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Full text of "Bulletin - United States National Museum"

SMITHSONIAN 

INSTITUTION 




UNITED STATES 

NATIONAL MUSEUM 

BULLETIN 243 

WASHINGTON, D.C. 

1965 



PUBLICATIONS OF THE UNITED STATES NATIONAL MUSEUM 

The scientific publications of the United States National Museum, which is 
comprised of the Museum of Natural History and the Museum of History and 
Technology, include two series, Proceedings of the United States National Museum and 
United States National Museum Bulletin. 

In these series are published original articles and monographs emanating from 
the two Museums and dealing with their collections and work. These publica- 
tions set forth newly acquired facts in the fields of anthropology, biology, geology, 
history, and technology. Copies of each publication are distributed to libraries 
and scientific organizations and to specialists and others interested in the various 
subjects. 

The Proceedings, begun in 1878, are intended for the presentation, in separate 
form, of shorter papers. These are gathered in volumes, octavo in size, with the 
publication date of each paper recorded in the table of contents of the volume. 

In the Bulletin series, the first of which was issued in 1875, appear longer, separate 
publications consisting of monographs (occasionally in several parts) and volumes 
in which are collected works on related subjects. Bulletins are either octavo or 
quarto in size, depending on the needs of the presentation. 

Since 1902 papers relating to the botanical collections of the Museum have been 
published in the Bulletin series under the heading Contributions from the United States 
National Herbarium. 

This work forms number 243 of the Bulletin series. 

Frank A. Taylor, 
Director, United States National Museum. 
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Corythucha championi Drake and Cobben 



MUSEUM OF NATURAL HISTORY 



Lacebugs of 
the World 

A Catalog 
'Hemiptera: Tingidae) 



CARL J. DRAKE 

Research Associate, Smithsonian Institution 

and 

FLORENCE A. RUHOFF 



SMITHSONIAN INSTITUTION 
WASHINGTON, 1965 



CONTENTS 

Page 

Introduction 3 

Catalog of the Tingidae 22 

Species wrongly classified as Tingidae 445 

Types and their location 446 

Host plants 449 

Host plants cited only as common names 479 

Literature cited 482 

Index 557 

Index to host plants 623 

v 



Illustrations 



Figures 

i a, Stephanitis ligyra Drake; b, c, Tingis cardui (Linnaeus) 5 

2 Leptopharsa mira Drake and Hambleton 7 

3 Vatiga illudens (Drake) 9 

4 a, Gargaphia tiliae (Walsh) ; b, Cantacader amydis Drake 11 

5 Phatnoma agviates Drake and Runoff 13 

6 a, b, Cantacader afzelii Stal; c, d, Tingis cardui (Linnaeus) 15 

Plates 

(1-56 follow page 444) 

Frontispiece Corythucha championi Drake and Cobben 

1 Alloeoderes davao Drake 

2 Phatnoma pacifica Kirkaldy 

3 Phatnoma baltica Drake 

4 ^etekella pulla Drake and Plaumann 

5 Ceratocader armatus (Hacker) 

6 Cantacader tener Bergroth 

7 Litadea delicatula China 

8 Litadea delicatula China 

9 Cysteochila poecilia Drake and Ruhoff 

10 Ambotingis senta (Drake and Hambleton) 

1 1 Eteoneus peroronus Drake 

1 2 Alloeocysta approba Drake 

13 Epimixia vittata Horvath 

1 4 Agramma onar Drake 

1 5 Lullius spinifemur Drake 

16 Perbrinckea brincki Drake 

17 Onymochila dichapetali (Horvath) 

18 Copium clavicorne (Linnaeus) 
1 g Copium clavicorne (Linnaeus) 

20 Copium clavicorne (Linnaeus) 

21 Copium teucrii (Host) 

22 Copium teucrii (Host) 

23 Paracopium hamadryas (Drake) 

24 Paracopium hamadryas (Drake) 

25 Perissonemia ecmeles Drake and Mohanasundarum 

26 Dicrotingis digitalis (Drake) 

27 Codotingis evansi Drake 

28 Diplocysta trilobata Drake and Poor 

29 Hegesidemus otiosus Drake 

VII 



VIII ILLUSTRATIONS 

30 Diconocoris capusi (Horvath) 

3 1 Diconocoris capusi (Horvath) 

32 Australotingis jranzeni Hacker 

33 Australotingis Jranzeni Hacker 

34 Dictyla picturata (Distant) 

35 Dictyla veterna (Scudder) 

36 Ammianus alberti (Schouteden) 

37 Ammianus alberti (Schouteden) 

38 Ammianus dilatatus (Guerin-Meneville) 

39 Mecopharsa hacker i Drake 

40 Planibyrsa montei Drake and Hambleton 

41 Nobarnus signatus (Distant) 

42 Inoma multispinosa Hacker 

43 Hyalochiton colpochilus (Horvath) 

44 Galeatus spinifrons (Fallen) 

45 Galeatus sinuatus (Herrich-Schaeffer) 

46 Galeatus maculatus (Herrich-Schaeffer) 

47 Stephanitis typica (Distant) 

48 Stephanitis takeyai Drake and Maa 

49 Holophygdon melanesica Kirkaldy 

50 Gargaphia sororia Hussey 

51 Chorotingis indigena Drake 

52 Ypsotingis luzonana Drake 

53 Thaumamannia manni Drake and Davis 

54 Anommatocoris minutissimus China 

55 Anommatocoris coleopteratus (Kormilev) 

56 A, B, Anommatocoris zeteki Drake and Froeschner; c, Anommatocoris minutissimus 

China 



Lacebugs of the World 

(Hemiptera: Tingidae) 



Introduction 

The members of the family Tingidae are purely plant-feeding bugs that 
live mostly on the lower surfaces of the leaves of living plants. The thin 
lacy outgrowths of the pronotum and the delicate lacelike texture of the 
forewings (pis. 36, 41, 42) have quite aptly earned for them the common 
name "lacebugs" all over the world. Expressed polyglotally, lacebugs are 
called "netzwanzen" or " gitterwanzen" in German, "netwantsen" or 
"netwerkwantsen" in Dutch, and "chinches de encaja" in Spanish. The 
vernacular name in French includes that of the host plant, such as"le tigre 
du poirier," "le tigre du cerisier," and "le tigre du caffeier." 

Systematically, lacebugs are classified in the family Tingidae of the insect 
order Hemiptera. As cataloged herein, the family consists of 1,820 species 
arranged in 3 subfamilies and 236 genera. These figures include both 
living and extinct forms as far as now recorded in the literature; subgenera, 
subspecies, varieties, and invalid taxa are not included in the totals. 

Geographically, lacebugs are widely distributed in the torrid and tem- 
perate zones on all continents and on most islands in the oceans. They are 
unknown in the frigid zones. 

Lacebugs are relatively small insects, some nearly miniature in size (pis. 
1, 14, 45) and the "giants" seldom exceed one-fourth inch in length 
(pis. 36, 38). The overall form varies from long and very slender (pis. 
15, 16) to extremely wide (pis. 38, 40). Of the ineffable wonders of insect 
kinds, none surpasses the lacebugs in the diversity and oddities of forms 
(pis. 28, 30, 49) adorned with fancy lace designs. The lacework is original, 
specific, and thus peculiar to each kind of lacebug. No two species are 
ever dressed exactly alike. Only the adult members of the same species are 
clothed in identical topcoats of living lace. In size, color, markings, and 
general aspect, there are scarcely any differences in the outward appear- 
ances between the sexes. The classification of tribes, genera, and species 
is based for the most part upon the lacy outgrowths, especially the for- 
mations of the pronotum and the elytra (figs. 1-6). The buccal, sternal, 
and hypocostal laminae (figs. 1A, 3, 4) likewise furnish characterizing features. 

Adults and nymphs alike obtain food by piercing the epidermis of the leaf 
blades of living plants and extracting the sap from the cellular tissue therein 
by means of the protrusile and retractile stylets operating from a freely 
movable haustellum. Only the very slender stylets of the mouth parts 
pierce and enter the leaves to take out the sap on which they feed. 

The egg, except in the floral-gall makers, is thrust deeply into the leaf 
of the food plants almost up to the operculum and coated with a dark 
brownish adhesive material that soon hardens and forms a protective 

3 



4 U.S. NATIONAL MUSEUM BULLETIN 243 

coating. After hatching and drying, the nymphs begin to feed at once in 
small clusters near the empty eggshells and adults. If disturbed and scat- 
tered, they will soon reassemble again in a cluster. 

Metamorphosis is gradual. The growth and development from egg 
to the unchanging imaginal stage consists of five instars separated from one 
another by successive moults. Most species in the temperate zones are uni- 
or bivoltene and overwinter either in the imaginal or egg stage. Not much 
has been printed concerning the biology and estivation of tingids 
in the tropical region. 

Many kinds of plants, mostly the terrestrial flowering species, serve as 
feeding and breeding hosts. All species are rather highly specialized in 
their food habits, and generation after generation live on the same kind of 
plant or closely related ones. The life cycle of each species is attuned so 
that its breeding, egg-laying, and development period parallels in the 
growing time with that of the unfolding of the foliar (leaf-feeding species) 
or floral (gall-making species) parts of its host plant. 

Much injury and plasmolysis of the foliage always accompanies the feeding 
activities. Damage to the foliar tissue increases in severity with the hatch- 
ing of the eggs and development of the broods. Heavily infested leaves be- 
come badly discolored, turn yellowish to whitish, wither, and often drop pre- 
maturely. Serious losses are effected on many cultivated and wild plants of 
prime importance in agriculture and horticulture. For example, among 
the useful plants upon which lacebugs breed and obtain their livelihood are 
maize, sugarcane, cotton, castor bean, pineapple, papaya, avocado, pear, 
cherry, olive, tea, pepper, coffee, cinnamon, camphor, cacao, banana, coco- 
nut, cassava, rubber, walnut, oak, hazelnut, azalea, and rhododendron. 

According to preliminary experiments by Nagaraj and Menon (1956), 
recent field tests by Shanta, et al. (I960), and by Mathen (1960), the lacebug 
Stephanitis typica (pi. 47) is accused of being a vector of a serious disease of 
coconut palms in Travancore-Cochin, India. This disease (probably a 
virus) is called "wilt [root] of coconut palm" and seriously affects each 
year about one-fourth of the coconut area there under cultivation. S. 
typica is also a pest of consequence on banana, camphor, and other 
plants in India and the East Indies. 

As dwellers on the undersides of the leaves, tingids feed, stand, and rest 
in a topsy-turvy upside-down position. In this inverted posture, the lacy 
topcoat serves to conceal the body, obliterate its shadow, and thus functions 
in ways similar to the nets or screens devised by man to hide and camou- 
flage military installations, equipment, and troops in wartime. Lace 
curtains hanging over a window provide the same sort of concealment for 
mankind and objects inside a room. Diaphanous veils are worn to obscure 
or protect the face. The inherent ability of tingids to stand or "sit" still 
and then remain motionless for long periods is vital to survival. The least 
movement by the bug betrays its presence on the leaf. 



LACEBUGS OF THE WORLD 




elytron 
N hypocostal lamina 



a 



buccula 



rostrum 



ostiolaY canal 



d 




- - rrjrr rJJ- ~-"- J " parameres 



--proctiger 
— pygophore 



9 th paratergite 
i 




3 r gonapophysis 



subgenital 
plate 



-2™ gonocoxopodite 



gonocoxopodite 



ovipositor^,, •- 
extended 



Figure 1. — Structures used in classification: a, Stephanitis ligyra Drake, lateral aspect; 
b, Tingis cardui (Linnaeus), ventral aspect of cf genital capsule; c, same, lateral aspect 
of 9 genital segments. 

Cryptic coloration enters into the living picture in various ways. In the 
species of many genera, the paranota and elytra are broadly produced 
outwardly beyond the body limits (pis. 38, 40, 46). In such forms, 
the areolae differ in size, shape, and in the degree of iridescence, 
translucence, and transparence. The areolae in the dark elytral bands 



6 U.S. NATIONAL MUSEUM BULLETIN 243 

and other color markings often vary in translucence, whereas those in the 
clear areas are quite hyaline. The thin film of the areolae is usually irides- 
cent. Thus the color of the foliage on which the tingid feeds and rests can 
always be seen through those windowlike areolae of the topcloak that 
possess the property of transmitting rays of light. 

As the feeding injury develops, the various color changes effected on the 
leaf — from green through brownish to yellowish white — are likewise 
reflected through the panelike areolae to the degree that the cells are 
pervious to light. Lacebugs themselves do not possess any ability to 
undergo chameleonic changes in color. 

The immature stages present an altogether different external appearance 
in yet other ways than those of the lacy topcoats of their parents. Instead 
of the network, the dorsal surface of the nymph is smooth and plain (pi. 8) 
or armed with simple or modified spines of various kinds and sizes (pis. 31, 
33, 37). The dissimilarities between the imaginal and nymphal stages are 
depicted in the illustrations of such forms as Litadea delicatula (pis. 7, 8) 
from Reunion, Diconocoris capusi (pis. 30, 31) from Viet-Nam, and Ammianus 
alberti (pis. 36, 37) from Africa, and Australotingis jranzeni (pis. 32, 33) 
from Australia. 

The ability of the nymphs to crouch low and remain stock-still with 
their bodies pressed close to the surface of the leaves are characteristics 
inherited from their parents. Besides these behavioral traits, the castoff 
skins, eggshells, scattered dots of brownish-black excrement, and the somber 
color of the body also tend to make them less noticeable. The spines 
(pis. 33, 37), tubercles, and hairs on the dorsal surface are likewise factors in 
concealment. The nymphal stages of some species also possess glandular 
spines. 

Neither adults nor nymphs imitate or simulate in color, form, or behavior 
any animate or inanimate object in their habits. The species, in all life 
stages, are relatively sluggish, completely defenseless, and without any 
means or ways to fight back or even the speed needed to evade hungry 
marauders. To escape enemies, they rely entirely on behavioral features 
and camouflage. 

The feeding activities of some species of lacebugs cause the young foliar 
leaves of their host plants to curl downwards. According to Horvath 
(1929), the African tingid Onymochila dichapetali (pi. 17) causes extensive 
leaf-curl on the foliage of its host, Dichapetalum cymosum (pi. 17), in the 
Transvaal. The attack of the Japanese tingid Tingis comosa on the foliage 
of its host plant, Artemesia sp., produces a similar type of down-curl (fide 
Takeya); and, in India, the feeding of adults and nymphs of Corythauma 
ayyari (Drake) produces a leaf-curl on Jasminum spp. (fide Livingston). 
These are the only lacebugs so far known to effect this type of leaf damage 
and thus may indicate their roles as vectors. 

The members of two genera of lacebugs are true gall-producing insects. 



LACEBUGS OF THE WORLD 




frontal spines 
median spine 
occipital spines 

hood 

callus 

paranotum 

--T-. lateral carinae 

-1(4- median carina 



— costal area 

— discoidal area 



cephalic 
spines 



subcostal area 



sutural area 



elytron 



Figure 2. — Structures used in classification: Leptopharsa mira Drake and Hambleton, 

dorsal aspect. 

The offspring, from the neonate nymph to full maturity, are confined in 
closed, one-celled chambers. Only the genera Copium and Paracopium of 
the tingines are producers and dwellers in closed galls. These genera are 
represented in many countries in the Old World, but are unknown in the 
Americas. 

The gall-making lacebugs are anthophagous and induce most profound 
reactions on the flowers they affect. In marked contrast to the foliar 
feeding species that live open and free on the foliage, the gallicolous tingids 
cause the host plant to produce malformations known as flower cecidia or 
galls (pis. 20, 22, 24), within which the nymphal stages live, feed, grow 
and moult until they attain the imaginal stage. From a domiciliary 
standpoint, the closed gall forms a safe shelter and luxuriant abode for the 



8 U.S. NATIONAL MUSEUM BULLETIN 24 3 

progressive development of the immature stages. There is no reduction in 
the size of the legs and antennae (pis. 18, 21, 23) in the cecidogenous 
tingids as compared with those of free -living species (pis. 10, 11, 26). 

The conformation and structure of the eggs and the method of oviposition 
of members of the genera Copium and Paracopium differ notably from those 
of the tingids that do not produce galls. According to the observations of 
Monod and Carayon (1958) the egg of C. cornutum (now clavicorne) has a 
campanular or domelike operculum, whose surface is smooth like that of 
the body of the egg. The time and method of laying and the fixation of 
the egg to the corolla are connected with floral cecidogenesis. During the 
laying operation, the egg is turned end for end (180°) so that the opercular 
pole is inserted in the corolla. Conversely, the abopercular pole is thrust 
into the tissue of the living leaf by the nongall-making lacebugs. 

Two and a quarter centuries ago the gifted naturalist Rene A. F. Reamur 
(1737) of France wrote a classic account of an insect that produced galls 
on the flower buds of the plant "camedrys." He included figures of the 
gall and other parts of the plant as well as those of the adult and last nymphal 
stage of the insect that produced the galls. After opening many galls, 
Reamur recorded that each gall formed a one-celled chamber and confined 
therein was a single insect, either in the nymphal or adult stage. Years 
later, Geoffroy (1762) published a similar article about the same insect 
and its unichambered gall on the inflorescence of " chamaedras." In the 
12th edition of his"Systema Naturae," Linnaeus (1767) cited the above 
references, clearly authenticating the identity of the species previously 
named and described by him as Cimex clavicornis (now in the genus Copium) 
from Europe. Historically, this is the first gall-making lacebug to be 
technically named and described (pis. 18, 19). 

The tingicecidium is an abnormal growth or excrescence of the corolla of 
the flowerbud (pis. 20, 22, 24). The hypertropic growth contains a fairly 
large, closed, monothalamous cell, or chamber. The immature stages of 
the lacebug are confined therein from hatching until the completion of 
metamorphosis. At that time, the gall itself has also reached full maturity 
and bursts open naturally by dehiscence at the tip, thus setting free its 
haustellate inmate {Copium spp.) or inmates {Paracopium spp.). The adults, 
after escaping from the gall, become free-living like the leaf-feeding species. 

An examination of 100 nearly mature cecidia of Copium clavicorne (pi. 20) 
and an equal number of galls of C. teucrii (pi. 22) showed each cecidium of 
these species to be one-chambered and inhabited by only one tingid in 
either the last nymphal or newly emerged imaginal stage. The cecidia 
were preserved in alcohol as picked in the field from the host plant, Teucrium 
chamaedrys, in Austria. Although the galls examined were almost fully 
grown, they were still tightly closed and showed no external signs of 
dehiscence. 

The adults of C. teucrii (pi. 21) and C. clavicorne (pi. 18) are very similar in 



LACEBUGS OF THE WORLD 



antenniferous 
process * 



buccula 



paranotum 



constricted-- 
rostral sulcus 

ostiolar 

canal 




costal area — 



hypocostal 

lamina 



metarhoracic -=.— 
wings 



Figure 3. — Structures used in classification: Vatiga illudens (Drake), ventral aspect. 

size and habitus, and their separation from each other is based largely on 
the antennal characters of the adults. The last nymphal stage (pi. 19) also 
furnishes additional characters in the pronotal carinae. However, the cecidia 
(pis. 20, 22) of these two species are very dissimilar and thus provide much 
better distinguishing characters than those of either the nymphal or imaginal 
forms. 

The dissection of 9 nearly mature cecidia of Paracopiam hamadryas (pi. 24) 
from Malagasy showed each cecidium to be monothalamous, closed, and 
to contain from four to seven lacebugs in the last nymphal and adult stages. 
A census of nine galls gave these figures: 2 galls with 2 nymphs and 2 adults; 
3 galls with 3 nymphs and 2 adults; 2 galls with 3 nymphs and 3 adults; 
and 2 galls with 3 nymphs and 4 adults. These figures give an average 
population of 5.4 lacebugs per gall. From one to three young larvae of 
nolid moths, assumed to be scavengers, were found in four cecidia. The 
genera Copium and Paracopium comprise a total of 45 species (10 -f 35) and, 
insofar as known, are all gall-forming tingids. 

681-552— ,65 2 



10 U.S. NATIONAL MUSEUM BULLETIN 243 

Lacebugs are no new creations. According to the geological time 
calendar, the petrified forms found in stone (pi. 35) and moulds engulfed in 
amber (pi. 3) represent bygone species that lived many million years ago. 
No present-day lacebug is also known as a fossil. As the illustrations 
depict, the extinct species as well as the forms living today wore topcoats 
of living lace. Although specifically different from the living forms, the 
zoolithic Phatnoma baltica (pi. 3) and Dictyla veterna (pi. 35) are typically 
members of these respective genera that were erected to hold living species 
(pis. 2, 7). Both of the genera are represented today by members of modern 
species in the Old and New Worlds. 

The present picture of the fossil tingids, because of the imperfect state 
of our knowledge, is much confused and in need of specific and generic 
revision. Four forms, each described simply as " Tingis sp." are without triv- 
ial names and probably do not even belong to this genus. Bachofen- 
Echt (1949, fig. 160) published an excellent photograph of an unidentified 
amber lacebug which may be Phatnoma baltica Drake (1950) described in 
Baltic amber. 

The lacebug species common to the Old and New Worlds are numerically 
low, totaling only nine species. A digest of the literature and studies of 
numerous collections have so far not revealed evidences of any natural 
faunal exchange through migration in either direction. Apparently, all 
the tingid species shared similarly between the two worlds made the sea 
voyage by ships of various kinds in transoceanic commerce. Paleontologic 
and chorologic data disclose no evidences of the so-called Amphiatlantic or 
Westarctic distribution in the Tingidae, and collections from eastern 
Siberia and northwestern North America (Alaska and Canada) have not 
as yet disclosed a single species common in these regions, even among the 
muscicolous genus Acalypta. The members of the latter genus are among 
the northernmost Tingidae in these regions. Apparently, perhaps, even 
in the present era, the Bering Strait plus unfavorable biotic factors such 
as low temperature and floral composition have formed a natural barrier 
for tingids between the Asian and North American continents. 

Ballast materials, used quite extensively on sailing vessels in the colonial 
times, played a significant role in the transportation and introduction of 
certain kinds of insects and other invertebrates as well as many plants into 
North America from Europe (Lindroth, 1957). However, our studies of 
the tingifauna collected at various times in the vicinity of ballast-dumping 
sites near old seaports along the Atlantic and Pacific coasts of North 
America indicate that earthen and other sorts of ballast seem to have 
played no findable part in the hemispheric exchange of Tingidae. 

Caloloma uhleri Drake and Bruner is an Australian indigene originally 
described from the West Indies (Antigua, Lesser Antilles). Specimens 
received recently from the Windward Islands (Saint Vincent) show that 
this lacebug has become established in the West Indies. From its native 



LACEBUGS OF THE WORLD 



11 



mesosterna 
lamina 

metasrernal 
lamina 



mesosrernum 



\ metasternum 



notal carinae (5) 




discoidal area 

— stenocostal area 

— costal area 

subcostal area 



surural area 



Figure 4,-Structures used in classification: a, Gargaphia tiliae (Walsh) interrupted rostra 
channel on metasternum; b, Cantacader amydis Drake, dorsal aspect. 



12 U.S. NATIONAL MUSEUM BULLETIN 24 3 

home far away, we have specimens from Queensland, New South Wales, 
and South Australia. This is the only known Australian tingid colonized 
in the Americas. 

Contrary to the usual statements, Stephanitis rhododendri Horvath is indig- 
enous in the United States and Canada. It is not a European element 
transported to North America from Holland as generally credited. Mu- 
seum specimens bearing locality and collecting dates prove that the rhodo- 
dendron lacebug was widely dispersed in eastern and central parts of the 
United States and in southern Canada at the time the species was 
originally described from Holland. This indigeneity of rhododendri is in 
agreement with the findings and hypothesis proposed by Johnson (1936) 
relative to its native patria. Furthermore, in general aspect, rhododendri 
is systematically nearer to S. blatchleyi Drake of Florida than it is to members 
of this genus in the Old World, or even to its South American congeners. 
Shipments of nursery stock of the decorative rhododendrons and azaleas 
account for its disjunct, synathropous dispersion in the Old World. Since 
the tingid rhododendri and also pyrioides overwinter in the egg stage, the erica- 
ceous hosts with persistent leaves serve as insidious conveyors of the eggs 
in plant shipments. 

Teleonemia scrupulosa Stal is indigenous in Mexico, West Indies, Central 
and most of South Americas. It was experimentally introduced in the 
Hawaiian Islands from Mexico for the "biological control" of the noxious 
lantana plant. As a result of the Hawaiian tests, scrupulosa was intentionally 
transported and released in numbers for the same biological end in the 
East Indies, Australia, India, and Africa, where it is now firmly colonized 
and becoming widely spread. This is the only tingid species employed in 
the biological control of troublesome plants and, furthermore, the only 
New World lacebug intentionally imported into the Eastern Hemisphere. 
No lacebug has been purposely introduced into the Americas from the Old 
World. 

Corythucha morrilli Osborn and Drake, a recent overseas transport into the 
Hawaiian Islands, is a native of southwestern United States, Mexico, 
Central America, and the West Indies. Records indicate that it was 
probably accidentally included in shipments of nursery plants from south- 
western United States to Hawaii. 

Tingis auriculata (Costa), indigenous in several European countries, was 
collected by Uhler many years ago in Maryland and has not been taken 
there since. It probably was unable to self-maintain its population in 
eastern United States. 

Dictyonota tricornis (Schrank) is an indigene in many countries in Europe, 
central Asia, and northern Africa. Although discovered in eastern Canada 
and New England States a half a century ago, its increase and spread for 
an introduced species have been slow. The North American specimens from 



LACEBUGS OF THE WORLD 



13 



dlscoidol area-- 



sutural area -- 




cephalic spines (7) ^c~-~^r.r 



paranotum--£TT 

collar 

callus 

pronotum 

median carina- - 

scutellum__/^ 
costal area 



subcostal — 
area 



Figure 5. — Structures used in classification: Phatnoma agviates Drake and Ruhoff, dorsal 

aspect. 

Nova Scotia and Maine are typical tricornis and not a variety or subspecies 
as described. 

Dictyonota fuliginosa Costa is indigenous in several countries in eastern 
Europe. Scudder (1960) recently collected several specimens on broom 
in western Canada (British Columbia), probably introduced there on 
nursery stock several years ago. 

Stephanitis pyrioides (Scott), originally described from Japan, is an Asian 
indigene and globally known as the azalea lacebug. Its synanthropic dis- 
persal, like that of S. rhododendri, is associated with the wide culture and im 
portation of azaleas and rhododendrons, both greenhouse and landscape 
forms and varieties, and likewise is closely bound to its ericaceous hosts. 



14 U.S. NATIONAL MUSEUM BULLETIN 243 

Records show that pyrioides was first introduced into Europe from Japan and 
then later into North America either from Europe or Japan, or perhaps 
both. Through the transportation of nursery stock, pyrioides has also 
gained entrance into Morocco, South Africa, and Australia. 

Stephanitis takeyai Drake and Maa (pi. 48) is a Japanese indigene rather 
recently imported into eastern United States, apparently on nursery stock 
from Japan. It is known from Connecticut and New Jersey in the United 
States. 

Galeatus spinifrons (Fallen) (pi. 44), originally described from Sweden, is 
indigenous in many countries in Europe and Asia. It was the first lacebug 
accidentally transported into North America (probably New England or 
Newfoundland) from Europe. The synonymity of G. angusticollis Reuter 
and G. spinifrons (Fallen) of Europe and of G. peckhami (Ashmead) and G. 
uhleri Horvath of the United States and Canada gives the survivor spinifrons 
circumglobal distribution. It is the only tingid species that has encircled 
the world in distribution. In Europe and Asia, both macropterous and 
brachypterous forms are fairly common, whereas an examination of more 
than 500 specimens (all long-winged) from northern United States and 
southern Canada, indicates that perhaps only the homozygous, macrop- 
terous form was probably imported into northern United States. Galeatus 
is an Old World genus, and G. spinifrons is pterygopolymorphic in Eurasia. 

Founded on morphologic structures from a global concept, Drake 
and Davis (1960) proposed the superfamily Miroidea Hahn to hold the 
families Tingidae Laporte and Miridae Hahn, and then systematized the 
Tingidae into the subfamilies Cantacaderinae Stal, Tinginae Laporte, and 
Vianaidinae Kormilev. In consummating these hierarchic changes, the 
subfamily Agrammatinae Douglas and Scott (formerly Serenthinae 
Stal) was suppressed as a synonym of Tinginae and the rank of the family 
Vianaididae Kormilev was lowered to that of subfamily and then, along 
with its generic taxa in totality, transferred to the Tingidae. The super- 
family Miroidea falls in the higher hierarchial Cimicomorpha (Leston et 
al. 1954). 

For almost a century, catalogs, synopses, textbooks, and other publi- 
cations had either treated the Piesmatidae as a subfamily of Tingidae (Stal 
1874, Horvath 1906) or as a separate family. Reuter (1912, pp. 49, 56) 
classified the Piesmatidae and Tingidae together in the superfamily Tingi- 
toidea. Recently, Leston et al. (1954) and Drake and Davis (1958, 1960) 
have clearly shown that these two families are not so closely related to each 
other and that they belong to different familial groups, the Piesmatidae to the 
Pentamorpha and the Tingidae, as stated above, to the Cimicomorpha. 
Among the structural differences between these families, it should be noted 
that the Piesmatidae (genus Piesma, ventral sterna V, VI) possess tricho- 
bothria on the underside of the abdomen, whereas trichobothrial hairs are 
lacking in the Tingidae. 



LACEBUGS OF THE WORLD 



15 



_ clavus 



subcosta 




hypocosta 




__ I s * anal 
vein 

_ post-cubitus 



_ secondary 
veins 

_ cubitus 



-radius 
-media 

-cubitus 

,_ subcosta 

_ radius- 
media 




clavus 



hypocosta 



.radius- 
media 



_L_cubitus 



post- 
cubitus 



subcosta 



secondary 
veins 

cubitus- 

media 




_ media 
_ subcosta 

_ fused 
radius & 
media 

— cubitus 



Figure 6. — Structures used in classification: a, Cantacader afzelii St&l, venation of fore- 
wing; b, same venation of hindwing; c, Tingis cardui (Linnaeus), venation of forewing; 
d, same venation of hindwing. 



16 U.S. NATIONAL MUSEUM BULLETIN 24 3 

The subfamilial and tribal keys are based upon fully mature specimens, 
are worldwide in scope, and use only those structural features that apply 
equally well to individuals of all species regardless of sex, brachyptery (pis. 
1, 45) or machroptery (pi. 46), and distribution. As dichotomized herein, 
the authors have embodied the higher categories formulated by Drake and 
Davis in their basic treatment of the Tingidae. 

For the sake of uniformity in tingitaxonymy, the structures (figs. 1-6) 
customarily used in the description of species, genera, and in the formation 
of taxonomic keys are semidiagramatically figured with their respective 
technical names affixed. Some nymphs (pis. 31, 33, 37) and many adults 
(pis. 2, 43, 52, 54) are illustrated to depict clearly the habitus of represent- 
ative forms found in various subfamilies and tribes. 



Key to Subfamilies of Tingidae 

Dorsal surface finely to coarsely punctate, never lacelike; compound eyes vestigial, 
each with a small number of poorly developed, usually scattered ommatidia; 
antenna with segment I shortest, II and III nearly equal, IV longest; pronotum 
without discal carinae, each lateral margin above with a very narrow low 
ridge; scutellum fairly large, triangular, exposed; ostiole and furcated ostiolar 
canal unusually prominent, somewhat Y-shaped, the vertical sulcus furnished 
with a backward projecting branch; evaporatorium enormously developed, 
covering entire metapleuron, hind part of mesopleuron, and then extending 
downward on the respective thoracic sterna to the laminar ridge of rostral 
sulcus; abdominal sterna II through IV fused, other sterna free. Macrop- 
terous form unknown. Myrmecetophiles (pis. 53, 54, 55, 56). 

Subfamily Vianaidinae Kormilev (p. 443) 

Dorsal surface finely to coarsely lacelike; compound eyes fully developed; each 
with a large number of normal ommatidia; antenna with segment I slightly 
to much longer than II, II usually shortest, sometimes I and II subequal, III 
customarily longest, rarely III and IV subequal lengthwise, IV commonly 
moderately long; pronotum rarely noncarinate (pi. 1), almost always carinate 
with carinae present in combinations of one (pis. 4, 1 4), three (pis. 3, 5, 9), or five 
(pi. 6); scutellum small, exposed (pi. 2) or concealed (pis. 10, 12, 13) beneath 
hind margin or backward process of pronotum; ostiole and ostiolar canal 
of each metapleuron either present or absent, when present (figs. \a, 3) 
with a slightly arcuate, nonbranching, nearly vertical canal, without distinct 
evaporatorium; abdominal sterna II and III fused, other sterna free. Pter- 
ygopolymorphic forms found in many species 2 

Head always long, much produced in front of compound eyes, nearly porrect, 
unarmed or armed with one to nine tubercular or spiniform processes; buc- 
culae long, wide, straight, with anterior ends extending forward beyond apex 
of clypeus, or shorter with ends curved inward and more or less meeting mesad 
in front of labium; antenna with segments I and II always short, apex of 
latter not reaching apex of clypeus; pronotum rarely noncarinate (pi. 1), 



LACEBUGS OF THE WORLD 17 

usually carina te with carinae arranged in combinations of one (pi. 4), three 
(pi. 5), or five (pi. 6); hind margin of pronotum little produced backward, 
wide, slightly sinuately or obtusely angulately truncate, never triangularly 
prolonged backward; scutellum small, either exposed or covered by hind 
margin of pronotum; clavus large, well developed, always entirely visible 
(pi. 2), often more or less fused with discoidal area (pi. 4); abdominal sterna 
II and III fused, other sterna free. 

Subfamily Cantacaderinae Stal (p. 22) 
Head either (1) very short (pi. 50), only slightly produced in front of compound 
eyes, sharply declivent in front; antennal segment I mostly surpassing apex 
of clypeus (pis. 9, 10), or, (2) very long (pi. 51), subporrect, much produced 
in front of compound eyes, apex of antennal segment I and sometimes also 
that of II not surpassing apex of clypeus (pi. 51); unarmed or armed with 
one to five (rarely seven) tubercular or spinelike processes; bucculae either (1) 
short, with anterior ends not extending beyond lateral sides of labium, or, 
(2) long, with ends curved inward and almost or meeting mesad in front of 
labium; pronotum carinate, carinae present in combinations of either one 
(pis. 7, 14, 40) or three (pis. 9, 12); hind margin of pronotum triangularly 
prolonged backward (pi. 10), occasionally with apex much abbreviated, 
always completely covering scutellum and clavi of elytra in resting posture; 
abdominal sterna II and III fused, other sterna free. 

Subfamily Tinginae Laporte (p. 42) 

The cantacaderines comprise a moderately large, sharply defined group 
and represent the oldest of the living lacebugs. Of the 21 known genera, 
Phatnoma is the only genus shared by the Old and New Worlds. The 
subfamily is unknown in the Americas north of Mexico. 

Key to Tribes of Cantacaderinae 

Elytron with a narrow, uniseriate stenocostal area lying between the costal area 
and outer margin of elytron (fig. 4; pis. 5, 6) .... Cantacaderini Stal (p. 22) 

Elytron without stenocostal area, the outer margin of costal area forming the 
exterior boundary of elytron (fig. 5; pis. 2, 4). 

Phatnomini Drake and Davis (p. 30) 

The tingines excel by far the other subfamilies both in number of species 
and populations, and are well represented in all continents. Almost all 
the lacebugs that live at the expense of cultivated, decorative, and wild 
plants of importance in human welfare belong to this subfamily. The 
tribes may be separated thus: 

Key to Tribes of Tinginae 

1. Head very long, greatly prolonged in front of compound eyes, subporrect; apex 
of antennal segment I (sometimes also that of II) not surpassing apex of 
clypeus (pis. 51, 52) Ypsotingini, new tribe (p. 428) 



18 U.S. NATIONAL MUSEUM BULLETIN 2 43 

Head very short, little produced in front of compound eyes, sharply declivent; 
antennal segment I surpassing by most of its length the apex of clypeus (pis. 

9»39) 2 

2. All tarsi slender, segment II at most only slightly swollen (pis. 9, 29). 

Tingini Laporte (p. 42) 

All tarsi greatly swollen; segment II enormously swollen, nearly ovate in outline, 

upper surface convex, beneath deeply concave, the concavity filled with short, 

bristlelike hairs (pis. 7, 8) Litadeini, new tribe (p. 42) 

The vianaidines are indigenous in the Neotropical region and known 
only from there. Four species are described, all being found inside the 
subterranean nests and underground passageways of ants. As to the 
natural history and relation between the vianaidines and ants, very littie 
is known. All species are myrmecophiles with obsolete ommatidia, and 
perhaps blind. The ostiole and ostiolar canal are very prominent and 
provided with an extremely large evaporatorium (pi. 56). 



Key to Genera and Species of Vianaidinae 

1. Broadly ovate, widest behind middle of elytra, there width about two-thirds 

that of median length; paranotum and costal area wide, equally explanate, 
each with large rounded punctures, base of pronotum and elytron distinctly 
punctate; hypocostal lamina wide, obtusely angulately widened near base, 
there biseriate, thence posteriorly uniseriate (pi. 53). 

Thaumamannia manni Drake and Davis (p. 444) 
Elongate-ovate, elytra at widest point less than half the median length; parano- 
tum and costal area absent; posterior part of pronotum and elytron distinctly 
punctate; hypocostal lamina very narrow (usually mistaken for costal area), 
composed of one row of tiny punctures 2 

2. Backward branch of the vertical ostiolar canal situated only slightly above the 

hind acetabulum so as to divide the posterior part of the metapleural evapora- 
torium into unequal divisions (pi. 56). 

Anommatocoris zeteki Drake and Froeschner (p. 444) 

Backward branch of the vertical ostiolar canal placed higher on the metapleural 

evaporatorium so as to divide the evaporatorium into two nearly equal 

divisions 3 

3. Compound eyes with only a small cluster of poorly developed ommatidia; 

elytron finely punctate for most of its length, shortly pubescent (pi. 55). 

Anommatocoris coleopteratus (Kormilev) (p. 444) 

Compound eyes with a few scattered, almost obsolete ommatidia; elytron 

punctate on basal part, thence posteriorly impunctate, inconspicuously 

pubescent (pi. 54) Anommatocoris rninutissimns China (p. 444) 

In this catalog an attempt has been made to record the scientific names 
and technical literature published on lacebugs since the beginning of 
systematic entomology, which dates from the publication of Linnaeus' 
Systema Naturae, 10th edition, 1758. 



LACEBUGS OF THE WORLD 19 

A uniform method is followed in citing references. In the text and 
synonymies the name of the author is followed by a year and page reference, 
with the full tide appearing in the complete bibliography at the end of the 
text. In case an author has published more than one paper in a given year, 
the chronologic order is indicated in every case by an alphabetic affixation 
to the years. 

The three subfamilies of Tingidae, their respective tribes, and the genera 
within the tribes are arranged in alphabetical sequence, each in their 
respective categories. The species are likewise arranged in alphabetic order 
in their respective genera. Synonyms, homonyms, and other errata are 
arranged chronologically under the valid specific name. Citations to other 
catalogs, faunal lists, monographs, and keys to genera or species are in- 
cluded in the generic citations. In addition to the original and later 
descriptions, specific citations include references to illustrations, morphology, 
biology, ecology, host plants, and distribution. Distributional records are 
in summary by countries. The original technical name of the type species 
of a genus always precedes the list of species. All elements in boldface type 
are regarded as of equal taxonomic value. 

The incorporation of references dealing with biology, morphology, and 
plant injury as well as those concerned with systematic entomology makes 
the catalog a wide source of subject matter in different technical spheres. 

To obtain the full literature for any one species, the generic references as 
well as those beneath the specific name should always be checked. The 
specific references occurring in such publications as catalogs, lists of species, 
and keys are listed beneath the generic names but not always beneath the 
trivial unless illustrations or morphologic or biologic data are involved. 
This combination of references (generic plus specific) includes the impor- 
tant citations to each species. 

Information on the type (holotype or lectotype) specimen is recorded 
when it was included in the original description, published later, or obtain- 
able from an institutional deposition. As will be noted in the text, the type 
specimens of a considerable number of species are either "lost" or "un- 
known." For full account of the history and type fixation of tingid genera, 
consult " Lacebug Genera of the World" (Drake and Ruhoff 1960a). 

The information on type locality is included as it was written in the 
original description or as it appears on the locality label of the type specimen. 
Political divisions, created by events in human history of the present day, 
have made it ever difficult to keep pace with the changes in the names of 
localities and countries for distributional records. Our efforts in up-to- 
dateness have not been entirely successful. However, for such information 
we have followed the maps recently published by the National Geographic 
Society (Washington, D.C.), The Columbia Lippincott Gazetteer of the 
World, Webster's Geographic Dictionary (rev. ed., 1957), and information, 
current and historical, obtained from the Division of Philately and Postal 



20 U.S. NATIONAL MUSEUM BULLETIN 243 

History of the United States National Museum. In the case of the Union 
of Soviet Socialist Republics, due to the uncertainties in some geographic 
boundaries, it will be noted that both general areas (Caucasus, Crimea, 
Siberia, etc.) and separate republics (Latvia, Dagestan, etc.) are used. 
The term "Ostindien," as used by Fieber (1844), is vague and can refer to 
either the East Indies or eastern India. Specimens from the Fieber collec- 
tion deposited in the Naturhistorisches Museum, Vienna, bear simply the 
label " Ostindien." 

As used in the text of this catalog, an asterisk (*) placed just before the 
scientific name means that the species is represented in the U.S. National 
Museum and a dagger ( |) indicates that it is a fossil species. Symbols or 
marks used in the index only are explained immediately beneath the index 
caption. 

Of the 1,820 valid species cataloged herein, 1,491 species are represented 
in the U.S. National Museum. This number includes 669 holotypes (or 
lectotypes) (of which 586 are in the Carl J. Drake collection and 83 in the 
Museum collection). The Drake collection contains 1,482 different species 
of lacebugs, of which 945 species are represented by a type of some kind. 

Acknowledgments 

For the loan of and comparison of Tingidae in their respective museums, 
the authors express here their most grateful thanks to Dr. S. L. Tuxen, 
Universitetets Zoologiske Museum, Copenhagen; Dr. E. Kjellander, 
Naturhistoriska Riksmuseet, Stockholm; Dr. W. E. China and Mr. R. J. 
Izzard, British Museum (Natural History), London; Dr. Max Beier, 
Naturhistorisches Museum, Vienna; Dr. Jacques Carayon, Museum 
National d'Histoire Naturelle, Paris; Dr. Ludvig Hoberlandt, Narodni 
Museum, Prague; Dr. Henri Schouteden, Musee Royal de l'Afrique 
Centrale, Tervuren, Belgium; Dr. John W. Beardsley, Experiment Station, 
Hawaiian Sugar Planters Association, Honolulu, Hawaii; Dr. J. Linsley 
Gressitt, Bishop Museum, Honolulu, Hawaii; Dr. Hugh B. Leech, Cali- 
fornia Academy of Sciences, San Francisco, Calif.; Dr. L. Kocher, Institut 
Scientifique Cherifien, Rabat, Morocco. 

Also to Dr. Richard Froeschner and Dr. Reece I. Sailer, U.S. Depart- 
ment of Agriculture, and Dr. J. F. G. Clarke, U.S. National Museum, 
for helpful suggestions and reading parts of the manuscript; Mr. Curtis 
Sabrosky, U.S. Department of Agriculture, for solving numerous nomen- 
clatorial problems and interpreting the Code of Zoological Nomenclature; 
Dr. Syoiti Miyamoto, Kyushu University, Fukuoka, Japan, for microfilms 
and for translations of papers written in Japanese; Dr. John J. Wurdack, 
U.S. National Museum, Department of Botany, for checking the botanical 
names and placing them into families; and to Mr. Theodore B. Ruhoff, 



LACEBUGS OF THE WORLD 21 

U.S. National Museum, Division of Philately and Postal History, for 
assistance with geographical problems. 

For the fine illustrations we are indebted to Elsie Froeschner and Caroline 
Lutz, both of Arlington, Va.; Maria Lisa Biganzoli and Elinor Stromberg, 
both of Washington, D.C.; Patricia and Robert Hogue, Alexandria, Va.; 
and to Arthur Smith, British Museum (Natural History), London. 

Special acknowledgment is also due to Emily Bennett and Ruth Blanchard 
and the staff of the Smithsonian Institution Library for assistance in locating 
and obtaining sources of references; and for translations of papers in foreign 
languages, we are indebted to the late Ruth Ericson, translator, U.S. 
Department of Agriculture. Mr. John S. Lea and Mrs. Mary M. Ingraham, 
of the Smithsonian editorial staff, have been most cooperative and helpful 
with suggestions on format and in seeing the manuscript through the press. 

This work was supported in part by National Science Foundation Grant 
18721. 



Family TING ID AE Laporte 

Cimicjdes Latreille 1807, p. 126 (Tingis, p. 139). 

Geocorises Latreille 1829, p. 201. 

Tingidites Laporte 1833, p. 47. 

Tingidae Westwood 1840a, p. 447; 1840b, p. 120. — Lethierry and 

Severin 1896, pp. 1, 4.— China and Miller 1955, p. 261; 1959, pp. 9, 

34.— Drake and Ruhoff 1960a, p. 3.— Drake and Davis 1960, p. 4 

(morphology and higher classification). 
Ductirostri, Group Tingides Amyot and Serville 1843, pp. 285, 295. 
Tingiditae Spinola 1852b, p. 27. 

Tingididae Fieber 1861, pp. 35, 116, 400.— Oshanin 1908, p. 395. 
Tingitidea Costa 1863, p. 6. 
Tingitidae Stal 1873, pp. 115, 116— Horvath 1906a, p. 1; 1911d, 

p. 14.— Oshanin 1912, p. 42.— China 1943, p. 245. 

Subfamily Cantacaderinae Stal 

Cantacaderaria Stal 1873, p. 116. — Horvath 1906a, p. 10. — Oshanin 

1908, p. 400 (as divisions). 
Cantacaderini Champion 1897, p. 2 (as group). 
Cantacaderaria Oshanin 1912, p. 42 (as tribe). 
Cantacaderinae Stichel 1926, p. 103. — China and Miller 1959, p. 

34.— Drake and RuhofT 1960a, p. 26.— Drake and Davis 1960, pp. 

74, 75. 
Type Genus: Cantacader Amyot and Serville. 

Tribe Cantacaderini Stal 

Cantacaderini Drake and Davis 1960, p. 77. 
Type Genus: Cantacader Amyot and Serville. 

Genus ALLOCADER Drake 

Allocader Drake 1950a, pp. 156, 163.— Drake and RuhofT 1960a, p. 26. 
Type Species: Cantacader leai Hacker. 
* Allocader cordatus (Hacker) 

Phatnoma cor data Hacker 1927, p. 19, fig. 9 [Queensland]. 
Cantacader cordatus: Hacker 1928, p. 174. 
Allocader cordata: Drake 1950a, p. 156. 
Type: Sex unknown; National Park, Queensland, Australia; Queensland 
Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
22 



LACEBUGS OF THE WORLD 23 

*Allocader leai (Hacker) 

Cantacader leai Hacker 1928, p. 176, pi. 20, fig. 3 [Tasmania]. 
Allocader leai: Drake 1950a, p. 156. 
Type: Sex unknown; Hobart, Tasmania, Australia; Queensland Mus. 
Distribution: Australia (Tasmania). 
Host Plant: Unrecorded. 

Note: Myrmecophile; adults and nymphs taken in nest of Amblyopone 
australis Erichson [Hacker]. 
Allocader nesiotes Drake and Runoff 

Allocader nesiotes Drake and Ruhoff 1962b, p. 249, pi. 7, fig. 2 [Lord 
Howe]. 
Type: Holotype d\ brachypterous; Lord Howe Island; So. Austr. Mus. 
Distribution: Australia (Lord Howe Island). 
Host Plant: Unrecorded. 

Genus CANTACADER Amyot and Serville 

Cantacader Amyot and Serville 1843, p. 299— Stal 1873, p. 116; 
1874, p. 46.— Lethierry and Severin 1896, p. 4.— Distant 1903b, 
p. 123 .— Horvath 1906a, p. 10.— Oshanin 1908, p. 400; 1912, p. 42 — 
Stichel 1935, p. 348; 1938a, p. 404; 1960a, p. 268; 1960b, p. 390; 
1960c, p. 129.— Drake 1950a, p. 163.— Drake and Ruhoff 1960a, 
p. 26. 
Taphrostethus Fieber 1844, p. 40. — Drake and Ruhoff 1960a, p. 30. 
Type Species: Piesma quadricornis Le Peletier and Serville. 
*Cantacader abdivitus Drake 

Cantacader abdivitus Drake 1950a, p. 161 [Queensland]. 
Type: Holotype 9, macropterous; Redlynch, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
♦Cantacader afzelii Stal Figure 6a, b 

Cantacader afzelii Stal 1873, p. 116 [Sierra Leone]. — Lethierry and 
Severin 1896, p. 4 [Guinea] .—Distant 1902a, p. 238, pi. 15, fig. 3 — 
Schouteden 1923, p. 83 [Congo].— Mancini 1939c, p. 308 [Somalia]; 
1956b, p. 78 [Ethiopia].— Drake 1958a, p. 101 [Kenya; Angola; 
Ivory Coast; Ghana]; 1958b, p. 25.— Drake and Mamet 1956, p. 300 
[Mauritius]. — Mamet 1957, p. 56. 
Type: Holotype o 71 ; Sierra Leone; Stockholm Mus. 
Distribution: Guinea; Sierra Leone; Ivory Coast; Liberia; Ghana; 
Ethiopia; Somalia; Kenya; Congo; Angola; Mascarene Islands (Mauritius). 
Host Plant: Unrecorded. 
Note: Taken at light [Drake 1958a]. 



24 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Cantacader agilis Drake 

Cantacader agilis Drake 1951, p. 166 [Deslac]; 1960, p. 341. 
Type: Holotype cf ; Deslac Island; Hungarian Mus. 
Distribution: New Guinea Territory (Deslac Island; Northeast New 
Guinea). 

Host Plant: Unrecorded. 
*Cantacader agilis var. tricarinatus Drake 

Cantacader agilis var. tricarinata Drake 1951, p. 167 [Deslac]; 1960, p. 342. 
Type: Holotype o 71 , macropterous; Deslac Island; Hungarian Mus. 
Distribution: New Guinea Territory (Deslac Island; Northeast New 
Guinea). 

Host Plant: Unrecorded. 
*Cantacader amydis Drake Figure Ab 

Cantacader amydis Drake 1960, p. 343, fig. 2 [Normanby]. 
Type: Holotype 9, macropterous; Wakaiuna, Normanby; Amer. Mus. 
Distribution: D'Entrecasteaux Islands (Normanby; Fergusson). 
Host Plant: Unrecorded. 
Cantacader angulipennis Horvath 

Cantacader quadricornis (not Le Peletier and Serville): Fieber 1861, p. 118 

[Spain]. 
Cantacader angulipennis Horvath 1906a, p. 12, fig. 3. — Lindberg 1960a, 
p. 13 [Tenerife]. 
Type: Spain; sex and deposition of type unknown. 
Distribution: Spain; Canary Islands (Tenerife). 
Host Plant: Unrecorded. 
Cantacader attenuatus Distant 

Cantacader attenuatus Distant 1902a, p. 238 [Cape Province]. 
Cantacader attenuatis [sic]: Drake 1950a, p. 163. 
Type: Sex unknown; Cape Colony, South Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
jCantacader avitus Drake 

Cantacader avitus Drake 1950a, p. 161, fig. 3 [fossil]. — Drake and Ruhoff 
1960a, p. 11. 
Type: Holotype, sex undeterminable; fossilized in Baltic amber; MCZ. 
Distribution: Fossil. 
*Cantacader basilewskyi Schouteden 

Canthacader [sic] basilewskyi Schouteden 1955b, p. 163 [Ruanda-Urundi]; 

1957c, p. 312. 
Cantacader basilewski [sic]: Drake 1958a, p. 102 [Congo; Angola], 
Type: Sex unknown; Ruanda; Cent. Afr. Mus. 
Distribution: Ruanda-Urundi; Congo; Angola. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 25 

*Cantacader claratis Drake 

Cantacader claratis Drake 1950a, p. 160, fig. 2 [Malaya]. 
Type: Holotype 9, macropterous; Perak, Federation of Malaya; Drake 
Coll. (USNM). 
Distribution: Federation of Malaya. 
Host Plant: Unrecorded. 
*Cantacader diffidentis Drake and Poor 

Cantacader diffidentis Drake and Poor 1936b, p. 141 [India]. 
Type: Holotype o 71 , macropterous; New Forest, Dehra Dun, India; 
Drake Coll. (USNM). 
Distribution: Greater Sunda Islands (Java); India. 
Host Plant: Unrecorded. 
* Cantacader divisus Bergroth 

Cantacader divisus Bergroth 1908, p. 108 [Ethiopia]. — Drake 1955b, p. 
85 [Angola]; 1958b, p. 26 [Kenya; Tanganyika; Congo]. — Mancini 
1956b, p. 78. 
Cantacader dividus [sic]: Drake 1950a, p. 163. 
Type: Ethiopia; sex and deposition of type unknown. 
Distribution: Ethiopia; Kenya; Tanganyika; Congo; Angola. 
Host Plant: Unrecorded. 
*Cantacader formosus Drake 

Cantacader Jormosus Drake 1950a, p. 159 [Taiwan]. 
Type: Holotype o 71 , macropterous; Formosa; Drake Coll. (USNM). 
Distribution: Taiwan. 
Host Plant: Unrecorded. 
Cantacader gerardi Schouteden 

Canthacader [sic] gerardi Schouteden 1955b, p. 162 [Congo]. 
Type: Sex unknown; Mulungu, Nyunzu, Katanga, Belgian Congo; Cent. 
Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 
^Cantacader infuscatus Distant 

Cantacader infuscatus Distant 1903b, p. 124 [Burma]. 
Type: Sex unknown; Rangoon, Burma; British Mus. 
Distribution: Burma; India. 
Host Plant: Unrecorded. 
Cantacader insularis Drake 

Cantacader insularis Drake 1957e, p. 399, fig. 1 [Reunion]. 
Type: Holotype 9; Plaine des Cafres, Reunion; Madagascar Sci. Inst. 
Distribution: Mascarene Islands (Reunion); Malagasy Republic. 
Host Plant: Unrecorded. 



681-552— ,65- 



26 U.S. NATIONAL MUSEUM BULLETIN 243 

*Cantacader japanicus Drake 

Cantacader japanicus Drake 1947d, p. 225, fig. [Japan; Viet-Nam; 

Thailand]; 1947f, fig. 68 (color). 
Cantacader japonicus [sic]: Takeya 1951a, p. 5. 
Type: Holotype o 71 , macropterous; Shimabara Peninsula, Japan; Drake 
Coll. (USNM). 
Distribution: Japan; Viet-Nam; Thailand. 
Host Plant: Unrecorded. 
*Cantacader laratanus Drake 

Cantacader laratanus Drake 1947d, p. 226 [Larat]. 
Type: Holotype d% macropterous; Larat, South Pacific; Drake Coll. 
(USNM). 
Distribution: Tanimbar Islands (Larat). 
Host Plant: Unrecorded. 
Cantacader laticollis Horvath 

Cantacader quadricornis var. staudingeri (not von Baerensprung) : Puton 

1879c, p. 88 [Algeria]. 
Cantacader laticollis Horvath 1906a, p. 11, fig. 1. 
Type: Sex unknown; Bone, Algeria; Paris Mus. 
Distribution: Algeria. 
Host Plant: Unrecorded. 
*Cantacader lethierryi Scott 

Cantacader lethierryi Scott 1874, pp. 291, 443 [Japan].— Uhler 1896, 
p. 265.— Esaki 1932, p. 1634, fig.; 1954, p. 235, fig. 606.— Drake 
1950a, p. 164 [China].— Takeya 1951a, p. 5; 1953d, p. 167. 
Type: Sex unknown; Japan; British Mus. 
Distribution: Japan; China. 
Host Plant: Unrecorded. 
*Cantacader nocturnis Hacker 

Cantacader nocturnis Hacker 1929, p. 324, pi. 32, fig. 1 [Queensland]. — 

Drake 1960, p. 341 [New Guinea]. 
Cantacader nocturnus [sic]: Drake 1950a, p. 164. 
Type: Sex unknown; Brisbane, Queensland, Australia; Queensland Mus. 
Distribution: Australia (Queensland); New Guinea (Northeast). 
Host Plant: Unrecorded. 
Note: Taken at light [Hacker]. 
*Cantacader quadricornis (Le Peletier and Serville) 

Piesma quadricornis Le Peletier and Serville 1828, p. 653. — Brulle 1835, 

p. 343. 
Cantacader quadricornis: Amyot and Serville 1843, p. 299 [Spain]. — 
Puton 1879c, p. 88 [France; Corsica; Greece; Caucasus]. — Bolivar 
and Chicote 1879, p. 166.— Horvath 1906a, p. 12 [Algeria; Mauri- 
tania]. — Oshanin 1908, p. 400 [Crimea; Morocco]. — Moroder Sala 
1920, p. 12.— de Seabra 1931, pp. 410, 444, figs. 482, 483 [Portu- 



LACEBUGS OF THE WORLD 27 

gal].— Vidal 1937, p. 197.— Guide 1938, p. 241, fig.— Drake 1950a, 
p. 164 [Turkey].— Kiritshenko 1951, p. 243, fig. 323.— Mancini 1952, 
p. 12. — Priesner and Alfieri 1953, p. 63 [Egypt]. — Gomez-Menor 
1956a, p. 109, fig. 5 (fig. only); 1956b, p. 81, fig.— Drake and Davis 
1960, figs. 6, 11, 13-15, 19, 20, 24, 28.— Drake and Ruhoff 1960a, 
pi. 2. 
Taphrostethus staudingeri von Baerensprung 1858, p. 205, pi. 2, fig. 10. 
Cantacader staudingeri: Fieber 1861, p. 118. — Ferrari 1878, p. 65. — 

Jakovlev 1880b, p. 99. 
Cantacader staudingeri var. doriae Ferrari 1874, p. 169 [Italy]. 
Cantacader cuadricornis [sic]: Gonzalez 1948, p. 49. 
Cantacader quadricornis quadricornis: Stichel 1960a, p. 268, fig. 32. 
Type: Unknown. 

Distribution: Portugal; Spain; France; Corsica; Italy; Greece; Turkey; 
U.S.S.R. (Crimea; Caucasus); Egypt; Algeria; Morocco; Mauritania. 
Host Plant: Unrecorded. 
Note: Taken at light [Priesner and Alfieri]. 
*Cantacader quadricornis var. nubilis Horvath 

Cantacader quadricornis var. nubilis Horvath 1906a, p. 12, fig. 2 [Cauca- 
sus; Turkey]. — Kiritshenko 1951, p. 243. — Priesner and Alfieri 1953, 
p. 63 [Egypt].— Hoberlandt 1955, p. 86. 
Cantacader quadricornis nubilus [sic]: Stichel 1960a, p. 268. 
Type: Unknown. 

Distribution: U.S.S.R. (Caucasus); Turkey; Egypt. 
Host Plant: Unrecorded. 
Note: Taken at light [Priesner and Alfieri]. 
f Cantacader quinquecarinatus (Germar and Berendt) 

Tingis quinquecarinata Germar and Berendt 1856, p. 23, pi. 3, fig. 19 

[fossil]. 
Eotingis quinquecarinata: Scudder 1890, p. 359; 1891, pp. 406, 449 

[Horizon: Ligurian]. 
Cantacader quinquecarinatus: Drake and Ruhoff 1960a, p. 10. 
Type: Sex undeterminable; fossilized in Prussian amber; deposition of 
type unknown. 

Distribution: Fossil. 
*Cantacader quinquecostatus (Fieber) 

Taphrostethus quinquecostatus Fieber 1844, p. 41, pi. 3, figs. 18-22 
[Ostindien].— Herrich-Schaeffer 1850, p. 150, pi. 311, figs, c-e — 
Walker 1873b, p. 2 [India]. 
Monanthia subovata Motschulsky 1863, p. 91 [Ceylon]. — Kirby 1891, 

p. 109. 
Cantacader subovatus: Stal 1873, p. 116. 

Cantacader quinquecostatus: Stal 1873, p. 117. — Distant 1903b, p. 123, 
fig. 88; 1910a, p. 100.— Kirkaldy 1908c, p. 12.— Horvath 1912c, p. 341 



28 U.S. NATIONAL MUSEUM BULLETIN 243 

[Java; Sumatra]; 1926, p. 327 [Malaya].— Bergroth 1921, p. 103. — 
Drake and Poor 1936b, p. 141; 1937a, p. 1 [Philippines].— Blote 
1945, p. 78.— Drake 1956d, p. 106, fig. 1 [Penang; Palau].— Drake 
and Davis 1960, fig. 25.— Drake 1960, p. 341, fig. 1 [New Guinea]. 
Type: "Ostindien"; sex and deposition of type unknown. 
Distribution: India; Ceylon; Burma; Greater Sunda Islands (Java; 
Sumatra); New Guinea; Federation of Malaya; Penang Island; Philippine 
Islands; Palau Islands. 
Host Plant: Unrecorded. 
Note: Taken at light [Distant 1910a]. 
*Cantacader tener Bergroth Plate 6 

Cantacader tener Bergroth 1894, p. 167 [Malagasy].— Drake 1958c, p. 315. 
Cantacader tenei [sic]: Drake 1950a, p. 164. 
Type: Holotype 6"; Madagascar; deposition unknown. 
Distribution: Malagasy Republic; Kenya. 
Host Plant: Unrecorded. 
*Cantacader tenuipes Stal 

Cantacader tenuipes Stal 1865, p. 26 [Sierra Leone]; 1873, p. 116. — 
Distant 1902a, p. 238, pi. 15, fig. 2.— Schouteden 1923, p. 83 
[Congo].— Mancini 1949a, p. 229 [Somalia].— Drake 1954e, p. 1; 
1957a, p. 415 [Aden]; 1958a, p. 102 [Guinea; Angola]; 1958b, p. 26 
[Kenya; Ghana]. 
Type: Holotype o 71 ; Sierra Leone; Stockholm Mus. 
Distribution: Aden Protectorate; Somalia; Sierra Leone; Guinea; 
Ghana (?); Congo; Angola; Kenya. 
Host Plant: Unrecorded. 
Note: Taken at light [Drake 1958a]. 
Cantacader tenuipes var. furtivus Drake 

Cantacader tenuipes var. infuscatus Schouteden 1916b, p. 290 [Congo]; 

1923, p. 83. 
Cantacader tenuipes var. furtivus Drake 1950a, p. 153 [Republique du 
Congo]. 
Type: Sex unknown; Vankerckhovenville, Belgian Congo; Cent. Afr. 
Mus. 
Distribution: Congo; Republique du Congo. 
Host Plant: Unrecorded. 
*Cantacader uniformis Distant 

Cantacader uniformis Distant 1902b, p. 353 [India]; 1903b, p. 124. — 
Drake 1950a, p. 164 [Burma]. 
Type: Sex unknown; India; British Mus. 
Distribution: India; Burma. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 29 

Cantacader vandenplasi Schouteden 

Cantacader vandenplasi Schouteden 1923, p. 83 [CongoJ. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Genus CERATOCADER Drake 

Ceratocader Drake 1950a, pp. 157, 164.— Drake and Ruhoff 1960a, p. 27. 
Type Species: Cantacader armatus Hacker. 
*Ceratocader armatus (Hacker) Plate 5 

Cantacader armatus Hacker 1928, p. 174, pi. 20, fig. 1 [South Australia]. 
Ceratocader armatus: Drake 1950a, pp. 158, 164. — Drake and Davis 
1960, fig. 30. 
Type: Sex unknown; Murray Bridge, South Australia, Australia; 
Queensland Mus. 
Distribution: Australia (South Australia). 
Host Plant: Unrecorded. 
Ceratocader dentatus (Hacker) 

Cantacader dentatus Hacker 1928, p. 175, pi. 20, fig. 2 [Tasmania]. 
Ceratocader dentatus: Drake 1950a, pp. 158, 164. 
Type: Sex unknown; Burnie, Tasmania, Australia; Queensland Mus. 
Distribution: Australia (Tasmania). 
Host Plant: Unrecorded. 

Genus NECTOCADER Drake 

Neclocader Drake 1928a, p. 41; 1944b, p. 141; 1950a, p. 165.— Monte 
1937d, p. Ill; 1941e, p. 71.— Drake and Ruhoff 1960a, p. 28. 
Type Species: Cantacader gounellei Drake. 
*Nectocader gounellei (Drake) 

Cantacader gounellei Drake 1923b, p. 81, fig. 1 [Brazil]. 
Nectocader gounellei: Drake 1928a, p. 42. — Drake and Hambleton 1934, 
p. 436.— Monte 1937a, p. 30, fig. 1 [Vernonia]; 1937d, p. 112, fig. 
10; 1939b, p. 63. 
Type: Holotype o 71 , macropterous; Novo Friburgo, near Rio de Janeiro, 
Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Vernonia sp. 

Genus TERATOCADER Drake 

Teratocader Drake 1950a, pp. 158, 166.— Drake and Ruhoff 1960a, p. 30. 
Type Species: Cantacader magnificus Drake. 



30 U.S. NATIONAL MUSEUM BULLETIN 243 

*Teratocader magnificus (Drake) 

Cantacader magnified Drake 1923b, p. 83 [Malaya]. 
Teratocader magnificus: Drake 1950a, pp. 158, 166. 
Type: Holotype o 71 , macropterous; Perak, Malay Peninsula; Drake Coll. 
(USNM). 
Distribution: Federation of Malaya. 
Host Plant: Unrecorded. 

Tribe Phatnomini Drake and Davis 

Phatnomini Drake and Davis 1960, p. 78. 
Type Genus: Phatnoma Fieber. 

Genus ALLOEODERES Drake 

Alloeoderes Drake 1961a, p. 115. 
Type Species: Alloeoderes davao Drake. 
Alloeoderes davao Drake Plate 1 

Alloeoderes davao Drake 1961a, p. 116, fig. 43 [Mindanao]. 
Type: Holotype o 71 ; Mt. McKinley, Davao, Mindanao, Philippine 
Islands; Chicago Museum Natural History. 
Distribution: Philippine Islands (Mindanao). 
Host Plant: Unrecorded. 

Genus ANGIOCADER Drake 

Angiocader Drake 1950a, pp. 159, 163.— Drake and Ruhoff 1960a, p. 26. 
Type Species: Phatnoma obesa Distant. 
Angiocader obesus (Distant) 

Phatnoma obesa Distant 1902a, p. 239, pi. 15, fig. 16 [Cape Province]. 

Gonycentrum obesum: Bergroth 1903, p. 297. 

Sinalda obesa: Distant 1904, p. 427. 

Angiocader obesus: Drake 1950a, pp. 159, 163. 
Type: Sex unknown; Cape Colony, South Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

Genus ASTOLPHOS Distant 

Astolphos Distant 1904, p. 428.— Drake 1950a, p. 163.— Drake and 
Ruhoff 1960a, p. 26. 
Type Species: Astolphos capitatus Distant. 
*Astolphos capitatus Distant 

Astolphos capitatus Distant 1904, p. 429, pi. 8, fig. 5 [Cape Province]. 
Type: Sex unknown; Cape Town, South Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 31 

Genus CNEMIANDRUS Distant 

Cnemiandrus Distant 1902a, p. 239.— Drake 1950a, p. 164.— Drake and 
RuhofT 1960a, p. 27. 
Type Species: Cnemiandrus typicus Distant. 
* Cnemiandrus typicus Distant 

Cnemiandrus typicus Distant 1902a, p. 240, pi. 15, fig. 18 [Cape Province]. 
Type: Sex unknown; Cape Colony, South Africa; So. Afr. Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

Genus CYCLOTYNASPIS Montandon 

Cyclotynaspis Montandon 1892, p. 265. — Lethierry and Severin 1896, 
p. 4.— Drake and RuhofT 1960a, p. 27. 
Type Species: Cyclotynaspis acalyptoides Montandon. 
Cyclotynaspis acalyptoides Montandon 

Cyclotynaspis acalyptoides Montandon 1892, p. 265 [Singapore]. — Drake 
1955a, p. 78, fig. 25. 
Type: Holotype 9, brachypterous; Singapore; Bucharest Mus. 
Distribution: Singapore. 
Host Plant: Unrecorded. 
Note: Type specimen redescribed and figured [Drake]. 

Genus CYPEROBIA Bergroth 

Cyperobia Bergroth 1927, p. 673.— Drake 1950a, p. 164.— Drake and 
Runoff 1960a, p. 27. 
Type Species: Cyperobia carectorum Bergroth. 
Cyperobia carectorum Bergroth 

Cyperobia carectorum Bergroth 1927, p. 674 [New Zealand; Cassinia 

leptophylla; sedges]. 
Cyperobia correctorum [sic]: Drake and Davis 1960, fig. 31. 
Type: Holotype 9; Gollan's Valley, Wellington, New Zealand; deposi- 
tion unknown. 
Distribution: New Zealand. 
Host Plants: Cassinia leptophylla; sedges. 

Genus EOCADER Drake and Hambleton 

Eocader Drake and Hambleton 1934, p. 436; 1944b, p. 121.— Monte 
1941e, p. 71; 1942d, p. 104.— Hurd 1946, p. 439.— Drake 1950a, 
p. 164.— Drake and Ruhoff 1960a, p. 27. 
Montea Bruner 1940, p. 246.— Drake and Ruhoff 1960a, p. 28. 
Type Species: Eocader vesrandis Drake and Hambleton. 



32 U.S. NATIONAL MUSEUM BULLETIN 2 43 

*Eocader bouclei (Bruner) 

Montea bouclei Bruner 1940, p. 246, pi. 43 [Cuba; Casuarina equisetifolia]. 
Eocader bouclei: Monte 1942d, p. 104. — Bruner, Scaramuzza and Otero 
1945, p. 33.— Hurd 1946, p. 439 [Casuarina bark]. 
Type: Holotype d", macropterous; Vedado, Havana, Cuba; USNM. 
Distribution: Cuba. 

Host Plants: Casuarina equisetifolia; "Casuarina bark". 
*Eocader vegrandis Drake and Hambleton 

Eocader vegrandis Drake and Hambleton 1934, p. 436, fig. 1 [Brazil; 
Bombax munguba]. — Araujo e Silva 1936, p. 7. — Costa Lima 1936, 
p. 126. 
Type: Holotype 9, brachypterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Bombax munguba. 

Genus GONYCENTRUM Bergroth 

Gonycenirum Bergroth 1898, p. 9.— Distant 1903b, p. 125.— Drake 1950a, 

p. 165.— Drake and Ruhoff 1960a, p. 28. 
Teleia Fieber 1844, p. 55.— Walker 1873b, p. 3.— Lethierry and 

Severin 1896, p. 4.— Drake and Ruhoff 1960a, p. 30. 
Sinalda Distant 1904, p. 426.— Drake and Ruhoff 1960a, p. 30. 
Type Species: Teleia coronata Fieber. 
*Gonycentrum aethiops (Distant) 

Phatnoma aethiops Distant 1902a, p. 238, pi. 15, fig. 12 [Cape Province]. 
Gonycentrum aethiops: Bergroth 1903, p. 297. — Drake 1956f, p. 421. 
Sinalda aethiops: Distant 1904, p. 427. 
Type: Sex unknown; Cape Colony, South Africa; So. Afr. Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
*Gonycentrum afrum Drake and Ruhoff 

Gonycentrum afrum Drake and Ruhoff 1961b, p. 126, fig. 1 [Cape 
Province]. 
Type: Holotype cf, macropterous; Grahamstown, South Africa; Drake 
Coll. (USNM). 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
^Gonycentrum angustatum Drake 

Gonycentrum angustatum Drake 1956c, p. 15 [Tanganyika]. 
Type: Holotype o 71 , macropterous; Arusha, Tanganyika Territory; 
Drake Coll. (USNM). 
Distribution : Tanganyika . 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 33 

Gonycentrum coronatum (Fieber) 

Teleia coronata Fieber 1844, p. 56, pi. 4, figs. 26-32 [Ostindien]. — 
Herrich-Schaeffer 1850, p. 149, pi. 311, figs. 954f-k. — Stal 1873, 
p. 117 [India orientalis]. 
Gonycentrum coronatum: Distant 1903b, p. 125, fig. 89. 
Type: "Ostindien"; sex and deposition of type unknown. 
Distribution: India. 
Host Plant: Unrecorded. 
Gonycentrum elegans (Distant) 

Sinalda elegans Distant 1904, p. 427, pi. 8, fig. 2 [Cape Province]. 
Gonycentrum elegans: Drake 1950a, p. 165. 
Type: Sex unknown; Cape Town, South Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
^Gonycentrum engistum Drake and Ruhoff 

Gonycentrum engistum Drake and Ruhoff 1961b, p. 127, fig. 3* [New 
Guinea]. 
Type: Holotype 9, brachypterous; Mt. Lamington, New Guinea; Drake 
Coll. (USNM). 
Distribution: New Guinea. 
Host Plant: Unrecorded. 

Note: * In Drake and Ruhoff 1961b, the illustration in figure 3 is that of 
engistum, not figure 2 as listed in text and figure citation. 
*Gonycentrum nebulosum (Distant) 

Sinalda nebulosa Distant 1904, p. 428, pi. 8, fig. 4 [Cape Province]. 
Gonycentrum nebulosum: Drake 1950a, p. 165. 
Type: Sex unknown; Grahamstown, Cape Colony, South Africa; 
British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
*Gonycentrum reticulatum (Distant) 

Sinalda reticulata Distant 1904, p. 427, pi. 8, fig. 3 [Cape Province]. 
Gonycentrum reticulatum: Drake 1950a, p. 165. 
Type: Sex unknown; Cape Colony, South Africa; So. Afr. Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
*Gonycentrum socium Drake and Ruhoff 

Gonycentrum socium Drake and Ruhoff 1961b, p. 128, fig. 2* [Tasmania]; 
1962b, p. 249, pi. 7, fig. 1 [South Australia]. 
Type: Holotype cf, brachypertous; Launceston, Tasmania, Australia; 
Drake Coll. (USNM). 



34 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Distribution: Australia (Tasmania; South Australia). 
Host Plant: Unrecorded. 

Note: *In Drake and Runoff 1961b, the illustration in figure 2 is that of 
socium, not figure 3 as listed in the text and figure citation. 
*Gonycentrum testaceum (Distant) 

Phatnoma testacea Distant 1902a, p. 238, pi. 15, fig. 13 [Gape Province]. 

Gonycentrum testaceum: Bergroth 1903, p. 297. 

Sinalda testacea: Distant 1904, p. 427. 
Type: Sex unknown; Cape Colony, South Africa; So. Afr. Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

*Gonycentrum thomasi Drake 

Gonycentrum thomasi Drake 1956c, p. 14 [Kenya]. 
Type: Holotype o 71 , macropterous ; Nairobi, Kenya Colony; Drake Coll. 
(USNM). 
Distribution: Kenya. 
Host Plant: Unrecorded. 
*Gony centrum tindalei (Hacker) 

Phatnoma tindalei Hacker 1928, p. 177, pi. 21, fig. 5 [South Australia; 

moss; lichens]. 
Sinalda tindalei: Hacker 1929, p. 333. 

Gonycentrum tindalei: Drake 1950a, p. 165. — Drake and Ruhoff 1962b, 
p. 249. 
Type: Sex unknown; Mount Lofty Ranges, South Australia, Australia; 
Queensland Mus. 
Distribution: Australia (South Australia). 
Host Plants: Moss; lichens. 

Genus MALALA Distant 

Malala Distant 1910a, p. 101. — Drake 1950a, p. 165. — Drake and 
Ruhoff 1960a, p. 28. 
Type Species: Malala bulliens Distant. 
*Malala bulliens Distant 

Malala bulliens Distant 1910a, p. 101, fig. 48 [Ceylon]. 
Type: Sex unknown; Peradeniya, Ceylon; British Mus. 
Distribution: Ceylon; India. 
Host Plants: Tobacco; "swept from short grass." 

Genus ORANOMA Drake 

Oranoma Drake 1951, p. 165.— Drake and Ruhoff 1960a, p. 29. 
Type Species: Oranoma biroi Drake. 
*Oranoma biroi Drake 

Oranoma biroi Drake 1951, p. 166 [New Guinea]; 1960, p. 344, fig. 4. — 
Drake and Davis 1960, fig. 29. 



LACEBUGS OF THE WORLD 35 

Type: Holotype 9, brachypterous; Sattelberg, Huon Gulf, New Guinea; 
Hungarian Mus. 
Distribution: New Guinea (Northeast). 
Host Plant: Unrecorded. 

Genus PHATNOMA Fieber 

Phatnoma Fieber 1844, p. 57. — Walker 1873b, p. 3. — Lethierry and 
Severin 1896, p. 5.— Champion 1897, p. 2.— Gibson 1919a, pp. 
181-185 (key).— Monte 1941e, p. 72; 1946d, pp. 247-254 (key).— 
Drake 1944b, p. 141; 1950a, p. 165.— Hurd 1946, p. 438.— Drake and 
Ruhoff 1960a, p. 29. 
Type Species: Phatnoma laciniata Fieber. 

* Phatnoma agviates Drake and Ruhoff Figure 5 

Phatnoma agviates Drake and Ruhoff 1961b, p. 130, fig. 5 [Bougainville]. 
Type: Holotype o 71 , brachypterous; Bougainville, Solomon Islands; 
Drake Coll. (USNM). 
Distribution: Solomon Islands (Bougainville). 
Host Plant: Unrecorded. 

* Phatnoma amazonica Drake and Hambleton 

Phatnoma amazonica Drake and Hambleton 1944b, p. 120 [Brazil]. 
Type: Holotype 9, macropterous; Para, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Phatnoma annulipes Champion 

Phatnoma annulipes Champion 1897, p. 4, pi. 1, fig. 2 [Mexico; Guate- 
mala; Panama]. — Gibson 1919a, p. 184. — Drake and Hambleton 
1945, p. 356 [Costa Rica; Vernonia]. — Swezey 1945, p. 371 
[Venezuela].— Hurd 1946, p. 438 [Peru].— Drake 1948c ; p. 21. 
Type: Sex unknown; Volcan de Chiriqui, Panama; British Mus. 
Distribution: Mexico; Honduras; Guatemala; Costa Rica; Panama; 
Venezuela; Peru. 
Host Plant: Vernonia sp. 

Note: Intercepted in orchid packing from Venezuela, at port-of-entry, 
Washington, D.C. [Swezey]. 

* Phatnoma annulipes var. concisa Drake 

Phatnoma annulata [sic] var. concisa Drake 1948c, p. 21 [Venezuela]. 
Type: Holotype cf, macropterous; Caracas, Venezuela; Drake Coll. 
(USNM). 
Distribution: Venezuela. 
Host Plant: Unrecorded. 
*fPhatnoma baltica Drake Plate 3 

Phatnoma baltica Drake 1950a, p. 153, fig. 1 [fossil]. — Drake and 
Ruhoff 1960a, p. 11. 
Type: Holotype 9; fossilized in Baltic amber; Drake Coll. (USNM). 
Distribution: Fossil. 



36 U.S. NATIONAL MUSEUM BULLETIN 243 

*Phatnoma barberi Drake 

Phatnoma barberi Drake 1941a, p. 141 [Colombia]. 
Type: Holotype 9, macropterous; Colombia; USNM. 
Distribution: Colombia. 
Host Plant: Unrecorded. 

Note: Intercepted on Cattleya schroederae at port-of-entry, San Francisco, 
Calif. [Drake]. 
*Phatnoma costalis Distant 

Phatnoma costalis Distant 1909a, p. 113 [Burma]; 1910a, p. 102, fig. 49 — 
Gibson 1919a, p. 184.— Drake and Poor 1936a, p. 439 [Hainan 
Island]. 
Type: Sex unknown; Myitta, Tenasserim, Burma; British Mus. 
Distribution: Burma; China (Hainan Island). 
Host Plant: Unrecorded. 

* Phatnoma coyazana Drake 

Phatnoma coyazana Drake 1948d, p. 15 [Brazil]. 
Type: Holotype 9, macropterous; Jutahy, Goyaz Province, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

* Phatnoma ecuadoris Drake 

Phatnoma ecuadoris Drake 1941a, p. 141 [Ecuador]. 
Type: Holotype o 71 , macropterous; Ecuador; USNM. 
Distribution: Ecuador. 
Host Plant: Unrecorded. 
Note: Intercepted on bananas at port-of-entry, New York City [Drake]. 

* Phatnoma guatemalana Drake 

Phatnoma guatemalana Drake 1948c, p. 20 [Guatemala]. 
Type: Holotype 9, macropterous; Guatemala City, Guatemala; USNM. 
Distribution: Guatemala. 
Host Plant: Unrecorded. 

Note: Intercepted on Epidendrum cochleatum at port-of-entry, San Fran- 
cisco, Calif. [Drake], 

* Phatnoma hacked Drake 

Phatnoma pacifica (not Kirkaldy): Hacker 1928, p. 176, pi. 20, fig. 4 

[Australia] . 
Phatnoma hackeri Drake 1950a, p. 154. 
Type: Holotype 9, brachypterous; Cairns District, Queensland, Aus- 
tralia; Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 37 

Phatnoma hova Schouteden 

Phatnoma hova Schouteden 1957a, p. 82 [Malagasy]. 
Type: Sex unknown; Madagascar; Paris Mus. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 
*Phatnoma jinjana Drake 

Phatnoma jinjana Drake 1956c, p. 13 [Uganda]. 
Type: Holotype 9, macropterous; Jinja, Uganda; Drake Coll. (USNM). 
Distribution: Uganda. 
Host Plant: Unrecorded. 
Phatnoma laciniata Fieber 

Phatnoma laciniata Fieber 1844, p. 57, pi. 4, figs. 33-38 [Ostindien], — 
Herrich-Schaeffer 1850, p. 149, pi. 311, figs. 952a-b.— St&l 1873, 
p. 117 [India orien talis]. —Distant 1903b, p. 126, fig. 90 [Ceylon].— 
Gibson 1919a, p. 183 [East Indies]. 
Type: "Ostindien"; sex and deposition of type unknown. 
Distribution: India; Ceylon; East Indies(?). 
Host Plant: Unrecorded. 
*Phatnoma maculata Monte 

Phatnoma maculata Monte 1946d, p. 252, figs. 1, 8 [Brazil]. — Kormilev 
1955a, p. 63 [Argentina]. 
Type: Holotype 9; Sao Paulo, Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil; Argentina. 
Host Plant: Unrecorded. 
* Phatnoma marmorata Champion 

Phatnoma marmorata Champion 1897, p. 3, pi. 1, figs. 1,1a [Panama]. — 
Gibson 1919a, p. 183.— Drake 1922c, p. 352 [Brazil].— Monte 1941d, 
p. 93 [Costa Rica]; 1941e, p. 72 [Honduras; cacao]. — Drake and 
Hambleton 1944b, p. 120 [Trinidad; pineapple]. 
Phatnoma filetia Gibson 1919a, p. 185. 
Phatnoma spinosa Gibson 1919a, p. 185. 
Type: Sex unknown; David, Chiriqui, Panama; British Mus. 
Distribution: Trinidad; Honduras; Costa Rica; Panama; Brazil; 
Ecuador. 

Host Plants: Cacao; cultivated pineapple. 
Phatnoma maynei Schouteden 

Phatnoma maynei Schouteden 1916b, p. 289 [Congo]. 
Phatnoma magnei [sic]: Drake 1950a, p. 165. 
Type: Sex unknown; Congo da Lemba, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 



38 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Phatnoma ovata Champion 

Phatnoma ovata Champion 1897, p. 4, pi. 1, figs. 3, 3a [Guatemala]. — 
Gibson 1919a, p. 182.— Monte 1943d, p. 263 [Brazil].— Drake 
1944b, p. 141 [Mexico]. 
Type: Sex unknown; San Isidro, Guatemala; British Mus. 
Distribution: Mexico; Guatemala; Brazil. 
Host Plant: Unrecorded. 

* Phatnoma pacifica Kirkaldy Plate 2 

Phatnoma pacifica Kirkaldy 1908b, p. 363 [Fiji]. — Drake and Poor 1943, 
p. 191. 
Type: Holotype 9; Fiji Islands; Hawaii. Sugar Plant. Assn. 
Distribution: Fiji Islands. 
Host Plant: Unrecorded. 

* Phatnoma takasago Takeya 

Phatnoma takasago Takeya 1933, p. 32, text fig. 1; pi. 3, fig. 1 [Taiwan]; 
1951a, p. 6. — Drake and Poor 1939c, p. 203 [India; Lantana]. 
Type: Holotype cf ; Makazayazaya, Heito, Formosa; Kyushu Univ. 
Distribution: Taiwan; India. 
Host Plant: Lantana sp. 
*Phatnoma togularis Drake 

Phatnoma togularis Drake 1950a, p. 154 [India]. 
Type: Holotype 9, macropterous; Dehra Dun, India; Drake Coll. 
(USNM). 
Distribution: India. 
Host Plant: Unrecorded. 
*Phatnoma tonkinana Drake and Maa 

Phatnoma tonkinana Drake and Maa 1955, p. 1 [Viet-Nam]. 
Type: Holotype 9, macropterous; Hoa Binh, Tonkin; Drake Coll. 
(USNM). 
Distribution: Viet-Nam. 
Host Plant: Unrecorded. 
*Phatnoma trinidadana Drake 

Phatnoma trinidadana Drake 1948c, p. 21 [Trinidad]. 
Type: Holotype 9, macropterous; Trinidad, British West Indies; Drake 
Coll. (USNM). 
Distribution: Trinidad. 
Host Plant: Cultivated pineapple. 

* Phatnoma uichancoi Drake 

Phatnoma pacifica (not Kirkaldy): Drake and Poor 1937b, p. 397 

[Philippines] . 
Phatnoma uichancoi Drake 1950a, p. 155; I960, p. 343 [New Guinea]. — 
Drake and Ruhoff 1962b, p. 249 [Papua]. 
Type: Holotype d", macropterous; Los Banos, Philippine Islands; 
Drake Coll. (USNM). 



LACEBUGS OF THE WORLD 39 

Distribution: Philippine Islands (Los Banos; Mindanao); Solomon 
Islands; New Guinea (Papua). 

Host Plant: Unrecorded. 
*Phatnoma varians Drake 

Phatnoma varians Drake 1922c, p. 352 [French Guiana]. — Swezey 1945, 
p. 371 [Colombia]. 
Type: Holotype 9; Env. de St. Georges, Oyapock, French Guiana; 
Paris Mus. 
Distribution: French Guiana; Colombia. 
Host Plant: Unrecorded. 

Note: Intercepted at port-of-entry, California, on Cattleya schroederae 
from Colombia [Swezey]. 
*Phatnoma varians var. unicarinata Drake 

Phatnoma varians var. unicarinata Drake 1922c, p. 353 [French Guiana]. 
Type: Holotype 9, macropterous; Env. de St. Georges, Oyapock, French 
Guiana; Drake Coll. (USNM). 
Distribution: French Guiana. 
Host Plant: Unrecorded. 
*Phatnoma veridica Drake and Maa 

Phatnoma takasago (not Takeya): Drake 1950a, p. 156 [New Guinea]. 

Phatnoma veridica Drake and Maa 1955, p. 2 [Palau; Papua; Glochi- 

dion\— Drake 1956d, p. 107, fig. 2 [Caroline Islands]; 1960, p. 344, 

fig. 3 [Pip turns]. 

Type: Holotype o 71 , macropterous; New Guinea; Drake Coll. (USNM). 

Distribution: New Guinea (Papua, Northeast; Netherlands); Palau 

Islands (Koror; Peleiu; Aurapusheparu; Babelthuap; Ngiramaous; Uleb- 

sehel; Ngarmalk); Caroline Islands. 

Host Plants: Pipturus sp.; taken on bark of Glochidion sp. 
*Phatnoma vernoniae Drake and Hambleton 

Phatnoma veroniae [sic] Drake and Hambleton 1938b, p. 51, pi. 9, fig. a 

[Brazil; Vernonia polyanthes]. 
Phatnoma vernoniae: Monte 1942d, p. 107. 
Type: Holotype cf , macropterous; Campinas, Sao Paulo, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Vernonia polyanthes. 

Genus PLESIONOMA Drake 

Plesionoma Drake 1950a, pp. 157, 166; 1954f, p. 2.— Drake and Ruhoff 
1960a, p. 29. 
Type Species: Phatnoma humcralis Distant. 
*Plesionoma eteosa Drake 

Plesionoma eteosa Drake 1954f, p. 2 [Cape Province]. 



40 U.S. NATIONAL MUSEUM BULLETIN 243 

Type: Holotype cf , brachypterous; Cape Recife, South Africa; Drake 
Coll. (USNM). 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
*Plesionoma humeralis (Distant) 

Phatnoma humeralis Distant 1902a, p. 239, pi. 15, fig. 14 [Cape Prov- 
ince].— Bergroth 1903, p. 297. 
Plesionoma humeralis: Drake 1950a, p. 157. 
Type: Sex unknown; Cape Colony, South Africa; So. Afr. Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
Plesionoma leroyi Schouteden 

Plesionoma leroyi Schouteden 1955a, p. 25 [Ruanda]. 
Type: Sex unknown; Ruanda; Cent. Afr. Mus. 
Distribution: Ruanda-Urundi. 
Host Plant: Unrecorded. 

Genus PSEUDOPHATNOMA Blote 

Pseudophatnoma Blote 1945, p. 78. — Drake 1950a, p. 166. — Drake and 
Ruhoff 1960a, p. 29. 
Type Species: Pseudophatnoma corniculata Blote. 
Pseudophatnoma corniculata Blote 

Pseudophatnoma corniculata Blote 1945, p. 78, fig. 1 [Rhio-Archipelago] . 
Type: Sex unknown; Durian, Rhio-Archipelago; Leiden Mus. 
Distribution: Rhio-Archipelago, Netherlands Indies. 
Host Plant: Unrecorded. 

Genus STENOCADER Drake and Hambleton 

Stenocader Drake and Hambleton 1944b, p. 120.— Drake 1944b, p. 142; 

1950a, p. 166.— Drake and Ruhoff 1960a, p. 30. 
Type Species: Piesma tingidoides Spinola. 
*Stenocader tingidoides (Spinola) 

Piesma tingidoides Spinola 1852a, p. 200 [Chile].— StM 1873, p. 134. 
Cantacader tingidoides: Signoret 1863, p. 575. — Walker 1873b, p. 1. — 

Lethierry and Severin 1896, p. 4. — Reed 1900, p. 176 (p. 86 separate). 
Cantacader ? germainii Signoret 1863, p. 586. — Stal 1873, p. 134. — 

Walker 1873b, p. 1. 
Cantacader germaini [sic]: Lethierry and Severin 1896, p. 25. — Reed 

1900, p. 179 (separate, p. 86). 
Nectocader germaini [sic]: Drake 1928a, p. 41. — Porter 1933, p. 236. 
Nectocader tingitoides [sic]: Drake 1928a, p. 42; 1939c, p. 332.— Porter 

1933, p. 236.— Monte 1937d, p. 114. 
Nectocader tingidoides: Monte 1942d, p. 107. 
Stenocader tingidoides: Drake and Hambleton 1944b, p. 120. 



LACEBUGS OF THE WORLD 41 

Type: Holotype o 71 ; Chile; deposition unknown. 
Distribution: Chile. 
Host Plant: Unrecorded. 

Genus ULMUS Distant 

Ulmus Distant 1904, p. 425.— Drake 1950a, p. 166.— Drake and Ruhoff 
1960a, p. 30. 
Type Species: Ulmus testudineatus Distant. 
*Ulmus engaeus Drake and Ruhoff 

Ulmus engaeus Drake and Ruhoff 1961b, p. 129, fig. 4 [Congo]. 
Type: Holotype o", brachypterous; Kasango, Kivu Province, Congo; 
Drake Coll. (USNM). 
Distribution: Congo. 
Host Plant: Unrecorded. 
*Ulmus testudineatus Distant 

Ulmus testudineatus Distant 1904, p. 426, pi. 8, figs. 1,1a [Transvaal] . — 
Drake 1956f, p. 421. 
Type: Sex unknown; Pretoria District, Transvaal, South Africa; British 
Mus. 
Distribution: South Africa (Transvaal). 
Host Plant: Unrecorded. 

Genus ZETEKELLA Drake 

Zetekella Drake 1944b, pp. 139, 142; 1950a, p. 166.— Hurd 1946, p. 

439.— Drake and Ruhoff 1960a, p. 31. 
Minitingis Barber 1954, p. 7.— Drake and Ruhoff 1960a, p. 28. 
Type Species: ^etekella zeteki Drake. 
*Zetekella minuscula (Barber) 

Minitingis minusculus Barber 1954, p. 7, fig. 1 [So. Bimini]. 
^etekella minuscula: Drake and Ruhoff 1960a, p. 28. 
Type: Holotype cf ; South Bimini Island, Bahamas; Amer. Mus. 
Distribution: Bahama Islands (So. Bimini Island). 
Host Plant: Unrecorded. 
*Zetekella pulla Drake and Plaumann Plate 4 

^etekella pulla Drake and Plaumann 1956, p. 17 [Brazil]. 
Type: Holotype cf, macropterous; Nova Teutonia, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Zetekella zeteki Drake 

^etekella zeteki Drake 1944b, p. 140, fig. 1 [Canal Zone]. 
Type: Holotype 9, brachypterous; Barro Colorado Island, Panama Canal 
Zone; USNM. 
Distribution: Panama (Canal Zone). 
Host Plant: Unrecorded. 

681-552—65 4 



42 U.S. NATIONAL MUSEUM BULLETIN 243 

Subfamily Tinginae Laporte 

Tingidites Laporte 1833, p. 47. 

Agrammidae Douglas and Scott 1865, pp. 24, 242 (as family). 
Tingedidae Douglas and Scott 1865, pp. 24, 243 (as family). 
Serenthiaria Stal 1873, pp. 116, 117; 1874, p. 46.— Distant 1903b, 

p. 126— Horvath 1906a, p. 107.— Oshanin 1908, p. 457; 1912, p. 

46 (as divisions). 
Tingitaria Stal 1873, p. 118; 1874, p. 47.— Distant 1903b, p. 130. — 

Horvath 1906a, p. 13.— Oshanin 1908, p. 401; 1912, p. 42 (as 

divisions) . 
Axiokersosaria Distant 1909a, p. 124 (as division). 
Aidoneusaria Distant 1909a, p. 125 (as division). 
Galeatini Blatchley 1926, p. 451 (as tribe). 
Acalyptini Blatchley 1926, p. 479 (as tribe). 
Physatocheilini Blatchley 1926, p. 483 (as tribe). 
Agramminae China and Miller 1955, p. 261; 1959, p. 9 (as subfamily). 
Agrammatinae Drake and Ruhoff 1960a, p. 23 (as subfamily). 
Tinginae China and Miller 1955, p. 261; 1959, p. 9.— Drake and 

Ruhoff 1960a, p. 31 (as subfamily). — Drake and Davis 1960, pp. 74, 

78 (new concept of subfamily). 
Aidoneusinae Menon and Hakk 1959a, p. 392 (as subfamily). 
Axiokersosinae Menon and Hakk 1959a, p. 392 (as subfamily). 
Type Genus: Tingis Fabricius. 

Tribe Litadeini, new tribe 
Type Genus: Litadea China. 

Genus LITADEA China 

Litadea China 1924, p. 438.— Drake and Ruhoff 1960a, p. 66. 
Type Species: Litadea delicatula China. 

*Litadea delicatula China Plates 7, 8 

Litadea delicatula China 1924, p. 439, figs. 2a, c [Rodriguez]. — Drake 
and Davis 1960, figs. 39, 40. 
Type: Sex unknown; Rodriguez Island; British Mus. 
Distribution: Mascarene Islands (Rodriguez). 
Host Plant: Unrecorded. 

Tribe Tingini Laporte 
Type Genus: Tingis Laporte. 



LACEBUGS OF THE WORLD 43 

Genus ABDASTARTUS Distant 

Abdastartus Distant 1910a, p. 103.— Bergroth 1911, p. 185.— Drake 
1956b, p. 110.— Drake and Ruhoff 1960a, p. 31. 
Type Species: Abdastartus tyrianus Distant =Monanthia atra Motschulsky. 

* Abdastartus atrus (Motschulsky) 

Monanthia atra Motschulsky 1863, p. 91 [Ceylon].— StM 1873, p. 134 — 

Kirby 1891, p. 109.— Lethierry and Severin 1896, p. 25. 
Teleonemia atra: Distant 1902b, p. 356; 1903b, p. 143.— Bergroth 1921, 

p. 104. 
Abdastartus tyrianus Distant 1910a, p. 103, fig. 50 [India]. — Drake 

1930c, p. 15. 
Abdastartus atrus: Drake 1956b, p. 110. 
Type: Sex unknown; Mount Patannas, Ceylon; Moscow Univ. 
Distribution: Ceylon; India. 
Host Plant: Unrecorded. 

* Abdastartus longulus Drake 

Abdastartus longulus Drake 1953b, p. 95 [India]. 
Type: Holotype o 71 , macropterous; Dehra Dun, India; Drake Coll. 
(USNM). 
Distribution: India. 
Host Plant: Unrecorded. 

♦Abdastartus sacchari Drake 

Abdastartus tyrianus (not Distant): Drake 1927d, p. 307 [Java; Taiwan; 

sugar]. 
Abdastartus sacchari Drake 1930c, p. 15 [Sumatra]. — Box 1953, p. 37. 
Type: Holotype d\ macropterous; Randoeblatoeng, Java; Drake Coll. 
(USNM). 
Distribution: Greater Sunda Islands (Java; Sumatra); Taiwan. 
Host Plant: Sugarcane. 

Genus ACALYPTA Westwood 

Acalypta Westwood 1840b, p. 121.— Stal 1873, pp. 118, 122; 1874, 
p." 51. —Saunders 1875, pp. 245, 251— Horvath 1906a, pp. 13,24 
(key); 1906e, p. 498.— Muzik 1907, pp. 50, 52 (key ) .— Oshanin 
1908, p. 406; 1912, p. 42.— Banks 1910, p. 55.— Jensen-Haarup 1912, 
p. 150 (key).— Osborn and Drake 1916a, p. 220 (key).— Van Duzee 
1916, p. 25; 1917b, p. 211.— Lindberg 1919, p. 42.— Torre-Bueno 
1924, p. 93.— Stichel 1926, pp. 103, 106 (key); 1935, p. 348; 1938a, 
p. 404; 1960a, p. 274; 1960b, p. 391; 1960c, p. 130.— Drake 1928c, 
pp. 1-9.— Borner 1935, pp. 73-75 (key) .— Scholte 1935, p. 43 (key).— 
Guide 1938, pp. 242, 249 (key).— Hoberlandt 1942, p. 124.— China 
1943, p. 245.— Blote 1945, p. 80.— Hurd 1946, p. 462.— Bailey 1951, 



44 U.S. NATIONAL MUSEUM BULLETIN 243 

p. 32.— Kiritshenko 1951, pp. 240, 244 (key).— Singer 1952, p. 48 — 

Drake and Ruhoffl959, p. 138; 1960a, p. 31. — Southwood and Leston 

1959, p. 141. 
Orthosteira Fieber 1844, p. 46.— Sahlberg 1848, p. 128.— China 1941, 

p. 130.— Drake and Ruhoff 1960a, p. 73. 
Orthostira [sic]: Fieber 1861, p. 130. — Douglas and Scott 1865, p. 260. — 

Thomson 1871, p. 398.— Ferrari 1878, p. 82 (key) .— Reuter 1882b, 

p. 114 (key). — Saunders 1892, p. 125. — Lethierry and Severin 1896, 

p. 6. 
Fenestrella Osborn and Drake 1916a, p. 222. — Parshley 1917a, p. 14. — 

Drake and Ruhoff 1960a, p. 57. 
Drakella Bergroth 1922, p. 152.— Blatchley 1926, p. 481.— Drake and 

Ruhoff 1960a, p. 54. 
Type Species: Tingis carinata Panzer. 

*Acalypta acutangula (Jakovlev) 

Orthostira acutangula Jakovlev 1880a, p. 127 [Russia]; 1893, pp. 293, 302. 
Acalypta acutangula: Horvath 1901, p. 475 [Turkey]; 1906a, p. 31 
[Siberia].— Kiritshenko 1951, p. 244.— Hoberlandt 1955, p. 86. 
Type: Holotype 9, macropterous; Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Siberia); Turkey. 
Host Plant: Unrecorded. 

*Acalypta barberi Drake 

Acalypta barberi Drake 1934a, p. 196 [N.Y.].— Drake and Ruhoff 1959, 

p. 137 [N.B.; moss; hops]. 
Acalypta mera Drake 1941a, p. 142 [B.C.; Oreg.]. 
Type: Holotype 9, brachypterous; Merrifield, New York, U.S.; USNM. 
Distribution: U.S. (N.Y.; Oreg.); Canada (B.C.; N.B.). 
Host Plants: Moss; hops. 

* Acalypta brunnea (Germar) 

Tingis brunnea Germar 1836, fasc. 18, tab. 23 [Austria]. 

Monanthia brunnea: Herrich-Schaeffer 1837, p. 25, pi. 118, fig. 374. 

Orthosteira brunnea: Fieber 1844, p. 48, pi. 3, figs. 43-45. — Scholz 1847, 
p. 118 [Germany].— Herrich-Schaeffer 1850, p. 158. 

Orthostira concinna Douglas and Scott 1863, p. 143, fig. 5 [England]; 
1865, p. 260 [moss on oak].— Fieber 1864, p. 211. 

Orthostira brunnea: Ferrari 1878, p. 82. — Puton 1879c, p. 95 [France]. — 
Saunders 1892, p. 126, pi. 11, fig. 7.— Hiieber 1893, p. 305. 

Acalypta brunnea: Stil 1874, p. 51. — Saunders 1875, p. 251 [moss]. — 
Horvath 1906a, p. 28 [Belgium; Switzerland].— Muzik 1907, p. 53 
[Czechoslovakia].— Butler 1923, p. 201 [Scotland; Ireland].— Stichel 
1926, p. 106; 1960a, p. 276 [Sweden].— Scholte 1935, p. 44 [Nether- 
lands].— Hoberlandt 1943b, pp. 120, 121, 122; 1944, pp. 33, 35. 
Type: Austria; sex and deposition of type unknown. 



LACEBUGS OF THE WORLD 45 

Distribution: Ireland; Scotland; England; Netherlands; Belgium; 
France; Switzerland; Austria; Czechoslovakia; Germany; Sweden. 
Host Plants: Moss; moss on oak. 
Note: Life history study [Butler]. 
*Acalypta carinata (Panzer) 

Tingis carinata Panzer 1806, heft 99, tab. 20 [Germany].— Le Peletier 
and Serville 1828, p. 653.— Curtis 1839, tab. 741 [England].— 
Fieber 1844, p. 52, pi. 4, fig. 10. 

Tingis cassidae Fallen 1807, p. 37 [Sweden]; 1829, p. 146— Herrich- 
Schaeffer 1835, p. 59.— Zetterstedt 1840, p. 269 [Lapland]. 

Monanthia carinata: Burmeister 1835, p. 262. 

Tingis cervina Germar 1836, fasc. 18, tab. 22. 

Monanthia cervina: Herrich-Schaeffer 1837, p. 26, pi. 118, fig. 375; pi. 
129, fig. G— Vollenhoven 1878, p. 279, pi. 9, figs. 11, 11a 
[Netherlands]. 

? Monanthia fr acta Herrich-Schaeffer 1838, pp. 53, 63. 

? Monanthia acuminata Herrich-Schaeffer 1838, p. 63. 

