:3 
2s 
a 
g& 
a3 
ans) 
oe 
Ace 
23s 
ge 
n 
dS 
Q 
3 
i= 
-) 


~~ 
° 
q 
fs) 
&p 
a 
[a= 
2s 
xO 
aay 
as 
9 &§ 
Ss 
i = 
oe. 
Ss 
WE 
gs 
o2 
Q 
nN 
b 
re) 


878. Dioryctria zimmermani (Grote), figured 


from a large western (Ariz.) example 


D. delectella (Hulst). 


of its synonym, 


875. 


3immermani 


cambiicola 


878. zimmermani 


518 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 879-882.— FEMALE GENITALIA. 


879. Dioryciria amaiella (Hulst), typical Flor- 
ida specimen. 

880. Dioryctria albovittella (Hiilst). 

881. Dioryctria baumhoferi Heinrich, new spe- 
cies. 

882. Dioryctria subtracta Heinrich, new species. 


882. subtracta 


880. albovittella 


881. baumhoferi 


879. amatella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 519 


Figures 883-885.—FEMALE GENITALIA. 


883. Dioryctria clarioralis (Walker), typical 
Florida specimen. 

884. Epischnia prodromella (Hiibner). 

885. Laodamia faecella (Zeller). 


a 
Me dy 
TAG 


% 
EES 


peice 1 tea 


884. prodromella 


883. clarioralis 


885. faecella 


520 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 886—890.—FEMALE GENITALIA. 


886. Sarata delia Heinrich, new species. 

887. Sarata kappa Heinrich, new species, type. 
888. Sarata alpha Heinrich, new species, type. 
889. Sarata beta Heinrich, new species, type. 
890. Sarata gamma Heinrich, new species. 


\ 


Sa (RE 


886. delta 


wee 
shee, 


BRIS 


892. epsilon 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 521 


rhoiella 


ots 


894.1 


ota 


Figures 891-895.—FEMALE GENITALIA. 


891. Sarata phi Heinrich, new species, type. 

892. Sarata epsilon Heinrich, new species, 
type. 

893. Sarata perfuscalis (Hulst), type. 

894. Sarata iota Heinrich, new species. 

895. Philodema rhoiella (Dyar), paratype from 
type locality. 


522 UNITED: STATES NATIONAL MUSEUM BULLETIN 207 


Ficures 896—900.—FEMALE GENITALIA. 


896. Lipographis fenestrella (Packard). 

897. Lipographis umbrella (Dyar). 

898. Lipographis truncatella (Wright), speci- 
men from San Diego, Calif. 

899. Tota galdinella (Schaus), type. 

900. Oryctometopia fossulatella Ragonot. 


896. \2v 


fenestrella galdinella 


898. 
truncatella 


900.fossulatella 


. umbrella 


903. lithosella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 523 


904.petrella 


902 


rubedinella 


905. 


Ficures 901-905.—FEMALE GENITALIA. 


901. Adelphia ochripunctella (Dyar), showing 
to the side of bursa a single spine of 
the signum spine cluster, greatly en- 
larged. 

902. Ufa rubedinella (Zeller). 

903. Ufa lithosella (Ragonot). 

904, Adelphia petrella (Zeller). 

905. Ufa senta Heinrich, new species, 


524 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 906—910.— FEMALE GENITALIA, 


906. Elasmopalpus lignosellus (Zeller). 

907. Acroncosa albiflavella Barnes and Mc- 
Dunnough. 

908. Passadena flavidorsella (Ragonot). 

909. Ulophora guarinella (Zeller), specimen 
from Cuba. 

910. [Myelozs] famula Zeller. 


ry 


TUNG 
NRA 30720) 
NCA 
ty fi 


Ke 


908. flavidorsella 909.guarinella _ 910. famula 


912.feriella 


AMERICAN MOTHS OF THE 


913. extrincica 


914. idiotes 


SUBFAMILY PHYCITINAE 525 


Figures 911-914.—FeMALE GENITALIA, 


911. [Nephopteryx] fuscifrontella Zeller, speci- 
men from Colombia, with dorsal view 
of ductus bursae shown in small sup- 
plemental figure. 

912. Tacoma feriella Hulst. 

913. Chorrera extrincica (Dyar). 

914. Chorrera idiotes Dyar. 


526 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 915-919. Mat AND FEMALE GENI- 
TALIA. 


915. [Myelois] grosstpunctella Ragonot, male 
genitalia, with aedeagus omitted; 915a, 
aedeagus and anellus; 915b, gnathos; 
915c, transtilla. 

916. [Myelozs] grossipunctella Ragonot, female 
genitalia. 

917. [Elasmopalpus] corrientellus Ragonot, 
type, female genitalia. 

918. [Salebria] nigricans Hulst, type, female 
genitalia. 

919. [Hypochalcia] cervinistrigalis Walker, 
type, female genitalia. 


918. nigricans 


917. corrientellus 919. cervinistrigalis 


mysiella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 527 


parvulella 


aureofasciella 


923. 
dryadella 


922. 
| ochrella 


Figures 920-925.— FEMALE GENITALIA. 


920. 
921. 
922. 
923. 


924, 
925. 


Eumysia mysiella (Dyar), paratype from 
type locality. 

Divitiaca parvulella Barnes and McDun- 
nough. 

Divitiaca ochrella Barnes and McDun- 
nough. 

Ocala dryadella Hulst. 

Macrorrhinia aureofasciella Ragonot. 

Protasia mirabilicornella (Dyar). 


528 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 926—931.—FEMALE GENITALIA. 


926. Valdivia lativittella (Ragonot), from 
Brownsville, Tex. 

927. Valdivia lativittella (Ragonot), from 
Pinal Mts., Ariz. 

928. Heterographis samaritanella (Zeller), 
bursa. 

929. Heterographis morrisonella Ragonot; 
929a, bursa of another example show- 
ing extent of variation in its spining. 

930. Staudingeria albtpenella (Hulst). 

931. Hulstia undulatella (Clemens). 


morrisonella 


926, 
lativittella 


921. 
lativitella 


931. undulatella 


930. albipenella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 529 


cundajensis 


semidiscella 


934. banhoti ulmiarrosorella 


Figures 932-936.—FEMALE GENITALIA, 


932. Cabotia cundajensis (Zeller). 

933. Cabotia rhythmatica Dyar, paratype. 

934. Cabotia bonhoti (Hampson), specimen 
from Nassau, Bahamas. 

935. Cabotia semidiscella Ragonot. 

936. Canarsia ulmiarrosorella (Clemens). 


530 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 937—941.—FEMALE GENITALIA, 


937. Honora mellinella Grote. 

938. Honora dotella Dyar, specimen from San 
Diego, Calif. 

939. Honora montinaiatella (Hulst), type. 

940. Honora perdubiella (Dyar), paratype 
from type locality, showing to side of 
bursa bases of some spines of signum 
patch, greatly enlarged. 

941. Honora subsciurella Ragonot, showing to 
each side of bursa examples of spines, 
greatly enlarged. 


937. 
mellinella 


subsciurella 


montinatatella 


anticella 


ignidorsella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 531 


rubripurpurco 


947. 
Fuliginosus 


Figures 942-947.—FEMALE GENITALIA. 


942. Oncolabis anticella Zeller, specimen from 
French Guiana; 942a, from Guatemala 
(these two figures showing extremes of 
variation). 

943. Hurythmidia ignidorsella (Ragonot). 

944, Oedothmia endopyrella Hampson, figured 
from a sketch by J. F. G. Clarke, of 
the type of its synonym Synothmia 
bahamasella Hampson. 

945. Eurythmasis ignifatua Dyar, paratype 
from type locality. 

946. Stylobasis rubripurpurea Hampson, speci- 
men from Juan Vifias, Costa Rica. 

947. Honorinus fuliginosus Heinrich: new 
species, lateral view; 947a, ventral 
view of genital opening. 


532 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 948-951.—FEMALE GENITALIA. 


948, 949. Wunderia neaeriatella Grossbeck, two 
examples, showing extent of variation. 

950. Cacozophera venosa Dyar, type. 

951. [Honora] dulciella Hulst, an unplaced 
species, type, showing to the side of Ss 
bursa two projections of a signum, 
greatly enlarged. 


94.8. 


neacriatella 


949, 
neaeriatella 


950. venosa 951. dulciella 


semicana, 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 533 


nymphaeella 


Fiaures 952-954.—FEMALE GENITALIA. 


952. Psorosina hammondi (Riley). 
953. Patriciola semicana Heinrich, new species. 
954. Palatka nymphaeella (Hulst). 


534 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 955-959.— FEMALE GENITALIA. 


955, 956. Cassiana malacella (Dyar), two 
specimens from Puerto Rico, showing 
variations in signum. 

957. Aptunga imperfecta (Dyar), type. 

958. Aptunga macropasa (Dyar), type. 

959. Anderida sonorella (Ragonot), type. 


958. 


macropasa 


957. import a 
rBeRneee a 1 vosg. aeieeelln 959. sonorella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 535 


Ficures 960-965.—FEMALE GENITALIA. 


960. Mescinia parvula (Zeller), with samples 
of the spines of signum shown to the 
side of bursa, greatly enlarged. 

961. Mescinia commatella (Zeller), type, bursa 
and part of ductus bursae. 

962. Mescinia bacerella Dyar, specimen from 
Sierra Maestra, Cuba., showing beside 
bursa samples of spines of signum, 
greatly enlarged. 

963. Mescinia estrella Barnes and McDun- 
nough, with samples of signum spines, 
greatly enlarged, shown to the side of 
bursa. 

964. Mescinia peruella Schaus, with enlarge- 
ment of some signum spines to the side 
of bursa; 964a, bursa of another speci- 
men showing variation in spining of 
signum. 

965. Mescinia triloses Dyar, with enlarged 
samples of signum spines shown to 
the side of bursa. 


peruella 


bacerella 


965. triloses 


963. estrella 


536 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 966—-974.—FEMALE GENITALIA. 


966. Mescinia berosa Dyar, type, with en- 
larged samples of signum spines shown 
to the side of bursa. 

967. Mescinia discella Hampson. 

968. Mescinia indecora Dyar, type, signum 
greatly enlarged. 

969. Nonia exiguella (Ragonot), specimen 
from Panamé. 

970. Phestinia costella Hampson; 970a, sig- 
num of Puerto Rican example, greatly 
enlarged; 970b, enlarged signum of 
type. 

971. Comotia sp., specimen from Puerto Rico. 

972. Bema neuricella (Zeller); 972a-c, various 
modifications of the signum, greatly 
enlarged; 972d, signa, greatly enlarged, 
of B. myja Dyar, a synonym of neuri- 
cella. 

973. Bema fifaca (Dyar), type, signum, 
greatly enlarged. 

974. Bema ydda (Dyar), type, signum, greatly 
enlarged. 


exiquella 


costella 


discella 


fifaca 


9724. 


Comotia sp. 


972. 


neuricella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 537 


oslarellum 


980.emendator 


sinuellum 


electellum 


982.inornatellum 


eB 


975a. opalescellum 


% 


976a.uncanale 


983. 


976.stypticellum albescentellum 


300329—55——35 


Ficures 975-983.— FEMALE GENITALIA. 


975. Homoeosoma electellum (Hulst); 975a, 
cephalic end of bursa and signum of 
H. opalescellum (Hulst), a synonym of 
electellum. 

976. Homoeosoma stypticellum Grote, bursa; 
976a, signum of type of H. uncanale 
Hulst, a synonym of stypticellum. 

977. Homoeosoma striatellum Dyar, bursa; 
977a, signum from another specimen, 
showing amount of variation. 

978. Homoeosoma sinuellum (Fabricius), bursa. 

979. Homoeosoma oslarellum Dyar, paratype, 
bursa. 

980. Homoeosoma illuviellum emendator Hein- 
rich, new race, paratype from Rich- 
field, Utah, bursa. 

981. Homoeosoma impressale Hulst, bursa. 

982. Homoeosoma inornatellum (Hulst), type, 
bursa. 

983. Homoeosoma aalbescentellum Ragonot, 
type, bursa. 


538 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 984—990.—FEMALE GENITALIA. 


984. 


985. 


986. 


987. 


988. 


989. 
990. 


Homoeosoma imitator Heinrich, new 
species. 

Homoeosoma deceptorium Heinrich, new 
species, bursa. 

Homoeosoma discrebile Heinrich, new 
species, bursa. 

Homoeosoma peregrinum Heinrich, new 
species. 

Homoeosoma assitum Heinrich, new spe- 
cies. 

Homoeosoma oconequensis (Dyar), type. 

Homoeosoma ditaeniatellum Ragonot, 


type. 


assitum 


988. 


peregrinum 


990. 


deceptorium oconequensis ditaeniatellum 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 539 


992. 


994..mucidella 


991. 
vepallidum : 
993. magellanella 


Fiaures 991-994.— FEMALE GENITALIA. 


991. Homoeosoma vepallidum Heinrich, new 
species, paratype. 

992. Homoeosoma vepallidum Heinrich, new 
species, specimen from Sierra de 
Cérdoba, Argentina. 

993. Patagonia magellanella (Ragonot), type. 

994, Rotruda mucidella mucidella (Ragonot); 
994a, samples of spines of signum, 
greatly enlarged and shown in two 
projections. 


540 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 995-1001.— FEMALE GENITALIA. 


995. Unadilla erronella (Zeller); 995a en- 
largements of individual signa. 

996. Unadilla maturella (Zeller), specimen 
from Guatemala. 

997. Laetilia coccidivora (Comstock); 997a, 
dorsal view of eighth-segment collar; 
997b, enlargement of signum. 

998. Laetilia coccidivora cardint Dyar, type; 
998a, dorsal view of eighth-segment 
collar. 

999. Laetilia coccidivora quadricolorella 
(Dyar); 999a, dorsal view of eighth- 
segment collar. 

1000. Laetilia obscura Dyar; 1000a, dorsal 
view of eighth-segment collar. 

1001. Laetilia portoricensis Dyar; 1001a, dor- 
sal view of eighth-segment collar. 


997. coccidivora 


quadricolorella * 1000. obscura 
1001. portoricensis 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 


10025 


zamacrella 
1004. 


1008. 
glabrella 


eremiella 


1006a, 


1005: fiskella 


1007. haywardi 


541 


Fiaures 1002-1008.—FEMALE GENITALIA. 


1002. 


1003. 


1004, 
1005. 


1006. 


1007. 


1008. 


Laetilia melanostathma (Meyrick); 
1002a, dorsal view of eighth-segment 
collar and apical portion of ductus 
bursae. 

Laetilia myersella Dyar, with enlarge- 
ment of signum shown to the side of 
bursa; 1003a, dorsal view of eighth- 
segment collar. 

Laetilia zamacrella Dyar. 

Laetilia fiskella Dyar, with enlargement 
of signum shown to the side of bursa. 

Baphala basimaculatella (Ragonot), fig- 
ured from its synonym Laetilia ere- 
miella Dyar; 1006a, enlargement of 
signum, 

Baphala haywardi Heinrich, new spe- 
cies, with enlargement of signum 
shown to the side of bursa. 

Baphala glabrella (Dyar), type, signum, 
greatly enlarged. 


542 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 1009-1015.— FEMALE GENITALIA. \ ty 


1009a, olivacea 


1009. Baphala goyensis (Ragonot), signa, goyensis 
1010a. 


showing extremes of variation. 

1010. Baphala goyensis olivacea Heinrich, new 
race; 1010a—d, various modifications 
of the signum in this race. 

1011. Baphala homoeosomella (Zeller): 1011— 
1011a, Varieties of signa of female 
genitalia in typical examples; 1011b- 
1011f, varieties of signa in 24 
synonym, sazssetiae (Dyar); 1011¢, 
signum of another synonym, bodkinz 
(Dyar); 1011h, apical part of ductus 
bursae, collar, and ovipositor of 
another synonym, rusto (Dyar). 

1012. Rhagea packardella (Ragonot). 

1013. Rhagea stigmella (Dyar). 

1014. Rhagea maculicula (Dyar), a synonym of 
R. stigmella (Dyar). 

1015. Zophodia convolutella (Hiibner). 


10llg. 
bodkni 


\ 
1010. olivacea 


convolutella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 543 


Figures 1016—1019.— FEMALE GENITALIA. 


1016. Melitara prodenialis Walker; 1016a, dor- 
sal view of eighth-segment collar. 

1017. Melitara dentata (Grote). 

1018. Olycella junctolineella (Hulst); 1018a, 
dorsal view of eighth-segment collar. 

1019. Olycella subumbrella (Dyar), paratype 
from type locality, collar of eighth 
abdominal segment, dorsal view. 

(Explanation of symbols applied 

to female genitalia: Bc, bursa copu- 
latrix; Clr, collar of eighth abdominal 
segment; Db, ductus bursae; dp, dor- 
sal plates behind genital opening; Ds, 
ductus seminalis; Go, genital open- 
ing; Sm, signum.) 


1017. dentata 


2 P1015 subumbrelia 
1018. junctolineella 


544 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 1020—-1023.— FEMALE GENITALIA. 


1020. Olyca phryganoides Walker; 1020a, dor- 
sal view of eighth-segment collar. 

1021. Alberada parabates (Dyar); 1021a, dor- 
sal view of eighth-segment collar. 

1022. Alberada bidentella (Dyar). 

1023. Alberada holochlora (Dyar). 


102].parabates 


1022. bidentella 


1023. holochlora 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 545 


Figures 1024-1027.— FEMALE GENITALIA. 


1024. Cactoblastis cactorum (Berg) ; 1024a, dor- 
sal view of eighth-segment collar. 

1025. Cactoblastis doddi Heinrich; 1025a, dor- 
sal view of eighth-segment collar. 

1026. Cactoblastis bucyrus Dyar; 1026a, dorsal 
view of eighth-segment collar. 

1027. Cahela ponderosella Barnes and McDun- 
nough, paratype from type locality; 
1027a, dorsal view of eighth-segment 
collar. 


1024. cactorum 


1026.bucyrus 1027 ponderosella 


300329—56——-86 


546 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 1028—1032.— FEMALE GENITALIA. 


1028. Rumatha glaucatella (Hulst), with 
eighth-segment collar and ovipositor 
omitted and with signum shown, 
much enlarged, to the side of bursa. 

1029. Rumatha polingella (Dyar), with eighth- 
segment collar and ovipositor omitted. 

1030. Rumatha bihinda (Dyar). 

1031. Eremberga leuconips (Dyar); 1031a, 
dorsal view of eighth-segment collar. 

1032. Yosemitia longipennella (Hulst), with 
eighth-segment collar and ovipositor 
omitted and with signum, much en- 
larged, shown to the side of bursa. 


1028. glaucatella 


1029. polingella 


1030. bihinda 


1032. longipennella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 547 


Fiaures 1033-1037.—F MALE GENITALIA. 


1033. Yosemitia graciella (Hulst). 

1034. Tucumania tapiacola Dyar. 

1035. Tucumania porrecia Dyar, with eighth- 
segment collar and ovipositor omitted. 

1036. Ozamia immorella (Dyar). 

1037. Ozamia punicans Heinrich. 


229099... 
Ps $ 
APS 


FITTS 

3 392 3 Sy 
3407 Ae 
282 +; 


Eeey 


ELA 

35328957 

beers 
2 


1033.graciella 1034. tapiacola 1037. punicans 


548 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fieures 1038-1043.—FEMALE GENITALIA. 


