' CHECKLIST OF AMERICAN PHYCITINAE
- Pages 317-329 from
United States National Museum Bulletin 207
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
By Carl Heinrich
Checklist of American Phycitinae
(Synonyms in italics)
1. CrypToBLaBEs Zeller
1.
gnidiella (Milliére): Europe, Africa, Asia,
Bermuda, Venezuela, Brazil
2. Acrosasis Zeller
Mineola Hulst
Seneca Hulst
Acrocaula Hulst
. indigenella (Zeller): Eastern U.S. and Can-
ada, California
nebulo (Walsh)
nebulella (Riley)
zelatella (Hulst)
3. grossbecki (Barnes and McDunnough), new
Doe
comb.: Florida
. vaccinii Riley: U.S.
. amplexella Ragonot: Eastern U.S.
. tricolorella Grote: U.S., Canada
scitulella Hulst
. comptella Ragonot: Western U.S.
. minimella Ragonot: LHastern U. S.
nigrosignella Hulst
. feltella Dyar: Eastern U. S., Canada
. palliolella Ragonot: Eastern U. S., Canada
albocapitella Hulst
. caryalbella Ely: U.S. (Connecticut)
. juglandis (LeBaron): Eastern U.S.
. sylviella Ely: Eastern U. S., Canada
. kearfottella Dyar: Eastern U. S.
. caryae Grote: Eastern U. S., Canada
. evanescentella Dyar: U.S. (Florida)
. stigmella Dyar: Eastern U.S.
. aurorella Ely: Eastern U. S.
. peplifera Dyar: Eastern U.S.
. exsulella (Zeller), new comb.: Eastern U.S.
septentrionella Dyar
. angusella Grote: Eastern U. S., Canada
eliella Dyar
. demotella Grote: Eastern U. S.
. latifasciella Dyar: Eastern U.S.
. irrubriella Ely: Eastern U. S.
. normella Dyar: Eastern U.S. (Connecticut)
. malipennella Dyar: Eastern U. S. (Con-
necticut)
. dyarella Ely: Eastern U. S. (Connecticut)
. ostryella Ely: Eastern U. S., Canada
. secundella Ely: Eastern U. S., Canada
. coryliella Dyar: Eastern U. S.
. hebescella Hulst: Eastern U. S. (New Jer-
sey)
. cirroferella Hulst: Eastern U. S. (Texas)
. cunulae Dyar and Heinrich: Eastern U. S.
. caryivorella Ragonot: Eastern and South-
western U.S.
35. comacornella (Hulst), new comb.: Eastern
U.S. (Texas)
36. betulella Hulst: Eastern and Western U.S.,
Canada
37. rubrifasciella Packard: Eastern U. S.,
Canada
alnella McDunnough
38. comptoniella Hulst: Eastern U. S., Canada
39. myricella Barnes and McDunnough: U. S.
(Florida)
40. tumidulella (Ragonot), new comb.: U. S.
(Florida)
3. RHopDOPHAEA Guénée
41. caliginella (Hulst), new comb.: U.S. (Cali-
fornia, Arizona)
caliginoidella (Dyar)
42. supposita (Heinrich), new comb.: Canada
(British Columbia)
. TRACHYCERA Ragonot
43. pallicornella (Ragonot): U.S. (Texas)
. Anazasis Heinrich, new genus
44. ochrodesma (Zeller), new comb.: U. S.,
(Florida), México, Guatemala, Panamé
Colombia, West Indies
crassisquamella (Hampson)
. Miuprrxra Dyar
45. constitutionella Dyar: México, Guatemala
. SEMATONEURA Ragonot
46. atrovenosella Ragonot: México, Costa Rica,
Colombia, Ecuador, Pert, Argentina
47. abitus Heinrich, new species: Ecuador
. Hypsipyta Ragonot
48. grandella (Zeller): U. S. (Florida), West
Indies and Tropical America to Argen-
tina
cnabella Dyar
49. ferrealis (Hampson), new comb.: ‘Tropical
America (Costa Rica to Brazil)
50. dorsimacula (Schaus), new comb.: Costa
Rica
51. fluviatella Schaus: Costa Rica
. Hemrpritocera Ragonot
52. chinographella Ragonot: French Guiana,
Brazil, Pert
53. bigrana (Zeller): México, Colombia
54. plumigerella (Ragonot), new comb.: ‘‘Amer.
Merid.”’
55. letharda (Schaus), new comb.: Panama,
México
56. jocarella (Schaus): Costa Rica, Panama,
Brazil
57. exoleta (Zeller): Colombia
317
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318
10.
11.
12.
13.
14.
15.
16.
17.
18.
19:
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Crocipomera Zeller
58. turbidella Zeller:
U.S. (Texas)
59. fissuralis (Walker):
Puerto Rico
adonea (Felder and Rogenhofer)
60. stenopteryx (Dyar), new comb.: México
Cuniperta Heinrich, new genus
61. subtinctella (Ragonot), new comb.:
ern U.S. and Canada
Horas Heinrich, new genus
62. disjunctus Heinrich, new species:
Apanarsa Heinrich, new genus
63. intransitella (Dyar), new comb.:
(Arizona, New Mexico)
Brrinus Heinrich, new genus
64. russeolus Heinrich, new species:
Guiana
Buertetia Barnes and McDunnough
65. grisella Barnes and McDunnough: U. S.
(Arizona)
Hypareynria Ragonot
Cuba, Jamaica, México,
Dominican Republic,
West-
Colombia
U.S.
British
66. definitella (Zeller): Puerto Rico, Virgin
Islands, Colombia, Brazil
67. slossonella (Hulst), new comb.: U. S.
(Florida), México
tenuella (Barnes and McDunnough)
Craranrica Heinrich, new genus
68. annuliferella (Dyar), new comb.: U. S.
(New Mexico, Arizona)
69. hystriculella (Hulst), new comb.: U. S.
(Texas, Florida)
70. bicolorella (Barnes and McDunnough), new
comb.: U.S. (Arizona, Nevada, Cali-
fornia)
Mye topsis Heinrich, new genus
71. coniella (Ragonot), new comb.:
ada, México
nefas (Dyar)
72. immundella (Hulst),
(Texas)
73. subtetricella (Ragonot), new comb.:
Canada
zonulella (Ragonot)
obnupsella (Hulst)
74. minutularia (Hulst),
(Texas)
75. alatella (Hulst), new comb.:
rectistrigella (Ragonot)
fragilella (Dyar)
piazzella (Dyar)
AnypsipyLa Dyar
76. univitella Dyar: Cuba, México, Guatemala,
Panam4, Venezuela, Brazil, Peri, Ecua-
dor, Jamaica
U.S., Can-
new comb.: U. S.
5s,
new comb.:
Western U.S.
20. ApomyEtots Heinrich, new genus
77. bistriatella (Hulst), new comb.: Eastern
U.S., Canada
bilineatella (Ragonot)
Uy Ss -
21.
22.
23.
24.
25.
26.
Ecromyetots Heinrich, new genus.
78. decolor (Zeller), new comb.: Tropical
America
ephestiella (Hampson)
79. ceratoniae (Zeller), new comb.: Europe,
U. S. (Florida), Puerto Rico, Jamaica,
Argentina
oporedestella (Dyar)
80. muriscis (Dyar), new comb.: Tropical
America
palpalis (Dyar)
81. furvidorsella (Ragonot), new comb.: Puerto
Rico
82. zeteki Heinrich, new species: Panam&é
ParAMYELO!Is Heinrich, new genus.
83. transitella (Walker), new comb.: U. &S.,
tropical America
notatalis (Walker)
solitella (Zeller)
duplipunctella (Ragonot)
venipars (Dyar)
cassiae (Dyar)
Psrupopivona Dyar
84. commensella Dyar: México
85. cispha Dyar: Guatemala, Costa Rica, Brit-
ish Honduras
86. santa-maria Dyar: Guatemala
87. carabayella Dyar: Pert, Bolivia, Colombia
ProToMOERBES Heinrich, new genus
88. aberrans Heinrich, new species:
89. separabilis Heinrich, new species:
Diatomocera Ragonot
Cabima Dyar
90. tenebricosa (Zeller):
ana, Costa Rica
91. dosia (Dyar), new comb.:
Colombia
Colombia
Colombia, French Gui-
Panama
92. excisalis (Hampson), new comb.: French
Guiana, Bolivia (?)
93. decurrens (Dyar), new comb.: Panam&
94. majuscula Heinrich, new species: Brazil
95. albosigno Heinrich, new species: Brazil
96. hoplidice (Dyar), new comb.: Panama
97. extracta Heinrich, new species: Costa Rica
98. mochlophleps (Dyar), new comb.: México
Psrupocasima Heinrich, new genus
99. castronalis Heinrich, new species: Brazil
100. fearnella (Schaus), new comb.: Costa Rica,
Guatemala
101. guianalis Heinrich, new species: French
Guiana, British Guiana
102. euzopherella (Dyar), new comb.: Panam&
103. pombra (Dyar), new comb.: Panama
104. nigristrigella (Ragonot), new comb.: Brazil
105. arizonensis Heinrich, new species: U. S.
(Arizona)
106. expunctrix (Dyar and Heinrich), new comb.:
Brazil
107. perrensiella (Ragonot, new comb.: Argen-
tina
Lae A
: + tie
27.
28.
29.
30.
31.
32.
AMERICAN MOTHS OF THE
108. rubrizonalis (Hampson), new comb.: French
Guiana, Brazil
Hyatosrita Ragonot
109. stictoneurella Ragonot: México, Guatemala,
Brazil
110. celiella Schaus: Costa Rica
111. insequens Heinrich, new species: Bolivia,
Colombia
112. majorina Heinrich, new species: México
113. fulgidula Heinrich, new species: Cuba
114. egenella (Ragonot), new comb.: Brazil
115. xanthoudemia (Dyar), new comb.: Panama,
Costa Rica
116. angulinella (Schaus), new comb.: Costa
Rica
117. clevelandella (Dyar): Panam4
118. semibrunneella Ragonot: Colombia
Funpewya Zeller
119. pellucens Zeller: U.S. (Florida), West In-
dies, Brazil, Bolivia
cistipennis (Dyar)
120. argentina Dyar: U. S. (Florida, Texas),
West Indies, Venezuela, Brazil, Argen-
tina
eucasis Dyar
121. agapella Schaus: Gal&pagos Islands
122. ignobilis Heinrich: México, Guatemala,
Costa Rica, Cuba, Puerto Rico, Haiti
123. ahemora Dyar: México, Guatemala, Costa
Rica
DirunpeEtua Dyar
124. corynophora Dyar:
French Guiana
125. subsutella (Schaus), new comb.:
Guatemala, Panamé,
Costa Rica
126. distractor Heinrich, new species: Puerto
Rico
127. tolerata Heinrich, new species: Bolivia
Coprarruria Ragonot
128. dasypyga (Zeller):
Promy.eEa Ragonot
129. lunigerella Ragonot: Western U. S. and
Canada.
130. lunigerella glendella (Dyar):
131. dyari Heinrich, new name:
zimmermani (Druce)
drucet (Dyar)
132. druceii (Ragonot), new comb.:
133. mindosis Dyar: México
134. dasystigma Dyar: México
ANADELOSEMIA Dyar
135. senesciella (Schaus): Costa Rica
136. tecmessella (Schaus): Costa Rica
137. fifria Dyar: Guatemala
138. base Dyar: Guatemala
139. obstitella (Schaus), new comb.: Costa Rica
140. texanella (Hulst), new comb.: U.S. (Texas,
Florida), Puerto Rico, Cuba
dulciella (Hulst)
condigna Heinrich, new species:
zona)
Colombia, Guatemala
Colorado
México
Guatemala
141. U.S. (Ari-
SUBFAMILY PHYCITINAE
33.
34.
35.
36.
37.
38.
39.
319
Dasypyea Ragonot
142. alternosquamella Ragonot:
Canada
stictophorella Ragonot
Rampyitua Dyar
143. orio Dyar: México
144. polydectella (Schaus):
145. subcaudata (Dyar), new comb.:
Costa Rica, Brazil
Western U. S.,
Costa Rica
Guatemala,
146. lophotalis Heinrich, new species: México,
Guatemala
Fuurapa Heinrich, new genus
147. querna (Dyar), new comb.: Panama
148. carpasella (Schaus), new comb.: Galipagos
Islands
Scoryuus Heinrich, new genus
149. cubensis Heinrich, new species: Cuba
Davara Walker
Homalopalpia Dyar
Eucardinia Dyar
caricae (Dyar), new comb.:
Tropical America
dalera (Dyar)
150. U. S. (Florida),
151. columnella (Zeller), new comb.: Colombia
152. nerthella (Schaus), new comb.: México,
Guatemala, Costa Rica
euthales (Dyar)
153. paranensis (Dyar), new comb.: Brazil
154. azonaxsalis Walker: Brazil
155. interjecta Heinrich, new species: Puerto
Rico, Dominican Republic
156. rufulella (Ragonot), new comb.: Puerto Rico
Sarasota Hulst
Cuba Dyar
157. plumigerella Hulst: U.S. (Florida)
158. furculella (Dyar), new comb.: Cuba, Puerto
Rico, Dominica, Virgin Islands
159. ptyonopoda (Hampson), new comb.:
ward Islands
Presmopopa Zeller
Discopalpia Ragonot
Amphycitopsis Dyar
160. rubicundella Zeller: Brazil
161. xanthomera Dyar: Guatemala,
Costa Rica, French Guiana
zanthozona Dyar
trichomata (Zeller): Colombia
flavicans (Zeller): Colombia, French Guiana
fratella Dyar
ragonoti (Dyar), new comb.:
temala, Costa Rica
isabella (Dyar), new comb.:
xanthopolys Dyar: Panama
parva Heinrich, new species: Panama
semirufella (Zeller): Colombia
apocerastes Dyar: México, Costa Rica,
French Guiana, Brazil, Dominica
montella Schaus: Costa Rica
Wind-
Panama
162.
163.
164. México, Gua-
165. Costa Rica
166.
167.
168.
169.
170.
+ Hi be
wf
ah
shiv
SDA,
320
UNITED STATES NATIONAL MUSEUM BULLETIN 207
40. AtHrtoca Heinrich, new genus
41.
42.
43.
44.
45.
46.
47.
48.
171. subrufella (Hulst), new comb.:
ida), Cuba, Virgin Islands
filiolella (Aulst). Virgin Islands
ptychis (Dyar)
U.S. (Flor-
172. bondari Heinrich, new species: Brazil
PRAEDONULA Heinrich, new genus
173. almonella (Dyar), new comb.: Panama
Peavus Heinrich, new genus
174. burdettellus (Schaus), new comb.: Costa
Rica, Guatemala
semproniella (Schaus)
175. dissitus Heinrich, new species: Brazil
176. subaquilellus (Ragonot), new comb.: Guate-
mala
Gasinius Heinrich, new genus
177. paulsoni (Ragonot), new comb.: Chile
CrracanTuHiA Ragonot
Procandiopa Dyar
178. mamella (Dyar), new comb.: Panama,
Guatemala
179. vepreculella Ragonot: Ecuador
Mecarruria Ragonot
180. peterseni (Zeller): Guatemala, Colombia,
Brazil, Pert
181. squamifera Heinrich, new species: Costa
Rica
182. frustrator Heinrich, new species: Costa Rica
183. schausi Heinrich, new species: Costa Rica
184. cervicalis Dyar: Cuba
185. alpha Heinrich, new species: Guatemala,
México, Costa Rica, Panamé, Bolivia,
Brazil
186. beta Heinrich, new species: México, Guate-
mala, Costa Rica, Trinidad
Drescoma Dyar
187. cyrdipsa Dyar: México, Guatemala, Pan-
ama, French Guiana
188. cinilixa Dyar: Guatemala, Panamé
Mowoptitora Hulst
189. pergratialis (Hulst): U.S.
grotella (Ragonot)
nubilella Hulst
Zamaetria Dyar
190. dixolophella Dyar: Panam&
191. pogerythrus Dyar: México, Guatemala
192. hospitabilis Dyar: Cuba
193. masculinus Dyar: Guatemala
194. australella (Hulst), new comb.: Was:
(Texas, Florida)
bumeliella (Barnes and McDunnough):
U.S. (Texas, Florida)
195. fraterna Heinrich, new species: Cuba
196. laidion (Zeller): U.S. (Florida), Tropical
America
deia Dyar
striella Dyar
197. ipsetona Dyar:
399671—56——2
Costa Rica
49.
ANEGCEPHALESIS Dyar
198. arctella (Ragonot), new comb.: U. S.
(Florida), Bahamas, Cuba
cathaeretes Dyar
50. Maatriopsts Heinrich, new genus
199. denticosella (Dyar), new comb.: ‘Tropical
51.
52.
53.
54.
55.
America
cristalis (Hampson)
AncyLostomi1a Ragonot
200. stercorea (Zeller):
America
ignobilis (Butler)
diffissella (Zeller)
201. sauciella (Zeller): Colombia
202. argyrophleps Dyar: México, Guatemala
203. euchroma Dyar: Brazil
Carisranius Heinrich, new genus
204. pellucidellus (Ragonot), new comb.: Puerto
Rico, St. Vincent, Jamaica, Surinam,
Brazil
melanoplaga (Hampson)
205. decoloralis (Walker), new comb.:
Us:
metagrammalis (Walker)
Surfurellus (Hulst)
floridellus (Hulst)
206. guatemalellus (Ragonot),
Guatemala
Ertexua Zeller
207. zinckenella (Treitschke):
U. S., Tropical America
etiella (Treitschke)
schisticolor Zeller
villosella Hulst
rubribasella Hulst
Guyptocera Ragonot
208. consobrinella (Zeller):
U.S. (Florida), Tropical
Southern
new comb.:
Europe, Asia,
Eastern U. S., Can-
ada,
busckella (Dyar)
Pima Hulst
209. boisduvaliella (Guénée), new comb.:
rope, Canada
albiplagiatella (Packard), new comb.: East-
ern U. S., Canada
Eu-
210.
211. albiplagiatella occidentalis Heinrich, new
race: Western U.S.
212. fosterella Hulst: Western U. S., Canada
213. vividella (McDunnough), new comb.: Can-
ada
214. albocostalialis (Hulst), new comb.: Western
U.S., Canada
215. albocostalialis subcostella (Ragonot), new
comb.: Southwestern U. S.
fulvirugella (Ragonot,) Newcomb.: Western
U.S. (California)
granitella (Ragonot), new comb.:
216.
PANE Western
piperella (Dyar)
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56.
57.
58.
59.
60.
61.
62.
63.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
218. parkerella (Schaus), new comb.: Western
U.S. (Montana)
InterseEctIo Heinrich, new genus
219. denticulella (Ragonot), new comb.: North-
western U. S., Canada
220. columbiella (McDunnough), new
Northwestern U. S., Canada
comb.:
221. ruderella (Ragnot), new comb.: ‘‘N.
Amer.” (California?)
222. niviella (Hulst) new comb.: U.S., Canada
AmpBeEsa Grote
223. laetella Grote: Western U. S., Canada
224, walsinghami (Ragonot): Western U. S.
monodon Dyar
walsinghami mirabella Dyar, new status:
U.S. (Southern California)
lallatalis (Hulst): . Western U. S. (Nevada,
Utah)
Carast1a Hiibner
225.
226.
227. bistriatella (Hulst), new comb.: Western
U.S. (California)
228. incorruscella (Hulst), new comb.: Western
U.S. (California)
229. actualis (Hulst), new comb.: Western U.S.,
Canada
Immyrua Dyar
230. nigrovittella Dyar:
Oreana Hulst
231. unicolorella (Hulst): Eastern U. S., Canada
leucophaeella (Hulst)
Oxysria Heinrich
Eastern U. S.
232. aliculella (Hulst), new comb.: Southwestern
U.S.
oberthuriella (Ragonot)
233. furciferella (Dyar) new comb.: Southwest-
ern U.S. (Arizona)
SaLesriacus Heinrich, new genus
234. odiosellus (Hulst), new comb.: Western
Weise
bakerella (Dyar)
yumaella (Dyar)
SaLEBrRiaRi4 Heinrich, new genus
235. turpidella (Ragonot), new comb.: Southern
WES:
ademptandella (Dyar)
nubiferella (Ragonot), new comb.:
engeli (Dyar) U.S.
annulosella (Ragonot), new comb.:
(Texas, North Carolina)
robustella (Dyar)
tenebrosella (Hulst), new comb.:
quercicolella (Ragonot)
heinrichalis (Dyar)
pumilella (Ragonot) new comb.:
ern U.S.
georgiella (Hulst)
fructetella (Hulst) new comb.:
rectistrigella (Dyar)
236.
237.
238.
U.S.
UPS:
239. Ves:
240. Southeast-
241. Uos;
64.
65.
66.
67.
68.
321
QUASISALEBRIA Henrich, new genus
242. admixta Heinrich, new species:
Uss:
OrtTHourPis Ragonot
243. jugosella Ragonot: Eastern U. S., Canada
244. pasadamia (Dyar), new comb.: U.S., Can-
ada
Potoprustis Ragonot
245. arctiella (Gibson):
Meroptera Grote
Emmerita Hampson
246. mirandella Ragonot:
247. cviatella Dyar:
248. pravella (Grote):
Western
Alaska, Canada
Western U. 8.
U.S. (illinois, Mississippi)
U.S., Canada
249. abditiva Heinrich, new species: Canada
NepHorrreryx Hiibner
250. subfuscella (Ragonot), new comb.: Eastern
U.S., Canada
semiobscurella (Hulst)
251. delassalis Hulst: Western U. S.
purpurella (Hulst)
pudibundella (Ragonot)
252. delassalis fraudifera Heinrich, new race:
Canada (British Columbia), U. S.(Wash-
ington)
253. rubescentella (Hulst): U.S.
254. fernaldi (Ragonot), new comb.: U. §&.,
Canada
255. dammersi Heinrich, new species: Western
U.S. (California, Arizona)
dammersi floridensis Heinrich, new race:
U.S. (Florida)
vetustella (Dyar), new comb.:
ada
256.
257. U. S., Can-
258. inconditella (Ragonot), new comb.: West-
ern U. S. (Arizona, Colorado)
259. subcaesiella (Clemens), new comb.: U. S.,
Canada
contatella (Grote)
260. virgatella (Clemens), new comb.: U. S.,
Canada ;
quinquepunctella (Grote)
261. carneella Hulst: U.S., Canada
inquilinella (Ragonot)
262. basilaris Zeller: U.S., Canada
263. termitalis (Hulst), new comb.: Western
U.S., Canada
levigatella (Hulst)
264. termitalis yuconella Dyar, new status:
Alaska
265. bifasciella Hulst: U.S. (Arizona)
nogalesella (Dyar)
266. uvinella (Ragonot), new comb.: Eastern
afflictella (Aulst)
liquidambarella (Dyar)
celtidella (Hulst), new comb.:
267. Uns:
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69.
70.
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72.
73.
74,
75.
76.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
268. rubrisparsella (Ragonot): U.S.
rufibasella (Ragonot)
croceella (Hulst)
texanella (Hulst)
269. gilvibasella Hulst:
lacteella (Hulst)
270. crassifasciella Ragonot:
decipientella Dyar
crataegella B. and McD.
271. bisra Dyar: México
Trascata Hulst
272. reductella (Walker): Eastern U.S.
gleditschiella (Fernald)
Tusa Heinrich, new genus
273. finitella (Walker), new comb.:
Canada
melanellus (Hulst)
274. umbripennis (Hulst), new comb.:
(Colorado)
gillettella (Dyar)
275. oregonella. (Barnes and McDunnough), new
comb.: U.S. (Oregon)
276. infinitella (Dyar), new comb.:
HomoroerapHa Ragonot
277. lanceolella Ragonot: Pert
TELETHusIA Heinrich, new genus
278. ovalis (Packard), new comb.:
latifasciatella (Packard)
geminipunctella (Ragonot)
modestella (Hulst)
279. rhypodella (Hulst), new comb.:
gon’’)
Puosus Heinrich, new genus
280. brucei (Hulst), new comb.:
281. funerellus (Dyar), new comb.:
Canada
U.S. (Texas)
Eastern U.S.
Eastern U.S.,
Uy: S:
México
U.S., Canada
U.S. (“Ore-
Western U. S.
Western U.S.,
282. curvatellus (Ragonot), new comb.: Western
283. incertus ‘Heinrich, new species: Western
U.S. (California)
Actrix Heinrich, new genus
284. nyssaecolella (Dyar), new comb.: Eastern
U.S.
285. dissimulatrix Heinrich, new species: Eastern
U.S. (Virginia)
Styitopautpr1a Hampson
West Indies, México
U.S. (Texas,
286. lunigerella Hampson:
287. scobiella (Grote), new comb.:
Colorado)
decimerella (Hulst)
288. argentinensis Heinrich, new species: Ar-
gentina
Pyza Grote
289. fasciolalis (Hulst), new comb.: Canada
(British Columbia)
290. impostor Heinrich, new species: Western
U.S., Canada
291. aequivoca Heinrich, new species: Western
Canada
399671—56—_3
ites
292. insinuatrix Heinrich, new species: Canada
(Manitoba)
aenigmatica Heinrich, new species: U. S.,
Canada
criddlella Dyar: Canada (Manitoba)
fusca (Haworth), new comb.: Holarctic
moestella (Walker)
frigidella (Packard)
cacabella (Hulst)
triplagiatella (Dyar)
hypochalciella (Ragonot), new comb.: North-
western U. S., Canada.
blackmorella (Dyar)
hanhamella Dyar: Canada (Manitoba)
scintillans (Grote): Western U. S. (Califor-
nia)
feella Dyar
293.
294.
295.
296.
297.
298.
299. sylphiella Dyar: Northwestern U.S., Canada
300. rainierella Dyar: Northwestern U.S. (Wash-
ington)
301. aeneella Hulst: Western U. S. (Colorado,
Utah)
302. aeneoviridella Ragonot: Western U. S.,
Canada
303. metalicella Hulst: Western U.S. (Colorado,
Utah)
304. fasciella Barnes and McDunnough: North-
western U. S. (California)
nigricula Heinrich, new species:
U.S. (Nevada)
viridisuffusella Barnes and McDunnough:
Western U.S. (California)
Diorycrria Zeller
Pinipestis Grote
307. abietella (Denis and Schiffermiiller):
ern Hemisphere
decuriella (Hisbner)
abietivorella (Grote)
elegantella (Hulst)
sysstratiotes Dyar:
reniculella (Grote):
ponderosae Dyar:
California)
majorella Dyar: México
muellerana Dyar
disclusa Heinrich: Eastern U.S.
auranticella (Grote): Western U.S., Canada
miniatella Ragonot
zanthaenobares Dyar
erythropasa (Dyar):
(Arizona)
315. horneana (Dyar):
305. Western
306.
North-
Guatemala
Northern U.S., Canada
Western U.S. (Montana,
308.
309.
310.
dll.
312.
313.
314. Southwestern U, S.
Cuba
316. pygmaeella Ragonot: Eastern U.S.
317. zimmermani (Grote): U.S., Canada
delectella (Hulst)
austriana (Cosens)
318. cambiicola (Dyar): Western U. S.
319. amatella (Hulst): Eastern U. S.
320. albovittella (Hulst): Western U.S.
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POG Tea at
miler hn:
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78.
a9.
80.
81.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
321. gulosella (Hulst), new comb.: Western U.S.
(Colorado, New Mexico)
322. baumhoferi Heinrich, new species: South-
western U.S. (Arizona)
323. subtracta Heinrich, new species: South-
western U. S. (New Mexico)
324. clarioralis (Walker): Eastern U. S.
brunneella (Dyar)
OrycToMEToPiIA Ragonot
325. fossulatella Ragonot:
ical America
moeschleri (Ragonot)
SaraTa Ragonot
U. S. (Texas), Trop-
326. edwardsialis (Hulst), new comb.: Western
Wass
polyphemella (Ragonot)
327. Western
pullatella (Ragonot), new comb.:
USS
328. punctella (Dyar), new comb.: México
329. punctella septentrionaria Heinrich, new race:
Western U.S.
330. incanella (Hulst), new comb.: Western U.S.
aridella (Dyar)
331. atrella (Hulst), new comb.: Western U. S.
(Colorado)
332. caudellella (Dyar), new comb.: Western
U.S., Canada
333. dnopherella Ragonot: Western U.S. (Cali-
fornia)
334. nigrifasciella Ragonot: Western U.S., Can-
ada
335. cinereella Hulst: Western U. S. (Colorado)
336. rubrithoracella (Barnes and McDunnough),
new comb.: Western U.S.
337. tephrella Ragonot: Western U. S. (Wash-
ington)
338. alpha Heinrich, new species: Canada (Sas-
katchewan)
339. beta Heinrich, new species: Western U. S.,
Canada
340. gamma Heinrich, new species: Western
U.S. (California)
341. iota Heinrich, new species: Western U. S.
(California)
342. perfuscalis (Hulst): Western U. S.
excantalis (Hulst)
343. epsilon Heinrich, new species: Western U.S.
344. phi Heinrich, new species: Western U.S.
345. kappa Heinrich, new species: Western U.S.
(Arizona)
346. delta Heinrich, new species: Western U. S.
PuitopEMa Heinrich, new genus
347. rhoiella (Dyar), new comb.: Western U. S.
Lipograpuis Ragonot
348. fenestrella (Packard):
fornia)
humilis Ragonot
349. leoninella (Packard):
Western U.S. (Cali-
Western U. S., Can-
ada
pallidella (Dyar)
82.
83.
84.
85.
86.
87.
88.
323
350. truncatella (Wright), new comb.: South-
western U. S. (California)
351. umbrella (Dyar), new comb.:
U.S. (California)
352. subosseella Hulst: Bahamas
Apvr.puta Heinrich, new genus
353. petrella (Zeller), new comb.:
rubiginella (Walker)
rufinalis (Walker)
hapsella (Hulst)
354. ochripunctella (Dyar), new comb.:
U.S. (California)
Tora Heinrich, new genus
355. galdinella (Schaus), new comb.:
Islands
Ura Walker
356. lithosella (Ragonot), new comb.:
western U. S., México
luteella Hulst)
roseitinctella (Dyar), new comb.:
western U. S., México
senta Heinrich, new species:
U.S. (Texas, Arizona)
rubedinella (Zeller), new comb.:
ida), Tropical America
translucida (Walker)
rufescentalis (Walker)
minualis (Walker)
deprivalis (Walker)
venezuelalis Walker
pyrrhochrellus (Ragonot)
Easmopatpus Blanchard
360. lignosellus (Zeller): U.S., Tropical America
angustellus Blanchard
tartarella (Zeller)
incautella (Zeller)
major (Zeller)
anthracellus Ragonot
carbonella (Hulst)
puer Dyar
Acroncosa Barnes and McDunnough
361. albiflavella Barnes and McDunnough: West-
ern U. S. (California)
362. albiflavella castrella Barnes and McDun-
nough: Western U.S. (New Mexico)
363. similella Barnes and McDunnough: West-
ern U.S. (Nevada, Utah)
PassaDENA Hulst
364. flavidorsella (Ragonot):
México
canescentella (Hulst)
constantella Hulst
cinctella (Hulst)
UxorHora Ragonot
Acromeseres Dyar
365. groteii Ragonot: Eastern U. S.
tephrosiella Dyar
366. guarinella (Zeller):
dialithus (Dyar)
Southwestern
Wes:
Western
Galipagos
South-
BY lc South-
358. Southwestern
359. U.S. (Flor-
Western U. S.,
Cuba, Colombia
aly
ee
5
'
Dia be
1:
4
Nia, Oy wig k
4 ee “
‘
324
89.
90.
91.
92.
93.
94.
95.
96.
oT.
98.
os
UNITED STATES NATIONAL MUSEUM
Cuorrera Dyar
367. idiotes Dyar: Panamé4
368. extrincica (Dyar), new comb.:
369. postica (Zeller), new comb.:
Tacoma Hulst
370. feriella Hulst: Southwestern U.S.
submedianella Dyar
ApELPERGA Heinrich, new genus
371. cordubensiella (Ragonot), new comb.:
gentina
Evumysia Dyar
372. mysiella (Dyar): Western U. S.
373. maidella (Dyar): Western U. S., Canada
374. pallidipennella (Hulst), new comb.: West-
ern U.S
fuscatella (Hulst):
fornia)
376. semicana Heinrich, new species:
U.S. (Washington)
Drvitraca Barnes and McDunnough
377. ochrella Barnes and McDunnough: South-
ern U.S. (Florida)
378. simulella Barnes and McDunnough: South-
ern U. S. (Florida)
379. parvulella Barnes and McDunnough: South-
ern U.S. (Florida)
380. parvulella consociata Heinrich, new race:
Colombia
Macrorrainia Ragonot
Dolichorrhinia Ragonot
381. aureofasciella Ragonot: Southwestern U.S.,
México
382. placidella (Zeller):
Ocata Hulst
383. dryadella Hulst: Southern U.S. (Florida)
platanella (Grossbeck)
Vaupivia Ragonot
Maricopa Hulst
384. coquimbella Ragonot:
385. lativittella (Ragonot):
México
aureomaculella (Dyar)
386. walkerella (Ragonot), new comb.:
ProrasiA Heinrich, new genus
387. mirabilicornella (Dyar), new comb.:
ern U.S. (California)
Hererocrapuis Ragonot
Mona Hulst
388. morrisonella Ragonot:
coloradensis Ragonot
olbiella (Hulst)
ignistrigella Ragonot
palloricostella (Walter)
STaupincerIA Ragonot
389. albipenella (Hulst):
olivacella Dyar
perluteella Dyar
Cuba
Colombia
Ar-
Sioe Western U. S. (Cali-
Western
Brazil
Chile
Southwestern U. S.,
Chile
West-
U.S., México
Western U.S.
100.
101.
102.
103.
104.
105.
106.
107.
108.
109.
110.
Lae
112.
BULLETIN 207
Houustia Ragonot
390. undulatella (Clemens):
rubiginalis (Walker)
obsipella (Hulst)
fumosella (Hulst)
Honora Grote
391. mellinella Grote: U.S.
ochrimaculella Ragonot
subsciurella Ragonot: Western U. §,
sciurella Ragonot: Western U. S.
fornia)
dotella Dyar: Western U. S. (California)
montinatatella (Hulst): Western U. S.
canicostella Ragonot
perdubiella (Dyar), new comb.: Western
Was:
U.S., Canada
392.
393. (Cali-
394,
395.
396.
Hownorinus Heinrich, new genus
397. fuliginosus Heinrich, new species:
Oncoxasis Zeller
Endommasis Hampson
398. anticella Zeller: Tropical America
nigritella (Hampson)
Casotia Ragonot
Encystia Hampson
semidiscella Ragonot: Argentina
schini (Berg): Argentina
rhythmatica Dyar: Panamé
cundajensis ‘(Zeller): Colombia
impeditella (Zeller)
bonhoti (Hampson), new comb.:
Jamaica
Canarsta Hulst
404. ulmiarrosorella (Clemens):
pneumatella (Hulst)
ulmella (Ragonot)
fuscatella (Hulst)
gracilella Hulst
feliculella Dyar
Harnocua Dyar
405. velessa Dyar:
EKurytamasis Dyar
406. ignifatua Dyar:
Cuba
Evurytamipia Ragonot
407. ignidorsella (Ragonot):
México, Panamé
Wonperra Grossbeck
408. neaeriatella Grossbeck: U.S. (Florida)
Oxrpotumia Hampson
Synothmia Hampson
409. endopyrella Hampson: México, Bahamas
bahamasella (Hampson)
Sryntosasts Hampson
410. rubripurpurea Hampson:
Drv1ana Ragonot
Dannemora Hulst
411. eudoreella Ragonot:
edentella (Hulst)
Pert
399.
400.
401.
402.
403. Bahamas,
U.S., Canada
Panamaé
Panamdé, Puerto Rico,
U. S. (Arizona),
México, Brazil
Eastern U. 8S.
A
rf
PROP hay |
113.
114,
115.
116.
ave
118.
119.
120.
121.
122.
123.
124.
125.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
PaxuatKa Hulst
412. nymphaeella (Hulst): Eastern U. S.
verecuntella (Grossbeck)
CacozopHEra Dyar
413. venosa Dyar: Guatemala
Psorosina Dyar
414. hammondi (Riley): Eastern and Central
U.S., Canada
angulella Dyar
Parriciota Heinrich, new genus
415. semicana Heinrich, new species:
Pacontus Heinrich, new genus
416. corniculatus Heinrich, new species:
Rico
AptunGa Heinrich, new genus
417. macropasa (Dyar), new comb.:
México
418. imperfecta (Dyar), new comb.:
ANDERIDA Heinrich, new genus
419. sonorella (Ragonot), new comb.:
U.S. (Arizona)
placidella (Dyar)
Cassrana Heinrich, new genus
420. malacella (Dyar), new comb.:
Puerto Rico, Virgin Islands
Mescrinia Ragonot
421. triloses Dyar:
mosces Dyar
pandessa Dyar: Guatemala
bacerella Dyar: Cuba
estrella Barnes and McDunnough: U. S.
(Florida)
moorei Heinrich, new species: British Guiana
parvula (Zeller): Colombia
commatella (Zeller): Colombia
berosa Dyar: Panam4, Puerto Rico
peruella Schaus: Peri
430. discella Hampson: México, Guatemala
431. indecora Dyar: México
Nonta Ragonot
Hypermescinia Dyar
432. exiguella (Ragonot):
lambella (Dyar)
Puestinia Hampson
433. costella Hampson:
Comotia Dyar
434. torsicornis Dyar: Panamé
435. convergens (Dyar), new comb.:
Bema Dyar
Relmis Dyar
436. neuricella (Zeller), new comb.:
America
myja Dyar
437. fritilla Dyar: Guatemala
438. ydda (Dyar), new comb.: Panamé, French
Guiana
439. yddiopsis (Dyar), new comb.: Cuba
440. fifaca (Dyar), new comb.: Panama
Utah
Puerto
Guatemala,
Guatemala
México,
México,
Panamé
422.
423.
424,
425.
426.
427.
428.
429.
-
Tropical America
Jamaica, Puerto Rico
Guatemala
Tropical
126.
127.
128.
129.
130.
325
Homorosoma Curtis
Phycidea Zeller
441. electellum (Hulst): U.S., México, Guate-
mala, Cuba, British West Indies
opalescellum (Hulst)
texanellum Ragonot
tenuipunctella Ragonot
differtella Barnes and McDunnough
442. stypticellum Grote: U.S., Canada
uncanale Hulst
443. striatellum Dyar: Southwestern U. S.
444, oslarellum Dyar: Western U.S.
445. oslarellum breviplicitum Heinrich, new race:
Southwestern U.S. (California)
446. illuviellum Ragonot: U.S. (Arizona, Colo-
rado), México
candidella Hulst
447, illuviellum emendator Heinrich, new race:
Western U.S.
448. imitator Heinrich, new species: Southwestern
U.S. (California)
449. longiventrellum Ragonot: Chile
noctividella Ragonot
450. albescentellum Ragonot: Western U.S.
elongellum Dyar
451. impressale Hulst: Western U. S., Canada
452. inornatellum (Hulst): Eastern U.S.
453. deceptorium Heinrich, new species: U. S.
(Pennsylvania), Canada
454. discrebile Heinrich, new species, Brazil
455. peregrinum Heinrich, new species: U. S.
(California), Costa Rica
456. vepallidum Heinrich, new species: Argentina
457. ditaeniatellum Ragonot: Chile
458. oconequensis (Dyar), new comb.: Pert
459. assitum Heinrich, new species: Pert
460. acmaeopterum Ragonot: Chile
461. nimbosellum Ragonot: Chile
462. unionellum Ragonot: México
Paraconia Ragonot
463. magellanella (Ragonot): Chile
Rorrupa Heinrich, new genus
464. mucidella mucidella (Ragonot), new comb.:
Western U.S., Canada
465. mucidella reliquella (Dyar), new comb.:
Eastern U. S., Canada
466. mucidella olivaceela (Ragonot), new comb.:
Tropical America
musiosum (Dyar)
cubella (Dyar)
467. mucidella affusella (Ragonot), new comb.:
Argentina
STREPHOMESCINIA Dyar
468. schausella Dyar:
Unapiuia Hulst
Strymaz Dyar
Cuba
Siti ress
Bias ts
te
ahs )
¢
Cf a aes estan
326
131.
132.
133.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
469. erronella (Zeller): Tropical America.
ubacensis (Zeller)
bipunctella (Hampson)
dorae (Dyar)
pyllis (Dyar)
470. maturella (Zeller): Colombia, Guatemala,
Cuba
471. albidiorella (Richards and Thomason); new
comb.: Pert
472. floridensis Heinrich, new species: U.S. (Flor-
ida)
473. nasutella Hulst: U.S. (New Mexico)
Laetitia Ragonot
Laosticha Hulst
474. coccidivora (Comstock):
pallida (Comstock)
dilatifasciella (Ragonot)
hulstiit Cockerell
U_S.
475. coccidivora quadricolorella (Dyar), new
comb.: Southwestern U. S.
476. coccidivora cardini Dyar: Cuba, U. S.
(Florida)
477. obscura Dyar: Cuba
478. portoricensis Dyar: Puerto Rico
479. melanostathma (Meyrick), new comb.: Ar-
gentina
480. amphimetra (Meyrick), new comb.: Argen-
tina
481. zamacrella Dyar: Western U.S. (California)
482. myersella Dyar: Hastern U.S.
483. ephestiella (Ragonot): Southwestern U. S.
(Arizona)
lustrella (Dyar)
484. fiskella Dyar: Eastern U. S. (North Caro-
lina)
485. glomis (Dyar), new comb.: Panamé
Bapnata Heinrich, new genus
486. basimaculatella (Ragonot), mew comb.:
Western U. S.
eremiella (Dyar)
487. goyensis (Ragonot), new comb.:
Uruguay, Argentina
488. goyensis olivacea Heinrich, new race: Argen-
tina
489. homoeosomella (Zeller), new comb.:
ical America.
bodkini (Dyar)
rusto (Dyar)
taboga (Dyar)
saissetiae (Dyar)
490. haywardi Heinrich, new species: Argentina
491. glabrella (Dyar), new comb.: Guatemala
492. squalida (Walker), new comb.: Brazil
Raacea Heinrich, new genus
493. packardella (Ragonot), new comb.:
ern U.S.
orobanchella (Dyar)
494. stigmella (Dyar), new comb.:
U.S. (California), México
maculicula (Dyar)
Brazil,
Trop-
West-
Southwestern
134.
135.
136.
137.
138.
139.
140.
141.
142.
143.
Zopuopia Hiibner
Dakruma Grote
495. convolutella (Hiibner):
ada
grossulariella (Hiibner)
turbatella (Grote)
grossulariae (Riley)
franconiella (Hulst)
bella Hulst
ihouna Dyar
dilativitta Dyar
magnificans Dyar
Metitara Walker
Europe, U.S., Can-
496. prodenialis Walker: U.S.
bolliz (Zeller)
497. dentata (Grote): U.S.
doddalis Dyar
Otycetia Dyar
498. junctolineella (Hulst): Southern U.S. (Texas)
499. junctolineella pectinatella (Hampson): Méx-
ico
500. nephelepasa (Dyar:) México
501. subumbrella Dyar: Western U.S.
Otyca Walker
502. phryganoides Walker:
lic, Haiti
ALBERADA Heinrich
503. parabates (Dyar):
504. bidentella (Dyar):
(Texas, Arizona)
505. holochlora (Dyar):
(Texas)
Nanaia Heinrich
506. substituta Heinrich: Pert
Cactostastis Ragonot
Neopyralis Bréthes
Dominican Repub-
U.S., México
Southwestern U. S.
Southwestern U. S.
507. cactorum (Berg): Argentina, Uruguay, Aus-
tralia
508. ronnai (Bréthes): Brazil
509. doddi Heinrich: Argentina
510. mundelli Heinrich: Pert
511. bucyrus Dyar:
CauEta Heinrich
512. ponderosella (Barnes and McDunnough):
Western U.S., México
purgatoria (Dyar)
interstitialis (Dyar)
phoenicis (Dyar)
Rumatua Heinrich
513. glaucatella (Hulst): Southern U. S.
514. bihinda (Dyar): Western U.S.
515. polingella (Dyar): Southwestern U.S. (Ari-
zona, Texas)
Yosremit1a Ragonot
516. graciella (Hulst): Western U.S.
517. longipennella (Hulst): Southwestern U. S.
(Texas)
518. fieldiella (Dyar):
Arizona)
519. didactica Dyar:
Argentina
Western U.S. (California,
México
144.
145.
146.
147.
148.
149.
150.
151.
154.
155.
156.
157.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Tucumania Dyar
520. tapiacola Dyar:
521. porrecta Dyar:
EREMBERGA Heinrich
522. leuconips (Dyar):
523. creabates (Dyar):
fornia)
524. insignis Heinrich: México
SALAMBONA Heinrich
525. analamprella (Dyar):
Parotyca Dyar
526. asthenosoma (Dyar):
Sieeueaira Heinrich
527. chilensis Heinrich: Chile
528. huanucensis Heinrich: Pert
529. transilis Heinrich: Pert
AMALAFRIDA Heinrich
530. leithella (Dyar):
Colombia
Ozamia Ragonot
531. lucidalis (Walker): West Indies
532. fuscomaculella (Wright): Southwestern
U.S. (California)
heliophila Dyar
fuscomaculella clarefacta Dyar: Uz. S.
(Texas), México
thalassophila Dyar:
immorella (Dyar), new comb.:
stigmaferella Dyar: Argentina
537. hemilutella Dyar: Argentina
538. punicans Heinrich: Argentina
Cactosrosis Dyar
539. fernaldialis (Hulst): Southwestern U. S.
gigantella (Ragonot)
cinerella (Hulst)
longipennella (Hampson):
elongatella (Hampson):
maculifera Dyar: México ©
insignatella Dyar: México
strigalis (Barnes and McDunnough): West-
ern U. S., México
Argentina
Uruguay
Western U. S. (Arizona)
Western U. S. (Cali-
Argentina
French Guiana
West Indies, Venezuela,
533.
534.
535.
536.
U.S. (California)
México
540. México
541.
542.
543.
. Drescomopsis Dyar
544. soraella (Druce):
drucella (Dyar)
subelisa Dyar
Tropical America
. Intatiza Dyar
545. gurbyris Dyar: Panamé
Lascetina Heinrich, new genus
546. canens Heinrich, new species:
U.S. (Texas), México
Metepuestia Ragonot
547. simplicula (Zeller): U.S. (Florida), Puerto
Rico, Colombia, British West Indies
Sretaa Heinrich, new genus
548. arizonella (Hulst), new comb.:
ern U.S. (Arizona)
Entmemacornis Dyar
549. proselytes Dyar: Guatemala
550. pulla Heinrich, new species: Brazil
Southern
Southwest-
158.
159.
160.
161.
162.
163.
164.
165.
166.
167.
168.
169.
170.
327
Cayrnni1a Hampson
551. rufitinctalis Hampson:
Riosa, Heinrich, new genus
French Guiana
552. nexa Heinrich, new species: Argentina
Moerrszgs Dyar
553. dryopella (Schaus): Costa Rica
554. alveolella (Ragonot), new comb.: Brazil
555. emendata Heinrich, new species: Panamé,
French Guiana
Moopnopsis Dyar
Campyloplesis Dyar
556. decipiens Dyar: México
557. perangusta (Dyar), new comb.: Trinidad
558. inornatella (Ragonot), new comb.: Costa
Rica, Brazil
559. parallela Heinrich, new species: Brazil, Pera
560. inveterella (Dyar), new comb.: Guatemala
561. portoricensis Heinrich, new species: Puerto
Rico
Eputica Ragonot
562. compedella (Zeller):
KuzorHera Zeller
Tropical America
563. semifuneralis (Walker): U. S., Canada,
México
aglaeella Ragonot
pallulella (Hulst)
564. ostricolorella Hulst: Eastern U. 8S.
565. nigricantella Ragonot: Southwestern U. S.,
México
griselda Dyar
Exvurprrius Heinrich, new genus
566. negator Heinrich, new species:
Evtoetra Heinrich, new genus
Pert
567. ochrifrontella (Zeller), new comb.: U. S.,
Canada
ferruginella (Ragonot)
ProsonuzopHeErRA Heinrich, new genus
568. impletella (Zeller), new comb.: Colombia,
Jamaica, Puerto Rico
Farnosia Heinrich, new genus
569. quadripuncta (Zeller), new comb.: Costa
Rica, Panam4, French Guiana, Colombia
Gennavius Heinrich, new genus
570. junctor Heinrich, new species:
ana
Micromescinia Dyar
571. pygmaea Dyar:
Epuestiopgs Ragonot
572. gilvescentella Ragonot:
ada, México
nigrella Hulst
infimella Ragonot:
erythrella Ragonot:
coloradella (Hulst)
benjaminella Dyar
mignonella Dyar: U.S. (Texas)
erasa Heinrich, new species: U.S. (Florida)
lucidibasella Ragonot: Chile
productella Ragonot: Colombia (?)
indentella Dyar: Bermuda
French Gui-
Panamaé
Western U.S., Can-
Eastern U. S.
Western U.S., Canada
573.
574.
575.
576.
577.
578.
579.
Hea
PRS
ants
TD ine
i De apdbritsy ~
FO hae a
ra A
328
171.
172.
173.
174.
175.
176.
177.
178.
179.
180.
UNITED STATES NATIONAL MUSEUM BULLETIN
580. plorella Dyar: Panam&
vestilla (Dyar)
581. stictella (Hampson), new comb.:
West Indies
uniformella Hampson
granulella Hampson
582. noniella Dyar: Panama
Azazura Schaus
Calamophleps Dyar
583. muciella Schaus: Costa Rica, Guatemala,
Panama
squalidella (Dyar)
584. nodoses (Dyar), new comb.:
Bahamas,
Panama
585. lophophera (Dyar), new comb.: Panama
Moopna Hulst
586. ostrinella (Clemens): U.S., Canada
obtusangulella (Ragonot)
pelviculella Hulst
587. bisinuella Hampson:
Vituta Ragonot
588. edmandsae (Packard): Eastern U. S., Can-
ada
dentosella Ragonot
México, U.S. (Texas)
589. edmandsae serratilineella Ragonot, new sta-
tus: Western U.S., Canada
590. lugubrella Ragonot, new comb.: Western
U.S. (California)
591. pinei Heinrich, new species: Western U. S.
(Utah, Nevada)
592. inanimella (Dyar), new comb.: México,
Guatemala
ticitoa (Dyar)
593. laura (Dyar), new comb.: Guatemala
Manuatta Hulst
Hornigia Ragonot
594. setonella (McDunnough), new comb.: U.S.
(Utah), Canada (British Columbia)
595. broweri Heinrich, new species: Eastern U.S.
(Maine)
Verna Heinrich, new genus
596. supplicella (Dyar), new comb.: México,
Guatemala, Panam4, Brazil
Vacozpanta Heinrich, new genus
597. divergens (Butler), new comb.: Chile
Moopnetua Heinrich, new genus
598. paula Heinrich, new species: Guatemala,
Brazil, Argentina
Vouatica Heinrich, new genus
599. pachytaeniella (Ragonot), new comb.: Bra-
zil
600. trinitatis Heinrich, new species: Trinidad
Vezina Heinrich, new genus
601. parasitaria Heinrich, new species:
tina, Brazil
Caupgtuia Dyar
602. apyrella Dyar: Eastern U. S. (Maryland)
603. albovittella Dyar: Eastern U. S.
604. nigrella (Hulst), new comb.: Western U.S.
arizonella (Walter)
Argen-
181.
182.
183.
184.
185.
186.
187.
207
605. declivella (Zeller), new comb.: Panama,
Colombia
animosella (Dyar)
606. colorella (Dyar), new comb.: Panama
607. clara Heinrich, new species: Puerto Rico
Micropuestia Dyar
608. animalcula Dyar: Panam&
Sosrpatra Heinrich, new genus
609. rileyella (Ragonot), new comb.: Western
U.S., México
micaceella (Hampson): México
anthophila (Dyar), new comb.:
U.S. (Texas)
610.
611. Western
612. thurberiae (Dyar), new comb.: Western U.S.
613. nonparilella (Dyar), new comb.: Western
U.S. (Arizona)
614. majorella (Dyar), new comb.: México
615. divergens (Dyar): Panamé
Berruuria Ragonot
616. championella Ragonot: Guatemala
Risva Heinrich
617. innoxia Heinrich: Cuba
618. contigua Heinrich, new species:
619. patriciella (Dyar), new comb.:
Propra Guénée
620. interpunctella (Hiibner):
interpunctalis (Hiibner)
zeae (Fitch)
latercula (Hampson)
glycinwora (Matsumura)
621. dolorosa Dyar: Guatemala
AnaGastA Heinrich, new genus
622. kiihniella (Zeller): Cosmopolitan
tuscofasciella (Ragonot)
gitonella Druce
Epuestia Guénée
Hyphantidium Scott
623. elutella (Hiibner): Cosmopolitan
elutea (Haworth)
semirufa (Haworth)
rufa (Haworth)
sericarium (Scott)
roxburghit Gregson
unicolorella Staudinger
amarella Dyar
624. cautella (Walker):
defectella (Walker)
desuetella (Walker)
cahiritella Zeller
passulella Barrett
formosella (Wileman and South)
625. figulilella Gregson: Europe, Asia, Africa,
Hawaii, Australia, North America (U.S.,
California), South America
~ ficulella Barrett
milleri Zeller
figuliella Forbes
Jigulella Curran
venosella 'Turati
ernestinella Turati
Puerto Rico
Cuba
Cosmopolitan
Cosmopolitan
ne Leer
Res Se,
Rey Pts
eer inna
PDE EU
i an
aD AAC on
188.
189.
190.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Nicrtioprs Schaus
626. apianella Schaus:
VaRNERIA Dyar
Galapagos Islands
627. postremella Dyar: Eastern U.S.
628. nannodes Dyar: Panama
629. atrifasciella Barnes and McDunnough:
Southern U. S. (Florida)
630. dubia Heinrich, new species:
Evrytumia Ragonot
631. hospitella (Zeller):
U.S
Puerto Rico
Southern and Western
spaldingella Dyar
632. hospitella yavapaella Dyar, new status:
Western U. S.
633. angulella Ely: Eastern U. S., Canada
diffusella Ely
634. fumella Ely: Eastern U.S. (Connecticut)
191.
192.
193.
194.
329
Ere.ieva Heinrich, new genus
635. quantulella (Hulst), new comb.:
U.S. (Texas), West Indies
santiagella (Dyar)
636. coca (Dyar), new comb.:
coquilla (Dyar)
mossa (Dyar)
uncta (Dyar)
637. parvulella (Ely), new comb.:
(Connecticut)
Casnia Dyar
638. myronella Dyar:
Microprycita Dyar
639. titillella Dyar: Panamé
Rasrria Heinrich, new genus
640. conops (Dyar), new comb.:
Southern
Panama
Eastern U. S.
Eastern U. S.
Panama
Species unplaced or unrecognized
brevistrigella Ragonot [Zophodia]
came Dyar [EHuzophera]
cervinistrigalis Walker [Hypochalcia]
clitellatella Ragonot [Hornigia]
corrientellus Ragonot [Elasmopalpus]
daedalella Ragonot [Luzophera]
disticta Zeller [Psorosa]
dulciella Hulst [Honora]
famula Zeller [Myelois}
flavicornella Ragonot [Phycitopsis]
formulella Schaus [Moodna]
fuscifrontella Zeller [Nephopteryz]
gais Dyar [Euzophera]
grossipunctella Ragonot [Myelois]
hulstiella Ragonot [Hypochalcia]
infusella Zeller [Myelois]
intextella Zeller [Huzophera]
trichampa Dyar [Anthropteryz]
megalopalis Hampson [Huzopherodes]
nigricans Hulst [Salebria]
olivella Hampson [Moodna]
postflavida Dyar [Euzophera]
putidella Schaus [Hucampyla]
rinmea Dyar [Huzophera]
subcanella Zeller [Zophodia]
U, S, GOVERNMENT PRINTING OFFICE; 1956
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UNITED STATES NATIONAL MUSEUM BULLETIN 207
AMERICAN MOTHS
OF THE SUBFAMILY
PHYCITINAE
By CARL HEINRICH
SMITHSONIAN INSTITUTION +¢ WASHINGTON, D.C., 1956
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II
Contents
Introuuctionr st sot) iene eo AES AL he UD NRE YA ERD g
ianuly Phymtidae see SOS SPNCMUSR |, FL, BO WOOT, BON ea
sl Pea a ONE ER cuter ten tienes ks ss aise 1s
Genera and species unplaced, unrecognized, or referred from the Phycitinae : . . .
@hecklishorAmericansehycitinae: ss tes SSS US PS A I Ss
Page
oy , om ah aaa “aw Mathonnd Bec.
ea A iN aes denne nen f
DR age a
a witey
bigs i he PAP is at hi
pei eRe es is AN: ye a a pais
AES, tis ata al et {anit ovine ination
‘hy bef TTA as ha
rs cartiolk ia. fas wi on
Sanh se nr
tigi coi mse a " nate aowhpohey ! ice
Aa Pucionhaidy oh Wetaral.” relyoitaehy: Entiat’ etalon’ geek ha oft real aN,
pane egtint vodele) qolitouona, ak obher tan) ogi of sigan mpetui,
nalitnemrae water “useing, dyad qoaaarbe ase ie Vj liye
Dv i pet a, aa pale. a Heidi be iy sia
Introduction
This paper completes a 25-year study of the New
World moths of the subfamily Phycitinae. It is based
chiefly on the collections in the United States National
Museum and the Hulst collection, formerly at Rutgers
University, supplemented by material from the Cornell
and Canadian national collections and specimens—
mostly tropical American—from the British Museum,
the Janse collection, and the collections of several South
American lepidopterists.
Recognized and included in the classification are 194
genera, 619 species, and 21 subspecies (local races). Of
these, 60 genera, 81 species, and 8 races are described
as new. The new species and races represent only a
fraction of the undescribed material examined. The
remainder consists mostly of females, chiefly from trop-
ical America and without authentically associated males
or host plants. Their description would have added
nothing to our scientific knowledge and the additional
names would have been only a nuisance to other workers.
Already too many names have been given such material.
Acknowledgments
A work of this kind could not be carried through
without generous assistance from other entomologists.
To each of them I owe a debt of gratitude: To Carl
Muesebeck, Chief of the Division of Insect Investi-
gation of the U. S. Bureau of Entomology and Plant
Quarantine, for his support and encouragement at all
stages of the project; toB. B. Pepper, State Entomologist
of New Jersey, and John B. Schmitt for permission to
examine the genitalia of the Hulst types and for their
courtesies to me at Rutgers University; to J. Bourgogne
of the Muséum d’Histoire Naturelle, Paris, for the
privilege of studying the genitalia of the Ragonot
types of American species; to N. D. Riley and to
W. H. T. Tams for the loan of unidentified tropical
American Phycitinae from the British Museum (Natural
History) and to Tams especially for photographs of
many types and their genitalia; to Martin Herring of the
Zoologisches Museum der Universitat, Berlin, for the
loan of Ragonot types; to W. T. M. Forbes, Department
of Entomology, Cornell University, for the loan of his
extensive collections from Puerto Rico, the Virgin Is-
lands, and Surinam; to J. McDunnough and T. N. Free-
man, Canadian National Museum, Ottawa, for the loan
of Canadian specimens; to A.J.T.Janse of the Transvaal
Museum, Pretoria, South Africa, for the loan of South
American Phycitinae from his collection and for much
valuable information; to A. da Costa-Lima, Escola
Nacional de Agronomia, Universidad Rural, Distrito
Federal, Brazil, for the loan of Brazilian specimens; to
Frank Morton Jones for a gift of Phycitinae collected at
Martha’s Vineyard, Mass.; to John A. Comstock for a
loan of southern California specimens; to my colleagues
at the U. S. National Museum—to J. F. Gates Clarke,
for extensive notes on the phycitid types in the Museums
of Paris, London, Oxford, and Berlin, and to Hahn
Capps, for assistance in the tedious business of slide
preparations.
My greatest debt is to the artists of the Bureau of
Entomology and Plant Quarantine for the drawings ac-
companying this paper. Where genitalia are used in in-
sect classification verbal descriptions are not enough.
Figures must accompany and supplement them to give
the reader a true picture of structural characters. The
drawings in this paper were begun in 1930 by Eleanor
A. Carlin and continued by her until October 30, 1940,
when she retired from the Bureau. From that time the
drawings were made by Sara H. De Bord, who has made
the majority of the drawings here published. Her con-
tribution was of especial value because she was not only
a capable artist but a trained entomologist as well, and
her interest in the paper and her devotion to her share
in it was so complete that she worked well on into her
last illness (she was retired on disability August 12,
1948, and died March 12, 1950). Since her death some
drawings were made by Arthur Cushman and Addie
Egbert, and the former did most of the assembling of
the plates. The drawings were all made under my
supervision and for any inaccuracies in them I am alone
responsible.
The indices were prepared by Mrs. Marguerite W.
Poole.
Abbreviation of references
To conserve space and eliminate useless repetition,
titles to certain publications frequently cited are here
abbreviated as follows:
The Ragonot ‘‘Monographie des Phycitidae et des
Galleriidae,” published as vol. 7 (1893) and vol. 8
(1901, completed by Hampson) of the Romanoff
“Mémoires sur les Lépidoptéres,”’ is cited as ‘‘Mono-
graph, pt. 1,” or “Monograph, pt. 2.”
Ragonot’s ‘“‘Diagnoses of North American Phycitidae
and Galleriidae,” 1887, is cited as ‘‘N. Amer. Phyciti-
dae,” and his ‘‘Nouveau Genera et Espéces de Phyci-
tidae et Galleriidae,” 1888, as ‘‘Nouv. Gen.”
Walker’s ‘List of the Specimens of Lepidopterous
Insects in the Collections of the British Museum,”
1854-66, is cited as “‘List.”
Hulst’s ‘‘The Phycitidae of North America,’”’ pub-
lished in the Transactions of the American Entomologi-
v
VI INTRODUCTION
cal Society, vol. 17, pp. 93-228, pls. 6-8, March—July
1890, is cited as “‘Phycitidae of N. Amer.”
Barnes and McDunnough, “Contributions to the
Natural History of the Lepidoptera of North America,”
vols. 2 (1913-1914) and 3 (1916-1917), is cited as
“Contributions.”
McDunnough’s “Check List of the Lepidoptera of
Canada and the United States of America, Part II,
Microlepidoptera,” published in the Memoirs of the
Southern California Academy of Science, vol. 2, No. 1,
1939, is cited as ‘Check list.”
Forbes’s ‘‘Lepidoptera of New York and Neighboring
States,” Cornell University Agricultural Experiment
Station Memoir 68, 1923, is cited as ‘Cornell Mem. 68.”
The distributional records for species in this paper are
obviously incomplete. They are based (with a few
exceptions noted in the text) solely on specimens I
have examined. This was the only safe procedure.
So many misidentifications have been made in the past,
even by lepidopterists of repute, that the records in
literature can not be accepted merely on the authority
of an author. Unless the specimens upon which his
statements were based can be examined and the state-
ments themselves verified, it is best to ignore them.
By taking them simply on faith and repeating them we
not only run the risk of perpetuating error, but do an
injustice to past workers who did not have or could not
use the evidence available to us.
This caution applies with even greater force to
“accepted” generic and specific synonymy. J have been
very fortunate in being able to examine the genitalia
of so many holotypes and in having authentic specimens
of most of the type species of described genera occurring
in the New World. All unqualified synonymy in this
paper is based upon genitalic examination of such
material. From the synonymy of some species common
to both the New and Old Worlds I have omitted some
names—chiefly of Old World synonyms—because I
could not examine their types and had no certainty as
to correctness of their synonymizing. Such omissions
are discussed in the text.
Classification and arrangement
A general revision has a twofold purpose, a taxonomic
and a practical one: To define accurately, to delineate
as nicely as possible, and to name categories which,
as far as our knowledge permits, represent objective
realities in nature; and to arrange these categories in
an order that permits their ready identification. Both
purposes must be served if the revision is to have any
value as a contribution to knowledge or to be of
practical use to other workers.
To satisfy both requirements I have adopted in this
paper a dual classification: a definition and division
into named categories of races, species, genera and
subfamilies; and an artificial, unnamed division,
between genus and subfamily, into groups of genera
or, in a few instances within a single genus, into groups
of species.
The named categories themselves are more or less
tentative. They are not adequate expressions of the
truth. They are only approximations to it. As we
learn more we shall have to amend or replace our defini-
tions and the categories will come a little closer to the
realities they represent. The names (except for
homonyms) will always be available; but the concepts
will change. There are several indications that taxo-
nomic groupings between genus and subfamily may
eventually be possible; and that when we have a clearer
picture of host relations and larval characters, and more
extensive collections from unexplored regions, we may
be able to establish tribes on a legitimate taxonomic
basis; but at present this is impossible. What few
definite derivations we can trace from genus to genus
show that tribal groupings would cut across the lines
of any artificial system we might be able to use.
The artificial system here adopted (based on vena-
tional characters) is proposed merely for key purposes.
The keys themselves, except for the one separating the
subfamilies, are in no true sense a part of the taxonomic
system. They are keys, and keys only. They are
intended merely to open a ready way to the descriptions
of the genera and have been constructed on the assump-
tion that they must work for all normal specimens.
I hope so, for a key that will unlock a door only 75
(or even 90) percent of the time is a tool of little worth.
Here, a word of caution. No possible key will work
for abnormal specimens. The worker in Phycitidae
must be always on the alert for them, for the family
contains an unusual number of freaks (chiefly vena-
tional abnormalities). Any one wishing to identify
phycitids must resign himself to the tedium of dissection
and slide making. Here, as in all the serious business of
science, there is no easy way, no short cut to knowledge.
The groupings of genera and species, prefaced by
brief summaries of their common characters, which I
have interposed within the text, are intended only to
assist the reader and are not.to be understood, in any
sense, as definitions of taxonomic (tribal or subgeneric)
groups. In a few instances they may be; but they are,
in intent, only divisions of convenience.
The only portion of the keys offered as a description
of taxonomic units is that separating the subfamilies
Anerastiinae and Phycitinae. This long-established
division of the family Phycitidae seems to be a sound
one, and in the main the subfamilies themselves appear
to be natural entities, although their definition leaves
much to be desired. Probably when the Anerastiinae
are thoroughly studied we may find other features more
constant than the reduced and concealed tongue.
There may even be some shifting of genera across the
subfamily lines. However, this is only hopeful antici-
pation for the future and will remain so until the Old
World genera and species of the family are thoroughly
revised. For the present we shall have to content
ourselves with an imperfect definition.
The chart opposite this page shows my interpretation
of the genera in their relation to each other and to the
system based on venational characters.
INTRODUCTION
GENERA OF AMERICAN PHYCITINAE GROUPED ACCORDING TO GENITALIA AND VENATION
Relationships on genitalic characters shown in horizontal arrangement. Venational groupings are vertical
Group 11 - H. W. vein 4s absent | Group 11
Acrobasis
Rhodophaea
Wanita cere Prosoeuzophera
Promylea
Anadelosemia
Dasypyga
(Rampylta 2 )
Monopt! lota
Caristanius
Patriciola
Tulsa
Telethusia Psorosina
Staudingeria
Heterographis|
Adelperga Stylobasis
Protasia Oedothmia
Valdivia
}Ocala
Macrorrhinia
Eumysia Divitiaca
VIII INTRODUCTION
The sequence of genera and of species within the
genus here offered isan attempt to bring together in linear
arrangement the forms showing affinities in structure
and development. It is only an attempt and I shall
quarrel with no one who objects to it.
When I began this study I had hoped to write a
monographic treatise and explore the phylogeny of the
family, but I now find that I know so much less than I
thought I did and that the accumulated knowledge of
others is so meager that any attempt along these lines
would be & vain and futile performance.
We don’t know what a primitive phycitid was like.
We don’t know which forms evolved from which, or
how. We weren’t there. We may surmise; but the
guess of one ignoramus is as good as that of another, and
there is nothing to be gained from either. I have had
to be content with a mere revision. Would that it were
more worthy.
Cari HEINRICH
Carl Heinrich (1880-1955)
This monograph was written by Carl Heinrich in the months following his retirement from
Government service in 1949. Upon it he focus>! the extensive knowledge gained during his 36
years as entomologist with the U. S. Department of Agriculture. Its publication, toward which
the Department of Agriculture has contributed substantially, was undertaken in 1954, and the
author had completed his review of the galley proofs at the time of his death, age 75, on May 31,
1955.
A biographical memoir of Car] Heinrich and a bibliography of his scientific writings appeared
in the Proceedings of the Entomological Society of Washington for October 1955 (vol. 57, No. 5,
pp- 249-255). In addition to the present bulletin, the U. S. National Museum has published a
number of his papers, of which several, as noted, are now out of print:
1921. On some forest Lepidoptera with descrip-
tions of new species, larvae, and pupae.
Proc. U. S. Nat. Mus. No. 2305, vol. 57,
pp- 53-96, 13 pls., June 17, 1920. (Out
of print.)
1923. Revision of the North American moths of
the subfamily Eucosminae of the family
Olethreutidae. U. S. Nat. Mus. Bull.
123, iv-+298 pp., 1 fig., 59 pls., Apr. 12,
1923. (Out of print.)
1926. Revision of the North American moths of
the subfamilies Laspeyresiinae and Ole-
threutinae. U.S. Nat. Mus. Bull. 132,
iv-+216 pp., 2 figs., 76 pls., Feb. 2, 1926.
(Out of print.)
1927. The American moths of the genus Diatraea
allies. Proc. U. S. Nat. Mus. No. 2691,
vol. 71, Art. 19, 48, pp., 20 pls., Aug. 23,
1927. Joint authorship with H. G. Dyar.
(Out of print.)
1929. Notes on some North American moths of the
subfamily Eucosminae. Proc. U.S. Nat.
Mus. No. 2779, vol. 75, Art. 8, 23 pp., 5
pls., Apr. 5, 1929.
1932. Notes on and descriptions of some American
moths. Proc. U.S. Nat. Mus. No. 2879,
vol. 79, Art. 13, 16 pp., 1 fig., 7 pls., Aug.
10, 1931.
1938. Moths of the genus Rupela (Pyralididae.
Schoenobiinae). Proc. U. S. Nat. Mus.
No. 3019, vol. 84, pp. 355-388. 12 pls.,
July 3, 1937.
1940. The cactus-feeding Phycitinae: A contribu-
tion toward a revision of the American
pyralidoid moths of the family Phycitidae.
Proc. U. S. Nat. Mus. No. 3053, vol. 86,
pp- 331-413, 29 pls., Mar. 16, 1939.
1945. The genus Fundella Zeller: A contribution
toward a revision of the American Pyrali-
deid moths of the family Phycitidae.
Proc. U. S. Nat. Mus. No. 3190, vol. 96,
pp. 105-114. 3 pls., May 18, 1945.
American Moths of the Subfamily Phycitinae
Family PayciripaE
Mors: Labial palpus well developed. Maxillary
palpus present, variously developed, rarely vestigial.
Tongue well developed or reduced, rarely absent; when
distinguishable, basal portion scaled. Forewing entire
(not divided); 11 veins or less; vein 7 always absent; 8
and 9 stalked or united; 1c absent (represented only by a
fold or crease in the wing membrane); no areole. Hind
wing with 8 veins or less; vein 8 closely approximate or
contiguous to or anastomosing or completely fused with
7 beyond cell; 1c always present; a fringe of pecten on
lower median vein at base; frenulum of female simple
(a single strong spine).
Larva: With primary setae only; two setae on
prespiracular shield of prothorax; setae IV and V ap-
proximate and under the spiracle on abdominal seg-
ments 1 to 8; normally a sclerotized, pigmented ring
1In Cactoblastis the aborted tongue is completely concealed by
the broadly scaled basal segments of the labial palpi. However,
the genitalia, habitus and larval affinities show that Cactoblastis
is ajtrue phycitine and must be placed with the other closely
related genera of the cactus-feeding Phycitinae.
encircling or partially encircling the tubercle of seta
IIb on mesothorax and a smilar ring encircling the
tubercle of seta IIT on eighth abdominal segment (this
character absent from the following genera of the
American Phycitinae; Htiella, Oryctometopia, Ulophora,
Rotruda, Rhagea, and Unadilia). Prolegs normal;
crochets in a complete circle.
The subfamilies of Phycitidae are separated by the
following key:
Tongue normally well developed; if sometimes reduced,
not concealed between the labial palpi (except in
Cactoblastis ); ocelli always present . . Phycitinae
Tongue reduced or vestigial; when merely reduced,
concealed between the labial palpi; if sometimes
exposed between the palpi (Bandera), then ocelli
absent Anerastiinae
Subfamily Phycitinae
The larger groups of the subfamily Phycitinae are
separated by the following key:
Key to the larger groups of Phycitinae
Hind wing with veins 3 and 4 both present .
Hind wing with vein 3 present, 4 absent . .
Hind wing with veins 3 and 4 both absent .
Group I
[Hind wing with veins 3 and 4 both present]
Keys To THE VENATIONAL Divisions AND GENERA OF Group I
Hind wing with vein 3 appreciably before the outer angle of cell; cell less than one-half the
wing length
BEVUTR UNTHRL HIS Venational division A
Cryptoblabes (p. 10)
Hind wing with vein 3 closely approximate to or from the angle of the cell (rarely shortly
stalked with 4-5); 7 and 8 approximate, contiguous, or shortly and weakly anastomosed
beyond cell; cell at lower angle nearly half as long as wing; if shorter, then vein 3 of
moderate length and the free (divergent) part of 3 decidedly shorter than vein 2. If
vein 3 sometimes appreciably before outer angle of cell (Acrobasis), then cell one-half
threrwanpylencthy.) (eye ey A ee sss
Big re neha Venational division B (key, p. 2)
Hind wing with vein 3 closely approximate to or from the angle of cell; veins 7-8 solidly
anastomosed beyond cell for at least three-fourths of their lengths.
300329—56——2
Venational division C (key, p. 7)
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Hind wing with vein 3 closely approximate to or from the angle of cell; veins 7-8 approxi-
12.
13.
tion.
mate or very weakly and shortly anastomosed beyond cell; 2 and 3 both long, the
divergent element of 3 nearly as long as 2; cell distinctly ie than half the length of
WI cr Peas atau ry inl: Ararat Sea rea eee te (tse ee a . Venational division D (key, p. 8)
Venational division B
. Hind wing with discocellular vein oblique. ...... Sa Coc taben net: Re Wie WLI Sik Yer 2
Hind wing with discocellular vein curved . . 3
. Forewing with subbasal ridge of raised scales; ed ae sii cell someinat ils dhe
one-third the length of the wing; eighth abdominal segment of male with midven-
tral hair tuftyy) «essde satomredo. - dehsReey vita. bedokyv: Mildrixia (p. 26)
Forewing smooth; hind wing with cell one-fifth the length of the wing; eighth abdominal
segment of male with sternite developed as a sclerotized, digitate pocket.
Drescoma (p. 88)
. Hind wing with vein 3 from before, but near, lower outer angle of cell. . . . . . . 4
Hind wing with vein 3 from the angle or the stalk of veins 4-5. . ........ 8
. Hind wing with veins 4-5 connate; vein 6 of forewing always straight. ...... 5
Hind wing with veins 4-5 approximate, contiguous, anastomosed or stalked for a short
distance from cell, if sometimes connate (on individuals of Hypsipyla) vein 6 of fore-
wing slightly bent towards base ........-..2.... Satis) ih ee eee eR
. Basal segment of male antenna triangulate. ........... Acrobasis (p. 11)
Basal segment of male antenna simple (cylindrical): ...... Rhodophaea? (p. 24)
. Forewing with subbasal ridge of raised scales; antenna of male pubescent (cilia distinctly
shorter than width of shaft) .............4... Anabasis (p. 25)
Forewing smooth; antenna of male ciliate (cilia distinctly longer than width of shaft) . 7
. Forewing with vein 6 straight, remote from veins 8-9 at base . . Sematoneura (p. 27)
Forewing with vein 6 bent, shortly separated from 8-9 at base . . . Hypsipyla (p. 27)
. Vestiture of head, thorax, labial palpi, and femora a mixture of scales and hairs; male with
harpe short (stubby), clasper absent, apical process of gnathos an inverted heart-
shaped lobe with short, slender spine; female with ventral surface of genitalia sclero-
tized throughout its length, ductus seminalis from near junction of bursa and ductus
DUESHED te tects eee ron yn icc ie Aa an |, Nena Beep On ane . Polopeustis (p. 120)
Vestiture entirely of scales; or; if occasionally mixed with hair (Saraéa airella), male with
harpe elongate, clasper present, knoblike, apical process an elongate, stout hook;
female with ductus bursae unsclerotized, ductus seminalis from bursa remote from
junction of bursa and ductusbursae .........2.2.+.-.-++-+++-+-+.-. 9
. Forewing with subbasal ridge ofraisedscales . ........-.4...- spay yl
Foréwimgismootll iis) 2 be ey GN Eee ala a INT URS Saleh ee a 16
. Hind wing with vein 3 from the stalk of vems4—5 =... ..... Passadena (p. 175)
Hind wing with vein 3 approximate to but not from the stalk of vems 4-5 ..... 11
. Labial palpus porrect, beaklike . . ..........++e+-- . KEtiella (p. 98)
Labial palpus obliqueorupturned . . 2. 2... ee ee ee 12
Forewing with vein 6 from the upper angle of the cell, bent towardsbase. ........
Hypargyria (p. ee
Forewing with vein 6 from well below the angle of cell, straight. . . .. . ay scree
Forewing with veins 4 and 5 slightly separated at base and approximate for a short ste
tance beyond 2 te cnndw aa wool ae-tad vitor eface eel deeiias o Paar beeoms 14
Forewing with veins 4 and 5 connate or stalked; if sometimes shortly separated at base
notapproximate beyond. ...... 2. ee eee ee es 15
2 The genus Trachycera is omitted from our key as the male is unknown except from Ragonot’s descrip-
The female before me has the venation of Rhodophaea. It is distinguished from other females of
Division B in haying two scobinate, cuplike signa similar to those of Davara in division D. (See p. 25.)]+
14,
15.
16.
17.
18.
19.
20.
21.
22.
23.
24,
25.
26.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Hind wing with veins 7-8 anastomosed beyond cell; maxillary palpus of male in the form
oftaneierette...... OR CLIk Were eReIUG aioth dir olan Immyrla (p. 111)
Hind wing with veins 7-8 approximate beyond cell; maxillary palpus of male squamous.
Ulophora (p. 176)
Labial palpus oblique, second segment on male grooved; male with eighth abdominal
segment simple; female with genitalopeningsimple .... . Ortholepis (p. 119)
Labial palpus upturned, second segment on male not grooved; eighth abdominal segment
of male with compound tufts; female with strongly sclerotized plate at genital open-
ing attached to supplemental eighth-segment collar ...... Tlascala (p. 133)
Labial palpus porrect, beaklike; male with aedeagus flanged and strongly spined; female
with ductus seminalis-from bursa .....-.+...-.» Polliyom phonyage ditore |) ily
Labial palpus oblique or upturned; if sometimes appearing porrect (Stylopalpia) due to
long, deflected third segment, male with aedeagus simple; female with ductus semi-
nahstirom ductus burske)) .Joge 2 Jol Seyieqg oy enigy aliwy yogis |. 21
Forewing with veins 4-5 approximate for a short distance from cell. . . . 2... 18
Forewing with veins4—5 stalked. . 2. 2 ek ee, 19
Labial palpi extending at least twice the length of head beyond it; harpe of male elongate;
female with ductus bursae much longer than bursa ........ Pima (p. 101)
Labial palpi extending little more than the length of head beyond it; harpe of male short;
female with ductus bursae little, if any, longer than bursa . . Interjectio (p. 106)
Male with second segment of labial palpus grooved on inner side; female with ductus
seminalis from bursa remote from junction of bursa and ductus bursae . Sarata (p. 159)
Male with second segment of labial palpus not grooved; female with ductus seminalis
from bursa near junction of bursaand ductus bursae . ............ 20
Female with a strongly sclerotized plate behind genital opening . . Philodemia (p. 165)
Female without sclerotization at genital opening ........ Lipographis (p. 166)
Hind wing with vein 3 from the stalk of 4-5 or closely approximate to it for some distance
from lower outer angle of cell; vein 2 alwaysratherneartheangle. ....... 22
Hind wing with vein 3 connate with the stalk of veins 4—5 or connected with it at base by
a very short spur; if sometimes approximate to the stalk of 4-5 (Megarthria, Acron-
cosa), vein 2 always from well before lower outer angle of cell; or, if vein 3 some-
times shortly fused with the stalk of veins 4-5 (Actriz), male with apical process of
gnathos developed as a square or inverted heart-shaped plate and female with
caudal half of bursa copulatrix densely spinose. . . ..... 2... ee 28
Hind wing of male with anal angle folded into a pocket; female with strong sclerotizations
behind or surrounding genital opening of genitalia... .........4.. 23
Hind wing of male without folded pocket at anal angle; genital opening of female
simple. ico, soley Sodan bestjowloe goods Miw has gerd er papi: 24
Male genitalia with uncus hammer-clawed (long, curved, constricted at middle and
broadly divided at apex); female with bursa containing strongly sclerotized folds
or stoutly spmedibandstsimer lagunohds sigan, .outd boozad, Fundella (p. 59)
Male genitalia with uncus otherwise (sometimes broadly divided at apex but the divided
elements small and spinelike and the middle of uncus not appreciably constricted) ;
bursa sometimes with a small granulate patch but otherwise membranous and
RUDAPITOG 5 hiss econ Shwe Ni Ay cisync socse Ae Mates es © OMQUNse SOONIOOR: Difundella (p. 62)
Hind wing with vein 3 approximate to the stalk of veins 4—5 for some distance from outer
pngle‘obealliisls et SvSiibogl SURRY Gy ugnooyiay Starts Wig ¢ Scorylus (p. 72)
Hind wing with vein 3 from the stalk of veins 4-5... 2... ee ee 25
Male genitalia with transtilla a sinuate, sclerotized scobinate band involved with gnathos
and with a long free spine involved with anellus; female with ductus seminalis from
ductusbirsaesioielor . .sialqmigs alivaayny. ine egdyq leds tig? gauge ig Bia, 26
Male genitalia with transtilla incomplete or absent; female with ductus seminalis from
bursa l! ...Ueo brayed aanntelh taods ww busompienin 2S ete? tipw wii DM 27
Maxillary palpus of male filiform. . ........2.2.200.. Coptarthria (p. 64)
Maxillary palpus of malesquamous. ............. Anadelosemia (p. 67)
27.
28.
29.
30.
31.
32.
33.
34,
35.
36.
37.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Male genitalia with divided element of incomplete transtilla strongly sclerotized, gnathos
absent; female with ductus bursae partially sclerotized .... . Gabinius (p. 84)
Male genitalia with transtilla absent, gnathos well developed and with apical process
a strong hook; female with ductus bursae membranous throughout.
Ceracanthia (p. 85)
Male genitalia with transtilla a sinuate sclerotized band involved with gnathos; female
with bursa small, membranous and ductus bursae much longer than bursa, signum
(if present) a small granulate patch or small plate with single minute thorn, genital
opening narrow (the ductus bursae never expanded into a widened opening) ; if bursa
sometimes large and ductus bursae proportionally shorter (Rampylla), collar of
eighth :sepment modified. \y.%5)..\)5))9 hea orleans Bee eben ae 29
Male genitalia without transtilla or, if present, otherwise; female genitalia never as
aboyeuin-all details} ents sanehaaulie shee heal: Duke betes Bas pees 31
Forewing with veims 4—5 approximate for a short distance from cell; hind wing with
vein 2 from well before lower outer angle of cell. . . . .......2...-. 30
Forewing with veins 4-5 stalked; hind wing with vein 2 from near outer angle of cell.
Dasypyga (p. 69)
Female with ductus seminalis from ductus bursae. ........ Promylea (p. 65)
Female with ductus seminalis from bursa copulatrix ........ Rampylla (p. 70)
(males: venation group D, couplet 8)
Male with transtilla of genitalia complete and strongly sclerotized; or, if incomplete, the
elements enlarged, strongly sclerotized and modified; when complete not in the
form of a squarish plate. Female with a single signum developed as a small, scobin-
ate or granulate cup or patch or (Adanarsa) as a single short, stout thorn; genital
openingvelways broady sty.) sd}isoss heilidoe alata aaatieise vimaciie et Ss cae 32
Male genitalia with transtilla incomplete or absent. Female with signa or signum, if
present, otherwise developed. If transtilla complete then weakly sclerotized or
developed as a square plate; and if signum of female a small scobinate patch (Zegar-
thria alpha) genital opening narrow... .......2..+.+-+.+.4+2-s 39
Forewing with vein 6 bent towards base. . ......-....-.+-++-+--- 33
Forewing with vein 6 straight. 2 2.0.0. 2 ee ee 36
Transtilla of male complete, strongly arched and with median areaforked ..... . 34
Transtilla of male complete, but not arched nor with median area forked. .... . 35
Antenna of male with shaft unipectinate. Ductus bursae of female very short, less than
one-half as long as bursa, scobinate-granulate and more or less sclerotized but not
transversely wrinkled ............2..26: Hemiptilocera (p. 30)
Antenna of male with shaft pubescent. Ductus bursae of female longer, about half as
long as bursa and with strong, sclerotized wrinkling before genital opening.
Crocidomera (p.32)
Maxillary palpus squamous. Forewing of male with costal fold and a fovea in cell
slightly beyond base. Highth abdominal segment with sternite developed as a
sclerotized pocket. . ...........24.4- ONG tits mt Heras (p.34)
(male only, female unknown)
Maxillary palpus filiform. Forewing of male without costal fold or fovea. Eighth
abdominal segment simple... ........ 52 ee eee Birinus (p. 36)
(male only, female unknown)
Male antenna with shaft unipectinate; transtilla incomplete, its elements long, stout with
their apices broadly and irregularly developed and hooked. Female with a large
semicircular sclerotized and scobinate plate on membrane behind genital opening.
Bertelia (p. 36)
Male antenna with shaft pubescent; transtilla complete. Sclerotized plate on membrane
behind genital opening of female, if present, not semicircular. . ....... 37
Hind wing with veins 7-8 anastomosed for a short distance beyond cell. Male with trans-
tilla strongly arced but with its median area bearing a smooth narrow crosspiece, not
forked. Female with signum a single, short, stout, hooked thorn. Adanarsa (p. 35)
38.
39.
40.
41.
42,
43.
44,
45.
46.
47.
48.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Hind wing with veins 7-8 approximate for a short distance from cell. Male transtilla
otherwise. Female with signum a single, small, scobinate or granulate, cup-shaped
patcliys, heels ier a sued ET “Qpdzly “Pl, Si 38
Male with apical process of gnathos a stout houk. Fennle with ductus burdie appreci-
ably longer than bursa; ductus seminalis from ductus bursae . . Cuniberta (p. 34)
Male without projecting apical process, the lateral arms supporting a thinly sclerotized
subanal plate. Female with ductus bursae much shorter than bursa; ductus semi-
nalis from lobe of bursa near junction of bursa and ductus bursae. Chararica (p. 38)
Male with sternite of eighth abdominal segment developed as a sclerotized pocket.
Female with signum a small, depressed, granulate-scobinate patch; or, if signum
absent, ductus bursae narrow throughout and sclerotized for about one-third of its
length from shortly beyond its junction with bursa, the sclerotized portion sharply
bentigveriteok.. - .. mous Croom Ha, viedioie, rut, Megarthria (p. 86)
Male with eighth abdominal ebeniaite ib developed as a sclerotized pocket. Female
with signum (or signa) developed as sclerotized and strongly spined plates or bands,
or entirely absent. If signa absent, ductus bursae not as above. If sometimes
(Olybria) a single more or less strongly sclerotized band in bursa at junction of bursa
and ductus bursae, the band finely serrate or edged with short spines . . . . .40
Antenna of male with shaft unipectinate. Female bursa with a single strongly spined
sipnammosele jon elagiquos asiouh Bogatvig adalat pagoteq Monoptilota (p. 89)
Antenna of male with shaft pubescent. Female bursa with two signa or none . . .41
Male genitalia with a pair of long, strong, sclerotized arms from the ventrolateral angles
of uncus; gnathos absent. Female with two signa consisting of stoutly and coarsely
spined bands: ductus bursae strongly sclerotized, fattened (ribbonlike).
Caristanius (p. 97)
Male genitalia with uncus otherwise; gnathos present and well developed. Female with-
out signa; or, when present, consisting of two strongly spined bands or plates; when
signa are present, ductus bursae not as above, if partially flattened and ribbonlike
thenhyerymarrowe “Ute wtiibe, viitigdel Dodlay®, oc, T-h elev AUPE Qniwe . .42
Male genitalia with aedeagus expanding to lateral, flanged projections before apex, the
flanges each bearing a cluster of strong spines. Female with ductus seminalis from
ducinsibursde: sTaheel Ha Base soit inuiels Vibe ais Stylopalpia (p. 140)
Male genitalia with aedeagus otherwise, if sometimes spined (Pyla), not flanged before
apex. Female with ductus seminalis from bursa copulatrix ......... 43
Masallany palpus ‘vestigial .. °°.0". Gaertn ond) oho. PG Wee Ophah URE tyne 44
Maxillary palpus squamous, or filiform, or (on some males) in the form of an aigrette . 45
Penis of male unarmed except for a eran cluster of very weak, short, slender spines.
Female with ovipositor strongly sclerotized. . . ..... ‘ Télethiusia (p. 186)
Penis of male armed with a single, strong cornutus. Female with ovipositor normal (not
sirosply ‘scleretized)d ier pets Sew Se Saskies sured al. Phobus (p. 138)
Forewing with some rough (raised) scaling in median area beyond outer border of ante-
median line, but without subbasal ridge of raised scales . . . . . Tulsa (p. 134)
Forewing Sfadath Yr eet om oee ated CUCSTLONIO COTY Ah pis Ae, TAG
Fore tibia with a long inner and a short outer claw ....... Nleei cask (p. 174)
Honatibin atherwasexisoiu, res eit, TRADE SEY Ailes Ws, WAT
Hind wing with veins 7-8 distinctly anastomosed for about half hein ddngths (more than
the free length of vein 8). Male with apical process of gnathos a broad shield
without central terminal spine .........2.2.. oy iter PAGtrix, (ps 139)
Hind wing with veins 7 and 8 approximate; if sometimes contiguous or anastomosed be-
yond cell, then weakly and very shortly so. Male with apical process of gnathos
otherwise al Sjad eased holla adchanibesritos <ihisciphagg CIN. LECENER CIOL, OFT... 48
Male genitalia with a pair of straight, strongly sclerotized arms [not to be confused with
similar projections from vinculum in Nephopteryz crassifasciella] projecting back-
ward from lower, posterior angles of tegumen. Female with ductus bursae of gen-
italia flattened (cibboniike), waved (twice bent) and sclerotized throughout, the
UNITED STATES NATIONAL MUSEUM BULLETIN 207
sclerotization developed at genital opening into a stout, squarish ventral
platejalycog-to.ntaniloon Sluis heals ge Beenie eM okepeas, ae Olybria (p. 113)
Male genitalia without projecting arms from tegumen. Ductus bursae of female
otherwisejs:'. dois’ elasne tl. alood tote 3 ROR, PY Ree OSG. Wield olely 49
49. Harpe of male genitalia with a long hair brush from inner surface, along lower edge of
basal half of sclerotized costa. Bursa copulatrix of female with the lobe giving off
ductus seminalis strongly sclerotized; or most of dorsal area of bursa strongly and
smoothly sclerotized; or bursa with conspicuous, round or oval, strongly pigmented
and sclerotized, densely granulate patches [absent in Nephopteryx subcaesiella (fic.
826)], the bursa otherwise spinose overitsmembranous areas. ........ 50
Harpe of male without such hair brush. Bursa of female not as above ..... 53
50. Male with sacculus of harpe strongly produced at apex. Female with most of dorsal
surface of bursa strongly and smoothly sclerotized..... . Glyptocera (p. 100)
Male with sacculus of harpe simple (not produced at apex). Female with only the lobe
giving off ductus seminalis sclerotized: or bursa with granulate sclerotized
patchesgvicais aa sou oneal aiogh Soesitn eee Es aisacte glares Ge 51
51, Penis of male armed with numerous strong, slender spines.. Female with lobe of bursa
giving off ductus seminalis strongly sclerotized. ......... Oreana (p. 112)
Penis of male armed with two stout spines or (very rarely) one spine. Bursa of female
with granulate patches, the lobe giving off ductus seminalis not sclerotized. . . 52
52. Harpe of male genitalia with an appressed, stout, thorny or serrate clasper. Female
with ductus bursae unsclerotized adjacent to bursa, but with strong sclerotization
dti-cenital openings; izcas via cat iy esi. takede pong Meroptera (p. 121)
Harpe of male with clasper digitate, slender, simple (without spining). Female with
ductus bursae sclerotized along ventral surface from junction with bursa, the sclero-
tization terminating before genital opening, the latter simple (unsclerotized).
Nephopteryx (p. 123)
53. Forewing with veins 4—5 stalked for nearly half their lengths. . . . . Tacoma (p. 178)
Forewing with veins 4-5 not stalked (slightly separated or closely approximate at
cell) tax neciios ier berated olemiol of palomneme eoasbes Aiiiy sileisivs.. . . 54
54. Male genitalia with transtilla complete but very weakly sclerotized (a narrow angulate
band); aedeagus slender, elongate, not spined or divided. Female genitalia with
bursa entirely membranous and smooth; ductus bursae rather narrow, tubular and
sclerotized. throughout, nowhere appreciably widened; genital opening narrow (no
wider than narrowest part of ductus bursae)........... Tota (p. 170)
Male genitalia with transtilla incomplete or absent; aedeagus moderately broad to stout,
if sometimes slender and elongate then spined or partially divided or (Chorrera)
vinculum with produced lateral lobes from terminal margin. Female with or with-
out signa in bursa; if without signa, bursa spinose or with same sclerotizations
adjacent to ductus bursae; if bursa entirely membranous and smooth, genital
opening decidedly broadened . . . .. 2... 2... ee ee ee ee ts 55
55. Male with aedeagus of genitalia spined or partially divided; if sometimes simple, then a
strong knoblike and spinose projection from harpe at base of costa. Female without
signa in bursa, the latter usually simple (unsclerotized and smooth), but sometimes
with sclerotized, convolute, longitudinal bands near junction of bursa and extending
into the ductus, the bursa never spinose . .......-..... Pyla (p. 142)
Male with aedeagus neither spined nor divided; if harpe sometimes with projection from
base of costa, the latter neither knoblike nor spinose. Female with or without signa
in bursa; if without signa, the bursa spinose (at lease partially so) or sclerotized at
junction of bursa and ductus bursae, but such sclerotization not in the form of
econvolute, Joneitudinal bands, 2. 2. ee ee ww ee ee OU , 38
56. Male with maxillary palpus squamous or filiform. Female without signa; a serrate or
minutely spined sclerotization at junction of bursa and ductus bursae, the latter
broad and strongly sclerotized or with a pair of elongate sclerotized plates behind
57.
58.
59.
60.
61
62.
63.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
genital opening; if signa present, the ductus bursae very narrow and expanded
into a membranous globe shortly before genital opening. . ......... 57
Male with maxillary palpus in the form of an aigrette. Female with or without signa;
bursa and ductus bursae not as above. . . 2... 6 ee ee ee 59
Male genitalia with aedeagus stout; penis armed with a single, long stout spine. Bursa
of female geriitaliawwithoutisignan oli i! soot lS ance sNodle lag spelled 58
Male genitalia with aedeagus very slender; penis unarmed. Bursa of female containing
two signa developed as opposed, longitudinal bands bearing a row of short stout
EPINES..;\. (sg cs le elt, aetdete bis (busolo).roits Chorrera (p. 177)
Maxillary palpus of male minute, filiform. Female without sclerotized plate or plates
behind «genitaliopaning ay. js cobindin, newt aecaud tical elenies Ambesa (p. 108)
Maxillary palpus of male squamous (broadly scaled). Female with a pair of elongate
sclerotized plates on inner dorsal surface of ductus bursae behind genital
openinewamayyis eesdst. leokje (sal .oiolayanimi-cam 20 obs Catastia (p. 110)
Male with sacculus of harpe slightly produced at apex. Female with ductus bursae of
genitalia cylindrical; narrow except at genital opening; appreciably longer than
length of bursa; sclerotized for half its length from junction with bursa, the sclero-
tization longitudinally ribbed ............. Elasmopalpus (p. 172)
Male with sacculus of harpe not produced at apex. Ductus bursae of female other-
Wiseweneti | .wanowiie idente! divtaiiniivads yore doisdiiocule laches sti tis . 60
Hind wing with vein 2 from before but rather near outer angle of cell Atiw qwatcucee 61
Hind wing with vein 2 from well before outer angle ofcell . . . 1... .... 62
Male with penis armed with a single stout spine. Female with ductus brusae of genitalia
sclerotized (at least towards genital opening). Forewing with veins 8-9 stalked for
less than two-thirds of their lengths. Hind wing with veins 4-5 stalked for approxi-
mately one-half their lengths. ... 2... 2... ee ae Salebriaria (p. 115)
Male with penis unarmed. Female with ductus bursae cartilagenous, except at its
junction with bursa. Forewing with veins 8-9 stalked for over two-thirds of their
lengths. Hind wing with veins 4-5 stalked for over three-fourths of their
lengths: Sein Gets. below orneth Insdusdie¥ a ctiie se - Quasisalebria (p. 118)
Male genitalia with a strong, straight or curved arm from base of costa of harpe; penis
unarmed. Female genitalia with ductus seminalis from bursa adjacent to junction
of bursa and ductus bursae (signa present). ......... Adelphia (p. 168)
Male genitalia without projecting arm from base of costa of harpe; penis armed with a
single strong spine. Female with ductus seminalis from bursa remote from junction
of bursa and ductus bursae (signa present or absent). .... Le ay S463
Male with cornutus on penis a long straight spine, over half as long as wed Garuke Female
with bursa densely spinose, signa absent; ductus seminalis from near anterior end of
bursa “sah: diate weeds Cine easier i sieeve leeeapetan eit Salebriacus (p. 114)
Male with cornutus on penis a short, curved spine, somewhat less than one-third as long
as aedeagus. Female with bursa smooth except for strongly spined signa (and in
one species a strongly spined collar at middle of bursa) ; ductus seminalis from bursa
Sa juceninboronevOh tie Siena Skee roe ele ene alee tantel esis tess Ufa (p. 170)
Venational division C
. Hind wing with discocellular vein straight, vertical. . . . . . . Homoeographa (p. 135)
Hind wing with discocellular vein straight, oblique. ......... Atheloca (p. 81)
Hind wing with discocellular vein fainted O.Otw. stand . Srad otalngar ori. ... 2
. Male with uncus of genitalia more or less spoon-shaped (the lateral margins deeply
concave at middle). Female with signa present, consisting of 2 or more sclerotized
disks or series of contiguous, bluntthorns. . . . 1... 2... 1 ee ee eee 3
Male with uncus triangulate or subtriangulate. Female with or without signa; if present
MOT ASIADOVOSs : RRR renee is cS AT, GI Nye boos se opie caer catteb ett 3 4
UNITED STATES NATIONAL MUSEUM BULLETIN 207
. Male with costal fold on forewing; shaft of antenna notched at base . Diatomocera (p. 50)
Male without costal fold on forewing; antennal shaft not notched . Pseudocabima (p. 53)
. Hind wing with vein 3 from the stalk of veins 4-5. ........ Anypsipyla (p. 42)
Hind wing with vein 3 from the angle of the cell; if sometimes approximate to the stalk
of veins 4—5 for a short distance, never actually fromit. ........ 2a NE XS
. Maxillary palpi of both sexes filiform. Male with complete transtilla ....... 6
Maxillary palpi squamous. Transtilla of maleimcomplete ........2.... 8
. Male with antennal shaft pubescent. Female with ductus seminalis from near middle or
towards anterior (closed) end of bursa .......2.2.2.2.2.2.4.22... : 7
Male with antennal shaft shortly ciliate (cilia as long as width of shaft). Female with
ductus seminalis from bursa near junction of bursa and ductus bursae.
Kcetomyelois (p. 43)
. Forewing with vein 2 from near outer angle of cell. Male with transtilla weakly sclerotized
(a thin band or sub-triangulate plate); apical process of gnathos broadly U-
shapedih sabivaiaen ts .. aeeiey tithe Devs Decrees enter shah: etd Tay: ret ge Myelopsis (p. 40)
Forewing with vein 2 from well before the angle. Male with transtilla strongly sclero-
tized and arched; apical process of gnathos a simple, stout hook . . Apomyelois (p. 42)
. Hind wing with cell moderately long (from a little over to slightly less than one-half the
length ciitheiswing)iingh) .. 2a. dk Roa bela aaa do. eoluad tin, eels. 9
Hind wing with cell short (not over one-third the length of wing). . Protomoerbes (p. 49)
. Forewing with vein 2 from near lower outer angle of cell; vein 10 from the cell. Male
with eighth abdominal segment simple. Female with signum in bursa (a cluster of
coarse scobinations); a sclerotized plate behind genital opening (on inner dorsal
surface of ductus bursae). ............2... Paramyelois (p. 46)
Forewing with vein 2 from well before the angle; vein 10 from the stalk of veins 8-9.
Male with a pair of ventrolateral hair tufts on eighth abdominal segment. Female
with bursa and ductus bursae simple (membranous throughout, with neither signum
in bursa nor plate behind genital opening) ......... Pseudodivona (p. 48)
Venational division D
1. Normal dark discal spots on forewing at end of cell replaced by a conspicuous white spot
or line (obscured only on clarioralis) on discocellular vein. [Male genitalia with
eostal area of harpe broadly sclerotized and produced at apex; clasper present,
erect; penis armed with numerous, straight spines. Female with signa developed as
2 or 3 clusters of strong, slender spines; ductus bursae flattened, strongly sclerotized
over most of its length, the sclerotization terminating just before simple genital
roy OLSSON kes el eal lA be SiN all A, ian) adh Seam ati eh on Saat 78 Dioryctria (p. 149)
No such white spot on discocellular vein. . .......2.2.+..4.+..4.-. sae
2. Forewing with ridge of raised scales preceding antemedial line .......... 3
Forewing smooth; if sometimes with a few roughened scales, no such subbasal ridge. . 4
3. Forewing with veins 4-5 closely approximate for a short distance from cell. Hind
wing with cell less than one-fourth the length of wing. Maxillary palpus of male
in) the form\ofjansierette)) 278. fine soe 4 6 jue ee es Zamagiria (p. 90)
Forewing with veins 4-5 connate or very shortly stalked. Hind wing with cell slightly
less than one-third the length of wing. Maxillary palpus of male subsquamous.
Anegcephalesis (p. 93)
4. Male genitalia with uncus and tegumen greatly reduced; uncus a narrow, weakly sclero-
tized, angulate band. Female with a single signum in bursa, consisting of a small,
sclerotized plate supporting a very short thornlike spine. . ... . Peadus (p. 83)
Male genitalia with uncus and tegumen well developed. Female with signum or signa
Gf present) otherwise’. 6... “eesti! corace ot) Ween tal BS 5
10.
11.
12.
13.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
. Male genitalia with transtilla complete, developed as a narrow, slightly arched band,
attached to harpes only by membrane. Female with signum a cluster of bluntly
pointed thorns, more or less surrounded by fine scobinations or strongly pigmented
Eraniuations s °S SOOM RN, STR Aibe artiunind Hyalospila (p. 56)
Male genitalia incomplete or absent; if complete (Magiriopsis) not a narrow band.
Female with signum or signa (if present) otherwise. .......2.2.... 6
. Hind wing with discocellular vein incomplete ........2... Fulrada (p. 71)
(based on male; female unknown)
Hind wing with discocellular vein complete. . . . 2... .... 2 ee ee ee 7
. Hind wing with cell very short (about one-fifth the length of wing). Male genitalia
with lateral arms of gnathos broad, expanded and curled at their extremities.
Praedonula (p. 82)
Hind wing with cell longer (from one-fourth to one-third the length of wing). Gnathos
oftmale! otlierwiselt! Hrw. OL 20 8 clay ve heal cn ny alts 8
. Hind wing of male with anal area (involving vein la) thicker? aad folded, Genie
a produced pocket; underside of wing with roughened scale or hair tufts on some
OLGHELVeMIS hr tii TS 4 ca pica, oc. « ACORN EheaILD, Abewes Rampylla (p. 70)
(females: Venation Group B, couplet 30)
Hind wing of male without such modification. . ......... IHS BOG
. Male with uncus of genitalia bifid. Female with one or two small signa developed
as granulate depressions in bursa; if signa sometimes absent (some species of
Piegnoppeo) puts: Melnbrapousy . . . 0... yes my ee ee ee 10
Uncus of male undivided (triangulate or benanonen iaene of rorinle developed as
strongly, spined bands or plates; if sometimes absent (Ancylostomia), bursa weakly
but, extensively sclenotizedy). QING? .. me actran-» ‘agasne ee taut repmatend oslanns 12
Gnathos complete, a thin, weakly sclerotized, transverse band. Female with two
SISTEM DUIS ee. ek ye te et a DONG AMAe sian orld jeeps MPa ded Ye bn Spy. elih
Gnathos incomplete, the lateral arms strong, broad; between their separated apices a
well sclerotized anal plate. Female with one signum or none. . Piesmopoda (p. 77)
Forewing with vein 6 from below upper angle of cell, separated at base from the stalk
of veins 8-9. Male genitalia with a stout free spine associated with anellus.
Davara (p. 73)
Forewing with vein 6 from upper angle of cell, connate with the stalk of veins 8-9.
Male genitalia without free spine associated with anellus . . . . Sarasota (p. 76)
Male antenna unipectinate for basal half of shaft, shortly ciliate beyond. Female
with two signa developed as strongly spined plates . ..... Magiriopsis (p. 94)
Male antenna with shaft pubescent. Female with one signum ornone ..... . 13
Male with maxillary palpus in the form of an aigrette; eighth abdominal segment with
compound ventral scale and hair tufts. Female genitalia without signum; the
bursa copulatrix weakly sclerotized throughout. . ..... Ancylostomia (p. 95)
Male with maxillary palpus squamous; eighth abdominal segment with paired ventro-
lateral hair tufts. Female with signum, consisting of a single round, curved plate,
densely armed with long stiff spines; bursa otherwise membranous.
Oryctometopia (p. 158)
10 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Genus 1: Cryptoblabes
[Venational division A. Hind wing with vein 3 distinctly before
lower outer angle of cell; 7 and 8 approximate, or weakly and
shortly anastomosed beyond cell. Forewing with vein 6 bent
towards base; 10 from the cell. Male genitalia with transtilla
complete; uncus bilobed.]
1. Genus Cryptoblabes Zeller
Cryptoblabes Zeller, Isis von Oken, 1848, p. 644.—Ragonot,
Monograph, pt. 1, pp. xliv, 12, 1893.—Staudinger and Rebel,
Catalog der Lepidopteren des palaearctischen Faunenge-
bietes, vol. 2, p. 42, 1901.—Mayrick, Revised handbook
of British Lepidoptera, p. 397, 1928.—Bisset, zn Pierce and
Metcalfe, Genitalia of the British Pyrales, p. 57, 1938.—
Janse, Journ. Ent. Soc. South Africa, vol. 14, p. 143, 1941.
(Type of genus: Cryptoblabes rutilella Zeller, a synonym of
bistriga (Haworth); figs. 2, 131, 638.)
Tongue well developed. Antenna pubescent; shaft
of male notched at base and with curved, horny hook
protruding from the notch. Labial palpus upturned,
slender, reaching a little above vertex; third segment
about two-thirds the length of second, acuminate.
Maxillary palpus squamous. Forewing smooth; 11
veins; vein 2 from well before the lower outer angle of
cell (from lower median vein of cell at about three-
fourths); vein 3 also before the angle (from lower
median at about five-sixths); 4 and 5 closely approxi-
mate at base, rarely (in individual specimens) connate;
6 bent towards base, from upper angle or from very
close to upper angle of cell; 10 from the cell, separated
from stalk of 8-9 at base; male without costal fold.
Hind wing with vein 2 from middle or just beyond mid-
dle of lower median vein of cell; 3 from before and
more or less removed from the outer angle of cell; 4
and 5 from the angle, closely approximate at base,
thence diverging, 7 and 8 closely approximate, con-
tiguous or weakly anastomosing for a short distance
beyond cell; cell less than one-half the length of wing,
but not ‘very short” as stated by Ragonot in his
generic key (Monograph, p. xliv); discocellular vein
curved. Highth abdominal segment of male simple.
Male genitalia with apical process of gnathos a small,
simple hook. Transtilla complete; developed as a
narrowly banded bridge with more or less elongate
central projection. Uncus broad, with apical margin
broadly rounded and invaginate, giving the uncus a
bilobed appearance. Harpe (in European species) with
strong hair tufts arising from articulated plates in
intersegmental area between base of sacculus and ter-
minal margin of eighth abdominal segment, or with
long hair tuft from sacculus near its base (rutvlelia).
Anellus with elongate, narrow, lateral lobes. Aedeagus
simple; penis with or without cornutus, latter, when
present, a single, long, spine. Vinculum broad and
with broad terminal margin more or less concave.
Female genitalia with ductus bursae membranous,
finely scobinate towards junction with bursa; genital
Opening simple except for a narrow, sclerotized band
behind and above the opening; bursa membranous
more or less finely scobinate; signum present, developed
as a stout, blunt, flattened thorn (rutilella) or a patch
of dense granulations (gnidiella); ductus seminalis
from bursa.
The foregoing description was drawn from European
species which are obviously congeneric. Numerous
other species have been described in the genus from
India, Formosa, Japan, Australia and the islands of
the Pacific. Whether these are all congeneric I do not
know. Two unidentified species before me from the
Philippines have genitalia similar in habitus to those
of the genotype (rutilella) except that the dorsal, inter-
segmental tufts at base of harpe are lacking. Their
unci have the same characteristic bilobed appearance.
The venational character which has been generally
accepted as defining the genus (i. e., the position of vein
3 of hind wing in relation to the lower outer angle of
cell) is variable and illusive, being closer to the angle
in gnidiella (fig. 1) than in rutilella (ig. 2) and still
closer in the Philippine species. Indeed, in American
examples of Acrobasis (=Mineola) vein 3 is often as far
from the angle as it is in gnidiella. The shorter cell of
Cryptoblabes, coupled with the position of vein 3, will,
however, suffice to maintain the group separation made
between the two genera in our key.
Cryptoblabes is a distinctly Old World genus with no
indigenous New World species. Itis represented in the
Western Hemisphere by only one introduced European
species (gnidiella).
1. Cryptoblabes gnidiella (Milliére)
Fieures 1, 132, 639
Ephesiia gnidiella Mulliére, Iconographie et description de
chenilles et Lépidoptéres inedits, vol. 2, p. 308, 1864.
Cryptoblabes gniediella (Millitre) Ragonot, Monograph, pt. 1,
p. 16, 1893.—Dyar, Ins. Inse. Menstr., vol. 3, p. 88, 1915.
Forewing pale brownish fuscous with a faint rosy
overcast due to more or less diffused longitudinal streaks
of reddish scales (in fresh and well marked specimens
especially along the fold, on the veins in outer area, and
below costa from base); some whitish dusting along
costa and in the cell, most pronounced as a pale shade
between the dark discal spots at end of cell; transverse
lines obscure and not sharply outlined, but distinguish-
able, whitish ocherous; the antemedial line oblique and
curved, set well out towards middle of wing; subter-
minal nearly straight, parallel with termen; discal dots
separate, blackish fuscous. Hind wing whitish, trans-
lucent, the veins darkened, a narrow dark shade along
costa and a narrower dark line along terminal margin.
Alar expanse, 11-16 mm.
Male genitalia with heavy hair tufts from plates
articulating with base of sacculus of harpe; penis armed
with spinelike cornutus about two-thirds as long as
aedeagus. Female genitalia with signum developed as
a dense scobinate-granulate patch; ductus seminalis
from bursa near attachment of ductus bursae.
TYPE LOCALITY: France (type in Paris Mus.).
Foop pLants: Fruits of Chaenomeles japonica, pome-
eranates, oranges, citron, grapes, raisins, etc. (often
fallen and desiccated fruit), onion seeds, leaves and
flowers of Daphne gnidium, flowers of Ricinus communis,
green corn stalks (reared moth, in USNM, from Hawaii),
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 11
young stems of Tamarix and the stems, leaves, and
flowers of Lythrum. Apparently has a various larval
habit and a wide variety of hosts. A moth Gn USNM)
from Muar Johore, Malaya (Clausen), was reported as
reared from a larval predator on Aleurocanthus.
Distrisution: Mediterranean countries of Europe,
Africa, and Asia. Presumably widely distributed in
the east and among the Pacific islands; but some of the
published records may apply to other species. New
World distribution: Brermupa (Jan., Feb., Apr., May).
VeneEzvuELA: El Valle (Aug.). Braz: Séio Paulo
(Feb.).
Genera 2-17: Acrobasis to Chararica
[Venational division B. Hind wing with vein 3 from the lower
outer angle of the cell or (if from before the angle) close to it;
7 and 8 approximate beyond cell, rarely shortly and weakly
anastomosed. Male genitalia with transtilla complete or,
where incomplete (Bertelia, Hypargyria), the elements strongly
developed and with expanded apices. Uncus triangulate, or
hoodlike with rounded apical margin, or spatulate (Birinus).]
2. Genus Acrobasis Zeller
Acrobasis Zeller, Isis von Oken, 1839, p. 176; 1848, p. 606.—
Herrich-Schaffer, Systematische Bearbeitung der Schmet-
terlinge von Europa, vol. 4, p. 99, 1849.—Heinemann, Die
Schmetterlinge Deutschlands und der Schweiz, vol. 1,
pt. 2, p. 175, 1865.—Hulst, Phycitidae of N. Amer., p. 120,
1890; U. S. Nat. Mus. Bull. 52, p. 418, 1902.—Ragonot,
Monograph, pt. 1, p. 85, 1893.—Dyar, Proc. Ent. Soc.
Washington, vol. 10, p. 41, 1908.—Spuler, Die Schmetter-
linge Europas, vol. 2, p. 214, 1910.—Barnes and McDun-
nough, Contributions, vol. 2, p. 221, 1914.—Forbes, Cornell
Mem. 68, p. 614, 1923.—Meyrick, Revised handbook of
British Lepidoptera, p. 397, 1928.—Pierce and Metcalfe,
Genitalia of the British Pyrales, p. 10, 1938.—Bisset, in
Pierce and Metcalfe, op. cit., p. 55, 1938.—Janse, Journ.
Ent. Soc. South Africa, vol. 4, p. 148, 1941. (Type of
genus: Phycis tumidella Zincken (=Acrobasis zelleri
Ragonot); figs. 133, 640.)
Mineola Hulst, Phycitidae of N. Amer., p. 126, 1890; U. S. Nat.
Mus. Bull. 52, p. 419, 1902.—Forbes, Cornell Mem. 68,
p. 618, 1923. (Type of genus: Myelois indigenella Zeller.)
Seneca Hulst, Phycitidae of N. Amer., p. 177, 1890. (Type of
genus: Cateremna tumidulella Ragonot. New synonymy.)
Acrocaula Hulst, Canadian Ent., vol. 32, p. 170, 1900. (Type
of genus: Acrocaula comacornella Hulst. New synonymy.)
Tongue well developed. Antenna pubescent; on
male, basal segment enlarged and angulate, the shaft
with a slight sinus at base. Labial palpus upturned,
reaching to or a trifle above vertex; third segment
slightly more than half the length of second, acuminate.
Maxillary palpus filiform. Forewing smooth or with
transverse antemedian ridge of raised scales; 11 veins;
vein 2 from well before the angle of the cell; 3 rather
well separated from 4, but somewhat nearer to 4 than
to 2; 4 and 5 closely approximate at base or connate
(rarely, in individual specimens, shortly stalked);
6 from below upper angle of cell, straight; 10 from the
cell, usually (except in individual specimens) separated
from stalk of 8-9 at base; male without costal fold.
Hind wing with vein 2 from well before lower outer
angle of cell (from outer two-thirds of lower median);
3 from before but near the angle; 4 and 5 from the
angle, connate; 7 and 8 shortly anastomosed beyond
cell; cell about half the wing length; discocellular vein
curved. Eighth abdominal segment of male with
midventral hair tuft.
Male genitalia with apical process of gnathos a simple,
elongate hook, or an elongate trifurcate hook. Trans-
tilla complete, sharply angulate and reaching as far
back as base of apical process of gnathos; terminal
margin narrow and indented. Uncus broadly triangu-
late. Harpe simple.’ Anellus a narrowly sclerotized
U- or V-shaped plate. Aedeagus simple, rather stout;
penis with numerous sclerotized wrinklings, otherwise
unarmed. Vinculum stout, a trifle longer than broad,
slightly tapering; terminal margin truncate and more
or less concave.
Female genitalia with ductus bursae and bursa popu-
latrix membranous except for a dorsal sclerotized plate
in genital opening; ductus and bursa more or less scobi-
nate; signum, if present, consisting of a granulate cup
or & minute central spine surrounded by a dense cluster
of scobinations; ductus seminalis from a lobe of bursa
near junction of bursa and ductus bursae.
Acrobasis as here defined is something of a composite
genus, dividing into two distinct groups on the develop-
ment of the apical process of gnathos. Typical Acro-
basis, comprising the European species with both
smooth-winged forms (including the type, twmidella)
and those with the raised-scale ridge on forewing, and
all smooth-winged American species (formerly under
Mineola), have the apical process of gnathos produced
as a simple, elongate hook (fig. 133b). All our Ameri-
can species with the raised-scale ridge on forewing
(except minimella Ragonot) have the apical process of
gnathos trifurcate, that is, produced as an elongate
hook with a lateral projection from each side before
apex. This latter group is strictly North American,
limited in distribution to the United States, and Canada
east of the Rocky Mountains (except for A. betulella
Hulst). As far as I know there are no Old World species
with a similar gnathos. I had hoped to distinguish
this distinctly American group as a separate genus under
one of Hulst’s available names (Seneca or Acrocauwla) on
the basis of the trifurcate projection of gnathos and the
raised-scale ridge on the forewing; but minimella Rago-
not prevents this, as it falls between the two groups,
having the raised-scale ridge and the simple projection
of gnathos. The females offer no characters that will
serve to differentiate the groups. Their genitalia are
so similar that they cannot be used, in many cases,
even for specific separation, much less for group division.
The males of the typical American group with raised-
scale ridge are somewhat more variable than the females,
exhibiting slight differences in the shape of the transtilla
and apical projection of gnathos. Figures of these
structures are given, for what they are worth, for all the
species represented by authentic males. The differ-
ences are comparative only, and I suspect that, when
extended series of the several species are available, they
12 UNITED STATES NATIONAL MUSEUM BULLETIN 207
will prove to be more individual than specific in
character.
The reasons for sinking Seneca and Acrocaula into the
synonymy of Acrobasis are given in the discussion of
their types (pp. 22 and 24).
Genus Acrobasis, Species 2-7: A. indigenella to
A. comptella
[Male with apical process of gnathos a simple hook; forewing
smooth.]
2. Acrobasis indigenella (Zeller)
Ficures 3, 134, 641
Myelois indigenella Zeller, Isis von Oken, 1848, p. 867.—Riley,
Fourth annual report on the noxious, beneficial and other
insects, of the State of Missouri, p. 38, 1872.
Phycita nebulo Walsh, Prairie Farmer, p. 308, 1860; Proc.
Boston Soc. Nat. Hist., vol. 9, p. 312, 1863.
Phycis indiginella (Zeller) Weed, in Forbes, Fifteenth report of
the State Entomologist on the noxious and beneficial insects
of the State of Illinois, p. 65, 1889.
Acrobasis indiginella (Zeller) Riley, Canadian Ent., vol. 16,
p. 237, 1884.—Ragonot, Monograph, pt. 2, p. 118, 1893.—
Forbes, Cornell Mem. 68, p. 619, 1923.
Phycita (Acrobasis) nebulo (Walsh) Riley, Fourth annual report
on the noxious, beneficial and other insects, of the State of
Missouri, p. 38, 1872.
Phycita (Acrobasis) nebulo nebulella Riley, Fourth annual report
on the noxious, beneficial and other insects, of the State of
Missouri, p. 42, 1872.
Myelois zelatella Hulst, Ent. Amer., vol. 3, p. 136, 1887.
Mineola indiginella (Zeller) Hulst, Phycitidae of N. Amer., p. 130,
1890.—Quaintance and Siegler, U. S. Dep. Agr. Farmers’
Bull. 1270, p. 49, 1922.—Essig and Keifer, Monthly Bull.
California Dep. Agr., vol. 22; p. 155, 1933.—McDonough,
Check list, No. 6115, 1939.
Mineola indigenella nebulella (Riley) Hulst, Phycitidae of
N. Amer., p. 131, 1890.
Forewing gray-brown densely dusted with white; the
white dusting concentrated on the upper half of wing
and somewhat between subterminal line and outer
margin, forming two strongly contrasted white patches,
one from costa in subbasal area, the other more or less
triangular and extending from costa, between the
transverse dark lines, into cell and including the blackish
discal spots; the whitish terminal area is less sharply
defined and on many specimens somewhat faint; trans-
verse antemedial line curving obliquely from basal third
of costa to middle of inner margin, indicated chiefly by
its outer dark margin, which begins as a conspicuous
black triangle on costa; from inner margin at one-third
a blackish line curves upward to meet the antemedial
line near costa; the area enclosed between them reddish
ocherous; a similar, smaller spot of the same color on
base of inner margin; extreme base of costa blackish;
subterminal line sinuate, bordered inwardly and out-
wardly by dark lines which begin as blackish spots on
costa; from the outer of these a dark band extends
transversely across to the base of antemedial line at
inner margin, somewhat obscured in the dark ground
color on all but the palest and most contrastingly
marked specimens; discal spots at end of cell black,
more or less confluent, usually a black bar along dis-
cocellular vein; a more or less broken, black line along
terminal margin. Hind wing subpellucid, pale smoky
fuscous; the veins very faintly, if at all, darkened;
a narrow, obscure, dark line along termen. Alar ex-
panse, 15-20mm. Male genitalia as given for the genus.
Female genitalia without signum.
Type Locatitizs: ‘‘Carolina”’ (indigenella, in BM);
Illinois? (nebulo, lost); Missouri? (nebulelia, lost); “New
York and Canada,” (zelatella; the supposed type, co’,
in AMNH, ex Rutgers, is labeled “Blanco County,
Texas’’).
Foop piants: Apple, crabapple, plum, prune, cherry,
quince, Crataegus, Cotoneaster, Pyrocantha coccinea.
Larva feeding on leaves and forming serpentine resting
and hibernating case of silk and frass.
Disrrisution: Unirep Starrs: Maine, Sebec Lake
(uly); New Hampshire, Hampton (July); Vermont,
Clarendon; Massachusetts, Newton Highlands; Connec-
ticut, Hast River (July); New York, Catskill Mts., Ilion;
New Jersey, Rutherford; Pennsylvania, Chambersburg
(June, July), Germantown (July), New Brighton (July),
Pittsburgh (June); Virginia, Colonial Beach (July),
Norfolk (May); North Carolina, Tryon (Aug.); Georgia,
Savannah (Apr.); Illinois, Chicago, Decatur (May,
June, July, Aug.); Mssourt, Mossele (June, July),
Norborne (Apr., June, July), St. Louis (Aug.); Nebraska,
Wahoo (May); Kansas, Onaga, Wichita (June); Missis-
sippi, ‘Agr. College” (May, June, July); Arkansas,
Siloam Springs (June), Washington County (July);
Texas, Abilene, Blanco County (Sept.), Fort Worth
(Sept.), Houston (May, June, Aug.), Kerrville, Victoria
(June); California, Lomita, Los Angeles County (Mar.)
Orange County (June). (The California records all
from plum.) Canapa: Ontario, Trenton (June, July);
Quebec, Meach Lake (July).
Apparently generally distributed east of the Rocky
Mountains and rather recently introduced into Cali-
fornia.
This species has been considered of economic impor-
tance as a defoliator of fruit trees in the Middle States,
and in the official list of common names approved by
the American Association of Economic Entomologists
is designated as the “leaf crumpler.”” However, it does
not seem to be more than a minor pest of local and
occasional concern. Several references are made to it
in the economic literature but none of these adds any-
thing of significance to our knowledge of the insect
beyond what is given in the early papers by Walsh and
Riley.
Riley’s nebulella was described by him as a variety
of nebulo and distinguished from the latter by the more
diffused dark shading and the separation of the discal
spots on forewing. In the series before me there is
considerable variation in the extent and intensity of the
dark coloring and the discal mark varies even more,
being sometimes divided into two distinct spots or fused
into a single bar on different sides of the same specimen,
so the varietal designation is hardly worth maintaining.
In 1908 (Proc. Ent. Soc. Washington, vol. 10, p. 45)
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 13
Dyar, on the evidence of a supposed type of nebulella
in the National Collection, resurrected the name and
applied it to the ‘‘pecan leaf casebearer.’’ Barnes and
McDunnough (Contributions, vol. 2, p. 222, 1914;
vol. 3, p. 221, 1917) called attention to the spuriousness
of the alleged “‘type” and gave the pecan casebearer its
proper reference (juglandis LeBaron); but on the
strength of Dyar’s identification the name nebulella
had already appeared, and continued to be used for some
years in economic publications for the pecan leaf case-
bearer. In his 1939 Check List McDunnough applies
the name in an entirely new sense, transferring it to
Meroptera with the well-known unicolorella Hulst as a
syhonym. This was most unfortunate and altogether
unnecessary. We know what unicolorella Hulst is, and
its type is at hand for reference. The type of nebulella
is nonexistent and McDunnough’s new reference has
nothing to back it but an entomologist’s interpretation
of Riley’s description and very poor and over-inked
figure of the forewing. I see nothing in either to rule
out the original interpretation, so shall let the name
sleep in synonymy.
3. Acrobasis grossbecki (Barnes and McDunnough), new combi-
nation
Mineola indigenella nebulella Grossbeck (not Riley), Bull. Amer.
Mus. Nat. Hist., vol. 37, p. 129, 1917.—Barnes and Mc-
Dunnough, Contributions, vol. 3, p. 220, 1917.
Mineola grossbecki Barnes and McDunnough, Contributions,
vol. 3, p. 221, 1917——McDunnough, Check list, No. 6116,
1939.
Forewing purplish brown, the dark ground color
more extended than in indigenella, obscuring the pale
antemedial line and completely obliterating the reddish
ocherous subbasal patch on inner margin usually present
in the genus; triangular black spot on costa, beginning
the outer dark border of antemedial line, distinct and
sharply contrasted as in indigenella; white areas
restricted more than in indigenella, the subbasal one
narrowly triangulate with its point on inner margin,
midcostal one extending to and including the discal
spots in its lower angle; whitish dusting in terminal
area very faint; subterminal line obscure; discal spots
at end of cell black, separated. Hind wing shiny,
smoky fuscous. Alar expanse, 15-16 mm.
Genitalia as in indigenella.
Tyre tocauity: Lakeland, Fla. (type in USNM).
Foop puant: Crataegus (larva feeding on the leaves).
Distrisution: Known only from the type locality.
May be a Florida race of indigenella, but appears to
be a distinct species despite the likeness of its genitalia
to those of indigenella.
4. Acrobasis vaccinii Riley
Fieure 642
Acrobasis vaccinii Riley, Canadian Ent., vol. 16, p. 237, 1884; in
Rep. [U. 8.] Comm. Agr. for 1884, p. 355, 1885.—Smith,
in Rep. [U. S.] Comm. Agr. for 1884, p. 394, 1885.—Saunders,
Insects injurious to fruits, p. 375, 1883 (as ‘“The Cranberry
Fruit-worm”).—Ragonot, Monograph, pt. 1, p. 121, 1893.—
Forbes, Cornell Mem. 68, p. 618, 1923.
Mineola vaccinii (Riley) Hulst, Phycitidae of N. Amer., p. 128,
1890.—Brown, Oregon Agr. Exp. Station Bull. 225, p. 19,
1927.—Crowley, Washington Agr, Exp. Station Bull. 230,
p- 24, 1929——McDunnough, Check list, No. 6114, 1939.—
Beckwith, Journ. Econ. Ent., vol. 34, p. 169, 1941.
Averaging smaller than indigenella; dark ground
color similar but more extended and without the con-
trasting black costal triangle; pale antemedial line
obliterated by a transverse extension of the ground
color, bordered inwardly by an almost vertical, rather
narrow white band which expands narrowly on costa
towards base (the remains of the much-reduced sub-
basal white area); midcostal white patch also much
restricted, barely including at its lower angle the
separated black discal spots; on fresh specimens some
sprinkling of rufous scaling is distinguished under high
magnification, but no reddish or other contrastingly
colored, angulate, subbasal patch on inner margin (as
in indigenella and tricolorella). Hind wing pale smoky
fuscous. Alar expanse 14-18 mm.
Male genitalia differing in no significant detail from
those of indigenella. Female genitalia with bursa
more or less heart-shaped (less elongate than that of
indigenella or grossbecki); signum present as a minute
granulate cuplike patch.
Typr Locauity: Massachusetts (type in USNM).
Foop puant: Cranberry, blueberry (larva in the
fruit).
Distripution: Massachusetts (type series, no exact
locality, June), Wareham (June, July); Connecticut, East
River (July); New Jersey, Pemberton (May), Whitesbog
(June); Wisconsin; Michigan; Georgia; Mississippi,
Biloxi, Poplarville; Washington, Long Beach (June),
Seaview (July).
Presumably generally distributed on the range of its
food plants in the United States and Canada. The
foregoing records are from reared and typical examples
in the National Collection.
This species, popularly known as the ‘cranberry
fruitworm,”’ is of some importance, especially to cran-
berry growers, and has a rather extensive economic
literature, mostly in annual reports, bulletins, and other
publications of state entomologists and experiment
stations. None of these adds anything of biological or
taxonomic significance to the earlier records of Riley
and Smith.
5. Acrobasis amplexella Ragonot
Acrobasis amplexella Ragonot, N. Amer. Phycitidae, p. 3, 1887;
Monograph, pt. 1, p. 97, 1893.—Forbes, Cornell, Mem. 68,
p. 618, 1923.
Mineola amplexella (Ragonot) Hulst, Phycitidae of N. Amer.,
p. 127, 1890.—McDunnough, Check list, No. 6112, 1939.
This is probably nothing but a color form of vaccinii.
I can find no difference from the latter except in the
greater extension of the basal dark area of forewing and
the consequent further restriction of the subbasal white
area which is a narrow band throughout, not expanding
along costa towards base.
Rearing will have to settle the status of amplezella.
In the material before me there are only collected
14 UNITED STATES NATIONAL MUSEUM BULLETIN 207
specimens. All reared examples we have from either
cranberry or blueberry are typical vaccinii.
Alar expanse, 12-18 mm.
Typz xocauity: North Carolina (type in Paris
Mus.).
Foop prant: Unknown.
Distrizution: Maine, Monmouth (June), Sebec
Lake (July); New Hampshire (June, July); Massachu-
setts, Cohasset (July), Framingham (June), Winchendon
(July), Worcester (July) ; Connecticut, Hast River (July);
New York, Liberty (June), Sullivan County (July);
Pennsylvania, Hazleton (May); North Carolina, Tryon
(July).
In the Barnes and old U. S. National Museum Col-
lections these examples were about equally divided
under the two names, amplexella and vaccinii.
6. Acrobasis tricolorella Grote
Figures 135, 643
Acrobasis iricolorella Grote, Bull. U. S. Geol. Geogr. Surv. Terr.,
vol. 4, p. 694, 1878.—Ragonot, Monograph, pt. 1, p. 93,
1893.—Forbes, Cornell Mem. 68, p. 618, 1923.
Mineola tricolorella (Grote) Hulst, Phycitidae of N. Amer., p.
127, 1890.—McDunnough, Check list, No. 6111, 1939.
Mineola scitulella Hulst, Canadian Ent., vol. 32, p. 169, 1900.—
Hungerford, Idaho Agr. Exp. Station Bull. 149, p. 29,
1927; Bull. 164, p. 29, 1929.—Pack and Dowdle, Journ.
Econ. Ent., vol. 23, p. 321, 1930.—Haegele, Journ. Heon.
Ent., vol. 25, p. 1078, 1982.—Essig and Kiefer, Monthly
Bull. California Dep. Agr., vol. 22, p. 153, 1983.—McDun-
nough, Check list, No. 6110, 1939. (New synonymy.)
Forewing grayish fuscous more or less dusted with
whitish scales, in some specimens well diffused over the
middle of wing, giving it a pale slate ground color, but
normally concentrated into a pale patch from costa
before subterminal line, including the discal mark, and
a pale terminal suffusion below apex; usual subbasal
white area constricted into a narrow, sharply defined
antemedial band, outwardly oblique from costa to top
of cell, thence vertical to inner margin and bordered
outwardly on its vertical portion by a more or less
triangular, contrasted orange or reddish orange patch;
bordering the white line and the orange patch out-
wardly, a black oblique angulate line extending to
near middle of inner margin and beginning on costa in
a more or less angulate and diffused blackish patch
(similar to but not so sharply defined nor contrasted as
the black costal patch on indigenella) ; subterminal line
distinct, narrow, white, angled inwardly at vein 6 and
lower fold and curved outwardly between, bordered
inwardly by a narrow black line and outwardly by a
black costal spot and a more or less pronounced orange
or reddish orange band (well marked in many eastern
and western specimens, but sometimes obscured by
dark scaling); discal spots usually fused into a slightly
curved, black bar along discocellular vein, rarely
separated. Hind wing smoky white to pale smoky
fuscous. Alar expanse, 18-22 mm.
Male genitalia with cucullus of harpe of more even
width throughout and apex more evenly rounded than
in other species of the genus. Eighth abdominal seg-
ment of male with a single central, ventral hair tuft
(supplementary ventral hair tufts on the other species).
Female genitalia with signum present as a small,
granulate, cup-shaped patch.
TYPE LOCALITIES: Oldtown, Maine (éricolorella, in
BM); Colorado (scitulella, in AMNH, ex Rutgers).
Foop Puants: Apple, apricot, catalina cherry, plum,
prune (larvae feeding in buds and fruits, probably also
on leaves), also recorded from galls on chokecherry
(Park and Dowdle).
Distrrsution: Unitep Status: Maine, Oldtown,
Bar Harbor (July), Orono; New Hampshire, Hampton
(July); Massachusetts, Amherst (Hatch Exper. Station,
July); New York, Catskill Mts., Ilion (July); New
Mexico, Albuquerque (July); Colorado, Denver, Glen-
wood Springs (May, July, Aug., Sept.), Grand Junction
(Aug.), Gunnison County (July); Utah, Bellevue (May),
Dividend (Aug.), Eureka (Aug.), Logan (July), Park
City (July), Provo (July); Zdaho, Boise (Sept.), Emmett
(July); California, Loma Linda (Aug.), Mount Lowe
(May, July), San Diego (June, July), Santa Barbara
(Aug.), Warner Mts. (Modoc County, July); Oregon
Lake View (Aug), The Dalles (June); Washington,
Prosser (June), Pullman (July), Walla Walla (Aug.),
Wenatchee (Aug.). Canapa: Ontario, Ottawa (July);
Manitoba, Cartwright; British Columbia, Arrowhead
Lake (June).
The species seems to be abundant in our western
states and relatively scarce im the east, to judge by
examples in collections, and has attracted some atten-
tion as a fruit pest in Utah, Idaho, and California.
There is nothing to distinguish western from eastern
specimens and the one detail that Hulst relied upon
for the separation of his scitulella (the presence of
an orange outer border to the subterminal line) does not
hold. It is present in eastern and western examples
and equally variable in both. I am therefore sinking
the name in the synonymy of tricolorella.
7. Acrobasis comptella Ragonot
Figure 646
Acrobasis compitella Ragonot, N. Amer. Phycitidae, p. 4, 1887.—
Hulst, Ent. Amer., vol. 5, p. 156, 1889 (makes synonym of
caliginella Hulst).
Forewing dark gray dusted with white, the white
dusting concentrated on basal area, on costal median
half of wing (forming a pale angulate patch which
includes the discal spots), and in terminal area beyond
subterminal line; in some specimens the white dusting
is more extended, making most of the basal, median,
and terminal areas pale ashy gray; outwardly bordering
basal pale area a black line (narrowing from a shallow
triangulate patch on costa) extends obliquely outward
to top of cell thence vertically to inner margin, bordered
outwardly on vertical part by a triangulate, tawny or
reddish brown patch; subterminal line white, sinuate,
bordered inwardly by a narrow, blackish line and out-
wardly, at costa, by a black smudge; discal spots at end
of cell distinct, separate, black. Hind wing pellucid,
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 15
whitish or pale smoky fuscous. Alar expanse, 14-21
mm.
Male genitalia exhibiting no distinctive specific
characters. Female genitalia with several short, paral-
lel lines of fine scobinations in bursa; signum present as
a small, granulate, cup-shaped patch.
Typr Locatity: California (type in Paris Mus.).
Foop puant: Unknown.
Distrisution: California, Cisco (July), Colfax (July),
Pasadena, San Diego (June), Santa Catalina Isl. (May),
Warners (San Diego County, Aug.); Arizona, Gila
County, Redington; New Mezico, Albuquerque (July),
Las Vegas; Utah, Provo (July).
Hulst in 1889 made comptella a synonym of his
caliginella and it has remained as such in our lists.
However, the two are generically as well as specifically
distinct, caliginella having the basal segment of the
male antenna cylindrical (not triangularly expanded at
apex as in Acrobasis). It is superficially similar in
color and markings to comptella; but the black line
bordering the whitish basal patch of forewing is
distinctly broken, its vertical portion not reaching to
inner margin. I am removing both caliginella and
Mineola supposita Heinrich to Rhodophaea.
Genus Acrobasis: Species 8. A. minimella
[Male with apical process of gnathos a simple hook; forewing
with raised-scale ridge.]
8. Acrobasis minimella Ragonot
Fieure 140
Acrobasis minimella Ragonot, Ent. Amer., vol. 5, p. 113, 1889;
Monograph, pt. 1, p. 105, 1898.—MeDunnough, Check list,
No. 6088, 1939.
Acrobasis nigrosignella Hulst, Phycitidae of N. Amer., p. 123,
Map ee Proc. Ent. Soc. Washington, vol. 10, p. 43,
Forewing grayish fuscous with a purplish suffusion;
some whitish dusting on basal area and, very faintly,
from costa before subterminal line to discal spots;
antemedial line obscure except towards inner margin,
where it is a narrow whitish line; a blackish triangular
costal patch following the antemedial line and continued
as thin black line on its outer border to inner margin;
vertical scale ridge black; area between scale ridge and
antemedial line ocherous or reddish; discal spots at
end of cell small, obscure, separated. Hind wing smoky
fuscous. Alar expanse, 13-16 mm.
Female genitalia exhibiting no specific difference to
distinguish them from those of other species having the
raised-scale ridge on forewing.
TypE LocaLity: Texas (minimella, 9, in Paris Mus.,
and nigrosignella, in AMNH, ex Rutgers).
Foop prant: Oak (this food plant record from speci-
men, in USNM, reared at Falls Church, Va., under
Hopkins No. 9847, C. F. Johansen).
Distrizution: Teras (Apr.); Mississippi, Starkville
(July); Louisiana, Winfield (June); North Carolina,
Southern Pines (Apr., May, June, Aug.), Tryon (May);
Virginia, Falls Church; District of Columbia, Washing-
ton (June); New Jersey, Lakehurst (July).
The species is easily recognized by its size, color, and
male characters. The sex-scaling is present and con-
sists of long broad black costal streaks on underside of
fore and hind wings.
Genus Acrobasis, Species 9-22: A. feltella to A.
demotella
[Male: Apical process of gnathos trifurcate; forewing with raised-
scale ridge; black sex-scaling beneath.]
9. Acrobasis feltella Dyar
Ficure 141
Acrobasis feltella Dyar, Proc. Ent. Soc. Washington, vol. 11,
p. 214, 1910.—Ely, Ins. Inse. Menstr., vol. 1, p. 51, 1913.—
McDunnough, Check list, No. 6080, 1939.
Head, basal segment of antenna, thorax, and basal
area of forewing white; a faint rosy tint on the posterior
of thorax and a more obvious rosy shading on lower
half of pale basal area of forewing (more intense and
extended on the female than on the male); remainder
of wing dark gray-brown (in fresh specimens blackish
brown) with a faint, pale grayish shading in terminal
area and a whitish spot on inner margin near tornus
(the lower end, and contrasted portion of the otherwise
obscure subterminal line); black discal dots at end of
cell distinguishable but somewhat obscured in the dark
ground color, usually separate, but occasionally fused;
antemedial scale ridge blackish. Hind wing of male
white at base, shading to smoky fuscous outwardly;
the veins in both sexes faintly darkened. Alar expanse,
14-18 mm.
Black sex-scaling consisting of a short patch at base
of costa on forewing.
Type tocatity: Warner, N. Y. (type in USNM).
Foop prant: Hickory (larva boring in petioles).
DisrrisuTion: Unirep States: New York, Warner
(July) ; Connecticut, Hast River (July) ; Zilinois, Putnam
County (June). Canapva: Ontario, Merivale (June).
This species, palliolella Ragonot, and caryalbella Ely
are identical in color, maculation, and all superficial
characters. They exhibit trifling differences in their
male genitalia, especially in the shapes of their trans-
tillae and the apical processes of their gnathi. These
differences are probably no more than individual in
character. We figure them for what they are worth.
Ely (1913) noted differences in the larval cases of
feltella and caryalbella which should be significant. He
also saw, or thought he saw, a difference in the sex-
scaling of Dyar’s type and the type of caryalbella. In
this he was in error; for the sex-scaling is identical in
both types and on the males of palliolella and juglandis
as well. I suspect that the three names (feltella,
palliolella, and caryalbella) apply to a single species; but
this cannot be determined until the biologies and larvae
of the various hickory-feeding forms of the genus are
more thoroughly studied. Until that is done it seems
best to keep the names separated.
16 UNITED STATES NATIONAL MUSEUM BULLETIN 207
10. Acrobasis palliolella Ragonot
Ficgure 142
Acrobasis palliolella Ragonot, N. Amer. Phycitidae, p. 4, 1887;
Monograph pt. 1, p. 92, 1893.—Hulst, Phycitidae of N.
Amer., p. 121, 1890.
Acrobasis albocapitella Hulst, Ent. Amer., vol. 4, p. 116, 1888.
Not distinguishable superficially from feltella. Tri-
fling differences in the male genitalia are shown in the
figure. They are probably not significant.
Type tocauities: North America (palliolella, in
Paris Mus.); Canada (albocapitella, in AMNH, ex
Rutgers).
Foop pLant: Presumably hickory. Life history not
known.
Distrisution: Unirep States: Illinois, Chicago
(July); Pennsylvania, New Brighton (July); North
Carolina, Plymouth (May); Connecticut, East River
(July). Canapa: Ontario, Ottawa (July).
The name palliolella has been variously misapplied
and has appeared frequently in economic literature for
the ‘‘pecan leaf casebearer”’ (juglandis LeBaron). In
our latest checklist (McDunnough, 19389) it appears as
a synonym of juglandis but I do not think this is correct.
A long series of juglandis before me shows consider-
able variation in color but at the same time consistent
differences from palliolella, whose closest affinities are
feliella Dyar and caryalbella Ely.
11. Acrobasis caryalbella Ely
Fiaure 143
Acrobasis caryalbella Ely, Ins. Insc. Menstr., vol. 1, p. 52,
1913.—McDunnough, Check list, No. 6081, 1939.
Acrobasis angusella Dyar (not Grote), Proc. Ent. Soc. Washing-
ton, vol. 10, p. 42, 1908.
Ely distinguishes his species from Dyar’s feltella
chiefly on the differences in their larval cases (‘‘co-
coons”). There is nothing else to separate them except
some slight and probably not significant differences in
their genitalia. These are shown in the figure.
Typxz Locauity: East River, Conn. (type in USNM).
Foop puant: Hickory.
Known only from reared examples from the type
locality and the female from hickory (June) without
locality label, bearing Riley’s No. 376 and referred by
Dyar to angusella Grote. The sex-scaling on the male
is the same as that on feltella and palliolella.
12. Acrobasis juglandis (LeBaron)
Fieurrs 138, 644
Phycita juglandis LeBaron, Second annual report on the noxious
insects of the State of Illinois, p. 23, 1872.
Acrobasis juglandis (LeBaron) Riley, Fourth annual report on
the noxious, beneficial and other insects of the State of
Missouri, p. 42.—Barnes and McDunnough, Contributions,
vol. 3, p. 221, 1917—McDunnough, Check list, No. 6082,
1939.—Moznette (and others), U. S. Dep. Agr. Farmers’
Bull. 1829, p. 16, 1940.—Craighead, U. S. Dep. Agr. Misc.
Publ. 657, p. 449, 1950.
Acrobasis nebulella Dyar (not Riley), Proc. Ent. Soc. Washing-
ton, vol. 10, p. 45, 1908.
Aerobasis palliolella Dyar (not Ragonot), Proc. Ent. Soc. Wash-
ington, vol. 10, p. 44, 1908.—Forbes (in part), Cornell
Mem. 68, p. 617, 1923.
Similar to the three preceding species except: White
basal area more or less shaded with ashy gray; without
rosy tints on lower basal and outer areas; generally
paler in outer area, mouse gray, with the white dusting
from midcosta somewhat more intense; a distinct
blackish costal triangle following the antemedial line.
Hind wing smoky fuscous shading to rather dull white
towards base on the male, darker and more uniformly
colored on the female. Alar expanse, 14-17 mm.
TypE Locairy: Illinois (type lost).
Foop puiants: Hickory, pecan, walnut, butternut
(larvae feeding on leaves, buds, and flowers).
Distrisution: Iilinois, Chicago (July); Missouri;
Mississippi, Wiggins (May); Texas, Black Springs,
Brownsville (May), Cuero (June), Kerrville (May,
June), Victoria (May, June); Georgia, Albany (July),
Atlanta, Blackshear (May, June), Cairo (May, June);
Florida, Monticello (May, June), Orlando (May),
Palatka (May), Tallahassee (May); South Carolina,
Mt. Pleasant (July); North Carolina, Edgecombe
County (May), Plymouth (May); District of Columbia,
Washington (June).
A large reared series in the National Museum is
mostly from pecan. Also before me a series reared
from walnut and butternut that appears to be a suffused,
dark form of juglandis. Two specimens of the latter
series are from Ontario, Canada.
In our Gulf States the species is of some importance
as a defoliator of pecan, and is popularly known as
the ‘‘pecan leaf casebearer.”” It has numerous refer-
ences in economic literature. I have retained only one
of these (Moznette, 1940), for it gives all the biological
information available on the species under its correct
specific name. Dyar’s unfortunate identifications have
greatly confused the nomenclature, with the result that
most economic references previous to 1939 are under
nebulella or palliolella. Hulst (Phycitidae of N. Amer.,
p. 131, 1890) and Ragonot (Monograph, pt. 1, p. 120,
1893) are also at fault in applying the name juglandis.
Their descriptions apply to examples of indigenella and
not to the “pecan leaf casebearer.”’
The sex-scaling of juglandis is like that on feltella.
13. Acrobasis sylviella Ely
Fieure 144
Acrobasis sylviella Ely, Proc. Ent. Soc. Washington, vol. 10,
p. 161, 1908.—McDunnough, Check list, No. 6094, 1939.
Forewing pale ashy gray; the basal area, thorax, and
head but slightly paler, not contrastingly whitish; no
triangular black spot on costa outside the antemedial
line, the dark outer border of the antemedial line a
narrow band or weak, diffused shade from costa. Hind
wing pale smoky fuscous on male, slightly darker on
female. Alar expanse, 19-21 mm.
Typr Locauity: East River, Conn. (type in USNM).
Foop PiLant: Ostrya.
Distrisution: Unirep Srares: Connecticut, East
River (July); Pennsylvania, New Brighton (May).
Canapa: Ontario, South March (June).
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 17
The Pennsylvania specimen was in the Barnes
Collection as cirroferella Hulst. This is incorrect,
however, as the male type of cirroferella is without
sex-scaling. The Canadian specimens (one male and
one female) were reared from Osirya and are responsible
for the food plant record. They were tentatively
identified by McDunnough as sylviella and I think
correctly. They are considerably darker than the type
series of the Pennsylvania specimen; but the fact that
they were reared, and probably under excess moisture,
would easily account for the difference.
The black sex-scaling of sylviella is similar to that of
feltella but slightly more extended, reaching slightly
beyond basal fourth of costa on the underside of fore-
wing.
14, Acrobasis kearfottella Dyar
Figure 145
Acrobasis kearfottella Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 34, 1905.—Ely, Ins. Inse. Menstr., vol. 1, p. 53, 1913.—
McDunnough, Check list, No, 6079, 1939.
p- ‘The most distinct and strikingly marked of the Ameri-
can Acrobasis species; costal half or third of basal area
of forewing snow white, this white area extending out
along costa to subterminal line and broadening to in-
clude the blackish discal spots; midcostal margin nar-
rowly black-edged; white area uncrossed at any place
by dark lines; subterminal line faint, but distinct, dull
white; remainder of forewing dark gray-brown. Hind
wing of male white shading to fuscous at apex and
terminal margin; hind wing of female pale glossy brown
throughout. Thorax and head of male show white; of
female concolorous with dark area of forewing. Alar
expanse, 18-21 mm.
TYPE Locatity: Cleveland, Ohio (type in USNM).
Foop piant: Hickory (larvae feeding on the leaves).
Distrisution: Ohio, Cleveland (June); New York,
lion (July); Connecticut, East River (July); Pennsyl-
vania, New Brighton (July); North Carolina, Black
Mountain; Jilinois, Oconee (Aug.).
Very little is known of the life history. Ely’s paper
describes the cocoon. The black sex-scaling on the
male is similar to that of juglandis and the preceding
hickory-feeding species.
15. Acrobasis caryae Grote
Fiaures 1387, 146
Acrobasis caryae Grote, Papilio, vol. 1, p. 18, 1881; Bull. U. S.
Geol. Geog. Surv. Terr., vol. 6, p. 591, 1882.—Hulst,
Phycitidae of N. Amer., p. 122, 1890.—Ragonot, Mono-
graph, pt. 1, p. 105, 1893——Barnes and McDunnough,
Contributions, vol. 2, p. 222, 1914.—Forbes, Cornell Mem.
68, p. 617, 1923—McDunnough, Check list, No. 6100,
1939.—Moznette and others, U. S. Dep. Agr. Farmers’
Bull. 1829, p. 2, 1940.—Craighead, U. S. Dep. Agr. Misc.
Publ. 657, p. 449, 1950.
Acrobasis caryaevorella Dyar (not Ragonot), Proc. Ent. Soc.
Washington, vol. 10, p. 44, 1908.
Acrobasis hebescella Dyar (not Hulst), Proc. Ent. Soc. Washing-
ton, vol. 10, p. 44, 1908.
Forewing glossy gray (in southern specimens from
pecans pale and with little darker shading except
narrowly along antemedial line); basal area concolorous
with median area except in some of the darker speci-
mens; northern specimens from hickory normally dark
grayish fuscous; antemedial line whitish towards inner
margin; raised-scale ridge black, preceded by some white
scaling and followed by a narrow, more or less obscured,
flesh-colored patch; subterminal line pale gray, obscure;
discal dots distinct and separate but not strongly
contrasted against ground color. Hind wings smoky
fuscous. Alar expanse, 18-20 mm.
Genitalia exhibiting no distinguishing specific char-
acters; figured from southern male reared from pecan
nut. The scale tufting on the eighth abdominal
segment of the male consists of a single, rather long,
central ventral tuft like that shown in figure 137 and
similar to that of the European type of the genus
(tumidella).
TypeE Locatity: Illinois (type in BM).
Foop piants: Hickory, pecan (overwintering larvae
feeding in early spring upon opening leaves and in the
stems of new growth; later generations in the nuts.
Larva does not make a case during feeding period).
Disrrisurion: Unrrep States: Florida, Monticello
(June, July, Aug.), Tallahassee (May) ; Georgia, Albany
(July); Mississippt, Goodman (July), Ocean Springs
(May, Sept.), Pascagoula (June), Wiggins (June, July) ;
Texas, Boerne (June), Bosque (May), Brownwood
(Apr., May, June, July, Aug., Sept.), Colorado River
(Apr., May), Cuero (Aug., Sept.), Dallas (May), Fort
McKevett (June), Pecan Bayou (July), Pioneer (Aug.),
San Saba (May), Texas A. and M. College Station
(June, July), Victoria (June, July, Sept.); Illinois,
Chicago, (July), Decatur (June); Pennsylvania, New
Brighton (July, Aug., Sept.); North Carolina, Mill
Brook; District of Columbia, Washington (May, June) ;
Connecticut, East River (July, Aug.). Canapa: On-
tario, Merivale (June).
This is the “pecan nut casebearer’”’ of economic
literature. It has a rather extended literature but is
of importance only as a pecan pest in the Gulf States.
Most of the economic references before 1929 are to
hebescella and caryaevorella as a result of Dyar’s mis-
identification of those species. I cite only one economic
reference here, as the Moznette (1940) paper gives all
the biological information available on the species as
@ pecan insect. Its biology as a hickory insect in the
north is imperfectly known.
The sex-scaling of the male is characteristic, consist-
ing of a short black patch on base of costa of forewing
(as in feltella) and a long black streak along the top of
cell on the underside of forewing. This combination
is peculiar to caryae and evanescentella.
16. Acrobasis evanescentella Dyar
Fieure 147
Acrobasis evanescentella Dyar, Proc. Ent. Soc. Washington, vol.
10, p. 44, 1908.—McDunnough, Check list, No. 6086, 1939.
Doubtfully distinct from caryae. The dark areas of
forewing beyond base have a purplish luster, and the
pale (whitish dusting) is more distinct, forming a pale
18 UNITED STATES NATIONAL MUSEUM BULLETIN 207
grayish white spot on costa before subterminal line
(and extended to include the discal spots); sex-scaling
on underside of wing as in caryae, except that the
streak along top of cell is somewhat obscured by an
overlay of pale scales. Under these it is black (not
“pale gray”as stated by Dyar) and no narrower than
that of caryae.
The only authentic specimens I have seen are those
of the original type series. They are in excellent con-
dition. The other specimens which Dyar later asso-
ciated with them are all typical caryae. The note
with the type lot (‘Chittenden 250”) tells nothing
about the larval habits, so we do not know whether
there are any biological characters to distinguish
evanescentella from caryae. We shall have to hold the
name until the biology is thoroughly studied.
Type wocauity: Orlando, Fla. (May) (type in
USNM).
Foop PLant: Pecan.
17. Acrobasis stigmella Dyar
Fiaure 148
Acrobasis stigmella Dyar, Proc. Ent. Soc. Washington, vol. 10,
p. 43, 1908.—Forbes, Cornell Mem. 68, p. 616, 1923.—
McDunnough, Check list, No. 6089, 1939.
Forewing purplish gray; the basal area very dark
(darker than remainder of wing); scale ridge scarcely
darker, its outer margin edged with dull red; discal dots
obscure; subterminal line very faint. Hind wing of
male pale smoky fuscous with a slight ocherous tint;
of female darker and without the ocherous tint on
upper surface.
On the male the upper surface of the head is yellowish
white, the thorax, purplish gray. On the female the
head and thorax are concolorous, purplish gray.
Alar expanse, 17-21 mm.
Typ tocauity: Fort Lee, N. J. (type lost).
Foon piant: Hickory.
Distrisution: New Jersey; Connecticut, Hast River
(Aug.); District of Columbia, Washington (May, June),
Virginia, Falls Church (June); Illinois, Decatur (May),
Putnam County (July).
In his description Dyar states that it is based upon
two males and one female from ‘‘Fort Lee, N. J., May
1896 (H. G. Dyar)” and one female from ‘‘Hast River,
Conn., Aug. 20, 1906 (Chas. R. Hly).” The female
from Hast River is in the National Collection but there
are no specimens from Fort Lee and none dated 1896.
The specimen (a male in good condition) bearing Dyar’s
type label is one reared by him at Washington, D. C.,
June 19, 1900. This probably was overlooked by him
at the time he prepared his description and not identified
or labeled until later. It is unquestionably stigmella
but, in the light of his published declaration, cannot be
accepted as the type.
The species is quite distinct and easily identified—
especially the males by their contrasting yellow-white
heads against their dark thoraces. The sex-scaling on
underside of the male and the contrasting dark basal
patch of forewing distinguish it from everything else
except aurorella Kly. The sex-scaling consists of a nar-
row black streak on forewing extending for about one-
fourth of costa from base, a strongly contrasted and
rather broad black midcostal streak on hind wing and
some black scaling on the extreme base of the veins of
cell of the hind wing. The underside of hind wing is
otherwise a uniform ocherous white.
The life history is also characteristic. In early spring
(Mar.) the young overwintering larva is found within
the unopened leaf-bud, its presence indicated by a
small round frass lid over the entrance hole. For a
short time the larva feed within the bud, chiefly upon
the bud sheath. When it opens the young leaves are
partially eaten and then the larva enters the new shoot.
Thereafter the entire feeding life and pupal period is
spent within the new growth. The larva makes a
larval case during this generation. The life history of
later summer generations is not known.
18. Acrebasis aurorella Ely
Fieure 149
Acrobasis aurorella Ely, Proc. Ent. Soc. Washington, vol. 12,
p. 67, 1910.—Forbes, Cornell Mem. 68, p. 616, 1923.—
McDunnough, Check list, No. 6090, 1939.
Close to and similar to stigmella but differing markedly
in the ground color of the forewing. The sex-scaling is
the same. ‘The upper surface of the head on the male
is also ocherous white but much duller and less con-
trasted against the dark gray thorax. Forewing with
basal area blackish gray sharply contrasted against re-
mainder of wing; median and outer areas of wing pale
pinkish ocherous or pale gray with a pinkish overcast;
subterminal line extremely faint; discal dots distinct
but not strongly contrasted. Hind wings as in stigmella.
Alar expanse, 19-22 mm.
Typ Locatiry: Washington, D.C. (typein USNM).
Foop prant: Unknown.
Distrisution: Connecticut, Hast River (Aug.); New
York, Ilion (Aug.); New Jersey, Montclair (June);
Pennsylvania, New Brighton (Sept.); District of Colum-
bia, Washington (June). Also two specimens from the
Fernald Collection, without locality and labeled ‘‘demo-
tella Grt.”
19. Acrobasis peplifera Dyar
Fieure 150
Acrobasis peplifera Dyar, Ins. Inse. Menstr., vol. 13, p. 13,
1925.—McDunnough, Check list, No. 6106, 1939
Forewing dark gray with a vinous tint; basal area a
dark wine red; scale ridge black, followed by a red line,
then by a whitish line or triangle on inner margin,
shading into black towards costa; discal dots obscured
in the dark ground color, more or less confluent. Hind
wings pale smoky fuscous, darker in the female than
male; veins faintly outlined by dark scaling. Alar ex-
panse, 13-17 mm.
The male sex-scaling on underside is as follows: On
forewing a rather wide black stripe on basal fifth of
costa; on hind wing a black streak from base along top
of cell, widening out to costa at its middle and putting
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 19
out a thin black branch along lower vein of cell (this
lower streak extending out as far as base of vein 2).
Tyre tocauity: Millbrook, N. C. (type in USNM).
Foop piants: Hickory, pecan.
Distrisution: North Carolina, Elizabeth City (Aug.),
Millbrook (Aug.), New Bern (Aug.), Tryon (Aug.);
Illinois, Putnam County (July); Arkansas, Washington
County (Aug.); Texas, Cuero (July); Georgia, Albany
(June); Florida, Monticello.
Doubtfully distinct from exsulella, but the names had
better be kept separate until more is known about the
biologies of the two color forms.
20. Acrobasis exsulella (Zeller), new combination
Figure 151
Myelois exsulella Zeller, Isis von Oken, 1848, p. 868.
Rhodophaea exsulella (Zeller) Ragonot, Ent. Amer., vol. 5, p. 114,
1889; Monograph, pt. 1, p. 80, 1893.—Hulst, Phycitidae of
N. Amer., p. 120, 1890.—McDunnough, Check list, No.
6076, 1939.
Acrobasis septentrionella Dyar, Ins. Insc. Menstr., vol. 13, p. 13,
1925.—McDunnough, Check list, No. 6107, 1939. (New
synonymy.)
Similar to peplifera except: Averaging larger; basal
area of forewing more decidedly and evenly reddish;
outer area somewhat paler; scale ridge not blackish,
red and concolorous with remainder of basal area, very
weak. Alar expanse, 14-20 mm.
Type LocauitiEs: North America [Georgia?] (ersulel-
la, in Zool. Mus. Univ. Berlin) ; Florida (septentrionella,
in USNM).
Foop piants: Hickory, pecan.
Distrisution: Florida (type, no other locality,
Apr.), Orlando (Apr.); Texas, Brownwood (Mar.);
North Carolina, Raleigh (May); Maryland, Plummers
Isl. (June).
The types of both ezsulella and septentrionella are
females. The former is supposed to be in Berlin.
The figure of it given by Ragonot (Monograph, pl. 5,
fig. 19) is a very good match for Dyar’s species and I
have no hesitation in synonymizing the latter. The
scale ridge is present on forewing but could be easily
overlooked, especially ona female that had been spread.
Even on the unspread and unrubbed females in the
National Collection it is not discernible except under
considerable magnification. The structure is more
prominent on the male.
Our Texas specimens were reared from larvae feeding
on the expanding buds of pecan. Dyar’s paratype from
North Carolina was reared from hickory. We have no
further information on the biology. The sex-scaling
of the male is like that of peplifera, which will probably
prove to be nothing more than a variety or color form
of exsulella.
21. Acrobasis angusella Grote
Figure 152
Acrobasis angusella Grote, North Amer. Ent., vol. 1, p. 51, 1880;
Papilio, vol. 1, p. 14, 1880; Bull. U. S. Geol. Geogr. Surv.
Terr., vol. 6, p. 590, 1882.—Ragonot, Monograph, pt. 1,
p. 104, 1893.—Barnes and McDunnough, Contributions,
vol. 2, p. 221, 1914.—Forbes, Cornell Mem. 68, p. 615,
1923—McDunnough, Check list, No. 6091, 1939.
Acrobasis eliella Dyar, Proc. Ent. Soc. Washington, vol. 10, p.
43, 1908.
Head and thorax whitish clay color, more or less
shaded with pinkish or reddish suffusion (darker on
females than on males). Forewing with basal area
reddish and with some dusting of black scales on darker
specimens; scale ridge black or black with an inter-
mixture of red scales, followed outwardly by a reddish
or reddish ocherous patch narrowing towards costa;
pale antemedial line chiefly indicated on lower half of
wing, obscure, followed on costa by a dark fuscous
triangulate shade; median area gray with a diffused
pale shade surrounding discal spots and extending to
costa; discal dots separate, distinct, but not strongly
contrasted against ground color; subterminal line
distinct, denticulate, preceded by a narrow dark border
and followed in terminal area by a broad reddish or
reddish ocherous suffusion. Hind wing pale smoky
fuscous. Alar expanse, 17-22 mm.
Type tocauities: West Farms, N. Y. (angusella,
in BM); East River, Conn. (eliella, in USNM).
Foop puant: Hickory (larvae boring in the leaf
stems).
DistrisuTion: Unitep States: Massachusetts, North
Adams (Aug.); New York, West Farms, Ilion (July,
Aug.) ; New Jersey (June) ; Pennsylvania, New Brighton
(May, Aug.); Maryland, Beltsville (May) ; Connecticut,
East River (July, Aug., Sept.). Canapa: Ontario,
Ottawa (July).
Barnes and McDunnough (1914) were correct in their
criticism of Dyar’s identification of angusella and in
their reference of eliella to synonomy, but their descrip-
tion of the male sex-scaling is at fault. It is more
correctly described by Ragonot in his monograph. It
consists of a very short black patch on costa of forewing;
a long black streak along the upper vein of cell, expand-
ing almost to the costal edge at middle and terminating
well beyond the end of the cell, and from the base of
this streak a second short black streak along lower line
of cell for about half its length. On some males there
are also a few black scales on vein 1c shortly beyond its
base.
22. Acrobasis demotella Grote
Figure 153
Acrobasis demotella Grote, Papilio, vol. 1, p. 14, 1881; Bull. U. S.
Geol. Geogr. Surv. Terr., vol. 6, p. 590, 1882.—Hulst,
Phycitidae of N. Amer., p. 122, 1890.—Ragonot, Mono-
graph, pt. 1, p. 103, 1893.—Dyar, Proc. Ent. Soc. Washing-
ton, vol. 10, p. 42, 1908.—Barnes and McDunnough, Con-
tributions, vol. 2, p. 221, 1914.—Forbes, Cornell Mem. 68,
p. 616, 1923.—McDunnough, Check list, No. 6092, 1939.
Color and markings in general similar to those of
angusella except: Central area of forewing a uniformly
suffused dark grayish fuscous; basal area a paler red-
dish shade without dark dusting except for some
fuscous smudging of the scale ridge on inner margin;
antemedial line more distinct, dull white and on most
specimens completely indicated to costa; discal dots
20 UNITED STATES NATIONAL MUSEUM BULLETIN 207
and subterminal line much obscured, the latter not
denticulate, and followed by 2@ rather faint reddish
shading, distinct only on dark unrubbed or unfaded
specimens. Hind wing pale smoky fuscous. Alar
expanse, 20-24 mm.
Typ Locauity: West Farms, N. Y. (typein BM).
Foop piant: Black walnut (larvae feeding in buds
and stems).
Distrisution: New Hampshire, Durham; New York,
West Farms, Long Island; Pennsylvania, New Brighton
(June); North Carolina, Black Mountain (June), Tryon
(May); Jliinois, Chicago, Decatur (May), Putnam
County (June); Missouri, St. Louis (June).
The sex-scaling of the male is of the same type as
that of angusella, differing as follows: On forewing the
black costal streak is longer, extending to basal fifth of
costa; the upper streak on hind wing is somewhat
shorter, narrow at base, swelling to an oblong patch at
middle; the streak along lower vein of cell is broader
and longer, extending to the lower outer angle of cell.
Hulst gives a description of the larva and life history
as supplied him by Fernald. Nothing substantial has
been added since then to our knowledge of the biology
of the species.
Genus Acrobasis, Species 23-40: A. latifasciella
to A. tumidulella
[Male: Apical process of gnathos trifurcate; forewing with
Taised-scale ridge; without sex scaling.]
23. Acrobasis latifasciella Dyar
Figure 139
Acrobasis latifasciella Dyar, Proc. Ent. Soc. Washington, vol.
10, p. 45, 1908.—Forbes, Cornell Mem. 68, p. 617, 1923.—
MeDunnough, Check list, No. 6096, 1939.
In color and maculation similar to angusella Grote;
but without black sex-scaling on underside of male
wings. Alar expanse, 18-21 mm.
Type tocauity: New Brighton, Pa. (type in USNM).
Foop puants: Hickory, walnut.
DistripuTion: New York, Ilion (Sept.); Pennsyl-
vania, New Brighton (Aug.); District of Columbia,
Washington; Maryland, Plummers Isl. (Aug.); Illinois,
Putnam County (June).
Nothing is known of the biology except the food
plants.
24, Acrobasis irrubriella Hly
Fiaure 154
Acrobasis irrubriella Ely, Proc. Ent. Soc. Washington, vol. 10;
p. 161, 1908.—Forbes, Cornell Mem. 68, p. 618, 1923.—
MecDunnough, Check list, No. 6095, 1939.
Color and maculation similar to those of wngusella
and latifasciella except: Thorax and basal area of fore-
wing showing little or no trace of reddish scaling;
reddish ocherous band on outer border of scale ridge
narrower and fainter; no reddish shading in terminal
area beyond subterminal line. Alar expanse, 19-21
mm.
Male genitalia with apex of uncus more rounded than
that of latifasciella—at best, a character of doubtful
specific value.
TypELocatity: Hast River, Conn. (typein USNM).
Foop puant: Unknown, probably Carya (hickory or
walnut).
Distrisution: Connecticut, East River (July); Indi-
ana, Mineral Springs (Aug.).
25. Acrobasis normella Dyar
Fieure 155
Acrobasis normella Dyar, Proc. Ent. Soc. Washington, vol. 10,
p. 46, 1908.—Forbes, Cornell Mem. 68, p. 617, 1923.—
McDunnough, Check list, No. 6097, 1989.
Similar to wrrubriella except: Averages smaller; fore-
wing of a more even glossy texture; more white dusting
on basal area, giving it a more decidedly pale gray ap-
pearance; the white dusting from outer half of costa to
and surrounding the discal dots also a trifle stronger.
Alar expanse, 17-19 mm.
Typ Locatity: Kast River, Conn. (type in USNM).
Foop pirant: Presumably hickory.
Represented in the National Collection by 15 speci-
mens from the type locality. In addition, there are
before me two smaller (15 mm.), darker, more suffused
specimens (male and female) from the Barnes collection
that McDunnough had identified as irrubriella. They
were reared from hickory (June) and are, I believe, only
a color form of normella. The male genitalia (fig. 1552)
show a striking departure in the assymetrical and greatly
reduced lateral elements of the trifurcate apical projec-
tion of gnathos, but apparently this is the result of a
deformation of the organ in this particular specimen.
26. Acrobasis malipennella Dyar
Ficure 156
Acrobasis malipennella Dyar, Proc. Ent. Soc. Washington, vol.
10, p. 47, 1908.—Forbes, Cornell Mem. 68, p. 618, 1923.—
McDunnough, Check list, No. 6093, 1939.
Similar to the species following (dyarella) in color and
markings except that the darker areas of forewing are
dark fuscous and lack the intense red suffusion charac-
teristic of the latter. The structural differences noted
by Dyar (the broader, shorter forewing, its broader
cell, the close approximation of veins 2 and 3, different
on each forewing, and the connate condition of 4 and 5
and their remoteness from 3) are due to deformation.
The male type never matured properly and the fore-
wings are not fully developed. I believe the specimen
is nothing but a color form of dyarella and a freak at
that. The male genitalia are very close, differing only
in the somewhat more broadly rounded apex of gnathos,
a, difference of no specific significance in this group. It
is a pity Dyar ever described it and a still greater pity
that we cannot ignore his name; for it will probably
have to replace dyarella Ely which was based on normal
specimens and is represented by types in good condition.
Typr Locauity: East River, Conn. (Aug.; type in
USNM).
Foop prant: Unknown.
Known only from the unique type.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE a1
27. Acrobasis dyarella Ely
Fieurs 157
Acrobasis dyarella Ely, Proc. Ent. Soc. Washington, vol. 12, p. 67,
1910.—McDunnough, Check list, No. 6099, 1939.
Head yellow gray. Thorax much suffused with red
scaling. Basal area of forewing red dusted with whitish
towards costa; scale ridge black, followed outwardly by
a broad red band which extends and diffuses outwardly
towards costa, obscuring and partially obliterating the
blackish costal triangle; red scaling generally scattered
over lower median area of wing; some obscure whitish
dusting on median costal area and about the small,
separated discal spots; subterminal line faint, bordered
inwardly and outwardly, except on costa, by reddish
lines; terminal area and dark markings otherwise, dark
gray. Hind wing glossy, pale smoky fuscous. Alar
expanse, 19-20 mm.
Type Locauity: East River, Conn. (type in USNM).
Foop pirant: Unknown.
Represented only by the male type (Aug.) and female
paratype (Sept.).
28. Acrobasis ostryella Ely
Fiaure 158
Acrobasis ostryella Ely, Ins. Insc. Menstr., vol. 1, p. 54, 1913.—
McDunnough, Check list, No. 6084, 1939.
Similar to dyarella except: Reddish color more gener-
ally diffused over outer areas of forewing, not forming
a strongly accented band following the scale ridge, and
of a purplish red shade; the coastal triangle and scale
ridge contrastingly black. Hind wing dark smoky
fuscous. Alar expanse, 15-18 mm.
Type Locauity: East River, Conn. (type in USNM).
Foop Piants: Ostrya virginiana, Carpinus.
Distrisution: Unirep States: Connecticut, East
River (July). Canapa: Ontario, South March (June).
29. Acrobasis secundella Ely
Figure 159
Acrobasis secundella Ely, Ins. Inse. Menstr., vol. 1, p. 55, 1913.—
McDunnough, Check list, No. 6083, 1939.
Doubtfully distinct from ostryella. The holotype
and one other reared male from the type locality and
two Canadian specimens before me are darker and a
nearly uniformly suffused purplish, with a pale dusting
on basal area of forewing, and about the discal spots a
rather faint and pale gray rather than white. However,
other reared specimens from hazel are a perfect match
for the type of ostryella. Alar expanse, 15-18 mm.
The male genitalia show a trifling difference in the
length of the lateral projections of the apical projection
of gnathos. Ely states that the larval case of secundella
is longer and more slender than that of ostryella; but
the life history needs further investigation before any
such difference can be evaluated.
Type tocauity: East River, Conn. (type in USNM).
Foop piant: Corylus.
Distrisution: Unitep States: Connecticut, East
River (July).
Canapa: Ontario, Merivale (June).
30. Acrobasis coryliella Dyar
Ficure 160
Acrobasis coryliella Dyar, Proc. Ent. Soc. Washington, vol. 10,
p. 47, 1908.—Ely, Ins. Inse. Menstr., vol. 1, p. 53, 1913.—
Forbes, Cornell Mem. 68, p. 618, 1923.—MeDunnough,
Check list, No. 6098, 1939.
Forewing a dull, ratber lusterless gray, paler in the
central area and without any reddish or purplish
shading; following scale ridge on inner margin a faintly
ocherous patch obscured by gray scaling; the scale ridge
and other dark markings blackish, but the usual black
triangle on costa following antemedial line replaced by
a narrow line; the usual transverse dark shade from
inner upper edge of subterminal line to middle of lower
margin; subterminal line bordered inwardly by a narrow
blackish line; discal dots normally confluent, forming a
curved line along discocellular vein of cell. Hind wing
pale smoky fuscous; darker on female. Alar expanse,
17-20 mm.
Type Locauity: Unspecified [New York?] (type in
USN).
Foop piant: Corylus.
Distrisution: New York; Connecticut, East River
(July, Aug.); Massachusetts, Newton Highlands; Jili-
bac Decatur (June, July), Putnam County (June,
July).
In addition to a long series reared from hazel at East
River, Conn., there are three specimens from the
Fernald and Brooklyn Museum Collections (one male
and two females from [llinois) labeled ‘‘Acrobasis
hebescella,” the two females labeled ‘type’; and one
male from Decatur, Ill., which McDunnough had
tentatively identified as A. sylviella Ely.
The species is easy to recognize from its rather uni-
form gray shade and strongly contrasted, short, black
scale ridge.
31. Acrobasis hebescella Hulst
Acrobasis hebescella Hulst, Phycitidae of N. Amer., p. 126, 1890.—
Ragonot, Monograph, pt. 1, p. 109, 1893.—Barnes and
MecDunnough, Contributions, vol. 3, p. 194, 1916.—
MeDunnough, Check list, No. 6085, 1939.
The only authentic representation of this species is the
female type, which is in very poor condition but ap-
parently does not, or did not originally, differ in any
significant detail from coryliella Dyar except as to its
host. It was reared from a cocoon found onoak. This
may or may not be significant. The name is just
another of those that must wait for clarification until
someone shall make a careful and more thorough
study of the life histories of the various Acrobasis
species. Alar expanse, 16.5 mm.
TypE tLocauiry: ‘Jersey pines, June”
AMNH, ex Rutgers).
Foop pLant: Oak.
(type in
32. Acrobasis cirroferella Hulst
Acrobasis cirroferella Hulst, Canadian Ent., vol. 24, p. 60, 1892.—
MeDunnough, Check list, No. 6109, 1939.
The type is a male without abdomen. There is no
sex-scaling. Close to coryliella, but with dark areas of
22 UNITED STATES NATIONAL MUSEUM BULLETIN 207
forewing more brownish gray and the whitish areas
more strongly contrasted; central costal area dis-
tinctly white; dark outer margin of antemedial line
brown, narrow on costa; costa before it white, entire
basal area having some white dusting; discal dots
distinct, separate. Hind wing pale fuscous. Alar ex-
panse, 18 mm.
TypE Locantiry: Austin, Tex. (type in AMNH, ex
Rutgers).
Foop prant: Unknown.
The above description is drawn from the type, which
is worn and faded. I have seen nothing that exactly
matches it.
33. Acrobasis cunulae Dyar and Heinrich
Fieure 162
Acrobasis cunulae Dyar and Heinrich, Proc. Ent. Soc. Washing-
ton, vol. 31, p. 37, 1929.—McDunnough, Check list, No.
6102, 1939.
Forewing pale slate gray; basal area paler; midcostal
area with some faint pale dusting, especially about the
discal spots; scale ridge weak and little or not at all
darker than the ground color of the wing; subterminal
line distinct, with a narrow, dentate, dark inner border,
neither the pale line itself nor its dark border strongly
contrasted; discal dots blackish, separated and rather
conspicuous. Hind wing smoky fuscous. Alar ex-
panse, 20-24 mm.
The male genitalia have what appears to be a dis-
tinguishing specific character in the decidedly broad-
ened lateral elements of the apical projection of gnathos.
TyvE tocatiry: Mobile, Ala. (type in USNM).
Foop piant: Pecan.
Distrizution: Florida, Monticello (May); Georgia,
Cairo (May), DeWitt (May); Alabama, Auburn (May),
Mobile (May); Mississippi, Wiggins (May).
Close to but apparently distinct from caryivorella.
34. Acrobasis caryivorella Ragonot
Fiaure 161
Acrobasis caryivorella Ragonot, N. Amer. Phycitidae, p. 4, 1887;
Monograph, pt. 1, p. 108, 1893.—Hulst, U. S. Nat. Mus,
Bull. 52, p. 419, 1903.—Hill, Florida Ent., vol. 21, p. 12,
1938.—McDunnough, Check list, No. 6087, 1989.—Craig-
head, U. S. Dep. Agr. Misc. Publ. 657, p. 450, 1950.
Acrobasis caryaevorella Hulst, Phycitidae of N. Amer., p. 121,
1890 (misspelling).
Acrobasis conjivorella Hulst, in J. B. Smith, List of the Lepi-
doptera of Boreal America, No. 4262, 1891 (misspelling).
Acrobasis caryae Dyar (not Grote), Proc. Ent. Soc. Washington,
vol. 10, p. 46, 1908.
Forewing dark bluish gray, nearly black; basal area
towards costa (above the scale ridge) and a small tri-
angular area on costa adjacent to subterminal line
powdered with grayish white; scale ridge black, on
some specimens bordered outwardly by a faint ocherous
red patch (especially on specimens reared from hickory) ;
subterminal line pale gray, faint; discal dots distinct,
separate or confluent. Hind wing smoky white to
smoky fuscous, darker on female than on male. Alar
expanse, 19-24 mm.
TypE Locauiry: Missouri (type in Paris Mus.).
Foop puants: Hickory, walnut, pecan. Larvae
boring in the buds and new growth of the stems.
Distrisution: Massachusetts, Melrose (June, July);
Maryland, Beltsville (May, July, Aug.), Hyattsville
(June), Prince Georges County, (June); North Carolina,
“N. Car. Dept. Agr.”; South Carolina, Summerton
(May); Georgia, Albany (May), Barnesville (May,
June) ; Florida, Monticello (May, July), Orlando (Apr.);
Mississippi, Biloxi (Aug.), State College; Missouri;
Texas, Austin (Aug.), Brownwood (May, June),
Menard (June), Victoria, Waco (Apr., May, June);
New Mexico, Carlsbad (Aug.).
The species is of some importance in the Gulf States
as an enemy of pecan and the name caryworella has
appeared several times in economic publications but
nearly always wrongly applied to specimens of caryae
Grote. Specimens of the true caryivorella have also
been identified as caryae on the basis of Dyar’s (1908)
misapplication of the two names.
35. Acrobasis comacornella (Hulst), new combination
Figure 136
Acrocaula comacornella Hulst, Canadian Ent., vol. 32, p. 170,
1900.—McDunnough, Check list, No. 6078, 1939.
Forewing with dark areas glossy, purplish brown;
extreme base of wing dark, followed by a rather narrow
subbasal whitish area; median costal area narrowly
whitish; outer area uniformly dark; subterminal line
obscure, not bordered by darker lines; discal dots
distinct, dark brown, the lower dot twice the size of
the upper; a little red on the antemedial line towards
costa. Hind wing whitish with a faint ocherous
fuscous tint; rather glossy; veins very faintly darkened;
a narrow dark line along termen. Alar expanse, 17 mm.
Detail of male genitalia figured from type.
Type tocatiry: Blanco County, Tex. (type in
AMNH, ex Rutgers).
Foop puant: Unknown.
The male type is obviously an Acrobasis without sex-
scaling. It resembles very much a specimen of caryi-
vorella (in USNM) from Victoria, Tex., except that the
hind wing is paler, the subterminal line less distinct,
and the discal dots larger and more contrasted. I sus-
pect that it is nothing more than a variety of caryi-
vorella.
36. Acrobasis betulella Hulst
Ficurs 164
Acrobasis betulella Hulst, Phycitidae of N. Amer., p. 125, 1890.—
Ragonot, Monograph, pt. 1, p. 107, 1893.—Dyar, Proc.
Ent. Soc. Washington, vol. 10, p. 47, 1908.—Forbes, Cornell
Mem. 68, p. 618, 1923.—McDunnough, Check list, No.
6101, 1939.
Forewing dark gray faintly tinted with reddish violet
and with white dusting on basal area and forming a
triangulate patch from costa before subterminal line,
the white dusting faint (less contrasted than on caryi-
vorella); scale ridge black, without any red bordering
patch or bar; discal dots at end of cell distinct, separate;
antemedial line obscure, almost obsolete; subterminal
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 23
line faint.
20-24 mm.
Male genitalia figured from specimen from the origi-
nal Hulst series in the National Collection (bearing a
Hulst ‘‘type”’ label and presumably a paratype).
Typr Locauity: New York (type, 2, in AMNH, ex
Rutgers).
Foop prant: Betula.
Distrisution: Unirep States: Maine (no further
locality, July), Sebec Lake (July); New Hampshire,
Center Harbor (July), Hampton (June); Massachusetts,
Amherst (June, July), Melrose; Connecticut, Hast River
(July); New York (no further locality, July); Colorado,
Platt Canon (July); California, Siskiyou County.
Canapva: Ontario, Trenton.
There is little or nothing to separate collected speci-
mens of betulella from comtoniella or rubrifasciella except
the complete absence of any reddish outer border to the
scale ridge on forewing (a distinction that does not hold
for all specimens of rubrifasciella) and a more glossy
sheen on the specimens of comptoniella and rubrifasciella
(a comparative difference that is displayed only when
series of the three species are seen side by side). The
thing that really distinguishes betulella is its host plant,
Betula. It also differs from the other species of Ameri-
can Acrobasis with the scale ridge on forewing in that it
has been found in the Rocky Mountain region and in
areas west thereof. The Colorado and California
records are from the specimens in the National Collec-
tion mentioned by Dyar. I have seen no later collec-
tions from any area west of the Rockies.
Hind wing smoky fuscous. Alar expanse,
37. Acrobasis rubrifasciella Packard
Figure 165
Acrobasis rubrifasciella Packard, Ann. Lyc. Nat. Hist. New
York, vol. 10, p. 267, 1873.—Grote, Bull. U. S. Geol. Geog.
Surv. Terr., vol. 4, p. 693, 1878.—Hulst, Phycitidae of N.
Amer., p. 124, 1890.—Ragonot, Monograph, pt. 1, p. 106,
1893.—Forbes, Cornell Mem. 68, p. 617, 1923.—McDun-
nough, Canadian Ent., vol. 65, p. 206, 1933; Check list,
No. 6103, 1939.
Phycis rubifasciella (Packard) Beutenmuller, Canadian Ent.,
vol. 22, p. 16, 1890 (spelling! and larva).
Acrobasis alnella McDunnough, Canadian Ent., vol. 54, p. 36,
1922.
Similar to betulella except that normally there is a
faint, but distinguishable, wine-red bar outwardly
bordering the scale ridge on forewing and that, in
series, the forewing surface has a more glossy appear-
ance. Neither of these differences holds for all speci-
mens; nor are the genitalic differences noted by Mc-
Dunnough (1933), the width of the lateral flanges of
the apical projection of gnathos, reliable as a specific
character. Alar expanse, 20-24 mm.
TypE Locatities: Orono, Maine (rubrifasciella, in
MCZ); Ottawa, Canada (alnella, Canadian Nat. Coll.).
Foop piant: Alnus.
Distrievution: Unitep States: Maine, Orono (June);
New Hampshire, Durham; Massachusetts, Amherst
(June) ; Connecticut, Hast River (July); New York, Cats-
kill Mts.; North Carolina, Black Mts. Canapa: On-
tario, Ottawa (July); Quebec, Meach Lake (July); Nova
Scotia, Truro (July, Aug.).
38. Acrobasis comptoniella Hulst
Acrobasis comptoniella Hulst, Phycitidae of N. Amer., p. 125,
1890.—Ragonot, Monograph, pt. 1, p. 108, 1893.—Dyar,
Proc. Ent. Soc. Wash., vol. 10, p. 46, 1908.—Forbes, Cornell
Mem, 68, p. 618, 1928.—MecDunnough, Check list, No. 6104,
1939.—Craighead, U. S. Dept. Agr. Misc. Pub. 657, p.
450, 1950.
Superficially like rubrifasciella except wine-red bar
bordering scale ridge of forewing always present and
of a more intense and darker shade. Hind wing gen-
erally darker. Alar expanse, 21-25 mm.
Type Locatity: Long Island, N. Y. (type, 9, in
AMNH, ex Rutgers; paratype, 9, from type locality
in USNM).
Foop piants: Comptonia and Myrica.
Disrripution: Unirep Srarss: Maine, Bar Harbor
(July), Kennebunk (July), Mount Desert (July); New
Hampshire, Center Harbor, Durham; Massachusetts,
Melrose (June), North Saugus; Connecticut, East River
(July); New York, Long Island; New Jersey, Bergenfield,
New Lisbon (June); Michigan, Dickinson County.
Canapa: Ontario, Trenton.
The best way to separate comptoniella from the two
preceding species is by rearing from their respective
food plants.
39. Acrobasis myricella Barnes and McDunnough
Fiaurs 163
Acrobasis comptoniella Grossbeck (not Hulst), Bull. Amer. Mus.
Nat. Hist., vol. 37, p. 129, 1917.
Acrobasis myricella Barnes and McDunnough, Contributions,
pee p. 221, 1917.—McDunnough, Check list, No. 6105,
Close to comptoniella Hulst, but superficially quite
different; smaller, white dusting in pale (basal and
central-costal) areas much denser and more contrasted;
dark areas blackish gray with very faint purplish over-
tint ; reddish bar bordering black scale ridge narrow and
inconspicuous; subterminal line distinct, white. Alar
expanse, 17-19 mm.
Typr Locatity: Fort Myers, Fla. (Apr., May; type
in USNM).
Foop puant: Myrica.
Distrisution: Florida.
Except for the type series from the type locality I
have seen only one specimen (a female from Royal Palm
State Park, Fla., Apr. 5, 1929, F. M. Jones, collector)
that can be definitely assigned to the name myricella.
We have, however, in the National Collection a series
of males and females reared by Chas. R. Ely at East
River, Conn., from Myrica cerifera, that are inter-
mediate between myricella and typical comptoniella
(reared from Comptonia), like the former in size and in
the intensity and extent of the white dusting on fore-
wing, but with the subterminal line obscure as in
comptoniella. I think they are only a variety of
comptoniella. Indeed, myricella may prove to be only
94 UNITED STATES NATIONAL MUSEUM BULLETIN 207
a race of Hulst’s species; but with our present knowl-
edge we must retain it as a distinct species. ©
40. Aecrobasis tumidulella (Ragonot), new combination
Fieure 645
cai tumidulella Ragonot, N. Amer. Phycitidae, p. 13,
1887.
Seneca tumidulella (Ragonot) Hulst, Phycitidae of N. Amer., p.
178, 1890.
Hyphantidium tumidulellum (Ragonot) Hampson, in Ragonot,
Monograph, pt. 2, p. 74, 1901.—McDunnough, Check list,
No. 6321, 1939.
This species was based on a single female with the
habitus and raised-scale ridge of an Acrobasis but with
vein 4 absent from hind wing. Bourgogne informs me
that the venation is alike on both hind wings. On the
strength of this venation the species was referred to
Group II of the Phycitinae and made the type of Hulst’s
Seneca. However, I am firmly convinced that the speci-
men is nothing but an Acrobasis with abnormal venation,
another of those freaks that turn up all too frequently in
the Phycitidae. Ihave examined the female genitalia of
the type (figured here) and can find nothing to distin-
guish them from those of caryivorella. I suspect that
tumidulella is nothing more than an abnormal specimen
of caryivorella; but we shall have to await final disposi-
tion of the name until a similar freak male is recovered
from the type locality.
Typr Locatity: Florida (type in Paris Mus.).
Foop prant: Unknown.
3. Genus Rhodophaea Guénée
Rhodophaea Guénée, Ann. Soc. Ent. France, ser. 2, vol, 3, p. 312,
1845; Europaeorum Microlepidopterorum index methodi-
cus... , p. 74, 1845.—Ragonot, Ent. Monthly Mag., vol.
22, p. 19, 1885; Monograph, pt. 1, p. 63, 68, 1898. (Type
of genus; Phycis advenella Zincken; figs. 166, 649.)
Characters of Acrobasis except: Male antenna simple,
basal segment cylindrical, no sinus in base of shaft;
forewing always smooth; vein 2 of forewing from cell
before lower outer angle, but somewhat nearer the
angle than in Acrobasis; male genitalia with apical
process of gnathos an elongate hook (partially divided
on advenella); eighth abdominal segment of male simple
or (on advenella) with midventral hair tuft.
This genus is distinguished from the smooth-winged
species of Acrobasis only by its simple male antenna.
Our two American species do not go any too well with
advenella; the European type of the genus, differing in
having an undivided apical projection from gnathos
and simple eighth abdominal segment. However, in
these characters they agree with otber obviously con-
generic European species, marmorea (Haworth), lega-
tella (Hiibner), suavella (Zincken). R. advenella has a
somewhat differently shaped transtilla from caliginella,
supposita, and the three aforementioned Huropean
species. In all of these the terminal margin of trans-
tilla is more or less indented (as in Acrobasis) while in
advenella it is rather deeply U-shaped.
None of the American species that hitherto have been
listed under Rhodophaea belongs there.
They have
entirely different genitalia. .
41. Rhodophaea caliginella (Hulst), new combination
Fieure 647
Nephopteryx caliginella Hulst, Ent. Amer., vol. 3, p. 131, 1887;
vol. 5, p. 156, 1889.
Mineola caliginella (Hulst), Phycitidae of N. Amer., p, 128,
1890; U. S. Nat. Mus. Bull. 52, p. 419, 1902.— Barnes and
McDunnough, Contributions, vol. 4, p. 174, 1918.—
McDunnough, Check list, No. 6113, 1939.
Acrobasis caliginella (Hulst) Ragonot, Monograph, pt. 1, p. 115,
1893.
Myelois caliginoidella Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 33, 1905.—McDunnough, Check list, No. 6072, 1939.
(New synonymy.)
Similar in color and markings to Acrobasis comptella
Ragonot except that the narrow black line outwardly
bordering the whitish basal area of forewing does not
extend all the way to inner margin. This slight dif-
ference in maculation seems to be constant and will
distinguish the females of the two species which, other-
wise, are difficult to tell apart. Alar expanse, 18-25
mm.
TypE LocaLities: Arizona (caliginella, in AMNH,
ex Rutgers); Santa Clara, Calif. (caliginoidella, in
USNM).
Foop puLant: Scrub oak. This host record from
reared specimens in National Collection received from
Commander Dammers, Riverside, Calif., June 1938.
Distripurion: California, Alma (Aug.), Atascadero
(July), Los Angeles County (July), Riverside (June),
San Diego (May, June, July, Aug.), Santa Clara;
Arizona.
In addition to the female type in the Rutgers Col-
lection there is also a female from Arizona (‘‘7810”) in
the National Collection bearing Hulst’s “‘type’’ label.
This specimen was originally in the Fernald Collection.
A female from California donated by Hulst. to the
Brooklyn Museum Collection and twice labeled ‘‘Acro-
basis comptelia” in his and Ragonot’s handwriting is
also in the National Museum. This specimen, except
that it lacks an abdomen, is in good condition. It is
certainly caliginella and presumably was responsible
for Hulst’s synonymizing of caliginella and compiella.
The males of caliginella have hitherto been known as
caliginoidella Dyar. Hulst evidently never saw a male
of his species.
42. Rhodophaea supposita (Heinrich), new combination
Fiaures 167, 648
Mineola supposita Heinrich, Proc. Ent. Soe. Washington, vol.
42, p. 33, 1940.
Forewing very dark grayish fuscous with a powder-
ing of white scales on basal and midcostal areas and
very faintly in the area bordering terman; antemedial
line narrow, slanting from inner third of costa to just
before middle of inner margin, slightly notched at
vein 1b, pale ashy gray bordered inwardly from top of
cell to inner margin by a dull red triangular patch which
has an obscure, straight, blackish line along its inner
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 25
edge; inner margin from base to antemedial line nar-
rowly bordered by reddish scales; subterminal line
narrow, slightly outcurved between vein 6 and lower
fold, pale gray, inwardly bordered by a narrow black
line; a blackish fuscous patch outwardly bordering the
antemedial line from costa to cell; a similar dark shade
on costa near apex; these blackish patches shading into
the dark central area of wing; black discal dots at end
of cell distinct and separate; some obscure dull red
shading in terminal area toward tornus; along termen
a narrow discontinuous black line. Hind wing pale
smoky fuscous with veins, terminal margin, and apical
area darker. Alar expanse, 16-20 mm.
Male genitalia similar to those of caliginella except
transtilla broader at apex and arms of anellus stouter.
Female genitalia differing from those of caliginella
chiefly in that there are no patches of small scobina-
tions in bursa near its junction with ductus bursae.
TypE .Locauity.: Vancouver, British Columbia
(type in Canadian Nat. Coll.).
Foop puant: Cotoneaster.
Known so far only from the type series from Van-
couver. It is distinguished from caliginella chiefly by
its generally darker color.
4, Genus Trachycera Ragonot
Trachycera Ragonot, Monograph, pt. 1, p. 2, 1893. (Type of
genus, Rhodophaea pallicornella Ragonot.)
This genus is close to Rhodophaea, being distinguished
from it chiefly by male characters. The male of
pallicornella from which these were drawn is apparently
lost. Clarke was unable to locate it at Paris, where it
should have been; and, as no other males agreeing with
Ragonot’s description or figure (Monograph, pl. 5,
fig. 20) are available, we are unable to check his charac-
ters. Ragonot separates Trachycera from Rhodophaea
widely in his generic key (Monograph, pt. 1, pp. xliii
and xliv) on the basis of the trifid or bifid condition of
the median vein of hind wing. This is an error, how-
ever, for the true Rhodophaea species are no more bifid
than is Trachycera. The female of pallicornella has
essentially the same venation as the type of Rhodophaea
(advenella).
The distinguishing male characters given by Ragonot
are: Serratiform male antenna; very short labial palpus
(scarcely reaching to middle of face); and minute maxil-
lary palpus.
The female has a pair of small signa in the bursa
copulatrix, developed as granulate cups (as in Davara).
43. Trachycera pallicornella (Ragonot)
Fiaure 650
Rhodophaea pallicornella Ragonot, N. Amer. Phycitidae, p. ,3
1887.—Hulst, Phycitidae of N. Amer., p. 119, 1890.
Trachycera pallicornella (Ragonot), Monograph, pt. 1, p. 2,
1893.—McDunnough, Check list, No. 6057, 1939.
The holotype of pallicornella is a female labeled in
Ragonot’s handwriting ‘‘ty. or. Pl. V, fig. 20.’ Accord-
ing to Clarke the Ragonot figure represents it accu-
300329—56——3
rately. Unfortunately it is a mended specimen and
the glued-on abdomen is spurious. Its genitalia are
pyraustine rather then phycitid.
I have before me a female which is an exact match for
Ragonot’s figure. It is somewhat smaller (15 mm.)
than the type (“19 mm.”) but this difference is easily
within the normal range for many species of medium-
sized Phycitinae. It was collected at Devils River,
Tex. (May). The genitalia are figured from this
specimen.
Forewing pale gray with some blackish dusting on
base, especially on base of costa; a faint purplish gray
shade on lower part of postmedian area; antemedial
band rather broad, red narrowly lined with black on
inner and outer sides, the back outer margin somewhat
widened at costa; subterminal line narrow, nearly
vertical, with an outward bulge between vein 6 and
lower fold, whitish, bordered inwardly by a narrow
black line and by a black outer patch at costa near apex;
discal and terminal dots obsolete. Hind wing dull
whitish with a faint yellow tint and shading to pale
fuscous towards apex. Alar expanse, 15-19 mm.
TypPE Locatity: Texas (type in Paris Mus.).
Foop piant: Unknown.
5. Anabasis, new genus
Typx or Genus: Myelois ochrodesma Zellar.
Tongue well developed. Antenna pubescent; on
male, basal segment enlarged and angulate (as in
Acrobasis), shaft simple. Labial palpus upturned
reaching to vertex (slightly longer on female than on
male). Maxillary palpus rather broadly scaled. Fore-
wing with a transverse, antemedian ridge of raised
scales; 11 veins; vein 2 from before but near lower
outer angle of cell; 3 from the angle, approximate to 2
at base, nearer to 2 than to 4; 4 and 5 closely approxi-
mate for some distance from base; 6 from below upper
angle of cell, slightly bent towards base; 10 from the
cell; male without costal fold. Hind wing with vein 2
from well before lower outer angle of cell; 3 from before
but very near the angle; 4 and 5 from the angle, closely
contiguous (or more or less anastomosed) for about
half their lengths; 7 and 8 contiguous or weakly anasto-
mosed for some distance beyond cell; cell one-third
the length of wing; discocellular vein curved. Eighth
abdominal segment of male with ventral hair tuft and
a pair of modified, ventrolateral tufts.
Genitalic characters as in Acrobasis except: Harpe
with a transverse, sclerotized ridge from base of costa
to lower outer angle of sacculus; a cluster of modified
scales on outer edge of inner margin in the angle be-
tween sacculus and cucullus; terminal margin of
vinculum more rounded. (These may be only specific
characters.)
A development from and quite close to Acrobasis,
which it replaces in tropical America; distinguished
from that genus chiefly by shorter cell and the con-
tiguous position of veins 4 and 5 of hind wing. Except
26 UNITED STATES NATIONAL MUSEUM BULLETIN 207
on denuded wings under strong magnification they
appear to be stalked for half their lengths. Contains
one tropical American species.
44. Anabasis ochrodesma (Zeller), new combination
Ficures 168, 652
Myelois ochrodesma Zeller, Horae Soc. Ent. Rossicae, vol. 16,
p. 209, 1881.
Piesmopoda ochrodesma (Zeller) Ragonot, Monograph, pt. 1,
p. 165, 1893.
Acrobasis crassisquamella Hampson, in Ragonot, Monograph,
pt. 2, p. 520, 1901 (mew synonymy).
Forewing grayish brown finely powdered with
blackish scales and with a faint rosy suffusion; ante-
medial line oblique, narrow, obscured by a heavy whit-
ish ocherous ridge of raised scales on its inner margin
and bordered outwardly by a narrow black line; sub-
terminal line obscure, narrow, when distinguishable,
sinuate, ocherous white, bordered inwardly by an
obscure, broken, black line; discal and terminal dots
obsolete. Hind wing smoky white; veins and a narrow
shade along termen, fuscous. Alar expanse, 13-16 mm.
Genitalia as given for the genus; bursa of female with
small signum.
TypE LocaLities: Honda, Colombia (ochrodesma, in
BM); Teapa, Tabasco, México (crassisquamella, in BM).
Foop piants: Cassia alata, Cassia nodosa, Cassia tora
(U. S. Dep. Agr. Florida rearings; larva a leaf-folder),
Sciacassia siamea.
Distrisution: Unitep Stratis: Florida, Coconut
Grove (May), Miami (May), St. Petersburg (June).
Puxrrto Rico: Bayamén (Sept.), Coamo Springs (Apr.),
Mayagtiez (Jan.), Puerto Real (Vieques Isl., Apr.), Rio
Piedras (Feb.). Virein Istanps: Kingshill (St. Croix,
Oct., Nov., Dec.). Cusa: Santiago de las Vegas
(Havana, Dec.). Grenada. JAMAICA. TRINIDAD:
Fyzabad (Feb.), Tunapuna (Apr.). México: Tabasco,
Teapa. Guatemana: Quirigué (May). Panam:
Corozal (July), Porto Bello (Apr.). Conomspra: Honda.
An easily recognized tropical American species whose
range has been extended into southern Florida.
6. Genus Mildrixia Dyar
Mildrizia Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 405, 1914.
(Type of genus; Mildrixia constitutionella Dyar.)
Tongue well developed. Antenna of male strongly
ciliate, the cilia over three times longer than width of
segments; basal segment elongate, subtubular, flat-
tened and broadening towards apex; first segment of
shaft swollen and with a short spine and scale tuft from
inner upper angle (this with the rough scaling of basal
segment gives the latter when fully scaled the appear-
ance of the triangulate first segment of Acrobasis) ;
antenna of female simple and very weakly pubescent.
Labial palpus obliquely ascending, reaching to slightly
above vertex; moderately rough scaled beneath; third
segment acuminate, about two-thirds the length of
second. Maxillary palpus moderately large, squamous.
Forewing narrowly elongate, with transverse, ante-
medial ridge of raised scales; 11 veins; vein 2 from
before, but near lower outer angle of cell; 3 from the
angle, approximate to 2 at base and for a short distance
beyond; 4 and 5 connate or very shortly stalked; 6
from very close to upper angle of cell, closely approxi-
mate to 8 at base, nearly straight (very slightly bent
towards base); 10 from the cell, closely approximate to
the stalk of 8-9 for some distance from cell; male with-
out costal fold. Hind wing with vein 2 from before
but close to lower outer angle of cell; 3 contiguous with
the stalk of 4-5 for some distance from angle, on
undenuded wings appears stalked with 4-5; 4 and 5
stalked for more than half their lengths; 7 and 8
contiguous or weakly anastomosed for a short distance
beyond cell; cell one-third the length of wing; disco-
cellular vein oblique. Eighth abdominal segment of
male with broad ventral hair tuft.
Male genitalia of the old world Acrobasis type
except: Uncus subtriangulate, its apex rather broadly
rounded; transtilla terminating posteriorly in a U-
shaped projection with elongate, slender, widely spaced
and divergent arms; vinculum longer than broad, evenly
tapering to bluntly pointed terminal margin; anellus
an elongate, semitubular plate with short lateral lobes
near base; penis armed with a short, sclerotized plate
and numerous sclerotized wrinklings.
Female genitalia of the Acrobasis type but without
any sclerotized plate or plates at genital opening; a
single signum in bursa, developed as a small, cupped,
granulate plate.
A distinct genus, distinguished at once by its male
antenna, venation, and transtilla. Contains one
tropical American species.
45. Mildrixia constitutionella Dyar
Figures 169, 651
Mildrizia constitutionella Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 405, 1914.
Forewing grayish fuscous with some whitish dusting,
especially in median area about the discal spots; ante-
medial scale ridge blackish preceded by a narrow white
line and followed by an indistinct dark shade; discal
dots at end of cell distinct, black; just beyond the lower
diseal dot, an outwardly angled mark from the upper
edge of which a narrow dark shade extends to the
inner costal edge of the subterminal line (distinct only
on well-marked and unrubbed specimens) ; subterminal
line narrow, denticulate, pale, bordered inwardly and
outwardly by somewhat broader dark lines; terminal
dots blackish, more or less confluent. Hind wing trans-
lucent, opalescent white, the veins faintly darkened
toward their outer extremities, especially on the females;
a dark shade along costa and a narrow one along termen.
Alar expanse, 19-22 mm.
Genitalia as given for the genus.
Tyrer Locanitry: Jalapa, México (type in USNM).
Foop puant: Unknown.
Distrizution: Méxtco: Jalapa.
cin Santa Maria (June, July, Oct.).
GUATEMALA: Vol-
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 27
7. Genus Sematoneura Ragonot
Sematoneura Ragonot, Nouv. Gen., p. 9, 1888; Monograph, pt.
1, p. 186, 1893. (Type of genus: Sematoneura atrovenosella
Ragonot.)
Tongue well developed. Antenna of male shortly
ciliate (cilia a trifle longer than width of shaft), of
female weakly pubescent. Labial palpus upturned,
reaching vertex, cylindrical, slender; third segment
about two-thirds length of second, acuminate. Max-
illary palpus filiform. Forewing smooth; 11 veins;
vein 2 from well before lower outer angle of cell;
3 from the angle, much nearer to 4 than to 2; 4 and 5
closely approximate for a short distance from cell;
6 from well below upper angle of cell, straight; 8 and 9
long stalked; 10 shortly stalked with 8-9; male without
costal fold. Hind wing with vein 2 from well before
outer angle of cell; 3 from before, but near the angle;
4 and 5 closely approximate for a short distance from
the angle; 7 and 8 closely approximate or contiguous
beyond cell; cell about half the length of wing; dis-
cocellular vein curved. Eighth abdominal segment of
male with a single, broad ventral hair tuft.
Male genitalia with apical process of gnathos a
simple, elongate hook, slightly notched at apex. Uncus
broadly triangulate. Transtilla complete, stout, arched,
its central area developed as a flat, broad lobe with
slightly concave terminal margin. Harpe with costa
sclerotized and produced at apex into a short project-
ing digitus. Anellus a slightly curved plate with
moderately long lateral arms. Aedeagus simple; penis
armed with a single elongate, moderately stout cornutus,
about one-third as long as aedeagus. Vinculum stout,
as broad as or a trifle broader than long, tapering to
broad, truncate terminal margin.
Female genitalia with bursa and ductus bursae
simple, without signum, smooth except for minute
granulations in bursa; ductus bursae shorter than bursa;
genital opening simple; ductus seminalis from bursa
near its junction with ductus bursae.
The foregoing description is drawn from the type
species (atrovenosella). In male genitalia the new
species (abitus), tentatively included in the genus,
departs in some apparently essential details of structure,
having a different type of transtilla and gnathos and
lacking the apical projection from costa of barpe; but
in all other structural characters it agrees with atroveno-
sella. When its female is discovered a new generic
placement may be necessary.
46. Sematoneura atrovenosella Ragonot
Fieures 4, 171, 653
Sematoneura atrovenosella Ragonot, Nouv. Gen., p. 10, 1888;
Monograph, pt. 1, p. 136, 1898.
Forewing gray with more or less ochraceous dusting
above inner margin and in outer area between the veins;
the veins conspicuously outlined by blackish scaling; a
similar narrow, dark line along the lower fold; these
dark lines expanded and intensified at basal third
indicating the remains of an antemedial band, and
broken in outer area by a rather broad, faint, pale
subterminal band; lower discal dot at end of cell
faintly indicated; a line of blackish dots along termen
between the vein ends. Hind wings dusky white,
translucent; the veins darkened and a narrow dark
line along termen. Alar expanse, 26-35 mm.
Genitalia as given for the genus.
Typn Locatity: Chanchamayo, Pert (type in Zool.
Mus. Univ. Berlin).
Foop prant: Unknown.
DistrisuTion: México: Coatepec. Costa Rica:
Huan Vifas (Jan., Feb., Nov.), Tuis (May). Conom-
BIA: La Selva (San Juan Chaco, Sept.), Juntas (San
Juan Chaco, Feb.). Ecuapor: Alpayacu (Rio Pas-
taza), Quito. Prrt: Chanchamayo, Santo Domingo
(Nov.). Argentina: Tucuman.
One example before me (a female from Santo Domin-
go, Peri, 6,000 ft.) differs in coloration from normal
specimens in having a dark suffusion over the basal area
to the antemedial line and a distinct antemedial pale
line with continuous, black outer border. It also lacks
any trace of ocherous dusting on the forewing. In
genitalia and otherwise in color and maculation it is
normal. I believe that it is only a color form. The
specimen was from the unplaced material in the British
Museum.
47, Sematoneura abitus, new species
Fiaurs 172
Similar in color and markings to atrovenosella except a
short strongly contrasted black streak just below costa
at base and a broad black streak along median fold
extending from base to end of cell. The ciliations of the
male antenna are also a trifle shorter than those of
atrovenosella. Alar expanse, 31 mm.
The male genitalia differ markedly from those of
atrovenosella in several details. Apical projection of
gnathos is a rather short triangulate, pointed hook.
The costa of harpe is broadly sclerotized but lacks the
projecting digitus at apex. The gnathos is developed
into a strongly sclerotized, hairpinlike, backwardly
projecting loop with dense scobinations along its inner
margin.
Typs Locatity: Alpayacu, Rio Pastaza, East Ecua-
dor (type in BM).
Foop prant: Unknown.
Described from unique male type collected by M. G.
Palmer at 6,000 ft. The specimen is not in good con-
dition but the essential features of the pattern are
distinguishable and the male genitalia are so distinctive
that description seems justified. In the absence of a
female the generic placement cannot be made with
absolute certainty. I expect, however, that the female
genitalia will exhibit no radical difference from those of
the type of the genus.
8. Genus Hypsipyla Ragonot
Aypsipyla Ragonot, Nouv. Gen., p. 10, 1888; Monograph, pt. 1,
p. 187, 1898. (Type of genus: Hypsipyla pagodella Ragonot,
synonym of Magiria robusta Moore; India; figs. 173, 656.)
Tongue well developed. Antenna of male shortly
28 UNITED STATES NATIONAL MUSEUM BULLETIN 207
ciliate (cilia but slightly longer than width of shaft,
except on male of dorsimacula where they are about
twice as long as width of shaft). Labial palpus of male
upturned, reaching vertex, slender; third segment about
half as long as second, acuminate; of female obliquely
ascending. Maxillary palpus filiform (moderately large
in pagodella and grandella, minute in other species).
Forewing smooth; 11 veins; vein 2 from well before
lower outer angle of cell; 3 from the angle, much closer
to 4 than to 2; 4 and 5 approximate for a short distance
from cell, occasionally connate; rarely short stalked; 6
more or less bent towards base and more or less approxi-
mate to upper angle of cell; 10 normally from the cell,
rarely connate or shortly stalked with 8-9; male without
costal fold. Hind wing with vein 2 from well before
outer angle of cell; 3 from before, but near the angle; 4
and 5 normally shortly stalked, occasionally connate,
partially anastomosed or (in some large females) closely
approximate for a short distance from cell; 7 and 8
closely approximate beyond cell; cell half or (males of
grandella) somewhat less than half the length of wing;
discocellular vein curved. Highth abdominal segment
of male simple or with two or three pairs of ventral hair
tufts.
Male genitalia with apical process of gnathos a hook,
forked at apex. Uncus more or less triangulate; apex
rounded. ‘Transtilla complete, stout, arched, its cen-
tral area produced into two widely spaced horns; the
latter stout in all species except pagodella. Harpe with
costa, strongly sclerotized but not produced at apex.
Anellus V- or U-shaped with long lateral arms. Aedea-
gus simple; penis armed with a single, more or less
twisted, fattened bladelike cornutus (except dorsima-
cula). Vinculum stout, short or but slightly longer
than broad, with truncate, broad terminal margin.
Female genitalia with bursa and ductus bursae
simple, unsclerotized except for a narrow band along
ventral margin of genital opening; with or without
signum; when present, the latter developed as a small,
scobinate, cup-shaped plate; ductus bursae shorter
than bursa; ductus seminalis from bursa at its junction
with ductus bursae.
The.genus is very close to and difficult to distinguish
from Sematoneura. It is characterized chiefly by the
bent condition of vein 6 of forewing. In Sematoneura
this vein is always perfectly straight and remote from
8-9 at base. In Hypsipyla 10 is also normally from the
cell and there is frequent stalking of 4 and 5 of hind
wing, neither of these conditions occurs in Sematoneura;
but the venation is so individually variable in Hypsipyla
that it cannot be trusted.
48, Hypsipyla grandella (Zeller)
Fieures 5, 174, 655
Nephoteryx grandella Zeller, Isis von Oken, 1848, p. 881.
Hypsipyla grandella (Zeller) Ragonot, Monograph, pt. 1, p. 189,
1893.—Dyar, Ins. Inse. Menstr., vol. 7, p. 41, 1919.—Monte,
Rev. de Ent., Brazil, vol. 3, p. 281, 1933.
Hypsipyla cnabella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 405,
1914.
Forewing grayish fuscous shaded (especiaily on lower
half of wing) with dull rust-red; veins outlined in black;
antemedial pale line narrow, incomplete, rounded out-
ward at middle and indented at vein 1b, bordered out-
wardly by a narrow, discontinuous, black line; beyond
this in median area between top of cell and vein 1b, an
expanded faint whitish patch; some faint whitish dust-
ing also in the middle-outer area between the veins;
subterminal line faint, indicated chiefly by the intensi-
fied black streaks bordering it inwardly on the veins,
sinuate, deeply notched at lower fold; discal spots
obsolete; terminal black dots between the vein ends
distinct. Hind wing hyaline white with a fuscous
shade along costa, some fuscous shading on the vein
ends and a narrow fuscous line along termen. Alar
expanse, 23-45 mm.
Male genitalia with uncus rather abruptly narrowed
beyond its broad base, the apex narrowly rounded;
apical hook of gnathos narrow, short; paired horns of
transtilla curving outward (away from each other);
vinculum distinctly broader than long, its terminal
margin very broad and but slightly convex, nearly
straight. Highth abdominal segment of male simple.
Female genitalia with signum.
TYPE LocaLitins: Brazil (grandella, location of type
unknown to me) ; Cérdoba, México (cnabella, in USNM).
Foop piants: Cedrela and Swietenia (larva bores it
fruits and branches).
Distripution: Unirep States: Florida, Miami
(Nov.). Mexico: Cérdoba (Feb., Sept.), Jalapa.
GuaTreMaLa: Cayuga (Apr., May, Oct.), Chejel (June),
Quirigué (June). Honpuras: La Cambra (Feb.).
Costa Rica: Avangarez (July), Juan Vifias (Jan.,
Feb., Nov.), San José (Jan.), San de Montes de Oca
(Oct.), Tuis (May, June). Panam: Almirante (Aug.),
Summit (C. Z., Mar.). Purrtro Rico: Cayey (May).
Cusa: Santiago de las Vegas (Apr.). Hartt: Pétion-
ville (June, Dec.). Jamaica. TRINIDAD (Dec.). Con-
ompia: ‘Above Rio Negro.” VEnEzuELA: El Valle
(July), Maracay, Trompillo (July). British GUIANA:
Georgetown (July). Brazit: Aragatuba (Sao Paulo,
Apr.), Baia (Feb.), Campo Bello, Castro (Paranda),
Espirito Santo, Nova Teutonia (May), Santa Catarina
(Aug., Sept.). Paraguay: Sapucay (Oct.), Villarrica
(Sept., Oct.). HEcuapor: Loja. ArGcentina: Tucuman.
Pert: Lima.
Generally distributed throughout tropical America
wherever its food plants occur.
A native American species close to and superficially
similar to the Indina H. robusta; but with different male
and female genitalia. It is apparently of some eco-
nomic importance in the West Indies and South America
as a pest of mahogany and the Cedrela species. Like
many borers it varies greatly in size, and the venation
is more than ordinarily unstable even for a phycitid.
Vein 10 of forewing may be from the cell, separated
from, closely approximate or connate with 8-9 or some-
times shortly stalked with them, Veins 4 and 5 may
be anything from approximate towards base to shortly
stalked. Vein 6 is always slightly bent towards base
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 29
but less so and more remote from the base of 8-9 on
large females than on the smaller females and average-
size males. On the hind wing 4 and 5 are usually
shortly stalked or connate but on some large specimens
are closely approximate for nearly half their lengths
beyond the lower angle of the cell.
49. Hypsipyla ferrealis (Hampson), new combination
Fiaures 176, 657
Crocidomera ferrealis Hampson, Ann. Mag. Nat. Hist., ser. 10,
vol. 4, p. 352, 1929.
Maculation of forewing similar to that of grandella
except: Brownish fuscous, the general color decidedly
more brown than gray, the dull reddish dusting giving
the wing a somewhat rosy brown tint; dark lining on the
veins less conspicuous and more discontinuous; ante-
medial pale line very faint, obsolete on many specimens,
indicated chiefly by its broken black outer margin; the
white spot beyond this black margin (conspicuous in
grandella) absent or indicated only by a faint ocherous
white shade; subterminal line very faint, indicated by a
black shading on the veins along its inner margin, sinu-
ate, rather deeply indentate at vein 6 and lower fold.
Hind wing smoky fuscous with a faint brownish or
ocherous tint, more or less smoky white towards base;
veins darkly outlined. Eighth abdominal segment of
male with three pairs of ventral hair tufts. Alar
expanse, 20-43 mm.
Male genitalia with uncus of the same shape, but
wider than that of grandella; vinculum narrower,
longer than broad; transtilla with horns of central area
divergent, forming a round-bottomed \V, the central
connecting part of the transtilla slender. Female geni-
talia without signum.
TypE Locatity: Sixaola River, Costa Rica (type in
BM).
Foop PLANT:
the seeds).
Distrisution: Costa Rica: Cachi, Cain, Juan
Vifias (May), Puerto Limén, Sixaola River (May,
Sept.), Tuis (May). Conompra: San Antonio (Dec.).
VENEZUELA: Maturaca (Sept.). FRencH GUIANA:
Cayenne, St. Jean Maroni, St. Laurent Maroni.
Trinipap: Caparo. Braziu: Pard.
A distinct species easily distinguished from grandella
by its dark hind wings. The foregoing food plant and
Venezuelan records are from a series of small reared
specimens (20-24 mm.) submitted by Dr. Ballou in
1942. These are not only considerably smaller than
average from the other localities listed; but are some-
what grayer in color. The genitalia, however, are
like those of typical Costa Rican examples. In
venation ferrealis appears somewhat less variable than
grandella. Vein 10 of forewing is always from the
cell and 4 and 5 usually shortly stalked, rarely con-
tiguous for a short distance beyond the cell.
Carapa guianensis (larvae feeding on
50. Hypsipyla dorsimacula (Schaus), new combination
Figures 175, 654
Myelois dorsimacula Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11,
p. 245, 1913.
The female type is badly rubbed and the markings
consequently obscured; ground color bronzy brown;
faint indications of a pale antemedial line rather far
out on wing; fainter indications of a subterminal line;
at end of cell a dark brown spot on discocellular vein
and shortly separated from it a similar spot in cell,
between them a pale spot (this marking at end of cell
seems the characteristic pattern character of the
species) ; below discocellular vein, on lower fold, a short
blackish streak. Hind wing semihyaline, lilacine,
darker towards apex; veins not appreciably darker.
Alar expanse, 40 mm.
Female genitalia without signum.
TypEe Locauity: Sixaola River, Costa Rica (Sept.)
(type in USNM).
Foop prant: Unknown.
In addition to the female type I have before me
what I believe to be a male of the same species from
the Janse Collection, taken at La Selva, San Juan, on
the Chaco slopes of Colombia (4,600 ft., Sept.). It is
as badly rubbed as the type but shows the same charac-
teristic markings at end of cell and on the fold beneath;
ground color of forewing rust-red; hind wing hyaline
white with a very faint ocherous tint; antennal ciliations
longer than on other species of the genus; at least
twice as long as width of shaft. Male genitalia with
stouter gnathos and quite differently shaped transtilla
from previous species, the prongs of transtilla rather
narrowly separated; penis without cornutus. The
venation is alike on both specimens except for vein
10 of forewing, which is from the cell on the female
and short-stalked with 8-9 on the male; 4 and 5 of
fore and hind wings are short-stalked; 6 of forewing
is sharply bent towards base and connate with 8-9.
Eighth abdominal segment of male simple.
51. Hypsipyla fluviatella Schaus
Fiaure 177
Hypsipyla fluviatella Schaus, Ann. Mag. Nat. Hist., ser. 8,
vol. 11, p. 246, 1913.
Forewing long, and narrower in proportion than other
species of the genus; reddish brown, darker brown in
basal area; antemedial line well out towards middle
of wing, indicated by its brown outer border, the
latter out-bent from costa, thence nearly vertical to
inner margin, forming three lunules, defined by narrow,
pale buff inner shadings (the remains of the pale
antemedial line), also preceded on inner margin by some
silvery gray dusting; a narrow, elongate, pale buff
patch on inner margin at tornus; a similar, wider, pale
streak from cell to outer margin, occupying the space
between veins 5 and 8 and bisected longitudinally by
a narrow red-brown streak along vein 6. Hind wing
hyaline white, inner margin rather broadly tinted with
ocherous, and a narrow ocherous line along outer
margin. Alar expanse, 45-46 mm.
Male genitalia with uncus broad throughout, its
terminal margin broadly rounded. Transtilla com-
plete but the central fusion weak; the horns widely
spaced, forming a broad, shallow U. Apical process
30 UNITED STATES NATIONAL MUSEUM BULLETIN 207
of gnathos rather broad, oval, flattened. Highth
abdominal segment of male simple. Female unknown.
TypE LocaLity: Sixaola River, Costa Rica (type in
USNM).
Foop prant: Unknown.
A distinct species easily distinguished by its narrow,
peculiarly marked, red-brown forewings and the oval,
flattened shape of the apical process of gnathos. The
male hind wings are distinctly triangulate, but this is
probably only a sex character.
The forewing venation is fairly stable; vein 10 from
the cell; 6 bent towards base and narrowly separated
from 8-9 at base; 4 and 5 closely approximate for a
short distance from cell. Hind wing with veins 4 and 5
shortly stalked or closely approximate for some distance
from the cell.
The species is represented only by the type series
of four males from the type locality.
9. Genus Hemiptilocera Ragonot
Hemiptilocera Ragonot, Nouv. Gen., p. 9, 1888; Monograph, pt.
1, p. 144, 1898. (Type of genus: Hemiptilocera chino-
graphella Ragonot.)
Tongue well developed. Antenna of male with basa!
segment elongated, cylindrical; a tuft of scales on base
of shaft (weak on chinographella); shaft unipectinate
for two-thirds, crenulate and pubescent beyond. An-
tenna of female like that of the male type (chino-
graphella) except for lack of scale tuft on shaft and for
shorter basal segment; on other species of the genus
shaft simple and pubescent. Labial palpus ascending;
reaching to or nearly to vertex (shorter on chinographella
than on other species); slender. Maxillary palpus
small, squamous. Forewing smooth; 11 veins; vein
2 from well before lower outer angle of cell; 3 from the
angle, much closer to 4 than to 2; 4 and 5 connate
(chinographella) or closely approximate at base and
for a very short distance from cell (other species of
genus); 6 bent towards base, close to or connate with
8 at base; 10 from the cell, closely approximate to the
stalk of 8-9 for some distance from cell; male without
costal fold. Hind wing with vein 2 from well before
outer angle of cell; 3 from the angle, connate with or
closely approximate to 4-5; 4 and 5 stalked for about
half their lengths; 7 and 8 closely approximate beyond
cell; cell one-half or slightly less than one-half the
length of wing; discocellular vein curved. Eighth
abdominal segment of male with one or more paired
hair tufts and sternal plates modified, one strongly
sclerotized element in the form of an open loop.
Male genitalia with apical process of gnathos a hook
with notched apex. Uncus triangulate, apex bluntly
pointed. Transtilla complete, stout, arched, its central
area produced into widely spaced horns. Harpe with
costa strongly sclerotized (produced at apex on bigrana
and plumigerella, not produced on chinographella).
Anellus with short, stout, lateral arms. Penis armed
with strongly sclerotized cornutus and numerous
sclerotized wrinklings. Vinculum stout, longer than
broad, terminal margin broad and more or less indented.
Female genitalia with signum developed as a small,
strongly scobinate pocket; bursa large; ductus bursae
short, more or less sclerotized and strongly scobinate-
granulate, the scobinations and granulations extending
into bursa for a short distance from place of junction
with ductus; genital opening with sclerotized and more
or less wrinkled plate on lower margin (except on
exoleta); ductus seminalis from bursa near junction of
bursa and ductus bursae.
This genus is distinguished by its pectinate male
antenna, the strong, stalking of veins 4—5 of hind wing,
the long vinculum with broad terminal margin, the
short arms of anellus and its short, granulate-scobinate
ductus bursae. Eventually it may have to be restricted
to its type species (chinographella) and a new generic
placement found for the other species now included.
All of these have simple pubescent female antennae;
while those of chinographella are pectinate in both sexes.
There are also several differences between males of
chinographella and those of plumigerella and bigrana
(notably in the shape of the transtilla, the costal
development of harpe, and the size of the antennal
tuft); but unfortunately we do not know the males of
three other species (letharda, jocarella, exoleta) and until
they are known it seems the wiser course not to attempt
further generic separation. All the species have similar
habitus and wing maculation and (except for exoleta)
female genitalia showing only specific differences.
52. Hemiptilocera chinographella Ragonot
Figures 178, 658
Hemiptilocera chinographella Ragonot, Nouv. Gen., p. 9, 1888;
Monograph, pt. 1, p. 144, 1893.
Male antenna with scale tuft on base of shaft weak,
clay colored. Antenna of female pectinate, the pecti-
nations a trifle shorter than those of the male. Thorax
and basal segment of antenna clay-yellow. Forewing
clay yellow (‘‘olivaceous ocherous” according to
Ragonot) dusted and shaded with dull reddish brown
and dark grayish fuscous, the ground color predomi-
nating in the basal area, along the costa and (more
faintly) bordering the termen and as a narrow longi-
tudinal streak between the transverse line and including
at its middle the lower discal spot; antemedial line
faint, indicated chiefly by the broken elements of its
outer dark border (a short notched blackish streak
slanting outwardly from costa, a blackish spot on top
of cell, another on lower vein of cell and a third on lower
fold, these three blackish spots in a vertical line out
near middle of wing, on a few well-marked specimens
connected by a very faint, twice-outcurved dark line);
subterminal line somewhat stronger, inwardly margined
by a black spot, outwardly margined by a duller dark
shade, more or less accented at costa and on the veins;
discal dots at end of cell separated, distinct, especially
the lower one, black; along termen a row of distinct,
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 31
well contrasted, black dots (these a rather characteristic
feature of most of the species of the genus); on the
female a conspicuous whitish patch on inner margin at
inner edge of the subterminal line (this whitish patch
not present on the males before me). Hind wing
semihyaline, shaded with smoky fuscous towards apex,
on the veins and narrowly along termen. Eighth
abdominal segment of male with a single moderately
long pair of ventrolateral hair tufts. Alar expanse,
22-26 mm.
Male genitalia with central part of transtilla quadri-
form (the horns flattened and with flattened lobes from
their bases, the space between the horns even through-
out); harpe with costa not produced at apex; apex of
cornutus enlarged, sharply bent and bearing a row of
thornlike spines. Female genitalia with ventral plate
at genital opening smooth or nearly so; granulations of
ductus bursae dense and forming a continuous sclero-
tized mass.
Typn Locatity: Chanchamayo, Peri (type in Zool.
Mus. Univ. Berlin).
Foop puant: Unknown.
Distripution: Frencu Guiana: Cayenne. Braziu:
Paré. Prrt: Chanchamayo.
53. Hemiptilocera bigrana (Zeller)
Fieures 180, 660
Myelois bigrana Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 200,
Hemiptilocera bigrana (Zeller) Ragonot, Monograph, pt. 1, p. 145,
1893.
Male antenna with hair tuft strong, black. Antenna
of female pubescent. Thorax and base of antenna dull
whitish dusted with fuscous. Forewing pale gray dusted
with dull rosy and dark grayish fuscous; the dark dust-
ing more evenly distributed than in chinographella;
lower discal spot at end of cell large, strongly contrasted;
upper spot weak or not distinguishable; antemedial line
not defined; subterminal defined by its dark borders,
which consist of blackish streaks on the veins. Hind
wing hyaline white with a faint smoky fuscous shade at
apex, on the outer half of the veins and narrowly along
termen. Eighth abdominal segment of male with a long,
strong pair of ventrolateral hair tufts and two other
pairs of modified scale tufts. Alar expanse, 25-29 mm.
Male genitalia with central part of gnathos a stout
crescent-shaped projection with the horns widely
spaced; harpe with apex of costa produced into a short
spine; cornutus a spatulate ribbed blade. Female with
ventral plate at genital opening deeply wrinkled; ductus
bursae with a central, elongate, irregular patch of scob-
inations, extending into adjacent area of bursa.
Typs Locauity: Honda, Colombia (type in BM).
Foop prant: Unknown.
DistripuTion: México: Guerrero (Aug.), Iguala
(Guerrero, June), Popocatépetl Parks (Distrito Federal,
June). Conomera: Honda.
The Mexican records are from specimens in the U. S.
National Museum.
54. Hemiptilocera plumigerella (Ragonot), new combination
Ficure 179
Nephopteryx plumigerella Ragonot, Nouy. Gen., p. 14, 1888;
Monograph, pt. 1, p. 261, 1893.
This species is known only from the male type. From
Ragonot’s description the forewing color and markings
must be similar to those of chinographella except much
more heavily overshaded with vinous brown. The lower
discal spot is conspicuous as in bigrana and the antenna]
tuft is stout and ochraceous in color. Hind wing iri-
descent, semitransparent, grayish brown. Eighth
abdominal segment of male with a single pair of long,
stout ‘ventrolateral hair tufts and another pair of
much shorter, central tufts. Alar expanse, 21 mm.
Male genitalia with transtilla as in bigrana, except
that the crescent-shaped central projection is much more
slender; cornutus a short bluntly pointed plate with a
row of short, blunt spines near apex.
Type tocatity: ‘America Meridionalis’” (type in
Paris Mus.).
Foop pirant: Unknown.
55. Hemiptilocera letharda (Schaus), new combination
Ficure 662
Chloropaschia letharda Schaus, Proc. Ent. Soc. Washington,
vol. 24, p. 237, 1922; Ann. Carnegie Mus., Pittsburgh, vol.
16, p. 112, 1925.
Forewing olive buff shaded with vinaceous fawn on
subcoastal, median, and basal areas; the wing markings
black, and some faint scatterings of black scales on fore-
wing and thorax; antemedial line indicated only by
broken fragments of its narrow outer border; discal
black spots at end of cell both conspicuous, the lower
one large; black dots on veins forming the inner margin
of the pale subterminal line and black dots along termen
also conspicuous. Hind wing semitransparent, smoky
white, the veins slightly darkened; a dark shade towards
apex and a narrow dark line along termen. Alar
expanse, 29 mm.
Female genitalia similar to those of bigrana but with
a heavier concentration of scobinations in ductus bursae
and a much stronger signum.
Type Locauiry: Cabima, Panamé (May; type in
USN).
Foop piant: Unknown.
In addition to the female type I have before me a
British Museum female from Presidio, México, origi-
nally identified as H. bigrana. It is paler than the
Schaus type, but this apparently is due to its faded and
slightly rubbed condition. Schaus recognized his
original misplacement of the species in the Epipaschiidae
and in his 1925 paper referred it to the Phycitinae, trans-
ferring the specimen to Hemiptilocera in the National
Collection; but I am unable to find any published
reference of his or Dyar’s giving of the generic reference.
32 UNITED STATES NATIONAL MUSEUM BULLETIN 207
56. Hemiptilocera jocarella (Schaus)
Figure 659
Acrobasis jocarella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11,
p. 245, 1913.
Hemiptilocera jocarella (Schaus) Dyar, Ins. Inse. Menstr, vol. 7,
p. 42, 1919.
Forewing reddish brown, the reddish (vinous) shade
predominant in cell and, broadly, along lower fold; a
dull olivaceous shade along costa and in terminal area;
antemedial line obsolete, indicated only by fragments of
its outer border (a black narrow streak from costa to
cell, a black spot on lower margin of cell, and a thin,
in-bent black streak from vein 1b to inner margin);
subterminal line indicated by a pale black margined
spot on costa and an outward series of short whitish
streaks on veins 6 to 1b, these spots inwardly and out-
wardly margined by black dots; a series of black dots
along termen (less distinct than in the other species of
the genus) ; the usual black discal spots, only the lower
one pronounced, and it but slightly so. Hind wing
glossy purplish or smoky brown; the veins but faintly
darkened; a fine dark line along termen. Alar expanse,
21-26 mm.
Female genitalia exhibit (in the amount of scobina-
tion of ductus bursae and the smaller size of signum)
but trifling differences from those of letharda. The male
of jocarella is unknown.
Type Ltocatity: Avangarez, Costa Rica (type in
USNM).
Foop PLANT: Unknown.
Distripution: Costa Rica: Avengarez (July).
Panami: Porto Bello (Dec.). Brazm: Nova Teutonia
(May).
These records from four females in the U. S. National
Museum. When males can be associated it is quite
ikely that jocarella will prove to be only the female
form of plumigerella.
57. Hemiptilocera exoleta (Zeller)
Fieure 661
Myelois exoleta Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 201,
Hemiptilocera exoleta (Zeller) Ragonot, Monograph, pt. 1, p. 146,
1893.
This species is represented only by the female type.
T have seen nothing that matches Ragonot’s description
or Zeller’s rather crude figure; but from both and from
details of the female genitalia the reference is doubtful.
Ragonot himself questions the correctness of his generic
placement; but in the absence of a male no better
placement could or can be made.
The forewing shows the usual distinctively contrasted
row of terminal dots and the maculation otherwise is
that of a Hemiptilocera except that the usual discal dots
are replaced by areddishlunule. According to Ragonot
the cell of hind wing is also short for a Hemiptilocera.
Alar expanse, 25 mm.
The genitalia show a peculiar development of the
eighth segment collar, a central-dorsal, invaginated,
sclerotized pocket flanked by a pair of irregular, elon-
gate, flattened lobes and on dorsum of ovipositor a pair
of shallow sclerotized pockets (fig. 661a). The genital
opening also is unsclerotized, lacking the usual ventral
shield.
Typr tocatity: Honda, Colombia (type in BM).
Foop prant: Unknown.
A male will be needed for certain generic placement.
10. Genus Crocidomera Zeller
Crocidomera Zeller, Isis von Oken, 1848, p. 865.—Ragonot, Mono-
graph, pt. 1, p. 132, 1893. (Type of genus: Crocidomera
turbidella Zeller.)
Tongue well developed. Antenna pubescent; male
with a short, blunt, spine from upper outer angle of
basal segment of shaft; basal segment of male antenna
swollen and broadly scaled. Labial palpus upcurved,
reaching to vertex or slightly above it; third segment
over half as long as second, acuminate. Maxillary
palpus small, squamous. Forewing smooth; 11 veins;
vein 2 from well before lower outer angle of cell; 3 from
the angle; 4 and 5 approximate for a short distance from
cell; 6 from below upper angle of cell, very slightly bent
towards base; 8-9 stalked for about half their lengths;
10 from the cell approximate to the stalk of 8-9; male
without costal fold. Hind wing with vein 2 from well
before lower outer angle of cell; 3 from the angle; 4—5
contiguous for about one-third their lengths beyond cell;
7 and 8 closely approximate beyond cell; cell slightly
less than half the length of wing; discocellular vein
curved. Eighth abdominal segment of male with com-
pound ventral and ventrolateral tufts.
Male genitalia with apical process of gnathos an
elongate hook, blunt and rounded or slightly notched at
apex. Uncus broad, hoodlike, apical margin broadly
rounded or broad and truncate. 'Transtilla complete,
stout, arched, with a strongly forked central projection.
Harpe with costa rather broadly sclerotized, but not
produced at apex, clasper more or less developed, simple,
erect. Anellus with broad, dorsoventrally flattened
lateral arms. Aedeagus with longitudinal rows of
thornlike spines towards apex; penis with sclerotized
wrinklings, but otherwise unarmed. Vinculum stout,
slightly constricted from middle to moderately broad
terminal margin; slightly longer than broad.
Female genitalia with signum developed as a small
scobinate cup-shaped patch; ductus bursae moderately
long (shorter than bursa), expanding gradually to the
wide genital opening and with some strong sclerotized
wrinklings before genital opening; lower margin of
genital opening sclerotized, wrinkled and more or less
finely scobinate; dense, fine scobinations on the dorsal
membrane behind genital opening; ductus seminalis
from bursa near junction of bursa and ductus bursae.
A tropical American genus ranging as far north as our
Texas border and probably into southern Florida; easily
distinguished by its genitalia and male antenna.
e
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 35
58. Crocidomera turbidella Zeller
Ficures 182, 664
Crocidomera turbidella Zeller, Isis von Oken, 1848, p. 865.—
Ragonot, Monograph, pt. 1, p. 132, 1893.
Ground color of forewing variable, pale ocherous gray
or white shaded with faintly reddish or reddish brown on
outer area and in a narrowing shade towards base along
inner margin; an indistinct blackish spot on costa
beyond base; two other blackish dots marking the place
of the obsolete antemedial line, one on costa, the other
at top of cell; a dark patch on inner margin at what
would be the inner margin of the antemedial line, on
well marked specimens containing one or two minute
black dots or dashes; subterminal pale line faint but
distinguishable, indicated chiefly by an inner border of
black spots on the veins and similar, fainter dark streaks
(or a confluent dark shade) bordering it outwardly; a
row of black dots along termen, quite marked on fresh
specimens; discal dots at end of cell small, black, more
or less confluent, when separated the lower not appreci-
ably enlarged; below these, on the fold of some speci-
mens, a larger spot of reddish or brown scales. Hind
wing transparent, opalescent white with a faint fuscous
shading at apex and at the vein ends and a fine dark
line along termen. Alar expanse, 20-25 mm.
Male genitalia with apical process of gnathos bluntly
rounded at apex; uncus subtriangulate, its apical mar-
gin evenly rounded; central projection of transtilla
V-shaped, the prongs divergent and rather slender;
harpe with outer margin of cucullus evenly rounded.
Female genitalia with sclerotization along lower margin
of genital opening narrow and but slightly wrinkled.
Tyrer Locaity: ‘South America” (type in BM).
Foop piant: Unknown.
DistrRipution: Cusa: Baracoa, Santiago (June, Oct.).
Jamaica. M*sxico: Jalapa. Unirep Statszs: Terzas,
San Benito (May, Oct.). These localities from examples
(in USNM) from which the foregoing description was
drawn.
I have omitted the Moschler reference cited by
Ragonot for I suspect that the Puerto Rican specimens
which he and Ragonot had and from which the Ragonot
description was partly drawn are not turbidella but
Jissuralis without the peculiar longitudinal black streak
on forewing characteristic of the type of fissuralis. I
have seen no specimens of Crocidomera from any South
American locality except the Bolivian example men-
tioned in the following discussion of fissuralis. I doubt
that this could be Zeller’s species.
59. Crocidomera fissuralis (Walker)
Figures 183, 665
Nephopteryx fissuralis Walker, List, vol. 27, p. 58, 1863.
Myelois (?) adonea Felder and Rogenhofer, Reise... Novara...,
Lepidoptera, pl. 137, fig. 8, 1874.
Crocidomera fissuralis (Walker) Méschler, Die Lepidopteren-
Fauna von Portorico, p. 327, 1890.—Ragonot, Monograph,
pt. 1, p. 138, 1893.
A photograph of the female type of fissuralis shows a
specimen with a strongly contrasted, black longitudinal
300329—56——4
streak on forewing extending along lower median vein
from base of wing to subterminal line and giving off two
short forks, one along vein 2, the other along vein 3.
The figure of adonea, presumably also a female, shows a
similar marking. I have seen nothing to match this
peculiar pattern except one female from the Janse col-
lection from the Provincia del Sara (Department of
Santa Cruz), Bolivia. The genitalia of this specimen
match those of fissuralis fairly well except for the
sclerotization of the ductus bursae, which is more like
that of stenopteryx. The specimen cannot be placed with
certainty until a male from the type locality is associated
with it. I doubt very much that the peculiar longi-
tudinal streaking represents anything more than an
aberrational or varietal character; for I have before me
a series of three males and three females from Puerto
Rico which lack the longitudinal streak, but are obvi-
ously distinct specifically from what I have recognized
as turbidella. Their female genitalia are identical in all
details with those of the type of fissuralis. On super-
ficial characters they differ from turbedilla chiefly in
having the lower discal spot at end of cell more pro-
nounced and distinctly enlarged in comparison to the
upper discal spot. Alar expanse, 25-30 mm.
Male genitalia with terminal margin of uncus notched ;
apical projection of gnathos slightly notched at apex;
central projection of transtilla with its prongs converging
towards their apices; harpe with cucullus triangulate,
its apex narrowly rounded; aedeagus much stouter and
its thornlike spines stronger and more numerous than
those of turbidella or stenopteryx; terminal margin of
vinculum nearly straight, terminal part of vinculum pro-
portionally about twice as wide as that of either turbi-
della or stenopteryx. Female genitalia with ventral
plate at genital opening deeply wrinkled.
TYPE LOCALITIES: Santo Domingo [Dominican Re-
public] (jissuralis, adonea, both in BM).
Foop prant: Unknown.
DistrisuTion: Dominican REPUBLIC. PuERTO
Rico: Aguirre Central (Apr.), Coamo Springs (Apr.),
Culebra Isl. (Feb.).
60. Crocidomera stenopteryx (Dyar), new combination
Ficures 181, 663
Dioryctria stenopteryx Dyar, Ins. Insc. Menstr., vol. 10, p. 16,
Q
Forewing ocherous gray with blackish markings; a
black dot at base of cell; three black streaks beyond on
lower median vein; a couple of black dots on costa near
base; a black discal spot at lower, outer angle of clel;
antemedial pale line indicated below median vein, pre-
ceded by a dark patch on inner margin (somewhat
tinted with reddish brown on female), margined out-
wardly by an obscure, blackish line, beyond which
on inner margin a somewhat diffused dark shade; vein
1b more or less outlined in black scaling; subterminal
line faint, bordered inwardly and outwardly by some
blackish dots or streaklets on the veins; a fine black line
along termen (formed by the confluent terminal dots).
Hind wing transparent, hyaline white with a faint
34 UNITED STATES NATIONAL MUSEUM BULLETIN 207
grayish shade along costa and a fine fuscous line along
termen. Alar expanse, 25-27 mm.
Male genitalia with terminal margin of uncus broad,
very slightly convex; apical process of gnathos slightly
notched at apex; prongs of central projection of trans-
tilla slightly convergent toward their apices; harpe with
terminal margin of cucullus oblique, apex bluntly
pointed. Female genitalia with ductus bursae smoothly
sclerotized between its sclerotized, wrinkled part and
the schrotized and wrinkled margin of genital opening.
TYPE LocALiITy: Tehuacan, Mexico (type in USNM).
Foon piant: Unknown.
Known only from the type locality.
11. Cuniberta, new genus
TypE or GENUS: Nephopteryx subtinctella Ragonot.
Tongue well developed. Antenna of male with basal
segment elongate, cylindrical; shaft weakly serrate and
pubescent (the cilia about as long as width of segments),
basal segments swollen and incurved, forming a sinus
containing a, row of minute thornlike spines and overlaid
with a spread of appressed scales; antenna of female
simple and very weakly pubescent. Labial palpus up-
turned, scarcely reaching vertex; third segment shorter
than second, acuminate. Maxillary palpus small, squa-
mous. Forewing smooth; 11 veins; vein 2 from before
(but rather near) lower outer angle of cell; 3 from the
angle, but little further from 2 at base than from 4; 4
and 5 short stalked; 6 from below upper angle of cell,
straight; 8 and 9 stalked for half their lengths; 10 from
the cell, closely approximate to basal half of the stalk
of 8-9; male without costal fold. Hind wing with vein 2
from well before lower outer angle of cell; 3 from the
angle, connate with 4-5; 4 and 5 stalked for approxi-
mately half their lengths; 7 and 8 closely approximate
beyond cell; cell slightly less than half the length of
wing; discocellular vein curved. Eighth abdominal seg-
ment with 2 pairs of ventrolateral hair tufts.
Male genitalia with apical process of gnathos an
elongate, stout, rather broad hook with blunt, notched
apex. Uncus triangulate. Transtilla complete, stout,
arched, produced at middle into a broad U-shaped pro-
jection. Harpe simple. Anellus with rather broad,
dorsoventrally flattened lateral arms. Aedeagus mod-
erately slender with a single row of very minute serra-
tions along one lateral margin towards apex, otherwise
simple; penis with fine sclerotized wrinklings, otherwise
unarmed. Vinculum stout, somewhat longer than
broad.
Female genitalia with signum developed as a small
granulate cup-shaped patch; bursa small; ductus bursae
considerably longer than bursa, unsclerotized except for
a narrow sclerotization along lower margin of genital
opening; ductus seminalis from ductus bursae near
genital opening.
The genus is close to both Hemiptilocera and Crocido-
mera and shares some of the characters of each but is
distinct from both in the definite stalking of veins 4-5
of forewing and in the attachment of the ductus semi-
nalis of the female genitalia.
Contains one North American species.
61. Cuniberta subtinctella (Ragonot), new combination
FieurEes 170, 666
Nephopteryx subtinctella Ragonot, N. Amer. Phycitidae, p. 7,
1887; Monograph, pt. 1, p. 302, 1893.—Hulst, Phycitidae of
N. Amer., p. 146, 1890.—McDunnough, Check list, No.
6175, 1939.
Forewing gray, more or less dusted with whitish on
basal and median areas; antemedial line nearly vertical,
out-angled at upper and lower margins of cell, bordered
outwardly by a black line which is expanded and
strongly accented on costa, bordered inwardly on lower
margin by a reddish or reddish olivaceous patch; on
some specimens a similar shade in fold beyond the ante-
medial line; subterminal line sinuous, bordered in-
wardly by a fine black line which, in most specimens,
expands on costa into a conspicuous black spot or
streak; on costa following the subterminal line a similar
more or less expanded black spot; discal dots at end of
cell usually confluent and forming a thin black lunule
along the discocellular vein; a thin black streaklet on
vein 2. Hind wing pale smoky fuscous; veins scarcely
darker; a faintly darkened line along terminal margin.
Alar expanse, 22-26 mm.
Genitalia as given for the genus; male with apex of
uncus narrowly rounded; vinculum evenly tapering to
rather broad terminal margin.
TypE LocaLity: California (type in Paris Mus.).
Foop Prant: Unknown.
Distripution: Unirep Srarus: Utah, Park City
(June), Provo (uly, Aug.); California, Shasta Retreat
(Siskiyou County, Aug.). Canapa: British Columiia,
Kaslo (June).
12. Heras, new genus
Types or GENus: Heras disjyunctus, new species.
Tongue well developed. Antenna of male with first
segment rather long, cylindrical; shaft with a sinus and
heavy scale tuft at base, otherwise weakly pubescent.
Labial palpus upceurved, reaching above vertex; dorso-
ventrally flattened; third segment somewhat shorter
than second. Maxillary palpus squamous. Forewing
smooth; 11 veins; vein 2 from well before lower outer
angle of cell; 3 from the angle; 4 and 5 connate or very
shortly stalked, shortly separated from 3 at base; 6
from slightly below upper angle of cell, very slightly
bent towards base; 10 from the cell, closely approximate
to the stalk of 8-9; on male, a long narrow costal fold
and, on upper surface of wing, a fovea (depressed
pocket) in cell slightly beyond base. Hind wing with
vein 2 from well before lower outer angle of cell; 3 from
the angle, connate with 4-5; 4 and 5 stalked for half
their lengths; 7 and 8 closely approximate beyond cell;
cell about half the length of wing; discocellular vein
curved. Eighth abdominal segment of male with ster-
nal plate developed as a narrow sclerotized pocket at
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 35
its center; in the intersegmental area two pairs of mem-
branous eversable lobes (not haired), one long ventro-
lateral pair and one shorter dorsolateral pair.
Male genitalia with apical process of gnathos a nar-
row, somewhat flattened hook with slightly forked apex.
Uncus triangulate. Transtilla complete, stout; a rather
short bridge with long widely spaced lateral arms pro-
jecting backward; and a similarly spaced, shorter pair
of arms projecting forward and articulating with the
anellus. Harpe with a strong, long hair tuft from outer
surface of base of sacculus, otherwise simple. Anellus
a narrow plate with very long, slender, strongly sclero-
tized, lateral arms. Aedeagus rather long and slender,
sclerotized only on dorsal half; penis with some weak
wrinklings and minute scobinations near apex, other-
wise unarmed. Vinculum approximately triangulate,
but slightly longer than its greatest width; its central,
ventral area unsclerotized.
I very much dislike to erect & new genus on a single
male; but the genitalic and secondary male characters
of this example are so striking and its distinctness from
any known genus is so obvious it seems best to give it
a name and separate designation. The genus is ap-
parently closest to Hemiptilocera.
62. Heras disjunctus, new species
Figure 184
Forewing rosy fuscous with costal area beyond ante-
medial line broadly clay colored (pale ocherous); the
rose shade predominant on upper part of wing, the fus-
cous shade more accented in lower fold and along inner
margin; antemedial line weak, indicated chiefly by a
distinct but small whitish ocherous spot near inner mar-
gin; subterminal more distinct, whitish ocherous, ter-
minating at inner margin in another pale spot similar
to the one on antemedial line, inner dark margin of
subterminal line narrow and very faint; discal dots at
end of cell confluent, blackish; terminal dots confluent,
some faint blackish streaking on the veins before and
beyond the subterminal line. Hind wing pale smoky
fuscous; darker along the veins and towards outer mar-
gin. Alar expanse, 22 mm.
Male genitalia with the long posteriorily projecting
arms of transtilla terminating in flattened lobes; apex
of uncus narrowly rounded. Vinculum tapering to
evenly rounded terminal margin. Female unknown.
TyPE tocatity. Don Amo, Colombia (200 ft., July)
(type in Janse Coll.).
Foop prant. Unknown.
Described from unique male type. Superficially (in
maculation and color) it strongly resembles Hyalospila
stictoneurella Ragonot.
13. Adanarsa, new genus
Typs or Genus: Rhodophaea intransitella Dyar.
Tongue well developed. Antenna simple and pubes-
cent in both sexes. Labial palpus upturned, reaching
to vertex; slightly flattened laterally; third segment
about half the length of second, blunted and slightly
broadened (ventrally) by scales at apex. Maxillary
palpus small, squamous. Forewing smooth; 11 veins;
vein 2 from well before lower outer angle of cell; 3 from
the angle; 4 and 5 shortly stalked, separated at base
from 3; 6 from below upper angle of cell, straight; 8 and
9 stalked for half their lengths; 10 from the cell, closely
approximate to the stalk of 8-9 for a short distance be-
yond cell; male without costal fold. Hind wing with
vein 2 from well before lower outer angle of cell; 3 from
the angle, approximate to 4-5 at base; 4 and 5 stalked
for half or nearly half their lengths; 7 and 8 weakly
anastomsed for a short distance beyond cell; cell half
the length of wing; discocellular vein curved. Eighth
abdominal segment of male with a small pair of ventro-
lateral hair tufts.
Male genitalia with apical process of gnathos a slen-
der, rather long hook with slightly forked apex. Uncus
semitriangulate. Transtilla complete, strongly sclero-
tized and arched, supporting at its center a rather nar-
row, smooth, curved crosspiece. Harpe with strongly
sclerotized, erect clasper, otherwise simple. Anellus
with short, broad, dorsoventrally flattened lateral arms.
Aedeagus with a row of very fine serrations along one
lateral edge towards apex; penis armed with a single,
slender, sinuate, cornutus. Vinculum stout, about as
long as greatest width; terminal margin broad.
Female genitalia with cornutus developed as a single,
short, stout, hooked thorn; ductus bursae much shorter
than bursa, broad, flattened and with a broad transverse
sclerotized band across it at junction of ductus and
bursa; genital opening weakly and narrowly sclerotized
along its lower margin and with a narrow, transverse
sclerotized band in the membrane just behind the open-
ing; ductus seminalis from bursa near its junction with
ductus bursae.
A distinct genus distinguished from related genera
with complete transtilla by the strongly sclerotized
clasper and the slight but definite anastomoning of
veins 7-8 of hind wing. The amount of anastomosis
varies in different specimens of the type species but is
always present and always for somewhat less than half
the length of the veins.
63. Adanarsa intransitella (Dyar), new combination
Ficures 185, 667
Rhodophaea intransitella Dyar, Proc. Ent. Soc. Washington, vol.
7, p. 33, 1905.—MeDunnough, Check list, No. 6075, 1939.
Forewing pale ash gray with a fine scattered dusting
of black scales and a very faint clouding of ocherous
fuscous above inner margin between the transverse
lines; antemedial line obsolete, indicated only by small
black spot on costa and a larger black spot on inner
margin at what would be the inner margin of the trans-
verse line; outer line faint, indicated chiefly by border-
ing black dashes on costa, a faint blackish line along
its outer border and a few inwardly bordering black
dots; lower discal spot black, followed outwardly by an
obscure dark streak; a row of black dots along termen.
Hind wing whitish, subpellucid; more or less shaded
36 UNITED STATES NATIONAL MUSEUM BULLETIN 207
with fuscous towards apex and with a dark line along
termen. Alar expanse, 16-19 mm.
Male genitalia with apex of uncus evenly but rather
narrowly rounded; clasper broadly flaring at apex; vin-
culum no longer than broad; terminal margin abruptly
truncate, nearly straight (very slightly concave at
middle); aedeagus stout. Female genitalia as given for
the genus.
Type tocauity: Albuquerque, N. Mex. (type in
USNM).
Foop prant: Unknown.
Distrisution: New Mexico, Albuquerque (July);
Arizona, Christmas, Kingman (Oct.), Phoenix (Mar.).
14. Birinus, new genus
TYPE OF GENUS: Birinus russeolus, new species.
Tongue well developed. Antenna simple and pubes-
cent. Labial palpus upturned, slender, barely reaching
to vertex; third segment shorter than second, acuminate.
Maxillary palpus minute, filiform. Forewing smooth;
11 veins; vein 2 from before (but moderately near)
lower outer angle of cell; 3 from the angle, closer to 4-5
than to 2; 4 and 5 very shortly stalked; 6 from very near
to upper angle of cell, bent towards base, approximate
at base to stalk of 8-9; 8 and 9 long stalked (for more
than two-thirds of their lengths) ; 10 from the cell, closely
approximate to stalk of 8-9 for a considerable distance
beyond cell. Male without costal fold. Hind wing with
vein 2 from well before lower outer angle of cell; 3 from
the angle, connate with the stalk of 4—5; 4 and 5 stalked
for a little over half their lengths; 7 and 8 closely approx-
imate for a short distance beyond cell; cell about half
the length of wing; discocellular vein curved. Highth
abdominal segment of male simple.
Male genitalia with apical projection of gnathos long,
nearly straight, needlelike. Uncus spatulate, broadest
at apical margin. Transtilla complete, a very slightly
curved band (nearly straight on posterior margin),
broad at its bases, narrow at middle. Harpe with sac-
culus strongly sclerotized throughout its length and with
apex produced as a short spur at lower outer angle of
harpe; costa very short, sclerotized; bent upward at a
sharp right angle a short distance from base, not pro-
duced; cucullus forming more than half of the harpe
area. Sclerotized part of anellus greatly reduced. Aede-
agus long, moderately stout, smooth; penis armed with
two narrow, blade like cornuti and numerous fine granu-
lations. Vinculum triangulate, sclerotized only along
margins.
The foregoing description is incomplete, as the female
is unknown; but the new genus seems to be justified by
the male genitalia, which are unlike anything else in the
American fauna.
64. Birinus russeolus, new species
Ficures 8, 186
Forewing reddish brown, the rust-red shading a little
more pronounced in outer costal and marginal areas and
along lower vein of cell; a faint pale, clay-colored blotch
in outer median area between vein 8 and the lower fold,
enclosing in its center a small patch of blackish brown
scales and at its inner margin bordered by a similar
blackish smudge formed by the confluent discal spots;
on the fold below and just before lower outer angle of
cell a somewhat larger blackish brown patch preceded
by a small clay-colored spot; a thin line of dark scales
along the remainder of the fold to base of wing; the usual
antemedial and subterminal lines obsolete; terminal dots
very faint. Hind wing pale smoky fuscous, the veins
brown and the cell filled with brown scaling. On the
underside of fore and hind wings dark brown sex-scaling
(a male character) covers the upper wing area (above
lower margin of cell) from base to somewhat beyond the
outer margin of the cell. Forefemora of male with a
strong, clay-colored, hair tuft from upper basal angle
(also a male character). Alar expanse, 22 mm.
Male genitalia as given for the genus. Female un-
known.
TYPE LocaLiry: Tumatumari, Potaro River, British
Guiana (type in Cornell).
Foop prant: Unknown.
Described from unique male type collected by W. T.
M. Forbes, June 20, 1927 (Cornell lot 760 sub. 114).
15. Genus Bertelia Barnes and McDunnough
Bertelia Barnes and McDunnough, Contributions, vol. 2, p. 140,
1913. (Type of genus: Bertelia grisella Barnes and McDun-
nough.)
Tongue well developed. Antenna of male with a
strong posterior scale tuft on basal segment; shaft with
a deep incurvation (sinus) at base, the sinus containing
a few minute spinelike thorns but no scale tuft, shaft
beyond sinus strongly unipectinate; antenna of female
simple and pubescent. Labial palpus upturned on male,
reaching a trifle higher than vertex; oblique on female;
third segment on male slender, acuminate, about half
the length of second, on female shorter and somewhat
expanded with scales at apex. Maxillary palpus squa-
mous. Forewing smooth; 11 veins; vein 2 from well
before lower outer angle of cell, 3 from the angle; 4 and
5 shortly stalked; 6 from below upper angle of cell,
straight; 8 and 9 stalked for a trifle more than half their
lengths; 10 from the cell, closely approximate to the
stalk of 8-9; male without costal fold. Hind wing with
vein 2 from well before lower outer angle of cell; 3 from
the angle; 4 and 5 contiguous, shortly anastomosed or
stalked, usually stalked for less than half their lengths;
7 and 8 closely approximate for half their lengths beyond
cell; cell about half the length of wing; discocellular vein
curved. Highth abdominal segment of male with a pair
of ventrolateral hair tufts.
Male genitalia with apical process of gnathos de-
veloped as a stout hook, slightly notched at apex.
Uncus triangulate. Transtilla incomplete, but with
the elements long and stout, their apices broadly and
irregularly developed and hooked. Harpe simple.
Anellus U-shaped, its lateral arms dorsolaterally flat-
tened. Aedeagus with a short row of minute scobina-
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 37
tions along one lateral edge towards apex, moderately
stout; penis with a few sclerotized wrinklings, otherwise
unarmed. Vinculum stout, slightly longer than great-
est width; terminal margin broad.
Female genitalia with bursa large and elongate,
signum present, developed as a small, scobinate, cup-
shaped depression, ductus bursae very short; genital
opening with a narrow, short, sclerotized plate on its
lower margin and a large semicircular, sclerotized and
scobinate dorsal plate in the membrane behind the
opening; © pair of ventral scobinate plates in the inter-
segmental area between eighth segment collar and
Ovipositor; ductus seminalis from lobe of bursa near
itsjunction with ductus bursae.
A distinct genus containing one described North
American species. The venation of the hind wing is
individually variable in the amount of stalking or
anastomosis of veins 4 and 5.
65. Bertelia grisella Barnes and McDunnough
Fieures 187, 669
Bertelia grisella Barnes and McDunnough, Contributions, vol. 2,
p. 140, 19183 —McDunnough, Check List, No. 6140, 1939.
Forewing pale ashy gray dusted with fuscous; faint
interrupted black streaking on upper and lower veins
of cell, the lower fold and some of the veins beyond
cell; antemedial line obscure, indicated chiefly by its
narrow blackish outer border (out-angled from costa)
and by a whitish incurved line between cell and inner
margin, preceding which is an obscure dark shading;
subterminal line nearly obsolete, followed on costa by
a blackish shade; discal dots obsolete on many speci-
mens, occasionally indicated by a small blackish dot
at lower outer angle of cell. Hind wing semihyaline
white with a faint ocherous tint; veins not appreciably
darkened; a faint fuscous line along outer margin. Alar
expanse, 24-30 mm.
Male genitalia with apex of uncus bluntly and nar-
rowly rounded; terminal margin of vinculum slightly
angled; lateral arms of anellus moderately long and
broad. Female genitalia as given for the genus.
Type Locatity: Redington, Ariz. (type in USNM).
Foop prant: Unknown.
DistrisutTion: Arizona, Redington, Santa Catalina
Mts. (Sept.).
16. Genus Hypargyria Ragonot
Hypargyria Ragonot, Nouv. Gen., p. 9, 1888, Monograph, pt. 1,
p. 122, 1893.—Janse, Journ. Ent. Soc. South Africa, vol. 4,
p. 149, 1941. (Type of genus: Hypargyria metalliferella
Ragonot; India.)
Tongue well developed. Antenna pubescent, basal
segment on male elongate with a short spur of scales
from its upper inner angle (giving the base of antenna
much the same appearance as that of the undenuded
two first segments of the male antenna of Mildrizia,
fig. 169f); male shaft with a deep sinus towards base
containing a longitudinal row of very minute teeth but
no scale tuft; antenna of female simple. Labial palpus
upturned, reaching a little above vertex; third segment
nearly as long as second, acuminate. Maxillary palpus
squamous. Forewing with transverse, antemedial ridge
of raised scales; 11 veins; vein 2 from before but near
lower outer angle of cell; 3 from the angle, at base about
equidistant from 4-5; 4 and 5 connate or very shortly
stalked; 6 from upper angle of cell, connate with stalk
of 8-9, straight or but slightly bent towards base; 10
from the cell, approximate to the stalk of 8-9; male
with a notch in costa very close to base and on under-
side at base of costa a small knot of modified scales and
(projecting into the costal notch) a very short brush of
stiff hairs. Hind wing with vein 2 from well before
lower outer angle of cell; 3 from the angle, connate with
stalk of 4-5; 4 and 5 stalked for slightly less than half
their lengths; 7 and 8 closely approximate for less than
half their lengths beyond cell; cell slightly more than
one-third the length of wing; discocellular vein curved.
Eighth abdominal segment of male with compound
ventral scale tufts.
Male genitalia with gnathos weak, lacking a central
projection, the lateral arms meeting at the base of a
rudimentary subanal plate. Uncustriangulate. Trans-
tilla incomplete, but with the elements long and stout,
their apices broadly developed. Harpe with costa
broadly and very strongly sclerotized and stoutly pro-
jecting at apex; a fine moderately long hair tuft from a
sclerotized disk attached to base of sacculus. Anellus a
broad, deep plate (bearing short, knoblike, stoutly
spined, lateral projections on the American species).
Aedeagus smooth; penis armed with two or more short,
rather stout, straight spines, a deeply wrinkled, sclero-
tized band, and a cluster of fine moderately long spines.
Vinculum very stout, considerably larger than uncus
and tegumen combined, longer than broad.
Female genitalia with signum developed as a small,
round, scobinate, cup-shaped plate; ductus bursae
shorter than bursa, a broad, strongly sclerotized band
at the junction of ductus and bursa and a narrower
sclerotized band at genital opening; behind genital
opening a conspicuous pair of strongly sclerotized, gran-
ulate, pocket like lobes; ductus seminalis from bursa
near junction of bursa and ductus bursae.
Presumably an Old World genus of tropical and prob-
ably African origin; possessing some structural char-
acters of Acrobasis, Mildrixia, and Bertelia but amply
distinct from any of them. It contains two American
species.
66. Hypargyria definitella (Zeller)
Figures 188, 668
Myelois definitella Zeller.
p. 205, 1881.
Hypargyria definitella (Zeller) Ragonot, Monograph, pt. 1, p. 124,
1893.
Horae Soc. Ent. Rossicae, vol. 16,
Forewing purplish ocherous to purplish brown, most
of basal area and costal half of median area white
sparsely dusted with red scales, the red dusting most
abundant along midcosta; a small ocherous patch on
inner margin near base; antemedial line evenly curved,
38 UNITED STATES NATIONAL MUSEUM BULLETIN 207
ocherous, bordered outwardly by a red or purplish line
continued from a rather pronounced costal dash, and
inwardly by the vertical scale ridge, the latter red,
reddish ocherous or purple with some admixture of
blackish scaling; subterminal line very faint with faint
purplish borders; discal spots at end of cell separated,
blackish; black terminal dots faint. Hind wing hyaline
white with a faint smoky tint on some specimens; the
veins darkened (brown) and a narrow brown line along
termen. Undersides of male fore and hind wings in
the area between vein 2 and costa and from near end of
cell outward covered with shining silvery scales; also
on forewing a short black median streak from base,
more or less extended into cell along lower edge of upper
vein of cell and on hind wing a similar black streak
on upper vein of cell; these black sex-scalings not con-
stant and altogether absent from occasional males.
Alar expanse, 16-20 mm.
Genitalia as given for the genus.
TypE Locatity: Honda, Colombia (type in BM).
Foop prant: Unknown.
Distrisution: Purrto Rico: Puerto Real (Vieques
Isl., Apr.), San Germdén. Virern Istanps: Kingshill
(St. Croix, June, Oct.). CouomBra: Honda, Valparaiso.
Braziu: Castro, Santa Catarina.
The males of this species can be distinguished at once
from any other American phycitid by the shining
silvery scaling on the undersides of the wings; a char-
acter, however, shared by the Old World type of the
genus. The Old World metalliferliae exhibits 2 number
of slight but consistent male genitalic differences: The
heavier and more abundant spining on the penis, a
different shape to the apical projection of costa of harpe,
a different shape to the apices of the elements of trans-
tilla (not developed into paired hooks as in the Ameri-
can species), and an anellus without spined lateral pro-
jections. Such differences are certainly specific but no
more. Through the courtesy of the British Museum I
have been able to examine males of metalliferella from
Pusa in India and Nyasaland in Africa. There were
no differences of any kind between them.