Acalypta carinata: Westwood 1840a, p. 477; 1840b, p. 121. — Horvath 
1906a, p. 29 [Rumania].— Oshanin 1908, p. 407 [Belgium].— Jensen- 
Haarup 1912, p. 152 [Denmark].— Sahlberg 1920, p. 78.— Stichel 
1926, p. 106; 1938a, p. 404 [Poland]; 1960a, p. 278, fig. 55 [Norway].— 
Drake 1928c, p. 1.— Lindberg 1932b, p. 211.— Scholte 1935, p. 44, 
fig. 8.— Hoberlandt 1943b, pp. 121, 122 [Czechoslovakia].— Gon- 
zalez 1948, p. 49 [Spain].— Kiritshenko 1951, p. 244.— Bator 1953, 
p. 324, pi. 2, fig. 2.— Southwood and Leston 1959, p. 142, pi. 21, 
fig. 2, text figs. 43, 48.— Stusak 1961, p. 80, fig. 20. 

Orthosteira cervina: Fieber 1844, p. 48, pi. 4, figs. 1-3 [Austria]. — Scholz 
1847, p. 118.— Sahlberg 1848, p. 129 [Finland].— Herrich-Schaeffer 
1850, p. 159.— Mayr 1858, p. 569.— China 1941, p. 130. 

Orthosteira cassidea: Mayr 1858, p. 569 [moss]. 

Monanthia {Orthosteira) cervina: Flor 1860, p. 341 [Latvia; Estonia]. 

Orthostira cervina: Fieber 1861, p. 130; 1864, p. 212. — Douglas and Scott 
1865, p. 262.— Frey-Gessner 1865, p. 233 [Switzerland].— Horvath 
1874b, p. 432 [Hungary].— Jakovlev 1876b, p. 70 [Russia]; 1893, 
p. 301.— Reiber and Puton 1876, p. 68.— Puton 1879c, p. 96 
[France].— Reuter 1882b, p. 114.— Dubois 1888, p. 120.— 
d' Antessanty 1890,'p. 31.— Saunders 1892, p. 126, pi. 11, fig. 8 — 
Hiieberl893, p. 305. 

Orthostira cassidea: Frey-Gessner 1865, p. 233. — Garbiglietti 1869, p. 275 
[Italy].— Thomson 1871, p. 399. 

Acalypta cassidea: Stal 1874, p. 52. 

Acalypta cervina: Reuter 1874, p. 563. — Saunders 1875, p. 252. — 
Reichensperger 1920, p. 61.— Butler 1923, p. 202, fig. [Wales; 
Scotland; Ireland]. — Blote 1945, p. 80 [Yugoslavia; Algeria]. 



46 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Type: Mannheim, Germany; sex and deposition of type unknown. 

Distribution: England; Scotland; Wales; Ireland; Netherlands; Bel- 
gium; France; Spain; Italy; Switzerland; Czechoslovakia; Austria; Hun- 
gary; Yugoslavia; Rumania; Germany; Denmark; Poland; U.S.S.R. 
(Latvia; Estonia); Finland; Sweden; Norway; (Lapland); Algeria. 

Host Plant: Moss. 

Note: Nymphs [Buder]; eggs [Stusak]. 
Acalypta carinata var. angustula Horvath 

Acalypta cervina var. angustula Horvath 1898a, p. 69 [Hungary; France]. 
Acalypta carinata var. angustula: Horvath 1906a, p. 30 [Germany; Aus- 
tria; Rumania; Finland; Estonia; Latvia; Lapland]. — Sahlberg 1920, 
p. 79.— Scholte 1935, p. 44 [Netherlands].— Stichel 1960a, p. 278. 
Type: Unknown. 

Distribution: Netherlands; France; Germany; Austria; Hungary; 
Rumania; U.S.S.R. (Estonia; Latvia); Finland; (Lapland). 

Host Plant: Unrecorded. 
Acalypta carpathica Horvath 

Acalypta carpathica Horvath 1905b, p. 561 [Rumania]; 1906a, p. 30. — 
Stichel 1960a, p. 278. 
Type: Sex unknown; Sinaia, Rumania; Hungarian Mus. 
Distribution: Rumania. 
Host Plant: Unrecorded. 
*Acalypta cooleyi Drake 

Acalypta cooleyi Drake 1917a, p. 213 [Mont.]; 1928c, p. 9. 
Type: Holotype 9, macropterous; Bozeman, Montana, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Mont.; Oreg.; Calif.). 
Host Plant: Unrecorded. 
*Acalypta duryi Drake 

Fenestrella ovata Osborn and Drake 1916a, p. 223, fig. 3 [Ohio]; 1917a, 

p. 155, pi. 8, fig. d. 
Drakella [ovata]: Bergroth 1922, p. 152. 
Drakella ovata: Blatchley 1926, p. 482, fig. 112. 
Acalypta ovata: Drake 1928c, p. 3, fig. la [moss]. 
Acalypta duryi Drake 1930d, p. 268. 
Type: Holotype 9, brachypterous; Cincinnati, Ohio, U.S.; Ohio State 
Univ. 
Distribution: U.S. (Ohio; N.C.; Tenn.). 
Host Plant: Moss. 
Acalypta elegans Horvath 

Acalypta elegans Horvath 1906a, p. 32 [Siberia]. 
Type: Sex unknown; Siberia, Russia; Helsin. Mus. 
Distribution: U.S.S.R. (Siberia). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 47 

Acalypta elinoides (Jakovlev) 

Orthostira elinoides Jakovlev 1893, pp. 294, 302 [Russia]. 

Acalypta elinoides: Horvath 1906a, p. 31 [Siberia]. 
Type: Sex and locality unknown; Leningrad Inst. 
Distribution: U.S.S.R. (Siberia). 
Host Plant: Unrecorded. 

Acalypta finitima (Puton) 

Orthostira finitima Puton 1884, p. 313 [France]. 
Acalypta finitima: Horvath 1906a, p. 29. 
Type: Sex unknown; St. Martin-Lantosque, France; Paris Mus. 
Distribution: France. 
Host Plant: Unrecorded. 
* Acalypta gracilis (Fieber) 

Orthosteira gracilis Fieber 1844, p. 54, pi. 4, figs. 19-21 [Czechoslovakia; 
Thymus serpy I his]. — Scholz 1847, p. 118 [Germany]. — Herrich-Schaef- 
fer 1850, p. 159. — Mayr 1858, p. 569 [Austria; Echium vulgare]. 
Monanthia (Orthosteira) parvula (not Fallen): Flor 1860, pp. 333, 335 

[Estonia; Latvia; Sweden]. 
Orthostira gracilis: Fieber 1861, p. 131 [Thymus]. — Horvath 1874b, 
p. 432 [Hungary].— Reiber and Puton 1876, p. 68.— Ferrari 1878, 
pp. 65, 83.— Puton 1879c, p. 99 [France].— Reuter 1882b, p. 116 
[Finland; moss].— Jakovlev 1893, p. 293 [Russia].— Hueber 1893, 
p. 310. 
Orthostira biseriata Thomson 1871, p. 402. 
Orthostira reticosta Thomson 1871, p. 402. 

Orthostira propinqua Ferrari 1874, p. 171 [Italy; Plantago cynops]. 
Acalypta gracilis: Puton 1874a, p. 227.— Reuter 1874, p. 564; 1880b, 
p. 165. — Horvath 1906a, p. 33 [Denmark; Rumania; Siberia]. — 
Jensen-Haarup 1912, p. 152. — Schumacher 1914, p. 257 [Artemisia; 
Calluna].— Sahlberg 1920, p. 80.— Stichel 1926, p. 107; 1938a, p. 404 
[Poland]; 1960a, p. 281 [Erodium cicutarium; Norway; Switzerland]. — 
Scholte 1935, p. 46, figs. 11, 12 [Netherlands].— Borner 1935, p. 75, 
figs. 113, 119a. — Guide 1938, p. 258, fig. [Artemisia campestris]. — 
Hoberlandt 1943b, p. 122.— Kiritshenko 1951, p. 244. 
Orthostira parvula var. biseriata: Ferrari 1878, p. 83. 
Orthostira parvula var. reticosta: Ferrari 1878, p. 83. 
Orthostira parvula var. propinqua: Ferrari 1878, p. 83. 
Type: Sex unknown; Prague, Czechoslovakia; Vienna Mus. 
Distribution: Netherlands; France; Switzerland; Germany; Czecho- 
slovakia; Italy; Austria; Hungary; Rumania; Poland; Denmark; Norway; 
Sweden; Finland; U.S.S.R. (Latvia; Estonia; Siberia). 

Host Plants: Artemisia campestris; Artemisia sp.; Calluna sp.; Echium 
vulgare; Erodium cicutarium; Plantago cynops; Thymus serpyllus; Thymus sp. ; moss. 



48 U.S. NATIONAL MUSEUM BULLETIN 24 3 

* Acalypta hellenica Reuter 

Acalypta hellenica Reuter 1888, p. 224 [Greece]; 1891a, p. 25— Horvath 
1889, p. 329 [Russia]; 1906a, p. 33 [France; Corsica; Italy; Yugo- 
slavia]; 1911b, p. 584 [Sicily].— Oshanin 1912, p. 42 [Austria].— 
Lindberg 1936b, p. 29 [Canary Islands].— Stichel 1938a, p. 404 
[Turkey]; 1960a, p. 281 [Sardinia].— Kiritshenko 1951, p. 245.— 
Mancini 1953d, p. 15. — Gomez-Menor 1955b, p. 249 [Spain; 
Quercus]. 
Type: Sex unknown; Greece; Helsin. Mus. 

Distribution: France; Corsica; Spain; Sardinia; Italy; Sicily; Austria; 
Yugoslavia; Greece; Turkey; U.S.S.R.; Canary Islands. 

Host Plant: Quercus sp. 
Acalypta heteropepla Horvath 

Acalypta heteropepla Horvath 1907, p. 303 [Algeria]. 
Type: Holotype 9; Mons Babor, Algeria; Oxford Mus. 
Distribution : Algeria. 
Host Plant: Unrecorded. 
Acalypta hoberlandti Roubal 

Acalypta hoberlandti Roubal 1958, p. 54, fig. 1 [France].— Stichel 1960a, 
p. 274, fig. 42. 
Type: Sex unknown; France; Roubal Coll. 
Distribution: France. 
Host Plant: Unrecorded. 
*Acalypta lillianis Torre-Bueno 

Acalypta lillianus Torre-Bueno 1916, p. 39 [N.Y.; Maine; Mich.]. — 
Osborn and Drake 1916a, p. 221, figs. 1, 2 [Ohio; N.H.;Ont.].— Van 
Duzee 1917b, p. 212 [N.J.].— Parshley 1917a, p. 14; 1917b, p. 53; 
1923b, p. 698.— Barber 1922a, p. 17 [moss].— McAtee 1923, p. 145 
[D.C.]. — Blatchley 1926, p. 481, fig. Ill [Ind.; Climacium american- 
um\— Drake 1928b, p. 100; 1928c, p. 6 [Iowa; 111.; Pa.; Que.].— 
Froeschner 1944, p. 669.— Hurd 1946, p. 463 [N. Dak.].— Bailey 
1951, p. 32 [Polytrichurn].— Lindberg 1958b, p. 14 [Nfld.; Dryas; 
Oxytropis].—Byers 1959, p. 191. 
Acalypta ovata Osborn and Drake 1916b, p. 9, fig. 1. — Drake 1932, 

p. 100 [Tenn.]. 
Acalypta grisea Heidemann 1917, p. 218, pi. 17, fig. 2 [Mass.; N.C.; 

Md.].— McAtee 1917b, p. 78. 
Acalypta modesta Parshley 1921, p. 16 [B.C.].— Downes 1925, p. 14; 
1927a, p. 10.— Drake 1928c, p. 7. 
Type: Lectotype 9; White Plains, New York, U.S.; Snow Mus. 
Distribution: U.S. (Maine; N.H.; Vt.; Mass.; Conn.; R.I.; N.Y.; N.J.; 
Pa.; Md.; D.C; Va.; N.C.; Tenn.; Ohio; 111.; Ind.; Mich.; Wis.; Nebr.; 
Iowa.; Minn.; N. Dak.; Idaho; Alaska); Canada (Nfld.; Que.; Ont.; 
B.C.). 



LACEBUGS OF THE WORLD 49 

Host Plants: Climacium americanam; Dryas sp.; Oxytropis sp.; Polytrichum 
sp.; moss. 

Note: Lectotype designated [Byers]. 
* Acalypta marginata (Wolff) 

Acanthia marginata Wolff 1804, p. 131, pi. 13, fig. 126 [Hungary]. — 

Fieber 1844, p. 50, pi. 4, fig. 8. 
Tingis pusilla Herrich-Schaeffer 1835, p. 59. 
Monanthia pusilla: Herrich-Schaeffer 1837, p. 24, pi. 118, fig. 373; 

1838, p. 64, pi. 129, fig. F. 
Orthosteira macrophthalma Fieber 1844, pp. 49, 110, pi. 4, figs. 4-7 
[Sweden; Germany; Czechoslovakia; Austria].— Sahlberg 1848, 
p. 129 [Finland].— Herrich-Schaeffer 1850, p. 159. 
Orthosteira pusilla: Fieber 1844, p. 51, pi. 4, fig. 9 [moss]. 
Monanthia (Orthosteira) cinerea (not Fieber): Flor 1860, pp. 333, 337 

[Latvia; Estonia]. 
Orthostira pusilla: Fieber 1861, p. 131. — Garbiglietti 1869, p. 275 

[Italy].— Ferrari 1878, p. 83. 
Orthostira cylindricornis Thomson 1871 p. 401. 
Acalypta macrophthalma: Reuter 1874, p. 564.— Saunders 1875, p. 253. — 

Buder 1923, p. 203. 
Orthostira macrophthalma: Reiber and Puton 1878, p. 68. — Puton 1879c, 
p. 98 [France].— Chicote 1880, p. 189 [Spain].— Reuter 1882b, 
p. 116 [Lapland].— Saunders 1892, p. 128.— Hueber 1893, p. 309 
[Switzerland; Thymus; Artemisia campestris]. — Jakovlev 1893, p. 293 
[Russia]. 
Orthostira acutispinis Reuter 1886, p. 233. 

Acalypta marginata: Horvath 1906a, pp. 26, 31, pi. 1, fig. 5 [Yugo- 
slavia; Bulgaria; Armenia]. — Oshanin 1908, p. 410 [Caucasus]. — 
Sahlberg 1920, p. 80.— Stichel 1926, p. 107, fig. 296; 1938a, p. 
404 [Norway]; 1938b, p. 454; 1960a, p. 280, fig. 60 [Portugal; Calluna 
vulgaris].— Scholte 1935, p. 46, fig. 9 [Netherlands].— Poisson 1938, 
p. 589 [Erodium].— China 1943, p. 246.— Kiritshenko 1951, p. 244.— 
Stehlik 1952, p. 205.— Bator 1953, p. 324, pi. 2, fig. 5.— Stusak 
1957b, p. 135, fig. 2; 1961a, p. 79, figs. 4a, b, 7; pi. 1, fig. c— Cobben 
1958b, p. 16 [Hieracium pilosella]. 
Type: Hungary; sex and deposition of type unknown. 
Distribution: Netherlands; France; Spain; Italy; Portugal; Switzerland; 
Czechoslovakia; Germany; Austria; Hungary; Bulgaria; Yugoslavia; 
Poland; Norway; Sweden; Finland; (Lapland); U.S.S.R. (Latvia; Estonia; 
Armenia; Siberia; Caucasus). 

Host Plants: Artemisia campestris; Calluna vulgaris; Erodium sp.; Hieracium 
pilosella; Thymus sp.; moss. 

Note: Records from England incorrect [China]. Associated with ants 
[Stichel I960]. 



50 U.S. NATIONAL MUSEUM BULLETIN 24 3 

* Acalypta mniophila Drake and Ruhoff 

Acalypta mniophila Drake and Ruhoff 1959, p. 136, fig. 1 [Mexico]. 
Type: Holotype 9, brachypterous; Mexico; USNM. 
Distribution: Mexico. 
Host Plant: Moss. 
Acalypta montana Hoberlandt 

Acalypta montana Hoberlandt 1944, p. 34, fig. [Czechoslovakia]. — 
Stichel 1960a, p. 276, fig. 49. 
Type: Holotype cf; Savinske Alps, Montes Alpinum, Czechoslovakia; 
Prague Mus. 

Distribution : Czechoslovakia. 
Host Plant: Unrecorded. 

* Acalypta musci (Schrank) 

Cimex musci Schrank 1781, p. 265 [Austria; moss]. 
Orthosteira cassidea (not Fallen): Fieber 1844, p. 47, pi. 3, figs. 39-42 
[Germany; Czechoslovakia; Italy]. — Herrich-Schaeffer 1850, p. 159. 
Campylosteira {Orthosteira) cassidea: Kolenati 1856, p. 430 [Caucasus]. 
Orthostira cassidea: Fieber 1861, p. 130 [France]. 

Orthostira musci: Reiber and Puton 1876, p. 68. — Ferrari 1878, p. 82. — 
Puton 1879c, p. 94 [Hungary] .— Chicote 1880, p. 189 [Spain].— 
Jakovlev 1880b, p. 100 [Russia].— Hiieber 1893, p. 303 [Switzerland]. 
Acalypta musci: Horvath 1906a, p. 28 [Yugoslavia; Rumania; Bulgaria; 
Belgium].— Muzik 1907, p. 53, fig. 3.— Stichel 1926, p. 106; 1960a, 
p. 275, fig. 46 [Andorra; Poland; Cor iolus versicolor; Trametes gibbosa; 
Polyporus unicolor]. — Scholte 1935, p. 44, fig. 7 [Netherlands]. — 
Hoberlandt 1943b, p. 122.— Kiritshenko 1951, p. 244, fig. 329 — 
Bator 1953, p. 324, pi. 2, fig. 1.— Mancini 1953c, p. 22; 1954b, 
p. 11.— Stusak 1957b, pp. 134, 140, figs. 1, 13b, 15c; 1958, p. 365, 
figs. 5-7. 
Type: Austria; sex and deposition of type unknown. 
Distribution: Andorra; Spain; France; Belgium; Netherlands; Ger- 
many; Czechoslovakia; Switzerland; Austria; Hungary; Italy; Yugoslavia; 
Rumania; Poland; U.S.S.R. (Caucasus); Sweden. 
Host Plants: Coriolus versicolor; Polyporus unicolor; Trametes gibbosa; moss. 
Note: Eggs [Stusak]. 

* Acalypta musci var. ditata (Puton) 

Acalypta musci (not Schrank): Stal 1874, p. 51. 

Orthostira musci var. ditata Puton 1879b, p. 297; 1879c, p. 94 [France; 
Austria; moss]. — Chicote 1880, p. 189 [Spain]. 

Acalypta musci var. ditata: Horvath 1906a, p. 28 [Switzerland; Italy; 
Germany; Hungary; Yugoslavia; Bulgaria; Rumania]. — Mu ik 
1907, p. 53 [Czechoslovakia].— Scholte 1935, p. 44 [Netherlands].— 
Blote 1945, p. 81 [Tunisia].— Bator 1953, p. 324.— Mancini 1954b, 
p. 11. 



LACEBUGS OF THE WORLD 51 

Acalypta musci var. dilatata [sic]: Gomez-Menor 1955b, p. 249 [Andorra]. 
Type: Unknown. 

Distribution: Spain; Andorra; France; Belgium; Netherlands; Ger- 
many; Czechoslovakia; Austria; Hungary; Switzerland; Italy; Yugoslavia; 
Bulgaria; Rumania; U.S.S.R. (Caucasus); Tunisia. 

Host Plant: Moss. 
Acalypta musci var. triseriata Stichel 

Acalypta musci var. triseriata Stichel 1960a, p. 275. 
Type: Unknown. 
Distribution: Unknown. 
Host Plant: Unknown. 
* Acalypta nigrina (Fallen) 

Tingis nigrina Fallen 1807, p. 37 [Finland]; 1829, p. 145.— Herrich- 
Schaeffer 1830, heft 118, tab. 16 [Germany].— Zetterstedt 1840, 
p. 269 [Lapland]. 
Tingis pusilla Fallen 1807, p. 38 [Sweden]; 1829, p. 146.— Curtis 

1839, tab. 741 [England]. 
Tingis nigrinus: Herrich-Schaeffer 1835, p. 58. 
Monanthia pusilla: Burmeister 1835, p. 262. 

Monanthia nigrina: Herrich-Schaeffer 1838, pp. 52, 62, pi. 125, fig. G. 
Monanthia (Phyllontocheila) tiigrina: Fieber 1844, p. 63, pi. 5, figs. 

23-24.— Herrich-Schaeffer 1850, p. 156. 
Orthosteira cinerea Fieber 1844, pp. 52, 110, pi. 4, figs. 11-14 [Czecho- 
slovakia; moss]. — Sahlberg 1848, p. 130. — Herrich-Schaeffer 1850, 
p. 159.— Mayr 1858, p. 569 [Austria]. 
Monanthia {Orthosteira) nigrina: Flor 1860, p. 334 [Estonia; Latvia]. 
Orthostira nigrina: Fieber 1861, p. 131. — Frey-Gessner 1865, p. 233 
[Switzerland]. — Jako vie v 1871, p. 26 [Russia]. — Thomson 1871, 
p. 400— Horvath 1874b, p. 432 [Hungary].— Ferrari 1874, p. 172 
[Italy; Spartiumjunceum]; 1878, p. 83. — Reiber and Puton 1876, p. 68. — 
Puton 1879c, p. 98 [France].— Saunders 1892, p. 127, pi. 11, fig. 9 — 
Hiieber 1893, p. 307. 
Orthostira pusilla: Frey-Gessner 1865, p. 233. — Gredler 1870, p. 76. 
Acalypta nigrina: Stal 1874, p. 52. — Reuter 1874, p. 564. — Saunders 
1875, p. 252.— Horvath 1906a, p. 31 [Thymus serpyllus]. —Reichcn- 
sperger 1920, p. 61.— Sahlberg 1920, p. 79.— Butler 1923, p. 203 
[Scotland]. — Stichel 1926, p. 107 [Hieracium murorum]; 1938a, 
p. 404 [Norway; Poland]; 1960a, p. 279 [Iceland; Denmark; Calluna 
vulgaris].— Poisson 1938, p. 589 [Erodium] .— Blote 1945, p. 81 [Tuni- 
sia] .— Kiritshenko 1951, p. 244.— Stehlik 1952, p. 205.— Bator 1953, 
p. 324, pi. 2, fig. 4.— Southwood and Leston 1959, p. 144, pi. 21, 
fig. 3, text fig. 49. 
Monanthia carinata (not Panzer): Vollenhoven 1878, p. 280, pi. 10, 
figs. 1, la [Netherlands]. 



52 U.S. NATIONAL MUSEUM BULLETIN 243 

Type: Sex unknown; Finland; Lund Zool. Inst. 

Distribution: England; Scotland; Netherlands; France; Germany; 
Italy; Switzerland; Czechoslovakia; Austria; Hungaiy; Poland; Denmark; 
Norway; Sweden; Finland; (Lapland); Iceland; U.S.S.R. (Estonia; 
Latvia); Tunisia. 

Host Plants: Calluna vulgaris; Er odium sp.; Hieracium murorum; Spartium 
junceum; Thymus serpyllus; moss. 

* Acalypta nigrinervis Stal 

Acalypta nigrinervis Stal 1874, p. 52 [Spain]. — Horvath 1906a, p. 28. 

Orthostira nigrinervis: Ferrari 1878, p. 82. — Puton 1879c, p. 95 
[France].— Chicote 1880, p. 189. 
Type: Sex unknown; Spain; Stockholm Mus. 
Distribution: Spain; France. 
Host Plant: Unrecorded. 

* Acalypta nyctalis Drake 

Acalypta nyctalis Drake 1928c, pp. 3, 5 [N.H.; Alta.]. — Bailey 1951, 
p. 35.— Lindberg 1958b, p. 13 [Nfld.; Alaska; moss]. 
Type: Holotype 9, brachypterous; Franconia, New Hampshire, U.S.; 
USNM. 
Distribution: U.S. (N.H.; Alaska); Canada (Alta.; Nfld.). 
Host Plant: Mosses. 

* Acalypta parvula (Fallen) 

Tingis parvula Fallen 1807, p. 37 [Sweden]; 1829, p. 145. — Curtis 
1839, tab. 741 [England].— Fieber 1861, p. 384. 

Monanthia obscura Herrich-Schaeffer 1837, p. 23, pi. 118, fig. 372 
[Germany; moss]. — Vollenhoven 1878, p. 281, pi. 10, fig. 2. 

Orthosteira obscura: Fieber 1844, p. 54, pi. 4, figs. 22-25 [Czechoslovakia; 
Austria; Thymus serpyllus'].- — Sahlberg 1848, p. 130.— Herrich- 
SchaefTer 1850, p. 160.— Mayr 1858, p. 569 [Hungary]. 

Monanthia (Phyllontocheila) parvula: Fieber 1844, p. 66. 

Orthostira obscura: Fieber 1861, p. 131. — Douglas and Scott 1865, p. 
263.— Frey-Gessner 1865, p. 233 [Switzerland].— Garbiglietti 1869, 
p. 275 [Italy]. 

Orthostira parvula: Thomson 1871, p. 401.- — Reiber and Puton 1876, 
p. 68.— Puton 1879c, p. 99 [France; Corsica].— Reuter 1882b, p. 116 
[Finland].— Saunders 1892, p. 127, pi. 11, figs. 10, 10a.— Hiieber 
1893, p. 312.— de Walsche 1960, p. 38. 

Acalypta parvula: Stal 1874, p. 52.— Reuter 1874, p. 564; 1880b, p. 
165.— Saunders 1875, p. 252.— Horvath 1906a, p. 33 [Spain; Bel- 
gium]. — Oshanin 1908, p. 412 [Portugal; Algeria]. — Jensen-Haarup 
1912, p. 152, fig. 95 [Denmark].— Reichensperger 1920, p. 61 — 
Sahlberg 1920, p. 81 [Thymus].— Butler 1923, p. 204 [Scotland; 
Ireland; Herniaria].— Stichel 1926, p. 107, fig. 297; 1938a, p. 404 



LACEBUGS OF THE WORLD 53 

[Norway; Russia; Poland; Yugoslavia]; 1960a, p. 282, fig. 65 [Bul- 
garia; Sardinia; Calluna vulgaris; Coriolus versicolor; Trametes gibbosa; 
Polyporus hirsutus; Pholiota mutabilis]. — de Seabra 1931, p. 412, figs. 
484, 485.— Scholte 1935, p. 46, fig. 10.— Borner 1935, p. 75, fig. 
119b.— Poisson 1938, p. 589 [Erodium].— China 1938, p. 20 
[Madeira]. — Kiritshenko 1951, p. 245. — Novak and Wagner 1951, 
p. 70.— Leston 1953, p. 132, fig. 1.— Mancini 1953d, p. 15; 1954b, 
p. 11.— Gomez-Menor 1955b, p. 249— Stusak 1957b, pp. 135, 140, 
figs. 3, 15d; 1961a, p. 80, fig. 17.— Southwood and Leston 1959, p. 
144, pi. 25, fig. 4; fig. 42.— Lindberg 1960b, p. 92. 
Orthostira parvula var. macroptera Ferrari 1878, p. 83. 
Orthostira parvula var. pallescens Ferrari 1878, p. 83. 
Orthostira parvula var. major Ferrari 1878, p. 83. 

Orthostira parvula var. minor Ferrari 1878, p. 83. — Puton 1879c, p. 99. 
Monanthia parvula: Vollenhoven 1878, p. 278, pi. 9, figs. 10, 10a 

[Netherlands]. 
Campylostira parvula vav.fuscicornis Rey 1893, p. 97. 
Campylostira parvula var. minor: Rey 1893, p. 97. 
Type: Sex unknown; Scania, Sweden; Lund Zool. Inst. 
Distribution: England; Scotland; Ireland; Netherlands; Belgium; 
France; Portugal; Spain; Italy; Sardinia; Corsica; Switzerland; Czecho- 
slovakia; Austria; Hungary; Bulgaria; Yugoslavia; Poland; Germany; 
Denmark; Norway; Sweden; Finland; U.S.S.R.; Algeria; Madeira Islands. 
Host Plants: Calluna vulgaris; Climacium dendroides; Erodium sp.; Herniaria 
sp.; Thymus serpyllus; Thymus sp.; moss; fungi {Coriolus versicolor; Pholiota 
mutabilis; Polyporus hirsutus; Trametes gibbosa). 

Note: Study of eggs [Leston; Stusak]. Found in association with ants 
[Stichel 1960a]. 
*Acalypta platycheila (Fieber) 

Orthosteira platycheila Fieber 1844, p. 53, pi. 4, figs. 15-18 [Austria; 

Czechoslovakia]. — Herrich-Schaeffer 1850, p. 159. 
Monanthia {Orthosteira) intermedia Flor 1860, p. 339 [Estonia; Latvia]. 
Orthostira platychila [sic]: Fieber 1861, p. 130. — Thomson 1871, p. 400 
[Sweden].— Ferrari 1878, p. 83.— Puton 1879c, p. 97 [France].— 
Reuter 1882b, p. 115 [Lapland; moss].— Jako vie v 1893, p. 301 
[Russia].— Hiieber 1893, p. 306. 
Acalypta platychila [sic]: Reuter 1874, p. 563. — Sahlberg 1878, p. 21 
[Germany; Finland]; 1920, p. 79.— Butler 1923, p. 203 [Carex].— 
Stichel 1938a, p. 404 [Denmark; Belgium]. — Southwood and Leston 
1959, p. 142, pi. 21, fig. 7; fig. 50. 
Monanthia nigrina (not Fallen): Vollenhoven 1878, p. 277, pi. 9, figs. 9, 9a 
[Netherlands]. 



54 U.S. NATIONAL MUSEUM BULLETIN 243 

Acalypta platycheila: Horvath 1906a, p. 30, pi. 1, fig. 4 [England; Hungary; 
Siberia].— Stichel 1926, p. 107, fig. 295; 1960a, p. 279 [Norway; 
Poland; Yugoslavia].— Kiritshenko 1951, p. 244.— Bator 1953, 
p. 324.— Massee 1954, p. 259.— Mancini 1954b, p. 11 [Italy].— 
Cobben 1958b, p. 16. 
Type: Unknown. 

Distribution: England; Netherlands; Belgium; France; Italy; Czecho- 
slovakia; Austria; Hungary; Yugoslavia; Poland; Germany; Denmark; Swe- 
den; Norway; (Lapland); Finland; U.S.S.R. (Estonia; Latvia; Siberia). 
Host Plants: Carex sp.; moss. 

* Acalypta pulchra Stusak 

Acalypta pulchr a Stusak 1961b, p. 261, figs. 1-3 [Bulgaria; moss]. 
Type: Holotype d\ brachypterous; Borovec, Rila Planina, Bulgaria; 
Stusak Coll., Prague, Czechoslovakia. 
Distribution: Bulgaria. 
Host Plant: Moss. 

* Acalypta samara (Puton) 

Orthostira samara Puton 1887, p. 98 [Switzerland]. 

Acalypta samara: Horvath 1906a, p. 27. — Roubal 1958, p. 54 [France]. — 
Stichel 1960a, p. 274, fig. 44 [Austria]. 
Type: Sex unknown; Lugano, Switzerland; Paris Mus. 
Distribution: Switzerland; France; Austria. 
Host Plant: Unrecorded. 

* Acalypta saundersi (Downes) 

Drakella saundersi Downes 1927b, p. 60 [B.C.]. 

Acalypta saundersi: Drake 1928c, p. 4, fig. lb; 1930d, p. 268 [moss; 
Wash.]. 
Type: Holotype 9, macropterous; Goldstream, British Columbia, 
Canada; Canada. Coll. 
Distribution: Canada (B.C.); U.S. (Wash.; Oreg.). 
Host Plant: Moss. 

* Acalypta sauteri Drake 

Acalypta sauteri Drake 1942a, p. 14 [Japan]. — Takeya 1951a, p. 6. 
Type: Holotype o 71 , brachypterous; Oayama, Japan; Drake Coll. 
(USNM). 
Distribution: Japan. 
Host Plant: Unrecorded. 
Note: Intercepted at port-of-entry, New York City, on packing mosses. 

Acalypta sejuncta Horvath 

Acalypta sejuncta Horvath 1905b, p. 561 [Armenia]; 1906a, p. 30 [Syria]. — 
Oshanin 1908, p. 409 [Transcaucasus] .— Hoberlandt 1955, p. 86 
[Turkey] . 



LACEBUGS OF THE WORLD 55 

Type: Sex unknown; Vallis Araxis, Armenia, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Armenia; Transcaucasus) ; Syria; Turkey. 
Host Plant: Unrecorded. 

Acalypta sibirica Jakovlev 

Acalypta sibirica Jakovlev 1903a, p. 4 [Siberia]. — Horvath 1906a, p. 30. 

Type: Holotype cf, brachypterous; Irkoutsk, Siberia, Russia; Leningrad 
Inst. 

Distribution: U.S.S.R. (Siberia). 

Host Plant: Unrecorded. 

Acalypta sordida (Jakovlev) 

Orthostira sordida Jakovlev 1893, pp. 293, 301, 302 [Siberia]. 

Acalypta sordida: Reuter 1902, p. 151. — Horvath 1906a, p. 30. 
Type: Sex unknown; Siberia, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Siberia). 
Host Plant: Unrecorded. 

Acalypta subtilis (Reuter) 

Orthosteira subtilis Reuter 1882a, p. cxxx [Transcaucasus]. 

Acalypta subtilis: Horvath 1906a, p. 34. 
Type: Sex unknown; Lenkoran, Russia; Helsin. Mus. 
Distribution: U.S.S.R. (Transcaucasus; Caucasus). 
Host Plant: Unrecorded. 

Acalypta suturalis (Puton) 

Orthostira suturalis Puton 1879b, p. 297 [Spain]; 1879c, p. 95. — Chicote 

1880, p. 189. 
Acalypta suturalis: Horvath 1906a, p. 29. — Stichel 1960a, p. 277. 
Type: Sex unknown; Irun, Spain; Paris Mus. 
Distribution: Spain. 
Host Plant: Unrecorded. 

*Acalypta thomsonii Stal 

Acalypta thomsonii Stal 1873, p. 122 [Carolina]. — Osborn and Drake 

1916a, p. 220 [S.C.; Va.].— Heidemann 1917, p. 220, pi. 17, fig. 3 

(fig. only).— Parshley 1923b, p. 699.— Blatchley 1926, p. 480, 

fig. 110.— Bailey 1951, p. 35 [R.I.; Md.]. 

Acalypta thomsoni [sic]: Drake 1926b, p. 377, pi. 34, fig. d; 1928c, p. 4, 

fig. 2 [D.C.; moss; sphagnum]. 
Acalypta madelinae Torre -Bueno 1926b, p. 117 [Mass.]. 
Type: Lectotype cf, brachypterous; "Carolina meridionalis," U.S.; 
Stockholm Mus. 

Distribution: U.S. (Mass.; R.I.; N.J.; Md.; D.C.; Va.; N.C.; S.C.; 
Ga.; Fla.). 
Host Plants: Moss; sphagnum. 



56 U.S. NATIONAL MUSEUM BULLETIN 243 

* Acalypta uniseriata (Puton) 

Orthostira uniseriata Puton 1879b, p. 297 [Caucasus]; 1879c, p. 94. 

Acalypta uniseriata: Horvath 1906a, p. 28. — Kiritshenko 1951, p. 244. — 
Stichel 1960a, p. 275. 
Type: Sex unknown; Caucasus, Russia; Paris Mus. 
Distribution: U.S.S.R. (Caucasus). 
Host Plant: Unrecorded. 

* Acalypta vanduzeei Drake 

Acalypta vanduzeei Drake 1928c, pp. 3, 8 [Calif.]. 
Type: Holotype d\ brachypterous; Green Point Ranch, Humboldt 
County, California, U.S.; Cal. Acad. 
Distribution: U.S. (Calif.). 
Host Plant: Moss. 

*Acalypta vandykei Drake 

Acalypta vandykei Drake 1928c, pp. 3, 8 [Calif.].— Hurd 1946, p. 463 
[Oreg.]. 
Type: Holotype cf ; San Francisco County, California, U.S.; Cal. Acad. 
Distribution: U.S. (Calif.; Oreg.). 
Host Plant: Unrecorded. 

Genus ACANTHOCHEILA Stal 

Monanthia {Acanthoeheila) Stal 1858, p. 61. 

Acanthochila [sic]: Stal 1873, p. 127. — Lethierry and Severin 1896, p. 

14.— Banks 1910, p. 55. 
Acanthoeheila: Van Duzee 1916, p. 26; 1917b, p. 219.— Blatchley 

1926, p. 479.— Monte 1939b, p. 64; 1941e, p. 73.— Hurd 1946, p. 

469.— Drake and Ruhoff 1960a, p. 32. 
Type Species: Monanthia {Acanthoeheila) armigera Stal. 

* Acanthoeheila abducta Buchanan-White 

Acanthoeheila abducta Buchanan-White 1879, p. 485 [Brazil]. 

Acanthoeheila kahavalu Kirkaldy 1905, p. 216 [Peru]. — Drake 1922b, p. 42 
[Bolivia].— Monte 1940e, p. 287— Blote 1945, p. 85. 

Acanthoeheila kahavala [sic]: Drake 1931b, p. 511. 
Type: Holotype 9; Brazil; deposition unknown. 
Distribution: Brazil; Peru; Bolivia. 
Host Plant: Unrecorded. 

* Acanthoeheila armigera (Stal) 

Monanthia {Acanthoeheila) armigera Stal 1858, p. 61 [Brazil]. 

Monanthia {Acanthoeheila) spinuligera Stal 1858, p. 61. 

Acanthochila armigera: Stal 1873, p. 127. — Distant 1888, p. lxxxiii. — 
Champion 1897, p. 28, pi. 2, figs. 19, 19a, 20 [Mexico; Guatemala; 
Panama]; 1898b, p. 60.— Van Duzee 1907, p. 20 [Jamaica].— 
Osborn and Drake 1915b, p. 536. — Drake and Poor 1937d, p. 306 



LACEBUGS OF THE WORLD 57 

[Haiti; Peru; Bolivia; Colombia; Nicaragua; Puerto Rico; Cuba; 
Tex.]. 
Acanthochila spinuligera: Stal 1873, p. 127. 

Acanthocheila armigera: Drake and Bruner 1924a, p. 146 [Trinidad]. — 
Drake 1926b, p. 377; 1928e, p. 5 [Honduras]. — Drake and Hamble- 
ton 1934, p. 442 [Pisonia]; 1945, p. 359 [Ecuador].— Costa Lima 
1936, p. 124.— Monte 1938b, p. 128 [Argentina; Venezuela]; 1940d, 
p. 100 [Nicotiana tabacum]; 1942d, p. 108. — Singh 1953, p. 117. — Silva 
1956, p. 15, fig. 1 [Ouratea]. 
Acanthocheila nigrescens (not Drake and Bondar): Monte 1937a, p. 31 
[Pisonia tomentosa]. 
Type: Sex unknown; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Argentina; Bolivia; Brazil; Peru; Ecuador; Colombia; 
Venezuela; Panama; Nicaragua; Honduras; Guatemala; Mexico; U.S. 
(Tex.); Cuba; Puerto Rico; Trinidad; Jamaica; Haiti. 

Host Plants: Nicotiana tabecum; Ouratea sp.; Pisonia tomentosa; Pisonia 
sp. 
*Acanthocheila comentis Drake 

Acanthochila comentis Drake 1953a, p. 13 [Brazil]. 
Type: Holotype cf, macropterous; Vicosa, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Acanthocheila comitis Drake 

Acanthocheila comitis Drake 1948c, p. 23 [Maria Magdalena Island]. 
Type: Holotype 9; Magdalena Island, Tres Marias; Cal. Acad. 
Distribution: Mexico, Los Tres Marias Islands (Maria Magdalena). 
Host Plant: Unrecorded. 

Acanthocheila denieri Monte 

Acanthocheila denieri Monte 1940e, p. 287, fig. 1 [Argentina]. 

Type: Holotype o"; Clorinda, Formosa, Argentina; Monte Coll. (Mus. 
Nacional) . 

Distribution: Argentina. 

Host Plant: Unrecorded. 

* Acanthocheila dira Drake and Hambleton 

Acanthocheila dira Drake and Hambleton 1945, p. 359 [Guatemala]. 

Type: Holotype 9, macropterous; El Porvenir, Guatemala; Drake Coll. 
(USNM). 

Distribution: Guatemala. 

Host Plant: Unrecorded. 



681-552—64- 



58 U.S. NATIONAL MUSEUM BULLETIN 243 

*Acanthocheila exquisita Uhler 

Acanthocheila exquisita Uhler 1889, p. 143 [Fla.]. — Barber 1914, p. 507. — 
Drake 1919a, p. 421— Blatchley 1926, p. 479.— Monte 1940e, p. 
288, fig. 2. 
Type: Holotype 9, macropterous; Cape Florida, Florida, U.S.; Cornell 
Univ. 
Distribution: U.S. (Fla.); Bahama Islands. 
Host Plant: Unrecorded. 

* Acanthocheila hollandi Drake 

Acanthocheila hollandi Drake 1935, p. 16 [Brazil; Paraguay]; 1947a, p. 

2. — Drake and Hambleton 1938a, p. 46 [Bignoniaceae] . — Monte 

1940e, p. 287 [Argentina]. 
Type: Holotype 9; Chapalo, Brazil; Carnegie Mus. 
Distribution: Brazil; Argentina; Paraguay. 
Host Plant: Bignoniaceae. 

* Acanthocheila nexa Drake 

Acanthocheila nexa Drake 1936b, p. 701, fig. 1 [Argentina]. — Monte 
1940e, p. 288. 
Type: Holotype 9, macropterous; Loreto, Argentina; Leningrad Inst. 
Distribution: Argentina; Brazil. 
Host Plant: Unrecorded. 
*Acanthocheila nigrescens Drake and Bondar 

Acanthocheila nigrescens Drake and Bondar 1932, p. 88 [Brazil; Sapotaceus 
tree]. — Costa Lima 1936, p. 124. 
Type: Holotype d 1 , macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Sapotaceus tree. 

* Acanthocheila rustica Monte 

Acanthocheila rustica Monte 1942, p. 91 [Brazil]. 
Type: Holotype cT; Palmeiras, Ipaussu, Sao Paulo, Brazil; Monte Coll. 
(Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Acanthocheila rustica var. plana Drake 

Acanthochila rustica var. plana Drake 1953a, p. 13 [Argentina]. 
Type: Holotype 9, macropterous; Loreto, Misiones, Argentina; Drake 
Coll. (USNM). 
Distribution : Argentina. 
Host Plant: Unrecorded. 
*Acanthocheila sigillata Drake and Bruner 

Acanthocheila sigillata Drake and Bruner 1924a, p. 147 [Cuba; Pisonia 
aculeata]. — Drake 1926a, p. 86. — Bruner, Scaramuzza, and Otero 
1945, p. 141. 



LACEBUGS OF THE WORLD 59 

Type: Holotype o 71 , macropterous; Taco Taco, Pinar del Rio, Cuba; 
Drake Coll. (USNM). 
Distribution: Cuba. 
Host Plant: Pisonia aculeata. 

* Acanthocheila spinfcosta Van Duzee 

Acanthochila spinicosta Van Duzee 1907, p. 20 [Jamaica]. 
Acanthocheila spinicosta: Drake and Bruner 1924a, p. 146, fig. 1 [Domini- 
can Republic]. — Barber 1939, p. 370 [St. Thomas; Haiti; Pisonia 
domingensis]. 
Acanthocheila spinocosta [sic]: Gowdy 1926, p. 35. 
Type: Sex unknown; Mandeville, Jamaica; Cal. Acad. 
Distribution: Jamaica; Haiti; Dominican Republic; Puerto Rico; 
Virgin Islands (St. Thomas). 
Host Plants: Pisonia domingensis; Torrubia fragrans. 

*Acanthocheila thaumana Drake and Cobben 

Acanthocheila thaumana Drake and Cobben 1960, pp. 67, 81, fig. 82 
[St. Eustatius; St. Martin]. 
Type: Holotype o", macropterous; Koolbaai, St. Martin; Drake Coll. 
(USNM). 
Distribution: Leeward Islands (St. Eustatius; St. Martin). 
Host Plant: Unrecorded. 

*Acanthocheila tumida Drake 

Acanthocheila tumida Drake 1924, p. 94 [Bolivia]. — Drake and Hambleton 
1938a, p. 46 [Brazil; Anemopaegma prostratum]. — Monte 1938b, p. 128. 
Type: Holotype 9, macropterous; Cochabamba, Bolivia; Drake Coll. 
(USNM). 
Distribution: Bolivia; Brazil. 
Host Plant: Anemopaegma prostratum. 

* Acanthocheila visenda Drake and Hambleton 

Acanthocheila visenda Drake and Hambleton 1934, p. 442 [Brazil]. — 
Monte 1937a, p. 32 [Bignonia exoleta]; 1940e, p. 287 [Argentina]; 
1943a, p. 105 [Peru].— Drake and Davis I960, fig. 34. 
Type: Holotype cf, macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Argentina; Brazil; Peru. 
Host Plant: Bignonia exoleta. 

Genus ACANTHOTINGIS Monte 

Acanthotingis Monte 1940a, p. 13. — Drake and Ruhoff 1960a, p. 32. 
Type Species: Acanthotingis apicicornis Monte. 



60 U.S. NATIONAL MUSEUM BULLETIN 243 

Acanthotingis apicicornis Monte 

Acantholingis apicicornis Monte 1940a, p. 15, fig. [Brazil]; 1940e, p. 300; 
1941e, p. 149 [Rubiaceae]. 
Type: Holotype cf ; Sao Paulo, Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Rubiaceae. 

Genus ACONCHUS Horvath 

Galeatus (Aconchus) Horvath 1905b, p. 566. 

Aconchus: Horvath 1906a, pp. 14, 54.— Oshanin 1908, p. 424; 1912, 
p. 43.— Stichel 1926, pp. 104, 109.— Drake and Ruhoff 1960a, p. 32. 
Type Species: Galeatus {Aconchus) urbanus Horvath. 

* Aconchus urbanus (Horvath) 

Galeatus {Aconchus) urbanus Wow kth 1905b, p. 565 [Italy]. 
Aconchus urbanus: Horvath 1906a, p. 54; 1912c, p. 341 [Java]. — Drake 
and Poor 1937a, p. 17 [Malaya].— Drake and Ruhoff 1962d, p. 497. 
Aconchus ghesquierei Schouteden 1923, p. 89 [Congo]. — Mayn6 and 
Ghesquiere 1934, p. 19 [papaya]. 
Type: Sex unknown; Turin, Italy; Paris Mus. 

Distribution: Italy; Congo; Kenya; South Africa; Mozambique; India; 
Ceylon; Federation of Malaya; Philippine Islands; Greater Sunda Islands 
(Java). 
Host Plants: Urochloa reptans; papaya. 

Genus ACYSTA Champion 

Acysta Champion 1898a, p. 46. — Costa Lima 1936, p. 124. — Monte 
1939b, p. 64; 1941e, p. 76.— Hurd 1946, p. 458.— Drake and Ruhoff 
1960a, p. 32. 
Type Species: Acysta Integra Champion. 

* Acysta australica Drake 

Acysta australica Drake 1942a, p. 15 [Australia]. 
Type: Holotype 9, macropterous; National Park, Australia; Drake Coll. 
(USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

* Acysta hubbelli Drake 

Acysta hubbelli Drake 1928e, p. 2, fig. 1 [Honduras]. 
Type: Holotype o 71 ; Rio Sangrelaya, Honduras; Michigan Univ. 
Distribution: Honduras; Mexico (Vera Cruz). 
Host Plant: Unrecorded. 
Note: Collected on banana bits about to be planted. 



LACEBUGS OF THE WORLD 61 

Acysta integra Champion 

Acysta integra Champion 1898a, p. 46, pi. 3, fig. 22 [Guatemala]. 
Type: Sex unknown; Cerro Zunil, Guatemala; British Mus. 
Distribution: Guatemala. 
Host Plant: Unrecorded. 

Acysta interrupta Champion 

Acysta interrupta Champion 1898a, p. 47, pi. 3, fig. 23 [Panama]. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

* Acysta myrocarpi Drake and Poor 

Acysta myrocarpi Drake and Poor 1938a, p. 31 [Brazil]. — Monte 1941e, 
p. 76 [Myrocarpus fastigiatus']. 
Type: Holotype d", macropterous; Belo Horizonte, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Myrocarpus fastigiatus. 

* Acysta nectandrae Drake and Hambleton 

Acysta nectandrae Drake and Hambleton 1934, p. 439 [Brazil; Nectandra]. 
Type: Holotype cf, macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Nectandra sp. 

* Acysta neivai Drake and Hambleton 

Acysta neivai Drake and Hambleton 1940, p. 534 [Brazil]. 
Type: Holotype cf, macropterous; Guaruja, Sao Paulo, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

* Acysta ocoteae Drake and Hambleton 

Acysta ocoteae Drake and Hambleton 1935, p. 144 [Brazil; Ocotea pretiosa]. 
Type: Holotype cf , macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Ocotea pretiosa. 

* Acysta praeclara Drake and Hambleton 

Acysta praeclar a Drake and Hambleton 1934, p. 439 [Brazil; Boragina- 
ceae]. — Drake and Davis 1960, fig. 70. 
Type: Holotype cf , macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Boraginaceae. 



62 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Genus AEPYCYSTA Drake and Bondar 

Aepycysta Drake and Bondar 1932, p. 93. — Monte 1939b, p. 65; 1941e, 
p. 77.— Hurd 1946, p. 478.— Drake and Ruhoff 1960a, p. 33. 
Type Species: Aepycysta undosa Drake and Bondar. 

Aepycysta decorata Monte 

Aepycysta decorata Monte 1941d, p. 93, fig. 1 [Costa Rica]. 

Type: Holotype cf ; San Isidro de Coronado, Alajuela, Costa Rica; 
Monte Coll. (Mus. Nacional). 

Distribution: Costa Rica. 

Host Plant: Unrecorded. 

* Aepycysta schwarzi (Drake) 

Galeatus schwarzi Drake 1922b, p. 39, fig. 1 [Canal Zone]. 

Aepycysta schwarzi: Drake and Hambleton 1935, p. 153. 

Aepycysta schwartzi [sic]: Drake and Davis 1960, fig. 59. 
Type: Holotype o", macropterous; Paraiso, Panama Canal Zone; 
USNM. 
Distribution: Panama (Canal Zone). 
Host Plant: Unrecorded. 

* Aepycysta undosa Drake and Bondar 

Aepycysta undosa Drake and Bondar 1932, p. 94, fig. 1 [Brazil; Ichnanthus 
leiocarpus]. — Drake and Hambleton 1935, p. 153. — Costa Lima 1936, 
p. 125.— Monte 1938b, p. 127.— Silva 1956, p. 16, fig. 2. 
Type: Holotype o" , macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil; Paraguay. 
Host Plant: Ichnanthus leiocarpus. 

Genus AFRAMIXIA Drake and Ruhoff 

Ajramixia Drake and RuhofF 1960a, p. 33. 
Type Species: Epimixia roboris Drake. 

*Aframixia roboris (Drake) 

Epimixia roboris Drake 1942a, p. 12 [Malagasy]. 

Ajramixia roboris: Drake and Ruhoff 1960a, p. 33. 
Type: Holotype 9, macropterous; Madagascar; Drake Coll. (USNM). 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

Genus AGACHILA Drake and Gomez-Menor 

Agachila Drake and Gomez-Menor 1954, p. 89. — Drake and Ruhoff 
1960a, p. 34. 
Type Species: Agachila biqfrana Drake and Gomez-Menor. 



LACEBUGS OF THE WORLD 63 

Agachila biafrana Drake and Gomez-Menor 

Agachila biafrana Drake and Gomez-Menor 1954, p. 90, fig. 1 [Spanish 
Guinea]. 
Type: Holotype 9; Biafra, Cabo San Juan, Escalera, Spanish Guinea; 
Inst. Ent. Madrid. 
Distribution: Spanish Guinea. 
Host Plant: Unrecorded. 

Genus AGAOTINGIS Drake 

Agaotingis Drake 1954b, p. 13.— Drake and Ruhoff 1960a, p. 34. 
Type Species: Tingis australis Montrouzier. 

* Agaotingis australis (Montrouzier) 

Tingis australis Montrouzier 1864, p. 235 [New Caledonia]. — Stal 1873, 

p. 134. 
Agaotingis australis: Drake 1954b, p. 13. 
Type: Holotype d\ macropterous; New Caledonia; Drake Coll. 
(USNM). 
Distribution: New Caledonia. 
Host Plant: Unrecorded. 

Genus AGLOTINGIS Drake 

Aglotingis Drake 1954a, p. 232.— Drake and Ruhoff 1960a, p. 34. 
Type Species: Aglotingis nimbana Drake. 
Aglotingis affinis Schouteden 

Aglotingis affinis Schouteden 1955a, p. 32 [Congo]. 
Type: Sex unknown; Kikongo, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Aglotingis basilewskyi Schouteden 

Aglotingis basilewskyi Schouteden 1955a, p. 31 [Congo; Ruanda]; 
1957c, p. 313. 
Type: Unknown. 

Distribution: Congo; Ruanda-Urundi. 
Host Plant: Unrecorded. 

* Aglotingis nimbana Drake 

Aglotingis nimbana Drake 1954a, p. 233 [French Guinea]; 1955b, p. 90 
[Angola]. 
Type: Holotype 9, macropterous; Mt. Nimba, French Guinea; Inst. Fr. 
Afr. N. 
Distribution: Guinea; Angola. 
Host Plant: Unrecorded. 



64 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Genus AGRAMMA Stephens 

Agramma Stephens 1829a, p. 64; 1829b, p. 336.— Westwood 1840b, 
p. 120.— Fieberl844, p. 36; 1861, p. 118.— Garbiglietti 1869, p. 272.— 
Walker 1873b, p. 2— Horvath 1874b, p. 432.— China 1943, 
p. 248.— Hoberlandt 1955, p. 96.— Drake 1955d, p. 1.— Drake and 
Maa 1955, p. 10.— Drake and Ruhoff 1960a, p. 23.— Stichel 1960a, 
p. 344 (key); 1960b, p. 402; 1960c, p. 141. 

Serenthia Spinola 1837, p. 168. — Herrich-Schaeffer 1838, p. 47. — 
Stal 1873, p. 117; 1874, p. 46.— Lethierry and Severin 1896, p. 5 — 
Horvath 1906a, pp. 13, 107 (key).— Muzik 1907, p. 49 (key).— 
Oshanin 1908, p. 457; 1912, p. 46.— Schumacher 1914, p. 259; 
1919, p. 202.— Stichel 1926, p. 116 (key); 1935, p. 349; 1938a, 
p. 408— Borner 1935, p. 78 (key).— Guide 1938, p. 321 (key).— 
Wagner 1941, pp. 1-27 (key; subgenera).— Blote 1945, p. 79.— 
Kiritshenko 1951, pp. 243, 254 (key) .— Mancini 1953b, p. 186; 
1953d, p. 16.— Drake 1954b, p. 13; 1956a, p. 7; 1958a, p. 107.— 
Gomez-Menor 1955b, p. 248.— Drake and Ruhoff 1960a, p. 25. 

Wombalia Schouteden 1919, p. 139.— Drake and Ruhoff 1960a, p. 25. 

Serenthia {Agramma): Wagner 1941, pp. 6, 26. 

Serenthia (Paraserenthia) Wagner 1941, pp. 6, 26. — Drake and Ruhoff 
1960a, p. 25. 

Serenthia (Set enthiella) Wagner 1941, pp. 6, 24, 26. — Drake and Ruhoff 
1960a, p. 25. 

Drakea Schouteden 1953d, p. 166. — Drake and Ruhoff 1960a, p. 24. 
Type Species: Tingis laeta Fallen. 

* Agramma afranum Drake and Ruhoff 

Agramma angolana Drake 1958a, p. 107, fig. (not 1955) [Angola]. 

Agramma qfrana Drake and Ruhoff 1960c, p. 36. 
Type: Holotype cf ; Alto Chicapa, Angola; Cent. Afr. Mus. 
Distribution: Angola. 
Host Plant: Unrecorded. 

* Agramma aliwalanum (Drake) 

Serenthia aliwalana Drake 1954b, p. 14 [Cape Province]. 

Agramma aliwalanum: Drake 1958a, p. 107. 
Type: Holotype d"; North Aliwal, Cape Province, South Africa; British 
Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

* Agramma angolanum (Drake) 

Serenthia angolana Drake 1955b, p. 90 [Angola]. 

Agramma angolanum: Drake 1958a, p. 107. 
Type: Holotype cf; Serra do Moco, Luimbale, Angola; Cent. Afr. Mus. 
Distribution: Angola. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 65 

*Agramma atricapillum (Spinola) 

Serenthia atricapilla Spinola 1837, p. 168 [Sardinia]. — Herrich-Schaeffer 
1838, p. 48.— Amyot and Serville 1843, p. 300.— Costa 1847a, p. 24; 
1847c, p. 164.— Garbiglietti 1869, p. 272 [Italy].— StM 1874, p. 47 — 
Puton 1879c, p. 89 [France; Corsica]. — -Bolivar and Chicote 1879, 
p. 166 [Spain].— Jakovlev 1880b, p. 100 [Caucasus].— Horvath 1889, 
p. 329; 1892b, p. 131; 1906d, p. 2; 1916, p. 9.— Reuter 1891a, p. 25 
[Greece; Albania; Crete]; 1908, p. 89 [Juncus]. — Lethierry and 
Severin 1896, p. 5 [Algeria]. — Oshanin 1908, p. 457 [Morocco]. — 
Moroder Sala 1920, p. 13.— de Seabra 1924, p. 18, fig. 32 [Portugal]; 
1931, p. 442, figs. 512(1-4), 513.— Lindberg 1932a, p. 45 [Scirpus 
holoschoenus]; 1948, p. 60 [Sicily; Egypt; Bulgaria; Juncus aculus]. — 
Wagner 1941, pp. 6, 13, 26, figs. 2d, 3e, 4d, 5f, 6d— Hoberlandt 
1949, p. 8 [Tunisia].— Kiritshenko 1951, p. 254 [Ukraine; Crimea].— 
Novak and Wagner 1951, p. 71 [Typha latifolia]. — Seidenstiicker 1954, 
p. 236. 
Agramma atricapilla: Fieber 1844, p. 39; 1861, p. 118 [Yugoslavia]. — ■ 
Mayr 1858, p. 569 [Hungary].— Jakovlev 1871, p. 7; 1876b, p. 66.— 
Hoberlandt 1955, p. 96 [Israel; Iraq] .— Gomez-Menor 1956b, p. 82, 
fig. — Stichel 1960a, p. 345, figs. 183-186 [Juncus marilumus], 
Serenthia brevirostr is Jakovlev 1901, p. 35. 

Serenthia antricapilla [sic]: Horvath 1906a, p. 110 [Rumania; Turkestan; 
Turkey]. 
Type: Sardinia; sex and deposition of type unknown. 
Distribution: Portugal; Spain; France; Corsica; Italy; Sicily; Sardinia; 
Hungary; Bulgaria; Rumania; Yugoslavia; Albania; Greece; Crete; Turkey; 
Iraq; Israel; U.S.S.R. (Caucasus; Ukraine; Crimea; Turkestan); Egypt; 
Morocco; Algeria; Tunisia. 

Host Plants: Juncus acutus; Juncus maritimus; Juncus sp.; Scirpus holoschoe- 
nus; Typha latifolia. 
Agramma atricapillum var. mendax (Horvath) 

Serenthia antricapilla [sic] var. mendax Horvath 1906a, pp. 107, 111 
[Turkestan]. 
Type: Sex unknown; Michailovo, Turkestan, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 
Agramma atricapillum var. pallens (Horvath) 

Serenthia antricapilla [sic] var. pallens Horvath 1906a, p. Ill [Egypt]. 
Serenthia atricapilla var. pallens: Priesner and Alfieri 1953, p. 66 
[Gramineae]. 
Type: Sex unknown; Fayum, Egypt; Hungarian Mus. 
Distribution: Egypt. 
Host Plant: Gramineae. 



66 U.S. NATIONAL MUSEUM BULLETIN 243 

*Agramma basilicorne (Drake) 

Serenthia basilicornis Drake 1951, p. 167 [Tanganyika]. 
Type: Holotype cf; Shirati, Africa orientalis; Hungarian Mus. 
Distribution: Tanganyika; Egypt. 
Host Plant: Unrecorded. 

* Agramma blandulum (Horvath) 

Serenthia blandula Horvath 1905b, p. 558 [Caucasus]; 1906a, p. 116 
[Turkey]. — Kiritshenko 1951, p. 255 [Crimea]. 

Serenthia (Serenthiella) blandula: Wagner 1941, p. 25. 

Agramma {Serenthiella) blandula: Hoberlandt 1955, p. 97. 

Agramma blandula: Stichel 1960a, p. 35, fig. 217 [Greece]. 
Type: Sex unknown; Caucasus, Russia; Paris Mus. 
Distribution: U.S.S.R. (Caucasus; Crimea); Turkey; Greece. 
Host Plant: Unrecorded. 

* Agramma carinatum (Distant) 

Serenthia carinata Distant 1911b, p. 269 [Ceylon; Juncus]. 
Type: Sex unknown; Ambalangoda Lake, Ceylon; British Mus. 
Distribution: Ceylon. 
Host Plant: Juncus sp. 

* Agramma confusum (Puton) 

Serenthia laeta (not Fallen): Herrich-Schaeffer 1838, p. 49, pi. 122, 
fig. 388. — Amyot and Serville 1843, p. 300 (in part) [France]. — Costa 
1847a, p. 24; 1847c, p. 164. 
Sercuthia [sic] jemoralis var. confusa Puton 1879b, p. 297. 
Serenthia femoralis var. confusa: Puton 1879c, p. 90 [Corsica]. — Chicote 

1880, p. 189 [Spain].— Horvath 1906d, p. 2. 
Serenthia confusa: Reuter 1885, p. 43. — Horvath 1906a, pp. 109, 114 
[Italy; Germany; Hungary; Yugoslavia; Greece; Rumania; Russia; 
Turkey; Juncus]; 1916, p. 9 [Albania]; 1918, p. 334.— Stichel 1926, 
p. 117 [Plantago maritima]. — Borner 1935, p. 79, fig. 121 [Juncus 
gerardi]. — Guide 1938, p. 326, fig. [Cyperaceae]. — Kiritshenko 1951, 
p. 254 [Ukraine; Crimea]. — Mancini 1953b, p. 186 [Turkestan]. 
Serenthia (Agramma) confusa: Wagner 1941, pp. 7, 15, 26, figs. 7a, 8a, 9a, 

11a.— Hoberlandt 1942, p. 126 [Czechoslovakia]. 
Agramma confusa: Hoberlandt 1955, p. 96 [Transcaucasus]. — Stichel 
1960a, p. 349, figs. 206-207 [Bulgaria; Poland; Austria]. 
Type: Unknown. 

Distribution: Spain; France; Germany; Austria; Czechoslovakia; Po- 
land; Hungary; Italy; Corsica; Rumania; Bulgaria; Albania; Yugoslavia; 
Greece; Turkey; U.S.S.R. (Caucasus; Ukraine; Crimea; Transcaucasus; 
Turkestan). 
Host Plants: Juncus gerardi; Juncus sp.; Plantago maritima; Cyperaceae. 



LACEBUGS OF THE WORLD 67 

Agramma confusum var. antennatum (Horvath) 

Serenthia jemoralis var. antennata Horvath 1905b, p. 556 [Rumania]. 
Serenthia conjusa var. antennata: Horvath 1906a, p. 114 [Hungary; 
Turkestan]. 
Type: Sex unknown; Rumania; Hungarian Mus. 
Distribution: Rumania; Hungary; U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 

Agramma confusum var. thoracicum (Horvath) 

Serenthia Jemoralis var. thoracica Horvath 1905b, p. 557 [Hungary]. 
Serenthia confusa var. thoracica: Horvath 1906a, p. 114. — Hoberlandt 
1943, pp. 116, 117 [Czechoslovakia]. 
Type: Sex unknown; Hungary; Hungarian Mus. 
Distribution: Hungary; Czechoslovakia. 
Host Plant: Unrecorded. 

* Agramma dilectulum (Drake) 

Serenthia dilectula Drake 1951, p. 168 [Ethiopia]. 

Agramma dilectula: Drake 1956f, p. 429 [Cape Province]; 1958a, p. 107. 
Type: Holotype cf ; Maraquo, Abyssinia; Hungarian Mus. 
Distribution: Ethiopia; South Africa (Cape Province). 
Host Plant: Unrecorded. 

* Agramma dubium (Horvath) 

Serenthia ruficornis var. dubia Horvath 1905b, p. 556 [Algeria]. 
Serenthia dubia: Horvath 1906a, p. 112. — Lindberg 1932a, p. 45 

[Spain]. 
Agramma dubia: Drake 1958a, p. 107 [Mauritania]. — Stichel 1960a, 
p. 347 [Morocco]. 
Type: Sex unknown; Philippe ville, Algeria; Hungarian Mus. 
Distribution: Algeria; Mauritania; Tunisia; Spain. 
Host Plant: Unrecorded. 
Agramma dubium var. imbecillum (Horvath) 

Serenthia Jemoralis jvar. imbecilla Horvath 1905b, p. 557 [Mauritania]. 
Serenthia dubia var. imbecilla: Horvath 1906a, p. 113. — Lindberg 1932a, 

p. 45 [Spain; Morocco; Scirpus holoschoenus]. 
Agramma dubia var. imbecilla: Stichel 1960a, p. 347. 
Type: Sex unknown; Mauritania; Hungarian Mus. 
Distribution: Mauritania; Morocco; Spain. 
Host Plant: Scirpus holoschoenus. 
Agramma ecmeles Drake and Ruhoff 

Agramma ecmeles Drake and Ruhoff 1962a, p. 164 [Western Australia]. 
Type: Holotype cf, macropterous; Kimberley District, north Western 
Australia, Australia; Stockholm Mus. 

Distribution: Australia (Western Australia). 
Host Plant: Unrecorded. 



68 U.S. NATIONAL MUSEUM BULLETIN 243 

*Agramma elegans Kiritshenko 

Agramma elegans Kiritshenko 1952, p. 180, fig. 26 [Russia]. 
Type: Sex unknown; Tadshikistan, Russia; Leningrad Inst. 
Distribution: U.S.S.R. 
Host Plant: Unrecorded. 
Agramma fallax (Horvath) 

Serenthia confusa var. fallax Horvath 1906a, p. 114 [Hungary; Russia]. 
Serenthia fallax: Stichel 1926, p. 117 [Germany]. — Borner 1935, p. 79 

[Juncus]. 
Serenthia (Agramma) fallax: Wagner 1941, pp. 7, 17, 26. 
Agramma fallax: Stichel 1960a, p. 349. 
Type: Unknown. 

Distribution: Germany; Hungary; U.S.S.R. 
Host Plant: Juncus sp. 
* Agramma femorale Thomson 

Agramma laetum (not Fallen): Flor 1860, p. 324 [Sweden; Germany; 

France; England; Italy; Spain; Schoenus albus]. 
Agramma femoralis Thomson 1871, p. 397. — Stichel 1960a, p. 351 

[Rhynchospora alba]. 
Serenthia femoralis: Stal 1874, p. 46. — Reuter 1874, p. 562 [Finland; 
Eriophorum angustifolium]; 1882b, p. Ill; 1885, p. 43; 1890a, p. 249.— 
Sahlberg 1878, p. 20 [Latvia]; 1920, p. 86— Hiieber 1893, p. 299 
[Hungary; Austria]. 
Serenthia femorata [sic]: Jakovlev 1893, p. 293 [Russia]. 
Serenthia laeta var. femoralis: Horvath 1906a, p. 115 [Siberia]. — Oshanin 

1908, p. 461 [Morocco; Rumania].— Kiritshenko 1951, p. 254. 
Serenthia (Agramma) laeta var. femoralis: Wagner 1941, p. 24, fig. lOe. 
Type: Unknown. 

Distribution: Sweden; Finland; U.S.S.R. (Latvia; Siberia); Hungary; 
Austria; Germany; France; Rumania; Italy; Spain; England; Morocco. 

Host Plants: Eriophorum angustifolium; Rhynchospora alba; Schoenus albus. 
Agramma femorale var. poppii (Horvath) 

Serenthia laeta var. poppii Horvath 1906a, p. 116 [Siberia]. — Stichel 1926, 

p. 118.— Guide 1938, p. 327. 
Serenthia femoralis var. poppiusi: Sahlberg 1920, p. 86 [Finland]. 
Serenthia laeta var. poppiusi: Stichel 1935, p. 350; 1938b, p. 455. 
Agramma femoralis wax. poppiusi: Stichel 1960a, p. 351. 
Type: Sex unknown; Saoneskje Kosmosero, Siberia, Russia; Helsin. Mus. 
Distribution: U.S.S.R. (Siberia); Finland. 
Host Plant: Unrecorded. 
Agramma formosanum (Matsumura) 

Serenthia formosana Matsumura 1910, p. 24, pi. 12, fig. 10 [Taiwan]; 
1911, p. 138.— Esaki 1926, p. 163; 1932, p. 1637, fig— Takeya 1951a, 
p. 23 [Saccharum officinarum]. — Box 1953, p. 37. 



LACEBUGS OF THE WORLD 69 

Type: Holotype 9; Rinkiho, Formosa; deposition unknown. 

Distribution: Taiwan. 

Host Plant: Saccharum officinarum. 

*Agramma gibbum Fieber 

Agramma gibba Fieber 1844, p. 38, pi. 3, figs. 7-11 [Ostindien]. 
Serenthia gibba: Stal 1873, p. 117 [India orientalis]. — Distant 1903b, 
p. 127, fig. 91 [India].— Drake 1938b, p. 195 [East Indies; Hainan 
Island].— Blote 1945, p. 79. 
Type: Sex unknown; "Ostindien"; Vienna Mus. 
Distribution: India; East Indies(?); China (Hainan Island); Ceylon. 
Host Plant: Unrecorded. 

Agramma globiceps (Horvath) 

Serenthia globiceps Horvath 1906a, p. Ill [Syria]. 
Type: Sex unknown; Jaffa, Syria; Hungarian Mus. 
Distribution: Syria. 
Host Plant: Unrecorded. 

* Agramma gracilicorne (Wagner) 

Serenthia (Paraserenthia) gracilicornis Wagner 1941, pp. 6, 11, 26, figs. 2b, 

3b, 4 c, f, 5b, 6b [Germany]. 
Agramma gracilicornis: Stichel 1960a, p. 346, figs. 193-196 [Austria; 
Poland]. 
Type: Sex unknown; Germany; Wagner Coll. 
Distribution: Germany; Austria; Poland. 
Host Plant: Unrecorded. 

* Agramma hupehanum (Drake and Maa) 

Serenthia hupehana Drake and Maa 1954, p. Ill [China]. 
Type: Holotype cf; Suisapa, Western Hupeh, China; Cal. Acad. 
Distribution: China; India (Bengal). 
Host Plant: Unrecorded. 

* Agramma intermedium (Wagner) 

Serenthia {Agramma) intermedia Wagner 1941, pp. 7, 20, 26, figs. 7c, 8c, 9c, 

10 a, c, lie [Trieste]. 
Serenthia intermedia: Novak and Wagner 1951, p. 71 [Yugoslavia; Typha 
latifolia]. — Mancini 1953b, p. 186 [Italy; Germany; Hungary; 
Greece] . 
Agramma intermedia: Hoberlandt 1955, p. 96 [Turkey]. — Stichel 1960a, 
p. 348, figs. 197-200 [France; Austria; Czechoslovakia; Poland]. 
Type: Sex unknown; Trieste; Wagner Coll. 

Distribution: Trieste; Yugoslavia; Greece; Turkey; Hungary; Austria; 
Italy; Czechoslovakia; Germany; Poland; France. 
Host Plant: Typha latifolia. 



70 U.S. NATIONAL MUSEUM BULLETIN 243 

*Agramma japonicum (Drake) 

Serenthia japonica Drake 1948g, p. 174 [Japan]. — Takeya 1951a, p. 23. — 
Drake and Maa 1953, p. 88 [China]. 
Type: Holotype 9, brachypterous; Sapporo, Japan: Drake Coll. (USNM). 
Distribution: Japan; China. 
Host Plant: Unrecorded. 

*Agramma karisimbiense (Schouteden) 

Serenthia karisimbiensis Schouteden 1953d, p. 165 [Ruanda]. 

Agramma karisimbiensis: Drake 1958a, p. 107. 
Type: Sex unknown; Nya Muzinga, Volcan Karisimibi, Ruanda; Cent. 
Afr. Mus. 
Distribution: Ruanda-Urundi; Kenya. 
Host Plant: Unrecorded. 

* Agramma kivuanum (Drake) 

Serenthia kivuana Drake 1956a, p. 8 [Congo]. 

Agramma kivuanum: Drake 1958a, p. 107. 
Type: Holotype cf; lac Magera, Kivu, Belgian Congo; Pares Nat. Inst. 
Distribution: Congo. 
Host Plant: Unrecorded. 

* Agramma laetum (Fallen) 

Tingis laeta Fallen 1807, p. 40 [Sweden; Germany]; 1829, p. 151.— 
Germar 1835, fasc. 10, tab. 14.— Herrich-Schaeffer 1835, p. 59 — 
Brulle 1835, p. 340.— Blanchard 1840, p. 112. 

Agramma laeta: Stephens 1829a, p. 64 [England]; 1829b, p. 336 — 
Westwood 1840b, p. 120.— Fieber 1844, p. 37, pi. 2, figs. 22-28 
[Czechoslovakia; Italy]; 1861, p. 119.— Mayr 1858, p. 569 [Hungary; 
Austria; moss]. — Frey-Gessner 1865, p. 231 [Switzerland; Carex]. — 
Douglas and Scott 1865, p. 242.— Jakovlev 1869, p. Ill [Russia].— 
Gredler 1870, p. 75— Stusak 1958, p. 362, figs. 1-4.— Southwood 
and Leston 1959, p. 151, pi. 18, fig. 1.— Drake and Ruhoff 1960a, 
pi. 1.— Stichel 1960a, p. 350, figs. 212-214 [Sardinia; Poland]. 

Piesma tricolor Laporte 1833, p. 48 [France]. 

Piesma laetum: Burmeister 1835, p. 257. 

Serenthia laeta: Herrich-Schaeffer 1838, p. 49, pi. 122, figs, b-f; pi. 125, 
fig. c— Amyot and Serville 1843, p. 300 (in part).— Stal 1874, p. 46 — 
Saunders 1875, p. 246; 1892a, p. 123, pi. 11, fig. 5 [heath].— Reiber 
and Puton 1876, p. 67.— Puton 1879c, p. 90 [Corsica].— Bolivar and 
Chicote 1879, p. 166 [Spain; Portugal].— Jakovlev 1880b, p. 100 
[Caucasus]; 1893, p. 293.— Reuter 1882b, p. Ill [Juncus]; 1890a, 
p. 249; 1891a, p. 25 [Greece].— Dubois 1888, p. 120.— Hueber 1893, 
p. 297— Rey 1893, p. 97.— Horvath 1905b, p. 557 [Trieste]; 1906a, 
pp. 110, 115 [Siberia; Schoenus albus].— Muzik 1907, p. 49, fig. 2 — 
Oshanin 1908, p. 460 [Belgium; Yugoslavia] .— Jensen-Haarup 1912 ? 



LACEBUGS OF THE WORLD 71 

p. 149, fig. 93 [Denmark]. — Reichensperger 1920, p. 61. — Butler 
1923, p. 200 [Wales; rushes].— Stichel 1926, p. 118, fig. 309 [Rhyncho- 
spora alba}; 1938a, p. 408 [Finland; Norway].— de Seabra 1931, p. 443, 
figs. 512(6), 515.— Scholte 1935, p. 92, fig. 27 [Luzula campestris].— 
Poisson 1938, p. 589 [Juncus rnaritimus]. — Blote 1945, p. 80. — Kirit- 
shenko 1951, p. 254, fig. 330.— Gomez-Menor 1956a, p. 115, fig. 10 
(fig. only). 
Agramma laetum: Vollenhoven 1878, p. 285, pi. 22, fig. 11 [Netherlands]. 
Serenthia {Agramma) laeta: Wagner 1941, pp. 6, 21, 26, figs. 1, 6g, 7d, 
8d, 9d, lOd, lid.— Hoberlandt 1942, p. 126. 
Type: Unknown. 

Distribution: England; Wales; Netherlands; Belgium; France; Portugal; 
Spain; Corsica; Sardinia; Italy; Switzerland; Germany; Austria; Czecho- 
slovakia; Hungary; Trieste; Yugoslavia; Greece; U.S.S.R. (Caucasus; 
Siberia); Poland; Finland; Denmark; Norway; Sweden; Mongolia 
Republic. 

Host Plants: Car ex sp.; Juncus rnaritimus; Juncus sp.; Luzula campestris; 
Rhynchospora alba; Schoenus albus; heath; rushes; moss. 
Note: Eggs [Stusak]. 

* Agramma laetum var. apicicorne (Horvath) 

Serenthia laeta var. apicicornis Horvath 1905b, p. 557 [Trieste]; 1906a, 
p. 115. 
Type: Holotype 9; Trieste; Hungarian Mus. 
Distribution: Trieste. 
Host Plant: Unrecorded. 

Agramma leleupi (Schouteden) 

Drakea leleupi Schouteden 1953d, p. 166 [Congo]. 
Type: Sex unknown; Kundelungu, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Agramma lineatum (Horvath) 

Serenthia lineata Horvath 1929, p. 326 [Natal]. 

Agramma lineatum: Drake 1958a, p. 107. 
Type: Sex unknown; Weenan, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

* Agramma longum (Drake) 

Serenthia longa Drake 1954b, p. 13 [Cape Province]. 

Agramma longum: Drake 1958a, p. 107. 
Type: Holotype o 71 ; Ceres, Cape Province, South Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 



72 U.S. NATIONAL MUSEUM BULLETIN 243 

* Agramma longurium Drake 

Agramma longuria Drake 1958b, p. 30 [Congo]. 
Type: Holotype cf ; Lusinga, Belgian Congo; Pares Nat. Inst. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Agramma maynei (Schouteden) 

Serenthia maynei Schouteden 1916b, p. 291 [Congo]; 1923, p. 84. 
Agramma maynei: Drake 1958a, p. 107. 
Type: Sex unknown; Congo da Lemba, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

* Agramma melanoscele (Horvath) 

Serenthia melanoscelis Horvath 1906a, p. 114 [Algeria].— Blote 1945, p. 80 

[Tunisia]. 
Serenthia {Agramma) melanoscelis: Wagner 1941, pp. 7, 18, 26, figs. 7b, 

8b, 9b, lib [Germany; Juncus]. 
Agramma malonascelis [sic]: Drake 1958a, p. 107. 
Agramma melanoscelis: Stichel 1960a, p. 348, figs. 201-204. 
Type: Sex unknown; Tlemcen, Algeria; Hungarian Mus. 
Distribution: Algeria; Tunisia; Upper Volta; Germany. 
Host Plant: Juncus sp. 

Agramma melanoscele var. muelleri (Wagner) 

Serenthia {Agramma) melanoscelis var. muelleri Wagner 1941, p. 19 
[Germany]. 

Agramma melanoscelis var. muelleri: Stichel 1960a, p. 348. 
Type: Sex unknown; Nurnburg, Germany; Wagner Coll. 
Distribution: Germany. 
Host Plant: Unrecorded. 

Agramma minor (Distant) 

Lullius? minor Distant 1904, p. 430, pi. 8, fig. 8 [Cape Province]. 
Agramma minor: Drake and Ruhoff 1960a, p. 24. 
Type: Sex unknown; Cape Colony, South Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

*Agramma minutum Horvath 

Agramma minutum Horvath 1874a, p. 333 [Hungary]. 

Agramma deprcssa Jakovlev 1874b, p. 265 [Russia]. 

Serenthia depressa: Puton 1876, p. 290.— Jakovlev 1893, p. 293. 

Serenthia minuta: Hiieber 1893, p. 300 [Austria]. — Horvath 1906a, p. 116 
[Siberia; Rumania]. — Oshanin 1908, p. 462 [Caucasus]. — Hober- 
landt 1943, pp. 116, 117.— Kiritshenko 1951, p. 255 [Ukraine].— 
Remold 1959, p. 1, fig. 1. 



LACEBUGS OF THE WORLD 73 

Serenthia (Serenthiella) minuta: Wagner 1941, pp. 8, 25, 26, figs. 7f, 8f, 

lie. — Hoberlandt 1942, p. 126 [Germany; Czechoslovakia]. 
Agramma minuta: Wagner 1958, p. 241 [France]. — Stichel 1960a, p. 351 
[Bulgaria]. 
Type: Sex unknown; Hungary; Hungarian Mus. 

Distribution: France; Germany; Czechoslovakia; Austria; Hungary; 
Rumania; Bulgaria; U.S.S.R. (Siberia; Caucasus; Ukraine). 
Host Plant: Unrecorded. 

* Agramma nexile (Drake) 

Serenthia nexilis Drake 1948g, p. 174 [Taiwan; Japan]. — Takeya 1951a, 
p. 23; 1953d, p. 167.— Esaki 1954, p. 238, fig. 615. 
Type: Holotype cT, macropterous; Formosa; Drake Coll. (USNM). 
Distribution: Taiwan; Japan. 
Host Plant: Unrecorded. 

Agramma nigrellum Drake 

Agramma nigrella Drake 1958a, p. 108, fig. [Angola]. 
Type: Holotype o" ; Vila Luso, Angola; Cent. Afr. Mus. 
Distribution: Angola. 
Host Plant: Unrecorded. 
Note: Taken in light trap [Drake]. 
*Agramma nigrum Fieber 

Agramma nigra Fieber 1844, p. 40, pi. 3, figs. 12-17 [Sicily; Statice]; 
1861, p. 118.— Herrich-Schaeffer 1850, p. 150.— Drake 1958a, p. 
107.— Stichel 1960a, p. 347. 
Serenthia nigra: Garbiglietti 1869, p. 272— Stal 1874, p. 46. — Horvath 
1906a, p. 113 [Tunisia; Algeria]. 
Type: Sex unknown; Sicily; Vienna Mus. 
Distribution: Sicily; Algeria; Tunisia. 
Host Plant: Statice sp. 
Agramma nigrum var. collare (Horvath) 

Serenthia nigra var. collaris Horvath 1905b, p. 558 [Sicily; Algeria]; 

1906a, p. 114. 
Agramma nigra var. collaris: Stichel 1960a, p. 347. 
Type: Unknown. 
Distribution: Sicily; Algeria. 
Host Plant: Unrecorded. 
* Agramma onar Drake Plate 14 

Agramma onar Drake 1961c, p. 130, pi. 9 [Natal]. 
Type: Holotype o 71 , brachypterous; Cathedral Peak Forestry Reserve, 
Natal Drakensberg, headwaters of the Indumeni River, South Africa; 
Natal Museum, Pietermaritzburg, Natal, South Africa. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

681-552—64 6 



74 U.S. NATIONAL MUSEUM BULLETIN 243 

Agramma pallidulum (Schouteden) 

Serenthia pallidula Schouteden 1955a, p. 26 [Congo]. 

Agramma pallida [sic]: Schouteden 1957c, p. 312 [Ruanda]. 

Agramma pallidulum: Drake 1958a, p. 107. 
Type: Sex unknown; Rutshuru, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Ruanda-Urundi. 
Host Plant: Unrecorded. 

Agramma pallidulum var. pictum (Schouteden) 

Serenthia pallidula var. picta Schouteden 1955a, p. 27 [Congo]. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

* Agramma peringueyi (Distant) 

Serenthia per ingueyi Distant 1904, p. 429, pi. 8, fig. 6 [South Africa]. — 

Drake 1956a, p. 7 [Ruanda]. 
Agramma peringueyi: Drake 1956f, p. 429 [Basutoland] ; 1958a, p. 107; 
1958c, p. 316 [Malagasy; Kenya]. 
Type: Sex unknown; South Africa; So. Afr. Mus. 

Distribution: South Africa; Basutoland; Kenya; Ruanda-Urundi; 
Malagasy Republic. 
Host Plant: Unrecorded. 

Agramma pictipenne (Horvath) 

Serenthia pictipennis Horvath 1902b, p. 604 [New South Wales]. 
Agramma pictipennis: Drake and Ruhoff 1961b, p. 180, fig. 21. 
Type: Sex unknown; Glen Innes, New South Wales, Australia; Hun- 
garian Mus. 
Distribution: Australia (New South Wales). 
Host Plant: Unrecorded. 

Agramma pullum Drake 

Agramma pulla Drake 1958b, p. 30, fig. 1 [Congo]. 

Type: Holotype cf; Mukana-Lusinga, Belgian Congo; Pares Nat. 
Mus. 

Distribution: Congo. 

Host Plant: Unrecorded. 

* Agramma ruficorne (Germar) 

Tingis ruficornis Germar 1835, fasc. 15, tab. 12 [Switzerland]. — Herrich- 
Schaeffer 1835, p. 59. 

Serenthia ruficornis: Herrich-Schaeffer 1838, p. 48 [Germany]. — Stal 
1874, p. 47.— Puton 1879c, p. 90 [France] .—Dubois 1888, p. 119 — 
Hueber 1893, p. 297.— Horvath 1906a, pp. 108, 112.— Oshanin 1908, 
p. 458 [Turkestan]. — Schumacher 1914, p. 259 [Carex; sphagnum]. — 
Stichel 1926, p. 117 [Juncus]; 1938a, p. 408 [Poland].— de Seabra 



LACEBUGS OF THE WORLD 75 

1931, p. 442, figs. 512(5), 514 [Portugal].— Scholte 1935, p. 92 
[Netherlands].— Bator 1953, p. 327, pi. 4, fig. 7. 
Agramma ruficornis: Fieber 1844, p. 38, pi. 3, figs. 1-6 [Italy; Austria; 
Czechoslovakia]; 1861, p. 118.— Mayr 1858, p. 569 [Hungary].— 
Frey-Gessner 1865, p. 231 [moss].— Stichel 1960a, p. 346, figs. 191-192 
[Sardinia]. 
Scraulia ruficornis: Lethierry and Severin 1896, p. 8. 

Serenthia (Paraserenthia) ruficornis: Wagner 1941, pp. 6, 10, 26, figs. 2c, 
3c, 4 b, e, 5c, 6c— Hoberlandt 1942, p. 126. 
Type: Switzerland; sex and deposition of type unknown. 
Distribution: Netherlands; France; Switzerland; Germany; Hungary; 
Czechoslovakia; Austria; Italy; Sardinia; Portugal; Poland; U.S.S.R. 
(Turkestan). 
Host Plants: Carex sp.; Juncus sp.; moss; sphagnum. 

Agramma rwindianum (Drake) 

Serenthia rwindiana Drake 1956a, p. 7 [Congo]. 
Agramma rwindianum: Drake 1958a, p. 107. 
Type: Holotype o 71 ; lac Edouard, East Rwindi, Belgian Congo; Pares 
Nat. Inst. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Agramma scitulum Drake and Maa 

Agramma scitula Drake and Maa 1955, p. 11 [India]. 
Type: Holotype cf, brachypterous; Tanjore District, South India; 
Drake Coll. (USNM). 
Distribution: India. 
Host Plant: Unrecorded. 

*Agramma sedale (Drake) 

Serenthia sedalis Drake 1927d, p. 312 [Luzon]. — Drake and Poor 1937a, 
p. 2; 1937b, p. 397. 
Type: Holotype o", macropterous; Manila, Philippine Islands; USNM. 
Distribution: Philippine Islands (Luzon); Ceylon; Greater Sunda 
Islands (Java). 
Host Plant: Unrecorded. 

* Agramma singulum (Drake) 

Serenthia singula Drake 1954b, p. 14 [Cape Province]. — Drake and 

Slater 1955, p. 49 [Zululand]. 
Agramma singulum: Drake 1958a, p. 107. 
Type: Holotype d"; Somerset East, Cape Province, South Africa; 
British Mus. 
Distribution: South Africa (Zululand; Cape Province). 
Host Plant: Unrecorded. 



76 U.S. NATIONAL MUSEUM BULLETIN 243 

*Agramma sociale (Drake) 

Serenthia socialis Drake 1951, p. 169 [Tanganyika; Ethiopia]. 

Agramma sociale: Drake 1958a, p. 107 [Kenya]. 
Type: Holotype cf; Arusha-Ju, Africa orientalis; Hungarian Mus. 
Distribution: Tanganyika; Kenya; Ethiopia. 
Host Plant: Unrecorded. 

* Agramma striolum (Drake) 

Serenthia striola Drake 1951, p. 168. 

Agramma striola: Drake 1955d, p. 1 [Urundi; Ethiopia]. 
Type: Holotype o" ; Hungarian Mus.; type locality unknown. 
Distribution: Ruanda-Urundi; Ethiopia. 
Host Plant: Unrecorded. 

* Agramma tropidopterum Flor 

Agramma tropidopterum Flor 1860, p. 326 [Estonia]. 
Serenthia tropidoptera: Stal 1874, p. 47. — Horvath 1906a, p. Ill, pi. 1, 
fig. 1 [Russia]. — Stichel 1926, p. 117, fig. 308 [Juncus; Eriophorum; 
Germany]. — Kiritshenko 1951, p. 254. 
Scraulia tropidoptera: Lethierry and Severin 1896, p. 9. 
Serenthia (Paraserenthia) tropidoptera: Wagner 1941, pp. 6, 9, 26, figs. 

2a, 3a, 4a, 5a, 6a. 
Agramma tropidoptera: Stichel 1960a, p. 345, figs. 187, 188. 
Type: Estonia; sex and deposition of type unknown. 
Distribution: U.S.S.R. (Estonia; Latvia); Germany. 
Host Plants: Juncus sp.; Eriophorum sp. 

* Agramma turanicum (Horvath) 

Serenthia turanica Horvath 1905b, p. 556 [Turkestan]; 1906a, p. 113. 
Type: Sex unknown; Ilysk, Turkestan, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 

* Agramma umbrosum (Horvath) 

Serenthia umbrosa Horvath 1906a, p. 113 [Algeria]. 

Agramma umbrosum: Drake 1958a, p. 107. 
Type: Holotype cf; Madjez-Amar, Algeria; Hungarian Mus. 
Distribution: Algeria; West Africa; Senegal. 
Host Plant: Unrecorded. 

Agramma vanderysti (Schouteden) 

Wombalia vanderysti Schouteden 1919, p. 140 [Congo]. 

Serenthia vanderysti: Drake 1954b, p. 13. 

Agramma vanderysti: Drake 1958a, p. 107. 
Type: Sex unknown; Wombali, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 77 

*Agramma vicinale (Drake) 

Serenthia vicinalis Drake 1927d, p. 311 [Luzon]; 1930e, p. 166.— Drake 
and Poor 1937a, p. 1. 
Type: Holotype o 71 , macropterous; Mount Maquiling, Luzon, Philippine 
Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

*Agramma vulturnum (Kirkaldy) 

Serenthia vulturna Kirkaldy 1908a, p. 778 [Queensland]. — Hacker 1927, 
p. 20, fig. 17. 
Type: Holotype o 71 ; Bundaberg, Queensland, Australia; Hawaii. Sugar 
Plant. Assn. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Genus AIDONEUS Distant 

Aidoneus Distant 1909a, p. 122; 1910a, p. 125.— Drake and Runoff 
1960a, p. 34. 
Type Species: Aidoneus dissimilis Distant. 

Aidoneus dissimilis Distant 

Aidoneus dissimilis Distant 1909a, p. 123 [India]; 1910a, p. 126, fig. 66. 
Type: Sex unknown; Bengal, India; British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 

Genus ALLOEOCYSTA Drake 

Alloeocysta Drake 1961b, p. 109. 
Type Species: Alloeocysta approba Drake. 

Alloeocysta approba Drake Plate 12 

Alloeocysta approba Drake 1961b, p. 109, pi. 5 [New South Wales]. 

Type: Holotype o 71 ; Bogan River, New South Wales, Australia; Austr. 
Mus. 

Distribution: Australia (New South Wales). 

Host Plant: Unrecorded. 

Genus ALLOIOTHUCHA Drake 

Alloiothucha Drake 1927a, p. 58.— Drake and Poor 1937a, p. 18 (in part); 
1939c, p. 207 (in part).— Drake and Ruhoff 1960a, p. 34. 
Type Species: Alloiothucha philippinensis Drake. 



78 U.S. NATIONAL MUSEUM BULLETIN 243 

*Alloiothucha artocarpi (Horvath) 

Holophygdon artocarpi Horvath 1926, p. 327, figs. 1-2 [Java; Artocarpus 

integrifolia]. — Poisson 1951, p. 1798, fig. 1591a (fig. only). 
Alloiothucha artocarpi: Drake and Poor 1939c, p. 207. 
Type: Holotype 9; Buitenzorg, Java; Hungarian Mus. 
Distribution: Greater Sunda Islands (Java; Sumatra). 
Host Plant: Artocarpus integrifolia. 

*Alloiothucha necopinata Drake 

Alloiothucha necopinata Drake 1927a, p. 59 [Palawan]. — Drake and Poor 

1939c, p. 207. 
Holophygdon necopinata: Drake and Poor 1937a, p. 18. 
Type: Holotype cf, macropterous; Puerto Princesa, Palawan, Philippine 
Islands; Drake Coll. (USNM). 

Distribution: Philippine Islands (Palawan); Greater Sunda Islands 
(Borneo). 
Host Plant: Unrecorded. 

*Alloiothucha philippinensis Drake 

Alloiothucha philippinensis Drake 1927a, p. 58 [Luzon]. — Drake and 

Poor 1939c, p. 207, fig. 1. 
Holophygdon philippinensis: Drake and Poor 1937a, p. 18. 
Type: Holotype o 71 , macropterous; Mount Maquiling, Luzon, Philippine 
Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

Genus ALLOTINGIS Drake 

Allotingis Drake 1930d, p. 269.— Hurd 1946, p. 471— Drake and RuhofF 
1960a, p. 35. 
Type Species: Leptobyrsa binotata Drake and Bruner. 
* Allotingis binotata (Drake and Bruner) 

Leptobyrsa binotata Drake and Bruner 1924b, p. 155 [Cuba; Thrinax 

wendlandiana]. 
Allotingis binotata: Drake 1930d, p. 270. 
Type: Holotype o 71 , macropterous; La Machorra, Peninsula de Guana- 
hacabibes, Cuba; Drake Coll. (USNM). 
Distribution: Cuba. 
Host Plant: Thrinax wendlandiana. 
*AUotingis insulicola Drake and Poor 

Allotingis insulicola Drake and Poor 1939a, p. 31 [Haiti]. 
Type: Holotype o 71 , macropterous; Fond-des-Negres, Haiti; Drake Coll. 
(USNM). 
Distribution: Haiti. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 79 

Genus ALVEOTINGIS Osborn and Drake 

Alveotingis Osborn and Drake 1916a, p. 245; 1917b, p. 305 (key). — Van 
Duzee 1917b, pp. 221, 818.— Parshley 1917a, p. 24; 1923b, p. 707 — 
Blatchley 1926, p. 486.— Hurd 1946, p. 445.— Bailey 1951, p. 20 — 
Drake and Runoff 1960a, p. 35. 
Type Species: Alveotingis grossocerata Osborn and Drake. 

*Alveotingis brevicornis Osborn and Drake 

Alveotingis brevicornis Osborn and Drake 1917b, p. 305, fig. 2c [Iowa]. — 
Froeschner 1944, p. 670 [Mo.]. 
Type: Holotype 9, macropterous; Little Rock, Iowa, U.S.; Drake Coll. 
(USNM). 

Distribution: U.S. (Iowa; Mo.; Minn.). 
Host Plant: Unrecorded. 

*Alveotingis grossocerata Osborn and Drake 

Alveotingis grossocerata Osborn and Drake 1916a, p. 245, fig. 9 [Maine]; 
1917b, pp. 305, 306, fig. 2a [N.H.].— Van Duzee 1917b, p. 221 
[Pa.].— Parshley 1917a, p. 25; 1917b, p. 57; 1920a, p. 274; 1922a, 
p. 236 [Conn.]; 1923b, p. 707.— Blatchley 1926, p. 486, fig. 113. — 
Drake 1928b, p. 103 [N.Y.; thisde] .— Torre-Bueno 1931, p. 149 
[Mass.].— Bailey 1951, p. 20.— Drake and Davis 1960, fig. 67. 
Type: Holotype o 71 , brachypterous; Orono, Maine, U.S.; Drake Coll. 
(USNM). 

Distribution: U.S. (Maine; N.H.; Mass.; Conn.; N.Y.; Pa.; Kans.). 
Host Plant: Unrecorded. 
Note: Netted on thistle. 

*Alveotingis minor Osborn and Drake 

Alveotingis minor Osborn and Drake 1917b, p. 305, fig. 2h [Iowa]. 

Type: Holotype cf, brachypterous; Ames, Iowa, U.S.; Drake Coll. 
(USNM). 

Distribution: U.S. (Iowa). 

Host Plant: Unrecorded. 

Genus AMBLYSTIRA Stal 

Amblystira Stal 1873, pp. 120, 129.— Champion 1897, p. 29 (key).— 
Monte 1939b, p. 65; 1941e, p. 77.— Hurd 1946, p. 455.— Drake and 
Ruhoff 1960a, p. 35. 
Type Species: Monanthia pallipes Stal. 

*Amblystira arnica Drake and Ruhoff 

Amblystira solida Drake 1942a, p. 17 (not p. 16) [Haiti]. 
Amblystira dozieri Drake and Hambleton 1944a, p. 94. 
Amblystira arnica Hurd 1946, p. 456 (nom. nud.). — Drake and Ruhoff 
1960c, p. 31. 



80 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Type: Holotype 9, macropterous; Hinche, Haiti; Drake Coll. (USNM). 
Distribution: Haiti. 
Host Plant: Unrecorded. 

Amblystira angella Drake and Runoff 

Amblystira angella Drake and Runoff 1962c, p. 137, fig. 1 [Peru]. 

Type: Holotype 9, macropterous; Monson Valley, Tingo Maria, Peru; 
Cal. Acad. 

Distribution: Peru. 

Host Plant: Unrecorded. 

* Amblystira atrinervis Champion 

Amblystira atrinervis Champion 1897, p. 31, pi. 2, fig. 24 [Mexico]. 
Type: Holotype o"; Atoyac, Vera Cruz, Mexico; British Mus. 
Distribution: Mexico. 
Host Plant: Unrecorded. 

*Amblystira fuscitarsis Champion 

Amblystira juscitar sis Champion 1897, p. 30, pi. 2, figs. 21-22 [Guate- 
mala; Panama]. — Drake and Bruner 1924a, p. 148 [Cuba]. — Monte 
1938b, p. 128 [Brazil]. — Bruner, Scaramuzza and Otero 1945, p. 100 
[Lonchocarpus sericeus]. — Drake and Hambleton 1945, p. 358 [Colom- 
bia; Haiti; Denis elliptica]. 
Type: Sex unknown; Volcan de Atitlan, Guatemala; British Mus. 
Distribution: Mexico; Guatemala; Panama; Colombia; Brazil; Haiti; 
Cuba. 

Host Plants: Denis elliptica; Longchocarpus sericeus. 

*Amblystira laevifrons Champion 

Amblystira laevifrons Champion 1897, p. 31, pi. 2, fig. 25 [Mexico]. 
Type: Sex unknown; Atoyac, Vera Cruz, Mexico; British Mus. 
Distribution: Mexico. 
Host Plant: Unrecorded. 

*Amblystira machalana Drake 

Amblystira machalana Drake 1948c, p. 22 [Ecuador]. 
Type: Holotype d 1 , macropterous; Machala, Ecuador; Drake Coll. 
(USNM). 
Distribution: Ecuador; Peru. 
Host Plants: Teak; cacao. 

Amblystira maculata Van Duzee 

Amblystira maculata Van Duzee 1907, p. 21 [Jamaica]. — Drake and 
Bruner 1924a, p. 148, fig. 2.— Gowdy 1926, p. 35. 
Type: Sex unknown; Richmond Village, Jamaica; Cal. Acad. 
Distribution: Jamaica. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 81 

* Amblystira marginata Drake 

Amblystira marginata Drake 1922c, p. 362, pi. 39, fig. 7 [Canal Zone]. — 
Hurd 1946, p. 456 [Costa Rica]. 
Type: Holotype d", brachypterous; Paraiso, Panama Canal Zone; 
USNM. 
Distribution : Panama (Canal Zone); Costa Rica. 
Host Plant: Unrecorded. 

* Amblystira melanosoma Monte 

Amblystira melanosoma Monte 1941d, p. 95, fig. 2 [Costa Rica]. 
Type: Holotype 9; San Isidro de Coronado, Alajuela, Costa Rica; 
Monte Coll. (Mus. Nacional). 
Distribution: Costa Rica. 
Host Plant: Unrecorded. 

* Amblystira morrisoni Drake 

Amblystira morrisoni Drake 1922c, p. 360 [Dominican Republic]. — Drake 
and Bruner 1924a, p. 148. 
Type: Holotype o 71 , macropterous; San Domingo City, San Domingo; 
USNM. 
Distribution: Dominican Republic. 
Host Plant: Unrecorded. 

* Amblystira nyctalis Drake 

Amblystira nyctalis Drake 1922c, p. 360 [Brazil]. 
Type: Holotype o 71 ; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Amblystira opaca Champion 

Amblystira opaca Champion 1897, p. 30, pi. 2, fig. 23 [Guatemala]. — 
Hurd 1946, p. 456 [Panama]. 
Type: Sex unknown; San Isidro, Guatemala; British Mus. 
Distribution: Guatemala; Panama; Nicaragua. 
Host Plant: Denis elliptica. 

* Amblystira pallipes (Stal) 

Monanthia (Tropidocheila) pallipes Stal 1858, p. 62 [Brazil]. 
Monanthia pallipes: Walker 1873a, p. 192. 

Amblystira pallipes: Stal 1873, p. 129.— Champion 1898b, p. 61, pi. 2, 
fig. 12.— Drake and Hambleton 1935, p. 141 [Serjania]; 1944b, p. 124 
[Venezuela]; 1945, p. 358 [Peru].— Drake and Poor 1937d, p. 304 
[Colombia].— Monte 1938b, p. 128; 1939b, p. 65 [Bredemeyera]. 
Type: Sex unknown; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil; Colombia; Peru; Venezuela. 
Host Plants: Serjania sp.; Bredemeyera sp. 



82 U.S. NATIONAL MUSEUM BULLETIN 243 

* Amblystira peltogyne Drake and Hambleton 

Amblystira peltogyne Drake and Hambleton 1935, p. 141, fig. 1 [Brazil; 
Peltogyne]. — Monte 1939b, p. 65 [Hymenaea stigonocarpa]. 
Type: Holotype o", macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plants: Peltogyne sp.; Hymenaea stigonocarpa. 

* Amblystira pensa Drake and Hambleton 

Amblystira pensa Drake and Hambleton 1939, p. 155 [Brazil]. — Monte 
1941e, p. 78 [Roupala]. 
Type: Holotype d\ macropterous; Belo Horizonte, Minas Gerais, 
Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Roupala sp. 

*Amblystira scita Drake and Hambleton 

Amblystira scita Drake and Hambleton 1944b, p. 124 [Costa Rica]. 

Type: Holotype 9, macropterous; Mercedes, Costa Rica; Drake Coll. 
(USNM). 

Distribution: Costa Rica. 

Host Plant: Unrecorded. 

* Amblystira silvicola Drake 

Amblystira silvicola Drake 1922c, p. 361, pi. 39, fig. 6 [Bolivia; Brazil; 
British Guiana]; 1935, p. 11 [Paraguay]; 1936b, p. 699 [Argentina].— 
Monte 1938b, p. 128; 1939b, p. 65 [Serjania; Bignoniaceae]. — Silva 
1956, p. 18, fig. 3. 
Type: Holotype cf ; Rio Machupe, Bolivia; Carnegie Mus. 
Distribution: Bolivia; Brazil; British Guiana; Argentina; Paraguay. 
Host Plants: Serjania sp.; Bignoniaceae. 
*Amblystira socia Drake 

Amblystira socia Drake 1942a, p. 17 [Paraguay]. 
Type: Holotype o 71 , macropterous; Chaco, Paraguay; Drake Coll. 
(USNM). 
Distribution: Paraguay; Peru; Bolivia. 
Host Plant: Unrecorded. 