1038. 


1039. 


1040. 


1041. 


1042. 


1043. 


Ozamia stigmaferella (Dyar), type; 
1038a, dorsal view of eighth-segment 
collar. 

Ozamia thalassophila Dyar, type; 1039a, 
dorsal view of eighth-segment collar. 

Ozamia fuscomaculella clarefacta Dyar; 
1040a, dorsal view of eighth-segment 
collar. 

Ozamia hemilutella Dyar; 1041a, dorsal 
view of eighth-segment collar. 

Ozamia lucidalis (Walker); 1042a, dor- 
sal view of eighth-segment collar. 

Ozamia fuscomaculella (Wright). 


1040, 


1038. stigmaferella clarafacta. fs: 


‘1041a. 


1041. hemilutella 1042. lucidalis 


1043. fuscomaculella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 549 


\ analamprella 


substituta 


1047a. 


fernaldialis 


1044. leithella 1047. huanucensis 


Fiaures 1044-1048.— FEMALE GENITALIA. 


1044, 


1045. 
1046. 


1047. 


1048. 


Amalafrida leithella (Dyar) ; 1044a, dor- 
sal view of eighth-segment collar. 

Salambona analamprella (Dyar). 

Sigelgaita chilensis Heinrich; 1046a, 
dorsal view of eighth-segment collar, 

Sigelgaita huanucensis Heinrich, type; 
1047a, dorsal view of eighth-segment 
collar, 

Nanaia substituta Heinrich; 1048a, dor- 
sal view of eighth-segment collar. 


550 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 1049-1052.— FEMALE GENITALIA. 


1049. Cactobrosis fernaldialis (Hulst); 1049a, 
dorsal view of eighth-segment collar. 

1050. Cactobrosis longipennella (Hampson), 
specimen from Oaxaca, México; 
1050a, dorsal view of eighth-segment 
collar. 

1051. Cactobrosis maculifera Dyar, eighth-seg- 
ment collar and apical portion of duc- 
tus bursae. 

1052. Cactobrosis insignatella Dyar, type; 
1052a, dorsal view of eighth-segment 
collar. 


SI 


maculifera 


1052. insignatella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 551 


Figures 1053-1057.— FEMALE GENITALIA. 


1053. Cactobrosis strigalis Barnes and Mc- 
Dunnough. 

1054. Illatila gurbyris Dyar, with enlarge- 
ment of signum shown to the side of 
bursa. 

1055. Lascelina canens Heinrich, new species. 

1056. Metephestia simplicula (Zeller). 

1057. Selga arizonella (Hulst); 1057a, signum, 
greatly enlarged. 


1054. gurbyris 


1053. 


057. 
arizonella 


1056. 
simplicula 


1055. canens 


552 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 1058—1062.— FEMALE GENITALIA. 


1058. Entmemacornis proselytes Dyar. 

1059. Anthropteryx irichampa Dyar, type. 

1060. Moodnopsis portoricensis Heinrich, new 
species. 

1061. Moodnopsis inornatella (Ragonot). 

1062. Moodnopsis decipiens Dyar. 


irichampa 


1059. 


1061. 


inornatella 


1062. decipiens 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 553 


Figures 1063-1067.—F eMALE GENITALIA. 


1063. Huzophera cinerosella (Zeller). 

1064, Euzophera semifuneralis (Walker). 

1065. Huzophera ostricolorella Hulst. 

1066. Huzophera nigricantella Ragonot. 

1067. Prosoeuzophera impletella (Zeller), speci- 
men from Puerto Rico (in Cornell 
Univ.). 


1064. 


semifuneralis 


1066. 
1065. ostricolorella nigricantella 1067, impletella 


554 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 1068-1071.— FEMALE GENITALIA. 


1068. Vezina parasitaria Heinrich, new spe- 
cies; 1068a, lateral view of part of 
genitalia showing sclerotizations at or 
near genital opening. 

1069. Eulogia ochrifrontella (Zeller); 1069a, 
enlargement of signum. 

1070. Farnobia quadripuncta (Zeller); 1070a, 
enlargement of signum. 

1071. Edulica compedella (Zeller). 


1068a. 


ochrifrontella: 


1068. parasitaria 


1071. 


compedella 


1070. ppadeipancta! 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 555 


Figures 1072-1078.— FEMALE GENITALIA. 


1072, 1073. Ephestiodes gilvescentella Ragonot, 
two examples, showing extent of vari- 
ation. 

1074. Ephestiodes erythrella Ragonot; 1074a-b, 
outlines of two shields behind genital 
opening, showing extent of variation. 

1075. Ephestiodes mignonella Dyar, type. 

1076. Ephestiodes indentella Dyar; 1076a, dor- 
sal view of eighth-segment collar. 

1077. Ephestiodes erasa Heinrich, new species; 
10778, dorsal view of eighth-segment 
collar. 

1078. Ephestiodes productella Ragonot, type 
(in Paris Mus.). 


1072. 
gilvescentella 


1073. 
_ gilvescentella 


1074 
erythrella. 


1077a. 


MUTT eccae 1078. productella 


506 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Friaurss 1079-1085.— FEMALE GENITALIA. 


1079. 
1080. 


1081. 


1082. 


1083. 
1084. 


1085. 


Ephestiodes noniella Dyar. 

Ephestiodes plorella Dyar; 1080a, dorsal 
view of eighth-segment collar. 

Ephestiodes lucidibasella Ragonot, para- 
type (nm BM), apical (sclerotized) 
part of ductus bursae. 

Ephestiodes stictella (Hampson); 1082a, 
dorsal portion of eighth-segment col- 
lar; 1082b, apical portion of ductus 
bursae, plate behind genital opening, 
and apophyses of eighth-segment col- 
lar. 

Micromescinia pygmaea Dyar. 

Azaera muciella Schaus; 1084a, dorsal 
view of eighth-segment collar. 

Azaera nodoses (Dyar), type; 1085a, 
dorsal view of eighth-segment collar. 


1080. plorella 


Pygmaea 


lucidibasella 


1084. muciella 


1085.nodoses 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 557 


1088. 


edmandsae serratilineella 


1089. inanimella. 


1087a.bisinuella 


1090.pinei 


YZ 
Hy 


1087. bisinuella 


Figures 1086-1090.— FEMALE GENITALIA. 


1086. Moodna ostrinella (Clemens); 1086a, 
lateral view of ovipositor and inter- 
segmental membrane with lobe, be- 
tween ovipositor and eighth-segment 
collar; 1086b, dorsal view of eighth- 
segment collar. 

1087. Moodna bisinuella Hampson; 1087a, 
lateral view of caudal portion of geni- 
talia, showing genital opening, eighth- 
segment collar, intersegmental lobe 
and ovipositor; 1087b, dorsal view of 
eighth-segment collar. 

1088. Vitula edmandsae serratilineella Rago- 
not. Female genitalia. 

1089. Vitula inanimella (Dyar), paratype from 
Orizaba, México; 1089a, female geni- 
talia figured from type of its synonym 
Euzophera ticitoa Dyar. 

1090. Vitula pinei Heinrich, new species. 


558 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 1091-1097.—FEMALE GENITALIA. 


1091. Vztula laura (Dyar), type. 

1092. Manhaita biviella (Zeller). 

1093. Manhatta setonella (McDunnough), para- 
type (in USNM). 

1094. Manhatia broweri Heinrich, new species, 
paratype from type locality. 

1095. [Eucampyla] putidella Schaus, type. 

1096. Moodnella paula Heinrich, new species. 

1097. Verina supplicella (Dyar), type. 


1092. biviella 


1091. laura: 


1097. 


1093. setonella 1094. broweri supplicella 


1098. 
pachytaeniella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 559 


1099. trinitalis 


Fiaures 1098-1100.—FrMALE GENITALIA. 


1098. Volatica pachytaeniella (Ragonot). 
1099. Volatica trinitatis Heinrich, new species. 
1100. Vagobanta divergens (Butler). 


l00.divergens 


560 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Fiaures 1101—1105.— FEMALE GENITALIA. 


1101. Caudellia colorella (Dyar), type. 

1102. Caudellia nigrella (Hulst) ; 1102a, dorsal 
view of eighth-segment collar. 

1108. Caudellia albovittella Dyar. 

1104. Caudellia declivella (Zeller), type. 

1105. Soszpatra rileyella (Ragonot) 


1102a. \ 
nigrella | 


colorella ao 
MOS a 
albovittella 


et We 


ly 


\ 2 
loz. 7 N04. 


rileyella nigrella declivella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 561 


Ficures 1106-1114.—FEMALB GENITALIA. 


1106. Sosipatra anthophila (Dyar). 

1107. Sosipatra thurberiae (Dyar). 

1108. Sosipatra majorella (Dyar). 

1109. Sosipatra divergens (Dyar), type. 

1110. Varneria postremella Dyar, type. 

1111. Varneria nannodes Dyar, type. 

1112. Varneria atrifasciella Barnes and Mc- 
Dunnough. 

1113. Varneria dubia Heinrich, new species. 

1114. Microphestia animacula Dyar, type. 


J109. divergens 
H06.onthophila —_ 107. thurberiae 1108. majorella 


14. 


animacula 


u10. ; : lil2. 
postremella _lll.nannodes atrifasciella 


113, dubia 


562 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Figures 1115—-1120.— FEMALE GENITALIA. 


1115. 
1116. 
1117. 
1118. 
1119. 
1120. 


Ribua innoxia Heinrich. 

Ribua patriciella (Dyar), type. 

Ribua contigua Heinrich, new species. 
Plodia interpunctella (Hiibner). 

Plodia dolorosa Dyar. 

Bethulia championella Ragonot, type. 


118. interpunctella 


1119. dolorosa 


1120.championella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 563 


Figures 1121-1125.—FemMALE GENITALIA. 


1121. Ephestia cautella (Walker). 

1122. Ephestia elutella (Hiibner). 

1123. Ephestia figulilella Gregson. 

1124. Anagasta kihniella (Zeller) ; 1124a, some 
variations in signa, enlarged. 

1125. Nicetiodes apianella Schaus, paratype 
from type locality. 


121. cautella 


1125. apianella 
1124. kihniella 


564 UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Ficgures 1126-1129.— FEMALE GENITALIA. 


1126. [Huzophera] came Dyar, type; 1126a, 
dorsal view of eighth-segment collar. 

1127. [Moodna] formulella Schaus, type. 

1128. [Euzophera] postflavida Dyar; 1128a, en- 
largement of signum. 

1129. [Euzophera] rinmea Dyar, type; 1129a, 
dorsal view of eighth-segment collar. 


1127. formulella 


1126. came 


29. 


postflavida rinmea 


1136, conops 


1137. titillella 


AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 565 


1133.fumella 


1138. myronella 


1134. quantulella 


1135. pohellelle 


Figures 1130-1138.—FrMALE GENITALIA. 


1130. 


1131. 


1132. 
1133. 
1134. 


1135. 
1136. 


1137. 
1138. 


Eurythmia hospitella (Zeller), figured 
from type; 1130a, bursa of a female 
from Texas, showing variation in 
number of signa. 

Eurythmia hospitella yavapaella Dyar, 
bursa of a female from Glenwood 
Springs, Colo.; 1131a, bursa of a fe- 
male from San Diego, Calif. 

Eurythmia angulella Ely, bursa. 

Eurythmia fumella Ely, type, bursa. 

Erelieva quantulella (Hulst), specimen 
from Blanco County, Tex., showing 
to the side of bursa a couple of signa, 
greatly enlarged. 

Erelieva parvulella (Ely), bursa. 

Rabiria conops (Dyar), specimen from 
type locality. 

Microphycita titillella Dyar. 

Cabnia myronella Dyar. 


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Index to the genera and species of Phycitinae 


(Page numbers of principal entries in boldface. 


Synonyms in italics. 


For species unknown, misplaced, or unrecognized, see page 580.) 


abditiva Heinrich, n. sp., Meroptera, 123 | Acrobasis Zeller—Continued 


aberrans Heinrich, n. sp., Protomoerbes, 
49 
abietella (Denis & Schiffermiiller), Di- 
oryctria, 149, 150, 151, 152, 156, 233 

abietivorella (Grote), Dioryctria, 150 
abitus Heinrich, Sematoneura, n. sp., 27 
acmaeopterum Ragonot, Homoeosoma, 225 
Acrobasis Zeller, 1, 2, 10, 11, 25, 26, 37 

albocapitella Hulst, 16 

alnella McDunnough, 23 

amplexella Ragonot, 13, 118 

angusella Dyar, 16 

angusella Grote, 19, 20 

aurorella Ely, 18 

betulella Hulst, 11, 22, 23 

caryae Dyar (not Grote), 22 

caryae Grote, 17, 18 

caryaevorella (Dyar, not Ragonot), 17 

earyalbella Ely, 15, 16 

caryivorella Ragonot, 22, 24 

cirroferella Hulst, 17, 21 

comacornella (Hulst), 22 

comptella Ragonot, 14, 24 

comptoniella Grossbeck (not Hulst,) 23 

comptoniella Hulst, 23 

coryliella Dyar, 21 

cunulae Dyar & Heinrich, 22 

demotella Grote, 19 

dyarella Ely, 20, 21 

eliella Dyar, 19 

evanescentella Dyar, 17 

exsulella (Zeller), 19 

feltella Dyar, 15, 17 

grossbecki (Barnes & McDunnough), 

13 

hebescella (Dyar, not Hulst), 17 

hebescella Hulst, 21 

indigenella (Zeller), 12, 14 

irrubriella Ely, 20 

juglandis (LeBaron), 15, 16, 17 

kearfottella Dyar, 17 

latifasciella Dyar, 20 

malipennella Dyar, 20 

minimella Ragonot, 11, 15 

myricella Barnes & McDunnough, 23 

nebulella Dyar (not Riley), 16 

nebulella (Grossbeck, not Riley), 13 

nebulella (Riley), 12, 13 

nebulo (Walsh), 12 

nigrosignella Hulst, 15 

normella Dyar, 20 

ostryella Ely, 21 

palliolella Dyar (not Ragonot), 16 

palliolella Ragonot, 15, 16 


peplifera Dyar, 18, 19 
rubrifasciella Packard, 23 
scitulella (Hulst), 14 
secundella Ely, 21 
septentrionella Dyar, 19 
stigmella Dyar, 18 
sylviella Ely, 16, 21 
tricolorella Grote, 13, 14 
tumidella (Zincken), 11, 17 
tumidulella (Ragonot), 24 
vaccinii Riley, 13 
zelatella (Hulst), 12 
zelleri Ragonot, 11 
Acrocaula Hulst, 11, 12 
Acromeseres Dyar, 176 
Acroncosa Barnes & McDunnough, 3, 5, 
174 
albiflavella Barnes & McDunnough, 
174 
albiflavella castrella Barnes & McDun- 
nough, 174 
similella Barnes & McDunnough, 174 
Actrix Heinrich, n. gen., 3, 5, 139 
dissimulatrix Heinrich, n. sp., 140 
nyssaecolella (Dyar), 139, 140 
actualis (Hulst), Catastia, 110, 111 
Adanarsa Heinrich, n. gen., 4, 35 
intransitella (Dyar), 35 
Adelperga Heinrich, n. gen., 182, 187 
cordubensiella (Ragonot), 187, 196 
Adelphia Heinrich, n., gen., 7, 168, 170 
hapsella (Hulst), 169 
ochripunctella (Dyar), 169 
petrella (Zeller), 168, 169, 170 
rubiginella (Walker), 169 
rufinalis (Walker), 169 
ademptandella (Dyar), Salebriaria, 115 
admixta Heinrich, n. sp., Quasisalebria, 
118, 119 
adonea (Felder & Rogenhofer), Crocido- 
mera, 33 
advenella (Zincken), Rhodophaea, 24, 25 
aeneella Hulst, Pyla, 147, 148 
aeneoviridella Ragonot, Pyla, 147, 148 
aenigmatica Heinrich, n. sp., Pyla, 144, 145 
aequivoca Heinrich, n. sp., Pyla, 143, 145 
afflictella (Hulst), Nephopteryx, 131 
affusella (Ragonot) (race of mucidella 
(Ragonot)), Rotruda, 219, 227 
agapella Schaus, Fundella, 60, 61 
aglaeella Ragonot, Euzophera, 273 
ahemora Dyar, Fundella, 59, 62 
ahenella (Zeller), Hypochalcia, 312 
alatella (Hulst), Myelopsis, 41 


Alberada Heinrich, 183, 244, 245 
bidentella (Dyar), 244, 245 
holochlora (Dyar), 245 
parabates (Dyar), 244, 245 


albescentellum Ragonot, Homoeosoma, 
222, 223 

albidiorella (Richards & Thomson), Una- 
dilla, 229 

albiflavella Barnes & McDunnough, 


Acroncosa, 174 
albipenella (Hulst), Staudingeria, 194, 195 
albiplagiatella Hulst (not Packard), Pima, 
102, 104 
albiplagiatella (Packard), Pima, 103, 104 
albiplagiatella (Ragonot, not Packard), 
Pima, 104 
albistrigella Staudinger, Myrlaea, 124 
albocapitella Hulst, Acrobasis, 16 
albocostalialis (Hulst), Pima, 103, 104 
albocostalis (Hulst), Pima, 104 
albocostella Hulst, ‘“Maricopa,’”’ 192, 316 
albosigno Heinrich, n. sp., Diatomocera, 52 
albovittella Dyar, Caudellia, 292 
albovittella (Hulst), Dioryctria, 127, 156 
aliculella (Hulst), Olybria, 113, 114 
almonella (Dyar), Praedonula, 82, 83 
alnella McDunnough, Acrobasis, 23 
alpha Heinrich, n. sp., Megarthria, 88 
alpha Heinrich, n. sp., Sarata, 164 
alternosquamella Ragonot, Dasypyga, 69 
alveolella (Ragonot), Moerbes, 269 
Amalafrida Heinrich, 183, 255, 256 
leithella (Dyar), 256 
amarella Dyar, Ephestia, 302 
amatella (Hulst), Dioryctria, 156 
Ambesa Grote, 7, 108, 110 
laetella Grote, 108 
lallatalis (Hulst), 109 
monodon Dyar, 108 
walsinghami (Ragonot), 108, 109 
walginghami mirabella Dyar, n. sta- 
tus, 109 
amphimetra (Meyrick), Laetilia, 230, 233 
Amphycitopsis Dyar, 78, 80 
amplexella Ragonot, Acrobasis, 13, 118 
Anabasis Heinrich, n. gen., 2, 25 
crassisquamella (Hampson), 26 
ochrodesma (Zeller), 25, 26 
Anadelosemia Dyar, 3, 65, 67, 69, 72, 73 
base Dyar, 68 
condigna Heinrich, n. sp., 69 
dulcella (Hulst), 68 
fifria Dyar, 68 
obstitella (Schaus), 68 
senesciella (Schaus), 67, 68, 69 