67. Hypargyria slossonella (Hulst), new combination
Salabria slossonella Hulst, Canadian Ent., vol. 32, p. 170, 1900.
Acrobasis tenuella Barnes and McDunnough, Contributions, vol.
2, p. 181, 1913.
Acrobasis slossonella (Hulst) Barnes and McDunnough, Contri-
butions, vol. 3, p. 195, 1916—McDunnough, Check list,
No. 6108, 1939.
Not distinguishable from definitella except that the
males lack entirely the silvery scaling on the under-
sides of the fore and hind wings.
I suspect that it is only a variety or race of definitella;
but until more material is available and something is
known of their life histories the two forms will have to
be kept as separate species. The genitalia of slossonella
exhibit no differences of any specific significance from
those of definitella.
TypsE LocaLities: Miami, Fla. (slossonella, in AMNH,
ex Rutgers); Everglades, Fla. (tenuella, in USNM).
Foop pLrant: Unknown.
DistrisutTion: Unrrep States: Florida, Everglades
(Apr.), Fort Myers (Apr.), Miami (‘‘February-March’’).
México: Oaxaca.
17. Chararica, new genus
Typ or Genus: Myelois annuliferella Dyar.
Tongue well developed. Antenna simple and pubes-
cent on both sexes. Labial palpus upturned, reaching
to vertex, slender; third segment about as long as sec-
ond, acuminate. Maxillary palpus squamous. Fore-
wing smooth; 11 veins; vein 2 from before, but near,
lower outer angle of cell; 3 from the angle; 4 and 5
separated at base, distance separating them slightly
less than that separating 3 and 4; 6 from below upper
angle of cell, straight; 8 and 9 stalked for slightly less
than half their lengths; 10 from the cell, approximate
to the stalk of 8-9 for a short distance; male without
costal fold. Hind wing with vein 2 from well before
lower outer angle of cell; 3 from the angle, connate
with or very closely approximate to 4 at base; 4 and 5
shortly stalked; 7 and 8 approximate for a short dis-
tance beyond cell; cell about half the length of wing;
discocellular vein curved. Eighth abdominal segment
of male with a pair of ventrolateral hair tufts.
Male with gnathos weak, lacking a central projec-
tion, the lateral arms articulating with a thinly sclero-
tized subanal plate. Uncus triangulate, sharply taper-
ing to a blunt point. Transtilla complete, with a cen-
tral sclerotized apron connecting transtilla and uncus,
and with a pair of widely spaced lateral arms each
bearing at its apex a clutter of slender spinelike setae.
Harpe with costa slightly concaved, strongly sclero-
tized, not produced at apex; cucullus large, forming
about two-thirds of the harpe, outer margin broadly
rounded. Anellus with stubby, stout, convergently
directed, lateral projections. Aedeagus with lateral
margins serrated toward apex; penis with a few weakly
sclerotized wrinklings or granulations, otherwise un-
armed. Vinculum stout, nearly square in outline.
Female genitalia with signum developed as a small,
finely granulate-scobinate, cup-shaped disk; bursa large;
ductus bursae, broad, short, less than half as long
as bursa, weakly sclerotized and transversely wrinkled
towards genital opening; genital opening broad, with
strongly sclerotized transverse plate along lower margin
and some weak scobinations on the membrane above
and behind the opening; ductus seminalis from bursa
near junction of bursa and ductus bursae.
This genus is erected for three North American spe-
cies now listed under Rhodophaea, but differing from
that genus in both venational and genitalic characters.
It is easily recognized by its male genitalia. The spe-
cies also have a pattern character which aids in iden-
tification: the usual discal dots on forewing at end of
cell are replaced by a small obicular marking. This is
weak on bicolorella but present and distinguishable on
most specimens.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 39
68. Chararica annuliferella (Dyar), new combination
Ficures 189, 670
Myelois annuliferella Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 33, 1905.
Rhodophaea annuliferella (Dyar) Barnes and McDunnough,
Check list of the Lepidoptera of Boreal America, No. 5516,
1917.—McDunnough, Check list, No. 6074, 1939.
Forewing dark gray with a faint, pale ocherous shade
along inner margin; antemedial line well out on wing,
outwardly arched in cell to slightly beyond middle of
wing, inbent from lower fold to inner margin, white,
bordered outwardly by a narrow black line; basal area
with veins black and faint intervenous whitish dusting;
subterming line rather near terminal margin, slightly
sinuous, fine, white with a narrow, inner, black border;
some faint white dusting in median area, especially
along costa; usual discal dots at end of cell replaced
by small black obicular mark with a whitish center.
Hind wing hyaline white; veins not appreciably dark-
ened; a faint, small fuscous shade at apex and a very
faint, dark line along termen. Alar expanse, 19-22 mm.
Genitalia of male with lateral arms of transtilla much
reduced, widely spaced, their spinelike hair tufts long.
Vinculum with terminal margin decidedly incurved at
middle. Female genitalia with transverse sclerotized
band on lower margin of genital opening narrow.
Typxz Locatity: Gallinas Canyon, N. Mex. (type in
USNM).
Foop prant: Unknown.
Distrisution: New Merico, Gallinas Canyon (July) ;
Arizona, Colorado Desert (Yuma County), ‘‘So. Ariz.,”’
Kingman (Oct.), Yavapai County.
69. Chararica hystriculella (Hulst), new combination
Figures 190, 671
Acrobasis hystriculella Hulst, Ent. Amer., vol. 3, p. 135, 1887.
Rhodophaea hystriculella (Hulst) Ragonot, Ent. Amer., vol. 5, p.
114, 1889; Monograph, pt. 1, p. 78, 1893.—McDunnough,
Check list, No. 6073, 1939.
Myelois hystriculella (Hulst) Hulst, Phycitidae of N. Amer., p.
119, 1890.
Forewing whitish gray with extreme base dark fuscous
and a similar fuscous shade over the outer area from
subterminal line on costa obliquely to inner margin near
antemedial line, and outward to termen; some black
streaking on upper and lower veins of cell and vein 1b
before the antemedial line; antemedial line well out
towards middle of wing, twice angled outwardly, indi-
cated chiefly by a fine black outwardly bordering line;
subterminal line faint, with narrow dark inner and outer
borders, beginning as blackish dashes on costa; obicular
spot at end of cell conspicuous, black with a narrow
whitish center; terminal dots black, more or less con-
fluent. Hind wing hyaline white with a faint fuscous
shade at apex and a narrow dark line along termen,
these dark shadings very slight on the males, more ex-
tended and stronger on some females; veins occasionally
darkened on females, not darkened on males. Alar
expanse, 17-23 mm.
Male genitalia with uncus narrowly triangulate; lat-
eral arms of transtilla rather long and their terminal
hair tufts correspondingly shortened, not so widely
spaced as those of annuliferella; terminal margin of
vinculum very slightly concave, nearly straight. Fe-
male genitalia with transverse sclerotized band on lower
margin of genital opening broad (at least twice as wide
as that of annuliferella).
Typ Locauity: Texas (type, 2, in AMNH, ex Rut-
gers).
Foop prant: Unknown.
Distrisution: Texas, Blanco County (Sept.), Browns-
ville (May, June), Chisos Mts. (June), Cotula (Mar.,
Apr., May), Devils River (May), Kenedy (May), Kerr-
ville (Aug.), Nueces River (Zavalla County, Apr.),
Sabinal (Mar.), San Antonio (June, July), San Benito
(Mar.), San Diego (Apr., May, June); Florida, Coconut
Grove, Miami.
70. Chararica bicolorella (Barnes and McDunnough), new
combination
Rhodophaea bicolorella Barnes and McDunnough, Canadian Ent.,
vol. 49, p. 404, 1917.—MceDunnough, Check list, No. 6077,
1939.
Forewing with costal half of basal area black, streaked
and peppered with white, giving this area of the wing
a slate-colored appearance to the naked eye; outer area
of wing from subterminal line to outer margin and costa
to lower fold a similarly dark shade; remainder of wing
ocherous, shading outwardly to tawney or ruddy ocher-
ous; antemedial line obsolete except along outer margin
of blackish basal patch; subterminal line faint, narrowly
and weakly bordered inwardly and outwardly by black-
ish lines; obicular mark at end of cell very faint but
distinguishable on most specimens; terminal dots con-
fluent.. Hind wing hyaline white with a very faint
ocherous line on outer margin for a short distance from
apex. Alar expanse, 20-24 mm.
Genitalia essentially like those of hystriculella.
Typr Locauity: Christmas, Gila County, Ariz. (type
in USNM).
Foop prant: Unknown.
Distrisution: Arizona, Christmas, Mohave County
(May, June, July, Aug., Sept.), Redington; Nevada,
Clark County (Apr.), ‘“‘So. Nevada” (July); California,
San Bernadino County (Apr.).
A striking species easily distinguished by its color pat-
tern, but not structurally different from hystriculella.
The original type series consists of two males and two
females, not four males as stated by the authors.
Both bicolorella and hystriculella have a strong hair
tuft on the metathorax of the male adjacent to the base
of the leg. This character is lacking in annuliferella.
40 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Genera 18-21: Myelopsis to Ectomyelois
[Venational division C. Forewing with 11 veins; 4and 5 stalked
for half their lengths or less. Hind wing with veins 7 and 8 anas-
tomosed for over half their lengths beyond cell (the free element
of 8 shorter than the anastomosed stalk of 7-8). Transtilla of
male genitalia complete.]
18. Myelopsis, new genus
Typr or GENus: Myelois coniella Ragonot.
Tongue well developed. Antenna simple, pubescent.
Labial palpus upturned, reaching slightly above ver-
tex; second segment somewhat roughly scaled in front;
third segment slightly shorter than second, acuminate.
Maxillary palpus filiform. Forewing smooth; 11 veins,
vein 2 from before but near lower outer angle of cell;
3 from the angle; 4 and 5 shortly stalked, the stalk at
base separated from 3 for a distance but slightly less
than that between 3 and 2; 6 from below upper angle
of cell, straight; 10 from the cell separated from 8-9
at base, and more or less divergent beyond; male with
out costal fold. Hind wing with vein 2 from well be-
fore outer angle of cell; 3 from the angle, connate with
the stalk of 4-5; 4 and 5 stalked for slightly less than
half their lengths; 7 and 8 strongly anatomosed beyond
cell, the free element of 8 short; cell half the length of
wing; discocellular vein curved. Eighth abdominal
segment of male simple.
Male genitalia with apical process of gnathos U-
shaped (consisting of a pair of widely spaced, short
arms). Uncus stout, more or less triangulate, apex
rather narrowly rounded. Transtilla complete, but
weakly sclerotized. Harpe simple; costa strongly scler-
otized and projecting at apex (except in subtetricella).
Anellus U-shaped, narrowly sclerotized throughout.
Aedeagus smooth; penis with sclerotized wrinklings.
Vinculum triangulate, tapering, longer than greatest
width.
Female genitalia with or without signum, latter when
present weak. Ductus bursae membranous. Genital
opening simple. Ductus seminalis from middle or
towards terminal end of bursa.
This genus comprises several North American species
that have been referred to Myelois Hibner. ‘The latter
a, heterogenous assemblage of species, very few of which
are actually congeneric with the type of genus (medu-
lallis Hiibner synonym of cribrella Hiibner). The lat-
ter has veins 7 and 8 of hind wing very shortly and
weakly anastomosed beyond the cell, the free element
of vein 8 correspondingly long and the cell itself over
half the length of the wing. It belongs properly in
our venational division B. None of the American and
very few of the Old World species that have been as-
signed to it are properly referrable to Myelois. The
European éetricella Schiffermueller belongs in Myelopsis.
71. Myelopsis coniella (Ragonot), new combination
Fieures 191, 673
Myelois coniella Ragonot, N. Amer. Phycitidae, p. 3, 1887;
Monograph, p. 1, p. 53, 1893.—Hulst, Phycitidae of N.
Amer., p. 118, 1890.—McDunnough, Check list, No. 6071,
1939.
Rampylla nefas Dyar, Ins. Insc. Menstr., col. 10, p. 172, 1922
(new synonymy).
Forewing pale ash gray (on Utah and Nevada spec-
imens) to blackish gray; on darker specimens the basal
and terminal areas are contrastingly paler than the
area between the transverse lines; at extreme base on
inner margin an obscure ocherous patch (not distin-
guishable on worn or faded specimens) ; antemedial line
oblique, white with a broad, black outer border; sub-
terminal line sinuate, more or less contrastingly pale
and inwardly bordered by a blackish line or varying
intensity; discal dots at end of cell usually distinct,
separated, black; terminal dots obscure, when distin-
guishable, more or less confluent. Hind wing hyaline
white to pale smoky fuscous with a fine dark line along
termen. Alar expanse, 16-22 mm.
Male genitalia with transtilla slender, arched and
very weakly sclerotized at the central attachment of
its elements. Harpe with costa narrowly sclerotized
and projecting a trifle beyond the apex of the cucullus.
Female genitalia without signum; bursa membranous;
ductus bursae with some minute scobinations near its
junction with bursa, otherwise smooth.
TYPE Locatities: Nevada (coniella, in Paris Mus.);
Mexico City, Mexico (nefas, in USNM).
Foop puant: Unknown.
DistriputTion: Unitep states: Nevada, Montgom-
ery Pass (Mineral County, Sept.), Utah, Dividend
(Aug.), Eureka (June, Aug.), Provo (July, Aug.,
Sept.), Stockton (Sept.), Trout Creek (Ibapah Mts.,
Sept.); Colorado, Glenwood Springs (Aug.); Arizona,
Pinal Mts. (July), no definite locality (Aug., Sept.);
New Mexico, Gallinas Canyon; Texas, Burnet County,
(Mar.), Kerrville (Mar.); Nebraska, Sioux County,
(July); Michigan, Dickinson County; Maine, Bar Har-
bor (Aug.), Mount Desert (Aug.). Cawnapa: British
Columbia, Kaslo (July, Aug.) ; Manitoba, Aweme (Aug.);
Ontario, Ottawa (July, Aug.). Mexico: Mexico City
(Sept.), Tehuacdn (Sept.).
A variable species in color but with remarkably con-
stant genitalia. Dyar’s nefas has much darker fore-
wings than specimens from Utah or Nevada but no
darker than some specimens from Arizona and New
Mexico.
72. Myelopsis immundella (Hulst), new combination
Myelois immundella Hulst, Phycitidae of N. Amer., p. 117,
1890.—Ragonot, Monograph, pt. 1, p. 49, 1893.—MceDun-
nough, Check list, No. 6068, 1939.
The type is without abdomen. In size, wing shape,
pattern, and general coloration it is like the following
species (subtetricella) except that the antemedial and
subterminal lines of forewing are more whitish and dis-
tinct and their dark borders (especially the outer border
of the antemedial line) blackish and more strongly con-
trasted against the dark ground color of the wing. The
discal dots are also more strongly contrasted.
The name may represent only a color form of subtetri-
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 41
cella, but until other specimens matching the type are
found and their genitalia studied this cannot be deter-
mined one way or the other.
TyprELocaLity: Texas (typein AMNH, ex Rutgers).
Foop puant: Unknown.
73. Myelopsis subtetricella (Ragonot), new combination
Fiaures 192, 672
Myelois subtetricella Ragonot, Ent. Amer., vol. 5, p. 113, 1889;
Monograph, pt. 1, p. 47, 1893.—Hulst, Phycitidae of N.
Amer., p. 117, 1890.—McDunnough, Check list, No. 6062,
1939.
Myelois zonulella Ragonot, Ent. Amer., vol. 5, p. 113, 1889;
Monograph, pt. 1, p. 49, 1893.—Forbes, Cornell Mem. 68, p.
613, 1923.—McDunnough, Check list, No. 6066, 1939. (New
synonymy.)
Myelois obnupsella Hulst, Phycitidae of N. Amer., p. 118, 1890.—
Ragonot, Monograph, pt. 1, p. 48, 1893.—Barnes and
McDunnough, Contributions, vol. 3, p. 193, 1916.—Forbes,
Cornell Mem. 68, p. 613, 1923.—McDunnough, Check list,
No. 6063, 1939. (New synonymy.)
Forewing brownish gray with some faint whitish dust-
ing on basal and median costal areas; antemedial line
but slightly oblique, rather faint, dull whitish with a
more or less obscured dark outer border; subterminal
line obsolete or very faintly indicated; discal dark dots
at end of cell separated, only the lower one distinct and
always distinguishable. Hind wings dull smoky white
to pale fuscous; veins darkened slightly in several speci-
mens; a narrow dark line along termen. Alar expanse,
20-24 mm.
Male genitalia similar to those of coniella except scle-
rotized costal margin of harpe abruptly terminated before
apex of cucullus and not projecting as a free spur at apex.
Female genitalia with a small weak signum; bursa
weakly sclerotized, finely scobinate, and with a longi-
tudinal sclex otized groove in area near ductus bursae;
ductus bursae very short; ductus seminalis from bursa
well towards its terminal end,
Typs Locauities: ‘North America” (subtetricella, in
Paris Mus.); north Illinois (zonulella, in BM); Canada
(obnupsella, in AMNH, ex Rutgers).
Foop piant: Unknown.
DisrriputTion: Unrrep States: New Hampshire,
Hampton; Massachusetts, Cohasset (July), Forest Hills
(May, June), Framington (May), Winchendon (May);
Pennsylvania, Beaver County (May), New Brighton
(May, June), Pittsburgh (May); Jilmois, Arlington
Heights (May), Chicago (May), Quincey (June); Ohio,
Calla; Florida (no specific locality, Mar.). Canapa:
Alberta, Bilby (June), Edmonton (May); Manitoba,
Aweme (May, June).
The species is quite distinct and easily identified by
male and female genitalic characters. The Florida rec-
ord cited above is from a spurious “type (male)’’ of
Myelois immundella Hulst, originally in the Fernald Col-
lection and now in the U. S. National Museum. It is
not immundella, and in genitalia, color and markings
agrees perfectly with other males of subtetricella. Rag-
onot’s zonulella was described from four females in the
British Museum labeled ‘‘N, Ill.” and bearing the num-
ber “82-54.” I have examined the genitalia of two of
these and they agree in all details with those of the type
of subtetricella. As Ragonot designated no holotype I am
selecting as lectotype one of the specimens J examined.
74, Myelopsis minutularia (Hulst), new combination
FiauRE 675
Dioryctria minutularia Hulst, Ent. Amer., vol. 3, p. 135, 1887.
Myelois minutulella Hulst, Phycitidae of N. Amer., p, 118, 1890.—
Ragonot, Monograph, pt 1, p. 48, 1893.
Myelois minutularia (Hulst) McDunnough, Check list, No. 6064,
1939.
The status of this species is in doubt. It is known
only from females which look like small dark examples
of coniella, of which it may be only a race or variety.
However, the ductus bursae of minutularia is longer than
that of typical coniella and the bursa shows considerably
more scobination. Hind wing semihyaline, smoky
white. Alar expanse, 11-13 mm.
Type xnocauity: Blanco County, Tex. (type in
AMNG, ex Rutgers).
Foop pLant: Unknown.
The only known distribution is Texas; examples be-
fore me are from Blanco and Burnet Counties. The
statement by Hulst in his original description that his
types are males is an error. The male is unknown.
75. Myelopsis alatella (Hulst), new combination
Ficures 193, 194, 195, 674
Acrobasis alatella Hulst, Ent. Amer., vol. 3, p. 135, 1887.
Myelois rectistrigella Ragonot, N. Amer. Phycitidae, p. 3, 1887.
Myelois alatella (Hulst) Hulst, Phycitidae of N. Amer., p. 118,
1890.—Ragonot, Monograph, pt. 1, p. 52, 1893—McDun-
nough, Check list, No. 6070, 1939.
Myelois fragilella Dyar, Proc. Ent. Soc. Washington, vol. 6, p
114, 1904.—McDunnough, Check list, No. 6060, 1939.
(New synonymy.)
Myelois piazzella Dyar, Ins. Inse. Menstr., vol. 13, p. 11, 1925.—
McDunnough, Check list, No. 6061, 1939. (New
synonymy.)
Forewing ash gray more or less dusted with fuscous,
general color varying from pale ash gray to grayish
fuscous (but not so dark as some specimens of coniella) ;
antemedial line oblique, indicated by its narrow, black
outer border which is shortly and sharply out-angled
at middle; subterminal line rather close and paralled
to outer margin, sinuate, sharply indented between
costa and vein 6, very slightly so at lower fold, often
obscure, sometimes with a distinct inwardly bordering
black line; discal dots separated, black, lower one (at
least) always distinct. Hind wing semihyaline smoky
white, somewhat darkened towards apex and with more
or less darkening of the veins; a fine dark line along
termen. Alar expanse, 20-26 mm.
Male genitalia with transtilla a thin, weakly
sclerotized subtriangulate plate. Harpe with costa
broadly sclerotized, produced at apex, but not extending
to apex of cucullus. Female genitalia with a small
signum; a rather large round area of dorsal surface of
bursa thinly sclerotized.
42 UNITED STATES NATIONAL MUSEUM BULLETIN 207
TyprE Locauities: Napa, Calif. (alatella, in AMNH,
ex Rutgers); California (rectistrigella, in Paris Mus.);
Pecos, N. Mex. (fragilella, in USNM); San Diego,
Calif. (piazzella, in USNM).
Foop puant: Unknown.
Distrisution: California, Clarkville (June), Monache
Meadows (July), Napa, Placerville (May), San Diego
(Mar., Aug.), San Francisco (Apr.); Utah, Bellevue
(Apr.); Colorado, Gunnison County (near Almont,
July); New Mexico, Fort Wingate (June, July), Jemez
Springs (June), Pecos (June).
An individually variable species in color and to some
extent in male genitalia. The actual holotypes of
alatella, fragilella, and piazzella seem different enough;
but there are all intergrades among them in a series
from any given locality. Indeed the two cotypes of
alatella from Napa, Calif. (alike in color and markings)
show considerable variation in details of male genitalia
(width of the sclerotized costa of harpe, shape of
transtilla, and spacing of the apical prongs of gnathos).
The genitalia of the cotype from Napa (in USNM, fig.
193) shows an extreme of variation. The other cotype
(in AMNH, ex Rutgers, the actual holotype) has
genitalia identical with those of piazzella shown in
figure 195. At most, the Dyar names represent forms
or varieties, but not species or local races.
19. Genus Anypsipyla Dyar
Anypsipyla Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 327, 1914.
(Type of genus: Anypsipyla univitella Dyar.)
Tongue well developed. Antenna of male shortly
ciliate, the cilia no longer than width of shaft; of female
pubescent. Labial palpus obliquely upturned, reach-
ing slightly above vertex; third segment about as long
as second, acuminate. Maxillary palpus very slightly
dilated with scales at apex (subsquamous). Forewing
smooth; 11 veins; vein 2 from before but close to lower
outer angle of cell; 3 from the angle, approximate to
2; 4 and 5 stalked for approximately half their lengths,
approximate (rarely connate) to 3 at base; 6 from below
upper angle of cell, straight; 8 and 9 stalked for about
half their lengths; 10 from the cell, at base closely
approximate to or connate with stalk of 8-9, thence
divergent; male with short narrow costal fold. Hind
wing with vein 2 from well before lower outer angle of
cell; vein 3 from the stalk of 4-5; 4 and 5 long stalked;
7 and 8 anastomosed beyond cell for appreciably more
than half their lengths; cell about half the length of
wing; discocellular vein curved. Hight abdominal
OME of male with a single pair of ventrolateral hair
tuits.
Male genitalia with apical projection of gnathos an
elongate hook with slightly notched apex. Uncus sub-
triangulate (hoodlike). Transtilla complete, strongly
arched. Harpe with costa strongly sclerotized through-
out and projecting at apex beyond apex of cucullus;
otherwise simple. Anellus a narrow band with slender
lateral arms. Aedeagus simple; penis with some
weakly sclerotized wrinklings, otherwise unarmed.
Female genitalia with or without signa, if present, in
the form of a row of very small, weak, thornlike spines;
bursa very finely scobinate, ductus bursae considerably
longer than bursa, simple; genital opening simple;
ductus seminalis from anterior (terminal) end of bursa.
A distinct genus with one tropical American species.
76. Anypsipyla univitella Dyar
Ficures 39, 196, 679
Anypsipyla univitella Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 327,
1914.
Forewing fuscous gray with a broad white subcostal
streak extending from near base to apex and touching
costa near base and at apex; a black streak along mid-
costal edge and on fresh specimens a fine black line along
lower vein of cell and some faint black streaking on the
outer veins; a fine powdering of reddish scales on the
white subcostal stripe; discal dots faint or absent; trans-
verse lines obsolete. Hind wing hyaline white with a
smoky tint along costa and at apex and a fine dark line
along termen. ‘Alar expanse, 20-32 mm.
Male genitalia with terminal margin of uncus rather
broadly round; apical projection of costa of harpe blunt;
transtilla truncately arched. Female genitalia as given
for the genus. The signa are usually absent and when
present consist of from 2 to 10 very weak spines.
TypE Locatity: Corozal, Canal Zone, Panama (type
in USNM).
Foop piants: Cassia brasiliensis, Samanea samdn
(larva feeding in pods), Pacae (larva in fruit).
Distrisution: Cusa: Victoria de las Tunas, San
Blas (Trinidad Mts., May). Mexico: Colima (May,
Nov.). Guatmmata: Cayuga (Apr.), Quirigué (Mar.).
PanamA: Corozal (Apr., Noy.), Las Sabanas (Apr.),
Porto Bello (May). VeEnezurza: El Valle (Apr.).
Braziu: “S. E. Brazil,” Tapera (Pernambuco). PERU:
Lima (Feb.). Ecuapor. Jamaica: Kingston (Dec.).
Probably generally distributed in tropical America,
where its host plants occur.
20. Apomyelois, new genus
Types oF Genus: Dioryctria bistriatella Hulst.
Tongue well developed. Antenna simple and pubes-
cent on both sexes. Labial palpus upturned, slender,
reaching to slightly above vertex; third segment slightly
shorter than second, acuminate. Maxillary palpus fili-
form. Forewing smooth; 11 veins; vein 2 from well
before lower outer angle of cell; 3 from the angle; 4 and
5 stalked for slightly less than half their lengths, the
stalk separated from 3 at base; 6 from below upper
angle of cell, straight; 8 and 9 long stalked (for over
two-thirds their lengths); 10 from the stalk of 8-9; male
without costal fold. Hind wing with vein 2 from well
before lower outer angle of cell; vein 3 from the angle;
4 and 5 stalked for two-thirds of their length, the stalk
connate with 3; 7 and 8 anastomosed beyond cell for
about half their lengths (the anastomoses slightly longer
than the free part of vein 8); cell a trifle more than half
the length of the wing; discocellular vein curved. Kighth
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 43
abdominal segment with a pair of weak, short, ventro-
lateral hair tufts.
Male genitalia with apical projection of gnathos an
elongate, slender, simple hook. Uncus subtriangulate,
apex rounded. Transtilla complete, well sclerotized
and strongly arched; produced at middle into a flat,
broadly and bluntly forked projection. Harpe with
costa sclerotized throughout and projection slightly at
apex; otherwise simple. Anellus U-shaped. Aedeagus
smooth, slightly flaring at apex; penis with a single,
straight, short, weakly sclerotized, spikelike cornutus
and a few minute and weak scobinations; otherwise
unarmed.
Female genitalia with signa present as an oval cluster
of thornlike scobinations; bursa otherwise smooth, large;
ductus bursae considerably shorter than bursa, simple;
genital opening simple; ductus seminalis from anterior
(terminal) end of bursa.
This genus is another subtraction from the composite
genus Myelois of Authors. Of all the American species
that have been referred to that genus it is the nearest
to the type of Myelois (medullalis Hiibner, a synonym
of cribreila Hiibner) of any American species, agreeing
with cribrelia in forewing venation, except that the stalk-
ing of vein 10 with 8-9 is less consistent in cribrella than
in bistriatella. In eribrella 10 is often short stalked (as
in fig. 38); but it is as often from the cell, connate with
or approximate to or distinctly separated from the stalk
of 8-9. In bistriatella it is from the stalk of 8-9 on all
specimens that I have seen. However, this difference
has no more than specific significance and would not of
itself justify any separation of bistriatella from Myelois.
There are some other differences that, in my judgment,
are of generic character and justify such separation.
The hind wing venation and length of cell are similar in
cribrella and bistriatella except for the anastomosis of
veins 7 and 8; in cribrella this anastomosis is very weak
and shorter than it is in bistriatella and would place
typical Myelois in our venational division B, while
Apomyelois would go definitely into division C. In male
genitalia cribrella (fig. 203) differs in having apical pro-
jection of gnathos developed as a deeply, strongly, and
narrowly forked process and the transtilla developed as
a simple, strongly arched, narrow band. The female
genitalia of cribrella (fig. 684) differs in more striking
fashion: the ductus bursae being much longer than
bursa and strongly granulate and partially sclerotized
throughout most of its length; and ductus seminalis is
from the bursa between the signum and the junction of
bursa and ductus bursae.
The new genus contains one North American species.
77. Apomyelois bistriatella (Hulst), new combination
Ficures 40, 197, 676
Dioryctria bistriatella Hulst, Ent. Amer., vol. 3, p. 136, 1887.
Myelois bilineatella Ragonot, N. Amer. Phycitidae, p. 3, 1887;
Monograph, pt. 1, p. 48, 1893.—Hulst, Phycitidae of
N. Amer., p. 117, 1890; U. S. Nat. Mus. Bull. 62, p. 418,
1902.
Myelois bistriatella (Hulst) Hulst, Phycitidae of N. Amer., p. 117,
1890.—Ragonot, Monograph, pt. 1, p. 51, 1893.—Barnes
and McDunnough, Contributions, vol. 3, p. 194, 1916.—
Forbes, Cornell Mem. 68, p. 613, 1923.—McDunnough,
Check list, No. 6067, 1939.
Forewing gray-brown faintly dusted with white on
costal half of basal area and in central area from mid-
costa to lower margin of cell; transverse lines white,
rather sharply contrasted, especially towards inner
margin and without appreciably contrasted blackish
borders; antemedial line transverse, from costa dis-
tinctly before middle, straight, except for an occasional
slight notch in cell; subterminal line somewhat narrower
and less distinct, sinuate; dark discal dots at end of
cell often fused into a single spot or line along discocel-
lular vein, usually set off by the surrounding white
dusting of the central area; terminal dots very faint,
more or less confluent. Hind wing dull smoky white,
the veins slightly darkened and a narrow dark line
along termen. Alar expanse, 19-22 mm.
Genitalia as given for the genus.
Type Locauities: Washington, D. C. (bistriatella,
type lost?); “America septentrionale” (bilineatella, in
Paris Mus.).
Foop prant: Unknown.
Distrisution: Unirep States: Florida (Mar.); Dis-
trict of Columbia, Washington; New York; Massachusetts,
Framingham (May); Jilinois, Edgebrook (May); Wis-
consin; Iowa, Ames (May). Canapa: Ontario, Ottawa
(June, July), Trenton (May, June).
Probably much more widely distributed throughout
eastern and central United States and Canada, nowhere
apparently a very abundant species.
The supposed type of bistriatella is labeled “Iowa,
H. S. Saunders, June 6, 1886.” It is definitely that
species but, unless it is mislabeled or the type locality
given by Hulst in his original description (“‘Washington,
D. C.”’) is wrong, it could not be the actual holotype.
I have seen no specimens anywhere labeled “‘Washing-
ton, D. C.”’ There is a female in the National Museum
from the Fernald Collection, bearing a Hulst type label
but no locality. This might be the true type. It is a
perfect match for the Iowa specimen in the Rutgers
Collection. Since there can be no reasonable doubt as
to what the name stands for we may as well consider
the holotype lost and forget it.
21. Ectomyelois, new genus
Tyrer or genus: Myelois decolor Zeller.
Tongue well developed. Antenna of male shortly
ciliate (cilia about the length of width of shaft or
slightly less), otherwise simple; of female simple and
pubescent. Labial palpus upturned, reaching to or
nearly to apex (not above it); second segment some-
what broadened with scales; third segment short,
distinctly shorter than second, acuminate. Maxillary
palpus filiform. Forewing smooth; 11 veins; vein 2
from well before angle; 3 from the angle, shortly
separated from the stalk of 4-5 at base; 4-5 shortly
stalked (very shortly stalked in most specimens and
never for more than half the length of the veins);
44 UNITED STATES NATIONAL MUSEUM BULLETIN 207
6 from below upper angle, straight; 8 and 9 long stalked,
for over two-thirds of their lengths; 10 from the cell,
closely approximate to the stalk of 8-9 for some distance
from cell; male without costal fold. Hind wing with
vein 2 from well before lower outer angle of cell; 3 from
the angle; 4-5 stalked for not over half their lengths
(usually for less), the stalk connate with or very closely
approximate to 3 at base; 7 and 8 strongly anastomosed
for most of their lengths beyond cell (free element of 8
very short); cell half the length of wing; discocellular
vein curved. Highth abdominal segment of male
simple or with a weak, short pair of ventrolateral hair
tufts.
Male genitalia similar to those of the foregoing
genus (Apomyelois) except: Apical process of gnathos
slightly notched at apex; costa of harpe not produced
at apex (except slightly in muriscis and zeteki); penis
without cornutus; vinculum more truncate and less
tapering.
Female genitalia with signum sometimes absent,
when present consisting of an elongate patch of scobina-
tions; ductus bursae normally longer than bursa, in-
dividually variable, simple, except for a weak scleroti-
zation at genital opening; ductus seminalis from bursa
near junction of bursa and ductus bursae.
In male genitalia there is little or nothing of a generic
character to separate Hctomyelois from Apomyelois.
The two genera are distinguished by the shorter cell
and much more extended anastomosis of veins 7 and 8
of hind wing and the different place of departure of
ductus seminalis from the bursa of the female.
The genus is erected for another group of species
(American and European) removed from Myelois of
Authors. These species all appear to be of tropical or
semitropical origin. They are distinguished from
typical Myelois by the much stronger anastomosis of
veins 7 and 8 of hind wing and the consequent reduction
of the free element of vein 8.
78. Ectomyelois decolor (Zeller), new combination
Ficures 198, 677
Myelois decolor Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 222,
1881.—Ragonot, Monograph, pt. 1, p. 58, 1893.—Dyar,
Proc. U. 8. Nat. Mus., vol. 47, p. 326, 1914.—Wolcoitt,
Journ. Agr. Univ. Puerto Rico, vol. 20, No. 1, p. 476, 1936.
Nephopieryx ephestiella Hampson, Ann. Mag. Nat. Hist., ser. 7,
vol. 7, p. 257, 1901 (new synonymy).
Forewing dark grayish fuscous with some white
powdering in basal area and considerably more in the
median area from slightly above inner margin and in
outer area between subterminal line and termen; trans-
verse lines white, well contrasted, especially the ante-
medial line which is rather wide, sharply oblique,
slightly indented at lower fold and (in some specimens)
in the cell, outwardly bordered by a more or less ob-
scure dark shade; subterminal line fainter, narrow,
sinuate, obscurely and narrowly dark margined; some
faint blackish streaking on the veins; discal dots at end
of cell distinct, separated; terminal dots normally well
contrasted and separate, blackish. Hind wings smoky
white to pale smoky fuscous; the veins darkened and a
narrow dark line along termen. Eighth abdominal
segment of male with hair tufts. Alar expanse, 19-30
mm,
Male genitalia with outer margin of uncus rather
evenly rounded; central area of transtilla produced into
a moderately broad plate with notched terminal margin;
anellus a broad plate with wide, flattened, incurved,
stubby arms; vinculum nearly square in outline, its
terminal margin very slightly concave.
Female genitalia exhibiting considerable individual
variation in the size of bursa and corresponding length
of ductus bursae which is usually considerably longer
than bursa; signum patch of variable shape but usually
elongate. The female genitalia exhibit no distinctively
specific characters.
Type LocaLities: Honda, Colombia (decolor, in
BM); Nassau, Bahamas (ephestiella, in BM).
Foop PLANTs: Annona squamosa, Ceratonia siliqua,
Hymenaea courbil; these records from reared specimens
in the U.S. National Museum. Presumably the species
has much the same hosts and habits as the closely re-
lated Ectomyelois ceratoniae. The larvae feed in the
fruits and are very difficult to separate from those of
ceratoniae.
Distripution: Cusa: Baracoa (Aug., Oct., Nov.),
Havana, Santiago de las Vegas (Mar.), “Santiago Prov-
ince” (Sept., Oct., Dec.). Purrro Rico: Arecibo,
San Germdn (Apr.). Jamaica. Banamas: Nassau.
GuaTEMALA: Cayuga (Mar., June, Aug.). PanamA:
Porto Bello (Mar., Dec.). Conomspra: Honda, ‘West
Slopes” (4,400 ft., Feb.). Venrzurxa: Aroa. BritisH
Gu1ana: Tumatumari (June). FREncH Guiana: Cay-
enne, St. Jean Maroni. Surtnam: Surinam River.
Braziu: Paré (June), Ponte Nova (Rio Xingu, Amazon-
as), Santa Catarina (July).
Ragonot considered decolor as a probable variety of
ceratoniae; but there is a consistent difference in the
shape of the transtilla between the two which indicates
more than varietal or racial difference; and in unrubbed
and unfaded specimens the color difference is obvious
and consistent. E. decolor seems to be confined to the
New World while ceratoniae occurs in both the New and
Old World. Hampson’s ephestiella is nothing more than
a rather large example of decolor. Like other species in
this genus, decolor varies greatly in size.
79. Ectomyelois ceratoniae (Zeller), new combination
Figures 199, 678
Myelois ceratoniae Zeller, Isis von Oken, 1839, p. 176; 1848, p.
675.—Ragonot, Monograph, pt. 1, p. 57, 1893.—Staudinger
and Rebel, Catalog der Lepidopteran des palaearctischen
Faunengebietes, vol. 2, No. 787, 1901.—Spuler, Die Schmet-
terlinge Europas, vol. 2, p. 216, 1910.—Forbes, Cornell
Mem. 68, p. 614, 1923.—Meyrick, Revised handbook of
British Lepidoptera, p. 395, 1928.—Wolcott, Journ. Agr.
Univ. Puerto Rico, vol. 20, No. 1, p. 475, 1936.—Corbet
and Tams, Proc. Zool. Soc. London, ser. B, vol. 113, p. 68,
1943.
Myelois oporedestella Dyar, Proc. Ent. Soc. Wash., vol. 18, p. 30,
1911.—McDunnough, Check list, No. 6065, 1939. (New
synonymy.)
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 45
Color and markings similar to decolor except: Fore-
wing more uniformly gray, with less of the white dust-
ing, especially on median area; antemedial line narrower
and usually more distinctly notched. The chief char-
acter, however, is in the transtilla of the male genitalia.
The central projection of this organ is more constricted
and decidedly narrower on ceratoniae than on decolor,
and this difference seems to be consistent. In several
preparations of each species from different rearings and
localities I have found no intergrading examples. The
female genitalia offer no satisfactory distinguishing
characters. As in decolor, individual differences (even
in the proportional length of the ductus bursae) are
greater than any difference between the two species.
Alar expanse, 16-24 mm.
Type Locaities: Laibach, Austria (ceratoniae, in
BM); Miami, Fla. (oporedestella, in USNM).
Foop piants: Carissa grandiflora, Cassia bicapsularis,
Ceratonia siliqua, Erisbotyra japonica (chiefly in mum-
mied fruits), Livistona chinensis, Robinia, Tamarindus
indica, Vachellia insularis. Also on dried figs, dates,
raisins, and nuts in storage. Primarily a leguminous
feeder. The favored host seems to be the pods and
seeds of the corob (Ceratonia siliqua).
Distrisution: Unitep Srares: Florida, Homestead
(May), Key West (Apr.), Miami (May, July, Aug.,
Noyv.). Purrto Rico: Arecibo (May), Trujillo Alto
(Mar., July). Jamaica (July). Ancentina: Buenos
Aires (Feb.), Catamarca (May, June). Also in the Old
World in the Mediterranean areas of Europe, Africa
and Asia and (by introduction in dried fruits) extending
into Central Europe and England.
Apparently of Mediterranean origin, introduced by
commerce and established in some tropical and semi-
tropical areas of the New World. Probably much more
widely distributed than indicated by the above records
from specimens before me. The species is of minor
importance as a feeder on the seeds of the corob. It
has been found rather frequently at our port quarantine
stations in shipments of English walnuts from Italy.
I have omitted all European synonymy as I have not
been able to verify its correctness. This, with further
references to European literature will be found in
Ragonot’s monograph and the Staudinger and Rebel
catalog. Myelois phoenicis Durrant may be only a
color variety or race of ceratoniae; a small series before
me reared from dates from Algeria has the ground color
of forewing white, but the male genitalic characters
of ceratoniae. Corbet and Tams list phoenicis as a
synonym.
80. Ectomyelois muriscis (Dyar), new combination
Figures 200, 680
Myelois transitella Dyar (not Walker), Proc. U. S. Nat. Mus.,
vol. 47, p. 326, 1914.
Hypsipyla muriscis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330,
1914,
Myelois palpalis Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 1919
(new synonymy).
Forewing dull rusty brownish ocherous to reddish
brown; costal third to half of wing strongly dusted with
white, the white area rather well contrasted against
dark ground color; antemedial line angulate, obscure,
indicated chiefly by a brown or blackish spot on its
outer margin at or just below costa; subterminal line
better defined, sinuate, margined inwardly and out-
wardly by narrow dark lines, the latter especially
emphasized at costa; discal spots usually distinct and
separated, sometimes one or the other obscured by the
white dusting or by an extension of the ground color,
rarely fused into a line along discocellular vein, blackish
brown; terminal dots more or less distinct. Hind wing
dull, translucent white to smoky fuscous (as a rule
darker on female than on male); a dark shade toward
apex, some dark shading on the veins and a fine dark
line along termen. Eighth abdominal segment of male
simple. Alar expanse, 16-26 mm.
Male genitalia with apical projection of gnathos slen-
der, very long, extending at least as far backward as
apex of uncus (when genitalia are in natural position it
extends well beyond the uncus); transtilla a rather nar-
row, sclerotized band, looped backward in a rounded
arch; sclerotized costa of harpe very slightly and
bluntly produced at apex.
Female genitalia with or without signa, when present
a patch of coarse scobinations, the patch varying in size
in different specimens; ductus bursae simple or very
weakly sclerotized on ventral surface at genital opening.
Type Ltocauities: Cabima, Panamé (muriscis, in
USNM); Cayuga, Guatemala (palpalis, in USNM).
Foop piants: Mammea americana (larvae feeding in
the fruit), Theobroma cacao (larvae in the pods).
Distrisution: Hartt. Puerto Rico, Mayagiiez
(July). Britis West Invres: Trinidad, several exam-
ples with no more specific locality, St. Clair (Mar.);
Grenada, several examples with no more specific locality;
Tobago (Apr.). GuatTemaa: Cayuga (Jan., Feb., Apr.,
May, June), Quirigué (Sept.). Costa Rica: Esperanza
(May, Aug.). PanamA: Alhajuelo (Apr.), Cabima
(May), Porto Bello (Apr., Oct.), Rio Trinidad (Mar.,
June). Cotomera: La Esperanza (Dec.), no specific
locality (June). Bonfivia: “East Bolivia’ (Oct.).
British Guiana: ‘“‘Mazaruni Clearing’? (Aug., Oct.).
Frenca Guiana: Cayenne, St. Jean Maroni, St.
Laurent Maroni. Braziu: Rio de Janeiro (June).
This species is primarily a feeder in the pods of the
cacao and is well distributed in tropical America wher-
ever its host occurs. All specimens in the National
Museum (except the holotypes of muriscis and palpalis)
had been identified by Dyar as ‘‘Myelois transitella
Walker.” The two species are easily confused on super-
ficial characters, especially among faded and stained
tropical specimens; but their genitalia are quite distinct.
Dyar’s types of muriscis and palpalis are males and
alike in genitalic and all other characters. It is very
probable that muriscis eventually will prove to be no
more than a variety (or synonym) of furvidorsella
Ragonot.
46 UNITED STATES NATIONAL MUSEUM BULLETIN 207
81. Ectomyelois furvidorsella (Ragonot), new combination
Figure 681
Myelois furvidorsella Ragonot, Nouv. Gen., p. 8, 1888; Mono-
graph, pt. 1, p. 56, 1893.
This is probably the same as muriscis. 'The genitalia
of the female type (fig. 681) are somewhat unusual in
that bursa and ductus bursae are perfectly smooth, with
no trace of signum or sclerotization of the ductus at
genital opening. However, I have seen similar geni-
talia in typical examples of muriscis from Central
America; but I have seen so few examples of muriscis
(only males) from Puerto Rico that I prefer to keep the
names apart till more material is available.
Alar expanse, 22 mm.
TypE Locauity: Puerto Rico (type in Paris Mus.).
Foop PLant: Unknown.
82. Ectomyelois zeteki, new species
FiaureEs 201, 682
Forewing pale brownish gray faintly dusted with
blackish fuscous; antemedial line obscure, angulate,
indicated chiefly by its blackish fuscous outer border,
the latter incomplete on many specimens; subterminal
line sinuate, rather close to termen, on well-marked
specimens consisting chiefly of white spots on the veins,
preceded and followed by dark streaks, obscure on
many specimens; also on well-marked specimens a me-
dian, longitudinal blackish fuscous streak from base of
wing to antemedial line; discal dots at end of cell very
faint or completely obliterated, when distinguishable
more or less confluent; terminal dark dots faint, con-
fluent. Hind wing translucent, white with a faint
smoky tint towards apex; a dark line along outer margin
and some darkening of the veins. Highth abdominal
segment of male simple. Alar expanse, 17-22 mm.
Male genitalia similar to those of muriscis except:
Uncus narrower; apical projection of gnathos shorter,
not reaching so far backward as apex of uncus; transtilla
a narrow band forming a truncated arch with slightly
concaved posterior margin; sclerotized costa of harpe
projecting somewhat further at its apex. Female geni-
talia with scobinations of signum patch stouter and
sclerotization of ductus bursae at genital opening fore-
wing a larger and more strongly pigmented shield than
those of any other species of the genus.
Type tocatity: Near Capira, Panama (type in
USNM, 61316; paratypes in USNM, Cornell Univ.,
Transvaal Mus. (Janse Coll.), Paris Mus., BM).
Foop puant: Cassia moschata.
Described from male type, and 16 male and 19
female paratypes from the type locality; all reared (May
1941, Zetek No. 4807) by James Zetek, who has con-
tributed much valuable material to the National Collec-
tion, and for whom the species is named.
It is easily distinguished from any other American
phycitid by its male genitalia.
Genera 22-24: Paramyelois to Protomoerbes
[Venational division C. Forewing with 11 veins; 4 and 5 stalked
for less than half their lengths. Hind wing with veins 7 and 8
anastomosed for most of their lengths (free element of 8 short).
Transtilla incomplete; but (except in Paramyelois) its free
elements well developed.]
22. Paramyelois, new genus
Type or Genus: Myelois solitella Zeller.
Tongue well developed. Antenna of male shorily
ciliate (cilia shorter than width of shaft), simple; of
female pubescent. Labial palpus oblique, laterally flat-
tened (broad and flat from lateral view) ; second segment
roughly scaled beneath; third segment shorter than
second, somewhat roughly scaled. Maxillary palpus
squamous (rather heavily and broadly scaled). Fore-
wing smooth; 11 veins; vein 2 from before but rather
near lower outer angle of cell; 3 from the angle, well
separated from the stalk of 4-5 at base, but nearer to
4—5 than to 2; 4 and 5 shortly stalked; 6 from below
upper angle of cell, straight; 8 and 9 long stalked; 10
from the cell, approximate to the stalk of 8-9 for a
short distance from base; male without costal fold.
Hind wing with vein 2 from well before lower outer
angle of cell; 3 from the angle, closely approximate to
or connate with the stalk of 4—5 at base; 4 and 5 nor-
mally stalked for half or slightly less than half their
lengths, rarely (in some small specimens) stalked for
over half their lengths: 7 and 8 strongly anastomosed
for most of their lengths, free element of 8 short; cell
slightly over half the length of wing in male (as in
Kuropean Myelois), half the length of wing in female;
discocellular vein curved. Highth abdominal segment
of male simple.
Male genitalia with apical process of gnathos a short,
rather broad, blunt hook, slightly notched at apex.
Uncus with broad base; narrowed and triangulate just
beyond; apex acutely rounded. Tegumen short and
broad. Transtilla incomplete. Harpe very broad at
base; costa strongly and broadly sclerotized and forming
a broad, pointed projection before middle, not appreci-
ably sclerotized beyond; sacculus large and strongly
sclerotized; cucullus greatly reduced. Anellus a curved
shield with long, strongly sclerotized, smooth, slender,
tapering and pointed lateral arms. Aedeagus scobinate
on one lateral edge at apex; penis with a few sclerotized
wrinklings, otherwise unarmed. Vinculum stout, slightly
longer than broad, truncate, scarcely tapering to broad
terminal margin.
Female genitalia with weak signum consisting of a
cluster of rather coarse scobinations; finer scobinations
scattered over the caudal half of bursa. Ductus bursae
shorter than bursa, broadened and sclerotized on inner
ventral and lateral surfaces towards genital opening; on
inner dorsal surface behind the opening a pair of small
sclerotized plates. Ductus seminalis from bursa near
its junction with ductus bursae.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 47
This genus is easily recognized and is distinguished
from other sections of the ‘‘Myelois’’ complex by its
labial palpi and male genitalia.
In their normal position the palpi are directed in a
straight line obliquely from the face; but many speci-
mens show the third segment more or less deflected for-
ward, and some with both the second and third seg-
ments more or less porrected, results of the death con-
tortions of the moths. Several European species listed
under Myelois have oblique palpi but they are all more
or less cylindrical and do not have the broadly scaled
and flattened lateral aspect of those of Paramyelois.
The male genitalia with their incomplete transtilla (its
elements reduced and well separated) are unique among
the groups nearly related to Myelois or any of the Ameri-
can species that previously have been referred to that
genus.
I have chosen a synonym as type of the new genus
advisedly, as the type specimen of the oldest name
(transitella) is a female, and there may be some question
of my application of the name to the species here treated.
There can be no such doubt in regard to solitella.
83. Paramyelois transitella (Walker), new combination
Figures 202, 683
Nephopteryx transitella Walker, List, pt. 27, p. 54, 1863.
Nephopteryx notatalis Walker, List, pt. 27, p. 57, 1863.
Myelois solitella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 217,
1881.—Ragonot, Monograph, pt. 1, p. 55, 1893. (New
synonymy.)
Myelois duplipunctella Ragonot, N. Amer. Phycitidae, p. 3, 1887;
Monograph, pt. 1, p. 56, 1893.—MceDunnough, Check list,
No. 6059, 1939. (New synonymy.)
Myelois transitella (Walker) Ragonot, Monograph, pt. 1, p. 42,
1893 (in part)—McDunnough, Check list, No. 6058, 1939
(in part).
Myelois venipars Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 404,
1914.—Mote, Monthly Bull. California Dep. Agr., vol. 11,
p. 628, 1922.—Glick, Arizona Comm. Agr. and Hort., Four-
teenth Ann. Rep., p. 78, 1922.—Essig, Insects of western
North America, p. 708, 1929.—Hixon, Journ. Econ. Ent.,
vol. 27, p. 547, 1934. (New synonymy.)
Emporia cassiae Dyar, Ins. Inse. Menstr., vol. 5, 91, 1917
(new synonymy).
Similar in color and maculation to Ectomyelois muris-
cis except: Ground color on lower half of wing darker;
the dark borders of the transverse lines and the discal
dots decidedly darker, blackish; the white areas more
strongly contrasted; the dark outer border of antemedial
complete in most examples and enlarged below costa
into a conspicuous blackish spot. Hind wings a clearer
white on the males; more or less smoky on the females.
Alar expanse, 15-28 mm.
Genitalia as given for the genus. The male genitalia
show little or no individual variation. Among the fe-
males, however, there is considerable variability in mi-
nor details, namely, the amount of sclerotization about
genital opening and the amount of scobination in the
bursa, but these are differences of no specific significance.
TYPE Locauitigs: ‘United States,” probably Florida
(transitella, in BM); Santo Domingo (notatalis, in BM);
Colombia (solitella, in BM); Florida (duplipunctella, in
Paris Mus.); Hermosillo, México (venipars, in USNM);
Georgetown, British Guiana (cassiae, in USNM).
Foop piants: Orange, grapefruit, peach, apple, dates,
figs, Acacia farnesiana, Aesculus glabra, Cassia grandis,
Genipa americana, Gleditsia triacanthos, Pithecolobium
flexicaule, Robinia, Sapindus drummondii, Yucca, English
walnut. These records from reared specimens in the
U.S. National Museum.
Distrisution: Unirep Srarzs: Arizona, Maricopa
County (Dec.), Mesa (Nov.), Phoenix (Aug., Sept.,
Nov., Dec.), Tempe, Yuma; Texas, Anahuae (March),
Brownsville (Dec.), Dallas (May), Fort Davis (Oct.),
Harlingen (July), Hidalgo County (Apr.), Kerrville,
Louise (Feb.), Mercedes (Feb.), Mission, San Antonio
(May), San Benito (Sept.) ; Oklahoma, Stillwater (June) ;
Louisiana, Forbing (Oct.); Alabama, Mobile (Apr.);
Georgia, St. Simons; Florida, Orlando (Oct., Nov.), Vero
Beach (Apr., May); North Carolina, Durham. CuBa.
Dominican Repusiic. Mexico: Hermosillo, Oaxaca.
GuaTEMAta: Cayuga (Mar., Apr.), Chejel (June). Pan-
amMA: El Cermeno (Apr., June). Contompia. Braziu:
Tapera (Pernambuco). Prrt: Lima, Rio Pacaya (June,
July, Aug.).
This species has attracted some attention in the south-
west as a minor orchard pest and is known to economic
entomologists as the “‘nayel-orange worm.” The larvae
feed on the nuts, in the seed pods, or on the fruits of
numerous trees but they seem to prefer the fallen and
mummied fruits or the dry seed pods or injured or
diseased fruits. Rarely do they attack sound fruit on
the trees. They have been reported as infesting sound
oranges, but such behavior is probably an exceptional
departure from normal habit.
The foregoing synonymy requires some comment. I
have not seen the types of transitella or duplipunctella
(both females), but from the original. descriptions and
the Ragonot figures they cannot apply to anything else
than the species we have hitherto known in the United
States as venipars Dyar. Of the synonymy of venipars
and solitella there is no possible doubt. The type of the
latter is a male (not a female as stated by Zeller) and
figures of its genitalia, supplied by Tams and Clarke,
show agreement in every detail with those of venipars.
Clarke has also furnished excellent photographs of the
female types of transitella and notatalis and of the geni-
talia of transitella. The latter show only trifling indi-
vidual differences from the genitalia of Dyar’s female
type of venipars. Unfortunately, the type of notatalis
lacks an abdomen; but photographs of the moths and
their palpi show no essential differences between the two
types; so Ragonot’s reference of notatalis to synonymy
must be accepted. In his specific key to the species of
“Myelois” Ragonot (Monograph, pt. 1, p. 27) places
transitella in a group with veins 7 and 8 of hind wing
approximate. This characterization was obviously
based upon a freak specimen. Dyar found one such
freak (a female from Grenada) among the examples of
muriscis which he misidentified as “‘transitella”’; but in
over a hundred examples of the true transitella before
me veins 7 and 8 are strongly anastomosed, and this is
48 UNITED STATES NATIONAL MUSEUM BULLETIN 207
a good character of much more than specific value
despite its lapse in individual specimens. Freaks of all
kinds can and do turn up anywhere in the Phycitinae.
Dyar’s cassiae was described from stained and faded
females; but, even so, it is strange that he did not see
their resemblance to his venipars, especially in their
palpi, and still more strange that he should refer them
to the Old World anerastiid genus H#mporia. They have
normal phycitine tongues and their genitalia agree with
those of the female type of venipars.
Bondar’s (Instituto de Cacau da Bahia Boletim 5,
p- 72, 1939) identification of a lepidoperon in cacao pods
as duplipunctella Ragonot (the genus given as “Myel-
osis’”’) is probably incorrect. What he had was pre-
sumably Hetomyelois muriscis.
23. Genus Pseudodivona Dyar
Pseudodivona Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 405, 1914.
(Type of genus: Pseudodivona commensella Dyar.)
Tongue well developed. Antenna shortly ciliate on
male, cilia about as long as width of shaft (longer on
carabayella). Labial palpus oblique, broadly scaled and
laterally flattened; third segment short, acuminate.
Maxillary palpus squamous. Forewing smooth; 11
veins; vein 2 from before lower outer angle of cell; 3
from the angle; 4 and 5 stalked for slightly less than
half their lengths, the shaft separated at base from 3;
6 from below upper angle of cell, slightly curved towards
base; 8 and 9 stalked for two-thirds their lengths; 10
from the stalk of 8-9; 11 from cell rather near outer
angle and running close to the stalk of 8-9-10; male
without costal fold. Hind wing from well before lower
outer angle of cell; 2 from very close to the angle, or
from the angle (cispha), closely approximate to or con-
nate with stalk of 4—5 at base; 4 and 5 stalked for about
half their lengths; 7 and 8 anastomosed for most of their
lengths (free element of 8 very short); cell less than one-
half (but more than a third) the length of wing; disco-
cellular vein curved. Highth abdominal segment of
male with a strong pair of ventrolateral hair tufts.
Male genitalia with apical process of gnathos de-
veloped as an elongate, stout, flattened hook with
forked or notched apex. Uncus subtriangulate, with
rounded terminal margin. Transtilla incomplete; its
elements well developed, elongate and knobbed at their
apices. Harpe simple with outer margin evenly rounded;
costa sclerotized for about four-fifths its length; but not
produced. Anellus a narrow curved band with slender
lateral lobes. Aedeagus moderately slender, nearly
straight; penis unarmed. Vinculum stout, decidedly
longer than broad, tapering, expanded towards angulate
terminal margin.
Female genitalia without signum; bursa, ductus bur-
sae, and genital opening simple; ductus seminalis from
bursa near junction of bursa and ductus bursae.
In genitalic and many other characters as well as
wing pattern and color, this genus resembles Moerbes,
to which it is apparently closely related. It differs
chiefly in having vein 4 present and well developed in
hind wing, a different development of the elements of
transtilla, and strong hair tufts on the eighth abdominal
segment of the male. Four tropical American species
are recognized. How many of these are really distinct
species it is impossible to determine from the scanty ma-
terial available. Nothing is known of their biology or
habits.
84. Pseudodivona commensella Dyar
Fieures 41, 204
Pseudodivona commensella Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 406, 1914.
Forewing dull white on area above lower margin of
cell; the area from cell to inner margin a glossy vinous
brown; a blackish brown smudge on costa at base;
antemedial line obscure except where it cuts the brown
shade and forms a contrasting white spot at inner mar-
gin, bordered outwardly by a blackish brown, out-
wardly angled line, obscure on all but fresh specimens
and frequently broken into two blackish spots, one on
costa, the other in the cell; subterminal line indicated
by a pair of narrow and narrowly spaced blackish dashes
from costa near apex and some inwardly bordering
blackish spots or streaks on the veins; the veins other-
wise more or less darkly streaked; lower discal spot at
end of cell enlarged, blackish, more or less confluent
with a smaller, much fainter upper spot; a row of black-
ish dots along termen. Hind wing pale, glossy, semi-
translucent grayish white; veins darkened and clearly
outlined; a narrow dark line along termen. Alar
expanse, 20-22 mm.
Male genitalia with trifling differences in the shape of
the apical projection of gnathos between this and the
following species of the genus and some differences in
the curve of the outer margin of the harpe, but I suspect
that these differences are individual rather than specific
in character.
TypEr Locatity: Jalapa, México (type in USNM).
Foop Prant: Unknown.
Distrisution: Mexico: Jalapa, Orizaba.
Known only from the four males of Dyar’s type
series.
85. Pseudodivona cispha Dyar
Fieure 205
Pseudodivona cispha Dyar, Ins. Insc. Menstr., vol. 7, p. 53, 1919.
A smaller, less distinctly marked species than the
preceding one (commensella); the brownish area of
forewing narrower and paler (not “reddish” as stated
by Dyar); the dark markings fewer, fainter and paler
and, except for the spot on base of costa and a short
wedge at apex, not blackish; discal dots inconspicuous,
light brown, the lower dot much smaller than on com-
mensella. Alar expanse, 16-18 mm.
Female genitalia like those of P. santa-maria.
TyprE LocaLity: Volcin Santa Maria, Guatemala
(type in USNM).
Foop PLaAnt: Unknown.
DistrisutTion: GUATEMALA: Cayuga (Aug.), Voleén
Santa Maria (July, Oct.). Costa Rica: Tuis (May).
British Honpuras: Rio Grande (Sept.), Punto Gorda
(July).
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 49
86. Pseudodivona santa-maria Dyar
Ficure 690
bea cee santa-maria Dyar, Ins. Insc. Menstr., vol. 7, p. 54,
1919
Known only from the two females of the type series.
The coloration and markings are more like those of
commensella except that the dark striping of the veins is
fainter and the discal dots smaller, paler and less con-
spicuous, like those of cispha. It is quite possible that
these specimens are only larger, darker, better marked,
female examples of cispha and equally possible that
both cispha and santa-maria are only varieties of com-
mensella. Alar expanse, 21 mm.
Type tocatity: Volcfn Santa Maria, Guatemala
(July; type in USNM). Paratype from Cayuga,
Guatemala (May).
Foop pLant: Unknown.
87. Pseudodivona carabayella Dyar
Ficurss 206, 691
Pseudodivona carabayella Dyar, Ins. Insc. Menstr. vol. 7, p. 54,
1919
Larger and more strikingly marked than any of the
preceding species. Forewing with pale areas pure
white; lower area of wing (between cell and inner mar-
gin) vinous brown (not “purplish red’’ as in Dyar’s
original description) except for an extension of the
white behind the antemedial line where it reaches
almost to inner margin; an elongate black patch on
costa at base; antemedial line indicated by the usual
white spot on inner margin and its black outer border,
the latter is rather broad from costa, strongly angled
and extends from costa to the lower margin of the cell;
a subbasal black spot in the cell; subterminal line indi-
cated above cell only by its widely spaced black inner
and outer borders; the inner black border a strong slant-
ing black dash extending from costa almost to the lower
discal dot at end of cell (indicating a deep angulation of
the subterming line); the outer black border a much
shorter, slanting, dash from apex to vein 6; from about
vein 5 the subterminal line is indicated by a faint white
line through the brownish ground color and is bordered
inwardly by a few rather faint blackish spots; discal
spots, distinct, black and somewhat enlarged, the lower
one particularly. Hind wing white, more or less tinted
with grayish on some specimens; veins darkly outlined;
a narrow dark line along termen. Alar expanse, 23-28
mm.
Apical process of gnathos of male genitalia figured
from type. Another male from Incachaca, Bolivia (in
BM), exhibits some variation from the type in the shape
of the apical process of gnathos (fig. 206a). It is a
small specimen (23 mm.) and seems to have the dark
areas and markings of forewing paler, but it is a rubbed
and faded example; other specimens from the Schaus
Collection in the National Museum and from the same
Bolivian locality are typical in all details. The British
Museum specimen is probably nothing more than an
individual variant.
TYPE LocALITy: Oconeque, Carabaya, Peri (type in
USNM).
Foop pLrant: Unknown.
DistrisutTion: Pert: Carabaya, Oconeque, Tinguri.
Bourvia: Cochabamba,¥Incachaca. Conompra: San
Antonio (Dec.).
All Peruvian examples in the U.S. National Museum,
British Museum, and Janse Collection are males. The
only female of the species that I have seen is the
specimen from Incachaca, Bolivia, from which the
genitalia are figured.
The species is undoubtedly a distinct one. The
widely spaced, strong, black dashes bordering the sub-
terminal line indicate this as well as the ciliations of the
male antenna, which are longer than those of any of
the preceding species, being somewhat longer than the
width of the antennal shaft.
24, Protomoerbes, new genus
Typr or genus: Protomoerbes aberrans, new species.
Characters of Pseudodivona except: Labial palpus
upturned (but otherwise as in Pseudodivona) ; forewing
with vein 3 closely approximate to the stalk of 4-5 at
base, male with narrow costal fold; hind wing with veins
4 and 5 stalked for at least three-fourths of their length,
cell one-third the length of wing; eighth abdominal
segment of male without hair tufts; transtilla of male
genitalia incomplete, its elements elongate-angulate,
their apices not knobbed or expanded.
In many details this genus is more like Moerbes of
Group II than Pseudodivona. It differs from both
genera in its upturned rather than oblique palpi. The
cilia of the male antenna are also shorter (slightly less
than the width of the shaft), but this is hardly a generic
character. Wing pattern, color, and general habitus
are like those of both Pseudodivona and Moerbes. All
three have the contrasting white spot on inner margin
of forewing indicating the base of the antemedial line.
Protomoerbes in every way seems to be an intermediate
and connecting link between Pseudodivona and Moerbes.
It is represented by only two species from Colombia.
Their females are unknown.
88. Protomoerbes aberrans, new species
Fiaure 208
Forewing white; basal area, median area below cell
and outer area below apex shaded with pale brown; a
yellow longitudinal median streak from base to end of
cell cutting the antemedial line; antemedial line a con-
spicuous white spot on inner margin and a fainter white
spot on costa, bordered outwardly below costa and on
inner margin by blackish scaling and inwardly by a
subcostal black streak reaching nearly to base of wing
and by scattered black dusting at inner margin; veins
and lower fold beyond antemedial more or less streaked
or dusted with black, the black streaks especially marked
and angulate at inner margin of subterminal line; lower
discal dot expanded and extended along lower vein of
cell, black; upper discal dot not distinguishable; sub-
50 UNITED STATES NATIONAL MUSEUM BULLETIN 207
terminal white line narrow, faint, sinuate and sharply
dentate below costa, bordered at costa by a pair of
narrow, short, blackish dashes; a row of separate black
dots along termen. Hind wing white, translucent.
The veins faintly darkened and a narrow dark shade
along termen. Alar expanse, 27-30 mm.
Male genitalia with aedeagus smooth and evenly
tapering from base.
TypE Locauity: Colombia (type in USNM, 61317;
paratype in BM).
Foop prant: Unknown.
Described from male type and two male paratypes
labeled ‘‘Colombia, Fassel.’? The specimens are only
in fair condition. Fresher examples would probably
show the black longitudinal streakings somewhat more
contrasted and extended. ‘The species has the general
habitus of Pseudodivona commensella but is larger. It
is easily separated from commensella and all the other
similarly colored and marked species of Pseudodivona
and Moerbes by the yellow longitudinal median streak
on forewing. ‘This is easily distinguished under slight
magnification, even on slightly rubbed specimens.
89. Protomoerbes separabilis, new species
Fieure 207
Similar to aberrans except: Less distinctly and exten-
sively streaked with black; median yellow longitudinal
streak missing from forewing, replaced by a narrow
extension of the white ground color, extended to the
base of the wing; aedeagus with a thornlike projection
from underside near apex (similar to the projection
from the aedeagus of Moerbes emendata but larger and
more blunt). The male genitalia also differ from those
of M. emendata in having a proportionally longer and
more evenly tapering vinculum. Hind wing smoky
white, semitranslucent; the veins distinctly darkened.
Alar expanse, 29 mm.
TYPE Locatity: San Antonio, Colombia (type in BM;
paratype in USNM, 61318).
Foop prant: Unknown.
Described from male type and one male paratype
from the type locality, labeled ‘San Antonio, W.
Colombia, Dec. 07, 5800 feet, M. G. Palmer.’ The
paratype lacks an abdomen.
Genera 25 and 26: Diatomocera and Pseudocabima
[Venational division C. Forewing with 11 veins; 4 and 5 stalked
for less than half their lengths. Hind wing with veins 7 and 8
anastomosed for most of their lengths (free element of 8 short).
Transtilla complete but weakly sclerotized. Uncus spoon- or
semispoon-shaped.]
25. Genus Diatomocera Ragonot
Diatomocera Ragonot, Monograph, pt. 1, p. 250, 1893.
genus; Homoeosoma tenebricosa Zeller.)
Cabima Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 329, 1914.
(Type of genus: Cabima dosia Dyar. New synonymy.)
Tongue well developed. Antenna of male shortly
ciliate (the cilia about as long as width of shaft); the
(Type of
shaft with notch at base; of female simple and pubes-
cent. Labial palpus of male upturned, reaching to or
almost to vertex, slender; third segment nearly as long
as second, acuminate; palpus of female oblique and
slightly longer than that of male. Maxillary palpus
filiform. Forewing smooth; 11 veins; vein 2 from well
before lower outer angle of cell; 3 from the angle; 4 and
5 shortly stalked (for half or less than half their lengths),
the stalk separated from 3 at base; 6 from below upper
angle of cell, very slightly bent towards base; 8 and 9
long stalked; 10 from the cell, but approximate to the
stalk of 8-9 for some distance; male with an elongate,
narrow costal fold. Hind wing with vein 2 from well
before lower outer angle of cell; 3 from the angle, connate
with the stalk of 4-5; 4 and 5 long stalked; 7 and 8
anastomosed for most of their lengths beyond cell, free
element of 8 short; cell about half the length of the cell
on male, on female longer; discocellular vein curved.
Highth abdominal segment of male with a pair of
ventrolateral hair tufts.
Male genitalia with apical process of gnathos U-
shaped, consisting of a pair of short, blunt arms. Uncus
spoon- or semispoon-shaped. ‘Transtilla complete but
weakly sclerotized (except along its lateral edges), con-
sisting of a broad, more or less finely scobinate plate;
weakly attached to harpes. Harpe simple; costa sclero-
tized for most of its length, but not produced. Anellus
a curved plate with short lateral lobes. Aedeagus stout,
straight (or but slightly bent near middle), more or less
tapering to apex, moderately long; penis with a few
weakly sclerotized wrinklings and more or less finely
spined, otherwise unarmed. Vinculum stout, elongate,
constricted towards angulate (or narrowly rounded)
terminal margin.
Female genitalia with bursa more or less finely
scobinate; signa present, consisting of a cluster of two
or more sclerotized disks; ductus bursae unsclerotized;
genital opening simple; ductus seminalis from bursa in
the neighborhood of the signa (sometimes between them
and the junction of bursa.and ductus bursae, but not
near the junction).
The genus is readily recognized by its combination of
male characters, the most striking feature of which is
the spoon= or semispoon-shaped uncus which is found
in only two other American genera—Pseudocabima,
which lacks the antennal notch and long costal fold of
forewing, and Entmemacornis, which has veins 4 and 5
of hind wing completely united. Diatomocera is ap-
parently confined to tropical America. Nine species
are here recognized. They are represented by scanty
and scattered material and nothing is known of their
life histories or habits. From the greasy condition of
some of the specimens it may be assumed that the larvae
are borers.
90. Diatomocera tenebricosa (Zeller)
Fiaures 42, 209, 565, 686
Homoeosoma tenebricosa Zeller, Horae Soc, Ent. Rossicae, vol. 16,
p. 242, 1881.
Diatomocera tenebricosa (Zeller) Ragonot, Monograph, pt. 1, p.
250, 1893.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 51
I have not seen any specimens from the type locality
but have before me a male and female from French
Guiana and a somewhat larger pair (20-21 mm.) from
Costa Rica compared by Schaus with the type. They
are either this species or varieties thereof. The follow-
ing description is drawn from them.
Ground color of forewing gray shaded with reddish
brown between the veins; the veins themselves outlined
with black, the blackish streaks broken by a very faint
dull whitish antemedial line and by a more distinct
subterminal line and more or less interrupted between;
at extreme base the vein markings fused into a blackish
patch; subterminal line close to outer margin, out-
wardly angled between the fork of veins 4 and 5; discal
dots small, separated, set obliquely, blackish; terminal
dots faint. Hind wings grayish brown, paler and semi-
translucent on the male; the veins faintly darkened,
brown; a narrow brown line along termen. Alar ex-
panse, 16-21 mm.
Male genitalia with uncus but slightly constricted
near its middle (wider in this area than in any of the
following species); penis finely spined. Female geni-
talia with signum a small cluster of bluntly rounded,
closely appressed disks.
Typ tocatity: Honda, Colombia (type in BM).
Foop prant: Unknown.
Distrisution: Cotomsr1a: Honda. Frencu Guiana:
Cayenne. Costa Rica: Juan Vifias (May), Sixaola
River (Mar.).
The species is easily distinguished from anything else
in the genus by its smaller size and the shape of its
uncus. There are several minor differences between the
genitalia of specimens from French Guiana and Costa
Rica; in the spacing between the apical prongs of gna-
thos, in the shape of the terminal margin of vinculum,
in the amount of spining on the median area of tran-
stilla, and in the number of disks forming the female
signum. These are shown in our figures. Such differ-
ences, however, do not seem to be of anything more
than individual or (at most) varietal significance. The
foldings of the median (membranous) area of the tran-
stilla shown in figure 209 are superficial and result from
the manner in which the slide preparation was made.
91. Diatomocera dosia (Dyar), new combination
Fiaures 48, 210, 685
Cabima dosia Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330, 1914.
Forewing dull white; the veins streaked with black
broken into short dashes and dots; antemedial line not
defined except by the black vein streakings on outer
margin; subterminal line faint, angulate, the apex of
angle between the fork of veins 4 and 5; discal and ter-
minal dots distinct, separate, black; a faint shading of
ocherous fuscous scales above and below vein 1b at
middle; costa at base black for most of the length of
fold on male. Hind wing dull semitransparent white,
slightly darker on female; the veins more or less out-
lined in pale brown; a smoky shade along costa and a
fine, brown line along termen. Alar expanse, 24-31
mm.
Male genitalia with penis finely granulate-scobinate.
Female genitalia with signum a chain of bluntly pointed
disks.
Typ tocatity: Cabima, Panama (type in USNM).
Foop piant: Unknown.
Known only from the type series (nine specimens)
from the type locality (May).
92. Diatomocera excisalis (Hampson), new combination
Ficurzs 211, 687
Crocidomera excisalis Hampson, Ann. Mag. Nat. Hist., ser. 10,
vol. 4, p. 353, 1929.
Similar to dosia Dyar and probably no more than a
variety of that species. Distinguished by its generally
smaller size, some trifling differences in male genitalia,
and a shorter chain of disks forming the signum of the
female.
Alar expanse, 23-25 mm.
Male genitalia considerably smaller than those of
dosia but otherwise similar.
Typ LocaLity: Cayenne, French Guiana (type in
BM).
Foop piant: Unknown.
DistrisutTion: FrencH Guiana: Cayenne, St. Lau-
rent Maroni.
Also before me is a female from the unidentified ma-
terial of the British Museum from eastern Bolivia
(“‘Aug.—Oct., 1920, T. Steinbach’’) which is superficially
a very good match for the female paratype of excisalis
and may be a variety of that species. Unfortunately
it lacks an abdomen, so positive identification cannot
be made.
93. Diatomocera decurrens (Dyar), new combination
Ficure 212
Cabima decurrens Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 330,
1914,
Forewing ‘‘luteous gray’ (the ground color of a dis-
tinctly yellowish tint); black markings on veins as in
excisalis but especially strong along vein 1b; the sub-
terminal line somewhat more distinct. Hind wing semi-
transparent white with a very faint ocherous tint; veins
very faintly darkened; a narrow, pale brown line along
termen. Alar expanse, 21-28 mm.
Male genitalia distinguished from those of dosia and
excisalis by its much narrower (more constricted)
vinculum; penis with a few weak, minute scobinations.
The sternite of eighth abdominal segment is also differ-
ently shaped from that of dosia or excisalis.
Type Locauiry: Rio Trinidad, Panamé (type in
USNM).
Foop prant: Unknown.
DistriputTion: Panami: Corazal (Mar.), La Chorrera
(May), Rio Trinidad (Mar.).
A distinct species easily distinguished by its male gen-
italia and ocherous forewing. Known only from males
of the original type series. I fail to see the difference
52 UNITED STATES NATIONAL MUSEUM BULLETIN 207
in the antennal notch from that of dosia which Dyar
mentions. His examination of the material before him
was obviously cursory for he refers to three of the La
Chorrera specimens as ‘“‘females.’’
94. Diatomocera majuscula, new species
Fiaure. 213
Forewing pale, dull, brownish gray; the veins outlined
by dark brown; entire basal area to antemedial line suf-
fused with blackish brown; antemedial line indicated by
three detached dull-white marks, a spot on costa, a
smaller one in cell, and a third, somewhat diffused, white
smudge on inner margin; subterminal line indicated by
a white spot on costa, a white spot on inner margin and
a much fainter, pale, outwardly curved line cutting the
darkened veins from vein 1 to vein 5; discal spots dis-
tinct, separated, black. Hind wing white, transparent;
the veins very slightly darkened; a gray brown shade
along costa, and a narrow pale-brown line along termen.
Alar expanse, 32 mm.
Male genitalia distinguished chiefly by their large
size. Highth segment tufts of abdomen also more ro-
bust than those of any other species of the genus.
Typz Locatiry: Ponta Nova, Rio Xingu, Amazonas,
Brazil (type in USNM, 61319).
Foop prant: Unknown.
Described from male type, from the Dognin Collec-
tion in the U. S. National Museum, that had been iden-
tified as Sematoneura atrovenosella. The species can be
easily identified by its large size and the contrasted,
blackish basal area of forewing.
95. Diatomocera albosigno, new species
Figure 214
Forewing dull ocherous brown; the veins outlined in
black; broken outwardly by a faint subterminal line
which is indicated by a small white spot on costa, a
smaller, much weaker spot on inner margin and a very
faint, pale outcurved line between veins 1 and 6; ante-
medial line replaced by a large white blotch, as broad as
long and extending from inner margin to or almost to
costa, bordered outwardly by a narrow blackish line;
discal spots replaced by a narrow blackish line along
the discocellular vein, Hind wing dull, smoky white,
semitranslucent; veins faintly darkened; a thin fuscous
line along termen. Alar expanse, 21-23 mm.
Typr tocanity: “S. E. Brazil” [probably Paran4]
(type in BM; paratype in USNM, 61320).
Foop prant: Unknown.
Described from male type and one male paratype
from the type locality labeled ‘‘S. E. Brazil, E. D. Jones,
1920-303.”’ Hasily recognized by the large white spot
on the subbasal area of forewing; the only species of
Diatomocera so marked.
96. Diatomocera hoplidice (Dyar), new combination
Fieure 215
a
Cabima hoplidice Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330”
1914.
Forewing dark gray; unicolorous except for faint
darker (blackish) shading on some of the veins and a
dark edging to the costa; antemedial and subterminal
lines obsolete; discal dots replaced by a very faint dark
line along the discocellular vein. Hind wings very dark,
smoky gray (almost black). Alar expanse, 26 mm.
Male genitalia with aedeagus broad and abruptly
tapered toward apex.
TypE LocALity: Puerto Bello, Panamé (Mar.; type
in USNM).
Foop prant: Unknown.
Known only from the male type. Hasily recognized
because of its nearly uniform dark coloration and lack
of any transverse markings on forewing.
97. Diatomocera extracta, new species
FI@urREsS 217, 688
Ground color of forewing gray with a faint ocherous
tint, especially in basal area; veins darkened; antemedial
line well out, near middle of wing, vertical, bordered
outwardly by a narrow reddish brown line and in-
wardly by a fainter line of the same color; subtermina
line indicated only by breaks in the dark lining of some
of the veins; discal spots replaced by an oblique brown
line along discocellular vein. Hind wing semitranslu-
cent, white with a very faint grayish ocherous tint,
slightly darker on female; veins very faintly darkened;
a fine dark (brownish) line along termen. Alar expanse,
21-25 mm.
Male genitalia with harpe narrow, not expanded to-
ward outer margin; aedeagus slightly bent; penis armed
with an elongate, dense cluster of slender dark spines
and a scattering of fine scobinations. Female genitalia
with bursa transversely elongate, weakly scloerotized in
the area about the signa and junction of ductus semin-
alis, also finely scobinate in this area, the scobinations
extending into the ductus bursae; signa consisting of
two or three very narrow, thin, elongate disks.
TypE Locatity: Tuis, Costa Rica (type in USNM,
61321).
Foop pirant: Unknown.
Described from male type from the type locality, col-
lected by W. Schaus (“May 28—June 4”), and one male
and one female paratypes, collected by Schaus and
Barnes (Nov.).
The species is closest to but quite distinct from moch-
lophleps Dyar. Most nearly resembles pale or faded
examples of Pseudocabima rubrizonalis (Hampson), with
which it was confused in the National Collection.
98. Diatomocera mochlophleps (Dyar), new combination
Fieurss 216, 689
Cabima mochlophleps Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 404, 1914
Forewing purplish gray with faint pale reddish brown
shading between the veins; the area from base to near
middle slightly paler, its outer margin faintly indicating
an antemedial line, somewhat curved and inwardly
oblique from costa; subterminal line well inward from
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 53
termen, angulate with apex of the angle within the fork
of veins 4—5, appears denticulate due to pale interrup-
tions on the veins, preceded by short black dashes on
the veins; veins otherwise faintly blackish; a distinct
and characteristic discal mark beginning as a black
streak or dot at lower outer angle of cell and continued
as a curved line along discocellular vein and for a short
distance inward along upper vein of cell; a row of black
dots along terminal margin. Hind wing translucent,
white; the veins darkened, a smoky shade along costa
and a narrow blackish line along termen. Alar expanse,
31 mm.
Male genitalia similar to those of extracta, except api-
cal part of uncus broader and vinculum stouter. Fe-
male genitalia with bursa globular; the signa consider-
ably larger and more triangulate than those of extracta.
Typr Locauity: Zacualpan, México (type in USNM).
Foop prant: Unknown.
In addition to the male type and a female from the
type locality there is before me a smaller (27 mm.) male
from the Janse Collection labeled simply ‘‘Mexico.” It
is more suffused, lacks the pale basal shade on forewing,
and shows scarcely any trace of asubterminal line. The
black discal mark, however, is present and strongly con-
trasted. This is the characteristic feature of the species.
It resembles an inverted comma, with the tail pointed
towards the base of the wing.
26. Pseudocabima, new genus
Typs or GENUS: Myelois euzopherella Dyar.
Characters of Diatomocera except: No notch in shaft
of male antenna; forewing of male without costal fold;
hind wing with vein 3 frequently stalked with the stalk
of veins 4-5 (apparently not a constant specific char-
acter). In one species (rubrizonalis) the apical process
of gnathos differs from that of any other species of either
Pseudocabima or Diatomocera in that it is developed as a
flattened hook with cleft apex and not as a U- or V-
shaped pair of prongs. Such a departure from type is
unusual within generic limits, but of no more than spe-
cific significance, because the species otherwise is per-
fectly normal.
The new genus is proposed with great reluctance; for
its species are very closely related to the bulk of those in
Diatomocera, though none has been previously associated
with them; but some separation must be made if we are
to define our superspecific groups with any exactness.
Pseudocabima, Diatomocera, and Entmemacornis are all
obviously closely related, but they are separable on con-
sistent, if slight, structural differences.
Ten species are here recognized as belonging to the
genus. One of these is North American (arizonensis).
The remainder are tropical and probably only a fraction
of the species inhabiting Central and South America.
Before me are five single examples of what appear to
be as many new species. It seems advisable to leave
them undescribed until more material is available, their
sexes can be associated, and more is known about the
individual and local ranges of variability within species
of the genus. There seems to be some variability, both
in color and in minor details of genitalic structure.
99. Pseudocabima castronalis, new species
Figures 218, 696
Forewing gray, the ground color lightened by white
dusting over much of the median area; antemedial line
distinguishable throughout, whitish, nearly vertical,
slightly out-bent from before middle of costa to middle
of cell, thence slanting inwardly very slightly to vein 1b
and thence outwardly to the inner margin, followed on
inner margin by a dark blotch; subterminal line from
outer fourth of costa, bluntly angulate at middle (the
line more curved than angled between vein 6 and lower
fold) ; discal dots replaced by a pale ocherous brown spot
covering the discocellular vein; fainter extension of this
brownish shade between some of the veins in outer area,
especially near costa beyond and before the subterminal
line; some blackish streaking on the veins, conspicuous
as three short black streaks following the brownish dis-
cal spot; hind wing smoky white, semitranslucent on
the male; darker, more brownish on female; the veins
but faintly darkened. Alar expanse, 23-27 mm.
Male genitalia with aedeagus tapering to narrow
apical end; penis without spining or scobinations. Fe-
male genitalia with signum a curved chain of more or
less bluntly pointed disks.
Type Locauity: Castro, Parand, Brazil (type in
USNM, 61322; paratypes in BM).
Foop prant: Unknown.
Described from male type from the type locality, and
one male paratype and one female paratype labeled
“S.E. Brazil, E. D. Jones, 1920-303,” the latter two from
unplaced material in the British Museum. They are
somewhat discolored and consequently appear more
yellowish than the type, which is in better condition,
showing no trace of grease. This specimen was de-
posited in the National Museum in 1905 by Schaus as
representative of a Hampson manuscript species. It
bears the name label (in Hampson’s handwriting):
“Coptarthria castroalis Hampson, Type o.’’ Apparently
Hampson never published a description.
The best character for recognition of the species is
the ocherous-brown discal spot with its outwardly bor-
dering contrasted black streaks.
100. Pseudocabima fearnella (Schaus), new combination
Figure 219
Myelois fearnella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11,
p. 245, 1913.
Forewing gray, a reddish brown shade along lower
fold and some dusting of the same color along inner mar-
gin; antemedial line obsolete; subterminal line faint but
distinguishable, bent as in castronalis, pale gray; veins
discontinuously lined with black; discal spots black,
separated or (at most) partially fused. Hind wing
white, semitransparent; the veins slightly darkened;
a darker more distinct line along termen. Alar expanse,
19-23 mm.
54 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Male genitalia differing only in trifling details from those
of castronalis, apical part of uncus being slightly broad-
er and a trace of fine scobination appearing on the
penis. The female is unknown.
TypE LocaLity: Avengarez, Costa Rica (July; type
in USNM).
Foop prant: Unknown.
In addition to the male type there are before me two
males from Cayuga, Guatemala (Feb., June). They
are smaller than the type and somewhat rubbed and
faded and, as a result, considerably paler. However,
they agree in all other details. The species is evidently
very close to castronalis.
101. Pseudocabima guianalis, new species
Fieurrs 220, 697
Forewing pale brownish gray; some darker dusting
for a wide area along inner margin; veins very faintly
and irregularly outlined by dark scaling thickened at
lower angle of cell, the curvation of the line inward;
antemedial line obsolete, very faintly indicated on one
or two specimens; subterminal line obscure, more
sharply angulate than on preceding species. Hind
wing translucent white; veins partially and faintly
darkened; a pale smoky brown line along termen. Alar
expanse, 25-33 mm.
Male genitalia with uncus narrower in middle than
that of castronalis or fearnella; penis armed with a
cluster of very fine weak spines and scobinations. Sig-
num of female genitalia a short cluster of blunt, thorn-
like disks.
TyPE Locatity: St. Jean Maroni, French Guiana
(type in USNM, 61323; paratypes in BM and Cornell
Univ.).
Foop prant: Unknown.
Described from male type and one male and two fe-
male paratypes from the type locality, collected by
W. Schaus; one female paratype from Rockstone, Hs-
sequebo, British Guiana (Schaus); one male paratype
from Tumatumari, Potaro River, British Guiana, June
26, 1927 (Cornell Lot 760, sub. 114); one male and one
female paratypes from Mackenzie, Demerara River,
British Guiana, June 21, 22, 1927 (Cornell lot 760, sub.
102, 104). One of the female paratypes from St. Jean
Maroni bears a Hampson “cotype” label inscribed
“Coptarthria guianalis.”’ Evidently another example
of an undescribed Hampson species.
102. Pseudocabima euzopherella (Dyar), new combination
Figures 228, 693
Myelois euzopherella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 326,
1914.
Forewing gray more or less dusted with reddish brown
(the specimens inclined to be greasy, giving a distinctly
brownish shade to the wing); transverse lines distinct,
whitish; antemedial line nearly vertical, a slight bend
in cell, a narrow dark, outer, bordering line; subterminal
line with a similar dark inner border, slightly and
bluntly angled or rounded at vein 5; the distinctive
mark a round blackish brown smudge, touching outer
margin of cell, consisting of a black discal dot surround-
ed by blackish or brown smudges; some broken black
streaks faintly indicated on the veins. Hind wing pale
smoky fuscous, subtranslucent; veins darkened; a fine
dark line along termen. Alar expanse, 19-24 mm.
Male genitalia with central area of uncus very narrow;
penis armed with numerous fine spines and scobinations.
Female genitalia exhibiting no distinctive specific char-
acters; signum a row of rather stout, thornlike disks.
TypE tocaLity: Rio Trinidad, Panamé (type in
USNM).
Foop piant: Unknown.
Distrisution: PanamMA; Cabima (May), Corozal
(May), Tabernilla, Rio Trinidad (June).
Vein 3 of hind wing seems to be consistently from the
stalk of veins 4 and 5, though the amount of stalking
varies in different specimens. In the forewing, 10 is
from the cell, approximate to the stalk of 8-9 but not
from it as stated by Dyar.
103. Pseudocabima pombra (Dyar), new combination
Fiaure 221
Myelois pombra Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 326,
1914.
Forewing pale ocherous gray; transverse lines faint,
whitish, the antemedial near middle of wing, vertical,
the subterminal rather well back from termen, slightly
angled at middle; no discal markings; some scattered
dark (brownish) dusting on the veins. Hind wing con-
colorous with forewing, semitranslucent; the vems not
appreciably darkened. Alar expanse, 17 mm.
Male genitalia similar to those of ewzopherella except
smaller and stem of uncus more slender, differences of
doubtful specific value.
Typr Locatiry: Cabima, Panamdé (May; type in
USNM).
Foop prant: Unknown.
Known only from the male type. Evidently close
to and possibly only a pale suffused variety of ewzophe-
rella. The type is somewhat rubbed. On a fresh
specimen in better condition the dark shading on the
veins of forewing presumably would be more apparent.
104. Pseudocabima nigristrigella (Ragonot), new combination
FIGures 222, 699
Myelois nigristrigella Ragonot, Nouv. Gen., p. 7, 1888; Mono-
graph, pt. 1, p. 41, 1893.
Forewing white dusted with blackish, giving it an
ashy gray color to the naked eye; under magnification
some rust colored scaling between the veins especially
above and along inner margin; antemedial line near
middle of wing, nearly vertical (very slightly convex),
thin, white, bordered outwardly by a narrow, strongly
contrasted, blackish brown line; subterminal line cury-
ing outward from costa to between veins 4-5, thence
vertical to inner margin, white, bordered inwardly by a
narrow blackish brown line; a black line along dis-
cocellular vein and continued for a short distance along
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 55
vein 3; a faint concentration of the black dusting on the
other veins. Hind wing semitransparent, more or less
smoky white; darker on female than on male; the veins
darkened, a broad dark shade along costa and a narrow
dark line along termen. Alar expanse, 22-24 mm.
Male genitalia figured from type. On another male
in the British Museum from the type locality (June) the
stem of uncus is broader, the terminal part of vinculum
less constricted and its terminal margin less sharply
angled. (Another example of the amount of individual
variation that may be expected within specific limits in
this genus.) Female genitalia figured from specimen in
British Museum. The signum consists of three closely
grouped, moderately large, bluntly rounded disks.
TypE Locauity: Rio Negro, Amazonas, Brazil (May;
type in Paris Mus.).
Foop puant: Unknown.
Known only from the type locality. A distinct
species easily identified by the contrasted black trans-
verse lines on the ash gray forewing.
105. Pseudocabima arizonensis, new species
Friaure 698
Similar in color and markings to the preceeding
species (nigristrigella) but darker, the black dusting
heavier (especially on basal area), giving the wing a
coarser, more pepper-and-salt appearance; no rust-
colored scaling between the veins; the antemedial and
subterminal lines blackish bordered on both sides but
the borders (especially of antemedial line) much less
contrasted than in nigristrigella; subterminal line acute-
ly angled at middle; somewhat expanded black streak-
ing on veins 2, 3, the stalk of 4, 5, and sometimes 6 for
a short distance from cell; terminal dots fused into a
continuous black line along termen. Hind wing shining
white on male, pale smoky white on female; veins very
faintly darkened; a fine dark line along termen. Alar
expanse, 24-27 mm.
Male genitalia exhibiting no specific characters.
Female genitalia with signum a compact cluster of
numerous, closely appressed disks.
Typr Locauity: Redington, Ariz. (type in USNM,
61324; paratypes in Paris Mus., Cornell Univ., Trans-
vaal Mus. (Janse Coll.), and BM).
Foop prant: Unknown.
Described from male type and 2 male and 6 female
paratypes from the type locality; 8 male and 11 female
paratypes from the Baboquivari Mts., Ariz. (June,
July, Aug., Sept.); and 4 male and 1 female paratypes
from Mohave County, Ariz. (Aug., Sept.).
So far, this is the only species of the genus recovered
from the United States. In general habitus it resembles
Euzophera nigricantella Ragonot, also from Arizona.
106. Pseudocabima expunctrix (Dyar and Heinrich), new combina-
tion
Figures 224, 692
Myelois expunctrix Dyar and Heinrich, Proc. Ent. Soc. Washing-
ton, vol. 31, p. 116, 1929.
Forewing slate gray; some black scaling on the veins
and (under magnification) a faint scattering of white
scales over outer area; antemedial and subterminal
lines and discal spots lacking; a row of black dots at
the vein ends along termen. Hind wing semitrans-
lucent white, a smoky shade at apex, along costa, and
on the veins (especially of the female, the veins of the
male wing not appreciably darkened); a fine dark line
along termen. Alar expanse, 22-30 mm.
Female genitalia with signum a long chain of pointed,
thornlike disks.
TypE LocALiTy: Baia, Brazil (type in USNM).
Foop piant: “Stems of leguminous tree.’
Known only from the type series. The only reared
species of the genus and the only one without any
trace of transverse markings. One of the male para-
types proves to be a specimen of Fundella argentina
Dyar. In the forewing veins 4 and 5 are somewhat
longer stalked than in other species of the genus except
perrensiella, the stalking being for a half to slightly
more than half their lengths.
107. Pseudocabima perrensiella (Ragonot), new combination
Fiecure 695
Myelois perrensiella Ragonot, Nouv. Gen., p. 8, 1888; Mono-
graph, pt. 1, p. 52, 1893.
From Ragonot’s description and figure the species
must be similar to erpunctriz except for the presence
of a distinct pale subterminal line, a trace of an ante-
medial line, and a thin dark line along discocellular vein.
Veins 4-5 are ‘long-stalked,” as in exrpunctriz. Alar
expanse, 28 mm.
The female genitalia differ from those of all other
described species of the genus in having a sharp, par-
tially sclerotized, deeply wrinkled, and densely scobi-
nate bend in ductus bursae near its junction with bursa
copulatrix; the signum consists of a curved band of
bluntly rounded, closely impacted, thornlike disks.
Typr Locauity: Goya, Argentina (type in Paris
Mus.).
Foop prant: Unknown.
Known only from the female type.
108. Pseudocabima rubrizonalis (Hampson), new combination
Fieurus 225, 694
Crocidomera rubrizonalis Hampson, Ann. Mag. Nat. Hist., ser.
10, vol. 4, 353, 1929.
Forewing pale gray; the costal edge and the veins
beyond cell purplish brown; the basal area suffused
with some faint dark dusting on the male, considerably
darker on the female; antemedial line at middle of wing,
vertical or nearly so, dull white, bordered on inner and
outer sides by reddish brown bands, the outer one the
wider and somewhat broadened in cell; subterminal
line faint, when distinguishable, grayish white, set well
back from outer margin (the space between it and
antemedial line correspondingly reduced), sharply out-
angled, the apex of the angle in the fork of veins 4-5; a
black line, slightly curved, along discocellular vein;
56 UNITED STATES NATIONAL MUSEUM BULLETIN 207
terminal dots faint, more or less confluent. Hind wing
(of male) translucent white, the veins pale brown in
outer area; a fine brown line along termen. Alar
expanse, 26-30 mm.
Male genitalia with apical process of gnathos a
single, narrow, flattened hook with notched apex.
TypE Locatity: St. Jean Maroni, French Guiana
(type in BM).
Foop prant: Unknown.
DistripuTion: French Gurana: Cayenne, St. Jean
Maroni. Brazi: Parand, Taperinha.
The foregoing description based on three males in
the National Museum. What I take to be a female from
Cayenne in the Janse Collection is also before me. It
differs from the males in haying broader forewing, the
antemedial line outwardly oblique from costa, the
subterminal line more distinct, and the hind wing a
glossy brown. ‘The bursa is large, as long as ductus
bursae; the signum a single straight line of rather sharp,
spinelike thorns.
Genus 27: Hyalospila
[Venational division D. Forewing with 11 veins; 4 and 5 closely
approximate for some distance from cell. Hind wing with all
veins very long; cell less than one-third the length of wing.
Transtilla complete, developed as a narrow, slightly arched band.]
27. Genus Hyalospila Ragonot
Hyalospila Ragonot, Nouv. Gen. p. 11, 1888; Monograph, pt. 1,
p. 168, 1898. (Type of genus: Hyalospila stictoneurella
Ragonot.)
Tongue well developed. Antenna of male simple;
shaft very weakly pubescent. Labial palpus upcurved,
reaching above vertex; slender; segment 3 as long as or a
trifle longer than 2, acuminate. Maxillary palpus sub-
squamous (scaling more or less dilated). Forewing
smooth; 11 veins; vein 2 from before lower outer angle
of cell; 3 from the angle, shortly separated from 4 at
base; 4 and 5 closely approximate for some distance
from cell (not stalked as stated by Ragonot); 6 from
below upper angle of cell, straight; 8 and 9 stalked for
not more (usually less) than half the length of 8; 10
from the cell, approximate to the stalk of 8-9; male
without costal fold. Hind wing with vein 2 from before
lower outer angle of cell; 8 from the angle, closely
approximate to the stalk of 4-5; 4 and 5 stalked for
about half their lengths; 7 and 8 contiguous or very
weakly anastomosed for not over half their lengths
beyond cell; all veins very long; cell less than one-third
the length of wing; discocellular vein curved. Highth
abdominal segment of male with a pair of ventrolateral
hair tufts.
Male genitalia with apical process of gnathos an
elongate, slender hook with slightly notched apex.
Uncus triangulate. Transtilla complete, a simple, nar-
row, slightly arched band. Harpe narrow; costa
strongly sclerotized throughout and projecting slightly
at apex; otherwise simple. Anellus a small shield with
long, slender lateral arms. Aedeagus simple; penis
armed with numerous fine scobinations or one or more
clusters of slender spines. ’
Female genitalia with signa present, consisting of a
single cluster of bluntly pointed thorns, frequently
surrounded by fine scobinations or strongly pigmented
granulations; bursa various, strongly granulate over
mich of one side, weakly sclerotized towards or at,junc-
tion with bursa, more or less finely scobinate or (except
for signum patch)! smooth; ductus bursae weakly
sclerotized towards genital opening or with genital
opening simple; ductus seminalis from bursa near
junction of bursa and ductus bursae.
A distinct, easily recognized genus apparently
limited to tropical America. The species also are easily
identified by their genitalia. It is difficult to, place
Hyalospila satisfactorily in any linear arrangement for
it partakes of the characters of two distinct groups. On
male genitalia, especially its complete bridgelike trans-
tilla, it should go with the genera of the main Acrobasis-
Myelois stem having this organ well developed, while
on other characters, general habitus, and the long veins
and short cell of hind wing it seems more closely related
to genera of the Piesmopoda group. ;
109. Hyalospila stictoneurella Ragonot
Fiaures 44, 226, 703
Hyalospila stictoneurella Ragonot, Nouv. Gen., p. 11, 1888;
Monograph, pt.1, p. 169, 1893.
Forewing purplish brown, a narrow ocherous-white
band along costa, extending from costa to upper vein of
cell and showing under magnification a scattered pow-
dering of reddish scales; the extreme costal edge at its
middle, darkened; the veins more or less streaked with
black scaling; antemedial line indistinct, indicated by
an oblique blackish streak from costa, a small whitish
dot in cell and another on vein 1b, each followed by a
black dot; subterminal line faint but distinguishable,
close to and parallel with termen, not dentate; discal
dots at end of cell confluent, blackish brown. Hind
wing semitransparent whitish; the veins brown, a brown
border along costa and a fine brown line along termen.
Alar expanse, 19-22 mm.
Male genitalia with transtilla a short, slightly arched
bridge, somewhat wider than that of any other known
species and with a minute, pointed projection at middle.
Penis armed with three small clusters of fine, short
spines. Vinculum but slightly longer than greatest
width, tapering to its rather broad, rounded terminal
margin.
Female genitalia with bursa unsclerotized, weakly
scobinate in the area about the signum; the latter a
small patch of small, thornlike spines (fig. 703a) ; ductus
bursae considerably longer than bursa, weakly sclero-
tized for a short distance from genital opening and with
a patch of scobinations at its middle, otherwise mem-
branous and simple.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 57
Type Locatity: Las Mercedes, ‘‘Amer. centr. mer.’
[Guatemala?] (type in Paris Mus.).
Foop prant: Unknown.
Distrisution: México: Jalapa, Misantla (May),
Orizaba. Guatemata: Las Mercedes [?], Volcin Santa
Maria (Aug., Oct., Nov.). Costa Rica: Juan Vifias
(Nov.). Braz: Campo Bello, Santa Catarina (Oct.),
“S. E. Brazil” [Paran4?].
110. Hyalospila celiella Schaus
Fiaures 227, 700
Hyalospila celiella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11,
p. 248, 1913.
Ground color in outer area between the veins and in
entire area between lower margin of cell and inner mar-
gin (except for vein 1b) pale ocherous brown; a narrow
pale band along costa (fainter than that on stictoneurella,
less whitish and, under magnification, showing a fine
irroration of purplish brown scales, especially on costal
edge) ; upper vein of cell white for a short distance from
subbasal area; lower vein of cell white from base to end
of cell except for a black spot near middle; a blackish
brown line bordering the lower edge of cell from base to
middle; a short white streak on vein 1b before middle,
preceded and followed by blackish dots (these and the
black spot on lower vein of cell all that remain to indi-
cate an obsolete antemedial line); vein 1b otherwise
more or less outlined by brown scaling; outer veins
shortly streaked with blackish brown; subterminal line
indicated only by an interruption of the blackish vein
streaks near outer margin; discal spots a pair of short
black streaklets at end of cell. Hind wing semihyaline,
white with a very faint smoky tint in outer area; veins
pale brown; a darker brown line along outer margin.
Alar expanse, 19-21 mm.
Male genitalia easily distinguished by the stout,
greatly elongated vinculum. The abdominal tuft and
sclerotized sternite of eighth segment similar to those
of stictoneurella.
Female genitalia with a weak sclerotization of bursa
in the area surrounding the ductus seminalis and ex-
tending for a short distance into the ductus bursae; no
spining at middle of ductus bursae; otherwise as in
stictoneurella. The sclerotization of the ductus bursae
at genital opening varies individually in extent and
amount as it does in stictoneurella.
TyPE Locatity: Juan Vifias, Costa Rica (type in
USNM).
Foop piant: Unknown.
Represented in the National Collection by the male
holotype (Jan.) and two females (June, Nov.), all from
the type locality.
111. Hyalospila insequens, new species
Similar to celiella except: Larger; ground color of fore-
wing darker, some rosy scaling overlaying most of the
pale area between cell and inner margin; dark streaking
300329—56——_5
on veins fainter; hind wing a clearer white, the veins
not appreciably darkened. Alar expanse, 23-24 mm.
Male genitalia similar to those of celiella except vin-
culum not greatly elongated, slightly less than twice as
long as its greatest width. Female unknown.
Type tocauity: Incachaca, Cochabamba, Bolivia
(type in USNM, 61325; paratype in BM).
Foop puant: Unknown.
Described from male type, collected by J. Steinbach,
and one male paratype from San Antonio, western
Colombia, 5,800 feet, Nov. 1907, M. G. Palmer,
collector.
The species has the same dark streak from base of
forewing along the under edge of the cell as celiella, but
not so strongly accented. The discal spots are slightly
more pronounced and more or less confluent. The
shorter vinculum at once distinguishes it.
112. Hyalospila majorina, new species
Fieure 701
Forewing pale gray-brown; costal area to and includ-
ing the cell dull white, the white shade narrowing
gradually beyond cell to apex of costa; lower discal dot
at end of cell enlarged, dark brown, completely en-
circled by white; antemedial line obsolete; subterminal
line very faint, close to and parallel with costa, indicated
chiefly by short, whitish streaklets on a few of the upper
veins and a slight paling of the ground color from vein
4 to inner margin. Hind wing semitranslucent, whitish
with a faint brown tint; the veins darker; a fine brown
line along termen. Alar expanse, 27 mm,
Female genitalia with ductus bursae considerably
broadened for most of its length, wrinkled and weakly
sclerotized on one side at junction with bursa; bursa
finely scobinate over part of one side, the scobinations
extending into ductus; genital opening simple.
TypE Locatity: Misantla, México (type in USNM,
61326).
Foop prant: Unknown.
Described from female type, collected by Robert
Miiller, Sept. 1914 (“4362”), and one female paratype,
the latter from Jalapa, México. The male is unknown.
I should not have described a new species from females
alone, but in this case a male from México should be
easily matched. The females are readily distinguished
by their large size and their genitalia.
113. Hyalospila fulgidula, new species
Fiaures 228, 702
Ground color of forewing a clear bright white, clouded
by a faint, pale drab shade along inner margin, this
shade beyond lower outer angle of cell extending ob-
liquely upward to apex; costa at base reddish; a minute
red dot on costal edge at one-third and below it a con-
spicuous, broad, oblique, black dash crosses the cell;
below this one or two black dots on vein 1b; a short,
black dash along lower margin of cell at base; some few
scattered black scales on edge of inner margin and on
58 UNITED STATES! NATIONAL MUSEUM BULLETIN 207
some of the veins and a very sparse and scattered dust-
ing of red scales in the outer white area; discal dots at
end of cell separate, black, the lower conspicuous, the
upper minute; subterminal line clearly indicated by a
row of black dots along its inner border and an out-
wardly bordering, short, black dash from costa; a row
of detached black dots from vein 6 to lower fold along
edge of termen. Hind wing glossy, smoky white, dark-
ening outwardly and with a brown shade along outer
margin. Abdominal tufts and eighth segment sternite
of male as in stictoneurella. Alar expanse, 12-13 mm.
Male genitalia with transtilla a very narrow, squarely
arched band; penis very finely scobinate for about the
length of aedeagus, otherwise unarmed. Female geni-
talia with ductus bursae somewhat swollen and densely
but minutely scobinate towards its junction with bursa,
the scobinations extending on one side into bursa;
ductus bursae also sclerotized for a short distance from
genital opening, on its dorsal surface the sclerotization
forming a shield projecting caudally beyond the opening.
TYPE LOCALITY: Santiago Province, Cuba (type in
USNM, 613827).
Foop ruant: Unknown.
Described from’ male type (June) and three female
paratypes (June, Sept., Nov.) from the type locality,
collected by W. Schaus. A distinct species, easily iden-
tified by its small size, the squarely arched transtilla,
and the bright white ground color and contrasted black
spotting of its forewing.
114. Hyalospila egenella (Ragonot), new combination
Fieure 704
Piesmopoda egenella Ragonot, Nouv. Gen., p. 11, 1888; Mono-
graph, pt. 1, p. 165, 1893.
Forewing grayish brown finely powdered with reddish
scales; the costal area white with scattered red scaling;
antemedial line not distinguishable; subterminal line in-
dicated chiefly by its slightly darkened inner and outer
borders, close to and parallel with termen, not sinuate.
Hind wing smoky white, semitranslucent, the veins
darkened and a dark line along termen; the smoky tint
accented somewhat towards apical area. Alar expanse,
15-20 mm.
Female genitalia distinguished chiefly by the en-
larged, wrinkled and scobinate ductus bursae and the
enlarged blunt thorns forming the signum (fig. 704a);
ductus bursae weakly and narrowly sclerotized at gen-
ital opening. The male is unknown.
Typ tocatity: Rio Negro, Brazil (type in Paris
Mus.).
Foop PLiant: Unknown.
A small (15 mm.), somewhat rubbed female in the
U. S. National Museum from Santa Catarina, Brazil
(July), seems to be this species. It has genitalia similar
to those of Ragonot’s type, differing only in minor indi-
vidual details, a somewhat more extended scobination
of ductus bursae, and faint traces of pale yellowish
sclerotization in some of the folds at junction of bursa
and ductus bursae. It also shows traces of a dark discal
spot on forewing at lower outer angle of cell which Rag-
onot states is absent from his type. None of these
differences is significant.
115. Hyalospila xanthoudemia (Dyar), new combination
Figures 229, 709
Piesmopoda xanthoudemia Dyar, Proc. U. 8. Nat. Mus., vol. 47,
p. 333, 1914.
Forewing olivaceous ocherous; the costal area beyond
extreme base and including the cell and the outer area
above vein 6, white with a scattered powdering of red-
dish scales; extreme costal edge red, this shade espe-
cially noticeable at base; antemedial line, discal and
terminal dots obsolete; subterminal line faint but visible,
whitish, very close to termen, not sinuate. Hind wing:
whitish with a faint ocherous fuscous tint, especially in
outer area, somewhat darker on female than on male;
veins faintly darkened; a fine, pale brown line along
termen. Alar expanse, 16-19 mm.
Male genitalia with penis armed with a band (about
one-third as long as aedeagus) of fine scobinations. Fe-
male genitalia with bursa simple except for the signum
patch; ductus bursae much longer than bursa, slender
for most of its length, without scobinations and unsclero=
tized except very weakly at genital opening.
Typr Locaniry: Rio Trinidad, Panamé (type in
USNM).
Foop Pirant: Unknown.
Distrisution: PanamAd: Chiriqui (April), Paraiso
(Jan.), Rio Trinidad (Mar.). Costa Rica: Experenza
(May), Juan Vifias (Oct.).
Doubttully distinct from angulineella (Schaus).
116. Hyalospila angulineella (Schaus), new combmation
Fieure 707
Piesmopoda angulineella Schaus, Ann. Mag. Nat. Hist., ser. 8,
vol. 11, p. 246, 1913.
Known only from the female type. Color and mark-
ings as on zanthoudemia except for faint traces of an
angulate dark antemedial line on forewing, and darker
(pale smoky brown) hind wing.
Ductus bursae of female genitalia longer than that of
zanthoudemia. Otherwise the ductus shows but trifling
differences which are somewhat exaggerated in the figure.
TYPE LOCALITY: Juan Viiias, Costa Rica (June; type
in USNM).
Foop Puant: Unknown.
117. Hyalospila clevelandella (Dyar)
Fiaures 230, 705, 706
Oryctometopia clevelandella Dyar, Proc. U. §. Nat, Mus., vol. 47,
p. 331, 1914. ee
Hyalospila clevelandella (Dyar) Dyar, Ins. Insc. Menstr., vol. 7,
p. 48, 1919. :
Forewing gray-brown from lower margin of cell and
(in outer area) below vein 5, costal area white sparsely.
irrovated with red scales; extreme base of costal edge
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 59
brown; costal edge beyond more or less reddish; ante-
medial line indicated only by fragments of its outer
border, a couple of red dots or dashes in the white area
and a faint, dark, gray-brown dot on lower fold of vein
1b; discal dots at end of cell weak, lower brown, upper
red; subterminal line faint but distinguishable, not sinu-
ate, close to and parallel with termen, dull white. Hind
wing smoky white with the veins and lower fold dis-
tinctly darkened and a dark smoky shade along termen.
Alar expanse, 13-16 mm.
Male genitalia with vinculum sharply constricted into
a digitate projection slightly beyond base; anellus an
elongate, irregularly shaped, curved plate with elongate,
very slender (almost threadlike) lateral lobes; penis
armed with a single, dense cluster of dark brown, slender
spines, the cluster as long or nearly as long as aedeagus.
Female genitalia with ventral surface of half of bursa
and ductus bursae covered with a mat of closely placed,
pigmented granulations, the granulations extending
around partly to dorsal surface; signum patch (on dor-
sum of bursa) surrounded by a teardrop-shaped mass of
granulations (fig. 705a); genital opening simple.
Typr nocauity: Porto Bello, Panamé (type in
USNM).
Foop pLant: Unknown.
Distrisution: PanamA: Porto Bello (Dec.), Taboga
Isl. (Feb.), Tabogilla Isl. (Feb.).
The species can be at once identified by its peculiar
genitalia. The female paratype has genitalia similar in
all but the most trifling details to those of semibrunneella
Ragonot, and if it is actually conspecific with the males
of clevelandella the latter name will fall as a variety or
synonym of semibrunneella. However, there is some
doubt that this is the case; for we have in the National
Collection a series of four females from Cayuga, Guate-
mala, and Jalapa, México, of the same size (16-17 mm.)
and identical color and maculation as the female para-
type of clevelandella, but with quite different genitalia
(fig. 706). Also in the collection are four other females
with the same color and markings and the same size as
the males of clevelandella (13-14 mm.) but with differ-
ent female genitalia. Either of these two groups of
specimens could be the females of Dyar’s species so, for
obvious reasons, I am not attempting to name them or
to propose any synonymy.
118. Hyalospila semibrunneella Ragonot
Ficure 708
Hyalospila semibrunneella Ragonot, Nouv. Gen., p. 12, 1888;
Monograph, pt. 1, p. 169, 1893.
Color and maculation similar to those of clevelandella
except antemedial line more distinct. The female gen-
italia agree substantially with those of the female para-
type of clevelandella. Weshall have to wait discovery of
a male of semibrunneella from the type locality before
the status of the two supposed species can be determined.
Type tocatity: “New Granada” [Colombia] (Mar.;
type in Paris Mus.).
Foop prant: Unknown.
Genus 28: Fundella
[Venational division B. Forewing with veins 4-5 connate or
approximate at base. Hindwing with vein 3 from the stalk of
4-5 or closely approximate to it for some distance; cell short;
on male anal area folded into a pocket. Male genitalia with uncus
hammer-clawed (long, curved, constricted at middle and broadly
divided at apex); transtilla absent; sacculus of harpe not pro-
duced; cornutus present, a single, strong spine. Eighth abdom-
inal segment of male with pair of hair tufts.]
28. Genus Fundella Zeller
Fundella Zeller, Isis von Oken, 1848, p. 866.—Ragonot, Mono-
graph, pt. 1, p. 210, 1893.—Janse, Journ. Ent. Soc. South
Africa, vol. 4, p. 163, 1941.—Heinrich, Proc. U.S. Nat. Mus.,
vol. 96, p. 105, 1945. (Type of genus: Fundeila pellucens
Zeller.)
Ballovia Dyar, Proc. U. S. Nat. Mus., vol. 44, p. 8328, 1913; Ins.
Inse. Menstr., vol. 7, p. 40, 1919. (Type of genus: Ballovia
cistipennis Dyar.)
Tongue well developed. Antenna of male weakly
pubescent, basal segment somewhat enlarged, shaft
laterally flattened and very slightly excavate at base
(fig. 231e) (except on ignobilis and ahemora) and with a
very small blackish scale tuft in the excavation (except
on ignobilis); of female, slender, simple. Front of male
head deeply grooved to hold labial palpi; of female,
rounded. Labial palpus upcurved, reaching to vertex,
clothed with broad appressed scales; on male closely
appressed to face, with second segment over three times
as long as first and with third segment very short (about
one-sixth the length of second); on female with second
segment shorter and third about one-third the length of
second. Maxillary palpus minute, filiform. Forewing
smooth; 11 veins; vein 2 from before lower outer angle
of cell; 3 from the angle, approximately equidistant
from 2 and 4; 4 and 5 connate or approximate at base;
6 from below upper angle of cell, straight; 8 and 9 long
stalked, 9 short; 10 from the cell, parallel for some dis-
tance but not closely approximate to the stalk of 8-9.
Hind wing with vein 2 from close to lower outer angle
of cell; 3 from the stalk of 4—5 or closely approximate to
it for some distance; 4 and 5 stalked for over half (about
two-thirds) their lengths; 7 and 8 closely approximate
beyond cell for less than half their lengths; cell short,
about one-third the length of wing; discocellular vein
curved; on male, anal area (involving veins la and 1b)
thickened and folded under to form a pocket enclosing
enlarged scales and hair tufts. Eighth abdominal seg-
ment of male bearing a thin, short pair of ventrolateral
hair tufts.
Male genitalia with uncus long, curved, strongly scler-
otized, constricted at middle and broadly divided at
apex (hammer-clawed) ; gnathos terminating in a short,
stout hook or a short, broad plate (achemora); transtilla
absent. Harpe rather short, with clasper. Vinculum
narrow, short. Aedeagus stout with long, stout, pro-
jecting, curved spine or spines at apex (except in
argentina); cornutus a single, strong spine.
Female genitalia without signum (pellucens) or with
signum well developed and consisting of a large oval or
pear-shaped cluster of thornlike spines (argentina, aga-
60 UNITED STATES NATIONAL MUSEUM BULLETIN 207
pella), or curved sclerotized bands armed with stout,
thornlike spines (ahemora, ignobilis) ; bursa large; ductus
bursae short, broad (narrowest in agapella); area sur-
rounding genital opening strongly sclerotized, the dorsal
sclerotization in the form of a band connected with the
supporting rods of eighth segment collar, and armed
with two or four spinelike projections (except in ignobilis
and some examples of argentina); ductus seminalis from
caudal area of bursa.
This genus is easily distinguished by its striking male
characters; the strongly sclerotized, long-stemmed, bi-
furcate (hammer-clawed) uncus; the large pocket on
anal area of hind wing; the long, embedded labial palpus
with very short third segment; and minute maxillary
palpus. A similar bifurcate uncus is not found in any
other American genus except Difundella Dyar. In the
type species of the latter (corynophora Dyar) the uncus
is somewhat produced and exhibits a slight bifurcation
at apex; but other species, which must also be referred
to Difundella, lack this character. Difundella separates
readily on other male structures—its strongly hooked,
partially free sacculus of harpe, its rounded frons, and
the narrow, strongly sclerotized, deeply invaginated
pocket of the sternite of the eighth abdominal segment.
In Fundella the wing pattern varies so much within
any given species that it affords no reliable character
for specific identification, and the several species can be
separated with certainty only by their genitalia.
119. Fundella pellucens Zeller
Figures 6, 231, 713
Fundella pellucens Zeller, Isis von Oken, vol. 41, p. 866, 1848;
Horae Soc. Ent. Rossicae, vol. 15, p. 236, 1881.—Ragonot,
Monograph, pt. 1, p. 210, 1893.—Heinrich, Proc. U. 8. Nat.
Mus., vol. 96, p. 107, 1945.
Ballovia cistipennis Dyar, Proc. U. 8. Nat. Mus., vol. 44, p. 323,
1913.
Fundella cistipennis (Dyar) Dyar, Ins. Insc. Menstr., vol. 7, p. 40,
1919.—Wolcott, Journ. Agr. Uniy. Puerto Rico, vol. 17,
pp. 241-255, 1933; Journ. Agr. Univ. Puerto Rico, vol. 18,
p. 432, 1934; vol. 20, p. 477, 1936.—Scott, Journ. Agr. Univ.
Puerto Rico, vol. 24, pp. 35-47, 1940.
Male antennal shaft with very small black basal tuft
(fig. 231e). Forewing grayish fuscous more or less dusted
with whitish and with interspersed reddish brown scales
(in ‘many specimens the ground color is reddish brown),
giving the moth a distinctly gray or gray-brown appear-
ance to the naked eye; a conspicuous, round, darker
brown or fuscous spot in the center of the area usually
occupied by the antemedian line, this dark spot more or
less obscured in some specimens, but in typical examples
outlined by whitish areas inwardly and outwardly and
not reaching to inner margin or costa of the wing; discal
mark at end of cell obscure, often absent; subterminal
line (when distinguishable) faint, white, indented at
vein 6 and at submedian fold; a row of dark spots along
termen (present only in specimens having an appreciable
dusting of white scales). Hind wing white, translucent,
a faint fuscous border along costa and (in some speci-
mens) a fuscous line on termen for a short distance from
apex; cilia white; anal pocket yellowish white. Mid-
tibia with a fringe of pale hairlike scales along dorsum.
Hind tibia with a rather long and slender tuft of pale
(whitish ocherous), hairlike scales from the knee joint
(fig. 231f).
Female essentially like the male in color and markings
except that the dark spot near the base of the forewing
is more diffused, sometimes reaching to the costa. Hind
wing usually with a dark shade along termen.
Alar expanse, 19-24 mm.
Male genitalia with a large, strongly sclerotized
subanal plate, constricted before and beyond its middle.
Harpe with apex notched below costa; clasper short,
curved, situated near middle of harpe, and armed with
several setae at its knobbed apex. Aedeagus with a
cluster of several long, curved spines from apex; cor-
nutus long, straight, stout.
Female genitalia with bursa copulatrix finely scobi-
nate but without signum; ductus bursae flattened,
broad, twisted and constricted near genital opening,
sclerotized throughout, the sclerotization involving
bursa adjacent to ductus bursae and ductus seminalis;
sclerotized band behind genital opening armed with
four long, stout, projecting spines; collar of eighth seg-
ment invaginated at dorsal margin to form a sclerotized
pocket (fig. 713a).
TYPE Locaitius: St. Thomas, British West Indies
(pellucens, in BM); Barbados (cistupennis, in USNM).
Foop piants: Vigna unguiculata (cowpeas, black-
eyed peas, and garden peas), Bauhinia variagata, Can-
avalia ensiformis (swordbeans), Canavalia maritima
(black bean), Cajan cajan (pigeon pea), Phaseolus luna-
tus (cultivated and wild limabeans), Phaseolus sp.
(Brazilian specimens), Cassia occidentalia (one reared
specimen from McCubbins Mills, Puerto Rico, before
me; most records from this last plant are doubtful and
probably the result of a misidentification of Fundella
argentina as cistipennis).
According to Scott the favored host in Puerto Rico is
the cowpea (Vigna unguiculata), and the species, while
frequent in limabeans, seldom does serious damage.
Potentially it is an insect of economic importance. The
larvae are primarily pod borers, but also bore into the
stems and feed on the flowers of their hosts. They
attack, as far as known, only Leguminosae.
DistrisuTion: Unirep States: Florida, Hobe Sound
(May), Miami, (Apr., May), Jupiter (Apr.), Coconut
Grove, Marco Isl., Tampa (Mar.); Walton, Jensen
(U. S. Dep. Agr. rearings from limabeans, Feb.
1944), Riviera Beach, Vero Beach (J. R. Malloch, Dec.
1941), Barsapos. Hartz: Damien (Dec., Feb.),
Port-au-Prince. MontTSERRAT (Jan.). CuBA: Santiago,
Matanzas. Vircin Isuanps: St. Croix (Mar., Oct.,
Nov.). Purrto Rico: San Juan, Rio Piedras (Mar.—
May), Isabella, Catano (July), Vieques Isl. (Apr.).
Braziu: Bafa (May), Cearé. Botrvra.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 61
120. Fundella argentina Dyar
Figures 234, 711
Fundella pellucens Zeller (in part, ‘‘var. b’’), Isis von Oken, vol.
41, p. 867, 1848; Horae Soc. Ent. Rossicae, vol. 16, p. 237,
fig. 41b, 1881.
Fundella argentina Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 1919.—
Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. 109, 1945.
Fundella eucasis Dyar, Ins. Insc. Menstr., vol. 7, p. 40, 1919.
Male antennal shaft with even smaller black basal
scale tuft than that of pellucens. Forewing gray with-
out the reddish brown, interspersed scaling character-
istic of typical examples of pellucens; entire basal area
to antemedian line dark fuscous gray (with but very
slight dusting of whitish scales toward base in some
specimens); this dark basal patch contrasted against
the paler gray color of the remainder of the wing, ex-
tending from costa to inner margin and bordered out-
wardly by a narrow whitish line. Otherwise not dis-
tinguishable, superficially, from pellucens.
Female essentially like the male in color and markings
except that the basal area of forewing is concolorous
with or contrastingly paler than the remainder of the
wing. A narrow dark line or a diffused dark shading
outwardly bordering the obscure antemedian line.
Alar expanse, 15-23 mm.
Male genitalia without sclerotized subanal plate.
Terminal projection of gnathos varying from round to
pointed (fig. 234b) at apex. Harpe tapering to bluntly
pointed apex; clasper a single, straight, slightly rough-
ened, appressed spine, situated beyond middle of harpe.
Aedeagus simple; cornutus a single, straight spine.
Female genitalia with signum well developed and
consisting of a large pear-shaped cluster of thornlike
spines; sclerotized band behind genital opening, divided
in the middle, simple (fig. 711a) in Argentinian and
Brazilian specimens, or armed with a pair of median,
spinelike projections (fig. 711), rather long in West In-
dian specimens or short and disappearing in Mexican
and Venezuelan specimens.
TYPE Locaitigs: Tucumén, Argentina (argentina, in
USNM); Caracas, Venezuela (eucasis, in USNM).
Foop puant: Cassia spp. (reared examples in Na-
tional Collection from Cassia bicapsularis and C. corym-
bosa), Poinciana gilliesi.
Distrisution: Unirep Srares: Florida, Biscayne
Bay (May), Coconut Grove (Apr.), Stock Island (Apr.);
Texas, Brownsville (Nov.). Mxico: Several examples
reared from pods and blossoms of Cassia bicapsularis
at Brownsville, Tex., quarantine station. CuBa: Bara-
gua (Mar.), Habana, Matanzas, Santiago Province.
Puerto Rico: Bayamén (Mar., Sept.), Vieques Isl.
(Apr., July), Coamo Springs (Apr.), Aguirre Central
(Aug.), San German (Aug.), San Juan (Noy.). Hartt:
Pétionville (June). Virein Isuanps: St. Croix (Oct.-
Nov.). Jamatca. VENEZUELA: El Valle (June).
Braziu: Bafa (May). Argentina: Tucumén (Mar.).
In collections this species has appeared most fre-
quently under the name pellucens. Both argentina and
pellucens have about the same distribution and are
abundant in the West Indies, though, from material
at hand, pellucens seems to be rarer on the mainland.
Throughout its range argentina shows considerable var-
iation in female genitalia. West Indian specimens have
rather conspicuous spinelike extensions of the sclero-
tized band behind the genital opening. These are en-
tirely lacking in Brazilian specimens, and if one had
only these extremes he would be justified in assuming
that they were at least racially distinct. However,
Venezuelan and Mexican examples show an intermedi-
ate form with very short projections, and Central Amer-
ican specimens, when recovered in sufficient numbers,
will probably show all intergradations. The male geni-
talia are remarkably uniform throughout the range of
the species, exhibiting only minor individual variations
in the shape of the terminal projection of the gnathos.
121. Fundella agapella Schaus
Figure 710
Fundella agapella Schaus, Zoologica, vol. 5, No. 2, p. 47, 1923.—
Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. 111, 1945.
Female palpi, head, thorax, and forewing whitish
gray; dark markings drab gray; transverse antemedian
line of forewing white, defined chiefly by its narrow,
dark outer border, sharply sinuate, indented a trifle
just below costa, more deeply at top of cell and still
more deeply at fold below cell; discal dot at end of cell
obscure; white subterminal line indented at vein 6 and
at submedian fold, bordered inwardly by a distinct dark
shade .as broad as the white line itself and outwardly
by a similar, fainter, dark shading, the latter conspic-
uous only at apex. Hind wing as in the other species
of Fundelia. Alar expanse, 12 mm;
Genitalia like those of intermediate examples of ar-
gentina except that the signum is considerably smaller
in proportion to the size of the bursa.
Typr Locarity: Tagus Cove, Albemarle, Galipagos
Islands (type in USNM).
Foop rrant: Unknown.
Known only from the female type. Superficially a
distinct species. The female genitalia, however, would
indicate that agapella is only a race of argentina. A
male will be needed for exact placement, and until it is
available we shall have to treat agapella as a species.
122. Fundella ignobilis Heinrich
Ficures 232, 712
Fundella ignobilis Heinrich, Proc. U. 8. Nat. Mus., vol. 96, p.
112, 1945.
Male antennal shaft without any trace of black basal
scale tuft. Otherwise partaking of the pattern mark-
ings of both pellucens and argentina; in some specimens
dark basal patch of forewing round and reaching neither
costa, nor inner margin (as in typical pellucens), in ma-
jority of specimens, however, basal patch occupying
whole basal area (as in typical argentina); median and
outer areas of wing averaging a trifle paler than in
argentina and without the reddish brown scaling of
pellucens.
62 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Female superficially similar to argentina except a
trifle paler, on the average.
Alar expanse, 13-22 mm,
Male genitalia with gnathos terminating in a short,
stout hook. Harpe with apex truncate; clasper mod-
erately long, curved, and weakly haired at apex. Ae-
deagus with a single, long, strong, curved spine from
below apex; cornutus a short, stout, curved thorn.
Female genitalia without spines adjacent to genital
opening. Bursa copulatrix with signa consisting of a
pair of partially fused bands, each armed with a row
of short, stout, thornlike spines; ductus bursae short
and broad, with median area unsclerotized; eighth seg-
ment collar completely sclerotized except for a small,
round, transparent spot on midventer, sclerotization
extending to and over area behind genital opening.
TypE LOcALity: Oaxaca, México (type in USNM).
Foop prant: Unknown.
Distrisution: México: Cérdoba (May), Guadala-
jara, Jalapa, Oaxaca, Orizaba, Tehuacén (May, June,
July). Guatemata: Cayuga. Costa Rica. Cusa:
Santiago (June), Sierra Miestra (May). Purrto Rico:
Aguirre Central (Aug.). Harri: Pétionville (June).
123. Fundella ahemora Dyar
Figures 233, 714
Fundella ahemora Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 403,
1914.—Heinrich, Proc. U. S. Nat. Mus., vol. 96, p. 113, 1945.
Antenna of male with small black scale tuft at base
of shaft. Forewing with no or a very faint dark basal
patch (when present covering basal area to antemedian
line); antemedian line whitish, very faint; subterminal
line white, faint but less obscure than antemedian,
without dark borders except for an inner and an outer
dark spot at inner margin of wing; veins from cell rather
strongly outlined by dark scaling (the most conspicuous
superficial character of the species). A thick, dark
(brownish) hair tuft covering outer surface of male
foretibia, a character not found in other species of the
genus. Alar expanse, 18-23 mm.
Male genitalia with gnathos terminating in a broad
tongue-like plate. Harpe somewhat tapering but with
apex truncate; a strong tuft of long scales from costa;
clasper long, curved, slender, with a few hairs at apex.
Aedeagus with a pair of long, curved, flattened spines
from apex; cornutus a long, straight, slender spine.
Female genitalia with a pair of long, widely spaced,
basally curved spines from sclerotized area immediately
behind genital opening. Bursa copulatrix with signa
consisting of two rather short bands, each armed with
a row of long spines. Ductus bursae bulged in the
middle and with a strongly sclerotized median collar.
Collar of eighth segment partially sclerotized and fused
ventrally.
TYPE LocaLity.—Orizaba, México (type in USNM).
Foop pLant.— Unknown.
DistRIBUTION: México: Orizaba, Jalapa, Teapa
(Dec.), Cérdoba (Apr., Dec.), Cuernavaca (July).
GUATEMALA: Quirigué (Mar.), Cayuga (Jan., May),
Parulhé (July). Costa Rica: Juan Vifas (Nov.).
Superficially the most easily distinguished species in
the genus. The large foretibial tuft at once identifies
the male, and both sexes can be separated by the rather
conspicuous dark outlining of the veins. The veins are
similarly dark scaled in the other species, but the con-
trast of the dark veins against the pale intervenular area
is more marked in ahemora.
Genus 29: Difundella
[Venational division B. Forewing with veins 4-5 connate or
closely approximate at base. Hind wing with vein 3 from stalk
of 4-5 or closely approximate to it for a short distance; on male,
anal area folded into a pocket. Male genitalia without transtilla;
sacculus of harpe strongly sclerotized and produced (free or
partially free); penis without cornutus or other armature.
Abdomen of male with lateral pockets and hair tufts between
segments 2 and 3; eighth abdominal sternite developed as a
narrow pocket; no hair tufts.]
This genus shows affinities to Fundella but in many
characters resembles more closely Rampylla and Cop-
tarthria. In general habitus (wing pattern, color, and
maculation) the moths of Difundella, Coptarthria, and
Rampylla are strikingly similar, but the three genera
are different on structural characters. Rampylla differs
from the other two in the free length of vein 3 of
male hind wing; Coptarthria in its notched male antenna;
and Difundelia in its anellus (a simple plate without the
long free spine of Rampylla and Coptarthria) and in the
possession of scaled pockets between the second and
third segments of the male abdomen.
The species of Difundella differ considerably from
each other on structural details, falling into two distinct
groups which divide as follows:
—Labial palpus reaching above vertex in both sexes. Hind
wing with cell less than one-third the length of wing;
vein 1b of male bent before middle and with a tuft of
yellow hairlike scales on its under side (within the bend).
Gnathos greatly reduced, its apical projection fine,
needlelike. Costa of harpe without projections.
—Labial palpus not reaching vertex on males, barely reaching
vertex on females. Hind wing with cell more than
one-third (but less than half) the length of wing; vein
1b of male not bent; rough sex-scaling bordering 1c on
under side of wing beyond base. Gnathos with apical
process enlarged and strongly sclerotized. Costa of
harpe with strongly sclerotized projection or projections.
The second group probably deserves a separate generic
designation; but the material before me representing its
two species is too scanty and not in good enough con-
dition, and the association of the females with their
proper males too uncertain, to permit proper evalua-
tion of generic characters for separation at this time.
29. Genus Difundella Dyar
Difundella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 327, 1914,
(Type of genus: Difundella corynophora Dyar.)
Tongue well developed. Antenna of male weakly
pubescent. Labial palpus ascending, recurved, slender,
smooth scaled; third segment acuminate. Maxillary
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 63
palpus with second segment slightly thickened with
scales. Forewing smooth; 11 veins; vein 2 from before
lower outer angle of cell; 3 from the angle, nearly
equidistant from 2. and 4; 4 and 5 connate or closely
approximate at base and approximate for a short dis-
tance beyond base; 6 from below upper angle of cell,
slightly bent towards base; 10 from the cell, more or less
approximate to the stalk of 8-9. Hind wing with vein
2 from close to lower outer angle of cell; 3 from the
stalk of 4-5 or closely approximate with it for a short
distance from angle of cell; 4 and 5 long stalked (for over
one-half their lengths); 7 and 8 closely approximate
beyond cell, or shortly and weakly anastomosed; cell
short, less than one-half the wing length; discocellular
vein curved, partially obsolescent; on male, anal angle
folded under to form a pocket enclosing a long hair-
pencil. Eighth abdominal segment of male with sternite
developed as a narrow, sclerotized pocket; on each side
between abdominal segments 2 and 3 a shallow pocket
containing a modified scale tuft.
Male genitalia with uncus stout, but variously shaped.
Transtilla absent. Harpe with sacculus very strongly
sclerotized, free or partially free and curved. Aedeagus
with strongly sclerotized and pointed apex; penis
unarmed. Vinculum stout. A long hair tuft from
intersegmental area adjacent to base of sacculus of
harpe.
Female genitalia with bursa copulatrix membranous;
signum, if present, a small patch of weak scobinations;
ductus bursae membranous (unsclerotized) except about
genital opening; ductus seminalis from bursa near
junction of bursa and ductus bursae.
The characteristic pattern features of the species of
Difundella (as of Coptarthria and most species of
Rampylia) are: The strongly contrasted, fine, blackish,
transverse lines forming the outer border of the ante-
medial and the inner border of subterminal lines of
forewing; the almost straight, oblique or vertical ante-
medial line, set well out from base of wing; the oval,
pale discal spot covering the discocellular vein; and the
black streaks on veins 2 to 6 just beyond it.
Genus Difundella, Species 124 and 125: D. cory-
nophora and D. subsutella
[Labial palpus reaching above vertex in both sexes. Hind wing
with cell less than one-third the length of wing; vein 1b of male
bent before middle and with a tuft of yellow hairlike scales on its
under side (within the bend). Gnathose greatly reduced, its
apical projection fine, needlelike. Costa of harpe without
projections.]
124. Difundella corynophora Dyar
Fiaures 7, 235, 715
Difundella corynophora Dyar, Proc. U. 8. Nat. Mus., vol. 47’
p. 327, 1914.
Forewing with basal area (to antemedial line), and
upper median area beyond antemedial line and from
lower margin of cell to costa, blackish fuscous; some
extension of this dark shade extends narrowly to inner
margin along the outer dark border of the antemedial
line; ground color of remainder of wing a ruddy ocherous;
some extension of this ocherous shade invades the dark
basal area along the lower fold and forms the centers of
the transverse lines; beyond the cell the ocherous shade
is broken by black streaks on veins 2 to 6 and beyond
subterminal line it is more or less clouded by blackish
fuscous; discal spot ruddy ocherous, covering disco-
cellular vein; antemedial line oblique; its narrow dark
borders black shading to reddish; the dark borders of
subterminal line also more reddish brown than black;
all the dark transverse lines less contrasted and con-
spicuous than those of other species in the genus. Hind
wing dark smoky fuscous; the veins and terminal edge
black and (on the female) a blackish shade at apex.
Alar expanse, 15-16 mm.
Male genitalia with uncus produced and slightly
bifurcate at apex. Free projection of sacculus curved
back towards lower margin of harpe. Aedeagus forked;
the longer element of the fork spined at apex. Female
genitalia with signum a small round patch; genital
opening surrounded by an oblong, strongly sclerotized
plate; in intersegmental area behind this plate a pair of
ventrolateral pockets (fig. 715b).
Typr tocauiry: La Chorrera, Panama (type in
USNM).
Foop piant: Unknown.
DistrisuTion: GuATEMALA: Cayuga (Aug.), Chejel
(June). Panam&: La Chorrera. Franco GUurIANA:
Cayenne.
Easily identified by its genitalia.
125. Difundella subsutella (Schaus), new combination
FicursE 236
Ulophora subsutella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11,
p. 248, 1913.
Rampylla subsutella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7,
p. 84, 1919.
The only representative of this species is the male
type which is somewhat rubbed and the markings con-
sequently obscured. It differs from corynophora in
having the pale ground color of the outer areas of fore-
wing more reddish than ocherous; the black outer
margin of antemedial line more sharply defined, black
throughout, outwardly oblique to lower fold and thence
slightly curved inward to lower margin. Hind wing
semihyaline white with a faint brownish ocherous tint;
veins not appreciably darkened; a fine brown line along
termen. Alar expanse, 17.5 mm.
Male genitalia with uncus triangulate. Gnathos
reduced even more than that of corynophora, the lateral
arms represented by mere stubs. Projecting part of
sacculus curving away from harpe. Aedeagus sickle
shaped; its apical half sharply curved, very strongly
sclerotized, and tapering to a sharp point.
Typr LOCALITY: Juan Vifias, Costa Rica (Jan.; type
in USNM).
Foop puant: Unknown.
64 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Genus Difundella, Species 126 and 127: D. dis-
tractor and D. tolerata
[Labial palpus not reaching vertex on males, barely reaching
vertex on females. Hind wing with cell more than one-third
(but less than half) the length of wing; vein 1b of male not bent;
rough sex-scaling bordering Ic on underside of wing beyond base.
Gnathos with apical process enlarged and strongly sclerotized.
Costa of harpe with strongly sclerotized projection or projections.]
126. Difundella distractor, new species
Figures 237, 716
Dark areas of forewing dark brownish gray; the pale
outer areas dull whitish; antemedial line well out
towards middle of wing, nearly vertical, slightly notched
at vein 1b, red-brown with a fine black line along its
outer edge and preceded by a rather broad whitish
blotch, extending from just below costa, nearly to inner
margin; a short black streak along lower fold for a short
distance from base of wing; the discal spot whitish;
blackish lining on the veins beyond cell very weak;
subterminal line very slightly bent between veins 4
and 6 otherwise nearly vertical, outwardly bordered by
a narrow shade of the dark ground color and inwardly
by a fine black line, Hind wing semihyaline white; a
blackish brown line along termen; the veins not dark-
ened; the male wing on the undersurface rather loosely
and coarsely covered with yellowish scales, especially
along the veins. Alar expanse, 14.5-15.5 mm.
Male genitalia with uncus semispoon-shaped. Apical
projection of gnathos large, triangulate, strongly sclero-
tized. Harpe with subtriangulate cucullus; a single,
long, stout, flat, curved, tapering projection from mid-
costa. Aedeagus long, slender, smooth, tapering to a
point. Vinculum’ about twice as long as greatest
width. Female genitalia without signum; genital
plate large, medially notched and supported by strongly
sclerotized, involuted seventh abdominal segment.
TyprE Locatity: Palmas Abajas, Puerto Rico (type
in Cornell Univ., paratype in USNM, 61328).
Foop PLant: Unknown.
Described from male type labeled ‘‘Palmas Abajas
(near Guayama), P. R., 1900 ft., [date unreadable],
W. A. Hoffman’; and one female paratype from Aguirre
Central, Puerto Rico, ‘‘Apr. 2-3, 31,’’ M. D. Leonard,
collector. The female is in good condition and served
for the foregoing color and pattern description. The
male is badly rubbed and has the palpi and antennae
broken off; but enough of the markings remain to show
that they were the same as those of the female.
127. Difundella tolerata, new species
Fieures 238, 717
Similar to the foregoing species (distractor) except
that pale areas of forewing’ are much more restricted,
limited to an irregular area bordering inner margin of
subterminal line ‘(extending back to cell above and
nearly to antemedial line below); some diffused pale
shading along the lower fold in basal area; the pale
areas very dull ocherous white, not as well contrasted
as in distractor; discal spot ocherous. Hind wing trans-
lucent, white with a very faint ocherous tint on male;
a faint dark line along termen. Alar expanse, 19 mm.
Male genitalia with uncus strongly sclerotized and
stout (longer than basal width), broadened and rounded
towards apex. Apical projection of gnathos a long,
broad, strongly sclerotized, obliquely bent band.
Harpe with narrow, spatulate cucullus; two projections
from costa, the first a double thornlike projection from
near middle, the second a rather slender spine from
outer third. Aedeagus with a couple of short spines
at apex. Vinculum no longer than greatest width.
Female genitalia with signum a narrow, elongate
patch of scobinations; genital opening simple; ductus
bursae about three times as long as bursa. From
the dorsocaudal margin of seventh segment an in-
vaginated, sclerotized shield supports at each lateral
margin a short, blunt, weakly pigmented hornlike
process.
Typr Locatity: Kast Bolivia (type in BM).
Foop puant: Unknown.
Described from two specimens from the British Mu-
seum Collection labeled “Ost Bolivia, Aug.—Oct., 1920,
T. Steinbach,’’ the male type and a female matching the
male in size, color, and markings. J do not make’ this
female a paratype (although the foregoing description
of the female genitalia is made from it) because there is
a slight doubt that it is the true female of the species.
Its genitalia are similar in all but the most trifling details
to those of a series of females from Cayuga, Guatemala,
associated in our collection with males of Coptarthria
dasypyga. One or the other of our identifications of
females (or both of them) may be in error. We shall
not know until more material is available.
Genera 30-33: Coptarthria to Dasypyga
[Venational division B. Male genitalia with transtilla a sinuate,
sclerotized, more or less scobinate band involved with gnathos;
a long, free spine associated with anellus. Highth abdominal
segment of male with sternite developed as a narrow sclerotized
pocket.] 4
30. Genus Coptarthria Ragonot
Coptarthria Ragonot, Monograph, pt. 1, p. 251, 18938. (Type of
genus: Myelois dasypyga Zeller.)
Tongue well developed. Antenna‘of male with basal
segment of shaft considerably elongated, flattened and
deeply notched at outer extremity; shaft pubescent.
Labial palpus upturned, slender, barely reaching vertex;
third segment about half the length of second, bluntly
pointed. Maxillary palpus filiform. Forewing smooth;
11 veins; vein 2 from before lower outer angle of cell; 3
from the angle, closer to 4 at base than to 2; 4 and 5
short stalked or connate and closely approximate for a
short distance from base; 6 from below upper angle of
cell, bent towards base; 10 from the cell; male without
costal fold. Hind wing with vein 2 from close to lower
outer angle of cell; 3 from middle of stalk of 4-5, or
closely approximate to it; 4. and 5 stalked for slightly
more than half their lengths; 7 and 8 closely approxi-
mate beyond cell; cell short, about one-third the length
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 65
of wing; discocellular vein curved. Highth abdominal
segment of male with sternite developed as a narrow,
sclerotized pocket.
Male genitalia with uncus moderately stout; deeply
concaved apically (probably only a specific character).
Transtilla a sinuate, sclerotized band involved with and
fusing into gnathos. Gnathos proper identifiable only
by its rather weak lateral arms. Harpe with apex of
cucullus slightly hooked; sacculus simple, not produced.
Anellus a small plate with greatly reduced lateral lobes,
the latter indicated chiefly by their short hair tufts;
dependent from near base of anellus plate and associated
with it a long, slender U-shaped band supporting from
the bottom of the U a long, strongly sclerotized, free
spine, the latter lying dorsad of the aedeagus. Aedeagus
small, simple; penis with a few weak scobinations, other-
wise unarmed.
The genus is distinguished from its nearest allies by
the notched shaft of its male antenna. Female char-
acters could not be included in the foregoing descrip-
tion, as examples of this sex have not been satisfactorily
associated with males of the type species, the only
known representative of the genus.
128. Coptarthria dasypyga (Zeller)
Ficures 10, 239, (?) 718
Myelois dasypyga Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 215,
1881
Coptarthria dasypyga (Zeller) Ragonot, Monograph, pt. 1, p. 251,
1893.
I have seen no Colombian examples, but there are six
males in the National Museum from Guatemala which
Dyar identified as dasypyga. There is no reason to
question his identification; for the specimens have the
typical Coptarthria antenna, are the right size, and agree
in color and maculation with Ragonot’s description of
the Zeller type.
Forewing gray-brown with a very slight intermixture
of ocherous shading above inner margin; the basal area
a trifle paler than remainder of wing; branches of median
vein (veins 2 to 5) faintly streaked with brown or black-
ish brown; transverse lines pale with dark borders and
faintly tinged with reddish scaling at middle and near
costa; antemedial line well out near middle of wing,
nearly vertical, straight except for a slight inward angu-
lation at vein 1b, its inner border a weak brown line, its
outer bordering line black; subterminal line well back
from termen, vertical to vein 4, thence oblique to inner
margin, its inner border a black line; a rather large, oval,
ochraceous spot on discocellular vein, margined by faint
black scaling; along termen a row of conspicuous, more
or less confluent black spots. Hindwing semitranslu-
cent white; the veins faintly darkened (pale, ocherous
brown) and a faint dark line along termen. Alar ex-
panse, 14-17 mm.
Male genitalia as given for the genus.
TYPE Locatity: Honda, Colombia (type, o, in BM).
Foop piant: Unknown.
300329—56——6
Distrisution: Coomera: Honda, GuaTeMata: Cay-
= om , May), Quirigué (Mar.), Volcén Santa Maria
y).
Associated with the males in the National Collection
are five females from Cayuga (Apr., May) identical with
the males in all superficial characters. Their genitalia
are like those of the Bolivian female I have associated
tentatively with the type of Difundella tolerata (fig. 717).
However, there is also a female with the same color and
markings from Juan Vifas, Costa Rica (Feb.), which
has different genitalia (fig. 718), similar to those of
Anadelosemia. From the limited material available
and the few and scattered distributional records it is im-
possible to determine which females go with which males,
31. Genus Promylea Ragonot
Promylea Ragonot, N. Amer. Phycitidae, p. 5, 1887; Monograph,
pt. 1, p. 207, 1893.—Hulst, Phycitidae of N. Amer., p. 139,
1890. (Type of genus: Promylea lunigerella Ragonot).
Tongue well developed. Antenna pubescent; shaft
of male without notch or other modifications. Labial
palpus upturned, slender, reaching vertex; third seg-
ment about half the length of second, acuminate. Max-
illary palpus squamous. Forewing smooth; 11 veins;
vein 2 from before, but rather near lower outer angle of
cell; 3 from the angle, closer to 4 than to 2 at base; 4
and 5 closely approximate for a short distance from
base; 6 from below upper angle of cell, straight; 8 and 9
long stalked, the free element of 9 short; 10 from the
stalk of 8-9, or from the cell, connate with or closely
approximate with it for a short distance beyond base
(definitely stalked with 8-9 in most of the specimens of
lunigerella); male without costal fold. Hind wing with
vein 2 from before lower outer angle of cell; 3 from the
angle, connate with 4; 4 and 5 closely approximate or
anastomosed for half their lengths beyond cell; 7 and 8
approximate or partially anastomosed for less than half
their lengths beyond cell; cell nearly half the length of
wing; discocellular vein curved. Eighth abdominal seg-
ment of male with sternite developed as a narrow sclero-
tized pocket.
Male genitalia asin Coptarthria except: Apical margin
of uncus rounded; lateral arms of gnathos more strongly
developed; penis sometimes with a weak cornutus.
Female genitalia with bursa and ductus bursae mem-
branous, bursa small, considerably shorter than ductus;
signum present but weak, a patch of scobinations or a
small plate supporting a very small thorn; genital open-
ing simple; ductus seminalis from ductus bursae. Col-
lar of eighth abdominal segment with a broad, flaring,
sclerotized apron projecting from center of anterior
dorsal margin; in the intersegmental area between collar
and seventh segment a sclerotized and coarsely granu-
late pocket (fig. 721a).
The genus is close to both Coptarthria and Anadelo-
semia but distinct, differing from the former in its simple
male antenna and from the latter and all the genera of
this immediate group having the sinuate, involved
transtilla and the free spine associated with anellus by
its peculiarly developed, female, eighth-segment collar.
66 UNITED STATES NATIONAL MUSEUM BULLETIN 207
In habitus the moths of Promylea differ strikingly from
those of Coptarthria; the transverse lines of forewing
being more widely separated and the antemedial line
decidedly oblique.
129. Promylea lunigerella Ragonot
Figures 9, 240, 721
Promylea lunigerella Ragonot, N. Amer. Phycitidae, p. 5, 1887;
Monograph, pt. 1, p. 20, 1893.—Hulst, Phycitidae of N.
Amer., p. 139, 1890.—McDunnough, Check list, No. 6146,
1939.
Forewing gray or brownish gray, pale in median and
most of basal areas and (in many specimens from Van-
couver and Washington State) with a faint rosy tint;
antemedial line indicated chiefly by its blackish outer
border, strongest towards costa and oblique from costa
before one-third to inner margin, preceded by reddish
brown or ocherous brown patch, broad and inwardly
dark-margined on inner margin, attenuated and paling
out towards costa; subterminal line pale gray, bordered
inwardly by a blackish brown line and outwardly by a
much fainter dark line (grayish or reddish brown), out-
wardly curved between veins 8 and 1b; discal dots rarely
separated, normally fused into a thin blackish lunule
on the discocellular vein; terminal dots obscure, when
distinguishable more or less fused into a line along ter-
minal margin. Hind wings pale smoky fuscous; the veins
little if any darkened; a very faint brownish line along
termen. Alar expanse, 20-24 mm.
TYPE Locatiry: Vancouver Isl., British Columbia
(type in Paris Mus.).
Foop puant: Unknown. :
Distripution: Canapa; British Columbia, Fitzgerald
(June), Duncans (Vancouver Isl.), Victoria (June, July,
Aug.), Unirep Statss: Washington, Bellingham (Aug.),
Friday Harbor (July, Aug.), Mt. Constitution (July);
California, Glen Alpine (Lake Tahoe, July).
130. Promylea lunigerella glendella (Dyar)
Figures 241, 720
Myelois glendella Dyar, Journ. New York Ent. Soc., vol. 14, p. 30,
1906.
Promylea glendella (Dyar) Barnes and McDunnough, Check list
of the Lepidoptera of Boreal America, No. 5584, 1917.—
McDunnough, Check list, No. 6147, 1939.
There is nothing to distinguish this from many of our
specimens of lunigerella from Washington and British
Columbia except some slight differences in genitalia
of very doubtful significance, and the name should prob-
ably go into synonymy; but until material is available
from intervening areas and something is known of the
life history, glendella may be retained as a possible local
race. It certainly is nothing more than that.
TypPE tocaLity: Glenwood Springs, Colo. (type in
USNM).
Foop prant: Unknown.
Represented in the National Museum by two males
and three females from the type locality (Aug., Sept.,
Oct.). In addition to the foregoing there are before me
five examples of another variety of lunigerella from Fal-
len Leaf Lake, Calif., Aug. 8 and 12, 1932, H. H. Keifer,
collector. They are quite different in color, having
paler gray forewings with much fainter transverse dark
markings and more whitish hind wings than our other
examples of lunigerella. They are probably nothing
but a color form and for obvious reasons are going with-
out a name at present.
131. Promylea dyari, new name
FIcuRE 243
Dioryctria zimmermani Druce (not Grote), Biologia Centrali
Americana, Lepidoptera Heterocera, vol. 2, p. 564, 1899.
Dioryctria drucet Dyar, Ins. Inse. Menstr., vol. 7, p. 44, 1919
(preoccupied).
Similar to lunigerella except: Larger and darker; the
paler areas of forewing a dark ashy gray; outer black
border of antemedial line broader, forming a small tri-
angle on costa; inner border of subterminal line also
slightly stronger, black; the subterminal line itself is
more sharply angled below costa and from about vein 6
proceeds to inner angle in a slanting almost straight line
(a difference that strikes the eye but is hardly signifi-
cant; for on some examples of Junigerella the subterminal
line is similarly shaped); terminal dots confluent, form-
ing a conspicuous black line along termen. Alar ex-
panse, 30 mm.
Male genitalia with tezumen and vinculum somewhat
stouter (broader in proportion to their width) than those
of lunigerella; penis armed with a weak cornutus.
TYPE Locatity: Rinconada, Vera Cruz, México (type
in USNM).
Foop prant: Unknown. sods
The species is known only from the male type which
both Druce and Dyar mistook for a female. ‘The refer-
ence to Dioryctria is difficult to understand in Dyar’s
case; for vein 3 of hind wing is appreciably too short for
that genus. The transfer of “Dioryctria drucer Dyar”
and “‘Nephopteryx druceit Ragonot”’ to the genus Promy-
lea makes the former a secondary homynym and necessi-
tates the newname. It is possible that the two “drucei”
represent only different sexes of one species; but this
cannot be determined without more material of each, so
for the present they must be treated as separate species.
132. Promylea druceii (Ragonot), new combination
FIcuRE 722
Nephopteryx drucezi Ragonot, Nouv. Gen., p. 15, 1888; Mono-
graph, pt. 1, p. 801, 1893.
Lhave not seen any specimens matching Ragonot’s de-
scription or figure (Monograph, pl. 9, fig. 17) ; but Ihave
before me an excellent photograph of the type supplied
by Tams. This shows a moth similar to dyari but with
the dark markings (especially the dark borders of the
transverse lines) much more|expanded, the outer border
of the antemedial line forming a large triangle on costa.
According to Ragonot these dark areas have a decided
purple tint, rather than the dull black or blackish brown
of dyart. The spot on the inner margin before the ante-
median line is also purplish black rather than orange (as
AMERICAN’ MOTHS OF THE SUBFAMILY PHYCITINAE 67
in dyari) and there is considerable purplish dusting on
the paler areas of the wing. The moth is also smaller
than Dyar’s type. Alar expanse, 24 mm.
Typz Locauity: Totonicapin, Guatemala (type in
BM).
Foop prant: Unknown.
Represented only by the type.
133. Promylea mindosis Dyar
Figures 242, 723
Promylea mindosis Dyar, Ins. Insc. Menstr., vol. 10, p. 172, 1922.
A narrow-winged, suffused, dark species. Ground
color of forewing very dark gray-brown; the blackish
transverse lines narrow and faint; antemedial line ob-
solete except for its faint outer border; subterminal line
distinguishable but faint; discal spots at end of cell
weak but apparently separated. Hind wing very pale
smoky fuscous, translucent. Alar expanse, 25-26 mm.
Male genitalia with penis with weak cornutus.
Female genitalia with siznum developed as a small
plate bearing a minute, knoblike projection.
Typr Locauity: Mexico City, Mexico (Aug.; type in
USN).
Foop pirant: Unknown.
Known only from the type series of two males and
one female from the type locality.
134. Promylea dasystigma Dyar
Fiaure 724
Promylea dasystigma Dyar, Ins. Insc. Menstr., vol. 10, p. 172,
1922.
Similar to mindosis, differing only in slight details:
The forewing is a trifle darker, more blackish than
brownish gray; the subterminal line ends on inner
margin in a small but distinct white spot; discal spots
fused into a thin, blackish lunule on discocellular vein.
The apron from the eighth-segment collar is differently
shaped and larger and the intersegmental pocket be-
tween seventh segment and collar proportionally wider
than those of mindosis (compare figs. 723a and 724a).
Alar expanse, 25-26 mm.
Female genitalia with signum a very weak patch of
scobinations.
Typ Locatity: Mexico City, Mexico (Aug.; type in
USNM).
Foop puant: Unknown.
Known only from the two females of the type series.
32. Genus Anadelosemia Dyar
Anadelosemia Dyar, Ins. Insc. Menstr., vol. 7, p. 51,1919. (Type
of genus: Nephopteryx senesciella Schaus).
Characters of Promylea except: Forewing with vein
10 always from the cell and not closely approximate to
the stalk of 8-9. Hind wing with vein 2 from close to
lower outer angle of cell; 3 from the stalk of 4-5 or at
least anastomosed with it for a short distance; 4 and 5
stalked for over half their lengths beyond cell; cell about
one-third the length of wing. Female abdomen with
eighth-segment collar simple; no dorsal pocket between
seventh and eighth segments. On male a hair-pencil
from lower, outer side of metathorax near base of leg.
Very close to Promylea but apparently distinct
enough. The general habitus of the moths is similar
except that the species of Anadelosemia (except for
obstitella) are decidedly smaller.
135. Anadelosemia senesciella (Schaus)
Figures 11, 244
Nephopteryx senesciella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol.
11, p. 251, 1913.
Anadelosemia senesciella (Schaus) Dyar, Ins. Insc. Menstr., vol.
7, p. 52, 1919,
Forewing ashy white, the basal area to antemedial
line stained with pale brown; costal edge at base black;
antemedial line white, broad and oblique from costa to
lower fold, thence crescentiform to inner margin, on
upper half bordered by a broad blackish spot, diffused
on costa and continued below fold as a narrow line to
inner margin, a weaker dark inner border on lower half;
a dark (brownish) shade extending obliquely across the
wing from costa just before subterminal line to near
middle of inner margin, irregular and more or less
diffused over remainder of outer area; discal spots dis-
tinct, blackish (on this and most other species of the
genus different on opposing forewings, separated or
fused into a line); subterminal line sinuate-dentate,
rather close to termen, bordered inwardly by a few
blackish dots and outwardly by a black line which be-
gins as a strong dash on costa; a row of blackish dots
along termen. Hind wing semihyaline tinted with
brown; the veins darkened and a narrow dark shade
along termen. Alar expanse, 15 mm.
Male genitalia distinguished chiefly by the shapes of
uncus, harpe, and vinculum (fig. 244); penis armed with
an elongate, narrow, flattened, bladelike cornutus.
TYPE LOCALITY: Juan Vifias, Costa Rica (Jan.; type
in USNM).
Foop prant: Unknown.
Known only from the male type.
136. Anadelosemia tecmessella (Schaus)
Ceracanthia tecmessella Schaus, Ann, Mag. Nat. Hist., ser. 8,
vol. 11, p. 251, 1913.
Anadelosemia tecmessella (Schaus) Dyar, Ins. Insc. Menstr., vol.
7, p. 52, 1919.
Forewing dull ashy white; an oblique blackish shade
at base; antemedial line narrow, white, a broken narrow
inner black border indicated; bordering the antemedian
line on costal half, a rather large triangular bronzy
brown spot; this color diffused outwardly along costal
edge; the oblique dark shade across wing beyond cell
extended to include the remainder of the outer area;
subterminal line but slightly paler than the brownish
color of outer area, otherwise as in senesciella; discal dots
distinct, blackish brown, well separated. Hind wing
pale smoky brown, veins and terminal margin darker.
Alar expanse, 14 mm.
68 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Typ LocaLity: Avangarez, Costa Rica (uly; type
in USNM).
Foop puant: Unknown.
Known only from the female type. It lacks an
abdomen so genitalia could not be studied. However,
the coloration of forewing suggests a species distinct
from anything else in the genus.
137. Anadelosemia fifria Dyar
FiauRE 726
Anadelosemia fifria Dyar, Ins. Insc. Menstr., vol. 7, p. 52, 1919.
Forewing similar to that of senesciella except: Whitish
ground color more extended filling most of outer area; a
dark brown shading at extreme base; dark shadings
fuscous brown rather than blackish; antemedial white
line narrow, its'dark borders also narrower; the post
media, transverse dark shade also narrower; terminal
dots more or less confluent. Alar expanse, 15 mm.
Female genitalia with signum, developed as a narrow,
small, shallow, granulate cup.
TYPE LocaLity: Cayuga, Guatemala (type in USNM).
Foop prant: Unknown.
Represented only by the female type and one other
female from the type locality (May).
138. Anadelosemia base Dyar
FIGURE 727
Anadelosemia base Dyar, Ins. Insc. Menstr., vol. 7, p. 52, 1919.
Similar to the foregoing species (jifria) except: Dark
markings a paler brown; the outer dark border of costal
half of antemedian line a small triangulate spot with
some extension outward on costal edge; terminal dots
rather weak but not confluent. Alar expanse, 15 mm.
Female genitalia without signum; ductus bursae pro-
portionally much longer than that of jifria; ductus
seminalis from near middle of ductus bursae. (In the
other species it branches off from the ductus bursae
very near its Junction with the bursa copulatrix.)
TYPE LocaLity: Cayuga, Guatemala (type in USNM).
Foop prant: Unknown.
Known only from the female type and one other
female from the type locality. Superficially hardly dis-
tinguishable from jifria. However, the differences in
their female genitalia suggest two distinct species.
139. Anadelosemia obstitella (Schaus), new combination
FicurE 728
Nephopteryzx obstitella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol.
11, p. 251, 1913.
The largest and most strikingly marked species in the
genus; the antemedial and subterminal lines shining
white; the inner, black bordering line of the former
continuous from costa to inner margin and strongly con-
trasted against the ashy gray ground color of the basal
area of the forewing; outer black border on costal half
of antemedial line and the black dashes preceding and
following the subterminal line enlarged and well con-
trasted. Hind wing semihyaline white with a very faint
brownish tint; veins faintly darkened; a narrow, pale
fuscous shading along termen, especially towards apex.
Alar expanse, 22 mm.
Female genitalia with signum present as a narrow,
weak, elongate, irregular scobinate patch (see enlarge-
ment). The shape and development of signum is a
character of very doubtful value in this genus and
probably subject to considerable individual variation.
Type LocaLity: Mount Pods, Costa Rica (type in
USNM).
Foop prant: Unknown.
Known only from the female type.
140. Anadelosemia texanella (Hulst), new combination
Figures 246, 729
Myelois texanella Hulst, Canadian Ent., vol. 24, p. 60, 1892.
Myelois dulciella Hulst, Canadian Ent., vol. 32, p. 176, 1900.
Tacoma texanella (Hulst) Barnes and McDunnough, Contribu-
tions, vol. 3, p. 193, 1916; vol. 4, p. 174, 1918—MceDun-
nough, Check list, No. 6143, 1939.
Forewing ashy white (due to a fine, sparse peppering
of fuscous scales on the white ground color); costal edge
at extreme base black; antemedial line not differenti-
ated except by the narrow, curved, black line forming
its outer border from costa to inner margin and by @
preceding brown spot on inner margin; subterminal line
sinuate, narrow, bordered inwardly by a continuous
black line and outwardly by a faint, narrow, brownish
shade continued from a blackish dash on costa; discal
spots black, separated. Hind wing smoky white, dark-
ening outwardly; a fine brown line along termen. Alar
expanse, 13-16 mm.
Male genitalia figured from type of dulciella. They
exhibit several specific characters: a long, slender,
strongly sclerotized, spinelike, apical projection from
enathos (the other species whose males are known show
no such structure, the only elements attached to the
lateral arms of gnathos at their junction being the
transtilla and the base of the more or less sclerotized
subanal plate); tezumen considerably elongated in pro-
portion to the vinculum; harpe short and broad; penis
finely spined at apex. Female genitalia without signum;
ductus seminalis from ductus bursae near junction of
bursa copulatrix. The distinctive female structural
character is the shape of the eighth-segment collar (fig.
729).
TypPE LocaLitixs: Blanco County, Tex. (texanella,
in AMNH, ex Rutgers); Hastings, Fla. (dulciella, in
AMNG, ex Rutgers).
Foop piant: Unknown.
DistrisuTion: UnitEpD States: Texas, Blanco
County, San Benito (Apr., Sept.); Florida, Hastings
(Oct.). Purrto Rico: San Germén (Apr.); Cusa: Santa
Clara, Central Soledad (‘“‘E. EH. A. Cuba, Ento. no.
10234,” May).
The Puerto Rican and Cuban examples are males
which agree in every detail with the type of dulciella.
The type of tezanella lacks an abdomen, so its genitalia
could not be checked (the other Texas examples are
females) ; but careful study of the two types discovered
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 69
no difference that would justify any doubt of the syn-
onymy proposed by Barnes and McDunnough (1918).
141. Anadelosemia condigna, new species
Fieures 245, 730
Forewing similar to that of teranella except: Outer
black border of antemedial line nearer middle of wing,
nearly vertical and more denticulate; subterminal line
with a discontinuous, blackish outer border; some red-
dish spotting in the postmedial area near inner margin;
costa at base not black, but a transverse black marking
at extreme base; discal spots confluent, forming a black-
ish or reddish brown line along discocellular vein. Hind
wing semihyaline white with a narrow fuscous shade
along termen. Alar expanse, 15-18 mm.
Male genitalia similar to those of senesciella, but dif-
fering in the shapes of uncus, harpe, and vinculum; penis
armed with a narrow, flat, bladelike cornutus similar
to that of senesciella. Female genitalia without signum,
similar to that of texana except for the shape of the
eighth-segment collar (fig. 730).
TyprrE Locauity: Prescott, Ariz. (type in USNM,
61329; paratypes in Cornell Univ. and BM).
Foop prant: Unknown.
Described from male type from the type locality
(July) and seven male and four female paratypes from
the Baboquivari Mts., Pima County, Ariz., collected by
O. C. Poling, May 1-15, 1924.
33. Genus Dasypyga Ragonot
Dasypyga Ragonot, N. Amer. Phycitidae, p. 5, 1887; Monograph,
pt. 1, p. 206, 1893.—Hulst, Phycitidae of N. Amer., p. 138,
1890.
Tongue well developed. Antenna weakly pubescent.
Labial palpus upcurved, reaching to vertex (female) or
nearly to it (male) ; third segment half as long as second,
acuminate. Maxillary palpus small, squamous. Fore-
wing with some raised scaling on basal area (probably
only a specific character); 11 veins; vein 2 from before
but near lower outer angle of cell; 3 from the angle,
closer to 4-5 than to 2; 4 and 5 shortly stalked; 6 from
below upper angle of cell, straight; 10 from the cell,
closely approximate for some distance with the stalk of
8-9; male without costal fold. Hind wing with vein 2
from near lower outer angle of cell; 3 from the angle,
connate with the stalk of 4-5; 4 and 5 long stalked (for
about two-thirds of their lengths); 7 and 8 closely ap-
proximate or contiguous for a short distance beyond
cell; cell one-third the length of wing; on male with anal
angle folded into a thickened pocket containing a hair
tuft. Eighth abdominal segment of male with sternite
developed as a narrow, sclerotized pocket.
Male genitalia similar to those of Anadelosemia ex-
cept: Sacculus of harpe strongly sclerotized and for half
its length developed as a free arm extending across
harpe; cornutus well developed; a pair of hair tufts from
intersegmental area adjacent to outer surfaces of the
sacculi at their bases.
Female with signum a small patch of scobinations;
ductus bursae considerably longer than bursa, tubular
and strongly sclerotized for about one-fifth its length
from genital opening, and for over half its remaining
length sclerotized, slightly flattened and bent (sinuate) ;
ductus seminalis from bursa near its junction with
ductus bursae.
A distinct genus distinguished from the other genera
with similar transtillae and venation by its stout, greatly
produced sacculus and sclerotized ductus bursae. Con-
tains one North American species.
142. Dasypyga alternosquamella Ragonot
Ficures 12, 247, 719
Dasypyga alternosquamella Ragonot, N. Amer. Phycitidae, p. 5,
1887; Monograph, pt. 1, p. 206, 1893.—Hulst, Phycitidae
of N. Amer., p. 138, 1890.—Heinrich, Proc. U. S. Nat. Mus.,
vol. 57, p. 84, 1920 (larva, pupa, life history).—Essig,
Insects of western North America, p. 709, 1926.—McDun-
nough, Check list, No. 6145, 1939.
Dasypyga alternosquamella stictophorella. Ragonot, N. Amer.
Phycitidae, p. 5, 1887; Monograph, pt. 1, p. 206, 1893.
Forewing pale salmon pink; basal area black dusted
with white, this black area (from the upper vein of cell
to inner margin) extending almost to middle of wing;
the black and white scaling coarse and more or less
raised in base and along outer margin of antemedial
line which cuts the black areas as a narrow, pale (och-
raceous red), obliquely curved line; subterminal line
absent, but along termen a rather broad border of coarse
black and whitish scales; on most specimens a clear
white line extends outward from lower angle of cell
along vein 5 towards and sometimes to the dark terminal
border and usually enclosing a detached patch of black
scales forming a discal spot at end of cell; above this
(below vein 6) a fine red line runs to near outer dark
margin and thence angles sharply to apex; on well
marked and fresh specimens traces of a similar red line
on the Jower fold in outer area. Hind wing very pale,
shining, smoky fuscous with a faint dark line along
termen. Alar expanse, 19-24 mm.
Genitalia as given for the genus; figured from reared
examples; cornutus of male penis about half as long as
aedeagus, somewhat flattened, twisted, and bluntly
pointed; penis also minutely scobinate towards apex.
TyprE LocaLity: California (type in Paris Mus.).
Foop puant: Razamofskya cryptopoda.
DistrisutTion: Unirep Status: California, Lake
Arrowhead (May); Arizona, Mohave County (Sept.),
Williams (June); Colorado, Glenwood Springs, Monu-
ment (May, June, Aug.); Washington, Bellingham
(June), Seattle. Canapva: British Columbia, Kaslo
(June).
A species that can be at once recognized by its pecu-
liar markings and coloration. The white longitudinal
line on forewing is of varying length on different speci-
mens but is present on all that I have seen, though
sometimes weak. Its presence is the characteristic
feature given by Ragonot for his variety stictophorella.
It is presumably absent from the type of alternosqua-
mella. Isuspect that the latter is an individual variant.
Notes on the life history and descriptions of larva and
pupa are given in my paper.
70 UNITED STATES ‘NATIONAL MUSEUM BULLETIN 207
Genus 34: Rampylla
[Male: Venational division D. Transtilla present but variously
modified. Harpe with apex of sacculus produced, strongly
sclerotized and pointed. Hind wing triangulate; anal angle
folded and produced; sex tufts and scalings on lower surface.
Female: Venational division B. Bursa and ductus bursae
simple; ductus seminalis from bursa. Hind wing with cell one-
third the length of wing.]
34. Genus Rampylla Dyar
Rampylla Dyar, Ins. Inse. Menstr., vol. 7, p. 84, 1919.
of genus: Rampzylla orto Dyar).
Tongue well developed. Antenna pubescent; shaft
of male slightly thickened. Labial palpus upturned,
not reaching vertex in male, a trifle longer in female;
second segment slightly rough scaled beneath; third
segment bluntly acuminate, about half the length of
second (shorter on male than female). Maxillary
palpus squamous, small. Forewing smooth except for
a slight, projecting scale tuft from inner margin near
base on male; 11 veins; vein 2 from before lower outer
angle of cell; 3 from the angle, well separated from 2;
4 and 5 approximate at base and for a very short dis-
tance beyond; 6 from below upper angle of cell, straight;
10 from the cell; male without costal fold. Hind wing
with vein 2 from before the lower outer angle of cell;
3 from the angle and in the male almost as long as vein
2; in the female considerably shorter; 4 and 5 very
shortly stalked or contiguous for a short distance from
cell, connate with 3; 7 and 8 contiguous or closely ap-
proximate for a short distance from cell; cell in male
one-fourth the length of wing, in female approximately
one-third; discocellular vein curved; on male anal area
(involving vein 1a) thickened and folded, forming a
produced pocket, enclosing a long hair pencil; under-
side of male wing with roughened scale or hair tufts on
some of the veins. Highth abdominal segment of male
with sternite developed as a narrow, sclerotized pocket.
Metathorax of male with a stout pencil of spatulate
scales from just above base of leg.
Male genitalia with transtilla present, variously modi-
fied (greatly reduced in lophotalis). Harpe with sac-
culus produced into a strong free hook at apex. A long
free spine associated with anellus (as in the four preced-
ing genera).
Female genitalia with bursa more or less finely scobin-
ate and with some concentration of these fine scobina-
tions but no. definable signum; ductus bursae simple,
short (shorter than bursa except in lophotalis). An
invaginated, sclerotized, dorsal pocket at apical end of
seventh abdominal segment or (in lophotalis) a sclero-
tized, granulate, dorsal pocket between ovipositor and
eighth-segment collar.
The length of vein 3 in proportion to 2 of hind wing
places the males in our venational division D and, in
conjunction with their decidedly triangulate hind wings
and the sex-scalings and tuftings on their under surfaces,
readily distinguishes the genus. The females on hind
wing venation fall into division B. The four species
here recognized are all tropical American. They ex-
(Type
hibit distinct specific differences in genitalia, color, and
maculation. The transverse lines of forewing, in their
rather close approximation and narrow black borders,
are similar to those of Coptarthria to which Rampylla
seems most nearly related.
143. Rampylla orio Dyar
Fieures 45, 248
Rampylla orio Dyar, Ins. Insc. Menstr., vol. 7, p. 84, 1919.
Forewing violaceous gray; under magnification ex-
treme base and a rather broad area along inner margin
to beyond middle tinted with purplish red; antemedial
line obsolete; subterminal line narrow, blackish, rather
close to and nearly parallel with termen, vertical from
costa to vein 8, inwardly angled between veins 8 and 5,
thence oblique to inner margin, bordered inwardly by a
faint ocherous shade and outwardly by a narrow ocher-
ous line; discal dots at end of cell separate, ocherous; a
thin dark line along terminal margin. Hind wing trans-
lucent white with a fuscous shade at apex and anal area
yellow; veins not appreciably darkened; terminal mar-
gin darkened only towards apex; on underside of hind
wing (male) a yellow hair tuft at origin of veins 4 and 5
from cell; a white fringe continuing outwardly on these
veins and a similar white fringe on vein 7 above the
yellow hair tuft. Alar expanse, 22 mm.
Male genitalia with lateral portion of uncus on each
side produced into a broad, strongly sclerotized, projec-
tion armed at apex with a cluster of long, slender, strong,
black spines. ‘Transtilla produced caudally into a bi-
lobed, scobinate process fusing with reduced arms of
gnathos. Tegumen with a long, stout, curved, free arm
arising from base at each ventrolateral angle. Harpe
with produced sacculus developed as a long, stout, taper-
ing hook, curved across face of harpe. Anellus heart-
shaped; associated spine straight. Aedeagus partially
sclerotized; penis with some faint, sclerotized wrinklings
and a few microscopic scobinations, otherwise unarmed.
Tyrr Locatity;: Zacualpaén, México (type in USNM).
Foop prant: Unknown.
Known only from the male type.
144, Rampylla polydectella (Schaus)
Figure 732
Salebria polydectella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11,
p. 250, 1913.
Rampylla polydectella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7,
1919.
Forewing brownish gray with a faint purplish tint; a
narrow border along inner margin between the trans-
verse lines and a somewhat broader area along termen
dusted with dull, grayish white; antemedial line narrow,
faint, pale gray, indicated chiefly by its narrow, black
outer-bordering line, the line well out towards middle of
wing and slightly sinnous, nearly vertical; subterminal
line equally thin and pale gray with a similar black inner
border, angled outward slightly at middle and nearly
parallel with termen; a black line along terminal margin;
discal spots small, confluent, ocherous; faint indication
AMERICAN MOTHS OF THE. SUBFAMILY PHYCITINAE vel
of black streaklets on veins 2, 3, and 4 just beyond cell.
Hind wing glossy brown-gray; the veins faintly dark-
ened and a fine dark line along termen. Alar expanse,
20 mm.
Female genitalia distinguished chiefly by the shape
of eighth-segment collar and the invaginated, sclero-
tized pocket from seventh segment.
TYPE Locatity: Juan Vifias, Costa Rica (June; type
in USNM).
Foop puant: Unknown.
Known only from the female type.
145. Rampylla subcaudata (Dyar), new combination
Ficures 249, 733
Cerocanthia subcaudata Dyar, Ins. Insc. Menstr., vol. 7, p. 42,
Forewing ocherous gray, the basal area to antemedial
line purplish tinted; a similar purplish shade along outer
border of the subterminal line; a large, somewhat
darker (fuscous) patch surrounding the discal spot and
extending from antemedial almost to subterminal line
and from vein 2 to costa, darkest on veins 2 to 4; ante-
medial line faint, narrow, ocherous, indicated by a thin
blackish brown line forming its outer border, the latter
near middle of wing, slightly sinuate and nearly vertical;
subterminal line with a narrow, blackish brown, inner
border, slightly outbent or angled at middle; discal
mark a narrow ocherous spot along discocellular vein;
a narrow blackish line along terminal margin. Hind
wing of male subpellucid white with a faint yellowish
tint, decidedly ocherous along inner margin; of female
with pale smoky tint; underside of male hind wing with
a yellowish scale tuft covering the bases of veins 2 to
5. Alar expanse, 16 mm.
Male genitalia with margins of uncus evenly rounded,
lacking any sclerotized projections. Harpe with
apical projection of sacculus a slender, upcurved
hook; a long, stout, hair and scale tuft from a pad ad-
jacent to base of sacculus. Penis armed with a narrow,
weakly sclerotized, bladelike cornutus. Female geni-
talia similar to those of polydectella, differing only in
slight details in the shape of the eighth-segment collar
and the sclerotized pocket from seventh segment (fig.
733a).
TypE Locauity: Cayuga, Guatemala (May; type in
USNM).
Foop PLant: Unknown.
In addition to the male type there is one other speci-
men in the National Collection from Quirigué, Guate-
mala (May), a female with slightly darker hind wings
whose genitalia are here figured. In addition there are
before me two somewhat larger specimens (19 mm.), a
male in the Janse Collection from San José, Costa Rica
(“H. Schmidt, 8-11”), and a female from the British
Museum labeled ‘‘S. E. Brazil, E. D. Jones, 1920-303.”
The genitalia of the latter are almost identical with
those of polydectella. The genitalia of the male are like
those of the type of subcaudata. The two specimens
are identical in size, color, and markings. Except for
size (which is not significant) they agree superficially
with the type of subcaudata. Despite the differences in
fore and hind wing coloration I suspect that subcaudata
may not be specifically distinct from polydectella.
146. Rampylla lophotalis, new species
Figures 250, 731
Similar to subcaudata except: Ground color of fore-
wing more yellowish; the dark areas, especially just
preceding the antemedial line and in the dark area’ sur-
rounding the discal spot, blackish; dark lines bordering
the transverse lines blackish rather than blackish brown;
antemedial line farther out on wing, extending from
midcosta to inner margin beyond middle, narrowing the
area between antemedial and subterminal lines. Hind
wing with the scale tuft on underside blackish rather
than yellow, the blackish shade extending to the costa
and visible through the wing from above. Alar ex-
panse, 17.5-18 mm.
Male genitalia with two pairs of strongly sclerotized,
curved, lateral projections from triangulate uncus.
Gnathos entirely absent (unless the lower pair of pro-
jecting arms from uncus can be interpreted as lateral
arms of a gnathos, which is very doubtful, as there is
no separation whatever between them and the uncus).
Transtilla reduced to a shortened and slender, trans-
verse, centrally bent band. Tegumen with a short,
two-pronged projection from each ventrolateral angle.
Anellus and harpe as in subcaudata. Female genitalia
with a thick, sclerotized roll on the back of ovipositor;
a broad, coarsely granulate pocket between ovipositor
and collar; eighth-segment collar narrow; ductus bursae
slightly longer than bursa.
Typr Locatity: Jalapa, México (type in USNM,
61330).
Foop piant: Unknown.
Described from male type and one female paratype
from the type locality and one male paratype from
Volcén Santa Maria, Guatemala (Nov., Schaus and
Barnes, collectors). The male holotype bears a label
in Hampson’s handwriting ‘‘Cerocanthia lophotalis
Hampson, o type.”’ As far as I know Hampson never
published a description of the species.
Genus 35: Fulrada
[Venational division D. Hind wing with veins 4 and 5 contiguous
or closely approximate beyond cell (not stalked) ; cell one-fourth
the length of wing; discocellular vein incomplete. Eighth ab-
dominal segment of male with broad ventral tuft. Transtilla
vestigial. Harpe with apex of sacculus not produced.]
35. Fulrada, new genus
Typr or Genus: Dasypyga querna Dyar.
Tongue well developed. Antenna weakly pubescent;
shaft of male simple. Labial palpus upturned, slender,
reaching to vertex; third segment slightly shorter than
second, acuminate. Maxillary palpus squamous, small,
appressed to face. Forewing smooth; 11 veins; vein 2
72 UNITED STATES NATIONAL MUSEUM BULLETIN 207
from before but near lower outer angle of cell; 3 from
the angle, nearly equidistant from bases of 2 and 4; 4
and 5 closely approximate or connate and contiguous,
for a short distance beyond base; 6 from below upper
angle, straight or very slightly bent towards base; 10
from the cell, approximate to the stalk of 8-9; male
without costal fold. Hind wing with vein 2 from before
lower outer angle of cell; 3 from the angle, long (its
free; length slightly shorter than 2); 4 and 5 contiguous
and. closely approximate for slightly less than half their
distances from angle of cell (not stalked); 7 and 8 closely
approximate beyond cell; cell one-fourth the length of
wing; discocellular vein incomplete. Highth abdominal
segment of male with a broad, ventral hair or scale tuft
and sternite developed as a narrow sclerotized pocket.
Male genitalia with uncus subtriangulate, its apical
margin truncate. Gnathos represented only by its
lateral arms, to which (at their junction) is attached
the base of subanal plate. Transtilla represented only
by a bilobed central vestige behind (above) the anellus.
Harpe with sacculus not produced. Anellus semi-
tubular (querna) or a broad slightly curved plate with
a greatly reduced, associated, free spine (carpasella).
Vinculum stout, as broad or nearly as broad as long;
terminal margin broad and slightly rounded.
The foregoing description is incomplete, as females
are unknown. ‘The genus is apparently close to both
Anadelosemia and Rampylla, differing from the former
chiefly on hind wing venation and from the latter on
secondary male characters.
147. Fulrada querna (Dyar), new combination
Fieure 251
Dasypyga querna Dyar, Proc. U. S: Nat. Mus., vol. 47, p. 331,
1914.
Forewing ocherous gray with a faint dusting of
reddish scales on basal area and a very faint reddish
tint over the area below the discal spots; a scattered
blackish powdering in costal areas; antemedial line
oblique, indicated only by a row of (4 or 5) well sepa-
rated black dots forming its inner border and an outer
black spot on costa; discal dots at end of cell, small,
separated, black; a row of small but distinct black dots
along termen. Hind wing translucent, smoky white,
darker towards apex; veins faintly darkened and a
narrow dark line along termen. Alar expanse, 11 mm.
Male genitalia with anellus semitubular, oblong,
bottle shaped, more or less involved with the vestigial
hairy-lobed transtilla. Harpe with a strongly haired,
transverse, sclerotized ridge extending from near base
of costa to lower outer angle of cucullus. Aedeagus
short; penis armed with a very weak, flattened cornutus.
Tuft on eighth abdominal segment a row of very fine,
hairlike scales.
Typn Locauity: La Chorrera, Panamé (May; type
in USNM).
Foop prant: Unknown.
Represented only by the male type.
148. Fulrada carpasella (Schaus), new combination
FigurE 252
Piesmopoda carpasella Schaus, Zoologica, vol. 5, No. 2, p. 47,
1923.
Forewing white finely irrorated with black and
brown; some pale tawny shading on the white ante-
medial line along the inner margin of its black outer
border, also postmedially below vein 2 and on discocel-
lular vein; base black, this shade expanding’ obliquely
to inner margin and outwardly edged by some pale
tawny scaling; antemedial line oblique, indicated
chiefly by its black outer border, the latter strongly
contrasted from costa to lower margin of cell, very faint
from cell to inner margin, slightly outcurved from costa;
subterminal line parallel with termen, slightly sinuous
and with well-contrasted dark outer and inner borders,
the inner one a narrow black line, the outer a black,
angulate costal dash continued as a rather broad
brownish shade to tornus; discal dots separated, small
black dots on the outer angles of cell; a row of well-
contrasted black dots along termen. Hind wing
whitish towards base, shading to fuscous outwardly, a
rather broad brownish shade along termen. Alar
expanse, 12 mm.
Male genitalia with a vestigial free spine associated
with the broad, large, slightly curved plate’ of anellus.
Harpe with rather long, slender, erect clasper: Aedea-
gus rather long, slender, straight. Highth abdominal
tuft consisting of long, spatulate scales.
Typr LocaLity: Conway Bay, Indefatigable, Galé-
pagos Islands (Apr.; type in USNM). .
Foop pirant: Unknown.
Known only from the male type.
Genus 36: Scorylus
[Venational division B. Male antenna with shallow sinus in
shaft at base. Hind wing with veins 4 and 5 stalked for about
two-thirds; cell about one-third the length of wing; discocellular
vein complete, curved. Highth abdominal segment of male with
broad, ventral scale tuft. Metathorax with stout scale and
hair tuft near base of leg. Transtilla vestigial. Anellus with-
out trace of associated free spine. Harpe with apex of sacculus
produced.]
36. Scorylus, new genus
TYPE OF GENUS: Scorylus cubensis, new species.
Tongue well developed. Antenna finely pubescent;
the male shaft with a shallow sinus at base (a slight ex-
cavation covering several basal segments); within the
sinus and extending slightly past it a thin layer of modi-
fied appressed scales. Labial palpus upturned, reaching
vertex, second segment slightly rough scaled; third seg-
ment slightly shorter than second, acuminate. Maxil-
lary palpus squamous. Forewing smooth; 11 veins;
vein 2 from before lower outer angle of cell; 3 from the
angle, nearer to 4 than to 2;4 and 5 closely approximate
for a short distance from cell; 6 from below upper angle
of cell, slightly bent towards base; 8 and 9 stalked for
slightly more than half their lengths; 10 from the cell,
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 73
approximate to the stalk of 8-9; male without costal
fold. Hind wing with vein 2 from before, but close to
lower outer angle of cell; 3 from the angle, contiguous to
the stalk of 4-5 for nearly half the length of the stalk,
the free length of 3 decidedly shorter than vein 2; 4 and
5 stalked for about two-thirds their lengths; 7 and 8
closely approximate for some distance beyond cell; cell
about one-third the length of wing; discocellular vein
curved. Highth abdominal segment with broad, stout
ventral scale tuft and sternite developed as a narrow
sclerotized pocket. A stout scale and hair tuft from
metathorax at base of leg.
Male genitalia with uncus hoodlike. Gnathos well
developed, its apical projection a long, tapering spike.
Transtilla represented by a modified central vestige.
Harpe with sacculus produced at apex into a sclerotized
hook; a strong hair tuft from lobe near base of sacculus;
anellus a triangulate plate with short, blunt, broad lat-
eral lobes; aedeagus moderately stout, slightly swollen
and bent towards base; penis armed with an elongate,
narrow cluster of short, thin spines. Vinculum stout,
slightly tapering and somewhat longer than basal width.
Female genitalia without signum but with a concen-
tration of fine granulations in bursa near its junction
with ductus bursae; ductus bursae and its genital open-
ing simple; ductus seminalis from ductus bursae.
This genus shows affinities to both Fulrada and
Anadelosemia but is distinct from both. Its venation
is similar to that of Anadelosemia, to which it appears to
be most closely allied.
149. Scorylus cubensis, new species
Figures 253, 725
Forewing white with blackish fuscous and reddish
markings; basal area strongly irrorated with red and
some scattered reddish scaling in the median white area;
antemedial line indicated only by a blackish outer line
from costa, expanding into a black spot in cell, continued
thence as a very thin blackish fuscous line to inner mar-
gin, bordered outwardly (from cell to inner margin) by
an olivaceous patch; a faint triangulate olivaceous-
fuscous shade over outer area from just beyond middle
of inner margin to subterminal line at costa; a small
black spot on midcosta; subterminal line sinuate,
bordered inwardly by a strong, rather broad, blackish
line and outwardly by a fainter reddish line continued
from a strong black spot on costa; discal dots black,
more or less confluent along discocellular vein; below
them on the olivaceous fuscous shade a patch of reddish
scaling; a row of black dots along termen. Hind wing
translucent white; veins very faintly darkened near
outer margin; a narrow dark shade along termen. Alar
expanse, 12-13 mm.
Male genitalia with outer margin of uncus evenly
rounded. ‘Transtilla fragment in the shape of two short,
oblong, pointed plates weakly joined at their bases.
Apical projection of sacculus sharply upturned, mod-
erately long, thornlike. Terminal margin of vinculum
truncate. Female genitalia with bursa large, as long as
ductus bursae.
TYPE Locauity: Santiago Province, Cuba (type in
USNM, 61331).
Foop piant: Unknown.
Described from male type and one male and one fe-
male paratypes from the type locality (June, Dec.,
Schaus and Barnes, collectors).
Genera 37-39: Davara to Piesmopoda
[Venational division D. Forewing with veins 4 and 5 approxi-
mate for a short distance from cell. Hind wing with cell less
than one-third the length of wing. Male genitalia with uncus
bifid (divided to base); harpe with strong, hooked clasper from
near apex of sacculus.]
37. Genus Davara Walker
Davara Walker, List, pt. 19, p. 1020, 1859.—Hampson, in
Ragonot, Monograph, pt. 2, p. 530, 1901. (Type of genus:
Davara azonazsalis Walker.)
Homalopalpia Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 332, 1914.
(Type of genus: Homalopalpia dalera Dyar. New synonymy.)
Eucardinia Dyar, Ins. Insc. Menstr., vol. 6, p. 188, 1918; vol. 7,
p- 50, 1919. (Type of genus: Ulophora caricae Dyar.)
Tongue well developed. Antenna of male (except in
rufulella) with basal segment enlarged, deeply notched
and with a brush of short fine bristles in the notch
(fig. 254d); male shaft simple or flattened and dilated
towards base, pubescent. Labial palpus upcurved; on
male (except in rufulella) reaching well above vertex;
broadly scaled; third segment considerably shorter than
second. Maxillary palpus squamous. Forewing more
or less rough scaled at base on male; 11 veins; vein 2
from before, but near lower outer angle of cell;3 from the
angle, nearer to 4 at base than to 2; 4 and 5 approximate
for a short distance from cell; 6 from below upper angle
of cell, straight or very slightly curved towards base; 8
and 9 long stalked; 10 from the cell, approximate to the
stall of 8-9; male without costal fold. Hind wing with
vein 2 from before the angle of the cell; 3 from the
angle, long (its free length slightly shorter than 2); 4 and
5 anastomosed for slightly less than half their lengths
beyond angle of cell; 7 and 8 contiguous or shortly
anastomosed beyond cell; cell less than one-third the
length of wing; discocellular vein curved. Eighth
abdominal segment of male with sternite developed as
a narrow, sclerotized pocket.
Male genitalia with uncus bifid (divided to base).
Gnathos weak, a thin narrow band. Transtilla absent
(except in interjecta). Harpe with strong, hooked
clasper from near apex of sacculus; a long hair tuft from
lobe near base of sacculus. Anellus a narrow curved
plate with long, lateral arms; a stout, free spine associ-
ated with anellus. Penis armed with a thin, narrow,
curved, flattened, bladelike cornutus.
Female genitalia with two signa, developed as small,
granulate depressions; ductus bursae shorter than bursa,
with paired cuplike plates behind genital opening or
with genital opening simple (interjecta) ; ductus seminalis
74 UNITED STATES NATIONAL MUSEUM BULLETIN 207
from bursa near its junction with ductus bursae. Collar
of eighth segment complete.
Davara and. the two following genera (Sarasota and
Piesmopoda) form a compact group distinguished from
all other American phycitid genera by the peculiar bifid
development of their unci. Davara was described by
Walker on the basis of a single female which he mistook
for a male; it was referred as a synonym of the Old
World Phycita by Hampson (1903). Dyar did not
recognize it. His Homalopalpia was erected on male
antennal and palpal structures which normally should
be of generic value, but in this particular instance do
not seem to hold as separating Davara from Piesmopoda.
They fall down in the case of rufulella, which on genitalic
characters of both male and female must be referred to
Davara. In my opinion Davara and Piesmopoda should
be retained as separate genera. Their species differ in
habitus as well as in genitalic structure. In Davara the
male transtilla is absent and the female bursa always
has two signa. In Piesmopoda the transtilla is present
and developed as two long, slender, curved free arms
and the female’ bursa has a single signum or none.
Unfortunately an anomalous species (interjecta) seems
to upset the division. It has the female and all the
secondary male characters of typical Davara but male
genitalia of the Piesmopoda type. However, there are
some minor differences in its male and female genitalia
which may eventually allow its separation from both
Davara and Piesmopoda under a separate generic desig-
nation. For the present I am referring it tentatively to
Davara. Its distinctive characters are discussed more
fully under the specific description.
150. Davara caricae (Dyar), new combination
Fiaures 46, 254, 735
Ulophora caricae Dyar, Proc. Ent. Soc. Washington, vol. 14, p.
218, 1913.
Homalopalpia dalera Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 332,
1914. (New synonymy.)
Eucardinia caricae (Dyar) Dyar, Ins. Insc. Menstr., vol. 6, p.
139, 1918; Ins. Insc. Menstr., vol. 7, p. 50, 1919.—McDun-
nough, Check list, No. 6119, 1939.
Antenna of male with the brush in notch of basal
segment black; the basal segment itself a very pale buff;
basal segments of shaft decidedly flattened and broad-
ened and with black serrations on upper edge. Labial
palpus of male very broadly scaled, the second segment
reaching well above vertex; less broadly scaled and
somewhat shorter on female; reddish brown with a
peppering of pale buff scaling on outer side and on male
more or less shaded with blackish brown on inner side.
Forewing tan-gray shaded with reddish brown; basal
area, (on male) blackish brown ‘and rough scaled; a
whitish or pale buff shade precedes the antemedial line
and a similar transverse shade crosses the disk and
includes the discocellular mark; the space between this
transverse pale shade‘and antemedial line suffused with
reddish brown (on some female examples almost purplish
fuscous); a similar reddish brown suffusion over outer
area; antemedial line oblique, slightly angled between
cell and inner margin, faint, ocherous, bordered in-
wardly and outwardly by narrow dark lines; subterminal
line indistinct except for its brown inner and outer
borders, sinuate; discal spots fused into a narrow lunu-
late line on the discocellular vein. Hind wing soiled
white, with a narrow fuscous shade along termen. Alar
expanse, 14-18 mm.
Male genitalia with a cluster of fine, long, hairlik
spines surrounding the strong, free, forked spine asso-
ciated with anellus; lateral arms of anellus considerably
shorter than in other species of the genus. Female
genitalia with a pair of strong ventrolateral ridges on the
invaginated portion of the eighth-segment collar.
Tyre tocauities: Miami, Fla. (caricae, in USNM);
LaChorrera, Panama (dalera, in USNM),.
Food pLant: Carica papayae (larvae in the fruit).
Distrizution: Unitep States: Florida, Florida City
(Apr., May), Fort Pierce (May), Miami (Apr., Dec.),
Royal Palm State Park. Purrtro Rico: Bayamén
(June, Dec.), El] Yunque (Apr.), Jajoma Alta (June),
Lares (June, Sept., Nov.). Cusa: Baracoa (July, Aug.)
Santiago de las Vegas (Feb., Mé4r.), Santiago Province
(@Jan., June, Oct.). Harri: Damien (Aug.). Domini-
can ReEpusiic: San Francisco Mts. (Aug., Sept.).
Trinipap: Mt. Harris. GuatEemata: Cayuga (Apr.,
May), Chejel (June, Aug.), Purulhi (June, July).
Costa Rica: Guapiles (May), Juan Vinias (Feb., May,
June), Silio (May). Ecuapor: Quevedo (‘‘Nov.-
Dec.”). ;
The types of both caricae and dalera are males. Dyar
(1919) recognized the generic synonymy of Hucardinia
and Homalopalpia but never admitted the specific
identity of their types, although he had every reason
to suspect it. His designation of new Cuban types for
caricae in 1918 is obviously invalid, for he had previ-
ously (1913) designated Florida types for what he ad-
mitted was the same species under the same name. I
believe he was correct in his surmise that caricae is the
same as columnella Zeller, but as I have never examined
any Colombian examples of any Davara species I
hesitate to propose the synonymy.
151. Davara columnella (Zeller), new combination
Figure 736
Myelois columnella Zeller, Horae Soc. Ent. Rossicae, vol. 16,
pp. 209, 210, 1881.
Piesmopoda columnella (Zeller) Ragonot, Monograph, pt. 1, p.
161, 1893.
Homalopalpia columnella (Zeller) Dyar, Ins. Insc. Menstr., vol.
7, p. 49, 1919.
I have seen no specimens from the type locality but
have before me a photograph of the female type which
agrees with the females of a series from Costa Rica
identified by Schaus as columnella and correctly referred
by Dyar to his dalera, and I have little doubt that the
names columnella and caricae (=dalera) stand for the
same species. The female genitalia of Zeller’s type
(here figured) show some trifling differences in the size
of the plates behind genital opening and in the eighth-
segment collar from those of typical caricae; but these
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 75
are probably only individual differences. Variations
as great are exhibited among reared examples of caricae
from different localities. Alar expanse, 15 mm.
Typxr Locatity: Honda, Colombia (type in BM).
Foop puant: Unknown (presumably papaya).
152. Davara nerthella (Schaus), new combination
Ficure 738
Piesmopoda nerthella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol.
11, p. 247, 1913.
Homalopalpia euthales Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 403, 1914.
Homalopalpia nerthella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7,
p. 49, 1919.
Similar to caricae except averaging somewhat larger;
brush in notch of basal segment of male antenna
ocherous, the segment itself smaller; the subbasal pale
shade before the antemedial line more extended on the
male and much more so on the female, reducing con-
siderably the blackish brown shading of the basal area
and forming with the slightly paler antemedial line a
broad pale pinkish ocherous band; antemedial line
straight, not angled below cell, its inner and outer
bordering lines very faint and narrow, reddish brown;
discal dots separate (never fused) and often only the
lower one distinguished; on females more or less of an
ocherous tint over the brownish median shade following
the antemedial line, especially towards costa. Alar
expanse, 19-22 mm.
Male genitalia like that of the following species
(paranensis) except tuft from near base of sacculus pale
yellow. Highth-segment collar without ridges on the
invaginated portion; narrower on venter than in other
species; posterior ventral margin without notch or but
slightly notched. The extent of this notching is indi-
vidually variable in all the species and is not a reliable
character for specific separation.
TYPE LOCALITIES: Juan Vifias, Costa Rica (nerthella,
in USNM); Orizaba, México (euthales; in USNM).
Foop prant: Unknown.
DistrisutTion: Mfxico: Jalapa, Orizaba. Guatr-
MALA: Volcdén Santa Maria (June, July). Costa Rica:
Juan Vinas (Jan.); other Costa Rican females without
further locality designation in Janse Collection.
Doubtfully distinct from paranensis. The chief dif-
ferences between the males are in coloration and be-
tween the females in the width and notching of the
eighth-segment collar. There is the same amount of
individual variation in wing color and maculation as in
other species, some specimens generally paler than
others, some with the subterminal line distinct, others
with it almost obsolete.
Dyar (1919) recognized the synonymy of his euthales
with nerthella. The type of the former is a male, of
the latter a female.
153. Davara paranensis (Dyar), new combination
Fieure 255
Homalopalpia paranensis Dyar, Ins. Insc. Menstr., vol. 7, p. 49,
1919,
Similar to nerthella except brush in notch of basal seg-
ment of male antenna brownish; subbasal pale shade
before antemedial line of forewing much narrower, re-
stricted (especially on females) by a greater extension
of the black basal scaling; median area along costa
rather strongly tinted with ocherous drab or reddish
ocherous (on the type and one female). Alar expanse,
18-19 mm.
Male genitalia with no fine spine cluster surrounding
the free, forked spine associated with anellus. Tuft
from near base of sacculus black. Female genitalia
similar to those of azonazsalis.
Type LocaLity: Castro, Parané, Brazil (type in
USN).
Foop prant: Unknown.
At first glance Dyar’s type seems quite distinct from
males of nerthella, its palpi and antennae being much
darker and the pale transverse antemedial shade less
contrasted against the ground color. However the
specimen is stained and none of the other examples of
the species before me is in very good condition. Be-
sides the type, I have before me a female from the type
locality, another female from Santa Catarina, Brazil,
and a male from the British Museum collection from
Sao Paulo, Brazil, that is without abdomen but a good
match for Dyar’s type. Both it and the type had been
originally identified by Hampson as columnella Zeller.
I suspect that when additional South American ma-
terial is available paranensis will prove to be nothing
more than a variety of nerthella, and that eventually
both nerthella and paranensis will fall to azonarsalis of
Walker.
154. Davara azonaxsalis Walker
FIaure 737
Davara azonazsalis Walker, List, pt. 19, p. 1020, 1859.
Phycita azonazsalis (Walker) Hampson, in Ragonot, Monograph,
pt. 2, p. 531, 1901.
I have seen no specimens of Davara from the type
locality and none from anywhere of the size of Walker’s
type (30 mm.). A photograph of the type and its geni-
talia supplied by Tams are before me. The antemedial
line of forewing shows an angulation between cell and
inner margin similar to that on typical caricae. The
subterminal line is rather distinctly marked and the
basal area much like that of females of nerthella, but
not so strongly contrasted. None of these features,
however, is enough for specific separation.
The female genitalia show an appreciably wider
eighth-segment collar than that of nerthella and a dis-
tinct notch in its ventroposterior margins. The much
smaller female of paranensis from Castro has similar
but somewhat smaller genitalia.
TypE Locauity: Rio de Janiero, Brazil (type in Ox-
ford Univ. Mus.).
Foop piant: Unknown.
155. Davara (?) interjecta, new species
Fiaurus 256, 734
Male antenna with basal segment enlarged and
notched as in caricae, but the usual brush of fine bristles
76 UNITED STATES NATIONAL MUSEUM BULLETIN 207
in the notch here replaced by smooth, appressed, silvery
and ocherous scales; first segment of shaft broadly
flattened, the shaft shortly ciliate (the cilia about as
long as width of shaft). Labial palpus broadly scaled,
the second segment reaching well above vertex, deep
red-brown to blackish brown on outer side, the third
segment black scaled, at least on inner surface. Thorax
deep brown mixed with blackish and buff scaling. Fore-
wing reddish brown more or less shaded with black in
median area, the raised scaling at base mixed black and
dark red-brown; antemedial line obsolete or very faintly
indi¢ated on the paler specimens, when distinguishable
oblique, straight, narrow, pale buff; lower discal spot
at end of cell usually distinct, blackish; subterminal
line obscure or obsolete; an interrupted row of partially
confluent black dots along termen. Hind wing trans-
lucent, white, the veins more or less darkened, a narrow
fuscous shade along termen.
Female generally paler than the male. Palpi, head,
thorax, and forewing light reddish brown with a faint
sprinkling of whitish scales, especially bordering ter-
men; ground color darkened in median area along costa;
usually a broad, olivaceous shade along inner margin
at base; no defined transverse markings.
Alar expanse, 17-19 mm.
Male genitalia of the Piesmopoda type with incom-
plete transtilla developed as a pair of opposed, long,
slender, curved, strongly sclerotized arms, their apices
curved towards each other; anellus a narrow, broadly
V-shaped band, with long, curved, slender, haired lat-
eral arms, their apices bulbed. Female genitalia with
genital opening simple, no sclerotized plates behind the
opening.
Type tocauity: El Yunque, Luquillo Mts., Puerto
Rico (type in Cornell Univ.; paratypes in Cornell and
USNM, 61332).
Foop prant: Unknown.
Described from male type and two male and five fe-
male paratypes from the type locality (1,500-2,000 ft.,
Apr. 22, 23, and Mar. 29,1930, Cornell lot 795, sub. 38,
40, and 9, 'W. T. M. Forbes, ‘collector) and two male
paratypes from San Francisco Mts., Santo Domingo
(Sept. 1905, A. Busck, collector). The males, with the
exception of the holotype and one paratype, are badly
rubbed. The females are in better condition.
This species is referred with reservations to Davara.
In its structural characters it straddles both Davara and
Piesmopoda and fits comfortably in neither genus. Its
female genitalia are those of Davara except that the
usual sclerotized plates behind genital opening are lack-
ing. Its male genitalia are those of Piesmopoda except
that the apices of the elements of transtilla pomt toward
(rather than away from) each other; and the apices of
the lateral arms of anellus are swollen (bulbed). In all
known species of Piesmopoda the apices of the elements
of transtilla point away from each other and the apices
of the lateral arms of transtilla are pointed. The male
antenna of interjecta also is abnormal for Davara in that
there is no brush of fine spines in the notch of the basal
segment. In my opinion these differences could permit
generic separation of interjecta from both Davara and
Piesmopoda; but they are so slight that, without further
evidence from biology or the early stages, a new generic
designation does not seem justified at this time.
156. Davara rufulella (Ragonot), new combination
FicureEs 257, 739
Piesmopoda rufulella Ragonot, Bull. Soc. Ent. France, 1888,
p. cxxxix; Monograph, pt. 1, p. 165, 1893.
Male antenna simple. Labial palpus upturned, not
broadly scaled, cylindrical; reaching to slightly above
vertex on male, nearly to vertex onfemale; terminal
segment acuminate. Forewing pale red-brown; the
basal area a trifle paler with some faint olivaceous shad-
ing in inner margin; antemedial line oblique, straight,
obscure, indicated chiefly by its outer dark margin,
which begins as a blackish smudge on costa and contin-
ues to inner margin as a slight darkening of the ground
color; more or less blackish dusting in the cell beyond
antemedial line; on paler specimens some peppering of
white scales in median and outer areas, especially on
midcosta; subterminal line obsolete or very faintly indi-
cated; discal spots obscure, confluent along discocellular
vein.. Hind wing translucent white; the veins dark-
ened; a faint, narrow fuscous shade along termen. Alar
expanse, 13-16 mm.
Male genitalia without cluster of fine spines surround-
ing free spine associated with anellus, the free spine
itself short, stout, broadly forked. Lateral arms of
anellus straight. Harpe with apex of cucullus pointed;
clasper long, stout, strongly curved. Female genitalia
with two pairs of contiguous sclerotized plates behind
genital opening; ductus bursae sclerotized at genital
opening.
Type Locauity: Puerto Rico (type in Zool. Mus.
Univ. Berlin).
Foop piant: Unknown.
Distrizution: Purrro Rico: Bayamén (Apr., Sept. ),
Comerio (Nov.), Jajoma Alto (June), Lares (Dec. ), La
Sardinera (Dorado, June), Palmas Abajas (June), San
German (Apr.).
I have seen no specimens from any but Puerto Rican
localities. As mentioned in the discussion of the genus,
rufulella is aberrant in that it lacks the modified basal
segment of male antenna and the characteristic broad
scaling of the male labial palpi. However, the genitalia,
both male and female, are characteristic of the genus,
showing only specific differences from. other species of
Davara.
38. Genus Sarasota Hulst
Sarasota Hulst, Journ. New York Ent. Soc., vol. 8, p. 222, 1900.
(Type of genus: Sarasota plumigerella Hulst).
Cuba Dyar, Ins. Inse: Menstr., vol. 7, p. 50, 1919.
genus: Cuba furculella Dyar. New synonymy.)
Tongue well developed. Antenna simple in both
sexes, shaft weakly pubescent. Labial palpus up-
turned, reaching to slightly above vertex; third segment
slightly over half the length of second, acuminate.
Maxillary palpus small, squamous. Forewing smooth;
11 veins; vein 2 from before but near lower outer angle
(Type of
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE TE
of cell; 3 from the angle, approximately equidistant at
base from 2 and 4, 4 and 5 approximate for a short
distance from cell; 6 from upper angle of cell, slightly
bent towards base, connate with the stalk of 8-9; 10
from the cell, well separated from the stalk of 8-9; male
without costal fold. Hind wing with vein 2 from before
lower outer angle of cell; 3 from the angle, nearly as
long as 2, connate with 4; all veins long; 4 and 5 con-
tiguous or weakly anastomosed for about half their
lengths from cell; 7 and 8 weakly anastomosed beyond
cell, their free elements long; cell slightly less than one-
fourth the length of wing; discocellular vein curved for
a short distance from lower angle, thence vertical.
Kighth abdominal segment of male with sternite de-
veloped as a narrow, sclerotized pocket.
Male genitalia similar to those of typical Davara
except: Free spine associated with anellus entirely
lacking; harpe with two-pronged clasper and apical end
of sacculus produced; penis with a pair of curved, flat-
tened, bladelike cornuti. Female genitalia with ductus
bursae sclerotized for a short distance from genitalia
opening; a single, angulate, projecting plate behind
genital opening; otherwise as in Davara.
157. Sarasota plumigerella Hulst
FIGURE 258
Sarasota plumigerella Hulst, Journ. New York Ent. Soc., vol. .8,
p. 222, 1900.—Grossbeck, Bull. Amer. Mus. Nat. Hist.,
vol. 37, p. 128, 1917.—McDunnough, Check list, No. 6120,
1939.
Basal segment of antenna red spotted with black.
Prothorax wine red; mesothorax and metathorax black.
Forewing black at base followed by a whitish ocherous
shade; antemedial line oblique, straight, ocherous
white, preceded by more blackish scaling, followed (es-
pecially on costa) by a strong black shade; remainder
of wing blackish fuscous stained with wine red, the
costa on outer half distinctly reddish; subterminal line
very faint, sinuate; discal spots obscure or absent;
terminal dots more or less confluent, blackish; cilia red.
Hind wing dull, translucent white; a dark shade at
apex and a narrow dark line along termen. Midtibia
with a strong yellow hair tuft. Alar expanse, 11-14
mm.
Male genitalia with terminal margin of vinculum not
produced at the edges but exhibiting no other specific
characters. Female genitalia slightly smaller but other-
wise not distinct from those of furculella.
Typrr Locauity: Palm Beach, Fla. (type in USNM).
Foop puiants: Laguncularia racemosa, Coccolobis
uvifera (larvae feeding under a light silk webbing on
the leaves and flower buds).
Distrrisution: Florida, Palm Beach, Ramrod Key
(Apr.), Stock Isl. (Apr.), Sugar Loaf Key (Apr.).
158. Sarasota furculella (Dyar), new combination
Ficures 48, 259, 740
Cuba furculella Dyar, Ins. Insc. Menstr., vol. 7, p. 50, 1919.
Larger and paler than plumigerella. Thorax and
forewing violaceous gray more or less tinted with red-
dish; the blackish shade of plumigerella replaced in
Surculella by red; antemedial line dull white with a
strong ocherous tint, especially towards inner margin,
its dark outer border red; some white dusting in median
costal area; discal dots more distinct, separate, red or
fuscous; subterminal line more distinct, narrow, sinuate,
dull white; terminal dots few, but distinct and separated,
blackish.’ Hind wings white with a faint smoky tint;
the veins very slightly darkened. Midtibial hair tuft
as in plumigerella. Alar expanse, 14-16 mm.
Male genitalia with terminal margin of vinculum
slightly produced at the sides; otherwise as in plumi-
gerella. Female genitalia a trifle larger than those of
plumigerella but showing no specific characters.
TyprE LocaLity: Santiago, Cuba (type in USNM).
Foop puant: Unknown.
Distrisution: Cusa: Baracoa, Matanzas, Santiago
(June). Purrtro Rico: Dorado (May), Puerto Real
(Vieques Isl., Apr.). Dominica (Dec.). Viren Is-
LANDS: Kingshill (St. Croix, June, Dec.).
Very close to plumigerella but apparently a distinct
species, separable chiefly on color.
159. Sarasota ptyonopoda (Hampson), new combination
Phycita ptyonopoda Hampson, Ann. Mag. Nat. Hist., ser. 6, vol.
16, p. 347, 1895.
Hyalospila ptyonopoda (Hampson) Hampson, in Ragonot, Mono-
graph, pt. 2, p. 530, pl. 56, fig. 9, 1901.
I have seen no examples of this species. From the
description and the figure in the Ragonot Monograph
the coloration and markings must be similar to those of
jurculella. According to Hampson the male has a
strong, long tuft of scales from the hind femur. Clarke
has reexamined the type and tells me that there is a
large expanded dorsal tuft from the base of the hind
femur and that there is no tuft on the midtibia. These
differences in tufting distinguish the species easily from
either plumigerella or furculella. A fine photograph of
the male genitalia, taken by Clarke, is before me.
They are like those of furculella Dyar. Unfortunately
the abdomen had been glued on the type so there will
remain some question of placement for the species until
another male from the type locality and with tufted
hind femora is recorded. Alar expanse, 16 mm.
Typr nocauity: St. Vincent, Windward Islands,
British West Indies (type in BM).
Foop puant: Unknown.
In the U. S. National Museum is a rubbed male from
Jalapa, México, labeled in Hampson’s handwriting
“Phycita ptyonopoda Hampson,” but it cannot be that
species. What remain of the legs show a yellow hair
tuft on midtibia and no trace of tuft on the hind femur
(the hind tibiae are missing). This specimen is an
Atheloca sp., close to bondari.
39. Genus Piesmopoda Zeller
Piesmopoda Zeller, Isis von Oken, 1848, p. 863.—Hulst, Phy-
citidae of N. Amer., p. 132, 1890.—Ragonot, Monograph,
pt. 1, p. 158, 18938. (Type of genus: Pzesmopoda rubicun-
della Zeller.)
78 UNITED STATES NATIONAL MUSEUM, BULLETIN. 207
Discopalpia Ragonot, Monograph, pt. 1, p. 167, 1893.
genus: Myelots flavicans Zeller. New synonymy.)
Amphycitopsis Dyar, Ins. Inse.’ Menstr., vol. 7, p. 45, 1919.
(Type of genus: Amphycitopsis isabella Dyar. New syn-
onymy.)
Tongue well developed. Antenna pubescent or
shortly ciliate (the cilia no longer than width of shaft) ;
on male, shaft simple, notched at base or with curved
excavation (a long sinus) towards base; basal segment
of male sometimes swollen but not notched or otherwise
modified. Labial palpus upturned, of varying length,
not reaching vertex (isabella, fratella) or extending to or
above vertex; third segment acuminate or (ragonott)
broadly dilated with scales. Maxillary palpus minute
(folded over tongue), filiform or subsquamous (third
segment slightly dilated with scales). Forewing smooth;
venation as in Sarasota except vein 8 normally consider-
ably closer to 4 at base than to 2; male without costal
fold. Hind wing as in Sarasota. Highth abdominal
segment ‘with sternite developed as a narrow, sclero-
tized pocket.
Male genitalia with uncus bifid. Gnathos incom-
plete, represented by its broad lateral arms, separated
at their apices, between which lies a rather well sclero-
tized subanal’ plate. Transtilla well developed but
incomplete, consisting of a pair of long, very slender,
curved arms whose sharply pointed apices are directed
away from each other. Harpe with @ strong, hooked
clasper from near apex of sacculus; a strong hair tuft
from lobe near base of sacculus. Anellus a curved plate
(U- or V-shaped) with long, slender, haired and bluntly
pointed lateral arms. Penis armed with a narrow,
flattened and more or less curved, bladelike cornutus;
sometimes with two such cornuti.
Female genitalia with single signum or none, signum
when present developed as a small granulate depres-
sion; ductus bursae with genital opening simple or more
or less sclerotized, but without the paired cuplike plates
found in Davara; ductus seminalis from bursa near its
junction with ductus bursae. EHighth-segment collar
more or less invaginate (except in apocerastes and
montella) .
A distinct genus defined and easily recognized by its
male and female genitalia. As used by Ragonot and
subsequent authors the name covered a composite of
disparate elements. The antennal and palpal char-
acters upon which Presmopoda, Discopalpia, and Am-
phycitopsis were distinguished are in this instance of no
more ‘than specific significance. They are discussed
more fully under their type species.
The genus is apparently limited to tropical America.
Nothing is known of the life history or host association
of any of the species.
(Type of
160. Piesmopoda rubicundella Zeller
Figure 260
Piesmopoda rubicundella Zeller, Isis von Oken, 1848, p. 864.—
Ragonot,; Monograph, pt. 1, p. 160, 1893.
Antenna of male with basal segment swollen, triangu-
late; shaft notched at base. Labial palpus slender,
third segment acuminate, reaching well above vertex.
Forewing pale (olivaceous ocherous) at base and for
some distance along inner margin; the median costal
area broadly white, heavily dusted with red scaling;
more or less of this red dusting also on costal half at
base; antemedial line not distinguishable; subterminal
line distinct, straight, oblique, pale reddish or ocherous
gray bordered by dark purplish fuscous lines; the entire
outer fourth of wing more or less suffused with purplish
fuscous; discal dots more or less distinct, separated, red
or, reddish fuscous. Hind wing pale smoky fuscous,
lighter towards base and semitransparent; veins dis-
tinctly darkened. Alar expanse, 13 mm.
Male genitalia having harpe with cucullus narrowly
elongate, evenly tapering. Anellus a broadly sclero-
tized, V-shaped band.
TyPE Locanity: Brazil (type in Mus. Univ. Berlin).
Foop piranr: Unknown.
I have examined the male type but have seen no
other examples from Brazil. The type is not distin-
guishable from males of the following species, (canthom-
era) except for minor differences in male. genitalia,
which may or may not be significant.
161. Piesmopoda xanthomera Dyar
Ficures 266, 745
Piesmopoda zanthomera Dyar, Proc. U. S. Nat. Mus., vol. 47;
p. 332, 1914.
Piesmopoda xanthozona Dyar, Ins. Insc. Menstr., vol. 7, p. 45,
1919 (new synonymy).
With the same male characters as rubicundella and
superficially not distinguishable from it. I suspect that
zanthomera is nothing more than a synonym or variety
of rubicundella, but this cannot be proven until females
of the latter from Brazil are available. Dyar’s two
names apply only to the sexes (canthomera to the females
and zanthozona to the males). Alar expanse, 13-17 mm.
Male genitalia, figured from type of zanthozona, differ
from those of rubicundella chiefly in the stronger tufting
from the base of the harpe, a character of very doubtful
specific value.’ Female genitalia, figured from type of
zanthomera and checked with those of females from all
localities here cited, distinguished chiefly by the shape
of the eighth-segment collar and its sclerotized invagi-
nate portion and the pair of corrugate patches on the
ventral surface of the latter. Bursa with signum:
Genital opening simple. MIO
TYPE LocAuitTiEes: La Chorrera, Panama (anthomera,
in USNM); St. Jean Maroni, French Guiana (xantho-
zona, in USNM).
Foop prant: Unknown.
DistRiBuTION: GUATEMALA: Cayuga (Apr., May).
Costa Rica: San José (July). Panam: La Chorrera
(May), Porto Bello (Oct.).. France Guiana: Cayenne,
St. Jean Maroni.
162. Piesmopoda trichomata (Zeller)
Figure 744
Myelois trichomata Zeller, Horae Soc. Ent. Rossicae, vol. 16, p.
194, 1881. :
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 79
Ragonot (Monograph, pt. 1, p. 160) makes this a
synonym of rubicundella, considering it merely the fe-
male of the latter. This is a very dubious placement.
The Zeller type material in the British Museum con-
sists of two female cotypes (photographs of which are
before me) alike in all details and similar to rubicundella
except that the antemedial line is indicated by broken
remnants of its outer border, a dark dash from costa,
and a more or less diffused dark spot on lower margin of
cell. The lower half of wing is somewhat suffused into
dark shading and contrasted against the whitish mid-
costal area, as in rubicundella. Zeller’s figure is mis-
leading in that it shows much of the wing bright yellow,
asin floricans. Alar expanse, 13 mm.
The genitalia of the female cotype here figured, ac-
cording to Tams and Clarke, agree in all details with
those of the other cotype. The latter should be con-
sidered the holotype, as it is the better preserved speci-
men. Bursa with signum. Genital opening simple.
The narrow eighth-segment collar easily identifies the
species. It is unlike any other in the genus that I have
seen.
Typsr Locautity: Honda, Colombia (type in BM).
Foop piant: Unknown.
Known only from the type specimens.
163. Piesmopoda flavicans (Zeller)
Fiaures 262, 746
Myelois flavicans Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 193,
Discopalpia flavicans (Zeller) Ragonot, Monograph, pt. 1, p. 167,
1893 (in part, 2).
Piesmopoda flavicans (Zeller) Dyar, Proc. U. 8. Nat. Mus., vol.
47, p. 333, 1914.
Piesmopoda fratella Dyar, Ins. Insc. Menstr., vol. 7, p. 45, 1919
(new synonymy).
Antenna of male simple. Labial palpus slender; on
male not reaching vertex; on female extending slightly
above vertex (as in isabella).
Forewing bright yellow; outer third purple dusted
with blackish towards apex; the yellow ground color ex-
tended further outward on costa and inner margin than
at middle; antemedial line obsolete; subterminal line
faint, narrow, whitish, straight, oblique and close to
outer margin. Hind wing whitish, stained with smoky
fuscous towards apex and on the outer parts of the veins.
Alar expanse, 14-15 mm.
Male genitalia like those of zanthopolys except that
elements of transtilla are stouter, lateral arms of anellus
are bent sharply away from each other, and terminal
margin of vinculum is acutely rounded (rather than
straight). Female genitalia similar to those of rantho-
polys except for slight differences in the structure of
eighth-segment collar.
Type tocauities: Honda, Colombia (flavicans, in
BM); St. Jean Maroni, French Guiana (fratella, in
USNM).
Foop prant: Unknown.
DistrisuTion: Frencu Guiana: Cayenne, St. Jean
Maroni (Mar.). Cotomspra: Honda.
Specimens of five different species in the U. S. Na-
tional Collection had been identified by Hampson,
Dyar, and Schaus as flavicans. Among them was one
female from French Guiana which Hampson identified
(correctly) as flavicans and which Dyar later included in
his series of fratella. Clarke and Tams have checked
our genitalic figures of the several Piesmopoda species
with the female type of flavicans in the British Museum
and inform me that the genitalia of fratella agree in all
details with those of favicans. Ragonot also misidenti-
fied Zeller’s species (at least as far as males are con-
cerned) and on the strength of their peculiar male palpi
erected the genus Discopalpia, with flavicans as type.
Dyar (1914) noted the misidentification and renamed
the males of flavicans Ragonot (not Zeller) as Discopal-
pia ragonoti. Later evidence from genitalia justifies
the new specific name; but nomenclatorily the type of
Discopalpia must remain flavicans Zeller, and Dyar’s
citation of a new type (flavicans Ragonot=ragonoti
Dyar) is not justified, despite Ragonot’s misidentifica-
tion and the characters derived therefrom for his genus.
Fortunately these characters (as far as Piesmopoda are
concerned) are of specific significance only, so Disco-
palpia would fall, however we interpreted its type;
but I for one hold that when a specific name is cited as
type of a new genus, that species remains the type of
the genus regardless of what specimens were before the
author of the genus or how he described his generic
concept. The decision is one of nomenclature and
not of zoology.
164. Piesmopoda ragonoti (Dyar), new combination
Figures 265, 747
Discopalpia ragonoti Dyar, Ins. Insc. Menstr., vol. 7, p. 44, 1919.
Discopalpia flavicans Ragonot (in part, co’; not Zeller), Mono-
graph, pt. 1, p. 167, 1893.
Antenna of male simple; basal segment cylindrical;
shaft without notch or other modification. Labial
palpus very long, the second segment reaching well
above vertex in both sexes; on male the third segment
broadly expanded with long scales, fan shaped; a strong
admixture of black scaling on outer sides of the palpi of
both sexes.
Forewing as in flavicans. Alar expanse, 14-16 mm.
Male genitalia distinguished chiefly by the very
heavy, black, broad-haired tuft from base of sacculus of
harpe. Female genitalia with signum; genital opening
simple; distinguished from those of other yellow-winged
species by minor differences in the configuration of the
eighth-segment collar; closest to those of zanthopolis
Dyar.
TYPE LOCALITY:
USNM).
Foop prant: Unknown.
Disrrisution: México: Distrito Federal. Guatr-
mata: Cayuga (Feb., Apr., May). Costa Rica: Juan
Vifias (Jan.).
Cayuga, Guatemala (type in
80 UNITED STATES NATIONAL MUSEUM BULLETIN 207
165. Piesmopoda isabella (Dyar), new combination
Figures 264, 749
Amphycitopsis isabella Dyar, Ins. Inse. Menstr., vol. 7, p. 45,
1919
Male antenna simple. Labial palpus short, hardly
reaching vertex on male and but a trifle beyond vertex
on female.
Forewing as in flavicans. Alar expanse, 18-20 mm.
Male genitalia with cucullus of harpe subtriangulate,
harpe wider in proportion to its length and less evenly
tapering than in other species except apocerastes which
has similar male genitalia. Female genitalia with
signum; genital opening simple; distinguished by the
conficuration of the eighth-segment collar.
TYPE LocALITy: Juan Vifias, Costa Rica (type in
USNM).
Foop prant: Unknown.
Represented by the male type and a slightly larger
female from the type locality (Jan.). The latter had
been identified by Schaus as “Piesmopoda flavicans
Zeller.”’ The species served as type for Dyar’s genus
Amphycitopsis, erected solely on the basis of the short
labial palpi of the male.
166. Piesmopoda xanthopolys Dyar
Figures 261, 748
Piesmopoda xanthopolys Dyar, Proc. U. S. Nat. Mus., vol. 47, p.
332, 1914
Male antenna simple. Labial palpi slender, extend-
ing above vertex in both sexes, somewhat longer on
female than on male.
Forewing as in flavicans. On the female from the
type locality the purplish shading on the outer area of
the forewing is somewhat more extended and the yellow
area of the wing proportionally more restricted than in
flavicans; but this difference seems to be individual
rather than specific. In zanthopolis as well as the other
species with coloration similar to flavicans the extent of
purple shading is variable between the sexes and even
among individuals of one sex. Alar expanse, 13-16 mm.
Male genitalia figured from specimen from La Chor-
rera, Panamd. Their most obvious feature seems to
be the rather short arms of the bifid uncus (proportion-
ally shorter than those of any other Piesmopoda except
flavicans which has stouter transtilla and differently
shaped anellus and vinculum). Female genitalia with
signum present. Genital opening simple. Very close
to those of flavicans, with incurvation of posterior-
ventral margin of eighth-segment collar similar and ex-
hibiting only minor differences in the collar otherwise.
TyPE Locauity: Porto Bello, Panam& (type in
USNM).
Foop prant: Unknown.
Distrisution: PanamA: Corozal (July), La ae
rera (May), Porto Bello (Sept., Dec.).
Known only from the original type series. Dyar in
1919 (Ins. Insc. Menstr., vol. 7, p. 44) placed zantho-
polys in the synonymy of flavicans. The two species
are very close, but apparently distinct.
167. Piesmopoda parva, new species
Figures 263, 750
Male antenna simple except for a very slight incur-
vation of the shaft towards base. Labial palpus slender,
reaching vertex; terminal segment acuminate.
Forewing yellow with a slight olivaceous tint; costa
tather broadly margined from base to near apex with
white faintly peppered with red scaling; a few red and
black scales at extreme base and for a short distance
from base along inner margin; no antemedial line; sub-
terminal line straight, oblique, close to termen, with
narrow purplish fuscous borders and preceded by a fus-
cous shade which extends, triangularly, almost to the
cell. Hind wings translucent white with a faint smoky
tint, darkening slightly towards apex. Alar expanse,
10-11 mm.
Male genitalia distinguished by the slender arms of
the divided uncus and the slender, naillike spine asso-
ciated with anellus. Female genitalia with signum;
genital opening simple. Distinguished by the broad
and deep excurvation in posteroventral margin of the
eighth-segment collar.
Typr tocauity: La Chorrera, Panamé (type in
USNM, 61333).
Foop prant: Unknown.
Described from male type from the type locality
(May), one male paratype from Cabima, Panam4 (May
1911), and one female paratype from Taboga Isl.,
Panamé (Feb. 1912), all collected by A. Busck. These
specimens had been included by Dyar among his para-
types of Piesmopoda xanthomera. They are somewhat
rubbed but otherwise in good condition. The species
is the smallest of the Presmopoda.
168. Piesmopoda semirufella (Zeller)
Figure 752
Myelois semirufella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p.
196, 1881.
Piesmopoda semirufella (Zeller) Ragonot, Monograph, pt. 1, p.
160, 1893.—Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 332,
1914; Ins. Insc. Menstr., vol. 7, p. 46, 1919.
Several different species have been identified as semi-
rufella.. I have seen but one example that can be defi-
nitely placed to Zeller’s name, a female from Cayuga,
Guatemala, collected by Schaus and Barnes (Jan.).
The genitalia of this specimen have been checked by
Clarke with the genitalia of the type of semirufella and
he finds them identical. They have the signum present
and a broad, strongly sclerotized plate at genital open-
ing. The latter structure at once identifies the female.
Males of the species have not been properly associated.
Superficially, semirufella is not distinguishable from
females of apocerastes Dyar. Indeed, several females of
the latter in both the British Museum and U. S. Na-
tional Collections had been identified as Zeller’s species.
Alar expanse, 16 mm.
TypEr Locatiry: Colombia (type, 2, in BM).
Foop piant: Unknown.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAEB 81
169. Piesmopoda apocerastes Dyar
Fieure 751
Piesmopoda apocerastes Dyar, Ins. Insc. Menstr., vol. 7, p. 45,
Male antenna with basal segment cylindrical, slender;
shaft with a long sinus (involving about eight of the
basal segments) lined by flattened blackish scales which
terminate in a slight tuft at the outer extremity of the
sinus. Labial palpus cylindrical, slender, reaching to
slightly above vertex.
Forewing, except for a whitish border along costa,
suffused reddish brown to the naked eye, very slightly
darkened towards outer margin (under magnification
the ground color shows a strong under tinting of oliva-
ceous ocherous); whitish costal border peppered with
scattered red scaling, a concentration of these along ex-
treme costal margin; discal dots separate, red; ante-
medial line obsolete or, at most, faintly indicated on
some specimens by an obscure, narrow, dark, trans-
verse shade; subterminal line faint, weakly bordered by
narrow dark (reddish fuscous) lines. Hind wings smoky
white to pale smoky fuscous, darkening towards apex
and outer margin; the veins darkened. Alar expanse,
15-16 mm.
Male genitalia similar to those of isabella Dyar. Fe-
male genitalia without signum; a narrow, strongly scle-
rotized plate at genital opening with weakly sclerotized,
anterior, lobelike projection.
TYPE LOCALITY: Juan Vifas, Costa Rica (type in
USNM).
Foop puant: Unknown.
Distrisution: México: Jalapa. Costa Rica: Juan
Vifias (May, Nov.). Dominica (British West Indies,
Feb.). Frencno Guiana: St. Jean Maroni. Brazit:
Parané, Castro.
A distinct species easily identified by its female geni-
talia. Several of the females before me in the National
Collection had been identified by Hampson as Piesmo-
poda semirufella. The genitalia of female specimens
from all the above-mentioned localities have been
checked.
170. Piesmopoda montella Schaus
Fiaurs 743
Piesmopoda montella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol.
11, p. 247, 1913.
Labial palpus of female slender, reaching slightly
higher than vertex.
Forewing light olivaceous brown; costal margin to
subterminal line broadly white irrorated with red-
brown, the extreme costal edge black at base, reddish
brown at middle; no trace of any antemedial line; sub-
terminal line slightly outcurved below vein 6, bordered
inwardly and outwardly from costa to vein 2 by blackish
bands into which some reddish scales are intermixed;
discal dots separated, reddish brown; a few blackish
dots on terminal margin. Hind wing pale, semihyaline
brown, darkening towards outer margin; the veins
darkly outlined. Alar expanse, 24 mm.
Female genitalia without signum; bursa copulatrix
small, oblong; ductus bursae very short and broad, al-
most as broad at middle as the bursa, weakly sclerotized
at genital opening, finely sclerotized otherwise.
Kighth-segment collar simple, not fused ventrally.
Typp tocauity: Mount Pods, Costa Rica (May;
type in USNM).
Foop prant: Unknown.
A distinct species easily identified by its large size
and distinctive genitalia; represented only by the fe-
male type.
Genus 40: Atheloca
[Venational division C. Forewing with veins 4 and 5 connate
and contiguous or partially anastomosed for one-third their
lengths from cell; 6 from upper angle of cell, connate with the
stalk of 8-9, bent towards base. Hind wing with cell less than
one-fifth the length of wing; discocellular vein oblique, straight.
Male genitalia with uncus hoodlike; lateral arms of gnathos fus-
ing into anellus; harpe broadly angled at base of cucullus.]
40. Atheloca, new genus
Typr or Genus: Nephopteryz subrufella Hulst.
Tongue well developed. Antenna of male simple,
shaft pubescent. Labial palpus slender, upturned,
reaching to or a trifle above vertex; third segment
shorter than second, acuminate. Maxillary palpus fili-
form. Forewing smooth; 11 veins, vein 2 from before
lower outer angle of cell; 3 from the angle, much nearer
to 4 at base than to 2; 4 and 5 connate, contiguous or
partially anastomosed beyond base for one-third of
their lengths; 6 from upper angle of cell, curved towards
base and connate with the stalk of 8-9; 10 from the cell,
approximate to the stalk of 8-9; male without costal
fold. Hind wing with vein 2 from before lower outer
angle of cell; 3 from the angle, nearly as long as vein 2,
closely approximate to the stalk of 4-5 at base; 4 and
5 stalked for half their lengths; 7 and 8 anastomosed
for most of their lengths beyond cell, the free element
of vein 8 very short; cell less than one-fifth the length
of wing; discocellular vein oblique, straight. Abdomen
of male with a pair of invaginated hair tufts at base;
sternite of eighth segment developed as a sclerotized
digitate pocket.
Male genitalia with uncus hoodlike, triangulate,
densely haired on outer surface. Gnathos represented
only by its lateral arms which fuse into anellus at their
apices. Transtilla absent. Harpe broadly angled at
base of cucullus. Anellus a stout, triangulate plate
with strongly sclerotized, sharply out-curved, smooth
lateral arms (these latter may possibly represent ele-
ments of a divided transtilla fused with the anellus, but
this is extremely doubtful). Aedeagus and penis simple.
Vinculum longer than greatest width, but slightly taper-
ing to truncate terminal margin.
Female genitalia with signa present in the form of
two small scobinate patches; ductus bursae with a small,
weakly sclerotized collar near the junction with bursa
copulatrix; genital opening simple, ductus seminalis
from bursa near the junction of bursa and ductus bur-
sae; eighth-segment collar completely fused ventrally.
82 UNITED STATES NATIONAL MUSEUM BULLETIN 207
171. Atheloca subrufella (Hulst), new combination
Figures 267, 741
Nephopteryx subrufella Hulst, Ent. Amer., vol. 3, p. 182, 1887.
Nephopteryz filiolella Hulst, Ent. Amer., vol. 4, p. 117, 1888 (new
synonymy).
Piesmopoda subrufella (Hulst) Hulst, Phycitidae of N. Amer.,
p. 133, 1890.—Ragonot, Monograph, pt. 1, p. 166, 1893.—
Grossbeck, Bull. Amer. Mus. Nat: Hist., vol. 37, p. 129, 1917.
Piesmopoda filiolella (Hulst) Hulst, Phycitidae of N. Amer., p.
133, 1890,—Ragonot, Monograph, pt. 1, p. 166, 1893.
Sarasota subrufella (Hulst) Barnes and McDunnough, Check list
of the Lepidoptera of Boreal America, No. 5558, 1917.—
McDunnough, Check list, No. 6121, 1939.
Sarasota filiolella (Hulst) Barnes and McDunnough, Check List
of the Lepidoptera of Boreal America, No. 5559, 1917.—
McDunnough, Check list, No. 6122, 1939."
Hyalospila ptychis Dyar, Ins. Insc. Menstr., vol. 7, p. 49, 1919
(new synonymy).
Forewing ocherous fuscous shaded with reddish or
purplish red except along costa; costa at extreme base
edged with black, between the transverse lines rather
broadly bordered by dull white sprinkled with red
scales; antemedial line sometimes obsolete, when pres-
ent indicated by a transverse black band interfused with
reddish and preceded by a narrow dusting of white
scales; subterminal line faint, pale, bordered inwardly
and outwardly by narrow blackish or purplish red
bands; discal spots usually distinct (at least the lower
one), well separated, blackish (rarely with a touch of
red); a more or less distinct row of blackish dots along
termen. Hind wing smoky white, translucent; the
veins darkened and a distinct dark shade along termen.
Midtibia of male with strong hair tuft from base on
inner side. Hind tibia of male without appreciable
hair tuft. Alar expanse, 12-19 mm.
Male genitalia as given for the genus. Female geni-
talia with sclerotized portion of ductus bursae very nar-
row.
TypE Locauities: Florida (subrufella, nm AMNH, ex
Rutgers); “Texas” (filiolella, in AMNH, ex Rutgers);
Cuba. (piychis, in USNM).
Foop prant: Unknown.
Distrisution: Unrrep States: Florida, Chokolos-
kee, Christmas Harbor (Mar.), Duardia (May), Fort
Drum, Hastings (Aug., Sept.), Lake Alfred (May),
Paradise Key (Mar.), Royal Palm State Park (June,
Sept.), Stemper (May, July, Aug.), Vero Beach (Apr.),
Winter Park (July, Aug.). Cusa: Pinar del Rio, no
specific locality (type of ptychis). VIRGIN ISLANDS:
Kingshill (St. Croix, ““Nov.—Dec.’’).
The types of both subrufella and jfiliolella are females,
neither of which bears a locality label; subrufella bears
a label containing only the number ‘‘61”’; filzolella only
a date label “March.” The latter is without abdomen
and in very poor condition; but exhibits no difference
from the type of subrufella except its somewhat larger
size. In the National Museum there is a female from
Christmas Harbor (‘‘March’’) labeled by Hulst ‘“Ne-
phopteryx filiolella, type.”” It is an exact match for the
type in the Rutgers Collection, as are three other large
females from Florida in the National Collection, obvi-
ously the same as subrufella. We have a large series of
the species but no Texas examples, and I doubt very
much the correctness of Hulst’s citation as the type
locality of his filiolella. The species is obviously a
tropical one which has extended its range to Florida.
Dyar’s ptychis is merely a Cuban example, differing in
no wise from typical subrufella.
172. Atheloca bondari, new species
Hyalospila ptychis Bondar (not Dyar), Rev. de Ent., Brazil, vol.
11, p. 199, 1940.—Lepesme, Les insectes des Palmier’s, p.
343, 1947 (Paris).
Similar to subrufella except that the pale costal area
of forewing is less contrasted and conspicuous. The
genitalia male and female exhibit no essential differences
from those of subrufella. The one distinguishing char-
acter, seems to be a strong, dorsal, yellow hair tuft from
the base of the male hind tibia. This is lacking from
all specimens of subrufella and is an addition to the
similar tuft on male midtibia, present on both subrufella
and bondari. Alar expanse, 15-16 mm.
TypE Locauity: Baia, Brazil (type in USNM, 61335).
Foop PLant: Cocos nucifera (Lepesme also records
C. coronata, C. vagans, Attalea funifera, and A. piassa-
bossu).
Described from male type and three male and five
female paratypes all from the type locality and reared
(June and July 1939, under Bondar Nos. 2521 and 2561)
from larvae feeding in the seeds and at the base of the
fruits of Cocos nucifera. These were received from Dr.
Gregorio Bondar, for whom the species is named. He
gives a good account of the habits of the species in the
above-cited paper. J am responsible for the misidenti-
fication to ptychis, for at the time I overlooked the differ-
ence in leg tuftings between ptychis and the Brazilian
specimens.
Genera 41 and 42: Praedonula and Peadus
[Venational division D. Forewing with veins 4 and 5 closely
approximate for a short distance from cell; vein 6 straight; 10
from the cell. Hind wing with cell one-fourth to one-fifth the
length of wing; discocellular vein more or less curved. Male
antenna with a shallow sinus in shaft involving the first half
dozen segments. Male genitalia with a stout hair tuft from near
base of sacculus.]
41. Praedonula, new genus
Type or cenus: Phycita almonella Dyar.
Tongue well developed. Male antenna with a shallow
sinus in shaft at base (involving the first six segments),
the sinus containing a scattering of minute papillalike
setae and a narrow ridge of scales along its outer edges,
otherwise pubescent. Labial palpus very slender,
upturned, reaching to or slightly above vertex; third
segment slightly shorter than second, acuminate.
Maxillary palpus subsquamous (the scales on second
segment expanded, on third rather long and drawing to
® point). Forewing smooth; 11 veins; vein 2 from
before but near lower outer angle of cell (nearer the
angle in male than in female); vein 3 from the angle;
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 83
4 and 5 closely approximate at base and for a short
distance beyond; 6 straight, from slightly below upper
angle of cell (male) or from the angle (female); 10 from
the cell, closely approximate to the stalk of 8-9 (male)
or slightly separated from it (female); male without
costal fold. Hind wing with vein 2 from before lower
outer angle of cell; 3 from the angle, nearly as long as
2, approximate at base to the stalk of 4-5; 4-5 stalked
for at least half their lengths (female), somewhat longer
stalked on male; 7 and 8 contiguous or weakly anas-
tomosed for a short distance beyond cell, the free
element of 8 long; cell short, about one-fifth the length
of wing; discocellular vein oblique, very slightly curved.
Eighth abdominal segment with sternite developed as
a narrow (digitate) sclerotized pocket.
Male genitalia with uncus hoodlike, rounded. Gna-
thos strongly developed, the lateral arms broad, ex-
panded and curled at their extremities and supporting
a sclerotized subanal plate with a short thornlike spur
at its base. Transtilla absent. Harpe stout, simple,
slightly broadened at middle; at base of cucullus a
stout hair tuft. Anellus a heart-shaped plate with
stubby lateral lobes. Penis unarmed. Vinculum stout
(but sclerotized narrowly along its margins); slightly
longer than broad; scarcely tapering.
Female genitalia with signa present in the form of two
small granulate patches; ductus bursae with genital
opening surrounded by a sclerotized plate; ductus
seminalis from bursa near junction of bursa and ductus
bursae. Eighth-segment collar completely fused
ventrally.
This genus is obviously closely related to the preced-
ing genus (Aftheloca) and that which follows (Peadus).
From the former it differs in the male antennal character
and the weak anastomosis of veins 7-8 of hind wing.
From Peadus it differs chiefly in the simple (undivided)
harpe and the much more strongly developed uncus and
tegumen of its male genitalia. It has no close relation-
ship to the Old World Phycita to which Dyar referred
its type species.
173. Praedonula almonella (Dyar), new combination
Fiaures 47, 268, 742
Phycita almonella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 333,
1914.
The type series (a male and two females) are some-
what rubbed and the coloration and markings of fore-
wing consequently obscured. Superficially the species
resembles Atheloca subrufella; the ground color of fore-
wing a gray brown, darkening in outer area and shading
to ocherous brown along inner margin at base; the
costal margin broadly margined with white rather
heavily dusted with red scales; antemedial line not dis-
tinguishable; subterminal line faint, oblique and close
to outer margin; discal dots very faint, separated.
Hind wing translucent white shaded with fuscous at
apex; the veins slightly darkened; on underside of male
& coarse yellow sex-scaling between costa and cell, along
lower margin of cell, extending for a short distance along
veins 2 and 3, and along’ vein lb from base for nearly
half its length. Alar expanse, 12-14 mm.
Male genitalia with terminal margin of vinculum
evenly rounded; aedeagus slightly bulged at middle, a
row of short teeth: along lateral edge towards apex.
Female genitalia simple and membranous except for a
fine sprinkling of minute scobinations and the two
small signa.
Type nocauity: Porto Bello, Panam& (type in
USNM).
Foop piant: Unknown.
Distrisution: PanamA: La Chorrera (May), Porto
Bello (May).
Known only from Dyar’s original type series.
42, Peadus, new genus
Type or genus: Piesmopoda burdettella Schaus.
Tongue well developed. Male antenna with a shal-
low sinus in shaft at base (as in Praedonula except here
the hollow of the sinus is overlaid with rather coarse,
appressed scales and without any indication of a tuft
or lateral scale ridge), pubescent. Labial palpus slen-
der, upturned, reaching vertex; third segment shorter
than second, acuminate. Maxillary palpus subsqua-
mous (as in Praedonula). Forewing smooth; venation
as in Praedonula except vein 2 further from lower outer
angle of cell, and 6 from below upper angle. Hind wing
with cell one-fourth the length of wing; discocellular
slightly but evenly curved. Eighth abdominal seg-
ment with sclerotized pocket of sternite long and
needlelike and with a large, flattened, fanlike tuft of
long slender scales.
Male genitalia with uncus and tegumen greatly re-
duced; the uncus a narrow, weakly sclerotized angulate
band. Gnathos indistinguishable (burdetellus) or rep-
resented only by a very weakly sclerotized, transverse
band (dissitus). Transtilla absent. Harpe short, stout;
sacculus broad and broadly produced at apex; giving
the harpe a partially divided appearance; the free costal
half of harpe strongly recurved and bearing two very
stout spines, one on outer lower margin near the angle
produced by the projection of sacculus and another at
lower angle of cucullus; the cucullus itself narrow and
greatly reduced; a strong hair tuft from intersegment
adjacent to base of sacculus. Anellus a shallow, curved
plate with stubby lateral arms. Aedeagus spined at
apex; penis armed with a couple of more or less curved
and crinkled sclerotized bands and a cluster of short,
stout spines. Vinculum long, stout and evenly taper-
ing; considerably longer than greatest width.
Female genitalia with signum developed as a short,
slender thorn; area of bursa immediately surrounding
ductus seminalis scobinate and weakly sclerotized;
ductus bursae much shorter than bursa, broad and
more or less sclerotized; behind genital opening a pair
of narrow elongate plates extending backward and fus-
ing into the ventrally divided eighth-segment collar
(except in subaquilellus); ductus seminalis from bursa
near junction of bursa and ductus bursae.
84 UNITED STATES NATIONAL MUSEUM BULLETIN 207
The genus is quite distinct from any other on male
genitalia. Its closest relatives seem to be Praedonula
and Hyalospila. On many genitalic characters it re-
sembles the following genus (Gabinius), especially in
the development of uncus and harpe; but separates
from it on hind wing venation, especially the length
and position of vein 2.
174. Peadus burdettellus (Schaus), new combination
Figures 269, 754
Piesmopoda burdetiella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol.
11, p. 247, 1913.
Discopalpia semproniella Schaus, Ann. Mag. Nat. Hist., ser. 8,
vol. 11, p. 249, 1913.
Hyalospila burdettella (Schaus) Dyar, Ins. Insc. Menstr., vol. 7,
p. 48, 1919.
Forewing pale brown shaded with red and blackish
scales; the costal border white sparsely dusted with
red scales and with medial costal edge reddish; ante-
medial line obscure except on lower half, far out, oblique
from costa to cell and below cell inwardly concave,
white, bordered outwardly on costa by a faint reddish
streak; some black scaling along basal half of vein 1b;
a distinct black dot on lower vein of cell at middle, and
on the vein 1b on the outer edge of the antemedial line;
subterminal line sinuous, bordered inwardly by a nar-
row, dark brownish shade and followed in outer area
(especially towards apex) by some white dusting; discal
dots distinct, separated, black, the lower one somewhat
elongated; an irregular black line along terminal mar-
gin reaching almost to apex. Hind wings translucent,
smoky white, darkening outwardly; the veins slightly
darkened and a fine dark line along termen.. Alar ex-
panse, 19.5—20 mm.
Male genitalia with no trace of sclerotized gnathos;
aedeagus with apex bluntly pointed, bearing a line of
short, coarse spines along its edge. Female genitalia
with ductus bursae weakly sclerotized near genital open-
ing; sclerotized plates behind genital opening narrow,
bladelike; sclerotization of bursa near ductus seminalis
slight.
TYPE LocaLitizus: Mount Pods (Juan Vinas), Costa
Rica (burdettella, in USNM); Juan Vifias (semproniella,
in USNM).
Foop prant: Unknown.
Distrisution: Costa Rica: Juan Vifias (Jan.),
Mount Pods (May). GuatEemaua: Volcén Santa Maria
(July).
Dyar established the above synonymy of Schaus’
species. The genitalia of their male types are identical.
175. Peadus dissitus, new species
Figures 270, 755
Similar to burdetiellus in color and markings except
for a distinct whitish longitudinal shade through the
cell of forewing and a stronger accentuation of the black
scaling; a thin black streak from base along half the
lower fold and a similar, shorter black streak on it just
before the subterminal line; outer margin of antemedian
line indicated by strong black dots on upper and lower
veins of cell and on vein 1b; inner dark margin of sub-
terminal line broadened by black streak. Alar expanse,
20-22 mm.
Male genitalia with gnathos a weakly sclerotized
transverse band; aedeagus with an expanded, flangelike,
densely and finely spined apex. Other minor differ-
ences from bwrdetiella (especially in the shapes of
cucullus and the projecting part of sacculus of the harpe)
are shown in the figures. Female genitalia with ductus
bursae much shorter than in burdettella, strongly sclero-
tized; sclerotized plates behind genital opening broad-
ened at their bases; sclerotization of bursa near
ductus seminalis appreciably stronger.
Typn tocauiry: “S. E. Brazil’? [Parané?] (type in
BM;; paratypes in BM and USNM, 61334).
Foop PLANT: Unknown.
Described from male type and three male and three
female paratypes from the type locality, “EH. D. Jones,
1920—803”’.
176. Peadus subaquilellus (Ragonot), new combination
Figure 753
Hyalospila subaquilella Ragonot, Nouv. Gen., p. 11, 1888; susie
graph pt. 1, p. 170, 1893; pt. 2, p. 38, fig. Ds, 1901.
This species is known only from the female'type. “A
drawing of its genitalia made by Clarke is figured: I
have seen nothing to match Ragonot’s figure and de-
scription which indicate a form with dark reddish brown
forewing shaded somewhat with black at base and on
the costa’ but without other markings; hind wing
“transparent,” smoky, the veins and terminal margin
appreciably darkened. Alar expanse, 20 mm.
The generic placement is tentative, pending discovery
ofamale. The female genitalia are not typical, lacking
the sclerotized plates behind genital opening, but seem
to indicate a closer relationship to, Peadus than to any
other genus.
Type LocaLiry: “Cerro Zunil, ” Guatemala (type i in
BM).
Foop purant: Unknown.
Genus 43 : Gabinius
[Venational division B. Forewing with veins 4 and 5 connate;
2 from near lower outer angle of cell. Hind wing with 2 from
close to lower outer angle of cell; 3 from the stalk of 4—5; cell less
than one-third the length of wing. Uncus reduced, weakly
sclerotized, triangulate. Tegumen greatly reduced but with
strong, projecting lateral arms. Transtilla incomplete. Harpe
reduced; apex of costa spined. Vinculum short, stout.]
43. Gabinius, new genus
Typr or Genus: Promylea paulsoni Ragonot.
Tongue well developed. Antenna of male simple,
pubescent. Labial: palpus upturned, reaching above
vertex; second segment somewhat broadly scaled; third
segment shorter than second, bluntly acuminate. Max-
illary palpus squamous. Forewing smooth; 11 veins;
vein 2 from before but near lower outer angle of cell,
nearly as close to 3:at base as 3 is to 4;3 from the angle;
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 85
4 and 5 connate; 6 from below upper angle of cell,
straight; 8 and 9 long stalked, the free element of 9 very
short; 10 from the cell, approximate to the stalk of 8-9
for a short distance; male without costal fold. Hind
wing with vein 2 from close to lower angle of cell; 3
from the stalk of 4-5; 4 and5 very long stalked; 7 and 8
closely approximate beyond cell; cell less than one-third
the length of wing; discocellular vein curved. Highth
abdominal segment with sternite developed as a short
(stubby) sclerotized pocket with thin, curved lateral
arms extending into a slender U-shaped plate, fringed
with moderately long scales.
Male genitalia with uncus reduced, broader than long,
triangulate and weakly sclerotized. Tegumen support-
ing a pair of long, broad, pointed, strongly sclerotized,
backwardly projecting arms; otherwise greatly reduced.
Gnathos absent. ‘Transtilla incomplete; its elements
pointed towards each other, their apices expanded and
nearly touching. Harpe short, nearly as broad as long;
costa broadly sclerotized, but shorter than remainder of
harpe, bearing a stout, rather long, projecting spine at
apex; cucullus much reduced, bearing a similar, stout
but shorter spine on costal edge at apex; sacculus
broadly sclerotized, but not produced. Anellus a broad
crescentiform plate; its lateral lobes reduced to weakly
haired knobs. Aedeagus smooth, straight; penis armed
with a pair of curved, more or less wrinkled, sclerotized
plates. Vinculum stout, as broad as long; its terminal
margin evenly rounded.
Female genitalia with signum developed as a thin,
keellike blade on a narrow, elongate plate; ductus bursae
shorter than bursa and with a sclerotized collar near
simple genital opening; ductus seminalis from’ bursa
near junction of bursa and ductus bursae.
A distinct genus, apparently most closely related to
Peadus but falling into a different venational division
(B). The shape and structure of the uncus, the re-
duced tegumen, and the short harpe with its stout
projecting spines suggest the relationship to Peadus.
The very short cell of hind wing separates it from most
genera of division B. It is only remotely related to
Promylea, to which Ragonot referred its type species.
177. Gabinius paulsoni (Ragonot), new combination
Ficures 271, 756
Promylea paulsoni Ragonot, Nouv. Gen., p. 12, 1888; Mono-
graph, pt. 1, p. 208, 18938.
Ground color of forewing olivaceous gray strongly
tinted with vinous brown in the dorsal area, this latter
shade extending obliquely almost to apex; costal area
from base to subterminal line, including the cell and
tapering to costa beyond it, white with a scattered
peppering of red-brown scales; costal edge from ante-
medial line to above end of: cell edged with blackish
brown; antemedial line distinct on lower half of wing,
vertical with a slight inward concavity, followed out-
wardly by a narrow blackish brown band and inwardly
by a blotch of the same shade; the antemedial line
indistinct and oblique on upper area of wing; subtermi-
nal line, narrow, faint, sinuate, whitish, followed and
preceded for a short distance from costa by blackish
brown shadings; a scattering of white scaling in the
terminal area below apex; lower discal spot blackish
brown, more or less distinct, the upper very small and
faint; a blackish irregular line along terminal margin,
not reaching to apex. Hind wing translucent, yellowish
white with a smoky tint towards apex and along ter-
minal margin. Alar expanse, 20-23 mm.
Male genitalia with characters as given for the
genus.
TYPE Locauity: Quillota, Chile (type in Paris Mus.).
Foop puant: Unknown.
The female in the National Collection matches in
every detail Ragonot’s description and figure of the
male type (Monograph, pl. 10, fig. 6). It is labeled
“Chile, Silva.’ Superficially, the maculation and color
resemble those of the Honora species.
Genera 44-46: Ceracanthia to Drescoma
[Venational division B. Forewing with veins 4 and 5 slightly
separated at cell. Hind wing with cell less than half the length
of wing (sometimes very short); 4 and 5 strongly stalked. Eighth
sternite of male developed as a digitate pocket, sometimes the
latter flattened and distorted. Male genitalia with vinculum
very long, sclerotized only along its margins, arched dorsally
(like a bent hairpin), its terminal margin broad; harpe with
tufts on a projecting arm from base of sacculus.]
44, Genus Ceracanthia Ragonot
Ceracanthia Ragonot, Monograph, pt. 1, p. 230, 1893. (Type of
genus: Ceracanthia vepreculella Ragonot.)
Procandiopa Dyar, Ins. Inse. Menstr., vol. 7, p. 50, 1919. (Type
of genus: Procandiopa mamella Dyar. . New synonymy.)
Tongue well developed. Antenna pubescent; basal
segment elongate, cylindrical (longer and more heavily
scaled in male than female); shaft of male with an
elongate, shallow sinus at base, from middle of sinus a
short, sharp, sclerotized spine, a similar, shorter spine
at apical end of sinus. Labial palpus upturned, reach-
ing above vertex, cylindrical; third segment nearly as
long as second, acuminate. Maxillary palpus sub-
squamous (scaling of second segment somewhat ex-
panded); folded across base of tongue. Forewing
smooth; 11 veins; vein 2 from near lower outer angle
of cell; 3 from the angle, about equidistant from 2 and
4; 4 and 5 slightly separated at base, thence divergent;
6 from slightly below upper angle of cell, straight; 10
from the cell, approximate'to the stalk of 8-9 for a very
short distance from cell; male without costal fold
Hind wing with vein 2 from before, but near lower outer
angle of cell; 8 from the stalk of 4-5; 4 and 5 stalked for
half their lengths; 7 and 8 closely approximate for a
short distance from cell; cell slightly over one-third the
length of wing; discocellular vein curved. Eighth ab-
dominal segment with sternite developed as a strongly
sclerotized, digitate pocket.
Male genitalia with uncus hoodlike (not tapering).
Apical process of gnathos a simple, elongate, slender
hook. Transtilla absent.’ Harpe broad, stout, sacculus
86 UNITED STATES NATIONAL MUSEUM BULLETIN 207
partially divided towards apex and with a projecting
arm from its base supporting a heavy hair tuft. Anellus
a rather narrow band with short, stubby, lateral arms.
Aedeagus somewhat curved and with a slightly more
sclerotized bulge from outer third; penis with a few
minute scobinations or some fine sclerotized wrinkles,
otherwise unarmed. Vinculum elongate (considerably
longer than greatest width), U-shaped, sclerotized only
(and narrowly) along its margins, arched dorsally (like
a bent hairpin).
Female genitalia with signum; ductus bursae and
genital opening simple; ductus seminalis from bursa
near junction of bursa and ductus bursae.
This genus is uncomfortably close to the genus: fol-
lowing (Megarthria), from which it differs chiefly in
the closer approximation of vein 2 of hind wing to the
lower outer angle of cell, the decided stalking of 3 with
4-5, and the slightly longer cell. The male antennal
character on which Ragonot and Dyar erected their
genera is probably here (as in Megarthria) of specific
rather than generic significance.
178. Ceracanthia mamella (Dyar), new combination
Fieaures 13, 272, 278, 757
Procandiopa mamella Dyar, Ins. Insc. Menstr., vol. 7, p. 51, 1919.
Forewing yellowish white along costa; remainder of
wing a purplish shade with a scattered peppering of
blackish scales especially along base of median fold;
antemedial line obscure, far out on wing and outwardly
angled in cell, indicated chiefly by a discontinuous, nar-
tow, blackish outer bordering line; subterminal line
somewhat more distinct, smuate, yellowish white, fol-
lowed on costa by a short blackish dash and bordered
inwardly by a black line which expands into patch at
middle; discal dots: blackish; terminal dots confluent
forming a blackish line along the outer margin. Hind
wing pale smoky brown; the veins faintly darkened and
a distinctly smoky shade along termen. Alar expanse,
15-16 mm.
Male genitalia with ontee margin of harpe evenly
rounded. Female genitalia with signum a flat, some-
what granulate plate, bearing a short thorn near its
center.
Tyre tocauity: Rio Trinidad, Panama (type in
USNM).
Foop piuAnt: Unknown.
Distrisution: PanaMA: Rio Trinidad (Mar., May).
GuatEmaLa: Cayuga (May).
Represented only by the original type series, a male
and female from the type locality and a male and female
from Guatemala.
The sinus in the male antenna (fig. 278) appears
smoothly scaled to the naked eye but under magnifica-
tion shows several minute, erect, papillalike scales, simi-
lar to those on Megarthria peterseni but less conspicuous.
179. Ceracanthia vepreculella Ragonot
Figure 273
Ceracanthia vepreculella Ragonot, Monograph, pt. 1, p. 230, 1893.
_I have seen no examples of this species but from
Ragonot’s description and figure it appears to be similar
to mamella except that the general color is a more uni-
form yellowish gray without the contrastingly paler
costa and the purplish shade on the remainder of the
wing, characteristic of mamella. It is also a larger
species. Alar expanse, 23 mm:
Male genitalia with outer margin of harpe angulate.
Typrr Locatity: Loja, Ecuador (type in Paris Mus.).
Foop prant: Unknown.
45. Genus Megarthria Ragonot
Megarthria Ragonot, Monograph, pt. 1, p. 156, 1893.
genus: Myelois petersenz Zeller.)
Characters similar to those of Ceracanthia except:
Vein 2 of hind wing further removed from lower outer
angle of cell; the cell itself shorter, lessithan one-third
the length of wing; vein 3 connate with the stalk of 4-5
or contiguous with it for a short distance from the angle
of cell, but not from the stalk; eighth abdominal seg-
ment of male with sternite developed usually as a
laterally flattened, more or less bent, digitate pocket,
the supporting lateral arms of the sternite bearing a
pair of scale tufts.
Male genitalia with anellus an elongate shield; trans-
tilla absent (except in schaust).
Female genitalia: with or without signum; ductus
seminalis from bursa adjacent to junction of bursa and
ductus bursae.
The shaft of the male antenna has a rather broad
sinus at base (as in Ceracanthia) but is specifically vari-
able. The lower outer angle of cell in the hind wing is
as far out as in Ceracanthia, but the discocellular vein
curves inward more deeply, making the cell itself BPEREs
ciably shorter than that of Ceracanthia.
In the National Collection all the specimens aoae
the male type of cervicalis Dyar had been identified as
peterseni Zeller. They are strikingly similar in color
and maculation, but their structures show that four
species are present among the males and at least two
among the females, With our. present knowledge the
sexes cannot be associated with any certainty; so until
something 1 is known of their host association and the
species are reared it seems a safer procedure to anticipate
later synonymy and give separate names to the males
and females rather than to link the females nomencla-
torily to any of the male types.
(Type of
180. Megarthria peterseni (Zeller)
Fieures 14, 274, 279
Myelois petersenit Zeller, Horae' Soc. Ent. Rossicae, vol. 16, P.
198, 1881.
Megarthria petersent (Zeller) Ragonot, Monograph, pt. 1, p. 187, }
1893
Giéountl color of forewing white on costal half of wing,
olivaceous brown on lower half, this brown shade
extending upward to costa at extreme base and obliquely
upward from lower outer angle of cell to costa at apex;
an oblique blackish brown band: from: costa at about
one-third extends to the paler brown ground color on
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 87
lower half of wing, dividing the white costal area into
two strongly contrasted white patches; dark basal area
also blackish towards costa; a similar blackish shade
from apex extends obliquely inward towards lower outer
angle of cell; some blackish shading along the outer
veins and a narrowly elongate, blackish brown spot on
midcosta; subterminal line very faint, except at costa,
dull white, some faint whitish dusting in the outer
brown area just below apex; a faint peppering of red
scales on the white areas and more or less over the
blackish brown markings on costal half of wing; discal
dots blackish brown, separated and usually distinct; a
row of black dots along termen. Hind wing semitrans-
parent, whitish with a smoky tint towards apex and
along costa and termen; the veins faintly darkened and
a blackish line along terminal margin. Eighth abdomi-
nal tufts more or less swollen hairlike scales. Alar
expanse, 22-26 mm.
Male genitalia with transtilla absent; harpe with
sacculus partially divided (towards its apex); hair tufts
from extended basal arm from sacculus, yellow; penis
bearing a patch of fine scobinations; vinculum con-
stricted near middle.
Antenna of male (fig. 279) with a broad sinus in base
of shaft occupying a half-dozen fused segments; the
sinus with many minute, papillalike setae (or scales) on
its inner surface, but without hair or scale tuft; a small
but strongly sclerotized spine from lateral edge of smus
beyond its base and a similar small spine from apex
of the sinus.
Typn LocaLiry: Honda, Colombia (type, @, in BM).
Foop prant: Unknown.
DistRisuTIoN: GuatemMaLa: Voleén Santa Maria
(June, July). Cotomp1a: Honda (Apr.). Braziu:
Santa Catarina (July). Pert: Oconeque (Carabaya).
I have seen no specimens from Colombia; but the
male examples before me from Guatemala, Brazil, and
Peri agree in antennal characters with the type of
petersenti as described by Ragonot. Evidently the
species has a wide distribution in Central and South
America.
181. Megarthria squamifera, new species
Figures 275, 280
Color and markings as in petersent. Male genitalia
also similar except lateral arms of gnathos stouter and
vinculum less constricted. Digitate pocket of eighth
abdominal sternite not appreciably flattened. Male
antenna (fig. 280) with a broad, elongate sinus; from
one lateral edge of sinus a flat brush of long stiff hairs;
the opposing edge concave for most of its length, the
concavity ending in a sharp projecting point at each
end; inner surface of sinus smooth.