*Amblystira solida Drake 

Amblystira solida Drake 1942a, p. 16 [Brazil]. — Drake and Davis 1960, 
fig. 66. 
Type: Holotype 9, macropterous; Guaruja, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 83 

Genus AMBOTINGIS Drake and Ruhoff 

Ambotingis Drake and Runoff 1960c, p. 29. 
Type Species: Monanthia senta Drake and Hambleton. 
*Ambotingis senta (Drake and Hambleton) Plate 10 

Monanthia senta Drake and Hambleton 1942a, p. 329 [Peru]; 1945, 

p. 356 [Ecuador]. 
Ambotingis senta: Drake and Ruhoff 1960c, p. 31, fig. 1; 1960e, p. 80. 
Type: Holotype cf , macropterous; Sullena, Peru; Drake Coll. (USNM). 
Distribution: Peru; Ecuador. 
Host Plant: Unrecorded. 

Genus AMBYCYSTA Drake and Hurd 

Ambycysta Drake and Hurd 1945b, p. 129.— Hurd 1946, p. 475.— Drake 
and Ruhoff 1960a, p. 36. 
Type Species: Megalocysta championi Drake. 

* Ambycysta championi (Drake) 

Megalocysta championi Drake 1922b, p. 38 [Brazil]. — Monte 1941e, 

p. 123; 1942c, p. 302. 
Ambycysta championi: Drake and Hurd 1945b, p. 130. 
Type: Holotype cf, macropterous; Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

Note: This species is known only from the holotype, which was errone- 
ously recorded as a female in the original description. 
Ambycysta cornuta (Monte) 

Megalocysta cornuta Monte 1942c, p. 302, figs. 1-3 [Peru]. 
Ambycysta cornuta: Drake and Hurd 1945b, p. 130. 
Type: Holotype o"; Hacienda San Juan, Col. Perene, Peru; Monte 
Coll. (Mus. Nacional). 
Distribution: Peru. 
Host Plant: Unrecorded. 

* Ambycysta enodis Drake 

Ambycysta enodis Drake 1948j, p. 430. 
Type: Holotype 9, macropterous; Pachitea, Peru; Drake Coll. (USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 

* Ambycysta gibbifera (Picado) 

Leptostyla gibbifera Picado 1913, pp. 303, 306, pi. 13, fig. 5; figs. 29, 
30 [Costa Rica; Aechmea]. 

Megalocysta gibbifera: Drake 1928d, p. 23.— Monte 1942c, p. 302. 

Ambycysta gibbifera: Drake and Hurd 1945b, p. 130. 
Type: Sex unknown; Orosi, Costa Rica; Paris Mus. 
Distribution: Costa Rica. 
Host Plant: Aechmea sp. 



84 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Genus AMMIANUS Distant 

Ammianus Distant 1903b, p. 136; 1910a, p. 115. — Drake 1955d, p. 5; 

1957c, p. 31.— Drake and Runoff 1960a, p. 36. 
Monanthia (Phyllontocheila) Fieber 1844, p. 59 (in part). 
Phyllontochila (emendation): Stal 1873, pp. 120, 128— Horvath 1909b, 

p. 632; 1910, p. 62. 
Sakuntala Kirkaldy 1902, p. 298.— Horvath 1909b, p. 632; 1910, p. 62.— 

Drake and Runoff 1960a, p. 80. 
Phyllontocheila: Horvath 1911a, p. 327 (key). — Schouteden 1923, p. 95. — 

Drake and Ruhoff 1960a, p. 76. 
Phyllontocheila (Kitoko) Schouteden 1923, p. 95. — Drake 1955c, p. 105. — 

Drake and Ruhoff 1960a, p. 63. 
Type Species: Monanthia (Phyllontocheila) erosa Fieber. 

* Ammianus admirandus (Drake) 

Phyllontocheila admiranda Drake 1931d, p. 96 [Borneo; Malaya]; 1934b, 
p. 107, fig. 

Ammianus admirandus: Drake 1955d, p. 5. 
Type: Sex unknown; Borneo; Paris Mus. 

Distribution: Greater Sunda Islands (Borneo); Federation of Malaya. 
Host Plant: Unrecorded. 

* Ammianus alaticollis (Stal) 

Phyllontocheila alaticollis Stal 1855a, p. 37 [Natal]. — Horvath 1911a, 
pp. 329, 331. 

Monanthia (Phyllontochila) alaticollis: Stal 1865, p. 27 [Caffraria]. 

Monanthia alaticollis: Walker 1873a, p. 195. 

Phyllontochila alaticollis: Stal 1873, p. 128. — Distant 1902a, p. 241, 
pi. 15, fig. 5. 

Phyllontochila (Sinuessa) alaticollis: Horvath 1910, p. 63 [Tanganyika]. 

Ammianus alaticollis: Drake 1957c, p. 32. 
Type: Sex unknown; Natal, South Africa; Stockholm Mus. 
Distribution: South Africa (Natal); Tanganyika; Mozambique. 
Host Plant: Unrecorded. 

* Ammianus alberti (Schouteden) Plates 36, 37 

Phyllontocheila alberti Schouteden 1916a, p. 275 [Congo]; 1916b, p. 293. — 

Drake 1954e, p. 4. 
Phyllontocheila (Kitoko) alberti: Schouteden 1923, p. 96.- — Mayne and 

Ghesquiere 1934, p. 20 {Vernonia conjerta; cacao]. 
Kitoko alberti: Schouteden 1953d, p. 170. 
Ammianus alberti: Drake 1955d, p. 5. 
Type: Sex unknown; Uele, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plants: Vernonia conjerta; cacao. 



LACEBUGS OF THE WORLD 85 

*Ammianus alberti var. tricarinatus (Schouteden) 

Kitoko alberti var. tricarinata Schouteden 1953d, p. 170 [Congo]. 
Phyllontocheila alberti subsp. tricarinata: Drake and Gomez-Menor 1954, 
p. 93 [Spanish Guinea]. — Drake 1955c, p. 105 [Cameroun; Vernonia]. 
Ammianus alberti var. triseriatus [sic]: Drake 1955d, p. 5. 
Piesma pupla (not Puton): Gomez-Menor 1956a, p. 119, fig. 15 (fig. 
only). 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Spanish Guinea; Cameroun. 
Host Plant: Vernonia sp. 

Ammianus basilewskyi Schouteden 

Ammianus basilewskyi Schouteden 1957c, p. 315 [Ruanda]. 
Type: Sex unknown; Ruanda; Cent. Afr. Mus. 
Distribution : Ruanda-Urundi. 
Host Plant: Unrecorded. 

* Ammianus bobangensis (Schouteden) 

Phyllontocheila bobangensis Schouteden 1923, p. 96 [Congo]. 

Ammianus bobangensis: Drake 1955d, p. 5; 1958b, p. 29. 
Type: Sex unknown; Bolobo, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

* Ammianus burgeoni (Schouteden) 

Phyllontocheila burgeoni Schouteden 1923, p. 97 [Congo]. 
Type: Sex unknown; Mabao, Haut Uele, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Ammianus corticinus (Horvath) 

Phyllontocheila corticina Horvath 1911a, pp. 328, 329 [Cameroun]. — 

Schouteden 1923, p. 99 [Congo]. 
Ammianus corticinus: Drake 1955d, p. 5. 
Type: Sex unknown; Vallis N'Goko, Cameroun; Hungarian Mus. 
Distribution: Cameroun; Congo. 
Host Plant: Unrecorded. 

* Ammianus depictus (Schouteden) 

Phyllontocheila burgeoni var. depicta Schouteden 1923, p. 98 [Congo]. 

Ammianus depictus: Drake and Ruhoff 1962a, p. 156. 
Type: Sex unknown; Belgian Congo; Cent. Afr. mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 



86 U.S. NATIONAL MUSEUM BULLETIN 243 

* Ammianus dilatatus (Guerin-Meneville) Plate 38 

Tingis dilatata Guerin-Meneville 1831, p. 8, fig. [Senegal]. — Stal 1873, 

p. 134. 
Phyllontochila dilatata: Distant 1902a, p. 241 [West Africa; Rhodesia]; 

1902b, p. 355 [Sierra Leone]. 
Phyllontocheila dilatata: Horvath 1911a, pp. 328, 331. — Drake 1954a, 
p. 232 [Ivory Coast]; 1954e, p. 4 [Congo]; 1955b, p. 90 [Angola;] 
1955c, p. 105 [Cameroun; Nigeria]. 
Ammianus dilatatus: Drake 1955d, p. 5. 
Type: Senegal; sex and deposition of type unknown. 
Distribution: Senegal; Sierra Leone; Guinea; Ivory Coast; Ghana; 
Nigeria; Cameroun; Spanish Guinea; Angola; Congo; Rhodesia; West 
Africa. 
Host Plant: Unrecorded. 

Ammianus dilatatus var. obscurus (Schumacher) 

Phyllontocheila dilatata var. obscura Schumacher 1912, p. 320 [Cameroons]. 
Type: Barombi, Cameroons; sex and deposition of type unknown. 
Distribution: Cameroons. 
Host Plant: Unrecorded. 

* Ammianus elisabethae (Schouteden) 

Phyllontocheila elisabethae Schouteden 1916a, p. 274 [Congo]; 1916b, 
p. 293; 1923, p. 99 [Tanganyika]. 

Ammianus elisabethae: Drake 1955d, p. 5. 
Type: Sex unknown; Belgian Congo; Cent. Air. Mus. 
Distribution: Congo; Tanganyika. 
Host Plant: Unrecorded. 

* Ammianus erosus (Fieber) 

Monanthia (Phyllontocheila) erosa Fieber 1844, p. 71, pi. 6, figs. 5-9 

[Ostindien]. — Herrich-Schaeffer 1850, p. 155. 
Tingis erosa: Stal 1870a, p. 671 [Philippines]. — Walker 1873a, p. 181 

[India]. 
Phyllontocheila erosa: Stal 1873, p. 128 [Penang]. — Distant 1902b, 
p. 355.— Drake and Poor 1937a, p. 15 [Luzon]. — Drake 1937a, p. 387 
[Sumatra]; 1938b, p. 196 [China]. 
Ammianus erosus: Distant 1903b, p. 137, fig. 100 [Burma]. — Drake 1955d, 

p. 5; 1957c, p. 32. 
Phyllontocheila erosa: Blote 1945, p. 92. 
Type: Sex unknown; "Ostindien"; Vienna Mus. 

Distribution: India; Burma; Thailand; Penang; China; Greater Sunda 
Islands (Sumatra; Java); Philippine Islands (Luzon). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 87 

* Ammianus flabilis (Bergroth) 

Phyllontochila flabilis Bergroth 1894, p. 167 [Malagasy]. 

Ammianus flabilis: Drake 1955d, p. 5. 

Ammianus flavilis [sic]: Drake 1957f, p. 131. 
Type: Sex unknown; Madagascar; Helsin. Mus. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

* Ammianus ghesquierei (Schouteden) 

Phyllontocheila ghesquierei Schouteden 1923, p. 99 [Congo]. — Drake 
1954e, p. 4. 
Type: Sex unknown; Luebo, Kasai, Belgian Congo; Cent. Air. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Ammianus junodi (Distant) 

Phyllontochila junodi Distant 1904, p. 431, pi. 8, fig. 10 [Transvaal]. 
Phyllontochila (Sinuessa) junodi: Horvath 1910, p. 64 [Tanganyika]. 
Phyllontocheila junodi: Horvath 1911a, pp. 329, 332. — Drake 1956f, p. 424 

[Cape Province; Uganda; Kenya; Natal; Zanzibar]. 
Ammianus janodi [sic]: Drake 1957c, p. 32. 
Type: Sex unknown; Shilouvane, Zoutpansberg District, Transvaal, 
South Africa; British Mus. 

Distribution: South Africa (Transvaal; Natal; Cape Province); Uganda; 
Kenya; Tanganyika; Zanzibar. 
Host Plant: Unrecorded. 

*Ammianus kassianoffi Drake 

Ammianus kassianoffi Drake 1955d, p. 4 [Congo; Uganda]. 
Type: Holotype o 71 ; Lukolo, Belgian Congo; Brussels Mus. 
Distribution: Congo; Uganda. 
Host Plant: Unrecorded. 

*Ammianus laminatus (Horvath) 

Phyllontocheila laminata Horvath 1911a, pp. 328, 330 [Uganda]. — 
Schouteden 1916b, p. 293 [Congo]; 1923, p.100— Drake and Gomez- 
Menor 1954, p. 93 [Cameroun]. 
Ammianus laminatus: Drake 1955d, p. 5. 
Type: Entebbe, Uganda; sex and deposition of type unknown. 
Distribution: Uganda; Cameroun; Congo; Tanganyika; Kenya. 
Host Plant: Unrecorded. 

Ammianus maseruanus (Drake) 

Phyllontocheila maseruana Drake 1956f, p. 424 [Basutoland]. 
Type: Holotype o 71 ; Mount Machache, Basutoland; Lund Zool. Inst. 
Distribution: Basutoland. 
Host Plant: Unrecorded. 



88 U.S. NATIONAL MUSEUM BULLETIN 243 

*Ammianus mayri (Haglund) 

Phyllontochila mayri Haglund 1895, p. 471 [Cameroun]. — Distant 1902a, 

p. 241 [West Africa]. 
Phyllontocheila mayri: Horvath 1911a, p. 331 [Dahomey]. 
Ammianus mayri: Drake 1955d, p. 5. 
Type: Sex unknown; Cameroun; Stockholm Mus. 
Distribution: Cameroun; Dahomey; West Africa. 
Host Plant: Unrecorded. 

*Ammianus mayri subsp. septuosus Drake and Ruhoff 

Phyllontocheila mayri var. obscura Schouteden 1916a, p. 276 [Congo]. 

Ammianus mayri var. obscura: Drake 1955d, p. 5. 

Ammianus mayri subsp. septuosus Drake and Ruhoff 1962a, p. 161. 

Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 

Distribution: Congo; Nigeria; Sierra Leone. 

Host Plant: Unrecorded. 

* Ammianus mussolinii (Mancini) 

Phyllontocheila mussolinii Mancini 1939a, p. 214, tab. 5, fig. 2 [Ethiopia]. 

Ammianus amoenus Drake 1956b, p. 112 [Kenya]. 

Ammianus mussolinii: Drake and Ruhoff 1961c, p. 145. 
Type: Holotype c? ; Neghelli, Ethiopia; Mancini Coll. 
Distribution: Ethiopia; Kenya. 
Host Plant: Unrecorded. 

* Ammianus perakensis (Distant) 

Phyllontochila perakensis Distant 1902b, p. 355 [Malaya]. 

Phyllontocheila perakensis: Blote 1945, p. 92 [Sumatra]. 

Ammianus par akensis [sic]: Drake 1955d, p. 5. 

Ammianus perakensis: Drake 1957c, p. 32. 
Type: Sex unknown; Perak, Malay Peninsula; British Mus. 
Distribution: Federation of Malaya; Greater Sunda Islands (Sumatra; 
Sarawak). 
Host Plant: Unrecorded. 

* Ammianus philippinensis (Distant) 

Phyllontochila philippinensis Distant 1902b, p. 355 [Philippines]. — Drake 

and Poor 1937a, p. 15 [Luzon]. 
Phyllontocheila philippinensis: Drake and Poor 1937b, p. 403. 
Ammianus philippinensis : Drake 1957c, p. 32. 
Type: Sex unknown; Philippine Islands; British Mus. 
Distribution: Philippine Islands (Luzon; Mindanao); Greater Sunda 
Islands (Java). 
Host Plant: Unrecorded. 



LACE BUGS OF THE WORLD 89 

*Ammianus ravanus (Kirkaldy) 

Sakuntala ravana Kirkaldy 1902, p. 299 [Ceylon].— Blote 1945, p. 86 

[Lombok]. 
Phyllontocheila ravana: Distant 1903a, p. 51; 1903b, p. 136, fig. 99 
[Vitrex trifolia]; 1910a, p. 115 [India].— Drake 1926e, p. 335 [Vitrex 
pubescens; Java]; 1933, p. 1016.— Drake and Poor 1936b, p. 146; 
1937a, p. 15 [Singapore; Philippines].— Singh 1953, p. 118. 
Ammianus ravanus: Drake 1957c, p. 31. 
Type: Peradeniya, Ceylon; sex and deposition of type unknown. 
Distribution: India; Ceylon; Singapore; Lesser Sunda Islands (Lom- 
bok); Greater Sunda Islands (Java; Sumatra); Philippine Islands. 
Host Plants: Vitrex pubescens; Vitrex trifolia. 

* Ammianus schoutedeni Distant 

Ammianus schoutedeni Distant 1908a, p. 220 [Congo]. — Drake 1955d, 

p. 5. 
Phyllontocheila schoutedeni: Horvath 1911a, p. 333. — Schouteden 1916a, 
p. 276; 1916b, p. 293; 1923, p. 101.— Mayne and Ghesquiere 1934, 
p. 20 [Triumfetta rhombeaefolia]. — Drake and Gomez-Menor 1954, 
p. 93 [Rio Muni]. 
Stephanitis pyrioides (not Scott): Gomez-Menor 1956a, p. 117, fig. 12 
(fig. only). 
Type: Sex unknown; Popocabacca, Belgian Congo; British Mus. 
Distribution: Congo; Rio Muni; Ghana; Liberia. 
Host Plant: Triumfetta rhombeaefolia. 

* Ammianus spinosus (Schouteden) 

Phyllontocheila spinosa Schouteden 1923, p. 100 [Congo]. 

Ammianus spinosus: Drake 1955d, p. 5. 
Type: Sex unknown; Madyu, Haut Uele, Belgian Congo; Cent. Afr. 
Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Ammianus tellinii (Schouteden) 

Phyllontocheila tellinii Schouteden 1916a, p. 273 [Eritrea]. 
Type: Sex unknown; Adi LTgri, Eritrea; Cent. Afr. Mus. 
Distribution: Eritrea. 
Host Plant: Unrecorded. 
: ' Ammianus toi (Drake) 

Phyllontochila toi Drake 1938b, p. 196 [China]; 1947d, p. 228 [Viet- 

Nam]. 
Ammianus toi: Drake 1955d, p. 5; 1957c, p. 32. 
Type: Holotype d"; Canton, P'an-yu District, China; Lingnan Surv. 
Distribution: China; Viet-Nam; Greater Sunda Islands (Java). 
Host Plant: Unrecorded. 

681-552—64 7 



90 U.S. NATIONAL MUSEUM BULLETIN 243 

Ammianus urundicus Schouteden 

Ammianus urundicus Schouteden 1957c, p. 313 [Urundi]. 
Type: Sex unknown; Urundi; Cent. Afr. Mus. 
Distribution : Ruanda-Urundi. 
Host Plant: Unrecorded. 
*Ammianus vanderijsti (Schouteden) 

Phyllontocheila vanderijsti Schouteden 1923, p. 101 [Congo]. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Kenya; Tanganyika. 
Host Plant: Unrecorded. 
Ammianus vicinus Schouteden 

Ammianus vicina Schouteden 1957c, p. 315 [Ruanda]. 
Type: Sex unknown; Ruanda; Cent. Afr. Mus. 
Distribution : Ruanda-Urundi. 
Host Plant: Unrecorded. 
* Ammianus wahlbergi (Stal) 

Phyllontocheila wahlbergi Stal 1855a, p. 37 [Natal]. — Horvath 1911a, 
p. 331.— Schouteden 1923, p. 103 [Congo].— Mancini 1939a, p. 214 
[Ethiopia].— Drake 1954a, p. 232 [Guinea]. 
Monanthia (Phyllontochila) wahlbergi: Stal 1865, p. 27 [Caffraria]. 
Monanthia wahlbergi: Walker 1873a, p. 195. 

Phyllontochila wahlbergii [sic]: Stal 1873, p. 128.— Distant 1902a, p. 241. 
Type: Sex unknown; Natal, South Africa; Stockholm Mus. 
Distribution: South Africa (Natal); Nigeria; Guinea; Congo; Ethiopia. 
Host Plant: Unrecorded. 

Genus ANGOLOTINGIS Drake 

Angolotingis Drake 1955b, p. 88. — Drake and Ruhoff 1960a, p. 37. 
Type Species: Angolotingis vilhenai Drake. 
*Angolotingis vilhenai Drake 

Angolotingis vilhenai Drake 1955b, p. 88, fig. 2 [Angola]; 1958a, p. 104, 
fig. 1. 
Type: Holotype o 71 ; Serra do Moco, Luimbale, Angola; Cent. Afr. Mus. 
Distribution: Angola; Southern Rhodesia. 
Host Plant: Unrecorded. 

Genus ANGOLUSA Drake 

Angolusa Drake 1958a, p. 104.— Drake and Ruhoff 1960a, p. 37. 
Type Species: Angolusa machadoi Drake. 
Angolusa machadoi Drake 

Angolusa machadoi Drake 1958a, p. 105. 
Type: Holotype 9; Vila Luso, Angola; Cent. Afr. Mus. 
Distribution: Angola. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 91 

Genus APHELOTINGIS Drake 

Aphelotingis Drake 1948g, p. 176.— Drake and Ruhoff 1960a, p. 37. 
Type Species: Abdastarus muiri Drake. 

* Aphelotingis muiri (Drake) 

Abdastartus muiri Drake 1927d, p. 307 [Java; Singapore]. 

Aphelotingis muri [sic]: Drake 1948g, p. 176. 
Type: Holotype cf, macropterous; Pekalongan, Java; Drake Coll. 
(USNM). 
Distribution: Singapore; Greater Sunda Islands (Java). 
Host Plant: Unrecorded. 

* Aphelotingis perinetana Drake 

Aphelotingis perinetana Drake 1957f, p. 130 [Malagasy]. 
Type: Holotype 9; Perinet, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded, 

Aphelotingis tacsa Drake 

Aphelotingis tacsa Drake 1957f, p. 129 [Malagasy]. 
Type: Holotype cf; Perinet, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

Genus ARISTOBYRSA Drake and Poor 

Aristobyrsa Drake and Poor 1937c, p. 164. — Monte 1941e, p. 79. — 
Hurd 1946, p. 472.— Drake and Ruhoff 1960a, p. 37. 
Type Species: Leptobyrsa latipennis Champion. 

*Aristobyrsa latipennis (Champion) 

Leptobyrsa latipennis Champion 1897, p. 25, pi. 2, figs. 13, 13a [Panama]. 
Aristobyrsa latipennis: Drake and Poor 1937c, p. 164 [Brazil; Peru]. — 
Monte 1940b, p. 375; 1943a, p. 105; 1947b, p. 432 [Lucuma sp.].— 
Silva 1956, p. 19. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Panama; Brazil; Peru. 
Host Plant: Lucuma sp. 

Genus ARUSHIA Drake 

Arushia Drake 1951, p. 170.— Drake and Ruhoff 1960a, p. 37. 
Type Species: Arushia horvathi Drake. 

*Arushia connata Drake 

Arushia connata Drake 1951, p. 170 [Tanganyika; Kenya]. 
Type: Holotype 9; Arusha-Ju, Africa orientalis; Hungarian Mus. 
Distribution: Tanganyika; Kenya. 
Host Plant: Unrecorded. 



92 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Arushia horvathi Drake 

Arushia horvathi Drake 1951, p. 170 [Tanganyika]; 1954g, p. 663 
[Ethiopia; Congo]; 1956a, p. 5. 
Type: Holotype 9; Arusha-Ju, Africa orientalis; Hungarian Mus. 
Distribution: Tanganyika; Congo; Ethiopia; Guinea. 
Host Plant: Gramineae. 

Genus ATHEAS Champion 

Atheas Champion 1898a, p. 44.— Heidemann 1909, p. 231 (key). — 
Banks 1910, p. 55.— Van Duzee 1916, p. 26; 1917b, p. 222.— Blatch- 
ley 1926, p. 494 (key).— Monte 1939b, p. 65; 1941e, p. 80.— Hurd 
1946, p. 460.— Drake and Ruhoff 1960a, p. 38. 
Type Species: Atheas nigricornis Champion. 
*Atheas austroriparius Heidemann 

Atheas austroriparius Heidemann 1909, p. 235, fig. [Tex.; Fla.]. — Barber 
1914, p. 507.— Drake 1925c, p. 39 [Miss.; Desmodium].— Blatchley 
1926, p. 496, fig. 119c— Froeschner 1944, p. 670 [Mo.]. 
Type: Holotype 9, brachypterous; Duval, Florida, U.S.; USNM. 
Distribution: U.S. (Fla.; Miss.; Tex.; Mo.; S.C.); Mexico. 
Host Plants: Desmodium sp.; Schrankia sp. 
Atheas cearanus Monte 

Atheas cearana Monte 1947b, p. 430 [Brazil; Manihot]. 
Type: Holotype d"; Baturite, Ceara, Brazil; Monte Coll. (Mus. 
Nacional). 
Distribution: Brazil. 
Host Plant: Manihot sp. 
*Atheas exiguus Heidemann 

Atheas exiguus Heidemann 1909, p. 233, fig. [Fla.]. — Barber 1914, p. 
507.— Blatchley 1926, p. 496, fig. 119b; 1928, p. 5. 
Type: Holotype cf, macropterous; Sevenoaks, Florida, U.S.; USNM. 
Distribution: U.S. (Fla.; Miss.; Tex.). 
Host Plant: Unrecorded. 
*Atheas flavipes Champion 

Atheas flavipes Champion 1898a, p. 45, pi. 3, figs. 19, 19a [Panama]. — 
Drake and Hambleton 1938a, p. 46, fig. 7 [Brazil; Machaerium 
angustifolium]. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Panama; Brazil. 
Host Plant: Machaerium angustifolium. 
*Atheas fuscipes Champion 

Atheas Juscipes Champion 1898a, p. 45, pi. 3, fig. 20 [Mexico; Guate- 
mala]. — Drake and Hambleton 1934, p. 438 [Brazil; Legumino- 
sae].— Costa Lima 1936, p. 125.— Drake 1942a, p. 15 [Bolivia; 
Paraguay]. 



LACE BUGS OF THE WORLD 93 

Type: Sex unknown; Rio Naranjo, Guatemala; British Mus. 
Distribution: Mexico; Guatemala; El Salvador; Brazil; Bolivia; 
Paraguay. 

Host Plant: Leguminosae. 
*Atheas insignis Heidemann 

Atheas insignis Heidmann 1909, p. 232, fig. [Md.; D.C.]. — Osborn and 
Drake 1917b, p. 295 [Va.].— McAtee 1923, p. 146 [Stylosanthes 
biflora]. — Drake 1925c, p. 39 [Miss.; Desmodium]. — Blatchley 1926, 
p. 495, fig. 119a. 
Type: Holotype 9, macropterous; Bladensburg, Maryland, U.S.; 
USNM. 
Distribution: U.S. (Md.; D.C; Va.; Miss.). 
Host Plants: Desmodium sp.; Stylosanthes biflora. 
*Atheas laetantis Drake and Hambleton 

Atheas laetantis Drake and Hambleton 1944b, p. 124 [Brazil; Machaeri- 
um angustifolium; Machaerium]. 
Type: Holotype cf, macropterous; Vigosa, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plants: Machaerium angustifolium; Machaerium sp. 

* Atheas mimeticus Heidemann 

Atheas mimeticus Heidemann 1909, p 234, fig. [Wis.; Minn.; Kans.]. — 
Osborn and Drake 1917b, p. 295 [Colo.; N. Mex.].— Drake 1925c, 
p. 39 [Miss.; Desmodium].— Blatchley 1926, p. 496, fig. 119d. — 
Froeschner 1944, p. 670 [Mo.; Petalostemon purpureum]. — Hurd 1946, 
p. 460 [Wyo.]. 
Atheas annulatus Osborn and Drake 1917b, p. 295 [Ark.]. — Froeschner 

1944, p. 670. 
Atheas sordidus Osborn and Drake 1917b, p. 296 [Iowa]. 
Type: Holotype o 71 , brachypterous; Onaga, Kansas, U.S.; USNM. 
Distribution: U.S. (Wyo.; Colo.; N. Mex.; Nebr.; Kans.; Iowa; Minn.; 
Wis.; Mo.; Ark.; La.; Miss.; Fla.). 

Host Plants: Desmodium sp.; Petalostemon purpureum. 
*Atheas mirabilis Drake 

Atheas mirabilis Drake 1938a, p. 70 [Mexico]. 
Type: Holotype o 71 ; Temescaltepec, Mexico; Cal. Acad. 
Distribution: Mexico. 
Host Plant: Unrecorded. 

* Atheas nigricornis Champion 

Atheas nigricornis Champion 1898a, p. 45, pi. 3, fig. 21 [Mexico; Guate- 
mala].— Osborn and Drake 1915b, p. 536; 1917b, p. 296 [Ariz.].— 
Drake 1938a, p. 70. — Hurd 1946, p. 461 [Alnus acuminata; Parosela 
citriodora]. 



94 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Type: Sex unknown; Cerro Zunil, Guatemala; British Mus. 
Distribution: Ecuador; Honduras; Guatemala; Mexico; U.S. (Tex.; 
Ariz.). 
Host Plants: Alnus acuminata; Parosela citriodora. 

*Atheas ornatipes Drake and Hambleton 

Atheas ornatipes Drake and Hambleton 1935, p. 143 [Brazil; Aeschynomene 
falcata]. 
Type: Holotype o 71 , macropterous; Guaruja, Sao Paulo, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Aeschynomene falcata. 

*Atheas paganus Drake 

Atheas paganus Drake 1942a, p. 15 [Brazil; Aeschynomene], 
Type: Holotype o" , macropterous; Vicpsa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Aeschynomene sp. 

*Atheas placentis Drake and Poor 

Atheas fuscipes (not Champion): Monte 1939b, p. 66 [Celtis brasiliensis]. 
Atheas placentis Drake and Poor 1940, p. 226 [Brazil]. — Drake and 
Hambleton 1944b, p. 124. 
Type: Holotype 9, macropterous; Belo Horizonte, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Celtis brasiliensis. 

* Atheas tristis Van Duzee 

Atheas tristis Van Duzee 1923, p. 143 [Mexico; Aeschynomene nivea]. 
Type: Holotype cf; Concepcion Bay, Lower California, Mexico; Cal. 
Acad. 
Distribution: Mexico. 
Host Plant: Aeschynomene nivea. 

Genus AULOTINGIS Drake and Poor 

Aulotingis Drake and Poor 1943, p. 194. — Drake and Ruhoff 1960a, 
p. 38. 
Type Species: Aulotingis moalae Drake and Poor. 

Aulotingis moalae Drake and Poor 

Aulotingis moalae Drake and Poor 1943, p. 195, fig. 2 [Fiji]. 
Type: Holotype cf; Naroi, Moala Island, Fiji Islands; Bishop Mus. 
Distribution: Fiji Islands (Moala). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 95 

Genus AUSTRALOTINGIS Hacker 

Australotingis Hacker 1927, p. 29.— Drake and Ruhoff 1960a, p. 38. 
Type Species: Australotingis franzeni Hacker. 

♦Australotingis franzeni Hacker Plates 32, 33 

Australotingis franzeni Hacker 1927, p. 29, pi. 10, fig. 15 [Queensland]. 
Type: Holotype 9; Nanango, Queensland, Australia; Queensland Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

♦Australotingis vinnula Drake 

Australotingis vinnula Drake 1953c, p. 152 [Queensland]. 
Type: Holotype o" ; Lankelly Creek, Cape York, Queensland, Australia; 
MCZ. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Genus AXIOKERSOS Distant 

Axiokersos Distant 1909a, p. 121; 1910a, p. 124.— Drake and Ruhoff 
1960a, p. 38. 
Type Species: Axiokersos ovalis Distant. 

♦Axiokersos ovalis Distant 

Axiokersos ovalis Distant 1909a, p. 122 [India]; 1910a, p. 124, fig. 65- 
Type: Sex unknown; Calcutta, India; Indian Mus. 
Distribution: India. 
Host Plant: Unrecorded. 

Genus BAEOCHILA Drake and Poor 

Cysteochila {Baeochila) Drake and Poor 1937b, p. 400. 
Baeochila: Drake 1948e, p. 151.— Drake and Ruhoff 1960a, p. 39. 
Type Species: Cysteochila elongata Distant. 

♦Baeochila dehrana Drake and Maa 

Baeochila dehrana Drake and Maa 1954, p. 113 [India]. 

Type: Holotype o 71 , macropterous; Dehra Dun, India; Drake Coll. 
(USNM). 

Distribution: India; Ceylon. 

Host Plant: Unrecorded. 

♦Baeochila elongata (Distant) 

Cysteochila elongata Distant 1903a, p. 49 [India]; 1903b, p. 138, fig. 

102.— Drake and Poor 1937a, p. 7. 
Cysteochila {Baeochila) elongata: Drake and Poor 1937b, p. 400 [Taiwan]. 
Baeochila elongata: Drake 1948e, p. 151. — Takeya 1951a, p. 21. 



96 U.S. NATIONAL MUSEUM BULLETIN 243 

Type: Sex unknown; Margherita, Assam, India; British Mus. 
Distribution: India; Burma; Taiwan. 
Host Plant: Unrecorded. 
Baeochila nexa (Distant) 

Cysteochila nexa Distant 1903a, p. 49 [India]; 1903b, p. 139. — Drake and 

Poor 1937a, p. 7. 
Baeochila nexa: Drake and Ruhoff 1960c, p. 29. — Drake and Davis 1960, 
fig. 53. 
Type: Sex unknown; Margherita, Assam, India; British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 
*Baeochila perinetana Drake 

Baeochila per inetana Drake 1958c, p. 320 [Malagasy]. 
Type: Holotype cf; Perinet, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 
*Baeochila scitula Drake 

Baeochila scitula Drake 1948e, p. 151 [Taiwan]. — Takeya 1951a, p. 21. 
Type: Holotype d% macropterous; Formosa; Drake Coll. (USNM). 
Distribution: Taiwan. 
Host Plant: Unrecorded. 

Genus BAEOTINGIS Drake and Poor 

Baeotingis Drake and Poor 1 939b, p. 96. — Drake and Ruhoff 1 960a, p. 39. 
Type Species: Baeotingis ogloblini Drake and Poor. 
*Baeotingis ogloblini Drake and Poor 

Baeotingis ogloblini Drake and Poor 1939b, p. 96, fig. [Argentina]. — 
Kormilev 1955a, p. 64. 
Type: Holotype o", brachypterous; San Ignacio, Misiones, Argentina; 
Drake Coll. (USNM). 
Distribution : Argentina. 
Host Plant: Unrecorded. 
*Baeotingis silvestrii Drake 

Baeotingis silvestrii Drake 1948e, p. 153 [Argentina]. 
Type: Holotype o 71 ; Rio Santa Cruz, Patagonia, Argentina; Hungarian 
Mus. 

Distribution : Argentina. 
Host Plant: Unrecorded. 
*Baeotingis vianai Kormilev 

Baeotingis vianai Kormilev 1955a, p. 65, fig. 1 [Argentina]. 
Type: Holotype 9, brachypterous; La Falda, Cordoba, Argentina; 
Drake Coll. (USNM). 
Distribution: Argentina. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 97 

Genus BAICHILA Drake and Slater 

Baichila Drake and Slater 1955, p. 50.— Drake and Runoff 1960a, p. 39. 
Type Species: Baichila capeneri Drake and Slater. 

*Baichila capeneri Drake and Slater 

Baichila capeneri Drake and Slater 1955, p. 51 [Transvaal]. 

Type: Holotype o", macropterous; Rustenburg, Union of South Africa; 
Drake Coll. (USNM). 

Distribution: South Africa (Transvaal). 

Host Plant: Unrecorded. 

*Baichila pulla Drake and Slater 

Baichila pulla Drake and Slater 1955, p. 52 [Transvaal]. 

Type: Holotype o 71 , macropterous; Rustenburg, Union of South Africa; 
Drake Coll. (USNM). 

Distribution: South Africa (Transvaal). 

Host Plant: Unrecorded. 

*Baichila vulsa Drake and Slater 

Baichila vulsa Drake and Slater 1955, p. 51 [Transvaal]. 
Type: Holotype 9, macropterous; Rustenburg, Union of South Africa; 
Drake Coll. (USNM). 
Distribution: South Africa (Transvaal). 
Host Plant: Unrecorded. 

Genus BAKO Schouteden 

Bako Schouteden 1923, p. 91.— Drake 1954f, p. 8; 1956e, pp. 63-66.— 

Drake and Ruhoff 1960a, p. 39. 
Galeotingis Drake 1947b, p. 1.— Drake and Ruhoff 1960a, p. 58. 
Type Species: Bako lebruni Schouteden. 

*Bako capeneri Drake 

Bako capeneri Drake 1956e, p. 65 [Transvaal]. 
Type: Holotype 9, macropterous; Johannesburg, South Africa; Drake 
Coll. (USNM). 
Distribution: South Africa (Transvaal). 
Host Plant: Unrecorded. 

*Bako dieides Drake and Ruhoff 

Bako dieides Drake and Ruhoff 1961b, p. 181, figs. 22 a, b [Cape 
Province]. 
Type: Holotype cf, macropterous; Grahamstown, South Africa; Drake 
Coll. (USNM). 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 



98 U.S. NATIONAL MUSEUM BULLETIN 243 

*Bako editus Drake 

Bako editus Drake 1956e, p. 65 [Transvaal]. 
Type: Holotype 9, macropterous; Johannesburg, South Africa; Drake 
Coll. (USNM). 
Distribution: South Africa (Transvaal); Malagasy Republic. 
Host Plant: Unrecorded. 
*Bako lebruni Schouteden 

Bako lebruni Schouteden 1923, p. 91 [Congo].— Drake 1954f, p. 8; 

1956e, p. 64.— Drake and Ruhoff 1962a, p. 156. 
Galeotingis usumburana Drake 1947b, p. 2, fig. 1. 
Bako usumburana: Drake 1954f, p. 8; 1956e, p. 64. 
Type: Sex unknowm; Kwamouth, Belgian Congo; Cent. Air. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 
*Bako malayanus (Drake) 

Galeotingis malayana Drake 1947b, p. 2 [Malaya]. 

Bako malayanus: Drake 1954f, p. 8; 1956e, p. 64. — Drake and Mohana- 

sundarum 1961, p. 113 [India; Panicum repens; Cynadon dactylon; 

Cenchurus glaucus; Oriza sativa]. 

Type: Holotype cf, macropterous; Parit Butan, Malay Peninsula; 

Drake Coll. (USNM). 

Distribution: Federation of Malaya; Philippine Islands (Luzon); India. 

Host Plants: Cenchrus glaucus; Cynodon dactylon; Oryza sativa; Panicum 

repens. 

Genus BANAHAONA Drake and Ruhoff 

Banahaona Drake and Ruhoff 1961b, p. 170. 
Type Species: Banahaona exalla Drake and Ruhoff. 
*Banahaona exalla Drake and Ruhoff 

Banahaona exalla Drake and Ruhoff 1961b, p. 171, figs. 16 a, b [Luzon]. 
Type: Holotype o 71 , macropterous; Mount Banahao, Luzon, Philippine 
Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

Genus BELENUS Distant 

Belenus Distant 1909a, p. 116; 1910a, p. 115.— Horvath 1910, p. 62.— 
Drake 1957c, pp. 31-35.— Drake and Ruhoff 1960a, p. 39. 
Type Species: Monanthia dentatus Fieber. 
*Belenus adocetus Drake 

Belenus adocetus Drake 1957c, p. 32 [Uganda]. 
Type: Holotype d% macropterous; Jinj a, Uganda; Drake Coll. (USNM). 
Distribution: Uganda. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 99 

Belenus angulatus Distant 

Belenus angulatus Distant 1909a, p. 116 [Burma]; 1910a, p. 116, fig. 59. 
Type: Sex unknown; Burma; British Mus. 
Distribution: Burma. 
Host Plant: Unrecorded. 

*Belenus bengalensis Distant 

Belenus bengalensis Distant 1909a, p. 117 [India]; 1910a, p. 117. 
Type: Sex unknown; India; British Mus. 
Distribution: India; Thailand. 
Host Plant: Unrecorded. 

*Belenus dentatus (Fieber) 

Monanthia {Phyllontocheila) dentata Fieber 1844, p. 71, pi. 6, figs. 2-4 

[Ostindien].— Herrich-Schaeffer 1850, p. 155. 
Phyllontochila dentata: Stal 1873, p. 128 [India orientalis]. — Distant 

1903b, p. 136. 
Belenus dentatus: Distant 1909a, p. 116 [India; Borneo]; 1910a, p. 116, 
fig. 58— Horvath 1912c, p. 342 [Java].— Esaki 1926, p. 163.— Drake 
1923a, p. 104 [Taiwan]; 1937a, p. 387 [Thailand; Viet-Nam; Su- 
matra]; 1957c, p. 32.— Drake and Poor 1936b, p. 147 [Malaya]; 
1937a, p. 15 [Luzon]; 1937b, p. 402.— Blote 1945, p. 86.— Takeya 
1951a, p. 15. 
Type: "Ostindien"; sex and deposition of type unknown. 
Distribution: India; Thailand; Viet-Nam; Federation of Malaya; 
Taiwan; Greater Sunda Islands (Borneo; Java; Sumatra); Philippine Is- 
lands (Luzon; Mindanao); Marianas Islands (Guam). 
Host Plant: Unrecorded. 
Note: Taken at light. 

*Belenus laplumei (Schouteden) 

Phyllontocheila laplumei Schouteden 1916a, p. 271 [Congo]; 1916b, p. 292. 
Belenus laplumei: Drake 1957c, p. 32. 
Type: Sex unknown; Api, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Sierra Leone; Central African Republic. 
Host Plant: Unrecorded. 
*Belenus thomasi Drake 

Belenus thomasi Drake 1957c, p. 33 [Kenya]. 
Type: Holotype o 71 , macropterous; Limuru, Kenya Colony; Drake Coll. 
(USNM). 
Distribution: Kenya. 
Host Plant: Unrecorded. 

Genus BEROTINGIS Drake 

Berotingis Drake 1956d, pp. 106, 113.— Drake and Ruhoff 1960a, p. 40. 
Type Species: Berotingis rugiana Drake. 



100 U.S. NATIONAL MUSEUM BULLETIN 243 

:! Berotingis guamensis (Drake) 

Tingis (Tingis) guamensis Drake 1941a, p. 142 [Guam]. 

Tingis guamensis: Usinger 1946a, p. 39 [Premna gaudichaudii]. 

Berotingis guamensis: Drake 1956d, pp. 106, 115. 
Type: Holotype 9, macropterous; Guam; USNM. 
Distribution: Marianas Islands (Guam). 
Host Plant: Premna gaudichaudii. 

*Berotingis rugiana Drake 

Berotingis rugiana Drake 1956d, pp. 106, 113, fig. 7b [Rota]. 
Type: Holotype o 71 , macropterous; Rugi, Rota, Marianas Islands; USNM. 
Distribution: Marianas Islands (Rota; Guam). 
Host Plant: Unrecorded. 

'••'Berotingis yapensis (Drake) 

Eteoneus yapensis Drake 1946, p. 27 [Yap; Premna gaudichaudii]. — Takeya 

1951a, p. 18. 
Berotingis yapensis: Drake 1956d, pp. 106, 113, fig. 7a. 
Type: Holotype cT; Dogor-Kanif-Rul, Yap; Kyushu Univ. 
Distribution: Caroline Islands (Yap). 
Host Plant: Premna gaudichaudii. 

Genus BIRABENA Drake and Hurd 

Birabena Drake and Hurd 1945b, p. 127.— Drake and Ruhoff 1960a, 
p. 40. 
Type Species: Birabena birabeni Drake and Hurd. 

*Birabena angusta (Drake and Hambleton) 

Atheas angusta Drake and Hambleton 1940, p. 535 [Brazil]. — Monte 

1941e, p. 80. 
Birabena angusta: Drake and Hurd 1945b, p. 128. 
Type: Holotype 9, macropterous; S. Jose dos Campos, Sao Paulo, 
Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Birabena birabeni Drake and Hurd 

Birabena birabeni Drake and Hurd 1945b, p. 127, fig. 1 [Argentina]. — 
Kormilev 1955a, p. 64. 
Type: Holotype 9, macropterous; Puna, S. del Estero, Argentina; La 
Plata Mus. 
Distribution: Argentina. 
Host Plant: Unrecorded. 



LACE BUGS OF THE WORLD 101 

Birabena carvalhoi Monte 

Birabena carvalhoi Monte 1947b, p. 429 [Brazil]. 
Type: Holotype cf; Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Birabena elongata (Drake) 

Atheas elongata Drake 1922c, p. 367, pi. 39, fig. 1 [Brazil]. — Monte 
1941e, p. 80; 1942a, p. 97. 

Birabena elongata: Drake and Hurd 1945b, p. 128. 
Type: Sex unknown; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

Genus BUNIA Schouteden 

Bunia Schouteden 1955b, p. 167. — Drake and Ruhoff 1960a, p. 41. 
Type Species: Bunia ituriensis Schouteden. 
Bunia ituriensis Schouteden 

Bunia ituriensis Schouteden 1955b, p. 167 [Congo]. 
Type: Sex unknown; Bunia, Ituri, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Genus BUNOTINGIS Drake 

Bunotingis Drake 1948e, p. 152.— Drake and Ruhoff 1960a, p. 41. 
Type Species: Cysteochila (Parada) camelinus Hacker. 
*Bunotingis camelina (Hacker) 

Cysteochila (Parada) camelinus Hacker 1927, p. 24, pi. 8, fig. 10 [New 

South Wales]. 
Bunotingis camelina: Drake 1948e, p. 152. 
Type: Sex unknown; Tooloom, New South Wales, Australia; Queens- 
land Mus. 
Distribution: Australia (New South Wales; Queensland). 
Host Plant: Unrecorded. 

Genus CALLITHRINCUS Horvath 

Callithr incus Horvath 1925a, p. 10.— Drake and Ruhoff 1960a, p. 42. 
Type Species: Callithr incus serratus Horvath. 
*Callithrincus serratus Horvath 

Callithrincus serratus Horvath 1925a, p. 10, fig. 6 [Queensland]. — Hacker 
1928, p. 181.— Drake 1942b, p. 361. 
Type: Holotype 9; Mt. Tambourine, Queensland, Australia; Stock- 
holm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 



102 U.S. NATIONAL MUSEUM BULLETIN 243 

*Callithrincus signatus Drake 

Callithrincus signatus Drake 1942b, p. 361 [Queensland]. 
Type: Holotype o 71 , macropterous; Mount Glorious, Queensland, 
Australia; Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Genus CALOLOMA Drake and Bruner 

Caloloma Drake and Bruner 1924a, p. 152.— Drake 1945, p. 98.— Hurd 
1946, p. 472.— Drake and Runoff 1960a, p. 42. 
Type Species: Caloloma uhleri Drake and Bruner. 

*Caloloma uhleri Drake and Bruner 

Caloloma uhleri Drake and Bruner 1924a, p. 153 [Antigua]. — Drake 
1945, p. 98 [Australia]; 1961b, p. 112.— Drake and Runoff 1961b, 
p. 183, fig. 24. 
Type: Holotype 9, macropterous; San Juan, Antigua, Lesser Antilles; 
USNM. 

Distribution: Leeward Islands (Antigua); Windward Islands (St. 
Vincent); Australia (Queensland; New South Wales; South Australia). 
Host Plant: Unrecorded. 

Note: Australian indigene. Contrary to previous statements in the 
literature, this species now appears to be permanently established in the 
West Indies. 

Genus CALOTINGIS Drake 

Calotingis Drake 1918d, p. 86.— Hacker 1929, p. 334.— Hurd 1946, 

p. 454.— Drake and Ruhoff 1960a, p. 42. 
Neopachycysta Hacker 1928, p. 183.— Drake and Ruhoff 1960a, p. 70. 
Type Species: Calotingis knighti Drake. 

*Calotingis knighti Drake 

Calotingis knighti Drake 1918d, p. 87 [Tex.; Malvaviscus drummondii]. — 
Van Duzee 1923, p. 141 [Mexico].— Drake and Davis 1960, fig. 68. 
Type: Holotype o 71 , macropterous; Helotes, Texas, U.S.; Drake Coll. 
(USNM). 
Distribution: U.S. (Tex.); Mexico. 
Host Plant: Malvaviscus drummondii. 

Calotingis subopaca (Hacker) 

Neopachycysta subopaca Hacker 1928, p. 184, pi. 22, figs. 11,12 [Queens- 
land]. 
Calotingis subopaca: Hacker 1929, p. 334. 
Type: Sex unknown; Maleny, Queensland, Australia; Queensland Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 103 

Genus CAMPYLOSTEIRA Fieber 

Campylosteira Fieber 1844, p. 42. — Horvath 1906a, pp. 13, 15 (key). — 
Muzik 1907, pp. 49, 51 (key).— Jensen-Haarup 1912, p. 150 — 
Stichel 1926, pp. 103, 105 (key); 1960a, p. 269 (key); 1960b, p. 
390; 1960c, p. 129.— Guide 1938, pp. 242, 245 (key).— China 1943, 
p. 245.— Kiritshenko 1951, pp. 240, 243 (key).— Drake and Ruhoff 
1960a, p. 43. 

Campylostira [sic]: Fieber 1861, p. 131. — Douglas and Scott 1865, 
p. 257.— Stal 1874, p. 52.— Puton 1887, p. 100 (key).— Horvath 
1892, p. 309.— Lethierry and Severin 1896, p. 9.— Oshanin 1908, 
p. 402; 1912, p. 42.— Stichel 1935, p. 348; 1938a, p. 404. 
Type Species: Campylosteira falleni Fieber. 

Campylosteira bosnica Horvath 

Campylostira bosnica Horvath 1892a, pp. 309, 311 [Yugoslavia]. 
Campylosteira bosnica: Horvath 1906a, pp. 15, 19. — Stichel 1926, p. 
105; 1960a, p. 269. 
Type: Sex unknown; Bosnia; Hungarian Mus. 
Distribution : Yugoslavia. 
Host Plant: Unrecorded. 

Campylosteira bosnica var. diluta Horvath 

Campylostira bosnica var. diluta Horvath 1892a, pp. 309, 312 [Yugoslavia]. 
Campylosteira bosnica var. diluta: Horvath 1906a, p. 19. — Stichel 1960a, 
p. 269. 
Type: Sex unknown; Bosnia; Hungarian Mus. 
Distribution : Yugoslavia. 
Host Plant: Unrecorded. 
Campylosteira ciliata Fieber 

Campylosteira ciliata Fieber 1844, p. 44, pi. 3, figs. 33-37 [Czechoslo- 
vakia].— Herrich-Schaeffer 1850, p. 160.— Horvath 1906a, p. 23 — 
Stichel 1926, p. 106; 1960a, p. 273, fig. 38a.— Hoberlandt 1943b, 
p. 120. 
Campylostira ciliata: Fieber 1861, p. 132 [moss]. — Puton 1887, p. 100. — 

Horvath 1892a, pp. 309, 311.— Hueber 1893, p. 300. 
Monanthia ciliata: Walker 1873a, p. 190. 
Type: Prague, Czechoslovakia; sex and deposition of type unknown. 
Distribution: Czechoslovakia. 
Host Plant: Moss. 
Campylosteira dispar Horvath 

Campylosteira dispar Horvath 1905b, p. 558 [Algeria; Tunisia]; 1906a, 
p. 22.— Hoberlandt 1955, p. 86 [Turkey]. 
Type: Unknown. 

Distribution: Algeria; Tunisia; Turkey. 
Host Plant: Unrecorded. 



104 U.S. NATIONAL MUSEUM BULLETIN 243 

*Campylosteira eximia Horvath 

Campylostira eximia Horvath 1892a, pp. 311, 313 [Turkestan]. 

Campylosteira eximia: Horvath 1906a, p. 22. 
Type: Holotype 9; Taschkent, Turkestan, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 

^Campylosteira falleni Fieber 

Campylosteira fal lent Fieber 1844, p. 43, pi. 3, figs. 23-26 [Czechoslo- 
vakia].— Horvath 1906a, p. 20.— Stichel 1926, p. 105; 1960a, p. 270, 
fig. 34a.— Kiritshenko 1951, p. 243 [Russia]. 
Campylosteira jallenii [sic]: Herrich-Schaeffer 1850, p. 160. 
Campylostira falleni: Fieber 1861, p. 132 [moss]. — Puton 1879c, p. 93. — 
Lethierry and Severin 1896, p. 9 [Hungary; Yugoslavia; Rumania; 
Caucasus].— Blote 1945, p. 82. 
Monanthia falleni: Walker 1873a, p. 190. 
Campylostira fallenii [sic]: Hiieber 1893, p. 302. 
Type: Sex unknown; Prague, Czechoslovakia; Vienna Mus. 
Distribution: Czechoslovakia; Hungary; Yugoslavia; Rumania; 
U.S.S.R. (Caucasus). 
Host Plant: Moss. 

Campylosteira horvathi Drake 

Campylosteira horvathi Drake 1951, p. 171 [Tanganyika]. 
Type: Holotype o", brachypterous; Arusha-Chini, Africa orientalis; 
Hungarian Mus. 
Distribution: Tanganyika. 
Host Plant: Unrecorded. 

Campylosteira libanotica Horvath 

Campylosteira libanotica Horvath 1906a, p. 21 [Lebanon]. 
Type: Sex unknown; Lebanon; Hungarian Mus. 
Distribution: Lebanon. 
Host Plant: Unrecorded. 

Campylosteira moroccana Puton 

Campylostira moroccana Puton 1887, p. 100 [Morocco]. 

Campylostira maroccana [sic] : Horvath 1892a, p. 310. 

Campylosteira maroccana [sic]: Horvath 1906a, p. 20 [Mauritania]. 
Type: Sex unknown; Morocco; Paris Mus. 
Distribution: Morocco; Mauritania. 
Host Plant: Unrecorded. 

* Campylosteira orientalis Horvath 

Campylostira orientalis Horvath 1881a, p. 223 [Hungary; Caucasus; 
Yugoslavia]. — Oshanin 1912, p. 42 [Austria; Crimea]. — Stichel 
1938a, p. 404 [Turkey; Albania]. 



LACEBUGS OF THE WORLD 105 

Campyl&stira falleni var. orien talis: Horvath 1892a, p. 310 [Rumania; 

Transcaucasus]. 
Campylosteira orientalis: Horvath 1906a, p. 20. — Kiritshenko 1951, 
p. 243.— Stichel 1960a, p. 270. 
Type: Unknown. 

Distribution: Hungary; Austria; Yugoslavia; Albania; Turkey; Ru- 
mania; U.S.S.R. (Caucasus; Transcaucasus; Crimea). 
Host Plant: Unrecorded. 

*Campylosteira orientalis var. miridita Horvath 

Campylosteira falleni var '. miridita Horvath 1905b, p. 558 [Albania]. 

Campylosteira orientalis var. miridita: Horvath 1906a, p. 20. — Mancini 
1953b, p. 185.— Stichel 1960a, p. 270. 

Campylostira orientalis v ar. miridita: Horvath 1916, p. 9. 
Type: Sex unknown; Velipoja, Albania; Hungarian Mus. 
Distribution: Albania. 
Host Plant: Unrecorded. 

Campylosteira orientalis var. suspecta Horvath 

Campylostira falleni (not Fieber) : Horvath 1892a, p. 310. 

Campylostira falleni var . suspecta Horvath 1892a, pp. 310, 312 [Rumania]. 

Campylosteira orientalis var. suspecta: Horvath 1906a, p. 20 [Hungary]. — 
Stichel 1960a, p. 271. 
Type: Sex unknown; Rumania; Hungarian Mus. 
Distribution: Rumania; Hungary. 
Host Plant: Unrecorded. 

Campylosteira parvula Ferrari 

Campylostira parvula Ferrari 1874, p. 172 [Iran]. — Horvath 1892a, p. 310. 

Serenthia parvula: Lethierry and Severin 1896, p. 5. 

Campylosteira parvula: Horvath 1906a, p. 23. 
Type: Sex unknown; Persia; Genova Mus. 
Distribution: Iran. 
Host Plant: Unrecorded. 

*Campylosteira perithrix Puton 

Campylostira perithrix Puton 1887, p. 99 [Algeria]. — Horvath 1892a, p. 
309. 

Serenthia perithrix: Lethierry and Severin 1896, p. 5. 

Campylosteira perithrix: Horvath 1906a, p. 23. 
Type: Sex unknown; Algeria; Paris Mus. 
Distribution : Algeria ; Tunisia . 
Host Plant: Unrecorded. 



681-552—64- 



106 U.S. NATIONAL MUSEUM BULLETIN 243 

Campylosteira perithrix var. cognata Horvath 

Campylosteira cognata Horvath 1905b, p. 559 [Algeria]. 

Campylosteira perithrix var. cognata: Horvath 1906a, p. 23. 
Type: Sex unknown; St. Charles, Algeria; Hungarian Mus. 
Distribution: Algeria. 
Host Plant: Unrecorded. 

*Campylosteira pilicornis Horvath 

Campylosteira pilicornis Horvath 1906a, p. 19 [Israel]. 
Type: Sex unknown; Jerusalem, Syria; Hungarian Mus. 
Distribution: Israel. 
Host Plant: Unrecorded. 

*Campylosteira pilifera Reuter 

Campylostira pilifera Reuter 1880a, p. 12 [Greece]; 1891a, p. 25. — Puton 

1887, p. 100.— Horvath 1892a, pp. 309, 311. 
Serenthia pilifera: Lethierry and Severin 1896, p. 5. 

Campylosteira pilifera: Reuter 1902, p. 151. — Horvath 1906a, p. 23. — 
Stichell960a,p.272. 
Type: Sex unknown; Olympus, Greece; Helsin. Mus. 
Distribution: Greece. 
Host Plant: Unrecorded. 

Campylosteira pilifera var. diminuta Horvath 

Campylosteira pilifera var. diminuta Horvath 1905b, p. 561 [Greece]; 
1906a, p. 23.— Stichel 1960a, p. 272. 
Type: Sex unknown; Attica, Greece; Hungarian Mus. 
Distribution: Greece. 
Host Plant: Unrecorded. 

*Campylosteira pilifera var. humilis Horvath 

Campylosteira pilifera var. humilis Horvath 1905b, p. 560 [Greece]; 
1906a, p. 23.— Stichel 1960a, p. 272. 
Type: Holotype 9; Greece; Hungarian Mus. 
Distribution: Greece. 
Host Plant: Unrecorded. 

^Campylosteira rotundata Takeya 

Campyrostira [sic] rotundata Takeya 1933, p. 34, pi. 3, fig. 2, text fig. 2 

[Japan]. 
Campylostira rotundata: Takeya 1951a, p. 6. — Esaki 1954, p. 235, fig. 607. 
Type: Holotype 9; Kyoto, Honshu, Japan; Kyushu Univ. 
Distribution: Japan; China (Hunan). 
Host Plant: Unrecorded. 

Campylosteira serena Horvath 

Campylosteira serena Horvath 1902a, p. 593 [Spain]; 1906a, p. 22. — 
Gomez-Menor 1955b, p. 249.— Stichel 1960a, p. 272 [Sardinia]. 



LACEBUGS OF THE WORLD 107 

Type: Holotype 9; Ciudad Real, Spain; Hungarian Mus. 
Distribution: Spain; Sardinia. 
Host Plant: Unrecorded. 

Campylosteira sinuata Fieber 

Campylostira sinuata Fieber 1861, p. 132 [Germany]. — Puton 1879c, 
p. 93. — Horvath 1892a, p. 311 [Transcaucasus]. — Hiieber 1893, 
p. 303— Borner 1935, p. 75. 
Campylosteira sinuata: Horvath 1906a, p. 22, pi. 1, fig. 3. — Stichel 1926, 

p. 106, fig. 294; 1960a, p. 272.— Kiritshenko 1951, p. 243. 
Monanthia sinuata: Walker 1873a, p. 190. 
Serenthia sinuata: Lethierry and Severin 1896, p. 5. 
Type: Sex unknown; Germany; Vienna Mus. 
Distribution: Germany; U.S.S.R. (Transcaucasus). 
Host Plant: Unrecorded. 

Campylosteira sororcula Horvath 

Campylosteira sororcula Horvath 1905b, p. 560 [Algeria]; 1906a, p. 23. 
Type: Holotype o 71 ; St. Charles, Algeria; Hungarian Mus. 
Distribution: Algeria. 
Host Plant: Unrecorded. 

*Campylosteira verna (Fallen) 

Tingis verna Fallen 1826, p. 16 [Sweden]; 1829, p. 147. 

Monanthia verna: Herrich-Schaeffer 1838, pp. 53, 64, pi. 127, fig. 398 
[Germany; Czechoslovakia]. 

Campylosteira brachycera Fieber 1844, p. 43, pi. 3, figs. 27-32 [moss]. — 
Herrich-Schaeffer 1850, p. 160.— Mayr 1858, p. 569 [Austria; 
Hungary]. 

Campylosteira verna: Fieber 1844, p. 45, pi. 3, fig. 38. — Kolenati 1856, 
p. 429 [Transcaucasus; Onosma echioides]. — Reuter 1874, p. 565. — 
Horvath 1898b, p. 279; 1906a, pp. 17, 21, pi. 1, fig. 2 [Denmark; 
Belgium; Italy; Rumania]. — Jensen-Haarup 1912, p. 150, fig. 94. — 
Reichensperger 1920, p. 61.— Butler 1923, p. 200.— Stichel 1926, 
p. 106, fig. 293; 1960a, p. 271, fig. 36 [Poland; Albania].— Guide 
1938, p. 247, fig.— Hoberlandt 1942, p. 124; 1943b, p. 122.— Kirit- 
shenko 1951, p. 243.— Bator 1953, p. 323, pi. 4, fig. 2.— Mancini 
1953c, p. 22. — Stusakl958, p. 367, figs. 8-11. — Southwood and Leston 
1959, p. 141, pi. 18, fig. 2, pi. 21, fig. 1. 

Campylostira brachycera: Fieber 1861, p. 132. — Douglas and Scott 1865, 
p. 259.— Frey-Gessner 1865, p. 233 [Switzerland].— Horvath 1874b, 
p. 432.— Saunders 1875, p. 246. 

Campylostira verna: Fieber 1861, p. 132. — Douglas and Scott 1865, 
p. 258. — Reiber and Puton 1876, p. 67 [France]. — Vollenhoven 
1878, p. 282, pi. 10, fig. 3 [Netherlands].— Puton 1879c, p. 92 — 
Jakovlev 1880b, p. 100 [Caucasus].— Reuter 1882b, p. 113.— Saun- 



108 U.S. NATIONAL MUSEUM BULLETIN 243 

ders 1892, p. 124, pi. 11, fig. 6.— Horvath 1892a, p. 310.— Hiieber 
1893, p. 300.— Oshanin 1908, p. 403 [Spain].— Schumacher 1914, 
p. 256.— Scholte 1935, p. 29, fig. 4.— Borner 1935, p. 75— Blote 
1945, p. 82 [Greece]. — Novak and Wagner 1951, p. 70 [Yugoslavia]. 
Serenthia verna: Lethierry and Severin 1896, p. 6. 
Type: Sex unknown; Sweden; Lund Zool. Inst. 

Distribution: England; Belgium; Netherlands; France; Spain; Italy; 
Switzerland; Germany; Denmark; Sweden; Poland; Austria; Czechoslo- 
vakia; Hungary; Yugoslavia; Albania; Rumania; Greece; U.S.S.R. 
(Caucasus; Transcaucasus). 
Host Plants: Onosma echioides; moss. 
Note: Eggs [Stusak]. 
Campylosteira verna var. latipennis Horvath 

Campylostira verna var. latipennis Horvath 1892a, pp. 310, 312 [Rumania; 

Hungary]. 
Campylosteira verna var. latipennis: Horvath 1906a, p. 22 [Yugoslavia; 

Germany; France].- — Muzik 1907, p. 52 [Czechoslovakia]. 
Serenthia verna var. latipennis: Lethierry and Severin 1896, p. 6. 
Type: Unknown. 

Distribution: France; Germany; Czechoslovakia; Hungary; Yugo- 
slavia; Rumania. 
Host Plant: Unrecorded. 

Genus CAMPYLOTINGIS Drake and Bondar 

Campylotingis Drake and Bondar 1932, p. 89. — Monte 1939b, p. 66; 
1941e, p. 81.— Drake and Ruhoff 1960a, p. 43. 
Type Species: Tigava mollicula Drake. 
*Campylotingis bondari (Drake) 

Tigava bondari Drake 1930b, p. 25 [Brazil]; 1931a, p. 405. 
Campylotingis bondari: Drake and Bondar 1932, p. 89. — Costa Lima 
1936, p. 125.— Monte 1941e, p. 82 [Machaerium].—Si\va. 1956, p. 
19, fig. 4 [Canavalia ensiformis; Dolichos lablab; Phaseolus lunatus; 
Phaseolus vulgaris; Vigna sinensis']. 
Type: Holotype d% macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 

Host Plants: Canavalia ensiformis; Dolichos lablab; Machaerium sp.; 
Phaseolus lunatus; Phaseolus vulgaris; Vigna sinensis. 
*Campylotingis carvalhoi Drake and Hambleton 

Campylotingis carvalhoi Drake and Hambleton 1938a, p. 47 [Brazil; 
Papilionaceae]. — Monte 1941e, p. 82 [Ferreirea spectabilis]. 
Type: Holotype o 71 , macropterous; Curvello, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plants: Ferreirea spectabilis; Papilionaceae. 



LACEBUGS OF THE WORLD 109 

^Campylotingis clara Drake and Hambleton 

Campylotingis clara Drake and Hambleton 1942b, p. 77 [Brazil]. 
Type: Holotype d\ macropterous; Belo Horizonte, Minas Gerais, 
Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Campylotingis clavata Drake and Hambleton 

Campylotingis clavata Drake and Hambleton 1939, p. 156 [Brazil]. — 
Monte 1941e, p. 82 [Myrocarpus fastigiatus]. 
Type: Holotype o 71 , macropterous; Belo Horizonte, Minas Gerais, 
Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Myrocarpus fastigiatus. 
*Campylotingis genetica Drake and Hambleton 

Campylotingis genetica Drake and Hambleton 1942b, p. 78 [Brazil]. 
Type: Holotype o 71 , macropterous; Vigosa, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
^Campylotingis integra Drake and Hambleton 

Campylotingis integra Drake and Hambleton 1942b, p. 79 [Brazil]. 
Type: Holotype d\ macropterous; Belo Horizonte, Minas Gerais, 
Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
* Campylotingis jansoni (Drake) 

Tigava jansoni Drake 1922c, p. 364, pi. 39, fig. 8 [Brazil]. 
Campylotingis jansoni: Drake and Bondar 1932, p. 89. — Drake and 
Hambleton 1938a, p. 46 [Mackaerium]. 
Type: Holotype cf; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Mackaerium sp. 
^Campylotingis lenatis Drake 

Camplotingis [sic] lenatis Drake 1935, p. 14 [Paraguay]. 
Campylotingis lenatis: Drake and Hambleton 1938a, p. 46 [Brazil; 
Mackaerium]. — Monte 1940e, p. 290, fig. 3. 
Type: Holotype o"; S. Bernadino, Paraguay; Vienna Mus. 
Distribution: Paraguay; Brazil. 
Host Plant: Mackaerium sp. 
*Campylotingis levis Drake and Hambleton 

Campylotingis levis Drake and Hambleton 1942b, p. 80 [Brazil]. 
Type: Holotype o*, macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 



110 U.S. NATIONAL MUSEUM BULLETIN 243 

*Campylotingis machaerii Drake and Hambleton 

Camplotingis [sic] machaerii Drake and Hambleton 1934, p. 442 [Brazil; 

Argentina; Machaeriurri]. — Drake 1936b, p. 700. 
Campylotingis machaerii: Costa Lima 1936, p. 125. — Bosq 1937, p. 

128.— Monte 1939b, p. 66 [Ferreirea spectabilis]; 1940b, p. 375. 
Campylotingis snipesi Drake and Hambleton 1944b, p. 125. 
Type: Holotype d\ macropterous; Vigosa, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil; Argentina. 
Host Plants: Ferreirea spectabilis; Machaerium sp. 

*Campylotingis mollicula (Drake) 

Tigava mollicula Drake 1922c, p. 365 [Brazil]. 

Campylotingis mollicula: Drake and Bondar 1932, p. 89. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Campylotingis mollis Drake and Bondar 

Campylotingis mollis Drake and Bondar 1932, p. 90 [Brazil; Cassia], — 
Costa Lima 1936, p. 125.— Silva 1956, p. 21. 
Type: Holotype o" , macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Cassia sp. 

*Campylotingis prudens Drake and Hambleton 

Campylotingis prudens Drake and Hambleton 1938a, p. 47 [Brazil; 
Machaerium]. — Monte 1939b, p. 66 [Machaerium villosum]; 1940e, 
p. 290, fig. 4. 
Type: Holotype d", macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil; Argentina. 
Host Plants: Machaerium villosum; Machaerium sp. 

*Campylotingis tantilla Drake 

Camplotingis [sic] tantilla Drake 1935, p. 14, fig. 2 [Paraguay]. 

Campylotingis tantilla: Drake and Hambleton 1938a, p. 46 [Brazil; 
Machaerium] . 
Type: Holotype cf ; S. Bernardino, Paraguay; Vienna Mus. 
Distribution: Paraguay; Brazil. 
Host Plant: Machaerium sp. 

Genus CANTINONA Distant 

Canlinona Distant 1913, p. 158. — Drake and Runoff 1960a, p. 43. 
Type Species: Cantinona praecellens Distant. 



LACEBUGS OF THE WORLD 111 

Cantinona praecellens Distant 

Cantinona praecellens Distant 1913, p. 159, pi. 11, fig. 15 [Seychelles]. — 
Drake and Ruhoff 1961b, p. 168, fig. 14. 
Type: Sex unknown; Seychelles; British Mus. 
Distribution: Seychelles Island. 
Host Plant: Unrecorded. 



Genus CATOPLATUS Spinola 

Catoplatus Spinola 1837, p. 137— Stal 1873, pp. 121, 128; 1874, p. 57 — 
Lethierry and Severin 1896, p. 20— Horvath 1906a, pp. 14, 85 
(key).— Oshanin 1908, p. 442; 1912, p. 45.— Kiritshenko 1918, p. 
109; 1951, pp. 242, 251 (key).— Stichel 1926, pp. 104, 113 (key); 
1935, p. 349; 1938a, p. 408; 1960a, p. 323 (key); 1960b, p. 398; 
1960c, p. 137.— Borner 1935, pp. 74,77 (key).— Guide 1938, pp. 244, 
300 (key).— Hoberlandt 1942, p. 125; 1943, p. 119; 1955, p. 92.— 
China 1943, p. 247— Blote 1945, p. 88.— Singer 1952, p. 53.— Drake 
and Ruhoff 1960a, p. 43. 

Catoplatus (Coscinopoea) Stal 1873, p. 128; 1874, p. 58.— Drake and Ru- 
hoff 1960a, p. 48. 

Monanthia (Catoplatus): Puton 1879c, p. 117. — Saunders 1892, p. 
134. 
Type Species: Tingis Jabricii Stal. 

*Catoplatus amicus (Reuter) 

Monanthia (Catoplatus) antica Reuter 1880, p. 11 [Greece]. 
Catoplatus anticus: Lethierry and Severin 1896, p. 20. — Horvath 1906a, 
p. 88 [Turkey]; 1916, p. 9 [Albania]; 1918, p. 334.— Kiritshenko 
1951, p. 251 [Russia].— Mancini 1953b, p. 186.— Hoberlandt 1955, 
p. 92 [Israel].— Stichel 1960a, p. 324. 
Type: Sex unknown; Greece; Helsin. Mus. 
Distribution: Albania; Greece; Turkey; Israel; U.S.S.R. 
Host Plant: Unrecorded. 

Catoplatus anticus var. syriacus Horvath 

Catoplatus anticus var. syriacus Horvath 1906a, p. 88 [Israel]. — Hober- 
landt 1955, p. 92 [Turkey; Russia]. 
Type: Sex unknown; Syria; Hungarian Mus. 
Distribution: Israel; Turkey; U.S.S.R. (Crimea). 
Host Plant: Unrecorded. 

Catoplatus bletoni Vidal 

Catoplatus bletoni Vidal 1937, p. 197, fig. 2 [Morocco]. 
Type: Sex unknown; Morocco; Vidal Coll. 
Distribution: Morocco. 
Host Plant: Unrecorded. 



112 U.S. NATIONAL MUSEUM BULLETIN 243 

*Catoplatus carthusianus (Goeze) 

Cimex carthusianus Goeze 1778, p. 268. — Fourcroy 1785, p. 212 [Eryn- 

gium\ 
Tingis eryngii Latrielle 1804, p. 253.— Vallot 1829, pp. 99, 113 

[Eryngium campestre]. 
Tingis melanocephala Panzer 1806, heft 100, tab. 21 [Germany]. — 

Herrich-Schaeffer 1835, p. 58. 
Monanthia melanocephala: Le Peletier and Serville 1828, p. 653. — 
Herrich-Schaeffer 1838, p. 54 [Hungary; Czechoslovakia]. — Mayr 
1858, p. 570 [Caucasus; Austria]. 
Piesma melanocephalum: Burmeister 1835, p. 258 [Portugal]. 
Dictyonota eryngii: Westwood 1840b, p. 121 [England]. 
Monanthia (Tropidocheila) melanocephala: Herrich-Schaeffer 1850, p. 151. 
Monanthia albida (not Herrich-Schaeffer): Fieber 1861, p. 124 [Seseli 

glaucum] . 
Monanthia nigripes Motschulsky 1863, p. 91 [Algeria]. — Kiritshenko 

1915, p. 300. 
Monanthia eryngii: Garbiglietti 1869, p. 273 [Italy]. — Jakovlev 1880b, 
p. 109 [Onopordon].—Chicote 1880, p. 189 [Spain].— Hueber 1893, 
p. 343. 
Catoplatus (Coscinopoea) eryngii: Stal 1873, p. 128; 1874a, p. 58. 
Catoplatus (Coscinopoea) albidus: Stal 1874, p. 58. 
Monanthia (Catoplatus) eryngii: Puton 1879c, p. 117 [France; Eryngium 

maritimum]. 
Catoplatus carthusianus: Reuter 1891a, p. 26 [Greece]. — Lambertie 
1906, p. 25— Horvath 1906a, pp. 86, 89 [Turkey].— Oshanin 1908, 
p. 443 [Morocco; Rumania; Iran]. — Moroder Sala 1920, p. 12. — 
Reichensperger 1920, p. 62.— de Seabra 1924, p. 16, fig. 28; 1931, 
p. 429, figs. 500 (2-4), 501.— Stichel 1926, p. 113; 1960a, p. 324, 
fig. 127 [Sardinia; Bulgaria]. — Lindberg 1932a, p. 43. — Scholte 
1935, p. 77, fig. 20 [Netherlands].— Hoberlandt 1942, p. 125; 1955, 
p. 92 [Transcaucasus]. — Kiritshenko 1951, p. 251. — Novak and 
Wagner 1951, p. 71 [Yugoslavia; Eryngium amethystinum]. — Mancini 
1953b, p. 186 [Albania]; 1953c, p. 22.— Seidenstiicker 1954, p. 236 — 
Gomez-Menor 1955b, p. 249.— Stusak 1957a, p. 21, figs. 4, 5; 1957b, 
p. 136, fig. 4; 1959a, p. 52, figs. 1-8. 
Catoplatus melanocephalus: Lethierry and Severin 1896, p. 20. 
Cantacader nigripes: Oshanin 1908, p. 401. — Drake 1950a, p. 164. 
Type: Unknown. 

Distribution: England; France; Netherlands; Spain; Portugal; Italy; 
Sardinia; Austria; Czechoslovakia; Germany; Albania; Hungary; Yugo- 
slavia; Bulgaria; Greece; Rumania; Turkey; Iran; U.S.S.R. (Caucasus; 
Transcaucasus); Algeria; Morocco. 



LACEBUGS OF THE WORLD 113 

Host Plants: Eryngium amethystinum; Eryngium campestre; Eryngium 
maritimwn; Eryngium sp.; Onopordon sp.; Seseli glaucum. 

Note: Life history study [Stusak 1959a]; eggs [Stusak 1957a]; nymphs 
and immature stages [Stusak 1957b; 1959a]. 
*Catoplatus carthusianus var. albidus (Herrich-Schaeffer) 

Monanthia albida Herrich-Schaeffer 1838, p. 54, pi. 125, fig. b; pi. 126, 

fig. 396 [Hungary].— Ferrari 1874, p. 171 [Italy]; 1878, p. 66.— 

Reiber and Puton 1876, p. 69.— Bolivar and Chicote 1879, p. 166 

[Spain].— Jakovlev 1880b, p. 109 [Caucasus]. 

Monanthia {Tropidocheila) schaefferi Fieber 1844, p. 78, pi. 6, figs. 31-35 

[Czechoslovakia; Austria; Seseli glaucum]. 
Monanthia {Tropidocheila) albida: Herrich-Schaeffer 1850, p. 151. 
Monanthia schaefferi: Mayr 1858, p. 570 [Lichtenstein; Greece; Yugo- 
slavia; Germany; Eryngium]. 
Catoplatus (Coscinopoea) albida: Stal 1873, p. 128. 
Catoplatus dacicus Montandon 1895, p. 160 [Rumania]. 
Catoplatus carthusianus var. albidus: Horvath 1906a, p. 89. — Stichel 
1926, p. 114; 1960a, p. 324 [Sardinia].— Novak and Wagner 1951, 
p. 71.— Kiritshenko 1951, p. 251.— Hoberlandt 1955, p. 92 [Trans- 
caucasus]. — Stusak 1959a, p. 52. 
Type: Unknown. 

Distribution: Italy; Sardinia; Austria; Germany; Czechoslovakia; 
Hungary; Yugoslavia; Rumania; Lichtenstein; Spain; Greece; U.S.S.R. 
(Caucasus; Transcaucasus). 

Host Plants: Seseli glaucum; Eryngium sp. 
Catoplatus carthusianus var. intermedius Vidal 

Catoplatus carthusianus var. intermedius Vidal 1937, p. 198 [Morocco]. — 
Stichel 1960a, p. 324. 
Type: Sex unknown; Oujda, Morocco; Vidal Coll. 
Distribution: Morocco. 
Host Plant: Unrecorded. 
*Catoplatus citrinus Horvath 

Catoplatus citrinus Horvath 1897a, p. 89 [Turkestan]; 1906a, p. 91. — 
Kiritshenko 1959, p. 105. 
Type: Sex unknown; Samarkand, Turkestan; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 
*Catoplatus crassipes (Fieber) 

Monanthia crassipes Fieber 1861, p. 123 [Yugoslavia]. 
Catoplatus crassipes: Lethierry and Severin 1896, p. 20. — Horvath 1906a, 
p. 88, fig. 4.— Stichel 1960a, p. 323. 
Type: Sex unknown; Serbia; Vienna Mus. 
Distribution: Yugoslavia. 
Host Plant: Unrecorded. 



114 U.S. NATIONAL MUSEUM BULLETIN 243 

*Catoplatus dilatatus (Jakovlev) 

Monanthia {Catoplatus) dilatata Jakovlev 1880a, p. 140 [Caucasus]. 
Monanthia dilatata: Jakovlev 1880b, p. 109. 
Monanthia {Catoplatus) krueperi Reuter 1880a, p. 11 [Turkey]. 
Catoplatus krueperi: Lethierry and Severin 1896, p. 20. 
Catoplatus dilatatus: Lethierry and Severin 1896, p. 20. — Horvath 
1906a, p. 87 [Armenia; Syria].— Lindberg 1948a, p. 59 [Olea; Cyprus; 
Tunisia; Greece]. — Kiritshenko 1951, p. 251. — Seidenstiicker 1954, 
p. 235.— Hoberlandt 1955, p. 92 [Transcaucasus].— Stichel 1960, 
p. 323. 
Type: Sex unknown; Derbent, Caucasus, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Armenia; Caucasus; Transcaucasus); Turkey; 
Syria; Cyprus; Greece; Tunisia. 

Host Plant: Olea sp. 
Catoplatus dispar Drake and Maa (emendation) 

Catoplatus disparis Drake and Maa 1954, p. 114 [China]. 
Type: Holotype o"; Suisapa, Western Hupeh, China; Cal. Acad. 
Distribution: China. 
Host Plant: Unrecorded. 

*Catoplatus distinctus Montandon 

Catoplatus distinctus Montandon 1895, p. 160 [Rumania]. — Horvath 
1906a, p. 91.— Stichel 1938a, p. 408 [Russia]; 1960a, p. 326.— Kirit- 
shenko 1951, p. 251.— Seidenstiicker 1954, p. 236 [Turkey].— Hober- 
landt 1955, p. 93. 
Type: Comana, District de Vlasca, Rumania; sex and deposition of 
type unknown. 
Distribution: Rumania; Turkey; U.S.S.R. 
Host Plant: Unrecorded. 

*Catoplatus fabricii (Stal) 

Tingis costata (not Fabricius): Fallen 1807, p. 36; 1829, p. 143— Zetter- 
stedt 1828, p. 480 [Lapland]; 1840, p. 269.— Herrich-Schaeffer 1835, 
p. 58. — Germar 1836, fasc. 18, tab. 25 [Germany; Sweden]. 

Catoplatus costatus: Spinola 1837, p. 167. — Stal 1874, p. 58. 

Monanthia costata: Herrich-Schaeffer 1838, p. 55, pi. 123, fig. 390. — 
Fieber 1861, p. 123.— Douglas and Scott 1865, p. 248.— Thomson 
1871, p. 398.— Saunders 1875, p. 248.— Vollenhoven 1878, pp. 270, 
272, pi. 22, fig. 10 [Netherlands].— Chicote 1880, p. 189 [Spain].— 
Hiieber 1893, p. 341 [Switzerland]. 

Monanthia (Tropidocheila) costata: Fieber 1844, p. 72, pi. 6, figs. 10-12 
[Italy; Hungary; England].— Sahlberg 1848, p. 132 [Finland].— 
Herrich-Schaeffer 1850, p. 152.— Flor 1860, p. 347 [France]. 

Tingis fabricii Stal 1868, p. 93. 

Monanthia oblonga Garbiglietti 1869, p. 274. 



LACEBUGS OF THE WORLD 115 

Catoplatus fabricii: Stal 1873, p. 128.— Reuter 1882b, p. 119— Horvath 
1906a, p. 87 [Chrysanthemum leucanthemum]. — Muzik 1907, p. 60 
[Czechoslovakia]. — Oshanin 1908, p. 442 [Belgium; Russia; Ru- 
mania; Austria].- — Schumacher 1914, p. 258. — Lindberg 1919, p. 
42.— Reichensperger 1920, p. 62.— Sahlberg 1920, p. 84.— Stichel 
1926, p. 113; 1938a, p. 408 [Norway; Albania]; 1960a, p. 323 [Poland; 
Bulgaria; Yugoslavia; Genista tinctoria]. — Scholte 1935, p. 76, fig. 19 
[Medicago lupulina].— Guide 1938, p. 302, fig— Blote 1945, p. 88 
[Tunisia; Algeria; Egypt].— Mancini 1949b, p. 37; 1949c, p. 137; 
1954b, p. 1 1 .— Southwood and Leston 1959, p. 150, pi. 19, fig. 4 — 
Stus'ak 1961a, p. 84, fig. 9; pi. 1, fig. d. 
Monanthia fabricii: Horvath 1874b, p. 432.— Butler 1923, p. 212 

[Hypericum; Spartium]. 
Monanthia (Catoplatus) costata: Puton 1879c, p. 117 [Chrysanthemum]. — 

Saunders 1892, p. 134, pi. 12, fig. 8. 
Catoplatus jabrici [sic]: Gomez-Menor 1955b, p. 249. 
Type: Unknown. 

Distribution: England; Belgium; Netherlands; France; Spain; Ger- 
many; Switzerland; Italy; Hungary; Austria; Czechoslovakia; Rumania; 
Albania; Yugoslavia; Bulgaria; Poland; Sweden; Norway; Finland; 
(Lapland); U.S.S.R.; Algeria; Egypt; Tunisia. 

Host Plants: Chrysanthemum leucanthemum; Chrysanthemum sp.; Genista 
tinctoria; Genista sp.; Hypericum sp.; Medicago lupulina; Spartium sp. 
Note: Eggs [Stusak]. 

*Catoplatus fulvicornis (Jakovlev) 

Monanthia (Catoplatus) fulvicornis Jakovlev 1890, p. 334 [Russia]. 
Catoplatus fulvicornis: Horvath 1906a, p. 91 [Caucasus; Turkmen; 
Turkistan; Afghanistan; Turkey]. — Hoberlandt 1955a, p. 92 [Trans- 
caucasus]. — Stichel 1960a, p. 325. 
Type: Sex unknown; Russia; Leningrad Inst. 

Distribution: U.S.S.R. (Turkmen; Caucasus; Transcaucasus; Turki- 
stan); Afghanistan; Turkey. 
Host Plant: Unrecorded. 

*Catoplatus hilaris Horvath 

Catoplatus hilaris Horvath 1906a, p. 90 [Israel]. — Seidenstiicker 1954, 
p. 236 [Turkey].— Hoberlandt 1955, p. 92 [Syria].— Stichel 1960a, 
p. 326. 
Type: Sex unknown; Syria; Hungarian Mus. 
Distribution: Israel; Syria; Lebanon; Turkey. 
Host Plant: Unrecorded. 



116 U.S. NATIONAL MUSEUM BULLETIN 243 

*Catoplatus horvathi (Puton) 

Monanthia flavipes Horvath 1874a, p. 334 [Hungary]; 1874b, p. 432. 
Monanthia {Catoplatus) horvathi Puton 1879c, p. 119 [France]. 
Monanthia horvathi: Horvath 1889, p. 329 [Austria; Eryngium campestre]. — 
d'Antcssanty 1890, p. 33 [Eryngium] . — Hiieber 1893, p. 343 
[Bulgaria]. 
Catoplatus flavipes: Lethierry and Severin 1896, p. 20 [Yugoslavia]. — ■ 
Horvath 1906a, p. 90 [Rumania].— Oshanin 1908, p. 444 [Crimea; 
Tauria].— Stichel 1926, p. 114; 1938a, p. 408 [Belgium; Albania; 
Turkey]; 1960a, p. 325.— Scholte 1935, p. 77 [Netherlands].— 
Hoberlandt 1943b, pp. 121, 124 [Czechoslovakia]. — Kiritshenko 
1951, p. 251. 
Catoplatus horvathi: Drake and Runoff 1961b, p. 167; 1961c, p. 149. 
Type: Sex unknown; France; Paris Mus. 

Distribution: France; Belgium; Netherlands; Austria; Czechoslovakia; 
Hungary; Bulgaria; Rumania; Yugoslavia; Albania; U.S.S.R. (Crimea; 
Tauria) ; Turkey. 
Host Plants: Eryngium campestre; Eryngium sp. 

Catoplatus horvathi var. pallens Dobsik 

Catoplatus flavipes var. pallens Dobsik 1951, p. 40 [Czechoslovakia]. 
Type: Holotype 9; Lazanecky, Czechoslovakia; deposition unknown. 
Distribution : Czechoslovakia. 
Host Plant: Unrecorded. 
*Catoplatus leucus Kiritshenko 

Catoplatus leucus Kiritshenko 1914, p. 195 [Turkestan]. 
Type: Sex unknown; Iskender, Turkestan, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 
Catoplatus mamorensis Bergevin 

Catoplatus mamorensis Bergevin 1922, p. 108, figs. 1, 2 [Morocco; Quercus 
suber]. 
Type: Sex unknown; Rabat, Morocco; Bergevin Coll. (Paris Mus.). 
Distribution: Morocco. 
Host Plant : Quercus suber. 
*Catoplatus mamorensis var. biseriatus Lindberg 

Catoplatus mamorensis var. biseriatus Lindberg 1936a, p. 85, fig. 3 [Mo- 
rocco] . 
Type: Holotype 9; Morocco; Therys Coll. 
Distribution: Morocco. 
Host Plant: Unrecorded. 
*Catoplatus nigriceps Horvath 

Monanthia (Tropidocheila) melanocephala (not Panzer): Fieber 1844, p. 
77, pi. 6, figs. 26-30 [Germany; Austria]. 



LACEBUGS OF THE WORLD 117 

Tingis melanocephala: Kolenati 1856, p. 425 [Transcaucasus]. 

Monanthia eryngii (not Latrielle): Fieber 1861, p. 124. 

Monanthia (Catoplatas) melanocephala: Puton 1879c, p. 118 [Eryngium 

campestre] . 
Monanthia melanocephala: Hueber 1893, p. 344. 

Catoplatus nigriceps Horvath 1905b, p. 569; 1906a, p. 89 [Hungary; 
Caucasus; Potentilla].—Stiche\ 1926, p. 114; 1960a, p. 325— Hober- 
landt 1943, p. 119 [Czechoslovakia].— Blote 1945, p. 88 [Tunisia].— 
Drake 1948a, p. 7 [Japan].— Takeya 1951a, p. 18.— Kiritshenko 
1951, p. 251. 
Type: Unknown. 

Distribution: Czechoslovakia; Austria; Hungary; Germany; U.S.S.R. 
(Caucasus; Transcaucasus); Japan; Tunisia. 
Host Plants: Eryngium campestre; Potentilla sp. 

Note: The record of C. nigriceps in Japan is based upon a 9 specimen 
labeled" Sappora-Matsum." kindly sent by the late Prof. Matsumura. 

*Catoplatus olivieri (Puton) 

Monanthia (Tropidochila) olivieri Puton 1873, p. 18 [Algeria]. 
Monanthia (Catoplatus) olivieri: Puton 1879c, p. 117 [Sardenia]. 
Catoplatus (Coscinopoea) olivieri: Stal 1874, p. 58. 

Catoplatus olivieri: Lethierry and Severin 1896, p. 21. — Horvath 1906a, 
p. 90 [Tunisia].— Lindberg 1932a, p. 43 [Morocco].— Stichel 1960a, 
p. 325. 
Type: Sex unknown; Bone, Algeria; Paris Mus. 
Distribution: Algeria; Tunisia; Morocco; Sardenia. 
Host Plant: Unrecorded. 

Genus CELANTIA Distant 

Celantia Distant 1903b, p. 137.— Drake and Ruhoff 1960a, p. 43. 
Type Species: Leptodictya vagans Distant. 

Celantia creta Drake 

Celantia creta Drake 1960, p. 366, fig. 17 [New Britain]. 
Type: Holotype 0"; Kerawat, New Britain; Bishop Mus. 
Distribution: New Britain Island, Bismarck Archipelago. 
Host Plant: Unrecorded. 

*Celantia nitidula (Stal) 

Tingis (Tingis) nitidula Stal 1873, p. 130 [Australia]. 

Phyllontochila nitidula: Lethierry and Severin 1896, p. 17. 

Celantia nitidula: Drake and Ruhoff 1961b, p. 167. 
Type: Sex unknown; Australia; Stockholm Mus. 
Distribution: Australia. 
Host Plant: Unrecorded. 



118 U.S. NATIONAL MUSEUM BULLETIN 243 

fCelantia seposita Cockerell 

Celantia (?) seposita Cockerell 1921, p. 542, fig. 1 [fossil; Oligocene; 
Isle of Wight].— Drake and Ruhoff 1960a, p. 11. 
Type: Sex undeterminable; Isle of Wight; British Mus. 
Distribution: Fossil. 
*Celantia teres Drake (emendation) 

Celantia teretis Drake 1951, p. 174 [India]. 
Type: Holotype 9; Trichinopoly, India; Hungarian Mus. 
Distribution: India. 
Host Plant: Unrecorded. 

Celantia vagans (Distant) 

Leptodictya vagans Distant 1903a, p. 48 [Ceylon]. 

Celantia vagans: Distant 1903b, p. 137, fig. 101. — Drake and Ruhoff 
1961b, p. 168, fig. 13. 
Type: Sex unknown; Ceylon; British Mus. 
Distribution: Ceylon. 
Host Plant: Unrecorded. 

Genus CERATINODERMA Stal 

Ceratinoderma Stal 1873, p. 117. — Lethierry and Severin 1896, p. 9. — 
Drake and Ruhoff 1960a, p. 23. 
Type Species: Ceratinoderma fornicata Stal. 

Ceratinoderma fornicata Stal 

Ceratinoderma fornicata Stal 1873, p. 117 [Caffraria]. — Distant 1902a, 
p. 240, pi. 15, fig. 4.— Drake 1954f, p. 1. 
Type: Holotype 9; Caffraria, South Africa; Stockholm Mus. 
Distribution: South Africa. 
Host Plant: Unrecorded. 

Ceratinoderma kivuensis Schouteden 

Ceratinoderma kivuensis Schouteden 1955a, p. 27 [Congo]. 
Type: Sex unknown; Burunga, Kivu, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Genus COCHLOCHILA Stal 

Monanthia {Cochlochila) Stal 1873, p. 133. 

Cochlochila: Horvath 1910, p. 67.— Drake 1948i, pp. 179-181.— Drake 

and Ruhoff 1960a, p. 44. 
Physodictyon Lindberg 1927, p. 16.— Drake and Ruhoff 1960a, p. 77. 
Type Species: Monanthia {Cochlochila) bullita Stal. 



LACEBUGS OF THE WORLD 119 

Cochlochila adenana Drake 

Cochlochila adenana Drake 1957a, p. 419, fig. 3 [Aden]. 
Type: Holotype cT; Jebel Jihaf, Western Aden Protectorate, Arabia; 
British Mus. 

Distribution: Aden Protectorate. 
Host Plant: Unrecorded. 

*Cochlochila aemula Drake 

Cochlochila aemula Drake 1958b, p. 27 [Congo]. 
Type: Holotype o 71 ; Lusinga, Belgian Congo; Pares Nat. Inst. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Cochlochila bequaerti Schouteden 

Cochlochila bequaerti Schouteden 1916b, p. 294 [Congo]. 

Type: Sex unknown; Mufungwa-Sampwe, Belgian Congo; Cent. Afr. 
Mus. 

Distribution: Congo; South Africa (Cape Province). 

Host Plant: Unrecorded. 

*Cochlochila bullita (Stal) 

Monanthia {Cochlochila) bullita Stal 1873, p. 133. 

Tingis globulifera Walker 1873a, p. 182 [India; heliotrope]. — Kirby 

1891, p. 110, pi. 4, fig. 11 [Ceylon]. 
Monanthia globulifera: Distant 1902b, p. 356; 1903b, p. 144, fig. 107; 
1910a, p. 123 [Coleus; sage].— Maxwell-Lefroy 1909, p. 692, fig. 460 
[Ocimum basilicum]. — Fletcher 1918, p. 267 [Ocimum sanctum]; 1920, 
p. 264 [mint; safflower] .— Iyengar 1924, p. 296, pis. 27, 28.— Singh 
1953, p. 118. — Sharga 1953, p. 885, figs. 1-5 [Ocimum canum; Ocimum 
kilimandscharicum; Salvia officinalis; Carthamus tinctorius; Mentha]; 1955, 
p. 284, figs. 1-7. — Thontadarya and Channa Basavanna 1959, p. 289. 
Monanthia mitrata Distant 1904, p. 433, pi. 8, figs. 14, 14a [Cape Prov- 
ince].— Hesse 1925, p. 90.— Blote 1945, p. 91.— Drake 1948i, pp. 
179, 181. 
Cochlochila bullita: Horvath 1909b, p. 632; 1910, p. 67.— Drake 1948i, 
p. 179; 1956a, p. 4 [China]; 1956f, p. 422 [Uganda; Kenya; Java].— 
Drake and Maa 1953, p. 91. 
Cochlochila mitrata: Horvath 1910, p. 67 [Tanganyika]. — Schouteden 

1923, p. 109 [Congo]. 
Monanthia bullita: Drake 1937a, p. 386 [Philippines]. 
Type: Unknown. 

Distribution: South Africa (Cape Province); Kenya; Uganda; Tan- 
ganyika; Congo; India; Ceylon; Greater Sunda Islands (Java); Philippine 
Islands; China. 



120 U.S. NATIONAL MUSEUM BULLETIN 243 

Host Plants: Carthamus tinctorius; Coleus sp. ; Mentha sp; Ocimum basilicum; 
Ocimum canum; Ocimum kilimandscharicum; Ocimum sanctum; Salvia officinalis; 
heliotrope. 

Note: Life history studies [Iyengar; Sharga 1953]; genitalia and repro- 
ductive organs [Sharga 1955]; predators [Sharga 1953]. 

*Cochlochila capeneri Drake and Slater 

Cochlochila capeneri Drake and Slater 1955, p. 49 [Transvaal]. — Drake 
1961c, p. 129 [Natal; Cape Province]. 
Type: Holotype 9, macropterous; Kaalfontein, Pretoria, South Africa; 
Drake Coll. (USNM). 
Distribution: South Africa (Transvaal; Natal; Cape Province). 
Host Plant: Unrecorded. 

Note: Type locality incorrectly given as "Rustenburg, South Africa" 
[Drake]. 

Cochlochila exolenta Drake 

Cochlochila exolenta Drake 1957a, p. 416, fig. [Aden; jasmine]. 
Type: Holotype 9; Jebel Jihaf, Western Aden Protectorate, Arabia; 
British Mus. 
Distribution: Aden Protectorate. 
Host Plant: Wild jasmine. 

*Cochlochila ituriensis Schouteden 

Cochlochila ituriensis Schouteden 1953d, p. 167 [Congo]. — Drake 1958b, 
p. 29. 
Type: Sex unknown; Yebo Moto, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Cochlochila kilimensis Horvath 

Cochlochila kilimensis Horvath 1910, p. 67 [Tanganyika]. — Drake 1948i, 
p. 181 [Kenya; Rhodesia]; 1954e, p. 3 [Congo]; 1958b, p. 29. 
Type: Sex unknown; Kibongoto, Kilimandjaro; Stockholm Mus. 
Distribution: Tanganyika; Kenya; Congo; Northern Rhodesia. 
Host Plant: Unrecorded. 

*Cochlochila lewisi (Scott) 

Leptodictya? lewisi Scott 1880, p. 314 [Japan]. — Oshanin 1908, p. 450. — 

Takeya 1951a, p. 20. 
Monanthia conchata Matsumura 1913, p. 151, pi. 14, fig. 16; 1931, p. 1202. 
Physodictyon vesicarius Lindberg 1927, p. 17, pi. 1, fig. 1 [Siberia]. 
Cochlochila conchata: Esaki 1932, p. 1636, fig.; 1954, p. 236, fig. 611. — 

Saito 1934, p. 69 [Korea].— Drake 1948i, p. 179 [China].— Takeya 

1951a, p. 22 [Petasites japonica; Ligularia sibirica; Phytolacca acinosa; 

Cacalia thunbergii]; 1953a, p. 2. 
Cochlochila lewisi: Drake and Ruhoff 1961c, p. 150. 



LACEBUGS OF THE WORLD 121 

Type: Japan; sex and deposition of type unknown, presumably lost. 
Distribution: Japan; China; U.S.S.R. (Siberia); Korea. 
Host Plants: Cacalia thunbergii; Ligularia sibirica; Petasites japonica; 
Phytolacca acinosa. 

Note: Accredited wrongly to Horvath by Matsumura (1913). 
*Cochlochila naivashae Drake 

Cochlochila naivashae Drake 1948i, p. 180 [Kenya]. 
Type: Holotype cf ; Naivasha, Kenya; Coryndon Mus. 
Distribution: Kenya. 
Host Pla^it: Unrecorded. 
^Cochlochila nilgiriensis (Distant) 

Monanthia nilgiriensis Distant 1903a, p. 50 [India]; 1903b, p. 144. — 

Drake 1933, p. 1015.— Singh 1953, p. 118. 
Cochlochila nilgiriensis: Drake 1948i, p. 181. — Gomez-Menor 1956a, 
fig. 4 (fig. only). 
Type: Sex unknown; Nilgiri Hills, India; British Mus. 
Distribution: India; Ceylon. 
Host Plant: Tectona grandis. 
Cochlochila yemenana Drake 

Cochlochila yemenana Drake 1957a, p. 417, fig. 2 [Yemen]. 
Type: Holotype o 71 ; Jebel Sumara, Yemen, Arabia; British Mus. 
Distribution: Yemen. 
Host Plant: Unrecorded. 
*Cochlochila zetana Drake 

Cochlochila zetana Drake 1954c, p. 70 [Orange Free State]. 
Type: Holotype 9, macropterous; Bloemfontein, Orange Free State; 
Drake Coll. (USNM). 
Distribution: South Africa (Orange Free State). 
Host Plant: Unrecorded. 

Genus CODOTINGIS Drake 

Codotingis Drake 1942b, p. 360.— Drake and Ruhoff 1960a, p. 44. 
Type Species: Codotingis recurva Drake. 
^Codotingis evansi Drake Plate 27 

Codotingis evansi Drake 1961b, p. 108, pi. 4 [Queensland]. 
Type: Holotype cf; Moruen District, Queensland, Australia; Austr. Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Codotingis recurva Drake 

Codotingis recurva Drake 1942b, p. 361 [Queensland]. 
Type: Holotype 9, macropterous; Nanango District, Queensland, 
Australia; Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

681-552—65 9 



122 U.S. NATIONAL MUSEUM BULLETIN 243 

Genus COLEOPTERODES Philippi 

Solenostoma Signoret 1863, p. 575. — Stal 1873, p. 117. — Lethierry and 
Severin 1896, p. 9.— Reed 1900, p. 178 (p. 85, separate).— Drake 
and Ruhoff 1960a, p. 25. 
Coleopterodes Philippi 1864, p. 306.— Kirkaldy 1900, p. 241— Bergroth 
1922, p. 152.— Drake and Ruhoff 1960a, p. 24. 
Type Species: Solenostoma liliputiana Signoret = Coleopterodes juscescens 
Philippi. 
*Coleopterodes brunnea Drake and Poor 

Coleopterodes liliputiana var. brunnea Drake and Poor 1938b, p. 104 

[Argentina; Baccharis pingraea]. 
Coleopterodes brunnea: Drake 1944a, p. 67. 
Type: Holotype 9; Argentina; La Plata Mus. 
Distribution : Argentina. 
Host Plant: Baccharis pingraea. 

*Coleopterodes liliputiana (Signoret) 

Solenostoma liliputiana Signoret 1863, p. 575, pi. 13, fig. 27 [Chile]. — 
Stal 1873, p. 117. — Berg 1884, p. 100 [Argentina; Baccharis pingraea]. — 
Reed 1900, p. 178 (p. 85, separate).— Pennington 1921, p. 20 — 
Bosq 1937, p. 131. 
Coleopterodes juscescens Philippi 1864, p. 308.— Walker 1873b, p. 4. 
Monanthia liliputiana: Walker 1873a, p. 194. 
Solenostoma lipiputianum [sic]: Champion 1898b, p. 56. 
Coleopterodes liliputianum: Drake 1922b, p. 50. 

Coleopterodes liliputiana: Drake 1922c, p. 353, pi. 39, fig. 13; 1939c, p. 
332; 1948h, p. 76 [Acacia cavenia]. 
Type: Chile; sex and deposition of type unknown. 
Distribution: Chile; Argentina. 
Host Plants: Acacia cavenia; Baccharis pingraea. 

Genus COLLINUTIUS Distant 

Collinutius Distant 1903b, p. 134.— Drake and Ruhoff 1960a, p. 44. 
Type Species: Tingis alicollis Walker. 

*Collinutius alicollis (Walker) 

Tingis alicollis Walker 1873a, p. 182 [India]. 
Phyllontochila alicollis: Distant 1902b, p. 354. 

Collinutius alicollis: Distant 1903b, p. 135, fig. 98.— Drake and Maa 
1954, p. 115 [China]. 
Type: Sex unknown; Hindostan; British Mus. 
Distribution: India; China. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 123 

Genus COMPSEUTA Stal 

Monanthia {Compseuta) Stal 1873, p. 133. 