567 


568 


Anadelosemia Dyar—Continued 
tecmessella (Schaus), 67 
texanella (Hulst), 68, 69 
Anagasta Heinrich, n. gen., 186, 299 
fuscofasciella (Ragonot), 300 
gitonella (Druce), 300 
kiihniella (Zeller), 291, 299, 300, 302 
sericaria authors (not Scott), 300 
analamprella (Dyar), Salambona, 254 
Ancylostomia Ragonot, 9, 95, 171 
argyrophleps Dyar, 96 
diffissella (Zeller), 95 
euchroma Dyar, 96 
ignobilis (Butler), 95 
sauciella (Zeller), 96 
stercorea (Zeller), 95, 96 
Anderida Heinrich, n. gen., 
213 
placidella (Dyar), 211 
senorella (Ragonot), 211 
sonorella (Ragonot), 211 
Anegcephalesis Dyar, 8, 93 
arctella (Ragonot), 94 
cathaeretes Dyar, 93, 94 
Anerastiinae, 1 
angulella Dyar, Psorosina, 208, 209 
angulella Ely, Eurythmia, 307, 308 
angulinella (Schaus), Hyalospila, 58 
angusella Dyar, Acrobasis, 16 
angusella Grote, Acrobasis, 19, 20 
angustellus Blanchard, Elasmopalpus, 172, 
173 
animaleula Dyar, Microphestia, 294 
animosella (Dyar), Caudellia, 293 
annulatella (Zetterstedt), Polopeustis, 120, 
121 
annuliferella (Dyar), Chararica, 38, 39 
annulosella (Ragonot), Salebriaria, 
117 
anthophila (Dyar), Sosipatra, 295 
anthracellus Ragonot, Elasmopalpus, 173 
anticella Zeller, Oncolabis, 199, 206 
Anypsipyla Dyar, 8, 42 
univitella Dyar, 42 
Anthropteryx Dyar, 208, 313 
apianella Schaus, Nicetiodes, 304, 314 
apocerastes Dyar, Piesmopoda, 78, 80, 
81 
Apomyelois Heinrich, n. gen., 8, 42, 44 
bilineatella (Ragonot), 43 
bistriatella (Hulst), 42, 43 
Aptunga Heinrich, n. gen., 181, 211, 212, 
213 
imperfecta (Dyar), 211 
macropasa (Dyar), 211 
apyrella Dyar, Caudellia, 292 
arctella (Ragonot), Anegcephalesis, 94 
arctiella (Gibson), Polopeustis, 121 


182, 211, 


116, 


argentina Dyar, Fundella, 55, 59, 60, 
61, 62 

argentinensis Heinrich, n. sp., Stylo- 
palpia, 141 


argyrophleps Dyar, Ancylostomia, 96 
aridella (Dyar), Sarata, 161 

arizonella (Hulst), Selga, 265 
arizonella (Walter), Caudellia, 293 


arizonensis Heinrich, n. sp., Pseudo- 
cabima, 55 

assitum Heinrich, n. sp., Homoeosoma, 
225 


asthenosoma (Dyar), Parolyca, 255 
Atheloca Heinrich, n. gen., 7, 81,'83 
bondari Heinrich, n. sp., 77, 82 
filiolella (Hulst), 82 
ptychis Bondar (not Dyar), 82 
ptychis (Dyar), 82 
subrufella (Hulst), 81, 82, 33 
atrella (Hulst), Sarata, 2, 162, 164, 165 
atrifasciella Barnes & McDunnough, Var- 
neria, 305, 306 
atristrigella Ragonot, ‘‘Myelois,’”’ 315 
atrovenosella Ragonot, Sematoneura, 27, 
51 
auranticella (Grote), Dioryctria, 152, 153 
aureofasciella Ragonot, Macrorrhinia, 190 
aureomaculella (Dyar), Valdivia, 192 
aurorella Ely, Acrobasis, 18 
australella (Hulst), Zamagiria, 92, 171 
austriana (Cosens), Dioryctria, 155 
Azaera Schaus, 180, 185, 282 
lophophora (Dyar), 283 
muciella Schaus, 282, 283 
nodoses (Dyar), 283 
Azaera squalidella (Dyar), 282 
azonaxsalis Walker, Davara, 73, 75 


bacerella Dyar, Mescinia, 213, 214 
bahamasella (Hampson), Oedothmia, 205 
bakerella (Dyar), Salebriacus, 114 
Ballovia Dyar, 59 
Bandera Ragonot, 1, 315, 316 
Baphala Heinrich, n. gen., 181, 234, 235 
basimaculatella (Ragonot), 235, 236 
bodkini (Dyar), 236 
eremiella (Dyar), 235 
glabrella (Dyar), 237 
goyensis (Ragonot), 235, 236 
goyensis olivacea Heinrich, n. race, 236 
haywardi Heinrich, n. sp., 236 
homoeosomella (Zeller), 235, 236, 237 
rusto (Dyar), 236 
saissetiae (Dyar), 236 
squalida (Walker), 236, 237 
taboga (Dyar), 236 
base Dyar, Anadelosemia, 68 
basilaris Zeller, Nephopteryx, 129 
basimaculatella (Ragonot), Baphala, 235, 
236 
baumhoferi Heinrich, n. sp., Dioryctria, 157 
bella Hulst, Zophodia, 239 
Bema Dyar, 180, 185, 217 
fifaca (Dyar), 218, 219 
fritilla Dyar, 218 
myja Dyar, 217, 218, 219 
neuricella (Zeller), 218 
ydda (Dyar), 217, 218 
yddiopsis (Dyar), 219 
benjaminella Dyar, Ephestiodes, 279 
berosa Dyar, Mescinia, 214 
Bertelia Barnes & McDunnough, 4, 11, 36 
grisella Barnes & McDunnough, 36, 37 
beta Heinrich, n. sp., Megarthria, 88 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


beta Heinrich, n. sp., Sarata, 164 
Bethulia Ragonot, 186, 296, 298 
championella Ragonot, 296, 297 

betulella Hulst, Acrobasis, 11, 22, 23 

bicolorella (Barnes & McDunnough), Cha- 
rarica, 39 

bidentella (Dyar), Alberada, 244, 245 

bifasciella Hulst, Nephopteryx, 130 

bigrana (Zeller), Hemiptilocera, 30, 31 

bihinda (Dyar), Rumatha, 248, 249 

bilineatella (Ragonot), Apomyelois, 43 

bipunctella (Hampson), Unadilla, 228 

Birinus Heinrich, n.-gen., 4, 11, 36 

russeolus Heinrich, n: sp., 36 

bisinuella Hampson, Moodna, 284 

bisra Dyar, Nephopteryx, 133 

bistriatella (Hulst), Apomyelois, 42, 43 

bistriatella (Hulst), Catastia, 110, 111 

bistriga (Haworth), Cryptoblabes, 10 

biviella (Zeller), Manhatta, 272, 284, 287 

blackmorella Dyar, Pyla, 145 

bodkini (Dyar), Baphala, 236 

boisduvaliella (Guénée), Pima, 102, 103, 
105 

bollaz Dodd (not Zeller), Melitara, 241 

bolliz (Zeller), Melitara, 240 

bondari Heinrich, n. sp., Atheloca, 77, 82 

bonhoti (Hampson), Cabotia, 200, 201 

breviplicitum Heinrich, n. race (race of 
oslarellum Dyar), Homoeosoma, 221 

brevistrigella Ragonot, ‘‘Zophodia,” 314 

broweri Heinrich, n. sp., Manhatta, 288 

brucei (Hulst), Phobus, 110, 138, 139 

brunneella (Dyar), Dioryctria, 158 

bucyrus Dodd (not Dyar), Cactoblastis, 246 

bucyrus Dyar, Cactoblastis, 247 

bumeliella (Barnes & McDunnough), Zama- 
giria, 92 

burdetellus (Schaus), Peadus, 83, 84 

busckella (Dyar), Glyptocera, 101 


Cabima Dyar, 50 

Cabnia Dyar, 310 
myronella Dyar, 310 

Cabotia Ragonot, 180, 200 
bonhoti (Hampson), 200, 201 
cundajensis (Zeller), 201 
impeditella (Zeller), 201 
rhythmatica Dyar, 200, 201 
schini (Berg), 201 
semidiscella Ragonot, 200, 201 

cacabella (Hulst), Pyla, 145 

Cacozophera Dyar, 185, 208 
venosa Dyar, 208 

Cactoblastis Ragonot, 1, 183, 245 
bucyrus Dodd (not Dyar), 246 
bucyrus Dyar, 247 
cactorum (Berg), 245, 246, 247, 257 
doddi Heinrich, 246, 247 
mundelli Heinrich, 247 
ronnai (Bréthes), 245, 246 

Cactobrosis Dyar, 183, 233, 240, 257, 260 
cinerella (Hulst), 260 
elongatella (Hampson), 261 
fernaldialis (Hulst), 260, 261 
gigantella (Ragonot), 260. 


Cactobrosis Dyar—Continued 
insignatella Dyar, 260, 261 
longipennella (Hampson), 260, 261 
maculifera Dyar, 260, 261 
strigalis (Barnes & McDunnough), 
248, 253, 260, 262 
cactorum (Berg), Cactoblastis, 245, 246, 
247, 257 
Cadra Walker, 301 
Cahela Heinrich, 182, 184, 247, 248, 249 
interstitialis (Dyar), 248 
phoenicis (Dyar), 248 
ponderosella (Barnes & McDunnough), 
247, 248, 249 
purgatoria (Dyar), 248 
cahiritella Zeller, Ephestia, 303 
Calamophleps Dyar, 282 
caliginella (Hulst), Rhodophaea, 15, 24, 25 
caliginoidella (Dyar), Rhodophaea, 24 
cambiicola (Dyar), Dioryctria, 150, 155, 
156, 157 
came Dyar, ‘‘Euzophera,’’ 313. 
Campyloplesis Dyar, 269 
Canarsia Hulst, 181, 201 
feliculella Dyar, 202 
fuscatella (Hulst), 202 
gracilella Hulst, 202 
pneumatella (Hulst), 202 
ulmella (Ragonot), 202 
ulmiarrosorella (Clemens), 201, 202 
candidella Hulst, Homoeosoma, 221 
canens Heinrich, n. sp., Lascelina, 264 
canescentelia (Hulst), Passadena, 175 
canicostella Ragonot, Honora, 198 
carabayella Dyar, Pseudodivona, 48, 49 
carbonella (Hulst), Elasmopalpus, 173 
eardini Dyar (race of coccidivora (Com- 
stock), Laetilia, 230, 232 
earicae (Dyar), Davara, 73, 74, 75 
Caristanius Heinrich, n. gen., 5, 97 
decoloralis (Walker), 98 
floridellus (Hulst), 98 
furfurella (Hulst), 98 
guatemalellus (Ragonot), 98 
melanoplaga (Hampson), 97 
metagrammalis (Walker), 98 
pellucidellus (Ragonot), 97, 98 
carneella Hulst, Nephopteryx, 128 
carpasella (Schaus), Fulrada, 72 
caryae Dyar (not Grote), Acrobasis, 22 
caryae Grote, Acrobasis, 17, 18 
caryaevorella (Dyar, not Ragonot), Acro- 
basis, 17 
caryalbella Ely, Acrobasis, 15, 16 
caryivorella Ragonot, Acrobasis, 22 °4 
cassiae (Dyar), Paramyelois, 47 
Cassiana Heinrich, n. gen., 181, 212, 213 
Cassiana malacella (Dyar), 212 
eastrella Barnes & McDunnough (race of 
albiflavella Barnes & McDunnough), 
Acroncosa, 174 
castronalis Heinrich, n. sp., Pseudocabima, 
53, 54 
Catastia Hiibner, 7, 110, 112 
actualis (Hulst), 110, 111 
bistriatella (Hulst), 110, 111 


300329—56——37 


INDEX 


Catastia Hiibner—Continued 
incorruscella (Hulst), 110, 111 
marginea (Schiffermiiller), 110 
cathaeretes Dyar, Anegcephalesis, 93, 94 
caudellella (Dyar), Sarata, 162, 165 
Caudellia Dyar, 186, 292, 294, 298 
albovittella Dyar, 292 
animosella (Dyar), 293 
apyrella Dyar, 292 
arizonella (Walter), 293 
clara Heinrich, n. sp., 294 
colorella (Dyar), 293 
declivella (Zeller), 292, 293, 294 
nigrella (Hulst), 283, 293 
cautella (Walker), Ephestia, 298, 302, 303, 
304 : 
Cayennia Hampson, 182, 267 
rufitinctalis Hampson, 267 
celiella Schaus, Hyalospila, 57 
celtidella (Hulst), Nephopteryx, 131, 132 
Ceracanthia Ragonot, 4, 85, 86 
mamella (Dyar), 86 
vepreculella Ragonot, 86 
ceratoniae (Zeller), Ectomyelois, 44 
cervicalis Dyar, Megarthria, 86, 87 
cervinistrigalis Walker, ‘Hypochalcia,”’ 
312 
championella Ragonot, Bethulia, 296, 297 
Chararica Heinrich, n. gen., 5, 38 
annuliferella (Dyar), 38, 39 
bicolorella (Barnes & McDunnough), 
39 
hystriculella (Hulst), 39 
chilensis Heinrich, Sigelgaita, 255 
chinographella Ragonot, Hemiptilocera, 
30, 31 
Chorrera Dyar, 6, 7, 177 
extrincica (Dyar), 177, 178 
idiotes Dyar, 177, 178 
postica (Zeller), 178 
cinctella (Hulst), Passadena, 175 
cinereella Hulst, Sarata, 162, 163 
cinerella (Hulst), Cactobrosis, 260 
cinerosella (Zeller), Euzophera, 272, 273 
cinilixa Dyar, Drescoma, 89 
cirroferella Hulst, Acrobasis, 17, 21 
cispha Dyar, Pseudodivona, 48, 49 
cistipennis (Dyar), Fundella, 59, 60 
clara Heinrich, n. sp., Caudellia, 294 
clarefacta Dyar (race of fuscomaculella 
(Wright)), Ozamia, 258 
clarioralis (Walker), Dioryctria, 150, 158 
clevelandella (Dyar), Hyalospila, 58 
climosa Dyar, ‘‘Euzophera,” 315 


clitellatella Ragonot, ‘Hornigia,” 284, 
314 

cnabella Dyar, Hypsipyla, 28 

coca (Dyar), Erelieva, 308, 309 

coccidivora (Comstock), Laetilia, 230, 


231, 232, 254 
coloradella (Hulst), Ephestiodes, 279, 280 
coloradensis Ragonot, Heterographis, 194 
colorella (Dyar), Caudellia, 293 
columbiella (McDunnough), Interjectio, 
106, 107 
columnella (Zeller), Davara, 74, 75 


569 


comacornella (Hulst), Acrobasis, 22 
commatella (Zeller), Mescinia, 212, 214 
commensella Dyar, Pseudodivona, 
49, 50 
Comotia Dyar, 185, 217 
convergens (Dyar), 217 
torsicornis Dyar, 217 
compedella (Zeller), Edulica, 271, 272 
comptella Ragonot, Acrobasis, 14, 24 
comptoniella Grossbeck (not Hulst), Acro- 
basis, 23 
comptoniella Hulst, Acrobasis, 23 
condigna Heinrich, n. sp., Anadelosemia, 69 
coniella (Ragonot), Myelopsis, 40, 41, 265 
conops (Dyar), Rabiria, 311 
conquistador Dyar, ‘‘Euzophera,” 315 
consobrinella (Zeller), Glyptocera, 100, 101 
consociata Heinrich, n. race (race of par- 
vulella Barnes & McDunnough), 
Divitiaca, 190 
constaniella Hulst, Passadena, 175 
constitutionella Dyar, Mildrixia, 26 
contatella (Grote), Nephopteryx, 124, 127, 
128 
contigua Heinrich, n. sp., Ribua, 297 
convergens (Dyar), Comotia, 217 
convolutella (Hiibner), Zophodia, 238, 239 
Coptarthria Ragonot, 3, 62, 63, 64, 66, 70 
dasypyga (Zeller), 64, 65 
coquilla (Dyar), Erelieva, 308 
coquimbella Ragonot, Valdivia, 191, 192 
cordubensiella (Ragonot), Adelperga, 187, 
196 
corniculatus Heinrich, n. sp., Paconius, 210 
corrientellus Ragonot, ‘‘Hlasmopalpus,”’ 
312 
coryliella Dyar, Acrobasis, 21 
corynophora Dyar, Difundella, 62, 63 
costella Hampson, Phestinia, 216 
crassifasciella Ragonot, Nephopteryx, 5, 
132 
crassisquamella (Hampson), Anabasis, 26 
crataegella Barnes & MceDunnough, Ne- 
phopteryx, 132 
creabates (Dyar), Eremberga, 253 
cribrella Hiibner, Myelois, 40, 43 
eriddlella Dyar, Pyla, 144, 145 
cristalis (Hampson), Magiriopsis, 94 
croceella (Hulst), Nephopteryx, 123, 131 
Crocidomera Zeller, 4, 32, 34, 276 
adonea (Felder & Rogenhofer), 33 
fissuralis (Walker), 33 
stenopteryx (Dyar), 33 
turbidella Zeller, 32, 33 
Cryptoblabes Zeller, 1, 10 
gnidiella (Millitre), 10 
bistriga (Haworth), 10 
gnidiella (Milliére), 10 
rutilella Zeller, 10 
Cuba Dyar, 76 
cubella Dyar, Homoeosoma, 219, 226 
cubensis Heinrich, n. sp., Scorylus, 72, 73 
cundajensis (Zeller), Cabotia, 201 
Cuniberta Heinrich, n. gen., 5, 34 
subtinctella (Ragonot), 34 
cunulae Dyar & Heinrich, Acrobasis, 22 


48, 


570 


curvatellus (Ragonot), Phobus, 137, 139 
cviatella Dyar, Meroptera, 122 
cyrdipsa Dyar, Drescoma, 88, 89, 263 


daedalella Ragonot, “‘Euzophera,” 314 
Dakruma Grote, 238 
dalera (Dyar), Davara, 73, 74 
dammersi Heinrich, n. sp., Nephopteryx, 
126, 127 
Dannemora Hulst, 206, 276 
Dasypyga Ragonot, 4, 69 
alternosquamella Ragonot, 69 
stictophorella Ragonot, 69 
dasypyga (Zeller), Coptarthria, 64, 65 
dasystigma Dyar, Promylea, 67 
Davara Walker, 2, 9, 73, 77, 78 
azonaxsalis Walker, 73, 75 
caricae (Dyar), 73, 74, 75 
columnella (Zeller), 74, 75 
dalera (Dyar), 73, 74 
euthales (Dyar), 75 
interjecta Heinrich, n. sp., 73, 74, 75 
nerthella (Schaus), 75 
paranensis (Dyar), 75 
rufulella (Ragonot), 73, 76 
deceptorium Heinrich, n. sp., Homoeo- 
soma, 223 
decimerella (Hulst), Stylopalpia, 141 
decipiens Dyar, Moodnopsis, 269, 270, 271 
decipientella Dyar, Nephopteryx, 132 
declivella (Zeller), Caudellia, 292, 293, 294 
decolor (Zeller), Ectomyelois, 43, 44 
decoloralis (Walker), Caristanius, 98 
decuriella (Hiibner), Dioryctria, 150 
decurrens (Dyar), Diatomocera, 51 
defectella (Walker), Ephestia, 301, 303 
definitella (Zeller), Hypargyria, 37, 38 
dea Dyar, Zamagiria, 93 
delassalis Hulst, Nephopteryx, 125, 127 
delassalis (Hulst, not Hulst), Nephopteryx, 
125, 126 
delectella}(Hulst), Dioryetria, 155 
delta Heinrich, n. sp., Sarata, 165 
demotella Grote, Acrobasis, 19 
dentata (Grote), Melitara, 241 
denticosella (Dyar), Magiriopsis, 94 
denticulella (Ragonot), Interjectio, 
107, 110, 138 
dentosella Ragonot, Vitula, 285 
deprivalis (Walker), Ufa, 171 
desuetella (Walker), Ephestia, 303 
dialithus (Dyar), Ulophora, 176, 177 
Diatomocera Ragonot, 8, 50, 53, 266 
albosigno Heinrich, n. sp., 52 
decurrens (Dyar), 51 
dosia (Dyar), 51 
excisalis (Hampson), 51 
extracta Heinrich, n. sp., 52, 53 
hoplidice (Dyar), 52 
majuscula Heinrich, n. sp., 52 
mochlophleps (Dyar), 52 
tenebricosa (Zeller), 50 
didactica Dyar, Yosemitia, 250, 251 
differtella Barnes & McDunnough, Homo- 
eosoma, 220 
diffissella (Zeller), Ancylostomia, 95 