Tyrr tocatity: Mount Pods, Costa Rica (type in
USNM, 61336).
Foop pirant: Unknown.
Described from male type, collected by Schaus and
Barnes (May).
182. Megarthria frustrator, new species
Color markings and male genitalia similar to those of
petersent. Male antenna like that of squamifera. Dif-
fers from other males of the genus in having a narrow
ridge of rough, protruding scales along the costa of
forewing for more than half its length from base. Alar
expanse, 20 mm.
TyPE LocaLiry: Juan Viiias, Costa Rica (type in
USNM, 61337).
Foop piant: Unknown.
Described from male type, collected by Schaus and
Barnes (Feb.).
183. Megarthria schausi, new species
FicurE 276
Color and markings as in peterseni. Male genitalia
with vinculum not constricted ; apical process of gnathos
terminating in a weak, short spine. Transtilla present
developed as a square, sclerotized plate, pendant from
protruding lobes from the costobasal area of the harpes
and with thin short projecting arms from its lower
(anterior) corners. Hair tufts from projecting basal
arm of sacculus short, yellow. Male antenna like that
of squamifera. Alar expanse, 18 mm.
TypE Locauity: Juan Vifias, Costa Rica (type in
USNM, 61338).
Foop pirant: Unknown.
Described from male type, collected by Schaus (Jan.)
and named in memory of him. The species is easily
identified by its platelike transtilla. I have seen nothing
resembling this structure in any other male of the genus.
184, Megarthria cervicalis Dyar
Ficures 277, 281
Megarthria cervicalis Dyar, Ins. Insc. Menstr., vol. 7, p. 42, 1919.
Forewing as on petersent except that dark areas are
paler; less blackish brown and with more red scaling
on the dark antemedial band and midcostal spot, the
latter almost entirely reddish. Eighth abdominal tuft
small, consisting of broadly flattened and contorted
scales; digitate pocket from sternite of eighth segment
strongly bent and decidedly flattened. Alar expanse,
20 mm.
Male genitalia with vinculum not constricted, of even
width throughout and with terminal margin evenly
rounded; penis armed with a small, flat, bladelike cor-
nutus; sacculus of harpe not divided at apex; hair tuft
from projecting basal arm of sacculus, black.
Male antenna (fig. 281) with a short, shallow sinus
at base of shaft and with a very small scale tuft from
the base of the sinus (under the lower magnification
looks like a small triangulate spine).
Tyrr tocaLity: Taénamo, Cuba (Aug.; type in
USNM).
Foop piant: Unknown.
Known only from its male type. <A distinct species
easily identified by its male antennae and genitalia.
88 UNITED STATES NATIONAL MUSEUM BULLETIN 207
185. Megarthria alpha, new species
Ficure 760
Female. Color and markings as in peterseni males.
Alar expanse, 17-26 mm.
Genitalia with signum developed as a teardrop
shaped, finely granulate-scobinate, depressed patch:
ductus bursae shorter than bursa with a narrow scle-
rotized band near genital opening. LHighth-segment
collar with a thin apron projecting anteriorly (the
shape and size of this individually variable; identical
in no two specimens; compare figures 760a and 760b).
Typr LocaLity: Volcén Santa Maria, Guatemala
(type in USNM, 61339).
Foop prant: Unknown.
Described from type (Sept.) and three paratypes
from the type locality (Schaus and Barnes, collectors,
July, Oct.) and one paratype from each of the following
localities: Purulhé, Guatemala (Schaus and Barnes,
July); Quirigué, Guatemala (Schaus and Barnes, Apr.),
Jalapa, México (Schaus); Orizaba, México (R. Miiller,
Mar. 13); Mount Pods, Costa Rica (Schaus, May);
Juan Vinas, Costa Rica (Schaus and Barnes, Jan.); Rio
Trinidad, Panamé (A. Busck, Mar. 1912); Incachaca,
Cochabamba, Bolivia (J. Steinbach); Santa Catarina,
Brazil (F. Hoffmann, July 12, 1935). All females;
genitalic preparations made of all specimens.
I believe that alpha will eventually prove to be the
female of petersent, but have no convincing evidence at
this time that it is so.
186. Megarthria beta, new species
Figure 759
Superficially indistinguishable from alpha but with
quite different genitalia. Alar expanse, 17-23 mm.
Bursa without trace of signum; ductus bursae much
longer than bursa, unsclerotized at genital opening ex-
cept for a narrow, very weakly sclerotized band on
lower margin ; anterodorsal projection of eighth-segment
collar slight.
TYPE LOCALITY: Orizaba, México (type in USNM,
61340).
Foop pirant: Unknown.
Described from type (Schaus Collector, no data); one
paratype from Jalapa, México (Schaus, no data); three
paratypes from Cayuga, Guatemala (Schaus and
Barnes, Feb., May, Oct.); one paratype from San José,
Costa Rica (H. Schmidt); two paratypes from Porto
Bello, Panamé (A. Busck, May 1912); and one para-
type from Caparo, west-central Trinidad (F. Birch, no
data). All females; the paratypes from Trinidad and
Costa Rica in the Js anse Collection.
46. Genus Drescoma Dyar
Drescoma Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 328, 1914.
(Type of genus: Drescoma cyrdipsa Dyar.)
Tongue well developed.. Antenna pubescent; basal
segment normal (ot swollen or elongated); shaft of
male simple. Labial palpus upturned, reaching vertex;
moderately slender, the second segment somewhat
broadly scaled; third segment nearly as long as second,
acuminate. Maxillary palpus subsquamous (scaling
of second segment somewhat expanded); folded across
base of tongue. Forewing smooth; 11 veins; vein 2
from near lower, outer angle of cell; 3 from the angle
about equidistant from 2 and 4; 4 and 5 slightly sepa-
rated at base, thence divergent; 6 from below (but near)
upper angle of cell, straight; 8 and 9 long stalked; 10
from the cell approximate to the stalk of 8-9 for some
distance; male without costal fold but with a distinct
notch in costa beyond base. Hind wing with vein 2
from lower outer angle of cell; 3 apparently from the
stalk of 4-5, but actually contiguous or weakly anas-
tomosed with it for half its length; 4 and 5 stalked for
over half their lengths; 7-8 contiguous or partially
anastomosed beyond cell, the free parts of the veins
very long; cell very short, one-fifth the length of wing
or less; discocellular vein oblique; on male upper vein
of cell notched just beyond base and with some modi-
fied sex-scaling above and beyond the notch; also on
male a short fold on anal margin enclosing a hair pencil.
Kighth abdominal segment without tuft; sternite de-
veloped as a short sclerotized, digitate pocket.
Male genitalia as in Megarthria except: Transtilla
present and in the form of a wide, very shallow U;
harpe with an appressed, clasperlike projection near
apex of sacculus; vinculum terminating in a strongly
sclerotized, sinuate, transverse bar.
Female genitalia with signum developed as a long,
strong, curved hook; ductus bursae shorter than bursa;
genital opening simple; ductus seminalis from bursa
near junction of bursa and ductus bursae.
A distinct genus closely related to Ceracanthia and .
Megarthria and to Drescomopsis in group II; but easily
distinguished by the bent (notched) upper vein of cell
in the male hind wing, the very short hind wing cell
in both sexes, the notched forewing of the male, the
peculiarly developed transtilla and terminal margin of
vinculum.
187. Drescoma cyrdipsa Dyar
Figures 15, 282, 758
Drescoma cyrdipsa Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 328,
1914.
Color and markings as in Megarthria peterseni except
for darker hind wings, especially those of the females
which are dark smoky fuscous. The average size is
also consistently smaller.
Hind wing of male with notched projection of upper
vein of cell into the cell deep and wide; on underside of
wing a patch of semimetallic scales above the notch;
a black patch preceding it, and following it a line of
yellow and black scales along the vein.
Male genitalia with transtilla triangularly broadened
toward harpes; clasperlike projection from sacculus
rather broadly triangulate. Female genitalia with
bursa minutely granulate; coarser granulations in ductus
bursae at its junction with bursa. ‘The long, strong,
thornlike cornutus may be a specific character also,
but is more probably generic.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 89
Type tocauity: La Chorrera, Panamaé (type in
USNM).
Foop piant: Unknown.
Disrrisution: México: Chiapas (May). GuatE-
MALA: Cayuga (Jan., May, June, Aug.). Panam:
Cabima (May), Corozal (Feb., Nov.), La Chorrera
(Apr., May), Rio Trinidad (Mar., June), Tabernilla.
Frenca Guiana: St. Jean Maroni.
188. Drescoma cinilixa Dyar
Figure 283
Drescoma cinilixa Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 329,
1914.
Markings and color of forewing as in cyrdipsa except
dark ground color and dark costal markings paler,
more ocherous fuscous than brown. Underside of male
hind wing with a patch of black sex-scaling above the
notch in cell, orange-yellow sex-scaling on the upper
vein of cell preceding the notch, on several veins follow-
ing the notch, and in the median fold at base of wing;
the indentation of upper vein of cell also much shallower
than on cyrdipsa, a concavity rather than a strongly
triangular notch. Hind wing paler; dull smoky white,
darkening towards termen and apex. Alar expanse,
15-16 mm.
Male genitalia with transtilla narrowed towards
harpes; clasperlike projection from sacculus slender,
sharply pointed. Female unknown.
Typn tocauiry: La Chorrera, Panamé (type in
USNM).
Foop pirant: Unknown.
DistrisuTion: GuaTEMALA: Cayuga (May). Pa-
naMA: La Chorrera (May).
Distinguishable from cyrdipsa by its paler color, the
secondary sexual characters of the male hind wing, and
the differently shaped transtilla and clasper.
Genus 47. Monoptilota
[Venational division B. Forewing with veins 4 and 5 closely ap-
proximate for a short distance from cell. Hind wing with vein 2
from well before angle of cell; 3 from the angle; cell one-third the
length of wing. Male genitalia with uncus bifid, its divided ele-
ments widely separated; transtilla incomplete; apical process of
gnathos a broadly triangulate hook. Male antenna unipectinate
with sinus and scale tuft in base of shaft.]
47. Genus Monoptilota Hulst
Monoptilota Hulst, Canadian Ent., vol. 32, p. 13, 1900.—Forbes,
Cornell Mem. 68, p. 621, 1923. (Type of genus: Monoptilota
nubilella Hulst.)
Tongue well developed. Antenna of male (figs.
285g—-h) with basal segment elongate, shaft unipectinate
and with a long shallow sinus at base containing a stout,
appressed scale tuft; of female simple, smooth. Labial
palpus obliquely upturned, reaching to vertex; second
segment broadly scaled; third segment small, acumi-
nate. Maxillary palpussquamous. Forewing smooth;
11 veins; vein 2 from well before lower outer angle of
cell; 3 from the angle, much closer at base to 4 than to
3003293—56——7
2; 4-5 closely approximate for a short distance from
cell; 6 from below upper angle of cell, straight; 8 and 9
stalked for less than half their lengths; 10 from the cell;
male without costal fold. Hind wing with vein 2 from
well before lower outer angle of cell; 3 from the angle,
connate with or very closely approximate at base to
stalk of 4-5; 4 and 5 stalked for about half their lengths;
7 and 8 closely approximate for a short distance from
cell; cell one-third the length of wing; discocellular vein
curved. Eighth abdominal segment of male with a
pair of fine, weak, hair tufts.
Male genitalia with uncus bifid, the divided parts
widely separated and hooked at their apices. Gnathos
terminating in a triangulate, sharply hooked central
process, its supporting lateral arms strongly arched and
arising well down from ventrolateral projections of
tegumen; an elaborate well-sclerotized subanal plate
attached to alimentary tube. Transtilla incomplete,
consisting of two, widely spaced, slender, weakly sclero-
tized plates. Anellus a simple, moderately broad, par-
tially curved band. Aedeagus short, stout; penis un-
armed. Vinculum short (shorter than broad) ; terminal
margin truncate.
Female genitalia with signum developed as a spined
plate (individually variable); bursa otherwise smooth;
ductus bursae rather stout, expanding gradually into
the bursa, smooth except for a broad sclerotized banding
near genital opening; ductus seminalis from bursa near
junction of bursa and ductus bursae.
A unique genus of doubtful affinities.
the one North American species.
Contains but
189. Monoptilota pergratialis (Hulst)
Ficurss 27, 285, 763, 764
Nephopteryx pergratialis Hulst, Trans. Amer. Ent. Soc., vol. 13,
p. 162, 1886; Phycitidae of N. Amer., p. 143, 1890.—Rago-
not, Monograph, pt. 1, p. 267, 1893.
Nephopteryx grotella Ragonot, N. Amer. Phycitidae, p. 6, 1887.
Monoptilota nubilella Hulst, Canadian Ent., vol. 32, p. 14, 1900.—
Chittenden, U. S. Dep. Agr. Div. Ent. Bull. 23, pp. 9-17,
1900.—Welden, Jour. Econ. Ent., vol. 1, p. 148, 1908.—
Forbes, Cornell Mem. 68, p. 621, 1923.
Monoptilota pergratialis (Hulst) Barnes and McDunnough, Con-
tributions, vol. 3, p. 195, 1916.—Brannon, Journ. Econ.
Ent., vol. 27, p. 719, 1984.—Brimley, Insects of North Caro-
lina, p. 300, 1988.—McDunnough, Check list, No. 6141,
1939.
Forewing dark fuscous (gray-brown) with blackish
shading on many of the veins and on costal half of wing;
along terminal margin a heavy dusting of white between
the veins, giving much of wing an ashy appearance;
antemedial line obscure, indicated chiefly by a diffused
blackish brown outer border, broadest and strongest
from costa to lower margin of cell, frequently inter-
rupted or obscured in the ground color towards inner
margin; subterminal line sinuate, grayish white with
dentate blackish brown inner and outer borders; discal
dots distinct, separated, blackish; a black line along
terminal margin more or less interrupted by white
streaklets at the vein ends. Hind wing of male semi-
hyaline white with veins slightly darkened and a black-
ish brown line along terminal margin; of female much
90 UNITED STATES NATIONAL MUSEUM BULLETIN 207
darker, dull smoky white to brown, with veins and ter-
minal edge correspondingly darker. Alar expanse,
17-30 mm.
Male genitalia with characters as given for the genus.
Female genitalia with signum extremely variable, rang-
ing from a small, weakly spined plate, like that of the
paratype (fig. 764), to a large plate with rather long
slender spines (fig. 763). In one large female of a reared
series from Virginia the signum is completely absent.
This variability in female structure is not matched by
anything in the male, where the genitalia are remark-
ably uniform for large and small specimens alike.
Typr Locatity: Florida (pergratialis, 9, m AMNH,
ex Rutgers; grotella, 9, in Paris Mus.); Auburn, Ala.
(nubilella, &', in USNM).
Foop PLANT: Limabean (larva a borer in the stems).
Distrisution: Maryland, Cabin John Bridge (Aug.,
Sept.), Salisbury (Sept.); Virginia, Norfolk (May), St.
Elmo (Jan., Feb., Mar.); North Carolina, Vance County
(Aug., Oct.); South Carolina, Florence (June, July),
Lyna Plantation (June); Georgia, Savannah (June);
Florida, Coconut Grove (Apr., May), Miami (Apr.);
Alabama, Auburn (July), Montgomery (July); Arizona,
Baboquivari Mts. (June, Aug., Sept.), Huachuca Mts.
(uly, Aug.), Nogales (uly), Palmerlee, Washington
Mts., White Mts. (June).
The species is of some importance as an enemy of
limabeans and is known in economic literature as the
‘dimabeam vine borer.”’ The Chittenden (1900) paper
cited gives what is known of the life history. There are
several later references in publications devoted to eco-
nomic entomology but they add nothing to our knowl-
edge of the insect. It is not known outside of the
United States; at least no specimens have been received
or identified from any of the tropical American regions
where the species might be expected to occur; and in
the United States its distribution seems to be limited
to the eastern area from the District of Columbia south
to Florida and adjacent Gulf States and to southern
Arizona. The only known host is the limabean. The
southern Arizona distribution raises a question as to
another possible host (probably a wild legume), for the
Arizona, localities are mostly out of the range of lima-
bean cultivation.
Genera 48-50: Zamagiria to Magariopsis
[Venational division D. Forewing with veins 4 and 5 connate,
shortly stalked or closely approximate for a short distance from
base; 8 close to 4-5 at base; 6 bent towards base connate with or
shortly separated from the stalk of 8-9 at base. Hind wing with
4 and 5 anastomosed from just beyond angle of cell for about
half their lengths; cell short. Eighth abdominal segment of
male with compound, ventral tufts. Labial palpi of male up-
curved; appressed to face or to each other; third segment greatly
reduced, acuminate. Gnathos with apical process broadly pro-
duced and lateral arms elongate and arising from ventrolateral
projection from tegumen. Female with ductus bursae short and
5 sclerotized towards genital opening and junction with
bursa.
48. Genus Zamagiria Dyar
Zamagiria Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 329, 1914.
(Type of genus: Zamagiria dixolophella Dyar.)
Tongue well developed. Antenna of male shortly
ciliate, a deep sinus containing heavy scale tuft at base
of shaft; of female simple and weakly pubescent. Male
head between the antennae deeply grooved to hold an
appressed, matted tuft of long scales arising from the
upper edge of frons. Labial palpus recurved-ascending;
second segment very long; broadly dilated and hol-
lowed within to hold the maxillary palpus; third seg-
ment short, acute. In repose the labial palpi are
appressed to each other and fit into the groove on head
covering both the maxillary palpi and the scale tuft
from frons. Maxillary palpus of male in the form of a
large aigrette; of the female minute, filiform. Forewing
with a ridge of roughened scales preceding the ante-
medial line but not reaching to costa; 11 veins; vein 2
from before the lower outer angle of cell; 3 from the
angle, close to 4—5 at base; 4 and 5 very closely approxi-
mate for a short distance from cell; 6 from upper angle
of cell, bent towards base, connate with 8-9; 8 and 9
long stalked; 10 from the cell, approximate to the stalk
of 8-9 for a short distance from cell; male without costal
fold but rough scaled on underside of costa at base.
Hind wing with vein 2 from well before the angle of the
cell; 3 from the angle; 4 and 5 anastomosed from just
beyond the angle for about half their lengths; 7 and 8
contiguous or weakly anastomosed for a short distance
beyond cell; cell less than one-fourth the length of wing;
discocellular vein curved. Highth abdominal segment
of male with compound ventral scale tufts.
Male genitalia with uncus triangulate. Gnathos with
apical process broadly produced and variously modified ;
lateral arms elongate and arising well down from ventro-
lateral projections of tegumen. Transtilla absent.
Harpe with sacculus more or less produced at apex;
clasper in some form usually present; cucullus narrow
and reduced. Aedeagus stout; penis armed with strong
spine or spines (except in hospitabilis).
Female genitalia with bursa armed with strong spine
cluster or clusters and usually partially sclerotized;
ductus bursae short, strongly sclerotized towards genital
opening and junction of bursa and ductus bursa (the
sclerotizations more or less contorted) ; genital opening
broad; ductus seminalis from bursa towards junction
of bursa and ductus bursae.
An easily recognized, compact genus exhibiting strik-
ing structural specific differences in genitalia.
190. Zamagiria dixolophella Dyar
Figure 286
Zamagiria dixolophella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p.
329, 1914.
Aigrette of male maxillary palpus reddish.
Forewing dark smoky gray, the basal area below costa
a trifle paler; antemedial line faint, narrow, oblique and
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 91
sinuate, well out on wing, bordered outwardly by a
narrow blackish line, inwardly (from top of cell to inner
margin by a broad salmon-ocherous patch filling the
space between antemedian line and the vertical scale
ridge; the latter thin, blackish, bordered inwardly by a
white line; subterminal line near outer margin, indis-
tinct, pale, bordered inwardly by some obscure blackish
streaklets on the veins; discal dots separated; all the
markings obscure except the white inner margin of the
subbasal scale ridge. Hind wing translucent, pale
smoky fuscous; the veins and terminal margin faintly
darkened. Alar expanse, 21 mm.
Male genitalia with flaring apical process of gnathos
terminating in a shortly forked hook. Harpe with a
broadly triangulate projection (clasper) from near apex
of sacculus, the sacculus otherwise not appreciably
produced at apex. Penis armed with three clusters of
straight, strong spines.
TypE Locauity: Corozal, Canal Zone, Panamd (Nov.;
type in USNM).
Foop prant: Unknown.
Represented only by the male type.
191. Zamagiria pogerythrus Dyar
Fiaures 49, 287, 765
Zamagiria pogerythrus Dyar, Ins. Insc. Menstr., vol. 7, p. 47,
1919.
Aigrette of male maxillary palpus red.
Forewing pale brownish gray dusted and blotched
with white, slightly darker along costa; the white scaling
most conspicuous about the lower discal spot, as a more
or less diffused blotch in median area following the
antemedial line, as an interrupted white line preceding
the subbasal scale ridge, and as a short, narrow shade
from apex in terminal area; antemedial pale line faint,
its outer blackish border interrupted on the males,
continuous on females; the subbasal scale ridge broken
into two or three patches of black intermixed with some
red raised scales; the patch between these and the ante-
medial line more restricted and fainter than on dixo-
lophella, ocherous fuscous; subterminal line well con-
trasted, indented at vein 6, white, margined inwardly by
blackish streakings on the veins and inwardly and out-
wardly by dark spots on costa; discal dots separate,
black, the lower one elongately enlarged and the most
conspicuous black marking on the wing; a row of 4 or
5 black dots along terminal margin. Hind wing whit-
ish, translucent; a dark shade along costa and a narrow
dark line along termen; the veins very faintly darkened.
Alar expanse, 20-22 mm.
Male genitalia with apical process of gnathos con-
torted. Harpe with apex of sacculus shortly and
bluntly produced at apex. Penis armed with two
clusters of straight, stout, moderately long spines.
Female genitalia with signum developed as a narrow,
elongate, strongly spined plate extending the length of
the bursa; bursa otherwise unsclerotized and minutely
spinose; ductus bursae very broad (as broad as long and
broader than bursa), its membrane thickened and
bearing on its inner dorsal surface, near genital opening,
a large pair of conjoined sclerotized plates, and on its
inner ventral surface, near junction of bursa and ductus,
a large, thickened, corrugate, triangulate, sclerotized
plate bearing minute spines over its inner surface.
Eighth-segment collar ventrally fused.
Type tocauity: Chejel, Guatemala (type @, in
USNM).
Foop prant: Unknown.
Distrisution: Méx1co: Campeche (July).
MALA: Chejel (June, Aug.), Purulhé (July).
GUATE-
192. Zamagiria hospitabilis Dyar
Ficure 288
Zamagiria hospitabilis Dyar, Ins. Insc. Menstr., vol. 7, p. 48,
1919.
Aigrette of maxillary palpus pale red.
Forewing ocherous brown heavily overlaid with black,
giving the wing a dark, blackish brown ground color;
two strongly contrasted white markings, the inner
border of the broken, black, subbasal scale ridge and a
large, irregular blotch on lower median area just beyond
the antemedial line; antemedial line not strongly con-
trasted but distinguishable throughout, sinuate, whit-
ish, bordered outwardly by a thin black line; subtermi-
nal line more contrasted, dull white, indented at vein 6
and at lower fold; discal dots confluent, forming a black
lunate mark on discocellular vein; outer area beyond
subterminal line very faintly dusted with white; ter-
minal dots confluent forming a narrow black line along
outer margin of wing. Hind wing semihyaline white
with a dark shade along costa; the veins and terminal
margin faintly darkened. Alar expanse, 21 mm.
Male genitalia with apical process of gnathos broadly
flaring and serrate. Harpe with sacculus shortly pro-
duced at apex, very long and with broadly triangulate,
median, clasperlike projection (similar to that of
dizolophella). Lateral margins of aedeagus armed with
short, stout spines; penis unarmed except for a small,
flat, weakly sclerotized plate.
Typx Locauity: Ténamo, Cuba (type in USNM).
Foop pirant: Unknown.
Represented only by the male type.
193. Zamagiria masculinus Dyar
FicurE 289
Zamagiria masculinus Dyar, Ins. Inse. Menstr., vol. 7, p. 46,
1919.
Aigrette of male maxillary palpus whitish ocherous.
Forewing very pale ocherous gray thinly dusted with
white in median area; the ocherous tint strongest in the
patch between scale ridge and antemedial line, along
the costal edge, and broadly along lower fold; ante-
medial and subterminal lines distinguishable, but not
strongly accented, whitish; black scaling of subbasal
scale ridge limited to one or two dots; blackish outer
92 UNITED STATES NATIONAL MUSEUM BULLETIN 207
border of antemedial line interrupted; inner blackish
border of subterminal line broken into short black
streaks on the veins; discal dots separated, blackish; a
row of small blackish dots (5 or 6) along termen between
the vein ends; on inner margin at lower outer edge of
antemedial line a small white spot; none of the white
markings strongly contrasted; the usual black markings
broken into dots and very short dashes; overall shade a
pale clay color. Hind wing translucent, white with a
faint ocherous tint; a narrow pale brownish line along
termen; the veins very faintly darkened. Alar expanse,
25 mm.
Male genitalia with gnathos terminating in a broad,
inverted, heart-shaped apical projection. Harpe with
sacculus short, sharply pointed and shortly projecting
at apex. Penis armed with a single stout spine about
one-third as long as aedeagus.
TypEr Locality: Cayuga, Guatemala (Apr.; type in
USNM).
Foop pLrant: Unknown.
The largest known species of Zamagiria.
only from the male type.
Known
194. Zamagiria australella (Hulst), new combination
Figures 292, 766
Selagia australella Hulst, Canadian Ent., vol. 32, p. 174, 1900.—
McDunnough, Check list, No. 6232, 1939.
Immyrla bumeliella Barnes and McDunnough, Contributions,
vol. 2, p. 182, 1913.—McDunnough, Check list, No. 6188,
1939. (New synonymy.)
Aigrette of male maxillary palpus ocherous white.
Forewing white dusted with black scales, giving the
wing a pale gray color, lightest in basal and through the
median areas, slightly darker along costa; subbasal scale
ridge conspicuous and normally unbroken, black with a
fine white inner border; the patch between scale ridge
and antemedial line olivaceous ocherous; antemedial
line oblique, dentate-sinuate, faint, indicated chiefly by
its black outer bordering line; subterminal line dentate-
sinuate, bordered inwardly and outwardly at costa by
blackish dashes, outwardly below costa by a narrow
brownish shade and inwardly by a fine black line; discal
dots distinct, separate, black; a small blackish or
brownish spot on inner margin a slight distance beyond
antemedial line and diffused black smudges on veins 2,
3 and 4 for a short distance from cell. Hind wing
semihyaline white; a darker line along termen and the
veins very slightly darkened. Alar expanse, 15-18 mm.
Male genitalia with apical process of gnathos a nearly
square plate. Harpe. with costa narrowly sclerotized
and shortly produced at’ apex; sacculus shortly and
bluntly produced. Penis armed with a single stout
cornutus about half as long as aedeagus.
Female genitalia with bursa sclerotized and densely
spined at posterior end, the sclerotization extending
into ductus bursae; occupying most of the remainder of
ductus bursae a funnel-shaped, convoluted, sclerotized
plate.
Typr LocaALitizs: Blanco County, Tex. (australella,
in AMNH, ex Rutgers); Fort Myers, Fla. (6umeliella
in USNM).
Foop piant: Bumelia microcarpa.
Distrisution: Texas, Blanco County; Florida, Fort
Myers (May), Miami.
The genitalia of the female type of australella are
identical with those of a reared female paratype of
bumeliella, and the two moths otherwise agree, so there
is DO question of the synonymy. However, I doubt
somewhat the correctness of the locality label on Hulst’s
type and suspect that it may be a Florida specimen.
195. Zamagiria fraterna, new species
Figure 291
Aigrette of male maxillary palpus white.
Forewing white on dorsal half and along inner margin
beyond antemedial line to tornal angle; extreme base of
costa rough scaled, black (similar black sex-scaling on
underside of wing at base) ; antemedial line at middle of
wing incomplete, indicated only by an angulate white
line on lower half, bordered inwardly and outwardly by
small black smudges; along lower margin to antemedial
line the ground color is pale brown; a faint dusting of
brown or purplish brown scales on the white ground for
a Darrow margin along costa; scale ridge interrupted, a
series of black dots with the inner white bordering line
very faintly indicated; the usual patch between scale
ridge and antemedial line pale brown, a trifle paler than
the ground color on lower basal area of wing; some simi-
larly colored scales in the lower postmedian area be-
tween the antemedian line and the end of cell and in the
interspaces of veins 3 to 5; subterminal line incomplete,
only its middle portion distinguishable; discal dots dis-
tinct, separated, black, the lower one enlarged; black
scaling along veins 3, 4 and 5. Hind wing hyaline
white, a pale brownish gray line along terminal margin
and a similar shade along costa; at extreme base of wing
a few black sex-scales on the veins. Alar expanse,
21.5 mm.
Male genitalia with apical process of gnathos roundly
spatulate. Harpe considerably broadened towards
middle, thence sharply tapered to the narrow cucullus;
costa strongly humped at middle; sacculus produced at
apex into a long, strongly sclerotized, curved, free,
hooklike arm. Anellus a U-shaped plate. Penis armed
with a single stout spine (about one-fourth as long as
aedeagus), a strongly sclerotized, corrugated plate and
a concentration of rather coarse granulations.
TYPE LOCALITY: Santiago de las Vegas, Cuba (type
in USNM, 61841).
Foop piant: ‘Caimitillo.”
Described from male type reared by A. Otero, Aug.
17, 1932, and labeled: ‘“‘Leaf tier on Caimitillo, E. E. A.
Cuba, Ento. No. 10006.”
196. Zamagiria laidion (Zeller)
Figures 290, 767
Myelois laidion Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 211,
Piesmopoda laidion (Zeller) Ragonot, Monograph, pt. 1, p. 162,
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 93
Zamagiria laidion (Zeller) Dyar, Ins. Inse. Menstr., vol. 7, p. 46,
1
Zamagiria deia Dyar, Ins. Insc. Menstr., vol. 7, p. 46, 1919 (new
synonymy).
Zamagiria striella Dyar, Ins. Insc. Menstr., vol. 7, p. 47, 1919
(new synonymy).
Aigrette of male maxillary palpus dull white.
Forewing pale gray, the overall tint shading from
grayish white to very pale bluish gray (in fresh reared
examples) ; costal border slightly darker; the patch be-
tween scale ridge and antemedial line reddish brown;
similar reddish brown scaling spread over basal area
bordering inner margin and frequently blotching the
median area over vein 1b and the lower fold (especially
on the males); also some scattering of reddish brown
streaking on veins 8 and 4 for a very short distance from
cell; black scale ridge more or less complete; on several
males a black or black and red-brown streak along vein
1b from scale ridge to base of wing; transverse lines
faint, the outer blackish border of the antemedial and
the inner dark border of subterminal more or less inter-
rupted (more so on males than females); discal dots
separated, black, the lower one slightly enlarged; a
row of 5 or 6 small black dots along termen. Hind
wing semihyaline white; a dark line along terminal mar-
gin and some faint darkening of the veins. Alar ex-
panse, 15-22 mm.
Male genitalia similar to those of fraterna except:
Costa of harpe convex but not decidedly humped; an-
nellus inverted;* penis with cornutus a single, very
stout, long spine (over two-thirds as long as aedeagus)
surrounded by a cluster of small granulations.
Female genitalia with a finely spined plate (signum)
near junction of bursa and ductus bursae, and a strongly
sclerotized lateral patch in bursa near its anterior end;
ductus bursae with a sclerotized plate occupying most
of its length, the posterior end of the plate folded over
into triangulate ventral lips.
Types LocaLitres: Honda, Colombia (/aidion, in BM);
Chejel, Guatemala (deia and striella, in USNM).
Foop piants: Achras sapota, Mimusops emarginata,
Eriobotyra japonica (larvae feeding on leaves and
flowers).
DisrrisuTion: Unirep States: Florida, Key West
(Apr., May), Miami (Jan., Dec.). Guatemaua: Chejel
(June). Panamd: Porto Bello (Feb.). Cotompta:
Honda. Braziu: Castro (Parand) Obidos (Amazon re-
gion, Sept.), Vigosa (Minas Geraes, Sept.). Boxrvia:
“Hast Bolivia” (“Aug.—Oct., T. Steinbach”).
Hitherto laidion has been recognized only from female
examples. A reared series in the National Museum
from Florida has enabled us to associate the sexes and
has established the synonymy of deia and striella, both
described from males. Dyar’s type of striella is merely
an extreme example of a common color variant with
3 This structure seems more like a transtilla than an anellus;
for its straight posterior margin lies between the costal bases of
the harpes, and it could be interpreted as a transtilla or com-
bination of transtilla and anellus, except that in other species of
the genus there is no trace of even the vestiges of a true transtilla.
more or less black streaking on the base of vein 1b of
forewing. The reared Florida specimens have also
given us the host records cited above.
197. Zamagiria ipsetona Dyar
Fiaure 768
Zamagiria ipsetona Dyar, Ins. Insc. Menstr., vol. 7, p. 47, 1919.
Forewing gray, more heavily marked with black than
in preceding species; the blackish streakings on the
veins (2 to 6 in this species) longer and stronger; lower
discal spot elongated into a black dash; dark dashes
(reddish brown) on the veins in outer area following the
faint subterminal line; the black scale ridge not con-
tinuous; antemedial line sinuate and nearly vertical;
the whitish areas limited to a pale oval patch surround-
ing the black-streaked veins and discal dots and a faint
shade preceding the black scale ridge; the usual red-
brown scaling limited to the patch between scale ridge
and antemedial line and weak shadings between some
of the veins in postmedian area. Hind wing translu-
cent white; a narrow dark shade along costa and a
blackish line along termen; veins appreciably darkened.
Alar expanse, 23-24 mm.
Female genitalia with two elongate spined plates in
bursa; another more weakly spined plate at junction of
bursa and ductus bursae; from the junction a scleroti-
zation extends along one side of bursa for about four-
fifths of its length; ductus bursae very short, strongly
sclerotized towards genital opening, the plate folded
over at the opening into ventrolateral lips. Eighth-
segment collar completely fused ventrally.
TYPE LOCALITY: Juan Vifias, Costa Rica (type in
USNM).
Foop pLant: Unknown.
Represented by three females from the type locality
(Feb., June). The male is unknown.
49. Genus Anegcephalesis Dyar
Anegcephalesis Dyar, Ins. Insc. Menstr., vol. 5, p. 46, 1917.—
Heinrich, Ins. Insc. Menstr., vol. 5, p. 48, 1917 (larva).
(Type of genus: Anegcephalesis cathaeretes Dyar.)
Characters of Zamagiria except: Maxillary palpus
minute, subsquamous in both sexes. Forewing with
veins 4 and 5 connate or very shortly stalked. Hind
wing with cell longer (slightly less than one-third the
length of wing).
The genus is very close to Zamagiria but its separa-
tion seems to be justified by the differences in the male
maxillary palpi and the longer cell of hind wing. The
venational differences noted above (between veins 4 and
4 of forewing) may be only a specific character. Dyar
distinguished Anegcephalesis from Zamagiria on the
difference in maxillary palpi, but mistook the appressed
scale tuft from frons for that organ. This scale tuft is
present and equally well developed in Zamagiria and
Anegcephalesis.
94. UNITED STATES NATIONAL MUSEUM BULLETIN 207
198. Anegcephalesis arctella (Ragonot), new combination
Figures 50, 294, 770
Phycita arctella Ragonot, N. Amer. Phycitidae, p. 4, 1887;
Monograph, pt. 1, p. 180, 1893.
Anegcephalesis cathaeretes Dyar, Ins. Insc. Menstr., vol. 5, p. 46,
1917 (new synonymy).
Anegcephalesis catheretes (Dyar), Ins. Insc. Menstr., vol. 7, p.
48, 1919 (food plant and altered spelling)—MceDunnough,
Check list, No. 6176, 1939.—Bruner, Scaramuzza, and
Otero, Bull. 63, Estaci6n Exp. Agron., Cuba, p. 69, 1945.
Forewing brownish gray dusted with white; the
white shading more pronounced (on the males) in the
median area above the base of the cell, as a small spot
on inner margin below the dark markings on veins 2
to 5, and in outer area beyond the dark outer border of
the subterminal line, on the female the white dusting
is more generally distributed over median and outer
areas and, on some specimens, on the basal area, giving
the wing an ashy gray appearance; subbasal ridge black
bordered inwardly by a white line and followed by a
dull olivaceous ocherous patch; antemedial line well
out towards middle of wing, oblique, notched at vein 1b
and evenly curved above it, outwardly bordered by a
continuous black line; subterminal line distinct; in-
dented just below costa and very slightly so at 1b;
discal spots fused into a lunate black line along dis-
cocellular vein; a dark brownish smudge over veins 1
to 5 adjacent to cell. Hind wings translucent white;
the veins very faintly darkened. A narrow brown line
along termen. Alar expanse, 20-23 mm.
Male genitalia with uncus narrowly triangulate.
Apical process of gnathos a thin, broadly ovate plate.
Harpe with appressed, elongate clasper; sacculus not
produced at apex. Aedeagus sharply bent; penis armed
with an elongate, flattened, partially bent, platelike
cornutus, and some sclerotized wrinklings and granu-
lations.
Female genitalia with signum developed as an elon-
gate, strongly spined plate (about half as long as bursa) ;
opposite the signum a narrower, much more weakly
sclerotized plate beginning at junction of bursa and
ductus bursae (this plate about half the length of
signum) ; bursa, otherwise smooth; ductus bursae tubu-
lar, sclerotized. Highth-segment collar fused ven-
trally.
Typ Locauity: Nassau, Bahamas (arctella, in Paris
Mus.; cathaeretes, mn USNM).
Foop piant: Dipholis salicifolia.
Distrisution: Unitep States: Florida, Miami,
Paradise Key (Mar.), Royal Palm State Park (Feb.).
Bawamas: Nassau (New Providence Isl., Feb., Mar.,
April, May, Sept.). Cusa: Baracoa (Oct.), Santiago
Province (Feb., June, Sept., Oct., Dec.).
Ragonot’s arctella was described from a single female.
His description and figure agree in every detail with
Dyar’s cathaeretes represented by a large series in the
U. S. National Museum. Both were described from
the same type locality. There is no doubt that both
names apply to the same species.
50. Magiriopsis, new genus
TyPE oF GENUS: Sematoneura denticosella Dyar.
Characters of Zamagiria except: Antenna of male
unipectinate for basal half of shaft, serrate and shortly
ciliate beyond; shaft with very shallow sinus near base,
containing a few roughened scales and a short row of
minute spines but without scale tuft. Labial palpi of
male broad, dorsoventrally flattened and appressed to
face (not to each other as in Zamagiria and Anegcephale-
sis); hollowed inner surface; third segment greatly re-
duced and completely hidden by scaling of second
segment. Male head without scale tuft from upper
edge of frons, not deeply grooved between antennae but
with an enlarged scale tuft behind antennae. Maxillary
palpus of female squamous (of male, as in Zamagiria,
in the form of an aigrette). Forewing smooth; vein 2
from well before outer angle of cell; 6 slightly bent
towards base but from below upper angle of cell, sepa-
rate from stalk of 8-9 at base. Hind wing with cell
less than one-third the length of wing.
Male genitalia with complete transtilla.
199. Magiriopsis denticosella (Dyar), new combination
Figures 293, 769
Sematoneura denticosella Dyar, Proc. U. S. Nat. Mus., vol. 42,
p. 105, 1912.
Hypsipyla denticosella (Dyar), Ins. Insc. Menstr., vol. 7, p. 41,
1919.
Crocidomera cristalis Hampson, Ann. Mag. Nat. Hist., ser. 10,
vol. 4, p. 352, 1929 (new synonymy).
Aigrette of male maxillary palpus whitish ocherous.
Forewing reddish (rust) brown, the costal third lightly
dusted with white, especially along the costa and be-
tween the veins, the dark color somewhat accented on
the veins and, on occasional female specimens, some
black scaling on the veins before the usual location of
subterminal line; the latter rarely indicated ; antemedial
line and discal spots obsolete. Hind wing translucent,
opalescent with a smoky shade at apex; a dark line along
termen and the outer veins faintly darkened. Alar
expanse, 32-40 mm.
Male genitalia with uncus narrowly triangulate.
Apical process of gnathos a long, strong spine, sharply
hooked at apex. Transtilla a weakly sclerotized, shield-
like bridge attached to costal bases of harpes. Harpe
with costa strongly sclerotized for basal third of harpe
and terminating in a broad projection; sacculus with a
broad projection from base and a broadly projecting
clasper from apex. Aedeagus long and stout; penis
armed with three clusters of strong spines (about as
long as width of aedeagus). Vinculum elongate (two
and one-half times as long as greatest width), stout, not
appreciably tapering; posterior margin reinforced and
squarely excised.
Female genitalia with two irregularly shaped and
strongly spined plates in bursa, and bursa at junction
with ductus bursa strongly sclerotized, the wrinkled
sclerotization extending part way into the ductus;
ductus bursae otherwise only sclerotized (weakly) at
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 95
genital opening. Highth-segment collar not fused
ventrally.
TypPE LocaLitiEs: Orizaba, México (denticosella, in
USNM); Juan Vifias, Costa Rica (cristalis, in BM).
Foop piant: Unknown.
Distrisution: México: Orizaba (Oct.), Misantla
(Aug.). Guatemata: Cayuga (Jan., Aug.), Chejel
(June). Costa Rica: Juan Vifias (Jan., Nov.). Co-
LomBiA: Upper Rio Negro. VeEnnzuEua: Aroa, San
Esteban Valley. British Guiana: Omai. Braziu:
Ponte Nova (Rio Xingu, Amazonas), Santos (Mar.).
Both denticosella and cristalis were described from
females. The synonymy is obvious. Both Dyar and
Hampson had seen males many years before but evi-
dently overlooked them when writing their descriptions.
Genus 51: Ancylostomia
Venational division D. Forewing with veins 4 and 5 contiguous
or stalked for about one-third their lengths from cell; 3 connate
with 4 at base; 6 bent towards base, connate with the stalk of
8-9. Hind wing with veins 4 and 5 stalked to middle; 3 closely
approximate to the stalk of 4-5 at base; cell short. Labial palpus
obliquely ascending with third segment long and porrect. Uncus
pentagonal. Gnathos produced at apex into a pair of long,
flattened, pointed, contorted, bandlike projections. Female
with bursa copulatrix weakly sclerotized throughout; eighth-
segment collar modified ventrally. Eighth abdominal segment
of male with compound ventral tufts.
51. Genus Ancylostomia Ragonot
Ancylostomia Ragonot, Monograph, pt. 1, p. 567, 1893. (Type
of genus: Myelois stercorea Zeller.)
Tongue well developed. Antenna of male with a
sinus and strong scale tuft in shaft at base, shaft other-
wise flattened, weakly serrate and pubescent; of female,
simple and pubescent. Labial palpus obliquely ascend-
ing, with third segment porrect; second segment long,
reaching well above vertex, on male hollowed to receive
maxillary palpus; third segment slender, acuminate, on
male half as long as second, on female about the same
length as second segment. Maxillary palpus of male
in the form of an aigrette; of female, squamous. Fore-
wing smooth; 11 veins; vein 2 from before but near
lower outer angle of cell; 3 from the angle, connate with
4; 4 and 5 contiguous or stalked for about one-third
their lengths from cell; 6 from upper angle of cell, con-
nate with the stalk of 8-9, bent towards base; 8 and 9
stalked for slightly more than half their lengths; 10 from
the cell, approximate for a short distance to the stalk of
8-9; male without costal fold. Hind wing with vein
2 from before lower outer angle of cell; 3 from the angle,
closely approximate to 4 at base; veins 2 and 3 very
long; 4 and 5 stalked to middle; 7 and 8 anastomosed or
contiguous for a short distance from cell (for less than
half their lengths); cell less than one-third (about one-
fourth) the length of wing; discocellular vein curved.
Eighth abdominal segment of male with compound,
ventral scale tufts.
Male genitalia with uncus pentagonal (the sides
parallel, the terminal margin angled). Gnathos pro-
duced at apex into a pair of long, flattened, sharply
pointed and slightly contorted, bandlike projections;
the lateral arms short and articulated to base of uncus.
Transtilla absent. Harpe elongate, slender, cucullus
reduced, its apex bluntly pointed; sacculus produced
at apex into a slender, free, curved hook. Penis armed
with a pair of thin, somewhat flattened and twisted
spines as long (or nearly as long) as aedeagus. Vincu-
lum slightly longer than greatest width; terminal margin
broadly rounded.
Female genitalia without signum; bursa weakly
sclerotized throughout; ductus bursa strongly sclero-
tized except for a narrow space beyond junction of
bursa, at junction with bursa finely ridged and spined,
the ridges and spines extending for a short distance into
bursa, at genital opening the margins of ductus fuse
into eighth-segment collar. Eighth-segment collar very
strongly sclerotized; enlarged but not fused ventrally;
its ventrocaudal angles produced and pointed; from its
ventrolateral angles a pair of invaginated sclerotized
pockets.
The genus is easily identified by its peculiarly de-
veloped gnathos and the eighth-segment collar of the
female. Its species are tropical American in distribu-
tion with (in the case of stercorea) a slight extension of
range into southern Florida.
200. Ancylostomia stercorea (Zeller)
Figures 295, 771
Myelois stercorea Zeller, Isis von Oken, 1848, p. 873.
Anerastia ignobilis Butler, Proc. Zool. Soc. London, 1878, p. 494.
Pempelia diffissella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p.
178, 1881.
Ancylostomia stercorea (Zeller) Ragonot, Monograph, pt. 1, p.
568, 1893.
Forewing pale ocherous along the costa, through the
median fold and over most of the basal area; some
pinkish brown shading between the veins in the pale
areas giving the ocherous ground color a rosy tint
(especially on reared and fresh specimens); the lower
outer half of wing heavily shaded with brown more or
less dusted with black (on reared examples this area is
decidedly blackish brown, its inner margin oblique
from lower outer angle of cell to basal third of inner
margin); a distinct brown or black streak under vein 8
from apex about half-way to cell and a similar longer
dark streak along vein 6 from terminal margin to the
cell; a thin whitish line along the lower margin of cell
from beyond base and continuing into vein 4 for a short
distance; a similar shorter white line on the subbasal
half of vein 1b; a small black dot on the middle of the
white streak on vein 1b and a similar black dot above
it on the white streak on lower margin of cell; a single,
large, conspicuous black discal spot at lower outer angle
of cell; antemedial and subterminal lines obsolete; a
few black dots or streaklets in outer areas about the
middle of veins 2, 3, and 4 and on dark specimens some
blackish or dark brown shading on upper and lower
96 UNITED STATES NATIONAL MUSEUM BULLETIN 207
veins of cell at base of wing. Hind wing translucent
white with a smoky shade along costa, towards apex
and along termen; the veins slightly darkened. Alar
expanse 16-28 mm.
Genitalic characters as given for the genus.
segment collar of female laterally corrugated.
TYPE LOCALITIES: Brazil (stercorea, in BM); Jamaica
(ignobilis, in BM); Honda, Colombia (diffissella, in
BM).
Foop pPLant: Cajanus cajan (larvae feeding in the
pods).
DistRipution: Unirep States: Florida, Cocanut
Grove (Apr., May), Goulds (May), Homestead (Apr.),
Jupiter (Apr.), Miami (May). Cua: Santiago Prov-
ince (May, June, Oct., Dec.), Santiago de las Vegas
(May). Harrr (June). Dominican Repusiic (Aug.).
Pusrto Rico: Isabela (Apr.), Puerto Real (Vieques
Isl., Apr.). Viremn Istanps: Kingshill (St. Croix;
June, “Noyv.—Dec.”). Jamatca (Mar.). BAHAMAS:
Nassau (May). Grenapa: Nevis (Jan.); St. Kitts
(June). Trinmpap (Mar., May). México: Cuerna-
vaca (July), Jalapa, Orizaba, Zacualpin (July). Gua-
TEMALA: Chejel (July, Aug.), Volcin Santa Maria
(June, July). Costa Rica: Juan Vifias (June.) Pan-
amA: La Chorrera (May), Rio Trinidad (Mar.). Co-
Lompia: Honda. Frencw Guiana: Cayenne. Bra-
ziL: Castro.
The food plant and Florida records are from a large
series reared by the Special Survey of the U.S. Bureau
of Entomology and Plant Quarantine in 1944. Wealso
have a couple of reared adults and several collected
larvae from chickpea (Cicer) and black-eyed pea
(Dolichos). The favored host, however, seems to be
the pigeonpea (Cajanus). In the pods of that plant
the larvae are abundant throughout the West Indies
and in southern Florida.
HKighth-
201. Ancylostomia sauciella (Zeller)
Figure 296
Pempelia sauciella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p.
183, 1881.
Ancylostomia sauciella (Zeller) Ragonot, Monograph, pt. 1, p.
569, 1893.
Forewing cinnamon red mixed with rose; costa paler;
a narrow subcostal reddish brown shade extending from
base to apex; the veins somewhat accented by reddish
scaling; a small blackish discal dot at lower outer angle
of cell; a similar dot on basal third of vein 1b in a white
streaklet on the vein; from apex a short oblique blackish
shade. Hind wing yellowish white, translucent; a thin
dark line along termen and the veins slightly darkened.
Alar expanse, 20 mm.
Male genitalia as in stercorea except uncus narrower
and apical projecting arms of gnathos shorter and
decidedly broader.
TypPE Locauity: Maraquita, Colombia (type in BM).
Foop prant: Unknown.
Known only from the type.
202. Ancylostomia argyrophleps Dyar
Ancylostomia argyrophleps Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 406, 1914.
Similar to sauciella except: Costa and lower half of
basal area carneous white; a broad subcostal band
extending from base to apex, red-brown shaded with
black especially towards apex; lower outer area of the
same color with a smoky fuscous shade along outer half
of inner margin from near tornus; a white line along
lower margin of cell and vein 5 from basal third, enclos-
ing a black dot at lower outer angle of cell; a similar
black dot on basal third of vein 1b. Hind wing trans-
lucent, semi-irridescent white with a faint smoky tint,
the latter more pronounced towards apex; veins dark-
ened in outer area on females, not appreciably so on
males. Alar expanse, 20-25 mm.
Male genitalia as in stercorea except apical processes
of gnathos a trifle broader (but not so broad as on
sauciella). Highth-segment collar of female smooth.
Female genitalis otherwise essentially as in stercorea.
TyprE Locatiry: Orizaba, México (type, co’, m
USNM).
Foop pranr: Unknown.
Distrisution: México: Orizaba (Aug.), Cuernavaca
(July). Guatemata: Chejel (July).
203. Ancylostomia euchroma Dyar
Ancylostomia euchroma Dyar, Ins. Inse. Menstr., vol. 7, p. 53,
Forewing below the cell from base to tornus red-
brown; the upper area white-lined between the veins
and along the median fold in cell; the veins red-brown;
a black dot at lower outer angle of cell, one on lower
margin of cell before its middle and another on basal
third of vein 1b; a diffused pale shade surrounds this
last dot and extends obliquely backward to inner
margin, an oblique line of black dots on veins 2, 3, and
4; on the female a smoky brown shade from apex
extending narrowly along costa to base; on male the
shade from apex is short and cinnamon red, and the
dark area on lower half of wing is a bright cinnamon
red. Hind wing in the male translucent white with a
faint smoky shade at apex; on the female the smoky
shade extends further back from apex and outer margin
and the veins in outer area are appreciably darkened.
Alar expanse, 24-25 mm.
Male genitalia with the projecting apical bands from
gnathos as broad as those of sauciella but longer (at
least as long as those of stercorea). Female genitalia
with the sclerotized ribbing and spining at junction of
bursa and ductus bursae slightly stronger than those of
either stercorea or argyrophleps. Highth-segment collar
very weakly corrugated.
Type Locatiry: Castro, Parand, Brazil (type in
USNM).
Foop prant: Unknown.
Represented only by the female type and male
paratype from the type locality. May be only a race
of sauciella.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 97
Genus 52: Caristanius
{Venational division B. Forewing with veins 4-5 separated at
base and divergent shortly beyond, smooth; hind wing with vein
3 approximate to the stalk of 4-5 at base. Highth abdominal
segment of male with compound tufts. Antenna of male with
sinus and scale tufts on base of shaft. Labial palpus obliquely
ascending; second segment of male grooved. Maxillary palpus
of male in the form of an aigrette. Male genitalia with a pair of
long, strongly sclerotized arms from ventrolateral margins of
uncus; transtilla absent; gnathos absent; a single strong cornutus
on penis. Female genitalia with signa present, developed as
coarsely spined plates; ductus bursae ribbonlike, sclerotized.]
52. Caristanius, new genus
Type or GENUs: Oligochroa pellucidella Ragonot.
Tongue well developed. Antenna of male with sinus
and stout scale tuft on base of shaft, finely pubescent;
of female simple. Labial palpus obliquely ascending;
on male stout and smooth scaled (the palpi appressed
to each other), second segment long, reaching well above
vertex, grooved to hold maxillary palpus, third segment
very short, hidden in scaling of second; on female slen-
der, shorter, reaching slightly above vertex, more
roughly scaled, third segment over half as long as sec-
ond, acuminate. Maxillary palpus of male in the form
of an aigrette; of female squamous. Forewing smooth;
11 veins; vein 2 from before lower outer angle of cell;
3 from the angle; 4 and 5 shortly separated at base and
divergent very shortly beyond; 6 from below upper
angle of cell, straight; 8 and 9 stalked for half or less
than half their lengths; 10 from the cell; male without
costal fold. Hind wing with vein 2 from before lower
outer angle of cell; 3 from the angle, approximate to
stalk of 4-5 at base; veins 2 and 3 of moderate length;
4 and 5 stalked for half their lengths; 7 and 8 approxi-
mate beyond cell (for less than half their lengths) ; cell
less than half the length of wing (but more than one-
third); discocellular vein curved. Eighth abdominal
segment of male with compound ventral scale tuft.
Male genitalia with uncus short, decidedly broader
than long; a pair of long, strong, sclerotized arms from
its ventrolateral angles. Gnathos absent (unless the
fused arms from uncus can be interpreted as parts of
this organ, which is extremely doubtful). Transtilla
absent. Harpe witb costa strongly sclerotized and with
a strong projection from its base or a strong clasperlike
projection from below it near base; sacculus short, nar-
row, weak, bluntly produced at apex; cucullus very nar-
row, weak and reduced. Anellus a narrow band with a
central, bifurcate, bandlike projection, flanked by elon-
gate, lateral lobes. Aedeagus elongate, moderately
slender, straight; penis armed with a single strong cornu-
tus and some fine spines and granulations. Vinculum
stout, as long as or longer than greatest width.
Female genitalia with signa present, developed as
elongate, curved, sclerotized, strongly and coarsely
spined plates; ductus bursae flattened, ribbonlike,
strongly sclerotized except for a narrow space near
middle, also granulate towards bursa, the granulations
extending into bursa; at genital opening ductus bursae
300329—56——8
very strongly and broadly sclerotized; ductus seminalis
from bursa near junction of bursa and ductus bursae.
The species referred here have been placed in Elasmo-
palpus. ‘They agree with the type of that genus (ligno-
sellus) on antennal, palpal, and venational characters;
but differ strikingly on male and female genitalic struc-
ture. The characteristic uncus with its long, produced,
basal arms, the reduced, weak sacculus, the lack of any
distinguishable gnathos, and the flattened, ribbonlike
ductus bursae with its strongly sclerotized development
at genital opening at once distinguish Caristanius from
Elasmopalpus.
204. Caristanius pellucidellus (Ragonot), new combination
Ficures 297, 775
Oligochroa pellucidella Ragonot, Bull. Soc. Ent. France, 1888,
p. exl.—Mbéschler, Die Lepidopteren—Fauna von Portorico,
p. 329, 1890.
Elasmopalpus pellucidellus (Ragonot), Monograph, pt. 1, p. 429,
1893.
Rhodophaea melanoplaga Hampson, in Ragonot, Monograph, pt.
2, p. 519, 1901 (mew synonymy).
neo SSS
oe a
Ground color of forewing variable, pale brownish gray
to dark gray with a faint purplish or reddish brown tint;
transverse markings obsolete on most specimens; on
some the antemedial line indicated by a narrow, very
faint, pale line between vein 1b and inner margin, pre-
ceded by a pale reddish or brownish patch more or less
overlaid by black scaling (on most specimens before me
this spot entirely absent); subterminal line distinguish-
able only on a few specimens, very faint, indicated
chiefly by some short blackish streaklets on the veins;
discal spots usually distinct but faint, blackish, sepa-
rated; a row of black dots along termen. Hind wing
transparent white with a dark shade along costa and at
apex and extending as a dark line downward along ter-
men; on the males this dark line extends only to about
middle of termen, on females to or nearly to anal angle
of wing. Alar expanse, 19-25 mm.
Male genitalia with costa of harpe broadly and stoutly
sclerotized, a large oval projection from base and its
apical end folded backward and contorted. Cornutus
about one-third as long as aedeagus. Vinculum about
as long as greatest width.
Female genitalia with granulations of ductus bursae
extending for a very short distance into bursa. Eighth-
segment collar not extended to ventral surface; at-
tached laterally to the extended sclerotization of ductus
bursae at genital opening; stoutly sclerotized dorsally
and with an inwardly projecting curved sclerotized
apron; between this apron and ductus bursae a gland
of unknown function extends into abdomen, terminat-
ing ina bulb with thickened membrane. This structure
not noted on other species of the genus.
Typx Locatities: Puerto Rico (pellucidellus, in Paris
Mus.); Sio Paulo, Brazil (melanoplaga, in BM).
Foop prant: Unknown.
DistriputTion: Puerto Rico: Puerto Real (Vieques
Isl., Apr., July), Rio Piedras (Aug., Sept.), San German
(Apr., Aug.). Sv. Vincent. Jamaica. Surinam: Zan-
98 UNITED STATES NATIONAL MUSEUM BULLETIN 207
derij I (Apr.). Braziz: Sio Paulo, “S. H. Brazil”
[probably Castro].
The species is somewhat variable in color and mark-
ings, but is easily identified by its genitalia. Hampson’s
melanoplaga was based on large Brazilian females (25
mm.). All specimens from the West Indies and Suri-
nam that J have seen are smaller (19 to 21 mm.). How-
ever, there does not seem to be any reason to keep
melanoplaga as a racial designation on a mere difference
in size, as there is nothing else to distinguish the Bra-
zilian examples.
205. Caristanius decoloralis (Walker), new combination
Figures 298, 773
Trachonitis decoloralis Walker, List, vol. 27, p. 42, 1863.
Nephopteryx metagrammalis Walker, List, vol. 27, p. 42, 1863.
Nephopteryx furfurella Hulst, Ent. Amer., vol. 3, p. 131, 1887;
Phycitidae of N. Amer., p. 143, 1890 (mew synonymy).
Elasmopalpus decoloralis (Walker) Ragonot, Ent. Amer., vol. 5,
p. 115, 1889; Monograph, pt. 1, p. 420, 1893.—MceDun-
nough, Check list, No. 6228, 1939.
Elasmopalpus floridellus Hulst, Phycitidae of N. Amer., p. 158,
1890.—Ragonot, Monograph, pt. 1, p. 423, 1893.
Elasmopalpus decoralis Hulst, Phycitidae of N. Amer., p. 158,
1890 (misspelling for decoloralis).
Elasmopalpus decorellus Hulst, in J. B. Smith, List of the Lepi-
doptera of Boreal America, No. 4341, 1891 (misspelling for
decoloralis). %
Elasmopalpus furfurellus (Hulst), Barnes and McDunnough,
Contributions, vol. 3, p. 196, 1916.—McDunnough, Check
list, No. 6230, 1939.
Forewing bluish gray or pale fawn gray with more or
less white dusting over median area; transverse lines
indistinct; antemedial line indicated chiefly by its
blackish outer border broken into dots on vein 1b, lower
vein of cell and a subcostal spot or short dark streak
from costa to top of cell; preceding antemedial line a
reddish patch on inner margin extending to or into cell
and more or less shaded by black scaling; subterminal
line sinuate, pale and very faint; lower discal spot at
end of cell distinct, the upper discal dot much smaller,
sometimes distinct but frequently obscured; a row of
black dots along termen. Hind wing translucent, whit-
ish with a pronounced smoky tint, especially over outer
half of wing; the veins more or less darkened and a fine
dark line along termen. Alar expanse, 21-25 mm.
Male genitalia with projecting arms from uncus
somewhat curved. Costa of harpe sclerotized for the
length of the harpe, narrowly sclerotized beyond base;
a strong, curved, clasperlike projection from below costa
near base, projected beyond costal edge. Cornutus as
long as aedeagus. Vinculum considerably longer than
greatest width.
Female genitalia with granulations of ductus bursae
extending deeply into bursa; ductus bursae scobinate
on lower surface at genital opening. Eighth-segment col-
lar narrowed dorsally, complete but not fusing ven-
trally, without sclerotized dorsal apron.
TYPE LocALItTIEs: United States (decoloralis and me-
tagrammalis, in BM); Florida (furfurellus and floridellus,
in AMNH, ex Rutgers).
Foop piant: Chamaecrista spp. (brachiata, fascicu-
lata, robusta) larvae feeding on the leaves. These rec-
ords from rearings by the Special Survey (1944) of the
Division of Foreign Plant Quarantine of the U. S.
Bureau of Entomology and Plant Quarantine.
Distrisution: Florida, Orlando (Feb.), no specific
locality (Mar.), Fort Myers (May), St. Petersburg
(June), Stuart (May), Tampa, Vero Beach (Apr., May,
June, Oct., Dec.); Texas, Brownsville; South Carolina,
Florence (June), Hiltonhead Isl. (Aug.), Pawleys Beach
(Sept.).
This species seems to be confined to the southeast-
ern United States. In his description of decoloralis
Walker mentions the protruding arms of the uncus.
Hulst noted similar structures in his floridellus and asso-
ciated them with the genitalia, suspecting the synonymy
of his and Walker’s species. I have before me a photo-
graph of the genitalia of the type of decoloralis supplied
by Clarke. They are identical with those of the male
type of floridellus. The synonymy of floridellus and
JSurfurellus was established by Barnes and McDunnough
(1916) and that of metagrammalis with decoloralis by
Ragonot (1889).
206. Caristanius guatemalellus (Ragonot), new combination
FIGURE 774
Salebria guatemalella Ragonot, Nouv. Gen., p. 18, 1888.
Laodamia guatemalella (Ragonot), Monograph, pt. 1, p. 414,
1893.
This species is apparently known only from the female
type. Ragonot’s description and figure suggest a large,
pale brownish, suffused form of pellucidellus; the fore-
wing pale ocherous brown tinted with reddish, trans-
verse lines absent, the lower discal spot distinct, and the
veins sparsely powdered with blackish scales; hind wing
semitransparent, white faintly tinted with ocherous.
Alar expanse, 26 mm.
The genitalia determined the present generic refer-
ence. The granulations of ductus bursae extend deeply
into the bursa as in decoloralis; the eighth-segment collar
has a dorsal, sclerotized, invaginated apron smaller
than and differently shaped from that of pellucidellus,
and the collar itself is completely sclerotized ventrally.
TypE LocaLity: San Geranimo, Guatemala (type in
BM).
Foop prant: Unknown.
Genus 53: Etiella
[Venational division B. Forewing with veins 4-5 separated and
divergent from base, a raised-scale ridge beyond base; hind wing
with vein 3 approximate to the stalk of 4-5 at base. Eighth
abdominal segment of male with paired tufts. Labial palpus
porrect, long; second segment of male grooved. Maxillary pal-
pus in the form of an aigrette. Male genitalia with apical process
of gnathos a simple, short hook; transtilla incomplete and
vestigial; harpe with a strong curved arm projecting the length
of the harpe from base of costa, harpe otherwise weakly sclero-
tized; two strong cornuti on penis. Female genitalia with signa
developed as curved, sclerotized bands armed with slender
spines; ductus bursae short, tubular, sclerotized.]
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 99
53. Genus Etiella Zeller
Etiella Zeller, Isis von Oken, 1839, p. 179; 1846, p. 733.—Heine-
mann, Die Schmetterlinge Deutschlands und der Schweiz,
Abt. 2, vol. 1, pt. 2, p. 154, 1865.—Meyrick, Proc. Linn.
Soc. New South Wales, vol. 3, p. 629, 1882.—Hulst, Phy-
citidae of N. Amer., p. 169, 1890; U. S. Nat. Mus. Bull. 52,
p. 428, 1902.—Ragonot, Monograph, pt. 1, p. 569, 1893.—
Spuler, Die Schmetterlinge Europas, vol. 2, p. 208, 1910.—
Forbes, Cornell Mem. 68, p. 629, 1923.—Janse, Journ. Ent.
Soc. South Africa, vol. 7, p. 15, 1944. (Type of genus:
Phycis zinckenella Treitschke.)
Ramphodes Guénée, Europaeorum Microlepidopterorum index
methodicus ..., p. 81, 1845. (Type of genus: Phycis
zinckenella Treitschke.)
Tongue well developed. Antenna of male with basal
segment enlarged and bearing a short, bluntly pointed
projection on inner side near base; shaft with sinus and
large hair and scale tuft at base, pubescent; antenna of
female simple. Labial palpus porrect; very long (the
length of head and thorax); smoothly scaled; second
segment about five times the length of third on male
and grooved to hold maxillary palpus; third segment
short on male, longer on female, acuminate, frequently
bent downward, especially on female. Maxillary palpus
of male in the form of an aigrette; of female small,
squamous. Forewing with ridge of raised scales beyond
base; 11 veins; vein 2 from before but near lower outer
angle of cell; 3 from the angle, approximately equi-
distant from 2 and 4; 4 and 5 shortly separated at base,
diverging from cell; 6 from below upper angle of cell,
straight; 8 and 9 stalked for half their lengths; 10 from
the cell, separated and divergent from the stalk of 8-9;
male without costal fold. Hind wing with vein 2 from
well before lower outer angle of cell; 3 from the angle,
appreciably shorter than 2, closely approximate to the
stalk of 4-5 at base (separated by a very short vein);
4 and 5 normally stalked (rarely weakly anastomosed
or contiguous) to about middle; 7 and 8 closely approxi-
mate for less than half their lengths from cell; cell about
one-third the length of wing on male, slightly longer on
female; discocellular vein curved. Eighth abdominal
segment of male with a pair of weak ventrolateral hair
tufts.
Male genitalia with uncus hoodlike, its apical margin
evenly rounded. Apical process of gnathos a simple,
sharp hook. ‘Transtilla incomplete and vestigial (its
elements rarely distinguishable except under high
magnification). Harpe with a strongly sclerotized,
curved, tapering, pointed arm projecting from base of
costa and as long as costa; remainder of harpe weakly
sclerotized and abruptly narrowed at middle (the shape
of harpe probably a specific character). Anellus V-
shaped; its arms rather broad, blunt and haired.
Aedeagus moderately long, stout; penis armed with
two strong cornuti. Vinculum stout; as long as greatest
width; evenly tapering to a blunt point.
Female genitalia with bursa elongate, finely scobinate,
armed with signa consisting of curved, sclerotized bands
armed with slender spines and situated near junction of
bursa and ductus bursae;a sclerotized lobe on bursa near
ductus bursae; ductus bursae short (much shorter than
bursa), tubular, sclerotized, ribbed and broadened
towards genital opening; ductus seminalis from bursa
adjacent to sclerotized lobe.
A distinct genus, not to be confused with anything
else and easily identified by its combination of antennal,
palpal, wing, and genitalic characters. Ragonot, Hulst,
and Janse list five Walker names in the generic synony-
my. These supposed genera were based on Old World,
tropical species which have all been referred as syno-
nyms of zinckenella. I have no reason to question this
synonymy but have omitted the references as I have
not been able to check the genitalia or their types. The
only species occurring in the New World is zinckenella.
The larva differs from typical phycitid larvae in that
it lacks altogether the sclerotized rings about seta IIb
of mesothorax and seta III of the eighth abdominal
segment.
207. Etiella zinckenella (Treitschke)
Figures 17, 326, 840
Phycis zinckenella Treitschke, Die Schmetterlinge von Europa,
vol. 9, pt. 1, p. 201, 1832.
Phycis etiella Treitschke, Die Schmetterlinge von Europa, vol.
10, p. 3, p. 174, 1835.—Duponchel, Histoire naturelle des
lépidoptéres, ou papillons de France, vol. 10, p. 180, 1836.—
Milliére, Iconographie et description de chenilles et lépidop-
téres inedits, vol. 1, p. 248, 1861. (Originally proposed as
new name for zinckenella.)
Pempelia Etiella zinckenella (Treitschke) Zeller, Isis von Oken,
1839, p. 179; 1846, p. 755.—Herrich-Schiffer, Systematische
Bearbeitung der Schmetterlinge von Europa, vol. 4, p.":72,
1849.
Ramphodes zinckenella (Treitschke) Guénée, Europaeorum
Microlepidopterorum index methodicus ..., p. 81, 1845.
Etiella zinckenella (Treitschke) Heinemann, Die Schmetterlinge
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 154,
1865.—Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 177,
1881.—Hulst, Phycitidae of N. Amer., p. 170, 1890; U. 8S.
Nat. Mus. Bull. 52, p. 428, 1902.—Ragonot, Monograph,
pt. 1, p. 572, 1893.—Chittenden, U. 8. Dep. Agr. Bur. Ent.
Bull. 82 (pt. 3), p. 25, 1909.—Essig, Insects of western
North America, p. 709, 1926.—Walcott, Journ. Agr. Univ.
Puerto Rico, vol. 20, no. 1, p. 476, 1936.—McDunnough,
Check list, No. 6274, 1939.
Etiella zinckenella schisticolor Zeller, Horae Soc. Ent. Rossicae,
vol. 16, p. 178, 1881.—Hyslop, U. S. Dep. Agr. Bur. Ent.
Bull. 95, pt. 6, p. 82, 1912.
Etiella villosella Hulst, Ent. Amer., vol. 3, p. 133, 1887.
Etiella schisticolor (Zeller) Ragonot, Ent. Amer., vol. 5, p. 116,
1889; Monograph, pt. 1, p. 274, 1893.—Hulst, Phycitidae
of N. Amer., p. 170, 1890.
Etiella rubribasella Hulst, Phycitidae of N. Amer., p. 170, 1890.—
Ragonot, Monograph, pt. 1, p. 572, 1893.
Forewing gray more or less lightened by white
scaling, especially in median areas; a broad white band
along costa, extending from base to or nearly to apex;
extreme costal edge dark gray to red; on occasional
specimens some red scaling at extreme base of wing and
in outer median area; transverse lines obsolete; the
antemedial line replaced by a ridge of raised scales near
basal third and extending from inner margin to top of
cell, the raised scales metallic ocherous or orange red
bordered outwardly by a broad ocherous or orange
patch; discal spots obsolete. Hind wing whitish with
a faint smoky tint, to dark smoky fuscous; the veins
100 UNITED STATES NATIONAL MUSEUM BULLETIN 207
and terminal margin darker. Alar expanse, 15-28 mm.
Genitalia as given for the genus.
TypE LOCALITIES: Sicily (zinckenella, in Hungarian
Nat. Mus., Budapest; etiella); California (schisticolor,
in BM); Colorado (villosella, in AMNH, ex Rutgers);
Florida (rubribasella, in AMNH, ex Rutgers).
Foop pPiants: Pods and seeds of various Leguminosae
(Astragalus, Cajanus, Colutea, Crotolaria, Dolichos,
Glycina, Lupinus, Phaseolus, Pisum, Vicina, Vigna).
Distrizsution: Throughout the tropical and sub-
tropical areas of the world and in the warmer temperate
regions of Hurope, Asia, and North America. The fol-
lowing American records are from specimens before me.
Unirep Statss: Florida, Archer (Mar.), Buena Vista
(May), Crescent City (Mar.), Dade City (Aug.), Eg-
mont (Apr., June), Everglade (Apr.), Jupiter (Apr.),
Lake Alfred (June, July, Nov.); Texas, Brownsville
(June, July, Aug.), Cotulla (May), Dallas (June),
Gainesville (Nov.), Kerrville, Logan (June); Arizona,
Baboquivari Mts. (June, July, Aug.), Nogales (July),
Woodruff (June); California, Alameda County (Sept.),
Garden City (Jan., Feb.), Loma Linda (June, Aug.,
Sept., Oct.), Palo Alto (Sept.), San Diego (May, June,
Sept.), San Gabriel Mts. (June), Santa Paula, upper
Ojai (uly); Washington, Pullman (May, July, Aug.),
Yakima (May, June), Walla Walla (June, Aug.),
Wenatchee (July); Idaho, Springfield (June); Nevada,
Pyramid Lake, Reno; Utah, Provo (July), Vineyard
(June); Colorado, Boulder Creek Canyon (May), Den-
ver (May); Oklahoma, Stillwater (Aug.); Rhode Island,
Weekapaugh (Aug.) ; New Hampshire, Hampton (Sept.).
Canava: Saskatchewan, Oxbow (June). Cusa: San-
tiago (Feb., June, Oct., Nov.). Purrtro Rico: Dorado
(May), Isabella (Jan.), Mayagiiez (Jan.), Palmas Aba-
jas (near Guayaman), Puerto Real (Vieques Isl., Apr.),
Rio Piedras (Apr.), San Germén (Apr.). GRENADA.
Jamaica: Newport (Feb.). México: Eldorado (Mar.),
Mexico City (Nov.), Oaxaca, Crizaba, Tehuacén (Apr.).
Frence Guiana: St. Laurent Maroni. Brazin: Santa
Catarina (July, Dec.), Sao Paulo (May). Unucuay:
Montevideo. Paraguay: Villarrica (Jan., Apr., July,
Sept., Nov.). Prrt: Angasmarca. GanApacos: Con-
way Bay (Apr.).
Presumably to be found in every Central and South
American country.
This species is of economic importance as an occa-
sional pest of cultivated beans and to American ento-
mologists is known as the “limabean pod borer.” It
has an extensive literature and has been described under
many names, having at least 13 Old World syn-
onyms. I have omitted these (for reasons given under
discussion of the genus) and listed only the American
synonyms. Some authors have treated schisticolor and
rubribasella as distinct species, others as races of
zinckenella. ‘They appear as the latter in our latest list
(McDunnough, 1939). However, they are no more
than color forms intergrading with the typical zincke-
nella and deserve no separate designation. The
accepted Old World synonymy is given in the Ragonot
Monograph (1893). The best and most complete
accounts of the life history and immature stages will be
found in the U. S. Department of Agriculture bulletins
cited here (Chittenden, 1909, and Hyslop, 1912). For
additional references the “Review of Applied Hnto-
mology” should be consulted.
Genus 54: Glyptocera
[Venational division B. Forewing with veins 4-5 connate,
smooth; hind wing with veins 4-5 distinctly stalked for over half
their lengths; 3 connate with the stalk of 4-5. Eighth abdominal
segment of male with compound tufts. Antenna of male with
shallow, spined sinus at base of shaft. Labial palpus upturned,
rough scaled beneath. Maxillary palpus squamous. Male
genitalia with sacculus of harpe produced at apex as a long, free
spine; apex of gnathos a short, stout hook; transtilla complete
but weakly sclerotized; a single strong cornutus on penis.
Female genitalia with bursa partly sclerotized, otherwise finely
spined but without definable signa; ductus bursae flattened and
partially sclerotized.]
54. Genus Glyptocera Ragonot
Glyptocera Ragonot, Ent. Amer., vol. 5, p. 114, 1889; Bull. Soc.
Ent. France, 1890, p. vii; Monograph, pt. 1, p. 209, 18938.—
Hulst, Phycitidae of N. Amer., p. 140, 1890.—Forbes,
Cornell Mem. 68, p. 621, 1923. (Type of genus: Nepho-
pteryz consobrinella Zeller.)
Tongue well developed. Antenna of male pubescent,
shaft with shallow sinus towards base containing a row
of short, toothlike spines; antenna of female simple.
Labial palpus upturned; second segment rough scaled
beneath; third segment about one-third as long as
second, acuminate. Maxillary palpus rather large,
squamous. Forewing smooth; 11 veins; vein 2 from
before but rather near lower outer angle of cell; 3 from
the angle, separated from 4—5 at base, but about half
as far from them as from 2; 4 and 5 connate; 6 from
below upper angle of cell, straight; 8 and 9 long stalked
(for slightly over two-thirds their lengths) ; 10 from the
cell, closely approximate to the stalk of 8-9 for nearly
half its length; male without costal fold. Hind wing
with vein 2 from before but near lower outer angle of
cell; 3 from the angle, connate with the stalk of 4-5;
4 and 5 distinctly stalked for over half their lengths; 7
and 8 closely approximate for a short distance from
cell; cell about half the length of wing; discocellular
vein curved. Highth abdominal segment of male with
compound ventral scale tuft.
Male genitalia with apical process of gnathos a short,
stout, simple hook. ‘Transtilla complete but very
weakly sclerotized, a simple, arched band. Harpe with
a row of fine, erect hairs from below costa near base; a
slender, short, clasperlike projection from just below
base of costa; sacculus produced at apex as a long, free,
spinelike projection. Vinculum a small V-shaped plate
with elongate, bandlike, central projection attaching
to apex of aedeagus. Penis armed with a single, strong
cornutus. Vinculum stout, longer than greatest width.
Female genitalia with bursa sclerotized over nearly
half of one surface, densely and finely spined over most
of remaining area; ductus bursae flattened, a broad
sclerotized band extending its length on ventral surface;
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 101
ductus seminalis from bursa near junction of bursa and
ductus bursae; genital opening simple.
The combination of male characters and the rather
broadly squamous maxillary palpi distinguish the
genus. The long, straight, free sacculus at once identi-
fies it and distinguishes it from the genera which follow,
and which are related to Nephopteryx and Salebria. The
weak transtilla also occurs in Meroptera and some species
of Nephopteryz. The female genitalia are similar to
those of Nephopteryr.
Glyptocera contains but the one North American
species.
208. Glyptocera consobrinella (Zeller)
Fieurss 327, 811
Nephopteryx consobrinella Zeller, Verh. zool.-bot. Ges. Wien, vol.
22, p. 548, 1872.
Glyptocera consobrinella (Zeller) Ragonot, Ent. Amer., vol. 5, p.
114, 1889; Monograph, pt. 1, p. 210, 1893.—Hulst, Phyciti-
dae of N. Amer., p. 140, 1893.—Forbes, Cornell Mem. 68,
p- 621, 1923.—McDunnough, Check list, No. 6148, 1939.
Ambesa busckella Dyar, Proc. Ent. Soe. Washington, vol. 6, p.
108, 1904.—Barnes and McDunnough, Contributions, vol.
3, p. 196, 1916.
Forewing ashy gray shaded and marked with black,
the blackish shade most conspicuous broadly bordering
the antemedial line on inner side, expanded narrowly
along inner margin to base and also broadening the
black outer border of the line at costa; basal area other-
wise pale clay color more or less shaded with pale sal-
mon especially in lower fold; on many specimens a
blotch of the same salmon shade over the middle of
inner margin; antemedial line well out beyond basal
third, slightly oblique, sharply sinuate, whitish gray
with black inner and outer bordering lines; subterminal
line outwardly rounded at middle and angled above
and below the bulge, margined inwardly and outwardly
by distinct black lines; discal spots confluent, forming
a black line along discocellular vein which expands be-
low into short black streaklets or smudges on the lower
veins; terminal dots confluent, forming a black line
along termen. Hind wing smoky white with a faint
ocherous tint; the veins not appreciably darkened; a
narrow dark line along termen. Alar expanse, 20-25
mm.
Genitalia as given for the genus. On some males in
addition to the strong cornutus there is a second much
smaller and weaker spine but this is variable and ap-
parently not a constant structure.
TYPE LOCALITIES: Texas (consobrinella, in MCZ);
Plummers Island, Md. (busckella, in USNM).
Foop piant: Viburnum, maple (Dyar and Ely
rearings).
Disrrisution: Unitep Statss: Maine, Lincolnville
(May, reared by Dyar on Viburnum), Orono, Sebec
Lake (July), Weld (July); New Hampshire, Hampton
(July); Vermont, Clarendon; Massachusetts, Framing-
ham (June), New York, Plattsburg (July), Valcour Isl.
(July); Connecticut, East River (June, July); New Jer-
sey, Basking Ridge; Pennsylvania, New Brighton (June,
July, Aug.), Pittsburgh (July); Maryland, Plummers
Isl. (May, Aug.); District of Columbia, Washington
(July, reared by Dyar from larva on maple); Tezas,
Kerrville (Mar.); Jilinois, Chicago (July). Canapa:
Ontario Trenton (July); Quebec, St. Johns (June); Nova
Scotia, Cape Breton Isl. (June); Newfoundland, Hum-
ber Mouth (Bay of Fundy, Aug.).
Presumably generally distributed over the eastern
section of the continent from Canada to Texas.
Genera 55-58: Pima to Catastia
[Venational division B. Forewing with veins 4-5 separate or
connate at base, smooth; hind wing with veins 4 and 5 anas-
tomosed for about half their lengths, discocellular vein consider-
ably extended at lower angle. Eighth abdominal segment of
male with paired tufts. Antenna of male with shallow, spined
sinus at base of shaft (also on Catastia a weak scale tuft). Labial
palpus porrect or oblique, not grooved. Maxillary palpus of
male squamous or subsquamous. Male genitalia with costa of
harpe strongly sclerotized throughout its length and slightly
produced at apex; gnathos terminating in a short, stout hook;
transtilla incomplete or absent; penis armed with two stout
cornuti (Pima, Interjectio) or a single strong cornutus (Ambesa,
Catastia). Female genitalia with bursa partially sclerotized and
sometimes (Pima, Interjectio) granulate-scobinate but without
definable signa; ductus bursae more or less sclerotized and con-
siderably broadened at genital opening.]
55. Genus Pima Hulst
Pima Hulst, Ent. Amer., vol. 4, p. 114, 1888; Phycitidae of N.
Amer., p. 164, 1890. (Type of genus: Pima fosterella Hulst.)
Epischnia Authors (not Hiibner) Ragonot, Ent. Amer., vol. 5,
p. 115, 1889; Monograph (in part) pt. 1, p. 493, 1893.—
Forbes, Cornell Mem. 68, p. 629, 1923.—McDunnough,
Canadian Ent., vol. 67, p. 176, 1935.
Tongue well developed. Antenna finely pubescent;
on male with a very slight incurvation in base of shaft
containing a row of minute, black, toothlike spines (6
to 8). Labial palpus porrect (the second segment
oblique, the third projected forward); extending at
least twice the length of the head beyond it; second
segment broadly (triangularly) scaled; third segment
as long as second. Maxillary palpus minute but rather
broadly scaled. Forewing smooth; 11 veins; vein 2
from before lower outer angle of cell; 3 from the angle,
closer to 4 than to 2; 4 and 5 separated at base; 6 from
well below upper angle of cell, straight; 8 and 9 stalked
for one-half of less than half their lengths; 10 from the
cell, more or less approximate to the stalk of 8-9; male
without costal fold. Hind wing with vein 2 from before
lower outer angle of cell; vein 3 from 4 well beyond 2
and considerably shorter than 2; 4 and 5 anastomosed
just beyond 3 for nearly half their lengths; 7 and 8
contiguous or closely approximate for a short distance
from cell; cell slightly less than half the length of wing;
discocellular vein curved, greatly extended at lower
angle (running into 4 just beyond base of vein 8).
Eighth abdominal segment of male with a pair of ven-
trolateral hair tufts.
Male genitalia with uncus hoodlike; its terminal mar-
gin broadly rounded. Apical process of gnathos a short,
stout hook. ‘Transtillaabsent. Harpe elongate, taper-
ing to bluntly pointed apex; costa broadly and strongly
102 UNITED STATES NATIONAL MUSEUM BULLETIN 207
sclerotized for the entire length of the harpe, its apex
blunt, slightly produced and usually forked; sacculus
short and narrow, its inner surface covered with dense,
spinelike hairs (probably the “‘strong spines’”’ mentioned
by Hulst in his original description of the genus). Vin-
culum a broad plate with weak lateral lobes. Aedeagus
long, moderately stout, smooth, slightly curved towards
base; penis armed with two stout cornuti. Vinculum
stout, tapering, considerably longer than greatest width.
Female genitalia with bursa stout, scobinate-granu-
late and usually with sclerotized patches or folds; ductus
bursae long (much longer than bursa), flat (ribbonlike),
sclerotized and granulate for its entire length, broadest
towards genital opening; ductus seminalis from lobe of
bursa near junction of bursa and ductus bursae. Eighth-
segment collar with middle of dorsal anterior margin
slightly produced.
Hulst erected the genus Pima for one species (foster-
ella) which he later (1890) synonymized with albiplagia-
tella. Ragonot (1889) referred fosterella and the other
congeneric American species to Hpischnia Hiibner. The
latter as defined by Ragonot (1893) is a composite of
several disparate elements and none of the species occur-
ring in the New World agrees with the type species of
Epischnia (prodromella, Hiibner). The latter has quite
different genitalia (figs. 426, 884). Its male antenna
has a deep sinus but the latter is without the row of
toothlike spines characteristic of Pima; and the third
segment of the labial palpus is much shorter (less than
half the length of the second segment). There is also
a heavy scale-and-hair tuft on the underside of the pro-
thorax. This may be only a specific character (as simi-
lar tuftings are in some other phycitid genera); but the
structure is entirely lacking on the New World species
and on the Huropean boisduvaliella Guénée, which is a
typical Pima on all characters. The venation of Pima
is like that of Epischinia (fig. 26).
The species here referred to Pima (except granitella
and parkerella) have a strikingly similar habitus; but are
individually variable in size and color, especially among
examples of western species, several of which exhibit
both pale and dark forms. The extent of the white
costal streak on forewing is also individually variable
and reliance on this and other color features has resulted
in considerable confusion in the application of names.
The most reliable specific characters are in the genitalia,
especially those of the females. The chief male differ-
ences are in the shape and size of the cornuti, the shape
of the anellus, and the configuration and width of the
apex of the sclerotized costa of harpe. These differ-
ences are trifling and also subject to some variation, es-
pecially the notching at the apex of costa of harpe.
The larvae feed in the flowers and seed pods of vari-
ous Leguminosae. From scattered reared examples in
the National Collection the species do not seem to be
confined to specific plants; but there have been no ex-
tensive and systematic rearings, and these will be needed
to clear up host relationships and to differentiate any
possible food-plant races.
Genus Pima, Species 209-216: P. boisduvaliella
to P. fulvirugella
[Forewing with strongly contrasted white costal stripe.]
209. Pima boisduvaliella (Guénée), new combination
Figures 299, 776
Epischnia boisduvaliella Guénée, Europaeorum Microlepidopter-
orum index methodicus .. . , p. 81, 1845.—Ragonot, Ent.
Monthly Mag., vol. 22, p. 23, 1885; Monograph, pt. 1, p.
518, 1893 (part).—Lafaury, Ann. Soc. Ent. France, ser. 6,
vol. 5, p. 398, 1885.—Spuler, Die Schmetterlinge Europas,
vol. 2, p. 209, 1910.—Meyrick, Revised handbook of British
Lepidoptera, p. 382, 1928.—McDunnough, Canadian Ent.,
vol. 67, p. 176, 1935 (part); Check list, No. 6251, 1939.
Forewing pale fawn on lower half shading into muddy
fawn along the lower border of the white costal stripe
where it forms a more or less distinct dark band through
the center of the wing from base to termen; no indica-
tion of transverse lines; extreme costal edge blackish
gray weakly peppered with white, this dark shade en-
croaching on the white costal stripe beyond middle;
white costal stripe extending to apex, its lower margin
edged by a fine gray-black line; discal dots at end of cell
minute, blackish, the upper one lying within the black-
ish edge of the white stripe and frequently indistinguish-
able, the lower one distinct but not conspicuous; some
faint scattered black dotting and dusting on and below
vein 1b beyond base. Hind wing pale ocherous brown,
on some specimens with a faint ocherous tint. Alar
expanse, 19-25 mm.
Male genitalia with apex of costa of harpe slightly
expended, concave or weakly notched, the upper angle
of the notch rounded, the apical margin oblique.
Cornuti both broadened for more than half their
lengths; the broad part of the shorter thorn longitudi-
nally grooved (fluted) ; the longer thorn but slightly less
than half the length of the aedeagus.
Female genitalia with bursa narrowly heart shaped; a
small rounded or oval sclerotized plate in bursa near its
middle; bursa longitudinally wrinkled, the wrinklings
weakly sclerotized; also a couple of broader, more
tortuous, sclerotized folds, the one curving about junc-
tion of bursa and ductus bursae serrate along its edge;
bursal granulations weak, scobinations in lobed area
adjacent to junction of ductus bursae fine but rather
dense; lower margin of ductus bursae at genital opening
straight, not produced.
TypE Locatity: Switzerland (location of type un-
known).
Foop pLants: Ononis, Anthyllis, Lotus, Astragalus
(European records).
DistRIBUTION: Europr. Canapa: Manitoba, Aweme
(May), Beulah (June, Aug.), Winnipeg; Saskatchewan,
Regina (June, Aug.); Alberta, Lethbridge (June).
This European species is definitely established in
North America, but most of the American references to
boisduvaliella apply to other native species. The only
American examples of the true boisduvaliella I have
seen are some ten specimens from the Canadian localities
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 103
cited above. Old World synonymns and doubtful
American references are omitted from the foregoing
literature citations. There can be no reasonable doubt
of the correctness of the European synonymy of farrella
(Curtis) or lafauriella (Constant) (their references will
be found in the Ragonot Monograph); but the status
of the supposed Asiatic variety tabulella Staudinger is
doubtful. Ragonot’s reference of albocostalialis Hulst
as a variety of boisduvaliella is obviously incorrect.
In size boisduvaliella averages appreciably smaller
than any other species of Pima. Occasional examples
have an expanse equal to that of small specimens of some
of our American species (24-25 mm.); but most speci-
mens expand 24 mm. or less, while in the other species
the average expanse is well over 25 mm.
The male genitalia are similar in all essential charac-
ters to those of albiplagiatella; but the female genitalia,
while of the same general habitus, differ in marked and
apparently consistent details—the folds of the bursa are
more weakly sclerotized, the bursa itself decidedly
smaller, and the granulations and scobinations in bursa
weaker. The contrasts are somewhat greater than
shown in the figures.
210. Pima albiplagiatella (Packard), new combination
Figures 305, 777
Myelois albiplagiatella Packard, Ann. Lyc. Nat. Hist. New York,
feud! vol. 10, p. 269, 1874.
Epischnia boisduvaliella albiplagiatella (Packard) McDunnough,
Canadian Ent., vol. 67, p. 70 (larva), p. 176 (part), 1935;
Check list, No. 6251, 1939.
Larger than boisduvaliella. The ground color on lower
half of forewing very pale fawn, paler than that of
boisduvaliella and containing little of no dark scaling,
the latter when present confined to a couple of faint
gray dots on basal third of vein 1b and some very
sparse gray scaling along inner margin near tornus; the
contrasted dark band bordering lower margin of the
white costal stripe brown rather than ruddy; a similar,
weaker, narrower, brown shade along lower fold for
most of its length; extreme costal edgepale brownish
gray, this color encroaching on the white stripe beyond
middle as in other species; lower discal dot distinguish-
able but very faint. Hind wing whitish ocherous. Alar
expanse, 26-31 mm.
Male genitalia like those of boisduvaliella. Female
genitalia similar to those of boisduvaliella but con-
sistently different in minor details. These differences
were noted in the discussion of boisdwvaliella and are
shown in the figures.
Typr Locatiry: New Hampshire (type in MCZ).
Foop puiant: Lathyrus maritima and presumably
other Leguminosae.
Distrisution: Unirep Stares: New Hampshire,
Hampton (June, July). Canapa: Nova Scotia, White
Point Beach (Queens County, July).
Typical albiplagiatella is probably generally distrib-
uted in eastern Canada and northeastern United States.
The foregoing records are from specimens whose
genitalia I have been able to examine, the Nova Scotia
record from a specimen supplied by McDunnough. He
has referred albiplagiatella as a race of boisduvaliella. It
is indeed very close to the European species, but I
believe the differences in female genitalia justify more
than racial separation.
211. Pima albiplagiatella occidentalis, new race
Figure 303
A variety occurring in the Rocky Mountain and
Pacific Coast States. Extremely variable in color and
the amount of blackish dusting on forewing. Average
specimens in Washington, Colorado, and New Mexico
with ground color very pale fawn (cream white in some
specimens) ; the dark border of the white costal stripe
ranging from pale ocherous brown to dark gray-brown;
two distinct black dots on basal third of vein 1b, enclos-
ing a white spot; the remainder of 1b more or less
streaked with black outwardly and a fine peppering of
black scales and white scales along inner margin near
tornus; costal edge brownish or blackish gray, encroach-
ing on and attenuating the white costal streak beyond
middle, and sometimes blotting it out before apex. In
southern California the paler specimens show more
blackish dusting expecially along the outer veins, and
sometimes a faint white subterminal line can be dis-
tinguished; the darkest specimens are almost a uniform
dark gray with the blackish gray dusting obliterating
all markings except the white spot on vein 1b and the
contrasted white costal streak; between these two ex-
tremes there is every intergrade in series from any given
locality; lower discal spot small, but usually distinct,
blackish. Hind wings ocherous white to dark smoky
gray. Alar expanse, 24-31 mm.
Male genitalia like those of typical albiplagiatella
except that apex of sclerotized costa of harpe is some-
what more swollen; in average specimens the outer
edge of apex is slightly concaved; one example from
Alamogordo, N. Mex., has the edge almost straight,
but there is a gradual intergradation from this to forms
with the apex as in our figure of albiplagiatella (fig.
305c). The cornuti are like those of typical albiplagia-
tella. The female genitalia are like those of eastern
albiplagiatella. A series of 24 preparations from the
various western localities exhibits no significant varia-
tion from type and nothing approaching the bursa of
boisduvaliella.
Type LocaLity: Pullman, Wash. (type in USNM,
61342).
Foop piants: Astragalus, Lathyrus.
Described from male type (C. V. Piper, collector,
May 1905) and one female paratype (J. F. G. Clarke,
May 30, 1924) from the type locality, and paratypes as
follows: One male from above Golden, Colo. (H. G.
Dyar No. 17468, May 29, 1901); one female, Beulah,
Colo. (June 21, 1900, W. D. Kearfott Collection) ; 2 male
and one female, Alamogordo, N. Mex. (May 15, 17,
1929, reared by M. W. Talbot from Astragalus wootont) ;
one female, Phoenix, Ariz. (Apr., Kunze, collector); one
male, Palm Springs, Calif.; one male and one female,
104 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Loma Linda, Calif. (May, June); one female, Clare-
mont, Calif. (Baker, no date); and one female, San
Diego, Calif. (H. G. Dyar, May 22, 1924). These are
from a series of 85 specimens in the U. S. National
Collection from the following localities: United States:
Colorado, Beulah (June), Denver, Golden (May); New
Mexico, Alamogordo, (May), Jemez Springs (Apr.,
May), Pecos; Arizona, Phoenix (Apr.). ‘Southern
Arizona”; California, Claremont, Loma Linda (Mar.,
Apr., May, June), ‘Los Angeles County” (May), Mir-
age Lake (San Bernardino County, Apr.), Olancha (Apr.,
May), Palm Springs (Mar.), San Diego (Mar., May),
“Shasta County,” ‘Sierra Nevada’; Oregon, Baker
(June); Washington, Copalis (a gray specimen with very
dark hind wings, reared under Special Survey No.
26286, Mar. 27, 1945, from Lathyrus sp.), Palouse Falls
(May), Pullman (May, June, July, Aug.), Walla Walla
(June, July), Yakima (May).
212. Pima fosterella Hulst
Fieures 300, 783
Pima fosterella Hulst, Ent. Amer., vol. 4, p. 114, 1888.
Pima albiplagiaiella Hulst (not Packard), Phycitidae of N. Amer.,
p. 164, 1890.
Epischnia albiplagiatella Ragonot (not Packard), Monograph,
pt. 1, p. 518, 1893.
Epischnia fulvirugella McDunnough (not Ragonot), Canadian
Ent., vol. 70, p. 178, 19385.
Epischnia fosterella (Hulst) MecDunnough, Check list, No. 6252,
1939.
A large, pale species similar in general appearance to
typical eastern albiplagiatella, but without any trace of
a dark shade in lower fold; the dark shade along lower
border of the white stripe also paler, a light drab brown;
a single small black spot on basal third of vein 1b, but
no white spot and seldom any further dark shading on
1b, but more or less gray dusting along outer two-thirds
of inner margin; white costal streak usually obliterated
before apex, rarely reaching apex; lower discal dot
usually well contrasted, but minute, blackish. Hind
wing whitish ocherous or pale smoky fuscous. Alar
expanse, 27-35 mm.
Male genitalia with harpe somewhat longer in pro-
portion to tegumen and uncas than in preceding species;
sclerotized costa broadened and forked at apex, the
prongs of the fork poimted. Cornuti spaced apart;
neither one appreciably flattened or ribbed towards
base; the longer slightly less than one-third the length
of aedeagus.
Female genitalia with little or no sclerotization of
bursa except immediately about junction of bursa and
ductus bursae. The extreme of sclerotization is shown
in figure 783, from an Arizona female; the female type
shows none except about the junction with ductus.
Bursa finely scobinate over entire inner surface. Duc-
tus bursae produced at apex into a projecting shield,
its apical margin variable, pointed to evenly rounded.
Typr wtocauity: Colorado (type in AMNH, ex
Rutgers).
Foop pirant: Unknown.
Distrrsution: Unirep States: Colorado, Baileys
(July), Denver (June), Durango (June), Golden (June),
Gunnison County (near Altmont, July); Arizona,
Williams; Utah, Hureka, (May), Park City (July); Mon-
tana, Miles City. Canapa: Nordegg (June).
A distinct species easily recognized by its female
genitalia. The type () at Rutgers bears only a number
label (‘43’) but is obviously a Colorado specimen and
an authentic type. A male paratype matching it is
in the National Museum. It and one other male and
two females labeled only ‘‘Colo.”? All Colorado exam-
ples are larger specimens. The Nordegg specimen (¢)
had been received from Dr. McDunnough and formed
part of the series he had treated as fulvirugella in his
1935 paper. It and a male from EHureka, Utah, show
some black scaling on the outer veins. They are super-
ficially very much like some specimens of the western
race of albiplagiatella and except for their genitalia
could easily be confused with them.
213. Pima vividella (McDunnough), new combination
Figures 302, 780
Epischnia vividella McDunnough, Canadian Ent., vol. 67, p.
179, 1935; Check list, No. 6256, 1939.
Forewing salmon pink below costal white stripe,
shading below and towards tornus into pinkish ocherous;
some gray dusting along outer two-thirds of inner
margin; a black dot, followed by an obscure white one,
on vein 1b at basal third; white costal stripe attenuated
by smoky costal scaling on its outer half and obliterated
before apex. Hind wing pale smoky with a faint
ocherous tint. Alar expanse, 27-30 mm.
Male genitalia with apex of costa of harpe forked as
in fosterella; but upper prong somewhat longer and
sharper than the lower one. Cornuti closely approxi-
mate; the longer one with flattened but not ribbed
basal part, a trifle longer than one-third of the aedeagus.
Female genitalia resemble those of fosterella except:
Bursa, proportionally smaller, with two pitted and
sclerotized patches; sclerotization at junction of bursa
and ductus bursae serrate along one edge.
Tyrer tocatity: Lethbridge, Alberta (type in Ca-
nadian Nat. Coll.).
Foop piant: Unknown.
Distrisution: Manitoba, Aweme (June); Saskatche-
wan, Saskatoon (June); Alberta, Lethbridge (June).
Also recorded by McDunnough from Beulah, Manitoba
(June) and Indian Head, Saskatchewan (July).
A good species, close to but distinct from fosterella;
easily identified by its male cornuti, female genitalia,
and salmon-colored forewings.
214. Pima albocostalialis (Hulst), new combination
Fiaures 301, 778, 779
Ephestia albocostalialis Hulst, Trans. Amer. Ent. Soc., vol. 13,
p. 64, 1886.
Epischnia albocostalis (Hulst), Phycitidae of N. Amer., p. 163,
1890. (Emended spelling).
Epischnia boisduvaliella albocostalis (Hulst) Ragonot, Mono-
graph, pt. 1, p. 520, 1893.
Epischnia albocostalialis (Hulst) McDunnough, Canadian Ent.,
vol. 67, p. 178, 1935; Check list, No. 6254, 1939.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 105
Forewing dark gray shading into very dark grayish
or blackish brown towards the white costal streak; the
latter ending just before apex; no spottings or other
markings on the wing, even the lower discal dot lost in
the dark suffusion of the wing. Hind wing whitish
at base, shading into a smoky outer area. Alar expanse,
26-34 mm.
Male genitalia with apex of costa of harpe narrow,
very slightly notched, reaching only to end of cucullus.
Harpe itself shorter in proportion to length of tegumen
and uncus than that of any preceding species except
boisduvaliella. Cornuti spaced apart as in fosterella,
the longer one about one-third the length of aedeagus.
Female genitalia with two rather large, irregularly
shaped, pitted and sclerotized patches in bursa, one at
the terminal end, the other on the left side (viewed
ventrally); bursa, otherwise membranous except for a
sclerotization about junction with ductus bursae;
ductus bursae not produced at genital opening, its
apical margin straight.
Type tocauity: California (type in AMNH, ex
Rutgers).
Foop prant: Unknown.
Distrisution: California, Claremont, ‘En route Im-
perial to Bishop” (May), Loma Linda (Mar.), River-
side (Apr.), Shasta Retreat (Siskiyou County, July);
Washington, Olympia (June), Rochester (June), Seattle,
Wenatchee (May). McDunnough also records the spe-
cies from Lethbridge, Alberta, Canada. His specimen
is undoubtedly this species, but may represent one of
the varieties that follow.
The type in the Rutgers Collection lacks antennae
and abdomen but matches otherwise the examples in
the National Collection, so there can be no question of
the application of Hulst’s name. He emended its spell-
ing to albocostalis in 1890, but gave no reason for doing
so, hence we shall probably have to perpetuate the
original barbarous spelling.
215. Pima albocostalialis subcostella (Ragonot), new combination
Epischnia subcostella Ragonot, N. Amer. Phycitidae, p. 10, 1887;
Monograph, pt. 1, p. 520, 1893.—MeDunnough, Check list,
No. 6255, 1939.
Forewing with white costal streak as in typical albo-
costalialis; below it a bordering band of blackish brown
with a lighter brown shade below it as far as lower fold;
these dark shades terminating beyond cell in what would
be the position of the antemedial line if one were pres-
ent, the outer margin of the dark shade outwardly
angled at middle; remainder of wing pale, ashy gray
with a dusting of blackish scales along outer two-thirds
of inner margin and (in some specimens) blackish
streaklets on the veins before termen; on basal third of
vein 1b a white dot preceded by a black shade. Hind
wing white; smoky shading limited to a narrow line
along termen and a very small area at apex; on the fe-
males the smoky tints slightly more extended. Alar
expanse, 25-27 mm.
Genitalia, male and female, as in typical albocostal-
talis.
Type Locaity: Utah (type in Paris Mus.).
Foop piant: Unknown.
A male and three females from Eureka, Utah (May,
June), a female from Bellevue, Washington County,
Utah (May) and a male from Pyramid Lake, Nev., are
before me. They match Ragonot’s description and
figure of subcostella in every detail. The name may
represent nothing more than a color variety of albo-
costalialis and if there were not an old name available
for it, and one that until now was supposed to repre-
sent a distinct species, I should not have named it. I
am holding subcostella as a trinomial against the possi-
bility that it may represent a valid local race.
There are also before me what appear to be two
other varieties with male genitalia identical to those of
albocostalialis:
Variety a: A large form with a reddish brown band
bordering the white costal stripe; the red-brown color
shading into ocherous fawn on lower and outer areas
of the wing; lower discal dot present, black; no white
spot or other appreciable marking on vein 1b. Hind
wing ocherous white with a pale broken line along ter-
men. Alar expanse, 34-35. Represented by three
males from Manitou, Colorado (H. G. Dyar, coll. No.
6062-6065, May 5, 1891).
Variety b: A variable variety ranging from dark gray,
suffused examples to a couple with the entire median
area of the forewing a ruddy fawn color; the white spot
is present on vein 1b, but very faint on the darker
specimens. Hind wings white to pale smoky fuscous.
Alar expanse, 26-27 mm.
The female genitalia differ rather markedly from
those of subcostella or typical albocostalialis in that the
sclerotized patches in bursa are situated on opposite
sides of that organ. This arrangement is consistent for
the females from both New Mexico and Arizona.
Distrisution: Colorado, Glenwood Springs (June);
New Mexico, Fort Wingate (June), Pecos (June); Ari-
zona, Huachuca Mts., White Mts. (Aug.).
Superficially this form looks like nothing but a color
variant of subcostella. However, if the single female
genitalic difference should hold through extended series
it will need further designation.
216. Pima fulvirugella (Ragonot), new combination
Ficure 304
Epischnia fulvirugella Ragonot, N. Amer. Phycitidae, p. 10,
1887; Monograph pt. 1, p. 521, 1893—MceDunnough, Check
list, No. 6253, 1939.
I have seen nothing that exactly matches Ragonot’s
description or figure (Monograph, pl. 16, fig. 43). A
male before me from San Francisco has a similar, at-
tenuated white subcostal streak and rather pronounced
black lining on the veins from cell. Its genitalia, how-
ever, do not match those of the type, and the moth
itself is smaller (27 mm.). Ragonot gives the alar
expanse of his type as 30 mm.
Type Locauity: California (type in Paris Mus.).
Foop pranr: Unknown.
106
Genus Pima, Species 217 and 218: P. granitella
and P. parkerella
[Forewing with white costal streak obscure or absent.]
217. Pima granitella (Ragonot), new combination
Fieures 307, 781
Epischnia granitella Ragonot, N. Amer. Phycitidae, p. 9, 1887;
Monograph, pt. 1, p. 523, 1893.—Hulst, Phycitidae of N.
Amer., p. 162, 1890.—McDunnough, Check list, No. 6258,
1939.
Megasis piperella Dyar, Proc. Ent. Soc. Washington, vol. 6, p.
110, 1904.
Forewing gray finely powdered with white, the white
dusting concentrated between the veins; the latter finely
lined with blackish brown giving the wing (to the naked
eye) a longitudinally dark-lined, over-all pale bluish
gray appearance; on some specimens a trace of the usual
costal white strip as a narrow white line along top of
cell for a short distance from base; the lower blackish
discal spot usually distinct but occasionally obliter-
ated; on a few specimens the subterminal line slightly
indicated by interruptions in the blackish streaks on the
veins, but, otherwise, transverse lines absent. Hind
wing pale brown, unicolorous, except for a narrow, very
slightly darker line along termen. Alar expanse, 22-
23 mm.
Male genitalia with sclerotized costa of harpe broad-
ened, slightly notched and produced at apex. Cornuti
lying one before the other, of nearly equal length, not
(or but very slightly) broadened and not flattened at
base; the longer one slightly less than one-third the
length of aedeagus. Female genitalia similar to those
of albocostalialis except sclerotized patches in bursa,
narrower, more elongate.
TypE Locatity: California (granitella, in Paris Mus.);
Pullman, Wash. (piperella, in USNM).
Foon piant: Crotolaria.
Distrisution: Colorado, Glenwood Springs (Apr.,
May); Utah, Eureka (May, June), Richfield (May),
Stockton (May); New Mexico, Jemez Springs (Apr.,
May); Arizona, Dewey (Apr.); Nevada, Baker (May);
California, Argus Mts. (May), Crows Landing (May),
Los Angeles County (May); Washington, Pullman,
Walla Walla (May).
A distinct species, easily recognized by its wing pat-
tern.
218. Pima parkerella (Schaus), new combination
Figures 306, 782
Epischnia parkerella Schaus, Proc. Ent. Soc. Washington, vol. 26,
p. 196, 1924.—MecDunnough, Check list, No. 6250, 1939.
Forewing olive buff, costal and inner margins whitish
more or less overlaid with a fine peppering of blackish
scales; on some specimens a scattering of similar dark
scales in the central area and near outer margin along
some of the veins; no contrasted white costal strip;
transverse lines well marked on most specimens, white,
rather broad (absent on one example before me); the
antimedial line outwardly oblique from costa to inner
margin, nearly straight, at most with a slight bend at
UNITED STATES NATIONAL MUSEUM BULLETIN 207
vein 1b; subterminal line inwardly oblique from costa
to inner margin, with a decided notch at lower fold, in-
wardly margined by a faint dark shade; discal spots usu-
ally obsolete, when distinguishable very faint. Hind
wing very pale fuscous with a narrow dark line along
termen; on the paler examples the hind wing has a slight
ocherous-gray tint. Alar expanse, 30-35 mm.
Male genitalia somewhat stouter than those of pre-
ceding species. Sclerotized costa of harpe not appre-
ciably broadened at apex nor projecting beyond apex
of cucullus; very slightly if any notched at apex. Cor-
nuti lying close to each other, the apex of one projecting
slightly beyond that of the other; about one-third as
long as aedeagus.
Female genitalia with bursa minutely scobinate, at
left posterior angle developed as a protruding lobe with
thickened membrane; ductus bursae developed into a
projecting shield at genital opening.
Typ Locatity: Bozeman, Mont. (type in USNM).
Foop priant: “Locoweed”’ (Astragalus).
Represented in the National Collection by nine ex-
amples from the type locality (July). The species is
easily recognized by its forewing markings. It is the
only Pima with anything approaching distinct trans-
verse lines.
56. Interjectio, new genus
Tyrer oF GENus: Ambesa columbiella McDunnough.
Characters as in Pima except: Labial palpus extend-
ing little more than the length of the head beyond it,
third segment less than half as long as second; apical
process of gnathos a broad, flanged plate terminating in
a short slender hook; harpe short, hardly extending be-
yond apical margin of uncus; vinculum little longer than
greatest width; genitalia generally broader andmore
chunky than those of Pima; ductus bursae of female
short, little, if any, longer than bursa.
This genus is intermediate between Pima and Ambesa.
From the latter it differs chiefly in having at least two
cornuti on the penis and the granulations of the ductus
bursae uninterrupted for its entire length.
219. Interjectio denticulella (Ragonot), new combination
Fieure 308
Pristophora denticulella Ragonot, N. Amer. Physicidae, p. 6, 1887.
Ambesa lallatalis Authors not Hulst (in part) Ragonot, Ent,
Amer., vol. 5, p. 115, 1889; Monograph, pt. 1, p. 241, 1893.
Ambesa denticulella (Ragonot) McDunnough, Canadian Ent.,
vol. 67, p. 174, 1985; Check list, No. 6158, 1939.
Forewing white dusted and marked with black,
making the ground color (of fresh specimens) white
with a very faint bluish tint; the dark markings strongly
contrasted, consisting of streaks and spots chiefly indi-
cating broken margins of the antemedial and postmedial
lines; the antemedial line itself obscure except between
lower fold and inner margin when it is an inwardly
curved white line preceded by a black spot and followed
by a thin black marginal line; above, indicated only by
its outwardly oblique, broken outer margin, consisting
of a short black line from costa and black streaklets on
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 107
upper and lower veins of cell; subterminal line indicated
by an oblique, deeply dentate, broken, black inner
border; black discal dots at end of cell distinct, the
upper minute, the lower a short but conspicuous streak;
a row of black dots along termen; the blackish shading
otherwise consists of very fine dusting between the
veins, supplemented by some faint brownish shading
in the folds. Hind wing whitish brown, the veins very
slightly dakened; a pale brown line along termen. Alar
expanse, 32-34 mm.
Male genitalia with apical process of gnathos with
the lateral angles of the flanged plate incurved and
broadly rounded. Apex of cucullus not projecting
beyond sclerotized costa of harpe. Cornuti two stout
thorns, less than half as long as aedeagus; the latter
short, stout, straight.
Type tocauity: North America, but otherwise
unspecified (type in Paris Mus.).
Foop pLant: Unknown.
Distrisution: Unirep States: Washington, Wenat-
chee (May). Canapa: British Columbia, Keremeos
(June).
This species was removed by McDunnough (1935)
from the synonymy of Ambesa lallatalis (Hulst) where
it was placed originally by Ragonot (1889), presumably
on the basis of specimens of supposed Jallatalis sent him
by Hulst. The latter species was misidentified by
Hulst himself as well as by later authors.
220. Interjectio columbiella (McDunnough), new combination
Fiaure 784
Ambesa columbiella McDunnough, Canadian Ent., vol. 67, p.
175, 1935; Check list, No. 6159, 1939.
Similar to denticulella but without the well-contrasted
dark markings of that species; the black spot on inner
margin, preceding the antemedial line, entirely lacking
and the dark markings themselves more brown than
blackish, except for the small, narrow black lower discal
dot, an occasional black dot on vein 1b near base, and
some blackish dots along termen; the brown markings
limited to streakings on the veins and (where the trans-
verse lines are distinguishable) to a narrow dark shade
along inner margin of the subterminal line. Hind
wings as in denticulella. Alar expanse, 23-34 mm.
The species averages smaller than denticulella, espe-
cially the females, which are even smaller than any of
the males before me.
Male genitalia similar to those of denticulella except
cornuti distinctly more slender. Female genitalia
(figured from a Pullman, Wash., specimen from a series
associated with males of the same locality) with
sclerotized ductus bursae projecting as a short shield
at genital opening.
Type Loca.ity: Oliver, British Columbia (type in
Canadian Nat. Coll.)
Foop prant: Unknown.
Distrizution: Unirep States: Utah, Stockton
(June); Washington, Pullman (May, June), Yakima
(June). Canava: British Columbia, Oliver (June);
Alberta, Lethbridge (July).
221. Interjectio ruderella (Ragonot), new combination
Fieurn 785
Epischnia ruderella Ragonot, N. Amer. Phycitidae, p. 9, 1887;
Monograph, pt. 1, p. 514, 1893.—Hulst, Phycitidae of N.
or p. 162, 1890.—McDunnough, Check list, No. 6249,
This species is known only from the female type,
which, from Ragonot’s description and figure, must be
very close to some of the small females of columbiella,
especially paler examples from Pullman, Wash., except
that the lower discal spot is a more contrasted black
streaklet. Alar expanse, 24 mm.
The genitalia show a broader and stronger sclerotiza-
tion of the ductus bursae at its junction with bursa than
typical columbiella; but this character is approached in
Utah examples of the latter species.
Type Locauity: “North America” [given as “without
doubt from California” in the Ragonot Monograph]
(type in Paris Mus.).
Foop pLant: Unknown.
222. Interjectio niviella (Hulst), new combination
Figure 309
Iipographis niviella Hulst, Ent. Amer., vol. 4, p. 117, 1888.
Ambesa niviella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115,
1889; Monograph, pt. 1, p. 241, 1893.—Hulst, Phycitidae
of N. Amer., p. 141, 1890.—McDunnough, Check list, No.
6157, 1939.
Forewing chalk white; a large black patch bordering
inner margin of antemedial line and extending from
inner margin to top of cell; antemedial line faint but
complete, oblique from costa to lower vein of cell, thence
concave to inner margin, bordered outwardly by a black
line, which is more or less broken into spots on upper
half; black streaks and wedges on the outer veins indi-
cating the deeply notched, otherwise obscure sub-
terminal line; both discal spots conspicuous, black; a
row of short black streaklets along outer margin; on
some specimens smears of a faint, pale, ocherous brown
tint towards apex; a fine brown line along termen.
Alar expanse, 25-28 mm.
Male genitalia with apical process of gnathos a broad-
ly crescentiform plate with the lateral angles slightly
produced; the terminal hook short and slender. Harpe
with apex of cucullus projecting beyond the sclerotized
costa. Cornuti a cluster of several short, stubby
thorns. Aedeagus rather slender, sinuously curved.
Type nocauity: Colorado (type in AMNH, ex-
Rutgers).
Foop prant: Unknown.
Distrisution: Unirep States: Colorado; Iowa,
Sioux City. Canapa: Manitoba, Aweme (July), Cart-
wright, Winnipeg.
The type is a female without abdomen. I have seen
but one other female and it too was without abdomen,
so the female genitalia could not be studied. These
specimens matched the males in every detail of color
and maculation.
108 UNITED STATES NATIONAL MUSEUM BULLETIN 207
57. Genus Ambesa Grote
Ambesa Grote, N. Amer. Ent., vol. 1, p. 98, 1880.—Hulst, Phy-
citidae of N. Amer., p. 141, 1890.—Ragonot, Monograph,
pt. 1, p. 237, 1893. (Type of genus: Ambesa laetella Grote.)
Tongue well developed. Antenna as in Pima and
Interjectio. Labial palpus oblique, extending above
vertex; second segment roughly and rather broadly
scaled; third segment not defected forward, about one-
third as long as second, acuminate. Maxillary palpus
minute, filiform. Forewing smooth; venation as in
Pima. Hind wing with vein 3 from the angle of the
cell (but separated from discocellular vein by a short
spur), longer in proportion to 2 than in Pima, 4 and 5
anastomosed for nearly half their lengths, 7 and 8 closely
approximate for a short distance from cell; cell one-
third the length of wing; discocellular vein curved, ex-
tended at lower angle but not so far asin Pima. Highth
abdominal segment of male simple or (/aetella) with a
pair of ventrolateral hair tufts.
Male genitalia similar to those of Interjectio except:
Harpe longer in proportion to combined tegumen and
uncus; its sclerotized costa with upper angle at apex
produced into a sharp point; sacculus finely haired, not
with coarse spinelike hairs of Pima or Interjectio.
Anellus a simple plate without lateral projections.
Penis armed with a single, long, stout cornutus; over
half as long as aedeagus. Vinculum as long as greatest
width.
Female genitalia with bursa unsclerotized except at
junction of bursa and ductus bursae; ductus bursae
sclerotized except for narrow space at middle, broad-
ened at genital opening.
A North American genus close to both Pima and
Interjectio, distinguished from both by its palpi, weakly
haired sacculus, single strong cornutus, and the inter-
rupted sclerotization of ductus bursae. The known
species occur only in the western parts of the United
States and Canada.
223. Ambesa laetella Grote
Fieurus 18, 310, 790
Ambesa laetella Grote, N. Amer. Ent., vol. 1, p. 98, 1880.—Hulst,
Phycitidae of N. Amer., p. 141, 1890.—Ragonot, Mono-
graph, pt. 1, p. 240, 1893.—McDunnough, Check list, No.
6156, 1939.
A brilliant, brightly colored species with clearly
marked pattern; forewing markings a rich red-brown
against a surrounding suffusion of ashy white; on mid-
costa a broad, elongate red-brown patch extending from
outer margin of antemedial line to end of cell and from
costal edge well into cell, shading into a blackish line on
its lower margin, and bordered on its outer edge by an
angled black discal mark formed of the fused discal
spots; upper half of basal area, median area from the
brown midcostal patch to vein 1b, and most of the area
beyond as far as subterminal line, ash white; a brownish
suffusion, paler than the costal patch, along inner mar-
gin below vein 1b, on some specimens intensified into a
constrastingly darkened patch above middle of inner
margin; the transverse lines well separated, clearly
marked ; antemedial line narrow, angled, white, bordered
outwardly by a narrow blackish brown line; subterminal
line sinuate, narrow, white, preceded and followed by
blackish brown costal spots, the inner one continued as
a brown inner bordering line, the outer expanding below
into a pale brownish suffusion filling tornal area; sub-
apical area dusted with white; terminal dots fused into
a black line along terminal edge. Hind wing pale
smoky fuscous with a very faint yellowish tint. Alar
expanse, 28-30 mm.
Male genitalia with apical process of gnathos con-
tinued below into a pair of narrow, converging, sclero-
tized lobes. ‘Transtilla absent. Harpe with cucullus
narrow; sclerotized costa terminating im a short, sharp
point at apex; an erect clasper arising from below costa
near base. ‘Terminal margin of vinculum narrowly
rounded. Highth abdominal segment of male with
paired tufts.
Female genitalia with terminal, sclerotized portion
of ductus bursae abruptly widened and transversely
wrinkled toward genital opening. Highth-segment col-
lar narrowly sclerotized.
Typs Locatity: Colorado (type in BM).
_ Foop prant: Unknown.
Disrrrpurion: Unirep States: Arizona, White Mts.
(July, Aug.), Williams; Colorado, Fort Collins (Aug.),
Glenwood Springs (Aug.); Montana, Bozeman (Aug.),
Cut Bank (July); Utah, Provo, ‘‘South Utah” (July);
Nevada, Verdi (June); California, Inyo County (June),
San Bernardino Mts. (July, Aug.), Sierra Nevada Mts.;
Washington, Pullman (June, July, Aug.). CANADA:
Manitoba, Aweme (Aug.); Alberta, Calgary (July).
An easily recognized species and one of the most
beautiful of the American Phycitidae.
224, Ambesa walsinghami (Ragonot)
Figures 312, 791, 792
Pristophora walsingham: Ragonot, N. Amer. Phycitidae, p. 6,
1887.
Ambesa walsinghami (Ragonot) Hulst, Phycitidae of N. Amer.,
p. 142, 1890.—Ragonot, Monograph, pt. 1, p. 239, 1893.—
McDunnough, Check list, No. 6153, 1939.
Ambesa monodon Dyar, Ins. Insc. Menstr., vol. 1, p. 34, 1913.—
MeDunnough, Check list, No. 6154, 1939. (New synonymy.)
Forewing blackish gray faintly tinted with purplish
fuscous over the lower half of the wing; on costal half
from base to antemedial line, white faintly streaked
along the veins with black, the outer margin of the
whitish area oblique from lower angle of cell to costa.
near beginning of subterminal line and, on some fresh
specimens, bordered outwardly by a transverse blackish
darkening of the ground color; some further ashy white
dusting in the subapical area beyond the subterminal
line; transverse lines well contrasted; the white ante-
medial line distinct only from inner margin to cell,
concave to lower fold thence inwardly oblique to the
cell, beyond which it is lost in the white dusting above,
its outer black margin begins as a black, oblique streak
from costa and continues outwardly as a fine black line
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 109
along upper edge of cell as far as discocellular vein,
whence it loops backward along the lower vein of cell
and thence along outer edge of the white line to inner
margin; subterminal line, parallel with termen, sinuate,
white, bordered inwardly by a black line and outwardly
by a narrow dark shade, both borders enlarged at
costa into contrasted black spots; discal dots not dis-
tinguishable; along termen a row of more or less con-
fluent black dots. Hind wings varying from smoky
white to pale brownish, the veins faintly darkened.
Alar expanse, 19-28 mm.
Male genitalia with uncas broadly elongate, the side
margins parallel. Apical process of gnathos without
pendant lobes. Transtilla present, incomplete. Harpe
with cucullus moderately broad; apex of sclerotized
costa produced into a long point at upper (outer) angle;
no erect clasper. Eighth abdominal segment of male
without tufts.
Female genitalia with terminal sclerotized portion of
ductus bursae gradually broadened to genital opening,
its terminal margin deeply concaved. LHighth-segment
collar broadly sclerotized, ventrally fused.
Tyre Locauiries: California (walsinghami, in Paris
Mus.); Stockton, Utah (monodon, in USNM).
Foop piant: Prunus virginiana melanocarpa.
Disrrisution: California, Cloverdale (June), Deer
Park Spring (Lake Tahoe), Plumas County (July, Aug.) ;
Utah, Bellevue (May), Eureka (June, July), Provo
(July), Stockton (July); Washington, Kamiack Butte
(May).
Dyar’s type of monodon is a small female whose
genitalia (fig. 792) show minor differences from typical
California specimens, but a series from Utah shows all
intergradations between the extremes displayed in the
figures. The food plant record is from a series reared
by J. F. G. Clarke in 1934 at Kamiack Butte, Wash.,
which I then identified as mirabella Dyar. Their
abdomens are distinctly gray and their hind wings (es-
pecially those of the females) are brownish. In exam-
ples of typical walsinghami from the coastal region of
California (Cloverdale) the hind wings are whitish and
the abdomens gray or grayish ocherous.
225. Ambesa walsinghami mirabella Dyar, new status
Fieurts 313, 793
Ambesa mirabella Dyar, Proce. Ent. Soc. Washington, vol. 10,
p. 59, 1908.—Essig, Insects of western North America, p.
709, 1826.—McDunnough, Check list, No. 6155, 1939.
Not distinguishable from typical walsinghami in color
or maculation of forewing. The hind wing of a dis-
tinctly ocherous tint and the abdomen ocherous over
the entire upper surface. Alar expanse, 25-28 mm.
The male genitalia show only a trifling difference
from those of typical walsinghami in forking of apex of
costa (fig. 313). Female genitalia with a shallower con-
cavity in terminal margin of the projecting ductus bur-
sae at genital opening. Figure 793 shows the extreme of
reduction in the concavity. Other examples of mira-
bella show intergrading approaches to the deep con-
cavity of walsinghami.
TYPE Locauity: San Diego, Calif. (type in USNM).
Foon piant: “Plum.”
Distrisution: California, Atascadero (San Luis
Obispo County, July), Camp Baldy (San Bernardino
Mts., July), Cloudburst Canyon (Los Angeles County,
July), Mount Lowe (July), Pasadena, Pine Valley (San
Diego County), San Diego (July). According to Essig
(1926) the larvae were taken in large numbers on prune
trees at Hopeland, Calif.
The name mirabella represents, at most, only a south-
ern California race of walsinghami.
226. Ambesa lallatalis (Hulst)
Fiaures 311, 789
Neophopteryz lallatalis Hulst, Trans. Amer. Ent. Soc., vol. 13,
p. 161, 1886.
Ambesa lallatalis (Hulst) Phycitidae of N. Amer., p. 142, 1890
(part).
Forewing white heavily dusted with blackish giving
the entire wing a pale ashy gray appearance; markings
as in walsinghami but much fainter; the transverse lines
obscure; antemedial line indicated by its broken outer
blackish border which has the same outer loop over the
cell so characteristic of walsinghami but much fainter
and often interrupted; on better marked examples a
whitish crescent on inner margin indicates the base of
the normal antemedial line; beyond this, dark lines
extend along vein 1b and the edge of inner margin as
far as base of subterminal line, defining a narrow oval
patch along inner margin; subterminal line sinuate, de-
fined by its black, dentate inner border, the latter inter-
rupted by a rather broad pale shade extending along
outer half of lower fold; the outer area (beyond sub-
terminal line) and the costal area at base also paler than
remainder of wing; a thin blackish line along outer mar-
gin. Hind wing white to pale smoky brown. Eighth
abdominal segment of male without tufts. Alar ex-
panse, 26-30 mm.
Male genitalia with apical process of gnathos a simple,
narrow, moderately long hook. Transtilla absent.
Harpe without clasper; cucullus moderately broad;
sclerotized costa terminating in a very shortly project-
ing point at apex. Anellus a narrow, elongate, plate.
Cornutus almost as long as aedeagus. Vinculum with
terminal margin broad.
Female genitalia with terminal margin of ductus
bursae at genital margin scobinate, convex, slightly
notched in the middle (giving the projecting lower sur-
face of the ductus a bilobed appearance). Highth-seg-
ment collar broadly sclerotized, wrinkled at lateral mar-
gins, not ventrally fused.
Typr tocauity: Nevada (type in AMNH, ex Rut-
gers).
Foop piant: Unknown.
In addition to the female type in the Rutgers Collec-
tion, I have seen only four other authentic specimens, a
series of three males and one female from Bellevue,
Washington County, Utah, in the National Collection,
collected by G. P. Engelhardt, June 21, 1917. The
110 UNITED STATES NATIONAL MUSEUM BULLETIN 207
genitalia of the Utah female agree in every detail with
those of the type.
The Ragonot and Dyar references to Jallatalis have
been omitted from the above synomy as they apply to
other species. For comments on the misapplication of
Hulst’s name see under Interjectio denticulella (p. 106)
and Phobus brucei (p. 138).
58. Genus Catastia Hitibner
Catastia Hiibner, Verzeichniss bekannter Schmetterlinge, p. 372,
1825.—Heinemann, Die Schmetterlinge Deutschlands und
der Schweiz, Abt. 2, vol. 1, pt. 2, p. 164, 1865.—Ragonot,
Monograph, pt. 1, p. 479, 1893.—Spuler, Die Schmetterlinge
Europas, vol. 2, p. 210, 1910.—Hemming, Hubner, vol. 2, p.
168, 1937.—Janse, Journ. Ent. Soc. South Africa, vol 7, p.
13, 1944 (description and figures). (Type of genus: Noctua
marginea Schiffermiiller; figs. 314, 796.)
Tongue well developed. Antenna finely pubescent;
on male with a shallow sinus in base of shaft containing
a row of minute black teeth and a short, weak scale
tuft. Labial palpus oblique, not extending above
vertex; broadly scaled, the scales tightly appressed
(except on ineorruscella and actualis); third segment
over one-third the length of second,‘ usually projected
forward and partially concealed in the scaling of second
segment, acuminate. Maxillary palpus squamous
(broadly scaled). Forewing smooth; 11 veins; vein 2
from before lower outer angle of cell; 3 from the angle,
closer to 4 than to 2; 4 and 5 separated at base; 6 from
below upper angle of cell, straight; 8 and 9 stalked for
nearly half their lengths; 10 from the cell; male without
costal fold. Hind wing as in Ambesa (except that in
the European marginea vein 3 is slightly longer in
proportion to 2; this species barely coming within our
venational group B). Eighth abdominal segment of
male with a pair of ventrolateral hair tufts.
Male genitalic characters as in Ambesa except:
Harpe not so long in proportion to combined length of
tegumen and uncus; cucullus narrow on all species.
Transtilla incomplete, but its elements more strongly
sclerotized. Vinculum as long or a trifle longer (mar-
ginea) than greatest width, stout.
Female genitalia with lower surface of ductus bursae
towards genital opening unsclerotized or very weakly
sclerotized (except in marginea); a pair of narrow,
elongate plates on inner dorsal surface of ductus bursae
at genital opening; otherwise as in Ambesa.
This genus is very close to Ambesa, differing chiefly
in its shorter labial palpus with deflected third segment;
its squamous maxillary palpus; stronger sclerotization
of the elements of transtilla; the presence of the two
elongate sclerotized plates in the ductus bursae towards
genital opening; and the slight scale tuft in the sinus
of the male antennal shaft.
The European type of the genus (figs. 314, 796) dif-
fers from our American species in having the lower
surface of the ductus bursae sclerotized and produced
“4 Denuded example of palps of the type species (marginea)
show the third segment half again as long as the figure (52b) in
Janse’s 1944 paper.
at genital opening, the pointed projection from apex
of costa a trifle longer, the vinculum narrowly rounded
at its extremity, and vein 3 of hind wing a trifle longer
in relation to 2. These differences, however, are more
specific than generic in character and do not seem to
justify a separate generic designation for our American
species, despite the obvious likenesses in structure.
The life history of none of the species is known.
227. Catastia bistriatella (Hulst), new combination
Figures 316, 797
Pyla bistriatella Hulst, Canadian Ent., vol. 27, p. 54, 1895.—
McDunnough, Check list, No. 6238, 1939.
Head, thorax, and forewing black with a deep greenish
blue iridescence; the forewing crossed by two rather
broad white lines; the antemedial line slightly oblique,
nearly straight; the subterminal line set well back from
termen, somewhat wavy. Hind wing a uniform, glossy
black-brown. Labial palpus not reaching to vertex
(shorter than the palpi of the other species of the
genus) ; second segment broadly scaled, the scales flatly
appressed ; third segment shorter than that of any other
species of the genus, almost completely hidden in the
scaling of second segment. Alar expanse, 23-25 mm.
Male genitalia with projecting spine at apex of
sclerotized costa very short. Terminal margin of
vinculum moderately broad, slightly produced at the
lateral edges, very slightly convex. Female genitalia
with ventral surface of ductus bursae at genital opening
not sclerotized; bursa copulatrix with an irregular,
lined, weakly sclerotized patch towards anterior end
(probably an individual rather than a specific charac-
ter).
TYPE LOcALITy: Yosemite, Calif. (type in AMNH,
ex Rutgers).
Foop prant: Unknown.
Distrizution: California, Humphreys Basin (Fresno
County, Aug.), Yosemite.
A striking, easily recognized species. Hulst is in
error in stating that the maxillary palpi are not scale
tufted. They are squamous like those of the other
cogeneric species but are difficult to see behind the
heavily scaled labial palpi. The genitalia of the male
type agree in every detail with those of the Humphreys
Basin male figured.
228. Catastia incorruscella (Hulst), new combination
Figures 315, 795
Pyla incorruscella Hulst, Canadian Ent., vol. 27, p. 55, 1895.—
McDunnough, Check list, No. 6240, 1939.
Fore and hind wings a deep, blackish brown, some-
what lustrous but without metallic iridescence; trans-
verse lines similar to those of bistriatella, but thinner and
a dull ocherous white. On thorax and palpi a scatter-
ing of whitish scales; the scaling on second segment of
labial palpus slightly roughened; third segment about
the same proportional length as on marginea and actu-
alis, longer than that of bistriatella. Alar expanse, 20-
22 mm.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE Gla!
Male genitalia differ from those of bistriatella only in
insignificant details. Female genitalia having bursa
without sclerotized patch. However, a sclerotized
patch similar to that shown for bistriatella (fig. 797) is
present in the bursa of a female in the National Collec-
tion from Slate Peak, Wash. On this specimen there
is also a weak sclerotization of the ventral surface of the
ductus bursae at genital opening.
Type Locauity: Colorado (type in AMNH, ex Rut-
gers).
Foop prant: Unknown.
Distrisution: Colorado; Arizona, White Mountains
(June); Washington, Slate Peak (Okanogan County,
Aug.).
I have seen no Colorado examples except the male
type, but the Arizona locality is represented in the
National Collection by a series of males and females in
excellent condition (collected by Grace M. and John L.
Sperry at Colter’s Ranch in the White Mountains, June
17-18, 1937). Their male genitalia are like those of the
type.
229. Catastia actualis (Hulst), new combination
Ficures 317, 794
Nephopteryx actualis Hulst, Trans. Amer. Ent. Soc., vol. 13, p.
161, 1886.
Dioryctria actualis (Hulst), Phycitidae of N. Amer., p. 135,
1890.—Ragonot, Monograph, pt. 1, p. 203, 1893.—MeDun-
nough, Check list, No. 6132, 1939.
Monoptilota actualis (Hulst), U. S. Nat. Mus. Bull. 52, p. 421,
1902 (this combination the result of an accidental misplace-
ment).
Forewing with basal and outer areas ocherous, the
basal area more or less dusted with ashy fuscous, espe-
cially towards inner margin, the outer area with some
blackish streaks on the veins; median area (between
the transverse lines) ashy fuscous, the whitish dusting
concentrated into a pale suffusion transversely across
the wing from costa before subterminal line to or almost
to inner margin at base of antemedial line; antemedial
line white, oblique, notched below cell and more or less
dentate above; bordered outwardly by a diffused black-
ish smudge at costa and below cell by a blackish line,
and preceded on inner margin by a black patch (except
on California examples); a blackish spot precedes and
one usually follows the sinuate subterminal white line
on costa, the inner costal spot continuing as a blackish
bordering line to inner margin; discal dots distinct,
separated, black; a row of blackish dots along termen.
Hind wing ocherous brown, darker brown on most fe-
males; a dark line along termen and some darkening of
the veins. Alar expanse, 24-28 mm.
Male genitalia differing only in insignificant details
from those of bistriatella and incorruscella.
Female genitalia with sclerotized wrinklings of bursa
more extended than in other species of the genus; duc-
tus bursae weakly granulate towards genital opening
(differences of little or no significance).
Type tocauity: Colorado (type in AMNH, ex Rut-
gers).
Foop piant: Unknown.
DistrisuTion: Unirep Strats: Colorado, Denver
(June), Florissant (July), Gunnison County (near Al-
mont, June, July), Platt Canyon (July), no specific lo-
cality (June) ; Nevada; Secret Pass (Elko County, June);
California, Deer Park Springs (Lake Tahoe, July), Tuo-
lumne Meadows (July); Washington, Easton, Walla
Walla (June). Canapva: Manitoba, Aweme (July);
British Columbia, Blue Lake (west of Lytton, Aug.).
The foregoing description was drawn from typical
Colorado examples. The British Columbia and Wash-
ington specimens before me are much darker, the ocher-
ous coloration of forewing replaced by deep brown and
the general color of the wing a suffused blackish fuscous
with only the white transverse lines and some whitish
dusting in the median area contrasted; the hind wings
deep brown to blackish brown with no ocherous tinting.
This is merely a color form, not a race, and probably
represents nothing more than individual response to a
moist condition. The Nevada record, cited above, is
from a single male in the Canadian National Collection
(Grace H. and John L. Sperry, collectors). It also rep-
resents a divergent color form with very dark brown
hind wing, a dark ground color on forewing and strongly
marked, white, transverse lines. The type in the Rut-
gers Collection is a male. Its genitalia agree in every
detail with those of our pale and dark forms.
Genera 59-64: Immyrla to Quasisalebria
[Venational division B. Veins 4 and 5 of forewing slightly
separated at base, in Oreana connate or, rarely, very shortly
stalked; vein 6 straight. Hind wing with veins 7-8 approximate
or very shortly anastomosed beyond cell. Antenna of male with
sinus and scale tuft at base of shaft. Labial palpus erect or
oblique, smooth scaled; on male second segment grooved to hold
maxillary palpus. Maxillary palpus of male in the form of an
aigrette. Male genitalia with transtilla absent, or incomplete
and with its elements rudimentary; penis normally armed with
a single strong cornutus (except Oreana which has several cornuti
and Quasisalebria which has none). Female genitalia with bursa
more or less finely and densely spined or scobinate.]
59. Genus Immyrla Dyar
Immyrla Dyar, Journ. New York Ent. Soc., vol. 14, p. 108,
1906.—Forbes, Cornell Mem. 68, p. 627, 1923. (Type of
genus: Immyrla nigrovittella Dyar.)
Tongue well developed. Antenna pubescent; a sinus
and large-scale tuft on base of shaft of male. Labial
palpus erect; reaching above vertex; smoothly scaled;
second segment on male grooved to hold the maxillary
palpus, appressed to face; third segment minute (less
than one-fifth the length of second and hidden in the
scaling of the latter on male, a trifle longer and partially
exposed on female), acuminate. Maxillary palpus of
male in the form of an aigrette; of female squamous.
Forewing with subbasal scale ridge; 11 veins; vein 2
from before lower outer angle of cell; 3 from the angle,
closer to 4 than to 2; 4 and 5 approximate for a short
distance from their bases; 6 from below upper angle of
cell, straight; 8 and 9 stalked for over half their lengths;
10 from the cell, approximate for a short distance to the
112
stalk of 8-9; male without costal fold. Hind wing with
vein 2 from before lower outer angle of cell; 3 from 4 well
beyond 2, in some specimens (fig. 19) connected with
discocellular by a short spur before its separation from
4; 4 and 5 anastomosed for less than half their lengths;
7 and 8 anastomosed for a short distance beyond cell;
cell less than half the length of wing; discocellular vein
curved and considerably extended at lower angle.
Highth abdominal segment of male with compound
ventral scale tufts.
Male genitalia as in Catastia except transtilla com-
pletely absent, a fine brush of long hairs arismg from
inner surface of harpe along lower edge of basal half of
the sclerotized costa (as in Meroptera).
Female genitalia with bursa copulatrix finely and
densely spined; ductus bursae flattened, granulated,
inbent at middle; not longer than bursa, its lateral
margins strongly and broadly sclerotized towards
genital opening, opening deeply concave; ductus semi-
nalis from a thickened (but not sclerotized) lobe of
bursa, near junction of bursa and ductus bursae.
This and the five genera following form a group
closely allied to Salebria Hiibner. All have grooved
and erect or oblique male labial palpi, the male maxil-
lary palpus in the form of an aigrette, a scale tuft in
sinus on base of shaft of male antenna, no transtilla or
only the greatly reduced rudiments of one, and (except
for Oreana and Quasisalebria) a single, long, strong
cornutus on penis. Jmmyrla is distinguished from all
nearly related genera by the raised scale ridge on fore-
wing. Ortholepis also has this character, but the costal
sclerotization of its harpe is much weaker, and it has a
complete transtilla.
230. Immyrla nigrovittella Dyar
Figures 19, 318, 798
Immyrla nigrovitiella Dyar, Journ. New York Ent. Soc., vol. 14,
p. 109, 1906.—Forbes, Cornell Mem. 68, p. 627, 1923.—
MecDunnough, Check list, No. 6187, 1989.
Forewing dark gray, the basal area darker than re-
mainder of wing; median and outer areas a paler
brownish gray with a faintly darker shade preceding
the subterminal line, a very sparse dusting of whitish
scales on the paler areas; antemedial line faint, narrow,
oblique and more or less curved, dull white, followed
by a narrow, blackish brown border and preceded by a
conspicuous black ridge of raised scales extending from
inner margin to top of cell; subterminal line obscure,
sinuate, pale, without dark bordering lines; discal spots
faint, blackish and confluent, forming a curved line
along discocellular vein. Hind wing pale fuscous, the
veins very slightly darkened. Both fore and hind
wings have a rather slick, glossy finish. Alar expanse,
20-21 mm.
Genitauia: As given for the genus.
Type Locatity: Pittsburgh, Pa. (type in USNM).
Foop prant: Hickory.
Disrrizution: Pennsylvania, New Brighton (June),
Pittsburgh (May); New York, Ilion (June).
UNITED STATES NATIONAL MUSEUM BULLETIN 207
The only species so far discovered referable to the
genus. The food plant record is from a female without
a locality label in the National Collection, reared from a
pupa. ‘The label reads simply ‘102, Hickory, pupated
ViI—17.” The specimen also bears an identification
label in Dyar’s handwriting.
60. Genus Oreana Hulst
Oreana Hulst, Ent. Amer., vol. 4, p. 115, 1888.
Dioryctria unicolorella Hulst.)
(Type of genus:
Characters as in Immyrla except: Forewing smooth;
veins 4 and 5 connate (rarely very shortly stalked) ; vein
10 from the stalk of 8-9 a short distance from cell.
Hind wing with 4 and 5 stalked for more than half their
lengths.
Male genitalia with numerous strong slender cornuti
on penis.
Female genitalia with bursa copulatrix strongly
sclerotized in the lobed area giving off the ductus semi-
nalis; sclerotized lateral margins of ductus bursae not
produced at genital opening and ventral margin of the
opening not appreciably concave (these differences in
the ductus bursae probably only of specific significance).
Oreana sank into the synonymy of Meroptera when
Ragonot (1889) referred its type species (wnicolorella) to
the latter genus, but it must be restored, as wnicolorella
is not a Meroptera on genitalic characters. Its chain of
numerous, strong cornuti distinguish it from any species
in Meroptera or the American genera, closely allied to
Salebria. Oreana, on most characters, seems nearest to
Immyrla, from which it is at once distinguished by its
smooth forewing. It contains but the one described
American species.
231. Oreana unicolorella (Hulst)
Fieures 319, 788
Dioryciria unicolorella Hulst, Ent. Amer., vol. 3, p. 136, 1887.
Oreana unicolorella (Hulst), Ent. Amer., vol. 4, p. 115, 1888.
Meroptera unicolorella (Hulst) Ragonot, Ent. Amer., vol. 5, p.
115, 1889; Monograph, pt. 1, p. 315, 1893.—Hulst, Phy-
citidae of N. Amer., p. 149, 1890.—Forbes, Cornell Mem.
68, p. 624, 1923.
Myelois leucophaeella Hulst, Canadian Ent., vol. 24, p. 60, 1892.
Meroptera leucophaeella (Hulst), Barnes and McDunnough, Con-
tributions, vol. 3, p. 194, 1916.
Meroptera nebulella McDunnough (not Riley), Check list, No.
6185, 1939.
Forewing mouse gray, the basal area slightly paler
than remainder of wing; transverse lines grayish white,
obscure; antemedial line oblique, notched below cell,
bordered outwardly from costa for a short distance by an
obscure blackish shade; subterminal line sinuate, with-
out appreciable dark borders; discal dots faint, brown,
usually separated but sometimes confluent; on underside
of male forewing a streak of blackish sex scaling along
basal third of costa. Hind wing pale smoky brown, the
veins slightly darkened. Alar expanse, 18-22 mm.
Male genitalia with harpe, tegumen, and vinculum
similar to those of Immyrla nigrovittella. Uncus some-
what smaller in proportion; in natural position inclined
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
downward, but when flattened out it shows an appre-
ciable constriction at middle. Anellus a simple shield-
shaped plate. Female genitalia as given for the genus;
spining of bursa not so dense or continuous as in
nigrovittella.
Type tocauities: Washington, D. C. [sic] (unicol-
orella, in AMNH, ex Rutgers); Iowa (leucophaeella, in
AMNH, ex Rutgers).
Foop piant: Apple (record from female in National
Collection labeled “bred from apple, emerged 25—V—
1904, Ottawa, J. Fletcher’).
Distrisution: Unirep Sratres: Maine, Augusta
(June); New Hampshire, Dublin; Connecticut, Hast
River (July); New Jersey, Montclair (July); Pennsyl-
vania, New Brighton (May, June, July, Aug.), Oak
Station (June), Pittsburgh (June, July) ; Iowa; Colorado;
Oregon, Milton (July); Washington, Pullman (June).
Canapa: Ontario, Ottawa (May), Trenton (July); Que-
bec, Montreal (July), St. Hilaire (July); Nova Scotia,
Cape Breton Isl. (July).
There is some mislabeling of the types or, what is more
likely, errors in the citation of type localities by Hulst.
In his original description of unicolorella he cites ““Wash-
ington, D. C.” and in his 1890 paper gives ‘‘Canada.”
His type is a male (without abdomen) from Iowa,
labeled: “HH. S. Sanders, June 13, 1886.” The type of
leucophaeella is a female with typical genitalia, labeled
“Colo., Gillette.”” The type locality citation of “Iowa”
in the original description can be written off as another
Hulst lapsus, although the species occurs there and is
represented by several examples in the National Collec-
tion. Barnes and McDunnough (1916) first put leuco-
phaeella into synonymy with unicolorella, and there does
not seem to be any reason to doubt the correctness of
that procedure. The two specimens at Rutgers labeled
“type” by Hulst certainly represent one and the same
species.
61. Olybria, new genus
Type or Genus: Myelois aliculella Hulst.
Tongue well developed. Antenna finely and densely
pubescent; on male a sinus and scale tuft in shaft at
base. Labial palpus oblique, reaching well above vertex;
second segment of male grooved to hold the maxillary
palpus; third segment short, about one-third the length
of second, acuminate, partially concealed by scaling of
second segment. Maxillary palpus of male in the form
of an aigrette; of female small, squamous. Forewing
smooth; 11 veins; vein 2 from before but near lower
outer angle of cell; 2, 3, and 4 equidistant at base; 4 and
5 shortly separated at base and thence approximate
(parallel) for a very short distance; 6 from below upper
angle of cell, straight; 8 and 9 stalked for about half
their lengths; 10 from the cell, shortly separated from
the stalk of 8-9 at base; male without costal fold. Hind
wing with vein 2 from well before lower outer angle of
cell; 3 from the angle, connate with the stalk of 4-5;
4 and 5 stalked for half their lengths; 7 and 8 closely
approximate for half their lengths beyond cell; cell
113
slightly less than half the length of wing; discocellular
vein curved, considerably extended at lower angle.
Eighth abdominal segment of male with a pair of thin,
weak, ventrolateral hair tufts.
Male genitalia with uncus small, hoodlike, projected
downward (at right angle to tegumen). Apical process
of gnathos a short, stout, curved hook. Tegumen with
a pair of straight, strongly sclerotized arms projecting
backward from its lower, posterior angles. Transtilla
absent. Harpe with costa strongly sclerotized through-
out its length and produced at apex into a sharp spine;
cucullus narrowly elongate, tapering to pointed apex.
Anellus a broadly U-shaped band, supplemented (in
JSurciferella) by some sclerotization of the membranous
tube surrounding the aedeagus. Aedeagus rather long
and stout; penis armed with a single long, strongly
sclerotized, rather slender cornutus. Vinculum stout,
longer than greatest width, narrowly truncate at
terminal margin; the latter very slightly concave.
Female genitalia with bursa smoothly sclerotized over
most of dorsal surface, its ventral surface finely sco-
binate with contorted, wrinkled and spined bands more
or less encircling the bursa at junction of bursa and
ductus bursae; ductus bursae at least as long as bursa,
rather broad, flattened (ribbonlike) and waved (twice
bent), sclerotized throughout, at genital opening the
sclerotization forming a stout, squarish ventral plate;
ductus seminalis from a lobe of bursa near junction of
bursa and ductus bursae.
This genus is distinguished from its nearest allies of
the Salebria complex by the projecting arms from
tegumen of the male genitalia, the ribbonlike, sclerotized
ductus bursae and heavy, squarish genital plate of the
female, and the simple, paired tufts of the eighth abdom-
inal segment of the male. This last character is
shared by the genus Salebriacus, which separates from
Olybria on other differences of genitalia and venation.
Two North American species, referred from Salebria
(of authors), represent the only known components of
the genus.
232. Olybria aliculella (Hulst), new combination
Figures 320, 786
Myelois aliculella Hulst, Ent. Amer., vol. 3, p. 135, 1887.
Salebria oberthuriella Ragonot, N. Amer. Phycitidae, p. 9, 1887.
Salebria aliculella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115,
1889; Monograph, pt. 1, p. 367, 1893.—Hulst, Phycitidae
of N. Amer., p. 154, 1890.—Barnes and McDunnough,
Contributions, vol. 3, p. 197, 1916—McDunnough, Check
list, No. 6217, 1939.
Forewing white dusted with black, making the general
color ashy gray, the black dusting concentrated on
extreme base and in short streaklets on the lower veins
at termen; antemedial line narrow, oblique, slightly
notched at vein 1b, white, bordered outwardly by a
black line which begins on costa as a conspicuous,
triangular, black spot; on inner margin, preceding the
antemedial line, a large orange spot; subterminal line
sinuate, narrow, white, bordered inwardly by a narrow
black line and outwardly by a broad orange band which
114 UNITED STATES NATIONAL MUSEUM BULLETIN 207
ends in a black spot at costa; along lower fold between
the transverse lines fresh specimens show a faint shading
of olivaceous ocherous; a similar shade often along
discocellular vein; completely surrounding the dis-
cocellular vein a large black ring (obicular). Hind
wing translucent white with a faint ocherous tint
especially towards outer margin and anal angle; the
veins very faintly darkened and a dark line along
termen. Alar expense, 19-22 mm.
Male genitalia showing only comparative differences
to distinguish them from those of furciferella. ‘These
are shown in the figures. Female genitalia with a row
of stiff, flattened setae along lower, posterior margin of
the eighth-segment collar.
Type LocaLity: Arizona (aliculella, in AMNH, ex
Rutgers; oberthuriella, in Paris Mus.).
Foop piuant: Ceanothus ? (Hulst, 1890).
Distrrpution: Arizona, Kingman (Oct.), White
Mts. (Aug.), Wilgus, Williams, state locality only
(June); New Mexico, Albuquerque, Fort Wingate
(June), Jemez Springs (June, July); Texas, Big Bend
region (May).
Easily identified by the orange spot preceding the
antemedial line and the large black obicular mark on
the disc of the forewing.
233. Olybria furciferella (Dyar), new combination
Ficurts 321, 787
Salebria furciferella Dyar, Journ. New York Ent. Soc., vol. 12,
p. 106, 1903.—Barnes and McDunnough, Contributions,
vol. 3, p. 197, 1916 —McDunnough, Check list, No. 6216,
1939.
Sunilar to aliculella except: Forewing more strongly
dusted with black, making ground color a decided ash
gray; the obicular spot on disk absent, replaced by a
couple of black streaks on upper and lower veins at end
of cell, forking from a black streak extending along the
upper vein of cell from the black costal dash bordering
the antemedial white line; the subbasal orange patch
on inner margin preceding the antemedial line and the
orange shade following the subterminal line reduced
and, on some specimens, obscured by black scaling.
Hind wing as in aliculella. Alar expanse, 21-23 mm.
Male genitalia similar to those of aliculella except
projecting arms of tegumen stouter; aedeagus and
cornutus longer. Female genitalia with a fringe of
fine hairlike setae along the lower posterior margin of
eighth-segment collar.
TypE tocaLity: Ashfork, Ariz. (type in USNM).
Foop prant: Unknown.
Distrisution: Arizona, Ashfork and Prescott (June,
type, o’, and 2 paratypes, 9), also 2 females with only
the state locality, from the Fernald and Brooklyn
Museum Collections and originally made cotypes of
aliculella Hulst.
The species is easily recognized by the black streak
along the upper vein of cell and is distinct from alicu-
lella though obviously very close to it.
62. Salebriacus, new genus
Typr or Genus: Nephopteryx odiosella Hulst.
Characters of Olybria except: Male antenna with
scale tuft in shallow sinus on base of shaft but greatly
reduced, a mere vestige. Forewing with vein 2 slightly
further from 3 at base than 3 is from 4; 8 and 9 stalked
for considerably more than half their lengths. Paired
tufts on eighth abdominal segment of male very weak,
mere vestiges.
Male genitalia with uncus hoodlike, elongate (longer
than wide) and not projected downward. Apical
hooked process of gnathos stout, long, reaching nearly
as far as apex of uncus. Tegumen simple. Transtilla
present but incomplete and its elements much reduced.
Harpe with apex of sclerotized costa produced as a
stout, free spine before apex of cucullus. Anellus a
simple shield. Entire genitalia more robust and
proportionally shorter than those of Olybria or the genus
Salebriaria, which follows.
Female genitalia with bursa very broad in proportion
to its length, not sclerotized, nearly half the inner sur-
face covered with a dense, spinose mat; ductus bursae
semitubular, broad and short, partially sclerotized (the
sclerotization interrupted near middle of the ductus);
ductus seminalis from near anterior end of bursa.
Highth-segment collar narrow ventrally and laterally,
dorsally produced as a spatulate apron.
This genus is another restriction from Salebria of
authors. It is distinguished chiefly by the weak tuft
of the male antenna, the squat, stout, male genitalia;
the robust, hooked projection of gnathos, the eighth-
segment collar of the female, and the place of departure
from bursa copulatrix of the ductus seminalis.
It contains but one North American species.
234. Salebriacus odiosellus (Hulst), new combination
Figures 322, 799
Nephopteryz odiosella Hulst, Ent. Amer., vol. 3, p. 132, 1887.
Salebria odiosella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115,
1889; Monograph, pt. 1, p. 366, 1893.—Hulst, Phycitidae
of N. Amer., p. 155, 1890.—Barnes and McDunnough,
Contributions, vol. 3, p. 197, 1916—MceDunnough, Check
list, No. 6219, 1939.
Salebria bakerella Dyar, Journ. New York Ent. Soc., vol. 12, p.
105, 1904.—McDunnough, Check list, No. 6220, 1939.
(New synonymy.)
Salebria yumaella Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 35, 1905—McDunnough, Check list No. 6218, 1939.
(New synonymy.)
Forewing white dusted with black, making the
ground color ashy gray, palest over the median area;
antemedial line narrow, oblique from costa to lower
margin of cell, thence slightly incurved to lower margin,
white, margined outwardly on upper half by a rather
broad black band and inwardly on lower margin by a
more or less expanded black patch; subterminal line
sinuate, narrow, white, bordered inwardly and, to a
lesser extent, outwardly by blackish lines which expand
into distinct black dashes at costa; discal dots at end
of cell, separated, usually distinct, blackish; on most
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
specimens a diffuse oblique shade from upper part of
subterminal line obliquely across wing toward inner
margin, and on a few of the more contrastingly marked
specimens a faint ocherous shading along the lower
fold; terminal dots along outer margin more or less
confluent, obscure on many specimens. Hind wing
white, translucent; faintly shaded with pale brown at
apex and along outer margin for a short distance from
apex. Alar expanse, 19-26 mm.
Male genitalia with vinculum rather abruptly nar-
rowed from middle to a truncate terminal margin.
Female genitalia with sclerotized portion of ductus
bursae at genital opening produced and broadened, its
terminal margin and lateral angles concave.
Type Locauities: “Colorado” [sic] (odiosellus, in
AMNH, ex Rutgers). Ormsby County, Nevada (baker-
ella, in USNM); Yuma County, Ariz. (ywmaella, in
USNM).
Foop prant: Unknown.
Disrrisution: Teras, Blanco County [?]; Arizona,
Mohave County (July, Sept.), Yavapai County, Yuma
County; Colorado; Utah, Bellevue (May, June, July);
Nevada, Ormsby County (July), Pyramid Lake, Reno;
California, Jacumba (May), Mexican Wells (Clarke
Mts., Sept.), Morongo Valley (May).
The types of odiosellus, bakerella, and yumaella are all
males with identical genitalia. That of yumaella is a
small, somewhat suffused specimen with the black
markings less strongly contrasted than those of typical
Nevada examples. In our series there is a complete
intergradation between the extreme forms.
Barnes and McDunnough (1916) called attention to
the confused citations of the type locality of odiosellus.
Hulst gives Colorado in his original description, but in
his 1893 paper cites ‘“‘Texas.”” The type at Rutgers is
labeled ‘‘Blanco Co., Texas.’’ I suspect that this is a
mislabeling, for Colorado seems a more likely locality
than central Texas. In my paper on the cactus-feeding
Phycitinae (Proc. U.S. Nat. Mus., vol. 86, p. 389, 1939)
I misapplied the Hulst name to a species described by
Dyar as Ozamia clarefacta. This error is discussed
further under the treatment of clarefacta (p. 258).
63. Salebriaria, new genus
Typsz or Genus: Salebria ademptandella Dyar.
Tongue well developed. Maleantenna pubescent with
strong scale tuft in sinus at base of shaft. Labial palpus
obliquely upturned, reaching above vertex, smoothly
scaled; second segment on male grooved to hold the
maxillary palpus, appressed to face; third segment mi-
nute and hidden in scaling of second on male, somewhat
longer and partially exposed on female, acuminate.
Maxillary palpus of male in the form of an aigrette; of
female subsquamous. Forewing smooth; 11 veins; vein
2 from before but near lower outer angle of cell; 3
usually somewhat nearer to 4 than to 2, sometimes equi-
distant from them; 4 and 5 shortly separated at base,
rarely (in smaller specimens) closely approximate; 6
from below upper angle of cell, straight; 8 and 9 stalked
115
for well over half their lengths (except in fructetella) ;
male without costal fold; on underside of male wing a
streak of appressed, black sex-scaling along basal third
of costa. Hind wing with vein 2 from before, but
rather near lower outer angle of cell; 3 from the angle,
connate with the stalk of 4—5; 4 and 5 stalked for approx-
imately half their lengths (for slightly over half in
Jructetella); 7 and 8 approximate beyond cell (except in
pumilella and fructetella where they are contiguous or
weakly anastomosed for a short distance); cell slightly
less than half the length of wing. Eighth abdominal
segment of male with compound ventral scale tufts.
Male genitalia with uncus subtriangulate (apex rather
broadly rounded). Apical process of gnathos a short
hook, slender (except on fructetella). Transtilla absent.
Harpe with apex of sclerotized costa produced as a short
free spine before apex of cucullus. Anellus a semitubu-
lar shield. Penis armed with a single, strongly sclero-
tized, moderately long cornutus. Vinculum stout, sub-
triangulate, slightly longer than greatest width.
Female genitalia with much of inner surface of bursa
covered with a dense matting of fine spines (especially
towards distal end); bursa also sometimes partially
sclerotized (pumilella, fructetella) ; ductus bursae shorter
than bursa, strongly sclerotized (at least towards genital
opening), partially flattened (ribbonlike in pumuilella);
ductus seminalis from lobe of bursa adjacent to the
junction of bursa and ductus bursae.
This genus is a further restriction from Salebria. It
is a compact group of very closely related species; in its
smooth forewings and male genitalia markedly distinct
from typical Salebria; closest to Salebriacus from which
it is at once distinguished by the compound scale tufts
on eighth abdominal segment of the male and the place
of departure of the ductus seminalis from bursa in the
female. How many valid species are represented by the
several names in our North American lists and definitely
referable to the genus cannot be exactly determined
until larger series of reared specimens are available.
The genitalia (except for those of pumilella and fruc-
tetella) offer little or nothing in the nature of trust-
worthy specific characters. The maculation and color
differences on forewings that have been used by previous
authors seem to be equally unreliable.
235. Salebriaria turpidella (Ragonot), new combination
Fiaures 323, 802
Salebria turpidella Ragonot, Nouv. Gen., p. 19, 1888; Monograph,
pt. 1, p. 346, 1893.—Forbes, Cornell Mem. 68, p. 625,
1923.—McDunnough, Check list, No. 6196, 1939.
Salebria ademptandella Dyar, Proc. Ent. Soc. Washington, vol.
10, p. 114, 1908.—McDunnough, Check list, No. 6198,
1939. (New synonymy.)
Forewing powdery gray, the ground color variable,
ranging from pale ash gray (ademptandella) to a more
suffused pale brownish gray; basal area usually some-
what paler than median area; the transverse lines but
little paler than the ground color, indicated chiefly by
their dark margins; the latter narrow, blackish, well
contrasted, especially on the paler examples; antemedial
116 UNITED STATES NATIONAL MUSEUM BULLETIN 207
line sinuate-angulate, bordered outwardly by a black
line and inwardly, on lower half, by a similar narrow
black line; on many specimens a small, faint, whitish
patch just beyond the antemedial line on inner margin;
subterminal line sinuate, bordered inwardly by a black-
ish line, the latter fainter than that bordering the ante-
medial line; discal spots black, confluent, normally
forming a black line along the discocellular vein, but on
individual specimens tending to separation on one or
the other forewing. Hind wing smoky white to brown,
variable in both sexes. Alar expanse, 17-18 mm.
Male genitalia show no distinguishable differences
from those of typical turpidella, nubiferella, or annulo-
sella. Female genitalia with the spining on anterior half
of bursa of a nearly uniform fineness (no dense concen-
tration of darker spines at the closed end); bursa with-
out appreciable sclerotization at middle.
TYPE LOCALITIES: Unitep Srares: (turpidella, 3,
in Paris Mus.); Kerrville, Tex. (ademptandella, in
USNM).
Foop prant: Unknown.
Disrrisution: Texas, Burnet County (Oct.), Kerr-
ville (Apr., May, June), Sabinal (Sept.), also one Texas
male from the Boll Collection (No. 558) labeled
‘“Kurope”’; Florida, Gotha (F. Rauterberg, collector,
2 specimens, of and Q).
The foregoing from typical examples. Also before
me are transitional examples between turpidella and
nubiferella from Texas, Blanco County (May, July)
and Shovel Mountain, and North Carolina, Southern
Pines (Apr., July, Aug.) and Tryon (May, Sept.).
These specimens are dark, with the size and markings
of turpidella but with female genitalia more like those of
anmulosella. Some of them had been identified as
annulosella. Others were under turpidella. I doubt
very much if turpidella is anything more than a variety
of nubiferella. Dyar’s ademptandella is nothing but a
paler form of the typical turpidella.
236. Salebriaria nubiferella (Ragonot), new combination
Salebria nubiferella Ragonot, N. Amer. Phycitidae, p. 8, 1887;
Monograph, pt. 1, p. 344, 1893.—Hulst, Phycitidae of N.
ane p. 150, 1890—McDunnough, Check list, No. 6191,
Salebria annulosella nubiferella (Ragonot) Forbes, Cornell Mem.
68, p. 625, 1923.
If this and the preceding “species” (turpidella) are
anywise distinct, the name nubiferella will apply to a
larger form (21 mm.) with the basal area of forewing
dark and concolorous with the darker shading in median
and outer areas and with a more strongly contrasted
white patch on the inner margin just beyond the ante-
medial line; but neither of these differences, nor those
used by Ragonot (Monograph, pp. 329, 345) to separate
turpidella and nubiferella are constant. According to
Clarke’s notes the species is represented in the Paris
Museum only by the unique type, a male labeled “‘type
orig., pl. XIV fig. 23, Amer. Sept. don. C. V. Riley.” I
have examined its genitalia and they show nothing to
distinguish them from those of turpidella, annulosella,
or engelt.
What appears to be a typical female in the National
Museum from Putnam County, Ill. (July) has the well-
contrasted white spot on inner margin, dark hind wings
and dark gray forewings, the latter with discal dots
fused and the dark (blackish) borders of antemedial and
subterminal lines as in turpidella, but somewhat weaker.
Its alar expanse is 20mm. The female genitalia have a
rather dense and dark concentration of fine spines at
the anterior end of bursa and a thickening and slight
sclerotization of the membrane at middle of bursa, the
sclerotized part showing what seems to be a fixed longi-
tudinal fold. In these particulars the genitalia are like
those of annulosella.
Type Locauity: “Amer. Sept.’’ [Texas] (type in Paris
Mus.).
Foop pranr: Unknown.
Besides the aforementioned female there are before
me a male (19 mm.) from Putnam County, Ill. July),
and a male (18 mm.) and female (19 mm.) from near
St. Louis, Mo. (June) which appear to be conspecific
with the Illinois female. They have the white spot on
inner margin less strongly marked, but the strength of
this marking does not seem to be of any significance.
Presumably several of the Texas examples from the
intermediate specimens mentioned under turpidella
should be referred here if a specific distinction can be
maintained between turpidela and nubiferella. We
shall have to have host plant association and good
reared series before such a distinction can be made with
any certainty.
237. Salebriaria engeli (Dyar), new combination
Salebria engeli Dyar, Journ. New York Ent. Soc., vol 14, p. 107,
1906.—Forbes, Cornell Mem. 68, p. 625, 1923.—McDun-
nough, Check list, No. 6195, 1939.
Forewing dark brownish gray, the transverse lines
obscured; antemedial line indicated only by its very
faint, narrow, angulate, outer black border, followed
outwardly on inner margin by a strongly contrasted
white patch; subterminal line faint, but slightly paler
than the ground color and with an obscure, narrow, dark
inner border, sinuate; discal dots more or less confluent,
blackish with some pale scaling on their outer margins;
a row of separated blackish dots along outer margin.
Hind wings smoky fuscous, on darker specimens with
a pale brownish tint; the veins darkened. Alar expanse,
18-20 mm.
Genitalia as in annulosella.
TypE Locauity: Oak Station, Pa. (type in USNM).
Foop prant: Unknown.
DistrisutTion: Pennsylvania, Hazleton, New
Brighton (July), Oak Station (July); Maryland, Plum-
mers Isl. (July); Illinois, Decatur (May); Tezas,
Kerrville (May, June, July).
An intermediate form between typical nubiferella and
annulosella, probably only a variety of the former.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 117
238. Salebriaria annulosella (Ragonot), new combination
Fiaure 800
Salebria annulosella Ragonot, N. Amer. Phycitidae, p. 7, 1887;
Monograph, pt. 1. p. 346, 1893.—Forbes, Cornell Mem. 68,
p. 625, 1923.—MeDunnough, Check list, No. 6197, 1939.
Salebria robustella Dyar, Proc. Ent. Soc. Washington, vol. 10,
p. 114, 1908.—McDunnough, Check list, No. 6211, 1939
(New synonymy.)
Forewing gray; antemedial line but slightly paler than
the ground color and not sharply defined, margined on
outer side at costa by a more or less triangulate blackish
patch and on inner side at inner margin by a somewhat
larger blackish patch which extends to or nearly to base
of wing; discal dots distinctly separated, black, sur-
rounded by pale dusting; subterminal line obscure; a
row of distinct blackish dots along terminal margin.
Alar expanse, 18-20 mm.
Female genitalia figured from specimen from Burnet
County, Tex. They are like those of the type in Paris
and differ in no essential details from those of females
of nubilella.
Type LocauitiEs: Texas (annulosella, in Paris Mus.) ;
Burnet County (robustella, in USNM).
Foop prant: Unknown.
Distrisution: Texas, Blanco County (Apr.); Burnet
County (Apr.); North Carolina, Raleigh (June).
According to Clarke’s notes, two examples, forming
the basis of Ragonot’s diagnosis of the species in his
Monograph, are in the Paris Museum. One is a female
from Texas, obviously the type but not so labeled. It
was the specimen figured in the Monograph (pl. 9, fig.
6). The other specimen is a male labeled: ‘Dallas,
Texas, Boll, 24-V-689.’’ A female in the U. S. Na-
tional Museum, also a Boll specimen from Texas (No.
559) and probably a mate to the Paris male, bears a
name label (Salebria annulosella) in Ragonot’s hand-
writing. I doubt that either of these Boll specimens is
a typical annulosella. Our female in markings is inter-
mediate between annulosella and tenebrosella with female
genitalia like the latter species. Dyar’s type of robus-
tella (a male) matches Ragonot’s figure and description
of annilosella in all details except that the discal spots
are obscured on one forewing. A female, obviously the
other sex of Dyar’s type, had been identified by him as
annulosella. ‘The remaining Texas examples before me
(Blanco County) had been identified by Hulst as either
nubiferella or pumilella.
239. Salebriaria tenebrosella (Hulst), new combination
Fieurss 801, 803
Nephopteryz tenebrosella Hulst, Ent. Amer., vol. 3, p. 131, 1887.
Nephopteryx quercicolella Ragonot, N. Amer. Phycitidae, p. 7,
1887.
Salebria tenebrosella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115,
1889; Monograph, pt. 1, p. 347, 1893.—Hulst, Phycitidae
of N. Amer., p. 151, 1890.—McDunnough, Check list, No.
6199, 1939.
Salebria heinrichalis Dyar, Ins. Insc. Menstr., vol. 5, p. 45,
1917.—McDunnough, Check list, No. 6192, 1989. (New
synonymy.)
Forewing dark gray more or less shaded with black,
especially in basal area; antemedial line whitish, dif-
fused, oblique, sometimes interrupted at middle, pre-
ceded by a dark red patch on lower half; cutting this
red patch an oblique black line which fuses into the
black basal shade on costal half of basal area; subter-
minal line obscure, sinuate, very slightly paler than
the ground color; discal dots coalesced into a black
lunulate line along discocellular vein, partially obscured
in the dark ground color over middle of wing but set
off by some pale (whitish) dusting along its outer edge;
separated blackish dots along terminal margin. Hind
wings brownish gray; the veins slightly darkened.
Alar expanse, 17-18 mm.
Male genitalia showing no distinguishing characters
from those of preceding species.
Type Locatities: Texas (tenebrosella, in AMNH, ex
Rutgers); ‘“‘America Septentrionalis” (querciocolella, in
Paris Mus.) ; Falls Church, Va. (heinrichalis, in USNM).
Foop piant: Oak (larva a leaf tier).
Distripution: Texas; Missouri (Aug.); Virginia,
Falls Church (Apr.).
In addition to typical examples from the above lo-
calities there is before me a series of males and females
of a slightly larger average size, 18-20 mm., with a
somewhat more diffused and contrasted whitish ante-
medial line and little or no trace of the subbasal red
patch on forewing and no red on the thorax. On
typical fenebrosella the tips of the patagia are shaded
with red. The females of these variant examples also
have larger genitalia (fig. 803). They may represent a
food plant race or a color form of tenebrosella but hardly
anything more. In view of the already obscure specific
limits of the described species I do not feel justified in
adding a further name.
The variety is represented in the National Collection
from the following localities: Massachusetts, Cohasset
(July), Martha’s Vineyard (July); New York, Utica
(Aug.); Maryland, Plummers Isl. (Aug.); District of
Columbia, Washington (June); North Carolina, Tryon
(May); Georgia, Atlanta (June); Jilinois, Decatur
(May), Lacon (June); Missouri, near St. Louis, and one
small example from the Murtfeldt Collection labeled
“130 M. apple, 5-8-89.”’ Several of these had been
previously misidentified as Oreana leucophaeella (Hulst).
240. Salebriaria pumilella (Ragonot), new combination
Fiaures 325, 804
Salebria pumilella Ragonot, N. Amer. Phycitidae, p. 8, 1887;
Monograph, pt. 1, p. 344, 1893—MceDunnough, Check list,
No. 6190, 1939.
Salebria georgiella Hulst, Canadian Ent., vol. 27, p. 57, 1895.—
McDunnough, Check list, No. 6210, 1939. (New synony-
my.)
Forewing dark gray shaded with brown; a rather pale
chocolate brown, triangulate patch on inner margin on
inner side of antemedial line; the latter narrow, white,
its blackish bordering lines broken, and obscure except
the inner one on lower half of wing; a similar black
border on the inner edge of the chocolate brown patch;
on some specimens (especially faded examples) a slight
ocherous shading at extreme base of wing; on most fresh
118 UNITED STATES NATIONAL MUSEUM BULLETIN 207
specimens a narrowly elongate, grayish fuscous patch
on midcosta; some white dusting on inner margin
beyond antemedial line and in cell towards its outer
margin; subterminal line fine, white, its dark borders
obscure; discal spots confluent, forming a narrow,
blackish line along discocellular vein; the blackish dots
along terminal margin weak, more or less confluent.
Hind wing pale smoky fuscous. Alar expanse, 15-17
mm.
Male genitalia with spine from apex of sclerotized
costa of harpe projecting straight out beyond apex of
cucullus (not curved dorsally as in the other species).
Female genitalia with ductus bursae flattened, ribbon-
like, sclerotized throughout and dorsoventrally folded
towards bursa copulatrix, its lower margin produced at
genital opening into a subtriangulate projecting shield
with somewhat rounded terminal margin; bursa with
nearly half of one side strongly and smoothly sclero-
tized.
TypE LOCALITIES: Texas (pumilella, in Paris Mus.);
Charlotte Harbor, Fla. (georgiella, mn AMNH, ex Rut-
gers).
Foop pLrant: Unknown.
Distrisution: Florida, Charlotte Harbor (Mar.);
Texas, Burnet County (Apr.), also 3 examples (co and
99) with only the state locality and without dates; North
Carolina, Southern Pines (June, Aug.).
Hulst’s references to pumilella in his Phycitidae of
N. Amer. (p. 150) are omitted from the above synonymy,
since he had misidentified Ragonot’s species. The
technical description he gives was simply copied from
Ragonot’s original description. There is no doubt
about the synonymy of georgiella.
Tn all examples I have seen, veins 4 and 5 of forewing
are rather closely approximate.
241. Salebriaria fructetella (Hulst), new combination
Ficures 324, 805
Myelois fructetella Hulst, Canadian Ent., vol. 24, p. 59, 1892.
Salebria rectistrigella Dyar, Proc. Ent. Soc. Washington, vol. 10,
p. 115, 1908.
Salebria fructetella (Hulst) Barnes and McDunnough, Contribu-
tions, vol. 3, p. 194, 1916—McDunnough, Check list, No.
6203, 1939.
Forewing glossy brownish gray shaded with blackish
brown and dusted with white; the black shading form-
ing a blotch on costa following, and a similar blotch on
inner margin preceding the antemedial line, and an
obscure dark shade extending from costa near apex
obliquely inward across the subterminal line; white
dusting limited to a more or less triangulate cloud from
midcosta surrounding the upper discal spot and a
smaller cloud on costa preceding the antemedial line;
antemedial line narrow, slightly oblique, straight or (on
some examples) with a slight notch at middle, white and
distinctly contrasted against the ground color but with-
out appreciable black bordering lines; discal dots sep-
arated, blackish; subterminal line obscure, whitish gray,
vertical except for a median outward bulge; terminal
dots obscure, brown. Hind wing smoky white shading
to fuscous along termen; veins very slightly darkened.
Alar expanse, 15-18 mm.
Male genitalia distinguished from those of other
species in the genus by the spoon-shaped apical projec-
tion of gnathos, the V-shape of the sclerotized part of
anellus, and the longer cornutus.
Female genitalia with ductus bursae very short, its
median area membranous, a moderately broad sclero-
tized and granulate band at genital opening, strongly
sclerotized and longitudinally ridged towards junction
with bursa; this peculiar sclerotization extending into
and occupying nearly half of the bursa, anterior (closed)
end of bursa fused into a thickened (but not sclerotized)
membrane; remainder of bursa covered by a mat of
fine spines.
TyPE LocALitius: Blanco County, Tex. (fructetella,
in AMNH, ex Rutgers); Kerrville, Tex. (rectistrigella,
in USNM).
Foop Piant: Oak (larva a leaf feeder).
Distrisution: Florida, Key West, Orlando (Mar.);
Louisiana, Natchitoches Parish (Aug.); Texas, Blanco
County (June), Kerrville (May, June), Sabinal (Sept.),
Shovel Mountain (@June, July); Arizona, Williams
(Sept.), Missourt (June, reared); District of Columbia
(May, Aug., reared); New York, Bellport (June, Sept.,
reared).
Superficially fructetella strikingly resembles Acrobasis
amplexella Ragonot (especially the females). The reared
examples before me had been identified to the latter
name. The male antennal character and the genitalia
of both sexes, however, easily separate the two species.
64. Quasisalebria, new genus
TYPE OF GENUS: Quasisalebria admixta, new species.
Characters of Salebriaria except: Labial palpus erect,
appressed to flattened face. Forewing with vein 8 and
9 stalked for slightly more than two-thirds of their
lengths. Hind wing with veins 4 and 5 stalked for over
three-fourths of their lengths; 7 and 8 shortly anas-
tomosed beyond cell.
Male genitalia with costa of harpe sclerotized for its
entire length, not appreciably produced at apex, but
with a strongly sclerotized, free, articulating arm from
base; shieldlike part of anellus with a pair of long,
divergent, lateral horns; penis without cornutus.
Female genitalis with ductus bursae thickened
(cartilaginous in texture) except at junction with bursa
copulatrix; bursa finely spined only in area adjacent to
junction with ductus bursae.
The type of this genus is, in many respects, close to
Salebriaria fructetella and appears to be an aberrant
offshoot of Salebriaria. I propose the new generic
designation reluctantly; but no other procedure is pos-
sible if we are to have any exact definition of genera for
the species allied to Salebria. Nothing in our American
fauna is properly referable to the latter genus, which is
characterized by a forewing with partial scale ridge in
the subbasal area; harpe (fig. 332) with erect clasper
from near middle and costal margin weakly sclerotized ;
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 119
bursa copulatrix of female without spining or granula-
tions (membranous and smooth in the type, palumbella).
242. Quasisalebria admixta, new species
Figures 328, 806
Forewing ashy white shaded with olivaceous brown
or grayish fuscous in outer area and on lower half of
basal area; the whitish ground color strongly contrasted
on costal half of basal area and in a more or less tri-
angulate area extending from median half of costa into
cell and including the discal spots; antemedial line
distinct, narrow, slightly curved, white, bordered out-
wardly on costa by a strongly contrasted, black, tri-
angulate patch and preceded on inner margin by a
similar quadrate spot; subterminal line narrow, white,
close to termen, and outwardly bulged at middle,
bordered by fine blackish lines which begin as strong
black smudges at costa; discal dots separated, the lower
one always distinct, the upper sometimes absent. Hind
wing semihyaline, white with a brownish shade at apex
and a narrow brown line along upper half of termen;
the veins not appreciably darkened. Alar expanse,
19-21 mm.
Male genitalia. Characters as given for the genus.
The peculiar development of the anellus may be only
of specific significance.
TypE Locautity: Provo, Utah (type in USNM,
61343).
Foop prant: Unknown.
Described from male type and two male and four
female paratypes from the type locality (July, Aug.);
one male paratype from Redington, Ariz.; two female
paratypes from Bellevue, Washington County, Utah
(May); and one female paratype from Glenwood
Springs, Colo. (Aug.).
The species is superficially similar to Salebriaria
fructetella Hulst, but with the white areas and blackish
markings of forewing more strongly contrasted.
Genus 65: Ortholepis
[Venational division B. Veins 4 and 5 of forewing connate
(rarely in individual specimens, slightly separated at base).
Hind wing with vein 2 from rather near lower outer angle of
cell. Male genitalia with transtilla complete but its median
area weakly sclerotized and granulate; costa of harpe strongly
sclerotized throughout but not produced at apex; penis armed
with a single, long, strong cornutus. Female genitalia without
signum or scobinations in bursa.]
65. Genus Ortholepis Ragonot
Ortholepis Ragonot, N. Amer. Phycitidae, p. 6, 1887; Monograph,
pt. 1, p. 214, 1893.—Hulst, Phycitidae of N. Amer., p. 140,
1890. (Type of genus: Ortholepis jugosella Ragonot.)
Tongue well developed. Antenna of male pubescent,
shaft with sinus towards base containing a row of black
toothlike spines more or less concealed under a weak
seale tuft; antenna of female simple. Labial palpus
obligue; second segment roughly and rather broadly
scaled, on male slightly grooved on inner side; third
segment very short, acuminate, reaching to height of
vertex. Maxillary palpus of male squamous (jugosella)
or in the form of an aigrette (pasadamia); of female
minute and filiform. Forewing with ridge of raised
scales on inner side of antemedial line, not reaching
costa or inner margin; 11 veins; 2 from before lower
outer angle of cell; 3 from the angle, separated at base
from 4-5; 4 and 5 connate, rarely (in individual speci-
mens) slightly separated at base; 6 from below upper
angle of cell, straight, 8 and 9 stalked for slightly more
than half their lengths; 10 from the cell, shortly sepa-
rated from 8-9 at base and thence divergent; male
without costal fold. Hind wing with vein 2 from before
but rather near lower outer angle of cell; 3 from the
angle, connate with the stalk of 4-5, short as compared
with 2; 4 and 5 stalked for half or a trifle over half their
lengths; 7 and 8 contiguous or anastomosed for a very
short distance beyond cell; cell less than half the length
of wing; discocellular vein curved, considerably extended
at lower angle. Eighth abdominal segment of male
simple.
Male genitalia with uncus subtriangulate; slightly
produced (lobed) near its lower, lateral angles. Apical
process of gnathos a short, simple hook. Transtilla
complete but its median area weakly sclerotized and
granulate; its lateral elements broadly sclerotized.
Harpe narrow; costal margin sclerotized throughout but
not produced. Anellus U-shaped, narrowly sclerotized.
Penis armed with a single stout cornutus nearly as long
as aedeagus. Vinculum stout, triangulate; about as
long as greatest width.
Female genitalia without signum; bursa elongate,
large, longitudinally wrinkled, partially sclerotized in
the lobed area bearing the ductus seminalis; ductus
bursae considerably shorter than bursa; flattened,
strongly sclerotized, at least near and at its junction
with bursa copulatrix; ductus seminalis from lobe of
bursa near junction of bursa and ductus bursae.
Eighth-segment collar with sclerotized part on dorsum
reduced to a U-shaped band.
An American genus showing affinities to Polopeustis
and the various genera of the Salebria complex; but
easily distinguished by its genitalia. Ragonot’s de-
scription is somewhat misleading. Veins 4 and 5 of
forewing are normally connate and not “nearly paral-
lel”? except well beyond base; from base to near middle
they are divergent.
243. Ortholepis jugosella Ragonot
Figures 23, 329, 808
Ortholepis jugosella Ragonot, N. Amer. Phycitidae, p. 6, 1887;
Monograph, pt. 1, p. 214, 1893.—Hulst, Phycitidae of N.
Amer., p. 140, 1890.—McDunnough, Check list, No. 6149,
1939.
Maxillary palpus of male squamous.
Forewing gray, rather shiny; palest (on some speci-
mens ash gray) on costal half of median area and on the
anterior costal half of basal area; remainder of basal
and median areas and the area beyond the subterminal
line brownish gray with a faint purplish suffusion;
120 UNITED STATES NATIONAL MUSEUM BULLETIN 207
antemedial line oblique from costa to lower margin of
cell, thence vertical to inner margin, its upper half
obscure, indicated chiefly by a blackish outer border,
lower half distinct, whitish; the antemedial line pre-
ceded by a raised tuft of brown and blackish scales
which are bordered inwardly by a more or less con-
trasted white line; subterminal line faint, narrow,
slightly bulged at middle; discal dots black, separated,
distinct and rather large, especially the upper one;
terminal dots reduced, obscure and more or less conflu-
ent. Hind wing light brown; a thin blackish line along
termen; the veins not appreciably darkened. Alar
expanse, 19-20 mm.
Male genitalia as given for the genus. Female geni-
talia with posterior half of ductus bursa very weakly
sclerotized.
Type Locaity: “‘America septentrionalis” (type in
Paris Mus.).
Foop pLiant: Hickory (Carya alba) and wild azalea
(Ragonot records). Hickory and walnut are the more
probable hosts.
Distripution: Unirep StratEs: Connecticut, Hast
River (July). Canapa: Nova Scotia, White Point
Beach (Queens County, July).
244. Ortholepis pasadamia (Dyar), new combination
Fiaure 807
Immyrla pasadamia Dyar, Ins. Insc. Menstr., vol. 5, p. 45,
1917.—McDunnough, Check list, No. 6189, 1939.
Maxillary palpus of male in the form of a short aig-
rette.
Forewing uniformly dark glossy gray with a purplish
tint; the transverse lines well marked, narrow, whitish; a
contrasted white inner margin to the subbasal tuft; dis-
cal dots usually confluent, distinguishable but not
strongly contrasted against the dark ground color.
Hind wing smoky fuscous; the veins faintly darkened.
Alar expanse, 17-20 mm.
Male genitalia as in jugosella. Female genitalia as in
jugosella except ductus bursae sclerotized along ventral
surface to genital opening.
TypPE LOcALITy: St. Johns, Quebec (type in USNM).
Foop Puant: Betula.
Disrrizution: Unirep States: Pennsylvania, Ha-
zleton (June); New Hampshire, Dublm, Hampton
(July) ; Maine, Mount Desert Island (July), Sebec Lake
(July); Washington, Meadow Creek (Grant County,
Apr.), reared specimen, Walla Walla (June). Canapba:
Ontario, Blacotasing (July), Ottawa (July), Waubamia
(Perry Sound, July); Quebec, St. Johns (June).
The foregoing description was drawn from typical
examples represented in the National Collection by a
series of 16 males and females. There are also before
me four specimens from Maine, New Hampshire, and
Quebec of what appears to be a color form in which the
transverse lines (except for the white inner border of the
raised-scale patch) are almost completely obliterated;
the ground color of the forewing is darker (more pur-
plish) and the hind wing is brown (as in jugosella) ; there
is also a faint narrow dusting of whitish scales in the
median area of forewing. The genitalia of these speci-
mens are identical with those of typical pasadamia.
Examples of both forms have been reared from Betula.
Genus 66: Polopeustis
[Venational division B. Vestiture of head, thorax, labial palpi,
and femora a mixture of scales and hairs.]
66. Genus Polopeustis Ragonot
Polopeustis Ragonot, Monograph, pt. 1, p. 233, 1893.—Staudin-
ger and Rebel, Catalog der Lepidopteren des palaearctichen
Faunengebietes, vol. 2, p. 30, 1901.—Spuler, Die Schmetter-
linge Europas, vol. 2, p. 212, 1910.—Forbes, Cornell Mem.
68, p. 622, 1923.—Janse, Journ. Ent. Soc. South Africa, vol.
5, p. 30, 1942. (Type of genus: Phycis annulatella Zetter-
stedt; figs. 24, 330, 809.)
Tongue well developed. Antenna of male ciliate-
pubescent (the cilia a trifle shorter than width of seg-
ments) the shaft curved towards base and containing
two or three short teeth in the incurvation; of female
simple. Labial palpus obliquely ascending, not reach-
ing height of vertex; third segment short, less than one-
third of second. Maxillary palpus minute, filiform.
Forewing smooth; 11 veins; vein 2 from before but near
lower outer angle of cell; 3 from the angle; 4 and 5
separated at base; 6 from below upper angle of cell,
straight; 8 and 9 stalked for over half their lengths; 10
from the cell, shortly separated from the stalk of 8-9 at
base; male without costal fold. Hind wing with vein 2
from well before outer angle of cell; 3 from the angle; 4
and 5 anastomosed for about half their lengths beyond
cell; 7 and 8 closely approximate for a short distance
beyond cell; cell less than half the length of wing; dis-
cocellular vein curved, produced atlowerangle. Highth
abdominal segment of male with pair of ventrolateral
hair tufts.
Vestiture of head, thorax, labial palpi, femora, and
and foretibiae a mixture of scales and hairs.
Male genitalia with uncus about as broad as long; at
apical margin broadly rounded. Apical process of
enathos (from ventral view) an inverted heart-shaped
lobe terminating in a short, slender, hooked spine.
Transtilla absent. Harpe short, stubby; its apex bluntly
rounded; costa broadly and strongly sclerotized through-
out, but not produced at apex; otherwise simple. Anel-
lus a U-shaped plate terminating in short lateral lobes.
Aedeagus slender; penis armed with moderately stout,
elongate cornutus or two similar cornuti. Vinculum
stout, slightly longer than greatest width; tapermg to
blunt and rather broad terminal margin.
Female genitalia without signum; bursa copulatrix
small and with a rather broad, fused cartilagenouslike
thickening around its lateral and anterior margins,
otherwise minutely granulate and containing a weak
elongate chitinized strip; ductus bursae twice as long as
bursa, flattened, its ventral surface sclerotized and gran-
ulate through its length, the sclerotization extending
into bursa; genital opening simple. Highth-segment
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 121
collar with sclerotized area reduced to a narrow, more
or less U-shaped dorsolateral band (similar to that of
Ortholepis).
An Old World genus of Holarctic distribution with
one European and one North American species; easily
identified by its genitalia and hairy vestiture.
245. Polopeustis arctiella (Gibson)
Fiaures 331, 810
Pyla arctiella Gibson, Report of the Canadian Arctic Expedition
(1913-18), vol. 3, pt. 1, p. 46, 1920.
Polopeustis annulatella arctiella (Gibson) McDunnough, Ca-
nadian Ent., vol 67, p. 174, 1935; Check list, No. 6150, 1939.
Forewing slate gray with a fine scattered sprinkling
of white, the whitish dusting more concentrated in basa]
area and the area beyond the subterminal line; trans-
verse lines rather broad, well contrasted, white; ante-
medial line oblique and slightly angulate; subterminal
line sinuate; a narrow blackish shade bordering the
antemedial outwardly and the subterminal inwardly;
discal dots obscure, more or less confluent. Hind wings
smoky white; the veins darkened; a narrow dark shade
along termen. Alar expanse, 21-25 mm.
Male genitalia with two cornuti on penis; otherwise as
in annulatella. Female genitalia differ from those of
annulatella chiefly in the shape of the sclerotized area of
eighth-segment collar (compare figs. 809a and 810a).
The differences in shape and extent of sclerotized area
of bursa are probably individual in character.
Type Locauity: Collinson Point, Alaska (type in
Canadian Nat. Coll.).
Foop piant: Unknown.
Distrisution: AuasKa: Collinson Point (July).
Canava: Labrador, Hopedale (July), Nain; Manitoba,
Fort Churchill (June, July).
Gibson’s name may represent no more than a New
World race of annulatella; but arctiella is at least that.
Good series of both are before me and their genitalic
differences appear to be constant; the male of annulat-
ella has but one cornutus, that of arctiella has consist-
ently two cornuti. If and when intergrading examples
of Polopeustis are recovered from northern Siberia the
name arctiella may be reduced to subspecific status or
referred as a synonym to annulatella. Meanwhile a
specific separation seems the safer procedure.
Genera 67-70: Meroptera to Tulsa
[Venational division B. Veins 4 and 5 of forewing very shortly
stalked, connate or closely approximate at base, in Tulsa approxi-
mate at base and for a short distance beyond; § and 9 long stalked;
10 frequently connate or shortly stalked with 8-9, if from the cell
approximate to the stalk of 8-9 for some distance from its base.
Antenna of male with sinus and scale tuft in base of shaft. Labial
palpus erect or obliquely upturned. Maxillary palpus of male
in the form of an aigrette or squamous. Male genitalia with
transtilla frequently complete, but if so, weakly sclerotized;
harpe with clasper always present and well sclerotized, digitate
or enlarged and spined; harpe with long hair brush from inner
surface along lower edge of basal half of sclerotized costa; penis
armed with two stout, rather short cornuti. Female genitalia
300329—56——_9
with bursa finely and densely spined, usually with one or more
sclerotized, granulate patches.]
67. Genus Meroptera Grote
Meroptera Grote, Canadian Ent., vol. 14, p. 29, 1882.—Hulst,
Phycitidae of N. Amer., p. 148, 1890.—Ragonot, Mono-
graph, pt. 1, p. 312, 1893.—Forbes, Cornell Mem. 68,
p. 624, 1923. (Type of genus: Pempelia pravella Grote.)
Emmerita Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 76,
1930. (Type of genus: Meroptera mirandella Ragonot. New
synonymy.)
Tongue well developed. Antenna weakly pubescent;
on male with sinus and scale tuft in base of shaft. La-
bial palpus upturned, closely appressed to face; smooth
scaled; reaching above vertex; second segment long, on
male hollowed to receive maxillary palpus; third seg-
ment short (about one-fourth of second), acuminate.
Maxillary palpus of male in the form an of aigrette;
of female squamous. Forewing smooth; 11 veins; vein
2 from before but rather near lower outer angle of cell;
3 from the angle, at base slightly nearer to 4-5 than to
2; 4 and 5 very shortly stalked, connate or closely ap-
proximate at base; 6 from below upper angle of cell,
straight or (rarely) very slightly bent towards base; 8
and 9 stalked for over two-thirds their lengths; 10
shortly stalked or connate with the stalk of 8-9 (indi-
vidually variable); male without costal fold. Hind
wing with vein 2 from before but near lower outer angle
of cell; 3 from the angle, connate with 4; 4 and 5 stalked
for slightly more than half their lengths; 7 and 8 closely
approximate for a short distance beyond cell; cell less
than one-half the length of wing; discocellular vein
curved, outwardly produced at lower angle of cell.
Highth abdominal segment of male with compound
scale tufts.
Male genitalia with uncus triangulate or subtriangu-
late, its apex bluntly rounded. Apical process of
gnathos a short, stout hook. Transtilla complete but
weakly sclerotized; a narrow, flatly arched band.
Harpe with costa sclerotized throughout but not pro-
duced at apex; a fine brush of long hairs arising from
inner surface along lower edge of basal half of sclero-
tized costa; cucullus simple, narrow, tapering slightly
to rounded apex; from near base of harpe an appressed,
stout, thorny or serrate clasper. Penis armed with
two, stout, rather short cornuti less than one-half the
length of aedeagus. Vinculum stout, as long as or
somewhat longer than greatest width, its terminal
margin bluntly rounded or narrowly truncate.
Female genitalia without signum; bursa with a couple
of conspicuous round or oval, strongly pigmented and
sclerotized, densely granulate patches, otherwise bursa
finely spinose over its membranous areas; ductus bursae
shorter than bursa, not sclerotized adjacent to bursa,
but with strong sclerotization at broadened genital
opening; ductus seminalis from a lobe of bursa near
junction of bursa and ductus bursae.
This genus, while distinct from, is very close to
Nephopteryz, from which it is distinguished only by the
122 UNITED STATES NATIONAL MUSEUM BULLETIN 207
peculiar development of the clasper on harpe of the
male genitalia and the strong sclerotization about the
genital opening of the female. Hampson’s Hmmerita
has not a single character to separate it from Meroptera.
The stalking of vein 10 of forewing with 8-9, upon
which Hampson evidently relied, is shared by the types
of both Meroptera and Nephopteryr. This stalking is
very short at most and is not even specifically constant
in either genus.
The genus as here defined contains but four North
American species, and (to the best of my knowledge)
no Old World representatives.
246. Meroptera mirandella Ragonot
Figures 21, 333, 816
Meroptera mirandella Ragonot, Monograph, pt. 1, p. 313, 1893.
Eimmerita mirandella (Ragonot) Hampson, Ann. Mag. Nat. Hist.,
ser. 10, vol. 5, p. 76, 1930.—McDunnough, Check list,
No. 6181, 1939.
Forewing ocherous white with a very faint and
scattered dusting of blackish scales in median area;
antemedial line narrow, oblique, indented at lower fold,
bordered outwardly by a black line which begins as an
enlarged, more or less triangular dash on costa, in-
wardly by a straight black line from top of cell to inner
margin; subterminal line obscure, dentate-sinuate,
margined inwardly and outwardly for a short distance
from costa by black lines, on well marked specimens the
inner line continued as a fine dark border to tornus;
also on well marked examples a faint blackish or
fuscous shade extending obliquely across the wing from
the inner costal edge of the antemedial line; discal dots
faint, but usually distinguishable, blackish, separated,
obliquely placed; terminal black dots more or less con-
fluent. Hind wing white with a faint creamy or smoky
tint; a very faint brownish line along termen; veins not
or very slightly darkened. Alar expanse, 23-26 mm.
Male genitalia with broad, irregularly fan-shaped
clasper. Female genitalia with sclerotization of genital
opening a narrow, corrugate, sclerotized band with
short lateral, inward projections; bursa with two
opposed, lateral, granulate patches.
TypE Locatity: Colorado (type in Paris Mus.).
Foop piant: Populus (this record from an Arizona
specimen in the National Collections reared from a
cocoon on a cottonwood leaf).
Distrisution: Colorado, Denver; Arizona, Douglas
(Aug.), Nogales (July), Phoenix (Apr.), Pinal Mts.
(May), Readington, Tucson (Apr.), Yuma (June).
One of the females before me from Phoenix is de-
cidedly abnormal in venation, the forewing having
vein 10 from the cell and closely approximate to the
stalk of 8-9, 6 bent at base and from very near the upper
angle of the cell and 4 and 5 closely approximate at base
and for some little distance from the cell. Its genitalia
are normal and it is clearly an individual aberration,
but an example of what occurs all too often in the
Phycitidae and which should caution us to use generic
keys with discretion and to place unusual specimens
only after examination of their genitalia.
247. Meroptera cviatella Dyar
Fieure 817
Meroptera cviatella Dyar, Proc. Ent. Soc. Washington, vol. 7, p.34,
1805.—MecDunnough, Check list, No. 6182, 1939.
Salebria cviatella (Dyar) Forbes, Cornell Mem. 68, p. 627, 1923.
Forewing bright reddish brown with a more or less
distinct, transverse purplish gray shading in outer area
and bordering outwardly the subterminal line; this
latter area sometimes dusted with blackish scales; the
central area near antemedial line more or less dusted
with whitish scales; antemedial line oblique, narrow,
strongly indented at lower fold, its upper half obscured;
the white line bordered outwardly by a conspicuous
black band, broad on costa and very gradually narrow-
ing towards inner margin; inwardly bordered on lower
half by a similar broad, vertical black band; subterminal
line diffused, whitish; discal dots confluent, black, the
lower one sometimes obscured; a row of more or less
confluent black dots along termen. Hind wing pale
smoky brown, darkened slightly towards termen. Alar
expanse, 23-25 mm.
Male genitalia similar to those of mirandella. Fe-
male genitalia with sclerotization of genital opening a
narrow, dark, granulate band, broken and with two
short, divergent extensions at middle, these extensions
fusing into a small, thin, triangulate, sclerotized patch
on the lower median surface of the ductus bursae.
Typx Locatity: Chicago, Ill. (type in USNM).
Foop pianr: Populus (cottonwood). Larva a borer
in buds and new shoots.
Distrizution: Illinois, Chicago (June, July), Lacon
(Aug.), Putnam County (July, Aug.); Mississippi,
Starkville (July).
The above food-plant record is from Putnam County
specimens reared by Mr. Murry O. Glenn. One of
his series is labeled ‘bred from larva on Amorpha
canescens.”’ IJ doubt if Amorpha is a true food plant or
the presence of the larva on this plant anything more
than an accidental last-stage migration from cotton-
wood.
248. Meroptera pravella (Grote)
Figures 22, 334, 812
Pempelia pravella Grote, Bull. U. S. Geol. Geogr. Surv. Terr.,
vol. 4, No. 3, p. 694, 1878.
Meroptera pravella (Grote), Canadian Ent., vol. 14, p. 30, 1882.—
Packard, U. S. Dep. Agr. Div. Ent. Bull. 13, p. 23, 1887;
U. S. Dep. Agr. Fifth Rep., Ent. Comm., p. 574, 1890.—
Ragonot (in part), Monograph, pt. 1, p. 314, 1893.—Mc-
Dunnough, Check list, No. 6184, 1939.
Forewing dark gray; the basal area, except for a
narrow dark gray shading at extreme base, pale ashy
gray and contrastingly paler; antemedial line obscure
and often partially obliterated, when distinguishable
it is oblique, narrow, dull white, distinguished chiefly
by its black borders, which tend to coalesce into a
broad, diffused, blackish band; subterminal line faint
but distinguishable, grayish white, sinuate, in fresh
specimens bordered inwardly by an obscure, narrow,
blackish line; discal dots separated, blackish, not con-
spicuous; a faint blackish line along termen. Hind
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 123
wing very pale smoky fuscous.
mm.
Male genitalia with an elongate, stout, tapering
clasper extending about half the length of harpe;
figured from male from Edmonton, Alberta, and com-
pared by Clarke with the genitalia of the type of
pravella in the British Museum. Female genitalia with
sclerotization at genital opening in the form of a broad,
stout, curved, granulate and wrinkled, lunate plate
with strong lateral arms projecting into the ductus
bursae.
TypPE Locatiry: Oldtown, Maine (type in BM).
Foop piant: Populus, Salix [?].
Disrrisution: Unitep States: Maine; New Hamp-
shire, Hampton (July); New York, Ilion (June); Colo-
rado, Chimney Gulch (Golden, June). Canapa: Que-
bec, Knowlton (Feb., reared in laboratory from larva
on aspen), Norway Bay (June); Ontario, Constance
Bay (Feb., from poplar), Grand Bend (July), Hymers
(June), Mer Bleue (June), Ottawa (Mar., from poplar),
Smoky Falls (Mattagami River, June), Trenton (June);
Manitoba, Aweme (May, June, July), Winnipeg; Sa-
skatchewan, Indian Head (June, July); Alberta, Bilby
(June, July), Edmonton (May, July), Nordegg (July);
British Columbia, Canim Lake (June), ‘100 Mile
House’”’ (June).
These records (except for the type locality) are from
specimens before me from the U. S. and Canadian
National Collections. The Salix plant record is from
Packard. I have not seen the reared Brunswick,
Maine, specimens upon which it was based but have
little doubt that the name pravella was correctly ap-
plied. Most of the specimens in our collections that
have been identified as pravella as well as many of the
references in literature are referable to Nephopteryx
subfuscella (Ragonot) (=semiobscurella (Hulst)). This
confusion is discussed under subfuscella. The two
species are easily confused, especially with worn exam-
ples, if their genitalia are not examined; but in unrubbed
specimens pravella is easily separated from subfuscella
by the lack of any reddish or reddish ocherous scaling
adjacent to inner margin at the base of forewing. Also
included under pravella in the U. S. and Canadian
National Collections were 21 Canadian examples of a
new species (hereinafter described as abditiva) similar
in all superficial characters to pravella but with dis-
tinctly different male and female genitalia.
Alar expanse, 20-22
249. Meroptera abditiva, new species
Ficures 335, 813, 814
In color and maculation like pravella but with con-
sistently different genitalia. Alar expanse, 19-22 mm.
Male genitalia with harpe having a stout but short,
stubby clasper, less than one-fourth the length of harpe.
Female genitalia with the sclerotization at genital open-
ing in the form of a rather narrow granulate, curved,
transverse band without inwardly projecting arms.
The pigmented, granulated area of the bursa is individ-
ually variable in extent and sometimes divided into two
distinct patches by a slight break at the anterior
(closed) end of the bursa. Extremes of variation are
shown in figures 813 and 814. However, there are no
intergradations whatever between abditiva and pravella
in the structure of the female genital plate nor in the
male clasper.
Typp Locatity: Knowlton, Quebec (type in Cana-
dian Nat. Coll.; paratypes in Canadian Nat. Coll. and
USNM, 61344).
Foop piant: Populus tremuloides.
Described from male type and 3 female paratypes
from the type locality reared (in laboratory) Feb. 12,
1 and 2, 1930, from larvae feeding on leaves of P. tre-
muloides (“aspen’’) by J. McDunnough; and 17 para-
types from the following Canadian localities: Quebec,
Mount St. Hilaire, June 30, 1908, G. Chagnon (9);
Norway Bay, June 4, 1938, E. G. Lester (@). New
Brunswick, Chamcook, June 23, 1938, T. N. Freeman
(9); Eel River, June 21, 1941, T. N. Freeman (<*).
Nova Scotia, Beddeck, June 23, 1936 and June 27, 1938,
T. N. Freeman (29); White Point Beach, Queens, Feb.
12 and 20, 1936, J. McDunnough (2 9, reared).
Ontario, Trenton, May 29 and June 25, 1908, Evans
(2 9); Vineland Station, June 15, 1936, W. L. Putnam
(2, reared). Saskatchewan, Christopher Lake, June
19, 1939, A. R. Brooks (). British Columbia,
Canim Lake, June 25, 1938, J. K. Jacob (9); Jesmond,
July 13, 1937, J. K. Jacob (2 9); Kaniloops, June 14,
1937, J. K. Jacob (@); Shingle Creek, Penticton,
June 25, 1935, A. K. Cartrell (9).
In as much as pravella and abditiva have the same
hosts and larval habits, an overlapping distribution,
and similar habitus, it is necessary to examine their
genitalia to distinguish them apart.
68. Genus Nephopteryx Hiibner
Nephopieryx Hiibner, Verzeichniss bekannter Schmett[er]linge,
p. 370, 1825.—Zeller, Isis von Oken, 1846, p. 731.—Grote,
Bull. U. 8. Geol. Geogr. Surv. Terr., vol. 4, p. 695, 1878;
North Amer. Ent., vol. 1, p. 11, 1879.—Ragonot, Ent.
Monthly Mag., vol. 22, p. 19, 1885 (citation of type); Mono-
graph, pt. 1, p. 254, 1893.—Hulst, Phycitidae of N. Amer.,
p. 142, 1890.—Forbes, Cornell Mem. 68, p. 622, 1923.—
Hemming, Hubner, vol. 2, p. 229, 1937.—Bisset, in Pierce
and Metcalfe, Genitalia of the British Pyrales, p. 61, 1938.—
Janse, Journ. Ent. Soc. South Africa, vol. 5, p. 34, 1942.
(Type of genus: Phycita rhenella Zincken; Europe; figs. 25,
336, 815.)
Sciota Hulst, Ent. Amer., vol. 4, p. 115, 1888.
Sciota croceella Hulst.)
Characters of Meroptera except: Labial palpus erect
or obliquely upturned. Maxillary palpus of male in
the form of an aigrette or squamous. Forewing with
10 usually connate with the stalk of 8-9 or closely
approximate to it, rarely stalked. Transtilla frequently
incomplete or absent. Clasper of harpe digitate, slen-
der, simple (without spining). One cornutus on penis in
uvinella, other species have two cornuti as in Meroptera.
Female genitalia with ductus bursae sclerotized along
ventral surface for most of its length from junction with
bursa, the sclerotization terminating before genital
opening, the latter simple (unsclerotized).
(Type of genus:
124
As here defined the genus includes a number of
species with two types of maxillary palpi, several of
which have been hitherto referred to either Salebria or
Myrlaea. The reference of some of our North American
species to the latter genus on the strength of a slight
obliqueness of their labial palpi is not warranted as
none of them agrees with the Old World type of Myrlaea
(albistrigella Staudinger) on either male or female
genitalic characters. The ductus bursae and bursa of
albistrigella are perfectly smooth, without granulations,
scobinations, or sclerotizations of any kind; and its
harpe lacks the hair brush characteristic of Meroptera
and Nephopterysx.
The two types of male maxillary palpi (aigrettelike
and squamous) do not justify generic separation of the
species here any more than they do in Dioryctria or
Ortholepis although such a difference should be, and in
the Phycitidae usually is, of generic significance. On
basilaris, the palpus is midway between the two types,
though somewhat more aigrettelike than squamous.
The uniform type of their male and female genitalia and
the similar habitus of the included species indicate a
distinct and natural group. Nephopteryz is very close
to Meroptera but is distinguished by its simple, slender
clasper, simple female genital opening, and differently
sclerotized ductus bursae.
Genus Nephopteryxz, Species 250-267: N. sub-
fuscella to N. celtidella
[Males with aigrettelike palpi.]
250. Nephopteryx subfuscella (Ragonot), new combination
Fieures 337, 821
Salebria subfuscella Ragonot, N. Amer. Phycitidae, p. 8, 1887;
Monograph, pt. 1, pp. 329, 350, 1893.—Hulst, Phycitidae
of N. Amer., p. 151, 1890.—McDunnough, Check list, No.
6123, 19389.
Salebria semiobscurella Hulst, Phycitidae of N. Amer., p. 151,
1890.—Ragonot, Monograph, pt. 1, p. 352, 1893.—Barnes
and McDunnough, Contributions, vol. 3, p. 197, 1916.—
Forbes, Cornell Mem. 68, p. 626, 1923.—McDunnough,
Check list, No. 6212, 1939.—Craighead, U. S. Dep. Agr.
Misc. Publ. 657, p. 454, 1950. (New synonymy.)
Meroptera pravella (authors not Grote) Hulst (in part), Phyci-
tidae of N. Amer., p. 148, 1890.—Ragonot (in part), Mono-
graph, pt. 1, p. 314, 1893.—Grossbeck, Bull. Amer. Mus.
Nat. Hist., vol. 37, p. 180, 1917.—Forbes, Cornell Mem. 68,
p. 624, 1923.
Color and maculation resembling those of Meroptera
pravella. Forewing gray; the basal area contrastingly
paler with some dark shading at extreme base and more
or less reddish or orange scaling on base of inner margin
(at least a trace on all except badly rubbed specimens);
antemedial line obscure, indicated chiefly by its fused
dark inner and outer borders which form a rather broad,
oblique, blackish band, the antemedial line itself distin-
guishable on most specimens only as an incomplete, me-
dian, pale (whitish) streak in the dark band ;subterminal
line obscure but complete, narrow, sinuate, pale gray with
fine dark bordering lines; some whitish dusting over
central area of wing, especially on pale examples; discal
UNITED STATES NATIONAL MUSEUM BULLETIN 207
dots blackish, occasionally separated, more often fused
into a curved line; dots along terminal line fine, weak,
blackish, usually separated, on some specimens con-
fluent. Hind wing pale brown to smoky fuscous; veins
darkened, especially on the darker females; a narrow
dark shade along termen. Alar expanse, 18-22 mm.
Male genitalia with apical process of gnathos triangu-
late. Clasper bent downward, parallel to surface of
harpe. Female genitalia with granulate sclerotized
patch on bursa a continuous band across posterior
ventral surface and forward on right side of bursa.
Type tocauity: Not given (subfuscella, in Paris
Mus.); Blanco County, Tex. (semiobscurella, in AMNH,
ex Rutgers).
Foop piants: Rhus, locust [7]. Larva a leaf-folder.
Distripution: Unitrp statrs: Maine, Augusta
(June); Massachusetts, Martha’s Vineyard (May, July);
Connecticut, East River (June); Rhode Island, Weeka-
paugh (Aug.); New York, Ilion (June), Rossville (Long
Island, Feb., Mar.), Shore of Lake Ontario (near
Rochester, May, June); Pennsylvania, Oak Station
(Aug.); District of Columbia, Washington; North Caro-
lina, Tryon (May); Florida, Fort Myers (May), Lake
Alfred (July); Texas, Burnet County (Apr.), Kerrville
(May, Aug.); Missouri, Kirkwood (Mar., Apr., May);
Iitinois, Decatur (July, Aug.); Washington, Almota
(July), Bellingham (Nov.), Pullman (Feb., May, June,
July, Aug., Nov.), Snake River (Jan., Feb., July),
Walla Walla (uly), Wawawai (May). Canapa: Que-
bec, Levis, Meach Lake (July), Mount St. Hilaire (July);
Ontario, Merivale (Mar.), Ottawa (Mar.), Trenton
(May, June, July).
Many of the above records are from reared examples
from sumac, and all such had been identified by Riley
and Dyar as Meroptera pravella. Barnes and McDun-
nough (1916) were the first to question and correct this
identification, assigning the name Salebria semiobscurella
Hulst to the sumac feeder. At that time subfuscella
had not been recognized by American lepidopterists.
It resembles pravella but has the reddish scaling on base
of inner margin of forewing. Unquestionably it is the
same as what Hulst later described as semiobscurella.
No other American species that could have been referred
to Meroptera or Salebria has this red-scale character
with the habitus, otherwise, of pravella. The food
plant record of dried peaches (‘‘peches desséchées’’) given
for subfuscella by Ragonot (Monograph, pt. 1, p. 352)
on the basis of two imperfect specimens sent him so
labeled by Riley is obviously incorrect and can be
ignored. I question also “locust” as a probable or even
occasional food plant. That record, given above, is
based on four males from the Fernald collection that
had been identified by Dyar as pravella. ‘They had been
reared (Apr. 1889, May 1893) from larvae collected by
Miss Murtfeldt at Kirkwood, Mo., onlocust. Her note
(“‘324M’’) states that most of the larvae in the lot were
“‘Salebria contatella Grote” but that a few seemed to be
a different species. I suspect that the latter (the sub-
fuscella larvae) had merely migrated to the locust after
feeding on nearby sumac.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 125
251. Nephopteryx delassalis Hulst
Fieure 818
Nephopteryx delassalis Hulst, Trans. Amer. Ent. Soc., vol. 13,
p. 161, 1886.
Salebria purpurella Hulst, Canadian Ent., vol. 24, p. 61, 1892.—
Forbes, Cornell Mem. 68, p. 627, 1923.
Salebria pudibundella Ragonot, Monograph, pt. 1, p. 350, 1893.—
McDunnough, Check list, No. 6208, 1939. (New synonymy.)
Myelaea delassalis (Hulst) Barnes and McDunnough, Contribu-
tions, vol. 3, p. 198, 1916.—McDunnough, Check list, No.
6225, 1939.
Thorax vinous red with a scattered dusting of white
and black scales. Forewing vinous red with a more or
less diffused bluish white shading in basal area immedi-
ately preceding inner border of the antemedial line;
this border a broad black band (the most conspicuous
marking on the wing), vertical, and fusing at costa with
the outer black border; the antemedial line itself faint,
but on most specimens its lower half distinct, narrow,
white, slightly oblique and inwardly angulate at lower
fold, bordered outwardly by a rather narrow, inter-
rupted black band; some blackish dusting on the whitish
subbasal area, black scaling along lower margin of cell,
on some of the lower veins from cell and on costa, espe-
cially towards apex; discal dots (when distinguishable)
separated, red, more or less shaded with black; dots
along termen very faint, blackish, more or less confluent.
Hind wing pale ocherous fuscous. Alar expanse, 22-
26 mm.
Male genitalia similar to those of fernaldi; transtilla
absent; apical process of gnathos narrow (not triangu-
late); clasper short, slightly curved. Female genitalia
with two granulate patches on bursa copulatrix, a small
one on posterior dorsal surface, near the left side of
bursa and a larger on ventral surface at the antero-
lateral margin (closed end) of bursa.
Typr Locatities: Nevada (delassalis, in AMNH, ex
Rutgers); New Mexico (purpurella, in AMNH, ex Rut-
gers); Colorado (pudibundelia, in Paris Mus.).
Foop piant: Unknown.
Distrisution: Unirep Sratrs: Arizona, White
Mts; Colorado, Beulah (June), Fort Collins; Utah,
Vineyard (June, July) ; Nevada; California, Inyo County
(May, June, July).
The types of delassalis and purpurella are both fe-
males. Their genitalia are alike. Barnes and McDun-
nough (1916) first noted this synonymy and the mis-
identification of his species by Hulst (1900) and the
consequent misapplication of the name delassalis to
specimens of fernaldi Ragonot. The descriptions under
delassalis in Hulst’s 1900 revision and in Ragonot’s
monograph apply to fernaldi and not delassalis. On
the other hand, Ragonot’s description of pudibundella
applies in detail to the true delassalis.
The venation of forewing is individually variable, as
with many species of Nephopteryz, vein 10 being either
connate or closely approximate at base to the stalk 8-9
(rarely short stalked with it) and veins 4 and 5 either
connate or closely approximate at base.
252. Nephopteryx delassalis fraudifera, new race
Superficially appears quite distinct from delassalis;
the entire median area and much of the basal area of
forewing being heavily dusted with whitish, giving the
general ground color a whitish blue-gray shade similar
to that of inconditella rather than the vinous red of
typical delassalis; the vinous red limited in fraudifera
to the thorax, extreme base of forewing, a broad band
outwardly bordering the subterminal line, and a faint
diffused shading just preceding it. The red shade
somewhat darker than in typical delassalis; the black
borders of antemedial line also somewhat broader, es-
pecially at costa. Alar expanse, 24-26 mm.
Male and female genitalia agreeing in all details with
those of delassalis.
Typr Locauiry: Oliver, British Columbia, (type in
Canadian Nat. Coll.; paratypes in USNM, 61345, and
Canadian Nat. Coll.).
Foop piant: Unknown.
Described from male type and one female paratype
from the type locality (June 8 and 9, 1923, G. B. Gar-
rett, collector) ; and paratypes from the following locali-
ties: “Shingle Cr. Road,” Keremeos, British Columbia
June 22, 1935, A. N. Cartrell (9); Salmon Arm, British
Columbia, June 20, 1916, ‘“W. R. B.”’ (9); Kaslo, Brit-
ish Columbia, June 13, 1903, H. G. Dyar, “19366” (9);
Alberni, British Columbia, June 20, 1922, ““‘W. R. B.”
(@), and Bellingham, Wash., May 30, 1922, J. F. G.
Clarke ().
Presumably a distinct food plant as well as local race.
253. Nephopteryx rubescentella (Hulst)
Mineola rubescentella Hulst, Canadian Ent., vol. 32, p. 169, 1900.
Nephopteryx rubescentella (Hulst), U. S. Nat. Mus. Bull. 52, p.
419, 1903.—MecDunnough, Check list, No. 6173, 1939.
Ground color of forewing slightly paler than that of
typical delassalis. Thorax also paler, pale purplish
gray or grayish ocherous. The dark bands bordering
antemedial line on forewing dull red or reddish orange,
containing no black except for occasional scattered
scales. Maculation otherwise as in typical delassalis.
Alar expanse, 26 mm.
Male genitalia similar to those of fernaldi.
Typm LOCALITY: Tennessee (type in USNM).
Foop prant: Unknown.
In addition to the male type, the National Collection
contains a male from Denver, Colo. (Aug.). I have
seen no other specimens. A female from the type
locality will be needed before the exact status of rubes-
centella can be determined. It may be no more than
a race or variety of fernaldi.
254. Nephopteryx fernaldi (Ragonot), new combination
Friaures 340, 819
Salebria fernaldi Ragonot, N. Amer. Phycitidae, p. 9, 1887.
Salebria delassalis Hulst (not Hulst), Phycitidae of N. Amer.,
p. 154, 1890.
Myrlaea delassalis (Hulst not Hulst) Ragonot, Monograph, pt. 1,
p. 402, 1893.
Myrlaea fernaldi (Ragonot) McDunnough, Check list, No. 6226,
1939.
126
Forewing whitish ocherous with basal half of costa
faintly washed with reddish ocherous, also some traces
of this shading on the pale ocherous thorax and along
inner margin of forewing from base to antemedial line,
strongest on costa just above the dark borders of the
antemedial line; some blackish dusting along costa near
apex and occasionally on a few of the veins; antemedial
line obscure, broken, frequently obscured entirely by
its black borders; the latter forming a broad, strongly
contrasted black blotch which extends from inner
margin to top of cell (not reaching costa); discal dots
much reduced or absent, if present more or less con-
fluent. Hind wing pale whitish ocherous; the veins
very slightly darkened. Alar expanse, 22-25 mm.
Male genitalia figured from Colorado specimen identi-
fied by Hulst as delassalis (one of his spurious “‘types’’).
There are no structural differences to distinguish the
male genitalia of the two species.
Female genitalia with a single, rather large, granulate
patch in bursa, on left side and extending onto ventral
surface near posterior end of bursa; the bursa copula-
trix itself as broad as long, approximately round, the
membrane at anterior end thickened (cartilaginous).
Type Locauity: Arizona (type in Paris Mus.).
Foop prant: Unknown (probably Amorpha sp.).
Distripution: Unirep States: Arizona; Colorado,
Denver, Glenwood Springs (July), also two males and
a female with only the state locality, identified by Hulst
as ‘delassalis’’; Kansas, Thomas County; Jowa, Ames.
Canava: Manitoba, Cartwright (June, July), Winnipeg.
The female genitalia and ground color of thorax and
forewing easily separate this species from delassalis,
with which it has been confused due to Hulst’s later
(1890) misidentification of his own species.
255. Nephopteryx dammersi, new species
Figures 338, 822
Ground color pale ashy gray strongly shaded with
pale rust red on thorax, at extreme base, on basal half
of costal edge and on basal third of inner margin of
forewing; more or less of this red shading on the black
inner border of the antemedial line and forming its
outer border near costa; an obscure, ill-defined, rusty
blotch on middle of lower fold; antemedial line distin-
guishable on most specimens, narrow, whitish, slightly
oblique and indented between cell and inner margin
(the antemedial line, where it can be distinguished for
any appreciable distance, has a similar configuration
and slant on most of our American Nephopteryz),
bordered outwardly on lower half by a narrow, inter-
rupted black line and inwardly by a broad black band,
the latter extending only from inner margin to middle
of cell and (as noted above) more or less shaded with
rust red; subterminal line obsolete or nearly so; discal
dots usually distinct, especially the lower one, sepa-
rated, blackish; terminal dots minute, very faint, not
UNITED STATES NATIONAL MUSEUM BULLETIN 207
confluent. Hind wing dull white with a faint ocherous
tint on male; pale smoky fuscous on female; the veins
faintly darkened and a faint, narrow, dark shade along
termen. Alar expanse, 25-26 mm.
Male genitalia similar to those of fernaldi. There is
some difference in the armature of the penis between the
two species; the cornuti are somewhat shorter and
stouter and there is a darker pigmentation of some of
the scobinations on penis in dammersi (fig. 338); but
there is so much individual variation of these structures
within any given species of Nephopteryx that they can
not be safely used to separate species. Female genitalia
with two small and one large granulate patch in bursa.
Type tocauity: Cajon Valley, San Bernardino
Valley, Calif. (type in USNM, 61346).
Foop pLant: Amorpha californica.
Described from male type and three male and six
female paratypes from the type locality (reared by
Commander J. Dammers Apr. 15, 1933, and July 20,
30, 1932), and one female from the Huachuca Mts.,
Ariz. (“July 8-15’). In addition I have before me a
male from Douglas County, Ariz. (July 20, 1940,
collected by Fritz Forbes), which appears to be con-
specific. It is a trifle smaller (24 mm.) than the speci-
mens of the type series. Without a matching female
from the same locality it appears unwise to include it
among the paratypes.
The species is named in honor of Commander
Dammers, who has given many fine reared and collected
specimens to our National Collection. It is close to
fernaldi, but on female genitalic characters seems to be
a distinct species and not a color form or local race.
256. Nephopteryx dammersi floridensis, new race
Smaller and considerably darker than typical dam-
mersi; the ground color dark gray finely peppered with
white especially in median and subbasal areas making
these areas a trifle paler than remainder of wing; the
rust red markings of typical dammersi replaced by
lavender-red in floridensis and this color more extended,
forming a faint suffusion over much of the median and
outer areas in addition to the stronger markings on
costa, inner margin and extreme base; subterminal line
distinct and with narrow dark borders. Hind wing
smoky fuscous; the veins darkened; a narrow blackish
line along termen. Alar expanse, 21-22 mm.
Genrrauia: Male and female as in typical dammersi.
TypE Locauity: Williamsburg, Fla. (type in USNM,
61347).
Foop PpLant: Amorpha herbacea.
Described from male type from the type locality,
reared under S. S. No. 16970A, June 25, 1944; and one
female paratype from Tampa, Fla., reared under S. S.
No. 16859, June 29, 1944. Larva collected and both
specimens reared by members of the Special Survey of
the Division of Foreign Plant Quarantine of the U. S.
Bureau of Entomology and Plant Quarantine.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 127
257. Nephopteryx vetustella (Dyar), new combination
FicurE 820
Salebria vetustella Dyar, Journ. New York Ent. Soc., vol. 12,
p. 106, 1904.—Forbes, Cornell Mem. 68, p. 626, 1923.
Myrlaea vetustella (Dyar) Barnes and McDunnough, Check list
of the Lepidoptera of Boreal America, No. 5650, 1916.—
Forbes, Cornell Mem. 68, p. 627, 1923.—McDunnough,
Check list, No. 6224, 1939.
Forewing pale ash gray with a slight purplish shading
over outer half; basal area almost to inner dark border
of antemedial line, Indian red or reddish orange, this
reddish shade also on collar and top of head; antemedial
line distinct from upper vein of cell to inner margin,
narrow, oblique, inwardly notched between cell and
inner margin, bordered inwardly by a broad black band
and outwardly by a narrow black line, the black borders
continuous to and fused at costa; subterminal line
usually distinct but faint and with a weak dark inner
bordering line, sinuate; discal dots separated, small,
blackish; a weak row of blackish dots along termen.
Hind wing pale smoky brown; the veins and terminal
margin slightly darkened. Alar expanse, 22-25 mm.
Male genitalia similar to those of fernaldi. Female
genitalia with a single long granulate patch in bursa
extending nearly the length of the bursa on its left side
(seen from below) and curving onto ventral surface near
junction of bursa and ductus bursae.
Typr Locauity: Plummers Isl., Md. (typein USNM).
Foop puant: Unknown (Amorpha sp. ?).
Disrrisution: Unirep Sratses: Maryland, Plum-
mers Isl. (Apr., May); Pennsylvania, Oak Station
(June), Pittsburgh (May, June) ; New York, Ilion (May) ;
Illinois, Edgebrook (June), Elkhart (Aug.), Palos Park
(June); Iowa, Iowa City (June), Sioux City (July);
Missouri, St. Louis; Florida, St. Petersburg (June).
Canapa: Ontario, Ottawa (June, July); Quebec, Meach
Lake (June).
This species forms one of a group of closely related
species (delassalis to vetustella) that feed upon Amorpha
and have similar male genitalia but differ markedly in
the number, arrangement, and relative sizes of the gran-
ulate patches of the female bursa. These differences
appear to be constant and the specific character holds
even in series of individuals from widely different
localities. The male genitalia exhibit some minor dif-
ferences, especially in the relative size and position of the
two cornuti; but here individual variation makes the
apparent differences untrustworthy for specific differ-
entiation. During dissection of the male organs a
simple transtilla can be faintly seen in all the species of
the group, but it is so weakly sclerotized that it is not
visible or but partially distinguishable in balsam mounts.
The structure is obsolescent and cannot be considered
“present”? in the sense in which it is in Meroptera for
example or even in a few other species of Nephopteryz.
258. Nephopteryx inconditella (Ragonot), new combination
Figure 825
Salebria contatella inconditella Ragonot, Monograph, pt. 1,
348, 350, 1893.—McDunnough, Check list, No. 6205b, 1939,
Thorax and forewing pale ash gray with a faint bluish
tint; some reddish scaling at extreme base of wing and a
small spot of the same shade at middle of lower fold;
pale antemedial line distinguishable from inner margin
to cell, deeply notched at vein 1b, its black borders
complete from inner margin to costa, the inner border
broad and fusing with the narrow outer one near costa;
subterminal line distinct, sinuate-serrate, narrowly bor-
dered by obscure blackish lines; discal spots obscure but
usually distinguishable, separated; terminal dots weak,
confluent; on most specimens a narrow, dark transverse
shade extending from costa at inner edge of subterminal
line to middle of inner margin. Hind wings translucent,
whitish with a faint ocherous tint, the veins slightly
darkened; a narrow brownish shade along terminal
margin. Alar expanse, 25-30 mm.
Male genitalia of the fernaldi type but clasper some-
what more strongly sclerotized than in the seven preced-
ing species. Female genitalia with a single, small,
round, granulate patch on ventral surface of bursa very
near junction of bursa and ductus bursae.
Type Locauity: Colorado (type in Paris Mus.).
Foop prant: Unknown.
Distrisution: Colorado; Arizona, Huachuca Mts.,
Palmerlee, and four Arizona specimens with only the
state locality and without dates, one of these a pseudo-
type of ‘Pinipestis albovittella Hulst.”’
The species was described as a pale western variety of
contatella (Grote), to which it is apparently most nearly
related and which, except for its paler ground color, it
resembles. Its female genitalia however clearly indi-
cate a distinct species. The ground color of forewing
is intermediate between that of subcaesiella (=conta-
tella) and that of dammersi.
259. Nephopteryx subcaesiella (Clemens), new combination
Fiaurus 339, 826
Pempelia subcaesiella Clemens, Proc. Acad. Nat. Sci. Philadel-
phia, p. 206, 1860.
Pempelia contatella Grote, North Amer. Ent., vol. 1, p. 49, 1880.—
Comstock, in Rep. [U. S.] Comm. Agr. for 1880, p. 261, 1881
(in part).
Salebria contatella (Grote), Bull. U. S. Geol. Geogr. Surv. Terr.,
vol. 6, no. 3, p. 590, 1882.—Beutenmiiller, Canadian Ent.
vol 22, p. 16, 1890 (larva).—Hulst, Phycitidae of N. Amer.,
p. 152, 1890; U. S. Nat. Mus. Bull. 52, p. 424, 1903.—
Packard, U. S. Dep. Agr. Fifth Rep. Ent. Comm., p. 361,
1890.—Ragonot, Monograph, pt. 1, p. 348, 1893.—Forbes,
Cornell Mem. 68, p. 626, 1923.—McDunnough, Canadian
Ent., vol. 78, p. 109, 1946.
Salebria virgatella subcaesiella (Clemens) Barnes and McDun-
nough, Check list of the Lepidoptera of Boreal America, No.
5631a, 1916—McDunnough, Check list, No. 6205a, 1939.—
Craighead, U. 8. Dep. Agr. Mise. Publ. 657, p. 453, 1950.
Color and maculation similar to those of inconditella
but the ground color of forewing a much darker gray,
basal area contrastingly paler gray; the reddish scaling
on base of wing ranging from wine to rusty ocherous,
always present but sometimes reduced to a few scales,
rarely extended for a short distance onto the thorax;
inner black border of antemedial line somewhat broader
and more diffused than on inconditella, complete to costa
128 UNITED STATES NATIONAL MUSEUM BULLETIN 207
(unbroken at any part of its length) ; following the ante-
medial line 2 pale grayish spot (sometimes very faint but
usually more contrasted than in inconditella); discal
spots distinct, black, separated. Hind wing smoky
white to smoky fuscous or brown, darker on females
than on males, a distinct dark shade along termen.
Alar expanse, 21-28 mmi
Male genitalia as in inconditella and virgatella except
for the armature of the penis. In both inconditella and
virgatella there are two moderately stout cornuti situ-
ated on penis, one to the side and slightly behind the
other. In subcaesiella the second cornutus is greatly
reduced. McDunnough (1946) states that there is only
one cornutus in subcaesiella (contatella). That could
easily be the case on individual specimens, but the
normal condition is two cornuti on penis. Every prep-
aration J have seen shows at least a vestige of the
second cornutus. In all three species the lateral ele-
ments of transtilla are indicated, but very weakly
sclerotized. Female genitalia without granulate patch
or patches on bursa.
Typr Locanitius: Not given, presumably Pennsyl-
vania (subcaesiella, in Acad. Nat. Sci. Philadelphia);
“New England” (contatella, i BM).
Foop piant: Robinia pseudoacacia. Wisteria also
recorded as food plant.
DistrisutTion: Unirep States: Maine, Augusta
(May, June), Orono; New Hampshire, Hampton (July);
Massachusetts, Amherst (June), Framingham (May),
Martha’s Vineyard (July, Aug.), Newton Highlands;
New Jersey, Essex County Park (June), New Lisbon
(Aug.) ; Pennsylvania, New Brighton (May, July, Aug.),
Oak Station (June), Pittsburgh (May, June, July);
Maryland, Plummers Isl. (June, Aug.), Somerset Heights
(Aug.); District of Columbia, Washington (Apr., May,
July); Virginia, Falls Church (Aug.), Snickers Gap
(July); North Carolina, Black Mountain (July), Tryon
(May); Tennessee, no specific locality (May); Iihnois,
Decatur (July), Elkhart, Oconee (July); Jowa, Iowa
City (Aug.), Sioux City (June) ; Missouri, “Cent. Mo.”’
(Aug.), Kirkwood (Apr., May), St. Louis (June); Ar-
kansas, Washington County (‘“‘July—Aug.”). CanNapa:
Nova Scotia, Smith’s Cove (recorded by McDunnough,
1946).
I have not seen any Canadian examples but Mc-
Dunnough’s description leaves no doubt of what he had.
He notes the differences in the cornuti and larval
characters between subcaestella (contatella) and virgatella
quinquepunctella) and treats them as a distinct species.
The difference in their female genitalia and the con-
sistent difference in maculation of forewing are added
evidence that they are not merely races of one variable
species.
260. Nephopteryx virgatella (Clemens), new combination
Figure 827
Pempelia virgatella Clemens, Proc. Acad. Nat. Sci.
p. 205, 1860.
Pempelia contatella quinquepunctella Grote, North Amer, Ent.,
vol. 1, p. 50, 1880.—Comstock, in Rep. [U. 8.] Comm. Agr.
for 1880, pp. 261-262, 1881 (part; larva).
Philadelphia,
Salebria contatella quinquepunctella (Grote) Hulst, Phycitidae of
North Amer., p. 152, 1890; U.S. Nat. Mus. Bull. 52, p. 424,
1903.—Ragonot, Monograph, pt. 1, p. 348, 1893.—Forbes,
Cornell Mem. 68, p. 626, 1923.
Salebria virgatella (Clemens) Barnes and MceDunnough, Check
list of the Lepidoptera of Boreal America, No. 5631, 1916.—
McDunnough, Check list, No. 6205, 1939.
Salebria quinquepunctella (Grote) McDunnough, Canadian Ent.,
vol. 78, p. 109, 1946 (larva).
Ground color of forewing (on most specimens) 2
trifle paler than that of subcaesiella, gray with a faint
brownish or purplish tint; a reddish (or pale purplish)
brown shade along lower fold, cutting the antemedial
line and its black borders; a similar, narrower, shorter
streak on median fold; outer black border of ante-
medial line between top of cell and inner margin reduced
to two black dots, one on the lower margin of cell, the
other on vein 1b and enclosed within the pale patch
following the antemedial line; both these dots and the
black discal dots at end of cell well contrasted; sub-
terminal line faint, not appreciably darkly bordered,
Interrupted at the folds. Hind wing smoky white to
brown; the veins more or less darkened and a smoky
brown shade along termen. Alar expanse, 22-26 mm.
Male genitalia similar to those of inconditella; both
cornuti moderately stout and situated one to the side
of and slightly behind the other. Female genitalia
with two strong granulate patches on bursa, a rather
large one on middle of dorsal surface curving around
left side onto ventral surface, and a smaller ventral
patch near junction of bursa and ductus bursae.
TypE Locatiries: Not given, presumably Pennsyl-
vania (virgatella, in Acad. Nat. Sci. Philadelphia); New
York (quinquepunctella, in BM).
Foop piant: Robinia pseudoacacia.
Distrisution: UnrTrep States: Maine; Massachusetts,
Amherst (June), Martha’s Vineyard (June, July, Aug.);
New York, Orient (Long Island, Aug., Sept.) and one
specimen with only state locality; New Jersey, Essex
County Park (Aug.); Pennsylvania, Buena Vista (Aug.);
New Brighton (May, July); West Virginia, Jefferson
County (Aug.); District of Oolumbia, Washington
(June); Virginia, Berryville (May); North Caroline,
Tryon; ilinois, Elkhart, Putnam County (May); Mis-
souri, St. Louis; Arkansas, Washington County (‘“‘July—
Aug.”). Canapa: Ontario, London, Trenton (June);
Nova Scotia, Bridgetown (July), Smith’s Cove.
The differences separating virgatella from subcaesiella,
of which it was long considered only a variety, are dis-
cussed under the latter species.
261. Nephopteryx carneella Hulst
° Figure 823
Nephopteryx carneella Hulst, Ent. Amer., vol. 3, p. 131, 1887.
Nephopteryx inquilinella Ragonot, N. Amer. Phycitidae, p. 8,
1887; Monograph, pt. 1, p. 290, 1893.—Hulst, Ent. Amer.,
vol. 5, p. 156, 1889; Phycitidae of N. Amer., p. 145, 1890.—
McDunnough, Check list, No. 6171, 1939.
Salebria carneella (Hulst), Phycitidae of N. Amer., p. 153, 1890.—
Ragonot, Monograph, pt. 1, p. 367, 1893.—Barnes and Mc-
Dunnough, Contributions, vol. 3, p. 197, 1916.—Forbes,
Cornell Mem. 68, p. 627, 1923.—McDunnough, Check list,
No. 6204, 1939.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 129
Ground color of forewing bluish gray, the gray shade
most obvious in median area and as a narrow band
preceding the inner border of the antemedial line; base
maroon red, this shade more or less suffusing the gray
thorax; antemedial line obscure, bordered inwardly by
a broad madder-red band which extends to costa; a
similar red shade in outer area bordering the faint
sinuate subterminal line and some red suffusion over
median part of the lower fold; discal spots distinct,
separated, black; terminal dots very faint more or less
confluent. Hind wing smoky white with a faint yellow-
ish tint; somewhat darker on females; a fine brown line
along termen. Alar expanse, 20-23 mm.
Male genitalia of the fernaldi type; the two cornuti
lie side by side, one slightly shorter and more slender
than the other. Female genitalia with bursa and re-
mainder of genitalia considerably smaller than in pre-
ceding species; two granulate patches, one large, one
considerably smaller, placed opposite each other on
lateral margins of the bursa.
Typr LocALitiEs: Maine [?] (carneella, in AMNH,
ex Rutgers; the male type bears no locality label, but
in his original description Hulst gives New Mexico as
the type locality; this, as pointed out by Barnes and
McDunnough, is probably an error or pure guesswork
on Hulst’s part); Wisconsin (inquilinella, in Paris Mus.).
Foop piant: Saliz (carneella); galls of sawfly (Huura
Salicisnodum) on willow.
Distrisution: Unirep Sratrs: Maine, Monmouth
(July), and two examples with only state locality; Mas-
sachusetts, Amherst (June), Martha’s Vineyard (Apr.);
Indiana, Hessville (June); Wisconsin; Michigan, Dick-
inson County. Canapa: Ontario, Ottawa (June),
Trenton (July); Manitoba, Aweme (May).
Hulst was correct the first time (1889) in making
inquilinella 2 synonym of his carneella. The genitalia
of their male types are identical.
262. Nephopteryx basilaris Zeller
Figures 341, 829
Nephopteryx basilaris Zeller, Verh. zool.-bot Ges. Wein, vol. 22,
p. 548, 1872.—Grote, N. Amer. Ent., vol. 1, p. 51, 1880.—
Hulst, N. Amer. Phycitidae, p. 145, 1890.
Salebria basilaris (Zeller) Ragonot, Monograph, pt. 1. p. 353,
1893.—Forbes, Cornell Mem. 68, p. 627, 1923.—McDun-
nough, Check list, No. 6209, 1939.
Forewing pale ash gray to dark gray with a faint
bluish tint; basal area pale wood brown or pale orange,
a black shade along its inner margin expanding upward
at antemedial line to middle of cell; the lower fold in
basal area streaked with red or reddish brown with
some scattering of similarly colored scales on the pale
area above; antemedial line distinct to top of cell, well
contrasted, grayish white, nearly vertical, inwardly
dentate between cell and inner margin; margined out-
wardly by a narrow black line; subterminal line faint
but distinguishable, sinuate-dentate, followed in outer
area (on some specimens) by a broad reddish shade;
discal dots obscured; a row of small blackish dots along
terminal margin. Hind wings white with a faint ocher-
300329—56——10
ous or smoky tint; the veins very faintly darkened; a
thin brownish line along termen. Alar expanse, 24—27
mm.
Male genitalia resembling those of subcaesiella; the
clasper and lateral elements of transtilla somewhat
stronger; the smaller cornutus on penis longer and
stouter but also on some specimens reduced to a mere
vestige. Female genitalia with two granulate patches
on ventral surface of bursa, a weaker granulation of the
surface connecting them.
TypE Locauity: Massachusetts (type in BM),
Foop prant: Unknown.
Distrisution: Unirep Sratres: Maine, Augusta
(June), Orono; New Hampshire, Hampton (June); Mas-
sachusetts, Martha’s Vineyard (July), Wilmington
(June); New York, Catskill Mts., Ilion (June); New
Jersey, Newark; Illinois, Lacon (June); Indiana, Hess-
ville (May, July); Michigan, one example, state locality
only; Colorado, Fort Collins (July), and one example
state locality only; Utah, Provo Canyon (July). Can-
ava: Ontario, Budbury, Hymers (July), Trenton (July);
Manitoba, Aweme (June), McCreary, Winnipeg.
A strikingly marked species, the most easily identified
in the genus. The maxillary palpus of the male is, as
noted by Ragonot (Monograph, p. 354), not strictly in
the form of an aigrette. The scales are moderately
long and slender, but not hairlike, intermediate between
those of a squamous and a typical aigrettelike palpus.
The labial palpus is grooved on inner surface of the
second segment as in most species having maxillary
palpi of the aigrette type.
263. Nephopteryx termitalis (Hulst), new combination
Fiaures 342, 828
Pempelia termitalis Hulst, Trans. Amer. Ent. Soc., vol 13, p. 162,
Salebria termitalis (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115,
1889,—Hulst, Phycitidae of N. Amer., p. 153, 1890.
Salebria levigatella Hulst, Canadian Ent., vol. 24, p. 61, 1892.—
Forbes, Cornell Mem. 68, p. 626, 1923.—McDunnough,
Check list, No. 6207, 1939. (New synonymy.)
Myrlaea termitalis (Hulst) Ragonot, Monograph, pt. 1, p. 401,
1893.—McDunnough, Check list, No. 6223, 1939.
Forewing dark suffused gray (darker than on basilaris) ;
basal area dull reddish orange with little black on most
specimens except along costal edge; on others some
black scaling at extreme base and, narrowly, along
inner margin; antemedial line nearly obliterated by its
black borders which are more or less fused and from a
broad, nearly vertical band from inner margin to costa,
not strongly contrasted against the dark ground color
of the wing; subterminal line, discal and terminal dots
obscure.
Thorax dark gray, on some specimens more or less
suffused by the orange color of the basal area of fore-
wing. One specimen before me (a male from Inyo
County, Calif.) has the entire thorax and base of fore-
wing to the black inner border of the antemedial line a
dull madder-red, and the outer third of wing faintly
suffused with the same reddish shade. Hind wing dull
130 UNITED STATES NATIONAL MUSEUM BULLETIN 207
ocherous white to smoky white.
mm
Alar expanse, 23-27 mm.
Male genitalia with apical process of gnathos tri-
angulate. Transtilla weak but distinguishable. Clasper
erect and slightly curved, slender, digitate. One
moderately sized and one much smaller cornutus on
penis. Female genitalia with a single moderately large
granulate patch on posterior lateral corner of bursa and
extending in weaker granulation transversely across the
middle of the lower surface.
Typ Locatitizs: Colorado (fermitalis, in AMNH,
ex Rutgers); “Amherst, Massachusetts” [sie] (leviga-
tella, in AMNH, ex Rutgers).
Foop prant: Unknown.
DistRipuTion: UNITED States: Colorado, Glenwood
Springs, Gunnison County near Altmont, and two
examples (c', 2) with only the state locality; Utah,
Spanish Fork (July); Arizona, Prescott (June); Cah-
fornia, Inyo County (June), Placer County (June).
Canava: Ontario, Trenton (July), Manitoba, Winni-
peg; Alberta, Bilby (June); British Columbia, Clinton
(June).
Hulst also gives Massachusetts and Wisconsin as
localities for his levigatella. His type of the latter how-
ever has no locality label. A female of levigatella from
the Fernald Collection is in the National Collection.
It also bears a Hulst “type” label, but no locality or
date. The type of termitalis is a male, not a female as
given in Hulst’s original description. Genitalia of
conspecific females from western localities agree in every
detail with those of the type of levigatella so there can
be no question of the synonymy of the two Hulst names.
The labial palpi appear a trifle oblique (less tightly
appressed to the face than on most of the preceding
species) which may account for Ragonot’s reference of
the species to Myrlaea.
Alar expanse, 23-27
264. Nephopteryx termitalis yuconella (Dyar), new status
Salebria yuconella Dyar, Ins. Insc. Menstr., vol. 13, p. 12, 1925.—
McDunnough, Check list, No. 6194, 1939.
A slightly darker more suffused local race of termitalis;
the basal area of forewing almost entirely suffused with
blackish except for a narrow grayish white shade along
inner margin of the black inner border of the antemedial
line; no appreciable red or orange shading anywhere on
forewing or thorax. Alar expanse, 25-27 mm.
Genitatta: Male and female as in typical termitalis.
Typz tocatity: Near Fort Yukon, Alaska (type in
USNM).
Foop rirant: Unknown.
Distrisution: ALasKa: Dawson (June), Fort Yukon.
Specimens of yuconella as of typical termitalis show
a strong tendency to become greasy with age, indicating
that their larvae are borers; but nothing is known about
the biology of either form.
265. Nephopteryx bifasciella Hulst
Figures 343, 831
Nephopteryx bifasciella Hulst, Ent. Amer., vol. 3, p. 132, 1887.
Salebria bifasciella (Hulst) Ragonot, Ent. Amer., vol. 5, p. 115,
1889; Monograph, pt. 1, p. 366, 1893.—Hulst, Phycitidae
of N. Amer., p. 154, 1890.—Barnes and McDunnough,
Contributions, vol. 3, p. 197, 1916.—McDunnough, Check
list, No. 6214, 1939.
Salebria nogalesella Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 35, 1905.—MecDunnough, Check list, No. 6215, 1939.
(New synonymy.)
Forewing ash gray with a fine powdering of black
scales, giving the wing a faint pale bluish tint; the black
borders of the transverse lines strongly contrasted;
antemedial line distinct, at least from below upper vein
of cell, narrow, slightly oblique and somewhat angulate
at middle, its outer black border complete, slightly
broadened at costa, its inner black border narrow,
extending from inner margin only to top of cell; sub-
terminal line sinuate, bordered inwardly by a narrow
black line, and outwardly by a much fainter, paler
dark line; discal dots usually distinguishable but faint,
separate or confluent (sometimes both ways on the
same specimen); dots along terminal margin very
faint, more or less confluent. Hind wing white with a
very faint ocherous or smoky tint, slightly darker on
female than on male. Alar expanse, 20-22 mm.
Male genitalia with apical process of gnathos tri-
angulate. Clasper sharply curved and running close
and parallel to surface of harpe, simple and strongly
sclerotized. Cornuti rather short, but individually
variable in size. There is also some slight variability
in the size and shape of the apical process of gnathos.
The genitalia of the type of bifasciella and those of its
synonym nogalesella are more nearly alike than those
of any other two males before me. Female genitalia
with bursa nearly round, armed with two large granulate
patches, arranged as in figure 831 but with the position
of the anterior patch (at closed end of bursa) somewhat
variable. In one specimen from Palmerlee there is a
third patch on the right side of bursa and the bursa
itself is narrower and considerably elongated. ‘These
differences probably represent nothing more than
individual aberrations. Males from the same locality
are normal.
Type Locauities: Arizona (bifasciella, in AMNH,
ex Rutgers); Nogales, Ariz. (nogalesella, in USNM).
Foop prant: Unknown.
DistRiBuTIon: Arizona, Baboquivari Mts. (July),
Huachuca Mts. (July, Aug.), Nogales (July), Palmerlee,
Redington, ‘‘Southern Arizona” (Aug.), and two ex-
amples with only the state locality.
266. Nephopteryx uvinella (Ragonot), new combination
Fieures 344, 824
Meroptera uvinella Ragonot, N. Amer. Phycitidae, p. 8, 1887;
Monograph, pt. 1, p. 315, 1893.—Hulst, Phycitidae of N.
Amer., p. 148, 1890.—McDunnough, Check list, No. 6183,
1939.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 131
Salebria afflictella Hulst, Canadian Ent., vol. 32, p. 170, 1900.—
Forbes, Cornell Mem. 68, p. 625, 1923.—McDunnough,
Check list, No. 6200, 1939. (New synonymy.)
Meroptera liquidambarella Dyar, Proc. Ent. Soc. Washington,
vol. 6, p. 108, 1904.
Meroptera afflictella (Hulst) Dyar, Proc. Ent. Soc. Washington,
vol. 7, p. 34, 1905.
Forewing fuscous (gray-brown), the median area
heavily dusted with white giving it a pale ash gray
color; basal area of the ground color, rarely with some
red scaling in the median fold; dark base followed by
an oblique, straight, whitish band; this latter bordered
outwardly by a broad, somewhat diffused, dark brown
or blackish band through which may be distinguished
faint white traces of the true antemedian line; subter-
minal line with a very slight central bulge, rarely some-
what crenulate, whitish gray, bordered inwardly by a
diffused brown shade and outwardly by a narrow brown
line; discal dots more or less fused; blackish terminal
dots confluent. Hind wings pale to dark smoky fus-
cous. Alar expanse, 14-18 mm.
Male genitalia distinguished chiefly by the armature
of the penis. On the paratype of afflictella from Mont-
clair, N. J., there appears to be two cornuti, one closely
appressed to the other; but in other preparations before
me (including the types of wvinella and liquidambarella)
the two cornuti are completely fused into a single rather
short and stout, longitudinally ribbed cornutus. The
clasper is a slender, curved, sharply pointed, smooth,
sclerotized hook. Vestiges of a divided transtilla
distinguishable on most preparations.
Female genitalia small (approximately the size and
form of those of carneella); bursa with two moderately
sized granulate patches, the bursal lobe giving off the
ductus seminalis also granulate and partially (smoothly)
sclerotized; this lobe as usual arises from the dorsum of
bursa but is deflected to the left.
TYPE LOCALITIES: United States (wvinella, in USNM)
Elizabeth, N. J. (affictella, in AMNH, ex Rutgers);
Washington, D. C., (liquidambarella, in USNM).
Foop piant: Liquidambar styraciflua (sweetgum).
Larva a leaf-tier.
DistrisutTion: Connecticut, East River (May), Stam-
ford (Aug.); New Jersey, Elizabeth (Aug.), Lakeland
(May), Montclair (June, Aug.), New Lisbon (June);
District of Columbia, Washington (May, Aug.) ; Virginia,
Skyland (July); North Carolina, Greensboro (Aug.);
Georgia, Savannah (Sept.); Florida, Lakeland (May,
June).
Ragonot’s wvinella has been an unknown entity ever
since its description and was suspected of being only a
variety of Meroptera pravella. Fortunately the type
was secured by Dr. Barnes when the Oberthiir Collec-
tion was sold. It lacks one forewing but is otherwise
intact and its habitus and genitalia leave no doubt that
it is the same as the sweet-gum feeder described by
Hulst and Dyar. The type of affictella is a female.
The types of uvinella and liquidambarella are both males.
The genitalia of all of them are before me.
267. Nephopteryx celtidella (Hulst), new combination
Figures 345, 830
Salebria celtidella Hulst, Phycitidae of N. Amer., p. 155, 1890.—
Beutenmiller, Canadian Ent., vol. 22, p. 17, 1890 (larva).—
Ragonot, Monograph, pt. 1, p. 354, 1893.—Forbes, Cornell
Mem. 68, p. 626, 1923.—McDunnough, Check list, No.
6206, 1939.
Ground color ocherous (clay color) more or less shaded
with fuscous gray over submedian and (on especially
dark females) outer basal areas; the ground color espe-
cially well contrasted at base and in the central area
about the discal spots, also on thorax, at extreme base
of wing and on thorax sometimes of a tawny shade; ante-
medial line far out towards middle of wing, oblique, sinu-
ate, faint, indicated chiefly by its narrow black borders
which are more or less broken into dots on the veins;
subterminal line sinuate-serrate, bordered inwardly by
a row of black (somewhat confluent) dots and outwardly
by a row of small black wedges on the veins; discal dots
distinct, well separated, black; a row of distinct black
dots along termen. Hind wing pale to rather dark
smoky fuscous. Alar expanse, 18-22 mm.
Male genitalia with apical process of gnathos smaller
than that of preceding species (not triangulate). Clasper
rather short, bent across surface of harpe, blunt. Sclero-
tized lateral elements of transtilla distinguishable,
rather long, slender.
Female genitalia with a round, moderately large
granulate patch on ventral surface of bursa and a smaller
patch near junction of bursa and ductus bursae and the
base of the lobe giving off the ductus seminalis.
Typr Locatity: New York (in AMNH, ex Rutgers).
Foop puant: Celtis. Larva a leaf-tier.
Distripution: Unitep Starrs: New York, Long
Island; Maryland, Plummers Isl. (May, July); Flor-
ida, Palm Beach (Feb.); Texas, Brownsville (July), Vic-
toria (May), Zavalla County (Apr.); Mississippi, “Agr.
College” (Apr.), Starkville (July); Missouri, St. Louis
(Aug.); Zilinois, Oconee (Aug.).
A distinct species distinguished from any of the spe-
cies with aigrettelike maxillar palpi on the male by its
clay-colored, black-mottled forewings. Its habitus is
nearest to that of rubrisparsella in the group with
squamous, male maxillary palpi.
Genus Nephopteryx, Species 268-271: N. rubri-
sparsella to N. bisra
[Males with squamousmaxillary palpi.]
268. Nephopteryx rubrisparsella (Ragonot)
Ficures 346, 832
Pristrophora rubrisparsella Ragonot, N. Amer. Phycitidae, p. 6,
87
Pristophora rufibasella Ragonot, N. Amer. Phycitidae, p. 7, 1887.
Sciota croceella Hulst, Ent. Amer., vol. 4, p. 115, 1888.
Nephopteryx rubrisparsella (Ragonot), Ent. Amer., vol. 5, p. 115,
1889; Monograph, pt. 1, p. 284, 1893.—Hulst, Phycitidae
of N. Amer., p. 145, 1890.—Forbes, Cornell Mem. 68, p. 623,
1923.—McDunnough, Check list, No. 6169, 1939.
132 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Psorosa texanella Hulst, Canadian Ent., vol. 32, p. 174, 1900.
Hulstea texanella (Hulst), U.S. Nat. Mus. Bull. 52, p. 432, 1903.—
MeDunnough, Check list, No. 6342, 1939. (Newsynonymy.)
Similar to celtidella in color and maculation except:
Ocherous basal area of forewing more or less shaded with
reddish purple and similar suffusions over much of the
median and outer areas; also a broad blackish suffusion
over the area of the antemedial line and extending some-
what beyond it; the inner dark border of subterminal
line more nearly continuous and the outer bordering
dashes much fainter except in dark and very well
marked examples. Alar expanse, 17-20 mm.
Male genitalia figured from type of texanella Hulst,
which agree in every detail with those of typical rubri-
sparsella and of the male type of croceella. ‘They differ
from those of celtidella in having a longer, more strongly
sclerotized clasper, and different cornuti (shown in lat-
eral view in our figures). The cornuti lie side by side
in both celtidella and rubrisparsella, but in the former
species they are of equal length while in rubrisparsella,
one cornutus is much shorter than the other—not too
reliable a character, but apparently consistent here.
Transtilla, on dissection, distinguishable as a complete
band, but central area very weak and in balsam prepa-
ration only the well sclerotized lateral elements easily
seen.
Female genitalia with a broad granulate band par-
tially encircling middle of bursa, similar to that in sub-
fuscella except that in subfuscella it is chiefly over the
ventral surface of bursa while in rubrisparsella it is on
the dorsal.
TYPE LOCALITIES: United States (rubrisparsella, in
Paris Mus.); Texas (rufibasella, in Paris Mus.); Blanco
County, Tex. (croceella and texanella, in AMNH, ex
Rutgers).
Foop pnant: Celtis.
Distrisution: Texas, Black Jack Springs, Blanco
County, Kerrville (June); Oklahoma; Missouri, St.
Louis (June); Jilinois, Lacon (July), Putnam County
(uly); West Virginia, Jefferson County (Aug.); Mary-
land, Plummers Isl.; District of Columbia, Washington
(May, June).
Very close to celfidella and distinguished from it
chiefly by its genitalia and squamous male maxillary
palpi. Hulst’s texanella was described from an ab-
normal specimen with vein 4 lacking in the hind wing.
I have before me two similar abnormal females (from
Oklahoma and Illinois) and in the following species
(gilvibasella) a couple of examples, out of a long series
of normal specimens, that also have vein 4 absent.
The type of croceella in addition to the Hulst name
label also bears the following label in Ragonot’s hand-
writing: ‘‘Nephopteryx rubrisparsella Rag.—rufibasella
Rag.”
269. Nephopteryx gilvibasella Hulst
Figure 836
Nephopteryx gilvibasella Hulst, Phycitidae of N. Amer., p. 145,
1890.—Ragonot, Monograph, pt. 1, p. 289, 1893.—McDun-
nough, Check list, No. 6172, 1939.
Salebria lacteella Hulst, Canadian Ent., vol. 32, p. 71, 1900.—
cia and McDunnough, Contributions, vol. 3, p. 197,
1916.
Much paler than the preceding species (rubrisparsella)
and without its blackish shadings. Ground color pale
gray with a very faint bluish tint; extreme base of wing
ocherous orange, this shade extended somewhat on
inner margin and indicated on many specimens in
median area over the lower fold; a broad band of the
same color immediately preceding the antemedial line
and extending from inner margin to middle of cell;
above it a blackish shade extending to costa; lower half
of antemedial line clearly indicated, narrow, whitish,
bordered outwardly by a narrow black line which fuses
towards costa into the blackish shade above the ocher-
ous patch; subterminal line faintly bordered inwardly
by a faint blackish line; discal dots small, separated,
sometimes obscure, but usually distinct; a row of weak
but discernible blackish dots along termen. Hind wing
subpellucid; whitish with a very faint ocherous tint; a
weak, pale brown line along termen; the veins not
appreciably darkened. Alar expanse, 17-20 mm.
Male genitalia similar to those of rubrisparsella. The
eighth-segment tufts much simpler, reduced to two
lateral pairs with scales of a uniform shape and size.
Female genitalia with a broad granulate band on
bursa similar to that in rubrisparsella; bursa bulged into
a small, strongly granulate lobe at its left posterior
angle (this lobe opposite to that giving off ductus
seminalis) ; also a narrow row of moderately long spines
across upper side of bursa at junction of bursa and
ductus bursae.
TyprE Locatitizs: Not given (gilvibasella, in AMNH,
ex Rutgers); Blanco County, Tex. (lacteella, in AMNH,
ex Rutgers).
Foop prant: Unknown.
Distrisution: Texas, Blanco County (Sept.),
Brownsville (Mar., July), San Benito (Mar., Apr.,
May, July, Aug., Sept.).
The species is close to but distinct from rubrisparsella.
The types of gilvibasella and lacteella are both females
with identical genitalia. A long series from Browns-
ville and San Benito are before me.
270. Nephopteryx crassifasciella Ragonot
Fieurts 347, 835
Nephopteryx crassifasciella Ragonot, N. Amer. Phycitidae, p. 8,
1887; Monograph, pt. 1, p. 285, 1893—Hulst, Phycitidae
of N. Amer., p. 146, 1890.—MceDunnough, Check list, No.
6170, 1939.
Nephopteryx decipientella Dyar, Proc. Ent. Soc. Washington, vol.
7, p. 34, 1905.—McDunnough, Check list, No. 6162, 1939.
(New synonymy.)
Nephopteryx crataegella Barnes and McDunnough, Contributions,
vol. 3, p. 222, 1917—McDunnough, Check list, No. 6164,
1939. (New synonymy.)
Forewing ash gray more or less suffused by somber
purplish or gray brown, shading into blackish brown on
the borders of the transverse lines; the pale color limited
to a rather narrow band preceding the inner border of
the antemedial line, part of the median area following
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 133
the antemedial line and surrounding the discal dots,
and some faint pale dusting immediately preceding
terminal margin; extreme base of wing purplish brown
with a shading (in some specimens) of reddish scales,
especially towards inner margin; antemedial line pale
gray, distinguishable only from inner margin to middle
of cell, bordered inwardly by a broad, vertical, blackish
brown band which extends to costa, and outwardly by
a narrow black line which fuses into the inner dark
border towards costa; subterminal line sinuate, pale
gray with narrow, black-brown borders; discal dots
distinct, sometimes partially fused. Hind wing light
to dark smoky brown; the veins slightly darkened and
a dark line along termen. Alar expanse, 16-17 mm.
Male genitalia with clasper short, blunt, curved.
Cornuti moderately long, approximately the same size
and lying side by side. A long slender arm projecting
from each dorsolateral angle of vinculum. Female
genitalia, with two large, opposing, roundly oval, gran-
ulate patches on bursa, and some rather strong, darkly
pigmented spining on the lobe giving off the ductus
seminalis.
Type LocauitiEs: California [?] (crassifasciella, lost?) ;
unknown (decipientella, in USNM); Lakeland, Fla.
(crataegella, in USNM).
Foop piants: Vaccinium, Crataegus.
Distrisution: Maryland, Plummers Isl. (May);
District of Columbia, Washington (May) ; Georgia, Sape-
loe (Sept.); Florida, Lakeland (May).
A male from the District of Columbia (reared from
Vaccinium by Chambliss, May 10, 1895) and a similar
female from Plummers Isl., Md., in the National Col-
lection agree in every detail with Ragonot’s figure and
description of crassifasciella and were so identified by
Dyar. His decipientella was described from a single
specimen without locality label. It is merely a dark-
suffused male with the pale coloration limited to a more
restricted area about the discal spots and some light
dusting between the outer veins. The Georgia example
(reared from Vaccinium) is a female and matches the
type of decipientella except for venation. It is another
of the all too frequent phycitid freaks with vein 4 miss-
ing from hind wing and 7-8 long stalked. The Barnes
and MeDunnough type of crataegella (also a male and
reared from Crataegus) differs from typical crassifasciella
only in a somewhat more extended and lighter colora-
tion of the pale areas of forewing. All three males
before me agree in genitalic structure.
The species is easily identified by the peculiar develop-
ment of the vinculum, not found in any other American
species in the genus.
Dr. Bourgogne informs me that the type of inquilinella
could not be found at Paris. It is probably lost, and I
suspect that the California citation in Ragonot’s Mono-
graph was probably a guess. I have seen nothing from
the Pacific Coast States that could possibly be his
species.
271. Nephopteryx bisra Dyar
Fievurse 833
Nephopteryx bisra Dyar, Ins. Inse. Menstr., vol 7, p. 51, 1919.
Forewing pale gray-brown; base shaded with black-
ish; a straight, narrow, oblique whitish line along inner
margin of the inner border of the antemedial line, the
latter a thin, oblique, notched, whitish line, obscured
towards costa; its inner border a wide blackish band
reaching to costa; its outer border a narrow black line;
a pale patch on inner margin following the antemedial
line; subterminal line distinct, whitish gray, sinuate,
margined by narrow blackish brown inner and outer
lines; discal spots fused into a lunate mark with a pale
surrounding shade; a blackish line along termen. Hind
wing pale gray-brown, paler than ground color of fore-
wing; veins not appreciably darkened; a narrow, dark
(brownish) line along termen. Alar expanse, 21 mm.
Female genitalia with a large granulate patch cover-
ing most of dorsal surface of bursa, a smaller latero-
ventral patch and some scattered granulations at left,
posterior angle on ventral side; bursa otherwise rather
densely and finely spinose.
Typs Locauity: Orizaba, México (type in USNM).
Foop prant: Unknown.
Known only from the female type. The placement
in the second group of Nephopteryz is provisional, pend-
ing discovery of a male. In the type, vein 10 of fore-
wing is shortly stalked with the stem of 8-9.
69. Genus Tlascala Hulst
Tlascala Hulst, Phycitidae of N. Amer., p. 146, 1890.—Forbes,
Cornell Mem. 68, p. 623, 1923. (Type of genus: Nepho-
pleryx reductella Walker.)
Tongue well developed. Antenna weakly pubescent;
on male, with sinus and enlarged scale tuft in base of
shaft. Labial palpus upturned, not appressed to face,
cylindrical, smooth scaled, reaching above vertex;
second segment of male not hollowed; third segment
moderately long, acuminate. Maxillary palpi of both
sexes squamous, appressed to face. Forewing with
most of inner border of antemedial line consisting of
raised scales; 11 veins; vein 2 from near lower outer
angle of cell; 3 from the angle, about equidistant from
2 and 4; 4 and 5 very shortly stalked or connate, rarely
(in individual specimens) closely approximate at base;
6 from below upper angle of cell, straight; 8-9 long
stalked (for three-fourths of their lengths) ; 10 from the
cell approximate to or connate with the stalk of 8-9 at
base, and approximate to it for some distance beyond
base; male without costal fold. Hind wing venation
similar to that of Meroptera and Nephopteryx. Highth
abdominal segment of male with compound scale tufts.
Male genitalia with strongly sclerotized, broadly tri-
angulate clasper with a row of irregular teeth along its
outer margin. Penis armed with two, equally sized,
stout, sharply curved, thornlike cornuti. Otherwise
as in Nephopteryz.
134 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Female genitalia without signum; bursa finely and
densely spined over most of inner surface and with a
cluster of longer and stronger spines near middle of
lateral margin; ductus bursae short (less than half the
length of bursa); armed on ventral surface by an elon-
gate pair of granulate plates; at genital opening a
strongly sclerotized, centrally interrupted genital plate,
attached to a narrow, sclerotized collar (incomplete
dorsally and ventrally) and supplemental to the regular
eighth-segment collar; ductus seminalis from bursa near
junction of bursa and ductus bursae.
The genus is close to and has several characters in
common with both Meroptera and Nephopteryz, resem-
bling the former in the well developed genital plate of
the female genitalia and the strong, serrate clasper of
the male, differing from both genera in the rough scaling
on forewing, the ungrooved labial palpi and much
stouter antennal tuft of the male, the sharply curved
cornuti on penis, and the supplemental collar attached
to the female genital plate. As here defined it contains
only its American type species.