Compseuta: Distant 1904, p. 433; 1910, p. 105.— Horvath 1910, p. 70 — 
Drake 1948f, pp. 197-204.— Drake and Runoff 1960a, p. 44. 
Type Species: Tropidocheila ornatella Stal. 
*Compseuta bioculata Drake 

Compseuta bioculata Drake 1948f, pp. 202, 203, fig. 1 [Nigeria]. 
Type: Holotype 9, macropterous; Cameroon Mts., Nigeria; Drake 
Coll. (USNM). 
Distribution: Nigeria; South Africa. 
Host Plant: Unrecorded. 
*Compseuta bispinosa Drake 

Compseuta bispinosa Drake 1951, p. 173 [Tanganyika]. — Drake and 

Ruhoff 1961c, p. 145. 
Compseuta kittenbergeri Drake 1951, p. 173. 
Type: Holotype cf ; Arusha, Africa orientalis; Hungarian Mus. 
Distribution: Tanganyika; Egypt. 
Host Plant: Unrecorded. 
*Compseuta capensis (Walker) 

Piesma capensis Walker 1873b, p. 6 [Natal]. 
Monanthia capensis: Distant 1902a, p. 243; 1902b, p. 356. 
Compseuta capensis: Drake 1948f, p. 202. 
Type: Sex unknown; Natal, South Africa; British Mus. 
Distribution: South Africa (Natal); Uganda. 
Host Plant: Unrecorded. 
*Compseuta comes Drake 

Compseuta comes Drake 1951, p. 172 [Tanganyika]. 
Type: Holotype o 71 ; Arusha, Africa orientalis; Hungarian Mus. 
Distribution : Tanganyika. 
Host Plant: Unrecorded. 
*Compseuta cordiae Drake 

Compseuta cordiae Drake 1948f, pp. 200, 203 [Uganda; Cordia]; 1954g, 
p. 665 [Ethiopia; Cordia abyssinica]. — Schouteden 1957c, p. 318 
[Congo]. 
Type: Holotype 9, macropterous; Kampola, Uganda; Drake Coll. 
(USNM). 
Distribution: Uganda; Congo; South Africa; Ethiopia; Tanganyika. 
Host Plants: Cordia abyssinica; Cordia sp. 
*Compseuta dispar Schouteden 

Compseuta dispar Schouteden 1923, p. 109 [Congo]. 
Type: Sex unknown; Madyu, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Uganda. 
Host Plant: Unrecorded. 



124 U.S. NATIONAL MUSEUM BULLETIN 243 

Compseuta expleta Drake 

Compseuta expleta Drake 1955b, p. 85 [Angola]. 
Type: Holotype &; Serra do Moco, Luimbale, Angola; Cent. Afr. Mus. 
Distribution: Angola. 
Host Plant: Unrecorded. 

*Compseuta holana Drake 

Compseuta holana Drake 1948f, pp. 199, 203 [Zanzibar; Ethiopia; 
Tanganyika]. 
Type: Holotype o 71 , macropterous; Zanzibar; Drake Coll. (USNM). 
Distribution: Zanzibar; Tanganyika; Ethiopia; Uganda; Congo; South 
Africa. 
Host Plant: Unrecorded. 

*Compseuta lamellata Drake 

Compseuta lamellata Drake 1958a, p. 105 [Transvaal]. 
Type: Holotype cf, macropterous; El Andshoek, Eastern Transvaal, 
South Africa; Drake Coll. (USNM). 
Distribution: South Africa (Transvaal). 
Host Plant: Unrecorded. 

*Compseuta latipennis Horvath 

Compseuta latipennis Horvath 1910, p. 70 [Tanganyika]. 
Type: Sex unknown; Kibonota, Kilimandjaro; Stockholm Mus. 
Distribution: Tanganyika. 
Host Plant: Unrecorded. 

*Compseuta lefroyi Distant 

Compseuta lejroyi Distant 1909a, p. 113 [India]; 1910a, p. 106, fig. 52 — 
Drake 1948a, p. 4 [Java; Lantana]. — Drake and Ruhoff 1962a, p. 
156; 1962b, p. 250. 
Compseuta tropica Hacker 1928, p. 184, fig. 1 [Queensland]. — Drake 

1948a, p. 4. 
Compseuta tessellata Drake and Poor 1936a, p. 441 [Hainan Island]. — 

Drake 1948a, p. 4 [China]. 
Compseuta lefroyi var. tropica: Drake 1948f, pp. 197, 203. 
Compseuta lefroyi var. tessellata: Drake 1948f, pp. 197, 203. 
Type: Sex unknown; Pusa, Bengal, India; British Mus. 
Distribution: India; Australia (Queensland); China (Hainan Island); 
Greater Sunda Islands (Java). 
Host Plants: Lantana sp. ; Cordia myxa. 

*Compseuta montandoni Distant 

Compseuta montandoni Distant 1904, p. 434, pi. 8, fig. 15 [South Africa]. — 
Horvath 1910, p. 71 [Tanganyika].— Drake 1948f, p. 201.— Priessner 
and Alfieri 1953, p. 65 [Egypt; Triumfetta flavescens].— Drake and 
Slater 1955, p. 49 [Natal]. 



LACEBUGS OF THE WORLD 125 

Type: Sex unknown; Rikatla, South Africa; British Mus. 
Distribution: South Africa (Natal) ; Tanganyika; Kenya; Egypt. 
Host Plant : Triumfetta flavescens. 
Compseuta montandoni var. simulans Horvath 

Compseuta montandoni var. simulans Horvath 1910, p. 71 [Tanganyika]. 
Type: Sex unknown; Kilimandjaro; Stockholm Mus. 
Distribution: Tanganyika. 
Host Plant: Unrecorded. 

Compseuta motoensis Schouteden 

Compseuta motoensis Schouteden 1955a, p. 30 [Congo]. 
Type: Sex unknown; Moto, Belgium Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Compseuta notialis Drake 

Compseuta notialis Brake 1948f, pp. 199, 204 [Uganda]. 
Type: Holotype 9, macropterous; Masaka, Uganda; Drake Coll. 
(USNM). 
Distribution: Uganda; Egypt; South Africa. 
Host Plant: Unrecorded. 

*Compseuta ornatella (Stal) 

Tropidocheila ornatella Stal 1855a, p. 37 [Caffraria]. 

Monanthia (Physatochila) ornatella: Stal 1865, p. 28. 

Monanthia {Compseuta) ornatella: Stal 1873, p. 133. 

Monanthia ornatella: Walker 1873a, p. 195.— Distant 1902a, p. 242, 

pi. 15, fig. 9. 
Compseuta ornatella: Distant 1904, p. 433.- — Schouteden 1916b, p. 292 
[Congo]; 1923, p. 108; 1957c, p. 318 [Ruanda].— Drake 1948f, 
p. 198; 1954e, p. 3; 1955b, p. 86 [Uganda; Kenya; Angola]; 1958b, 
p. 28 [Rhodesia]. — Drake and Gomez-Menor 1954, p. 91 [Spanish 
Guinea]. 
Monanthia nigristernum Horvath 1910, p. 69 [Tanganyika]. 
Type: Sex unknown; Caffraria, South Africa; Stockholm Mus. 
Distribution: South Africa; Northern Rhodesia; Congo; Kenya; 
Tanganyika; Angola; Uganda; Ruanda-Urundi; Spanish Guinea. 
Host Plant: Unrecorded. 

♦Compseuta ornatella var. biseriata Drake 

Compseuta ornatella var. biseriata Drake 1948f, pp. 198, 204 [Uganda]. 
Type: Holotype 9, macropterous; Entebbe, Uganda; Drake Coll. 
(USNM). 
Distribution: Uganda; South Africa. 
Host Plant: Unrecorded. 



126 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Compseuta ornatella var. carinata Drake 

Compseuta tenella [sic] subsp. carinata Drake 1958b, p. 28 [Congo]. 
Type: Holotype 9; Lusinga, Belgian Congo; Pares Nat. Inst. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Compseuta ornatella var. teres Drake (emendation) 

Compseuta ornatella var. teretis Drake 1948f, pp. 199, 204 [Uganda]; 
1954a, p. 232 [Guinea]. 
Type: Holotype 9, macropterous; W. Nile, Uganda; Drake Coll. 
(USNM). 
Distribution: Uganda; Guinea; South Africa. 
Host Plant: Unrecorded. 

*Compseuta picta Schouteden 

Compseuta picta Schouteden 1923, p. 108 [Congo]. — Drake 1954e, p. 3; 
1955b, p. 86 [Angola]. — Drake and Gomez-Menor 1954, p. 91 
[Spanish Guinea; Cameroons]. — Gomez-Menor 1956a, p. 112 
[Malvaceae]. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Angola; Cameroons; Spanish Guinea. 
Host Plant: Malvaceae. 

Compseuta picta var. brevicarinata Schouteden 

Compseuta picta var. brevicarinata Schouteden 1923, p. 108 [Congo]. 

Type: Sex unknown; Kamaiembi, Kasai, Belgian Congo; Cent. Afr. 
Mus. 

Distribution: Congo. 

Host Plant: Unrecorded. 

*Compseuta picta var. funebris Schouteden 

Compseuta picta var. funebris Schouteden 1923, p. 109 [Congo]. 
Type: Sex unknown; Kamaiembi, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Compseuta sejuncta Drake 

Compseuta sejuncta Drake 1948f, pp. 201, 204 [Egypt]. 
Type: Holotype o 71 , macropterous; Shenki, Egypt; Drake Coll. (USNM). 
Distribution: Egypt. 
Host Plant: Unrecorded. 

Genus CONCHOTINGIS Drake 

Conchotingis Drake 1954c, p. 71. — Drake and Ruhoff 1960a, p. 45. 
Conchochila Drake 1958c, p. 329.— Drake and Ruhoff 1960a, p. 45. 
Type Species: Xenotingis trepidantis Drake. 



LACEBUGS OF THE WORLD 127 

♦Conchotingis borneoana Drake and Ruhoff 

Conchotingis borneoana Drake and Ruhoff 1961 b, p. 172, fig. 17a [Borneo]. 

Type: Holotype 9, macropterous; Sandakan, Borneo; Drake Coll. 
(USNM). 

Distribution: Greater Sunda Islands (Borneo). 

Host Plant: Unrecorded. 

Conchotingis insulana (Drake) 

Conchochila insulana Drake 1958c, p. 331 [Malagasy]. 

Conchotingis insulana: Drake and Ruhoff 1960a, p. 45. 
Type: Holotype c? ; Perinet, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

♦Conchotingis sundra (Drake) 

Conchochila sundra Drake 1958c, p. 329, fig. 1 [Malagasy]. 

Conchotingis sundra: Drake and Ruhoff 1960a, p. 45. 
Type: Holotype o 71 ; Perinet, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

♦Conchotingis trepidans (Drake) emendation 

Xenotingis trepidantis Drake 1927d, p. 310 [Malagasy]. 
Conchotingis trepidantis: Drake 1954c, p. 71; 1958c, p. 332. 

Type: Holotype 9, macropterous; Madagascar; Drake Coll. (USNM). 

Distribution: Malagasy Republic. 

Host Plant: Unrecorded. 

Genus CONGOCHILA Drake 

Congochila Drake 1954e, p. 8. — Drake and Ruhoff 1960a, p. 45. 
Type Species: Congochila congoana Drake. 

♦Congochila congoana Drake 

Congochila congoana Drake 1954e, p. 9 [Congo]. 
Type: Holotype o"; Libenge, Savane, Liki-Bembe, Belgian Congo; 
Brussels Mus. 

Distribution: Congo. 
Host Plant: Unrecorded. 

Genus COPIUM Thunberg 

Copium Thunberg 1822, p. 8. — Lethierry and Severin 1896, p. 15. — 
Horvath 1906a, pp. 14, 91 (key).— Oshanin 1908, p. 445; 1912, 
p. 45.— Stichel 1926, pp. 104, 114 (key); 1935, p. 349; 1938a, p. 408; 
1960a, p. 326; 1960b, p. 399.— de Seabra 1931, p. 430.— Guide 
1938, pp. 244, 304 (key) .— Kiritshenko 1951, pp. 242, 251.— Takeya 
1951a, p. 18.— Singer 1952, p. 53.— Wagner 1954, p. 200 (key); 



128 U.S. NATIONAL MUSEUM BULLETIN 243 

1956, p. 86 (key). — Hoberlandt 1955, p. 93. — Monod and Carayon 

1958, pp. 1-30.— Drake and Ruhoff 1960a, p. 45. 
Monanthia Le Peletier and Serville 1828, p. 653. — Drake and Runoff 

1960a, p. 68. 
Eurycera Laporte 1833, p. 49— Stal 1874, p. 57.— Rey 1888, p. 190. — 

Drake and Ruhoff 1960a, p. 56. 
Laccometopus Fieber 1844, pp. 30, 96; 1861, p. 119. — Drake and Ruhoff 

1960a, p. 63. 
Type Species: Copium cornutum Thunberg = Cimex clavicornis Linnaeus. 

*Copium adumbratum (Horvath) 

Eurycera adumbrata Horvath 1891, p. 79 [Armenia]. 

Copium adumbratum: Lethierry and Severin 1896, p. 15. — Horvath 
1906a, p. 93 [Syria].— Oshanin 1908, p. 446 [Transcaucasus]. — 
Seidenstiicker 1954, p. 236 [Turkey].— Wagner 1954, pp. 202, 209, 
figs. 1, 8, 15, 22; 1956, p. 86.— Hoberlandt 1955, p. 93.— Stichel 
1960a, p. 326. 
Paracopium adumbratum: Horvath 1929, p. 322. 
Type: Holotype 9; Armenia; Hungarian Mus. 

Distribution: U.S.S.R. (Armenia; Transcaucasus); Syria; Turkey. 
Host Plant: Unrecorded. 

*Copium bernardi Wagner 

Copium bernardi Wagner 1954, pp. 203, 209, figs. 5, 13, 20, 27 [Algeria; 
Teucrium polium]; 1956, p. 86. — Monod and Carayon 1958, p. 5. — 
Stichel 1960a, p. 329. 
Type: Holotype cf; T'in Taradjeli, Tassili-Plateau, Sahara; Wagner 
Coll. 
Distribution: Algeria. 
Host Plant: Teucrium polium. 

*Copium brevicorne (Jakovlev) 

Eurycera brevicornis Jakovlev 1880b, p. 103 [Caucasus]. — Rey 1888, p. 

191. 
Copium brevicorne: Lethierry and Severin 1896, p. 15. — Wagner 1954, 
pp. 200, 203, 206, 209, figs. 6, 12, 19, 26 [Greece; Yugoslavia]; 1956, 
p. 86.— Monod and Carayon 1958, p. 5.— Stichel 1960a, p. 331 
[Cyprus]. 
Copium teucrii var. brevicorne: Horvath 1906a, p. 93 [Tunisia; Armenia; 
Turkey; Syria]. — Oshanin 1908, p. 446 [Rumania]. — Kiritshenko 
1951, p. 252.— Hoberlandt 1955, p. 93 [Transcaucasus; Israel]. 
Type: Sex unknown; Derbent, Dagestan, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Dagestan; Caucasus; Transcaucasus; Armenia); 
Turkey; Syria; Israel; Rumania; Yugoslavia; Greece; Cyprus; Tunisia. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 129 

*Copium clavicorne (Linnaeus) Plates 18, 19, 20 

Cimex clavicornis Linnaeus 1758, p. 442, (no. 12); 1761, p. 246 (no. 911); 
1767, p. 717 (no. 16).— Houttuyn 1765, p. 339.— Muller 1774, p. 
482. — Fourcroy 1785, p. 212 [Teucrium chameadrys]. — Gmelin 1790, 
p. 2125. 

Acanthia clavicornis: Fabricius 1775, p. 694; 1781, p. 336; 1787, p. 
278; 1794, p. 70.— Roemer 1789, p. 79, fig. 4.— Rossi 1790, p. 223 
[Italy].— Panzer 1794, heft 23, tab. 23.— Walckenaer 1802, p. 337 
[France]. 

Tingis clavicornis: Fabricius 1803, p. 124. — Latreille 1804, p. 251. — 
Vallot 1829, p. 99 [Teucrium montanum]. — Herrich-Schaeffer 1835, 
p. 57. 

Tingis punctata Lamarck 1816, p. 504. 

Copium cornutum Thunberg 1822, p. 8; 1825, p. 9. — Horvath 1906a, 
p. 92 [Greece; Rumania; Turkey]; 1916, p. 9 [Albania]. — Muzik 
1907, p. 61, fig. 5.— Stichell926, p. 114; 1938a, p. 408 [Poland].— 
de Seabra 1931, pp. 431, 444, figs. 502(1-4), 503 [Portugal].— Bor- 
ner 1935, p. 78, fig. 115.— Scholte 1935, p. 77, fig. 21 [Netherlands].— 
Guide 1938, p. 305, fig.— Hoberlandt 1942, p. 126; 1943a, p. 119; 
1943b, p. 124; 1955, p. 93 [Israel].— Blote 1945, p. 85.— Kiritshenko 
1951, p. 252.— Behr 1952, p. 326, figs. 1-6.— Bator 1953, p. 326, 
pi. 3, fig. 2.— Mancini 1953a, p. 186.— Seidenstiicker 1954, p. 236 — 
Wagner 1954, pp. 200, 202, 209, figs. 3, 10, 17, 24; 1956a, p. 86— Go- 
mez-Menor 1955b, p. 249 [Spain]. — Monod and Carayon 1958, 
pp. 5, 8, figs. 1-7 [Teucrium flavum; Teucrium marum]. 

Monanthia clavicornis: Le Peletier and Serville 1828, p. 653. — Westwood 
1840a, p. 478; 1840b, p. 121 [England]. 

Eurycera nigricornis Laporte 1833, p. 49. — Burmeister 1835, p. 258. — 
Spinola 1837, p. 167.— Amyot and Serville 1843, p. 296. 

Eurycera clavicornis: Brulle 1835, p. 341, pi. 26, fig. 2. — Herrich-Schaeffer 
1838, p. 65, pi. 129, fig. 400a.— Blanchard 1840, p. 113.— Jakovlev 
1875b, p. 262; 1880b, p. 103— Reiber and Puton 1876, p. 69.— Andre 
1876, p. 34; 1878a, p. 25.— Ferrari 1878, p. 66.— Puton 1879c, p. 106 
[Teucrium scorodonium]. — Rey 1888, p. 190, fig. — Dubois 1888, p. 
120.— d'Antessanty 1890, p. 31— Hiieber 1893, p. 326. 

Laccometopus clavicornis: Fieber 1844, p. 97, pi. 8, figs. 10-16 [Czecho- 
slovakia]; 1861, p. 119.— Frauenfeld 1853, p. 160.— Mayr 1858, 
p. 571 [Austria].— Garbiglietti 1869, p. 272.— Gredler 1870, p. 75 
[Germany].— Horvath 1874b, p. 432 [Hungary].— Douglas 1877, 
p. 236.— Buchanan-White 1877, p. 283.— Rubsaamen 1896, p. 428, 
pi. 15, fig. 8.— Drake and Ruhoff 1960a, p. 63. 

Tingis (Laccometopus) clavicornis: Kolenati 1856, p. 426 [Caucasus; 
Transcaucasus; Teucrium canum; Teucrium scordium]. 

Laccometohus [sic] clavicornis: Frey-Gessner 1865, p. 231 [Switzerland] 



130 U.S. NATIONAL MUSEUM BULLETIN 243 

Dictyonota nigricornis: Walker 1873a, p. 176. 
Eurycera cornuta: Reuter 1881, p. 170. 

Copium clavicorne: Lethierry and Severin 1896, p. 15. — Reichensperger 
1920, p. 61.— Ross and Hedicke 1927, p. 281.— Stichel 1960a, p. 
328, fig. 137 [Belgium; Sardinia; Yugoslavia; Bulgaria]. 
Copium clavicornis: Wagner 1954, pp. 200, 209. — Drake and Davis 1960, 
fig. 2.— Drake and Runoff 1960a, pp. 45-47, pi. 4. 
Type: "Europa"; Stockholm Mus.; sex unknown. 

Distribution: England; Belgium; Netherlands; France; Spain; Portugal; 
Sardinia; Italy; Switzerland; Germany; Austria; Czechoslovakia; Hungary; 
Rumania; Greece; Albania; Poland; Bulgaria; Yugoslavia; Turkey; 
U.S.S.R. (Caucasus; Transcaucasus) ; Israel. 

Host Plants: Teucrium chamaedrys ; Teucrium canum; Teucrium flavum; 
Teucrium marum; Teucrium montanum; Teucrium scordium; Teucrium scorodonium; 
Teucrium sp. 

Note: 1. The following references concern Cimex clavicornis: Cimex sp. 
687 Linnaeus 1746, p. 212; Cimex sp. 56 Geoffroy 1762, p. 461 (cited Lin- 
naean references and that specimens were feeding on flowers of chamaedry); 
(unnamed insect) Reaumur 1737, p. 427a, pi. 34, fig. 3, 4 (contains a good 
discussion of gall-making insect and figure of galls on " Fluer du camedry"). 
2. Host plant damage [Douglas; Behr]. 3. Discussion on nomenclature 
and history [Drake and Ruhoff]. 
*Copium horvathi Wagner 

Copium horvathi Wagner 1957, p. 28, figs. 1-5 [Turkey; Greece]. — Stichel 
1960a, p. 331 [Yugoslavia; Bulgaria; Teucrium]. 
Type: Unknown. 

Distribution: Turkey; Greece; Yugoslavia; Bulgaria. 
Host Plant: Teucrium sp. 

*Copium intermedium (Rey) 

Eurycera intermedia Rey 1888, p. 191, fig. [Corsica; Sardinia]. 
Copium intermedium: Lethierry and Severin 1896, p. 16. — Wagner 1954, 
pp. 200, 207, 209, figs. 7, 14, 21; 1956, pp. 84, 86, figs. 1-6.— Stichel 
1960a, p. 330. 
Copium teucrii var. intermedium: Horvath 1906a, p. 93 [Bulgaria]. — 
Mancini 1935, p. 9 [Capraia]. — Novak and Wagner 1951, p. 71 
[Yugoslavia; Juniperus phoenicea; Helichrysum italicum], — Gomez-Menor 
1955b, p. 249 [Spain]. 
Type: Unknown. 

Distribution: Spain; Italy (Capraia Island); Corsica; Sardinia; 
Bulgaria; Yugoslavia. 
Host Plants: Helichrysum italicum; Juniperus phoenicea. 



LACEBUGS OF THE WORLD 131 

*Copium japonicum Esaki 

Laccometopus clavicornis (not Linnaeus): Scott 1874, p. 291 [Japan; 

Keiskea japonica] . 
Copium japonicum Esaki 1931, pp. 244, 251, fig. 2; 1932, p. 1637, fig.; 
1954, p. 237, fig. 614.— Takeya 1931, p. 81; 1951a, p. 19 [Taiwan; 
Mosla punctata]. — Monod and Carayon 1958, p. 5. 
Paracopium japonicum: Drake 1937b, p. 591 [China]. 
Type: Holotype cf; Mino, Honshu, Japan; Kyushu Univ. 
Distribution: Japan; China; Taiwan. 
Host Plants: Keiskea japonica; Mosla punctata. 

*Copium magnicorne (Rey) 

Eurycera magnicornis Rey 1888, p. 191, fig. [France]. 
Copium magnicorne: Lethierry and Severin 1896, p. 16. — Wagner 1954, 
p. 208; 1955, p. 182, figs. 1-6; 1956, p. 86; 1958, p. 241 [Teucrium 
montanum; Teucrium chamaedrys]. — Monod and Carayon 1958, p. 5. — 
Stichel 1960a, p. 330. 
Type: Sex unknown; Rouen, France; Lyon Mus. 
Distribution: France. 
Host Plants: Teucrium chamaedrys; Teucrium montanum. 

*Copium reyi Wagner 

Copium reyi Wagner 1954, pp. 203, 205, 209, figs. 2, 9, 16, 23 [France; 

Teucrium scorodonium]; 1956, p. 86. — Monod and Carayon 1958, p. 

5.— Stichel 1960a, p. 327. 
Type: Holotype cf; Madeloc, France; Wagner Coll. 
Distribution: France. 
Host Plant: Teucrium scorodonium. 

*Copium teucrii (Host) Plates 21, 22 

Cimex teucrii Host 1788, p. 255, fig. [Teucrium supinum]. 

Tingis teucrii: Westwood 1840a, p. 478. 

Laccometopus teucrii: Frauenfeld 1853, p. 160 [Teucrium montanum]. — 
Mayr 1858, p. 571 [Austria; Russia].— Fieber 1861, p. 119 [Italy].— 
Garbiglietti 1869, p. 272. — Gredler 1870, p. 75 [Germany; Teucrium 
chamaedrys]. — Horvath 1874b, p. 432 [Hungary]. — Riibsaamen 1896, 
p. 429, pi. 16, figs. 1, 4, 37, 38 [Teucrium polium]. 

Monanthia teucrii: Walker 1873a, p. 188. 

Eurycera teucrii: Puton 1879c, p. 106 [Corsica; France]. — Jakovlev 
1880b, p. 103.— Rey 1888, p. 190, fig.; 1893, p. 97.— Dubois 1888, 
p. 121.— d'Antessanty 1890, p. 31.— Hueber 1893, p. 328. 

Copium teucrii: Lethierry and Severin 1896, p. 16. — Lambertie 1906, 
p. 25. — Horvath 1906a, p. 93 [Teucrium capitatum; Switzerland; 
Yugoslavia]. — Oshanin 1908, p. 445 [Spain; Caucasus; Crimea]. — 
Reuter 1908, p. 89.— Moroder Sala 1920, p. 12.— Stichel 1926, p. 
114; 1938a, p. 408 [Albania]; 1960a, p. 329 [Helichrysum angustifolium; 



132 U.S. NATIONAL MUSEUM BULLETIN 243 

Belgium; Sardinia; Greece; Bulgaria]. — Lindberg 1932a, p. 43 
[Morocco]; 1948, p. 59 [Cyprus].— Scholte 1935, p. 77 [Nether- 
lands] .— Borner 1935, p. 78.— Ross and Hedicke 1927, p. 281 — 
Vidal 1937, p. 199.— Blote 1945, p. 85.— Gonzalez 1948, p. 51.— 
Kiritshenko 1951, p. 252.— Bator 1953, p. 326.— Priesner and Alfieri 
1953, p. 65 [Egypt].— Seidenstucker 1954, p. 236 [Turkey].— 
Wagner 1954, pp. 203, 209, figs. 4, 11, 18, 25; 1956, p. 86— Hober- 
landt 1955, p. 93 [Transcaucasus; Israel]. — Monod and Carayon 
1958, p. 5, figs. 8-16. 
Copium lusitanicum de Seabra 1924, p. 17, figs. 29, 30 [Portugal]. — 

Wagner 1954, p. 209. 
Copium teucri [sic]: Gomez-Menor 1956a, p. 115, fig. 11 (fig. only). 
Type: Unknown. 

Distribution: Portugal; Spain; France; Netherlands; Belgium; Ger- 
many; Austria; Italy; Corsica; Sardinia; Switzerland; Hungary; Yugo- 
slavia; Greece; Cyprus; Albania; Bulgaria; Turkey; U.S.S.R. (Caucasus; 
Transcaucasus; Crimea); Israel; Morocco; Alberia; Egypt. 

Host Plants: Helichrysum angustijolium; Teucrium capitatum; Teucrium 
chamaedrys; Teucrium montanum; Teucrium polium; Teucrium supinum; Teucrium 
sp. 

Genus CORINTHUS Distant 

Corinthus Distant 1920, p. 155.— Drake and Ruhoff 1960a, p. 47. 
Type Species: Corinthus typicus Distant. 

Corinthus typicus Distant 

Corinthus typicus Distant 1920, p. 156 [New Caledonia]. — Drake and 
Davis 1960, fig. 42. 
Type: Sex unknown; Houadou, New Caledonia; British Mus. 
Distribution: New Caledonia. 
Host Plant: Unrecorded. 

Genus CORYCERA Drake 

Corycera Drake 1922c, p. 368.— Monte 1939b, p. 66; 1941e, p. 84 — 
Hurd 1946, p. 457.— Drake and RuhofF 1960a, p. 47. 
Type Species: Corycera comptula Drake. 

*Corycera alboater Drake and Hambleton 

Corycera alboater Drake and Hambleton 1935, p. 143 [Brazil; Dahlstedtia 
pinnata] . 
Type: Holotype cf, macropterous; Estacao Biologica, Alta da Serra, 
Sao Paulo, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Dahlstedtia pinnata. 



LACEBUGS OF THE WORLD 133 

*Corycera albocosta (Drake) 

Gelchossa albocosta Drake 1922c, p. 372, pi. 39, fig. 2 [Brazil]. 

Leptopharsa albocosta: Monte 1941e, p. 109. 

Corycera albocosta: Drake and Runoff 1961c, p. 145. 
Type: Holotype o"; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Corycera comptula Drake 

Corycera comptula Drake 1922c, p. 369, pi. 39, fig. 3 [Brazil]; 1930a, p. 1; 
1935, p. 12 [Paraguay].— Drake and Hambleton 1944b, p. 124. 
Type: Holotype o"; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil; Paraguay. 
Host Plant: Unrecorded. 

Corycera fallax Monte 

Corycera fallax Monte 1942a, p. 92 [Brazil]. 

Type: Holotype cf ; Palmeiras, Ipaussu, Sao Paulo, Brazil; Monte Coll. 
(Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

* Corycera fortis Drake 

Corycera jortis Drake 1942a, p. 18 [Paraguay]. 
Type: Holotype d% macropterous; Chaco, Paraguay; Drake Coll. 
(USNM). 
Distribution: Paraguay. 
Host Plant: Unrecorded. 

*Corycera gibbosa Monte 

Corycera gibbosa Monte 1938c, p. 64, fig. [Brazil; Roupala]; 1940c, p. 191, 

fig. 25; 1940e, p. 296. 
Corycera rhopalae Drake and Poor 1938a, p. 29. 
Type: Holotype o 71 ; Belo Horizon te, Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Roupala sp. 

*Corycera juncta Drake and Hambleton 

Corycera juncta Drake and Hambleton 1944b, p. 124 [Brazil]. 

Type: Holotype 9, macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 



134 U.S. NATIONAL MUSEUM BULLETIN 243 

*Corycera machaerii Drake and Hambleton 

Corycera machaerii Drake and Hambleton 1935, p. 142 [Brazil; Machae- 
rium]. 
Type: Holotype cf, macropterous; Ponte Nova, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Machaerium sp. 

:1: Corycera panamensis Drake and Poor 

Corycera panamensis Drake and Poor 1938a, p. 30 [Canal Zone]. 

Type: Holotype 9, macropterous; Panama Canal Zone; Drake Coll. 
(USNM). 

Distribution: Panama (Canal Zone). 

Host Plant: Unrecorded. 

*Corycera pusilla Monte 

Corycera pusilla Monte 1942a, p. 92 [Brazil]. 

Type: Holotype cf; Bocaina, Sao Paulo, Brazil; Monte Coll. (Mus. 
Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

Corycera rochalimai Monte 

Corycera rocha-limai Monte 1940e, p. 296, fig. 6 [Brazil]. 

Type: Holotype o 71 ; Belo Horizonte, Minas Gerais, Brazil; Monte Coll. 
(Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Corycera rugulosa Drake 

Corycera rugulosa Drake 1922c, p. 369 [Brazil]. — Monte 1939b, p. 67 
[Erythroxylon]. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Erythroxylon sp. 

Corycera schubarti Monte 

Corycera schubarti Monte 1946b, p. 282 [Brazil]. 

Type: Holotype o*; Pirassununga, Sao Paulo, Brazil; Monte Coll. 
(Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Corycera separata (Drake and Hambleton) 

AmbJystira separata Drake and Hambleton 1935, p. 142 [Brazil; Ery- 
throxylon].— Monte 1938b, p. 128. 
Corycera separata: Monte 1941e, p. 86. 



LACEBUGS OF THE WORLD 135 

Type: Holotype o 71 , macropterous; Itarare, Sao Paulo, Brazil; Drake 
Coll. (USNM). 

Distribution: Brazil; British Guiana. 

Host Plant: Erythroxylon sp. 
*Corycera spissa Drake 

Corycera spissa Drake 1935, p. 12, fig. 1 [Paraguay]. — Monte 1939b, 
p. 67 [Brazil; Croton anlisyphiliticus]; 1940c, p. 193. 

Type: Holotype o 71 ; S. Bernardino, Paraguay; Vienna Mus. 

Distribution: Paraguay; Brazil. 

Host Plant: Croton anti syphiliticus. 
*Corycera vallaris Drake 

Corycera vallaris Drake 1942, p. 18 [Paraguay]. 

Type: Holotype o 71 , macropterous; Horqueta, Paraguay; Drake Coll. 
(USNM). 

Distribution: Paraguay. 

Host Plant: Unrecorded. 

Genus CORYTHAICA Stal 

Corythaica Stal 1873, pp. 120, 128.— Champion 1897, p. 9; 1898b, p. 
58.— Van Duzee 1917b, p. 817.— Gibson 1919c, p. 98 (key).— 
Blatchley 1926, p. 470 (key).— Monte 1939b, p. 67; 1941e, p. 86; 
1942d, p. 104.— Hurd 1945, pp. 79-99 (monograph); 1946, p. 
480.— Bailey 1951, p. 47.— Drake and Runoff 1960a, p. 47. 
Typonotus Uhler 1893a, p. 716.— Drake and Ruhoff 1960a, p. 87. 
Dolichocysta Champion 1898b, p. 56.— Van Duzee 1916, p. 25; 1917b, 
p. 215.— Drake 1917a, p. 214 (key).— Gibson 1919c, p. 101 (key).— 
Drake and Ruhoff 1960a, p. 54. 
Leptotingis Monte 1938b, p. 128; 1939b, p. 75; 1941e, p. 121.— Drake 
and Ruhoff 1960a, p. 65. 
Type Species: Tingis monacha Stal. 
*Corythaica acuta (Drake) 

Dolichocysta acuta Drake 1917a, p. 214 [Colo.; Mont.]. 
Corythaica acuta: Gibson 1919c, p. 100.— Hurd 1945, pp. 81, 93, pi. 1, 
fig. 3. 
Type: Holotype cf , macropterous; Fort Collins, Colorado, U.S.; Drake 
Coll. (USNM). 

Distribution: U.S. (Colo.; Mont.; Nev.). 
Host Plant: Unrecorded. 
*Corythaica bellula Torre-Bueno 

Corythaica bellula Torre-Bueno 1917, p. 19 [N.Y.].— Gibson 1919c, p. 
100.— Blatchley 1926, p. 471.— Drake 1928b, p. 102; 1930d, p. 
268.— Hurd 1945, pp. 81, 93, pi. 1, fig. 2.— Bailey 1951, p. 48, fig. 2 
[Conn.; R.I.; Mass.; N.H.; Panicum lindheimeri]; 1959, p. 64 
[Maine].— Byers 1959, p. 193. 



130 U.S. NATIONAL Ml'SKlM HI I.LKTIN 243 

Corythaica thndann Blatchley 1926, p. 472 [Fla.]; 1928, p. 5. 
Type: Lcctotype v; White Plains, New York, U.S.; Snow Mus. 
Distribution: U.S. ( NIL; M Maine; Conn.; K.I.; N.Y.; Fla.; 

Nri.r.). 

Host I'i \m: Panicum lindheimeri. 

\< ite: Life history study [Bailey]; notes on type specimen [Byers]. 

*Corythaica bosqi Monte 

Corythaica bosqi Monte 1938d, p. 391 [Argentina]. — Hurd 1945, pp. 82, 
98, fig. 5. 
Type: Holotype d"; Santiago del Estero, Fortin Inca, Argentina; Monte 
Coll. (Mus. Nacional). 
Distribution: Argentina. 
Host Plant: Unrecorded. 

*Corythaica caestri (Reed) 

Tingis caestri Reed 1900, p. 181 (reprint, p. 88) [Chile]. 

Corythaica cucallata (not Berg): Drake and Poor 1938b, p. 108 (in part). 

Corythaica caestri: Drake 1939c, p. 333.— Hurd 1945, pp. 82, 91, fig. 3. 
Type: Holotype 9, macroptcrous; Chile; Drake Coll. (USNM). 
Distribuih >\: Chile. 
Host Plant: Unrecorded. 

*Corythaica carinata Uhler 

Corythaica carinata Uhler 1886, p. 22 (nov. nud.); 1894a, p. 203 [Gre- 
nada]. — Champion 1897, p. 9, pi. 1, figs. 11, 11a [Guatemala]; 
18981), p. 59 [St. Vincent].— Van Duzee 1907, p. 19 [Jamaica].— 
Gibson 1919c, p. 100 [Tex.; eggplant].— Wolcott 1923, p. 246 
[Puerto Rico].— Drake and Bruner 1924a, p. 151 [Cuba; Haiti]. — 
Gowdy 1926, p. 35.— Barber 1939, p. 369.— Monte 1943c, p. 114 
[Solarium melongena}. — Hurd 1945, pp. 82, 88, pi. 1, fig. 1 [Honduras; 
Dominican Republic].- — Bruner, Scaramuzza and Otero 1945, p. 11 
[Althaea rosea]. — Drake and Cobben 1960, p. 87, fig. 87 [Fla.; Mexico; 
Aruba; Curacao; St. Eustatius; St. Martin; Saba; Passiflora joctida; 
Corchorus hirsutus; Achyranthcs aspcra; Piriqucta ovata; Sida procumbens]. 
Corythaica constricta Osborn and Drake 1917b, p. 304 [Colo.]. 
Dolichocysta constrii ta: Gibson 1919c, p. 103. 
Type: Sex unknown; Mount Gay Estate, Grenada, British West Indies; 
Bi itish Mus. 

Distribution: Windward Islands (Grenada; St. Vincent); Haiti: 
Dominican Republic; Jamaica; Puerto Rico; Trinidad; Cuba; Netherlands 
Antilles (Aruba; Curacao); Leeward Islands (St. Eustatius; Saba; St. 
Martin); Honduras; Guatemala; Mexico; U S. rex.; Colo.; Fla.). 

Hosi Plants: Achyranthcs aspcra; Althaea rosea; Corchorus hirsutus; Passiflora 
foetida; Piriqucta ovata; Sida procumbens; So/anum melongena. 



LACEBUGS OF THE WORLD 137 

*Corythaica costata Gibson 

Corythaica costata Gibson 1919c, p. 99 [Peru; cotton]. — Fenton 1934, p. 
199. — Monte 1940e, p. 284 [Ecuador; Lycopersicum esculentum]; 
1943c, p. 114 [Gossypium].— Drake and Hambleton 1942a, p. 330; 
1945, p. 366 [Colombia].— Hurd 1945, pp. 82, 94, pi. 1, fig. 4. 
Type: Holotype d", macropterous; Santa Clara, Peru; USNM. 
Distribution: Peru; Ecuador; Colombia. 
Host Plants: Gossypium sp.; Lycopersicum esculentum. 

*Corythaica cucullata (Berg) 

Leptobyrsa cucullata Berg 1879a, p. 41 [Argentina]; 1879b, p. 135. — 

Pennington 1921, p. 20. 
Corythaica cucullata: Drake and Poor 1938b, p. 108, fig. 2. — Monte 
1940e, p. 284 [Sphaeralcea miniata]; 1943c, p. 114.— Hurd 1945, pp. 
82, 87, fig. 1. 
Type: Sex unknown; Buenos Aires Province, Argentina; La Plata Mus. 
Distribution: Argentina. 
Host Plant: Sphaeralcea miniata. 

♦Corythaica cyathicollis (Costa) 

Tingis cyathicollis Costa 1864, p. 146, pi. 2, fig. 4. 

Corythaica monacha (not Stal 1858): Stal 1873, p. 128 (in part).— Jones 
1915, p. 4 [Puerto Rico; eggplant; Solanum torvum]. — Cotton 1917, p. 
170. — Gibson 1919c, p. 99 [Solanum melongena; Ricinus communis] (in 
part).— Wolcott 1923, p. 246.— Townsend 1928, p. 19, fig. 20 — 
Bosql937,p. 128. 

Leptobyrsa passiflorae Berg 1884, p. 102 [Argentina; Passiflora caerulea]. — 
Pennington 1921, p. 20. 

Typonotus planaris Uhler 1893a, p. 716 [St. Vincent]; 1894a, p. 203 
[Grenada]. — Champion 1897, p. 9. 
Corythaica planaris: Drake and Bruner 1924a, p. 151. — Drake 
1932, p. 100 [Canal Zone].— Drake and Hambleton 1934, p. 451 
[Brazil; Solanum tuberosum; Solanum sisymbrifolium; Lycopersicum esculen- 
tum; Solanum racemiflorum]. — Costa Lima 1936, p. 126. — Monte 1937a, 
p. 31, fig. 9 [Solanum balbisii]; 1937b, p. 79, fig. [Solanum lycopersicum; 
Solanum gilo]; 1937c, p. 72; 1938d, p. 391.— Drake and Poor 1937d, 
p. 311. — Bosq 1937, p. 128 [Solanum bonariense; Solanum elaeagnifolium]. — 
Barber 1939, p. 369 [St. Thomas].— Soares 1941, p. 262, figs. 
2, 5, 6, 7.— Gomez- Menor 1956a, p. 112. 

Corythaica planaria [sic]: Drake 1935, p. 20. 

Corythaica cyathicollis: Drake and Poor 1938b, p. 108. — Monte 1939b, 
p. 67 [Solanum juripeba]; 1940d, p. 105 [Solanum quitoense]; 1941e, p. 
86 [Solanum variabile]. — Drake and Hambleton 1945, p. 366 [Colom- 
bia]. — Hurd 1945, pp. 82, 84, pi. 1, fig. 5 [Martinique; tobacco; 
cabbage]. — Silva 1956, p. 21, fig. 5 [Solanum paniculatum; Solanum 

681-552—65 10 



138 U.S. NATIONAL MUSEUM BULLETIN 243 

fndverulentum]. — Drake and Cobben 1060, p. 89 [Aruba; Curacao; 
St. Eustatius; Saba; St. Martin; Solarium racemonsum; Solarium argil- 
licolum; Solarium nigrum americanum]. 
Corylhaica passijlorae: Monte 1942d, p. 110; 1943c, pp. 113-120, fig. 
[Cuba; Solarium hirtum; Solarium grandijlorurn]; 1945b, p. 36, fig. 
[Solarium tabacijolium]; 1947a, p. 233, fig. — Bruncr, Scaramuzza 
and Otero 1945, p. 168. 
Type: Presumably lost. 

Distribution: Windward Islands (Martinique; St. Vincent; Grenada); 
Cuba; Puerto Rico; Virgin Islands (St. Thomas); Netherlands Antilles 
(Aruba; Curasao); Leeward Islands (Saba; St. Martin; St. Eustatius); 
Panama (Canal Zone); Colombia; Venezuela; Brazil; Peru; Argentina. 

Host Plants: Lycopersicum esculentum; Passijlora caerulea; Ricinus communis; 
Solanum argillicolum; Solanum balbisii; Solanum bonariense; Solanum elaeagni- 
folium; Solanum gilo; Solanum grandijlorum; Solanum hirtum; Solanum juripeba; 
Solanum lycopersicum; Solanum mclongena; Solanum nigrum v. americanum; 
Solanum paniculatum; Solanum pulverulentum; Solanum quitoense; Solanum racemi- 
florum; Solanum racemosum; Solanum sisymbrijolium; Solanum tabacijolium; 
Solanum torvum; Solanum tuberosum; Solanum variabile; cabbage; tobacco. 

Note: Life history studies [Cotton; Monte 1943c, 1945b]; predators and 
parasites [Cotton; Soares]. 

*Corythaica cytharina (Butler) 

Monanthia cytharina Butler 1877, p. 90 [James Island]. 
Leptostyla cytharina: Distant 1902b, p. 354.— Champion 1924, p. 260. 
Corylhaica renormata Barber 1925, p. 251 [Daphne Major]. 
Corylhaica cytharina: Barber 1934, p. 286.— Hurd 1945, pp. 82, 86, 
figs. 2a, b. 
Type: Sex unknown; James Island, Galapagos Islands; British Mus. 
Distribution: Galapagos Islands (James; Daphne Major). 
Host Plant: Unrecorded. 

*Corythaica monacha (Stal) 

Tingis monacha Stal 1858, p. 64 [Brazil]. 

Corylhaica monacha: Stal 1873, p. 128 (in part). — Gibson 1919c, p. 99 
(in part).— Drake and Bruner 1924a, p. 151.— Drake 1926b, p. 378; 
1936b, p. 700 [Argentina].— Costa Lima 1930, p. 84; 1936, p. 126. — 
Drake and Hambleton 1934, p. 450 [Sida glomerata]. — Drake and 
Poor 1937d, p. 311 [Chile].— Monte 1937a, p. 30, fig. 8 [Sida cordi- 
Jolia; Gossypium); 1943c, p. 115 [Sida rhombijolia; Richardia brasilien- 
sis}.— Soares 1941, p. 263, figs. 1, 3, 4.— Hurd 1945, p. 82, pi. 1, 
fig. 6 [Venezuela].— Blote 1945, p. 85.— Silva 1956, p. 24, fig. 6 
[Sida ulmi/olia]. 

Corylhaica monancha [sic]: Drake 1930a, p. 1; 1935, p. 20 [Paraguay]. 
Type: Sex unknown; Brazil; Stockholm Mus. 



LACEBUGS OF THE WORLD 139 

Distribution: Brazil; Paraguay; Venezuela; Argentina; Chile; Peru. 
Host Plants: Gossypium sp. ; Richardia brasiliensis; Sida cordifolia; Sida 
glomerata; Sida rhombifolia; Sida ulmifolia; tomato. 
Note: Parasites [Soares]. 

*Corythaica smithi Drake 

Corythaica smithi Drake 1921b, p. 50, pi. 1, figs, a, a' [Colombia]. — Hurd 
1945, pp. 81, 95, pi. 1, figs. 7, 7a. 
Type: Sex unknown; Bonda, Colombia; Carnegie Mus. 
Distribution: Colombia. 
Host Plant: Unrecorded. 

*Corythaica umbrosa (Monte) 

Leptotingis umbrosa Monte 1938b, p. 128 [Brazil; Richardia brasiliensis]; 

1939b, p. 76 [Diodia conferta]; 1940c, p. 193. 
Corythaica umbrosa: Monte 1942d, p. 105, fig. 1; 1943c, p. 115. — Drake 
and Hambleton 1942a, p. 330.— Hurd 1945, pp. 81, 95, fig. 4 
[Paraguay] . 
Type: Holotype d* ; Belo Horizon te, Minas Gerais, Brazil; Monte Coll. 
(Mus. Nacional). 
Distribution: Brazil; Paraguay. 
Host Plants: Diodia conferta; Richardia brasiliensis. 
*Corythaica venusta (Champion) 

Dolichocysta venusta Champion 1898b, p. 57, pi. 2, figs. 1,1a [Mexico]. — 
Heidemann 1899a, p. 339 [Mont.; Tex.].— Drake 1917a, p. 214 
[Colo.].— Gibson 1919c, p. 102 [Calif.].— Monte 1942d, p. 112. — 
Perkins and Swezey 1924, p. 52 [Lantana]. 
Dolichocysta magna Gibson 1919c, p. 102 [S. Dak.; Nebr.]. 
Dolichocysta densata Gibson 1919c, p. 102.— Monte 1942d, p. 112, fig. 3. 
Dolichocysta obscura Van Duzee 1923, p. 140. 

Corythaica venusta: Hurd 1945, pp. 81, 90, pi. 1, figs. 8, 8a [Ariz.; 
N. Mex.; Kans.; Eriogonum; Salsola pestifer]. 
Type: Sex unknown; Guadelupe, Lower California, Mexico; Vienna 
Mus. 

Distribution: Mexico; U.S. (Ariz.; N. Mex.; Tex.; Calif.; Colo.; 
S. Dak.; Kans.; Nebr.; Mont.); Canada (B.C.). 
Host Plants: Eriogonum sp.; Lantana sp.; Salsola pestifer. 

Genus CORYTHAUMA Drake and Poor 

Corythauma Drake and Poor 1939c, p. 206. — Drake and Runoff 1960a, 
p. 48. 
Type Species: Leptopharsa ayyari Drake. 



140 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Corythauma ayyari (Drake) 

Leptopharsa ayyari Drake 1933, p. 1016 [India; jasmine]. 
Corythauma ayyari: Drake and Poor 1939c, p. 206 [Lantana], 
Type: Holotype 9, macropterous; S. Arcot, Cuddalore, Madras 
Presidency, India: Drake Coll. (USNM). 
Distribution: India: Penang Island. 
1 [ost l'i an is: Jasminum pubescens; Lantana sp.; chamcla leaves. 

♦Corythauma varia Drake and Maa 

Corythauma ayyari var. varia Drake and Maa 1953, p. 95 [India]. 
Corythauma varia: Drake and Runoff 1962a, p. 156. 
Type: Holotype 9, macropterous; Trichinopoly, Madras, India; Drake 
Coll. (USNM). 
Distribution: India. 
Host Plant: Unrecorded. 

Genus CORYTHOTINGIS Drake and Poor 

Corythotingis Drake and Poor 1943, p. 195. — Drake and Runoff 1960a, 
p. 48. 
Type Species: Corythotingis zimmermani Drake and Poor. 

Corythotingis zimmermani Drake and Poor 

Corythotingis zimmermani Drake and Poor 1943, p. 196, fig. 3 [Fiji]. 
Type: Holotype cf; Mt. Korobamba, Viti Levu, Fiji; Bishop Mus. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Unrecorded. 

Genus CORYTHUCHA Stal 

Corythucha Stal 1873, pp. 119, 122.— Champion 1897, p. 6 (key).— 
Osborn and Drake 1916a, p. 224 (key).— Van Duzee 1916, p. 25; 
1917b, p. 212— Gibson 1918, pp. 64-104 (key).— Parshley 1919b, 
p. 70; 1923b, p. 699.— Britton 1920, p. 77.— Barber 1922a, p. 16; 
1922b, p. 19.— Drake 1922d, p. 64; 1928b, p. 100— McAtee 1923, 
p. 147.— Downes 1925, p. 14; 1927a, p. 10.— Torre-Bueno 1929, p. 
311.— Knowlton 1933, p. 262.— Monte 1941e, p. 88.— Froeschner 
1944, pp. 646, 667.— Hurd 1946, p. 482.— Bailey 1951, p. 62.— 
Ash 1954, p. 185.— Gibson and Carrello 1959, p. 19.— Drake and 
Ruhoff 1960a, p. 48. 

Corythuca [sic]: Uhler 1886, p. 22 — Lethierry and Severin 1896, p. 10 — 
Torre-Bueno 1908, p. 232; 1910, p. 30.— Banks 1910, p. 55.— Smith 
1910, p. 148.— Blatchley 1926, p. 453 (key).— Blote 1945, p. 82. 
Type Species: Tingis fuscigera Stal. 



LACEBUGS OF THE WORLD 141 

*Corythucha abdita Drake 

Corythucha abdita Drake 1948j, p. 435 [Guatemala]. 
Type: Holotype o 71 , macropterous; S. Geronimo, Guatemala; Drake 
Coll. (USNM). 
Distribution: Guatemala. 
Host Plant: Unrecorded. 

*Corythucha acculta Drake and Poor 

Corythucha acculta Drake and Poor 1942, p. 302 [Argentina]. 
Type: Holotype cf ; Tucuman, Argentina; Argentina Mus. 
Distribution: Argentina. 
Host Plant: Unrecorded. 

*Corythucha aesculi Osborn and Drake 

Corythucha aesculi Osborn and Drake 1916a, p. 232, pi. 8, fig. g [Aesculus 
glabra; Ohio]; 1917b, p. 304 [III.; Ky.].— Gibson 1918, pp. 71, 76, 
78 [Aesculus hippocastanum].— -Drake 1928b, p. 101 [N.Y.].— Stehr 
1938, p. 13, figs. 1-3, tables 1-10 [Aesculus octandra]. — Froeschner 
1944, p. 668 [Mo.]. 
Corythucha fuscigera (not Stal): Gibson 1918, p. 78 (in part). 
Corythuca aesculi: Blatchley 1926, p. 465 [Ind.; Fla.]. 
Type: Holotype 9, macropterous; Columbus, Ohio, U.S.; Drake Coll. 
(USNM). 

Distribution: U.S. (Ohio; Ind.; 111.; Iowa; Mo.; Ky.; Ark.; W. Va.; 
Va.; D.C.; Md.; N.C.; Fla.; N.Y.; Conn.). 
Host Plants: Aesculus glabra; Aesculus hippocastanum; Aesculus octandra. 
Note: Life history study [Stehr]; predators [Stehr]. 

*Corythucha agalma Drake and Cobben 

Corythucha decens (not Stal): Champion 1897, p. 7 (in part). 
Corythucha agalma Drake and Cobben 1960, pp. 68, 96, fig. 88b [Saba; 
Guatemala; Vernonia]. 
Type: Holotype 9, macropterous; Hellsgate, Saba; Drake Coll. 
(USNM). 

Distribution: Leeward Islands (Saba); Guatemala. 
Host Plant: Vernonia sp. 
*Corythucha arcuata (Say) 

Tingis arcuata Say 1832, p. 27.— Fitch 1858, p. 794.— Le Conte 1859, 

p. 350.— Walker 1873a, p. 180.— Uhler 1878a, p. 415 (in part). 
Corythucha arcuata: Stal 1873, p. 123 [Tex.; 111.]. — Osborn and Drake 
1916a, p. 227, pi. 9 [Ohio]; 1917b, p. 297 [Wis.; Iowa; Quercus 
macrocarpa]. — Parshley 1917b, p. 54 [N.H.; Mass.; R.I.; Conn.]; 
1922b, p. 11 [S. Dak.].— Van Duzee 1917b, p. 212 [Que.; Ont; 
N.J.; D.C.; Ala.; N.C.].— Gibson 1918, pp. 72, 75, 101.— Barber 
and Weiss 1922, p. 5, fig. 1.— Hussey 1922a, p. 11 [N. Dak.]; 1922b, 
p. 23 [Quercus muehlenbergii; Mich.]. — Knowlton 1933, p. 262 



142 U.S. NATIONAL MUSEUM BULLETIN 243 

[Utah].— Froeschner 1944, p. 667 [Mo.].— Bailey 1951, p. 63 [Va.; 
Vt.; Quercus prinoides].— Feldman and Bailey 1952, pp. 98, 99, 102, 
103, fig. 6. 
Corythuca arcuata: Uhler 1884, p. 284, fig. 327 [Canada; oaks].— 
Summers 1891, p. 90, fig. 11 (fig. only).— Gillette and Baker 1895, 
p. 57 [Colo.]. — Morrill 1903, p. 127, pi. 3 [Quercus alba; Quercus 
acuminata; Qjtercus prinus; Quercus rubra]. — Parshley 1914, p. 145 
[Maine].— Blatchley 1926, p. 464, fig. 103 [Ind.]. 
Corythuca ciliata (not Say): Provancher 1886, p. 158. — Van Duzee 

1912, p. 323. 
Corythuca juglandis (not Fitch): Provancher 1886, p. 158. — Van Duzee 

1912, p. 323. 
Corythucha polygrapha Lintner 1888, p. 109 [N.Y.; Md.; Castanea 

americana] . 
Corythuca arquata [sic]: Van Duzee 1889, p. 5; 1894, p. 181. 
Corythuca polygraphia [sic]: Banks 1910, p. 56. 
Corythucha mali Gibson 1918, pp. 71, 77, 98. 
Corythucha pier cei Gibson 1918, pp. 72, 77, 85 [Ariz.; Quercus]. 
Corythucha arcuata var. mali: Drake 1921b, p. 54 [apple; maple]. — 

Parshley 1922b, p. 11. 
Corythuca arcuata var. mali: Blatchley 1926, p. 465 [rose]. 
Type: ApparenUy lost. 

Distribution: U.S. (Maine; N.H.; Mass.; Vt.; R.I.; Conn.; N.Y.; N.J.; 
Pa.; Md.; D.C.; Va.; N.C.; S.C.; Ga.; Ala.; Miss.; Ohio; 111.; Ind.; Mo.; 
Mich.; Wis.; Nebr.; Minn.; Iowa; N. Dak.; S. Dak.; Colo.; Utah; Ariz.; 
N. Mex.; Tex.); Canada (Que.; Ont.). 

Host Plants: Primary host plants — Quercus acuminata; Quercus alba; 
Quercus macrocarpa; Quercus muehlenbergii; Quercus prinoides; Quercus prinus; 
Quercus rubra; Quercus sp.; Castanea americana; occasionally found feeding and 
breeding on Pyrus malus; maple; wild rose. 

Note: Life history studies [Morrill; Bailey]; nymphs [Barber and Weiss]; 
ovipositors [Feldman and Bailey]. 

Corythucha argentinensis Monte 

Corythucha argentinensis Monte 1940e, p. 297 [Argentina]. 
Type: Holotype cf; Resistencia, Argentina; Monte Coll. (Mus. 
Nacional). 

Distribution: Argentina. 
Host Plant: Unrecorded. 
♦Corythucha associata Osborn and Drake 

Corythucha associata Osborn and Drake 1916b, p. 14 [Tcnn.; Primus 
serotina]; 1917b, p. 304 [Ga.; N.Y.; D.C.].— Gibson 1918, pp. 71, 74, 
79 [peach].— Drake 1921b, p. 54 [Ohio; Md.]; 1925c, p. 36 [Miss.].— 



LACEBUGS OF THE WORLD 143 

Barber and Weiss 1922, p. 9.— McAtee 1923, p. 147 [Va.].— Bailey 
1951, p. 66 [Conn.]. 
Corythucha spinulosa Gibson 1918, pp. 71, 75, 79 [N.J.]. — Dickerson and 

Weiss 1918, p. 121, pi. 7. 
Corythucha fusciger a (not Stal): Gibson 1918, p. 78 (in part). 
Corythuca associata: Blatchley 1926, p. 456 [Ind.; Pa.]. 
Type: Holotype cf, macropterous; Clarksville, Tennessee, U.S.; Drake 
Coll. (USNM). 

Distribution: U.S. (R.I.; Conn.; N.Y.; N.J.; Md.; D.C.; Va.; Ga.; 
Ala.; S.C.; Miss.; Tenn.; Ohio; Pa.; Ind.; 111.; Iowa). 
Host Plants: Prunus serotina; Prunus per ska. 

Note: Life history studies [Bailey; Dickerson and Weiss; Barber and 
Weiss]. 
*Corythucha baccharidis Drake 

Corythucha baccharidis Drake 1922b, p. 37 [Fla.; Baccharis]. 
Corythuca baccharidis: Blatchley 1926, p. 463 [Baccharis halimifolia]. 
Type: Holotype d\ macropterous; Paradise Keys, Florida, U.S.; USNM. 
Distribution: U.S. (Fla.; Miss.; Tex.). 
Host Plants: Baccharis halimifolia; Baccharis sp. 

*Corythucha bellula Gibson 

Corythucha bellula Gibson 1918, pp. 72, 75, 93 [Ohio; Crataegus]. — Well- 
house 1919, p. 441, figs. 24, 25 [N.Y.; Crataegus neofluvialis; Crataegus 
albicans; Crataegus punctata]. — Drake 1922d, p. 64 [alder]; 1928b, 
p. 101 [Crataegus pruinosa; Alnus incana; Ribes oxyacanthoides]. — Bailey 
1951, p. 67 [Vt.; Maine; Corylus]; 1959, p. 65 [N.H.]. 
Corythuca bellula: Blatchley 1926, p. 458. 
Type: Holotype 9, macropterous; Tiffin, Ohio, U.S.; USNM. 
Distribution: U.S. (Maine; N.H.; Vt.; N.Y.; Ohio). 
Host Plants: Alnus incana; Corylus sp.; Crataegus albicans; Crataegus 
neofluvialis; Crataegus pruinosa; Crataegus punctata; Crataegus sp.; Ribes oxyacan- 
thoides. 

Note: Life history study [Wellhouse]. 
Corythucha boliviana Monte 

Corythucha boliviana Monte 1946c, p. 27 [Bolivia]. 
Type: Holotype o 71 ; La Paz, Bolivia; Monte Coll. (Mus. Nacional). 
Distribution: Bolivia. 
Host Plant: Unrecorded. 

*Corythucha brunnea Gibson 

Corythucha brunnea Gibson 1918, pp. 72, 75, 93 [La.; Tex.; Crataegus]. 
Type: Holotype 9, macropterous; Alexandria, Louisiana, U.S.; USNM. 
Distribution: U.S. (La.; Tex.; Miss.; 111.). 
Host Plant: Crataegus sp. 



144 U.S. NATIONAL MUSEUM BULLETIN 24 3 

♦Corythucha bulbosa Osborn and Drake 

Corythucha bulbosa Osborn and Drake 1916a, p. 232, pi. 8, fig. e [Ohio; 
Md.]; 1917b, ]>. 304 [Staphylec tnjolia].— McAtce 1917b, p. 79; 1923a, 
p. 147 [D.C.].— Gibson 1918, pp. 71, 74, 77 [Va.].— Barber and \\ 
1922, p. 9.— Barber 1922a, p. 16 [X.J.]. 
Corythuca bulbosa: Weiss 1919a, p. 17 [Pa.].— Blatchley 1926, p. 455 
find.]. 
Type: Holotypc 9, macropterous; Sugar Grove, Ohio, U.S.; Drake Coll. 
(USNM). 
Distribution: U.S. (Ohio; Mo.; Ind.; 111.; Pa.; N.J.; Md.; D.C.; Va.). 
Host Plant: Staphylea trifolia. 
Note: Life history study [Weiss]. 
*Corythucha caelata Uhler 

Corythuca caelata Uhler 1894b, p. 279 [Mexico].— Banks 1910, p. 55 

[Calif.]. 
Corythucha caelata: Gibson 1918, pp. 71, 77, 101 (in part). 
Corythucha compta Drake and Hambleton 1944a, p. 96. 
I ype: Lcctotype o", macropterous; "S. Cal."; Cal. Acad. 
Distribution: Mexico (San Esteban, Baja California); U.S. (Calif.). 
Host Plant: Taken on pods of Yucca arbor escens. 

♦Corythucha caryae Bailey 

Corythucha caryae Bailey 1951, p. 68, fig. 3 [Mass.; N.H.; Carya ovata]; 
1959, p. 65 [R.I.; Carya].— Feldman and Bailey 1952, pp. 98, 100, 102, 
103, fig. 12. 
Type: Holotypc o 71 ; Newbury, Massachusetts, U.S.; MCZ. 
Distribution: U.S. (Mass.; N.H.; R.I.). 
Host Plants: Carya ovata; Carya sp. 

Note: Parasites [Bailey 1951]; ovipositors [Feldman and Bailey]. 
*Corythucha celtidis Osborn and Drake 

Corythucha celtidis Osborn and Drake 1916a, p. 227 [Ohio; hackberry]. — 
Gibson 1918, pp. 72, 77, 84.— Drake 1919a, p. 418 [Ky.; Tenn.; 
S.C.]; 1925c, p. 37 [Celtis occidentalis].— Weiss 1921, p. 104 [N.J.].— 
Barber and Weiss 1922, p. 8.— McAtce 1923, p. 148 [Md.]. 
Corythuca celtidis: Blatchley 1926, p. 468 [Ind.]. 
Type: Sex unknown; Columbus, Ohio, U.S.; Ohio State Univ. 
Distribution: U.S. (Ohio; Iowa; 111.; Ind.; Ky.; Tenn.; Ga.; N.C.; 
S.C.;Md.;NJ.). 
Host Plant: Celtis occidentalis. 
No'ii : Life history study [Weiss]. 
*Corythucha celtidis var. mississippiensis Drake 

Corythucha celtidis var. mississippiensis Drake 1925c, p. 36 [Miss.; Ga.; 

S . C . ; Te n n . ; Celtis m is sis si pp i ens is] . 
Corythuca celtidis var. mississippiensis: Blatchley 1926, p. 468. 



LACEBUGS OF THE WORLD 145 

Type: Holotype o", macropterous; Columbus, Mississippi, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Miss.; Ga.; Term.; S.C.). 
Host Plant: Celtis mississippiensis. 

*Corythucha cerasi Drake 

Corythucha cerasi~Drake 1948j, p. 435 [Wash.; Ind.; cherry; oak]. 

Type: Holotype cf, macropterous; Wenatchee, Washington, U.S.; 
Drake Coll. (USNM). 

Distribution: U.S. (Wash.; Ind.). 

Host Plants: Cherry; oak. 

*Corythucha championi Drake and Cobben Frontispiece 

Corythucha decens (not Stal): Champion 1897, p. 7, pi. 1, figs. 7, 7a (in 
part) [Guatemala]. — Gibson 1918, pp. 74, 96 [Panama]. — Drake 
1926b, p. 379.— Drake and Hambleton 1945, p. 366; 1946b, p. 123. 
Corythucha championi Drake and Cobben 1960, pp. 68, 93, figs. 88a, 89 
[Curacao; Abutilon umbellatum; Ayenia magna]. 
Type: Holotype o 71 , macropterous; Groot St. Jeris, Curasao, Nether- 
lands Antilles; Drake Coll. (USNM). 

Distribution: Netherlands Antilles (Curasao); Mexico; Guatemala; 
Colombia; El Salvador; U.S. (Pa.). 
Host Plants: Abutilon umbellatum; Ayenia magna. 

*Corythucha ciliata (Say) 

Tingis ciliata Say 1832, p. 27.— Fitch 1858, p. 793.— Le Conte 1859, p. 
348.— Walker 1873a, p. 180.— Uhler 1878, p. 414 [Ala.; Md.; 
sycamore]. 

Tingis hyalina Herrich-Schaeffer 1840, p. 84, pi. 173, fig. 532 [Carolina]; 
1850, p. 161.— Fieber 1844, p. 103, pi. 9, figs. 1-4.— Walker 1873a, 
p. 180. 

Corythucha ciliata: Stal 1873, p. 123 [111.; N.J.]. — Osborn and Drake 
1916a, p. 225, pi. 8, fig. h [Ohio]; 1917b, p. 297 [Iowa; Tex.].— 
Parshley 1917b, p. 53 [Conn.; R.I.]; 1923b, p. 702.— Van Duzee 
1917b, p. 212 [Que.; Ont.; D.C.].— Wade 1917, p. 3, figs. 3-7 
[Okla.].— Gibson 1918, pp. 75, 102.— Drake 1919a, p. 418 [Mo.; 
Fraxinus; Carya ovata; Broussonetia papyri/era]. — Kotinsky 1921, p. 74, 
fig. 53.— Barber and Weiss 1922, p. 4.— Hussey 1922b, p. 23 [Mich.].— 
Felt 1933, p. 49.— Froeschner 1944, p. 667.— Bailey 1951, p. 72 
[Vt.; Chamaedaphne}.— Feldman and Bailey 1952, pp. 98, 100, 102, 
103, fig. 13. 

Corythucha hyalina: Stal 1873, p. 123. 

Corythuca ciliata: Lintner 1888, p. 107 [N.Y.; Platanus occidentalis]. — 
Cockerell 1893, p. 364 [Colo.].— Van Duzee 1894, p. 181.— Gillette 
and Baker 1895, p. 57.— Morrill 1903, p. 133 [Mass.].— Wirtner 
1904, p. 202 [Pa.].— Smith 1910, p. 148.— Weiss 1913, p. 407 — 



146 U.S. NATIONAL MUSEUM BULLETIN 243 

Parshley 1914, p. 145 [Maine].— Blatchley 1926, p. 458 [Ind.].— 
MOte 1945, p. 82 [Va.]. 

Corythuca ciliati [sic]: Summers 1891, p. 90 [Term.]. 

Type: Unknown. 

Distribution: U.S. (Colo.; Okla.; Tex.; Iowa; Mo.; Mich.; 111.; Ohio; 
Ind.; Pa.; N.Y.; N.J.; Md.; D.C.; Va.; N.C.; Ala.; Ga.; Fla.; Tenn.; 
Mass.; R.I.; Conn.: Yt.; Maine): Canada (Ont.; Que.). 

Host Plants: Primary host Platanus occidentalis; also found on Brous- 
sonetia papyri/era; Carya ovata; Chamaedaphne sp.; Fraxinus sp. 

Note: Life history studies [Morrill; Wade; Bailey]; ovipositor [Feldman 
and Bailey]. 

*Corythucha clara Drake and Hambleton 

Corythucha clara Drake and Hambleton 1938a, p. 66 [Brazil]. — Monte 
1941e, p. 88 [Urticaceae]. 
Type: Holotype d\ macropterous; Rio Claro, Sao Paulo, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Urticaceae. 
♦Corythucha confraterna Gibson 

Corythucha confraterna Gibson 1918, pp. 72, 75, 102 [Calif.; Mexico; 
Platanus]. 
Type: Holotype 9, macropterous; Los Angeles, California, U.S.; 
USNM. 
Distribution : U.S. (Calif.; Ariz.); Mexico. 
Host Plant: Platanus sp. 

*Corythucha coryli Osborn and Drake 

Corythucha coryli Osborn and Drake 1917b, p. 299 [Md.; Corylus ameri- 
cana].— Gibson 1918, pp. 72, 74, 92.— McAtee 1923, p. 148 [Va.].— 
Bailey 1951, p. 73 [Mass.; Ostrya virginiana]; 1959, p. 66 [Conn.; 
N.H.; R.I.; Corylus].— Feldman and Bailey 1952, pp. 98, 100, 102, 
fig. 15. 
Corythuca coryli: Blatchley 1926, p. 455 [N.J.; Ind.]. 
Type: Holotype 9, macropterous; Plummer's Island, Maryland, U.S.; 
Drake Coll. (USNM). 

Distribution: U.S. (Md.; D.C.; Va.; Iowa; 111.; Kans.; Nebr.; Ind.; 
N.J.; Mass.; Conn.; R.I.; N.H.). 

Host Plants: Corylus americana; Corylus sp.; Ostrya virginiana. 
♦Corythucha cydoniae (Fitch) 

Tingis cydoniae Fitch 1861, p. 114, fig. [Mass.; quince]. 
Corythuca arcuata (not Say): Comstock 1880, p. 221 [D.C.; Crataegus 
cor data; Crataegus coccinea; Crataegus tomentosa; Crataegus crusgalli; 
Crataegus parvijolia]. — Lintner 1888, p. 108, figs. 42, 43. — Wirtner 
1904, p. 202 [Pa.]. 



LACEBUGS OF THE WORLD 147 

Corythuca arcuata var. crataegi Morrill 1903, p. 132. 

Corythuca cydoniae: Banks 1910, p. 56 [N.Y.].— Blatchley 1926, p. 455, 

fig. 99 [Ind.; Ont.]. 
Corythucha crataegi: Osborn and Drake 1916a, p. 229, pi. 8, fig. f [Ohio]; 
1917b, p. 300 [Tex.; Colo.; Md.; Va.; S.C.; Ga.; S. Dak.; Iowa; 
Nebr.; Wis.].— Parshley 1917b, p. 54 [Conn.]; 1917c, p. 47.— McAtee 
1917b, p. 78 [Amelanchier ; Cephalanthus]. 
Corythucha cydoniae: Gibson 1918, pp. 72, 75, 87 [Canada; Amelanchier 
intermedia; Cydonia vulgaris]. — Barber 1922a, p. 16 [N.J.]. — Hussey 
1922a, p. 11 [N. Dak.; Mich.; Amelanchier canadensis]. — McAtee 1923, 
p. 148 [Cephalanthus occidentalis]. — Torre-Bueno 1926a, p. 54; 1933, 
p. 228 [Quercus; Ark.].— Froeschner 1944, p. 667 [Mo.].— Sailer 
1945, p. 81. — Bailey 1951, p. 74, graphs 1, 2 [Chaenomeles; Pyrus 
americana; Pyrus melanocarpa; Pyrus aucuparia; Pyracantha coccinea; 
Mains; loquat; pear]; 1959, p. 66 [R.I.; Cotoneaster hupehensis]. — Feld- 
man and Bailey 1952, pp. 96, 98, 100, 102, 103, figs. 3, 7.— Ash 
1954, p. 185. 
Corythucha occidentalis Drake (in Gibson 1918, pp. 74, 91) [Calif.]. 
Type: Leominster, Massachusetts, U.S.; sex and deposition of type 
unknown. 

Distribution: U.S. (Maine; R.I.; Mass.; Conn.; Vt.; N.Y.; N.J.; Pa. 
Md.; D.C.; Va.; S.C.; Ala.; Ga.; Ark.; Tex.; Nebr.; Mich.; Wis.; Ohio 
Ind.; Mo.; Idaho; Iowa; Colo.; S. Dak.; N. Dak.; Utah; Wash.; Calif.) 
Canada (Ont.); Mexico. 

Host Plants: Amelanchier canadensis; Amelanchier intermedia; Amelanchier 
sp.; Cephalanthus occidentalis; Cephalanthus sp.; Chaenomeles sp.; Cotoneaster 
hupehensis; Crataegus coccinea; Crataegus cordata; Crataegus crusgalli; Crataegus 
parvifolia; Crataegus tomentosa; Crataegus sp.; Cydonia vulgaris; Pyracantha 
coccinea; Pyrus americana; Pyrus aucuparia; Pyrus melanocarpa; Malus sp.; 
Quercus sp.; loquat; pear. 
Note: Life history studies [Comstock; Bailey]; bite to man [Sailer]. 

:i: Corythucha decepta Drake 

Corythucha decepta Drake 1932, p. 100 [Mexico]; 1938a, p. 70 [Alnus 

acuminata]. — Monte 1947a, p. 237. 
Corythucha deceptiva [sic]: Drake and Hambleton 1945, p. 366 [Guate- 
mala]. 
Type: Holotype o* ; San Miguel, Hidalgo, Mexico; MCZ. 
Distribution: Mexico; Guatemala. 
Host Plant: Alnus acuminata. 

*Corythucha distincta Osborn and Drake 

Corythucha sp. Cockerell 1893, p. 364 [Colo.; Cnicus]. 
Corythuca fuscigera (not Stal): Gillette and Baker 1895, p. 57. 
Corythucha distincta Osborn and Drake 1916b, p. 13; 1917b, p. 301 



148 U.S. NATIONAL MI/SLUM BULLETIN 243 

[Wash.; Munt.: Wyo.J Utah; Calif.; Carduus lanceolatus]. — Gibson 1918, 

pp. 72,75,81.— Parshley 1919a, p. 18 [B.C.; hollyhock]; 1922b, p. 11 

[S. Dak.].- Downes 1925, p. 15, fig. 5; 1927a, p. 10 [Lathyrus nuttallu}. 
Type: Sex unknown; Colorado, U.S.; Ohio State Univ. 
Distribution: U.S. (Calif.; Wash.; S. Dak.; \\ >...; Mont.; Utah; Colo.); 
Canada (B.C.). 

Host Plants: Carduus lanceolatus; Cnicus sp.; Lathyrus nuttallii; hollyhock. 

*Corythucha distincta var. spinata Osborn and Drake 

Corythucha distincta var. spinata Osborn and Drake 1917b, p. 301, figs. 
1 a, b [Mont.; Carduus]. 
Type: Holotype 9, macropterous; Florence, Montana, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Mont.). 
Host Plant: Carduus sp. 
Note: This may not be a good variety. 

*Corythucha elegans Drake 

Corythucha elegans Drake {in Gibson 1918, pp. 73, 75, 89) [Colo.; N.Y.; 
Salix]; 1919b, p. 159 [Ont.; Mich.; Populus balsamifera]; 1922d, p. 65, 
figs. 25 a, b, 26 [Populus tremuloides; Populus grandidentata]. — Torre- 
Bueno 1929, p. 311 [Mass.].— Hurd 1946, p. 483 [Wis.]. 
Corylhuca elegans: Blatchley 1926, p. 456, fig. 102a. 
Type: Holotype 9, macropterous; Poudre Canyon, Colorado, U.S.; 
Drake Coll. (USNM). 

Distribution: U.S. (Colo.; Wis.; Mich.; N.Y.; Mass.); Canada (Ont.; 
B.C.; Sask.; Man.). 

Host Plants: Primary host Salix sp.; also breeds to some extent on 
Populus balsamifera; Populus grandidentata; Populus tremuloides. 
Note: Eggs and instars [Drake 1919b]; predators [Drake 1919b]. 

*Corythucha eriodictyonae Osborn and Drake 

Corythucha eriodictyonae Osborn and Drake 1917b, p. 302 [Calif.; Erio- 
dictyon californicum]. — Van Duzce 1917a, p. 258. — Gibson 1918, 
pp. 77, 99. 
Corythucha erydictyonae [sic]: Drake and Davis 1960, fig. 71. 
Type: Holotype o", macropterous; San Francisquito Cr., San Mateo Co., 
California, U.S.; Drake Coll. (USNM). 
Distribution: U.S. (Calif.). 
Host Plant: Eriodictyon californicum. 

*Corythucha floridana Heidemann 

Corylhuca floridana Heidemann 1909, p. 236, fig. [Fla.J Cephalanthus]. — 

Van Duzee 1909, p. 173.— Blatchley 1926, p. 459, fig. 101. 
Corythucha floridana: Osborn and Drake 1917b, p. 300 [oak].— Gibson 
1918, pp. 76, 101. 
Type: Holotype d\ macropterous; Biscaync Bay, Florida, U.S.; USNM. 



LACEBUGS OF THE WORLD 149 

Distribution: U.S. (Fla). 

Host Plants: Cephalanthus occidentalis; Cephalanthus sp.; oak. 

*Corythucha fuscigera (Stal) 

Tingis fuscigera Stal 1862, p. 323 [Mexico]. 
Monanthia lucida Walker 1873a, p. 191. 

Corythucha fuscigera: Stal 1873, p. 122. — Champion 1897, p. 7, pi. 1, 
figs. 6, 6a [Calif.; Guatemala]; 1898b, p. 57.— Van Duzee 1914, 
p. 11 [chaparral]; 1917a, p. 258 [Hosackia crassifolia]. — Osborn and 
Drake 1917b, p. 303.— Gibson 1918, p. 78 (in part).— Drake 1919a, 
p. 419 [Ariz.]; 1926b, p. 378. 
Leptostyla ? lucida: Uhler 1886, p. 22. 

Corythuca fuscigera: Uhler 1894b, p. 278.— Distant 1902b, p. 357. 
Type: Holotype 9; Mexico; Stockholm Mus. 
Distribution: Mexico; Guatemala; U.S. (Calif.; Ariz.). 
Host Plants: Hosackia crassifolia; chaparral. 

Note: Most records of C. fuscigera in the United States are referable 
to other species. 

*Corythucha fuscomaculata (Stal) 

Tingis fusco-maculata Stal 1858, p. 63 [Brazil]. 

Corythucha fuscomaculata: Stal 1873, p. 123. — Distant 1888, p. lxxxiii. — 
Champion 1898b, p. 57, pi. 2, fig. 2.— Gibson 1918, pp. 76, 89 — 
Drake and Hambleton 1934, p. 451 [Triumfetta semitriloba]. — Drake 
and Poor 1937d, p. 312 [Peru] .—Monte 1939b, p. 68 [Colombia]; 
1941e, p. 88 [Mexico; Argentina]. — Kormilev 1955a, p. 67 [Para- 
guay; Bolivia]. 
Corythuca fuscomaculata: Costa Lima 1936, p. 126. — Monte 1937a, p. 31 
[Triumfetta rhomboidea]. — Silva 1956, p. 25, fig. 7. 
Type: Sex unknown; Brazil; Stockholm Mus. 

Distribution: Brazil; Peru; Colombia; Bolivia; Argentina; Paraguay; 
Mexico. 

Host Plants: Triumfetta rhomboidea; Triumfetta semitriloba; Solarium sp. 
*Corythucha globigera Breddin 

Corythucha glob iger a Breddin 1901, p. 82 [Ecuador]. — Drake and Hamble- 
ton 1944b, p. 129 [Peru].— Drake and Runoff 1961c, p. 145. 
Corythucha globulif era Monte 1940d, p. 102, fig. 14 [Venezuela; Solanum]. 
Type: Holotype o"; Santa Inez, Ecuador; Berlin Mus. 
Distribution: Ecuador; Peru; Venezuela. 
Host Plant: Solanum sp. 
*Corythucha gossypii (Fabricius) 

Acanthia gossypii Fabricius 1794, p. 78. 

Tingis gossypii: Fabricius 1803, p. 126. — Le Peletier and Serville 1828, 
p. 653. — Burmeister 1835, p. 259 [Mexico]. — Herrich-Schaeffer 



150 U.S. NATIONAL MUSEUM BULLETIN 243 

1840, p. 85, pi. 173, fig. 534 [St. Thomas]; 1850, p. 162.— Fiebcr 
1844, p. 104, pi. 9, fig. 5.— Guerin-Meneville 1857, p. 409 [Cuba; 
cotton]. -Walker 1873a, p. 180. 

Tingis decens Stal 1862, p. 324. 

Galeatus gossypii: Stal 1868, p. 93. 

Corythucha decens: Stal 1873, p. 123. — Champion 1897, p. 7 (in part). — 
Lutz 1929, p. 233 [Pa.]. 

Corythucha gossypii: Stal 1873, p. 123.— Barber 1914, p. 507 [Fla.]; 
1939, p. 368.— Osborn and Drake 1917b, p. 300 [Grenada; Ichthyo- 
methia piscipula].— Gibson 1918, pp. 72, 77, 96.— Drake 1919a, p. 
418; 1926a, p. 87; 1928e, p. 5 [Honduras; soursop]. — Van Duzee 
1923, p. 140 [Coronados Islands; Atamisquea emarginata]. — Wilson 
1923, p. 9 [St. Croix].— Gowdy 1925, p. 19 [congo pea; Hibiscus]; 
1926, p. 35.— Leonard and Mills 1931, pp. 309-321 [Trinidad; 
N. Mex.; Tex.; Venezuela; Antigua; Guatemala; Costa Rica; 
Panama; Haiti; Montserrat; Dominican Republic; St. Vincent; 
Union; orange; grapefruit; Jatropha; Dahlia; Prunus persicus; Citrus 
medica; Cassia emarginata; Annona diver sifolia]. — Fenton 1934, p. 198 
[mango; Cajanus indicus; Solanum torvum]. — Monte 1940d, p. 102 
[Colombia]. — Drake and Hambleton 1945, p. 366 [Salvador; Ecua- 
dor; Nicaragua; eggplant]; 1946b, p. 123. — Bruner, Scaramuzza and 
Otero 1945, pp. 14, 15, 29, 45, 72, 84, 114, 132, 152, 155, 188 [Annona 
squamosa; Arachis hypogaea; Erythrina berteroana; Gossypium; Musa 
paradisiaca; Phaseolus limensis; Roystonea regia; ^anthoxylum martini- 
cense]. — Ramos 1946, p. 25 [Mona Island; Capparis flexuosa]. — Gomez - 
Menor 1956a, p. 112.— Drake and Cobbcn 1960, p. 91, fig. 88d 
[Aruba; Curacao; Bonaire; St. Eustatius; Saba; St. Martin; Ja- 
tropha gossypifolia; Jatropha multifida; Capparis cynophallophora; Hi- 
biscus rosa-sinensis; Hibiscus esculentus; Solanum lycopersicum; Solanum 
melongena; Lagenaria leucantha]. 

Corythuca gossypii: Van Duzee 1907, p. 18 [Jamaica]. — Jones 1915, p. 
4 [yautia; Canavalia ensiformis; Ricinus communis]. — Wolcott 1923, p. 
246 [Puerto Rico; Carica papaya; Annona muricata]. — Blatchley 1926, 
p. 460. 

Type: Holotype 9, macropterous; "Americae mcridionalis, Insulis 
Dom."; Kiel Coll. (Copenhagen Mus.). 

Distribution: Jamaica; Haiti; Dominican Republic; Trinidad; Leeward 
Islands (Antigua; St. Eustatius; Saba; St. Martin; Montserrat); Windward 
Islands (St. Vincent; Grenada); Virgin Islands (St. Croix; St. Thomas; 
St. John); Cuba; Puerto Rico; Netherlands Antilles (Bonaire; Aruba; 
Curacao); Ecuador; Colombia; Venezuela; Panama; Costa Rica; Nica- 
ragua; El Salvador; Honduras; Guatemala; Mexico (Coronados, San 
Esteban Islands); Mona Island; U.S. (Tex.; N. Mex.; Fla.; Pa.). 



LACE BUGS OF THE WORLD 151 

Host Plants: Annona divers if olia; Annona muricata; Annona squamosa; 
Arachis hypogaea; Atamisquea emarginata; Cajanus indicus; Canavalia ensiformis; 
Capparis cynophallophora; Capparis flexuosa; Carica papaya; Cassia emarginata; 
Citrus medica; Dahlia sp.; Erythrina berteroana; Gossypium sp.; Hibiscus escu- 
lentus; Hibiscus rosa-sinensis; Hibiscus sp.; Ichthyomethia piscipula; Jatropha 
gossypifolia; Jatropha multifida; Jatropha sp.; Lagenaria leucantha; Musa para- 
disiaca; Phaseolus limensis; Prunus persicus; Ricinus communis; Roystonea regia; 
Solanum lycopersicum; Solanum melongena; Solanum torvum; ^anthoxylum marti- 
nicense; mango; lemon; grapefruit; orange; yautia; soursop. 

Note: Life history studies [Leonard and Mills]; predators [Leonard 
and Mills]. 

*Corythucha heidemanni Drake 

Corythucha heidemanni Drake (in Gibson 1918, pp. 75, 87) [N.Y.]; 1919b, 
p. 159 [Ont.; birch]; 1928b, p. 101 [Ulmus americana; Ulmus fulva]. — 
Parshley 1920b, p. 142 [Vt.; alder]; 1922a, p. 236; 1923b, p. 701. — 
Proctor 1946, p. 74.— Bailey 1951, p. 82 [Mass.; N.H.; Conn.; 
N.B.; Sorbus; Alnus].— Feldman and Bailey 1952, pp. 97-100, 102, 
103, figs. 16, 18.— Lindberg 1958b, p. 14 [Nfld.; N.S.; Alnus crispa]. 
Corythucha borealis Parshley (in Gibson 1918, pp. 75, 92) [Maine]. 
Corythuca heidemanni: Blatchley 1926, p. 457. 
Type: Holotype 9, macropterous; Cranberry Lake, New York, U.S.; 
Drake Coll. (USNM). 

Distribution: U.S. (N.Y.; Conn.; Mass.; Maine; N.H.; Vt.); Canada 
(N.S.; Nfld.; Ont.; N.B.). 

Host Plants: Alnus crispa; Alnus sp.; Betula lutea; Sorbus sp.; Ulmus 
americana; Ulmus fulva. 

*Corythucha hewitti Drake 

Corythucha hewitti Drake 1919b, p. 159 [Manitoba; Cory lus americana], — 
Parshley 1921, p. 17. — Downes 1925, p. 15, fig. 4 [Cory lus calif ornica]. — 
Hurd 1946, p. 483 [Pa.; Iowa; Colo.].— Bailey 1951, p. 84 [Conn.]. 
Corythucha hesperia Parshley 1919a, p. 23 [B.C.]. 
Type: Sex unknown; Aweme, Manitoba, Canada; Canada, Coll. 
Distribution: Canada (Man.; Ont.; B.C.); U.S. (Minn.; Wis.; Iowa; 
Colo.; Pa.; Conn.). 

Host Plants: Cory lus americana; Cory lus calif ornica. 

*Corythucha hispida Uhler 

Corythuca hispida Uhler 1894b, p. 279 [Mexico].— Banks 1910, p. 56 

[Calif.]. 
Corythucha hispida: Gibson 1918, pp. 77, 104 (in part). — Van Duzee 1923, 
p. 140 [Abutilon]. 
Type: Holotype d\ macropterous; San Esteban, Lower California, 
Mexico; Cal. Acad. 



152 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Distribution: Mexico (Baja ( lalif. I; U.S. (Calif.). 
Hosi l'i \\ i : Abutilon sp. 

(.'orythucha immaculata Osborn and Drake 

Corytkucha immaculata Osborn and Drake 19161), p. 11 [Calif.]; 1917b, 
p. 301 [Oreg.; Colo.].— Drake 1918d, p. 86 [Idaho; Mont.]; 1919a, 
p. 419; 1919b, p. 159 [B.C.].— Gibson 1918, pp. 76, 103.— Parshley 
1921, p. 17.— Downes 1925, p. 16, fig. 7.— Knowlton 1933, p. 263 
[Utah]. 
Corythuca fmra Gibson 1917, p. 258 [Wash.; Balsamorhiza sagittata]. 

thucha pura: Gibson 1918, pp. 72, 77, 103.— Parshley 1919a, p. 24. 
Type: Sex unknown; Alameda, ( ialifornia, U.S.; Ohio State Univ. 
Distribution: U.S. (Calif.; Oreg.; Wash.; Idaho; Mont.; Utah; Colo.); 
Canada (B.C.). 
Host Plant: Balsamorhiza sagittata. 

*Corythucha incurvata Uhler 

Corythuca incurvata Uhler 18941), p. 280 [Mexico; Ariz.]. 

Corythuca arcuata (not Say) : Pembertonl911, p. 339, pis. 12-1 4 [Heteromeles 

arbutifolia; Calif.]. 
Corythucha bullata Van Duzee 1917a, p. 258. 

Corytkucha incurvata: Van Duzee 1917b, p. 213 [Oreg.]. — McAtee 1917b, 
p. 78.— Gibson 1918, pp. 72, 74, 92.— Drake and Ruhoff 1962a, p. 
156. 
Corythucha heteromelecola Drake 1920, p. 50. 
Type: Sex unknown; "Cal. 7"; Cal. Acad. 
Distribution: U.S. (Calif.; Ariz.; Oreg.); Mexico. 
Host Plant: Heteromeles arbutifolia. 
Note: Life history study [Pemberton]. 

*Corythucha juglandis (Fitch) 

Tingisjuglandis Fitch 1857, p. 466, item 193 [N.Y.; basswood]. 
Corythucha juglandis: Stal 1873, p. 123 [111.; N.J.].— Osborn and Drake 

1916a, p. 226 [walnut; Ohio]; 1917b, p. 297 [Ga.; Tenn.; Iowa; 

Wis.]. — Gibson 1918, pp. 71, 75, 80 [Kans.; Tex.; Tilia americana]. — 

Myers 1926, p. 111.— Proctor 1946, p. 74.— Bailey 1951, p. 84 [Vt.; 

N.B.; Carya illinoensis; Rubus; Sorbus]. — Feldman and Bailey 1952, 

pp. 98, 101,102, fig. 10. 
Tingis arcuata (not Say) : Uhler 1 878, p. 41 5 (in part) . 
Corythuca juglandis: Uhler 1884, p. 285 [butternut]. — Van Duzee 

1905, p. 549.— Blatchley 1926, p. 466. 
Corythuca arcuata: Smith 1910, p. 148.— Parshley 1914, p. 145 [Maine]; 

1917b, p. 55 [N.H.; Mass.; R.I. ; Conn.]. 
Corythucha conlracla Osborn and Drake 1916a, p. 230. — Gibson 1918, 

p. 104.— Drake 1921b, p. 51 [Ind.].— Barber and Weiss 1922, p. 8 — 

Proctor 1946, p. 74 [Sorbus americana]. 



LACEBUGS OF THE WORLD 153 

Corythucha parshleyi Gibson 1918, pp. 72, 76, 83 [Carya olivaeformis; 
Amelanchur intermedia]. — Weiss and Dickerson 1918b, p. 401 [Juglans 
cinerea; Juglans sieboldiana; Juglans nigra]. — Drake 1919b, p. 159 
[N.C.]. 
Corythuca contracta: Blatchley 1926, p. 467. — Blote 1945, p. 83 [Md.; 
Va.]. 
Type: Holotype cf, macropterous; New York State, U.S.; USNM. 
Distribution: U.S. (Maine; N.H.; Vt.; Mass.; R.I.; Conn.; N.Y.; N.J.; 
Md.; Va.; N.C; S.C.; Ga.; Tenn.; Ohio; Ind.; 111.; Wis.; Iowa; Kans.; 
Tex.; Miss.); Canada (N.B.). 

Host Plants: Amelanchier intermedia; Carya illinoensis; Carya olivaeformis; 
Juglans cinerea; Juglans nigra; Juglans sieboldiana; Rubus sp.; Sorbus sp.; 
Sorbus americana; Tilia americana. 

Note: Life history studies [Barber and Weiss; Weiss and Dickerson]; 
ovipositors [Feldman and Bailey]. 

*Corythucha lowryi Drake 

Corythucha lowryi Drake 1948j, p. 434 [Wis.; Carpinus caroliniana]. 
Type: Holotype cf, macropterous; Frederic, Wisconsin, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Wis.). 
Host Plant: Carpinus caroliniana. 

*Corythucha marmorata (Uhler) 

Tingis marmorata Uhler 1878, p. 415 [N.C.]. 

Corythuca marmorata: Van Duzee 1889, p. 5 [Canada]; 1894, p. 181 
[N.Y.]; 1907, p. 19 [Jamaica]; 1909, p. 173 [Fla.].— Howard 1904, 
p. 89.— Felt 1904, p. 125 [ragweed].— Smith 1910, p. 148 [N.J.].— 
Osborn and Drake 1915a, p. 506 [Ohio].— Blatchley 1926, p. 462, 
pi. 4, fig. 6.— Blote 1945, p. 83 [Va.]. 

Corythuca irrorata Howard 1898, p. 99 [Ala.; Chrysanthemum]. 

Corythuca gossypii (not Fabricius): Torre-Bueno 1908, p. 232. 

Corythucha marmorata: Van Duzee 1914, p. 11 [Calif.; sunflower]; 1917b, 
p. 214 [Ont.; N. Mex.].— Barber 1914, p. 507.— Osborn and Drake 
1916a, p. 226; 1917b, p. 298 [Nebr.; Ind.; Mich.; Pa.; Maine; Mass.; 
Ga.; Tenn.; La.; Iowa; Colo.].— Parshley 1917b, p. 54 [N.H.; Vt.; 
R.I.; Conn.]; 1917c, p. 47; 1919a, p. 20 [B.C.]; 1922b, p. 11 [S. 
Dak.]; 1923a, p. 103; 1923b, p. 702, pi. 17, fig. 6.— McAtee 1917b, 
p. 78 [D.C.; Aster]; 1923, p. 148 [Solidago; Ambrosa trifida].— Gibson 
1918, pp. 76, 94.— Barber and Weiss 1922, p. 7, fig. 3.— Hussey 
1922a, p. 11 [N. Dak.]; 1922b, p. 23 [goldenrod].— Weiss 1924, p. 367 
[Solidago semper vir ens]. — Weiss and Lott 1924, p. 68. — Downes 1925, 
p. 16, fig. 6.— Torre-Bueno 1925, p. 179.— Drake 1926b, p. 379 — 
Knowlton 1933, p. 263.— Abbott 1935, p. 13.— Froeschner 1944, 
p. 667 [Mo.; Helianthus].— Proctor 1946, p. 74.— Bailey 1951, p. 86 

681-552—65 11 



154 U.S. .NATIONAL MUSEUM BULLETIN 213 

[ Tanacetum] ; 1959, p. 67 [Rudbeckia serotina; Echinops sphaerocephalus]. — 
Fcldman and Bailey 1952, pp. 96, 98, 101-103, figs. 2, 8. 
Corythucha decens (not Stal): Van Duzee 1917b, p. 214 (in part). 
Corythucha lactea Drake (in Gibson 1918, pp. 74, 94) [Utah]. 
Corythucha marmorata var. informis Parshlcy 1919a, p. 20 [N.S.]; 1922b, 

p. 11.— Downcs 1925, p. 16.— Bailey 1951, p. 86. 
Corythucha marmorala var. minutissima Drake 1920, p. 50. 
1 w\.: North Carolina, U.S.; sex and deposition of type unknown. 
Distribution: U.S. (Maine; N.H.; Vt.; Mass.; R.I.; Conn.; N.Y.; N.J. 
Pa.; Md.; D.C.; Va.; N.C.; Ga.; Miss.; Fla.; Ala.; La.; Tenn.; W. Va. 
Ohio; Ind.; 111.; Iowa; Kans.; Mo.; Mich.; Wis.; Nebr.; N. Dak.; S. Dak. 
Colo.; Utah; Idaho; Oreg.; N. Mex.; Ariz.; Tex.; Calif.); Canada (Ont. 
B.C.; N.S.) ; Jamaica; Mexico. 

Host Plants: Ambrosia triftda; Aster sp.; Chrysanthemum sp.; Echinops 
sphaerocephalus; Helianthus tuberosus; Helianthus sp. ; Rudbeckia serotina; Solidago 
sempervirens; Solidago sp.; Tanacelum sp. 

Note: Life history studies [Felt; Barber and Weiss; Bailey]; eggs 
[Abbott]; causes damage in greenhouses [Osborn and Drake.]. 

*Corythucha mcelfreshi Drake 

Corythucha mcelfreshi Drake 1921b, p. 50, pi. 1, figs, b, b' [Mexico]. 
Type: Holotype o* , macropterous; Mexico; Drake Coll. (USNM). 
Distribution: Mexico. 
Host Plant: Unrecorded. 

*Corythucha mollicula Osborn and Drake 

Corythucha mollicula Osborn and Drake 1916b, p. 12, fig. 2 [Mich.]. — 
Drake 1921b, p. 53 [S.C.; poplar]; 1922d, p. 64, figs. 25c,d.— Hussey 
1922b, p. 23.— Parshley 1922a, p. 236; 1923b, p. 701 [Conn.].— 
McAtee 1923, p. 148 [Md.; Va.].— Froeschner 1944, p. 667 [Mo.].— 
Proctor 1946, p. 74 [Maine].— Bailey 1951, p. 90 [R.I.; Populus]; 
1959a, p. 67.— Feldman and Bailey 1952, pp. 98, 101-103, fig. 5. 
Corythucha salicis Osborn and Drake 1917b, p. 298 [Mass.; Wis.; Mont.; 
currant].— Gibson 1918, pp. 73, 76, 85 [Fla.; Ribes].— Drake 1919b, 
p. 159 [N.J.; N.Y.; Ont.; Man.; Salix discolor].— Parshley 1920a, 
p. 273 [Salix]; 1921, p. 17 [Salix hookeriana].— Downes 1925, p. 15, 
fig. 1. 

Corythucha molliculata [sic]: Gibson 1918, pp. 85, 90. 
Corythucha canadensis Parshley 1919a, p. 18 [B.C.]. 
Corythuca mollicula: Blatchley 1926, p. 461, fig. 102c. 
Type: Holotype 9, macropterous; East Lansing, Michigan, U.S.; Drake 
Coll. (USNM). 

Distribution: U.S. (Maine; N.H.; Mass.; R.I.; Conn.; N.Y.; N.J.; 
Pa.;Md.; Va.; S.G.; Ga.; Fla.; Mich.; Mo.; 111.; Ind.; Wis.; Kans.; N. Dak.; 



LACEBUGS OF THE WORLD 155 

S. Dak.; Nebr.; Idaho.; Mont.; Oreg.; Wash.); Canada (Man.; B.C.; Ont.). 
Host Plants: Populus sp.; Ribes sp.; Salix discolor; Salix hooker iana; Salix sp. 
*Corythucha montivaga Drake 

Corythucha montivaga Drake 1919a, p. 417 [Mont.]. — Hurd 1946, p. 484 
[Wyo.]. 
Type: Holotype 9, macropterous; Bear Pw. Mt., Montana, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Mont.; Wyo.; Utah). 
Host Plant: Unrecorded. 

Note: The holotype was erroneouly recorded as a o" in the original 
description. 
*Corythucha morrilli Osborn and Drake 

Corythucha morrilli Osborn and Drake 1917b, p. 298 [Colo; Ariz.]. — 
Gibson 1918, pp. 74, 95.— Drake 1919a, p. 419 [N. Mex.; Tex.; 
Calif.; Helianthus]; 1921b, p. 51 [ebony]; 1926b, p. 379; 1938a, p. 70.— 
Drake and Bruner 1924a, p. 154 [Jamaica; Cuba]. — Downes 1925, 
p. 16 [B.C.; Artemisia dracunculoides]. — Blatchley 1928, p. 4 [Fla.]. — 
Tilden 1950, p. 135 [Baccharis pilularis]. — Drake and Cobben 1960, 
p. 93, fig. 88c [Aruba; Curasao; Bonaire; St. Eustatius; Egletes 
prostrata; Ipomoea batatas; Helianthus annuus; Lagascea mollis; Synedrella 
nodiflora; Wedelia jacquini] . 
Corythucha decens (not Stal): Van Duzee 1917b, p. 214 (in part). 
Corythucha mexicana Gibson 1918, pp. 76, 95 [Mexico]. 
Corythucha morrillii [sic]: Gowdy 1926, p. 35. 
Corythucha morelli [sic]: Knowlton 1933, p. 263 [Utah]. 
Corythuca morrilli: Hardy 1954, p. 282 [Hawaii; Xanthium; Ambrosia 
artemisiijolia; Verbesina encelioides]. — Bianchi 1955a, p. 378 [Pluchea 
indica]; 1955b, p. 383 [Xanthium canadense]. 
Type: Holotype o 71 , macropterous; Fort Collins, Colorado, U.S.; Drake 
Coll. (USNM). 

Distribution: U.S. (Colo.; Utah; Calif.; Ariz.; N. Mex.; Tex.; Fla.; 
Hawaii); Canada (B.C.); Mexico; Guatemala; Cuba; Jamaica; Nether- 
lands Antilles (Aruba; Curagao; Bonaire); Leeward Islands (St. Eustatius). 
Host Plants: Ambrosia artemisiijolia; Artemisia dracunculoides; Baccharis 
pilularis; Erigeron canadensis; Egletes prostrata; Helianthus annuus; Helianthus 
sp.; Ipomoea batatas; Lagascea mollis; Pluchea indica; Synedrella nodiflora; Ver- 
besina encelioides; Wedelia jacquini; Xanthium canadense; Xanthium sp.; ebony. 

Note: Biological notes [Tilden]. Introduced into Hawaii. 
*Corythucha nicholi Drake 

Corythucha nicholi Drake 1928f, p. 3 [Ariz.]. 
Type: Holotype o", macropterous; Santa Rita Mountains, Arizona, 
U.S.; Drake Coll. (USNM). 
Distribution: U.S. (Ariz.). 
Host Plant: Unrecorded. 



156 U.S. NATIONAL MUSEUM BULLETIN 243 

*Corythucha nobilis Drake and Bondar 

Corythucha nobilis Drake and Bondar 1 '» ^2, p. ( )2 [Brazil; Solarium]. — 

a Lima 1936, p. \2(>. 
Corythuca nobilis: Silva L956, p. 27. 
I , ri : 1 [olotypc d", macroptcrous; Bahia, Brazil; Drake Coll. (L'SN'M . 
1 tei Kim noN: Brazil; Paraguay. 

I It «T l'l w r: Solarium sp. 
*Corythucha nocens Drake and Hamblcton (emendation) 

I rytkucha nocentis Drake and Hamblcton 1942a, p. 330 [Peru]; 1945, 
p. 366 [Ecuador]. 
Type: Holotype d\ macropterous; Canete, Peru; Drake Coll. (USNM). 
Distribution: Peru; Ecuador. 
Host l'i w i : Unrecorded. 

*Corythucha obliqua Osborn and Drake 

Corythucha obliqua Osborn and Drake 1916b, p. 11 [Calif.]; 1917b, p. 
301 [Oreg.; Idaho].— Gibson 1918, pp. 72, 76, 84 [Ceanothus].— 
Drake 1919a, p. 418. 
Corythucha maculata Van Duzee 1917a, p. 257 [Utah; Colo.]. 
Corythucha coelata [sic] (not Uhlcr): Van Duzee 1917a, p. 259. 
Corythucha fuscigera (not Stal): Van Duzee 1917b, p. 213 (in part). 
Corythucha contaminata Gibson 1918, pp. 72, 76, 82 [Ceanothus cardulatus]. 
Type: Holotype 9, macropterous; Dutch Flats, California, U.S.; Drake 
Coll. (USNM). 

Distribution: U.S. (Calif.; Oreg.; Idaho; Wash.; Utah.; Colo.). 
Host Plants: Ceanothus cardulatus; Ceanothus sanguineus; Ceanothus sp. 

*Corythucha omani Drake 

Corythucha omani Drake 1941a, p. 144 [Ariz.]. 
Type: Holotype o 71 , macropterous; NogaJes, Arizona, U.S.; USNM. 
Distribution: U.S. (Ariz.). 
Host Plant: Unrecorded. 