106, 


UNITED 


STATES NATIONAL MUSEUM BULLETIN 207 


diffusella Ely, Eurythmia, 306, 307 
Difundella Dyar, 3, 60, 62, 263 
corynophora Dyar, 62, 63 
distractor Heinrich, n. sp., 64 
subsutella (Schaus), 63 
tolerata Heinrich, n. sp., 64, 65 
dilatifasciella (Ragonot), Laetilia, 230 
dilativitia Dyar, Zophodia, 239 
Dioryctria Zeller, 8, 66, 124, 149, 191, 233 
abietella (Denis & Schiffermiiller), 149, 
150, 151, 152, 156, 233 
abtetivorella (Grote), 150 
albovittella (Hulst), 127, 156 
amatella (Hulst), 156 
auranticella (Grote), 152, 153 
austriana (Cosens), 155 
baumhoferi Heinrich, n. sp., 157 
brunneella (Dyar), 158 
cambiicola (Dyar), 150, 155, 156, 157 
clarioralis (Walker), 8, 150, 158 
decuriella (Hiibner), 150 
delectella (Hulst), 155 
disclusa Heinrich, 152, 153 
elegentella (Hulst), 150 
erythropasa (Dyar), 152, 153 
gulosella (Hulst), 149, 157, 158 
horneana (Dyar), 154 
laurata (Heinrich), 153 
majorella Dyar, 152 
mendacella Staudinger, 149 
miniatella Ragonot, 153 
muellerana Dyar, 152 
pineae Staudinger, 149 
ponderosae Dyar, 152 
ponderosae Heinrich (not Dyar), 155, 
156 
pryeri Ragonot, 153 
pygmaeella Ragonot, 154 
reniculella (Grote), 151, 152 
reniculella (Packard, not Grote), 150 
subtracta Heinrich, n. sp., 157, 158 
sysstratiotes Dyar, 151, 152 
zanthaenobares Dyar, 153 
zimmermani (Grote), 149, 150, 152, 
154, 155, 156, 157 
discella Hampson, Mescinia, 213, 215 
disclusa Heinrich, Dioryctria, 152, 153 
Discopalpia Ragonot, 78, 79 
discrebile Heinrich, n. sp., Homoeosoma, 
224 
disjunctus Heinrich, n. sp., Heras, 34, 35 
dissimulatrix Heinrich, n. sp., Actrix, 140 
dissitus Heinrich, n. sp., Peadus, 84 
disticta (Zeller), ‘‘Psorosa,”’ 314 
distractor Heinrich, n. sp., Difundella, 64 
ditaeniatellum Ragonot, Homoeosoma, 224 
divergens (Butler), Vagobanta, 289 
divergens (Dyar), Sosipatra, 296 
Diviana Ragonot, 185, 206, 207 
edentella (Hulst), 206, 207 
eudoreella Ragonot, 206, 207 


Divitiaca Barnes& McDunnough, 185, 
189, 190 
ochrella Barnes & McDunnough, 189, 
190 


parvulella Barnes & McDunnough, 190 


Divitiaca Barnes & McDunnough—Cont. 
parvulella consociata, Heinrich, n. 
race, 190 
simulella Barnes & McDunnough, 189, 
190 
Divona Ragonot, 166 
ilignella (Zeller), 166 
dixolophella Dyar, Zamagiria, 90, 91 
dnopherella Ragonot, Sarata, 159, 162, 163, 
164, 165 
doddalis Dyar, Melitara, 241 
doddi Heinrich, Cactoblastis, 246, 247 
Dolichorrhinia Ragonot, 190 
dolorosa Dyar, Plodia, 298, 299 
dorae (Dyar), Unadilla, 227, 228 
dorsimacula (Schaus), Hypsipyla, 28, 29 
dosia (Dyar), Diatomocera, 51 
dotella Dyar, Honora, 197, 198 
Drescoma, Dyar, 2, 88, 262, 266 
cinilixa Dyar, 89 
cyrdipsa Dyar, 88, 89, 263 
Drescomopsis Dyar, 88, 180, 262, 266 
drucella (Dyar), 263 
soraella (Druce), 263 
subelisa Dyar, 262, 263 
drucet (Dyar), Promylea, 66 
druceii (Ragonot), Promylea, 66 
drucella (Dyar), Drescomopsis, 263 
dryadella Hulst, Ocala, 191 
dryopella Dyar (not Schaus), Moerbes 269 
dryopella (Schaus), Moerbes, 268, 269 
dubia Heinrich, n. sp., Varneria, 306 
dulciella Hulst, ‘“Honora,’’ 197, 313 
dulciella (Hulst), Anadelosemia, 68 
duplipunctella (Ragonot), Paramyelois, 47 
dyarella Ely, Acrobasis, 20, 21 
dyari Heinrich, n. name, Promylea, 66, 67 


Eccopsia Hulst, 285 
Ectomyelois Heinrich, n. gen., 8, 43 
ceratoniae (Zeller), 44 
decolor (Zeller), 43, 44 
ephestiella (Hampson), 44 
furvidorsella (Ragonot), 45, 46 
muriscis (Dyar), 45, 46, 47, 48 
oporedestella (Dyar), 44 
palpalis (Dyar), 45 
phoenicis (Durrant), 45 
transitella (Dyar, not Walker), 45 
zeteki Heinrich, n. sp., 46 
edentella (Hulst), Diviana, 206, 207 
edmandsae (Packard), Vitula, 285, 286, 
287, 288 
Edulica Ragonot, 182, 271 
compedella (Zeller), 271, 272 
edwardsialis (Hulst), Sarata, 160 
egenella (Ragonot), Hyalospila, 58 
Elasmopalpus Blanchard, 7, 97, 169, 170, 
172, 204, 312 
angustellus Blanchard, 172, 173 
anthracellus Ragonot, 173 
carbonella (Hulst), 173 
incautella (Zeller), 173 
lignosellus (Zeller), 172, 173, 204 
major (Zeller), 173 


Elasmopalpus Blanchard—Continued 
puer Dyar, 173 
tartarella (Zeller), 173 
electellum (Hulst), Homoeosoma, 220, 223, 
224 
elegentella (Hulst), Dioryctria, 150 
eliella Dyar, Acrobasis, 19 
elongatella (Hampson), Cactobrosis, 261 
elongellum Dyar, Homoeosoma, 222 
elutea (Haworth), Ephestia, 302 
elutella (Hitbner), Ephestia, 227, 301, 302, 
303, 304 
emendata Heinrich, n. sp., Moerbes, 50, 
268, 269 
emendator Heinrich n. race (variety of 
illuviellum Ragonot), Homoeosoma, 
222 
Emmerita Hampson, 121 
Encystia Hampson, 200 
Endommasis Hampson, 199 
endopyrella Hampson, Oedothmia, 205 
engeli (Dyar), Salebriaria, 116 
Entmemacornis, Dyar, 50, 53, 181, 266, 
267 
proselytes Dyar, 266, 267 
pulla Heinrich, n. sp., 266 
Ephestia Guénée, 10, 186, 202, 228, 279, 
284, 287, 293, 299, 300, 301 
amarella Dyar, 302 
cahiritella Zeller, 303 
cautella (Walker), 298, 302, 303, 304 
defectella (Walker), 301, 303 
desuetella (Walker), 303 
elutea (Haworth), 302 
elutella (Hiibner), 227, 301, 302, 303, 
304 
ernestinella Turati, 304 
ficulella Barrett, 304 
figulella Curran, 304 
figulilella Gregson, 302, 303, 304 
formosella (Wileman & South), 303 
millert Zeller, 304 
passulella Barrett, 303 
roxzburghit Gregson, 302 
rufa (Haworth), 302 
semirufa (Haworth), 302 
sericarium (Scott), 301, 302 
unicolorella Staudinger, 302 
venosella Turati, 304 
vitivora Filipjev, 302 
ephestiella (Hampson), Ectomyelois, 44 
ephestiella (Ragonot), Laetilia, 230, 234 
Ephestiodes Ragonot, 185, 277, 278, 282, 
306 
benjaminella Dyar, 279 
coloradella (Hulst), 279, 280 
erasa Heinrich, n. sp., 280 
erythrella Ragonot, 279, 280 
gilvescentella Ragonot, 278, 279, 280, 
281, 282 
granulella Hampson, 281 
indentella Dyar, 281 
infimella Ragonot, 279, 280, 281 
lucidibasella Ragonot, 278, 280 
mignonella Dyar, 280 
nigrella Hulst, 278, 283 


INDEX 


Ephestiodes Ragonot—Continued 
noniella Dyar, 278, 282, 306 


571 


Eutrichocera Hampson, 315 
paurolepidalis Hampson, 315 


plorella Dyar, 278, 281, 282, 306, 309 | Euzophera Zeller, 181, 207, 233, 272, 274, 


productella Ragonot, 281 
stictella (Hampson), 278, 281, 306 
uniformella Hampson, 281 
vestilla (Dyar), 281, 309 
Epischnia authors, 101, 102 
Epischnia Hiibner, 102 
prodromella Hiibner, 102 
epischnioides Hulst, ‘‘Zophodia,” 315 
epsilon Heinrich, n. sp., Sarata, 165 


275, 276, 287, 313 
aglaeella Ragonot, 273 
cinerosella (Zeller), 272, 273 
griselda Dyar, 274 
nigricantella Ragonot, 55, 274 
ostricolorella Hulst, 273, 274 
pallulella (Hulst), 273 
pinguis (Haworth), 272, 273 
semifuneralis (Walker), 273, 276 


erasa Heinrich, n. sp., Ephestiodes, 280 | euzopherella (Dyar), Pseudocabima, 53, 54 
Erelieva Heinrich, n. gen., 186, 307, 308 | evanescentella Dyar, Acrobasis, 17 


coca (Dyar), 308, 309 

coquilla (Dyar), 308 

hospitella (Dyar, not Zeller), 308 

mossa (Dyar), 308 

parvulella (Ely), 307, 308, 309 

quantulella (Hulst), 307, 308, 309 

santiagella (Dyar), 308 

uncta (Dyar), 308 
Eremberga Heinrich, 183, 252, 253 

creabates (Dyar), 253 

insignis Heinrich, 254 

leuconips (Dyar), 248, 253, 262 
eremiella (Dyar), Baphala, 235 
ernestinella Turati, Ephestia, 304 
erronella (Zeller), Unadilla, 228, 229 
erythrella Ragonot, Ephestiodes, 279, 280 
erythropasa (Dyar), Dioryctria, 152, 153 
estrella Barnes & McDunnough, Mescinia, 

213 

Etiella Zeller, 1, 2, 98, 99 

etiella (Treitschke), 99 

rubribasella Hulst, 99 

schisticolor Zeller, 99 

villosella Hulst, 99 

zinckenella (Treitschke), 99, 238 
etiella (Treitschke), Etiella, 99 
Eucardinia Dyar, 73, 74 
eucasis Dyar, Fundella, 61 
euchroma Dyar, Ancylostomia, 96 
eudoreella Ragonot, Diviana, 206, 207 
Eulogia Heinrich, n. gen., 181, 275 

ferruginella (Ragonot), 275 

ochrifrontella (Zeller), 275 
Eumysia Dyar, 182, 187 

fuscatella (Hulst), 188 

maidella (Dyar), 187, 188, 189 

mysiella (Dyar), 187, 188 

pallidipennella (Hulst), 187, 188, 191 

semicana Heinrich, n. sp., 188 
Eurythmasis Dyar, 184, 203, 204, 206 

ignifatua Dyar, 203, 204 
Eurythmia Ragonot, 186, 281, 306, 308 

angulella Ely, 307, 308 

diffusella Ely, 306, 307 

fumella Ely, 308 

hospitella (Zeller), 306, 307 

hospitella yavapaella Dyar, 307 

spaldingella Dyar, 307 
Eurythmidia Ragonot, 184, 204, 205 

ignidorsella (Ragonot), 204, 205 


euthales (Dyar), Davara, 75 


excantalis (Hulst), Sarata, 161, 163, 164, 
165 

excisalis (Hampson), Diatomocera, 51 

exiguella (Ragonot), Nonia, 215, 216 

exoleta (Zeller), Hemiptilocera, 30, 32 

expunctrix (Dyar & Heinrich), Pseudo- 
cabima, 55 

exsulella (Zeller), Acrobasis, 19 

extracta Heinrich, n. sp., Diatomocera, 52, 
53 

extrincica (Dyer), Chorrera, 177, 178 

Exuperius Heinrich, n. gen., 181, 274 

negator Heinrich, n. sp., 274 


faecella (Zeller), Laodamia, 145 

famula Zeller, ‘“‘“Myelois,” 312 

Farnobia Heinrich, n. gen., 184, 276, 277 
quadripuncta (Zeller), 207, 276 

fasciella Barnes & McDunnough, Pyla, 

147, 148 

fasciolalis (Hulst), Pyla, 142, 143, 149 

fearnella (Schaus), Pseudocabima, 53, 54 

feella Dyar, Pyla. 146 

feliculella Dyar, Canarsia, 202 

feltella Dyar, Acrobasis, 15, 17 


fenestrella (Packard), Lipographis, 166, 
167, 168 

feriella Hulst, Tacoma, 178 

fernaldi (Ragonot), Nephopteryx, 125, 
126, 127, 129 


fernaldialis (Hulst), Cactobrosis, 260, 261 

ferrealis (Hampson), Hypsipyla, 29 

ferruginella (Ragonot), Eulogia, 275 

ficulella Barrett, Ephestia, 304 

fieldiella (Dyar), Yosemitia, 250, 251 

fifaca (Dyar), Bema, 218, 219 

fifria Dyar, Anadelosemia, 68 

figulella Curran, Ephestia, 304 

figulilella Gregson, Ephestia, 302, 303, 304 

filiolella (Hulst), Atheloca, 82 

finitella (Walker), Tulsa, 134, 135 

fiskella Dyar, Laetilia, 230, 234, 235 

fissuralis (Walker), Crocidomera, 33 

flavicans (Ragonot, not Zeller), Piesmo- 
poda, 79 

flavicans (Zeller), Piesmopoda, 78, 79, 80 

flavicornella Ragonot, ‘‘Phycitopsis,’’ 314 

flavidorsella (Ragonot), Passadena, 175 

flavipicta Hampson, Psammia, 315 

floridellus (Hulst), Caristanius, 98 

floridensis Heinrich, n. race (race of dam- 
mersi Heinrich), Nephopteryx, 126 


572 


floridensis Heinrich, n. sp., Unadilla, 229 
fluviatella Schaus, Hypsipyla, 29 
formosella (Wileman & South), Ephestia, 
303 
formulella Schaus, ‘“‘Moodna,” 313 
fossulatella Ragonot, Oryctometopia, 158, 
159 
fosterella Hulst, Pima, 101, 102, 104, 105 
fragilella (Dyar), Myelopsis, 41 
franconiella Hulst, Zophodia, 239 
fraiella Dyar, Piesmopoda, 78, 79 
fraterna Heinrich, n. sp., Zamagiria, 92, 93 
fraudifera Heinrich n. race (race of delas- 
salis Hulst), Nephopteryx, 125 
frigidella (Packard), Pyla, 144 
fritilla Dyar, Bema, 218 
fructetella (Hulst), Salebriaria, 115, 118, 
119 
frustrator Heinrich, n. sp., Megarthria, 87 
fulgidula Heinrich, n. sp., Hyalospila, 57 
fuliginosus Heinrich, n. sp., Honorinus, 199 
Fulrada Heinrich, n. gen., 9, 71, 73 
carpasella (Schaus), 72 
querna (Dyar), 71, 72 
fulvirugella (McDunnough, not Ragonot), 
Pima, 104 
fulvirugella (Ragonot), Pima, 104, 105 
fumella Ely, Eurythmia, 308 
fumosella (Hulst), Hulstia, 196 
Fundella Zeller, 3, 59, 62, 312 
agapella Schaus, 60, 61 
ahemora Dyar, 59, 62 
argentina Dyar, 55, 59, 60, 61, 62 
cistipennis (Dyar), 59, 60 
eucasis Dyar, 61 
ignobilis Heinrich, 60, 61 
pellucens Zeller, 59, 60, 61 
pellucens Zeller (not Zeller), 61 
funerellus (Dyar), Phobus, 138, 139 
furciferella (Dyar), Olybria, 113 114 
furculella (Dyar), Sarasota, 76, 77 
furfurella (Hulst), Caristanius, 98 
furvidorsella (Ragonot), Ectomyelois, 45, 
46 
fusca (Haworth), Pyla, 144 
fuscatella (Hulst), Canarsia, 202 
fuscatella (Hulst), Eumysia, 188 
fuscifrontella Zeller, ‘“‘Nephopteryx,” 312 
fuscofasciella (Ragonot), Anagasta, 300 
fuscomaculella (Wright), Ozamia, 257 


Gabinius Heinrich, n. gen., 4, 84 
paulsoni (Ragonot), 84, 85 
gais Dyar, ‘‘Euzophera,”’ 313 
galdinella (Schaus), Tota, 170 
gamma, Heinrich, n. sp., Sarata, 164 
gemina (Haworth), Homoeosoma, 219 
geminipunctella (Ragonot), Telethusia, 137 
Gennadius Heinrich, n. gen., 184, 276, 
277 
junctor Heinrich, n. sp., 277 
georgiella (Hulst), Salebriaria, 117 
gigantella (Ragonot), Cactobrosis, 260 
gillettella (Dyar), Tulsa, 135 
gilvescentella Ragonot, Ephestiodes, 278, 
279, 280, 281, 282 


gilvibasella Hulst, Nephopteryx, 132 
gitonella (Druce), Anagasta, 300 
glabrella (Dyar), Baphala, 237 
glaucatella (Hulst), Rumatha, 248, 249 
gleditschiella (Fernald), Tlascala, 134 
glendella (Dyar) (race of lunigerella Rag- 
onot), Promylea, 66 
glomis (Dyar), Laetilia, 230, 234 
glycinivora (Matsumura), Plodia, 299 
Glyptocera Ragonot, 6, 100 
busckella (Dyar), 101 
consobrinella (Zeller), 100, 101 
gnidiella (Milliére), Cryptoblabes, 10 
goyensis (Ragonot), Baphala, 235, 236 
eraciella (Hulst), Yosemitia, 250, 251 
graciella (Hulst, not Hulst), Yosemitia, 
251 
gracilella Hulst, Canarsia, 202 
grandella (Zeller), Hypsipyla, 28, 29 
granitella (Ragonot), Pima, 106 
granulella Hampson, Ephestiodes, 281 
griselda Dyar, Euzophera, 274 
grisella Barnes & McDunnough, Bertelia, 
36, 37 
grossbecki (Barnes & McDunnough), Ac- 
robasis, 13 
grossipunctella Ragonot, “‘Myelois,” 312 
grossulariae (Riley), Zophodia, 239 
grossularialis Hiibner, Zophodia, 239 
grossulariella (Hiibner), Zophodia, 239 
groteii Ragonot, Ulophora, 176, 177 
grotella (Ragonot), Monoptilota, 89 
guarinella (Zeller), Ulophora, 177 
guatemalellus (Ragonot), Caristanius, 98 
guianalis Heinrich, n. sp., Pseudocabima, 
54 
gulosella (Hulst), 
158 
gurbyris Dyar, Ilatila, 263 