272. Tlascala reductella (Walker)
FIGURES 28, 348, 834
Nephopteryx reductella Walker, List, vol. 27, p. 638, 1863.—
Ragonot, Ent. Amer., vol. 5, p. 115, 1889; Monograph, pt. 1,
p. 283, 1893.
Pempelia gleditschiella Fernald, in Comstock, in Rep. [U. §.]
Comm. Dep. Agr. for 1880, p. 262, 1881.—Packard, U. S.
Dep. Agr. Fifth Rep. Ent. Com., p. 652, 1890.
Tlascala reductella (Walker) Hulst, Phycitidae of N. Amer., p.
146, 1890.—Forbes, Cornell Mem. 68, p. 624, 1923.—
McDunnough, Check list, No. 6180, 1939.
In color and maculation similar to the Kuropean
Nephopteryz rhenella except for the raised scaling of the
subbasal black bar.
Forewing dull ash gray with purplish brown shading
at extreme base and paler brownish shading on outer
median and terminal areas; antemedial line complete
but faint, narrow, oblique, slightly indented on lower
half, preceded by a broad black band of rough scales
and followed by a narrow black band; subterminal line
obscure, sinuate-dentate, faintly bordered by dark inner
and outer lines; discal dots distinct, blackish, separated;
a row of faint blackish dots along termen. Hind wing
pale smoky fuscous; the veins very slightly darkened
and a narrow dark line along termen. Alar expanse,
17-23 mm.
Male genitalia with cornuti set side by side near outer
end of penis, their apices turned away from each other.
Female genitalia as given for the genus.
TypE LOCALITIES: Honduras [sic] (reductella, in BM);
District of Columbia (gleditschiella, in USNM).
Foop Piant: Gleditsia. Larva a leaf-tier.
DistrisutTion: District of Columbia (May, July);
Maryland, Plummers Isl. (June); North Carolina, Hil-
tonhead Isl. (Aug.); Pennsylvania; Illinois, Decatur
(Mar., Apr., May, July), Quincy (May); Jowa, Ames;
Kansas, Lawrence (May); Missouri, St. Louis (May);
Texas, Kenedy (Apr.), Paris (May), Victoria (July),
Zavalla County (Apr.); Louisiana, Crown Point (June,
larva), New Orleans (larva, June).
Walker gives Honduras as the type locality. I
rather doubt the correctness of this citation, for I have
never seen anything from Central America that even
remotely resembled the species. I have not seen his
types, but have no reason to question the correctness of
Ragonot’s reference of gleditschiella to synonymy.
70. Tulsa, new genus
Type of genus: Nephopteryx finitella Walker.
Characters of Tlascala except:
Forewing with some rough scaling in the median
area beyond the outer margin of the antemedian line,
sometimes a small tuft on lower fold, always a few
roughened scales in the discal spots; veins 4—5 approxi-
mate at base and for a short distance beyond. Male
genitalia with sacculus of harpe considerably enlarged,
strongly sclerotized, densely and finely spined along
entire lower margin, and produced at apex; clasper a
thin, dished plate produced into an elongate, curved,
sharply pointed claw at each lower angle. Uncus
broadened at apex. Transtilla complete, very weakly
but evenly sclerotized throughout. Cornuti straight,
set one before the other. Female genitalia with several
lines of fine spines running from bursa into ductus
bursae almost to genital opening; genital plate and its
attached supplemental collar strongly wrinkled (more
so than shown in the figures) ; no granulations in ductus
bursae.
The genus is very close to Tlascala and I propose the
new name with some misgiving; but the habitus of the
moths and their genitalia, male and female, differ so
much from those of the type of Tiascala that something
more than a species-group difference is indicated. When
larvae and host relations of the Tulsa species are known
we shall probably find additional supporting characters
for the genus. Specifically the genitalia are remarkably
similar, offering little or nothing to distinguish the
species. Four are here recognized.
273. Tulsa finitella (Walker), new combination
Figure 349
Nephopteryx finitella Walker, List, vol. 27, p. 53, 1863.—Ragonot,
Monograph, pt. 1, p. 282, 1893.
Tlascala finitella (Walker) Hulst, Phycitidae of N. Amer., p. 147,
1830.—Forbes, Cornell Mem. 68, p. 624, 1923.—McDun-
nough, Check list, No. 6177, 1939.
Elasmopalpus melanellus Hulst, Phycitidae of N. Amer., p. 157,
1890.—Barnes and McDunnough, Contributions, vol. 3,
p. 199, 1916 (makes synonym of finitella).
Forewing very dark gray with the blackish borders of
the antemedial and postmedial lines but slightly con-
trasted; some slight dusting of white on subbasal, me-
dial, and terminal areas; antemedial line somewhat
stronger, its inner blackish bordering line more or less
interrupted on the veins, its outer border continuous
but faint; discal dots tending to coalesce; an obscure
row of blackish dots along termen; raised scales con-
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 135
spicuous on lower half of inner border of antemedial
line and as a patch on middle of lower fold; a raised
scale or two in the discal spots. Hind wing pale smoky
fuscous with a darker shading towards termen; the
veins slightly darkened; the entire wing darker on
northern examples. Alar expanse, 21-25 mm.
Male genitalia with no distinguishing specific features.
Female genitalia similar to those of wmbripennis but
smaller (about the size of infinitella) and with more
decided wrinkling of the genital plate and its supple-
mental collar.
Typr LocatitiEs: United States (jinitella, in BM);
Florida (melanellus, in AMNH, ex Rutgers).
Foop prant.—Blueberry. Thisrecord isfromareared
female (Brunswick, Ga., Quaintance No. 31501) in the
National Collection. The larva, however, may have
been accidentally on that plant.
DistrisutTion: Unirep States: Florida, Charlotte
Harbor (Mar.), Fort Myers (Apr.), Miami (Mar.), St.
Petersburg (Apr.), Tampa, also examples with only
state locality; Georgia, Brunswick (June); North Caro-
lina, Raleigh (June); Virginia, Richmond (May); Dis-
trict of Columbia, Washington (May, June); New Jersey,
Essex County (May), Newark (May); Massachusetts,
Martha’s Vineyard (Aug.); Jndiana, Hessville (June).
Canava: Ontario, Trenton; Quebec, Kazubazua (June).
Walker also reports the species from Nova Scotia.
The type of melanellus in the Rutgers Collection is a
female without abdomen. Matching cotypes (o and 9)
from Mrs. Slosson’s material are in the National Collec-
tion. There can be no question of the synonymical
reference by Barnes and McDunnough.
274. Tulsa umbripennis (Hulst), new combination
Fieures 350, 842
Pinipestis umbripennis Hulst, Canadian Ent., vol. 27, p. 57, 1895.
Ortholepis gillettella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 107, 1904.
Tlascala umbripennis (Hulst) Barnes and McDunnough, Contri-
butions, vol. 3, p. 195, 1916.—McDunnough, Check list,
No. 6178, 1939.
Fore and hind wings dark brown with a somewhat
glossy sheen not possessed by the other species of the
genus; discal dots on forewing confluent, forming a bar
on discocellular vein; raised scales and maculation as in
Jjinitella. Alar expanse, 25-26 mm.
Male genitalia, figured from type of gillettella, agree
in every detail with those of the male type of umbri-
pennis. Female genitalia with bursa copulatrix
slightly larger than that of any of the other Tulsa
species except oregonella.
TypEtocauitizs: Colorado (umbripennis, in AMNH,
ex Rutgers; gillettella, in USNM).
Foop prant: Unknown.
Known only from the type locality. A series before
me is from Chimney Gulch, Colo. (June and July).
The types of umbripennis and gillettella have only the
state locality, but the latter was probably from the
neighborhood of Fort Collins.
275. Tulsa oregonella (Barnes and McDunnough), new combination
FicurE 351
Tlascala oregonella Barnes and McDunnough, Contributions,
ve p. 175, 1918.—McDunnough, Check list, No. 6179,
Forewing a dull, powdery, slate gray; basal area a
trifle paler; the transverse lines somewhat more distinct
and better defined than on wmbripennis; inner black
border of antemedial line slightly narrower than in
preceding species; dark borders of subterminal line very
faint; discal dots separated. Hind wing smoky gray-
brown, paler than that of wmbripennis and not glossy.
Alar expanse, 26-28 mm.
Female genitalia similar to those of wmbripennis.
Type Locauity: Crater Lake, Oreg. (July; type in
USNM).
Foop puant: Unknown.
Known only from the type series.
276. Tulsa infinitella (Dyar), new combination
Figure 841
Tlascala infinitella Dyar, Ins. Insc. Menstr., vol. 7, p. 52, 1919.
Similar to oregonella except: Forewing slightly darker;
antemedial line obscure; the discal dots confluent, form-
ing a blackish line of slightly roughened scales. Hind
wing as in southern specimens of finitella; pale smoky
fuscous at base with a darker shading towards termen.
Alar expanse, 27 mm.
Female genitalia like those of finitella except lateral
elements of genital plate smoother.
Tyrer Locauity: Orizaba, México (type in USNM).
Foop piant: Unknown.
Known only from the female type.
Genus 71: Homoeographa
[Venational division C. veins 4 and 5 shortly stalked or weakly
anastomosed for a short distance beyond cell. Hind wing with
discocellular vein of cell vertical, straight. Antenna of male
with sinus and scale tuft in base of shaft. Labial palpus obliquely
upturned, second segment laterally flattened and broadly scaled,
on male grooved to hold maxillary palpus. Maxillary palpus of
male in the form of an aigrette. Highth abdominal segment of
male with compound scale tufts. Male genitalia without trans-
tilla; clasper present, digitate; penis armed with two moderately
stout cornuti. Female genitalia without signum; bursa with
deep, convoluted, sclerotized folds; genital opening simple.]
71. Genus Homoeographa Ragonot
Homoeographa Ragonot, Nouv. Gen., p. 24, 1888; Monograph,
pt. 1, pp. xlvi, 432, 1893. (Type of genus: Homoeographa
lanceolella Ragonot.)
Tongue well developed. Antenna pubescent; male
with sinus and strong tuft in base of shaft. Labial
palpus obliquely upturned, reaching well above vertex;
second segment flattened and broadly scaled, on male
grooved to hold the maxillary palpus; third segment
short, partially hidden in scaling of second. Maxillary
palpus of male in the form of an aigrette; of female
136
squamous. Forewing smooth; 11 veins; vem 2 from
before the lower outer angle of cell; 3 from the angle,
separated from 4-5; 4 and 5 shortly stalked or weakly
anastomosed for less than half their lengths from cell;
6 from below upper angle of cell, slightly bent at base;
8 and 9 stalked for over half their lengths; 10 from the
cell, connate or closely anastomosed at base with the
stalk of 8-9. Hind wing with vein 2 from well before
lower outer angle of cell; 3 from the angle, long; 4 and 5
stalked for about four-fifths of their lengths; 7 and 8
anastomosed for three-fourths of their lengths; cell
short, one-third the length of wing; discocellular vein
vertical, straight. Highth abdominal segment of male
with compound ventral scale tufts.
Male genitalia with uncus subtriangulate and with
blunt, moderately broad, notched apical margin. Apical
process of gnathos a moderately stout hook with a
slender, digitate basal projection. Transtilla absent.
Harpe with costa sclerotized throughout, but not
produced at apex; cucullus simple, curved, apex
bluntly pointed; clasper present, simple, erect, digitate.
Penis armed with two moderately stout cornuti, about
one-third as long as aedeagus. Vinculum about twice
as long as broad, evenly tapering to its truncate,
strongly sclerotized, anterior margin. Vinculum U-
shaped with somewhat enlarged base.
Female genitalia with bursa elongate, narrow, signum
absent, two or three deep, convolute sclerotized folds
at posterior half, the sclerotization extending for a
short distance into ductus bursae; genital opening
simple. Ductus seminalis from bursa, near junction of
bursa and ductus bursae.
The genus is easily distinguished by its hind wing
venation and genitalia. The male genitalia indicate
a close relationship to the Nephopteryx group of genera,
However the characteristic hair brush on the harpe of
the latter are absent from Homoeographa. It contains
only one known tropical American species.
277. Homoeographa lanceolella Ragonot
Ficures 352, 839
Homoeographa lanceolella Ragonot, Nouv. Gen., p. 25, 1888;
Monograph, pt. 1, p. 483, 1893.
Forewing gray heavily dusted with white on costal
half; antemedial line indicated by a narrow, white, in-
wardly notched line between cell and inner margin and
above that by its incompleted outer border, a black
line oblique from costa to cell thence inwardly angled to
lower vein of cell; subterminal line faint, sinuate, in-
dicated chiefly by blackish gray bordering streaks from
costa, the inner one the longer and continued as a weak
blackish shading to inner margin; discal dots small,
separated, blackish gray; in outer area black streaklets
bordering vein 1b above and veins 3 and 6 below.
Hind wing semitranslucent smoky white; the veins
slightly darkened and a narrow dark line along termen.
Head ashy white. Alar expanse, 21 mm.
Female genitalic characters as given for the genus.
TypE LocaLity: Callao, Peri (type in Paris Mus.).
Foop piant: Unknown.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Known only from the type series in the Muséum
National d’Histoire Naturelle, Paris, and the British
Museum.
Genera 72-76: Telethusia to Pyla
[Venational division B. Veins 4 and 5 of forewing usually
separated at base (shortly stalked in Actriz); 10 from the cell,
separated at base from stalk of 8-9. Hind wing with cell less
than half the length of wing (about one-thid in Pyla). Antenna
of male with sinus and scale tuft in base of shaft. Labial palpus
oblique or upturned. Maxillary palpus various (minute,
squamous or aigrettelike). Male genitalia with transtilla
usually absent, if present (Phobus, Stylopalpia) incomplete or
its median area very weakly sclerotized; harpe with sclerotized
costa sometimes produced at base, never at apex; clasper absent
or more or less developed (strongly so in many species of Pyla);
aedeagus frequently divided (bifid) or spined; penis unarmed or
finely scobinate or finely and weakly spined, rarely (Phobus)
with a single cornutus. Female genitalia without signum; bursa
frequently smooth or weakly spined, occasionally with some
sclerotized folds continued from ductus bursae; the latter more
or less sclerotized in part, in many Pyla species broadly expanded
towards genital opening.]
72. Telethusia, new genus
Typr or Genus: Pempelia ovalis Packard.
Tongue well developed. Antenna pubescent; on male
with sinus and enlarged scale tuft in base of shaft.
Labial palpus obliquely upturned, reaching to vertex
on male, above vertex on female; laterally flattened and
broadly scaled; second segment of male grooved on
inner side to hold the tongue; third segment consider-
ably shorter than second, bluntly pointed, more or less
deflected forward and partially hidden in scaling of
second segment. Maxillary palpus minute (a mere
vestige). Forewing smooth; 11 veins; vem 2 from
before lower outer angle of cell; 3 from the angle,
nearer to 4 than to 2; 4 and 5 separated at base, parallel
for a short distance beyond; 6 from below upper angle
of cell, straight; 8 and 9 stalked for half or nearly half
their lengths; 10 from the cell separated from the stalk
of 8-9 at base and divergent from it shortly beyond;
male without costal fold. Hind wing with vein 2 from
before lower outer angle of cell; 3 from the angle,
connate at base with the stalk of 4-5; 4 and 5 stalked
for about half their lengths; 7 and 8 contiguous or
weakly anastomosed for a short distance beyond cell;
cell less than half the length of wing; discocellular vein
curved, outwardly produced at lower angle of cell.
Eighth abdominal segment of male with compound
scale tufts.
Male genitalia with uncus subtriangulate; apex
bluntly rounded. Apical process of gnathos a short,
stout hook. Transtilla absent. Harpe simple; clasper
rudimentary. Aedeagus simple, straight, not taper-
ing; penis unarmed except for a small comb of very
weak, short, slender spines. Vinculum stout, longer
than greatest width, tapering slightly to truncated
terminal margin.
Female genitalia with ovipositor strongly sclerotized;
apophyses (supporting rods) of ovipositor and eighth
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 137
segment collar stout; bursa elongate, finely scobinate
over two-thirds of its inner surface but without signum
or other sclerotization; ductus bursae unsclerotized
except for a narrow, weak band near genital opening;
ductus seminalis from junction of bursa and ductus
bursae. At least half of membrane between collar and
ovipositor finely and densely spinose.
The genus is erected for a species hitherto referred to
Nephopteryz. It differs from the latter in its vestigial
maxillary palpi, the lack of cornutus or cornuti of penis
or the hair brush on harpe of its male genitalia, the
hardened ovipositor of female, the shorter stalking of
veins 8 and 9, and the somewhat more separated condi-
tion at base of veins 4 and 5 of forewing.
278. Telethusia ovalis (Packard), new combination
Ficures 353, 843
Pempelia ovalis Packard, Ann. Lyc. Nat. Hist. New York, vol. 10,
p. 269, 1873.
Nephopteryz latifasciatella Packard, Ann. Lyc. Nat. Hist., New
York, vol. 10, p. 269, 1873.
Nephopteryz ovalis (Packard) Grote, Bull., U. 8. Geol. Geogr.
Surv. Terr., vol. 4, p. 696, 1878; North Amer. Ent., vol. 1,
p. 11, 1879.—Hulst, Phycitidae of N. Amer., p. 144, 1890.—
Ragonot, Monograph, pt. 1, p. 269, 1893.—Forbes, Cornell
Mem. 68, p. 623, 1923——McDunnough, Check list, No.
6163, 1939.
Nephopteryz ovalis geminipunctella Ragonot, Monograph, pt. 1,
p. 270, 1893.
Nephopteryx modestella Hulst, Canadian Ent., vol. 34, p. 170
1900.—Barnes and McDunnough, Contributions, vol. 3,
p. 196, 1916.
Forewing ashy gray-white with a distinct powdery
appearance, the white dusting conspicuous on median
and basal areas; antemedial line oblique, zigzag (twice
notched), narrow, white, bordered inwardly by a broad
dark fuscous (blackish gray) band which is interrupted
by a streak of dull ocherous orange at lower fold (this
ocherous shade also continued along fold in median
area) ; antemedial line bordered outwardly by a blackish
gray bar at costa and similarly colored dots on cell and
just above inner margin; subterminal line sinuate
bordered inwardly by a more or less broken blackish
gray line and outwardly by a broader blackish gray,
brownish, or brownish ocherous shade (the latter when
present interrupted by blackish streaklets on the veins),
both borders strongly accented at costa; on most speci-
mens a narrow, oblique, dark shading across median
area from costal inner margin of subterminal line; discal
dots distinct, separated, blackish, the lower sometimes
expanded into a black smudge; a row of black dots along
termen. Hind wing pale smoky fuscous, more whitish
in some specimens, darker and somewhat brownish gray
in others. Alar expanse, 20-28 mm.
Genitalia as given for the genus.
Type Locauities: Maine (ovalis and latifasciatella,
in MCZ); Washington State (geminipunctella, in Paris
Mus.); Massachusetts (modestella, in AMNH, ex Rut-
gers).
Foop puants: Antennaria, Eriophyllum ignotum.
These records from Washington specimens reared by
J. F. G. Clarke. Presumably on other Compositae.
Distrisution: Unitep Srates: Maine, Orono (July),
Wales (July); New Hampshire, Durham, Hampton
(Aug.), Fort Washington (July); Vermont, Clarenton;
Massachusetts; Connecticut, East River (July); New
York, Catskill Mts., Ilion (June, July); Colorado, Gun-
nison County (near Altmont, July); Utah, Stockton
(June, July); Montana, Missoula (Aug.); Washington,
Bellingham (June), Chuckanut Bay (Whatcom County,
June), Godman Springs (Blue Mts., July), Kamiack
Butte (May), Pullman (June, July); California, Placer
County (June), San Jacinto Mts. (July), Tuolumne
Meadows (July, Aug.). Canapa: Ontario, Trenton
(July); Alberta, Banff (June, July); British Columbia,
Wellington.
The species is variable in color, especially in the
Western areas of the United States and Canada. Most
of the specimens from Washington and British Colum-
bia have the white dusting on forewing more conspicu-
ous and the dark markings more strongly contrasted
than on eastern examples. However, there are inter-
grades, and no sharp line can be drawn on color between
the two areas. In the Tuolumne Meadows of Cali-
fornia there is a larger (26-28 mm.) and somewhat paler
form. Specimens from Colorado, Alberta, and occa-
sionally from Washington form another darker and
duller variety, the forewing showing little or no brown
shading, the pattern markings a dull black, and the
pale areas more gray than whitish and less strongly
contrasted against the dark markings than in other
Washington or eastern specimens. ‘Three specimens
before me from California have the transverse dark
lines much weaker and the over-all color an ashy gray
with a slight bluish tint. The Utah examples are the
most distinctive of all the forms, their hind wings de-
cidedly paler, the forewing a very pale ashy gray and
all the darker pattern markings more or less obscured
or obliterated. I do not think that these varieties
represent anything but color forms or that any one of
them is entitled to a subspecific designation. More
and wider collecting throughout the Middle and Far
West will probably turn up still other color variants.
The species itself, despite its variability, is easily identi-
fied by its genitalia.
279. Telethusia rhypodella (Hulst), new combination
Glyptoteles rhypodella Hulst, Ent. Amer., vol. 3, p. 137, 1887.
Nephopteryx rhypodella (Hulst), Phycitidae of N. Amer., p. 144,
1890.—Ragonot, Monograph, p. 1, p. 270, 1893.—Barnes
and McDunnough, Contributions, vol. 3, p. 196, 1916.—
McDunnough, Check list, No. 6165, 1939.
There are no specimens in the Rutgers Collection or
elsewhere that I have seen matching Hulst’s description,
nor any available Oregon examples that could be re-
ferred to rhypodella. The alleged type at Rutgers, a
female without locality label and bearing only the num-
ber 42, is a typical representative of Phobus curvatella
(Ragonot). Unfortunately the Hulst ‘“types’’ are fre-
quently as unreliable as his type designations and this
particular type is probably spurious. Hulst’s descrip-
tions, on the other hand, are usually more reliable and I
138 UNITED STATES NATIONAL MUSEUM BULLETIN 207
suspect that when sufficient Oregon material is available
we shall find that rhypodella is merely one of the nu-
merous color forms of ovalis.
Typr Locauity: ‘Oregon’ (type lost?).
Foop prant: Unknown.
73. Phobus, new genus
Type or GENus: Dioryctria brucei Hulst.
Characters of Telethusia except: Eighth abdominal
segment of male with a pair of ventrolateral hair tufts.
Penis of male genitalia armed with a single cornutus;
usually also a cluster of very fine, minute, slender spines
at apex of aedeagus (but not on membranous penis).
Transtilla represented at least by its lateral elements,
sometimes the median area is recognizable but is very
weakly sclerotized and the completed band not a con-
stant character. Female genitalia with ovipositor nor-
mal (not strongly sclerotized) ; apophyses of ovipositor
and eighth-segment collar slender; bursa copulatrix
simple (smooth); membrane between collar and ovi-
positor smooth.
The foregoing differences seem to justify separation
from Telethusia, with which the genus is very closely
related.
280. Phobus brucei (Hulst), new combination
Figures 354, 844
Dioryctria brucei Hulst, Canadian Ent., vol. 27, p. 55, 1895.
Ambesa lallatalis Hulst (not Hulst), U. S. Nat. Mus. Bull. 52, p.
422, 1903.
Tacoma lallatalis Dyar (not Hulst), Proc. Ent. Soc. Washington,
vol. 6, p. 227, 1904.
Nephopteryz lallatalis brucei (Hulst) McDunnough, Check list,
No. 6160, 1939.
Forewing whitish more or less dusted and shaded with
blackish or fuscous scales, making the general color
cream white (with some pale brownish shading on paler,
weakly marked examples) to ashy gray with a faint
bluish gray tint (on well-marked specimens) ; the trans-
verse lines irregular and more or less interrupted and
not strongly contrasted; antemedial line oblique, ser-
rate, interrupted at lower fold by a pale olivaceous-
ocherous shade which extends rather broadly the length
of the fold and also cuts the subterminal line; a similar
but somewhat weaker shade fills the cell; outer margin
of antemedial line consisting of a thin blackish line
curving outwardly from costa to top of cell, a small
blackish dot or dash on lower vein of cell and a similar
blackish marking on vein 1b; subterminal line markedly
serrate, deeply indented (almost to cell) below costa, on
well-marked (darker) specimens bordered inwardly by a
black line from costa at least to cell, this line frequently
continued along top of cell to the black outer marking
of antemedial line, forming a continuous, long, narrow
hook along the median and postmedian subcostal area;
below the lower fold the blackish outer border of the
subterminal line is also continued back, as a black line
under vein 1b to and fusing with a narrow blackish line
on the outer edge of the antemedial line forming a nar-
row, oval marking on lower margin between the trans-
verse lines; on dark examples a more or less conspicuous,
blackish, quadrate patch inwardly bordering the ante-
medial line at inner margin; on pale specimens this
patch pale brown, more or less obscured; discal dots
obscured, rarely distinguishable; a row of small narrow
black or brownish dots along termen. Hind wings
subpellucid with a faint ocherous tint; the veins not
appreciably darkened; a faint narrow dark shade along
termen. Alar expanse, 26-29 mm.
Male genitalia with cornutus small, slender. Female
genitalia, with bursa very short, not much longer than
ductus bursae.
Type tocaLiry: Colorado (type in AMNH, ex Rut-
gers).
Foop prant: Unknown.
Distrisution: Colorado; Utah, Eureka (June, July),
Stockton (June, July, Sept.); Nevada, Ormsby, and one
female with only the state locality (a pseudotype of
lallatalis Hulst); California, San Luis Obispo; Washing-
ton, Pullman (July).
All specimens of this species in the National Collec-
tion had been identified as lallatalis Hulst on Dyar’s
misidentification of the latter species. As a result
Dyar referred brucei as a synonym of lallatalis. Hulst,
however, was primarily to blame for the confusion; for
he had identified and sent out as “types” of Jallatalis
specimens of both brucei and Interjectio denticulella.
The true brucei resembles superficially both lallatalis
and denticulella in some of its more striking details of
maculation, but is easily distinguished from both by
its male and female genitalia.
281. Phobus funerellus (Dyar), new combination
Salebria funerella Dyar, Ins. Inse. Menstr., vol. 13, p. 12, 1925.—
McDunnough, Check list, No. 6193, 1939.
Forewing blackish to dark brownish gray more or less
dusted with white on basal, median, and terminal areas;
the white dusting very faint and scattered on the type
series from Southern California, which have a uniform
blackish gray ground color, much stronger on specimens
from Washington, British Columbia, and New Mexico;
transverse lines and thin blackish borders complete (not
interrupted as in brucei), sinuate; the antemedial line
oblique, sharply serrate, narrow, whitish gray, bordered
outwardly by a narrow black line and inwardly by a
broad, unbroken blackish or (on paler examples) dark
gray-brown band, this band distinguishable and con-
trasted even on the darkest, most suffused examples;
subterminal line serrate but not deeply indented below
costa, bordered inwardly by a narrow, continuous black
line, the latter not continued inwardly below costa or
on vein 1b as it is on brucei; subterminal line bordered
outwardly by a rather broad dark band; discal dots
usually distinct, black, more or less confluent; a row of
blackish dots along termen. Hind wing brown; the
veins darkened and a narrow blackish line along termen.
Alar expanse, 24.5-30 mm.
Genitalia similar to those of bruce.
Typz Locauity: Southern California (type in USNM).
Foop puant: Unknown.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 139
Distrisution: Unirep States; California (Southern
California without more definite locality, the type
series), Clarksville (El Dorado County, June); Washing-
ton, Pullman; New Mexico, Fort Wingate (July).
Canava: British Columbia, Departure Bay (Aug.),
Duncans (Vancouver Isl., July), Nicola (July), Welling-
ton (June). Also one male without state locality
labeled ‘‘Larima Co. [sic], Aug., 1901, Schaus collector.”’
The species is distinct and easily distinguished from
brucei on the color and pattern of forewing, especially
by the broad black band extending from inner margin
to costa before the antemedial line and by the shallow
indentation of the subterminal line below costa.
282. Phobus curvatellus (Ragonot), new combination
Ficures 355, 845
Nephopteryx curvatella Ragonot, N. Amer. Phycitidae, p. 7,
1887.—Barnes and McDunnough, Contributions, vol. 3,
p. 196, 1916.—McDunnough, Check list, No. 6166, 1939.
Nephopteryx rhypodella Ragonot (not Hulst), Monograph, pt. 1,
p. 270, 1893.
Forewing ashy bluish gray; the transverse lines com-
plete, narrow, white, obscure except on the well-marked
darker examples, indicated chiefly by the fine, black
outer border of the antemedial line and similar black
inner border of the subterminal] line; preceding the ante-
medial line a quadrate blackish spot on inner margin;
limited above by a weak, smaller, pale, somewhat oilva-
ceous shade in the lower fold, this pale shade not
interrupting the antemedial line itself; indentations of
subterminal line as in funerellus; discal dots obscure,
the lower one sometimes distinct (under magnification)
and frequently with a dark shade below it which forms
a round dark spot, to the naked eye one of the more
conspicuous markings on the wing; a row of more or
less confluent black dots along termen. Hind wing
translucent, whitish with a smoky shade towards apex;
the veins darkened; a fine brown line along termen.
Alar expanse, 26-30 mm.
Male genitalia with cornutus of the same length as
that of brucei and funerellus but somewhat stouter.
Genitalia otherwise like those of the species following
(incertus). Female genitalia with bursa very long, and
narrow throughout its length, but little wider than the
ductus bursae.
TypE Locatity: America Septentrionalis (type im
Paris Mus.).
Foop puant: Unknown.
Distrisution: California, Loma Linda (June, July),
Los Angeles County (1,060 ft., June), Monachee
Meadows (Tulare County, July), Mount Lowe (July),
San Gabriel Mts. (1,700 ft., July); Arizona, Nogales
(May), Santa Rita Mts. (May); Utah, Bellevue
(Washington County, May, June); Colorado, Silverton
(July).
The spurious type of Telethusia rhypodella (Hulst)
in the Rutgers Collection belongs here. It is discussed
under the treatment of rhypodella (p. 137).
283. Phobus incertus, new species
Ficures 356, 846
Color and markings of forewing as in curvatellus
except duller, lacking the bluish tint of the latter
species; the quadrate dark spot preceding the ante-
medial line also continued as a broad band to costa,
though frequently interrupted by a pale shading at
lower fold.
Male genitalia with cornutus appreciably stouter and
longer than that of any of the other species of the genus.
Bursa copulatrix of the female genitalia less than half
the length of that of curvatellus but twice the length of
that of brucei or funerellus.
Type Locauity: Strawberry Valley (6,000 ft.), San
Jacinto Mts., Calif. (type in USNM, 61348).
Described from male type and five male and five
female paratypes from the type locality, collected by F.
Grinnell, Jr., June 16, 17, and 18, 1908.
Except for the genitalic differences this might easily
be a higher altitude race of curvatellus, but the differ-
ences in size of cornutus and length of bursa seem to
be constant characters and greater than to be expected
in variants of one species.
74, Actrix, new genus
TypzE or Genus: Tacoma nyssaecolella Dyar.
Tongue well developed. Antenna weakly pubescent;
on male with sinus and scale tuft in base of shaft.
Labial palpus upcurved, slender, reaching above vertex;
second segment somewhat flattened and very slightly
rough scaled, not grooved on male; third segment about
two-thirds the length of second, acuminate. Maxillary
palpus squamous. Forewing smooth; 11 veins; vein 2
from near lower outer angle of cell; 3 from the angle,
approximately equidistant from 2 and 4 at base; 4 and
5 shortly stalked; 6 from below upper angle of cell,
straight; 8 and 9 very long stalked; 10 from the cell,
approximate to the stalk of 8-9 for some distance; male
without costal fold. Hind wing with vein 2 from before
but near lower outer angle of cell; 3 from the angle,
connate or (on occasional specimens) shortly fused
with the stalk of 4-5; 4 and 5 stalked for two-thirds of
their lengths; 7 and 8 strongly anastomosed beyond
cell for half or a trifle more than half their lengths;
cell slightly less than half the length of wing; dis-
cocellular vein curved, extended outwardly at lower
angle of cell. Eighth abdominal segment of male with
pair of ventrolateral hair tufts.
Male genitalia with uncus broader than long, its
outer lateral angles slightly lobed. Apical process of
gnathos broad, shieldlike, its lateral arms greatly
reduced. ‘Transtilla absent. Harpe without clasper.
Aedeagus straight, divided towards apex, the projecting
divided elements strongly sclerotized; penis with some
very weak scobinations, but otherwise unarmed.
Vinculum short, stout, about as long as greatest width,
evenly rounded to blunt terminal margin.
140 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Female genitalia with bursa small, covered with
dense slender spines on posterior half, otherwise very
finely spinose, without signum; ductus bursae granu-
late, short, expanded into a broad, strongly sclerotized,
contorted plate towards genital opening; ductus semi-
nalis from lobe of bursa near junction of bursa and
ductus bursae.
The genus is a further restriction from Nephopteryz
of authors. Its nearest relationship seems to be Pyla,
which it resembles in the peculiar modification of the
ductus bursae of the female and the aedeagus of the
male. Its venation, however, is quite distinct and,
except for the strong anastomosis of veins 7 and 8 of
hind wing, closer to that of Tacoma.
284. Actrix nyssaecolella (Dyar), new combination
Ficures 357, 838
Tacoma nyssaecolella Dyar, Proc. Ent. Soc. Washington vol. 6,
p. 112, 1904.—Forbes, Cornell Mem. 68, p. 621, 1923.—
Craighead, U. S. Dep. Agr. Misc. Publ. 657, p. 453, 1950.
Nephopteryx nyssaecolella (Dyar) Barnes and McDunnough,
Contributions, vol. 3, p. 196, 1916—McDunnough, Check
list, No. 6174, 1939.
Forewing brownish gray (“lilaceous gray”) paler in
basal area beyond extreme base, in the half of median
area just beyond antemedial line, and to a lesser extent
in outer area (beyond the subterminal line), these pale
areas of an ashy hue, due to a faint peppering of white
scales; antemedial line narrow, oblique and very slightly
angled at middle, dull white, preceded on inner margin
by a subquadrate blackish brown patch and bordered
outwardly at costa by a triangulate black patch which
(on most specimens) continues as a more or less inter-
rupted black line to inner margin; subterminal line
sinuate (bulged at middle) and weakly serrate, dull
white, bordered inwardly by a dark shade and outwardly
by a narrower dark line, these dark borders especially
marked and blackish at costa; discal dots confluent,
blackish; a row of more or less confluent black dots
along termen. Hind wing pale smoky brown, with a
glossy sheen; veins very faintly darkened; a narrow
brown line along termen. Alar expanse, 15-18 mm.
Male genitalia with terminal margin of uncus con-
cave, its lateral lobes turned laterally outward. Apical
process of gnathos a convex shield, slightly longer than
broad and with apical, lateral angles produced backward
into bluntly pointed spines. Divided elements of aedea-
gus produced as short, sharply and oppositely curved
hooks. Cucullus of harpe sharply curved towards its
apex. Female genitalia distinguished at once by the
shape of the ventral sclerotized plate of ductus bursae
and the eighth-segment collar, which is complete and
strongly sclerotized ventrally.
Typr Locatity: Washington, D.C. (typein USNM).
Foop prant: Nyssa sylvatica (larva a leaf-folder).
Distrisution: Massachusetts, Nantucket (July);
Connecticut, Kast River (July, Aug.); Rhode Island,
Weekapaugh (July) ; New Jersey, Anglesea (May, June);
Pennsylvania, New Brighton (July, Aug.), Oak Station
(Aug.) ; District of Columbia, Washington (Aug.); North
Carolina, Southern Pines (Aug.), Tryon (Aug.).
285. Actrix dissimulatrix, new species
Fieures 358, 837
Superficially not distinguishable from nyssaecolella
except (on the specimens before me) a slightly stronger
white dusting on median area of forewing especially
over inner margin immediately following the antemedial
line. This difference can hardly be expected to hold
for the species.
The genitalia of both sexes are very different from
those of nyssaecolella. Male genitalia with terminal
margin of uncus evenly rounded, its lateral lobes turned
inward and downward, partially encircling the anal
tube. Apical process of gnathos heart shaped, without
produced angles. Divided elements of aedeagus pro-
duced as long, straight, stout thornlike projections, one
of which is coarsely scobinate. Female genitalia with
a stout pair of strongly sclerotized, curved, lateral arms
projecting forward from the ventral sclerotized plate
of the ductus bursae.
Typ Locatity: Cape Henry, Va. (type in USNM,
61349).
Foop prant: Nyssa sylvatica.
Described from male type and two male and one
female paratypes from the type locality, reared by A.
Busck, Aug. 10, 16 and 18, 1927, from larvae feeding on
the leaves of Nyssa sylvatica. Four larvae were pre-
served from the collection. Three of these are blackish
brown in color and are undoubtedly nyssaecolella. One
larva is yellow with a pale yellowish head and thoracic
shield. Itis probably dissimulatriz. No difference was
noted in larval habits between the two forms.
75. Genus Stylopalpia Hampson
Stylopalpia Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7,
p. 257, 1901. (Lype of genus: Stylopalpia lunigerella
Hampson.)
Tongue well developed. Antenna very shortly pubes-
cent, shaft of male somewhat flattened and with a
shallow sinus at base containing a row of short spines
more or less concealed by a small, weak scale tuft.
Labial palpus obliquely upturned; third sesment very
long (nearly twice the length of second), slender and
porrect in lunigerella, much shorter (about half the
length of second) and oblique in the other two species,
in these reaching a little above vertex, acuminate in all
species. Maxillary palpus minute, its scaling slightly
expanded. Forewing smooth; 11 veins; vein 2 from
before the lower outer angle of cell; 3 from the angle;
4 and 5 separated at base and divergent beyond, 4 but
slightly nearer to 5 at base than to 3; 6 from below upper
angle of cell, straight; 8 and 9 stalked for slightly more
than half their lengths; 10 from the cell, well separated
from the stalk of 8-9 at base but just beyond approach-
ing it for a very short distance; male without costal
fold. Hind wing with vein 2 from well before lower
outer angle of cell; 3 from the angle; 4 and 5 stalked
for about half their lengths; 7 and 8 approximate or
contiguous for a very short distance beyond cell; cell
less than half the length of wing; discocellular vein
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 141
curved, produced at lower angle, a short spur connecting
it and vein 8. Highth abdominal segment with weak
compound tufts in lwnigerella, paired ventral tufts in
the other two species.
Male genitalia with uncus subtriangulate, its terminal
margin more or less broadly rounded. Apical process
of gnathos a simple, rather short, stout hook. Trans-
tilla incomplete, sclerotized only in its reduced lateral
elements (the central portion, as shown in the figures,
distinguishable but not sclerotized). Harpe with large,
strongly sclerotized, erect, scoop-shaped clasper, situ-
ated towards base, below costa; cucullus narrow,
elongate, very slightly tapering to rounded apex.
Anellus shield-shaped, with small lateral lobes. Aedea-
gus expanding to lateral, flanged projections before
apex, the latter flanges, each bearing a cluster of strong
spines; penis unarmed. Vinculum stout; slightly longer
than broad; tapering slightly to broadly rounded ter-
minal margin.
Female genitalia without signum. Bursa mem-
branous with only a little fine spining at junction of
bursa and ductus bursae; ductus bursae weakly scle-
rotized at, and just before, genital opening, the latter
broad; ductus seminalis from middle of ductus bursae.
The genus was originally erected on the peculiar
palps of its type species. This character, however,
proves to be specific rather than generic. The two
species here included do not have it; but agree with
the type species on every other detail of venation,
antennal structure, and genitalia. The genus can
easily be maintained on its combination of male and
female genitalic characters.
286. Stylopalpia lunigerella Hampson
Ficures 30, 359, 848
Stylopalpia lunigerella Hampson, Ann. Mag. Nat. Hist., ser. 7,
vol. 7, p. 258, 1901.
Third segment of labial palpus very long, slender
and porrect. Eighth abdominal segment of male with
compound tufts.
Forewing ocherous (clay color) dusted with blackish,
making the general shade dark gray, the ocherous color
forming a contrasting band along costa and more or
less lightening the lower median area and the base of
inner margin; antemedial line indicated by a pale
lunulate line between cell and inner margin and, on
its upper half, by faint traces of its narrow, blackish,
outer border; subterminal line very close to outer
margin, slightly bulged at middle, not serrate, pre-
ceded by some black streaklets on the veins and whitish
or pale ocherous streaklets between them; discal dots
separated, blackish; a few of black dots along termen.
On female a somewhat broader brownish shade at apex
and along termen and some darkening of the outer
parts of the veins. Alar expanse, 18-24 mm.
Male genitalia with clasper of harpe considerably
longer than deep, serrate along lower and inner margins.
Female genitalia with genital opening very broad.
Type Locauity: Jamaica (type in BM).
Foop pirant: Unknown.
DistriputTion: Banamas, Nassau. Purrto Rico:
Aguirre Central (Apr., June, Aug.), Camuy (Apr.),
Coamo Springs (Apr.), Ponce (Sept.), San Germén
(June). Cua: Santiago Province (June, July, Oct.).
Jamaica. M*&xtco: Colima (Jan., June, July).
The species is easily identified by its peculiar palpi,
alike in both sexes.
287. Stylopalpia scobiella (Grote), new combination
Ficure 360, 847
Nephopteryx scobiella Grote, N. Amer. Ent., vol. 1, p. 5, 1890.—
Hulst, Phycitidae of N. Amer., p. 148, 1890.—Ragonot, Ent.
Amer., vol. 5, p. 115, 1889; Monograph, pt. 1, p. 266, 1893.—
McDunnough, Check list, No. 6161, 1939.
Lipographis decimerella Hulst, Ent. Amer., vol. 4, p. 117, 1888.
Third segment of labial palpus short, oblique. Except
on a few of the grayer specimens, head and thorax
distinctly ocherous.
Forewing pale gray to grayish ocherous, extreme base
of wing ocherous and a similar pale ocherous shade
rather broadly bordering the costa; transverse markings
nearly obliterated; antemedial line indicated only by a
small Junate white spot on vein 1b, preceded and fol-
lowed by black dots, a similar blackish dot or streaklet
at lower margin of cell (representing a median fragment
of the usual black outer border of the antemedial line) ;
subterminal line obscure, a very faint pale line weakly
bordered inwardly by a darker shade; lower discal spot
a blackish streaklet, upper discal dot usually absent, if
present very faint; a row of fine blackish dots along
termen. Hind wings whitish with a faint ocherous or
smoky tint; the veins little if any darkened; a thin
brownish line along termen. Alar expanse, 24-26 mm.
Male genitalia with clasper of harpe broadly oval,
not serrate. Terminal margins of uncus and vinculum
very broadly rounded, the vinculum not appreciably
tapering. Eighth abdominal segment of male with
paired ventral tufts. Female with genital opening more
constricted than that of lunigerella.
Typ LocaLitiEs: Bosque County, Tex. (scobiella, in
BM); Blanco County, Tex. (decimerella, in AMNH,
ex Rutgers).
Foop pirant: Unknown.
Distrisution: Texas, Barber (Apr.), Beeville (May),
Blanco County (May), Bosque County, Burnet County
(Apr., Sept., Oct.), Kerrville, Sabinal (Mar., Apr.),
Sapulpa (May), San Diego (Apr.), Victoria (Apr., Sept.),
Zavalla County (Apr.); Colorado, Glenwood Springs.
Also four specimens from Texas with only the state
locality. The species probably also occurs in northern
México.
288. Stylopalpia argentinensis, new species
Ficure 361
Labial palpus and eighth abdominal tufts as in scobi-
ella. 'The head and thorax brown.
Forewing pale brown with a strong dusting of white
scales faintly peppered with black in median area; basal
142 UNITED STATES NATIONAL MUSEUM BULLETIN 207
area brownish ocherous, shading to brown at extreme
base and with a clouding of white in midbasal area; ante-
medial line complete, well out on wing, oblique, inwardly
notched at vein 1b, white, bordered outwardly by a
narrow blackish line; subterminal line also complete,
further back from termen than in the two preceding
species, oblique (parallel to termen), whitish ocherous
with a narrow blackish inner border; discal dots distinct,
separated, black. Hind wing dark brown; the fringe
whitish ocherous with a dark median band. Alar ex-
panse, 24 mm.
Male genitalia similar to those of scobiella except:
Uncus narrower and tapering to more narrowly rounded
terminal margin; vinculum longer in proportion to its
width; spines on aedeagus fewer and coarser.
TyrEeocauity: “Villa Anna, F.S.C. Fe., Argentina”’
(type in BM).
Foop prant: Unknown.
Described from unique male, collected by K. J.
Hayward, Dec. 1925.
76. Genus Pyla Grote
Pyla Grote, New check list of North American moths, p. 55,
1882.—Ragonot, N. Amer. Phycitidae, p. 9, 1887; Mono-
graph, pt. 1, p. 481, 1893.—Hulst, Phycitidae of N. Amer.,
p. 161,1890. (Typeof genus: Nephopterysx scintillans Grote.)
Tongue well developed. Antenna finely pubescent;
on male with sinus and scale tuft in base of shaft.
Labial palpus oblique; second segment broadly scaled,
somewhat flattened laterally, reaching above vertex; on
male with a slight groove to hold the maxillary palpus;
third segment short, less than one-third the length of
second, porrect. Maxillary palpus of male in the form
of an aigrette, semiaigrette (i. e., the scales hairlike
but short), or more or less squamous. Forewing smooth;
11 veins; vein 2 from near lower outer angle of cell; 3
from the angle, slightly nearer to 4 at base than to 2;
4 and 5 slightly separated at base, parallel for a short
distance beyond cell; 6 from below upper angle of cell
straight; 8 and 9 stalked from one-half to a third of
their lengths; 10 from the cell, slightly separated at
base from the stalk of 8-9; male without costal fold.
Hind wing with vein 2 from before but rather near
lower outer angle of cell; 3 from the angle, connate with
4; 4 and 5 stalked or anastomosed for half or slightly
less than half of their lengths; 7 and 8 contiguous or
very weakly anastomosed for a short distance beyond
cell; cell less than half the length of wing; discocellular
vem curved, outwardly produced at lower angle and
connected with vein 3 by ashort spur. Eighth abdom-
inal segment of male with a pair of ventrolateral hair
tufts (fig. 372b), or two or three pairs containing some
modified scales (fig. 367c).
Male genitalia with uncus broad, more or less trian-
gulate. Apical process of gnathos a short, stout hook.
Transtilla absent. Harpe usually with base of costa
produced into a strong projecting hook, or spine, or
spined lobe; frequently a strong hooked or spined clasper
from median basal area; sacculus simple; costa strongly
sclerotized but sclerotization rather abruptly terminated
before apex of harpe. Aedeagus usually partially divided
(bifurcate) or armed with projecting spine or spines,
rarely simple; penis unarmed (except for a very weak
cornutus in fusca). Vinculum stout, slightly tapered to
truncate or more or less broadly rounded terminal
margin.
Female genitalia without signum; bursa copulatrix
usually simple, sometimes with strongly sclerotized,
convolute, longitudinal bands near junction of bursa
and ductus bursae and extending a short distance into
the ductus; ductus bursae short, widening to broad gen-
ital opening, usually strongly and elaborately sclero-
tized towards genital opening; genital opening rarely
simple (fasciolalis, mridisuffusella); ductus seminalis
from bursa near junction of bursa and ductus bursae.
The genus as here defined includes what superficially
appears to be two distinct entities, one group of species
with gray forewings and another with shiny brown
wings, the latter the typical Pyla of authors. For con-
venience of identification I am designating them as
species groups. The division is not supported by any
consistent structural character or combination of char-
acters. The differences exhibited by the several species
in male and female genitalia, male maxillary palpi, and
male abdominal tufts are striking, but apparently only
of specific significance.
Nothing is known of the food habits or early stages of
any of the species except fusca, which is recorded from
Erica in the Old World. I suspect that the Hricaceae
will prove to be the chief hosts of the genus.
Genus Pyla, Species 289-297: P. fasciolalis to
P. hanhamella
[Ground color of forewing gray.]
289. Pyla fasciolalis (Hulst), new combination
Figures 362, 3638, 849
Pinipestis fasciolalis Hulst, Trans. Amer. Ent. Soce., vol. 13, p.
162, 1886.
Nephopteryx fasciolalis (Hulst) Ragonot, Ent. Amer., vol. 5, p.
115, 1889; Monograph, pt. 1, p. 271, 1893 (?).—Hulst,
Phycitidae of N. Amer., p. 144, 1890.—McDunnough, Check
list, No. 6167, 1890.
Maxillary palpus of male squamous.
Forewing gray finely powdered with white, giving the
wing an ashy gray appearance; antemedial line distinct
throughout, whitish, oblique, notched at top of cell and
on lower fold, bordered outwardly by a black line begin-
ning as a black smudge on costa, inwardly by a mod-
erately broad black line extending from inner margin to
cell; subterminal line well marked, bulged at middle and
more or less dentate, bordered inwardly by a blackish
band and outwardly by a somewhat broader, fainter
dark band, these dark borders strongest near costa;
discal dots distinct, small, normally separated, occa-
sionally partially coalesced; a row of small black dots
along termen. Hind wing smoky white with a pale
brownish tint; veins very faintly darkened ; a narrow dark
shade along termen. Alar expanse, 27-30 mm.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 143
Male genitalia with harpe simple; clasper vestigial.
Aedeagus deeply divided; one of the divided elements
slightly forked at apex. Female genitalia with strongly
sclerotized, convulute bands in bursa; genital opening
simple.
TYPE Locauity: British Columbia (type in AMNH,
ex Rutgers).
Foop prant: Unknown.
Distrizution: British Columbia, Goldstream (Aug.),
Necola (July), Saunichton (July).
The only specimen in the Rutgers Collection is a large
male (30 mm.) from British Columbia labeled “Dioryc-
tria fasciolalis Hulst, Type.” I think we may safely
assume this to be the actual type although in his original
description Hulst gives ‘‘Nevada”’ as the type locality,
probably one of his characteristic lapses. A perfect
match for the type in color, markings, and genitalia, is
found in a specimen from Necola, in the Canadian Na-
tional Collection. The other records cited above (Gold-
stream and Saanichton) are from specimens in the U. 8.
National Collection.
Also before me are four examples of what I take to be
a variety of fasciolalis—two males from Gunnison
County, Colo. (July), and 2 females from Wallace,
Idaho (Aug.). Their blackish markings on forewing
are a trifle stronger, the vinculum of male genitalia (fig.
363) is somewhat shorter than in typical fasciolalis. The
convolute bands in the bursa of the female are also a
trifle longer. I doubt very much if these differences
ndicate anything more than a possible local race.
290. Pyla impostor, new species
Fieures 364, 850
Maxillary palpus of male in the form of an aigrette.
Forewing color and markings similar to those of
fasciolalis except: Somewhat duller and darker; white
dusting sparser; the transverse pale lines fainter, in
some specimens much obscured; their blackish borders
less strongly contrasted against the ground color of the
wing. Hind wing pale smoky fuscous, the brownish
tint of fasciolalis very faint or altogether lacking. Alar
expanse, 23-30 mm.
Male genitalia with a long, slender, somewhat flat-
tened, outwardly curved clasper on harpe; base of costa
not modified. Aedeagus divided for about half its
length, moderately slender, the divided elements rigid,
pointed and unforked at their apices. Female with
convolute, sclerotized folds extending from bursa shortly
into ductus bursae; ductus bursae itself partially sclero-
tized, the sclerotization forming broad ventrolateral
bands extending from just beyond the convolute folds
of the bursa to genital opening.
TYPE Loca.ity: Slate Peak, Whatcom County, Wash.
(6,000-7,000 ft.; type in USNM, 61350; paratypes in
USNM and Canadian Nat. Coll.).
Foop prant: Unknown.
Described from male type and one female paratype
from the type locality, collected by J. F. G. Clarke, Aug.
2, 1940; and paratypes from the following localities:
Bogachiel Peak, Olympic Mts., Wash., Aug. 9, 1936,
Dr. A. F. Braun (?); Chimney Gulch, Golden, Colo.,
Oslar (oc); Colorado with only the state locality (a
pseudotype, o, of fascialis Hulst, from the Fernald
Collection); Big Belt Mts., Mont., July 18, 1928, J.
McDunnough (co); upper Gallatin Canyon, Mont.,
7,000 ft., July 4, 1928, J. McDunnough (c’); Banff,
Alberta, July 20, 1925, Owen Bryant, (co); Lethbridge,
Alberta, July 3, 1922, H. L. Seamans (c’); Moraine
Lake, Alberta, July 3, 4, 7, 1923, J. McDunnough (4 &*
and 2 °); Waterton Lakes, Alberta, July 23, 28, 1923,
J. McDunnough (co and 2); Hope Mts., British Co-
lumbia, July 22, 1932, A. N. Gartrell (o&’); Mount Rey-
elstoke, British Columbia, 6,000 ft., July 12, 1923, E.
E. Buckell (0).
Most of the foregoing were in our collections as
fasciolalis on the basis of the false Hulst type in the
National Museum. It is superficially like fasciolalis,
but a distinct mountain-top species easily identified by
its genitalia.
291. Pyla aequivoca, new species
Fiaures 366, 855
Maxillary palpus of male in the form of an aigrette.
Superficially like ampostor; the ground color of the
male forewing a trifle more brownish gray, and the dark
outer border of antemedial line somewhat broader and
diffused into the ground color at costa, differences that
could not be expected to hold in any extended series.
Distinguished by its genitalia. Alar expanse, 26-29 mm.
Male genitalia with clasper arising from midbasal
area of harpe as in impostor, but much shorter and angu-
late. Anellus U-shaped, its lateral arms long and
slender. Aedeagus much shorter and somewhat stouter;
its divided elements broader, decidedly flattened, termi-
nating in sharp spines and more or less laterally spined
towards apices. Eighth abdominal segment of male
with two pairs of hair tufts.
Typr Ltocauity: Banff, Alberta, Canada (type in
Canadian Nat. Coll., paratype in USNM, 61351).
Foop puant: Unknown.
Described from male type and three male paratypes
from the type locality, collected by C. B. D. Garrett,
June 21 and 30 and July 6, 1922; and one male paratype
from Hymers, Ontario June 8, 1915. In addition to the
type series I have before me a female from the Canadian
National Collection collected at Aweme, Manitoba,
Aug. 26, 1921, by Norman Criddle. Its genitalia differ
markedly from those of impostor. The membrane of
bursa is thickened near junction with ductus bursae but
lacks the sclerotizations of impostor; the ventrolateral
bands of the ductus bursae are differently shaped and
fuse into the ventrolateral sclerotizations of the inter-
segmental area before the eighth-segment collar. While
I have little doubt that this female is conspecific with
the males, I am not designating it as a paratype.
144
292. Pyla insinuatrix, new species
Fiaures 365, 856
Maxillary palpus of male in the form of an aigrette.
Forewing paler than in the preceding species, due to
stronger white dusting, giving the paler areas a faint
bluish tint; transverse lines and dark markings more
strongly contrasted; the white antemedial line especially
well marked on its lower half, its inner black border
below cell expanded into a moderately wider blackish
band or patch; blackish inner and outer borders of sub-
terminal line well marked, especially the former; discal
dots conspicuous, tending to coalesce. Hind wing
subpellucid smoky white, darkening towards apex and
termen; the veins very faintly darkened and a narrow
dark line along termen. Alar expanse, 24-26 mm.
Male genitalia with uncus hoodlike, constricted to-
wards base. Harpe with a strong, outwardly produced,
spined, knob from base of costa; no appreciable clasper.
Aedeagus slender, with a very slight bifurcation at
apex; the bifurcate projections straight. ‘Two pairs of
ventrolateral tufts on eighth abdominal segment; some
of the hairs broadly expanded at their apices (as in
aenigmatica, fig. 367¢c).
Female genitalia with bursa copulatrix membranous
except for a very faint sclerotization of the lobe giving
off the ductus seminalis; ductus bursae flattened,
weakly sclerotized, expanding at genital opening into
sclerotized, scobinate ventrolateral lobes.
Type Locatiry: Aweme, Manitoba, Canada (type in
Canadian Nat. Coll.; paratypes in USNM, 61352).
Foop pranr: Unknown.
Described from male type and four male and two
female paratypes from the type locality, collected by
Norman Criddle July 13 and Aug. 10, 1925; Aug. 10,
1921; Aug. 10, 1925; Aug. 19, 1915).
293. Pyla aenigmatica, new species
Figures 367, 853
Mazxillary palpus of male in the form of a semiaigrette
(the hairs short).
Forewing as on insinuatriz except darker, the ground
color like that of impostor; the transverse lines distinct;
lower half of antemedial line bordered inwardly by a
subquadrate blackish patch, costal half of the outer
dark border rather broad and well contrasted, blackish;
the dark borders of subterminal line well contrasted
towards costa; discal dots confluent; dots along termen
minute, weak. Alar expanse, 25-28 mm.
Male genitalia with tegumen having two, strongly
spined, protruding lobes on each ventrolateral margin.
Harpe with base of costa enlarged and coarsely scobi-
nate; clasper small, semicircular, erect. Aedeagus
slender; shortly bifurcate; bifurcate elements at apex
spinelike, bent sharply at right angles to the aedeagus.
Anellus a semitubular shield with rather long, strongly
sclerotized lateral arms. Eighth abdominal segment
with two pairs of ventrolateral hair tufts; one pair with
the hairs expanded at their apices (forming knoblike
clusters).
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Female genitalia with bursa membranous. Ductus
bursae short, strongly sclerotized along lateral margins,
the sclerotizations expanding laterally and at right
angles at genital opening into a pair of convolute, finely
scobinate lobes.
Typrz LocaLity: Wellington, British Columbia (type
in USNM, 61353; paratype in Canadian Nat. Coll.).
Foop prant: Unknown.
Described from male type and one male paratype
from the type locality, “‘21-VI-04,” G. W. Taylor, and
paratypes from the following localities: Goldstream,
British Columbia, “30-VIII-20” (9); Salmon Arm,
British Columbia, ‘‘22-6-20, W. R. B.” (co); Pine
Grove, Colo., July 8, 1901, H. G. Dyar, “17310” (&);
East River, Conn., Aug. 21 and Sept. 3, 1908, C. R.
Ely (f and 9); Oak Station, Pa., Aug. 20, 1911, Fred
Marloff (#); Watchung Mts., N. J., “64-99,” W. D.
Kearfott ().
The hind wings are a trifle darker on the eastern
examples, which were, in our collection, identified as
Susca.
The species is evidently closely related to insinuatriz,
but is distinct and easily distinguished by its genitalia.
294. Pyla criddlella Dyar
FieurEe 368
Pyla criddlella Dyar, Journ. New York Ent. Soc., vol. 15, p. 110,
1907.—McDunnough, Check list, No. 6241.
Maxillary palpus of male squamous.
Forewing blackish gray, semilustrous, unicolorous
except for a slight darkening of the ground color border-
ing the transverse lines; the latter very faint, but slightly
lighter than the ground color; discal and terminal dots
obscured. Hind wing brownish gray. Alar expanse,
18 mm.
Male genitalia with harpe simple except for a greatly
reduced, upcurving, triangulate clasper. Aedeagus
deeply bifurcate; the right divided element (in ventral
view) produced into a sharp abruptly curved hook. A
single pair of ventrolateral hair tufts on eighth abdom-
inal segment.
Typ Locatity: Aweme, Manitoba, Canada (June;
type in USNM).
Foop ruant: Unknown.
Known only from the male type.
295. Pyla fusca (Haworth), new combination
Ficures 369, 852
Phycis fusca Haworth, Lepidoptera Brittanica, pt. 3, p. 493, 1828.
Phycita fusca (Haworth) Stephens, Illustrations of British ento-
mology, Haustellata, vol. 4, p. 310, 1834.
Pempelia fusca (Haworth) Stainton, Manual of British butterflies
and moths, vol. 2, p. 176, 1859.—Packard, Ann. Lye. Nat.
Hist. New York, vol. 10, p. 271, 1873.
Nephopteryx moestella Walker, List, vol. 27, p. 53, 1863.
Eudorea (?) frigidella Packard, Proc. Boston Soc. Nat. Hist.,
vol. 11, p. 53, 1866.
Salebria fusca (Haworth) Heinemann, Die Schmetterlinge
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 156,
1865.—Grote, Bull. U. S. Geol. Geogr. Surv. Terr., vol. 4,
p. 695, 1878; North Amer. Ent., vol. 1, p. 11, 1879.—
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 145
Staudinger and Rebel, Catalog der Lepidopteren des palae-
arctischen Faunengebietes, vol. 2, p. 34, 1901.—Spuler, Die
Schmetterlinge Europas, vol. 2, p. 211, 1910.—Meyrick,
Revised handbook of British Lepidoptera, p. 380, 1928.—
Ford, Guide to the smaller British Lepidoptera, p. 10, 1949.
Pinipestis cacabella Hulst, Ent. Amer., vol. 3, p. 133, 1887.
Laodamia fusca (Haworth) Ragonot, Ent. Amer., vol. 5, p. 115,
1889; Monograph, pt. 1, p. 408, 1893.—Hulst, Phycitidae
of N. Amer., p. 156, 1890; U. 8. Nat. Mus. Bull. 52, p. 425,
1902.—Forbes, Cornell Mem. 68, p. 628, 1923.—MceDun-
nough, Check list, No. 6227, 1939.
Salebria triplagiatella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 109, 1904.—Barnes and McDunnough, Contributions,
vol. 3, p. 196, 1916.
Dioryctria fusca (Haworth) Pierce and Metcalfe, The genitalia of
the British Pyrales, p. 3, pl. 2, 1938.
Maxillary palpus of male in the form of an aigrette.
Forewing blackish gray, generally of a duskier hue
than that of any of the preceding species; transverse
lines usually faint and a dull whitish gray, rarely con-
trasted against the ground color and when so, chiefly
the lower half of antemedial line; the latter bordered
outwardly at costa and inwardly at inner margin by
blackish patches more or less contrasted against the
ground color of the wing; a similar dark shade inwardly
bordering the subterminal line; discal and terminal dots
tending to coalesce, black. Hind wing dusky white
between the veins; the latter appreciably darkened; a
smoky shade along termen. Alar expanse, 25-30 mm.
Male genitalia with harpe simple except for a thin,
saucer-shaped, erect clasper with a toothlike projection
from its upper inner angle. Aedeagus slender with its
anterior end abruptly expanded and anterior margin
straight (as in hypochalctella); one side produced into
an extended, strongly sclerotized arm, sharply bent and
pointed at apex (as in criddlella)*; penis armed with a
single, moderately long, hairlike cornutus. Three pairs
of ventrolateral hair tufts on eighth abdominal segment
of male, some of the hairs broadly expanded at their
apices.
Female genitalia with bursa membranous; ductus
bursae sclerotized for most of its length, the sclerotiza-
tions extending for a short distance into the bursa;
genital opening simple except for some weak granula-
tions on and behind the ductus bursae.
TYPE LOCALITIES: England (fusca, in BM); eastern
Canada (moestella, in BM); Caribou Is., Labrador
(frigidella, in MCZ); “New York’ ® (cacabella, in
AMNH, ex Rutgers); Winnipeg, Manitoba (triplagia-
tella, in USNM).
Foon piants: Erica and probably some other Hrica-
ceae. The only authentic Old World record is Erica
(Meyrick 1938, Ford 1949). Ragonot (Monograph,
p. 408) records Vaccinium myrtellus and Salix caprea as
probabilities; but these plants only on the basis of food
accepted in the laboratory by larvae hatched from eggs
from gravid females by Porritt (Ent. Monthly Mag.,
5 This extension of aedeagus was misidentified by Pierce and
Metcalfe (1938) as a cornutus. They overlooked the true
cornutus attached to the vesica.
6 So given in Hulst’s original description.
ever, bears no locality label.
The male type, how-
vol. 19, p. 11, 1882). A female in the U. S. National
Museum from Ottawa, Canada, was reared by James
Fletcher (Aug. 1889) from a “black larva’’ found on
Betula. Isuspect, however, that the larva had migrated
to that plant. We have no other New World rearing
records.
Distrrisution: Holarctic. In the Old World from
Great Britain to Japan. The American records from
specimens are: Unirep Srarus: Maine, Orono; New
Hampshire, Hampton (June), Mount Washington (July) ;
Massachusetts, Framingham (July), Martha’s Vineyard
(Aug.); New York, Rochester (June), Waterville (Aug.);
Colorado, Glenwood Springs (Aug.); Washington, Pull-
man. Canapa: Newfoundland, Port aux Basque (Aug.),
St. George Bay (Harry’s River and Stephenville, Aug.),
Spruce Brook (Aug.); Labrador, Caribou Isl., Hopedale,
Nain; Nova Scotia, Baddeck (Cape Breton Isl., Aug.);
Quebec, Chelesea (May); Ontario, Albany River (St.
Martin’s Falls), Hymers (Aug.), Ottawa (June, Aug.);
Manitoba, Aweme (June, July, Aug.), Winnpieg; Alberta,
Banff (July), Calgary (Aug.), Edmonton (May); British
Columbia, Fraser Mills (June), Kaslo (July, Aug.),
Shawnigan Lake (Aug.), Victoria (July). AwasKa:
Cordova, Fort Yukon, Juneau (July), Rampart (July).
The species can be readily distinguished by its aede-
agus, threadlike cornutus, and the peculiar sclerotization
of its ductus bursae. On habitus and all its structural
characters it is closely related to the gray-winged
species of Pyla. Superficially it could easily be con-
fused with impostor, equivoca, or aenigmatica. It is not
congeneric with faecella (Zeller), the type of Laodama,
to which genus Ragonot referred it. The latter differs
markedly in male and female genitalia (figs. 427 and
885), and on venation falls into our venational group D.
Both Pyla and Laodamia have the cell of hind wing
short; but in Laodamia vein 3 is appreciably longer in
relation to vein 2 (fig. 52).
Packard’s frigidella was retained by Ragonot as a
separable variety from fusca, but it is at most only one
of its color variants and is not entitled to any trinomial
designation as a race.
Several Old World references and synonyms have
been omitted from the above synonymy. I do not
question them, but have not been able to verify them.
Anyone interested will find the names and references in
Hulst (Phycitidae of N. Amer., 1890) and Ragonot
(Monograph, 1893).
296. Pyla hypochalciella (Ragonot), new combination
Fieures 370, 854
Nephopteryx ovalis hypochalciella Ragonot, N. Amer. Phycitidae,
p. 7, 1887.—Hulst, Phycitidae of N. Amer., p. 144, 1890.
Nephopteryx hypochalciella Ragonot, Monograph, pt. 1, p. 272,
1893.—McDunnough, Check list, No. 6168, 1939.
Pyla blackmorella Dyar, Ins. Inse. Menstr., vol. 9, p. 68, 1921.—
McDunnough, Check list, No. 6248, 1939. (New syn-
onymy.)
Maxillary palpus of male in the form of an aigrette.
Forewing very dark gray-brown, the dark areas of
some of the darkest specimens almost black;a very faint
146 UNITED STATES NATIONAL MUSEUM BULLETIN 207
powdering of white on costal half of median area and,
narrowly, along terminal margin; the transverse lines
powdery, grayish white, obscure on some specimens;
antemedial line oblique, expanded slightly towards costa,
without distinct inner dark border and with but faint
indication of a blackish brown outer bordering shade
towards costa; subterminal line more or less obscured, on
well-marked examples preceded by a thin blackish line
and followed by a broad band of the darkest prevailing
ground color; discal dots black, well separated; a row of
small black dots along termen, tending to fuse and on a
few specimens forming a fine black line. Hind wing a
uniform very dark satiny brown; the cilia paler, shading
from pale brown to white at their tips. Alar expanse,
22-26 mm.
Male genitalia having harpe with an erect clasper
armed along its outer margin with a row of stout spines
(in the figure this looks like an enlargement of the base
of costa, but it arises below costa and the base of the
costa itself is simple). Anellus bearing two pairs of
comblike, heavy spines, one pair ventral, one dorsal, the
latter situated behind the former. Aedeagus slender;
its anterior end abruptly expanded and the anterior
margin straight; apical fourth bifid, the divided elements
terminating in laterally curved horns. A single pair
of simple hair tufts on eighth abdominal segment of
male.
Female genitalia with bursa membranous; ductus
bursae partially flattened, sclerotized throughout, the
sclerotization expanding abruptly into a wide funnel at
genital opening.
Type LocaLitiEs: ‘Washington Territory” (hypo-
chalciella, in Paris Mus.); Mount Tzouhalem, southern
Vancouver Isl., British Columbia (6lackmorella, in
USNM).
Foop prant: Unknown.
Distrisution: Unitep Starrs: Washington, Friday
Harbor (June, July). Canava: British Columbia (south-
ern Vancouver Isl.), Cowichan District (June), Duncans
(June), Mount Malahat (June), Mount Tzouhalem
(June).
The species is easily identified by its peculiarly armed
anellus. In ground color of forewing it and the species
following (hanhamella) are intermediate between typical
gray- and brown-winged members of the two Pyla
species groups. However, except for a very faint trace
of it in hanhamella, they both lack the bronzy luster on
forewing so characteristic of the typical brown group.
297. Pyla hanhamella Dyar
Figures 371, 860
Pyla hanhamella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 109,
1904.—McDunnough, Check list, No. 6239, 1939.
Maxillary palpus of male subsquamous (a short brush
of hairs mixed with flattened scales).
Forewing color and markings as in hypochalciella
except for traces of a metallic sheen at base. Hind
wing pale brown; cilia white with a fine dark subbasal
line. Alar expanse, 20-24 mm.
Male genitalia having clasper of harpe a moderately
long, erect spike; costa at base simple (not produced).
Aedeagus slightly bent towards middle; shortly bifid at
apex, the divided elements coarsely scobinate. A single
pair of simple hair tufts on eighth abdominal segment.
Female genitalia with the lobe of bursa giving off the
ductus seminalis partially sclerotized, otherwise mem-
branous; ductus bursae flattened, sclerotized through-
out, concavely bent at middle, the sclerotization termi-
nating in a sinuate, thickened, narrow, liplike band
along the lower margin of the genital opening.
TYPE LOCALITY: Winnipeg, Manitoba (type in
USNM).
Foop prant: Unknown.
Distrisution: Manitoba, Aweme (June, July), Win-
nipeg (June).
Genus Pyla, Species 298-306: P. scintillans to
P. viridisuffusella
[Ground color of forewing bronzy brown.]
298. Pyla scintillans (Grote)
Figures 29, 372, 373, 857
Nephopteryzr scintillans Grote, Papilio, vol. 1, p. 18, 1881.
Pyla scintillans (Grote), New check list of North American
moths, p. 55, 1882.—Hulst, Phycitidae of N. Amer., p. 161,
1890.—Ragonot, Monograph, pt. 1, p. 482, 1893.—MeDun-
nough, Check list, No. 6235, 1939.
Pyla feella Dyar, Ins. Insc. Menstr., vol. 9, p. 68, 1921.—Mc-
Dunnough, Check list, No. 6247, 1939. (New synonymy.)
Maxillary palpus of male subsquamous (the scales
short, flattened, forming a small expanded brush).
Forewing dark bronzy brown; the scaling shiny,
metallic; transverse lines absent, indicated only on well
marked specimens by very faint, moderately broad,
blackish brown bands (vestiges of their dark borders) ;
discal and terminal dots obsolete. Alar expanse,
20-26 mm.
Male genitalia having harpe with strong clasper,
developed as a stout outwardly curved hook with an
extended, elongate, bladelike base, the latter more or
less serrate. Considerable individual variation is shown
in the clasper and the shape ofits base. Inone example,
from Inyo County (presumably a variety of scintillans
but possibly a distinct species), the clasper hook is
markedly longer than in the examples figured, and the
bladelike base narrower. Costa of harpe produced at
base into a pointed, stout, very coarsely spined projec-
tion. Aedeagus bifid for less than half its length; one
of the divided elements with a short, thornlike spine
projecting from lateral margin before apex; the other
with 2 or 3 similar spines from lateral margin near apex
(usually 3, rarely 2, a single specimen from El Dorado
County, exhibiting only one). A single pair of ventro-
lateral abdominal hair tufts on eighth segment.
Female genitalia with bursa small, membranous
throughout; ductus bursae very short, expanded abruptly
into a sclerotized cup, its lower surface developed as a
pair of flattened, pointed, elongate-oval blades which
project beyond genital opening. Only trifling individual
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
differences can be distinguished between Dyar’s feella
and females from other California localities.
TYPE LOCALITIES: Summit, Sierra Nevada Mts.,
Calif. (scintillans, in BM); Bullfrog Lake (10,634 ft.),
Sierra Nevada Mts., Calif. (feella, in USNM).
Foop prant: Unknown.
Disrrisution: California, Cisco (Placer County,
July), Deer Park Springs (Lake Tahoe, July), El
Dorado County (July), Inyo County (July), Mineral-
king (Tulare County, July, Aug.), Sierra Nevada Mts.
(Bullfrog Lake and Summit, Aug.), Tuolumne Meadows
(July).
299. Pyla sylphiella Dyar
Fieures 375, 858
Pyla sylphiella Dyar, Ins. Insc. Menstr., vol. 9, p. 68, 1921.—
McDunnough, Check list, No. 6246, 1939.
Maxillary palpus of male subsquamous.
Superficially like scintillans, averaging a trifle darker;
but distinguished only by its genitalia. Alar expanse,
19-25 mm.
Male genitalia with clasper of harpe similar to that
of scintillans; produced enlargement of base of costa
considerably stouter and more coarsely spined. Aede-
agus with only a pair of lateral spines from adedeagus
near its apex (one spine from each of the divided
elements opposite and pointed away from each other).
These differences are slight but appear to be consistent
through long series. Female genitalia with ventral
surface of the cup-shaped portion of ductus bursae
bent into broad, deep, strongly sclerotized folds.
Type tocatity: Mount Rainier, Wash. (type in
USNM).
Foop piant: Unknown.
Distripution: Unirep States: Washington, Mount
Rainier (Aug.), Paradise Valley (Mount Rainier,
Aug.), Sheep Lake (Yakima County, Aug.), Skyline
Ridge (Mount Baker District, Aug.), Slate Peak
(Whatcom County, Aug.). Canapa: British Columbia,
Mount Cheam (Aug.), Mount McLean (Aug.).
The species is very close to scintillans but apparently
distinct. The male genitalia differ only in minor details
and the color and maculation offer little if anything to
separate the two; but the female genitalia are markedly
different and, from the specimens available, sylphiella
appears to have a more northerly distribution.
300. Pyla rainierella Dyar
Fiaures 374, 859
Pyla rainierella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 109,
1904.—McDunnough, Check list, No. 6243, 1939.
Maxillary palpus of male subsquamous.
Moths averaging a trifle smaller than the preceding
species (sylphiella), but certainly distinguished from it
only by genitalia. Alar expanse, 16-20 mm.
Male genitalia with production from base of costa
of harpe considerably smaller and less coarsely spined
than that of either sylphiella or scintillans. Aedeagus
short, the apices of its divided elements bent abruptly
downward as sharp, parallel, spinelike hooks. Female
147
genitalia with the sclerotized portion of the cup-shaped
area of ductus bursae developed laterally as triangulate
plates.
Type tocatiry: Mount Rainier, Wash. (type in
USN).
Foop pirant: Unknown.
Distrisution: Washington, Mount Rainier (Aug.),
Paradise Valley (Mount Rainier, July), Sheep Lake
(Yakima County, Aug.), Slate Peak (Whatcom
County, Aug.), Table Mountain (Aug.).
Like sylphiella, this species is chiefly distinguished by
its female genitalia, the sclerotized area of ductus
bursae at genital opening resembles somewhat that of
Jasciella but differs in shape and is like that of no other
species in the genus. The aedeagus easily separates
the male of fasciella from either scintillans or sylphiella
301. Pyla aeneella Hulst
Ficures 376, 864
Pyla aeneella Hulst, Canadian Ent., vol. 27, p. 55, 1895.—Mc-
Dunnough, Check list, No. 6242, 1939.
Maxillary palpus of male in the form of a semiaigrette
(the hairs short).
Forewing unicolorous, without any trace of dark
transverse shadings or discal spots; brown with a bronzy
green irridescence. Hind wing concolorous with fore-
wing. Alar expanse 23-25 mm.
Male genitalia with costa of harpe at base produced
into a thin, rounded lobe with finely serrate edge;
clasper erect, short, stout, thornlike. Aedeagus short,
broadest at middle, divided to middle; the divided ele-
ments rather broadly flattened and abruptly, asym-
metrically bent at their apices. Female genitalia with
ductus bursae broadly cup-shaped from shortly beyond
its junction with bursa; the ventral surface weakly
sclerotized, granulate, and with slight infoldings on the
lower median area.
Typr Locaity: Colorado (in AMNH, ex Rutgers).
Distrisution: Colorado; Utah, Silver Falls (July),
Stockton (May, June).
A good series of the Utah specimens is in the National
Collection. The genitalia of the females agree in every
detail with those of the Colorado type in the Rutgers
Collection. The metallic iridescence of the forewings
is conspicuous but its greenish tint is very faint.
302. Pyla aeneoviridella Ragonot
Ficures 378, 862
Pyla aeneoviridella Ragonot, N. Amer. Phycitidae, p. 9, 1887;
Monograph, pt. 1, p. 482, 1893.—Hulst, Phycitidae of N.
Amer., p. 161, 1890.—Barnes and McDunnough, Contribu-
tions, vol. 2, p. 222, 1914—McDunnough, Check list, No.
6237, 1939.
Maxillary palpus in the form of a semiaigrette (the
scales somewhat flattened, not so decidedly hairlike as
in aeneella).
Forewing unicolorous bronzy brown, without dark
markings of any kind; the iridescent scaling with a
faint greenish tint and somewhat more strongly con-
centrated at the base of the wing than in the median
148 UNITED STATES NATIONAL MUSEUM BULLETIN 207
and outer areas. The color of both the fore and hind
wings is similar to that of aeneella and the two species
can only be safely distinguished by their genitalia.
Alar expanse, 23-29 mm.
Male genitalia with an angulate, serrate and coarsely
spined projection from costal base of harpe; clasper out-
bent, strongly spined and more or less serrate. Aedea-
gus short, slightly bent at middle and with a single stout,
stubby spine projecting from one side (extent of indi-
vidual variation shown in figs. 378a, b). Female geni-
talia with a weak sclerotization of the lobe of bursa
giving off the ductus seminalis; ductus bursae with
ventral surface of cup-shaped area more strongly
sclerotized than that of aeneella and differently sculp-
tured.
TYPE LocaLity: Evanston, Wyo. (type in Paris Mus.)-
Foop prant: Unknown.
Distrisution: Unitep States: Colorado, Tennessee
Pass (July); Wyoming, Big Horn Mts. (July),
Evanston, Yellowstone Park (July, Aug.); Montana,
Bozeman (July), Glacier Park (June); Oregon, Wallowa
Mts. (Arnold Lake, July); Washington, Olympic Mts.
(Hurricane Ridge, June, July). Canapa: Alberta,
Laggan (July).
The Washington and Oregon specimens are consider-
ably darker than those from the other localities, the
specimens from Oregon having almost black hind wings
and blackish brown forewings.
In his original description and in his Monograph
Ragonot gives “N. Y.’’ as the type locality. This was
a misreading of the label of his type. ‘The correction
was made by Barnes and McDunnough in the reference
cited above.
303. Pyla metalicella Hulst
Figures 377, 863
Pyla metalicella Hulst, Canadian Ent., vol. 27, p. 54, 1895.—
McDunnough, Check list, No. 6236, 1939.
Maxillary palpus of male in the form of a semiaigrette
(the hairs rather short).
Appreciably lighter than aeneoviridella. The fore-
wing a unicolorous light bronzy brown with greenish
yellow iridescence; discal dots more or less distinct,
separated, blackish; no other markings. Average size
larger than that of aeneoviridella. Alar expanse, 25-32
mm.
Male genitalia with a long, somewhat flattened, out-
wardly curved clasper on harpe (similar to that of
wmpostor, but proportionally longer); costa of harpe at
base simple. Aedeagus very shortly divided at apex;
a pair of very short, sharp, straight spines on ventral
surface near apex.
Female genitalia with lobe of bursa giving off ductus
seminalis weakly sclerotized; cuplike area of ductus
bursae, funnel shaped (triangulate), strongly sclerotized
over its entire ventral surface and containing a broad,
centrally located, funnel-shaped fold. Individual vari-
ations in this fold are shown in figs. 863 and 863a.
Type Locauity: Colorado (type in AMNH, ex
Rutgers).
Foop prant: Unknown.
DistriBuTion: Colorado, Silverton (July, Aug.), and
two specimens with only the state locality; Utah, Silver
Lake (July). The Silverton locality is represented in
the National Collection by 17 specimens. Also in the
National Collection is a female from Colorado (Bruce),
labeled ‘“Pyla aeneella Hulst, Type,” another of Hulst’s
pseudotypes and possibly part of his original ‘“‘type”
series of aeneella. ‘The actual type of metalicella is a
male with only the state locality. Its genitalia agree
in every detail with those from Silverton specimens.
304, Pyla fasciella Barnes and McDunnough
Figures 379, 861
Pyla fasciella Barnes and McDunnough, Canadian Ent., vol. 49,
p. 405, 1917.—McDunnough, Check list, No. 6244, 1939.
Maxillary palpus of male minute, squamous.
Forewing blue-black, some paler bluish iridescence
over basal area and (in strong light) a faint, brownish
iridescence in outer area; antemedial line represented
by a narrow oblique black band near, but before middle;
subterminal line a similar curved band well back from
and parallel with termen; the area between the two
black bands darker than remainder of wing, forming a
faint, broad, median, black fascia; discal dots obsolete.
Hind wings very dark brown, shiny. Alar expanse,
21-24 mm.
Male genitalia without clasper on harpe; base of
costa of harpe produced into a knoblike projection,
finely spinose along margin. Aedeagus simple. A
single pair of ventrolateral hair tufts on eighth segment.
Female genitalia similar to those of rainierella except
that the paired plates of ductus bursae at genital open-
ing are narrower and differently shaped.
Typrk tocatity: Mount Shasta, Calif. (type in
USNM).
Foop prant: Unknown.
Known only from northern California. Represented
in the National Collection by the type series from
Mount Shasta, 7,000 ft., July—three males and two
females (not four males and two females as given in the
original description) ; and one male from Bartle, Calif.
(June 14, 1939, Grace H. and John L. Sperry). In
their original description the authors give the expanse
as “24-31 mm.” This is probably a printer’s error,
for the largest specimen before me is a scant 24 mm.
305. Pyla nigricula, new species
Fieure 380
Maxillary palpus of male minute, squamous.
Superficially like fasciella except: Transverse dark
lines of forewing obsolete, only the antemedial black
band very faintly indicated; no contrasted dark median
fascia; the entire median and outer areas a dark
purplish brown. Alar expanse, 26 mm.
Male genitalia with uncus broader and squattier than
that of fasciella. Projection from costal base of harpe
differently shaped, bluntly poimted; clasper developed
as a stout, smooth, curved, pointed hook. Aedeagus
simple.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 149
Typrxocauity: Verdi, Nev. (type in USNM, 61354).
Foop piant: Unknown.
Described from unique male collected by A. H.
Vachell, “June 1 to 10.”” This specimen had been in
our collection under scintillans Grote.
306. Pyla viridisuffusella Barnes and McDunnough
Figures 381, 851
Pyla viridisuffusella Barnes and McDunnough, Canadian Ent.,
Mayan’ p. 406, 1917—McDunnough, Check list, No. 6245,
Maxillary palpus of male in the form of a short
aigrette.
Forewing heavily suffused with light, bronzy green,
irridescent scaling, especially strong over basal area and
in a line indicating the subterminal line; two transverse
blackish bands, an oblique, antemedial one and another
forming an inner border to the subterminal line; discal
dots, when distinguishable, confluent, forming a line
along discocellular vein. Hind wing very dark brown.
Alar expanse, 17-20 mm.
Male genitalia with harpe simple. Aedeagus bifur-
cate to middle, the divided elements asymmetrical (one
longer than the other). Female genitalia with a pair
of strongly sclerotized, convolute bands extending from
posterior end of bursa well into ductus bursae; genital
opening simple.
Tyrer ocauity: Tuolumne Meadows,
County, Calif. (type in USNM).
Foop prant: Unknown.
Distrisution: California, Humphreys Basin (Fresno
County, Aug.), Johnsons Park (Sierra Nevada Mts.),
Kernick Meadows (9,250 ft., July), Mineralking (Tulare
County, July, Aug.), Tuolumne Meadows (July, Aug.).
The most brilliant of the Pyla species. Its genitalia,
both male and female most resemble those of fasciolalis
Hulst.
Tuolumne
Genera 77 and 78: Dioryctria and Oryctometopia
[Venational division D. Forewing with veins 4 and 5 closely
approximate for a short distance from cell (Dioryctria), or connate
or very shortly stalked (Oryctometopia); vein 6 straight or bent
towards base; 10 from the cell. Hind wing with cell less than
one-third the length of wing; discocellular vein curved; veins 4
and 5 stalked for at least half their lengths. Male antenna with
@ shallow sinus or slight incurvation in base of shaft, containing
& row of fine spines or a weak scale tuft. Male genitalia with
transtilla incomplete or absent; harpe with costa strongly scle-
rotized and produced at apex (Dioryctria) or with one or more
short, stout, thornlike spines from lower margin of sacculus.]
77. Genus Dioryctria Zeller
Dioryctria Zeller, Isis von Oken, 1846, p. 632.—Heinemann, Die
Schmetterlinge Deutschlands und der Schweiz, Abt. 2, vol.
1, pt. 2, p. 148, 1865.—Ragonot, Ent. Monthly Mag., vol.
22, pp. 52, 56, 1885; Monograph, pt. 1, p. 187, 1893.—Hulst,
Phycitidae of N, Amer., p. 134, 1890.—Spuler, Die Schmet-
terlinge Europas, vol. 2, p. 213, 1910.—Forbes, Cornell Univ.
Agr. Exp. Station, Mem. 68, p. 619, 1923.—Meyrick, Re-
vised Handbook of British Lepidoptera, p. 383, 1928.—
Bisset, in Pierce and Metcalfe, Genitalia of the British
Pyrales, p. 57, 1938 (notes Ragonot’s fixation (1885) of
type of genus).—Janse, Journ. Ent. Soc. South Africa, vol.
4, p. 161, 1941. (Type of genus: Tinea abietella Denis and
Schiffermiiller.)
Pinipestis Grote, Canadian Ent. vol. 10, p. 19, 1878; Bull. U. 8S.
Geol. Geogr. Surv. Terr., vol. 4, p. 699, 1878.—Hulst,
Phycitidae of N. Amer., p. 136, 1890. (Type of genus:
Nephopteryx zimmermani Grote.)
Tongue well developed. Antenna finely pubescent
or very shortly ciliate, rarely unipectinate (in males of
some Old World species, pineae Staudinger, mendacella
Standinger); on all males a shallow sinus in base of
shaft, containing a short row of minute black, thornlike
spines, more or less concealed by rough scaling. Labial
palpus upturned, reaching to or a trifle above vertex;
second segment grooved on inner side; third segment
short (less than one-third the length of second), acumi-
nate. Maxillary palpus of male small and squamous or
(rarely) in the form of an aigrette. Forewing smooth
or with two or more tufts or raised (ruffed) scales; 11
veins; vein 2 from before lower outer angle of cell; 3 from
the angle, slightly nearer to 4 than to 2 at base; 4 and 5
from the cell, approximate at and for about one-fourth
their distance from cell; 6 bent towards base and from
close to upper angle of cell (in type species, straight and
from well below the angle on some specimens); 10 from
the cell, approximate to the stalk of 8-9 for some dis-
tance beyond cell; 8 and 9 long stalked; male without
costal fold. Hind wing with vein 2 from before lower
outer angle of cell; 3 from the angle, long (almost as long
as 2); 4 and 5 stalked for half or slightly more than half
their lengths; 7 and 8 closely approximate for less than
half their lengths beyond cell; rarely shortly anasto-
mosed; cell Jess than one-third the length of wing; dis-
cocellular vein curved, outwardly extended at lower
angle. Eighth abdominal segment of male with com-
pound ventral scale tufts.
Male genitalia decidedly elongated (least so in gulo-
sella). Uncus broad, stout, its terminal margin more or
less broadly rounded. Apical process of gnathos a
short, stout hook. ‘Transtilla incomplete or absent; its
lateral elements, when distinguishable, slender and
usually attached to costal base of harpe. Harpe with
costal area broadly sclerotized and produced at apex;
cucullus narrowly elongate, bluntly pointed or very
narrowly rounded at apex; sacculus short, simple;
clasper present, erect, usually finely haired at or near
apex, but not bearing strong spines, thorns, or serra-
tions. Anellus with well-developed lateral lobes. Ae-
deagus long, moderately stout; penis with strong cor-
nuti, consisting of numerous straight, slender spines (as
long or nearly as long as width of aedeagus) and usually
one or more longer, stouter spines placed back of them
on the penis. Vinculum stout; longer than broad
(frequently considerably elongated), gradually tapering
to a moderately broad, rounded, or abruptly angled
terminal margin.
Female genitalia with well-developed signa, consisting
of two or three clusters of strong, slender spines, their
bases in each of the clusters more or less fused into
sclerotized plates; the clusters in end of bursa near
150 UNITED STATES NATIONAL MUSEUM BULLETIN 207
ductus bursae and one of them frequently in or extend-
ing into the ductus; ductus bursae flattened, strongly
sclerotized over most of its length, the sclerotization
more or less longitudinally wrinkled and terminating just
before the simple genital opening; ductus seminalis from
bursa near the junction of bursa and ductus bursae.
Dioryctria is one of the most, if not the most, distinct
and sharply defined genus in the Phycitidae; and is so
despite the variations in structure exhibited by its male
antennae, male maxillary palpi, and its wing scaling.
Its male and female genitalia have a characteristic
habitus, difficult to describe, but easily observed in slides
or figures. There is also a maculation character of the
forewing common to nearly all the species; the usual
black dots at end of cell are absent, being replaced by a
white spot or line on the discocellular vem. The only
North American species without such a marking is
clarioralis, where a considerable area about the end of
cell is clouded with a dark suffusion. The raised-scale
character upon which Grote distinguished his genus
Pinipestis from Dioryciria consists of a mere rufiing of
the scales on two or three spots on the forewing. It is
at best an elusive character, subject to opinion as much
as to observation, and in some of the forms(zimmermani,
cambiicola) the tufts may be either raised or flattened on
unspread specimens. Naturally, on spread specimens
they are usually flattened, whatever was their condition
in nature. Ragonot was perfectly justified (1893) in
relegating Pinipestis to synonymy.
As here defined the genus is of world-wide distribution
in the Northern Hemisphere and all the species whose
larval habits are known are borers in conifers. Most of
the American species are represented in the National
Collection by numerous specimens reared in connection
with the forest-insect investigations of the U.S. Bureau
of Entomology and Plant Quarantine.
307. Dioryctria abietella (Denis and Schiffermiiller)
Fieures 51, 382, 865
Tinea abietella Denis and Schiffermiiller, Systematische Ver-
zeichniss der Schmetterlinge der Wienergegend . . . , p. 138,
1776.—Fabricius, Mantissa insectorum . . . , vol. 2, p. 245,
1787; Entomologica systematica . . . , vol. 3, pt. 2, p. 302,
1793.
Tinea decuriella Hibner, Sammlung europaischer Schmetterlinge,
p- 35 and Lepidoptera 8, Tineae 2, pl. 11, fig. 74, 1796.
Phycis abietella (Denis and Schiffermiiller) Zincken, in Germar
and Zincken, Mag. der Ent., vol. 3, p. 160, 1818.—Treitschke,
Die Schmetterlinge von Europa, vol. 9, p. 177, 1832.—
Ratzeburg, Die Forst-Insecten . . . , vol. 2, p. 244, pl. 15,
fig. 2, 1840.
Dioryctria abietella (Denis and Schiffermiiller) Zeller, Isis von
Oken, p. 736, 1846——Heinemann, Die Schmetterlinge
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 148,
1865.—Ragonot, Ent. Monthly Mag., vol. 22, p. 52, 1885;
Monograph, pt. 1, p. 198, 1893.—Hulst, Phycitidae of N.
Amer., p. 135, 1890.—Spuler, Die Schmetterlinge Europas,
vol. 2, p. 213, 1910.—Joannis, Ann. Soc. Ent. France, vol.
85, p. 259, 1916; Bull. de la Station de Recherches forestiérs
du Nord de 1’ Afrique, vol. 1, p. 192, 1921.—Forbes, Cornell
Mem. 68, p. 621, 1923.—Meyrick, Revised handbook of
British Lepidoptera, p. 384, 1928.—Pierce and Metcalfe,
Genitalia of the British Pyrales, p. 2, pl. 1, 1938.—Keen,
U.S. Dep. Agr. Mise. Publ. 273, p. 39, 19838.—McDunnough,
Check list, No. 6129, 1939.—Janse, Journ. Ent. Soc. South
Africa, vol. 4, p. 161, 1941—MacKay, Canadian Ent., vol.
75, p. 94, 1943.—Craighead, U. S. Dep. Agr. Misc. Publ.
657, p. 451, 1950.
Pinipestis abietivorella Grote, Bull. U. S. Geogr. Geol. Surv. Terr.,
vol. 4, p. 701, 1878.
Pinipestis reniculella Packard (not Grote), U. 8S. Dep. Agr. Ent.
Bull. 13, p. 21, 1887; U.S. Dep. Agr. Fifth Rep. Ent. Comm.,
p. 854, 1890.
Myelois elegantella Hulst, Canadian Ent., vol. 24, p. 59, 1892.
Maxillary palpus of male squamous.
Forewing smooth; bluish gray, dusted with white, the
white dusting of variable intensity, when pronounced,
more or less concentrated in basal area, along the outer
border of antemedial line, the inner border of subter-
minal line and, weakly, along terminal margin; trans-
verse lines white, distinct, narrow; antemedial line
oblique, inwardly angled at cell and vein 1b, preceded
on costa by a blackish shade and bordered outwardly
by a black line, and usually preceded by a pale patch
on inner margin, this often no more than a smear of
olivaceous white scales and never so conspicuous or
well contrasted as in reniculella; subterminal line
sinuate-angulate, preceded and followed by blackish
bordering lines; discal mark a white, lunate spot; a fine
black line along terminal margin. Hind wing dusky
white, darkened slightly towards outer margin and on
the veins. Alar expanse, 20-30 mm.
Male genitalia with uncus subtriangulate and rather
narrow in normal position; a short, slight incurvation
of the lateral margins near base, and the lateral margins
themselves infolded. (When uncus is flattened in prep-
arations and the lateral folds pushed out, the uncus
appears as in fig. 382, but never takes the form of the
flattened unci of the species which have a longer incurva-
tion of the lateral margins, such as zimmermani). Harpe
with one or more spines projecting from the terminal
margin of the sclerotized costal area below its apex
(there is considerable individual variation in this feature,
a few examples of which, from small American speci-
mens, are shown in figs. 882c-e). Penis armed with a
single stout spine behind anterior spine cluster.
Female genitalia chiefly distinguished by a longitu-
dinal fissure on the ventral surface of the sclerotized
portion of ductus bursae, variations of which are shown
in figures 865 and 865a. The females of sysstratiotes
from Guatemala also show traces of such a fissure but
this species is only doubtfully distinct from abietella.
TYPE LOCALITIES: Austria (abietella, location un-
known); Germany (decuriella, type lost); Amherst,
Mass. (abietivorella, in BM); Seattle, Wash. (elegantella,
in AMNH, ex Rutgers). ;
Foop PLants: Pine, spruce, fir; in the spruces and firs
(Abies, Pseudotsugha, etc.) chiefly in the cones. The
favored host seems to be Pinus of which it attacks all
species. The larvae exhibit a variety of habits. They
are both primary and secondary. ‘They bore into new
and otherwise uninfested terminals and into terminals
that have been attacked by Rhyacionia buoliana or the
white pine weevil (Pissodes strobi.) They attack both
healthy and diseased cones. They bore into and feed
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 151
on the cambium of smooth bark on the trunks and
branches; and also feed in the galls on any part of the
tree.
Disrrisution: Apparently throughout the range of
the genus Pinus in the Northern Hemisphere. Ameri-
can records from specimens (moths) before me as
follows: Unirep States: Maine, Orono (Aug.); Massa-
chusetts, Amherst, Framingham (Oct.), Martha’s Vine-
yard (Sept.), Pepperell (Aug.); Connecticut, Bradford,
Lyme (Aug.), New Haven (June); New York, Long
Island (Garden City, Oct., Great Neck, July), Warrens-
burg (Sept.); New Jersey, Lakehurst (May); District
of Columbia, Washington (July, Aug.); Florida, Alton
(June), Eustis (June, July), Gainesville (June), Orlando
(June), ‘So. Florida” (June, July, Aug.); Jdlinois, Dun-
dee; Nebraska, Halsey (Apr., June, Aug.); Montana,
Dillon (July), Elliston, Evaro (Mar.); Colorado, Glen-
wood Springs; Arizona, Prescott (July); California,
Berkeley, Patrick’s Creek (Sept.), Sacramento, Shasta
Retreat (July); Oregon, Ashland (July, Aug., Sept.,
Oct.), Salem (Aug.), Colestin (June), Silver Lake (Aug.),
Sprague River (July); Washington, Hoquiam, Pullman,
Rock Lake (Whitman County, Sept.); Seattle. Can-
apa: Labrador, Dublin Shore (Lunenburg County) ;
Quebec, Montreal (June); Saskatchewan, Lutherland
(June, Aug., Sept.); British Columbia, Kaslo (June).
GuateMaLa: A series of males and females in the U. S.
National Museum, reared from pine cones, May 1927,
by J. G. Salas and labeled simply “Guatemala, C. A.’
The species is of considerable economic importance,
especially to young pine trees in our Western States,
and particularly in reforestation areas. It has an
extensive literature in the Old World. I have listed
here only the more important references and have
omitted purely European synonyms. For additional
references the reader is referred to Ragonot (Mono-
graph, p. 198), Hulst (Phycitidae of N. Amer., 1890),
the Journal of Economic Entomology, and the Review
of Applied Entomology. The most satisfactory infor-
mation on life history and larval habits will be found in
the MacKay (1943) and Craighead (1950) papers.
308. Dioryctria sysstratiotes Dyar
Ficure 866
Dioryctria sysstratiotes Dyar, Ins. Insc. Menstr., vol. 7, p. 48,
Forewing smooth, similar to that of abietella except:
A pale brownish suffusion in median area forming a
rather large patch below discal spot; a similar brownish
shade outwardly bordering subterminal line; the patch
preceding antemedial line on inner margin, more dis-
tinct, larger, pale olivaceous brown. Hind wing some-
what darker, translucent smoky white with a very faint
brownish tint towards outer margin.
Alar expanse, 23-28 mm.
Female genitalia as in abietella except no (or only a
faint trace of) median ventral cleft in ductus bursae.
TyPE Locauity: Cayuga, Guatemala (type in
USNM).
Foop PLant: presumably Pinus. No rearing records.
Disrripution: GuaTEMALA: Cayuga (June), Chejel
(June), Purulhé (July).
Known only from females. Doubtfully distinct from
abietella except as arace or colorform. Its exact status
will have to await discovery of a male.
309. Dioryctria reniculella (Grote)
Figures 383, 867
Pinipestis reniculella Grote, North Amer. Ent., vol. 1, p. 67,
Dioryctria reniculella (Grote), Ragonot, Monograph, pt. 1, p. 200
1893.—Forbes, Cornell Mem. 68, p. 620, 1923.—McDun™
nough, Check list, No. 6131, 1939.—Brown, Canada Dep-
Agr. Publ. 712, Techn. Bull. 31, p. 13, 1941.—MacKay, Cana-
dian Ent., vol. 75, p. 94, 1943.—Craighead, U. 8. Dep.
Agr. Misc. Publ. 657, p. 451, 1950.
Maxillary palpus of male squamous.
Forewing smooth; in color and maculation similar to
that of abietella except: Ground color pale brownish
gray; the transverse lines and discal spot more sharply
contrasted, more distinctly white; usually a rather large
olivaceous patch on inner margin preceding the ante-
medial line; hind wing darker, pale smoky fuscous.
Alar expanse, 22-26 mm.
Male genitalia with uncus tonguelike; more elongate
and narrower than that of any other American species;
its terminal margin very narrowly rounded; no incur-
vation of the lateral margins (its shape not appreciably
altered by flattening in slide preparations). A very
slight spur from the lower outer angle of the sclerotized
costal area of harpe, but no other spines from below
apex of costa. Penis armed only with anterior spine
cluster; no single stout spine on penis behind the an-
terior cluster.
Female genitalia with only one strong spine cluster,
that at junction of bursa and ductus bursae; girdle of
spines in bursa before junction with ductus, weak,
broken, and the spines themselves greatly reduced.
Type Locauity: New York (type in BM).
Foop Puants: Various spruces; rarely in balsam fir
and tamarack. Reported as occasional in jack pine,
but such records are doubtful. The larvae feed in
terminals and cones and to a lesser extent upon the
foliage of the terminals.
Distrisution: Unirep Srates: Maine, Blue Hill
(July), Sebee Lake (July); Connecticut, East River
(July), New Haven (June, July); New York; Illinois,
Putnam County (July) ; Michigan, East Lansing (Aug.) ;
Colorado, Estes Park (July); California, Fallen Leaf
Lake (Aug.), Mount Lowe (July); Washington, Belling-
ham (Aug.), Hoquiam, Kamiack Butte (Aug.), Pullman
(July, Aug.). Canapa: Nova Scotia, Cape Breton
(Aug.); Quebec, St. Therese Isl. (St. John’s County,
July); Ontario, Westree; Saskatchewan (June); British
Columbia, Seton Lake (June, July), Victoria (July).
The species is easily distinguished by its genitalia.
In the past it has been frequently confused with
abietella and until 1893 was treated by Hulst and
Ragonot as a synonym of the latter. The larvae of the
two species often occur together in spruce cones, so
there is no safe way to separate them on their larval
152 UNITED STATES NATIONAL MUSEUM BULLETIN 207
habits. However, reniculella is primarily a spruce
feeder, while abietella shows a marked preference for
the pines. Both species are borers, as are all the
Dioryciria species. It is unfortunate that Miss Mac-
Kay in her otherwise excellent paper (1943) should refer
to reniculella as ““The spruce foliage worm.” It does
feed to some extent upon the foliage of terminal shoots;
but likewise, and more frequently within the terminals
themselves and in the cones, and in economic literature
is quite properly referred to as ‘“‘the spruce cone moth.”
It is a strictly American species limited in distribution,
apparently, to the northern United States and Canada.
310. Dioryctria ponderosae Dyar
Figures 384, 868
Dioryciria ponderosae Dyar, Ins. Insc. Menstr., vol. 2, p. 2,
1914.—McDunnough, Check list, No. 6130, 1939.
Maxillary palpus of male squamous.
Forewing smooth; like that of abietella except: Black-
ish markings more strongly contrasted; a broad black
band inwardly bordering the antemedial line, a similar
band on some specimens of abietella but not so broad
nor so well contrasted; subterminal line outwardly
angled at middle; the white transverse lines and discal
spot well contrasted. Hind wing white, clouded with
smoky fuscous towards outer margin; the veins slightly
darkened. Alar expanse, 27-30 mm.
Male genitalia with uncus considerably shorter and
broader than that of either abietella or reniculella; its
terminal margin broadly rounded; lateral margins in-
curved near its base. Harpe with apex of sclerotized
costa produced into a blunt, curved hook, no spine from
lower outer angle of the sclerotized costal area; clasper
reduced. Female genitalia distinguished by its short
ductus bursae:
TypE Locatity: Lamedeer, Mont. (type in USNM).
Foop prant: Pinus ponderosa. Larva a borer in the
cambium of the bark.
Distrisution: Montana, Lamedeer (June), Colorado,
Glenwood Springs (June, July, Aug.); California,
American River (May).
A distinct species known only from a half-dozen
specimens from the above localities. It belongs defi-
nitely with the smooth-winged Dioryctria, although a
few roughened scales can be distinguished in the black
inner borders of the antemedial line on a couple of the
specimens. Its genitalia separate it easily from all
other smooth-winged species. The name has been
misapplied to a color variety of zimmermani. This
misidentification is discussed under the latter name.
The Missoula, Mont., female (Hopkins U.S. No. 11508)
mentioned by Dyar in his description of ponderosae
belongs to this variety of zimmermani.
311. Dioryctria majorella Dyar
Fiaures 385, 871
Dioryctria majorella Dyar, Ins. Insc. Menstr., vol, 7, p. 43, 1919.
Dioryctria muellerana Dyar, Ins. Insc. Menstr., vol. 7, p. 85, 1919
(new synonymy).
Maxillary palpus of male squamous.
Forewing smooth, similar in color and markings to
that of sysstratiotes except: Somewhat more suffused;
the whitish markings fainter and a duller, more sordid
white; the pale discal spot obscure; the blackish inner
border of the subterminal line and the blackish band
preceding the antemedial line broader. Hind wing
smoky white, the veins faintly darkened. Alar expanse,
28-33 mm.
Male genitalia distinguished by greatly increased
width of the sclerotized costal area of harpe; a short
spur projects from the outer margin of this sclerotized
area just below its apex. D. erythropasa has a similar
harpe; but differs markedly in other details of the geni-
talia—differently shaped uncus, narrower clasper, and
different spining of penis.
Female genitalia differ from those of sysstratiotes only
in insignificant details. The female genitalia of both
species differ from those of abietella chiefly in the greater
broadening and thickening of the membrane of ductus
bursae near its junction with bursa copulatrix.
TYPE Locauities: Jalapa, México (majorella, 9, in
USNM); Zacualpén, México (muellerana, o, in USNM).
Foop pPLant: Pine.
Known only from Dyar’s two types and a pair (o' and
9) from British Honduras. The Honduran specimens
are smaller than the Mexican types and in wretched
condition, but their genitalia are a perfect match in all
details. In his description of muellerana Dyar sug-
gested that it might be the male of majorella. This
synonymy is proven by the Honduran examples. They
were reared from cones of Pinus caribaea.
312. Dioryctria disclusa Heinrich
Fiaure 372
Dioryctria disclusa Heinrich, im Farrier and Tauber, Iowa State
Coll. Journ. Sci, vol. 27, p. 495, 1953.
Maxillary palpus of male squamous.
Forewing smooth; basal area to antemedial line
orange yellow; area beyond brownish red, more or less
shaded or suffused with yellowish orange (on some
specimens the ground color of the entire wing yellowish
orange), usually the red shade most conspicuous in the
area between subterminal line and termen; transverse
lines narrow, white; a white streak along lower margin
of cell between the transverse lines; antemedial line
faint, oblique, nearly straight ; subterminal line stronger,
set well out, rather near terminal margin, outwardly
angulate between veins 6 and 1b; discal mark (when dis-
tinguishable) a white line along discocellular vein; some
very short white dashes on terminal margin; cilia smoky
white. Hind wing smoky white to pale smoky gray, the
paler examples showing a very faint ocherous tint;
veins slightly darkened; cilia whitish. Alar expanse,
24-29 mm.
Male genitalia like those of auranticella. Female
genitalia essentially like those of auranticella. The
differences shown in the figures for the two species
are the extremes and represent, at most, individual
variations.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 153
Typrr Locauity: Tryon, N.C. (type in USNM).
Foop piants: Pinus spp. Larvae feeding in the
cones.
DistrisuTion: Massachusetts, Martha’s Vineyard
(June) ; New Jersey, Lakehurst (July) ; District of Colum-
bia; West Virginia, Roosevelt (June), North Carolina,
Raleigh (June), Tryon (June); Jowa, Ames (June).
Examples of this species have been in the National
Collection as auranticella and it was on the basis of this
misapplication of Grote’s name that Dyar described his
zanthaenobares. The true auranticella is strictly a
western species, while the distribution of disclusa, as
far as I know, is limited to the eastern and central areas
of the United States.
The paper by Farrier and Tauber gives all the infor-
mation known on the life history, food habits, and
behavior of the insect.
313. Dioryctria auranticella (Grote)
Ficures 386, 873
Nephopteryz auranticella Grote, Ann. Mag. Nat. Hist., ser. 5, vol.
11, p. 57, 1883; Trans. Kansas Acad. Sci., vol. 8, p. 57, 1883.
Dioryctria miniatella Ragonot, N. Amer. Phycitidae, p. 4, 1887.—
Hulst, Ent. Amer., vol. 5, p. 156, 1889.
Dioryctria auranticella (Grote) Hulst, Phycitidae of N. Amer., p.
134, 1890.—Ragonot, Monograph, pt. 1, p. 194, 1893.
McDunnough, Check list, No. 6126, 1939.
Dioryctria zanthaenobares Dyar, Proc. Ent. Soc. Washington, vol.
13, p. 81, 1911.—Keen, U. S. Dep. Agr. Misc. Publ. 273, p.
38, 1938.—McDunnough, Check list, No. 6127, 1939.—
Heinrich, in Farrier and Tauber, Iowa State Coll. Journ.
Sci., vol. 27, p. 495, 1953.
Maxillary palpus of the male in the form of an
aigrette.
Forewing smooth; color and markings similar to those
of disclusa, but on the average somewhat paler, the
ground color ranging from yellowish orange to brownish
red; on the darker suffused, reddish examples the yel-
lowish color of the basal area is less contrasted than in
ordinary disclusa. Very pale examples of the two
species cannot be distinguished superficially. In size
auranticella averages larger. Alar expanse, 27-33 mm.
Male genitalia with apex of cucullus of harpe extend-
ing beyond apex of the sclerotized costa. Female
genitalia exhibiting only slight and comparative differ-
ences from those of dasclusa.
Tyrer Locaities: New Mexico (auranticella, in Univ.
Kansas); Arizona (miniatella, in Paris Mus.); Kaslo,
British Columbia (zanthaenobares, in USNM).
Foop piants: Pinus spp. Larvae feed chiefly in the
cones, sometimes in the twigs. The favored host seems
to be Pinus ponderosa.
DistrisutTion: Unitsep Strats: New Merico, state
locality only; Arizona, Mohave County (July); Colo-
rado, Glenwood Springs (Aug.), Rocky Mountain Na-
tional Park (July); Utah, Eureka (July, Aug.); Zdaho,
Coeur d’Alene (July); Montana, Bitterroot River
(July), Camas (July); California, Gasquets (May),
Pasadena, Warner Mts. (July); Oregon, Monumental
Pass (Aug.), Silver Lake (Aug.); Washington, Pullman
(July), Rock Lake (June, July), Seattle; Nebraska,
300329—56——11
Halsey (Aug.), Valentine (July, Aug.). CaNnapa:
British Columbia; Kaslo (Aug.), Trout Creek (Ibapah
Mts.). The Nebraska records, our most easterly, are
from examples reared from Pinus ponderosa in refor-
ested areas. The insect was probably introduced there
on western nursery stock.
The species is represented by a large series of speci-
mens in the National Collection, a majority of them
reared from cones in connection with the forest insect
investigations of the U. S. Bureau of Entomology and
Plant Quarantine. The type of auranticella in the Snow
Collection at the University of Kansas is a pale female
in rather poor condition. Three other similar females,
in better condition are in the Rutger’s College Collec-
tion. All of these are labeled simply “(New Mexico,
Snow, Coll.” Females of auranticella are not separable
from those of disclusa, especially the pale, yellowish
examples, except by their locality labels. On average
specimens the white streak along the lower margin of
the cell of forewing is shorter in auranticella, not reach-
ing beyond the outer angle of the cell; but this feature is
not constant in either species. Between males there
is never any need for confusion; for auranticella is the
only known American species with aigrettelike maxillary
palpi.
A similar maxillary palpus occurs in Dioryctria laurata
(Heinrich) from Japan (described as a Salebria in Proc.
Ent. Soc. Washington, vol. 30, p. 61, 1928). The unique
male type has the labial palpi erect and appressed
close to the face and a strong scale tuft in the sinus of
the antennal shaft. Hence the original reference to
Salebria. It is probably a snyonym of Dvioryctria
pryert Ragonot (Monograph, pt. 1, p. 194, 1923),
described from a single female from the Holland Col-
lection, now in the Carnegie Museum, Pittsburgh. I
have never seen this specimen.
314. Dioryctria erythropasa (Dyar)
Figures 387, 869
Pinipestis erythropasa Dyar, Ins. Inse. Menstr., vol. 2, p. 112,
1914.
Dioryciria erythropasa (Dyar) Barnes and McDunnough, Check
list of the Lepidoptera of Boreal America, No. 5564, 1917.—
McDunnough, Check list, No. 6128, 1939.
Maxillary palpus of male squamous.
Forewing with some roughened (raised) scale tufts in
median area (one in lower fold and another in cell just
beyond antemedial line and a slight roughening of the
white scales of the discal spot), otherwise smooth;
ground color red-brown of a somewhat darker, more
rosy shade and lacking any of the orange suffusion com-
mon to auranticella and disclusa; more or less dusted
with white in median area, the white concentrated into
a broad patch extending from inner margin to top of cell
just beyond antemedial line; the transverse lines thin,
white; antemedial line oblique, irregularly and very
weakly dentate; subterminal line nearly vertical, slightly
denticulate, bordered inwardly by a dark red-brown
line; a similar dark line forms an outer border to the
antemedial line; discal spot a slightly enlarged, lunate,
154 UNITED STATES NATIONAL MUSEUM BULLETIN 207
white line on discocellular vein; outer area beyond sub-
terminal line red-brown, terminal dots confluent, form-
ing a more or less continuous blackish line along termen;
cilia reddish brown. Hind wing smoky white; the veins
darkened; a very narrow dark shade along termen; cilia
whitish, cut by a dark subbasal line. Alar expanse,
23-28 mm.
Male genitalia of the majorella type but with uncus
short, broad, its terminal margin angulate. Harpe with
slender, digitate clasper. Vinculum narrower, more
gradually tapered. Penis with numerous anterior
spines, but without the usual enlarged posterior cornu-
tus. Female genitalia with bursa copulatrix greatly
reduced, much shorter than ductus bursae.
Tyrr LocaLity: Chiricahua National Forest, Ariz.
(type in USNM).
Foop puant; Pinus chihuahuana. Larvae feeding
in the cones.
Distripution: Arizona, Chiricahua National For-
est (May); Redington (Aug.).
This species, with pygmaeella, forms a connecting link
between the smooth-winged Dioryctria species and those
with distinctly roughened scales formerly referred to
Pinipestis.
315. Dioryctria horneana (Dyar)
Fieure 874
Pinipestis horneana Dyar, Ins. Inse. Menstr., vol. 7, p. 43, 1919.
Maxillary palpus of male squamous.
Forewing bright red-brown shaded with white, the
ground color brighter and more on the red shade than
that of any of the foregoing species; a rather broad,
diffused, white shade preceding the antemedial line
and two rather large, confluent patches following it, one
in the cell and another in lower fold; a strong broad
oblique white shade extending from inner margin near
subterminal line to costal beginning of that line and
fusing with and more or less obscuring it; the scales of
these white areas as well as those of the white discal
spot decidedly roughened; a narrow band of appressed
white scales along terminal margin; antemedial line
narrow, white, nearly vertical, notched above and below
its middle, followed on basal half by a faint, narrow,
gray outer bordering line; subterminal white line rather
close to outer margin, vertical with a slight bulge at
middle; terminal black dots narrow weak, confluent.
Hind wing yellowish white with a very faint smoky tint;
veins but slightly darkened; a fine, pale brown line along
termen. ‘Top of head and collar of thorax red-brown;
remainder of thorax whitish. Alar expanse, 25-28 mm.
Genitalia similar to those of zimmermant.
Type tocauiry: Herradura, Pinar del Rio, Cuba
(type in USNM).
Foop piant: Pinus sp.
Known only from the female type and a male from
thesame rearing. Dyar in his original description gives
Santiago de las Vegas as the type locality, but was evi-
dently in error. Dr. S. C. Bruner, Chief of the De-
partmenta Agronémica of Cuba, has given us the cor-
rect locality. The species is easily distinguished from
others of the zimmermani complex by the bright reddish
ground color of its forewings and from the other red-
winged American species by its strongly tufted fore-
wings.
The labial palpi of the male are more closely appressed.
to the face than those of most Dioryctria species.
316. Dioryctria pygmaeella Ragonot
Ficures 388, 877
Dioryciria pygmaeella Ragonot, N. Amer. Phycitidae, p. 5, 1887;
Monograph, pt. 1, p. 192, 1893.
Pinipestis pygmaeella (Ragonot) Hulst, Phycitidae of N. Amer.,
p. 136, 1890.—Forbes, Cornell Mem. 68, p. 620, 1923.—
McDunnough, Check list, No. 6136, 1939.
Maxillary palpus of male squamous.
Forewing smooth except for a slight roughening of
the white scales of the transverse lines and discal spot;
purplish gray with irregular patches of dull dark red
and a more or less extended white dusting; the red
patches consisting of a rather broad band preceding the
antemedial line, a similar band from middle of inner
margin to cell and a rather broad shade outwardly
bordering the subterminal line; the white dusting con-
centrated over the median areas not occupied by the
red median band, in subbasal area just behind the red
band, and along termen following the red subterminal
shade; transverse lines thin, whitish, rather faint (under
magnification, the scales silvery and somewhat rough-
ened); antemedial line oblique, notched at vein 1b,
bordered outwardly by a black line which expands at
costa into a black patch; subterminal line sinuate-
angulate with a broad, black, inner, bordering line;
terminal dots fused into a fine black line along termen;
white discal spot an oblique lunule; a fine black line
along the lower half of the inner margin of the red band
preceding antemedial line (but not a raised-scale ridge
as stated by Ragonot); also some dusting of black scales
on the red of extreme basal area. Hind wing pale smoky
gray; the veins darkened and a narrow dark shade
along termen. Alar expanse, 15-21 mm.
Male genitalia with a cluster of long strong posterior
spines and one or two weak anterior spines on penis.
Female genitalia with the spine clusters in bursa con-
sisting of narrow, straight bands of slender spines;
ductus bursae sclerotized from junction with bursa
almost to genital opening.
Type Locatity: Florida (type in Paris Mus.).
Foop pLant: Taxodium distichum. Larvae in the
cones. ‘This record from reared series from Maryland
(Heinrich, 1920) and Virginia (Busck, 1927) in the U.S.
National Museum. .
Distrisution: Florida, Winter Park (July); Virginia,
Cape Henry (Aug.); Maryland, Pokomoke (Aug., Sept.).
Probably over the range of its host plant.
‘A distinct, easily recognized species, intermediate be-
tween the smooth-winged species and. mbGEe with definite
meee Seale TCE !
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 155
317. Dioryctria zimmermani (Grote)
Ficures 389, 875, 878
Nephopteryx (Dioryctria) zimmermani Grote, Canadian Ent.,
vol. 9, p. 163, 1877.
Nephopteryx (Pinipestis) zimmermani (Grote), Canadian Ent.,
vol. 10, p. 19, 1878.—Packard, U. S. Dep. Agr. Fifth Rep.
Ent. Comm., p. 73, 1890.
Pinipestis zimmermani (Grote), Bull. U. S. Geol. Geogr. Surv.
Terr., vol. 4, p. 699, 1878; op. cit., vol. 6, p. 589, 1882; N.
Amer. Ent., vol. 1, p. 11, pl. 2, fig. 10, 1879.
Nephopteryx zimmermani (Grote) Kellicott, Canadian Ent., vol.
11, p. 114, 1879; Ent. Amer., vol. 1, p. 173, 1885.
Dioryctria zimmermanni (Grote) Ragonot, Ent. Amer., vol. 5,
p. 114, 1889; Monograph, pt. 1, p. 190, 1893 (emended
spelling of specific name).—Forbes, Cornell Mem. 68, p. 620,
1923.
Pinipestis zimmermanni (Grote) Hulst, Phycitidae of N. Amer.,
p. 137, 1890.—McDunnough, Check list, No. 6133, 1939.
Salebria delectella Hulst, Canadian Ent., vol. 27, p. 57, 1895.—
Ragonot, Monograph, pt. 2, p. 550, 1901. (New synonymy.)
Dioryctria delectella (Hulst) Dyar, Proc. Ent. Soc. Washington,
vol. 6, p. 227, 1904.
Retinia austriana Cosens, Canadian Ent., vol. 38, p. 362, 1906.—
Busck, Journ. New York Ent. Soc., vol. 15, p. 236, 1907.
Pinipestis delectella (Hulst) Barnes and McDunnough, Check
list of the Lepidoptera of Boreal America, No. 5570, 1916.—
MeDunnough, Check list, No. 6134, 1939.
Dioryctria ponderosae Heinrich (not Dyar), in Keen, U. S. Dep.
Agr. Misc. Pub. 273, p. 38, 1938.
Dioryctria zimmermani (Grote) Craighead, U. 8S. Dep. Agr.
Misc. Publ. 657, p. 452, 1950.
Maxillary palpus of male squamous.
Forewing with a ridge of raised (roughened) scales
preceding and one following the antemedial line, some
rough scaling of the discal spots and on some specimens
in the outer median area above inner margin and im-
mediately before subterminal line, the raised scaling
somewhat variable and nowhere reaching to costa,
easily and frequently flattened in the spreading and
setting of specimens; color variable, blackish gray with
a rather broad, but faint, whitish dusting before the
subterminal line and, on some specimens but to a lesser
extent, immediately following the antemedial line and
narrowly and faintly along the inner margin of the
subbasal scale ridge; basal and terminal areas normally
shaded with red, the extent and tint of the shading
extremely variable and on some specimens almost
obliterated or confined to dull patches of an oblivace-
ous hue; when strongly accented, extended over base
and onto the collar of the prothorax, usually most dis-
tinct between subbasal scale ridge and antemedial line;
the raised-scale ridges themselves, black; transverse
lines dull white, always distinguishable but sometimes
faint, bordered inwardly and outwardly by black lines
which broaden into dark wedges on costa; white discal
spot usually distinct; a black line along terminal margin.
Hind wing white more or less shaded with smoky fuscous
along costa and termen, less so on male than female; a
fine dark line along termen; veins only faintly darkened.
Alar expanse, 25-33 mm. '
Male genitalia with uncus but slightly longer than
broad, the lateral margins slightly concave; terminal
margin rounded; when flattened in preparation; as in
figure 390a of cambiicola. .Harpe with costa broadly
sclerotized and terminating at apex in a long curved
pointed hook, a short spine from its lower outer angle;
clasper digitate; cucullus narrow, pointed at apex.
Penis with posterior spine, long, strong, straight, evenly
tapering to a sharp point. Vinculum stout, consider-
ably longer than broad, evenly tapering to roundly
angulate terminal margin.
In the female genitalia the variation in the spining
of bursa shown in the figures is merely individual and
is equaled or exceeded in any series of eastern or western
specimens. Ductus bursae much longer than bursa,
sclerotized for its entire length except for a short dis-
tance from genital opening, the sclerotization ribbon-
like, broadening and bent towards bursa, longitudinally
ribbed on caudal half and terminating caudally in a
produced, bluntly pointed or acutely rounded central
projection. Bursa proportionally small; the spine
clusters closely grouped at its posterior half; the en-
larged lobe giving off the ductus seminalis appreciably
thickened.
Typ Locauities: Buffalo, N. Y. (zimmermani, in
BM; paratype, o, in USNM); Colorado (delectella, in
AMNH, ex Rutgers); Toronto, Ontario, Canada (aus-
triana, in Royal Ontario Mus.).
Foop prant: Pinus spp. Most if not all species of
pine in this country are attacked. The spruce records
given by Packard (1895) have never been verified.
They were probably based upon misidentified larvae.
I doubt very much that zimmermanz feeds on anything
but pine. The larvae bore into the cambium of the
trunk, branches, and twigs, causing considerable dam-
age to the new growth of older trees and sometimes even
killing younger trees (8 inches or less in diameter) by
completely girdling their boles. The place of attack is
usually indicated by a resinous mass of exuded pitch
mixed with frass and larval exuviae.
Distripution: Unirep States: New York, Buffalo,
Coram (Long Island, Aug.), Warrensburg (Aug.); Con-
necticut, Woodstock (Aug.); Rhode Island, Washington
County (June); Massachusetts, Dover (Aug.), Martha’s
Vineyard (Aug.); New Hampshire, Hampton (Sept.);
North Carolina, Tryon (Aug.); Ohio, Akron (July, Aug.),
Lake County (July, Aug.), Mentor (June), Scioto
County (July) ; Ziinois, Oregon (July, Aug.) ; Nebraska,
Ainsworth (Aug.), Halsey (May, June, July, Aug.),
Meadville (Aug.), Norden (Aug.), Wyoming, Wyoming
National Forest (June, July) ; Montana, Banner (July),
Missoula (July); Colorado, only the state locality (fe-
male cotypes of delectella, Bruce, collector) ; New Mezico,
Taos Junction (July); Arizona, Santa Catalina Mts.
(Bear Canyon, July), White Mts. (Aug.); California,
Placerville, San Mateo (June), Ventura; Oregon, Butte
Falls, Coletin; Washington, Friday Harbor (Aug.),
Rock Lake (July). Canapa: Ontario, Toronto.
The foregoing records are from specimens before me,
most of them reared. The range of the species probably
extends over the entire northern areas of the United
States wherever its hosts occur and presumably over
a considerable area in southern Canada.
Hulst’s delectella was described from, Colorado females
156 UNITED STATES NATIONAL MUSEUM BULLETIN 207
which, except for somewhat larger expanse (82 mm.)
than average eastern specimens, are typical zimmermani
in coloration and genitalia. The unfortunate reference
to ponderosae in the Keen paper (1938) was due to my
misidentification of reared specimens from the reforesta-
tion areas of Nebraska. These came to us in good
series but were spread and the raised scales on the fore-
wings had been flattened, and on most of the examples
(as also on some eastern specimens) the characteristic
red shading on basal area of forewing was lacking.
Superficially they looked like ponderosae; but had the
typical zimmermani genitalia, and unrubbed examples
clearly showed the raised scaling.
Economically zimmermani is our most important
Dioryctria. In this country it does more serious dam-
age, especially to young trees, in both the East and the
West, than abietella. As far as I know it is strictly a
bark borer and does not attack the cones. The most
complete and accurate account of the life history is that
in the Craighead (1950) paper. The life history of the
insect in Nebraska is also treated in the Keen (1938)
paper under “‘ponderosae.”
318. Dioryctria cambiicola (Dyar)
Figures 390, 876
Pinipestis cambticola Dyar, Ins. Insc. Menstr., vol. 2, p. 2,
1914.—MecDunnough, Check list, No. 6137, 1939.
Maxillary palpus of male squamous.
Forewing as in zimmermani except: Ground color of
basal, submedial and terminal areas a dark, dull, reddish
brown (somewhat paler on Arizona specimens); the
black scaling greatly reduced; the white scaling follow-
ing antemedial and preceding subterminal lines and
bordering terminal margin; the transverse lines and
the discal mark, dull silvery. Hind wing smoky fuscous;
the veins darkened and a dark line along termen. Alar
expanse, 28-32 mm.
Genitalia, male and female, show no specific differ-
ences from those of zimmermant. The extent of the
bend in the female ductus bursae shown by the figures
is purely individual and can be easily accented or
diminished in preparing the slides.
TypE Locatity: Flathead Reservation, Mont. (type
in USNM).
Foop pLants: Pinus ponderosae, P. scopulorum, P.
coulteri. The larvae are cambium borers in new growth,
causing pitch exudations like those of zimmermant.
DistriBution: Montana, Flathead Reservation (July,
Aug.); Missoula (July); Colorado, Boulder (Sept.),
Palmer Park (July, Aug.); New Mexico, Las Vegas;
Arizona, Flagstaff, White Mts. (Aug.); California,
Julian (Sept.).
The species is doubtfully distinct from zimmermani
except as a possible race. It differs chiefly in its dark
hind wings. It apparently has a limited distribution
in our Western States. In his original description Dyar
mentions an eastern specimen (presumably from Wash-
ington, D. C.) reared from a cone of Pinus taeda, Aug.
14, 1882. I have also before me a similar female from
Cape Henry, reared June 9, 1927. I suspect that both
these examples may be hybrids of zimmermani and
amatella.
319. Dioryctria amatella (Hulst)
Fiaure 879
Nephopteryz amatella Hulst, Ent. Amer., vol. 3, p. 131, 1887.
Dioryctria amatella (Hulst), Ragonot, Ent. Amer., vol. 5, p. 114,
1889; Monograph, pt. 1, p. 191, 1893.—Craighead, U. S.
Dep. Agr. Mise. Publ. 657, 1950.
Pinipestis amatella (Hulst), Phycitidae of N. Amer., p. 136,
1890.—McDunnough, Check list, No. 6135, 1939.
Maxillary palpus of male squamous.
Forewing with raised scales as in zimmermani but the
scale ridges somewhat narrower; ground color a dark
wood brown; transverse lines, discal spot, a thin trans-
verse band preceding the subbasal ridge, a blotch fol-
lowing the antemedial line, a similar shade near inner
margin of subterminal line on lower half of wing, and
a thin zigzag pale shade just within terminal margin,
silvery white and strongly contrasted against the
ground color; black scaling limited to the subbasal and
submedian scale ridges, the thin borders of the trans-
verse lines, a spot at extreme base, and the thin terminal
line. Hind wing smoky with a somewhat glossy pale
brownish tint; the veins slightly darkened and a thin
dark line along terminal margin. Alar expanse, 27—
32 mm.
Genitalia essentially like those of zimmermani.
TypE Locauity: Florida (type in AMNH, ex Rutgers).
Foop pLants: Pinus spp. Larvae feeding in cones
and terminals, more often in the cones than terminals and
apparently favoring diseased cones.
Disrrisution: Florida, Alton (June), Camp Pinchot
(June), Fort Mead (May), Lake City (May, June),
Monticello (Sept.), Orlando (June), ‘Southern Florida’’
(June, July), Starke (May); ZLowiszana, New Orleans
(Sept., Nov.), Woodworth (June) ; Texas, Conroe (May);
Mississippi, Hattiesburg (June), Picayune (May);
Maryland, Baltimore (Aug.); District of Columbia,
Washington (July).
Close to but apparently distinct from zimmermani
and cambiicola, distinguished from both by the strong
contrast of its white markings. Generally distributed
in the Gulf States where its abundance, especially in
Florida and Louisiana, makes it something of a pest. A
brief account of what is known of its life history is given
in the Craighead (1950) paper.
320. Dioryctria albovittella (Hulst)
Fiaure 880
Pinipestis albovittella Hulst, Phycitidae of N. Amer. p. 138,
1900.—Barnes and McDunnough, Contributions, vol. 4, p.
174, 1918.—MeDunnough, Check list, No. 6138, 1939.
Dioryciria albovittella Ragonot, Monograph, pt. 1, p. 193, 1893.
Maxillary palpus of male squamous.
Forewing with a narrow black subbasal scale ridge,
little if any raised scaling otherwise; forewing gray
densely dusted with white, making the extreme basal,
median, and terminal areas a pale ash color; transverse
lines and discal mark well contrasted, white; antemedial
line nearly vertical, slightly notched above and below
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 157
middle, bordered outwardly by a thin black line and
inwardly by a broad dark shade which includes the
black raised-scale ridge; a white blotch on lower half of
wing just beyond the antemedial line; subterminal line
well defined, median section broadly triangulate, bor-
dered inwardly by a blackish line and outwardly by a
narrow dark shade; discal marking a white spot covering
discocellular vein; a row of confluent black dots along
termen. Hind wing white with a faint ocherous tint; a
fuscous shade at apex and, narrowly, along termen.
Alar expanse, 23-30 mm.
Male genitalia like those of zimmermani. Female
genitalia similar to those of cambiicola, differing only in
trifling details of the spining in the bursa, not exhibiting
any consistent differences of a specific character.
Typr tocatiry: Colorado (type in AMNH, ex
Rutgers).
Foop piant: Cones of Pinus monophylla. This rec-
ord from reared Nevada specimens received from the
Forest Insect Division of the U. S. Bureau of Entomol-
ogy and Plant Quarantine under Hopkins U. S. No.
32009.
Distripution: New Mexico, Jemez Mts. (Aug.),
Jemez Springs (Aug.); Arizona, Mohave County (July,
Aug.) ; Colorado; Utah, Dividend (Aug.), Eureka (Sept.),
Ibapah Mts. (Trout Creek, July); Nevada, Topaz Lake
(Aug.).
Another close relative of zimmermani and cambiicola,
distinguished chiefly by its much paler forewings. In
his original description Hulst gives Hot Springs, N.
Mex., as the type locality; but this, as pointed out by
Barnes and McDunnough (1918) is evidentally a lapsus.
The type is a male, labeled ‘‘Colo. Bruce.”
321. Dioryctria gulosella (Hulst), new combination
Figure 392
Acrobasis gulosella Hulst, Phycitidae of N. Amer., p. 126, 1890.—
Ragonot, Monograph, pt. 1, p. 109, 1893; pt. 2, p. 520, pl.
50, fig. 11, 1901.
Pinipestis gulosella (Hulst) Barnes and McDunnough, Check
list of the Lepidoptera of Boreal America, No. 5575, 1916.—
MceDunnough, Check list, No. 6139, 1939.
Maxillary palpus of male squamous.
Forewing with a subbasal scale ridge and small patch
of raised scales in median area; dark gray with a fine,
sparse, white dusting, making the general color a dark
ash gray; transverse lines whitish gray, distinct; ante-
medial line twice notched, edged outwardly by a thin
black line, preceded by an obscure pale patch on lower
half of wing, this followed on its inner margin by a black
scale ridge continued as a thin black line to costa; sub-
terminal line outwardly angled at middle, bordered
inwardly by a black line and outwardly by a narrow
dark shade; discal spot white; a narrow black line along
termen. Hind wing white, smoky at apex and some-
what along termen (especially on females); the veins
more or less darkened; a fine brown line along terminal
margin. Alar expanse, 21-27 mm.
Genitalia similar to those of bawmhoferi except for the
shape of the uncus. The latter has a more broadly
rounded terminal margin on gulosella (compare figs.
391 and 392).
Type tocauity: Hot Springs, N. Mex. (type in
AMNH, ex Rutgers).
Foop prant: Not definitely known, but undoubtedly
pine.
Disrrisution: New Mexico, Hot Springs (Aug.);
Colorado, Glenwood Springs (July, Aug.), also one
female with only the state locality (Bruce, collector)
and bearing a Hulst “type” label. The type from New
Mexico is also a female. The Colorado specimens (2
of and 4 9) are all in the National Collection. They
are a perfect match for Ragonot’s figure.
322. Dioryctria baumhoferi, new species
Figures 391, 881
Maxillary palpus of male squamous.
Forewing with raised scaling as on typical zimmer-
mani; blackish gray with the usual black markings indi-
cated by a more intense darkening of the ground color;
the transverse lines faint but distinguishable, gray;
antemedial line bordered inwardly by a broad black
band including the subbasal scale ridge; a similar
blackish, transverse shade across the middle of the
wing; subterminal line outwardly angulate at middle,
bordered inwardly by a black line and outwardly by a
rather broad black shade extended into streaks on some
of the veins; a black line along terminal margin; discal
spot whitish gray, sometimes very faint. Hind wing
smoky gray; the veins darkened and the smoky shade
intensified along termen. Alar expanse, 25-28 mm.
Male genitalia having uncus triangulate with nar-
rowly rounded apical margin. Female genitalia of the
zimmermani type but with ductus bursae shorter in
proportion to length of bursa and somewhat broader.
Typr LocaLity: Prescott, Ariz. (type in USNM,
61355).
Foop piant: Pinus ponderosa.
new growth.
Described from male type and one male and nine
female paratypes from the type locality, reared under
Hopkins Nos. 9932C and 18506, June 5, 6, and 11,
1928, by the late L. G. Baumhofer of the Forest Insect
Division of the U. S. Bureau of Entomology and Plant
Quarantine. Baumhofer’s extensive rearings and field
studies, especially in the Nebraska National Forest,
have contributed what knowledge we have of the
biology of the Dioryctria species in that area.
The new species is close to gulosella Hulst, from which
it is distinguished by the much darker color of its fore
and hind wings, differently shaped male uncus, and the
somewhat stouter spining of the male penis. It may
prove to be a local race of guloselia but is at least as
distinct from it as cambiicola is from zimmermant.
Larvae feeding in
323. Dioryctria subtracta, new species
FiaureEs 393, 882
Maxillary palpus of male squamous.
Forewing with a subbasal ridge of raised scales and
158 UNITED STATES NATIONAL MUSEUM BULLETIN 207
a small tuft in lower fold just beyond antemedial line;
dark gray finely peppered with white making the ground
color a dark ash gray; the subbasal scale ridge, outer
border of antemedial and inner border of subterminal
lines, and the small raised patch following antemedial
line, black strongly contrasted against the ground
color, the outer border of antemedial line somewhat
fainter than the other black markings; subbasal scale
ridge narrow, reaching almost to costa; between it and
antemedial line a broad pale band; antemedial line
obscure, indicated chiefly by its blackish outer bordering
line which is almost vertical and parallel with the scale
ridge; subterminal line also faint, pale gray, outwardly
angled at middle, bordered inwardly by a narrow black
line; discal spot obscure, a narrow grayish white line on
discocellular vein; a fine, strongly contrasted, black line
along terminal margin. Hind wing white, on female a
faint smoky tint towards apex; the outer parts of the
veins faintly darkened and a narrow brown line along
terminal margin. Alar expanse, 23-25 mm.
Male genitalia with uncus rather short in proportion
to width; its terminal margin broadly rounded. Harpe
narrow and rather short. Penis armed with two
groups of subterminal spines and a single, straight,
slender, rather short posterior spine; one of the anterior
groups consisting of a line of short spines along lateral
margin of penis near its apex.
TypH Locatity: Fort Wingate, N. Mex. (type in
USNM, 61356).
Foop pirant: Unknown.
Described from male type and one female paratype
from the type locality (July), and one female from Glen-
wood Springs, Colo. (Sept.) which McDunnough had
identified as gulosella Hulst. They are superficially
similar to the type of that species except for the stronger
contrast of the black markings on forewing; but are
easily separated by their radically different genitalia.
The expanded (bulbous) shape of the caudal end of the
female ductus bursae is found in only one other Ameri-
can species of Dioryctria (clarioralis) and there in a
lesser degree.
324, Dioryctria clarioralis (Walker)
Fiaures 394, 883
Nephopteryz clarioralis Walker, List, vol. 27, p. 54, 1863.
Dioryctria clarioralis (Walker) Ragonot, Ent. Amer., vol. 5, p.
114, 1889; Monograph, pt. 1, p. 193, 1893.—Hulst, Phy-
citidae of N. Amer., p. 136, 1890—Forbes, Cornell Mem.
68, p. 620, 1923.—MeDunnough, Check list, No. 6125, 1939.
Ulophora brunneella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 106, 1904.—Barnes and McDunnough, Contributions,
vol. 3, p. 195, 1916 (make synonym of clarioralis).
Maxillary palpus of male squamous.
Forewing with a weak subbasal ridge of raised scales
(the species has always been included in the smooth-
winged Dioryctria group, but unrubbed and unpressed
examples always show some traces of a raised subbasal
ridge); ground color grayish brown with black patches
and more or less white shading in the median and ter-
minal areas; a broad blackish band preceding the ante-
medial line, paling towards inner margin, bordered in-
wardly by the black raised-scale ridge and outwardly by
the thin, black inner border of the antemedial line; the
latter thin, oblique, sometimes weakly notched below
costa and more rarely at lower fold, white without (or
with only an occasional trace, near inner margin) the
normal black outer bordering line; a more or less ex-
tended black smudge in cell, sometimes extended as
far as the black inner border of subterminal line and
usually completely obliterating any trace of a white
discal spot; some white streaking on lower vein of cell;
subterminal line distinct, sharply indented between
costa and vein 5, thence vertical and straight to inner
margin, whitish with a thin black inner border; a fine
_line of confluent black dots along terminal margin.
Hind wing smoky gray or brownish; the veins more or
less darkened; a fine dark line along termen. Alar
expanse, 22-29 mm.
Male genitalia with uncus triangulate, appreciably
longer than greatest width, evenly tapering to very
narrowly rounded apex. Female with ductus bursae
broadened near genital opening but less so than in
subtracta; bursa much larger and more heavily spined
than that of subtracta, at least as long as ductus bursae.
TYPE LOCALITIES: “United States” (clarioralis, in
BM); Tryon, N. C. (brunneella, in USNM).
Foop piant: Pinus palustris. This record from a
specimen received from L. A. Hetrick reared from larva
feeding in the cone.
Distrisution: Florida, Dunedin (Mar.), Fort Myers
(Apr.), Miami; North Carolina, Tryon (May, June);
Massachusetts, Martha’s Vineyard (June, July).
Another intermediate between the smooth-winged
and rough-scaled species, easily distinguished by its
genitalia and wing pattern.
78. Genus Oryctometopia Ragonot
Oryctometopia Ragonot, Nouv. Gen., p. 11, 1888; Monograph,
pt. 1, p. 153, 1893.—Janse, Journ. Ent. Soc. South Africa,
vol. 4, p. 156, 1941. (Type of genus: Oryctometopia fossula-
tella Ragonot.)
Tongue well developed. Antenna of male with a
slight sinus and scale tuft in base of shaft; pubescent.
Labial palpus obliquely upturned, reaching to or a trifle
above vertex; third segment about two-thirds the length
of second, bluntly pointed, more or less deflected for-
ward. Maxillary palpi of both sexes broadly squamous,
the scales forming a flat cover over the face. Forewing
smooth; 11 veins; vein 2 from before but near lower
outer angle of cell; 3 from the angle, equidistant at base
from 2 and 4, parallel with 2 from just beyond base for
its remaining length; 4 and 5 connate or very shortly
stalked; 6 from below upper angle of cell, straight; 8
and 9 stalked for about two-thirds the length of 8; 10
from the cell, approximate to stalk of 8-9 at base; male
with costal fold containing a row of coarse scales. Hind
wing with vein 2 from before lower outer angle of cell;
3 from the angle; 4 and 5 stalked for about half their
lengths; 7 and 8 closely approximate beyond cell at least
for half their lengths; all veins long; cell about one-
fourth the length of wing; discocellular vein slightly
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 159
curved, not extended at lower angle. Highth abdom-
inal segment of male with a pair of long, slender, ventro-
lateral hair tufts.
Male genitalia with uncus subtriangulate. Apical
process of gnathos a short, stout, hook. Transtilla ab-
sent. Harpe with one or more short stout thornlike
spines projecting from lower margin of sacculus. Aedea-
gus slender, rather long, sinuate. Penis armed with a
single thornlike cornutus. Vinculum stout, longer
than broad, subtriangulate, narrowed from middle to
terminal margin.
Female genitalia with signum consisting of a single
round curved plate, densely armed with long, stiff
spines and covering the ventral and lateral caudal half
of the bursa copulatrix; bursa otherwise membranous;
a narrow sclerotized collar about ductus bursae at its
junction with bursa, the ductus bursae otherwise un-
sclerotized; genital opening simple; ductus seminalis
from bursa near junction of bursa and ductus bursae.
The genus, so far as we know, is confined to the New
World and contains but one tropical American species.
325. Oryctometopia fossulatella Ragonot
Ficures 53, 395, 900
Oryctomeiopia fossulatella Ragonot, Nouv. Gen. p. 11, 1888; Mon-
ograph, pt. 1, p. 153, 1893.
Phycita moeschleri Ragonot, Nouv. Gen., p. 12, 1888; Monograph,
pt. 1, p. 182, 1893—Méschler, Die Lepidopteren-Fauna
von Portorico, p. 328, 1890. (New synonymy.)
Forewing gray to pale brownish gray; the transverse
lines obscure but usually distinguishable; antemedial
line nearly vertical, far out from base, on well marked
specimens irregularly serrate and bordered outwardly
by a thin dark line which is more or less broken, pre-
ceded by an obscure reddish olivaceous (or reddish
brown) patch on inner margin, the latter bordered
inwardly by same blackish scaling; a similar obscure
reddish brown patch over lower fold just before the sub-
terminal line; subterminal line more distinct than ante-
medial, sinuate (outwardly bulged at middle), dull
whitish gray, bordered inwardly and outwardly by
narrow dark lines; discal dots, when distinguishable,
small, separated, blackish; a row of obscure blackish
dots along termen; on most specimens a dull whitish
patch over middle of inner margin. Hind wing trans-
lucent white; the veins not appreciably darkened; a con-
trasting dark shade along costa and narrowly along ter-
men; on males, a fine dark subbasal line through the
white cilia. Alar expanse, 15-20 mm.
Genitalia as given for the genus. The thornlike
spines on sacculus of the male harpe are individually
variable, consisting of one stout, hooked spine and two
or more slenderer spines, their number and size varying
not only in individuals but on opposite harpes of the
same specimen. Cornutus of penis with a flattened
platelike base.
Type LocauitiEs: “Irazu’’? [Mount Irazi], Costa
Rica (fossulatella, in BM); Puerto Rico (moeschleri, in
Paris Mus.).
Foop piant: Bauhinia mexicana. Larvae in the
pods. This record from Brownsville, Tex., specimens
reared by the Division of Foreign Plant Quarantine of
the U. S. Bureau of Entomology and Plant Quarantine.
Distrisution: Unitep States: Texas, Brownsville
(June). M*sxico: Chiapas (May), Jalapa, Oaxaca, Te-
huacin (Apr.). GuatTeMaua: Cayuga (May, June,
Sept.), Chejel (June, July, Aug.), Purulhé (July), Vol-
cin Santa Maria (May, June, July). Costa Rica:
Esperanza (May), Mount Irazi. Panami: Cabima
(May), Corazal (Apr.), La Chorrera (May), Porto
Bello (Oct.). VenrEzunLA: Aragus (Rancho Grande,
May). Braz: Rio de Janeiro (‘10-1-31”’). Puzrtro
Rico: Coamo Springs (Apr.), Covado (May), Puerto
Real (Vieques Isl., Apr. ), San German (Apr.). Vira@in
Isuanps: Kingshill (St. Croix, June).
The species is easily distinguished by its genitalia;
but in color and maculation is variable, as the foregoing
description indicates. Such variability is purely indi-
vidual and has no racial or local significance whatsoever.
Ragonot described fossulatella from a large (20 mm.)
male and his moeschleri from a small (17 mm.) female
color variant, which accounts for his two names and
their placement in different genera.
Genera 79-81: Sarata to Lipographis
[Venational division B. Forewing smooth; veins 4 and 5 sepa-
rated at base. Hind wing with veins 4 and 5 stalked; cell usually
short, about one-third the length of wing (longer in Lipographis,
about one-half). Labial palpus porrect, broadly scaled, beak-
like. Male genitalia with transtilla incomplete or absent; harpe
with costa partially sclerotized, not produced, clasper reduced or
absent, otherwise simple. Female genitalia with bursa smooth
or scobinate, without signum; ductus seminalis from bursa.]
79. Genus Sarata Ragonot
Sarata Ragonot, N. Amer. Phycitidae, p. 11, 1887; Monograph,
pt. 1, p. 614, 1893.—Hulst, Phycitidae of N. Amer. p. 168,
1890. (Type of genus: Sarata dnopherella Ragonot.)
Tongue well developed. Antenna pubescent; shaft
of male cylindrical, slightly swollen at base and smooth-
ly scaled or with a very slight ridge of roughened scales
along a few of the basal segments. Labial palpus
porrect, beaklike; second segment oblique, laterally
flattened, broadly scaled; third segment deflected for-
ward, about the length of second (sometimes a trifle
shorter or longer), bluntly acuminate. Maxillary pal-
pus minute, filiform. Forewing smooth; 11 veins;
vein 2 from well before lower outer angle of cell; 3 from
the angle; 4 and 5 separated at base; 6 from below upper
angle of cell, straight; 8 and 9 stalked for from one-half
to two-thirds their lengths; 10 from the cell, approxi-
mate to the stalk of 8-9 for a considerable distance from
base; male without costal fold. Hind wing with vein 2
from well before lower outer angle of cell; 3 from the
angle or separated from it by a very short spur; 4 and
5 stalked for two-thirds their lengths; 7 and 8 closely
approximate beyond cell for nearly half their lengths;
cell about one-third the length of wing; discocellular
160
vein curved, outwardly produced at lower angle of cell.
Eighth abdominal segment of male with a pair of weak,
ventrolateral hair tufts.
Male genitalia with uncus triangulate, its apex
rounded. Apical process of gnathos terminating in an
elongate hook, moderately long and bladelike (except in
incanella where it is short, and digitate from an enlarged
base); lateral arms of gnathos broad and stout. Trans-
tilla incomplete; its lateral elements well sclerotized and
short or moderately long and slender, their apices
pointed. Harpe, elongate, slender; apex of cucullus
rounded; clasper present as a very short, blunt, wartlike
projection from near middle of basal margin of cucullus
(except in incanella where it is broader and more scoop-
like), the size and shape of the wartlike clasper indi-
vidually variable; costa not produced, strongly sclero-
tized only on basal half. Anellus a shallow, broadly
U-shaped shield. Aedeagus long, straight, not appre-
ciably tapering or expanded towards apex, stout to
moderately slender (incanella); penis (except in incan-
ella) armed with a single, long, stout, spikelike cornutus,
rarely a second elongate slenderer spine (on penis of
incanella the single cornutus is @ short, stout thorn sit-
uated near apex). Vinculum stout, as long as or but a
trifle longer than broad; terminal margin broad.
Female genitalia with bursa strongly scobinate over
much of inner surface and more or less thickened
(cartilaginous) at or near anterior end; ductus bursae
simple (unsclerotized and unspined throughout), nor-
mally distinctly shorter than bursa; ductus seminalis
from bursa near its middle well forward of junction of
bursa and ductus bursae).
The species here referred to Sarata form a homogene-
ous group. Some of them on the basis of a smooth
male antennal shaft have hitherto been listed under the
Old World genus Megasis; but none agrees with the
type of the latter (rippertella (Zeller), fig. 428) on
genitalic characters or the stalking of veins 4—5 of hind
wing, which is always shorter (and frequently incom-
plete, a mere approximation or anastomosis of the basal
half of the veins in rippertella). The difference between
a smooth antennal shaft and one with some roughened
scales towards base is very slight, and should have no
weight against the uniformity of the genitalic and vena-
tional characters of Sarata otherwise. Indeed the
species of Sarata are much closer to those of Lipographis
than to the type of Megasis.
None of our species has been reared and nothing is
known of the food plants of any of them, so that any
association of females with males is purely speculative.
Such associations as have been made are open to grave
suspicion. The females differ from the males not only
in size but also in pattern and color, and within any
given species of females the color varies more between
individuals than it does between the species themselves.
For this reason I have treated the males and females
separately, giving to the latter new temporary names
which can go into synonymy when the sexes are properly
associated.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
The following three species hitherto listed in Megasis
or Sarata must be referred elsewhere:
Sarata rhoiella Dyar becomes the type of a new genus
(Philodema).
Sarata umbrella Dyar goes to Lipographis.
Megasis indianella Dyar is an anerastiine and a
synonym (see p. 315) of Ragonotia olivella (Hulst).
Genus Sarata (males), Species 326-330:
S. edwardsialis to S. incanella
[Antennal shaft at base smooth scaled.]
326. Sarata edwardsialis (Hulst), new combination
Figure 396
Megaphysis edwardsialis Hulst, Trans. Amer. Ent. Soc., vol. 13,
p. 163, 1886.
Megasis polyphemella Ragonot, N. Amer. Phycitidae, p. 10, 1887;
Monograph, pt. 1, p. 545, 1893.
Megasis edwardsialis (Hulst), Ent. Amer., vol. 5, p. 156, 1889;
Phycitidae of N. Amer., p. 165, 1890.—McDunnough, Check
list, No. 6259, 1939.
Forewing pale grayish brown more or less smudged
with darker gray; blackish streaks on several of the
veins, especially marked on vein 1b, the lower vein of
cell, and the veins immediately preceding and following
the subterminal line; the latter faintly indicated; ante-
medial line obsolete; discal spots poorly defined, often
obliterated, where distinguishable, separated; a row of
narrow black dots along termen, more or less accented.
Hind wing a little lighter grayish brown than ground
color of forewing, the veins not appreciably darkened,
a slightly darker line along termen; cilia paler, a dull
white with a faint fuscous tint. Alar expanse, 35-46
mm.
Male genitalia with uncus evenly tapering to narrowly
rounded apex. Apical process of gnathos about half as
long as uncus. Elements of transtilla very short.
Aedeagus long, stout; penis armed with a single, straight
cornutus, nearly as long as aedeagus, also a small sup-
plemental sclerotized patch. Terminal margin of vin-
culum concave.
TyprE LocaLitiEs: Nevada (edwardsialis, in AMNH,
ex Rutgers) ; California (polyphemella, in Paris Mus.).
Foop prant: Unknown.
Distrisution: Colorado, Boulder (Mar.), Glenwood
Springs (Mar., Apr.), Salida; Utah, Dividend (Mar.),
Eureka (Apr., May); Nevada; California, ““Middle Cali-
fornia’”’; Washington, Grand Coulee (Apr.), Pullman
(Apr.).
Average specimens (40 mm. or more) make this the
largest species in the genus. The Ragonot figure of
polyphemella (Monograph, pl. 19, fig. 8a) is a very good
likeness of normal examples, except that hind wing is a
trifle too dark.
327. Sarata pullatella (Ragonot), new combination
FIGurE 397
Megasis pullatella Ragonot, N. Amer. Phycitidae, p. 10, 1887;
Monograph, pt. 1, p. 547, 1893.
Smaller on the average than edwardsialis and darker.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 161
Forewing almost uniformly"suffused dark grayish fus-
cous (blackish gray on some examples); on some speci-
mens the basal and terminal areas paler by contrast
and the antemedial line indicated, but very faint, nearly
vertical ; on occasional specimens a paler brownish shade
in the cell; faint blackish streakings on the veins in
terminal area (especially on specimens with pale outer
area) and more or less of a black streak on lower vein
of cell; subterminal line very faint, often completely
obscured, when distinguishable indicated chiefly by a
broken black shading forming its inner border; discal
dots obscured. Hind wing pale to dark gray-brown;
the cilia whitish. Alar expanse, 29-36 mm.
Male genitalia figured from type and a typical speci-
men from Dividend, Utah, to show extent of individual
variation. Uncus not so evenly tapering as in edward-
sialis; its lateral margins slightly angled at middle.
Aedeagus less stout and the single long cornutus on penis
more slender than those of edwardsialis; no supplemental
patch on penis.
Type Locatity: California (type in Paris Mus.).
Foop piant: Unknown.
Distrrisution: Utah, Dividend (Apr.); California,
San Diego (Jan.); Jdaho, Malta; Washington, Kamiack
Butte (Feb.), Pullman (Apr.).
Easily confused with smaller specimens of edwardsialis,
and all specimens in the National Collection had been so
identified; but otherwise distinguished by its genitalia.
Hulst in his Phycitidae of North America (1890)
made pullatella a synonym of his excantalis and it has
since appeared as such in our lists. Since the type of
excantalis is a female, the synonymy is doubtful, to say
the least.
328. Sarata punctella (Dyar), new combination
Figure 398
Megasis punctella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 404,
1915.
Forewing pale brownish gray with a slight rufous tint
through the cell; antemedial line indicated by an outer
border of three black dots, one below costa, a second on
lower vein of cell, and a third on vein 1b; subterminal
line obscure but usually distinguishable, a whitish spot
on costa, preceded and followed by blackish dots and,
below, bordered inwardly by short blackish streaklet on
veins; on typical specimens some faint, blackish streak-
lets on the veins of outer area (following the subterminal
line); a row of very faint blackish dots along termen;
discal dots obsolete or very faint. Some specimens
show little or no trace of the blackish markings. Hind
wings pale brownish gray (“mouse gray”’); the cilia but
slightly paler, a narrow dark line along termen. Alar
expanse, 25-30 mm.
Male genitalia with uncus elongate, slightly and
evenly tapering to rounded apex. Apical process of
gnathos appreciably shorter than in preceding species,
slender. Elements of transtilla long, slender. Penis
armed with two cornuti—one stout, slightly bent or
300329—56—12
sinuate, about half as long as aedeagus; the other a
slender, flattened spine, as long as aedeagus.
TypE Locauity: Tehuacin, México (type in USNM).
Foop prant: Unknown.
Distrisution: M&xico: ‘Las Vigas’’ [probably Las
Viagas, Vera Cruz], Tehuacin (Oazaca, Sept.), Uruapan
(Michoacdn, Mar.). Also three specimens labeled sim-
ply “V.5.” One of the latter was before Dyar when he
described hisspecies. It had been identified by Druceas
“Zophodia inornatella Rag.”
Dyar’s short original description is thoroughly ade-
quate and accurate for the type but takes no account
of suffused examples which lack the characteristic
blackish markings.
329. Sarata punctella septentrionaria, new race
Like typical punctella but larger; the Arizona, New
Mexico, and Colorado examples a suffused pale gray-
brown with the usual dark markings obsolete or nearly
so; the Wyoming example more grayish with most of the
veins faintly streaked with blackish scaling. The
genitalia agree with those of the type of punctella. Alar
expanse, 32-35 mm.
Typr Locauity: Palmerlee, Ariz. (type in USNM,
61357).
Foop prant: Unknown.
Described from male type from the type locality,
“Apr. 1-7,” and one male paratype from each of the
following localities: Fort Wingate, N. Mex. (Apr. 14,
1908); Golden, Colo., Dyar and Caudell, No. 16259;
Medicine Bow, Wyo. July 4, 1936, I. H. Blake, No.
321-11.”
Possibly a distinct species but probably only a larger
northern variety of punctella. When collections are
made in the poorly explored areas of northern México
the species should show a continuous distribution.
330. Sarata incanella (Hulst), new combination
Ficure 399
Epischnia incanella Hulst, Canadian Ent., vol. 27, p. 56, 1895.—
McDunnough, Check list, No. 6257, 1939.
Megasis aridella Dyar, Proc. Ent. Soc., Washington, vol. 7, p.
35, 1905.—McDunnough, Check list, No. 6263, 1939. (New
synonymy.)
Forewing gray heavily dusted with white especially
in outer area and along costa at base and middle; the
median area between the transverse lines darker than
remainder of wing; dark shading also on basal area below
costa; black streaking on the veins, especially pro-
nounced in outer area, on vein 1b and upper and lower
veins of cell; transverse lines white, distinctly outlined ;
antemedial line zigzag, nearly vertical, with some black-
ish shading along outer margin; subterminal line sharply
indented at veins 6 and 1b; discal dots separated, black-
ish; terminal dots faint but distinguishable, separated.
Hind wing whitish with a faint smoky tint; a very faint
dark line along termen; cilia concolorous with wing.
Alar expanse, 31-36 mm.
Male genitalia with uncus as long as greatest width;
its apical margin broadly rounded. Apical process of
162 UNITED STATES NATIONAL MUSEUM BULLETIN 207
gnathos a short, slender hook arising from a thickened
base. Clasper of harpe broad, squarish. Aedeagus
rather short; penis armed with a single small thornlike
cornutus. Vinculum with terminal margin broadly and
evenly rounded; in all other species of the genus the
terminal margin distinctly concave.
TypE LocaLities: Colorado (incanella, in USNM);
Stockton, Utah (aridella, in USNM).
Foop prant: Unknown.
Distrizution: Colorado; Utah, Stockton (May);
California, Inyo County (May).
The palest of the Sarata species, except tephrella Rag-
onot. The genitalia of the types of incanella and ari-
della are identical.
Genus Sarata (males) Species 331-337: S. atrella
to S. tephrella
[Antennal shaft with a ridge of roughened scales at base.]
331. Sarata atrella (Hulst), new combination
Ficure 400
Megasis atrella Hulst, Phycitidae of N. Amer., p. 166, 1890.—
Ragonot, Monograph, pt. 1, p. 548, 1893.—McDunnough,
Check list, No. 6268, 1939.
Vestiture of labial palpi, head, and prothorax a mix-
ture of scales and long hairs.
Forewing blackish gray; transverse lines whitish gray,
dull; the antemedial line obscure; subterminal line
stronger, always distinguishable, bordered inwardly and
outwardly by black streaklets on the veins; discal spots
obsolete, obscured in the dark ground color; blackish
dots along termen confluent. Hind wing pale to rather
dark smoky fuscous, a thin dark line along termen. Alar
expanse, 30-33 mm.
Male genitalia with uncus elongate, evenly tapering
to narrowly rounded apex. Apical process of gnathos
about half as long as uncus, ventrally flattened. Ele-
ments of transtilla short. Penis armed with a single
strong cornutus, as long as aedeagus, no supplemental
sclerotized plate. —
Type Locauiry: West Cliff, Colo. (type in AMNH,
ex Rutgers).
Foop prant: Unknown.
Distrizution: Colorado, Custer County, West Cliff,
and two specimens with only the state locality.
The species is easily distinguished by the hairy vesti-
ture not possessed by any other known species in the
genus.
332. Sarata caudellella (Dyar), new combination
Ficure 401
Megasis caudellella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p- 110, 1904.—McDunnough, Check list, No. 6261, 1939.
Similar to the foregoing species (atrella) except: With-
out hairy vestiture and with some fine white powdering
on the blackish gray ground color of forewing; the
transverse lines distinct, oblique and nearly straight;
antemedial line followed on costa and preceded at inner
margin by obscure dark blotches; subterminal line bor-
dered inwardly by a continuous irregular blackish
shade, outwardly by a short, faint, dark streaking of
the veins; the veins otherwise not appreciably streaked;
discal dots faint, but usually distinguishable, more or
less confluent; dots along terminal margin weak, fused
into a faint blackish line. Hind wing pale brownish
gray, semilustrous; a dark line along termen. Alar ex-
panse, 28-32 mm.
Male genitalia differ from those of atrella only in
trifling details.
Typz Locatitry: Golden, Colo. (type in USNM).
Foop piant: Unknown.
Disrrisution: Unirep Starrs: Colorado, Golden
(May); Washington, Grand Coulee (Apr.). CANADA:
Saskatchewan, Oxbow (June) ; Manitoba, Aweme (Apr.),
Miniota.
Close to but distinct from atrella, from which it is
distinguished chiefly by its smooth-scaled vestiture
and the strong contrast of the whitish transverse lines
of forewing, especially the well-marked antemedial line;
333. Sarata dnopherella Ragonot
Fieure 402
Sarata dnopherella Ragonot, N. Amer. Phycitidae, p. 11, 1887;
Monograph, pt. 1, p. 616, 1893.
This and the two species following (nigrifasciella and
cinereelia) are very close, separable only by minor color
differences and trifling variations in the genitalia of their
types. Such variations are more than covered in the
series of nigrifasciella and cinereella before me. I sus-
pect that the names represent nothing but color varie-
ties of one variable species; but am keeping them sepa-
rate until life-history information and more extensive
collections are available and more exact definitions of
species and possible races can be made.
Ragonot’s dnopherella is authentically represented
only by its type. His description and figure suggest a
grayish brown form suffused with blackish brown and
with the transverse lines very weakly contrasted and
poorly defined against the ground color. Alar expanse,
32 mm.
Male genitalia with uncus broadly and bluntly tri-
angulate, evenly tapering to narrowly rounded apex.
Elements of transtilla moderately long, slender (about
half the length of those of atrella). Penis armed with a
single stout cornutus, slightly more than half as long
as aedeagus and preceded basally by a small, weakly
sclerotized patch.
TyprE Locaity: California (type in Paris Mus.).
Foop piant: Unknown.
Hulst (Phycitidae of N. Amer., p. 168, 1890) treated
dnopherella as a synonym of his perfuscalis and it so
appears in our lists. As perfuscalis was described from
a female the synonymising of the two names was
arbitrary and, under the circumstances, unwarranted.
Ragonot (Monograph, p. 616, 1893) very rightly ques-
tioned it.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 163
334. Sarata nigrifasciella Ragonot
Fiaure 403
Sarata nigrifasciella Ragonot, N. Amer. Phycitidae, p. 11, 1887;
Monograph, pt. 1, p. 615, 1893.—Hulst, Phycitidae of N.
eae p. 169, 1890.—McDunnough, Check list, No. 6266,
1939.
Forewing pale to dark ashy gray, some specimens
tinted with a faint brownish shade (one specimen before
me from Mineralking, Calif., a very pale, sordid
brownish gray). The transverse lines distinct, but
indicated chiefly by the blackish outer border of the
antemedial line and a similar inner border to the sub-
terminal line. This character, however, is variable and
on the Idaho specimen before me is almost obsolete.
Hind wing very pale brown, with a fine dark terminal
line. Alar expanse, 29-32 mm.
Male genitalia similar to those of dnopherella except
for the narrower apical process of gnathos (viewed
ventrally). This character, however, is not reliable;
for in a typical series of cinereella all intergrades are
found between the gnathos of typical nigrifasciella and
that of dnopherella.
Type Locatity: America Septentrionalis (type in
Paris Mus.).
Foop puant: Unknown.
Disrrisution: Unirep Srares: New Mexico, Fort
Wingate (Mar.); Colorado, Chimney Gulch (June),
Platte Canyon (June); Jdaho, Wallace (Apr.); Cali-
fornia, Mineralking (July), Yosemite (this specimen
labeled nigrifasciella in Ragonot’s handwriting). Can-
ADA: Manitoba, Aweme (Apr.).
This species, if such it be, is distinguished from the
preceding and following species chiefly by its more
marked transverse lines in forewing and the narrower
apical process of its gnathos. Ragonot’s description
of the females (also from “Amer. Sept.’’) associated
with his male type can be ignored, for they represent
two distinct species.
335. Sarata cinereella Hulst
Sarata cinereella Hulst, Canadian Ent., vol. 32, p. 172, 1900.—
McDunnough, Check list, No. 6271, 1939.
Forewing dark ash gray, nearly concolorous; the
transverse lines obsolete or nearly so; the subterminal
line faintly indicated on most specimens and, on one or
two, very faint traces of the antemedial line; as on
dnopherella and nigrifasciella there is more or less
blackish dusting on the veins. Hind wing pale brown-
ish gray. Alar expanse, 29-33 mm.
Male genitalia of type show a slightly longer cornutus
than that of nigrifasciella, but other examples inter-
grade in all characters between the two types.
Type Locauity: Salida, Colo. (type in AMNH, ex
Rutgers).
Foop piant: Unknown.
Disrrisution: Colorado, Denver (Mar.), Glenwood
Springs, Salida, also three examples with only state
locality, two of them bearing Hulst ‘‘type’’ labels
(Bruce, collector), and the other a pseudotype of
‘“Anerastia excantalis Hulst.”’
Probably only a suffused form of nigrifasciella.
336. Sarata rubrithoracella (Barnes and McDunnough), new
combination
Fieurs 404
Megasis rubrithoracella Barnes and McDunnough, Contributions,
veer p. 140, 1913 —McDunnough, Check list, No. 6268,
Forewing pale brownish gray rather evenly dusted
with white intermixed with a sparse peppering of
blackish scales; the transverse lines distinct, but faint,
whitish; antemedial line bordered outwardly by a thin,
broken, blackish shade; subterminal line bordered in-
wardly by a continuous narrow blackish band; no ap-
preciable streaking on the veins; discal spots obsolete;
terminal dots confluent, forming a faint, dark line.
Hind wing pale grayish brown; a thin dark line along
termen. Alar expanse, 22-29 mm.
Male genitalia show little to distinguish them from
those of the preceding three species except for the very
short lateral elements of transtilla and a slightly longer
cornutus (our drawing of the aedeagus is in reversed
position from that of other species; if drawn as were
the other aedeagi the apex of cornutus would point to
the right).
Tyrer Locatity: White Mts., Ariz. (type in USNM).
Foop pLant: Unknown.
Distrisution: Arizona, White Mts.; New Mezico,
Fort Wingate (Apr., July).
Apparently a distinct species. Differs from the
others by a distinct rufous-ocherous shading on the
thorax and, in genitalia, from the species of the dnoph-
erella—cinereella group by the short transtilla elements
of its genitalia. The rufous ocherous thorax occurs
also in two species of females (kappa and phi).
337. Sarata tephrella Ragonot
Fiaure 405
Sarata tephrella Ragonot, Monograph, pt. 2, p. 616, 1893.—
McDunnough, Check list, No. 6268, 1939.
Known to me only from Ragonot’s description and
figure, and the genitalia of its type. Evidently a much
paler species than any other of those with a rough-scaled
antennal shaft, and having much the general habitus of
a Lipographis. Forewing heavily dusted with white;
some ocherous brown shading in median and outer
areas; transverse lines whitish, distinguishable but
poorly defined. Hind wing dark gray. Alar expanse,
28 mm.
Male genitalia similar to those of the dnopherella-
cinereella group except elements of transtilla larger and
cornutus more slender and more evenly tapering.
Type tocauity: ‘Washington Territory” (type in
Paris Mus.).
Foop prant: Unknown.
Described and so far known only from its male type.
164 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Genus Sarata (females), Species 338-346:
S. alpha to S. delta
[The females are of a uniform pattern and similar coloration; the
ground color gray, sometimes tinted with pale brown and more
or less dusted with white, variations in color more individual
than specific; the transverse lines strongly contrasted, white with
strongly accented blackish or brownish borders on outer margin
of antemedial and inner margin of subterminal lines; the lines
oblique and straight or notched, the notching also more individual
than specific in character. The only reliable specific characters
are in the genitalia, and for certain identification it is necessary
to dissect nearly all females.]
338. Sarata alpha, new species
FIGURE 888
A bright species with the white lines and their black
borders sharply contrasted. Forewing blackish gray
with a strong dusting of white (however, in one speci-
men from the type locality, the median area distinctly
darker than basal or outer areas); antemedial line
slightly curved ; subterminal line straight or with a very
slight bend at lower fold; discal dots distinct, more or
less confluent. Hind wing whitish with a faint brown
tint; the veins slightly darkened; a broadened blackish
brown line along termen; cilia white. Alar expanse,
21-24 mm.
Female genitalia with bursa large and greatly elon-
gated; densely and finely spined over most of interior
surface, the denser spining in longitudinal rows, partial-
ly divided by lines of the clear membrane; anterior end
thickened (cartilaginous), the amount of thickening
individually variable; ductus bursae very short.
TYPE LOCALITY: Oxbow, Saskatchewan, Canada (type
in USNM, 61358).
Foop pirant: Unknown.
Described from female type and two female paratypes
from the type locality, May 14, 1907, Frederick Knab;
and additional female paratypes as follows: One from
Aweme, Manitoba, Apr. 12, 1908. N. Criddle; one from
Regina, Saskatchewan, June 5, 1907; and two from
Chimney Gulch, Golden, Colo., July, Oslar. A female
in the Rutgers Collection (C. H. slide No. 2186) from
Colorado identified as atrella also goes here. Needless
to say there is no trace on any of these females of the
hairy vestiture of the male of atredla.
339. Sarata beta, new species
Figure 889
Similar to alpha except less glossy. Forewing duller;
less white dusting; transverse lines and their black
borders less strongly contrasted; discal dots obscured.
Hind wing dark smoky gray; the veins not darkened.
Alar expanse, 23-26 mm.
Female genitalia like those of alpha in shape and
proportions except that spining covers appreciably less
of the bursa surface, leaving half or more than half of
the latter membranous and unspined.
Type Locauity: Colorado (type in USNM, 61359).
Foop prant: Unknown.
Described from female type with only a state locality
label; one female paratype from Custer County, Colo.;
and one female paratype from Chilcotin, British
Columbia, May 2, 1920, E. K. Buckell No. 137. The
two Colorado examples were in the Barnes and National
Museum Collections as females of atrella Hulst.
340. Sarata gamma, new species
Figure 890
This is the female figured in the Ragonot Monograph
(pt. 1, pl. 23, fig. 2b.) as a paratype of his dnopher-
ella. Its genitalia, here figured, are similar to those of
alpha and beta except for slight differences in the spiming
of the bursa, as shown in the figure.
Type Locatity: California (type in Paris Mus.).
Foop prant: Unknown.
341. Sarata iota, new species
FIGURE 894
This name is proposed for the female paratype of
pullatella Ragonot, described and figured by him
(Monograph, pt. 1, p. 547, pl. 19, fig. 9b, 1893).
Forewing blackish gray with very little pale dusting;
the antemedial line broader and more strongly con-
trasted than the subterminal. Hind wing dark grayish
brown. Alar expanse, 24 mm.
Genitalia (C. H. slide No. 3113) with bursa, consider-
ably smaller and more sparsely spined than that of any
of the preceding species; ductus bursze about half as
long as bursa.
Tyrer Locauity: California (type in Paris Mus.).
Foop puant: Unknown.
It is very likely that this is the female of pullatella;
but at the present time there is no certainty about any
of the sex associations in the genus.
342. Sarata perfuscalis (Hulst)
Figure 893
Nephopteryx perfuscalis Hulst, Trans. Amer. Ent. Soc., vol. 13,
p. 161, 1886
Anerastia excantalis Hulst, Trans, Amer. Ent. Soc., vol. 13,
p. 163, 1886 (mew synonymy).
Megasis excantalis (Hulst), Ent. Amer., vol. 5, p. 156, 1889;
Phycitidae of N. Amer., p. 165, 1890.—McDunnough,
Check list, No. 6260, 1939.
Sarata perfuscalis (Hulst), Phycitidae of N. Amer., p. 168, 1890.—
MceDunnough, Check list, No. 6269, 1939.
Forewing dull, dark gray more or less dusted with
white; the terminal and (usually) the median areas the
paler, the basal area the darker; transverse lines dis-
tinct, sordid white, their dark borders well contrasted,
especially on specimens with considerable white dusting.
Hind wing smoky grayish brown. Alar expanse, 25—
29 mm.
Female genitalia distinguished by the spining of the
large bursa. These spines are arranged in an elongate,
ribbed band which extends most of the length of the
inner dorsolateral surface, curving onto ventral surface
at anterior end; the area of bursa under the spines more
or less sclerotized.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 165
TyprE Locauity: California (perfuscalis, excantalis, in
AMNGH, ex Rutgers).
Foop prant: Unknown.
Distrisution: California (state locality only);
Washington, Seattle; Utah, Eureka (Mar., Apr.),
Stockton (Apr.).
Hulst associated his perfuscalis with dnopherella
Ragonot; and excantalis with puillatella Ragonot. So
much for superficial sex associations. Their genitalia
show the two females to be obviously conspecific.
343. Sarata epsilon, new species
Figure 892
The smallest of the female species. Similar in colora-
tion to perfuscalis except that the dark borders of the
transverse whitish lines are somewhat broader and more
strongly contrasted. Alar expanse, 19-21 mm.
Female genitalia with bursa moderately large, evenly
and finely spined on anterolateral half and extreme
anterior end.
TypE Locauity: Yosemite, Calif. (type in USNM,
61360).
Foop prant: Unknown.
Described from female type from the type locality
identified by Ragonot as a female of nigrifasciella and
bearing that name label in his handwriting and female
paratypes as follows: One from Colorado, Cockerell,
collector, identified by Hulst as atrella; two from Golden,
Colo., May, Dyar and Caudell Nos. 16252 and 16253,
and identified by Dyar as paratypes of caudellella;
seven from Chimney Gulch, Golden, Colo., June,
Oslar; and one from Fort Wingate, N. Mex., March.
Here also is referrable one of the two female paratypes of
nigrifasciella Ragonot in the Paris Museum (C. H. slide
No. 2891).
Most of the foregoing examples show at least one of
the black discal spots. On each of the seven specimens
from Chimney Gulch, Colo., is 8 minute white spot on
discocellular vein between the black dots and on these
also there is some very dark brown shading on the
otherwise blackish borders of the white transverse lines.
344. Sarata phi, new species
Fiaure 891
Thorax shaded with rufus ocherous.
Forewing more evenly dusted with white; dark
borders of the transverse lines distinctly brownish; dis-
cal spots obsolete, replaced by a faint, white line or spot
on the discocellular vein. Alar expanse, 23-27 mm.
Female genitalia essentially like those of epsilon. The
figure shows the bursa twisted into a reverse position
from that of epsilon to show the somewhat greater
development of the thickened (cartilagenous) lateral
margin (a variable and probably only an individual
character).
TypE LocaLity: White Mts., Ariz.
61361).
Foop piant: Unknown.
Described from female type from the type locality
(type in USNM,
(the female paratype of rubithoracella Barnes and
McDunnough); and five female paratypes from Fort
Wingate, N. Mex. March, June, July. Also before
me, but not included among the paratypes, is a large
female (26.5 mm.) from Denver, Colo., Apr. 1, 1904,
Oslar. On this specimen the patagia are more putty
white than rufus. Its genitalia, however, agree in
detail with those of typical phi from Arizona and New
Mexico. The species is uncomfortably close to epsilon.
It probably does represent the female of rubithoracella
but the verification of that relation will have to wait.
upon rearing evidence.
345. Sarata kappa, new species
FIGURE 887
Forewing dull, as in beta, but with considerable white
dusting, rather evenly distributed; the transverse lines
more irregular and their black borders more strongly
contrasted; subterminal line with slight notches at vein
6 and lower fold; lower discal dot faint, but distinguish-
able. The thorax of the type is strongly shaded with
rufus-ocherous. Alar expanse, 23 mm.
Female genitalia with bursa copulatrix very small
(the smallest of any of the Sarata species); the greater
part of its inner dorsal surface covered with a dense mat
of very fine spines. Ductus bursae as long as bursa.
Tyrer Locauity: Arizona (type in USNM, 61362).
Foop piant: Unknown.
Described from a pseudotype of perfuscalis Hulst
from the Fernald Collection, bearing only a state
locality and numbered ‘‘7820.”
In coloration similar to phi except for the blackish
borders of the transverse lines of forewing. Dis-
tinguished from that and other species of the genus by
its genitalia.
346. Sarata delta, new species
Ficure 886
This name is proposed for the second of the female
paratypes of nigrifasciella Ragonot (in Paris Mus., C.
H. slide No. 3111) whose genitalia are here figured.
80. Philodema, new genus
Tyre or Genus: Sarata rhoiella Dyar.
Tongue well developed. Antenna pubescent; shaft
of male slightly flattened and with a shallow sinus near
base, the latter containing some slightly roughened
scales and a few, minute serrations. Labial palpus
porrect (as in Sarata but shorter). Maxillary palpus
vestigial. Venation as in Sarata except veins 4 and 5
of hind wing stalked for half or less than half their
lengths and cell a short one-third the length of wing.
Eighth abdominal segment of male with ventrolateral
hair tufts.
Male genitalia as in Sarata except: More squat,
broader in proportion to their length; harpe short in
proportion to its width; no erect clasper; anellus
strongly sclerotized, its central area developed into a
pair of produced, pointed, bladelike arms, the usual
166
lateral lobes absent; aedeagus slender, its basal end
broadened and flattened; penis without cornutus or
other armature except for a few weak scobinations at
apex; vinculum stout, shorter than its greatest width.
Female genitalia with bursa small, simple, membran-
ous; ductus bursae short with a strongly sclerotized,
curved, wide, centrally notched, dorsal plate behind
genital opening; ductus seminalis from a small lobe of
bursa near junction of bursa and ductus bursae.
The genus falls between Sarata and Hypochalcia and
has several features of each of these genera but can go
into neither of them on the sum of its characters. It
appears to be a New World analogue of the Old World
Hypochalcia, agreeing with the latter on most genitalic
characters except for its short vinculum and peculiarly
developed anellus. The type of Hypochalcia (ahenella
(Zeller), fig. 54) has an elongate vinculum. It also
differs from Philodema in having much longer, smoother
and slenderer labial palpi, rather broad, squamous
maxillary palpi and smoother more glossy wing vesti-
ture.
Philodema differs markedly from Sarata in that there
is no sexual dimorphism, the males and females being
alike in color and markings.
347, Philodema rhoiella (Dyar), new combination
Fiaures 406, 895
Sarata rhoiella Dyar, Journ. New York Ent. Soc., vol. 12, p. 105,
1903.—McDunnough, Check list, No. 6267, 1939.
Forewing pale, sordid, brownish gray; extreme basal
area dark smoky gray; the transverse lines indicated
chiefly by their dark borders, the latter dark smoky
gray; the outer border of the antemedial line more or
less broken and diffused; subterminal line bordered
inwardly by an irregular (zigzag) border, somewhat ac-
cented on the veins, and outwardly by a more obscure
dark shade; discal dots distinct, separated. Hind wing
smoky gray; the veins slightly darkened; a narrow dark
shade along termen. Alar expanse, 23-33 mm.
Genitalia as given for the genus; figured from para-
types from the type locality. The male holotype was
without an abdomen.
Tyre tocauity: Platte Canyon, Colo. (type in
USNM).
Foop puant: Rhus toxicodendron.
Distripution: Colorado, Platte Canyon (July) ; Utah,
“So. Utah” (July).
The Utah specimens (2 <) are larger (82-33 mm.)
than any of the Colorado examples; but have identical
genitalia and wing maculation; nothing is known of the
life history except Dyar’s statement that two specimens
of the type series were reared from larvae on poison-ivy.
81. Genus Lipographis Ragonot
Lipographis Ragonot, N. Amer. Phycitidae, p. 10, 1887; Mono-
graph, pt. 1, p. 562, 1893.—Hulst, Phycitidae of N. Amer.,
p. 166, 1890. (Type of genus: Pempelia fenestrella Packard.)
Tongue well developed. Antenna weakly pubescent;
on male, shaft flattened, serrate, and with sinus and
UNITED STATES NATIONAL MUSEUM BULLETIN 207
strong scale tuft at base (except in umbrella and subos-
seella where the shallow sinus has a weak tuft of rough-
ened scales). Labial palpus porrect, second segment
oblique, laterally flattened, broadly scaled; third seg-
ment deflected forward, decidedly shorter than second,
its proportions obscured by its long scaling and the
extended scaling of second segment. Maxillary palpus
subsquamous (small but broadly scaled, vestigial in
umbrella). Forewing smooth; venation as in Sarata
except for a somewhat shorter stalking of veins 8 and 9
of forewing and a longer cell in hind wing (nearly one-
half the length of the wing); 4 and 5 are also shorter
stalked, about one-half their lengths.’ Highth abdomi-
nal segment of male with a pair of ventrolateral hair or
scale tufts (absent in subosseella).
Male genitalia with apical projection of gnathos a
short stout hook (except in subosseella). Transtilla
absent except in fruncatella and subosseella where it is
represented by its short, weak, divided elements. Anel-
lus with short, weak, lateral lobes (except in subosseella).
Aedeagus broadly expanded towards apex (except in
truncatella); penis armed with one or more strongly
sclerotized, curved, spinelike cornuti (the latter always
decidedly less than half as long as the aedeagus). Geni-
talia otherwise as in Sarata.
Female genitalia with bursa membranous and greatly
reduced, if sometimes elongate (truncatella) narrow;
ductus bursae scobinate and partially sclerotized near
its junction with bursa copulatrix, greatly broadened in
proportion to width of bursa (except in éruncatella);
genital opening simple, unsclerotized; ductus seminalis
from bursa near junction of bursa and ductus bursae.
Lipographis agrees with Philodema and differs from
Sarata in that the males and females are alike in color
and markings. It is distinguished from both Sarata
and Philodema chiefly by its female genitalia. The
latter resemble those of the type of the Old World
Divona Ragonot (ilignella (Zeller)) except that the
bursa of 2lignella is strongly scobinate, partially sclero-
tized, and proportionally much larger.
In his original description of Lipographis, Ragonot
designated fenestrella as type of the genus. Later
(Monograph, 1898) he cites humilis as its type. ‘This
substituted designation is invalid, regardless of the fact
that humilis was an originally included species and may
have served as the basis for the original generic descrip-
tion. That humilis now proves to be a synonym of
fenestrella is also immaterial and irrelevant.
One species (swbosseella) originally described in Iipo-
graphis is here provisionally retained in the genus. It
may eventually have to have a new generic placement
as its only representation (the male type) is aberrant
in several genitalic details.
7 The venation of fenestrella and leoninella exhibit considerable
individual variation; veins 4 and 5 of forewing are normally ap-
proximate for a short distance from cell but sometimes divergent
and (rarely) even shortly stalked. In one freak specimen before
me vein 4 is also absent from hind wing, another example which
advises caution against relying too much upon one structure for
the identification of phycitids.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 167
348. Lipographis fenestrella (Packard)
Ficurrs 31, 407, 896
Pempelia fenestrella Packard, Ann. New York Lyc. Nat. Hist.,
vol. 10, p. 259, 1873.
Nephopteryx fenestrella (Packard) Grote, Bull. U. 8S. Geol. Geogr.
Surv. Terr., p. 697, 1878.
Lipographis humilis Ragonot, N. Amer. Phycitidae, p. 11, 1887;
Monograph, pt. 1, p. 563, 1893.—Hulst, Phycitidae of N.
Amer., p. 167, 1890. (New synonymy.)
Lipographis fenestrella (Packard) Ragonot, N. Amer. Phycitidae,
p- 10, 1887; Monograph, pt. 1, p. 564, 1893.—Hulst, Phy-
citidae of N. Amer. p. 166, 1890.—McDunnough, Check
list, No. 6272, 1939.
Forewing ash gray, dusted with white and shaded
with brownish ocherous; the transverse lines narrow,
white; antemedial line oblique, nearly straight, bordered
inwardly by a broad brownish ocherous band marked
by black dots or streaklets on vein 1b and upper and
lower veins of cell, followed outwardly by two or three
similar black dots; white dusting along lower vein of
cell, median part of vein 1b and along some of the veins
preceding the subterminal line; subterminal line parallel
to termen, very slightly indented at veins 6 and 1b,
bordered outwardly by a broad brownish ocherous band
(the latter interrupted by blackish streaklets on the
veins) and from costa by short, faint, narrow, inner and
outer, blackish bordering lines; along termen a narrow
dusting of white; terminal dots more or less confluent,
individually variable, forming sometimes a straight,
sometimes a scalloped, black line; discal dots separated,
small, blackish; usually a brownish ocherous shade
along median area of lower fold. Hind wing dull white
with a faint, smoky tint towards apex and termen;a fine
blackish line along terminal margin; the veins not
appreciably darkened. Alar expanse, 19.5-24 mm.
Male genitalia with aedeagus decidedly bulged from
shortly beyond base; penis armed with a comb of 5
stout, curved spines of a graduating length. Female
genitalia with bursa greatly reduced and but slightly
longer than ductus bursae; the latter appreciably
broader than the bursa.
Type tocauity: California (fenestrella, in MCZ; hu-
milis, in Paris Mus.).
Foop prant: Unknown.
Distrizution: California, ‘Middle California,’ Palo
Alto (May), San Diego (Apr., May, June, July, Aug.,
Oct.), San Francisco.
The type of Ragonot’s humilis is merely a small,
rather dark male of fenestrella with identical genitalia.
In any considerable series of fenestrella the palpal differ-
ences cited by Ragonot can be observed. They are
indeed more apparent than real and more due to pro-
portionate differences in the sizes of the individual
specimens and to differing positions of the palps. In
our latest checklists humilis is listed as a subspecies or
variety of fenestrella. It is not even that.
349. Lipographis leoninella (Packard)
Pempelia leoninella Packard, Ann. New York Lyc. Nat. Hist.,
vol. 10, p. 259, 1873.
Nephopteryzx leoninella (Packard) Grote, Bull. U. S. Geol. Geogr.
Surv. Terr., vol. 4, p. 697, 1878.
Lipographis leoninella (Packard) Ragonot, Ent. Amer., vol. 5,
p. 115, 1889; Monograph, pt. 1, p. 565, 1893.—MceDunnough
Check list, No. 6273, 1939.
Lipographis fenestrella leoninella (Packard) Hulst, Phycitidae of
N. Amer., p. 167, 1890.
Pyla pallidella Dyar, Journ. New York Ent. Soc., vol. 12, p, 107,
1903.—Barnes and McDunnough, Contributions, vol. 3, p.
199, 1916 (make synonym of leoninella).
Forewing similar in maculation to that of fenestrella
except: General color more ocherous than gray, the gray
shading limited to the median area between the trans-
verse line; basal area of wing pale ocherous; the
inner border of the antemedial and outer border of the
subterminal lines yellow; lower fold between the trans-
verse lines pale ocherous; no appreciable black streaking
on the veins of outer area; discal spot at lower outer
angle of cell larger, more conspicuous. Hind wing paler,
with a faint ocherous tint towards apex and termen.
Alar expanse, 21-24 mm.
Male and female genitalia like those of fenestrella.
Type tocaities: California (leoninella, in MCZ);
Salt Lake, Utah (pallidella, in USNM).
Foop piant: Unknown.
Distrisution: Unitep States: California, ‘“Mid-
dle Calif.,”” Olancha (Inyo County, June), Palo Alto
(May), Sonoma County (May); Utah, Richfield (Aug.),
Salt Lake, Stockton (July), Vineyard (June, July).
Canava: Manitoba, Cartwright (Aug.).
Hulst treated leoninella as a variety of fenestrella, and
probably correctly. There is nothing to separate the
two except coloration. However, as nothing isknown
about their biology, it seems the better part of wisdom
to keep the two names apart. Larval characters and
habits and hosts may indicate separate species or at
least distinct races.
350. Lipographis truncatella (Wright), new combination
Figures 408, 898
Hypochalcia truncatella Wright, Ent. News, vol. 27, p. 25, 1916.—
McDunnough, Check list, No. 6276, 1939.
Forewing a dull, pale, brownish ocherous, dusted with
white and a fine peppering of black scales, making the
general color an ashy gray with a strong suffusion of the
ground color, the latter most pronounced in basal area
and in lower half of median area; antemedial line faint,
without any appreciable inner border, its outer border
indicated by black dots on costa, upper and lower veins
of cell, and on vein 1b; subterminal line distinct, with a
narrow, faint, but distinguishable and continuous inner,
black, bordering line. Hind wing pale gray, very faintly
tinted with ocherous toward base and shading into a
smoky hue towards apex and termen; a strong narrow
dark shade along termen. Alar expanse, 22-25 mm.
Male genitalia distinguished chiefly by its much
slenderer aedeagus, narrowing at apex, and the single,
very short, thornlike, curved cornutus on penis. Dif-
ferences also in the shape of the sclerotization of the
eighth segment tergite of abdomen are shown in the
figure. Female genitalia figured from a San Diego
specimen in the National Collection (W. S. Wright,
June 23, 1911). Bursa narrowly elongate; ductus bur-
168 UNITED STATES NATIONAL
sae much shorter than bursa, and narrow (no wider than
bursa, except at genital opening).
TypPE LocaLity: San Diego, Calif. (type probably
lost).
Foop prant: Unknown.
Distrisution: California, Chula Vista (June), San
Diego (June),
Despite its striking specific differences in genitalia
and more broadly scaled labial palpi this species fits
well into Lipographis. It is certainly not a Hypochal-
cia. The latter, an Old World genus, as far as I know
is not represented in our fauna.
351. Lipographis umbrella (Dyar), new combination
Ficures 410, 897
Sarata umbrella Dyar, Proc. Ent. Soc. Washington, vol. 10, p. 59,
1908.—McDunnough, Check list, No. 6270, 1939.
Male antenna with a weak ridge of roughened scales
in shallow sinus at base of shaft.
Forewing orange yellow; transverse lines narrow,
white; antemedial line oblique, somewhat curved, set
well out on wing and with only the faintest indication
of a dark outer border, the latter sometimes containing
a few black scales; subterminal line nearly straight, with
only a slight median bulge, inwardly more or less bor-
dered with black, the latter color varying from a thin,
weak line to large smudges extending well into the
median area of the wing; on some specimens a narrow
oblique blackish shade just beyond basal attachment of
wing; discal dots obscure, often obliterated by streaks
of white scaling or extensions of the black border of the
antemedial line. Hind wing semilustrous, ocherous
with a smoky suffusion, the latter most pronounced on
dark specimens; veins not appreciably darkened. Alar
expanse, 26.5-31 mm.
Male genitalia similar to those of fenestrella and
leoninella; differing from them chiefly in the armature
of the penis; the latter consists of a comb of 6 or 7 curved
spines and another straight spine, near but distinctly
separated from the comb. Female genitalia differing
only in minor details from those of fenestrella.
TYPE LOCALITY: San Diego, Calif. (type in USNM).
Foop prant: Unknown.
Distrisution: California, Laguna, Long Beach
(Sept.), Los Angeles (Sept.), Petaluma (Sept.), San
Diego (Aug., Sept.).
Dyar placed the species in Sarata on the basis of its
male antenna character; but its genitalia as well as the
lack of any sexual dimorphism in wing maculation or
color show that it belongs in Lipographis.
352. Lipographis (?) subosseella Hulst
Fieure 409
Iipographis subosseella Hulst, Canadian Ent., vol. 24, p. 62,
1893.—Ragonot, Monograph, pt. 1, p. 565, 1893.
Male antenna with a very weak scale ridge in shallow
sinus at base of shaft.
Thorax and forewing sordid white overshaded with
dull ocherous; the whitish ground color most noticeable
MUSEUM BULLETIN 207
along costa, the ocherous shade strongest along lower
fold and in outer area, making the general color of the
wing (to the naked eye) a pale brownish yellow; trans-
verse lines nearly obsolete; antemedial line distinguish-
able only as an oblique whitish streak from lower vein
of cell to inner margin, preceded on inner margin by a
blackish brown smudge; subterminal line indicated only
by its dark borders, a pale brownish, rather broad outer
band and a fainter, narrower, broken inner line; discal
dots separated, weak, blackish brown; a half-dozen
narrow blackish dots along termen. Hind wing white
with some fuscous shading at apex, on the outer veins,
and, narrowly, along termen; cilia shining white. Alar
expanse, 16 mm.
Male genitalia uncus triangulate, its apex narrowly
rounded. Apical process of gnathos an elongate, rather
narrow, veutrally flattened hook. Harpe elongate,
slender, with very small thornlike clasper. Anellus
without lateral lobes. Aedeagus broadly expanded at
apex; penis armed with a single, stout, curved cornutus
and a small supplemental detached sclerotized plate.
Kighth abdominal segment simple (without tufts).
TypE LocaLity: Bahama Islands (type in AMNH,
ex Rutgers).
Foop prant: Unknown.
An anomalous species, differing in its tropical habitat,
wing pattern, and many details of male genitalia from
other species of Lipographis, in which genus it is tenta-
tively retained. Eventually, when more material is
available, especially some female examples, a new ge-
neric reference may be needed. At present the species
is known only from its unique male type.
Genera 82-86: Adelphia to Acroncosa
[Venational division B. Forewing smooth; vein 2 from near lower
outer angle of cell; 4 and 5 separated at base. Hind wing with
7 and 8 closely approximate for a short distance from cell, rarely
(in individual specimens) weakly anastomosed; cell short, less
than one-half the length of cell. Female genitalia with signa
developed as opposed, strongly spined plates (except in Tota,
where bursa is smooth); ductus seminalis from bursa.]
82. Adelphia, new genus
Typr or Genus: Pempelia petrella Zeller.
Tongue well developed. Antenna pubescent; on male
a short, shallow sinus with small scale tuft in base of
shaft. Labial palpus oblique on male, upturned on
female, reaching to, but not appreciably above vertex;
second segment long, on male grooved to hold the maxil-
lary palpus; third segment minute, acuminate. Maxil-
lary palpus of male in the form of an aigrette; of female
squamous. Forewing smooth; 11 veins; vein 2 from
before but rather near lower outer angle of cell; 3 from
the angle, but little further at base from 2 than from 4;
4 and 5 separated at cell and divergent very shortly be-
yond it; 6 from below upper angle of cell, straight; 8
and 9 stalked for slightly less than half their lengths;
10 from the cell, approximate to the stalk of 8-9 for a
short distance from cell; male without costal fold. Hind
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 169
wing with vein 2 from well before lower outer angle of
cell; 3 from the angle connected with 4 by a short
spur; 4 and 5 contiguous or anastomosed for about
half their lengths; 7 and 8 closely approximate for less
than half their lengths; cell less than half the length of
wing; discocellular vein curved, produced outwardly at
lower angle. Eighth abdominal segment of male with
two pairs of ventrolateral hair tufts (the tufts stouter
and the hairs broadened and flattened in ochripunctella).
Male genitalia with uncus broad, hoodlike. Apical
process of gnathos a simple, strongly sclerotized hook.
Transtilla absent. Harpe elongate, slender; costa
strongly sclerotized on basal half; a strongly sclerotized
arm or hook arising from the lower edge of the sclero-
tized costa at base; cucullus narrow, tapering to a blunt
point. Anellus with short, weak lateral lobes. Penis
without cornuti or other appreciable armature. Vin-
culum stout.
Female genitalia with signa strongly developed, con-
sisting of two opposed, densely spined plates (in petrella
an additional collar of similar spines about the posterior
third of bursa); ductus bursae short, funnel-shaped,
sclerotized except at junction with bursa where it is
more or less finely scobinate; genital opening simple;
ductus seminalis from bursa at junction of the bursa and
ductus bursae.
This and the two genera following (Tota and Ufa) are
closely related to Elasmopalpus, and each contains a
species withdrawn from the latter genus. Such with-
drawal was necessary if Hlasmopalpus was to be ac-
curately defined. With its previous inclusions such
definition was impossible.
3538. Adelphia petrella (Zeller), new combination
Fieurss 411, 904
Pempelia petrella Zeller, Isis von Oken, 1846, p. 771; 1848, p. 886;
Verh. zool.—bot. Ges. Wien, vol. 22, p. 545, 1872.
Nephopteryx rubiginella Walker, List, vol. 27, p. 55, 1863.
Nephopteryz rufinalis Walker, List, vol. 27, p. 56, 1863.
Nephopteryx hapsella Hulst, Ent. Amer., vol. 3, p. 132, 1887.
Elasmopalpus petrellus (Zeller) Hulst, Phycitidae of N. Amer.,
p. 158, 1890.—Ragonot, Monograph, pt. 1, p. 421, 1893.—
Forbes, Cornell Mem. 68, p. 628, 1923.—McDunnough,
Check list, No. 6229, 1939.
Head and thorax reddish brown or brownish ocherous
with some gray shading on the mesothorax.
Forewing brownish gray with a fine white dusting in
median area especially toward costa; antemedial line
very faint, an irregular, interrupted, fine white line well
out on the wing, bordered outwardly by two or three
small black dots and preceded by a broad brownish
ocherous or reddish brown band; the inner edge of the
latter more or less shaded with black heavily dusted
with white; extreme base of wing shaded with blackish
gray; subterminal line usually distinct, narrow, and
finely dentate, white with a very weak dark inner border
except at costa but, on all well-marked specimens,
followed outwardly by some black streaklets on the
veins; discal dots separated, black, the lower one always
distinct, the upper weaker and sometimes obscured; a
row of small black dots along terminal margin set off
by a narrow dusting of white. Hind wing pale grayish
brown; the veins slightly darkened; on most specimens
a narrow smoky shade along outer margin. Alar ex-
panse, 20-27 mm.
Male genitalia with terminal margin of uncus rounded
and with a very slight notch at apex. Apical process
of gnathos a very stout hook. Projecting hook from
subcostal base of harpe a long, stout, nearly straight
arm, nearly as long as costa of harpe. Aedeagus not
appreciably widened towards apex.’ Female genitalia
with bursa considerably elongated, slender for over half
its length and with a collar of strong spines near its
middle.
Type tocauities: North America (petrella, in Mus.
Univ. Berlin); United States (rubiginella and rufinalis,
in BM); Florida (hapsella, in AMNH, ex Rutgers).
Foop prant: Unknown.
Distrisution: Florida, Coconut Grove, Glenwood,
Lake Alfred (July), Orlando (Feb., Mar., Apr.), St.
Petersburg (Feb.), Tampa, Vero Beach (Apr., June,
Sept., Oct., Dec.); Georgia (Feb., Mar.); Texas, Browns-
ville, San Benito, Victoria (Mar.); North Carolina,
Raleigh (Apr.), Southern Pines (Aug.), Tryon (May,
Aug.); Virginia, Virginia Beach (Aug.); District of
Columbia (May, July); New Jersey, Anglesea (May),
Woodbury (May); Jowa, Iowa City (July).
The Walker species (rubiginella and rujinalis) are
included in the synonymy on the strength of Ragonot’s
reference which is probably correct, for he presumably
examined their types. I have not. Hulst’s hapsella
agrees in genitalia and all other characters with typical
petrella. Its type is a female, not a male as stated by
Hulst. Honora obsipella Hulst is also listed as a
synonym of petrella in our lists, but incorrectly. It is
a synonym of Hulstia undulatella (see p. 196).
354. Adelphia ochripunctella (Dyar), new combination
Figures 412, 901
Salebria ochripunctella Dyar, Proc. Ent. Soc. Washington, vol.
10, p. 59, 1908 —MeDunnough, Check list, No. 6202, 1939.
Forewing mouse gray; the transverse lines obsolete
except for a faint indication of the antemedial line which
is oblique, nearly straight, and a dull pale ocherous gray;
a similarly faint ocherous discal spot at end of cell. Ex-
cept for these markings the wing is unicolorous. Hind
wing subpellucid, whitish with a faint smoky tint at
apex and narrowly along terminal margin; the veins
not appreciably darkened. Alar expanse, 17-21 mm.
Male genitalia with uncus slightly constricted towards
terminal margin, which is slightly concave; its lateral,
apical angles produced into short, ventrally projecting
lobes. Apical process of gnathos a small hook. Pro-
jecting hook from subcostal base of harpe, curved, about
half the length of costa of harpe. Aedeagus enlarged
(bulging) towards apex, and with some minute scobina-
8 The lateral projection from near apex of aedeagus shown in
our drawing (fig. 411a) is merely a partially sclerotized fragment
of the membrane connecting aedeagus and anellus and not a
projecting thorn or spine such as occurs in some species of Pyla.
170 UNITED STATES NATIONAL MUSEUM BULLETIN 207
tions on its outer surface (the latter are visible only
under high magnification and are somewhat exaggerated
in fig. 412a). Female genitalia with bursa much re-
duced as compared to that of petrella; armed only with
two opposed, spined plates.
TypE Locauity: San Diego, Calif. (type in USNM).
Foop piant: Eremocarpus setigerus. ‘This record
from an El Segundo specimen reared by W. D. Pierce.
Distrizution: California, El Segundo (Oct.), San
Diego (Oct., Nov.).
A distinct species easily identified by its peculiar
wing markings and male genitalia.
83. Tota, new genus
Typr or Genus: Elasmopalpus galdinella Schaus.
Characters of Adelphia except: Labial palpus con-
siderably longer, reaching well above vertex, especially
on the male. Hind wing with veins 4 and 5 stalked for
two-thirds of their lengths. Eighth abdominal segment
of male with compound ventral tufts. Male genitalia
with complete transtilla (a narrow, angulate, band);
aedeagus slender, elongate; penis armed with an elon-
gate, sclerotized plate bearing a row of six minute,
thornlike spines. Female genitalia with bursa simple,
without signa, spines, or other sclerotization; ductus
bursae narrow, tubular, sclerotized throughout its
length, not expanded or funnel shaped; genital opening
narrow.
The genus is distinguished from the others in this im-
mediate group by its genitalia and the rather long stalk-
ing of veins 4 and 5 of hind wing. Like the genus pre-
ceding (Adelphia) and that following (Ufa), it appears
to be closely related to Elasmopalpus, in which its type
and only known species was placed by Schaus.
355. Tota galdinella (Schaus), new combination
Fiaures 413, 899
Elasmopalpus galdinella Schaus, Zoologica, vol. 5, no. 2, p. 46,
Forewing blackish brown with the transverse lines
strongly contrasted, narrow, white; some scattered
white dusting in the subbasal and outer areas and, on
the male, appreciable white dusting over the medial
area; antemedial line oblique, irregularly dentate; sub-
terminal line zigzag, nearly vertical; no appreciable dis-
cal spots; a row of faint, separated, black dots along
terminal margin. Hind wing pale brownish (more
whitish on the male); the veins faintly darkened and a
smoky shade along outer margin. Alar expanse, 18-
24 mm.
Male genitalia with tegumen longer than greatest
width, its terminal margin straight and slightly pro-
duced at the lateral angles. Female genitalia with char-
acters as given for the genus.
Type Locatiry: Conway Bay, Indefatigable, Galé-
pagos Islands (type in USNM).
Foop puant: Unknown.
Distrisution: GauApagos Isuanps: “Camp Beta”
(Jan.), Conway Bay (Apr.), South Seymore (Apr.).
84. Genus Ufa Walker
Ufa Walker, List, vol. 27, p. 79, 1863.
venezuelalis Walker.)
Tongue well developed. Antenna pubescent; on
male with a short, shallow sinus with small scale tuft
in base of shaft. Labial palpus oblique in both sexes
(except in the female of rubedinella where it is some-
what upcurved); third segment nearly half the length
of second; extending well above vertex. Maxillary
palpi as in Adelphia. Venation as in Adelphia except:
Vein 3 of forewing on the average, closer to 4 than to 2;
hind wings with vein 4 and 5 anastomosed for about
half their lengths (or in some specimens of rubedinellus
and senta stalked for a trifle over half), cell one-third
the length of wing. Highth abdominal segment of male
with compound, ventral hair and scale tufts (except on
lithosella which has simple hair tufts).
Male genitalia without any sclerotized arm or hook
arising from base of costa of harpe; the costa strongly
sclerotized (in lithosella this sclerotized part of costa
terminating well before the apex of the harpe); clasper
present as a digitate or weak, rounded or triangulate,
erect projection from upper edge of sacculus. Anellus
a broad plate with short, lateral lobes. Aedeagus stout.
Penis armed with a single, stout, rather short, curved
spines.
Female genitalia similar to those of Adelphia; ductus
bursae short and broad, more or less funnel shaped,
with broad genital opening; ductus seminalis from
bursa well before junction of bursa and ductus bursae.
This genus, while sharing many of the characters of
Adelphia is closest to Elasmopalpus, from which it is
distinguished by its more oblique female labial palpi,
the strongly sclerotized costa of harpe, the shape of its
anellus, the stouter, much shorter apical projection of
gnathos, and the broader female ductus brusae.
(Type of genus: Ufa
356. Ufa lithosella (Ragonot), new combination
Fiaures 414, 903
Selagia lithosella Ragonot, N. Amer. Phycitidae, p. 9, 1887;
Monograph, pt. 1, p. 474, 1893.—Hulst, Phycitidae of N.
Amer., p. 160, 1890.—McDunnough, Check list, No. 6233,
1939.
Honora luteella Hulst, Journ. New York Ent. Soc., vol. 8, p. 223,
1900 p
Ancylostomia lithosella (Ragonot) Dyar, Ins. Insc. Menstr., vol.
7, p. 53, 1919.
Forewing pale ocherous; the ground color interrupted
by a narrow, paler, cream-colored shade along costal
edge, along lower margin of the cell, sometimes along
lower fold and (in a few examples) between the veins
in outer area; transverse lines obsolete; on some speci-
mens a dark grayish spot on vein 1b indicates what
remains of a dark outer margin to the antemedial line;
on occasional examples a dark grayish shade on middle
of inner margin; discal dots sometimes absent but the
lower one usually distinguishable, minute, blackish;
the usual dark dots along outer margin rarely dis-
tinguishable and when so only as a slight darkening of
the ground color. Hind wing semitransparent white
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 171
with a more or less pale smoky tint; the veins and
terminal margin not appreciably darkened, except on
the darkest specimens. Alar expanse, 26.5-32 mm.
Male genitalia with harpe very short, its costa
broadly and strongly sclerotized, the sclerotized costal
area terminating in an abrupt projection, well before
apex of cucullus. Lateral lobes of anellus knoblike.
Cornutus a short, stout, slightly curved, bluntly
pointed, hornlike thorn with a few minute serrate
projections on one side. Eighth abdominal segment
of male with a single pair of ventrolateral hair tufts.
Female genitalia with bursa armed with one large and
one small, round, strongly spined plate; the ductus
seminalis from bursa near the caudal margin of the
smaller plate; ductus bursae smooth except for a
slight, irregularly shaped sclerotization near genital
opening.
TypeE Locatities: Arizona (lithosella, in Paris Mus.);
Santa Rita Mts., Ariz. (luteella, in USNM).
Foop prant: Unknown.
Distrisution: Unirep States: Arizona, Baboqui-
vari Mts. (July, Oct.), Chiricahua Mts., Huachuca
Mts., Nogales (May), Palmerlee, Paradise (Cochise
County, Mar.), Santa Rita Mts. (June), Wilgus (Co-
chise County); New Mexico, Albuquerque. Mfxtco:
Cuernavaca (June), Durango (city), México (city, Oct.),
Venadio (Sinaloa), Zacualpin (June).
A distinct species, easily recognized by its genitalia
and obviously not closely related to Selagia where it
was placed by Ragonot, nor to Ancylostomia where it
was placed in the National Collection by Dyar.
357. Ufa roseitinctella (Dyar), new combination
Ancylostomia roseitinctella Dyar, Proc. U. S. Nat. Mus., vol. 42,
p. 105, 1912.
Forewing pale ocherous, the ground color suffused
with pale rose red broadly along inner margin and costa
and over most of median and outer areas; the ocherous
color on most specimens limited to the basal area and
(on a few examples) to a narrow, pale, longitudal shade
through the middle of the wing; transverse lines obso-
lete; discal dots minute, but usually distinguishable
(at least the lower one), blackish, separated; terminal
dots absent. Hind wing pale smoky fuscous, with a
very faint ocherous tint; the veins very slightly dark-
ened and a faint, dark line along termen. Alar expanse,
26-32 mm.
Female genitalia similar to those of lithosella.
Type tocauiry: Cuernavaca, México (type in
USNM).
Foop prant: Unknown.
Distrrisution: Unrrep Status: Arizona, Huachuca
Mts. (Sept.), Paradise (Cochise County, July, Oct.),
Pea Mts., Wilgus Mts. México: Cuernavaca
y).
This species is known only from females. It is very
close to and may only be a color form of lithosella, but
this cannot be determined until a male is recovered.
The Arizona specimens were in our collection under
(Selagia) Zamagiria australella (Hulst).
358. Ufa senta, new species
Fiecurss 415, 905
Forewing pale canary yellow with a narrow brownish
red shade along inner margin; costa pale on outer half,
whitish with a fine, sparse peppering of fuscous scales;
from apex inward along vein 6 a reddish fuscous streak
continued, on most specimens, as a faint, more or less
interrupted, dark shade along top of cell to base of wing;
lower discal spot minute, but usually distinct, dull red;
cilia peppered, ashy gray white. Hind wing translucent
white with a smoky shade towards outer margin; veins
in outer area of wing faintly darkened; a narrow fuscous
shade along termen. Alar expanse, 22-25 mm.
Male genitalia having harpe with costa sclerotized to
apex; clasper moderately long, slender, digitate. Anellus
with lateral lobes moderately long, strongly sclerotized,
tapering to blunt points. Cornutus a sharply curved,
strong, rather slender spine with a broad base.
Female genitalia with bursa rather narrowly elongate
with a lateral lobe near junction of bursa and ductus
bursa; signa consisting of two greatly elongated oval
lobes densely armed with long, slender spines; a second
cluster of minute spines adjacent to the posterior,
lateral lobe; ductus bursa short, triangulate, sclerotized
almost to genital opening.
Type tocauity: Big Bend, Tex. (type in USNM,
61363; paratype, co’, in BM).
Foop prant: Unknown.
Described from male type from the type locality
(Apr. 15, 1926); four male and one female paratypes
from Palmerlee, Ariz.; and one female paratype from
the Huachuca Mts., Ariz.
A distinct, easily recognized species, apparently near
to rubedinella Zeller.
359. Ufa rubedinella (Zeller), new combination
Ficurss 416, 902
Pempelia rubedinella Zeller, Isis von Oken, 1848, p. 885; Verh.
zool.-bot. Ges. Wien., vol. 24, p. 480, 1874; Horae Soc. Ent.
Rossicae, vol. 16, p. 181, 1881.
Acrobasis translucida Walker, List, vol. 27, p. 29, 1863.
Nephopteryx rufescentalis Walker, List, vol. 27, p. 58, 1863.
Nephopteryx minualis Walker, List, vol. 27, p. 60, 1863.
Nephopteryx deprivalis Walker, List, vol. 27, p. 60, 1863.
Ufa venezuelalis Walker, List, vol. 27, p. 60, 1863.
Elasmopalpus pyrrhochrellus Ragonot, Nouv. Gen., p. 23, 1888;
Monograph, pt. 1, p. 429, 1893 (new synonymy).
Elasmopalpus rubedinellus (Zeller) Ragonot, Nouv. Gen., p. 23,
1888; Monograph, pt. 1, p. 340, 1893.—Wolcott, Journ.
Agr. Univ. Puerto Rico, vol. 25, no. 2, p. 134, 1941.
Forewing a pale, glossy ocherous, more or less shaded
and marked with red or reddish brown; the females
usually darker and showing more of the reddish shading
than the males, some examples entirely suffused with
red except for a contrasted, whitish or very pale ocher-
ous shade along costa; three small but conspicuous
blackish or dark reddish dots in median area, two well
out from base, obliquely placed on lower vein of cell
and vein 1b, and one at lower outer angle of cell (on
some specimens indications of another dot at costa
opposite the one on vein ib and, very rarely, faint
172 UNITED STATES NATIONAL MUSEUM BULLETIN 207
traces of an upper discal dot at end of cell); from apex
a short oblique, reddish shade more or less peppered
with fuscous; faint tracings of red on the outer extrem-
ities of the veins and, on darker examples, a reddish
blush over the entire outer area; subterminal line in-
dicated by a narrow, oblique, twice-indented, red or
reddish fuscous line rather close to terminal margin.
Hind wings shiny, translucent white with a fine brown
line along termen; the veins not appreciably darkened.
Alar expanse, 16-22 mm.
Male genitalia with erect clasper of harpe short,
rounded, knoblike. Anellus with lateral lobes short,
stubby. Cornutus shaped like that of senta but appre-
ciably stouter. Terminal margin of vinculum slightly
produced at lateral angles. Female genitalia with
rather small narrow bursa containing a collar of fine
spines at junction of bursa and ductus bursae in addi-
tion to the normal] pair of spined signa; the broad, short
ductus bursae sclerotized throughout, its lower terminal
margin at genital opening sharply concave.
TYPE LOCALITIES: Brazil (rubedinella, in BM); Santo
Domingo (translucida and rufescentalis, in BM); Hon-
duras (minualis, in BM); Venezuela (deprivalis and
venezuelalis, in BM); Corrientes, Argentina (pyrrhochrel-
lus, in Paris Mus.).
Foop piants: Limabeans, black-eyed peas. These
records from Florida specimens reared by the Special
Survey of the Division of Foreign Plant Quarantine of
the U. S. Bureau of Entomology and Plant Quarantine.
Larvae feeding on the leaves.
DistRIBUTION.—UniTED States: Florida, Egmont
(Apr.), Hobe Sound (May), Hypoluxo, Lake Beach
(Feb.), Palm Beach, Palmetto, Vero Beach (Sept.,
Oct., Dec.), Winter Park (July). Cusa: Havana,
Matanzas (June), Santiago Province (Oct.). Domrni-
can Repusiic. Puerto Rico: Catafio July), Coamo
Springs (Apr.), Dovado (May), Isabela Substation
(Apr.), Palmas Abajas (Apr.), Puerto Real (Vieques Isl.,
Apr.), San German (Apr.). Viren Isuanps: Kingshill
(St. Croix, Mar., Apr., June). Jamaica. TRINIDAD:
Quare River Valley (Jan.). M*#x1co: Chiapas (May),
Guadalajara, Guerrero (Dec.), Oaxaca, Sierra de Gue-
rrero (Nov.), ‘Mexican Sub-region.” GUATEMALA:
Cayuga (Feb., Mar., Apr., May, June, Dec.), Chejel
(Aug.), Guatemala (city, Mar.), Quirigué (Mar.).
Honpuras. Costa Rica: Juan Vifias (Jan., June,
Dec.). Panamd: Alhajuela (Mar., Apr.), Corazal
(May, June, July), La Chorrera (Apr.), Obispo, Paraiso
(Apr., May), Porto Bello (Apr., May, Oct.), Taboga
Isl. (Jan., July), Tabernilla. Venrzunta. FRENcH
Gurana: St. Jean Maroni, St. Laurent Maroni. Bo-
Livia: Esperanza, Prov. del Saré (Dept. Santa Cruz,
May). Braz: Rio de Janiero (Nov.). Paraguay:
Villarrica (Jan., Mar., Oct., Nov., Dec.). AnrgEn-
TINA: Corrientes, Metin (Prov. del Salta, Feb.).
Pert: Chanchamayo. Generally distributed through-
out the tropical and subtropical regions of the New
World.
In many respects (its habitus, upturned female labial
palpi, sexual dimorphism, and similarity of larval
habit) this species seems to go with Elasmopalpus
lignosellus, with which it has been associated; but on
male and female genitalic characters it belongs definitely
with the species here assigned to Ufa, and on all larval
characters it is radically different from lignosellus. In
structural and pattern characters its larva resembles
that of Caristanius decoloralis.
85. Genus Elasmopalpus Blanchard
Elasmopalpus Blanchard, in Gay, Historia fisca y politica de
Chile. Zoologia, vol. 7, p. 104, 1852.—Hulst, Phycitidae of
N. Amer., p. 157, 1890.—Ragonot, Monograph, pt. 1, p. 418,
1893.—Forbes, Cornell Mem. 68, p. 628, 1923.—Janse,
Journ. Ent. Soc. South Africa, vol. 7, p. 4, 1944. (Type of
genus: Hlasmopalpus angustellus Blanchard.)
Tongue well developed. Antenna weakly pubescent;
on male a sinus and a short scale tuft in base of shaft.
Labial palpus of male erect and appressed to face, reach-
ing well above vertex; third segment minute, hidden in
scaling of second segment; of female upcurved, reaching
slightly above vertex; third segment but slightly shorter
than second, acuminate. Maxillary palpus of male in
the form of an aigrette; of female, squamous. Forewing
smooth; 11 veins; vein 2 from close to lower outer angle
of cell; 3 from the angle; 2, 3 and 4 approximately
equidistant at base (the position of 2 individually vari-
able, on some specimens very close and occasionally
connate with 3); 4 and 5 separated at base, approximate
for a short distance from cell; 6 from below upper angle
of cell, straight; 8 and 9 stalked for less than half their
lengths; 10 from the cell; male without costal fold.
Hind wing with vein 2 from well before lower outer angle
of cell; 3 from the angle, separated from 4 by a very
short spur; 4 and 5 stalked for at least two thirds of their
lengths; 7 and 8 closely approximate, contiguous or
weakly anastomosed fora very short distance from cell;
cell about one-third the length of wing; discocellular
vein curved, produced at lower angle. Highth abdom-
inal segment of male with compound ventral scale and
hair tufts.
Male genitalia with uncus rather narrowly triangu-
late, its apex narrowly rounded. Transtilla absent.
Apical process of gnathos a slender, elongate hook.
Harpe with costa not appreciably sclerotized; sacculus
with upper margin an erect, irregularly serrate ridge,
slightly produced at apex. Anellus U-shaped, with
lateral arms strongly sclerotized and produced as
curved pointed horns. Aedeagus slender; penis armed
with a single, slightly curved, slender, strongly sclero-
tized cornutus, from slightly more than one-third to
one-half as long as aedeagus. Vinculum stout, longer
than greatest width, tapering to narrowly rounded or
bluntly pointed terminal margin.
Female genitalia with signa consisting of two opposed,
strongly spined plates; ductus bursae cylindrical, longer
than bursa, sclerotized for half its length from junction
with bursa (the sclerotized part longitudinally ribbed),
membranous on posterior half, expanding to wide
genital opening, weakly sclerotized on inner dorsal
surface behind genital opening; ductus seminalis from a
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 173
lobe of bursa near junction of bursa and ductus bursae.
As here defined the genus is limited to its type
species. None of the other species that have hitherto
been assigned to Hlasmopalpus fits comfortably into the
genus.
360. Elasmopalpus lignosellus (Zeller)
Fiaures 33, 417, 906
Pempelia lignosella Zeller, Isis von Oken, 1848, p. 885; Verh. zool.-
bot. Ges. Wien, vol. 22; p. 544, 1872; vol. 24, p. 430, 1874;
Horae Soc. Ent. Rossicae, vol. 16, p. 180, 1881.—Riley, in
Rep. [U. 8.] Comm. Agr. for 1881, pp. 142-145, 1882.
Elasmopalpus angustellus Blanchard, in Gay, Historia fisca y
politica de Chile. Zoologica, vol. 7, p. 105, 1852.—Berg, Bull.
Soc. Imp. Nat., Moscou, p. 228, 1875; Anales Soc. Cient.
Argentina, vol. 4, pt. 4, p. 209, 1877 (makes synonym of
Pempelia lignosella).
Pempelia lignosella tartarella Zeller, Verh. zool.-bot. Ges. Wien,
vol. 22, p. 544, 1872; Horae Soc. Ent. Rossicae, vol. 16, p.
180, 1881.
Pempelia lignosella incautella Zeller, Verh. zool.-bot. Ges. Wien,
vol. 22, p. 544, 1872; Horae Soc. Ent. Rossicae, vol. 16, p.
180, 1881.
Pempelia lignosella major Zeller, Verh. zool.-bot. Ges. Wien, vol.
24, p. 430, 1874.
Elasmopalpus anthracellus Ragonot, Nouv. Gen., p. 23, 1888;
Monograph, pt. 1, p. 428, 1893 (new synonymy).
Dasypyga carbonella Hulst, Ent. Amer., vol. 4, p. 114, 1888.
Elasmopalpus lignosellus (Zeller) Ragonot, Ent. Amer., vol. 5, p.
115, 1889; Monograph, pt. 1, p. 425, 1893.—Hulst, Phyci-
tidae of N. Amer., p. 159, 1890.—Chittenden, U. 8S. Dep.
Agr. Div. Ent. Bull. 23, pp. 17-22, 1900.—Luginbill and
Ainslie, U. S. Dep. Agr. Bull. 539, 27 pp., 1917.—Forbes,
Cornell Mem. 68, p. 628, 1923.—Metcalf and Flint, Destruc-
tive and useful insects, ed 1, pp. 338-339, 1928; ed 2, pp. 367—
368, 1939.—McDunnough, Check list, No. 6231, 1939.—
Costa-Lima, Insetos do Brazil, p. 2, p. 93, 1950.—Craighead,
U.S. Dep. Agr. Misc. Publ. 657, p. 454, 1950.
Elasmopalpus puer Dyar, Ins. Insc. Menstr., vol. 7, p. 53, 1919
(new synonymy),
Ground color of forewing (on male) dull ocherous
fawn, transverse lines obsolete; on paler specimens a
subbasal black spot on vein 1b, one on lower vein of cell
beyond middle and another at lower outer angle of cell;
a row of faint, more or less confluent blackish dots along
terminal margin; a blackish shade of varying width
along costa and inner and outer margins; on females the
dark shading more extended, reducing the ocherous
ground color to a narrow, longitudinal shade, or com-
pletely suffusing the wing; many female examples en-
tirely black, with some occasional sparse reddish scaling
at extreme base of wing. Hind wing translucent white,
with a faint darkening of the outer veins and a narrow
brownish shade along terminal margin. Alar expanse,
16-24 mm.
Male and female genitalia: as given for the genus.
TYPE Locauities: Brazil (lignosellus, in BM); Rio
Negro, Brazil (anthracellus, in Paris Mus.); Concepcién,
Chile (angustellus, in Paris Mus.); Valparaiso, Chile
(major, in BM); Santiago Province, Cuba (pwer, in
USNM); Texas (tartarella and incautella, in BM; car-
bonella, in AMNH, ex Rutgers).
Foop piants: Beans, corn, cowpeas, chufa (Cyperus
exculentus), crabgrass, sudangrass, Johnsongrass, Japa-
nese cane, milo maze, sugar cane, sorghum, peanuts,
turnips, wheat, strawberry plants, flax, cotton, black
locust. Larvae boring into the stems of growing plants
and, to a lesser extent, feeding on the leaves.
Disrrizution.—Unitep States: Florida, Everglade
(Apr.), Fort Myers (Apr.), Hastings (Sept., Oct.), Lake
Alfred (July), Lakeland (Oct.), Miami (Oct.); Georgia,
Savannah (July); Alabama, Auburn (Aug.), Eufaula;
Lowsiana, Baton Rouge (July, Aug.), New Orleans
(June); Texas, Blanco County (Mar.), Brownsville
(Mar., June, Oct.), Burnet County (Mar.), College
Station, Dallas (Oct.), Dickinson (Oct.), Gainesville
(Aug.); Kerrville (Aug.), San Benito (Apr., June, July),
Smith Point (Sept.), Victoria (Apr., July); Arizona,
Baboquivari Mts. (Oct.), Mesa, (Aug.), Phoenix (Aug.),
Tempe (Oct.); California, San Diego (Aug.); South
Carolina, Columbia; Virginia, Cape Henry (June), Nor-
folk (Sept.); District of Columbia, Washington (Aug.,
Oct., Nov., Dec.); New Jersey, Montclair (Sept.);
Massachusetts, Cohasset (July, Aug.), Newton High-
lands. Brrmupa (Mar., Apr., May). Cusa: Havana,
Matanzas, Santiago de las Vegas (Apr.), Santiago
Province (Oct.). Purrro Rico: Anasco, Coamo Springs
(Apr.), Puerto Real (Vieques Isl., Apr.), Rio Piedras
(Aug.), San German (Apr.), Villalba (July). Viren
Isuanps: St. Croix (Apr.). Topsaco (June). JAMAIca.
México: Barmos (Sinaloa, Mar.), Chiapas, Colima
(Mar.), Guadalajara, Izuala (June), Mexico City (Nov.),
Orizaba, Tehuacin (May, June, Aug.). GUATEMALA:
Cayuga (Apr., June, July), Chejel (June, July, Aug.),
Guatemala City (Mar.), Perulhi (July), Quirigud
(Mar.), Volecén Santa Maria (Nov.), PanamA: Alha-
juela (Apr.), Corozal (May, July), La Chorrera (Apr.,
May), Paraiso (May), Porto Bello (Mar., Apr., May,
Sept., Oct.), Rio Trinidad (Mar.). VEnEzuEva: Las
Cruces, Colén (Dec.). Frencu Guiana: Cayenne, St.
Jean Maroni. Braziu: Castro (Parand), Rio Negro,
Santa Catarina (Oct.), Sdo Paulo (Sept.). Paraguay:
Paraguayan Chaco (Makthlawaiya, Nov., Nanahua,
Mar.), Villarrica (Feb., Sept., Oct., Nov.). ARGENTINA:
Tucumin (Nov.), Villa Cina (Feb., Mar., Nov.).
Curie: Valparaiso (Apr.). Prrt: Cafiete (Oct.). Gen-
erally distributed throughout the tropical and temperate
regions of the New World.
An insect of some economic importance in our
Southern States and known in economic literature as
the “lesser cornstalk borer.’”’ It is individually variable
in color and the sexes are markedly dimorphic. As a
result it has received several names. The latter have
no taxonomic value for they represent at most only
color forms. As far as I know the species is confined
to the New World, where it is abundant and widely
distributed.
The Luginbill and Ainslie paper (1917) gives all avail-
able information on the life history of the species.
There has been no contribution of any importance since
its publication.
174 UNITED STATES NATIONAL MUSEUM BULLETIN 207
86. Genus Acroncosa Barnes and McDunnough
Acroncosa Barnes and McDunnough, Canadian Ent., vol. 49,
p. 404, 1917. (Type of genus: Acroncosa albiflavella Barnes
and McDunnough.)
Tongue well developed. Antenna simple except for
a few minute thornlike projections on base of shaft of
the male (no appreciable sinus or scale tuft); very
shortly and weakly pubescent in both sexes. Labial
palpus short, slender, oblique, third segment shorter
than second, blunt, projected slightly forward; not
reaching to height of vertex. Maxillary palpus
squamous. Forewing smooth; 11 veins; vein 2 from
before but near lower outer angle of cell; 3 from the
angle; 4 and 5 separated at base; 6 from below upper
angle of cell, straight; 8 and 9 stalked for half or a
trifle over half their lengths; 10 from the cell, separated
at base from 8-9 and not approximate to its stalk;
male without costal fold. Hind wing with vein 2 from
well before lower outer angle of cell; 3 from the angle
and approximate to 4 for a short distance beyond the
angle; 4 and 5 contiguous or anastomosed beyond the
angle for about half their lengths (superficially, on
undenuded specimens, they appear long stalked); 7 and
8 closely approximate for a short distance from cell;
cell less than half the length of wing; discocellular vein
curved, considerable produced outwardly at lower angle.
Eighth abdominal segment of male with a pair of ventro-
lateral hair tufts (the hairs somewhat flattened).
Male genitalia with uncus broadly triangulate, its
apex bluntly pointed. Transtilla incomplete, its ele-
ments moderately sized, irregularly triangulate plates.
Harpe simple; costa sclerotized nearly to apex, not
produced. Anellus a shallow U-shaped plate with small
lateral lobes. Aedeagus long, stout; penis armed with
masses of spines varying from long (one-third the length
of aedeagus) to minute. (Figs. 418c-e show the shape
and character of the longer spines, greatly enlarged.)
Vinculum stout, longer than greatest width, tapering
to truncate terminal margin.
Female genitalia with signa consisting of two small,
opposed, strongly spined, round plates; bursa otherwise
membranous except for a thickening of the lobe giving
off the ductus seminalis; ductus bursae long, slender,
tubular, unsclerotized except for a couple of weak spine
clusters near junction with bursa; genital opening
simple; ductus seminalis from bursa near its junction
with ductus bursae.
Foretibia with a long inner and short outer claw. I
should suspect this character to be merely specific
except that it is equally developed in both sexes of both
species of the genus.
361. Acroncosa albiflavella Barnes and McDunnough
Ficurss 35, 418, 907
Acroncosa albiflavella Barnes and McDunnough, Canadian Ent.,
vol. 49, p. 405, 1917.—McDunnough, Check list, No. 6151,
1939.
Forewing white; the transverse white lines lost in the
ground color; antemedial line indicated only by a broad
oblique inner orange band extending from inner margin
to costa; subterminal line indicated by a similar,
narrower, sinuate, outer orange band, the latter termi-
nating at costa in a small blackish spot; a well-con-
trasted black discal dot at lower, outer angle of cell and
a smaller black dot on inner margin of the subbasal
orange band at vein 1b; on same specimens a few
widely scattered black scales on the white ground color
of median area. Hind wings whitish with a very faint
ocherous or smoky tint; the veins not appreciably
darkened. Alar expanse, 18-21 mm.
Genitalia as given for the genus.
Type Locatiry: Loma Linda, San Bernardino Coun-
ty, Calif. (type in USNM).
Foop piant: Unknown.
Distrisution: California, Loma Linda (July, Aug.),
Mona Lake (July).
362. Acroncosa albiflavella castrella Barnes and McDunnough
Acroncosa albiflavella castrella Barnes and McDunnough,
Canadian Ent., vol. 49, p. 405, 1917.
A local race of albiflavella differing only in having a
stronger peppering of black scales in the median area of
forewing. Alar expanse, 20-22 mm.
Genitalia like those of typical albiflavella.
Type tocatity: Fort Wingate, N. Mex. (type in
USNM).
Foop prant: Unknown.
pe usenuaeNs New Mexico, Fort Wingate (July,
ug.).
Apparently a distinguishable local race, but can be
identified only by locality label and a somewhat
stronger dusting of black scales on forewing.
363. Acroncosa similella Barnes and McDunnough
Acroncosa similella Barnes and McDunnough, Canadian Ent.,
vol. 49, p. 405, 1917——MceDunnough, Check list No. 6152,
1939.
Forewing similar to albiflavella except: A much
heavier peppering of black scales on forewing; orange
inner border of antemedial line extending only from
inner margin to top of cell; a concentration of black
scaling forming a more or less broken line along outer
margin of the antemedial line; a similar black line
inwardly bordering the subterminal line; basal, median,
and outer areas dusted with black scales. Alar expanse,
21-26 mm.
Genitalia like those of albiflavella.
Type LocaLity: Pyramid Lake, Nev.
USNM).
Foop Prant: Unknown.
Distrisution: Nevada, Pyramid Lake; Utah, Eureka
(June).
Not structurally distinct from and possibly only a
local race of albiflavella. The incomplete subbasal
orange band on forewing, however, suggests a distinct
species.
(type in
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 175
Genus 87: Passadena
[Venational division B. Forewing with weak subbasal scale
ridge; veins 4 and 5 shortly stalked; 10 from the cell. Hind
wing with vein 2 near angle of cell; 3 from the stalk of 4-5; 4 and
5 long stalked. Labial palpus short, porrect. Male genitalia
with a strongly developed clasper on harpe. Female genitalia
with signa developed as opposed, strongly spined plates; ductus
seminalis from bursa.]
87. Genus Passadena Hulst
Passadena Hulst, Canadian Ent., vol. 32, p. 171, 1900.
genus: Passadena constantella Hulst).
Tongue well developed. Antenna weakly pubescent;
on male with a shallow sinus and scale tuft in base of
shaft. Labial palpus short, porrect; second segment
oblique, the third deflected forward, minute, more or
less hidden in the thick scaling of second segment; the
latter grooved on male to hold the maxillary palpus.
Maxillary palpus of male in the form of an aigrette; of
female, squamous. Forewing with weak subbasal
ridge of raised scales; 11 veins; vein 2 from before but
near lower outer angle of cell; 3 from the angle; 4 and 5
shortly stalked; 6 from below upper angle of cell,
straight; 8 and 9 stalked for about half their lengths;
10 from the cell; male without costal fold. Hind wing
with vein 2 from near lower outer angle of cell; 3 from
the stalk of 4-5; 4 and 5 long stalked (for over two-
thirds their lengths; 7 and 8 approximate or contiguous
for a very short distance from cell; cell about half the
length of wing; discocellular vein curved, outwardly
produced at lower angle. Eighth abdominal segment
of male with a weak pair of ventrolateral hair tufts.
Male genitalia with uncus triangulate; apical process
of gnathos a strong, narrow, elongate hook. Transtilla
absent. Harpe with costa narrowly sclerotized for its
entire length; a strong, projecting clasper from upper
edge of sacculus near base (the clasper a striking feature;
its peculiar shape probably a specific character). Penis
armed with several short rows of weak spines with a
mat of fine scobinations between them. Vinculum
stout, stubby, slightly broader than long, terminal mar-
gin broadly truncate.
Female genitalia with signa developed as a pair of
opposed, strongly spined plates, one (ventral) rather
large, elongately oval, the other (dorsal) very small and
bearing only two or three spines; ductus bursae short,
broad, sclerotized for most of its length, and bearing on
midventral surface a row of minute spines; genital open-
ing simple; ductus seminalis from bursa near junction
of bursa and ductus bursae.
A distinct, monotypical, New World genus with one
western North American species.
(Type of
364. Passadena flavidorsella (Ragonot)
Figures 36, 419, 908
Anoristia flavidorsella Ragonot, N. Amer. Phycitidae, p. 9,
1887.—Hulst, Phycitidae of N. Amer., p. 160, 1890.
Meroptera canescentella Hulst, Phycitidae of N. Amer., p. 149,
1890.—Ragonot, Monograph, pt. 1, p. 319, 1893.—MceDun-
nough, Check list, No. 6186, 1939. (New synonymy.)
Getulia flavidorsella (Ragonot), Monograph, pt. 1, p. 528, 1893.
Passadena constantella Hulst, Canadian Ent., vol. 32, p. 171, 1900.
Megasis cinctella Hulst, Canadian Ent., vol. 32, p. 172, 1900 (new
synonymy).
Passadena cinctella (Hulst) Barnes and McDunnough, Contribu-
tions, vol. 3, p. 198, 1916.—MeDunnough, Check list, No.
6222, 1939.
Passadena flavidorsella Barnes and McDunnough, Check list of
the Lepidoptera of Boreal America, No. 5647, 1917.—
McDunnough, Check list, No. 6221, 1939.
Forewing whitish gray, more or less finely dusted with
black scales; transverse lines narrow, whitish; ante-
medial line somewhat oblique, twice indented, bordered
outwardly by a narrow black line, inwardly by a broader
black band of roughened scales, the black borders form-
ing a conspicuous black band divided by a narrow pale
line; subterminal line sinuate-angulate, bordered in-
wardly by a black line (most strongly accented at
costa) and outwardly at costa by a short black streak;
discal dots faint, only distinct on specimens with a pale
(whitish) ground color; a row of inconspicuous black
dots along termen. Hind wing white to pale smoky
fuscous, frequently with a slightly smoky shade towards
apex; veins not appreciably darkened; a fine brownish
line along outer margin. Alar expanse, 15-20 mm.
Genitalia as given for the genus. The male can be
readily identified by the peculiar shape of the clasper
of harpe (fig. 419e).
Type Locauitigs: Arizona (flavidorsella, in Paris
Mus.); Texas (canescentella, in AMNH, ex Rutgers);
California (constantella, in AMNH, ex Rutgers); Argus
Mts., Calif. (cinctella, in USNM).
Foop puant: Unknown.
Disrrisution: Texas, Brownsville (May), “Central
Texas” (the types of canescentella, 2 and o’, without
date); New Mexico, state locality only (July, Cockerell,
collector); Arizona, Ajo (Pima County, Mar.), Babo-
quivari Mts. (Apr., May, July, Aug., Sept.), Catalina
Springs (Apr.), Oracle (July), Phoenix (Apr.), Reding-
ton, Sells Post Office (Pima County, Apr.), “Salt River
Mts.” (Sept.), “Southern Arizona” (Aug.), Tempe
(Feb., May), Tucson (May), Wenden (Yuma County,
Aug.), Yuma County (‘Colorado Desert,’ Mar.);
California, Argus Mts. (Apr., May), Inyo County
(June, July), Jacumba (June), La Puerta (July), Los
Angeles (May), Mason Valley (San Diego County,
Apr.); Utah, Richfield (May). Also reported by Rago-
not from Sonora, México.
The species is somewhat variable in the ground color
and amount of blackish dusting on forewing, some
specimens being considerably paler than others; but
the pattern markings are constant, the most conspicu-
ous of which is the broad black subbasal band with
roughened scales, formed by the borders of the ante-
medial line.
The female type of canescentella Hulst in the Rut-
gers Collection lacks an abdomen, as does the matching
male paratype in the National Museum (originally from
the Fernald Collection). Both specimens are rubbed;
but the pattern characters are plainly distinguishable
and their palps, antennae, and venation are unmistak-
ably those of Passadena.
176 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Genus 88: Ulophora
[Venational division B. Forewing with subbasal scale ridge;
veins 4 and 5 approximate for a short distance from cell. Hind
wing with vein 2 from well before angle; 4 and 5 stalked for about
half their lengths; cell short (less than one-third the length of
wing). Male antenna simple. Labial palpus uncurved, long,
slender. Maxillary palpus squamous. Male genitalia with
transtilla incomplete or absent; penis armed with a longitudinal
row of short, slender spines.]
88. Genus Ulophora Ragonot
Ulophora Ragonot, Ann. Soc. Ent. France, ser. 6, vol. 10, Buil.,
p. vii, 1890; Monograph, pt. 1, p. 155, 1893.—Forbes,
Cornell Mem. 68, p. 619, 1923. (Type of genus: Ulophora
grotezz Ragonot.)
Acromeseres Dyar, Ins. Insc. Menstr., vol. 7, p. 41, 1919. (Type
of genus: Acromeseres dialithus Dyar. New synonymy.)
Tongue well developed. Antenna simple; shaft
somewhat broadly flattened towards base on male but
without sinus or scale tuft or thornlike spines, on female
threadlike, weakly pubescent on both sexes. Labial
palpus upcurved, reaching well above vertex, slender,
smoothly scaled; third segment about one-third the
length of second, acuminate. Maxillary palpus squa-
mous, more broadly so on male than on female. Fore-
wing with subbasal ridge of raised scales; 11 veins; vein
2 from before but rather near lower outer angle of cell;
3 from the angle, at base nearer to 4 than to 2; 4 and 5
approximate at base and for a very short distance
beyond; 6 from below upper angle of cell, straight; 8
and 9 stalked for slightly more than half their lengths;
10 from the cell, nowhere approximate to the stalk of
8-9; male without costal fold. Hind wing with vein 2
from well before lower outer angle of cell; 3 from the
angle connate with or very closely approximate to the
stalk of 4-5 at base; 4 and 5 stalked for about half their
lengths; 7 and 8 closely approximate for a short dis-
tance from cell; cell slightly less than one-third the
length of wing; discocellular vein curved, not appreci-
ably produced at lower angle. Highth abdominal
segment of male with a weak pair of ventrolateral hair
tufts.
Male genitalia with uncus triangulate, its apex
narrowly rounded. Apical process of gnathos a stout,
curved hook. ‘Transtill2 incomplete or absent (its
elements, when distinguishable, very small and weakly
sclerotized). Harpe simple; costa broadly but weakly
sclerotized; clasper vestigial. Aedeagus straight; penis
armed with a longitudinal row of short, slender,
straight spines (16 to 20, the number individually vari-
able). Vinculum stout, considerably longer than great-
est width, but slightly tapered to rounded terminal
margin.
Female genitalia without signa; bursa with a mat of
fine scobinations at and near junction of bursa and
ductus bursae, otherwise smooth and membranous;
ductus bursae longer than bursa, slender, tubular,
unsclerotized, genital opening simple, narrow; ductus
seminalis from middle of bursa.
A distinct genus of uncertain affinities; contains one
North American and one very closely related neo-
tropical species.
365. Ulophora groteti Ragonot
Ficure 420
Ulophora groteit Ragonot, Ann. Soc. Ent. France, ser. 6, vol. 10,
Bull., p. vii, 1890; Monograph, pt. 1, p. 156, 1893.—Hulst,
Phycitidae of N. Amer., p. 222, 1890.—Forbes, Cornell
Mem. 68, p. 619, 1923.—McDunnough, Check list, No.
6117, 1939.
Ulophora tephrosiella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 107, 1904.—Forbes, Cornell Mem. 68, p. 619, 1923.—
MeDunnough, Check list, No. 6118, 1989. (New synonymy.)
Forewing brown dusted with white on basal and
median areas and narrowly along terminal margin,
giving the paler areas an ashy gray appearance and ac-
centing the brown shade along costa and bordering the
transverse lines; the latter narrow, faint, silvery scaled ;
antemedial line at middle of wing, oblique, somewhat
sinuate-dentate, bordered outwardly by a narrow brown
line and inwardly by a broad brown band, the inner
edge of which forms a stout raised-scale ridge extending
from inner margin to top of cell, the inner margin rather
strongly dusted with white; subterminal line sinuate,
with rather broad inner and outer brown borders; discal
dots obscure, more or less confluent, brown; the terminal
dots confluent, forming a fine brown line along outer
margin. Hind wing pale to dark smoky brown; the
veins somewhat darker; a fine dark brown line along
termen. Alar expanse, 11-19 mm.
Male genitalia distinguished from those of the species
following (guarinella) chiefly by the broader harpes and
more narrowly rounded apex of uncus. The number
and length of the spines on penis is an individual charac-
ter. Our figure 420a shows the minimum in number
and the maximum in length. Female genitalia like
those of guarinella.
Typr LocaLities: North Carolina (grote, in Paris
Mus.); Washington, D. C. (ephrosiella, in USNM).
Foon eiant: Tephrosia spp. Larvae feeding on pods
and seeds.
DistrisutTion: New Jersey, Woodbine (Sept.); Dis-
trict of Columbia (July, Aug.); Virginia, Pendleton
(Aug.), Skyland (July); North Carolina, Southern Pines
(June, July, Aug., Sept.), Tryon (July, Aug.); South
Carolina, Clemson College (July); Georgia, Atlanta
(June), Spalding County (Nov.); Florida, Glenwood,
Lake Alfred (May, July), Lakeland (Sept.), Polk
County (Aug.), Tampa (June), Winter Haven (July);
Alabama, Auburn; Louisiana, Vernon Parish (Aug.);
Texas, Herne, Sandflat.
Dyar’s tephrosiella was described from small speci-
mens. ‘There are no structural or color differences to
distinguish these from typical grotew. The larva lacks
the sclerotized rings around seta IIb of mesothorax and
seta III of the eighth abdominal segment, normally
characteristic of phycitid larvae.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE Al¥pye
366. Ulophora guarinella (Zeller)
Ficures 421, 909
Myelois guarinella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p.
208, 1881.
Ulophora guarinelia (Zeller) Ragonot, Monograph, pt. 1, p. 155,
1893.
Acromeseres dialithus Dyar, Ins. Insc. Menstr., vol. 7, p. 42, 1919.
(New synonymy.)
Forewing decidedly paler than that of grotei and,
except for the band preceding the antemedial line, an
almost uniform pale gray; the transverse lines obscure;
antemedial line bordered inwardly by an orange-red
band with more or less black shading on its lower half
and a black scale ridge along its inner edge; subterminal
line nearly obsolete, indicated chiefly by a very faint,
narrow, dark gray inner border; discal dots more or less
obscured, when distinguishable, blackish and either
separated or coalesced; terminal dots faint, separated.
Hind wing whitish with a faint brownish or smoky tint
towards apex and along outer margin; the veins very
slightly darkened. Alar expanse, 13-18 mm.
Male genitalia are similar to those of groteii except
for narrower harpes and a more bluntly pointed uncus.
The female genitalia show no distinguishing characters.
TypPE Locatitizes: Honda, Colombia (guarinella, in
BM); Santiago, Cuba (dialithus, in USNM).
Foop piant: Unknown.
Distrisution: Cusa: Santiago (Jan., Oct.), Baracoa.
Cotompia: Honda.
In structure, color, and maculation Dyar’s dialithus
is a perfect match to the male type of guarinella. The
species is very close to groteii but apparently distinct.
Nothing is known of its life history.
Genus 89: Chorrera
[Venational division B. Forewing smooth; veins 3, 4 and 5 equi-
distant and approximate towards base. Hind wing with vein 2
well before angle; 4 and 5 stalked for over half their lengths.
Male genitalia with transtilla incomplete; aedeagus needlelike;
penis unarmed; vinculum with pair of anteriorally produced
lateral lobes from terminal margin. Female genitalia developed
as two elongate, narrow, opposed plates, armed with short, longi-
tudinally arranged spines; ductus bursae slender, globularly ex-
panded near genital opening.]
89. Genus Chorrera Dyar
Chorrera Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 330, 1914.
(Type of genus: Chorrera idiotes Dyar.)
Tongue well developed. Antenna weakly pubescent;
on male a slight scale tuft on shaft shortly beyond basal
segment, no appreciable sinus. Labial palpus obliquely
ascending, reaching to slightly above vertex, slender;
second segment roughly scaled; third less than one-half
the length of second, acuminate. Maxillary palpus
squamous. Forewing smooth; 11 veins; vein 2 from
before but rather near lower outer angle of cell; 3 from
the angle; 3, 4 and 5 equidistant and approximate
towards base; 6 from below upper angle of cell, straight ;
8 and 9 stalked for half or less than half their length;
10 from the cell; male without costal fold. Hind wing
with vein 2 from well before outer angle of cell; 3 from
the angle; 4 and 5 stalked for over half their lengths
(very long stalked in extrincica) ; 7 and 8 closely approxi-
mate for a short distance beyond cell; cell less than one-
half the length of wing; discocellular vein curved, con-
siderably produced at lower angle. Eighth abdominal
segment of male with a pair of very weak ventrolateral
hair tufts.
Male genitalia with uncus rather narrowly triangu-
late, tapering abruptly at apex. Apical process of
gnathos a short, stout hook. Transtilla incomplete, the
divided elements small. Harpe short, with strongly
sclerotized, thornlike, more or less appressed clasper,
about half as long as harpe. Anellus a shallow U- or
V-shaped plate with somewhat produced lateral lobes.
Aedeagus, long, slender (needlelike), slightly curved;
penis unarmed. Vinculum stout, about twice as long
as greatest width; slightly tapering to a moderately
broad terminal margin with a pair of anteriorly pro-
duced lateral lobes.
Female genitalia with signa developed as two
elongate, narrow, opposed plates armed with short,
stout, thornlike spines, longitudinally arranged; bursa
otherwise membranous, large, more or less pear shaped;
ductus bursae for half its length from bursa, very
narrow, ribbonlike, sclerotized and bent at middle,
globular and membranous beyond, and thence triangu-
larly expanding into the simple genital opening; ductus
seminalis from bursa near its junction with ductus
bursae.
The genus is certainly distinct and easily distin-
guished from other phycetine genera by its peculiar
genitalia. It contains what appear to be three tropical
American species. These may eventually prove to be
no more than races of a single variable species, but at
present we are not justified in such a grouping. We
know nothing of their life history and our knowledge of
their distribution is too fragmentary to permit more
than speculation as to their status.
367. Chorrera idiotes Dyar
Ficures 34, 422, 914
Chorrera idiotes Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 331,
1914.
Forewing gray with a fine dusting of white scales,
making the ground color a dark ashy gray; transverse
lines narrow, faint, whitish; antemedial line slightly
oblique, sinuate-angulate, with a narrow, obscured,
outer edging of black scales; subterminal line somewhat
more distinct, sinuate, with narrow inner and outer
dark borders, pronounced and black at costa; discal
dots very faint, when distinguishable, black and sepa-
rated; a row of faint, confluent, blackish dots along
terminal margin. Hind wing translucent white; a dark
smoky shade along costa and a narrower smoky shade
along terminal margin; the veins not darkened except
178
UNITED STATES NATIONAL
(on some females) at their marginal extremities. Alar
expanse, 15.5-19 mm.
Male genitalia with a short knoblike projection from
base of clasper of harpe. Lateral lobes of anellus
short, stubby. Female genitalia as given for the genus.
Typ tocauity: La Chorrera, Panama (May; type
in USNM).
Koop prant: Unknown.
Known only from the type locality.
368. Chorrera extrincica (Dyar), new combination
Figures 423, 913
Rhodophaea extrincica Dyar, Ins. Insc. Menstr. vol. 7, p. 41, 1919.
Similar to idiotes except: Forewing with less whitish
dusting; the transverse lines and their dark borders more
obscure; ground color a nearly uniform brownish gray
(fuscous). Hind wing with narrower smoky borders
along costa and terminal margin; veins 4 and 5 longer
stalked (4 very short and on some specimens vestigial or
altogether absent). Alar expanse, 13-15 mm.
Male genitalia with clasper of harpe a simple hook (as
in postica); lateral lobes of anellus longer and more
slender. Female genitalia with appreciably smaller
bursa and entire genitalia shorter.
TYPE LOCALITY: Santiago, Cuba (type in USNM).
Foop prant: Unknown.
Known only from the type locality (May, June, Oct.).
Represented in the National Collection by 12 males and
2 females.
369. Chorrera postica (Zeller), new combination
Fieure 424
Myelois postica Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 213,
1881.
Nephopteryx postica (Zeller) Ragonot, Monograph, pt. 1, p. 292,
1893.
Similar to idiotes except: Thorax and basal two-
thirds of forewing heavily dusted with white, making
ground color an ashy white as far on forewing as the
dark gray transverse shade extending from costa at
beginning of subterminal line to outer third of inner
margin. Hind wing without dark shade along costa
and with only a faint, narrow, dark shade on inner
Margin at apex. Alar expanse, 18 mm.
Type LocaLity: Honda, Colombia (type in BM).
Foop prant: Unknown.
Known only from the type locality.
Genus 90: Tacoma
[Venational division B. Forewing with veins 4 and 5 stalked for
nearly half their lengths. Hind wing with vein 2 near lower angle
of cell; 4 and 5 long stalked; cell about one-third the length of
wing. Antenna of male simple. Labial palpus upturned, slen-
der. Maxillary palpus squamous. Male genitalia with apical
process of gnathos a pair of converging bladelike projections;
transtilla absent; harpe simple; penis unarmed. Female genitalia
with signa developed as two plates armed with long, curved,
clawlike spines; ductus bursae from middle of bursa, long, slender,
membranous; ductus seminalis from bursa remote from junction
of ductus bursae and bursa.]
MUSEUM BULLETIN 207
90. Genus Tacoma Hulst
Tacoma Hulst, Ent. Amer., vol. 4, p. 115, 1888; Phycitidae of
N. Amer., p. 139, 1890.—Ragonot, Monograph, pt. 1, p. 205,
1893. (Type of genus: Tacoma feriella Hulst.)
Tongue well developed. Antenna simple. Labial
palpus upturned, cylindrical, slender, reaching to vertex;
third segment one-third the length of second, acuminate.
Maxillary palpus squamous. Forewing smooth; 11
veins; vein 2 from before but rather near lower outer
angle of cell; 3 from the angle; 4 and 5 stalked for
slightly less than half their lengths; 6 from below upper
angle, straight; 8 and 9 stalked for a half or a trifle more
than half their lengths; 10 from the cell; male without
costal fold. Hind wing with vein 2 from before but near
lower outer angle of cell; 3 from the angle, connate with
the stalk of 4-5; 4 and 5 long stalked (for over two-thirds
their lengths); 7 and 8 closely approximate for a short
distance from cell; cell about one-third the length of
wing; discocellular vein curved and outwardly produced
at lower angle of cell. Eighth abdominal segment of
male simple.
Male genitalia with uncus triangulate, its apex bluntly
rounded. Apical process of gnathos a pair of converg-
ing, flattened, pointed, bladelike projections. Trans-
tilla absent. Harpe simple, narrowly elongate. Anel-
lus a simple shield. Aedeagus simple, straight; penis
unarmed. Vinculum stout, subtriangulate with nar-
rowly rounded terminal margin; about as long as
greatest width.
Female genitalia with bursa shaped like an elongate
potato, with ductus bursae arising from its middle;
signa developed as two strongly spined plates, one of
irregular shape armed with numerous, slender, long,
curved spines and situated at junction of bursa and
ductus bursae, the other a narrow curved band with a
row of well spaced, strong, curved, clawlike spines
along one edge; also in bursa, several (6 or more) small,
detached, curved, thornlike spines; ductus bursae much
longer than bursa, slender, membranous throughout;
genital opening simple, small; ductus seminalis from
bursa remote from junction of bursa and ductus bursae.
A monotypical genus of unmistakable distinctness,
easily identified by its genitalia. Except for their
genitalia the sexes are not distinguishable on any
external characters. The spining of the female abdo-
men mentioned by Hulst (1890) is nonexistent. It is
impossible to conceive what he saw or thought he saw,
for there is no such structure on either the female or the
male.
370. Tacoma feriella Hulst
Ficures 37, 425, 912
Tacoma feriella Hulst, Ent. Amer., vol. 4, p. 115, 1888; Phycitidae
of N. Amer., p. 189, 1890.—Ragonot, Monograph, p. 1, p.
205, 1893.—McDunnough, Check list, No. 6142, 1939.
Tacoma submedianella Dyar, Ins. Insc. Menstr., vol. 1, p. 34,
1913.—McDunnough, Check list, No. 6144, 1939. (New
synonymy.)
Forewing gray, with an irregularly intermixed pow-
dering of black and white scales, the black predomi-
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 179
nating, making the over-all color dark gray with a faint
bluish tint; the transverse lines grayish white, faint
but distinguishable; antemedial line oblique, narrow
from costa to lower margin of cell, thence (on most
specimens) more or less expanded into a conspicuous
white blotch of variable but usually quadrate shape;
on some examples the white blotch is reduced and on a
few completely absent; in the latter the antemedial is a
narrow white line throughout, notched below costa and
at lower fold and with a complete, narrow, outer, black
border; on examples with the antemedial line expanded
into a white blotch, the blackish outer border is broken
or obliterated below the cell; subterminal line inwardly
notched at veins 6 and 1b, shaded inwardly by a
narrow, blackish border; discal dots, when distinguish-
able, confluent, forming, on well marked specimens, a
narrow black line along discocellular vein; terminal dots
more or Jess confluent, usually forming a blackish line
along outer margin. Hind wing whitish with a more
or less smoky brown tint; veins faintly darkened and a
narrow brown shade or line along outer margin. Alar
expanse, 15-20 mm.
Genitalia as given for the genus.
Types.—In Rutgers University (feriella); U. S.
National Museum (submedianella)
Type Locauitins: Texas (feriella, in AMNH, ex
Rutgers) ; La Puerta, Calif. (submedianella, in USNM).
Foop piant: Mistletoe. This record from speci-
mens from Riverside, Calif., reared by Commander
Dammers.
Distribution —Tezas, Blanco County (Sept.), Burnet
County (Sept.), Kerrville, Plano (Aug., Sept.), Rio Frio
(May); New Mexico; Mesilla Park (May); Arizona,
Fish Creek Station (Sept.), Redington, Santa Rita Mts.
(June), Wilgus; California, La Puerta (July), Mount
Lowe (July), Riverside (July, Sept.).
As indicated by the foregoing description the species
is individually variable. Dyar’s submedianella was
described from rather small California examples in
which the characteristic white patch over the lower
half of the antemedial line was partially or wholly
obliterated by dark scaling. In the series before me
there is every intergrade between the form with a
conspicuous white patch and that without it; the
extremes are not peculiar to any locality, nor do they
exhibit any genitalic differences.
180
Group II
[Hind wing with vein 3 present, 4 absent.]
K&rys TO THE VENATIONAL DIVISIONS AND GENERA OF GROUP II
Forewing with 11 veins; vein 2 from the cell . . . . . Venational division A (key, p. 180)
Forewing with 11, 10, or 9 veins; vein 2 stalked or united with 3.
Venational division B® (key, p. 184)
Forewing with 10 veins; veins 8 and 9 ee 4 and 5 stalked; 2 from the cell or [Azaera]
from the stalk of 4-5 . . ..... =... . Venational division C (key, p. 184)
Forewing with 10 veins; 8 and 9 stalked; 4 and 5 united; 2 from the cell.
Venational division D (key, p. 185)
Forewing with 9 veins; 8 and 9 united"; 4 and 5 united; 2 and 3 from the cell.
Venational division E (key, p. 185)
Forewing with 9 veins; 8 and 9 united; 4 absent; 3 and 5 stalked; 2 from the cell.
Venational division F (key, p. 186)
Venational division A
1. Cell of hind wing partially open; only a fragment of the discocellular vein distinguishable.
Psorosina (p. 208)
Cell closed; discocellular vein complete and curved ............... 2
2. Hind wing with vein 2 from lower outer angle ofcell ........2..2.2...8
Hind wing with vein 2 from before the angle. .......... 8
3. Forewing with vein 2 from the lower outer angle of wall: 2 emabnnte Meae Hoses (. 196)
Forewing with vein 2 from before the angle ava ;
4, Forewing with vein 2 from well before the angle; ein Albdamizel seamnent a wt
without hair tufts; transtilla absent (elements not distinguishable); ductus bursae
sclerotized for most of itslength ... . ... . . Ilatila (p. 263)
Forewing with vein 2 from near lower angle; sielnian opdbmeadl segment of male with
hair tufts; transtilla present Gameenied at least by Sh a: elements) ;
ductus mene membranous. . . 2 sree
5. Labial palpus upturned; maxillary wala a walle squamous; Sema a feansail a de-
pressed, small cluster of blunt, stout, thornlike spies . . . Drescomopsis (p. 262)
Labial palpus oblique; maxillary palpus 2 male in the form of an a signum or
signa of female otherwise .. . 6
6. Male genitalia with apical process ae eaminos on annie aie! hooting a sinelll anes
signa of female consisting of a chain of thornlike spines on bulbous bases and a
scattering of similarspines . . . . .. . . Cabotia (p. 200)
Male genitalia with apical process of anailias a gett. ieae hook; signa of female
consisting of large, round, densely spined plates... .....-...+.+--7
9 The genus Nonia in this division could easily be confused on forewing venation with the species and
genera of division F. However, in Nonia it is vein 2 that is absent (united with 3) and 4 and 5 that are
stalked; while in group F vein 2 is present, 4 absent, and 3 stalked with 5, two radically different developments
to the same end result. The genitalia of Nonia show its close relationship to forms with veins 2 and 3 stalked
and vein 2 tending to disappear.
10Tn occasional specimens of Vitula a vestige of vein 9 may be present on one forewing or the other,
but the normal condition is for vein 9 to be absent.
11 Some specimens of Bema show a trace of vein 9 on one side or the other, but this is an abnormal
condition.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
7. Male antenna with a stout hook from basal segment of shaft; tufts on eighth abdominal
segment of male compound; transtilla incomplete. . ..... Oncolabis (p. 199)
Male antenna with shaft simple; eighth abdominal segment of male with paired tufts;
transtila completes {exam Ae Qo. Ole ons aatels| De Honorinus (p. 199)
8. Veins 3 and 5 of hind wing approximate at base (or contiguous for a short distance
IexOViCl) eS * OE eS comes ee REN rte) ap artes were aeccee arn CCRT (25. | 9
Veins 3 and 5 of hind wing connate (rarely very shortly stalked)... ...... 15
Veins 3 and 5 of hind wing appreciably stalked. . ............2.2.. 24
0.2 Maxillary palpus!squamoush Saver a? HERA iO emo inode ww sige: yao
Midxillary: palpus«filiform™ fein... .Oha 80% enohei ane & gieveliin.geivew. 12
10. Labial palpus upturned; shaft of male antenna with sinus and scale tuft at base . .11
Labial palpus porrect; shaft of male antenna simple. ....... Patriciola (p. 209)
11. Forewing with vein 10 stalked with 8-9; eighth abdominal segment of male with com-
pound! ventrallituitsy, Pars HO Sl aries. Bilge’? vn. Canarsia (p. 201)
Forewing with vein 10 from the cell; eighth abdominal segment of male with paired
ventrolateral ttufts@ TiRG% 10 ort 20). beeccrohaia 308 Teciey. Paconius (p. 210)
12. Apical process of gnathos bifid or produced as large, partially fused lobes; ductus bursae
of female sclerotized only at genital opening . ............2... 13
Apical process of gnathos a short, blunt hook; ductus bursae sclerotized for most of its
length SO FReRIOVIO Die geen Je loletog. Uso old arost Ani awed’ eo. 14
13. Antennal shaft of male simple; ductus seminalis of female from middle of bursa; signum
BbsenG) SH SLND Dc a meh Yr Pet og er BILE BULA CRO, ae Aptunga (p. 211)
Antennal shaft of male with some rough scaling in a shallow sinus towards base; ductus
seminalis from anterior end of bursa; signum present .... . Baphala (p. 235)
14. Labial palpus porrect; eighth abdominal segment of male simple . . Volatica (p. 290)
Labial palpus oblique; eighth abdominal segment of male with paired dorsal tufts.
Vezina (p. 291)
15, Antennae bipectinate (in both sexes) ; veins 7 and 8 of hind wing closely approximate. . 16
Antennae pubescent; veins 7 and 8 of hind wing anastomosed. . ......... 17
16. Labial palpus of male porrect; bursa of female without signum . . . Melitara (p. 240)
Labial palpus of male obliquely ascending; bursa of female with signum . Olycella (p. 241)
17. Veins 7 and 8 of hind wing anastomosing beyond cell for not more than half the length
Ofsvenn’ 72511)! Uy Teme ee ae eae ate lo, wees la@eahnn wileliosysolalt. . 18
Veins 7 and 8 anastomosing for most of their lengths beyond cell. . . . ..... 21
18. Cell of hind wing not more than half thelength of wing. ............ 19
Cell of hind wing definitely more than half thelength of wing .......... 20
19. Maxillary palpus filiform; antennal shaft of male with modified, papillalike setae on basal
segments; female withsignum .......2...2..2... Zophodia (p. 238)
Maxillary palpus squamous; antennal shaft of male simple; female without signum.
Olyca (p. 243)
20. Maxillary palpus squamous; male with transtilla complete, forewing without costal
Folds AR NUR eae Mea BOSC, Chl Lees 5 Ona Euzophera (p. 272)
Maxillary palpus filiform; male with transtilla incomplete, forewing with costal fold.
Eulogia (p. 275)
2ishabialipalpus' upturned! OuhaMnt (io Asie), WDE Ames eh ep Geet loekneeliok . 22
Labial palpus oblique or obliquely ascending. . ...........0.52285 23
22. Forewing with vein 10 shortly stalked with 8-9; male with transtilla incomplete, but
elements considerably enlarged. . . ...........4. Moerbes (p. 268)
Forewing with vein 10 from the cell (usually separated, rarely connate, at base, with
8-9); male with transtilla complete. .......2.2... Moodnopsis (p. 269)
23. Cell of hind wing less than half (more than one-third) the length of cell; forewing of male
with costal fold; eighth abdominal segment of male with paired ventrolateral hair
tufts, |. . REN RR pow to Ru.BOLOR GAO, Bots Entmemacornis (p. 266)
Cell of hind wing more than half the length of cell; forewing of male without costal fold;
eighth abdominal segment of male simple .......... Exuperius (p. 274)
181
182
24.
25.
26.
27.
28.
29.
30.
31.
32.
33.
34.
35.
36.
37.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Hind wing with cell one-third the length of wg. ......... Edulica (p. 271)
Hind wing with cell approximately half the length of wmg. ........... 25
Penis of male armed with two stout thornlike cornute; signa of female consisting of two
large, opposed plates armed with long stout spines, a scattering of similar spines
between the plates, and a band of shorter spines at junction of bursa and ductus
bursae.. 3 SAAD APUG RN 00, oe, SO Ae eve Adelperga (p. 187)
Penis of male smooth or armed only with weak spines or sclerotized wrinklings; signa of
female (if present) otherwise; iacthiieny Hiasoenene yaihae de ie ee eae 26
Forewing with vein 2 from lower outer aele of cell... .scocieupe sued eee 2 27
Forewing with vein 2 from before the angle ...........2.2.4..... 28
Forewing with veins 2 and 3 closely approximate at base; males with strong costal fold on
forewing, eighth segment with paired ventrolateral hair tufts, transtilla complete.