*Corythucha padi Drake 

Corythucha padi Drake 1917a, p. 215 [Mont.; Oreg.; choke cherry]; 
1919b, p. 159 [Idaho]; 1932, p. 100.— Osborn and Drake 1917b, 
p. 303 [Wash.; B.C.; Primus dem issa] .—Gibson 1918, pp. 71, 75, 88 — 
Parshlcy 1919a, p. 19 [alder; hazel].— Dovvncs 1925, p. 15, fig. 3 
[walnut].— Knovvlton 1933, p. 262 [Utah]. 
Corythucha coloradensis Gibson 1918, pp. 72, 76, 89 [Colo.; Nebr.; Rhus 

toxicodendron]. 
Corythucha fuscigera (not Stal): Gibson 1918, p. 78 (in part). 
Type: Holotype 9, macropterous; Missoula, Montana, U.S.; Drake 
Coll. (USNM 

DlSTRlBl H'j\: U.S. (Mont.; Colo.; Utah; Ores?.; Wash.; Idaho; Nebr.); 
Canada (B.C.). 



LACEBUGS OF THE WORLD 157 

Host Plants: Primary host Prunus demissa; also recorded from Rhus 
toxicodendron; hazel; alder; walnut. 

*Corythucha pallida Osborn and Drake 

Corythucha pallida Osborn and Drake 1916a, p. 230 [Ohio; linden]; 
1917b, p. 300 [Md.; Tenn.; Va.; Ariz.; Tilia americana; Morus 
rubra].— McAtee 1917b, p. 78 [D.C.]; 1923a, p. 148.— Gibson 1918, 
pp. 76, 100.— Drake 1925c, p. 36 [Miss.].— Froeschner 1944, p. 667 
[Mo.].— Proctor 1946, p. 74 [Maine].— Bailey 1951, p. 91 [Podophyl- 
lum peltatum] . 
Corythuca pallida: Blatchley 1926, p. 459 [Ind.].— Blote 1945, p. 83 
[N.C.]. 
Type: Sex unknown; Columbus, Ohio, U.S.; Ohio State Univ. 
Distribution: U.S. (Maine; Md.; D.G.; Va.; W. Va.; Ohio; 111.; Tenn.; 
Miss.; Mo.; Ind.; Ga.; N.C.; S.C.; Ala.; Ariz.). 

Host Plants: Morus rubra; Podophyllum peltatum; Tilia americana. 

*Corythucha pallipes Parshley 

Corythucha juglandis (not Fitch): Van Duzee 1917b, p. 215 (in part). 
Corythucha pallipes Parshley (in Gibson 1918, pp. 73, 77, 82) [Conn.; 
Salix sieboldiana]; 1919b, p. 70; 1923b, p. 701 [Salix] .—Drake 1922d, 
p. 65; 1922f, pp. 111-116, pi. 4, fig. 44 [Mich.; Wis.; Oreg.; Wash.; 
Man.; Betula alba; Fagus grandiflora; Ostrya virginiana; Pyrus americana; 
Acer spicatum; Acer saccharinum; Acer saccharum; Acer pennsylvanicum]. — - 
Bailey 1951, p. 91 [Vt.; Que.; N.B.; Betula papyri/era].— Fcldman 
and Bailey 1952, pp. 98, 101-103, fig. 17.— Lindberg 1958b, p. 14 
[Nfld.]. 
Corythucha betulae Drake (in Gibson 1918, pp. 71, 75, 86) [N.Y.; Betula 

lutea]; 1919b, p. 159 [Ont.; Betida lento].— Parshley 1919b, p. 70. 
Corythucha cyrta Parshley {in Gibson 1918, pp. 75, 86) [Maine; Mass.; 

N.H.]; 1919b, p. 70; 1920a, p. 272. 
Corythucha paleipes [sic]: Barber 1922a, p. 16 [N.J.]. 
Corythuca pallipes: Blatchley 1926, p. 457, fig. 100. — Drake and Davis 
1960, fig. 18. 
Type: Holotype 9; Stamford, Connecticut, U.S.; Cal. Acad. 
Distribution: U.S. (Maine; Mass.; Vt.; N.H.; Conn.; N.Y.; N.J.; 
Oreg.; Wis.; Mich.; Wash.); Canada (Que.; Man.; N.B.; Ont.; Nfld.). 
Host Plants: Acer pennsylvanicum; Acer saccharinum; Acer saccharum; 
Acer spicatum; Betula alba; Betula lenta; Betula lutea; Betula papyri/era; Fagus 
grandifolia; Ostrya virginiana; Pyrus americana; Salix sieboldiana; Salix sp. 
Note: Life history studies [Drake 1922b; Bailey]; predaceous insects 
[Bailey; Drake]. 



l.'tS U.S. NATIONAL MUSEUM BULLETIN 213 

*Corythucha polmatifl Drake 

Corythucha palmatis Drake 1929, p. 37 [Costa Rica]. 
Type: Holotypc d"; San Jose, Costa Rica; Hamburg Mus. 
Distribii [i >\: ( dm. i Rica. 
Host Pi. an i : I fare* orded. 
*Corythucha pellucida Drake and Hambleton 

Corythucha pellucida Drake and Hambleton 1938a, p. 67 [Brazil; Celtis 
ferruginea]. — Monte 193913, p. 68 [Celtis brasiliensis]. 
Type: Holotypc d\ macropterous; Campinas, Sao Paulo, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plants: Celtis brasiliensis; Celtisjerruginea. 

*Corythucha pergandei Heidemann 

Corythuca pergandei Heidemann 1906, p. 10, figs. 2, 3 [D.C.; Pa.; Va.; 

Mass.; 111.; Kans.; elms; crab apple; hazel; Alnus]. — Smith 1910, p. 

148 [N.J.; N.Y.].— Parshley 1914, p. 145 [Maine].— Blatchley 

1926, p. 466, fig. 104 [Ga.]. 

Corythucha pergandei: Osborn and Drake 1916a, p. 228, figs. 4, 5 [Ohio]; 

1917b, p. 300 [Wis.; Md.; Tenn.].— Parshley 1917b, p. 54 [N.H.; 

Conn.]; 1923a, p. 102 [N.S.].— Van Duzee 1917b, p. 213 [Ont.].— 

Gibson 1918, pp. 71, 75, 91 [Calif.; Tex.; Betula nigra; Betula lenta; 

Alnus rugosa; Pyrus prunifolia; Ulmus]. — Weiss and Dickerson 1918a, 

p. 205, fig. [Alnus glutinosa; Betula lutea; Betula populijolia]. — Barber 

and Weiss 1922, p. 6, fig. 2.— Hussey 1922b, p. 23 [Mich.; Alnus 

incana]. — McAtee 1923, p. 148 [Prunus americana; Celtis crassifolia]. — 

Torre-Bueno 1926a, p. 54.— Drake 1928b, p. 100.— Bailey 1951, p. 

94 [Vt.].— Feldman and Bailey 1952, pp. 96, 98, 101-103, figs. 1, 2. 

Type: Holotype d\ macropterous; Washington, D.C., U.S.; USNM. 

Distribution: U.S. (Maine; N.H.; Vt.; Mass.; Conn.; N.Y.; N.J.; 

Pa.; Md.; D.C.; Va.; Ga.; Tenn.; Ky.; Ohio; Ind.; 111.; W. Va.; Mich.; 

Wis.; Minn.; Iowa; Mo.; Nebr.; Kans.; Okla.;Tex.; Calif.); Canada (Ont.: 

.VS.; B.C.). 

HOST Plants: Primary — Alnus glutinosa; Alnus incana; Alnus rugosa; 
Alnus sp.; also recorded from: Betula lenta; Betula lutea; Betula nigra; Betula 
populijolia; Celtis crassifolia; Corylus americana; Prunus americana; Pyrus prum- 
folia; I'lmus sp. 

Note: Life history studies [Weiss and Dickerson; Bailey]; taken at light 
[McAtee]; ovipositor [Feldman and Bailey]. 

*Corythucha pruni Osborn and Drake 

■ mi < >sborrj and Drake 1916a, p. 231 [D.C.; Prunus serotina]; 
I'M 7b, p. 303.— McAtee 1917b, p. 79.— Gibson 1918, pp. 71, 76, 
80 [N.Y.; N.C.; Ohio].— Hussey 1922b, p. 23 [Mich.].— Bailey 



LACE BUGS OF THE WORLD 159 

1951, p. 96, fig. 4, graphs 3-5 [Vt.; Mass.]; 1959, p. 67 [Conn.; 
R.I.].— Feldman and Bailey 1952, pp. 96, 98, 102, 103, figs. 4, 9. 
Corythucha hoodiana Osborn and Drake 1917b, p. 302 [Oreg.]. — Gibson 

1918, pp. 76,82. 
Corythucha exigaa Drake (in Gibson 1918, pp. 76, 83). 
Corythucha pyriformis Parshley 1920c, p. 81 [Maine; N.H.]; 1922a, p. 

236. 
Corythuca pruni: Blatchley 1926, p. 465. 
Corythuca exigua: Blatchley 1926, p. 467. 
Type: Sex unknown; Washington, D.C., U.S.; Ohio State Univ. 
Distribution: U.S. (Maine; N.H.; Conn.; R.I.; Mass.; Vt.; N.Y.; 
Md.; D.C.; Va.; N.C.; Ga.; Ohio; Mich.; Ind.; 111.; Mo.; Iowa; Nebr.; 
Utah; Oreg.). 
Host Plant: Prunus serotina. 

Note: Life history study [Bailey]; leaf injury [Bailey]; ovipositor [Feld- 
man and Bailey]. 
*Corythucha sagillata Drake 

Corythucha sagillata Drake 1932, p. 101 [Ariz.; Vauquelinia calif ornica]. 
Type: Holotype 9, macropterous; Santa Catalina Mountains, Arizona, 
U.S.; Drake Coll. (USNM). 
Distribution: U.S. (Ariz.). 
Host Plant: Vauquelinia calif ornica. 

*Corythucha salicata Gibson 

Corythucha salicata Gibson 1918, pp. 77, 90 [Oreg.].— Parshley 1919a, 
p. 20 [B.C.]; 1921, p. 17 [Salix lasiandra].— Drake 1921b, p. 53 [Wash.; 
Man.; poplar; alder]. — Downes 1925, p. 16, fig. 2 [Salix scouleriana]. — ■ 
Thompson and Wong 1933, pp. 1090-1095, fig. 95. 
Corythucha drakei Gibson 1918, pp. 71, 77, 98 [Calif.; Pyrus malus]. 
Corythucha essigi Drake 1918c, p. 385 [maize]. 
Corythucha platini Drake 1920, p. 49 [sycamore]. 
Type: Holotype 9, macropterous; Hood River, Oregon, U.S.; Drake 
Coll. (USNM). 

Distribution: U.S. (Oreg.; Wash.; Calif.; Idaho; Utah); Canada 
(Man.; B.C.). 

Host Plants: Salix lasiandra; Salix scouleriana; Pyrus malus; £ea mays; 
poplar; alder; sycamore. 
Note: Life history study [Thompson and Wong]. 

*Corythucha scitula Drake 

Corythucha scitula Drake 1948c, p. 24 [Oreg.; Corylus rostrata]. 
Type: Holotype o 71 , macropterous; Stayton, Oregon, U.S.; Drake Coll. 
(USNM). 
Distribution: U.S. (Oreg.). 
Host Plant: Corylus rostrata. 



160 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Corythucha seguyi I):akc 

lyi Drake 1921b, p. 51, pi. 1, fig. c [Bolivia]. — Drake and 
M imbleton 1945, p. 366 [Peru]. 
[Vpe: Sex unknown; Cochabamba, Bolivia; Paris Mus. 
Distribi noN: Bolivia; Peru. 
Hon I'i \\ i : I Unrecorded. 

*Corythucha serta Drake and I lamblcton 

'/ Drake and Hamblcton 1945, p. 366 [Guatemala]. 

Type: Holotype o\ macropterous; Lake Atitlan, Guatemala; Drake 
C.ll. (USNM . 

Distribution: Guatemala; Mexico. 

Host Plant: Unrecorded. 

*Corythucha setosa Champion 

Corythucha setosa Champion 1897, p. 8, pi. 1, figs. 10, 10a [Guatemala]. — 
Van Duzee 1917b, p. 215 [Calif.].— Gibson 1918, pp. 77, 100 [Mex- 
ico]. — Drake and Hambleton 1946b, p. 122. 
Type: Sex unknown; Cerro Zunil, Guatemala; British Mus. 
Distribution: Guatemala; Mexico; U.S. (Calif.). 
Host Plant: Unrecorded. 

Corythucha socia Monte 

Corythucha socia Monte 1940e, p. 296 [Brazil; Euphorbiaceae]. 

Type: Holotype o"; Lapinha, Lagoa Santa, Brazil; Monte Coll. (Mus. 
Nacional). 

Distribution: Brazil. 

Host Plant: Euphorbiaceae. 

*Corythucha sphaeralceae Drake 

Corythucha hispida (not Uhler) : Van Duzee 1917b, p. 214 (in part) 

[Ariz.].— Knowlton 1931, p. 43 [Utah]. 
Corythucha sphaeralceae Drake 1920, p. 51 [Calif.; Sphaeralcea]. 
Corythucha pacific a Drake 1920, p. 52 [Wash.]. 
Type: Holotype 9, macropterous; Deep Springs Valley, Inyo County, 
California, U.S.; Drake Coll. (USNM). 
Distribution: U.S. (Calif.; Ariz.; Utah). 
Host Plant: Sphaeralcea sp. 

( orythucha spinosa (Duges) 

Tingis spinosa Duges 1889, p. 207, pi. 18 [Mexico; Ricinus communis]. 
Tingis hyalina (not Hcrrich-Schaeffer) : Guerin-Mrnrville 1857, p. 408 
[Cuba]. 

'hucha spinosa: Champion 1897, p. 8, pi. 1, figs. 9, 9a; 1898b, p. 
57— Van Duzee 1917b, p. 214 [Calif.].— Gibson 1918, pp. 77, 
99. — Drake and Bruner 1924a, p. 154 [Bambusa]. Perkins and Swezey 

1924, p. 52 [Lantana camara]. — Drake 1926a, p. 87. — Bruner, Scara- 



LACEBUGS OF THE WORLD 161 

muzza and Otero 1945, pp. 97, 168, 184 [Solarium torvum; Triumfetta 
grossulariaefolia] . 
Corythucha unifasciata (not Champion): Gibson 1918, pp. 76, 97. 
Type: Silao de la Victoria, Guanajuato, Mexico; sex and deposition of 
type unknown. 
Distribution: Mexico; Cuba; Trinidad; Costa Rica; U.S. (Calif.). 
Host Plants: Lantana camara; Bambusa sp.; Solarium torvum; Ricinus 
communis; Triumfetta grossulariaefolia. 
Corythucha translucida Monte 

Corythucha translucida Monte 1946c, p. 28 [Bolivia]. 
Type: Holotype o 71 ; La Paz, Bolivia; Monte Coll. (Mus. Nacional). 
Distribution: Bolivia. 
Host Plant: Unrecorded. 

*Corythucha tuthilli Drake 

Corythucha tuthilli Drake 1940a, p. 172 [Colo.]. 

Type: Holotype d\ macropterous; Creede, Colorado, U.S.; Drake 
Coll. (USNM). 

Distribution: U.S. (Colo.). 

Host Plant: Unrecorded. 

*Corythucha ulmi Osborn and Drake 

Corythucha pallida var. ulmi ? Osborn and Drake 1916a, p. 231 [Ohio; 

Ulmus americana]. 
Corythucha ulmi: Gibson 1918, pp. 77, 97 [Md.; N.Y.; S.C.].— Parshley 
1922a, p. 236 [N.H.]; 1922b, p. 11 [S. Dak.]; 1923b, p. 702 
[Conn.].— McAtee 1923, p. 148 [Va.; Ulmus fulva].— Felt 1933, p. 
49.— Froeschner 1944, p. 668 [Mo.].— Hurd 1946, p. 484 [Nebr.; 
Minn.]. — Bailey 1951, p. 105 [Vt.; Mass.; Ulmus thomasii]; 1959, p. 
69.— Feldman and Bailey 1952, pp. 98, 102, 103, fig. 11.— Ash 
1954, p. 185 [Ind.; Ulmus glabra camperdownii]. 
Corythuca ulmi: Blatchley 1926, p. 464. 
Type: Neotype 9, macropterous; Plummer's Island, Maryland, U.S.; 
Drake Coll. (USNM). 

Distribution: U.S. (N.H.; Vt.; Mass.; Conn.; N.Y.; Md.; Va.; S.C; 
Miss.; Ohio; Ind.; Mich.; 111.; Wis.; Minn.; Iowa; Mo.; Nebr.; S. Dak.). 
Host Plants: Ulmus americana; Ulmus fulva; Ulmus glabra camperdownii; 
Ulmus thomasii. 

*Corythucha unifasciata Champion 

Corythucha unifasciata Champion 1897, p. 7, pi. 1, figs. 8, 8a [Mexico; 
Guatemala; Panama]. 
Type: Sex unknown; Las Mercedes, Guatemala; British Mus. 
Distribution: Mexico; Guatemala; Panama. 
Host Plant: Unrecorded. 



L62 U.S. NATIONAL Ml SKIM BULLETIN 243 

Genus COTTOTHUCHA Drake and Poor 

Cottothucha Drake and Poor 1941, p. 162. -Drake and Ruhoff 1960a, 
p. 49. 
Type Species: Cottothucha oceanae Drake and Poor. 

(uttothucha oceanae Drake and Poor 

Cottothucha oi, u u hi. ike and Poor 1941, p. 163 [Amboina]. — Drake 
and Ruhoff 1961b, |). 182, Bg. 23 [Luzon]. 
Type: Eiolotype o"; Amboina Inland; Cal. Acad. 
Distribution: Moluccas (Amboina); Philippine Islands (Luzon). 
Hosi Plant: Raphidophora merrUlii. 

Genus CROMERUS Distant 

Cromerus Distant 1902b, p. 355.— Drake 1926c, p. 334.— Drake and 
Ruhoff 1960a, p. 49. 
Type Species: Monanthia invar ia Walker. 

*Cromerus bakeri Drake 

Cromerus bakeri Drake 1930e, p. 166 [Samar]. 

Type: Holotype 9, macropterous; Samar Island, Philippine Islands; 
Drake Coll. (USNM). 

Distribution: Phillippinc Islands (Samar). 

Host Plant: Unrecorded. 

*Cromerus excelans Drake 

Cromerus invarius (not Walker): Drake 1930e, p. 167 [Samar; 

Mindanao]. 
Cromerus excelans Drake 1953b, p. 91. 
Type: Holotype o 71 , macropterous; Butuan, Mindanao, Philippine 
Islands; Drake Coll. (USNM). 

Distribution: Philippine Islands (Samar; Mindanao). 
I Iost Plant: Unrecorded. 

*Cromerus gressitti Drake 

Cromerus gressitti Drake 1937b, p. 591 [China]. 

Type: Holotype o" , macropterous; Yim-na Shan, E. Kwangtung, China; 
I hake Coll. (USNM). 

Distribution: China. 

1 1< >■-] Pi \n i : I nrecorded. 

♦Cromerus hackeri Drake and Poor 

Cromerus hacker i Drake and Poor 1939c, p. 203 [Australia]. 

Type: Eiolotype -\ macropterous; Cedar Creek, Queensland, Australia; 
Drake Coll. (I SNM 

Distribution: Australia (Queensland). 

I [( 8T Plant: Unrecorded. 



LACEBUGS OF THE WORLD 163 

*Cromerus invarius (Walker) 

Monanthia invaria Walker 1873a, p. 196 [New Guinea]. 

Cromerus invaria: Distant 1902b, p. 356. — Drake 1960a, p. 355. 
Type: Sex unknown; New Guinea; British Mus. 
Distribution : New Guinea. 
Host Plant: Unrecorded. 

*Cromerus kalshoveni Drake 

Cromerus kalshoveni Drake 1926e, p. 334 [Java; Vitex heterophylla]; 1930e, 
p. 167 [Mindanao]. 
Type: Holotype 9, macropterous; Kediri, Java; Drake Coll. (USNM). 
Distribution: Greater Sunda Islands (Java); Philippine Islands (Min- 
danao). 
Host Plant: Vitex heterophylla. 

*Cromerus palawanus Drake and Maa 

Cromerus palawanus Drake and Maa 1955, p. 6 [Palawan]. 

Type: Holotype 9, macropterous; Binaluan, Palawan, Philippine Is- 
lands; Drake Coll. (USNM). 

Distribution : Philippine Islands (Palawan) . 

Host Plant: Unrecorded. 

*Cromerus viadoi Drake and Maa 

Cromerus viadoi Drake and Maa 1955, p. 5 [Mindanao]. 

Type: Holotype o", macropterous; Mindanao, Philippine Islands; Drake 
Coll. (USNM). 

Distribution: Philippine Islands (Mindanao). 

Host Plant: Unrecorded. 

Genus CYSTEOCHILA Stal 

Cysteochila Stal 1873, pp. 121, 129. — Lethierry and Severin 1896, p. 
22.— Distant 1903b, p. 138.— Bergroth 1921, p. 104— Horvath 
1925a, p. 3.— Drake and Poor 1937a, p. 6.— Drake 1956b, p. 110. — 
Drake and Runoff 1960a, p. 49.— Stichel 1960b, p. 400. 

Bredenbachius Distant 1903a, p. 50; 1903b, p. 139. — Drake and Ruhoff 
1960a, p. 40. 
Type Species: Monanthia ? tingoides Motschulsky. 

*Cysteochila abetti Schouteden 

Cysteochila abetti Schouteden 1919, p. 140 [Congo]; 1923, p. 103. — 
Drake 1956a, p. 4 [Uganda]. 
Type: Sex unknown; Kilo, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Uganda. 
Host Plant: Unrecorded. 



164 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*CysteochiIa ablusa Drake 

I . teochila ablusa Drake 1948a, p. 6 [India]. 
Typi i Holotype o", macropterous; Barwai, India; Drake Coll. (USNM). 
1 tei ribution: India. 

Il('si l'i \m: Iuu/hinia variegata. 

*Cysteochila abundans Drake and Poor (emendation) 

teochila abundantis Drake and Poor l ( J37a, p. 8 [Mindanao]; 1937b, 
p. 400 [Luzon]. Singh 1953, p. 118. 
Type: Holotype o"; Tangkulan, Bukidnon Province, Mindanao, Philip- 
pine Islands; Drake Coll. (USNM). 

1 ta i i-ii'i i [( in : Philippine Islands (Mindanao; Luzon). 
Host Plant: Unrecorded. 

*CysteochiIa aletheia Drake and Ruhoff 

teochila aletheia Drake and Runoff 1962b, p. 251 [Papua]. 

Type: Holotype o 71 , macropterous; Bisiatabu, Port Moresby, Papua 
Territory, New Guinea; So. Austr. Mus. 

Distribution: New Guinea (Papua). 

Host Plant: Unrecorded. 

Cysteochila angulipennis Horvath 

Cysteochila angulipennis Horvath 1929, p. 323 [Zanzibar]. 
Type: Holotype d"; Zanzibar; British Mus. 
Distribution: Zanzibar. 
Host Plant: Unrecorded. 

Cysteochila annandalei (Distant) 

Bredenbachius annandalei Distant 1909a, p. 119 [India]; 1910a, p. 120. 
Type: Sex unknown; Calcutta, India; British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 

Cysteochila aspera Drake and Poor 

Cysteochila aspera Drake and Poor 1941, p. 160 [Amboina]. 
Type: Holotype d"; Amboina Island; Cal. Acad. 
Distribution: Moluccas (Amboina). 
Host Plant: Unrecorded. 

*Cysteochila bakeri Drake and Poor 

Cysteochila bakeri Drake and Poor 1937a, p. 9 [Mindanao; Luzon]; 1937b, 
p. 399. 

Type: Holotype 9, macropterous; Surigao, Mindanao, Philippine Islands; 
USNM. 

DISTRIBUTION: Philippine Islands ( Mindanao; Luzon). 

I [OST l'i \nt: Unrecorded. 



LACEBUGS OF THE WORLD 165 

Cysteochila basilewskyi Schouteden 

Cysteochila basilewskyi Schouteden 1957c, p. 316 [Ruanda]. 
Type: Sex unknown; Ruanda; Gent. Afr. Mus. 
Distribution: Ruanda-Urundi. 
Host Plant: Unrecorded. 

*Cysteochila biseriata Schouteden 

Cysteochila biseriata Schouteden 1916b, p. 295 [Congo]; 1923, p. 103. — 
Mayne and Ghesquiere 1934, p. 20 [Hevea]. 
Type: Sex unknown; Congo da Lemba, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Hevea sp. 

Cysteochila bredoi Schouteden 

Cysteochila bredoi Schouteden 1953b, p. 117 [Congo]. 
Type: Sex unknown; Bunia, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Cysteochila brunnea Hacker 

Cysteochila brunnea Hacker 1928, p. 180, pi. 23, fig. 13 [Queensland]. 
Type: Sex unknown; Cairns District, Australia; Queensland Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Cysteochila burgeoni Schouteden 

Cysteochila burgeoni Schouteden 1953b, p. 115 [Congo]. 
Type: Sex unknown; Rutshuru, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Cysteochila caffra Stal 

Cysteochila caffra Stal 1873, p. 129 [Caffraria].— Distant 1902a, p. 242, 
pi. 15, fig. 8. 
Type: Holotype 9; Caffraria, South Africa; Stockholm Mus. 
Distribution: South Africa; Malagasy Republic. 
Host Plant: Unrecorded. 

*Cysteochila chiniana Drake 

Cysteochila chiniana Drake 1942a, p. 5 [China]. 

Physatochila fieberi (not Scott): Takeya 1951a, p. 20 [Japan; Cissus 

japonica]. 
Physatocheila fieberi: Takeya 1953d, p. 175. 
Type: Holotype cf, macropterous; Tien Sun Shan, China; Drake Coll. 
(USNM). 
Distribution: China; Japan; Taiwan. 
Host Plant: Cissus japonica. 



166 U.S. NATIONAL MUSEUM BULLETIN 243 

Cysteochila collarti Schouteden 

teochUa collaiti Schouteden 1953b, p. 116 [Congo]. 
Type: Sex unknown; Stanleyville, Belgian Congo; Cent. Afr. Mus. 
Distribui [i >\ I 
Host Plant: Unrecorded. 

*Cysteochila consanguinea (Distant) 

Unbachius consangtuneus Distant 1909a, p. 118 [Burma]; 1910a, p. 
119. 
Cysteochila consanguiensis [sic]: Drake and Poor 1936a, p. 439 [Hainan]. 
Cysteochila consanguineus: Drake 1937b, p. 593 [China]. 
Type: Sex unknown; Tenasserim, Burma; British Mus. 
Distribution: Burma; India; China (Hainan Island). 
Host Plant: Unrecorded. 

*Cysteochila constans (Drake) emendation 

Oncophysa constantis Drake 1927a, p. 54 [Hong Kong]. — Wu 1935, p. 

449. 
Cysteochila constantis: Drake and Poor 1937a, p. 11. 
Type: Holotypc 9, macropterous; Hong Kong; Drake Coll. (USNM). 
Distribution: Hong Kong. 
Host Plant: Unrecorded. 
*Cysteochila consueta Drake 

Cysteochila consueta Drake 1948a, p. 5, fig. 2 [Taiwan; Viet-Nam]. — 
Takeya 1951a, p. 21. 
Type: Holotype o"; Formosa; Drake Coll. (USNM). 
Distribution: Taiwan; Viet-Nam. 
Host Plant: Unrecorded. 
Cysteochila cracens Drake (emendation) 

Cysteochila cracentis Drake 1954b, p. 3 [Western Australia]. 
Type: Holotype d"; Yancep, Western Australia, Australia; British Mus. 
Distribution: Australia (Western Australia). 
Host Plant: Unrecorded. 
*Cysteochila cremeri Drake 

Cysteochila cremeri Drake 1955d, p. 2, fig. 1 [Congo]. 
Type: Holotype o*; Libenge, Belgian Congo; Brussels Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 
^Cysteochila delineata (Distant) 

Bredenbachius delineatus Distant 1909a, p. 119 [India]; 1910a, p. 119, 

fig. 61. 
Cysteochila delineatus: Drake 1933, p. 1015. — Drake and Poor 1936b, 
p. 114 [Bauhinia purpurea]. — Singh 1953, p. 118. 
Type: Sex unknown; Calcutta, India; Indian Mus. 
Distribution: India. 
Host Plant: Bauhinia purpurea. 



LACEBUGS OF THE WORLD 167 

*Cysteochila dikoana Drake 

Cysteochila dikoana Drake 1954b, p. 4 [Nigeria]. 
Type: Holotype 9, macropterous; Dikoa, Nigeria; Drake Coll. (USNM). 
Distribution: Nigeria; Congo. 
Host Plant: Unrecorded. 

Cysteochila emmelia Drake 

Cysteochila emmelia Drake 1960, p. 347 [New Guinea]. 
Type: Holotype o 71 ; Mt. Wilhelm, Northeast New Guinea; Bishop Mus. 
Distribution: New Guinea (Northeast). 
Host Plant: Unrecorded. 

♦Cysteochila endeca Drake 

Cysteochila endeca Drake 1954b, p. 4 [Senegal; Ghana; Uganda; Cape 

Province]. 
Cysteochila endica [sic]: Lindberg 1958a, p. 79 [Cape Verde; Tamarindus 
indica]. 
Type: Holotype o 71 ; Bambey, Senegal; British Mus. 
Distribution: Senegal; Ghana; Uganda; South Africa (Cape Province); 
Cape Verde Islands. 
Host Plant: Tamarindus indica. 

*Cysteochila euphues Drake and Ruhoff 

Cysteochila euphues Drake and Ruhoff 1962a, p. 157 [Larat]. 

Type: Holotype 9, macropterous; Larat Island, Tanimbar Islands, E. 
Malay Archipelago; Cal. Acad. 

Distribution: Tanimbar Islands (Larat). 

Host Plant: Unrecorded. 

Cysteochila exigua Drake 

Cysteochila exigua Drake 195 7f , p. 125 [Malagasy]. 
Type: Holotype 9; Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

*Cysteochila expleta Drake and Maa 

Cysteochila expleta Drake and Maa 1954, p. 112 [India]. 
Type: Holotype o 71 ; Kodai Kanal, Madras, India; British Mus. 
Distribution: India; Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

Cysteochila faceta Drake 

Cysteochila faceta Drake 1958c, p. 316 [Malagasy]. 
Type: Holotype 9; Mont Tsaratanana, Madagascar; Madagascar Sci. 
Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 



ltiS l\S. NATIONAL Ml SKIM BULLETIN 243 

*Cysteochila fieberi (Scott) 

Monanthiafieberi Scott 1874, pp. 291, 442 [Japan]. 

Physatochila fieberi: LethieXTy and Scverin 1896, |>. 21. 

Physatochila montrosa (not Scott): Takeya 1951a, p. 20. 

Physatocheila monstrosa: Takeya 1953d, p. 173, pi. 6, fig. 1 [Boehmeria 
spicata] . 

CysUochila fieberi: Drake and Maa 1954, p. 112 [India; China; Taiwan]. 
Tvi'i : Sex unknown; Japan; British Mus. 
Disi Kim i [i in: Japan; China; Taiwan; India. 
Host PLANT: Boehmeria spicata. 

Cysteochila ghesquierei Schonteden 

leochila ghesqtderei Schouteden 1953b, p. 117 [Congo; Pseudospondias 
microcarpa]. 
Type: Sex unknown; Bengamissa, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Pseudospondias microcarpa. 

*Cysteochila gibbula (Horvath) 

Physatocheila gibbula Horvath 1912c, p. 342, fig. 2 [Java]. 

Cysteochila gibbula: Drake and Runoff 1961c, p. 145. 
Type: Holotype o"; Banjuwangi, Java; Hungarian Mus. 
Distribution: Greater Sunda Islands (Java). 
Host Plant: Unrecorded. 

*Cysteochila hackeri Drake 

Cysteochila hackeri Drake 1939a, p. 87 [South Australia]. 
Type: Holotype 9, macropterous; Melrose, Australia; Drake Coll. 
(USNM). 
Distribution: Australia (South Australia). 
Host Plant: Unrecorded. 

:!: Cysteochila humeralis (Distant) 

Bredenbachius humeralis Distant 1909a, p. 120 [India]; 1910a, p. 121, 
fig. 62. 
Type: Sex unknown; Dhikala, Naini Tal, Upper Province, India; 
Indian Mus. 
Distribution: India. 
Host Plant: Unrecorded. 

*Cysteochila humerella Drake 

CysUochila humerella Drake 1958c, p. 317 [Malagasy]. 
Type: Holotype cf; Anal avclona, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 

I ]• >< i l'i \n i : I "in (< < i] c 1<-(1. 



LACEBUGS OF THE WORLD 169 

*Cysteochila idonea Drake 

Cysteochila idonea Drake 1956d, p. 109. fig. 4 [Kusaie; Three Sisters]; 
1960, p. 345, fig. 5 [New Guinea]. 
Type: Holotype d", macropterous; Mt. Matante, Kusaie, Caroline 
Islands; USNM. 

Distribution: Caroline Islands (Kusaie); Solomon Islands (Three 
Sisters); New Guinea (Northeast). 
Host Plant: Unrecorded. 

*Cysteochila impressa Horvath 

Cysteochila impressa Horvath 1910, p. 66 [Tanganyika]. — Drake 1954a, 
p. 232 [Senegal]; 1957f, p. 127 [Malagasy]. 
Type: Holotype o" ; Kibongoto, Kilimandjaro; Stockholm Mus. 
Distribution: Tanganyika; Kenya; Senegal; Malagasy Republic. 
Host Plant: Unrecorded. 

*Cysteochila incolana Drake 

Cysteochila incolana Drake 1956h, p. 18 [Cape Province]; 1961c, p. 129, 
pi. 8. 
Type: Holotype o 71 ; Lion's Head, Capetown, South Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

Cysteochila ituriensis Schouteden 

Cysteochila ituriensis Schouteden 1953b, p. 121 [Congo]. 
Type: Sex unknown; Bondo Mabe, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Cysteochila jacobsoni (Blbte) 

Bredenbachius jacobsoni Blote 1945, p. 89 [Java]. 
Type: Sex unknown; Gunung Gedeh, Java; Leiden Mus. 
Distribution: Greater Sunda Islands (Java). 
Host Plant: Unrecorded. 

*Cysteochila javensis Drake and Poor 

Cysteochila javensis Drake and Poor 1937b, p. 400 [Java]. 

Type: Holotype 9, macropterous; Pekalongan, Java; Drake Coll. 
(USNM). 

Distribution: Greater Sunda Islands (Java); India. 

Host Plant: Unrecorded. 

Cysteochila jimmina Drake 

Cysteochila jimmina Drake 1960, p. 349 [New Guinea]. 
Type: Holotype 9; Kumur, Upper Jimmi Valley, Northeast New 
Guinea; Bishop Mus. 
Distribution: New Guinea (Northeast). 
Host Plant: Unrecorded. 

681-552 — 65 12 



170 U.S. NATIONAL MUSEUM BULLETIN 243 

Cysteochila josephinae (Schouteden) 

Physatochila josephinae Schouteden 1923, p. 106 [Congo]. 
teochila josephmat: Drake and Runoff 1961c, p. 145. 
I \ i'i .: Sex unknown; Vista, Belgian ( Songo; ( lent. Afr. Mus. 
Distribution: Congo. 
Him Plant: Unrecorded. 
Cysteochila katangae Schouteden 

rUochila katangae Schouteden 1953b, p. 123 [Congo]. 
Type: Sex unknown; Elizabethvillc, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Hosi Plant: Unrecorded. 
Cysteochila kintambo Schouteden 

Cysteochila kintambo Schouteden 1953b, p. 118 [Congo]. 
Type: Sex unknown; Leopoldvillc, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 
*Cysteochila lecta Drake and Poor 

Cysteochila lecta Drake and Poor 1937a, p. 7 [Borneo]. 
Type: Holotype 9, macropterous; Sandakan, Borneo; USNM. 
Distribution: Greater Sunda Islands (Borneo). 
Host Plant: Unrecorded. 
Cysteochila lita Drake 

Cysteochila lita Drake 1960, p. 345, fig. 6 [New Guinea]. 
Type: Holotype cT; Wisselmeren, Enarotadi, Netherlands New Guinea; 
Bishop Mus. 
Distribution: New Guinea (Netherlands). 
Host Plant: Unrecorded. 
Cysteochila lueboensis Schouteden 

Cysteochila lueboensis Schouteden 1953b, p. 119 [Congo; Hevea]. 
Type: Sex unknown; Luebo, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Hevea sp. 
*Cysteochila luzona Drake and Maa 

Cysteochila luzona Drake and Maa 1955, p. 4 [Luzon]. 
Type: Holotype 9, macropterous; Mt. Makiling, Luzon, Philippine 
Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Hosi I'i wt: Unrecorded. 
*Cysteochila malaisaei Drake and Ruhoff 

tochila malaisaei Drake and Ruhoff 1962a, p. 158 [Burma]. 
Type: Holotype d\ macropterous; Kambaiti, Burma; Stockholm Mus. 
Distribution: Burma. 
Hosi Pi wt: Unrecorded. 



LACEBUGS OF THE WORLD 171 

*Cysteochila malayana Drake 

Cysteochila malayana Drake 1956h, p. 19 [Malaya]. 
Type: Holotype cf, macropterous; Salangor, Malaya; British Mus. 
Distribution: Federation of Malaya; Laut Island. 
Host Plant: Unrecorded. 

Cysteochila maynei Schouteden 

Cysteochila maynei Schouteden 1916b, p. 296 [Congo]. 
Type: Sex unknown; Congo da Lemba, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Cysteochila mokuensis Schouteden 

Cysteochila mokuensis Schouteden 1953b, p. 120 [Congo]. 
Type: Sex unknown; Moku, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 
*Cysteochila monstrosa (Scott) 

Monanthia monstrosa Scott 1874, pp. 291, 441 [Japan]. 

Physatochila monstrosa: Lethierry and Severin 1896, p. 21. — Oshanin 

1908, p. 449. 
Cysteochila monostrosa [sic]: Drake and Ruhoff 1961c, p. 145. 
Type: Sex unknown; Japan; apparently lost. 
Distribution: Japan; China (Fukien). 
Host Plant: Unrecorded. 
*Cysteochila munda Drake 

Cysteochila munda Drake 1954b, p. 5 [Cape Province]. 
Type: Holotype cf 1 , macropterous; Pondoland, Cape Province, South 
Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
*Cysteochila natalensis (Stal) 

Physatocheila natalensis Stal 1855a, p. 38 [Natal]. 

Monanthia {Physatochila) natalensis: Stal 1865, p. 28 [Caffraria]. 

Monanthia {Compseuta) natalensis: Stal 1873, p. 133. 

Monanthia natalensis: Walker 1873a, p. 195. — Distant 1902a, p. 242, pi. 

15, fig. 10. 
Physatochila sordidula Horvath 1910, p. 66 [Tanganyika]. 
Cysteochila humeralis (not Distant) Schouteden 1953b, p. 122 [Congo]. 
Cysteochila gibbosula Schouteden 1955b, p. 168. 
Cysteochila natalensis: Drake 1956b, p. 109. 
Type: Sex unknown; Natal, South Africa; Stockholm Mus. 
Distribution: South Africa (Natal); Congo; Tanganyika; Uganda; 
Kenya. 
Host Plant: Unrecorded. 



172 U.Q. NATIONAL MUSEUM BULLETIN 243 

♦Cysteochila oativa Drake 

clula nativa Drake I960, p. 147, fig. 7 [New Guinea]. 

Type: Holotype 9, macropterous; Finchhafen, Northeasl New Guinea; 

Drake Coll. I I 5NM). 

Distribi noN: .\< w Guinea (Northeast) 
Host Plan i i Unrecorded. 
*Cysteochil.i aervosana Drake 

eochila nervosum Dr. ike L956b, p. 11" [Natal; ('ape Province]. 
Type: Holotype 9; Weenan, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal; Cape Province). 
Hosi I'i an i : Unrecorded. 
Cysteochila nigriceps (Signoret) 

Monanthia nigriceps Signoret 1861, p. 955 [Malagasy]. — Stal 1873, p. 134. 
Monanthia {Physatochila) nigriceps: Stal 1865, p. 29. 

eochila nigriceps: Lcthicrry and Severin 1896, p. 22. — Drake and 
Runoff 1960c, p. 31. 
I \ ii : Sex unknown; Madagascar; Vienna Mus. 
Distribution: Malagasy Republic. 
Most Plant: Unrecorded. 
♦Cysteochila nitens (Drake and Poor) 

Oncophysa nilentis Drake and Poor 1937a, p. 1 1 [Luzon]. 
Cysteochila nitens: Drake and RuhofT 1961c, p. 145. 
Type: Holotype 9, macropterous; Mount Banahao, Luzon, Philippine 
Islands; Drake Coll. (USNM). 
I )i>iribution: Philippine Islands (Luzon). 
1 Ii >m Pi w i : Unrecorded. 
♦Cysteochila orta Drake and Maa 

Cysteochila orta Drake and Maa 1953, p. 93 [China]. 
Type: Holotype 9, macropterous; Foochow, Fukien, China; Drake 
Coll. (USNM). 

ribution: China. 
HOST I'i \\t: Unrecorded. 
::: Cysteochila oscitans Drake and Maa (emendation) 

( . 'rochila oscitantis Drake and Maa 1953, p. 93 [China]. 
Type: Holotype o 71 , macropterous; Shaowu I Lien, Fukien, China; 
Drake Coll. (USNM). 
I )i- 1 in ' i [( in: China. 
Host Pi an i : Unrecorded. 
♦Cysteochila otaviana Drake 

leochila otaviana Drake l ( )54h, p. 6 [South-West Africa]. 
Type: Holotype o", macropterous; Otavi, South-West Africa; Drake 
Coll. (USNM). 
Distribution: South-West Africa. 

H<»i Pi \m : I 'n recorded. 



LACEBUGS OF THE WORLD 173 

*Cysteochila pallens Horvath 

Cysteochila pallens Horvath 1915, p. 331 [Eritrea]. — Silvestri 1915, p. 
241 [Oleachrysophylla]; 1934, p. 259, fig. 223. 
Type: Sex unknown; Nefasit, Eritrea; Hungarian Mus. 
Distribution: Eritrea. 
Host Plant: Olea chrysophylla. 

*Cysteochila phae Drake and Ruhoff 

Cysteochila phae Drake and Ruhoff 1961c, p. 148, fig. 3 [Uganda; 
French Somaliland]. 
Type: Holotype cf, macropterous; Dijibouti, French Somaliland; 
Paris Mus. 
Distribution: Uganda; French Somaliland. 
Host Plant: Unrecorded. 

Cysteochila phiala Drake 

Cysteochila phiala Drake 1957f, p. 126 [Malagasy]. 
Type: Holotype o 71 ; Tananarive-Tsimbazaza, Madagascar; Madagascar 
Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 
*Cysteochila picta (Distant) 

Bredenbachius pictus Distant 1903a, p. 50 [Burma]; 1903b, p. 140, fig. 

103. 
Cysteochila pictus: Drake 1930e, p. 168 [Luzon]. 
Cysteochila picta: Drake and Poor 1937a, p. 7 [Mindanao]. 
Type: Sex unknown; Myitta, Tenasserim, Burma; British Mus. 
Distribution: Burma; Philippine Islands (Luzon; Mindanao). 
Host Plant: Unrecorded. 
*Cysteochila poecilia Drake and Ruhoff Plate 9 

Cysteochila poecilia Drake and Ruhoff 1961c, p. 146, fig. 2 [Chad]. 
Type: Holotype 9; Sahara, south of Tibesti, Chad; Paris Mus. 
Distribution: Republic of Chad. 
Host Plant: Unrecorded. 
*Cysteochila ponda Drake 

Cysteochila ponda Drake 1937b, p. 593, fig. 18 [China]. 
Type: Holotype 9, macropterous; Hong Shan, Kiangsi, China; Drake 
Coll. (USNM). 
Distribution: China. 
Host Plant: Unrecorded. 
Cysteochila propria Drake 

Cysteochila propria Drake 1958c, p. 318 [Malagasy]. 
Type: Holotype 9; Perinet, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 



17 1 U.8. NATIONAL MUSEUM BULLETIN 243 

♦Cysteochila rhodesiae 1 ):. 

leochilarhoc Drake L954b,p.7 I Rhodesia]. 
tchila rhodtsia [sic]: Drake 1958a, p. 103 [Angola]. 
Type: Holotype ipterous; Southern Rhodesia; Hungarian Mus. 

Dbtpjbi now Southern Rhodesia; Angola. 
Ili l'i w i : (Jnrecoi ded. 
Note: Tab n in light trap [Drake 1958a]. 

♦Cysteochila rubida Horv&th 

sltochila rubida Horvath 1924, p. 193 [Malagasy]. 
Type: Sex unknown; Sinus Antongil, Madagascar; Hungarian Mus. 
Distribution: Malagasy Republic. 

Host Plant: Unrecorded. 

♦Cysteochila rustica Drake 

Cysteochila rustica Drake 1957e, p. 400 [Reunion]. 

Type: Holotype d\ macropterous; Saint-Denis, La Montagne, Re- 
union; Madagascar Sci. Inst. 

Distribution: Mascarene Islands (Reunion). 

Host Plant: Unrecorded. 

Cysteochila seydeli Schouteden 

Cysteochila seydeli Schouteden 1953b, p. 119 [Congo]. 
Type: Sex unknown; Elizabethville, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 
*Cysteochila sordida (Stal) 

Monanthia (Physatocheila) sordida Stal 1859, p. 259 [Cape Province]; 

1865, p. 29. 
Monanthia sordida: Walker 1873a, p. 195. 

Cysteochila sordida: Stal 1873, p. 129.— Distant 1902a, p. 242.— Silvestri 
1915, p. 241 [Olea verrucosa]; 1934a, p. 259.— Horvath 1925a, p. 3. 
Type: Sex unknown; Cape of Good Hope, South Africa; Stockholm 
Mus. 

1 )istribution: South Africa (Cape Province). 
I 1 1 >sT Plant: Olea verrucosa. 

*Cysteochila suensoni Drake 

Cysteochila suensoni Drake 1942a, p. 5 [China]. 
Type: Holotype 9, macropterous; Tien Sun Shan, China; Drake Coll. 
I SNM). 

Distribution: China. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 175 

Cysteochila suspecta Schouteden 

Cysteochila ? suspecta Schouteden 1953b, p. 121 [Congo]. 
Type: Sex unknown; Uvira, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Cysteochila syscena Drake 

Cysteochila syscena Drake 1960, p. 350 [New Britain]. 
Type: Holotype 9; Ti, Nakanai Mts., New Britain; Bishop Mus. 
Distribution: New Britain Island, Bismarck Archipelago. 
Host Plant: Unrecorded. 

*Cysteochila taprobanes Kirkaldy 

Cysteochila taphrobanes Kirkaldy 1908c, p. 12 [Ceylon]. — Distant 1910a, 
p. 126. 
Type: Galle, Ceylon; sex and deposition of type unknown. 
Distribution: Ceylon; India. 
Host Plant: Unrecorded. 

*Cysteochila tarda Drake 

Cysteochila tarda Drake 1954a, p. 235 [Senegal; Sudan]. 
Type: Holotype d"; Bambey, Senegal; Inst. Fr. Afr. N. 
Distribution: Senegal; Sudan. 
Host Plant: "Kittirbush." 

*Cysteochila terminalis Drake 

Cysteochila terminalis Drake 1948a, p. 6 [India]. 
Type: Holotype 9, macropterous; Samsingh, Kalimpong, India; Drake 
Coll. (USNM). 
Distribution: India. 
Host Plant: Unrecorded. 

*Cysteochila tingoides (Motschulsky) 

Monanthia ? tingoides Motschulsky 1863, p. 92 [Ceylon]. — Kirby 1891, 

p. 109. 
Cysteochila tingoides: Stal 1873, p. 129.— Distant 1903b, p. 139— Ber- 
groth 1921, p. 103.— Drake 1948a, p. 4 [India]. 
Type: Sex unknown; Montagnes de Nura-Ellia, Ceylon; Leningrad Inst. 
Distribution: Ceylon; India. 
Host Plant: Unrecorded. 

*Cysteochila tombeuri Schouteden 

Cysteochila tombeuri Schouteden 1923, p. 103 [Tanganyika]. — Drake 
1958a, p. 103 [Angola; So. Rhodesia; Mozambique; Kenya]. 
Type: Sex unknown; Kigoma, Tanganyika; Cent. Afr. Mus. 
Distribution: Tanganyika; Kenya; Angola; Mozambique; Southern 
Rhodesia. 
Host Plant: Unrecorded. 



176 OA, NATIONAL MUSEUM BULLETIN 243 

^Cysteochila tricolor (HackeJ 

rcilliger tricolm Hacker 1929, p. 332, text fig. 1 [Queensland], 

■ leochila fran&ni Drake 1942a, p. 4. 

teochila hicolor [sic]: Drake 1942a, p. 1. 

teochila tricolor: Drake and Maa 1953, |>. 92. 
I -. ri : Sex unknown; Mount Glorious, Queensland, Australia; Queens- 
land Mus. 
Distribution: Australia (Queensland). 
II* is'j I'i \n i : I Inrecorded. 

*Cysteochila tzitzikamana Drake 

leochila tzitzikamana Drake 1956f, p. 422 [Cape Province]. 

Type: Holotype d 71 ; Tzitzikama Forest, Stormrivierpiek, Cape Province, 
South Africa; Lund Zool. Inst. 
Distribution: South Africa (Cape Province); Congo. 
Host Plant: Unrecorded. 

*Cysteochila undosa Drake 

Cysteochila undosa Drake 1942a, p. 6 [Burma]. 
Type: Holotype d\ macropterous; Kambaiti, Burma; Drake Coll. 
(USNM). 
Distribution: Burma. 
Host Plant: Unrecorded. 

*Cysteochila vadosa Drake 

Cysteochila vadosa Drake 1956h, p. 18 [Natal]. 
Type: Holotype cf, macropterous; Umkomaas, Natal, South Africa; 
Drake Coll. (USNM). 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

^Cysteochila visenda Drake and Poor 

Cysteochila visenda Drake and Poor 1937a, p. 8 [Ncgros]. 
I ri : Holotype d\ macropterous; Cucrnos Mountains, Oriental Ncgros 
P ince, Negros, Philippine Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Negros). 
Host Plant: Unrecorded. 

^Cysteochila vitilevuana Drake and Poor 

anihia natalensis (not Stal): Kirkaldy 1908b, p. 366 [Fiji]. 
Cysteochila vitilevuana Drake and Poor 1943, p. 193. — Drake and Hurd 
1945a, p. 287, fig. 1. 
Type: Holotype o 1 ; Nandarivatu, Yiti Levu, Fiji; Bishop Mus. 
Distribution: Fiji Islands (Viti Levu). 

1 1< >sr Plan i : I 'nrecorded. 



LACEBUGS OF THE WORLD 177 

*Cysteochila vota Drake 

Cysteochila vota Drake 1948a, p. 7 [Japan]. — Takeya 1951a, p. 21. 
Type: Holotype 9, macropterous; Moji, Japan; Drake Coll. (USNM). 
Distribution: Japan. 
Host Plant: Unrecorded. 

*Cysteochila vota var. fida Drake and Maa 

Cysteochila vota var.jida Drake and Maa 1953, p. 92 [China]. 

Type: Holotype cf , macropterous; Shaowu City, Fukien, China; Drake 
Coll. (USNM). 

Distribution: China. 

Host Plant: Unrecorded. 

*Cysteochila wechinai Drake 

Cysteochila chiniana Drake 1954b, p. 5 (not 1942) [Luzon]. 

Cysteochila wechinai Drake 195 4f , p. 5. 
Type: Holotype cf ; Trinidad, Luzon, Philippine Islands; British Mus. 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

*Cysteochila yungana Drake and Maa 

Cysteochila yungana Drake and Maa 1955, p. 4 [China]. 

Type: Holotype 9, macropterous; Yungan City, Fukien, China; Drake 
Coll. (USNM). 

Distribution: China. 

Host Plant: Unrecorded. 

Cysteochila zavattarii Mancini 

Cysteochila zavattarii Mancini 1953a, p. 184 [Ethiopia]. 
Type: Holotype 9; Mega, Ethiopia; Mancini Coll. 
Distribution: Ethiopia. 
Host Plant: Unrecorded. 

Genus DASYTINGIS Drake and Poor 

Dasytingis Drake and Poor 1936b, p. 145. — Drake and Ruhoff 1960a, 
p. 49. 
Type Species: Dasytingis rudis Drake and Poor. 

*Dasytingis rudis Drake and Poor 

Dasytingis rudis Drake and Poor 1936b, p. 145, fig. 1 [India]. — Drake 

and Ruhoff 1962c, p. 133. 
Tingis bengalana Drake 1956h, p. 20. 
Type: Holotype cf, macropterous; Hoshangabad, Rahatgaon, India; 
Drake Coll. (USNM). 
Distribution: India. 
Host Plant: Vitex ruganda. 



178 0.8. NATIONAL MUSEUM BULLETIN 243 

*Dasytingis semota Drake and Lutz 

Dasytingis semota Drake and Lutz 1953, p. 104 [India]. 
I \ n : I [olotypc o\ macroptcrous; Karikal, India; I trake ( loll. (USNM). 
Distribution: India. 
Host Plant: Unrecorded. 

Genus DICHOCYSTA Champion 

Champion 1898a, p. 33. — Banks 1910, p. 56. — Van Duzce 
1916, p. 26; r»ri), p. 221. Blatchley 1926, p. 485.— Hurd 1946, 
p. 453. Drake and Ruhoff 1960a, p. 50. 
\ \ pe Species: Dichocysta pictipes Champion. 

*Dichocysta pictipes Champion 

Dichocysta pictipes Champion 1898a, p. 34, pi. 3, figs. 1, la, b, 2 [Guate- 
mala; Panama]. — Heidemann 1899a, p. 339 [Ariz.]. — Barber 1910, 
p. 38.— Drake 1918d, p. 88 [Fla.]; 1928c, p. 2 [Honduras; cacao]; 
1938a, p. 70 [Mexico].— Blatchley 1926, p. 485 [Costa Rica]. 
Type: Sex unknown: David, Chiriqui, Panama; British Mus. 
Distribution: U.S. (Fla.; Tex.; Ariz.); Mexico; Guatemala; Honduras; 
Costa Rica; Panama. 
Host Plant: Cacao. 

Genus DICONOCORIS Mayr 

Diconocoris Mayr 1865, p. 442— Stal 1873, p. 134.— Lethierry and 
Severin 1896, p. 25.— Drake 1937a, p. 386.— Drake and Ruhoff 
1960a, p. 50. 

Diplogomphtts Horvath 1906c, p. 296.— Drake and Poor 1937a, p. 10 — 
Drake and Ruhoff 1960a, p. 53. 
Type Species: Diconocoris javanus Mayr. 

*Diconocoris capusi (Horvath) Plates 30, 31 

Diplogomphus capusi Horvath 1906c, p. 296, fig. [Viet-Nam]. — Drake 

and Poor 1937a, p. 10. 
Diconocoris capusi: Drake 1937a, p. 386. — Drake and Davis 1960, fig. 54. 
Type: Indochina; sex and deposition of type unknown. 
Distribution: Viet-Nam. 
Host Plant: Unrecorded. 

^Diconocoris distant! Drake 

Elasmognathus greem (not Kirby): Distant 1903b, p. 142, fig. 105 [Ceylon; 
Piper nigrum]. 
Diconocoris distanti Drake 1954f, p. 4. 
Type: Holotype d 1 ; Peradeniya, Ceylon; British Mus. 
Distribution: Ceylon. 
Host Plant: Piper nigrum. 



LACEBUGS OF THE WORLD 179 

Diconocoris greeni (Kirby) 

Elasmognathus greeni Kirby 1891, p. 109, pi. 4, fig. 5. [Ceylon]. — 

Kirkaldy 1902, p. 297. 
Elasmognathus pallida Kirby 1891, p. 110. 
Diplogomphus greeni: Drake and Poor 1937a, p. 10. 
Diconocoris greeni: Drake 1937a, p. 386. 
Type: Sex unknown; Ceylon; British Mus. 
Distribution: Ceylon. 
Host Plant: Wild pepper. 

*Diconocoris hewetti (Distant) 

Elasmognathus hewetti Distant 1908b, p. 127, pi. 7, figs. 2, 2a [Borneo]. 
Elasmognathus picturatus Distant 1909c, p. 165, pi. 10, fig. 4, 4a. 
Elasmognathus hewitti [sic]: van der Vecht 1931, pp. 824, 828 [Sumatra; 

pepper]; 1935, pp. 484, 492, pi. 1 [Bangka Islands]. 
Diplogomphus hewitti [sic]: Drake and Poor 1937a, p. 10. — Blote 1945, 

p. 89. 
Cysteochila picturatus: Drake 1937a, p. 386. 

Diconocoris hewetti: Drake 1937a, p. 386. — -Drake and Ruhoff 1960c, p. 
36. 
Type: Sex unknown; Kuching, Borneo; British Mus. 
Distribution: Greater Sunda Islands (Borneo; Sumatra); Bangka 
Islands. 
Host Plant: Pepper. 
Note: Biology [van der Vecht 1935]. 

*Diconocoris inusitatus (Drake) 

Elasmognathus inusitatus Drake 1927d, p. 309 [Luzon]. 
Diplogomphus inusitatus: Drake and Poor 1937a, p. 10. 
Diconocoris inusitatus: Drake 1937a, p. 386. 
Type: Holotype cf, macropterous; Mount Maquiling, Luzon, Philip- 
pine Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

*Diconocoris javanus Mayr 

Diconocoris javanus Mayr 1865, p. 442 [Java]. — Stal 1873, p. 134. — 
Drake 1937a, p. 386, figs, la, b — Drake and Poor 1941, p. 160 
[Amboina]. 
Monanthia javana: Walker 1873a, p. 196. 
Type: Sex unknown; Java; Vienna Mus. 

Distribution: Greater Sunda Islands (Java); Moluccas (Amboina). 
Host Plant: Unrecorded. 



ISO l.S. NATIONAL MUSEUM BULLETIN 243 

Diconocoris nepalensis Distant) 

■nognuthus nepalensis Distant 1909a, p. 120 [Nepal]; 1910a, p. 122, 
fig. 63. 
Diplogompkus neftulensis: Bergroth 1911, p. 186. 
Diplogomphus napalensis [sic]: Drake and Poor 1937a, p. 10 [India]. 
D\ onocoris nepalensis: Drake 1937a, p. 386. 
Type: Sex unknown; Thamaspur, Nepal; Indian Mus. 
I hsi Rim mon: Nepal; India. 
Host Plant: Unrecorded. 

Genus DICROTINGIS Drake and Ruhoff 

Dicrotingis Drake and Ruhoff 1960c, p. 34. 
Type Species: Leptopharsa digitalis Drake. 
*Dicrotingis digitalis (Drake) Plate 26 

Leptopharsa digitalis Drake 1928d, p. 22 [Haiti; Phyllostylon rhamnoides]. 
Dicrotingis digitalis: Drake and Ruhoff 1960c, p. 35, fig. 4a, b. 
Type: Lectotype cf, macropterous; Crox des Missiones, Plaine Guide 
Sac, Haiti; Drake Coll. (USNM). 
Distribution: Haiti. 
Host Plant: Phyllostylon rhamnoides. 

Note: Holotype apparently lost. Paratype d", same locality and date 
as holotype, here designated as lectotype. 

Genus DICTYLA Stal 

Dictyla Stal 1874, p. 57.— Drake and Ruhoff 1960a, p. 50.— Stichel 
1960a, p. 336; 1960b, p. 400; 1960c, p. 139. 

Monanthia (of most authors, but not Le Peletier and Serville; see Drake 
and Ruhoff 1960a, pp. 45, 50).— Mayr 1858, p. 571.— Stal 1873, 
pp. 122, 133— Horvath 1874b, p. 432; 1906a, pp. 15, 97 (key).— 
d'Antessanty 1890, p. 32. — Lethierry and Severin 1896, p. 23. — 
Oshanin 1908, p. 450; 1912, p. 45.— Banks 1910, p. 57— Jensen- 
Haarup 1912, p. 155 (key).— Van Duzee 1917b, pp. 223, 819.— Drake 
1917c, pp. 49-52 (key); 1943a, p. 142.— Sahlberg 1920, p. 84 — 
Stichel 1926, pp. 105, 115 (key); 1935, p. 349; 1938a, p. 408.— de 
Seabra 1931, pp. 408, 434 (key).— Lindberg 1932a, p. 44— Borner 
1935, pp. 74, 78 (key).— Scholte 1935, p. 91 (key).— Drake and 
Poor 1936c, p. 388.— Guide 1938, pp. 245, 311 (key).— Monte 1939b, 
p. 76; 1941e, p. 123.— China 1943, p. 248.— Hoberlandt 1943, 
p. 124.— Blote 1945, p. 90.— Hurd 1946, p. 450.— Kiritshenko 
1951, pp. 242, 253 (key).— Singer 1952, p. 54. 
Type Species: Monanthia platyoma Ficbcr. 



LACEBUGS OF THE WORLD 181 

Dictyla abyssinica (Drake) 

Monanthia abyssinica Drake 1954g, p. 658, fig. 1 [Ethiopia; Cordia 
abyssinica]. 

Dictyla abyssinica: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9; Doukam, Ethiopia; British Mus. 
Distribution: Ethiopia. 
Host Plant: Cordia abyssinica. 

*Dictyla aima Drake 

Dictyla aima Drake 1961b, p. 107, pi. 3 [Queensland]. 
Type: Holotype o 71 ; Queensland, Australia; Austr. Mus. 
Distribution: Australia (Queensland). 
Host Plant: Collected on "cuolahah box." 

*Dictyla ainsliei (Drake and Poor) 

Monanthia ainsliei Drake and Poor 1938a, p. 28, fig. 1 [Guatemala]. 
Dictyla ainsliei: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9, macropterous; Concepcion, Guatemala; Drake Coll. 
(USNM). 
Distribution: Guatemala. 
Host Plant: Unrecorded. 

*Dictyla alia Drake and Cobben 

Dictyla alia Drake and Cobben 1960, pp. 67, 69, figs. 74a, 75a, 76a, 77 
[Curacao; Aruba; Bonaire; Cordia alba]. 
Type: Holotype d", macropterous; St. Martha, Curacao, Netherlands 
Antilles; Drake Coll. (USNM). 
Distribution: Netherlands Antilles (Curagao; Aruba; Bonaire). 
Host Plant: Cordia alba. 

*Dictyla amitina (Horvath) 

Monanthia amitina Horvath 1925a, p. 13 [Queensland]. — Hacker 1927 

p. 25, fig. 18; 1928a, p. 181. 
Dictyla amitina: Drake and Ruhoff 1960a, p. 51. — Drake 1961b, p. 107 
pi. 3. 
Type: Sex unknown; Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Dictyla aurigana (Drake) 

Monanthia aurigana Drake 1954b, p. 2 [Natal]. 

Dictyla aurigana: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype o* ; Shepstone, Natal, South Africa; British Mus. 
Distribution : South Africa (Natal) . 
Host Plant: Unrecorded. 



182 0.8. NATIONAL MUSEUM BULLETIN 243 

*Dictyla aurigana subsp. discoris (Drake) 

Monanthia aurigana Bubsp. dis orii Drake 1954b, p. 2 [Tanganyika]. 

Dictyla aurigana subsp. discoris: Drake and Ruhoff 1960a, p. 51. 
\w\. Holotype ?, macropterous; Lindi, Tanganyika; Drake Coll. 
(USNM 
Distribution : Tanganyika. 
Host Plant: Unrecorded. 
*Dictyla balli (Drake) 

Monanthia balli Drake 1922c, p. 355 [Haiti]. — Drake and Bruner 1924a, 

p. 144. 
Dictyla balli: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9, macropterous; Port au Prince, Haiti; Drake Coll. 
I s\M, 

Distribution: Haiti; Bahama Islands (Cat and Harbor Islands). 
Host Plant: Unrecorded. 
*Dictyla berryi (Drake) 

Monanthia berryi Drake 1943a, pp. 141, 142. — Drake and Hamblcton 

1944b, p. 121 [Peru]; 1945, p. 356 [Ecuador]. 
Dictyla berryi: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype e? , macropterous; Chanchaqui, Peru; USNM. 
Distribution: Peru; Ecuador. 
Host Plant: Unrecorded. 

Note: Drake and Hambleton 1944b corrected the type locality to 
"Chanchaqui, Peru" not "Montevideo, Uruguay" as published in the 
original description. 
♦Dictyla burgeoni (Schouteden) 

Monanthia burgeoni Schouteden 1923, p. 107 [Congo]. 
Type: Sex unknown; Bas Uele, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Note: Further study reveals this to be a valid species, not a synonym of 
nodipennis as designated in Drake 1956a, p. 4. 
Dictyla burgeoni vur. satanas (Schouteden) 

Monanthia burgeoni var. satanas Schouteden 1923, p. 107 [Congo]. 
Type: Sex unknown; Bas Uele, Belgian Congo; Cent. Afr. Mus. 
DisTRiBiTu in: ( longo. 
Host Plant: Unrecorded. 
*Dictyla cheriani (Drake) 

Monanthia cheriani Drake 1936a, p. 145 [India; Corida]. 
Dictyla cheriani: Drake and Ruhoff 1960a, p. 51; 1961b, p. 131, fig. 7. 
Type: Holotype d\ macropterous; Coimbatorc, India; Drake Coll. 
(USNM). 
Distribution: India. 
Host Plant: Corida sp. 



LACEBUGS OF THE WORLD 183 

*Dictyla c-nigrum (Champion) 

Monanthia c-nigrum Champion 1898a, p. 47, pi. 3, fig. 25 [Mexico; 
Guatemala; Nicaragua]. — Van Duzee 1907, p. 23 [Jamaica]. — Drake 
1922c, p. 355 [Haiti]; 1929, p. 35 [Costa Rica]; 1943a, p. 142 
[Brazil].— Barber 1939, p. 372.— Monte 1941d, p. 100.— Drake and 
Hambleton 1945, p. 356. 
Dictyla c-nigrum: Drake and Runoff 1960a, p. 51. 
Type: Sex unknown; Atoyac, Mexico; British Mus. 
Distribution: Mexico; Guatemala; Nicaragua; Costa Rica; Haiti; Ja- 
maica; Brazil; Guadeloupe Island. 
Host Plant: Unrecorded. 

Dictyla collarti (Schouteden) 

Monanthia collarti Schouteden 1953e, p. 196 [Congo]. 

Dictyla collarti: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Mayumbe, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Dictyla coloradensis (Drake) 

Monanthia coloradensis Drake 1917c, pp. 50, 51, fig. lb [Colo.]. 

Dictyla coloradensis: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9, macropterous; Colorado, U.S.; Drake Coll. (USNM). 
Distribution: U.S. (Colo.; Tex.; Oreg.). 
Host Plant: Unrecorded. 

*Dictyla comes (Drake) 

Monanthia comes Drake 1948a, p. 2 [China]; 1954g, p. 658 [Ethiopia; 
Cape Province; Orange Free State; Natal; Uganda]; 1956a, p. 4 
[Congo; India; Arabia; Egypt; Morocco; Senegal; Kenya; Nigeria; 
West Africa]. 
Dictyla comes: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype o 71 , macropterous; Kumming, Tausan-fu, Yunnan 
Province, China; Drake Coll. (USNM). 

Distribution: China; India; Arabia; Morocco; Egypt; Ethiopia; Congo; 
Senegal; Kenya; Uganda; Nigeria; South Africa (Orange Free State; Cape 
Province; Natal); West Africa. 
Host Plant: Unrecorded. 

*Dictyla convergens (Herrich-Schaeffer) 

Tingis convergens Herrich-Schaeffer 1835, p. 59. 

Monanthia convergens: Burmeister 1835, p. 261 [Myosotis palustris]. — 

Herrich-Schaeffer 1837, p. 15, pi. 114, p. 361; 1838, p. 58. 
Monanthia (Physatocheila) humuli (not Fabricius): Fieber 1844, p. 84, 

pi. 7, figs. 17-18 [Austria; Germany; Czechoslovakia; England]. — 

Herrich-Schaeffer 1850, p. 153.— Flor 1860, p. 355 [Estonia; Latvia; 

Sweden; France]. 



1st [J.S. NATIONAL MUSEUM BULLETIN 243 

Monanthia humuli: Scholz 1847, p. 120. Fieber 1861, p. 125. — Douglas 
and Scott 1865, p. 244. — Frcy-Gessner 1865, p. 232 [Switzerland]. — 
Jakovlev 1869, p. ill [Russia], Garbiglictti 1869, p. 273 [Italy].— 
Gredler 1870, p. 75.— Thomson 1871, p. 398.— Saunders 1875, p. 
24'); 1892, p. 136, pi. 13, fig. 1 [Scotland; Wales; Myosotis].— Reiber 
and Puton 1876, p. 69. Vollenhoven 1878, p. 274, pi. 9, fig. 6 
[Netherlands].- Puton 18~'>c, |». 122; 1886a, p. 12.— Chicotc 1880, 
p. 189 [Spain].— Rcutcr 1882b, p. 163 [Finland; Myosoiis lingu- 
lata}.- Ferrari 1884, p. 475 [Tunisia].— Hiieber 1893, p. 352 — 
Horvath 1906a, pp. 99, 102— Muzik 1907, p. 63.— Oshanin 1908, 
p. 452 [Belgium; Hungary; Rumania; Portugal]. — Jensen-Haarup 
1912, pp. 155, 157, fig. 98c [Denmark].— Schumacher 1914, p. 259 — 
Reichensperger 1920, p. 62.— Butler 1923, p. 215.— Stichel 1926, 
p. 116, fig. 306.— Ross and Hedickc 1927, p. 188.— de Seabra 1931, 
p. 436, fig. 507. — Scholte 1935, p. 91, fig. 25 [Myosotis cespitosa). — 
Guide 1938, p. 315, fig.— Cobben 1948, p. 82; 1958a, p. 14, fig. 17 — 
Kiritshcnko 1951, p. 253.— Lcston 1954, p. 101, fig. 5— Stusak 
1957b, pp. 138, 140, figs. 8, 10b, 12b.— Southwood and Lesion 
1959, p. 151, pi. 21, fig. 12; pi. 25, fig. 2. 

Dxclyla humuli: Drake and Ruhoff 1960a, p. 51.— Stichel 1960a, p. 341, 
fig. 175 [Poland]. 

Dutyla convergens: Drake and Ruhoff 1960b, p. 74, pi. 21. 

Type: Unknown. 

Distribution: Germany; France; England; Scotland; Wales; Switzerland; 
Czechoslovakia; Austria; Italy; Poland; Sweden; Denmark; Finland; 
Hungary; Rumania; Belgium; Portugal; Spain; Netherlands; U.S.S.R. 
(Estonia; Latvia); Tunisia. 

Host Plants: Myosotis cespitosa; Myosotis lingulata; Myosotis palustris: 
Myosotis sp. 

Note: Study of immature stages [Leston; Stusak]. Described by 
Herrich-Schaeffer but wrongly accredited by him and others to Klug 
[Hrrrich-Schaeffer 1837, p. 15]. 

:;: Dictyla echii (Schrank) 

Cimex clavkornis (not Linnaeus): Schrank 1781, p. 265. — Villers 1789, 

p. 485, pi. 3, fig. 
Cimex echii Schrank 17S2, p. 276. 
Acanthia aim: Schrank 1801, p. 65— Wolff 1804, p. 130, pi. 13, fig. 

124 [Echium vulgar e\. 
Tingis humuli (not Fabricius): Fallen 1807, p. 36; 1829, p. 144.— Brull6 

1835, p. 340, pi. 26, fig. 1 [Greece].— Blanchard 1840, p. 113. 
Tingis echii: Herrich-Schaeflfer 1835, p. 59.— Schilling 1838, p. 105. 
Mnnanthia humuli: Burmeister 1835, p. 261. — Costa 1847a, p. 22; 

1847c, p. 162. 



LACEBUGS OF THE WORLD 185 

Monanthia echii: Herrich-Schaeffer 1837, p. 14, pi. 114, fig. 360, 124c; 
1838, p. 58. — Jakovlev 1880b, p. 110 [Echium rubrum; Echium itali- 
cum\— Reuter 1891a, p. 26 [Crete].— Horvath 1906a, pp. 99, 103 
[Armenia; Turkestan; Siberia; Turkey; Algeria]; 1918a, p. 334. — 
Oshanin 1908, p. 453 [Belgium; Rumania; Yugoslavia; Iran]. — 
Jensen-Haarup 1912, pp. 155, 157 [Denmark]. — Schumacher 1914, 
p. 259.— Moroder Sala 1920, p. 13.— Stichel 1926, p. 116; 1938a, 
p. 408 [Albania; Israel; Poland; Norway]. — Ross and Hedicke 1927, 
pp. 86, 136, 181 [Lycopsis arvensis]. — de Seabra 1931, p. 437, figs. 506 
(2-5), 509.— Lindberg 1932a, p. 44 [Morocco]; 1948, p. 60 [Bul- 
garia].— Scholte 1935, p. 91, fig. 24.— Vidal 1937, p. 199 [Echium 
creticum]. — Blote 1945, p. 90 [Egypt; Tunisia; Corsica]. — Cobben 
1948, p. 82; 1958a, p. 14, fig. 18.— Kiritshenko 1951, p. 254.— Novak 
and Wagner 1951, p. 71 [Echium altissimum; Echium pustulatum]. — 
Mancini 1953a, p. 186.— Bator 1953, p. 327, pi. 4, fig. 6— Hober- 
landt 1955, p. 95 [Cyprus].— Stusak 1957b, pp. 138, 140, fig. 7; 1958, 
p. 369, fig. 16. 
Monanthia clavicornis: Amyot and Serville 1843, p. 298 [France; Sicily]. 
Monanthia (Physatocheila) wolffii Fieber 1844, p. 86, pi. 7, figs. 22-24 
[Germany; Italy; Hungary; Sweden]. — Herrich-Schaeffer 1850, p. 
153. — Flor 1860, p. 352 [Estonia; Latvia; Caucasus; Symphytum; 
Pulmonaria]. 
Monanthia (Tropidocheila) wolffii: Sahlberg 1848, p. 133 [Finland]. 
Monanthia wolffii: Mayr 1858, p. 571 [Austria; Czechoslovakia; Cyno- 
glossum].— Fieher 1861, p. 125.— Frey-Gessner 1865, p. 232 [Switzer- 
land].— Jakovlev 1869, p. Ill; 1880b, p. 109.— Gredler 1870, p. 75 
[Anchusa officinalis].— Stal 1874, p. 59.— Ferrari 1874, p. 171; 1878, 
p. 66.— Vollenhoven 1878, p. 275, pi. 9, fig. 7 [Netherlands].— Puton 
1879c, p. 120; 1881, p. 123 [Syria].— Bolivar and Chicote 1879, p. 
166 [Spain; Portugal].— Reuter 1882b, p. 120.— d'Antessanty 1890, 
p. 33 [Cynoglossum officinale]. — Hiieber 1893, p. 349. 
Dictyla echii: Drake and Ruhoff 1960a, p. 51.— Stichel 1960a, p. 342, 
fig. 178 [Poland; Sardinia; Anchusa arvensis]. 
Type: Unknown. 

Distribution: Netherlands; France; Belgium; Spain; Portugal; 
Germany; Austria; Switzerland; Czechoslovakia; Hungary; Italy; Sicily; 
Sardinia; Corsica; Poland; Denmark; Sweden; Finland; Norway; Rumania; 
Yugoslavia; Albania; Bulgaria; Greece; Crete; Cyprus; Turkey; U.S.S.R. 
(Latvia; Estonia; Siberia; Turkestan; Transcaucasus; Caucasus; Armenia); 
Syria; Israel; Iran; Morocco; Algeria; Egypt; Tunisia. 

Host Plants: Anchusa arvensis; Anchusa officinalis; Cynoglossum officinale; 
Cynoglossum sp.; Echium altissimum; Echium creticum; Echium italicum; Echium 



681-552—64 13 



1SG U.S. NATIONAL MUSEUM BULLETIN 243 

pustulatum; Echitim ruin urn; lu hium vulgare; Echium sp.; I.vcopus arvensis; 
Pulmonaii'i >(>.; Symphytum sp. 
Noil: Study of immature stages [Stusak 1957b]. 

Dictyla cchii var. nigricans (Hoberlandt) 

Monanthia ecliii var. nigricans Hoberlandt 1943, pp. 116, 117, 119 

[( izechoslovakia]. 

hu var. nigricans: Drake- and Runoff 1960a, p. 51. — SdcheJ 
1960a, p. 342. 
Type: Holotype o 71 ; Hodonm, Moravia, Czechoslovakia; Prague Mus. 
Distribution : Czechoslovakia. 
Host Plant: Unrecorded. 

:;: Dictyla echii var. rufina (Seidenstiicker) 

Monanthia cchii subvar. rufina Seidenstucker 1954, pp. 230, 235 [Tur- 

key]. Hoberlandt 1955, p. 95 [Syria]. 
Dictyla cchii var. rufina: Drake and Ruhoff 1960a, p. 51. — Sti 
1960a, p. 342. 
Type: Turkey; sex and deposition of type unknown. 
Distribution: Turkey; Syria. 
Host Plant: Unrecorded. 

*Dictyla ehrethiae (Gibson) 

Monanthia ehrethiae Gibson {in Drake 1917c, pp. 49, 50) [Tex.; Ehrctia 

elliptica] . 
Monanthia ehrethia [sic]: Drake 1943a, p. 142 [Mexico]. 
Dictyla ehrethiae: Drake and RuhofT 1960a, p. 51. 
Type: Holotype 9, macropterous; Brownsville, Texas, U.S.; USNM. 
Distribution: U.S. (Tex.); Mexico. 
Host Plant: Ehretia elliplica. 

*Dictyla evidens (Drake) (New Status) 

Monanthia formosa Drake 1923a, p. 102 [Taiwan]. — Esaki 1926, p. 

163.— Takeya 1951a, p. 22; 1953b, p. 4, fig. 1 [Japan; Ehretia thyrsi- 

flora']. — Maa 1957, pp. 119, 131 [Ehretia taiwaniana; Ehretia dickstmu}. 

Monanthia evidens Drake 1927a, p. 53 [Luzon]; 1937a, p. 385. — Drake 

and Poor 1937b, p. 397 [Ehretia acuminata]. — Takeya 1951a, p. 23. 
Monanthia formosana [sic]: Drake and Maa 1 ''53, p. 90 [China]. 
Dictyla formosa: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 0"; Kankau, formosa; Horn Coll. 
DISTRIBUTION: Taiwan; China; Japan; Philippine Islands (Luzon). 
II ' 1 l'i wts: Ehretia acuminata; Ehrctia dicksonii; Ehretia taiwaniana; 
Ehretia thyrsiflora. 

*Dictyla femoralis (Stal) 

Monanthia (Compseuta) femoralis Stal 1873, p. 133 [Caffraria]. 



LACEBUGS OF THE WORLD 187 

Monanthia femoralis: Lethierry and Severin 1896, p. 23. — Distant 1902a, 
p. 243. 

Dictyla femoralis: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9; Caffraria, South Africa; Stockholm Mus. 
Distribution: South Africa (Zululand). 
Host Plant: Unrecorded. 

*Dictyla figurata (Drake) 

Monanthia figurata Drake 1922c, p. 354, pi. 39, fig. 12 [Brazil]; 1930a, 
p. 1; 1931b, p. 510.— Drake and Hambleton 1938b, p. 52 [Cordia]. 
Dictyla figurata: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype o 71 ; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Cordia sp. 

*Dictyla flavipes (Signoret) 

Monanthia flavipes Signoret 1861, p. 956 [Malagasy].— Stal 1873, p. 134. 
Dictyla flavipes: Drake and Ruhoff 1960a, p. 51; 1960c, p. 29; 1961b, 
p. 131, fig. 6. 
Type: Lectotype cf; Mayotte, Comoro Islands; Vienna Mus. 
Distribution: Malagasy Republic; Comoro Islands (Mayotte). 
Host Plant: Unrecorded. 

*Dictyla fulvescens (Kiritshenko) 

Monanthia fulvescens Kiritshenko 1952, p. 179 [Russia]. 

Dictyla fulvescens: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Tadzhikistan, Russia; Leningrad Inst. 
Distribution: U.S.S.R. 
Host Plant: Unrecorded. 

Dictyla gerardi (Schouteden) 

Monanthia gerardi Schouteden 1953e, p. 198 [Congo]. 
Dictyla gerardi: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Katompe, Katanga, Belgian Congo; Cent. Afr. 
Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Dictyla haitiensis (Drake and Poor) 

Monanthia haitiensis Drake and Poor 1938a, p. 28 [Haiti]. — Hurd 

1946, p. 451 [Puerto Rico]. 
Dictyla haitiensis: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9, macropterous; Port au Prince, Haiti; Drake Coll. 
(USNM). 
Distribution: Haiti; Puerto Rico. 
Host Plant: Cordia serrata. 



1S8 U.S. NATIONAL Ml SLUM BULLETIN 243 

*Dictyla humuli ( Fabric ius) 

A mth.ia humuli Fabrichu 1794, p. 77 [Germany; Hamulus]. 

Tingis humuli: Fabrichu 1803, p. 126. — Herrich-Schaeffer 1835, p. 

58. 
Tingis symphyti Vallot 1S2 ( >, p. 99 [Symphytum officinale], 

anthia costata (not Fabricius): Burmeister 1835, p. 261. — Herrich- 
Schaeffer 1837, p. 15, pi. 114, fig. 362. 

■ nithia (Physatocheila) vesiculijera Fiebcr 1844, p. 87, pi. 7, figs. 25- 
26 [Sicily; Italy; Austria].— Hen ich-Schaeffcr 1850, p. 153. 
Monanthia vesiculijera: Scholz 1847, p. 121. — Mayr 1858, p. 571 [Hun- 
ry]. — Fiebcr 1861, p. 126. — Frey-Gcssner 1865, p. 232 [Switzer- 
land].— Stal 1874, p. 59.— Jakovlev 1876b, p. 66; 1880b, p. 110 
[Caucasus].— Vollenhoven 1878, p. 276, pi. 22, fig. 9 [Netherlands]. 
— Sahlberg 1878, p. 21.— Puton 1879c, p. 123 [France].— Dubois 
1888, p. 122.— Rey 1893, p. 97.— Hueber 1893, p. 354 [Czecho- 
slovakia]. 
Monanthia symphyti: Ilorvath 1889, p. 326; 1906a, pp. 99, 101 [Yugo- 
slavia; Rumania; Siberia]. — Rcuter 1891b, p. 184. — Muzik 1907, 
p. 63. — Oshanin 1908, p. 452 [Algeria]. — Reichensperger 1920, p. 
62.— Stichel 1926, p. 116.— Scholte 1935, p. 92, fig. 26.— Guide 
1938, p. 317 [Symphytum].— BlSte 1945, p. 91 [Tunisia].— Cobben 
1948, p. 82; 1958, p. 14, fig. 16.— Mancini 1949b, p. 37.— Kiritshenko 
1951, p. 253.— Bator 1953, p. 327, pi. 4, fig. 3.— Gomez-Menor 
1955b, p. 249 [Spain].— Stusak and Stys 1959, pp. 178-187, 191, 
figs. 1, 2, 7, 8, 13-18, 25-28, 30, 32, 33-35, 37-39.— Stusak 1961a, 
p. 86, figs. 2, 3, 5d, 12, 13, 19, 21. 
Dictyla symphyti: Drake and Ruhoff 1960a, p. 51.— Stichel 1960a, p. 

340, fig. 174 [Poland; Bulgaria]. 
Dictyla humuli: Drake and Ruhoff 1960b, pp. 70-72, pi. 20. 
Type: Holotype 9, brachypterous; Germany; Kiel Coll. (Copenhagen 
Mus.). 

Distribution: Netherlands; France; Spain; Germany; Switzerland; 
Czechoslovakia; Austria; Hungary; Italy; Sicily; Yugoslavia; Rumania; 
Bulgaria; Poland; U.S.S.R. (Caucasus; Siberia); Algeria; Tunisia. 
Host Plants: Humulus sp.; Symphytum officinale; Symphytum sp. 
Note: Study of morphology and nymphs [Stusak and Stys]; eggs [Stusak]. 
•Dictyla imparis (Drake) 

Monanthia imparis Drake 1954b, p. 1 [Kenya]. 
Dictyla imparis: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9; Ngong, Kenya; Hungarian Mus. 
Distribution: Kenya. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 189 

*Dictyla indigena (Wollaston) 

Tingis indigena Wollaston 1858, p. 124 [Madeira; Echium candicans]. 
Monanthia indigena: Stal 1874, p. 58. — Horvath 1906a, p. 102 [Canary 
Islands]; 1909a, p. 292.— Lindberg 1936b, p. 30; 1958a, p. 79 [Cape 
Verde Islands; Echium stenosiphon; Echium hyper tr op icum]. — China 1938, 
p. 21, fig. 4.— Gomez-Menor 1955b, p. 249. 
Dictyla indigena: Drake and Ruhoff 1960a, p. 51. — Stichel 1960a, p. 336. 
Type: Sex unknown; Feijaa de Corte, Madeira; British Mus. 
Distribution: Madeira Islands; Cape Verde Islands; Canary Islands 
Host Plants: Echium candicans; Echium hypertropicum; Echium stenosiphon 

Dictyla indigena subsp. bugioensis (China) 

Monanthia indigena subsp. bugioensis China 1938, p. 22, fig. 4 [Madeira]. 

Dictyla indigena subsp. bugioensis: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Southern Dezerta, Madeira; British Mus. 
Distribution: Madeira Islands. 
Host Plant: Unrecorded. 

*Dictyla labeculata (Uhler) 

Monanthia labeculata Uhler 1893b, p. 264 [Calif.; Pinus monophylla]. — 

Drake 1917c, pp. 49, 51, fig. la.— Hurd 1946, p. 451 [N. Mex.]. 
Monanthia labecula [sic]: Van Duzee 1916, p. 26; 1917a, p. 261. 
Monanthia lobeculata [sic]: Drake 1943a, p. 142 [Colo.; Ariz.]. 
Dictyla labeculata: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype cf, macropterous; Argus Mountains, California, U.S.; 
USNM. 
Distribution: U.S. (Calif; Ariz.; Colo.; N. Mex.); Canada (B.C.). 
Host Plant: Pinus monophylla. 
*Dictyla leporis (Drake) 

Monanthia leporis Drake 1937a, p. 385 [China]. 
Dictyla leporis: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9; Kuling, Kiangsi Province, China; Vienna Mus. 
Distribution: China. 
Host Plant: Unrecorded. 
Dictyla leroyi (Schouteden) 

Monanthia leroyi Schouteden 1953e, p. 197 [Congo; Cordia chrysocarpa]. 
Dictyla leroyi: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Kivu, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Cordia chrysocarpa. 
*Dictyla loricata (Distant) 

Monanthia loricata Distant 1888, p. Ixxxiii [Brazil]. — Berg 1892, p. 201 
[Argentina], (reprint p. 101). — Champion 1898b, p. 64, pi. 3, fig. 
11.— Pennington 1921, p. 20.— Drake 1922c, p. 354; 1943a, p. 142 
[Bolivia; Paraguay]; 1947a, p. 1. — Monte 1937a, p. 33, fig. 12 [Cordia 



190 U.S. NATIONAL MUSEUM BULLETIN 24 3 

corymbosa]; 1939b, p. 76 [Venezuela; Cordia]; 1940d, p. 101. — Drake 
and Hambleton 1938a, p. 45 [Tournefortia]; 1945b, p. 356 [Co- 
lombia].— Drake and Poor 1938b, p. 106.— SUva 1956, p. 43, fig. 15. 
Dictyla loricata: Drake and Runoff 1960a, p. 51. 
Type: Sex unknown; Entrc Rios, Brazil; Brussels Mus. 
Distribution : Brazil; Colombia; Argentina; Venezuela; Bolivia; Para- 
guay; Ecuador. 
Host Plants: Cordia corymbosa; Cordia sp.; Tournefortia sp. 

* Dictyla lupata (Drake and Poor) 

Monanthia lupata Drake and Poor 1936b, p. 142 [India]. 
Dictyla lupata: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype cf, macroptcrous; Gujranwala, India; Drake Coll. 
(USNM). 
Distribution: India. 
Host Plant: Unrecorded. 

*Dictyla lupuli (Herrich-Schaeffer) 

Monanthia lupuli Herrich-Schaeffer 1837, p. 13, pi. 114, fig. 359; 1838, 
p. 58.— Scholz 1847, p. 120.— Mayr 1858, p. 571 [Austria].— Fieber 
1861, p. 126.— Frey-Gessner 1865, p. 232 [Switzerland].— Stal 1874, 
p. 59.— Puton 1879c, p. 121 [France].— Jakovlev 1880b, p. 109 
[Russia]; 1893, p. 294.— Reuter 1882b, pp. 120, 121 [Finland].— 
Hueber 1893, p. 351— Horvath 1906a, pp. 99, 102 [Italy; Bulgaria; 
Siberia].— Muzik 1907, p. 63.— Oshanin 1908, p. 453 [Spain; 
Rumania; Caucasus]. — Stichel 1926, p. 116 [Myosotis]; 1938a, p. 408 
[Poland].— Scholte 1935, p. 92 [Netherlands].— Blote 1945, p. 91 
[Yugoslavia].— Kiritshenko 1951, p. 253.— Bator 1953, p. 327, pi. 4, 
fig. 5.— Hoberlandt 1955, p. 94 [Turkey]. 
Monanthia (Physalocheila) lupuli: Fieber 1844, p. 85, pi. 7, figs. 19-21 
[Czechoslovakia; Hungary; Germany]. — Herrich-Schaeffer 1850, p. 
153. — Flor 1860, p. 354 [Estonia; Latvia; Algeria; Myosotis palustris]. 
Dictyla lupuli: Drake and Ruhoff 1960a, p. 51.— Stichel 1960a, p. 341 
[Sardinia]. 
Type: Unknown. 

Distribution: Netherlands; France; Spain; Germany; Switzerland; 
Czechoslovakia; Italy; Sardinia; Hungary; Austria; Bulgaria; Yugoslavia; 
Rumania; Turkey; Poland; Finland; U.S.S.R. (Estonia; Latvia; Siberia; 
Caucasus); Algeria. 
Host Plants: Myosotis palustris; Myosotis sp. 

Note: Described by Herrich-Schaeffer but wrongly accredited by him to 
Kunze [Herrich-Schaeffer 1837, p. 13]. 

*Dictyla monotropidia (Stal) 

Monanthia {Physalocheila) monotropidia Stal 1858, p. 63 [Brazil]. 
Monanthia monotropidia: Stal 1873, p. 133 [Colombia].— Distant 1888, 



LACEBUGS OF THE WORLD 191 

p. lxxxiii. — Champion 1898a, p. 47, pi. 3, figs. 24, a, b [Guatemala; 
Panama]; 1898b, p. 64.— Van Duzee 1907, p. 23 [Jamaica].— Osborn 
and Drake 1915b, p. 536.— Drake 1922c, p. 354 [Cuba; Haiti; 
Mexico]; 1926a, p. 86 [Cordia gerascanthus]; 1926b, p. 375; 1928e, p. 1 
[Honduras]; 1929, p. 35 [Costa Rica]; 1931b, p. 510; 1935, p. 9 
[Paraguay]; 1943a, p. 142 [Bolivia; Venezuela]; 1947a, p. 1.— 
Wolcott 1923, p. 247 [Puerto Rico] .— Fenton 1934, p. 199 [Argentina; 
cotton].— Costa Lima 1936, p. 129.— Bosq 1937, p. 130.— Drake and 
Poor 1937d, p. 301 [Peru; Cordia]; 1939b, p. 95.— Monte 1937a, p. 
33; 1937c, pp. 71, 72 [Cordia tomentosa]. — Drake and Hambleton 
1938a, p. 44; 1945, p. 356 [El Salvador].— Barber 1939, p. 371 — 
Bruner, Scaramuzza and Otero 1945, p. 59. — Swezey 1945, p. 372. — 
Hurd 1946, p. 451 [Trinidad; Cordia alliodora].— Singh 1953, p. 118. — 
Gibson and Carrillo 1959, p. 19. 
Dictyla monotropidia: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9; Brazil; Stockholm Mus. 

Distribution: Brazil; Colombia; Peru; Bolivia; Paraguay; Venezuela; 
Argentina; Panama; El Salvador; Guatemala; Mexico; Honduras; Costa 
Rica; Haiti; Cuba; Jamaica; Puerto Rico; Trinidad. 

Host Plants: Cordia alliodora; Cordia gerascanthus; Cordia tomentosa; Cordia 
sp.; cotton. 

Note: Intercepted on orchids from Mexico at port-of-entry, Texas 
[Swezey] . 

*Dictyla montandoni (Horvath) 

Monanthia montandoni Horvath 1885, p. 322 [Rumania]; 1906a, p. 99 
[Russia].— Kiritshenko 1951, p. 253.— Putshkov 1960, p. 304 [Tourne- 
jortia sibirica]. 
Dictyla montandoni: Drake and Ruhoff 1960a, p. 51. — Stichel 1960a, p. 
337. 
Type: Holotype 9; Canstantia, Rumania; Hungarian Mus. 
Distribution: Rumania; U.S.S.R. 
Host Plant: Tournefortia sibirica. 
Note: Life history study [Putshkov]. 

Dictyla montandoni var. rivalis (Horvath) 

Monanthia montandoni var. rivalis Horvath 1905b, p. 570 [Russia]; 1906a, 

p. 99. 
Dictyla montandoni var. rivalis: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9; Sarepta, Russia; Hungarian Mus. 
Distribution: U.S.S.R. 
Host Plant: Unrecorded. 



192 U.S. NATIONAL MUSEUM BULLETIN 243 

:i Dictyla nassata (Puton) 

Monanthia reticulata Rambur 1830, p. 166 [Spain]. 

Monanthia convergens (not Hcrrich-Schaeffcr): Costa 1847b, p. 19; 1847d, 

p. 255. 
Monanthia (Physatocheila) reticulata: Ficbcr 1844, p. 90. 
Monanthia nassata Puton 1874b, p. 436 [Algeria; France; Corsica]; 
1876, p. 279; 1879c, p. 1 22; 1 886a, p. 5 [Tunisia].— Bolivar and Chicote 
1879, p. 166 [Portugal].— Horvath 1906a, p. 101 [Eritrea; Turkey; 
Greece; Yugoslavia; Hungary; Italy; Egypt; Canary Islands]; 
L906d, p. 2; 1909a, p. 292.— Stichcl 1926, p. 115; 1938a, p. 408 
[Albania].— de Seabra 1931, p. 436, figs. 506(1), 508.— Lindberg 
1932a, p. 44; 1936b, p. 30; 1948, p. 60 [Cyprus; Sicily; Eckium 
sericcum]. — Mancini 1935, p. 10 [Giglio Islands]. — Vidal 1937, p. 
199 [Morocco]. — Kiritshcnko 1951, p. 253 [Russia]. — Hoberlandt 
1952, p. 112; 1955a, p. 94 [Israel; Caucasus].— Bator 1953, p. 327, 
pi. 2, fig. 8. — Priesncr and Alficri 1953, p. 65. — Stusak and Stys 
1959, pp. 178-184, 186, 197, figs. 3-6, 9-12, 19-24, 29, 31, 40. 
Dictyla nassata: Drake and Runoff 1960a, p. 51. — Stichel 1960a, p. 
339 [Malta; Sardinia; Lybia; Turkestan; Cynoglossum; Onosma]. 
Type: Unknown. 

Distribution: Spain; Portugal; Corsica; Sardinia: France; Sicily; 
Malta; Italy; Hungary; Albania; Yugoslavia; Greece; Cyprus; Turkey; 
Israel; U.S.S.R. (Turkestan; Caucasus); Morocco; Algeria; Tunisia; 
1 ;>t: Lybia; Eritrea; Giglio Islands; Canary Islands. 

Host Plants: Cynoglossum sp.; Echium sericcum; Halogclon satirus; Echium 
plantagineum; Onosma sp. 
Note: Study of immature forms [Stusak and Stys]. 

*Dictyla nodipennis (Horvath) 

Monanthia nodipennis Horvath 1910, p. 68 [Tanganyika]. Drake 1956a, 
p. 4 [Congo; Kenya; Uganda]. 

Monanthia ravaltarii Mancini 1953a, p. 185, fig. 2 [Ethiopia]. 

Dictyla nodipennis: Drake and Ruhoff 1960a, p. 51; 1961c, p. 115. 

D tyla zavattarii: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype ,; Kibongoto, Kilimandjaro; Stockholm Mus. 
Distribution: Tanganyika; Kenya; Uganda; Congo; Ethiopia. 
Host Plant: Unrecorded. 

IDictyla novaki Drake and Ruhoff 

mthia Jlexuosa Novak 1877a, p. 79, pi. 2, figs. 8-11 [Bohemia]; 

18771), p. 9, pi. 2, figs. 8-11.— Scudder IXQO, p. 357 [Horizon: 

Aquitanian]; 1891a, p. -121. 
Dictyla Jlexuosa: Drake and Ruhoff 1960a, pp. 11, 51. 
Dictyla novaki Drake and Ruhoff 1962a, p. 156. 



LACEBUGS OF THE WORLD 193 

Type: Sex undeterminable; fossil; Krottensee, Bohemia; Tertiary; 
Prague Mus. 
Distribution: Fossil. 

*Dictyla parilis (Drake) 

Monanthia parilis Drake 1936b, p. 700 [Argentina]. — Drake and Poor 

1939b, p. 95. 
Monanthia paritis [sic]: Drake 1943a, p. 142 [Paraguay; Brazil]. 
Dictyla parilis: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9, macropterous; Loreto, Argentina; Leningrad Inst. 
Distribution: Argentina; Paraguay; Brazil. 
Host Plant: Unrecorded. 

*Dictyla parmata (Distant) 

Monanthia parmata Distant 1888, p. Ixxxiii [Brazil]. — Champion 1898b> 
p. 64, pi. 3, fig. 12.— Drake 1922c, p. 355; 1930a, p. 1; 1931b, p. 
510 [Argentina]; 1935, p. 9 [Paraguay]; 1943a, p. 142 [Peru].— 
Drake and Hambleton 1934, p. 437 [Cordia corymbosa]. — Costa Lima 
1936, p. 130. — Monte 1937a, p. 33 [Cordia; Cordia verbenacea]; 1938d, 
p. 387; 1940d, p. 101 [Venezuela]; 1941e, p. 125 [Cochranea auchusae- 
folia]. — Silva 1956, p. 43, fig. 16 [Cordia curassavica]. 
Dictyla parmata: Drake and Ruhoff 1960a, p. 51. — Drake and Cobben 
1960, p. 69, figs. 74b, 75b, 76b [Aruba; Curacao; Bonaire; Cordia 
cylindristachya]. 
Type: Sex unknown; Botafogo, Brazil; Brussels Mus. 
Distribution: Brazil; Argentina; Paraguay; Peru; Venezuela; Bolivia; 
Netherlands Antilles (Aruba; Bonaire; Curacao). 

Host Plants: Cochranea auchusaefolia; Cordia corymbosa; Cordia curassavica; 
Cordia verbenacea; Cordia cylindristachya; Cordia sp. 

*Dictyla patquiana (Drake) 

Monanthia patquiana Drake 1955a, p. 79 [Argentina]. 

Dictyla patquiana: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype cf; Patquia, Argentina; British Mus. 
Distribution: Argentina. 
Host Plant: Unrecorded. 

*Dictyla picturata (Distant) Plate 34 

Monanthia picturata Distant 1902a, p. 243, pi. 15, fig. 11 [Cape Prov- 
ince].— Drake 1961c, p. 129, pi. 7. 
Dictyla picturata: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Cape Colony, South Africa; So. Afr. Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 



I'M U.S. NATIONAL MUSEUM BULLETIN 243 

♦Dictyla platyoraa (Ficbcr) 

inthia pUtiyoma Ficbcr 1861, p. 125 [Czechoslovakia; Austria; 
Myosotis palustris; Symphytum], — Jakovlcv 1869, p. Ill [Russia]; 1871, 
p. 24; 188Ul», [>. H>8 [Cauca^,. ; 1893, p. 294— Horvath 1874b, 
p. 432 [Hungary]; 1906a, p. 100 [Turkey; Siberia; Turkestan; Greece; 
Rumania; Echinospermum lapptda\- Renter 1890a, p. 250; 1891b, 
p. 1 84 ; 1 902, p. 1 51 .— Hucber 1 893, p. 349 [Germany].— Lctbierry and 
Severin 1896, p. 24 [Turk«>in.mia].— Muzik 1907, p. 63.— Stiche] 
1926, i-. 115. -Lindberg 1934, p. 29 [China].— Guide 1938, p. 314 
[Myosotis].— Kiritshenko 1951, p. 253.— Hoberlandt 1952, p. 112 
[Cvprus]; 1955, p. 94. 
Dictyla platyomia [sic]: Stal 1874, p. 57. 

Dictyla platyoma: Drake and Ruhoff 1960a, p. 51, pi. 5.— Stichel 1960a, 
p. 338 [Bulgaria]. 
Type: Unknown. 

Distribution: Germany; Austria; Czechoslovakia; Hungary; Rumania; 
Bulgaria; Greece; Turkey; Cyprus; U.S.S.R. (Turkestan; Tukmen; 
Siberia; Caucasus); China. 

Host Plants: Echinospermum lappula; Myosotis palustris; Myosotis sp.; 
Symphytum sp. 

"Dictyla pongana (Drake) 

Monanthia pongana Drake 1953d, p. 211 [South-West Africa]. 
Dictyla pongana: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype cf ; Okahandja, South- West Africa; British Mus. 
Distribution: South- West Africa. 
Host Plant: Unrecorded. 

"Dictyla pucallpana (Drake and Hambleton) 

Monanthia puc all pana Drake and Hambleton 1945, p. 356 [Peru]. 

Dictyla pucallpana: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype o", macropterous; Pucallpa, Peru; Drake Coll. 
(USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 

*Dictyla putoni (Montandon) 

Monanthia putoni Montandon 1895, p. 161 [Rumania; Echium]. — Hor- 
vath 1906a, p. 101 [Hungary; Russia; Turkey].— Stichel 1938a, 
p. 408 [Albania; Bulgaria]. — Kiritshcnkol951, p. 253. — Scidcnstiicker 
1954, p. 236.— Hoberlandt 1955, p. 94 [Caucasus].— Stusak and 
Styi 1959, p. 185. 

Dictyla putom: Drake and Ruhoff 1960a, p. 51.— Stichel 1960a, p. 339 
[Franc ej Egypt]. 
Type: Zorleni, Moldavic, Rumania; sex and deposition of type unknown. 



LACEBUGS OF THE WORLD 195 

Distribution: France; Hungary; Rumania; Bulgaria; Turkey; Albania; 
U.S.S.R. (Caucasus); Egypt. 
Host Plant: Echium sp. 

*Dictyla putoni var. pulla (Horvath) 

Monanthia putoni var. pulla Horvath 1905b, p. 572 [Hungary]; 1906a, 
p. 101 [Rumania; Greece; Sardinia; Tunisia]. — Novak and Wagner 
1951, p. 71 [Echium altissimum; Cynoglossum creticum; Onosma visianii]. — 
Priesner and Alfieri 1953, p. 65 [Egypt]. 
Diclyla putoni var. pulla: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Hungary; Hungarian Mus. 

Distribution: Hungary; Rumania; Yugoslavia; Greece; Sardinia; 
Egypt; Tunisia. 