Dioryctria, 149, 157, 


hammondi (Riley), Psorosina, | 209 
hanhamella Dyar, Pyla, 146 
hapsella (Hulst), Adelphia, 169 
Harnocha Dyar, 184, 202 
velessa Dyar, 202, 203 
Harnochina Dyar, 315 
rectilinea Dyar, 315 
haywardi Heinrich, n. sp., Baphala, 236 
hebescella (Dyar, not Hulst), Acrobasis, 17 
hebescella Hulst, Acrobasis, 21 
heinrichalis (Dyar), Salebriaria, 117 
heliophila Dyar, Ozamia, 257 
hemilutella Dyar, Ozamia, 257, 259, 260 
Hemiptilocera Ragonot, 4, 30, 34, 35, 276 
bigrana (Zeller), 30, 31 
chinographella Ragonot, 30, 31 
exoleta (Zeller), 30, 32 
jecarella (Schaus), 30, 32 
letharda (Schaus), 30, 31, 32 
plumigerella (Ragonot), 30, 31, 32 
Heras Heinrich, n. gen., 4, 34 
disjunctus Heinrich, n. sp., 34, 35 
Heterographis Ragonot, 183, 193, 194, 195 
coloradensis Ragonot, 194 
ignistrigella Ragonot, 194 
morrisonella Ragonot, 193, 194 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Heterographis Ragonot—Continued 
olbiella (Hulst), 198, 194 
palloricostella (Walter), 194 
samaritanella (Zeller), 193 

holochlora (Dyar), Alberada, 245 

Homalopalpia Dyar, 73, 74 

Homoeographa Ragonot, 7, 135 
lanceolella Ragonot, 135, 136 

Homoeosoma Curtis, 184, 219, 225, 227, 

228 
acmaeopterum Ragonot, 225 
albescentellum Ragonot, 222, 223 
assitum Heinrich, n. sp., 225 
candidella Hulst, 221 
cubella Dyar, 219, 226 
deceptorium, Heinrich, n. sp., 223 
differtella Barnes & McDunnough, 
220 
discrebile Heinrich, n. sp., 224 
ditaeniatellum Ragonot, 224 
electellum (Hulst), 220, 223, 224 
elongellum Dyar, 222 
gemina (Haworth), 219 
illuviellum Ragonot, 221, 222, 223 
illuviellum emendator Heinrich, n. 
race, 222 
imitator Heinrich, n. sp., 222 
impressale Hulst, 223 
inornatellum (Hulst), 223 
longiventrellum Ragonot, 222 
musiosum Dyar, 219, 226 
nimbosellum Ragonot, 225 
noctividella Ragonot, 222 
oconequensis (Dyar), 224 
olectella (Hulst), 220. 
opalescella (Hulst), 220 
ogslarellum Dyar, 221 
oslarellum breviplicitum Heinrich, n. 
race, 221 
peregrinum Heinrich, n. sp., 224 
sinuella (Fabricius), 219 
striatellum Dyar, 221, 222 
stypticellum Grote, 220, 223 
tenuipunctella Ragonot, 220 
tezanella Ragonot, 220 
uncanale Hulst, 220 
uncanalis Ragonot (not Hulst), 223 
unionellum Ragonot, 225 
vepallidum Heinrich, n. sp., 224, 227 
homoeosomella (Zeller), Baphala, 235, 236, 
237 

Honora Grote, 85, 180, 196, 199, 210, 313 
canicostella Ragonot, 198 
dotella Dyar, 197, 198 
mellinella Grote, 197, 198 
montinatatella (Hulst), 198 
ochrimaculella (Ragonot), 197 
perdubiella (Dyar), 197, 198 
sciurella Ragonot, 198 
subsciurella Ragonot, 197, 198 

Honorinus Heinrich, n. gen., 181, 199 
fuliginosus Heinrich, n. sp., 199 

hoplidice (Dyar), Diatomocera, 52 

horneana (Dyar), Dioryctria, 154 

Hornigia Ragonot, 287 

hospitabilis Dyar, Zamagiria, 90, 91 


hospitella (Dyar, not Zeller), Erelieva, 308 
hospitella (Zeller), Eurythmia, 306, 307 
huanucensis Heinrich, Sigelgaita, 256 
Hulstia Ragonot, 183, 195 
fumosella (Hulst), 196 
oblitella (Ragonot, not Hulst), 196 
obsipella (Hulst), 169, 196 
rubiginalis (Walker), 196 
undulatella (Clemens), 169, 195, 196 
hulstiella Ragonot, ‘“Hypochaleia,’’ 312 
hulstii Cockerell, Laetilia, 230, 232 
humilis Ragonot, Lipographis, 166, 167 
Hyalospila Ragonot, 9, 56, 84 
angulinella (Schaus), 58 
celiella Schaus, 57 
clevelandella (Dyar), 58 
egenella (Ragonot), 58 
fulgidula Heinrich, n. sp., 57 
insequens Heinrich, n. sp., 57 
majorina Heinrich, n. sp., 57 
semibrunneella Ragonot, 59 
stictoneurella Ragonot, 35, 56, 57, 58 
xanthoudemia (Dyar), 58 
Hypargyria Ragonot, 2, 11, 37 
definitella (Zeller), 37, 38 
metalliferella Ragonot, 37, 38 
slossenella (Hulst), 38 
tenuella (Barnes & McDunnough), 38 
Hypermescinia Dyar, 215 
Hyphantidium Scott, 301 
Hypochaleia Hiibner, 312 
ahenella (Zeller), 312 
hypoehalciella (Ragonot), Pyla, 145, 146 
Hypsipyla Ragonot, 2, 27 
cnabella Dyar, 28 
dorsimacula (Schaus), 28, 29 
ferrealis (Hampson), 29 
fluviatella Schaus, 29 
grandella (Zeller), 28, 29 
pagodella Ragonot, 27, 28 
robusta (Moore), 27, 28 
hystriculella (Hulst), Chararica, 39 
idiotes Dyar, Chorrera, 177, 178 chill 
ignidorsella (Ragonot), Eurythmidia, 204, 
205 
ignifatua Dyar, Eurythmasis, 203, 204 
ignistrigella Ragonot, Heterographis, 194 
ignobilis (Butler), Ancylostomia, 95 
ignobilis Heinrich, Fundella, 60, 61 
ithouna Dyar, Zophodia, 239 
ilignella (Zeller), Divona, 166 
Illatila Dyar, 180, 263 
gurbyris Dyar, 263 
illuviellum Ragonot, Homoeosoma, 221, 
222, 223 
imitator Heinrich, n. sp., Homoeosoma, 222 
immorella (Dyar), Ozamia, 258 
immundella (Hulst), Myelopsis, 40 
Immyrla Dyar, 3, 111 
nigrovittella Dyar, 111, 112 
impeditella (Zeller), Cabotia, 201 
imperfecta (Dyar), Aptunga, 211 
impletella (Zeller), Prosoeuzophera, 275, 
276 


INDEX 


impostor Heinrich, n. sp., Pyla, 143, 144, 
145, 148 
impressale Hulst, Homoeosoma, 223 
inanimella (Dyar), Vitula, 287 
incanella (Hulst), Sarata, 160, 161 
incautella (Zeller), Elasmopalpus, 173 
incertus Heinrich, n. sp., Phobus, 139 
inconditella (Ragonot), Nephopteryx, 125, 
127, 128 
incorruscella (Hulst), Catastia, 110, 111 
indecora Dyar, Mescinia, 212, 213, 215 
indentella Dyar, Ephestiodes, 281 
indianella Dyar, ‘‘Megasis,’”’ 160, 315 
indigenella (Zeller), Acrobasis, 12, 14 
infimella Ragonot, Ephestiodes, 279, 280, 
281 
infinitella (Dyar), Tulsa, 135 
infusella Zeller, ‘‘Myelois,” 314 
innoxia Heinrich, Ribua, 297, 298 
inornatella (Ragonot), Moodnopsis, 270, 
271 
inornatellum (Hulst), Homoeosoma, 223 
inquilinella Ragonot, Nephopteryx, 128, 
133 
insequens Heinrich, n. sp., Hyalospila, 57 
insignatella Dyar, Cactobrosis, 260, 261 
insignis Heinrich, Eremberga, 254 
insinuatrix Heinrich, n. sp., Pyla, 144 
interjecta Heinrich, n. sp., Davara, 73, 
74, 75 
Interjectio Heinrich, n. gen., 3, 106, 108 
columbiella (McDunnough), 106, 107 
denticulella (Ragonot), 106, 107, 110, 
138 
lallatalis authors (not Hulst), 106, 107 
niviella (Hulst), 107 
ruderella (Ragonot), 107 
interpunctalis (Hiibner), Plodia, 298 
interpunctella (Hiibner), Plodia, 298, 299, 
303 
interstitialis (Dyar), Cahela, 248 
intextella (Zeller), ‘“Euzophera,” 314 
intransitella (Dyar), Adanarsa, 35 
inveterella (Dyar), Moodnopsis, 268, 271 
iota Heinrich, n. sp., Sarata, 164 
ipsetona Dyar, Zamagiria, 93 
irichampa Dyar, ‘‘Anthropteryx,” 313 
irrubriella Ely, Acrobasis, 20 
isabella (Dyar), Piesmopoda, 78, 79, 80, 81 


jocarella (Schaus), Hemiptilocera, 30, 32 

juglandis (LeBaron), Acrobasis, 15, 16, 17 

jugosella Ragonot, Ortholepsis, 119, 120 

junctolineella (Hulst), Olycella, 241, 242, 
243 

junctor Heinrich, n. sp., Gennadius, 277 


kappa Heinrich, n. sp., Sarata, 163, 165 

kearfottella Dyar, Acrobasis, 17 

kiihniella (Zeller), Anagasta, 291, 299, 300, 
302 


lacteella (Hulst), Nephopteryx, 132 

laetella Grote, Ambesa, 108 

Laetilia Ragonot, 183, 230, 231, 235, 240, 
255 


573 


Laetilia Ragonot—Continued 
amphimetra (Mayrick), 230, 233 
coccidivora (Comstock), 230, 231, 232, 
254 
coccidivora cardini Dyar, 230, 232 
coccidivora quadricolorella (Dyar), 
231, 232 
dilatifasciella (Ragonot), 230 
ephestiella (Ragonot), 230, 234 
fiskella Dyar, 230, 234, 235 
glomis (Dyar), 230, 234 
hulstii Cockerell, 230, 232 
lustrella (Dyar), 234 
melanostathma (Meyrick), 230, 232, 
233, 234 
myersella Dyar, 230, 233, 234 
obscura Dyar, 232 
pallida (Comstock), 230 
portoricensis Dyar, 232 
zamacrella Dyar, 230, 233, 234, 286 
laidion (Zeller), Zamagiria, 92 
lallatalis authors (not Hulst), Interjectio, 
106, 107 
lallatalis (Dyar, not Hulst), Phobus, 138 
lallatalis (Hulst), Ambesa, 109 
lallatalis (Hulst, not Hulst), Phobus, 138 
lambella (Dyar), Nonia, 215, 216 
lanceolella Ragonot, Homoeographa, 135, 
136 
Laodamia Ragonot, 145 
faecella (Zeller), 145 
Laosticha Hulst, 230 
Lascelina Heinrich, n. gen., 182, 264 
canens Heinrich, n. sp., 264 
latercula (Hampson), Plodia, 299 
latifasciella Dyar, Acrobasis, 20 
latifasciatella (Packard), Telethusia, 137 
lativittella (Ragonot), Valdivia, 191, 192 
laura (Dyar), Vitula, 285, 287 
laurata (Heinrich), Dioryctria, 153 
legatella (Hiibner), Rhodophaea, 24 
leithella (Dyar), Amalafrida, 256 


leoninella (Packard), Lipographis, 166, 
167, 168 

letharda (Schaus), Hemiptilocera, 30, 31, 
32 

leuconips (Dyar), Eremberga, 248, 253, 
262 


leucophaeella (Hulst), Oreana, 112, 117 
levigatella (Hulst), Nephopteryx, 129 
lignosellus (Zeller), Elasmopalpus, 
173, 204 
Lipographis Ragonot, 3, 160, 163, 166 
fenestrella (Packard), 166, 167, 168 
humilis Ragonot, 166, 167 
leoninella (Packard), 166, 167, 168 
pallidella (Dyar), 167 
subosseella Hulst, 168 
truncatella (Wright), 166, 167 
umbrella (Dyar), 160, 166, 168 
liquidambarella (Dyar), Nephopteryx, 131 
lithosella (Ragonot), Ufa, 170, 171 
longipennella (Hampson), Cactobrosis, 260, 
261 
longipennella 
251 


172, 


(Hulst), Yosemitia, 250, 


574 


longiventrellum Ragonot, 
222 

lophophora (Dyar), Azaera, 283 

lophotalis Heinrich, n. sp., Rampylla, 
70, 71 

lucidalis (Walker), Ozamia, 257, 258 

lucidibasella Ragonot, Ephestiodes, 278, 
280 

lugubrella (Ragonot), Vitula, 286 

lunigerella Hampson, Stylopalpia, 140, 141 

lunigerella Ragonot, Promylea, 65, 66 

lustrella (Dyar), Laetilia, 234 

luteella (Hulst), Ufa, 170 


Homoeosoma, 


mabes Dyar, ‘“‘Euzophera,” 315 
macropasa (Dyar), Aptunga, 211 
Macrorrhinia Ragonot, 182, 189, 190, 191, 
192, 194 
aureofasciella Ragonot, 190 
placidella (Zeller), 191 
maculicula (Dyar), Rhagea, 238 
maculifera Dyar, Cactobrosis, 260, 261 
magellanella (Ragonot), Patagonia, 225 
Magiriopsis Heinrich, n. gen., 9, 94 
cristalis (Hampson), 94 
denticosella (Dyar), 94 
magnificans Dyar, Zophodia, 239 
maidella (Dyar), Eumysia, 187, 188, 189 
major (Zeller), Elasmopalpus, 173 
majorella Dyar, Dioryctria, 152 
majorella (Dyar), Sosipatra, 296 
majorina Heinrich, n. sp., Hyalospila, 57 
majuscula Heinrich, n. sp., Diatomocera, 52 
malacella Dyar, Cassiana, 212 
malipennella Dyar, Acrobasis, 20 
mamella (Dyar), Ceracanthia, 86 
Manhatta Hulst, 185, 283, 284, 287 
biviella (Zeller), 272, 284, 287 
broweri Heinrich, n. sp., 288 
setonella (McDunnough), 287 
marginea (Schiffermiiller), Catastia, 110 
Maricopa Hulst, 191 
marmorea (Haworth), Rhodophaea, 24 
masculinus Dyar, Zamagiria, 91 
maturella (Zeller), Unadilla, 218, 228, 229 
medulallis Hubner, Myelois, 40, 43 
megalopalis Hampson, ‘‘Euzopherodes,”’ 
314 
Megaphycis Grote, 240 
Megarthria Ragonot, 3, 5, 86 
alpha Heinrich, n. sp., 88 
beta Heinrich, n. sp., 88 
cervicalis Dyar, 86, 87 
frustrator Heinrich, n. sp., 87 
peterseni (Zeller), 86, 87, 88 
schausi Heinrich, n. sp., 87 
squamifera Heinrich, n. sp., 87 
Megasis Guenée, 160 
rippertella (Zeller), 160 
melanellus (Hulst), Tulsa, 134 
melanoplaga (Hampson), Caristanius, 97 
melanostathma (Meyrick), Laetilia, 230, 
232, 233, 234 
Melia Heinemann, 272 


UNITED 


STATES NATIONAL MUSEUM BULLETIN 207 


Melitara Walker, 
244, 252 
bollit Dodd (not Zeller), 241 
bolliz (Zeller), 240 
dentata (Grote), 241 
doddalis Dyar, 241 
prodenialis Walker, 240, 241 
mellinella Grote, Honora, 197, 198 
mendacella Staudinger, Dioryctria, 149 
Meroptera Grote, 6, 13, 101, 112, 121, 
128, 124, 127, 138, 134 
abditiva Heinrich, n. sp., 123 
eviatella Dyar, 122 
mirandella Ragonot, 121, 122 
pravella (Grote), 121, 122, 123, 124, 
131 
Mescinia Ragonot, 184, 211, 212, 215, 216, 
217, 227, 293 
bacerella Dyar, 213, 214 
berosa Dyar, 214 
commatella (Zeller), 212, 214 
discella Hampson, 213, 215 
estrella Barnes & McDunnough, 213 
indecora Dyar, 212, 213, 215 
moorei Heinrich, n. sp., 214 
mosces Dyar, 213 
pandessa Dyar, 213 
parvula (Zeller), 214 
peruella Schaus, 214 
triloses Dyar, 213 
metagrammalis (Walker), Caristanius, 98 
metalicella Hulst, Pyla, 148 
metalliferella Ragonot, Hypargyria, 37, 38 
Metephestia Ragonot, 185, 264 
simplicula (Zeller), 264, 265 
micaceella (Hampson), Sosipatra, 295, 296 
Micromescinia Dyar, 184, 277 
pygmaea Dyar, 277 
Microphestia Dyar, 186, 294 
animalcula Dyar, 294 
Microphycita Dyar, 310, 311 
titillella Dyar, 310, 311 
mignonella Dyar, Ephestiodes, 280 
Mildrixia Dyar, 2, 26, 37 
constitutionella Dyar, 26 
Mineola Hulst, 10, 11 
milleri Zeller, Ephestia, 304 
mindosis Dyar, Promylea, 67 
miniatella Ragonot, Dioryctria, 153 
minimella Ragonot, Acrobasis, 11, 15 
minualis (Walker), Ufa, 171 
minutularia (Hulst), Myelopsis, 41 
minutulella (Hulst), Myelopsis, 41 
mirabella Dyar (race of walsinghami 
(Ragonot)), Ambesa, 109 
mirabilicornella (Dyar), Protasia, 193 
mirandella Ragonot, Meroptera, 121, 122 
mochlophleps (Dyar), Diatomocera, 52 
modestella (Hulst), Telethusia, 137 
Moerbes Dyar, 50, 181, 267, 268 
alveolella (Ragonot), 269 
dryopella Dyar (not Schaus), 269 
dryopella (Schaus), 268, 269 
emendata Heinrich, n. sp., 50, 268, 269 
moeschlert (Ragonot), Oryctometopia, 159 


moestella (Walker), Pyla, 144 


181, 240, 241, 243, 


Mona Hulst, 193 
monodon Dyar, Ambesa, 108 
Monoptilota Hulst, 5, 89 
grotella (Ragonot), 89 
nubilella Hulst, 69, 117 
pergratialis (Hulst), 89 
montella Schaus, Piesmopoda, 78, 81 
montinatatella (Hulst), Honora, 198 
Moodna Hulst, 185, 283, 285, 287, 288, 289, 
290, 292, 293, 313, 314 
bisinuella Hampson, 284 
obtusangulella (Ragonot), 284 
ostrinella (Clemens), 209, 284, 293 
pelviculella Hulst, 283, 284 
Moodnella Heinrich, n. gen., 184, 289 
paula Heinrich, n. sp., 289 
Moodnopsis Dyar, 181, 268, 269, 313, 314 
decipiens Dyar, 269, 270, 271 
inornatella (Ragonot), 270, 271 
inveterella (Dyar), 268, 271 
parallela Heinrich, n. sp., 271 
perangusta (Dyar), 270 
portoricensis Heinrich, n. sp., 230, 271 
moorei Heinrich, n. sp., Mescinia, 214 
morbosella (Staudinger), Staudingeria, 194 
morrisonella Ragonot, Heterographis, 193, 
194 
mosces Dyar, Mescinia, 213 
mossa (Dyar), Erelieva, 308 
mucidella (Ragonot), Rotruda, 219, 225, 
226, 227 
muciella Schaus, Azaera, 282, 283 
muellerana Dyar, Dioryctria, 152 
mundelli Heinrich, Cactoblastis, 247 
muriscis (Dyar), Ectomyelois, 45, 46, 47, 48 
musiosum Dyar, Homoeosoma, 219, 226 
Myelois Hiibner, 40, 46, 47, 312 
cribrella Hiibner, 40, 43 
medulallis Hiibner, 40, 43 
Myelopsis Heinrich, n. gen., 8, 40 
alatella (Hulst), 41 
coniella (Ragonot), 40, 41, 265 
fragilella (Dyar), 41 
immundella (Hulst), 40 
minutularia (Hulst), 41 
minutulella (Hulst), 41 
nefas (Dyar), 40 
obnupsella (Hulst), 41 
piazzella (Dyar), 41 
rectistrigella (Ragonot), 41 
subtetricella (Ragonot), 40, 41 
tetricella (Schiffermiiller), 40 
zonulella (Ragonot), 41 
myersella Dyar, Laetilia, 230, 233, 234 
myja Dyar, Bema, 217, 218, 219 
myricella Barnes & McDunnough, Acro- 
basis, 23 
Myrlaea Ragonot, 124, 130 
albistrigella Staudinger, 124 
myronella Dyar, Cabnia, 310 
mysiella (Dyar), Eumysia, 187, 188 