Cassiana (p. 212)
Forewing with veins 2 and 3 connate; male without costal fold, eighth abdominal segment
without hair tufts, transtilla complete. ............ Rioja (p. 267)
Hind wing with veins 7 and 8 anastomosed for most of their lengths beyond cell. . . . 29
Hind wing with veins 7 and 8 approximate, contiguous or anastomosed beyond cell; but if
anastomosed, then for less than two-thirds the length of vein7........ 31
Forewing with vein 10 stalked with 8-9. ............ Cayennia (p. 267)
Forewing with vein 10 from the cell, separated at base and divergent from stem of 8-9 . 30
Male forewing with costal fold; eighth abdominal segment of male simple; female genitalia
with ductus seminalis from bursa between signum and ductus bursae. Selga (p. 265)
Male forewing without costal fold; eighth abdominal segment of male with paired ventro-
lateral hair tufts; female genital with ductus seminalis from anterior end of
bursa .. .. .okpeu. alu to duensies lininobds Mia sasinoe Anderida (p. 211)
Male genitalia with apical process of Briathos greatly reduced and fused into subanal
plate; female genitalia with signum a single, long, stout, curved spine.
Lascelina (p. 264)
Male genitalia with apical process of gnathos well developed; female genitalia with sig-
num (if present) otherwise .*.)) 020.) ee eee we a ale. 32
Male genitalia with apical process of gnathos developed as a stout, elongate hook; female
genitalia with ductus seminalis from ductus bursae. ........2.... 33
Male genitalia with apical process of gnathos usually flanged or lobed, frequently bifid,
sometimes fused or partially fused, if completely fused (Cahela, Rumatha) consider-
ably enlarged SESE female pominle with ductus seminalis from bursa copulatrix.
40
Hapa palpi jporrect enor eins: oo tele dete ene eee atis/lod ventana & 34
Labial palpi oblique or upturned . . . . ~~... 2 ee eee eee ee wes 38
Highth abdominal segment of male with ventrolateral hair tufts and lateral eversible
HOES ee ee se AES Toh OM a Sey foe) cr se Ae eae mea 35
Eighth abdominal segment of male without tufts or lobes, ventrolateral hair tufts only
(no eversible lobes)... .... AU MLE ie ricco Geo MRI SNE cb eS - Cet ex%h'c 36
Maxillary palpus of male filiform. ............. Macrorrhinia (p. 190)
Maxillary palpus of male in the form ofanaigrette. ......... Ocala (p. 191)
Aedeagus of male of nearly even width throughout; female genitalia with ductus bursae
simple, and ductus seminalis from ductus bursaenearitsmiddle ........ 37
Aedeagus of male abruptly tapered from base and very slender therefrom; female geni-
talia very slender for half its length from genital opening, and thence to bursa
swollen and spiraled, armed throughout the spiraled portion with a line of fine, sharp
BPINES WOM: «, s. a. s,s tee sy oy op AMOReEO< Roots pene Valdivia © 191)
Kighth abdominal segment of male with a pair of short ventrolateral hair tufts; harpe
with clasper present, developed as a strongly sclerotized, nearly straight hook; Gamal
genitalia with ductus bursae as long as, or longer than, bursa. . . Eumysia (p. 187)
Eighth abdominal segment. of male without hair tufts; harpe without clasper; ductus
bursae considerably shorter than bursa. . .......... Protasia (p. 193)
38.
39.
40.
41.
42.
43.
44.
45.
46.
47.
48.
49.
50.
51.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Maxillary palpus of male squamous; female genitalia with an acutely bent, sclerotized
elbow in ductus bursae before genital opening. ......... Hulstia (p. 195)
Maxillary palpus of male filiform or in the form of an aigrette; female genitalia with
duchus bursacistraightmyin ye meee. Nem eos Cleese wlio, nilasinon oll . 39
Maxillary palpus of male filiform; female genitalia with ductus bursae long (as long or
longer -thantbursa) loge anne, Oly i) SaBotTR) wees Heterographis (p. 193)
Maxillary palpus of male in the form of an aigrette; female genitalia with ductus bursae
shorti(much'shorter than’ bursa)iiyili-ieteedesey es ee Staudingeria (p. 194)
Tongue considerably reduced, exposed or completely hidden by labial palpi . . . .41
Tongue well developed and exposed.) WS" 20. NON S RpY Ow Raggio. , 43
Hind wing with veins 7 and 8 anastomosed for at least half their distances from cell;
female genitalia with ductus seminalis from anterior end of bursa copulatrix.
Laetilia (p. 230)
Hind wing with veins 7 and 8 very shortly anastomosed (for less than half their distances
from cell); female genitalia with ductus seminalis from bursa near its junction with
ductus*bursae% 24 os = MAY Sho Le Sopte aay Mane pital bo te? , 42
Male genitalia with apical process of gnathos partially fused; female genitalia with
SISTA Oe AIRE DILL) OLS 1) Soe A PaaS Cactoblastis (p. 245)
Male genitalia with apical process of gnathos bifid; female genitalia without signum.
Eremberga (p. 253)
Mieixill aneya Al MSPETLOMUN voy co ws iee 5) aso) lay cis) ce! ea ah “Fes Semen heh Fomeepe Subd alemaey ot Boy ot 44
Maxillary palpus squamous, fan shaped or flamboyant ..........2... 45
Labial palpi of both sexes porrect; eighth abdominal segment of male simple.
Rhagea (p. 237)
Labial palpi of male upturned, of female oblique; eighth abdominal segment of male
with paired ventrolateral hair tufts. ........... Cactobrosis (p. 260)
Maxillary palpus flamboyant (extending above front). ...... Sigelgaita (p. 255)
Maxillary palpus fan shaped and held vertical to the face. . . .. Yosemitia (p. 250)
Maxillary palpus: squamous « «+ oo: e06 ws SEY YS Oo Roy Ca pyroqon » 46
Shaft of male antenna pubescent and with a series of modified papillalike setae on several
basal segments; female with labial palpus obliquely ascending . . Ozamia (p. 257)
Shaft of male antenna pubescent (but without modified setae on basal segments) or
pectinate; female with labial palpus porrect or obliquely porrect . ...... 47
Vinculum of male long (distinctly longer than broad) ; female genitalia without signum. 48
Vinculum short (little, if any, longer than broad); signum present ........ 50
Maxillary palpus extending well above middle of face; eighth abdominal segment of male
SUING Ve ter ir eR RE EN Scr. Wi cat Nanaia (p. 245)
Maxillary palpus not extending above middle of face; eighth abdominal segment of male
Withyventrolateral ham itifisgss webie AioelVsuisins: serusswsturn sf ideue a thustienee eater ° 49
Antenna of male pubescent; ductus seminalis of female from near anterior end of bursa;
bursaveryismiallis raieeesik See ee LO Ss ee SE EIN Salambona (p. 254)
Antenna of male bipectinate; ductus seminalis of female from middle of bursa; bursa
lareeig wrt MRR Ley nee 1s el ce SS AL iy, a, Amalafrida (p. 256)
Antenna of male unipectinate; (female unknown) ........ . Parolyca (p. 255)
Antenna of male bipectinate; ductus seminalis of female from middle of bursa.
Alberada (p. 244)
Antenna of male pubescent; ductus seminalis from bursa towards (near) junction of
bursa with ductussbursnewe vis eit) Fe Le to ee dn tatu Hebe tat 51
Habitat, South America; male genitalia with apical process of gnathos bifid.
Tucumania (p. 252)
_ Habitat, southwestern United States and northern México; male genitalia with apical
process\of gnathos uscd ee AL iin ha paie, chink anny © cios diies entwehait « 52
183
184
52.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Male genitalia with aedeagus moderately long and sclerotized throughout, harpe without
subbasal sclerotized pocket, anellus with base of plate narrowly sclerotized; female
with dark markings on forewing longitudinal, no discal spot . . . Cahela (p. 247)
Male genitalia with aedeagus short and partially sclerotized, harpe with subbasal sclero-
tized pocket, anellus with base of plate broadly sclerotized; female with dark mark-
ings of forewing transverse (at least in part), discal spot prominent.
Rumatha (p. 248)
Venational division B
. Forewing with veins 2 and 3 of forewing stalked see
Forewing with veins 2 and 3 of forewing united. . 6
. Forewing with veins 4 and 5 stalked. . +8
Forewing with veins 4 and 5 united . 5 8
. Forewing with veins 8 and 9 stalked. Lie cceucsie FF fo way Ciara acl ae anes e cy regenera
Forewing with veins 8andQ9 united. ......... . . . Micromescinia (p. 277)
. Cell of forewing open (no trace of disease vein). . .. . . . Gennadius (p. 277)
Cell of forewing closed . ..........2... . . . Mescinia (p. 212)
. Forewing with veins 8 and 9 stalked; hind wing with 3 and 5 iptolleed:
Eurythmasis (p. 203)
Forewing with veins 8 and 9 united; hind wing with 3 and 5 closely approximate at base.
eirephonyegints © on
, Hindi wing with yeins)3 and aystalked -)) sq-Uem ane y-li se ieciielsinlii mele
Hind wing with veins 3 and 5 closely approximate or connate at base. . .... . :
. Cell of forewing open (no trace of discocellular vein) . . .. . . . Farnobia (p. 276)
Cell of forewing closed . ...-.. 2... +22 - 2s e Ge Sige test wpa t att gee
. Cell of hind wing open (no trace of discocellular vein). . . . . . . Wunderia (p. 204)
Cell of hind wing closed. ............ ... . . .Hurythmidia (p. 204)
. Forewing with veins 8 and 9 stalked... ....... =... . . Phestinia (p. 216)
Forewing with veins 8and9 united. ........=...... . Nonia (p. 215)
Venational division C
. Hind wing with discocellular vein straight and vertical . . . . . . . BY, Sea,
Hind wang with’ discocellular vem curved: s.) -ete) -n sien eee cee et rauem
Horewine with) veins 4 and orstalkcedss 5). ee) sien tere mete nny nts) ee eee
Forewing with veins 4and 5connate.............. Patagonia (p. 225)
. Male genitalia with a comb of strong spines along side of aedeagus; female genitalia with
two large signa composed of opposed, cup-shaped, strongly spined, concave plates;
ductus seminalis from ductus bursae. .........-... Rotruda (p. 225)
Male genitalia with aedeagus smooth or with only minute scobinations at apex; female
genitalia with signum a single, coarsely spined plate, or absent; ductus seminalis
fromm bursa.’ Oyen rita En hiak aia SSE L are MPA reat sy te goed HgnivedsOmnA (p. 219)
. Hind wing with veins 3 and 5 approximate at base. .... . Pataca are sory itty scat te 5
Hind wing with veins 3 and 5 connate orstalked. ..............-.7
. Cell of hind wing one-third the length of wing Breed iin) TELE ORE SRE tee 6
Cell of hind wing one-fourth the length of wing. .......... Gvenna (p. 288)
. Eighth abdominal segment of male with compound diossall tufts; shaft of male antenna
SIMPLE ce ee eee. nee, Nhe Sty eae Tet Cot Baran ae ce Ree ata Vitula (p. 285)
Eighth abdominal segment of male simple; shaft of male antenna with shallow smus
towardstbases tee Tee gee ay SRE Dinteeiee eo, erie haere eee Moodnella (p. 289)
. Hind wing with vein 2 oe the lower outer angle of cell. . . . . Harnocha (p. 202)
Hind wing with vein 2 from before theangle. .........-+-+-+-+-+-+-+-s 8
. Hind wing with vein 2 near lower outer angle of cell. . . 2... . +++ ---- 9
Hind wing with vein 2 from well before the angle... .........2-.--. 10
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 185
9. Forewing with vein 3 separate from, connate with, or rarely (in individual specimens)
very shortly stalked with the stem of 4-5 . . . . . .Ephestiodes (p. 278)
Forewing with vein 3 from the stalk of 4-5 (well stalked maith it) . . . Azaera (p. 282)
10. Cell of hind wing approximately one-third the length of wing. . . 2478) Tal
Cell of hind wing slightly more than one-half the length of wing . . Vagobanta (p. 289)
11. Male with transtilla incomplete; shaft of antenna with shallow sinus towards base.
Moodna (p. 283)
Male with transtilla complete; shaft of antenna simple. . . . . . Manhatta (p. 287)
Venational division D
1. Hind wing with discocellular vein straight and vertical; cell very short (about one-fifth
the length of wing) . . . . Metephestia (p. 264)
Hind wing with dintdealittanevein Gaeved: call tonger ania slightly less than one-third
to one-half the length of wing) . SM tee eee ee ae et ee
2. Hind wing with vein 2 from lower outer ariel of cell Pili re Cee ale at ee ote:
Hind wing with 2 from before the angle. ... . . 4
3. Veins 7 and 8 of hind wing contiguous for some chesney hepend eet (but sa: werent
sing) ; labial palpus upturned; antennal shaft of male simple . Oedothmia (p. 205)
Veins 7 and 8 anastomosed beyond cell; labial palpus oblique; shaft of male antenna
with a hook from its basal segment and deeply curved for several segments beyond.
Stylobasis (p. 205)
4, Hind wing with vein 2 from near the angle ofcell. . 2. 2. 2... 2 ee
Hind wing with vein 2 from well before the angle. ....... 6
5. Labial palpus porrect; male penis without armature. .. . tes cD iaitiasts es 189)
Labial palpus upturned; penis armed with two stout, straighit cornuti. Diviana (p. 206)
6. Hind wing with veins 3 and 5 shortly stalked. . . . . . . . Prosoeuzophera (p. 275)
Hind wing with veins 3 and 5 connate or very closely approximate at base. . . . . 7
7. Labial palpus oblique, extending above vertex of head; veins 7 and 8 of hind wing closely
approximate for half their distance beyond cell. . . . . . . . Palatka (p. 207)
Labial palpus upturned, short, not reaching vertex; veins 7 and 8 anastomosed for most
of their distance beyond cell. . . . . . . 2... . . . Cacozophera (p. 208)
Venational division E
1. Hind wing with discocellular vein straight and vertical. . . . . .. . 2
Hind wing with discocellular vein straight and oblique. . ...... Beay (pr 217)
Hind wing with discocellular vein curved. . . Sako
2. Cell of hind wing less than one-fourth the wing length! Gali ious dude fold. on fore-
wing, with a weak pair of ventrolateral hair tufts on eighth abdominal segment.
Comotia (p. 217)
Cell of hind wing about one-third the wing length; male with costal fold, but with eighth
segment SIMplO oes wwe news Unaaiitia Grr)
3. Hind wing with vein 2 from near foere Peer snes ‘of ene . . . . Nicetiodes (p. 304)
Hind wing with vein 2 from well before angle of cell. . . . 4
4, Male with apical process of gnathos enlarged (broadened), ped ted: septic or fonaede
female with ductus seminalis from bursa near its junction ae ductus bursae.
Sosipatra (p. 294)
Male with apical process of gnathos otherwise; female with ductus seminalis from bursa
well removed from the junction of bursa ate ductus bursae. . . . 5
5. Labial palpus upturned; ductus bursae of female membranous for a or ‘all of its
length, not flattened. . . 6
Labial palpus oblique or parece actus ipa of feats peleronized fOr a eonsidersbio
Pant Onis lenrthvandanatroneds: 4. 2 secttucetae © easy. eee ee eet
300329—56——_13
186
10.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
. Male with transtilla complete or its elements greatly enlarged and approximate at their
apices; forewing with a strong costal fold; eighth abdominal segment with com-
pound dorsal tufts; female with apophyses of ovipositor and intersegmental area
between ovipositor and eighth-segment collar, short. . . . . . Ephestia (p. 301)
Male with transtilla incomplete, its elements well separated at their apices, not enlarged;
forewing without costal fold; eighth segment with paired dorsal hair tufts; female
with apophyses of ovipositor and intersegmental area between ovipositor and
eighth-segment collar, verylong. .........2.+.2... Anagasta (p. 299)
. Male genitalia with apical process of gnathos asec’: ductus seminalis of female
from very near anterior end of bursa. . ............ Ribua (p. 297)
Male genitalia with apical process of gnathos svmmetmieal: ductus seminalis of female
from middle or near middle of bursa. . . .......2-.-..2.+-..4:+-.-- 8
. Labial palpi of both sexes porrect. .......-..2...-. . . . . Plodia (p. 298)
Labial palpi of both sexes oblique. . .......2.2.2..4.++++e48020448-8 9
. Tongue reduced; alarexpanse8mm.. ........... ‘ Caan (p. 294)
Tongue well devdlaei: alar expanse 10mm.ormore. . . . area 1(0)
Bursa of female with signum (antenna of male with a sheila: sinus in | ghett jonas
DESC), koe tin Biyeye tic BeBe 3 Ae ass te iis . . Caudellia (p. 292)
Bursa of ennel’s without Siac, Gosia imino) - SRY eo . . Bethulia (p. 296)
Venational division F
. Male genitalia with a projecting spur from base of costa of harpe; apical process of gna-
thos U-shaped; female with ductus bursae sclerotized and flattened for most of its
lenethy ee 2 oe se ye) SRR BEEET FAT OUW. BEOSD AUR cranes Varneria (p. 305)
Male genitalia with ane! pela from costa of harpe near or -teyand middle; apical process
of gnathos (if present) not U-shaped; female with ductus bursae membranous and
tubular throughout... ............. 2
. Male with eighth abdominal segment simple; gnathos ammina fae | ma very dee cou
narrowly forked projection; aedeagus with bluntly pointed apex; penis pened sili
an elongate, spiraled, suleviiiza and spined plate; female genitalia with signa ar-
ranged as a series of (3 to 6) detached, broadly based, thornlike spines and narrow-
bladed disks situated near anterior end of bursa and a cluster of several smaller
spines near junction of bursa and ductus bursae. . . . . . . Eurythmia (p. 306)
Male with compound dorsolateral tufts on eighth abdominal segment; gnathos without
apical projection, its arms fusing at their distal ends into the sclerotized subanal
plate; aedeagus with apex enlarged and flaring; penis unarmed except for a few sclero-
tized wrinklings; signa of female consisting of a single elongate series of (5 to 14)
thornlike spines... .....-++-++.-. ....... . Erelieva (p. 308)
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Genera 91 and 92: Adelperga and Eumysia
[Venational division A. Forewing with 11 veins; 10 from cell,
8 and 9 stalked, 2 and 3 from the cell, 4 and 5 stalked or connate.
Hind wing with veins 7 and 8 closely approximate or contiguous;
2 from close to lower, outer angle of cell; discocellular vein
curved. Harpe of male genitalia with an elongate, strongly
sclerotized clasper; gnathos terminating in a stout, short or
moderately long, hooked process; transtilla incomplete; vinculum
stout, as broad or broader than long.]
91. Adelperga, new genus
Tyrer or Genus: Heterographis cordubensiella Ragonot.
Tongue well developed. Antenna simple, pubescent.
Labial palpus obliquely upturned, reaching vertex,
third segment short, projected forward. Maxillary
palpus squamous. Forewing with vein 2 from or from
very near outer angle of cell; 2 and 3 approximate at
base; 4 and 5 stalked; 6 straight from below upper
outer angle of cell; 10 from cell, closely approximate to
8-9; male without costal fold. Hind wimg with vein 2
from very close to angle of cell; 3 and 5 stalked; 7 and 8
approximate; cell less than half (more than one-third)
the length of the wing; discocellular vein curved.
Eighth abdominal segment with a pair of short ventro-
lateral hair tufts.
Male genitalia with apical process of gnathos a rather
short hook. Uncus broadly and bluntly rounded at
apex. Harpe with a decided incurvation between sac-
culus and cucullus; clasper present, developed as a
long, strongly sclerotized and apically curved and
swollen arm, projecting into the incurvation between
sacculus and cucullus. Anellus semitubular. Aedea-
gus long, stout, cleft and strongly sclerotized towards
apex; penis armed with a pair of short stout thornlike
cornuti and numerous granulations.
Female genitalia with bursa copulatrix thickened and
sclerotized at junction with ductus bursae; signa
strongly developed, consisting of two large, opposed
plates armed with long, stout spines, a scattering of
similar spines between the plates, and a band of shorter
spines where ductus and bursa join; ductus bursae
short, flattened, very broad and sclerotized throughout;
genital opening very large; ductus seminalis from
bursa near junction with ductus bursae.
This genus is easily identified by its genitalia. It
shows striking affinities to Passadena of group I in shape
of harpe, with incurvation between sacculus and cucul-
lus, and in the development of an enlarged, strongly
sclerotized and projecting clasper. Passadena also has
opposed signa similarly spined.
371. Adelperga cordubensiella (Ragonot), new combination
Ficures 429, 772
Heterographis cordubensiella Ragonot, Nouv. Gen., p. 30, 1888.
Hulstia cordubensiella (Ragonot), Monograph, pt. 2, p. 128, 1901.
Forewing with color and markings of Huzophera semi-
funeralis (especially the pale color form of its synonym
aglaeella Ragonot; see p. 273); the antemedial line far
out (at or very near middle of wing), nearly vertical,
slightly notched at top of cell and at lower fold, white
187
bordered outwardly by a black line; subterminal white
line sinuate, incurved at vein 6 and the lower fold,
bordered inwardly by black; the two transverse lines
rather close (as in typical Huzophera) and the space
between them dusted with blackish scales; otherwise
the ground color of the wing is ash gray strongly shaded
with reddish ocherous, especially in the enlarged basal
area; a blackish spot on inner margin near base; a
smaller blackish spot on costa just beyond the sub-
terminal line and a row of black dots along termen.
Hind wing whitish, shading to pale smoky fuscous
towards apex and outer margin. Alar expanse, 15-18
mm,
Genitalia with characters as given for the genus.
Type Locauity: Cérdoba, Argentina (type in Paris
Mus.).
Foop prant: Unknown.
Distrisution: Argentina: Cérdoba, Las Vasquez,
Tucuman.
The figure in the Ragonot Monograph (pl. 22, fig. 20)
is misleading. It shows the hind wing much too dark
and uniformly colored, and shows none of the strong
blackish dusting in the area between the antemedial
and subterminal lines.
92. Genus Eumysia Dyar
Eumysia Dyar, Ins. Insc. Menstr., vol. 18, p. 220, 1925.
of genus: Yosemitia mysiella Dyar.)
Tongue well developed. Antenna strongly ciliate in
male (cilia about three times the width of shaft), simple
in female. Labial palpus porrect, long and beaklike
(projecting about three times the length of the head
beyond it). Maxillary palpus minute, filiform. Fore-
wing smooth; 11 veins; vein 2 from before angle of cell; 3
from the angle; 4 and 5 very shortly stalked or con-
nate; 6 from below upper angle of cell, straight; 10 from
cell, approximate to stalk of 8-9 at base; male without
costal fold. Hind wing with vein 2 close to lower outer
angle of cell; 3 and 5 stalked; 7 and 8 closely approxi-
mate or contiguous for some distance from cell; cell
slightly less than one-half the length of wing; disco-
cellular vein curved. Eighth abdominal segment of
male with a pair of short, weak hair tufts.
Male genitalia with gnathos terminating in an
elongate hook. Uncus narrowly and bluntly rounded
at apex. Harpe with lower margin evenly curved, no
incurvation between sacculus and cucullus; clasper
present, developed as a strongly sclerotized, nearly
straight hook. Anellus a simple plate. Aedeagus
short, straight, moderately slender; penis unarmed.
Vinculum broad, short, truncate.
Female genitalia with bursa and ductus bursae
simple; ductus seminalis from middle of ductus bursae.
The genus was originally erected for nine species
which Dyar removed from Zophodia and Yosemitia.
As here restricted it includes only three of these (mysi-
ella, maidella, and fuscella), a fourth species (pallidi-
pennella), which Dyar had referred from Yosemitia to
Zophodia, and a new species from Washington State.
(Type
188 UNITED STATES NATIONAL MUSEUM BULLETIN 207.
There are no structural differences in genitalia that can
be used to distinguish these supposed species. The
form of the clasper of the harpe distinguishes Humysia
from any other genus in group II. A similar develop-
ment of the clasper occurs in Nephopteryx of group I.
In general habitus Humysia most resembles Ragonotia
in the Anerastiinae.
372. Eumysia mysiella (Dyar)
Fieures 430, 920
Yosemitia mysiella Dyar, Proc. Ent. Soc. Washington, vol. 7, p.
36, 1905.—Barnes and McDunnough, Contributions, vol. 3,
p. 199, 1916.
Eumysia mysiella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221,
1925.—McDunnough, Check list No. 6295, 1939.
Forewing white dusted with blackish fuscous, making
the general color a light ashen gray; antemedial white
line sinuate, nearly vertical, bordered inwardly from
inner margin to top of cell by a broad ocherous bar,
this bar preceded by a blackish margin, especially
towards inner margin; similar blackish scaling out-
wardly bordering the antemedial white line, sometimes
faint, but usually distinct towards costa; subterminal
line obscure, more or less shaded inwardly by dark
scaling; blackish discal dots (especially the one at lower,
outer angle of cell) usually distinct. Hind wing semi-
hyaline, white. Alar expanse, 21-27 mm.
Genitalia with characters as given for the genus.
TypE Locauity: Stockton, Utah (type in USNM).
Foop prant: Unknown.
Distrisution: Utah, Stockton (Sept.), Hureka
(June); Arizona, Phoenix (Aug.), Tempe (Aug.), Red-
ington; New Mexico, Las Vegas (July), Jemez Springs
(July).
373. Eumysia maidella (Dyar)
Figure 431
Yosemitia maidella Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 36, 1905.
Eumysia maidella (Dyar), Ins. Inse. Menstr., vol. 13, p. 221,
1925.—McDunnough, Check list No. 6297, 1939.
Similar to mysiella but averaging larger, with dark
markings more intensified along the veins and bordering
antemedial and subterminal lines, ocherous subbasal
bar fainter and in some specimens absent. Alar ex-
panse, 27-32 mm.
Genitalia similar to those of mysvella.
TYPE LOCALITY: Stockton, Utah (in USNM).
Foop Piant: Unknown.
Distrisution: Unitep Starrs: Utah, Stockton
(June, July), Eureka (June), Cedar Mts. (Iron County,
July); Colorado, Fort Collins; Arizona (Sept.); New
Mexico, Fort Wingate (May); California, Loma Linda
(Aug.). Canapa: British Columbia, Kaslo (Aug.).
Probably not specifically distinct from mysvella.
There are no structural differences between them. The
character given by Dyar, costa of male forewing ‘‘con-
cave” (maidella) or ‘‘not concave’ (mysiella) is purely
imaginary. Some females as well as some males of
both ‘‘species” appear to have the costa slightly concave
at middle but this is an optical illusion due to imperfect
flattening of the wings in their spreading. The differ-
ences in size and intensity of markings are slight. How-
ever, since we know nothing of the biology of any
Eumysia and since such differences may coincide with
differences in host plants it seems advisable to keep the
names separated for the present.
374. Eumysia pallidipennella (Hulst), new combination
Volusia pallidipennella Hulst, Canadian Ent., vol. 27, p. 56, 1895.
Trivolusia pallidipennata Hulst, U.S. Nat. Mus. Bull. 52, p. 438,
1903 (misspelling).
Yosemitia pallidipennella (Hulst) Barnes and McDunnough,
Contributions, vol. 3, p. 199, 1916.
Zophodia pallidipennella (Hulst) McDunnough, Check list, No.
3606, 1939.
Paler and averaging smaller than mysiella with dark
dusting fainter, giving the wing a more ocherous gray
than ashy gray tint; ocherous subbasal bar somewhat
more strongly accented. The genitalia of the female
type show no appreciable differences from those of
mysiella or maidella. The name may represent nothing
more than a race or variety but it should be kept specifi-
cally distinct until closer relationship is proven. In a
series of some 40-odd specimens before me the color
seems fairly constant. Alar expanse, 19-24 mm.
TypE Locauity: Colorado (type in AMNH, ex Rut-
gers).
Foop prant: Unknown.
Distrisution: Colorado; Utah, Stockton (June, Aug.,
Sept.), Vineyard (June, July); New Mexico, Mesilla
Park (May); California, San Francisco (Apr.), Loma
Linda (Apr., Aug., Oct.); Washington, Pullman (July,
Aug.). .
375. Kumysia fuscatella (Hulst)
Zophodia fuscatella Hulst, Canadian Ent., vol. 32, p. 173, 1900.
Eumysia fuscatella (Hulst) Dyar, Ins. Insc. Menstr., vol. 13, p.
221, 1925.—McDunnough, Check list, No. 6298, 1939.
Forewing mouse gray; transverse white lines but
faintly indicated. Hind wings pale smoky fuscous.
Alar expanse, 21 mm.
Typr Locatity: Los Angeles County, Calif. (type in
USNM).
Foop puant: Unknown.
Known only from the male type.
376. Eumysia semicana, new species
Antenna white annulated with black. Head, palpi,
thorax, and forewing white densely dusted with blackish
fuscous, giving the moth a predominately slate-gray
color with ashy white streakings on forewing in the
interspaces between the veins, through the middle of
the cell and bordering the upper and lower veins of cell;
antemedial and subterminal lines obsolete, indicated
only by an obscure, transverse, angulate, dark shading
at basal third and a couple of dark spots on costa near
apex, from the inner of which a dark shade extends
toward cell; base of wing (before transverse shade) more
or less clouded by dark scaling; discal spots at end of
cell, small, blackish; some obscure blackish dots along
termen; cilia fuscous, with a median white band.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 189
Hind wing smoky fuscous; cilia slightly paler, with a
whitish median band. Alar expanse, 30 mm.
TypE Locatity: Yakima, Wash. (type in USNM,
61364).
Foop pirant: Unknown.
Described from female type and one female paratype
from the type locality, collected by A. Rolfs, May 20,
1932, and received from J. F. G. Clarke under his Nos.
4008 and 4009.
A large, dark species. It may eventually prove to be
only an extremely dark color form or race of maidella,
but superficially it is so different in color and so unlike
any other described species in the genus that a separate
specific designation seems warranted.
Genus 93: Divitiaca
[Venational division D. Forewing with 10 veins; 10 from cell,
8 and 9 stalked; 4 and 5 united; 2 and 3 from the cell. Hind
wing with 7 and 8 approximate; 2 from close to lower outer angle
of cell; discocellular vein curved. Labial palpi porrect. Male
genitalia with uncus triangulate, tapering evenly to rounded or
bluntly pointed extremity, large in proportion to reduced
tegumen; transtilla incomplete; harpe with a transverse sclero-
tized ridge from base of costa to lower angle of cucullus; penis
without cornuti or other appreciable sclerotizations; vinculum
broad, stout, short. Female genitalia with ductus seminalis
from ductus bursae.]
93. Genus Divitiaca Barnes and McDunnough
Divitiaca Barnes and McDunnough, Contributions, vol. 2, p. 183,
1913. (Type of genus: Divitiaca ochreila Barnes and
McDunnough.)
Tongue well developed. Antenna pubescent; in
male a shallow sinus and slight scale tuft at base of
shaft; in female simple. Labial palpus porrect; long,
extending over twice the length of the head beyond it;
broadly scaled; third segment about half the length of
second. Maxillary palpus minute, filiform. Forewing
smooth; 10 veins; vein 2 from slightly before angle of
cell; 3 from angle; 4 and 5 united; 6 from below upper
angle of cell, straight; 10 from cell, separated from the
stalk of 8-9; male without costal fold. Hind wing
with vein 2 from very close to angle of cell; 3 and 5
stalked; 7 and 8 closely approximate for some distance
from cell; cell slightly less than one-half the length of
the wing; discocellular vein curved. Eighth abdominal
segment of male with a pair of short ventrolateral hair
tufts and a lateral pair of eversible lobes with long hair
tufts
Male genitalia with apical process of gnathos a stout
hook. Harpe with broadened cucullus, its outer
margin straight and vertical or slightly rounded; a
transverse sclerotized ridge extending from base of costa
to lower angle of cucullus. Anellus a slightly curved
plate with short lateral lobes. Aedeagus short, slender,
needlelike; penis without armature. Vinculum short,
extremity narrowly rounded.
Female genitalia with bursa and ductus bursae mem-
branous; bursa large, sausage shaped or trilobed (twice
constricted near junction with ductus bursae), without
signum; ductus bursae long, very slender, with or with-
out a few fine spines at junction with bursa; ductus
seminalis from ductus bursae nearer to bursa than to
genital opening.
This genus is closely related to Macrorrhinia and
Ocala, agreeing with them in genitalic and most other
structural characters except for the absence of vein 4 of
forewing, a consistent character for all the species and
specimens of the genus.
377. Divitiaca ochrella Barnes and McDunnough
Ficures 101, 433, 922
Divitiaca ochrella Barnes and McDunnough, Contributions, vol.
2, p. 188, 1918—McDunnough, Check list, No. 6357, 1939.
Forewing pale ocherous; veins faintly outlined,
whitish; antemedial line indicated by an outwardly
curved and slanting row of blackish spots; between this
and base of wing, above inner margin, a small, more or
less diffused, blackish spot; subterminal line faintly
indicated by a broken series of blackish streaks or spots
on the veins; a dark discal spot at lower outer angle of
cell, the discal spot at upper outer angle very faint or
absent. Hind wing semihyaline, whitish. Alar ex-
panse, 12-17 mm.
Male genitalia with apical process of gnathos a
moderately long, stout, slightly curved and evenly
tapering hook. Female genitalia with ductus bursae
weakly spined at junction with bursa copulatrix; bursa
sausage-shaped, in part minutely scobinate.
Type tocauity: Everglade, Fla. (Apr., type in
USNM).
Foop prant: Unknown.
Known only from the type series and one other male
from the type locality, and one male from Marco, Fla.,
all in the National Collection.
378. Divitiaca simulella Barnes and McDunnough
Divitiaca simulella Barnes and McDunnough, Contributions,
vol. 2, p. 183, 1913—McDunnough, Check list, No. 6356,
1939.
Similar to ochrella, differing only in its smaller average
size, and somewhat darker fore and hind wings. The
hind wing is a pale smoky hyaline hue. Alar expanse,
12-14 mm.
The male genitalia of the type are a trifle smaller than
those of the type of ochrella and the apex of uncus
seems slightly more pointed; but these are scarcely
more than individual differences. The female genitalia
are essentially like those of ochrella.
Typr tocatity: Everglade, Fla. (Apr., type in
USNM).
Foop piant: Unknown.
Represented in the National Collection by the type
series and one other male from the Everglades. Very
doubtfully distinct from ochrella. Ina reared series the
color differences would probably disappear and speci-
mens reared under moist conditions would show more
190 UNITED STATES NATIONAL MUSEUM BULLETIN 207
intense and diffused markings and a considerably darker
coloration.
379. Divitiaca parvulella Barnes and McDunnough
Fieuries 432, 921
Divitiaca parvulella Barnes and McDunnough, Contributions, vol.
2, p. 183, 1913 —McDunnough, Check list, No. 6355, 1939.
Averages smaller than simulella or ochrella. Ground
color and dark markings of forewing similar to those of
simulella but with a rather broad whitish band along
costa. Hind wings pale to dark smoky fuscous. Alar
expanse, 9-12 mm.
Male genitalia with apical process of gnathos a long,
stout arm, swollen and abruptly hooked at the end.
Female genitalia with bursa and ductus bursae smooth;
bursa trilobed (twice constricted near junction with
ductus bursae).
Typr Locatity: Marco, Fla. (type m USNM).
Foop prant: Achyranthus ramosissima.
Distrisution: Florida, Marco (Apr.), Vero Beach
(Apr.), Key West (Apr.).
Easily distinguished from other species in the genus
by the contrastingly white costa of forewing and the
peculiar development of gnathos and bursa. The Key
West specimens (1 ¢' and 3 9) were reared Apr. 7, 1945,
in connection with the Special Survey of the Division of
Foreign Plant Quarantine, U. S. Department of Ari-
culture, from larvae feeding in the flowers of Achyranthus.
They have given us our only food-plant record for the
genus. The male is slightly darker and somewhat more
strongly marked than either of the two males of the type
series. The reared females and a collected female from
Vero Beach are even darker, their hind wings being a
dark smoky fuscous. The larva lacks the sclerotized
rings about seta Iib of mesothorax and seta III of
eighth abdominal segment characteristic of most phyci-
tine larvae.
380. Divitiaca parvulella consociata, new race
Similar to typical parvulella except somewhat paler
and larger. The hind wing of the female is darker than
that of the male but not so dark as in the reared females
of parvulella from Key West, a difference of little or no
significance as between collected and reared specimens.
The genitalia, male and female, are also slightly larger
than those of Florida parvulella but not structurally
different. Alar expanse, 11-13.5 mm.
TypE Locauity: Valle de Medellin, Colombia (type
in USNM, 61365).
Foop pLrant: Unknown.
Described from male type and one male and one
female paratype from the type locality, received from
F. L. Gallego M. under his No. 111, and dated “January
1942.”
I name these examples with great reluctance and do
so only as a precautionary measure, since we have no
examples of Divitiaca from any intervening area between
Colombia and the United States.
Genera 94-97: Macrorrhinia to Protasia
[Venational division A. Forewing with 11 veins; 10 from cell, 8
and 9 stalked; 4 and 5 stalked; 2 and 3 from cell. Characters
otherwise as for previous group (Dzvitiaca).]
94. Genus Macrorrhinia Ragonot
Macrorrhinia Ragonot, N. Amer. Phycitidae, p. 13, 1887.
of genus: Macrorrhinia aureofasciella Ragonot.)
Dolichorrhinia Ragonot, Nouv. Gen., p. 28, 1888; Monograph,
pt. 2, pp. xi, 190, 1901—Hulst, Phycitidae of N. Amer., p.
190, 1890.
Tongue well developed. Antenna pubescent (cilia
about as long as width of shaft); in male a shallow sinus
with a very small tuft at base of shaft; in female simple.
Labial palpus porrect, downcurved; long, extending at
least three times length of head beyond it. Maxillary
palpus minute, filiform. Forewing smooth; 11 veins;
vein 2 from slightly before angle of cell; 3 from angle;
4 and 5 short stalked; 6 from below upper angle of cell,
straight; 10 from cell, at base approximate to stem of
8-9; male without costal fold. Hind wing with vein 2
from close to angle of cell; 3 and 5 stalked; 7 and 8
approximate or weakly anastomosed beyond cell; cell
slightly less than one-half the length of wing; discocel-
lular vein curved. Eighth abdominal segment of male
with a pair of short ventrolateral hair tufts and a lateral
pair of eversible lobes with long hair tufts (as in Diiti-
aca).
Male genitalia with apical process of gnathos a stout
hook. Harpe with broadened cucullus; a transverse
sclerotized ridge extending from base of costa to lower
angle of cucullus. Anellus a slightly curved plate with
short, thick, lateral lobes. Aedeagus short, slender,
needlelike; penis without armature. Vinculum short
(as broad as long); extremity rounded.
Female genitalia with bursa and ductus bursae mem-
branous throughout, except for a slight sclerotization of
the genital opening; bursa large, sausage shaped, with-
out signum; ductus bursae long, very slender for two-
thirds of its length, suddenly and considerably expanded
before junction with bursa; ductus seminalis from ductus
bursae near genital opening.
Ragonot (1888) proposed Dolichorrhinia as a new
name for Macrorrhinia Ragonot (1887) since the latter
was, in his opinion, a homonym, “resembling too much
the name Macrorhinus already employed twice.” The
name is uncomfortably close to and much nearer a
homonym of Macrorrhina Berthold (1827); but under
our interpretation of the International Code the name
Macrorrhinia is homonymous with neither and will have
to replace Dolichorrhinia which lepidopterists since
Ragonot have been using for the genus.
(Type
381. Macrorrhinia aureofasciella Ragonot
Fiaures 61, 437, 924
Macrorrhinia aureofasciella Ragonot, N. Amer. Phycitidae, p. 13,
1887.
Dolichorrhinia aureofasciella (Ragonot) Hulst, Phycitidae of N.
Amer., p. 190, 1890; U.S. Nat. Mus. Bull. 52, p. 433, 1902.—
Hampson, 7m Ragonot, Monograph, pt. 2, p. 190, 1901.—
McDunnough, Check list, No. 6351, 1939.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 191
Forewing ocherous gray; antemedial line broad,
slightly oblique, orange, preceded by a blackish line
which extends from middle of cell to inner margin;
discal dots at end of cell, blackish, the lower one always
distinct; subterminal line very faint, whitish, indis-
tinctly shaded inwardly by dark scaling, indented at
vein 6 and lower fold; terminal dots faint. Hind wing
translucent, whitish with a pale smoky tint. Alar
expanse, 13-22 mm.
Male genitalia with apical hooked process of gnathos
but slightly curved; aedeagus with a row of 4 or 5 very
small spines near apex. Female genitalia as given for
the genus.
Typm Locauity: Arizona (type in Paris Mus.).
Foop prant: Unknown.
Distrisution: Unirep Srates: Arizona, Baboqui-
vari Mts. (Apr. to Dec.), Greaterville, Palmerlee (Apr.),
Santa Rita Mts. (May, June, July), Wilgus Mts.;
New Mexico, Albuquerque (July), Las Vegas; Tezas,
Alice (June), Blanco County (July, Oct.), Brownsville,
Burnet County, Kerrville (Apr., July), Shovel Moun-
tain. México: Sonora.
Superficially aureofasciella is easily confused with
Eumysia pallidipennella, especially the female. Their
genitalia, however, readily separate the two species.
382. Macrorrhinia placidella (Zeller)
Myelois placidella Zeller, Isis von Oken, p. 874, 1848.
Dolichorrinia placidella (Zeller) Ragonot, Monograph, pt. 2,
p. 190, 1901.
This species is represented only by the female type in
Berlin. According to Ragonot the specimen is in poor
condition. I have seen nothing from tropical America
that matches Zeller’s description or Ragonot’s figure
(Monograph, pl. 31, fig. 20). Very likely the species
does not belong in Macrorrhinia, but without evidence
to the contrary must be left where Ragonot placed it.
Type Locauity: Alegrete, Brazil (type in Mus. Univ.
Berlin).
Foop pirant: Unknown.
95. Genus Ocala Hulst
Ocala Hulst, Canadian Ent., vol. 24, p. 61, 1892.
genus: Ocala dryadelia Hulst.)
(Type of
Characters of Macrorrhinia except: Labial palpus of
male grooved to hold maxillary palpus; maxillary pal-
pus of male in the form of an aigrette. There is no
structural difference between the two genera in the
tufting of the male abdomen. Hulst made no dissec-
tions and consequently failed to see the eversible tufts
in Macrorrhinia.
The palpal differences may be of no more than specific
significance here as they are, for example, in Dioryctria;
but until more material is available, other possible
species discovered, and something known about their
life histories it seems advisible to keep the generic
names separate.
383. Ocala dryadella Hulst
Ficures 59, 438, 923
Ocala dryadella Hulst, Canadian Ent., vol. 24, p. 61, 1892.—
MeDunnough, Check list, No. 6359, 1939.
Dolichorrhinia platanella Grossbeck, Bull. Amer. Mus. Nat.
Hist., vol. 37, art. 1, p. 131, 1917.—Barnes and McDun-
nough, Contributions, vol. 3, p. 222, 1917.
Forewing pale fuscous gray; antemedial white line
obscure, indicated chiefly by a narrow outer bordering
of blackish scales, angled slightly at middle and slanting,
bordered inwardly on lower half by a pale orange spot
behind which, on inner margin, is a rather conspicuous
blackish dot; discal spots at end of cell distinct, black-
ish; subterminal line faint, with a blackish inner border,
inwardly angled at vein 6 and (slightly) at lower fold; a
row of blackish dots along termen. Hind wings trans-
lucent, pale smoky white, with some dark streaking on
the veins, especially on the females. Alar expanse,
14-17 mm.
Male genitalia with outer margin of cucullus straight;
aedeagus smooth. Female genitalia with ductus bursae
sclerotized for a short distance from genital opening and
swollen for more than a third of its length from junction
with bursa copulatrix.
Tyrer Locauities: Charlotte Harbor, Fla. (dryadella,
in AMNH, ex Rutgers); Fort Myers (platanella, in
AMNBE).
Foop prant: Unknown.
Distrrisution: Florida, Charlotte Harbor,
glade (Apr.), Fort Myers (Apr.), Marco (Apr.).
Ever-
96. Genus Valdivia Ragonot
Valdivia Ragonot, Nouv. Gen., p. 27, 1888; Monograph, pt. 2,
p. xi, 1901.—Hampson, in Ragonot, Monograph, pt. 2, p.
191, 1901. (Type of genus: Valdivia coquimbella Ragonot.)
Maricopa Hulst, Phycitidae of N. Amer., p. 205, 1890. (Type of
genus: Ciris lativitella Ragonot.)
Tongue well developed. Antenna ciliate in male
(cilia over twice the width of the shaft); simple in
female. Labial palpus porrect (downcurved), long
(projecting about 3 times the length of the head beyond
it). Maxillary palpus minute, filiform. Forewing
smooth; 11 veins; vein 2 from before outer angle of
cell; 3 from the angle; 4 and 5 stalked; 6 from below
upper angle of cell, straight; 10 from cell, closely ap-
proximate at base or connate with the stem of 8-9; male
without costal fold. Hind wing with vein 2 from very
close to outer angle of cell; 3 and 5 stalked; 7 and 8
approximate or weakly anastomosed just beyond cell;
cell slightly less than one-half the length of the wing.
Eighth abdominal segment of male with a short pair of
ventrolateral hair tufts.
Male genitalia with apical process of gnathos a rather
short, stout hook. Harpe with broadened cucullus, its
outer margin rounded; a transverse sclerotized ridge
extending from base of costa to lower angle of cucullus.
Anellus an elongate, curved plate with lateral lobes.
Aedeagus short, slender; penis without armature.
Vinculum short, terminal margin rather broadly
rounded.
192 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Female genitalia with bursa simple, membranous;
ductus bursae very slender for half its length from
genital opening, thence to bursa swollen, spiraled and
armed throughout the spiraled portion with a line of
fine, sharp spines, sclerotized only at genital opening;
ductus seminalis from ductus bursae near genital
opening.
Valdivia is close to Macrorrhinia and Ocala, dis-
tinguished by a slightly wider spacing of veins 2, 3 and
the stem of 4-5 in forewing, the longer cilia of the
male antenna, the slightly stouter aedeagus, the absence
of eversible tufts on the male abdomen, the smaller,
oval bursa and the spiraled and spined ductus bursae.
Three species are recognized as belonging to the genus.
A fourth now listed in Valdivia (Maricopa albocostella
Hulst) will have to be referred to the Anerastiinae. Its
type (co) in the Rutgers Collection has a greatly reduced
tongue, completely enclosed by the labial palpi, vein 4
of forewing absent and a slight excavation in the base
of the antennal shaft.
384. Valdivia coquimbella Ragonot
FIgurE 436
Valdivia coquimbella Ragonot, Nouv. Gen., p. 27, 1888.—Hamp-
son, 7n Ragonot, Monograph, pt. 2, p. 191, 1901.
Forewing fuscous gray; transverse pale lines obscure,
the antemedial oblique line preceded by a diffused dark
shade, the subterminal line indicated only by a broken,
weakly indented line of blackish scaling running parallel
to the outer margin; discal dots obscure, blackish; a
row of faint black dots along termen. Hind wing very
pale smoky fuscous. Alar expanse, 22 mm.
Male genitalia figured from paratype in the British
Museum from the type locality. The type in the Paris
Museum is also a male (not a female as stated by Hamp-
son) but is without an abdomen. ‘These are the only
two examples of the species known. The anellus is
much shorter than that in lativiltella. V. coquimbella
also lacks any trace of the yellowish spot edging the
antemedial line, characteristic of the other two species.
TypE LocaLity: Coquimba, Chile (type in Paris
Mus.).
Foop prant: Unknown.
385. Valdivia lativittella (Ragonot)
Figures 434, 485, 926, 927
Ciris lativitiella Ragonot, N. Amer. Phycitidae, p. 18, 1887.
Ragonotia lativitella (Ragonot), Ent. Amer., p. 117, 1889.
Maricopa lativitella (Ragonot) Hulst, Phycitidae of N. Amer.,
p- 206, 1890.
Valdivia lativittella (Ragonot) Hampson, zn Ragonot, Mono-
graph, pt. 2, p. 191, 1901.—McDunnough, Check list, No.
6352, 1939.
Zophodia aureomaculella Dyar, Journ. New York Ent. Soc., vol.
12, p. 107, 1808. (New synonymy.)
Eumysia aureomaculella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221,
1925.—McDunnough, Check list, No. 6296, 1939.
Paler than coquimbella, the forewing very pale ashy
gray; on inner margin at base a pale ocherous spot; a
similar somewhat larger spot extending halfway across
the wing and bordering the antemedial line; on inner
margin between the two ocherous patches a blackish
fuscous spot (rather pronounced on the male, less so on
the female); antemedial line vertical, faint, mdicated
chiefly by some irregular and broken, outer, blackish
fuscous shading; subterminal whitish line more distinct,
indented at vein 6 and lower fold and margined inwardly
by a narrow dark line; discal and terminal dots blackish
fuscous, the latter faint. Hind wing very pale smoky
fuscous with a faint ocherous tint; a narrow dark line
along termen; veins faintly darkened. Alar expanse,
16-21 mm.
Male genitalia differ from those of coqguimbella chiefly
in their much longer anellus.
Type Locatitigs: Arizona (lativittella, in Paris Mus.) ;
Bremond, Tex. (aureomaculella, in USNM).
Foop purant: Unknown.
Distrisution: Unirep Statss: Arizona, Baboqui-
vari Mts. (July), Greaterville (June), Phoenix (Aug.),
Pinal Mts. (Aug.); Texas, Bremond (Apr.), Brownsville
(Mar.), San Benito (Mar., May). Mexico: Durango.
Twenty-five specimens are before me. A male from
Durango, Dyar’s type of aureomaculella from east-
central Texas and a series from Arizona exhibit no es-
sential differences. The Texan specimens from Browns-
ville and San Benito (1 @ and 3 6) show some trifling
differences in male genitalia and considerably weaker
spining of the ductus bursae in the female (fig. 926),
but nothing, however, that would justify specific or
even racial separation. Dyar’s supposed species is an
obvious synonym. He had not recognized either
Valdivia or lativittella when he described aureomaculella
or, later, transferred it to Humysia. The key vena-
tional character given by Hampson for the separation
of latiittella from coquimbella (vein 8 of hind wing “free”
from or anastomosed with 7) does not hold. In our
Pinal Mountains series of lativittella ves 7 and 8 occur
both ways, closely approximate or partially anasto-
mosed.
386. Valdivia walkerella (Ragonot), new combination
Fieure 439
Saluria walkerella Ragonot, Nouv. Gen., p. 44, 1888.
Hypogryphia walkerella (Ragonot) Hampson, in Ragonot, Mono-
graph, pt. 2, p. 194, 1901.
I have seen nothing from Chile that exactly matches
Ragonot’s description or the figure in his Monograph
(pl. 37, fig. 21), but the genitalia of the male type clearly
shows it to be a Valdivia and not a member of the Old
World genus Hypogryphia where Hampson placed it.
Veins 4 and 5 of forewing are shortly stalked and the
ocherous spot on inner margin before the antemedial
line is present, as in Jativitiella which it strongly re-
sembles in genitalia and general habitus. Alar expanse,
18 mm.
Typr Locanity: Valparaiso, Chile (type in Paris
Mus.).
Foop prant: Unknown.
Known only from the type.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 193
97. Protasia, new genus
Typ or Genus: Valdivia mirabilicornella Dyar.
Tongue short (but somewhat exposed between the
palpi). Antenna unipectinate in male, simple in fe-
male. Labial palpus porrect, long (projecting more than
3 times the length of the head beyond it). Maxillary
palpus minute, filiform. Forewing smooth; 11 veins;
vein 2 from before outer angle of cell; 3 from the angle;
4 and 5 very shortly stalked; 6 from below upper angle
of cell, straight; 10 from cell, approximate to 8-9 at
base; male without costal fold. Hind wing with vein
2 from very close to outer angle of cell; 3 and 5 stalked;
7 and 8 approximate or weakly anastomosed for a short
distance beyond cell; cell less than one-half the length
of the wing. Eighth abdominal segment of male
simple.
Male genitalia with apical process of gnathos a stout,
bluntly pointed hook. Harpe with lower margin
evenly curved; cucullus elongate, tapering slightly to
somewhat narrowly rounded apex; a transverse, weakly
sclerotized ridge extending from slightly below base of
costa to lower angle of cucullus. Anellus a simple,
nearly flat plate, without lateral lobes. Aedeagus
short, stout; penis without armature. Vinculum short,
terminal margin broadly rounded.
Female genitalia with bursa simple, membranous,
elongate; ductus bursae short, simple, membranous
throughout; ductus seminalis from ductus bursae.
The genus is readily distinguished from the others in
this immediate group by its unipectinate male antenna,
short tongue, lack of hair tufts on eighth abdominal
segment of male, and short and simple ductus bursae.
387. Protasia mirabilicornella (Dyar), new combination
Ficures 440, 925
Valdivia mirabilicornella Dyar, Proc. Ent. Soc. Washington, vol.
10, p. 60, 1908.—McDunnough, Check list, No. 6353, 1939.
Whitish gray with a sprinkling of dark scales and a
faint ocherous shading in the median and lower folds of
forewing; antemedial line dentate, outwardly bordered
by a thin blackish fuscous line, the latter sharply out-
angled between costa and middle of cell, very slightly
out-angled at lower margin of cell and at inner margin,
preceded by a more or less obscure, broad ocherous
orange shade, bordered basally on inner margin of wing
by a blackish fuscous patch, the latter encroaching upon
and more or less obscuring the ocherous shade; subter-
minal line with a narrow blackish inner border, sharply
indented to vein 6, thence bulging outward and again
indented at lower fold; discal dots distinct, blackish;
terminal dark dots more or less confluent. Hind wing
pale brassy whitish (pale smoky fuscous in some speci-
mens), sheeny; a slightly darker line along termen;
cilia somewhat paler. Alar expanse, 18-26 mm.
Male genitalia with uncus evenly tapering from broad
base to rather narrowly rounded apex; elements of
transtilla small, weak, recurved (fig. 440b). Female
genitalia with bursa narrow, nearly three times as long
as ductus bursae.
300329—56——14
Type Locatity: San Diego, Calif. (type in USNM).
Foop pirant: Unknown.
Represented by 25 specimens in the National Collec-
tion, all from the type locality (July, Oct., Nov., Dec.,
Feb.).
In his description Dyar describes the male antenna
as “bipectinate.” It is distinctly unipectinate, with
the pectinations and the shaft covered with long cilia.
Genera 98-100: Heterographis to Hulstia
[Venational division A. Forewing with 11 veins; 10 from cell;
2 from near lower outer angle of cell. Hind wing with 7 and 8
approximate or very weakly anastomosed; 2 from close to lower
outer angle of cell; discocellular vein curved. Labial palpi up-
turned. Male genitalia with uncus triangulate (or spoon-shaped),
large in proportion to reduced tegumen; tegumen constricted
before uncus, its lateral margins concave; transtilla incomplete
(except in Heterographis samaritanella); aedeagus straight,
slender; penis without cornuti or other armature. Female
genitalia with signa consisting of many straight disconnected
thornlike spines; ductus seminalis from ductus bursae.]
98. Genus Heterographis Ragonot
Heterographis Ragonot, Ent. Monthly Mag., vol. 22, p. 31, 1885;
Monograph, pt. 2, p. x, 1901.—Hulst, Phycitidae of N.
Amer., p. 186, 1890.—Hampson, in Ragonot, Monograph,
pt. 2, p. 140, 1901.—Bisset, in Pierce and Metcalf, Genitalia
of the British Pyrales, p. 59, 1938 (fixes type of genus).—
Janse, Journ. Ent. Soc. South Africa, vol. 8, p. 37, 1945.
(Type of genus: Huzophera samaritanella Zeller; figs. 441,
928).
Mona Peat Ent. Amer., vol. 4, p. 115, 1888.
Mona olbiella Hulst.)
Tongue well developed. Antenna simple, pubescent;
in the male the shaft slightly curved towards base.
Labial palpus upcurved, reaching above vertex. Max-
illary palpus minute, filiform. Forewing smooth; 11
veins; vein 2 from near lower outer angle of cell; 3 from
the angle; 4 and 5 connate or very shortly stalked,
closely approximate to 3; 6 from below upper angle of
cell, straight; 10 from cell, separated from the stalk of
8-9; male without costal fold. Hind wing with 2 from
close to outer angle of cell; 3 and 5 stalked; 7 and 8
closely approximate (occasionally contiguous or weakly
anastomosed for a short distance) beyond cell; cell less
than one-half (over one-third) the length of the wing.
Eighth abdominal segment of male with a strong pair
of ventrolateral hair tufts.
Male genitalia with apical process of gnathos a stout
hook. Elements of transtilla long, slender, curved,
their apices touching and weakly fused (samaritanella)
or narrowly separated (morrisonella) ; deeply and round-
ly arched behind aedeagus. Harpe simple, narrowly
elongate; apex rounded. Anellus a slightly curved
plate with lateral arms. Aedeagus long, slender; penis
without armature. Vinculum at least as long as broad;
truncate; terminal margin straight or slightly notched.
Female genitalia with bursa copulatrix and ductus
bursae membranous except for a slight sclerotization of
the tube of ductus near genital opening; bursa elongate
with signa consisting of numerous straight, sharp, broadly
(Type of genus,
194
based, disconnected spines, a few of these spines in
ductus near its junction with bursa; ductus seminalis
from ductus bursae near genital opening.
Represented in North America by a single species.
388. Heterographis morrisonella Ragonot
Figures 442, 929
Heterographis morrisonella Ragonot, N. Amer. Phycitidae, p. 11,
1887.—Hulst, Phycitidae of N. Amer., p. 186, 1890.—Hamp-
son, zz Ragonot, Monograph, pt. 2, p. 169, 1901.—Dyar,
Proe. Ent. Soc. Washington, vol. 7, p. 38, 1905—McDun-
nough, Check list, No. 6339, 1939.
Heterographis colorandensis Ragonot, N. Amer. Phycitidae, p. 12,
1887
Mona olbiella Hulst, Ent. Amer., vol. 4, p. 116, 1888.
Heterographis morrisonella coloradensis (Ragonot) Hulst, Phy-
citidae of N. Amer., p. 186, 1890.—Hampson, in Ragonot,
Monograph, pt. 2, p. 169, 1901.
Heterographis morrisonella olbiella (Hulst), Phycitidae of N.
Amer., p. 186, 1890.—Hampson, 7m Ragonot, Monograph,
pt. 2, p. 169, 1901.
Heterographis ignistrigella Ragonot, Monograph, pt. 2, p. 166,
1901.—McDunnough, Check list, No. 6338, 1939. (New
synonymy.)
Honora palloricostella Walter, Proc. Ent. Soc. Washington, vol.
30, p. 141, 1928.—McDunnough, Check list, No. 6344, 1939.
(New synonymy.)
Extremely variable in color and intensity of mark-
ings, ranging from a creamy ocherous form with only
the faintest dark dusting to forms much suffused with
reddish or blackish fuscous. Forewing with costa some-
what paler (from base to outer line) than the remainder
of the wing in nearly all specimens, contrastingly whit-
ish in some; transverse pale lines more or less distinct;
antemedial line outwardly slanting to lower margin of
cell, indented below at fold; subterminal line nearly
parallel with outer margin, indented at vein 6 and lower
fold; in some specimens an ocherous band, a blackish
fuscous patch, or a diffused reddish shade preceding the
antemedial line; similar colorations frequently following
the subterminal line; the area between the transverse
lines frequently grayish from the heavy dusting of dark
scales; usually the blackish dusting is more conspicuous
along some of the veins. Hind wings from very pale to
dark smoky fuscous; cilia whitish; veins more or less
outlined by fuscous shading. Alar expanse, 13-23 mm.
Male genitalia with lateral margins of uncus straight,
apex bluntly rounded. Harpe with costa and lower
margin of harpe nearly parallel; cucullus slightly curved
near apex. Female genitalia with bursa long; spines of
signa arranged in more or less regular, longitudinal
series, the number and arrangement of spines differing
in different specimens, hardly any two examples being
identical (extremes of variation are shown in the fig-
ures) ; ductus bursae long, slender throughout its length,
weakly sclerotized between junction of ductus seminalis
and genital opening.
TypE Locatitius: Texas (morrisonella and ignistri-
gella, in Paris Mus.); Colorado (coloradensis, in Paris
Mus.); Salida, Colo. (olbiella, in AMNH, ex Rutgers)
Tempe, Ariz. (palloricostella, in USNM).
Foop pruant: Franseria bipinnatifida. A single fe-
male reared by F. N. Pierce, Sept. 6, 1938, at El Se-
UNITED STATES NATIONAL MUSEUM BULLETIN 207
gundo, Calif., our only food-plant record. The species
presumably feeds on other composites.
Distrisution: Unirep States: Texas, Beeville
(Oct.), Brownsville (June), Burnet County (Apr.),
Sabinal (Apr., May, June), San Benito (June, July,
Sept.), San Diego (May), Zavalla County (Apr.); New
Mexico, Albuquerque (July), Deming (July), Fort Win-
gate (June, July), Hot Springs (Aug.), Jemez Springs,
Las Vegas (May), Mesilla Park (Apr., May), Santa Fe;
Arizona, Baboquivari Mts. (July), Phoenix (Mar.,
Apr., Aug., Oct.), Tempe (Apr., Aug., Sept., Oct.),
White Mts. (Aug.), Yuma County (Mar.); Colorado,
Denver, Glenwood Springs (Aug.); Utah, Provo (July),
Scover Valley (May), Stockton (June, July, Aug., Oct.),
Vineyard (June); Nevada, Pyramid Lake; California,
Balboa, (Sept.), Death Valley (Apr.), El Segundo
(Sept.), Ibanpa Mts. (Oct.), Jacumba (June), Laguna
(Sept.), La Puerta (July), Loma Linda (Mar., Apr.,
May, Aug., Sept.), Narrows (Mar.), Palm Springs
(Mar., Apr., June), San Diego (June); Washington,
Blue Mts. (Bone Springs, July), Pullman (June, July),
Walla Walla (May, June), Yakima (June). M*xrco:
Durango, Sonora.
The color variations are so many, the color forms so
widely distributed, and the intergradations so gradual
that none of the synonymical names can be used as a
racial designation. The species, however, is clearly
defined and easily identified by its genitalic characters.
99. Genus Staudingeria Ragonot
Staudingeria Ragonot, Ann. Soc. Ent. France, ser. 6, vol. 7, p.
249, 1887; Monograph, pt. 2, p. x, 1901.—Hulst, Phycitidae
of N. Amer., p. 185, 1890—Hampson, in Ragonot, Mono-
graph, pt. 2, p. 181, 1901. (Type of genus: Ancylosis morbo-
sella Staudinger.)
Characters of Heterographis except: Labial palpus
obliquely ascending, third segment very short, less than
one-third the length of second, in male second segment
grooved to hold the maxillary palpus; maxillary palpus
of male in the form of an aigrette; female genitalia with
bursa very narrowly elongate, ductus bursae much
shorter than bursa.
The status of this genus in relation to Heterographis
is similar to that of Ocala in relation to Macrorrhinia.
If only our North American species were involved I
should be inclined to consider the differences in male
maxillary palpi and the female bursae to be merely spe-
cific; but apparently some of the species under each
generic name in the Old World are properly assigned
and the two groups of species there are separable on the
male palpal character; so, for the present at least, it
seems advisable to keep the names separate. I have
not seen the type of Staudingeria, but there is no reason
to doubt that our American species is congeneric. Many
specimens of albipenella (especially what Dyar described
as olivacella) are an exact match for Ragonot’s figure of
morbosella (Monograph, pl. 27, fig. 1). The most striking
difference between our American representatives of
Staudingeria and Heterographis is in the comparative
length of the ductus bursae of the female. In Hefero-
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 195
graphis it is at least as long as the bursa. In Staudin-
geria it is much shorter than the bursa.
389. Staudingeria albipenella (Hulst)
Fiagures 443, 930
Pempelia albipenella Hulst, Ent. Amer., vol. 3, p. 133, 1887.
Staudingeria albipennella (Hulst) Ragonot, Ent. Amer., vol. 5, p.
116, 1889; Monograph, pt. 2, p. 186, 1901.—Dyar, Proc. Ent.
Soc. Washington, vol. 7, p. 38, 1905.—Hulst, Phycitidae of
N. Amer., p. 186, 1890.
Staudingeria olivacella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 111, 1904.
Siaudingeria perluteella Dyar, Proc.
vol. 6, p. 111, 1904.
Staudingeria albipenella (Hulst), McDunnough, Check list, No.
6337, 1939.
Variable in color and intensity of markings, ranging
from a form with forewing a uniform luteous ocherous
without definite markings, though forms more or less
dusted and clouded with blackish fuscous, to a form
with a clear red ground color and gradations of this
with the red more or less overlaid with blackish fuscous
scaling; costal edge white or whitish from base to just
before apex, the whitish costal margin sometimes very
thin and obscure but more often contrasted against the
ground color; transverse lines obsolete or obscure, in
better marked specimens chiefly indicated by their dark
marginal shadings; such shadings on subterminal line
deeply indented at vein 6 and slightly indented at lower
fold; cilia pale luteous ocherous, ocherous fuscous, or red,
concolorous with the ground color. Hind wing from
very pale to dark smoky fuscous. Fore and hind wings
have a slick, glossy sheen. Alar expanse, 18-27 mm.
Male genitalia with uncus tapering to a blunt point,
its lateral margins curved; anellus with very short
lateral lobes; aedeagus somewhat stouter than that of
Heterographis; vinculum very slightly tapered, terminal
margin broad and with a shallow indentation. Female
genitalia with bursa copulatrix a long tube terminating
in a small round bulb; signum consisting of an irregular
cluster of separate spines in the bulbous part of bursa,
a scattering of similar spines in the anterior half of the
tube and a small cluster of much shorter spines in the
tube near junction of ductus bursae; ductus not appreci-
ably sclerotized near genital opening.
Type Locauities: Southern California (albipenella,
in AMNH, ex Rutgers); Pullman, Wash. (olivacella, in
USNM); Fort Collins, Colo. (perluteella, in USNM).
Foon piant: “Loco weed.”
Distrisution: New Mexico, Fort Wingate (May);
Arizona, Phoenix (Apr.), Woodruff (June); California,
San Diego (June); Colorado, Denver (July, Sept.), Fort
Collins, Glenwood Springs (June, July); Utah, Provo,
Richfield (June), Stockton (July, Aug., Sept.); Wyo-
ming, Yellowstone Park (July) ; Idaho, Blackfoot (June),
Sawtooth (June); Oregon, Bone Springs (Blue Mts.,
July), Crater Lake (July); Washington, Godman
Spring (Blue Mts., July), Pullman (Aug., Sept.),
Vantage Ferry (May), Walla Walla (May, June, July),
Wenatchee (May, June, July, Aug.), Yakima (May).
Dyar’s two names (olivacella and perluteella) appear
Ent. Soc. Washington,
as racial designations in our lists but the differentiation
is not justified, for there is complete intergradation
between them and albipenella in any considerable
series from a given locality. The most outstanding
color form is exemplified in a series from Crater Lake,
Oreg. This is a clear, shining red variety, matched
only in some specimens from Idaho and Washington.
However there are intergrades in these localities and
one in the Crater Lake series between the red form and
the type of albipenella, so I do not feel justified in
adding another name. As in Heterographis we appear
to have only one clear-cut American species of the Old
World genus Staudingeria, and it seems limited in its
distribution to the Western States. It is easily identi-
fied by its genitalia.
100. Genus Hulstia Ragonot
Hulstia Ragonot, Monograph, pt. 2, p. x, 1901.—Hampson, in
Ragonot, Monograph, pt. 2, p. 127, 1901.
Hulstea Hulst, U. S. Nat. Mus. Bull. 52, p. 482, 1903. (Spelling.
Type of genus: Nephopteryx undulatella Clemens.)
Tongue well developed. Antenna simple, finely pu-
bescent. Labial palpus upturned; rather broadly and
smoothly scaled; third segment short, pointed. Mazxil-
lary palpus small, squamous, appressed to face. Fore-
wing smooth; 11 veins; vein 2 from near lower outer
angle of cell; 3 from the angle; 4 and 5 stalked, the
stalk closely approximate to 3 at base; 6 from below
upper angle of cell, straight; 10 from the cell, at base
connate or very closely approximate to 8-9; male with-
out costal fold. Hind wing with vein 2 from close to
outer angle of cell; 3 and 5 stalked; 7 and 8 contiguous
or partially anastomosed for a short distance beyond
cell; cell slightly less than one-half the length of wing;
discocellular vein curved. Eighth abdominal segment
of male with a pair of short, ventrolateral hair tufts.
Male genitalia with apical process of gnathos a stout,
blunt hook. Elements of incomplete transtilla, small
and weak. Harpe simple, narrowly elongate; apex
rounded. Anellus a narrowly elongate plate with
strongly sclerotized margin and short lateral lobes (re-
sembling an elongate and closed U). Aedeagus long,
moderately slender. Vinculum as broad as long, squar-
ish, terminal margin nearly straight.
Female genitalia with bursa copulatrix long, tapering
and expanding gradually from ductus bursae; demarca-
tion of bursa from ductus obscure; signa consisting of
a dense mass of straight, short, rather slender spines
with small bases and filling about half the bursa.
Ductus bursae short, with a strongly sclerotized, acutely
bent elbow before genital opening; genital opening
simple. Ductus seminalis from ductus bursae just
before sclerotized elbow.
Hulstia is very close to Heterographis and Staudin-
geria, agreeing with them in most structural characters
and differing chiefly in the more pronounced stalking
of veins 4 and 5 of forewing, the closer approximation
of vein 10 to the stalk of 8-9 at base, its much reduced
transtilla, the shape of the female bursa, the densely
196 UNITED STATES NATIONAL MUSEUM BULLETIN 207
clustered and narrowly based cornuti, and the sclero-
tized elbow in ductus bursae. It contains but one
known species. The other American species that have
been assigned to it by Hampson (texanella and cordu-
bensiella) are dissimilar in both structure and habitus.
390. Hulstia undulatella (Clemens)
Ficures 64, 444, 931
Nephopteryx undulatella Clemens, Proc. Acad. Nat. Sci. Phila-
delphia, p. 205, 1860.
Scoparia rubiginalis Walker, List, vol. 34, p. 1498, 1865.
Honora obsipella Hulst, Ent. Amer., vol. 4, p. 118, 1888 (new
synonymy).
Heterographis oblitella Ragonot (not Zeller), Ent. Amer., vol. 5,
p. 116, 1889.
Honora oblitella undulatella (Clemens) Hulst, Phycitidae of N.
Amer., p. 187, 1890.
Honora fumosella Hulst, Canadian Ent., vol. 32, p. 174, 1900.—
Forbes, Cornell Mem. 68, p. 6338, 1920.—McDunnough,
Check list, No. 6350, 1939. (New synonymy.)
Hulstia undulatella (Clemens) Hampson, in Ragonot, Mono-
graph, pt. 2, p. 128, 1901.—Hssig, Insects of western North
America, p. 711, 1926.—McDunnough, Check list, No. 6341,
1939.—Darlington, Trans. Amer. Ent. Soc., vol. 73, p. 91,
1947.
Hulstea undulatella (Clemens) Hulst, U. S. Nat. Mus. Bull. 52,
p- 432, 1903.
Forewing whitish ocherous, pale gray or dark grayish
fuscous, more or less dusted with white scaling especially
in median area of wing, in darkest specimens (color form
jfumosella) pattern mostly obliterated by dark scaling;
transverse lines normally distinct, narrow, white; ante-
medial line oblique, notched at cell and lower fold,
broadly bordered inwardly by fuscous or fuscous and
ocherous, especially toward inner margin, and outward-
ly by a narrow dark shade especially from costa to cell;
subterminal line sinuate, parallel to and rather near
termen, bordered inwardly by a narrow dark line and
outwardly by a fuscous or ocherous fuscous shade; discal
spots, especially the lower one, usually distinct. Hind
wings dark smoky fuscous. Palpi, underside of body,
and legs whitish, more or less shaded with fuscous in
darker specimens. Alar expanse, 16-22 mm.
Genitalic characters as given for the genus.
TYPE LOCALITIES: Pennsylvania (undulatella, lost);
North America (rubiginalis, in BM); ‘Texas’ [sic]
(obstpella, in AMNH, ex Rutgers); Newark, N. J.
(fumosella, in AMNH, ex Rutgers).
Foop prants: Elm [?], sugar beets.
Distrizution: Unitep States: Maine; Massachu-
setts, Martha’s Vineyard (July), Vineyard Haven
(Aug.); Connecticut, East River (July, Sept.); New
Hampshire, Hampton (June, Aug., Sept., Oct.); New
Jersey, Anglesea (June), Newark; North Carolina,
Southern Pines (Oct.); Florida, Glenwood, Hastings
(Apr.); Wisconsin, Cranmoor; Montana, Bozeman
(Aug.); Colorado, Clear Creek, Denver (Aug., Sept.),
Florisant (July), Glenwood Springs (Aug.), Windsor
(July) ; Utah, Delta (July), Logan (Aug.), Ogden (July),
Provo (July), Salt Lake, “Southern Utah” (July),
Stockton (Aug.), Vineyard (June); Nevada, Clark
County, Reno (July); New Mexico, Fort Wingate
(June), Hot Springs, Jemez Springs (July), Sapello
Canyon (July) ; Arizona, Flagstaff, ‘‘Southern Arizona,”
Williams (July); California, Alameda (Aug., Oct.),
Chino (Nov.), Folsom (July), Palo Alto, Sacramento
(Oct.), San Diego (Sept., Oct., Nov.), Siskiyou (Sept.),
Truckee (Aug.), Woodland; Washington, Bellingham
(July, Aug.), Friday Harbor (July), Pullman (May,
July, Aug., Sept.), Snake River (May), Walla Walla
(June, July), Wenatchee (July, Aug.). CAaNapa: Que-
bec, Montreal; Alberta, Calgary (July, Aug.); Manitoba,
Aweme (July, Aug.); British Columbia, Arrowhead
Lake, Goldstream (July), Kaslo, Victoria (Aug.).
This species was described from eastern specimens,
but seems to be much more abundant in our Western
States. Very little is known of its life history. Clem-
ens states that he took larvae and pupae of undulatella
on elm at Niagara Falls, Canada. He did not rear any
moths from the larvae and his statement is ambiguous
as to the rearing of moths from the pupae (which were
found under the bark of the tree), so the association of
undulatella with elm as a host is by no means estab-
lished. In the National Museum we have a half-dozen
reared specimens from Colorado, Utah, and California
reared from larvae feeding on sugar-beet foliage. ‘This
is the only authenticated food-plant record with which
IT am acquainted.
The name fumosella represents nothing more than a
much suffused, dark color form. There are four
examples in the National Collection matching Hulst’s
type and several specimens from widely scattered
localities intergrading between it and typical wndula-
tella. None of these (including the type of fumosella)
exhibits any genitalic differences.
The type of obsipella is a freak specimen with vein 4
present in one hind wing. On the strength of this
venation it was referred to the synonymy of Hlasmo-
palpus petrellus by Hulst in 1890 and so appears in our
lists. It is a ragged female with only one hind wing
complete enough to show venation and is labeled
“Colorado” a more probable locality than Texas as
given in the original description. Its genitalia are
those of undulatella.
Genera 101-104: Honora to Cabotia
[Venational division A. Forewing with 11 veins; 10 from cell;
2 from or from very near lower outer angle of cell; 3 closely
approximate to or connate with stalk of 4-5; 4-5 stalked at least
for one-half their lengths. Hind wing with 7—8 anastomosed or
contiguous (Oncolabis) beyond cell; 2 from or from very near
lower outer angle of cell; discocellular vein curved. Labial palpi
oblique. Male genitalia with uncus triangulate; tegumen not
appreciably reduced in proportion to uncus; harpe with short
erect clasper; transtilla incomplete (except in Honorinus) ; aedea-
gus short, stout; penis armed with sclerotized folds or granula-
tions or short spines. Female genitalia with signa consisting of
densely spined round plate, detached thornlike spines or both;
ductus seminalis from bursa copulatrix.]
101. Genus Honora Grote
Honora Grote, Bull U. 8S. Geol. Geogr. Surv. Terr., vol. 4, p. 702,
1878; North Amer. Ent., vol. 1, p. 11, 1879.—Hulst, Phycitidae
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 197
of N. Amer., p. 187, 1890.—Ragonot, Monograph, pt. 2,
p- x, 1901—Hampson, in Ragonot, Monograph, pt. 2, p.
183, 1901. (Type of genus: Honora mellinella Grote.)
Tongue well developed. Antenna simple, pubescent.
Labial palpus oblique, the third segment porrect (de-
flected forward). Maxillary palpus small, squamous,
appressed to face. Forewing smooth; 11 veins; veins
2 and 3 closely approximate and from the lower outer
angle of cell; 4 and 5 stalked, the stalk shortly sepa-
rated from 3 at base; 6 from below upper angle of cell,
straight; 10 from the cell, at base shortly separated
from 8-9; male without costal fold. Hind wing with
vein 2 from the lower outer angle of cell, connate with
3, 3 and 5 stalked; 7 and 8 anastomosed for nearly half
their lengths beyond cell; cell one-third the length of
wing; discocellular vein curved. Eighth abdominal
segment of male with a pair of ventrolateral hair tufts.
Male genitalia with apical process of gnathos a
stout hook. Harpe narrowly elongate; apex rounded
or bluntly pointed; an erect, narrow, short clasper
arising from sacculus near base of cucullus. Anellus a
nearly flat plate with a cupped depression at base and
posterior margin more or less notched. Aedeagus
short, moderately stout, slightly bent (elbowed) near
middle; penis armed with a narrow row of fine, spine-
like cornuti. Vinculum stout, as broad or slightly
broader than long; terminal margin broad.
Female genitalia with signa strongly developed, con-
sisting of a single large, round, curved, densely spined
plate and a scattering of detached spines (except in
dotella) opposite the plate; ductus bursae as long as or
somewhat shorter than length of bursa, finely scobinate
for two-thirds of its length from bursa; a narrow sclero-
tized plate in area above genital opening; ductus semi-
nalis from a lobed projection of bursa near its junction
with ductus bursae.
The genus is apparently confined to North America.
The species perdubiella (Dyar) is referred here from
Zophodia. Another, dulciella (Hulst), described in
Honora, must find placement elsewhere. The genitalia
of the unique female type (fig. 951) show that it is not
a Honora; but its proper generic placement will have
to wait upon discovery of a male. It is treated briefly
at the end of the paper (see p. 313).
The species are very close and the genitalic differences
separating them, especially among the males, are
trifling. The female genitalia show more obvious varia-
tion in the amount of spining in the bursa, but this is a
character of doubtful value. Such differences as exist
are shown in the several drawings.
391. Honora mellinella Grote
Figures 57, 445, 937
Honora mellinella Grote, Bull. U. S. Geol. Geogr. Surv. Terr.,
vol. 4, p. 702, 1890.—Hulst, Phycitidae of N. Amer., p. 188,
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 185,
1901.—McDunnough, Check list, No. 6343, 1939.
Honora ochrimaculella Ragonot, N. Amer. Phycitidae, p. 12,
1887.—Hulst, Phycitidae of N. Amer., pp. 188, 189, 1890.—
Hampson, in Ragonot, Monograph, pt. 2, p. 185, 1901.
Forewing dark grayish fuscous; costal area from
upper margin of cell whitish, in some specimens this
pale costal shading somewhat obscure, in most speci-
mens rather well contrasted against ground color; ante-
medial line nearly vertical, white, distinct and slightly
concave towards inner margin, bordered inwardly by
faint blackish shading and outwardly on inner margin
by a yellowish patch; a similar yellowish patch at base
of wing; subterminal white line, narrow, sinuate, ob-
scure and frequently obliterated; discal dots at end of
cell blackish, the upper in the pale costal area, the lower
in the dark area but normally with a faint, narrow,
whitish border. Hind wing very pale smoky fuscous,
with a silky sheen. Alar expanse, 16-24mm. Male
genitalia with apex of harpe rounded; cucullus not
appreciably broadened toward apex. Female with a
rather dense cluster of spines in bursa opposite spined
plate.
TyprE LOCALITIES: Texas (mellinella, in BM); Cali-
fornia (ochrimaculella, in Paris Mus.).
Foop puant: Palafoxia.
Distrisution: North Carolina, Southern Pines (Aug.,
Sept.); Zouwisiana, Orange, Sabine Parish (June);
Florida, Lakeland (Jan.), Tampa; Teras, Blanco
County, Burnet County, Kerrville (Mar., July, Sept.,
Oct.), New Braunfels (May), San Benito (Mar., Apr.) ;
New Mexico, Hot Springs (Sept.); Arizona, Palmerlee,
Phoenix (Oct.), Redington, Santa Rita Mts. (June,
July); California, Indian Wells (Jan.); Washington,
Pullman (June), Walla Walla (May, June, July),
Yakima (May).
The food plant record given above is from a female
from Indian Wells, Calif., reared by Commander C. M.
Dammers.
392. Honora subsciurella Ragonot
Fieures 447, 941
Honora subsciurella Ragonot, N. Amer. Phycitidae, p. 12, 1887.—
Hulst, Phycitidae of N. Amer., p. 189, 1880.—Hampson, in
Ragonot, Monograph, pt. 2, p. 186, 1901.—McDunnough,
Check list, No. 6347, 1939.
Doubtfully distinct from mellinella except, possibly,
as a local race. Distinguished chiefly by the paler
ground color of forewing beyond antemedial line, due
to a fine powdering of white scales; a broader diffusion
of the pale costal area (usually extended to include the
lower discal spot); the bluntly pointed apex of harpe
(fig. 447); and the sparser spining of the female bursa
(compare figs. 941 and 937). Alar expanse, 22-28 mm.
Typr Locauity: Colorado (type in Paris Mus.).
Foop piant: Unknown.
Distripution: Colorado, Glenwood Springs (Sept.);
Utah, Eureka (June, July, Aug.), Stockton (July, Aug.),
Vineyard (July); California, San Bernardino Mts.
(Sept.).
The “red” ground color of forewing is exaggerated by
Ragonot in his original description and in the figure
(pl. 31, fig. 12) in his Monograph. Hampson’s de-
scription (in Ragonot) is more accurate.
198
393. Honora sciurella Ragonot
Ficure 446
Honora sciurella Ragonot, N. Amer. Phycitidae, p. 12, 1887.—
Hulst, Phycitidae of N. Amer., p. 189, 1890.—Hampson, in
Ragonot, Monograph, pt. 2, p. 185, 1901—McDunnough,
Check list, No. 6346, 1939.
Known to me only from descriptions of Ragonot and
Hampson, the Ragonot figures (plate 37, figs. 16, 17)
and the genitalia of the male type. The harpes of the
latter (fig. 446) are somewhat longer in proportion to
the remainder of the genitalia than in the other species
of the genus. In the National Collection is a female
(24 mm.) from Walla Walla, Wash. (June), that is
almost an exact match, even to the dark fuscous hind
wing, to Ragonot’s figure 17 of the female in Paris. A
larger female (82 mm.) in the National Collection from
Monachee Meadows, Calif., is a good match for
Ragonot’s figure 16, of the male type, except that the
hind wing is whitish towards the base as described by
Hampson. In these two examples the spining of the
bursa is similar to that of montinatatella (fig. 939). The
exact status of sciwrella cannot be determined until it
is reared and more specimens in substantial series are
assembled.
Alar expanse, ‘32 mm.”’
Tyrer Locatity: California (type in Paris Mus.).
Foop pirant: Unknown.
394. Honora dotella Dyar
FIGURES 448, 938
Honora dotella Dyar, Pomona College Journ. Ent., vol. 2, p. 377,
1910.—McDunnough, Check list, No. 6345, 1939.
Distinguished from mellinella by the darker, more
suffused (blackish fuscous) ground color and the more
strongly contrasted whitish costal area of forewing;
antemedial line thin, white, not continued to costa;
ocherous patch beyond antemedial line obsolete or
represented only by a pale trace; basal pale spot clay
colored, concolorous with thorax; subterminal line ob-
solete; upper discal spot obliterated; lower discal spot
black, elongate, completely enclosed by the dark ground
color and with no trace of a white border. Hind wing
semihyaline white with a fuscous shade along termen
and some fuscous shading on the veins, especially in
female. Alar expanse, 17-24 mm.
Male genitalia distinguished chiefly by the somewhat
widened apical portion of cucullus. Bursa of female
genitalia without detached spines opposite the large,
spined plate (signum).
Typr Locatiry: Claremont, Calif. (type in USNM).
Foop prant: Unknown. F
Distrisution: California, Claremont, Loma Linda
(Mar.), San Diego (Mar., May, June).
Apparently a distinct species, but close to mellinella.
395. Honora montinatatella (Hulst)
FicureE 939
Spermatophthora montinatatella Hulst, Ent. Amer., vol. 3, p. 134,
1887.
Honora canicostella Ragonot, N. Amer. Phycitidae, p. 12, 1887.
UNITED STATES NATIONAL
MUSEUM BULLETIN 207
Honora montinatatella (Hulst), Ent. Amer., vol. 5, p. 156, 1889;
Phycitidae of N. Amer., p. 189, 1890.—Hampson, in
Ragonot, Monograph, pt. 2, p. 186, 1901.—McDunnough,
Check list, No. 6348, 1939.
Forewing brownish red with costal area white dusted
with reddish scales, the pale costal color spreading into
the cell at middle of wing and enclosing the blackish
discal spots, broken near middle of costa by a narrow
band of the ground color which slants outwardly into
cell; antemedial line incomplete, a thin line of whitish
scales from inner margin to cell, set further out on wing
than in other species; subterminal line obsolete or nearly
so, when present well back from terminal margin so
that space between antemedial and subterminal lines is
considerably reduced (less than one-third the wing
area); ON some specimens an obscure ocherous patch
bordermg the antemedial line outwardly near inner
margin. Hind wing palesmokyfuscous. Alar expanse,
24-28 mm.
Male genitalia similar to those of mellinella except
vinculum narrower at terminal margin, shape similar to
that of subsciurella (fig. 941).
TYPE LOCALITIES: Sierra Nevada, Calif. (montinata-
tella, i AMNH, ex Rutgers); California (canicostella,
in Paris Mus.).
Foop puant: Unknown.
In addition to the Hulst type I have seen two other
specimens (co' and 9, in USNM), from a subalpine
meadow on Bogachiel Peak of the Olympic Mts., Wash.
Apparently montinatatella is a high-altitude species.
396. Honora perdubiella (Dyar), new combination
FicurEs 449, 940
Zophodia perdubiella Dyar, Proc. Ent. Soc. Washington, vol. 7,
. 37, 1905.
Pie perdubiella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221,
1925.—McDunnough, Check list, No. 6299, 1989.
Pale ashy gray; costal area sparsely dusted with
whitish scales; transverse lines obsolete; lower discal
spot at end of cell faintly indicated, blackish. Hind
wing slightly paler than forewing, unmarked, shiny.
Alar expanse, 22-26 mm.
Male genitalia distinguished chiefly by the shape of
the apical projection of gnathos. Female genitalia
figured from paratype from the type locality; similar
to those of montinatatella, especially the genitalia of the
Mount Olympus female of Hulst’s species.
TypE LocaLiry: Stockton, Utah (June, type in
USNM).
Foop pLiant: Unknown.
Apparently a distinct species. Dyar’s types are
rubbed so the extremely pale color of forewing may not
be true of fresh examples. A female in the National
Collection from Baker, Nev. (May), is much darker,
shows faint traces of the normal transverse lines and
indications of the blackish discal and terminal dots.
However, there is scarcely any whitish dusting in the
costal area. The forewing is dark grayish fuscous and
the hind wing fuscous, with a faintly darker line along
termen and some dark shading on the veins.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 199
102. Honorinus, new genus
TyprE or genus: Honorinus fuliginosus, new species.
Characters of Honora except: Labial palpus of male
grooved to hold the maxillary palpus; maxillary palpus
of male in the form of an aigrette; forewing with vein 2
from before but near outer angle of cell; hind wing with
cell shorter, somewhat less than one-third the length of
wing; transtilla complete; abdomen of male with two
pairs of ventrolateral hair tufts.
Very close to Honora. In the absence of any typical
species of Honora from tropical America and because of
the divergence of the Peruvian species on male charac-
ters it seems better to give it a separate generic desig-
nation than to expand and loosen our definition of
Honora.
397. Honorinus fuliginosus, new species
Figures 450, 947
Forewing sooty fuscous; transverse lines obsolete or
nearly so; antemedial line faintly indicated in the male
near inner margin; beyond it a faint ocherous shade;
some pale scaling between the veins beyond cell; a
narrow, obscure ocherous shade along costa from base
to middle; discal and terminal dots obscure. Hind
wing semihyaline, whitish fuscous with a dark line
along termen and some dark shading on the veins;
veins 3 and 5 stalked for more than half their lengths.
Alar expanse, 24-26 mm.
Male genitalia with elements of transtilla solidly
fused at their apices to form a high, narrowly rounded
arch. Female genitalia with the membrane of posterior
lobe of bursa thickened.
Typ Locatity: Angasmarca, Pert (type in USNM).
Foop prant: Unknown.
Described from male type and three male and one
female paratypes, all from the type locality. In addi-
tion I have before me another female from Mima, Pert,
which I take to be the same species; but as it has veins
3 and 5 of hind wing somewhat longer stalked and a
slightly smaller genitalia than the female paratype I do
not include it among the types. The males are rubbed
and do not show the dark ground color as plainly as the
female, which is in good condition.
103. Genus Oncolabis Zeller
Oncolabis Zeller, Isis von Oken, p. 877, 1848.—Ragonot, Mono-
graph, pt. 1, pl. 1, fig. 26, 1893; pt. 2, p. xi, 1901—Hampson,
in Ragonot, Monograph, pt. 2, p. 187, 1901. (Type of genus:
Oncolabis anticella Zeller.)
Endommasis Hampson, in Ragonot, Monograph, pt. 2, pp. ix,
124, 1901. (Type of genus: Endommasis nigritella Hamp-
son. New synonymy.)
Tongue well developed. Antenna pubescent; basal
segment elongate; on male, shaft with a stout hook from
its basal segment and strongly curved for a few seg-
ments beyond (fig. 454d); on female simple. Labial
palpus obliquely ascending; long, extending half its
length above vertex; smooth scaled; segment 2 in male
grooved to hold maxillary palpus; segment 3 very short,
acuminate, not deflected forward. Maxillary palpus
of male in the form of an aigrette; of female filiform.
Forewing smooth; narrowly elongate; 11 veins; vein 2
from near lower outer angle of cell; 3 from the angle;
4-5 stalked for half their lengths, their stalk very closely
approximate or connate with 3 at base; 6 from below
upper angle of cell, straight; 10 from cell, closely ap-
proximate to the stalk of 8-9 for a short distance; male
without costal fold. Hind wing with vein 2 from very
close to lower outer angle of cell; 3 and 5 stalked for
half their lengths; 7 and 8 contiguous or closely approxi-
mate for a short distance beyond cell; cell one-third the
wing length; discocellular vein curved. Eighth ab-
dominal segment of male with compound ventral tufts.
Male genitalia with apical process of gnathos a
stout, elongate, pointed hook. Uncus triangulate; apex
pointed. Harpe elongate, slender; apex rounded;
clasper vertical, pointed. Anellus a small triangulate
cupped plate without lateral lobes. Aedeagus stout,
straight, moderately long; penis armed with a few short
sclerotized folds and numerous granulations. Vin-
culum stout, a trifle longer than broad, not appreciably
tapering to broad terminal margin.
Female genitalia with signa strongly developed,
consisting of a large, round, curved, densely spined
plate and a varying number of detached spines opposite
the plate; ductus bursae about the same length as
bursa, with a ring of sclerotized spinose ridges at its
junction with bursa, minutely granulose for a short
distance beyond; genital opening simple; ductus
seminalis from bursa near its junction with ductus
bursae.
Hampson distinguished his Hndommasis from
Oncolabis chiefly on the shape of the labial palpus,
upturned (‘‘redressé”) in Hndommasis and obliquely
erect with third segment projected forward in Oncolabis.
On the strength of this supposed difference the genera
were widely separated in the generic key in Ragonot’s
Monograph (pp. ix and xi). This difference was solely
due to the position of the palpi at time of death of the
particular specimens. The normal position of the labial
palpus is appressed to face with third segment erect.
Similar male antennae and female genitalia occur in
Stylobasis Hampson. The latter, however, is distin-
guished by its forewing venation, vein 4 being absent.
398. Oncolabis anticella Zeller
Figures 56, 454, 942
Oncolabis anticella Zeller, Isis von Oken, p. 877, 1848.—Hampson,
in Ragonot, Monograph, pt. 2, p. 187, 1901.
Endommasis nigritella Hampson, in Ragonot, Monograph, pt.
2, p. 124, 1901 (new synonymy).
Forewing dark brownish fuscous with basal area
darker, blackish; costal border (except in suffused
specimens) whitish finely dusted (under magnification)
with scattered reddish scales; antemedial line obsolete,
indicated only by a white spot extending from lower
fold to inner margin and having a small, central, black
dot; in fresh specimens the white spot narrowly
bordered inwardly and outwardly by a faint ocherous
shading. Suffused specimens show little or no trace of
200
the white spot or the white border of costa. Hind
wing hyaline white with a narrow fuscous shade along
termen and a similar shading on some of the veins,
especially on the females. Alar expanse, 13-21 mm.
Male genitalia with lateral margins of vinculum
slightly concave; terminal margin very slightly angled;
clasper sharply pointed. Female genitalia exhibiting
considerable variety in size of bursa, size of the large
spined plate of signa, and the number of scattered spines
opposite the spined plate. Extremes of variation are
shown in figure 942.
TYPE LocauitiEs: South Brazil (anticella, in BM);
Santos, Brazil (ngritella, in BM).
Foop pirant: Hlephantopus sp.
DisTRIBUTION: CuBA: Santiago de las Vegas (Sept.).
GuatemaLa: Cayuga (Aug.), Chejel (Aug.), Volcdin
Santa Maria (July). Costa Rica: La Florida, Juan
Vinas (May, Nov.). Panamé: Corozal (Apr.), La-
Chorrera (May), Parafso (May), Porto Bello (Apr.,
May, Oct., Dec.), Rio Trinidad (Mar.). FREencu
Guiana: St. Jean Maroni. Contompra: Minca.
Bottvia: Santa Cruz, Prov. del Sara (Jan., Nov.);
Brazit: Sao Paulo, Amparo, Santos, Ypianga (Sept.) ;
Santa Catarina, Santa Catarina Isl. Ecuapor:
Quevedo (Nov. Dec.). Paraguay: Villarrica (July,
Aug., Sept.). Argrnrina: “Villa Ana, F. C. S. F.”
(Jan.).
Over 60 specimens before me from the National
Museum, British Museum, Cornell, and Janse Collec-
tions exhibiting wide variation in size and color, from
suffused examples corresponding to the figure of
nigritella in the Ragonot Monograph (pl. 49, fig. 20)
to that of typical anticella (pl. 48, fig. 25). The male
genitalia are remarkably uniform. Female genitalia
vary in individual specimens as indicated above. Such
variations bear no relation to locality or pattern. The
food plant record is from two reared specimens (o" and @)
in the National Museum from Santiago de las Vegas
(HE. H. A. de Cuba, No. 9627). This is our only known
host record.
104. Genus Cabotia Ragonot
Cabotia Ragonot, Nouv. Gen., p. 30, 1888; Monograph, pt. 2,
pp. xi, 187, 19801.—Hampson, in Ragonot, Monograph,
pt. 2, p. 188, 1901. (Type of genus: Cabotia semidiscella
Ragonot.)
Encystia Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7, p. 256,
1901. (Type of genus: Encystia bonhoti Hampson. New
synonymy.)
Tongue well developed. Antenna pubescent; basal
segment elongate; on male shaft compressed and
strongly curved at base (fig. 452d); on female shaft
simple. Labial palpus oblique; segment 2 reaching
almost to vertex, in male grooved to hold the maxillary
palpus; segment 3 short, deflected forward or slightly
downward. Maxillary palpus of male in the form of an
aigrette; of female filiform, somewhat broadly scaled.
Forewing smooth; 11 veins; 2 from very close to lower
outer angle of cell; 3 from the angle; 4-5 stalked for at
UNITED STATES NATIONAL MUSEUM BULLETIN 207
least half their lengths, stalk connate with 3; 6 from
below upper angle of cell, straight; 10 from the cell,
at base shortly separated from 8-9; male without
costal fold. Hind wing with vein 2 from lower outer
angle of cell, connate with 3; 3 and 5 long stalked (for
over half their lengths); 7 and 8 anastomosed for
slightly more than half their lengths beyond cell; cell
about one-third the length of wing; discocellular vein
curved. Highth abdominal segment of male with a pair
of ventrolateral hair tufts.
Male genitalia with apical process of gnathos an ovate
plate bearing a small thornlike spine near its posterior
end. Uncus triangulate; apex pointed. Harpe elon-
gate; apex evenly rounded; clasper vertical, knobbed.
Anellus a triangulate, cupped plate. Aedeagus short,
stout, sharply angled at apex; penis with a few scattered
granulations, otherwise unarmed. Vinculum stout, as
long as or a trifle longer than broad, truncate, and of
nearly equal width throughout.
Female genitalia with signa well developed, consisting
of a chain of short thornlike spines on bulbous bases and
a varying number of similar, scattered, discontinuous
spines; bursa, otherwise smooth; ductus bursae mem-
branous; genital opening simple, unsclerotized; ductus
seminalis from bursa near its junction with ductus
bursae.
The genus is compact and sharply defined, easily
distinguished from nearly related genera by its male
antenna, signa, and the shape of the terminal projection
of gnathos. Hampson’s description of Encystia would
indicate something entirely different, but the description
is erroneous in a number of details: vein 9 of forewing is
not absent, but well developed; vein 2 of forewing is
close to, but not from the angle, and the discocellular
vein of hind wing is curved and not oblique. I have
before me two females of his series of bonhoti from
Nassau and photographs of his holotype and its male
genitalia, which clearly show that it is a typical Cabotia.
Specific limits within the genus are difficult to deter-
mine from the limited and scattered material available.
I suspect that most of the names will eventually fall
into synonymy. The genitalic differences exhibited by
the supposed species are trifling, consisting chiefly of
the terminal plate of gnathos and the number of spines
composing the signa, all very doubtful characters sub-
ject to individual variation. The coloration and pat-
tern—likewise somewhat individually variable in speci-
mens from a given locality—are much the same for all
the species except Dyar’s rhythmatica: Forewing luteous
more or less shaded with fuscous; antemedial line very
faintly indicated, well out towards middle of wing,
bordered outwardly at or near costa by a fuscous blotch
and inwardly near inner margin by a similar shade;
subterminal line obscure, parallel and close to termen;
costal area whitish; a sparse scattering of reddish scales
over most of the wing; discal dots faint; terminal dots
distinguishable except at tornus and near apex. Hind
wing pale smoky fuscous, proportionally darker in dark
females.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 201
399. Cabotia semidiscella Ragonot
Figure 55, 451, 935
Cabotia semidiscella Ragonot, Nouv. Gen., p. 30, 1888.—Hamp-
son, in Ragonot, Monograph, pt. 2, p. 188, 1901.
The alar expanse given by Ragonot is 16 mm. In
the National Museum are three smaller specimens (11
mm.) identified by Hampson—a male and female from
St. Jean Maroni, French Guiana (the genitalia of which
are figured), and a male from St. Laurent du Maroni,
French Guiana. This last is an abnormal specimen,
having veins 2 and 3 of forewing long stalked. Also in
the National Collection are six females from Los Vas-
quez, Argentina (15-16 mm.), agreeing with the fore-
going, and two somewhat darker (fresher) females that
I take to be the same species from Villarrica, Paraguay
(Feb., Oct.). Inthe British Museum there is a male and
nine females from southeastern Brazil (E. D. Jones,
1920-303”) and three females from Villa Ana, Ar-
gentina (Oct.).
Typr Locauity: Goya, Argentina (type in Paris
Mus.).
Foop pirant: Unknown.
400. Cabotia schini (Berg)
Spermatophthora schini Berg, Anales Soc. Cient. Argentina, vol.
19, p. 275, 1885.
Cabotia schini (Berg) Ragonot, Nouv. Gen., p. 30, 1888.—Hamp-
son, 7n Ragonot, Monograph, pt. 2, p. 188, 1901.
I have seen nothing identified as this species. How-
ever, it should be readily identifiable if Argentinian
examples of Cabotia are ever reared from the pepper-
tree. Alar expanse, 20 mm.
Type Locaity: Buenos Aires, Argentina (type lost).
Foop piant: Schinus molle Linnaeus.
Apparently known only from the type specimens
reared from galls on the peppertree. This is our only
food-plant record for the genus.
401. Cabotia rhythmatica Dyar
Ficures 453, 933
Cabotia rhythmatica Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 339,
1914,
Forewings less distinctly marked with fuscous and
with a more rosy suffusion than those of other species
inthe genus. Uncus shorter and male genitalia stockier
than in our examples identified as semidiscella or than in
males of bonhoti. Alar expanse, 13-14 mm.
Typr Locauity: Porto Bello, Panamdé (Mar., Apr.;
type in USNM).
Foop prant: Unknown.
Known only from the type specimens.
402. Cabotia cundajensis (Zeller)
Fiaure 932
Euzophera cundajensis Zeller, Horae Soc. Ent. Rossicae, vol. 16,
p. 227, 1881.
Euzophera impeditella Zeller, Horae Soc. Ent. Rossicae, vol. 16,
p. 229, 1881.
Cabotia cundajensis (Zeller) Ragonot, Monograph, pt. 2, p. 189,
1901.
I have seen no Cabotia from Colombia; but in the
National Museum are three females (15-20 mm.) from
Castro, Parand, Brazil, identified by both Hampson
and Schaus as cundajensis. The genitalia are figured
from one of these. They exhibit nothing that can be
definitely identified as a specific character. Alar ex-
panse, 16-22 mm.
Types: In British Museum (cundajensis, impeditella).
TypE LocauitiEes: Cundai, Colombia (cundajensis, in
BM); Viani, Colombia (impeditella, in BM).
Foop prant: Unknown.
I suspect that this as well as semidiscella will eventu-
ally prove to be the same as schini (Berg).
403. Cabotia bonhoti (Hampson), new combination
Figures 452, 934
Encystia bonhoti Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7,
p. 256, 1901.
Not appreciably different from what we have identi-
fied as semidiscella Ragonot from French Guiana,
except for trifling differences in genitalia which are
probably not of more than individual significance.
Male genitalia figured from specimen from Trelawney
Parish, Jamaica; female genitalia from Zeller specimen
in British Museum, from Nassau. Alar expanse,
13-16 mm.
Typr Locatity: Nassau, Bahamas (type in BM).
Foop piant: Unknown.
Distrisution: Bawamas: Nassau (July). Jamaica:
Newport (Feb.), St. Andrews Parish, Trelawney Parish.
The Jamaican material consists of 36 specimens in the
National Collection.
Genus 105: Canarsia
[Venational division A. Forewing with 11 veins; 10 from the
stalk of 8-9; 2 from well before lower outer angle of cell; stalk
of 4-5 separate from 3 at base. Hind wing with 7 and 8 anas-
tomosed for a short distance beyond cell; vein 2 from before
lower outer angle of cell; discocellular vein curved. Labial
palpus upecurved. Male genitalia with uncus triangulate; harpe
with short erect clasper; transtilla absent; aedeagus short, stout;
penis armed with several short spines. Female with a girdle of
slender spines in bursa; bursa partially sclerotized; ductus
seminalis from bursa couplatrix.]
105. Genus Canarsia Hulst
Canarsia Hulst, Phycitidae of N. Amer., p. 179, 1890.—Ragonot,
Monograph, pt. 2, p. ix, 1901—Hampson, in Ragonot,
Monograph, pt. 2, p. 119, 1901. (Type of genus: Ne-
phopteryx ulmiarrosorella Clemens.)
Tongue well developed. Antenna pubescent; on
male shaft with a deep sinus and strong scale tuft at
base; on female simple. Labial palpus upcurved, rough
scaled, reaching to vertex; third segment short (less
than half the length of second), pointed. Maxillary
palpus squamous, appressed to face. Forewing smooth;
11 veins, vein 2 from well before lower outer angle of
cell; 3 from angle; 4 and 5 stalked, the stalk well sepa-
rated from 3; 6 from below upper angle of cell, straight;
10 stalked with 8-9; male without costal fold. Hind
202 UNITED STATES NATIONAL MUSEUM BULLETIN 207
wing with vein 2 before lower outer angle of cell; 3 from
angle, closely approximate for some distance with 5;
7 and 8 anastomosed for a short distance beyond cell;
cell about one-half the wing length; discocellular vein
curved. Highth abdominal segment of male with
compound ventral hair tufts.
Male genitalia and apical process of gnathos a stout
hook. Elements of transtilla not distinguishable (un-
sclerotized); uncus triangulate; apex narrowly rounded.
Harpe narrowly elongate, of nearly even width through-
out; apex rounded; a small erect blunt clasper arising
from sacculus near middle of harpe. Anellus a rather
large flattened plate with prominent lateral lobes.
Aedeagus short, stout, strongly elbowed near middle
and with a scattering of fine spines toward apex; penis
armed with a short row of spinelike cornuti. Vinculum
stout, about as broad as long; tapering slightly to
broadly rounded terminal margin.
Female genitalia with signa consisting of a girdle of
long, slender spines with enlarged bases; bursa copu-
latrix partially sclerotized at junction with ductus
bursae and emergence of ductus seminalis; ductus
bursae short (shorter than bursa), sclerotized just before
genital opening, otherwise simple; genital opening
simple; ductus seminalis from bursa near its junction
with ductus brusae.
An easily recognized genus containing one North
American species.
404. Canarsia ulmiarrosorella (Clemens)
Figures 80, 455, 936
Nephopteryx (?) ulmiarrosorella Clemens, Proc. Acad. Nat. Sci.
Philadelphia, 1860, p. 205.
Stenoptycha pneumatella Hulst, Ent. Amer., vol. 3, p. 137, 1887.
Psorosa ulmella Ragonot, N. Amer. Phycitidae, p. 13, 1887.
Honora fuscatella Hulst, Ent. Amer., vol. 4, p. 118, 1888.
Canarsia ulmiarrosorella (Clemens) Hulst, Phycitidae of N.
Amer., p. 180, 1890; U. S. Nat. Mus. Bull. 52, p. 431,
1902.—Hampson, in Ragonot, Monograph, pt. 2, p: 120,
1901.—Forbes, Cornell Mem. 68, p. 632, 1923—MeDun-
nough, Check list No. 6332, 1939.—Darlington, Trans.
Amer. Ent. Soc., vol. 78, p. 91, 1947.—Craighead, U. S.
Dep. Agr. Mise. Publ. 657, p. 454, 1950.
Canarsia gracilella Hulst, Canadian Ent., vol. 32; p. 174, 1900.—
Forbes, Cornell Mem. 68, p. 632, 1923—McDunnough,
Check list, No. 6333, 1939. (New synonymy.)
Canarsia feliculella Dyar, Journ. New York Ent. Soc., vol. 15, p.
110, 1907.—McDunnough, Check list No. 6334, 1939. (New
synonymy.)
Forewing pale to dark gray; antemedial line well out
towards middle of wing, nearly vertical, slightly in-
dented at vein 1b and above the lower fold, white bor-
dered outwardly by a blackish line; subterminal line
parallel with termen, indented at vein 6 and lower fold
and outwardly angled between them, white, inwardly
bordered by a blackish line; discal spots confluent, form-
ing a blackish line along discocellular vein; terminal
dots confluent forming a more or less continuous black
line along termen. Hind wings pale smoky fuscous,
somewhat darker on dark specimens. Alar expanse,
15-20 mm.
Genitalic characters as given for the genus, the thin
spines of the signa slightly bent.
TypEz Locauities: None indicated (ulmiarrosorella,
type lost; pnewmatella, in AMNH, ex Rutgers) ; Wiscon-
sin (ulmella, in Paris Mus.); Blanco County, Tex.
(fuscatella, in AMNH, ex Rutgers); Montclair, N. J.
(gracilella, rn AMNH, ex Rutgers); Brownsville, Tex.
(feliculella, nm USNM).
Foop pLant: Elm (larva a leaf-tier). Also recorded
from hickory (Forbes). In the National Museum there
is a series reared from larvae taken under bark of hack-
berry; but as larvae wander about after feeding, records
other than elm should be discounted.
Distrisution: Unitep Starrs: Maine; New Hamp-
shire, Hampton (July); Massachusetts, Amherst (June,
July), Framingham (May); Connecticut, East River,
New Haven (Aug.) ; New York, Otto (July) ; New Jersey,
Essex County (June), Montclair (Aug.), New Bruns-
wick; Pennsylvania, New Brighton (May), Pittsburgh
(May); Maryland, Hyattsville, Plummers Isl. (May);
District of Columbia, Washington (Mar., Apr., May);
Ohio, Dayton (Aug.); Illinois, Chicago, Decatur (Apr.,
June), Lacon (Aug.), Oconee (Aug.); Wisconsin; Iowa,
Ames (May), Sioux City (May, July); Kansas, Law-
rence (Aug.), Manhattan (May), Onaga; Tennessee,
Knoxville (May); Missouri, St. Louis (June, Aug.) ;
Texas, Blanco County (May, June), Brownsville (May),
Burnet County, Kerrville (Apr.), Plano (July), San
Benito (Aug., Sept.), Victoria (May). Canapa: Ontario,
Trenton (June, July); Quebec, St. Hilaire (June, July) ;
Nova Scotia, Cape Breton Isl. (July).
Presumably generally distributed east of the Rocky
Mountains wherever the elm occurs.
The names gracilella and feliculella represent nothing
more than pale color forms and have no racial signifi-
cance. The holotype of gracilella is a male without ab-
domen, but. is obviously conspecific with ulmiarrosorelia.
The female paratype (also in the Rutgers Collection) is
an Hiphestia. Dyar’s type is a male agreeing in all de-
tails of genitalia with typical wlmiarrosorella.
Genus 106: Harnocha
[Venational division C. Forewing with 10 veins; 9 absent; male
without costal fold. Hind wing with vein 2 from angle of cell;
3 and 5 stalked; discocellular vein curved. Labial palpi porrect.
Transtilla incomplete.]
106. Genus Harnocha Dyar
Harnocha Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 337, 1914.
(Type of genus: Harnocha velessa Dyar.)
Tongue well developed. Antenna pubescent, shaft
very slightly bent at base. Labial palpus porrect,
downcurved; extending slightly over twice the length
of the head beyond it. Maxillary palpus minute, fili-
form. Forewing smooth; 10 veins; vein 2 from very
close to lower outer angle of cell, approximate to 3; 3
from the angle; 4 and 5 stalked for half their lengths,
the stalk connate with 3; 6 from below upper angle of
cell, straight; 8 and 9 united (9 absent); 10 from the
cell, shortly separated from 8 at base; male without cos-
tal fold. Hind wing with vein 2 from lower outer angle
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
of cell, connate with 3; 3 and 5 stalked for at least half
their lengths; 7 and 8 anastomosed for at least half their
lengths beyond cell; cell less than one-half (nearer one-
third) the length of the wing; discocellular vein curved.
Eighth abdominal segment of male simple.
Male genitalia with apical process of gnathos an ovate
plate terminating in a spinelike point. Uncus triangu-
late; apex bluntly pointed. Harpe narrowly elongate;
apex rounded; clasper absent. Anellus a triangulate,
flattened plate. Aedeagus moderately stout, sinuate
(slightly bent at one-third and again at two-thirds);
penis with a few weak scobinations, otherwise unarmed.
Vinculum stout, as broad as long, not tapering; terminal
margin evenly rounded.
This genus is distinct from but apparently very close
to Cabotia, with which it agrees on a great many struc-
tural details of venation and male genitalia. It differs
in having the antennal shaft less strongly bent, the
labial palpus porrect rather than oblique, vein 9 of fore-
wing absent, eighth abdominal segment of male without
paired hair tufts, and harpe without clasper. It con-
tains but one known tropical American species.
405. Harnocha velessa Dyar
Ficures 111, 456
Harnocha velessa Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 338,
1914.
Forewing luteous with a very faint rosy tint; costa
from base to apex rather broadly margined with white
very sparsely dusted with dark fuscous scales; an
oblique blackish band near base and extending from
inner margin to white costal stripe; a rather conspicuous
patch of blackish scales near middle, between cell and
lower fold; a similar smaller blackish spot at lower outer
angle of cell. Hind wing semihyaline, whitish, shaded
with pale fuscous along costa and at apex. Alar ex-
panse, 11-13 mm.
Typp tocaniry: La Chorrera, Panamd (May, type
in USNM).
Foop puant: Unknown.
Represented in the National Collection by the type,
eight other males from the type locality, and one male
from Rio Trinidad, Panama (June). The female is
unknown.
Genera 107-109: Eurythmasis to Wunderia
{Venational division B. Forewing with 10 veins; 9 present; 2
and 3 stalked or united, from lower outer angle of cell. Hind
‘wing with 2 from lower outer angle of cell; 3 and 5 stalked for at
least half their lengths; 7 and 8 strongly anastomosed. Uncus
triangulate. Transtilla incomplete.]
107. Genus Eurythmasis Dyar
Eurythmasis Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 338, 1914.
(Type of genus: Hurythmasis ignifatua Dyar.)
Tongue well developed. Antenna pubescent; basal
segment elongate; on male shaft with a hook from its
203
basal segment and curved for a few segments beyond;
on female simple. Labial palpus oblique, long, extend-
ing more than half its length above vertex; smooth;
segment 2 on male grooved to hold the maxillary palpus;
segment 3 very short, acuminate. Maxillary palpus
of male in the form of an aigrette; of female filiform.
Forewing smooth; 10 veins; veins 2 and 3 long stalked,
from lower outer angle of cell; 4 absent; 5 separated
from 2-3 at base; 6 from below upper angle of cell,
straight; 8 and 9 stalked; 10 from cell, not approximate
to 8-9; male without costal fold. Hind wing with vein
2 from lower outer angle of cell, connate with the stalk
of 3-5; 3 and 5 stalked for half their lengths; 7 and 8
anastomosed for half their lengths; cell one-third the
wing length; discocellular vein curved. Highth ab-
dominal segment of male with compound ventral tufts.
Male genitalia with apical process of gnathos an
elongate, rather narrow hook. Uncus triangulate; apex
pointed. Harpe elongate, slender; apex rounded;
clasper vertical, pointed, slightly curved. Anellus a
small, weakly sclerotized, cupped plate. Aedeagus
moderately stout, nearly straight; penis with a few
minute spines and granulations, otherwise unarmed.
Vinculum stout, a trifle longer than broad, not appre-
ciably tapering to broad terminal margin.
Female genitalia with signa strongly developed, con-
sisting of a large, curved, oval (or round), densely spined
plate and a few detached spines near or opposite the
plate; ductus bursae approximately the same length as
bursa, minutely scobinate for a short distance from
junction with bursa; genital opening simple; ductus
seminalis from bursa near its junction with ductus bursae.
In habitus and all structural characters except vena-
tion this genus resembles Oncolabis, to which it is ap-
parently closely related. It contains one tropical
American species.
406. Eurythmasis ignifatua Dyar
Ficures 95, 457, 945
Eurythmasis ignifatua Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 338, 1914.
Forewing gray brown; a white border along costa
sparsely dusted with reddish scales; on middle of inner
margin a narrowly elongate, dull ocherous patch with a
knoblike projection at each end extending to the cell;
a few reddish scales on dark ground color at base of
wing and bordering inner margin; transverse lines ob-
solete. Hind wing semihyaline; in female with some
fuscous shading on the veins. Alar expanse, 13-14 mm.
Genitalia as given for the genus; male with terminal
margin of vinculum slightly angled.
Type tocatiry: La Chorrera, Panamaé (type in
USNM).
Foop prant: Unknown.
DistriputTion: PanamA: La Chorrera (May). Purr-
to Rico: Adjuntas (June), Bayamén (Mar., Aug.),
Lavis (Oct.), San German (Apr., Aug.), Utuado. Cusa:
Oriente, Loma del Gato (Sept.).
204 UNITED STATES NATIONAL MUSEUM BULLETIN 207
108. Genus Eurythmidia Ragonot
Eurythmidia Ragonot, Monograph pt. 2, p. xii, 1901. Hampson,
in Ragonot, Monograph, pt. 2, p. 208, 1901. Janse (in
part), Journ. Ent. Soc. South Africa, vol. 7, p. 4, 1944.
(Type of genus: Hurythmia ignidorsella Ragonot.)
Tongue well developed. Antenna pubescent; on
male, a row of 4 or 5 rough scales on shaft beyond basal
segment; of female simple. Labial palpus, upturned,
extending well above vertex; third segment about half
the length of second, pointed. Maxillary palpus
squamous, appressed to face. Forewing smooth; 10
veins; veins 2 and 3 united, from lower outer angle of cell,
connate with the stalk of 4-5; 4 and 5 stalked for at
least half their lengths; 6 from below upper angle of
cell, straight; 8 and 9 stalked; 10 from cell approximate
to 8-9 at base, thence divergent; male without costal
fold. Hind wing with vein 2 from the lower outer angle
of cell, connate with the stalk of 3-5; 3 and 5 long
stalked (for more than half their lengths); 7 and 8
anastomosed for nearly two-thirds of their lengths; cell
short, less than one-third the wing length; discocellular
vein curved. Eighth abdominal segment of male simple.
Male genitalia with apical process of gnathos an
elongate, narrow hook. Uncus triangulate; apex nar-
rowly rounded. Harpe elongate, slender; apex rounded;
clasper vertical, short, pointed, weakly sclerotized.
Anellus a flattened plate. Aedeagus moderately slen-
der, straight; penis unarmed. Vinculum stout, as
broad as long, not tapering, truncate.
Female genitalia with bursa copulatrix elongate;
signa strongly developed, consisting of a curved, round,
densely spined plate and a few detached spines near the
plate; ductus bursae somewhat shorter than bursa,
minutely scobinate near junction with bursa, the sco-
binations extending into bursa for a short distance; a
narrow sclerotized plate behind (and above) genital
Opening; genital opening otherwise simple; ductus
seminalis from bursa near its junction with ductus
bursae.
The genus is obviously closely related to Hurythmasis,
with similar habitus and genitalia, differing chiefly in
venation, the simple eighth segment of male, and the
simple shaft of the male antenna. It contains one
species common to the southwestern United States and
Central America. I mterpret the venation of forewing
differently from Hampson and Ragonot. They con-
sider that 4 is absent and 3 and 5 stalked. From the
affinities of the genus I believe that 4 is present and
stalked with 5 and 2 and 3 united. By either interpre-
tation the end result would be the same; but Huryth-
midia seems to belong with the genera where the
tendency is for 2 and 38 to fuse.
The description and figures published by Janse apply
to Hurythmidia only so far as the female characters are
concerned. The two specimens from Amula, México,
which he had before him were from the Druce Collection
and bore Hampson’s identification labels. The female
is Hurythmidia ignidorsella but the male is not. It is
a somewhat rubbed example of Elasmopalpus lignosellus
Zeller. Evidently Hampson did not check the vena-
tion, for it is normal for Hlasmopalpus, vein 4 being
present in both hind wings. Dr. Janse kindly sent me
the specimens and his slides for examination and has
asked me to make the correction to his description.
407. Eurythmidia ignidorsella (Ragonot)
Ficures 91, 459, 943
Eurythmia ignidorsella Ragonot, N. Amer. Phycitidae, p. 16,
1887.—Hulst, Phycitidae of N. Amer., p. 196, 1890.
Eurythmidia ignidorsella Hampson, in Ragonot, Monograph, pt.
2, p- 208, 1901.—Dyar, Proc. U. S. Nat. Mus., vol. 47, p.
338, 1914—McDunnough, Check list, No. 6361, 1939.
Forewing blackish gray; a white border along costa,
sparsely dusted with reddish scales; on middle of inner
margin a whitish orange patch, the orange shade ex-
tending more or less along inner margin toward base of
wing; antemedial line obsolete; subterminal line very
faint (frequently absent), when distinguishable, whitish.
Hind wing semihyaline, more or less shaded with fuscous
along costa and outer margin. Alar expanse, 12-16
mm.
Genitalia as given for the genus; terminal margin of
vinculum straight; spined plate of signa rather small.
Typr LocaLiry: Arizona (type in Paris Mus.).
Foop piant: Unknown.
Distrisution: Unirep Staves: Arizona, Paradise
(Sept.), Prescott (Sept.), M#xtco: Orizaba, Guerrero,
Amula (May), Cayuga (Apr., May, Aug.), Panamé:
Alhajuelo (Mar., Apr.), Cafio Saddle (Gatin Lake,
May), Corazal (Mar.), Paraiso (Apr.), Porto Bello
(Feb., Mar., May). Reported from additional locali-
ties in México by Ragonot and Hampson.
Strikingly similar in color and markings to Huryth-
masis ignifatua but easily separable on genitalia and
venation.
109. Genus Wunderia Grossbeck
Wumnderia Grossbeck, Bull. Amer. Mus. Nat. Hist., vol. 37, art. 1,
p. 133, 1917. (Type of genus: Wunderia neaeriatella Gross-
beck.)
Tongue well developed. Antenna of female simple,
weakly pubescent. Labial palpus slender, upturned,
reaching above vertex, third segment acuminate,
slightly more than half as long as second. Forewing
smooth; 10 veins; veins 2 and 3 united, from lower
outer angle of cell, connate with the stalk of 4-5; 4 and 5
stalked for half their lengths; 6 from below upper angle
of cell, straight; 8 and 9 stalked; 10 from the cell,
separate from 8-9 at base. Hind wing with vein 2
from the stalk of 3-5; 3 and 5 long stalked; 7 and 8
anastomosed for nearly two-thirds of their lengths; cell
open (discocellular vein absent).
Female genitalia with bursa copulatrix without sig-
num and simple except for some minute scobinations at
its posterior end, the scobinations extending into the
ductus bursae for over half its length; ductus bursae
with a narrow sclerotized plate behind (and above)
genital opening; genital opening otherwise simple;
ductus seminalis from bursa near its junction with
ductus bursae.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
The genus and its type species were described from
a single female. The male is unknown, so the above
diagnosis is incomplete. The habitus of neaeriatella is
that of a pale ignidorsella so the placement of Wunderia
near Hurythmidia seems safe enough.
The genus is readily identified by the open cell of hind
wing, the otherwise Hurythmidia venation, and the
absence of asignum. Grossbeck’s description is faulty
in one respect. The front is not “projected forward in
the form of a cone.”’ The scaling projects in a conical
tuft, but the front itself is evenly rounded.
408. Wunderia neaeriatella Grossbeck
Figures 90, 948, 949
Wunderia neaeriatella Grossbeck, Bull. Amer. Mus. Nat. Hist.,
vol. 37, art. 1, p. 133, 1917—McDunnough, Check list,
No. 6361-1, 1939.
Markings and color of forewing similar to those of
Eurythmidia ignidorsella except slightly paler. The
female type is somewhat rubbed, which accounts for
the traces of a dark transverse antemedial line and the
dark discal markings mentioned by Grossbeck. Before
me are three females from the Cornell Collection, col-
lected at San German, Puerto Rico, Apr. 16 and 17,
1930. In these the entire area below the whitish costal
stripe is pale grayish fuscous without transverse lines
or appreciable discal spots, and there is a very faint
orange blotch on middle of dorsal margin (as in 2gni-
dorsella) distinguishable on one of the specimens. The
whitish costal stripe is sparsely dusted with reddish
scales and there is a scattering of similar scales on the
dark area. The Puerto Rican examples are certainly
congeneric and I believe conspecific with the Florida
type.
There are some differences in the female genitalia
(shown in figs. 948, 949); but these are merely in the
size and shape of the bursa, differences which are prob-
ably of no more than individual significance. The
Puerto Rican specimens are rather small (12-12.5 mm.)
compared with the type (14.5 mm.).
Type wocauity: Everglades, Fla. (Apr., type in
AMNB#).
Foop piant: Unknown.
The male is unknown.
Genera 110-114: Oedothmia to Cacozophera
[Venational division D. Forewing with 10 veins; 9 present; 4
absent; vein 2 from before but near lower outer angle of cell.
Hind wing with discocellular vein curved. Labial palpi up-
turned or oblique. Transtilla incomplete or altogether absent]
110. Genus Oedothmia Hampson
Oedothmia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 60,
1930. (Type of genus: Oedothmia endopyrella Hampson.)
Synothmia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 61,
1930. (Type of genus: Synothmia bahamasella Hampson.
New synonymy.)
Clarke and Tams have compared the types of Hamp-
son’s two supposed genera and inform me that they
205
appear to be no more than sexes of one species. Hamp-
son’s chief character for separating Oedothmia and
Synothmia was the shape of the frons, conical on
Oedothmia and round on Synothmia; but Clarke states
that his description of the former is at variance with
the type in several details—the male antenna is simple,
the shaft not excised at base, and the frons isround, not
pointed. The venation of the types of the two genera
is similar.
Tongue well developed. Antenna pubescent; shaft
simple. Labial palpus upturned; the second segment
reaching above vertex; third segment “thickly scaled.”
Maxillary palpus filiform. Forewing smooth; 10 veins;
vein 2 from very near to lower outer angle of cell; 3
from the angle, approximate to 2 for some distance from
cell; 4 and 5 united, shortly separated from 3 at base;
6 from below upper angle of cell, straight; 8 and 9
stalked for about half their lengths; 10 from the cell,
separated from 8-9 at base; male without costal fold.
Hind wing with vein 2 from lower outer angle of cell;
3 from the angle, contiguous (but not fused) to 5 for
half their lengths; 7 and 8 contiguous beyond cell for
some distance, but not anastomosed; cell short, less
than one-third the length of wing; discocellular vein
curved.
Male genitalia unknown; the type of O. endopyrella,
and only known male, is without abdomen.
Female genitalia resembling those of Hurythmidia
and Oncolabis; with the signa strongly developed, the
large spined plate situated in bursa like that of Oncolabis
but the collar of strong spines at junction of bursa and
ductus is lacking; genital opening simple; ductus semi-
nalis from bursa near its junction with ductus bursae.
The genus is easily distinguishable from its nearest
relatives in venational division D (except Stylobasis) by
the approximate condition of veins 2 and 3 of forewing.
From Stylobasis it is distinguished by the contiguous
rather than anastomosed condition of veins 7 and 8 of
hind wing, its upturned labial palpi and simple male
antennae.
409. Oedothmia endopyrella Hampson
Figures 102, 944
Oedothmia endopyrella Hampson, Ann. Mag. Nat. Hist., ser. 10,
vol. 5, p. 61, 1930.
Synothmia bahamasella Hampson, Ann. Mag. Nat. Hist., ser. 10,
vol. 5, p. 61, 1930 (new synonymy).
Markings and color similar to those of Oncolabis
anticella Zeller, from which it can be distinguished most
readily by its venation. Alar expanse, 16-18 mm.
Typr Locauitizs: Vera Cruz, México (endopyrella,
in BM); Nassau, Bahamas (bahamasella, in BM).
Known only from the two types.
111. Genus Stylobasis Hampson
Stylobasis Hampson, in Ragonot, Monograph, pt. 2, pp. xii, 198,
1901. (Type of genus: Stylobasis rubripurpurea Hampson.)
Tongue well developed. Antenna pubescent; basal
segment enlarged; on male shaft with a hook from its
basal segment and deeply curved for several segments
206
beyond (as in Oncolabis), otherwise flattened, on female
simple. Labial palpus oblique; long, second segment
extending above vertex, on male grooved to hold the
maxillary palpus; third segment porrect (deflected for-
ward). Maxillary palpus filiform. Forewing smooth;
10 veins; vein 2 from near lower outer angle of cell; 3
from the angle, approximate to 2 in the male, (closely
approximate or connate with 5 in the female); 4 absent;
6 from slightly below upper angle of cell, slightly curved
in male (straight in the female); 8 and 9 stalked; 10
from the cell, approximate to 8-9 at base; male without
costal fold. Hind wing with vein 2 from lower outer
angle of cell connate with the stalk of 3-5; 3 and 5
stalked for half or (in female) slightly more than half
their lengths; 7 and 8 anastomosed for about half their
lengths (more shortly anastomosed in female) beyond
cell; cell slightly under one-third the wing length; dis-
cocellular vem curved. The venation except for the
absence of vein 4 of forewing is strikingly similar to that
of Oncolabis, especially in the female. Eighth abdom-
inal segment of male with compound ventral tuft.
Male genitalia similar to those of Oncolabis and
Hurythmasis except for slightly longer vinculum (a
difference of, at most, specific significance).
Female genitalia similar to those of Oncolabis; signa
strongly developed, consisting of a large, round, curved,
densely spined plate and a few similar detached spines
near the plate; a fused collar of similar spines surround-
ing the ductus bursae at its junction with bursa copu-
latrix; ductus bursae as long as bursa, minutely scob-
inate for about half its length beyond the collar; genital
opening simple; ductus seminalis from bursa near its
junction with ductus bursae.
410. Stylobasis rubripurpurea Hampson
Ficurss 106, 458, 946
Stylobasis rubripurpurea Hampson, in Ragonot, Monograph, pt.
2, p. 198, 1901.
The moth has the general habitus of Oncolabis
anticella; forewing purplish gray (or dark grayish
fuscous) with whitish costal streak sparsely dusted with
reddish scales; transverse lines obsolete, the antemedial
indicated below cell only by an outwardly bordering
pale yellowish patch. Hampson’s description men-
tions faint blackish streaks from the base of costa and
on the median vein and one distinct discal spot. The
figure in Ragonot (pl. 49, fig. 19) is misleading. It
shows a form with the veins darkly outlined, a distinct
whitish subterminal line, and no trace of the yellowish
patch on inner margin, or of the pale costal streak. Our
female in the National Collection is rubbed so that the
dark ground color shows plainly only along the veins,
thus corresponding with Ragonot’s figure. Alar ex-
panse, 16-19 mm.
Genitalia as given for the genus.
TYPE Locauity: “Jrazu, Mexique” (type in Trans-
vaal Mus.).
Foop eiant: Unknown.
In addition to the male type Hampson mentions a
UNITED STATES NATIONAL MUSEUM BULLETIN 207
male from Santos, Brazil. These, in addition to the
female from Juan Vinas, Costa Rica (Apr., in USNM),
are the only specimens of the species known to me.
Through the courtesy of Dr. Janse I was able to see
and figure the genitalia of the male type. He also
submitted a drawing of the wing venation which is re-
produced here. The femele in the National Museum
tentatively identified by Schaus as rubripurpurea differs
in slight details of venation from the male type, as indi-
cated in the foregoimg generic description, and may or
may not be conspecific. However, from its labial palpi
and the sum of its characters I believe it is properly
placed.
112. Genus Diviana Ragonot
Diviana Ragonot, Nouv. Gen., p. 27, 1888; Monograph, pt. 2,
pp. xii, 201, 1901.—Hulst, Phycitidae of N. Amer., p. 190,
1890. (Type of genus: Diviana eudoreella Ragonot.)
Dannemora Hulst, Phycitidae of N. Amer., p. 212, 1890.—Hamp-
son, 7m Ragonot, Monograph, pt. 2, pp. xiii, 209, 1901.
ean of genus: Dannemora edentella Hulst. New synony-
my.
Tongue short and weak, but exposed. Antenna
pubescent; on male, shaft curved toward base, some
rough scaling in the curve (but no teeth or other spine-
like projections). Labial palpus upturned, short, barely
reaching to vertex; the second segment tufted in front;
the third short, acuminate. Maxillary palpus squa-
mous (not filiform as stated by Ragonot and Hampson).
Forewing smooth; subtriangular, rather broad towards
termen; 10 veins; vein 2 from before the angle of the
cell; 3 and 5 closely approximate or connate from the
angle; 4 absent; 6 from below upper angle of cell,
straight; 8 and 9 stalked; 10 normally from the cell and
approximate to the stalk of 8-9, rarely connate or very
shortly stalked with 8-9; male without costal fold.
Hind wing with vein 2 from before but near lower outer
angle of cell; 3 and 5 shortly stalked, from the angle;
7 and 8 anastomosed for a short distance beyond cell
(for half or less than half their lengths); cell less than
half the length of wing; discocellular vein curved.
Eighth abdominal segment of male simple.
Male genitalia with apical process of gnathos a small
knob terminating in a short spine. Uncus as broad as
long, lateral margins parallel, terminal margin bluntly
angled. Transtilla absent (elements not distinguish-
able). Harpe short, slender, very slightly tapering to
narrowly rounded apex; costa’ broadly sclerotized
throughout and terminating in a short, slender spine.
Anellus a broad, deeply and widely cleft plate. Ae-
deagus long, moderately stout, strongly sclerotized,
with a few scobinations toward apex; penis armed with
two stout, straight cornuti (slightly less than one-third
as long as aedeagus). Vinculum very stout, longer than
combined tegumen and uncus and considerably longer
than broad; very slightly tapering; terminal margin
broad, straight.
Both Diviana and Dannemora were described from
males. As far as I know there are no known females of
either type species. In the Ragonot Monograph the
two supposed genera are widely separated in the generic
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 207
key on the alternative of vein 10 of forewing from the
cell or from the stalk of 8-9, an unreliable character at
best and in this instance thoroughly misleading. In
two males before me (perfect matches for the figure of
eudoreella in the Monograph, pl. 23, fig. 12) vein 10 is
both ways, from the cell on three forewings and dis-
tinctly stalked on one (compare figs. 103 and 1032).
On the Hulst type of edentella it is stalked.
Hampson’s characterization of Dannemora in the
Monograph is false in a number of details. He included
in the genus (in addition to its type species) Huzophera
quadripuncta Zeller, of which he had only females, and
selected ‘‘characters’”’ from both species and added a
“character” possessed by neither, namely, a circular
hyaline depression in the cell on the underside of hind
wing. There is no such structure in edentella or quadri-
puncta. The male of the latter has a depression in the
cell of hind wing, but it is not hyaline, is on the upper
surface of the wing, and is filled with dense, modified
sex-scaling. Hampson did not know the male, and the
hind wing of the female is simple, so it is difficult to see
where he got this ‘“‘character.”’ Zeller’s species is not
even closely related to edentella. I am treating it else-
where (p. 276). Ragonot’s characterization of Diviana
is misleading in one particular. He states that there are
some teeth (‘‘dents’’) in the sinus of the shaft of the
male antenna. Such a character is present in Palatka
nymphaeella (Hulst); but I do not think he could have
had an example of this species before him unless his
description of ewdoreella is incorrect and his figure of it
completely false.
411. Diviana eudoreella Ragonot
Figures 103, 463
Diviana eudoreella Ragonot, Nouv. Gen., p. 27, 1888; Monograph,
pt. 2, p. 201, 1901.—Hulst, Phycitidae of N. Amer., p. 190,
1900.—McDunnough, Check list, No. 6358, 1939.
Dannemora edentella Hulst, Phycitidae of N. Amer., p. 212,
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 209,
1901.—McDunnough, Check list, No. 6362, 1939. (New
synonymy.)
Diviana eudoriella Hulst, U. 8. Nat. Mus. Bull. 52, p. 4383, 1902
(misspelling).
Forewing blackish brown, the median area heavily
dusted with white, giving it an ashy gray appearance;
antemedial white line nearly straight, slanting out-
wardly from costa to inner margin, slightly notched at
lower fold, bordered outwardly by a blackish line which
expands to a triangular blotch at costa, a similar, some-
what smaller blotch bordering the line at inner margin;
subterminal white line parallel to termen, notched at
vein 6 and at lower fold, towards costa bordered in-
wardly by a fuscous streak; discal dots at end of cell
distinct, blackish; entire outer area between subterminal
line and termen blackish brown. Hind wing pale smoky
fuscous, with a darker line along termen and some very
faint dark shading on the veins. Alar expanse, 16-19
mm.
Male genitalia as given for the genus.
Type Locauities: ‘‘America Septentrionalis’ (eu-
doreella, in Paris Mus.); Florida (Apr.; edentella, in
AMNH, ex Rutgers).
Foop pLant: Unknown.
Known only from males. The Hulst type is badly
tubbed and shows little trace of original markings; but
its genitalia agree in every detail with those of two well-
marked specimens (in USNM) from Orlando, Fla.
(Feb.), and Plummers Isl., Md. (July). These two
specimens are in perfect agreement with Ragonot’s
figure of eudoreella.
113. Genus Palatka Hulst
Palatka Hulst, Canadian Ent., vol. 24, p. 62, 1892; U. S. Nat.
Mus. Bull. 52, p. 433, 1902. (Type of genus: Divinia nym-
phaeella Hulst.)
Tongue reduced (better developed in female than
male). Antenna pubescent; shaft of male with a
shallow sinus at base containing a weak scale tuft and
a couple of short teeth concealed within the tuft (these
teeth are strongly sclerotized, pointed, dorsal projections
from two of the segments); shaft of female simple.
Labial palpus oblique, extending well above vertex;
broadly scaled; segment 2 long; segment 3 about one-
third the length of 2, acuminate. Maxillary palpus
filiform. Forewing smooth; 10 veins; vein 2 from before
the angle of the cell; 3 from the angle; 4 absent; 5 from
slightly above the lower angle and separate from 3; 6
from below upper angle, straight; 8 and 9 stalked; 10
from the cell separate at base from stalk of 8-9; male
without costal fold. Hind wing with vein 2 from well
before lower outer angle of cell; 3 and 5 from the angle,
very closely approximate towards base; 7 and 8 closely
approximate for half their distance beyond cell; cell
about one-half the length of wing; discocellular vein
curved. Eighth abdominal segment of male simple.
Male genitalia with apical process of gnathos an
elongate, narrow hook. Uncus longer than broad, with
terminal margin broadly rounded. ‘Transtilla incom-
plete, elements minute. Harpe elongate, narrow, grad-
ually tapering to bluntly pointed apex; sacculus pro-
duced at extremity into blunt, free arm directed towards
costa. Anellus a broad flattened plate. Aedeagus
long, moderately stout, smooth; penis armed with two
stout cornuti, about half as long as aedeagus. Vincu-
lum stout, slightly longer than broad, tapering to
broadly rounded terminal margin.
Female genitalia with bursa copulatrix scobinate but
without signa, thickened (but not sclerotized) at ter-
minal end; ductus bursae shorter than bursa, strongly
sclerotized from junction with bursa for a little more
than half its length, on ventral surface at genital open-
ing a broad, strongly sclerotized, triangulate plate;
ductus seminalis from bursa near its junction with
ductus bursae.
The genus is quite distinct from others of venational
group D and easily identified by its male and female
genitalia. It is closest to Diviana, from which it is
readily separable by its hind wing venation, veins 3 and
208 UNITED STATES NATIONAL MUSEUM BULLETIN 207
5 approximate rather than stalked. It contains only
one North American species.
412. Palatka nymphaeella (Hulst)
Ficures 104, 462, 954
Diiana nymphaeella Hulst, Canadian Ent., vol. 24, p. 62, 1892.
Palatka nymphaeella (Hulst), Canadian Ent., vol. 24, p. 62, 1892;
U. S, Nat. Mus. Bull. 52, p. 433, 1902.—Barnes and Mc-
Dunnough, Contributions, vol. 3, p. 222, 1917.—McDun-
nough, Check list, No. 6360, 1939.
Diviana verecuntella Grossbeck, Bull. Amer. Mus. Nat. Hist.,
vol. 37, art. 1, p. 132, 1917.
Forewing grayish ocherous with whitish transverse
line and more or less marked with brownish fuscous; in
paler specimens the ocherous tint predominates, in
darker one, the gray; antemedial line outwardly trans-
verse from costa, indented at median and lower folds,
in well-marked (pale) specimens bordered inwardly on
lower half by a large dark patch; subterminal line
rather near and parallel to termen, very slightly in-
dented at vein 6 and lower fold and with some dark
streaks bordering it near costa; discal spots distinct,
blackish; a row of blackish dots along termen. Hind
wings pale smoky fuscous. Alar expanse, 16-18 mm.
Genitalia as given for the genus.
Typr Locatities: Charlotte Harbor, Fla. (nymphae-
ella, in AMNH, ex Rutgers); Fort Myers, Fla. (vere-
cuntella, i AMNH).
Foop piuant: Unknown.
Distrisution: Florida, Charlotte Harbor (Apr.),
Everglades (Apr.), Fort Myers (Apr.); Texas, San Be-
nito (Apr.); Connecticut, East River (July).
The Hulst type in the Rutgers Collection is only a
fragment. One pair of wings, the antennae, and the
abdomen are missing; but there is no doubt of what it
represents. The alar expanse given by Hulst (21 mm.)
is too great. It is nearer 18 mm. The Connecticut
specimens (one male and six females in the National
Collection) are darker than those from Florida and
Texas, more grayish, and with little or no dark shading
or blotches bordering the transverse lines. They show
no genitalic differences.
114. Genus Cacozophera Dyar
Cacozophera Dyar, Ins. Insc. Menstr., vol. 7, p. 58, 1919.
(Type of genus: Cacozophera venosa Dyar.)
Tongue reduced but exposed. Antenna of female
pubescent, simple. Labial palpus upturned, short,
not reaching vertex; third segment acuminate, shorter
than second. Maxillary palpus filiform. Forewing
smooth; narrowly elongate; 10 veins; vein 2 from well
before outer angle of cell; 3 and 5 connate from the angle;
4 absent; 6 from below upper angle of cell, straight; 8
and 9 stalked; 10 from the cell, approximate to the
stalk of 8-9 for some distance. Hind wing with vein
2 from well before lower outer angle of cell; 3 and 5 con-
nate from the angle;7 and 8 anastomosed for most of their
lengths (8 very short); cell less than one-half the
length of the wing; discocellular vein curved.
Female genitalia with bursa copulatrix large, finely
scobinate throughout, the scobinations extending part
way into the ductus bursae; signa present, consisting
of a cluster of small, slender disks (three in the only
specimen available); ductus bursaze slightly shorter
than bursa, unsclerotized; genital opening simple;
ductus siminalis from bursa in the neighborhood of the
signa.
The genus and its type species were erected on a
single female. Dyar characterized Cacozophera merely
as having the venation of his genus Anthropteryz,
differing only in having the “wings long and narrow,
trigonate, the apex pointed, not short and square.”
Anthropteryx itself was also erected on a single female
and unfortunately is a freak with vein 4 absent from
one forewing. Dyar did not notice the other forewing
in which vein 4 was present and stalked with 5, the
normal condition for the specimen. For further dis-
cussion of Anthropteryx see page 313. The placement of
Cacozophera is tentative. Its relationship to the other
genera cannot be determined until a male is discovered.
413. Cacozophera venosa Dyar
Fiaures 105, 950
Cacozophera venosa Dyar, Ins. Insc. Menstr., vol. 7, p. 58, 1919.
Forewing brownish fuscous, the area between costa
and cell and vein 6 uniformly dark; below vein 6 and
from end of cell the veins darkly outlined and the inter-
spaces between them contrastingly paler; subterminal
line distinct from vein 6 to inner margin, whitish,
evenly curved and parallel with termen; no dis-
tinguishable antemedial line or discal spots; a faint
yellowish white shade along inner margin and (under
magnification) a scattered dusting of dull rosy scales on
the dark areas; terminal dots blackish, large, inter-
venular. Hind wing smoky fuscous, the veims and
terminal margin darker. Alar expanse, 19 mm.
Genitalia as given for the genus.
TyprE Locatity: Cayuga, Guatemala (May, type in
USNM).
Foop piant: Unknown.
Known only from the female type.
species but of uncertain affinities.
Obviously a good
Genera 115-117: Psorosina to Paconius
[Venational division A, Forewing with 11 veins; 10 from the
cell or connate or shortly stalked with stalk of 8-9; 4 and 5
approximate, connate or very shortly stalked. Hind wing with
vein 2 from before lower outer angle of cell; discocellular vein
curved or incomplete. Male genitalia with costa of harpe
produced; cornuti developed; transtilla incomplete or absent.]
115. Genus Psorosina Dyar
Psorosina Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 113,
1904.—Forbes, Cornell Mem. 68, p. 632, 1923. (Type of
genus: Psorosina angulella Dyar.)
Tongue well developed. Antenna pubescent; shaft
of male with sinus and large scale tuft at base; shaft of
female simple. Labial palpus upcurved, slender, rough
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 209
scaled, reaching to vertex in male, to slightly above in
female; third segment about half the length of second,
acuminate. Maxillary palpus squamous. Forewing
smooth; 11 veins; vein 2 from before lower outer angle
of cell; 3 from the angle; 4 and 5 connate (or very
shortly stalked), separated from 3 at base; 6 from
below upper angle of cell, straight; 8 and 9 stalked;
10 from cell, separated from stalk of 8-9; male without
costal fold. Hind wing with vein 2 from before (but
near) lower outer angle of cell; 3 and 5 stalked; 7 and
8 anastomosed for half their lengths beyond cell; cell
about half the length of wing, partially open; dis-
cocellular vein incomplete. Eighth abdominal segment
of male with two pairs of ventrolateral hair tufts.
Male genitalia with apical process of gnathos a stout
hook. Uncus triangulate. Transtilla incomplete, ele-
ments minute. Harpe elongate, narrow, not appre-
ciably tapering; apex rounded; strongly sclerotized
costa projecting at apex into a short, sharp spine;
clasper erect, short, weakly sclerotized. Anellus a
curved U-shaped plate with broad base. Aedeagus
short, stout, nearly straight; penis armed with a single,
long, moderately slender cornutus (nearly as long as
aedeagus). Vinculum stout, about as broad as long,
slightly tapering to truncate terminal margin.
Female genitalia with bursa copulatrix scobinate
over most of its inner surface; signum developed as an
irregularly shaped spined plate near junction of ductus
bursae; ductus bursae shorter than bursa, with a pair
of narrow, sclerotized, strongly spined plates along the
lateral margins, some minute scobinations in the inter-
vening area, a fingerlike, ventral, sclerotized plate from
genital opening, and a few granulations on the inner
wall above genital opening; ductus seminalis from
bursa near its junction with ductus bursae.
A distinct genus with one North American species;
easily distinguished by venation and genitalia.
414. Psorosina hammondi (Riley)
Fieurts 62, 461, 952
Pempelia hammondi Riley, Fourth annual report on the noxious,
beneficial and other insects, of the State of Missouri, p. 44,
1872.
Psorosa hammondi (Riley) Ragonot, Ent. Amer., vol. 5, p. 116,
1889.
Canarsia hammondi (Riley) Hulst, Phycitidae of N. Amer., p.
180, 1890.—Hampson, in Ragonot, Monograph, pt. 2, p.
120, 1901.
Psorosina hammondi (Riley) Dyar, Proc. Ent. Soc. Washington,
vol. 6, p. 113, 1°04.—Forbes, Cornell Mem. 68, p. 682,
1923.—McDunnough, Check list, No. 6335, 1939.
Psorosina angulella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 113,
1904.—Forbes, Cornell Mem. 68, p. 632, 1923.—McDun-
nough, Check list, No. 6336, 1939. (New synonymy.)
Forewing glossy brownish fuscous (purplish brown
in some lights) ; antemedial line grayish white, straight,
slightly curved or weakly angled at cell, slanting a trifle
from costa to inner margin, of varying width depending
on the amount of white scaling and sometimes (but
rarely) partially divided by a fine median dark line;
subterminal line obscure or indistinguishable, fine,
whitish, nearly vertical from costa to tornus, very
faintly bordered by blackish lines; on costa adjacent to
inner border of the subterminal line a more or less
extended and triangulate grayish white patch; blackish
discal and terminal dots obscure, the discal dots more
or less confluent. Hind wings smoky fuscous, glossy.
Alar expanse, 13-14 mm.
Genitalia as given for the genus; terminal margin of
vinculum broadly and shallowly notched.
Type tocauitins: Illinois (hammondi, in USNM);
Iowa (angulella, in USNM).
Foop pxiants: Apple, pear, Prunus maritima.
(Specimens in USNM also labeled ‘from sycamore”
and ‘‘from acorns,”’ but these are very dubious records.)
The larva is a leaf skeletonizer.
DistrrBuTion: Connecticut, East River (Aug.) ; Rhode
Island, Weekapaug (Aug., Sept.); Ohio, Cincinnati
(Aug.) ; Indiana, Bedford (July) ; Jilinois, Oconee (July,
Aug.); Iowa; Kansas, Wathena (July), Wichita (June) ;
Missouri, several specimens with no further locality,
St. Louis (Aug.). Rather generally distributed
throughout the eastern and central United States and
Canada.
Worn females of this species are superficially similar
to small specimens of Moodna ostrinella (Clemens) and
the two species have been occasionally confused. They
are quite distinct, however, on both venation and
genitalia. The character on which Dyar separated his
angulella (its “strongly angled’ antemedial line) is
rather imaginary than real. The pale markings on the
forewing of hammondi are formed by rather thinly
spread, whitish scales and vary from specimen to
specimen in size, shape, and intensity. Dyar’s type is
rubbed, a condition which accounts in part for the
angulate condition of the antemedial line, a very slight
angulation at that. Its genitalia agree in every detail
with those of typical hammondt.
116. Patriciola, new genus
Typr or Genus: Patriciola semicana, new species.
Tongue well developed. Antennae simple in both
sexes, shortly pubescent. Labial palpus porrect (beak-
like), long, projecting about four times the length of
head beyond it; alike in both sexes. Maxillary palpus
squamous, large; alike in both sexes. Forewing smooth;
11 veins; vein 2 from well before the lower outer angle
of the cell; vein 3 remote from 2 but before the angle of
the cell; 4 from the angle approximate to 5 for some
distance from cell; 6 from below upper angle of cell,
straight; 8 and 9 stalked; 10 connate or very shortly
stalked with 8-9; male without costal fold. Hind wing
with vein 2 from well before the lower outer angle of the
cell; 3 from near but before the angle; 5 from the angle;
7 and 8 contiguous (touching but not completely fused)
for nearly half their lengths beyond cell; cell about half
the length of the wing; discocellular vein curved.
Eighth abdominal segment of male simple.
Male genitalia with apical process of gnathos a rather
broadly based, blunt hook produced anteriorly into
a long, flat, sclerotized apron (the whole process resem-
210
bling, in outline, a bulbus-necked bottle). Uncus
broader than long; sides nearly parallel ; terminal margin
broadly rounded. Transtilla incomplete; elements mi-
nute, folded. Harpe short; appressed clasper well out
towards outer margin of harpe; from base of costa a
long, stout, curved, strongly sclerotized and pointed
clasperlike horn projects across face of harpe. Anellus
a flattened plate. Aedeagus short, stout, slightly sinu-
ate, and with apex sharply constricted; armed with a few
short spines near apex; penis with a small patch of fine
scobinations, otherwise unarmed. Vinculum stout, as
broad as long, tapering to truncate terminal margin.
Female genitalia with bursa copulatrix finely scob-
inate throughout, otherwise simple, without signum;
ductus bursae sclerotized, broad and very short, broad-
ening abruptly into the sclerotized and thickened lower
lip to an exceptionally wide genital opening; ductus
seminalis from bursa near its junction with ductus
bursae.
A striking genus easily identified by venation and
genitalia. Contains one North American species.
415. Patriciola semicana, new species
Fieures 84, 460, 953
Forewing ashy grayish white on costal half; shaded
with faun brown rather heavily dusted with blackish
fuscous on lower half, especially from base to antemedial
line and slightly beyond; veins partially outlined by
dark scaling; antemedial line distinct only from cell to
inner margin, far out towards middle of wing, white,
narrow, evenly incurved; subterminal line very faint,
nearly obsolete, parallel to and rather far in from
termen; lower discal dot at end of cell, large, distinct
and blackish, a similar smaller black spot in cell over
the antemedial line. Hind wing semihyaline, smoky
white; veins not darkened and terminal margin only
faintly so; cilia concolorous with wing. Alar expanse,
25-26 mm.
Genitalia as given for the genus.
TYPE Locality: Provo, Utah (type in USNM,
61367).
Foop prant: Unknown.
Described from male type and two female paratypes
from the type locality, collected by Tom Spalding July
16, 1909 (co), and Aug. 4, 1908 (¢@). In habitus sem-
icana resembles most some specimens of Honora but is
easily separable from anything in that genus.
117. Paconius, new genus
TypPE oF GENus: Paconius corniculatus, new species.
Tongue well developed. Antenna pubescent; shaft
of male with sinus and small scale tuft at base. Labial
palpus upcurved, rough scaled, reaching to vertex; third
segment about half the length of second, acuminate.
Maxillary palpus squamous. Forewing smooth; 11
veins; vein 2 from before the lower outer angle of cell;
3 from very close to angle; 4 and 5 approximate for some
UNITED STATES NATIONAL MUSEUM BULLETIN 207
distance from cell; 6 from below upper angle of cell,
straight; 8 and 9 stalked; 10 from the cell, shortly
separated from the stalk of 8-9; male without costal
fold. Hind wing with vein 2 from before lower outer
angle of cell; veins 3 and 5 contiguous for a short dis-
tance from angle (touching but not fused at any point);
7 and 8 contiguous for less than half their lengths beyond
cell; cell about half the length of wing; discocellular
vein curved. Highth abdominal segment of male with
a weak pair of ventrolateral hair tufts.
Male genitalia with apical process of gnathos nar-
rowly triangulate and acutely pointed. Uncus irregu-
larly triangulate, tapering to narrowly rounded apex.
Transtilla absent; elements not distinguishable. Harpe
divided to base with entire costa developed as a strongly
sclerotized, long, hornlike projection; remainder of
harpe much reduced, triangulate and tapering to a
point; clasper absent. Anellus a narrow, slightly
curved plate with very long, slender, lateral arms.
Aedeagus long, slender, abruptly bent toward apex;
armed at apex with two thornlike spurs; penis armed
with a cluster of short, rather stout spines. Vinculum
stout, slightly longer than broad, very slightly tapering
to broad, evenly rounded, terminal margin.
Female unknown.
Paconius has much the same venation, palpi, and
antennae as the European genus Psorosa; but differs
markedly in genitalia. In Psorosa the harpe is simple
and the gnathos, vinculum, and anellus entirely different.
Paconius has several features in common with Patriciola,
with which it appears to be most closely related, dif-
fering most from that genus in its upcurved rather than
porrect palpi. I dislike very much describing a new
genus without having females for completion of the
diagnosis on genitalic characters; but it is so obviously
new and the male genitalia so different from anything
else in our fauna that it seems advisable to give it some
designation.
416. Paconius corniculatus, new species
FIGURE 464
Forewing pale ashy gray; transverse lines nearly ob-
solete; antemedial line faintly outlined, nearly straight,
slanting outwardly from costa to inner margin, indicated
chiefly by a small blackish spot on its inner border at
inner margin and a couple of short blackish streaks on
its outer border near costa; costal edge at base of wing
blackish; subterminal line not defined except by a few
blackish, inwardly bordering streaks near costa. Hind
wing semihyaline white with a narrow fuscous shade
along costa and outer margin. Alar expanse, 19 mm.
Male genitalia as given for the genus.
Typr Locatity: San Germdn, Puerto Rico (type in
Cornell Univ.; paratype in USNM, 61368).
Foop prant: Unknown.
Described from male type and one male paratype
from the type locality collected Apr. 16 and 17, 1930,
under Cornell lot 795, sub. 34 and 36.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 211
Genera 118-120: Aptunga to Cassiana
[Venational division A. Forewing with 11 veins; 10 from cell,
8 and 9 stalked, 6 straight, 4 and 5 stalked, 2 and 3 separate or
approximate. Hind wing with 7 veins; 7 and 8 strongly anasto-
mosed, 3 and 5 approximate at base or stalked, discocellular vein
curved. Male genitalia with uncus subtriangular, tapering
abruptly, apical half narrow, bluntly pointed; gnathos ter-
minating in a small, bifid, hooked process; transtilla incomplete,
represented by a pair of separate, elongate plates; harpe simple,
elongate, apex obliquely rounded; vinculum stout, decidedly
longer than broad; aedeagus moderately stout, straight; penis
without cornutus. Female genitalia with ductus seminalis from
bursa.]
118. Aptunga, new genus
Typs or Genus: Vitula macropasa Dyar.
Tongue well developed. Antenna pubescent, shaft
simple in both sexes. Labial palpus obliquely up-
turned. Maxillary palpus filiform. Forewing nar-
rowly elongate, smooth; veins 2 and 3 closely approxi-
mate from lower outer angle of cell; stem of 4-5 approxi-
mate to 3; vein 10 from the cell, rather well separated
from stalk of 8-9; male without costal frold or other
sexual modifications. Hind wing with vein 2 from well
before angle of cell; 3 and 5 closely approximate at
lower, outer angle of cell; 7-8 anastomosed for most of
their lengths beyond cell (free end of vein 8 very short
and weak); cell less than half the length of wing.
Abdomen of male with a strong pair of ventrolateral
hair tufts from eighth segment.
Male genitalia with apical process of gnathos small,
bifid; aedeagus simple (without longitudinal ventral
sclerotized ridge).
Female genitalia without signum; bursa copulatrix
more or less finely scobinate in the area about attach-
ment of ductus seminalis, otherwise smooth; ductus
bursae smooth except for a weak sclerotization near
genital opening and some fine scobinations at junction
with bursa; ductus seminalis from middle or near
middle of bursa.
This genus and the two genera following are closely
related to each other and Mescinia, agreeing in nearly
all male genitalic characters but differing in venation.
Aptunga is similar in hind wing venation to Mescinia,
differing from the latter in having veins 2 and 3 of
forewing closely approximate at base rather than
stalked, and lacking the signum in bursa of the female.
417. Aptunga macropasa (Dyar), new combination
Ficures 85, 465, 958
Vitula macropasa Dyar, Ins. Inse. Menstr., vol. 7, p. 61, 1919.
Forewing grayish fuscous; antemedial line obscure,
whitish, bordered outwardly by a diffused dark shade;
subterminal line narrow, whitish, decidedly slanting
and slightly angled near middle, shaded inwardly and
outwardly by dark streaks on the veins; a row of small
blackish dots along termen; discal dots at end of cell
separate, somewhat elongate; under magnification costa
shows a peppering of reddish scales. Hind wing
whitish, semihyaline, the veins outlined with fuscous
and a fuscous shade bordering costa and along terminal
margin. Alar expanse, 19-23 mm.
Male genitalia with weak sclerotized granulations
and wrinklings on penis. Female genitalia with a
narrow band of sclerotized granulations on inner
dorsal surface of ductus bursae at genital opening.
TypE tocauity: Purulhé, Guatemala (type in
USNM).
Foop prant: Unknown.
Distripution: GuatemMAua: Chejel (Aug.), Purulha
(July), Voleén Santa Maria (Nov.). M*x1co: Orizaba.
418. Aptunga imperfecta (Dyar), new combination
FicurE 957
Mescinia inperfecta Dyar, Ins. Insc. Menstr., vol. 7, p. 60, 1919.
Considerably shorter and somewhat paler than
macropasa, the general color (in the rubbed type) more
brownish than grayish fuscous; no trace of reddish
scaling in costal area of forewing; discal dots fused into
a single round dark spot. Hind wing darker, less
hyaline than that of macropasa. Alar expanse, 14 mm.
Female genitalia with the ductus bursae appreciably
but not strongly sclerotized and granulate for a short
distance from genital opening.
Typr Locatity: Cayuga, Guatemala (Apr., type in
USNM).
Foop piant: Unknown.
Known only from the female type; described by
Dyar in Mescinia but ruled out of that genus by its
forewing venation and. lack of signum.
119. Anderida, new genus
Typr or Genus: Euzophera sonorella Ragonot.
Tongue well developed. Antenna pubescent, shaft
simple in both sexes. Labial palpus obliquely up-
turned. Maxillary palpus filiform. Forewing narrowly
elongate, smooth; vein 2 from near angle of cell but
well separated from 3; 10 from cell, separate from
stalk of 8-9; male without costal fold or other sexual
modifications. Hind wing with 3 and 5 stalked; 7-8
anastomosed for at least two-thirds the length of vein
7; cell approximately half the length of wing. Abdomen
of male with strong pair of ventrolateral hair tufts
from eighth segment.
Male genitalia as in Aptunga except apical process of
gnathos partially fused.
Female genitalia with signum; ductus seminalis from
anterior end of bursa; ductus bursae with a thin
sclerotized shield on venter at genital opening and more
or less finely scobinate towards bursa, otherwise smooth.
419. Anderida sonorella (Ragonot), new combination
Ficurss 467, 959
Euzophera senorella Ragonot, N. Amer. Phycitidae, p. 14, 1887;
Monograph, pt. 2, p. 59, 1901.
Euzophera sonorella Ragonot, Nouv. Gen., p. 4, 1888 (correction
of spelling).
Eyzophera placidella Dyar, Proc. Ent. Soc. Washington, vol. 10,
212 UNITED STATES NATIONAL MUSEUM BULLETIN 207
p. 115, 1908.—MeDunnough, Check list, No. 6314, 1939.
(New synonymy.)
Forewing ashy gray on costal half shading to white
on costa; lower half of wing shaded with ocherous gray;
rather narrow antemedian and subterminal bands
strongly contrasted, blackish fuscous, antemedian band
slightly curved, nearly vertical, ending just before costa;
discal dots small, blackish, the upper one obscure and
frequently absent. Hind wing whitish with a faint
gray or ocherous tint. Alar expanse, 16-22 mm.
Female genitalia with signum very weak, consisting
of a cluster of 3 to 5 small disks; bursa copulatrix finely
scobinate.
TypE Locauitiss: ‘Senora”’ [sic], México (sonorella,
in Paris Mus.); Yuma County, Ariz. (placidella, in
USNM).
Foop piant: Unknown.
Disrrisution: Unirep Starrs: Arizona, Huachuca
Mts., Santa Catalina Mts., Redington, Yuma County
(Colorado Desert). México: Sonora.
In some of the female specimens before me from
Redington, Ariz., the transverse dark markings are
rather faint, almost obsolete in a few examples and
there is considerable variation in size, but the general
habitus and genitalia easily distinguish the species.
120. Cassiana, new genus
Type or Genus: Vitula malacella Dyar
Tongue well developed. Antenna pubescent; in male
with a row of 6 or 7 short slender spines along outer side
of shaft towards base, shaft somewhat swollen. Labial
palpus upturned; in female slightly oblique. Maxillary
palpus filiform. Forewing smooth; venation as in
Aptunga, veins 2-3 closely approximate from angle of
cell; male with a strong costal fold terminating at basal
third in an outwardly projecting brush of scales. Hind
wing with veins 3-5 stalked; 7-8 anastomosed for most
of their lengths (free end of vein 8 a short, weak spine);
cell about half the length of wing. Abdomen of male
with a strong pair of ventrolateral hair tufts from eighth
segment.
Male genitalia with apical process of gnathos small,
bifid; aedeagus with a sclerotized, longitudinal, ventral
ridge for over half its length from base.
Female genitalia with sisnum; ductus seminalis from
anterior end of bursa.
420. Cassiana malacella (Dyar), new combination
Ficures 466, 955, 956
ieee malacella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 408,
914.
Forewing pale grayish fuscous, the area between cell
and costa white with a faint, scattered dusting of red-
dish scales; costal edge from base to one-third (the extent
of the costal fold) blackish; transverse antemedial line
obsolete; subterminal line but faintly indicated. Hind
wing whitish, the veins outlined by dark scales, and a
narrow smoky border along termen and costa. Alar
expanse, 15-17 mm.
Male genitalia with lateral margins of vinculum con-
cave, terminal margin straight. Female genitalia with
signum variable, consisting of from one small platelike
projecting disk to a line of similar disks (examples of
the extreme types in two Puerto Rican specimens are
shown in figs. 955, 956); bursa and considerable part of
ductus bursae finely scobinate.
Typz Locatity: Tehuacén, México (type in USNM).
Foop pranr: Unknown.
Disrrisution: Mrxico: Tehuacén (May). PuErro
Rico: Bayamén (Sept.), Puerto Real (Apr.), Rio
Piedras (Sept.). Virrern Isuanps: Kingshill (St. Croix,
Mar., Dec.).
Genus 121: Mescinia
[Venational division B. Forewing with 11 veins: 10 from cell,
8 and 9 stalked, 4 and 5 stalked, 2and 3 stalked. Male genitalia
with transtilla incomplete.]
121. Genus Mescinia Ragonot
Mescinia Ragonot, Monograph, pt. 2, p. viii, 1901—Hampson,
in Ragonot, Monograph, pt. 2, p. 83, 1901. (Type of genus:
Ephestia commatella Zeller.)
Tongue well developed. Antenna pubescent; shaft
slightly thickened towards base in male, simple in
female. Labial palpus obliquely upturned, reaching to
vertex. Maxillary palpus filiform. Forewing smooth,
narrowly elongate; termen slanting; 11 veins; veins 2
and 3 from angle of cell, stalked; 4 and 5 stalked; 6
straight; 8 and 9 stalked; 10 from the cell, separate from
the stalk of 8-9; male with a short costal fold at base of
wing. Hind wing with vein 2 from well before lower
angle of cell; 3 and 5 from angle and closely approximate
at base; 6 from upper angle of cell; 7 and 8 strongly
anastomosed beyond cell (in small species completely
fused or showing the free part of vein 8 as a short spur
to costa); discocellular vein curved; cell one-third to
two-fifths of wing length.
Male genitalia with uncus subtriangular, tapering
abruptly from broad base, terminal half narrow and
apex bluntly pointed (except in indecora), outer surface
covered with bristlelike scales. Gnathos terminating
in a small, or moderately small, bifid apical process
(except in indecora). Harpe simple, with costa sclero-
tized for most of its length. Anellus U-shaped with
long lateralarms. Aedeagus straight, moderately stout,
a strong, longitudinal, sclerotized ridge on ventral side.
Penis more or less scobinate and frequently with sclero-
tized patch and wrinklings. Vinculum stout, consider-
ably longer than broad (except in indecora) and with
terminal margin truncate.
Female genitalia with bursa copulatrix membranous,
finely scobinate over part of inner surface (especially
about signum and near ductus bursae); signum nor-
mally an oval or round sclerotized plate covered with
short, blunt or bluntly pointed thorns, or a single stout
platelike thorn (indecora), or absent (discella). Ductus
bursae membranous except for a slight sclerotization
near genital opening, finely scobinate toward bursa.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 213
Ductus seminalis from anterior end of bursa or from
bursa near its juncture with ductus bursae.
The genus Mescinia is closely related to the foregoing
three genera (Aptunga, Anderida, Cassiana) and to
Nonia, all of which have similar genitalia, especially in
the male, but which are readily separable on venation.
Mescinia is an American genus without, as far as I know,
any properly included exotic species. Hampson (in
Ragonot, Monograph, pt. 2, pp. 84, 86) includes one
Indian species, micans Hampson, but this obviously is
wrongly referred for it lacks vein 10 of forewing and
has 4 and 5 from the cell and not stalked as they should
be for Mescinia.
Our species divide into three groups on size as follows:
Alar expanse less than 15 mm.
Alar expanse more than 15 but less than 20 mm.
Alar expanse more than 20 mm,
The first two groups are typical Mescinia on all struc-
tural characters, but the two large species in the third
group are somewhat abnormal, especially indecora,
which is aberrant on both male and female genitalia.
The other species (discella) has normal male genitalia,
but the female lacks a signum, and in both the anasto-
mosis of veins 7 and 8 of hind wing is considerably
shorter than in normal Mescinia.
Among the species of the first group the color and
pattern differences are trifling, and the distinctions
stressed by Dyar between his species are as much due
to the condition of the specimens as to differences in
color or intensity of dark scaling. Genitalic differences
are also slight, but apparently more reliable. The best
characters are found in the signa, the apical process of
gnathos, and the armature of the penis. With the few
specimens from widely scattered localities and the
scanty biological information available, it is almost im-
possible to define specific limits in this group with any
certainty.
Genus Mescinia, Species 421-428: M. triloses to
M. berosa
[Alar expanse less than 15 mm.]
421. Mescinia triloses Dyar
Figures 468, 965
Mescinia triloses Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 341,
914.
Mescinia mosces Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 341,
1914 (new synonymy).
Forewing sordid white, the costal area paler and con-
trasted; dark markings pale brownish fuscous; discal
dots more or less confluent; antemedian line obscure;
subterminal dark line rather well marked in unrubbed
specimens; in fresh examples a dark streak along outer
half of fold (replacing the absent vein 1c). Hind wing
whitish, the veins faintly outlined by dark scaling;
discocellular vein slightly curved. Veins 2 and 3 of
forewing are stalked for about one-third and veins 4-5
for approximately one-half their lengths. Alar ex-
panse, 10-14 mm.
Tyrer Locauity: Taboga Isl., Panama (Triloses and
Mosces, in USNM).
Foop prant: Unknown.
DistripuTion: PanamA: Alhajuelo (Apr.), Corazal
(Feb.), Taboga Isl. (Feb.).
The specimens of Dyar’s mosces are a trifle darker and
more strongly marked than those of triloses, but they
represent only the least rubbed of a series collected on
the same date at the same locality and exhibit no struc-
tural differences of any significance in genitalia. I am
unable to distinguish any trace of the ‘‘faintly pinkish”
color which Dyar mentions in his descriptions.
422. Mescinia pandessa Dyar
Figure 469
Mescinia pandessa Dyar, Ins. Inse. Menstr., vol. 7, p. 60, 1919.
In color and markings similar to the females of
mosces. Possibly nothing more than a variety or race
of triloses, but exhibiting a few slight structural differ-
ences: Narrower harpes, a somewhat stronger scobina-
tion on penis, and a wider spacing of the bifid apical
process of gnathos. Such differences are hardly signifi-
cant, but in the absence of females it is safer to retain
the names in specific rank for the present. Veins 2 and
3 of forewing stalked for two-thirds and 4 and 5 for
one-half or slightly more than one-half of their lengths.
Discocellular vein of hind wing as in triloses. Alar
expanse, 12-13 mm.
Type tocauity: Cayuga, Guatemala (Apr., type in
USNM).
Foop puant: Unknown.
Known only from the types series from Cayuga.
The specimens in the National Collection are all males.
423. Mescinia bacerella Dyar
Ficures 94, 470, 962
Mescinia bacerella Dyar, Ins. Inse. Menstr., vol. 7, p. 59, 1919.
General color somewhat darker than triloses or
pandessa; costal area of forewing more contrasted,
whitish with a scattering of reddish brown scales.
Genitalia showing only minute differences from those of
preceding species. Veins 2 and 8 of forewing stalked
for two-thirds and 4 and 5 stalked for one-half their
lengths. Discocellular vein of hind wing very slightly
curved. Alar expanse, 12 mm.
Typr LocaLity: Havana, Cuba (type in USNM).
Foop piant: Unknown.
DisrrisuTion: Cusa: Havana, Sierra Maestra (Jan.).
424, Mescinia estrella Barnes and McDunnough
Figures 473, 963
Mescinia estrella Barnes and MeDunnough, Contributions, vol. 2,
p. 182, 1913.—McDunnough, Check list, No. 6322, 1939.
Wing pattern similar to that of other species in this
group, but (in reared examples especially) dark mark-
ings more intense and whitish costal area of forewing
more conspicuously dusted with reddish scales. Male
genitalia similar to those of pandessa except bifid apical
214
process of gnathos (fig. 473) somewhat more widely divi-
ded (as in parvula). 'The spining of the female signum is
almost identical to that of triloses (compare figs. 468
and 963). Veins 2 and 3 and 4 and 5 of forewing
stalked from slightly less than one-half to about one-
half their lengths, discocellular of hind wing decidedly
curved. The length of the stalking of the forewing
veins should not be given too much importance in
identifying species, for it is a variable character in any
considerable series of any given species. Alar expanse,
12-14 mm.
Typr Locatity: Hverglade, Fla. (type in USNM).
Foop pruants: Melanthara radiata (rearings of the
Special Survey of the Div. Foreign Plant Quar., U. S.
Bureau of Entomology and Plant Quarantine, Nos.
24851, 24867); Bidens (larva in the flower heads).
DistrisuTion: Florida, Everglade (Apr.), Key West
(Apr.), Sugar Loaf Key (Mar.), Sarasota (Jan., Feb.,
Mar.).
425. Mescinia moorei, new species
Ficure 472
In color and superficial appearance most resembling
bacarella but differing in details of genitalia, a somewhat
broader extension of the whitish costal margin of forewing
and less contrasted dark border of the subterminal line.
The outstanding character seems to be the narrow,
evenly spaced, parallel, and somewhat elongate prongs of
the bifid apical process of gnathos (fig.472c). Veins 2 and
3 of forewing are stalked for about one-third and veins
4-5 for approximately half their lengths. Alar expanse,
11 mm.
TypEr Locauity: Mon Repos, British Guiana (type in
USNM, 61369).
Foop puant: Seeds of Wulffia.
Described from male type and female paratype from
the type locality reared by H. W. B. Moore, for whom the
species is named. There are no dates on the specimens
and they are in poor condition, somewhat broken though
not badly rubbed. I should not have described them
had they not been reared and been previously identified
as parvula Zeller and that name probably been given to
Mr. Moore. The species should be easily recognized if
other specimens from the host plant are discovered in
British Guiana.
426. Mescinia parvula (Zeller)
Fieures 471, 960
Ephestia parvula Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 249,
1881
Mescinia parvula (Zeller) Hampson, in Ragonot, Monograph,
pt. 2, p. 85, 1901.
Similar to the other species of the group in color and
markings, except for the much darker, more contrasted
lining of the veins of hind wing. Differs strikingly from
all other described species of the genus in the angulate
projection from the costa of harpe. Alar expanse, 12
mm.
Type Locatiry: Honda, Colombia (type in BM).
Foop PLant: Unknown.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Disrrisution: Cotomsta: Honda.
There are no examples of parvula in the U.S. National
Museum and the series in the British Museum is prob-
ably mixed, as Hampson mentions a specimen from
Colombia with veins 2 and 3 from the cell.
427. Mescinia commatella (Zeller)
Ficure 961
Ephestia commatella Zeller, Horae Soc. Ent. Rossicae, vol. 16,
p. 247, 1881.
Mescinia comatella (Zeller) Ragonot, Monograph, pt. 2, p. 84,
1901.
Known to me only from description and figure of the
female type. Apparently distinguished by the incom-
plete but strong white antemedian fascia, slanting
obliquely from inner margin to top of cell and parallel
with outer margin. Alar expanse, 13 mm.
The female genitalia of the type shows a single small
disklike signum.
Tyrer Locatity: Ceiba, Colombia (type in BM).
Foop prant: Unknown.
428. Mescinia berosa Dyar
Figure 966
Mescinia berosa Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 341, 1914.
The female type from which the original description
was drawn is faded and somewhat rubbed. A fresher
female example from Puerto Rico shows the dark areas
of forewing a dark vinous fuscous and the whitish costal
area somewhat speckled with rufous scales; discal and
terminal dots blackish. Hind wing smoky, the veins,
apical area, and terminal margin darker. Female geni-
talia with bursa copulatrix narrowly elongate; signum a
large elongate patch of many stout, bluntly pointed,
thornlike spines; ductus seminalis from bursa near junc-
ture of bursa and ductus bursae. Veins 2 and 3 of fore-
wing stalked for about two-thirds and 4 and 5 for
slightly more than half their lengths. Discocellular
vein of hind wing decidedly curved. Alar expanse, 12
mm.
Typr uocatity: Rio Trinidad, Panamé (type in
USNM).
Foop puant: Unknown.
DistrisutTion: PanamAé: Rio Trinidad (Mar.).
Purrto Rico: El Semil, near Villalba, elevation 1,700
feet (May).
Known to me only from two female examples in the
National Collection.
Genus Mescinia, Species 429: M. peruella
[Alar expanse more than 15, less than 20 mm.]
429. Mescinia peruella Schaus
Fieures 474, 964
Mescinia peruella Schaus, Proc. Ent. Soc. Washington, vol. 29,.
p. 186, 1927.—Wille, Rey. de Ent., Brazil, vol. 4, p. 455,
1934.
Forewing gray with costa whitish nearly to apex;
under magnification showing a sparse, scattered dusting:
of rufous scales; transverse markings and discal spots:
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 215
hardly distinguishable. Hind wing white, translucent,
extreme apex and terminal margin faintly smoky.
Male genitalia with apical process of gnathos mod-
erately large (proportionally about twice the size of
that organ in the foregoing species group). Female
genitalia with spines of signum bluntly and broadly
rounded. Veins 2 and 3 and 4 and 5 of forewing stalked
for about one-third their lengths (in some specimens 2-3
stalked for nearly one-half). Discocellular vein of hind
wing very slightly curved. Alar expanse, 15-18 mm.
Typr Locauity: Cafiete Valley, Peri (type in
USNM).
Foop pxiant: Cotton (larvae feeding in bolls).
Distrisution: Pert: Cafiete Valley (Mar.), Lima
(Sept.).
Genus Mescinia, Species 430 and 431: M. discella
and M. indecora
[Alar expanse over 20 mm.]
430. Mescinia discella Hampson
Ficures 475, 967
Mescinia discella Hampson, in Ragonot, Monograph, pt. 2, p. 84,
1901.
Forewing a pale purplish fuscous on lower half,
yellowish (bronzy) above, shading to white on outer
half of subcostal area; costal edge yellowish brown
with some blackish scales towards base; a bright
orange-yellow patch towards end of cell; along median
fold a fine line of blackish scales; a similar blackish line
along the fold replacing vein 1c; from end of cell to sub-
terminal line blackish dashes on veins 2 and 3, 4 and 5
and 6, more or less fusing to form a large, conspicuous,
dark blotch; discal spots fused into a blackish line along
discocellular vein; a row of faint black dots along
termen; antemedial line very faint, narrow, white, out-
wardly angled at lower margin of cell and inwardly
angled at vein 1b; subterminal line sharply angled
inward at vein 6, thence curved outward around the
blackish blotch, disappearing near costa and inner
margin. Hind wing whitish, semihyaline; the veins
and terminal margins faintly and finely outlined with
purplish ocherous; veins 7 and 8 anastomosed for little
more than half the length of vein 7; discocellular vein
decidedly curved. Alar expanse, 20-21 mm.
Male genitalia with bifid apical process of gnathos
elongate, rather large; anellus with enlarged base and
curved lateral arms; penis with a strong cluster of
sclerotized folds. Female genitalia with signum re-
placed by a concentration of scobinations; ductus
bursae weakly sclerotized towards genital opening;
ductus seminalis from bursa near its junction with
ductus bursae.
TYPE LOcALITY: Jalapa, México (type in USNM).
Foop pirant: Unknown.
Disrrisution: Méxtco: Jalapa, Orizaba.
MALA: Volcfén Santa Maria (Nov.).
The species is easily recognized by its bronzy color
and the conspicuous blackish markings beyond the cell.
GUATE-
431. Mescinia indecora Dyar
Ficures 476, 968
Mescinia indecora Dyar, Ins. Insc. Menstr., vol. 8, p. 35, 1920.
Forewing grayish fuscous with a faint purplish tint;
the median area between faint, dark, antemedian and
subterminal lines weakly shaded with blackish fuscous;
costal area narrowly sordid whitish; discal dots blackish.
Hind wings semihyaline, the veins faintly outlined in
ocherous fuscous, a similar narrow shade along termen;
veins 7 and 8 anastomosed for little more than half the
length of vein 7; discocellular vein curved. Alar
expanse, 22—23 mm.
Male genitalia with uncus broadly triangulate,
slightly tapering and with apex broadly rounded; apical
process of gnathos completely fused, large, roughly
triangular and with angles more or less rounded; anellus
with lateral arms broad and narrowly divided; vin-
culum stout and about as long as broad. Female
genitalia of type damaged, so that little remains of
bursa except the signum, which is a rather large,
flattened, platelike thorn.
Typr Locatity: Zacualpin, México (type in USNM).
Foop piant: Unknown.
Disrrisution: México: Zacualpdén (July), Techua-
cin (July).
The species fits badly in Mescinia on genitalic char-
acters. Uncus, anellus, gnathos, the short vinculum,
and the female signum are unlike any of these structures
in the other species of the genus and suggest a separate
generic placement, which may be necessary when more
material can be studied and something is known of the
earlier stages and biology. At present indecora is
represented by only two specimens in the National
Museum.
Genus 122: Nonia
[Venational division B. Forewing with 9 veins: 10 from the
cell, 8 and 9 united, 4 and 5 stalked, 2 and 3 united. Male
genitalia with transtilla incomplete.]
122. Genus Nonia Ragonot
Nonia Ragonot, Monograph, pt. 2, p. xiii, 1901.—Hampson, in
Ragonot, Monograph, pt. 2, p. 260, 1901. (Type of genus:
Homoeosoma exiguella Ragonot.)
Hypermescinia Dyar, Proc. Ent. Soc. Washington, vol. 47, p.
341, 1914. (Type of genus: Hypermescinia lambella Dyar.
New synonymy.)
Tongue well developed. Antenna pubescent; shaft
simple in both sexes. Labial palpus obliquely up-
turned. Maxillary palpus filiform. Forewing smooth;
9 veins; veins 2 and 8 united from very near angle of
cell; 4 and 5 stalked; 8 and 9 united; 10 from the cell,
separate from 8-9; male with fringe of hair or scales
from underside of costa towards base, but without
costal fold. Hind wing with vein 2 from well before
lower angle of cell; 3 and 5 from angle and closely ap-
proximate at base; 6 from upper angle of cell; 7 and 8
completely fused beyond cell; discocellular vein curved;
cell one-third of wing length. Abdomen of male with
216
a strong pair of ventrolateral tufts from eighth segment.
Male genitalia of the Mescinia type but with all the
parts more slender. Female genitalia with signa a
cluster of several very small disks; ductus seminalis
from bursa near its juncture with ductus bursae.
The male genitalia show the close affinity of Nonia to
Mescinia and show that Dyar’s interpretation of the
forewing venation rather than that of Ragonot and
Hampson is the correct one; that is, 2 and 3 united and
4 and 5 stalked rather than 4 and 5 united and 3 stalked
with 5. Ragonot and Hampson are also in error in
regard to the discocellular vein of hind wing. I have
examined the type of exiguella, and it has the disco-
cellular vein decidedly curved, as have all the Central
and South American and Puerto Rican specimens in
the National Museum and Cornell University Collec-
tions. The genus contains only one described species.
432. Nonia exiguella (Ragonot)
Figures 89, 477, 969
Homocosoma exiguella Ragonot, Nouv. Gen., p. 35, 1888.
Nonia exiguella (Ragonot), Monograph, pt. 2, p. 260, 1901.
Hypermescinia lambella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p.
341, 1914 (new synonymy).
Forewing whitish gray overshaded (especially in fresh
specimens) with vinaceous fawn, whitish along costa;
antemedial line from costa beyond one-third, broken,
represented (except in the darkest, most strongly
marked specimens) by two or three blackish spots; a
dark fuscous shade on inner margin at base; subterminal
line, narrow, blackish, slanting, from costa to outer
fourth of inner margin; discal spots conspicuous and
confluent, blackish. Hind wings whitish in male,
somewhat fuscous in female, veins and termen slightly
darker. Alar expanse, 10-12 mm.
Male genitalia figured from types of exiguella and
lambella, which are identical in all details.
TYPE LocaALities: Colombia (eriguella, in Mus. Univ.
Berlin); Tabernilla, Panam4 (lambella, m USNM).
Foop pirant: Unknown.
Distripution: Pusrro Rico: Dorado (Dorado,
May), La Sardinera (June); San German (Mayagiiez,
Apr.); Isabela and Puerto Real (Vieques Isl., Apr.).
Virein Istanps: Kingshill (St. Croix, Nov., Dec.).
Jamaica. GuatEMALA: Cayuga (Feb., May). Pan-
AMA: La Chorrera (May), Porto Bello (Feb.), Taber-
nilla, Rio Trinidad (June). Cotompi1a. BRaziu:
Vicosa (Minas Gerais, Oct.). Paraguay: Villarrica
(July).
Genus 123: Phestinia
[Venational division B. Forewing with 10 veins; 10 from the
cell, 8 and 9 long stalked, 2 and 3 united.]
123. Genus Phestinia Hampson
Phestinia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 57,
1930. (Type of genus: Phestinia costella Hampson.)
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Tongue well developed. Antenna of female simple,
a few projecting scales at each of the joints. Labial
palpus upturned, slender, somewhat rough scaled;
second segment reaching to vertex; third segment nearly
as long as second, bluntly pointed. Maxillary palpus
small but rather broadly and flatly scaled (squamous)
and appressed to face. Forewing smooth; 10 veins;
veins 2 and 3 united, from just before lower outer angle
of cell; 4 and 5 from the angle, stalked for nearly half
their lengths, the stalk separated at base from 2-3; 6
from below upper angle of cell, straight; 8 and 9 long
stalked; 10 from the cell, separate at base from stalk of
8-9. Hind wing with vein 2 from well before lower
outer angle of cell; 3 and 5 from the angle, connate (or
very closely approximate) at base; 7 and 8 anastomosed
for most of their lengths beyond cell; cell about one-
third the length of wing; discocellular vein curved.
Female genitalia with bursa copulatrix smooth except
for signum; signum a small disk with short projecting
spine (or spines) from lower margin; ductus bursae con-
siderably longer than bursa, simple except for a mod-
erately broad sclerotization of the tube at genital open-
ing; ductus seminalis from bursa near its junction with
ductus bursae.
The genus was erected for a single species described
from a single female specimen (not a male as stated by
Hampson); so the foregoing diagnosis is incomplete.
However, the venation and female genitalia are sufii-
cient to show its distinctness from and close relationship
to Nonia. It differs from the latter chiefly in having
veins 9 of forewing and 8 of hind wing both present.
433. Phestinia costella Hampson
Fieures 92, $70
Phestinia costella Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5,
p. 57, 1930.
In general habitus similar to Nonia exiguella, but
with darker areas more suffused and dark antemedial
and discal markings less strongly contrasted. Hamp-
son’s description of the dark suffusion as “red-brown”
is somewhat exaggerated. There is a faint vinaceous
tint to the brown, but it is hardly describable as red.
Clarke has examined the genitalia of the female type
and furnished me with a sketch of the signum (fig. 970b)
and venation (fig. 92). I have seen no examples from
the type locality, but have before me a rather worn
female from the Cornell Collection, taken at San Ger-
man, Puerto Rico, Apr. 17, 1930, which is certainly
congeneric, and probably conspecific, with costella. It
differs in that the signum (fig. 970a) has only one spine
from the lower margin of signum. There are two spines
in the type of costella. It is also smaller. Otherwise
the Puerto Rican example agrees with the type. The
venational details of the two specimens are identical.
Alar expanse, 16-20 mm.
Tyrr Locatity: Constant Springs, Jamaica (type in
BM).
Foop prant: Unknown.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 217
Genera 124 and 125: Comotia and Bema
[Venational division E. Forewing with veins 9 and 4 absent,
rarely (in some specimens of Bema) with a vestigial indication of
9 from 8 near apex. Hind wing with discocellular vein straight,
vertical or slanting. Male genitalia with hooked apical process
of gnathos small, bifid or partially fused; transtilla incomplete;
uncus slender.]
124. Genus Comotia Dyar
Comotia Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 343, 1914.
(Type of genus: Comotia torsicornis Dyar.)
Antenna of male (fig. 478d) pubescent; basal segment
enlarged and elongate, flattened on inner side (not tri-
angular or armed with a spine as stated by Dyar); shaft
flattened, the first half-dozen segments enlarged and
excavated into a pocket (sinus) enclosing modified
scales and followed by a ridge of crests on the following
three or four segments; female antenna simple. Tongue
well developed. Labial palpus upturned. Maxillary
palpus filiform. Forewing narrowly elongate, smooth;
nine veins; vein 2 from before angle; 3 and 5 shortly
stalked; 10 from cell, close to 8; male without costal
fold. Hind wing with vein 2 from well before lower
angle of cell; 3 and 5 from the angle and approximate
at base; 7 and 8 completely fused beyond cell (rarely
a vestige of 8 discernible as a short spur near apex);
discocellular vein straight and vertical; cell very short,
less than one-fourth of wing length. Abdomen of male
with a pair of weak ventrolateral tufts from eighth
segment.
Male genitalia of the Mescinia type, but with vincu-
lum shorter (but slightly longer than broad), apical
process of gnathos bifid, aedeagus constricted towards
apex. Female with signum a single projecting disk,
ductus bursae unsclerotized at genital opening, ductus
seminalis from bursa near its junction with ductus
bursae.
434, Comotia torsicornis Dyar
Ficures 117, 478
Comotia torsicornis Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 343,
1914,
Forewing sordid white, lightly dusted with fuscous;
costal area not appreciably paler; discal dots at end of
cell separate, dark fuscous; some dark fuscous dots
along termen. Hind wing translucent, smoky white
with veins and termen faintly darkened. Alar expanse,
13 mm.
Male genitalia with some fine scobinations on penis.
TypE Loca.ity: Porto Bello, Panam& (Mar., type in
USNM).
Foop prant: Unknown.
Known only from the male type which is somewhat
rubbed. Fresh specimens would probably exhibit pat-
tern markings of a more distinctive type and contrasting
color. The female from Taboga Isl. which Dyar asso-
ciated with his type does not belong in Comotia. It is
& specimen of Bema myja.
300329—56—_15
435. Comotia conyergens (Dyar), new combination
Bema convergens Dyar, Ins. Insc. Menstr., vol. 7, p. 59, 1919.
A trifle larger than torsicornis and with slightly longer
labial palpi (they reach to vertex in torsicornis and
slightly beyond in convergens), but otherwise similar.
The species is known only from the male type and one
other male from the type locality (not a female, as
stated by Dyar). These specimens are somewhat
rubbed, so no distinct wing pattern is discernible. The
exact status of convergens will have to wait upon more
and better material of both it and torsicornis. I doubt
that it is specifically distinct. Dyar’s placement of it
in Bema was obviously a blunder. Alar expanse, 15
mm.
Type Locatity: Cayuga, Guatemala (May, type in
USNM).
Foop piant: Unknown.
Comotia sp.
Fiaure 971
The only female of the genus that I have seen is a
rubbed specimen in the Cornell Collection from Jajoma
Alta, Puerto Rico (collected by W. A. Hoffman, June 18,
1930). Enough of the scaling remains to indicate that
fresh specimens of Comotia will exhibit a rather well-
marked pattern with a sprinkling of rufous scales among
the fuscous scaling of the darker areas. The venation
of fore and hind wings is identical with that of the male.
I suspect that this specimen is a female of torsicornis,
but exact specific identification must wait upon receipt
of more material from Puerto Rico and Central America.
Alar expanse, 15 mm.
125. Genus Bema Dyar
Bema Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 356, 1914.
of genus: Bema myja Dyar.)
Relmis Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 336, 1914. (Type
of genus: Relmis ydda Dyar. New synonymy.)
Antenna of male pubescent, with basal segment en-
larged, triangular, and with shaft projecting from one
corner (fig. 479e); shaft slightly swollen towards base,
otherwise simple. Antenna of female simple. Male
head (fig. 479d) hollowed between antennae and with a
strongly sclerotized, hollowed tubercle projecting up-
ward from front and bearing a ring of modified scales;
these structures absent from female. Labial palpus
upturned. Maxillary palpus filiform. Forewing nar-
rowly elongate, smooth; normally with nine veins (in
some specimens, see figs. 119 and 120, a vestige of vein
9 present); vein 10 from the cell closely approximate to
8; 2 from before lower angle of cell; 3 and 5 separate;
male with costal fold. Hind wing with 2 from well be-
fore lower angle of cell; 3 and 5 from the angle and ap-
proximate at base; 7 and 8 completely fused beyond cell
(rarely with a vestige of vein 8 near terminal end of 7);
discocellular vein straight and slanting inward from
lower angle; cell short, less than one-third the wing
(Type
218
length. In male a pair of hair tufts on underside of
prothorax and lying within grooved forecoxae. Ab-
domen of male with two pairs of dorsolateral hair tufts
from eighth segment; ventral membranous area of eighth
segment covered with enlarged scales, these and the hair
tufts set in elongate sclerotized tubercles (fig. 479c).
Male genitalia similar to those of Nona except harpe
broadly oval, sclerotized costa strongly arched and ter-
minating in a short, free spur at apex; apical process of
gnathos partially fused; vinculum long and narrow, with
lateral margins subparallel (very slightly concave), and
anterior margin indented.
Female genitalia with signum a single projecting
digitate disk; ductus bursae unsclerotized at genital
opening; ductus seminalis from anterior end of bursa.
The genus is easily identified by its characteristic gen-
italia. It is difficult to see how Dyar justified his
separation of species into two genera (Bema and Relmis)
since he had a female of myja and females of ydda and
jfifaca before him and not a single structural character
to separate them except a trifling difference in the length
of the labial palpi, at best specific, and here no greater
between females of myja and ydda than between the
females and males of myja.
436. Bema neuricella (Zeller), new combination
Figures 118, 119, 120, 479, 972
Ephestia neuricella Zeller, Isis von Oken, p. 862, 1848.—Ragonot,
Monograph, pt. 2, p. 288, 1901.
Bema myja Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 336, 1914
(new synonymy).
Forewing grayish fuscous, basal area paler; ante-
medial white line straight and slanting outwardly from
costa to inner margin and with a blackish shading along
its outer margin; subterminal line obscure, whitish, in-
dented below costa (at vein 8) and, very slightly, at
submedian fold; discal dots obscure, blackish; veins
more or less outlined by dark scaling. In older speci-
mens the dark markings are a pale brownish fuscous.
Hind wings semitransparent, smoky white with veins
darker and a dark shade at apex and along termen.
Alar expanse, 15-19 mm.
Specific differences in male genitalia are slight in the
genus and comparative only, mostly in the size of the
aedeagi, the harpes, and the width of the vinculum.
Female genitalia with signum very small and of nearly
uniform size but variable in shape and number of digi-
tate projections. Variations in signa shown in figures
972a-d. Equal variability exhibited in a series from
one locality.
Type tocauities: “St. Thomas,’”’ [Virgin Islands]
(neuricella, 9, in Mus. Univ. Berlin); Taboga Isl.,
Panamé (myja, o', in USNM).
Foop PLant: Inga sp.
Distrizution: Cusa: Habana, Santiago (8 moths
reared under E. E. A. de Cuba No. 9600, from larvae
boring in seeds of Inga sp.; A. Otero, collector; only
host record available). Banamas: Cay Santo Domingo
(Sept.). Purrto Rico: Lares (July, Dec.). Viren
Isuanps: St. Thomas, Kingshill (St. Croix, Oct., Nov.,
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Dec.). Guaremata: Cayuga (Apr., May., June).
PanamA: Taboga Isl. (Feb.). Trinipap. FRENCH
Gurana: St. Laurent du Maroni.
This species is represented by 65 specimens (11 &
and 54 9) in the National and Cornell University Col-
lections. All the males are more or less rubbed and
show very little of the original color or pattern. The
females are in better condition, especially those from
Guatemala and St. Croix, and once the sexes are prop-
erly associated it is not difficult to identify the species.
437. Bema fritilla Dyar
Bema fritilla Dyar, Ins. Insc. Menstr., vol. 7, p. 59, 1919.
The only representative of this species is the male
type. It is badly rubbed and shows no trace of the
original pattern. I suspect that it is nothing but a
runted specimen of neuricella (=myja). There is
nothing to distinguish it from the type of myja except
its smaller size (11 mm.) and proportionally smaller
genitalia, which are otherwise identical. The difference
in anal tufts which Dyar thought he saw was purely
imaginary. The white scaling he mentions is that on
the membranous area of the eighth segment and is
identical on the two types. I am retaining the name
for the present, pending receipt of similar small males
from the type locality. The female from Cayuga
(May) which Dyar associated with his type is not con-
specific or even congeneric. Itis a Unadilla and prob-
ably a small specimen of maturella Zeller, if what we
have under that name is correctly identified.
TypE Locatiry: Cayuga, Guatemala (June, type in
USNM).
Foop prant: Unknown.
438. Bema ydda (Dyar), new combination
Figure 974
Relmis ydda Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 337, 1914.
Forewing dark gray (in fresh specimens with a faint
violaceous tint); base (and thorax) darker; antemedial
pale line faint, bordered outwardly by a broad dark
band; subterminal line pale, indented as in myja,
bordered inwardly by a broad dark shade reaching to
cell; outer area beyond subterminal line of a similar
dark shade; discal dots obscured, more or less fused.
Hind wing smoky fuscous. Abdominal tufts of eighth
segment appreciably stouter than those of other species
of Bema. Alar expanse, 19-20 mm.
Male genitalia with vinculum broader in proportion
to its length than that of myja or the other species of
the genus. The entire genitalia are larger than those
of myja, otherwise similar. Female genitalia as in myja
except for enlarged signum.
TypE Locatity: Rio Trinidad, Panamsé (May, June;
type in USNM).
Foop pirant: Unknown.
Represented in the National Collection only by the
female type and one other female from the type locality
and a male from St. Jean Maroni, French Guiana.
The outer border of the antemedial and the inner border
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 219
of the subterminal lines have the appearance, to the
naked eye, of broad, dark, weakly contrasted, trans-
verse fasciae.
439. Bema yddiopsis (Dyar)
Relmis yddiopsis Dyar, Ins. Insc. Menstr., vol. 7, p. 58, 1919.
Similar to ydda except ground color paler. Thorax
and base of forewing pale clay color; terminal area of
forewing also paler than the dark shade inwardly border-
ing the subterminal line. Alar expanse, 20-21 mm.
Male genitalia similar to those of myja except some-
what larger; vinculum as narrow in proportion as that
of myja.
Type Locauity: Ténamo, Cuba (Mar., type in
USNM).
Foop piant: Unknown.
Known only from the type and one other male from
the type locality. The type (in fair condition) shows
a few reddish scales in the dark area bordering the sub-
terminal line, but I am unable to find any trace of the
“red streak along submedian fold” mentioned by Dyar.
440. Bema fifaca (Dyar), new combination
Figure 973
Relmis fifaca Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 337, 1914.
Known only from the unique female type. This
specimen similar in color and markings to fresh speci-
mens of myja, of which it may be only a variety. The
labial palpi seem a, trifle longer than those of myja and
the signum is somewhat larger but not different enough
to rule it out of that species. The outward dentations
of the antemedial whitish line, mentioned by Dyar,
also appear in some specimens of myja, depending upon
how much the specimen is rubbed or how much the
dark outer shading encroaches upon the white line.
They are not constant, and the line properly should be
described as slanting and straight. Alar expanse,
18 mm.
Type tocauity: Porto Bello, Panama (Dec., type
in USNM).
Foop piant: Unknown.
Genera 126-128: Homoeosoma to Rotruda
[Venational division C. Forewing with 10 veins: 10 from cell,
8 and 9 united, 4 and 5 stalked or connate, 2 and 3 from cell.
Hind wing with veins 7 and 8 strongly anastomosed; 3 and 5
from cell; discocellular straight, vertical. Male genitalia with
uncus broad, subtriangular; apical process of gnathos fused;
vinculum stout, broad. Female genitalia with or without
Signum; bursa, except for signum and some weak scobinations,
not sclerotized; ductus seminalis from bursa or ductus bursae.]
126. Genus Homoeosoma Curtis
Homoeosoma Curtis, Ent. Mag., vol. 1, p. 190, 1883.—Hulst,
Phycitidae of N. Amer., p. 191, 1890.—Ragonot, Mono-
graph, pt. 2, p. xiii, 1901—Hampson in Ragonot, Mono-
graph, pt. 2, p. 227, 1901.—Janse, Journ. Ent. Soc. South
Africa, vol. 8, p. 26, 1945. (Type of genus: Phycis gemina
Haworth, synonym of Tinea sinuella Fabricius; figs. 112,
480, 978.)
Phycidea Zeller, Isis von Oken, p. 178, 1839.
Tinea sinuella Fabricius.)
Tongue well developed. Antenna of male pubescent
and with a slight notch at base of shaft; of female,
simple. Ocelli present but small and lenses flattened
in some specimens. Frons rounded, labial palpus up-
curved; somewhat rough scaled; reaching to or slightly
above vertex; third segment slightly shorter than
second. Maxillary palpus filiform. Forewing smooth;
narrowly elongate; termen slanting; 10 veins: veins 2
and 3 from near lower outer angle of cell, separate; 4 and
5 stalked for at least half the length of vein 5; 6 from
below upper angle of cell, straight; 9 absent; 10 from
cell; male with a slight costal fold at base of wing.
Hind wing with seven veins; vein 2 from before lower
angle of cell; 3 and 5 from angle of cell and closely
approximate at base; 6 from upper angle of cell; 7 and
8 fused beyond cell almost to costa; discocellular vein
straight, nearly vertical and obsolescent; cell one-third
to two-fifths of wing length. Eighth abdominal seg-
ment of male simple (fig. 482e) with a single stout,
central thorn associated with sternite (figs. 483f-g), or
with a pair of weak ventrolateral hair tufts (fig. 489e).
Male genitalia with uncus broad, subtriangular,
outer surface densely covered with bristlelike scales.
Gnathos terminating in a broadly triangulate or tear-
shaped fused apical process with a very small spine at
caudal end. Harpe simple; costa strongly sclerotized
for most or all of its length; apex broadly rounded.
Anellus U-shaped with narrow basal plate and long
slender lateral arms, or semitubular. Aedeagus more
or less sinuate, smooth or with a few minute scobina-
tions at apex. Penis with a patch or elongate roll of
sclerotized wrinklings, otherwise unarmed. Vinculum
stout and broad, short to long and with terminal
margin broadly rounded.
Female genitalia with bursa copulatrix membranous,
finely scobinate over most of its inner surface; signum
a coarsely spined plate or altogether absent. Ductus
bursae membranous throughout, finely scobinate only
at or near junction with bursa copulatrix. Genital
opening simple (unsclerotized and without adjacent
sclerotized plates or other armature). Ductus semi-
nalis from bursa near its junction with ductus bursae.
Ovipositor lobes membranous or sclerotized.
The genus, as here defined, is composed of two quite
dissimilar groups which eventually will require generic
separation. However, until the exact status of Pata-
goma is established they may remain together in
Homoeosoma, for I believe that the name Patagonia can
be used eventually to include the whole of one species
group. From Homoeosoma I am removing mucidel-
lum Ragonot, reliqueella Dyar, olivaceella Ragonot,
musiosum Dyar, cubellum Dyar, affusellum Ragonot, and
the European nimbellum to a new genus (Rotruda), since
they differ strikingly on both male and female genitalic
characters from other species generally referred to
Homoeosoma. Further generic division of Homoeosoma
may be necessary when the genitalia of Old World
(Type of genus:
220
species can be thoroughly studied. Our American
species divide as follows:
Male with apical process of gnathos broadly triangulate
(figs. 481, 975), anellus U-shaped; female with signum,
ovipositor lobes unsclerotized.
Male with apical process of gnathos considerably smaller
and tear-shaped, anellus semitubular; female without
signum, ovipositor lobes sclerotized.
The species of the first group are further divisible into
subgroups on the following male characters:
Highth abdominal segment simple.
Eighth abdominal segment with a strong thornlike process
associated with sternite.
Highth abdominal segment with a pair of ventrolateral scale
tufts.
Genus Homoeosoma, Species 441-447: H. electel-
lum to H. illuviellum emendator
[Male with apical process of gnathos broadly triangulate, anellus
U-shaped, eighth abdominal segment simple; female with signum,
ovipostor lobes unsclerotized.]
44). Homoeosoma electellum (Hulst)
Figures 481, 975
Anerastia electella Hulst, Ent. Amer., vol. 3, p. 137, 1887.
Ephestia opalescella Hulst, Ent. Amer., vol. 3, p. 138, 1887 (new
synonymy).
Homoeosoma texanella Ragonot, N. Amer. Phycitidae, p. 15, 1887.
Homoeosoma tenuipunctella Ragonot, N. Amer. Phycitidae, p. 15,
1887.
Homoeosoma olectella (Hulst), Phycitidae of N. Amer., p. 198,
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 243,
1901.
Homoeosoma epalescella (Hulst), Phycitidae of N. Amer., p. 192,
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 243,
1901,
Homoeosoma electellum (Hulst), U. 8. Nat. Mus. Bull. 52, p. 434,
1903.—Kearfott, Canadian Ent., vol. 37, p. 123, 1905 (de-
scription of larva).—Barnes and McDunnough, Contribu-
tions, vol. 8, p. 200, 1916.—Forbes, Cornell Mem. 68, p. 634,
1920.—McDunnough, Check list, No. 6374, 1939.—Satter-
thwait and Swain, Journ. Econ. Ent., vol. 39, p. 575, 1946.
Homoeosoma opalescellum (Hulst), U. S. Nat. Mus. Bull. 52,
p. 434, 1903—McDunnough, Check list, No. 6375, 1939.
Homoeosoma differtella Barnes and McDunnough, Contributions,
vol. 2, p. 184, pl. 4, fig. 9, 1913.
This and the three species following have similar
genitalia and cannot be separated by any consistent
characters in these organs. In all, the vinculum is pro-
duced dorsolaterally into a pair of free arms (fig. 481a),
the lateral margins of the apical process of gnathos are
deeply concave (fig. 481c), and the signum is situated
near the cephalic end of the bursa. There is consider-
able variation in the gnathi and signa, but it is more
individual than specific in character. H. electellum is
readily separable from the other three species on its
forewing pattern. The contrasting white subcostal
streak (when distinguishable) is diagnostic. It is more
or less obsolescent in the paler specimens. Wing color
varies from pale powdery gray to whitish ocherous.
The name opalescellum applies to the more yellowish
specimens but represents nothing more than a color
form. H. electellum is close to the Kuropean nebulellum
UNITED STATES NATIONAL MUSEUM BULLETIN 207
which has similar genitalia and larval habits, nebulellum
being an important enemy of sunflower in Russia (see
V. Schzegoleff, Journ. Oil Industry, Moscow, p. 44,
November 1928). However, there appear to be enough
differences in habitus to distinguish the two as distinct
species. Alar expanse, 15-25 mm.
TyeE Locatities: Blanco County, Tex. (electellum,
in AMNH, ex Rutgers); Texas (texanellum, in Paris
Mus.); California (opalescellum, in USNM; tenuipunc-
tellum, in Paris Mus.) ; Everglades, Fla. (differtellum, in
USNM).
Foop piants: Flower heads of various asteraceous
plants (Anthemis, Aster, Bidens, Brauneria, Chry-
santhemum, Coreopsis, Dahlia, Helianthus, Heliopsis,
Rudbeckia, Tagetes, Viguiera, Ximenesia); also from
flowers of Opuntia and cotton and several U. S. Dep.
Agr. rearings (at Mission, Hidalgo, and Brownsville,
Tex.) from fruit of orange (on the trees). Sunflower
seems to be the most favored food plant.
Distrisution: Unitep States: Florida, Miami
(Apr.), Orlando (June); Lowisiana, Baton Rouge
(July), Vernon Parish (July); Texas, Blanco County,
Brownsville (Mar., Apr., Aug., Sept.), College Station
(June), Hidalgo, Liberty (July), Mercedes (Feb.),
Mission (Jan.), New Braunfels (May), Presidio (Sept.),
Sabinal (Mar.), San Benito (July, Aug., Sept.), San
Diego (May), Smith Point (Sept.), Victoria (Mar.,
May, Sept.), Zavilla (Apr.); Mississippi, Natchez
(May); Missouri, New Madrid (Aug.), Scott County
(Oct.); Jowa, Ames (Aug.), Sioux City (June); Kansas,
Onaga; Nebraska, Fort Calhoun; South Dakota, Elk
Point (Aug.); Colorado, Boulder (Sept.), Denver
(June), Pueblo (Sept.); New Mexico, Fort Wingate
(May, June), Jemez Springs (Apr.), Las Vegas, Pecos
(June); Arizona, South Liberty (Sept.), Santa Rita
Mts. (Aug.); California, Sacramento (Aug.); Wash-
ington, Bonneville (July), Pullman (June, July, Aug.,
Sept.), Walla Walla (June, July), Wenatchee (Aug.).
México: Cuernavaca (July), Orizaba (May), Tehuacin
(May, June). GuatTemata: Guatemala City. CuBsa:
Havana (Sept.). Bermupa (Jan., Mar., Apr., May,
July).
The foregoing localities are for specimens before me.
The species is generally distributed throughout the
United States.
442. Homoeosoma stypticellum Grote
Figure 976
Homoeosoma stypticella Grote, Bull. U. S. Geol. Geogr. Surv.
Terr. vol. 4, p. 703, 1878.—Hulst, Phycitidae of N. Amer.,
p- 193, 1890.—Hampson, tn Ragonot, Monograph, pt. 2,
p. 252, 1901.
Homoeosoma uncanale Hulst, Trans. Amer. Ent. Soc., vol. 3,
p. 162, 1886; U. S. Nat. Mus. Bull. 52, p. 434, 1903.—
Barnes and McDunnough, Contributions, vol. 4, p. 175,
1918.—McDunnough, Check list, No. 6324, 1939. (New
synonymy.)
Homoeosoma uncanalis Hulst, Phycitidae of N. Amer., p. 192,
1890.
Homoeosoma stypicellum Hulst, U. 8S. Nat. Mus. Bull. 52, p. 434,
1903 (misspelling).
Homoeosoma stypticellum (Grote), Forbes, Cornell Mem. 68,
p. 634, 1920.—McDunnough, Check list, No. 6371, 1939.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 221
Distinguished from other species with the electellum
type of genitalia by the broad, transverse, antemedial
dark brown band and the more or less extended dark
shading on inner margin of subterminal line of forewing,
this latter dark shading sometimes extending to the
dark discal spots at outer margin of cell. Hind wings
smoky fuscous. Alar expanse, 18-25 mm.
I agree with McDunnough that the female from
Colorado in the Hulst Collection which McDunnough
has so labeled is the probable type of uncanale. The
differences shown in figure 976 between it and the
signum of typical eastern siypticellum are only individual
in character and no more than those exhibited by other
eastern examples of stypticellum. I find nothing on
which to distinguish uncanale, even as a western race,
from stypticellum.
Typrr tocauities: Maine (stypticellum, in BM [?]);
Custer County, Colo. (uncanale, in AMNH, ex Rutgers).
Foop piant: Cirsium, Rhus.
Distrisution: Unrrep States: Maine, Orono, Bar
Harbor (July), Wales (July); Massachusetts, Martha’s
Vineyard (July, Aug.); Connecticut, East River (July,
Aug.); Rhode Island, Weekapaug (July, Aug.); New
York, Mlion (June); Pennsylvania, Oak Station (Mar.),
New Brighton (July); Jilinois, Palos Park (Aug.);
Arkansas, Washington County (July); Texas; Colorado,
Custer County, Glenwood Springs (Aug.), Logan’s
Peak (July). Canapa: Quebec, St. Hilaire (July);
Ontario, Ottawa (June), Trenton (June); Manitoba,
Aweme (June, July), Rounthwaite (Aug.); Saskatch-
ewan, Regina (July).
Among the foregoing, one example (from Palos Park,
Til., Aug. 6, 1939) was reared by A. K. Wyatt from
larvae feeding in the flower heads of ‘‘swamp thistle.”
443. Homoeosoma striatellum Dyar
Fiaurt 977
Homoeosoma striatellum Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 38, 1905.—McDunnough, Check list, No. 6367, 1939.
Pale slate gray; the veins of forewing outlined by
blackish scaling, transverse lines nearly obsolete, in
some specimens indicated. by a narrow, dark, angulate,
antemedial band and some dark shading towards costa,
on inner side of subterminal oblique line; discal spots
faintly indicated or absent. Hind wings whitish to very
pale smoky fuscous. Signum of female somewhat vari-
able, the extent of variation shown in the figure. Alar
expanse, 18-23 mm.
Typr Locauity: Phoenix, Ariz. (type in USNM).
Foop puant: Unknown.
Distripution: Arizona, Baboquivari Mts., Ajo,
Pima County (Mar.), Phoenix (Mar., Apr.), Tempe
(Feb.); California, Death Valley (Apr.).
444, Homoeosoma oslarellum Dyar
Figure 979
Homoeosoma oslarellum Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 38, 1905.—McDunnough, Check list, No. 6373, 1939.
Dark grayish fuscous with a scattering of whitish
scales along costa of forewing; transverse marking obso-
lete. Hind wing pale smoky fuscous. Alar expanse,
17-23 mm.
Female genitalia with signum similar to that of
striatellum, considerably larger than that of either
stypticellum or electellum, situated somewhat nearer the
middle of the bursa.
Type tocauiry: Chimney Gulch, Golden, Colo.
(type in USNM).
Foop piant: Unknown.
Distrisution: Colorado, Golden; New Mexico, Beulah
(July); California, San Francisco (Apr.).
445. Homoeosoma oslarellum breviplicitum, new race
A Southern California race differing from typical
oslarellum by the much shorter roll of sclerotized wrink-
lings of penis (but half the length of that in oslarellum
or the other preceding species), its paler wing color and
forewing markings. Color of forewing slate gray (inter-
mediate between that of oslarellum and striatellum); a
faint but distinguishable and sharply angulate, narrow,
dark, antemedial band; veins more or less outlined by
blackish scaling, especially in outer area of wing; hind
wing more whitish and with the veins more darkly con-
trasted than in typical oslarellum. Alar expanse, 20-25
mm.
Type tocauity: San Diego, Calif. (type in USNM,
61370).
Foop prant: Unknown.
Described from male type and male paratype from
the type locality (May 3, 1924, H. G. Dyar); one female
paratype from San Diego (June 14, 1924, J. M. Dam-
mers) and three female paratypes from Riverside,
Calif. (Apr. 14 and 17, 1937, Grace B. and John L.
Sperry).
446. Homoeosoma illuviellum Ragonot
Figure 482
Homoeosoma illuviella Ragonot, Nouv. Gen., p. 33, 1888; Mono-
graph, pt. 2, p. 245, 1901—Hulst, Phycitidae of N. Amer.,
p. 192, 1890.
Homoeosoma candidella Hulst, Ent. Amer., vol. 4, p. 118, 1888.
Homoeosoma illuviellum Ragonot, Hulst, U. 8. Nat. Mus. Bull.
52, p. 484, 1903.—McDunnough, Check list, No. 6366, 1929.
White with a faint scattering of dark scales on fore-
wing; transverse lines obsolete, their usual position
indicated only by a couple of dark dots in the ante-
medial and a very faint transverse shading of dark dust-
ing in the subterminal areas; costal edge dark beyond
middle; a small but rather conspicuous dark dot at
lower outer angle of cell and a much smaller, more
obscure dot at upper outer angle. Hind wings pale
smoky fuscous; cilia white. Alar expanse, 22-28 mm.
Male genitalia with apical process of gnathos stouter
and with less deeply concave lateral margins than in
the preceding species; vinculum not produced dorso-
laterally.
Typr tocaities: Sonora, México (iluviellum, in
Paris Mus.); Arizona (candidellum, in AMNH, ex
Rutgers).
Foop puant: Unknown.
222
DistRipution: Unirep Sratres: Arizona (no further
locality), Prescott (July); Colorado. M*#xtco: Sonora.
The type and only specimen in Paris is from “Senora”
[sic], México, Morrison, 1883, labeled in Ragonot’s
handwriting “Homoeosoma illunella Rag. pl. xxxii,
fig. 11, type original.” It is a female and not a male as
stated in the original description. I have examined the
genitalia. The type of candidellum has been examined
by Ragonot as the label indicates. It also is a female
with a glued-on abdomen. However, there is no doubt
that it is conspecific with dlunellum.
447. Homoeosoma illuviellum emendator, new race
Fiaure 980
Differs from typical dluviellum only in color and in-
tensity of markings; head, thorax, and forewing a pale
slate gray rather than white; dark spots as in Wluviellum
but more conspicuous and the dark shading along outer
line more pronounced; hind wings darker. Alar ex-
panse; 25-29 mm.
Genitalia as in dluviellum.
The new name is given with considerable hesitation
as this may prove to be only a colorform. However, as
several distinct species in Homoeosoma cannot be sepa-
rated by genitalia and the specimens before me are so
different in color from true dluviellum, I suspect that
they represent something more than a mere color form,
possibly a food-plant race.
Type Locauity: Chimney Gulch, Golden, Colo.
(May; type in USNM, 61371).
Foop piant: Unknown.
Described from male type, one male paratype from
Breckenridge, Colo. (June), one female paratype from
Eureka, Utah (June 2, 1917, Tom Spalding), and two
female paratypes from Richfield, Utah (May 20, 22,
1930).
Genus Homoeosoma, Species 448-453: H. imitator
to H. deceptorium
[Male with apical process of gnathos broadly triangulate, anellus
U-shaped, eighth abdominal segment with a strong thornlike
process associated with sternite; female with signum, ovipositor
lobes unsclerotized.]
448. Homoeosoma imitator, new species
FIGURES 483, 984
In color and markings similar to striatellum Dyar but
strikingly different in structure. The presence of the
thornlike process on eighth abdominal segment at once
distinguishes it from all the species of the foregoing sub-
group, and the produced lateral arms of vinculum from
all the species of this group except longiventrellum. The
eighth segment process varies in length in different
specimens. Extremes of variation are shown in figures
483f and 483g.
Alar expanse, 19-24 mm.
Type Locatiry: Palm Springs, Riverside County,
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Calif. (type in USNM, 61372; paratypes in Los An-
geles County Mus., Calif.).
Foop prant: Unknown.
Described from male type and three male and three
female paratypes from the type locality (Mar.), one
male paratype from West Riverside, Calif. (Feb. 16,
1906), and two female paratypes from Palo Verde,
Imperial County, Calif. (Feb. 10, 1940).
449. Homoeosoma longiventrellum Ragonot
Figure 484
Homoeosoma longiventrella Ragonot, Nouv. Gen., p. 34, 1888;
Monograph, pt. 2, p. 253, 1901.
Homoeosoma longiventrella noctividella Ragonot, Monograph, pt.
2, p. 253, 1901.
There are no specimens from tropical America agree-
ing with Ragonot’s description or figure of longiventrella
in the National Collection. The type is in the British
Museum but lacks an abdomen. In Paris, Clarke lo-
cated but one specimen in the Ragonot Collection, a
male from Chiriqui, Panamé, labeled Gn Ragonot’s
handwriting) “var. noctividella Rag.” If the genitalia
(figured) of this specimen represent, as they presumably
do, the true longiventrella the species should be easily
recognized, for it is the only one of the tropical species
with projecting arms from vinculum and a thornlike
projection from the eighth abdominal segment. The
projecting arms of vinculum are exceptionally long.
TypE Locatities: Chile (?) (longiventrellum, in BM);
locality not known to me (noctividellum, in Mus. Univ.
Berlin).
Foop prant: Unknown.
450. Homoeosoma albescentellum Ragonot
Fiaures 487, 983
Homoeosoma albescentella Ragonot, N. Amer. Phycitidae, p. 15,
1887.—Hulst, Phycitidae of N. Amer., p. 192, 1890.—
Hampson, in Ragonot, Monograph, pt. 2, p. 250, 1901.
Homoeosoma elongellum Dyar, Proc. Ent. Soc. Washington, vol.
5, p. 227, 1903.—McDunnough, Check list, No. 6365, 1929.
(New synonymy.)
Homoeosoma albescentellum (Ragonot) Hulst, U. S. Nat. Mus.
Bull. 52, p. 434, 1903.—McDunnough, Check list, No. 6368,
1939.
Rather pale brownish fuscous, dusted with white
scales and with darker markings accented. The white
scaling concentrated on the costal half of forewing,
giving the lower half a contrasting brownish shade;
transverse antemedial band blackish brown, normally
angulate but outwardly rounded in occasional speci-
mens; subterminal line inwardly angled at vein 6 and
inwardly shaded with blackish brown, especially towards
costa; discal spots at end of cell small, but distinct.
Hind wing smoky fuscous.
From others in the subgroup having a ventral, thorn-
like process from the eighth abdominal segment of the
male, this and the three species following are distin-
guished by their simple vinculi (without produced lat-
eral arms). Alar expanse, 21-30 mm.
Male genitalia figured from type of elongellum. Fe-
male genitalia figured from the type of albescentellum
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
in the Paris Museum. This latter is the only specimen
of the species in the Ragonot Collection. There are no
characters of structure or pattern by which elongellum
can be separated from albescentellum.
Type Locauitiss: California (albescentellum, in Paris
Mus.); Williams, Ariz. (elongellum, in USNM).
Foop pLant: Unknown.
Distrisution: California; Arizona, Williams (July),
Redington, Santa Catalina Mts.; Colorado, Silverton
(July).
451. Homoeosoma impressale Hulst
Ficures 486, 981
Homoeosoma impressalis Hulst, Trans. Amer. Ent. Soc., vol. 13,
p- 163, 1886; Phycitidae of N. Amer., p. 191, 1890.—
Hampson, in Ragonot, Monograph, pt. 2, p. 239, 1901.
Homoeosoma uncanalis Ragonot (not Hulst), Monograph, pt. 2,
p. 253, 1901.
Homoeosoma impressale Hulst, U. S. Nat. Mus. Bull. 52, p. 434,
1903.—Barnes and McDunnough, Contributions, vol. 3, p.
200, 1916; vol. 4, p. 174, 1918 —McDunnough, Check list,
No. 6363, 1939.
McDunnough’s identification (Contributions, vol. 4,
pt. 2, p. 174, 1918) of the true type of this species in the
Rutgers Collection is without any doubt the correct
one. The specimen is a male without abdomen,
labeled Nevada, ‘3838.’ It also bears a Hulst name
label ““Homoeosoma uncanale,’”’ which is obviously in-
correct and can be ignored as an error in labeling. The
Nevada specimen is one of the well-marked examples
of the species and agrees with the original description.
The markings are variable in the extreme, ranging
from pure white examples with only a couple of small
blackish discal dots at end of cell and a row of faint
blackish dots along termen to white examples with a
broad, slanting, blackish, antemedial band and a narrow
blackish subterminal band. On the right side of one
specimen before me from Manitoba the blackish bands
are fused, filling the entire center of the wing. The
specimens without transverse markings are superficially
like those of tlluviellum, usually showing the discal spot
at upper outer angle of cell a trifle stronger but other-
wise only separable on genitalia. The two figures in
the Ragonot Monograph (pl. 33, figs. 7, 18) are good
illustrations of the normal variation in pattern. Alar
expanse, 23-34 mm.
TYPE LOCALITY:
Rutgers).
Foop prant: Unknown.
Distrizution: Unirep States: Arizona; Nevada;
Colorado; Utah, Stockton (June, July); California,
Davis Creek, Modoc County (June); Washington,
Pullman (Aug.), Walla Walla (June). Canapa: Mani-
toba, Cartwright, Rounthwaite (Aug.).
Nevada (type in AMNH, ex
452. Homoeosoma inornatellum (Hulst)
Figures 485, 982
Euzophera inornatella Hulst, Canadian Ent., vol. 32, p. 173,
1900.—Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 228,
1904.
223
Homoeosoma inornatellum (Hulst), McDunnough, Check list,
No. 6369, 1939,
A very faintly marked species, especially in the male;
forewing pale gray or grayish fuscous with a faint, white
subcostal streak (much as in electellum); antemedial
markings consisting of one or two obscure dark dots;
discal dots at end of cell faint. In the female the
markings are somewhat stronger. Hind wings pale
smoky fuscous. Superficially the males look much like
some examples of electellum, but the two species are
easily separated on structure. In addition to the
differences in male genitalia and the eighth abdominal
segment, inornatellum has veins 2 and 3 of forewing
approximate at origin while in electellum these veins are
well separated, about as far apart as 3 and 4. Alar
expanse 23-25 mm.
From other species with simple vinculi (albescentellum,
impressale, deceptorium), inornatellum is readily dis-
tinguished by the shape of the harpe and the apical
process of gnathos. The signum of the female is
situated very near the anterior end of the bursa, as in
electellum. Dyar’s identification of the type of inorna-
tellum with stypticellum is an obvious error. The two
species are not even superficially similar.
Tyrr Locauity: Anglesea, N. J. (type in AMNH, ex
Rutgers).
Foop piant: Unknown.
Distrisution: New Jersey, Anglesea (May, June);
Pennsylvania, Mount Airy.
A series of males from the type locality is in the
National Collection.
453. Homoeosoma deceptorium, new species
Figures 488, 985
Color and markings as in the larger, most strongly
marked specimens of stypticellum, except that the white
dusting on forewing is more restricted to the costal half
of the wing; transverse antemedial band broad, slanting,
not appreciably angulate, blackish brown; subterminal
dark band narrower but distinct; discal spots at outer
angles of cell distinct. Hind wings pale smoky fuscous
with veins outlined by darker scaling. Alar expanse
23-27 mm.
Male genitalia chiefly distinguished from those of
other associated species in this subgroup by the shape
of the apical process of gnathos. Signum of female
located near middle of bursa.
TyprE Locauity: New Brighton, Pa. (type in USNM,
61373; paratypes in Canadian Nat. Coll.).
Foop piant: Unknown.
Described from male type and four male and one
female paratypes from the type locality (July, Aug.);
one male and one female paratype from Alcove, Quebec
(July 7, 1936, F. A. Urquart).
224
Genus Homoeosoma, Species 454: H. discrebile
[Male with apical process of gnathos broadly triangulate, anellus
U-shaped, eighth abdominal segment with a pair of ventrolateral
scale tufts; female with signum, ovipositor lobes unsclerotized.]
454, Homoeosoma discrebile, new species
Figures 489, 986
| White with a very faint cream tint and with three
trongly contrasted brown markings on forewing, a
transverse slanting antemedial band, a somewhat
narrower but equally distinct subterminal band and a
large discal spot formed by the fusion of the two dots
at outer angles of cell. Hind wings white; veins not
outlined by dark shading. Alar expanse, 15-17 mm.
Male genitalia similar to those of electellum; vinculum
with produced dorsolateral arms. Signum of female
large, situated well back of the anterior end of bursa.
Typr tocauity: “S. KH. Brazil” (F. D. Jones, ‘‘1920-
303”; type in BM; paratypes in BM and USNM,
61374).
Foop PLiant: Unknown.
Described from male type and three female paratypes
from the type locality, and one female paratype from
Castro, Parand, Brazil.
An easily recognized species.
Genus Homoeosoma, Species 455-461: H. pere-
grinum to H. nimbosellum
{Male with apical process of gnathos considerably smaller and
tear-shaped, anellus semitubular; female without signum, ovi-
postor lobes sclerotized.]
455. Homoeosoma peregrinum, new species
Fieurrs 490, 987
Forewing very narrow in the male (8 by 1/4 mm.); in
female somewhat broader. Color brown, evenly pep-
pered with whitish scales, making the general color
grayish fuscous; markings obscure, only the faintest
indication of a narrow antemedial dark line and the
discal spot at end of cell. Hind wing of a very pale
smoky hue with darkened veins and a fuscous line along
termen. Alar expanse, 15-17 mm.
Male genitalia chiefly distinguished from those of
other species in the group (except assitum) by the very
small, semicircular sclerotized area of penis (assitum is
distinguished by its smooth aedeagus and slenderer,
more evenly tapering harpes); aedeagus in peregrinum
finely scobinate at apex. The structural differences
among the species of this group are slight and hardly of
specific value. In the female the amount and extent
of scobination of the bursa varies somewhat from species
to species, but almost as much from specimen to speci-
men. These scobinations (except in oconequensis) are
more concentrated in a single circular area where they
have the appearance of a weak signum, which they are
not.
TYPE Locaity: Carmel, Calif. (typein USNM, 61375;
paratypes in USNM and Lange Coll.).
Foop PLant: Anaphalis margaritosa.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Described from male type and three male and two
female paratypes from the type locality, reared by
W. H. Lange (Mar. 4, 1938, and Mar. 25, 1939); and
one male paratype from Juan Vinas, Costa Rica (May).
H. peregrinum is the only species of the group: known
from the United States, and since it also occurs in Costa
Rica, the natural assumption is that it is a wanderer
from tropical America. It may be the species that
Hampson misidentified with his Old World ephestidiella
(in Ragonot, Monograph, pt. 2, p. 256, 1901). ‘The
latter belongs to quite a different species group which is
not found in the New World.
456. Homoeosoma vepallidum, new species
Figures 491, 991, 992
Pale brownish fuscous dusted with white, giving the
insect a sordid whitish tint; antemedial markings of
forewing reduced to a dark spot extending from imner
margin to vein 1b; discal dots at outer angles of cell
present but ebscure; subterminal line obscure, slanting,
narrow, white, and margined imwardly by a few very
faint, dark spots; a faint but distinct white subcostal
streak. Hind wing smoky white, veins faintly dark-
lined. Alar expanse, 18-20 mm.
Genitalia without outstanding specific characters.
Typz Locauity: “Villa Ana, F. C. S. F.,”’ Argentina
(type and paratype in BM).
Foop prant: Unknown.
Described from male type and female paratype from
the type locality, Nov. 29, 1923, R. J. Hayward. A
female from the Cornell University Collection, taken in
the Sierra de Cérdoba, Argentina, Mar. 9, 1920, is not
included among the types because it is paler in color
and has a larger bursa, copulatrix (fig. 992) than the
paratype from Villa Ana, but it is undoubtedly the
same species.
457. Homoeosoma ditaeniatellum Ragonot
Ficurs 990
Homoeosoma ditaeniatella Ragonot, Nouv. Gen., p. 33, 1888;
Monograph, pt. 2, p. 236, 1901.
According to Clarke’s notes this specimen is badly
rubbed and stained and the figure and description drawn
from it consequently somewhat misleading. The species
is probably considerably more whitish than the type
would seem to indicate. Its identification will have to
wait upon more material from the type locality. Alar
expanse, 18 mm.
TYPE LOCALITY: Quillota, Chile (Paulson, 1887; type
in Paris Mus.).
Known only from the female type.
458. Homoeosoma oconequensis (Dyar), new combination
FigurE 989
Eucampyla oconequensis Dyar, Ins. Insc. Menstr., vol. 7, p. 60,
A pale fuscous gray species almost unmarked; the
faintest indication of a narrow, transverse, dark ante-
medial line and a dark discal dot at lower outer angle
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
of cell. Hind wing pale smoky white; veins faintly
outlined. Alar expanse, 22 mm.
Genitalia distinguished by the very sparse scobina-
tion of bursa copulatrix without any trace of a concen-
tration of these scobinations into a conspicuous spot.
Tyrer Locauity: Oconeque, Carabaya, Peri (Schaus,
collector; type in USNM).
Foop prant: Unknown.
Known only from the female type.
459. Homoeosoma assitum, new species
Fiaures 492, 988
A small pale gray species, very faintly but appreciably
marked; forewing with a narrow, slanting, dark ante-
medial line; a very faint, straight, slanting, white, sub-
terminal line; two small, obscure discal dots at outer
angles of cell. Hind wing smoky white; veins darkly
outlined; a narrow dark line along termen; cilia white.
The species resembles peregrinum but is smaller and
paler. Alar expanse, 13-17 mm.
Male genitalia distinguished by their narrow, taper-
ing harpes and smooth aedeagus.
TYPE LocaLity: Cafiete, Pera (type in USNM, 61376;
paratypes in USNM, BM, Paris Mus.).
Foop pirant: Unknown.
Described from male type and seven male and four
female paratypes from the type locality, taken Mar. 2,
1943, on the foliage of a composite by E. J. Hambleton.
460. Homoeosoma acmaeopterum Ragonot
Ficure 493
Homoeosoma acmaeoptera Ragonot, Nov. Gen., p. 34, 1888;
Monograph, pt. 2, p. 255, 1901.
According to Clarke’s notes there are five specimens
under this name in the Ragonot Collection at Paris
representing what appears to be an assortment of spe-
cies. The male type, so labeled by Ragonot and from
which the figure in his Monograph (pl. 33, fig. 21) was
made, is the only specimen that can definitely be called
acmaeopterum. According to Clarke, the Ragonot
figure is a fair representation except that the forewing
is too short in proportion to its width and its median
dark area too strongly contrasted. Alar expanse, 22
mm.
Type Locauity: Coquimbo, Chile (type in Paris
Mus.).
Foop pirant: Unknown.
461. Homoeosoma nimbosellum Ragonot
Fiaure 494
Homoeosoma nimbosella Ragonot, Nouv. Gen., p. 34, 1888;
Monograph, pt. 2, p. 255, 1901.
The type is a male, and not a female, as stated by
Ragonot. According to Clarke, the Ragonot figure
(Monograph, pl. 30, fig. 22) is somewhat inaccurate,
being too dark and with the subterminal line too well
defined. Alar expanse, 18 mm.
Type xocatity: Valparaiso, Chile (type in Paris
Mus.).
Foop prant: Unknown.
Known only from the type in Paris.
8003293—bé6——16
225
Genus Homoeosoma, Species 462 (unplaced):
H. unionellum
462. Homoeosoma unionellum Ragonot
Homoeosoma unionella Ragonot, Nouv. Gen., p. 33, 1888.—
Hampson, in Ragonot, Monograph, pt. 2, p. 235, 1901.
A photograph of the type shows a large, narrow-
winged, pale specimen (yellowish white, according to
description) with costal edge of forewing beyond middle,
blackish. A sketch of the female genitalia shows that
the species belongs in the first group (Homoeosoma
Species 441-454), but more exact placement is impos-
sible without a male. Alar expanse, 28 mm.
TyrE Locatity: Milpas, México (type in BM).
Foop prant: Unknown.
Known only from the female type.
127. Genus Patagonia Ragonot
Patagonia Ragonot, Monograph, pt. 2, p. xiii, 1901—Hampson,
in Ragonot, Monograph, pt. 2, p. 236, 1901. (Type of
genus: Homoeosoma magellanella Ragonot.)
Characters of the second Homoeosoma species group,
except veins 4 and 5 of forewing connate rather than
stalked.
So far represented only by the single female example
of the type species. J suspect that this is an individual,
venational aberrant and that when further examples
are taken they will show the normal Homoeosoma vena-
tion of forewing, 4 and 5 appreciably stalked. If,
however, the connate condition of 4-5 were to prove
constant and the genitalia of the male show no marked
variation from those of the second Homoeosoma group,
the venational difference would have a purely specific
significance and Patagonia could be retained as a valid
genus on the genitalic characters of the group; and the
species of the group could be referred to it.
463. Patagonia magellanella (Ragonot)
Figure 993
Homoeosoma magellanella Ragonot, Nouv. Gen., p. 35, 1888.
Patagonia magellanella (Ragonot), Monograph, pt. 2, p. 226,
1901.
According to Clarke’s notes, the figure of the type in
Ragonot’s Monograph (pl. 33, fig. 25) is faulty in that
the forewing is too broad, the whitish costal streak not
extensive or white enough, the subterminal striations
too much accented, and the wing generally too dark.
However, the pattern should make specimens from the
type locality easily identifiable. Alar expanse, 19 mm.
Tyrer Locatity: Punta Arenas, Straits of Magellan,
Chile (type in Paris Mus.).
Foop pirant: Unknown.
128. Rotruda, new genus
Tyrer oF GeNus: Homocosoma mucidellum Ragonot.
Tongue, antenna, palpi, venation, and habitus as in
Homoeosoma. Male genitalia with a comb of strong
spines along side of aedeagus. Female genitalia with
226
two large signa composed of opposed, cup-shaped,
strongly spined concave plates; ductus seminalis from
ductus bursae.
In addition to mucidella and its American allies the
European Homoeosoma nimbella Zeller should be re-
ferred here (fig. 496).
464. Rotruda mucidella mucidella (Ragonot), new combination
Figures 495, 994
Homoeosoma mucidella Ragonot, N. Amer. Phycitidae, p. 15,
1887; Monograph, pt. 2, p. 258, 1901.—Hulst, Phycitidae
of N. Amer., p. 193, 1890.
Homoeosoma mucidellum Ragonot, Hulst, U. S. Nat. Mus. Bull.
52, p. 434, 1903.—Dyar, Pomona College Journ. Ent., vol. 2,
p- 378, 1910.—Barnes and McDunnough, Contributions,
vol. 2, p. 223, 1914.—McDunnough, Check list, No. 6370,
1939.
There is nothing except color differences to separate
typical mucidella from the following supposed species
which I am treating as races or synonyms, and these
differences are not clear-cut between tropical examples
and dark specimens from eastern United States. What
color differences there are depend upon the amount of
white dusting and the intensity of the dark markings.
Typical mucidellum is the most heavily dusted with
white and has the dark markings of forewing most
sharply contrasted (frequently blackish), and to the
naked eye seems a paler form than any of the others,
although there are intergrading examples between it
and religuella. Alar expanse, 15-22 mm.
There is considerable variation in the number of
spines in the comb along the side of aedeagus (10 to 15),
but this variation is as great between examples of
typical mucidella as it is between it and any of its races.
Tyrer Locatity: California (type in Paris Mus.).
Foop prants: Aster (and presumably other com-
posites).
Distrisution: Unirep Strarss: California, Clare-
mont, Colfax (July), Deer Park Springs (Lake Tahoe),
La Puerta Valley (July), Loma Linda (July), Los
Angeles County (Aug.), Polente (Sept.), Riverside
(Nov.), Sacramento (Sept.), San Bernardino Mts.
(July), San Diego (July), Shasta Retreat (Siskiyou
County, July, Aug., Sept.), Warner Mts. (Modoc
County, July); New Mexico, Jemez Springs, Las Vegas,
Hot Springs (June, Sept.); Arizona, Palmerlee, Tempe
(June); Colorado, Denver (July), Glenwood Springs
(Aug., Oct.); Utah, Provo (July), Salt Lake, Stockton
(June, Aug.); Wyoming, Jackson Hole (July); Oregon,
Crater Lake (July); Washington, Bellingham (June),
Pullman (May, July, Aug., Oct.), Snake River (July),
Walla Walla (June, July), Wenatchee (Aug.). Can-
ADA: British Columbia, Kaslo (June).
The range of distribution is probably the entire
United States, southern Canada, and northern México
from the Rocky Mountains westward to the Pacific
Ocean.
The European nimbella (fig. 496) is very close to
mucidella, differing only in the greater number of spines
on aedeagus (19 to 23). Were it not for this apparently
UNITED STATES NATIONAL MUSEUM BULLETIN 207
consistent difference I should be inclined to treat
mucidella as an American race of the Old World species.
465. Rotruda mucidella reliquella (Dyar), new combination
Homoeosoma reliquellum Dyar, Proc. Ent. Soc. Washington, vol. 6,
p- 112, 1904.—Forbes, Cornell Mem. 68, p. 634, 1920.—
MeDunnough, Check list, No. 6372, 1939.
Darker than typical mucidella with forewing some-
what suffused, the dark markings less sharply con-
trasted and the whitish dusting confined mostly to the
costal half of the wing. Apparently confined to the
areas of Canada and the United States east of the
Rocky Mountains.
Alar expanse, 14-19 mm.
TypE Locauity: Center Harbor, N. H. (type in
USNM).
Foop piants: Aster, Cirsium spinosisimum, Sitihas
caroliniana, Sonchus asper, lettuce.
Distrisution: Unirep Srares: New Hampshire,
Center Harbor (July), Hampton (June, July); Massa-
chusetts, Amherst (June), Cohasset, Framingham (May),
Winchendon (June); Connecticut, East River (July,
Aug.); New Jersey, Montclair (Aug.), Newark (Aug.);
Pennsylvania, New Brighton (May, June, July), Oak
Station (June), Pittsburgh (May, June, Aug.); Illinois,
Edgebrook (Oct.), Chicago (July) ; District of Columbia,
Washington (May); Maryland, Plummer’s Isl. (May);
Virginia, Great Falls (July); North Carolina, Tryon
(May); Georgia, Savannah (May); Florida, Big Pine
Key (Apr.), Chokoloskee, Everglades (Apr.), Ft.
Myers (Apr.), Lake Alfred (July), Miami (Apr.), Pom-
pano (Mar.). Cawnapa: Ontario, Trenton (May, June,
July, Aug.).
466. Rotruda mucidella olivaceella (Ragonot), new combination
Homoeosoma olivaceella Ragonot, Nouv. Gen., p. 33, 1888; Mono-
graph, pt. 2, p. 244, 1901.
Homoeosoma musiosum Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 342, 1915 (new synonymy).
Homoeosoma mucidella Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 342, 1915.
Homoeosoma cubella Dyar, Ins. Insc. Menstr., vol. 7, p. 62, 1919
(mew synonymy).
There is nothing, structural or otherwise, to separate
the female types of musiosum and cubella from each
other or from what Dyar correctly identified as muci-
della from Central America. I have examined the type
of olivaceella (&, in Mus. Univ. Berlin) as well as its
genitalia. It is merely a small (13 mm.), somewhat
rubbed and faded specimen. I am holding olivaceella
as a tropical American race of mucidella more on geo-
graphical distribution than anything else. It shows, in
contrast to typical mucidella, the same color variations
and obscurity of pattern markings as the eastern United
States race reliqguella. When examples are recovered
from the intermediate areas of northeastern México it
will probably be necessary to sink reliquella into the
synonymy of olivaceella. The races at best are dubious
entities, and I doubt that even the distinctness of
mucidella mucidella will survive extensive collection in
the central areas of United States.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Alar expanse, 12-18 mm.
Type Locauitizs: “St. Thomas,’ [Virgin Islands]
(oliwaceella, in Mus. Univ. Berlin); Porto Bello, Panama
(musiosum, in. USNM); Baracoa, Cuba (cubella, in
USNM).
Foop piants: Unknown.
Distrisution: Cusa: Baracoa. VircGin IsLanps: St.
Thomas. GuateMaLa: Cayuga (Apr., May), Quirigué
(Feb.). Costa Rica: Juan Vifias (June). PanamA:
Cabima (May), Corozal (Nov.), Porto Bello (Feb.,
Mar.), Rio Trinidad (Mar., May). Brazi: Sania Cata-
rina, New Bremen; Minas Gerais, Vicosa (Oct.). Para-
Guay: Villarrica (Sept., Oct.).
467. Rotruda mucidella affusella (Ragonot), new combination
Homoeosoma affusella Ragonot, Nouv. Gen., p. 34, 1888; Mono-
graph, pt. 2, p. 260, 1901.
The name affusella is doubtfully placed here in associ-
ation with mucidella, for Ragonot’s description and
figure suggest something quite different, possibly similar
to what I have described as Homocosoma vepallidum.
However, Clarke’s notes on the type discount the figure
(Monograph, pl. 33, fig. 24) and, to a considerable
degree, the original description. He states that “the
figure is poor. The dorsal spot at basal third is dis-
tinct but the excurved line costad is very faint. The
s. t. line of dashes, which is so prominent in the figure,
is scarcely discernible in the specimen. On the fore-
wing there is a heavier dusting of white than is indi-
cated in the drawing.’’ This suggests a rather pale,
not too well marked example of mucidella. We shall
have to await recovery of examples matching the type
from Argentina before the status of the name affusella
can be definitely settled. The abdomen of the type
was no help. It has been glued on and is obviously
spurious. I examined the genitalia (?). They are
those of E’phestia elutella.
Alar expanse, 18 mm.
Typr Locauity: Cérdoba, Argentina (type in Paris
Mus.).
Foop prant: Unknown.
Genus 129: Stirephomescinia
[Venational division B. Forewing with 9 veins: 10 from cell;
8 and 9 united; 4 and 5 united; 2 and 3 stalked. Male genitalia
with transtilla incomplete.]
129, Genus Strephomescinia Dyar
Strephomescinia Dyar, Ins. Insc. Menstr., vol. 7, pl 60, 1919.
(Type of genus: Strephomescinia schausella Dyar.)
Tongue well developed. Antenna pubescent; shaft
simple. Labial palpus upturned. Maxillary palpus
filiform. Forewing smooth; 9 veins; veins 2 and 3 long
stalked from angle of cell; 4 and 5 united; 6 straight; 8
and 9 united; 10 from the cell but closely approximate
to 8-9 for some distance; male with weak costal fold.
Hind wing with vein 2 from well before angle of cell,
straight and parallel with 3; 3 and 5 from angle and
227
closely approximate at base; 6 from upper angle of cell;
7 and 8 completely fused beyond cell; discocellular vein
curved; cell about one-fourth the wing length. Abdo-
men of male with a pair of ventrolateral hair tufts.
Male genitalia with uncus triangulate. Gnathos
terminating in a small bifid apical process. Harpe
simple; costa broadly sclerotized for most of its length.
Aedeagus straight, simple, moderately slender. Penis
very weakly scobinate, otherwise unarmed. Vinculum
stout, short, terminal margin truncate.
This genus is known only from the male of the type
species but is easily identified by its venation. The
genitalia show the close affinity of Strephomescinia to
both Homoeosoma and Mescinia.
468. Strephomescinia Schausella Dyar
Figures 93, 497
Strephomescinia schausella Dyar, Ins. Insc. Menstr., vol. 7, p. 60,
1919.
Forewing whitish gray, irregularly and faintly
marked with fuscous; a dark streak along submedian
fold and some dark streaking on the veins toward
termen; discal spots elongate, separate; no transverse
lines distinguishable. Hind wing translucent, with the
veins outlined by dark scaling; terminal and costal
margins narrowly dark-margined. The one available
specimen is badly rubbed. Fresh specimens would
probably exhibit a more definable pattern. Alar ex-
panse, 9.5 mm.
Male genitalia with apex of uncus bluntly pointed;
cucculus of harpe oval, apex evenly rounded; elements
of divided transtilla long and slender.
Type tocauiry: Santiago, Cuba (June; type in
USNM).
Foop prant: Unknown.
So far represented only by the unique male type.
Genus 130: Unadilla
[Venational division E. Forewing with 9 veins: 10 from cell;
9 absent (8 and 9 united): 4 absent (4 and 5 united); 2 and 3
from cell. Hind wing with discocellular vein straight and verti-
cal. Male genitalia with transtilla incomplete; uncus broad
throughout. Female genitalia with genital opening simple
(unsclerotized and without adjacent armature).]
130. Genus Unadilla Hulst
Unadilla Hulst, Phycitidae of N. Amer., p. 197, 1890.—Ragonot,
Monograph, pt. 2, p. xiii, 1901—Hampson, in Ragonot,
Monograph, pt. 2, p. 261, 1901.—Janse, Journ. Ent. Soc.
South Africa, vol. 8, p. 25,1945. (Type of genus: Unadilla
nasutella Hulst.)
Strymax Dyar, Proc. Ent. Soc. Washington, vol. 47, p. 344,
1914.—Richards and Thomson, Trans. Ent. Soc. London,
vol. 80, p. 201, 1932. (Type of genus: Sirymax dorae Dyar.
New synonymy.)
Tongue well developed. Antenna pubescent; shaft
simple. Labial palpus upturned, somewhat obliquely
so in the female. Maxillary palpus filiform. Forewing
smooth; 9 veins; veins 2 and 3 separate, 2 near and 3
from the angle of cell; 3 and 5 closely approximate at
base; 4 absent; 6 straight; 8 and 9 united; 10 from cell,
228
well separated from 8-9; male with a short costal fold.
Hind wing with vein 2 from well before angle of cell; 3
from before but near angle, approximate to 5; 6 from
near upper angle of cell; 7 and 8 completely fused be-
yond cell (in occasional specimens a rudiment of 8 visible
as a short spur from outer fifth of vein 7) ; discocellular
vein straight and vertical; cell approximately one-third
the length of the wing. Eighth abdominal segment of
male simple.
Male genitalia with uncus broad, subtriangular,
outer surface densely covered with bristlelike scales.
Gnathos terminating in a moderately large, fused or
partially fused, hooked apical process. Harpe simple;
costa, strongly sclerotized for most of its length; apex
broadly rounded. Anellus U-shaped; lateral arms
curved and partially encircling aedeagus. Aedeagus
straight; somewhat constricted at apex; smooth or
with a patch of minute scobinations at apex. Penis
with some scobinate wrinklings, otherwise unarmed.
Vinculum stout and broad, long and tapering slightly
to more or less narrowly rounded terminal margin.
Female genitalia with bursa copulatrix membranous,
finely scobinate over caudal half of inner surface;
signum consisting of a girdle of fine, narrow, serrate,
ridgelike disks about middle of bursa, or completely
absent. Ductus bursae membranous throughout, finely
scobinate towards junction with bursa. Genital open-
ing unsclerotized. Ductus seminalis from bursa near
its junction with ductus bursae.
Richards and Thomson (1932) referred Strymax
Dyar to Ephesiia as a subgenus. They did not treat
Unadilla. Hulst erected the latter for a single species
(nasutella Hulst) which Hampson placed with two other
Zeller species (ubacensis and maturella) in the synonymy
of erronella Zeller. Upon the basis of this synonymy,
which is doubtful, to say the least, Hampson cites
erronella as the type of Unadilla. Janse follows him
in this. Nomenclatorily nasutella must be the type of
Unadilla whether nasutella proves to be a synonym of
erronella or not.
Strymazx is an obvious synonym of Unadilla. It has
no close affinity to Ephestia even in the broad sense in
which that genus is interpreted by Richards and
Thomson. Hind wing venation and genitalia seem to
indicate much closer relation to Homoeosoma.
On structures of the male and female genitalia the
species divide into two distinct groups, as follows:
Male with apical process of gnathos partially fused; female
with an encircling band of signa.
Male with apical process of gnathos completely fused;
female without signa.
Genus Unadiila, Species 469: U. erronella
[Male with apical process of gnathos partially fused; female
with an encircling band of signa.]
469. Unadilla erronella (Zeller)
Fiaures 121, 498, 995
Homoeosoma erronella Zeller, Horae Soc. Ent. Rossicae, vol. 16,
p. 238, 1881.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Homoeosoma ubacensis Zeller, Horae Soc. Ent. Rossicae, vol. 16,
p. 239, 1881.
Unadilla erronella (Zeller) Hampson, in Ragonot, Monograph,
pt. 2, p. 262, 1901.
Ephestia bipunctella Hampson, Ann. Mag. Nat. Hist., ser. 7, vol.
7, p. 255, 1901 (new synonymy).
Strymazx dorae Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 344, 1914
(new synonymy).
Strymaz pyllis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 344, 1914
(new synonymy).
Ephestia (Strymazx) bipunctella (Hampson) Richards and Thom-
son, Trans. Ent. Soc. London, vol. 80, p. 201, 1932.
Ephestia (Strymaz) pyllis (Dyar) Richards and Thomson,
Trans. Ent. Soc. London, vol. 80, p. 202, 1932.
Hampson referred ubacensis as a synonym of erronella,
and, I believe, correctly so, judging from photographs
of the male types and their genitalia. The only differ-
ence between the two is in the more intensified macu-
lation of ubacensis. The same difference is exhibited
by the male types of dorae and pyllis, the latter being a
pale example with the normal subbasal markings of
forewing almost obsolete. However, in a series of
dorae from Panamé there are several intergrading
examples. U. erronella is also variable in size and to
some extent in structure, the vinculum being shorter in
some specimens from Puerto Rico than in those from
Panamé4 and Colombia, but otherwise the genitalia are
remarkably constant. In careful preparations of the
genitalia of Dyar’s supposed two species (dorae and
pyllis) I am unable to find any of the differences shown
in the figures published by Richards and Thomson.
The general color of the forewing is whitish gray; sub-
basal markings, when present, consisting of from one to
three pale brownish spots forming a broken antemedial
transverse shade; discal dots faint; hind wing white in
the male, more or less shaded with pale smoky fuscous
in the female. Alar expanse, 9-14 mm.
TYPE LOCALITIES: Honda, Colombia (erronel/a, in
BM); Ubaque, Colombia (ubacensis, in BM); Nassau,
Bahamas (bipunctella, in BM); La Chorrera, Panamd
(dorae, in USNM); Porto Bello, Panama (pyllis, in
USNM).
Foop piant: Unknown.
DistrisuTion: Conompia; Honda, Mariquita,
Ubaque. Panamd: Corazal (Mar., Apr., May), La
Chorrera (Apr., May), Paraiso (Apr.), Porto Bello
(Feb.), Rio Trinidad (Mar., May), Taboga Isl. (Feb.,
June). Banamas: Nassau. Pusrto Rico: Aguirre
Central (Apr.), Cataéno (Apr., June), Coamo Springs
(Apr.), Dorado (May, June), Isabela (Apr.), San
German (Aug.); Puerto Real (Vieques Isl., Apr., July).
Virein Isuanps: Kingshill (St. Croix, Mar., Dec.).
The Puerto Rican specimens (some 40-odd in the
Cornell University Collection) may possibly represent
a distinct race from the mainland form, but I am
unable to find any valid character for their separation.
The somewhat shorter vinculum of the male is matched
in some Panamé specimens and is of very doubtful
significance. U. erronella is easily identified as it is
the only described species with a partially divided
apical process of gnathos and a girdle of signa. The
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
remaining species (with the possible exception of
nasutella) belong in the following group.
Genus Unadilla, Species 470-472: U. maturella
to U. floridensis
[Male with apical process of gnathos completely fused; female
without signa.]
470. Unadilla maturella (Zeller)
Fieures 499, 996
Homoecosoma maturella Zeller, Horae Ent. Soc. Rossicae, vol. 16,
p. 240, 1881.
Hampson made maturella a synonym of erronella,
but a sketch by Clarke of the male genitalia of the male
type shows a completely fused apical process of gnathos
which places maturella in this group of species. A
photograph of the type also shows a pale grayish white
form with a strong, dark, transverse, antemedial shade
extending on a straight, outward slant from below costa
almost to inner margin. I have before me a series of
similarly marked specimens from Guatemala and Cuba
with genitalia similar to those of Zeller’s type, and
presumably conspecific. Alar expanse, 14-16 mm.
TypE Ltocatity: Colombia (type in BM).
Foop prant: Unknown.
Distrisution: Cotompia. GuaTEMALA: Cayuga
(Mar., Apr., May). Cusa: Baracoa (May), Havana.
471. Unadilla albidiorella (Richards and Thomson), new combination
Ephestia (Strymaz) albidiorella Richards and Thomson, Trans,
Ent. Soc. London, vol. 80, p. 201, 1932.
Described from a single male. From description and
figure published by the authors it must be very close to
maturella if not a race of that species. I have seen no
Peruvian examples of Unadilla. Alar expanse, 17 mm.
Typr tocatity: Rio Ucayale, Contamino, Pert
(“Oct.—Dec.,”’ type in BM).
Foop pirant: Unknown.
472. Unadilla floridensis, new species
Darker and more decidedly gray than the other
species of the genus; dark pattern markings faint, in
some specimens almost obsolete, in others conspicuous
only as a single antemedial shade or spot and a narrow
fainter dark inner shade along subterminal line of fore-
wing; white dusting rather sparse. Alar expanse,
8-13 mm.
The genitalia (co and 9) are similar to those of
maturella, exhibiting no differences of a specific nature.
Type Locatity: Key West, Fla. (type in USNM,
61377).
Foop piants: Pulchea odorata, Melanthera radiata.
Described from male type and 10 male and 4 female
paratypes from the type locality, reared Apr. 23 to May
1, 1945, by the Special Survey of the Division of Foreign
Plant Quarantine, U. S. Bureau of Entomology and
Plant Quarantine, from larvae feeding in the blossoms
229
and seed pods of Pulchea and Melanthera. These are
the only food-plant records we have for the genus
Unadilla. The larvae are similar in markings and
habitus to those of Rotruda mucidella and like them lack
the sclerotized rings about seta IIb of mesothorax and
seta IIT of the eighth abdominal segment.
Genus Unadilla, Species 473 (unplaced): U.
nasutella
473. Unadilla nasutella Hulst
Unadilla nasutella Hulst, Phycitidae of N. Amer., p. 197, 1890.
The male type in the Rutgers Collection lacks an
abdomen, so it will be impossible to place nasutella in
its proper species group or to determine its synonymy
until more specimens from the southwestern United
States are discovered. Except for the Hulst type I
have seen no example of Unadilla from New Mexico,
Arizona, or Texas. Alar expense, 13.5 mm.
Type tocauity: Hot Springs, N. Mex. (type in
AMNH, ex Rutgers).
Foop piant: Unknown.
Hampson referred nasutella as a synonym of erronella
Zeller and it so stands in our lists. This synonymy is
probably incorrect and is certainly unwarranted on the
evidence before us.
Genera 131-151: Laetilia to Cactobrosis
{Venational division A. Forewing smooth, oblong, broadest
toward termen; color alike in both sexes; 11 veins; 10 from cell,
8 and 9 stalked, 6 straight, 4 and 5 stalked, 2 and 3 separate or
approximate, 2 from near lower outer angle of cell; no costal
fold or other secondary sexual modifications. Hind wing with
7 veins; 3 and 5 connate or stalked; 2 from before lower outer
angle of cell; discocellular vein curved, complete; no sex-scaling
or other sexual modifications; cell one-half or somewhat less
than one-half the length of the wing. Abdomen of male with a
pair (rarely two pairs) of ventrolateral hair tufts at base of
eighth segment or with eighth segment simple. Male genitalia
with uncus broad, subtriangular, never hook-shaped or otherwise
modified, apex broadly rounded, its outer (dorsal) surface densely
covered with bristlelike scales; gnathos terminating in a flanged
(or lobed) and hooked apical process, frequently bifid, sometimes
fused or partially fused; transtilla incomplete (represented by 2a
pair of separate, more or less elongate-triangulate plates), never
forming a bridge or otherwise modified; harpe simple, without
clasper or extensions from sacculus or costa; anellus U-shaped;
aedeagus straight or slightly sinuate, usually smooth, but occa-
sionally with a few very small scobinations at apex; penis smooth,
finely scobinate or with sclerotized wrinklings but not other-
wise armed; vinculum stout and broad, short or long, and with
terminal margin normally broadly rounded. Female genitalia
with bursa copulatrix membranous, smooth or more or less mi-
nutely scobinate on inner surface, never strongly sclerotized or
pigmented; signum frequently absent, when present consisting
of a small, ribbed, weakly serrate, finely scobinate or cupped
plate; ductus bursae membranous throughout or scobinate for a
short distance from genital opening, gradually widening into and
not sharply differentiated from bursa copulatrix; genital opening
normally simple, sometimes minutely scobinate, occasionally
with sclerotized dorsal or ventral plates or a few setae on the inner
surfaces of the ductus at the opening, otherwise unmodified or
unarmed; ductus seminalis from bursa, rarely from ductus bursae
near junction with bursa.]
230
131. Genus Laetilia Ragonot
Laetitia Ragonot, Entomologica Americana, vol. 5, p. 116, 1889;
Bull. Soc. Ent. France, p. viii, January 1890.—Hulst,
Phycitidae of N. Amer., p. 182, 1890.—Hampson, in Rago-
not, Monograph, pt. 2, pp. 116, 560, 1901.—Forbes, Cornell
Mem. 68, p. 631, 1920. (Type of genus: Dakruma cocci-
divora Comstock.)
Laosticha Hulst, U. S. Nat. Mus. Bull. 52, p. 431, 1902.—Dyar,
Proc. Ent. Soc. Washington, vol. 6, p. 159, 1904. (Type of
genus: Dakruma ephestiella Ragonot.)
Tongue short (greatly reduced in melanostathma and
glomis, but not completely enclosed and hidden by the
labial palpi). Antenna pubescent, shaft simple in both
sexes. Labial palpus oblique or obliquely upturned
(males of melanostathma and zamacrella). Maxillary
palpus filiform or subsquamous (coccidivora). Hind
wing with veins 7 and 8 anastomosing for at least half
their distance beyond cell; 3 and 5 stalked. Highth
abdominal segment of male simple.
Male genitalia with apex of uncus normally broadly
rounded (somewhat narrowly rounded in coccidivora,
cardini, and myersella) ; apical flanged process of gnathos
stout, the lobes fusing posteriorly; aedeagus smooth,
moderately stout; penis without cornuti, rarely with
a few sclerotized folds (zamacrella, myersella), otherwise
smooth; vinculum stout, rarely longer than broad
(jiskella).
Female genitalia with signum consisting of a narrow,
flanged or a small rounded, bluntly dentate plate; bursa
copulatrix, except for signum and occasional minute
scobinations surrounding the signum, smooth; ductus
bursae with a sclerotized ventral plate and a pair of
dorsal sclerotized plates at genital opening, otherwise
smooth; ductus seminalis from anterior end of bursa
copulatrix.
Larvae predaceous on scale insects.
The genus as here defined is structurally a somewhat
composite group but is, I think, a natural one, linking
in one direction with Rhagaea, Zophodia, and the cactus-
feeding phycitine genera and on another with two or
three as yet undefined genera of coccid feeders in the
Anerastiinae. Typical ZLaetilia (coccidivora to portori-
censis) are as much an erastiine as phycitine on male
genitalic characters (especially the general habitus of
the organs as well as the peculiar development of the
gnathos), but the tongue, while much reduced, is dis-
tinetly exposed as in most other short-tongued Phyci-
tinae. Three of the transition species (melanostathma,
amphimetra, and glomis) have the tongue rudimentary
and on this structure could go into the Anerastiinae,
were it not for their genitalia. The remaining transi-
tion species (zamacrella, myersella, ephestiella and jfis-
kella), all nearctic species, are definitely phycitine on
all characters. On the sum of its characters Laetillia
must be included in the Phycitinae. Upon one or more
structural differences it can be distinguished from any
other phycitine or anerastiine genus.
474. Laetilia coccidivora (Comstock)
Fiaures 63, 502, 503, 997
Dakruma coccidivora Comstock, North Amer. Ent., vol. 1, p. 26,
1879; Rep. Comm. Agr. for 1879, p. 241, 1880.—Packard,
UNITED STATES NATIONAL MUSEUM BULLETIN 207
U. 8. Ent. Comm. Bull. 7, p. 54, 1881; U.S. Dep. Agr. Fifth
Rep. Ent. Comm., p. 413, 1890.
Dakrum pallida Comstock, Rep. Comm. Agr. for 1879, p. 243,
1880.
Laetilia coccidivora (Comstock) Ragonot, Ent. Amer., vol. 5, p.
116, 1889; Bull. Soc. Ent. France, p. viii, 1890.—Hulst,
Phycitidae of N. Amer., p. 182, 1890; U. S. Nat. Mus. Bull.
52, p. 431, 1903.—Hampson, in Ragonot, Monograph, pt. 2,
p. 117, 1901.—Hunter, Pratt, and Mitchell, U. S. Dept.
Agr. Bur. Ent. Bull. 113, pp. 24, 26, 1912.—Forbes, Cornell
Mem. 68, p.632, 1920.—McDunnough, Check list, No. 6328,
1939.
Zophodia dilatifasciella Ragonot, N. Amer. Phycitidae, p. 13,
1887.—Hulst, Phycitidae of N. Amer., p. 174, 1890; U. S.
Nat. Mus. Bull. 52, p. 429, 1903.—Hampson, im Ragonot,
Monograph, pt. 2, p. 21, 1901.—Hunter, Pratt, and Mitchell,
U.S. Dep. Agr. Bur. Ent. Bull. 113, pp. 24, 26, 1912.—Mc-
Dunnough, Check list, No. 6309, 1939. (New synonymy.)
Laetilia coccidivora hulstit Cockerell, Amer. Nat., vol. 31, p. 588,
1897.—Hulst, U. S. Nat. Mus. Bull. 52, p. 431, 1903.—
MeDunnough, Check list, No. 6328a, 19389.
Laetilia hulstii (Cockerell) Dyar, Proc. Ent. Soc. Washington,
vol. 6, p. 228, 1904.
Palpi, face, head, and thorax brownish fuscous dusted
with white, the white more pronounced on undersurface
of second segment of labial palpus, on head, and on
collar. Forewing pale brownish fuscous heavily dusted
with white and with dark brown bordering the trans-
verse lines; the white dusting concentrated on the costal
half of wing, giving this area (especially between cell
and costa) a distinctly whitish appearance; area between
cell and inner margin (and, beyond cell, between vein
3 and tornal margin) more or less suffused with pale
ocherous fuscous; costa for a short distance from base
blackish brown; in most specimens a rather obscure
transverse pale brownish basal band extending from
just below base of costa to inner margin near antemedial
line (only observable on unfaded specimens, sometimes
obsolete); from middle of basal band a short blackish
streak; antemedial line slanting outwardly from costa,
slightly angled or bent at fold, white and margined in-
wardly by a very narrow blackish brown band which
fades out towards costa, and outwardly margined by a
broad blackish brown band, especially contrasted to-
wards costa; subterminal line, faint, smuate (inwardly
angled opposite cell and also, but very slightly, at fold),
inwardly margined by a narrow blackish brown band
and outwardly edged for a short distance from costa
by a broader, more or less triangular, blackish shading;
a small blackish dot or line at upper angle of cell and
another at lower outer angle, these occasionally fused
but normally well separated; along termen at the vem
ends a row of obscure brownish dots; cilia ashy gray.
Hind wing whitish with a somewhat smoky tint, a dark
shading along costa, a narrow dark line along termen
and some dark shading on the veins; cilia white with a
broad, slightly darker, basal band. Alar expanse,
10-20 mm.
Male genitalia with gnathos terminating in a flanged
process with drooping lobes and a pair of short, strongly
sclerotized, divergent posterior horns; apex of uncus
truncate; penis simple, without cornuti or sclerotized
folds.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 231
Female genitalia with signum rather large, a single,
elongate, strongly sclerotized, pocketlike projection into
bursa; bursa copulatrix, except for signum, smooth.
Type Locauities: Washington, D. C. (coccidivora, in
USNM); Sanford, Fla. (pallida, in USNM); Las Cruces,
N. Mex. (hulstti, in AMNH, ex Rutgers); Arizona
(dilatifacsiella, in Paris Mus.).
Foop: Various Coccidae.'2 Specific records include
Pulvinaria vitis (L.) (originally identified as Lecanium
acericola or Pulvinaria innumerabilis), Pulvinaria amyg-
dali Ck11., Pulvinaria bigeloviae Ck11., Toumeyella
numismaticum (Ck11.), Toumeyella spp., Coccus hesperi-
dum L., Eriococcus quercus (Coms.), Saissetia oleae
(Bern.), Kermes spp., Lecaniodiaspis sp., Cerococcus
quercus Coms., Lecanium arizonensis King, Dactylopius
sp., Dactylopius confusus Ck11., Neolecanium cornu-
parvum (Tho.), Dactylopius tormentosus (Lam.), Pseu-
dococcus sp. The larvae will also feed in flowers of
cactus (Platypuntia spp.)