Host Plants: Cynoglossum creticum; Echium altissimum; Onosma visianii. 
*Dictyla rasilis (Drake and Maa) 

Monanthia rasilis Drake and Maa 1955, p. 3 [Taiwan; Solanum ver- 

bascifolium]. 
Dictyla rasilis: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype d\ macropterous; Koshun, Formosa; Drake Coll. 
(USNM). 
Distribution: Taiwan. 
Host Plant: Solanum verbascifolium. 
Dictyla ruandae (Schouteden) 

Monanthia ruandae Schouteden 1953e, p. 199 [Ruanda]. 
Dictyla ruandae: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Bugarama, Ruanda; Cent. Afr. Mus. 
Distribution: Ruanda-Urundi. 
Host Plant: Unrecorded. 
*Dictyla ruficeps (Horvath) 

Monanthia ruficeps Horvath 1905b, p. 571 [Tunisia]; 1906a, p. 99. 
Dictyla ruficeps: Drake and Ruhoff 1960a, p. 51. — Stichel 1960a, p. 337. 
Type: Holotype 9; Cherichera, Tunisia; Hungarian Mus. 
Distribution: Tunisia. 
Host Plant: Unrecorded. 
*Dictyla sahlbergi (Horvath) 

Monanthia sahlbergi Horvath 1906a, p. 100 [Turkestan]. 
Dictyla sahlbergi: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Bekljar-bek, Turkestan, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 
*Dictyla salicorum (Baba) 

Stephanitis salicorum Baba 1925, p. 3. 

Monanthia salicorum: Takeya 1930a, pp. 67-72, 1 pi. (8 figs.) [Japan; 
Salix pupurea]; 1951a, p. 22.— Maa 1957, p. 132. 



196 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Dictyla salicorum: Drake and Runoff 1960a, p. 51. 
Type: Sea unknown; Shenano Province, Honshu, Japan; Nagano 
Agricultural Experiment Station, Japan. 
Distribvii' n : Japan. 

1 1 < ■>. i PLANT: Salix fnirfmrea. 

Note: Redeacription of type specimen [Takeya 1930]. 

♦Dictyla sauteri (Drake) 

Monanthia sauteri Drake 1923a, p. 103 [Taiwan]; 1927a, p. 53 [Philip- 
pines].- — Esaki 1926, p. 163. — Drake and Poor 1937b, p. 397 [Luzon; 
Cordia myxa]. — Takeya 1951a, p. 22. 
Dictyla sauteri: Drake and Runoff 1960a, p. 51. 
Type: Holotype 9; Kankau, Formosa; Horn Coll. 
Distribution: Taiwan; Philippine Islands (Luzon). 
Host Plant: Cordia myxa. 

*Dictyla seorsa (Drake and Poor) 

Monanthia seorsa Drake and Poor 19371), p. 398 [Taiwan]. — Drake 
1938b, p. 195 [China].— Takeya 1951a, p. 23.— Maa 1957, pp. 124, 
132 [India]. 
Dictyla seorsa: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype d\ macropterous; Formosa; Drake Coll. (USNM). 
Distribution: Taiwan; China; India. 
Host Plant: Unrecorded. 
*Dictyla seorsa var. inflata (Drake and Poor) 

Monanthia seorsa var. inflata Drake and Poor 1937b, p. 398 [Luzon]. 
Dictyla seorsa var. inflata: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9, macropterous; Los Banos, Luzon, Philippine Is- 
lands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 
*Dictyla sessoris (Drake and Poor) 

Monanthia sessoris Drake and Poor 1937b, p. 398 [Luzon]. 
Dictyla sessoris: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype 9, macropterous; Mt. Makiling, Luzon, Philippine 
Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon; Mindanao). 
Host Plant: L T nrecorded. 
*Dictyla sjostedti (HorvatM 

Monanthia sjostedti Horvath 1910, p. 69 [Tanganyika]. — Drake 1956a, 

p. 3 [Congo; Kenya; Uganda]; 1956f, p. 422 [Rhodesia; Hoslundia], 
Dictyla sjostedti: Drake and Ruhoff 1060a, p. 51. 
I yit.: Holotype 9; Kibongoto, Kilimandjaro; Stockholm Mus. 
Distribution: Tanganyika; Congo; Uganda; Kenya; Southern Rhodesia. 
Host Pi am : Jloslundia sp. 



LACEBUGS OF THE WORLD 197 

*Dictyla subdola (Horvath) 

Monanthia subdola Horvath 1905b, p. 571 [Armenia]; 1906a, p. 100 
[Turkestan]. — Oshanin 1908, p. 451 [Transcaucasus]. 

Dictyla subdola: Drake and Runoff 1960a, p. 51. — Stichel 1960a, p. 338. 
Type: Holotype 9; Vallis Araxis, Armenia; Hungarian Mus. 
Distribution: U.S.S.R. (Armenia; Turkestan; Transcaucasus). 
Host Plant: Unrecorded. 

*Dictyla triconula (Seidenstiicker) 

Monanthia triconula Seidenstiicker 1954, pp. 230, 233, 236, figs, lb, 2 
[Turkey; Cynoglossum officinale]. — Hoberlandt 1955, p. 95 [Cynoglos- 
sum]. 
Dictyla triconula: Drake and Ruhoff 1960a, p. 51. — Stichel 1960a, p. 
342, fig. 180. 
Type: Holotype cf; Namrun, Turkey; Seidenstiicker Coll. 
Distribution: Turkey. 
Host Plants: Cynoglossum officinale; Cynoglossum sp. 

*Dictyla tuberosa (Horvath) 

Monanthia tuberosa Horvath 1929, p. 324 [Natal], 

Dictyla tuberosa: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Weenen, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

*Dictyla uichancoi (Drake and Poor) 

Monanthia uichancoi Drake and Poor 1937b, p. 399 [Luzon]. 
Dictyla uichancoi: Drake and Ruhoff 1960a, p. 51. 
Type: Holotype cf, macropterous; Calamba, Luzon, Philippine Islands; 
Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon); Taiwan. 
Host Plant: Unrecorded. 

f Dictyla veterna (Scudder) Plate 35 

Monanthia veterna Scudder 1890, p. 359, pi. 23, figs. 5, 9 [Colo.]; 1891, p. 
422 [Horizon: Oliogocene]. 

Dictyla veterna: Drake and Ruhoff 1960a, pp. 11, 51. 
Type: Sex undeterminable; fossil; Florissant, Colorado, U.S.; MCZ. 
Distribution: Fossil. 

*Dictyla vulcanorum (Schouteden) 

Monanthia vulcanorum Schouteden 1953e, p. 200 [Ruanda]; 1957c, p. 

318.— Drake 1958b, p. 26 [Kenya]. 
Dictyla vulcanorum: Drake and Ruhoff 1960a, p. 51. 
Type: Sex unknown; Ruanda; Cent. Afr. Mus. 
Distribution: Ruanda-Urundi; Kenya. 
Host Plant: Unrecorded. 



198 U.S. NATIONAL MUSEUM BULLETIN 243 

Genus DICYSTA Champion 

Dicysta Champion 1897a, p. 5.— Drake 1922a, pp. 269-273 (key).— 
Monte 1939b, p. 68; 1941e, p. 92.— Hurd 1946, p. 477.— Drake and 
Ruhoff 1960a, p. 53. 
Type Species: Dicysta vitrea Champion. 

*Dicysta arnica Drake and Hambleton 

Dicysta arnica Drake and Hambleton 1939, p. 162 [Brazil]. — Monte 
1941e, p. 92 [Bignoniaceae]. 
Type: Holotype 9, macropterous; Belo Horizonte, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Bignoniaceae. 

*Dicysta braziliensis Drake 

Dicysta braziliensis Drake 1922a, p. 272 [Brazil]; 1931b, p. 512 [Peru]. 
Type: Holotype o 71 ; Santarem, Brazil; Carnegie Mus. 
Distribution: Brazil; Peru. 
Host Plant: Unrecorded. 

*Dicysta cara Drake 

Dicysta cara Drake 1939a, p. 87 [Queensland]. 

Type: Holotype 9, macropterous; Maleny, Queensland, Australia; 
Drake Coll. (USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

*Dicysta fonsecai Monte 

Dicysta fonsecai Monte 1940e, p. 299 [Brazil]. 
Type: Holotype o 71 ; Guaruja, Santos, Sao Paulo, Brazil; Monte Coll. 
(Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Dicysta hoi land i Drake 

Dicysta hollandi Drake 1922a, p. 271, fig. 1 [Brazil]. — Drake and Hamble- 
ton 1945, p. 365 [Peru]. 
Type: Holotype 9, macropterous; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil; Peru. 
Host Plant: Unrecorded. 
*Dicysta lauta Drake and Hambleton 

Dicysta lauta Drake and Hambleton 1935, p. 152, fig. 6 [Brazil; Adeno- 
calymna] . 
Type: Holotype cf, macropterous; Ponte Nova, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Adenocalymna sp. 



LACEBUGS OF THE WORLD 199 

*Dicysta limata Drake and Hambleton 

Dicysta limata Drake and Hambleton 1939, p. 162 [Brazil]. 

Type: Holotype cf, macropterous; Belem, Para, Brazil; Drake Coll. 
(USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Dicysta parilis Drake 

Dicysta parilis Drake 1939, p. 88 [Queensland]. 
Type: Holotype 9, macropterous; Mackay, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Dicysta peruviana Drake and Poor 

Dicysta peruviana Drake and Poor 1940, p. 231 [Peru]. 
Type: Holotype 9, macropterous; Lima, Peru; Drake Coll. (USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 

*Dicysta sagillata Drake 

Dicysta sagillata Drake 1922c, p. 376 [Panama]. — Monte 1939b, p. 68 
[Brazil; Bignoniaceae]. 
Type: Holotype 9, macropterous; Porto Bello, Panama; USNM. 
Distribution: Panama; Brazil. 
Host Plant: Bignoniaceae. 

*Dicysta smithi Drake 

Dicysta smithi Drake 1922a, p. 273 [Brazil]. — Drake and Hambleton 
1938b, p. 57.— Monte 1940c, p. 193 [Adenocalymna]. 
Type: Holotype 9, macropterous; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Adenocalymna sp. 

*Dicysta vitrea Champion 

Dicysta vitrea Champion 1897, p. 5, pi. 1, figs. 4, 4a [Panama]. — 
Drake 1922a, p. 270 [Brazil]; 1935, p. 20 [Paraguay].— Drake and 
Bondar 1932, p. 93 [Mansoa glaziovii]. — Drake and Hambleton 1935, 
p. 152 [Adenocalymna] ; 1945, p. 365 [Peru]. — Costa Lima 1936, p. 
126. — Monte 1937a, p. 34 [Petastoma formosum; Petastoma samydoides]; 
1937c, p. 71.— Singh 1953, p. 118.— Kormilev 1955a, p. 67 — 
Silva 1956, p. 27. 
Type: Holotype o 71 ; Volcan de Chiriqui, Panama; British Mus. 
Distribution: Panama; Brazil; Peru; Paraguay. 

Host Plants: Adenocalymna sp.; Mansoa glaziovii; Petastoma formosum; 
Petastoma samydoides. 



200 U.S. NATIONAL MUSEUM BULLETIN 243 

Genus DIPLOCYSTA Horvath 

Diplocysta Horvath 1925a, p. 11. — Drake and Runoff 1960a, p. 53. 
Type Species: Diplocysta bilobata Horvath. 

*Diplocysta bilobata Horvath 

Diplocysta bilobata Horvath 1925a, p. 12, fig. 7 [Australia]. — Hacker 
1928, p. 179. 
Type: Sex unknown; Geraldton, Australia; Stockholm Mus. 
Distribution: Australia (Western Australia). 
Host Plant: Unrecorded. 

*Diplocysta globuliformis Hacker 

Diplocysta globulif or mis Hacker 1928, p. 179, pi. 22, figs. 9, 10 [Thursday 
Island; Queensland]. 
Type: Sex unknown; Thursday Island; Queensland Mus. 
Distribution: Thursday Island; Magnetic Island; Australia (Queens- 
land). 

Host Plant: Unrecorded. 

Diplocysta papuana Drake 

Diplocysta papuana Drake 1960, p. 379 [New Guinea]. 

Type: Holotype cf; Bisianumu, E. of Port Moresby, New Guinea 
(Papua); Bishop Mus. 

Distribution: New Guinea (Papua). 

Host Plant: Unrecorded. 

*Diplocysta rustica Drake 

Diplocysta rustica Drake 1960, p. 380 [New Guinea]. 

Type: Holotype d% macropterous; Eubenangee, New Guinea; Drake 
Coll. (USNM). 

Distribution: New Guinea. 

Host Plant: Unrecorded. 

*Diplocysta trilobata Drake and Poor Plate 28 

Diplocysta trilobata Drake and Poor 1939c, p. 205 [Australia]. — Drake 
and Davis 1960, figs. 43, 44.— Drake 1961b, p. 108 [Tasmania, 
South Australia].— Drake and Ruhoff 1962b, p. 250. 
Type: Holotype 9, brachypterous; Kiata, Victoria, Australia; Drake 
Coll. (USNM). 
Distribution: Australia (Victoria; South Australia; Tasmania). 
Host Plant: Unrecorded. 

Genus DULINIUS Distant 

Dulinius Distant 1903a, p. 48; 1903b, p. 133.— Drake 1953b, p. 95 — 

Drake and Ruhoff 1960a, p. 54. 
Sankisia Schouteden 1916b, p. 293.— Drake and Ruhoff 1960a, p. 81. 
Type Species: Dulinius conchatus Distant. 



LACEBUGS OF THE WORLD 201 

*Dulinius bellus Drake 

Dulinus [sic] bellus Drake 1948g, p. 175 [Mozambique]. 
Type: Holotype 9, macropterous; Delagga Bay, Mozambique; Drake 
Coll. (USNM). 
Distribution : Mozambique. 
Host Plant: Unrecorded. 

Dulinius burgeoni Schouteden 

Dulinius burgeoni Schouteden 1953d, p. 170 [Congo]. 
Type: Sex unknown; Albertville, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Dulinius conchatus Distant 

Dulinius conchatus Distant 1903a, p. 48 [Ceylon]; 1903b, p. 133, fig.96; 
1910a, p. 110 [India; Morinda] .—Drake and Poor 1937a, p. 18 
[Java; Negros; Luzon]. — Drake 1937a, p. 387 [Malaya]. — Maa 1957, 
pp. 130, 132 [China]. 
Type: Sex unknown; Batticaloa, Ceylon; British Mus. 
Distribution: Ceylon; India; China; Federation of Malaya; Greater 
Sunda Islands (Java); Philippine Islands (Luzon; Negros). 
Host Plant: Morinda sp. 

*Dulinius congruus Drake 

Dulinius congruus Drake 1953b, p. 96 [Cape Province]. 

Type: Holotype 9, macropterous; Grahamstown, South Africa; Drake 
Coll. (USNM). 

Distribution: South Africa (Cape Province). 

Host Plant: Unrecorded. 

*Dulinius kraussi Drake 

Dulinius kraussi Drake 1953b, p. 95 (nom. nud.); 1954c, p. 72 [Kenya]. 

Type: Holotype cf, macropterous; Diani Beach, Kenya; Drake Coll. 
(USNM). 

Distribution: Kenya. 

Host Plant: Unrecorded. 

*Dulinius pulchrus (Schouteden) 

Sankisia pulchra Schouteden 1916b, p. 294 [Congo]. 

Dulinius pulchrus: Drake 1953b, p. 95. 
Type: Sex unknown; Sankisia, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Southern Rhodesia. 
Host Plant: Unrecorded. 
Dulinius unicolor (Signoret) 

Tingis unicolor Signoret 1861, p. 956 [Malagasy]. — Stal 1873, p. 134. 

Dulinius nigrolineatus Distant 1913, p. 158, pi. 11, fig. 16 [Seychelles]. 

Galeatus involutus Drake 1925b, p. 108. 

681-552—64 14 



202 U.S. NATIONAL MUSEUM BULLETIN 243 

Dulinius unicolor: Drake and Ruhoff 1960c, p. 32, figs. 2, 3; 1960e, p. 
80.— Carayon 1960, p. 110. 
Type: Sex unknown; Madagascar; Vienna Mus. 
Distribution: Malagasy Republic; Seychelles; Uganda. 
Host Plant: Taken on "musenosa." 
Note: Predators [Carayon]. 

Genus DYSPHARSA Drake and Hambleton 

Dyspharsa Drake and Hambleton 1944b, p. 127.— Hurd 1946a, p. 466. — 
Drake and Ruhoff 1960a, p. 54. 
Type Species: Leptopharsa myersi Drake. 

*Dyspharsa myersi (Drake) 

Leptopharsa myersi Drake 1926a, p. 87 [Trinidad]. 

Dyspharsa myersi: Drake and Hambleton 1944b, p. 128. — Hurd 1946, 
p. 467 [Cuba]. 
Type: Holotype 9, macropterous; Mina Carlotta, Trinidad; Drake Coll. 
(USNM). 
Distribution: Trinidad; Cuba. 
Host Plant: Unrecorded. 

Genus ELASMOGNATHUS Fieber 

Elasmognathus Fieber 1844, p. 90— Stal 1873, pp. 121, 129.— Lethierry 
and Severin 1896, p. 22.— Distant 1903b, p. 141.— Drake and Poor 
1937a, p. 10.— Drake and Ruhoff 1960a, p. 54. 
Type Spectes: Elasmognathus helferi Fieber. 

Elasmognathus fieberi Stal 

Elasmognathus fieberi Stal 1855a, p. 38 [Natal]; 1873, p. 129.— Distant 

1902a, p. 242, pi. 15, fig. 7. 
Monanthia {Elasmognathus) fieberi: Stal 1865, p. 29. 
Monanthia fieberi: Walker 1873a, p. 195. 
Type: Holotype o 71 ; Natal, South Africa; Stockholm Mus. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

*Elasmognathus helferi Fieber 

Elasmognathus helferi Fieber 1844, p. 91, pi. 7, figs. 33-41 [Ostindien]. — 
Herrich-Schaeffer 1850, p. 156.— Stal 1873, p. 129 [India orien- 
talise—Distant 1903b, p. 142.— Drake and Davis 1960, figs. 55, 56. 
Monanthia helferi: Walker 1873a, p. 195. 
Elasmognathus helfneri [sic]: Drake 1927d, p. 309 [Luzon]. 
Type: "Ostindien"; sex and deposition of type unknown. 
Distribution: India; Philippine Islands (Luzon). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 203 

Genus ELASMOTROPIS Stal 

Elasmotropis Stal 1874, p. 54— Horvath 1906a, pp. 14, 56 (key).— 
Muzik 1907, pp. 50, 57.— Oshanin 1908, p. 426; 1912, p. 44.— Stichel 
1926, pp. 104, 110 (key); 1935, p. 349; 1938a, p. 406; 1960a, p. 304; 
1960b, p. 395; 1960c, p. 1 34.— Kiritshenko 1951, pp. 242, 248 
(key).— Drake and Runoff 1960a, p. 55. 
Phyllontochila [sic] {Elasmotropis): Lethierry and Severin 1896, p. 16. 
Type Species: Monanthia {Phyllontocheila) echinopsis Fieber= Tingis testacea 
Herrich-Schaeffer. 
Elasmotropis abimva (Schouteden) 

Elasmognathus abimva Schouteden 1953d, p. 169 [Congo]. 
Type: Sex unknown; Abimva, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Note: From the description, the species seems to belong to Elasmotropis 
rather than Elasmognathus and is here so transferred. 
Elasmotropis burgeoni (Schouteden) 

Elasmognathus burgeoni Schouteden 1953d, p. 167 [Congo]. 
Type: Sex unknown; Yebo Moto, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Note: From the description, the species seems to belong to Elasmotropis 
rather than Elasmognathus and is here so transferred. 
Elasmotropis distans (Jakovlev) 

Monanthia {Platychila) distans Jakovlev 1903a, p. 2 [Mongolia]. 
Elasmotropis distans: Horvath 1906a, p. 59.— Wu 1935, p. 449. 
Type: Holotype 9, macropterous; Khangai, Mongolia; Leningrad Inst. 
Distribution: Mongolia Republic. 
Host Plant: Unrecorded. 
Elasmotropis motoensis (Schouteden) 

Elasmognathus motoensis Schouteden 1953d, p. 168 [Congo]. 
Type: Sex unknown; Yebo Moto, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Note: From the description, the species seems to belong to Elasmotropis 
rather than Elasmognathus and is here so transferred. 
*Elasmotropis selecta (Horvath) 

Phyllontocheila testacea var. selecta Horvath 1891, p. 79. 
Elasmotropis selecta: Horvath 1906a, p. 58 [Russia; Armenia]. — Oshanin 
1908, p. 426 [Transcaucasus].— Kiritshenko 1951, p. 248.— Stichel 
1960a, p. 304. 
Type: Unknown. 

Distribution: U.S.S.R. (Armenia; Transcaucasus). 
Host Plant: Unrecorded. 



204 U.S. NATIONAL MUSEUM BULLETIN 243 

*Elasmotropis testacea (Herrich-Schaeffer) 

Tingis testacea Herrich-Schaeffer 1830, heft 118, tab. 23 [Germany]; 

1835, p. 58. 
Monanthia testacea: Herrich-Schaeffer 1838, pp. 52, 60, pi. 125, figs. 
h,i. [Czechoslovakia]. — Puton 1879c, p. Ill [France, Algeria]. — 
Hueber 1893, p. 333. 
Monanthia (Phyllontocheila) echinopsis Fieber 1844, p. 62, pi. 5, figs. 19-22 

[Echinops sphaerocephala] . 
Monanthia (Phyllontocheila) testacea: Herrich-Schaeffer 1850, p. 155. 
Monanthia echinopsis: Mayr 1858, p. 570 [Austria]. 
Monanthia echinopsidis [sic]: Fieber 1861, p. 122. — Horvath 1874b, p. 
432 [Hungary].— Jakovlev 1876b, p. 67 [Russia]; 1880b, p. 104 
[Caucasus]. 
Elasmotropis echinopsidis: Stal 1874, p. 54. 
Phyllontochila testacea: Reuter 1891a, p. 26 [Greece]. — Lethierry and 

Severin 1896, p. 17 [Armenia]. 
Elasmotropis testacea: Horvath 1906a, p. 58. — Stichel 1926, p. 110; 
1960a, p. 304, fig. 108a [Yugoslavia].— Vidal 1937, p. 197 [Mo- 
rocco].— Guide 1938, p. 282, fig.— Lindberg 1948, p. 58 [Cyprus; 
Pistacia; Ceratonia siliqua]. — Hoberlandt 1951, p. 22 [Poland; Bul- 
garia; Syria; Egypt; Tunisia; Israel]. — Priesner and Alfieri 1953, 
p. 64 [Echinops].— Stusak 1961a, p. 82, figs. 5f, 8; pi. 1, fig. B. 
Type: Sex unknown; Regensburg, Germany; Munich Mus. 
Distribution: France; Germany; Czechoslovakia; Austria; Poland; 
Hungary; Bulgaria; Yugoslavia; Greece; Cyprus; Syria; Israel; 
U.S.S.R. (Armenia; Caucasus); Egypt; Algeria; Morocco; Tunisia. 
Host Plants: Ceratonia siliqua; Echinops sphaerocephala; Echinops sp.; 
Pistacia sp. 

Note: Eggs (Stusak). 
*Elasmotropis testacea var. egena (Puton) 

Monanthia testacea var. egena Puton 1879c, p. Ill [Algeria]. 
Elasmotropis testacea var. egena: Horvath 1906a, p. 58 [Tunisia]. — 
Lindberg 1932a, p. 41 [Morocco] .—Vidal 1937, p. 197— Blote 1945, 
p. 86. — Priesner and Alfieri 1953, p. 64. 
Type: Sex unknown; Oran, Algeria; Paris Mus. 
Distribution: Algeria; Tunisia; Morocco; Egypt. 
Host Plant: Unrecorded. 

*Elasmotropis testacea var. platydera Horvath 

Elasmotropis testacea var. platydera Horvath 1906a, p. 58 [Hungary; Bul- 
garia].— Stichel 1926, p. 110; 1960a, p. 305 [Echinops]. 
Type: Unknown. 
Distribution: Hungary; Bulgaria. 
Host Plant: Echinops sp. 



LACEBUGS OF THE WORLD 205 

*Elasmotropis testacea var. vicina Horvath 

Elasmotropis testacea var. vicina Horvath 1906a, p. 58 [Russia; Syria; 
Egypt].— Stichel 1926, p. 110; 1960a, p. 305 [Pistacia; Ceratonia; 
Cupressus].—Vida\ 1937, p. 197 [Morocco].— Lindberg 1948, p. 58 
[Cyprus; Caucasus]. — Priesner and Alfieri 1953, p. 64. — Seiden- 
stucker 1954, p. 235 [Turkey].— Hoberlandt 1955, p. 88. 
Type: Unknown. 

Distribution: Syria; Cyprus; Turkey; U.S.S.R. (Caucasus); Egypt; 
Morocco. 
Host Plants: Ceratonia sp.; Cupressus sp.; Pistacia sp. 

Genus ENGYNOMA Drake 

Engynoma Drake 1942b, p. 362.— Drake and Ruhoff 1960a, p. 55. 
Type Species: Perissonemia tasmaniae Drake and Poor. 

*Engynoma angulata (Hacker) 

Tingis (Tingis) angulata Hacker 1929, p. 329, pi. 34, fig. 9 [Queensland]. 
Engynoma angulata: Drake 1942b, p. 362. 
Type: Sex unknown; Upper Brookfield, Queensland, Australia; Queens- 
land Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Engynoma deaba Drake 

Engynoma deaba Drake 1942b, p. 363 [Queensland]. 
Type: Holotype 9, macropterous; National Park, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Engynoma immaculata Drake 

Engynoma immaculata Drake 1942b, p. 362 [Queensland]. 
Type: Holotype d\ macropterous; Cedar Creek, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Engynoma isolata Drake and Ruhoff 

Tingis insularis Hacker 1929, p. 330, pi. 34, fig. 10 [Queensland]. 

Engynoma insularis: Drake 1942b, p. 362. 

Engynoma isolata Drake and Ruhoff 1962c, p. 134. 
Type: Sex unknown; Dunk Island, North Queensland, Australia; 
Queensland Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 



206 U.S. NATIONAL MUSEUM BULLETIN 243 

*Engynoma spinicollis (Horvath) 

Tingis (Tingis) spinicollis Horvath 1925a, p. 6, fig. 3 [Queensland]. 

Tingis spinicollis: Hacker 1928, p. 183. 

Engynoma spinicollis: Drake 1942b, p. 362. 
Type: Sex unknown; Malanda, Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Engynoma tasmaniae (Drake and Poor) 

Perissonemia tasmaniae Drake and Poor 1937b, p. 402 [Tasmania]. 

Engynoma tasmaniae: Drake 1942b, p. 362. 
Type: Holotype cT, macropterous; Launceston, Tasmania, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Tasmania). 
Host Plant: Unrecorded. 

Genus ENGYOTINGIS Drake and Ruhoff 

Engyotingis Drake and Ruhoff 1961b, p. 175. 
Type Species: Engyotingis cybele Drake and Ruhoff. 

*Engyotingis cybele Drake and Ruhoff 

Engyotingis cybele Drake and Ruhoff 1961b, p. 176 [Basilan]. 
Type: Holotype o" , macropterous; Basilan Island; Drake Coll. (USNM). 
Distribution: Basilan Islands (Basilan). 
Host Plant: Unrecorded. 

*Engyotingis tonkinana (Drake and Maa) 

Tpsotingis tonkinana Drake and Maa 1955, p. 7 [Viet-Nam]. — Drake 

1958d, p. 153. 
Engyotingis tonkinana: Drake and Ruhoff 1961b, p. 177. 
Type: Holotype cf, macropterous; Tonkin, Hoa-Binh, Indochina; 
Drake Coll. (USNM). 
Distribution: Viet-Nam. 
Host Plant: Unrecorded. 

Genus EOTINGIS Scudder 

Eotingis Scudder 1890, p. 359.— Hurd 1946, p. 455.— Drake and 
Ruhoff 1960a, pp. 10, 55. 
Type Species: Eotingis antennata Scudder. 

fEotingis antennata Scudder 

Eotingis antennata Scudder 1890, p. 360, pi. 23, figs. 1, 3 [Colo.]; 1891, 
p. 406 [Horizon: Oligocene]. — Drake and Ruhoff 1960a, pp. 10, 11. 
Type: Sex undeterminable; fossil; Florissant, Colorado, U.S.; MCZ. 
DiSTRmuTiON : Fossil. 



LACEBUGS OF THE WORLD 207 

Genus EPIMIXIA Kirkaldy 

Epimixia Kirkaldy 1908a, p. 779— Horvath 1925a, p. 15.— Drake and 
Runoff 1960a, p. 55; 1961a, pp. 1-6 (key). 
Type Species: Epimixia alitophrosyne Kirkaldy. 

*Epimixia alitophrosyne Kirkaldy 

Epimixia alitophrosyne Kirkaldy 1908a, p. 780 [New South Wales]. — 
Drake and Ruhoff 1961a, p. 3, fig. 1 a, b [Western Australia]. 
Type: Holotype 9, macropterous; Sydney, New South Wales, Aus- 
tralia; Hawaii. Sugar Plant. Assn. 
Distribution: Australia (New South Wales; Western Australia). 
Host Plant: Unrecorded. 

Epimixia nigriceps (Signoret) 

Agramma nigriceps Signoret 1881, p. L [New Caledonia]. 

Serenthia nigriceps: Lethierry and Severin 1896, p. 5. 

Epimixia nigriceps: Drake and Ruhoff 1960a, p. 55; 1961a, p. 5. 
Type: Sex unknown; New Caledonia; Vienna Mus. 
Distribution: New Caledonia. 
Host Plant: Unrecorded. 

Epimixia nigripes Horvath 

Epimixia alitophrosyne var. nigripes Horvath 1925a, p. 16 [Queensland]. 

Epimixia nigripes: Drake and Ruhoff 1961a, p. 3. 
Type: Holotype cf; Evelyne, Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Epimixia nigripes subsp. dysmica Drake and Ruhoff 

Epimixia alitophrosyne (not Kirkaldy): Horvath 1925a, p. 16 [Queens- 
land].— Hacker 1927, p. 20, fig. 16; 1928, p. 186 [South Australia; 
New South Wales]. 
Epimixia nigripes subsp. dysmica Drake and Ruhoff 1961a, p. 4. 
Type: Holotype o 71 ; Colosseum, Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland; New South Wales; South Aus- 
tralia) . 
Host Plant: Unrecorded. 

*Epimixia veteris Drake 

Epimixia veteris Drake 1944a, p. 71 [Queensland]; 1961b, p. 112 [Casu- 
arina]. — Drake and Ruhoff 1961a, p. 4, fig. 2 [South Australia; 
Western Australia]. 
Type: Holotype o* , macropterous; Samsonvale, Queensland, Australia; 
Drake Coll. (USNM). 

Distribution: Australia (Queensland; South Australia; Western Aus- 
tralia). 
Host Plant: Casuarina sp. 



208 U.S. NATIONAL MUSEUM BULLETIN 243 

*Epimixia vittata Horvath Plate 13 

Epimixia vittata Horvath 1925a, p. 16 [Queensland]. — Hacker 1928, p. 
186 [Tasmania; South Australia; New South Wales]. — Drake and 
Runoff 1961a, p. 4, fig. 3. 
Type: Holotype 9; Colosseum, Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland; Tasmania; New South Wales; 
South Australia). 
Host Plant: Unrecorded. 

*Epimixia vulturna (Kirkaldy) 

Teleonemia vulturna Kirkaldy 1908a, p. 781 [Queensland]. — Hacker 

1927, p. 22. 
Epimixia vulturna: Drake 1944a, p. 67. — Drake and Ruhoff 1961a, p. 5, 

fig. 4 [New South Wales; South Australia]. 
Epimixia tenuatis Drake 1944a, p. 71; 1961b, p. 112. 
Epimixia evansi Drake 1944a, p. 72 [Tasmania]. 
Type: Holotype 9, macropterous; Kuranda, Queensland, Australia; 
Hawaii. Sugar Plant. Assn. 

Distribution: Australia (Queensland; New South Wales; South Aus- 
tralia; Tasmania). 
Host Plant: Unrecorded. 

Genus ERITINGIS Drake and Ruhoff 

Eritingis Drake and Ruhoff 1962d, p. 497. 
Type Species: Teleonemia pacifica Kirkaldy. 
*Eritingis agyiates Drake and Ruhoff 

Eritingis agyiates Drake and Ruhoff 1962d, p. 498 [Queensland]. 
Type: Holotype cf, macropterous; North Pine, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Eritingis amoena Drake and Ruhoff 

Eritingis amoena Drake and Ruhoff 1962d, p. 504 [New South Wales; 
Tasmania; South Australia]. 
Type: Holotype 9, macropterous; Mittagong, New South Wales, 
Australia; Drake Coll. (USNM). 
Distribution: Australia (New South Wales; South Australia; Tasmania). 
Host Plant: Unrecorded. 
*Eritingis aporema Drake and Ruhoff 

Eritingis aporema Drake and Ruhoff 1962b, p. 250 (nom. nud.) 
[South Australia; Flinders Island]; 1962d, p. 503 [Queensland]. 
Type: Holotype cf, macropterous; Stanthorpe, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland; South Australia; Flinders Island). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 209 

*Eritingis exalla (Drake) 

Tingis exalla Drake 1961b, p. 110 [New South Wales]. 
Eritingis exalla: Drake and Ruhoff 1962d, p. 497. 
Type: Holotype cf; Kosciusko, New South Wales, Australia; Austr. 
Mus. 
Distribution: Australia (New South Wales). 
Host Plant: Unrecorded. 

*Eritingis hylaea Drake and Ruhoff 

Eritingis hylaea Drake and Ruhoff 1962d, p. 502 [Queensland]. 

Type: Holotype o 71 , macropterous; Roma, Queensland, Australia; 
Drake Coll. (USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

*Eritingis koebeli (Drake) 

Nether sia koebeli Drake 1944a, p. 74 [Australia]. 

Eritingis (nom. nud.) koebeli: Drake and Ruhoff 1962b, p. 250 

[South Australia]. 
Eritingis koebeli: Drake and Ruhoff 1962d, p. 497. 
Type: Holotype 9, macropterous; Australia; Drake Coll. (USNM). 
Distribution: Australia (South Australia). 
Host Plant: Unrecorded. 

*Eritingis nostratis (Drake) 

Nether sia nostratis Drake 1944a, p. 74 [Queensland]. 
Eritingis nostratis: Drake and Ruhoff 1962d, p. 497. 
Type: Holotype 9, macropterous; Cedar Creek, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Eritingis pacifica (Kirkaldy) 

Teleonemia pacifica Kirkaldy 1908a, p. 780 [Viti Levu] (in part, specimen 

from Rewa only). 
Ulonemia pacifica: Drake and Poor 1943, p. 193 (in part). — Drake and 

Hurd 1945a, p. 288 (in part). 
Eritingis pacifica: Drake and Ruhoff 1962d, pp. 497, 498. 
Type: Lectotype 9; Rewa, Viti Levu, Fiji Islands; Hawaii. Sugar Plant. 
Assn. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Unrecorded. 
Note: Lectotype [Drake and Ruhoff]. 

*Eritingis recens (Drake and Poor) 

Perissonemia {Ulonemia) recentis Drake and Poor 1937a, p. 5 [Singapore!. 
Perissonemia recentis: Drake and Poor 1937b, p. 400. 



210 U.S. NATIONAL MUSEUM BULLETIN 243 

Ulonemia recens: Drake 1947d, p. 229 [Viet-Nam]. 
Eritingis recens: Drake and Runoff 1962d, p. 497, fig. 3. 
Type: Holotype cf, macropterous; Singapore, Straits Settlements; 
USNM. 
Distribution: Singapore; Viet-Nam. 
Host Plant: Unrecorded. 

*Eritingis trivirgata (Horvath) 

Teleonemia pacifica Kirkaldy 1908a, p. 780 [Queensland] (in part, speci- 
men from Kuranda, Queensland only). 
Tingis (Tropidocheila) trivirgata Horvath 1925a, p. 6. — Hacker 1927, p. 

24. 
Ulonemia pacifica: Drake and Poor 1943, p. 193 (in part). — Drake and 

Hurd 1945a, p. 288 (in part).— Drake 1961b, p. 112. 
Tingis virgata [sic]: Drake 1947c, p. 113 (fig. only). 
Eritingis (nom. nud.) trivirgata: Drake and Ruhoff 1962b, p. 250. 
Eritingis trivirgata: Drake and Ruhoff 1962d, p. 497. 
Type: Lectotype 9; Mt. Tambourine, Australia; Stockholm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
Note: Lectotype [Drake and Ruhoff 1962d]. 

*Eritingis violina Drake and Ruhoff 

Eritingis violina Drake and Ruhoff 1962d, p. 500 [New South Wales]. 

Type: Holotype cf, macropterous; Glouchester, New South Wales, 
Australia; Drake Coll. (USNM). 

Distribution: Australia (New South Wales). 

Host Plant: Unrecorded. 

Genus ESOCAMPYLIA Hacker 

Esocampylia Hacker 1929, p. 326. — Drake and Ruhoff 1960a, p. 56. 
Type Spectes: Esocampylia incarinata Hacker. 

*Esocampylia hackeri Drake 

Esocampylia hackeri Drake 1942a, p. 14 [Australia; moss; lichens]. 
Type: Holotype 9; Mt. Lofty Ridge, Australia; Drake Coll. (USNM). 
Distribution: Australia. 
Host Plants: Moss; lichens. 

*Esocampylia incarinata Hacker 

Esocampylia incarinata Hacker 1929, p. 326, pi. 32, fig. 4 [Queensland]. 

Type: Sex unknown; Blackbutt Range, Queensland, Australia; Queens- 
land Mus. 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 211 

Genus ETEONEUS Distant 

Eteoneus Distant 1903b, p. 129.— Drake 1946, p. 28.— Drake and 
Runoff 1960a, p. 56. 
Type Species: Serenthia dilatata Distant. 

Eteoneus angolensis Drake 

Eteoneus congolensis (not Schouteden) : Drake 1955b, p. 87, fig. 1 [Angola]. 

Eteoneus angolensis Drake 1956b, p. 113. 
Type: Sex unknown; Dundo, Angola; Cent. Afr. Mus. 
Distribution: Angola. 
Host Plant: Unrecorded. 

*Eteoneus angulatus Drake and Maa 

Eteoneus angulatus Drake and Maa 1953, p. 89 [China]. 

Type: Holotype 9, macropterous; Siyingpan, Yungan Hsien, Fukien, 
China; Drake Coll. (USNM). 

Distribution: China. 

Host Plant: Unknown. 

Note: Found "resting on leaf of Ulmus sp." [Drake and Maa]. 

*Eteoneus boops (Blote) 

Compseuta boopsBlote 1945, p. 89 [Sumatra]. 
Eteoneus boops: Drake 1948h, p. 75. 
Type: Sex unknown; Lubuk Sikaping, Sumatra; Leiden Mus. 
Distribution: Greater Sunda Islands (Sumatra); Philippine Islands 
(Mindanao). 

Host Plant: Unrecorded. 
*Eteoneus confectus Drake 

Eteoneus confectus Drake 1942a, p. 7 [Malaya]. 
Type: Holotype 9, macropterous; "Malacca, Perak"; Drake Coll. 
(USNM). 
Distribution: Federation of Malaya; Greater Sunda Islands (Java). 
Host Plant: Unrecorded. 

Note: Type locality taken from label on type specimen. 
Eteoneus congolensis Schouteden 

Eteoneus congolensis Schouteden 1923, p. 85 [Congo]. 
Type: Sex unknown; Watsa, Haut-Uele, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 
*Eteoneus dilatatus (Distant) 

Serenthia dilatata Distant 1903a, p. 46 [Burma]. 

Eteoneus dilatatus: Distant 1903b, p. 129, fig. 93.— Takeya 1931, p. 82, 
pi. 9, figs. 11-14 [Taiwan]; 1951a, p. 17. 
Type: Sex unknown; Tenasserim, Burma; British Mus. 
Distribution: Burma; Taiwan. 
Host Plant: Unrecorded. 



212 U.S. NATIONAL MUSEUM BULLETIN 243 

*Eteoneus esakii Drake 

Eteoneus esakii Drake 1939e, p. 102 [Peleliu; Premna]; 1956d, p. 110, 
fig. 5 [Koror]. — Takeya 1951a, p. 17. 
Type: Holotype cf ; Akarokuru-Shiasu, Peliliou, Palau Islands; Kyushu 
Univ. 
Distribution: Palau Islands (Peleliu; Koror). 
Host Plant: Premna sp. 
Note: Taken at light [Drake 1939f]. 

Eteoneus homelys Drake and Ruhoff 

Eteoneus homelys Drake and Ruhoff 1962a, p. 159, pi. 41 [Amboina]. 

Type: Holotype cT, macropterous; Amboina, Malay Archipelago; Cal. 
Acad. 

Distribution: Moluccas (Amboina Island). 

Host Plant: Unrecorded. 

*Eteoneus inopinus Drake 

Eteoneus inopinus Drake 1945, p. 99 [Rhodesia]; 1958b, p. 29 [Congo]. 
Type: Holotype o", macropterous; Bulamayo, Southern Rhodesia; Drake 
Coll. (USNM). 
Distribution: Southern Rhodesia; Congo. 
Host Plant: Unrecorded. 

*Eteoneus lectus Drake 

Eteoneus lectus Drake 1960, p. 353, fig. 9 [New Britain]. 
Type: Holotype cf ; Kerawat, Gazelle Peninsula, New Britain; Bishop 
Mus. 
Distribution: New Britain Island, Bismarck Archipelago. 
Host Plant: Unrecorded. 

Eteoneus minor Schouteden 

Eteonus [sic] minor Schouteden 1955a, p. 29 [Congo]. 
Type: Sex unknown; Moto, Ituri, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Eteoneus peroronus Drake Plate 11 

Eteoneus peroronus Drake 1942a, p. 7 [Malaya]; 1960, p. 353 [New 
Guinea].— Drake and Ruhoff 1962a, p. 160, pi. 42 (fig. only). 
Type: Holotype 9, macropterous; Parit Buntar, Malaya; Drake Coll. 
(USNM). 

Distribution: Federation of Malaya; New Guinea (Papua; Northeast 
Netherlands). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 213 

*Eteoneus sarptus Drake and Poor 

Eteonus [sic] sarptus Drake and Poor 1937a, p. 12 [Mindanao]. 

Type: Holotype 9, macropterous; Surigao, Mindanao, Philippine 
Islands; Drake Coll. (USNM). 

Distribution: Philippine Islands (Mindanao). 

Host Plant: Unrecorded. 

*Eteoneus sigillatus Drake and Poor 

Eteoneus sigillatus Drake and Poor 1936b, p. 144 [India]. — Singh 1953, 

p. 118. 
Eteoneus sagillatus [sic]: Drake 1946, p. 28. 
Type: Holotype 9, macropterous; Haldgaddi, Kalagarh Division, United 
Provinces; Drake Coll. (USNM). 
Distribution: India. 
Host Plant: Unrecorded. 

*Eteoneus virtutis Drake and Poor 

Eteonus [sic] virtutis Drake and Poor 1937a, p. 13 [Luzon]. 
Eteoneus virtutis: Drake and Poor 1937b, p. 402. 
Type: Holotype o 71 , macropterous; Mount Maquiling, Laguna Province, 
Luzon, Philippine Islands; USNM. 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

*Eteoneus visendus Drake and Poor 

Eteoneus visendus Drake and Poor 1937a, p. 14 [Luzon]. 

Type: Holotype o", macropterous; Imugan, Nueva Vizcaya Province, 
Luzon, Philippine Islands; Drake Coll. (USNM). 

Distribution: Philippine Islands (Luzon). 

Host Plant: Unrecorded. 

Genus EUAHANES Distant 

Euahanes Distant 1911a, p. 42. — Drake and RuhofF 1960a, p. 56. 
Type Species : Euahanes inflatus Distant. 

*Euahanes inflatus Distant 

Euahanes inflatus Distant 1911a, p. 43, fig. [Cape Province]. — Blote 
1945, p. 84. 
Type: Sex unknown; Grahamstown, Cape Colony, South Africa; So. 
Afr. Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

Genus EURYPHARSA Stal 

Eurypharsa Stal 1873, pp. 122, 133.— Monte 1939b, p. 69; 1941e. p. 
94.— Hurd 1946, p. 468.— Drake and RuhofF 1960a, p. 56. 
Type Species: Tingis nobilis Guerin-Meneville. 



214 U.S. NATIONAL MUSEUM BULLETIN 243 

*Eurypharsa championi Bergroth 

Eurypharsa championi Bergroth 1922, p. 151 [Brazil]. — Drake and Ham- 
bleton 1938b, p. 53.— Monte 1940b, p. 375.— Silva 1956, p. 27. 
Type: Sex unknown; Minas Gerais, Brazil; Helsin. Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

Eurypharsa farouki Silva 

Eurypharsa farouki Silva 1956, p. 27, fig. 8 [Brazil]. 

Type: Holotype 9, macropterous; Correntina, Estado da Bahia, Brazil; 
Instituto Biologico da Bahia. 

Distribution: Brazil. 

Host Plant: Found "sucking the foliage of 'cipo' (Malpighiaceae?)." 

Eurypharsa fenestrata Champion 

Eurypharsa jenestrata Champion 1898a, p. 44, pi. 3, figs. 17, 17a [Pan- 
ama]. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

*Eurypharsa nobilis (Guerin-Meneville) 

Tingis nobilis Guerin-Meneville 1844, p. 349 [Colombia; Bolivia]. 
Tingis circumdata Blanchard 1846, p. 219, pi. 29, fig. 7. — Stal 1873, p. 

134.— Lethierry and Severin 1896, p. 26. 
Eurypharsa nobilis: Stal 1873, p. 133 [Brazil]. — Lethierry and Severin 
1896, p. 23.— Champion 1898a, p. 44; 1898b, p. 63— Horvath 
1925b, p. 220 [Peru].— Drake 1931c, p. 226.— Drake and Hambleton 
1938b, p. 53. 
Type: Unknown. 

Distribution: Brazil; Colombia; Bolivia; Peru. 
Host Plant: Unrecorded. 

*Eurypharsa phyllophila Drake 

Eurypharsa phyllophila Drake 1922c, p. 359 [Brazil]. 
Type: Holotype cf, macropterous; Rio Guapore, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Eurypharsa quadrifenestrata Bergroth 

Eurypharsa quadrifenestrata Bergroth 1898, p. 9 [Brazil]; 1922, p. 151. — 
Drake and Hambleton 1938b, p. 53, pi. 9, fig. c. — Silva 1956, p. 30. 
Type: Sex unknown; Brazil; Helsin. Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 21 5 

Genus FROGGATTIA Froggatt 

Froggattia Froggatt 1901, p. 1601— Horvath 1902b, p. 605.— Drake 
and Runoff 1960a, p. 57. 
Type Species: Froggattia olivinia Froggatt. 

*Froggattia disticha Drake 

Froggattia disticha Drake 1947c, p. 120 [Queensland]. 

Type: Holotype 9, macropterous; Cedar Creek, Queensland, Australia; 
Drake Coll. (USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

*Froggattia hargreavesi Drake 

Froggattia hargreavesi Drake 1945, p. 100 [Uganda; Jasminum dichotomum]. 
Type: Holotype o 71 ; Kampala, Uganda; British Mus. 
Distribution: Uganda. 
Host Plant: Jasminum dichotomum. 

*Froggattia olivinia Froggatt 

Froggattia olivinia Froggatt 1901, p. 1601, fig. 14 [New South Wales; 
olive; Notelaea longifolia]. ■ — Drake and Ruhoff 1960a, p. 57. — Drake 
1961b, p. 108. 
Froggattia olivina Horvath 1902b, p. 605. — Hacker 1927, p. 25 
[Queensland]; 1928, p. 181 [Tasmania].— Silvestri 1934, p. 259 
[Olea europaea]. 
Type: Sex unknown; Camden Park, New South Wales, Australia; 
Hungarian Mus. 
Distribution: Australia (New South Wales; Tasmania; Queensland). 
Host Plants: Notelaea longifolia; Olea europaea. 
Note: See Drake and Ruhoff re authorship priority. 

Genus FURCILLIGER Horvath 

Furcilliger Horvath 1925a, p. 3.— Drake and Ruhoff 1960a, p. 57. 
Type Species: Furcilliger asperulus Horvath. 

Furcilliger asperulus Horvath 

Furcilliger asperulus Horvath 1925a, p. 4, fig. 2 [Queensland]. 
Type: Sex unknown; Herberton, Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Furcilliger cheesmanae Drake 

Furcilliger cheesmanae Drake 1954b, p. 10 [Papua]. — Drake and Davis 
1960, fig. 45.— Drake 1960, p. 368, fig. 20. 
Type: Holotype cf ; Kokoda, Papua; British Mus. 
Distribution: New Guinea (Papua). 
Host Plant: Unrecorded. 



216 U.S. NATIONAL MUSEUM BULLETIN 243 

*Furcilliger orestes Drake and Ruhoff 

Furcilliger orestes Drake and Runoff 1962d, p. 492 [New Guinea]. 

Type: Holotype 9, macropterous; Medang, New Guinea; Drake Coll. 
(USNM). 

Distribution: New Guinea. 

Host Plant: Unrecorded. 

Genus GABIROBIUS Schouteden 

Gabirobius Schouteden 1955b, p. 166.— Drake and Ruhoff 1960a, p. 58. 
Type Species: Gabirobius basilewskyi Schouteden. 

Gabirobius basilewskyi Schouteden 

Gabirobius basilewskyi Schouteden 1955b, p. 167 [Ruanda]; 1957c, p. 313. 
Type: Sex unknown; Gabiro, Ruanda; Cent. Afr. Mus. 
Distribution: Ruanda-Urundi. 
Host Plant: Unrecorded. 

Genus GALEATUS Curtis 

Galeatus Curtis 1833, p. 196— Stal 1874, p. 48.— Jakovlev 1880a, p. 
129.— Horvath 1897b, pp. 455-460 (key); 1906a, pp. 13, 49 (key).— 
Puton 1899, p. 40. — Lethierry and Severin 1896, p. 11.- — Oshanin 
1908, p. 421; 1912, p. 43.— Osborn and Drake 1916a, p. 236.— 
Stichel 1926, pp. 104, 109 (key); 1935, p. 349; 1938a, p. 406; 1960a, 
p. 295 (key); 1960b, p. 393; 1960c, p. 132.— Borner 1935, pp. 73, 76 
(key).— Guide 1938, pp. 243, 271 (key).— Hurd 1946, p. 478.— 
Kiritshenko 1951, pp. 241, 246 (key).— Singer 1952, p. 50.— Drake 
and Ruhoff 1960a, p. 58. 

Cadmilos Distant 1909a, p. 113; 1910a, p. 107.— Drake and Ruhoff 
1960a, p. 41. 
Type Species: Tingis spinifrons Fallen. 

Galeatus armatus Takeya 

Galeatus armatus Takeya 1931, p. 68, pi. 8, figs. 4, 5 [Japan]; 1951a, p. 7. 
Type: Holotype cf; Fukuoka, Kyushu, Japan; Kyushu Univ. 
Distribution: Japan. 
Host Plant: Unrecorded. 

*Galeatus cellularis Jakovlev 

Galeatus cellularis Jakovlev 1884, p. 121 [Turkestan]. — Horvath 1897b, 
pp. 455, 458; 1906a, p. 52.— Kiritshenko 1951, p. 247. 
Type: Holotype 9, macropterous; Turkestan, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 217 

*Galeatus clara Drake 

Galeatus clara Drake 1948e, p. 155 [China]. 

Type: Holotype o", macropterous; Shanghai; China; Drake Coll. 
(USNM). 

Distribution: China. 

Host Plant: Unrecorded. 

*Galeatus decorus Jakovlev 

Galeatus decorus J akovlev 1880a, pp. 131, 134 [Russia]. — Horvath 1905b, 
p. 566 [Hungary; Rumania]; 1906a, p. 54.— Lindberg 1934, p. 29 
[China].— Kiritshenko 1951, p. 247.— Stichel 1960a, p. 298 [Bul- 
garia]. 
Type: Holotype 9, brachypterous; Chvalynsk, Saratovskaja, Russia; 
Leningrad Inst. 
Distribution: U.S.S.R.; Rumania; Hungary; Bulgaria; China. 
Host Plant: Unrecorded. 

*Galeatus inermis (Jakovlev) 

Tingis inermis Jakovlev 1876b, p. 68, pi. 2, fig. 8 [Russia]. 
Galeatus inermis: Jakovlev 1880a, p. 130. — Horvath 1897b, pp. 455, 
457 [Turkestan]; 1906a, p. 51.— Oshanin 1908, p. 421 [Caucasus].— 
Blote 1945, p. 83.— Kiritshenko 1951, p. 246; 1959a, p. 105.— Stichel 
1960a, p. 295. 
Type: Sex unknown; Astrakhan, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Turkestan; Caucasus). 
Host Plant: Unrecorded. 

^Galeatus maculatus (Herrich-Schaeffer) Plate 46 

Tingis pyri (not Fabricius): Herrich-Schaeffer 1835, p. 58; 1836, p. 74, 
fig. 291.— Schilling 1838, p. 105. 

Tingis cristata (not Panzer): Schilling 1838, p. 106. 

Tingis maculata Herrich-Schaeffer 1838, p. 68. — Fieber 1844, p. 107, 
pi. 9, fig. 21; 1861, p. 130.— Mayr 1858, p. 572 [Hungary] —Frey- 
Gessner 1865, p. 233 [Switzerland].— Jakovlev 1869, p. 112 [Russia]. 

Tingis sub glob osa Herrich-Schaeffer 1838, p. 68. — Fieber 1844, p. 106, 
pi. 9, figs. 17-20 [Czechoslovakia; Austria].— Scholz 1847, p. 122. 

Campylosteira (Diciyonota) subglobosa: Kolenati 1856, p. 435 [Caucasus]. 

Galeatus maculatus: Stal 1874, p. 48. — Puton 1879c, p. 105 [France; 
Hieracium pilosella]. — Bolivar and Chicote 1879, p. 166 [Portugal; 
Spain].— Jakovlev 1880a, p. 131; 1880b, p. 101.— Reuter 1885, p. 
44. — Lethierry and Severin 1896, p. 11 [Algeria; Tunisia]. — d'Antes- 
santy 1890, p. 31.— Hiieber 1893, p. 322— Horvath 1897b, pp. 457, 
460 [Belgium; Netherlands; Germany; Transcaucasus] ; 1906a, p. 
53 [Potentilla]; 1923b, p. 190.— Muzik 1907, p. 57.— Reichensperger 
1920, p. 61.— Moroder Sala 1920, p. 12.— Stichel 1926, p. 109 
[Hieracium]; 1960a, p. 297, fig. 92 [Poland].— Jordan 1933, p. 229, 

CSl-552— 64 15 



218 U.S. NATIONAL MUSEUM BULLETIN 243 

figs. 25-30 [Statice armeria; Herniaria glabra; Erica vulgaris]. — Scholte 
1935, p. 31, fig. 6.— Guide 1938, p. 275, fig.— Hoberlandt 1942, p. 
125; 1943a, p. 118; 1943b, p. 123.— Blote 1945, p. 83.— Gonzalez 
1948, p. 50.— Kiritshenko 1951, p. 247.— Gomez-Menor 1955b, p. 
249.— Drake and Ruhoff 1962c, p. 135. 
Type: Unknown. 

Distribution: Spain; Portugal; France; Netherlands; Belgium; Switzer- 
land; Germany; Czechoslovakia; Austria; Poland; Hungary; U.S.S.R. 
(Transcaucasus; Caucasus); Algeria; Tunisia. 

Host Plants: Erica vulgaris; Herniaria glabra; Hieracium pilosella; Hieracium 
sp.; Potentilla sp.; Statice armeria. 

Note: Life history study [Jordan], 
*Galeatus major Puton 

Galeatus maculatus var. major Puton 1886a, p. 5 [Tunisia]. 
Galeatus major: Horvath 1897b, pp. 457, 460 [Algeria]; 1906a, p. 53. — 
Gomez-Menor 1955b, p. 249 [Spain].— Stichel 1960a, p. 298 [Sicily; 
Italy].— Drake and RuhofF 1962d, p. 489. 
Galeatus maroccanus Lindberg 1936a, p. 81, fig. 1 [Morocco]. 
Type: Sex unknown; Tunisia; Paris Mus. 
Distribution: Tunisia; Morocco; Algeria; Spain; Sicily; Italy. 
Host Plant: Unrecorded. 
*Galeatus scitulus Drake and Maa 

Galeatus scitulus Drake and Maa 1953, p. 90 [China; Chrysanthemum]. 
Type: Holotype 6% macropterous; Siyingpan, Yungan Hsien, Fukien, 
China; Drake Coll. (USNM). 
Distribution: China. 
Host Plant: Wild Chrysanthemum. 
*Galeatus scrophicus Saunders 

Galeatus scrophicus Saunders 1876, p. 103 [Greece]. — Lethierry and 
Severin 1896, p. 12 [Egypt].— Horvath 1897b, pp. 455, 457 [Armenia]; 
1906a, p. 52 [Crete; Algeria]; 1911c, p. 108.— Oshanin 1908, p. 422 
[Caucasus].— Lindberg 1948, p. 58 [Cyprus]; 1958a, p. 78 [Cape 
Verde]. — Priesner and Alfieri 1953, p. 63 [Echinops spinosa]. — Drake 
1954f, p. 7 [Sudan; Transvaal; Senegal; South-West Africa; sun- 
flower].— Gomez-Menor 1955b, p. 249 [Spain]; 1956a, p. 109, fig. 3 
(fig. only). — Hoberlandt 1955, p. 87 [Turkey; Iraq; Israel; Tadzhik; 
Transcaucasus; Artemisia; Astragalus; Plantago]. — Stichel 1960a, p. 
295 [Morocco; Canary Islands; Syria; Launaea spinosa], 
Galeatus semilucidus J akovlev 1884, p. 121 [Turkestan]. 
Galeathus [sic] semilucidus: Reuter 1891a, p. 26. 
Cadmilos retiarius Distant 1909a, p. 114 [India; Chrysanthemum]; 1910a, 

p. 107, fig. 53. 
Galeatus retiarius: Fletcher 1920, p. 263. 
Galeatus serophicus [sic]: Gonzalez 1948, p. 50. 



LACEBUGS OF THE WORLD 219 

Type: Sex unknown; Point Scophia, Greece; British Mus. 

Distribution: Spain; Greece; Crete; Cyprus; Israel; Iraq; Turkey; 
Syria; U.S.S.R. (Armenia; Transcaucasus; Caucasus; Turkestan; Tadzhik); 
India; Algeria; Egypt; Sudan; Senegal; South Africa (Transvaal; South- 
West Africa); Morocco; Cape Verde Islands; Canary Islands. 

Host Plants: Artemisia sp.; Astragalus sp.; Chrysanthemum sp.; Echinops 
spinosa; Launaea spinosa; Plantago sp.; sunflower. 
*Galeatus sinuatus (Herrich-Schaeffer) Plate 45 

Tingis sinuata Herrich-Schaeffer 1838, p. 68, pi. 126, fig. 394 [Hungary]; 
1850, p. 162.— Fieber 1844, p. 108, pi. 9, fig. 22; 1861, p. 129 — 
Scholz 1847, p. 122.— Mayr 1858, p. 572 [Austria] .—Walker 1873a, 
p. 179. 
Galeatus sinuatus: Stal 1874, p. 48. — Hiieber 1893, p. 324 [Hieracium 
pilosella].—YLoxvkth. 1897b, pp. 455, 457; 1906a, p. 52.— Oshanin 
1908, p. 422 [Germany].— Stichel 1926, p. 109; 1960a, p. 295, fig. 
85 [Bulgaria].— Kiritshenko 1951, p. 247.— Putshkov 1960, p. 302 
[Phlomis tuber osa], 
Tingis per spicuus J akovlev 1875a, p. 162 [Russia]. 
Galeatus bvevispinus [sic] Jakovlev 1880a, p. 130. — Puton 1886b, p. 33. 
Galeatus brevispinus Jakovlev 1880a, p. 131. — Reuter 1890a, p. 249. — 
Horvath 1897b, pp. 455, 457; 1906a, p. 52.— Puton 1899, p. 40 — 
Kiritshenko 1951, p. 246. 
Galeatus per spicuus: Jakovlev 1880a, p. 130. 
Type: Hungary; sex and deposition of type unknown. 
Distribution: Hungary; Germany; Austria; Poland; Bulgaria; U.S.S.R. 
Host Plants: Hieracium pilosella; Phlomis tuberosa. 

Note: Life history study [Putshkov]. Synonymy of brevispinus Jakovlev 
fide I. M. Kerzhner. 

*Galeatus spinifrons (Fallen) Plate 44 

Tingis spinifrons Fallen 1807, p. 38 [Sweden; Solanum dulcamara]; 1829, 
p. 148.— Germar 1817, fasc. 13, tab. 18— Herrich-Schaeffer 1835, 
p. 58; 1838, p. 67, pi. 130, figs, a-c, g-h— Schilling 1838, p. 106 — 
Amyot and Serville 1843, p. 297.— Fieber 1844, p. 104, pi. 9, figs. 
6-12 [England; Czechoslovakia; Austria; Italy; Artemisia campestris]; 
1861, p. 129— Scholz 1847, p. 121.— Mayr 1858, p. 572— Frey- 
Gessner 1865, p. 233 [Switzerland].— Garbiglietti 1869, p. 275 — 
Horvath 1874b, p. 432 [Hungary]. 

Galeatus spinifrons: Curtis 1833, p. 196. — Spinola 1837, p. 164. — Stal 
1874, p. 48.— Jakovlev 1880a, p. 131; 1893, p. 294.— Reuter 1882b, 
p. 118 [Calluna; Hieracium].— Hiieber 1893, p. 320.— Horvath 1897b, 
pp. 456, 459 [Turkestan]; 1906a, p. 53 [Siberia; Medicago]. — Muzik 
1907, p. 57, fig. 4.— Oshanin 1908, p. 423 [Japan].— Lindberg 1919, 
p. 42 [Antennaria dioica]. — Reichensperger 1920, p. 61. — Sahlberg 



220 U.S. NATIONAL MUSEUM BULLETIN 243 

1920, p. 82 [Calluna vulgaris].— -Drake 1923, p. 104.— Stichel 1926, 
p. 109; 1960a, p. 297, fig. 89 [Denmark].— Esaki 1932, p. 1634, 
fig.; 1954, p. 235, fig. 608.— Kato 1933, pi. 31, fig. 5.— Scholte 
1935, p. 31 [Netherlands].— Fukui 1938, pi. 50, fig. 34— Kirit- 
shenko 1951, p. 247, fig. 328.— Takeya 1951a, p. 6 [Taiwan; Ar- 
temisia vulgaris indica; Chrysanthemum indicum]; 1953d, p. 168. — 
Putshkov 1960, p. 303 [Helichrysum arenarium\ — Drake and Ruhoff 
1962c, p. 135; 1962d, p. 490, fig. 2. 
Tingis affinis Herrich-Schaeffer 1835, p. 58; 1836, p. 73, pi. 290 [Ger- 
many]; 1838, p. 67; 1850, p. 162.— Fieber 1844, p. 106, pi. 9, figs. 
13-16; 1861, p. 129.— Sahlberg 1848, p. 136 [Finland].— Flor 1860, 
p. 366 [Estonia; Latvia]. — Garbiglietti 1869, p. 275. — Gredler 
1870, p. 76. 
Galeathus [sic] angusticollis Reuter 1874, p. 563. 
Galeatus affinis: Stal 1874, p. 48. 
Galeathus [sic] spinifrons: Reuter 1874, p. 562. 

Galeatus angusticollis: Jakovlev 1880a, p. 130 [Russia]. — Hiieber 1893, 
p. 321— Horvath 1897b, pp. 456, 459; 1906a, pp. 50, 52— Muzik 
1907, p. 56.— Oshanin 1908, p. 423 [Belgium].— Stichel 1926, p. 
109; 1938a, p. 406 [Poland]; 1960a, p. 296.— Hoberlandt 1942, p. 
125.— Kiritshenko 1951, p. 247.— Drake and Ruhoff 1962a, p. 163, 
pi. 43. 
Sphaerocysta peckhami Ashmead 1887, p. 156 [Wis.]. 
Galeatus peckhami: Van Duzee 1889, p. 5 [Canada]; 1917b, p. 216 
[Ont.; Man.].— Uhler 1904, p. 362.— Parshley 1916b, p. 105 [N.H.]; 
1917a, p. 15 [Maine]; 1917b, p. 55; 1923b, p. 703.— Osborn and 
Drake 1916a, p. 236, pi. 10, fig. c [Mass.].— Drake 1918d, p. 86; 
1922d, p. 66 [N.Y.; Aster macrophyllus; Eupatorium]; 1922e, pp. 105— 
110, pi. 5 [Minn.; Mich.]; 1923a, p. 102; 1928b, p. 101— Blatchley 
1926, p. 453, fig. 98.— Proctor 1946, p. 74.— Bailey 1951, p. 39; 
1959, p. 64. 
Galeatus uhleri Horvath 1923a, p. 108 [N. Mex.].— Hurd 1946, p. 478 

[Colo.; Aha.]. 
Copium artemisifolium Shinji 1938, p. 316 [Artemisia vulgaris]. — Takeya 

1951a, p. 19. 
Galeatus angusticollis subsp. uhleri: Drake and Ruhoff 1962a, p. 163. 
Type: Unknown. 

Distribution: England; Netherlands; Belgium; Germany; Switzerland; 
Czechoslovakia; Hungary; Austria; Italy; Poland; Sweden; Finland; 
Norway; Denmark; U.S.S.R. (Latvia; Estonia; Turkestan; Siberia); 
Japan; Taiwan; U.S. (Maine; Wis.; N.H.; N.Y.; Mass.; Minn.; Mich.; 
Colo.; Utah; N. Mex.); Canada (Ont.; Man.; Alta.). 

Host Plants: Antennaria dioica; Artemisia campestris; Artemisia vulgaris; 
Artemisia vulgaris indica; Aster macrophyllus; Calluna vulgaris; Calluna sp.; 



LACEBUGS OF THE WORLD 221 

Chrysanthemum indicum; Eupatorium sp.; Helichrysum arenarium; Hieracium sp.; 
Medicago sp.; Solarium dulcamara. 
Note: Life history studies [Putshkov; Drake 1922e; Bailey]. 

Genus GARGAPHIA Stal 

Monanthia (Gargaphia) Stal 1862, p. 324. 

Gargaphia: Stal 1873, pp. 119, 124.— Uhler 1886, p. 22.— Lethierry and 
Severin 1896, p. 13.— Champion 1897, p. 9 (key).— Banks 1910, 
p. 56.— Van Duzee 1916, p. 25; 1917b, p. 217.— Osborn and Drake 
1916a, p. 233 (key).— Drake 1917b, pp. 227-228 (key ) .—Gibson 
1919b, pp. 187-201 (key).— Blatchley 1926, p. 472 (key).— Costa 
Lima 1936, p. 126.— Monte 1939b, p. 69; 1941e, p. 95.— Hurd 1946, 
p. 479.— Drake and Runoff 1960a, p. 58. 
Type Species: Monanthia {Phyllontochila) patricia Stal. 

*Gargaphia acmonis Drake and Hambleton 

Gargaphia acmonis Drake and Hambleton 1945, p. 366 [Peru; Colombia]. 

Type: Holotype d", macropterous; Tin go Maria, Peru; Drake Coll. 
(USNM). 

Distribution: Peru; Colombia; Surinam. 

Host Plant: Hibiscus esculentus. 

*Gargaphia albescens Drake 

Gargaphia albescens Drake 1917b, p. 228 [Calif.].— Gibson 1919b, p. 197. 
Type: Holotype 9, macropterous; Sacramento, California, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Calif.). 
Host Plant: Unrecorded. 

*Gargaphia amorphae (Walsh) 

Tingis amorphae Walsh 1864, p. 409 [Amorpha fruticosa]. — Walker 1873a, 

p. 180 [111.]. 
Gargaphia amorphae: Uhler 1886, p. 22.— Bergroth 1892, p. 264 — 
Smith 1910, p. 149 [NJ.].— Osborn and Drake 1916a, p.235.— McAtee 
1917b, p. 79.— Gibson 1919b, p. 195 [N.C.; W. Va.].— Drake 1925c, 
p. 37 [Miss.]; 1928b, p. 102 [N.Y.] .— Blatchley 1926, p. 474 [Ind.].— 
Froeschner 1944, p. 668 [Mo.].— Hurd 1946, p. 480 [Iowa]. 
Gargaphia tiliae var. amorphae: Van Duzee 1917b, p. 217. 
Type: Lectotype 9, macropterous; U.S.; USNM. 

Distribution: U.S. (N.Y.; Pa.; N.J.; Va.; N.C.; Miss.; Ohio; Mo.; 
Ind.; W. Va.; 111.; Iowa). 
Host Plant: Amorpha fruticosa. 

Note: Walsh failed to state the type locality for this species, and the 
type specimen bears only the label "Collection C. V. Reiley." We believe 
that the type locality is Illinois. 



222 U.S. NATIONAL MUSEUM BULLETIN 243 

*Gargaphia angulata Heidemann 

Gargaphia angulata Heidemann 1899b, p. 301 [Ala.; D.C.; Mo.; bean]. — 
Chittenden 1900, p. 32, fig. 8.— Smith 1910, p. 149, fig. 63 (fig. 
only). — Osborn and Drake 1916a, p. 233, fig. 6 [Tenn.; Mass.; Pa.; 
N.C.].— Van Duzee 1917b, p. 218 [Ohio] .— Parshley 1917b, p. 56 
[Conn.]; 1923b, p. 704, pi. 17, fig. 7.— McAtee 1917b, p. 79 [Ceanothus 
americanus]; 1923, p. 147 [Va.].— Gibson 1919b, pp. 189, 191 [Ariz.; 
N.J.].— Barber 1922a, p. 17.— Blatchley 1926, p. 474, fig. 106; pi. 4, 
fig. 7 [N.Y.].— Drake 1928b, p. 102 [Solanum].— Froeschner 1944, 
p. 669.— Hurd 1946, p. 480 [Colo.; Minn.].— Bailey 1951, p. 42.— 
Drake and Ruhoff 1962a, p. 161. 
Gargaphia undulata Heidemann 1901, p. 493 (nom. nud). — Van Duzee 
1917b, p. 218. 
Type: Holotype d", macropterous; Auburn, Alabama, U.S.; USNM. 
Distribution: U.S. (Mass.; Conn.; N.Y.; Pa.; N.J.; Md.; D.C.; Va.; 
N.C; S.C.; Ala.; Miss.; Tenn.; Ohio; Mich.; Mo.; Kans.; Minn.; Colo.; 
Ariz.). 
Host Plants: Ceanothus americanus; Solanum sp.; bean. 

*Gargaphia argillacea Monte 

Gargaphia argillacea Monte 1943d, p. 263, fig. 1 [Brazil]. 
Type: Holotype cf; Guarapuava, Parana, Brazil; Monte Coll. (Mus. 
Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Gargaphia arizonica Drake and Carvalho 

Gargaphia arizonica Drake and Carvalho 1944, p. 43 [Tex.; Ariz.; 
N. Mex.; Mexico]. 
Type: Holotype o 71 , macropterous; Huachuca Mountains, Arizona, 
U.S.; Drake Coll. (USNM). 
Distribution: U.S. (Ariz.; N. Mex.; Tex.); Mexico. 
Host Plant: Unrecorded. 

*Gargaphia balli Drake and Carvalho 

Gargaphia balli Drake and Carvalho 1944, p. 42 [Ariz.; N. Mex.]. 
Type: Holotype o 71 , macropterous; Naco, Arizona, U.S.; Drake Coll. 
(USNM). 
Distribution: U.S. (Ariz.; N. Mex.). 
Host Plant: Unrecorded. 

*Gargaphia bimaculata Parshley 

Gargaphia bimaculata Parshley 1920a, p. 271 [Fla.]. — Blatchley 1926, p. 

475 [thistle]. 
Gargaphia binotata [sic]: Drake 1925c, p. 37. 



LACEBUGS OF THE WORLD 223 

Type: Holotype 9, macropterous; Biscayne Bay, Florida, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Fla.). 
Host Plant: Thistle. 

Gargaphia boliviana Monte 

Gargaphia boliviana Monte 1947b, p. 431 [Bolivia]. 
Type: Holotype o 71 ; Cochabamba, Bolivia; Monte Coll. (Mus. Nacional). 
Distribution: Bolivia. 
Host Plant: Unrecorded. 

*Gargaphia brunfelsiae Monte 

Gargaphia brunfelsiae Monte 1938c, p. 64, fig. [Brazil; Brunjelsia]; 1938d> 
p. 389 [Argentina; Brunjelsia hopeana]; 1940c, p. 193, fig. 26; 1940e> 
p. 295.— Singh 1953, p. 118. 
Type: Holotype c? ; Belo Horizonte, Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil; Argentina. 
Host Plants: Brunjelsia hopeana; Brunjelsia sp. 

*Gargaphia comosa Monte 

Gargaphia comosa Monte 1941a, p. 374, fig. 2 [Brazil]. 
Type: Holotype cf; Catanduva, Sao Paulo, Brazil; Monte Coll. (Mus. 
Nacional) . 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Gargaphia concursa Drake 

Gargaphia concursa Drake 1930b, p. 25 [Brazil]. — Monte 1939b, p. 69 
[Croton; Annonaceae]. — Silva 1956, p. 30. 

Gargaphia concusa [sic]: Drake 1947a, p. 2. 
Type: Holotype cf , macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plants: Croton sp.; Annonaceae. 

*Gargaphia condensa Gibson 

Gargaphia condensa Gibson 1919b, pp. 189, 197 [Ariz.; Dahlia parryi]. — 

Drake 1922b, p. 41. 
Gargaphia carinata Gibson 1919b, p. 199. 
Type: Holotype 9, macropterous; Santa Rita Mountains, Arizona, 
U.S.; USNM. 
Distribution: U.S. (Ariz.). 
Host Plant: Dahlia parryi. 

*Gargaphia costalimai Monte 

Gargaphia costa-limai Monte 1 938b, p. 130, fig. 1 [Brazil; Croton]. 
Type: Holotype o 71 ; Belo Horizonte, Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Croton sp. 



224 U.S. NATIONAL MUSEUM BULLETIN 243 

*Gargaphia crotonae Drake and Hambleton 

Gargaphia crotonae Drake and Hambleton 1938a, p. 66 [Brazil; Croton 
arucurana]. — Monte 1939b, p. 69 [Solarium]; 1940c, p. 193. 
Type: Holotype o 71 , macropterous; Atibaia, Sao Paulo, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Croton urucurana; Solanum sp. 

*Gargaphia deceptiva (Drake and Bruner) 

Gelchossa deceptiva Drake and Bruner 1924a, p. 150 [British Guiana; 

Trinidad]. 
Leptopharsa deceptiva: Monte 1942a, p. 95 [Ipomoea]. 
Gargaphia deceptiva: Hurd 1946, p. 480. 
Type: Holotype 9, macropterous; Demarara R., British Guiana; Drake 
Coll. (USNM). 
Distribution: British Guiana; Trinidad; Venezuela; Colombia. 
Host Plant: Ipomoea sp. 

*Gargaphia decoris Drake 

Gargaphia decoris Drake 1931b, p. 514 [Brazil]; 1935, p. 20. — Drake and 
Poor 1939b, p. 98 [Solanum]. — Monte 1939b, p. 69 [Croton urucurana]; 
1940b, p. 376. 
Gargaphia decorie [sic]: Monte 1938b, p. 131 [Argentina]. 
Type: Holotype cf, macropterous; Belo Horizonte, Minas Gerais, 
Brazil; Cornell Univ. 
Distribution: Brazil; Argentina. 
Host Plants: Croton urucurana; Solanum sp. ("probably auricalatum") . 

*Gargaphia differitas Drake (emendation) 

Gargaphia differatis Drake 1935, p. 19 [Paraguay]. — Monte 1939b, p. 70 
[Brazil]. — Silva 1956, p. 30 [Dalechampia ficifolia]. 
Type: Holotype d"; S. Bernardina, Paraguay; Vienna Mus. 
Distribution: Paraguay; Brazil. 
Host Plant: Dalechampia ficifolia. 

*Gargaphia dissortis Drake 

Gargaphia dissortis Drake 1930a, p. 3 [Brazil]. — Monte 1942a, p. 94 
[Argentina]. 
Type: Holotype o*; Chapada, Brazil; Amer. Mus. 
Distribution: Brazil; Argentina. 
Host Plant: Unrecorded. 

*Gargaphia flexuosa (Stal) 

Monanthia (Phyllontocheila) flexuosa Stal 1858, p. 61 [Brazil]. 
Monanthia {Gargaphia) flexuosa: Stal 1862, p. 324. 
Monanthia flexuosa: Walker 1873a, p. 192. 



LACEBUGS OF THE WORLD 225 

Gargaphia flexuosa: Stal 1873, p. 124. — Drake and Hambleton 1934, 
p. 450 [Cissampelos glaberrima; Cissampelos tamoides}. — Drake and Poor 
1937d, p. 309, pi. 36, fig. 10.— Silva 1956, p. 30, fig. 9 [Aristolochia sp.]. 

Type: Holotype $; Brazil; Stockholm Mus. 

Distribution: Brazil. 

Host Plants: Aristolochia sp.; Cissampelos glaberrima; Cissampelos tamoides 

(?)• 

*Gargaphia formosa (Stal) 

Monanthia {Phyllontocheila) formosa Stal 1858, p. 61 [Brazil]. 

Monanthia {Gargaphia) formosa: Stal 1862, p. 324. 

Monanthia formosa: Walker 1873a, p. 192. 

Gargaphia formosa: Stal 1873, p. 125. — Drake and Poor 1937d, p. 310. — 

Monte 1940e, p. 294, fig. 8 [Solanum concinnum]. 
Tingis formosa (not Stal) Goldi 1886, pp. 234-241, figs. 
Type: Holotype 9; Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Solanum concinnum. 
Note: Morphology and life study [Goldi]. 

*Gargaphia gentilis Van Duzee 

Gargaphia gentilis Van Duzee 1923, p. 141 [Carmen Island; Solanum 
hindsianum]. 
Type: Holotype o 71 ; Carmen Island, Gulf of California, Mexico; Cal. 
Acad. 
Distribution: Mexico (Carmen Island). 
Host Plant: Solanum hindsianum. 

*Gargaphia gracilenta Drake 

Gargaphia gracilenta Drake 1928i, p. 74 [Argentina]. — Monte 1940e, 
p. 296. 
Type: Holotype cT, macropterous; Cordoba, Alta Gracia, Argentina; 
Drake Coll. (USNM). 
Distribution: Argentina. 
Host Plant: Unrecorded. 

Gargaphia holoxantha Monte 

Gargaphia holoxantha Monte 1942a, p. 93, fig. 1 [Brazil], 
Type: Holotype o 71 ; Campos, Estado do Rio, Brazil; Monte Coll. (Mus. 
Nacional) . 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Gargaphia implicata Drake and Hambleton 

Gargaphia concur sa var. implicata Drake and Hambleton 1940, p. 535 
[Brazil]. 



226 U.S. NATIONAL MUSEUM BULLETIN 243 

Gargaphia implicata: Drake and Hambleton 1944b, p. 127 [Argentina; 
Paraguay]. 
Type: Holotype d\ macropterous; Vicosa, Est. de Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil; Argentina; Paraguay. 
Host Plant: Unrecorded. 
*Gargaphia insularis Van Duzee 

Gargaphia insularis Van Duzee 1923, p. 141 [Angel de la Guardia Island; 
Parosela emoryi]. 
Type: Holotype c? 1 ; Angel de la Guardia Island, Gulf of California, 
Mexico; Cal. Acad. 
Distribution: Mexico (Angel de la Guardia Island). 
Host Plant: Parosela emoryi. 
Gargaphia interrogationis Monte 

Gargaphia interrogationis Monte 1941d, p. 99, fig. 4 [Costa Rica]. 
Type: Holotype d"; Pacora, Costa Rica; Monte Coll. (Mus. Nacional). 
Distribution: Costa Rica. 
Host Plant: Unrecorded. 
*Gargaphia iridescens Champion 

Gargaphia iridescens Champion 1897, p. 10, pi. 2, figs. 1,1a [Mexico]. — 
Banks 1910, p. 56 [Tex.].— Van Duzee 1917b, p. 217 [Colo.]; 1923, 
p. 141 [San Marcos Island].— Gibson 1919b, pp. 189, 197 [Calif.; 
Ariz.; N. Mex.; Ambrosia; Solanum; Malva; sand nettle]. — Torre- 
Bueno 1942, p. 131 [hollyhock].— Gibson and Carrillo 1959, p. 19. 
Type: Sex unknown; Juarez, Mexico; British Mus. 
Distribution: Mexico (San Marcos Island); U.S. (Tex.; N. Mex.; Ariz.; 
Calif.; Colo.). 
Host Plants: Ambrosia sp.; Malva sp.; Solanum sp.; sand nettle; hollyhock. 
Note: "Maternal solicitude" [Torre-Bueno] . 
*Gargaphia jucunda Drake and Hambleton 

Gargaphia jucunda Drake and Hambleton 1942b, p. 80 [Panama]. 
Type: Holotype c\ macropterous; Panama City, Panama; Drake Coll. 
(USNM). 
Distribution: Panama. 
Host Plant: Unrecorded. 

*Gargaphia lanei Monte 

Gargaphia lanei Monte 1940b, p. 376 [Brazil]. — Drake and Hambleton 

1944b, p. 127. 
Gargaphia limata Drake and Poor 1940, p. 228. 
Gargaphia limitata [sic]: Monte 1941e, pp. 98, 171. 
Type: Holotype cf; Belo Horizonte, Minas Gerais, Brazil; Monte Coll. 
(Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 227 

*Gargaphia lunulata (Mayr) 

Monanthia lunulata Mayr 1865, p. 441 [Brazil]; 1866, p. 163. — Walker 

1873a, p. 193. 
Gargaphia lunulata: Stal 1873, p. 124. — Berg 1884, p. 190 [Argentina; 
Passiflora caerulea]. — Pennington 1921, p. 20. — Drake and Hambleton 
1934, p. 450 [Canavalia ensiformis; Phaseolus vulgaris; Hibiscus esculentus; 
Glycine max; Passiflora violacea; Ricinus communis; Pyrus communis; 
Gossypium arbor eum; Sligmaphyllon; Cassia fistula; Urena lobata; Manihot 
utilissima; Chorisia speciosa']. — Drake 1935, p. 19 [Paraguay]. — Costa 
Lima 1936, p. 127 [Doliciws lablab; Phaseolus lunatus; Cassia imperialist. — 
Bosq 1937, p. 128 [Psidium; Uruguay] .—Monte 1937a, p. 36, fig. 6; 
1938b, p. 131; 1938d, p. 388.— Drake and Poor 1938b, p. 109 — 
Araujo Mendonca 1949, pp. 183-187, figs.— Silva 1956, p. 32, fig. 
[Rosa; Abelmoschus esculentus; Euphorbia heterophylla; %ornia diphylla; 
Cnidoscolus; Meibomia adscendens]. 
Gargaphia lasciva Gibson 1919b, p. 198.— Fiuza 1946, pp. 338, 340, 341, 
343. 
Type: Rio de Janeiro, Brazil; sex and deposition of type unknown. 
Distribution: Argentina; Colombia; Brazil; Paraguay; Uruguay. 
Host Plants: Abelmoschus esculentus; Canavalia ensiformis; Cassia fistula; 
Cassia imperialis; Chorisia speciosa; Cnidoscolus sp.; Dolichos lablab; Euphorbia 
heterophylla; Glycine max; Gossypium arboreum; Hibiscus esculentus; Manihot 
utilissima; Meibomia adscendens; Passiflora caerulea; Passiflora violacea; Phaseolus 
lunatus; Phaseolus vulgaris; Psidium sp.; Pyrus communis; Ricinus communis; 
Rosa sp.; Stigmaphyllon sp.; Urena lobata; ^prnia diphylla. 

*Gargaphia manni Drake and Hurd 

Gargaphia manni Drake and Hurd 1945b, p. 130 [Bolivia]. 
Type: Holotype cf , macropterous; Waschi, Bolivia; USNM. 
Distribution: Bolivia. 
Host Plant: Unrecorded. 

*Gargaphia mexicana Drake 

Gargaphia mexicana Drake 1922b, p. 40 [Mexico]. — Hurd 1946, p. 480 
[Tex.]. 
Type: Holotype 9, macropterous; Tampico, Mexico; USNM. 
Distribution: Mexico; U.S. (Tex.). 
Host Plant: Unrecorded. 

*Gargaphia mirabilis Monte 

Gargaphia mirabilis Monte 1938a, p. 292, fig. 32 [Venezuela; Croton 
pungens] . 
Type: Holotype cf; Antimano, districto Federal, Venezuela; Monte 
Coll. (Mus. Nacional). 
Distribution: Venezuela. 
Host Plant: Croton pungens. 



228 U.S. NATIONAL MUSEUM BULLETIN 243 

*Gargaphia munda (Stal) 

Monanthia {Phyllontocheila) munda Stal 1858, p. 60 [Brazil]. 
Monanthia (Gargaphia) munda: Stal 1862, p. 324. 
Monanthia munda: Walker 1873a, p. 192. 

Gargaphia munda: Stal 1873, p. 124. — Distant 1888, p. lxxxiii. — Penning- 
ton 1921, p. 20 [Argentina].— Drake 1922b, p. 41; 1935a, p. 20 — 
Drake and Hambleton 1934, p. 450 [Brunfelsia; Solanum]; 1944b, p. 
127.— Araujo Silva 1936, p. 7.— Bosq 1937, p. 129 [Brunfelsia hope- 
ana].— Drake and Poor 1937d, p. 310, pi. 36, fig. 12 [Peru]; 1938b, 
p. 109.— Monte 1944b, p. 466. 
Monanthia lineifera Walker 1873a, p. 194. 

Leptostyla lineifera: Champion 1898b, p. 60. — Distant 1902b, p. 356. 
Gargaphia magna Gibson 1919b, p. 194 [Paraguay]. 
Type: Holotype 9; Brazil; Stockholm Mus. 
Distribution: Brazil; Argentina; Paraguay; Peru. 
Host Plants: Brunfelsia hopeana; Brunfelsia sp.; Solanum sp. 

*Gargaphia neivai Drake and Poor 

Gargaphia neivai Drake and Poor 1940, p. 227 [Paraguay; Ecuador]. — ■ 
Drake and Hambleton 1945, p. 365. 
Type: Holotype cf, macropterous; Horqueta, Paraguay; Drake Coll. 
(USNM). 
Distribution: Ecuador; Paraguay. 
Host Plant: Unrecorded. 

*Gargaphia nexilis Drake and Hambleton 

Gargaphia nexilis Drake and Hambleton 1940, p. 537 [Brazil]. 

Type: Holotype 9, macropterous; Campinas, Sao Paulo, Brazil; Drake 
Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Gargaphia nigrinervis Stal 

Gargaphia nigrinervis Stal 1873, p. 125 [Colombia]. — Champion 1897, 
p. 10, pi. 1, figs. 13, 13a [Panama]. — Osborn and Drake 1915b, p. 
536 [Guatemala].— Gibson 1919b, p. 192.— Drake 1926b, p. 378; 
1932, p. 100; 1935, p. 20 [Venezuela].— Drake and Poor 1937d, p. 
311.— Monte 1940d, p. 104 [Solanum] .—Drake and Hambleton 1945, 
p. 365 [Peru].— Drake and Cobben 1960, p. 87, fig. 86 [Aruba; Cu- 
rasao; Bonaire; Jatropha wrens']. 
Type: Holotype o" ; Bogota, Colombia; Stockholm Mus. 
Distribution: Colombia; Venezuela; Peru; Panama; Guatemala; 
Netherlands Antilles (Aruba; Curacao; Bonaire). 
Host Plants: Jatropha urens, Solanum sp. 



LACEBUGS OF THE WORLD 229 

*Gargaphia nigrinervis var. irnpedita Drake and Hambleton 

Gargaphia nigrinervis var. irnpedita Drake and Hambleton 1944b, p. 127 
[Colombia]. 
Type: Holotype cf, macropterous; Rio Frio, Colombia; Drake Coll. 
(USNM). 
Distribution: Colombia. 
Host Plant: Unrecorded. 

*Gargaphia nociva Drake and Hambleton 

Gargaphia nociva Drake and Hambleton 1940, p. 536 [Brazil]. 

Type: Holotype o", macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Gargaphia obliqua Stal 

Gargaphia obliqua Stal 1873, p. 124 [Brazil]. — Drake and Hambleton 
1934, p. 450 [Serjania].— Drake 1936b, p. 700 [Argentina].— Drake 
and Poor 1937d, p. 310, pi. 36, fig. 9.— Monte 1947a, p. 236 — 
Drake and Davis 1960, fig. 33. 
Gargaphia stigma Monte 1946b, p. 284. 
Type: Holotype o 71 ; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil; Argentina. 
Host Plant: Serjania sp. 

*Gargaphia opacula Uhler 

Gargaphia opacula Uhler 1893b, p. 263 [Calif.]; 1894b, p. 278 [Magda- 

lena]. — Townsend 1894, p. 313 (N. Mex.; eggplant]. — Van Duzee 

1914, p. 11 [willow].— Gibson 1919b, pp. 189, 199 [Utah; Kans.; 

Dahlia spinosa; Solanum melongena]. 

Type: Holotype 9, macropterous; Argus Mountains, California, U.S.; 

USNM. 

Distribution: U.S. (Calif.; N. Mex.; Utah; Kans.); Mexico (Magda- 
lena Island). 
Host Plants: Dahlia spinosa; Solanum melongena; willow. 

*Gargaphia opima Drake 

Gargaphia opima Drake 1931b, p. 513 [Peru]. — Drake and Hambleton 

1945, p. 365 [Colombia; Canavalia ensiformis]. 
Gargaphia inca Monte 1943, p. 105, fig. 1; 1947a, p. 232, fig. 
Type: Holotype cf, macropterous; Miriatiriani, Cam. del Pichis, 
Peru; Cornell Univ. 
Distribution: Peru; Colombia; Bolivia. 
Host Plant: Canavalia ensiformis. 



230 U.S. NATIONAL MUSEUM BULLETIN 243 

*Gargaphia oreades Drake 

Gargaphia oreades Drake 1928i, p. 75 [Argentina]; 1935, p. 20 [Brazil]; 
1936b, p. 700. — Bosq 1937, p. 129 [Solanum nigrum; Solarium capsicas- 
trum].— Monte 1938d, p. 388. 
Type: Holotype cf, macropterous; Buenos Aires, Argentina; Drake 
Coll. (USNM). 
Distribution: Argentina; Brazil. 
Host Plants: Solanum capsicastrum; Solanum nigrum. 

*Gargaphia oregona Drake and Hurd 

Gargaphia oregona Drake and Hurd 1945b, p. 131 [Oreg.]. 

Type: Holotype cf, macropterous; Corvallis, Oregon, U.S.; Drake 
Coll. (USNM). 

Distribution: U.S. (Oreg.). 

Host Plant: Unrecorded. 

Gargaphia panamensis Champion 

Gargaphia panamensis Champion 1897, p. 10, pi. 1, figs. 14, 14a [Pana- 
ma].— Gibson 1919b, pp. 190, 193. 
Type: Sex unknown; Caldera, Chiriqui, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

*Gargaphia paraguayensis Drake and Poor 

Gargaphia paraguayensis Drake and Poor 1940, p. 229 [Paraguay]. 

Type: Holotype cf, macropterous; Horqueta, Paraguay; Drake Coll. 
(USNM). 

Distribution: Paraguay. 

Host Plant: Unrecorded. 

*Gargaphia patria (Drake and Hambleton) 

Leptopharsa patria Drake and Hambleton 1938a, p. 58 [Brazil; Bathysa 

stipulata] . 
Gargaphia patria: Drake 1942a, p. 1. — Silva 1956, p. 34 [Sickingia 
tinctoria\ . 
Type: Holotype o", macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plants: Bathysa stipulata; Sickingia tinctoria. 

*Gargaphia patricia (Stal) 

Monanthia (Phyllontochila) patricia Stal 1862, p. 324 [Mexico]. 
Monanthia {Gargaphia) patricia: Stal 1862, p. 324. 
Monanthia patricia: Walker 1873a, p. 191. 

Gargaphia patricia: Stal 1873, p. 125. — Champion 1897, p. 9, pi. 1, 
figs. 12, 12a [Panama; Guatemala]. — Gibson 1919b, p. 196. — Drake 
1926b, p. 377; 1936b, p. 700 [Argentina]; 1948h, p. 76.— Drake and 



LACEBUGS OF THE WORLD 231 

Poor 1937d, p. 309 [Colombia].— Monte 1940d, p. 104 [Venezuela; 
Euphorbiaceae] ; 1941d, p. 98 [Costa Rica]. 
Gargaphia biolleyi Blote 1945, p. 85 (nom. nud. of Montandon). 
Type: Sex unknown; Mexico; Stockholm Mus. 

Distribution: Mexico; Costa Rica; Guatemala; Panama; Colombia; 
Venezuela; Argentina. 

Host Plant: Euphorbiaceae. 
*Gargaphia paula Drake 

Gargaphia paula Drake 1939f, p. 68 [Canal Zone]. — Drake and Hamble- 
tonl945, p. 366 [Peru]. 
Type: Holotype o 71 , macropterous; Barro Colorado Island, Panama 
Canal Zone; Drake Coll. (USNM). 
Distribution: Panama (Canal Zone); Brazil; Peru. 
Host Plant: Unrecorded. 
*Gargaphia penningtoni Drake 

Gargaphia penningtoni Drake 1928i, p. 75 [Argentina]; 1935, p. 20 
[Paraguay]. — Drake and Hambleton 1934, p. 450 [Sida acuta; Abutilum; 
Acalypha]. — Bosq 1937, p. 129. — Monte 1938d, p. 388 [Cajanus indicus; 
Brazil]; 1939b, p. 70 [Sida; Phaseolus]; 1940c, p. 193 [Wissadula; 
Croton lobatus]. — Drake and Poor 1939b, p. 98. 
Gargaphia sabpilosa (not Berg): Monte 1943d, p. 264, fig. 2. 
Type: Holotype cf , macropterous; Buenos Aires, Argentina; Drake Coll. 
(USNM). 
Distribution: Argentina; Paraguay; Brazil. 

Host Plants: Abutilum sp.; Acalypha sp.; Cajanus indicus; Croton lobatus; 
Phaseolus sp.; Sida acuta; Sida sp.; Wissadula sp. 
*Gargaphia schulzei Drake 

Gargaphia schulzei Drake 1954d, p. 76 [Paraguay]. 
Type: Holotype o", macropterous; Asuncion, Paraguay; Drake Coll. 
(USNM). 
Distribution : Paraguay. 
Host Plant: Unrecorded. 
*Gargaphia seorsa Drake and Hambleton 

Gargaphia seorsa Drake and Hambleton 1945, p. 365 [Peru]. 
Type: Holotype c 71 , macropterous; Aguaytia, Peru; Drake Coll. (USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 
*Gargaphia serjaniae Drake and Hambleton 

Gargaphia serjaniae Drake and Hambleton 1938b, p. 57, pi. 10, fig. G 
[Brazil; Serjania]. 
Type: Holotype o 71 , macropterous; Pedro Leopoldo, Minas Gerais, 
Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Serjania sp. 



232 U.S. NATIONAL MUSEUM BULLETIN 243 

*Gargaphia shelfordi Drake and Hambleton 

Gargaphia shelfordi Drake and Hambleton 1944a, p. 95 [Mexico]. 

Type: Holotype o 71 , macropterous; Victoria, Mexico; Drake Coll. 
(USNM). 

Distribution: Mexico. 

Host Plant: Unrecorded. 

*Gargaphia socorrona Drake 

Gargaphia socorrona Drake 1954d, p. 77 [Revilla Gigedo]. 

Type: Holotype o", macropterous; Socorro Island, Revilla Gigedo 
Islands, Mexico; Drake Coll. (USNM). 

Distribution: Mexico (Revilla Gigedo Islands). 

Host Plant: Unrecorded. 

*Gargaphia solani Heidemann 

Gargaphia solani Heidemann 1914, p. 136, fig. 1 [Mo.; Tex.; Okla.; 

Va.; Solanum carolinense; Solanum elaeagnifolium; eggplant; potato; 

coffee weed]. — Fink 1915, p. 1, figs. 1, 2 [D.C.; Md.; Cassia]. — Osborn 

and Drake 1916a, p. 235, fig. 7 [Ohio].— Somes 1916, p. 42.— Gibson 

1919b, pp. 189, 192 [Ariz.; Solanum tuberosum; Solanum melongena; 

cotton; Amphiachyris; Salvia pitcheri]. — Parshley 1923b, p. 704. — 

McAtee 1923, p. 147.— Blatchley 1926, p. 474 [N.G.].— Fenton 1934, 

p. 199 [S.C.; Mexico; Salvia azurea; hollyhock]. — Froeschner 1944, 

p. 668— Hurd 1946, p. 480 [Canada].— Bailey 1951, p. 43 [Conn.; 

Lycopersicum esculentum; Gossypium herbaceum]. — -Ash 1954, p. 185 [Ind.]. 

Type: Holotype 9, macropterous; Lavaca County, Texas, U.S.; USNM. 

Distribution: U.S. (Conn.; N.J.; Pa.; Md.; D.C.; Va.; N.C.; S.G.; Ala.; 

Tenn.; Miss.; Mo.; Ark.; Ohio; Ind.; 111.; Kans.; Iowa; Okla.; Tex.; 

Ariz.); Canada (B.C.); Mexico. 

Host Plants: Amphiachyris sp.; Cassia sp.; Gossypium herbaceum; Lycopersi- 
cum esculentum; Salvia azurea; Salvia pitcheri; Solanum carolinense; Solanum 
elaeagnifolium; Solanum melongena; Solanum tuberosum; Solanum sp.; hollyhock. 
Note: Life history studies [Fink; Bailey]; predators [Fink; Bailey]. 

Gargaphia sororia Hussey Plate 50 

Gargaphia sororia Hussey 1957, p. 175 [Fla.]. 
Type: Holotype cf ; Gadsden County, Florida, U.S.; Florida Univ. 
Distribution: U.S. (Fla.). 
Host Plant: Unrecorded. 

*Gargaphia subpilosa Berg 

Gargaphia subpilosa Berg 1879a, p. 42 [Argentina; Mikania auricularis]; 
1879b, p. 136.— Pennington 1921, p. 20.— Drake 1931b, p. 512 
[Brazil]; 1948j, p. 432.— Fenton 1934, p. 199 [Bolivia; cotton; peas].— 
Bosq 1937, p. 129 [Helianthus annus; Ipomoea bonariensis; Phaseolus 
vulgaris; Prunus cerasus; Prunus persica; Pyrus communis; Solanum tube- 



LACEBUGS OF THE WORLD 233 

rosum].— Monte 1937a, p. 36 [Wissadula]; 1938d, p. 387; 1940f, pp. 
301-306.— Drake and Poor 1938b, p. 109. 
Gargaphia iridescens (not Champion): Bergroth 1922, p. 149. — Monte 

1938d, p. 389 [Ambrosia tenuifolia], 
Gargaphia bergi Monte 1940f, p. 306, fig. 1. 
Type: Sex unknown; Argentina; La Plata Mus. 
Distribution: Argentina; Bolivia; Brazil. 

Host Plants: Ambrosia tenuifolia; Gossypium sp.; Helianthus annus; Ipomoea 
bonariensis; Mikania auricularis; Phaseolus vulgaris; Prunus cerasus; Primus 
per ska; Pyrus communis; Solanum tuberosum; Wissadula sp.; peas. 

*Gargaphia tiliae (Walsh) Figure 4a 

Tingis tiliae Walsh 1864, p. 408 [basswood; cherry]. — -Walker 1873a, p. 

180. 
Gargaphia fasciata Stal 1873, p. 125 [111.].— Uhler 1886, p. 22 [Mexico].— 

Wirtner 1904, p. 202 [Pa.].— Gibson 1919b, p. 195 [Ala.]. 
Gargaphia tiliae: Uhler 1886, p. 22. — Provancher 1886, p. 159 [Can- 
ada].— Bergroth 1892, p. 264.— Van Duzee 1894, p. 181 [N.Y.]; 
1912, p. 323.— Gillette and Baker 1895, p. 57 [Colo.].— Torre-Bueno 
1910, p. 31.— Smith 1910, p. 149 [N.J.].— Osborn and Drake 1915a, 
p. 506 [Ohio]; 1916a, p. 234.— Parshley 1917b, p. 56 [N.H.; Conn.; 
Mass.]; 1922b, p. 11 [S. Dak.]; 1923b, p. 704.— Weiss 1919b, p. 165 
[Va.; Kans.].— Gibson 1919b, pp. 189, 194 [N.C.; Md.; Tenn.; Mo.; 
Wis.; Tilia pubescens; Prunus serotina]. — Hussey 1922a, p. 22 [Mich.; 
Tilia americana].— Barber and Weiss 1922, p. 13. — Barber 1922a, p. 
17.— Drake 1922b, p. 41; 1926b, p. 378; 1928b, p. 102.— McAtee 
1923, p. 147. — Blatchley 1926, p. 473 [Ind.; Cercis canadensis; syca- 
more].— Felt 1933, p. 49.— Froeschner 1944, p. 668.— Hurd 1946, p. 
480 [Fla.; Ont.; Nebr.].— Bailey 1951, p. 45 [Maine; Vt.; N.B.; 
Que.; Ariz.; Tilia]; 1959, p. 64.— Ash 1954, p. 185. 
Type: Holotype cf , macropterous; Rock Island [assumed Illinois], U.S.; 
USNM. 

Distribution: U.S. (Maine; Vt.; N.H.; Mass.; Conn.; N.Y.; Pa.; N.J 
Md ; Va.; W. Va. N.C.; S.C.; Ga.; Fla.; Ala.; Miss.; Ky.; Tenn.; Mo 
Ohio; Ind.; 111.; Mich.; Wis.; Nebr.; Kans.; Minn.; Iowa; S. Dak.; Colo 
Ariz.); Mexico; Canada (Ont.; N.B.; Que.). 

Host Plants: Cercis canadensis; Prunus serotina; Tilia americana; Tilia 
pubescens; Tilia sp. 

Notes: Life history studies [Weiss; Barber and Weiss; Bailey]; taken at 
light [McAtee]. 

There is no locality label on type specimen, only the label reading 
"Collection C. V. Reiley" and in his article Walsh gives only the following 
information "Very abundant near Rock Island and on the basswood." 



6S1-552— 64- 



234 U.S. NATIONAL MUSEUM BULLETIN 243 

*Gargaphia torresi Costa Lima 

Gargaphia torresi Costa Lima 1922, p. 112 [Brazil]; 1930, p. 84 [Croton; 
Triumfetta; Ipomoea]; 1936, p. 127. — Bondar 8, p. 244, figs. 

[Phaseolus].— Drake 1931b, p. 512 [cotton]; 1935, p.. 20.— Drake and 
Hambleton 1934, p. 450 [maize; sweet potato].— Fenton 1934, p. 199 
[Argentina].— Bosq 1937, p. 130.— Monte 1937a, p. 35, fig. 7; 1939b, 
p. 70 [Paraguay]. — Fiuza 1946, pp. 338, 341 [Phaseolus vulgaris; 
Phaseolus lunatus]. — Silva 1956, p. 34, fig. 11 [Gossypium; Canavalia 
obtusijolia; Zinnia elegans; Hibiscus syriacus"]. 
Type: Sex unknown; Brazil; Ent. Agr. Brazil. 
Distribution: Brazil; Argentina; Paraguay; Bolivia. 
Host Plants: Canavalia obtusijolia; Croton sp.; Gossypium sp.; Hibiscus 
syriacus; Ipomoea sp.; Phaseolus lunatus; Phaseolus vulgaris; Phaseolus sp.; Sida 
sp.; Triumfetta sp.; Zinnia elegans; sweet potato; maize. 
*Gargaphia trichoptera Stal 

Gargaphia trichoptera Stal 1873, p. 125 [Colombia]. — Champion 1898b, 
p. 58, pi. 2, fig. 5.— Drake and Poor 1937d, p. 311.— Monte 1947b, 
p. 432 [Peru]. 
Type: Sex unknown; Bogota, Colombia; Stockholm Mus. 
Distribution: Colombia; Peru; Brazil. 
Host Plant: Unrecorded. 
*Gargaphia tuthilli Drake and Carvalho 

Gargaphia tuthilli Drake and Carvalho 1944, p. 42 [Colo.]. 
Type: Holotype d\ macropterous; Mesa Verde, Colorado, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Colo.). 
Host Plant: Unrecorded. 