Nanaia Heinrich, 183, 245 

substituta Heinrich, 245 
nannodes Dyar, Varneria, 305 
nasutella Hulst, Unadilla, 227, 228, 229 


Nasutes Hampson, 315 
venata Hampson, 315, 316 
neaeriatella Grossbeck, Wunderia, 204, 205 
nebulella Dyar (not Riley), Acrobasis, 16 
nebulella (Grossbeck, not Riley), Acrobasis, 
13 
nebulella (McDunnough, not Riley), Ore- 
ana, 112 
nebulella (Riley), Acrobasis, 12, 13 
nebulo (Walsh), Acrobasis, 12 
nefas (Dyar), Myelopsis, 40 
negator Heinrich, Exuperius, n. sp., 274 
Neopyralis Bréthes, 245 
nephelepasa (Dyar), Olycella, 242, 243 
nephelepasa (Dyar, in part), Olycella, 243 
Nephopteryx Hiibner, 6, 101, 121, 123, 133, 
134, 136, 137, 188, 312 
afflictella (Hulst), 131 
basilaris Zeller, 129 
bifasciella Hulst, 130 
bisra Dyar, 133 
carneella Hulst, 128 
celtidella (Hulst), 131, 132 
contatella (Grote), 124, 127, 128 
crassifasciella Ragonot, 5, 132 
crataegella Barnes & MecDunnough, 132 
croceella (Hulst), 123, 131 
dammersi Heinrich, n. sp., 126, 127 
dammersi floridensis Heinrich, n. race, 
126 
decipientella Dyar, 132 
delassalis Hulst, 125, 127 
delassalis (Hulst, not Hulst), 125, 126 
delassalis fraudifera Heinrich, n. race, 
125 
fernaldi (Ragonot), 125, 126, 127, 129 
gilvibasella Hulst, 132 
inconditella (Ragonot), 125, 127, 128 
inquilinella Ragonot, 128, 133 
lacteella (Hulst), 132 
levigatella (Hulst), 129 
liquidambarella (Dyar), 131 
nogalesella (Dyar), 130 
pravella authors (not Grote), 124 
pudibundella (Ragonot), 125 
purpurella (Hulst), 124 
quinquepunctella (Grote), 128 
rhenella (Zincken), 123, 134 
rubescentella (Hulst), 125 
rubrisparsella (Ragonot), 131, 132 
rufibasella (Ragonot), 131 
semiobscurella (Hulst), 123, 124 
subcaesiella (Clemens), 6, 127, 128, 129 
subfuscella (Ragonot), 123, 124 
termitalis (Hulst), 129, 130 
termitalis yuconella (Dyar), 130 
teranella (Hulst), 132 
uvinella (Ragonot), 123, 130 
vetustella (Dyar), 127 
virgatella (Clemens), 128 
nerthella (Schaus), Davara, 75 
neuricella (Zeller), Bema, 218 
nexa Heinrich, n. sp., Rioja, 267, 268 
Nicetiodes Schaus, 185, 304 
apianella Schaus, 304, 314 
nigrella (Hulst), Caudellia, 283, 293 


obtusangulella (Ragonot), 


INDEX 


nigrella Hulst, Ephestiodes, 278, 283 
nigricantella Ragonot, Euzophera, 55, 274 
nigricans Hulst, ‘“‘Salebria,’’? 312 
nigricula Heinrich, n. sp., Pyla, 148 
nigrifasciella Ragonot, Sarata, 162, 163, 165 
nigristrigella (Ragonot), Pseudocabima, 
54, 55 
nigritella (Hampson), Oncolabis, 199 
nigrosignella Hulst, Acrobasis, 15 
nigrovittella Dyar, Immyrla, 111, 112 
nimbella (Zeller), Rotruda, 219, 226 
nimbosellum Ragonot, Homoeosoma, 225 
niviella (Hulst), Interjectio, 107 
noctividella Ragonot, Homoeosoma, 222 
nodoses (Dyar), Azaera, 283 
nogalesella (Dyar), Nephopteryx, 130 
Nonia Ragonot, 180, 184, 213, 215, 216 
exiguella (Ragonot), 215, 216 
lambella (Dyar), 215, 216 
noniella Dyar, Ephestiodes, 
306 
nonparilella (Dyar), Sosipatra, 296 
normella Dyar, Acrobasis, 20 
notatalis (Walker), Paramyelois, 47 
nubiferella (Ragonot, Salebriaria, 
117 
nubilella Hulst, Monoptilota, 89, 117 
nymphaeella (Hulst), Palatka, 207, 208 
nyssaecolella (Dyar), Actrix, 139, 140 


278, 282, 


116, 


oberthuriella (Ragonot), Olybria, 113 
oblitella (Ragonot, not Hulst), Hulstia, 196 
obnupsella (Hulst), Myelopsis, 41 
obscura Dyar, Laetilia, 232 
obsipella (Hulst), Hulstia, 169, 196 
obstitella (Schaus), Anadelosemia, 68 
Moodna, 
Ocala Hulst, 189, 191, 192, 194 

dryadella Hulst, 191 

platanella (Grossbeck), 191 
occidentalis Heinrich, n. race (race of 


284 


albiplagiatella (Packard)), Pima, 
103 

ochrella Barnes & McDunnough, Divitiaca, 
189, 190 

ochrifrontella (Zeller), Eulogia, 275 

ochrimaculella (Ragonot), Honora, 197 


ochripunctella (Dyar), Adelphia, 169 
ochrodesma (Zeller), Anabasis, 25, 26 
oconequensis (Dyar), Homoeosoma, 224 
odiosella Weinrich (not Hulst), Ozamia, 
257, 258 
odiosellus (Hulst), Salebriacus, 114 
Oedothmia Hampson, 185, 205 
bahamasella (Hampson), 205 
endopyrella Hampson, 205 
olbiella (Hulst), Heterographis, 193, 194 
olectella (Hulst), Homoeosoma, 220 
olivacea Heinrich, n. race (race of goyensis 
(Ragonot)), Baphala, 236 
olivaceella (Ragonot), race of mucidella 
(Ragonot)), Rotruda, 226 
olivacella Dyar, Staudingeria, 194, 195 
olivella Hampson, ‘‘Moodna,” 284, 314 
olivella (Hulst), Ragonotia, 160, 315 


575 


Olybria, Heinrich, 6, 113, 114 
aliculella (Hulst), 113, 114 
furciferella (Dyar), 113, 114 
oberthuriella (Ragonot), 113 
Olyca Walker, 181, 243, 253 
phryganoides Walker, 243 
Olycella Dyar, 240, 241, 243, 256 
junctolineella (Hulst), 241, 242, 243 
junctolineella pectinatella (Hampson), 
242 
nephelepasa (Dyar), 242, 243 
nephelepasa (Dyar, in part), 243 
subumbrella (Dyar), 242, 243 
Oncolabis Zeller, 181, 199, 203, 206 
anticella Zeller, 199, 206 
nigritella (Hampson), 199 
opalescella (Hulst), Homoeosoma, 220 
oporedestella (Dyar), Ectomyelois, 44 
Orena Hulst, 6, 111, 112 
leucophaeella (Hulst), 112, 117 
nebulella (McDunnough, not Riley), 


112 
unicolorella (Hulst), 13, 112 
oregonella (Barnes & MeDunnough), 
Tulsa, 135 


orio Dyar, Rampylla, 70 
orobanchella (Dyar), Rhagea, 237 
Ortholepis Ragonot, 3, 112, 119, 121, 124 
jugosella Ragonot, 119, 120 
pasadamia (Dyar), 120 
Oryctometopia Ragonot, 1, 9, 158 
fossulatella Ragonot, 158, 159 
moeschleri (Ragonot), 159 
oslarellum Dyar, Homoeosoma, 221 
ostricolorella Hulst, Euzophera, 273,274 
ostrinella (Clemens), Moodna, 209, 284, 
293 
ostryella Ely, Acrobasis, 21 
ovalis (Packard), Telethusia, 
138 
Ragonot, 183, 257 
fuscomaculella (Wright), 257 
fuscomaculella clarefacta Dyar, 258 
heliophila Dyar, 257 
hemilutella Dyar, 257, 259, 260 
immorella (Dyar), 258 
lucidalis (Walker), 257, 258 
odiosella Heinrich (not Hulst), 257, 258 
punicans Heinrich, 257, 259 
stigmaferella Dyar, 258 
thalassophila Dyar, 258 


136, 137, 


pachytaeniella (Ragonot), Volatica, 290, 
291 
packardella (Ragonot), Rhagea, 237, 238 
Paconius Heinrich, n. gen., 181, 210 
corniculatus Heinrich, n. sp., 210 
pagodella Ragonot, Hypsipyla, 27, 28 
Palatka Hulst, 185, 207 
nymphaeella (Hulst), 207, 208 
verecuntella (Grossbeck), 208 
pallicornella (Ragonot), Trachycera, 25 
pallida (Comstock), Laetilia, 230 
pallidella (Dyar), Lipographis, 167 
pallidipennella (Hulst), Eumysia, 187, 188, 
191 


576 


palliolella Dyar (not Ragonot), Acrobasis, 
16 
palliolella Ragonot, Acrobasis, 15, 16 
palloricostella (Walter), Heterographis, 194 
pallulella (Hulst), Euzophora, 273 
palpalis (Dyar), Ectomyelois, 45 
pandessa, Dyar, Mescinia, 213 
parabates (Dyar), Alberada, 244, 245 
parallela Heinrich, n. sp., Moodnopsis, 271 
Paramyelois Heinrich, n. gen., 8, 46 
cassiae (Dyar), 47 
duplipunctella (Ragonot), 47 
notatalis (Walker), 47 
soliiella (Zeller), 46, 47 
transitella (Walker), 47 
ventpars (Dyar), 47 
paranensis (Dyar), Davara, 75 
parasitaria Heinrich, n. sp., Vezina, 291 
parkerella (Schaus), Pima, 102, 106 
Parolyca Dyar, 183, 255 
asthenosoma (Dyar), 255 
parva Heinrich, n. sp., Piesmopoda, 80 
parvula (Zeller), Mescinia, 214 
parvulella Barnes & MceDunnough, Diviti- 
aca, 198 
parvulella (Ely), Erelieva, 307, 308, 309 
pasadamia (Dyar), Ortholepis, 120 
Passadena Hulst, 2, 175, 187 
canescentella (Hulst), 175 
cinctella (Hulst), 175 
constantella Hulst, 175 
flavidorsella (Ragonot), 175 
passulella Barrett, Ephestia, 303 
Patagonia Ragonot, 184, 219, 225 
magellanella (Ragonot), 225 
patriciella (Dyar), Ribua, 298 
Patriciola Heinrich, n. gen., 181, 209, 210 
semicana Heinrich, n. sp., 209, 210 
paula Heinrich, n. sp., Moodnella, 289 
paulsoni (Ragonot), Gabinius, 84, 85 
paurolepidalis Hampson, Eutrichocera, 315 
Peadus Heinrich, n. gen., 8, 83, 85 
burdetellus (Schaus), 83, 84 
dissitus Heinrich, n. sp., 84 
semproniella (Schaus), 84 
subaquilellus (Ragonot), 84 
pectinatella (Hampson) (race of junctolin- 
eella (Hulst)), Olycella, 242 
pellucens Zeller, Fundella, 59, 60, 61 
pellucens Zeller (not Zeller), Fundella, 61 
pellucidellus (Ragonot), Caristanius, 97, 98 
pelviculella Hulst, Moodna, 283, 284 
peplifera Dyar, Acrobasis, 18, 19 
perangusta (Dyar), Moodnopsis, 270 
perdubiella (Dyar), Honora, 197, 198 
peregrinum Heinrich, n. sp., Homoeosoma, 
224 
perfuscalis Hulst, Sarata, 162, 164, 165 
pergratialis (Hulst), Monoptilota, 89 
perluteella Dyar, Staudingeria, 195 
perrensiella (Ragonot), Pseudocabima, 55 
peruella Schaus, Mescinia, 214 
peterseni (Zeller), Megarthria, 86, 87, 88 
petrella (Zeller), Adelphia, 168, 169, 170 
Phestinia Hampson, 184, 216 
costella Hampson, 216 


phi Heinrich, n. sp., Sarata, 163, 165 
Philodema, Heinrich, n. gen., 3, 165, 166 
rhoidella (Dyar), 160, 165, 166 

Phobus, Heinrich, n. gen., 5, 138 
brucei (Hulst), 110, 138, 139 
curvatellus (Ragonot), 137, 139 
funerellus (Dyar), 138, 139 
incertus Heinrich, n. sp., 139 
lallatalis (Dyar, not Hulst), 138 
lallatalis (Hulst, not Hulst), 138 
rhypodella (Ragonot, not Hulst), 139 
phoenicis (Durrant), Ectomyelois, 45 
phoenicis (Dyar), Cahela, 248 
phryganoides Walker, Olyca, 243 
phycidea Zeller, 219 
Phycitidae, 1, 150 
Phycitinae, 1 
Phycitopsis Ragonot, 314 
piazzella (Dyar), Myelopsis, 41 
Piesmopoda Zeller, 9, 56, 74, 76, 77 
apocerastes Dyar, 78, 80, 81 
flavicans (Ragonot, not Zeller), 79 
flavicans (Zeller), 78, 79, 80 
fratella Dyar, 78, 79 
isabella (Dyar), 78, 79, 80, 81 
montella Schaus, 78, 81 
parva Heinrich, n. sp., 80 
ragonoti (Dyar), 78, 79 
rubicundella Zeller, 77, 78, 79 
semirufella (Zeller), 80, 81 
trichomata (Zeller), 78 
xanthomera Dyar, 78 
xanthopolys Dyar, 79, 80 
zanthozona Dyar, 78 
Pima, Hulst, 3, 101, 106, 108 
albiplagiaiella Hulst (not Packard), 102, 
104 
albiplagiatella (Packard), 103, 104 
albiplagiatella (Ragonot, not Packard), 
104 
albiplagiatella occidentalis Heinrich, 
n. race, 103 
albocostalialis (Hulst), 103, 104 
albocostalialis subcostella (Ragonot), 
105 
albocostalis (Hulst), 104 
boisduvaliella (Guénée), 102, 103, 105 
fosterella Hulst, 101, 102, 104, 105 
fulvirugella (McDunnough, not Rag- 
onot), 104 
fulvirugella (Ragonot), 104, 105 
granitella (Ragonot), 106 
parkerella (Schaus), 102, 106 
piperella (Dyar), 106 
vividella (McDunnough), 104 
pineae Staudinger, Dioryctria, 149 
pinei Heinrich, n. sp., Vitula, 286, 288 
pinguis (Haworth), Euzophera, 272, 273 
Pinipestis Grote, 150, 154 
piperella (Dyar), Pima, 106 
placidella (Dyar), Anderida, 211 
placidella (Zeller), Macrorrhinia, 191 
platanella (Grossbeck), Ocala, 191 
Plodia Guénée, 186, 292, 297, 298 
dolorosa Dyar, 298, 299 
glycinivora (Matsumura), 299 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Plodia Guénée—Continued 
interpunctalis (Hiibner), 298 
interpunctella (Hiibner), 298, 299, 303 
latercula (Hampson), 299 
zeae (Fitch), 299 
plorella Dyar, Eiphestiodes, 278, 281, 282, 
306, 309 
plumigerella (Ragonot), Hemiptilocera, 30, 
31, 32 
plumigerella Hulst, Sarasota, 76, 77 
pneumatella (Hulst), Canarsia, 202 
pogerythrus Dyar, Zamagiria, 91 
polingella (Dyar), Rumatha, 249, 250 
Polopeustis, Ragonot, 2, 119, 120 
annulatella (Zetterstedt), 120, 121 
aretiella (Gibson), 121 
polydectella (Schaus), Rampylla, 70, 71 
polyphemella (Ragonot), Sarata, 160 
pombra (Dyar), Pseudocabima, 54 
ponderosae Dyar, Dioryctria, 152 
ponderosae Heinrich (not Dyar), Dioryctria, 
155, 156 
ponderosella (Barnes & McDunnough), 
Cahela, 247, 248, 249 
porrecta Dyar, Tucumania, 252 
portoricensis Dyar, Laetilia, 232 
portoricensis Heinrich, n. sp., Moodnopsis, 
230, 271 
postflavida Dyar, ‘“Huzophera,”’ 312 
postica (Zeller), Chorrera, 178 
postremella Dyar, Varneria, 305 
Praedonula Heinrich, n. gen., 9, 82, 84 
almonella (Dyar), 82, 83 
pravella (authors, not Grote), Neph- 
opteryx, 124 
pravella (Grote), Meroptera, 121, 122, 123, 
124, 131 
Procandiopa Dyar, 85 
prodenialis Walker, Melitara, 240, 241 
prodromella Hiibner, Hpischnia, 102 
productella Ragonot, Ephestiodes, 
Promylea Ragonot, 4, 65, 85 
dasystigma Dyar, 67 
drucei (Dyar), 66 
drucei (Ragonot), 66 
dyari Heinrich, n. name, 66, 67 
lunigerella Ragonot, 65, 66 
lunigerella glendella (Dyar), 66 
mindosis Dyar, 67 
zimmermannt (Druce, not Grote), 66 