Distrisution: District of Columbia (July); Virginia,
Falls Church (Aug.), Vienna (July); Maryland, Plum-
mers Isl. (June); Pennsylvania, Longhorne (Oct.),
Rockville (Apr.); Ohio, Newark (Nov.); Florida, no
specific locality (May), Everglades (Apr.), Chokoloskee,
Miami (Feb.); Alabama, Mobile (June); Mississippi,
Biloxi (June); Louisiana, Baton Rouge (Aug., Nov.),
New Orleans; Texas, Sabinal (Mar.), Victoria (Feb.),
Brownsville (Jan., Feb., Apr.), Kerrville (Apr., May),
San Antonio (Oct.), Uvalde (May, Aug.), Sterling City
(Oct.), Dallas (Nov.), Corpus Christi (Feb.); New
Mexico, Mesilla Park (June), Hot Springs; Arizona,
Tucson (June, Nov.), Paradise (Mar., Sept., Oct.),
Redington, Palmerlee, Huachuca Mts., Wilgus Mts.,
Baboquivari Mts. (July, Sept., Oct.); California, San
Bernardino (Nov.), San Diego (Oct.), Palm Springs,
no locality (Mar.).
There is considerable variation in size in this species
and corresponding differences in the shape of the
flanged processes of the gnathos and the length of the
lateral arms of anellus, but nothing that would justify
separation into races; for the extremes of difference in
size and structure are present in the reared cotype series
from Washington, D. C. The name dilatifasciella ap-
plies to Arizona specimens which average larger than
those from eastern localities and show some occasional
traces of ocherous shading on the lower half of forewing.
Here also there is no consistency which would justify
even a racial separation. The name has escaped
synonymy so far only because specimens identified to
it have been placed in the wrong genus. The larvae of
coccidivora like those of other species of Laetilia are
predaceous on the Coccidae. For obvious reasons they
do not attack the smaller hard-scaled species but other-
wise are not restricted, feeding on any of the larger
scales and mealy bugs that occur in colonies of sufficient
number to provide food. They seem to be fond of the
122 Dr. Harold Morrison was good enough to check the coccid
records in connection with rearings of L. coccidivora and has
supplied the names of coccids here given.
cochineal scale on cactus and on this plant will occa-
sionally vary their diet by feeding upon the flowers.
Several moths have been reared from larvae which fed
altogether upon the flowers of Platypuntia. As a conse-
quence coccidivora has been listed as a cactus insect; but
this plant feeding is, I believe, only a departure from
the normal predaceous habit. Certainly coccidivora
does not favor the coccids on cactus above those on
other plants. Its impartiality in diet and lack of host
restriction are probably accountable for its ineffective-
ness as an agent of control in scale infestations.
475. Laetilia coccidivora quadricolorella (Dyar), new combination
Fiaures 504, 999
Atascosa quadricolorella Dyar, Proc. Ent. Soc, Washington, vol. 6,
p. 116, 1904.
Poujadia quadricolorella (Dyar), Barnes and McDunnough,
Check list of the Lepidoptera of Boreal America, no. 5819,
1917.—MecDunnough, Check list, No. 6422, 1939.
Differs from that of typical coccidivora in that the
white dusting is more abundant on head, thorax, and
forewing, and the brownish fuscous shading largely re-
placed by ocherous; the basal patch ocherous with a
narrow outer border of blackish scales; band outwardly
bordering the antemedial line ocherous with a blackish
brown dash or narrow smudge at top of cell; area
between cell and inner margin and between antemedial
and subterminal lines more or less ocherous (in Cali-
fornia specimens before me completely so) ; subterminal
line narrowly bordered inwardly and outwardly by
blackish scales, especially towards costa; black scales
more or less lining the upper margin of cell; blackish
discal dots at end of cell and along termen at vein ends
rather sharply contrasted against the white dusting of
the ground color. Hind wing clear white with no dark
scaling on the veins; along termen a narrow brown line.
Alar expanse, 15-20 mm.
Male genitalia with apex of gnathos more rounded,
posterior horns of apical process of gnathos somewhat
shorter and closer together than those of typical
coccidivora; but otherwise substantially the same.
Female genitalia not essentially different from those of
typical coccidivora.
Type tocauiry: Las Cruces, N. Mex. (type in
USNM).
Foop: Unknown (none of the specimens before me
has been reared but the larvae presumably feed upon
more than one species of coccids).
Distrisution: New Mezico, Las Cruces, Fort Win-
gate (May); Arizona, Redington; California, Mojave
(Aug.), Olancha (May, June).
This local color variety (it is hardly more than that)
has probably escaped synonymy because it was de-
scribed in the wrong genus and subfamily. When it is
reared we may discover that it has no right even to
racial distinction. In New Mexico and Arizona it and
typical coccidivora occur in the same localities. The
California specimens before me, however, show such
marked contrasts in color (intensity of the white dusting
and light ocherous tint of the normally brownish areas
232
of forewing) that I hesitate to sink the name completely.
It is possible that hulstii Cockerell and quadricolorella
stand for the same form (types of both are from Las
Cruces and have clear white hind wings); but the type
of the former (9) is unrecognizable and the name may
very well be left where it is.
Dyar’s type is a male and not a female as he stated
in his description.
476. Laetilia coccidivora cardini Dyar
Fiaures 505, 998
Laetilia cardini Dyar, Ins. Inse, Menstr., vol. 6, p. 139, 1918.
Not distinguishable from small specimens of typical
coccidivora except that the hind wing is somewhat
darker. In addition to the female type I have before
me another female from the type locality and three
males and eight females from Orlando, Fla. The
Florida specimens were reared from larvae feeding on
mealy bugs on grapefruit.
The female genitalia differ only in their smaller size
from genitalia of normal coccidivora, but even in this
detail do not differ from equally small examples of
coccidwora. The male genitalia show apex of uncus
rather more pointed and the posterior horns of apical
process of gnathos more widely spaced than those of
typical coccidivora. However there is considerable
variability in these structures in coccidivora and the
differences exhibited by the male of cardini may not be
constant. In my opinion they do not justify more than
racial separaton.
TypE LocaLity: Santiago de las Vegas, Cuba (type
in USNM).
Foop: Mealy bugs.
Distrizution: Cusa: Havana, Santiago de las
Vegas. Unrtep Statss: Florida, Orlando (June).
477. Laetilia obscura Dyar
Ficures 506, 1000
Laetilia obscura Dyar, Ins. Insc. Menstr., vol. 6, p. 140, 1918.
Brownish fuscous with a sparse scattering of whitish
scales on head, thorax, and subcostal area of forewing.
Usual transverse markings on forewing nearly obsolete;
antemedial line very faint, straight, shaded outwardly
towards costa by dark brown; subterminal line a mere
trace; discal dot obscure; cilia very little paler than
ground color. Hind wing pale smoky fuscous, darkened
outwardly and with blackish brown terminal margin;
cilia slightly paler with dark basal band. Alar expanse,
9-11 mm.
Male genitalia similar to those of coccidivora cardini
but with posterior horns of gnathos more widely
spaced. Female genitalia with signum similar to but
weaker and narrower than that of coccidwora or any of
its varieties.
Typ Locauity: Santiago de las Vegas, Cuba (type
in USNM).
Foop: Presumably Coccidae (type series apparently
reared but without labels to indicate food or dates of
emergence).
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Distrizution: Cusa.
Represented in the National Collection only by
specimens from the type locality. It may prove to be
merely a dark, suffused race of coccidivora; but the
uniformly dark color, the almost complete obliteration
of the usual transverse markings, and the much narrower
signum suggest a distinct species.
478. Laetilia portoricensis Dyar
Fiaures 500, 1001
Laetilia portoricensis Dyar, Ins. Insc. Menstr., vol. 3, p. 62,
1915.—Wolcott, Journ. Agr. Univ. Puerto Rico, vol. 20,
No. 1, p. 479, 1936.
Palpi, face, head, and thorax pale brownish fuscous
with a few white scales on inner side of palpus and on
face. Forewing pale brownish fuscous with area
between costa and upper vein of cell and antemedial
and subterminal lines white; on this white area a
rather conspicuous midcostal, blackish brown dash;
basal area paler than ground color, whitish toward
costa; transverse pale lines obscure, indicated chiefly
by their dark outer borders; antemedial line slightly
angled at cell, bordered outwardly by blackish brown
which is conspicuous on costa and forms a patch at the
cell; subterminal line outcurved between veins 2 and 5,
rather broadly bordered with blackish brown towards
costa; a blackish brown discal dot at upper outer angle
of cell; more or less blackish shading along submedian
fold and upper vein of cell; a few indistinct dark spots
on pale termen between the veins; cilia concolorous
with pale terminal margin. Hind wing pale smoky
brown; terminal margin dark brown; cilia sordid
whitish with dark basal band. Alar expanse, 10-13
mm.
Male genitalia with gnathos terminating in a, pair of
rather weakly sclerotized, liplike lobes without posterior
horns; uncus rather broad but laterally folded, apical
margin rounded; penis without cornuti. Female geni-
talia with signum a rather small, nearly round, sclero-
tized, scobinate disk; bursa copulatrix, except for
signum and area immediately adjoining it, smooth;
vinculum long.
Typ Locality: Rio Piedras, Puerto Rico (type in
USNM).
Foon: Saissetia oleae, Lecanium sp.
Distrisution: Purrto Rico: Rio Piedras (Nov.),
Bayamén (Aug.), Comerio (Aug.).
The genitalia and the midcostal dark dash on fore-
wing readily distinguish this species from anything else
in the genus. Dyar described it as “gray,” but the
ground color and general effect, except for the contrasted
whitish subcostal area, is more brown than gray.
479. Laetilia melanostathma (Meyrick), new combination
Fieures 501, 507, 1002
Euzophera melanostathma Meyrick, Exotic Microlepitoptera, vol.
5, pt. 5, p. 134, 19387,
Palpi, head, and thorax pale grayish brown. Fore-
wing light gray with a whitish suffusion in subcostal
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
area and a faint lilac tint due to sparsely scattered,
reddish brown scales; antemedial line very faintly in-
dicated, straight, slanting inwardly from costa, and
bordered outwardly in cell by a blackish blotch, this
black marking not continued to costa; at outer end of
cell a similar conspicuous, large, round black spot; a
faint dark streak on midcosta; subterminal line faint,
sinuous, bordered outwardly and inwardly by blackish
bands, the latter rather broad and well defined from
costa but fading out toward inner margin; a few irregu-
lar black dots on termen between the veins; cilia
slightly paler than ground color of the wing. Hind
wing very pale smoky fuscous, translucent and almost
white towards base, somewhat darker in the female;
terminal margins dark; cilia slightly paler than wing.
Alar expanse, 11-12 mm.
Male genitalia with gnathos terminating in a broad
heart-shaped process composed of two partially cleft,
partially fused lobes ending in a single, slender posterior
horn; vinculum rather long and broad; penis without
cornuti.
Female genitalia with ductus bursae strongly and
broadly sclerotized on ventral side toward genital open-
ing; bursa copulatrix minutely scobinate; signum
similar to that of coccidivora, but smaller.
Type Locatiry: Concordia, Entre Rios, Argentina
(type in BM).
Foon: Tachardiella argentina.
As yet known only from the type locality.
The four specimens examined had been reared (Nov.
1936) and were submitted by K. J. Hayward, and pre-
sumably formed part of the original series from which
the type and paratypes were selected.
This species is a somewhat anomalous Laetilia. The
very short tongue, short and acuminate third segment
of labial palpus, and definitely filiform maxillary palpus
suggest other generic placement; but the genitalia show
that it is closely allied to typical Laetilia. It certainly
does not belong in Euzophera where it was originally
placed. The two conspicuous black spots on the cell
of forewing will identify it specifically, and distinguish
it from other described species of the genus with the
possible exception of amphimetra.
480. Laetilia amphimetra (Meyrick), new combination
Euzophera amphimetra Meyrick, Trans. Ent. Soc. London, vol.
89, pt. 4, p. 47, 1939.
This species is known to me only from the description.
In the National Collection there are five badly rubbed
specimens (one o' and four 9) reared from larvae
feeding on a Ceroplastes sp. at Valemar, Chile, by
P. A. Barry (“9-21-40”) which might be amphimetra.
What is left of the pattern matches well enough with
Meyrick’s description. The markings are similar to
those of melanostathma and the male genitalia are
identical, except that the vinculum is shorter in the
Chilean specimen. It and its accompanying females,
however, are appreciably darker than typical examples
of melanostathma. If the Chilean specimens are
233
amphimetra, the latter is probably no more than a color
variety of melanostathma.
Typr tocauity: Concordia, Entre Rios, Argentina
(type in BM).
Foon: Ceroplastes sp. on guava.
481. Laetilia zamacrella Dyar
Ficures 508, 1004
Laetilia zamacrella Dyar, Ins. Insc. Menstr., vol. 13, p. 12,
1925.—McDunnough, Check list, No. 6327, 1939.
Palpi, face, head, thorax, and forewing grayish
fuscous rather evenly dusted with white (the head
densely so) making the general color to the naked eye a
pale slate gray. Forewing blackish at base for a very
short distance; antemedial line nearly vertical, slightly
indented at fold, white, bordered within and without by
blackish brown, outer dark border appreciably broad-
ened at top of cell; a conspicuous blackish discal spot
at outer margin of cell; subterminal line sinuate (out-
wardly angled between veins 1 and 5), white, narrowly
bordered inwardly and outwardly by blackish fuscous;
along termen a row of more or less fused blackish spots;
cilia ashy gray. Hind wing semihyaline with a pale
fuscous shade bordering costa and a fine fuscous line
along termen from apex to vein 1b; veins slightly darker
than wing membrane; cilia white. Alar expanse, 24-25
mm.
Male genitalia with apical process of gnathos par-
tially fused, large, the lobes broadly flaring, the posterior
projections stubby and close together; penis with
several long sclerotized folds but without cornuti.
Female genitalia with a pair of strongly sclerotized
dorsal plates in ductus bursae at genital opening;
ventral plate at opening large; signum a small flanged
plate; bursa copulatrix smooth except for a few minute
scobinations surrounding signum.
Typrgz xocauity: Mount Wilson, Calif.
USNM).
Foon: Scales on pine.
Distrisution: California, Mount Wilson (July),
Mount Hamilton (Apr., May).
The seven Mount Hamilton specimens were reared
by H. E. Burke, under Hopkins No. 19013d, from
larvae feeding on an unidentified scale on Pinus radiata.
The moth superficially resembles Dioryctria abietella
and might easily be mistaken for a Dioryctria species
except for its reduced hind wing venation and the
black rather than white discal spot on forewing. Its
genitalia, except for their squat appearance, resemble
those of Zophodia and Cactobrosis and show the close
affinity of Laetilia to the cactus-feeding phycitine
genera,
(type in
482, Laetilia myersella Dyar
Fiaures 509, 1003
Laetilia myersella Dyar, Proc. Ent. Soc. Washington, vol. 12,
p. 54, 1910.—Forbes, Cornell Mem. 68, p. 632, 1920.—
McDunnough, Check list, No. 6330, 1939.
Palpi, face, head, and thorax brownish fuscous sparse-
ly dusted with white, the white dusting most conspic-
234
uous on the middle third of tegulae, throwing into con-
trast their dark basal and even darker apical areas.
Forewing ashy white with brownish areas forming a
rather strongly contrasted pattern; basal area brownish
fuscous with a narrow pale subbasal border; antemedial
line well marked, somewhat sinuate and slanting out-
ward from costa to immer margin, white, with a narrow
dark inner border and a broad darker outer border
which is appreciably broadened towards inner margin;
discal dots fused into a large, dark spot at end of cell;
subterminal line slightly sinuate (outwardly angled be-
tween veins 1 and 5), white, rather broadly bordered
inwardly and outwardly by dark brownish fuscous;
terminal area of the pale ground color with terminal
margin dark brown; cilia pale brownish fuscous. Hind
wing pale fuscous with a dark line along termen; cilia
whitish. Alar expanse, 14-16 mm.
Male genitalia with apical process of gnathos partially
fused, large, the lobes flaring but not so broadly so as
those of zamacrella, the posterior projections sharp and
appreciably separated; penis with several long sclero-
tized folds but without cornuti (similar to that of
zamacrella) .
Female genitalia with a pair of strongly sclerotized
plates in ductus bursae at genital opening; ventral
plate at opening large; signum a small plate with from
two to four bladelike projections; bursa copulatrix
smooth except for a few minute scobinations surround-
ing signum.
Typn LocaLity: Rockville, Pa. (type in USNM).
Foon: Scales on pine.
Distrisution: Pennsylvania, Rockville (May) ; North
Carolina, Tryon, Southern Pines (Sept.).
Twelve specimens examined.
This species is apparently confined to the Eastern
States. It is readily separable on habitus and structure
from the western zamacrella but is obviously closely
related to that species.
483. Laetilia ephestiella (Ragonot)
FicureE 510
Dakruma ephestiella Ragonot, N. Amer. Phycitidae, p. 13, 1887.
Laetilia ephestiella (Ragonot) Hulst, Phycitidae of N. Amer.,
p. 185, 1890.—Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 159, 1904; vol. 7, p. 37, 1905.—McDunnough, Check list,
No. 6326, 1939.
Lasiosticha ephestiella (Ragonot), Monograph, pt. 2, p. 109, 1901.
Laosticha ephestiella (Ragonot) Hulst, U. S. Nat. Mus. Bull. 52,
p. 431, 1903.
Maricopa lustrella Dyar, Proc. Ent. Soc. Washington, vol. 5,
p. 227, 1903.
Ashy fuscous, resembling myersella very closely in
color and markings but with the transverse pale lines
and dark areas of forewing less strongly contrasted;
antemedial and subterminal lines faint but distinguish-
able and otherwise as on myersella, their dark borders
a paler brown, not so broad and more obscured than
those on myersella. Hind wing translucent, with a very
pale ocherous tint; termen hardly darker; cilia con-
colorous. Alar expanse, 17-20 mm.
Male genitalia with apical process of gnathos partially
UNITED STATES NATIONAL MUSEUM BULLETIN 207
fused, large, the lobes flaring and similar to those of
myersella, the posterior projections short, blunt and
close together as in zamacrella. Uncus narrower than
that of either myersella or zamacrella.
TYPE LOCALITIES: Arizona (ephestiella, in Paris Mus.) ;
Williams, Ariz. (June; lustrella, in USNM).
Foop: Unknown, presumably scales.
Distripurion: Arizona.
This species is known to me only from Dyar’s male
type. I have no reason to question the synonymy
which he proposed in 1905. The three species, zama-
crella, myersella, and ephestiella, can readily be separated
on color and genitalic characters but they are obviously
very closely related and form a distinct subgroup within
the genus.
484. Laetilia fiskella Dyar
Figures 511, 1005
Laetilia fiskella Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 221,
1904.—McDunnough, Check list, No. 6331, 1939.
Similar in color and markings to myersella except that
central area of forewing (between antemedial and sub-
terminal lines) is much darker, blackish brown; the basal
area is paler than the median area, but the entire wing
lacks the dusting of white so characteristic of myersella;
antemedial white line outwardly angulate at middle;
discal dot as in myersella, but sometimes obscured by
the general dark suffusion. Hind wing dark smoky
fuscous; cilia slightly paler. Alar expanse, 16-20 mm.
Male genitalia of the Baphala type; uncus with lateral
edges concave; apical process of gnathos consisting of a
pair of tear-shaped lobes fused at apex; vinculum long;
penis smooth. Female genitalia with pair of sclerotized
plates in ductus bursae at genital opening; ventral plate
at genital opening large; ductus bursae long and much
more slender than in myersella; siznum a small plate
armed with several irregularly shaped, thornlike projec-
tions; bursa copulatrix triangulate, smooth except for a
few minute scobinations surrounding signum.
Typr Locatity: Tryon, N. C. (type in USNM).
Foop: Unknown, presumably scale insects.
Known to me only from the type series, two males
and three females from the type locality (not one male
and four females as stated by Dyar). The species is a
transition one linking typical Laetilia and Baphala; it
has Baphala genitalia but lacks the other diagnostic
characters of that genus, the eighth abdominal segment
of the male being simple, the male antenna without
sinus or sex-scaling on the base of shaft, and veins 3 and
5 of hind wing strongly stalked.
485. Laetilia glomis (Dyar), new combination
FieurE 512
Euzophera glomis Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 335,
1914.
Clay colored, with extreme base of forewing blackish;
antemedial line slanting outwardly to inner margin and
with a slight notch at vein 1b, distinct and blackish;
subterminal line faint, narrow, subdenticulate, blackish,
bordered outwardly by 8 somewhat wider pale band;
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 235
two faint, dark discal dots at outer angles of cell; along
termen a row of faint blackish dots. Hind wing glossy
smoke brown; cilia very slightly paler, nearly concolor-
ous with wing. Alar expanse, 16 mm.
Male genitalia with lateral edges of uncus concave
(but not so deeply as in jiskella); apical process of
gnathos partially fused, the lobes small, rounded, the
posterior fused portion rather broad and forked at apex;
vinculum short (as wide as long); penis with a few
weakly sclerotized folds, otherwise unarmed.
Tyrer Locauity: Taboga Isl., Panama (Feb., type in
USNM).
Foop: Unknown, presumably scales.
Represented only by the unique type, a male (not
female, as stated by Dyar). The tongue is greatly re-
duced and less exposed by the labial palpi than that of
melanostathma Meyrick, indicating glomis as another
direct link between Laetilia and the coccid-feeding
Anerastiinae. Its genitalia, however, show that glomis
is better placed in the Phycitinae and in Laetilia than
with the coccid-feeding complex referred to the Aner-
astiinae.
Baphala, new genus
Type oF Genus: Huzophera homoeosomella Zeller.
Tongue well developed. Antenna pubescent; in male
some rough scaling in a very shallow sinus toward base
of shaft. Labial palpus obliquely upturned, reaching
to level of or slightly above vertex. Maxillary palpus
filiform. Hind wing with veins 7 and 8 anastomosing
for most of their length (free end of 8 very short and
weak); 3 and 5 closely approximate or connate at lower
angle of cell; cell less than one-half (little more than
one-third) the length of the wing. Abdomen of male
with a pair of ventrolateral hair tufts from eighth
segment.
Male genitalia with uncus narrowed at middle, the
lateral edges deeply concave, apex rounded; apical
process of gnathos a pair of moderately large lobes
fusing and hooked posteriorly; aedeagus smooth; penis
with weakly sclerotized wrinklings, otherwise unarmed;
vinculum stout, longer than broad.
Female genitalia with signum a small coarsely dentate
plate; bursa copulatrix more or less finely scobinate on
inner surface; ductus bursae with a sclerotized ventral
plate and internal scobinations at genital opening,
otherwise smooth; ductus seminalis from anterior end
of bursa.
Larvae predaceous on scale insects.
The genus is close to but distinct from Laetilia, easily
distinguished by its stronger tongue, the sex-scaling on
shaft of male antenna, the narrower median area of
uncus, the abdominal tufts on eighth segment of the male,
and the shorter cell and the approximate (rarely connate)
condition of veins 8 and 5of hind wing. Structural differ-
ences separating the species are slight and chiefly in the
size and shape of the terminal process of gnathos.
Differences in the female signa (usually of specific
value) are unreliable in Baphala, for they are as great
among individuals of a given species as they are be-
tween any two of the species themselves. The hind
wings are similarly colored and exhibit the same sexual
difference in the several species, whitish and semihyaline
in the male, pale smoky fuscous in the female.
486. Baphala basimaculatella (Ragonot), new combination
Figures 514, 1006
Vitula basimaculatella Ragonot, N. Amer. Phycitidae, p. 15,
1887.—Hampson, in Ragonot, Monograph, pt. 2, p. 82,
1901.—McDunnough, Check list, No. 6325, 1939.
Vitula basimaculella Hulst, Phycitidae of N. Amer., p. 179, 1890;
U.S. Nat. Mus. Bull. 52, p. 431, 1902 (misspelling).
Laetilia eremiella Dyar, Proc. Ent. Soc. Washington, vol. 12, p.
54, 1910.—McDunnough, Check list, No. 6329, 1939.
(New synonymy.)
Forewing pale whitish gray, very little darker along
inner margin; discal dots and transverse dark markings
blackish and strongly contrasted especially toward
costa; the discal dots normally separated but occasion-
ally partially fused. The palest species in the genus
and the only one known from the United States. The
male type in Paris lacks an abdomen but is obviously
what Dyar later described as eremiella. It was sent to
Ragonot by C. V. Riley and is labeled ‘‘Am. Sept.”
In the Stainton Collection in the British Museum are
two matching specimens from Texas. The male geni-
talic figure was drawn from one of these. The two
specimens in Paris from Loja, Ecuador, associated by
Hampson with basimaculatella are not that species. I
have examined the genitalia (male) of one of these Loja
specimens but am unable to match them with those of
any described species of Phycitinae known to me.
Alar expanse, 15-19 mm.
TYPE LOCALITIES: United States (basimaculatella, in
Paris Mus.); Stockton, Utah (eremiella, in USNM).
Foop: Unknown. Larvae presumably predaceous
on coccids.
Distrisution: Teras; Utah, Stockton (Aug.), Provo
(July, Aug.).
487. Baphala goyensis (Ragonot), new combination
Fieures 513, 1009
Zophodia goyensis Ragonot, Nouv. Gen., p. 31, 1888; Monograph,
pt. 2, p. 25, 1901.
Extreme base of forewing blackish fuscous; contrast-
ingly whitish beyond to the broad blackish fuscous
antemedial band; latter somewhat irregular, more or
less angulate; ground color of wing between lower
margin of cell and costa and between antemedial and
subterminal transverse markings, whitish; remainder
of wing shading to smoky fuscous with only a faint
dusting of white at apex; dark border of subter-
minal line well contrasted especially towards costa;
midcosta contrastingly edged with blackish fuscous; a
single rather large discal spot at end of cell (formed by
the complete fusion of the ordinary pair of discal dots).
Easily identified by the conspicuous discal spot and
other blackish fuscous markings. Alar expanse 18-20
mm.
236
Typr Locauity: Goya, Argentina (type in Paris
Mus:).
Foop: Black and wax scales (Ceroplastes and Saissetia
spp.).
Distrisution: Southeastern Brazil, Uruguay, Ar-
gentina. Six specimens are before me: A male collected
by Schaus at Castro, Parand, Brazil; another Brazilian
male without locality label, reared from a larva feeding
on a Ceroplastes and received from Dr. Costa Lima; and
four females reared at the South American Parasite
Laboratory of the U. S. Bureau of Entomology and
Plant Quarantine, at Montevideo, Uruguay, from
larvae feeding on black and wax scales.
488. Baphala goyensis olivacea, new race
Ficure 1010
Similar in color and markings to goyensis except that
the thorax and the dark markings on forewing are
olivaceous rather than blackish fuscous, and the discal
dots at end of cell are usually separated and not fused
into a single conspicuous dark spot. Alar expanse,
18-20 mm.
The genitalia show no characters of specific signifi-
cance to distinguish them from those of typical goyensis.
Typr Locatity: Posadas, Argentina (type in USNM,
61378; paratypes in Paris Mus. and BM).
Foon: Scale insects (not identified).
Described from male type and two male and six
female paratypes from type locality, reared (Dec., Jan.)
under No. 578-7, at the South American Parasite
Laboratory of the U. 8. Bureau of Entomology and
Plant Quarantine, at Montevideo, Uruguay, from larvae
predaceous on scales; and one female paratype from
Metaén, Argentina (Apr.). This last is from the col-
lection of the British Museum and had been identified
as squalida Walker. Judging from a photograph of the
type of Walker’s species I do not think it could possibly
be that species. Even in its somewhat faded condition
the olivaceous color of the thorax and dark forewing
markings are distinctly discernible. If this color had
been present in the type of squalida, Walker or Ragonot
would certainly have mentioned it.
489. Baphala homoeosomella (Zeller), new combination
Figures 517, 1011
Euzophera homocosomella Zeller, Horae Soc. Ent. Rossicae, vol.
16, p. 231, 1881—Hampson, in Ragonot, Monograph, pt. 2,
p. 61, 1901.
Vitula bodkint Dyar, Ins. Insc. Menstr., vol. 1, p. 99, 1913; Proc.
U.S. Nat. Mus., vol. 47, p. 339, 1914 (mew synonymy).
Vitula rusto Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 338, 1914
(new synonymy).
Vitula taboga Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 339, 1914
(mew synonymy).
Vitula saissetiae Dyar, Proc. Ent. Soc. Washington, vol. 31, p.
16, 1929 (new synonymy).
A pale brownish gray species, averaging smaller than
either basimaculatella or goyensis and with the trans-
verse dark markings on forewing weakly contrasted,
much obscured in some specimens; discal dots when
distinguishable small, not fused; the whitish dusting
UNITED STATES NATIONAL MUSEUM BULLETIN 207
limited chiefly to a shade along costa and some pale
dusting in basal area. Alar expanse, 11-16 mm.
There are no male genitalic structural characters to
separate any of Dyar’s supposed species from typical
homoeosomella and only trifling and inconsistent color
differences to distinguish them from each other. The
females show considerable variation in their signa, but
as much or more among specimens of any one of the
forms as between typical females of the forms them-
selves. The greatest variation is shown among speci-
mens from a reared series of typical saissetiae (figs.
1011b-4).
Tyre LocaLities: Honda, Colombia (homoeosomella,
in BM); Georgetown, British Guiana (bodkini, in
USNM); Parafso (rusto, in USNM), Taboga Isl. (taboga,
in USNM), Barro Colorado Isl. (saissetiae, in USNM),
all in Canal Zone, Panamé.
Foon: Saissetia spp. and Ceroplastes spp. (wax and
black scales), Toumayella sp.
Distrizution: Cupa. VireGin Istanps: St. Croix
(July). PanamA Canat Zone: Barro Colorado Isl.
(May), Paraiso (May), Taboga Isl. (Feb.), Rio Trinidad
(May). Brivise Guiana: Georgetown (June). Co-
LomBIA: Honda (Apr., Aug., Sept.). Brazin: Sado
Paulo (Feb.).
The most widely distributed and apparently the
commonest phycitine predaceous on scales in tropical
America.
490. Baphala haywardi, new species
Fieures 516, 1007
In color and markings similar to the more strongly
marked color forms of homoeosomella, the female hardly
distinguishable from the female paratype of saissetiae;
basal area of forewing, midcostal area above cell, and
terminal area beyond subterminal line rather strongly
dusted with whitish scales; dark transverse antemedial
band pale grayish brown, outwardly angulate and
diffusing into a similar shade through most of area be-
tween inner margin and cell; smuate subterminal white
line bordered inwardly and outwardly by pale grayish
brown; discal dots at.end of cell similarly colored; a row
of somewhat darker dots along termen. Alar expanse,
15-16 mm.
Male genitalia with vinculum shorter and apical
process of gnathos stouter than those of homocosomella
or any of the other described species of the genus.
Female genitalia with fine scobinations of bursa dis-
tributed over most of inner surface; sclerotization of
ductus bursae at genital opening weaker than that of
homoeosomella (compare figs. 1011h and 1007).
Typr Locauity: Concordia, Entre Rios, Argentina
(type in USNM, 61879).
Foon: Ceroplastes grandis.
Described from male type and female paratype from
type locality, reared by K. J. Hayward from larvae
feeding on the large wax scale (co, ‘9—-4-1935,” 9,
“14-VIT-1935,” Hayward No. 3185). They were sub-
mitted as examples of homoeosomella, which they could
easily be on color and markings; but their genitalia,
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
especially those of the male, seem to rule them out
from that species.
491. Baphala glabrella (Dyar), new combination
Fiaurss 515, 1008
Euzophera glabrella Dyar, Ins. Insc. Menstr., vol. 7, p. 57, 1919.
A suffused, grayish brown species with markings ob-
scure, the pale transverse antemedial and subterminal
lines and the dark discal dots faintly indicated; general
color darker (more brownish) than that of homoeo-
somella. Probably only a variety of that species, but
a reared series would be needed to determine this. The
genitalia show some trifling differences from homoco-
somella, but none that could be classed as specific.
Alar expanse, 15-16 mm.
Type Locauity: Cayuga, Guatemala (Apr., May,
June; type in USNM).
Foop: Unknown.
The species is known only from the original series of
five specimens upon which the name was based.
492. Baphala squalida (Walker), new combination
Nephopteryz squalida Walker, List, pt. 27, p. 59, 1863.
Zophodia squalida (Walker) Ragonot, Monograph, pt. 2, p. 25,
1901.
I know this species only from photographs of the male
type and its genitalia submitted by Tams. The photo-
graph of the moth shows a specimen almost completely
suffused with blackish fuscous except for a small area
from outer part of cell to costa. The subterminal pale
line of forewing, the two discal dots, and the row of
terminal dots are distinct, but the antemedial line and
the usually contrasted dark transverse markings are
indistinguishable in the general dark suffusion. The
hind wing shows the veins strongly outlined by dark
scaling and the apical area of the wing darkly clouded
(unusual markings for a male of this genus). Alar ex-
panse, 20 mm.
Typr Locauity: Rio de Janeiro, Brazil (type in BM).
Foop: Unknown.
Certain specific application of Walker’s name will
have to wait upon recovery of more material in reared
series from the type locality. If, as I suspect, a series of
rather dark homoeosomelia in the United States National
Museum from Sao Paulo (four males and eight females
reared from larvae on wax scales) should prove to be
squalida, Walker’s name will take precedence and
homoeosomella will fall as a synonym. However, none
of our Brazilian specimens is over 16 mm. in expanse
and their small size seems to rule them out.
133. Rhagea, new genus
Typ or GeNus: Zophodia packardella Ragonot.
Tongue well developed. Antenna pubescent, shaft
simple in both sexes. Labial palpus porrect. Maxil-
lary palpus filiform. Hind wing with veins 7 and 8
anastomosing for about half their distance beyond cell;
3 and 5 stalked. Eighth abdominal segment of male
simple.
237
Male genitalia with apex of uncus broad, apical
process of gnathos bifid, large; aedeagus moderately
long and stout; penis with some very weakly sclerotized
scobinations or pubescence but without cornutus.
Female genitalia without signum; bursa copulatrix
smooth or (stigmella) minutely scobinate-granulate;
ductus bursae with a weak bandlike sclerotized ventral
plate or scobinate patch on inner surface at genital
opening, otherwise membranous; ductus seminalis from
anterior end of bursa.
Larvae feeding in roots of Crassulaceae or flowers of
Orobanchaceae.
This genus is close to Zophodia Hiibner. It differs in
having porrect labial palpi in both sexes, the eighth seg-
ment of male simple, the antennal shaft of the male
simple and the female bursa without trace of signum.
493. Rhagea packardella (Ragonot), new combination
Ficures 518, 1012
Zophodia packardella Ragonot, N. Amer. Phycitidae, p. 12,
1887.—Hulst, Phycitidae of N. Amer., p. 173, 1890; U. 8.
Nat. Mus. Bull. 52, p. 430, 1903—Hampson, in Ragonot,
Monograph, pt. 2, p. 22, 1901.—McDunnough, Check list,
No. 6305, 1939.
Zophodia orobanchella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 111, 1904—-McDunnough, Check list, No. 6307, 1939.
(New synonymy.)
Palpi, head, thorax, and forewings white dusted with
brown, giving the moth a pale ash gray appearance.
Forewing with costal area paler than remainder of
wing; no trace of any transverse antemedial line; dark
dustings outlining most of the veins and forming rather
conspicuous longitudinal lines through center of cell, on
basal third of lower vein of cell, and along the basal
third of the fold; a large conspicuous discal spot at end
of cell covering the entire crossvein between veins 3
and 8; subterminal line indicated by dark shading which
extends from costa near apex to about vein 2, indented
between veins 4 and 6 and distinct only towards costa;
terminal margin dark; cilia ash gray. Hind wing pale
smoky white, with veins and terminal area slightly
darker and a fine dark line along termen; cilia sordid
white with slightly darker basal band. Alar expanse
18.5-25 mm.
Male genitalia with vinculum shorter and divided
apical process of gnathos larger in proportion to re-
mainder of genitalia than those of stigmella. Female
genitalia with inner surface of ductus bursae finely
scobinate-granulate at genital opening; bursa copula-
trix minutely and sparsely scobinate.
Typr tocauities: California (packardella, in Paris
Mus.); Wawawai, Wash. (orobanchella, in USNM).
Foop piant: Orobanche ludoviciana Nutall (larvae
feeding in the flowers).
Distrisution: Washington, Wawawai (Aug.); Utah,
Provo (July), Eureka (July); California, Dixieland
(Apr.); Arizona, Phoenix (Mar.); Colorado, Fort Collins
(Sept.).
Dyar had his specimens of orobanchella under pack-
ardella in the National Collection but, to the best of my
knowledge, the synonymy had not been published.
238
The species is easily recognized by the large, dark,
discal spot on the forewing, the most conspicuous mark-
ing on the insect.
494. Rhagea stigmella (Dyar), new combination
Ficures 519, 520, 521, 1013, 1014
Zophodia stigmella Dyar, Pomona Coll. Journ. Ent., vol. 2, No. 4,
p. 378, 1910.—Barnes and McDunnough, Check list of the
Lepidoptera of Boreal America, No. 5716, 1917.
Yosemitia maculicula Dyar, Ins. Insc. Menstr., vol. 1, p. 34,
1918.—Barnes and McDunnough, Contributions, vol. 3,
p. 200, 1916.
Zophodia stigmella maculicula (Dyar), Barnes and McDunnough,
Check list of the Lepidoptera of Boreal America, No. 5716,
1917.
Eumysia stigmella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221,
1925.—McDunnough, Check list, No. 6300, 1939.
Eumysia maculiella Dyar, Ins. Insc. Menstr., vol. 13, p. 221,
1925 (misspelling for maculicula).
Eumysia stigmella maculicula (Dyar), McDunnough, Check list,
No. 6300, 1939.
Palpi, face, and head pale brown dusted with white,
the scales brown with their tips white. Thorax grayish
white shading to pale argillaceous. Forewing grayish
(ashy) white with blackish brown markings and a
rather broad pale argillaceous shading along the fold
and fainter traces of the same color between the veins
beyond cell; veins, except on lower half of wing, faintly
lined with blackish brown; antemedial line incomplete,
acutely angled, in most specimens indicated only by a
broad blackish spot on the basal third of inner margin
extending from inner margin to vein 1b, a rather con-
spicuous blackish patch on lower vein of cell just beyond
middle and an obscure blackish shade on basal third of
costa; subterminal line often obsolete, otherwise indi-
cated by a narrow blackish streak from costa to vein 8,
a more or less broken line between veins 5 and 2, and a
small blackish patch or spot on vein 1b; rarely are the
transverse markings strongly indicated; a more or less
pronounced blackish discal spot at lower outer angle of
cell and a few obscure blackish spots along termen
between the vein ends; cilia argillaceous with the scale
tips white. Hind wing semihyaline white with a
smoky shade toward apex, a fine dark line along termen,
and the veins faintly outlined by smoky scaling; cilia
white with a very faint ocherous basal shade. Alar
expanse, 21-30 mm.
Male genitalia with vinculum appreciably longer and
apical process of gnathos proportionately smaller than
those of packardella. The type of stigmella (fig. 519) is
abnormal in that the apical process of gnathos is par-
tially fused. The normal bifid condition isshownin figures
520 and 521. Female genitalia with a weak, bandlike,
sclerotized ventral plate on imner surface of ductus
bursae at genital opening; bursa copulatrix smooth.
Larva unusual for the family in that the sclerotized
rings about setal tubercles IIb on mesothorax and III
of eighth abdominal segment are absent. In this
respect it is similar to the caterpillar of Hiiella zincken-
ella which it otherwise resembles in superficial appear-
ance.
TYPE LOCALITIES: San Diego, Calif. (stigmella, in
UNITED STATES NATIONAL MUSEUM BULLETIN 207
USNM); La Puerta, Calif. (maculicula, in USNM).
Foop puant: (Sedum) Hasseanthus elongatus (larvae
boring in the roots; probably also in the roots of other
succulents).
Distrizution: Unitep Srares: California, San
Diego (Mar., May, June, July, Aug., Oct.), Loma Linda
(June), La Puerta (July), Laguna (May), Half Moon
Bay (Aug.). México: Baja California (June).
This is a variable species in size, markings, and struc-
ture. Extremes of difference in genitalia are shown in
the figures. Barnes and McDunnough (1916) pointed
out that maculicula was only a color variety of stigmella
and in their Check List reduced the name to subspecific
rank. JI do not believe that it deserves even this status
for it represents only one of many color variants. In
some specimens the transverse markings on forewing
are nearly obsolete, in others more or less indicated but
incomplete, and in one specimen before me (Half Moon
Bay, Calif., Aug. 11, 1937, W. H. Lange No. 27) the
discal spot and transverse lines are conspicuous and
the latter complete except at the fold, the subterminal
line running from outer fourth of costa transversely to
outer third of inner margin, broken between veins 2 and
1b and with a sharp outward angulation near yein 5.
The genitalic differences shown in our figures at first
glance seem rather striking; but they are not consistent
and represent only individual variation.
This species seems to be native to southern California
and northwestern México. We have a series of reared
specimens from Laguna, Calif. A few larvae and pupae
have been intercepted at quarantine ports in California
from roots of succulents (‘“Sedum sp.’’) shipped from
México and from another similar interception at San
Francisco a large (85 mm.) male was reared.
134. Genus Zophodia Hiibner
Zophodia Hiibner, Verzeichniss bekannter Schmettfer]linge, p.
370 [1825].—Ragonot, Ent. Monthly Mag., vol. 22, p. 19,
1885.—Hulst, Phycitidae of N. Amer., p. 172, 1890.—
Hampson, in Ragonot, Monograph, pt. 2, p. 18, 1901.—
Spuler, Die Schmetterlinge Europas, vol. 2, p. 207, 1910.—
Dyar, Ins. Insc. Menstr., vol. 13, p. 220, 1925.—Heinrich,
Proc. U. 8. Nat. Mus., vol. 86, p. 401, 1939.—Janse, Journ.
Ent. Soc. South Africa, vol. 8, p. 39, 1945. (Type of genus:
Tinea convolutella Hubner.)
Dakruma Grote, Bull. U. 8. Geol. Geogr. Surv. Terr., vol. 4, p.
702, 1878. (Type of genus: Dakruma turbatella Grote.)
Tongue well developed. Antenna of male pubescent
and with a series of modified, papillalike setae on the
inner sides of several basal segments of the shaft; of
female simple and very shortly pubescent. Labial
palpus oblique in the male, porrect in the female.
Maxillary palpus filiform. Hind wing with veins 7 and
8 anastomosing beyond the cell; 3 and 5 connate (in
occasional specimens very shortly stalked). Highth
abdominal segment with a pair of weak ventrolateral
hair tufts.
Male genitalia with apical process of gnathos bifid,
large; apex of harpe evenly rounded; vinculum long;
anellus with base of plate narrowly sclerotized, arms
moderately long, slender, slightly twisted; aedeagus
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 239
moderately long and stout; penis partially ribbed and
pubescent.
Female genitalia with a small weak signum developed
as a plate with an inwardly projecting flange; bursa
copulatrix small, minutely and very weakly scobinate;
ductus bursae minutely scobinate, with two rather large,
sclerotized, dorsal plates at genital opening; ductus
seminalis from bursa near signum.
As here defined the genus is limited to its type species.
Its distribution is central and southern Europe, the
northern part of the United States and southern Canada.
495. Zophodia convolutella (Hiibner)
Ficures 522, 1015
Tinea convolutella Hiibner, Sammlung europiischer Schmetter-
linge, Lepidoptera 8, Tineae 2, pl. 5, fig. 34, 1796.
Tinea grossulariella Hiibner, Geschichte europaischer Schmetter-
linge, Tinea II, pl. C.a.b., fig. 2.a.b.c., [1807-1809] (larva).
Phycis grossulariella (Hiibner) Zincken, in German and Zincken,
Mag. der Ent., vol. 3, p. 144, 1818.—Treitschke, Die Schmet-
terlinge von Europa, vol. 9, pt. 1, p. 172, 1832; op. cit., vol.
10, pt. 3, p. 275, 1835.—Duponchel, Histoire naturelle des
Lépidoptéres, ou papillons de France, vol. 10, p. 206, pl.
279, fig. 9, 1836.
Zophodia grossularialis Hiibner, Verzeichniss bekannter Schmet-
tferllinge p. 370, [1825] (emended spelling for grossulariella
and to replace convolutella).
Zophodia convolutella (Hiibner), Verzeichniss bekannter Schmet-
tferllinge (sic), p. 370, [1825]—Heinemann, Schmetterlinge
Deutschlands und der Schweiz, Abt. 2, vol. 1, no. 2, p. 190,
1865.—Ragonot, Ent. Monthly Mag., vol. 22, p. 19, 1865.—
Hampson, in Ragonot, Monograph, pt. 2, p. 20, 1901.—
Staudinger and Rebel, Catalog der Lepidopteren des palae-
arctischen Faunengebietes, vol. 2, p. 25, 1901.—Spuler,
Die Schmetterlinge Europas, vol. 2, p. 207, 1910.—Heinrich,
Proc. U. 8. Nat. Mus., vol. 86, p. 402, 1939.
Myelois (Zophodia) convolutella (Hiibner) Zeller, Isis von Oken,
1839, p. 178; 1848, p. 679.
Homoeosoma convolutella (Hiibner) Herrich-Schaffer, Systema-
tische Bearbeitung der Schmetterlinge von Europa, vol. 4,
p. 107, 1849.
Pempelia grossulariae Riley, First annual report on the noxious,
beneficial and other insects of the State of Missouri, p. 140,
1869; Papilio, vol. 1, p. 108, 1881 (suggests synonymy with
convolutella).—Packard, Guide to the study of insects, p.
331, 1869.
Dakruma turbatella Grote, Bull. U. 8. Geol. Geogr. Surv. Terr.
vol. 4, pp. 702, 703, 1878; North Amer. Ent., vol. 1, p. 11,
1879.
Myelots convolutella (Hiibner) Packard, Guide to the study of
insects, ed. 7, p. 331, 1880.
Dakruma grossulariae (Riley) Grote, North Amer. Ent., vol. 1,
p. 68, 1880.
Dakruma convolutella (Hiibner) Grote, New check list of North
American moths, p. 55, 1882 (gives grossulariae and turba-
tella as synonyms).
Zophodia grossulariae (Riley) Hulst, Phycitidae of N. Amer., p.
173, 1880; U.S. Nat. Mus. Bull. 52, p. 429, 1903.—Hampson,
in Ragonot, Monograph, pt. 2, p. 21, 1$01.—Dyar, Proc.
Ent. Soc. Washington, vol. 7, p. 37, 1905; Ins. Insc. Menstr.,
vol. 13, p. 221, 1925.—Pack, Utah Agr. Exp. Station. Bull.
216, pp. 1-12, 1930.—McDunnough, Check list, No. 6303,
1939.
Euzophera franconiella Hulst, Phycitidae of N. Amer., p. 177,
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 61, 1901.
Zophodia bella Hulst, Canadian Ent., vol. 24, p. 61, 1892.—Dyar,
Proc. Ent. Soc. Washington, vol. 6, p. 228, 1904.
Zophodia franconiella (Hulst) Barnes and MecDunnough, Check
list of the Lepidoptera of Boreal America, No. 5706, 1917.
Zophodia grossulariae franconiella (Hulst) Dyar, Ins. Insc.
Menstr., vol. 13, p. 221, 1925.
Zophodia grossulariae thouna Dyar, Ins. Insc. Menstr., vol. 13,
p. 221, 1925.
Zophodia grossulariae dilativitta Dyar, Ins. Insc. Menstr., vol.
13, p. 222, 1925.
Zophodia grossulariae magnificans Dyar, Ins. Inse. Menstr., vol.
13, p. 222, 1925.
Forewing fuscous dusted with white, the white color
strongest in central costal area, the general color gray;
antemedial line outwardly oblique to lower vein of cell
and notched between cell and inner margin, white,
bordered outwardly by a more or less extended black
shade; subterminal line oblique, slightly dentate and
sinuate, white, bordered inwardly by a black line and
outwardly by a narrow black line for a short distance
from costa; the fold and veins at extreme base of wing
and in area beyond subterminal line faintly outlined in
black; discal mark at end of cell black, curved, rarely
replaced by a pair of dots; a row of black dots along
termen between the vein ends. Hind wing pale smoky
white with a narrow dark line along termen. Alar
expanse, 25-36 mm.
Male genitalia with lateral margins of vinculum
broadly and shallowly excavate, its posterior margins
straight. Female genitalia with signum small and
weak. The eighth-segment collar is subject to con-
siderable variation in the size and shape of the unsclero-
tized dorsal area; but these variations do not conform to
the varieties that have been named and can be found
in any series from one locality.
TYPE LocaLities: Germany (convolutella and grossu-
lariella, no known existing types); Missouri (grossu-
lariae, no known existing type); Oldtown, Maine
(turbatella, in BM); Franconia, N. H. (franconiella, in
AMNH, ex Rutgers); Massachusetts (bella, in AMNH,
ex Rutgers); southern Utah (ihowna, in USNM); San
Diego, Calif. (dilativitta, in USNM) Seattle, Wash.
(magnificans, in USNM).
Foon piants: Ribes grossularia Linnaeus and other
Ribes species (larva feeding in the fruit).
Distrisution: Evurorr (central and southern).
Unitep States: Maine, Orono; New Hampshire,
Hampton (May), Durham; Missouri; Colorado, Mani-
tou, Denver (Apr.), Fort Collins (Mar., Apr.); Utah,
Logan (‘June’), Beaver Canyon (‘vii’); Oregon;
California, San Diego; Washington, Seattle, Bellingham
(Apr.). Canapa: Quebec, St. Johns County (Apr.),
Mount St. Hilaire (May); Ontario, Hymers; Alberta,
Edmonton (May), Bilby (May); British Columbia,
Kaslo (Apr.), Wellington (Apr.), Alberni (May),
Goldstream (May), Vancouver Isl. (Apr.).
The foregoing localities are for the specimens before
me. The species is generally distributed over the
northern part of the United States and southern
Canada.
The species is more extensively treated in my paper
on the cactus-feeding Phycitinae (Proc. U.S. Nat. Mus.,
vol. 86, pp. 402-405, 1939). In this country it is popu-
larly known as the ‘gooseberry fruit-worm.” It has a
rather extensive economic literature and is the most im-
240 UNITED STATES NATIONAL MUSEUM BULLETIN 207
portant lepidopterous pest of the gooseberry here and
abroad, often doing serious injury. It is also recorded
as an occasional enemy of currants.
There is one generation a year, the moths flying from
mid-April to early June. About 10 months are passed
in the pupal stage, the insects overwintering as pupae in
loose cocoons on the ground under fallen leaves and
other rubbish.
135. Genus Melitara Walker
Melitara Walker, List, pt. 27, p. 136, 1863.—Hulst, Phycitidae of
N. Amer., p. 171, 1890.—Ragonot, Monograph, pt. 2, p. 12,
1901.—Dyar, Proc. Ent. Soc. Washington, vol. 30, p. 133,
1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 338,
1939.—Dodd, Biological campaign against prickly-pear,
Brisbane, Australia, pp. 38, 39, 58, 1940. (Type of genus:
Melitara prodenialis Walker.)
Megaphycis Grote, Canadian Ent., vol. 14, p. 30, 1882.
of genus: Zophodia bollii Zeller.)
Tongue developed (stout, but rather short). Antenna
of male bipectinate, of female shortly bipectinate. La-
bial palpus porrect. Maxillary palpus squamous. Hind
wing with veins 7 and 8 approximate beyond cell; 3 and
5 connate (rarely very shortly stalked). Eighth ab-
dominal segment of male simple.
Male genitalia with apical process of gnathos bifid;
harpe with apex evenly rounded; vinculum short; anel-
lus with base of plate narrowly sclerotized, arms mod-
erately long and stout; aedeagus stout, moderately long.
Female genitalia without signum, the latter replaced
by a few very fine scobinations (not distinguishable in
most preparations except under high magnification),
bursa copulatrix otherwise simple; ductus seminalis from
bursa near junction of ductus bursae and bursa copula-
trix.
Larva bluish, not banded; sclerotized plates sur-
rounding body setae rather small; 3 setae in group VII
on abdominal segments 7 and 8.
The larvae feed gregariously in the joints of various
species of Platypuntia.
Eggs laid in chains (‘egg sticks”).
Melitara and the 16 genera following (Olycella to Cac-
tobrosis) form what appears to be a natural group of
genera and species more closely related to each other
than to other Phycitinae, although they also show dis-
tinct affinities to Zophodia and the coccid feeders of the
Laetilia complex. They all have a common host asso-
ciation, their larvae being internal feeders in the fruits
and stems of various cacti. A fuller treatment of the
several species will be found in my paper on the cactus-
feeding Phycitinae (Proc. U. S. Nat. Mus., vol. 86,
pp. 331-413, 1939).
The genus Melitara as here defined is separated from
the other cactus-feeding Phycitinae by the following
combination of characters: Antennae bipectinate in
both sexes; labial palpi porrect in both sexes; veins 7 and
8 of hind wing approximate; veins 3 and 5 of hind wing
connate; eighth abdomial segment of male simple;
larvae not banded or conspicuously spotted, gregarious
in habit throughout feeding period.
(Type
Two species are recognized as belonging to the genus,
and its distribution is apparently limited to the United
States and adjacent areas in northern México.
496. Melitara prodenialis Walker
Fieures 74, 527, 1016
Melitara prodenialis Walker, List, vol. 27, p. 137, 1863.—Hulst,
Phycitidae of N. Amer., vol. 17, p. 171, 1890; U. S. Nat,
Mus. Bull. 52, p. 429, 1903.—Ragonot, Monograph, pt. 2,
p- 13, 1901.—Hunter, Pratt, and Mitchell, U. S. Dep. Agr.
Bur. Ent. Bull. 113, p. 28, 1912.—Dodd, Council for Sci.
and Ind. Res., Australia, Bull. 34. p. 27, 1927; Biological
campaign against prickly-pear, Brisbane, Australia, p. 77,
1940.—Dyar, Proc. Ent. Soc. Washington, vol. 30, p. 133,
1928.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 339,
1939.—MeDunnough, Check list, No. 6277, 1939.
Zophodia bollit Zeller, Verh. zool.-bot. Ges. Wien, p. 550, pl. 3,
fig. 21, 1872.
Megaphycis bollit (Zeller) Grote, Canadian Ent., vol. 14, p. 30,
1882.
Melitara prodenialis bollii (Zeller) Dyar, Proc. Ent. Soc. Wash-
ington, vol. 30, p. 133, 1928.
Palpi, head, and thorax cinereous fuscous sparsely
dusted with white, especially on basal segments of Jabial
palpi; posterior margin of thorax blackish. Forewing
cinereous fuscous with a heavy dusting of white on costal
half; the fuscous and whitish areas contrasted but not
sharply defined, the white dusting most pronounced
between antemedial and subterminal lines and in
subapical area beyond subterminal line; a few black
scales scattered over entire wing; antemedial line
narrow, black, outwardly angled from basal fourth of
costal margin to 1b, less sharply defined from there to
inner margin; subterminal line narrow, black out-
wardly margined by a narrow border of white, beyond
which is a faintly dark shading which forms an obscure
line paralleling the subterminal line, the parallel black
lines most pronounced from costal margin near apex to
vein 6; subterminal line irregularly dentate and smuate,
the angulations rather shallow; discal dots fused, form-
ing a black line or smudge along discocellular vein; a
row of black dots along termen at the vein ends; cilia
grayish fuscous; underside of wing grayish fuscous, in
some specimens with a more ocherous tint. Hind wing
white, semihyaline with more or less fuscous suffusion
at apex and along costal and terminal margins, the
fuscous shading more extended in female than male;
cilia white with a narrow, dark subbasal line. Alar
expanse, 31-45 mm.
Male genitalia with outer margins of vinculum evenly
curved; elements of transtilla rather broad. Female
genitalia with ductus bursae rather slender for most of
its length.
Type Locatitizs: ‘United States” (prodenialis, in
BM); Texas (bollvi, Cambridge Mus. Nat. Hist.).
Foop piants: Opuntia (Platypuntia) spp.
Distrisution: Teras, Dallas, Freeport, Utopia, Col-
lege Station (Oct.), Brownsville; Mississippi, Biloxi
(Sept.); Oklahoma, Wichita National Forest (June);
Florida, Altamont (Sept., Oct.), Key West, Lakeland
(Apr.), Crescent City (May), Miami (Oct.), St. Peters-
burg (Mar., June, Sept., Oct.), Fort Meade (Apr.), Fort
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Myers (Apr.), Venice (May); North Carolina, Southern
Pines (June); Delaware, Indian River Bay (July); New
Jersey, Lakehurst (Sept.); New York, Rye (July).
497. Melitara dentata (Grote)
Fieures 528, 1017
Zophodia dentata Grote, Canadian Ent., vol. 8, p. 158, 1876;
Bull. U. S. Geol. Geogr. Surv. Terr., vol. 3, p. 799, 1877.
Megaphycis dentata (Grote), Canadian Ent., vol. 14, p. 30, 1882.
Melitara deniaia (Grote) Hulst, Phycitidae of N. Amer., p. 172,
1890; U. S. Nat. Mus. Bull. 52, p. 429, 1903.—Kellogg,
Kansas Univ. Quart., vol. 1, p. 39, 1892.—Ragonot, Mono-
graph, pt. 2, p. 14, 1901.—Hunter, Pratt, and Mitchell,
U. S. Dep. Agr. Bur. Ent. Bull. 113, p. 28, 1912.—Dodd,
Council for Sci. and Ind. Res., Australia, Bull. 34, p. 29,
1927; Biological campaign against prickly-pear, Brisbane,
Australia, p. 79, 1940.—Dyar, Proc. Ent. Soc. Washington,
vol. 30, p. 134, 1928.—Heinrich, Proc. U. 8. Nat. Mus. vol.
86, p. 341, 1989.—McDunnough, Check list No. 6278, 1939.
Melitara doddalis Dyar, Ins. Insc. Menstr., vol. 13, p. 13, 1925;
Proc. Ent. Soc. Washington, vol. 30, p. 184, 1928.—Dodd,
Council for Sci. and Ind. Res., Australia, Bull. 34, p. 29,
1927; Biological campaign against prickly-pear, Brisbane,
Australia, p. 80, 1940.
Melitara junctolineella Hulst (in part), Canadian Ent., vol. 32,
p. 173, 1900.—Barnes and McDunnough, Contributions,
vol. 3, p. 199, 1916.
Melitara bollit Dodd (not Zeller), Council for Sci. and Ind. Res.,
Australia, Bull. 34, p. 29, 1927.
General color and pattern as in prodenialis except:
Blackish shading on posterior margin of thorax less pro-
nounced and in some specimens not distinguishable.
Forewing with white dusting rather evenly distributed
over the entire wing, the whitish and fuscous areas not
contrasted except (in some specimens) for a rather
narrow pale suffusion along costal margin and a more
or less pronounced dark shade from end of cell to middle
of inner margin; the subterminal line with no black
shading beyond its white border except for a short dis-
tance from apex, markedly dentate and sinuate, the
angulations deep, the angulation between veins 5 and
6 reaching to the cell. Hind wing semihyaline, in the
male almost pure white with little or no fuscous shading,
the latter, when present, confined to a narrow band
along costal margin and a fine line along termen; in the
female fuscous shadings nearly always pronounced,
though limited to a narrow line along costal margin, a
slight clouding at apex, and a thin line along termen;
cilia white. Alar expanse, 33-50 mm.
Male genitalia with outer margins of vinculum slight-
ly sinuate; elements of transtilla slightly narrower than
those of prodenialis. Slight differences in the shape of
the anellus between the two species shown in figures
527e-f and 528b. Female genitalia essentially like
those of prodenialis except that ductus bursae is nor-
mally stouter.
Typr Locaities: Clear Creek Canyon, Colo. (den-
tata, in BM); Mesilla Park, N. Mex. (doddalis, in
USNM).
Foop puants: Opuntia (Platypuntia) spp.
DistrisuTion: Unirep States: Wyoming, Evanston
(July), Medicine Bow (July); Colorado, Glenwood
Springs (July, Aug., Sept.), Fort Collins, Denver, Rocky
241
Ford (Sept.); Utah, Buckskin Valley (Iron County,
June, July), Eureka (Aug.), Dividend (Aug., Sept.);
Kansas, Manhattan (Sept.); Arizona, Mormon Lake
(July), Douglas (Aug., Sept.), Oracle (Sept.), Globe
(Sept.), Quijotoa (Oct.), Chiricahua Mts.; New Mexico,
Mesilla Park (Sept.), Silver City (Sept.), Jemez Springs
(July, Aug., Sept.); Texas, Uvalde (Sept., Oct., Nov.),
Henrietta (Oct.), Trent (Oct.), Rock Springs, Laredo
(Sept.), Shafter (Sept.), Albany, Panhandle (Aug.).
México: Chihuahua (city), Julimes (Sept., Oct.), Mo-
relia, (Oct.).
Shortly after the publication of my 1939 paper on
the cactus-feeding phycitids (in which I discussed the
synonymy of dentata and doddalis) we received through
Mr. Leith Hitchcock a series of moths from Uvalde,
Tex., labeled Melitara bollii. They were evidently the
specimens that Dodd distinguished from prodenialis in
treating bolli as a distinct species. Dyar was respon-
sible for the original identification of bollii with pro-
denialis as a race or variety. He apparently was con-
fused by Dodd’s biological notes on the Uvalde moths.
Some of these moths he placed with other Texas speci-
mens of true bollii, while others from the same Uvalde
rearing he included under his doddalis. M. bollit
Zeller is a synonym of prodenialis while bollii Dodd (not
Zeller) is merely a southern Texas form of the exceed-
ingly variable dentata Grote.
136. Genus Olycella Dyar
Olycella Dyar, Proc. Ent. Soc. Washington, vol. 30, p. 134,
1928.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 348,
1939.—Janse, Journ. Ent. Soc. South Africa, vol. 8, p. 40,
1945. (Type of genus: Melitara junctolineella Hulst.)
Tongue developed (as in Melitara). Antenna of male
bipectinate, of female shortly bipectinate. Labial pal-
pus obliquely ascending (sometimes in the female the
third segment is bent forward, which gives the palpus a
porrect appearance, but the second segment is always
deflected upward and reaches nearly as high as the top
of the head). Maxillary palpus squamous. Hind wing
with veins 7 and 8 approximate beyond the cell; 3 and
5 connate. Eighth abdominal segment of male simple,
Male genitalia with apical process of gnathos bifid;
harpe with apex evenly rounded; vinculum short;
anellus with base of plate narrowly sclerotized, arms
moderately long and stout; aedeagus stout, moderately
long.
Female genitalia with signum, the latter a small
ridged plate; bursa copulatrix wrinkled, otherwise
simple and without scobinations; ductus bursae with a
pair of sclerotized plates on inner wall at genital open-
ing; ductus seminalis from center of bursa.
Larva white with broad blackish or purplish cross
bands on the caudal margins of the segments; sclero-
tized plates surrounding setae rather small; three setae
in group VII on abdominal segments 7 and 8.
The larvae feed gregariously for a short period after
hatching (probably during the first instar) but there-
after are solitary in habit. They feed in the joints of
various Platypuntias.
242
The genus is close to Melitara, distinguished from it
only by the following characters: Labial palpi obliquely
ascending; larvae transversely banded and solitary in
habit during most of the feeding period.
Three species and one local race are here recognized.
They are remarkably alike in structure, the genitalia
exhibiting difference of only an individual character.
The species, however, can be distinguished by color
differences which seem to be constant.
aa 498. Olycella junctolineella (Hulst)
Fieures 75, 529, 1018
Melitara junctolineella Hulst, Canadian Ent., vol. 32, p. 173, 1900;
U.S. Nat. Mus. Bull. 52, p. 429, 1803.—Hunter, Pratt, and
Mitchell, U. S. Dep. Agr. Bur. Ent. Bull. 113, p. 25, 1912—
Barnes and McDunnough, Contributions, vol. 3, p. 199,
1916.—Dodd, Council for Sci. and Ind. Res., Australia,
Bull. 34, p. 27, 1927.
Olyca junctolineella (Hulst) Barnes and McDunnough, Check list
of the Lepidoptera of Boreal America, No. 5695, 1917.
Olycella junctolineella (Hulst) Dyar, Proc. Ent. Soc. Washington,
vol. 30, p. 134, 1928.—Heinrich, Proc. U. 8. Nat. Mus.,
vol. 86, p. 344, 1939.—MeDunnough, Check list, No. 6281,
1939.—Dodd, Biological campaign against prickly-pear,
Brisbane, Australia, pp. 39, 59, 81, 1940.
Head, thorax, and forewing ocherous fuscous dusted
with white and marked with patches and lines of black
scales. Labial palpus with the apical ends of the seg-
ments blackish. Maxillary palpus crossbanded with
black scales. Thorax with some black dusting on
posterior margin: Forewing with whitish dusting
slightly mtensified in costal area; veins faintly outlined
in black; a row of more or less obscure black dots on
termen between the vein ends; antemedial line inter-
rupted, sometimes obscure, in fresh, well-marked speci-
mens its outer dentation much extended and meeting a
shade from the inner angulation of the subterminal line
at the fold (which indicates the normal position of the
absent vein 1c); subterminal line interrupted, strongly
indicated only between veins 5 and the fold and for a
short distance from inner margin; black discal dots at
end of cell more or less fused and pronounced; cilia
ocherous fuscous. Hind wing pure white in male; in
female generally suffused with fuscous. Alar expanse,
38-55 mm.
The genitalia presenting no outstanding specific
characters.
TYPE LOcALITy: Texas (type in AMNH, ex Rutgers).
Foon piants: Opuntia (Platypuntia) spp.
Distrisution: Texas, Brownsville (Apr., June, July,
Aug.), Corpus Christi (Sept., Oct.), Burnet County,
San Benito (Mar., Aug., Sept.), Shovel Mountain
(May), Kerrville (Apr.), Victoria (Oct., Nov.), Laredo
(Sept.).
This species is most readily distinguished from the
others in the genus by its ocherous fuscous color, which
seems to be constant. It is remarkably so in the
specimens before me. O. nephelepasa and subumbrella
are decidely gray in appearance.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Rather full notes on the life history and larval habits
of junctolineella are given by Dodd and by Hunter,
Mitchell, and Pratt. The latter also give descriptions
of the larva and pupa.
The species has two generations a year.
499. Olycella junctolineella pectinatella (Hampson)
Fiaure 530
Olyca pectinatella Hampson, 7n Ragonot, Monograph, pt. 2, p. 35,
1901.
Olyca junctolineella (Hulst) Barnes and McDunnough, Check list
of the Lepidoptera of Boreal America, No. 5695, 1917 (in
art).
Olycella cote (Huist) Dyar, Proc. Ent. Soc. Washington,
vol. 30, p. 1384, 1928 (in part).
Olycella junctolineella pectinatella (Hampson) Heinrich, Proc.
U.S. Nat. Mus., vol. 86, p. 345, 1989.
Known only from two males from the type locality.
These two specimens are a trifle paler than normal
junetolineella and are less clearly marked except for the
pronounced discal spots. The transverse lines on the
forewing are almost obsolete and the veins very slightly
indicated by dark shading.
Inasmuch as we have no larvae or females, or any
information on the life history of the form from Jalapa
(which is far south of the known range of typical
junctolineella), I do not think we are justified in treating
it as @ Mere synonym; or, on the evidence before us,
as a distinct species.
TYPE Locauity: Jalapa, State of Veracruz, México
(type in USNM).
Foop puant: Opuntia (Platypuntia) spp.
500. Olycella nephelepasa (Dyar)
Olyca nephelepasa Dyar, Ins. Insc. Menstr., vol. 7, p. 55, 1919.
Olycella nephelepasa (Dyar), Proc. Ent. Soc. Washington, vol. 30,
p. 134, 1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p.
346, 1989.
Similar in pattern and general appearance to junc-
tolineella, but darker. The forewing is grayish fuscous
with a slight brownish tint, but decidedly more grayish
brown than ocherous fuscous. Also the hind tibiae of
nephelepasa are heavily dusted with fuscous, while those
of junctolineella are nearly pure white. Alar expanse,
42-52 mm.
The genitalia cannot be distinguished from those of
junctolineella.
Type Locatity: Tehuacén, México (type in USNM),.
Foop piants: Opuntia (Platypuntia) spp.
Disrrisution: México: Tehuacin (Sept.), México
(city), Cuernavaca, Aguascalientes, San Luis Potosi
(June).
The known distribution of this species is confined to
the central plateau of México. In the National Col-
lection there is one female (determined as nephelepasa)
from Monclova, México. This specimen is colored like
typical subwmbrella and is, I think, only a southern
example of that species.
O. nephelepasa has two generations a year.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 243
501. Olycella subumbrella (Dyar)
Ficure 1019
Olyca subumbrella Dyar, Ins. Insc. Menstr., vol. 13, p. 14, 1925.
Olycella nephelepasa (Dyar), Proc. Ent. Soc. Washington, vol. 30,
p. 134, 1928 (in part).
Olycella subumbrella (Dyar) Heinrich, Proc. U. S Nat. Mus.,
vol. 86, p. 347, 1939.—Dodd, Biological campaign against
prickly-pear, Brisbane, Australia, p. 39, 1940.
Similar to nephelepasa except that the forewing is less
brownish and more grayish. The general color is de-
cidedly gray rather than brownish or ocherous. Alar
expanse, 40-55 mm.
Male genitalia as in nephelepasa and junctolineella.
Female genitalia like those of junctolineella. The figure
shows the extreme variation from typical junctolineella;
but the differences in the shape of the sclerotized areas
of the collar of the eighth segment and the length of the
supporting rods of the collar are not specific. Every
intergrade between this and typical junctolineella may
be found in each of the species in the genus.
Type Locauiry: Carlsbad, N. Mex. (type in USNM).
Foop pLants: Opuntia (Platypuntia) spp.
Distrisution: Texas, El Paso (Mar.); New Mexico,
Carlsbad (Sept.); Arizona, Dewey, Redington, Palmer-
lee, Paradise (Cochise County, Mar., Apr., May, June),
Douglas (May, Aug.,) Pinal Mts. (Apr.), Haualapai
Mts. (May); California, Warner (Sept.), Santa Clara
(Apr.); Utah, Dividend (May, June), Stockton (May),
Richfield (May); Nebraska, Scotts Bluff (June).
In addition to the above there are before me two
specimens from Monclova, Coahuila, México (E.
Mortensen Collection, Sept. 1926), which probably are
referable here. One (a male) was in the collection
under junctolineella, the other (a female) under nephele-
pasa. The male is in very poor condition but obviously
belongs with the female. The latter is in fair shape,
and its color is that of typical swbumbrella. More
material is needed from northern México before we can
determine what species inhabits that region.
In 1928 Dyar sank subumbrella in the synonymy of
nephelepasa; but Dodd informs me that the larval
habits of the two are quite different. In nephelepasa
“the larvae are banded with blue or blue-black and do
not keep entrance hole in plant open for discharge of
frass,’”’ while in subumbrella the larvae have ‘“‘rather pale
purplish bands and maintain the hole open for the dis-
charge of frass.’”’ These differences in larval habit,
coupled with the slight but apparently consistent color
differences in the moths seem to warrant the separation
of subumbrella from nephelapasa.
O. subumbrella has one generation a year.
137. Genus Olyca Walker
Olyca Walker, List, pt. 11, p. 725, 1857.—Hampson, in Ragonot,
Monograph, pt. 2, p. 34, 1901.—Dyar, Proc. Ent. Soc.
Washington, vol. 30, p. 133, 1928.—Heinrich, Proc. U. 8S.
Nat. Mus., vol. 86, p. 348, 1939. (Type of genus: Olyca
phryganoides Walker.)
Tongue reduced (shorter than in Méelitara). An-
tennae pubescent in both sexes (the pubescence longer
in the male than in the female), slightly serrate in the
male. Labial palpus of the male obliquely ascending;
of the female porrect and downcurved. Maxillary
palpus squamous. Hind wing with veins 7 and 8
shortly anastomosed beyond cell; 3 and 5 connate (oc-
casionally very shortly stalked). Eighth abdominal
segment of male simple.
Male genitalia with apical process of gnathos bifid;
harpe with apex evenly rounded; vinculum short;
anellus with base of plate narrowly sclerotized, arms
moderately long and stout; aedeagus stout, moderately
long, apex armed with many minute, hairlike spines.
Female genitalia without signum; bursa copulatrix
simple except for a few microscopic scobinations; ductus
bursae short; ductus seminalis from bursa somewhat
caudad of middle.
Larvae not banded, solitary in habit, feeding in
Platypuntias (presumably in the stems).
Eggs unknown.
Olyca is readily separated from other genera of the
cactus-feeding Phycitinae having veins 3:and 5 of hind
wing connate by having the antennae pubescent in both
sexes.
The male genitalia are similar to those of Olycella,
differing only in slight details; the vinculum is slightly
shorter, the uncus broader in proportion to its length,
the cleft apical process of gnathos smaller, the elements
of transtilla longer and straighter.
The females differ chiefly in that they lack the signum
in the bursa and the sclerotized plates in the opening
of the ductus bursae.
The genus as here defined contains only the type
species from the West Indies.
502. Olyca phryganoides Walker
Figures 71, 531, 1020
Olyca phryganoides Walker, List, pt. 11, p. 726, 1857—Hampson,
in Ragonot, Monograph, pt. 2, p. 35, 1901.—Dyar, Proc.
Ent. Soc. Washington, vol. 30, p. 134, 1928.—Heinrich,
Proc. U. S. Nat. Mus., vol. 86, p. 348, 1939.—Dodd, Bio-
logical campaign against prickly-pear, Brisbane, Australia,
p. 40, 1940.
General color (except hind wings) pinkish white, more
or less spotted and suffused with black. Palpi, thorax,
and underside of body heavily dusted and shaded with
black. Forewing of male with no distinguishable ante-
medial line, the latter being replaced by two more or
less transversely extended black spots; subterminal line
only partially and faintly indicated, irregularly dentate;
vein ends marked with blackish dots or dashes; black
discal spot large, conspicuous; below the discal dot a
more or less extended black smudge.
On the female about three-fourths of the forewing is
suffused with black, the pinkish white color being
strongly contrasted and limited to a rather narrow area
along the costa, with a triangular projection at the end
of the cell; terminal area and a patch on inner margin
opposite discal spot also pale, but duller and less con-
trasted than the costal color. Hind wing white with a
blackish fuscous shade on terminal margin and a some-
244
what wider dark shade along costal margin; these dark
shadings broader in the female than the male; also in
the female the veins are outlined by fuscous scaling;
cilia white with a dark subbasal line. Alar expanse,
35-47 mm.
Male genitalia figured from specimen from Azuda.
Female genitalia with bursa copulatrix containing a
scattering of microscopic scobinations, otherwise simple;
scobinations in genital opening stronger and more
dense, also in genital opening a few fine setae (the latter
probably a generic character).
Eggs unknown.
Larva ‘cream or buff colored, with dark spiracular
markings,” according to Dodd.
Typ LocaLity: Hispaniola (type in BM).
Foop piant: Opuntia (Platypuntia) spp.
Distrinution: Dominican Repusuic: Azuda (Jan.).
Hartt: Port-au-Prince (Jan.).
Nothing has been published on the life history of this
species, and little is known about it. ‘The larvae are
presumably solitary in habit and confined to the Platy-
puntias. The distribution of phryganoides is probably
confined to the West Indies.
138. Genus Alberada Heinrich
Alberada Heinrich, Proce. U. S. Nat. Mus., vol. 86, p. 350; 1939.
(Type of genus: Melitara parabates Dyar.)
Tongue as in Melitara. Antenna of male> bipecti-
nate, of female pubescent. Labial palpus porrect and
downcurved. Maxillary palpussquamous. Hind wing
with veins 7 and 8 anastomosing beyond cell; 3 and 5
stalked. Highth abdominal segment of male simple.
Male genitalia with apical process of gnathos bifid,
the two prongs rather widely separated; harpe with the
apex evenly rounded; vinculum short; anellus with base
of plate narrowly sclerotized, arms moderately long and
stout; aedeagus stout, weakly sclerotized in middle
except on midventer.
Female genitalia with sigsnum developed as a ridged
plate; bursa copulatrix finely scobinate and wrinkled;
ductus seminalis from middle of bursa.
Larvae bluish, not banded; solitary feeders in the
joints of Cylindropuntias.
Eggs laid singly or in masses of two or three.
The genus is close to Melitara, differing in the follow-
ing characters: Veins 7 and 8 of hind wing anastomosed,
3 and 5 stalked, aedeagus only partially sclerotized,
apical process of gnathos with prongs well separated,
bursa copulatrix with signum.
The distribution is apparently limited to México and
the southwestern part of the United States.
503. Alberada parabates (Dyar)
Fieures 533, 1021
Melitara parabates Dyar, Proc. U. S. Nat. Mus., vol. 44, p. 322,
1913; Proc. Ent. Soc. Washington, vol. 30, p. 184, 1928.—
Barnes and McDunnough, Contributions, vol. 4, p. 175,
1918.—Dodd, Council for Sci. and Ind: Res., Australia,
Bull. 34, p. 27, 1927.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Alberada parabates (Dyar) Heinrich, Proc. U. 8. Nat. Mus., vol.
86, p. 350, 1939.
Forewing fuscous with area between lower vein of
cell and costal margin and from antemedial to sub-
terminal lines heavily dusted with white; area between
lower vein of cell and inner margin and from base to
subterminal line suffused with ocherous fuscous; on the
middle of this area a more or less extended smudge of
blackish brown; antemedial line black, bordered inwardly
by a line of white scales, dentate and sinuate, a sharp
dentation at vein 11, a longer one in the cell (extending
nearly to middle of wing), another equally long and
acute dentation at the fold, and two very slight denta-
tions between 1b and inner margin; subterminal line
black with a white outer border, dentate and sinuate,
the angulations deep, the angulation between 5 and 6
reaching almost to cell; area beyond subterminal line
dark fuscous, paler in some specimens; along termen a
row of black dots at the vein ends; discal black dot at
end of cell conspicuous in most specimens. Hind wing
white, semihyaline; costal margin bordered with
fuscous and a fine dark fuscous line on termen for a
short distance from apex; in many females a stronger
fuscous shading in apical area. Alar expanse, 35-48
mm.
Male genitalia over twice as large as those of biden-
tella; aedeagus more extensively sclerotized. Female
genitalia larger than those of the other species of the
genus and with scobinations in bursa finer.
Eggs laid singly or in masses of two or three.
Typr tocatiry: Cerritos, San Luis Potosi, México
(type in USNM).
Foop prants: Opuntia (Cylindropuntia) imbricata
(Haworth) and probably several other Cylindropuntias.
DistrizpuTion: Unirep states: California, San
Diego, Warner (Sept.), Palm Springs (Apr.), Oceanside
(Aug.), Riverside (Oct.); Arizona, Christmas (Gila
County), Fort Grant (July), Oracle (July), Redington,
Santa Catalina Mts. (Sept.), Baboquivari Mts. (Apr.,
June, July, Aug., Sept., Oct.), Sells Post Office (Indian
Oasis, Apr.), Douglas (June, Sept.), Mohave County
(Aug.); Texas, Presidio County (July), Brewster County.
México: San Luis Potosi, Cerritos (Aug.), Tamaulipas,
Tula (June).
504. Alberada bidentella (Dyar)
Ficures 532, 1022
Zophodia bidentella Dyar, Proc. Ent. Soc. Washington, vol. 10,
p. 114, 1908.
Eumysia bidentella (Dyar), Ins. Insc. Menstr., vol. 13, p. 221,
192
Alberada didentella (Dyar) Heinrich, Proc. U. S. Nat. Mus., vol.
86, p. 352, 1939.
Much smaller and paler than parabates but with simi-
lar pattern, the ground color more ocherous than fus-
cous, the white dusting on forewing heavier, the denta-
tions of antemedial and subterminal lines shorter; discal
dots distinct and not fused as is frequently the case in
parabates. Alar expanse, 19-24 mm.
Male genitalia similar to those of parabates but much
smaller and with central ventral part of aedeagus more
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
narrowly sclerotized. Female genitalia appreciably
smaller than those of parabates; bursa wrinkled and
more coarsely scobinate.
Typs Locaity: San Antonio, Tex. (type in USNM).
Foop piant: Unknown.
Distrisution: TZexras, San Antonio (July), San
Benito (June, Aug., Sept.), Brownsville (June) ; Arizona,
Phoenix, ‘‘route between Dewey and Salome.”
A uniformly marked and colored species, known only
from collected specimens.
505. Alberada holochlora (Dyar)
Figure 1023
Zophodia holochlora Dyar, Ins. Insc. Menstr., vol. 13, p. 15, 1925.
Alberada holochlora (Dyar) Heinrich, Proc. U. S, Nat. Mus., vol.
86, p. 352, 1939.
This is probably a synonym or, at most, a variety of
bidentella. 'The three females of the type series are the
only specimens I have seen. They are a trifle smaller
than typical bidentella, and there are some slight, though
hardly significant, differences in the female genitalia
(shown in figs. 1022 and 1023). However, until males
of holochlora are discovered and bidentella has been
reared, it will be wiser to keep the two as separate
species.
According to Dodd the larvae are solitary in habit
and dark blue and the eggs laid singly.
Alar expanse, 18 mm.
Types Locauity: Uvalde, Tex. (type in USNM).
Foop piant: Opuntia (Cylindropuntia) leptocaulis De
Candolle.
139. Genus Nanaia Heinrich
Nanaia Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 353, 1939.
(Type of genus: Nanaia substituta Heinrich.)
Tongue well developed. Antenna of male bipec-
tinate; of female pubescent. Labial palpus obliquely
porrect (second segment obliquely upturned nearly to
top of face and third segment bent forward or slightly
downcurved) ; third segment long (in the female as long
as second segment), pointed in the male, blunt in the
female. Maxillary palpus large, developed as a broad,
curved, somewhat flattened tuft of scales which reaches
well above middle of face. Hind wing with veins 7 and
8 anastomosing beyond cell; 3 and 5 stalked. Eighth
abdominal segment of male simple.
Male genitalia with apical process of gnathos bifid,
the two prongs narrowly separated; harpe with the apex
somewhat tapering but bluntly rounded; vinculum
moderately long (longer than broad); anellus with base
of plate narrowly sclerotized, arms long, tapering, and
slightly twisted; aedeagus moderately stout, strongly
sclerotized throughout.
Female genitalia without signum; ductus bursae and
bursa copulatrix simple except for fine scobinations in
ductus at genital opening; ductus bursae long; bursa
copulatrix small; ductus seminalis from middle of bursa.
Larva bluish, not banded; sclerotized plates sur-
rounding body setae small; 2 setae in group VII on ab-
dominal segments 7 and 8.
245
The larvae are solitary feeders in the trunks of Cylin-
dropuntia and Trichocereus.
Ege and egg-laying habits unknown.
This genus is close to Alberada but distinguished by
several characters: The forewings are distinctly nar-
rower, the vinculum is longer in proportion to its width,
the aedeagus more evenly sclerotized, the apical process
of gnathos more narrowly cleft, the anellus more de-
cidedly curved, the bursa simple, without signum or
scobinations, the transverse markings on forewing al-
most obliterated, and the maxillary palpi much larger.
The maxillary palpi are similar to those of Sigelgaita,
the moths of which resemble in general habitus those of
Nanaia. The two genera, however, are easily dis-
tinguished by their different labial palpi, porrect in
Nanaia, upturned in the males of Sigelgaita.
Known only from Pert.
506. Nanaia substituta Heinrich
Fieures 534, 1048
Nanaia substituta Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 354
General color (except hind wings) ocherous fuscous
peppered with black and white; the type darker than
most of the paratypes. Forewing with pale color con-
fined to costal half of wing; terminal area and the area
between cell and inner margin darker, with very little
white dusting; in most specimens a rather pronounced,
broad, longitudinal, ocherous fuscous shade in the fold;
transverse and discal markings almost obsolete, in a few
specimens the antemedial line faintly indicated and in
the palest of the paratypes the discal black dots dis-
tinguishable, also some black scaling along the veins.
Hind wing white (whitish ocherous on female) with a
smoky tint toward apex and termen; terminal margin
blackish fuscous; cilia smoky white with a dark sub-
basal line. Alar expanse, 37-40 mm.
Female genitalia with no appreciable scobinations or
granulations in bursa; ductus bursae minutely scobinate
at genital opening, otherwise smooth.
TypE Locauiry: Cuzco. Pert (type in USNM).
Foop piant: Opuntia (Cylindropuntia) exaltata
Berger.
140. Genus Cactoblastis Ragonot
Cactoblastis Ragonot, Monograph pt. 2, p. 15, 1901.—Dyar, Proc.
Ent. Soc. Washington, vo. 30, p. 135, 1928.—Heinrich, Proc.
U. S. Nat. Mus., vol. 86, p. 354, 1939.—Janse, Journ. Ent.
Soc. South Africa, vol. 8, p. 45, 1945. (Type of genus:
Zophodia cactorum Berg.)
Neopyralis Bréthes, in Ronna, Chacaras e Quinaes, vol. 20, No. 1,
p. 18, 1920. (Type of genus: Neopyralis ronnai Bréthes.)
Tongue considerably reduced. Antenna of male
pubescent, of female shortly pubescent. Labial palpus
of male ascending (upcurved), of female porrect. Hind
wing with veins 7 and 8 shortly anastomosed beyond
cell; 3 and 5 shortly stalked. Eighth abdominal seg-
ment of male simple.
Male genitalia with apical process of gnathos partially
fused, the prongs separated only for a short distance;
246
harpe with apex evenly rounded; vinculum short;
anellus with base of plate narrowly sclerotized, arms
moderately long and rather slender, slightly twisted,
very finely serrate on outer edges toward apices;
aedeagus stout, moderately long.
Female genitalia with sigsnum developed as a series of
more or less fused plates; bursa copulatrix weakly and
very finely scobinate; ductus seminalis from bursa at
junction of ductus bursae and bursa copulatrix.
Larva bright orange or red, with rows of large black
spots resembling broken crossbands; two setae in group
VII on abdominal segments 7 and 8.
The larvae feed gregariously in the joints of Platy-
puntia, Oylindropuntia, Trichocereus, Echinopsis, and
Denmoza.
Eggs laid in long chains.
The genus as here defined is distinguished from other
cactus-feeding phycitids by the following combination
of characters: Antennae of both sexes pubescent; labial
palpi upcurved in the male, porrect in the female; veins
7 and 8 of hind wing shortly anastomosed; veins 3 and 5
shortly stalked; apical process of gnathos partially
fused; eighth abdominal segment of male simple; larvae
bright orange or red, with rows of large black spots
resembling broken crossbands, gregarious in habit.
Four (possibly five) species are recognized as belong-
ing to the genus. Its natural distribution is apparently
limited to South America south of the Equator, but at
least one of its species (cactorum) has been introduced
and become established in Australia.
507. Cactoblastis cactorum (Berg)
Ficures 66, 535, 1024
Zophodia cactorum Berg, Anales Soc. Cient. Argentina, vol. 19, p.
276, 1885.
Cactoblastis cactorum (Berg) Ragonot, Monograph, pt. 2, p. 16,
1901.— Dodd, Council for Sci. and Ind. Res., Australia, Bull.
34, p. 30, 1927; Bull. Ent. Res., vol. 27, p. 509, 1936; Bio-
logical campaign against prickly-pear, Brisbane, Australia,
177 pp., 1940.—Dyar, Proc. Ent. Soc. Washington, vol. 30,
p. 135, 1928.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p.
356, 1939.
Head sordid whitish ocherous. Palpi pale cinereous,
the tips of the maxillary palpi and the ends of the seg-
ments of the labial palpi blackish fuscous. Thorax dull
ocherous fuscous rather heavily dusted with blackish
fuscous on posterior half. Forewing ocherous fuscous
more or less dusted with white on costal half between
antemedial line and apex; antemedial line black, angu-
late, the apex of angle at vein 1b, sometimes obscure
except on costal half; subterminal line black, ‘with a
narrow whitish outer border and beyond this a faint
fuscous band, the black line straight from near apex to
vein 6, thence sinuate and dentate to inner margin, the
ends of the dentations rounded; a black spot at end of
cell and a few scattered black scales on disc; along
termen at vein ends a row of seven distinct black dots.
Hind wing white, semihyaline, costal margin narrowly
bordered with fuscous and on termen a fine black line,
the latter not extending to inner angle, on the female
the fuscous shading is rather broad at apex and along
UNITED STATES NATIONAL MUSEUM BULLETIN 207
termen for a short distance from apex. Midtibia pale
cinereous with a narrow, black, transverse band at
outer fourth. Alar expanse, 23-40 mm.
Genitalia not exhibiting any marked specific char-
acters.
TYPE LOCALITY:
known).
Foop puants: Opuntia (Platypuntia) spp. Appar-
ently limited to the Platypuntias.
Distrisution: Arecentina: La Plata, Concordia,
Tacanitas, Santiago del Estero. Unrucauay: Piriapolis.
AUSTRALIA (introduced and established).
According to Dodd “‘cactorum is a native of Uruguay
and the northern Argentine provinces of Entre Rios,
Corrientes, Sante Fe, Santiago del Estero, Tucumén,
Salta, and Chaco.” He also includes Paraguay and
southern Brazil in its possible range, but we have no
adult specimens from the latter localities.
This is the species that has been used with such
remarkable success in the biological campaign against
the prickly-pear in Queensland and New South Wales.
All available information on the species is given in
Dodd’s 1940 paper.
Argentina (location of type un-
508. Cactoblastis ronnai (Bréthes)
Neopyralis ronnai Bréthes, in Ronna, Chacaras e Quinaes, vol.
20, No. 1, p. 18, 1920.—Costa-Lima, Terceiro catalogo dos
insectos que vivem nas plantas do Brazil, p. 268, No. 1031,
1936.
Cactoblastis ronnai (Bréthes) Heinrich, Proc. U. 8. Nat. Mus.,
vol. 86, p. 357, 1939.
Probably a synonym of cactorum. Its identity will
have to wait upon rearing of Cactoblastis from southern
Brazil.
TyPE tocatity: Rio Grande do Sul, Brazil (type
lost?).
Foop pian: “Spineless cactus.”
509. Cactoblastis doddi Heinrich
Fieures 536, 1025
Cactoblastis bucyrus Dodd (not Dyar), Council for Sci. and Ind.
Res., Australia, Bull. 34, p. 30, 1927.
Cactoblastis doddi Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p.
358, 1939.—Dodd, Biological campaign against prickly-pear,
Brisbane, Australia, pp. 39, 59, 75, 1940,
Similar to that of cactorum except as follows: White
dusting on forewing less contrasted, sparser; general
color darker, decidedly grayish fuscous in specimens
from Tucumén; dentations of subterminal line of fore-
wing acute and their ends pointed; black dots along
termen very faintly imdicated, normally altogether
absent. Himd wing of male semihyaline white; of
female dark smoky fuscous throughout. Alar expanse,
31-41 mm.
Male genitalia similar to those of cactorum, differing
chiefly in the shorter cleft between the prongs at apex
of gnathos. This character, however, is subject to
some individual variation, and should be used with
discretion. Female genitalia with scobinations of
bursa. somewhat more uniformly distributed than in
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
other species of Cactoblastis, not an altogether reliable
or satisfactory character in this genus.
Typr Locauity: Tapia, Tucumén, Argentina (type
in USNM).
Foop puants: Opuntia (Platypuntia) sulphurea G.
Don, Opuntia (Platypuntia) ficus-indica (Linnaeus).
Represented only by the type series from the type
locality.
According to Dodd, this species ‘‘is distributed along
the eastern edge and foothills of the Andes from Men-
doza right to the northern boundary of the Republic in
O. sulphurea, and almost certainly into southern Bolivia
at altitudes to 8,000 fect and probably more. Hence, as
far as our information goes, No. 49 (doddi) inhabits
territory lying in between that of cactorum and the
Peruvian insect (mundelli).”’
O. sulphurea seems to be the favored host of doddi.
Dodd tells me that cactorum does not attack this cactus
although it is abundant in territory within the range
of that insect. He also states that there are consistent
differences in the eggs and egg sticks between the two
species and that their larvae can be distinguished in the
field. I am unable to separate alcoholic specimens of
the larvae with any certainty. The moths can be dis-
tinguished easily enough by the following combination
of characters: Forewing without terminal row of dots or
with but 3 or 4 very faintly indicated; hind wing of
male semihyaline white, of female dark smoky fuscous.
510. Cactoblastis mundelli Heinrich
Figure 537
Cactoblastis mundelli Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p.
359, 1939.
Head ocherous. Palpi cinereous, dusted with black.
Thorax ocherous fuscous, heavily dusted with white and
black scales, especially on tegulae and posterior margin.
Forewing ocherous fuscous with a fine dusting of white
scales in costal area from base to apex; some black scal-
ing on the veins; discal dot at end of cell somewhat
obscured by a dark smudge which extends beyond the
cell toward vein 1b; transverse black lines distinct and
well contrasted against the ground color; dentations of
subterminal line as in doddi; a row of seven small black
dots on termen at the vein ends. Hind wing of male
dull white with a faint smoky tint; veins faintly out-
lined in fuscous ocherous; some fuscous shading along
costa and a fine fuscous line on termen from apex to
about vein 1b; hind wing of female pale smoky fuscous
throughout. Alar expanse, 38-42 mm.
Male genitalia with base of apical process of gnathos
nearly square when viewed from beneath. Female
genitalia not specifically different from those of cac-
torum except for a somewhat shorter ductus bursae.
Type Locatity: Arequipa, Peri (type in USNM).
Foop piant: Opuntia (Cylindropuntia) exaltata Berger
(apparently does not attack the Platypuntias).
The species is known only from the type series from
the type locality.
It is easily recognized by the color of the hind wings
and the ocherous suffusion on the forewings.
247
511. Cactoblastis bucyrus Dyar
Fiaures 538, 1026
Cactoblastis bucyrus Dyar, Ins. Insc. Menstr., vol. 10, p. 16, 1922;
Proc. Ent. Soc. Washington, vol. 30, p. 135, 1928.—Heinrich,
Proce. U. S. Nat. Mus., vol. 86, p. 360, 1939.—Dodd, Bio-
logical campaign against prickly-pear, Brisbane, Australia,
p. 39, 1940.
The male is much darker than the males of other
species of Cactoblastis. Palpi and thorax heavily dusted
with blackish scales. Head and collar ocherous. Fore-
wing brownish fuscous; white dusting inconspicuous,
the pale scales more ocherous than white in the male,
in female more whitish than ocherous and a trifle more
noticeable; black antemedial and subterminal lines
somewhat obscured by the dark ground color, conspicu-
ous only toward costa, dentations of subterminal line
as in doddi; a row of seven black dots along termen at
vein ends. Hind wing brownish fuscous in both sexes.
Alar expanse, 30-41 mm.
Male genitalia with arms of anellus somewhat shorter
than those of cactorum, doddi, and mundelli. Female
genitalia with bursa very minutely and sparsely scobin-
ate. The signum, like that of other species of Cacto-
blastis, is individually variable. Extremes of variation
are shown in figures 1026 and 1027.
Typr tocauity: Mendoza, Argentina
USNM).
Foop piants: Trichocereus, Echinopsis, Denmoza.
Distripution: ARGENTINA: Tucumén, Tapia, Men-
doza, Catamarca, Andalgalé.
(type in
141. Genus Cahela Heinrich
Cahela Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 361, 1939.
(Type of genus: Olyca ponderosella Barnes and McDun-
nough.)
Tongue short but stout. Antennae of male and fe-
male pubescent, the pubescence shorter in the female.
Labial palpus of male obliquely upturned, of female
porrect, with third segment downcurved and second and
third segments longer than those of male; male palpus
not extending above middle of front and with third
segment short. Maxillary palpus squamous. Hind
wing with veins 7 and 8 shortly anastomosing beyond
cell; 3 and 5 stalked. Eighth abdominal segment of
male simple.
Male genitalia with apical process of gnathos fused;
harpe with apex evenly rounded; vinculum short;
anellus with base of plate narrowly sclerotized, arms
long and broad, slightly twisted; aedeagus stout,
sclerotized throughout.
Female genitalia with signum developed as a ridged
plate (a hollow, blunt, flattened, more or less thornlike
projection into the bursa) ; bursa copulatrix large, finely
scobinate expecially in the neighborhood of the signum;
ductus bursae scobinate at genital opening; ductus
seminalis from bursa near signum.
Larvae whitish, not banded or conspicuously spotted;
solitary in habit; stem borers in Cylindropuntias.
Eggs laid singly.
248 UNITED STATES NATIONAL MUSEUM BULLETIN 207
This genus and the following (Rumatha) are dis-
tinguished from all others in the cactus-feeding group
by the complete fusion of the apical process of the
gnathos. Several male characters distinguish the two
genera from each other, but Cahela is most easily recog-
nized by the black longitudinal lines between the veins
on the forewing.
The genus is apparently limited in distribution to the
southwestern part of the United States and northern
México.
512, Cahela ponderosella (Barnes and McDunnough)
Figures 539, 1027
Olyca ponderosella Barnes and MecDunnough, Contributions, vol.
4, No. 2, p. 175, 1918.
Zophodia purgatoria Dyar, Ins. Insc. Menstr., vol. 13, p. 222, 1925.
Cactobrosis interstitialis Dyar, Ins. Insc. Menstr., vol. 13, p. 223,
1925; Proc. Ent. Soc. Washington, vol. 30, p. 1386, 1928.
Cactobrosis phoenicis Dvyar, Ins. Insc. Menstr., vol. 18, p. 223,
1925; Proc. Ent. Soc. Washington, vol. 30, p. 136, 1938.
Cactobrosis (?) ponderosella (Barnes and McDunnough) Dyar,
Proc. Ent. Soc. Washington, vol. 30, p. 136, 1928.—Mc-
Dunnough, Check list, No. 6284, 1939.
Cahela ponderosella (Barnes and McDunnough) Heinrich, Proc.
U.S. Nat. Mus., vol. 86, p. 361, 1939.
Head, thorax, forewings, and body dark fuscous-gray
peppered with white and with pronounced, longitudinal,
black lines on the forewing; a long black line through
the cell and extending from near base of wing to termen;
another long black line from base to tornus running
parallel and very close to the fold; in outer area, from
beyond cell to apex and termen, five other shorter black
lines, the largest and most pronounced above vein 6;
all the black lines between and not on the veins; ante-
medial and subterminal lines normally obsolete; in very
few specimens a faint indication of a partial, black,
antemedial line and in several specimens a dark shade
from end of cell to middle of inner margin, but no
trace of any transverse subterminal line; body some-
what paler than forewing or thorax. Hind wing white,
semihyaline, termen for a short distance from costa very
faintly and narrowly edged with fuscous; a more pro-
nounced fuscous shading along termen on the female.
Alar expanse, 30-42 mm.
Male genitalia show some variation in the shape and
size of the terminal process of the gnathos in different
specimens from any given locality. The extremes of
variation are shown in the figure. In the female geni-
talia the size of the signum and bursa varies somewhat
in different specimens but the variations are slight and
can be found in any series from one locality.
TYPE LocaLities: Palm Springs, Calif. (ponderosella
and phoenicis, in USNM); Colorado Desert, Yuma
County, Ariz. (purgatoria, in USNM); Presidio, Tex.
(nterstitialis, in USNM).
Foop pPuant: Opuntia (Cylindropuntia) imbricata
(Haworth) and probably other Cylindropuntias.
Distrizution: California, Palm Springs (Apr., Aug.),
San Bernardino (Apr., May); Utah, St. George (May,
June); Nevada, Charlestown Mts. (July), Clark County
(Apr., May, June); Arizona, Yuma County, Mohave
County (Apr., May), Dewey (June), “‘en route from
Dewey to Salome” (Apr.), Maricopa County (July),
Prescott (Apr., June), Redington, Baboquivari Mts.
(Pima County, May, July, Aug.), Phoenix (May),
Tucson: (June), Douglas (May), Christmas (Gila
County), Paradise (Cochise County, June); Texas,
Brewster County, Alpine (Apr.), Presidio.
Dodd states that the range of the species includes
the central plateau of México, which is what we should
expect. JI have seen no specimens from México.
The species is remarkably uniform in color and mark-
ing but varies considerably in size, which accounts for
some of the synonymy. Superficially ponderosella is
similar to both Eremberga leuconips (Dyar) and Cacto-
brosis strigalis (Barnes and McDunnough). They also
are dark gray with conspicuous black longitudinal lines
on their forewings, but in leuconips and strigalis the
black lines are on the veins, while in ponderosella they
lie between the veins. This character at once dis-
tinguishes it from other known cactus-feeding phycitids.
142. Genus Rumatha Heinrich
Rumatha Heinrich, Proc. U.S. Nat. Mus., vol. 86, p. 363, 1939.
(Type of genus: Zophodia bihinda Dyar.)
Tongue more or less developed (very short in glauca-
tella, but not hidden by palpi). Antenna of male
shortly serrate and pubescent, of the female simple and
pubescent, the pubescence shorter in the female than in
the male. Labial palpi porrect in both sexes; third
segment of palpus about half as long as second. Max-
illary palpus squamous. Hind wing with veins 7 and 8
anastomosing beyond cell; 3 and 5 stalked. Highth
abdominal segment of male simple.
Male genitalia with apical process of gnathos fused;
harpe with apex evenly rounded and with a subbasal
sclerotized pocket (pkt., fig. 540) between sacculus and
costa; uncus truncate and short in proportion to its
breadth; vinculum short, truncate, almost square in
outline; anellus with base rather broadly sclerotized,
arms short, broad, slightly twisted; aedeagus very short,
stout, partially sclerotized (on ventral half only). The
entire genitalia have a squat appearance that is charac-
teristic.
Female genitalia with signum developed as a ridged
plate with inwardly projecting ridge bluntly serrate
(except in glaucatella, in which the signum is as in
Cahela ponderosella); bursa copulatrix large, finely
scobinate, especially in neighborhood of signum; ductus
bursae scobinate at genital opening and with a pair of
more or less defined sclerotized plates on the dorsal
membrane of the ductus at the opening; ductus semi-
nalis from bursa near signum (but somewhat farther
removed than in Cahela).
Larva of only one species (glaucatella) known; whit-
ish, not banded or conspicuously spotted; solitary im
habit; stem borer in Cylindropuntia.
Eggs laid singly.
This genus is very close to Cahela. Both genera have
similar larvae and host associations and a like structure
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
of the gnathos; but there are too many other differences
in male characters and adult habitus to permit their
lumping. The partially sclerotized aedeagus, the short
stout arms of the anellus, the squat appearance of the
whole male genitalia, the porrect male labial palpi, and
the serrate male antennae at once distinguish the males
of Rumatha from those of Cahela; and the wing patterns
readily separate both sexes. In Rumatha the discal
dot is prominent and the transverse lines on the fore-
wing are well defined for at least half their length. In
Cahela the distinctive wing markings are longitudinal.
Three species are recognized as belonging to the
genus. Its distribution is limited apparently to the
southwestern part of the United States and possibly the
adjacent regions of northern México, although as yet
no specimens have been received from México.
513. Rumatha glaucatella (Hulst)
Fieures 542, 1028
Honora glaucatella Hulst, Ent. Amer., vol. 4, p. 117, 1888.
Zophodia glaucatella (Hulst) Phycitidae of North Amer., p. 174,
1890; U. S. Nat. Mus. Bull. 52, p. 4380, 1903.—Ragonot,
Monograph, pt. 2, p. 23, 1901—McDunnough Check list,
No. 6311, 1939.
Rumatha glaucatella (Hulst) Heinrich, Proc. U. 8. Nat. Mus.,
vol. 86, p. 365, 1939.
Palpi, head, and thorax pale fuscous, sparsely
sprinkled with white; posterior margin of thorax edged
with blackish fuscous. Forewing dull white, sparsely
sprinkled with fuscous and with a very pale fuscous
stain in a broad area bordering inner margin; ante-
medial line angulate, fuscous, rather faint but complete
and always distinguishable; subterminal line double,
consisting of two parallel, faint, pale fuscous lines,
almost vertical and but very slightly dentate; discal
spot at end of cell blackish fuscous, prominent; a row of
small blackish dots along termen between the vein ends.
Hind wing whitish with a very pale fuscous line edging
termen. Alar expanse, 15-20 mm.
Male genitalia much smaller than those of either
bihinda or polingella; basal portion of aedeagus narrower
in proportion; harpe with apex more bluntly rounded
than that of polingella but with width of harpe less in
proportion to its length than that of bihinda.
Female genitalia with signum similar to that of
Cahela ponderosella, the inwardly projecting edge not
appreciably serrate; sclerotized plates in genital open-
ing very weak, hardly distinguishable except under
very high magnification.
Larvae solitary in habit, white, not banded or conspic-
uously spotted.
TypE Locatity: Texas (type in AMNH, ex Rutgers).
Foop prant: Opuntia (Cylindropuntia) leptocaulis De
Candolle.
Distrisution: Texas, San Benito (May, June, July,
Aug.), Brownsville (June), San Diego (May), Laredo
(July), San Antonio; Florida (one female, so labeled and
without other locality, from the Fernald Collection, in
USNM).
The labial palpus of the male is somewhat misleading.
3003293—56——17
249
In natural position the third segment is projected
forward, but in relaxed and badly prepared specimens
it may be bent upward.
514. Rumatha bihinda (Dyar)
Figures 540, 1030
Zophodia bihinda Dyar, Ins. Insc. Menstr., vol. 10, p. 173, 1922.
Eumysia bihinda (Dyar), Ins. Insc. Menstr., vol. 13, p. 221,
1925.—McDunnough, Check list No. 6301, 1939.
Rumatha bihinda (Dyar) Heinrich, Proc. U. 8S. Nat. Mus., vol. 86,
p. 366, 1939.
Palpi, head, thorax, forewing, and abdomen dark
fuscous, dusted with white, giving a decidedly grayish
fuscous appearance to the moth; the white dusting
heavy on costal half of forewing and upper surface of
abdomen; discal spots and transverse markings on
forewings blackish fuscous. Forewing with area be-
tween cell and inner margin brownish, with little or no
white dusting and with transverse lines obscured; costal
half (especially above cell) strongly suffused with white;
transverse antemedial line blackish, distinct only from
costa to fold; subterminal line markedly dentate and
sinuate, blackish, oblique, broad and conspicuous from
costa to vein 8, with a slight dentation between veins
8 and 6 and a deep angulation between veins 5 and 6
extending to cell, between vein 5 and the fold straight
and inwardly slanting, thence obscure to inner margin
of forewing; discal dots at end of cell normally con-
spicuous and fused into a single black spot, obscure in a
few specimens; a row of black dots along termen at the
vein ends; in some specimens faint traces of a black
longitudinal line through center of cell and a line of
blackish scales along the fold. Hind wing white,
semihyaline, with a fine, faint, fuscous line along
termen and some fuscous shading on costal margin.
Undersurface of abdomen decidedly brownish fuscous,
sparsely dusted with white. Legs with femora whitish,
with some fuscous spotting; coxae uniformly dark
brown, with no white dustings or markings, strongly
contrasted against femora. Alar expanse, 30-36 mm.
Male genitalia appreciably larger than those of other
species in the genus; harpe broader in proportion to its
length and with apex more broadly rounded. Female
genitalia similar to those of polingella and hardly to be
distinguished; signum with inner projecting edge irregu-
larly and bluntly serrate.
Typr Locauity: Jemez Springs, N. Mex. (type in
USNM).
Foop piant: Unknown.
Distrrisution: Zeras, Alpine (Apr.); New Mezico,
Jemez Springs (June, July); Arizona, Yuma County
(Apr.), “en route from Dewey to Salome’ (Apr.),
Dewey (May), Mohave County (Mar.); Nevada, Clark
County (Mar., Apr., May), Bellevue (Washington
County, May).
This species has never been reared and its larva is
unknown. From its close relationship to glaucatella we
may expect that its host will prove to be one of the
Cylindropuntias.
250
515. Rumatha polingella (Dyar)
Fieures 541, 1029
Zophodia polingella Dyar, Journ. New York Ent. Soc., vol. 14,
p- 31, 1806.—McDunnough, Check list, No. 6312, 1939.
Rumatha polingella (Dyar) Heinrich, Proc. U. 8. Nat. Mus., vol.
86, p. 367, 1939.
Similar in appearance to that of bihinda but with
transverse antemedial and subterminal lines more dis-
tinctly continued to inner margin of forewing; indenta-
tions of subterminal line not so deep as in bihinda and
not extending to cell; a scattering of pinkish scales
among the white scales on costal area of forewing. Alar
expanse, 23-35 mm.
Male genitalia similar to those of bihinda but with
harpe narrower, apex of harpe more acutely rounded,
and aedeagus slightly narrower in proportion to its
length. Female genitalia essentially like those of 67-
hinda except that the sclerotized plates in genital open-
ing are not so distinct and the signum is on the average
smaller.
TYPE Locatiry: Southern Arizona (type in USNM).
Foop puant: Opuntia (Cylindropuntia) leptocaulis
De Candolle.
Distrisution: Arizona, Douglas (June, Aug.), Red-
ington, Palmerlee, Paradise (Cochise County, July,
Sept.), Pinal Mts. (Apr.), Baboquivari Mts. (June, July,
Aug., Sept.), Santa Catalina Mts. (Aug.), ‘southern
Arizona”’ (Apr.); Texas, Presidio (Aug.).
The Texas record is from a small reared male, giving
us our first food plant record for the species.
143. Genus Yosemitia Ragonot
Yosemitia Ragonot, Monograph, pt. 2, p. 17, 1901.—Heinrich,
Proc. U.S. Nat. Mus., vol. 86, p. 368, 1939. (Type of genus:
Spermatopthora graciella Hulst.)
Yosemetia Hulst, U. S. Nat. Mus. Bull. 52, p. 429, 1903.—Dyar,
Ins. Insc. Menstr., vol. 18, p. 220, 1925. (Misspelling.)
Tongue well developed. Antenna of male weakly
serrate and pubescent, of female simple and shortly
pubescent. Labial palpi obliquely porrect. Maxillary
palpus fan shaped and held vertically to the face. Hind
wing with veins 7 and 8 anastomosing beyond cell;
veins 3 and 5 stalked. Highth abdominal segment of
male simple.
Male genitalia with apical process of gnathos bifid;
harpe with apex evenly rounded; vinculum moderately
long; anellus with arms broad, short, slightly twisted,
and base of plate broadly sclerotized; aedeagus short
and slender, sclerotized throughout.
Female genitalia with signum developed as a small,
shortly spined plate; ductus bursae short; bursa copu-
latrix finely scobinate, especially in area about signum;
ductus seminalis from bursa near junction of bursa and
ductus bursae.
Larvae bluish, dark, not banded, or conspicuously
spotted.
The larvae feed gregariously (sometimes singly) in
Echinocereus, Coryphantha, Homalocephala, and pre-
sumably also in Echinocactus and Neomamillaria.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Eggs laid singly.
This genus as here defined is distinguished by the fol-
lowing combination of characters: Male antenna ser-
rate and pubescent; labial palpi porrect in both sexes;
maxillary palpi fan shaped; male genitalia with vincu-
lum moderately long and rather narrow, apical process
of gnathos bifid, anellus small and stout with broad
short arms, aedeagus slender; eighth abdominal sezsment
of male simple; female genitalia with signum a small,
shortly spined plate, ductus bursae short and ductus
seminalis from bursa copulatrix near junction of bursa
and ductus bursae; larvae not banded and normally
gregarious.
The male genitalia have a characteristic habitus
which makes them easy to place generically; but the
differences between species are very slight and not alto-
gether trustworthy, hardly more than might be expected
within specific limits.
Four species are recognized as belonging to the genus.
Its distribution is the southwestern part of the United
States and México.
516. Yosemitia graciella (Hulst)
Figures 68, 547, 1033
Spermatopthora graciella Hulst, Ent. Amer., vol. 3, p. 184, 1887.
Zophodia graciella (Hulst), Phycitidae of N. Amer., p. 173, 1890.
Yosemitia graciella (Hulst) Ragonot, Monograph, pt. 2, p. 13,
1901.—Barnes and McDunnough, Contributions, vol. 3, p.
199, 1916.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 369,
1939.—MeDunnough, Check list, No. 6293, 1939.
Yosemeiza graciella (Hulst), U. S. Nat. Mus. Bull. 52, p. 429,
1908.—Dyar, Ins. Insc. Menstr., vol. 13, p. 220, 1925.
Forewing pale brownish fuscous dusted and streaked
with black and dusted with white scales; the white
scaling concentrated on and strongly whitening the
costal half of the wing; the black scaling thinly dusted
over the lower half of the wing (between cell and inner
margin) and outlining the veins; transverse lines incom-
plete, blackish; the antemedial line indicated only by a
transverse dash in the cell and a dot or very short streak
on inner margin; subterminal line prominent from costa
near apex to vein 8 (Sometimes to vein 6), inwardly
slanting, interrupted between veins 6 and 5, obscure
between vein 2 and inner margin; discal dots fused into
a line of black scales on discocellular vein; a row of
small black dots along termen between the vein ends.
Hind wing whitish to very pale smoky fuscous, faintly
darker at apex, along costa, and narrowly along termen;
cilia white with a very fine, pale fuscous, subbasal line.
Alar expanse, 25-30 mm.
Male genitalia very little different from those of other
species in the genus; the vinculum is not so broad as
that of fieldiella or so long as that of didactica; the anelli
of the several species seem to offer the best characters
for separating the species on genitalic characters; those
of graciella and longipennelia are much alike, but in
didactica the arms appear to be more sharply twisted
and in fieldiella the basal portion is more narrowly
sclerotized and the free arms, therefore, correspondingly
longer. These characters, however, may not be con-
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 251
stant in long series. Female genitalia with signum a
trifle smaller than that of any other Yosemitia, other-
wise not specifically distinguished.
Larvae dark, dull blue and solitary or gregarious in
habit, according to Dodd.
Type tLocauity: Blanco County, Tex.
AMNHG, ex Rutgers).
Foop puants: Echinocereus viridiflorus Engelmann,
E. polyacanthus Engelmann, and Coryphantha aggregata
(Engelmann).
Distrrsution: Colorado, Denver (July), and one
specimen with only the State designation; Nevada,
Clark County (Apr., May); California, San Bernardino
County (Apr.), Providence Mts. (May), one specimen
with only State designation (Apr.); Arizona, Yavapai
County, Ajo (Pima County, Mar.), Baboquivari Mts.
(May), White Mts. (June), Pinal Mts. (Apr.), Quijotoa
Mts, (June), Santa Rita Mts. (June), Sells Post Office
(Pima County, May), ‘‘en route from Dewey to Salome”
(Apr.), Mojave County (May), Roosevelt (June),
Phoenix (Mar., Apr.), Redington, no locality except
the State (2 specimens reared from Coryphantha aggre-
gata, June); New Mexico (Mar.); Texas (no specific
locality, one specimen reared from Echinocereus viridi-
florus; Apr.).
(type in
517. Yosemitia longipennella (Hulst)
Ficures 546, 1032
Zophodia longipennella Hulst, Ent. Amer., vol. 4, p. 118, 1888.
Zophodia graciella (Hulst, in part), Phycitidae of N. Amer., p.
173, 1890.
Yosemitia graciella (Hulst, in part) Ragonot, Monograph, pt. 2,
p. 13, 1901.
Yosemetia graciella longipennella (Hulst) Barnes and McDun-
nough, Check list of the Lepidoptera of Boreal America,
No. 5699, 1917.
Yosemetia longipennella (Hulst) Dyar, Ins. Insc. Menstr., vol. 13,
p. 220, 1925.
Yosemitia longipennella (Hulst) Heinrich, Proc. U. S. Nat. Mus.,
vol. 86, p. 370, 1939.—McDunnough, Check list, No. 6294,
1939.
Similar to graciella except black dusting on forewing
sparse, veins not or but very faintly outlined by black
scales, transverse subterminal line not interrupted
between veins 6 and 5, dentate. Alar expanse, 21-26
mm.
Genitalia similar to those of graciella but with signum
of female a trifle larger.
Larvae dark dull blue and gregarious in habit, accord-
ing to Dodd.
Tyrer Locauity: Texas (type in AMNH, ex Rutgers).
Foop piant: Homalocephala texensis (Hopffer). Ac-
cording to Dodd the larvae also feed in Neomamillaria.
Disrrisution: Texas, Uvalde (June), “Big Bend”
(Apr.), San Antonio (June), Van Horn (June), San
Diego (Apr.), San Benito (Apr., May).
518. Yosemitia fieldiella (Dyar)
Figure 544
Zophodia fieldiella Dyar, Ins. Inse. Menstr., vol. 1, p. 35, 1913.—
McDunnough, Check list, No. 6310, 1939.
Yosemitia fieldiella (Dyar) Heinrich, Proc. U. S. Nat. Mus.,
vol. 86, p. 371, 1939.
Forewing heavily dusted with white on costal half
and with some scattered white scales on remainder of
wing; area between inner margin and cell pale brownish;
antemedial line incomplete, distinct only from costa to
lower vein of cell; subterminal line complete but obscure
except for the blackish costal dash, dentate, the incurva-
tion between veins 6 and 5 shallow; from antemedial
line, where it meets the cell, a thin, obscure, curved,
blackish line extends to upper outer angle of cell;
discal dot small, obscure; on some specimens a few black
scales outlining vein 6; on termen a row of obscure
blackish dots lying between the vein ends. Hind
wing white, in female faintly tinted with smoky fuscous;
a very pale fuscous line along termen. Alar expanse,
22-25 mm.
Male genitalia with basal portion of anellus more
narrowly sclerotized and arms correspondingly longer
than in the other species of the genus; vinculum also
broader and shorter. Female genitalia similar to those
of longipennella, but signum somewhat larger.
Type Locauity: La Puerta Valley, Calif. (type in
USNM).
Foop piant: Unknown
Distrizution: California, La Puerta Valley (July);
Arizona, Catalina Springs (May).
519. Yosemitia didactica Dyar
Figure 545
Yosemitia didactica Dyar, Proc, U.S. Nat. Mus., vol. 47, p.
408, 1915.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p.
372, 1939.
Forewing heavily dusted with white on costal half;
a short blackish line on midcosta; lower half of wing
concolorous with thorax; antemedial line obscure,
incomplete; discal dot at outer end of cell distinct,
blackish; subterminal line complete, dentate, double for
a short distance from apex and thence outwardly
margined by a narrow pale shade, obscure toward inner
margin, parallel to termen; veins 5 to 9 very faintly
outlined by dark scaling; terminal row of dots almost
obsolete. Hind wing whitish, faintly smoke tinted,
somewhat darker toward apex and along termen; cilia
with a pale smoky subbasal line. Alar expanse,
22-23 mm.
Male genitalia with vinculum rather longer than that
of any other species in the genus; anellus with arms
bent about aedeagus. Female genitalia similar to
those of graciella.
Type Locauity: Tehuacin, México (type in USNM).
Foop piant: Unknown.
Distrisution: Mixico: Tehuacén (May, June),
Orizaba.
This species resembles graciella but is somewhat paler
and the male has slightly darker (smoky) hind wings.
It is at once distinguished by the dark line on the
midcosta of the forewing. Its life history is unknown.
252 UNITED STATES NATIONAL MUSEUM BULLETIN 207
144. Genus Tucumania Dyar
Tucumania Dyar, Ins. Insc. Menstr., vol. 13, p. 224, 1925.—
Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 373, 1939.
(Type of genus: Tucumania tapiacola Dyar.)
Tongue rather short, but stout (as im Melitara).
Antenna of male shortly serrate and pubescent, of
female simple and shortly pubescent. Labial palpus
of male upturned, reaching almost to level of top of
eye; of female porrect (the second segment oblique,
the third slightly downcurved). Maxillary palpus
squamous. Hind wing with veins 7 and 8 anastomos-
ing for a short distance beyond cell; 3 and 5 stalked.
Highth abdominal segment of male simple.
Male genitalia with apical process of gnathos bifid;
harpe with apex bluntly pointed or elliptically rounded;
vinculum moderately long (it is somewhat foreshortened
in fig. 543), broad; anellus with base of plate moderately
sclerotized, arms rather broad and long, slightly
twisted; aedeagus long, slender; penis weakly scobinate
toward outer extremity.
Female genitalia with signum a small ridged or
granulate plate; bursa copulatrix with some fine
scobinations in the area about signum; ductus seminalis
from bursa near junction of bursa and ductus bursaae.
Larva purplish or wine colored with sclerotized areas
about body tubercles dark brown and large; two setae in
group VII on abdominal segments 7 and 8.
The larvae are solitary feeders in the joints of Platy-
puntias.
Eggs laid singly.
This genus is distinguished from others having serrate
and pubescent male antennae and squamous maxillary
palpi by its host association, its upturned male palpi,
slender aedeagus, female genitalia with signum, and
with ductus seminalis from the bursa. It is nearest to
Eremberga, but that genus is broad-winged and has a
flat, more strongly sclerotized anellus, a scobinate
aedeagus, stouter male genitalia, no signum, and the
ductus seminalis coming from the ductus bursae. In
Tucumania the wings are long and rather narrow.
The known distribution is Argentina and Uruguay.
520. Tucumania tapiacola Dyar
Ficures 65, 5438, 1034
Tucumania tapiacola Dyar, Ins. Insc. Menstr., vol. 13, p. 225,
1925.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 374,
1939.—Dodd, Biological campaign against prickly-pear,
Brisbane, Australia, pp. 39, 59, 83, 1940.
Palpi, face, head, thorax, and forewing dark grayish
fuscous with a sparse scattering of obscure whitish
scales (Dyar states that the coxae and parts of the
femora and tibiae of the legs are black, but even on
these parts there is some scattered pale scaling and the
ground color is fuscous rather than black). Forewing
almost uniformly dark, sometimes a very faint luteous
tint in the median area and a slight pale suffusion in
terminal area; transverse lines black but not strongly
contrasted against the dark ground color; antemedia
line bidentate, its apex extending almost to center of
cell; subterminal line dentate, sinuate, the dentations
short, bordered outwardly by a pale line and beyond
this by a rather broad blackish band, from costa well
before apex; discal spot at end of cell large; veins beyond
cell faintly outlined by dark scaling; a row of black dots
along termen at the vein ends. Hind wing whitish,
semihyaline, strongly shaded with fuscous at apex and
narrowly along margin of termen almost to anal angle,
this fuscous shading more extended on the female. Alar
expanse, 27-30 mm.
Male genitalia with apex of harpe bluntly pointed;
anellus with the apices of the arms appreciably broad-
ened. These are presumably specific characters. I
have seen no males of any other species of Tucumama.
Female genitalia with scobinations of bursa very weak
and distinguishable only in area surrounding signum;
signum somewhat granulate.
TyrE Locatiry: Tapia, Tucum4n, Argentina (type in
USNM).
Foop piants: Opuntia (Platypuntia) discolor Britton
and Rose, O. (Platypuntia) aurantiaca Lindley.
DistRIBUTION: ARGENTINA.
Only three specimens are before me, the male type
and a pair (o' and @) reared in Australia from Argen-
tine stock and sent me by Dodd.
521. Tucumania porrecta Dyar
Ficuré 1035
Tucumania porrecta Dyar, Ins. Insc. Menstr., vol. 13, p. 225,
1925.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 375, 1939.
Large and paler than fapiacola. ‘Thorax pale fawn
color. Forewing pale purplish fuscous with black
markings diffused; antemedial and subterminal lines
narrow, black, irregularly dentate, distinguishable
throughout but somewhat interrupted; discal dots at
end of cell rather large but not sharply contrasted
against ground color of the wing because of scattered
black dusting in the surrounding area; dots along termen
distinct; a short black streak from base through middle
of cell to apex of angulate antemedial line. Hind wing
white, faintly smoke tinted, especially toward apex.
Legs pale purplish fuscous; femora and tibiae trans-
versely banded with blackish fuscous on outer sides.
Alar expanse, 32-35 mm.
Female genitalia with scobinations of bursa very fine
but denser than in tapiacola; a small patch of somewhat
larger scobinations in neck of bursa; signum larger, with
a thin even keel but no granulations.
Typr Locauity: Paysandi, Uruguay
USNM).
Foop prant: Opuntia (Platypuntia) sp.
Distrisution: Urueuay.
Represented in the National Collection only by the
type and paratype from the type locality (Dodd, Feb.
1925), both females. The male is unknown.
(type in
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 253
145. Genus Eremberga Heinrich
Eremberga Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 375, 1939.
(Type of genus: Cactobrosis leuconips Dyar.)
Tongue reduced (small and weak). Antenna of male
serrate and pubescent, of female simple and shortly
pubescent. Labial palpus of male upturned, of female
obliquely porrect. Maxillary palpus squamous. Hind
wing with veins 7 and 8 very shortly anastomosed be-
yond cell: 3 and 5 stalked. Highth abdominal segment
of male simple.
Male genitalia with apical process of gnathos bifid;
harpe with apex evenly rounded; vinculum broad and
short; anellus with base of plate broadly and strongly
sclerotized, arms short, broad, not twisted or bent and
with apices pointed; aedeagus moderately long, rather
slender, sclerotized throughout and with a minutely
scobinate flange at apex.
Female genitalia without signum; bursa copulatrix
smooth or with a few scattered microscopic scobina-
tions; ductus bursae short, scobinate at genital open-
ing; ductus seminalis from ductus bursae.
Larva white with dark spots forming incomplete
crossbands; two setae in group VII on abdominal seg-
ments 7 and 8.
The larvae are solitary or semigregarious feeders in
Echinocereus. The larva of only one species (leuconips)
is known but the characters here given presumably
apply to the genus.
Eggs laid singly.
This genus is close to Tucumania and has many
characters in common with Olyca. The latter, however,
has veins 3 and 5 of the hind wing connate, the ductus
seminalis from the bursa rather than from the ductus
bursae, the male labial palpus oblique, the aedeagus
stout, and the basal plate of the anellus narrowly
sclerotized. The characters separating Hremberga from
Tucumania have been discussed in connection with the
latter genus.
Three species are here recognized as belonging to
Eremberga. Its distribution is the southwestern part
of the United States and México.
522. Eremberga leuconips (Dyar)
Fiaures 548, 1031
Cactobrosis leuconips Dyar, Ins. Inse. Menstr., vol. 13, p. 224,
1925; Proc. Ent. Soc. Washington, vol. 30, p. 136, 1928.—
McDunnough, Check list, No. 6288, 1939.
Eremberga leuconips (Dyar) Heinrich, Proc. U. 8. Nat. Mus.,
vol. 86, p. 376, 1939.
Palpi, head, thorax, and forewing grayish fuscous
densely sprinkled with white, giving the insect a pale
slate color. Labial palpus banded with blackish fuscous
toward the ends of the segments. Forewing with vein
3 to 10 outlined in black, the black scaling especially
strong on lower vein of cell; antemedial and subterminal
lines very fine and faint but usually discernible, black;
antemedial line acutely angulate and irregularly sinuate
and dentate, more or less broken and normally obliter-
ated at costa; subterminal line also irregularly sinuate
and dentate, decidedly slanting, obscured toward costa;
no distinct discal marks at end of cell and no dots along
termen. Hind wing of male glistening white, semi-
hyaline with a band of fuscous shading along costa and
a fine, pale fuscous line along termen for a short distance
from apex; of female, dark smoky fuscous, the fuscous
shading extending into the cilia and strongly outlining
most of the veins, and the wing paler towards base.
Alar expanse, 26-27 mm.
Male genitalia with lateral edges of anellus finely and
irregularly serrate; vinculum with terminal margin
evenly rounded, lateral margins not concave or exca-
vate. Female genitalia without any trace of signum;
bursa nearly smooth; ductus seminalis from ductus
bursae a short distance from genital opening.
Typr Locauity: Baboquivari Mts., Ariz. (type in
USNM).
Foop piant: Echinocereus polyacanthus Engelmann.
Distrisution: Arizona, Baboquivari Mts. (July,
Sept.), Roosevelt (July), Oracle (July), Huachuca Mts.
(Aug.), Chiricahua Mts., Mohave County (Sept.).
This species bears a strong resemblance to Cactobrosis
strigalis (Barnes and McDunnough) and might easily
be confused with it. The latter, however, has filiform
maxillary palpi while those of leuconips are distinctly
squamous. Dyar seems to have overlooked this char-
acter in placing many of his species. Also there is a
difference in the longitudinal markings. In sétrigalis
the strongest black longitudinal line is that along the
top of the cell and vein 6, while in leuconips the strongest
line is that along the lower vein of the cell.
In two males and some of the females of lewconips
there is a faint brownish fuscous suffusion on the lower
third of the forewing (bordering the inner margin), but
this is not distinguishable on all specimens and does
not seem to be a specific character.
523. Eremberga creabates (Dyar)
Fiaure 550
Olyca creabates Dyar, Ins. Inst. Menstr., vol. 11, p. 29, 1923.
Cactobrosis creabates (Dyar), Proc. Ent. Soc. Washington, vol. 30,
p. 136, 1928—McDunnough, Check list, No. 6289, 1939.
Eremberga creabates (Dyar) Heinrich, Proc. U. 8. Nat. Mus., vol.
86, p. 377, 1939.
Palpi grayish fuscous. Head grayish fuscous shaded
with white. Thorax luteous, whitish toward anterior
margin. Forewing with basal area (to antemedial line),
and all the area between antemedial and subterminal
transverse dark shade, and the fold and costa, white;
area between fold and inner margin luteous; outer area
(beyond subterminal dark shade) ashy white, shading
to luteous at tornus; transverse antemedial line well
contrasted against ground color, thin black, irrorate,
forming a sharp angle at the fold, the apex of the angle
extending almost to the middle of the fold; subterminal
line obsolete, replaced by a dark, transverse shade be-
low end of cell and some scattered blackish dusting
toward apex; veins 2 to 10 and upper and lower veins of
cell more or less outlined in black, the lines very faint
254
on all the veins except vein 4; along termen, between the
vein ends, a row of very faint blackish dots; no discal
marks at end of cell, Hind wing shiny white, semihya-
line, with a faint pale fuscous shading along costa, on
veins 6, 7, and 8, and at extreme apex. Alar expanse,
34 mm.
Genitalia with lateral margins of anellus smooth;
vinculum with terminal margin straight, rather broad,
lateral margins excavate.
Typr LocaLiry: San Diego, Calif. (July; type in
USNM).
Foop prant: Unknown.
Known only from the unique male type. It is a
striking species and should be easily recognized from
the description and genitalic features.
524, Eremberga insignis Heinrich
Figure 549
Eremberga insignis Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p.
378, 1939.
Palpi, face, head, thorax, and forewing dark grayish
fuscous. Forewing very faintly dusted with white on
costal half; lower half of wing faintly shaded with dull
luteous ocherous; antemedial and subterminal lines as
in leuconips, except antemedial not obliterated toward
costa; veins 2 to 9 very faintly outlined in black, the
black lining most distinct on lower vein of cell; a con-
spicuous black spot at end of cell; along termen, be-
tween the vein ends, a row of rather conspicuous black
dots. Hind wing shiny white, semihyaline, with a
fuscous shade bordering costa and a pale fuscous line
on termen for a short distance from apex. Alar expanse,
35 mm.
Male genitalia with lateral margins of anellus smooth;
vinculum with terminal margin straight and narrow,
lateral margins outwardly angled.
Typr LocaLity: San Luis Potosi, México (type in
USNM).
Foop piant: Unknown.
This species is known only from the male type. It is
easily distinguished from the other two species in the
genus by the conspicuous discal spot on forewing.
146. Genus Salambona Heinrich
Salambona Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 379, 1939.
(Type of genus: Zophodia analamprella Dyar.)
Tongue well developed. Antenna of male pubescent
and slightly serrate, of female simple and shortly pubes-
cent. Labial palpi of both sexes porrect with the third
segment downcurved, the third segment slightly longer
in the female than in the male. Maxillary palpus
squamous. Hind wing with veins 7 and 8 anastomosed
for more than one-half their lengths beyond the cell; 3
and 5 stalked. Eighth abdominal segment with a pair
of strong ventrolateral hair tufts.
Male genitalia with apical process of gnathos bifid,
small; uncus constricted toward apex; harpe with apex
oblique; vinculum long; anellus with base of plate nar-
UNITED STATES NATIONAL MUSEUM BULLETIN 207
rowly sclerotized, arms long, curved and twisted part
way around aedeagus; aedeagus long, stout.
Female genitalia without signum or scobinations in
bursa; bursa small, smooth; ductus bursae long, slender,
smooth; ductus seminalis from near end of bursa.
Larvae grayish green or blackish, according to Dodd;
not banded or conspicuously spotted; solitary feeders in
fruits of Platypuntia.
Eggs laid singly.
The genus is distinguished from other genera in the
cactus-feeding group by the following combination of
characters: Antenna of male serrate and pubescent;
labial palpi of both sexes porrect and downcurved;
maxillary palpus squamous; harpe of genitalia with
apex oblique; vinculum long; eighth abdominal segment
of male bearing a pair of ventrolateral tufts; bursa
copulatrix of female small and without signum or
scobonations (smooth); ductus seminalis from near end
of bursa; larvae unbanded, dark, fruit feeders in
Platypuntias.
Only the type species is recognized as belonging to
the genus. It is known only from Argentina.
525. Salambona analamprella (Dyar)
Fieures 552, 1045
Zophodia analamprella Dyar, Ins. Insc. Menstr., vol. 10, p. 17,
1922.
Salambona analamprella (Dyar) Heinrich, Proc. U. 8. Nat. Mus.,
vol. 86, p. 380, 1939.—Dodd, Biological campaign against
prickly-pear, Brisbane, Australia, p. 40, 1940.
Palpi, head, thorax, and forewings dark stone gray;
the scales under magnification dark grayish fuscous
tipped with dull white. Forewing with the costa
broadly margined (to top of cell) with white, the white
streak diminishing toward base of wing and terminating
before apex; no transverse lines, or discal or terminal
dots. Hind wing semihyaline with a smoky shade
along costa and a narrow smoke-brown line along
termen; the smoky shade somewhat more extended on
the female. Alar expanse 25-27 mm.
Male genitalia characters as given for the genus.
Female genitalia with bursa very small and ductus
bursae long and very slender.
TypE LocaLity: Carmen Patagones, Argentina (type
in USNM).
Foop pLant: Opuntia (Platypuntia) sulphurea G.
Don and probably other species of Platypuntia.
Distrisution: ARGENTINA: Carmen Patagones
(Jan.), Andalgalé (Mar.), La Rioja.
According to Dodd this insect is the chief enemy of
the cochineal scales in Argentina and is usually preda-
ceous on cochineal (Dactylopius spp.) but not un-
commonly the larvae feed in Opuntia fruit and flower
buds. From the genitalic and other structural charac-
ters of the moth I am inclined to doubt this. I think
analamprella will prove to be primarily a cactus feeder
and only secondarily predaceous on the cochineal scales
on the cactus. It is the other way around with Laetila
coccidivora (Comstock). The latter is a true predator
and follows its coccid hosts no matter to what plant
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
they may go. It also varies its diet somewhat by
occasional feedings on buds and flowers. According to
Dodd it sometimes feeds in Opuntia flowers; but this is
a secondary habit and the association with Opuntia
accidental. Laetilia is close to but not a part of the
cactus-feeding group of Phycitinae. Salambona, on the
other hand, is, in all adult characters, definitely a
member of the group.
The species is a striking one, easily recognized by the
strongly contrasted, white costal stripe on forewing.
147. Genus Parolyea Dyar
Parolyca Dyar, Proc. Ent. Soc. Washington, vol. 30, p. 17,
1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 381,
1939.—(Type of genus: Olyca asthenosoma Dyar.)
Tongue well developed. Antenna of male unipecti-
nate. Labial palpus of male upcurved. Maxillary
palpus squamous. Hind wing with veins 7 and 8
shortly anastomosed beyond cell; 8 and 5 shortly
stalked. Eighth abdominal segment with a strong
pair of ventrolateral tufts.
Male genitalia with apical portion of gnathos bifid;
uncus narrowed well before apex; harpe with apex
oblique; vinculum long, its terminal margin rounded,
its lateral margins excavate (probably a specific charac-
ter only); anellus with base of plate broadly sclerotized,
arms long, rather broad and slightly twisted; aedeagus
long, stout; penis weakly scobinate.
The genus is known only from the male of its type
species. Its biology is unknown, but from the genitalic
and other structural characters of the adult its larvae
are presumed to be cactus feeders. It is easily recog-
nized, for it is the only genus in the cactus-feeding
group with unipectinate antenna. The habitat is
French Guiana.
526. Parolyca asthenosoma (Dyar)
Figures 69, 551
Olyca asthenosoma Dyar, Ins. Insc. Menstr., vol. 7, p. 55, 1919.
Parolyca asthenosoma (Dyar), Proc. Ent. Soc. Washington, vol.
30, p. 137, 1929.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86,
p. 381, 1939.
Palpi, head, and thorax sordid white. Forewing
white, with a yellowish tint on area between fold and
inner margin; antemedial band angulate, consisting of
parallel black lines and a central white line; a black
oblique dash in median area from inner margin to dashes
at apex, black dots on the veins, and a black spot on
inner margin; a black discal dot at end of cell and some
black scaling on bases of veins 2 to 4; a row of small
black dots along termen, between the vein ends. Hind
wing white, semihyaline, with a pale fuscous shade
along costa, a narrow fuscous line along termen, and
some pale fuscous scaling on veins 2 to 8. Alar expanse,
30 mm.
Male genitalia with lateral margins of vinculum
excavate.
Type Locatity: Maroni River, French Guiana (type
in USNM).
255
Foop prant: Unknown.
Known only from the unique male type.
148. Genus Sigelgaita Heinrich
Sigelgaita Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 382, 1939.
(Type of genus: Sigelgaita chilensis Heinrich.)
Tongue well developed. Antenna of male bipectinate
(in transilis with a few flattened setae on the inner row
of pectinations of the first five or six segments of the
shaft); antenna of female shortly pubescent. Labial
palpus of male upcurved, of female porrect (the second
segment obliquely upturned, the third bent forward).
Maxillary palpus large, extending above front, flam-
boyant. Hind wing with veins 7 and 8 anastomosing
beyond cell; 3 and 5 stalked. Eighth abdominal seg-
ment with two pairs of thin hair tufts (very slight in
chilensis).
Male genitalia with apical process of gnathos small,
bifid; harpe with apex oblique; vinculum long; anellus
with base of plate somewhat broadly sclerotized, arms
moderately long, slightly twisted (in transilis) ; aedeagus
moderately stout, long; penis weakly scobinate.
Female genitalia with signum weak or absent; bursa
small and finely scobinate; ductus bursae moderately
long, finely scobinate toward bursa and genital opening;
ductus seminalis from middle of bursa.
Larva blue or blue-green according to Dodd; not
banded or conspicuously spotted; two setae in group
VII on abdominal segments 7 and 8.
The larvae are solitary feeders in the fruits of Zuly-
chnia, Trichocereus, and Platypuntia.
Egg and egg-laying habits unknown.
This genus is closest to Amalafrida but in many
characters more nearly resembles Nanaia. The max-
illary palpi are long in both Sigelgaita and Nanaia but
are not so closely appressed to the face in the former as
in the latter. The labial palpi of the males (upcurved
in Sigelgaita, porrect in Nanaia) readily separate the
two genera.
527. Sigelgaita chilensis Heinrich
Fieures 554, 1046
Sigelgaita chilensis Heinrich, Proc. U. 8S. Nat. Mus., vol. 86, p.
383, 1939.
Palpus, head, and thorax fuscous, strongly irrorated
with white; head and collar more whitish than fuscous;
posterior margin of thorax shaded with black. Fore-
wing fuscous, dusted with white, giving the wing an
ashy gray (in some specimens a bluish gray) color; a
white suffusion filling the cell; antemedial line near
middle of wing, black, outwardly angulate; from upper
angle of cell to middle of inner margin a more or less
prominent blackish shade; subterminal band dentate,
consisting of a thin, black, inner line, a parallel outer
black line, and a central pale line, the dentations of the
outer line acute and extended in short dashes onto the
veins; a row of black dots along termen between the
vein ends. Hind wing whitish, smoky fuscous toward
termen, apex, and costa, and on the veins; cilia white
256
with a pale fuscous subbasal line. Alar expanse, 31-44
mm.
Male genitalia with harpe fairly broad, aedeagus
somewhat stouter than in ¢ransilis, vinculum shorter.
Female genitalia with signum present, the latter con-
sisting of three or four minute, more or less coalesced,
blunt spines.
TypPE Locauity: Ovalle, Chile (type in USNM).
Foop piant: Hulychnia acida Philippi, Trichocereus
chiloensis (Colla).
Distrizution: Curie: La Serena, Ovalle, Tofa.
Superficially this species and huanucensis resembles
Nanaia substituta. The latter, however, lacks alto-
gether the dark shade between the outer angle of the
cell and the inner margin so characteristic of chilensis
and huanucensis. The forewings of the three species
are similar—long, and narrow and of about the same size
and shape.
528. Sigelgaita huanucensis Heinrich
Ficure 1047
Sigelgaita huanucensis Heinrich, Proc. U. S. Nat. Mus., vol. 86,
p. 383, 1939.
Similar to chilensis, except as follows: Paler, white
dusting on head, thorax, and forewing more pro-
nounced; general color of forewing brownish rather than
gray; transverse dark shade from outer upper angle of
cell to inner margin pale brown; antemedial and sub-
terminal lines interrupted, the latter indicated only by
blackish scaling on the veins; a pale brownish shade in
area bordering inner margin. Hind wing pure white,
with a very faint fuscous shade along costa and a
thin, pale fuscous line on termen for a short distance
from apex, this line a trifle broader on female than on
male and extending nearly to anal angle of wing. Alar
expanse, 45 mm.
Female genitalia without signum.
TypE LocaLity: Hudnuco, Peri (type in USNM).
Foop prant: Opuntia (Platypuntia) ficus-indica (Lin-
naeus).
Known only from the female type and male paratype
from the type locality.
529. Sigelgaita transilis Heinrich
Figure 553
Sigelgaita transilis Heinrich, Proc. U. S. Nat. Mus., vol. 86,
p. 384, 1939.
Palpi, head, thorax, and forewing dark grayish
fuscous finely peppered with white, giving them a slate-
gray color. Forewing with antemedial and subterminal
lines obscured, indicated by faint whitish lines bordered,
for a short distance from costa, by blackish streaks;
discal spot at end of cell blackish, rather large; a row of
black dots along termen between the vein ends. Alar
expanse, 26 mm.
Genitalia with harpe narrower than that of chilensis,
vinculum considerably longer, and aedeagus slenderer
and appreciably tapering toward apex.
TYPE Locatity: Santa Eulalia, Pera (type in USNM).
Foop puant: Trichocereus sp.
Known only from the male type.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
149. Genus Amalafrida Heinrich
Amalafrida Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 385, 1939.
(Type of genus: Cactoblastis leithella Dyar.)
Tongue well developed. Antenna of male bipec-
tinate; on each of the inner pectinations of the first five
segments a row of from three to five flattened, spinelike
setae; antenna of female simple and finely pubescent.
Labial palpus of male obliquely ascending, of female
obliquely porrect. Maxillary palpus squamous. Hind
wing with veins 7 and 8 anastomosing for over half
their lengths beyond cell; veins 3 and 5 stalked. Eighth
abdominal segment of male with two pairs of ventro-
lateral hair tufts.
Male genitalia with apical process of gnathos bifid;
harpe with apex oblique; vinculum long; anellus with
base of plate rather broadly sclerotized, arms long,
slightly twisted; aedeagus long, moderately stout;
Penis scobinate.
Female genitalia without signum; bursa copulatrix
large, weakly and scatteringly scobinate; ductus bursae
long, slender; ductus seminalis from about middle of
bursa.
Larva grayish in color with a tendency toward pale
transverse bands‘after the manner of Olycella larvae,
according to Dodd; solitary tunnelers in Platypuntia.
Eggs unknown.
The genus at present is represented by only the type
species. It is closest to Sigelgaita, one species of which
(transits) also has setiferous pectinations on some of
the basal segments of the male antennal shaft. The
form of the maxillary palpi, as well as the shape of the
forewings, distinguish the two genera. In Sigelgaita
the forewing is much longer in proportion to its width
and the termen more rounded than is the case in
Amalafrida. According to Dodd, leithella differs mark-
edly from the species of Sigelgaita in larval and pupal
habits.
530. Amalafrida leithella (Dyar)
Ficures 555, 1044
Cactoblastis leithella Dyar, Proc. Ent. Soc. Washington, vol. 30,
p. 135, 1928.
Amalafrida leithella (Dyar) Heinrich, Proc. U. S. Nat. Mus., vol.
86, p. 386, 1939.—Dodd, Biological campaign against
prickly-pear, Brisbane, Australia, p. 40, 1940.
Forewing with the areas between cell and costa, be-
tween vein 1b and inner margin for a short distance,
and along costal half of termen, white with a scattering
of black scales; ground color of remaining areas ocherous
fuscous, very faintly shaded with rufous above inner
margin; transverse and discal markings black; ante-
medial line incomplete, indicated by a thin, blackish,
irregular line from inner margin to cell and a broad
black streak from costa to about middle of the fold (in
some specimens this fuses with a black streak, which
extends from middle of vein 1b to end of cell) ; subter-
minal line black, faint (obscured below vein 6 in some
specimens), sinuate and dentate, outwardly bordered
by a whitish line and beyond this by a second, very
faint, parallel, pale fuscous line; at end of cell a large,
irregular, black spot; a line of distinct black dots along
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
termen between the vein ends. Hind wing of male
white, semihyaline, with a narrow pale fuscous shade
along costa and termen; of female dark smoky fuscous
shading to white towards base; cilia white with a
fuscous basal band. Alar expanse, 30-33 mm.
Male genitalia as given for the genus. Female geni-
talia with bursa very large and irregularly shaped,
minutely scobinate.
Type LocaLity: Curacao, Dutch West Indies (type
in USNM).
Foop piant: Opuntia (Platypuntia) sp.
Distrisution: Dutcu West Inpres: Curacao (Jan.).
VENEZUELA. Caracas (Jan.). Coxomsta: Province of
Colombia (Jan.)
Superficially leithella resembles Cactoblastis cactorum
but is easily distinguished on structural characters of
the male and female genitalia and of the male antennae.
150. Genus Ozamia Ragonot
Ozamia Ragonot, Monograph, pt. 2, p. 34, 1901.—Heinrich, Proc.
U. S. Nat. Mus., vol. 86, p. 386, 1939. (Type of genus:
Trachonitis lucidalis Walker.)
Antenna of male serrate (except in hemilutella and
punicans, where it is simple) and pubescent with a series
of modified, papillalike setae on the inner side of several
basal segments of the shaft (fig. 557d); antenna of the
female simple and pubescent. Labial palpi obliquely
ascending in both sexes. Maxillary palpus squamous.
Hind wing with veins 7 and 8 anastomosing beyond the
cell; 3 and 5 stalked. Eighth abdominal segment bear-
ing one pair or two (odiosella) pairs of ventrolateral
hair tufts.
Male genitalia with apex of gnathos small or moder-
ately large, bifid; apex of harpe oblique (except in
punicans); vinculum long; anellus with base of plate
broadly rather than narrowly sclerotized, arms long,
slightly twisted and curved; aedeagus rather long and
moderately stout (except in lucidalis); penis scobinate.
Female genitalia with signum weak or absent (luci-
dalis), when present developed as a thin, short, scobi-
nate or shortly thorned plate or a series or cluster of
small, weak spines; bursa copulatrix minutely scobinate,
at least toward ductus bursae (wrinkled in the South
American species); ductus bursae long or moderately
long, scobinate toward bursa; ductus seminalis from
bursa near signum.
Larva wine colored, olive green, or blackish, not
banded or conspicuously spotted; with two setae in
group VII of abdominal segments 7 and 8; solitary
feeders in fruits and flower buds of Opuntia and Cereus,
sometimes (some South American species) in the stems
of Cereus.
This genus divides into two natural groups: The
North American species with unwrinkled bursa and
minutely scobinate ductus bursae, and all fruit or bud
feeders; and the South American species with wrinkled
bursa and coarsely scobinate ductus bursae and either
fruit or stem feeders. The West Indian species
(lucidalis) is anomalous in some genitalic characters
300329—56——18
257
(small abdominal tufts, rather slender aedeagus, long
ductus bursae, and no signum), but on habitus and
other characters it appears closely allied to the North
American group. When males of all the species are
known it may be possible to give a separate generic
designation to the South American forms, but in the
absence of definitive male characters that does not seem
justified.
The papillalike setae on the male antennal shaft of
Ozamia also occur in Cactobrosis and Zophodia, but
the last two genera are distinguished by filiform maxil-
lary palpi.
Eight species are here recognized as belonging to the
genus. Its distribution appears to be the southwestern
part of the United States, Central and South America,
and the West Indies.
531. Ozamia lucidalis (Walker)
Ficures 556, 1042
Trachonitis lucidalis Walker, List, vol. 27, p. 39, 1863.
Ozamia lucidalis (Walker) Ragonot, Monograph, pt. 2, p. 34,
1801.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 388,
1939.—McDunnough, Check list, No. 6290, 1939.
Palpi, head, thorax, and forewing sordid white.
Forewing with ferruginous-fuscous spottings on the area
bordering inner margin; transverse markings black,
shading to ferruginous fuscous toward inner margin;
antemedial line angulate, white, bordered on inner and
outer sides by black or ferruginous, the outer black
marking at costa a broad spot; subterminal line dentate,
slanting from costa near apex to outer fourth of inner
margin, bordered inwardly and outwardly by dark lines,
shading from black to ferruginous; discal spot at end of
cell irregular, frequently extended beyond cell into two
short dashes, black; a row of black dots along termen
at the vein ends. Hind wing white, semihyaline, with
a fine fuscous line along termen; cilia white with a faint,
dark, subbasal line. Abdominal tufts small. Alar
expanse, 25-30 mm.
Male genitalia with apical process of gnathos small;
end of vinculum bluntly rounded. Female genitalia
without signum; bursa and part of ductus bursae
minutely scobinate; ductus bursae long, slender, bent
at middle.
TypPE Locatity: Santo Domingo [Dominican Repub-
lic] (type in BM).
Foop puant: Opuntia (Platypuntia) sp.
Distripution: Cusa; JAmMarca, Kingston (Jan.).
I have seen no specimens from the type locality.
532. Ozamia fuscomaculella (Wright)
Fiaures 557, 1043
Euzophera fuscomaculella Wright, Ent. News, vol. 27, p. 27,
1916.—MceDunnough, Check list, No. 6320, 1939.
Ozamia heliophila Dyar, Ins. Inse. Menstr., vol. 18, p. 222, 1925.
Ozamia odiosella fuscomaculella (Wright) Heinrich, Proc. U. S.
Nat. Mus., vol. 86, pl 390, 1939.
Ground color and markings of forewing similar to
those of lucidalis except that transverse markings are
blackish throughout, paling somewhat toward inner
258
margin but not shading into ferruginous; no ferruginous
coloring on forewing. Abdominal tufts much stronger
than in lucidalis and in two distinct pairs. Alar
expanse, 23-28 mm.
Male genitalia with apical process of gnathos rather
large; terminal margin of vinculum bluntly angulate.
Female genitalia with signum a thin short line of
minute spines; bursa copulatrix smooth except toward
ductus bursae, where it is finely scobinate; ductus
bursae of moderate length, swollen toward and gradually
expanding into bursa.
TYPE LOcALITIES: San Diego, Calif. (fuscomaculella,
in W. S. Wright Collection); Los Angeles, Calif. (helio-
pila, in USNM).
Foon pian: Opuntia (Platypuntia) spp.
Disrripution: California, San Diego (May, June,
Aug.), Los Angeles (July), Pasadena (Aug.).
533. Ozamia fuscomaculella clarefacta Dyar
Fieures 67, 559, 1040
Ozamia clarefacta, Dyar, Ins. Inse. Menstr., vol. 7, p. 55, 1919—
McDunnough, Check list, No. 6292, 1939.
Ozamia odiosella Heinrich (not Hulst), Proc. U. 8. Nat. Mus.,
vol. 86, p. 389, 1939.—Dodd, Biological campaign against
prickly-pear, Brisbane, Australia, pp. 39, 60, 84, 1940.
The variety (or race) is apparently limited to south-
eastern Texas and eastern México. It is distinguished
by the presence of a greenish tint over the area bordering
the inner margin of forewing. According to Dodd this
green shade is quite distinct on live or freshly killed
moths, but is not observable on most pinned examples.
There is also a slight difference in the female signum.
In clarefacta it consists of a narrow, minutely spined
plate. Otherwise clarefacta has nothing to distinguish
it from typical fuscomaculella.
TYPE LOCALITY: Orizaba, México (type in USNM).
Foop PLants: Opuntia (Platypuntia) spp.
Distrisution: Unirep Starrs: Texas, Brownsville,
Victoria (May), Burnet County (Oct.), Uvalde (June,
July), Kerrville (May, June), San Benito (Aug.).
Mexico: Orizaba (Apr.), Jalapa.
In my 1939 paper I misidentified this variety with
Hulst’s odiosella on the basis of a female from Texas in
the National Collection which Hulst had labeled ‘‘Saleb-
ria odiosella Hulst, Type” and which both Dyar and I
had supposed was the true type of that species. It is
not the actual type, which I had previously overlooked.
This is in the Rutgers Collection. It is a male, super-
ficially similar to the spurious ‘‘type”’ in the National
Museum but structurally quite distinct; it has 8 veins in
the hind wing and the maxillary palpus in the form of an
aigrette (as in Salebria where Hulst placed it). The
true odiosella is treated on page 114.
534. Ozamia thalassophila Dyar
Fiaure 1039
Ozamia thalassophila Dyar, Ins. Insc. Menstr., vol. 18, p. 15,
1925.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 391,
1939.—McDunnough, Check list, No. 6291, 1939.
Palpi, head, thorax, and forewing dark grayish
UNITED STATES NATIONAL MUSEUM BULLETIN 207
fuscous, lightly sprinkled with white. Forewing with
some white dusting on costal half and a slightly more
brownish shade on inner half; antemedial and subter-
minal transverse lines whitish, bordered with black or
blackish fuscous, the pattern as in lucidalis and odio-
sella; discal spot at end of cell curved, black; between
this and subterminal lines one or two small, obscure,
blackish dots; upper and lower veins of cell faintly out-
lined by white scales; a row of black dots along termen
at or close to the vein ends; cilia pale ocherous fuscous.
Hind wing white with a narrow fuscous shade along
termen; cilia white, with a fuscous subbasal line. Alar
expanse, 28 mm.
Female genitalia, with signum a small cluster of two
or three more or less fused and minute spines; bursa
copulatrix partially wrinkled (im the region of the sig-
num); ductus bursae of moderate length, finely sco-
binate toward bursa.
TYPE LocaLity: Oceanside, Calif. (type in USNM).
Foop piant: Opuntia (Cylindropuntia) sp.
Known only from the unique female type, reared
(Aug. 1924) from larva in a Cylindropuntia, presumably
feeding in the fruit.
535. Ozamia immorella (Dyar), new combination
Fieure 1036
Euzophera immorella Dyar, Proc. U. 8. Nat. Mus., vol. 44, p. 322,
1913.
Similar in color and markings to thalassophila except
for a faint purplish red suffusion over the ground color
of forewing (especially noticeable on freshly reared
examples) and less white dusting on costal area of wing.
Alar expanse, 25-31 mm.
Male genitalia with apical process of gnathos rather
large (as in fuscomaculella); terminal margin of vin-
culum bluntly rounded. Female genitalia with signum
a narrow, bluntly spined plate, the signum longer than
any other in the genus; bursa copulatrix somewhat
wrinkled in the neighborhood of signum, finely scobi-
nate towards ductus bursae, the scobinations extending
for a short distance into ductus. iq
Typr Locatity: Tehuacén, México (type in Oe
Foop PLant: Opuntia (Platypuntia) sp.
Distripution: México: Caxaxa, Tehuacin (uly),
Zacatecas (Dec.)
This species is very close to thalassophila but appar-
ently distinct. I overlooked it when treating the
cactus-feeding Phycitinae in 1939. Since that time a
series (including one male) was reared on Dec. 10, 1946,
by George Callaghan of the U.S. Bureau of Entomology
and Plant Quarantine from larvae feeding in the fruits
of prickly-pear at Zacatecas, México.
536. Ozamia stigmaferella Dyar
Fieure 1038
Ozamia stigmaferella Dyar, Ins. Insc. Menstr., vol. 10, p. 17,
1922.—Heinrich, Proc. U. S. Nat. Mus., vol. 86, p. 391,
1939.
Palpi, head, and thorax dark grayish fuscous sparsely
sprinkled with white, forewing dark grayish fuscous;
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
extreme base black; remainder of basal area and area
between cell and costa and to the subterminal line
white lightly dusted with black; antemedial line obso-
lete, indicated only by a large black spot on costa
(corresponding to the black shade outwardly bordering
the white antemedial line in the North American
species of Ozamia); subterminal line whitish, dentate,
bordered inwardly and outwardly by black; a short,
dentate, black line from vein 8 to vein 2, midway
between the end of cell and the subterminal line and
parallel with the latter; a black curved mark at end of
cell; a row of black dots along termen at or near the
vein ends; a faint whitish color dusted with black in
apical area. Hind wing white, semihyaline, with a
fuscous shade at apex and for a short distance along
termen; cilia whitish with a very faint fuscous subbasal
line. Alar expanse, 26 mm.
Female genitalia with signum a small buttonlike
thorn; bursa copulatrix wrinkled and finely scobinate;
ductus bursae long, coarsely scobinate toward bursa.
Type tocatity: Catamarca, Argentina (type in
USNM).
Foop piant: Cereus validus Haworth.
This species is known only from the female type,
reared Mar. 7, 1921, by W. B. Alexander from a larva
that had excavated a hollow in a stem of Cereus validus.
O. stigmaferelia and the two following species (hemilu-
tella and punicans) are the South American repre-
sentatives of the genus and differ from those from the
United States, México, and the West Indies in having
the ductus bursae of the female coarsely scobinate and
the bursa copulatrix decidedly wrinkled.
537. Ozamia hemilutella Dyar
Fiaures 560, 1041
Ozamia hemilutella Dyar, Ins. Insc. Menstr., vol. 10, p. 17,
Pdi ta Proc. U. S. Nat. Mus., vol. 86, p. 392,
1939,
Palpi, head, and thorax brownish fuscous, finely and
evenly sprinkled with white, making the general color
(to the naked eye) pale gray; collar of thorax with a
slight yellowish tint. Forewing with area between
lower vein of cell, vein 2, and inner margin pale yellow
without any markings; remainder of wing pale gray,
concolorous with head and thorax; antemedial line
obsolete; from costa just before middle to middle of
lower vein of cell, a rather broad, transverse, brownish
shade; a brown discal dot at end of cell and some brown
shading just beyond; subterminal line faint, weakly
dentate, parallel with termen, bordered inwardly and
outwardly by thin faintly brownish lines; a row of
minute black dots along termen at the vein ends; cilia
pale gray. Hind wing white, semihyaline with a thin
fuscous line along termen and some fuscous shading
bordering the costa. Abdomen with one pair of strong
tufts. Alar expanse, 27-30 mm.
Male genitalia with apical process of gnathos small;
end of vinculum bluntly angulate. Female genitalia
with signum a single, weak, irregular, thornlike patch;
259
bursa copulatrix wrinkled and finely scobinate; ductus
bursae coarsely scobinate toward bursa.
Type ocauity: La Rioja, Argentina (type in
USNM).
Foop piant: Cereus validus Haworth.
DistRIBUTION: ARGENTINA: La Rioja, Santiago del
Estro.
The moth is easily identified by the clear yellow inner
area of the forewing. The larvae feed in the fruits and
flower buds and, possibly, to some extent, in the stems
of Cereus. Dodd states that they also attack fruits of
Platypuntias.
538. Ozamia punicans Heinrich
Fieures 558, 1037
Ozamia punicans Heinrich, Proc. U. S. Nat, Mus., vol. 86, p. 393,
939.
Palpi fuscous sprinkled with white. Head and thorax
fuscous heavily dusted with white and more or less
shaded with pale rust color, especially on top of head
and on collar of thorax. Forewing pale gray (fuscous
heavily dusted with white) marked with darker gray
and with large blotches of pale rust color; the rust shade
filling about one-fourth of the basal area and nearly all
the area between antemedial and subterminal lines,
lower vein of cell, vein 2, and vein 1b; antemedial line
obscure, indicated chiefly by a rather broad dark gray
shade from costa to lower vein of cell and a thin dark
gray line thence to inner margin; subterminal line faint,
somewhat sinuate but not dentate, approximately paral-
lel with termen, bordered inwardly and outwardly by
obscure dark gray; apical mark at end of cell irregular,
dark gray; between cell and subterminal line some faint
rust shading in the interspaces between the veins; a row
of black dots along termen between the vein ends; cilia
pale rust red. Hind wing white, semihyaline, with a
fuscous shade in costal area to top of cell and vein 8,
some fuscous shading on the vein ends, and a fine fuscous
line along termen to vein 1b; cilia shiny white. Abdom-
inal tufts as in hemilutella. Alar expanse, 36-40 mm.
Male genitalia with apical process of gnathos moder-
ately large; apex of harpe more rounded than in other
species of Ozamia; end of vinculum more rounded than
angulate and lateral margins excavate; penis bearing a
number of coarse spines. Female genitalia larger than
those of any other Ozamia; with signum a small granu-
lose plate containing a stubby central thorn; bursa
copulatrix wrinkled and finely scobinate; ductus bursae
long, very coarsely scobinate toward bursa.
TyprE LocaLity: Tapia, Tucumdn, Argentina (type
in USNM).
Foop piant: Cereus validus Haworth.
According to Dodd, punicans differs from other
species of Ozamia in that it is a stem borer and appar-
ently does not attack the fruits or flower buds. It
differs also in that the apex of the harpe is not definitely
oblique, and the maxillary palpi are somewhat narrowly
scaled. However, the latter are of the squamous
rather than the filiform type, and from its general
habitus the species is obviously closely related to
260 UNITED STATES NATIONAL MUSEUM BULLETIN 207
hemilutella. The moth can be easily identified by the
rust-red cilia and blotches on the forewing. It, so far,
is known only from the type series from Tucumdén.
151. Genus Cactobrosis Dyar
Cactobrosis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 406, 1915;
Proc. Ent. Soc. Washington, vol. 30, p. 135, 1928.—Hein-
rich, Proc. U.S. Nat. Mus., vol. 86, p. 394, 1939.
Tongue well developed. Antenna of male with a
series of modified, papillalike setae on the inner sides of
several basal segments of the shaft, bipectinate (fer-
naldialis, longipennella) orstrongly serrate and pubescent
(maculifera, strigahis); antenna of female simple and
shortly pubescent. Labial palpus upturned in the
male, oblique in the female. Maxillary palpus filiform.
Hind wing with veins 7 and 8 anastomosing beyond
the cell; 3 and 5 shortly stalked. Highth abdominal
segment bearing a pair of ventrolateral hair tufts (the
tufts long and dense except in strigalis).
Male genitalia with apex of gnathos large, bifid; apex
of harpe evenly rounded; vinculum long (moderately
long in strigalis); anellus with base of plate narrowly
sclerotized, arms long, slender, slightly twisted ; aedeagus
long, stout (shorter and less stout in sfrigalis); penis
more or less densely pubescent (armed with short, hair-
like spines).
Female genitalia without sisnum; ductus bursae long,
finely scobinate only at genital opening or (in séfrigalis
only) sparsely so at junction of bursa copulatrix and
ductus bursae, with two small sclerotized dorsal plates
and a single ventral plate at genital opening (the ventral
plate absent in strigalis); bursa copulatrix large, smooth
(except in strigalis, in which it has a few minute scobina-
tions) ; ductus seminalis from near end of bursa.
Larvae bluish, not banded or conspicuously spotted;
with two setae in group VII on abdominal segments 7
and 8; gregarious feeders in Ferocactus, Echinocereus,
Peniocereus, and, probably, Carnegiea.
Eggs laid singly.
The genus as here defined is distinguished from all]
other genera of the cactus-feeding group by its filiform
maxillary palpi. Zophodia, which it resembles in most
structural characters, is not a cactus-feeding genus, has
the male antenna unserrate, the labial palpus of female
porrect, and a small signum in the bursa, copulatrix.
Five species are recognized as belonging to the genus.
They are fairly easy to distinguish but subject to much
individual variation in wing markings.
The known distribution is the southwestern part of
the United States and México.
539. Cactobrosis fernaldialis (Hulst)
Fiaures 73, 524, 1049
Megaphycis fernaldialis Hulst, Trans. Amer. Ent. Soc., vol. 13,
p. 163, 1886.
Euzophera gigantella Ragonot, Nouv. Gen., p. 32, 1888; Mono-
graph, pt. 2, p. 51, 1901.—Dyar, Proc. Ent. Soc. Washing-
ton, p. 158, 1904.
Melitara fernaldialis (Hulst), Phycitidae of N. Amer., p. 172,
1890; U. S. Nat. Mus. Bull. 52, p. 429, 1903.—Schwarz,
Psyche, vol. 8, p. 15, 1901.—Hunter, Pratt, and Mitchell,
U.S. Dept. Agr. Bur. Ent. Bull. 113, p. 29, 1912.
Honora cinerella Hulst, Journ. New York Ent. Soc., vol. 8,
p. 223, 1901; U. S. Nat. Mus. Bull. 52, p. 433, 1903.
Melitara fernaldalis Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 36, 1905 (misspelling for fernaldialis Hulst).
Cactobrosis fernaldalis (Dyar), Proc. U. S. Nat. Mus., vol. 47,
p- 407, 1914; Ins. Insc. Menstr., vol. 13, p. 223, 1925 (in
part); Proc. Ent. Soc. Washington, vol. 30, p. 135, 1928
(in part).
Cactobrosis fernaldialis (Hulst) Heinrich, Proc. U. S. Nat. Mus.,
vol. 86, p. 396, 1939—McDunnough, Check list, No.
6283, 1939.—Dodd, Biological campaign against prickly-
pear, Brisbane, Australia, p. 40, 1940.
Mats: Antenna bipectinate. Palpi, head, and thorax
grayish fuscous dusted with white. Forewing grayish
fuscous dusted with white and more or less blotched
with black; some specimens with a faint ocherous-
fuscous tint in the middle of the cell and on the area
between vein 1b and the cell; normally with antemedial
and subterminal transverse markings indistinct, but
indicated by whitish angulate and dentate bands shaded
inwardly and outwardly by black; a blackish shade at
end of cell, often extending to costa; below it on inner
margin a similar dark spot; veins 2 to 8 faintly lined
with black and in many specimens the fold to a little
beyond its middle. Hind wing white, semihyaline,
shaded in costal area above vein 6 and cell with pale
fuscous, with some fuscous scaling on the veins and a
fine fuscous line along termen; anal margin and adjoin-
ing cilia faintly ocherous; cilia otherwise white, with a
narrow, fuscous subbasal line. Alar expanse, 36—47
mm
Male genitalia essentially like those of longipennella
but somewhat larger, in size and habitus like those of
maculifera.
Fematz: In color and markings like the male except
that there is never any black streak on the fold of the
forewing; some specimens are heavily dusted with black
over the entire base of the forewing as far as the ante-
medial line; others have the transverse lines and con-
trasted dark spots almost obliterated and the wing of a
pale slate color with only the faintest remnants of the
normal markings. Alar expanse, 34-50 mm.
Female genitalia with the sclerotized ventral plate
in ductus bursae at opening smaller than those in
longipennella, insignatella, and maculifera.
TYPE Locauities: Arizona (fernaldialis, in AMNH,
ex Rutgers; gigantella, in Paris Mus.); Santa Rita Mts.,
Ariz. (cinerella, in USNM).
Foop piants: Ferocactus wislizent (Engelmann) and
probably other species of Ferocactus; Peniocereus greggu
(Engelmann) ; Carnegiea gigantea (Englemann) ; Homalo-
cephala texensis (Hopffer).
Distrisution: Arizona, Catalina Springs (Apr.),
Oracle (July), Tuscon (June), Baboquivari Mts. (Apr.,
May, June, July, Aug., Oct., Nov.), Christmas, Reding-
ton, Pinal Mts., Santa Rita Mts. (May, June), Hua-
chuca Mts. (Aug.), Douglas (Apr., May), Mohave
County (May), Sells Post Office (Indian Oasis, Apr.),
Dewey (June), Maricopa County (July), “en route from
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 261
Dewey to Salome’ (Apr., May); California, San Diego
(May, Oct.); Texas, Brownsville (Jan., U. S. Dep. Agr.
rearing).
540. Cactobrosis longipennella (Hampson)
Fiaures 523, 1050
Euzophera longipennella Hampson, in Ragonot, Monograph, pt.
2, p. 52, 1901.
Moodna elongatella Hampson, in Ragonot, Monograph, pt. 2, p.
269, 1901.
Cactobrosis longipennella (Hampson) Dyar, Proc. U.S. Nat. Mus.,
vol. 47, p. 407, 1914.—Heinrich, Proc. U. 8. Nat. Mus., vol.
86, p. 397, 1939.
Cactobrosis elongatella (Hampson) Dyar, Proc. U. 8. Nat. Mus.,
vol. 47, p. 407, 1914.
Cactobrosis fernaldialis Dyar (part), Ins. Insc. Menstr., vol. 13,
p. 223, 1925; Proc. Ent. Soc. Washington, vol. 30, p. 135,
1928.
Mats: Like that of fernaldialis except that pectina-
tions of antenna are about half the length of those on
fernaldialis; transverse pale markings on forewing obso-
lete in some specimens. Alar expanse, 34-40 mm.
Male genitalia similar to those of fernaldialis except
smaller; harpe not so markedly creased.
Frmate: Similar in color and markings to the female
of fernaldialis. Alar expanse, 33-43 mm.
Female genitalia with sclerotized ventral plate in
ductus bursae at genitalic opening larger and the oppos-
ing small plates on the dorsal wall of the ductus
narrower than those of fernaldialis.
Typr tocauities: Tres Marfas Isl., México (longi-
pennella, in BM); Orizaba, México (elongatella, in
USNM).
Foop puant: Unknown, probably Ferocactus.
Distrisution: México: Orizaba, Oaxaca, Tehuacén
(June), Cuernavaca (June, July), Zacualpin (Mar.,
Oct.).
I have seen no examples from the type locality of
longipennella.
541. Cactobrosis maculifera Dyar
Ficures 525, 1051
Cactobrosis maculifera Dyar, Proc. U. S. Nat. Mus., vol. 47, p.
407, 1914; Proc. Ent. Soc. Washington, vol. 30, p. 136,
1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 398, 1939.
Mate: Antenna strongly serrate and fasciculate.
Palpi, head, and thorax pale clay color (‘‘luteous’’).
Forewing luteous gray shaded and spotted with dark
grayish fuscous, the luteous tint pronounced on basal
third of costa and over the submedian area of the wing;
transverse antemedial and subterminal lines obsolete;
a fuscous shade from costa before middle to cell, another
from costa at middle, and, below these, corresponding
streaks or spots on lower vein of cell and on vein 1b;
a thin blackish line on the fold from its base to near its
middle; a similar dark streak on vein 1b at outer third;
short, broken, dark streaks on the veins at or near the
cell; a clouded fuscous spot at end of cell; outer half of
costa shaded with fuscous; a row of dark spots along
termen at or very close to the vein ends. Hind wing
white, semihyaline with only the faintest indication of
a fuscous line on termen toward apex. Alar expanse,
32-45 mm.
Male genitalia agreeing in size and nearly all details
with those of longipennella except that the arms of the
anellus are a trifle longer in maculifera.
Frmate: In color and pattern like the male except
that there is some gray shading on the head and thorax
and considerably more gray on the forewing (the single
specimen before me is in much better condition than
the males, which may account for some of the differ-
ences); basal third of wing clouded with dark fuscous;
terminal area more faintly clouded; subterminal line
faintly indicated, sharply angulate at middle, broken
below; the luteous shade more contrasted than in the
male, but restricted to middle of cell and the area be-
tween veins 1b and the fold. Hind wing white, semi-
hyaline with a narrow fuscous shade along termen and
on the veins near their apices. Alar expanse, 37 mm.
Female genitalia similar to those of insignatella but
with sclerotized ventral plate in ductus bursae at gen-
ital opening smaller.
TypE LocaLity: Oaxaca, México (type in USNM).
Foop prant: Unknown.
Distrisution: M*£xico:
(Sept.).
This species may be distinguished from other species
of Cactobrosis by the strong luteous (pale clay) shade
on the forewing and the serrate-fasciculate male
antenna.
Oaxaca, Salina Cruz
542. Cactobrosis insignatella Dyar
FicureE 1052
Cactobrosis insignatella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p.
407, 1914; Proc. Ent. Soc. Washington, vol. 30, p. 136,
1928.—Heinrich, Proc. U. 8. Nat. Mus., vol. 86, p. 399, 1939.
Mate: Unknown.
Frema.e: Palpi, head, thorax, and forewing of a soft,
nearly uniform grayish fuscous (with a more brownish
than slate-gray tint). Forewing without discal spot or
transverse dark markings; costa at base very slightly
paler than ground color of wing, concolorous with collar
of thorax; an obscure pale shade on midcosta and the
faintest indication of a pale subterminal line, the latter
broadly angulate at middle; some faint dark shading
on the veins from cell to termen and a row of small,
dark dots along termen near the vein ends. Hind wing
white, semihyaline, with a pale fuscous line along ter-
men; cilia white with a narrow, pale fuscous, subbasal
line. Alar expanse, 37-40 mm.
Genitalia with the dorsal plates in ductus bursae at
genital opening strongly sclerotized; ventral plate at
opening slightly larger than in any of the other species.
Type Locauity: Oaxaca, México (type in USNM).
Foop prant: Unknown.
Known only from the female type and paratype from
the type locality. These specimens resemble suffused
specimens of fernaldialis and longipennella except that
the latter are more slate colored. The slight genitalic
262
differences seem to indicate that insignatella is a good
species and not a mere color form.
543. Cactobrosis strigalis (Barnes and McDunnough)
Figures 526, 1053
Euzophera strigalis Barnes and McDunnough, Canadian Ent.,
vol. 44, p. 127, 1912; Contributions, vol. 1, No. 4, pl. 1,
fig. 14, 1912.
Caciobrosis strigalis (Barnes and McDunnough), Check list of
the Lepidoptera of Boreal America, No. 5697, 1917.—Dyar,
Ins. Inse. Menstr., vol. 13, p. 224, 1925; Proc. Ent. Soc.
Washington, vol. 30, p. 136, 1928.—Heinrich, Proc. U. 8.
Nat. Mus., vol. 86, p. 400, 1989.—McDunnough, Check
list, No. 6287, 1939.
Mats: Antenna strongly serrate. Palpi, head, thorax,
and forewing grayish fuscous sprinkled with whitish
(the ends of the scales white) making the ground color
a pale slate-gray. Forewing with the veins outlined in
black, the strongest black line being that along upper
vein of cell and vein 6; transverse lines and discal mark
absent; no dots along termen. Hind wing white, semi-
hyaline, with a faint fuscous shade bordering costa, and
a fine fuscous line along termen for a short distance from
apex. Tufts on eighth abdominal segment weak. Alar
expanse, 30-43 mm.
Male genitalia with vinculum moderately long, but
considerably shorter than in other species of Cactobrosis;
aedeagus also shorter.
Frmaue: Similar to the male in color and markings
except that hind wing is more or less suffused with
smoky fuscous beyond the base, especially along the
veins and termen. Alar expanse, 33-44 mm.
Female genitalia with sclerotized plates on dorsal wall
of ductus bursae behind the genital opening well de-
veloped, but with opposing ventral sclerotized plate
absent, replaced by minute scobinations; ductus bursae
shorter than in other Cactobrosis species; bursa copula-
trix not entirely smooth, having a few weak scobinations
toward ductus.
TypE LOCALITY: Hureka, Utah (type in USNM).
Foop piant: Echinocereus rigidissimus (Engelmann),
E. pectinatus (Scheidweiler), and probably a number of
other species of H-chinocereus.
Distripution: Unirep States: Utah, Eureka (Aug.,
Sept.). Dividend (Sept.); Cahfornia, San Gorgonio
Pass (July); Arizona, Tucson (Apr., July); Texas, Brew-
ster County (July, Aug.), Alpine (Apr.). Mexico:
Distrito Federal, México (National University, male
reared from £. pectinatus, June 3, 1931).
In a number of respects (its shorter vinculum and
ductus bursae, its weak abdominal tufts, and its par-
tially scobinate bursa copulatrix) this species fits badly
into Cactobrosis. Eventually it may need a separate
generic designation, but this had better be postponed
until the life histories of the other species of Cactobrosis
are more fully known.
The forewing markings of strigalis resemble those of
Euremberga leuconips (Dyar). The latter, however, is
easily distinguished by its squamous maxillary palpi.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Genus 152: Drescomopsis
[Venational division A. Forewing with 11 veins: 4 and 5 approxi-
mate or connate. Hind wing with vein 2 from the lower outer
angle of cell; discocellular vein curved. Male genitalia with
transtilla complete; harpe with base of sacculus produced and
bearing stout hair tuft. Labial palpus upturned.]
152. Genus Drescomopsis Dyar
Drescomopsis Dyar, Ins. Insc. Menstr., vol. 7, p. 61, 1919. (Type
of genus: Drescomopsis subelisa Dyar.)
Tongue well developed. Antenna pubescent; shaft
of male slightly swollen at base, otherwise simple. La-
bial palpus upcurved, broadly scaled; reaching a little
above vertex. Maxillary palpus squamous, appressed
to face. Forewing smooth except for a projecting tuft
of rough scales from base of costa in male; 11 veins; vein
2 from very close to lower outer angle of cell; 3 from the
angle, shortly separated from 4; 4 and 5 connate or very
closely approximate at bases; 6 from below upper angle
of cell, straight; 8 and 9 stalked; 10 from the cell, ap-
proximate to the stalk of 8-9 for some distance; male
without costal fold. Hind wing with vein 2 from lower
outer angle of cell; 3 and 5 shortly stalked; 7 and 8 ap-
proximate (or contiguous) for a short distance (less than
half their lengths) beyond cell; cell about one-third the
length of wing, shorter in male than female; on under
side of male wing, between cell and vein 1c, an elongate
pocket enclosing hair tuft and a mass of modified scales;
discocellular vein curved. Wighth abdominal segment
of male with a pair of ventrolateral hair tufts and ster-
nite produced as a sclerotized pocket (projecting into
abdomen).
Male genitalia with apical process of gnathos a slen-
der, elongate, flattened hook. Uncus rounded, hook-
like. Transtillas complete, thrust anteriorally from
costal margins of harpes. Harpe broad, apex broadly
and evenly rounded; clasper appressed to harpe; base of
sacculus produced and bearing a stout hair tuft. Anel-
lus a slightly curved, elongate plate. Aedeagus slender,
smooth, moderately long, sharply bent beyond one-
third from base; penis without cornuti or other appreci-
able armature. Vinculum long, narrow, strongly
arched dorsally, strongly sclerotized only on lateral and
terminal margins; constricted laterally and flaring to a
broad lateral margin.
Female genitalia with signum consisting of a de-
pressed, tear-shaped patch of short scobinations with a
sparser scattering of weaker scobinations in surrounding
area, (signa shown in frontal and lateral views in figs.
761 and 762); bursa otherwise simple; ductus bursae
shorter than bursa, simple; genital opening simple; duc-
tus seminalis from bursa near its junction with ductus
bursae.
The genus is very closely related to and apparently
derived from Drescoma in group I, agreeing with it on
most important genitalic characters, differing chiefly in
the absence of vein 4 of hind wing.
Contains but one tropical American species.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 2
544. Drescomopsis soraella (Druce)
Ficures 16, 284, 761, 762
Homoeosoma soraella Druce, Biologia Centrali-Americana, Lepi-
doptera Heterocera, vol. 2, p. 565, 1899.
Drescoma soraella (Druce) Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 328, 1914.
Drescoma drucella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 328,
1914 (new synonymy).
Drescomopsis subelisa Dyar, Ins. Inst. Menstr., vol. 7, p. 62, 1919
(new synonymy).
Forewing with costal area above middle of cell and
from just beyond base to subterminal line white; re-
mainder of wing a dull purplish fuscous; a rather broad
glossy dark brown oblique antemedial band dividing
the white area and extending half way across wing; ex-
treme base of wing on costal half similarly colored; a
narrowly lunulate brown patch on midcosta and a simi-
lar shade extending from apex transversely toward cell,
broken toward costa by faint indication of a subterminal
pale line; discal dots small but distinct; under magnifi-
cation the dark brown markings show an admixture of
reddish scales. Hind wings dark smoky fuscous, the
veins and terminal margin slightly darker. Alar ex-
panse, 14-16 mm.
Male genitalia with apical process of gnathos sharply
pointed; clasper a short, sharp spine; transtilla a
narrow, short, arched band with lateral ends produced,
pointed, and directed anteriorally. Female genitalia as
given for the genus. There is considerable individual
variation in the size of the bursa, but as this is an
expansible organ (when unsclerotized) its size and shape
mean little or nothing.
TypE LocauitiEs: Jalapa, México (soraelia, in BM);
Porto Bello, Panama (drucella, in USNM); Cayuga,
Guatemala (swbelisa, in USNM).
Foop piant: Unknown.
Distripution: M*f&xico: Cérdoba (May), Jalapa.
GuatemaLa: Cayuga (Apr., June). Costa Rica:
Juan Vifias (June, Nov.), Sitio. PanamA: Paraiso
(May), Porto Bello (Mar., Oct.), Rio Trinidad (Mar.).
Ecuapor: Zaruma. Braziu: Santa Catarina, Santa
Catarina Isl. (June).
Dyar evidently did not look very carefully at his
male type of drucella or the specimens from Costa Rica
he had as soraella or he never would have put them into
a genus with eight veins in hind wing, nor associated
with his males of drucella some small females of Dres-
coma cyrdipsa. The true females of his drucella he
later described as Drescomopsis subelisa.
Genera 153 and 154: Illatila and Lascelina
[Venational division A. Forewing with 11 veins; 4 and 5 stalked.
Hind wing with vein 2 close to or from lower outer angle of cell;
discocellular vein curved. Male genitalia with transtilla
absent; apical process of gnathos considerably reduced. Labial
palpus upturned.]
153. Genus Illatila Dyar
Illatila Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 334, 1914.
of genus: Jllatila gurbyris Dyar.)
(Type
Tongue well developed. Antenna pubescent, simple.
Labial palpus upturned, slender; reaching a little above
vertex; third segment acuminate, more than half as long
assecond. Maxillary palpus filiform, appressed to face.
Forewing smooth; 11 veins; vein 2 from before the lower
outer angle of cell, well separated from 3; 3 from the
angle; 4 and 5 short stalked, the stalk separate from 3
at base; 6 from below upper angle of cell, straight; 8 and
9 stalked; 10 from the cell, closely approximate to 8-9
for a short distance from cell; male without costal fold.
Hind wing with vein 2 from lower outer angle of cell;
3 and 5 stalked for less than half their lengths; 7 and 8
anastomosed beyond cell for a trifle less than half their
lengths; cell short, less than one-third the length of
wing; discocellular vein curved; on male the inner
margin thickened and folded with a fringe of hairs
enclosed within the fold. Eighth abdominal segment
of male with sternite produced as a sclerotized pocket,
projecting into abdomen.
Male genitalia with apical process of gnathos weak,
straight, very slender (needlelike). Uncus subtriangu-
late. Transtilla absent (elements not distinguishable).
Harpe rather narrow, apex evenly rounded; sacculus
produced at extremity into a short spine; base of
sacculus hairy, but not produced. Anellus a curved
plate. Adeagus short, moderately stout; cornuti pres-
ent, consisting of rather short stout spines in clusters.
Vinculum stout, triangulate; terminal margin rounded.
Female genitalia with strong signa, developed as a
small cluster of blunt, stout, thornlike spines; ductus
bursae sclerotized throughout, tubular, stout; genital
opening simple; ductus seminalis from bursa between
signa and ductus bursae.
This genus is quite distinct in genitalic characters
from anything else in group II. Its nearest affinities
seem to be with Difundella and related genera in group I.
It contains but one tropical American species.
545. Illatila gurbyris Dyar
Ficures 60, 561, 1054
Illatila gurbyris Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 334, 1914,
Dyar’s original description is accurate and adequate
except that the ground color is a rusty brown rather
than “brownish gray,’’ discocellular vein of forewing
outlined by blackish scaling with a broader pale outer
border (Dyar’s “discal spot”). Alar expanse, 12-13
mm.
Male genitalia with gnathos weakly attached to
tegumen at base of uncus, its lateral arms produced
anteriorly (in relation to head of insect) into widely
spaced ribbonlike bands. Apex or uncus_ broadly
rounded. Cornuti consisting of three spine clusters, two
of them of stout and one (apical) of slender spines.
Vinculum a trifle longer than broad (foreshortened in
fig. 561); tapering; terminal margin narrowly rounded.
Tegumen with lateral margin at base produced into a
rounded projection with serrate edge.
Female genitalia with bursa finely scobinate over
most of its inner surface; ductus bursae bulging and
264
heavily rugose near junction with bursa, nearly as long
as bursa.
TyPE LocaLity: Taboga Isl., Panamd (Feb.; type in
USNM).
Foop prant: Unknown.
Represented only by the original type series. Hasily
distinguished from anything else in the family by its
unique male and female genitalia.
154. Lascelina, new genus
Typr or GENus: Lascelina canens, new species.
Tongue well developed. Antenna pubescent; shaft
of male with sinus and strong scale tuft at base; of fe-
male simple. Labial palpus upturned, reaching vertex
in male, well above vertex in female; slender; rough
scaled; third segment slightly shorter than second,
bluntly pointed. Maxillary palpus minute, filiform.
Forewing smooth; 11 veins; vein 2 from before, but near,
lower outer angle of cell; 3 from the angle; 4—5 shortly
stalked, the stalk separate from 3 at base; 6 from below
upper angle of cell, straight; 8-9 stalked; 10 from the
cell, approximate to stalk of 8-9 for some distance from
cell; male without costal fold. Hind wing with vein 2
from very close to lower outer angle of cell; 3 and 5
stalked for half their lengths; 7 and 8 anastomosed for
half their lengths; cell slightly less than half the wing
length; discocellular vein curved. Highth abdominal
segment with sternite produced as a sclerotized pocket
projecting into abdomen.
Male genitalia with gnathos and its apical process
greatly reduced, latter fused with subanal plate. Uncus
triangulate. Transtilla absent (elements not distin-
guishable). Harpe broad, evenly curved; clasper pres-
ent as a short erect lobe; a pair of strong, detached hair
tufts adjacent to bases of sacculi. Anellus a triangu-
late, slightly curved plate. Aedeagus short, stout;
penis armed with minute spines and fine scobinations.
Vinculum long (longer than combined tegumen and
uncus) ; U-shaped; slightly tapering to rounded terminal
margin.
Female genitalia with bursa copulatrix elongate;
signum present, consisting of a single long stout curved
spine; ductus bursae very short, sclerotized near junc-
tion with bursa, the sclerotizations extending into the
neck of bursa; genital opening simple.
A distinct genus of uncertain affinities, easily distin-
guished by its male and female genitalia. Represented
only by its type species.
546. Lascelina canens, new species
Fieures 562, 1055
Forewing grayish white (the ground color that of
wood ashes); antemedial and subterminal lines faint,
indicated chiefly by the dark markings bordering them;
antemedial line bordered inwardly by a thin broken
blackish line and terminating at costa in a small reddish
brown patch; a similar, smaller, reddish brown patch on
midcosta; subterminal line bordered on both sides
UNITED STATES NATIONAL MUSEUM BULLETIN 207
towards costa by blackish fuscous; discal spots at end of
cell, small, distinct, blackish; along termen a confluent
row of blackish dots. Some reddish brown and blackish
scaling on the labial palpi and a strong admixture of
blackish scales in the antennal tuft of the male. Hind
wing smoky white, with a dark line along termen. Alar
expanse, 11-16 mm.
Male genitalia with uncus very narrowly rounded at
apex; costa of harpe broadly sclerotized; terminal mar-
gin oblique, straight; anellus with lateral margins
slightly produced; aedeagus bulged before apex. Fe-
male genitalia as given for the genus.
Typr Locatity: Brownsville, Tex. (type m USNM,
61380; paratypes in BM, Paris Mus., and Cornell.
Canadian National, and Janse Collections).
Foop piant: Maytenus phyllanthoides.
Described from male type and 24 male and 17 female
paratypes from the type locality (40 of these reared
Apr. 29, Aug. 4, and Dec. 7, 1938, and June 16, 1943, by
members of the Foreign Plant Quarantine Division of the
U. S. Department of Agriculture at Brownsville, Tex.,
from larvae feeding on the leaves of Maytenus) ;1maleand
41 female paratypes from San Benito, Tex. (Mar., Apr.,
May, Aug., Sept.); 1 female paratype from Redington,
Ariz., and 1 female paratype from Prescott, Ariz. (July).
In addition to the foregoing I have also seen a male from
Los Mochis, Sinaloa, México, intercepted in quarantine
at Nogales, Ariz. (Feb. 27, 1941).
Genus 155: Metephestia
[Venational division D. Forewing with 10 veins; 4 absent; 10
from the cell; hind wing with 3-5 approximate at base; discocel-
lular vein straight, vertical. Male genitalia with uncus bifur-
cate.]
155. Genus Metephestia Ragonot
Meiephestia Ragonot, Monograph, pt. 2, p. viii, 1901.—Hamp-
son, in Ragonot, Monograph, pt. 2, p. 87, 1901. (Type of
genus: Ephestia simplicula Zeller.)
Tongue well developed. Antenna very shortly pu-
bescent;shaft of male with a shallow sinus and small scale
tuft at base; shaft of female simple. Labial palpus up-
turned, slender, reaching to vertex; third segment acu-
minate, about half as long as second. Maxillary palpus
filiform. Forewing smooth; 10 veins; vein 2 from before
but near lower outer angle of cell; 3 from the angle; 4
absent (united with 5); 5 separated slightly from 3 at
base; 6 from below upper angle of cell, very slightly
curved (nearly straight); 8 and 9 stalked; 10 from the
cell, separate from the stalk of 8-9; male without costal
fold. Hind wing with vein 2 from well before lower
outer angle of cell; 3 and 5 from the angle, approximate
at base; 7 and 8 anastomosed for less than half their
lengths beyond cell; cell very short, about one-fifth the
length of wing; discocellular vein straight, vertical.
Abdomen of male with a strong pair of ventrolateral hair
tufts.
Male genitalia with apical projection of gnathos pro-
duced into a long, slender hook with notched apex.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Uncus bifurcate. Transtilla absent. Harpe simple
except for short, bluntly rounded clasper; slender; sac-
culus reduced; costa not appreciably sclerotized; apex
rounded. Anellus a narrow curved band with long
lateral lobes and long, slender, sclerotized lat-
eral attachments to tegumen. Aedeagus long, stout,
partially sclerotized; penis armed with a single, stout
cornutus. Vinculum elongate, narrowing abruptly be-
yond base.
Female genitalia without signum; bursa copulatrix
with some fine sclerotized wrinklings and a small lobe of
thickened membrane near ductus bursae; ductus bursae
much shorter than bursa, expanding abruptly to en-
larged genital opening; a pair of weakly sclerotized
plates on membrane behind genital opening; ductus and
genital opening otherwise simple; ductus seminalis from
bursa near thickened lobe.
A distinct genus of uncertain affinities, with male gen-
italia unlike anything else in group II. Contains but
one described species.
547. Metephestia simplicula (Zeller)
Fiaures 100, 563, 1056
Ephestia simplicula Zeller, Horae Soc. Ent. Rossicae, vol. 16, p.
246, 1881.
Metephestia simplicula (Zeller) Hampson, in Ragonot, Mono-
graph, pt. 2, p. 87, 1901.
Forewing unicolorous light gray brown to dark gray
(reared examples much darker than flown specimens);
transverse lines obsolete; in fresh specimens faint traces
of dark terminal and discal dots, otherwise unmarked.
Hind wing semihyaline white with a faint ocherous tint
towards apex and termen. Alar expanse, 11-16 mm.
Genitalia as given for the genus; apex of harpe nar-
rowly rounded; vinculum longitudinally grooved and
with notched apex.
Typr Locauity: Honda, Colombia (type in BM).
Foop puants: Indigofera tinctoria and I. verbasifo-
lium.
Distrisution: Unirep Sratss: Florida, Key West
(Mar., Apr.). Pusrro Rico: Catafio (Apr., July),
Coamo Springs (Apr.), Dovado (May), Rio Piedras
(Aug.), San German (Apr.); Puerto Real (Vieques Isl.,
Apr.). Cotomsra: Honda (Apr.). Also reported by
Hampson from St. Vincent Isl., British West Indies.
The above food-plant records are from a series of
moths reared by the Special Survey of the Division of
Foreign Plant Quarantine of the U. S. Bureau of Ento-
mology and Plant Quarantine from larvae webbing the
leaves of Indigofera. This is the first record we have of
the species from the United States or of its host plant.
Genera 156-159: Selga to Rioja
[Venational division A. Forewing with 11 veins; 4 and 5 stalked.
Hind wing with 2 from before lower outer angle of cell; 3 and 5
stalked or connate (Entmemacornis); discocellular vein curved.
Male genitalia with apical process of gnathos bifid; transtilla
complete (except in Selga). Labial palpi oblique (Selga) or
upturned.]
265
156. Selga, new genus
Typ or GENUS: Heterographis arizonella Hulst.
Tongue well developed. Antenna pubescent; shaft
of male simple. Labial palpus oblique, reaching height
of vertex; slender; third segment nearly as long as second,
apex bluntly pointed. Maxillary palpus minute, fili-
form. Forewing smooth; 11 veins; vein 2 from before
but near lower, outer angle of cell; 3 from the angle; 4
and 5 shortly stalked (for less than half their lengths),
stalk separated at base from 3; 6 from below upper angle
of cell, straight; 8 and 9 stalked; 10 from the cell, sepa-
rate and divergent from the stalk of 8-9; male with a
narrow costal fold, extending for over one-fourth of costa
from base. Hind wing with vein 2 from before lower
outer angle of cell; 3 and 5 from the angle, shortly
stalked (for less than half their lengths); 7 and 8 strongly
anastomosed for most of their lengths beyond cell; cell
one-half the Jength of wing; discocellular vein curved.
Kighth abdominal segment of male simple.
Male genitalia with apical process of gnathos U-
shaped (consisting of a pair of widely spaced, short,
blunt arms). Uncus stout, broadly rounded. Trans-
tilla incomplete. Harpe simple, apex rounded; costa
sclerotized throughout and projecting slightly at apex.
Anellus a broad, curved plate. Aedeagus short, stout;
penis with sclerotized wrinklings, otherwise unarmed.
Vinculum stout, rather short (broader than long);
terminal margin broadly rounded.
Female genitalia with bursa copulatrix elongate,
minutely scobinate; signum present, consisting of a
small coarsely and bluntly spined plate. Ductus bursae
unsclerotized, shorter than bursa; with a pair of narrow
elongate dorsal plates and minute scobinations at genital
opening, otherwise simple. Ductus seminalis from
bursa about half-way between signum and junction of
ductus bursae.
A distinct genus easily distinguished on male genitalia
from anything else in group IT with 11 veins in forewing.
Its male genitalia are strikingly similar to those of
Myelopsis coniella in group I to which it apparently is
closely related. It contains one North American
species.
548. Selga arizonella (Hulst), new combination
Ficurss 564, 1057
Heterographis arizonella Hulst, Journ. New York Ent. Soc., vol.
8, p. 222, 1900.—McDunnough, Check list, No. 6340, 1939.
Forewing white, finely and evenly peppered with
blackish scales giving the wing a pale ashy gray color;
antemedial line white, slightly notched in cell and more
deeply notched at lower fold, outwardly margined by a
blackish line and inwardly margined by a broader
blackish shade; subterminal line parallel with and
rather near to terminal margin, indented slightly at vein
6 and lower fold, white bordered inwardly and out-
wardly by blackish lines; blackish discal dots at end of
cell small, more or less confluent, in more strongly
marked specimens a fine blackish line extends from
them to middle of inner margin; a row of small blackish
266
dots along terminal margin. Hind wing glistening
white with some faint fuscous shading on the veins and
along terminal and costal margins. Alar expanse,
14-19 mm.
Genitalia as given for the genus; aedeagus sharply
constricted towards apex; apical margin of anellus
broadly notched; elements of transtilla long, slender.
Type LocaLity: Catalina Springs, Ariz. (type in
USNM).
Foop prant: Unknown.
Distrisution: Arizona, Catalina Springs (Apr.),
Redington, ‘‘Southern Arizona” (May), Tempe (June).
157. Genus Entmemacornis Dyar
Entmemacornis Dyar, Ins. Insc. Menstr., vol. 7, p. 57, 1919.
(Type of genus: Entmemacornis proselytes Dyar.)
Tongue well developed. Antenna pubescent; of
male (fig. 566f) with basal segment broadened and shaft
with notch at base; of female simple. Labial palpus
upturned, reaching to vertex, slender; third segment
nearly as long as second, acuminate. Maxillary palpus
minute, filiform. Forewing smooth; 11 veins; vein 2
from well before lower, outer angle of cell; 3 from the
angle; 4 and 5 shortly stalked (for less than half their
lengths), the stalk separated from 3 at base; 6 from
below upper angle of cell, slightly curved towards base;
8 and 9 long stalked; 10 from the cell but approximate
to the stalk of 8-9 for some distance; male with a
narrow costal fold. Hind wing with vein 2 from well
before lower outer angle of cell; 3 and 5 connate from
the angle; 7 and 8 anastomosed almost to costa (free
element of 8 very short); cell less than half the length
of the wing; discocellular vein curved. Eighth abdom-
inal segment of male with a pair of ventrolateral hair
tufts.
Male genitalia with apical process of gnathos bifid,
consisting of a pair of short blunt arms (similar to but
not so widely separated as those of Selga). Uncus
semispoon-shaped (of the Diatomocera type). Tran-
stilla complete but weakly sclerotized, a broad, finely
scobinate plate, weakly attached to harpes. Harpe
simple; terminal margin evenly rounded; costa sclero-
tized throughout but not produced; on outer surface
bordering lower margin a row of modified, flattened
setae. Anellus a curved plate with short lateral lobes.
Aedeagus stout, straight, moderately long; penis with a
few sclerotized wrinklings and several minute spines
(the latter distinguishable only under high magnifica-
tion), otherwise unarmed. Vinculum stout, elongate,
constricted towards angulate terminal margin.
Female genitalia with bursa copulatrix long, minutely
but sparsely scobinate, the scobinations extending into
ductus bursae; signa present, consisting of small
sclerotized disks; ductus bursae long, slender, un-
sclerotized; genital opening simple; ductus seminalis
from bursa near its anterior end.
On nearly all characters except hind-wing venation
this genus agrees with Diatomocera in group I. Were
it not for the fact that the four specimens of the type
UNITED STATES NATIONAL MUSEUM BULLETIN 207
series of Hntmemacornis proselytes are consistent in
venation, I should be inclined to discount the absence
of vein 4 in hind wing as an individual aberration and
unite the two genera; but apparently the same relation-
ship prevails here as that between Drescoma and Dresco-
mopsis. Entmemacornis is closely related to and
obviously derived from Diatomocera but is apparently
distinct.
549. Entmemacornis proselytes Dyar
Fieures 81, 566, 1058
Enimemacornis proselytes Dyar, Ins. Insc. Menstr., vol. 7, p. 58,
Thorax gray brown; tegulae tips black. Forewing
eray-brown dusted with dull white and reddish scales
(the latter only distinguishable under magnification) ;
antemedial line slanting, notched at lower fold, whitish
and concolorous with the general color of the basal area,
defined inwardly by a narrow, faint, dark border and
outwardly by a diffused dark shade; from base of wing
a median longitudinal black streak extending to but
not cutting the antemedial pale line; subterminal line
faint, parallel and rather close to termen, outcurved
between veins 6 and lower fold, formed by whitish
scaling on the veins and bordered by narrow inner and
outer dark lines more or less interrupted between the
veins; distal and terminal dots, small, faint, blackish;
between cell and subterminal line black streaks on veins
5 and 6; costal fold of male about one-third the length
of costa. Hind wing pale, glossy, brownish gray; the
veins and terminal margin darker. Alar expanse,
15-17 mm.
Male genitalia with terminal margin of uncus convex;
apex of harpe narrowly rounded; arms of bifid apical
process of gnathos slightly convergent toward their
apices; plate of anellus narrow. Female genitalia with
bursa bent sharply at middle, its terminal margin
straight.
TypELocatiry: Cayuga, Guatemala (typein USNM).
Foop piant: Unknown.
Known only from the type series, two males and two
females from the type locality.
550. Entmemacornis pulla, new species
FicurE 567
Forewing blackish gray; antemedial line slanting,
notched at lower fold, dull white with a very faint,
narrow, blackish outer border; subterminal line some-
what irregular, slanting sharply inwardly from costa to
vein 8, then outwardly convex to lower fold, thence
straight to inner margin, dull white with some blackish
streaks on the veins bordering it on both sides; discal
and terminal dots small, faint, blackish; basal area of
wing before antemedial line and between cell and inner
margin dull reddish ocherous; a dusting of similar color
along the fold beyond antemedial line; from base of
wing a median longitudinal black streak extends to
antemedial line; bordering termen a narrow faint dust-
ing of whitish scales; costal fold of male less than one-
fourth the length of costa. Hind wing very pale
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
brownish gray; veins and terminal margin but little
darker. Alar expanse, 19.5 mm.
Male genitalia with terminal margin of uncus slightly
concave; apex of harpe rather broadly rounded; arms
of bifid apical process of gnathos divergent toward their
apices; plate of anellus broad.
Type Locatiry: Santa Catarina, Brazil (type in
USN, 61381).
Foop prant: Unknown.
Described from unique male type collected by Fritz
Hoffmann Sept. 27, 1934.
The species is easily separated from proselytes by its
darker (blackish gray) ground color, more strongly con-
trasted transverse pale lines, shorter costal fold, and dif-
ferent genitalia. The tegulae are black tipped like those
of proselytes, but their blackness is less contrasted
against the dull reddish ocherous of the remainder of the
thorax.
158. Genus Cayennia Hampson
Cayennia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 62,
1930. (Type of genus: Cayennia rufitinctalis Hampson.)
Tongue well developed. Antenna pubescent (cilia-
tions less than the width of shaft). Labial palpus up-
turned, reaching to vertex, slender, third segment nearly
as long as second. Maxillary palpus minute, filiform.
Forewing smooth; 11 veins; vein 2 from well before lower
outer angle of cell; 3 from the angle; 4-5 stalked for
nearly half their lengths, approximate at base to 3; 6
from below upper angle of cell, very slightly curved; 8
and 9 long stalked; 10 from the stalk of 8-9; 11 from
well out on cell, running close to stalk of 8-9 for a short
distance; male without costal fold. Hind wing with
vein 2 from before lower outer angle of cell; 3 and 5 from
the angle, stalked (for approximately one-half) ; 7 and 8
anastomosed for most of their lengths beyond cell (free
element of 8 a very short spur); cell nearly one-half the
length of the wing; discocellular vein curved. Eighth
abdominal segment of male with a pair of ventrolateral
hair tufts.
Male genitalic characters same as those for Entmema-
cornis except penis with only sclerotized wrinklings.
A distinct genus close to Hntmemacornis and agreeing
with it in all male genitalic characters, differing only in
having simple antennae and different venation (vein 10
of forewing stalked with 8-9, and 3-5 of hind wing
stalked) and in lacking costal fold on male forewing.
551. Cayennia rufitinctalis Hampson
Ficurss 72, 568
Cayennia rufitinctalis Hampson, Ann. Mag. Nat. Hist., ser. 10,
vol. 5, p. 62, 1930.
Forewing pale ashy gray peppered with fuscous, the
basal half of inner area and tornal half of terminal area
tinged with rufous; antemedial line near middle of wing,
narrow, slanting outwardly from costa to middle of cell
thence inwardly to lower fold and thence outwardly to
inner margin, whitish with blackish lines before and
beyond it on the veins; subterminal line inwardly
oblique and slightly concave from apex to inner margin
267
before tornus, whitish, margined inwardly by short
blackish dashes on the veins; discal dots small, blackish,
below them on an inwardly slanting line blackish dots
on veins 3 and 2b; a row of small blackish dots along
termen. Hind wing semihyaline white with veins,
terminal and costal margins shaded with pale brown.
Alar expanse, 17.5 mm.
Male genitalia with apical portion of uncus narrow
(considerably reduced as compared with that of
Entmemacornis proselytes); transtilla triangulate; vin-
culum tapering, but slightly constricted before narrow,
angulate terminal margin.
Typr LocaLity: Cayenne, French Guiana (type in
BM).
Foop piant: Unknown.
The female is unknown. Hampson mentions only
the male type in his description; but in the U. S.
National Museum there is a male “‘cotype’”’ so labeled
by Hampson from the type locality. The species is
easily identified by its genitalia and the medial position
of the antemedial line of forewing.
159. Rioja, new genus
TYPE OF GENUS: Rioja nexa, new species.
Tongue well developed. Antenna pubescent. Labial
palpus upturned, reaching vertex, slender; third seg-
ment almost as long as second. Maxillary palpus
minute, filiform. Forewing smooth, 11 veins; veins 2
and 3 connate from lower outer angle of cell; 4 and 5
short stalked (for decidedly less than half their lengths),
approximate at base to 3; 6 from below upper angle of
cell, straight; 8 and 9 long stalked; 10 from cell, ap-
proximate to stalk of 8-9 at base; male without costal
fold. Hind wing with 2 from before lower outer angle
of cell; 3 and 5 stalked for less than half their lengths;
7 and 8 anastomosed beyond cell for half their lengths;
cell a trifle less than half as long as wing; discocellular
vein curved. Eighth abdominal segment of male with-
out hair tufts; sternite developed as a triangulate,
sclerotized pocket.
Male genitalia with apical process of gnathos bifid.
Uncus semispoon-shaped, constricted before apical end.
Transtilla a complete arched bridge. Harpe simple;
cucullus broad and with terminal margin evenly
rounded; costa sclerotized but not produced. Anellus
a narrow curved band with long lateral arms partially
encircling the aedeagus. Aedeagus stout, moderately
long; penis minutely scobinate, otherwise simple.
Vinculum stout, longer than broad, slightly tapering.
This genus forms a link between the two preceding
genera and Moerbes Dyar. I should hesitate to describe
it upon a single male if it were not obvious that the
specimen is not a freak, and if it did not have characters
that prevented its inclusion in Entmemacornis, Cayen-
nia, or Moerbes.
Its uncus and gnathos are of the Entmemacornis
type, its eighth-segment pocket similar to that of
Moerbes; but its venation is unique. The connate con-
268 UNITED STATES NATIONAL MUSEUM BULLETIN 207
dition of veins 2 and 3 (from the angle of the cell) at
once distinguishes it from all near relations.
552. Rioja nexa, new species
Figures 70, 569
Forewing blackish with basal area to antemedial area
dusted with whitish scaling and contrastingly paler than
remainder of wing; also some whitish dusting forming a
pale transverse shade across wing at end of cell; ante-
medial line outwardly slanting from costa to inner mar-
gin with a slight notch at lower fold, white, outwardly
bordered by a narrow black line; subterminal line faint,
irregularly dentate, pale (whitish gray), bordered in-
wardly by a broad blackish suffusion; discal dots con-
fluent, forming a blackish line along discocellular vein;
a row of faint blackish dots along termen. Hind wing
hyaline white with a faint fuscous shade on costa and
on termen towards apex. Alar expanse, 17 mm.
Male genitalia with arms of bifid apical process of
gnathos divergent toward their pointed apices; aedeagus
bulging and slightly bent before its broad, truncate
apex.
Type LOCALITY: La Rioja, Argentina (type in USNM,
61382).
Foop piant: Unknown
Described from a single male collected by Schaus.
The specimen is undated.
Genera 160-162: Moerbes to Edulica
[Venational division A. Forewing with 11 veins; 4 and 5 stalked.
Hind wing with 2 from before lower outer angle of cell; 3 and
5 connate or stalked (Edulica); discocellular vein curved. Male
genitalia with apical process of gnathos a stout hook with forked
apex; transtilla complete or, if incomplete (Moerbes), elements well
developed and their apices approximate. Labial palpi oblique or
upturned (Edulica).]
160. Genus Moerbes Dyar
Moerbes Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 337, 1914. (Type
of genus: Zophodia dryopella Schaus.)
Tongue well developed. Antenna shortly ciliate on
male (cilia as long as width of shaft). Labial palpus
oblique, broadly scaled, reaching to level of vertex;
third segment short (one-fourth the length of second),
acuminate. Maxillary palpus moderately large, some-
what rough scaled. Forewing smooth; 11 veins; vein 2
from well before lower, outer angle of cell; 3 from the
angle; 4 and 5 stalked (for a trifle less than half their
lengths), the stalk shortly separated from 3 at base;
6 from below upper angle of cell, slightly curved; 8 and
9 stalked for about two-thirds their lengths; 10 stalked
with 8-9; male with a short, narrow costal fold. Hind
wing with vein 2 from well before lower, outer angle of
cell; 3 and 5 from the angle, connate; 7 and 8 anasto-
mosed for most of their lengths (free element of 8 very
short) ; cell approximately one-third the length of wing;
discocellular vein curved. Eighth abdominal segment
of male without hair tufts, but with sternite developed
as a shallow, triangulate, sclerotized pocket.
Male genitalia with apical process of gnathos devel-
oped as an elongate, stout, flattened hook with forked
apex. Uncus subtriangulate with broad terminal mar-
gin. ‘Transtilla incomplete, its elements elongate-angu-
late and well developed. Harpe simple with terminal
margin evenly rounded; costa sclerotized for four-fifths
of its length, but not produced. Anellus a narrow,
curved band with minute lateral lobes. Aedeagus mod-
erately stout, nearly straight; penis simple. Vinculum
stout, longer than broad, slightly tapering to blunt ter-
minal margin.
A distinct genus showing strong affinities in general
habitus (wing markings and color) and male genitalic
structure to Pseudodiwona of group I. However, the
venational difference (the absence or presence of vein 4)
is consistent in both genera and the association of
Moerbes with the genus following (Moodnopsis) seems a
natural and proper one. It contains three tropical
American species represented in the National Museum
by eight males. I have seen no females.
553. Moerbes dryopella (Schaus)
Ficures 82, 570
Zophodia dryopella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11,
p. 249, 1913.
Moerbes dryopella (Schaus) Dyar (part), Proc. U. S. Nat. Mus.,
vol. 47, p. 337, 1914; Ins. Insc. Menstr., vol. 7, p. 54, 1919.
Forewing white with a broad, dull, ocherous brown
shade along inner margin, and a sparse peppering of
black scales in the white areas; antemedial line rather
broad, defined chiefly by thin inner and outer bordering
black lines, and conspicuous as a white spot on inner
margin, outwardly slanting from costa, inwardly angled
at lower part of cell and thence incurved to inner mar-
gin; subterminal line faint, defined by two black dashes
before and beyond it on costa and thin, broken, blackish
bordering lines, inwardly angled at vein 6, thence paral-
lel to termen; costa from base to antemedial line black,
and a fine black transverse subbasal line; lower discal
spot at end of cell present, black; below it a similar black
spot or dash merging into the brown shade bordering
inner margin; discal spot at upper outer angle of cell
obsolete; terminal blackish dots faint. Hind wing very
pale smoky fuscous, veins and terminal margin darker.
Alar expanse, 23 mm.
Male genitalia with apex of apical projection of
gnathos considerably broadened and rather deeply bi-
furcate. Apices of transtilla elements well separated.
Aedeagus smooth, stout. Terminal margin of vinculum
angulate.
TYPE LOCALITY: Juan Vinas, Costa Rica (Apr.; type
in USNM).
Foop piant: Unknown.
Represented only by the unique male type. The
specimens from Panamé referred by Dyar to Schaus’
species (Proc. U.S. Nat. Mus., vol. 47, p. 337, 1914) are
congeneric but not conspecific. I am describing them
in this paper as Moerbes emendata.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 269
554. Moerbes alveolella (Ragonot), new combination
Fiaure 571
Zophodia alveolella Ragonot, Nouv. Gen., p. 31, 1888; Mono-
graph, pt. 2, p. 25, 1901.
The type in Paris is a female according to Ragonot.
I have not seen it; but have before me two males from
Santa Catarina, Brazil (Sept. 2, and Oct. 26, 1934, col-
lected by Fritz Hoffman), which are a perfect match for
Ragonot’s excellent figure (Monograph, pl. 25, fig. 7).
They are identical in color and maculation with the type
of dryopella but markedly different in structure: 8 and
9 of forewing longer stalked (over two-thirds); apical
process of gnathos longer, its apex narrow and shallowly
bifurcate; apices of transtilla elements touching (but
not fused) and forming a high arch over aedeagus;
aedeagus more slender; terminal margin of vinculum
bluntly rounded (almost straight). Alar expanse, 21-23
mm.
TyprE Loca.ity: Rio de Janeiro, Brazil (type in Paris
Mus.).
Foop puant: Unknown.
555. Moerbes emendata, new species
Figure 572
Moerbes dryopella Dyar (not Schaus), Proc. U. S. Nat. Mus., vol.
47, p. 337, 1914.
Similar in markings and color to the two preceding
species except that the ground color of forewing is a
more soiled white (less contrasted, except for the white
spot on inner margin formed by the base of the ante-
medial line), and both upper and lower discal spots are
present and distinct. The male genitalia are also
markedly different: Uncus appreciably narrower than
that of alveolella or dryopella; aedeagus with a thornlike
projection from undersurface before apex; vinculum
sharply tapering from middle to angulate terminal
margin. Alar expanse, 16-20 mm.
Tyrer Locatity: Cabima, Panamé (type in USNM,
61383).
Foop piant: Unknown.
Described from male type from the type locality
(May); one male paratype from La Chorrera, Panamé;
one male paratype from Porto Bello, Panama (Mar.);
one male from Lino, Panamé (from the Janse Collec-
tion); one male from 6 miles up the Maroni River,
French Guiana (Schaus, collector), and one male without
locality label, collected by Schaus and presumably also
from French Guiana. The Panama specimens, except
for the one from Lino, were collected by Busck and are
those referred by Dyar to dryopella Schaus. ‘The Porto
Bello specimen is abnormal in that veins 8 and 9 are
united on both forewings; but this is a freak. Other-
wise the specimen is normal and on the other specimens
the venation is normal. The species can be distin-
guished at once by the thornlike projection from its
aedeagus.
161. Genus Moodnopsis Dyar
Moodnopsis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 408, 1914.
(Type of genus: Moodnopsis decipiens Dyar.)
Campyloplesis Dyar, Ins. Insc. Menstr., vol. 7, p. 61, 1919. (Type
of genus: Campyloplesis inveterella Dyar. New synonymy.)
Tongue well developed. Antenna of male shortly
ciliate (cilia no longer than width of shaft); of female
pubescent. Labial palpus of male obliquely ascending,
cylindrical, reaching nearly to vertex, third segment less
than half as long as second; of female porrect, laterally
somewhat flattened, second segment oblique, long, ex-
tending to the level of vertex, third segment deflected
forward, about half the length of second. Maxillary
palpus filiform. Forewing smooth; 11 veins; vein 2
from well before lower outer angle of cell; 3 from the
angle; 4 and 5 stalked, the stalk approximate at base
(and in some specimens for a short distance from base)
with 3; 6 from below upper angle, slightly curved to-
wards base; 8 and 9 long stalked; 10 from the cell, more
or less approximate (rarely connate) to the stalk of 8-9
at base; male with a long costal fold (extending for half
or more than half the length of costa). Hind wing with
vein 2 from well before lower, outer angle of cell; 3 and 5
connate from the angle; 7 and 8 anastomosed beyond
cell almost to apex (completely anastomosed in a couple
of males); cell about one-third the length of the wing;
discocellular vein curved. Eighth abdominal segment
of male with a short pair of ventrolateral hair tufts and
sternite developed as a triangulate sclerotized pocket.
Male genitalia with apical process of gnathos de-
veloped as a stout, flattened hook with forked apex.
Transtilla complete, the apices of its elements enlarged
and fused into a spreading, bulbous knob. Harpe with
terminal margin evenly rounded; costa strongly sclero-
tized for four-fifths of its length, but not produced; on
outer surface, bordering inner margin, a row of modified,
flattened setae. Anellus a narrow, curved band with
short lateral lobes. Aedeagus smooth, straight or
slightly curved, tapering slightly from base to apex.
Vinculum stout, longer than broad, tapering or abruptly
constricted towards angulate terminal margin.
Female genitalia with bursa copulatrix finely scobi-
nate; signum present as an elongate narrow projecting
plate with serrate edge; ductus bursae shorter than
bursa, simple; genital opening simple; ductus seminalis
from bursa towards (but not near) its junction with
ductus bursae.
Dyar described Moodnopsis from two large dark fe-
male specimens and Campyloplesis from two small pale
males. On the evidence of these alone the generic sep-
aration would seem valid enough; but the evidence of
associated males and females of species other than the
type of genus shows no consistent character for such
separation. The palpal differences are purely sexual
and the trifling venational differences are either individ-
ual or, at most, specific in character. Moodnopsis is a
distinct genus easily identified by its male genitalia, and
is somewhat more closely related to Moerbes than to
270
Moodna, from which Dyar distinguished it. His diag-
nosis of Campyloplesis is in error in two important par-
ticulars: The male labial palpi are not ‘“‘porrect” but
obliquely ascenting, almost upcurved; veins 3, 4, and 5
of forewing are not “stalked,” 3 being only approximate
to the stalk of 4-5 at base and for a very short distance
beyond.
556. Moodnopsis decipiens Dyar
Fieures 83, 1062
Moodnopsis decipiens Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 408,
1914.
The two females from which the species was described
(and the only specimens available) are rubbed so that
the forewing pattern cannot be clearly distinguished.
The ground color is a brownish gray (rather dark) with
a faint dusting of whitish scales along costa and (under
magnification) a scattered peppering of reddish scales
over most of the wing; antemedial line indicated only by
a rather broad dark outer border, this broken; the sub-
terminal pale line faintly indicated, defined chiefly by
dark streaks on the veins before and beyond it, the inner
streaks rather long; discal dots faint but distinguish-
able, blackish; dots along terminal margin very faint
and more or less confluent. Hind wing pale fuscous;
the veins and terminal margin darker. Alar expanse,
27-28 mm.
Female genitalia with signum large, the serrations
along projecting edge bluntly and rather broadly
rounded.
TYPE LocaLity: Orizaba, México (type in USNM).
Foop erant: Unknown.
In unrubbed specimens the dark streaking along the
veins would be more emphasized. Such lining of the
veins is characteristic of all the species in the genus; but
decipiens stands out because of its greater size and
darker ground color.
557. Moodnopsis perangusta (Dyar), new combination
Ficure 573
Euzophera perangusta Dyar, Ins. Insc. Menstr., vol. 7, p. 57, 1919.
Ground color of forewing as in decipiens but trans-
verse lines more distinct; the antemedial line deeply
notched at lower fold; the subterminal irregularly den-
tate and parallel with termen; dark borders of the trans-
verse pale lines narrower and dark lining of the veins
much less pronounced than in decipiens; blackish discal
spots confluent; veins 4 and 5 longer stalked (for more
than half their lengths); 8 and 9 longer stalked and 10
closely approximate to the stalk of 8-9 for some distance
beyond cell; costal fold extending to well beyond middle
of costa. Hind wings translucent white, gray toward
apex and along upper half of terminal margin. Alar
expanse, 25 mm.
Male genitalia with bifurcation of apical process of
gnathos deeper, aedeagus smaller and less tapering,
vinculum broader at, and less constricted before, ter-
minal margins than those of other species in the genus.
The genitalic differences among the several species are
UNITED STATES NATIONAL MUSEUM BULLETIN 207
better illustrated than described and can be readily
distinguished in the drawings.
TypE LocaLity: Montserrat, Trinidad, British West
Indies (type in USNM).
Foop prant: Unknown.
Known only from the male type. It is possibly the
male of decipiens. The size and coloration of forewing
would suggest this. The white hind wings and vena-
tion, however, would seem to indicate a distinct species,
though the differences noted may be only sexual. The
length of the stalking of veins 4 and 5 of forewing is
certainly unreliable, varying in individuals of species
where we have more than two specimens.
558. Moodnopsis inornatella (Ragonot), new combination
Ficures 574, 1061
Zophodia inornatella Ragonot, Nouv. Gen., p. 31, 1888; Mono-
graph, pt. 2, p. 25, 1901.
The type of this species is a worn female from Costa
Rica without abdomen. In his original description
Ragonot gives the type locality as Brazil; but according
to Clarke this could easily have been a misreading of a
small pin label with ‘‘Druce” inscribed on it. The
female in the British Museum is labeled ‘ty. original’
in Ragonot’s handwriting and bears the further infor-
mation, ‘Irazu, 6-7,000 ft., H. Rogers.” This must be
the type, for Dr. Burgogne states that it is not in the
Ragonot Collection at the Paris Museum.
A photograph of the type agrees well with a series of
specimens in the National Museum (one male and three
females) from Juan Vifias, Costa Rica.
The general color of the forewing is paler than that
of either decipiens or perangusta; but the markings are
similar, the dark lining of the veins, if anything, more
distinct. Worn specimens show no trace of an ante-
medial line, but in one of the females before me (in
better condition than the others) it is faintly outlined.
The male costal fold is shorter than in perangusta,
extending only to middle of costa. Hind wing of
female very pale fuscous; of male soiled white; veins
and edge of terminal margin darker. Alar expanse,
20-28 mm.
The distinguishing characters of the male genitalia
are the shape of terminal projection of gnathos, the
shape of apical lobe of transtilla, and the constricted
vinculum. The female genitalia are similar to those of
decipiens except that the signum is smaller and has
weaker, less rounded serrations along its projecting edge,
trifling differences of rather doubtful value.
TypE Locauity: Irazt, Costa Rica (type in BM).
Foop piant: Unknown.
In addition to the Costa Rican specimens I have
before me a2 male from Santa Catarina, Brazil (July 22,
1935, Fritz Hoffmann, collector) which I take to be
anornatella or a variety of it. The fore and hind wings
are considerably paler (more whitish) and the ante-
medial and subterminal lines more clearly indicated.
It is the same size as the male from Juan Vinas (20
mm.), has a costal fold of the same length, and agrees
with it in every genitalic detail except that the forking
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 271
of the apical process of gnathos is a trifle shallower. It
certainly is not conspecific with the males of the follow-
ing species from the same locality and collection.
Upon superficial examination vein 3 of forewing seems
to be connate with the stalk of 4-5 in inornatella and in
most specimens of the other species, and is so described
by Ragonot; but if the wings are partially denuded on
underside at outer end of cell, more or less of a sepa-
ration shows between 8 and 4—5 at their bases.
559. Moodnopsis parallela, new species
Fiaure 575
Forewing similar in color and markings to that of the
females of inornatella; costal fold very long, extending
for at least two-thirds the length of costa. Male geni-
talia with apical process of gnathos longer than that of
other species of Moodnopsis except inveterella and its
apical bifurcation U-shaped; aedeagus very stout,
straight and evenly tapering; apical lobe of transtilla
considerably enlarged; vinculum constricted before
terminal margin, similar to that of inornatella. Alar
expanse, 21-23 mm.
Type Locauity: Santa Catarina, Brazil (type in
USNM, 61384).
Foop prant: Unknown.
Described from male type and four male paratypes
from the type locality (Oct. 2, 1934, July 7, 8, 12, 1935,
Fritz Hoffmann, collector). In addition to these I have
before me a somewhat larger male (26 mm.) with identi-
cal genitalia from Tinguri, Carabaya, Peri (Schaus,
collector)
The female is unknown.
560. Moodnopsis inveterella (Dyar), new combination
Ficures 86, 576
Campyloplesis inveterella Dyar, Ins. Insc. Menstr., vol. 7, p. 61,
Similar in color and maculation to inornatella Ragonot
but much smaller and with quite different genitalia.
The venation of the type is figured (fig. 86) to show
extent of variation in venation within the genus between
it and the large female of decipiens (fig. 83). The com-
plete fusion of veins 7 and 8 of hind wing in the type of
inveterella is apparently an abnormality, a short spur of
8 showing in one of the hind wings of the paratype.
The costal fold extends for half the length of costa.
Alar expanse, 14.5 mm.
The distinguishing characters of the genitalia are:
The stout, flat-faced apical process of gnathos (fig. 576b)
with shallow V-shaped bifurcation at its apex; the nar-
row, looped knob at apex of transtilla (fig. 576c); and the
very abruptly tapered vinculum with its narrow termi-
nal margin. The aedeagus is moderately stout and
tapering.
TypE tLocauity: Cayuga, Guatemala (type in
USNM).
Foop prant: Unknown.
Known only from the male type (June) and a male
paratype (Apr.) from the type locality.
561. Moodnopsis portoricensis, new species
Fiaures 577, 1060
In size, color, and maculation similar to inveterella but
with differently shaped uncus, apical process of gnathos,
and transtilla. Alar expanse, 14.5-16 mm.
The female genitalia are distinguished only by a very
narrow, short signum, with sharply serrate edge and set
far towards the anterior end of the bursa. The position
of the signum is a character of very doubtful value.
Type tocauity: Lares, Puerto Rico (type in Cornell
Univ.; paratypes in USNM, 61385).
Foop prant: Unknown.
Described from male type and one male and one
female paratype from the type locality, collected by
Francesco Sein, Jr., July, 1931.
162. Genus Edulica Ragonot
Edulica Ragonot, Monograph, pt. 2, p. ix, 1901.—Hampson, in
Ragonot, Monograph, pt. 2, p. 122, 1901. (Type of genus:
Euzophera compedella Zeller.)
Tongue well developed. Antenna pubescent; basal
joint enlarged on male. Labial palpus upturned; third
segment approximately half as long as second, dorsally
flattened. Maxillary palpus squamous, appressed to
face. Forewing smooth; 11 veins; vein 2 from near
lower outer angle of cell; 4 and 5 shortly stalked, from
the angle; 3 connate or very shortly stalked with 4-5;
6 from upper angle of cell, curved; 8 and 9 stalked; 10
from the stalk of 8-9; male without costal fold. Hind
wing with vein 2 from very close to lower outer angle of
cell; 3 and 5 stalked for at least half their lengths; 7 and
8 anastomosed beyond cell for approximately half their
lengths; cell one-third the length of wing; discocellular
vein curved. EKighth abdominal segment of male with
sternite developed as a shallow, triangulate, sclerotized
pocket, otherwise simple.
Male genitalia with apical process of gnathos devel-
oped as an elongate, stout, flattened hook, with very
slightly notched apex. Uncus stout with broadly
rounded terminal margin. Transtilla complete, devel-
oped as a strongly sclerotized arch with broad, flaring
apical crossband. Harpe with a strong hair tuft from
base of sacculus; constricted between sacculus and
cucullus; terminal margin rounded; costa sclerotized for
two-thirds its length, not produced; clasper short, ap-
pressed. Anellus a slightly curved, narrow band with
well developed lateral lobes. Aedeagus short, stout;
penis armed with many strongly sclerotized folds and
a cluster of moderately stout, straight, elongate spines
(cornuti) about one-fourth as long as aedeagus. Vincu-
lum stout, as broad as long, triangulate; terminal end
pointed.
Female genitalia with bursa copulatrix sclerotized
towards junction with ductus bursae and with several
sclerotized folds extending well into the ductus, finely
scobinate over most of inner surface; signum present as
a small, cupped, scobinate plate; ductus bursae much
shorter than bursa; genital opening simple.
The genus is apparently close to but distinct from
272 UNITED STATES NATIONAL MUSEUM BULLETIN 2:07
Euzophera, easily distinguished from that genus by the
close association of veins 3, 4 and 5 of forewing, and the
short cell of hind wing. Contains one tropical Ameri-
can species. Hampson includes a species from Mada-
gascar but this is probably improperly placed.
562. Edulica compedella (Zeller)
Fieures 58, 585, 1071
Euzophera compedella Zeller, Horae Soc. Ent. Rossicae, vol. 16,
p. 224, 1881.
Edulica compedella (Zeller) Ragonot, Monograph, pt. 2, p. ix,
1901.—Hampson, zn Ragonot, Monograph, pt. 2, p. 122,
1901.
Ground color of forewing varying from somber brown
to gray-brown; costal half of wing dusted with ashy
grayish white; the discal spots blackish, confluent; some
scattered blackish dots on several of the veins; ante-
medial line not defined; the subterminal one weakly so
by its dark inner and outer bordering lines, irregularly
dentate; terminal dots distinct, blackish. Hind wing
of male whitish with dark brown shading on veins and
along termen; of female smoky fuscous, darkening
towards apex and termen. Alar expanse, 20-25 mm.
Genitalia as given for the genus.
TyPE Locatity: Honda, Colombia (type in BM).
Koop Pprant: Unknown.
Distrisution: PanamA: Porto Bello (Apr., May).
Cotompia: Honda. French Guiana: St. Jean
Maroni. Braziu: Amazonas, Ponte Nova (Rio
Xingu).
Genera 163 and 164: Huzophera and Exuperius
[Venational division A. Forewing with 11 veins; 4 and 5 stalked.
Hind wing with vein 2 from well before lower outer angle of cell;
3 and 5 connate; cell long; discocellular vein curved. Male geni-
talia with apical process of gnathos a stout, elongate hook with
pointed apex; transtilla complete. Labial palpi upturned]
163. Genus Euzophera Zeller
Stenoptycha Heinemann (not Zeller), Schmetterlinge Deutsch-
lands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 190, 1866.
Melia Heinemann, Schmetierlinge Deutschlands und der
Schweiz, Abt. 2, vol. 1, pt. 2, p. 209, 1866 (mew name, pre-
occupied, proposed for Stenoptycha Heinemann).
Euzophera Zeller, Trans. Ent. Soc. London, ser. 3, vol. 5, p. 456,
1867 (mew name for Stenoptycha Heinemann); Stettiner
Ent. Zeit., vol. 28, p. 377, 1867.—Hulst, Phycitidae of
N. Amer., p. 174, 1880 (cites type as Myelozs cinerosella
Zeller).—Hampson, Moths, vol. 4, p. 72, 1896, in Blanford,
Fauna of British India (cites type as Ephestia biviella
Zeller); 7n Ragonot, Monograph, pt. 2, p. 36, 1901.—
Ragonot, Monograph, pt. 2, p. vii, 1901 (cites type as
Phycis pinguis Haworth).—Forbes, Cornell Mem. 68, p.
630, 1923.—Bisset, zn Pierce and Metcalf, Genitalia of the
British Pyrales, p. 59, 1938 (notes fixation of type by
Hulst).—Janse, Journ. Ent. Soc. South Africa, vol. 8, p. 31,
1945. (Type of genus: Myelois cinerosella Zeller; figs. 77,
578, 1063.)
Tongue well developed. Antenna simple, shaft
pubescent (ciliations very short). Labial palpus up-
turned, reaching to or almost to vertex; second segment
rough scaled; third segment about half as long as
second, acuminate. Maxillary palpus subsquamous
(somewhat rough scaled), appressed to face. Forewing
smooth; 11 veins; vem 2 from well before lower, outer
angle of cell; 3 from the angle; 4 and 5 stalked (for less
than half the lengths) ; 6 from below upper angle of cell,
straight; 8 and 9 long stalked; 10 normally from the
cell, separated from or approximate to (rarely connate
with or from the stalk of) 8-9; male without costal fold.
Hind wing with vein 2 from well before lower outer
angle of cell; 3 and 5 from the angle, connate, rarely (in
individual specimens) very shortly stalked; 7 and 8
strongly anastomosed beyond cell (for more than half
their lengths); cell long, slightly more than half the
length of wing; discocellular vein curved. Wighth
abdominal segment of male simple except in cinerosella
which has a pair of short ventrolateral hair tufts.
Male genitalia with apical process of gnathos devel-
oped as an elongate, stout, simple hook with pointed
apex. Uncus stout, broadly triangulate. Transtilla
complete, developed as a strongly sclerotized arch with
prominent, caudally projecting, lateral lobes. Harpe of
nearly equal width throughout; terminal margin
rounded and with a short rounded projection from apex
of costa; costa otherwise, for most of its length, strongly
sclerotized; clasper not developed. Anellus a slightly
curved U-shaped shield, the lateral lobes well developed
and strongly sclerotized in the American species.
Aedeagus moderately long, stout; penis armed with
numerous, short, sharp spines. Vinculum stout, as long
as (or very little longer than) broad; terminal margin
rounded.
Female genitalia with numerous sclerotized or scob-
inate folds in bursa copulatrix adjacent to or at junction
with ductus bursae (except im the European type,
cinerosella), finely scobinate over most of inner surface;
signum present as a small, cupped, scobinate plate;
ductus bursae with genital opening simple; ductus
seminalis from bursa near signum (in American species)
or from terminal end of bursa (in cinerosella). Dorsal
sclerotized area of eighth segment collar a narrow band
with central tonguelike projection.
Euzophera, like Zophodia, has served as the receptacle
for a heterogeneous lot of species, most of which must
be referred to various other genera. As here defined
the genus contains only three described American
species that can be referred to it with any certainty.
These form a compact group with constant genitalic
characters and similar forewing pattern: antemedian
line far out on wing, subterminal line well back from the
terminal margin, the space between the lines corre-
spondingly reduced, and a white spot on discocellular
vein. On venation, palpi, and male genitalia they
agree with the European type of genus, cinerosella, but
differ from it in wing pattern and two details of struc-
ture, cinerosella having hair tufts on the eighth abdom-
inal segment of the male, and the ductus seminalis from
the anterior end of the female bursa (figs. 578b, 1063).
Eventually it may be possible to give our American
species a separate generic designation; but upon present
knowledge they must be retained in Huzophera.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Venation in Huzophera is subject to considerable
individual variation especially as regards the relation
of vein 10 of forewing to 8-9. In our American species
and the European cinerosella (fig. 77) vein 10 is from the
cell and normally distinctly separated from the stalk of
8-9, though in individual specimens of any American
species it may be either connate with or approximate,
for some distance, to the stalk. In the European
pinguis Haworth, it may be from the middle of the
stalk, short stalked, or connate with it, or (rarely) from
the cell and distinctly separate.
563. Euzophera semifuneralis (Walker)
Ficures 579, 1064
Nephopteryx semifuneralis Walker, List, pt. 27, p. 57, 1863.
Euzophera aglacella Ragonot, N. Amer. Phycitidae, p. 14, 1887;
Monograph, pt. 2, p. 65, 1901.—Hulst, Phycitidae of N.
Amer., p. 177, 1890.—Essig, Insects of western North Amer-
ica, p. 710, 1929.—McDunnough, Check list, No. 6316, 1939.
Stenoptycha pallulella Hulst, Ent. Amer., vol. 3, p. 137, 1887.
Euzophera semifuneralis (Walker) Ragonot, Ent. Amer., vol. 5, p.
116, 1889.—Forbes, 8. A., Psyche, vol. 5, p. 295, 1890.—
Hulst, Phycitidae of N. Amer., p. 175, 1890.—Hampson, in
Ragonot, Monograph, pt. 2, p. 66, 1901.—Blakeslee, U. S.
Dep. Agr. Bull. 261, 13 pp., 1915.—Forbes, W. T. M.,
Cornel Mem. 68, p. 631, 1923.—HEssig, Insects of western
North America, p. 710, 1929.—Keifer, Monthly Bull.
Dep. Agr. California, vol. 20, p. 618, 1931.—McDun-
nough, Check list, No. 6317, 1939.
Forewing with basal and terminal areas dark to pale
reddish brown more or less dusted with white along
costa and (in very pale specimens) in apical-terminal
area; area between the transverse lines densely dusted
with blackish scales; antemedial line more or less ver-
tical to lower margin of cell, inwardly angled at lower
fold, white bordered outwardly by a black line; sub-
terminal line somewhat irregular, usually out-angled
evenly at middle, white, bordered inwardly by a black
line; discal black dots more or less obscured in the black
dusting of median area, usually a white mark on dis-
cocellular vein; a line of black dots along terminal
margin, confluent in some specimens, distinctly sepa-
rated in others. Hind wing white to smoky fuscous,
more or less darkened along terminal margin, at apex,
and along some of the veins. Alar expanse, 16.5-28
mm. (Hampson gives extreme expanse as 34 mm.).
Male genitalia with lateral, projecting lobes of trans-
tilla and lateral lobes of anellus more slender than those
of other American species; cucullus of harpe narrower
and more elongate; vinculum somewhat longer than
broad; uncus evenly tapering. Female genitalia with
bursa rather small and wrinkled over most of its surface.
Type tocauitiss: “North America” (semifuneralis,
in BM); Sonora, México (aglaeella, in Paris Mus.);
Colorado (pallulella, in AMNH, ex Rutgers).
Foop piants: Various deciduous trees (apple, apri-
cot, pear, peach, plum, persimmon, cherry, mulberry,
walnut, pecan, olive, linden, poplar, Liquidambar,
Ginkgo, mountain ash). Larva a bark borer. Also
stems of cotton and corn.
Distrisution: Unirep States: New Hampshire,
Hampton; Connecticut, East River (July, Sept.); New
273
York (no exact locality); New Jersey, Hackensack
(Nov.), Montclair (May, Aug.), Morristown (June);
Maryland, Plummers Isl. (May); District of Columbia,
Washington (May, Sept.); Virginia, Cape Henry (July),
Vienna (May); North Carolina, Southern Pines (Mar.,
Apr.), Tryon (May); South Carolina, Anderson (Oct.);
Georgia, Fort Valley; Michigan, Monroe; Indiana, Bed-
ford (Apr.); Illinois, Decatur (Apr.); Kansas, Lawrence
(May), Onaga; Missouri, St. Louis (June); Mississippi,
Jackson (Feb.), Tishomingo (Nov.); Texas, Blanco
County, Brownsville, Brownwood (Oct.), Justine (June),
Kerrville (Apr.), Paris, Plano (Sept.), San Benito (July),
San Diego (May), Shovel Mountain, Snyder (Mar.,
Apr.), Victoria (Apr.), Zavalla County (Apr.); New
Mexico, Mesilla, Roswell (Apr.); Arizona, Baboquivari
Mts. (Apr., May), Chiricahua Mts., Garces, Huachuca
Mts., Palmerlee, Scotsdale (May), Yavapai County;
Colorado, Denver (June); Utah, Bellevue (May); Cali-
fornia, Corningi (Jan.), Inyo County (June, July),
Loma Linda (Sept.), Piru (Ventura County, July), Pla-
cerville (Jan., May), Putah Canyon (Yolo County,
Nov.). Canapa: British Columbia, Alberni (July),
Duncans (Vancouver Isl., Aug.). Mzs&xico: Sonora.
The species is variable in color but is easily distin-
guished by the reddish basal and terminal areas of fore-
wing and the narrowed, black-dusted area between the
transverse lines. The name aglaeella represents a color
form, with paler red-brown areas and more strongly
contrasted, blackish median dustings and markings.
It is commoner in the Western and Southwestern States
and México than elsewhere but cannot be maintained
asarace;forit also occurs in the Kast and there are inter-
grades between it and the typical dark form, common to
the Eastern and Central States, but occurring through-
out the range of the insect. The species is of some im-
portance as an orchard insect, the larvae often domg
considerable damage as a bark borer in plum, olive, and
walnut trees. It is known in economic literature as
the ‘‘American plum borer.”
564. Euzophera ostricolorella Hulst
Figures 580, 1065
Euzophera ostricolorella Hulst, Phycitidae of N. Amer. p. 175,
1890.—Hampson, in Ragonot, Monograph, pt. 2, p. 67,
1901.—Heinrich, Proc. U. 8. Nat. Mus., vol. 57, p. 87, 1920
(life history, larva and pupa).—Forbes, Cornell Mem. 68,
p. 631, 1923.—McDunnough, Check list, No. 6319, 1939.
Forewing purplish brown, dark grayish fuscous along
costa and with some grayish dusting in apical area;
transverse lines whitish, not darkly bordered; anteme-
dial line vertical to lower margin of cell, deeply angled
below, at fold; subterminal line sinuate; a whitish line
along discocellular vein at end of cell; along termen a
row of obscure, blackish dots; in some specimens a suf-
fusion of dark red brown in basal area and above inner
margins beyond base, especially on the folds. Hind
wing rather pale smoky fuscous with a fine dark line
along terminal margin. Alar expanse, 29-40 mm.
Male genitalia with lateral, projecting lobes of trans-
tilla broader than those of semifuneralis; anellus very
274
broad, its lateral lobes wide and widely spaced, their
apices narrowly rounded; harpe with cucullus broaden-
ing toward apex; vinculum about as broad as long;
uncus abruptly narrowed beyond base, thence tapering.
Female genitalia with bursa large, densely spined and
wrinkled towards junction with ductus bursae; ductus
bursae very short, deeply wrinkled.
Type Locatity: New York (in AMNH, ex Rutgers).
Foop puant: Liriodendron tulipifera.
Distrisution: New York, Long Island; District of
Columbia, Washington (June); Maryland, Oxon Hill,
Plummers Isl. (Oct.); Virginia, Leesburg; South Caro-
lina, Greenville (June), Hendersonville (June), Paris
Mountain (June), Rockhill (June), Yorkville (June);
Georgia, Atlanta (Oct.).
Probably distributed throughout the range of its host.
This distinct species apparently has only one host, the
‘tulip tree.’”’? The larva bores in the bark on the damp
side near the base of the tree.
565. Euzophera nigricantella Ragonot
Ficures 581, 1066
Euzophera nigricantella Ragonot, N. Amer. Phycitidae. p. 14,
1887.—Hulst, Phycitidae of N. Amer., p. 177, 1890.—Hamp-
son, in Ragonot, Monograph, pt. 2, p. 63, 1901.—McDun-
nough, Check list, No. 6315, 1939.
Euzophera griselda Dyar, Proc. U. 8. Nat. Mus. vol. 44, p. 322,
1913 (new synonymy).
Forewing gray evenly dusted with blackish; trans-
verse pale lines paler, but obscure, shaped as in the pre-
ceding species; antemedial line with an obscure narrow
blackish outer border; subterminal with faint dark bor-
ders on inner and outer sides; a whitish spot on disco-
cellular vein; terminal dots, faint, blackish. Hind wings
white with some dark shading on the veins and along
terminal margin. Alar expanse, 21-27 mm.
Male genitalia with lateral projecting lobes of trans-
tilla largest of any known American Huzophera; anellus
longer than broad, the lateral lobes untapering, nar-
rowly spaced, and with outer margins at apices inwardly
hooked; harpe with cucullus stubby, its lower margin
slightly concave; vinculum as broad as long; uncus as in
ostricolorella. Female genitalia with a wrinkled sclero-
tized collar about ductus bursae at its junction with
bursa copulatrix.
Type tocauities: Arizona (nigricantella; in Paris
Mus.); Tehuacén, México (griselda, in USNM).
Foop piant: Unknown.
Distrisution: Untrep Staves: Arizona, Baboqui-
vari Mts. (Apr., June, July, Aug., Sept., Oct.), Reding-
ton, Tempe (Mar.), Yavapai County; New Mexico, Al-
buquerque (July) ; Texas, San Benito (May). M*xico:
Sonora; Tehuacén (May, Sept.), San Jose de Guaymas
(Apr.).
Dyar’s griselda was described from females which
match specimens of nigricantella from Arizona in every
detail of maculation, color, and genitalia. At the time
he described griselda Dyar had not recognized Ragonot’s
species.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
164. Exuperius, new genus
TYPE OF GENUS: Hruperius negator, new species.
Tongue well developed. Antenna simple, pubescent.
Labial palpus upturned, barely reaching to vertex; first
segment broadly scaled; third segment nearly as long as
second, acuminate. Mazxillary palpus minute, filiform.
Forewing smooth; 11 veins; vein 2 from before lower
outer angle of cell; 3 from the angle; 4 and 5 stalked (for
less than half their lengths), separate from 3 at base: 6
slightly bent, from below, but near upper angle of cell;
8 and 9 rather long stalked (for two-thirds their lengths) ;
10 from the cell, approximate to the stalk of 8-9 for
some distance; male without costal fold. Hind wing
with vein 2 from well before lower outer angle of cell;
3 and 5 connate, from the angle; 7 and 8 anastomosed
beyond cell for most of their lengths (free segment of 8
short); cell long, slightly over half the length of wing;
discocellular vein curved. Highth abdominal segment
of male simple.
Male genitalia with apical process of gnathos de-
veloped as an elongate, stout, simple hook with pointed
apex. Uncus narrow, constricted towards base, shaped
like a narrow, bluntly pointed arrowhead. ‘Transtilla
complete, but median portion a narrow, rather weakly
sclerotized band. Harpe with broadened, evenly
rounded cucullus; spoon-shaped; costa broadly sclero-
tized but not produced; clasper present, short, knobbed
at apex; from near base of cucullus a strong, long, semi-
detached hair tuft. Anellus a broadly U-shaped nar-
row band with long, flattened, lateral projections.
Aedeagus long, stout; penis armed with a dense mass of
minute granulations. Vinculum considerably shorter
than broad, terminal margin broadly and evenly
rounded.
Female unknown.
This genus is close to Huzophera and agrees with it in
characters of venation, labial palpi, and gnathos; but its
peculiar genitalia (aside from the apical projection of
enathos) differ in every detail from those of typical
Euzophera and suggest a separate generic designation.
They resemble those of no other American group that
I know. The partial forking of vein 1b of forewing at
base (shown in fig. 76) is present on both forewings but
may be an individual abnormality. It is most unusual
for a phycetine.
566. Exuperius negator, new species
Ficures 76, 584
Forewing brown, dusted with grayish white over basal
area and for a short distance beyond antemedial line
near costa and inner margin; some blackish dusting in
cell beyond its middle and a black spot on costa on each
side of the subterminal line; antemedial line indicated
by a blackish band slanting slightly inward from slightly
beyond basal third of costa to inner margin, straight;
subterminal line somewhat wavy, obscure except on
costal half where it is distinctly white; a small but con-
spicuous white spot above the stalk of veins 4—5 at end
of cell; terminal dots blackish, more or less confluent.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 275
Hind wing hyaline white shaded with blackish fuscous,
broadly along costa and narrowly along terminal margin;
veins faintly outlined by dark shading. Alar expanse,
22 mm.
Male genitalia as given for the genus; aedeagus
broadened at base, sharply constricted just beyond.
Typr Locauity: La Chorerra, Patamayo District,
Pert (type in Cornell Univ.).
Foop piant: Unknown.
Described from unique male type collected on the
Cornell University Expedition (Apr. 1920) under lot
No. 607.
The white discal spot on forewing and the genitalia
should easily identify the species. The former is also
common to our American species of Huzophera but in
that genus the transverse lines are much closer together
than in negator.
Genus 165: Eulogia
[Venational division A. Forewing with 11 veins; 4-5 approx-
imate, connate or very shortly stalked. Hind wing with 3 and 5
connate; cell long; discocellular vein curved. Male genitalia
with apical process of gnathos a short, flat, furcate plate; trans-
tilla incomplete. Labial palpus upturned.]
165. Eulogia, new genus
Typz or Gunus: Ephestia ochrifrontella Zeller.
Characters of Huzophera except: Labial palpus with
third segment as long or nearly as long as second. Fore-
wing of male with a narrow, elongate costal fold; veins
4 and 5 approximate, connate or very shortly stalked.
Male genitalia with apical process of gnathos a short,
flat plate, furcate at apex; transtilla incomplete; aede-
agus slender; penis armed with a single curved cornutus.
Female genitalia with bursa copulatrix smooth except
for a very few scobinations surrounding signum; signum
a cluster of small, short, thornlike spines (not ona plate) ;
ductus bursae strongly sclerotized at and just before
genital opening, the sclerotized part of ductus tubular.
On venation, wing maculation, and palpal characters
the type species (ochrifrontella) could go well enough in
Euzophera, where it has been placed; but the genitalia
rule it out. All the really congeneric species of Europe
and North America placed in Huzophera have similarly
constructed, complete transtillae, the same type of
hooked projection from gnathos, similar multiple cor-
nuti on the penis, and similar wide-mouthed, unsclero-
tized genital openings in the female. On all of these
characters ochrifrontella is ruled out.
567. Eulogia ochrifrontella (Zeller), new combination
Ficures 78, 79, 583, 1069
Ephestia ochrifrontella Zeller, Verh. zool.-bot. Ges. Wien, vol. 25,
p. 337, 1876.
Euzophera ferruginella Ragonot, N. Amer. Phycitidae, p. 14,
Euzophera ochrifrontella (Zeller) Hulst, Phycitidae of N. Amer.,
p. 177, 1890.—Hampson, in Ragonot, Monograph, pt. 2,
p- 67, 1901.—Ely, Proc. Ent. Soc. Washington, vol. 12, p.
203, 1910.—Forbes, Cornell Mem. 68, p. 631, 1920.—
McDunnough, Check list, No. 6318, 1939.
Forewing copper colored, with the area between the
transverse lines heavily dusted with black; antemedial
line far out (slightly beyond middle) on wing, pale
ocherous, angulate; subterminal line, parallel with
termen, slightly indented at vein 6 and lower fold; an
obscure pale ocherous line along discocellular vein; in
some specimens the blackish dusting extends beyond
the transverse lines over most of the basal and terminal
areas of the wing, but this is not a normal condition.
Hind wing pale smoky fuscous; veins and terminal
margin slightly darker. Alar expanse, 11-15 mm.
Genitalia as given for the genus. In the female the
ductus bursae has a slightly sclerotized band near its
junction with bursa.
Typr tocauities: Texas (ochrifrontella, in MCZ);
North Carolina (ferruginella, in Paris Mus.).
Foop puants: Pecan, oak, apple.
Distrisution: Unitrep Statrs: Massachusetts, Co-
hasset (July), Framingham (Sept., Oct.), Martha’s
Vineyard (July, Aug.); Connecticut, East River (July,
Aug., Sept., Oct.), Stamford (July); New Jersey, Eliza-
beth (Aug.); Pennsylvania, Adams County (June),
Arendtsville (Aug.), Hazleton (July), New Brighton
(June, July); Maryland, Plummers Isl. (May, June,
July); North Carolina; Florida, Orlando (Mar.); Texas;
Illinois, Decatur (May, June); Kansas, Onaga. Can-
ADA: Manitoba, Rounthwaite (July), Winnipeg; British
Columbia, Duncans (Vancouver Isl., June), Kaslo
‘@uly).
The foregoing food-plant records are from specimens
in the National Museum. Nothing, as far as I know,
has been published on the life history.
The venation is variable in different specimens and
sometimes on opposing wings of the same insect. On
forewing vein 10 may be separate from, approximate to,
or connate with the stalk of 8 and 9; 4 and 5 may be
approximate or connate or, sometimes, very shortly
stalked. On the hind wing the anastomisis of 7 and 8
varies, but is normally short. There is no appreciable
individual variation in genitalia.
Genus 166: Prosoeuzophera
[Venational division D. Forewing with 10 veins; 4 absent; 3 and
5 connate. Hind wing with veins 3 and 5 connate or shortly
stalked. Male genitalia with apical process of gnathos a stout,
elongate hook with pointed apex; transtilla complete. Labial
palpi upturned.]
166. Prosoeuzophera, new genus
Type or Genus: Euzophera impletella Zeller.
Characters of Huzophera except: Forewing with 10
veins; vein 4 absent; 3 and 5 connate from lower outer
angle of cell.
A direct derivative of Huzophera but with advanced
venation, replacing Luzophera in tropical America.
276
568. Prosoeuzophera impletella (Zeller), new combination
Figures 107, 582, 1067
Euzophera impletella Zeller, Horae Soc. Ent. Rossicae, Vol. 16,
p. 234, 1881.
In color and maculation like small, pale examples of
Euzophera semifuneralis with which it was synonymized
by Hampson (Ragonot, Monograph, pt. 2, p. 66,
1901). He either overlooked or ignored the distinct
venational difference between impletella and the North
American species of Huzophera. The genitalia of
impletella (o' and ) exhibit differences of only a
specific character from the Huzophera, quite distinct
specifically from semifuneralis, as our figures show; but
the absence of vein 4 and the connate condition of
veins 3 and 5 of forewing are constant. Alar expanse,
15-16.5 mm.
TyrE Locality: Honda, Colombia (Gn BM).
Foop piant: Unknown.
Distrisution: Cotompra: Honda, Maraquita. Ja-
marca (2, in USNM). Purrto Rico: Villa Margarita
(Catano, Apr.)
Genera 167-169: Farnobia to Micromescinia
[Venational division B. Forewing with 10 or 11 veins; 2 and 3
stalked or united. Cell open or incomplete in fore or hind wings.
Male genitalia with transtilla complete.]
167. Farnobia, new genus
TyrE oF Genus: Huzophera quadripuncta Zeller.
Tongue well developed. Antenna of male with basal
segment narrowly elongate, bearing a short spur from
inner side near apex; shaft deeply curved toward base,
pubescent; of female simple. Labial palpus upturned,
scarcely reaching vertex; third segment shorter than
second. Mazxillary palpus minute, rather broadly and
flatly scaled, appressed to face. Forewing smooth; 10
veins; veins 2 and 3 united; 4 and 5 short stalked; 6
curved and connate with the stalk of 8-9-10; 8 and 9
stalked; 10 from the stalk of 8-9; cell open (discocellular
vein absent); in male costa enlarged, lobed and sinuate,
and outer margin concave between veins 7 and 5; an
elongate discal fovea in male between veins 6 and the
stalk of 3-5. Sternite of eighth abdominal segment of
male developed as a broadly triangulate sclerotized
pocket.
Male genitalia with apical process of gnathos a short,
stout, blunt hook. Uncustriangulate. Transtilla com-
plete, a heavy arched bridge with enlarged lateral lobes.
Harpe with incurvation between sacculus and cucullus;
cucullus narrow and narrowly rounded at apex; costa
sclerotized for most of its length, but not produced.
Anellus a narrow curved band with broad, curved,
bandlike lateral projections (clasping the sides of
aedeagus). Aedeagus stout, somewhat bent; penis
armed with numerous sclerotized folds and two or three
stout, very short, thornlike spine clusters. Vinculum
UNITED STATES NATIONAL MUSEUM BULLETIN 207
stout, as broad as long, bluntly triangulate; terminal
margin reinforced and broadly rounded.
Female genitalia with bursa copulatrix large and
elongate, finely scobinate over surface of anterior half,
thickened and partially sclerotized at junction with
ductus bursae; signum present as a sclerotized cup
evenly covered with short, stout, blunt spines; ductus
bursae very short, a mere neck between genital opening
and bursa; genital opening with a curled-over, strongly
sclerotized and centrally emarginate lower margin;
ductus seminalis from a lobe of bursa near its junction
with ductus bursae.
In genitalia this genus resembles the Hemiptiolocera—
Crocidomera complex of group I with which it is appar-
ently related. It is even more nearly related to the
genus following (Gennadius) which also has the cell of
forewing open and (except for specific differences) sim-
ilar male genitalia. However, Farnobia is so different
in venational characters that it need not be confused
with anything else in the Phycitidae. The forewing
venation might be interpreted differently from what
T indicate (i. e., vein 4 rather than 3 absent and 3 and
5 short stalked); but in view of the outward position
of 2 and the close association of Farnobia with
Gennadius (which has 2 and 3 distinctly stalked) the
correct interpretation appears to be 2 and 3 united.
569. Farnobia quadripuncta (Zeller), new combination
FicuREs 98, 99, 586, 1070
Euzophera quadripuncta Zeller, Horae Soc. Ent. Rossicae, vol. 16,
p. 226.
Dannemora quadripuncta (Zeller) Hampson, in Ragonot, Mono-
graph, pt. 2, p. 209, 1901.
Forewing brown with a purplish tint; the costal area
dusted with ocherous gray, this pale shading extending
into the cell at middle; antemedial line obscure, blackish,
dentate and outwardly oblique from costa to inner mar-
gin; subterminal line better defined, blackish, sharply
dentate at the veins; three rather conspicuous blackish
dots forming an angle near extremity of cell and, in some
specimens, traces of a fourth dot on costa marking be-
ginning of antemedial line. Hind wings whitish, semi-
hyaline, with a narrow dark line along termen and dark
shading on some of the veins. On the underside of the
male hind wing the costal area is filled with coarse
ocherous scaling terminating in a blackish fuscous patch
at the outer angle of thelobe. Alar expanse, 21-24 mm.
Genitalia as given for the genus.
TyprE Locatity: Honda, Colombia (type in B. M).
Foop prant: Unknown.
Distrizution: Costa Rica: Esperanza (May).
Pawnam&: La Chorrera (May), Cabima (May). France
Guiana: St. Laurent du Maroni. Cotompra: Honda.
Also recorded by Hampson from Brazil.
When Hampson placed quadripuncta in Dannemora
he had only females of Zeller’s species and no authentic
examples of the type of Hulst’s genus. The two species
have very little in common, structurally or otherwise.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 277
168. Gennadius, new genus
Type or Genus: Gennadius junctor, new species.
Tongue well developed. Antenna of male with basal
segment narrowly elongate, bearing a short spur from
middle of inner side; shaft with a short spur from basal
joint, deeply curved for several segments beyond, pu-
bescent. Labial palpus oblique, slender, dorsoventrally
flattened, reaching above vertex; third segment about
as long as second. Maxillary palpus moderately sized,
flatly scaled and appressed to face. Forewing with a
subbasal clump of raised scales (possibly a male or spe-
cific character) ; 11 veins; veins 2 and 3 stalked; 4 and 5
stalked (for about half their lengths); 6 curved, and
connate with the stalk of 8-9-10; 8 and 9 stalked; 10
from the stalk of 8-9; cell open (discocellular vein ab-
sent); male without costal fold. Hind wing of male
with vein 2 from very close to angle of cell; 3 and 5
stalked; 7 and 8 united; in male costa triangularly pro-
duced and outer margin concave between veins 7 and 5;
cell closed, very short (less than one-fourth the length
of wing in male); discocellular vein curved. Sternite of
eighth abdominal segment of male developed as a
triangulate, sclerotized pocket.
Male genitalia as in Farnobia except: Apical hook of
gnathos slightly furcate at apex; lateral lobes of trans-
tilla bluntly pointed rather than rounded; lateral arms
of anellus shorter, narrower and tapering; harpe not
incurvate between succulus and cucullus. Most of
these differences are probably specific rather than
generic in character.
Female unknown.
The genus is very close to Farnobia as indicated by
the antennal and genitalic structures and the open cell
of forewing, but is easily separated by the stalking rather
than fusion of veins 2 and 3 of forewing and by the
closed cell and the absence of vein 8 in hind wing.
570. Gennadius junctor, new species
Figures 97, 587
Forewing brown with a purplish tint; the costal area
dusted with ocherous gray; some blackish streaking on
upper vein of cell and along median fold and a few
blackish spots indicating the broken margins of the
transverse lines; a conspicuous white patch along inner
margin between antemedial and subterminal lines and
extending from inner margin to cell; transverse lines
obscure. Hind wing very pale brown; terminal margin
and veins little if any darker; in the male rather coarsely
scaled over most of undersurface and with a border of
coarse, thick, ocherous scaling along costa. Alar
expanse, 22.5 mm.
Male genitalia as given for the genus; figure 587b
shows the penis extruded to display the cornuti.
Typr Locauity: St. Jean Maroni, French Guiana
(type in USNM, 61886).
Foop piant: Unknown.
Described from male type and one male paratype
from the type locality (Schaus, collector, no date given).
The species is easily identified by its structural
characters and the elongate white patch on inner
margin of forewing.
169. Genus Micromescinia Dyar
Micromescinia Dyar, Proc. U. 8S. Nat. Mus., vol. 47, p. 347, 1914.
(Type of genus: Micromescinia pygmaea Dyar.)
Tongue well developed. Antenna simple. Labial
palpus oblique, reaching to or a trifle above vertex; slen-
der; third segment nearly as long as second, acuminate.
Maxillary palpus minute, filiform. Forewing smooth;
10 veins; veins 2 and 3 stalked; 4 and 5 long stalked; 6
from below upper angle of cell, very slightly curved; 8
and 9 united (9 absent); male with a short costal fold
enclosing a hair tuft. Hind wing with 2 from before
(but near) lower outer angle of cell; 3 and 5 stalked,
from the angle; 7 and 8 completely fused beyond cell;
cell partially open (only a rudiment of discocellular
vein), long, a trifle more than half as long as wing.
Eighth abdominal segment of male simple.
Male genitalia with apical process of gnathos devel-
oped as a stout, curved, bluntly pointed hook. Uncus
stout, hoodlike, scarcely tapering and with broadly
rounded terminalmargin. Transtilla complete, a bridge
with central furcate projection, not appreciably arched.
Harpe with costa produced and angulate at middle.
Anellus a U-shaped plate with rather short lateral arms.
Aedeagus rather slender, simple, slightly curved; penis
armed with a thin elongate sclerotized band. Vincu-
lum moderately stout, about as broad as long, triangu-
late.
Female genitalia with bursa copulatrix weakly and
sparsely scobinate; signa present as a cluster of (3 or 4)
small, weakly sclerotized discs. Ductus bursae sclero-
tized, granulate and flattened for two-thirds of its length
from genital opening and with a produced, angulate,
sclerotized shield behind genital opening. Ductus sem-
inalis from bursa near signa (near anterior end of bursa).
A distinct genus showing affinities, especially in fe-
male genitalia, to Hphestiodes and several of the follow-
ing genera. Contains one tropical American species.
571. Micromescinia pygmaea Dyar
Ficures 96, 595, 1083
Micromescinia pygmaea Dyar, Proc. U. 8S. Nat. Mus., vol. 47,
p. 347, 1914.
Forewing pale ocherous with a narrow purplish red
shade along costa and a similarly colored broad suffu-
sion along termen. Hind wing pale smoky fuscous, paler
towards base. Alar expanse, 9-10 mm.
Genitalia as given for the genus; cucullus of harpe
angulate, apex bluntly pointed.
Type tocauity: Porto Bello, Panamé (type in
USNM).
Foop piant: Unknown.
Distripution: PanamA: Porto Bello (Mar.), Taber-
nilla, Taboga Isl. (Feb.).
Known only from the original type series (two males
and one female) in the National Collection.
278 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Genera 170-177: Ephestiodes to Moodnella
[Venational division C. Forewing with 10 veins; 8 and 9 united
(except in aberrant individuals); male with costal fold. Labial
palpi oblique. Male abdomen with compound dorsal tufts on
eighth segment or simple. Transtilla complete or incomplete;
if complete, an angulate bridge; if incomplete, its elements well
developed. Ductus bursae sclerotized for at least part of its
length from genital opening and with projecting sclerotized shield
behind genital opening.]
170. Genus Ephestiodes Ragonot
Ephestiodes Ragonot, N. Amer. Phycitidae, p. 16, 1887; Mono-
graph, pt. 2, pp. xiii, 264, 1901.—Hulst, Phycitidae of N.
Amer., p. 194, 1890.—Forbes, Cornell Mem. 68, p. 635, 1923.
(Type of genus: Ephestiodes gilvescentella Ragonot.)
Tongue well developed. Antenna simple, pubescent.
Labial palpus oblique, reaching to vertex, somewhat
rough scaled; segment 3 about aslong as 2. Mazxillary
palpus filiform. Forewing smooth; 10 veins; vein 2
from very near to lower outer angle of cell; 3 from the
angle; 4 and 5 stalked, the stalk approximate to or con-
nate with 3 at base; 6 from below upper angle of cell,
straight or very slightly curved; 8 and 9 united (9 ab-
sent); 10 from the cell, separated from 8; male with a
short costal fold. Hind wing with vein 2 from before
(but near) lower outer angle of cell; 3 and 5 long stalked;
7 and 8 anastomosed almost to costa (8 a mere vestige) ;
cell about half the length of wing; discocellular vein
curved. Highth abdominal segment of male with com-
pound dorsal scale tufts (except in noniella).
Male genitalia with apical projection of gnathos
forked, U- or V-shaped. Uncus broad and with apical
margin broadly rounded. ‘Transtilla incomplete (ex-
cept in stictella and noniella) but with the elements
strongly developed, their apices approximate and broad-
ly flared. Harpe normally (except in Jucidibasella,
stictella, and noniella) with a transverse sclerotized ridge
across base of cucullus; costa strongly sclerotized but
not produced. Anellus a narrow, semicircular band
with flanged base. Aedeagus straight, moderately long;
penis armed (except in noniella) with a weak, thin, flat
bladelike cornutus or a pair of similar cornuti. Vincu-
lum stout, triangulate.
Female genitalia with signum developed as a short
comb of from two to six blunt, short, thornlike teeth, or
as a flat plate with a row of such teeth along one edge
(plorella); bursa copulatrix otherwise simple except for
a few weak scobinations surrounding signum or at
junction of ductus bursae. Ductus bursae longer than
bursa (except in stictella), sclerotized, granulate and
flattened for at least half (normally for most) of its
length from genital opening, with a produced, strongly
sclerotized shield behind genital opening and usually
with a sclerotized band or shield below the opening;
ductus seminalis from bursa close to signum.
The North American species of the genus form a com-
pact group agreeing on all structural characters. In
tropical America, however, there is considerable varia-
tion from type and two of the species (stictella and
noniella) are distinctly aberrant on male characters.