*Gargaphia valerioi Drake 

Gargaphia valerioi Drake 1941a, p. 144 [Costa Rica]. 
Type: Holotype d", macropterous; La Gloria, Costa Rica; USNM. 
Distribution: Costa Rica. 
Host Plant: Unrecorded. 
*Gargaphia vanduzeei Gibson 

Gargaphia vanduzeei Gibson 1919b, p. 198 [Costa Rica].— Hurd 1946, p. 
480 [Cuba]. 
Type: Holotype cf; Costa Rica; deposition of type unknown. 
Distribution: Costa Rica; Cuba. 
Host Plant: Unrecorded. 
Note: Type damaged [Gibson]. 
*Gargaphia venosa Drake and Poor 

Gargaphia venosa Drake and Poor 1942, p. 301 [Argentina]. 
Type: Holotype <?; Resistencia, Chaco, Argentina; Argentina Mus. 
Distribution: Argentina. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 235 

Genus GITAVA Drake 

Gitava Drake 1948e, p. 149.— Drake and RuhofT 1960a, p. 59. 
Type Species: Tigava ugandana Drake. 

Gitava basilewskyi Schouteden 

Gitava basilewskyi Schouteden 1953c, p. 130 [Congo]. 
Type: Sex unknown; Bondo Mabe, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Gitava benoiti Schouteden 

Gitava benoiti Schouteden 1953c, p. 131 [Congo]. 
Type: Sex unknown; Basoko, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Gitava ? dispar Schouteden 

Gitava ? dispar Schouteden 1957a, p. 88 [Malagasy]. 
Type: Sex unknown; Madagascar; Paris Mus. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

Gitava fusca Schouteden 

Gitava jusca Schouteden 1957a, p. 87 [Malagasy]. 
Type: Sex unknown; Madagascar; Paris Mus. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

Gitava madagascariensis Schouteden 

Gitava madagascariensis Schouteden 1957a, p. 86 [Malagasy]. 
Type: Sex unknown; Madagascar; Paris Mus. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

Gitava petila Drake 

Gitava petila Drake 1954g, p. 663, fig. 4 [Ethiopia]. 
Type: Sex unknown; Mulu, Ethiopia; British Mus. 
Distribution: Ethiopia. 
Host Plant: Unrecorded. 

*Gitava procera Drake 

Gitava procera Drake 1958c, p. 321 [Malagasy]. 
Type: Holotype 9; Ankazoabo, Hera, Madagascar; Madagascar Sci. 
Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 



236 U.S. NATIONAL MUSEUM BULLETIN 243 

*Gitava ugandana (Drake) 

Tigava ugandana Drake 1942a, p. 11 [Uganda]. 

Gitava Uganda [sic]: Drake 1948e, p. 149. — Drake and Runoff 1960a, 

p. 59. 
Gitava ugandana: Schouteden 1953c, p. 130 [Congo]. — Drake 1956a, 
p. 5. ^ 
Type: Holotype d", macropterous; Uganda; Drake Coll. (USNM). 
Distribution: Uganda; Congo. 
Host Plant: Unrecorded. 

*Gitava ugandana subsp. pallens Schouteden 

Gitava ugandana subsp. pallens Schouteden 1953c, p. 130 [Congo]. 
Type: Sex unknown; Lubero, Kivu, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Genus GYALOTINGIS Drake 

Gyalotingis Drake 1960, p. 376. 
Type Species: Gyalotingis gressitti Drake. 

*Gyalotingis gressitti Drake 

Gyalotingis gressitti Drake 1960, p. 377, fig. 26 [New Guinea]. 
Type: Holotype 9; Wisselmeren, Enarotadi, Netherlands New Guinea; 
Bishop Mus. 
Distribution: New Guinea (Netherlands). 
Host Plant: Unrecorded. 

Genus GYMNOTINGIS Hacker 

Gymnotingis Hacker 1928, p. 181.— Drake and Ruhoff 1960a, p. 59. 
Type Species: Gymnotingis serrulata Hacker. 

Gymnotingis serrulata Hacker 

Gymnotingis serrulata Hacker 1928, p. 182, pi. 23, fig. 14 [Queensland]. 
Type: Sex unknown; Cairns District, Australia; Queensland Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Genus HABROCHILA Horvath 

Habrochila Horvath 1912a, p. 353.— Drake and Ruhoff 1960a, p. 59. 
Type Species: Habrochila placida Horvath. 

*Habrochila africana Drake 

Habrochila africana Drake 1948e, p. 153 [Uganda; Kenya; Natal; 
Transvaal; Malagasy; Lantana]. — Schouteden 1953a, p. 104 [Congo]. 
Type: Holotype o 71 ; Kampal, Uganda; British Mus. 
Distribution: Uganda; Kenya; Congo; South Africa (Natal; Trans- 
vaal) ; Malagasy Republic. 
Host Plant: Lantana sp. 



LACEBUGS OF THE WORLD 237 

*Habrochila chinensis Drake 

Habrochila chinensis Drake 1947d, p. 231 [China; tea]. 
Type: Holotype 9, macropterous; Meitonhsein, China; Drake Coll. 
(USNM). 
Distribution: China. 
Host Plant: Tea (?). 

^Habrochila clivosa Drake 

Habrochila clivosa Drake 1954b, p. 11 [Natal]. 
Type: Holotype o*, brachypterous; Port Shepstone, Natal, South 
Africa; British Mus. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

Habrochila darthula (Kirkaldy) 

Galeatus darthula Kirkaldy 1902, p. 297 [Ceylon; Barleria cristata], — 

Distant 1903b, p. 131, fig. 94 [Barleria strigosa]. 
Habrochila darthula: Drake and Ruhoff 1961c, p. 151 [India]. 
Type: Sex unknown; Peradeniya, Ceylon; British Mus. 
Distribution: Ceylon; India. 
Host Plants: Barleria cristata; Barleria strigosa. 

*Habrochila ghesquierei Schouteden 

Habrochila ghesquierei Schouteden 1953a, p. 105 [Congo; Uganda; 
Ruanda; coffee]; 1957c, p. 313. 
Type: Unknown. 

Distribution: Congo; Uganda; Ruanda-Urundi. 
Host Plant: Coffee. 

Habrochila horvathi Schouteden 

Habrochila horvathi Schouteden 1923, p. 92 [Congo]; 1953a, p. 105. 
Type: Sex unknown; Watsa, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Habrochila iolana Drake 

Habrochila iolana Drake 1955a, p. 81 [Malagasy]. 

Type: Holotype o 71 , macropterous; Perinet, Analamasotra Province, 
Madagascar; Drake Coll. (USNM). 

Distribution: Malagasy Republic. 

Host Plant: Unrecorded. 

*Habrochila laeta Drake 

Habrochila laeta Drake 1954b, p. 12 [India], 
Type: Holotype o"; Bengalore, Madras, India; British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 



238 U.S. NATIONAL MUSEUM BULLETIN 243 

*Habrochila monticola Horvath 

Habrochila monticola Horvath 1929, p. 319 [Natal]. — Drake and Slater 
1955, p. 49.— Drake 1961c, p. 129. 
Type: Holotype 9; Weenen, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

*Habrochila natalana Drake 

Habrochila natalana Drake 1956f, p. 426 [Natal]. 
Type: Holotype o 71 ; Albert Falls, Umgeni River, E. of Pietermaritz- 
berg, Natal, South Africa; Lund Zool. Inst. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

*Habrochila placida Horvath 

Habrochila placida Horvath 1912a, p. 354 [Congo; cacao]. — Schouteden 
1923, p. 92; 1953a, p. 105 [Uganda].— Mayne and Ghesquiere 
1934, p. 20 [coffee ].— Caray on 1960, p. 119. 
Type: Sex unknown; Kondue, Kasai, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Uganda. 
Host Plants: Coffee; cacao. 

Genus HAEDUS Distant 

Haedus Distant 1904, p. 432.— Drake 1953b, p. 93.— Drake and Ruhoff 

1960a, p. 59. 
Hormisdas Distant 1910b, p. 59.— Drake and Ruhoff 1960a, p. 60. 
Type Species: Haedus clypeatus Distant. 

*Haedus bellus (Drake) 

Hormisdas bella Drake 1948g, p. 176 [Sierra Leone]. 
Haedus bellus: Drake 1953b, p. 93. — Schouteden 1953c, p. 129 [Congo]. 
Type: Holotype 9, macropterous; Makump, Sierra Leone; Drake 
Coll. (USNM). 
Distribution: Sierra Leone; Congo. 
Host Plant: Unrecorded. 

*Haedus burungus Schouteden 

Haedus burunga Schouteden 1953c, p. 128 [Congo]. 
Type: Sex unknown; Burunga, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Haedus clypeatus Distant 

Haedus clypeatus Distant 1904, p. 432, pi. 8, fig. 12 [Cape Province]. 

Type: Sex unknown; Wynberg, Cape Colony, South Africa; So. Afr. 
Mus. 

Distribution: South Africa (Cape Province). 

Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 239 

Haedus decellei Schouteden 

Haedus decellei Schouteden 1957b, p. 219 [Congo; Croton]. 
Type: Sex unknown; Yangambi, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Croton sp. 

*Haedus diversitas (Drake) (emendation) 

Harmidas [sic] diver satis Drake 1927d, p. 309 [Malagasy]. 
Haedus diver sitatus [sic]: Drake 1953b, p. 93. 

Type: Holotype 9, macropterous; Madagascar; Drake Coll. (USNM). 

Distribution: Malagasy Republic. 

Host Plant: Unrecorded. 

* Haedus elongatus (Drake) 

Hormistes [sic] elongata Drake 1948g, p. 177 [Tanganyika]. 
Haedus elongatus: Drake 1953b, p. 93. 
Type: Sex unknown; Arusha-Ju, Africa; Hungarian Mus. 
Distribution: Tanganyika; South Africa (Natal). 
Host Plant: Unrecorded. 

*Haedus javancus Drake 

Haedus javancus Drake 1953b, p. 94 [Java]. 
Type: Sex unknown; Bandoeng, Preonger, Java; British Mus. 
Distribution: Greater Sunda Islands (Java). 
Host Plant: Unrecorded. 

♦Haedus lectus (Drake) 

Hormisdas lectus Drake 1937a, p. 387, fig. lc [India; "Elephantia"]. 

Haedus lectus: Drake 1953b, p. 93. 
Type: Holotype o 71 ; Bombay, India; Vienna Mus. 
Distribution: India. 
Host Plant: "Elephantia." 

*Haedus oios Drake and Ruhoff 

Haedus oios Drake and Ruhoff 1962c, p. 139, fig. 2 [Southern Rhodesia]. 

Type: Holotype o 71 , macropterous; Salisbury, Southern Rhodesia; 
Drake Coll. (USNM). 

Distribution: Southern Rhodesia. 

Host Plant: Unrecorded. 

*Haedus otiosus Drake 

Haedus otiosus Drake 1953d, p. 215 [Northern Rhodesia]. 

Type: Holotype 9, macropterous; N'Dola, Northern Rhodesia; Drake 
Coll. (USNM). 

Distribution: Northern Rhodesia. 

Host Plant: Unrecorded. 



240 U.S. NATIONAL MUSEUM BULLETIN 243 

*Haedus pallens Schouteden 

Haedus bellus subsp. pallens Schouteden 1953c, p. 129 [Congo; Annona 

chrysophylla; Annona muricata]. 
Haedus pallens: Drake and Ruhoff 1962a, p. 156. 
Type: Sex unknown; Ituri, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Kenya. 
Host Plants: Annona chrysophylla; Annona muricata. 

*Haedus pictus (Distant) 

Hormisdas pictus Distant 1910b, p. 60, pi. 1, figs. 1 a, b [Luzon]. — Drake 

and Poor 1937a, p. 16. 
Haedus pictus: Drake 1953b, p. 93. 
Type: Sex unknown; Manila, Philippine Islands; British Mus. 
Distribution: Philippine Islands (Luzon). 
Host Plant: Lantana sp. 

*Haedus polulus Drake 

Haedus polulus Drake 1953d, p. 216 [Senegal]. 
Type: Holotype cf; Bambey, Senegal; British Mus. 
Distribution: Senegal. 
Host Plant: Unrecorded. 

*Haedus sidae (Drake and Poor) 

Hormisdas sidae Drake and Poor 1939c, p. 206 [Sida; Uganda]. 

Haedus sideus [sic]: Drake 1953b, p. 93; 1954e, p. 7 [Congo]. 

Haedus sidus [sic]: Drake 1956a, p. 5. 
Type: Holotype o 71 ; Kampala, Uganda; British Mus. 
Distribution: Uganda; Congo. 
Host Plant: Sida sp. 

*Haedus vicarius (Drake) 

Hormisdas vicarius Drake 1927a, p. 56 [Luzon; Larat]; 1936a, p. 145 
[India; Urena lobata]; 1938b, p. 197 [China]; 1948g, p. 176.— Drake 
and Poor 1937a, p. 16 [Negros]; 1937b, p. 403 [Urena lobata sinuata]. 
Haedus vicarius: Drake 1953b, p. 93. 
Type: Holotype d", macropterous; Larat Island; Drake Coll. (USNM). 
Distribution: Tanimbar Islands (Larat); Philippine Islands (Luzon; 
Negros); China; India. 
Host Plants: Urena lobata; Urena lobata sinuata. 

*Haedus villiersi (Drake) 

Haedus villiers [sic] Drake 1953b, p. 93 (nom. nud.). 

Hormisdas villiersi Drake 1954a, p. 236 [Ivory Coast]. 
Type: Holotype o 71 ; Bouake, Ivory Coast; Inst. Fr. Afr. N. 
Distribution: Ivory Coast; Congo. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 241 

Genus HEBETINGIS Drake 

Hebetingis Drake 1960a, p. 360. 
Type Species: Hebetingis adeia Drake. 

Hebetingis adeia Drake 

Hebetingis adeia Drake 1960, p. 360, fig. 13 [New Guinea]. 
Type: Holotype cf; Asaro-Chimbu Divide, E. Highlands, Northeast 
New Guinea; Bishop Mus. 
Distribution: New Guinea (Northeast). 
Host Plant: Unrecorded. 

Genus HEGESIDEMUS Distant 

Hegesidemus Distant 1911b, p. 270. — Drake and Ruhoff 1960a, p. 59. 
Type Species: Hegesidemus eliyanus Distant. 

*Hegesidemus elegantulus (Distant) 

Teleonemia ? elegantula Distant 1909c, p. 167, pi. 10, figs. 3, 3a [Borneo]. 

Compseuta elegantula: Bergroth 1921, p. 105. — Drake 1948f, p. 202. 

Hegesidemus elegantulus: Drake and Ruhoff 1961c, p. 145. 
Type: Sex unknown; Kuching, Borneo; British Mus. 
Distribution: Greater Sunda Islands (Borneo). 
Host Plant: Unrecorded. 

*Hegesidemus eliyanus Distant 

Hegesidemus eliyanus Distant 1911b, p. 270 [Ceylon; Strobilanthes]. 
Type: Sex unknown; Nuwera Eliya, Ceylon; British Mus. 
Distribution: Ceylon. 
Host Plant: Strobilanthes sp. 

*Hegesidemus otiosus Drake Plate 29 

Hegesidemus otiosus Drake 1953d, p. 221 [India]. 
Type: Holotype d*; Nilgiri Hills, India; British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 

^Hegesidemus pauliani Drake 

Hegesidemus pauliani Drake 1957e, p. 402 [Reunion]. 
Type: Holotype cf; Bebour, Reunion; Madagascar Sci. Inst. 
Distribution: Mascarene Islands (Reunion). 
Host Plant: Unrecorded. 

Genus HENRIKUS Drake 

Henrikus Drake 1955e, p. 280.— Drake and Ruhoff 1960a, p. 60. 
Type Species: Henrikus schouiedeni Drake. 



242 U.S. NATIONAL MUSEUM BULLETIN 243 

*Henrikus schoutedeni Drake 

Henrikus schoutedeni Drake 1955e, p. 282, fig. 1 [Congo]. 
Type: Holotype o 71 ; Kmango, Kimbau, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Genus HESPEROTINGIS Parshley 

Hesperotingis Parshley 1917a, p. 21; 1923b, p. 707.— Van Duzee 1917b, 
p. 818.— Blatchley 1926, p. 493 (key).— Hurd 1946, p. 446.— Drake 
and RuhofT 1960a, p. 60. 
Type Species: Hesperotingis antennata Parshley. 

*Hesperotingis antennata Parshley 

Hesperotingis antennata Parshley 1917a, p. 21, fig. 2 [N.J.; N.Y.; Pa.; 

Conn.]; 1917b, p. 57; 1923b, p. 707.— Barber 1922a, p. 17; 1922b, 

p. 23.— Olsen 1923, p. 163.— McAtee 1923, p. 145 [D.C.].— Blatchley 

1926, p. 493, fig. 118; 1928, p. 5 [Fla.].— Drake 1928b, p. 102 — 

Froeschner 1944, p. 670 [Mo.].- Torre-Bueno 1946, p. 94.— Bailey 

1951, p. 22 [N.H.; Mass.; Andropogon]. 

Type: Holotype 9, macropterous; Lakehurst, New Jersey, U.S.; USNM. 

Distribution: U.S. (N.H.; Mass.; Conn.; N.Y.; N.J.; Pa.; D.C; Fla.; 

Mo.). 

Host Plants: Andropogon sp.; "beaten from Spanish moss hanging from 
limbs of Quercus rubra 1 " [Blatchley 1928]. 

*Hesperotingis antennata var. borealis Parshley 

Hesperotingis antennata var. borealis Parshley 1917a, p. 24 [N.H.]; 1917b, 
p. 57.— Froeschner 1944, p. 670 [Mo.].— Hurd 1946, p. 447 [D.C].— 
Bailey 1951, p. 22. 
Type: Sex unknown; Hampton, New Hampshire, U.S.; MCZ. 
Distribution: U.S. (N.H.; Mo.; D.C). 
Host Plant: Unrecorded. 

*Hesperotingis duryi (Osborn and Drake) 

Melanorhopala duryi Osborn and Drake 1916b, p. 15, fig. 3d [Tex.]; 

1917a, p. 159, fig. 2; pi. 8, fig. e— Parshley 1917a, p. 19. 
Hesperotingis duryi: Hurd 1946, p. 447 [Fla.]. 
Type: Holotype 9, brachypterous; Brownsville, Texas, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Tex.; Fla.). 
Host Plant: Unrecorded. 

*Hesperotingis duryi var. confusa Drake 

Hesperotingis duryi var. confusa Drake 1922b, p. 48 [Tex.]. 
Type: Holotype 9, brachypterous; Marfa, Texas, U.S.; USNM. 
Distribution: U.S. (Tex.). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 243 

*Hesperotingis floridana Drake 

Hesperotingis floridana Drake 1928f, p. 4 [Fla.]. 
Type: Holotype 9, macropterous; Florida, U.S.; USNM. 
Distribution: U.S. (Fla.). 
Host Plant: Unrecorded. 

*Hesperotingis fuscata Parshley 

Hesperotingis juscata Parshley 1917a, p. 24 [Colo.]. — Hurd 1946, p. 447 
[Kans.]. 
Type: Holotype 9, brachypterous ; Golden, Colorado, U.S.; USNM. 
Distribution: U.S. (Colo.; Kans.). 
Host Plant: Unrecorded. 

^Hesperotingis illinoiensis Drake 

Hesperotingis illinoiensis Drake 1918d, p. 88 [111.]. — Blatchley 1926, p. 
494.— Bailey 1951, p. 24 [Conn.]. 
Type: Holotype 9, macropterous; Palos Park, Illinois, U.S.; Drake Coll. 
(USNM). 
Distribution: U.S. (111.; Ind.; Conn.). 
Host Plant: Unrecorded. 

^Hesperotingis mississippiensis Drake 

Hesperotingis mississippiensis Drake 1928f, p. 4 [Miss.]. 
Type: Holotype d\ brachypterous; Charleston, Mississippi, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Miss.; Fla.). 
Host Plant: Unrecorded. 

*Hesperotingis occidentalis Drake 

Hesperotingis occidentalis Drake 1922b, p. 49 [Colo.]. 
Type: Holotype 9, brachypterous; Colorado, U.S.; Drake Coll. 
(USNM). 
Distribution: U.S. (Colo.; Calif.; Idaho). 
Host Plant: Unrecorded. 

Genus HOLOPHYGDON Kirkaldy 

Holophygdon Kirkaldy 1908b, p. 364.— Drake and Poor 1937a, p. 18; 
1939c, p. 207.— Drake and Ruhoff 1960a, p. 60. 
Type Species: Holophygdon melanesica Kirkaldy. 
*Holophygdon melanesica Kirkaldy Plate 49 

Holophygdon melanesica Kirkaldy 1908b, p. 364, pi. 4, figs. 10, 11 [Fiji]. — 
Drake and Poor 1943, p. 205 [Viti Levu].— Drake and Davis 1960, 
fig. 58. 
Type: Holotype 9; Fiji Islands; Hawaii. Sugar Plant. Assn. 
Distribution: Fiji Islands [Viti Levu]. 
Host Plant: Unrecorded. 



244 U.S. NATIONAL MUSEUM BULLETIN 2 43 

*Holophygdon melanesica var. fusca Drake and Poor 

Holophygdon melanesica var. fusca Drake and Poor 1943, p. 205 [Viti 
Levu; Ovalau; Raphidophora merrillii]. 
Type: Holotype cf ; Wainiloka, Ovalau, Fiji Islands; Bishop Mus. 
Distribution: Fiji Islands (Viti Levu; Ovalau). 
Host Plant: Raphidophora merrillii. 

Genus HORVATHULA Schouteden 

Horvathula Schouteden 1957c, p. 317. 
Type Species: Sankisia uniseriata Horvath. 

*Horvathula uniseriata (Horvath) 

Sankisia uniseriata Horvath 1929, p. 319 [Nigeria; Cordia millenii]. 
Monanthia uniseriata: Drake 1954g, p. 658 [Ethiopia; Cordia abyssinica]. 
Horvathula uniseriata: Schouteden 1957c, p. 318 [Ruanda; Cordia 
chrysocarpa] . 
Type: Holotype o 71 ; Ibadan, Nigeria; British Mus. 

Distribution: Nigeria; Ruanda-Urundi; Ethiopia; Kenya; Congo; 
Uganda. 
Host Plants: Cordia abyssinica; Cordia chrysocarpa; Cordia millenii. 

Genus HOV ATLAS Schouteden 

Hovatlas Schouteden 1957a, p. 85. — Drake and Ruhoff 1960a, p. 60. 
Type Species: Hovatlas elegantulus Schouteden. 

Hovatlas elegantulus Schouteden 

Hovatlas elegantulus Schouteden 1957a, p. 85 [Malagasy]. 
Type: Holotype 9; Madagascar; Paris Mus. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

Genus HURDCHILA Drake 

Hurdchila Drake 1953b, p. 92.— Drake and Runoff 1960a, p. 60. 
Type Species: Jannaeus togularis Drake and Poor. 

*Hurdchila mira (Drake and Poor) 

Jannaeus mirus Drake and Poor 1936a, p. 442 [Hainan]. — Drake 1937b, 

p. 387 [Ceylon]. 
Hurdchila mira: Drake 1953b, p. 93. 
Type: Holotype d", macropterous; Ta Han, Hainan Island; Drake Coll. 
(USNM). 
Distribution: China (Hainan Island); Ceylon. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 245 

*Hurdchila togularis (Drake and Poor) 

Jannaeas togularis Drake and Poor 1936a, p. 441 [Hainan]. 
Hurdchila torgularis [sic]: Drake 1953b, p. 93. 
Type: Holotype cf, macropterous; Hainan Island, China; Drake Coll. 
(USNM). 
Distribution: China (Hainan Island). 
Host Plant: Unrecorded. 

Genus HYALOCHITON Horvath 

Galeatus (Hyalochiton) Horvath 1905b, p. 566. 

Hyalochiton: Horvath 1906a, pp.13, 48 (key) .— Oshanin 1908, p. 420; 

1912, p. 43— Stichel 1926, p. 109; 1935, p. 348; 1938a, p. 406; 

1960a, p. 293; 1960b, p. 393; 1960c, p. 132.— Guide 1938, pp. 243, 

270.— Kiritshenko 1951, pp. 241, 246.— Drake and Ruhoff 1960a, p. 

60. 
Type Species: Galeatus komarqffii J akovlev. 

*Hyalochiton colpochilus (Horvath) Plate 43 

Galeatus colpochilus Horvath 1897b, pp. 456, 459 [Spain]. 
Hyalochiton colpochilus: Horvath 1906a, p. 49. — Lindberg 1932a, p. 41 
[Morocco].— Stichel 1938a, p. 406 [France]; 1960a, p. 293 [Greece; 
Phlomis lynchitis]. — -Wagner 1958, p. 240 [Betonica officinalis]. 
Type: Sex unknown; Ciudad Real, Spain; Hungarian Mus. 
Distribution: Spain; France; Morocco. 
Host Plants: Betonica officinalis; Phlomis lynchitis. 

Hyalochiton colpochilus var. consimilis (Horvath) 

Galeatus consimilis Horvath 1905a, p. 273 [Spain]. 

Hyalochiton colpochilus var. consimilis: Horvath 1906a, p. 49. 

Hyalochiton syrmiensis var. consimilis: Vidal 1937, p. 197 [Morocco]. 
Type: Sex unknown; Spain; Hungarian Mus. 
Distribution: Spain; Morocco. 
Host Plant: Unrecorded. 

*Hyalochiton komaroffii (Jakovlev) 

Galeatus komaroffii Jakovlev 1880a, pp. 130, 133 [Caucasus]. — Horvath 

1897b, pp. 456, 458 [Hungary; Austria]. 
Galeatus comaroffii [sic]: Jakovlev 1880b, p. 102. 
Galeatus debilis Montandon 1887, p. 65 [Yugoslavia; Teucrium]. 
Galeatus (Hyalochiton) komaroffii: Horvath 1905b, p. 56. 
Hyalochiton komaroffii: Horvath 1906a, p. 49 [France]. — Guide 1938, 

p. 271, fig.— Stichel 1938a, p. 406 [Turkey]; 1960a, p. 293.— Drake 

and Ruhoff 1962d, p. 489. 
Hyalochiton algiricus Lindberg 1936a, p. 83, fig. 2 [Morocco]. 
Hyalochiton komarovii [sic]: Kiritshenko 1951, p. 246. — Gomez-Menor 

1955b, p. 249 [Spain], 



246 U.S. NATIONAL MUSEUM BULLETIN 243 

Type: Holotype d\ brachypterous; Derbent, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Caucasus); Yugoslavia; Hungary; Austria; 
France; Spain; Turkey; Morocco. 
Host Plant: Teucrium sp. 

*Hyalochiton multiseriatus (Reuter) 

Galeathus [sic] multiseriatus Reuter 1888, p. 225 [Greece]; 1891a, p. 26. 
Galeatus multiseriatus: Horvath 1897b, pp. 456, 459. 
Hyalochiton multiseriatus: Horvath 1906a, p. 49. — Stichel 1960a, p. 294 
[Turkey; Iraq; Syria; Phlomis]. 
Type: Sex unknown; Attica, Greece; Helsin. Mus. 
Distribution: Greece; Turkey; Iraq; Syria. 
Host Plant: Phlomis sp. 

*Hyalochiton strumosus (Horvath) 

Galeatus strumosus Horvath 1902a, p. 594 [Turkey]. 
Hyalochiton strumosus: Horvath 1906a, p. 49. — Seidenstiicker 1954, 
p. 235.— Hoberlandt 1955, p. 87.— Stichel 1960a, p. 294 [Syria]. 
Type: Sex unknown; Ak-Chehir, Anatolia, Turkey; Hungarian Mus. 
Distribution: Turkey; Syria. 
Host Plant: Unrecorded. 

Hyalochiton syrmiensis (Horvath) 

Galeatus syrmiensis Horvath 1897a, p. 88 [Hungary]; 1897b, pp. 456, 

458. 
Hyalochiton syrmiensis: Horvath 1906a, p. 49. — Vidal 1937, p. 197 
[Morocco].— Gonzalez 1948, p. 50 [Spain].— Stichel 1960a, p. 293. 
Type: Holotype o 71 ; Hungary; Hungarian Mus. 
Distribution: Hungary; Spain; Morocco. 
Host Plant: Unrecorded. 

Genus HYBOPHARSA Hurd 

Hybopharsa Hurd 1946, p. 467.— Drake and Ruhoff 1960a, p. 61. 
Type Species: Leptostyla colubra Van Duzee. 

*Hybopharsa colubra (Van Duzee) 

Leptostyla colubra Van Duzee 1907, p. 19 [Jamaica]. 

Gelchossa colubra: Drake and Bruner 1924a, p. 149; 1924b, p. 155 

[Cuba; Eugenia rhombea]. — Gowdey 1926, p. 35. 
Leptopharsa colubra: Bruner, Scaramuzza and Otero 1945, p. 74. 
Hybopharsa colubra: Hurd 1946, p. 468 [pimento]. 
Type: Sex unknown; Jamaica; Cal. Acad. 
Distribution: Jamaica; Cuba. 
Host Plants: Eugenia rhombea; Pimenta pimenta. 



LACEBUGS OF THE WORLD 247 

Genus HYPSIPYRGIAS Kirkaldy 

Hypsipyrgias Kirkaldy 1908a, p. 779. — Drake and Ruhoff 1960a, p. 61. 
Type Species: Hypsipyrgias telamonides Kirkaldy. 

♦Hypsipyrgias euphues Drake and Ruhoff 

Hypsipyrgias euphues Drake and Ruhoff 1962b, p. 251 [Lord Howe]. 
Type: Holotype d", macropterous; Lord Howe Island; So. Austr. Mus. 
Distribution: Australia (Lord Howe Island). 
Host Plant: Unrecorded. 

♦Hypsipyrgias telamonides Kirkaldy 

Hypsipyrgias telamonides Kirkaldy 1 908a, p. 779, pi. 43, figs. 4, 5 [Queens- 
land].— Hacker 1927, p. 25; 1928a, p. 179.— Drake and Ruhoff 
1962b, p. 250. 
Type: Holotype 9; Kuranda, Queensland, Australia; Hawaii. Sugar 
Plant. Assn. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Note: Type specimen bears: type label, identification label in Kirkaldy's 
handwriting, and printed locality label for "Cairns District." Kuranda is a 
small waystation about 15 miles from Cairns, the latter label probably 
used for convenience. 

Genus HYPSOTINGIS Drake 

Hypsotingis Drake 1960, p. 362. 
Type Species: Hypsotingis columna Drake. 

Hypsotingis columna Drake 

Hypsotingis columna Drake 1960, p. 363, fig. 15 [New Guinea]. 
Type: Holotype 9; Urapura, Kamo Valley, Wisselmeren, Netherlands 
New Guinea; Bishop Mus. 
Distribution: New Guinea (Netherlands). 
Host Plant: Unrecorded. 

Genus IDIOCYSTA China 

Idiocysta China 1930, p. 141.— Drake and Ruhoff 1960a, p. 61. 
Type Species: Idiocysta hackeri China. 

Idiocysta bicolor Drake and Poor 

Idiocysta bicolor Drake and Poor 1943, p. 199, fig. 4 [Viti Levu]. 
Type: Holotype 9; Nandarivatu, Viti Levu, Fiji Islands; Bishop Mus. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Unrecorded. 



248 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Idiocysta dryadis Drake and Poor 

Idiocysta dryadis Drake and Poor 1943, p. 200, fig. 5 [Viti Levu]. 
Type: Holotype 9; Nandarivatu, Viti Levu, Fiji Islands; Bishop Mus. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Unrecorded. 

*Idiocysta fijiana Drake and Poor 

Idiocysta fijiana Drake and Poor 1943, p. 198 [Viti Levu]. 

Type: Holotype cf ; Mt. Victoria, Tholo North, Viti Levu, Fiji Islands; 
Bishop Mus. 

Distribution: Fiji Islands (Viti Levu). 

Host Plant: Unrecorded. 

*Idiocysta floris Drake and Poor 

Idiocysta floris Drake and Poor 1943, p. 198 [Viti Levu]. 
Type: Holotype 9; Nandarivatu, Viti Levu, Fiji Islands; Bishop Mus. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Unrecorded. 

Idiocysta hackeri China 

Idiocysta hackeri China 1930, p. 142, fig. 22 [Upolu]. 
Type: Sex unknown; Malololelei, Upolu, Samoa Islands; British Mus. 
Distribution: Territory Western Samoa Islands (Upolu). 
Host Plant: Unrecorded. 

Genus IDIOSTYLA Drake 

Idiostyla Drake 1945, p. 97.— Drake and Ruhoff 1960a, p. 61. 
Type Species: Tigava anonae Drake and Hambleton. 

*Idiostyla anonae (Drake and Hambleton) 

Tigava anonae Drake and Hambleton 1938a, p. 45 [Brazil; Annona 

coriacea]. — Monte 1941e, p. 143. 
Idiostyla anomae [sic]: Drake 1945, p. 98. 
Type: Holotype d", macropterous; Cerqueira Cesar, Sao Paulo, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Annona coriacea. 

*Idiostyla rolliniae (Drake and Hambleton) 

Tigava rolliniae Drake and Hambleton 1934, p. 441 [Brazil; Rollinia 

silvatica].—Montz 1939b, p. 81; 1941e, p. 145. 
Idiostyla rollinae [sic]: Drake 1945, p. 98. — Drake and Davis 1960, fig. 46. 
Type: Holotype d" , macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Rollinia silvatica. 



LACEBUGS OF THE WORLD 249 

Genus ILDEFONSUS Distant 

Ildefonsus Distant 1910a, p. 110.— Drake and Ruhoff 1960a, p. 61. 
Type Species: Ildefonsus provorsus Distant. 

*Ildefonsus nexus Drake and Ruhoff 

Ildefonsus nexus Drake and Ruhoff 1961b, p. 180 [Burma]. 

Type: Holotype 9, macropterous; Kambaiti, Burma; Drake Coll. 
(USNM). 

Distribution: Burma. 

Host Plant: Unrecorded. 

*Ildefonsus provorsus Distant 

Ildefonsus provorsus Distant 1910a, p. 110, fig. 54 [India]. 
Type: Sex unknown; E. Himalayas, Ghoom, India; British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 

Genus INOMA Hacker 

Inoma Hacker 1927, p. 25.— Drake and Ruhoff 1960a, p. 62. 
Type Species: Inoma multispinosa Hacker. 

*Inorna angusta Drake 

Inoma angusta Drake 1942a, p. 16 [New South Wales]. 
Type: Holotype cf, macropterous; Dorrizo, New South Wales, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (New South Wales). 
Host Plant: Unrecorded. 

*Inoma multispinosa Hacker Plate 42 

Inoma multispinosa Hacker 1927, p. 25, pi. 9, fig. 12 [Queensland]. 

Type: Sex unknown; Sunnybank, Queensland, Australia; Queensland 
Mus. 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

Genus INONEMIA Drake 

Inonemia Drake 1942b, p. 361.— Drake and Ruhoff 1960a, p. 62. 
Type Species: Inonemia mussiva Drake. 

*Inonemia mussiva Drake 

Inonemia mussiva Drake 1942b, p. 362 [Queensland]. 

Type: Holotype o 71 , macropterous; Roma, Queensland, Australia; 
Drake Coll. (USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

681-552—65 17 



250 U.S. NATIONAL MUSEUM BULLETIN 243 

*Inonemia mussiva subsp. brevis Drake 

Inonemia mussiva subsp. brevis Drake 1942b, p. 362 [Queensland]. 

Type: Holotype 9, macropterous; Roma, Queensland, Australia; Drake 
Coll. (USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

Genus ISCHNOTINGIS Horvath 

Ischnotingis Horvath 1925a, p. 7. — Drake and Ruhoff 1960a, p. 62. 
Type Species: Ischnotingis prolixa Horvath. 

*Ischnotingis fasciata Horvath 

Ischnotingis fasciata Horvath 1925a, p. 9, fig. 5 [Western Australia]. 

Type: Sex unknown; Geraldton, Australia occidentalis; Stockholm Mus. 

Distribution: Australia (Western Australia; New South Wales). 

Host Plant: Unrecorded. 
*Ischnotingis horvathi Drake 

Ischnotingis horvathi Drake 1954c, p. 69 [New South Wales]; 1961b, p. 112. 

Type: Holotype <?; Mt. Victoria, New South Wales, Australia; Hungar- 
ian Mus. 

Distribution: Australia (New South Wales). 

Host Plant: Unrecorded. 

*Ischnotingis prolixa Horvath 

Ischnotingis prolixa Horvath 1925a, p. 8, fig. 4 [Western Australia]. — 

Drake and Ruhoff 1962a, p. 156. 
Ischnotingis yanchepana Drake 1953d, p. 215. 
Type: Holotype 9; Freemantle, Australia occidentalis; Stockholm Mus. 
Distribution: Australia (Western Australia). 
Host Plant: Unrecorded. 

Genus KAPIRIELLA Schouteden 

Kapiriella Schouteden 1919, p. 138.— Drake 1953b, p. 93; 1957b, pp. 

205-218 (key).— Drake and Ruhoff 1960a, p. 62. 
Lembella Schouteden 1919, p. 141.— Drake and Ruhoff 1960a, p. 63. 
Type Species: Kapiriella leplaei Schouteden. 
*Kapiriella conradsi Drake 

Kapiriella conradsi Drake 1957b, p. 215 [Tanganyika]. 
Cysteochila conradsi: Schouteden 1957c, p. 316 [Ruanda]. 
Type: Holotype cf; Victoria-Nyanza, Ukerewe Island, Tanganyika; 
Cent. Afr. Mus. 
Distribution: Tanganyika; Ruanda-Urundi. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 251 

*Kapiriella denigrata Drake 

Kapiriella denigrata Drake 1956a, p. 5, fig. 1 [Congo]; 1957b, p. 217. 
Type: Holotype cf ; Ngoma, Nord du lac Kivu, Belgian Congo; Pares 
Nat. Inst. 

Distribution: Congo. 
Host Plant: Unrecorded. 

*Kapiriella desaegeri Drake 

Kapiriella desaegeri Drake 1957b, p. 216 [Congo]. 
Type: Holotype cf; National Garamba Park, Belgian Congo; Pares 
Nat. Inst. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Kapiriella leplaei Schouteden 

Kapiriella leplaei Schouteden 1919. p. 139 [Congo]; 1923, p. 107 — 
Drake 1953b, p. 93; 1954e, p. 8; 1957b, p. 212. 
Type: Sex unknown; Kapiri, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

* Kapiriella maynei (Schouteden) 

Lembella maynei Schouteden 1919, p. 142 [Congo]; 1923, p. 110. 

Lembella maculigera Horvath 1929, p. 325 [Uganda]. 

Kapiriella maynei: Drake 1953b, p. 93; 1955b, p. 87 [Angola]; 1957b, 
p. 209, fig. lb [Cameroons] . 
Type: Sex unknown; Congo da Lemba, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Cameroons; Uganda; Angola. 
Host Plant: Unrecorded. 

*Kapiriella natalana Drake 

Kapiriella natalana Drake 1956f, p. 423 [Natal]; 1957b, p. 214. 
Type: Holotype o" ; Malvern, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

*Kapiriella polita (Drake) 

Lembella polita Drake 1948e, p. 150 [Sudan]; 1953b, p. 93 (in part). 

Lambella [sic] polita: Drake 1948h, p. 76. 

Lembella iturica Schouteden 1954, p. 141 [Congo]. 

Kapiriella polita: Drake 1957b, p. 210 [Senegal; Uganda; millet]. 
Type: Holotvpe cf, macropterous; Kavlugh, Sudan; Drake Coll. 
(USNM). 
Distribution: Sudan; Uganda; Congo; Senegal. 
Host Plant: Millet. 



252 U.S. NATIONAL MUSEUM BULLETIN 243 

*Kapiriella saetula Drake 

Kapiriella saetula Drake 1957b, p. 214, fig. la [Congo]. 
Type: Holotype cf; Kaziba, P.N.U., Belgian Congo; Pares Nat. Inst. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Kapiriella schoutedeni Drake 

Kapiriella schoutedeni Drake 1957b, p. 213, fig. Id [Malagasy], 
Type: Holotype o 71 ; Madagascar; Cent. Afr. Mus. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

*Kapiriella wechinai Drake 

Kapiriella wechinai Drake 1957b, p. 211, fig. lc [Northern Rhodesia; 
Congo; Kenya; Tanganyika]. 
Type: Holotype cf ; Lake Bangweulu, N'Chiti Island, Northern Rho- 
desia; British Mus. 
Distribution: Northern Rhodesia; Congo; Kenya; Tanganyika. 
Host Plant: Unrecorded. 

Genus LAROTINGIS Drake 

Larotingis Drake 1960, p. 357. 
Type Species: Larotingis aporia Drake. 
Larotingis aporia Drake 

Larotingis aporia Drake 1960, p. 357, fig. 11 [New Guinea]. 
Type: Holotype o* ; Wisselmeren, Okaitadi, Netherlands New Guinea; 
Bishop Mus. 
Distribution: New Guinea (Netherlands). 
Host Plant: Unrecorded. 

*Larotingis etes Drake and Ruhoff 

Larotingis etes Drake and Ruhoff 1961b, p. 165 [Mindanao]. 
Type: Holotype 9, macropterous; Surigao, Mindanao, Philippine 
Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Mindanao). 
Host Plant: Unrecorded. 

Genus LASIACANTHA Stal 

Tingis (Lasiacantha) Stal 1873, p. 130. 

Lasiacantha: Stal 1874, p. 56. — Lethierry and Severin 1896, p. 18. — 
Horvath 1906a, pp. 14, 59 (key).— Muzik 1907, pp. 50, 57.— Oshanin 
1908, p. 427; 1912, p. 44.— Stichel 1926, pp. 104, 109 (key); 1935, 
p. 349; 1938a, p. 406; 1960a, p. 306; 1960b, p. 395; 1960c, p. 134. — 
Hacker 1929, p. 334.— Borner 1935, pp. 74, 76.— Guide 1938, 
pp. 244, 283 (key).— China 1943, p. 246.— Kiritshenko 1951, pp. 



LACEBUGS OF THE WORLD 253 

242, 248 (key).— Singer 1952, p. 51.— Drake 1953b, p. 92.— Drake 

and Runoff 1960a, p. 63. 
Monanthia {Lasiacantha): Puton 1886b, p. 33. 
Jannaeus Distant 1909a, p. 118; 1910a, p. 117.— Drake and Ruhoff 

1960a, p. 62. 
Myrmecotingis Hacker 1928, p. 182. — Drake and Ruhoff 1960a, p. 69. 
Type Species: Tingis {Lasiacantha) hedenborgii Stal. 

*Lasiacantha absimilis Drake 

Lasiacantha absimilis'Drake 1951, p. 177 [Ethiopia]; 1954a, p. 232 [Niger]. 
Type: Holotype cf; Bubassa, Abyssinia; Hungarian Mus. 
Distribution: Ethiopia; Niger; Sudan. 
Host Plant: Unrecorded. 

♦Lasiacantha altimitrata (Takeya) 

Jannaeus altimitratus Takeya 1933, p. 35, pi. 3, fig. 3; text fig. 3 [Taiwan]; 

1951a, p. 15.— Drake and Maa 1953, p. 88 [China; Labiatae]. 
Lasiacantha altimitratus: Drake 1953b, p. 92. 
Type: Holotype d" ; Taihoku, Formosa; Kyushu Univ. 
Distribution: Taiwan; China; India. 
Host Plant: Labiatae. 

*Lasiacantha capucina (Germar) 

Acanthia cardui (not Linnaeus): Schellenberg 1800, p. 21, pi. 6, fig. 2. 

Tingis capucina Germar 1836, fasc. 18, tab. 24 [Germany]. 

Monanthia {Phyllontocheila) setulosa Fieber 1844, p. 68, pi. 5, figs. 34-36 
[Czechoslovakia; Thymus serpyllus]. — Herrich-Schaeffer 1850, p. 154. 

Camypotsteira (Derephysia) setulosa var. gracilis (not Herrich-Schaeffer): 
Kolenati 1856, p. 431 [Caucasus; Thymus montanus]. 

Monanthia {Phyllontocheila) gracilis: Flor 1860, p. 343 [Austria; France; 
Estonia; Latvia]. 

Monanthia setulosa: Fieber 1861, p. 122. — Frey-Gessner 1865, p. 231 
[Switzerland].— Garbiglietti 1869, p. 272 [Italy].— Jakovlev 1869, 
p. Ill; 1880b, p. 106.— Horvath 1874b, p. 432 [Hungary].— Puton 
1874a, p. 226.— Reiber and Puton 1876, p. 69.— Ferrari 1878, p. 66. 

Lasiacantha setulosa: Stal 1874, p. 56. 

Monanthia capucina: Puton 1879c, p. 113.— d'Antessanty 1890, p. 32. — 
Hiieber 1893, p. 337.— Rey 1893, p. 97. 

Monanthia {Lasiacantha) capucina: Puton 1886b, p. 33. 

Lasiacantha capucina: Lethierry and Severin 1896, p. 18. — Horvath 
1906a, p. 60 [Rumania; Yugoslavia].— Butler 1923, p. 208 [Eng- 
land]. — Stichel 1926, p. 109 [Ajuga; Galium; Hieracium]; 1960a, p. 307 
[Poland; Bulgaria].— Scholte 1935, p. 76, fig. 18 [Netherlands].— 
Guide 1938, p. 285, fig.— Hoberlandt 1942, p. 125 [Transcaucasus] .— 
Blote 1945, p. 87 [Tunisia].— Kiritshenko 1951, p. 248.— Novak and 
Wagner 1951, p. 70.— Bator 1953, p. 325, pi. 2, fig. 6.— Gomez- 



254 U.S. NATIONAL MUSEUM BULLETIN 243 

Menor 1955b, p. 249 [Spain].— Stusak 1957a, p. 20, figs. 1-3; 1957b, 
pp. 137, 139, figs. 6, 10a, lib; 1961a, p. 77, fig. 5e. — Southwood and 
Leston 1959, p. 147, pi. 21, fig. 10.— Mancini 1959, p. 260. 
Phyllontocheila capucina: Reichensperger 1920, p. 62. 
Type: "Saxonia"; sex and deposition of type unknown. 
Distribution: England; Netherlands; France; Spain; Germany; Switzer- 
land; Czechoslovakia; Poland; Hungary; Austria; Italy; Yugoslavia; 
Bulgaria; Rumania; U.S.S.R. (Estonia; Latvia; Caucasus; Transcaucasus) ; 
Tunisia. 

Host Plants: Ajuga sp.; Galium sp.; Hieracium sp.; Thymus montanus; 
Thymus serpyllus; moss. 

Note: Study of eggs and immature stages [Stusak 1957a, 1957b, 1961a]. 
*Lasiacantha comans Drake (emendation) 

Lasiacanthia [sic] comantis Drake 1953d, p. 220 [Senegal]. 
Type: Holotype <?', Bambey, Senegal; British Mus. 
Distribution: Senegal. 
Host Plant: Unrecorded. 
*Lasiacantha compta (Drake) 

Furcilliger comptus Drake 1942b, p. 363 [Queensland; Gmelina leichardtii]. 
Lasiacantha comptus: Drake and RuhofF 1961c, p. 145. 
Type: Holotype o 71 , macropterous; Imbil, Queensland, Australia; Drake 
Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Gmelina leichardtii. 
*Lasiacantha crassicornis Horvath 

Lasiacantha crassicornis Horvath 1929, p. 320 [Natal]. 
Type: Holotype 9; Weenen, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 
*Lasiacantha cuneata (Distant) 

Jannaeus cuneatus Distant 1909a, p. 118 [India]; 1910a, p. 118, fig. 
60.— Drake and Poor 1936b, p. 146.— Drake 1937a, p. 387 [Ceylon].— 
Singh 1953, p. 118. 
Lasiacantha cuneautus [sic]: Drake 1953b, p. 92. 
Type: Sex unknown; North Bengal, India; British Mus. 
Distribution: India; Ceylon; China. 
Host Plant: Unrecorded. 
*Lasiacantha discors Drake (emendation) 

Lasiacantha discordis Drake 1955a, p. 80 [Western Australia], 
Type: Holotype 9, macropterous; Kalgoorie, Western Australia, Aus- 
tralia; Drake Coll. (USNM). 
Distribution: Australia (Western Australia). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 255 

*Lasiacantha gambiana Drake 

Lasiacantha gambiana Drake 1954b, p. 11 [Gambia]. 

Type: Holotype cf, brachypterous; Bathurst, Gambia, West Africa; 
Drake Coll. (USNM). 

Distribution: Gambia. 

Host Plant: Unrecorded. 

*Lasiacantha gracilis (Herrich-Schaeffer) 

Tingis gracilis Herrich-Schaeffer 1830, heft 118, tab. 20 [Germany]; 

1835, p. 58.— Horvath 1905b, p. 569. 
Derephysia gracilis: Herrich-Schaeffer 1838, p. 72, pi. 128, figs. d-h. 
Monanthia (Phyllontocheila) setulosa var. gracilis: Fieber 1844, p. 69, 

pi. 5, figs. 37, 38 [Austria]. 
Campylosteira {Derephysia) setulosa var. capucina (not Germar): Kolenati 

1856, p. 431. 
Monanthia setulosa var. gracilis: Fieber 1861, p. 122. — Frey-Gessner 

1865, p. 231 [Switzerland]. 
Monanthia piligera Garbiglietti 1869, p. 273 [Italy]. 
Monanthia cucullifera Puton 1888, p. 105. 
Lasiacantha cucullifera: Lethierry and Severin 1896, p. 19. 
Lasiacantha gracilis: Horvath 1906a, pp. 59, 60 [Hungary; Caucasus]. — 
Stichel 1938a, p. 406 [France; Poland]; 1960a, p. 306 [Turkey; 
Bulgaria; Satureja montana; Thymus]. — Hoberlandt 1942, p. 125 
[Czechoslovakia; Transcaucasus] ; 1943, p. 118. — Kiritshenko 1951, 
p. 248.— Wagner 1958, p. 240. 
Lasiacantha cracilis [sic]: Stichel 1935, p. 349. 
Type: Regensburg, Germany; sex and deposition of type unknown. 
Distribution: Germany; France; Austria; Hungary; Czechoslovakia; 
Switzerland; Italy; Poland; Bulgaria; Turkey; U.S.S.R. (Caucasus; 
Transcaucasus) . 
Host Plants: Satureja montana; Thymus sp. 

* Lasiacantha hedenborgii (Stal) 

Tingis (Lasiacantha) hedenborgii Stal 1873, p. 130 [Cyprus]. 
Lasiacantha hedenborgii: Stal 1874, p. 56. — Horvath 1906a, p. 60 
[Israel].— Seidenstucker 1954, p. 235 [Turkey] .—Hoberlandt 1955, 
p. 88. 
Monanthia hedenborgi [sic]: Puton 1881, p. 123 [Lebanon]. 
Monanthia (Lasiacantha) hedenborgi [sic]: Puton 1886b, p. 33. 
Type: Holotype 9; Cyprus; Stockholm Mus. 
Distribution: Cyprus; Lebanon; Israel; Turkey. 
Host Plant: Unrecorded. 



256 U.S. NATIONAL MUSEUM BULLETIN 243 

*Lasiacantha histricula (Puton) 

Monanthia {Lasiacantha) histricula Puton 1878, p. lxvii [Spain]; 1886b, 

p. 33. 
Monanthia histricula: Puton 1880, p. 243 [France]. 

Lasiacantha histricula: Lethierry and Severin 1896, p. 19. — Horvath 
1906a, p. 61.— Gonzalez 1948, p. 50— Gomez-Menor 1955b, p. 249. 
Type: Sex unknown; Madrid, Spain; Vienna Mus. 
Distribution: Spain; France. 
Host Plant: Unrecorded. 

*Lasiacantha horvathi Drake 

Lasiacantha horvathi Drake 1951, p. 176 [Tanganyika]. 
Type: Holotype 9; Meschi, Africa Orientalis; Hungarian Mus. 
Distribution: Tanganyika; Sudan; South Africa (Cape Province). 
Host Plant: Unrecorded. 

Lasiacantha insularis Schouteden 

Lasiacantha insularis Schouteden 1957a, p. 84 [Malagasy]. 
Type: Sex unknown; Tananarive, Madagascar; Paris Mus. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

*Lasiacantha kin Schouteden 

Lasiacantha kin Schouteden 1955a, p. 28 [Congo]. 

Type: Sex unknown; Kinchassa, Bas-Congo, Belgian Congo; Cent. Afr. 
Mus. 

Distribution: Congo. 

Host Plant: Unrecorded. 

Lasiacantha leai (Hacker) 

Myrmecotingis leai Hacker 1928, p. 182, pi. 23. fig. 15 [Western Aus- 
tralia]. 
Lasiacantha leai: Hacker 1929, p. 334. 
Type: Sex unknown; Swan River, Western Australia, Australia; Queens- 
land Mus. 
Distribution : Australia (Western Australia) . 
Host Plant: Unrecorded. 

Note: Inquiline, "originally carded with specimens of the Dolichoderine 
ant Iridomyrmex conifer Forel" [Hacker 1928]. 

*Lasiacantha merita Drake 

Lasiacantha merita Drake 1958c, p. 322 [Malagasy]. 

Type: Sex unknown; Tulear, Anakao du Bas, Madagascar; Madagascar 
Sci. Inst. 

Distribution: Malagasy Republic. 

Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 257 

Lasiacantha odontostoma (Stal) 

Tingis (Lasiacantha) odontostoma Stal 1873, p. 130 [Sierra Leone]. 

Lasiacantha odontostoma: Lethierry and Severin 1896, p. 19 [Guinea]. 

Lasiacantha odontosoma [sic]: Distant 1902a, p. 242, pi. 15, fig. 6. 
Type: Holotype cf; Sierra Leone; Stockholm Mus. 
Distribution: Sierra Leone; Guinea. 
Host Plant: Unrecorded. 

*Lasiacantha sideris Drake 

Lasiacantha sideris Drake 1951, p. 175 [Kenya]. 
Type: Holotype 9, macropterous; Murnias, Kenya; Hungarian Mus. 
Distribution: Kenya. 
Host Plant: Unrecorded. 

*Lasiacantha turned Drake 

Lasiacanthia [sic] turneri Drake 1953d, p. 219 [South-West Africa]. 
Type: Holotype o 71 ; Okahandja, South-West Africa; British Mus. 
Distribution: South-West Africa. 
Host Plant: Unrecorded. 

*Lasiacantha yebo Schouteden 

Lasiacantha yebo Schouteden 1955a, p. 28 [Congo]. 
Type: Sex unknown; Yebo, Ituri, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Kenya. 
Host Plant: Unrecorded. 

Genus LEPTOBYRSA Stal 

Leptobyrsa Stal 1873, pp. 119, 123. — Lethierry and Severin 1896, p. 
13.— Drake and Poor 1937c, pp. 163-165.— Monte 1939b, p. 71; 
1941e, p. 101.— Drake and Ruhoff 1960a, p. 64. 
Type Species: Tingis steini Stal. 

*Leptobyrsa ardua Drake 

Leptobyrsa ardua Drake 1922c, p. 376 [Brazil]. — Drake and Hambleton 
1938b, p. 54.— Monte 1939b, p. 71 [Symphyoppapus reticulars]. 
Type: Holotype o 71 , macropterous; Minas Gerais, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil; Argentina. 
Host Plant: Symphyoppapus reticulatus. 

*Leptobyrsa baccharidis Drake and Hambleton 

Leptobyrsa baccharidis Drake and Hambleton 1938a, p. 65 [Brazil; 
Baccharis mesoneura; Eupatorium intermedium]. — Silva 1956, p. 36. 
Type: Holotype o 71 , macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plants: Baccharis mesoneura; Eupatorium intermedium. 



258 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptobyrsa bruchi Drake 

Leptobyrsa bruchi Drake 1928i, p. 73 [Argentina]. 
Type: Hole-type d\ macropterous; Cordoba, Argentina; Drake Coll. 
(USNM). 
Distribution: Argentina. 
Host Plant: Unrecorded. 

*Leptobyrsa decora Drake 

Leptobyrsa decora Drake 1922c, p. 375 [Colombia; Ecuador]. — Monte 
1938b, p. 130 [Citrus aurantium]. 
Type: Holotype 0% macropterous; Lacumbre, Colombia; Drake Coll. 
(USNM). 
Distribution: Colombia; Ecuador. 
Host Plant: Citrus aurantium. 

*Leptobyrsa mendocina Pennington 

Leptobyrsa mendocina Pennington 1919, p. 526, fig. 1 [Argentina]; 1921, 

p. 20. 
Leptobyrsa mendecina [sic]: Drake 1931b, p. 511. 
Type: Holotype 9, macropterous; Mendosa, Argentina; Drake Coll. 
(USNM). 
Distribution: Argentina. 
Host Plant: Unrecorded. 

*Leptobyrsa pulchra Monte 

Leptobyrsa pulchra Monte 1940e, p. 286 [Brazil]. 

Type: Holotype o 71 ; Nova Teutonia, Santa Catarina, Brazil; Monte 
Coll. (Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Leptobyrsa steini (Stal) 

Tingis steini Stal 1858, p. 64 [Brazil].— Walker 1873a, p. 181. 
Leptobyrsa steinii [sic]: Stal 1873, p. 123. — Pennington 1919, p. 527, fig. 
Leptobyrsa steini: Champion 1898b, p. 58, pi. 2, fig. 4. — Drake and 
Hambleton 1935, p. 148 [Eupatorium intermedium; Baccharis mesoneura]; 
1938a, p. 65 [Symphyopappus reticulatus]; 1944b, p. 126. — Drake 1936b, 
p. 700 [Argentina].— Drake and Poor 1937d, p. 309.— Monte 1941c, 
p. 204, fig. 1; 1942d, p. 108; 1944b, p. 461, figs. 4-8.— Singh 1953, 
p. 118. 
Leptobyrsa nigritarsis Monte 1937a, p. 32, fig. 10; 1937c, p. 72; 1941c, 
p. 207, fig. 2. 
Type: Holotype cf ; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil; Argentina. 

Host Plants: Baccharis mesoneura; Eupatorium intermedium; Symphyopappus 
reticulatus. 



LACEBUGS OF THE WORLD 259 

*Leptobyrsa tersa Drake and Hamblcton 

Leptobyrsa tersa Drake and Hambleton 1935, p. 148 [Brazil; Moquinia 
polymorpha]. — Monte 1937a, p. 32 [Moquinia]. 
Type: Holotypeo 71 , macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plants: Moquinia polymorpha; Moquinia sp. 

Genus LEPTOCYSTA Stal 

Leptocysta Stal 1873, pp. 121, 127.— Monte 1939b, p. 71; 1941e, p. 103; 
1946a, p. 325 (key).— Drake and Runoff 1960a, p. 64. 
Type Species: Tingis sexnebulosa Stal. 

*Leptocysta notialis Drake 

Leptocysta notialis Drake 1948d, p. 18 [Argentina]. 

Type: Holotypeo 71 , macropterous; Rosas, F. C. Sud, Buenos Aires, 
Argentina; Drake Coll. (USNM). 

Distribution : Argentina. 

Host Plant: Unrecorded. 

*Leptocysta novatis Drake 

Leptocysta sexnebulosa (not Stal): Berg 1884, p. 103. 
Leptocysta novatis Drake 1928i, p. 72 [Argentina]; 1935, p. 17 [Para- 
guay].— Drake and Poor 1938b, p. 107.— Monte 1938d, p. 387; 
1946a, p. 328, figs. 3, 4. 
Type: Holotypeo 71 , macropterous; La Plata, Argentina; Drake Coll. 
(USNM). 
Distribution: Argentina; Paraguay. 
Host Plant: Unrecorded. 

*Leptocysta sexnebulosa (Stal) 

Tingis sex-nebulosa Stal 1858, p. 64 [Brazil].— Walker 1873, p. 181. 
Leptocysta sexnebulosa: Stal 1873, p. 127. — Lethierry and Severin 
1896, p. 15 [Argentina].— Champion 1898b, p. 61, pi. 2, figs. 10, 
10a.— Pennington 1921, p. 20.— Drake 1930a, p. 1; 1931b, p. 511 
[Colombia]; 1935, p. 17 [Paraguay]. — Drake and Hambleton 
1934, p. 442 [Antennaria; Ipomoea batatas']; 1945, p. 359. — Costa 
Lima 1936, p. 128.— Bosq 1937, p. 130.— Drake and Poor 1937d, 
p. 306, pi. 36, fig. 13; 1939b, p. 98.— Monte 1937, p. 34, fig. 3 
[Mikania]; 1938b, p. 131; 1939b, p. 71 [Venezuela; Vernonia]; 1940c, 
p. 193; 1940d, p. 101; 1946a, p. 326, figs. 1, 2 [Peru]. 
Leptocysta sexnolata [sic]: Drake 1936b, p. 699. 
Type: Holotypeo 71 ; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil; Venezuela; Peru; Colombia; Argentina; Paraguay. 
Host Plants: Antennaria sp.; Ipomoea batatas; Mikania sp.; Vernonia sp. 



260 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptocysta tertia Monte 

Leptocysta tertia Monte 1946a, p. 330, figs. 5, 6 [Argentina]. 

Type: Holotype cf; Puerto Tirol, Argentina; Monte Coll. (Mus. 
Nacional). 

Distribution: Argentina. 

Host Plant: Unrecorded. 

Genus LEPTODICTYA Stal 

Leptodictya Stal 1873, pp. 121, 127. — Lethierry and Severin 1896, p. 
14.— Champion 1897, p. 23 (key).— Van Duzee 1916, p. 26; 
1917b, p. 219.— Blatchley 1926, p. 477 (key).— Drake 1931e, pp. 
119-121.— Monte 1939b, p. 71; 1941e, p. 103.— Hurd 1946, p. 
452.— Drake and Runoff 1960a, p. 64. 
Type Species: Monanthia {Physatocheila) ochropa Stal. 

Subgenus LEPTODICTYA (LEPTODICTYA) Stal 

Type Species: Monanthia {Physatocheila) ochropa Stal. 
* Leptodictya ochropa (Stal) 

Monanthia {Physatocheila) ochropa Stal 1858, p. 62 [Brazil]. 
Monanthia ochropa: Walker 1873a, p. 192. 

Leptodictya ochropa: Stal 1873a, p. 127. — Drake 1931e, pp. 119, 121; 
1936b, p. 699 [Argentina].— Drake and Poor 1937d, p. 306, pi. 36, 
fig. 4. — Drake and Hambleton 1938b, p. 53 [bamboo]. — Monte 
1938b, p. 131.— Silva 1956, p. 38, fig. 13.— Drake and Davis 1960, 
fig. 60. 
Type: Holotype cf; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Argentina; Brazil. 
Host Plant: Bamboo. 

Subgenus LEPTODICTYA (HANUALA) Kirkaldy 

Hanuala Kirkaldy 1905, p. 217. 

Leptodictya {Hanuala): Drake 1931e, pp. 119-121.— Monte 1939b, p. 
72; 1941e, p. 104.— Drake and Ruhoff 1960a, p. 65. 
Type Species: Hanuala leinahoni Kirkaldy. 
*Leptodictya (Hanuala) approximata (Stal) 

Monanthia {Physatocheila) approximata Stal 1858, p. 63 [Brazil]. 
Monanthia approximata: Walker 1873a, p. 193. 

Leptodictya approximata: Stal 1873, p. 127.— Drake 1928a, p. 46 — 
Drake and Poor 1937d, p. 307, pi. 36, fig. 3 [British Guiana].— Drake 
and Hambleton 1938b, p. 53. 
Leptodictya {Hanuala) approximata: Drake 1931e, p. 121. — Monte 1941e, 
p. 104 [Bambusa]. 
Type: Sex unknown; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil; British Guiana; Bolivia. 
Host Plant: Bambusa sp. 



LACEBUGS OF THE WORLD 261 

*Leptodictya (Hanuala) austrina Drake and Hambleton 

Leptodictya austrina Drake and Hambleton 1939, p. 157 [Brazil; Olyra], 
Leptodictya {Hanuala) austrina: Monte 1941e, p. 104. 
Type: Holotype cf, macropterous; Ribeirao Preto, Sao Paulo, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Olyra sp. 
*Leptodictya (Hanuala) bambusae Drake 

Leptodictya bambusae Drake 1918a, p. 175 [Puerto Rico; Bambusa vul- 
garis]; 1926a, p. 86 [Cuba; sugar] .— Wolcott 1923, p. 247.— Drake 
and Bruner 1924a, p. 149 [Mexico].— Barber 1939, p. 370 [Haiti].— 
Bruner, Scaramuzza and Otero 1945, pp. 19, 158 [Saccharum offici- 
narum]. — Drake and Hambleton 1945, p. 362 [El Salvador; Guate- 
mala; Tex.; Peru; maize]; 1946b, p. 122. — Box 1953, p. 37. 
Leptodictya (Hanuala) bambusae: Drake 1931e, pp. 119, 121. 
Type: Holotype o 71 , macropterous; Mayaguez, Puerto Rico; Drake Coll. 
(USNM). 

Distribution: U.S. (Tex.); Puerto Rico; Cuba; Haiti; Mexico; Guate- 
mala; El Salvador; Colombia; Ecuador; Venezuela; Peru. 

Host Plants: Bambusa vulgaris; Dendrocalamus sir ictus; Saccharum qffici- 
narum; maize. 
*Leptodictya (Hanuala) championi Drake 

Leptodictya championi Drake 1928a, p. 46 [Guatemala]. 

Leptodictya (Hanuala) championi: Drake 1931 e, p. 121. 
Type: Holotype o 71 , macropterous; Purula, Vera Paz, Guatemala; 
Drake Coll. (USNM). 
Distribution : Guatemala. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) circumcincta Champion 

Leptodictya circumcincta Champion 1897, p. 24, pi. 2, figs. 12, 12a [Pana- 
ma].— Drake and Ruhoff 1962d, p. 489. 

Leptodictya fusca Drake 1928a, p. 50 [Canal Zone]. 

Leptodictya (Hanuala) fusca: Drake 1931e, p. 121. 

Leptodictya (Hanuala) circumcincta: Drake 1931e, p. 121. 
Type: Sex unknown; San Felix, Chiriqui, Panama; British Mus. 
Distribution: Panama; Canal Zone; Ecuador. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) colombiana Drake 

Leptodictya colombiana Drake 1928a, p. 48 [Colombia]. 

Leptodictya (Hanuala) colombiana: Drake 1931 e, p. 121. 
Type: Holotype o 71 , macropterous; Colombia; Drake Coll. (USNM). 
Distribution: Colombia; Peru; Ecuador. 
Host Plant: Kidney bean. 



262 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptodictya (Hanuala) comes Drake and Hambleton (emendation) 
Leptodictya comitis Drake and Hambleton 1938b, p. 54, pi. 10, fig. e 

[Brazil; Olyra; bamboo]. 
Leptodictya {Hanuala) comitis: Monte 1939b, p. 72; 1941e, p. 105. 
Type: Holotype o 71 , macropterous; Ponte Nova, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plants: Olyra sp.; bamboo. 

*Leptodictya (Hanuala) cretata Champion 

Leptodictya cretata Champion 1897, p. 23, pi. 2, figs. 11, 11a, b [Guate- 
mala]. 
Leptodictya {Hanuala) cretata: Drake 1931e, pp. 120, 121 [Costa Rica; 
bamboo]. 
Type: Sex unknown; Panajachel, Guatemala; British Mus. 
Distribution: Guatemala; Costa Rica. 
Host Plant: Bamboo. 

*Leptodictya (Hanuala) decor Drake and Hambleton (emendation) 

Leptodictya decor is Drake and Hambleton 1945, p. 364 [Peru; bamboo]. 

Type: Holotype d\ macropterous; Tingo Maria, Peru; Drake Coll. 
(USNM). 

Distribution: Peru. 

Host Plant: Bamboo. 

Leptodictya (Hanuala) dilatata Monte 

Leptodictya dilatata Monte 1942a, p. 97 [Brazil]. 
Type: Holotype 9; Nova Friburgo, Estado do Rio, Brazil; Monte Coll. 
(Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Leptodictya (Hanuala) dohrni (Stal) 

Monanthia {Physatocheila) dohrni Stal 1858, p. 62 [Brazil]. 

Monanthia dohrni: Walker 1873a, p. 193. 

Leptodictya dohrnii [sic]: Stal 1873, p. 127.— Drake 1931b, p. 511. 

Leptodictya dohrni: Lethierry and Severin 1896, p. 14. — Drake and Poor 
1937d, p. 307, pi. 36, fig. 1.— Monte 1941b, pp. 101-106, fig. 

Leptodictya {Hanuala) dohrnii [sic]: Drake 1931e, p. 121. 
Type: Holotype 9; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil; Peru. 
Host Plant: Unrecorded. 

*Leptodictya (Hanuala) dola Drake and Hambleton 

Leptodictya dola Drake and Hambleton 1939, p. 159 [Brazil; Panicum 

maximum]. 
Leptodictya {Hanuala) dola: Monte 1941e, p. 105. 



LACEBUGS OF THE WORLD 263 

Type: Holotype o 71 , macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Panicum maximum. 
*Leptodictya (Hanuala) ecuadoris Drake and Hambleton 

Leptodictya ecuadoris Drake and Hambleton 1945, p. 362 [Ecuador; 
bamboo]. 
Type: Holotype d% macropterous; Vinces, Ecuador; Drake Coll. 
(USNM). 
Distribution: Ecuador; Brazil. 
Host Plant: Bamboo. 
*Leptodictya (Hanuala) elitha Drake and Runoff 

Leptodictya elitha Drake and Ruhoff 1962c, p. 136 [Peru]. 
Type: Holotype cf, macropterous; Queros, Rio Cosnipata, Dept. Cuszo, 
Peru; Drake Coll. (USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) evidens Drake 

Leptodictya evidens Drake 1928a, p. 49 [Panama]. 
Leptodictya {Hanuala) evidens: Drake 1931a, p. 121. 
Type: Holotype 9, macropterous; Tabernilla, Panama; Drake Coll. 
(USNM). 
Distribution: Panama: Peru. 
Host Plant: Unrecorded. 
Leptodictya (Hanuala) faceta Monte 

Leptodictya faceta Monte 1943d, p. 270 [Brazil]. 
Type: Holotype cf; Palmerias, Ipaussu, Sao Paulo, Brazil; Monte Coll. 
(Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) formositis Drake (emendation) 
Leptodictya formosatis Drake 1928a, p. 50 [Ecuador]. 
Leptodictya {Hanuala) formosatis: Drake 1931e, p. 121. 
Type: Holotype 9, macropterous; Mera, Ecuador; Drake Coll. (USNM). 
Distribution: Ecuador. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) fraterna Monte 

Leptodictya fralerna Monte 1941d, p. 96, fig. 3 [Costa Rica]. — Drake and 
Hambleton 1945, p. 362 [bamboo]. 
Type: Holotype cf, macropterous; San Isidro de Coronado, Alajuela, 
Costa Rica; Monte Coll. (Mus. Nacional). 
Distribution: Costa Rica. 
Host Plant: Bamboo. 



264 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptodictya (Hanuala) fuscocincta (Stal) 

Monanthia {Physatocheila) fusco-cincta Stal 1858, p. 62 [Brazil]. 
Monanthia fuscocincta: Walker 1873a, p. 193. 

Leptodictya fuscocincta: Stal 1873, p. 127. — Champion 1898b, p. 60, pi. 2, 
fig. 9. — Drake and Poor 1937d, p. 307. — Drake and Hambleton 
1938b, p. 53 [bamboo].— Monte 1938b, p. 131; 1941b, p. 104. 
Leptodictya {Hanuala) fuscocincta: Drake 1931e, p. 121. 
Type: Holotype 9; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Bamboo. 
*Leptodictya (Hanuala) galerita Drake 

Leptodictya galerita Drake 1942a, p. 19 [Paraguay]. 
Type: Holotype d", macropterous; Horqueta, Paraguay; Drake Coll. 
(USNM). 
Distribution : Paraguay. 
Host Plant: Unrecorded. 
:!: Leptodictya (Hanuala) grandatis Drake 

Leptodictya grandatis Drake 1931b, p. 511 [Peru]. 

Type: Holotype cf, macropterous; Tambo Enefias, Cam. del Pichis, 
Peru; Cornell Univ. 

Distribution: Peru. 

Host Plant: Unrecorded. 
Leptodictya (Hanuala) intermedia Monte 

Leptodictya intermedia Monte 1943d, p. 272 [Brazil]. 

Type: Holotypecf; Nova Teutonia, S. Catarina, Brazil; Monte Coll. 
(Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 
*Leptodictya (Hanuala) laidis Drake and Hambleton 

Leptodictya laidis Drake and Hambleton 1945, p. 362 [Colombia; 
bamboo]. 
Type: Holotype o 71 , macropterous; Villavicencio, Colombia; Drake Coll. 
(USNM). 
Distribution: Colombia. 
Host Plant: Bamboo. 
*Leptodictya (Hanuala) leinahoni (Kirkaldy) 

Hanuala leinahoni Kirkaldy 1905, p. 217 [Peru; Bolivia]. 
Leptodictya leinahoni: Drake 1922b, p. 42. — Monte 1938b, p. 131. 
Leptodictya {Hanuala) leinahoni: Drake 1931c, p. 226; 1931e, p. 121 
[Ecuador]. — Drake and Davis 1960, fig. 61. 
Type: Lectotype 9, macropterous; Mapiri, Bolivia; USNM. 
Distribution: Bolivia; Peru; Ecuador. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 265 

Leptodictya (Hanuala) lepida (Stal) 

Monanthia {Physatocheila) lepida Stal 1858, p. 63 [Brazil]. 

Monanthia lepida: Walker 1873a, p. 193. 

Leptodictya lepida: Stal 1873, p. 127.— Drake and Poor 1937d, p. 307, 

pi. 36, fig. 2. 
Leptodictya {Hanuala) lepida: Drake 1931 e, p. 121. 
Type: Sex unknown; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) litigiosa Monte 

Leptodictya litigiosa Monte 1940b, p. 380 [Brazil; bamboo]. — Silva 

1956, p. 36, fig. 12 [Phyllostachys castilloni]. 
Leptodictya {Hanuala) litigiosa: Monte 1941e, p. 106. 
Type: Holotypecf; Agua Preta, Estado da Baia, Brazil; Monte Coll. 
(Mus. Nacional). 
Distribution: Brazil; British Guiana. 
Host Plant: Phyllostachys castilloni; bamboo. 
*Leptodictya (Hanuala) lucida Drake and Hambleton 

Leptodictya lucida Drake and Hambleton 1945, p. 364 [Peru; bamboo]. 

Type: Holotypecf, macropterous; Tingo Maria, Peru; Drake Coll. 
(USNM). 

Distribution: Peru. 

Host Plant: Bamboo. 
*Leptodictya (Hanuala) luculenta Drake 

Leptodictya luculenta Drake 1928a, p. 49 [Ecuador]. 
Leptodictya {Hanuala) luculenta: Drake 1931e, p. 121. 
Type: Holotypecf, macropterous; Mera, Ecuador; Drake Coll. (USNM). 
Distribution: Ecuador; Peru. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) madelinae Drake 

Leptodictya madelinae Drake 1928a, p. 47 [Ecuador]; 1931b, p. 511 

[Peru]. 
Leptodictya {Hanuala) madelinae: Drake 1931 e, p. 121 [grass]. 
Type: Holotypecf, macropterous; Banos, Ecuador; Drake Coll. (USNM). 
Distribution: Ecuador; Peru. 
Host Plant: Grass. 
*Leptodictya (Hanuala) madra Drake and Hambleton 

Leptodictya madra Drake and Hambleton 1939, p. 158 [Brazil; Olyra]. 
Leptodictya {Hanuala) madra: Monte 1941e, p. 106. 
Type: Holotypecf, macropterous; Belem, Para, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Olyra sp. 

681-552—65 18 



266 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptodictya (Hanuala) nema Drake and Hambleton 

Leptodictya nema Drake and Hambleton 1939, p. 158 [Brazil]. 

Leptodictya {Hanuala) nema: Monte 1941 e, p. 106. 
Type: Holotype 9, macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Leptodictya (Hanuala) nicholi Drake 

Leptodictya nicholi Drake 1926c, p. 126 [Ariz.]. 

Leptodictya {Hanuala) nicholi: Drake 1931e, p. 121. 
Type: Holotype o 71 , macropterous; Santa Rita Mountains, Arizona, 
U.S.; Drake Coll. (USNM). 
Distribution: U.S. (Ariz.). 
Host Plant: Unrecorded. 

Leptodictya (Hanuala) nigra Monte 

Leptodictya nigra Monte 1941d, p. 97 [Costa Rica]. 
Type: Holotype cf; San Isidro de Coronado, Alajuela, Costa Rica; 
Monte Coll. (Mus. Nacional). 
Distribution: Costa Rica. 
Host Plant: Unrecorded. 

*Leptodictya (Hanuala) nigrosis Drake and Hambleton 

Leptodictya nigrosis Drake and Hambleton 1945, p. 363 [Peru; bamboo]. 
Type: Holotype cf, macropterous; Tingo Maria, Peru; Drake Coll. 
(USNM). 
Distribution: Peru. 
Host Plant: Bamboo. 

*Leptodictya (Hanuala) nota Drake and Hambleton 

Leptodictya nota Drake and Hambleton 1939, p. 157 [Brazil; Olyra]. 

Leptodictya {Hanuala) nota: Monte 1941e, p. 106. 
Type: Holotype d\ macropterous; Belem, Para, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Olyra sp. 

* Leptodictya (Hanuala) olyrae Drake 

Leptodictya {Hanuala) olyrae Drake 1931 e, p. 121 [Brazil; Olyra]. 
Leptodictya olyrae: Costa Lima 1936, p. 128. — Drake and Hambleton 
1938a, p. 48; 1938b, p. 54 —Monte 1938b, p. 131 [Olyra latifolia].— 
Silva 1956, p. 38. 
Type: Holotype 9, macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. ( 
Host Plants: Olyra latifolia; Olyra sp. 



LACEBUGS OF THE WORLD 267 

*Leptodictya (Hanuala) parilis Drake and Hambleton 

Leptodictya parilis Drake and Hambleton 1945, p. 363 [El Salvador; 
bamboo]. 
Type: Holotype o 71 , macropterous; San Andres, El Salvador; Drake Coll. 
(USNM). 
Distribution: El Salvador. 
Host Plant: Bamboo. 
*Leptodictya (Hanuala) paspalii Drake and Hambleton 

Leptodictya paspalii Drake and Hambleton 1934, p. 443 [Brazil; 

Paspalum]. — Costa Lima 1936, p. 128. 
Leptodictya (Hanuala) paspalii: Monte 1939b, p. 72; 1941e, p. 107. 
Type: Holotype o 71 , macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Paspalum sp. 
*Leptodictya (Hanuala) paulana Drake and Hambleton 

Leptodictya paulana Drake and Hambleton 1944b, p. 125 [Brazil]. 
Type: Holotype o 71 , macropterous; Taquaretinga, Sao Paulo, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) perita Drake 

Leptodictya perita Drake 1935, p. 17 [Brazil].— Monte 1940b, p. 380. 
Leptodictya (Hanuala) perita: Monte 1939b, p. 72; 1941e, p. 107. 
Type: Holotype o 71 ; Rio Grande do Sul, Brazil; Vienna Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) plana Heidemann 

Leptodictya plana Heidemann 1913b, p. 1, fig. 1 [Okla.]. — Drake 1925c, 

p. 38 [Miss.].— Blatchley 1926, p. 478, fig. 108. 
Leptodictya (Hanuala) plana: Drake 1931e, p. 121 [Fla.; Ala.; Tex.]. 
Type: Holotype d\ macropterous; Wistar, Indian Territory, U.S.; 
USNM. 

Distribution: U.S. (Okla.; Tex.; Ariz.; Kans.; Ala.; Miss.; Fla.); 
Mexico. 

Host Plant: Gramineae. 
*Leptodictya (Hanuala) simulans Heidemann 

Leptodictya simulans Heidemann 1913b, p. 3, fig. 2 [Va.; Ala.; S.C.]. — 

Blatchley 1926, p. 478, fig. 109 [Miss.]. 
Leptodictya (Hanuala) simulans: Drake 1931e, pp. 120, 121 [Tex.; Fla.; 
N.C.; bamboo]. 
Type: Holotype o 71 , macropterous; Drummond, Virginia, U.S.; USNM. 
Distribution: U.S. (Va.; N.C.; S.C; Fla.; Ala.; Miss.; Tex.). 
Host Plant: Bamboo. 



268 U.S. NATIONAL MUSEUM BULLETIN 243 

^Leptodictya (Hanuala) sinaloana Drake 

Leptodictya sinaloana Drake 1954d, p. 75 [Mexico]. 

Type: Holotype cf, macropterous; Sinaloa, Mexico; USNM. 

Distribution: Mexico. 

Host Plant: Unrecorded. 

Note: Found on orchid plants at port-of-entry, Nogales, Ariz. [Drake 
1954d]; found on Tillandsia usneoides from Mexico, at port-of-entry, San 
Pedro, Calif. 
*Leptodictya (Hanuala) socorrona Drake 

Leptodictya socorrona Drake 1948c, p. 23 [Revilla Gigedo]. 

Type: Holotype o 71 ; Socorro Island, Mexico; Cal. Acad. 

Distribution: Mexico (Revilla Gigedo Islands). 

Host Plant: Unrecorded. 

*Leptodictya (Hanuala) sodalatis Drake 

Leptodictya sodalatis Drake 1928a, p. 48 [Bolivia]. 
Leptodictya (Hanuala) sodalatis: Drake 1931 e, p. 121. 
Type: Holotypecf, macropterous; Cochabamba, Bolivia; Drake Coll. 
(USNM). 
Distribution: Bolivia. 
Host Plant: Unrecorded. 

*Leptodictya (Hanuala) solita Drake and Hambleton 

Leptodictya solita Drake and Hambleton 1938b, p. 53, pi. 9, fig. d 

[Brazil; Olyra]. 
Leptodictya (Hanuala) solita: Monte 1939b, p. 72; 1941e, p. 107. 
Type: Holotype 9, macropterous; Ponte Nova, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Olyra sp. 

* Leptodictya (Hanuala) tabida ( Herri ch-Schaeffer) 

Monanthia tabida Herrich-Schaeffer 1840, p. 86, pi. 173, fig. 535 [Mex- 
ico].— Stal 1873, p. 134.— Walker 1873a, p. 191. 

Monanthia (Phyllontocheila) tabida: Fieber 1844, p. 70, pi. 6, fig. 1. — 
Herrich-Schaeffer 1850, p. 155. 

Leptostyla tabida: Uhler 1886, p. 22. 

Leptodictya tabida: Champion 1897, p. 23, pi. 2, figs. 10, 10a [Guate- 
mala]; 1898b, p. 60.— Heidemann 1913a, p. 249, fig. 1 [Panama; 
sugar].— Drake 1925c, p. 38 [Tex.].— Drake and Hambleton 1946b, 
p. 122.— Box 1953, p. 37 [Venezuela].— Berry 1959, p. 145, fig. 126 
[El Salvador]. 

Leptodictya (Hanuala) tabida: Drake 1931e, pp. 119, 121 [Cuba; maize; 
bamboo]. 



LACEBUGS OF THE WORLD 269 

Type: Mexico; sex and deposition of type unknown. 
Distribution: Panama; El Salvador; Venezuela; Cuba; Guatemala; 
Mexico; U.S. (Tex.). 
Host Plants: Bamboo; maize; sugar. 

Leptodictya (Hanuala) tegeticula Monte 

Leptodictya tegeticula Monte 1943d, p. 271 [Brazil]. 
Type: Holotypeo"; Sao Paulo, Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Leptodictya (Hanuala) venezolana Monte 

Leptodictya venezolana Monte 1940d, p. 104 [Venezuela; Bambusa]. 

Type: HolotypecT; Caobos, Caracas, Venezuela; Monte Coll. (Mus. 
Nacional). 

Distribution: Venezuela. 

Host Plant: Bambusa sp. 

*Leptodictya (Hanuala) vulgata Drake 

Leptodictya vulgata Drake 1928a, p. 46 [Ecuador]. 
Leptodictya {Hanuala) vulgata: Drake 1931e, p. 121. 
Type: Holotypeo 71 , macropterous; Naranjapata, Ecuador; Drake Coll. 
(USNM). 
Distribution: Ecuador; Bolivia. 
Host Plant: Unrecorded. 
*Leptodictya (Hanuala) williamsi Drake 

Leptodictya williamsi Drake 1928a, p. 47 [Ecuador]. 
Leptodictya {Hanuala) williamsi: Drake 1931e, p. 121. 
Type: Holotypeo 71 , macropterous; Banos, Ecuador; Drake Coll. (USNM). 
Distribution: Ecuador. 
Host Plant: Unrecorded. 

Genus LEPTOPHARSA Stal 

Leptopharsa Stal 1873, pp. 122, 126. — Lethierry and Severin 1896, p. 

14.— Champion 1897, p. 21.— Drake 1928d, p. 21.— Costa Lima 

1936, p. 128.— Monte 1939b, p. 72; 1941e, p. 108.— Hurd 1946, p. 

465.— Bailey 1951, p. 35.— Silva 1956, pp. 38-41.— Drake and 

Ruhoff 1960a, p. 65. 
Leptostyla Stal 1873, pp. 120, 125. — Lethierry and Severin 1896, p. 

14. — Champion 1897, p. 11 (key). — Osborn and Drake 1916a, p. 

237.— Van Duzee 1917b, pp. 218, 817.— McAtee 1917a, p. 60 

(key).— Drake and Runoff 1960a, p. 65. 
Gelchossa Kirkaldy 1904, p. 280.— Drake 1922c, p. 372.— Parshley 

1923b, p. 704.— Blatchley 1926, p. 475 (key).— Drake and Ruhoff 

1960a, p. 58. 
Type Species: Leptopharsa elegantula Stal. 



270 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptopharsa albella Drake 

Leptopharsa albella Drake 1935, p. 18 [Paraguay]. 
Type: Holotype d"; S. Bernardino, Paraguay; Vienna Mus. 
Distribution : Paraguay. 
Host Plant: Unrecorded. 
*Leptopharsa angustata (Champion) 

Leptostyla angustata Champion 1897, p. 17, pi. 1, figs. 24, 24a [Guate- 
mala]. — Van Duzee 1907, p. 19 [Jamaica]. 
Gelchossa angustata: Drake and Bruner 1924a, p. 149. — Gowdy 1926a, 

p. 35. 
Leptopharsa angustata: Drake and Hambleton 1945, p. 360. — Hurd 1946, 
p. 466 [Artocarpus integrifolia]. 
Type: Sex unknown; San Geronimo, Guatemala; British Mus. 
Distribution: Guatemala; Jamaica. 
Host Plant: Artocarpus integrifolia. 
*Leptopharsa arta Drake and Poor 

Leptopharsa arta Drake and Poor 1942, p. 300 [Argentina; Brazil]. 
Type: Holotype cf; Loreto, Misiones, Argentina; Argentina Mus. 
Distribution: Argentina; Brazil. . 
Host Plant: Unrecorded. 
*Leptopharsa artocarpi Drake and Hambleton 

Leptopharsa artocarpi Drake and Hambleton 1938a, p. 48, fig. 1 [Brazil; 
Artocarpus integrifolia]. 
Type: Holotype 9, macropterous; Deodoro, E. do Rio, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Artocarpus integrifolia. 
*Leptopharsa atibaiae Drake and Ruhoff 

Leptopharsa atibaiae Drake and Ruhoff 1962d, p. 489 [Brazil]. 
Type: Holotype cf, macropterous; Atibaia, Sao Paulo, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
^Leptopharsa avia Drake 

Leptopharsa avia Drake 1953a, p. 15 [Brazil]. 
Type: Holotype d", macropterous; Belo Horizon te, Minas Gerais, 
Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
Leptopharsa bifasciata (Champion) 

Leptostyla bifasciata Champion 1897, p. 19, pi. 2, figs. 3,3a [Guatemala]. 
Leptopharsa bifasciata: Hurd 1946, p. 466. 
Type: Sex unknown; Senahu, Vera Paz, Guatemala; British Mus. 
Distribution: Guatemala. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 271 

*Leptopharsa bondari Drake and Poor 

Leptopharsa bondari Drake and Poor 1939a, p. 33 [Brazil]. 
Type: Holotype d", macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

^Leptopharsa bradleyi (Drake) 

Leptobyrsa bradleyi Drake 1931b, p. 513 [Peru]. 

Leptopharsa bradleyi: Drake and Poor 1937c, p. 163. 
Type: Holotype d% macropterous; Matucana, Peru; Cornell Univ. 
Distribution: Peru. 
Host Plant: Unrecorded. 

*Leptopharsa callangae Drake and Poor 

Leptopharsa callangae Drake and Poor 1940, p. 230 [Peru], 
Type: Holotype o 71 ; Callanga, Peru; Stockholm Mus. 
Distribution: Peru. 
Host Plant: Unrecorded. 

*Leptopharsa calopa Drake 

Leptopharsa calopa Drake 1928a, p. 54 [Brazil]. 
Type: Holotype o 71 ; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Leptopharsa clitoriae (Heidemann) 

Leptostyla clitoriae Heidemann 1911a, p. 137, pi. 10, fig. 6; 1911b, p. 180, 
fig. 4 [D.C.; Md.; Tex.; Clitoria mariana]. — Osborn and Drake 1916a, 
p. 239, fig. 8 [S.C.].— Van Duzee 1917b, p. 218 [Mass.; N.Y.; Pa.; 
Ark.].— McAtee 1917a, p. 61 [Lespedeza; Meibomia]; 1919, p. 143 
[Ind.].— Hussey 1922b, p. 23 [Mich.]. 
Leptostyla costofasciata Drake 1916, p. 326, fig. 1 [Tenn.]. 
Gelchossa clitoriae: McAtee 1923, p. 146.— Blatchley 1926, p. 476, fig. 

107— Blote 1945, p. 85. 
Leptopharsa clitoriae: Froeschner 1944, p. 669 [Mo.]. — Proctor 1946, 
p. 74 [Maine; alder]. — Bailey 1951, p. 36 [Conn.; Lappula]. 
Type: Holotype cf, macropterous; Rock Creek, District of Columbia, 
U.S.; USNM. 

Distribution: U.S. (Maine; Mass.; N.Y.; Pa.; Md.; D.C.; Va.; W. Va.; 
N.C.; S.C; Ark.; Miss.; Tenn.; Tex.; Kans.; Mo.; Iowa; Nebr.; Wis.; 
Mich.; 111.; Ind.; Ohio). 

Host Plants: Clitoria mariana; Meibomia sp.; Lespedeza sp.; Lappula sp.; 
alder. 
Note: Eggs [Heidemann 1911a; Bailey]. Taken at light [Bailey]. 



272 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptopharsa cognata Drake and Hambleton 

Leptopharsa cognata Drake and Hambleton 1934, p. 448 [Brazil; Miconia]. 
Type: Holotype 9, macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Miconia sp. 
*Leptopharsa constricta (Champion) 

Leptostyla constricta Champion 1897, p. 20, pi. 2, figs. 6, 6a [Guatemala; 

Panama]. — Van Duzee 1907, p. 19 [Jamaica]. 
Gelchossa constricta: Drake and Bruner 1924a, p. 149. — Gowdy 1926, 

p. 35. 
Leptopharsa constricta: Hurd 1946, p. 466. 
Type: Sex unknown; Pantaleon, Guatemala; British Mus. 
Distribution: Jamaica; Panama; Guatemala. 
Host Plant: Unrecorded. 

Note: Synonymy mentioned in Drake and Bruner incorrect. 
* Leptopharsa dampfi (Drake) 

Gelchossa dampfi Drake 1927b, p. 117 [Mexico]. 
Leptopharsa dampfi: Hurd 1946, p. 466. 
Type: Holotype cf, macropterous; Montana de Sumidero, Chiapas, 
Mexico; Drake Coll. (USNM). 
Distribution: Mexico. 
Host Plant: Unrecorded. 
^Leptopharsa dapsilis Drake and Hambleton 

Leptopharsa dapsilis Drake and Hambleton 1945, p. 359 [Guatemala; 
Olmediella betschleriana]. 
Type: Holotype o" , macropterous; Guatemala City, Guatemala; Drake 
Coll. (USNM). 
Distribution : Guatemala. 
Host Plant: Olmediella betschleriana. 
*Leptopharsa deca Drake and Hambleton 

Leptopharsa deca Drake and Hambleton 1945, p. 359 [Ecuador]. 
Type: Holotype o 71 , macropterous; Pichilingue, Ecuador; Drake Coll. 
(USNM). 
Distribution: Ecuador. 
Host Plant: Unrecorded. 
*Leptopharsa decens Drake and Hambleton 

Leptopharsa decens Drake and Hambleton 1938a, p. 58, fig. 4 [Brazil; 
Erythroxylon deciduum]. 
Type: Holotype cf, macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Erythroxylon deciduum. 



LACEBUGS OF THE WORLD 273 

*Leptopharsa delicata Monte 

Leptopharsa delicata Monte 1945a, p. 249, fig. 1 [Brazil]. 

Type: Holotype cf; Carmo do Rio Claro, Minas Gerais, Brazil; Monte 
Coll. (Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Leptopharsa difficilis Drake and Hambleton 

Leptopharsa difficilis Drake and Hambleton 1938b, p. 56, pi. 10, fig. F 
[Brazil; Chlorophora tinctoria]. — Monte 1942d, p. 108. 
Type: Holotype d\ macropterous; Campinas, Sao Paulo, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Chlorophora tinctoria. 

Leptopharsa dilaticollis (Champion) 

Leptostyla dilaticollis Champion 1897, p. 18, pi. 2, figs. 2, 2a [Guatemala]. 

Leptopharsa dilaticollis: Hurd 1946, p. 466. 
Type: Sex unknown; Cahabon, Vera Paz, Guatemala; British Mus. 
Distribution: Guatemala. 
Host Plant: Unrecorded. 

*Leptopharsa distans Drake (emendation) 

Leptopharsa distantis Drake 1928a, p. 53 [Mexico]. 

Type: Holotype cf, macropterous; Tamasopa, Mexico; Drake Coll. 
(USNM). 

Distribution: Mexico. 

Host Plant: Unrecorded. 

*Leptopharsa distinconis Drake 

Leptopharsa distinconis Drake 1928a, p. 54 [Brazil]. — Monte 1939b, p. 
73 [Rubiaceae]; 1947b, p. 432 [Siparana guianensis]. — Drake and 
Hambleton 1944b, p. 128. 
Leptopharsa iridis Drake 1930a, p. 2. 
Type: Holotype cf ; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plants: Siparuna guianensis; Rubiaceae. 

* Leptopharsa divisa (Champion) 

Leptostyla divisa Champion 1897, p. 19, pi. 2, figs. 4, 4a [Panama]. 
Leptopharsa divisa: Monte 1941d, p. 98 [Costa Rica]. — Drake and 
Hambleton 1945, p. 362 [Guatemala]; 1946b, p. 122 [Rubiaceae]. 
Type: Sex unknown; Volcan de Chiriqui, Panama; British Mus. 
Distribution: Panama; Guatemala; Costa Rica. 
Host Plant: Rubiaceae. 



274 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptopharsa elachys Drake and Ruhoff 

Leptopharsa elachys Drake and Ruhoff 1961b, p. 177, fig. 17b [South- 
West Africa]. 
Type: Holotype o 71 , macropterous; Windhoek, South- West Africa; 
Drake Coll. (USNM). 
Distribution: South- West Africa. 
Host Plant: Unrecorded. 

*Leptopharsa elata (Champion) 

Leptostyla elata Champion 1897, p. 16, pi. 1, figs. 21, 21 a, b 

[Mexico; Guatemala]. 
Leptopharsa elata: Monte 1942a, p. 94 [Costa Rica]. 
Type: Sex unknown; San Geronimo, Guatemala; British Mus. 
Distribution: Mexico; Guatemala; Costa Rica; El Salvador. 
Host Plant: Unrecorded. 

*Leptopharsa elegans (Hacker) 

Gelchossa elegans Hacker 1927, p. 27, pi. 7, fig. 7 [Queensland]. 

Type: Sex unknown; Upper Brookfield, Queensland, Australia; Queens- 
land Mus. 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

*Leptopharsa elegantula Stal 

Leptopharsa elegantula Stal 1873, p. 126 [Colombia]. — Champion 1898b, 
p. 60, pi. 2, fig. 8.— Drake and Poor 1937d, p. 308 [Bolivia; Brazil]. 
Type: Holotype o 71 ; Bogota, Colombia; Stockholm Mus. 
Distribution: Colombia; Bolivia; Brazil; Ecuador. 
Host Plant: Unrecorded. 

*Leptopharsa enodata Drake 

Leptopharsa enodata Drake 1942b, p. 364 [Queensland]. 
Type: Holotype o 71 ; North Pine River, Queensland, Australia; Queens- 
land Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Leptopharsa farameae Drake and Hambleton 

Leptopharsa farameae Drake and Hambleton 1938a, p. 62, fig. 11 [Brazil; 
Faramea] . 
Type: Holotype o", macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Faramea sp. 



LACEBUGS OF THE WORLD 275 

*Leptopharsa fici Drake and Hambleton 

Leptopharsa fici Drake and Hambleton 1938a, p. 50, fig. 2 [Brazil; Ficus 
elastica]. — Monte 1941e, p. 110 [Ficus subtriplinervia]. 
Type: Holotype o 71 , macropterous; Campinas, Sao Paulo, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 

Host Plants: Ficus elastica; Ficus subtriplinervia. 
* Leptopharsa fimbriata (Champion) 

Leptostyla fimbriata Champion 1897, p. 15, pi. 1, figs. 19, 19a [Mexico], 
Leptopharsa fimbriata: Hurd 1946, p. 466. 
Type: Sex unknown; Atoyac, Vera Cruz, Mexico; British Mus. 
Distribution: Mexico. 
Host Plant: Unrecorded. 
*Leptopharsa firma Drake and Hambleton 

Leptopharsa firma Drake and Hambleton 1938a, p. 53 [Brazil]. 
Type: Holotype cf, macropterous; Araraquara, Sao Paulo, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

Leptopharsa flava Monte 

Leptopharsa flava Monte 1940e, p. 291 [Brazil]. — Silva 1956, p. 40 
[coffee; Rubiaceae]. 
Type: Holotype o 71 ; Agua Preta, Baia, Brazil; Monte Coll. (Mus. 
Nacional) . 
Distribution: Brazil. 
Host Plants: Coffee; Rubiaceae. 

*Leptopharsa forsteroniae Drake and Hambleton 

Leptopharsa forsteroniae Drake and Hambleton 1938a, p. 51, fig. 6 
[Brazil; Forsteronia]. 
Type: Holotype cf , macropterous; Vicpsa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Forsteronia sp. 

*Leptopharsa fortis Drake and Hambleton 

Leptopharsa fortis Drake and Hambleton 1934, p. 447 [Brazil; Cephaelis 
hastisepala]. — Monte 1939b, p. 73 [Psychotria]. 
Type: Holotype o 71 . macropterous; Vicpsa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plants: Cephaelis hastisepala; Psychotria sp. 



276 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptopharsa furcata (Stal) 

Leptostyla furcata Stal 1873, p. 126 [Brazil].— Champion 1898b, p. 60, 

pi. 2, figs. 7, 7a. 
Gelchossa furcata: Costa Lima 1930, p. 84. 

Leptopharsa furcata: Costa Lima 1936, p. 128. — Drake and Poor 1937d, 
p. 308, pi. 36, fig. 7.— Drake and Hambleton 1938a, p. 55, fig. 5 — 
Monte 1940e, p. 292 [Rudgea villiflora]. 
Type: Sex unknown; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Rudgea villiflora. 
Leptopharsa furcata var. immatura (Stal) 

Leptostyla furcata var. immatura Stal 1873, p. 126 [Brazil]. 
Type: Sex unknown; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Leptopharsa furculata (Champion) 

Leptostyla fur culata Champion 1897, p. 20, pi. 2, figs. 5, 5 a, b [Panama; 

Guatemala] . 
Leptopharsa furculata: Hurd 1946, p. 466 [Rubiaceae]. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Panama; Guatemala; Mexico; Costa Rica. 
Host Plant: Rubiaceae. 
Leptopharsa fuscofasciata (Champion) 

Leptostyla fuscofasciata Champion 1897, p. 15, pi. 1, figs. 20, 20a 

[Panama]. 
Leptopharsa fuscofasciata: Hurd 1946, p. 466. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 
*Leptopharsa gracilenta (Champion) 

Leptostyla gracilenta Champion 1897, p. 17, pi. 1, figs. 23, 23a [Guate- 
mala] . 
Leptopharsa gracilenta: Drake and Bondar 1932, p. 91 [Brazil]. — Drake 
and Hambleton 1938a, p. 51 [Machaerium stipitatum]. 
Type: Sex unknown; Pantaleon, Guatemala; British Mus. 
Distribution: Guatemala; Brazil. 
Host Plant: Machaerium stipitatum. 
*Leptopharsa guatemalensis Drake and Poor 

Leptopharsa guatemalensis Drake and Poor 1939a, p. 34 [Guatemala]. 
Type: Holotype o", macropterous; Polochic River, Guatemala; Drake 
Coll. (USNM). 
Distribution: Guatemala. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 277 

*Leptopharsa heidemanni (Osborn and Drake) 

Leptostyla heidemanni Osborn and Drake 1916a, p. 238 [Ark.; D.C.; 
Mass.]. — McAtee 1917a, p. 63 [N.J.; La.; Baptisia tinctoria]; 1919, p. 
144.— Parshley 1917b, p. 56; 1919b, p. 70.— Barber 1922a, p. 17; 
1922b, p. 22. 
Gelchossa heidemanni: McAtee 1923, p. 146 [Md.]. — Parshley 1923b, 
p. 705, pi. 17, fig. 5 [Conn.].— Weiss and West 1924, p. 56 [Pa.].— 
Blatchley 1926, p. 477 [N.Y.].— Drake 1928b, p. 102.— Torre-Bueno 
1929, p. 311— Blote 1945, p. 85 [Va.]. 
Leptopharsa heidemanni: Froeschner 1944, p. 669 [Mo.]. — Hurd 1946, 
p. 466 [Ohio].— Bailey 1951, p. 37 [R.I.]. 
Type: Sex unknown; Arkansas, U.S.; Ohio State Univ. 
Distribution: U.S. (Mass.; R.I.; Conn.; N.Y.; N.J.; Pa.; Md.; D.C.; 

Va.; Ohio; Ind.; 111.; Mo.; Iowa; Ark.; Okla.; La.; Miss.). 
Host Plant: Baptisia tinctoria. 
Note: Life history studies [Weiss and West; Bailey]. 

*Leptopharsa heveae Drake and Poor 

Leptopharsa heveae Drake and Poor 1935, p. 283, fig. 1 [Brazil; Hevea 
braziliensis]. — Monte 1940b, p. 378. 
Type: Holotype cf, macropterous; Boa Vista Rio Tapajo, Brazil; 
USNM. 
Distribution: Brazil. 
Host Plant: Hevea braziliensis. 

*Leptopharsa hintoni Drake 

Leptopharsa hintoni Drake 1938a, p. 71 [Mexico]. — Hurd 1946, p. 466 
[Ariz.; Tex.]. 
Type: Holotype o"; Tejupilco, Temascaltepec, Mexico; Cal. Acad. 
Distribution: Mexico; U.S. (Ariz.; Tex.). 
Host Plant: Unrecorded. 

*Leptopharsa hoffmani Drake 

Leptopharsa hoffmani Drake 1928d, p. 21 [Haiti]. 
Type: Holotype cf , macropterous; Haiti; USNM. 
Distribution: Haiti. 
Host Plant: Unrecorded. 