281 


proselytes Dyar, Entmemacornis, 266, 
267 

Prosoeuzophera Heinrich, n. gen., 185, 
275 


impletella (Zeller), 275, 276 
Protasia Heinrich, n. gen, 182, 193 
mirabilicornella (Dyar), 193 
Protomoerbes, Heinrich, n. gen., 8, 49 
aberrans Heinrich, n. sp., 49 
separabilis Heinrich, n. sp., 50 
pryeri Ragonot, Dioryctria, 153 
Psammia Hampson, 315 
flavipicta Hampson, 315 
Pseudocabima Heinrich, n. gen., 8, 50, 53 
arizonensis Henrich, n. sp., 55 
castronalis Heinrich, n. sp., 53, 54 


Pseudocabima Heinrich—Continued 
euzopherella (Dyar), 53, 54 
expunctrix (Dyar & Heinrich), 55 
fearnella (Schaus), 53, 54 
guianalis Heinrich, n. sp., 54 
nigristrigella (Ragonot), 54, 55 
perrensiella (Ragonot), 55 
pombra (Dyar), 54 
rubrizonalis (Hampson), 52, 53, 55 

Pseudodivona, Dyar, 8, 48, 49, 268 
carabayella Dyar, 48, 49 
cispha Dyar, 48, 49 
commensella Dyar, 48, 49, 50 
santa-maria Dyar, 48, 49 

Psorosa Zeller, 210 

Psorosina Dyar, 180, 208 
angulella Dyar, 208, 209 
hammondi (Riley), 209 

ptychis (Bondar, not Dyar), Atheloca, 82 

ptychis (Dyar), Atheloca, 82 

ptyonopoda (Hampson), Sarasota, 77 

pudibundella (Ragonot), Nephopteryx, 125 

puer Dyar, Elasmopalpus, 173 

pulla Heinrich, n. sp., Entmemacornis, 

266 
pullatella (Raganot), 
165 
pumilella (Ragonot), Salebriaria, 115, 117 
punctella (Dyar), Sarata, 161 
punicans Heinrich, Ozamia, 257, 259 
purgatoria (Dyar), Cahela, 248 
purpurella (Hulst), Nephopteryx, 124 
putidella Schaus, ‘“Eucampyla,” 313 
pygmaea Dyar, Micromescinia, 277 
pygmaeella Ragonot, Dioryctria, 154 
Pyla, Grote, 5, 6, 140, 142, 145, 169 
aeneella Hulst, 147, 148 
aeneoviridella Ragonot, 147, 148 
aenigmatica Heinrich, n. sp., 144, 145 
aequivoca Heinrich, n. sp., 143, 145 
blackmorella Dyar, 145 
cacabella (Hulst), 145 
criddlella Dyar, 144, 145 
fasciella Barnes & McDunnough, 147, 
148 

fasciolalis (Hulst), 142, 143, 149 

feella Dyar, 146 

frigidella (Packard), 144. 

fusca (Haworth), 144 

hanhamella Dyar, 146 

hypochalciella (Ragonot), 145, 146 

impostor Heinrich, n. sp., 143, 144, 
145, 148 

insinuatrix Heinrich, n. sp., 144 

metalicella Hulst, 148 

moestella (Walker), 144 

nigricula Heinrich, n. sp., 148 

rainierella Dyar, 147, 148 

scintillans (Grote), 142, 146, 147, 148 

sylphiella Dyar, 147 

triplagiatella (Dyar), 145 

viridisuffusella Barnes & McDun- 
nough, 142, 149 

pyllis (Dyar), Unadilla, 228 
pytrhochrellus (Ragonot), Ufa, 171 


Sarata, 160, 164, 


INDEX 


quadricolorella (Dyar) (race of coccidivora 
(Comstock)), Laetilia, 231, 232 
quadripuncta (Zeller), Farnobia, 207, 276 
quantulella (Hulst), Erelieva, 307, 308, 
309 
Quasisalebria Heinrich, n. gen., 7, 111, 
112, 118 
admixta Heinrich, n. sp., 118, 119 
quercicolella (Ragonot), Salebriaria, 117 
querna (Dyar), Fulrada, 71, 72 
quinquepunctella (Grote), Nephopteryx, 
128 


Rabiria Heinrich, n. gen., 310, 311 
conops (Dyar), 311 
ragonoti (Dyar), Piesmopoda, 78, 79 
Ragonotia olivella (Hulst), 160, 315 
rainierella Dyar, Pyla, 147, 148 
Ramphodes Guénée, 99 
Rampylla Dyar, 4, 9, 62, 63, 70, 72 
lophotalis Heinrich, n. sp., 70, 71 
orio Dyar, 70 
polydectella (Schaus), 70, 71 
subcaudata (Dyar), 71 
rectilinea Dyar, Harnochina, 315 
rectistrigella (Dyar), Salebriaria, 118 
rectistrigella (Ragonot), Myelopsis, 41 
reductella (Walker), Tlascala, 133, 134 
reliquella (Dyar) (race of mucidella (Rag- 
onot)), Rotruda, 219, 226 
Relmis Dyar, 218 
reniculella (Grote), Dioryetria, 151, 152 
reniculella (Packard, not Grote), Dioryctria 
150 
restrictella Zeller, ‘‘Myelois,”’ 312 
Rhagea Heinrich, n. gen., 1, 183, 230, 237 
maculicula (Dyar), 238 
orobanchella (Dyar), 237 
packardella (Ragonot), 237, 238 
stigmella (Dyar), 237, 238 
rhenella (Zincken), Nephopteryx, 123, 134 
Rhodophaea Guénée, 2, 24, 25 
advenella (Zincken), 24, 25 
caliginella (Hulst), 15, 24, 25 
caliginoidella (Dyar), 24 
legatella (Hiibner), 24 
marmorea (Haworth), 24 
suavella (Zincken), 24 
supposita (Heinrich), 15, 24 
rhoiella (Dyar), Philodema, 160, 165, 166 
rhypodella (Hulst), Telethusia, 137 
rhypodella (Ragonot, not Hulst), Phobus, 
139 
rhythmatica Dyar, Cabotia, 200, 201 
Ribua Heinrich, n. gen., 186, 297, 298 
contigua Heinrich, n. sp., 297 
innoxia Heinrich, 297, 298 
patriciella (Dyar), 298 
rileyella (Ragonot), Sosipatra, 294, 295, 297 
rinmea Dyar, ‘‘Euzophera,’’ 313 
Rioja Heinrich, n. gen., 182, 267 
nexa Heinrich, n. sp., 267, 268 
rippertella (Zeller), Megasis, 160 
robusta (Moore), Hypsipyla, 27, 28 
robustella (Dyar), Salebriaria, 116 
ronnai (Bréthes), Cactoblastis, 245, 246 


577 


roseitinctella (Dyar), Ufa, 171 
Rotruda Heinrich, n. gen., 184, 225 
mucidella (Ragonot), 219, 225, 226, 
227 
mucidella affusella (Ragonot), 219, 227 
mucidella olivaceella (Ragonot), 226 
mucidella reliquella (Dyar), 219, 226 
nimbella (Zeller), 219, 226 
roxburghii Gregson, Ephestia, 302 
rubedinella (Zeller), Ufa, 170, 171 
rubescentella (Hulst), Nephopteryx, 125 
rubicundella Zeller, Piesmopoda, 77, 78, 79 
rubiginalis (Walker), Hulstia, 196 
rubiginella (Walker), Adelphia, 169 
rubribasella Hulst, Etiella, 99 
rubrifasciella Packard, Acrobasis, 23 
rubripurpurea Hampson, Stylobasis, 205, 
206 
rubrisparsella 
131, 132 
rubrithoracella (Barnes & McDunnough), 
Sarata, 163 
rubrizonalis (Hampson), 
52, 53, 55 
ruderella (Ragonot), Interjectio, 107 
rufa (Haworth), Ephestia, 302 
rufescentalis (Walker), Ufa, 171 
rufibasella (Ragonot), Nephopteryx, 131 
rufinalis (Walker), Adelphia, 169 
rufitinctalis Hampson, Cayennia, 267 
rufulella (Ragonot), Davara, 73, 76 
Rumatha Heinrich, 182, 184, 248 
bihinda (Dyar), 248, 249 
glaucatella (Hulst), 248, 249 
polingella (Dyar), 249, 250 
russeolus Heinrich, n. sp., Birinus, 36 
rusto (Dyar), Baphala, 236 
rutilella Zeller, Cryptoblabes, 10 


(Ragonot), Nephopteryx, 


Pseudocabima, 


saissetiae (Dyar), Baphala, 236 
Salambona Heinrich, 1838, 254, 255 
analamprella (Dyar), 254 
Salebria Hiibner, 101, 112, 113, 114, 115, 
119, 124, 153, 158, 312 
Salebriacus Heinrich, n. gen., 7, 113, 
115 
bakerella (Dyar), 114 
odiosellus (Hulst), 114 
yumaella (Dyar), 114 
Salebriaria Heinrich, n. gen., 7, 114, 
118 
ademptandella (Dyar), 115 
annulosella (Ragonot), 116, 117 
engeli (Dyar), 116 
fructetella (Hulst), 115, 118, 119 
georgiella (Hulst), 117 
heinrichalis (Dyar), 117 
nudiferella (Ragonot), 116, 117 
pumilella (Ragonot), 115, 117 
quercicolella (Ragonot), 117 
rectistrigella (Dyar), 118 
robustella (Dyar), 116 
tenebrosella (Hulst), 117 
turbidella (Ragonot), 115, 116 
samaritanella (Zeller), Heterographis, 193 
santa-maria Dyar, Pseudodivona, 48, 49 


114, 


115, 


578 


santiagella (Dyar), Erelieva, 308 
Sarasota, Hulst, 9, 74, 76 
furculella (Dyar), 76, 77 
plumigerella Hulst, 76, 77 
ptyonopoda (Hampson), 77 
Sarata, Ragonot, 3, 159, 165, 166, 168 
alpha Heinrich, n. sp., 164 
aridella (Dyar), 161 
atrella (Hulst), 2, 162, 164, 165 
beta Heinrich, n. sp., 164 
caudellella (Dyar), 162, 165 
cinereella Hulst, 162, 163 
delta Heinrich, n. sp., 165 
dnopherella Ragonot, 159, 162, 163, 
164, 165 
edwardsialis (Hulst), 160 
epsilon Heinrich, n. sp., 165 
excantalis (Hulst), 161, 163, 164, 165 
gamma Heinrich, n. sp., 164 
incanella (Hulst), 160, 161 
iota Heinrich, n. sp., 164 
kappa Heinrich, n. sp., 163, 165 
nigrifasciella Ragonot, 162, 163, 165 
perfuscalis Hulst, 162, 164, 165 
phi Heinrich, n. sp., 163, 165 
polyphemella (Ragonot), 160 
pullatella (Ragonot), 160, 164, 165 
punctella (Dyar,) 161 
punctella septentrionaria Heinrich, n. 
race, 161 
tubrithoracella (Barnes & McDun- 
nough), 163 
tephrella Ragonot, 162, 163 
sauciella (Zeller), Ancylostomia, 96 
schausella Dyar, Strephomescinia, 227 
schausi Heinrich, n. sp., Megarthria, 87 
schini (Berg), Cabotia, 201 
schisticolor Zeller, Etiella, 99 
scintillans (Grote), Pyla, 142, 146, 147, 148 
Sciota Hulst, 123 
scitulella (Hulst), Acrobasis, 14 
sciurella Ragonot, Honora, 198 
scobiella (Grote), Stylopalpia, 141, 142 
Scorylus, Heinrich, n. gen., 3, 72 
cubensis Heinrich, n. sp., 72, 73 
secundella Ely, Acrobasis, 21 
Selga Heinrich, n. gen., 182, 265, 266 
arizonella (Hulst), 265 
Sematoneura, Ragonot, 2, 27, 28 
abitus Heinrich, n. sp., 27 
atrovenosella Ragonot, 27, 51 
semibrunneella Ragonot, Hyalospila, 59 
semicana Heinrich, n. sp., Humysia, 188 
semicana Heinrich, n. sp., Patriciola, 209, 
210 
semidiscella Ragonot, Cabotia, 200, 201 
semifuneralis (Walker), Euzophera, 273, 
276 
semiobscurella (Hulst), Nephopteryx, 123, 
124 
semirufa (Haworth), Ephestia, 302 
semirufella (Zeller), Piesmopoda, 80, 81 
semtproniella (Schaus), Peadus, 84 
Seneca Hulst, 11, 12 
senesciella (Schaus), 
68, 69 


Anadelosemia, 67, 


senorella (Ragonot), Anderida, 211 
senta Heinrich, n. sp., Ufa, 170, 171 
separabilis Heinrich, n. sp. Protomoerbes, 
50 
septentrionaria Heinrich, n. race (race of 
punctella (Dyar)), Sarata, 161 
septentrionella Dyar, Acrobasis, 19 
sericaria (authors, not Scott), Anagasta, 
300 
sericartum (Scott), Ephestia, 301, 302 
serratilineella Ragonot, n. status (race of 
edmandsae (Packard)), Vitula, 285, 
286 
setonella (McDunnough), Manhatta, 287 
Sigelgaita Heinrich, 183, 255 
chilensis Heinrich, 255 
huanucensis Heinrich, 256 
transilis Heinrich, 256 
similella Barnes & McDunnough, Acron- 
cosa, 174 
simplicula (Zeller), Metephestia, 264, 265 
simulella Barnes & McDunnough, Divi- 
tiaca, 189, 190 
sinuella (Fabricius), Homoeosoma, 219 
slossonella (Hulst), Hypargyria, 38 
solitella (Zeller), Paramyelois, 46, 47 
sonorella (Ragonot), Anderida, 211 
soraella (Druce), Drescomopsis, 263 
Sosipatra Heinrich, n. gen., 185, 294 
anthophila (Dyar), 295 
divergens (Dyar), 296 
majorella (Dyar), 296 
micaceella (Hampson), 295, 296 
nonparilella (Dyar), 296 
rileyella (Ragonot), 294, 295, 297 
thurberiae (Dyar), 296 
spaldingella Dyar, Eurythmia, 307 
squalida (Walker), Baphala, 236, 237 
squalidella (Dyar), Azaera, 282 
squamifera Heinrich, n. sp., Megarthria, 87 
Staudingeria Ragonot, 183, 194, 195 
albipenella (Hulst), 194, 195 
morbosella (Staudinger), 194 
olivacella Dyar, 194, 195 
perluteella Dyar, 195 
stenopteryx (Dyar), Crocidomera, 33 
Stenoptycha Heinemann (not Zeller), 272 
stercorea (Zeller), Ancylostomia, 95, 96 


stictella (Hampson), Ephestiodes, 278, 
281, 306 

stictoneurella Ragonot, Hyalospila, 35, 
56, 57, 58 


stictophorella Ragonot, Dasypyga, 69 
stigmaferella Dyar, Ozamia, 258 
stigmella Dyar, Acrobasis, 18 
stigmella (Dyar), Rhagea, 237, 238 
Strephomescinia Dyar, 184, 227 
schausella, Dyar, 227 
striatellum Dyar, Homoeosoma, 221, 222 
striella Dyar, Zamagiria, 93 
strigalis (Barnes & McDunnough), Cacto- 
brosis, 248, 253, 260, 262 
Strymaz Dyar, 227 
Stylobasis Hampson, 185, 205 
rubripurpurea Hampson, 205, 206 


UNITED STATES NATIONAL MUSEUM BULLETIN 207 


Stylopalpia Hampson, 3, 5, 140 
argentinensis Heinrich, n. sp., 141 
decimerella (Hulst), 141 
lunigerella Hampson, 140, 141 
scobiella (Grote), 141, 142 

stypticellum Grote, Homoeosoma, 

223 

suavella (Zincken), Rhodophaea, 24 

subaquilellus (Ragonot), Peadus, 84 

subcaesiella (Clemens), Nephopteryx, 6, 

127, 128, 129 

subcanella (Zeller), “‘Zophodia,” 314 

subcaudata (Dyar), Rampylla, 71 

subcostella (Ragonot) (race of albocostal- 

ialis (Hulst)), Pima, 105 
subelisa Dyar, Drescomopsis, 262, 263 
subfuscella (Ragonot), Nephopteryx, 123, 
124 

submedianella Dyar, Tacoma, 178 

subosseella Hulst, Lipographis, 166, 168 

subrufella (Hulst), Atheloca, 81, 82, 83 

subsciurella Ragonot, Honora, 197, 198 

substituta Heinrich, Nanaia, 245 

subsutella (Schaus), Difundella, 63 

subtetricella (Ragonot), Myelopsis, 

Al 
subtinctella (Ragonot), Cuniberta, 34 
subtracta) Heinrich, n. sp., Dioryctria, 
157, 158 

subumbrella (Dyar), Olycella, 242, 243 

supplicella (Dyar), Verina, 288 

supposita (Heinrich), Rhodophaea, 

24 

sylphiella Dyar, Pyla, 147 

sylviella Ely, Acrobasis, 16, 21 

Synothmia Hampson, 205 

sysstratiotes Dyar, Dioryctria, 151, 152 


220, 


40, 


15. 


taboga (Dyar), Baphala, 236 
Tacoma Hulst, 178 
feriella Hulst, 178 
submedianella Dyar, 178 
tapiacola Dyar, Tucumania, 252 
tartarella (Zeller), Elasmopalpus, 173 
tecmessella (Schaus), Anadelosemia, 67 
Telethusia Heinrich, n. gen., 5, 136 
geminipunctella (Ragonot), 137 
latifasciatella (Packard), 137 
modestella (Hulst), 137 
ovalis (Packard), 136, 137, 138 
rhypodella (Hulst), 137 
tenebricosa (Zeller), Diatomosera, 50 
tenebrosella (Hulst), Salebriaria, 117 
tenuella (Barnes & McDunnough), 
pargyria, 38 
tenuipunctella Ragonot, Homoeosoma, 220 
tephrella Ragonot, Sarata, 162, 163 
tephrosiella Dyar, Ulophora, 176 
termitalis (Hulst), Nephopteryx, 129, 130 
tetricella (Schiffermiiller), Myelopsis, 40 
texanella (Hulst), Anadelosemia, 68, 69 
texanella (Hulst), Nephopteryx, 132 
tecanella Ragonot, Homoeosoma, 220 
thalassophila Dyar, Ozamia, 258 
thurberiae (Dyar), Sosipatra, 296 
ticitoa (Dyar), Vitula, 287 