Eventually it may be possible to give them separate
generic designations, but at present there are no charac-
ters to be found (apart from male genitalia) to permit
such separation. The species divide into three groups
as follows:
Transtilla incomplete, but strongly developed, with apices
flared and approximate. Harpe normally with a trans-
verse sclerotized ridge at base of cucullus. Eighth ab-
dominal segment of male with tufts. (Comprising all
the typical E’phestiodes.)
Transtilla complete, with greatly produced, asymmetrical,
caudal projections. Harpe with apex of sacculus pro-
duced into a free arm. Highth abdominal segment of
male with tufts. (Tropical only.)
Transtilla complete, a narrow band with central loop.
Harpe simple. Eighth abdominal segment of male
simple. (Tropical only.)
While adult specimens seem to be abundant and are
often taken in numbers, little if anything is known of
life histories within the genus and none of the species
has, so far, proven to be of any economic importance.
Most of the species are similar in color and maculation,
but each shows some distinguishing difference in male
genitalia. These are not easy to describe, but can be
seen readily enough in the drawings. The female geni-
talia of the North American species are remarkably
uniform in general structure, exhibiting more variation
within than between species. In all of them the dorsal
sclerotized area of the eighth-segment collar is reduced
to a narrow band, the sclerotized shield behind genital
opening has a central angulate projection from its pos-
terior margin, and the sclerotized band below the open-
ing is narrow.
Genus Ephestiodes, Species 572-580: E. gilves-
centella to E. plorella
[Transtilla incomplete but strongly developed, with apices flared
and approximate. Harpe normally with a transverse sclerotized
ridge at base of cucullus. Eighth abdominal segment of male
with tufts.]
572. Ephestiodes gilvescentella Ragonot
Figures 108, 589, 1072, 1073
Ephestiodes gilvescentella Ragonot, N. Amer. Phycitidae, p. 16,
1887; (Ragonot and Hampson), Monograph, pt. 2, p. 264,
1901.—Hulst, Phycitidae of N. Amer., p. 194, 1890.—
McDunnough, Check list, No. 6379, 1939.
Ephestiodes nigrella Hulst, Journ. New York Ent. Soc., vol. 8,
p. 224, 1800.—Donohoe and Barnes, Journ. Econ. Ent.,
vol. 27, p. 1071, 1984.—McDunnough, Check list, No. 6382,
1939. (New synonymy.)
Forewing pale gray, the median area (between the
transverse lines) dark gray; antemedial line outwardly
angled at middle and with a very slight notch at lower
fold, white with a narrow blackish outer border; sub-
terminal line fainter, slightly sinuate and parallel with
termen, whitish with a narrow blackish inner border;
discal dots at end of cell, separate, distinct, blackish;
terminal dots obscure, more or less confluent. Hind
wing white to pale smoky fuscous; the veins but faintly
darkened. Alar expanse, 12-16 mm.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 279
Male genitalia with apical process of gnathos
V-shaped, the prongs narrowly triangulate. Harpe
with strong transverse sclerotized ridge at base of
cucullus and projecting slightly beyond lower margin
of harpe; costa smooth; apex angled. ‘Transtilla with
flaring apices of its elements divergent. Aedeagus
moderately stout.
Female genitalia show some individual variation in
the number of teeth in the signum (4 to 6). The shape
of the shield behind genital opening is also variable
(compare figs. 1072 and 1078).
Type Locaitizs: California (gilvescentella, in Paris
Mus.); Los Angeles, Calif. (nigrella, in USNM).
Foop piants: Raisins, cotton, Gaura parviflora.
These food-plant records from reared specimens in the
National Collection. Moths have been frequently seen
about stored unprocessed raisins in California and have
been reared from larvae feeding on the raisins, but
apparently doing only minor injury. The life history
has not been worked out. The habits of the larvae are
presumably similar to those of Vitula and Ephestia.
Distrisution: Unitep States: Colorado, Denver;
Utah, Eureka (July, Aug.), Provo (June, July, Aug.,
Sept.), Stockton (July), Vineyard (June); New Mexico,
Mesilla (Apr., July); Arizona, Fish Creek Station
(Sept.), Tempe (Apr.); California, Blythe (Sept.),
Claremont, Fresno (July), Laguna Beach, Loma Linda
(July), Los Angeles (Mar., Apr., July), Pasadena
(June), Riverside (Apr., June, July), Sacramento
(Mar.), San Diego (Mar., May, June), San Gabriel
(July) ; Washington, Almota (Apr.), Bellingham (Aug.),
Pullman (June, July, Aug., Sept.), Snake River (June),
Wenatchee (Aug.). Canapa: British Columbia, Kere-
meos (Aug.), Seton Lake (Aug.), Victoria, Wellington.
México: Baja California, Ensenada (Apr.)
The name nigrella applies only to a suffused color
form in which the dark dusting on the central area of
forewing extends into the terminal area. In series from
any given locality intergrades occur between it and
typical gilvescentella. The species appears to be con-
fined to the North American region from the Rocky
Mountains westward.
573. Ephestiodes infimella Ragonot
Fieure 590
Ephestiodes infimella Ragonot, N. Amer. Phycitidae, p. 16, 1887;
(Ragonot and Hampson), Monograph, pt. 2, pp. 264, 265,
1901.—Hulst, Phycitidae of N. Amer., p. 194, 1900.—
Forbes, Cornell Mem. 68, p. 635, 1923.—McDunnough,
Check list, No. 6380, 1939.
Forewing similar in pattern and color to the more
suffused specimens of gilvescentella but averaging
smaller. In many specimens, especially those where the
basal area is pale and contrasted against the rest of the
wing, there is more or less shading of reddish luteous;
color as variable as in gilvescentella. Hind wing pale to
dark smoky fuscous. It is the common species in
eastern and central United States. Alar expanse,
10-15 mm.
Male genitalia with the prongs of apical] process of
gnathos slender, cylindrical, and widely spaced at base.
Harpe with the transverse, sclerotized ridge at base of
costa not projecting beyond lower margin of harpe;
costa irregularly serrate at middle (a variable and not
too reliable character, costa however never entirely
smooth under high magnification); apex rounded.
Transtilla with apices of its elements divergent at their
apices but less flaring than those of gilvescentella.
Aedeagus moderately stout. Female genitalia not
essentially different from those of gilvescentella.
Type Locauity: North Carolina (type in Paris Mus.).
Foop pxrants: Wild cherry, seeds of Ambrosia
(Forbes). The wild cherry record is from a specimen
reared from larvae collected by A. Busck at Cape
Henry, Va.
Distripution: Unitep States: Massachusetts, Mar-
tha’s Vineyard (Aug.); Connecticut, East River; New
Jersey, Anglesea (June), Montclair (Aug.); Pennsyl-
vania, Hazleton (July), Oak Station (July) ; Maryland,
Hyattsville (Aug.), Plummers Isl. (May, July) ; District
of Columbia, Washington (Aug.); Virginia, Cape Henry
(July) ; North Carolina, Tryon (Aug.); Kentucky (Aug.) ;
Illinois, Chicago (June), Putnam County (May, Sept.) ;
Iowa; Missouri, St. Louis (June); Arkansas, Washing-
ton County (July); Teras, Burnet County. Also re-
corded by Hampson from Colombia; but this record is
undoubtedly based on a misidentification. From all
available evidence the species is limited in its distribu-
tion to North America.
574, Ephestiodes erythrella Ragonot
Figures 591, 1074
Ephestiodes erythrella Ragonot, N. Amer. Phycitidae, p. 16, 1887;
(Ragonot and Hampson), Monograph, pt. 2, pp. 264, 266,
1901.—Hulst, Phycitidae of N. Amer., p. 195, 1900.—
McDunnough, Check list, No. 6384, 1939.
Eurythmia coloradella Hulst, Canadian Ent., vol. 32, p. 175,
1900.—McDunnough, Check list, No. 6390, 1939. (New
synonymy.)
Ephestiodes benjaminella Dyar, Proc. U. 8S. Nat. Mus., vol. 27,
p. 922, 1905.—McDunnough, Check list, No. 6383, 1939.
(New synonymy.)
Forewing vinous red, dusted with whitish gray in the
basal area and with some faint blackish dusting along
costa and on the veins; in the dry areas of Colorado and
Utah the ground color somewhat paler, with consider-
able whitish gray dusting in the area between the trans-
verse lines and little or no appreciable blackish dusting
on costa and veins; transverse lines whitish, somewhat
more distinct than in gilvescentella and similarly shaped
and dark margined (in some specimens they appear
curved, in others straight, due to the extent of the black-
ish shade bordering them, but in reality slightly angled
as in the other North American species) ; discal dots ob-
scure. Hind wings pale to moderately dark smoky
fuscous. Alar expanse, 13-17 mm.
Male genitalia similar to those of infimella except
apex of harpe angulate. Female genitalia showing no
distinctive specific characters.
280
Typr wocatitius: California (erythrella, in Paris
Mus.); Colorado (Coloradella, in AMNH, ex Rutgers);
Kaslo, British Columbia (benjaminella, in USNM).
Foop pLant: Unknown.
DistrisutTion: Unitep States: Colorado; Wyoming,
Jackson Hole (June); Utah, Eureka (May), Provo
(June, July); California, Alfa (Placer County, July),
Baxter (Placer County, July), Camp Baldy (San Ber-
nardino Mts., July), Makehlumne Hill (June); Wash-
ington, Pullman (June, July, Aug.). Canapa: British
Columbia, Goldstream (Sept.), Kaslo (June, July, Aug.),
Wellington. A long series in the National Collection
from Utah.
Like gilvescentella and infimella, erythrella is variable
in coloration and intensity of markings and in the shape
of the terminal margin of the sclerotized shield behind
genital opening of the female. Extremes of individual
variation in the shield are shown in figures 1074a, b.
The same amount of variation occurs in gilvescentella.
It has much the same range as the latter, but need not
be confused with it, for in dark or light specimens it can
be readily identified by its distinctly reddish ground
color. Occasional specimens of injimella are also red-
dish but that species does not occur in the same areas
as erythrella. I am unable to find any character of
specific value to distinguish either coloradella or ben-
gaminella from California examples of erythrella,so I am
sinking the names in synonymy.
575. Ephestiodes mignonella Dyar
Fiaures 592, 1075
Ephestiodes mignonella Dyar, Proc. Ent. Soc. Washington, vol.
10, p. 113, 1908.—McDunnough, Check list, No. 6381,
1939.
Forewing very pale grayish white; a blackish streak
on costa at base and, in many specimens, a subbasal
black streak on median vein; transverse lines as in pre-
ceding species, white, the antemedial with a strongly
contrasted, outer black band and the subterminal with a
narrower inner black band, especially pronounced on
costal half of wing (sometimes the black borders partially
obscured on lower half of wing by pale putty-colored
scaling); discal dots fused, black; terminal dots fainter,
confluent. Hind wings whitish, with a smoky tinge
at apex, along termen, and faintly on the veins, espe-
cially on the females. Alar expanse, 12-14 mm.
Male genitalia similar to those of injfimella except:
Harpe with apex angled; enlarged apices of elements of
transtilla parallel and facing each other. Female gen-
italia showing no distinctive specific characters.
TypE Locauity: Kerrville, Tex. (type in USNM).
Foop pLrant: Unknown.
Represented by 13 specimens from the type locality
in the National Collection (Apr.).
The species is easily distinguished by the very pale
ground color and strongly contrasted black markings of
forewing.
UNITED STATES NATIONAL
MUSEUM BULLETIN 207
576. Ephestiodes erasa, new species
Fieures 588, 1077
Forewing very pale gray with some pale putty-
colored scaling on lower half, especially towards base;
transverse lines faint, distinguishable only under low-
powered magnification; their dark borders nearly
obliterated; discal dots very faint, obsolescent in some
specimens. Hind wings smoky white, little paler than
forewing. Alar expanse, 11-13 mm.
Male genitalia with transverse ridge projecting
beyond lower margin of harpe (more so than in gilvescen-
tella); lower margin of cucullus incurved just beyond
transverse ridge; cucullus somewhat reduced. Female
genitalia exhibiting no distinguishing specific characters.
Tyrer tocatity: Lake Alfred, Fla. (type m USNM,
61387).
Foop prant: Unknown.
Described from male type and six female paratypes
from the type locality, May 6 and July 2, 1929, L. J.
Bottimer, collector. In addition to the type series I
have before me a female from Gainesville, Fla. (Apr.),
D. M. Bates, and one other female from Winter Park,
Fla. Guly 23, 1939), H. Fernald. All the foregomg
were collected at light.
The species is at once distinguished by its pale, almost
unmarked forewings.
577. Ephestiodes lucidibasella Ragonot
Fieures 593, 1081
Ephestiodaes lucidibasella Ragonot, Nouv. Gen., p. 35, 1888;
Monograph, pt. 2, pp. 264, 265, 1901.
Forewing rosy gray, the median and terminal areas
heavily and evenly dusted with blackish; the transverse
lines decidedly oblique, widely spaced on costa. Rago-
not’s figure (Monograph, pl. 34, fig. 6) shows a species
with an extension of the pale ground color along the
lower fold, and according to Clarke’s notes the figure is
a good representation of the type. Hind wings semi-
transparent, whitish with a slight smoky tint. Alar
expanse, 16 mm.
Gnathos of male genitalia with prongs of apical
process somewhat longer than those of the North Amer-
ican species, slender, widely spaced at base and curving
apart slightly toward their apices. Harpe without
trace of asclerotized, transverse ridge at base of cucullus;
edge of costa minutely and irregularly serrate; apex
evenly rounded. ‘Transtilla with the enlarged apices
of its elements concave, parallel, and facing each other.
Female genitalia distinguished by the shape of the
sclerotized shield from ductus bursae behind genital
opening.
TYPE LOCALITY: Quillota, Chile (type in Paris Mus.).
Foop piant: Unknown.
All known specimens are from Chile. In the Paris
Museum, according to Clarke’s notes, are three males
from the type locality and one male from Valparaiso.
A female paratype is in the British Museum. There
are no examples in the U. S. National Museum.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 281
578. Ephestiodes productella Ragonot
Ficure 1078
Ephestiodes productella Ragonot, Nouv. Gen., p. 36, 1888; Mono-
graph, pt. 2, pp. 264, 265, 1901.
This species is known only from the female type in
Paris. According to Clarke’s note it answers very well
to Ragonot’s description but not to his figure in the
Monograph (pl. 35, fig. 1).
The genitalia are characteristic and should easily
identify the species when other females are discovered.
The shield behind genital opening has an angulate pro-
jection from posterior margin as in North American
species, but is appreciably larger and the sclerotized
band below genital opening is much stouter and broader.
Alar expanse, 15 mm.
TYPE LocaLity: Given by Ragonot as “Am. Mér.”
in original description, and as Chiriqui, Colombia, in
Monograph (type in Paris Mus.). Nothing on type to
indicate the exact locality, but undoubtedly from
tropical America.
Foop prant: Unknown.
579. Ephestiodes indentella Dyar
Figure 1076
Ephestiodes indentella Dyar, Ins. Insc. Menstr., vol. 3, p. 89, 1915.
Forewing pale gray, the median area shading with
blackish scaling; the transverse lines widely spaced on
costa, strongly contrasted (especially the antemedial),
white; antemedial line decidedly oblique, notched at
lower fold and with a black outer border; subterminal
indented at vein 6 and slightly at lower fold, and with
a narrow black inner border; discal dots separate; ter-
minal dots confluent, not extending to either costa or
tornus. Hind wing pale smoky fuscous; veins and
terminal margin but slightly darkened. Alar expanse,
12.5-17 mm.
Female genitalia with a wide, deep notch in posterior
margin of shield behind genital opening; sclerotized
band below genital opening narrow, sinuous; eighth-seg-
ment collar broadly and deeply sclerotized on dorsum,
its anterior margin produced and rounded.
Male unknown.
TyprE Locauity: Bermuda (type in USNM).
Foop prant: Unknown.
Known only from Bermuda. In addition to the type
series (Apr.) I have before me 6 females from the
British Museum (Mar., Apr.). The species is readily
identified by its genitalia. It and plorella are tenta-
tively assigned to the first Ephestiodes species group,
but accurate placement of them will have to wait upon
discovery of males.
580. Ephestiodes plorella Dyar
Ficures 109, 1080
Ephestiodes plorella Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 344,
1914.
Eurythmia vestilla Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 339,
1914 (new synonymy).
Forewing rather pale brownish gray, uniformly
300329—56——19
colored except for a somewhat paler basal area and a
rather broad brownish outer band along the antemedial
line; antemedial band but slightly oblique; subterminal
line obscure, whitish with only faint trace of a dark
inner border; discal dots obscure, when distinguishable,
more or less confluent; terminal dots faint, confluent.
Hind wing whitish, some of the veins and terminal area
very faintly darkened. Alar expanse, 11-14 mm.
Female genitalia with signum developed as a flat plate
with serrate edge, the ductus seminalis arising from a
hole in the plate; bursa with a few scobinations at junc-
tion with ductus bursae; ductus bursae sclerotized for
only half its length; no sclerotized band below genital
opening; posterior margin of large sclerotized shield
behind genital opening evenly rounded except for a very
slight narrow central notch; collar of eighth segment
natrow but deeply sclerotized on dorsum, its produced
anterior margin concave and with asmall shallow central
notch.
Male unknown.
Type Locauity: Corazal, Canal Zone, Panama
(plorella and vestilla, in USNM).
Foop pirant: Unknown.
A rather suffused, poorly marked species without much
to distinguish it superficially from faded examples of our
North American gilvescentella or infimella. The female
genitalia, however, are markedly different from those of
any other species in the genus. The absence of a sclero-
tized band below genital opening, the shapes of the dor-
sal genital plate and eighth-segment collar, and the pe-
culiar signum at once distinguish them. The venation
also differs in that vein 3 of forewing is very closely ap-
proximate to, connate, or short stalked with 4-5, and
the latter are very long stalked. Dyar’s vestilla was
placed by him in Zurythmia on the belief that veins 4
and 5 of forewing were united as they appear to be on
superficial examination; but denuded wings (fig. 109)
show them long stalked. The female genitalia of his
type of vestella are identical with those of his plorella.
Genus Ephestiodes, Species 581: ZH. stictella
[Transtilla complete, with greatly produced, asymmetrical,
caudal projections. Harpe with apex of sacculus produced into
afree arm. Highth abdominal segment of male with tufts.]
581. Ephestiodes stictella (Hampson), new combination
Figure 596, 1082
Unadilla stictella Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7,
p. 255, 1901.
Ephestiodes uniformella Hampson, Ann. Mag. Nat. Hist., ser. 7,
vol. 7, p. 256, 1901 (new synonymy).
Ephestiodes granulella Hampson, Ann. Mag. Nat. Hist., ser. 7, vol.
7, p. 256, 1901 (new synonymy).
The type of stictella is afemale. A photograph of its
genitalia definitely fixes the species to what I here iden-
tify in long series from the Cornell Collection from Puerto
Rico and St. Croix. The type of uniformella is a male
without abdomen and according to Clarke a stained spec-
imen., A photograph shows it to be a dark specimen but
no darker than many examples of the St. Croix series.
282
The type of granulella is a male and a photograph of its
genitalia fixed it. I have before me other specimens of
stictella from the Bahamas and of granulella from Ja-
maica that agree in all characters. Hampson’s reference
of stictella to Unadilla ignored the diagnostic venational
character of that genus, the vertical discocellular vein of
hind wing. In stictella the discocellular vein is obviously
curved. ‘The species is variable in color, ranging from
very dark to whitish gray, the ground color (dark or
light) being rather uniform over the forewing, the basal
area no darker or lighter than the median and terminal
areas. Antemedial band rather broad, whitish, oblique
and nearly straight, outwardly bordered on costal half
by a narrow blackish line; subterminal line narrow, paral-
lel and near to termen, slightly irregular, whitish bor-
dered inwardly towards costa by a thin, faint, blackish
line; discocellular spots more or less obsolescent, when
distinct, separate and blackish. Hind wing whitish to
pale smoky fuscous, shaded with smoky fuscous towards
apex and termen. Alar expanse, 10-12 mm.
Male genitalia with transtilla elements fusing at a
point and thence developed into two extended, curving,
asymmetrical arms; harpe with sacculus produced at ex-
tremity into a clasperlike free arm; costa smooth; apex
of harpe bluntly pointed; vinculum triangulate, shorter
than broad; aedeagus rather slender. Female genitalia
with ductus bursae shorter than bursa, sclerotized
throughout its length; a narrow flaring sclerotized plate
below genital opening; eighth-segment collar divided, its
apophyses separated from the dorsal part and firmly at-
tached to the broad shield behind genital opening (fig.
1082b).; dorsal portion of collar (ig. 1082a) produced an-
teriorly into an elongate tongue and laterally into curved,
projecting arms.
Type Locaurry: Nassau, Bahamas (stictella, uniform-
ella, and granulella, in BM).
Foop prant: Unknown.
DistriputTion: Banamas: Nassau. Jamaica, Runa-
way Bay (Mar.). Purrro Rico: Coamo Springs (Apr.),
Palmas Abajas (June, July), San German (Apr., July),
Puerto Real (Vieques Isl., Apr.). Virein IsLanps:
Kingshill (St. Croix; May, Oct., Nov., Dec.).
This species on characters of the genitalia seems to de-
serve a separate generic designation; but until the males
of the other tropical species (described from females) are
known it seems best to retain it in H’phestiodes, with
which it agrees on all other than genitalic characters.
Genus Hphestiodes, Species 582: EH. noniella
[Transtilla complete, a narrow band with central loop. Harpe
simple. Eighth abdominal segment of male simple.]
582. Ephestiodes noniella Dyar
Figures 597, 1079
Ephestiodes noniella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 345,
1914.
Forewing (to the naked eye) gray, the basal area with
an olivaceous tint; under magnification the remainder
of the wing shows a strong dusting of rosy scaling; ante-
UNITED STATES NATIONAL MUSEUM BULLETIN 207
medial line faint, oblique, narrow, whitish, and faintly
bordered outwardly near costa by a narrow dark line;
subterminal line almost obliterated; discal dots not dis-
tinguishable. Hind wing pale smoky fuscous; the veins
and terminal margin but slightly darkened. Wighth
abdominal segment of male simple. Alar expanse,
9.5-10 mm.
Male genitalia with the prongs of apical process of
gnathos well separated but somewhat reduced. Trans-
tilla complete, a narrow band with a central loop.
Harpe simple; costa smooth; apex narrowly rounded.
Aedeagus short; penis without armature. Female geni-
talia like those of typical Ephestiodes except: Shield
behind genital opening smaller; sclerotized band below
genital opening developed as a half oval, caudally pro-
jecting shield.
Typn tocauity: La Chorrera, Panamd (type in
USNM).
Foop prant: Unknown.
Known only from the type series, two females from
the type locality (May) and one malefrom Rio Trinidad,
Panam4 (Sept.).
The species is an abnormal Hphestiodes on male char-
acters (the complete, thin, looped transtilla, lack of
cornutus on penis, and absence of tufts on eighth
segment), but separate generic designation does not
seem warranted at this time.
171. Genus Azaera Schaus
Azaera Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11, p. 250,
1918. (Type of genus: Azaera muciella Schaus.)
Calamophleps Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 342, 1914.
(Type of genus: Calamophleps squalidella Dyar.)
Characters of typical H’phestiodes except vein 3 of
forewing from the stalk of 4—5.
This genus is of doubtful status. There are nothing
but differences of specific value in the genitalia to
separate it from the type of Hphestiodes and, in the
latter genus, occasional specimens of gilvescentella have
vein 3 of forewing connate cr even very shortly stalked
with 4-5. However, none that I have seen (and | have
examined long series of H’phestiodes) shows such short
stalking on both forewings and none exhibits the con-
siderable stalking of Azaera (see fig. 110). Asa definite
stalking of 3 with 4 and 5 is a rare character in New
World Phycitinae, we had better retain the generic
separation for the present.
Dyar was in error in assuming that Azaera Schaus
was a homonym of Azara D’Orbigny, so Schaus’ name
must take precedence over Calamophleps.
583. Azaera muciella Schaus
Figures 110, 594, 1084
Azaera muciella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol. 11,
p- 250, 1913.
Calamophleps squalidella Dyar, Proc. U. 8. Nat. Mus., vol. 47,
p. 342, 1914.
Forewing color and markings similar to those of
Ephestiodes plorella Dyar except that subterminal line
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB
is always distinguishable. The species can be identified
satisfactorily only by its venation and genitalia. Hind
wing pale smoky fuscous; veins and terminal margin
darker. Alar expanse, 11-13 mm.
Male genitalia with prongs of apical process of gnathos
rather stout and curved towards each other at their
apices. Apices of elements of transtilla greatly de-
veloped, widely flaring. Harpe with transverse ridge
at base of cucullus projecting beyond lower margin of
harpe; lower margin of cucullus with a slight notch at
apex. Female genitalia with sclerotized band below
genital opening forming a rather broad oval shield;
ductus bursae much longer than bursa, sclerotized for
no more than half its length; dorsal sclerotized portion
of eighth-segment collar a rather broad band.
TYPE LOCALITIES: Juan Vifias, Costa Rica (muciella,
in USNM), Porto Bello, Panama (squalidella, in USNM).
Foop prant: Unknown.
DistrisuTion: Costa Rica: Juan Vifias (Feb.).
GuatEmata: Cayuga (May). PanamA: Corozal (Mar.
Apr., May), La Chorrera (May), Porto Bello (May,
Dec.).
Dyar acknowledged the synonymy of muciella and
squalidella in his description of the latter. It is borne
out by their genitalia. Both types are females.
584. Azaera nodoses (Dyar), new combination
Ficure 1085
mea nodoses Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 348,
4,
Smaller and paler than muciella, with the dark borders
indicating the transverse lines fragmentary and obscure;
but chiefly distinguished by its genitalia. Alar expanse,
10 mm.
Female genitalia with ductus bursae short (no longer
than bursa) and sclerotized for its entire length; dorsal
sclerotized portion of eighth-segment collar a narrow
band.
Type tocauity: Corozal, Panamé (Apr., type in
USNM).
Foop prant: Unknown.
Known only from the unique female type.
585. Azaera lophophora (Dyar), new combination
Calamophleps lophophora Dyar, Proc. U. 8. Nat. Mus., vol. 47,
p. 343, 1914
Similar to muciella but darker, smoky gray; the
transverse lines and their dark borders more distinct.
May be only a dark form of muciella; but this cannot
be determined until more specimens are available, as
the two females of the type series (the only specimens
known) lack abdomens. Alar expanse, 11-12 mm.
Typ Locatity: Porto Bello, Panamdé (May, type in
USNM).
Foop piant: Unknown.
The female paratype is from La Chorrera, Panama
(May).
283
172. Genus Moodna Hulst
Moodna Hulst, Phycitidae of N. Amer., p. 193, 1890.—Ragonot,
Monograph, pt. 2, p. xiii, 1901—Hampson, in Ragonot,
Monograph, pt. 2, p. 267, 1901.—Forbes, Cornell Mem. 68,
p. 636, 1923. (Type of genus: Moodna pelviculella Hulst.)
Tongue well developed. Antenna pubescent; shaft
of male with a short, shallow sinus towards base.
Labial palpus oblique, subcylindrical, reaching vertex,
somewhat rough scaled; third segment nearly as long
as second. Maxillary palpus filiform, short. Fore-
wing smooth; 10 veins; vein 2 from before, but rather
near, lower outer angle of cell; 3 from the angle; 4 and 5
stalked (for at least half their lengths), the stalk shortly
separated from 3 at base; 6 from below upper angle!of
cell, straight; 8 and 9 united (9 absent); 10 from the
cell, separated from 8 at base; male with a strong costal
fold enclosing hair tuft. Hind wing with vein 2 from
well before lower outer angle of cell; 3 and 5 from the
angle, connate, rarely (in individual specimens) very
shortly stalked; 7 and 8 anastomosed to or almost to
costa (8 absent or a mere vestige at costa); cell less than
half (but more than a third) the length of the wing;
discocellular vein curved. Eighth abdominal segment
of male with compound, dorsal tufts.
Male genitalia with apical projection of gnathos a
broad, short, pointed hook. Uncus broad; terminal
margin broad, straight or but slightly rounded. Trans-
tilla incomplete. Harpe with apex broadly rounded;
two strongly sclerotized subbasal projections from
costa. Anellus a narrow semicircular sclerotized band.
Aedeagus long, straight, not appreciably tapering,
simple; penis bearing one or two elongate, thin, weakly
sclerotized plates and a few sclerotized wrinklings,
otherwise unarmed. Vinculum long, triagulate.
Female genitalia with signum a small cluster of
rounded, short, projecting discs; bursa copulatrix small,
wrinkled, the wrinklings weakly sclerotized; ductus
bursae long (much longer than bursa), sclerotized for a
short distance from genital openings; ductus seminalis
from bursa close to signum; an extended ventral lobe
from membranous area between ovipositor and eighth-
segment collar (figs. 1086a and 1087a).
The species of the Moodna-Vitula complex offer a
difficult problem in generic placement. Typical species
of either genus are obviously different in characters of
venation, genitalia, and male antennae; but there are
a number of aberrant species that possess characters of
both Moodna and Vitula and will fit in neither genus.
If we should attempt to unite all under one genus, we
should have a group that could not be defined. I have
therefore divided the complex into small units, resur-
recting Hulst’s Manhatta and erecting separate genera
for some new species and the aberrant tropical species
that have been referred to Moodna, or misplaced else-
where. One name that appears under Moodna in our
lists (Z’phestia nigrella Hulst) has veins 4 and 5 of fore-
wing united and is treated elsewhere in this paper as
Caudellia nigrella (p. 293). It has the habitus of
Moodna, similar male antennae, is similar in color and
maculation to its type, and is apparently derived from
284
Moodna; but is structurally an advanced development.
As here defined, Moodna is limited to two species from
the United States and México (ostrinella Clemens and
bisinuella Hampson). The genus is easily distinguished
from other genera in venational group C by the follow-
ing combination of characters: Male antenna with a
shallow sinus in shaft; eighth abdominal segment of
male with tuft; hind wing with vein 2 from well before
angle of cell and veins 3-5 connate or very shortly
stalked; gnathos terminating in a short, simple hook;
transtilla incomplete.
Two species (Moodna olivella Hampson and Hornigia
clitellatella Ragonot) that Hampson (Ragonot Mono-
graph, pt. 2, pp. 268, 269) includes in the genus are un-
known to me, and I am unable to place them generically.
They are treated briefly at the end of this paper.
On the strength of the supposed synonymy of Man-
hatta Hulst (=Hornigia Ragonot, 1887), Hampson
cites the European biviella Zeller as the type of Moodna.
This, of course, is inadmissible, for not only was biviella
not among the species originally included in Moodna,
but in describing his genus Hulst designated pelviculella
as its type.
586. Moodna osirinella (Clemens)
Fieures 114, 599, 1086
Ephestia ostrinella Clemens, Proc. Acad. Nat. Sci. Philadelphia,
1860, p. 206.—Hulst, Phycitidae of N. Amer., p. 220, 1890.
Horginia obtusangulella Ragonot, N. Amer. Phycitidae, p. 16,
1887.
Moodna pelviculella Hulst, Phycitidae of N. Amer., p. 194, 1890.
Manhatta obtusangulella (Ragonot) Hulst, Phycitidae of N.
Amer., p. 197, 1890.
Moodna obtusangulella (Ragonot) Hampson, in Ragonot, Mono-
graph, pt. 2, p. 269, 1901.
Manhatta ostrinella (Clemens), Hulst, U. S. Nat. Mus., Bull. 52,
p. 486, 1903.
Moodna ostrinella (Clemens) Barnes and McDunnough, Check
list of the Lepidoptera of Boreal America, No. 5795, 1917.—
Forbes, Cornell Mem. 68, p. 636, 1923.—McDunnough,
Check list, No. 6896, 1939.—Darlington, Trans. Amer. Ent.
Soce., vol. 73, p. 91, 1947.
Forewing blackish fuscous; basal area strongly shaded
with a deep violet red; terminal area also more or less
shaded with the same reddish color; antemedial le
distinct, rather wide, nearly straight, white; subterminal
line obscure, parallel to termen, slightly and irregularly
denticulate; discal spots distinct, more or less confluent
and usually set off by some surrounding whitish dusting
which extends, in well marked specimens, to costa.
Hind wing smoky white to pale fuscous; veins darkly
outlined and a narrow dark shade along terminal mar-
gin. Alar expanse, 11-17 mm.
Male genitalia with inner costal projection from harpe
long, slender, tapering, vertical from costa; outer pro-
jection curved outwardly, the two projections pointed
away from each other. Uncus with sides nearly paral-
lel. Vinculum evenly tapering. Female genitalia with
ductus bursae weakly sclerotized toward genital opening.
Typ LOCALITIES: Pennsylvania [?] (ostrinella, type
lost); Texas (obtusangulella, in Paris Mus.); Newburgh,
N. Y. (pelviculella, in AMNH, ex Rutgers).
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Foop puiants: Betula, Rhus, Quercus, rose, pear,
peach, apple, loquat, iris, cotton, Pinus. The larva is
more or less a scavenger, feeding on dried seeds and
mummied fruits, on dry rose buds, and in galls on roses,
in acorns and old cotton bolls. Its favorite food in the
neighborhood of Washington, D. C., seems to be the
racemes of Rhus. It has much the habits of an E’phestia
except that it is an outdoor insect and does not attack
stored products to any extent.
Distrisution: Unirep Srares: Maine; Vermont,
Highgate (June); Massachusetts, Cohasset (July); Con-
necticut, Kast River (Aug.); New York, East Aurora
(Aug.), Ilion (Aug.), Newburgh (July), Rochester (Aug.);
New Jersey, Dayton (July), Greenwood Lake (June),
Pine Brook; Pennsylvania, Oak Station (Aug.), West
Chester (July); Maryland, Hyattsville; District of
Columbia, Washington (Mar., Apr., July); Virgina,
Arlington County (Aug.), Blacksburg (June), Cape
Henry (Aug.), Great Falls (Mar., Apr., May); North
Carolina, Southern Pines; Florida, Camp Pinchot
(Dec.), Hastings (Mar.), Lake Alfred (Aug.), Miami
(July, Aug.), Monticello (Mar.), ‘Southern Florida”
(June); Texas, Houston (July), Kountze (Apr.), Vic-
toria (Mar., May); Ohio, New Richmond (Apr.);
Iitinois, Oconee (Aug.); Jowa, Ames (Aug. Sept.).
Canapva: Quebec, Chelsea (Apr.), Meach Lake; On-
tario, Trenton (July). Distribution apparently limited
to United States and Canada east of the Rockies.
587. Moodna bisinuella Hampson
Ficures 600, 1087
Moodna bisinuella Hampson, in Ragonot, Monograph, pt. 2,
p. 268, 1901.
Forewing blackish fuscous with transverse lines
obscure; on fresh specimens some reddish scaling in
basal area and along the folds (visible only under
magnification) ; antemedial line as in ostrinella but very
faint; discal spots usually distinguishable and more or
less confluent. Hind wing whitish with apical area
somewhat smoky; the veins and a line along terminal
margin considerably darker. Alar expanse, 17-22 mm.
Male genitalia with inner costal projection from
harpe curved outwardly; outer projection curved in-
wardly, the two projections pointed toward each other.
Uncus with sides strongly convex. Vinculum sharply
angled at anterior end. Aedeagus considerably stouter
than that of osirinella. Female genitalia with ductus
bursae strongly sclerotized and striated towards genital
opening.
TYPE LOCALITY: Orizaba, México (type in USNM;
paratypes in BM).
Foop pLtant: Corn.
Distrisution: México: Matamoros (July, Aug.),
Orizaba, Tehuacén (Sept., Oct.). Unitmsp Sratzs:
Texas, Crystal Springs (Oct.), Weslaco (June).
Larvae of bisinuella have been frequently intercepted
in ears of green corn from México at border ports by
the Division of Foreign Plant Quarantine of the
U. S. Bureau of Entomology and Plant Quarantine.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
The species has apparently invaded the United States
from México and has become established in a few Texas
localities. A larva indistinguishable from bisinuella
has also been intercepted in green corn from British
Honduras but no adults from that locality have been
reared to verify the identification.
Hampson’s description of the male antenna is mis-
leading. He states that it has ‘deux sinus profonds 4
la base de la tige.’’ The shaft is slightly swollen from
the second to the seventh segments and beyond the
seventh segment has a single short shallow sinus. His
male type and several reared males are before me and
each of them shows an antennal sinus like that of
ostrinella.
173. Genus Vitula Ragonot
Vitula Ragonot, N. Amer. Phycitidae, p. 14, 1887; Monograph,
pt. 2, p. 81, 1901.—Hulst, Phycitidae of N. Amer., p. 178,
1890.—Forbes, Cornell Mem. 68, p. 631, 1923. (Type of
genus: Vitula dentosella Ragonot).
Eccopsia Hulst, U. 8. Nat. Mus. Bull. 52, p. 430, 1903.—Dyar,
Proc. Ent. Soc. Washington, vol. 6, p. 158, 1904. (Type of
genus: Vitula serratilineella Ragonot.)
Tongue well developed. Antenna simple. Labial
palpus upturned in male, more oblique in female,
reaching a little above vertex; subcylindrical, somewhat
rough scaled; third segment slightly shorter than
second. Maxillary palpus filiform, short. Forewing
smooth; 10 veins; vein 2 from before, but rather near
lower outer angle of cell; 3 from the angle; 4 and 5
stalked for half their lengths, the stalk separated from
3 at base; 6 from below upper angle of cell, straight;
8 and 9 united (a vestige of 9 present occasionally on
one side or another of individual specimens); 10 from
the cell, approximate at base to 8; male with a strong
costal fold enclosing hair tuft. Hind wing with vein 2
from well before lower outer angle of cell: 3 and 5 from
the angle, approximate at base; 7 and 8 anastomosed
for all or most of their lengths beyond cell (if present,
8 a mere vestige at costa); cell about one-third the
length of the wing (with veins 2, 3, and 5 appreciably
long); discocellular vein curved. Eighth abdominal
segment of male with compound dorsal tufts.
Male genitalia with apical process of gnathos a short,
bluntly pointed hook. Uncus broad; terminal margin
broadly rounded. ‘Transtilla complete, an angulate
bridge. Harpe with apex rounded; costa slightly
humped near middle but without projections. Anellus
a narrow, semicircular, sclerotized band with very short
lateral lobes. Aedeagus long, straight, not tapering;
penis bearing a single elongate, thin, weakly sclerotized
plate, otherwise unarmed. Vinculum stout, slightly
longer than broad, not appreciably tapering; terminal
margin broad.
Female genitalia with signum a small cuplike dise or
a cluster of two or more such discs; bursa copulatrix
small, more or less scobinate, the scobinations (except
in laura) prominent asa partial girdlenearsignum;ductus
bursae appreciably longer than bursa, strongly sclero-
tized for a short distance from genital opening and with
285
a projecting, angulate, sclerotized plate behind genital
opening; ductus seminalis from bursa close to signum;
ventral membrane between eighth-segment collar and
ovipositor not extruded.
As here defined Vitula is distinguished from Moodna
by its simple male antenna, complete transtilla, simple
harpe (without costal projection) and the approximate,
rather than connate, condition of veins 3 and 5 of hind
wing. On the strength of the synonymy of dentosella
and edmandsw Hulst cites edmandsii as type of the
genus and is followed in this by Ragonot in his Mono-
graph. This is not admissible nomenclatorially, as only
dentosella was included under the original description
of the genus and was definitely named at that time as
type by Ragonot.
588. Vitula edmandsae (Packard)
Figures 115, 605
Nephopteryx edmandsii Packard, Proc. Essex Inst., vol. 4, p. 120,
1864; Guide to the study of insects, p. 331, 1869.
Vitula dentosella Ragonot, N. Amer. Phycitidae, p. 14, 1887.
Vitula edmandsii (Packard) Hulst, Ent. Amer., vol. 5, p. 156,
1889; Phycitidae of N. Amer., p. 178, 1890.—Ragonot,
Monograph, p. 2, p. 82, 1901.—Forbes, Cornell Mem. 68,
p. 631, 1923.—Frison, Ann. Ent. Soc. Amer. vol. 19, p. 226,
1926.—MceDunnough, Check list, No. 6323, 1939.—Corbet
and Tams, Proc. Zool. Soc. London, vol. 113 (B), p. 64, 1943.
Forewing gray more or less dusted with blackish
fuscous, especially in the median area (between the
transverse lines) ; on some specimens a reddish ocherous
shade along lower fold and inner margin; pale ante-
medial line obscure, defined chiefly by its contrasted,
blackish outer border, the latter from costa near middle,
outwardly angled at cell and usually with a slight notch
at lower fold; subterminal line parallel to and well in
from terminal margin, rather deeply notched at vein 6,
and slightly so at lower fold, bordered inwardly by a
thin black line; discal dots black, rarely separated,
usually fused into a line along discocellular vein. Hind
wing pale smoky fuscous, veins darker, a narrow dark
line along terminal margin. Alar expanse, 15-22 mm.
Genitalia as given for the genus. The female bursa
shows more or less minute scaling near to and at junc-
tion of bursa and ductus bursae.
TyPE LocauitiEs: Bridport, Vt. (edmandsae, ia
MCZ); North Carolina (dentosella, in Paris Mus.).
Foop: Honeycomb of bees (larvae feeding on wax,
pollen, and comb).
DistriputTion: Unirep Sratrs: Vermont, Bridport;
Massachusetts, Boston (May), Framingham (Sept.);
Connecticut, East River (July); New Jersey, New Lisbon
(Sept.); Pennsylvania, Oak Station (Sept.), New
Brighton (June, Aug., Sept., Oct.) ; District of Columbia,
Washington (June, July, Aug.); Maryland, Plummers
Isl. (June, July, Aug.) ; North Carolina, Black Mountain
(June), Tryon (June, Aug., Sept.) ; Kentucky, Lexington
(Mar.); Missouri, St. Louis (June); Jilinois, Lacon
(Aug.), Oconee (July); Arkansas, Washington County
(July); Florida, Archer (Mar.). Canapa: Ontario,
Trenton (Aug., Sept.) ; Quebec, St. Hilaire (June, Sept.).
286
Occasionally examples of edmandsae and its variety
serratilineella show a vestige of vein 9 on forewing; but
in normal specimens the fusion of 8 and 9 beyond the
cell is complete, and the species obviously belongs in the
group with vein 9 absent. Large pale examples are
quite similar in habitus to Anagasta kuhniella and have
been confused with that species in some collections.
The mistake is very easy to make if one does not exam-
ine the venation of all specimens before him. In his
original description Packard stated that he was naming
the species after Miss A. M. Edmands of Cambridge. I
am therefore emending his name to give it the feminine
ending required by the International Code.
589. Vitula edmandsae serratilineella Ragonot, new status
Figure 1088
Vitula serratilineella Ragonot, N. Amer. Phycitidae, p. 15,
1887.—Hulst, Phycitidae of N. Amer., p. 179, 1890.—
Hampson, in Ragonot, Monograph, pt. 2, p. 83, 1901.—
Dyar, Proc. Ent. Soc. Washington, vol. 5, p. 104, 1903 (de-
scribes egg and larva); Proc. Ent. Soc. Washington, vol. 6,
p. 158, 1904.—Hamlin and Reed, Journ. Econ. Ent., vol. 20,
p. 840, 1927.—EHssig, Insects of western North America,
p. 710, 1929.—Simmons, Reed, and McGregor, U. 8S. Dep.
Agr. cire. 157, p. 38, 1931—McDunnough, Check list, No.
6324, 1939.
Eccopisa serratilineella (Ragonot), Monograph, pt. 2, p. 33,
1901.—Hampson, in Ragonot, Monograph, pt. 2, p. 560,
1901.
Eccopsia serratilineella (Ragonot) Hulst, U. S. Nat. Mus. Bull.
52, p. 430, 1903.
Not structurally different from typical edmandsae and
superficially distinguishable only by its somewhat paler
(more whitish) hind wings. J am keeping the name to
designate a western race which is of some importance
as a minor pest of dried stored fruits in California; but
it is probably not entitled even to this distinction. It
also attacks the honeycombs of bees in the West, but,
as in the Kastern States, does not seem to do any serious
damage to thriving bee colonies. In the literature of
economic entomology it is known as the “dried fruit
moth.” Alar expanse, 14-25 mm.
_Typr tocauity: North America (probably Southern
California; type in Paris Mus.).
Foop: Honeycombs of bees, dried fruits (apples, figs,
raisins, prunes).
Distrisution: Unitep States: California, Alameda
County (Apr.), Claremont, El Segundo (sand dunes,
Mar.), Fresno (May, June, July, Dec.), Humboldt, Los
Angeles (Apr.), Mount View (Apr.), San Diego (Apr.,
June, July), Santa Clara County (Apr.), Santa Cruz
County (Feb., June); Oregon (no definite locality);
Washington, Pullman (Feb., Mar., May, July), Wen-
atchee (Aug., Sept.); Wyoming, Cody (July); Nevada,
Verdi (June); Utah, Stockton (Sept.); Colorado, Fort
Collins; New Mexico, Pecos (June), Raton (Oct.); Ari-
zona, Baboquivari Mts. (July). Camnapa: British Co-
lumbia, Kaslo (Aug., Sept.), Victoria (June, July),
Wellington (July).
UNITED STATES NATIONAL MUSEUM BULLETIN 207
590. Vitula lugubrella (Ragonot), new combination
Fiaure 607
Hornigia lugubrella Ragonot, N. Amer. Phycitidae, p. 17, 1887.
Manhatia lugubrella (Ragonot) Hulst, Phycitidae of N. Amer.,
p. 197, 1890.
Moodna lugubrella (Ragonot) Hampson, 7n Ragonot, Monograph,
pt. 2, p. 270, 1901.—MecDunnough, Check list, No. 6395,
1939.
I have seen no California specimens that match
Ragonot’s description or figure. Evidently the fore-
wing shows some appreciable dusting of reddish scales
and has the antemedial line straight and approximately
vertical (more or less oblique from costa to inner mar-
gin). The genitalia of one of Ragonot’s male paratypes
(from an abdomen loaned by the Paris Museum) are
similar to those of edmandsae except that the vinculum
is longer (like that of pinei) and the harpe tapering from
middle to a more narrowly rounded apex, as shown in
figure 607.
TypxE Locatity: California (type in Paris Mus.).
Foop PLant: Unknown.
591. Vitula pinei, new species
Figures 606, 1090
Forewing white finely dusted with fuscous, giving the
wing a pale gray ground color to the naked eye; trans-
verse lines white bordered inwardly and outwardly by
sharply contrasted blackish lines; antemedial line
slightly angulate; subterminal line bent. inward slightly
for ashort distance from costa, shortly and sharply angled
out at middle and slightly notched at lower fold; discal
dots fused into a black line along discocellular vein and
extending (in fresh specimens) to the black inner border
of subterminal line; terminal dots fused into a black line
along termen; hair tuft enclosed by costal fold, white.
Hind wing whitish, semihyaline with a smoky line along
termen and some smoky shading on the veins. Alar
expanse, 19-21 mm.
Genitalia differing in slight details from those of
edmandsae; vinculum longer, its terminal margin more
evenly rounded; hump of costa of harpe more angu-
late; aedeagus stouter; female genitalia considerably
longer and sclerotization of ductus bursae somewhat
more extended; comparative characters only.
Tyrer nLocauity: Hureka, Utah (type m USNM,
61388).
Foop pLant: Pine cones.
Described from male type and one male paratype
from the type locality, July 14 and 17, 1911, collected
by Tom Spalding; and one female paratype from Baker,
Nev., reared by T. O. Thacher, May 28, 1940, from larva
feeding in cone of Pinus monophylla. The larva was
presumably feeding on the seeds or bracts.
The species is easily distinguished from edmandsae by
the sharply contrasted black double borders of the ante-
medial and subterminal lines. Superficially it bears a
rather striking resemblance to Laetilia zamacrella Dyar.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
592. Vitula inanimella (Dyar), new combination
Fieure 1089
Moodna inanimella Dyar, Proc. U. S. Nat. Mus., vol. 54, p. 372,
Euzophera ticitoa Dyar, Ins. Insc. Menstr., vol. 7, p. 56, 1919
(new synonymy).
A suffused gray-brown species with transverse lines
faintly indicated by their very slightly darker borders;
similar in maculation to edmandsae, but more uniformly
colored. The genitalia are also similar, those of the
males differing only in insignificant details, the differ-
ences no greater than those between individuals of
edmandsae. Female genitalia have the bursae smooth
except for the band of scobinations near signum and the
signa themselves smaller. The type of ticitoa (fig.
1089a) has one signum, the female paratype of inani-
mella from Orizaba, México (fig. 1089) has two signa, a
difference of not specific significance in this genus or the
allied Ephestia groups. The type of ticitoa shows a
vestige of vein 9 on one forewing which probably ac-
counts for Dyar’s placement of it in Huzophera. I am
unable to find a valid character for separating Dyar’s
supposed species and am therefore sinking ficitoa into
the synonymy of inanimella. Alar expanse, 20-21 mm.
Tyre Locauities: Zacualpin, México (inanimella, in
USNM); Volcaén Santa Maria, Guatemala (ticitoa, in
USNM).
Foop puants: Unknown.
Distrisution: México: Orizaba, Zacualpin (May).
Guatema.a: Volcin Santa Maria (May).
Represented in the National Collection only by the
original type series of two males and three females.
593. Vitula laura (Dyar), new combination
Fieure 1091
Euzophera laura Dyar, Ins. Insc. Mensir., vol. 7, p. 56, 1919.
Forewing dark purplish gray, costal area white;
antemedial line indicated by an oblique blackish band
extending across the white area and obsolete below;
subterminal line white, thin, parallel with and rather
close to costa, evenly curved, bordered before and
beyond by narrow bands of the dark ground color; some
faint white dusting bordering termen; discal dots dis-
tinct, separate, black; a row of faint blackish dots along
terminal margin. Hind wing a glossy smoky brown;
veins and terminal margin darker. Alar expanse, 20
mm.
Female genitalia with bursa copulatrix finely, evenly,
and sparsely scobinate; signum a cluster of a half-dozen
small discs; ductus bursae with a sclerotized collar for-
ward of the short sclerotized area from genital opening.
TypE Locatity: Cayuga, Guatemala (July; type in
USNM).
Foop prant: Unknown.
Known only from the unique female type. It is
obviously not a Euzophera, and in habitus and female
genitalia is not too good a Vitula, in which genus it is
placed only provisionally. A male will be necessary
for definite placement. Vein 9 of forewing is absent
287
and the venation otherwise and the female genitalia
indicate that it belongs somewhere in the Moodna-—
Vitula complex.
174. Genus Manhatta Hulst
Hornigia Ragonot, N. Amer. Phycitidae, p. 16, 1887. (Type of
genus: Ephestia biviella Zeller; Europe; figs. 113, 601, 1092.)
Manhatta Hulst, Phycitidae of N. Amer., p. 196, 1890 (mew
name for Hornigia Ragonot 1887, preoccupied by Hornigia
Ragonot 1885 in Galleriidae).
Characters of Vitula except veins 3 and 5 of hind
wing connate.
Hampson (in Ragonot, Monograph, pt. 2, p. 267,
1901) made Manhatta a synonym of Moodna, apparently
on the connate condition of veins 3 and 5 of the hind
wing of biviella. This synonymy cannot stand if we
are to maintain any generic separation between Moodna
and Vitula; for Manhatta has only the hind wing vena-
tion and somewhat longer hind wing cell of Moodna. In
other characters (simple male antenna and complete
transtilla) it agrees with Vitula, thus occupying an inter-
mediate position between the two genera, but appar-
ently more closely related to Vitula than to Moodna.
In addition to its type (biviella), the only European
representative of the Moodna—Viiula complex, it contains
two North American species.
594. Manhatta setonella (McDunnough), new combination
Figures 602, 1093
Moodna setonella McDunnough, Canadian Ent., vol. 59, p. 270,
1927; Check list, No. 6398, 1939.
Forewing whitish gray, more or less dusted with
blackish scales on lower half of wing, giving that area a
somewhat smoky tint; black discal spots and borders of
the transverse lines strongly contrasted, the transverse
lines themselves not distinguishable from the ground
color of wing and indicated only by their black borders;
antemedial line bordered outwardly only, the black
border normally straight and oblique, but in some
specimens slightly angled; subterminal line bulging
slightly at middle, bordered inwardly by a narrow black
line and outwardly by a black dash at costa, continu-
ing as a paler shade to inner margin; discal dots distinct,
separate; a few obscure blackish dots on terminal mar-
gin, beginning below apex and ending above tornus.
Hind wings semihyaline, whitish with veins and terminal
margin pale smoky fuscous. Alar expanse, 14-16 mm.
Male genitalia with costa of harpe produced at apex
into a short free spur. Female genitalia with ductus
bursae sclerotized for half its length from genital opening,
the sclerotized portion constricted at middle; a small
sclerotized patch at junction of ductus and bursa; spin-
ing of bursa as in typical Vitula.
Typr LocaLity: Seton Lake, British Columbia (June;
type in Canadian Nat. Coll.).
Foop puant: Unknown.
In addition to female paratypes from the type locality
there are in the U.S. National Collection two males from
Eureka and Provo, Utah (June). The species is easily
288
recognized byits genitalia; the apically produced costa of
harpe and the patch in the neck of the bursa at once
distinguishing it from anything else in the Moodna—
Vitula group. Superficially the moths resemble those
of Vitula pine but are smaller and lack the dark border
on inner side of the antemedial line.
595. Manhatta broweri, new species
Fieure 1094
Forewing pale brownish fuscous with a white-powdered
area about the discal dots and extending to costa, and
some white dusting at tornus; transverse lines white,
narrow; at base below fold and extending beyond ante-
medial line along fold a faint reddish ocherous shade;
dark ground color concentrated and intensified as rather
diffused broad bands outwardly bordering the anteme-
dial and inwardly bordering the subterminal lines; ante-
medial line slightly angulate; subterminal line irregu-
larly dentate, parallel with termen; discal dots more or
less confluent, brown. Hind wing pale fuscous, veins
and terminal margin slightly darker. Alar expanse, 14—
16 mm.
Genitalia, (male and female) differing very little from
those of Vitula edmandsae and exhibiting no distinguish-
ing specific characters.
TYPE LocALITY: Bar Harbor, Maine (type in USNM,
61389).
Foop prant: Unknown.
Described from male type and one male and five female
paratypes from the type locality, July 25-30, 1933, and
one male paratype from Mount Desert Isl., Maine, July
17, 1934, all collected by Dr. E. A. Brower for whom the
species isnamed. Paratypes deposited in Dr. Brower’s
collection.
The species is easily identified by its color and macu-
lation. Its male genitalia (of the edmandsae type) dis-
tinguish it from the other two species of Manhatia and
its hind wing venation from any species of Vitula. It
is the only known species of Manhatta in the eastern
United States.
175. Verina, new genus
TypE or Genus: Moodna supplicellaiDyar
Tongue well developed. Antenna pubescent (in male
the cilia less that the width of shaft in length); shaft of
male with a few rough scales at base, above, and just
beyond, a slight, very shallow sinus. Labial palpus
oblique, slender, reaching above vertex; third segment as
long assecond. Maxillary palpus filiform,short. Fore-
wing smooth; 10 veins; vein 2 from before, but near,
lower outer angle of cell; 3 from the angle; 4 and 5 long
stalked; 6 from below upper angle of cell, straight; 8
and 9 united; 10 from the cell approximate to 8 at base;
male with a strong costal fold enclosing hair tuft. Hind
wing with vein 2 from well before lower outer angle of
cell; 3 and 5 from the angle, closely approximate at base;
7 and 8 anastomosed for all or most of their lengths be-
yond cell; cell very short (about one-fourth the length
of wing), a trifle longer in female than in male; disco-
UNITED STATES NATIONAL MUSEUM BULLETIN 207
cellular vein slightly curved (nearly straight in male).
Kighth abdominal segment of male with compound dor-
sal tufts.
Male genitalia with apical process of gnathos forked,
U-shaped. Uncus broad, terminal margin broadly
rounded. Transtilla mcomplete. Harpe with a short
blunt digitate projection from near middle of costa.
Anellus a U-shaped sclerotized band. Aedeagus long,
not appreciably tapering; penis bearing a couple of
elongate, thin, weakly sclerotized plates, otherwise un-
armed. Vinculum elongate-angulate, tapering.
Female genitalia with ductus bursae sclerotized for
most of its length from genital opening; sclerotized por-
tion slightly flattened; bursa copulatrix sparsely and
finely scobinate; an extruded lobe from membranous
area between eighth-segment collar and ovipositor (as
in Moodna).
The genus is distinguished from the other genera in
group C by the following’ combination of characters:
Male antenna with slight sinus in shaft; eighth abdom-
inal segment of male with tuft; veins 3 and 5 of hind
wing approximate at base, cell very short and discocel-
lular vein very slightly curved; gnathos terminating in a
forked process; harpe with digitate projection from costa;
transtilla incomplete.
It contains one tropical American species.
596. Verina supplicella (Dyar), new combination
Fieures 116, 603, 1097
Moodna supplicella Dyar, Proc. U. 8S. Nat. Mus., vol. 47, p. 342,
1914.
Forewing dark vinous, dusted with black especially
in area between the transverse lines; basal and tornal
areas slightly paler; transverse lines white, slender,
powdery and broken, the antemedial line far out and
oblique, the subterminal parallel with and close to
termen; discal and terminal dots not distinguishable.
Hind wing translucent; whitish to pale fuscous; the
veins and terminal margin darker. Head and thorax
ocherous, more or less shaded with reddish or blackish
scaling. Labial palpus ocherous with reddish scaling
on terminal segment. Alar expanse, 13-16 mm.
Male genitalia with terminal margin of cucullus of
harpe oblique and slightly concave, its lower angle
produced, vinculum evenly tapering to abruptly pointed
extremity; arms of forked process of gnathos approx-
imate at their apices. Female genitalia with sclerotized
portion of ductus bursae strongly striated at genital
opening, finely granulate otherwise.
Typr Locatiry: Rio Trinidad, Panamé (type in
USNM).
Foop puant: Dried leaves.
Distrisution: Mexico: No specific locality (Aug.,
reared from leaf trash in banana cargo). GUATEMALA:
No specific locality (Apr., reared from dried leaves
intercepted at quarantine), Guatemala City (Mar.).
Panami: Cabima (May), Porto Bello (Apr., May,
Oct.), Rio Trinidad (June). Braziu: Santa Catarina
(July).
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
This species has been intercepted a number of times
at our quarantine ports in banana trash. The larva is
apparently a feeder on dry vegetable refuse. The moth
is easily identified by its male genitalia and (in fresh
specimens) by the broken powdery transverse lines and
wine color of its forewings.
176. Vagobanta, new genus
Type oF GENus: Cryptoblabes divergens Butler.
Characters of Verina except: Antenna of male simple;
eighth abdominal segment of male simple; forewing with
vein 2 shortly separated from, approximate to, or con-
nate with 3 (more separated in females than in males);
hind wing with veins 3 and 5 shortly stalked; cell long
(one-half or a trifle over one-half the length of wing);
discocellular vein decidedly curved.
Contains one tropical American species.
597. Vagobanta divergens (Butler), new combination
Fiaures 604, 1100
Cryptoblabes divergens Butler, Trans. Ent. Soc. London, p. 60,
1883.
Moodna divergens (Butler) Hampson, in Ragonot, Monograph,
pt. 2, p. 269, 1901.
Forewing powder gray, the basal area distinctly paler;
in fresh specimens an obscure shading of reddish scales
on middle of lower fold; antemedial line distinct, de-
cidedly oblique, white, bordered outwardly by a black-
ish band, nearly straight but in some specimens with a
notch at lower fold; subterminal line obscure, sinuate,
very faintly bordered by dark line on inner side. Hind
wing translucent, white, with a pale brown line along
termen and the veins slightly darkened. Alar expanse,
21-25 mm.
Male genitalia with arms of U-shaped apical projec-
tion of gnathos very short and widely spaced. Ele-
ments of divided transtilla long and stout. Harpe with
digitate projection from about middle of costa; outer
margin of cucullus rounded and turned up slightly at
apex. Anellus V-shaped, with stout base. Aedeagus
stout. Vinculum stout, but slightly tapering; terminal
end broadly and abruptly angulate, reinforced by a
sclerotized, pocketlike fold.
Female genitalia with bursa copulatrix large, finely
and sparsely scobinate; signum weak; ductus bursae
sclerotized for half its length, the sclerotized portion
flattened and bent.
TypxE LocaLity: Coquimbo, Chile (type in BM).
Foop piant: Puya alpestris.
Distrisution: Cute: Angol, Coquimbo.
A reared series of eight specimens from Angol, Chile,
is in the National Collection, received from D. S.
Bullock but undated.
177. Moodnella, new genus
Typr or Genus: Moodnella paula, new species.
Tongue well developed. Antenna with slight sinus
in shaft of male near base. Labial palpus oblique,
slender, reaching a trifle above vertex, third segment
800329—56——20
289
shorter than second. Maxillary palpus filiform, rather
long (as long as third segment of labial palpus and about
twice the length of the maxillary palpi of the other
genera in the Moodna-Vitula group). Wing venation
asin Vitula. Forewing with strong costal fold enclosing
scale tuft. Hind wing with cell one-third the length of
wing; discocellular vein curved. Eighth abdominal
segment of male simple.
Male genitalia as in Vitula.
Female genitalia with ductus bursae very weakly
sclerotized for a very short distance from genital open-
Differs from Vitula in having a sinus in the shaft of
male antenna, no tufts on eighth abdominal segment of
the male; from Moodna in its complete transtilla, simple
harpe, simple eighth abdominal segment, approximate
condition of veins 3 and 5 of hind wing; from Verina
in its complete transtilla, small hooked, apical process
of gnathos, simple harpe, simple eighth abdominal seg-
ment, longer cell and more distinctly curved discocellu-
lar vein of hind wing; and from all three of these genera
by its longer maxillary palpi.
Contains one tropical American species.
598. Moodnella paula, new species
Ficures 608, 1096
Forewing gray suffused with rufous and shaded with
blackish, the blackish shade extending in a narrow band
along costa and diffused irregularly in the median area;
antemedial white line far out on wing, at or a trifle
beyond middle, distinct on lower half, fading out
towards costa, bordered outwardly by an irregularly
diffused black, band; subterminal line narrow, parallel
with termen, nearly straight, slightly oblique from costa
to vein 6, shortly out-angled just below, thence straight
to inner margin, white, bordered inwardly by a narrow
blackish band; the area between the transverse lines
greatly restricted. Hind wings white to smoky fuscous;
the veins faintly darkened and a dark line along termen.
Head and thorax reddish ocherous with a scattered
dusting of blackish scales. Alar expanse, 14-15 mm.
Male genitalia with uncus narrowly rounded at apex;
vinculum long, stout, scarcely tapering, terminal margin
broad but very slightly convex (nearly straight).
TypE Ltocauity: Guatemala City, Guatemala (type
in USNM, 61390).
Foop prant: Unknown.
Described from male type and one male paratype
from the type locality collected by C. N. Ainslie, Mar.
1932; one female paratype from Santa Catarina, Brazil,
collected by Fritz Hoffmann, July 9, 1935; one female
paratype from Tigre, Argentina, Aug. 1939, from the
collection of Fernando Bourquin; and one female para-
type from Vicosa, Minas Gerais, Brazil, E. J. Hamble-
ton, collector, ‘‘12—-1-34,” from the Cornell Collection.
A pretty little species easily recognized by its reddish
fuscous color, the narrow interspace between the trans-
verse lines of forewing and the distinct whiteness of
these lines on their lower halves.
290
* Genera 178 and 179: Volatica and Vezina
[Venational division A. ‘Forewing with 11 veins; 8 and 9 long
stalked (9 rather weak). Hind wing with veins 3 and 5 approxi-
mate and from lower angle of cell. Labial palpi porrect or
oblique. Maxillary palpi minute. Transtilla complete and de-
veloped as an angulate bridge, or incomplete. Ductus bursae
with projecting sclerotized shield behind genital opening.]
178. Volatica, new genus
TyPE oF GENUS: Zophodia pachytaeniella Ragonot.
Tongue well developed. Antenna pubescent, simple.
Labial palpus porrect (second segment oblique, third
deflected forward), extending twice the length of head
beyond it, broadly scaled, third segment less than half
the length of second. Maxillary palpus minute, fili-
form. Forewing smooth; 11 veins; vein 2 from before,
but near, lower outer angle of cell; 3 from the angle; 4
and 5 stalked (for at least half their lengths), the stalk
separated from 3 at base; 6 from below upper angle of
cell, straight; 8 and 9 long stalked (free element of 9
sometimes weak, but always present); 10 from the cell,
separated from the stalk of 8-9 at base; male without
costal fold. Hind wing with vein 2 from well before
lower outer angle of cell; 3 and 5 from the angle, approxi-
mate at base; 7 and 8 anastomosed beyond cell for at
least half their lengths; cell about half the length of
wing; discocellular vein curved. Highth abdominal
segment of male simple.
Male genitalia of the Vitula type but larger and more
robust.
Female genitalia with or without signa; bursa copu-
latrix more or less finely scobinate; ductus bursae shorter
Gn pachytaeniella) or very little longer than bursa,
(trinitatis), sclerotized for over half its length, the
sclerotized portion flattened except just before genital
opening and with a more or less rounded, projecting,
sclerotized shield behind genital opening; ductus
seminalis from bursa near signum.
The genitalia, male and female, show the close rela-
tionship of this genus to the Vitula group despite the
forewing venation which is definitely that of venational
division A. Vein 9 while sometimes weak is always
present. In Vitula on the other hand vein 9 is nor-
mally absent, being present and weak only in occasional
specimens. Volatica is distinguished from all the other
genera of the Moodna—Vitula complex by its porrect,
broadly scaled labial palpi.
599. Volatica pachytaeniella (Ragonot), new combination
Figures 610, 1098
Zophodia pachytaeniella Ragonot, Nouv. Gen., p. 31, 1888.—
Hampson, zn Ragonot, Monograph, pt. 2, p. 23, 1901.
Forewing white lightly sprinkled with brownish
fuscous scales giving the costal half of the wing a soiled
ashy white appearance; lower half of wing smeared with
a pale drab shade; on fresh specimens a pink streak
along lower margin at base; antemedial line angulate,
indicated chiefly by a rather broad blackish brown outer
UNITED STATES NATIONAL MUSEUM BULLETIN 207
border; subterminal line more or less irregularly dentate,
bordered inwardly by a narrow blackish fuscous ‘line;
discal spot at lower outer angle of cell distinct, upper’
discal spot occasionally distinct but often obscure or
absent. Hind wing white; a broad fuscous shade along
costa; some darkening of the veins and a distinct dark
line along termen, broadening in most specimens. Alar
expanse, 26-33 mm.
Male genitalia with vinculum slightly constricted
before terminal margin; apical end of harpe somewhat
spatulate. Female genitalia with bursa copulatrix
elongate, much longer than ductus. bursae; signum
present, consisting of a short band of partially fused
discs; ductus bursaesclerotized for its entire length.
TypE LocaAuity: Rio de Janeiro, Brazil (type in Paris
Mus.).
Foop PLANT: Unknown.
Distrisution: Braziu: Parand, Castro, Ypiranga;
Rio de Janeiro, Petrépolis; Santa Catarina (no dates on
any specimens before me).
A large, distinct species so far only known from
Brazil, easily identified by maculation, color, and
genitalic structure.
600. Volatica trinitatis, new species
Fieures 609, 1099
Somewhat narrower winged than pachytaeniella,
similarly colored, but with whitish area of forewing
restricted to a narrower border along costa; transverse
lines obscured, only the antemedial line indicated in
male by a partial dark outer border, very faint in the
males before me, absent in the female; the latter shows
some dark scaling on the veins and has fuscous hind
wings; hind wing of male white with little or no dark
shading along costa or termen. Alar expanse, 24-26
mm.
Male genitalia with apical process of gnathos longer
than in pachyiaeniella (over twice as long as broad);
terminal margin of harpe evenly rounded; vinculum
not appreciably constricted towards terminal margin.
Female genitalia with bursa copulatrix somewhat
shorter than ductus bursae, without signum; ductus
bursae sclerotized for not more than two-thirds of its
length, bent slightly near membranous part of the
ductus. ‘There are also differences in the shape of the
dorsal projecting shield at genital opening between
trimiatis and pachytaeniella, but these differences may
not be significant.
Typr Locatity: Fyzabad, Trinidad (type in Cornell
Univ. Coll.).
Foop prant: Unknown.
Described from male type and one male and one
female paratype from the type locality. (Feb. 17, 19,
21, 1928) from the Cornell Collection, and one male
paratype (USNM 61391) from Trinidad, without date
or more exact locality, collected by A. Busck.
The species is very close to but apparently distinct
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 291
from pachytaeniella, easily distinguished by its genitalia,
the more extended drab suffusion and less distinct
transverse lines on forewing.
179. Vezina, new genus
TypE oF Genus: Vezina parasitaria, new species.
Tongue well developed. Antenna pubescent, simple.
Labial palpus oblique; reaching as high as vertex;
rather broadly and roughly scaled; third segment
shorter than second. Maxillary palpus short, filiform.
Forewing smooth; 11 veins; vein 2 from before but
rather near lower outer angle of cell; 3 from the angle;
4 and 5 stalked for half their lengths, separated from 3
at base; 6 from below upper angle of cell, straight; 8
and 9 long stalked; 10 from the cell, separated from 8
at base; male with a strong costal fold enclosing hair
tuft. Hind wing with vein 2 from well before lower
outer angle of cell; 3 and 5 from the angle, approximate
at base; 7 and 8 anastomosed for most of their lengths
beyond cell; cell about one-third the length of the wing
in male, one-half in female; discocellular vein curved.
Eighth abdominal segment of male with two pairs of
dorsolateral hair tufts.
Male genitalia with apical process of gnathos a short,
bluntly pointed hook. Uncus broad; terminal margin
broadly rounded. ‘Transtilla incomplete; its elements
weakly sclerotized. Harpe with a short angulate pro-
jection from costa near middle and with apex of costa
produced into a free hook at apex (as in Anagasta
kithniella). Anellus a stout, broad, semicircular band
with broad, deep, V-shaped ventral incision. Aedeagus
long, stout; penis armed with a thin, narrow, sclerotized
band and a stout, thornlike, broadly based spine.
Vinculum stout; tapering slightly; terminal margin
moderately broad.
Female genitalia with signum a small cuplike disc or
discs; bursa minutely and sparsely granulate; ductus
bursae granulate towards junction with bursa, with a
strongly sclerotized collar surrounding a broad genital
opening and with a broad, projecting, sclerotized shield
behind genital opening; ductus seminalis from bursa
close to signum.
This genus, like Volatica, is close to and obviously
related to Vitula despite its forewing venation. It is
distinguished by the following combination of charac-
ters: Vein 9 of forewing present; eighth abdominal
segment of male with paired dorsal tufts; transtilla in-
complete; penis armed with a stout, thornlike cornutus;
anellus a broad, semicircular band with V-shaped ven-
tral incision; ductus bursae with broad sclerotization
surrounding broad genital opening.
601. Vezina parasitaria, new species
Figures 598, 1068
Forewing dark gray (grayish brown on worn and
faded specimens); transverse lines white and strongly
contrasted from inner margin to cell, thence to costa
pale but more obscure, narrowly bordered inwardly and
outwardly by black lines; these most obvious from lower
margin of cell to costa; antemedial line at middle of
wing and nearly vertical; subterminal line oblique and
well back from termen; the interspace between the lines
short; discal spots at end of cell, confluent and forming
a thin black line along discocellular vein; terminal dots
obscure. Hind wing shining white, with a fuscous
shade along costa, some fuscous shading at apex and a
narrow dark line along termen. Alar expanse, 17-25
mm.
Male genitalia with vinculum but slightly longer
than broad; terminal margin angulate. Female geni-
talia with a short, thornlike pouch projecting from
venter of ductus bursae near genital opening.
Typr tocauity: José C. Paz, Province of Buenos
Aires, Argentina (type in USNM, 61392; paratypes in
Mus. Argentino Cienc. Nat., Buenos Aires, and BM).
Foop: Larvae feeding in larval cases of Oiketicus
kirbyi Guilding.
Described from male type and one male and one
female paratype from the type locality; one male para-
type from Juan B. Gonnet, Province of Buenos Aires;
five female paratypes from Tigre, Argentina; and one
female paratype from southeast Brazil, F. D. Jones,
1920, this last from the British Museum Collection.
The Argentinian specimens were received from Dr.
Everard E. Blanchard, Director and Sefor José A.
Pastrana, Ingeniero, of the Instituto de Sanidad
Vegetal of the Ministerio de Agricultura in Argentina.
They had been reared by Sefior Pastrana and Dr. Pablo
Kohler from small larvae in the larval cases of the
psychid Otketicus kirbyi Guilding. Dr. Kébler identi-
fied them as a new species to which he gave the manu-
script name ‘‘Canarsia parasitaria” upon the assump-
tion that the species was parasitic on the Ovketicus.
With Dr. Koéhler’s permission I am adopting his manu-
script specific name, but am in doubt as to the parasitic
nature of the larva. According to Sefior Pastrana the
actual feeding habits have not been noted, and I am
inclined to believe (from its close affinities to the
Moodna-Vitula complex) that parasitaria is a scavenger
rather than a parasite or predator upon the living
Otketicus larva or pupa. In a letter of Dec. 29, 1947,
Dr. Blanchard states that, in the Province of José C.
Paz, Sefior N. Jauch has observed larvae of parasitaria
on the remains of bagworms and a few days later all
traces of bagworms had disappeared, suggesting that
the larva are feeders upon the bagworms at least to the
extent of eating the larval and pupal exuviae or the
remains of the dead females of Oiketicus.
The species is a striking one easily recognized by its
bright white transverse lines with narrow black borders
contrasted against the otherwise dark gray ground color
of the forewing. The female genitalia are different in
the armature about genital opening from any other in
the subfamily.
292
Genera 180-185: Caudellia to Plodia
[Venational division E. Forewing with 9 veins; 10 from the
cell; 9 absent; 4 absent; 2 and 3 from the cell. Hind wing with
discocellular vein curved. Male genitalia with transtilla com-
plete or its elements greatly enlarged. Female genitalia with
ductus bursae sclerotized for a considerable part of its length
and usually flattened.]
180. Genus Caudellia Dyar
Caudellia Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 116,
1904.—Forbes, Cornell Mem. 68, p. 636, 1923. (Type of
genus: Caudellia apyrella Dyar.)
Tongue well developed. Antenna pubescent; shaft
of male with a short, shallow sinus near base (as in
Moodna). Labial palpus oblique, reaching to or slightly
above vertex, somewhat flattened laterally; third seg-
ment nearly as long as second (somewhat broadly scaled
and deflected forward in apyrella and albovittella). Max-
illary palpus minute, filiform. Forewing smooth; 9
veins; vein 2 from before but rather near lower outer
angle of cell; 3 from the angle; 4 absent; 5 more or less
approximate to 3 at base; 6 from below upper angle of
cell, straight; 8 and 9 united (9 absent); 10 from the
cell, separated from 8 at base; male with costal fold
enclosing a scale tuft. Hind wing with vein 2 from well
before lower outer angle of cell; 3 and 5 connate, rarely
very shortly stalked; 7 and 8 anastomosed for most or
all of their lengths beyond cell (8 when present very
short) ; cell one-half or slightly less than half the length
of wing; discocellular vein curved. Wighth abdominal
segment of male with compound dorsal tufts.
Male genitalia with transtilla complete (declivella) or
its elements broadened and often fused with arms of
gnathos. Aedeagus straight, smooth; penis with cor-
nutus developed as a weakly sclerotized, narrow, flat-
tened band, otherwise unarmed.
Female genitalia with ductus bursae sclerotized for a
third or more of its length from genital opening; signa
present as a cluster of small, bluntly rounded, projecting
discs or spines more or less fused at their bases; ductus
seminalis from bursa adjacent to signa (approximately
at middle of bursa).
This genus and the following genera with sclerotized
ductus bursae form a compact, strictly American group
closely related to and evidently derived from the
Moodna-Vitula complex. The moths of Caudellia also
resemble Moodna in color and maculation and male
antennal structures. The species differ markedly from
each other in male genitalia. On labial palpi they
divide into two groups:
Palpus somewhat broadly scaled and third segment deflected
forward.
Palpus slender and third segment not deflected.
The deflection of the third segment in our two species
from the Eastern United States is rather slight and does
not give the palpus nearly so pronouncedly porrect an
appearance as, for example, that of Plodia, and I do not
UNITED STATES NATIONAL MUSEUM BULLETIN 207
believe justifies any generic separation of the two
species groups.
Genus Caudellia, Species 602 and 603: C.. apyrella
and C. albovittella
[Labial palpus somewhat broadly scaled and third segment de-
tected forward.]
602. Caudellia apyrella Dyar
Figures 122, 613
Caudellia apyrella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 116, 1904.—Forbes, Cornell Mem. 68, p. 636, 1923.—
McDunnough, Check list, No. 6376, 1939.
Forewing vinous brown overlaid with blackish brown,
the blackish shading most obvious at apex; a faint pale
patch on outer third of costa; transverse lines and discal
spots obscured, the antemedial line very faintly indi-
cated, oblique, pale vinous brown. Hind wing whitish,
faintly tinted with fuscous; veins not appreciably
darkened. Alar expanse, 14-15 mm.
Male genitalia with gnathos terminating in a stubby,
angulate projection; harpe simple.
Tyrer LocaLity: Plummers Island, Md. (type in
USNM).
Foop prant: Unknown.
Known only from the type (co, July) and paratype
(o@, June) from the type locality.
603. Caudellia albovittella Dyar
Ficure 1103
Caudellia albovitiella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p- 116, 1904.—Forbes, Cornell Mem. 68, p. 636, 1923.—
MeDunnough, Check list, No. 6377, 1939.
Forewing vinous brown shaded with blackish brown,
the blackish shading less diffused than in apyrella, con-
centrated as a dark median streak at base, a dark blotch
on outer side of antemedial line and a fainter shade at
apex; antemedial line oblique, strongly contrasted,
white, preceded by a short white dash on inner margin;
subterminal line obscure, distinguishable on fresh speci-
mens as a thin irregular pale line near termen; discal
dots faint, blackish, separate; terminal dots confluent.
Hind wing pale fuscous, veins very slightly darkened.
Alar expanse, 13-21 mm.
Male genitalia not distinguishable from those of
apyrella. Female genitalia with ductus bursae scler-
otized for slightly less than half its length from genital
opening; signa a double line of 6 to 8 short discs, close
together and fusing at their bases.
Typ Ltocatity: Plummers Island, Md.
Foop prant: Unknown.
Distrisution: Maryland, Hyattsville (July), Plum-
mers Isl. (June, July); Missouri, St. Louis (Aug.).
Probably not specifically distinct from apyrella which
may be only a suffused color form or food-plant race; but
as nothing is known of the biology and no females cor-
responding to the males of apyrella are available for
genitalic comparison, the two will have to be kept as
separate species for the time being.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Genus Caudellia, Species 604-607: C. nigrella to
C. clara
[Labial palpus slender and third segment not deflected.]
604. Caudellia nigrella (Hulst), new combination
Ficures 611, 1102
Ephestia nigrella Hulst, Phycitidae of N. Amer., p. 200, 1890.
Mescinia nigrella (Hulst) Ragonot, Monograph, pt. 2, p. 85, 1901.
Moodna nigrella (Hulst) Barnes and McDunnough, Check list of
the Lepidoptera of Boreal America, No. 5796, 1917.—
McDunnough, Check list, No. 6397, 1939.
Ephestia arizonella Walter, Proc, Ent. Soc. Washington, vol. 30,
p. 141, 1928.—McDunnough, Check list, No. 6405, 1939.
(New synonymy.)
In color and markings resembles Moodna ostrinella.
Forewing blackish fuscous; basal area more or less
shaded with red or reddish ocherous, especially on lower
half, this reddish shade sometimes extending outward
for a short distance along lower fold, reappearing in
tornal area; antemedial line distinct, rather wide, nearly
straight, and in many specimens apparently vertical but
actually a trifle oblique; subterminal line obscure, paral-
lel to termen, slightly and irregularly denticulate; discal
dots sometimes obscured by the blackish dusting of
median area but usually distinct, more or less confluent
and set off by some surrounding pale dusting which
extends in well marked specimens to costa. Hind wing
white, smoky white or pale fuscous; veins darkly out-
lined, some dark shading and a narrow dark line along
termen. Alar expanse, 14-20 mm.
Male genitalia with gnathos terminating in a short,
bluntly pointed, straight spike; elements of transtilla
thin, broad, lightly sclerotized, fusing with arms of
gnathos and extending beyond them to subanal plate
(fig. 6112); harpe with a very short digitate projection
from basal third of costa, apex broadly rounded; vin-
culum long, its terminal margin sharply angled; aedea-
gus long and stout. Female genitalia with ductus
bursae sclerotized for about half its length, the sclero-
tized portion longitudinally wrinkled for its entire
length; signum a longitudinal series of four or five small
discs.
Tyrr Locaities: Blanco County, Tex. (nigrella, in
USNM); Tempe, Ariz. (arizonella, in USNM).
Foop prant: Unknown.
Distrrsution: Teras, Blanco County (Aug.), Browns-
ville (Mar.), San Benito (Mar., June, July, Sept.);
Arizona, Catalina Springs (May), Mohave County
(Sept.), Redington, Tempe (Aug., Sept.), Yuma (June) ;
California, Death Valley (Apr.), La Puerta Valley
(July), Palm Springs (Mar.).
Like Moodna ostrinella a variable species in color.
The name arizonella refers only to a color form with
pale reddish ocherous shading on the basal area of fore-
wing. Its genitalia (male and female) agree in every
detail with those of typical nigrella. The latter name
has been “kicked around” rather carelessly by later
authors since its original placement by Hulst. He
alone seems to have examined the venation, which would
allow reference to Ephestia but not to Moodna and
certainly not to Mescinia.
293
605. Caudellia declivella (Zeller), new combination
Ficures 612, 1104
Ephestia declivella Zeller, Horae Soc. Ent. Rossicae, vol. 16, p.
244, 1881.—Hampson, in Ragonot, Monograph, pt. 2, p.
305, 1901.—Dyar, Proc. Ent. Soc. Washington, vol. 31, p.
17, 1929.—Richards and Thomson, Trans. Ent. Soc. London,
vol. 80, p. 175, 1932.
Ephestia animosella Dyar, Proc. U. 8S. Nat. Mus., vol. 47, p. 345,
1914.
Forewing red heavily dusted with black especially in
the median area, the red shade more obvious in basal
and terminal areas and along the fold; antemedial line
strongly contrasted, white, narrow, straight, oblique;
from antemedial line to base a subcostal streak of white
scaling (only distinguishable in fresh specimens); sub-
terminal line faint and very narrow, near to and parallel
with termen; discal dots obscure, some white dusting in
the area surrounding them. Hind wing pale trans-
lucent fuscous, darker on the veins and towards terminal
margin. Alar expanse, 10-13 mm.
Male genitalia with transtilla complete, developed as
a bridge with humped central projection; apical projec-
tion of gnathos a very short, blunt, knoblike hook;
harpe simple; aedeagus rather slender.
Female genitalia with ductus bursae sclerotized for
most of its length, some sclerotized ridges at the junc-
tion with bursa copulatrix; signa a cluster of minute,
blunt discs.
Typr Locauitigs: Honda, Colombia (declivella, in
BM); Porto Bello, Panama (animosella, in USNM).
Foop puant: Unknown. Larva probably a scavenger
on dried vegetable matter.
Distrinution: PanamA: La Chorrera (Apr.), Porto
Bello, (May), Rio Trinidad (Mar., May, June).
CotomeraA: Honda, Maraquita.
The species is easily recognized by its male genitalia
which are similar to those of Manhatia biviella of
Europe, but unlike anything else from the Americas.
606. Caudellia colorella (Dyar), new combination
Ficures 614, 1101
Ephestia colorella Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 345,
1914.—Richards and Thomson, Trans. Ent. Soc. London,
vol. 80, p. 192, 1932.
Superficially similar in every respect to declivella
except that reddish areas of forewing are paler, ocherous
red rather than dull red as in declivella. In the hind
wing veins 3 and 5 are sometimes very shortly stalked.
They are occasionally so in declivella, but in the latter
normally are connate (not separate as Dyar states in
his original description of animosella). Alar expanse,
11-14 mm.
Male genitalia with transtilla completely fused with
gnathos and greatly broadened posteriorly. Harpe
with short, blunt projections from costa at middle and
apex; uncus broadly triangulate. Female genitalia
with ductus bursae sclerotized for two-thirds its length
from genital opening, not sclerotized at junction with
bursa,
‘294
Typr LocaLity: Taboga Island, Panamé (type in
USNM).
Foop pirant: Unknown.
DistRiBuTIoN: Panamd: Cabima (May), Corozal
(Mar.), Porto Bello (May), Taboga Isl. (Feb.).
607. Caudellia clara, new species
Figure 615
Pattern and color of forewing much as in the two pre-
ceding species except the red shading more confined to
‘streaks along the folds and rather inconspicuous; white
antemedial line and the white extension from it along
costa to base more strongly contrasted; general ground
color purplish fuscous; discal dots rather well marked
‘and set in a clear white field. Hind wing whitish; the
veins and a narrow line along termen pale fuscous.
Alar expanse, 13.5 mm.
Male genitalia with elements of transtilla very broad
and completely fused with arms of transtilla; gnathos
terminating in an elongate, stout, blunt hook; uncus
broad throughout, its terminal margin straight; harpe
with an enlarged flattened scooplike projection from
base of costa; costa broadly sclerotized and slightly and
bluntly projecting at apex. Female unknown.
TypE Locality: Hl Yunque, Luquillo Mts., Puerto
Rico (type in Cornell Univ. Coll.; paratype in USNM,
61393).
Foop eLant: Unknown.
Described from male type and one male paratype
from the type locality, 1,500 to 2,000 ft., Cornell lot No.
795, sub. 38, Apr. 22, 1930.
_ A distinct species, but in color and maculation hardly
separable from declivella. However, the male genitalia
are distinctive and easily identify the species.
181. Genus Microphestia Dyar
Microphestia Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 346, 1914.
(Type of genus: Microphestia animalcula Dyar.)
Tongue short (but not completely enclosed by labial
palpi). Antenna of female roughly scaled. Labial
palpus oblique, slender, reaching nearly to vertex; third
segment slightly shorter than second. Mazxillar palpus
minute, filiform. Forewing smooth; 9 veins; vein 2
from very near lower, outer angle of cell; 2, 3, and 5
slightly separated and approximately equidistant at
base; 4 absent; 6 from below upper angle of cell,
straight; 9 absent; 10 from cell, separated from 8 at
base. Hind wing with 2 from well before lower angle
of cell; 3 and 5 shortly stalked (not long stalked as
stated by Dyar); 7 and 8 completely anastomosed
beyond cell; cell one-half the length of wing; discocellu-
lar vein curved.
Female genitalia with ductus bursae sclerotized for
most of its length from genital opening, the sclerotized
area flattened; a triangulate, projecting, sclerotized
shield behind genital opening.
The above diagnosis is incomplete, as the male is
unknown. The genus is doubtfully distinct from
“UNITED STATES NATIONAL MUSEUM BULLETIN 207
Caudellia. It is distinguished from the latter chiefly
by its reduced tongue and the small size of its type, 8
mm.
608. Microphestia animalcula Dyar
Figure 1114
Microphestia animaleula Dyar, Proc. U. S. Nat. Mus., vol. 47,
p. 346, 1914.
Forewing unicolorous, dark glossy brown; under
magnification the faintest indication of a thin, pale,
oblique antemedial line; no other markings. Hind wing
pale fuscous. Alar expanse, 8 mm.
Female genitalia having bursa without signum;
ductus bursae no longer than bursa; ductus seminalis
from approximately middle of bursa.
Typr LocaLity: Rio Trinidad, Panam4é (Mar.; type
in USNM).
Foop prant: Unknown.
Known only from the unique female type.
182. Sosipatra, new genus
TyprE oF GENus: Ephestia rileyella Ragonot
Characters of Caudellia except: Shaft of male antenna
simple; labial palpus oblique or erect; apical process of
gnathos enlarged (broadened), undivided, knobbed or
looped; harpe with apex of costa produced as short
spine at apex, or apex of costa and cucullus upturned;
transtilla complete, an angulate bridge entirely free of
gnathos; ductus seminalis from bursa copulatrix near
its junction with ductus bursae; eighth abdominal seg-
ment of male with compound dorsal tufts or simple. In
the hind wing, veins 3 and 5 are either connate or
shortly stalked. Signum, when present, a single, blunt,
thornlike disc, adjacent to ductus seminalis.
The new genus brings together a group of American
species having a consistent female character in the
position of ductus seminalis in relation to the bursa and
a male character in the broadened apical process of
gnathos. On the harpe and the eighth abdominal seg-
ment of the male it divides into two groups as follows:
Harpe with costa produced at apex into a short spine; eighth
abdominal segment of male with compound tufts.
Harpe with apex of costa and cucullus upturned; eighth ab-
dominal segment of male simple.
Genus Sosipatra, Species 609-612: S. rileyella to
S. thurberiae
[Harpe with costa produced at apex into a short spine; eighth
abdominal segment of male with compound tufts.]
609. Sosipatra rileyella (Ragonot), new combination
Figures 616, 1105
Ephestia rileyella Ragonot, N. Amer. Phycitidae, p. 17, 1887.—
Hulst, Phycitidae of N. Amer., p. 198, 1890.—Hampson, in
Ragonot, Monograph, pt. 2, p. 294, 1901.—Richards and
Thomson, Trans. Ent. Soc. London, vol. 80, p. 182, 1932.—
McDunnough, Check list, No. 6400, 1939.
Forewing cream white very sparsely sprinkled with
black scales; costal edge for half the distance from base
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
to antemedial line black; antemedial line indicated by
two black spots, one above the other, on median and
lower folds, rarely with additional black dots above and
below them; subterminal line indicated by an oblique
series of black spots on the veins; discal spot on lower
outer angle of cell conspicuous, black, the discal spot
on upper angle of cell more or less obsolescent. Hind
wings translucent, shining white; a fine brown line
along termen; veins faintly, if at all, darkened. Alar
expanse, 15-23 mm.
Male genitalia distinguished chiefly by the shapes of
transtilla (fig. 616b) and apical process of gnathos
(fig. 616a). Female genitalia with signum.
Type Locauity: Utah (type in Paris Mus.).
Foop piants: Yucca, Nolina.
Distrisution: Unirep States: Utah, Hureka (June),
Penah (Feb.); Colorado, Grand Junction (July); Cali-
fornia, Loma Linda (Mar.), Los Angeles County (June,
July, Sept.), Mohave (May), Morongo (Apr.), Phelan
(May), Pipes Canyon (San Bernardino Mts.; Mar.,
Apr., May), San Pasqual (Apr.); Arizona, Catalina
Springs (May), Chiricahua Mts. (Apr., May), Mohave
County (Sept., Oct.), Phoenix (Apr.), Pinal Mts. (May);
New Mexico, Albuquerque (July), Valencia County;
Texas, Brownsville. M*xtco: Sonora.
A species easily recognized by its wing color and
maculation and its close association with Yucca. The
larva feeds upon the dry seeds in the pods of several
species of that genus. There are also in the National
Collection three specimens reared in 1939 by W. D.
Pierce from larvae feeding in the seed pods of Nolina
parryi at Pipes Canyon, San Bernardino Mts., Calif.
N. parryi is a plant closely related to Yucca and by some
botanists classified under the latter genus.
610. Sosipatra micaceella (Hampson), new combination
Figure 617
Ephestia micaceella Hampson, in Ragonot, Monograph, pt. 2,
p. 298, 1901.
Forewing gray-brown powdered with black, the black
dusting heaviest on the costal half of wing, under mag-
nification some white scaling that gives the costal mar-
gin the bluish gray tint mentioned by Hampson; two
black spots on costa near base; antemedian line faint
but distinguishable, pale gray, well out towards middle
of wing and nearly vertical, bordered outwardly by
some black dashes, the latter most pronounced on upper
margin and middle of cell and on lower fold, the two
upper dashes more or less confluent and extended into
the cell; subterminal line oblique, parallel to termen,
inwardly angled at vein 6 and slightly so at lower fold,
bordered inwardly by a black line (broken more or less
into short dashes on the veins); discal spots confluent,
forming a black line along discocellular vein; an obscure
blackish line along termen. Hind wing translucent,
white; the veins outlined by pale fuscous, a fuscous
295
shade along costa and narrowly bordering the termen.
Alar expanse, 18-19 mm.
Male genitalia similar to those of rileyella except for
slight differences in the shapes of transtilla and apical
process of gnathos.
The female is unknown.
Type Locauity: México (type in the collection of the
Abbé Joannis).
Foop prant: Unknown. The type was reared from
a larva found in a cocoon of (Attacus) Rothschildia ori-
zaba (Westwood) but the larval habits were not noted.
Known only from México. There are two males in
the National Collection from the city of México (Nov.).
One of these is badly rubbed; but the other is in fair con-
dition, only the lower half of the forewing being some-
what rubbed, and it answers well to Hampson’s descrip-
tion. Richards and Thomson in their paper on Hphestia
(Trans. Ent. Soc. London, vol. 80, p. 183, 1932) give a
description of micaceella from a male so determined by
Hampson in the British Museum. They publish no
figures, but describe the genitalia; and their description
raises some doubt as to what they had, for they state
that the ‘dorsal thickenings” of anellus (our transtilla)
are “absent, replaced by broad ventral thickenings of
the tegumen itself.” If they had an example of mica-
ceella before them, this is obviously a misinterpretation
of structure from a poor preparation; for micaceella has
as distinct a transtilla as rileyella and no broad ventral
thickening of the tegumen. No phycitid in any nearly
related group has such a tegumen.
611. Sosipatra anthophila (Dyar), new combination
Fiaures 618, 1106
Eurythmia anthophila Dyar, Ins. Insc. Menstr., vol. 13, p. 226,
1925.—McDunnough, Check list, No. 6394, 1939.
Forewing bluish gray, rather broadly shaded with
white along costa and with some faint white dusting in
terminal area; antemedial line slightly angulate, white
outwardly bordered by blackish shading, especially be-
low the white costal suffusion; subterminal line thin,
white, practically straight, bordered inwardly by a nar-
row blackish shade and outwardly for a short distance
from costa by a similar blackish shade; discal dots dis-
tinct, separate; terminal dots obscure. Hind wing
white, translucent; a narrow, pale yellowish fuscous line
along termen. Alar expanse 15-16 mm.
Male genitalia with apical process of gnathos a short,
stout, blunt, oval, scooplike hook; transtilla arched,
flattened at middle; vinculum with terminal margin
angulate. Female genitalia with the bursa somewhat
more strongly scobinate than in other species of the
genus, especially near and at junction of bursa and duc-
tus bursae; signum present.
Type Locatity: Uvalde, Tex. (type in USNM).
Foop piant: Opuntia (larva feeding in the flowers).
Known only from the type series, a male from Uvalde
(May) and one male and one female from Devils River,
Tex. (May).
296
612. Sosipatra thurberiae (Dyar), new combination
Ficures 619, 1107
Eurythmia thurberiae Dyar, Ins. Inse. Mensir., vol. 5, p. 46,
1917.—McDunnough, Check list No. 6393, 1939.
Forewing bluish gray more or less dusted with black,
in some specimens the entire area between the trans-
verse lines suffused with black, a fine whitish dusting
usually sets off the entire basal area; antemedial line
well out towards middle of wing, narrow and nearly
vertical, narrowly bordered outwardly by some accentu-
ation of the black dusting; subterminal line parallel to
termen, slightly angled inwardly at vein 6 and lower
fold; discal dots more or less distinct (at least the lower
one) and from them a broadening wedge of pale scaling,
extending to costa; terminal dots obscure, more or less
confluent. Hind wing white to pale smoky fuscous;
the veins more or less darkened and a narrow dark line
along termen. Alar expanse, 11-19 mm.
Male genitalia similar to those of anthophila but
easily distinguished by the shapes of transtilla and
apical process of gnathos; terminal margin of vinculum
evenly rounded. Female genitalia with very slender
signum.
TYPE LOCALITY: Bowie, Ariz. (type in USNM).
Foop puants: Cercis occidentalis (larvae in pods),
Quercus (larvae in “oak-apple” galls on leaves), Thur-
beria (larvae in seed pods).
Distrisution: Arizona, Bowie (June); California,
Applegate (July, Aug.), Buelton (July), Calpella (July),
Gasquet (July), Hopeland (July), San Felipe Wash
(San Diego County, June), Three Rivers (Apr.);
Oregon, Dundee (Aug.), Eugene (July), Woodburn
(Aug.).
Genus Soszpatra, Species 613-615: S. nonparilella
to S. diwergens
[Harpe with apex of costa and cucullus upturned; eighth ab-
dominal segment of male simple.]
613. Sosipatra nonparilella (Dyar), new combination
Figure 621
Ephestia nonparilella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 118, 1904.—McDunnough, Check list No. 6404, 1939.
Forewing white dusted with black, giving the wing a
pale ashy gray ground color; discal spots and borders of
transverse lines black and strongly contrasted; ante-
medial line indicated chiefly by its outer border, far out
on wing (about middle), just below costa angled inward
and running parallel with subterminal line; the latter
white and straight except for a very slight notch at
lower fold, parallel with and rather close to termen;
discal dots confluent, forming a black streak along
discocellular vein. Hind wing white, translucent; its
terminal margin but slightly darkened. Alar expanse,
18 mm.
Male genitalia distinguished by shape of apical
process of gnathos (fig. 621a).
TYPE LOCALITY: Santa Rita Mts., Ariz. (June; type
in USNM).
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Foop prant: Unknown.
Known only from the male type. Dyar’s original
description is in error in one particular. He states that
veins 3 and 5 of hind wing are “separate, but approxi-
mate at base.’’ They are distinctly connate.
614. Sosipatra majorella (Dyar), new combination
Fieures 620, 1108
Ephestia majorella Dyar, Ins. Insc. Menstr., vol. 10, p. 173, 1922.
Forewing similar to that of micaceella but with less
contrast between the costal and inner areas, and the
black borders of the transverse lines broader, more
complete (not broken into dashes); ground color dark
gray-brown with some lighter rust-brown scaling in the
folds; antemedial line slightly angled just below costa;
the black inner border of subterminal line angled out-
wardly at middle; discal spots confluent, black. Hind
wing of male light brown, of female, whitish fuscous;
the veins and terminal margin darker. Alar expanse,
20 mm.
Male genitalia similar to those of nonparilella except
for a somewhat broader transtilla, more slender aedea-
gus and differently shaped apical projection of gnathos.
Female genitalia with signum; bursa otherwise smooth.
Type nocanity: Guadalajara, México (type in
USNM).
Foop puant: Unknown.
Tn addition to the male type there is in the National
Museum from Mexico City (Aug.) a female which
Schaus or Dyar had associated with micaceella. Its
maculation is a much better match for that of majorella,
with which I associate it. The two species differ radi-
cally in male genitalia butaresimilarin colorand pattern
and easily confused. On the other hand, while they
differ markedly in color, the types of majorella and non-
parilella exhibit only minor structural differences.
615. Sosipatra divergens (Dyar)
Figure 1109
Ephestia divergens Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 345,
1914.
Forewing dark gray-brown; antemedial line vertical,
at middle of wing, bordered outwardly by black; sub-
terminal line oblique, slightly curved at fold, margined
within by a thin black line; discal spots confluent, form-
ing a blackish line along discocellular vem. Hind wing
pale fuscous, veins and terminal margin very slightly
darker. Alar expanse, 16 mm.
Female genitalia without signum. Male unknown.
Type tocatity: Taboga Isl., Panama (type in
USNM).
Foop prant: Unknown.
Known only from the unique female type.
183. Genus Bethulia Ragonot
Bethulia Ragonot, Nouv. Gen., p. 36, 1888.—Hampson, in Rag-
onot, Monograph, pt. 2, pp. xiv, 804, 1901. (Type.of genus:
Bethulia championella Ragonot.)
The generic descriptions of Ragonot and Hampson are
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 297
faulty in several details. The type of championella (and
only known example of the genus) is a female and not a
male as stated by Ragonot and Hampson. The vena-
tion is similar to that of Plodia—veins3 and 5 areslightly
separated at base, not from a point, and 8 and 5 of hind
wings are connate and not shortly stalked. The female
genitalia are also similar to those of Plodia except that
there is no signum. The labial palpi are oblique.
The genus is obviously very close to Plodia and Ribua
but in the absence of a male it is impossible to charac-
terize it properly or determine its status accurately.
616. Bethulia championella Ragonot
Ficures 123, 1120
Bethulia championella Ragonot, Nouv. Gen., p. 37, 1888; Mono-
graph, pt. 2, p. 304, 1901.—Druce, Biologia Centrali-Amer-
icana, Lepidoptera, Heterocera, vol. 2, p. 287, 1896.
The Ragonot figure of the type (Monograph, pl. 35,
fig. 16) is somewhat misleading. An enlarged photo-
graph, before me, shows the blackish margins defining
the antemedial and subterminal lines stronger and some-
what broader than those in the published figure and a
distinct scattering of blackish scales over the white areas
of the forewing; the antemedial line is angulate (not
“oblique’’) and the black border of the subterminal is
complete (not broken as in the figure) and shows a
sharp angulation at vein 6.
In female genitalia the projecting shield behind geni-
talia is subtriangulate and broadly flaring to its wide,
slightly convex terminal margin.
If other specimens are ever received from Central
America they should be easily identified; for champion-
ella is the only described white species, except Socipatra
rileyella, in any of the genera, of similar venation and
with the ductus bursae sclerotized and flattened for part
ofitslength. From rileyella it is easily distinguished by
its genitalia.
Type tocauity: San Joaquin, Vera Paz, Guatemala
(type in BM).
Foop pLant: Unknown.
Known only from the type specimen.
184. Genus Ribua Heinrich
Ribua Heinrich, Proc. Ent. Soc. Washington, vol. 42, p. 31, 1940.
(Type of genus; Ribua innozxia Heinrich.)
Tongue well developed. Antenna simple and pubes-
cent. Labial palpus oblique in male, porrect in female
(the third segment downcurved). Maxillary palpus
minute, filiform. Forewing smooth; 9 veins; vein 2
from just before lower outer angle of cell; 3 from the
angle; 4 absent; 5 closely approximate to 3 at base; 6
from below upper angle of cell, straight; 8 and 9 united;
10 from the cell, closely approximate to 8 for some dis-
tance from cell; male with costal fold enclosing a scale
tuft. Hind wing with vein 2 from well before outer
angle of cell; 3 and 5 approximate at base; 7 and 8 com-
pletely anastomosed beyond cell (rarely, in individual
specimens, with 8 represented as a short spur at costa) ;
cell slightly less than one-half the length of wing; disco-
cellular vein curved. Abdomen of male with a pair of
dorsolateral hair tufts on eighth segment.
Male genitalia with apical process of gnathos asym-
metrical, bearing two short hooks. Transtilla a narrow
band with flattened central process fusing to apical proc-
ess of gnathos (fig. 622). Harpe with costa slightly pro-
duced at apex, otherwise simple. Vinculum long and
broad.
Female genitalia with ductus bursae flattened and
sclerotized for most of its length; bursa copulatrix with
or without signa, latter when present consisting of one
or two small, bluntly pointed, disclike spines, ductus
seminalis from near anterior end of bursa.
The genus is close to but distinct from Plodia, dis-
tinguished by the following combination of characters:
Veins 3 and 5, and 8 and 10 closely approximate at base;
labial palpus of male oblique; tufts on eighth abdominal
segment of male simple; apical process of gnathos asym-
metrical and bifurcate; anellus fusing with gnathos;
ductus seminalis from near anterior end of bursa.
617. Ribua innoxia Heinrich
Figures 622, 1115
Ribua innozia Heinrich, Proc. Ent. Soc. Washington, vol. 42,
p. 32, 1940,
Forewing ash gray, obscurely marked and shaded with
blackish fuscous; an ill-defined dark basal patch; ante-
medial line faintly indicated, oblique, shaded outwardly
by a blackish band; subterminal line obscure, pale,
straight and parallel with termen, narrowly shaded
inwardly and outwardly by dark scaling; the veins
faintly dark-shaded; a scattering of dull reddish scales
on the wing, especially along lower fold, on upper vein
of cell and bordering subterminal line; all marking
obscure and dark shading more or less diffused. Hind
wing dull white; a narrow fuscous shade along costa and
termen, most conspicuous at apex; veins slightly dark-
ened. Alar expanse, 12-16 mm.
Male genitalia with prongs of apical process of
enathos separated (space between them U-shaped);
vinculum nearly twice as long as tegumen. Female
genitalia with two signa.
TYPE Locatity: Cuba (type in USNM).
Foop piant: Fungus on pineapple.
The larvae of this species are frequently intercepted
at our southern ports on pineapples from Cuba, but
apparently do no damage to the fruit. A moth was
recently received from Cuba which had been reared
from a larva in decaying sugarcane. Apparently the
species is more of a scavenger than anything else.
618. Ribua contigua, new species
Figures 623, 1117
Forewing reddish brown; black dusting sparser than
on innoria and largely replaced by red scaling; trans-
verse lines obliterated. Alar expanse, 12-14 mm.
Male genitalia with prongs of apical process of
gnathos close together, the space between them narrowly
V-shaped and the prongs much shorter than those of
298
innoxia; vinculum shorter (notover oneand one-half times
the length of tegumen) and broader throughout. than
that.of innoxia. Female genitalia not exhibiting any
specific characters to distinguish them from ‘those of
innoxia.
TypE LocaLity: Dorado, Puerto Rico (type in Cor-
nell Univ. Coll.; paratype, o, in USNM, 61894).
Foop prant: Unknown.
Described from male type and female paratype from
the type locality, collected May 30, 1930, by W. A.
Hoffman, and two male paratypes from Catafio, Puerto
Rico, collected by Leonard and Mills, May 16 and July
24, 1930.
Nothing is known of its life history, but its larval
habits are probably similar to those of innomia.
619. Ribua patriciella (Dyar), new combination
Fieure 1116
Ephestia patriciella Dyar, Ins. Insc. Menstr., vol. 6, p. 140, 1918.
Forewing coloration similar to that of contigua, but
the reddish dusting more towards a brown shade and
less generally distributed; the veins outlined in blackish
gray which predominates over the red-brown on most of
the wing; transverse lines and discal spots obsolete.
Alar expanse, 12 mm.
Female genitalia without signum.
TypE wLocauity: Baracoa, Cuba (Oct.;
USNM).
Foop piant: Unknown.
Known only from two specimens in poor condition,
the type and one female paratype from Santiago, Cuba
(May). The labial palpi seem more oblique than por-
rect, but they are twisted so much that their shape can
not be accurately determined. ‘The male is unknown.
The three males that Dyar included among his para-
types are Hphestia cautella.
type in
185. Genus Plodia Guénée
Plodia Guénée, Europaeorum Microlepidopterorum index me-
thodicus . . . p. 80, 1845.—Hinemann, Die Schmetterlinge
Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 202,
1865.—Snellen, De Vlinders van Nederland, Microlepidop-
tera, vol. 1, p. 163, 1882.—Hulst, Phycitidae of N. Amer., p.
200, 1890.—Ragonot, Monograph, pt. 2, p. xiv, 1901.—
Hampson, in Ragonot, Monograph, pt. 2, p. 305, 1901.—
Spuler, Die Schmetterlinge Europas, vol. 2, p. 201, 1910.—
Forbes, Cornell Mem. 68, p. 636, 1923.—Meyrick, Revised
handbook of British Lepidoptera, p. 386, 1938.—Richards
and Thomson, Trans. Ent. Soc. London, vol. 80, p. 203,
1932.—Pierce and Metcalf, Genitalia of the British Pyrales
p. 6, 1938.—Janse, Journ. Ent. Soc. South Africa, vol. 8, p.
26, 1945. (Type of genus: Tinea interpunctella Hubner.)
Tongue well developed. Antenna simple and pu-
bescent in both sexes. Labial palpus porrect. Maxil-
lary palpus minute, filiform. Forewing smooth; 9
veins; vein 2 from before, but near, lower outer angle
of cell; 3 from the angle; 4 absent; 5 appreciably sepa-
rated from 3 at base; 6 from below upper angle of cell,
straight; 8 and 9 united; 10 from the cell, separated from
8 at base; male with costal fold enclosing a scale tuft.
Hind wing with 2 from well before outer angle of cell;
UNITED STATES NATIONAL MUSEUM BULLETIN 207
3 and 5 connate; 7 and 8 anastomosed from most or all
of their lengths beyond cell; cell about half the length
of wing; discocellular vein curved. Abdomen of male
with two pairs of dorsal hair tufts, or compound dorsal
tufts on eighth segment.
Male genitalia with apical projection of gnathos a
small knob terminating in a very short, simple, blunt
hook. ‘Transtilla a narrow, curved, sclerotized band,
not in any way fusing to gnathos. Uncus for most of
its length rather narrow, its sides straight or but slightly
tapering to bluntly apical margin. Harpe with a very
slight, rounded projection from costa at middle and
a short thornlike projection at apex, otherwise simple.
Female genitalia with ductus bursae flattened and
sclerotized for over half its length, shorter than bursa;
signa, present, developed as a cluster of 3 to 5 contig-
uous, small, blunt, thornlike projections; ductus semi-
nalis from about middle of bursa, near signa.
The genus is distinguished from Ribua by the follow-
ing combination of characters: Labial palpi of both
Sexes porrect; veins 3 and 5 of forewing distinctly
separated at base; 10 from cell and not close to 8 at
base; 3 and 5 of hind wing connate; eighth abdominal
segment of male with two or more pairs of dorsal tufts;
ductus seminalis from middle of bursa copulatrix. In
the Ragonot key to genera (Monograph, p. xiv) veins 3
and 5 of hind wing are said to be separate; but this is an
error. ‘They are definitely connate.
Plodia is a genus of definitely American origin. Its
type species has become cosmopolitan through trans-
portation in ship stores and the commercial distribution
of dried grains and fruits; but all its nearest relatives
(Ribua, Caudellia, Bethulia, etc.), and the only other
known congeneric species (dolorosa), are confined to the
New World.
620. Plodia interpunctella (Hiibner)
Fiegures 124, 624, 1118
Tinea interpunctella Hiibner, Sammlung europdischer Schmetter-
linge, Lepidoptera 8, Tineae 5, pl. 45, fig. 310 [1810]-[1813].
Elucita interpunctalis Hiibner, Verzeichnis bekannter Schmett-
[erJlinge, p. 347, 1825 (change in spelling for interpunctella).
Phycita interpunctella (Hiibner) Treitschke, Die Schmetterlinge
von Europa, vol. 10, p. 196, 18382.—Duponchel, Histoire
naturelle des lépidoptéres .. . de France, vol. 10, p. 224,
1836.
Myelois interpunctella (Hiibner) Zeller, Isis von Oken, 1839, p.
176.
Plodia interpunctella (Hiibner) Guénée, Europaeorum Microlepi-
dopterorum index methodicus ... , p. 80, 1845.—Heine-
mann, Die Schmetterlinge Deutschlands und der Schweiz,
Abt. 2, vol. 1, pt. 2, p. 202, 1865.—Snellen, De Vlinders van
Nederland, Microlepidoptera, vol. 1, p. 163, 1882.—Ragonot,
Ent. Monthly Mag., ser. 2, vol. 22, p. 25, 1885.—Hulst,
Phycitidae of N. Amer., p. 200, 1900; U. S. Nat. Mus. Bull.
52, p. 487, 1903.— Hampson, in Ragonot, Monograph, pt.
2, p. 305, 1901.—Spuler, Die Schmetterlinge Europas, vol.
2, p. 201, 1910.—Forbes, Cornell Mem. 68, p. 636, 1923.—
Curran, Sci. Agr., vol. 6, p. 386, 1926.—Hudson, Butterflies
and moths of New Zealand, p. 156, 1928.—Meyrick, Revised
handbook of British Lepidoptera, p. 386, 1928.—Noyes,
Bull. Ent. Res., vol. 21, p. 81, 1930.—Richards and Herford,
Ann, Appl. Biol., vol. 17, p. 380, 1930.—Hamlin, Reed, and
Phillips, U. S. Dep. Agr. Techn. Bull. 242, 26 pp., 1931.—
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Richards and Thomson, Trans. Ent. Soc. London, vol. 80,
p. 205, 1932.—Norris, Proc. Zool. Soc. London, pp. 597-611,
1932.—Dickins, Trans. Ent. Soc. London, vol. 85, p. 338,
1936.—Barth, Zool. Jarb., Jena, vol. 58, pp. 297-329, 1937;
Zeitschr. Wiss. Zool., Leipzig, vol. 150, p. 27, 1938.—Leh-
mensick and Liebers, Zeits. Angew. Ent., Berlin, vol. 24,
p. 441, 1937; p. 582, 1938.—McDunnough, Check list, No.
6408, 1939.—Hinton, Bull. Ent. Res., vol. 34, p. 185, 1943.—
Corbet and Tams, Proc. Zool. Soc. London, p. 64, 1943.
Ephestia interpunclella (Hiibner) Herrich-Schiffer, Systematische
Bearbeitung der Schmetterlinge von Europa, vol. 4, p. 110,
1849.— Westwood and Humphreys, British moths and their
transformations, p. 274, 1854.—Stainton, Manual of
British butterflies and moths, vol. 2, p. 169, 1859.—Zeller,
Verh. zool.-bot. Ges. Wien, vol. 25, p. 336, 1876; vol. 34, p.
310, 1884.—Riley and Howard, Insect Life, vol. 2, p. 277,
1890.
Tinea zeae Fitch, Second report on the noxious, beneficial and
other insects, of the State of New York, p. 320, 1856.
Ephestia zeae (Fitch) Clemens, Proc. Acad. Nat. Sci. Philadel-
phia, p. 206, 1860.
Ephestia interpunctalis (Hiibner) Butler, Ent. Monthly Mag.,
vol. 15, p. 278, 1879.—Druce, Biologia Centrali-Americana,
Lepidoptera, Heterocera, vol. 2, p. 286, 1896.
Unadilla latercula Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 7,
p. 255, 1901 (New synonymy).
Ephestia glycinivora Matsumura, ‘“‘Oyo-Kinchyugakii”’ (Applied
entomology), pt. 1, p. 561, 1917; Dai-Nippon Gaichyu
Zensho (Injurious insects of the Japanese Empire), ed. 2,
vol. 1, p. 529, 1920 (spelled here as glycinivorella).—Shibuya,
Konchu Sekai, vol. 36, p. 225, 1982 (makes synonym of
interpunctella).
Ephestia (Strymaz) latercula (Hampson) Richards and Thomson,
Trans. Ent. Soc. London, vol. 80, p. 202, 1932.
Forewing with basal area ocherous white to pale
ocherous, in fresh specimens well contrasted against re-
mainder of wing; area beyond antemedial line rusty
ocherous to reddish fuscous, the red shading marked
along costal edge; central area more or less heavily
dusted with leaden fuscous, the fuscous scaling forming
an irregular blotch surrounding an ocherous line along
the discocellular vein; transverse lines sublustrous,
leaden lined; antemedial line rather broad, irregular,
set well out towards middle of wing, oblique and
slightly curved; subterminal line obscure, somewhat
sinuate, more or less parallel with termen; a faint leaden
line along termen. Hind wing pale, shining, fuscous;
veins and terminal margin faintly darkened. The red
scaling is pronounced on thorax and labial palpi as well
as on costa of forewing. Alar expanse, 12-19 mm.
Male genitalia with transtilla a smooth, uninterrupted
band; uncus not at all expanded at apex. Female geni-
talia with sclerotized projection of ductus bursae behind
genital opening rather broad, variously shaped in indi-
vidual specimens, angulate or rounded. Number of
spines of signa also variable.
TYPE LocauitTiEs: Europe (interpunctella, type lost);
New York (zeae, in USNM); Nassau, Bahamas (later-
cula, in BM); Japan (glycinivora, in Hokkaido Imperial
Univ., Sapporo, Japan).
Foon: All kinds of stored and dried vegetable prod-
ucts.
_ Distrrpution: Cosmopolitan.
This well-known and ubiquitous pest (‘‘the%Indian
meal moth” of economic literature) requires little com-
299
ment. It has an enormous literature. I have cited
above only the more important systematic references
and papers dealing with the physiology, morphology
and biology of the insect. The fullest treatment of
Plodia interpunctella and the common Ephestia species
will be found in the paper by Richards and Thomson
and that by Lehmensick and Liebers (1938). They
contain an extended bibliography, as does also the paper
by Hamlin, Reed, and Phillips. For additional refer-
ences the “Index of American Economic Entomology”
and the ‘Review of Applied Entomology” should be
consulted.
Hampson’s latercula, here placed in synonymy, has
none of the diagnostic characters of the genus Unadilla
in which he placed it. Itis obviously a Plodia. Ihave
seen the moth from Colombia he associated with his
type and have examined its genitalia; and Clarke and
Tams have compared the latter with the genitalia of the
type from the Bahamas. The moths themselves are in
very poor condition and show the basal area of the wing
somewhat darker than in typical interpunctella, but the
female genitalia show variation only of an individual
nature.
621. Plodia dolorosa Dyar
Figures 625, 1119
Plodia dolorosa Dyar, Ins. Inse. Menstr., vol. 7, p. 63, 1919.—
Richards and Thomson, Trans. Ent. Soc. London, vol. 80,
p. 204, 1932.
Forewing dark gray-brown (on the female type, the
only unrubbed specimen before me, with a purplish
sheen), the veins faintly darkened by black scaling;
transverse lines and discal marks obsolete. Hind wings
whitish fuscous, darker on the female, with veins clearly
outlined by dark scaling and a fine, dark fuscous line
along terminal margin. Alar expanse, 15-18 mm.
Male genitalia with the elements of transtilla slightly
curled and weakly united at their apices (not the smooth
band of interpunctella); apical process of gnathos some-
what larger; uncus a trifle broader and widened slightly
at apex. Female genitalia with ductus bursae narrower;
the dorsal projection at genital opening narrowly tri-
angulate and pointed; signa more reduced.
TypE Locauity: Cayuga, Guatemala (typein USNM).
Foop piant: Unknown.
Represented in the National Collection by a series of
12 specimens, both males and females, from the type
locality (Mar., Apr., May, Sept., Oct.).
Genera 186-188: Anagasta to Nicetiodes
[Venational division E. Forewing with 9 veins; 10 from the
cell; 9 absent; 4 absent; 2 and 3 from the cell. Hind wing with
discocellular vein curved. Male genitalia with transtilla com-
plete or its elements greatly enlarged and touching at apices
(except in Anagasia). Female genitalia with ductus bursae un-
sclerotized except, occasionally, immediately adjacent to genital
opening.]
186. Anagasta, new genus
Typs or Genus: Ephestia kiihniella Zeller.
Tongue well developed. Antenna simple and pubes-
300
cent in both sexes. Labial palpus upturned, not reach-
ing vertex in male, extending slightly above in female.
Maxillary palpus minute, filiform. Forewing smooth;
9 veins; venation individually variable; vein 2 from
well before lower outer angle of cell; 4 absent; 3 and 5
from the lower angle of cell, normally closely approxi-
mate at base, occasionally connate or shortly stalked;
6 normally from below upper angle of cell and only
slightly curved towards base (nearly straight), occa-
sionally curved upward to the angle and closely approxi-
mate to 8; 8 and 9 united; 10 from the cell, separated
from 8 at base; male without costal fold. Hind wing
with vein 2 from well before lower outer angle of cell;
3 and 5 normally approximate at base, rarely connate
or very shortly stalked and, when so, usually on only
one hind wing of an individual specimen; 7 and § anasto-
mosed for most or all their lengths beyond cell; cell
about one-third the length of wing; discocellular vein
curved. Abdomen of male with two pairs of short, sim-
ple, rather weak, dorsal hair tufts on eighth segment.
Male genitalia with apical projection of gnathos a
very small, shortly bifurcate hook. Transtilla incom-
plete, its elements long, slender, and widely separated
at their apices. Uncus long and moderately broad
(longer than tegumen), beyond its broad base slightly
tapering to rounded apical margin. Harpe elongate;
costa produced at apex into a short, free hook. Aedeagus
enlarged and ventrally cleft at apex. Vinculum stout,
but slightly longer than greatest width, tapering slightly
to broad terminal margin.
Female genitalia with apophyses of ovipositor and
intersegmental area between ovipositor and eighth seg-
ment collar very long; ductus bursae membranous
throughout, strongly spined at junction with bursa;
genital opening simple and unsclerotized; signa present,
consisting of from one to four separate, more or less elon-
gate and angled discs; ductus seminalis from near
middle of bursa, approximate to signa.
The generic separation of kuhniella from the other
stored-product insects of the Ephestia group requires
some justification in view of the close association of all
of them in habits, distribution, and indoor association;
but in any nice definition of E/phestia, kihniella fits
very badly. On female genitalia it could squeeze into
Ephestia, even though its long extruded ovipositor sets
it apart; but on male characters it differs radically. It
has no trace of a costal fold, while this structure is
strongly developed in all the true H’phestia. Its trans-
tilla is not only incomplete but the elements are slender
and no wise enlarged, connected, or even approximate
at their apices, quite the opposite of the development of
that organ in Hphestia. Also its venation is erratic,
even for a phycitid. Veins 6 and 8 of forewing are
parallel from a short distance beyond cell as in H’phestia
as defined in this paper; but the condition of 3 and 5 of
fore and hind wings is variable to a degree not found in
the true Ephestias. In addition to its adult characters
kithniella has the dorsum of the pupal thorax rugose.
In Ephestia the thorax of the pupa is smooth. Alto-
gether kiihniella is an aberrant species, probably of a
/UNITED STATES NATIONAL MUSEUM BULLETIN 207
different faunal origin from some, at least, of the
stored-product Ephestia species. Both they and kih-
mella are obviously of Old World origin. From all
available evidence the true home of kiihniella seems to
be the Mediterranean region, probably Asia Minor, as
suggested in 1930 by both Lebedev and Klemm.
622. Anagasta kiihniella (Zeller)
Figures 626, 1124
Ephestia kihniella Zeller, Stettiner, Ent. Zeit., vol. 40, p. 466,
1879.—Snellen, Tijds. voor Ent., vol. 28, p. 237, 1885.—
Barrett, Ent. Monthly Mag., vol. 23, p. 255, 1887.—Klein,
Proc. Ent. Soc. London, p. lii, 1887.—Omerod, Insect Life,
vol. 1, p. 314, 1889.—Fletcher, Ent. Soc. Ontario, Twentieth
Ann. Rept., p. 95, 1889. Insect Life, vol 2, p. 187, 1889;
Canadian Ent., vol. 22, p. 41, 1890.—Hulst, Phycitidae of
N. Amer., p. 198, 1890; U. S. Nat. Mus. Bull. 52, p. 435,
1903.—Riley, Insect Life, vol. 5, p. 276, 1892.—Danysz,
Mém. de Lab. Parasit. yég. Bourse de Commerce, vol. 1,
pp. viii-58, 1893.—Howard, Insect Life, vol. 7, p. 416,
1895.—Johnson, in Forbes, Nineteenth report of the State
Entomologist on the noxious and beneficial insects of the
State of Illinois, 65 pp., 1895-1896.—Druce, Biologia-Cen-
trali Americana, Lepidoptera, Heterocera, vol. 2, p. 286,
1896.—Fuller, Agr. Gaz. New South Wales, vol. 7, pp. 444—453,
1896.—Lounsbury, Ent. News, vol. 10, p. 291, 1899.—
Hampson in Ragonot, Monograph, pt. 2, p. 279, 1901.—
Carpenter, Econ. Proc. Roy. Dublin Soc., vol. 1, p. 209,
1903.—Barrett, Lepidoptera of the British Islands, vol. 10,
p. 54, 1905.—Spuler, Die Schmetterlinge Europas, vol. 2,
p. 201, 1910.—Mosher, Bull. Illinois State Lab. Nat. Hist.,
vol. 12, p. 24, fig. 74, 1916.—Durant and Beveridge, Journ.
Roy. Army Med. Corps., vol. 20, pp. 615-634, 1913; reprint
with notes, Brit. Mus. (Nat. Hist.), London, 1918.—Burk-
hardt, Zeitschr. Angew. Ent., Berlin, vol. 6, pp. 25-60,
1919 (biology).—Whiting, Journ. Exp. Zool., vol. 28, pp.
413-441 (genetics).—Forbes, Cornell Mem. 68, p. 635,
1923.—Curran, Sci. Agr., vol. 6, p. 385, 1926.—Hering, Anz.
fiir Schadlingsk., vol. 2, p. 139, 1926.—Richardson, Journ.
Agr. Res., vol. 32, p. 895, 1926.—White, Proc. Ent. Soe.
Washington, vol. 29, p. 147, 1927.—Candura, Boll. Lab.
Zool. Portici, vol. 21, p. 149, 1928.—Hudson, Butterflies and
moths of New Zealand, p. 156, 1928:—Meyrick, Revised
handbook of British Lepidoptera, p. 889, 1928.—Kiihn and
Henke, Ges. Wiss. Géttingen, Math. Phys., Abh., new ser.,
vol. 15, 121 pp., 1929 (physiology).—Brindley, Ent. Soc.
Amer. Ann., vol. 23, p. 740, 1930.—Klemm, Mitt. Ges.
Vorratsschutz, vol. 6, p. 26, 1930.—Lebedev, Zeitschr.
Angew. Ent., Berlin, vol. 6, pp. 597—605, 1930.—Noyes, Bull.
Ent. Res., vol. 21, p. 77, 1930.—Richards and Herford, Ann.
Appl. Biol., vol. 17, p. 380, 1980.—Clausen, U. 8. Dep. Agr.
circ. 168, p. 92, 1931 (Japanese record).—Richards and
Thomson, Trans. Ent. Soc. London, vol. 80, p. 177, 1932.—
Norris, Proc. Zool. Soc. London, pp. 597-611, 1932; pp.
903-934, 1933; pp. 333-360, 1934.—Dickins, Trans. Ent.
Soc. London, vol. 85, p. 333, 1986.—Busnel, Rev. Path.
Veg. et Ent. Agr. France, pp. 187-162, Apr. 1937 (arval
anatomy and physiology).—Lehmensick and Liebers, Zeits.
angew. Ent., Berlin, vol. 24, p. 442, 1937 (egg).—Pierce and
Metcalf, Genitalia of the British Pyrales, p. 7, 1938.—Mc-
Dunnough, Check list, No. 6399, 1939.—Corbet and Tams,
Entomologist, vol. 78, p. 87, 1945.
Ephestia fuscofasciella Ragonot, N. Amer. Phycitidae, p. 17,
1887.—Hulst, Phycitidae of N. Amer., p. 198, 1890.—
Hampson, in Ragonot, Monograph, pt. 2, p. 278, 1901.
Ephestia gitonella Druce, Biologia Centrali-Americana, Lepi-
doptera, Heterocera, vol. 2, p. 286, 1896.
Ephestia sericaria Authors (not Scott), Ragonot, Bull. Soc. Ent.
France, vol. 61, p. cclxxiv, 1892.—Richards and Thomson,
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 301
Trans. Ent. Soc. London, vol. 80, pp. 171, 177, 1932.—
Corbet and Tams, Proc. Ent. Soc. London (ser. B), vol. 113,
p. 68, 1943; Entomologist, vol. 76, p. 15, 1943.—Hinton,
Bull. Ent. Res., vol. 34, p. 195, 1943.
Forewing gray, more or less dusted with black and
with blackish markings; antemedial line grayish white
(this pale shade on some specimens extending over the
remainder of the basal area), slightly oblique and
irregularly dentate, bordered outwardly by a black
shade varying from a narrow line to a broad suffu-
sion; subterminal line pale gray, obscure on most
specimens, variable and irregularly dentate, usually
with a dark inner border; veins in outer area of wing
somewhat darkly outlined; discal dots at end of cell
pronounced, sometimes confluent along discocellular
vein but usually well separated and the lower dot
elongated slightly along veins 3 and 5; a row of faint
dark dots along termen. Hind wings translucent,
whitish, the veins and terminal margin pale brownish
fuscous. Alar expanse, 18-28 mm.
Genitalic characters as given for the genus. The
male genitalia are remarkably uniform and exhibit little
individual variation, considering the variability of the
species otherwise. The female genitalia are variable
(see fig. 1124) in the number and shape of the signa.
Typr Locauities: Germany (kihniella, in BM);
Durango, Mexico (gitonella, in BM); Texas (fusco-
Jfasciella, in Paris Mus.).
Foon: Flour, grains, nuts, dried fruits, chocolate,
seeds, biscuits, cakes, jellies, candy, and a host of other
dried and stored vegetable products. The favored
foods are wheat and other grain flours. Other records,
unless based upon reared adults, should be received
with caution; for the larvae are very difficult to dis-
tinguish from those of a number of other phycitids
feeding on dried vegetable matter, and records based
upon larval identifications are as apt to be incorrect as
not. The species is of major economic importance only
as a flour and milled-grain feeder and is often a serious
pest in flour mills. It is apparently an indoor insect
and requires a high temperature throughout the year
for maximum development.
DistrisutTion: Nearly cosmopolitan.
“The Mediterranean flour moth” has probably as
extensive a literature as any other lepidopteron. Only
the more important systematic, biological, and physio-
logical references are given above. For further refer-
ences the papers by Noyes (1931) and Richards and
Thomson (1932), the ‘Index of American Economic
Entomology,” “Zoological Record,” and ‘‘Review of
Applied Entomology’’ should be consulted.
Considering its variability kiihniella has remarkably
little synonymy except for numerous misspellings of its
specific name. A list of these will be found in the
Richards and Thomson paper (1932). Most modern
authors use the equivalent spelling kuehniella in place
of kiihniella; but as the latter was the original ortho-
graphic form and the species was named in honor of
Professor Kiihn of Halle we are obliged by the inter-
national rules to follow it.
The name fuscofasciella Ragonot applies to a color
form with broad blackish borders to the transverse
lines and a contrasted pale area of forewing. Such
examples occur rather frequently in our Southwestern
States. Unless venation is noted, it is easy to confuse
kithniella with large examples of Vitula edmandsae.
Pale specimens of the two species are strikingly similar
in habitus.
The original specimens before Zeller when he described
kithniella are in the British Museum but, according to
Clarke and Tams, no one specimen bears a type desig-
nation so I am here designating a male from the original
series as lectotype. This specimen bears three labels,
the first a small white one with the inscription,
“excl. 4/7/77’; a second label. inscribed in Zeller’s
handwriting “Zphestia kiihnii Z,’’ “MSS.Z”; and a
third label with the inscription ‘‘Zell. Coll. 1884.’”’ Tams
concurs in this selection.
187 Genus Ephestia Guénée
Ephestia Guénée, Europaeorum Microlepidopterorum index
methodicus . . . p. 81, 1845; Ann. Soc. Ent. France, ser. 2,
vol. 3, p. 319, 1845.—Zeller, Isis von Oken, 1848, pp. 585,
592.—Herrich-Schaffer, Systematische Bearbeitung der
Schmetterlinge von Europa, vol. 4, p. 110, 1849.—Heine-
mann, Schmetterlinge Deutschlands und der Schweiz, Abt.
2, vol. 1, pt. 2, p. 201, 1865.—Meyrick, Proc. Linn. Soc.
New South Wales, vol. 3, p. 215, 1878; vol. 4, p. 234, 1879;
vol. 7, p. 160, 1882; Revised handbook of British Lepidop-
tera, p. 387, 1928.—Hulst, Phycitidae of N. Amer., p. 197,
1890; U. S. Nat. Mus. Bull. 52, p. 434, 1903.—Ragonot,
Monograph, pt. 2, p. xiii, 1901.—Hampson, in Ragonot,
Monograph, pt. 2, p. 271, 1901.—Forbes, Cornell Mem. 68,
p. 634, 1923.—Richards and Thomson, Trans. Ent. Soc.
London, vol. 80, p. 171, 1932.—Bisset in Pierce and Metcalfe,
Genitalia of the British Pyrales, p. 58, 1938. (Type of
genus: Tinea elutella Hiibner.)
Hyphantidium Scott, Proc. Zool. Soc. London, vol. 27, p. 207,
1859, (Type of genus: Hyphantidium sericariwm Scott.)
Cadra Walker, List, vol. 30, p. 961, 1864 (Type of genus: Cadra
defectella Walker.)
Tongue well developed. Antenna simple and pubes-
cent in both sexes. Labial palpus upturned, reaching
slightly above vertex. Maxillary palpus minute, fili-
form. Forewing smooth; 9 veins; vein 2 from before
but rather near lower outer angle of cell; 3 from outer
angle; 4 absent; 5 more or less approximate to 3 at base;
6 from below upper angle of cell, straight or slightly
curved towards base (from a slight distance beyond base
running parallel with 8) ; 9 absent; 10 from the cell; male
with a strong costal fold enclosing scale tuft. Hind
wing with vein 2 from well before lower outer angle of
cell; 3 and 5 approximate at base or short stalked
(elutella); 7 and 8 anastomosed for most or all of their
lengths beyond cell; cell one-half or slightly less than
one-half the length of wing; discocellular vein curved.
Abdomen of male with compound dorsal tufts.
Male genitalia with gnathos terminating in a pair of
well-spaced prongs. Transtilla complete or, if elements
slightly separated, their apices broadly developed, if
complete with flaring lateral projections. Uncus short,
broad; its terminal margin bluntly rounded. Aedeagus
not appreciably enlarged at apex; penis armed with a
302
sclerotized band or a row) of sclerotized wrinklings.
Vinculum stout, longer than broad, tapering slightly to
blunt terminal margin.
Female genitalia with apophyses of ovipositor and
intersegmental area between ovipositor and eighth seg-
ment collar short; ductus bursae strongly spined towards
bursa or bearing a longitudinal row of sclerotized ridges;
signa present, consisting of a row of narrow sclerotized
discs; ductus seminalis from bursa approximate to signa.
Ephestia.as here defined is an Old World genus. Even
with kihniella and the formerly included American
species removed, it remains somewhat of a heteroge-
neous group. I suspect that the type species (elutella)
has a different faunal origin than cautella, figulilella,
callideila, and affatella, which, because of their similar
habits, present distribution and genitalic structures are
associated with it; but until the Old World species now
in Ephestia can be thoroughly studied and the incon-
gruous ones given a better generic placement we shall
have to keep the more-or less cosmopolitan species
together. However, elutella has a different hind wing
venation and a different type of harpe from the others
and I believe will eventually be generically distin-
guished.% Three species are represented in the Amer-
icas, all apparently imports from the Old World.
They separate into groups as follows:
Hind wing with veins 3 and 5:stalked; costa of harpe smooth.
Hind wing with veins 3 and 5 approximate at base; harpe
with digitate projection from middle or near middle
of costa.
Color and markings are of little help in distinguishing
the species. There is so much individual variation and
so little specific difference, even between typical speci-
mens, that superficial habitus can not be trusted.
Genitalia on the other hand do readily identify and
distinguish the species. Their specific characters are
several, obvious, and consistent. These organs should
be examined in every instance when specific identifica-
tion is of any importance.
Genus E’phestia, Species 623: E. elutella
[Hind wing with veins 3 and 5 stalked; costa of harpe smooth.]
623. Ephestia elutella (Hiibner)
Ficures 126, 628, 1122
Tinea elutella Hubner, Sammlung europaischer Schmetterlinge,
Lepidoptera 8, Tinea 5, pl. 24, fig. 163, 1796.
Phycis elutea Haworth, Lepidoptera Britannica, p. 496, 1811
(amended spelling for elutelia).
Phycis semirufa Haworth, Lepidoptera Britannica, p. 496, 1811.
Phycis rufa Haworth, Lepidoptera Britannica, p. 497, 1811.
Phycis elutella (Hiibner) Zincken, in Germar and Zincken, Mag.
der Ent., vol. 3, p. 175, 1818.—Treitschke, Die Schmetter-
linge von Kuropa, vol. 9, p. 194, 1832.—Duponchel, Histoire
naturelle des lépidoptéres, ou papillons de France, vol. 10,
p. 204, 1836.
8 Ephestia vitivora Filipjev (Deuts, Ent. Zeit. “Iris,” Dresden,
vol. 45, p. 70, 1931) from Transcaucasia is distinct from, but
obviously very close to, and in the event of any further division
of Ephestia, congeneric with elutella—suggesting a western Asiatic
origin for the latter.
(
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Phycita semirufa (Haworth) Stephens, Illustrations of British
entomology, Haustellata, vol. 4, p. 305, 1834.— Wood, Index
entomologicus, p. 210, pl. 46, fig. 1457, 1839.—Westwood
and Humphreys, British moths and their transformations,
vol. 2, p. 229, 1845.
Phycita elutella (Hibner) Stephens, Mlustrations-of British ento-
mology, Haustellata, vol. 4, p. 804, 1884—Wood, Index
entomologicus, p. 210, pl. 46, fig. 1454, 1839.—Westwood
and Humphreys, British moths and their transformations,
vol. 2, p. 229, 1845.
Myelois elutella (Hiibner) Zeller, Isis von Oken, 1839, pp. 176,
343.
Ephestia elutella (Hiibner) Guénée, Europaeorum Microlepidop-
terorum index methodicus..., p. 81, 1845.—Herrich-
' Schaffer, Systematische Bearbeitung der Schmetterlinge von
Europa, vol. 4, p. 110, 1849.—Stainton, Manual of British
butterflies and moths, vol. 2, p. 168, 1859.—Heinemann, Die
Schmetterlinge Deutschlands und der Schweiz, Abt. 2, vol. 1,
pt. 2, p. 201, 1865.—Zeller, Verh. zool.-bot. Ges. Wien, vol.
25, p. 338, 1876.—Hulst, Phycitidae of N. Amer., p. 200,
1900; U.S. Nat. Mus. Bull. 52, p. 435, 1903.—Hampson, in
Ragonot, Monograph, pt. 2, p. 300, 1901.—Forbes, Cornell
Mem. 68, p. 635, 1923.—Curran, Sci. Agr.; vol. 6, p. 386,
1926.—Meyrick, Revised handbook of British Lepidoptera,
p. 388, 1928.—Munro and Thomson, Empire Marketing
Board, No. 24, London, H. M. Stationery Office, p. 22,
1929.—Noyes, Bull. Ent. Res., p. 80, 1930.—Richards and
Herford, Ann. Appl. Biol., vol. 17, p. 380, 1930.—Back and
Reed, Journ. Econ. Ent., vol. 23, p. 1004, 1930.—Richards
and Thomson, Trans. Ent. Soc. London, vol. 80, p. 185,
1932.—Bovingdon, Empire Marketing Board, No. 67, pp. 1—
88, 1933.—Norris, Proc. Zool. Soc. London, 1934, pp. 333—
360, 1934.—Dickins, Trans. Ent. Soc. London, vol. 85,
p. 348, 1936.—Lehmensick and Liebers, Zeitschr. angew.
Ent., Berlin, vol. 24, p. 442, 1937.—Pierce and Metcalfe,
Genitalia of the British Pyrales, p. 6, 1938.—McDunnough,
Check list, No. 6401, 1939.—Hinton, Bull. Ent. Res., vol.
34, p. 196, 1943.—Corbet and Tams, Proc. Zool. Soc. Lon-
don, vol. 113, ser. B, p. 64, 1943.
Hypantidium sericariwm Scott, Proc. Zool. Soc. London, vol. 27,
p. 207, 1859.—Corbet and Tams, Entomologist, vol. 78,
p. 87, 1945 (establish synonymy with elutella).
Hphestia semirufa (Haworth) Staudinger, Horae Soc. Ent. Ros-
sicae, vol. 15, p. 229, 1879.
Ephestia roxburghii Gregson, Entomologist, vol. 6, p. 318, 1873.—
Ragonot, Monograph, pt. 2, p. 301, 1901.—Richards and
Thomson, Trans. Ent. Soc. Lond., vol. 80, p. 187, 1932.
Ephestia elutella unicolorella Staudinger, Horae Soc. Ent. Ros-
sicae, vol. 15, p. 228, 1879.
Ephestia unicolorella Staudinger, Horae Soc. Ent. Rossicae, vol.
16, p. 89, 1881.—Ragonot and Hampson, in Ragonot, Mono-
graph, pt.2, p. 229, 1901.—Richards and Thomson, Trans.
Ent. Soc. London, vol. 80, p. 187, 1932.
Ephestia amarella Dyar, Proc. U. 8. Nat. Mus., vol. 27, p. 921,
1904.
Forewing dark brown to pale gray, sometimes with a
paler shade between lower fold and inner margin; color
variable; transverse lines usually distinguishable; ante-
medial line obscure, oblique, bordered outwardly by a
narrow dark shade, the latter approaching middle of
wing; subterminal line somewhat sinuate, faintly bor-
dered on both sides by dark lines. Hind wing smoky
white to pale fuscous. Alar expanse, 13-18 mm.
Male genitalia with prongs of apical process of
gnathos forming a V, more narrowly spaced than in
other species of the genus. Harpe without projections
from costa. Female genitalia with ductus bursae rather
coarsely spined for about half its length towards bursa;
bursa evenly and minutely spined.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
TypE Locaitizs: Germany (elutella, type lost); Eng-
land (semirufa, type lost; rufa and roxburghii, in BM);
Malatia, Asia Minor (unicolorella, in Mus. Univ. Ber-
lin); Australia (sericarium, in BM); Kalso, Kootenai
District, British Columbia (amarella), in USNM).
Foon: Dried fruits, seeds, nuts, chocolate, cocoa,
tobacco, and a number of other dried vegetable prod-
ucts. Feeding records based on larval identifications
should be accepted with extreme caution; for the larvae
of elutella, are often impossible to distinguish from
those of cautella, and have been frequently confused
with them. The so-called characters given in literature
for separation of the larvae of the two species are not
constant and should not be depended upon for specific
identification. In both Europe and the United States
elutella seems to breed out-of-doors to some extent but
has been noticed as a pest only in storage houses, in
England chiefly on stored cocoa and in America on
tobacco. In our economic literature it is known as
“the tobacco moth.” Aside from its depredations in
tobacco warehouses it appears to be only a minor and
occasional pest on other stored products in the United
States.
Distripution: Practically cosmopolitan, but appar-
ently less numerous and not so generally distributed as
either Plodia interpunctella or E’phestia cautella.
The moths are easily separable from the other two
species found here (cautella and figulilella) and from
other truly congeneric European species by the short
stalking of veins 3 and 5 of hind wing. In other true
Ephestia these veins are always approximate at base.
They should be examined in every instance where
specific identification is of any importance.
The above synonymy gives only the more important
references. Some Old World references cited by
Richards and Thomson (1932) and Hulst (1903) have
been omitted and, except for a few essential articles,
no attempt has been made to cover the economic
literature, which is both extensive and repetitious.
Genus Ephestia, Species 624 and 625: E. cautella
and #. figulilella
[Hind wing with veins 2 and 5 approximate at base; harpe with
digitate projection from middle or near middle of costa.]
624, Ephestia cautella (Walker)
Fieurns 125, 629, 1121
Pempelia cautella Walker, List, vol. 27, p. 73, 1863.
Cadra defectella Walker, List, vol. 30, p. 962, 1864.
Nephopteryx desuetella Walker, List, vol. 35, p. 1719, 1866.
Ephestia cahiritella Zeller, Stettiner, Ent. Zeit., vol. 28, p. 384,
1867.—South, Entomologist, vol. 23, p. 304, 1890.—Van
Deventer, Tijdschr. voor Ent., vol. 46, p. 80, 1904.—
Barrett, Lepidoptera of the British Islands, vol. 10, p. 56,
1905.
Ephestia passulella Barrett, Ent. Monthly Mag., vol. 11, p. 271,
1875.—Ragonot, Ent. Monthly Mag., vol. 22, p. 24, 1885.
Salebria cautella (Walker) Cotes and Swinhoe, Catalogue of the
moths of India, pt. 6, p. 675, 1889.
303
Ephestia cautella (Walker) Hampson, Moths, vol. 4, p."66, 1896,
in Blanford, Fauna of British India.—Ragonot, Monograph,
pt. 2, p. 292, 1901.—Hulst, U. S. Nat. Mus. Bull. 52, p. 434,
1903.—Spuler, Die Schmetterlinge Europas, vol. 2, p. 202,
1910.—Chittenden, U. 8. Dep. Agr. Bur. Ent. Bull. 104,
40 pp., 1911.—Dyar, Proc. U. S. Nat. Mus., vol. 47, p.'345,
1914.— Forbes, Cornell Mem. 68, p. 635, 1923.—Curran,
Sci. Agr., vol. 6, p. 386, 1926.—Meyrick, Revised Handbook
of British Lepidoptera, p. 388, 1928.—Shibuya, Journ.
Faculty Agric. Hokkaido Imperial Univ., Sapporo, vol. 22,
p. 77, 1298.—Richards and Herford, Ann. Appl. Biol., vol.
17, p. 380, 1930.—Noyes, Bull. Ent. Res., p. 80, 1930.—
Simmons, Reed, and McGregor, U. 8. Dep. Agr. Cir. 157,
p. 36, 1931.—Keifer, Monthly Bull. Dep. Agr. California,
vol. 20, p. 619, 1931.—Richards and Thomson, Trans. Ent.
Soe. London, vol. 80, p. 197, 1932.—Bovingdon, Empire
Marketing Board, No. 67, pp. 1-88, 1933.—Norris, Proc.
Zool. Soc. London, 1934, pp. 333-360, 1934.—Dickins,
Trans. Ent. Soc. London, vol. 85, p. 342, 1936.—Lehmensick
and Liebers, Zeitschr. angew. Ent., Berlin, vol 24, p. 443,
1937.—Pierce and Metcalfe, Genitalia of the British Pyrales,
p. 7, 1938.—McDunnough, Check list, No. 6403, 1939.—
Hinton, Bull. Ent. Res., vol. 34, p. 193, 1943.—Corbet
and Tams, Proc. Zool. Soc. London, vol. 113, ser. B, p. 64,
1943.
Cryptoblabes formosella Wileman and South (<), Entomologist,
vol. 51, p. 219, 1918.—Shibuya, Journ. Faculty Agric.
Hokkaido Imperial Univ., Sapporo, vol. 22, pp. 17, 88, 1928.
Color and markings similar to those of elutella and as
variable, expecially on the females. On the average
the antemedial line is less oblique in both cautella and
Jigulilella than in elutella, but this difference does not
hold for all specimens. £. cavutella is distinguished by
its genitalia, both male and female. They are quite
distinct from those of any other phycitid.
Alar expanse, 11-18 mm.
Male genitalia with prongs of apical process of
gnathos forming a broad U, widely spaced. Harpe
with a short, digitate projection from near middle of
costa. Transtilla with a broad fusion at middle and
prominent, round, flattened, lateral lobes.
Female genitalia with ductus bursae bearing a longi-
tudinal row of strongly sclerotized and pigmented
ridges, the latter forming 9, narrow, fluted band some-
what over half as long as the ductus. Bursa copulatrix
rather coarsely spined over about half its surface.
Typr tocauitiges: Ceylon, India (cautella and de-
fectella, in BM); Moreton Bay, Queensland, Australia
(desuetella, in BM); Cairo, Egypt (cahiritelia, in BM);
England (passulella, in BM); Takow, Formosa (for-
mosella, in BM).
Foon: All kinds of dried, stored vegetable products.
Distrizution: Cosmopolitan.
This species is as abundant, and probably as destruc-
tive to dry fruits, grains, nuts, and other stored vege-
table products, as Plodia interpunctella. Surprisingly
enough it has fewer economic references than either
elutella or figulella though it appears to be more widely
distributed and more of a general nuisance than either
of them. In the list of common names approved by
the American Association of Economic Entomologists
it is known as the “almond moth.” In economic litera-
ture generally it has also been referred to as the ‘‘dried
currant moth” and the “fig moth.”
304
625. Ephestia figulilella Gregson
Fieures 630, 1123
Ephestia figulilella Gregson, Entomologist, vol. 5, p. 385, 1871.—
Ragonot, Ent. Monthly Mag., vol. 22, p. 25, 1885.—Hampson
and Ragonot, in Ragonot, Monograph, pt. 2, p. 282, 1901.—
Hulst, U. S. Nat. Mus. Bull. 52, p. 435, 1903.—Caradja,
Deutsche Ent. Zeitschr., ‘‘Iris,’’ vol. 24, p. 210, 1910.—Spuler,
Die Schmetterlinge Europas, vol. 2, p. 202, 1910.—Turati and
Zanon, Atti Soc. Italiana Sci. nat., vol. 61, p. 164, 1922.—
Keifer, Monthly Bull. Dep. Agr. California, vol. 20, p. 619,
1931.—Richards and Thomson, Trans. Ent. Soc. London,
vol. 80, p. 194, 1982.—Donohoe and Barnes, Journ. Econ.
Ent., vol. 27, pp. 1070, 1075, 1934.—Pierce and Metcalfe,
Genitalia of the British Pyrales, p. 6, 19838.—McDunnough,
Check list, No. 6402, 1939.—Hinton, Bull. Ent. Res., vol.
34, p. 194, 1943.—Corbet and Tams, Proc. Zool. Soc. Lon-
don, vol. 113, ser. B, p. 68, 1948.—Donohoe et al., U. S.
Dep. Agr. Techn. Bull. 994, pp. 1-23, 1949.
Ephestia ficulella Barrett, Ent. Monthly Mag., vol. 11, p. 271,
1875 (emended spelling for jigulilella)—Meyrick, Proc.
Linn. Soe. New South Wales, vol. 4, p. 244, 1880.—Chittenden,
U. S. Dep. Agr. Div. Ent. Bull. 8 (new ser.), p. 10, 1897.—
Meyrick, Revised hand book of British Lepidoptera, p. 388,
1928.
Ephestia millerit Zeller, Verh. zool.-bot. Ges. Wien., vol. 25, p.
339, 1876.—Druce, Biologia Centrali Americana, Lepidop-
tera, Heterocera, vol. 2, p. 287, 1890.
Ephestia figuliella Forbes, Cornell Mem. 68, p. 635, 1923 (mis-
spelling).
Ephestia figulella Curran, Sci. Agr., vol. 6, p. 386, 1926 (mis-
spelling).
Ephestia venosella Turati, Atti Soc. Italiana Sci. Nat., vol. 65,
p. 58, 1926.
Ephestia ernestinella Turati, Atti Soc. Italiana Sci. Nat., vol. 66,
p. 330, 1927.
The genitalia are distinctive but there are no con-
sistent superficial characters for the separation of figu-
hilella from cautella.
Alar expanse, 12-17 mm.
Male genitalia with prongs of apical process of
gnathos U-shaped with the base of the U flattened and
the prongs well separated. Harpe with a long digitate
projection from midcosta. Transtilla with its elements
broadened and touching (but not fusing) at their apices.
Female genitalia with ductus bursae armed with a
spiral of short, broadly based spines; a cluster of similar
spines at terminal end of bursa copulatrix; a pair of
fan-shaped scale tufts from intersegmental area ad-
jacent to genital opening; signa consisting of a row of
a half dozen elongate, narrow discs.
Typ tocatitizs: Liverpool, England (jfigulilella, in
BM); Central America (milleri, in BM); Bengasi,
Cyrenaica (venosella, [?] Turati Coll.); Giarabub,
Cyrenaica (ernestinella, [?] Turati Coll.).
Foop: Dried fruits, nuts, seeds, meal, beans, etc.
DistrrBution: Europe, Asia, North and West Africa,
Hawaii, Australia, North and South America. In the
United States it seems to have established itself only in
California where it is a minor pest of dried raisins,
though it has been intercepted at various ports of entry
in stored products. It is apparently less widely dis-
tributed and of considerably less economic importance
than either cautella or elutella. In our economic litera-
ture it is known as the “‘raisin moth,” and has a growing
list of economic references, for which the Review of
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Applied Entomology should be consulted. Some Old
World references cited by Richards and Thomson (1932)
have been omitted from the foregoing synonymy.
188. Genus Nicetiodes Schaus
Nicetiodes Schaus, Zoologica (Contr. New York Zool. Soc.),
vol. 5, No. 2, p. 48, 1923 (Type of genus: Nicetiodes
apianella Schaus).
Tongue well developed. Antenna pubescent; shaft
of male dilated and with a shallow, ventral sinus
towards base; shaft of female simple. Labial palpus
slender, upturned, reaching slightly above vertex; third
segment acuminate, about half the length of second.
Maxillary palpus rather broadly dilated with scales on’
female, less so on male. Forewing of male with some
raised scales alone upper margin of cell towards base
(possibly a specific character), of female smooth; 9
veins; vein 2 from before but near lower outer angle of
cell; 3 and 5 from the angle, closely approximate or
connate at base; 4 absent; 6 from below upper angle,
straight; 8 and 9 united; 10 from the cell, separate from
8 at base; male with a strong costal fold enclosing hair
tuft. Hind wing with vein 2 from very close to lower
outer angle of cell; 3 and 5 stalked for less than half
their lengths; 7 and 8 anastomosed for most of their
lengths beyond cell; cell about half the length of wing;
discocellular vein curved. Abdomen of male with
compound dorsal tufts.
Male genitalia with gnathos terminating in an en-
larged, angulate knob. ‘Transtilla complete, a narrow,
low-arched bridge with slight central projection. Uncus
narrow stemmed, abruptly expanding and flatly bilobed
at extremity; a short, broad scaphium projecting from
its base. Harpe with sacculus slightly produced at
extremity, otherwise simple. Anellus tubular with at-
tached ventral shield bearing short lateral lobes.
Aedeagus long, rather stout, flaring and sharply angled
at apex; penis with a few weakly sclerotized wrinklings,
otherwise unarmed. Vinculum triangulate, longer than
broad.
Female genitalia with ductus bursae scobinate
towards bursa and narrowly sclerotized at genital open-
ing; signa present, consisting of a row of small sclero-
tized discs; ductus seminalis from bursa approximate to
signa.
A distinct genus at once distinguished by its peculiar
uncus and gnathos and projecting sacculus. Contains
but one described species.
626. Nicetiodes apianella Schaus
Ficures 127, 637, 1125
Nicetiodes apianella Schaus, Zoologica, vol. 5, No. 2, p. 48, 1923.
Forewing blackish fuscous with a few dull whitish
scales at base and a shading of them surrounding dis-
cocellular vein and in area beyond subterminal line;
transverse lines white, well separated, the antemedial
line narrow, straight, and nearly vertical; subterminal
line faint, irregularly and shortly dentate, rather near
termen; discal dots obscure. Hind wing smoky white;
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
the veins slightly darkened and a similar narrow dark
shade along terminal margin. Alar expanse, 13 mm.
Genitalic characters as given for the genus; the
tubular part of anellus partially scobinate.
Type Locatity: Conway Bay, Indefatigable, South
Seymore, Tower Island, Galipagos (type in USNM).
Foop prant: Unknown.
Known only from the male type and female paratype
from the type locality. A rather striking species con-
spicuous because of its blackish forewings with con-
trasted white antemedial line.
Genera 189-191: Varneria to Erelieva
[Venational division F. Forewing with 9 veins; 2 from the cell;
3 and 5 stalked; 4 absent; 9 absent; 10 from the cell.]
189. Genus Varneria Dyar
Varneria Dyar, Proc. Ent. Soc. Washington, vol. 6, p. 114,
1904.—Forbes, Cornell Mem. 68, p. 639, 1923. (Type of
genus: Varneria postremella Dyar.)
Tongue well developed. Antenna weakly pubescent;
shaft simple, sometimes showing a slight bend at base
(but no sinus and the bend as obvious in female as
male). Labial palpus oblique, slender, laterally flat-
tened, reaching as high as vertex; third segment slightly
less than half as long as second. Maxillary palpus
filiform. Forewing smooth; 9 veins; vein 2 from lower
outer angle of cell or from very close to the angle, con-
nate with or approximate to the stalk of 3-5 at base;
4 absent; 3 and 5 stalked; 6 from below upper angle of
cell, straight; 8 and 9 united (9 absent) ; 10 from the cell,
separated from 8 at base; male with strong costal fold
containing projecting hair tuft. Hind wing with vein
2 from close to lower outer angle of cell; 3 and 5 long
stalked; 7 and 8 completely anastomosed beyond cell;
cell one-half the length of wing; discocellular vein
curved. Abdomen of male with rather weak, com-
pound, dorsolateral scale tufts.
Male genitalia with apical process of gnathos U-
shaped, the prongs rather short and moderately stout.
Transtilla complete, a narrow curved band. Harpe
with projecting spur from base of costa, otherwise
simple. Anellus a simple shield. Aedeagus straight,
simple, not appreciably tapering; penis bearing a flat,
weakly sclerotized plate, otherwise unarmed. Vincu-
lum subtriangulate, stout, no longer or but slightly
longer than greatest width.
Female genitalia with ductus bursae sclerotized and
flattened for more than half its length from genital
opening and with sclerotized, projecting, dorsal shield
behind genital opening; bursa copulatrix with or with-
out signum, latter when present a small flat plate with
a few toothlike projections along one edge; ductus
seminalis from near middle of bursa.
Four species are here recognized as belonging to the
genus; but two of these are of doubtful specific status.
The absence of males in two of the species and the
scarcity of specimens in those known only from females
305
makes the evaluation of characters for specific separa-
tion very difficult. All of the species have a superficial
character in common in the vinous red ground color of
the forewings.
627. Varneria postremella Dyar
Ficures 88, 631, 1110
Varneria postremella Dyar, Proc. Ent. Soc. Washington, vol. 6,
p. 115, 1904.—Ely, Proc. Ent. Soc. Washington, vol. 12,
p. 203, 1910.—Forbes, Cornell Mem. 68, p. 639, 1923.—
McDunnough, Check list, No. 6406, 1939.
Forewing vinous red; basal half of costa overlaid with
blackish scales; a line of blackish scales along discal
fold, spreading somewhat between veins 3 and 6, this
blackish dusting variable and sparse in some specimens;
no transverse lines or discal spots distinguishable.
Hind wing smoky fuscous, costal and terminal margins
darker. Alar expanse, 9-10.5 mm.
Male genitalia with costal process from harpe directed
forward, hooked at apex; lower margin of harpe incurved
near apex. Terminal margin of vinculum angulate.
Female genitalia with signum present; projecting shield
behind genital opening subtriangulate, sides incurved,
apex truncate.
Type LocaLity: Kentucky (type in USNM).
Foop piant: Unknown.
Disrripution: Connecticut, East River (July) ; Mary-
land, Hyattsville (Aug.), Plummers Isl. (July); Vir-
ginia, Mountain Lake (July); Kentucky (July).
In postremella vein 2 of forewing is connate with the
stalk of 3-5. In the remaining species of Varneria it is
closely approximate. This character, however, might
be expected to vary in individual specimens if more
were available.
628. Varneria nannodes Dyar
Figure 1111
Varneria nannodes Dyar, Proc. Ent. Soc. Washington, vol. 47,
p. 346, 1914.
This species is represented only by the female type,
which in color, markings, and genitalia exhibits nothing
to distinguish it specifically from females of postremella.
It is probably a synonym of the latter; but in the ab-
sence of a male of nannodes or any matching examples
between Virginia and Panamé it were wiser to hold any
synonymy in abeyance. Alar expanse, 8.5 mm.
Typr Locatity: Cabima, Panamé (type in USNM).
Foop piant: Unknown.
629. Varneria atrifasciella Barnes and McDunnough
Figure 1112
Varneria atrifasciella Barnes and McDunnough, Contributions,
vol. 2, p. 184, 19138.—McDunnough, Check list, No. 6407,
1939.
Forewing dark vinous red rather heavily dusted with
blackish scales; in the paratype faint indications of pale
(yellowish) antemedial and postmedial lines, these, how-
ever, not distinguishable in the type and their position
chiefly indicated by the concentration of blackish scaling
306 UNITED STATES NATIONAL MUSEUM BULLETIN 207
between them. Hind wing smoky white, the veins
slightly darker. Alar expanse, 9-9.5 mm.
Female genitalia without signum; projecting shield
behind genital opening large and approximately square;
an eversible lobe in ventral membrane between oviposi-
tor and eighth-segment collar.
TyYpE Locautity: Everglades, Fla. (type in USNM).
Foop prant: Unknown.
Represented only by the type and one paratype from
the type locality (Apr.), both females.
630. Varneria dubia, new species
Fiaures 632, 1113
Similar to atrifasciella except that forewing is less
abundantly shaded with black and female genitalia show
slight differences in the shape of the shield at genital
opening and the eighth-segment collar (compare figs.
1112 and 1118). Alar expanse, 8.5-11 mm.
Male genitalia with costal process from harpe directed
backward, curved, pointed at apex; lower margin of
harpe evenly convex toward apex. Female genitalia
with projecting shield behind genital opening triangu-
late; eighth-segment collar appreciably narrower to-
wards base on dorsum; otherwise as in atrifasciella.
Typz Locatity: El Yunque, Luquillo Mts., Puerto
Rico (type in Cornell Univ. Coll.; paratypes in Cornell
Univ. and USNM, 61395).
Foon. prant: Unknown.
Described from male type and one male and four fe-
male paratypes from the type locality, Mar. 29 to Apr.
23, 1930, Cornell University lot 795 sub. 9.and 40. In
addition to the types there are fragments of a broken
specimen in the Cornell Collection from Jajome Alta,
Puerto Rico, June 18, 1930, J. A. Hoffman, collector.
This species is uncomfortably close to atrifasciella and
may be nothing more than a race or variety of it, but
in all the females before me the genitalic differences,
slight though they be, are consistent, and in the absence
of any authentic males of aétrifasciella or sufficient fe-
male examples of any Varneria species to determine the
extent of individual variation in genitalia it seems safest
to define dubia as a distinct species.
190. Genus Eurythmia Ragonot
Hurythmia Ragonot, N. Amer. Phycitidae, p. 16, 1887.—Hulst,
Phycitidae of N. Amer., p. 195, 1900.—Hampson, zn Rago-
not, Monograph, pt. 2, pp. xiii, 266, 1901. (Type of genus:
Ephestia hospitella Zeller.)
Tongue well developed. Antenna weakly pubescent,
simple. Labial palpus upturned, reaching vertex, lat-
erally flattened and somewhat rough scaled; third seg-
ment slightly more than half as long as second. Maxil-
lary palpus filiform. Forewing smooth; 9 veins; vein 2
from very close to lower outer angle of cell; 3 from the
angle, shortly stalked with 5; vein 4 absent; 6 from be-
low but near upper angle, very slightly bent; 8 and 9
united; 10 from the cell, separated from 8 at base; male
with short costal fold. Hind wing with 2 from close to
lower outer angle of cell; 3 and 5 stalked for at least half
their lengths; 7 and 8 anastomosed for most of their
lengths beyond cell (free element of 8 very short); cell
less than half the length of wing; discocellular vein
curved. Highth abdominal segment of male simple.
Male genitalia with gnathos terminating in a very
short, stout, narrowly forked projection. Transtilla
complete, arched and produced at middle into a narrowly
forked projection with shorter central spur. Harpe with
costa sclerotized for half its length, the apex of the
sclerotized margin projecting as a free spur. Anellus a
curved shield with short lateral lobes. Aedeagus taper-
ing sharply from enlarged base to bluntly pointed apex;
penis armed with an elongate sclerotized spiral band
covered with short saw-toothed spines. Vineulum tri-
angulate, longer than greatest width and tapering to a
blunt point.
Female genitalia with ductus bursae membranous
throughout; genital opening simple; bursa copulatrix
with signa a series of (8 to 6) detached, broad-based,
short, thornlike spines and narrow-bladed discs situated
near anterior end of the bursa and a cluster of several
similar, much smaller spines near junction of bursa and
ductus bursae; ductus seminalis from bursa in the neigh-
borhood of the larger signa.
Eurythmia is distinguished chiefly by the armature of
the penis, the arrangement of the signa in bursa, and the
simple eighth abdominal segment of the male. The
venation is similar to that of Varneria except for the
shorter stalking of veins 3 and 5 of forewing, normally a
character of, at most, specific significance. Ragonot
and Hampson interpret the forewing venation differ-
ently from that given here; namely, 2-3 united and 4-5
stalked rather than 4 absent and 3-5 stalked. The end
result would be the same, but the male genitalia do not
indicate any affinities to the genera having 2 and 3
stalked and tending to fuse and do show affinities to
E’phestiodes (especially the aberrant species noniella and
stictella) where veins 4 and 5 are stalked and 4 shows &
tendency to disappear (compare venation of EL. plorella,
fig. 109).
Apparently the genus as here defined is confined to
North America; at least, no tropical species have as
yet been discovered. Several have been described from
the United States on trifling differences of color and
maculation. They exhibit no differences in male geni-
talia of any significance and no consistent characters in
the female genitalia. There is great variability in the
number, arrangement, and shape of the spines and discs
forming the signa; but this is individual and as great
between females of a given series as between it and any
of the other named forms. Pending some information
on the biology, and more material from intervening
areas of the distribution, we shall have to keep some of
the names; but I suspect that they all represent but one
variable species. Figures 633c-d show extremes of
variation in the forked central projection from transtilla
between a small Texas example of hospitella and the type
of diffusella Ely. Such difference is not of more than
individual significance as we have both extremes and
all intergrades in male examples of hospitella.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
631. Eurythmia hospitella (Zeller)
Frioursrs 87, 633, 1130
Ephestia hospitella Zeller, Verh. zool.-bot. Ges. Wien, vol. 25,
p. 338, 1875.
Eurythmia hospitella (Zeller) Ragonot, N. Amer. Phycitidae, p.
16, 1887.—Hulst, Phycitidae of N. Amer., p. 196, 1900;
U. S. Nat. Mus. Bull. 52, p. 486, 1903—Hampson, in
Ragonot, Monograph, pt. 2, p. 266, 1901.—McDunnough,
Check list, No. 6385, 1939 (in part).
Eurythmia spaldingella Dyar, Proc. Ent. Soc. Washington, vol. 7,
p. 39, 1905.—McDunnough, Check list, No. 6391, 1939,
(New synonymy.)
Forewing white more or less dusted with fuscous, the
concentrated brown dusting appearing more or less
blackish to the naked eye; antemedial line transverse,
indicated by a dark outer border which is frequently
produced into a sharp, outward angle at top of cell;
subterminal line near and parallel to termen, bordered
inwardly by a dark border and outwardly by some dark
dusting; discal dots at end of cell small, blackish,
separate. Hind wing whitish, subpellucid; the veins
but faintly darkened; a fuscous line along termen.
Alar expanse, 10-15 mm.
Male genitalia as given for the genus: Female geni-
talia with the larger spines and discs, comprising the
signa, varying from 4 to 6. The type of hospitella has
5. One of the paratypes of spaldingella has 4, as in
yavapaella (fig. 1131). The other female paratype of
spaldingella has 5, as has a specimen from Winter Park,
Fla. A female of hospitella in the National Museum
from Texas (fig. 1180a) shows 6. There is no corre-
spondence between locality or pattern variation and the
number of the larger signa.
TYPE LOCALITIES: Texas (hospitella, in BM); Stock-
ton, Utah (spaldingella, in USNM).
Foop prant: Unknown.
Distrisution: Florida, Winter Park (July); Tezas,
Bosque County (May, July); Brownsville (May), Kerr-
ville (July); Arizona, Maricopa County, Fish Creek
(Sept.), Phoenix (Sept.), Liberty (Sept.); Utah, Stock-
ton (Aug., Sept.).
Our series (14 specimens). under hospitella in the
National Collection. contained two species, not dis-
tinguishable on superficial characters but radically dif-
ferent in male and female genitalia. Eight of these
specimens were the true hospitella. The remaining six
are quantulella Hulst which Ragonot (1889), Hulst
(1900), Hampson (1901), and later authors have treated
as a synonym of hospitella, but which, along with par-
vulella Ely and the tropical forms Dyar described under
Eurythmia, I am removing to form the new genus
Erelieva. ‘From Clarke’s notes I suspect that the orig-
inal type series of hospitella in the British Museum is
also a mixture. It consists of four females and two
males. As no one of the cotypes was designated by
Zeller or any later worker as the actual holotype I am
here designating the female of his type series dated
“17-V” as lectotype. It bears the following labels: A
small white label with the numerals “17/5” to indicate
307
date; another white, rectangular label with the inscrip-
tion ‘‘Bosque Co. Texas’’; the usual Zeller rectangular
green label inscribed “‘hospitella Z. Texas Stt. 68”; and
another white label with ‘‘Zell. Coll. 1884.” A sketch
of the genitalia of this lectotype shows five of the larger
signa in bursa (2 thorns and 8 discs) similar to the
arrangement shown in figure 1130.
632. Eurythmia hospitella yavapaella Dyar, new status
Figure 1131
Eurythmia yavapaella Dyar, Journ. New York Ent. Soc., vol. 14,
p. 108, 1906.—McDunnough, Check list, No. 6392, 1939.
Forewing more heavily dusted with dark scaling than
in typical spaldingella; general color, to the naked eye,
more brownish fuscous than ash gray and with the dark
borders of the transverse lines more or less obscured in
the dark overscaling of the wing; hind wing with a faint
smoky tint. Alar expanse, 13.5-15.5 mm,
Female genitalia of specimen from Glenwood Springs,
Colo. (fig. 1131) show four large signa. Another female
from the same locality and collection shows six of the
larger signa. A San Diego, Calif., female (fig. 1131a)
shows five.
Typr nocauity: Yavapai County, Ariz. (type in
USN).
Foop prant: Unknown.
DistrisutTion: Arizona, Yavapai County; Colorado,
Glenwood Springs; California, San Diego (May, July),
San Felipe Wash, San Diego County (June).
This is at most a race and probably no more than a
color variety of hospitella. I am holding the name
merely as a precaution against possible future separa-
tion.
633. Eurythmia angulella Ely
FieurE 1132
Eurythmia angulella Ely, Proc. Ent. Soc. Washington, vol. 12,
p- 202, 1910.—McDunnough, Check list, No. 6387, 1939.
Eurythmia diffusella Ely, Proc. Ent. Soc. Washington, vol. 12,
p. 202, 1910.—McDunnough, Check list, No. 6388, 1939.
(New synonymy.)
Similar to the dark western form, yavapaella, except
dark margins of transverse lines of forewing somewhat
more contrasted and hind wings darker. Bursa copu-
latrix of female also larger than that of any western
specimens. The name angulella may apply only to a
northeastern race of hospitella but in the absence of any
intergrading examples from any central United States
localities we must keep angulella specifically separated.
Ely’s diffusella, represented only by his male type, is
simply a slightly more brownish specimen of angulella
with slightly wider dark margin to the antemedial line.
Alar expanse, 12-14 mm.
Typm Ltocauity: East River, Conn. (angulella and
diffusella, in USNM).
Foop piant: Unknown.
Represented in the National Museum by five males
and nine females from the type locality (July) and one
male from Trenton, Ontario, Canada (July).
308
634. Hurythmia fumella Ely
Fieurn 1133
EHurythmia fumella Ely, Proc. Ent. Soc. Washington, vol. 12, p.
202, 1910.—McDunnough, Check list, No. 6389, 1939.
Probably nothing more than an aberrant form of
angulella; but known only from the unique female type,
and differing from angulella in the wider diffusion of
the whitish transverse lines of forewing, especially the
antemedial line and in the blackish suffusion over the
area between the lines. Bursa of female with but three
of the larger signa. Alar expanse, 14 mm.
Tyr Locauity: Hast River, Conn. (July, im USNM).
Foop PLant: Unknown.
191. Erelieva, new genus
Typs or Genus: Pempelia quantulella Hulst.
Characters of Hurythmia except: Male genitalia
without an apical process to gnathos, the arms of
gnathos fusing at their distal ends into the sclerotized
subanal plate (subscaphium); aedeagus not tapering,
expanding into a wide mouth at apex; penis unarmed
except for some sclerotized wrinklings; transtilla pro-
duced at middle into a stout, widely forked projection.
Female genitalia with bursa copulatrix evenly and finely
scobinate over entire inner surface; signa a single
elongate series of (5 to 14) thornlike spines; no cluster
of smaller spines at junction of bursa and ductus
bursae, except in parvulella where such spines are
arranged in a spiral extending into the ductus. Highth
abdominal segment of male with compound: dorso-
lateral scale tufts.
The species included in this new genus are removed
from Hurythmia because ‘of their radically different
genitalia. They are a tropical group with a couple of
extensions into the United States, while Hurythmia, as
now restricted, seems to be confined to the continental
United States. I suspect that, as in Hurythmia, the
several specific names (with the exception of parvulella)
represent no more than varieties of one widely dis-
tributed, superficially variable species; but this cannot
be determined from the specimens now available.
635. Erelieva quantulella (Hulet), new combination
Fieures 634, 1134
Pempelia quantulella Hulst, Ent. Amer., vol. 8, p. 134, 1887.
Eurythmia quantulella (Hulst) Ragonot, Ent. Amer., vol. 5, p.
116, 1889 (referred as synonym of E. hospitella).
Eurythmia santiagella Dyar, Ins. Insc. Menstr., vol. 7, p. 62,
1919. (New synonymy.)
Eurythmia hospitella Dyar (not Zeller), Ins. Insc. Menstr., vol.
13, p. 226, 1925.
Forewing fuscous more or less dusted with white,
giving the wing a pale gray cloor, usually paler and
more contrasting on the basal area; antemedial line
slightly oblique, bordered outwardly by blackish fus-
cous; subterminal line faint with a narrow, blackish,
inner border; discal dots faint, but distinct and separate.
Hind wing dusky white, with veins perceptibly dark-
UNITED STATES NATIONAL MUSEUM BULLETIN 207
ened and dark fuscous borders along costa and
outer margin. Alar’ expanse, 11-15 mm.
Male genitalia as given for the genus.
talia with from 10 to 14 signa.
TYPE LOCALITIES: Blanco County, Tex. (quantulella,
in AMNH, ex Rutgers); Santiago, Cuba (santiagella, in
USNM).
Foop puants: Opuntia (flowers), bell pepper, sor-
ghum. The larvae are evidently flower or seed feeders
in the plants attacked.
Disrrizution: Unirep States: Texas, Blanco
County (Aug.), Carrizo Springs (Oct.), Uvalde (June).
Purrto Rico: Aguirre Central (Apr.), Coamo Springs
(Apr.), Palmas Abajas (near Guayama, July), Puerto
Real (Vieques Isl., Apr., July), San, Germdn (Apr.,
Aug.). Virein Istanps: Kingshill. (St. Croix; Mar.,
Oct., Nov., Dec.), no specific locality (Apr.): Cua:
Santiago. Harr: Port au Prince (May).
The type of quantulella in the Rutgers Collection is
indeed a “‘very frail” specimen as Hulst said. It con-
sists of a head, thorax, and a fragment of one forewing
and is a female (not a male as stated by Hulst). A
similar female from Blanco County, Tex. (nm USNM), is
also labeled ‘‘type”’ by Hulst. The two specimens are
obviously conspecific. Dyar’s santiagella has nothing
to distinguish it from Texas specimens except the
slightly larger size of his female type (14 mm.). The
mele associated with it and also from Santiago is con-
siderably smaller (11.5 mm.). A long series of Cornell
specimens from Puerto Rico and the Virgin Islands
exhibits considerable variation in size and the distinct-
ness of the pale basal area of forewing and the extremes
of variation in the number of spines (signa) in the bursa.
There are no consistent genitalic or pattern differences
to distinguish even local races within the species, which
is apparently a tropical one that has extended its range
or been introduced into Texas.
The only Texas specimens I have seen are those in the
National Museum, three males and five females. Three
of the females came originally from Hulst through the
Brooklyn and Fernald Collections. One female, from
Uvalde, was reared June 1925 by A. P. Dodd from a
larva feeding in the flowers of an Opuntia. ‘The remain-
ing female and the three males were reared at Carrizo
Springs, Oct. 28, 1944, by members of the Special
Survey of the Division of Plant Quarantine of the U.S.
Bureau of Entomology and Plant Quarantine from lar-
vae feeding in the ripened pods of bell pepper. The
sorghum record is from reared examples in the National
Collection from St. Croix.
Female geni-
636. Erelieva coca (Dyar), new combination
Eurythmia coca Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 339, 1914.
Eurythmia coquilla Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 339,
1914 (mew synonymy).
Eurythmia mossa Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 340,
1914 (new synonymy).
Eurythmia uncta Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 340,
1914 (new synonymy).
Differs from quantulella only in its somewhat smaller
average size (10-12 mm.) and the more suffused brown-
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 309
ish color of forewing, the basal area being but slightly if
any paler than the remainder of the wing. The color
differences noted by Dyar among his supposed Panamé
species are not so great as the variation exhibited by the
Puerto Rican series of guantulella and the palpal differ-
ences he stresses are purely imaginary. The heads of his
specimens were so badly battered and the palpi so dis-
torted it would be impossible to determine their exact
lengths, shapes, or scaling even if there were such differ-
ences, which there are not. I have grave doubt that
coca is anything but a local form of quantulella; but in the
absence of any examples of Frelieva from the mainland
between Panama and Texas it seems best to retain coca
as a specific designation until more material is available.
The genitalia are not significantly different from those
of guantulella.
Typr Locaitius: Taboga Island, Panama (coca and
uncta, in USNM); La Chorrera, Panama (coguilla, in
USNM); Taboguilla Island, Panama (mossa, in USNM).
Foop piant: Unknown.
The only specimens IJ have seen are the 13 specimens
(a male and 12 females dated Feb. and May) originally
associated by Dyar with his several names and one
female from Corazal, Panama (Mar.), which he had
erroneously associated with his female type of “Hu-
rythmia vestilla,” the latter, itself, a synonymy of
Ephestiodes plorelia.
637. Erelieva parvulella (Ely), new combination
Ficure 1135
Eurythmia parvulella Ely, Proc. Ent. Soc. Washington, vol. 12,
p- 202, 1910.—McDunnough, Check list, No. 6386, 1939.
Forewing brown with a very sparse dusting of whitish
scales and a glossy sheen in contrast to the duller ap-
pearance of quantulella and coca; transverse lines
straight, narrow, the antemedial line more distinct than
in the other two species; veins 3 and 5 very shortly
stalked. Hind wing glossy fuscous; veins not appreci-
ably darker and terminal margin but faintly so. Alar
expanse, 11—-12.5 mm.
Male genitalia not appreciably different from those
of quantulella. Female genitalia with from 5 to 7 signa
and a spiral of small spines at junction of bursa and
ductus bursae and extending into the ductus.
Typ LocaLiry: East River, Conn. (type in USNM).
Foop prant: Unknown.
A distinct species, distinguished by its glossy brown
forewing, its darker hind wing, fewer signa, and the spiral
spining at junction of bursa and ductus bursae. It is
represented in the National Museum by a good series
(37 specimens) collected by Ely in July 1909 and 1910
at the type locality; but, so far as I know, has not turned
up since from any other locality. It may be indigenous
to the northeastern United States or an invader from
the Tropics. I suspect that it is the latter.
Group II
[Hind wing with veins 3 and 4 both absent.]}
Key to the genera of group III
1. Forewing with veins 2 and 3 united; 4 and 5stalked. ....... OSES be DUE ES
Forewing with veins 2 and 3stalked;4 and 5 united. . .
2. Hind wing with discocellular vein absent; 6 greatly Petuced
Hind wing with discocellular vein present; 6 normal .
192. Genus Cabnia Dyar
Cabnia Dyar, Journ. New York Ent. Soe., vol. 12, p. 108, 1904.—
Forbes, Cornell Mem. 68, p. 639, 1923. (Lype of genus:
Cabnia myronella Dyar.)
Tongue very short; but exposed between the well-
spaced labial palpi. Antenna pubescent; on male with a
broadly scaled basal sezsment anda short, deep incurva-
tion in base of shaft. Labial palpus upturned, short,
not reaching to vertex; second segment triangularly
scaled; third segment very short and broadly scaled.
Maxillary palpus minute, hardly distinguishable. Fore-
wing smooth; 9 veins; veins 2 and 3 stalked; 4 and 5
united, separated from the stalk of 2-3 at base; 6 from
below upper angle of cell, straight; 8 and 9 united; 10
from the cell, well separated from 8 at base; male with
a strong costal fold. Hind wing with vein 2 from well
before lower outer angle of cell; 5 from the angle; 7 and
8 completely anastomosed beyond cell; discocellular
vein very weak, curved. Highth abdominal segment
of male simple.
Male genitalia with apical process of gnathos a very
short, blunt hook. ‘Transtilla complete. Uncus long
(but not longer than tegumen), moderately broad,
beyond its broad base but slightly tapering to rounded
apical margin. Harpe elongate; costa produced at
apex into a short, free hook. Aedeagus simple, not
tapering, and not appreciably enlarged at apex. Penis
unarmed.
Female genitalia with ductus bursae membranous,
short (much shorter than bursa); bursa copulatrix
elongate, membranous, weakly scobinate over most of
its inner surface; signa present but very weak, consist-
ing of a pair of minute disks; genital opening simple;
ductus seminalis from anterior end of bursa.
On its abbreviated tongue Cabnia could go in the
Anerastiinae where it is placed in our recent list of
North American Lepidoptera. However, on male
genitalia and habitus it seems to fit better ito the
Phycitinae. The ocelli are present and the tongue not
310
Seas Cabnia (p. 310)
. Microphycita (p. 310)
Rabiria (p. 311)
entirely concealed by the palps. The genus contains
one North American species.
638. Cabnia myronella Dyar
Fiaures 128, 627, 1138
Cabnia myronella Dyar, Journ. New York Ent. Soc., vol.'12, p.
108, 1904.—Forbes, Cornell Mem. 68, p. 689, 1923.—
McDunnough, Check list, No. 6427, 1939.
Forewing dark cinereus, uniformly colored, many of
the scales tipped with dull white, giving the wing a
slightly frosted appearance; transverse lines obsolete,
or nearly so. Hind wing whitish, subpellucid; a very
faint dark line along termen. Alar expanse 10-11 mm.
Genitalic characters as given for the genus. The
male genitalia are remarkably like those of Anagasta
kiihniella, differing chiefly in the proportionally shorter
uncus and the presence of a complete transtilla.
TypE Locauity: Washington, D. C. (type in USNM).
Foop piant: Unknown.
Distripution: Massachusetts, Vinyard Haven (Aug.);
District of Columbia (June) ; Florida, Lake Placid (Mar.,
Apr.).
193. Genus Microphycita Dyar
Microphycita Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 346, 1914.
(Type of genus: Microphycita titillella Dyar.)
Tongue well developed. Antenna pubescent, simple.
Labial palpus upturned, slender; reaching to vertex;
third segment as long as second, acuminate. Maxillary
palpus minute, filiform. Forewing smooth; 9 veins;
veins 2 and 3 united; 4 and 5 stalked, the stalk separated
from 2 and 3 at base; 6 from below upper angle of cell,
straight; 9 absent; 10 stalked with 8; male with costal
fold. Hind wing with veins 2 and 5 rather short; 6
greatly reduced and obsolescent at base; 7 absent; cell
open (discocellular vein absent). Highth abdominal
segment of male simple.
Male genitalia without apical projection from gnathos,
the slender lateral arms fusing into sclerotized subanal
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 311
plate. Transtilla complete and greatly developed. Un-
cusabsent. Harpe with cucullus projecting into a blunt
point at apex. Anellus semitubular with elongate,
slender, lateral lobes. Aedeagus curved, slightly ta-
pered. Penis without armature. Vinculum as broad
as long.
Female genitalia with ductus bursae membranous,
slender, longer than bursa; bursa copulatrix round, mem-
branous; signum a finely, densely spined plate; genital
opening simple; ductus seminalis from middle of bursa.
A tropical genus of uncertain affinities. Dyar’s inter-
pretation of the forewing venation differs from mine.
He considered vein 4 absent and veins 3 and 5 stalked.
The end result would be the same, whichever interpreta-
tion is accepted.
639. Microphycita titillella Dyar
Figures 130, 636, 1137
Microphycita titillella Dyar, Proc. U.S. Nat. Mus., vol. 47, p. 346,
1914.
Forewing dull ocherous with a narrow dark (fuscous)
shade along costa and termen, expanding somewhat at
tornal area; transverse lines obscure; the antemedial line
near middle of wing, vertical and straight, indicated
chiefly by a narrow, fuscous, outer border; subterminal
line well back from terminal margin, also indicated by a
narrow, inner border, subparallel to antemedial line; no
distinguishable discal dots. Hind wing pale smoky fus-
cous. Alar expanse, 6-8 mm.
Genital characters as given for the genus.
Type tocauiry: Rio Trinidad, Panamé (type in
USNM).
Foop prant: Unknown.
Distrisution: PanamA: Cabima (May), Rio Trinidad
(Mar., May, June).
194, Rabiria, new genus
Typs or Genus: Microphycita conops Dyar
Characters of Microphycita except: Forewing with
vein 10 from the cell, separated from 8 at base. Hind
wing with veins of normal length; vein 2 from before
lower outer angle of cell; cell closed, discocellular vein
curved. Male genitalia with apical process of gnathos
well developed, U-shaped; uncus strongly developed
(similar to that of Cabnia); harpe simple, elongate, api-
cal margin evenly rounded; anellus a simple curved band,
without lateral lobes; aedeagus straight, not tapering;
transtilla a narrow, arched bridge. Female genitalia
with signum a small, smooth disk.
This genus is erected to receive conops originally in-
cluded in Microphycita by Dyar along with the type
species (titilella) of the latter genus. Why Dyar ever
included the two species under one generic heading—
even without examination of their genitalia—is difficult
to understand; for the hind wing venation of the two
is radically different.
640. Rabiria conops (Dyar), new combination
Figures 129, 635, 1136
me pag conops Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 346,
4,
Forewing dark fuscous with a faint bronzy sheen;
without transverse lines or discal spots. Hind wing
pale translucent fuscous. Alar expanse, 8 mm.
Genitalic characters as given for the genus.
Type tocaurry: La Chorrera, Panamd (type in
USNM).
Foop prant: Unknown.
DistRipution: PanaMA: Cabima (May), La Chorrera
(Apr., May) Rio Trinidad (May).
Represented so far only by the original type series,
three males and one female. The type is a male (not a
female as stated by Dyar). The genitalia are figured
from this specimen and from a female (May) from the
type locality.
Genera and species unplaced, unrecognized, or referred
from the Phycitinae
Unplaced
[Myelois] grossipunctella Ragonot
Figures 915, 916
Myelois grossipunctella Ragonot, Nouv. Gen., p. 8, 1888; Ent.
Amer., vol. 6, p. 64, 1890; Monograph, pt. 1, p. 51, 1893.—
McDunnough, Check list, No. 6069, 1939.
This species was described from a male in the Ober-
thiir Collection without locality label, but presumably
from Australia. There is a series in the U.S. National
Museum from California reared from larvae feeding on
the “cottony cushion scale’ and undoubtedly intro-
duced with that insect. This series was reared under
No. 869-09 by Coquillet, April 1889, and the species
has not been recovered since in California, so appar-
ently it did not become established there. It should
be dropped from our lists.
It is not a Myelois as the genitalia clearly indicate;
but its proper placement could only be made by one
thoroughly acquainted with the Australian Phycitidae.
Tyrer tocatiry: Australia [?] (location of type
unknown to me).
Foon: Icerya purchasi. Larva a predator.
[Myelois] famula Zeller
Figure 910
Myelois famula Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 218,
1881.
Myelois restrictella Zeller, Horae Soc. Ent. Rossicae, vol. 16,
p. 220, 1881.
Salebria famula (Zeller) Ragonot, Monograph, pt. 1, p. 365, 1893.
This species and its synonym (restrictella) are known
only from their female types. Their genitalia are
identical and suggest a placement much closer to
Fundella than to either Myelois or Salebria. It will
have to wait upon discovery of a male.
TYPE LOCALITIES: Barranquila, Colombia (famula,
in BM); Honda, Colombia (restrictella, in BM).
[Nephopteryx] fuscifrontella Zeller
Figure 911
Nephopteryz fuscifrontella Zeller, Horae Soc. Ent. Rossicae, vol.
16, p. 183, 1881.—Ragonot, Monograph, pt. 1, p, 282, 1893.
In the U.S. National Museum are two females from
Colombia identified by Ragonot as fuscifrontella. The
genitalia of one of these are figured. A male will be
needed for generic placement. Whatever it is, the
species is not a Nephopteryz.
Typ Locatiry: Honda, Colombia (type in BM).
312
{Salebria] nigricans Hulst
FicurE 918
Salebria nigricans Hulst, Canadian Ent., vol. 32, p. 171, 1900.—
MeDunnough, Check list, No. 6201, 1989.
I have seen nothing resembling the female type,
whose genitalia are figured. A male will be needed
for proper generic placement.
Typr Locauity: Phoenix, Ariz. (type n AMNH, ex
Rutgers).
[Hypochalcia] cervinistrigalis Walker
Ficure 919
Hypochalcia cervinistrigalis Walker, List, pt. 27, p. 45, 1863.—
Ragonot, Monograph, pt. 1, p. 585, 1893.
Photographs of the female type and its genitalia are
before me. The latter are figured. I have seen
nothing like them. A male will be needed for generic
placement.
TyYpE LocaLiry: Santo Domingo [Dominican Repub-
lic] (type in BM).
Hypochalcia hulstiella Ragonot
Hypochalcia hulstiella Ragonot, N. Amer. Phycitidae, p. 11, 1887;
Monograph, pt. 1, p. 603, 1893.—Hulst, Phycitidae of N.
nae p: 168, 1890.—MecDunnough, Check list, No. 6275,
1939.
According to Clarke the type lacks an abdomen and
is otherwise in very poor condition. It is very likely
not an American example. Ragonot’s figure (Mono-
graph, pl. 21, fig. 5) strongly resembles one of the vari-
ants of the European Hypochalcia ahenella with well-
marked lines.
TyprE Locanitry: ‘Texas’? (type in Paris Mus.).
[Elasmopalpus] corrientellus Ragonot
Fieure 917
Elasmopalpus corrientellus Ragonot, Nouv. Gen., p. 23, 1888;
Monograph, pt. 1, p. 423, 1893.
This is not an Hlasmopalpus, but generic placement
cannot be made without an authentic male. The type
is a female. Its genitalia are figured.
Typr LocaLity: Corrientes, Argentina (type in Paris
Mus.).
[Euzophera] postflavida Dyar
Fiaure 1128
Euzophera postflavida Dyar, Ins. Inse. Menstr., vol. 11, p. 29,
1928.
This species is represented by the female type and a
female paratype (in USNM) from St. Laurent Maroni,
UNITED STATES NATIONAL MUSEUM BULLETIN 207
French Guiana, and a female (in Cornell Collection)
from Tumatumari, Potaro River, British Guiana (June).
The male is unknown. The distinctive genetalia and
coloration readily identify the species. The basal half
of hind wing and the abdomen (except for the last two
segments) are ochre yellow; apical half of hind wing and
the two caudal segments of abdomen blackish. A male,
however, will be needed before the species can be
properly placed generically. It is not a Huzophera.
Typr tocatity: Nouveau Chantier, French Guiana
(Sept.; type in USNM).
Foop prant: Unknown.
[Euzophera] rinmea Dyar
Figure 1129
Euzophera rinmea Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 335,
1914.
Known only from the female type.
place without male.
Typr Locauity: Rio Trinidad, Panama (Mar.; type in
USNM).
Foop piant: Unknown.
Impossible to
[Euzophera] came Dyar
Figure 1126
Euzophera came Dyar, Ins. Inse. Menstr., vol. 7, p. 56, 1919.
Represented in the National Collection by the female
type and five other females from the type locality.
Not a Euzophera but cannot be placed without male.
TyYpx Locauity: Cayuga, Guatemala (type in USNM).
Foop prant: Unknown.
[Euzophera] gais Dyar
Euzophera gais Dyar, Ins. Insc. Menstr., vol. 7, p. 57, 1919.
Represented only by the unique female type, which is
probably a synonym of Anthropteryx irichampa Dyar.
Both species have similar genitalia. Synonymy and
proper placement will have to wait upon recovery of a
male.
TypE LocaLity: Cayuga, Guatemala (June; type in
USNM).
Foop prant: Unknown.
Genus Anthropteryx Dyar
FiaurE 1059
Anthropteryx Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 335, 1914.
Nels of genus: Anthropteryx irichampa Dyar, loc. cit., p.
336.
This genus is probably a good one, but was erected
upon a freak specimen and the venational ‘‘characters”’
as given are incorrect. Vein 4 is absent from the left
forewing of irichampa but present on the right wing and
long stalked with 5, and vein 3 is connate with the stalk
of 4-5, not stalked as stated by Dyar. The species
belongs, therefore, in venational group A (11 veins in
forewing and vein 2 from the cell). Its genitalia indi-
cate close relationship to Moodnopsis. A male will be
needed before the status of Anthropteryx can be deter-
300329—56——21
313
mined. The type of genus is represented only by its
unique female type.
TyrpELocauity: Taboga Island, Panama (Feb.; type in
USNM).
Foop pirant: Unknown.
[Moodna] formulella Schaus
Figure 1127
Moodna formuleila Schaus, Ann. Mag. Nat. Hist. ser. 8, vol. 11,
p. 252, 1913.
A large (28 mm.), strikingly marked species which
should be easy to recognize if more specimens are dis-
covered. So far known only from the unique female
type. A male will be needed for proper placement. It
is obviously not a Moodna, having 11 veins in the fore-
wing, with 4 and 5 stalked for a little more than half
their lengths; 2 from before the lower outer angle of cell;
3 from the angle and slightly separated from the stalk
of 4-5; 6 straight from below upper angle of cell; 8 and 9
long stalked; 10 from the cell but rather close to the
stalk of 8-9. Hind wing with 2 from well before the
angle of the cell; 3 and 5 from the angle, approximate at
base; 7 and 8 anastomiosed from most of their lengths
beyond cell; cell one-third the length of wing; discocellu-
lar vein curved. The transverse lines of forewing are
rather close together, the antemedial line being near the
middle of the wing. The dark areas and markings of
the wing are a reddish brown rather than the usual
fuscous shades.
Typ Locauity: Juan Vifias, Costa Rica (Feb.; type
in USNM).
Foop prant: Unknown.
[Eucampyla] putidella Schaus
Fieure 1095
Eucampyla putidella Schaus, Ann. Mag. Nat. Hist., ser. 8, vol.
11, p. 250, 1913.
Known only from the female type. A male is needed
for proper placement. Certainly does not belong in
Eucampyla, which is an Australian genus not represented
in the American faunas. May be an aberrant exam-
ple of Vitula. The venation and female genitalia are
those of Vitula except for the presence of vein 9 in both
forewings (8 and 9 being long stalked) and the absence
of scobinations in the bursa.
TypE Locatiry: Mount Poas, Costa Rica (May; type
in USNM).
Foop puant: Unknown.
[Honora] dulciella Hulst
Figure 951
Honora dulciella Hulst, Journ. New York Ent. Soc.. vol. 8, p. 223,
1900.—McDunnough, Check list, No. 6349, 1939.
This species has the venation of Honora but the fe-
male genitalia are altogether wrong for that genus. A
male will be needed for proper placement. At present
dulciella is represented only by the female type.
Typx Locatity: Palm Beach, Fla. (type in USNM).
Foop prant: Unknown.
314
Unrecognized
Myelois infusella Zeller
Myelois infusella Zeller, Isis von Oken, 1848, p. 869.
Salebria infusella (Zeller) Ragonot, Monograph, pt. 1, p. 352,
1893.
Unknown to me. The type is supposed to be in the
Berlin Museum, but Dr. Hering was unable to locate it.
TYPE LocaLiry: Port-au-Prince, Haiti.
Genus Phycitopsis Ragonot
Phycitopsis Ragonot, N. Amer. Phycitidae, p. 4, 1887; Mono-
graph, pt. 1, p. 185, 1893.—Hulst, Phycitidae of N. Amer.,
p. 1388, 1890. (Type of genus: Phycitopsis flavicornella
Ragonot, loc. cit.—Hulst, op. cit., p. 134—MeDunnough,
Check list, No. 6124, 1939.)
T have seen nothing from America that could be this
genus or species. According to Clarke’s notes the fe-
male type is not in either the Paris or British Museum
Collections. From the Ragonot descriptions the genus
should go into venational division D and the species
must strongly resemble the Old World Phycita spissi-
cella (Fabricus). I suspect that flavicornella is a Kuro-
pean specimen and the ‘“Texas’’ locality is 2 mislabeling.
This is quite likely if the type was a Boll specimen.
There is a discrepancy between Ragonot’s description
and his figure of the labial palpus (Monograph, pl. 8,
fig. 10). The figure shows an oblique, moderately long
(not @ very short) ascending palp.
Tyerr Locauity: ‘“Texas’’ (type lost?).
Euzophera intextella (Zeller)
Myelois intextella Zeller, Isis von Oken, 1848, p. 871.
Euzophera intextella (Zeller) Ragonot, Monograph, pt. 2, p. 53,
1901.
Unknown to me. The type is supposed to be in the
Berlin Museum but Dr. Hering has been unable to
locate it.
Typrr Locatity: St. Thomas, British West Indies.
Euzophera daedalella Ragonot
Euzophera daedalella Ragonot, Nouv. Gen. p. 32, 1888; Mono-
graph, pt. 2, p. 56, 1910.
Unknown to me. This type is also supposed to be in
the Berlin Museum. Dr. Hering was unable to locate
it. :
Typr Locatity: Chanchamayo, Peri.
Zophodia brevistrigella Ragonot
Zophodia brevistrigella Ragonot, Nouv. Gen., p. 31, 1888; Mono-
graph, pt. 2, p. 24, 1901.
According to Ragonot the type (2) was deposited in
the Staudinger Collection in Berlin. It has not been
located and so far I have seen nothing that agrees with
the description and figure published in the Ragonot
monograph.
TypxE Locauity: Nova Friburgo, Brazil.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Zophodia subcanella (Zeller)
Myelois subcanella Zeller, Isis von Oken, 1848, p. 873.
Zophodia subcanella (Zeller) Ragonot, Monograph, pt. 2, p. 22,
1901.
Unknown to me. Type supposed to be in Berlin
Museum but not located.
TYPE LocaLity: Brazil.
Psorosa disticta (Zeller)
Euzophera disticta Zeller, Horae Soc. Ent. Rossicae, vol. 16, p.
232, 1881.
Psorosa disticta (Zeller) Ragonot, Monograph, pt. 2, p. 107, 1901.
A photograph of the genitalia of the female type
shows two large opposed signa composed of cup-
shaped, strongly spined concave plates (similar to those
of Rotruda mucidella). I have seen nothing that
matches the type exactly either in structure or super-
ficially. It is obviously not a Psorosa or a Huzophera;
but generic placement cannot be made without a male.
Tyrr Locauity: Maraquita, Colombia (type in BM).
Hornigia clitellatella Ragonot
Hornigia clitellatella Ragonot, Nouv. Gen., p. 36, 1888.
In the Ragonot monograph (pt. 2, p. 269, 1901)
Hampson places clitellatella in Moodna. It apparently
belongs somewhere near Moodna, in venational group
C, but accurate placement must wait upon discovery of
a male from Chile. The female type is in the Paris
Museum. I have seen nothing agreeing with Ragonot’s
description or figure (Monograph, pl. 34, fig. 12).
Typr Locatity: Callao, Chile (type in Paris Mus.).
Moodna olivella Hampson
Moodna olivella Hampson, in Ragonot, Monograph, pt. 2, p. 268,
1901.
I have seen nothing agreeing with either the descrip-
tion or figure (pl. 49, fig. 12) in the Monograph. Clarke’s
notes on the genitalia of the unique male type in the
British Museum show that olivella cannot be a Moodna.
The gnathos apparently is more like that of Moodnopsis.
Placement will have to wait upon recovery of additional
specimens.
TypsE LocaLiry: Petrépolis, Brazil.
Euzopherodes megalopalis Hampson
Euzopherodes megalopalis Hampson, Ann. Mag. Nat. Hist., ser. 7,
vol. 14, p. 181, 1904.
According to Clarke’s notes the type im the Bzitish
Museum is a female (not a male as stated by Hampson)
and has a glued-on abdomen, which may or may not
belong with the remainder of the specimen. <A photo-
graph of the type shows a moth superficially similar to
Nicetiodes apianella Schaus in markings. Its venation
(if correctly given) places it in venational group D
where we have nothing resembling it. We shall have
to wait upon additional specimens (male and female)
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 815
for proper placement. Hampson’s reference to the
Old World Euzopherodes is obviously wrong.
Type Locauity: Nassau, Bahamas.
Genera and species referred from Phycitinae
Genus Eutrichocera Hampson
Eutrichocera Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 14, p.
182, 1904. (Type of genus: Eutrichocera paurolepidalis
Hampson, loc. cit.)
This is an Epipaschiid. The genus is a synonym of
Pococera and its type species a synonym of P. insularella
Ragonot.
Tyrr Locatity: Abaco, Bahamas (type, o’, in BM).
Myelois atristrigella Ragonot
Myelois atristrigella Ragonot, Monograph, pt. 1, p. 40, pl. 18,
fig. 3, 1893.
This species goes to the Hpipaschiidae. I have
examined the type. It has the normal epipaschiid
venation, but will require a new genus, for it does not
fit comfortably in any of our described epipaschiid
genera. Its peculiar forewing pattern should make it
easy to identify if more specimens are recovered. At
present it is known only from the unique male type.
Typr Locatity: Puerto Rico (type in Zool. Mus.
Univ. Berlin).
Genus Psammia Hampson
Psammia Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 71,
1930. (Type of genus: Psammia flavipicta Hampson, loc.
cit—McDunnough, Check list, No. 6234, 1939.)
Genus and species go to Anerastiinae.
TypE Locauity: Florida (type in BM).
Megasis indianella Dyar
Megasis indianella Dyar, Ins. Inse. Menstr., vol. 11, p. 28, 1923.—
MeDunnough, Check list, No. 6265, 1939.
Goes to Anerastiinae. Dyar’s species is a synonym
of Ragonotia olivella Hulst.
TypE Locauity: Indian Wells, Calif. (type in USNM).
Euzophera tintilla Dyar
Euzophera tintilla Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 334,
1914.
Goes to Anerastiinae. Described from single male.
Generic placement uncertain.
Typr tocauiry: Porto Bello, Panama (Apr.; type in
USNM).
Foop: Unknown. Larva probably a coccid feeder.
Euzophera conquistador Dyar
Euzophera conquistador Dyar, Proc. U. 8. Nat. Mus., vol. 47,
p. 335, 1914.
Goes to Anerastiinae. Described from single female.
Generic placement uncertain.
TyprrE LocaLity: Cabima, Panamé& (May; type in
USNM).
Foop: Unknown.
Euzophera mabes Dyar
Euzophera mabes Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 335,
1914.
Goes to Anerastiinae. Described from female. Ge-
neric placement uncertain.
Types Locauity: Taboga Island, Panam (Feb.; type
in USNM).
Foop: Unknown.
Euzophera climosa Dyar
Euzophera climosa Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 335,
1914,
The original series before Dyar consisted of three
males and four females (not one male and five females as
stated) and only the male holotype and one male from
the type locality can be definitely assigned to the name.
The other male from the type locality is congeneric but
a distinct, undescribed species. The four females repre-
sent two species, each of which consists of one example
from Rio Trinidad and one from Porto Bello. Which
of these female species, if either, represents the other
sex of climosa is not possible to determine. All these
specimens belong in the Anerastiinae and will require
reference to a new genus when that subfamily is revised.
Typs Locauity: Rio Trinidad, Panamé' (Mar.; type
in USNM).
Foopv: Unknown. Probably a coccid feeder.
Zophodia epischnioides Hulst
Zophodia epischnioides Hulst, Canadian Ent. vol. 32, p. 173, 1900.
I have been unable to locate the type of this species in
the Rutgers Collection or elsewhere. From the descrip-
tion I suspect that it is an Anerastiine. It is obviously
not a Zophodia. Disposition of the name can wait upon
a revision of the Anerastiinae.
Typr Locatity: Not given (type lost?).
Foop prant: Unknown.
Genus Harnochina Dyar
Harnochina Dyar, Proc. U. 8. Nat. Mus., vol. 47, p. 340, 1914.
(Type of genus: Harnochina rectilinea Dyar, loc. cit.)
Goes to the Anerastiinae. The tongue is absent.
Typr Locauity: Corozal, Canal Zone, Panamé (Mar.;
type in USNM).
Foop piant: Unknown.
Genus Nasutes Hampson
Nasutes Hampson, Ann. Mag. Nat. Hist., ser. 10, vol. 5, p. 53,
1930. (Type of genus: Nasutes venata Hampson, op. cit. p.
54.)
This genus is a synonym of Bandera Ragonot and its
type species (venata) a synonym of B. cupidinella
(Hulst).
316 UNITED STATES NATIONAL MUSEUM BULLETIN 207
I have before me an enlarged photograph of the geni-
talia of the male type of venata and a slide of the genitalia
of the male type of cupidinella; the two, both from Colo-
rado, are identical in all details. The genus Bandera is
a true anerastiid on habitus and all characters except
the tongue. This organ is reduced, but not concealed
between the labial palpi as in typical Anerastiinae.
However, Bandera lacks any trace of an ocellus, as do
several other Anerastiinae. This organ is present in all
genera of Phycitinae in the Americas.
Typr Locauity: Colorado (type in BM).
Foop prant: Unknown.
Maricopa albocosiella Hulst
Maricopa albocostella Hulst, Canadian Ent., vol. 32, p. 176, 1900.
Valdiria albocostella (Hulst) Barnes and McDunnough, Check
list of the Lepidoptera of Boreal America, No. 5755.—
McecDunnough, Check list, No. 6354, 1939.
Not a Valdivia. Goes to Anerastiinae. Has no
tongue visible. The type is a male with veins 4 and 5
united in forewing and with a slight excavation in the
base of the antennal shaft.
TypE LtocaLity: Anglesea, N. J. (type in AMNH, ex
Rutgers).
Foop piant: Unknown.
Checklist of American Phycitinae
1. CryprosLaBEs Zeller
ile
gnidiella (Milliére): Europe, Africa, Asia,
Bermuda, Venezuela, Brazil
2. Acrosasis Zeller
ore
. Indigenella (Zeller):
. feltella Dyar:
. palliolella Ragonot:
. caryalbella Ely:
. juglandis (LeBaron): Eastern U. S.
. sylviella Ely: Eastern U. S., Canada
. kearfottella Dyar:
. caryae Grote:
. evanescentella Dyar:
. stigmella Dyar:
. aurorella Ely: Eastern U.S.
. peplifera Dyar:
. exsulella (Zeller), new comb.:
. angusella Grote:
. demotella Grote:
. latifasciella Dyar:
. irrubriella Ely: Eastern U.S.
. normella Dyar:
. Malipennella Dyar: Eastern U. S. (Con-
. cirroferella, Hulst:
. cunulae Dyar and Heinrich: Eastern U. S.
. caryivorella Ragonot:
Mineola Hulst
Seneca Hulst
Acrocaula Hulst
Eastern U.S. and Can-
ada, California
nebulo (Walsh)
nebulella (Riley)
zelatella (Hulst)
. grossbecki (Barnes and McDunnough), new
comb.: Florida
. vaccinii Riley: U.S.
. amplexella Ragonot: Eastern U.S.
. tricolorella Grote: U.S., Canada
scitulella Hulst
. comptella Ragonot: Western U.S.
. minimella Ragonot: Eastern U.S.
nigrosignella Hulst
Eastern U. S., Canada
Eastern U. S., Canada
albocapitella Hulst
U.S. (Connecticut)
Eastern U. S.
Eastern U. S., Canada
U.S. (Florida)
Eastern U.S.
Eastern U. S.
Eastern U. 8S.
septentrionella Dyar
Eastern U. S., Canada
eliella Dyar
Eastern U. S.
Eastern U.S.
Eastern U.S. (Connecticut)
necticut)
. dyarella Ely: Eastern U.S. (Connecticut)
. ostryella Ely: Eastern U.S., Canada
. secundella Ely: Eastern U.S., Canada
. coryliella Dyar: Eastern U.S.
. hebescella Hulst:
Eastern U. S. (New Jer-
sey)
Eastern U.S. (Texas)
Eastern and South-
western U.S.
“I
35. comacornella (Hulst), new comb.: Eastern
U.S. (Texas)
36. betulella Hulst: Eastern and Western U.S.,
Canada
37. rubrifasciella Packard: Eastern U. S.,
Canada
alnella McDunnough
38. comptoniella Hulst: Eastern U. S., Canada
39. myricella Barnes and McDunnough: U. S.
(Florida)
40. tumidulella (Ragonot), new comb.:
(Florida)
USS:
. ReopopHakEa Guénée
41. caliginella (Hulst), new comb.:
fornia, Arizona)
caliginoidella (Dyar)
42. supposita (Heinrich), new comb.:
(British Columbia)
U.S. (Cali-
Canada
. TracHycrra Ragonot
43. pallicornella (Ragonot): U.S. (Texas)
. ANABASIS Heinrich, new genus
44. ochrodesma (Zeller), new comb.: U. S.,
(Florida), México, Guatemala, Panama
Colombia, West Indies
crassisquamella (Hampson)
. Mitpreixra Dyar
45. constitutionella Dyar: México, Guatemala
. SEMATONEURA Ragonot
46. atrovenosella Ragonot: México, Costa Rica,
Colombia, Ecuador, Pert, Argentina
47. abitus Heinrich, new species: Ecuador
. HypsrpyLa Ragonot
48. grandella (Zeller): U. S. (Florida), West
Indies and Tropical America to Argen-
tina
cnabella Dyar
49. ferrealis (Hampson), new comb.: ‘Tropical
America (Costa Rica to Brazil)
50. dorsimacula (Schaus), new comb.: Costa
Rica
51. fluviatella Schaus: Costa Rica
. Hemipritocera Ragonot
52. chinographella Ragonot: French Guiana,
Brazil, Pera
53. bigrana (Zeller): México, Colombia
54, plumigerella (Ragonot), new comb.: ‘Amer.
Merid.”
55. letharda (Schaus), new comb.: Panamé,
México
56. jocarella (Schaus): Costa Rica, Panamé,
Brazil
57. exoleta (Zeller): Colombia
317
318
10.
11.
12.
13.
14.
15.
16.
17.
18.
i).
20.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Crocipomera Zeller
58. turbidella Zeller:
U.S. (Texas)
59. fissuralis (Walker):
Puerto Rico
adonea (Felder and Rogenhofer)
60. stenopteryx (Dyar), new comb.: México
CunisEertTa Heinrich, new genus
61. subtinctella (Ragonot), new comb.:
ern U.S. and Canada
Heras Heinrich, new genus
62. disjunctus Heinrich, new species:
Apanarsa Heinrich, new genus
63. intransitella (Dyar), new comb.:
(Arizona, New Mexico)
Birinus Heinrich, new genus
64. russeolus Heinrich, new species:
Guiana
BeErreLia Barnes and McDunnough
65. grisella Barnes and McDunnough: U. 5.
(Arizona)
Hypareyria Ragonot
Cuba, Jamaica, México,
Dominican Republic,
West-
Colombia
WES:
British
66. definitella (Zeller): Puerto Rico, Virgin
Islands, Colombia, Brazil
67. slossonella (Hulst), new comb.: U. S.
(Florida), México
tenuella (Barnes and McDunnough)
CuaAraAnrica Heinrich, new genus
68. annuliferella (Dyar), new comb.: U. S.
(New Mexico, Arizona)
69. hystriculella (Hulst), new comb.: U. S.
(Texas, Florida)
70. bicolorella (Barnes and McDunnough), new
comb.: U.S. (Arizona, Nevada, Cali-
fornia)
Myetopstis Heinrich, new genus
71. coniella (Ragonot), new comb.:
ada, México
nefas (Dyar)
72. immundella (Hulst),
(Texas)
73. subtetricella (Ragonot), new comb.:
Canada
zonulella (Ragonot)
obnupsella (Hulst)
U.S., Can-
new comb.: U. S.
wese
74. minutularia (Hulst), new comb.: U. S.
(Texas)
75. alatella (Hulst), new comb.: Western U.S.
rectistrigella (Ragonot)
fragilella (Dyar)
piazzella (Dyar)
AnypsIpyLa Dyar
76. univitella Dyar: Cuba, México, Guatemala,
Panamé, Venezuela, Brazil, Peri, Heua-
dor, Jamaica
ApomyYrEtois Heinrich, new genus
77. bistriatella (Hulst), new comb.:
U.S., Canada
bilineatella (Ragonot)
Eastern
21. Ecromyxrtots Heinrich, new genus.
22.
23.
24.
25.
78. decolor (Zeller), new comb.: ‘Tropical
America,
ephestiella (Hampson)
79. ceratoniae (Zeller), new comb.: Hurope,
U. S. (@lorida), Puerto Rico, Jamaica,
Argentina
oporedestella (Dyar)
80. muriscis (Dyar), new comb.: ‘Tropical
America
palpalis (Dyar)
81. furvidorsella (Ragonot), new comb.: Puerto
Rico
82. zeteki Heinrich, new species: Panama
PaRAMYELot!s Heinrich, new genus.
83. transitella (Walker), new comb.: U. S.,
tropical America
notatalis (Walker)
solitella (Zeller)
duplipunctella (Ragonot)
venipars (Dyar)
cassiae (Dyar)
Psrupopivona Dyar
84. commensella Dyar: México
85. cispha Dyar: Guatemala, Costa Rica, Brit-
ish Honduras
86. santa-maria Dyar: Guatemala
87. carabayella Dyar: Pert, Bolivia, Colombia
PROTOMOERBES Heinrich, new genus
88. aberrans Heinrich, new species:
89. separabilis Heinrich, new species:
Dratomocrera Ragonot
Cabima Dyar
90. tenebricosa (Zeller):
ana, Costa Rica
91. dosia (Dyar), new comb.: Panamé
92. excisalis (Hampson), new comb.:
Guiana, Bolivia (?)
93. decurrens (Dyar), new comb.:
94. majuscula Heinrich, new species:
95. albosigno Heinrich, new species:
96. hoplidice (Dyar), new comb.:
97. extracta Heinrich, new species:
98. mochlophleps (Dyar), new comb.:
Colombia
Colombia
Colombia, French Gui-
French
Panamaé
Brazil
Brazil
Panama
Costa Rica
México
. Psrupocasima Heinrich, new genus
99. castronalis Heinrich, new species: Brazil
100. fearnella (Schaus), new comb.: Costa Rica,
Guatemala
101. guianalis Heinrich, new species:
Guiana, British Guiana
euzopherella (Dyar), new comb.: Panama
Panama
French
102.
103. pombra (Dyar), new comb.:
104. nigristrigella (Ragonot), new comb.: Brazil
105. arizonensis Heinrich, new species: U. S.
(Arizona)
106. expunctrix (Dyar and Heinrich), new comb.:
Brazil
107. perrensiella (Ragonot, new comb.: Argen-
tina
27.
28.
29.
30.
31.
32.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
108. rubrizonalis (Hampson), new comb.: French
Guiana, Brazil
Hyatospita Ragonot
109. stictoneurella Ragonot: México, Guatemala,
Brazil
110. celiella Schaus: Costa Rica
111. insequens Heinrich, new species: Bolivia,
Colombia
112. majorina Heinrich, new species: México
113. fulgidula Heinrich, new species: Cuba
114. egenella (Ragonot), new comb.: Brazil
115. xanthoudemia (Dyar), new comb.: Panama,
Costa Rica
116. angulinella (Schaus), new comb.: Costa
Rica
117. clevelandella (Dyar): Panam&
118. semibrunneella Ragonot: Colombia
Funpeuia Zeller
119. pellucens Zeller: U.S. (Florida), West In-
dies, Brazil, Bolivia
cistipennis (Dyar)
argentina Dyar: U. S. (Florida, Texas),
West Indies, Venezuela, Brazil, Argen-
tina
eucasis Dyar
121. agapella Schaus: Galapagos Islands
122. ignobilis Heinrich: México, Guatemala,
Costa Rica, Cuba, Puerto Rico, Haiti
ahemora Dyar: México, Guatemala, Costa
Rica
DirunpDELia Dyar
124. corynophora Dyar:
French Guiana
125. subsutella (Schaus), new comb.:
120.
123.
Guatemala, Panamé,
Costa Rica
126. distractor Heinrich, new species: Puerto
Rico
127. tolerata Heinrich, new species: Bolivia
CopTarTHRIA Ragonot
128. dasypyga (Zeller):
PromMyYLEA Ragonot
129. lunigerella Ragonot:
Canada.
130. lunigerella glendella (Dyar):
131. dyari Heinrich, new name:
zimmermani (Druce)
drucei (Dyar)
132. druceii (Ragonot), new comb.:
133. mindosis Dyar: México
134. dasystigma Dyar: México
ANADELOSEMIA Dyar
135. senesciella (Schaus): Costa Rica
136. tecmessella (Schaus): Costa Rica
137. fifria Dyar: Guatemala
138. base Dyar: Guatemala
139. obstitella (Schaus), new comb.: Costa Rica
140. texanella (Hulst), new comb.: U.S. (Texas,
Florida), Puerto Rico, Cuba
dulciella (Hulst)
condigna Heinrich, new species:
zona)
Colombia, Guatemala
Western U. S. and
Colorado
México
Guatemala
141. U.S. (Ari-
33.
34.
35.
36.
37.
38.
39.
319
Dasypyea Ragonot
142. alternosquamella Ragonot:
Canada
stictophorella Ragonot
Rampyuua Dyar
143. orio Dyar: México
144. polydectella (Schaus): Costa Rica
145. subcaudata (Dyar), newcomb.: Guatemala,
Costa Rica, Brazil
146. lophotalis Heinrich, new species:
Guatemala
Fuurapa Heinrich, new genus
147. querna (Dyar), new comb.: Panamé
148. carpasella (Schaus), new comb.: Galdpagos
Islands
Scoryuus Heinrich, new genus
149. cubensis Heinrich, new species:
Davara Walker
Homalopalpia Dyar
Eucardinia Dyar
150. caricae (Dyar), new comb.:
Tropical America
dalera (Dyar)
Western U. S.,
México,
Cuba
U.S. (Florida),
151. columnella (Zeller), new comb.: Colombia
152. nerthella (Schaus), new comb.: México,
Guatemala, Costa Rica
euthales (Dyar)
153. paranensis (Dyar), new comb.: Brazil
154. azonaxsalis Walker: Brazil
155. interjecta Heinrich, new species: Puerto
Rico, Dominican Republic
156. rufulella (Ragonot), newcomb.: Puerto Rico
Sarasora Hulst
Cuba Dyar
157. plumigerella Hulst: U.S. (Florida)
158. furculella (Dyar), new comb.: Cuba, Puerto
Rico, Dominica, Virgin Islands
159. ptyonopoda (Hampson), new comb.:
ward Islands
Presmopopa Zeller
Discopalpia Ragonot
Amphycitopsis Dyar
160. rubicundella Zeller: Brazil
161. xanthomera Dyar: Guatemala,
Costa Rica, French Guiana
zanthozona Dyar
trichomata (Zeller): Colombia
flavicans (Zeller): Colombia, French Guiana
fratella Dyar
ragonoti (Dyar), new comb.:
temala, Costa Rica
isabella (Dyar), new comb.:
xanthopolys Dyar: Panama
parva Heinrich, new species: Panama
semirufella (Zeller): Colombia
apocerastes Dyar: México, Costa Rica,
French Guiana, Brazil, Dominica
montella Schaus: Costa Rica
Wind-
Panam 4
162.
163.
164. México, Gua-
165. Costa Rica
166.
167.
168.
169.
170.
320
40.
41.
42.
43.
44,
45.
46.
47.
48.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Atuertoca Heinrich, new genus
171. subrufella (Hulst), new comb.:
ida), Cuba, Virgin Islands
jiliolella (Aulst). Virgin Islands
ptychis (Dyar)
U.S. (Flor-
172. bondari Heinrich, new species: Brazil
PRAEDONULA Heinrich, new genus
173. almonella (Dyar), new comb.: Panama
Prapus Heinrich, new genus
174. burdettellus (Schaus), new comb.:
Rica, Guatemala
semproniella (Schaus)
175. dissitus Heinrich, new species: Brazil
176. subaquilellus (Ragonot), new comb.: Guate-
mala,
Gasinivs Heinrich, new genus
177. paulsoni (Ragonot), new comb.:
Crracantuia Ragonot
Procandiopa Dyar
178. mamella (Dyar),
Guatemala
179. vepreculella Ragonot:
Merearraria Ragonot
180. peterseni (Zeller):
Brazil, Pert
Costa
Chile
new comb.: Panama,
Ecuador
Guatemala, Colombia,
181. squamifera Heinrich, new species: Costa
Rica
182. frustrator Heinrich, new species: Costa Rica
183. schausi Heinrich, new species: Costa Rica
184. cervicalis Dyar: Cuba
185. alpha Heimrich, new species: Guatemala,
México, Costa Rica, Panam4, Bolivia,
Brazil
beta Heinrich, new species: México, Guate-
mala, Costa Rica, Trinidad
Drescoma Dyar
187. cyrdipsa Dyar: México, Guatemala, Pan-
ama, French Guiana
188. cinilixa Dyar: Guatemala, Panamé
186.
Mownopritora Hulst
189. pergratialis (Hulst): U.S.
grotella (Ragonot)
nubilella Hulst
ZAMAGIRIA Dyar
190. dixolophella Dyar: Panama
191. pogerythrus Dyar: México, Guatemala
192. hospitabilis Dyar: Cuba
193. masculinus Dyar: Guatemala
194. australella (Hulst), new comb.: Weis:
(Texas, Florida)
bumeliella (Barnes and McDunnough):
U.S. (Texas, Florida)
fraterna Heinrich, new species:
laidion (Zeller):
America
deva Dyar
striella Dyar
ipsetona Dyar:
195.
196.
Cuba
U. S. (Florida), Tropical
197. Costa Rica
49. ANEGCEPHALESIS Dyar
198. arctella (Ragonot), new comb.: U. S.
(Florida), Bahamas, Cuba
cathaeretes Dyar
50. Maeiriopsis Heinrich, new genus
199. denticosella (Dyar), new comb.: Tropical
51.
52.
53.
54.
55.
America
cristalis (Hampson)
ANcYLOSToMIA Ragonot
200. stercorea (Zeller):
America
agnobilis (Butler)
diffissella (Zeller)
201. sauciella (Zeller): Colombia
202. argyrophleps Dyar: México, Guatemala
203. euchroma Dyar: Brazil
Caristanius Heinrich, new genus
204. pellucidellus (Ragonot), new comb.: Puerto
Rico, St. Vincent, Jamaica, Surinam,
Brazil
melanoplaga (Hampson)
205. decoloralis (Walker), new comb.:
U.S.
metagrammalis (Walker)
furfurellus (Aulst)
floridellus (Hulst)
206. guatemalellus (Ragonot),
Guatemala
Erreia Zeller
207. zinckenella (Treitschke):
U.S., Tropical America
etiella (Treitschke)
schisticolor Zeller
villosella Hulst
rubribasella Hulst
GuyprTocera Ragonot
208. consobrinella (Zeller):
U.S. (florida), Tropical
Southern
new comb.:
Europe, Asia,
Eastern U. S., Can-
ada
busckella (Dyar)
Pima Hulst
209. boisduvaliella (Guénée), new comb.: EKu-
rope, Canada
210. albiplagiatella (Packard), new comb.: EHast-
ern U. S., Canada
211. albiplagiatella occidentalis Heinrich, new
race: Western U.S.
212. fosterella Hulst: Western U.S., Canada
213. vividella (McDunnough), new comb.: Can-
ada
214. albocostalialis (Hulst), new comb.: Western
U.S., Canada
215. albocostalialis subcostella (Ragonot), new
comb.: Southwestern U.S.
216. fulvirugella (Ragonot,) Newcomb.: Western
U.S. (California)
217. granitella (Ragonot), new comb.: Western
piperella (Dyar)
56.
57.
58.
59.
60.
61.
62.
63.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
218. parkerella (Schaus), new comb.: Western
U.S. (Montana)
InterRsEctiIo Heinrich, new genus
219. denticulella (Ragonot), new comb.: North-
western U.S., Canada
220. columbiella (McDunnough), new comb.:
Northwestern U.S., Canada
221. ruderella (Ragnot), new comb.: ‘‘N.
Amer.”’ (California?)
222. niviella (Hulst) new comb.:
Ambesa Grote
223. laetella Grote: Western U. S., Canada
224. walsinghami (Ragonot): Western U. S.
monodon Dyar
225. walsinghami mirabella Dyar, new status:
U.S. (Southern California)
226. lallatalis (Hulst): Western U. S. (Nevada,
Utah)
U.S., Canada
Carastia Hiibner
227. bistriatella (Hulst), new comb.: Western
U.S. (California)
228. incorruscella (Hulst), new comb.: Western
U.S. (California)
229. actualis (Hulst), new comb.:
Canada
Immyrua Dyar
230. nigrovittella Dyar:
Oreana Hulst
231. unicolorella (Hulst): Eastern U. S., Canada
leucophaeella (Hulst)
Otysrta Heinrich
232. aliculella (Hulst), new comb.:
U.S
Western U.S.,
Eastern U. S.
Southwestern
oberthuriella (Ragonot)
233. furciferella (Dyar) new comb.:
ern U.S. (Arizona)
SALEBRIACUS Heinrich, new genus
234. odiosellus (Hulst), new comb.:
U.S.
bakerella (Dyar)
yumaella (Dyar)
SaLesBriaria Heinrich, new genus
Southwest-
Western
235. turpidella (Ragonot), new comb.: Southern
U.S.
ademptandella (Dyar)
236. nubiferella (Ragonot), new comb.: U.S.
237. engeli (Dyar) U.S.
238. annulosella (Ragonot), new comb.: U. S.
(Texas, North Carolina)
robustella (Dyar)
239. tenebrosella (Hulst), new comb.: U.S.
quercicolella (Ragonot)
heinrichalis (Dyar)
240. pumilella (Ragonot) new comb.: Southeast-
ern U.S.
georgiella (Hulst)
241. fructetella (Hulst) new comb.:
rectistrigella (Dyar)
U.S.
300329—56——22
64.
65.
66.
67.
68.
321
QUASISALEBRIA Henrich, new genus
242. admixta Heinrich, new species:
LOTS
OrtTHOLEPIS Ragonot
243. jugosella Ragonot: Eastern U. S., Canada
244, pasadamia (Dyar), new comb.: U. S., Can-
ada
Poxoprustis Ragonot
245. arctiella (Gibson):
Merropterra Grote
Emmerita Hampson
246. mirandella Ragonot: Western U.S.
247. cviatella Dyar: U.S. (Illinois, Mississippi)
248. pravella (Grote): U.S., Canada
Western
Alaska, Canada
249. abditiva Heinrich, new species: Canada
Nepuorteryx Hiibner
250. subfuscella (Ragonot), new comb.: Eastern
U.S., Canada
semiobscurella (Hulst)
251. delassalis Hulst: Western U. S.
purpurella (Hulst)
pudibundella (Ragonot)
252. delassalis fraudifera Heinrich, new race:
Canada (British Columbia), U. S.(Wash-
ington)
253. rubescentella (Hulst): U.S.
254. fernaldi (Ragonot), new comb.: U. S.,
Canada
255. dammersi Heinrich, new species: Western
U.S. (California, Arizona)
dammersi floridensis Heinrich, new race:
U.S. (Florida)
vetustella (Dyar), new comb.:
ada
inconditella (Ragonot), new comb.:
ern U.S. (Arizona, Colorado)
subcaesiella (Clemens), new comb.:
Canada
contatella (Grote)
virgatella (Clemens), new comb.:
Canada
quinquepunctella (Grote)
carneella Hulst: U.S., Canada
inquilinella (Ragonot)
basilaris Zeller: U.S., Canada
termitalis (Hulst), new comb.:
U.S., Canada
levigatella (Hulst)
termitalis yuconella
Alaska
bifasciella Hulst: U.S. (Arizona)
nogalesella (Dyar)
256.
257. U. S., Can-
258. West-
259. U.S,
260. UESS:,
261.
262.
263. Western
264. Dyar, new status:
265.
266. uvinella (Ragonot), new comb.: Eastern
afflictella (Hulst)
liquidambarella (Dyar)
267. celtidella (Hulst), new comb.: U.S.
322
69.
70.
72.
73.
74.
75.
76.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
268. rubrisparsella (Ragonot): U.S.
rufibasella (Ragonot)
croceella (Hulst)
texanella (Hulst)
269. gilvibasella Hulst:
lacteella (Hulst)
270. crassifasciella Ragonot:
decupientella Dyar
crataegella B. and McD.
271. bisra Dyar: México
TiascaLa Hulst
272. reductella (Walker): Eastern U. S.
gleditschiella (Fernald)
Tusa Heinrich, new genus
273. finitella (Walker), new comb.:
Canada
melanellus (Hulst)
274. umbripennis (Hulst), new comb::
(Colorado)
gillettella (Dyar)
275. oregonella. (Barnes and McDunnough), new
comb.: U.S. (Oregon)
infinitella (Dyar), new comb.:
U.S. (Texas)
Eastern U.S.
Eastern U.S.,
UW. ©:
276. México
. Homorocraprua Ragonot
277. lanceolella Ragonot: Pert
TELETHuUsIA Heinrich, new genus
278. ovalis (Packard), new comb.:
latifasciatella (Packard)
geminipunctella (Ragonot)
modestella (Hulst)
279. rhypodella (Hulst), new comb.:
gon’’)
Pxosus Heinrich, new genus
280. brucei (Hulst), new comb.:
281. funerellus (Dyar), new comb.:
Canada
curvatellus (Ragonot), new comb.:
U.S.
incertus Heinrich, new species:
U.S. (California)
Actrix Heinrich, new genus
284. nyssaecolella (Dyar), new comb.:
U.S., Canada
U.S. (“Ore-
Western U. S.
Western U.S.,
282. Western
283. Western
Hastern
285. dissimulatrix Heinrich, new species: Eastern
U.S. (Virginia)
SryLopaLpia Hampson
286. lunigerella Hampson: West Indies, México
287. scobiella (Grote), new comb.: U.S. (Texas,
Colorado)
decimerella (Hulst)
288. argentinensis Heinrich, new species: Ar-
gentina
Pyta Grote
289. fasciolalis (Hulst), new comb.: Canada
(British Columbia)
290. impostor Heinrich, new species: Western
U.S., Canada
291. aequivoca Heinrich, new species: Western
Canada
77.
292. insinuatrix Heinrich, new species: Canada
(Manitoba)
293. aenigmatica Heinrich, new species: U. S.,
Canada
294.
295.
criddlella Dyar: Canada (Manitoba)
fusca (Haworth), new comb.: Holarctic
moestella (Walker)
frigidella (Packard)
cacabella (Hulst)
triplagiatella (Dyar)
hypochalciella (Ragonot), new comb.: North-
western U. S., Canada.
blackmorella (Dyar)
296.
297. hanhamella Dyar: Canada (Manitoba)
298. scintillans (Grote): Western U. S. (Califor-
nia)
feella Dyar
299. sylphiella Dyar: Northwestern U. S., Canada
300. rainierella Dyar: Northwestern U.S. (Wash-
ington)
301. aeneella Hulst: Western U. S. (Colorado,
Utah)
302. aeneoviridella Ragonot: Western U. S.,
Canada
303. metalicella Hulst: Western U.S. (Colorado,
Utah)
304. fasciella Barnes and McDunnough: North-
western U.S. (California)
nigricula Heinrich, new species:
U.S. (Nevada)
viridisuffusella Barnes and McDunnough:
Western U.S. (California)
Diorycrria Zeller
Pinipestis Grote
307. abietella (Denis and Schiffermiiller):
ern Hemisphere
decuriella (Hisbner)
abietivorella (Grote)
elegantella (Hulst)
sysstratiotes Dyar:
reniculella (Grote):
ponderosae Dyar:
California)
majorella Dyar: México
muellerana Dyar
disclusa Heinrich: Hastern U. S.
auranticella (Grote): Western U.S., Canada
miniatella Ragonot
zanthaenobares Dyar
erythropasa (Dyar):
(Arizona)
315. horneana (Dyar):
305. Western
306.
North-
308.
309.
310.
Guatemala
Northern U.S., Canada
Western U.S. (Montana,
311.
312.
313.
314. Southwestern U. S.
Cuba
316. pygmaeella Ragonot: Hastern U.S.
317. zimmermani (Grote): U.S., Canada
delectella (Hulst)
austriana (Cosens)
318. cambiicola (Dyar): Western U.S.
319. amatella (Hulst): Eastern U.S.
320. albovittella (Hulst): Western U.S.
78.
80.
81.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
321. gulosella (Hulst), new comb.: Western U.S.
(Colorado, New Mexico)
322. baumhoferi Heinrich, new species: South-
western U.S. (Arizona)
323. subtracta Heinrich, new species: South-
western U. S. (New Mexico)
clarioralis (Walker): Eastern U.S.
brunneella (Dyar)
Orycrometoria Ragonot
325. fossulatella Ragonot:
ical America
moeschleri (Ragonot)
324.
U. S. (Texas), Trop-
. Saravta Ragonot
326. edwardsialis (Hulst), new comb.: Western
AISE
polyphemella (Ragonot)
327. pullatella (Ragonot), new comb.: Western
U.S.
punctella (Dyar), new comb.: México
punctella septentrionaria Heinrich, new race:
_ Western U. S.
incanella (Hulst), new comb.:
aridella (Dyar)
328.
329.
330. Western U.S.
331. atrella (Hulst), new comb.: Western U. S.
(Colorado)
332. caudellella (Dyar), new comb.: Western
U.S., Canada
333. dnopherella Ragonot: Western U. S. (Cali-
fornia)
334. nigrifasciella Ragonot: Western U.S., Can-
ada
335. cinereella Hulst: Western U. S. (Colorado)
336. rubrithoracella (Barnes and McDunnough),
new comb.: Western U.S.
337. tephrella Ragonot: Western U. S. (Wash-
ington)
338. alpha Heinrich, new species: Canada (Sas-
katchewan)
339. beta Heinrich, new species: Western U. S.,
Canada
340. gamma Heinrich, new species: Western
U.S. (California)
341. iota Heinrich, new species: Western U. S.
(California)
perfuscalis (Hulst):
excantalis (Hulst)
epsilon Heinrich, new species: Western U.S.
phi Heinrich, new species: Western U. S.
342. Western U. S.
343.
344,
345. kappa Heinrich, new species: Western U.S.
(Arizona)
346. delta Heinrich, new species: Western U. S.
PuitopEMa Heinrich, new genus
347. rhoiella (Dyar), new comb.: Western U. S.
Lipoerapuis Ragonot
348. fenestrella (Packard):
fornia)
humilis Ragonot
349. leoninella (Packard):
ada
pallidella (Dyar)
Western U. S. (Cali-
Western U. S., Can-
82.
83.
84.
85.
86.
87.
88.
323
350. truncatella (Wright), new comb.: South-
western U.S. (California)
351. umbrella (Dyar), new comb.:
U.S. (California)
352. subosseella Hulst: Bahamas
AveE.puiA Heinrich, new genus
353. petrella (Zeller), new comb.:
rubiginella (Walker)
rufinalis (Walker)
hapsella (Hulst)
354. ochripunctella (Dyar), new comb.:
U.S. (California)
Tora Heinrich, new genus
355. galdinella (Schaus), new comb.:
Islands
Ura Walker
356. lithosella (Ragonot), new comb.:
western U. S., México
luteella Hulst)
roseitinctella (Dyar), new comb.:
western U. S., México
senta Heinrich, new species:
U.S. (Texas, Arizona)
rubedinella (Zeller), new comb.:
ida), Tropical America
translucida (Walker)
rufescentalis (Walker)
minualis (Walker)
deprivalis (Walker)
venezuelalis Walker
pyrrhochrellus (Ragonot)
Exasmopaupus Blanchard
360. lignosellus (Zeller): U.S., Tropical America
angustellus Blanchard
tartarella (Zeller)
incautella (Zeller)
major (Zeller)
anthracellus Ragonot
carbonella (Hulst)
puer Dyar
Acroncosa Barnes and McDunnough
361. albiflavella Barnes and McDunnough: West-
ern U. S. (California)
362. albiflavella castrella Barnes and McDun-
nough: Western U.S. (New Mexico)
363. similella Barnes and McDunnough: West-
ern U.S. (Nevada, Utah)
PassapDENA Hulst
364. flavidorsella (Ragonot):
México
canescentella (Hulst)
constantella Hulst
cinctella (Hulst)
Utopnora Ragonot
Acromeseres Dyar
365. groteii Ragonot: Eastern U.S.
tephrosiella Dyar
366. guarinella (Zeller):
dialithus (Dyar)
Southwestern
U.S.
Western
Galdpagos
South-
357. South-
358. Southwestern
359. U.S. (Flor-
Western U. S.,
Cuba, Colombia
324 UNITED STATES NATIONAL MUSEUM BULLETIN 207
89. CHorrEeRA Dyar
367. idiotes Dyar: Panama
368. extrincica (Dyar), new comb.:
369. postica (Zeller), new comb.:
90. Tacoma Hulst
370. feriella Hulst: Southwestern U.S.
submedianella Dyar
91. ApELPERGA Heinrich, new genus
371. cordubensiella (Ragonot), new comb.: Ar-
gentina
92. Humysta Dyar
372. mysiella (Dyar):
373. maidella (Dyar):
Cuba
Colombia
Western U. S.
Western U. S., Canada
374. pallidipennella (Hulst), new comb.: West-
ern U.S.
375. fuscatella (Hulst): Western U. S. (Cali-
fornia)
376. semicana Heinrich, new species: Western
U. S. (Washington)
93. Drvrt1aca Barnes and McDunnough
377. ochrella Barnes and McDunnough: South-
ern U.S. (Florida)
378. simulella Barnes and McDunnough: South-
ern U.S. (Florida)
379. parvulella Barnes and McDunnough: South-
ern U.S. (Florida)
380. parvulella consociata Heinrich, new race:
Colombia
94. MacrorrHInia Ragonot
Dolichorrhinia Ragonot
381. aureofasciella Ragonot: Southwestern U.S.,
México
382. placidella (Zeller):
95. Ocata Hulst
383. dryadella Hulst: Southern U.S. (Florida)
platanella (Grossbeck)
96. Vauprvia Ragonot
Maricopa Hulst
384. coquimbella Ragonot:
385. lativittella (Ragonot):
México
aureomaculella (Dyar)
386. walkerella (Ragonot), new comb.:
97. Prorasta Heinrich, new genus
387. mirabilicornella (Dyar), new comb.:
ern U.S. (California)
98. HreTrRoGRAPHis Ragonot
Mona Hulst
388. morrisonella Ragonot:
coloradensis Ragonot
olbiella (Hulst)
ignistrigella Ragonot
palloricostella (Walter)
99. StauDINGERIA Ragonot
389. albipenella (Hulst):
olivacella Dyar
perluteella Dyar
Brazil
Chile
Southwestern U. S.,
Chile
West-
U.S., México
Western U. S.
100. Huustia Ragonot
390. undulatella (Clemens):
rubiginalis (Walker)
obsipella (Hulst)
fumosella (Aulst)
101. Honora Grote
391. mellinella Grote: U.S.
ochrimaculella Ragonot
392. subsciurella Ragonot: Western U.S.
393. sciurella Ragonot: Western U. S. (Cali-
fornia)
394. dotella Dyar: Western U.S. (California)
395. montinatatella (Hulst): Western U. S.
canicostella Ragonot
396. perdubiella (Dyar), new comb.: Western
U.S.
U.S., Canada
102. Honorinus Heinrich, new genus
397. fuliginosus Heinrich, new species:
103. Oncoxasis Zeller
Endommasis Hampson
398. anticella Zeller: Tropical America
migritella (Hampson)
104. Casotia Ragonot
Encystia Hampson
399. semidiscella Ragonot: Argentina
400. schini (Berg): Argentina
401. rhythmatica Dyar: Panamé
402. cundajensis (Zeller): Colombia
impeditella (Zeller)
403. bonhoti (Hampson), new comb.:
Jamaica
105. Canarsta Hulst
404. ulmiarrosorella (Clemens):
pneumatella (Hulst)
ulmella (Ragonot)
fuscatella (Hulst)
gracilella Hulst
feliculella Dyar
106. Harnocua Dyar
405. velessa Dyar:
107. Eurytamasis Dyar
406. ignifatua Dyar:
Cuba
108. Euryramipia Ragonot
407. ignidorsella (Ragonot):
México, Panamé
109. Wunper1A Grossbeck
408. neaeriatella Grossbeck: U.S. (Florida)
110. OzpotTHm1a Hampson
Synothmia Hampson
409. endopyrella Hampson: México, Bahamas
bahamasella (Hampson)
111. StynoBasis Hampson
410. rubripurpurea Hampson: México, Brazil
112. Diviana Ragonot
Dannemora Hulst
411. eudoreella Ragonot:
edentella (Hulst)
Peri
Bahamas,
U.S., Canada
Panamdé
Panamé, Puerto Rico,
U. S. (Arizona),
Eastern U. S.
113.
114.
115.
116.
117.
118.
119.
120.
121.
122.
123.
124.
125.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
PanatKa Hulst
412. nymphaeella (Hulst): Eastern U. S.
verecuntella (Grossbeck)
CacozopHEera Dyar
413. venosa Dyar: Guatemala
Psorosina Dyar
414. hammondi (Riley): Eastern and Central
U.S., Canada
angulella Dyar
Patricroua Heinrich, new genus
415. semicana Heinrich, new species: Utah
Pacontus Heinrich, new genus
416. corniculatus Heinrich, new species:
Rico
Aptunea Heinrich, new genus
417. macropasa (Dyar), new comb.:
México
418. imperfecta (Dyar), new comb.:
ANDERIDA Heinrich, new genus
419. sonorella (Ragonot), new comb.:
U.S. (Arizona)
placidella (Dyar)
Casstana Heinrich, new genus
420. malacella (Dyar), new comb.:
Puerto Rico, Virgin Islands
Mescinra Ragonot
421. triloses Dyar:
mosces Dyar
pandessa Dyar: Guatemala
bacerella Dyar: Cuba
estrella Barnes and McDunnough: U. S.
(Florida)
moorei Heinrich, new species: British Guiana
parvula (Zeller): Colombia
commatella (Zeller): Colombia
berosa Dyar: Panam4, Puerto Rico
peruella Schaus: Pert
430. discella Hampson: México, Guatemala
431. indecora Dyar: México
Nona Ragonot
Hypermescinia Dyar
432. exiguella (Ragonot):
lambella (Dyar)
Puestinia Hampson
433. costella Hampson:
Comotta Dyar
434. torsicornis Dyar: Panamé
435. convergens (Dyar), new comb.:
Bema Dyar
Relmis Dyar
neuricella (Zeller), new comb.:
America
myja Dyar
437. fritilla Dyar: Guatemala
438. ydda (Dyar), new comb.:
Guiana
439. yddiopsis (Dyar), new comb.: Cuba
440. fifaca (Dyar), new comb.: Panama
Puerto
Guatemala,
Guatemala
México,
México,
Panam&
422.
423.
424.
425,
426.
427.
428.
429.
Tropical America
Jamaica, Puerto Rico
Guatemala
436. Tropical
Panamd, French
126.
127.
128.
129.
130.
325
Homogosoma Curtis
Phycidea Zeller
electellum (Hulst): U. S., México, Guate-
mala, Cuba, British West Indies
opalescellum (Hulst)
texanellum Ragonot
tenuipunctella Ragonot
differtella Barnes and McDunnough
. stypticellum Grote: U.S., Canada
uncanale Hulst
. striatellum Dyar: Southwestern U. S.
. oslarellum Dyar: Western U. S.
. oslarellum breviplicitum Heinrich, new race:
Southwestern U. S. (California)
. illuviellum Ragonot: U.S. (Arizona, Colo-
rado), México
candidella Hulst
. iluviellum emendator Heinrich, new race:
Western U.S.
. Imitator Heinrich, new species: Southwestern
U.S. (California)
. longiventrellum Ragonot:
noctividella Ragonot
. albescentellum Ragonot: Western U.S.
elongellum Dyar
. impressale Hulst: Western U. S., Canada
. inornatellum (Hulst): Eastern U.S.
. deceptorium Heinrich, new species: U. S.
(Pennsylvania), Canada
. discrebile Heinrich, new species, Brazil
. peregrinum Heinrich, new species:
(California), Costa Rica
. vepallidum Heinrich, new species: Argentina
. ditaeniatellum Ragonot: Chile
. oconequensis (Dyar), new comb.: Pert
. assitum Heinrich, new species: Pert
. acmacopterum Ragonot: Chile
. nimbosellum Ragonot: Chile
462. unionellum Ragonot: México
Pataconia Ragonot
463. magellanella (Ragonot):
Rorrupa Heinrich, new genus
464. mucidella mucidella (Ragonot), new comb.:
Western U. S., Canada
465. mucidella reliquella (Dyar), new comb.:
Eastern U. S., Canada
466. mucidella olivaceela (Ragonot), new comb.:
Tropical America
musiosum (Dyar)
cubella (Dyar)
467. mucidella affusella (Ragonot), new comb.:
Argentina
STREPHOMESCINIA Dyar
468. schausella Dyar:
UnapiLua Hulst
Strymaz Dyar
441.
Chile
De Ss
Chile
Cuba
326
131.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
469. erronella (Zeller): Tropical America.
ubacensis (Zeller)
bipunctella (Hampson)
dorae (Dyar)
pyllis (Dyar)
470. maturella (Zeller): Colombia, Guatemala,
Cuba
471. albidiorella (Richards and Thomason); new
comb.: Pert
472. floridensis Heinrich, new species: U.S. (Flor-
ida)
473. nasutella Hulst: U.S. (New Mexico)
Laetitia Ragonot
Laosticha Hulst
474. coccidivora, (Comstock):
pallida (Comstock)
dilatifasciella (Ragonot)
hulstii Cockerell
WS.
475. coccidivora quadricolorella (Dyar), new
comb.: Southwestern U. S.
476. coccidivora cardini Dyar: Cuba, U. S.
(Florida)
477. obscura Dyar: Cuba
478. portoricensis Dyar: Puerto Rico
479. melanostathma (Meyrick), new comb.: Ar-
dl gentina
480. amphimetra (Meyrick), new comb.: Argen-
tina
481. zamacrella Dyar: Western U.S. (California)
482. myersella Dyar: Eastern U.S.
483. ephestiella (Ragonot): Southwestern U. S.
(Arizona)
lustrella (Dyar)
484. fiskella Dyar: Eastern U. S. (North Caro-
lina)
485. glomis (Dyar), new comb.: Panamé
132.3BarHaa Heinrich, new genus
486. basimaculatella (Ragonot), new comb.:
133.
Western U.S.
eremella (Dyar)
487. goyensis (Ragonot), new comb.:
Uruguay, Argentina
488. goyensis olivacea Heinrich, new race: Argen-
tina
489. homoeosomella, (Zeller), new comb.:
ical America,
bodkini (Dyar)
rusto (Dyar)
taboga (Dyar)
saissetiae (Dyar)
490. haywardi Heinrich, new species:
491. glabrella (Dyar), new comb.:
492. squalida (Walker), new comb.:
Ruacea Heinrich, new genus
493. packardella (Ragonot), new comb.:
ern U.S.
orobanchella (Dyar)
494, stigmella (Dyar), new comb.:
U.S. (California), México
maculicula (Dyar)
Brazil,
Trop-
Argentina
Guatemala
Brazil
West-
Southwestern
134. ZorHopia Hubner
136.
137.
138.
139.
140.
141.
143.
Dakruma Grote
495. convolutella (Hiibner):
ada
grossulariella (Hiibner)
turbatella (Grote)
grossularvae (Riley)
franconiella (Hulst)
bella Hulst
thouna Dyar
dilativitta Dyar
magmyicans Dyar
Europe, U.S., Can-
. Menitara Walker
496. prodenialis Walker: U.S.
bolliz (Zeller)
497. dentata (Grote): U.S.
doddalis Dyar
OtycEeLLa Dyar
498. junctolineella (Hulst): Southern U.S. (Texas)
499. junctolineella pectinatella (Hampson): Méx-
ico
500. nephelepasa (Dyar:) México
501. subumbrella Dyar: Western U.S.
Ouyca Walker
502. phryganoides Walker: Dominican Repub-
lic, Haiti
ALBERADA Heinrich
503. parabates (Dyar):
504. bidentella (Dyar):
(Texas, Arizona)
505. holochlora (Dyar):
(Texas)
Nawata Heinrich
506. substituta Heinrich: Pert
Cactosiastis Ragonot
Neopyralis Bréthes
cactorum (Berg): Argentina, Uruguay, Aus-
tralia
. ronnai (Bréthes): Brazil
. doddi Heinrich: Argentina
. mundelli Heinrich: Pert
511. bucyrus Dyar: Argentina
CaHELa Heinrich
512. ponderosella (Barnes and McDunnough):
Western U. S., México
purgatoria (Dyar)
interstitialis (Dyar)
phoenicis (Dyar)
U.S., México
Southwestern U. S.
Southwestern U. S.
507.
. Rumatua Heinrich
Southern U. S.
Western U.S.
Southwestern U.S. (Ari-
513. glaucatella (Hulst):
514. bihinda (Dyar):
515. polingella (Dyar):
zona, Texas)
Yosrmitia Ragonot
516. graciella (Hulst): Western U.S.
517. longipennella (Hulst): Southwestern U. S.
(Texas)
518. fieldiella (Dyar):
Arizona)
519. didactica Dyar:
Western U.S. (California,
México
144,
145.
146.
147.
148.
149.
150.
151.
152.
156.
157.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Tucumania Dyar
520. tapiacola Dyar:
521. porrecta Dyar:
EremsBerca Heinrich
522. leuconips (Dyar):
523. creabates (Dyar):
fornia)
524. insignis Heinrich: México
SatamBona Heinrich
525. analamprella (Dyar):
Parotyca Dyar
526. asthenosoma (Dyar):
Sigeveaira Heinrich
527. chilensis Heinrich: Chile
528. huanucensis Heinrich: Pert
529. transilis Heinrich: Pert
AMALAFRIDA Heinrich
530. leithella (Dyar):
Colombia
Ozamia Ragonot
531. lucidalis (Walker): West Indies
532. fuscomaculella (Wright): Southwestern
U.S. (California)
heliophila Dyar
fuscomaculella clarefacta Dyar: U. S.
(Texas), México
thalassophila Dyar: U.S. (California)
immorella (Dyar), new comb.: México
stigmaferella Dyar: Argentina
537. hemilutella Dyar: Argentina
538. punicans Heinrich: Argentina
Cacrosrosis Dyar
539. fernaldialis (Hulst): Southwestern U. S.
gigantella (Ragonot)
cinerella (Hulst)
longipennella (Hampson):
elongatella (Hampson):
maculifera Dyar: México
insignatella Dyar: México
strigalis (Barnes and McDunnough): West-
ern U. S., México
Drescomopsis Dyar
544. soraella (Druce):
drucella (Dyar)
subelisa Dyar
Argentina
Uruguay
Western U.S. (Arizona)
Western U. S. (Cali-
Argentina
French Guiana
West Indies, Venezuela,
533.
534,
535.
536.
540. México
541.
542.
543.
Tropical America
. Intatiza Dyar
545. gurbyris Dyar: Panamé
. Lascrtina Heinrich, new genus
546. canens Heinrich, new species: Southern
U.S. (Texas), México
. MerepuHestia Ragonot
547. simplicula (Zeller): U.S. (Florida), Puerto
Rico, Colombia, British West Indies
Srxea Heinrich, new genus
548. arizonella (Hulst), new comb.:
ern U.S. (Arizona)
Entmemacornis Dyar
549. proselytes Dyar: Guatemala
550. pulla Heinrich, new species: Brazil
Southwest-
158.
159.
160.
161.
162.
163.
164.
165.
166.
167.
168.
169.
170.
327
Cayrennia Hampson
551. rufitinctalis Hampson: French Guiana
Rioga, Heinrich, new genus
552. nexa Heinrich, new species: Argentina
Moerrses Dyar
553. dryopella (Schaus): Costa Rica
554. alveolella (Ragonot), new comb.: Brazil
555. emendata Heinrich, new species: Panama,
French Guiana
Moopnopsis Dyar
Campyloplesis Dyar
556. decipiens Dyar: México
557. perangusta (Dyar), new comb.: Trinidad
558. inornatella (Ragonot), new comb.: Costa
Rica, Brazil
559. parallela Heinrich, new species: Brazil, Pera
560. inveterella (Dyar), new comb.: Guatemala
561. portoricensis Heinrich, new species: Puerto
Rico
Eputica Ragonot
562. compedella (Zeller):
KuzopHera Zeller
Tropical America
563. semifuneralis (Walker): U. S., Canada,
México
aglaeella Ragonot
pallulella (Hulst)
564. ostricolorella Hulst: Eastern U. S.
565. nigricantella Ragonot:
México
griselda Dyar
Exvurertivus Heinrich, new genus
566. negator Heinrich, new species:
Evutoera Heinrich, new genus
567. ochrifrontella (Zeller), new comb.:
Canada
Serruginella (Ragonot)
PROSOEUZOPHERA Heinrich, new genus
568. impletella (Zeller), new comb.:
Jamaica, Puerto Rico
Farnosia Heinrich, new genus
569. quadripuncta (Zeller), new comb.: Costa
Rica, Panama, French Guiana, Colombia
Gennapius Heinrich, new genus
570. junctor Heinrich, new species:
ana
Micromescinia Dyar
571. pygmaea Dyar:
Epuestioprs Ragonot
572. gilvescentella Ragonot:
ada, México
nigrella Hulst
573. infimella Ragonot:
574. erythrella Ragonot:
coloradella (Hulst)
benjaminella Dyar
mignonella Dyar: U.S. (Texas)
erasa Heinrich, new species: U.S. (Florida)
lucidibasella Ragonot: Chile
productella Ragonot: Colombia (?)
indentella Dyar: Bermuda
Southwestern U. S.,
Pert
U. S.,
Colombia,
French Gui-
Panama
Western U.S., Can-
Eastern U.S.
Western U.S., Canada
575.
576.
577.
578.
579.
328
171.
172.
173.
174.
175.
176.
177.
178.
179.
180.
UNITED STATES NATIONAL MUSEUM BULLETIN 207
580. plorella Dyar:
vestilla (Dyar)
581. stictella (Hampson), new comb.:
West Indies
uniformella Hampson
granulella Hampson
582. noniella Dyar: Panam4
AZAERA Schaus
Calamophleps Dyar
583. muciella Schaus: Costa Rica, Guatemala,
Panama
squalidella (Dyar)
584. nodoses (Dyar), new comb.: Panam’
585. lophophera (Dyar), new comb.: Panamé
Moopna Hulst
586. ostrinella (Clemens): U.S., Canada
obtusangulella (Ragonot)
pelviculella Hulst
587. bisinuella Hampson:
Viruta Ragonot
588. edmandsae (Packard):
ada
dentosella Ragonot
Panama
Bahamas,
México, U.S. (Texas)
Eastern U. S., Can-
589. edmandsae serratilineella Ragonot, new sta-
tus: Western U.S., Canada
590. lugubrella Ragonot, new comb.: Western
U.S. (California)
591. pinei Heinrich, new species: Western U. S.
(Utah, Nevada)
592. inanimella (Dyar), new comb.: México,
Guatemala
ticitoa (Dyar)
593. laura (Dyar), new comb.: Guatemala
Manuatta Hulst
Hornigia Ragonot
594. setonella (McDunnough), new comb.: U.S.
(Utah), Canada (British Columbia)
595. broweri Heinrich, new species: Hastern U.S.
(Maine)
Verina Heinrich, new genus
596. supplicella (Dyar), new comb.: México,
Guatemala, Panam4, Brazil
Vacozsanta Heinrich, new genus
597. divergens (Butler), new comb.: Chile
Moopnetua Heinrich, new genus
598. paula Heinrich, new species: Guatemala,
Brazil, Argentina
Vouatica Heinrich, new genus
599. pachytaeniella (Ragonot), new comb.: Bra-
zil
600. trinitatis Heinrich, new species: Trinidad
Vezina Heinrich, new genus
601. parasitaria Heinrich, new species:
tina, Brazil
CaubELui1a Dyar
602. apyrella Dyar: Eastern U. S. (Maryland)
603. albovittella Dyar: Eastern U. S.
604. nigrella (Hulst), new comb.: Western U.S.
arizonella (Walter)
Argen-
181.
182.
183.
184.
185.
186.
187.
605. declivella (Zeller), new comb.: Panama,
Colombia
animosella (Dyar)
606. colorella (Dyar), new comb.: Panam&
607. clara Heinrich, new species: Puerto Rico
MicropHestia Dyar
608. animalcula Dyar: Panama
SosipaTRa Heinrich, new genus
609. rileyella (Ragonot), new comb.: Western
U.S., México
micaceella (Hampson): México
anthophila (Dyar), new comb.:
U.S. (Texas)
thurberiae (Dyar), new comb.: Western U. S.
nonparilella (Dyar), new comb.: Western
U.S. (Arizona)
614. majorella (Dyar), new comb.:
615. divergens (Dyar): Panamé
Beruutia Ragonot
616. championella Ragonot:
Risua Heinrich
617. mnoxia Heinrich: Cuba
618. contigua Heinrich, new species:
619. patriciella (Dyar), new comb.:
Prop1a Guénée
620. interpunctella (Hiibner):
interpunctalis (Hiibner)
zeae (Fitch)
latercula (Hampson)
glycinwwora (Matsumura)
621. dolorosa Dyar: Guatemala,
Awacasta Heinrich, new genus
622. kiihniella (Zeller): Cosmopolitan
fuscofasciella (Ragonot)
gitonella Druce
Epuestia Guénée
Hyphantidium Scott
623. elutella (Hitibner): Cosmopolitan
elutea (Haworth)
semirufa (Haworth)
rufa (Haworth)
sericarium (Scott)
roxburghi Gregson
unicolorella Staudinger
amarella Dyar
624. cautella (Walker):
defectella (Walker)
desuetella (Walker)
cahiritella Zeller
passulella Barrett
formosella (Wileman and South)
625. figulilella Gregson: Europe, Asia, Africa,
Hawaii, Australia, North America (U.S.,
California), South America
jficulella Barrett
milleri Zeller
jiguliella Forbes
jigulella Curran
venosella Turati
ernestinella 'Turati
610.
611. Western
612.
613.
México
Guatemala
Puerto Rico
Cuba
Cosmopolitan
Cosmopolitan
188.
189.
190.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
NIcETIODES Schaus
626. apianella Schaus:
Varneria Dyar
Galfpagos Islands
627. postremella Dyar: Eastern U. S.
628. nannodes Dyar: Panam&
629. atrifasciella Barnes and McDunnough:
Southern U. S. (Florida)
630. dubia Heinrich, new species:
Evrytumia Ragonot
631. hospitella (Zeller): Southern and Western
U.S.
Puerto Rico
spaldingella Dyar
632. hospitella yavapaella Dyar, new status:
Western U.S.
633. angulella Ely: Eastern U. S., Canada
diffusella Ely
634. fumella Ely: Eastern U.S. (Connecticut)
191.
192.
193.
194.
329
Ereieva Heinrich, new genus
635. quantulella (Hulst), new comb.: Southern
U.S. (Texas), West Indies
santiagella (Dyar)
636. coca (Dyar), new comb.:~ Panam4
coquilla (Dyar)
mossa (Dyar)
uncta (Dyar)
637. parvulella (Ely), new comb.: Eastern U. S.
(Connecticut)
Casnia Dyar
638. myronella Dyar: Eastern U. S.
Micropuycira Dyar
639. titillella Dyar: Panama
Rasreia Heinrich, new genus
640. conops (Dyar), new comb.: Panam&é
Species unplaced or unrecognized
brevistrigella Ragonot [Zophodia]
came Dyar [Huzophera]
cervinistrigalis Walker [Hypochalcia]
clitellatella Ragonot [Hornigia]
corrientellus Ragonot [Elasmopalpus]
daedalella Ragonot [Euzophera]
disticta Zeller [Psorosa]
dulciella Hulst [Honora]
Jamula Zeller [Myelois]
flavicornella Ragonot [Phycitopsis]
formulella Schaus [Moodna]
Suscifrontella Zeller [Nephopteryz]
gais Dyar [Huzophera]
grossipunctella Ragonot [Myelois]
hulstiella Ragonot [Hypochalcia]
infusella Zeller [Myelois]
intextella Zeller [Huzophera]
trichampa Dyar [Anthropteryz]
megalopalis Hampson [Huzopherodes]
migricans Hulst [Salebria]
olwella Hampson [Moodna]
postflavida Dyar [Huzophera]
putidella Schaus [Hucampyla]
rinmea Dyar [Huzophera]
subcanella Zeller [Zophodia]
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Figures
The figures on the following pages are not drawn throughout to any fixed scale, the
smaller genitalia being shown in greater enlargement than those of the larger species and the
female genitalia on a uniformly smaller scale than the male genitalia. However, for the
species of any given genus a uniform scale within the sexes has been attempted.
The illustrations of venation in figures 1-53 and 55-130 are of males unless otherwise
‘stated. The illustrations of male genitalia in figures 54, 131-637, and 915 show first under
each figure, unless otherwise stated, a ventral view of the genitalia with one harpe and the
aedeagus omitted, and this view is usually accompanied by a drawing of the aedeagus and
other associated parts. Illustrations of female genitalia are shown in figures 638-914 and
‘915-1138. For a few species, details of antennae and eyes have been added.
331
332 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 1-8.—VENATION (MALE UNLESS
Oarwnr
OOF
. Cryptoblabes gnidiella (Milliére).
. Cryptoblabes rutilella Zeller.
. Acrobasis indigenella (Zeller).
. Hypsipyla grandella (Zeller).
. Fundella pellucens Zeller; 6a, underside of
OTHERWISE NOTED).
Sematoneura atrovenosella Ragonot.
anal angle of hind wing with pocket un- 1.Cryptoblabes
folded to show sex-scaling. Ie 2. Cryptoblabes
Difundella corynophora Dyar.
Birinus russeolus Heinrich, new species.
4. Sematoneura
6. Fundella
5. Hypsipyla
8. Birinus
7. Difundella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 333
Fieures 9—-16.—VENATION (MALE UNLESS
OTHERWISE NOTED).
9. Promylea lunigerella lunigerella Ragonot.
10. Coptarthria dasypyga (Zeller).
11. Anadelosemia senesciella (Schaus).
12. Dasypyga alternosquamella Ragonot.
13. Ceracanthia mamella (Dyar).
14, Megarthria peterseni (Zeller).
Coptarthria 15. Drescoma cyrdipsa Dyar.
16. Drescomopsis soraella (Druce).
Se
11. Anadelosemia
9. Promylea
12. Dasypyga
13. Ceracanthia
16, Drescomopsis
334 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 17-24.—VENATION (MALE UNLESS
OTHERWISE NOTED).
17. Etiella zinckenella (Treitschke).
18. Ambesa laetella Grote.
19. Immyrla nigrovittella Dyar.
20. Salebria plumbella (Schiffermiller), type
of the genus Salebria Zeller, not repre-
sented in the American fauna.
21. Emmerita mirandella (Ragonot).
22. Meropiera pravella (Grote).
23. Ortholepis jugosella Ragonot.
24. Polopeustis annulatella (Zetterstedt).
18.Ambesa
17. Etiella
20.
Salebria
Ortholepis
22Meroptera Polopeustis
AMERICAN MOTHS OF THE SUBFAMILY
25:
Nephopteryx
26.
Epischnia
28.
Tlascala
27. Monoptilota
30.Stylopalpia
Lipographis
PHYCITINAE 335
Figures 25-32.—VENATION (MALE UNLESS
25.
26.
27.
28.
29.
30.
31,
32.
OTHERWISE NOTED).
Nephopteryz rhenella (Zincken).
Epischnia prodromella (Hiibner).
Monoptilota pergratialis (Hulst).
Tlascala reductella (Walker).
Pyla scintillans (Grote).
Stylopalpia lunigerella Hampson.
Lipographis fenestrella (Packard).
Selagia argyrella (Schiffermiller). This
genus occurs in our lists, but is not rep-
resented in the American fauna.
336 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 33-40.—VENATION (MALE UNLESS <
OTHERWISE NOTED).
33. Elasmopalpus lignosellus (Zeller).
34. Chorrera idiotes Dyar.
35. Acroncosa albiflavella Barnes and McDun-
nough.
36. Passadena flavidorsella (Ragonot).
37. Tacoma feriella Hulst.
38. Myelois cribrella (Hubner), an Old World 33. Elasmopal pus 34. Chorrera
genus, not represented in the American
fauna.
39. Anypsipyla univitella Dyar.
40. Apomyelois bistriatella (Hulst).
36.Passadena
35. Acroncosa
4-0. Apomyelois
AMERICAN MOTHS
41. Pseudodivona
SS —— —
S
42. Diatomocera
45. Rampylla
47. Praedonula
43. Cabima
44. Hyalospila
46. Davara
48. Sarasota
OF THE SUBFAMILY PHYCITINAE 337
Figures 41-48.—VENATION (MALE UNLESS
OTHERWISE NOTED) AND MALE GENITALIA.
41. Pseudodivona commensella Dyar. Vena-
tion.
42. Diatomocera tenebricosa (Zeller). Vena-
tion.
43. Cabima dosia Dyar, venation, type of
Cabima Dyar, asynonym of Diatomocera
Ragonot.
44, Hyalospila stictoneurella Ragonot. Vena-
tion.
45. Rampylla orio Dyar. Venation.
46. Davara caricae (Dyar). Venation.
47. Praedonula almonella (Dyar). Venation.
48. Sarasota furculella (Dyar). Venation.
338
Figures 49-54.—VENATION (MALE UNLESS
49.
50.
51.
52.
53.
54.
OTHERWISE NOTED).
Zamagiria pogerythrus Dyar. Venation.
Anegcephalesis arctella (Ragonot), figured
from its synonym, cathaeretes Dyar.
Venation.
Dioryctria abietella (Denis and Schiffer-
miller). Venation.
Laodamia faecella (Zeller). Venation.
Oryctometopia fossulatella Ragonot. Vena-
tion.
Hypochalcia ahenella (Schiffermiiller), male
genitalia; male genitalia 54a, detached
harpe in ventral view; 54b, dorsal view
of tegumen and uncus; 54c, aedeagus;
54d, gnathos; 54e, anellus; 54f, sternite
and tergite of eighth abdominal seg-
ment. (Type of an Old World genus
not represented in the New World.)
UNITED STATES NATIONAL MUSEUM BULLETIN 207
49.
Ee SS ete Anagcephalesis
51. Dioryctria
52.
Laodamia
53.
Oryctometopia
ahenella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 339
Fieures 55-62.—VENATION (MALE UNLESS
OTHERWISE NOTED).
55. Cabotia semidiscella Ragonot.
56. Oncolabis anticella Zeller.
57. Honora mellinella Grote.
58. Edulica compedella (Zeller).
56. Oncolabis 59. Ocala dryadella Hulst.
60. [llatila gurbyris Dyar.
61. Macrorrhinia aureofasciella Ragonot.
62. Psorosina hammondi (Riley).
~$8. Edulica
NS = Y
=
Cre Sf
se
a
. :
a Sean Cacia — 60. Illatila
61. Macrorrhinia 62. Psorosina
340 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figtres 63-70.—VENATION (MALE UNLESS
OTHERWISE NOTED).
63. Laetilia coccidivora (Comstock).
64. Hulstia undulatella (Clemens).
65. Tucumania tapiacola Dyar.
66. Cactoblastis cactorum (Berg), female.
67. Ozamia fuscomaculella clarefacta Dyar.
68. Yosemitia graciella (Hulst).
69. Parolyca asthenosoma (Dyar).
70. Rioja nera Heinrich, new species.
63. Laetilia 64. Hulstia
66. Cactoblastis
es
67. Ozamia 68.Yosemitia
69. Parolyca 70 Rioja
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 341
Ficures 71-79.—VENATION (MALE UNLESS
OTHERWISE NOTED).
71. Olyca phryganoides Walker.
72. Cayennia rufitinctalis Hampson.
73. Cactobrosis fernaldialis (Hulst).
74. Melitara prodenialis Walker.
75. Olycella junctolineella (Hulst).
76. Hxuperius negator Heinrich.
77. Euzophera cinerosella (Zeller).
78. Hulogia ochrifrontella (Zeller), female.
79. Hulogia ochrifrontella (Zeller), hind wing of
female, showing variation in venation.
73. Cactobrosis
74. Melitara
6. Exuperius
%S on
75. Olycella
Euzophera 79. Eulogia
342
Figures 80-89.—VENATION (MALE UNLESS
80.
81.
82.
83.
84.
85.
86.
87.
88.
89.
OTHERWISE NOTED).
Canarsia ulmiarrosorella (Clemens).
Entmemacornis proselytes Dyar.
Moerbes dryopella (Schaus).
Moodnopsis decipiens Dyar, female.
Patriciola semicana Heinrich, new species.
Aptunga macropasa (Dyar), female. + ys
Moodnopsis inveterella (Dyar).
Eurythmia hospitella (Zeller).
Varneria postremella Dyar.
Nonia exiguella (Ragonot).
UNITED STATES NATIONAL MUSEUM BULLETIN 207
6]. Entmemacornis.
82. Moerbes
83.Moodnopsis 2
—~
—=-
85. Aptunga
=
87. Eurythmia 86.Moodnopsis
88 .Vorneria
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 343
SF
Eurythmidia
Phestinia
Strephomescinia
Micromescinia
ma é
id SS
awh 2; 97. Gennadius P 99. SS 46 Farnobia?
98. Farnobiaw”
AN SS
Figures 90-99.—VENATION (MALE UNLESS
OTHERWISE NOTED).
90. Wunderia neaeriatella Grossbeck, female.
91. Eurythmidia ignidorsella (Ragonot).
92. Phestinia costella Hampson.
93. Strephomescinia schausella Dyar.
94, Mescinia bacerella Dyar.
95. Eurythmasis ignifatua Dyar.
96. Micromescinia pygmaea Dyar.
97. Gennadius junctor Heinrich, new species.
98. Farnobia quadripuncta (Zeller), male.
99. Farnobia quadripuncta (Zeller), hind wing
of female.
344 UNITED
Fieures 100-110.—VENATION (MALE UNLESS
OTHERWISH NOTED).
100. Metephestia simplicula (Zeller).
101. Divitiaca ochrella Barnes and McDun-
nough.
102. Oedothmia endopyrella Hampson, figured
from a sketch, by J. F. G. Clarke, of
the type of its synonym Synothomia
bahamasella Hampson.
103. Diviana eudoreella Ragonot; 103a, fore-
wing showing variation in venation.
104. Palatka nymphaeella (Hulst).
105. Cacozophera venosa Dyar.
106. Stylobasis rubripurpurea Hampson.
107. Prosoeuzophera impletella (Zeller).
108. Ephestiodes gilvescentella Ragonot.
109. Ephestiodes plorella Dyar, Forewing of
its synonym Eurythmia vestilla Dyar.
110. Azaera muciella Schaus.
STATES NATIONAL MUSEUM BULLETIN 207
100. Metephestia
1 10 9
2
103. Diviana
107.
Prosoeuzophera,
Ol. Divitiaca
Se
102. Oedothmia
110. Azaera
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 345
=
112. Homoeosoma
SS
f
15.Vitula 116. Verina
117. Comotia
—————
119. Bema 2
~
™
120. Bema 2
300329—56——23
Figures 111-120.—VENATION (MALE UNLESS
OTHERWISE NOTED).
111. Harnocha velessa Dyar.
112. Homoeosoma sinuellum (Fabricius).
113. Manhatta biviella (Zeller), female.
114. Moodna ostrinella (Clemens).
115. Vitula edmandsae (Packard).
116. Verina supplicella (Dyar).
117. Comotia torsicornis Dyar.
118. Bema neuricella (Zeller), male.
119, 120. Bema neuricella (Zeller), females,
showing vestiges of vein 9 in forewings.
346 UNITED STATES NATIONAL MUSEUM BULLETIN 207.
Figures 121—-130.—VENATION (MALE UNLESS
OTHERWISE NOTED).
121. Unadilla erronella (Zeller).
122. Caudellia apyrella Dyar.
123. Bethulia championella Ragonot, female.
124. Plodia interpunctella (Hiibner).
125. Ephestia cautella (Walker).
126. Ephestia elutella (Hiibner).
127. Nicetiodes apianella Schaus.
126. Cabnia myronella Dyar.
129. Ribiria conops (Dyar).
130. Microphycita titillella Dyar.
121. Unadilla 122. Caudellia
ee
SS
123. Bethulia 24. Plodia
125. Z
Ephestia 126. Ephestia
Saaaaa—
127.
Nicetiodes
128. Cabnia
N.
130. Microphycita
2
129. Ribiria
1340. ew
134. indigenella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 347
‘134d,
Figures 131-134.—Mates.
131. Cryptoblabes rutilella Zeller, right harpe
denuded and aedeagus omitted; 131a,
vinculum, tegumen, and uncus; 131b,
harpe, denuded except for ventral tuft;
131c, aedeagus; 131d, gnathos; 13le,
transtilla and anellus; 131f, two views
(scaled and denuded) of basal segments
of antenna.
132. Cryptoblabes gnidiella (Millitre), left
harpe detached and aedeagus omitted;
132a, anellus; 132b, transtilla; 132c,
gnathos; 132d, aedeagus.
133. Acrobasis tumidella (Zincken); 133a,
gnathos; 133b, transtilla; 133c, anellus;
133d aedeagus; 133e, ventral tuft of
eighth abdominal segment.
134. Acrobasis indigenella (Zeller) ; 134a, aede-
agus; 134b, transtilla; 134c,d, basal
segments of antenna, denuded and
scaled.
348 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieurres 135-139.—Ma tess.
135. Acrobasis tricolorella Grote, 135a, aedea-
gus; 135b, gnathos; 135c, transtilla;
135d, ventral tuft of eighth abdominal
segment.
136. Acrobasis comacornella (Hulst), type;
136a, aedeagus; 136b, gnathos; 136c,
transtilla; 136d, anellus; 136e, ventral
tuft of eighth abdominal segment.
137. Acrobasis caryae Grote, specimen reared
from pecan nut, tuft of eighth ab-
dominal segment.
138. Acrobasis juglandis (LeBaron), specimen
Treared from pecan; 138a, aedeagus;
138b, gnathos; 138c, transtilla; 138d,
anellus.
139. Acrobasis latifasciella Dyar, type; 139a,
aedeagus.
iss encolarella 135c.
136.comacornella
136d.
138. juglandis 139, latifasciella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 349
142.
palliolella
lake, 142a.
144,
sylviella
14.6.
evanescentella,
149 a. 147a.
149. \
aurorella
151.
exsulella
152a.
Fiaures 140-154.—mates or AcROBASIS
SPECIES: UNCUS AND GNATHOS AND (a)
TRANSTILLA:
140.
141.
142,
143.
144,
145.
146.
147.
148.
149.
150.
151.
152.
153.
154.
Acrobasis minimella Ragonot, figured
from type of its synonym A. nigro-
signella Hulst.
Acrobasis feltella Dyar, type.
Acrobasis palliolella Ragonot, male from
Chicago, Ill.
Acrobasis caryalbella Ely, type.
Acrobasis sylviella Ely.
Acrobasis kearfottella Dyar, type.
Acrobasis caryae Grote, southern male,
reared from pecan nut.
Acrobasis evanescentella Dyar.
Acrobasis stigmella Dyar, reared specimen
from Falls Church, Va. (Hopk. U. S.
No. 12121d).
Acrobasis aurorella Ely, type.
Acrobasis peplifera Dyar, type.
Acrobasis exsulella (Zeller).
Acrobasis angusella Grote, figured from
type of its synonym A. eliella, Dyar.
Acrobasis demotella Grote, specimen
from New Brighton, Pa.
Acrobasis irrubriella Ely, type.
350 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 155-166.—Matzs.
155. Acrobasis normella Dyar, type, uncus
and gnathos; 155a, another male,
showing variation in apical process of
gnathos; 155b, type, transtilla.
156. Acrobasis malipennella Dyar, type, uncus
and gnathos; 156a, transtilla.
157. Acrobasis dyarella Ely, type, uncus and
gnathos; 157a, transtilla.
158. Acrobasis ostryella Ely, type, uncus and
gnathos; 158a, transtilla.
159. Acrobasis secundella Ely, type, uncus and
gnathos; 159a, transtilla.
160. Acrobasis coryliella Dyar, type, uncus
and gnathos; 160a, transtilla.
161. Acrobasis caryivorella Ragonot, specimen
reared on pecan at Austin, Tex.
uncus and gnathos; 161a, transtilla.
162. Acrobasis cunulae Dyar and Heinrich:
162-162a, Ventral and lateral views of
uncus and gnathos; 162b, apical proc-
ess of gnathos from a male showing
the broadest development in this
structure; 162c, transtilla.
163. Acrobasis myricella Barnes and McDun-
nough, type, uncus and gnathos; 163a,
transtilla.
164. Acrobasis betulella Hulst, uncus and
gnathos; 164a, transtilla.
165. Acrobasis rubrifasciella Packard, reared
male from New York, uncus and
enathos; 165a, transtilla.
166. Rhodophaea advenella (Zincken), aedea-
gus omitted; 166a, aedeagus; 166b,
transtilla; 166c, ventral tuft on eighth
abdominal segment.
ostryella
162.
162.
cunulae
163.
We, aN Ally
Wz ate C/I AS. art Wa
Ny Al ( a wi sy i SS S/
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 351
168.ochrodesma tN
169.
Ficures 167-170.— Mates.
167. Rhodophaea supposita (Heinrich), aedeag-
us omitted; 167a; aedeagus; 167b,
gnathos; 167c, transtilla.
168. Anabasis ochrodesma (Zeller), aedeagus
omitted; 168a, aedeagus; 168b, trans-
tilla; 168c, sternite and tergite of
eighth abdominal segment, showing
ventral tufts.
169. Mildrizia constitutionella Dyar; 169a,
aedeagus; 169b, anellus; 169c, gnathos;
169d, transtilla; 169e, sternite and
tergite of eighth abdominal segment,
showing ventral tuft; 169f, basal
segments of antenna, partially de-
nuded.
170. Cuniberta subtinctella (Ragonot); 170a,
aedeagus; 170b, gnathos; 170c, trans-
tilla; 170d, anellus.
352 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 171, 172.—Mates.
171. Sematoneura atrovenosella Ragonot, ae-
deagus omitted; 17la, aedeagus: 171b,
vinculum, tegumen, gnathos and uncus,
ventral view; 171c, transtilla; 171d,
anellus; 17le, ventral hair tuft of
eighth abdominal segment.
172. Sematoneura abitus Heinrich, new species,
type; 172a, aedeagus; 172b, gnathos;
172c, transtilla.
171. atrovenosello
172b.
172.abitus
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 353
173. a : | V y
\
300329—56——24
Ficurss 173, 174.—Matss.
173. Hypsipyla pagodella Ragonot; 173a,
aedeagus; 173b, gnathos; 173c, trans-
tilla; 173d, anellus; 178e, sternite,
tergite and ventrolateral tufts of
eighth abdominal segment.
174. Hypsipyla grandella (Zeller), aedeagus
omitted; 174a, aedeagus; 174b, gna-
thos; 174c, transtilla; 174d, sternite
and tergite of eighth abdominal seg-
ment.
354 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 175-177. MALEs.
175. Hypsipyla dorsimacula (Schaus); 175a,
aedeagus; 175b, gnathos; 175c, trans-
tilla.
176. Hypsipyla ferrealis (Hampson); 176a,
aedeagus; 176b, gnathos; 176c, trans-
tilla; 176d, anellus; 176e, eighth
abdominal segment, showing tufts and
modifications of sternite and tergite.
177. Hypsipyla fluviatella Schaus; 177a, ae- 175 x
deagus; 177b, sternite and tergite of e: Ss
eighth abdominal segment; 177c, gna- 175. dorsimacula
thos; 177d, transtilla; 177e, anellus.
“x \T1b.
MI
Me
WS
YY iy,
yy
NY
177. fluviatella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 355
178
chinographella
SZ
br
1798
A plumigerella
bigrana
Figures 178-180.—Matzs,
178. Hemiptilocera chinographella Ragonot;
178a, aedeagus; 178b, apex of aedea-
gus, showing bent and spined cornutus
in another view; 178c, transtilla; 178d,
modified sternite and hair tufts of
eighth abdominal segment; 178e, basal
segments of antenna.
179. Hemiptilocera plumigerella (Ragonot),
type; 179a, aedeagus; 179b, transtilla;
179c, sternite and tufts of eighth
abdominal segment.
180. Hemiptilocera bigrana (Zeller); 180a,
aedeagus; 180b, transtilla; 180c, tufts
and modified sternal plates of eighth
abdominal segment.
356 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 181-183.— Mats.
181. Crocidomera stenopteryx (Dyar), type;
18la, aedeagus.
182. Crocidomera turbidella Zeller, specimen
from Cuba, aedeagus omitted; 182a,
aedeagus; 182b, transtilla; 182c, an-
ellus; 182d, scale and hair tufts of
eighth abdominal segment.
183. Crocidomera fissuralis (Walker), speci-
men from Puerto Rico (in Cornell);
183a, aedeagus; 183b, transtilla; 183c,
anellus; 183d, scale and hair tufts of
eighth abdominal segment.
NIN
PE FAW
ey
183. fissuralis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 357
Figures 184—187.—Matss.
184. Heras disjunctus Heinrich, new species,
type; 184a, aedeagus; 184b, gnathos;
184c, transtilla; 184d, anellus.
185. Adanarsa intransitella (Dyar), aedeagus
omitted; 185a, aedeagus; 185b, anellus;
185c, gnathos; 185d, transtilla; 185e,
tufts on eighth abdominal segment.
186. Birinus russeolus Heinrich, new species,
type; 186a, aedeagus; 186b, gnathos;
186c, transtilla and anellus.
187. Bertelia grisella Barnes and McDun-
nough, type, aedeagus omitted; 187a,
aedeagus; 187b, transtilla; 187c, tufts
on eighth abdominal segment.
186.
russeolus
358 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 188-190.— Mates.
188. Hypargyria definitella (Zeller), aedeagus
omitted; 188a, aedeagus; 188b, gna-
thos; 188c, transtilla; 188d, anellus;
188e, eighth abdominal segment, show-
ing ventral scale tufts.
189. Chararica annuliferella (Dyar), aedeagus
omitted; 189a, aedeagus; 189b, par-
tially fused gnathos and transtilla;
189c, anellus.
190. Chararica hystriculella (Hulst), aedeagus
omitted; 190a, aedeagus; 190b, par-
tially fused gnathos and transtilla; ‘ 188. definitella
190c, anellus; 190d, hair tufts on
eighth abdominal segment.
189. annuliferella
190. hystriculella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
The
192. subtetricella
193. clatella
g
/195.
piazzella
195a.
359
Fiagures 191—195.—Mat.zs.
191, Myelopsis coniella (Ragonot); 19la,
aedeagus; 191b, gnathos; 191c, trans-
tilla; 191d, anellus.
192. Myelopsis subtetricella (Ragonot); 192a,
aedeagus; 192b, gnathos; 192c, trans-
tilla.
193. Myelopsis alatella (Hulst); 193a, aedea-
gus; 193b, gnathos; 193c, transtilla.
194. Myelopsis fragilella (Dyar), type,
synonym of M., alatella (Hulst); 194a,
transtilla.
195. Myelopsis piazzella (Dyar), type, @ syno-
nym of M. alatella (Hulst); 195a,
transtilla,
360 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 196-199.—Matss.
196. Anypsipyla wnivitella Dyar, aedeagus
omitted; 196a, aedeagus, 196b; trans-
tilla; 196c, ventrolateral tufts on
eighth abdominal segment.
197. Apomyelois bistriatella (Hulst); 197a,
aedeagus; 197b, gnathos; 197c, trans-
tilla.
198. Ectomyelois decolor (Zeller), aedeagus
omitted; 198a, aedeagus; 198b, trans-
tilla.
199. Ectomyelois ceratoniae (Zeller), aedeagus
omitted; 199a, aedeagus; 199b, trans-
tilla.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 361
Ficurrs 200-203.—Matszs.
200. Hctomyelois muriscis (Dyar), aedeagus
omitted; 200a, aedeagus; 200b, trans-
tilla.
201. Ectomyelois zeteki Heinrich, new species,
type; 201a, aedeagus.
202. Paramyelois transitella (Walker); 202a,
aedeagus; 202b, gnathos; 202c, ele-
ments of divided transtilla; 202d,
anellus.
203. Myelois cribrella (Hiibner), aedeagus
omitted; 203a, aedeagus; 203b, trans-
tilla.
SS ee
wy PP
203b.
362 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 204-209.— Mates.
204, Pseudodivona commensella Dyar, type,
aedeagus omitted; 204a, aedeagus;
204b, elements of transtilla; 204c, ter-
gite, sternite, and ventrolateral tufts
of eighth abdominal segment.
205. Pseudodivona cispha Dyar, type, harpe;
, . 205a, uncus; 205b, gnathos.
206. Pseudodivona carabayella Dyar, type,
uncus; 206a, specimen from Incachaca,
Bolivia, gnathos.
207. Protomoerbes separabilis Heinrich, new
species, type; 207a, aedeagus; 207b,
tergite and sternite of eighth abdom-
inal segment.
208. Protomoerbes aberrans Heinrich, new
species, type; 208a, aedeagus; 208b,
elements of transtilla.
209. Diatomocera tenebricosa (Zeller); 209a,
transtilla. (See also fig. 565.)
206a.
carabayella
ww
209.tenebricosa
AMERICAN MOTHS OF THE SUBFAMILY
210.dosia
Diz dectecens
213.
majuscula
PHYCITINAE 363
Ficurs 210-213.—Ma tes.
210. Diatomocera dosia (Dyar), type; 210a,
aedeagus; 210b, hair tufts of eighth
abdominal segment.
211. Diatomocera excisalis (Hampson), speci-
men from St. Laurent Maroni, French
Guiana (in USNM); 211a, aedeagus.
212. Diatomocera decurrens (Dyar), type;
212a, aedeagus.
213. Diatomocera majuscula Heinrich, new
species, type; 213a, aedeagus; 213b,
transtilla; 213c, tergite, sternite, and
tufts of eighth abdominal segment.
364 UNITED STATES NATIONAL: MUSEUM BULLETIN 207
Figures 214-219.—Matss.
214. Diatomocera albosigno Heinrich, new
species, type, one harpe omitted; 214a,
transtilla; 214b, tergite, sternite, and
hair tufts of eighth abdominal seg-
ment.
215. Diatomocera hoplidice (Dyar), type; 215a, 1 UNS
aedeagus. Wy,
216. Diatomocera mochlophleps (Dyar), type;
216a, aedeagus.
217. Diatomocera extracta Heinrich, new spe-
cies, type; 217a, aedeagus.
218. Pseudocabima castronalis Heinrich, new
species, type; 218a, aedeagus.
219. Pseudocabima fearnella (Schaus), type;
219a, aedeagus.
214. albosigno
211.
extracta
216.”
mochlophleps
castronalis
219. fearnella
224%
expunctrix
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 365
225.
rubrizonalis
220.
221,
222.
223.
224,
225.
Ficures 220-225.— Mates.
Pseudocabima guianalis Heinrich, new
species, type; 220a, aedeagus.
Pseudocabima pombra (Dyar), type, un-
cus.
Pseudocabima nigristrigella (Ragonot),
type; 222a, aedeagus; 222b, tergite,
sternite, and hair tufts of eighth ab-
dominal segment.
Pseudocabima euzopherella (Dyar); 223a,
aedeagus.
Pseudocabima expunctrix Dyar and Hein-
rich, type; 224a, aedeagus.
Pseudocabima rubrizonalis (Hampson),
“cotype” (in USNM) from Cayenne,
French Guiana; 225a, aedeagus.
366 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 226-230—MatgEs.
226. Hyalospila stictoneurella Ragonot; 226a,
aedeagus; 226b, transtilla; 226c, an-
ellus; 226d, tergite, sternite, and hair
tufts of eighth abdominal segment.
227. Hyalospila celiella Schaus, type; 227a,
aedeagus; 227b, transtilla; 227c, an-
ellus.
228. Hyalospila fulgidula Heinrich, new spe- ig
cies, type; 228a, aedeagus; 228b, trans- 226.
tilla; 228c, anellus. 2260. reves Sorelle
229. Hyalospila xsanthoudemia (Dyar), type;
229a, aedeagus; 229b, transtilla; 229c,
anellus; 229d, tergite, sternite, and hair
tufts of eighth abdominal segment.
230. Hyalospila clevelandella (Dyar), type;
230a, aedeagus; 230b, transtilla; 230c,
anellus; 230d, tergite, sternite, and hair
tufts of eighth abdominal segment.
ee,
228.
fulgidula
229. xanthoudemia
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 367
Fiaurres 231-233.—Ma tgs.
231. Fundella pellucens Zeller, aedeagus omit-
ted; 231a, aedeagus; 231b, lateral view
of uncus, gnathos, and tegumen; 231c,
tergite, sternite, and hair tufts of
eighth abdominal segment; 231d, basal
segments of antenna, scaled; 23le,
basal segments of male antenna, de-
nuded; 231f, hind tibia, showing hair
tuft.
232. Fundella ignobilis Heinrich, new species,
aedeagus omitted; 232a, aedeagus;
232b, anellus; 232c, lateral view of
uncus, gnathos, and tegumen.
233. Fundella ahemora Dyar, aedeagus omit-
ted; 233a, aedeagus; 233b, lateral view
of uncus, gnathos, and tegumen; 233c,
foretibia, showing hair tuft.
HAO ue
"
< ig
ahemora
368 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficgures 234-238.—Matus.
234. Fundella argentina Dyar, aedeagus) omit-
ted; 234a, aedeagus; 234b, apical
projection of gnathos, showing the
extreme of variability in this structure;
234c, hair tufts on eighth abdominal
segment.
235. Difundella corynophora Dyar, aedeagus
omitted; 235a, aedeagus; 235b, eighth
abdominal segment, showing sclero-
tized pocket developed from sternite.
236. Difundella subsutella (Schaus), type,
aedeagus omitted; 236a, aedeagus.
237. Difundella distractor Heinrich, new spe-
cies, both harpes and aedeagus omit-
ted; 237a, aedeagus; 237b, harpe, 237c,
sclerotized parts of eighth abdominal
segment, showing sclerotized pocket of
sternite.
238. Difundella tolerata Heinrich, new species,
type, both harpes, anellus, and aedea-
gus omitted; 238a, aedeagus; 238b,
anellus; 238c, harpe; 238d, scaled
pocket between second and third
segments of abdomen.
237. distractor
238 .tolerata
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 369
24a.
243a.
Figures 239-243.—Ma tgs.
239. Coplarthria dasypyga (Zeller); 239a,
aedeagus; 239b, anellus; 239c, sclero-
tizations of eighth abdominal segment,
showing development from sternite.
240. Promylea lunigerella Ragonot, specimen
from Duncans, Vancouver Island,
aedeagus omitted; 240a, aedeagus;
240b, anellus; 240c, sclerotizations of
eighth abdominal segment.
241. Promylea lunigerella glendella (Dyar);
241a, aedeagus; 241b, sclerotizations
of eighth abdominal segment.
242. Promylea mindosis Dyar, type; 242a,
aedeagus; 242b, sclerotizations of
eighth abdominal segment.
243. Promylea dyari Heinrich, type; 248a,
aedeagus.
370 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 244-247.—Matsgs.
244. Anadelosemia senesciella (Schaus), type,
one harpe and aedeagus omitted and \
one harpe detached; 244a, aedeagus;
244), anellus; 244c, sclerotization of
eighth abdominal segment.
245. Anadelosemia condigna Heinrich, new
species, type; 245a, aedeagus; 245b, ue
sclerotization of eighth abdominal seg- Gi.
ment.
246. Anadelosemia dulciella (Hulst); a syno- 244, senesciella
nym of A. texanella; 246a, aedeagus;
246b, anellus; 246c, sclerotization of
eighth abdominal segment.
247: Dasypyga alternosquamella Ragonot,
aedeagus omitted; 247a, aedeagus;
247b, combined gnathos and anellus;
247c, sclerotization of eighth abdomi-
nal segment.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 371
Ficurrs 248-250.—Matzs.
248. Rampylla orio Dyar, type; 248a, dorsal
view of uncus, gnathos and tegumen;
248b, aedeagus; 248c, anellus; 248d,
harpe; 248e, sclerotization of eighth
abdominal segment.
249. Rampylla subcaudata (Dyar), type; 249a,
aedeagus; 249b, sclerotization of eighth
abdominal segment.
250. Rampylla lophotalis Heinrich, new spe-
cies; 250a, aedeagus; 250b, harpe;
250c, transtilla; 250d, anellus; 250e,
sclerotization of eighth abdominal seg-
ment; 250f, uncus; 250g, tegumen;
250h, vinculum.
250.5.
lophotalis
372 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 251-254.—Mates.
251. Fulrada querna (Dyar), type; 25la,
aedeagus; 251b, sclerotization and tuft
of eighth abdominal segment.
252. Fulrada carpasella (Schaus), type; 252a,
aedeagus; 252b, anellus; 252c, scale
tufts and sclerotization of sternite of
eighth abdominal segment.
253. Scorylus cubensis Heinrich, new species;
253a, aedeagus; 253b, sternite and
tufts of eighth abdominal segment.
254. Davara caricase (Dyar), aedeagus omit-
ted; 2548, aedeagus; 254b, anellus; 254c,
sternite of eighth abdominal segment;
254d, basal segments of antenna.
eS
253. cubensis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 373
Figures 255-259.—Matzs.
255. Davara paranensis (Dyar), type; 255a,
aedeagus.
256. Davara interjecta Heinrich, new species;
256a, aedeagus.
257. Davara rufulella (Ragonot), aedeagus
omitted; 257a, aedeagus; 257b, forked
spine associated with anellus; 257e,
anellus; 257d, sternite and tufts of
eighth abdominal segment.
258. Sarasota plumigerella Hulst, type, aede-
agus omitted; 258a, aedeagus; 258b,
sternite of eighth abdominal segment.
259. Sarasota furculella (Dyar), aedeagus
omitted; 259a, aedeagus; 259b, anellus;
259c, sternite of eighth abdominal
segment.
259.furculella
374 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 260—264.— Mates. \ ; i
260. Piesmopoda rubicundella Zeller, type;
260a, aedeagus.
261. Piesmopoda xanthopolys Dyar, specimen
from La Chorrera, Panamd; 261a,
aedeagus; 261b, dorsal view of uncus
and tegumen.
262. Piesmopoda flavicans (Zeller), figured
from type of its synonym P. fratella;
262a, aedeagus.
263. Piesmopoda parva Heinrich, new species,
type; 263a, aedeagus. 7. ss Bees =
264. Piesmopoda isabella (Dyar), type; 264a, SS ae 26 ail
ae 260. rubicundella \ aa 261.xanthopolys
aedeagus; 264b, anellus; 264c, sternite
of eighth abdominal segment.
264.isabella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 875
——————
268.almonella
sf \ SS ;
266. — ey,
Figures 265—268.— Mauss.
265. Piesmopoda ragonoti (Dyar), type; 265a,
aedeagus.
266. Piesmopoda sxanthomera Dyar, figured
from the type of its synonym P.,
zanthozona Dyar; 266a, aedeagus.
267. Atheloca subrufella (Hulst), figured from
type of its synonym Hyalospila ptychis
Dyar, aedeagus omitted; 267a, aede-
agus; 267b, anellus and fusing arms of
gnathos; 267c, harpe; 267d, sternite of
eighth abdominal segment.
268. Praedonula almonella (Dyar), type, aede-
agus omitted; 268a, aedeagus; 268b,
anellus; 268c, transtilla and gnathos;
268d, sternite of eighth abdominal
segment,
376 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 269-271. Matss.
269. Peadus burdettellus (Schaus), type; 269a,
aedeagus; 269b, anellus; 269c, ventral
view of (detached) left harpe; 269d,
ventral view of left harpe, denuded;
269e, sternite and tuft of eighth ab-
dominal segment.
270. Peadus dissitus Heinrich, new species,
type; 270a, aedeagus.
271. Gabinius paulsoni (Ragonot), type; 271a,
aedeagus; 271b, dorsal view of uncus,
tegumen, and vinculum; 271c, trans-
tilla; 271d, anellus; 27le, sternite of
eighth abdominal segment.
270a.
270.dissitus
271. pauilsoni
27\b.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 377
Ficures 272-274.—Matss.
272. Ceracanthia mamella (Dyar), specimen
from type locality, aedeagus omitted;
272a, aedeagus; 272b, sternite of
eighth abdominal segment.
273. Ceracanthia vepreculella Ragonot, type;
273a, aedeagus; 273b, gnathos; 273c,
anellus; 273d, sternite of eighth ab-
dominal segment.
274. Megarthria peterseni (Zeller), specimen
from Guatemala; 274a, aedeagus;
274b, anellus; 274c, ventrolateral tufts
and sternite of eighth abdominal seg-
. ment, showing two views of the bent,
272.mamella : digitate pocket of sternite.
=) 274. peterseni
800329—56——25
378 UNITED STATES NATIONAL MUSEUM BULLETIN 207
YieuREs 275—-281.— Mates.
275. Megarthria squamifera Heinrich, new
- species; 275a, aedeagus; 275b, gnathos.
276. Megarthria schaust Heinrich, new species;
276a, aedeagus; 276b, gnathos; 276c,
transtilla; 276d, anellus.
277. Megarthria cervicalis Dyar, type; 277a,
aedeagus; 277b, anellus; 277c, sternite
and ventral tuft of eighth abdominal
segment.
278. Ceracanthia mamella (Dyar), basal seg-
ments of antenna, scaled and denuded.
279. Megarthria peterseni (Zeller), basal seg-
ments of antenna, scaled and denuded.
280. Megarthria squamifera Heinrich, new
species, basal segments of antenna,
scaled and denuded.
281. Megarthria cervicalis Dyar, type, basal
segments of antenna, scaled.
275. Squamifera
cervicalis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 379
284.drucella
Figures 282—284.—MA.gEs.
282. Drescoma cyrdipsa Dyar, type, aedeagus
omitted; 282a, aedeagus; 282b, trans-
tilla; 282c, sternite of eighth abdom-
inal segment,
283. Drescoma ciniliza Dyar, type, aedeagus
omitted; 283a, aedeagus; 283b, trans-
tilla; 283c, sternite of eighth abdominal
a segment.
284. Drescoma drucella Dyar, a synonym of
Drescomopsis soraella (Druce), aedea-
gus omitted; 284a, aedeagus; 284b,
transtilla; 284c, sternite and scale
tufts of eighth abdominal segment.
380 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fi@ures 285-287.—Matss.
285. Monoptilota pergratialis (Hulst), geni-
talia figured from type of its synonym
M. nubilella Hulst; 285a, aedeagus;
285b, elements of transtilla; 285c,
anellus; 285d, lateral view of gnathos;
285e, ventral view of gnathos; 285f,
sclerotization and tuftings of eighth
abdominal segment of male; 285g,
basal segments of male antenna,
denuded; 285h, basal segments of male
antenna, scaled.
286. Zamagiria dixolophella Dyar, type, aedea-
gus omitted and tegumen, uncus, and
gnathos slightly turned to show them
in ventrolateral view; 286a, aedeagus;
286b, male genitalia shown in full
ventral view with both harpes and
aedeagus omitted; 286c, compound
tufts of eighth abdominal segment of
abdomen.
287. Zamagiria pogerythrus Dyar; 287a, aedea-
gus.
287. pogerythrus
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 381
Ficurzis 288-290.—Matgs.
288. Zamagiria hospitabilis Dyar, type, aedea-
gus omitted; 288a, aedeagus; 288b,
anellus.
289. Zamagiria masculinus Dyar, type, aedea-
gus omitted; 289a, aedeagus.
. 290. Zamagiria laidion (Zeller), aedeagus
omitted ; 290a, aedeagus; 290b, anellus.
290. laidion 290b.
382 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 291-294.—Matszs.
291. Zamagiria fraterna Heinrich, new species;
291a, aedeagus.
292. Zamagiria australella (Hulst); 292a,
aedeagus.
293. Magiriopsis denticosella (Dyar), one harpe
and aedeagus omitted and uncus and
gnathos bent to show lateral view;
293a, aedeagus.
294. Anegcephalesis arctella (Ragonot); 294,
aedeagus; 294b, harpe, ventral view;
294c, compound tufts of eighth abdom-
inal segment.
293. denticosel
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 383
Ficures 295-298.—Matss.
295. Ancylostomia stercorea (Zeller), aedeagus
omitted; 295a, aedeagus; 295b, com-
pound tufts of eighth abdominal seg-
ment.
296. Ancylostomia sauciella (Zeller), type; 296a,
aedeagus; 296b, dorsal view of uncus.
297. Caristanius pellucidellus (Ragonot), aede-
agus omitted and right harpe detached;
297a, aedeagus; 297b, anellus; 297c,
compound tufts of eighth abdominal
segment.
298. Caristanius decoloralis (Walker), aede-
agus omitted; 298a, aedeagus; 298b.
anellus,
384 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 299-305.—Matss.
299. Pima boisduvaliella (Guénée), anellus; 299. boisduvaliella
299a, apical portion of harpe; 299b, fi
cornuti. 302.vividella
300. Pima fosterella Hulst, anellus; 300a, api-
cal portion of harpe; 300b, cornuti.
301. Pima albocostalialis (Hulst), apical por-
tion of harpe; 301a, cornuti.
302. Pima vividella (McDunnough), paratype
from Saskatoon, Saskatchewan (in
USNM); anellus; 302a, apical portion
of harpe; 302b, cornuti.
303. Pima albiplagiatella occidentalis Heinrich:
New race, anellus.
304. Pima fulvirugella (Ragonot), type; 304a,
apical portion of harpe; 304b, cornuti.
305. Pima albiplagiatella (Packard) ; 305a, ae-
deagus; 305b, anellus; 305c, ventral
view of right harpe; 305d, tufts of
eighth abdominal segment.
301.
albocostalialis
a
7
305. |
albiplagiatella
AMERICAN MOTHS OF THE SUBFAMILY
308. denticulella
307. granitella
300329—36——26
PHYCITINAE 385
Ficures 306-308.—Ma.zs.
306. Pima parkerella (Schaus), type; 306a, ae-
deagus; 306b, ventral view of harpe.
307. Pima granitella (Ragonot), figured from
type of its synonym Megasis piperella
Dyar; 307a, aedeagus; 307b, anellus;
307c, ventral view of right harpe.
308. Interjectio denticulella (Ragonot), speci-
men from British Columbia, aedeagus
omitted and right harpe detached and
shown in ventral view; 308a, aedeagus;
308b, anellus.
386 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieurres 309-311.—Mates.
309. Interjectio niviella (Hulst), aedeagus
omitted and right harpe detached and
shown in ventral view; 309a, aedeagus.
310. Ambesa laetella Grote, aedeagus omitted;
310a, aedeagus; 310b, tufts of eighth
abdominal segment.
311. Ambesa lallatalis (Hulst); 311a, aedeagus;
311b, ventral view of right harpe; 311c,
anellus.
<—S
‘
By FLT
A WR + Ah MY i:
ANZ SAU
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 387
312.
313.
314.
315.
316.
Ficures 312-316.— Mates.
Ambesa walsinghami (Ragonot), speci-
men from Plumas County, Calif.;
312a, aedeagus; 312b, elements of
transtilla; 312¢, apical portion of harpe.
Ambesa walsinghami mirabella Dyar,
apical portion of harpe.
Catastia marginea (Schiffermiiller): aede-
agus omitted; 3l4a, aedeagus; 314b,
transtilla.
Catastia incorruscella (Hulst), type; 315a,
aedeagus; 315b, gnathos; 315c, ele-
ments of transtilla; 315d, anellus; 315e,
tufts and sclerotization of eighth ab-
dominal segment.
Catastia bistriatella (Hulst), specimen
from Humphreys Basin, Calif.; 316a,
aedeagus.
388 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 317-321.—Matzs.
317. Catastia actualis (Hulst) ; 317a, aedeagus;
317b, elements of transtilla.
318. Immyrla nigrovitiella Dyar, aedeagus
omitted; 318a, aedeagus; 318b, com-
pound tufts of eighth abdominal seg-
ment.
319. Oreana unicolorella (Hulst), two views of
uncus and gnathos; 319a, aedeagus;
319b, anellus.
320. Olybria aliculella (Hulst) ; 320a, aedeagus.
321. Olybria furctferella (Dyar), type; 321a,
aedeagus.
320.
aliculella < furciferella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 389
Figures 322-325.—Matzs,
322. Salebriacus odiosellus (Hulst); 322a,
aedeagus.
323. Salebriaria turpidella (Ragonot), figured
from type of its synonym Salebria
ademptandella Dyar; 323a, aedeagus;
323b, anellus.
324. Salebriaria fructetella (Hulst), figured
from type of its synonym Salebria
rectistrigella Dyar; 3242, aedeagus;
324b, anellus.
325. Salebriaria pumilella (Ragonot), figured
from type of its synonym Salebria
georgiella Hulst; 325a, aedeagus; 325b,
gnathos; 325c, elements of transtilla;
325d, anellus; 325e, compound tufts of
eighth abdominal segment.
fructetella:
S 2) 325,
\ if pumilella
325d.
390
326.
327.
328.
329.
330.
331.
332,
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 326-332.—Matss.
Etiella zinckenella (Treitschke); 326a,
ventral view of harpe; 326b, aedeagus.
Glyplocera consobrinella (Zeller), aedeagus
omitted; 327a, aedeagus; 327b, trans-
tilla.
Quasisalebria admixta Heinrich, type;
328a, aedeagus.
Oriholepis jugosella Ragonot; 329a, aede-
agus.
Polopeustis annulatella (Zetterstedt) ;
330a, aedeagus.
Polopeustis arctiella (Gibson), aedeagus.
Salebria palumbella (Schiffermiller);
332a, aedeagus.
331.
arctiella
. ~ ZB
~ 330.annulatella
327b.
332.palu
i. 327. consobrinella
mbella.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 391
Freurss 333-336.— Matzs.
333. Meroptera mirandella Ragonot, aedeagus
omitted; 333a, aedeagus; 333b, trans-
tilla.
334. Meroptera pravella (Grote), specimen
from Edmonton, Alberta; 334a, aede-
agus.
335. Meroptera abditiva Heinrich, new species,
type; 335a, aedeagus,
336. Nephopteryz rhenella (Zincken), aedeagus
omitted; 336a, aedeagus; 336b, basal
segments of antenna, denuded.
392 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 337-342.—MaA.zs.
337. Nephopteryz subfuscella (Ragonot), ae-
deagus omitted; 337a, aedeagus; 337b,
transtilla.
338. Nephopteryx dammersi Heinrich, new
species, aedeagus.
339. Nephopteryz subcaesiella (Clemens), ae-
deagus.
340. Nephopteryz fernaldi (Ragonot), speci-
men from Colorado, aedeagus omitted;
340a, aedeagus.
341. Nephopteryz basilaris Zeller; 34la, ae-
deagus.
342. Nephopteryx termitalis (Hulst), type;
3428, aedeagus.
339. \
subcaesiella 340. fernaldi
320 teematalis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 393
Molt
344d-
346.
rubrisparsella
343.
344,
345.
346.
347.
348.
Fiaures 343-348.—Matgzs.
Nephopteryzx bifasciella Hulst, type; 348a,
aedeagus.
Nephopteryx uvinella (Ragonot); 344a,
aedeagus; 344b, elements of trans-
tilla; 344c, gnathos; 344d, anellus;
344e, compound tufts of eighth ab-
dominal segment.
Nephopteryx celtidella (Hulst); 345a,
aedeagus.
Nephopteryz rubrisparsella (Ragonot),
figured from type of its synonym
texanella Hulst, 346a, aedeagus; 346b,
gnathos; 346c, transtilla; 346d, anellus.
Nephopteryz crassifasciella Ragonot, one
harpe omitted, one detached, and
aedeagus omitted; 347a, aedeagus.
Tlascala reductella (Walker), figured from
type of its synonym Pempelia gledit-
schiella Fernald, aedeagus omitted;
348a, aedeagus; 348b, compound ven-
tral tufts of eighth abdominal segment.
394 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 349-352.—Matzs.
349. Tulsa finitella (Walker), specimen from
Florida, harpe; 349a, aedeagus.
350. Tulsa umbripennis (Hulst), figured from
type of its synonym Ortholepis gillet-
tella Dyar; 350a, aedeagus; 350b,
harpe; 350c, compound tufts of eighth
abdominal segment.
351. Tulsa oregonella (Barnes and MeDun-
nough), type; harpe; 351a, aedeagus.
352. Homeographa lanceolella Ragonot, type,
one harpe detached and aedeagus
omitted; 352a, aedeagus; 352b, gna-
thos; 352c, anellus; 352d, compound
tufts of eighth abdominal segment.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 395
354
355
356
357
i 355.
, curvatellus
4d.
Bhi bcucei
356b.
3538.
Fieurns 353-357.— Ma tzs.
Telethusia ovalis (Packard), aedeagus
omitted; 353a, aedeagus.
. Phobus brucei (Hulst), type; 354a,
aedeagus; 354b, elements of trans-
tilla; 354c, anellus; 354d, gnathos;
354e, sclerotizations aud tufts of
eighth abdominal segment.
. Phobus curvatellus (Ragonot), anellus.
. Phobus incertus Heinrich, new species;
356a, aedeagus; 356b, gnathos; 356c,
anellus.
. Actriz nyssaecolella (Dyar), type; 357a,
aedeagus; 357b, anellus; 357c, gnathos;
357d, uncus.
396 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 358-361.—Matzs.
358. Aciriz dissimulatriz Heinrich, type; 358a,
aedeagus, 358b, gnathos; 358c, anellus.
359. Stylopalpia lunigerella Hampson, aedea-
gus omitted; 359a, aedeagus; 359b,
transtilla; 359c, tufts of eighth abdomi-
nal segment.
360. Stylopalpia scobiella (Grote), 360a, aedea-
gus; 360b, vestiges of transtilla; 360c,
anellus.
361. Siylopalpia argentinensis Heinrich, type;
361a, aedeagus.
358. dissimulatrix
361. argentinensis
364. impostor
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 397
366.aequivoca
362.
363.
364.
365.
366.
Figures 362—366.—Ma tgs.
Pyla fasciolalis (Hulst), type; 362a,
aedeagus; 362b, anellus.
Pyla fasciolalis (Hulst), variety with
shorter harpe and vinculum.
Pyla impostor Heinrich, new species;
364a, aedeagus.
Pyla insinuatriz Heinrich, new species,
type; 365a, aedeagus.
Pyla aequivoca Heinrich, new species,
type; 366a, aedeagus.
398 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 367—370.—Ma.ss.
367. Pyla aenigmatica Heinrich, new species;
genitalia dissected and one harpe,
aedeagus, and anellus omitted; 367a,
anellus; 367b, aedeagus; 367c, tufts of
eighth abdominal segment.
368. Pyla criddlella Dyar, type, aedeagus
omitted; 368a, aedeagus.
369. Pyla fusca (Haworth), aedeagus omitted;
369a, aedeagus; 369b, tufts of eighth
abdominal segment.
370. Pyla hypochalciella (Ragonot), figured
from type of its synonym P. blackmor-
ella Dyar, aedeagus omitted; 370a,
aedeagus; 370b, dorsal view of uncus,
tegumen, and vinculum; 370c, anellus.
7
7 (43100.
hypochalciclla
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 399
Fieures 371-375.— Ma.gs.
371. Pyla hanhamella Dyar, type, aedeagus
omitted; 371la, aedeagus.
372. Pyla scintillans (Grote), specimen from
Tuolumne Meadows, Calif., aedeagus
omitted; 372a, aedeagus; 372b, hair
tufts of eighth abdominal segment.
373. Pyla scintillans (Grote), figured from
type of its synonym, P. feella Dyar,
aedeagus omitted; 373a, aedeagus.
374. Pyla rainierella Dyar, type, aedeagus
puma ae Ky" i omitted; 374a, aedeagus.
; 375. Pyla sylphiella Dyar, type, aedeagus
pared bY omitted; 375a, aedeagus.
\
Py sen Dr :
rene ERT
id
375. sylphiella
374. rainierella
400 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 376-381.— Ma tes.
376. Pyla aeneella Hulst, left harpe deformed,
tight harpe detached, and aedeagus
omitted; 376a, aedeagus.
377. Pyla metalicella Hulst: Ventral view of
unecus, gnathos, tegumen, and vincu-
lum; 377a, anellus and harpe; 377b,
aedeagus.
378. Pyla aeneoviridella Ragonot, aedeagus
omitted; 278a-b, two variations of the
aedeagus.
379. Pyla fasciella Barnes and McDunnough,
aedeagus omitted; 379a, aedeagus.
380. Pyla nigricula Heinrich, new species,
aedeagus omitted; 380a, aedeagus.
381. Pyla viridisuffusella Barnes and Mc-
Dunnough, one harpe detached and
one harpe and aedeagus omitted; 381a,
aedeagus.
381.
viridisuffusella
i
380.nigricula
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 401
tks j , i
iia
He
epee:
Oe
(
or
Ne
ca
Fieures 382—385.—Mauus
382. Dioryctria abietella (Denis and Schiffer-
miiller); 382a, aedeagus; 382b, uncus,
gnathos, and tegumen of a European
example; 383c-e, variations in the
costo-terminal part of harpe in Amer-
can specimens; 383f, compound tufts
on eighth abdominal segment.
383. Dioryctria reniculella Grote; 383a, aedea-
gus.
384. Dioryctria ponderosae Dyar, type; 384a,
aedeagus.
385. Dioryctria majorella Dyar; figured from
type of its synonym D. muellerana
Dyar; 385a, aedeagus.
402 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 386-389.—MaAteEs.
386. Dioryctria auranticella (Grote) ; 386a, ae-
deagus.
387. Dioryctria erythropasa (Dyar), type; 3872,
aedeagus.
388. Dioryctria pygmaeella Ragonot; 388a, ae-
deagus.
389. Dioryctria zimmermani (Grote), typical
eastern example; 389a, aedeagus.
=) 386. auranticella
388.pygmacella
\
NN J 389. zimmermani
393. subtracta
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 403
Figures 390-394.—Matgs.
. Dioryctria cambiicola (Dyar), type; 390a,
uncus, with lateral margins flattened;
390b, aedeagus.
. Dioryctria baumhoferi Heinrich, new spe-
cies, type; 391a, aedeagus.
. Dioryctria gulosella (Hulst), uncus.
. Dioryctria subtracta Heinrich, new spe-
cies, type; 393a, aedeagus.
. Dioryetria clarioralis (Walker), typical
Florida specimen; 394a, aedeagus.
404 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieurts 395-398.—MALgEs.
395. Oryctometopia fossulatella Ragonot, one
harpe detached and aedeagus omitted;
395a, aedeagus; 395b, gnathos; 395c,
eighth abdominal segment, showing
hair tufts.
396. Sarata edwardsialis (Hulst), type, one
harpe detached and aedeagus omitted;
396a, aedeagus; 396b, clasper of harpe
(small example), elements of transtilla,
and anellus.
397. Sarata pullatella (Ragonot), uncus,
gnathos, and tegumen of type; 397a,
aedeagus of type; 397b, uncus, gnathos,
and tegumen of example from Utah;
397c, aedeagus of example from Utah;
397d, elements of transtilla and anellus.
398. Sarata punctella (Dyar), uncus, gnathos,
and tegumen; 398a, elements of trans-
tilla; 398b-d, variations in clasper of
harpe; 398e, anellus; 398f, aedeagus.
396b,
a
396. edwacdsialis:
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
4.02.dnopherella
402a.
x A
AWAY
eT i vy
BAS 404. rubrithoracella
}405.tephrella
399.
400.
401.
402.
403.
404.
405.
405
Fiaures 399-405.— Mates.
Saraia incanella (Hulst), aedeagus
omitted; 399a, aedeagus; 399b, anellus.
Sarata atrella (Hulst), uncus, gnathos,
and tegumen; 400a, clasper of harpe;
400b, aedeagus; 400c, elements of
transtilla and anellus.
Sarata caudellella (Dyar), type, uncus,
gnathos, and tegumen; 401a, clasper of
harpe, elements of transtilla and
anellus; 401b, aedeagus.
Sarata dnopherella Ragonot, type, uncus,
gnathos, and tegumen; 402a, elements
of transtilla and anellus; 402b, clasper
of harpe; 402c, aedeagus.
Sarata nigrifasciella Ragonot, type,
uncus, gnathos, and tegumen; 403a,
aedeagus; 403b, clasper of harpe;
403c, anellus and elements of transtilla.
Sarata rubrithoracella (Barnes and
MecDunnough), type; 404a, aedeagus.
Saraia tephrella Ragonot, type; 405a, ae-
deagus; 405b, eighth abdominal seg-
ment, showing ventrolateral tufts.
406 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficurrs 406—410.—Mauzs.
406. Philodema rhoiella (Dyar), paratype from
type locality; 406a, ventral view of
right harpe; 406b, anellus; 406c, aede-
agus.
407. Lipographis fenestrella (Packard); 407a,
aedeagus; 407b, tergite and sternite of
eighth abdominal segment.
408. Lipographis truncatella (Wright), speci-
men from Chula Vista, Calif., 408a,
aedeagus; 408b, elements of transtilla
and anellus; 408¢, tergite and sternite
of eighth abdominal segment.
409. Lipographis subosseella Hulst, type; 409a,
gnathos; 409b, aedeagus; 409c, anellus;
409d, elements of transtilla.
410. Lipographis wmbrella (Dyar), type; 410a,
aedeagus.
406. rhoiella
ag /\N
410. umbrella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 407
Figures 411-415.—Matrs.
411. Adelphia petrella (Zeller), aedeagus
omitted; 411la, aedeagus; 411b, anellus;
41le, tufts of eighth abdominal seg-
ment.
412. Adelphia ochripunctella (Dyar), type;
412a, aedeagus; 412b, ventral view of
harpe; 412c, anellus; 412d, gnathos.
413. Tota galdinella (Schaus), aedeagus omit-
ted; 413a, aedeagus; 413b, transtilla;
413c, anellus; 413d, tufts of eighth
abdominal segment.
414. Ufa lithosella (Ragonot) ; 414a, aedeagus.
415. Ufa senia Heinrich, new species, type;
415a, aedeagus.
ZA
411. petrella \
408 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 416—418.— Mass.
416. Ufa rubedinella (Zeller); 4162, aedeagus;
416b, harpe; 416c, anellus; 416d, com-
pound tufts of eighth abdominal seg-
ment.
417. Elasmopalpus lignosellus (Zeller), aede-
agus omitted; 417a, aedeagus; 417b,
tufts of eighth abdominal segment,
ventral view; 417c, lateral view of
tufts of eighth abdominal segment.
418. Acroncosa albiflavella Barnes and Mc-
Dunnough; 4182, aedeagus, dorsal
view; 418b, aedeagus, ventral view;
418c-e, cornuti of penis, much en-
larged; 418f, ventrolateral tufts of
eighth abdominal segment; 418g, an-
ellus.
418b. 4.18. albiflavella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 409
Figures 419-425.— Matgs.
419. Passadena flavidorsella (Ragonot); 419a,
aedeagus; 419b, sclerotizations and
tufts of eighth abdominal segment;
419c, elements of transtilla and anellus;
419d, gnathos; 419e, clasper of harpe.
420. Ulophora groteit Ragonot, aedeagus omit-
ted; 420a, aedeagus; 420b, elements of
transtilla; 420c, hair tufts of eighth
abdominal segment.
421. Ulophora guarinella (Zeller), specimen
from Cuba, aedeagus omitted; 421a,
aedeagus.
422. Chorrera idiotes Dyar, one harpe detached
and aedeagus omitted; 422a, elements
of transtilla and anellus; 422b, aedea-
gus; 422c, hair tufts of eighth abdom-
inal segment.
423. Chorrera extrincica (Dyar), harpe; 423a,
anellus.
424. Chorrera postica (Zeller), type, one harpe
detached and aedeagus omitted; 424a,
gnathos, elements of transtilla and
anellus; 424b, tufts of eighth abdom-
inal segment; 424c, aedeagus.
425. Tacoma feriella Hulst; 425a, aedeagus.
oN
420b.
421.guarinella
Lh
\Y..
423. extrincica
=
ZZ, |
1 425.teriella
%
424. postica
300328—5é6——27
410 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 426—-428.—Mates.
426. Epischnia prodromella (Hiibner), aedea-
gus omitted; 426a, aedeagus; 426b,
compound tufts of eighth abdominal
segment.
427. Laodamia faecella (Zeller), one harpe
detached and aedeagus omitted; 427a,
aedeagus; 427b, anellus; 427c, com-
pound tufts of eighth abdominal seg-
ment.
428. Megasis rippertella (Zeller) ; 428a, aedea-
gus; 428b, elements of transtilla; 428c,
harpe; 428d, eighth abdominal seg-
ment.
F: E880. \;
4 GC 427.
2 4 aecella
‘
42 Ta.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 41]
Figures 429-433.—Mates.
429. Adelperga cordubensiella _ (Ragonot),
specimen from Tucumén (in USNM);
429a, aedeagus; 429b, elements of
transtilla and anellus; 429c, tufts of
/ Sy eighth abdominal segment.
429c. 430. Eumysia mysiella (Dyar), type; 430a,
aedeagus.
431. Eumysia maidella (Dyar), sclerotization
and tufts of eighth abdominal segment,
431la, elements of transtilla and anel-
lus.
432. Divitiaca parvulella Barnes and McDun-
nough; 432a, aedeagus; 432b, anellus;
432c, eighth abdominal segment, show-
ing sclerotizations, haired lobes, and
scale tufts.
433. Divitiaca ochrella Barnes and McDun-
nough, type; 433a, dorsal view of
uncus and tegumen; 433b, aedeagus;
433c, anellus and elements of trans-
tilla; 433d, eighth abdominal segments,
showing sclerotization, haired lobes,
and scale tufts.
>
cordubensiella
parvulella
432b.
SSS
> SS
~ ochrella
412 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 434—438.—Matess.
434. Valdivia lativittella (Ragonot), figured
from the type of its synonym Zophodia
aureomaculella Dyar; 434a, anellus;
434b, aedeagus; 434c, sclerotization
and tufts of eighth abdominal seg-
ment.
435. Valdivia lativittella (Ragonot), Browns-
ville, Tex., example; 435a, anellus;
435b, sclerotization and tufts of
eighth abdominal segment.
436. Valdivia coquimbella Ragonot, paratype
from type locality (in BM); 436a,
aedeagus; 436b, anellus; 436c, sclero-
tization and tufts of eighth abdominal
segment.
437. Macrorrhinia aureofasciella Ragonot;
437a, anellus; 437b, aedeagus; 437c,
sclerotization, lobes and tufts of
eighth abdominal segment.
438. Ocala dryadella Hulst, type; 438a, anellus
and elements of transtilla; 438b, aedea-
gus; 438c, sclerotization, lobes and
tufts of eighth abdominal segment.
wv
435. \ativittella
y
“35a.
434 lativittella
/ 4
43
aureofasciella
anyadalian
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 413
440
mirabilicornella
eS 441.
samaritanella
morrisonella
Fiaurses 439-443.—Matzs.
439. Valdivia walkerella (Ragonot), type;
439a, aedeagus.
440, Protasia mirabilicornella (Dyar), type;
440a, aedeagus; 440b, anellus and
elements of transtilla; 440c, scleroti-
zations of eighth abdominal segment.
441. Helerographis samaritanella (Zeller) ;
44la, aedeagus; 441b, transtilla and
anellus; 441c, tufts of eighth abdomi-
nal segment.
442. Heterographis morrisonella Ragonot;
442a, anellus; 442b, aedeagus.
443. Staudingeria olivacella Dyar, a synonym
of S. albipenella (Hulst), type; 443a,
transtilla and anellus; 443b, aedeagus.
414 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 444-448 —MaAtes.
444, Hulstia undulatella (Clemens), figured
from the type of its synonym Honora
fumosella Hulst; 4442, aedeagus; 444b,
anellus and elements of transtilla;
444c, sclerotization and tufts of eighth
abdominal segment.
445. Honora mellinella Grote, specimen from
Texas; 445a, aedeagus; 445b, anellus
and elements of transtilla; 445c,
sclerotization and tufts of eighth
abdominal segment.
446. Honora sciurella Ragonot, type; 446a,
: aedeagus; 446b, anellus and elements
of transtilla; 446c, sclerotization and
tufts of eighth abdominal segment.
447. Honora subsciurella Ragonot; 447a,
aedeagus; 447b, anellus and elements
of transtilla; 447c, sclerotization and
tufts of eighth abdominal segment.
448. Honora dotella Dyar, type; 448a, aedea-
gus; 448b, anellus and elements of
transtilla; 448c, sclerotization and
tufts of eighth abdominal segment.
7168,
dotella.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 415
Fiaures 449-453.—Matgs.
449. Honora perdubiella (Dyar) type; 449a,
aedeagus; 449b, anellus and elements
of transtilla; 449¢, sclerotization and
tufts of eighth abdominal segment.
450. Honorinus fuliginosus Heinrich, new
species, type; 450a, aedeagus; 450b,
anellus; 450c, transtilla; 450d, sclero-
tization and tufts of eighth abdominal
segment.
Sms My 451. Cabotia semidiscella Ragonot; 45la, ele-
= ; 450.fuliginosus a ments of transtilla; 451b, aedeagus;
449, ey } \ i { Ay: 451c, eighth abdominal segment, show-
perdubiella a) \ ‘gs ing tufts.
452. Cabotia bonhoti (Hampson), specimen
from Trelawney Parish, Jamaica;
452a, anellus and elements of transtilla;
452b, aedeagus; 452c, tufts of eighth
abdominal segment; 452d, basal seg-
ments of male antenna.
453. Cabotia rhythmatica Dyar, type; 453a,
anellus and elements of transtilla;
453b, aedeagus; 453c, tufts of eighth
abdominal segment.
453. “Ss
rhythmatica
416 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 454—459.— Matss.
454. Oncolabis anticella Zeller; 454a, aedeagus;
454b, tufts of eighth abdominal seg-
ment; 454c, elements of transtilla and
anellus; 454d, basal segments of an-
tenna.
455. Canarsia ulmiarrosorella (Clemens) ; 455a,
aedeagus; 455b, anellus; 455c, tufts of
eighth abdominal segment.
456. Harnocha velessa Dyar, type, aedeagus
omitted; 456a, aedeagus; 456b, ele-
ments of transtilla.
457. Eurythmasis ignifatua Dyar, type, aedea-
gus omitted; 457a, dorsal view of uncus
and tegumen; 467b, gnathos and ele-
ments of transtilla; 457c, aedeagus;
457d, tufts of eighth abdominal seg-
ment.
458. Stylobasis rubripurpurea Hampson, type,
three-quarter view; 458a, aedeagus.
459. Eurythmidia ignidorsella (Ragonot),
aedeagus omitted; 459a, aedeagus; Ve 4 455.
459b, elements of transtilla; 459c, ter- iy = ulmiarrosorella
gite and sternite of eighth abdominal i 456. velessa
segment.
ry
ignifatua
be
cornic:
300329—56——28
AMERICAN MOTHS OF THE
SUBFAMILY
PHYCITINAE 417
460.
461.
462.
463.
464.
Figures 460—464.— Ma.gs.
Patriciola semicana Heinrich, new spe-
cies; 460a, aedeagus; 460b, ventral
view of right harpe; 460c, gnathos;
460d, elements of transtilla; 460e,
anellus.
Psorosina hammondi (Riley); 46la,
aedeagus; 461b, anellus and elements
of transtilla; 461c, tufts of eighth
abdominal segment of abdomen.
Palatka nymphaeella (Hulst); 462a,
aedeagus; 462b, sclerotization of eighth
abdominal segment.
Diviana eudoreella Ragonot, figured from
type of itssynonym, Dannemora eden-
tella Hulst; 463a, aedeagus; 463b,
gnathos; 463c, anellus.
Paconius corniculatus Heinrich, new
species, type; 464a, aedeagus; 464b,
anellus; 464c, tufts of eighth abdom-
inal segment.
418 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 465-467.—Ma tgs.
465. Aptunga macropasa (Dyar); 465a, ele-
ments of transtilla; 465b, gnathos;
465e, anellus; 465d, aedeagus; 465e,
eighth abdominal segment, showing
tufts and sclerotizations.
466. Cassiana malacella (Dyar), type; 466a,
anellus; 466b, aedeagus; 466c, gnathos;
466d, elements of transtilla; 466e,
eighth abdominal segment, showing
tufts and sclerotizations.
467. Anderida sonorella (Ragonot), figured
from type of its synonym Hyzophera
placidella Dyar; 467a, anellus; 467b,
elements of transtilla; 467c, gnathos;
467d, aedeagus; 467e, eighth abdom-
inal segment, showing tufts and scle-
rotizations.
(23 b | i
malacella hy ay F sepa
66c.
sonorella
470
bacerella
471. parvula
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 419
ine
moorei
Figurrs 468-473.—Mates.
468. Mescinia triloses Dyar, type; 468a, aede-
agus; 468b, gnathos; 468c, elements of
transtilla; 468d, anellus; 468e, eighth
abdominal segment, showing tufts and
sclerotizations.
469. Mescinia pandessa Dyar, type; 469a,
gnathos; 469b, elements of transtilla;
469c, anellus; 469d, aedeagus.
470. Mescinia bacerella Dyar, type; 470a, ae-
deagus; 470b, gnathos; 470c, elements
of transtilla; 470d, anellus.
471. Mescinia parvula (Zeller), paratype (in
BM); 471a, aedeagus; 471b, gnathos;
471c, elements of transtilla; 471d,
anellus.
472. Mescinia moorei Heinrich, new species;
472a, aedeagus; 472b, elements of
transtilla and anellus; 472c, gnathos.
473. Mescinia estrella Barnes and McDun-
nough, anellus and gnathos.
420 UNITED STATES NATIONAL MUSEUM BULLETIN 207 \
Figures 474—476.— Mates.
474. Mescinia peruella Schaus; 474a, aede-
agus; 474b, gnathos; 474c, elements of
transtilla; 474d, anellus.
475. Mescinia discella Hampson; 475a, aede-
agus; 475b, gnathos; 475c, elements of
transtilla; 475d, anellus.
476. Mescinia indecora Dyar; 476a, aedeagus;
476b, gnathos; 476c, elements of trans-
tilla; 476d, anellus; 476e, eighth ab-
dominal segment, showing tufts and
sclerotizations.
74
peruella
476. indecora
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 421
Fiagures 477-479.—Ma tess.
477. Nonia exiguella (Ragonot), figured from
type and from type of its synonym
Hypermescinia lambella Dyar, aedeagus
omitted; 477a, aedeagus; 477b, dorsal
view of uncus and tegumen; 477c,
gnathos; 477d, elements of transtilla;
477e, anellus; 477f, eighth abdominal
segment, showing tufts and sclerotiza-
tions.
478. Comotia torsicornis Dyar, type, aedeagus
omitted; 478a, aedeagus; 478b, ele-
ments of transtilla; 478c, tufts and
sclerotizations of eighth abdominal
segment; 478d, basal segments of an-
tenna.
479. Bema neuricella (Zeller), figured from
type of its synonym B. myja Dyar, ae-
deagus omitted; 479a, elements of
transtilla; 479b, aedeagus; 479c, eighth
abdominal segment, showing tuftings
and sclerotizations; 479d, front view of
head; 479e, front view of head, de-
nuded.
478.
torsicornis
479 \
Nneuricella
‘422 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 480—-483.— Mates.
. 480. Homoeosoma sinuellum (Fabricius) ;
480a, gnathos; 480b, elements of trans-
tilla; 480c, anellus; 480d, aedeagus;
480e, sclerotizations of eighth abdom-
inal segment.
481. Homoeosoma electellum (Hulst); 481a,
vinculum; 481b, aedeagus; 481c, gna-
thos; 481d, elements of transtilla;
481e, anellus.
- 482. Homoeosoma illuviellum Ragonot; 482a,
- aedeagus; 482b, gnathos; 482c, anellus;
482d, elements of transtilla; 482e,
sclerotizations of eighth abdominal
segment.
483. Homoeosomaimitator Heinrich, new species;
488a, lateral view of uncus, gnathos, and
tegumen; 483b, aedeagus; 483c, gna-
thos; 483d, elements of transtilla; 483e,
anellus; 483f, sclerotizations of eighth
abdominal segment, showing shortest
form of ventral process; 483g, longest
form of ventral process from eighth
abdominal segment.
480. sinue\lum
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 423
485. inornatellum
impressale
Fiaures 484-486.—Matxgs.
484. Homoeosoma longiventrellum Ragonot,
3 specimens from Chiriqui, Panamé4;
484a, aedeagus; 484b, gnathos; 484c,
anellus; 484d, elements of transtilla;
484e, sclerotizations of eighth abdom-
inal segment.
485. Homoeosoma inornatellum (Hulst) ; 485a,
aedeagus.
486. Homoeosoma impressale Hulst; 486a,
aedeagus; 486b, gnathos; 486c, ele-
ments of transtilla; 486d, anellus,
486e, sclerotizations of eighth abdom-
inal segment.
424 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 487—490.— Ma tgs.
487. Homoeosoma albescentellum Ragonot, fig-
ured from type of its synonym H.
elongellum Dyar; 487a, aedeagus.
488. Homoeosoma deceptorium Heinrich, new
species, paratype from Alcove, Quebec;
488a, gnathos with apical projection
shown in ventral and ventrolateral
views; 488b, elements of transtilla;
A88c, anellus; 488d, aedeagus; 488e,
sclerotizations of eighth abdominal
segment.
489. Homoeosoma discrebile Heinrich, new
species; 489a, aedeagus; 489b, gnathos;
489c, anellus; 489d, elements of trans-
tilla; 489e, tufts and sclerotizations of
eighth abdominal segment.
490. Homocosoma peregrinum Heinrich, new
species; 490a-b, two views of aedeagus.
discrebile
( 489d,\
peregrinum
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 425
Fieures 491-496.—Matgs.
491. Homoeosoma vepallidum Heinrich, new
species, type; 491a, aedeagus.
492. Homoeosoma assitum Heinrich, new spe-
cies, type; 491a, aedeagus.
493. Homoeosoma acmaeopterum Ragonot,
type; 498a, gnathos; 493b, elements of
transtilla; 493c, anellus; 493d, aedea-
gus; 498e, sclerotizations of eighth ab-
dominal segment.
494, Homoeosoma nimbosellum Ragonot, type;
494a, aedeagus; 494b, gnathos and
elements of transtilla; 494c, anellus.
495. Rotruda mucidella (Ragonot); 495a, gna-
thos; 495b, anellus; 495c, elements of
transtilla; 495d, aedeagus; 495e, lat-
eral view of uncus, gnathos, and tegu-
men; 495f, tufts and sclerotizations of
eighth abdominal segment.
496. Rotruda nimbella (Zeller), aedeagus.
491. vepallidum
493e.
695. ee
mucidella
426 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 497—502.—Matss.
497. Strephomescinia schausella Dyar, type,
aedeagus omitted; 497a, aedeagus;
497b, anellus.
498. Unadilla erronella (Zeller); 498a, aedea-
gus; 498b, gnathos, elements of trans-
tilla, and anellus; 498c-f, variations in
vinculum.
499. Unadilla maturella (Zeller), specimen
from Guatemala; 499a, aedeagus;
499b, gnathos; 499c, elements of
transtilla; 499d, anellus.
500. Laetilia portoricensis Dyar; 500a, aedea-
gus; 500b, anellus; 500c, gnathos;
500d, lateral view of female head.
501. Laetilia melanostathma (Meyrick), lateral
view of male head.
502. Laetilia coccidivora (Comstock), lateral
view of male head.
498. erronella
500. Sz
ortoricensis
P melanostathma
coccidivora
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 427
Figures 503-507.—Matgss.
503. Laetilia coccidivora (Comstock); 508a,
aedeagus; 503b, elements of trans-
tilla; 503c, gnathos; 508d, anellus.
504. Laetilia coccidivora quadricolorella
(Dyar); 504a, aedeagus; 504b, ele-
ments of transtilla; 504c, gnathos;
504d, anellus.
505. Laetilia coccidivora cardini Dyar, speci-
men from Florida; 505a, aedeagus;
505b, elements of transtilla; 505c,
gnathos; 505d, anellus.
506. Laetilia obscura Dyar; 506a, aedeagus;
506b, elements of transtilla; 506c,
gnathos; 506d, anellus.
507. Laetilia melanostathma (Meyrick); 507a,
aedeagus; 507b, elements of trans-
tilla; 507c, gnathos; 507d, anellus.
504.
quadricolorella
506
obscura
melanostathma
428 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 508-511.— Mates.
508. Laetilia zamacrella Dyar; 508a, anellus;
508b, gnathos; 508c, elements of
transtilla; 508d, aedeagus.
509. Laetilia myersella Dyar; 509a, aedeagus;
509b, gnathos; 509c, elements of
transtilla; 509d, anellus.
510. Laetilia ephestiella (Ragonot); 510a,
gnathos; 510b, elements of trans-
tilla; 510c, anellus.
511. Laetilia fiskella Dyar; 511a, aedeagus;
511b, gnathos; 511c, elements of trans-
tilla; 511d, anellus.
ephestiella
511.
Fiskella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 429
Ja
J se. ¢ 513c. >
513d.
My!
maculatellae
Fiaurses 512-514.—Ma.ss.
512. Laetilia glomis (Dyar), type; 512a,
aedeagus; 512b, gnathos; 512c, ele-
ments of transtilla; 512d, anellus.
518. Baphala goyensis (Ragonot), specimen
from Parand, Brazil; 513a, aedeagus;
513b, gnathos; 513c, elements of trans-
tilla; 513d, anellus.
514. Baphala basimaculatella (Ragonot), speci-
men from Texas (in BM); 514a,
aedeagus; 514b, gnathos; 514c, ele-
ments of transtilla; 514d, anellus; 514e,
sclerotizations and tufts of eighth
abdominal segment.
430 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 515-517.—Matzs.
515. Baphala glabrella (Dyar), specimen from
type locality; 515a, aedeagus; 515b,
gnathos; 515c, elements of transtilla;
515d, anellus.
516. Baphala haywardi Heinrich, new species;
516a, aedeagus; 516b, gnathos; 516c,
elements of transtilla; 516d, anellus.
517. Baphala homoeosomella (Zeller), specimen
from type locality (in BM, Zeller,
collection); 517a, aedeagus; 517b-c,
gnathos with apical projection in
ventrolateral and ventral views; 517d,
elements of transtilla; 517e, anellus;
517£, tufts and sclerotizations of eighth
abdominal segment; 517g, basal seg-
ments of antenna.
homoeosomella
519.stigmella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 431
519e.
Ficurss 518, 519.—Matss.
518. Rhagea packardella (Ragonot), figured
from type of its synonym Zophodia
orobanchella Dyar; 518a, aedeagus;
518b, gnathos; 518c, elements of trans-
tilla; 518d, anellus.
519. Rhagea stigmella (Dyar), type; 519a,
aedeagus; 519b, gnathos; 519c, ele-
ments of transtilla; 519d, anellus;
519e, sclerotizations of eighth abdom-
inal segment.
432 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 520, 521.—Mates.
NT TT ah Iie I mV ossiad r oS Sai]
520. Rhagea stigmella (Dyar), figured from its
synonym Yosemitia maculicula Dyar;
520a, aedeagus; 520b, gnathos; 520c,
elements of transtilla; 520d, anellus.
521. Rhagea stigmella (Dyar), variety from
Half Moon Bay, Calif.; 521a, aedeagus;
521b, gnathos; 521c, elements of trans-
tilla; 521d, anellus.
Ss
SR
=
521. stigmella i)
52ld.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 433
Figures 522-524.—Matzs.
522. Zophodia convolutella (Hiibner), aedeagus
omitted; 522a, aedeagus; 522b, anellus;
522c, elements of transtilla; 522d, api-
cal process of gnathos; 522e, scleroti-
zations and tufts of eighth abdominal
segment.
523. Cactobrosis longipennella (Hampson), fig-
ured from type of its synonym
Moodna elongatella Hampson, aedegus
omitted; 523a, dorsal view of uncus
and tegumen; 523b, elements of trans-
tilla; 523c, anellus; 523d, aedeagus;
523e, tufts of eighth abdominal seg-
ment.
524. Cactobrosis fernaldialis (Hulst), part of
shaft of antenna, ventral view.
523. longipennella
J
434 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 525, 526.—MA.gzs.
525. Cactobrosis maculifera Dyar; 5252, part of
shaft of antenna, ventral view; 525b,
basal segments of antenna, lateral
view; 525c, elements of transtilla; 525d,
anellus; 525e, aedeagus.
526. Cactobrosis strigalis (Barnes and Mc-
Dunnough); 526a, elements of trans-
tilla; 526b, anellus; 526c, aedeagus.
525. maculifera
526. strigalis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 435
Ficures 527, 528.—Matss.
527. Melitara prodenialis Walker; 527a, dorsal
view of uncus and tegumen; 527b,
gnathos; 527c, elements of transtilla;
527d, anellus; 527e-f, lateral and ven-
tral views of aedeagus.
528. Melitara dentata (Grote); 528a, ele-
ments of transtilla; 528b, anellus;
528c, aedeagus.
528. dentata
436 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 529-531.—Matss.
529. Olycella junctolineella (Hulst); 529a,
anellus; 529b, elements of transtilla;
529¢c, aedeagus.
530. Olycella junctolineella pectinatella (Hamp-
son), gnathos.
531. Olyca phryganoides Walker, specimen
from Azuda, Santo Domingo; 531a,
elements of transtilla; 531b, aedeagus;
531c, anellus.
SWS
SS QQ SS -\
SSS ANY
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 437
534. substituta
Ficures 532-534.—Matezs.
532. Alberada bidentella (Dyar), aedeagus
omitted; 532a, elements of transtilla;
532b, anellus; 532c, aedeagus.
533. Alberada parabates (Dyar); 533a, ele-
ments of transtilla; 523b, anellus;
533c, aedeagus.
534. Nanaia substituta Heinrich, type; 534a,
elements of transtilla; 534b, anellus;
534c, aedeagus; 534d, sternite and
tergite of eighth abdominal segment.
438 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 535-539.— Mates.
535. Cactoblastis cactorwm (Berg); 535a, ele-
ments of transtilla; 535b, anellus;
535c, aedeagus.
536. Cactoblastis doddi Heinrich, gnathos.
537. Cactoblastis mundelli Heinrich, gnathos.
538. Cactoblastis bucyrus Dyar: 538a, elements
of transtilla; 538b, anellus; 538c,
aedeagus.
539. Cahela ponderosella (Barnes and Mc-
Dunnough), type; 539a-c, various
modifications of apical process of
gnathos; 539d, elements of transtilla;
539e anellus; 539f, aedeagus.
535. cactorum Rick 536. doddi
537. mundelli
538. bucyrus
539 ponderosella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 439
1, polingella
543. tapiacola
Figures 540-543.—Matgs.
540. Rumatha bihinda (Dyar), showing sub-
basal sclerotized pocket (pkt.); 540a,
elements of transtilla; 540b, anellus;
540c, aedeagus.
541. Rumatha polingella (Dyar); 541a, ele-
ments of transtilla; 541b, anellus; 541c¢,
aedeagus.
542. Rumatha glaucatella (Hulst), aedeagus
omitted; 542a, elements of transtilla;
542b, anellus; 542c, aedeagus.
543. Tucumania tapiacola Dyar, aedeagus
omitted, vinculum somewhat fore-
shortened; 543a, elements of transtilla;
543b, anellus; 548c, aedeagus; 543d,
sternite and tergite of eighth abdom-
inal segment.
440 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 544-547.—Mates.
544, Yosemitia fieldiella (Dyar), aedeagus
omitted; 544a, elements of transtilla;
544b, anellus; 544c, aedeagus.
545. Yosemitia didactica Dyar, aedeagus omit-
ted; 545a, elements of transtilla; 545b,
anellus; 545c, aedeagus.
546. Yosemitia longipennella (Hulst), aedeagus
omitted; 546a, elements of transtilla;
546b, anellus; 546c, aedeagus.
547. Yosemitia graciella (Hulst), aedeagus
omitted; 5472, elements of transtilla;
547b, anellus; 547c, aedeagus; 547d,
sternite and tergite of eighth abdom-
inal segment.
547. graciella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 44]
Fiaures 548-550.—Ma.gs.
548, Eremberga leuconips (Dyar); 548a, ele-
ments of transtilla; 548b, anellus; 548e,
aedeagus.
549. Eremberga insignis Heinrich, type; 549a,
elements of transtilla; 549b, anellus;
549c, aedeagus.
550. Hremberga creabates (Dyar), type; 550a,
elements of transtilla; 550b, anellus;
550c, aedeagus,
548. leuconips
300329—56——29
442 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 551-553.—Ma tes.
551. Parolyca asthenosoma (Dyar), type; 551a,
elements of transtilla; 551b, anellus;
551c, aedeagus; 551d, part of shaft of
antenna.
552. Salambona analamprella (Dyar); 552a,
elements of transtilla; 552b, anellus;
552c, aedeagus.
558. Sigelgaita transilis Heinrich, type; 553a,
elements of transtilla; 553b, anellus;
558c, aedeagus; 553d, eighth abdom-
inal segment, showing hair tufts.
553. transilis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 443
Ficures 554, 555.—Matgzs.
554. Sigelgaita chilensis Heinrich; 554a, ele-
ments of transtilla; 554b, anellus; 554,
aedeagus.
555. Amalafrida leithella (Dyar); 555a, ele-
ments of transtilla; 555b, anellus;
555c, aedeagus; 555d, eighth abdominal
segment, showing hair tufts; 555e,
basal segments of antenna; 555f, inner
pectination from one of basal segments
of antennal shaft, showing attachment
of modified setae (greatly enlarged).
555. leithella
444 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 556—-558.—Matss.
556. Ozamia lucidalis (Walker) ; 556a, gnathos,
ventral view; 556b, elements of trans-
tilla; 556c, anellus; 556d, aedeagus;
556e, eighth abdominal segment, show-
ing hair tufts.
557. Ozamia fuscomaculella (Wright); 5957a,
elements of transtilla; 557b, anellus;
557c, aedeagus; 557d, basal segments
of antenna.
558. Ozamia punicans Heinrich: 558a, ele-
ments of transtilla; 558b, anellus;
558c, aedeagus.
556. lucidalis
558. punicans
557. fuscomaculella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 445
Fieurss 559, 560.—Matzs.
559. Ozamia fuscomaculella clarefacta Dyar;
559a, elements of transtilla; 559b,
anellus; 559c, aedeagus; 559d, eighth
abdominal segment, showing hair tufts;
559e, lateral view of genitalia with one
harpe omitted.
560. Ozamia hemilutella Dyar; 560a, elements
of transtilla; 560b, anellus; 560c, aede-
agus; 560d, eighth abdominal segment,
showing hair tufts.
559. clarefacta
560. hemilutella
446 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 561—564.— Mats.
561. Illatila gurbyris Dyar, aedeagus omitted,
vinculum foreshortened; 561a, aede-
agus; 561b, lateral view of tegumen
showing serrate production from base;
561c, gnathos; 561d, sternite of eighth
abdominal segment.
562. Lascelina canens Heinrich, new species,
type; 562a, aedeagus; 562b, sternite of
eighth abdominal segment.
563. Meitlphestia simplicula (Zeller); 563a,
uncus; 563b, aedeagus; 563c, gnathos
and anellus; 563d, tufts of eighth ab-
dominal segment.
564. Selga arizonella (Hulst); 564a, aedeagus;
564b, gnathos; 564c, elements of trans-
tilla; 564d, anellus.
~ 563.
simplicula
564.arizonella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 447
Fieures 565-567.—Matgs.
565. Diatomocera tenebricosa (Zeller), example
from French Guiana, placed here to
show affinities in genitalic structure of
Diatomocera and Entmemacornis; 565a,
aedeagus; 565b, gnathos; 565c, trans-
tilla; 565d, anellus; 565e, tufts of
eighth abdominal segment.
566. Entmemacornis proselytes Dyar; 566a,
aedeagus; 566b, gnathos; 566c, trans-
tilla; 566d, anellus; 566e, tufts of
eighth abdominal segment; 566f, basal
segments of antenna, denuded.
567. Entmemacornis pulla Heinrich, new spe-
Sky : Z| iy ‘ cies, type; 567a, aedeagus; 567b,
nebricosa =, | ANG gnathos; 567c, transtilla; 567d, anellus;
iS 567e, tufts of eighth abdominal seg-
ment.
i
567. pulla
448 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 568-570.— Mates.
568. Cayennia rufitinctalis Hampson; 568a,
aedeagus; 568b, gnathos; 568c, trans-
tilla; 568d, anellus.
569. Rioja neza Heinrich, new species, type;
569a, aedeagus; 569b, tergite and ster-
nite of eighth abdominal segment; 569c,
gnathos; 569d, transtilla; 569e, anellus.
570. Moerbes dryopella (Schaus), type; 570a,
aedeagus; 570b, gnathos; 570c, ele-
ments of transtilla; 570d, anellus;
570e, tergite and sternite of eighth
abdominal segment.
569. nexa
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 449
572b.
To
d
573. paenaista
300329—56——30
Fieures 571-573.—Matzs.
571. Moerbes alveolella (Ragonot); 571a, aede-
agus; 571b, gnathos; 571c, transtilla;
571d, anellus; 57le, tergite and ster-
nite of eighth abdominal segment.
572. Moerbes emendata Heinrich, new species;
572a, aedeagus; 572b, gnathos; 572c,
transtilla; 572d, anellus; 572e, tergite
and sternite of eighth abdominal seg-
ment.
573. Moodnopsis perangusta (Dyar), type;
573a, aedeagus; 573b, gnathos; 5738c,
transtilla; 573d, anellus; 573e, tergite,
sternite and hair tufts of eighth ab-
dominal segment.
450 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 574-577.—MA.LEs.
574. Moodnopsis inornatella (Ragonot); 574a,
aedeagus; 574b, gnathos; 574c, trans-
tilla; 574d, anellus; 574e, tergite, ster-
nite, and tufts of eighth abdominal
segment.
575. Moodnopsis parallela Heinrich, new spe-
cies; 575a, aedeagus; 575b, gnathos;
575c, transtilla; 575d, anellus; 575e,
tergite, sternite, and tufts of eighth
abdominal segment.
576. Moodnopsis <inveterella (Dyar); 576a,
aedeagus; 576b, gnathos; 576c, trans-
tilla; 576d, anellus; 576e, tergite,
sternite, and tufts of eighth abdominal
segment.
577. Moodnopsis portoricensis Heinrich, new
species, uncus, gnathos, and tegumen
of male genitalia; 577a, transtilla.
inornatella
575.parallela
575c.
577.
portoricensis
2
TAS i
BAe
VW 576d.
576.inveterella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 451
Figures 578-580.— Mats.
578. Euzophera cinerosella (Zeller), aedeagus
omitted; 578a, aedeagus; 578b, tufts of
eighth abdominal segment; 578c,
gnathos; 578d, transtilla; 578e, anellus.
579. Euzophera semifuneralis (Walker); 579a,
gnathos; 579b, transtilla; 579c, anellus;
579d, aedeagus,
580. Euzophera osiricolorella Hulst; 580a,
aedeagus; 580b, gnathos; 580c, anellus;
580d, transtilla.
its
579. semifuneralis
580. ostricolorella
452 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 581-584.— Mates.
581. Euzophera nigricantella Ragonot; 581a,
gnathos; 581b, transtilla, 581c, anellus;
581d, aedeagus.
582. Prosoeuzophera impletella (Zeller), para-
type (in BM); 582a, aedeagus; 582b,
gnathos; 582c, transtilla; 582d, anellus.
583. Eulogia ochrifrontella (Zeller); 583a,
aedeagus; 583b, tufts of eighth abdom-
inal segment of male; 583c, gnathos;
583d, elements of transtilla; 583e,
anellus.
584. Exuperius negator Heinrich, type; 584a,
aedeagus.
581. nigricantella.
SS
5830. 583.ochrifrontella
negator
584. 5 583e.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 4538
Figures 585-587.— Mates.
585. Edulica compedella (Zeller); 585a, gna-
thos; 585b, transtilla; 585c, aedeagus,
586. Farnobia quadripuncta (Zeller); 586a,
anellus; 586b, gnathos; 586c; transtilla '
586d, aedeagus; 586e, sternite of eighth
abdominal segment.
587. Gennadius junctor Heinrich, new species;
587a, sternite of eighth abdominal seg-
ment; 587b, aedeagus with penis ex-
truded; 587c, aedeagus with penis in
normal position; 587d, gnathos; 587e,
transtilla; 587f, anellus,
586. quadripunc a
587. junctor
454 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 588-591.—MaA.ss.
588. Ephestiodes erasa Heinrich, new species,
type; 588a, anellus; 588b, gnathos;
588c, elements of transtilla; 588d,
aedeagus.
589. Ephestiodes gilvescentella Ragonot; 589a,
aedeagus; 589b, gnathos; 589c, ele-
ments of transtilla; 589d, anellus;
589e, tufts of eighth abdominal
segment.
590. Ephestiodes infimella Ragonot; 590a,
gnathos; 590b, elements of transtilla;
590c, anellus; 590d, aedeagus.
591. Eurythmia coloradella Hulst, a synonym
of Ephestiodes erythrella Ragonot;
591a, gnathos; 591b, elements of trans-
tilla; 591c, anellus; 591d, aedeagus.
591. coloradella
592. mignonella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 455
592a.
Ficures 592-594.—Matzs.
592. Ephestiodes mignonella Dyar; 592a, gna-
thos, 592b, elements of transtilla;
592c, anellus; 592d, aedeagus.
593. Ephestiodes lucidibasella Ragonot, type;
593a, gnathos; 593b, elements of
transtilla; 593c, anellus; 593d, aedea-
gus.
594, Azaera muciella Schaus; 594a, gnathos;
594b, elements of transtilla; 594c,
anellus; 594d, aedeagus.
456 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 595-598.—Mates.
595. Micromescinia pygmaea Dyar, aedeagus
omitted; 595a, anellus; 595b, gnathos;
595c, transtilla; 595d, aedeagus; 595e,
tergite and sternite of eighth abdomi-
nal segment.
596. Ephestiodes stictella (Hampson), one
harpe detached and aedeagus omitted;
596a, aedeagus; 596b, gnathos; 596c,
transtilla; 596d, tufts of eighth abdomi-
nal segment.
597. Ephestiodes noniella Dyar, 597a, aedea-
gus; 597b, gnathos; 597c, transtilla;
597d, anellus; 597e, tergite and sternite
of eighth abdominal segment.
598. Vezina parasitaria Heinrich, new species,
type; 598a, anellus; 598b, gnathos;
598c, elements of transtilla; 598d, ae-
deagus.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 457
599.
ostrinella
600.
bisinuella
ae,
601.
biviella
Fieurres 599-601.—Matgzs.
599. Moodna ostrinella (Clemens); 599a, gna-
thos; 599b, elements of transtilla;
599c, aedeagus; 599d, tufts of eighth
abdominal segment; 599e, basal seg-
ment of antenna.
600. Moodna bisinuella Hampson; 600a, ae-
deagus; 600b, gnathos; 600c, elements
of transtilla; 600d, anellus.
601. Manhatta biviella (Zeller); 601a, aedea-
gus; 601b, gnathos; 601c, transtilla;
601d, anellus; 60le, tufts on eighth
abdominal segment.
458 UNITED STATES NATIONAL MUSEUM! BULLETIN 207
Figures 602-604.—Matss.
602. Manhaita setonella (McDunnough), speci- 1. His ON
: : Leen
men from Utah; 602a, gnathos; 602b,
transtilla; 602c, anellus; 602d, aedea-
gus.
603. Verina supplicella (Dyar); 603a, tergite, “602R.S
sternite, and tufts of eighth abdominal Lorall i
segment; 603b, aedeagus; 608c, gna- Sexonene css
thos; 603d, elements of transtilla; 602c.
603e, anellus.
604. Vagobanta divergens (Butler); 604a, ter-
gite and sternite of eighth abdominal
segment; 604b, aedeagus; 604c, gna-
thos; 604d, elements of transtilla;
604e, anellus.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 459
Ficures 605-608.—Matgzs.
605. Vitula edmandsae (Packard) ; 605a, aedea-
gus; 605b, gnathos; 605c, transtilla;
605d, anellus; 605e, tergite, sternite,
and tufts of eighth abdominal segment.
606. Vitula pinei Heinrich, new species; 606a,
gnathos; 606b, transtilla; 606c, anel-
lus; 606d, aedeagus.
607. Vitula lugubrella (Ragonot), harpe.
608. Moodnella paula Heinrich, type; 608a,
aedeagus; 608b, gnathos; 608c, trans-
tilla; 608d, anellus.
6050. 605. edmandsae
608. paula
460 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 609, 610.—Mates.
609. Volatica trinitatis Heinrich, new species;
609a, aedeagus; 609b, gnathos, 609c,
transtilla; 609d, anellus; 609e, tergite
and sternite of eighth abdominal seg-
ment.
610. Volatica pachytaeniella (Ragonot); 610a,
gnathos; 610b, transtilla; 610c, anellus;
610d, aedeagus.
610. pachytaeniella
610c.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 461
Fiaures 611-614.—Matzs.
611. Caudellia nigrella (Hulst), type; 611a,
gnathos plus transtilla; 611b, anellus;
61lc, aedeagus.
612. Caudellia declivella (Zeller); 612a, aedea-
gus; 612b, gnathos; 612c, transtilla.
613. Caudellia apyrella Dyar, type; 613, ae-
deagus; 613b, elements of transtilla.
614. Caudellia colorella (Dyar), one harpe de-
tached and aedeagus omitted; 614a,
aedeagus; 614b, transtilla.
612.declivella
613. apyrella
Bow
élécolbralla
462 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 615-619.— Mates.
615. Caudellia clara Heinrich, type; 615a,
gnathos; 615b, eighth abdominal seg-
ment, showing dorsal tufts.
616. Sosipatra rileyella (Ragonot) ; 616a, gna-
thos; 616b, transtilla; 616c, anellus;
616d, aedeagus.
617. Sosipatra micaceella (Hampson); 6172,
aedeagus; 617b, gnathos; 617c, trans-
tilla; 617d, eighth abdominal segment,
showing dorsal tufts.
618. Sosipatra anthophila (Dyar), type; 618a,
aedeagus; 618b, gnathos with side
sketch showing apical projection in
lateral view; 618c, transtilla; 618d,
anellus.
619. Sosipatra thurberiae (Dyar), type; 6198,
aedeagus; 619b, gnathos; 619c, trans-
tilla; 619d, anellus.
l
ye RES)
He
Sa
617. micaceella
618.
anthophila
AMERICAN
620827
majorella
ed 622.innoxia
622c.
MOTHS OF THE SUBFAMILY PHYCITINAE 463
Figures 620-623.—Ma.gs.
620. Sosipaira majorella (Dyar), aedeagus
omitted; 620a, aedeagus; 620b, gna-
thos; 620c, transtilla; 620d, tergite and
sternite of eighth abdominal segment.
621. Sostpatra nonparilella (Dyar), type, ae-
deagus omitted; 621a, gnathos; 621b,
transtilla; 621c, anellus; 621d, aedea-
gus.
622. Ribua innoxia Heinrich, aedeagus omit-
ted; 622a, aedeagus; 622b, apical pro-
jection of gnathos; 622c, anellus; 622d,
tergite, sternite, and dorsal tufts of
eighth abdominal segment.
623. Ribua contigua Heinrich, new species,
aedeagus omitted; 623a, aedeagus;
623b, fused gnathos and transtilla.
464 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fia@ures 624-627.— Matss.
624. Plodia interpunctella (Hiibner); 624a,
aedeagus; 624b, transtilla.
625. Plodia dolorosa Dyar; 625a, aedeagus;
625b, transtilla; 625c, tergite, sternite,
and dorsal tufts of eighth abdominal
segment.
626. Anagasta kiuhniella (Zeller), aedeagus
omitted; 626a, aedeagus; 626b, gna-
thos; 626c, elements of transtilla; 626d,
anellus; 626e, tergite, sternite, and
dorsal tufts of eighth abdominal seg-
ment.
627.—Cabnia myronella Dyar, aedeagus omit-
ted; 627a, aedeagus; 627b, gnathos;
627c, transtilla; 627d, anellus; 627e,
lateral view of head.
624.2"
interpunctella
626. kuhniella
627. myronella
AMERICAN MOTHS OF
629a. Ss
629.cautella
630. figulilella
THE SUBFAMILY .PHYCITINAE 465
Figures 628-630.—Matzs.
628. Ephestia elutella (Hiibner); 628a, aedea-
gus; 628b, gnathos; 628c, transtilla;
628d, eighth abdominal segment, show-
ing dorsal tufts.
629. Ephestia cautella (Walker); 629a, aedea-
gus; 629b, gnathos; 629c, transtilla;
629d, anellus; 629e, dorsal view of
uncus and tegumen; 629f, eighth ab-
dominal segment, showing dorsal tufts.
630. Ephestia figulilella Gregson; 630a, aedea-
gus; 630b, gnathos; 630c, transtilla:
630d, anellus.
466 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 631-634.—Mat.es
631. Varneria postremella Dyar, one harpe de-
tached ‘and aedeagus omitted; 631a,
transtilla; 631b, aedeagus; 631c, eighth
abdominal segment, showing dorsal
tufts.
632. Varneria dubia Heinrich, new species,
aedeagus omitted; 632a, aedeagus;
632b, gnathos; 632c, transtilla; 632d,
eighth abdominal segment, showing
dorsal tufts.
633. Eurythmia hospitella (Zeller); 633a, ae-
deagus; 633b, gnathos; 633c, transtilla;
633d, a slight modification of the
central projection of transtilla drawn
from an example of angulella Ely
(-diffusella Ely); 633e, anellus; 633f,
sclerotizations of eighth abdominal
segment.
634. Hrelieva quantulella (Hulst); 6342, aede-
agus; 634b, gnathos; 634c, transtilla;
634d, eighth abdominal segment, show-
ing dorsal tufts.
Sat
63lc.
RE
VJ %
guantutella
634b.
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 467
635. conops Z\
~ 635c.
637. ~
apianella
Figures 635-637.—MA.zs
635. Rabiria conops (Dyar), type, aedeagus
omitted; 635a, gnathos; 635b, eighth
abdominal segment; 635c, transtilla;
635d, aedeagus.
636. Microphycita titillella Dyar, aedeagus
omitted; 636a, gnathos; 636b, trans-
tilla; 636c, aedeagus and anellus.
637. Nicetiodes apianella Schaus, type; 637a,
gnathos; 637b, transtilla; 637c, anellus;
637d, aedeagus; 637e, eighth abdom-
inal segment showing tufts.
468 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 638-644.—FEMALE GENITALIA.
638. Cryptoblades rutilella Zeller.
639. Cryptoblades gnidiella (Milliére).
640. Acrobasis tumidella (Zincken).
641. Acrobasis indigenella (Zeller).
642. Acrobasis vaccinii Riley.
643. Acrobasis tricolorella Grote.
644. Acrobasis juglandis (LeBaron), specimen
reared from pecan.
642. : 644,
indiginella vaccinii tricolorella juglandis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 469
Fieures 645-650.—FEMALE GENITALIA.
645. Acrobasis tumidulella (Ragonot), type.
646. Acrobasis comptella Ragonot.
647. Rhodophaea caliginella (Hulst), type.
648. Rhodophaea supposita (Heinrich).
649. Rhodophaea advenella (Zincken).
650. Trachycera pallicornella (Ragonot).
16 a otella
tumidulella
649.
supposita .- _—_ advenella pallicornella
470 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 651-654.— FEMALE GENITALIA.
651. Mildrizia constitutionella Dyar; 651a, sig-
num of bursa, enlarged.
652. Anabasis ochrodesma (Zeller).
653. Sematoneura atrovenosella (Ragonot).
654. Hypsipyla dorsimacula (Schaus).
653. atrovenosella
654
. dorsimacula
652.ochrodesma
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 471
Figures 655-657.—FEMALE GHNITALIA.
655. Hypsipyla grandella (Zeller),
656. Hypsipyla pagodella Ragonot.
657. Hypsipyla ferrealis (Hampson).
655.grandella
656. pagodella 657. ferrealis
472 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 658-660.— FEMALE GENITALIA.
658. Hemiptilocera chinographella Ragonot.
659. Hemiptilocera jocarella (Schaus), type.
660. Hemiptilocera bigrana (Zeller).
658,chinographella
659. 3
jecarella 660. bigrana
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 473
Fieures 661-663.—FEMALE GBNITALIA.
661. Hemiptilocera exoleta (Zeller), type; 66la,
dorsal view of eighth abdominal seg-
ment.
662. Hemiptilocera letharda (Schaus), showing
to the side of bursa a greatly enlarged
figure of the spines of signum.
663. Crocidomera stenopteryx (Dyar), specimen
from type locality.
661. exoleta
662.
letharda
663. stenopteryx
300329—56——31 stots Suiibaihicgs = ="
474 UNITED STATES NATIONAL MUSEUM BULLETIN 207.
Ficures 664—-667.— FEMALE GENITALIA.
664. Crocidomera turbidella Zeller.
665. Crocidomera fissuralis (Walker), type.
666. Cuniberia subtinctella (Ragonot).
667. Adanarsa iniransitella (Dyar), showing a
great enlargement of signum to one side
of bursa.
666.
subtinctella
"ences anorenens
664. turbidella
667. intransitella 665. fissuralis
670.
annuliferella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 475
1 668.
definitella
671. hystriculella
669.grisella
Fieures 668-671.—FEMALH GENITALIA.
668. Hypargyria definitella (Zeller).
669. Bertelia grisella Barnes and McDun-
nough.
670. Chararica annuliferella (Dyar).
671. Chararica hystriculella (Hulst).
476 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 672-675.— FEMALE GENITALIA.
672. Myelopsis subtetricella (Ragonot), type.
673. Myelopsis coniella (Ragonot).
674. Myelopsis fragilella (Dyar), a2 synonym
of M. alatella (Hulst).
675. Myelopsis minutularia (Hulst), type.
673. coniella 675. minutularia
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 477
676.
bistriatella
677. decolor
678, ceratoniae 679. univitella
Ficurres 676-679.—FEMALE GENITALIA.
676. Apomyelois bistriatella (Hulst).
677. Ectomyelois decolor (Zeller).
678. Pctomyelois ceratoniae (Zeller).
679. Anypsipyla univitella Dyar.
478 UNITED STATES NATIONAL MUSEUM BULLETIN. 207
Fraures 680-684.—FSMALE GENITALIA.
680. Ectomyelois muriscis (Dyar).
681. Ectomyelois furvidorsella (Ragonot), type.
682. Ectomyelois zeteki Heinrich, new species.
683. Paramyelois transitella (Walker).
684. Myelois cribrella (Hiibner).
680. muriscis
furvidorsella
684. cribrella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 479
Fiaurses 685-691.—FEMALE GENITALIA.
685. Diatomocera dosia (Dyar), paratype;
685a, enlargement of signa.
686. Diatomocera tenebricosa (Zeller); 686a,
enlargement of signa.
687. Diatomocera excisalis (Hampson), para-
type from Cayenne, French Guiana
(in USNM), signa in bursa copulatrix.
688. Diatomocera extracta Heinrich, new spe-
cies, bursa copulatrix.
689. Diatomocera mochlophleps (Dyar), bursa
copulatrix; 689a, enlargement of sig-
num in bursa.
690. Pseudodivona santa-maria Dyar, type,
bursa copulatrix.
691. Pseudodivona carabayella Dyar, specimen
from Incachaca, Bolivia.
690.
santa-maria
687.
excisalis
691.
carabayella
689. mochlophleps
’
1
At
ah
Nay
688. extracta
480 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 692-699.— FEMALE GENITALIA. $ :
rubrizonalis
693.
692. Pseudocabima expunctriz (Dyar and Hein- he
euzopherella 694.
rich), signa of female genitalia.
693. Pseudocabima euzopherella (Dyar), signa
of female genitalia.
694. Pseudocabima rubrizonalis (Hampson),
signa of female genitalia.
695. Pseudocabima perrensiella (Ragonot),
type; 695a, enlargement of signa.
696. Pseudocabima castronalis Heinrich, new
species, paratype; 696a, enlargement
of signa.
697. Pseudocabima guianalis Heinrich, new
species, signa of female genitalia.
698. Pseudocabima arizonensis Heinrich, new
species, signa of female genitalia.
699. Pseudocabima nigristrigella (Ragonot),
specimen in BM; 699a, enlargement
of signa.
arizonensis
698.
guianalis
695, castronalis
perrensiella sie nee
nigristrigella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 481
701.
majorina
700.celiella
702. 703.
fulgidula stictoneurella egenella
300329—56——32
Ficures 700-704. FEMALE GENITALIA AND
(a) ENLARGEMENT OF SIGNA.
700. Hyalospila celiella Schaus.
701. Hyalospila majorina Heinrich, new spe-
cies.
702. Hyalospila fulgidula Heinrich, new spe-
cies.
703. Hyalospila stictoneurella Ragonot.
704. Hyalospila egenella (Ragonot), type.
482 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieaures 705-709. FaMALE GENITALIA.
705. Hyalospila clevelandella (Dyar), paratype
from Porto Bello, Panamdé; 705a,
enlargement of signum patch.
706. Hyalospila clevelandella (Dyar), speci-
mens from México and Guatemala;
706a, enlargement of signum.
707. Hyalospila angulineella (Schaus), type;
707a, enlargement of signa.
708. Hyalospila semibrunneella (Ragonot),
type.
709. Hyalospila xanthoudemia (Dyar); 709a,
enlargement of signum.
106a.
clevelandella ?
<=
709a.
709,
708. semibrunncella Je Roudsmra
ignobilis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 483
pellucens
ahemora
Figures 710-714.— FEMALE GENITALIA.
710. Fundella agapella Schaus, type.
711. Fundella argentina Dyar; 711a, armature
of genital opening (in South American,
Argentine, and Brazilian specimens).
712. Fundella ignobilis Heinrich, new species.
713. Fundella pellucens Zeller; 713a, dorsal
view of eighth segment collar.
714. Fundella ahemora Dyar.
484. UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiqures 715-719.— FEMALE GENITALIA.
715. Difundella corynophora Dyar; 715a, dor-
sal view of collar of eighth abdominal
segment; 715b, ventrolateral pockets in
intersegmental area between seventh
and eighth abdominal segments, shown
from dorsal view.
716. Difundella distractor Heinrich, new spe-
cies.
717. Difundella tolerata Heinrich, new species;
717a, invaginated, sclerotized, dorsal
shield of seventh segment of abdomen.
718. ?Coptarthria dasypyga (Zeller), example
from Costa Rica; 718a, enlargement of
signum.
719. Dasypyga alternosquamella Ragonot.
distractor
718. dasypyga ? 719, alternosqaamella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 485
Figurts 720-725.—FEMALE GENITALIA.
720. Promylea lunigerella glendella (Dyar),
showing enlargement of signum beside
bursa; 721a, dorsal view of sclerotized
apron and pocket attached to collar.
721, Promylea lunigerella lunigerella Ragonot,
specimen from Duncans, Vancouver
Isl., showing enlargement of signum
beside bursa; 721la, dorsal view of
sclerotized apron and pocket attached
to collar; 721b, lateral view of same,
also showing genital opening of ductus
bursae.
722. Promylea druceii (Ragonot), type; 722a,
dorsal view of sclerotized apron and
pocket attached to collar.
723. Promylea mindosis Dyar; 728a, dorsal
view of sclerotized apron and pocket
attached to collar.
724. Promylea dasystigma Dyar; 724a, dorsal
view of sclerotized apron and pocket
attached to collar.
725. Scorylus cubensis Heinrich, new species.
721.
glendella lunigerella
722.
druceii 723. mindosis 724. deasystigma
725. cubensis
486 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 726-733.— FEMALE GENITALIA.
726. Anadelosemia fifria Dyar, showing en-
largement of signum to side of bursa.
727. Anadelosemia base Dyar.
728. Anadelosemia obstitella (Schaus), type,
showing enlargement of signum beside
bursa; 728a, dorsal view of eighth-
segment collar.
729. Anadelosemia texanella (Hulst), dorsal
view of eighth-segment collar.
730. Anadelosemia condigna Heinrich, new spe-
cies, dorsal view of eighth-segment
collar.
731. Rampylla lophotalis Heinrich, new spe-
cies; 731a, dorsal views of ovipositor
and eighth-segment collar.
732, Rampylla polydectella (Schaus), type;
732a, dorsal view of eighth-segment
collar; 732b, invaginated, sclerotized
pocket from seventh abdominal seg-
ment.
733. Rampylla subcaudata (Dyar), specimen
from Quirigus, Guatemala; 733a, dor-
sal views of eighth-segment collar and
sclerotized pocket of seventh segment.
726. fifria
729.
texanella
728. obstitella
727. base
733.
subcaudata
732.
731. lophotalis polydectella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 487
735. caricae
736.
columnella
737, azonaxsalis
Ficures 734-738.—FEMALE GENITALIA,
734.
735.
736.
737.
738.
Davara interjecta Heinrich, new species.
Davara caricae (Dyar).
Davara columnella (Zeller), type.
Davara azonazsalis (Walker), type.
Davara nerthella (Schaus), type.
488 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 739-744.— FEMALE GENITALIA.
739. Davara rufulella (Ragonot).
740. Sarasota furculella (Dyar), specimen
from Cuba.
741. Atheloca subrufella (Hulst), type.
742. Praedonula almonella (Dyar); 742a, dor-
salview of eighth-segment collar,
showing genital opening and its
attachments.
743. Piesmopoda montella Schaus, type.
744. Piesmopoda trichomata (Zeller), cotype.
740.furculella
742. almonella 743. montella trichomata
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 489
745. xanthomera 746.flavicans 748. xanthopol ys
750. parva
751 apocerastes
752. semirufella
745
746.
747.
748.
749.
750.
751.
752.
Ficures 745-752.— FEMALE GENITALIA.
. Piesmopoda sxanthomera Dyar, type,
detail to the side of collar shows its
invagination; similar details shown in
figures 746 to 750.
Piesmopoda flavicans (Zeller).
Piesmopoda ragonoti (Dyar).
Piesmopoda xanthopolys Dyar, type.
Piesmopoda isabella (Dyar).
Piesmopoda parva Heinrich.
Piesmopoda apocerastes Dyar.
Piesmopoda semirufella (Zeller).
490 UNITED STATES NATIONAL MUSEUM BULLETIN 207
. Fieures 753-758.— FEMALE GENITALIA.
. 753. Peadus subaquilellus (Ragonot), type,
with enlargement of signum shown to
the side of bursa.
754. Peadus burdettellus (Schaus).
755. Peadus dissitus Heinrich, new species.
756. Gabinius paulsoni (Ragonot), specimen
from ‘“‘Chile, Silva.’
757. Ceracanthia mamella (Dyar), type.
758. Drescoma cyrdipsa Dyar.
755. dissitus
756. paul soni
TST. momella 758. cyrdipsa
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 491
761. drucella
760.alpha
762: subelisa 764..nubilella 765. pogerythrus
761.
762.
763.
764.
765.
Fiaures 759-765.— FEMALE GENITALIA.
759.
760.
Megarthria beta Heinrich, type.
Megarthria alpha Heinrich, type; 760a, b,
collar in dorsal view showing variations
in the apron.
Drescoma drucella Dyar, a synonym of
Drescomopsis soraella (Druce).
Drescomopsis subelisa Dyar, a synonym
of Drescomopsis soraella (Druce), with
signum shown in full lateral view.
Monoptilota pergratialis (Hulst), bursa
copulatrix of female genitalia.
Monoptilota nubilella Hulst, paratype, a
synonym of M. pergratialis (Hulst).
Zamagiria pogerythrus Dyar.
492 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 766—770.— FEMALE GENITALIA.
766. Zamagiria australella (Hulst).
767. Zamagiria laidion (Zeller).
768. Zamagiria ipsetona Dyar.
769. Magiriopsis denticosella (Dyar).
770. Anegcephaleszs arctella (Ragonot).
australella
768. ipsetona - 769. denticosella 770. arctella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 493
Ficures 771-775.— FEMALE GENITALIA.
771. Ancylostomia stercorea (Zeller); 771a,
ventral view of ductus bursae and
eighth-segment collar with membrane
of seventh segment removed.
772. Adelperga cordubensiella (Ragonot), type.
773. Caristanius decoloralis (Walker); 773a,
ventral view of eighth-segment collar
and apical portion of ductus bursae.
774. Caristanius guatemalellus (Ragonot),
type.
775. Caristanius pellucidellus (Ragonot) ; 775a,
ventral view of apical part of ductus
bursae and eighth-segment collar;
775b, dorsal view of eighth-segment
collar and its attachments to ductus
bursae.
“y j 4 74
ff af
i ie
I #
774.
775. pellucidellus
temalellus
494 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 766—780.— FEMALE GENITALIA.
776. Pima boisduvaliella (Guénée), bursa cop-
ulatrix, apical part of ductus bursae,
and dorsal outline of eighth-segment
collar of female genitalia.
777. Pima albiplagiatella (Packard).
778. Pima albocostalialis (Hulst), bursa copu-
latrix, apical part of ductus bursae, and
dorsal outline of eighth-segment collar
of female genitalia.
779. Pima albocostalialis (Hulst), a specimen
showing extent of variation in patches
of bursa and shape of eighth-segment
collar.
780. Pima vwidella (McDunnough), specimen
from Aweme, Manitoba, bursa copula-
trix, apical part of ductus bursae, and
dorsal outline of eighth-segment collar
of female genitalia.
780. vividella 779. albocostalialis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 495
Ficures 781-783.— FEMALE GENITALIA
781. Pima granitella (Ragonot).
782. Pima parkerella (Schaus).
783. Pima fosterella Hulst, specimen from
Arizona, bursa copulatrix, apical part
of ductus bursae, and dorsal outline of
eighth-segment collar of female geni-
talia.
7; 781.
granitella
782. parkerella 783. fosterella
496 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 784—788.—FEMALE GENITALIA.
784. Interjectio columbiella (McDunnough),
specimen from Pullman, Wash,
785. Interjectio ruderella (Ragonot), type.
786. Olybria aliculella (Hulst).
787. Olybria furciferella (Dyar).
788. Oreana unicolorella (Hulst).
786. aliculella
= 788. unicolorella
787. furciferella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 497
Figures 789-790.—FEMALE GENITALIA.
789. Ambesa lallatalis (Hulst); 789a, dorsal
view of eighth-segment collar.
790. Ambesa laetella Grote.
790. laetella
789. lallatalis
498 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 791-793.—FEMALE GENITALIA.
791. Ambesa walsinghami (Ragonot).
792. Ambesa walsinghami (Ragonot), figured
from type of its synonym, A. monodon
Dyar.
793. Ambesa walsinghami mirabella Dyar.
792.walsinghami
791. walsinghami 793. mirabella
796. marginea
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 499
794. actualis
797. bistriatella
798. nigrovittella
Ficures 794-798.—FEMALE GENITALIA.
794. Catastia actualis (Hulst).
795. Catastia incorruscella (Hulst), specimen
from Arizona.
796. Catastia marginea (Schiffermiiller).
797. Catastia bistriatella (Hulst), specimen
from type locality.
798. Immyrla nigrovittella Dyar.
500 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 799-803.—FEMALE GENITALIA.
799. Salebriacus odiosellus (Hulst); 799a,
dorsal view of eighth-segment collar.
800. Salebriaria annulosella (Ragonot), speci-
men from Burnet County, Tex.
801. Salebriaria tenebrosella (Hulst), type.
802. Salebriaria turpidella (Ragonot).
803. Salebriaria tenebrosella (Hulst), figured
from large example.
tenebrosella
NG
799. odiosellus 803. tencbrosella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 501
fruc
807. pasadamia
808. jugosella
Figures 804-808.—FEMALE GENITALIA.
804. Salebriaria pumilella (Ragonot).
805. Salebriaria fructetella (Hulst), type.
806. Quasisalebria admizta Heinrich, paratype
from type locality.
807. Ortholepis pasadamia (Dyar); 807a, dor-
sal view of eighth-segment collar; 807b,
dorsal view of junction of bursa and
ductus bursae.
808. Ortholepis jugosella Ragonot; 808a, dorsal
view of eighth-segment collar.
502 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaurts 809-812.— FEMALE GENITALIA.
809. Polopeustis annulatella (Zetterstedt);
809a, dorsal view of eighth-segment
collar.
810. Polopeustis arctiella (Gibson), specimen
from Labrador; 810a, dorsal view of
; eighth-segment collar.
811. Glyptocera consobrinella (Zeller).
812. Meroptera pravella (Grote).
611, consobrinella 612. pravella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 503
Figures 813-817.— FEMALE GENITALIA.
813. Meroptera abditiva Heinrich, new species,
paratype from type locality, with
eighth-segment collar and ovipositor
omitted.
814. Meroptera abditiva Heinrich, new species,
paratype from type locality.
815. Nephopteryx rhenella (Zincken).
816. Meroptera mirandella Ragonot.
817. Meroptera cviatella Dyar.
815. rhenella
816. mirandella
504 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 818-822.—FEMALE GENITALIA.
818. Nephopteryx delassalis Hulst, type.
819. Nephopteryx fernaldi (Ragonot).
820. Nephopteryx vetustella (Dyar).
$21. Nephopteryx subfuscella (Ragonot), fig-
ured from type of its synonym Salebria
semiobscurella Hulst.
822. Nephopteryz dammerst Heinrich. new
species.
818. delassalis
820. vetustella
821. subfuscella 822 . dammersi
sts
826. subcaesiella
800329—56——-33
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 505
827. virgatella
Figures 823-827.—FEMALE GENITALIA.
823. Nephopteryx carneella Hulst, reared speci-
men from Maine.
824. Nephopteryz uvinella (Ragonot).
825. Nephopteryx inconditella (Ragonot).
826. Nephopteryx subcaesiella (Clemens),
827. Nephopteryx virgatella (Clemens).
506 UNITED STATES NATIONAL MUSEUM BULLETIN’ 207
Figures 828-831.—FEMALE GENITALIA.
828. Nephopteryx termitalis (Hulst), figured
from type of its synonym Salabria
levigatella Hulst.
829. Nephopteryx basilaris Zeller.
830. Nephopteryx celtidella (Hulst).
831. Nephopieryx bifasciella Hulst. 831a, ab-
normal specimen from Yuma, Ariz.
830. celtidella
<B%
831. bifasciella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 507
Figures 832-836.—FEMALE GENITALIA.
832. Nephopteryx rubrisparsella (Ragonot).
833. Nephopteryx bisra Dyar, type.
834. Tlascala reductella (Walker).
835. Nephopteryx crassifasciella Ragonot.
836. Nephopteryx gilvibasella Hulst.
834. reductella
835. crassifasciella 836. gilvibasella
508 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 837—-842.— FEMALE GENITALIA.
837. Actriz dissimulairix Heinrich.
838. Actriz nyssaecolella (Dyar).
839. Homoeographa lanceolella Ragonot, para-
type (in BM, “Peru, Coll. Walker,
84-72”).
840. Etiella zinckenella (Treitschke).
841. Tulsa infinitella (Dyar), type.
842. Tulsa umbripennis (Hulst), specimen
from Chimney Gulch, Colo.
841. infinitella
PI
g
z
=
@
840. 3inckenella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 509
Ficures 843-848.— FEMALE GENITALIA,
843. Telethusia ovalis (Packard).
844. Phobus brucet (Hulst).
845. Phobus curvatellus (Ragonot).
846. Phobus incertus Heinrich, new species.
847. Stylopalpia scobiella (Grote).
848. Stylopalpia lunigerella Hampson.
843. ovalis
curvatellus
84.7. scobiella
UNITED STATES NATIONAL ‘MUSEUM BULLETIN 207
510
xe
Figures 849-852.—FEMALE GENITALIA.
849, Pyla fasciolalis (Hulst), with ovipositor
omitted; 849a, female genitalia of a
variety from Gunnison County, Colo.
850, Pyla impostor Heinrich, new species.
851. Pyla viridisuffusella Barnes and McDun-
nough.
852. Pyla fusca (Haworth).
mr sae
ROOT OAV ANE Cv nyse ted sem uta pan IVETE
849a. fasciolalis
851. viridisuffusella
852 fusca
AMERICAN MOTHS OF THE > SUBFAMILY PHYCITINAE 511
Figures 853-856.—FBMALE GENITALIA.
853. Pyla aenigmatica Heinrich, new species.
854. Pyla hypochalciella (Ragonot).
855. Pyla aequivoca Heinrich, new species.
856. Pyla insinuatriz Heinrich, new species,
MOLINA BEA NE RMweserets etry,
a
~Stam
econ
—
855. aequivoca 856. insinuatrix
512 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 857—860.—FEMALE GENITALIA.
857. Pyla scintillans (Grote), figured from
type of its synonym P. feella Dyar;
857a, female genitalia of a specimen
from Tuolumne Meadows, Calif.
858. Pyla sylphiella Dyar.
859. Pyla rainierella Dyar.
860. Pyla hanhamella Dyar.
ean e7es
ESenimenn
858.sylphiella
Yi S; y
Ke By A
= WA NM NZ
Hee ie Jt
PS scintillans iS;
es : Hates
yg z
bog 4 5
a-i \ 2d _ 857a,
bey
cent
bell
NZ
ays
859. rainierella 860. hanhamella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 513
Ficures 861-864.—FEMALE GENITALIA.
861. Pyla fasciella Barnes and McDunnough.
862. Pyla aeneoviridella Ragonot.
863. Pyla metalicella Hulst; 863a, specimen
from Silverton, Colo.
864. Pyla aeneella Hulst, type.
861. fascicella
<
f
PREERTO Cocet aE
at seseue LOMAATEA NES OI RPC COMATOSE
8630. 863. metalicella 864.aeneella
300328—56——34
UNITED: STATES NATIONAL MUSEUM BULLETIN ‘207
514)
Fieures 865—867.— FEMALE GENITALIA.’
865. Dioryctria abietella (Denis and Schiffer-
miiller) ; 865a, part of female genitalia
of a specimen showing variation in the
suture of the sclerotization of ductus
bursae.
866. Dioryctria sysstratiotes Dyar.
867. Dioryctria reniculella Grote.
ALORA NAOII OS BLAS) pn ERNEL
Be otiey
f
866. sysstratiotes
1
/
865. abictella 867. reniculella
868. ponderosae
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 552
nemoniaa
suits
862 erythropasa
870. Dioryctria
Sp.
871. majorella
Ficures 868-871.—FEMALE GENITALIA:
868. Dioryctria ponderosae Dyar, paratype
from California.
869. Dioryctria erythropasa (Dyar), paratype:
870. Dioryctria sp., a probable hybrid of au-
ranticella and erythropasa.
871. Dioryctria majorella Dyar, type.
516 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficures 872—874.—F2EMALE GENITALIA.
872.. Dioryctria disclusa Heinrich, paratype
from New Jersey.
873. Dioryctria auranticella (Grote).
874. Dioryctria horneana (Dyar), type.
| 873. auranticella
872. disclusa 874. horneana
517
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
Figures 875-878.—FEMALE GENITALIA.
ir
o
s
Q
ive)
_
S
_
o
-
S}
iS)
'-
i=
3
=
n
*)
RS
=
=
3
=
~~
=
o
>
i]
‘=
Q
1S
iw
eo
®
2,
lal
~
zg
3
a
3
cal
Q
~—
38
=
:3
2s
a
g&
a3
ans)
oe
Ace
23s
ge
n
dS
Q
3
i=
-)
~~
°
q
fs)
&p
a
[a=
2s
xO
aay
as
9 &§
Ss
i =
oe.
Ss
WE
gs
o2
Q
nN
b
re)
878. Dioryctria zimmermani (Grote), figured
from a large western (Ariz.) example
D. delectella (Hulst).
of its synonym,
875.
3immermani
cambiicola
878. zimmermani
518 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 879-882.— FEMALE GENITALIA.
879. Dioryciria amaiella (Hulst), typical Flor-
ida specimen.
880. Dioryctria albovittella (Hiilst).
881. Dioryctria baumhoferi Heinrich, new spe-
cies.
882. Dioryctria subtracta Heinrich, new species.
882. subtracta
880. albovittella
881. baumhoferi
879. amatella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 519
Figures 883-885.—FEMALE GENITALIA.
883. Dioryctria clarioralis (Walker), typical
Florida specimen.
884. Epischnia prodromella (Hiibner).
885. Laodamia faecella (Zeller).
a
Me dy
TAG
%
EES
peice 1 tea
884. prodromella
883. clarioralis
885. faecella
520 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 886—890.—FEMALE GENITALIA.
886. Sarata delia Heinrich, new species.
887. Sarata kappa Heinrich, new species, type.
888. Sarata alpha Heinrich, new species, type.
889. Sarata beta Heinrich, new species, type.
890. Sarata gamma Heinrich, new species.
\
Sa (RE
886. delta
wee
shee,
BRIS
892. epsilon
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 521
rhoiella
ots
894.1
ota
Figures 891-895.—FEMALE GENITALIA.
891. Sarata phi Heinrich, new species, type.
892. Sarata epsilon Heinrich, new species,
type.
893. Sarata perfuscalis (Hulst), type.
894. Sarata iota Heinrich, new species.
895. Philodema rhoiella (Dyar), paratype from
type locality.
522 UNITED: STATES NATIONAL MUSEUM BULLETIN 207
Ficures 896—900.—FEMALE GENITALIA.
896. Lipographis fenestrella (Packard).
897. Lipographis umbrella (Dyar).
898. Lipographis truncatella (Wright), speci-
men from San Diego, Calif.
899. Tota galdinella (Schaus), type.
900. Oryctometopia fossulatella Ragonot.
896. \2v
fenestrella galdinella
898.
truncatella
900.fossulatella
. umbrella
903. lithosella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 523
904.petrella
902
rubedinella
905.
Ficures 901-905.—FEMALE GENITALIA.
901. Adelphia ochripunctella (Dyar), showing
to the side of bursa a single spine of
the signum spine cluster, greatly en-
larged.
902. Ufa rubedinella (Zeller).
903. Ufa lithosella (Ragonot).
904, Adelphia petrella (Zeller).
905. Ufa senta Heinrich, new species,
524 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 906—910.— FEMALE GENITALIA,
906. Elasmopalpus lignosellus (Zeller).
907. Acroncosa albiflavella Barnes and Mc-
Dunnough.
908. Passadena flavidorsella (Ragonot).
909. Ulophora guarinella (Zeller), specimen
from Cuba.
910. [Myelozs] famula Zeller.
ry
TUNG
NRA 30720)
NCA
ty fi
Ke
908. flavidorsella 909.guarinella _ 910. famula
912.feriella
AMERICAN MOTHS OF THE
913. extrincica
914. idiotes
SUBFAMILY PHYCITINAE 525
Figures 911-914.—FeMALE GENITALIA,
911. [Nephopteryx] fuscifrontella Zeller, speci-
men from Colombia, with dorsal view
of ductus bursae shown in small sup-
plemental figure.
912. Tacoma feriella Hulst.
913. Chorrera extrincica (Dyar).
914. Chorrera idiotes Dyar.
526 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 915-919. Mat AND FEMALE GENI-
TALIA.
915. [Myelois] grosstpunctella Ragonot, male
genitalia, with aedeagus omitted; 915a,
aedeagus and anellus; 915b, gnathos;
915c, transtilla.
916. [Myelozs] grossipunctella Ragonot, female
genitalia.
917. [Elasmopalpus] corrientellus Ragonot,
type, female genitalia.
918. [Salebria] nigricans Hulst, type, female
genitalia.
919. [Hypochalcia] cervinistrigalis Walker,
type, female genitalia.
918. nigricans
917. corrientellus 919. cervinistrigalis
mysiella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 527
parvulella
aureofasciella
923.
dryadella
922.
| ochrella
Figures 920-925.— FEMALE GENITALIA.
920.
921.
922.
923.
924,
925.
Eumysia mysiella (Dyar), paratype from
type locality.
Divitiaca parvulella Barnes and McDun-
nough.
Divitiaca ochrella Barnes and McDun-
nough.
Ocala dryadella Hulst.
Macrorrhinia aureofasciella Ragonot.
Protasia mirabilicornella (Dyar).
528 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 926—931.—FEMALE GENITALIA.
926. Valdivia lativittella (Ragonot), from
Brownsville, Tex.
927. Valdivia lativittella (Ragonot), from
Pinal Mts., Ariz.
928. Heterographis samaritanella (Zeller),
bursa.
929. Heterographis morrisonella Ragonot;
929a, bursa of another example show-
ing extent of variation in its spining.
930. Staudingeria albtpenella (Hulst).
931. Hulstia undulatella (Clemens).
morrisonella
926,
lativittella
921.
lativitella
931. undulatella
930. albipenella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 529
cundajensis
semidiscella
934. banhoti ulmiarrosorella
Figures 932-936.—FEMALE GENITALIA,
932. Cabotia cundajensis (Zeller).
933. Cabotia rhythmatica Dyar, paratype.
934. Cabotia bonhoti (Hampson), specimen
from Nassau, Bahamas.
935. Cabotia semidiscella Ragonot.
936. Canarsia ulmiarrosorella (Clemens).
530 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 937—941.—FEMALE GENITALIA,
937. Honora mellinella Grote.
938. Honora dotella Dyar, specimen from San
Diego, Calif.
939. Honora montinaiatella (Hulst), type.
940. Honora perdubiella (Dyar), paratype
from type locality, showing to side of
bursa bases of some spines of signum
patch, greatly enlarged.
941. Honora subsciurella Ragonot, showing to
each side of bursa examples of spines,
greatly enlarged.
937.
mellinella
subsciurella
montinatatella
anticella
ignidorsella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 531
rubripurpurco
947.
Fuliginosus
Figures 942-947.—FEMALE GENITALIA.
942. Oncolabis anticella Zeller, specimen from
French Guiana; 942a, from Guatemala
(these two figures showing extremes of
variation).
943. Hurythmidia ignidorsella (Ragonot).
944, Oedothmia endopyrella Hampson, figured
from a sketch by J. F. G. Clarke, of
the type of its synonym Synothmia
bahamasella Hampson.
945. Eurythmasis ignifatua Dyar, paratype
from type locality.
946. Stylobasis rubripurpurea Hampson, speci-
men from Juan Vifias, Costa Rica.
947. Honorinus fuliginosus Heinrich: new
species, lateral view; 947a, ventral
view of genital opening.
532 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 948-951.—FEMALE GENITALIA.
948, 949. Wunderia neaeriatella Grossbeck, two
examples, showing extent of variation.
950. Cacozophera venosa Dyar, type.
951. [Honora] dulciella Hulst, an unplaced
species, type, showing to the side of Ss
bursa two projections of a signum,
greatly enlarged.
94.8.
neacriatella
949,
neaeriatella
950. venosa 951. dulciella
semicana,
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 533
nymphaeella
Fiaures 952-954.—FEMALE GENITALIA.
952. Psorosina hammondi (Riley).
953. Patriciola semicana Heinrich, new species.
954. Palatka nymphaeella (Hulst).
534 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 955-959.— FEMALE GENITALIA.
955, 956. Cassiana malacella (Dyar), two
specimens from Puerto Rico, showing
variations in signum.
957. Aptunga imperfecta (Dyar), type.
958. Aptunga macropasa (Dyar), type.
959. Anderida sonorella (Ragonot), type.
958.
macropasa
957. import a
rBeRneee a 1 vosg. aeieeelln 959. sonorella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 535
Ficures 960-965.—FEMALE GENITALIA.
960. Mescinia parvula (Zeller), with samples
of the spines of signum shown to the
side of bursa, greatly enlarged.
961. Mescinia commatella (Zeller), type, bursa
and part of ductus bursae.
962. Mescinia bacerella Dyar, specimen from
Sierra Maestra, Cuba., showing beside
bursa samples of spines of signum,
greatly enlarged.
963. Mescinia estrella Barnes and McDun-
nough, with samples of signum spines,
greatly enlarged, shown to the side of
bursa.
964. Mescinia peruella Schaus, with enlarge-
ment of some signum spines to the side
of bursa; 964a, bursa of another speci-
men showing variation in spining of
signum.
965. Mescinia triloses Dyar, with enlarged
samples of signum spines shown to
the side of bursa.
peruella
bacerella
965. triloses
963. estrella
536 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 966—-974.—FEMALE GENITALIA.
966. Mescinia berosa Dyar, type, with en-
larged samples of signum spines shown
to the side of bursa.
967. Mescinia discella Hampson.
968. Mescinia indecora Dyar, type, signum
greatly enlarged.
969. Nonia exiguella (Ragonot), specimen
from Panamé.
970. Phestinia costella Hampson; 970a, sig-
num of Puerto Rican example, greatly
enlarged; 970b, enlarged signum of
type.
971. Comotia sp., specimen from Puerto Rico.
972. Bema neuricella (Zeller); 972a-c, various
modifications of the signum, greatly
enlarged; 972d, signa, greatly enlarged,
of B. myja Dyar, a synonym of neuri-
cella.
973. Bema fifaca (Dyar), type, signum,
greatly enlarged.
974. Bema ydda (Dyar), type, signum, greatly
enlarged.
exiquella
costella
discella
fifaca
9724.
Comotia sp.
972.
neuricella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 537
oslarellum
980.emendator
sinuellum
electellum
982.inornatellum
eB
975a. opalescellum
%
976a.uncanale
983.
976.stypticellum albescentellum
300329—55——35
Ficures 975-983.— FEMALE GENITALIA.
975. Homoeosoma electellum (Hulst); 975a,
cephalic end of bursa and signum of
H. opalescellum (Hulst), a synonym of
electellum.
976. Homoeosoma stypticellum Grote, bursa;
976a, signum of type of H. uncanale
Hulst, a synonym of stypticellum.
977. Homoeosoma striatellum Dyar, bursa;
977a, signum from another specimen,
showing amount of variation.
978. Homoeosoma sinuellum (Fabricius), bursa.
979. Homoeosoma oslarellum Dyar, paratype,
bursa.
980. Homoeosoma illuviellum emendator Hein-
rich, new race, paratype from Rich-
field, Utah, bursa.
981. Homoeosoma impressale Hulst, bursa.
982. Homoeosoma inornatellum (Hulst), type,
bursa.
983. Homoeosoma aalbescentellum Ragonot,
type, bursa.
538
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 984—990.—FEMALE GENITALIA.
984.
985.
986.
987.
988.
989.
990.
Homoeosoma imitator Heinrich, new
species.
Homoeosoma deceptorium Heinrich, new
species, bursa.
Homoeosoma discrebile Heinrich, new
species, bursa.
Homoeosoma peregrinum Heinrich, new
species.
Homoeosoma assitum Heinrich, new spe-
cies.
Homoeosoma oconequensis (Dyar), type.
Homoeosoma ditaeniatellum Ragonot,
type.
assitum
988.
peregrinum
990.
deceptorium oconequensis ditaeniatellum
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 539
992.
994..mucidella
991.
vepallidum :
993. magellanella
Fiaures 991-994.— FEMALE GENITALIA.
991. Homoeosoma vepallidum Heinrich, new
species, paratype.
992. Homoeosoma vepallidum Heinrich, new
species, specimen from Sierra de
Cérdoba, Argentina.
993. Patagonia magellanella (Ragonot), type.
994, Rotruda mucidella mucidella (Ragonot);
994a, samples of spines of signum,
greatly enlarged and shown in two
projections.
540 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 995-1001.— FEMALE GENITALIA.
995. Unadilla erronella (Zeller); 995a en-
largements of individual signa.
996. Unadilla maturella (Zeller), specimen
from Guatemala.
997. Laetilia coccidivora (Comstock); 997a,
dorsal view of eighth-segment collar;
997b, enlargement of signum.
998. Laetilia coccidivora cardint Dyar, type;
998a, dorsal view of eighth-segment
collar.
999. Laetilia coccidivora quadricolorella
(Dyar); 999a, dorsal view of eighth-
segment collar.
1000. Laetilia obscura Dyar; 1000a, dorsal
view of eighth-segment collar.
1001. Laetilia portoricensis Dyar; 1001a, dor-
sal view of eighth-segment collar.
997. coccidivora
quadricolorella * 1000. obscura
1001. portoricensis
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE
10025
zamacrella
1004.
1008.
glabrella
eremiella
1006a,
1005: fiskella
1007. haywardi
541
Fiaures 1002-1008.—FEMALE GENITALIA.
1002.
1003.
1004,
1005.
1006.
1007.
1008.
Laetilia melanostathma (Meyrick);
1002a, dorsal view of eighth-segment
collar and apical portion of ductus
bursae.
Laetilia myersella Dyar, with enlarge-
ment of signum shown to the side of
bursa; 1003a, dorsal view of eighth-
segment collar.
Laetilia zamacrella Dyar.
Laetilia fiskella Dyar, with enlargement
of signum shown to the side of bursa.
Baphala basimaculatella (Ragonot), fig-
ured from its synonym Laetilia ere-
miella Dyar; 1006a, enlargement of
signum,
Baphala haywardi Heinrich, new spe-
cies, with enlargement of signum
shown to the side of bursa.
Baphala glabrella (Dyar), type, signum,
greatly enlarged.
542 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 1009-1015.— FEMALE GENITALIA. \ ty
1009a, olivacea
1009. Baphala goyensis (Ragonot), signa, goyensis
1010a.
showing extremes of variation.
1010. Baphala goyensis olivacea Heinrich, new
race; 1010a—d, various modifications
of the signum in this race.
1011. Baphala homoeosomella (Zeller): 1011—
1011a, Varieties of signa of female
genitalia in typical examples; 1011b-
1011f, varieties of signa in 24
synonym, sazssetiae (Dyar); 1011¢,
signum of another synonym, bodkinz
(Dyar); 1011h, apical part of ductus
bursae, collar, and ovipositor of
another synonym, rusto (Dyar).
1012. Rhagea packardella (Ragonot).
1013. Rhagea stigmella (Dyar).
1014. Rhagea maculicula (Dyar), a synonym of
R. stigmella (Dyar).
1015. Zophodia convolutella (Hiibner).
10llg.
bodkni
\
1010. olivacea
convolutella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAB 543
Figures 1016—1019.— FEMALE GENITALIA.
1016. Melitara prodenialis Walker; 1016a, dor-
sal view of eighth-segment collar.
1017. Melitara dentata (Grote).
1018. Olycella junctolineella (Hulst); 1018a,
dorsal view of eighth-segment collar.
1019. Olycella subumbrella (Dyar), paratype
from type locality, collar of eighth
abdominal segment, dorsal view.
(Explanation of symbols applied
to female genitalia: Bc, bursa copu-
latrix; Clr, collar of eighth abdominal
segment; Db, ductus bursae; dp, dor-
sal plates behind genital opening; Ds,
ductus seminalis; Go, genital open-
ing; Sm, signum.)
1017. dentata
2 P1015 subumbrelia
1018. junctolineella
544 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 1020—-1023.— FEMALE GENITALIA.
1020. Olyca phryganoides Walker; 1020a, dor-
sal view of eighth-segment collar.
1021. Alberada parabates (Dyar); 1021a, dor-
sal view of eighth-segment collar.
1022. Alberada bidentella (Dyar).
1023. Alberada holochlora (Dyar).
102].parabates
1022. bidentella
1023. holochlora
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 545
Figures 1024-1027.— FEMALE GENITALIA.
1024. Cactoblastis cactorum (Berg) ; 1024a, dor-
sal view of eighth-segment collar.
1025. Cactoblastis doddi Heinrich; 1025a, dor-
sal view of eighth-segment collar.
1026. Cactoblastis bucyrus Dyar; 1026a, dorsal
view of eighth-segment collar.
1027. Cahela ponderosella Barnes and McDun-
nough, paratype from type locality;
1027a, dorsal view of eighth-segment
collar.
1024. cactorum
1026.bucyrus 1027 ponderosella
300329—56——-86
546 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 1028—1032.— FEMALE GENITALIA.
1028. Rumatha glaucatella (Hulst), with
eighth-segment collar and ovipositor
omitted and with signum shown,
much enlarged, to the side of bursa.
1029. Rumatha polingella (Dyar), with eighth-
segment collar and ovipositor omitted.
1030. Rumatha bihinda (Dyar).
1031. Eremberga leuconips (Dyar); 1031a,
dorsal view of eighth-segment collar.
1032. Yosemitia longipennella (Hulst), with
eighth-segment collar and ovipositor
omitted and with signum, much en-
larged, shown to the side of bursa.
1028. glaucatella
1029. polingella
1030. bihinda
1032. longipennella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 547
Fiaures 1033-1037.—F MALE GENITALIA.
1033. Yosemitia graciella (Hulst).
1034. Tucumania tapiacola Dyar.
1035. Tucumania porrecia Dyar, with eighth-
segment collar and ovipositor omitted.
1036. Ozamia immorella (Dyar).
1037. Ozamia punicans Heinrich.
229099...
Ps $
APS
FITTS
3 392 3 Sy
3407 Ae
282 +;
Eeey
ELA
35328957
beers
2
1033.graciella 1034. tapiacola 1037. punicans
548
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fieures 1038-1043.—FEMALE GENITALIA.
1038.
1039.
1040.
1041.
1042.
1043.
Ozamia stigmaferella (Dyar), type;
1038a, dorsal view of eighth-segment
collar.
Ozamia thalassophila Dyar, type; 1039a,
dorsal view of eighth-segment collar.
Ozamia fuscomaculella clarefacta Dyar;
1040a, dorsal view of eighth-segment
collar.
Ozamia hemilutella Dyar; 1041a, dorsal
view of eighth-segment collar.
Ozamia lucidalis (Walker); 1042a, dor-
sal view of eighth-segment collar.
Ozamia fuscomaculella (Wright).
1040,
1038. stigmaferella clarafacta. fs:
‘1041a.
1041. hemilutella 1042. lucidalis
1043. fuscomaculella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 549
\ analamprella
substituta
1047a.
fernaldialis
1044. leithella 1047. huanucensis
Fiaures 1044-1048.— FEMALE GENITALIA.
1044,
1045.
1046.
1047.
1048.
Amalafrida leithella (Dyar) ; 1044a, dor-
sal view of eighth-segment collar.
Salambona analamprella (Dyar).
Sigelgaita chilensis Heinrich; 1046a,
dorsal view of eighth-segment collar,
Sigelgaita huanucensis Heinrich, type;
1047a, dorsal view of eighth-segment
collar,
Nanaia substituta Heinrich; 1048a, dor-
sal view of eighth-segment collar.
550 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 1049-1052.— FEMALE GENITALIA.
1049. Cactobrosis fernaldialis (Hulst); 1049a,
dorsal view of eighth-segment collar.
1050. Cactobrosis longipennella (Hampson),
specimen from Oaxaca, México;
1050a, dorsal view of eighth-segment
collar.
1051. Cactobrosis maculifera Dyar, eighth-seg-
ment collar and apical portion of duc-
tus bursae.
1052. Cactobrosis insignatella Dyar, type;
1052a, dorsal view of eighth-segment
collar.
SI
maculifera
1052. insignatella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 551
Figures 1053-1057.— FEMALE GENITALIA.
1053. Cactobrosis strigalis Barnes and Mc-
Dunnough.
1054. Illatila gurbyris Dyar, with enlarge-
ment of signum shown to the side of
bursa.
1055. Lascelina canens Heinrich, new species.
1056. Metephestia simplicula (Zeller).
1057. Selga arizonella (Hulst); 1057a, signum,
greatly enlarged.
1054. gurbyris
1053.
057.
arizonella
1056.
simplicula
1055. canens
552 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 1058—1062.— FEMALE GENITALIA.
1058. Entmemacornis proselytes Dyar.
1059. Anthropteryx irichampa Dyar, type.
1060. Moodnopsis portoricensis Heinrich, new
species.
1061. Moodnopsis inornatella (Ragonot).
1062. Moodnopsis decipiens Dyar.
irichampa
1059.
1061.
inornatella
1062. decipiens
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 553
Figures 1063-1067.—F eMALE GENITALIA.
1063. Huzophera cinerosella (Zeller).
1064, Euzophera semifuneralis (Walker).
1065. Huzophera ostricolorella Hulst.
1066. Huzophera nigricantella Ragonot.
1067. Prosoeuzophera impletella (Zeller), speci-
men from Puerto Rico (in Cornell
Univ.).
1064.
semifuneralis
1066.
1065. ostricolorella nigricantella 1067, impletella
554 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 1068-1071.— FEMALE GENITALIA.
1068. Vezina parasitaria Heinrich, new spe-
cies; 1068a, lateral view of part of
genitalia showing sclerotizations at or
near genital opening.
1069. Eulogia ochrifrontella (Zeller); 1069a,
enlargement of signum.
1070. Farnobia quadripuncta (Zeller); 1070a,
enlargement of signum.
1071. Edulica compedella (Zeller).
1068a.
ochrifrontella:
1068. parasitaria
1071.
compedella
1070. ppadeipancta!
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 555
Figures 1072-1078.— FEMALE GENITALIA.
1072, 1073. Ephestiodes gilvescentella Ragonot,
two examples, showing extent of vari-
ation.
1074. Ephestiodes erythrella Ragonot; 1074a-b,
outlines of two shields behind genital
opening, showing extent of variation.
1075. Ephestiodes mignonella Dyar, type.
1076. Ephestiodes indentella Dyar; 1076a, dor-
sal view of eighth-segment collar.
1077. Ephestiodes erasa Heinrich, new species;
10778, dorsal view of eighth-segment
collar.
1078. Ephestiodes productella Ragonot, type
(in Paris Mus.).
1072.
gilvescentella
1073.
_ gilvescentella
1074
erythrella.
1077a.
MUTT eccae 1078. productella
506
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Friaurss 1079-1085.— FEMALE GENITALIA.
1079.
1080.
1081.
1082.
1083.
1084.
1085.
Ephestiodes noniella Dyar.
Ephestiodes plorella Dyar; 1080a, dorsal
view of eighth-segment collar.
Ephestiodes lucidibasella Ragonot, para-
type (nm BM), apical (sclerotized)
part of ductus bursae.
Ephestiodes stictella (Hampson); 1082a,
dorsal portion of eighth-segment col-
lar; 1082b, apical portion of ductus
bursae, plate behind genital opening,
and apophyses of eighth-segment col-
lar.
Micromescinia pygmaea Dyar.
Azaera muciella Schaus; 1084a, dorsal
view of eighth-segment collar.
Azaera nodoses (Dyar), type; 1085a,
dorsal view of eighth-segment collar.
1080. plorella
Pygmaea
lucidibasella
1084. muciella
1085.nodoses
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 557
1088.
edmandsae serratilineella
1089. inanimella.
1087a.bisinuella
1090.pinei
YZ
Hy
1087. bisinuella
Figures 1086-1090.— FEMALE GENITALIA.
1086. Moodna ostrinella (Clemens); 1086a,
lateral view of ovipositor and inter-
segmental membrane with lobe, be-
tween ovipositor and eighth-segment
collar; 1086b, dorsal view of eighth-
segment collar.
1087. Moodna bisinuella Hampson; 1087a,
lateral view of caudal portion of geni-
talia, showing genital opening, eighth-
segment collar, intersegmental lobe
and ovipositor; 1087b, dorsal view of
eighth-segment collar.
1088. Vitula edmandsae serratilineella Rago-
not. Female genitalia.
1089. Vitula inanimella (Dyar), paratype from
Orizaba, México; 1089a, female geni-
talia figured from type of its synonym
Euzophera ticitoa Dyar.
1090. Vitula pinei Heinrich, new species.
558 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 1091-1097.—FEMALE GENITALIA.
1091. Vztula laura (Dyar), type.
1092. Manhaita biviella (Zeller).
1093. Manhatta setonella (McDunnough), para-
type (in USNM).
1094. Manhatia broweri Heinrich, new species,
paratype from type locality.
1095. [Eucampyla] putidella Schaus, type.
1096. Moodnella paula Heinrich, new species.
1097. Verina supplicella (Dyar), type.
1092. biviella
1091. laura:
1097.
1093. setonella 1094. broweri supplicella
1098.
pachytaeniella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 559
1099. trinitalis
Fiaures 1098-1100.—FrMALE GENITALIA.
1098. Volatica pachytaeniella (Ragonot).
1099. Volatica trinitatis Heinrich, new species.
1100. Vagobanta divergens (Butler).
l00.divergens
560 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Fiaures 1101—1105.— FEMALE GENITALIA.
1101. Caudellia colorella (Dyar), type.
1102. Caudellia nigrella (Hulst) ; 1102a, dorsal
view of eighth-segment collar.
1108. Caudellia albovittella Dyar.
1104. Caudellia declivella (Zeller), type.
1105. Soszpatra rileyella (Ragonot)
1102a. \
nigrella |
colorella ao
MOS a
albovittella
et We
ly
\ 2
loz. 7 N04.
rileyella nigrella declivella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 561
Ficures 1106-1114.—FEMALB GENITALIA.
1106. Sosipatra anthophila (Dyar).
1107. Sosipatra thurberiae (Dyar).
1108. Sosipatra majorella (Dyar).
1109. Sosipatra divergens (Dyar), type.
1110. Varneria postremella Dyar, type.
1111. Varneria nannodes Dyar, type.
1112. Varneria atrifasciella Barnes and Mc-
Dunnough.
1113. Varneria dubia Heinrich, new species.
1114. Microphestia animacula Dyar, type.
J109. divergens
H06.onthophila —_ 107. thurberiae 1108. majorella
14.
animacula
u10. ; : lil2.
postremella _lll.nannodes atrifasciella
113, dubia
562
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Figures 1115—-1120.— FEMALE GENITALIA.
1115.
1116.
1117.
1118.
1119.
1120.
Ribua innoxia Heinrich.
Ribua patriciella (Dyar), type.
Ribua contigua Heinrich, new species.
Plodia interpunctella (Hiibner).
Plodia dolorosa Dyar.
Bethulia championella Ragonot, type.
118. interpunctella
1119. dolorosa
1120.championella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 563
Figures 1121-1125.—FemMALE GENITALIA.
1121. Ephestia cautella (Walker).
1122. Ephestia elutella (Hiibner).
1123. Ephestia figulilella Gregson.
1124. Anagasta kihniella (Zeller) ; 1124a, some
variations in signa, enlarged.
1125. Nicetiodes apianella Schaus, paratype
from type locality.
121. cautella
1125. apianella
1124. kihniella
564 UNITED STATES NATIONAL MUSEUM BULLETIN 207
Ficgures 1126-1129.— FEMALE GENITALIA.
1126. [Huzophera] came Dyar, type; 1126a,
dorsal view of eighth-segment collar.
1127. [Moodna] formulella Schaus, type.
1128. [Euzophera] postflavida Dyar; 1128a, en-
largement of signum.
1129. [Euzophera] rinmea Dyar, type; 1129a,
dorsal view of eighth-segment collar.
1127. formulella
1126. came
29.
postflavida rinmea
1136, conops
1137. titillella
AMERICAN MOTHS OF THE SUBFAMILY PHYCITINAE 565
1133.fumella
1138. myronella
1134. quantulella
1135. pohellelle
Figures 1130-1138.—FrMALE GENITALIA.
1130.
1131.
1132.
1133.
1134.
1135.
1136.
1137.
1138.
Eurythmia hospitella (Zeller), figured
from type; 1130a, bursa of a female
from Texas, showing variation in
number of signa.
Eurythmia hospitella yavapaella Dyar,
bursa of a female from Glenwood
Springs, Colo.; 1131a, bursa of a fe-
male from San Diego, Calif.
Eurythmia angulella Ely, bursa.
Eurythmia fumella Ely, type, bursa.
Erelieva quantulella (Hulst), specimen
from Blanco County, Tex., showing
to the side of bursa a couple of signa,
greatly enlarged.
Erelieva parvulella (Ely), bursa.
Rabiria conops (Dyar), specimen from
type locality.
Microphycita titillella Dyar.
Cabnia myronella Dyar.
shies oe TAH SITE TERR
ana Ales ws
; my y i
r i az — R 4
Bey at as 5 Oh lds
Poe
iat ve (
+ Ta i * aaa :
F rp 5
: Ke rae |
r f
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Abie tke torau't CEH
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vibiaiedena at ihe
‘ es
Index to the genera and species of Phycitinae
(Page numbers of principal entries in boldface.
Synonyms in italics.
For species unknown, misplaced, or unrecognized, see page 580.)
abditiva Heinrich, n. sp., Meroptera, 123 | Acrobasis Zeller—Continued
aberrans Heinrich, n. sp., Protomoerbes,
49
abietella (Denis & Schiffermiiller), Di-
oryctria, 149, 150, 151, 152, 156, 233
abietivorella (Grote), Dioryctria, 150
abitus Heinrich, Sematoneura, n. sp., 27
acmaeopterum Ragonot, Homoeosoma, 225
Acrobasis Zeller, 1, 2, 10, 11, 25, 26, 37
albocapitella Hulst, 16
alnella McDunnough, 23
amplexella Ragonot, 13, 118
angusella Dyar, 16
angusella Grote, 19, 20
aurorella Ely, 18
betulella Hulst, 11, 22, 23
caryae Dyar (not Grote), 22
caryae Grote, 17, 18
caryaevorella (Dyar, not Ragonot), 17
earyalbella Ely, 15, 16
caryivorella Ragonot, 22, 24
cirroferella Hulst, 17, 21
comacornella (Hulst), 22
comptella Ragonot, 14, 24
comptoniella Grossbeck (not Hulst,) 23
comptoniella Hulst, 23
coryliella Dyar, 21
cunulae Dyar & Heinrich, 22
demotella Grote, 19
dyarella Ely, 20, 21
eliella Dyar, 19
evanescentella Dyar, 17
exsulella (Zeller), 19
feltella Dyar, 15, 17
grossbecki (Barnes & McDunnough),
13
hebescella (Dyar, not Hulst), 17
hebescella Hulst, 21
indigenella (Zeller), 12, 14
irrubriella Ely, 20
juglandis (LeBaron), 15, 16, 17
kearfottella Dyar, 17
latifasciella Dyar, 20
malipennella Dyar, 20
minimella Ragonot, 11, 15
myricella Barnes & McDunnough, 23
nebulella Dyar (not Riley), 16
nebulella (Grossbeck, not Riley), 13
nebulella (Riley), 12, 13
nebulo (Walsh), 12
nigrosignella Hulst, 15
normella Dyar, 20
ostryella Ely, 21
palliolella Dyar (not Ragonot), 16
palliolella Ragonot, 15, 16
peplifera Dyar, 18, 19
rubrifasciella Packard, 23
scitulella (Hulst), 14
secundella Ely, 21
septentrionella Dyar, 19
stigmella Dyar, 18
sylviella Ely, 16, 21
tricolorella Grote, 13, 14
tumidella (Zincken), 11, 17
tumidulella (Ragonot), 24
vaccinii Riley, 13
zelatella (Hulst), 12
zelleri Ragonot, 11
Acrocaula Hulst, 11, 12
Acromeseres Dyar, 176
Acroncosa Barnes & McDunnough, 3, 5,
174
albiflavella Barnes & McDunnough,
174
albiflavella castrella Barnes & McDun-
nough, 174
similella Barnes & McDunnough, 174
Actrix Heinrich, n. gen., 3, 5, 139
dissimulatrix Heinrich, n. sp., 140
nyssaecolella (Dyar), 139, 140
actualis (Hulst), Catastia, 110, 111
Adanarsa Heinrich, n. gen., 4, 35
intransitella (Dyar), 35
Adelperga Heinrich, n. gen., 182, 187
cordubensiella (Ragonot), 187, 196
Adelphia Heinrich, n., gen., 7, 168, 170
hapsella (Hulst), 169
ochripunctella (Dyar), 169
petrella (Zeller), 168, 169, 170
rubiginella (Walker), 169
rufinalis (Walker), 169
ademptandella (Dyar), Salebriaria, 115
admixta Heinrich, n. sp., Quasisalebria,
118, 119
adonea (Felder & Rogenhofer), Crocido-
mera, 33
advenella (Zincken), Rhodophaea, 24, 25
aeneella Hulst, Pyla, 147, 148
aeneoviridella Ragonot, Pyla, 147, 148
aenigmatica Heinrich, n. sp., Pyla, 144, 145
aequivoca Heinrich, n. sp., Pyla, 143, 145
afflictella (Hulst), Nephopteryx, 131
affusella (Ragonot) (race of mucidella
(Ragonot)), Rotruda, 219, 227
agapella Schaus, Fundella, 60, 61
aglaeella Ragonot, Euzophera, 273
ahemora Dyar, Fundella, 59, 62
ahenella (Zeller), Hypochalcia, 312
alatella (Hulst), Myelopsis, 41
Alberada Heinrich, 183, 244, 245
bidentella (Dyar), 244, 245
holochlora (Dyar), 245
parabates (Dyar), 244, 245
albescentellum Ragonot, Homoeosoma,
222, 223
albidiorella (Richards & Thomson), Una-
dilla, 229
albiflavella Barnes & McDunnough,
Acroncosa, 174
albipenella (Hulst), Staudingeria, 194, 195
albiplagiatella Hulst (not Packard), Pima,
102, 104
albiplagiatella (Packard), Pima, 103, 104
albiplagiatella (Ragonot, not Packard),
Pima, 104
albistrigella Staudinger, Myrlaea, 124
albocapitella Hulst, Acrobasis, 16
albocostalialis (Hulst), Pima, 103, 104
albocostalis (Hulst), Pima, 104
albocostella Hulst, ‘“Maricopa,’”’ 192, 316
albosigno Heinrich, n. sp., Diatomocera, 52
albovittella Dyar, Caudellia, 292
albovittella (Hulst), Dioryctria, 127, 156
aliculella (Hulst), Olybria, 113, 114
almonella (Dyar), Praedonula, 82, 83
alnella McDunnough, Acrobasis, 23
alpha Heinrich, n. sp., Megarthria, 88
alpha Heinrich, n. sp., Sarata, 164
alternosquamella Ragonot, Dasypyga, 69
alveolella (Ragonot), Moerbes, 269
Amalafrida Heinrich, 183, 255, 256
leithella (Dyar), 256
amarella Dyar, Ephestia, 302
amatella (Hulst), Dioryctria, 156
Ambesa Grote, 7, 108, 110
laetella Grote, 108
lallatalis (Hulst), 109
monodon Dyar, 108
walsinghami (Ragonot), 108, 109
walginghami mirabella Dyar, n. sta-
tus, 109
amphimetra (Meyrick), Laetilia, 230, 233
Amphycitopsis Dyar, 78, 80
amplexella Ragonot, Acrobasis, 13, 118
Anabasis Heinrich, n. gen., 2, 25
crassisquamella (Hampson), 26
ochrodesma (Zeller), 25, 26
Anadelosemia Dyar, 3, 65, 67, 69, 72, 73
base Dyar, 68
condigna Heinrich, n. sp., 69
dulcella (Hulst), 68
fifria Dyar, 68
obstitella (Schaus), 68
senesciella (Schaus), 67, 68, 69
567
568
Anadelosemia Dyar—Continued
tecmessella (Schaus), 67
texanella (Hulst), 68, 69
Anagasta Heinrich, n. gen., 186, 299
fuscofasciella (Ragonot), 300
gitonella (Druce), 300
kiihniella (Zeller), 291, 299, 300, 302
sericaria authors (not Scott), 300
analamprella (Dyar), Salambona, 254
Ancylostomia Ragonot, 9, 95, 171
argyrophleps Dyar, 96
diffissella (Zeller), 95
euchroma Dyar, 96
ignobilis (Butler), 95
sauciella (Zeller), 96
stercorea (Zeller), 95, 96
Anderida Heinrich, n. gen.,
213
placidella (Dyar), 211
senorella (Ragonot), 211
sonorella (Ragonot), 211
Anegcephalesis Dyar, 8, 93
arctella (Ragonot), 94
cathaeretes Dyar, 93, 94
Anerastiinae, 1
angulella Dyar, Psorosina, 208, 209
angulella Ely, Eurythmia, 307, 308
angulinella (Schaus), Hyalospila, 58
angusella Dyar, Acrobasis, 16
angusella Grote, Acrobasis, 19, 20
angustellus Blanchard, Elasmopalpus, 172,
173
animaleula Dyar, Microphestia, 294
animosella (Dyar), Caudellia, 293
annulatella (Zetterstedt), Polopeustis, 120,
121
annuliferella (Dyar), Chararica, 38, 39
annulosella (Ragonot), Salebriaria,
117
anthophila (Dyar), Sosipatra, 295
anthracellus Ragonot, Elasmopalpus, 173
anticella Zeller, Oncolabis, 199, 206
Anypsipyla Dyar, 8, 42
univitella Dyar, 42
Anthropteryx Dyar, 208, 313
apianella Schaus, Nicetiodes, 304, 314
apocerastes Dyar, Piesmopoda, 78, 80,
81
Apomyelois Heinrich, n. gen., 8, 42, 44
bilineatella (Ragonot), 43
bistriatella (Hulst), 42, 43
Aptunga Heinrich, n. gen., 181, 211, 212,
213
imperfecta (Dyar), 211
macropasa (Dyar), 211
apyrella Dyar, Caudellia, 292
arctella (Ragonot), Anegcephalesis, 94
arctiella (Gibson), Polopeustis, 121
182, 211,
116,
argentina Dyar, Fundella, 55, 59, 60,
61, 62
argentinensis Heinrich, n. sp., Stylo-
palpia, 141
argyrophleps Dyar, Ancylostomia, 96
aridella (Dyar), Sarata, 161
arizonella (Hulst), Selga, 265
arizonella (Walter), Caudellia, 293
arizonensis Heinrich, n. sp., Pseudo-
cabima, 55
assitum Heinrich, n. sp., Homoeosoma,
225
asthenosoma (Dyar), Parolyca, 255
Atheloca Heinrich, n. gen., 7, 81,'83
bondari Heinrich, n. sp., 77, 82
filiolella (Hulst), 82
ptychis Bondar (not Dyar), 82
ptychis (Dyar), 82
subrufella (Hulst), 81, 82, 33
atrella (Hulst), Sarata, 2, 162, 164, 165
atrifasciella Barnes & McDunnough, Var-
neria, 305, 306
atristrigella Ragonot, ‘‘Myelois,’”’ 315
atrovenosella Ragonot, Sematoneura, 27,
51
auranticella (Grote), Dioryctria, 152, 153
aureofasciella Ragonot, Macrorrhinia, 190
aureomaculella (Dyar), Valdivia, 192
aurorella Ely, Acrobasis, 18
australella (Hulst), Zamagiria, 92, 171
austriana (Cosens), Dioryctria, 155
Azaera Schaus, 180, 185, 282
lophophora (Dyar), 283
muciella Schaus, 282, 283
nodoses (Dyar), 283
Azaera squalidella (Dyar), 282
azonaxsalis Walker, Davara, 73, 75
bacerella Dyar, Mescinia, 213, 214
bahamasella (Hampson), Oedothmia, 205
bakerella (Dyar), Salebriacus, 114
Ballovia Dyar, 59
Bandera Ragonot, 1, 315, 316
Baphala Heinrich, n. gen., 181, 234, 235
basimaculatella (Ragonot), 235, 236
bodkini (Dyar), 236
eremiella (Dyar), 235
glabrella (Dyar), 237
goyensis (Ragonot), 235, 236
goyensis olivacea Heinrich, n. race, 236
haywardi Heinrich, n. sp., 236
homoeosomella (Zeller), 235, 236, 237
rusto (Dyar), 236
saissetiae (Dyar), 236
squalida (Walker), 236, 237
taboga (Dyar), 236
base Dyar, Anadelosemia, 68
basilaris Zeller, Nephopteryx, 129
basimaculatella (Ragonot), Baphala, 235,
236
baumhoferi Heinrich, n. sp., Dioryctria, 157
bella Hulst, Zophodia, 239
Bema Dyar, 180, 185, 217
fifaca (Dyar), 218, 219
fritilla Dyar, 218
myja Dyar, 217, 218, 219
neuricella (Zeller), 218
ydda (Dyar), 217, 218
yddiopsis (Dyar), 219
benjaminella Dyar, Ephestiodes, 279
berosa Dyar, Mescinia, 214
Bertelia Barnes & McDunnough, 4, 11, 36
grisella Barnes & McDunnough, 36, 37
beta Heinrich, n. sp., Megarthria, 88
UNITED STATES NATIONAL MUSEUM BULLETIN 207
beta Heinrich, n. sp., Sarata, 164
Bethulia Ragonot, 186, 296, 298
championella Ragonot, 296, 297
betulella Hulst, Acrobasis, 11, 22, 23
bicolorella (Barnes & McDunnough), Cha-
rarica, 39
bidentella (Dyar), Alberada, 244, 245
bifasciella Hulst, Nephopteryx, 130
bigrana (Zeller), Hemiptilocera, 30, 31
bihinda (Dyar), Rumatha, 248, 249
bilineatella (Ragonot), Apomyelois, 43
bipunctella (Hampson), Unadilla, 228
Birinus Heinrich, n.-gen., 4, 11, 36
russeolus Heinrich, n: sp., 36
bisinuella Hampson, Moodna, 284
bisra Dyar, Nephopteryx, 133
bistriatella (Hulst), Apomyelois, 42, 43
bistriatella (Hulst), Catastia, 110, 111
bistriga (Haworth), Cryptoblabes, 10
biviella (Zeller), Manhatta, 272, 284, 287
blackmorella Dyar, Pyla, 145
bodkini (Dyar), Baphala, 236
boisduvaliella (Guénée), Pima, 102, 103,
105
bollaz Dodd (not Zeller), Melitara, 241
bolliz (Zeller), Melitara, 240
bondari Heinrich, n. sp., Atheloca, 77, 82
bonhoti (Hampson), Cabotia, 200, 201
breviplicitum Heinrich, n. race (race of
oslarellum Dyar), Homoeosoma, 221
brevistrigella Ragonot, ‘‘Zophodia,” 314
broweri Heinrich, n. sp., Manhatta, 288
brucei (Hulst), Phobus, 110, 138, 139
brunneella (Dyar), Dioryctria, 158
bucyrus Dodd (not Dyar), Cactoblastis, 246
bucyrus Dyar, Cactoblastis, 247
bumeliella (Barnes & McDunnough), Zama-
giria, 92
burdetellus (Schaus), Peadus, 83, 84
busckella (Dyar), Glyptocera, 101
Cabima Dyar, 50
Cabnia Dyar, 310
myronella Dyar, 310
Cabotia Ragonot, 180, 200
bonhoti (Hampson), 200, 201
cundajensis (Zeller), 201
impeditella (Zeller), 201
rhythmatica Dyar, 200, 201
schini (Berg), 201
semidiscella Ragonot, 200, 201
cacabella (Hulst), Pyla, 145
Cacozophera Dyar, 185, 208
venosa Dyar, 208
Cactoblastis Ragonot, 1, 183, 245
bucyrus Dodd (not Dyar), 246
bucyrus Dyar, 247
cactorum (Berg), 245, 246, 247, 257
doddi Heinrich, 246, 247
mundelli Heinrich, 247
ronnai (Bréthes), 245, 246
Cactobrosis Dyar, 183, 233, 240, 257, 260
cinerella (Hulst), 260
elongatella (Hampson), 261
fernaldialis (Hulst), 260, 261
gigantella (Ragonot), 260.
Cactobrosis Dyar—Continued
insignatella Dyar, 260, 261
longipennella (Hampson), 260, 261
maculifera Dyar, 260, 261
strigalis (Barnes & McDunnough),
248, 253, 260, 262
cactorum (Berg), Cactoblastis, 245, 246,
247, 257
Cadra Walker, 301
Cahela Heinrich, 182, 184, 247, 248, 249
interstitialis (Dyar), 248
phoenicis (Dyar), 248
ponderosella (Barnes & McDunnough),
247, 248, 249
purgatoria (Dyar), 248
cahiritella Zeller, Ephestia, 303
Calamophleps Dyar, 282
caliginella (Hulst), Rhodophaea, 15, 24, 25
caliginoidella (Dyar), Rhodophaea, 24
cambiicola (Dyar), Dioryctria, 150, 155,
156, 157
came Dyar, ‘‘Euzophera,’’ 313.
Campyloplesis Dyar, 269
Canarsia Hulst, 181, 201
feliculella Dyar, 202
fuscatella (Hulst), 202
gracilella Hulst, 202
pneumatella (Hulst), 202
ulmella (Ragonot), 202
ulmiarrosorella (Clemens), 201, 202
candidella Hulst, Homoeosoma, 221
canens Heinrich, n. sp., Lascelina, 264
canescentelia (Hulst), Passadena, 175
canicostella Ragonot, Honora, 198
carabayella Dyar, Pseudodivona, 48, 49
carbonella (Hulst), Elasmopalpus, 173
eardini Dyar (race of coccidivora (Com-
stock), Laetilia, 230, 232
earicae (Dyar), Davara, 73, 74, 75
Caristanius Heinrich, n. gen., 5, 97
decoloralis (Walker), 98
floridellus (Hulst), 98
furfurella (Hulst), 98
guatemalellus (Ragonot), 98
melanoplaga (Hampson), 97
metagrammalis (Walker), 98
pellucidellus (Ragonot), 97, 98
carneella Hulst, Nephopteryx, 128
carpasella (Schaus), Fulrada, 72
caryae Dyar (not Grote), Acrobasis, 22
caryae Grote, Acrobasis, 17, 18
caryaevorella (Dyar, not Ragonot), Acro-
basis, 17
caryalbella Ely, Acrobasis, 15, 16
caryivorella Ragonot, Acrobasis, 22 °4
cassiae (Dyar), Paramyelois, 47
Cassiana Heinrich, n. gen., 181, 212, 213
Cassiana malacella (Dyar), 212
eastrella Barnes & McDunnough (race of
albiflavella Barnes & McDunnough),
Acroncosa, 174
castronalis Heinrich, n. sp., Pseudocabima,
53, 54
Catastia Hiibner, 7, 110, 112
actualis (Hulst), 110, 111
bistriatella (Hulst), 110, 111
300329—56——37
INDEX
Catastia Hiibner—Continued
incorruscella (Hulst), 110, 111
marginea (Schiffermiiller), 110
cathaeretes Dyar, Anegcephalesis, 93, 94
caudellella (Dyar), Sarata, 162, 165
Caudellia Dyar, 186, 292, 294, 298
albovittella Dyar, 292
animosella (Dyar), 293
apyrella Dyar, 292
arizonella (Walter), 293
clara Heinrich, n. sp., 294
colorella (Dyar), 293
declivella (Zeller), 292, 293, 294
nigrella (Hulst), 283, 293
cautella (Walker), Ephestia, 298, 302, 303,
304 :
Cayennia Hampson, 182, 267
rufitinctalis Hampson, 267
celiella Schaus, Hyalospila, 57
celtidella (Hulst), Nephopteryx, 131, 132
Ceracanthia Ragonot, 4, 85, 86
mamella (Dyar), 86
vepreculella Ragonot, 86
ceratoniae (Zeller), Ectomyelois, 44
cervicalis Dyar, Megarthria, 86, 87
cervinistrigalis Walker, ‘Hypochalcia,”’
312
championella Ragonot, Bethulia, 296, 297
Chararica Heinrich, n. gen., 5, 38
annuliferella (Dyar), 38, 39
bicolorella (Barnes & McDunnough),
39
hystriculella (Hulst), 39
chilensis Heinrich, Sigelgaita, 255
chinographella Ragonot, Hemiptilocera,
30, 31
Chorrera Dyar, 6, 7, 177
extrincica (Dyar), 177, 178
idiotes Dyar, 177, 178
postica (Zeller), 178
cinctella (Hulst), Passadena, 175
cinereella Hulst, Sarata, 162, 163
cinerella (Hulst), Cactobrosis, 260
cinerosella (Zeller), Euzophera, 272, 273
cinilixa Dyar, Drescoma, 89
cirroferella Hulst, Acrobasis, 17, 21
cispha Dyar, Pseudodivona, 48, 49
cistipennis (Dyar), Fundella, 59, 60
clara Heinrich, n. sp., Caudellia, 294
clarefacta Dyar (race of fuscomaculella
(Wright)), Ozamia, 258
clarioralis (Walker), Dioryctria, 150, 158
clevelandella (Dyar), Hyalospila, 58
climosa Dyar, ‘‘Euzophera,” 315
clitellatella Ragonot, ‘Hornigia,” 284,
314
cnabella Dyar, Hypsipyla, 28
coca (Dyar), Erelieva, 308, 309
coccidivora (Comstock), Laetilia, 230,
231, 232, 254
coloradella (Hulst), Ephestiodes, 279, 280
coloradensis Ragonot, Heterographis, 194
colorella (Dyar), Caudellia, 293
columbiella (McDunnough), Interjectio,
106, 107
columnella (Zeller), Davara, 74, 75
569
comacornella (Hulst), Acrobasis, 22
commatella (Zeller), Mescinia, 212, 214
commensella Dyar, Pseudodivona,
49, 50
Comotia Dyar, 185, 217
convergens (Dyar), 217
torsicornis Dyar, 217
compedella (Zeller), Edulica, 271, 272
comptella Ragonot, Acrobasis, 14, 24
comptoniella Grossbeck (not Hulst), Acro-
basis, 23
comptoniella Hulst, Acrobasis, 23
condigna Heinrich, n. sp., Anadelosemia, 69
coniella (Ragonot), Myelopsis, 40, 41, 265
conops (Dyar), Rabiria, 311
conquistador Dyar, ‘‘Euzophera,” 315
consobrinella (Zeller), Glyptocera, 100, 101
consociata Heinrich, n. race (race of par-
vulella Barnes & McDunnough),
Divitiaca, 190
constaniella Hulst, Passadena, 175
constitutionella Dyar, Mildrixia, 26
contatella (Grote), Nephopteryx, 124, 127,
128
contigua Heinrich, n. sp., Ribua, 297
convergens (Dyar), Comotia, 217
convolutella (Hiibner), Zophodia, 238, 239
Coptarthria Ragonot, 3, 62, 63, 64, 66, 70
dasypyga (Zeller), 64, 65
coquilla (Dyar), Erelieva, 308
coquimbella Ragonot, Valdivia, 191, 192
cordubensiella (Ragonot), Adelperga, 187,
196
corniculatus Heinrich, n. sp., Paconius, 210
corrientellus Ragonot, ‘‘Hlasmopalpus,”’
312
coryliella Dyar, Acrobasis, 21
corynophora Dyar, Difundella, 62, 63
costella Hampson, Phestinia, 216
crassifasciella Ragonot, Nephopteryx, 5,
132
crassisquamella (Hampson), Anabasis, 26
crataegella Barnes & MceDunnough, Ne-
phopteryx, 132
creabates (Dyar), Eremberga, 253
cribrella Hiibner, Myelois, 40, 43
eriddlella Dyar, Pyla, 144, 145
cristalis (Hampson), Magiriopsis, 94
croceella (Hulst), Nephopteryx, 123, 131
Crocidomera Zeller, 4, 32, 34, 276
adonea (Felder & Rogenhofer), 33
fissuralis (Walker), 33
stenopteryx (Dyar), 33
turbidella Zeller, 32, 33
Cryptoblabes Zeller, 1, 10
gnidiella (Millitre), 10
bistriga (Haworth), 10
gnidiella (Milliére), 10
rutilella Zeller, 10
Cuba Dyar, 76
cubella Dyar, Homoeosoma, 219, 226
cubensis Heinrich, n. sp., Scorylus, 72, 73
cundajensis (Zeller), Cabotia, 201
Cuniberta Heinrich, n. gen., 5, 34
subtinctella (Ragonot), 34
cunulae Dyar & Heinrich, Acrobasis, 22
48,
570
curvatellus (Ragonot), Phobus, 137, 139
cviatella Dyar, Meroptera, 122
cyrdipsa Dyar, Drescoma, 88, 89, 263
daedalella Ragonot, “‘Euzophera,” 314
Dakruma Grote, 238
dalera (Dyar), Davara, 73, 74
dammersi Heinrich, n. sp., Nephopteryx,
126, 127
Dannemora Hulst, 206, 276
Dasypyga Ragonot, 4, 69
alternosquamella Ragonot, 69
stictophorella Ragonot, 69
dasypyga (Zeller), Coptarthria, 64, 65
dasystigma Dyar, Promylea, 67
Davara Walker, 2, 9, 73, 77, 78
azonaxsalis Walker, 73, 75
caricae (Dyar), 73, 74, 75
columnella (Zeller), 74, 75
dalera (Dyar), 73, 74
euthales (Dyar), 75
interjecta Heinrich, n. sp., 73, 74, 75
nerthella (Schaus), 75
paranensis (Dyar), 75
rufulella (Ragonot), 73, 76
deceptorium Heinrich, n. sp., Homoeo-
soma, 223
decimerella (Hulst), Stylopalpia, 141
decipiens Dyar, Moodnopsis, 269, 270, 271
decipientella Dyar, Nephopteryx, 132
declivella (Zeller), Caudellia, 292, 293, 294
decolor (Zeller), Ectomyelois, 43, 44
decoloralis (Walker), Caristanius, 98
decuriella (Hiibner), Dioryctria, 150
decurrens (Dyar), Diatomocera, 51
defectella (Walker), Ephestia, 301, 303
definitella (Zeller), Hypargyria, 37, 38
dea Dyar, Zamagiria, 93
delassalis Hulst, Nephopteryx, 125, 127
delassalis (Hulst, not Hulst), Nephopteryx,
125, 126
delectella}(Hulst), Dioryetria, 155
delta Heinrich, n. sp., Sarata, 165
demotella Grote, Acrobasis, 19
dentata (Grote), Melitara, 241
denticosella (Dyar), Magiriopsis, 94
denticulella (Ragonot), Interjectio,
107, 110, 138
dentosella Ragonot, Vitula, 285
deprivalis (Walker), Ufa, 171
desuetella (Walker), Ephestia, 303
dialithus (Dyar), Ulophora, 176, 177
Diatomocera Ragonot, 8, 50, 53, 266
albosigno Heinrich, n. sp., 52
decurrens (Dyar), 51
dosia (Dyar), 51
excisalis (Hampson), 51
extracta Heinrich, n. sp., 52, 53
hoplidice (Dyar), 52
majuscula Heinrich, n. sp., 52
mochlophleps (Dyar), 52
tenebricosa (Zeller), 50
didactica Dyar, Yosemitia, 250, 251
differtella Barnes & McDunnough, Homo-
eosoma, 220
diffissella (Zeller), Ancylostomia, 95
106,
UNITED
STATES NATIONAL MUSEUM BULLETIN 207
diffusella Ely, Eurythmia, 306, 307
Difundella Dyar, 3, 60, 62, 263
corynophora Dyar, 62, 63
distractor Heinrich, n. sp., 64
subsutella (Schaus), 63
tolerata Heinrich, n. sp., 64, 65
dilatifasciella (Ragonot), Laetilia, 230
dilativitia Dyar, Zophodia, 239
Dioryctria Zeller, 8, 66, 124, 149, 191, 233
abietella (Denis & Schiffermiiller), 149,
150, 151, 152, 156, 233
abtetivorella (Grote), 150
albovittella (Hulst), 127, 156
amatella (Hulst), 156
auranticella (Grote), 152, 153
austriana (Cosens), 155
baumhoferi Heinrich, n. sp., 157
brunneella (Dyar), 158
cambiicola (Dyar), 150, 155, 156, 157
clarioralis (Walker), 8, 150, 158
decuriella (Hiibner), 150
delectella (Hulst), 155
disclusa Heinrich, 152, 153
elegentella (Hulst), 150
erythropasa (Dyar), 152, 153
gulosella (Hulst), 149, 157, 158
horneana (Dyar), 154
laurata (Heinrich), 153
majorella Dyar, 152
mendacella Staudinger, 149
miniatella Ragonot, 153
muellerana Dyar, 152
pineae Staudinger, 149
ponderosae Dyar, 152
ponderosae Heinrich (not Dyar), 155,
156
pryeri Ragonot, 153
pygmaeella Ragonot, 154
reniculella (Grote), 151, 152
reniculella (Packard, not Grote), 150
subtracta Heinrich, n. sp., 157, 158
sysstratiotes Dyar, 151, 152
zanthaenobares Dyar, 153
zimmermani (Grote), 149, 150, 152,
154, 155, 156, 157
discella Hampson, Mescinia, 213, 215
disclusa Heinrich, Dioryctria, 152, 153
Discopalpia Ragonot, 78, 79
discrebile Heinrich, n. sp., Homoeosoma,
224
disjunctus Heinrich, n. sp., Heras, 34, 35
dissimulatrix Heinrich, n. sp., Actrix, 140
dissitus Heinrich, n. sp., Peadus, 84
disticta (Zeller), ‘‘Psorosa,”’ 314
distractor Heinrich, n. sp., Difundella, 64
ditaeniatellum Ragonot, Homoeosoma, 224
divergens (Butler), Vagobanta, 289
divergens (Dyar), Sosipatra, 296
Diviana Ragonot, 185, 206, 207
edentella (Hulst), 206, 207
eudoreella Ragonot, 206, 207
Divitiaca Barnes& McDunnough, 185,
189, 190
ochrella Barnes & McDunnough, 189,
190
parvulella Barnes & McDunnough, 190
Divitiaca Barnes & McDunnough—Cont.
parvulella consociata, Heinrich, n.
race, 190
simulella Barnes & McDunnough, 189,
190
Divona Ragonot, 166
ilignella (Zeller), 166
dixolophella Dyar, Zamagiria, 90, 91
dnopherella Ragonot, Sarata, 159, 162, 163,
164, 165
doddalis Dyar, Melitara, 241
doddi Heinrich, Cactoblastis, 246, 247
Dolichorrhinia Ragonot, 190
dolorosa Dyar, Plodia, 298, 299
dorae (Dyar), Unadilla, 227, 228
dorsimacula (Schaus), Hypsipyla, 28, 29
dosia (Dyar), Diatomocera, 51
dotella Dyar, Honora, 197, 198
Drescoma, Dyar, 2, 88, 262, 266
cinilixa Dyar, 89
cyrdipsa Dyar, 88, 89, 263
Drescomopsis Dyar, 88, 180, 262, 266
drucella (Dyar), 263
soraella (Druce), 263
subelisa Dyar, 262, 263
drucet (Dyar), Promylea, 66
druceii (Ragonot), Promylea, 66
drucella (Dyar), Drescomopsis, 263
dryadella Hulst, Ocala, 191
dryopella Dyar (not Schaus), Moerbes 269
dryopella (Schaus), Moerbes, 268, 269
dubia Heinrich, n. sp., Varneria, 306
dulciella Hulst, ‘“Honora,’’ 197, 313
dulciella (Hulst), Anadelosemia, 68
duplipunctella (Ragonot), Paramyelois, 47
dyarella Ely, Acrobasis, 20, 21
dyari Heinrich, n. name, Promylea, 66, 67
Eccopsia Hulst, 285
Ectomyelois Heinrich, n. gen., 8, 43
ceratoniae (Zeller), 44
decolor (Zeller), 43, 44
ephestiella (Hampson), 44
furvidorsella (Ragonot), 45, 46
muriscis (Dyar), 45, 46, 47, 48
oporedestella (Dyar), 44
palpalis (Dyar), 45
phoenicis (Durrant), 45
transitella (Dyar, not Walker), 45
zeteki Heinrich, n. sp., 46
edentella (Hulst), Diviana, 206, 207
edmandsae (Packard), Vitula, 285, 286,
287, 288
Edulica Ragonot, 182, 271
compedella (Zeller), 271, 272
edwardsialis (Hulst), Sarata, 160
egenella (Ragonot), Hyalospila, 58
Elasmopalpus Blanchard, 7, 97, 169, 170,
172, 204, 312
angustellus Blanchard, 172, 173
anthracellus Ragonot, 173
carbonella (Hulst), 173
incautella (Zeller), 173
lignosellus (Zeller), 172, 173, 204
major (Zeller), 173
Elasmopalpus Blanchard—Continued
puer Dyar, 173
tartarella (Zeller), 173
electellum (Hulst), Homoeosoma, 220, 223,
224
elegentella (Hulst), Dioryctria, 150
eliella Dyar, Acrobasis, 19
elongatella (Hampson), Cactobrosis, 261
elongellum Dyar, Homoeosoma, 222
elutea (Haworth), Ephestia, 302
elutella (Hitbner), Ephestia, 227, 301, 302,
303, 304
emendata Heinrich, n. sp., Moerbes, 50,
268, 269
emendator Heinrich n. race (variety of
illuviellum Ragonot), Homoeosoma,
222
Emmerita Hampson, 121
Encystia Hampson, 200
Endommasis Hampson, 199
endopyrella Hampson, Oedothmia, 205
engeli (Dyar), Salebriaria, 116
Entmemacornis, Dyar, 50, 53, 181, 266,
267
proselytes Dyar, 266, 267
pulla Heinrich, n. sp., 266
Ephestia Guénée, 10, 186, 202, 228, 279,
284, 287, 293, 299, 300, 301
amarella Dyar, 302
cahiritella Zeller, 303
cautella (Walker), 298, 302, 303, 304
defectella (Walker), 301, 303
desuetella (Walker), 303
elutea (Haworth), 302
elutella (Hiibner), 227, 301, 302, 303,
304
ernestinella Turati, 304
ficulella Barrett, 304
figulella Curran, 304
figulilella Gregson, 302, 303, 304
formosella (Wileman & South), 303
millert Zeller, 304
passulella Barrett, 303
roxzburghit Gregson, 302
rufa (Haworth), 302
semirufa (Haworth), 302
sericarium (Scott), 301, 302
unicolorella Staudinger, 302
venosella Turati, 304
vitivora Filipjev, 302
ephestiella (Hampson), Ectomyelois, 44
ephestiella (Ragonot), Laetilia, 230, 234
Ephestiodes Ragonot, 185, 277, 278, 282,
306
benjaminella Dyar, 279
coloradella (Hulst), 279, 280
erasa Heinrich, n. sp., 280
erythrella Ragonot, 279, 280
gilvescentella Ragonot, 278, 279, 280,
281, 282
granulella Hampson, 281
indentella Dyar, 281
infimella Ragonot, 279, 280, 281
lucidibasella Ragonot, 278, 280
mignonella Dyar, 280
nigrella Hulst, 278, 283
INDEX
Ephestiodes Ragonot—Continued
noniella Dyar, 278, 282, 306
571
Eutrichocera Hampson, 315
paurolepidalis Hampson, 315
plorella Dyar, 278, 281, 282, 306, 309 | Euzophera Zeller, 181, 207, 233, 272, 274,
productella Ragonot, 281
stictella (Hampson), 278, 281, 306
uniformella Hampson, 281
vestilla (Dyar), 281, 309
Epischnia authors, 101, 102
Epischnia Hiibner, 102
prodromella Hiibner, 102
epischnioides Hulst, ‘‘Zophodia,” 315
epsilon Heinrich, n. sp., Sarata, 165
275, 276, 287, 313
aglaeella Ragonot, 273
cinerosella (Zeller), 272, 273
griselda Dyar, 274
nigricantella Ragonot, 55, 274
ostricolorella Hulst, 273, 274
pallulella (Hulst), 273
pinguis (Haworth), 272, 273
semifuneralis (Walker), 273, 276
erasa Heinrich, n. sp., Ephestiodes, 280 | euzopherella (Dyar), Pseudocabima, 53, 54
Erelieva Heinrich, n. gen., 186, 307, 308 | evanescentella Dyar, Acrobasis, 17
coca (Dyar), 308, 309
coquilla (Dyar), 308
hospitella (Dyar, not Zeller), 308
mossa (Dyar), 308
parvulella (Ely), 307, 308, 309
quantulella (Hulst), 307, 308, 309
santiagella (Dyar), 308
uncta (Dyar), 308
Eremberga Heinrich, 183, 252, 253
creabates (Dyar), 253
insignis Heinrich, 254
leuconips (Dyar), 248, 253, 262
eremiella (Dyar), Baphala, 235
ernestinella Turati, Ephestia, 304
erronella (Zeller), Unadilla, 228, 229
erythrella Ragonot, Ephestiodes, 279, 280
erythropasa (Dyar), Dioryctria, 152, 153
estrella Barnes & McDunnough, Mescinia,
213
Etiella Zeller, 1, 2, 98, 99
etiella (Treitschke), 99
rubribasella Hulst, 99
schisticolor Zeller, 99
villosella Hulst, 99
zinckenella (Treitschke), 99, 238
etiella (Treitschke), Etiella, 99
Eucardinia Dyar, 73, 74
eucasis Dyar, Fundella, 61
euchroma Dyar, Ancylostomia, 96
eudoreella Ragonot, Diviana, 206, 207
Eulogia Heinrich, n. gen., 181, 275
ferruginella (Ragonot), 275
ochrifrontella (Zeller), 275
Eumysia Dyar, 182, 187
fuscatella (Hulst), 188
maidella (Dyar), 187, 188, 189
mysiella (Dyar), 187, 188
pallidipennella (Hulst), 187, 188, 191
semicana Heinrich, n. sp., 188
Eurythmasis Dyar, 184, 203, 204, 206
ignifatua Dyar, 203, 204
Eurythmia Ragonot, 186, 281, 306, 308
angulella Ely, 307, 308
diffusella Ely, 306, 307
fumella Ely, 308
hospitella (Zeller), 306, 307
hospitella yavapaella Dyar, 307
spaldingella Dyar, 307
Eurythmidia Ragonot, 184, 204, 205
ignidorsella (Ragonot), 204, 205
euthales (Dyar), Davara, 75
excantalis (Hulst), Sarata, 161, 163, 164,
165
excisalis (Hampson), Diatomocera, 51
exiguella (Ragonot), Nonia, 215, 216
exoleta (Zeller), Hemiptilocera, 30, 32
expunctrix (Dyar & Heinrich), Pseudo-
cabima, 55
exsulella (Zeller), Acrobasis, 19
extracta Heinrich, n. sp., Diatomocera, 52,
53
extrincica (Dyer), Chorrera, 177, 178
Exuperius Heinrich, n. gen., 181, 274
negator Heinrich, n. sp., 274
faecella (Zeller), Laodamia, 145
famula Zeller, ‘“‘“Myelois,” 312
Farnobia Heinrich, n. gen., 184, 276, 277
quadripuncta (Zeller), 207, 276
fasciella Barnes & McDunnough, Pyla,
147, 148
fasciolalis (Hulst), Pyla, 142, 143, 149
fearnella (Schaus), Pseudocabima, 53, 54
feella Dyar, Pyla. 146
feliculella Dyar, Canarsia, 202
feltella Dyar, Acrobasis, 15, 17
fenestrella (Packard), Lipographis, 166,
167, 168
feriella Hulst, Tacoma, 178
fernaldi (Ragonot), Nephopteryx, 125,
126, 127, 129
fernaldialis (Hulst), Cactobrosis, 260, 261
ferrealis (Hampson), Hypsipyla, 29
ferruginella (Ragonot), Eulogia, 275
ficulella Barrett, Ephestia, 304
fieldiella (Dyar), Yosemitia, 250, 251
fifaca (Dyar), Bema, 218, 219
fifria Dyar, Anadelosemia, 68
figulella Curran, Ephestia, 304
figulilella Gregson, Ephestia, 302, 303, 304
filiolella (Hulst), Atheloca, 82
finitella (Walker), Tulsa, 134, 135
fiskella Dyar, Laetilia, 230, 234, 235
fissuralis (Walker), Crocidomera, 33
flavicans (Ragonot, not Zeller), Piesmo-
poda, 79
flavicans (Zeller), Piesmopoda, 78, 79, 80
flavicornella Ragonot, ‘‘Phycitopsis,’’ 314
flavidorsella (Ragonot), Passadena, 175
flavipicta Hampson, Psammia, 315
floridellus (Hulst), Caristanius, 98
floridensis Heinrich, n. race (race of dam-
mersi Heinrich), Nephopteryx, 126
572
floridensis Heinrich, n. sp., Unadilla, 229
fluviatella Schaus, Hypsipyla, 29
formosella (Wileman & South), Ephestia,
303
formulella Schaus, ‘“‘Moodna,” 313
fossulatella Ragonot, Oryctometopia, 158,
159
fosterella Hulst, Pima, 101, 102, 104, 105
fragilella (Dyar), Myelopsis, 41
franconiella Hulst, Zophodia, 239
fraiella Dyar, Piesmopoda, 78, 79
fraterna Heinrich, n. sp., Zamagiria, 92, 93
fraudifera Heinrich n. race (race of delas-
salis Hulst), Nephopteryx, 125
frigidella (Packard), Pyla, 144
fritilla Dyar, Bema, 218
fructetella (Hulst), Salebriaria, 115, 118,
119
frustrator Heinrich, n. sp., Megarthria, 87
fulgidula Heinrich, n. sp., Hyalospila, 57
fuliginosus Heinrich, n. sp., Honorinus, 199
Fulrada Heinrich, n. gen., 9, 71, 73
carpasella (Schaus), 72
querna (Dyar), 71, 72
fulvirugella (McDunnough, not Ragonot),
Pima, 104
fulvirugella (Ragonot), Pima, 104, 105
fumella Ely, Eurythmia, 308
fumosella (Hulst), Hulstia, 196
Fundella Zeller, 3, 59, 62, 312
agapella Schaus, 60, 61
ahemora Dyar, 59, 62
argentina Dyar, 55, 59, 60, 61, 62
cistipennis (Dyar), 59, 60
eucasis Dyar, 61
ignobilis Heinrich, 60, 61
pellucens Zeller, 59, 60, 61
pellucens Zeller (not Zeller), 61
funerellus (Dyar), Phobus, 138, 139
furciferella (Dyar), Olybria, 113 114
furculella (Dyar), Sarasota, 76, 77
furfurella (Hulst), Caristanius, 98
furvidorsella (Ragonot), Ectomyelois, 45,
46
fusca (Haworth), Pyla, 144
fuscatella (Hulst), Canarsia, 202
fuscatella (Hulst), Eumysia, 188
fuscifrontella Zeller, ‘“‘Nephopteryx,” 312
fuscofasciella (Ragonot), Anagasta, 300
fuscomaculella (Wright), Ozamia, 257
Gabinius Heinrich, n. gen., 4, 84
paulsoni (Ragonot), 84, 85
gais Dyar, ‘‘Euzophera,”’ 313
galdinella (Schaus), Tota, 170
gamma, Heinrich, n. sp., Sarata, 164
gemina (Haworth), Homoeosoma, 219
geminipunctella (Ragonot), Telethusia, 137
Gennadius Heinrich, n. gen., 184, 276,
277
junctor Heinrich, n. sp., 277
georgiella (Hulst), Salebriaria, 117
gigantella (Ragonot), Cactobrosis, 260
gillettella (Dyar), Tulsa, 135
gilvescentella Ragonot, Ephestiodes, 278,
279, 280, 281, 282
gilvibasella Hulst, Nephopteryx, 132
gitonella (Druce), Anagasta, 300
glabrella (Dyar), Baphala, 237
glaucatella (Hulst), Rumatha, 248, 249
gleditschiella (Fernald), Tlascala, 134
glendella (Dyar) (race of lunigerella Rag-
onot), Promylea, 66
glomis (Dyar), Laetilia, 230, 234
glycinivora (Matsumura), Plodia, 299
Glyptocera Ragonot, 6, 100
busckella (Dyar), 101
consobrinella (Zeller), 100, 101
gnidiella (Milliére), Cryptoblabes, 10
goyensis (Ragonot), Baphala, 235, 236
eraciella (Hulst), Yosemitia, 250, 251
graciella (Hulst, not Hulst), Yosemitia,
251
gracilella Hulst, Canarsia, 202
grandella (Zeller), Hypsipyla, 28, 29
granitella (Ragonot), Pima, 106
granulella Hampson, Ephestiodes, 281
griselda Dyar, Euzophera, 274
grisella Barnes & McDunnough, Bertelia,
36, 37
grossbecki (Barnes & McDunnough), Ac-
robasis, 13
grossipunctella Ragonot, “‘Myelois,” 312
grossulariae (Riley), Zophodia, 239
grossularialis Hiibner, Zophodia, 239
grossulariella (Hiibner), Zophodia, 239
groteii Ragonot, Ulophora, 176, 177
grotella (Ragonot), Monoptilota, 89
guarinella (Zeller), Ulophora, 177
guatemalellus (Ragonot), Caristanius, 98
guianalis Heinrich, n. sp., Pseudocabima,
54
gulosella (Hulst),
158
gurbyris Dyar, Ilatila, 263
Dioryctria, 149, 157,
hammondi (Riley), Psorosina, | 209
hanhamella Dyar, Pyla, 146
hapsella (Hulst), Adelphia, 169
Harnocha Dyar, 184, 202
velessa Dyar, 202, 203
Harnochina Dyar, 315
rectilinea Dyar, 315
haywardi Heinrich, n. sp., Baphala, 236
hebescella (Dyar, not Hulst), Acrobasis, 17
hebescella Hulst, Acrobasis, 21
heinrichalis (Dyar), Salebriaria, 117
heliophila Dyar, Ozamia, 257
hemilutella Dyar, Ozamia, 257, 259, 260
Hemiptilocera Ragonot, 4, 30, 34, 35, 276
bigrana (Zeller), 30, 31
chinographella Ragonot, 30, 31
exoleta (Zeller), 30, 32
jecarella (Schaus), 30, 32
letharda (Schaus), 30, 31, 32
plumigerella (Ragonot), 30, 31, 32
Heras Heinrich, n. gen., 4, 34
disjunctus Heinrich, n. sp., 34, 35
Heterographis Ragonot, 183, 193, 194, 195
coloradensis Ragonot, 194
ignistrigella Ragonot, 194
morrisonella Ragonot, 193, 194
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Heterographis Ragonot—Continued
olbiella (Hulst), 198, 194
palloricostella (Walter), 194
samaritanella (Zeller), 193
holochlora (Dyar), Alberada, 245
Homalopalpia Dyar, 73, 74
Homoeographa Ragonot, 7, 135
lanceolella Ragonot, 135, 136
Homoeosoma Curtis, 184, 219, 225, 227,
228
acmaeopterum Ragonot, 225
albescentellum Ragonot, 222, 223
assitum Heinrich, n. sp., 225
candidella Hulst, 221
cubella Dyar, 219, 226
deceptorium, Heinrich, n. sp., 223
differtella Barnes & McDunnough,
220
discrebile Heinrich, n. sp., 224
ditaeniatellum Ragonot, 224
electellum (Hulst), 220, 223, 224
elongellum Dyar, 222
gemina (Haworth), 219
illuviellum Ragonot, 221, 222, 223
illuviellum emendator Heinrich, n.
race, 222
imitator Heinrich, n. sp., 222
impressale Hulst, 223
inornatellum (Hulst), 223
longiventrellum Ragonot, 222
musiosum Dyar, 219, 226
nimbosellum Ragonot, 225
noctividella Ragonot, 222
oconequensis (Dyar), 224
olectella (Hulst), 220.
opalescella (Hulst), 220
ogslarellum Dyar, 221
oslarellum breviplicitum Heinrich, n.
race, 221
peregrinum Heinrich, n. sp., 224
sinuella (Fabricius), 219
striatellum Dyar, 221, 222
stypticellum Grote, 220, 223
tenuipunctella Ragonot, 220
tezanella Ragonot, 220
uncanale Hulst, 220
uncanalis Ragonot (not Hulst), 223
unionellum Ragonot, 225
vepallidum Heinrich, n. sp., 224, 227
homoeosomella (Zeller), Baphala, 235, 236,
237
Honora Grote, 85, 180, 196, 199, 210, 313
canicostella Ragonot, 198
dotella Dyar, 197, 198
mellinella Grote, 197, 198
montinatatella (Hulst), 198
ochrimaculella (Ragonot), 197
perdubiella (Dyar), 197, 198
sciurella Ragonot, 198
subsciurella Ragonot, 197, 198
Honorinus Heinrich, n. gen., 181, 199
fuliginosus Heinrich, n. sp., 199
hoplidice (Dyar), Diatomocera, 52
horneana (Dyar), Dioryctria, 154
Hornigia Ragonot, 287
hospitabilis Dyar, Zamagiria, 90, 91
hospitella (Dyar, not Zeller), Erelieva, 308
hospitella (Zeller), Eurythmia, 306, 307
huanucensis Heinrich, Sigelgaita, 256
Hulstia Ragonot, 183, 195
fumosella (Hulst), 196
oblitella (Ragonot, not Hulst), 196
obsipella (Hulst), 169, 196
rubiginalis (Walker), 196
undulatella (Clemens), 169, 195, 196
hulstiella Ragonot, ‘“Hypochaleia,’’ 312
hulstii Cockerell, Laetilia, 230, 232
humilis Ragonot, Lipographis, 166, 167
Hyalospila Ragonot, 9, 56, 84
angulinella (Schaus), 58
celiella Schaus, 57
clevelandella (Dyar), 58
egenella (Ragonot), 58
fulgidula Heinrich, n. sp., 57
insequens Heinrich, n. sp., 57
majorina Heinrich, n. sp., 57
semibrunneella Ragonot, 59
stictoneurella Ragonot, 35, 56, 57, 58
xanthoudemia (Dyar), 58
Hypargyria Ragonot, 2, 11, 37
definitella (Zeller), 37, 38
metalliferella Ragonot, 37, 38
slossenella (Hulst), 38
tenuella (Barnes & McDunnough), 38
Hypermescinia Dyar, 215
Hyphantidium Scott, 301
Hypochaleia Hiibner, 312
ahenella (Zeller), 312
hypoehalciella (Ragonot), Pyla, 145, 146
Hypsipyla Ragonot, 2, 27
cnabella Dyar, 28
dorsimacula (Schaus), 28, 29
ferrealis (Hampson), 29
fluviatella Schaus, 29
grandella (Zeller), 28, 29
pagodella Ragonot, 27, 28
robusta (Moore), 27, 28
hystriculella (Hulst), Chararica, 39
idiotes Dyar, Chorrera, 177, 178 chill
ignidorsella (Ragonot), Eurythmidia, 204,
205
ignifatua Dyar, Eurythmasis, 203, 204
ignistrigella Ragonot, Heterographis, 194
ignobilis (Butler), Ancylostomia, 95
ignobilis Heinrich, Fundella, 60, 61
ithouna Dyar, Zophodia, 239
ilignella (Zeller), Divona, 166
Illatila Dyar, 180, 263
gurbyris Dyar, 263
illuviellum Ragonot, Homoeosoma, 221,
222, 223
imitator Heinrich, n. sp., Homoeosoma, 222
immorella (Dyar), Ozamia, 258
immundella (Hulst), Myelopsis, 40
Immyrla Dyar, 3, 111
nigrovittella Dyar, 111, 112
impeditella (Zeller), Cabotia, 201
imperfecta (Dyar), Aptunga, 211
impletella (Zeller), Prosoeuzophera, 275,
276
INDEX
impostor Heinrich, n. sp., Pyla, 143, 144,
145, 148
impressale Hulst, Homoeosoma, 223
inanimella (Dyar), Vitula, 287
incanella (Hulst), Sarata, 160, 161
incautella (Zeller), Elasmopalpus, 173
incertus Heinrich, n. sp., Phobus, 139
inconditella (Ragonot), Nephopteryx, 125,
127, 128
incorruscella (Hulst), Catastia, 110, 111
indecora Dyar, Mescinia, 212, 213, 215
indentella Dyar, Ephestiodes, 281
indianella Dyar, ‘‘Megasis,’”’ 160, 315
indigenella (Zeller), Acrobasis, 12, 14
infimella Ragonot, Ephestiodes, 279, 280,
281
infinitella (Dyar), Tulsa, 135
infusella Zeller, ‘‘Myelois,” 314
innoxia Heinrich, Ribua, 297, 298
inornatella (Ragonot), Moodnopsis, 270,
271
inornatellum (Hulst), Homoeosoma, 223
inquilinella Ragonot, Nephopteryx, 128,
133
insequens Heinrich, n. sp., Hyalospila, 57
insignatella Dyar, Cactobrosis, 260, 261
insignis Heinrich, Eremberga, 254
insinuatrix Heinrich, n. sp., Pyla, 144
interjecta Heinrich, n. sp., Davara, 73,
74, 75
Interjectio Heinrich, n. gen., 3, 106, 108
columbiella (McDunnough), 106, 107
denticulella (Ragonot), 106, 107, 110,
138
lallatalis authors (not Hulst), 106, 107
niviella (Hulst), 107
ruderella (Ragonot), 107
interpunctalis (Hiibner), Plodia, 298
interpunctella (Hiibner), Plodia, 298, 299,
303
interstitialis (Dyar), Cahela, 248
intextella (Zeller), ‘“Euzophera,” 314
intransitella (Dyar), Adanarsa, 35
inveterella (Dyar), Moodnopsis, 268, 271
iota Heinrich, n. sp., Sarata, 164
ipsetona Dyar, Zamagiria, 93
irichampa Dyar, ‘‘Anthropteryx,” 313
irrubriella Ely, Acrobasis, 20
isabella (Dyar), Piesmopoda, 78, 79, 80, 81
jocarella (Schaus), Hemiptilocera, 30, 32
juglandis (LeBaron), Acrobasis, 15, 16, 17
jugosella Ragonot, Ortholepsis, 119, 120
junctolineella (Hulst), Olycella, 241, 242,
243
junctor Heinrich, n. sp., Gennadius, 277
kappa Heinrich, n. sp., Sarata, 163, 165
kearfottella Dyar, Acrobasis, 17
kiihniella (Zeller), Anagasta, 291, 299, 300,
302
lacteella (Hulst), Nephopteryx, 132
laetella Grote, Ambesa, 108
Laetilia Ragonot, 183, 230, 231, 235, 240,
255
573
Laetilia Ragonot—Continued
amphimetra (Mayrick), 230, 233
coccidivora (Comstock), 230, 231, 232,
254
coccidivora cardini Dyar, 230, 232
coccidivora quadricolorella (Dyar),
231, 232
dilatifasciella (Ragonot), 230
ephestiella (Ragonot), 230, 234
fiskella Dyar, 230, 234, 235
glomis (Dyar), 230, 234
hulstii Cockerell, 230, 232
lustrella (Dyar), 234
melanostathma (Meyrick), 230, 232,
233, 234
myersella Dyar, 230, 233, 234
obscura Dyar, 232
pallida (Comstock), 230
portoricensis Dyar, 232
zamacrella Dyar, 230, 233, 234, 286
laidion (Zeller), Zamagiria, 92
lallatalis authors (not Hulst), Interjectio,
106, 107
lallatalis (Dyar, not Hulst), Phobus, 138
lallatalis (Hulst), Ambesa, 109
lallatalis (Hulst, not Hulst), Phobus, 138
lambella (Dyar), Nonia, 215, 216
lanceolella Ragonot, Homoeographa, 135,
136
Laodamia Ragonot, 145
faecella (Zeller), 145
Laosticha Hulst, 230
Lascelina Heinrich, n. gen., 182, 264
canens Heinrich, n. sp., 264
latercula (Hampson), Plodia, 299
latifasciella Dyar, Acrobasis, 20
latifasciatella (Packard), Telethusia, 137
lativittella (Ragonot), Valdivia, 191, 192
laura (Dyar), Vitula, 285, 287
laurata (Heinrich), Dioryctria, 153
legatella (Hiibner), Rhodophaea, 24
leithella (Dyar), Amalafrida, 256
leoninella (Packard), Lipographis, 166,
167, 168
letharda (Schaus), Hemiptilocera, 30, 31,
32
leuconips (Dyar), Eremberga, 248, 253,
262
leucophaeella (Hulst), Oreana, 112, 117
levigatella (Hulst), Nephopteryx, 129
lignosellus (Zeller), Elasmopalpus,
173, 204
Lipographis Ragonot, 3, 160, 163, 166
fenestrella (Packard), 166, 167, 168
humilis Ragonot, 166, 167
leoninella (Packard), 166, 167, 168
pallidella (Dyar), 167
subosseella Hulst, 168
truncatella (Wright), 166, 167
umbrella (Dyar), 160, 166, 168
liquidambarella (Dyar), Nephopteryx, 131
lithosella (Ragonot), Ufa, 170, 171
longipennella (Hampson), Cactobrosis, 260,
261
longipennella
251
172,
(Hulst), Yosemitia, 250,
574
longiventrellum Ragonot,
222
lophophora (Dyar), Azaera, 283
lophotalis Heinrich, n. sp., Rampylla,
70, 71
lucidalis (Walker), Ozamia, 257, 258
lucidibasella Ragonot, Ephestiodes, 278,
280
lugubrella (Ragonot), Vitula, 286
lunigerella Hampson, Stylopalpia, 140, 141
lunigerella Ragonot, Promylea, 65, 66
lustrella (Dyar), Laetilia, 234
luteella (Hulst), Ufa, 170
Homoeosoma,
mabes Dyar, ‘“‘Euzophera,” 315
macropasa (Dyar), Aptunga, 211
Macrorrhinia Ragonot, 182, 189, 190, 191,
192, 194
aureofasciella Ragonot, 190
placidella (Zeller), 191
maculicula (Dyar), Rhagea, 238
maculifera Dyar, Cactobrosis, 260, 261
magellanella (Ragonot), Patagonia, 225
Magiriopsis Heinrich, n. gen., 9, 94
cristalis (Hampson), 94
denticosella (Dyar), 94
magnificans Dyar, Zophodia, 239
maidella (Dyar), Eumysia, 187, 188, 189
major (Zeller), Elasmopalpus, 173
majorella Dyar, Dioryctria, 152
majorella (Dyar), Sosipatra, 296
majorina Heinrich, n. sp., Hyalospila, 57
majuscula Heinrich, n. sp., Diatomocera, 52
malacella Dyar, Cassiana, 212
malipennella Dyar, Acrobasis, 20
mamella (Dyar), Ceracanthia, 86
Manhatta Hulst, 185, 283, 284, 287
biviella (Zeller), 272, 284, 287
broweri Heinrich, n. sp., 288
setonella (McDunnough), 287
marginea (Schiffermiiller), Catastia, 110
Maricopa Hulst, 191
marmorea (Haworth), Rhodophaea, 24
masculinus Dyar, Zamagiria, 91
maturella (Zeller), Unadilla, 218, 228, 229
medulallis Hubner, Myelois, 40, 43
megalopalis Hampson, ‘‘Euzopherodes,”’
314
Megaphycis Grote, 240
Megarthria Ragonot, 3, 5, 86
alpha Heinrich, n. sp., 88
beta Heinrich, n. sp., 88
cervicalis Dyar, 86, 87
frustrator Heinrich, n. sp., 87
peterseni (Zeller), 86, 87, 88
schausi Heinrich, n. sp., 87
squamifera Heinrich, n. sp., 87
Megasis Guenée, 160
rippertella (Zeller), 160
melanellus (Hulst), Tulsa, 134
melanoplaga (Hampson), Caristanius, 97
melanostathma (Meyrick), Laetilia, 230,
232, 233, 234
Melia Heinemann, 272
UNITED
STATES NATIONAL MUSEUM BULLETIN 207
Melitara Walker,
244, 252
bollit Dodd (not Zeller), 241
bolliz (Zeller), 240
dentata (Grote), 241
doddalis Dyar, 241
prodenialis Walker, 240, 241
mellinella Grote, Honora, 197, 198
mendacella Staudinger, Dioryctria, 149
Meroptera Grote, 6, 13, 101, 112, 121,
128, 124, 127, 138, 134
abditiva Heinrich, n. sp., 123
eviatella Dyar, 122
mirandella Ragonot, 121, 122
pravella (Grote), 121, 122, 123, 124,
131
Mescinia Ragonot, 184, 211, 212, 215, 216,
217, 227, 293
bacerella Dyar, 213, 214
berosa Dyar, 214
commatella (Zeller), 212, 214
discella Hampson, 213, 215
estrella Barnes & McDunnough, 213
indecora Dyar, 212, 213, 215
moorei Heinrich, n. sp., 214
mosces Dyar, 213
pandessa Dyar, 213
parvula (Zeller), 214
peruella Schaus, 214
triloses Dyar, 213
metagrammalis (Walker), Caristanius, 98
metalicella Hulst, Pyla, 148
metalliferella Ragonot, Hypargyria, 37, 38
Metephestia Ragonot, 185, 264
simplicula (Zeller), 264, 265
micaceella (Hampson), Sosipatra, 295, 296
Micromescinia Dyar, 184, 277
pygmaea Dyar, 277
Microphestia Dyar, 186, 294
animalcula Dyar, 294
Microphycita Dyar, 310, 311
titillella Dyar, 310, 311
mignonella Dyar, Ephestiodes, 280
Mildrixia Dyar, 2, 26, 37
constitutionella Dyar, 26
Mineola Hulst, 10, 11
milleri Zeller, Ephestia, 304
mindosis Dyar, Promylea, 67
miniatella Ragonot, Dioryctria, 153
minimella Ragonot, Acrobasis, 11, 15
minualis (Walker), Ufa, 171
minutularia (Hulst), Myelopsis, 41
minutulella (Hulst), Myelopsis, 41
mirabella Dyar (race of walsinghami
(Ragonot)), Ambesa, 109
mirabilicornella (Dyar), Protasia, 193
mirandella Ragonot, Meroptera, 121, 122
mochlophleps (Dyar), Diatomocera, 52
modestella (Hulst), Telethusia, 137
Moerbes Dyar, 50, 181, 267, 268
alveolella (Ragonot), 269
dryopella Dyar (not Schaus), 269
dryopella (Schaus), 268, 269
emendata Heinrich, n. sp., 50, 268, 269
moeschlert (Ragonot), Oryctometopia, 159
moestella (Walker), Pyla, 144
181, 240, 241, 243,
Mona Hulst, 193
monodon Dyar, Ambesa, 108
Monoptilota Hulst, 5, 89
grotella (Ragonot), 89
nubilella Hulst, 69, 117
pergratialis (Hulst), 89
montella Schaus, Piesmopoda, 78, 81
montinatatella (Hulst), Honora, 198
Moodna Hulst, 185, 283, 285, 287, 288, 289,
290, 292, 293, 313, 314
bisinuella Hampson, 284
obtusangulella (Ragonot), 284
ostrinella (Clemens), 209, 284, 293
pelviculella Hulst, 283, 284
Moodnella Heinrich, n. gen., 184, 289
paula Heinrich, n. sp., 289
Moodnopsis Dyar, 181, 268, 269, 313, 314
decipiens Dyar, 269, 270, 271
inornatella (Ragonot), 270, 271
inveterella (Dyar), 268, 271
parallela Heinrich, n. sp., 271
perangusta (Dyar), 270
portoricensis Heinrich, n. sp., 230, 271
moorei Heinrich, n. sp., Mescinia, 214
morbosella (Staudinger), Staudingeria, 194
morrisonella Ragonot, Heterographis, 193,
194
mosces Dyar, Mescinia, 213
mossa (Dyar), Erelieva, 308
mucidella (Ragonot), Rotruda, 219, 225,
226, 227
muciella Schaus, Azaera, 282, 283
muellerana Dyar, Dioryctria, 152
mundelli Heinrich, Cactoblastis, 247
muriscis (Dyar), Ectomyelois, 45, 46, 47, 48
musiosum Dyar, Homoeosoma, 219, 226
Myelois Hiibner, 40, 46, 47, 312
cribrella Hiibner, 40, 43
medulallis Hiibner, 40, 43
Myelopsis Heinrich, n. gen., 8, 40
alatella (Hulst), 41
coniella (Ragonot), 40, 41, 265
fragilella (Dyar), 41
immundella (Hulst), 40
minutularia (Hulst), 41
minutulella (Hulst), 41
nefas (Dyar), 40
obnupsella (Hulst), 41
piazzella (Dyar), 41
rectistrigella (Ragonot), 41
subtetricella (Ragonot), 40, 41
tetricella (Schiffermiiller), 40
zonulella (Ragonot), 41
myersella Dyar, Laetilia, 230, 233, 234
myja Dyar, Bema, 217, 218, 219
myricella Barnes & McDunnough, Acro-
basis, 23
Myrlaea Ragonot, 124, 130
albistrigella Staudinger, 124
myronella Dyar, Cabnia, 310
mysiella (Dyar), Eumysia, 187, 188
Nanaia Heinrich, 183, 245
substituta Heinrich, 245
nannodes Dyar, Varneria, 305
nasutella Hulst, Unadilla, 227, 228, 229
Nasutes Hampson, 315
venata Hampson, 315, 316
neaeriatella Grossbeck, Wunderia, 204, 205
nebulella Dyar (not Riley), Acrobasis, 16
nebulella (Grossbeck, not Riley), Acrobasis,
13
nebulella (McDunnough, not Riley), Ore-
ana, 112
nebulella (Riley), Acrobasis, 12, 13
nebulo (Walsh), Acrobasis, 12
nefas (Dyar), Myelopsis, 40
negator Heinrich, Exuperius, n. sp., 274
Neopyralis Bréthes, 245
nephelepasa (Dyar), Olycella, 242, 243
nephelepasa (Dyar, in part), Olycella, 243
Nephopteryx Hiibner, 6, 101, 121, 123, 133,
134, 136, 137, 188, 312
afflictella (Hulst), 131
basilaris Zeller, 129
bifasciella Hulst, 130
bisra Dyar, 133
carneella Hulst, 128
celtidella (Hulst), 131, 132
contatella (Grote), 124, 127, 128
crassifasciella Ragonot, 5, 132
crataegella Barnes & MecDunnough, 132
croceella (Hulst), 123, 131
dammersi Heinrich, n. sp., 126, 127
dammersi floridensis Heinrich, n. race,
126
decipientella Dyar, 132
delassalis Hulst, 125, 127
delassalis (Hulst, not Hulst), 125, 126
delassalis fraudifera Heinrich, n. race,
125
fernaldi (Ragonot), 125, 126, 127, 129
gilvibasella Hulst, 132
inconditella (Ragonot), 125, 127, 128
inquilinella Ragonot, 128, 133
lacteella (Hulst), 132
levigatella (Hulst), 129
liquidambarella (Dyar), 131
nogalesella (Dyar), 130
pravella authors (not Grote), 124
pudibundella (Ragonot), 125
purpurella (Hulst), 124
quinquepunctella (Grote), 128
rhenella (Zincken), 123, 134
rubescentella (Hulst), 125
rubrisparsella (Ragonot), 131, 132
rufibasella (Ragonot), 131
semiobscurella (Hulst), 123, 124
subcaesiella (Clemens), 6, 127, 128, 129
subfuscella (Ragonot), 123, 124
termitalis (Hulst), 129, 130
termitalis yuconella (Dyar), 130
teranella (Hulst), 132
uvinella (Ragonot), 123, 130
vetustella (Dyar), 127
virgatella (Clemens), 128
nerthella (Schaus), Davara, 75
neuricella (Zeller), Bema, 218
nexa Heinrich, n. sp., Rioja, 267, 268
Nicetiodes Schaus, 185, 304
apianella Schaus, 304, 314
nigrella (Hulst), Caudellia, 283, 293
obtusangulella (Ragonot),
INDEX
nigrella Hulst, Ephestiodes, 278, 283
nigricantella Ragonot, Euzophera, 55, 274
nigricans Hulst, ‘“‘Salebria,’’? 312
nigricula Heinrich, n. sp., Pyla, 148
nigrifasciella Ragonot, Sarata, 162, 163, 165
nigristrigella (Ragonot), Pseudocabima,
54, 55
nigritella (Hampson), Oncolabis, 199
nigrosignella Hulst, Acrobasis, 15
nigrovittella Dyar, Immyrla, 111, 112
nimbella (Zeller), Rotruda, 219, 226
nimbosellum Ragonot, Homoeosoma, 225
niviella (Hulst), Interjectio, 107
noctividella Ragonot, Homoeosoma, 222
nodoses (Dyar), Azaera, 283
nogalesella (Dyar), Nephopteryx, 130
Nonia Ragonot, 180, 184, 213, 215, 216
exiguella (Ragonot), 215, 216
lambella (Dyar), 215, 216
noniella Dyar, Ephestiodes,
306
nonparilella (Dyar), Sosipatra, 296
normella Dyar, Acrobasis, 20
notatalis (Walker), Paramyelois, 47
nubiferella (Ragonot, Salebriaria,
117
nubilella Hulst, Monoptilota, 89, 117
nymphaeella (Hulst), Palatka, 207, 208
nyssaecolella (Dyar), Actrix, 139, 140
278, 282,
116,
oberthuriella (Ragonot), Olybria, 113
oblitella (Ragonot, not Hulst), Hulstia, 196
obnupsella (Hulst), Myelopsis, 41
obscura Dyar, Laetilia, 232
obsipella (Hulst), Hulstia, 169, 196
obstitella (Schaus), Anadelosemia, 68
Moodna,
Ocala Hulst, 189, 191, 192, 194
dryadella Hulst, 191
platanella (Grossbeck), 191
occidentalis Heinrich, n. race (race of
284
albiplagiatella (Packard)), Pima,
103
ochrella Barnes & McDunnough, Divitiaca,
189, 190
ochrifrontella (Zeller), Eulogia, 275
ochrimaculella (Ragonot), Honora, 197
ochripunctella (Dyar), Adelphia, 169
ochrodesma (Zeller), Anabasis, 25, 26
oconequensis (Dyar), Homoeosoma, 224
odiosella Weinrich (not Hulst), Ozamia,
257, 258
odiosellus (Hulst), Salebriacus, 114
Oedothmia Hampson, 185, 205
bahamasella (Hampson), 205
endopyrella Hampson, 205
olbiella (Hulst), Heterographis, 193, 194
olectella (Hulst), Homoeosoma, 220
olivacea Heinrich, n. race (race of goyensis
(Ragonot)), Baphala, 236
olivaceella (Ragonot), race of mucidella
(Ragonot)), Rotruda, 226
olivacella Dyar, Staudingeria, 194, 195
olivella Hampson, ‘‘Moodna,” 284, 314
olivella (Hulst), Ragonotia, 160, 315
575
Olybria, Heinrich, 6, 113, 114
aliculella (Hulst), 113, 114
furciferella (Dyar), 113, 114
oberthuriella (Ragonot), 113
Olyca Walker, 181, 243, 253
phryganoides Walker, 243
Olycella Dyar, 240, 241, 243, 256
junctolineella (Hulst), 241, 242, 243
junctolineella pectinatella (Hampson),
242
nephelepasa (Dyar), 242, 243
nephelepasa (Dyar, in part), 243
subumbrella (Dyar), 242, 243
Oncolabis Zeller, 181, 199, 203, 206
anticella Zeller, 199, 206
nigritella (Hampson), 199
opalescella (Hulst), Homoeosoma, 220
oporedestella (Dyar), Ectomyelois, 44
Orena Hulst, 6, 111, 112
leucophaeella (Hulst), 112, 117
nebulella (McDunnough, not Riley),
112
unicolorella (Hulst), 13, 112
oregonella (Barnes & MeDunnough),
Tulsa, 135
orio Dyar, Rampylla, 70
orobanchella (Dyar), Rhagea, 237
Ortholepis Ragonot, 3, 112, 119, 121, 124
jugosella Ragonot, 119, 120
pasadamia (Dyar), 120
Oryctometopia Ragonot, 1, 9, 158
fossulatella Ragonot, 158, 159
moeschleri (Ragonot), 159
oslarellum Dyar, Homoeosoma, 221
ostricolorella Hulst, Euzophera, 273,274
ostrinella (Clemens), Moodna, 209, 284,
293
ostryella Ely, Acrobasis, 21
ovalis (Packard), Telethusia,
138
Ragonot, 183, 257
fuscomaculella (Wright), 257
fuscomaculella clarefacta Dyar, 258
heliophila Dyar, 257
hemilutella Dyar, 257, 259, 260
immorella (Dyar), 258
lucidalis (Walker), 257, 258
odiosella Heinrich (not Hulst), 257, 258
punicans Heinrich, 257, 259
stigmaferella Dyar, 258
thalassophila Dyar, 258
136, 137,
pachytaeniella (Ragonot), Volatica, 290,
291
packardella (Ragonot), Rhagea, 237, 238
Paconius Heinrich, n. gen., 181, 210
corniculatus Heinrich, n. sp., 210
pagodella Ragonot, Hypsipyla, 27, 28
Palatka Hulst, 185, 207
nymphaeella (Hulst), 207, 208
verecuntella (Grossbeck), 208
pallicornella (Ragonot), Trachycera, 25
pallida (Comstock), Laetilia, 230
pallidella (Dyar), Lipographis, 167
pallidipennella (Hulst), Eumysia, 187, 188,
191
576
palliolella Dyar (not Ragonot), Acrobasis,
16
palliolella Ragonot, Acrobasis, 15, 16
palloricostella (Walter), Heterographis, 194
pallulella (Hulst), Euzophora, 273
palpalis (Dyar), Ectomyelois, 45
pandessa, Dyar, Mescinia, 213
parabates (Dyar), Alberada, 244, 245
parallela Heinrich, n. sp., Moodnopsis, 271
Paramyelois Heinrich, n. gen., 8, 46
cassiae (Dyar), 47
duplipunctella (Ragonot), 47
notatalis (Walker), 47
soliiella (Zeller), 46, 47
transitella (Walker), 47
ventpars (Dyar), 47
paranensis (Dyar), Davara, 75
parasitaria Heinrich, n. sp., Vezina, 291
parkerella (Schaus), Pima, 102, 106
Parolyca Dyar, 183, 255
asthenosoma (Dyar), 255
parva Heinrich, n. sp., Piesmopoda, 80
parvula (Zeller), Mescinia, 214
parvulella Barnes & MceDunnough, Diviti-
aca, 198
parvulella (Ely), Erelieva, 307, 308, 309
pasadamia (Dyar), Ortholepis, 120
Passadena Hulst, 2, 175, 187
canescentella (Hulst), 175
cinctella (Hulst), 175
constantella Hulst, 175
flavidorsella (Ragonot), 175
passulella Barrett, Ephestia, 303
Patagonia Ragonot, 184, 219, 225
magellanella (Ragonot), 225
patriciella (Dyar), Ribua, 298
Patriciola Heinrich, n. gen., 181, 209, 210
semicana Heinrich, n. sp., 209, 210
paula Heinrich, n. sp., Moodnella, 289
paulsoni (Ragonot), Gabinius, 84, 85
paurolepidalis Hampson, Eutrichocera, 315
Peadus Heinrich, n. gen., 8, 83, 85
burdetellus (Schaus), 83, 84
dissitus Heinrich, n. sp., 84
semproniella (Schaus), 84
subaquilellus (Ragonot), 84
pectinatella (Hampson) (race of junctolin-
eella (Hulst)), Olycella, 242
pellucens Zeller, Fundella, 59, 60, 61
pellucens Zeller (not Zeller), Fundella, 61
pellucidellus (Ragonot), Caristanius, 97, 98
pelviculella Hulst, Moodna, 283, 284
peplifera Dyar, Acrobasis, 18, 19
perangusta (Dyar), Moodnopsis, 270
perdubiella (Dyar), Honora, 197, 198
peregrinum Heinrich, n. sp., Homoeosoma,
224
perfuscalis Hulst, Sarata, 162, 164, 165
pergratialis (Hulst), Monoptilota, 89
perluteella Dyar, Staudingeria, 195
perrensiella (Ragonot), Pseudocabima, 55
peruella Schaus, Mescinia, 214
peterseni (Zeller), Megarthria, 86, 87, 88
petrella (Zeller), Adelphia, 168, 169, 170
Phestinia Hampson, 184, 216
costella Hampson, 216
phi Heinrich, n. sp., Sarata, 163, 165
Philodema, Heinrich, n. gen., 3, 165, 166
rhoidella (Dyar), 160, 165, 166
Phobus, Heinrich, n. gen., 5, 138
brucei (Hulst), 110, 138, 139
curvatellus (Ragonot), 137, 139
funerellus (Dyar), 138, 139
incertus Heinrich, n. sp., 139
lallatalis (Dyar, not Hulst), 138
lallatalis (Hulst, not Hulst), 138
rhypodella (Ragonot, not Hulst), 139
phoenicis (Durrant), Ectomyelois, 45
phoenicis (Dyar), Cahela, 248
phryganoides Walker, Olyca, 243
phycidea Zeller, 219
Phycitidae, 1, 150
Phycitinae, 1
Phycitopsis Ragonot, 314
piazzella (Dyar), Myelopsis, 41
Piesmopoda Zeller, 9, 56, 74, 76, 77
apocerastes Dyar, 78, 80, 81
flavicans (Ragonot, not Zeller), 79
flavicans (Zeller), 78, 79, 80
fratella Dyar, 78, 79
isabella (Dyar), 78, 79, 80, 81
montella Schaus, 78, 81
parva Heinrich, n. sp., 80
ragonoti (Dyar), 78, 79
rubicundella Zeller, 77, 78, 79
semirufella (Zeller), 80, 81
trichomata (Zeller), 78
xanthomera Dyar, 78
xanthopolys Dyar, 79, 80
zanthozona Dyar, 78
Pima, Hulst, 3, 101, 106, 108
albiplagiaiella Hulst (not Packard), 102,
104
albiplagiatella (Packard), 103, 104
albiplagiatella (Ragonot, not Packard),
104
albiplagiatella occidentalis Heinrich,
n. race, 103
albocostalialis (Hulst), 103, 104
albocostalialis subcostella (Ragonot),
105
albocostalis (Hulst), 104
boisduvaliella (Guénée), 102, 103, 105
fosterella Hulst, 101, 102, 104, 105
fulvirugella (McDunnough, not Rag-
onot), 104
fulvirugella (Ragonot), 104, 105
granitella (Ragonot), 106
parkerella (Schaus), 102, 106
piperella (Dyar), 106
vividella (McDunnough), 104
pineae Staudinger, Dioryctria, 149
pinei Heinrich, n. sp., Vitula, 286, 288
pinguis (Haworth), Euzophera, 272, 273
Pinipestis Grote, 150, 154
piperella (Dyar), Pima, 106
placidella (Dyar), Anderida, 211
placidella (Zeller), Macrorrhinia, 191
platanella (Grossbeck), Ocala, 191
Plodia Guénée, 186, 292, 297, 298
dolorosa Dyar, 298, 299
glycinivora (Matsumura), 299
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Plodia Guénée—Continued
interpunctalis (Hiibner), 298
interpunctella (Hiibner), 298, 299, 303
latercula (Hampson), 299
zeae (Fitch), 299
plorella Dyar, Eiphestiodes, 278, 281, 282,
306, 309
plumigerella (Ragonot), Hemiptilocera, 30,
31, 32
plumigerella Hulst, Sarasota, 76, 77
pneumatella (Hulst), Canarsia, 202
pogerythrus Dyar, Zamagiria, 91
polingella (Dyar), Rumatha, 249, 250
Polopeustis, Ragonot, 2, 119, 120
annulatella (Zetterstedt), 120, 121
aretiella (Gibson), 121
polydectella (Schaus), Rampylla, 70, 71
polyphemella (Ragonot), Sarata, 160
pombra (Dyar), Pseudocabima, 54
ponderosae Dyar, Dioryctria, 152
ponderosae Heinrich (not Dyar), Dioryctria,
155, 156
ponderosella (Barnes & McDunnough),
Cahela, 247, 248, 249
porrecta Dyar, Tucumania, 252
portoricensis Dyar, Laetilia, 232
portoricensis Heinrich, n. sp., Moodnopsis,
230, 271
postflavida Dyar, ‘“Huzophera,”’ 312
postica (Zeller), Chorrera, 178
postremella Dyar, Varneria, 305
Praedonula Heinrich, n. gen., 9, 82, 84
almonella (Dyar), 82, 83
pravella (authors, not Grote), Neph-
opteryx, 124
pravella (Grote), Meroptera, 121, 122, 123,
124, 131
Procandiopa Dyar, 85
prodenialis Walker, Melitara, 240, 241
prodromella Hiibner, Hpischnia, 102
productella Ragonot, Ephestiodes,
Promylea Ragonot, 4, 65, 85
dasystigma Dyar, 67
drucei (Dyar), 66
drucei (Ragonot), 66
dyari Heinrich, n. name, 66, 67
lunigerella Ragonot, 65, 66
lunigerella glendella (Dyar), 66
mindosis Dyar, 67
zimmermannt (Druce, not Grote), 66
281
proselytes Dyar, Entmemacornis, 266,
267
Prosoeuzophera Heinrich, n. gen., 185,
275
impletella (Zeller), 275, 276
Protasia Heinrich, n. gen, 182, 193
mirabilicornella (Dyar), 193
Protomoerbes, Heinrich, n. gen., 8, 49
aberrans Heinrich, n. sp., 49
separabilis Heinrich, n. sp., 50
pryeri Ragonot, Dioryctria, 153
Psammia Hampson, 315
flavipicta Hampson, 315
Pseudocabima Heinrich, n. gen., 8, 50, 53
arizonensis Henrich, n. sp., 55
castronalis Heinrich, n. sp., 53, 54
Pseudocabima Heinrich—Continued
euzopherella (Dyar), 53, 54
expunctrix (Dyar & Heinrich), 55
fearnella (Schaus), 53, 54
guianalis Heinrich, n. sp., 54
nigristrigella (Ragonot), 54, 55
perrensiella (Ragonot), 55
pombra (Dyar), 54
rubrizonalis (Hampson), 52, 53, 55
Pseudodivona, Dyar, 8, 48, 49, 268
carabayella Dyar, 48, 49
cispha Dyar, 48, 49
commensella Dyar, 48, 49, 50
santa-maria Dyar, 48, 49
Psorosa Zeller, 210
Psorosina Dyar, 180, 208
angulella Dyar, 208, 209
hammondi (Riley), 209
ptychis (Bondar, not Dyar), Atheloca, 82
ptychis (Dyar), Atheloca, 82
ptyonopoda (Hampson), Sarasota, 77
pudibundella (Ragonot), Nephopteryx, 125
puer Dyar, Elasmopalpus, 173
pulla Heinrich, n. sp., Entmemacornis,
266
pullatella (Raganot),
165
pumilella (Ragonot), Salebriaria, 115, 117
punctella (Dyar), Sarata, 161
punicans Heinrich, Ozamia, 257, 259
purgatoria (Dyar), Cahela, 248
purpurella (Hulst), Nephopteryx, 124
putidella Schaus, ‘“Eucampyla,” 313
pygmaea Dyar, Micromescinia, 277
pygmaeella Ragonot, Dioryctria, 154
Pyla, Grote, 5, 6, 140, 142, 145, 169
aeneella Hulst, 147, 148
aeneoviridella Ragonot, 147, 148
aenigmatica Heinrich, n. sp., 144, 145
aequivoca Heinrich, n. sp., 143, 145
blackmorella Dyar, 145
cacabella (Hulst), 145
criddlella Dyar, 144, 145
fasciella Barnes & McDunnough, 147,
148
fasciolalis (Hulst), 142, 143, 149
feella Dyar, 146
frigidella (Packard), 144.
fusca (Haworth), 144
hanhamella Dyar, 146
hypochalciella (Ragonot), 145, 146
impostor Heinrich, n. sp., 143, 144,
145, 148
insinuatrix Heinrich, n. sp., 144
metalicella Hulst, 148
moestella (Walker), 144
nigricula Heinrich, n. sp., 148
rainierella Dyar, 147, 148
scintillans (Grote), 142, 146, 147, 148
sylphiella Dyar, 147
triplagiatella (Dyar), 145
viridisuffusella Barnes & McDun-
nough, 142, 149
pyllis (Dyar), Unadilla, 228
pytrhochrellus (Ragonot), Ufa, 171
Sarata, 160, 164,
INDEX
quadricolorella (Dyar) (race of coccidivora
(Comstock)), Laetilia, 231, 232
quadripuncta (Zeller), Farnobia, 207, 276
quantulella (Hulst), Erelieva, 307, 308,
309
Quasisalebria Heinrich, n. gen., 7, 111,
112, 118
admixta Heinrich, n. sp., 118, 119
quercicolella (Ragonot), Salebriaria, 117
querna (Dyar), Fulrada, 71, 72
quinquepunctella (Grote), Nephopteryx,
128
Rabiria Heinrich, n. gen., 310, 311
conops (Dyar), 311
ragonoti (Dyar), Piesmopoda, 78, 79
Ragonotia olivella (Hulst), 160, 315
rainierella Dyar, Pyla, 147, 148
Ramphodes Guénée, 99
Rampylla Dyar, 4, 9, 62, 63, 70, 72
lophotalis Heinrich, n. sp., 70, 71
orio Dyar, 70
polydectella (Schaus), 70, 71
subcaudata (Dyar), 71
rectilinea Dyar, Harnochina, 315
rectistrigella (Dyar), Salebriaria, 118
rectistrigella (Ragonot), Myelopsis, 41
reductella (Walker), Tlascala, 133, 134
reliquella (Dyar) (race of mucidella (Rag-
onot)), Rotruda, 219, 226
Relmis Dyar, 218
reniculella (Grote), Dioryetria, 151, 152
reniculella (Packard, not Grote), Dioryctria
150
restrictella Zeller, ‘‘Myelois,”’ 312
Rhagea Heinrich, n. gen., 1, 183, 230, 237
maculicula (Dyar), 238
orobanchella (Dyar), 237
packardella (Ragonot), 237, 238
stigmella (Dyar), 237, 238
rhenella (Zincken), Nephopteryx, 123, 134
Rhodophaea Guénée, 2, 24, 25
advenella (Zincken), 24, 25
caliginella (Hulst), 15, 24, 25
caliginoidella (Dyar), 24
legatella (Hiibner), 24
marmorea (Haworth), 24
suavella (Zincken), 24
supposita (Heinrich), 15, 24
rhoiella (Dyar), Philodema, 160, 165, 166
rhypodella (Hulst), Telethusia, 137
rhypodella (Ragonot, not Hulst), Phobus,
139
rhythmatica Dyar, Cabotia, 200, 201
Ribua Heinrich, n. gen., 186, 297, 298
contigua Heinrich, n. sp., 297
innoxia Heinrich, 297, 298
patriciella (Dyar), 298
rileyella (Ragonot), Sosipatra, 294, 295, 297
rinmea Dyar, ‘‘Euzophera,’’ 313
Rioja Heinrich, n. gen., 182, 267
nexa Heinrich, n. sp., 267, 268
rippertella (Zeller), Megasis, 160
robusta (Moore), Hypsipyla, 27, 28
robustella (Dyar), Salebriaria, 116
ronnai (Bréthes), Cactoblastis, 245, 246
577
roseitinctella (Dyar), Ufa, 171
Rotruda Heinrich, n. gen., 184, 225
mucidella (Ragonot), 219, 225, 226,
227
mucidella affusella (Ragonot), 219, 227
mucidella olivaceella (Ragonot), 226
mucidella reliquella (Dyar), 219, 226
nimbella (Zeller), 219, 226
roxburghii Gregson, Ephestia, 302
rubedinella (Zeller), Ufa, 170, 171
rubescentella (Hulst), Nephopteryx, 125
rubicundella Zeller, Piesmopoda, 77, 78, 79
rubiginalis (Walker), Hulstia, 196
rubiginella (Walker), Adelphia, 169
rubribasella Hulst, Etiella, 99
rubrifasciella Packard, Acrobasis, 23
rubripurpurea Hampson, Stylobasis, 205,
206
rubrisparsella
131, 132
rubrithoracella (Barnes & McDunnough),
Sarata, 163
rubrizonalis (Hampson),
52, 53, 55
ruderella (Ragonot), Interjectio, 107
rufa (Haworth), Ephestia, 302
rufescentalis (Walker), Ufa, 171
rufibasella (Ragonot), Nephopteryx, 131
rufinalis (Walker), Adelphia, 169
rufitinctalis Hampson, Cayennia, 267
rufulella (Ragonot), Davara, 73, 76
Rumatha Heinrich, 182, 184, 248
bihinda (Dyar), 248, 249
glaucatella (Hulst), 248, 249
polingella (Dyar), 249, 250
russeolus Heinrich, n. sp., Birinus, 36
rusto (Dyar), Baphala, 236
rutilella Zeller, Cryptoblabes, 10
(Ragonot), Nephopteryx,
Pseudocabima,
saissetiae (Dyar), Baphala, 236
Salambona Heinrich, 1838, 254, 255
analamprella (Dyar), 254
Salebria Hiibner, 101, 112, 113, 114, 115,
119, 124, 153, 158, 312
Salebriacus Heinrich, n. gen., 7, 113,
115
bakerella (Dyar), 114
odiosellus (Hulst), 114
yumaella (Dyar), 114
Salebriaria Heinrich, n. gen., 7, 114,
118
ademptandella (Dyar), 115
annulosella (Ragonot), 116, 117
engeli (Dyar), 116
fructetella (Hulst), 115, 118, 119
georgiella (Hulst), 117
heinrichalis (Dyar), 117
nudiferella (Ragonot), 116, 117
pumilella (Ragonot), 115, 117
quercicolella (Ragonot), 117
rectistrigella (Dyar), 118
robustella (Dyar), 116
tenebrosella (Hulst), 117
turbidella (Ragonot), 115, 116
samaritanella (Zeller), Heterographis, 193
santa-maria Dyar, Pseudodivona, 48, 49
114,
115,
578
santiagella (Dyar), Erelieva, 308
Sarasota, Hulst, 9, 74, 76
furculella (Dyar), 76, 77
plumigerella Hulst, 76, 77
ptyonopoda (Hampson), 77
Sarata, Ragonot, 3, 159, 165, 166, 168
alpha Heinrich, n. sp., 164
aridella (Dyar), 161
atrella (Hulst), 2, 162, 164, 165
beta Heinrich, n. sp., 164
caudellella (Dyar), 162, 165
cinereella Hulst, 162, 163
delta Heinrich, n. sp., 165
dnopherella Ragonot, 159, 162, 163,
164, 165
edwardsialis (Hulst), 160
epsilon Heinrich, n. sp., 165
excantalis (Hulst), 161, 163, 164, 165
gamma Heinrich, n. sp., 164
incanella (Hulst), 160, 161
iota Heinrich, n. sp., 164
kappa Heinrich, n. sp., 163, 165
nigrifasciella Ragonot, 162, 163, 165
perfuscalis Hulst, 162, 164, 165
phi Heinrich, n. sp., 163, 165
polyphemella (Ragonot), 160
pullatella (Ragonot), 160, 164, 165
punctella (Dyar,) 161
punctella septentrionaria Heinrich, n.
race, 161
tubrithoracella (Barnes & McDun-
nough), 163
tephrella Ragonot, 162, 163
sauciella (Zeller), Ancylostomia, 96
schausella Dyar, Strephomescinia, 227
schausi Heinrich, n. sp., Megarthria, 87
schini (Berg), Cabotia, 201
schisticolor Zeller, Etiella, 99
scintillans (Grote), Pyla, 142, 146, 147, 148
Sciota Hulst, 123
scitulella (Hulst), Acrobasis, 14
sciurella Ragonot, Honora, 198
scobiella (Grote), Stylopalpia, 141, 142
Scorylus, Heinrich, n. gen., 3, 72
cubensis Heinrich, n. sp., 72, 73
secundella Ely, Acrobasis, 21
Selga Heinrich, n. gen., 182, 265, 266
arizonella (Hulst), 265
Sematoneura, Ragonot, 2, 27, 28
abitus Heinrich, n. sp., 27
atrovenosella Ragonot, 27, 51
semibrunneella Ragonot, Hyalospila, 59
semicana Heinrich, n. sp., Humysia, 188
semicana Heinrich, n. sp., Patriciola, 209,
210
semidiscella Ragonot, Cabotia, 200, 201
semifuneralis (Walker), Euzophera, 273,
276
semiobscurella (Hulst), Nephopteryx, 123,
124
semirufa (Haworth), Ephestia, 302
semirufella (Zeller), Piesmopoda, 80, 81
semtproniella (Schaus), Peadus, 84
Seneca Hulst, 11, 12
senesciella (Schaus),
68, 69
Anadelosemia, 67,
senorella (Ragonot), Anderida, 211
senta Heinrich, n. sp., Ufa, 170, 171
separabilis Heinrich, n. sp. Protomoerbes,
50
septentrionaria Heinrich, n. race (race of
punctella (Dyar)), Sarata, 161
septentrionella Dyar, Acrobasis, 19
sericaria (authors, not Scott), Anagasta,
300
sericartum (Scott), Ephestia, 301, 302
serratilineella Ragonot, n. status (race of
edmandsae (Packard)), Vitula, 285,
286
setonella (McDunnough), Manhatta, 287
Sigelgaita Heinrich, 183, 255
chilensis Heinrich, 255
huanucensis Heinrich, 256
transilis Heinrich, 256
similella Barnes & McDunnough, Acron-
cosa, 174
simplicula (Zeller), Metephestia, 264, 265
simulella Barnes & McDunnough, Divi-
tiaca, 189, 190
sinuella (Fabricius), Homoeosoma, 219
slossonella (Hulst), Hypargyria, 38
solitella (Zeller), Paramyelois, 46, 47
sonorella (Ragonot), Anderida, 211
soraella (Druce), Drescomopsis, 263
Sosipatra Heinrich, n. gen., 185, 294
anthophila (Dyar), 295
divergens (Dyar), 296
majorella (Dyar), 296
micaceella (Hampson), 295, 296
nonparilella (Dyar), 296
rileyella (Ragonot), 294, 295, 297
thurberiae (Dyar), 296
spaldingella Dyar, Eurythmia, 307
squalida (Walker), Baphala, 236, 237
squalidella (Dyar), Azaera, 282
squamifera Heinrich, n. sp., Megarthria, 87
Staudingeria Ragonot, 183, 194, 195
albipenella (Hulst), 194, 195
morbosella (Staudinger), 194
olivacella Dyar, 194, 195
perluteella Dyar, 195
stenopteryx (Dyar), Crocidomera, 33
Stenoptycha Heinemann (not Zeller), 272
stercorea (Zeller), Ancylostomia, 95, 96
stictella (Hampson), Ephestiodes, 278,
281, 306
stictoneurella Ragonot, Hyalospila, 35,
56, 57, 58
stictophorella Ragonot, Dasypyga, 69
stigmaferella Dyar, Ozamia, 258
stigmella Dyar, Acrobasis, 18
stigmella (Dyar), Rhagea, 237, 238
Strephomescinia Dyar, 184, 227
schausella, Dyar, 227
striatellum Dyar, Homoeosoma, 221, 222
striella Dyar, Zamagiria, 93
strigalis (Barnes & McDunnough), Cacto-
brosis, 248, 253, 260, 262
Strymaz Dyar, 227
Stylobasis Hampson, 185, 205
rubripurpurea Hampson, 205, 206
UNITED STATES NATIONAL MUSEUM BULLETIN 207
Stylopalpia Hampson, 3, 5, 140
argentinensis Heinrich, n. sp., 141
decimerella (Hulst), 141
lunigerella Hampson, 140, 141
scobiella (Grote), 141, 142
stypticellum Grote, Homoeosoma,
223
suavella (Zincken), Rhodophaea, 24
subaquilellus (Ragonot), Peadus, 84
subcaesiella (Clemens), Nephopteryx, 6,
127, 128, 129
subcanella (Zeller), “‘Zophodia,” 314
subcaudata (Dyar), Rampylla, 71
subcostella (Ragonot) (race of albocostal-
ialis (Hulst)), Pima, 105
subelisa Dyar, Drescomopsis, 262, 263
subfuscella (Ragonot), Nephopteryx, 123,
124
submedianella Dyar, Tacoma, 178
subosseella Hulst, Lipographis, 166, 168
subrufella (Hulst), Atheloca, 81, 82, 83
subsciurella Ragonot, Honora, 197, 198
substituta Heinrich, Nanaia, 245
subsutella (Schaus), Difundella, 63
subtetricella (Ragonot), Myelopsis,
Al
subtinctella (Ragonot), Cuniberta, 34
subtracta) Heinrich, n. sp., Dioryctria,
157, 158
subumbrella (Dyar), Olycella, 242, 243
supplicella (Dyar), Verina, 288
supposita (Heinrich), Rhodophaea,
24
sylphiella Dyar, Pyla, 147
sylviella Ely, Acrobasis, 16, 21
Synothmia Hampson, 205
sysstratiotes Dyar, Dioryctria, 151, 152
220,
40,
15.
taboga (Dyar), Baphala, 236
Tacoma Hulst, 178
feriella Hulst, 178
submedianella Dyar, 178
tapiacola Dyar, Tucumania, 252
tartarella (Zeller), Elasmopalpus, 173
tecmessella (Schaus), Anadelosemia, 67
Telethusia Heinrich, n. gen., 5, 136
geminipunctella (Ragonot), 137
latifasciatella (Packard), 137
modestella (Hulst), 137
ovalis (Packard), 136, 137, 138
rhypodella (Hulst), 137
tenebricosa (Zeller), Diatomosera, 50
tenebrosella (Hulst), Salebriaria, 117
tenuella (Barnes & McDunnough),
pargyria, 38
tenuipunctella Ragonot, Homoeosoma, 220
tephrella Ragonot, Sarata, 162, 163
tephrosiella Dyar, Ulophora, 176
termitalis (Hulst), Nephopteryx, 129, 130
tetricella (Schiffermiiller), Myelopsis, 40
texanella (Hulst), Anadelosemia, 68, 69
texanella (Hulst), Nephopteryx, 132
tecanella Ragonot, Homoeosoma, 220
thalassophila Dyar, Ozamia, 258
thurberiae (Dyar), Sosipatra, 296
ticitoa (Dyar), Vitula, 287
Hy-
tintilla Dyar, ‘“‘Euzophera,” 315
titillella Dyar, Microphycita, 310, 311
Tlascala Hulst, 3, 133
gleditschiella (Fernald), 134
reductella (Walker), 133, 134
tolerata Heinrich, n. sp., Difundella, 64, 65
torsicornis Dyar, Comotia, 217
Tota Heinrich, n. gen., 6, 169, 170
galdinella (Schaus), 170
Trachycera Ragonot, 2, 25
pallicornella (Ragonot), 25
transilis Heinrich, Sigelgaita, 256
transitella (Dyar, not Walker),
myelois, 45
transitella (Walker), Paramyelois, 47
translucida (Walker), Ufa, 171
trichomata (Zeller), Piesmopoda, 78
tricolorella Grote, Acrobasis, 13, 14
triloses Dyar, Mescinia, 213
trinitatis Heinrich, n. sp., Volatica, 290
triplagiatella (Dyar), Pyla, 145
truncatella (Wright), Lipographis, 166, 167
Tucumania Dyar, 183, 252, 253
porrecta Dyar, 252
tapiacola Dyar, 252
Tulsa Heinrich, n. gen., 5, 134
finitella (Walker), 134, 135
gillettella (Dyar), 135
infinitella (Dyar), 135
melanellus (Hulst), 134
oregonella (Barnes & McDunnough),
135
umbripennis (Hulst), 135
tumidella (Zincken), Acrobasis, 11, 17
tumidulella (Ragonot), Acrobasis, 24
turbatella (Grote), Zophodia, 238, 239
turbidella Zeller, Crocidomera, 32, 33
turpidella (Ragonot), Salebriaria, 115, 116
Ecto-
ubacensis (Zeller), Unadilla, 228
Ufa Walker, 7, 169, 170
deprivalis (Walker), 171
lithosella (Ragonot), 170, 171
luteella (Hulst), 170
minualis (Walker), 171
pyrrhochrellus (Ragonot), 171
roseitinctella (Dyar), 171
rubedinella (Zeller), 170, 171
rufescentalis (Walker), 171
senta Heinrich, n. sp., 170, 171
translucida (Walker), 171
venezuelalis Walker, 170, 171
ulmella (Ragonot), Carnarsia, 202
ulmiarrosorella (Clemens), Canarsia, 201, 202
Ulophora Ragonot, 1, 2, 176
dialithus (Dyar), 176, 177
groteii Ragonot, 176, 177
guarinella (Zeller), 177
tephrosiella Dyar, 176
umbrella (Dyar), Lipographis, 160, 166, 168
umbripennis (Hulst), Tulsa, 135
Unadilla Hulst, 1, 185, 218, 227, 282, 299
albidiorella (Richards & Thomson),
229
bipunctella (Hampson), 228
dorea (Dyar), 227, 228
INDEX
Unadilla Hulst—Continued
erronella (Zeller), 228, 229
floridensis Heinrich, n. sp., 229
maturella (Zeller), 218, 228, 229
masutella Hulst, 227, 228, 229
pyllis (Dyar), 228
ubacensis (Zeller), 228
uncanale Hulst, Homoeosoma, 220
uncanalis Ragonot (not Hulst), Homoe-
osoma, 223
uncta (Dyar), Erelieva, 308
undulatella (Clemens), Hulstia, 169,195, 196
unicolorella (Hulst), Oreana, 13, 112
unicolorella Staudinger, Ephestia, 302
uniformella Hampson, Ephestiodes, 281
unionellum Ragonot, Homoeosoma, 225
univitella Dyar, Anypsipyla, 42
uvinella (Ragonot), Nephopteryx, 123, 130
vaccinii Riley, Acrobasis, 13
Vagobanta Heinrich, n. gen., 185, 289
divergens (Butler), 289
Valdivia Ragonot, 182, 191, 316
aureomaculella (Dyar), 192
coquimbella Ragonot, 191, 192
lativittella (Ragonot), 191, 192
walkerella (Ragonot), 192
Varneria Dyar, 186, 305
atrifasciella Barnes & McDunnough,
305, 306
dubia Heinrich, n. sp., 306
nannodes Dyar, 305
postremella Dyar, 305
velessa Dyar, Harnocha, 202, 203
venata Hampson, Nasutes, 315, 316
venezuelalis Walker, Ufa, 170, 171
venipars (Dyar), Paramyelois, 47
venosa Dyar, Cacozophera, 208
venosella Turati, Ephestia, 304
vepallidum Heinrich, n. sp., Homoeosoma,
224, 227
vepreculella Ragonot, Ceracanthia, 86
verecuntella (Grossbeck), Palatka, 208
Verina Heinrich, n. gen., 184, 288, 289
Verina supplicella (Dyar), 288
vestilla (Dyar), Ephestiodes, 281, 309
vetustella (Dyar), Nephopteryx, 127
Vezina Heinrich, n. gen., 181, 291
parasitaria Heinrich, n. sp., 291
villosella Hulst, Etiella, 99
virgatella (Clemens), Nephopteryx, 128
viridisuffusella Barnes & MeDunnough,
Pyla, 142, 149
vitivora Filipjev, Ephestia, 302
Vitula Ragonot, 180, 184, 279, 283, 285,
287, 288, 289, 290, 291, 292, 313
dentosella Ragonot, 285
edmandsae (Packard), 285, 286, 287,
288
edmandsae serratilineella Ragonot, n.
status, 285, 286
inanimella (Dyar), 287
laura (Dyar), 285, 287
lugubrella (Ragonot), 286
pinei Heinrich, n. sp., 286, 288
ticitoa (Dyar), 287
579
vividella (McDunnough), Pima, 104
Volatica Heinrich, n. gen., 181, 290, 291
pachytaeniella (Ragonot), 290, 291
trinitatis Heinrich, n. sp., 290
walkerella (Ragonot), Valdivia, 192
walsinghami (Ragonot), Ambesa, 108, 109
Wunderia Grossbeck, 184, 204
neaeriatella Grossbeck, 204, 205
xanthaenobares Dyar, Dioryctria, 153
xanthomera Dyar, Piesmopoda, 78
xanthopolys Dyar, Piesmopoda, 79, 80
xanthoudemia (Dyar), Hyalospila, 58
zanthozona Dyar, Piesmopoda, 78
yavapaella Dyar (race of
(Zeller)), Eurythmia, 307
ydda (Dyar), Bema, 217, 218
yddiopsis (Dyar), Bema, 219
Yosemitia Ragonot, 183, 187, 250
didactica Dyar, 250, 251
fieldiella (Dyar), 250, 251
graciella (Hulst), 250, 251
graciella (Hulst, not Hulst), 251
longipennella (Hulst), 250, 251
yuconella (Dyar) (race of termitalis
(Hulst)), Nephopteryx, 130
yumaella (Dyar), Salebriacus, 114
hospitella
zamacrella Dyar, Laetilia, 230, 233, 234,
286
Zamagiria Dyar, 8, 90, 93, 94
australella (Hulst), 92, 171
bumeliella (Barnes & McDunnough),
92
deia Dyar, 93
dixolophella Dyar, 90, 91
fraterna Heinrich, n. sp., 92, 93
hospitabilis Dyar, 90, 91
ipsetona Dyar, 93
laidion (Zeller), 92
masculinus Dyar, 91
pogerythrus Dyar, 91
striella Dyar, 93
zeae (Fitch), Plodia, 299
zelatella (Hulst), Acrobasis, 12
zelleri, Ragonot, Acrobasis, 11
zeteki Heinrich, n. sp., Ectomyelois, 46
zimmermanni (Druce, not Grote), Promy-
lea, 66
zimmermani (Grote), Dioryctria, 149, 150,
152, 154, 155, 156, 157
zinekenella (Treitschke), Etiella, 99, 238
zonulella (Ragonot), Myelopsis, 41
Zophodia Hiibner, 181, 187, 197, 230, 233,
237, 238, 240, 257, 272, 315
bella Hulst, 239
convolutella (Hiibner), 238, 239
dilativitta Dyar, 239
franconiella Hulst, 239
grossularae (Riley), 239
grossularialis Hiibner, 239
grossulariella (Hiibner), 239
thouna Dyar, 239
magnificans Dyar, 239
turbatella (Grote), 238, 239
580 UNITED STATES NATIONAL’ MUSEUM BULLETIN 207
Species unknown, misplaced or
unrecognized
“Anthropteryx” irichampa Dyar, 313
“Hlasmopalpus” corrientellus Ragonot, 312
“Hucampyla” putidella Schaus, 313
“Wuzophera” came Dyar, 313
climosa Dyar, 315
conquistador, Dyar, 315
daedalella Ragonot, 314
gais Dyar, 313
intextella (Zeller), 314
mabes Dyar, 315
postflavida Dyar, 312
rinmea Dyar, 313
tintilla Dyar, 315
“Huzopherodes” megalopalis Hampson,
314
“Honora” dulciella Hulst, 197, 313
“Hornigia” clitellatella Ragonot, 234, 314
“Hypochalcia”’ cervinistrigalis Walker, 312
hulstiella Ragonot, 312
“Maricopa” albocostella Hulst, 192, 316
“Megasis” indianella Dyar, 160, 315
“Moodna” formulella Schaus, 313
olivella Hampson, 284, 314
“Myelois” atristrigella Ragonot, 315
famula Zeller, 312
grossipunctella Ragonot, 312
infusella Zeller, 314
restricteila Zeller, 312
“Nephopteryx” fuscifrontella Zeller, 312
“Phycitopsis” flavicornella Ragonot, 314
“Psorosa”’ disticta (Zeller), 314
“Salebria” nigricans Hulst, 312
“Zophodia” brevistrigella Ragonot, 314
epischnioides Hulst, 315
subcanella (Zeller), 314
Hosts
Abies, 150
Acacia farnesiana, 47
Achras sapota, 93
Achyranthus ramosissima, 190
Aesculus glabra, 47
Aleurocanthus, 11
Alnus, 23
Ambrosia, 279
Amorpha, 126, 127
californica, 126
herbacea, 126
Anaphalis margaritosa, 224
Annona squamosa, 44
Antennaria, 137
Anthemis, 220
Anthyllis, 102
Apple, 12, 14, 47, 113, 209, 273, 275, 284,
286
Apricot, 14, 273
Aster, 220, 226
Astragalus, 100, 102, 103, 106
Attalea funifera, 82
piassabossu, 82
Azalea, wild, 120
Balsam Fir (see Abies), 151
Bauhinia mexicana, 159
variegata, 60
Bean, 173
black, 60
Bell pepper, 308
Betula, 23, 120, 145, 284
Bidens, 214, 220
Black-eyed Peas, 60, 96, 172
Blueberry, 13, 135
Brauneria, 220
Bumelia microcarpa, 92
Butternut, 16
“Caimitillo,” 92
Cajanus, 96, 100
cajan, 60, 96
Canavalia ensiformis, 60
maritima, 60
Carapa guianensis, 29
Carica papayae, 74
Carissa, grandiflora, 45
Carnegiae gigantea, 260
Carya, 20
alba, 120
Cassia alata, 26
bicapsularis, 45, 61
brasiliensis, 42
corymbosa, 61
grandis, 47
meschata, 46
nodosa, 26
occidentalia, 60
spp., 61
tora, 26
Ceanothus, 114
Cedrela, 28
Celtis, 131, 132
Ceratonia siliqua, 44, 45
Cercis occidentalis, 296
Cereus validus, 259
Chaenomeles japonica, 10
Chamaecrista brachiata, 98
fasciculata, 98
robusta, 98
Cherry 12, 273
Catalina, 14
wild, 279
Chickpea, 96
Chokecherry, 13
Chrysanthemum, 220
Cicer, 96
Cirsium, 221
spinosisimum, 226
Citron, 10
Coccidae, 231, 232, 235
Coccolobis uvifera, 77
Cocos coronata, 82
nucifera, 82
vagans, 82
Colutea, 100
Comptonia, 23
Coreopsis, 220
Corn, 10, 173, 278, 284
Corylus, 21
Coryphantha aggregata, 251
Cotoneaster, 12, 25
Cotton, 173, 215, 220, 273, 279, 284
Cottonwood, 122
Cowpea, 60, 173
Crabgrass, 173
Crabapple, 12
Cranberry, 13
Crataegus 12, 13, 133
Crotolaria, 100, 106
Cylindropuntia, 245, 246, 248, 258
Cyperus exculentus, 173
Dahlia, 220
Daphne gnidium, 10
Dates, 45, 47
Denmoza, 246, 247
Dipholis salicifolia, 94
Dolichos, 96, 100
Hehinocereus, 250, 260, 262
pectinatus, 262
polyacanthus, 251, 253
Tigidissimus, 262
viridiflorus, 251
Echinopsis, 246, 247
Elephantopus, 199
Elm, 196, 202
Eremocarpus setigerus, 170
Erica, 145
Eriophyllum ignotum, 137
Erisbotyra japonica, 45, 93
Hulychnia acida, 256
Ferocactus wislizeni, 260, 261
Fig, 45, 47, 286, 303
Fir, 150
Flax, 173
Franseria bipinnatifida, 194
Fungus or pineapple, 297
Galls, of sawfly on willow, 129
on chokecherry, 14
Gaura parviflora, 279
Ginkgo, 273
Genipa americana, 47
Gleditsia, 134
triacanthos, 47
Glycina, 100
Grain, 301, 303
Grapefruit, 47
Grapes, 10
Hackberry, 202
Hasseanthus elongatus, 238
Helianthus, 220
Heliopsis, 220
Hickory, 15, 16, 17, 18, 19, 20, 22, 112,
120, 202
Homalocephala texensis, 250, 251, 260
Honeycomb, 285, 286
Hymenaea courbil, 44
Indigofera tinctoria, 265
Indigofera verbasifolium, 265
Inga, 218
Tris, 284
Jack pine, 151
Japanese cane, 173
Johnsongrass, 173
INDEX 581
Languncularia racemosa, 77 Orange, 10, 47, 220 Rose, 284
Lathyrus, 103 Orobanche ludovicana, 237 Rudbeckia, 220
maritima, 103 Ostrya, 16, 17 Salix, 123, 129
Leaves, dried, 288 virginiana, 21 caprea, 145
Lecanium sp. 232 Pacae, 42 Samanea samédn, 42
Lettuce, 226 Palafoxia, 197 Scale (see Coccidae), 233, 234, 235, 236
Limabeans, cultivated, 60, 90, 172 Peach, 47, 273, 284 Schinus molle, 201
wild, 60 Peanut, 173 Sciacassia siamea, 26
Linden, 273 Pear, 209, 273, 284 Seeds, 214, 284
Liquidambar, 273 Peas, garden, 60 Sitilias caroliniana, 226
styraciflua, 131 Pecan, 16, 17, 18, 19, 22, 273, 275 Sonchus asper, 226
Liriodendron tulipifera, 274 Peniocereus greggii, 260 Sorghum, 173, 308
Livistona chinensis, 45 Persimmon, 273 Sapindus drummondii, 47
Loco-weed, 106, 195 Phaseolus, 60, 100 “Spineless cactus,” 246
Locust, 124 Phaseolus lunatus, 60 Spruce, 150, 151
black, 173 Pigeon pea, 60, 96 “Stems of leguminous tree,” 55
Loquat, 284 Pine, 150, 152, 286 Strawberry, 173
Lotus, 102 Pinus, 150, 151, 153, 154, 155, 156, 284 Sudangrass, 173
Lupinus, 100 caribaea, 152 Sugar beet, 196
Lythrum, 11 chichuahuana, 154 Sugarcane, 173
Mammea americana, 45 coulteri, 156 Sunflower, 220
Maple, 101 monophylla, 157, 286 Sweetgum, 131
Maytenus phyllanthoides, 264 palustris, 158 Swietenia, 28
Mealy bugs (see Coccidae), 232 ponderosa, 152, 153, 156, 157 Swordbean, 60
Melanthera radiata, 214, 229 scopulorum, 156 Tachardiella argentina, 233
Milo Maize, 173 taeda, 156 Tagetes, 220
Mimusops emarginata, 93 Pisum, 100 Tamarack, 151
Mistletoe, 179 Pithecolobium flexicaule,”47 Tamarindus indica, 45
Mountain Ash, 273 Platypuntia, 240, 246, 254, 256 Taxodium distichum, 154
Mulberry, 273 Plum, 12, 14, 109, 273 Tamarix, 11
Myrica, 23 Poinciana gilliesi, 61 Tephrosia, 176
Myrica cerifera, 23 Pomegranate, 10 Theobroma cacao, 45
Neomamillaria, 251 Poplar, 273 Thurberia, 296 3
Nolina, 295 Populus, 122, 123 Trichocereus, 245, 246,"247, 256
Nolina parryi, 295 tremuloides, 123 Tobacco, 303
Nuts, 17, 45, 301, 303, 304 Prune, 12, 14, 286 chiloensis, 256
Nyssa sylvatica, 140 Prunus maritima, 209 Turnip, 173
Oak, 15, 21, 117, 118, 275 virginiana melanoearpa, 109 Vaccinium, 133
scrub, 24 Pseudotsuga, 150 myrtellus, 145
Oiketicus kirbyi, 291 Pulchea odorata, 229 Vachellia insularis,"45
Olive, 273 Puya alpestris, 289 Vegetable products,{dried, 284, 293, 299,
Onion, 10 301, 303
Pyracantha coccinea, 12
Ononis, 102 Viburnum, 101
Opuntia, 220, 254, 255, 257, 295, 308 I eee ld Vicina, 100
(Cylindropuntia) exaltata, 245, 247 Quince, 12 Vigna, 100
(Cylindropuntia) imbricata, 244, 248 | Raisin, 10, 45, 279, 286 unguiculata, 60
(Cylindropuntia) leptocaulis, 245, 249, | Razamofskya eryptopoda, 69 Viguiere, 220
250 5 7908
‘ : Rhus, 124, 221, 284 Walnut, 16, 20, 22,7120,8273
(Cyclindropuntia) sp., 258 toxicodendron, 166 black, 20
(Platypuntia) aurantiaca, 252
(Platypuntia) discolor, 252 Ribes, 239 f = ea
(Platypuntia) fiscus-indica, 247, 256 grossularia, 239 eat, 173,
(Platypuntia) sulphurea, 247, 254 Ricinus communis, 10 Wulffia, 214
(Platypuntia) sp., 240, 241, 242, 243, | Robinia, 45, 47 Ximenesia, 220
244, 246, 252, 257, 258 pseudoacacia, 128 Yucca, 47, 295
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