Hy- 


tintilla Dyar, ‘“‘Euzophera,” 315 
titillella Dyar, Microphycita, 310, 311 
Tlascala Hulst, 3, 133 

gleditschiella (Fernald), 134 

reductella (Walker), 133, 134 
tolerata Heinrich, n. sp., Difundella, 64, 65 
torsicornis Dyar, Comotia, 217 
Tota Heinrich, n. gen., 6, 169, 170 

galdinella (Schaus), 170 
Trachycera Ragonot, 2, 25 

pallicornella (Ragonot), 25 
transilis Heinrich, Sigelgaita, 256 
transitella  (Dyar, not Walker), 

myelois, 45 

transitella (Walker), Paramyelois, 47 
translucida (Walker), Ufa, 171 
trichomata (Zeller), Piesmopoda, 78 
tricolorella Grote, Acrobasis, 13, 14 
triloses Dyar, Mescinia, 213 
trinitatis Heinrich, n. sp., Volatica, 290 
triplagiatella (Dyar), Pyla, 145 
truncatella (Wright), Lipographis, 166, 167 
Tucumania Dyar, 183, 252, 253 

porrecta Dyar, 252 

tapiacola Dyar, 252 
Tulsa Heinrich, n. gen., 5, 134 

finitella (Walker), 134, 135 

gillettella (Dyar), 135 

infinitella (Dyar), 135 

melanellus (Hulst), 134 

oregonella (Barnes & McDunnough), 

135 

umbripennis (Hulst), 135 
tumidella (Zincken), Acrobasis, 11, 17 
tumidulella (Ragonot), Acrobasis, 24 
turbatella (Grote), Zophodia, 238, 239 
turbidella Zeller, Crocidomera, 32, 33 
turpidella (Ragonot), Salebriaria, 115, 116 


Ecto- 


ubacensis (Zeller), Unadilla, 228 
Ufa Walker, 7, 169, 170 
deprivalis (Walker), 171 
lithosella (Ragonot), 170, 171 
luteella (Hulst), 170 
minualis (Walker), 171 
pyrrhochrellus (Ragonot), 171 
roseitinctella (Dyar), 171 
rubedinella (Zeller), 170, 171 
rufescentalis (Walker), 171 
senta Heinrich, n. sp., 170, 171 
translucida (Walker), 171 
venezuelalis Walker, 170, 171 
ulmella (Ragonot), Carnarsia, 202 
ulmiarrosorella (Clemens), Canarsia, 201, 202 
Ulophora Ragonot, 1, 2, 176 
dialithus (Dyar), 176, 177 
groteii Ragonot, 176, 177 
guarinella (Zeller), 177 
tephrosiella Dyar, 176 
umbrella (Dyar), Lipographis, 160, 166, 168 
umbripennis (Hulst), Tulsa, 135 
Unadilla Hulst, 1, 185, 218, 227, 282, 299 
albidiorella (Richards & Thomson), 
229 
bipunctella (Hampson), 228 
dorea (Dyar), 227, 228 


INDEX 


Unadilla Hulst—Continued 
erronella (Zeller), 228, 229 
floridensis Heinrich, n. sp., 229 
maturella (Zeller), 218, 228, 229 
masutella Hulst, 227, 228, 229 
pyllis (Dyar), 228 
ubacensis (Zeller), 228 
uncanale Hulst, Homoeosoma, 220 
uncanalis Ragonot (not Hulst), Homoe- 
osoma, 223 
uncta (Dyar), Erelieva, 308 
undulatella (Clemens), Hulstia, 169,195, 196 
unicolorella (Hulst), Oreana, 13, 112 
unicolorella Staudinger, Ephestia, 302 
uniformella Hampson, Ephestiodes, 281 
unionellum Ragonot, Homoeosoma, 225 
univitella Dyar, Anypsipyla, 42 
uvinella (Ragonot), Nephopteryx, 123, 130 


vaccinii Riley, Acrobasis, 13 
Vagobanta Heinrich, n. gen., 185, 289 
divergens (Butler), 289 
Valdivia Ragonot, 182, 191, 316 
aureomaculella (Dyar), 192 
coquimbella Ragonot, 191, 192 
lativittella (Ragonot), 191, 192 
walkerella (Ragonot), 192 
Varneria Dyar, 186, 305 
atrifasciella Barnes & McDunnough, 
305, 306 
dubia Heinrich, n. sp., 306 
nannodes Dyar, 305 
postremella Dyar, 305 
velessa Dyar, Harnocha, 202, 203 
venata Hampson, Nasutes, 315, 316 
venezuelalis Walker, Ufa, 170, 171 
venipars (Dyar), Paramyelois, 47 
venosa Dyar, Cacozophera, 208 
venosella Turati, Ephestia, 304 
vepallidum Heinrich, n. sp., Homoeosoma, 
224, 227 
vepreculella Ragonot, Ceracanthia, 86 
verecuntella (Grossbeck), Palatka, 208 
Verina Heinrich, n. gen., 184, 288, 289 
Verina supplicella (Dyar), 288 
vestilla (Dyar), Ephestiodes, 281, 309 
vetustella (Dyar), Nephopteryx, 127 
Vezina Heinrich, n. gen., 181, 291 
parasitaria Heinrich, n. sp., 291 
villosella Hulst, Etiella, 99 
virgatella (Clemens), Nephopteryx, 128 
viridisuffusella Barnes & MeDunnough, 
Pyla, 142, 149 
vitivora Filipjev, Ephestia, 302 
Vitula Ragonot, 180, 184, 279, 283, 285, 
287, 288, 289, 290, 291, 292, 313 
dentosella Ragonot, 285 
edmandsae (Packard), 285, 286, 287, 
288 
edmandsae serratilineella Ragonot, n. 
status, 285, 286 
inanimella (Dyar), 287 
laura (Dyar), 285, 287 
lugubrella (Ragonot), 286 
pinei Heinrich, n. sp., 286, 288 
ticitoa (Dyar), 287 


579 


vividella (McDunnough), Pima, 104 

Volatica Heinrich, n. gen., 181, 290, 291 
pachytaeniella (Ragonot), 290, 291 
trinitatis Heinrich, n. sp., 290 


walkerella (Ragonot), Valdivia, 192 

walsinghami (Ragonot), Ambesa, 108, 109 

Wunderia Grossbeck, 184, 204 
neaeriatella Grossbeck, 204, 205 


xanthaenobares Dyar, Dioryctria, 153 
xanthomera Dyar, Piesmopoda, 78 
xanthopolys Dyar, Piesmopoda, 79, 80 
xanthoudemia (Dyar), Hyalospila, 58 
zanthozona Dyar, Piesmopoda, 78 


yavapaella Dyar (race of 
(Zeller)), Eurythmia, 307 
ydda (Dyar), Bema, 217, 218 
yddiopsis (Dyar), Bema, 219 
Yosemitia Ragonot, 183, 187, 250 
didactica Dyar, 250, 251 
fieldiella (Dyar), 250, 251 
graciella (Hulst), 250, 251 
graciella (Hulst, not Hulst), 251 
longipennella (Hulst), 250, 251 
yuconella (Dyar) (race of termitalis 
(Hulst)), Nephopteryx, 130 
yumaella (Dyar), Salebriacus, 114 


hospitella 


zamacrella Dyar, Laetilia, 230, 233, 234, 
286 
Zamagiria Dyar, 8, 90, 93, 94 
australella (Hulst), 92, 171 
bumeliella (Barnes & McDunnough), 
92 
deia Dyar, 93 
dixolophella Dyar, 90, 91 
fraterna Heinrich, n. sp., 92, 93 
hospitabilis Dyar, 90, 91 
ipsetona Dyar, 93 
laidion (Zeller), 92 
masculinus Dyar, 91 
pogerythrus Dyar, 91 
striella Dyar, 93 
zeae (Fitch), Plodia, 299 
zelatella (Hulst), Acrobasis, 12 
zelleri, Ragonot, Acrobasis, 11 
zeteki Heinrich, n. sp., Ectomyelois, 46 
zimmermanni (Druce, not Grote), Promy- 
lea, 66 
zimmermani (Grote), Dioryctria, 149, 150, 
152, 154, 155, 156, 157 
zinekenella (Treitschke), Etiella, 99, 238 
zonulella (Ragonot), Myelopsis, 41 
Zophodia Hiibner, 181, 187, 197, 230, 233, 
237, 238, 240, 257, 272, 315 
bella Hulst, 239 
convolutella (Hiibner), 238, 239 
dilativitta Dyar, 239 
franconiella Hulst, 239 
grossularae (Riley), 239 
grossularialis Hiibner, 239 
grossulariella (Hiibner), 239 
thouna Dyar, 239 
magnificans Dyar, 239 
turbatella (Grote), 238, 239 


580 UNITED STATES NATIONAL’ MUSEUM BULLETIN 207 


Species unknown, misplaced or 
unrecognized 


“Anthropteryx” irichampa Dyar, 313 
“Hlasmopalpus” corrientellus Ragonot, 312 
“Hucampyla” putidella Schaus, 313 
“Wuzophera” came Dyar, 313 

climosa Dyar, 315 

conquistador, Dyar, 315 

daedalella Ragonot, 314 

gais Dyar, 313 

intextella (Zeller), 314 

mabes Dyar, 315 

postflavida Dyar, 312 

rinmea Dyar, 313 

tintilla Dyar, 315 
“Huzopherodes” megalopalis Hampson, 

314 

“Honora” dulciella Hulst, 197, 313 
“Hornigia” clitellatella Ragonot, 234, 314 
“Hypochalcia”’ cervinistrigalis Walker, 312 

hulstiella Ragonot, 312 
“Maricopa” albocostella Hulst, 192, 316 
“Megasis” indianella Dyar, 160, 315 
“Moodna” formulella Schaus, 313 

olivella Hampson, 284, 314 
“Myelois” atristrigella Ragonot, 315 

famula Zeller, 312 

grossipunctella Ragonot, 312 

infusella Zeller, 314 

restricteila Zeller, 312 
“Nephopteryx” fuscifrontella Zeller, 312 
“Phycitopsis” flavicornella Ragonot, 314 
“Psorosa”’ disticta (Zeller), 314 
“Salebria” nigricans Hulst, 312 
“Zophodia” brevistrigella Ragonot, 314 

epischnioides Hulst, 315 

subcanella (Zeller), 314 


Hosts 


Abies, 150 
Acacia farnesiana, 47 
Achras sapota, 93 
Achyranthus ramosissima, 190 
Aesculus glabra, 47 
Aleurocanthus, 11 
Alnus, 23 
Ambrosia, 279 
Amorpha, 126, 127 
californica, 126 
herbacea, 126 
Anaphalis margaritosa, 224 
Annona squamosa, 44 
Antennaria, 137 
Anthemis, 220 
Anthyllis, 102 
Apple, 12, 14, 47, 113, 209, 273, 275, 284, 
286 
Apricot, 14, 273 
Aster, 220, 226 
Astragalus, 100, 102, 103, 106 


Attalea funifera, 82 
piassabossu, 82 
Azalea, wild, 120 
Balsam Fir (see Abies), 151 
Bauhinia mexicana, 159 
variegata, 60 
Bean, 173 
black, 60 
Bell pepper, 308 
Betula, 23, 120, 145, 284 
Bidens, 214, 220 
Black-eyed Peas, 60, 96, 172 
Blueberry, 13, 135 
Brauneria, 220 
Bumelia microcarpa, 92 
Butternut, 16 
“Caimitillo,” 92 
Cajanus, 96, 100 
cajan, 60, 96 
Canavalia ensiformis, 60 
maritima, 60 
Carapa guianensis, 29 
Carica papayae, 74 
Carissa, grandiflora, 45 
Carnegiae gigantea, 260 
Carya, 20 
alba, 120 
Cassia alata, 26 
bicapsularis, 45, 61 
brasiliensis, 42 
corymbosa, 61 
grandis, 47 
meschata, 46 
nodosa, 26 
occidentalia, 60 
spp., 61 
tora, 26 
Ceanothus, 114 
Cedrela, 28 
Celtis, 131, 132 
Ceratonia siliqua, 44, 45 
Cercis occidentalis, 296 
Cereus validus, 259 
Chaenomeles japonica, 10 
Chamaecrista brachiata, 98 
fasciculata, 98 
robusta, 98 
Cherry 12, 273 
Catalina, 14 
wild, 279 
Chickpea, 96 
Chokecherry, 13 
Chrysanthemum, 220 
Cicer, 96 
Cirsium, 221 
spinosisimum, 226 
Citron, 10 
Coccidae, 231, 232, 235 
Coccolobis uvifera, 77 
Cocos coronata, 82 
nucifera, 82 
vagans, 82 
Colutea, 100 
Comptonia, 23 
Coreopsis, 220 


Corn, 10, 173, 278, 284 

Corylus, 21 

Coryphantha aggregata, 251 

Cotoneaster, 12, 25 

Cotton, 173, 215, 220, 273, 279, 284 

Cottonwood, 122 

Cowpea, 60, 173 

Crabgrass, 173 

Crabapple, 12 

Cranberry, 13 

Crataegus 12, 13, 133 

Crotolaria, 100, 106 

Cylindropuntia, 245, 246, 248, 258 

Cyperus exculentus, 173 

Dahlia, 220 

Daphne gnidium, 10 

Dates, 45, 47 

Denmoza, 246, 247 

Dipholis salicifolia, 94 

Dolichos, 96, 100 

Hehinocereus, 250, 260, 262 
pectinatus, 262 
polyacanthus, 251, 253 
Tigidissimus, 262 
viridiflorus, 251 

Echinopsis, 246, 247 

Elephantopus, 199 

Elm, 196, 202 

Eremocarpus setigerus, 170 

Erica, 145 

Eriophyllum ignotum, 137 

Erisbotyra japonica, 45, 93 

Hulychnia acida, 256 

Ferocactus wislizeni, 260, 261 

Fig, 45, 47, 286, 303 

Fir, 150 

Flax, 173 

Franseria bipinnatifida, 194 

Fungus or pineapple, 297 

Galls, of sawfly on willow, 129 
on chokecherry, 14 

Gaura parviflora, 279 

Ginkgo, 273 

Genipa americana, 47 

Gleditsia, 134 
triacanthos, 47 

Glycina, 100 

Grain, 301, 303 

Grapefruit, 47 

Grapes, 10 

Hackberry, 202 

Hasseanthus elongatus, 238 

Helianthus, 220 

Heliopsis, 220 

Hickory, 15, 16, 17, 18, 19, 20, 22, 112, 

120, 202 

Homalocephala texensis, 250, 251, 260 

Honeycomb, 285, 286 

Hymenaea courbil, 44 

Indigofera tinctoria, 265 

Indigofera verbasifolium, 265 

Inga, 218 

Tris, 284 

Jack pine, 151 

Japanese cane, 173 

Johnsongrass, 173 


INDEX 581 


Languncularia racemosa, 77 Orange, 10, 47, 220 Rose, 284 
Lathyrus, 103 Orobanche ludovicana, 237 Rudbeckia, 220 
maritima, 103 Ostrya, 16, 17 Salix, 123, 129 
Leaves, dried, 288 virginiana, 21 caprea, 145 
Lecanium sp. 232 Pacae, 42 Samanea samédn, 42 
Lettuce, 226 Palafoxia, 197 Scale (see Coccidae), 233, 234, 235, 236 
Limabeans, cultivated, 60, 90, 172 Peach, 47, 273, 284 Schinus molle, 201 
wild, 60 Peanut, 173 Sciacassia siamea, 26 
Linden, 273 Pear, 209, 273, 284 Seeds, 214, 284 
Liquidambar, 273 Peas, garden, 60 Sitilias caroliniana, 226 
styraciflua, 131 Pecan, 16, 17, 18, 19, 22, 273, 275 Sonchus asper, 226 
Liriodendron tulipifera, 274 Peniocereus greggii, 260 Sorghum, 173, 308 
Livistona chinensis, 45 Persimmon, 273 Sapindus drummondii, 47 
Loco-weed, 106, 195 Phaseolus, 60, 100 “Spineless cactus,” 246 
Locust, 124 Phaseolus lunatus, 60 Spruce, 150, 151 
black, 173 Pigeon pea, 60, 96 “Stems of leguminous tree,” 55 
Loquat, 284 Pine, 150, 152, 286 Strawberry, 173 
Lotus, 102 Pinus, 150, 151, 153, 154, 155, 156, 284 Sudangrass, 173 
Lupinus, 100 caribaea, 152 Sugar beet, 196 
Lythrum, 11 chichuahuana, 154 Sugarcane, 173 
Mammea americana, 45 coulteri, 156 Sunflower, 220 
Maple, 101 monophylla, 157, 286 Sweetgum, 131 
Maytenus phyllanthoides, 264 palustris, 158 Swietenia, 28 
Mealy bugs (see Coccidae), 232 ponderosa, 152, 153, 156, 157 Swordbean, 60 
Melanthera radiata, 214, 229 scopulorum, 156 Tachardiella argentina, 233 
Milo Maize, 173 taeda, 156 Tagetes, 220 
Mimusops emarginata, 93 Pisum, 100 Tamarack, 151 
Mistletoe, 179 Pithecolobium flexicaule,”47 Tamarindus indica, 45 
Mountain Ash, 273 Platypuntia, 240, 246, 254, 256 Taxodium distichum, 154 
Mulberry, 273 Plum, 12, 14, 109, 273 Tamarix, 11 
Myrica, 23 Poinciana gilliesi, 61 Tephrosia, 176 
Myrica cerifera, 23 Pomegranate, 10 Theobroma cacao, 45 
Neomamillaria, 251 Poplar, 273 Thurberia, 296 3 
Nolina, 295 Populus, 122, 123 Trichocereus, 245, 246,"247, 256 
Nolina parryi, 295 tremuloides, 123 Tobacco, 303 
Nuts, 17, 45, 301, 303, 304 Prune, 12, 14, 286 chiloensis, 256 
Nyssa sylvatica, 140 Prunus maritima, 209 Turnip, 173 
Oak, 15, 21, 117, 118, 275 virginiana melanoearpa, 109 Vaccinium, 133 
scrub, 24 Pseudotsuga, 150 myrtellus, 145 
Oiketicus kirbyi, 291 Pulchea odorata, 229 Vachellia insularis,"45 
Olive, 273 Puya alpestris, 289 Vegetable products,{dried, 284, 293, 299, 
Onion, 10 301, 303 


Pyracantha coccinea, 12 


Ononis, 102 Viburnum, 101 

Opuntia, 220, 254, 255, 257, 295, 308 I eee ld Vicina, 100 
(Cylindropuntia) exaltata, 245, 247 Quince, 12 Vigna, 100 
(Cylindropuntia) imbricata, 244, 248 | Raisin, 10, 45, 279, 286 unguiculata, 60 
(Cylindropuntia) leptocaulis, 245, 249, | Razamofskya eryptopoda, 69 Viguiere, 220 

250 5 7908 
‘ : Rhus, 124, 221, 284 Walnut, 16, 20, 22,7120,8273 

(Cyclindropuntia) sp., 258 toxicodendron, 166 black, 20 


(Platypuntia) aurantiaca, 252 


(Platypuntia) discolor, 252 Ribes, 239 f = ea 
(Platypuntia) fiscus-indica, 247, 256 grossularia, 239 eat, 173, 
(Platypuntia) sulphurea, 247, 254 Ricinus communis, 10 Wulffia, 214 
(Platypuntia) sp., 240, 241, 242, 243, | Robinia, 45, 47 Ximenesia, 220 
244, 246, 252, 257, 258 pseudoacacia, 128 Yucca, 47, 295 


U. S. GOVERNMENT PRINTING OFFICE: 1956 


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