*Leptopharsa hyaloptera (Stal) 

Leptostyla hyaloptera Stal 1873, p. 126 [Brazil]. 

Leptopharsa hyaloptera: Drake and Poor 1937d, p. 308. — Drake and 
Hambleton 1938a, p. 56 [Psychotria suterella]. 
Type: Sex unknown; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Psychotria suterella. 



278 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptopharsa ignota Drake and Hambleton 

Leptopharsa ignota Drake and Hambleton 1934, p. 449 [Brazil; Psychotria 
hancorniaefolia] . 
Type: Holotypecf, macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Psychotria hancorniaefolia. 

* Leptopharsa inannana Drake 

Leptopharsa inannana Drake 1953a, p. 13 [Cuba]. 
Type: Holotypecf, macropterous; Imias, Cuba; Drake Coll. (USNM). 
Distribution: Cuba. 
Host Plant: Unrecorded. 

*Leptopharsa inaudita Drake and Hambleton 

Leptopharsa inaudita Drake and Hambleton 1938a, p. 62 [Brazil]. 

Type: Holotypecf, macropterous; . Itarare, Sao Paulo, Brazil; Drake 
Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Leptopharsa jubaris Drake and Hambleton 

Leptopharsa jubaris Drake and Hambleton 1945, p. 361 [Ecuador]. 

Type: Holotypecf, macropterous; Pichilingue, Ecuador; Drake Coll. 
(USNM). 

Distribution : Ecuador. 

Host Plant: Unrecorded. 

*Leptopharsa laureata Drake and Hambleton 

Leptopharsa laureata Drake and Hambleton 1945, p. 360 [Peru]. 
Type: Holotypecf, macropterous; Pucallpa, Peru; Drake Coll. (USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 

*Leptopharsa lauta Drake and Hambleton 

Leptopharsa lauta Drake and Hambleton 1945, p. 361 [Ecuador]. 

Type: Holotypecf, macropterous; Vinces, Ecuador; Drake Coll. 
(USNM). 

Distribution: Ecuador. 

Host Plant: Unrecorded. 

* Leptopharsa lenitis Drake (emendation) 

Leptopharsa lenatis Drake 1930d, p. 272 [Peru; Brazil]. 
Type: Holotype 9, macropterous; Peru; Drake Coll. (USNM). 
Distribution: Peru; Brazil. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 279 

*Leptopharsa lineata (Champion) 

Leptostyla lineata Champion 1897, p. 17, pi. 1, figs. 22, 22a [Guate- 
mala]. — Osborn and Drake 1915b, p. 536. 
Leptopharsa lineata: Monte 1939b, p. 74 [Gramineae; Brazil]. — Drake 
and Hambleton 1946b, p. 122. 
Type: Sex unknown; San Isidro, Guatemala; British Mus. 
Distribution: Guatemala; Brazil; Peru; Argentina. 
Host Plant: Gramineae. 

Leptopharsa livida Monte 

Leptopharsa livida Monte 1943d, p. 265, fig. 3 [Brazil]. 

Type: Holotypeo 71 ; Vigosa, Minas Gerais, Brazil; Monte Coll. (Mus. 
Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

* Leptopharsa longipennis (Champion) 

Leptostyla longipennis Champion 1897, p. 13, pi. 1, figs. 16, 16a,b 

[Guatemala] . 
Leptopharsa longipennis: Hurd 1946, p. 466. 
Type: Sex unknown; Panajachel, Guatemala; British Mus. 
Distribution: Guatemala. 
Host Plant: Unrecorded. 

*Leptopharsa luxa Drake and Hambleton 

Leptopharsa luxa Drake and Hambleton 1945, p. 361 [Ecuador]. 

Type: Holotypecf, macropterous; Vinces, Ecuador; Drake Coll. 
(USNM). 

Distribution: Ecuador. 

Host Plant: Unrecorded. 

^Leptopharsa machaerii Drake and Hambleton 

Leptopharsa machaerii Drake and Hambleton 1934, p. 446 [Brazil; 
Machaerium nictitans]. 
Type: Holotypecf, macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Machaerium nictitans. 

*Leptopharsa machalana Drake and Hambleton 

Leptopharsa machalana Drake and Hambleton 1946a, p. 12 [Ecuador; 
Desmodium] . 
Type: Holotype cf, macropterous; Machala, Ecuador; Drake Coll. 
(USNM). 
Distribution: Ecuador. 
Host Plant: Desmodium sp. 



280 U.S. NATIONAL MUSEUM BULLETIN 243 

:!: Leptopharsa machalana var. vinnula Drake and Hambleton 

Lcptopharsa machalana var. vinnula Drake and Hambleton 1946a, p. 13 
[Fla.; Guatemala; Desmodium; Caesar's burr; beggars' lice]. — Stahler 
1946, p. 545, fig. 1 [Boehmeria cylindrica drummondiana; Urena lobala; 
Bidens pilosa radiata]. 
Leptophersa [sic] vinnula: Gibson and Carrillo 1959, p. 19 [Mexico]. 
Type: Holotype d% macropterous; Fort Pierce, Florida, U.S.; USNM. 
Distribution: U.S. (Fla.); Mexico; Guatemala. 

Host Plants: Bidens pilosa radiata; Boehmeria cylindrica drummondiana; 
Desmodium sp.; Urena lobala. 

Note: Fungus Hirsutella sp. found on adults [Stahler]. 
Leptopharsa marginella (Stal) 

Monanthia (Tropidocheila) marginella Stal 1858, p. 62 [Brazil], 
Monanthia marginella: Walker 1873a, p. 192. 

Leptopharsa marginella: Stal 1873, p. 126. — Drake and Poor 1937d, 
p. 309, pi. 36, fig. 6. 
Type: Sex unknown; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Leptopharsa miconiae Drake and Hambleton 

Leptopharsa miconiae Drake and Hambleton 1938b, p. 55, fig. 2 [Brazil; 
Miconia]. 
Type: Holotype o", macropterous; Alto da Serra, Sao Paulo, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Miconia sp. 
*Leptopharsa milled Drake 

Leptopharsa milleri Drake 1954f, p. 6 [Congo; Rhodesia; polony]. 
Type: Holotype 9; Salisbury, Southern Rhodesia; British Mus. 
Distribution: Southern Rhodesia; Congo. 
Host Plant: Polony tree. 
*Leptopharsa mira Drake and Hambleton Figure 2 

Leptopharsa mira Drake and Hambleton 1934, p. 446 [Brazil; Swartzia] . — 
Monte 1939b, p. 74 [Cassia; Annonaceae]. 
Type: Holotype cf , macropterous; Vicpsa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 

Host Plants: Cassia sp.; Swartzia sp.; Annonaceae. 
*Leptopharsa modica Drake and Hambleton 

Leptopharsa modica Drake and Hambleton 1939, p. 161 [Brazil]. 
Type: Holotype cf, macropterous; Sao Paulo, Brazil ; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 281 

*Leptopharsa nota Drake and Hambleton 

Leptopharsa nota Drake and Hambleton 1938a, p. 52 [Brazil; Rudgea 
blanchetiana]. 
Type: Holotype d", macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Rudgea blanchetiana. 

*Leptopharsa oblonga (Say) 

Tingis oblonga Say 1825, p. 325.— Le Conte 1859, p. 248 [Mo.]. 
Leptostyla oblonga: Stal 1873, p. 126 [NJ.].— Provancher 1886, p. 159 
[Canada].— Van Duzee 1912, p. 323.— Drake 1916, p. 327 [Ark.; 
D.C.].— Osborn and Drake 1916a, p. 238.— McAtee 1917a, p. 62 
[Nebr. Iowa; 111.; Wis.; Md.; Falcata comosa; Petalostemon]; 1919, 
p. 144.— Parshley 1922b, p. 11 [S. Dak.]. 
Gelchossa oblonga: McAtee 1923, p. 147 [Va.].— Drake 1925c, p. 37 
[Amorpha fruticosa]. — Blatchley 1926, p. 477 [Ind.; Kuhnistera], — 
Blote 1945, p. 85. 
Leptopharsa oblonga: Froeschner 1944, p. 669. — Bailey 1951, p. 39. 
Type: Missouri, U.S.; sex and deposition of type unknown. 
Distribution: U.S. (NJ.; Md.; D.C; Va.; Ind.; 111.; Wis.; Ark.; Mo.; 
Nebr.; Iowa; S. Dak.); Canada. 

Host Plants: Amorpha fruticosa; Falcata comosa; Kuhnistera sp.; Petalo- 
stemon sp. 

*Leptopharsa ocoteae Drake and Hambleton 

Leptopharsa ocoteae Drake and Hambleton 1938a, p. 59, fig. 9 [Brazil; 
Ocotea pretiosa]. 
Type: Holotype cf, macropterous; Vicpsa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Ocotea pretiosa. 

*Leptopharsa ogloblini Drake 

Leptopharsa ogloblini Drake 1936b, p. 701 [Argentina]. — Drake and 
Hambleton 1938a, p. 51 [Brazil; Machaerium pedicillatum]. 
Type: Holotype 9, macropterous; Loreto, Argentina; Leningrad Inst. 
Distribution: Argentina; Brazil. 
Host Plant: Machaerium pedicillatum. 

Leptopharsa ornata Monte 

Leptopharsa ornata Monte 1940e, p. 294 [Bolivia]. 

Type: Holotype 9; Sur Yungas, Chulumani, Bolivia; Monte Coll. (Mus. 
Nacional). 

Distribution: Bolivia. 

Host Plant: Unrecorded. 

681-552—65 19 



282 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptopharsa ovantis Drake and Hambleton 

Leptopharsa ovantis Drake and Hambleton 1945, p. 360 [Peru]. 
Type: Holotype d\ macropterous; Tingo Maria, Peru; Drake Coll. 
(USNM). 
Distribution: Peru; Guatemala. 
Host Plant: Unrecorded. 
*Leptopharsa pacis Drake and Hambleton 

Leptopharsa pads Drake and Hambleton 1939a, p. 160 [Brazil]. — Drake 

1948j, p. 434.— Silva 1956, p. 40. 
Leptopharsa perbona (not Drake): Monte 1944b, p. 461, figs. 2, 3. 
Type: Holotype 9, macropterous; Baia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
Leptopharsa pallens Monte 

Leptopharsa pal lens Monte 1943d, p. 267 [Peru]. 
Type: Holotype 9; Satipo, Dep. Junin, Peru; Monte Coll. (Mus. 
Nacional). 
Distribution: Peru. 
Host Plant: Unrecorded. 
*Leptopharsa papella Drake 

Leptopharsa papella Drake 1941a, p. 143 [Md.]. — Hurd 1946, p. 466 
[Ind.]. 
Type: Holotype 9, macropterous; Ashton, Maryland, U.S.; USNM. 
Distribution: U.S. (Ind.; Md.). 
Host Plant: Unrecorded. 
* Leptopharsa partita (Champion) 

Leptostyla partita Champion 1898a, p. 48, pi. 3, fig. 26 [Mexico]. 
Leptopharsa partita: Hurd 1946, p. 466. 
Type: Sex unknown; Pedregal, Mexico; British Mus. 
Distribution: Mexico. 
Host Plant: Unrecorded. 
*Leptopharsa paulana Drake 

Leptopharsa paulana Drake 1953a, p. 16 [Brazil]. 
Type: Holotype d", macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Leptopharsa pensa Drake and Hambleton 

Leptopharsa pensa Drake and Hambleton 1939, p. 160 [Brazil; Scleria 
myricocarpa]. 
Type: Holotype cf, macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Scleria myricocarpa. 



LACEBUGS OF THE WORLD 283 

*Leptopharsa perbona Drake 

Leptopharsa perbona Drake 1930a, p. 2 [Brazil]; 1948j, p. 434. — Drake 
and Bondar 1932, p. 91 [Rudgea].— Drake and Hambleton 1944b, 
p. 128. 
Leptopharsa spectabilis Monte 1940e, p. 290, fig. 7. 
Type: Holotype cf ; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Rudgea sp. 

*Leptopharsa peruensis Drake 

Leptopharsa peruensis Drake 1928a, p. 55 [Peru]. 
Type: Holotype 9, macropterous; Peru; Drake Coll. (USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 

*Leptopharsa posoqueriae Drake and Hambleton 

Leptopharsa posoqueriae Drake and Hambleton 1938a, p. 56, fig. 8 [Brazil; 
Posoqueria acutifolia]. 
Type: Holotype o 71 , macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Posoqueria acutifolia. 

*Leptopharsa principis Drake and Hambleton 

Leptopharsa principis Drake and Hambleton 1938a, p. 54 [Brazil], — 
Monte 1939b, p. 75 [Cassia]. 
Type: Holotype cf , macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Cassia sp. 

*Leptopharsa probala Drake and Hambleton 

Leptopharsa probata Drake and Hambleton 1938a, p. 57, fig. 3 [Brazil]. — 
Monte 1939b, p. 75 [Leguminosae]. 
Type: Holotype 9, macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Leguminosae. 

*Leptopharsa psychotriae Drake and Hambleton 

Leptopharsa psychotriae Drake and Hambleton 1939, p. 161 [Brazil; 
Psychotria] . 
Type: Holotype cf, macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Psychotria sp. 



2S4 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptopharsa pudens Drake and Hambleton 

Leptopharsa pudens Drake and Hambleton 1938a, p. 52 [Brazil; Machaer- 
itari] . 
Type: Holotype d\ macropterous; Sao Paulo, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Machaerium sp. 

*Leptopharsa quadrata Drake 

Leptopharsa quadrata Drake 1954f, p. 7 [Natal]. 
Type: Holotype 9, macropterous; Durban, Natal, South Africa; Drake 
Coll. (USNM). 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

*Leptopharsa retrusa Drake and Hambleton 

Leptopharsa retrusa Drake and Hambleton 1939, p. 159 [Brazil]. 
Leptodidya (Hanuala) retrusa: Monte 1941e, p. 107. 
Type: Holotype d\ macropterous; Belem, Para, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Leptopharsa reuniona Drake 

Leptopharsa reuniona Drake 1957e, p. 404 [Reunion]. 
Type: Holotype cF; Beloune, Reunion; Madagascar Sci. Inst. 
Distribution: Mascarene Islands (Reunion). 
Host Plant: Unrecorded. 

*Leptopharsa rudgeae Drake and Hambleton 

Leptopharsa rudgeae Drake and Hambleton 1934, p. 447 [Brazil; 
Rudgea].— Costa Lima 1936, p. 129. 
Type: Holotype cf, macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Rudgea sp. 

*Leptopharsa rumiana Drake and Hambleton 

Leptopharsa rumiana Drake and Hambleton 1946a, p. 11 [El Salvador; 
Guatemala; Afalvaviscus arbor ens]. 
Type: Holotype o", macropterous; San Andres, El Salvador; Drake Coll. 
(USNM). 
Distribution: El Salvador; Guatemala. 
Host Plant: Malvaviscus arboreus. 



LACEBUGS OF THE WORLD 285 

*Leptopharsa ruris Drake 

Leptopharsa ruris Drake 1942a, p. 20 [Antigua]. — Drake and Cobben 
1960, p. 83, fig. 83 [St. Martin; Crotonfiavens}. 
Type: Holotype cf, macropterous; Antigua, British West Indies; Drake 
Coll. (USNM). 
Distribution: Leeward Islands (Antigua; St. Martin). 
Host Plant: Crotonfiavens. 

*Leptopharsa satipona Drake and Hambleton 

Leptopharsa satipona Drake and Hambleton 1944b, p. 128 [Peru]. 
Type: Holotype d\ macropterous; Satipo, Peru; Drake Coll. (USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 

*Leptopharsa sera Drake and Poor 

Leptopharsa sera Drake and Poor 1939a, p. 35 [Brazil; Bolivia]. 
Type: Holotype 9, macropterous; Chapada, Brazil; Drake Coll. (USNM). 
Distribution: Brazil; Bolivia. 
Host Plant: Unrecorded. 

Leptopharsa setigera (Champion) 

Leptostyla setigera Champion 1897, p. 14, pi. 1, figs. 18, 18a [Panama]. 

Leptopharsa setigera: Hurd 1946, p. 466. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

*Leptopharsa siderea Drake and Hambleton 

Leptopharsa siderea Drake and Hambleton 1946a, p. 13 [Guatemala]. 

Type: Holotype 9, macropterous; Esquintla, Guatemala; Drake Coll. 
(USNM). 

Distribution: Guatemala. 

Host Plant: Unrecorded. 

*Leptopharsa simulans (Stal) 

Monanthia {Phyllontocheila) simulans Stal 1858, p. 61 [Brazil], 
Monanthia (Gargaphia) simulans: Stal 1862, p. 324. 
Monanthia simulans: Walker 1873a, p. 192. 

Gargaphia simulans: Stal 1873, p. 124. — Lethierry and Severin 1896, 
p. 13.— Drake and Poor 1937d, p. 310, pi. 36, fig. 8.— Monte 1941e, 
p. 119. 
Leptopharsa scita Drake and Hambleton 1939, p. 160. 
Leptopharsa simulans: Monte 1947a, p. 236. 
Type: Holotype 9; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 



286 U.S. NATIONAL MUSEUM BULLETIN 243 

Leptopharsa sobrina Monte 

Leptopharsa sobrina Monte 1940b, p. 377 [Brazil; Aspidosperma warmingii]. 
Type: Holotype 9; Belo Horizonte, Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Aspidosperma warmingii. 

*Leptopharsa tenuatis Drake 

Leptopharsa tenuatis Drake 1928a, p. 52 [Brazil] .—Monte 1939b, p. 75 
[Dalbergia]. 
Type: Holotype o" , macropterous; Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Dalbergia sp. 

* Leptopharsa tenuis (Champion) 

Leptostyla tenuis Champion 1897, p. 18, pi. 1, figs. 25, 25 a, b [Guatemala]. 
Leptopharsa tenuis: Drake and Hambleton 1945, p. 360 [Ichthyomethia 
grandifolia] . 
Type: Sex unknown; Capetillo, Guatemala; British Mus. 
Distribution: Guatemala. 
Host Plant: Ichthyomethia grandifolia. 

Leptopharsa unicarinata Champion 

Leptopharsa unicarinata Champion 1897, p. 21, pi. 2, figs. 7, 7a [Panama]. 
Type: Sex unknown; Volcan de Chiriqui, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

*Leptopharsa usingeri Drake 

Leptopharsa usingeri Drake 1938a, p. 72 [Mexico]. 
Type: Holotype 9; Temascal tepee, Mexico; Cal. Acad. 
Distribution: Mexico. 
Host Plant: Unrecorded. 

^Leptopharsa valida Drake and Hambleton 

Leptopharsa valida Drake and Hambleton 1938a, p. 54 [Brazil; Condylo- 
carpon rauwolfiae]. 
Type: Holotype cf, macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Condylocarpon rauwolfiae. 

Leptopharsa variegata Monte 

Leptopharsa variegata Monte 1943d, p. 267 [Brazil]. 
Type: Holotype cf; Sao Paulo, Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 287 

*Leptopharsa velifer (McAtee) 

Leptostyla velifer McAtee 1917a, p. 60 [Ariz.]. 
Leptopharsa velifer: Hurd 1946, p. 466. 
Type: Holotype 9, macropterous; Arizona, U.S.; USNM. 
Distribution: U.S. (Ariz.). 
Host Plant: Unrecorded. 
*Leptopharsa vicina Drake and Poor 

Leptopharsa vicina Drake and Poor 1939a, p. 33 [Haiti]. 
Type: Holotype d% macropterous; Williamson, Haiti; Drake Coll. 
(USNM). 
Distribution: Haiti. 
Host Plant: Unrecorded. 
Leptopharsa vittipennis (Stal) 

Leptostyla vittipennis Stal 1873, p. 126 [Brazil]. 

Leptopharsa vittipennis: Drake and Poor 1937d, p. 307, pi. 36, fig. 11 
[Peru].— Drake and Hambleton 1938a, p. 52.— Monte 1940e, p. 292. 
Leptopharsa vitipennis [sic]: Monte 1941e, p. 120. — Silva 1956, p. 41, fig. 
14 [Artocarpus integrifolia]. 
Type: Holotype cf; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil; Peru. 
Host Plant: Artocarpus integrifolia. 
^Leptopharsa zeteki Drake 

Leptopharsa zeteki Drake 1939f, p. 69 [Canal Zone]. 
Type: Holotype cf, macropterous; Barro Colorado Island, Panama 
Canal Zone; Drake Coll. (USNM). 
Distribution: Panama (Canal Zone). 
Host Plant: Unrecorded. 

Genus LEPTOYPHA Stal 

Leptoypha Stal 1873, pp. 121, 129. — Lethierry and Severin 1896, p 
21.— Van Duzee 1916, p. 26; 1917b, pp. 220, 817.— McAtee 1917a 
p. 55 (key).— Blatchley 1926, p. 498 (key).— Monte 1941e, p. 121.— 
Hurd 1946, p. 456.— Bailey 1951, p. 27.— Drake and Maa 1953, p 
94.— Drake and Runoff 1960a, p. 65.— Stichel 1960b, p. 400; 1960c 
p. 139. 
Tingis (Birgitta) Lindberg 1927, p. 18.— Drake and Ruhoff 1960a 
p. 40. 
Type Species: Tingis mutica Say. 
* Leptoypha anceps (Horvath) 

Paracopium ? anceps Horvath 1925a, p. 9 [Queensland]. 
Leptoypha anceps: Drake 1947c, p. 112. 
Type: Holotype o 71 ; Yarrabah, Queensland, Australia; Stockholm Mus 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 



288 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptoypha barberi Drake and Ruhoff 

Leptoypha brevicornis (not Champion): Barber 1910, p. 38 [Ariz.]. — Van 

Duzee 1917b, p. 220 (in part). 
Leptoypha minor (not McAtee 1917a): McAtee 1919, p. 142 (in part). — 

Hurd 1946, p. 457 [Fraxinus berlandierana]. 
Leptoypha barberi Drake and Ruhoff 1960d, p. 152, fig. 1. 
Type: Holotype o 71 , macropterous; Huachuca Mountains, Arizona, 
U.S.; Barber Coll. (USNM). 
Distribution: U.S. (Ariz.). 
Host Plant: Fraxinus berlandierana. 
*Leptoypha binotata Champion 

Leptoypha binotata Champion 1897, p. 32, pi. 2, fig. 27 [Guatemala]. 
Type: Sex unknown; Cerro Zunil, Guatemala; British Mus. 
Distribution: Guatemala. 
Host Plant: Unrecorded. 
*Leptoypha braziliensis Drake and Hambleton 

Leptoypha braziliensis Drake and Hambleton 1939, p. 154 [Brazil]. 
Type: Holotype 9, macropterous; Amazon region, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Leptoypha brevicornis Champion 

Leptoypha brevicornis Champion 1897, p. 32, pi. 2, fig. 28 [Mexico]. — Van 
Duzee 1917b, p. 220 (in part). 
Type: Sex unknown; Omilteme, Guerrero, Mexico; British Mus. 
Distribution: Mexico. 
Host Plant: Unrecorded. 
*Leptoypha capitata (Jakovlev) 

Monanthia capitata Jakovlev 1876a, p. 110 [Siberia]. 

Monosteira capitata: Lethierry and Severin 1896, p. 25. — Horvath 

1906a, p. 106.— Kiritshenko 1913b, p. 482. 
Monostira capitata: Oshanin 1908, p. 456; 1912, p. 46. 
Tingis (Birgitta) tenuimarginata Lindberg 1927, p. 19, fig. 5. 
Leptoypha capitata: Kiritshenko 1931, p. 269 [Malus manschurica; Pyrus 
ussuriensis]. — Drake and Maa 1953, p. 94 [China; Japan]. 
Type: Holotype 9, macropterous; Ussuri, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Siberia); Japan; China. 
Host Plants: Malus manschurica; Pyrus ussuriensis. 
*Leptoypha costata Parshley 

Leptoypha costata Parshley 1917a, 16 [Md.].— McAtee 1917a, p. 57; 
1919, p. 143 [ash]; 1923, p. 146 [Franxinus caroliniana].— Drake 1918d, 
p. 87 [Colo.; Ark.; 111.]; 1925c, p. 38 [Miss.].— Blatchley 1926, 
p. 498.— Bailey 1951, p. 27 [Conn.; La.; Hicoria alba}. 



LACEBUGS OF THE WORLD 289 

Leptoypha distinguenda Heidemann 1917, p. 218, pi. 17, fig. 1 [D.C.; 
Va.; Hamamelis]. 
Type: Holotype 9; Marshall Hall, Maryland, U.S.; MCZ. 
Distribution: U.S. (Conn.; Md.; D.C.; Va.; Miss.; La.; Ark.; Ill; 
Colo.). 
Host Plants: Fraxinus caroliniana; Hicoria alba; Hamamelis sp. 

*Leptoypha drakei McAtee 

Leptoypha mutica (not Say): Uhler 1893b, p. 264. 

Leptoypha brevicornis (not Champion): McAtee 1917a, p. 59; 1919, 

p. 143. 
Leptoypha drakei McAtee 1919, p. 143 [Calif.; Tex.].— Hurd 1946, 
p. 457 [Ariz.]. 
Type: Holotype cf, macropterous; Argus Mountains, California, U.S.; 
USNM. 
Distribution: U.S. (Calif.; Ariz.; Tex.). 
Host Plant: Ash. 
*Leptoypha elliptica McAtee 

Leptoypha elliptica McAtee 1917a, p. 57 [Tex.]; 1919, p. 143.— Drake 
1918d, p. 87 [Ga.; Fla.; Ilex].— Blatchley 1926, p. 499 [Ind.].— 
Froeschner 1944, p. 670 [Mo.]. 
Type: Holotype d\ macropterous; Texas, U.S.; USNM. 
Distribution: U.S. (Tex.; Fla.; Ga.; Tenn.; Ind.; Mo.). 
Host Plant: Ilex sp. 
^Leptoypha hospita Drake and Poor 

Leptoypha hospita Drake and Poor 1937a, p. 12 [Penang]; 1937b, p. 402. 
Type: Holotype 9, macropterous; Penang Island, Straits Settlements; 
USNM. 
Distribution: Federation of Malaya (Penang Island); China. 
Host Plant: Unrecorded. 
*Leptoypha ilicis Drake 

Leptoypha ilicis Drake 1919a, p. 420 [Ga.; Ilex].— Blatchley 1926, p. 500 
[Fla.]; 1928, p. 5 [Vaccinium].— Hurd 1946, p. 457 [Tex.; Okla.].— 
Bailey 1951, p. 28. 
Type: Holotype d^, macropterous; Stone Mountain, Georgia, U.S.; 
Drake Coll. (USNM). 
Distribution: U.S. (Ga.; Fla.; Tex.; Okla.). 
Host Plants: Ilex sp.; Vaccinium sp. 
*Leptoypha luzona Drake and Ruhoff 

Leptoypha luzona Drake and Ruhoff 1961b, p. 140 [Luzon]. 
Type: Holotype o 71 , macropterous; Los Banos, Philippines; Drake Coll. 
(USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 



290 U.S. NATIONAL MUSEUM BULLETIN 243 

*Leptoypha mcateei Drake 

Leptoypha mcateei Drake 1921b, p. 49 [Fla.; Osmanthus americana]. — 

Blatchley 1926, p. 499 [Xolisma jerruginea]. 
Leptoypha mcatella [sic]: Drake 1925c, p. 38. 
Type: Holotype d% macropterous; Gainesville, Florida, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Fla.). 
Host Plants: Osmanthus americana; Xolisma Jerruginea. 

*Leptoypha minor McAtee 

Leptoypha minor McAtee 1917a, p. 56 [Calif.]; 1919, p. 142 (in part).— 
Van Duzee 1917a, p. 260.— Drake 1918d, p. 87 (in part) .— Usinger 
1946b, p. 286, figs. [Populus candicans; Fraxinus velutina; Fraxinus 
oregona; olive]. — Drake and Ruhoff 1962a, p. 156. 
Leptoypha nubilis Drake 1941b, p. 141 [Ceanothus]. 
Type: Holotype cf, macropterous; Siskiyou County, California, U.S.; 
USNM. 
Distribution: U.S. (Calif.). 

Host Plants: Ceanothus sp.; Fraxinus oregona; Fraxinus velutina; Populus 
candicans; "olive tree next to infested ash." 
Note: Life history study [Usinger]. 

*Leptoypha morrisoni Drake 

Leptoypha morrisoni Drake 1922b, p. 43 [Dominican Republic]. — Drake 

and Bruner 1924a, p. 146.— Gowdy 1926, p. 35.— Drake and Ham- 

bleton 1939, p. 155 [Canal Zone].— Hurd 1946, p. 457 [Fla.]. 

Leptoypha binotata ? (not Champion): Van Duzee 1907, p. 21 [Jamaica]. 

Type: Holotype cf, macropterous; San Pedro de Macoris, Dominican 

Republic; USNM. 

Distribution: Dominican Republic; Panama (Canal Zone); Jamaica; 
Haiti; Cuba; U.S. (Fla.). 
Host Plant: Mangle. 

*Leptoypha mutica (Say) 

Tingis mutica Say 1832, p. 27 [Ind.].— Fitch 1858, p. 794.— Le Conte 
1859, p. 349. 

Monanthia mutica: Walker 1873a, p. 190. 

Leptoypha mutica: Stal 1873, p. 129 [Tex.].— Provancher 1886, p. 160 
[Que.].— Uhler 1886, p. 22.— Champion 1898b, p. 61.— Wirtner 
1904, p. 203 [Pa.].— Smith 1910, p. 149 [N.J.] .— Banks 1910, p. 57.— 
Van Duzee 1912, p. 323.— Dickerson and Weiss 1916, p. 308, pi. 16 
[Chionanthus virginica]. — McAtee 1917a, p. 58 [D.C.; Nebr.; Wis.; 
Ohio; Ont.; Tenn.; Fraxinus]; 1919, p. 143; 1923, p. 146.— Drake 
1918d, p. 87 [Mich.; 111.; Md.; Va.; Ga.; Adelia acuminata]; 1928b, 
p. 102 [N.Y.] — Barber 1914, p. 507 [Fla.].— Barber and Weiss 1922, 
p. 14, fig. 8.— Hussey 1922a, p. 11 [N. Dak.]; 1922b, p. 22 [Fraxinus 



LACEBUGS OF THE WORLD 291 

americana].— Parshley 1922a, p. 236 [Mass.]; 1922b, p. 11 [S. Dak.]; 
1923b, p. 706 [Conn.].— Blatchley 1926, p. 500, fig. 121— Froeschner 
1944, p. 670 [Mo.].— Bailey 1951, p. 28 [Maine; N.H.]; 1959, p. 63. 
Leptophya [sic] mutica: Osborn and Drake 1916a, p. 241, pi. 10, fig. b. 
Type: Indiana, U.S.; type apparentiy lost. 

Distribution: U.S. (Maine; N.H.; Mass.; Conn.; N.Y.; N.J.; Pa.; Md.; 
D.C.; Va.; N.C.; S.C.; Ga.; Fla.; Tenn.; Ind.; Ohio; 111.; Nebr.; Kans.; 
Wis.; Mich.; Mo.; Iowa; S. Dak.; N. Dak.; Tex.; Minn.); Canada (Ont.; 
Que.); Mexico (Socorro Island). 

Host Plants: Adelia acuminata; Chionanthus virginica; Fraxinus americana; 
Fraxinus sp. 

Note: Taken at light [Drake 1918d]. Life history studies [Dickerson and 
Weiss; Bailey]. 

*Leptoypha wuorentausi (Lindberg) 

Tingis (Birgitta) wuorentausi Lindberg 1927, p. 18, fig. 4 [Russia]. 
Tingis (Birgitta) crispifolii Takeya 1932, p. 12, pi. 1, fig. 3 [Korea; 

Fraxinus mandshurica japonica]; 1951a, p. 17. 
Tingis crispifolii: Saito 1933, p. 7 [Fraxinus pubinervis; Fraxinus rhyncho- 

phylla]. 
Leptoypha wuorentausi: Drake and Maa 1953, p. 95. 
Type: Sex unknown; Ussuri, Spasskaja, Russia; Helsin. Mus. 
Distribution: U.S.S.R. (Siberia); Korea. 

Host Plants: Fraxinus mandshurica japonica; Fraxinus pubinervis; Fraxinus 
rhynchophylla.\ 

Genus LEPTURGA Stal 

Lepturga Stal 1873, pp. 119, 124.— Drake 1943b, pp. 175-177.— Drake 
and Ruhoff 1960a, p. 66. 
Type Species: Lepturga nigritarsis Stal. 

*Lepturga dignata Drake 

Lepturga dignata Drake 1943b, p. 176 [New Guinea]; 1960, p. 367, fig. 19. 

Type: Holotype 9, macropterous; Madang, New Guinea; Drake Coll. 
(USNM). 

Distribution: New Guinea. 

Host Plant: Unrecorded. 

Note: The type locality (Drake 1943b) is incorrectly given as "Pedang, 
Pedang, New Guinea." The label on the type specimen reads "F. Wil- 
helmshaven, Germ. N. Guinea" which today is known as "Madang." 
The distribution record (Drake 1960) of "Penang I., Malaya" is also in 
error. 



292 U.S. NATIONAL MUSEUM BULLETIN 243 

*Lepturga magnifica (Hacker) 

Leptobyrsa magnifica Hacker 1929, p. 331, pi. 35, fig. 13 [Queensland]. 
Lepturga magnifica: Drake 1943b, p. 176. 
Type: Sex unknown; Mt. Tambourine, Queensland, Australia; Queens- 
land Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
:i Lepturga major (Hacker) 

Leptobyrsa major Hacker 1929, p. 332, pi. 35, fig. 14 [Queensland]. 
Lepturga major: Drake 1943b, p. 176. 
Type: Sex unknown; Blackbutt, Queensland, Australia; Queensland Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Lepturga nigritarsis Stal 

Lepturga nigritarsis Stal 1873, p. 124 [Queensland]. — Lethierry and 
Severin 1896, p. 13.— Drake 1943b, p. 175. 
Type: Holotype 9; Cape York, Australia; Stockholm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Genus LIOTINGIS Drake 

Liotingis Drake 1930d, p. 270.— Drake and Hambleton 1935, p. 146. — 
Monte 1939b, p. 76; 1941e, p. 122.— Drake and Ruhoff 1960a, p. 66. 
Type Species: Liotingis evidens Drake. 

*Liotingis affinata Drake and Hambleton 

Liotingis affinatus Drake and Hambleton 1935, p. 147 [Brazil; 
Aspidosperma]. 
Type: Holotype o 71 , macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Aspidosperma sp. 

*Liotingis aspidospermae Drake and Hambleton 

Liotingis aspidospermae Drake and Hambleton 1935, p. 147, fig. 3 [Brazil; 
Aspidosperma]. 
Type: Holotype cf, macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Aspidosperma sp. 

*Liotingis evidens Drake (emendation) 

Liotingis evidentis Drake 1930d, p. 271 [Brazil]. — Drake and Hambleton 
1935, p. 146; 1938a, p. 63.— Monte 1939b, p. 76 [Aspidosperma]. 



LACEBUGS OF THE WORLD 293 

Type: Holotype 9, macropterous; Minas Gerais, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Aspidosperma sp. 

*Liotingis immaculata (Drake and Hambleton) 

Allotingis immaculata Drake and Hambleton 1934, p. 443 [Brazil; 

Aspidosperma]. — Costa Lima 1936, p. 125. 
Liotingis immaculata: Drake and Hambleton 1935, p. 146. 
Type: Holotype 9 5 macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Aspidosperma sp. 

Genus LULLIUS Distant 

Lullius Distant 1904, p. 429.— Drake and Ruhoff 1960a, p. 24. 
Type Species: Lullius major Distant. 

*Lullius insolens Drake 

Lullius insolens Drake 1944a, p. 68 [Cape Province]. 

Type: Holotype o* , macropterous; Camps Bay, Cape Peninsula, Africa; 
Drake Coll. (USNM). 

Distribution: South Africa (Cape Province). 

Host Plant: Unrecorded. 

*Lullius major Distant 

Lullius major Distant 1904, p. 430, pi. 8, fig. 7 [Cape Province]. — Drake 
1956f, p. 429; 1961c, p. 130. 
Type: Sex unknown; Cape Colony, South Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

*Lullius spinifemur Drake Plate 15 

Lullius spinifemur Drake 1961c, p. 130, pi. 10 [Cape Province]. 

Type: Holotype o 71 ; Seven Weeks Poort, Laingsburg District, West Cape 
Province, South Africa; Natal Museum, Pietermaritzberg, Natal, South 
Africa. 

Distribution : South Africa (Cape Province) . 

Host Plant: Unrecorded. 

Genus MACROCORYTHA Stal 

Corythucha (Macrocorytha) Stal 1873, p. 123. 
Macrocorytha: Drake and Ruhoff 1960a, p. 66. 
Type Species: Tingis rhomboptera Fieber. 



294 U.S. NATIONAL MUSEUM BULLETIN 243 

Macrocorytha rhomboptera (Ficber) 

Tingis rhomboptera Fieber 1844, p. 103, pi. 8, figs. 37, 38 [Luzon; Mal- 
vaceae]. — Herrich-Schaeffer 1850, p. 161. 
Corythucha {Macrocorytha) rhomboptera: Stal 1873, p. 123. 
Corythucha rhomboptera: Lethierry and Severin 1896, p. 11. 
Macrocorytha rhomboptera: Drake and Runoff 1960a, p. 66. 
Type: Sex unknown; Luzon, Philippine Islands; Vienna Mus. 
Distribution: Philippine Islands (Luzon). 
Host Plant: Malvaceae. 

Genus MACROTINGIS Champion 

Macrotingis Champion 1897, p. 22. — Hurd 1946, p. 469. — Drake and 
Ruhoff 1960a, p. 66. 
Type Species: Macrotingis biseriata Champion. 

*Macrotingis biseriata Champion 

Macrotingis biseriata Champion 1897, p. 22, pi. 2, figs. 8, 8a, b [Pan- 
ama]. — Drake 1928e, p. 4 [Honduras]. 
Type: Sex unknown; Panama; British Mus. 
Distribution: Panama; Honduras. 
Host Plant: Unrecorded. 

*Macrotingis biseriata var. novicis Drake 

Macrotingis biseriata var. novicis Drake 1928e, p. 4 [Honduras]. 
Type: Holotype cf; Progreso, Honduras; Michigan Univ. 
Distribution: Honduras. 
Host Plant: Unrecorded. 
Note: Taken at light. 

Macrotingis uniseriata Champion 

Macrotingis uniseriata Champion 1897, p. 22, pi. 2, figs. 9, 9a [Guate- 
mala]. 
Type: Sex unknown; Capetilla, Guatemala; British Mus. 
Distribution: Guatemala. 
Host Plant: Unrecorded. 

*Macrotingis zeteki Drake 

Macrotingis zeteki Drake 1950b, p. 299 [Canal Zone]. 
Type: Holotype d", macropterous; Barro Colorado Island, Panama 

Canal Zone; USNM. 
Distribution: Panama (Canal Zone). 
Host Plant: Unrecorded. 

Genus MAFA Hesse 

Mafa Hesse 1925, p. 88.— Drake and Ruhoff 1960a, p. 67. 
Type Species: Mafa lanceolata Hesse. 



LACEBUGS OF THE WORLD 295 

*Mafa lanceolata Hesse 

Mqfa lanceolata Hesse 1925, p. 89, pi. 4, fig. 4 [South- West Africa]. 
Type: Sex unknown; brachypterous form; Mafa, Ovamboland, South- 
West Africa; So. Afr. Mus. 
Distribution: South- West Africa. 
Host Plant: Unrecorded. 

Genus MALANDIOLA Horvath 

Malandiola Horvath 1925a, p. 13.— Drake and Ruhoff 1960a, p. 67. 
Type Species: Malandiola simplex Horvath. 

♦Malandiola acares Drake and Ruhoff 

Malandiola acares Drake and Ruhoff 1962d, p. 494 [Queensland]. 

Type: Holotype cf , macropterous; Bluff, Queensland, Australia; Drake 
Coll. (USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

♦Malandiola minys Drake and Ruhoff 

Malandiola minys Drake and Ruhoff 1962d, p. 493 [Queensland]. 

Type: Holotype o 71 ; Castle Hill, Queensland, Australia; Drake Coll. 
(USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

♦Malandiola semota Drake 

Malandiola semota Drake 1942b, p. 360 [Victoria; South Australia]. — 
Drake and Ruhoff 1962b, p. 250. 
Type: Holotype o 71 , macropterous; Murray Bridge, South Australia; 
Drake Coll. (USNM). 
Distribution: Australia (South Australia; Victoria; Western Australia). 
Host Plant: Tea tree (myrtaceous shrubs). 

♦Malandiola similis Hacker 

Malandiola similis Hacker 1927, p. 21, pi. 9, fig. 11 [Queensland]. 
Melandiola [sic] similis: Drake and Poor 1937b, p. 402 [New South 
Wales]. 
Type: Sex unknown; Sunnybank, Queensland, Australia; Queensland 

Mus. 
Distribution: Australia (Queensland; Bribie Island; New South Wales). 
Host Plant: Unrecorded. 

Malandiola simplex Horvath 

Malandiola simplex Horvath 1925a, p. 14, fig. 8 [Queensland]. 
Type: Holotype c? ; Malanda, Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 



296 U.S. NATIONAL MUSEUM BULLETIN 243 

*Malandiola syscena Drake and Runoff 

Malandiola syscena Drake and Ruhoff 1962d, p. 788 [Northern Territory]. 

Type: Holotype 9, macropterous; Darwin, Northern Territory, Aus- 
tralia; Drake Coll. (USNM). 

Distribution: Australia (Northern Territory). 

Host Plant: Unrecorded. 

Genus MECOPHARSA Drake 

Mecopharsa Drake 1953b, p. 96. — Drake and Ruhoff 1960a, p. 67. 
Type Species: Mecopharsa hackeri Drake. 

*Mecopharsa hackeri Drake Plate 39 

Macropharsa [sic] hackeri Drake 1953b, p. 97 [Queensland]. — Drake and 
Davis 1960, fig. 48. 
Type: Holotype 9, macropterous; Connondal, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Note: Through typographical errors the generic name Mecopharsa was 
spelled Mecropharsa and Macropharsa (Drake 1953b, p. 97). 

*Mecopharsa secta Drake 

Mecopharsa secta Drake 1960, p. 366, fig. 18 [New Guinea]. 
Type: Holotype 9, macropterous; Eubenangee, New Guinea; Drake 
Coll. (USNM). 
Distribution: New Guinea. 
Host Plant: Unrecorded. 

Genus MEGALOCYSTA Champion 

Megalocysta Champion 1897, p. 5.— Monte 1942c, p. 301. — Hurd 1946, 
p. 474.— Drake and Ruhoff 1960a, p. 67. 
Type Species: Megalocysta pellucida Champion. 

*Megalocysta pellucida Champion 

Megalocysta pellucida Champion 1897, p. 6, pi. 1, figs. 5, 5a, b, c 
[Panama].— Monte 1942c, p. 301. 
Type: Sex unknown; Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

Genus MELANORHOPALA Stal 

Tingis (Melanorhopala) Stal 1873, p. 130. 

Tingis (not Fabricius): Uhler 1886, p. 22.— Banks 1910, p. 57. 

Melanorhopala: Horvath 1908, p. 564.— Van Duzee 1916, p. 26; 1917b, 
pp. 220, 818.— Parshley 1917a, p. 17 (key) .— Blatchley 1926, p. 490 
(key).— Hurd 1946, p. 446.— Drake and Ruhoff 1960a, p. 68. 
Type Species: Tingis (Melanorhopala) clavata Stal. 



LACEBUGS OF THE WORLD 297 

*Melanorhopala balli Drake 

Melanorhopala balli Drake 1928f, p. 3 [Colo.]. 
Type: Holotype 9, brachypterous; Fort Collins, Colorado, U.S.; Drake 
Coll. (USNM). 
Distribution: U.S. (Colo.). 
Host Plant: Unrecorded. 

*Melanorhopala clavata (Stal) 

Tingis (Melanorhopala) clavata Stal 1873, p. 130 [N.Y.; Wis.]. — Parshley 
1914, p. 145 [Maine]. 

Tingis (Melanorhopala) lurida Stal 1873, p. 131 [111.]. 

Tingis (Melanorhopala) uniformis Stal 1873, p. 131. 

Cantacader henshawi Ashmead 1886, p. 20 [Mass.]. — Lethierry and 
Severin 1896, p. 4. 

Lasiacantha clavata: Lethierry and Severin 1896, p. 19. 

Lasiacantha lurida: Lethierry and Severin 1896, p. 19. 

Lasiacantha uniformis: Lethierry and Severin 1896, p. 19. 

Melanorhopala clavata: Horvath 1908, p. 564. — Torre-Bueno 1908, p. 
231; 1910, p. 30.— Smith 1910, p. 149 [N.J.].— Osborn and Drake 
1916a, p. 244, pi. 8, fig. a; pi. 10, fig. d; 1917a, p. 159, pi. 8, figs, a-c 
[Iowa; Nebr.].— Van Duzee 1917b, p. 220 [Man.].— Parshley 1917b, 
p. 56 [ELI.; Conn.]; 1919c, p. 102; 1920a, p. 274; 1923b, p. 706, pi. 
17, fig. 3.— Barber 1922a, p. 17.— Drake 1922d, p. 66; 1926b, p. 376, 
pi. 34, fig. b [Colo.; oak]; 1928b, p. 102; 1930d, p. 269— Blatchley 
1926, p. 491, pi. 4, fig. 3; figs. 116, 117 [Ind.].— Froeschner 1944, 
p. 670 [Mo.].— Proctor 1946, p. 75.— Hurd 1946, p. 446 [Wyo.].— 
Bailey 1951, p. 21 [Fla.; N.H.; Solidago]. 

Tingis clavata: Osborn and Drake 1915a, p. 506 [Ohio]. 

Melanorhopala lurida: Osborn and Drake 1916a, p. 244, pi. 8, fig. b; 
1917a, p. 160, pi. 8, figs, g, h — Van Duzee 1917b, p. 220 [Kans.].— 
Parshley 1917a, p. 19.— Hussey 1922a, p. 11 [N. Dak.].— Drake 
1926b, pi. 34, fig. a. 

Melanorhopala uniformis: Osborn and Drake 1916a, p. 245, pi. 8, fig. c 
[S. Dak.]; 1917a, p. 160, pi. 8, fig. f.— Parshley 1917a, p. 19.— Drake 
1926b, pi. 34, fig. c. 

Melanorhopala obscura Parshley 1916a, p. 167; 1917b, p. 57; 1917c, p. 
47. 

Melanorhopala reflexa Blatchley 1926, p. 492. 
Type: Unknown. 

Distribution: U.S. (Maine; N.H.; Mass.; R.I.; Conn.; N.Y.; N.J.; Fla.; 
Ohio; 111.; Ind.; Kans.; Nebr.; Mo.; Wis.; Iowa; Wyo.; N. Dak.; 
S. Dak; Colo.); Canada (Man.). 
Host Plants: Solidago sp.; oak. 
Note: Discovered in ocean drift [Parshley 1916a], 

681-552—65 20 



298 U.S. NATIONAL MUSEUM BULLETIN 243 

*Melanorhopala infuscata Parshley 

Melanorhopala infuscata Parshley 1917a, p. 19, fig. la [Va.; Liriodendron]; 
1920a, p. 274. — McAtee 1923, p. 145 [D.G.; Ceanothus americanus}. — 
Blatchley 1926, p. 493 [Md.; Ceanothus}.— Bailey 1951, p. 22. 
Type: Holotype cf; Falls Church, Virginia, U.S.; MCZ. 
Distribution: U.S. (Md.; D.C.; Va.). 
Host Plants: Ceanothus americanus; Ceanothus sp.; Liriodendron sp. 

Genus MONOSTEIRA Costa 

Monosteira Costa 1863, p. 7. — Lethierry and Severin 1896, p. 25. — 

Horvath 1906a, pp. 15, 104 (key).— Stichel 1926, pp. 105, 116 (key); 

1960a, p. 343; 1960b, p. 401; 1960c, p. 140.— de Seabra 1931, p. 

438.— Silvestri 1934, p. 259.— Kiritshenko 1951, pp. 243, 254 — 

Drake and Ruhoff 1960a, p. 69. 
Monostira [sic]: Oshanin 1908, p. 454; 1912, p. 46.— Stichel 1935, p. 

349; 1938a, p. 408.— Guide 1938, pp. 245, 319 (key).— Gomez- 

Menor 1949, p. 158; 1950, p. 2; 1954, p. 382; 1955a, p. 257.— Priesner 

and Alfieri 1953, pp. 63, 65. 
Type Species: Monanthia unicornis Mulsant and Rey. 

*Monosteira discoidalis (Jakovlev) 

Monanthia {Monosteira) discoidalis Jakovlev 1883, p. 107 [Russia]. 

Monosteira discoidalis: Lethierry and Severin 1896, p. 25 [Turkmen]. — 
Horvath 1906a, p. 106 [Turkestan]. 
Type: Holotype o 71 , macropterous; Tajikistan, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Turkestan; Turkmen). 
Host Plant: Unrecorded. 

*Monosteira inermis Horvath 

Monosteira inermis Horvath 1899, p. 449 [Turkestan]; 1906a, p. 106. — 
Kiritshenko 1959, p. 105. 
Type: Holotype 9; Turkestan, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 

*Monosteira lobulifera Reuter 

Monosteira lobulifera Reuter 1888, p. 225 [Greece]; 1891a, p. 27. — 
Horvath 1906a, p. 105 [Syria].— Hoberlandt 1955, p. 95 [Turkey; 
Israel; Transcaucasus; Quercus cocci/era; Quercus ilex]. 
Monostira lobulifera: Priesner and Alfieri 1953, p. 66 [Egypt; Salix]. 
Type: Attica, Greece; sex and deposition of type unknown. 
Distribution: Greece; Turkey; Syria; Israel; U.S.S.R. (Transcaucasus); 
Egypt. 
Host Plants: Quercus cocci/era; Qiiercus ilex; Salix sp. 



LACEBUGS OF THE WORLD 299 

*Monosteira minutula Montandon 

Monosteira minutula Montandon 1897, p. 101 [Algeria].— Horvath 1906a, 
p. 106 [Tunisia; Egypt].— Drake 1957a, p. 420 [Aden; Yemen; 
Indigofera] . 
Monostira minutula: Priesner and Alfieri 1953, p. 66 [Qzyphus spina- 
ckristi]. 
Type: Sex unknown; Taguin, Algeria; Bucharest Mus. 
Distribution: Algeria; Tunisia; Egypt; Senegal; Yemen; Aden Protec- 
torate. 
Host Plants: Indigofera sp.; ^izyphus jujuba; ^izyphus spinachristi. 
Note: Taken at light [Priesner and Alfieri]. 

*Monosteira unicostata (Mulsant and Rey) 

Monanthia unicostata Mulsant and Rey 1852a, p. 134 [France]; 1852b, 
p. 153.— Fieber 1861, p. 383.— Walker 1873a, p. 187.— Stal 1874, p. 
58.— Horvath 1874b, p. 432 [Hungary].— Ferrari 1878, p. 66 — 
Bolivar and Chicote 1879, p. 166. 

Monanthia aliena Fieber 1861, p. 124 [Turkey; Syria]. — Walker 1873a, 
p. 185.— Jakovlev 1876b, p. 67.— Ferrari 1874, p. 170; 1878, p. 66. 

Monosteira unicostata: Costa 1863, p. 7, pi. 1, fig. 3. — Garbiglietti 
1869, p. 274 [Italy].— Reuter 1891a, p. 27 [Greece]; 1908, p. 88 — 
Horvath 1892b, p. 131 [Pinus halepensis; Spartium scoparium]; 1906a, 
p. 106 [Yugoslavia; Rumania; Caucasus; Armenia; Turkmen; 
Tunisia]. — Lethierry and Severin 1896, p. 25 [Morocco]. — Stichel 
1926, p. 116; 1960a, p. 344, fig. 181 [Sardinia; Bulgaria; Prunus 
persica; Prunus cerasus]. — de Seabra 1931, pp. 439, 444, figs. 510 (1-4), 
511 [Portugal; Pyrus malus].— Silvestri 1934, p. 259, fig. 222 [Sicily].— 
Hoberlandt 1942, p. 126 [Czechoslovakia; Transcaucasus] ; 1955, p. 
96— Blote 1945, p. 92.— Kiritshenko 1951, p. 254. 

Monanthia (Monosteira) unicostata: Puton 1879c, p. 124 [Algeria; 
Russia; Populus alba]. 

Monostira unicostata: Horvath 1906d, p. 2; 1916, p. 9 [Albania]. — 
Lindberg 1932a, p. 44 [Alnus glutinosa]; 1948, p. 60 [Cyprus]. — 
Vidal 1937, p. 199; 1939, p. 27, fig. 1 [Pyrus communis].— Bremond 
1938, p. 294, pis. 1-3; figs. 1, 2 [Populus nigra; Populus tremula; Pyrus 
mamorensis; Salix; Cydonia vulgaris; Prunus; Crataegus monogyna; Cerasus; 
Amygdalus communis]. — Guide 1938, p. 320, fig. — Gonzalez 1948, p. 
51.— Gomez-Menor 1950, p. 99, figs. 1-5; 1954, p. 383, figs. 5-7 
[Prunus amygdalus]; 1955a, p. 257, figs. 34, 35 (a-e); 1955b, p. 249; 
1956a, pp. Ill, 112, pi. 1, figs. 6, 8.— Novak and Wagner 1951, p. 
71.— Mancini 1953b, p. 186; 1953c, p. 22; 1953d, p. 16.— Seiden- 
stiicker 1954, p. 236. 
Type: Languedoc, France; sex and deposition of type unknown. 
Distribution: Portugal; Spain; France; Italy; Sicily; Hungary; Czecho- 
slovakia; Yugoslavia; Greece; Albania; Sardinia; Bulgaria; Rumania; 



300 U.S. NATIONAL MUSEUM BULLETIN 243 

Turkey; Cyprus; Syria; U.S.S.R. (Armenia; Caucasus; Turkmen; 
Transcaucasus) ; Algeria; Tunisia; Morocco. 

Host Plants: Alnus glutinosa; Amygdalus communis; Cerasus sp.; Crataegus 
monogyna; Cydonia vulgaris; Pinus halepensis; Populus alba; Populus nigra; 
Populus tremula; Populus sp.; Prunus amygdalus; Prunus cerasus; Prunus per ska; 
Prunus sp.; Pyrus communis; Pyrus malus; Pyrus mamorensis; Salix sp.; Spar Hum 
scoparium. 

Note: Life history studies [Vidal 1939; Gomez-Menor 1950; 1955a; 
Bremond 1938]. 

*Monosteira unicostata var. buccata Horvath 
Monosteira buccata Horvath 1902a, p. 600 [Spain]. 
Monosteira unicostata var. buccata: Horvath 1906a, p. 106. — Stichel 

1960a, p. 344. 
Monostira unicostata var. buccata: Lindberg 1932a, p. 44 [Nerium olean- 
der].— Guide 1938, p. 320.— Gomez-Menor 1955b, p. 249 [Persica 
vulgaris]. 
Monostira var. bucata [sic]: Gonzalez 1948, p. 51. 
Type: Sex unknown; Ciudad Real, Spain; Hungarian Mus. 
Distribution: Spain. 
Host Plants: Persica vulgaris; Nerium oleander. 

Genus MUMMIUS Horvath 

Mummius Horvath 1910, p. 65. — Drake and Ruhoff 1960a, p. 69. 
Type Species: Mummius bicorniger Horvath. 

Mummius bicorniger Horvath 

Mummius bicorniger Horvath 1910, p. 65 [Tanganyika]. 

Mummius corniger [sic]: Drake 1954e, p. 5 [Congo]. 
Type: Holotype 9; Kibongoto, Kilimandjaro; Stockholm Mus. 
Distribution: Tanganyika; Congo. 
Host Plant: Unrecorded. 

Mummius denigratus Drake 

Mummius denigratus Drake 1954e, p. 5, fig. 1 [Congo]. 
Type: Holotype 9; Dipidi, Pare National de l'Upemba, Belgian Congo; 
Brussels Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Genus NAITINGIS Drake and Ruhoff 

Naitingis Drake and Ruhoff 1962c, p. 133. 
Type Species: Tropidocheila maynei Schouteden. 



LACEBUGS OF THE WORLD 301 

Naitingis blukwana (Drake) 

Tingis {Tropidocheila) blukwana Drake 1954e, p. 2 [Congo]. 
Naitingis blukwana: Drake and Runoff 1962c, p. 134. 
Type: Holotypecf ; Blukwa, Belgian Congo; Brussels Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Naitingis maynei (Schouteden) 

Tropidocheila maynei Schouteden 1923, p. 105 [Tanganyika]. 

Tingis {Tropidocheila) maynei: Drake 1954e, p. 2 [Congo]. 

Tingis (Tropidochila) maynei: Drake 1958a, p. 104 [Angola]. 

Naitingis maynei: Drake and Ruhoff 1962c, p. 134. 
Type: Sex unknown; Albertville, Tanganyika; Cent. Afr. Mus. 
Distribution: Tanganyika; Congo; Angola. 
Host Plant: Unrecorded. 

Naitingis maynei var. biseriata (Schouteden) 

Tropidocheila maynei var. biseriata Schouteden 1923, p. 106 [Tanganyika], 
Naitingis maynei var. biseriata: Drake and Ruhoff 1962c, p. 134. 
Type: Sex unknown; Albertville, Tanganyika; Cent. Afr. Mus. 
Distribution: Tanganyika. 
Host Plant: Unrecorded. 

Naitingis nyanzae (Schouteden) 

Tropidocheila nyanzae Schouteden 1955a, p. 30 [Congo], 
Naitingis nyanzae: Drake and Ruhoff 1962c, p. 134. 
Type: Sex unknown; Ukewere, Victoria-Nyanza, Belgian Congo; Cent. 
Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Genus NAOCHILA Drake 

Naochila Drake 1957f, p. 127.— Drake and Ruhoff 1960a, p. 70. 
Type Species: Cochlochila boxiana Drake. 

*Naochila arete Drake and Mohanasundarum 

Naochila arete Drake and Mohanasundarum 1961, p. 108, fig. 1 [India; 
Cordia] . 
Type: Holotype cf, brachypterous; Coimbatore, India; Drake Coll. 
(USNM). 
Distribution: India. 
Host Plant: Cordia sp. 

*Naochila boxiana (Drake) 

Cochlochila boxiana Drake 1953d, p. 214 [Ghana; Hoslundia oppositifolia; 

Uganda]; 1955b, p. 85 [Angola]. 
Naochila boxiana: Drake 1957f, p. 128. 



302 U.S. NATIONAL MUSEUM BULLETIN 2 43 

Type: Holotype cf; Tafo, Gold Coast; British Mus. 
Distribution: Ghana; Uganda; Angola. 
Host Plant: Hoslundia oppositijolia. 

*Naochila bukobana (Drake) 

Cochlochila bukobana Drake 1953d, p. 214 [Tanganyika]. 
Naochila bukobana: Drake 1957f, p. 128. 
Type: Holotype d", macropterous; Lake Victoria, Tanganyika; Drake 
Coll. (USNM). 
Distribution: Tanganyika. 
Host Plant: Unrecorded. 

*Naochila bullana (Drake) 

Cysteochila bullana Drake 1958c, p. 319 [Malagasy]. 
Naochila bullana: Drake and Ruhoff 1961c, p. 145. 
Type: Holotype d 71 ; Ankaratra, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

*Naochila kivuensis (Schouteden) 

Monanthia kivuensis Schouteden l953e, p. 199 [Congo; Acanthus; Ficus]; 

1957c, p. 318 [Ruanda]. 
Naochila kivuensis: Drake 1957f, p. 128. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Ruanda-Urundi. 
Host Plants: Acanthus sp.; Ficus sp. 

*Naochila natalana (Drake) 

Cochlochila natalana Drake 1954b, p. 8 [Natal; Pondoland]. 

Naochila natalana: Drake 1957f, p. 128. 
Type: Holotype d* ; Port Shepstone, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal; Pondoland); Kenya. 
Host Plant: Unrecorded. 

*Naochila parvella (Drake) 

Cochlochila parvella Drake 1954b, p. 7 [Natal; Pondoland; Cape Prov- 
ince]; 1958c, p. 320 [Malagasy]. 
Naochila parallela [sic]: Drake 1957f, p. 128. 
Type: Holotype o 71 ; Weenan, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal; Pondoland; Cape Province); Mala- 
gasy Republic. 
Host Plant: Unrecorded. 

*Naochila sufflata (Drake and Poor) 

Monanthia sufflata Drake and Poor 1939c, p. 203 [India; Lantana], 
Dictyla sufflata: Drake and Ruhoff 1960a, p. 51. 
Naochila sufflata: Drake and Ruhoff 1960c, p. 29. 



LACEBUGS OF THE WORLD 303 

Type: Holotype 9, macropterous; Dharampur, Dehra Dun, India; Drake 
Coll. (USNM). 
Distribution: India. 
Host Plants: Ehretia laevis; Lantana sp. 

*Naochila turgida Drake 

Naochila turgida Drake 1957f, p. 128 [Malagasy]. 
Type: Holotype o 71 ; Ankaratra, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

Genus NEOTINGIS Drake 

Neotingis Drake 1922c. p. 366.— Drake and Runoff 1960a, p. 70. 
Type Species: Neotingis hollandi Drake. 

*Neotingis hollandi Drake 

Neotingis hollandi Drake 1922c, p. 367, pi. 39, fig. 5 [Brazil]. — Monte 
1941e, p. 126. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

Genus NESOCYPSELAS Kirkaldy 

Nesocypselas Kirkaldy 1908b, p. 364.— Drake and Runoff 1960a, p. 70. 
Type Species: Nesocypselas dicysta Kirkaldy. 

Nesocypselas bellatula Drake 

Nesocypselas bellatula Drake 1960, p. 375, fig. 25 [New Guinea]. 
Type: Holotype 9; Wesselmeren, Duroto, E. of Enarotadi, Netherlands 
New Guinea; Bishop Mus. 
Distribution: New Guinea (Netherlands). 
Host Plant: Unrecorded. 

*Nesocypselas dicysta Kirkaldy 

Nesocypselas dicysta Kirkaldy 1908b, p. 365, pi. 4, figs. 8, 9 [Viti Levu; 
Artocarpus incisa]. — Drake and Poor 1943, p. 200. 
Type: Holotype o 71 ; Viti Levu, Fiji Islands; Hawaii. Sugar Plant. Assn. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Artocarpus incisa. 

*Nesocypselas evansi Drake 

Nesocypselas evansi Drake 1953b, p. 98 [Taveuni]. 
Type: Holotype o 71 ; Waiyero, Taveuni, Fiji Islands; British Mus. 
Distribution: Fiji Islands (Taveuni). 
Host Plant: Unrecorded. 



304 U.S. NATIONAL MUSEUM BULLETIN 243 

^Nesocypselas evansi subsp. aemulus Drake 

Nesocypselas evansi subsp. aemulus Drake 1953b, p. 99 [Fiji]. 
Type: Holotype 9; Bau, Fiji Islands; British Mus. 
Distribution: Fiji Islands. 
Host Plant: Unrecorded. 
^Nesocypselas inannana Drake 

Nesocypselas inannana Drake 1953d, p. 222 [Espiritu Santo]. 
Type: Holotype 9, macropterous; Espiritu Santo, New Hebrides; USNM. 
Distribution: New Hebrides Islands (Espiritu Santo). 
Host Plant: Unrecorded. 
Nesocypselas muiri Drake and Poor (emendation) 

Nesocypselas muri Drake and Poor 1943, p. 203, fig. 7 [Viti Levu]. 
Type: Holotype 9; Nandarivatu, Viti Levu, Fiji Islands; Bishop Mus. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Unrecorded. 
*Nesocypselas piperica Drake 

Nesocypselas piperica Drake 1957d, p. 203, figs. 1, 2 [New Britain; 
pepper]; 1960, p. 374, fig. 24 [Papua].— Drake and Ruhoff 1960a, 
p. 5, fig. 1. 
Type: Holotype c? ; Keravat, New Britain, Melanesia; British Mus. 
Distribution: New Britain Island, Bismarck Archipelago; New Guinea 
(Papua). 

Host Plant: Cultivated pepper. 
Nesocypselas simulis Drake and Poor 

Nesocypselas simulis Drake and Poor 1943, p. 203, fig. 6 [Viti Levu]. 
Type: Holotype 9; Nandarivatu, Viti Levu, Fiji Islands; Bishop Mus. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Unrecorded. 
^Nesocypselas vicinalis Drake and Poor (emendation) 

Nesocypselas vicinatis Drake and Poor 1943, p. 201 [Viti Levu]. 
Type: Holotype 9; Nandarivatu, Viti Levu, Fiji Islands; Bishop Mus. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Unrecorded. 

Genus NESOCYSTA Kirkaldy 

Nesocysta Kirkaldy 1908b, p. 365.— Drake and Ruhoff 1960a, p. 70. 
Type Species: Nesocysta rugata Kirkaldy. 
*Nesocysta rugata Kirkaldy 

Nesocysta rugata Kirkaldy 1908b, p. 366 [Viti Levu]. — Drake and Poor 
1943, p. 204. 
Type: Holotype 9; Rewa, Fiji Islands; Hawaii. Sugar Plant. Assn. 
Distribution: Fiji Islands (Viti Levu). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 305 

Genus NESOTINGIS Drake 

Nesotingis Drake 1957e, p. 402.— Drake and Runoff 1960a, p. 70. 
Type Species: Nesotingis pauliani Drake. 

Nesotingis pauliani Drake 

Nesotingis pauliani Drake 1957e, p. 403 [Reunion]. 
Type: Holotype 9; Belouve, Reunion; Madagascar Sci. Inst. 
Distribution : Mascarene Islands (Reunion). 
Host Plant: Unrecorded. 

*Nesotingis vinsoni (Drake and Mamet) 

Eteoneus vinsoni Drake and Mamet 1956, p. 301, fig. 6 [Mauritius]. — 

Mamet 1957, p. 57. 
Nesotingis vinsoni: Drake 1957e, p. 404 [Reunion]. 
Type: Holotype 9, macropterous; Les Mares, Mauritius Island; Drake 
Coll. (USNM). 
Distribution: Mascarene Islands (Mauritius; Reunion). 
Host Plant: Unrecorded. 

Genus NETHERSIA Horvath 

Nethersia Horvath 1925a, p. 14.— Drake and Ruhoff 1960a, p. 71. 
Type Species: Nethersia maculosa Horvath. 

*Nethersia absimilis Drake 

Nethersia absimilis Drake 1944a, p. 74 [Queensland]. 
Type: Holotype 9, macropterous; Roma, Queensland, Australia; Drake 
Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Nethersia haplotes Drake and Ruhoff 

Nethersia haplotes Drake and Ruhoff 1962d, p. 505 [New South Wales]. 

Type: Holotype cf , macropterous; Mullahey, New South Wales, Aus- 
tralia; Drake Coll. (USNM). 

Distribution: Australia (New South Wales). 

Host Plant: Unrecorded. 

*Nethersia maculosa Horvath 

Nethersia maculosa Horvath 1925a, p. 15, fig. 9 [Western Australia]. — 
Hacker 1928, p. 186 [South Australia; Queensland]. — Drake and 
Ruhoff 1962a, p. 156; 1962b, p. 250; 1962d, p. 505. 
Nethersia pugna Drake 1944a, p. 76. 
Nathersea [sic] maculosa: Drake 1947c, p. 111. 
Type: Holotype 9; Broome, Australis occidentalis; Stockholm Mus. 
Distribution: Australia (Queensland; Western Australia; Northern Ter- 
ritory; South Australia). 
Host Plant: Unrecorded. 



306 U.S. NATIONAL MUSEUM BULLETIN 243 

*Nethersia nigritarsis (Horvath) 

Tingis (Tropidochila) nigritarsis Horvath 1925a, p. 7 [Queensland]. 
N ether sia poor ae Drake 1944a, p. 72, fig. 1. 
Nether sia nigritarsis: Drake and RuhofF 1962d, p. 505. 
Type: Holotype 9; Herberton, Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Nethersia setosa (Hacker) 

Ischnotingis setosus Hacker 1927, p. 23, pi. 6, fig. 3 [Queensland; Acacia]. 
JV r ether sia setosa: Hacker 1928, p. 186. 
Type: Sex unknown; Mount Coot-tha, Queensland, Australia; Queens- 
land Mus. 
Distribution: Australia (Queensland; New South Wales). 
Host Plant: Acacia sp. 

Genus NOBARNUS Distant 

Nobarnus Distant 1920, p. 156.— Drake and RuhofF 1960a, p. 71. 
Type Species: Nobarnus typicus Distant. 

Nobarnus signatus (Distant) Plate 41 

Compseuta signata Distant 1920, p. 156 [New Caledonia]. 
Nobarnus signatus: China 1926, p. 228. — Drake and Davis 1960, fig. 38. 

Type: Sex unknown; Paompai, New Caledonia; British Mus. 

Distribution: New Caledonia. 

Host Plant: Unrecorded. 

Nobarnus typicus Distant 

Nobarnus typicus Distant 1920, p. 157 [New Caledonia]. 
Type: Sex unknown; Mt. Arago, New Caledonia; British Mus. 
Distribution: New Caledonia. 
Host Plant: Unrecorded. 

Genus NYCTOTINGIS Drake 

Nyctotingis Drake 1922c, p. 362.— Drake and RuhofF 1960a, p. 71. 
Type Species: Nyctotingis osborni Drake. 

*Nyctotingis nexilis Drake 

Nyctotingis nexilis Drake 1948J, p. 431 [Peru]. 
Type: Holotype 9, macropterous; Tingo Maria, Peru; Drake Coll. 
(USNM). 
Distribution: Peru; Bolivia. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 307 

*Nyctotingis osborni Drake 

Nyctotingis osborni Drake 1922c, p. 363, fig. 1 [Brazil]; 1928a, p. 42 
[Ecuador].— Monte 1938b, p. 132; 1941e, p. 126 [Bambusa].— Drake 
and Hambleton 1945, p. 357 [Peru].— Silva 1956, p. 46, fig. 17. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil; Ecuador; Peru. 
Host Plant: Bambusa sp. 

Genus OCTACYSTA Drake and Runoff 

Octacysta Drake and Ruhoff 1960a, p. 71.— Stichel 1960b, p. 401; 
1960c, p. 140. 
Type Species: Tingis rotundata Herrich-Schaeffer = Tingis echii Fabricius 
(not Schrank). 
*Octacysta echii (Fabricius) 

Tingis echii Fabricius 1803, p. 126 [Austria; Echium]. — Burmeister 
1835, p. 259 [Echium vulgare].—Amyot and Serville 1843, p. 297 
[France]. 

Tingis rotundata Herrich-Schaeffer 1835, p. 59. 

Monanthia rotundata: Herrich-Schaeffer 1838, pp. 52, 59, pi. 124, figs. 
392, f-g; pi. 125, fig. 2 [Czechoslovakia].— Horvath 1906a, pp. 99, 
103 [Hungary; Rumania; Russia; Caucasus]. — Muzik 1907, p. 64. — 
Oshanin 1908, p. 454 [Turkestan]; 1912, p. 46.— Moroder Sala 
1920, p. 13 [Spain].— Stichel 1926, p. 115; 1935, p. 349; 1938a, p. 
408— Borner 1935, p. 78.— Guide 1938, p. 318.— Hoberlandt 1942, 
p. 126; 1943a, p. 119; 1943b, p. 124; 1955, p. 95 [Transcaucasus].— 
Blote 1945, p. 9 1 .— Kiritshenko 1951, p. 254.— Seidenstiicker 1954, 
p. 236 [Turkey]. — Stusak and Stys 1959, p. 199, figs. 41-46 [Anchusa 
officinalis]. — Putshkov 1960, p. 305 [Cynoglossum]. — Stusak 1961a, p. 
86, figs. 5c, 14, 15; pi. 2. 

Monanthia (Physatocheila) echii: Fieber 1844, p. 88, pi. 7, figs. 27-32 
[Italy].— Herrich-Schaeffer 1850, p. 153. 

Tingis (Physatocheila) echii: Kolenati 1856, p. 428 [Echium rubrum; 
Echium italicum; Armenia]. 

Monanthia echii: Fieber 1861, p. 126. — Garbiglietti 1869, p. 273. — 
Walker 1873a, p. 187— Stal 1874, p. 59.— Puton 1879c, p. 123 — 
Hiieber 1893, p. 355 [Germany; Switzerland]. 

Octacysta rotundata: Drake and Ruhoff 1960a, p. 71, pis. 6, 7. — Stichel 
1960b, p. 401; 1960c, p. 140. 

Dictyla rotundata: Stichel 1960a, p. 342, fig. 180a [Poland; Bulgaria]. 

Octacysta echii: Drake and Ruhoff 1960b, p. 74, fig. 22. 
Type: Lectotype 9, brachypterous; Austria; Copenhagen Mus. 
Distribution: Spain; France; Germany; Switzerland; Czechoslovakia; 
Austria; Italy; Hungary; Rumania; Bulgaria; Poland; U.S.S.R. (Armenia; 
Caucasus; Transcaucasus; Turkestan); Turkey. 



308 U.S. NATIONAL MUSEUM BULLETIN 243 

Host Plants: Anchusa officinalis; Cynoglossum sp.; Echium italicum; Echium 
rubrum; Echium vulgare; Echium sp. 

Note: Study of nymphs and eggs [Stusak and Stys; Stusak]. Resurrec- 
tion of name [Drake and Ruhoff 1960b]. 

Genus OEDOTINGIS Drake 

Oedotingis Drake 1942a, p. 19.— Drake and Ruhoff 1960a, p. 72. 
Type Species: Australotingis williamsi Drake. 

*Oedotingis mexicana Drake 

Odeotingis [sic] mexicana Drake 1948d, p. 17 [Mexico]. 

Type: Holotype 9, macropterous; Guerrero, Mexico; USNM. 

Distribution: Mexico. 

Host Plant: Unrecorded. 

Note: Intercepted on orchid plants at port-of-entry, Laredo, Texas. 
The sex of the type species was erroneously reported as male in the original 
description. 

*Oedotingis williamsi (Drake) 

Australotingis williamsi Drake 1928a, p. 51 [Ecuador]. 

Oedotingis williamsi: Drake 1942a, p. 20. 
Type: Holotype 9, macropterous; Mera, Ecuador; Drake Coll. (USNM). 
Distribution: Ecuador. 
Host Plant: Unrecorded. 

Genus OGYGOTINGIS Drake 

Ogygotingis Drake 1948e, p. 149.— Drake and Ruhoff 1960a, p. 72. 
Type Species: Teleonemia insularis China. 

*Ogygotingis insularis (China) 

Teleonemia insularis China 1924, p. 436, fig. 2b [Rodriguez]. 

Ogygotingis insularis: Drake 1948e, p. 149. 

Orygotingis [sic] insularis: Drake and Davis 1960, fig. 41. 
Type: Sex unknown; Rodriguez Island; British Mus. 
Distribution: Mascarene Islands (Rodriguez). 
Host Plant: Unrecorded. 

Genus OLASTRIDA Schouteden 

Olastrida Schouteden 1956, p. 205.— Drake and Ruhoff 1960a, p. 72. 
Type Species: Olastrida oleae Schouteden. 

*01astrida oleae Schouteden 

Olastrida oleae Schouteden 1956, p. 205 [Ruanda; Olea europea]. 
Type: Sex unknown; Astrida, Ruanda; Cent. Afr. Mus. 
Distribution : Ruanda-Urundi. 
Host Plant: Olea europea. 



LACEBUGS OF THE WORLD 309 

Genus ONCOCHILA Stal 

Oncochila Stal 1873, p. 121; 1874, p. 57.— Kirkaldy 1904, p. 281.— 
Horvath 1906a, pp. 15, 96 (key).— Oshanin 1908, p. 449; 1912, p. 
45— Stichel 1926, pp. 105, 114 (key); 1935, p. 349; 1938a, p. 408; 
1960a, p. 336; 1960b, p. 400; 1960c, p. 139.— Borner 1935, pp. 74, 
78— Guide 1938, pp. 245, 309 (key).— China 1943, p. 248 — 
Kiritshenko 1951, pp. 242, 252.— Drake and Ruhoff 1960a, p. 72. 
Type Species: Monanthia (Physatocheila) scapular is Fieber. 
*Oncochila scapularis (Fieber) 

Monanthia {Physatocheila) scapularis Fieber 1844, p. 80, pi. 6, figs. 38-40 
[Czechoslovakia; Austria; Italy; Germany; Senecio jacobaea]. 

Monanthia scapularis: Scholz 1847, p. 120. — Mayr 1858, p. 571 [Hun- 
gary] .—Fieber 1861, p. 125.— Garbiglietti 1869, p. 273.— Walker 
1873a, p. 186.— Reiber and Puton 1876, p. 69.— Horvath 1889, p. 
326. — Hiieber 1893, p. 349 [Yugoslavia; Rumania]. 

Oncochila scapularis: Stal 1873, p. 121; 1874, p. 57. — Horvath 1906a, 
p. 97 [Russia; Siberia; Euphorbia gerardiana]. — Muzik 1907, p. 62. — 
Stichel 1926, p. 115; 1938a, p. 408 [Albania; Poland]; 1960a, p. 
336 [Bulgaria].— Hoberlandt 1942, p. 126; 1943b, pp. 119, 124.— 
Stusak 1958, p. 367, figs. 12-14. 

Physatochila scapularis: Lethierry and Sever in 1896, p. 21. 
Type: Unknown. 

Distribution: Czechoslovakia; Hungary; Austria; Italy; Germany; 
Rumania; Yugoslavia; Albania; Poland; Bulgaria; U.S.S.R. (Siberia). 
Host Plants: Euphorbia gerardiana; Senecio jacobaea. 
Note: Eggs [Stusak]. 
*Oncochila simplex (Herrich-Schaeffer) 

Tingis simplex Herrich-Schaeffer 1830, heft 118, tab. 21 [Germany]; 
1835, p. 59. 

Monanthia simplex: Herrich-Schaeffer 1838, p. 59. — Douglas and Scott 
1865, p. 245 [England].— Horvath 1874b, p. 432 [Hungary].— 
Jakovlev 1876b, p. 66 [Russia]; 1880b, p. 108 [Caucasus]. — Saunders 
1875, p. 250. — Puton 1879c, p. 119 [France; Euphorbia cyparissias]. — 
Chicote 1880, p. 189 [Spain].— Dubois 1888, p. 121.— d'Antessanty 
1890, p. 33.— Hiieber 1893, p. 347 [Austria; Switzerland].— Butler 
1923, p. 214 [Turkestan]. 

Monanthia (Physatocheila) simplex: Herrich-Schaeffer 1850, p. 153. — 
Saunders 1892, p. 136, pi. 12, fig. 10. 

Physatochila (Oncochila) simplex: Sahlberg 1878, p. 21. 

Physatochila simplex: Lethierry and Severin 1896, p. 21. — Reichensperger 
1920, p. 62. 



310 U.S. NATIONAL MUSEUM BULLETIN 243 

Oncochila simplex: Horvath 1906a, pp. 96, 97 [Belgium; Italy; Yugo- 
slavia; Rumania; Armenia]. — Muzik 1907, p. 62 [Czechoslovakia]. — 
Stichel 1926, p. 115; 1960a, p. 336 [Poland; Bulgaria; Thymus].— 
Lindberg 1927, p. 20.— Scholte 1935, p. 90, fig. 23 [Netherlands].— 
Guide 1938, p. 310, fig. [Denmark].— Hoberlandt 1942, p. 126; 
1943b, p. 124.— Blote 1945, p. 89 [Tunisia].— Bator 1953, p. 326, 
pi. 4, fig. 1. — Cobben 1958a, p. 12, fig. 15 [Euphorbia gerardiana]. — 
Southwood and Leston 1959, p. 151, pi. 19, fig. 3; pi. 21, fig. 6. 
Type: Regensburg, Germany; sex and deposition of type unknown. 
Distribution: Germany; France; England; Italy; Austria; Poland; 
Switzerland; Yugoslavia; Hungary; Belgium; Rumania; Bulgaria; Czecho- 
slovakia; Netherlands; Spain; Denmark; Albania; U.S.S.R. (Armenia; 
Turkestan; Caucasus); Tunisia. 
Host Plants: Euphorbia cyparissias; Euphorbia gerardiana; Thymus sp. 

fOncochila wollastoni (Heer) 

Tingis wollastoni Heer 1865, fig. 307; 1872, fig. 307; 1876, fig. 307; 

1879, fig. 307.— Scudder 1891, p. 449. 
Monanthia wollastoni: Heer 1865, p. 392 [Baden]; 1872, p. 480; 1876, 

p. 50; 1879, p. 417.— Scudder 1891, p. 422 [Horizon: Tortonian]. 
Dictyla wollastoni: Drake and Ruhoff 1960a, pp. 11, 51. 
Oncochila wollastoni: Drake and Ruhoff 1960c, p. 32. 
Type: Sex undeterminable; fossil; Oeningen, Baden; deposition unknown. 
Distribution: Fossil. 

Note: In the above publications by Heer, wollastoni was placed in the 
genus Tingis in figure captions (which preceded the text) and in Monanthia 
in the description. 

Genus ONCOPHYSA Stal 

Oncophysa Stal 1873, pp. 121, 129.— Drake and Ruhoff 1960a, p. 73. 
Type Species: Monanthia (Physatocheila) vesiculata Stal. 

*Oncophysa leai Drake 

Oncophysa leai Drake 1948e, p. 154 [Western Australia], 
Type: Holotype 9; Australia; Hungarian Mus. 
Distribution: Australia (Western Australia). 
Host Plant: Unrecorded. 

^Oncophysa rufescens Hacker 

Oncophysa rufescens Hacker 1928, p. 178, pi. 21, fig. 8 [South Australia]. 

Type: Sex unknown; Port Noarlunga, South Australia, Australia; 
Queensland Mus. 

Distribution: Australia (South Australia). 

Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 31 1 

*Oncophysa vesiculata (Stal) 

Monanthia (Physatocheila) vesiculata Stal 1859, p. 259 [New South Wales]. 
Monanthia vesiculata: Walker 1873a, p. 197. 

Oncophysa vesiculata: Stal 1873, p. 129. — Lethierry and Severin 1896, 
p. 22— Horvath 1925a, p. 2 [Queensland] .—Hacker 1927, p. 25 
[Pimelea]; 1928, p. 177 [Victoria].— Drake 1961b, p. 108. 
Type: Holotype 9; New Holland, Sydney, Australia; Stockholm Mus. 
Distribution: Australia (New South Wales; Queensland; Victoria). 
Host Plant: Pimelea sp. 

Oncophysa vesiculata var. gracilis Hacker 

Oncophysa vesiculata var. gracilis Hacker 1928, p. 178, pi. 21, fig. 7 [South 
Australia].— Drake 1961b, p. 108 [New South Wales]. 
Type: Sex unknown; Cape Jarvis, South Australia, Australia; Queens- 
land Mus. 
Distribution: Australia (South Australia; New South Wales). 
Host Plant: Unrecorded. 

*Oncophysa vesiculata var. nigra Hacker 

Oncophysa vesiculata var. nigra Hacker 1928, p. 178, pi. 21, fig. 6 [Tas- 
mania; Victoria].— Drake 1961b, p. 108 [New South Wales].— 
Drake and Runoff 1962b, p. 250. 
Type: Sex unknown; Mount Arthur, Tasmania, Australia; Queensland 
Mus. 
Distribution: Australia (Tasmania; New South Wales; Victoria). 
Host Plant: Unrecorded. 

Genus ONYMOCHILA Drake 

Onymochila Drake 1948e, p. 152.— Drake and Ruhoff 1960a, p. 73. 
Type Species: Cysteochila dichapetali Horvath. 

*Onymochila dichapetali (Horvath) (emendation) Plate 17 

Cysteochila dichopetali Horvath 1929, p. 324 [Transvaal; Dichapetalum 

cymosum]. 
Onymochila dichopalali [sic]: Drake 1948e, p. 152. 
Onymochila dichopetali: Drake and Davis 1960, figs. 50, 51. 
Type: Sex unknown; Pretoria, Transvaal, South Africa; British Mus. 
Distribution: South Africa (Transvaal). 

Host Plant: Causes extensive down -curl of the leaves of Dichapetalum 
cymosum (see pi. 17). 

Genus OROTINGIS Drake and Poor 

Orotingis Drake and Poor 1941, p. 161. — Drake and Ruhoff 1960a, 
p. 73. 
Type Species: Orotingis muiri Drake and Poor. 



312 U.S. NATIONAL MUSEUM BULLETIN 243 

Orotingis eueides Drake 

Orotingis eueides Drake 1960, p. 364, fig. 16 [New Guinea]. 
Type: Holotype o"; Wisselmeren, Duroto, Netherlands New Guinea; 
Bishop Mus. 
Distribution: New Guinea (Netherlands). 
Host Plant: Unrecorded. 

*Orotingis muiri Drake and Poor 

Orotingis muiri Drake and Poor 1941, p. 161 [Amboina]. — Drake and 
Davis 1960, fig. 64. 
Type: Holotype cf ; Amboina Island; Cal. Acad. 
Distribution: Moluccas (Amboina Island). 
Host Plant: Unrecorded. 

Genus OTTOICUS Drake 

Ottoicus Drake 1960, p. 358. 
Type Species: Ottoicus dissitus Drake. 

*Ottoicus dissitus Drake 

Ottoicus dissitus Drake 1960, p. 358, fig. 12 [New Guinea]. 
Type: Holotype cf ; Mt. Otto, above Kabebe, New Guinea; Bishop Mus. 
Distribution: New Guinea. 
Host Plant: Unrecorded. 

Genus PACHYCYSTA Champion 

Pachycysta Champion 1898b, p. 59.— Drake 1928g, pp. 184-185 (key).— 
Monte 194le, p. 126.— Hurd 1946, p. 474.— Drake and Ruhoff 
1960a, p. 74. 
Type Species: Pachycysta diaphana Champion. 

*Pachycysta adolpha Drake 

Pachycysta adolpha Drake 1948j, p. 431 [Peru]. 
Type: Holotype 9, macropterous; Maracapata, Peru; Drake Coll. 
(USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 

*Pachycysta championi Drake 

Pachycysta championi Drake 1921a, p. 344, figs, a, b [Bolivia]; 1928g, p. 
184.— Drake and Davis 1960, fig. 57. 
Type: Sex unknown; Cochabamba, Bolivia; Paris Mus. 
Distribution: Boliva. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 313 

*Pachycysta diaphana Champion 

Pachycysta diaphana Champion 1898b, p. 59, pi. 2, figs. 6, 6a [Brazil]. — 
Drake 1928g, p. 184.— Monte 1940e, p. 300; 1942a, p. 97.— Drake 
and Hambleton 1944b, p. 124 [Venezuela]. 
Type: Holotype 9; Amazons; Oxford Mus. 
Distribution: Brazil; Venezuela. 
Host Plant: Unrecorded. 

*Pachycysta hambletoni Drake and Poor 

Pachycysta hambletoni Drake and Poor 1938a, p. 31 [Brazil]. — Monte 
1941e, p. 127 [Guettarda viburnoides]. 
Type: Holotype cf, macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Guettarda viburnoides. 

*Pachycysta schildi Drake 

Pachycysta schildi Drake 1928g, p. 185 [Costa Rica]. 

Pachycysta shildi [sic]: Monte 1942a, p. 96 [Venezuela]. 
Type: Holotype o 71 , macropterous; Suize Tur'lba, Costa Rica; Drake Coll. 
(USNM). 
Distribution: Costa Rica; Venezuela. 
Host Plant: Unrecorded. 

Genus PALAUELLA Drake 

Palauella Drake 1956d, pp. 105, 110.— Drake and Runoff 1960a, p. 74. 
Type Species: Palauella gressitti Drake. 

*Palauella gressitti Drake 

Palauella gressitti Drake 1956d, pp. 105, 112, fig. 6 [Palau]. 

Type: Holotype 9, macropterous; Ulimang, Babelthuap, Palau Islands; 
USNM. 

Distribution: Palau Islands. 

Host Plant: Unrecorded. 

Genus PARACOPIUM Distant 

Paracopium Distant 1902b, p. 354; 1903b, p. 128— Horvath 1929, p. 
322.— Drake and Ruhoff 1960a, p. 74. 
Type Species: Dictyonota cingalensis Walker. 

*Paracopium africum Drake 

Paracopium africum Drake 1956g, p. 472 [Natal]; 1961c, p. 129. 
Type: Holotype o", macropterous; Natal, South Africa; Drake Coll. 
(USNM). 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

681-552—64 21 



314 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Paracopium albofasciatum Hacker 

Paracopium albqfasciata Hacker 1927, p. 21, pi. 6, fig. 2 [Queensland]. — 
Drake 1961b, p. 110. 
Type: Sex unknown; Prairie, Queensland, Australia; Queensland Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Paracopium antennatum (Schouteden) 

Copium antennatum Schouteden 1923, p. 86 [Congo]. 
Paracopium antennatum: Drake 1954f, p. 4. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Cameroun. 
Host Plant: Unrecorded. 

*Paracopium assimile Horvath 

Paracopium assimile Horvath 1929, pp. 321, 322 [Uganda]. 
Type: Holotype 9; Nagunga, Uganda; British Mus. 
Distribution: Uganda; Nigeria; Tanganyika. 
Host Plant: Unrecorded. 

*Paracopium australicum (Stal) 

Catoplatus austr aliens Stal 1873, p. 128 [Queensland]. — Lethierry and 

Severin 1896, p. 20. 
Paracopium austr aliens: Hacker 1927, p. 20, pi. 6, fig. 1; 1928, p. 183 

[New South Wales; Melville; Moa]. 
Paracopium australicum: Horvath 1929, p. 322. — Drake 1961b, p. 110. — 
Drake and Ruhoff 1962b, p. 250. 
Type: Holotype 9; Port Denison, Cape York, Australia; Stockholm Mus. 
Distribution: Australia (Queensland; New South Wales; Moa and 
Melville Islands). 

Host Plant: Unrecorded. 

*Paracopium bequaerti (Schouteden) 

Copium bequaerti Schouteden 1923, p. 86 [Congo]. 

Copium {Paracopium) bequaerti: Mayne and Ghesquiere 1934, p. 20 

[Clerodendron giletii] . 
Paracopium bequaerti: Drake 1954f, p. 4; 1958b, p. 29. 
Type: Sex unknown; Sankisia, Katanga, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Clerodendron giletii. 

Paracopium burgeoni (Schouteden) 

Catoplatus burgeoni Schouteden 1923, p. 89 [Congo]. 

Paracopium burgeoni: Drake 1954f, p. 4. 
Type: Sex unknown; Kindu, Manyema, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 315 

*Paracopium caledonicum Drake 

Paracopium caledonicum Drake 1956g, p. 474 [New Caledonia]. 
Type: Holotype o 71 , macropterous; New Caledonia; Drake Coll. (USNM). 
Distribution: New Caledonia. 
Host Plant: Unrecorded. 
*Paracopium cingalense (Walker) 

Dictyonota cingalensis Walker 1873a, p. 178 [Ceylon]. — Kirby 1891, 

p. 109. — Lethierry and Severin 1896, p. 25. 
Paracopium cingalensis: Distant 1902b, p. 354. 

Paracopium cingalense: Distant 1903b, p. 128, fig. 92. — Maxwell-Lefroy 
1909, p. 693 [India; Clerodendron phlomoides]. — Horvath 1929, p. 322. 
Type: Sex unknown; Ceylon; British Mus. 
Distribution: Ceylon; India. 
Host Plant: Clerodendron phlomoides. 
Paracopium coloratura Schouteden 

Paracopium coloratum Schouteden 1955b, p. 164 [Congo]. 
Type: Sex unknown; Lukenie, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 
*Paracopium comatum Drake (emendation) 

Paracopium comantis Drake 1953d, p. 217 [India]. 
Type: Holotype cf ; West Almora, United Province, India; British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 
*Paracopium congruum Drake 

Paracopium congruum Drake 1956g, p. 471, fig. 1 [Congo; Rhodesia]. 
Type: Holotype cf, macropterous; Victoria Falls, Rhodesia; Drake Coll. 
(USNM). 
Distribution: Rhodesia; Congo. 
Host Plant: Unrecorded. 
*Paracopium dauphinicum Drake 

Paracopium dauphinicum Drake 1956g, p. 473, fig. 2 [Malagasy]. 
Type: Holotype 9, macropterous; Fort Dauphin, Madagascar; Drake 
Coll. (USNM). 
Distribution : Malagasy Republic. 
Host Plant: Clerodendron micans. 
*Paracopium fenestellatum (Bergroth) 

Copium fenestellatum Bergroth 1894, p. 168 [Malagasy]. — Lethierry and 

Severin 1896, p. 16. 
Paracopium fenestellatum: Horvath 1929, p. 322. — Drake 1958c, p. 325. 
Type: Sex unknown; Madagascar; Helsin. Mus. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 



316 "U.S. NATIONAL MUSEUM BULLETIN 243 

*Paracopium floricolum (Horvath) 

Copium floricola Horvath 1910, p. 61 [Tanganyika]. 
Paracopium floricola: Horvath 1929, p. 322. 
Type: Sex unknown; Usambara, Tanganyika; Stockholm Mus. 
Distribution: Tanganyika. 
Host Plant: Unrecorded. 

Paracopium fraterculum Schouteden 

Paracopium f rater culum Schouteden 1955b, p. 166 [Congo]. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Paracopium furvum (Horvath) 

Copium furvum Horvath 1925a, p. 2 [Queensland]. 

Paracopium furvum: Horvath 1929, p. 322. 
Type: Holotype o 71 ; Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Paracopium ghesquierei (Schouteden) 

Copium ghesquierei Schouteden 1923, p. 87 [Congo]. 

Paracopium ghesquierei: Drake 1954f, p. 4. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Paracopium gigantos Drake 

Paracopium gigantos Drake 1954f, p. 3 [Malagasy]. 
Type: Holotype 9, macropterous; Perinet, Analamasotra, Madagascar; 
Drake Coll. (USNM). 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

*Paracopium glabricorne (Montandon) 

Eurycera glabricornis Montandon 1892, p. 267 [Mozambique]. — Distant 

1904, p. 430, pi. 8, fig. 9 [Cape Province; Natal; Transvaal]. 
Copium glabricorne: Lethierry and Severin 1896, p. 16. — Distant 1902a, 
p. 241.— Schouteden 1916b, p. 292 [Congo]; 1923, p. 88.— Mancini 
1939c, p. 308 [Somali].— Blote 1945, p. 85. 
Paracopium glabricorne: Horvath 1929, p. 322.- — Drake and Slater 
1955, p. 49 [Zululand]. 
Type: Sex unknown; Rikada, Mozambique; Montandon Coll. (Bucha- 
rest Mus.). 

Distribution: Mozambique; Congo; South Africa (Cape Province; 
Natal; Transvaal; Zululand); No. Rhodesia; Somali. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 317 

*Paracopium hamadryas (Drake) Plates 23, 24 

Copium hamadryas Drake 1925a, p. 1 [Congo; Clerodendron]. 
Paracopium hamadryas: Horvath 1929, p. 322 [Ghana; Clerodendron 

buchholzi]. — Drake and Davis 1960, fig. 52 [Uganda]. 
Copium {Paracopium) hamadryas: Mayne and Ghesquiere 1934, p. 20 
[Clerodendron subreniforme]. 
Type: Holotype 9; Thysville, Belgian Congo; American Mus. 
Distribution: Congo; Ghana; Uganda; Tanganyika. 
Host Plants: Clerodendron buchholzi; Clerodendron subreniforme; Clerodendron 
sp. 

Note: Figures of floral cecidia formed on Clerodendron sp. [Drake and 
Davis] . 
*Paracopium hirsutum Drake 

Paracopium hirsutum Drake 1953d, p. 218 [Malagasy]; 1958c, p. 326. 
Type: Holotype d% macropterous; Perinet, Analamasotra, Madagascar; 
Drake Coll. (USNM). 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

*Paracopium hirticorne (Bergroth) 

Copium hirticorne Bergroth 1912, p. 145 [Guinea]. 

Paracopium hirticorne: Horvath 1929, p. 323. 

Paracopium histocorne [sic]: Drake 1954a, p. 232 [Ghana]. 
Type: Holotype cT; Assini, Guinea; deposition unknown. 
Distribution: Guinea; Ghana. 
Host Plant: Unrecorded. 

*Paracopium insigne Schouteden 

Paracopium ? insigne Schouteden 1955b, p. 163 [Congo; Ruanda]; 
1957c, p. 313. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Ruanda-Urundi. 
Host Plant: Unrecorded. 

Paracopium kollari (Fieber) 

Laccometopus kollari Fieber 1844, p. 98, pi. 8, figs. 17-22 [Mauritius]. — 

Herrich-Schaeffer 1850, p. 158. 
Eurycera kollari: Stal 1873, p. 129. 
Copium kollari: Lethierry and Severin 1896, p. 16. 
Paracopium kollari: Horvath 1929, p. 323. — Drake and Mamet 1956, p. 
300; 1961, p. 223, fig.— Orian 1956, p. 647.— Mamet 1957, p. 56. 
Type: Holotype o", macropterous; Mauritius Island; Vienna Mus. 
Distribution: Mascarene Islands (Mauritius). 
Host Plant: Unrecorded. 
Note: Taken at light [Drake and Mamet 1961]. 



318 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Paracopium lewisi Distant 

Paracopium lewisi Distant 1903a, p. 48 [Ceylon]; 1903b, p. 128.— Horvath 
1929, p. 323. 
Type: Sex unknown; Ceylon; British Mus. 
Distribution: Ceylon; Greater Sunda Islands (Sumatra). 
Host Plant: Unrecorded. 

*Paracopium longulum Drake 

Paracopium longulum Drake 1958c, p. 323 [Malagasy]. 
Type: Holotype 9; Foret au Nord d'Anosibe, Madagascar; Madagascar 
Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

*Paracopium lupakense (Schouteden) 

Copium lupakense Schouteden 1923, p. 88 [Congo]. 
Paracopium lupakense: Drake 1954f, p. 4. 
Type: Sex unknown; Lupaka, Luapula, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

Paracopium modestum Schouteden 

Paracopium modestum Schouteden 1955b, p. 165 [Congo], 
Type: Sex unknown; Moto, Ituri, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 

*Paracopium parvum Drake 

Paracopium parvum Drake 1958c, p. 326 [Malagasy]. 
Type: Holotype o*; Tongobory, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 

*Paracopium philippinense Drake 

Paracopium philippinensis Drake 1930e, p. 165 [Sibuyan]. 
Type: Holotype o 71 , macropterous; Sibuyan Island, Philippine Islands; 
USNM. 
Distribution: Philippine Islands (Sibuyan; Negros). 
Host Plant: Unrecorded. 

*Paracopium proprium Drake 

Paracopium proprium Drake 1958c, p. 325 [Malagasy]. 

Type: Holotype cf ; Reserve Naturelle, Nosivola, Madagascar; Mada- 
gascar Sci. Inst. 

Distribution: Malagasy Republic. 

Host Plant: Unrecorded. 



LACEBTJGS OF THE WORLD 319 

*Paracopium sauteri Drake 

Paracopium sauteri Drake 1951, p. 171 [Taiwan]. 
Type: Holotype cf; Anping, Formosa; Hungarian Mus. 
Distribution: Taiwan. 
Host Plant: Unrecorded. 

*Paracopium stolidum (Horvath) 

Copium stolidum Horvath 1912a, p. 355 [Congo]. — Schouteden 1916b, 
p. 292; 1923, p. 88— Mayne and Ghesquiere 1934, p. 20 [Kalaharia 
(Clerodendron) spinescens]. 
Paracopium stolidum: Horvath 1929, p. 323.— Drake 1955b, p. 89 
[Angola]. 
Type: Holotype 9; Leopoldville, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Angola. 

Host Plants: Kalaharia (Clerodendron) spinescens; "in galls of old fallen 
flowers of the herb called by the Batshick { kakolu-ua-hachi' " [Drake 
1955b]. 
*Paracopium summervillei (Hacker) 

Teleonemia summervillei Hacker 1927, p. 22, pi. 6, fig. 4 [Queensland; 

Scaevola koenigii], 
Paracopium summetvillei [sic]: Drake 1947c, p. 112 [Dunk; Prince Wales 
Islands; New Hebrides]. 
Type: Sex unknown; Palm Island, Queensland, Australia; Queensland 
Mus. 

Distribution: Australia (Queensland; Palm, Dunk, Prince Wales 
Islands); New Hebrides Islands. 
Host Plant: Scaevola koenigii. 

Genus PARADA Horvath 

Cysteochila (Parada) Horvath 1925a, p. 3. 

Parada: Drake 1942a, p. 4; 1952a, pp. 143-147.— Drake and Ruhoff 
1960a, p. 74. 
Type Species: Cysteochila (Parada) taeniophora Horvath. 

*Parada absona Drake 

Parada absona Drake 1952, p. 146 [Queensland]. 
Type: Holotype cf; Lake Barrine, Queensland, Australia; MCZ. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Parada darlingtoni Drake 

Parada darlingtoni Drake 1952, p. 143 [Queensland]. 
Type: Holotype d"; National Park, McPherson Range, Queensland, 
Australia; MCZ. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 



320 U.S. NATIONAL MUSEUM BULLETIN 243 

*Parada hackeri Drake 

Parada hackeri Drake 1952, p. 145 [Queensland]. 
Type: Holotype d", macropterous; Mt. Gipps, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Parada popla Drake 

Cysteochila {Parada) taeniophora (not Horvath): Hacker 1927, p. 24 

[New South Wales; Queensland]; 1928, p. 180. 
Parada popla Drake 1942a, p. 3; 1952, p. 147; 1961b, p. 110. 
Type: Holotype cf, macropterous; National Park, Queensland, Aus- 
tralia; Drake Coll. (USNM). 
Distribution: Australia (Queensland; New South Wales). 
Host Plant: Unrecorded. 
*Parada popula var. nigrans Drake (emendation) 

Parada popla var. nigrantis Drake 1942a, p. 4 [Queensland]. 
Type: Holotype cf, macropterous; Mount Glorious, Queensland, 
Australia; Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Parada solla Drake and Ruhoff 

Parada solla Drake and Ruhoff 1961b, p. 139 [Queensland]. 
Type: Holotype cf; Mt. Spurgeon, Queensland, Australia; MCZ. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Parada taeniophora (Horvath) 

Cysteochila {Parada) taeniophora Horvath 1925a, p. 2, fig. 1 [Queensland]. 
Parada taeniophora: Drake 1952, p. 147. — Drake and Ruhoff 1962b, p. 
250 [New South Wales]. 
Type: Sex unknown; Malanda, Queensland, Australia; Stockholm Mus. 
Distribution: Australia (Queensland; New South Wales). 
Host Plant: Unrecorded. 
*Parada torta Drake 

Parada torta Drake 1942a, p. 4 [Queensland]; 1952, p. 147 [New South 
Wales]; 1961b, p. 110. 
Type: Holotype d\ macropterous; Mt. Gipps, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland; New South Wales). 
Host Plant: Unrecorded. 
Parada torta var. pulla Drake 

Parada torta was. pulla Drake 1952, p. 147 [New South Wales]. 
Type: Holotype 9; Mt. Wilson, New South Wales, Australia; MCZ. 
Distribution: Australia (New South Wales). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 321 

Genus PASEALA Schouteden 

Paseala Schouteden 1923, p. 93.— Drake and Runoff 1960a, p. 74. 
Type Species: Paseala arnoldi Schouteden. 

*Paseala arnoldi Schouteden 

Paseala arnoldi Schouteden 1923, p. 94 [Congo]. — Mayne and Ghes- 
quiere 1934, p. 20 [Apocynaceae]. — Drake and Gomez-Menor 1954, 
p. 92 [Spanish Guinea]. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Spanish Guinea. 
Host Plant: Apocynaceae. 

Genus PENOTTUS Distant 

Penottus Distant 1903c, p. 254.— Drake and Ruhoff 1960a, p. 75. 
Cetiocysta Drake and Poor 1939c, p. 205. — Drake and Ruhoff 1960a, p. 
44. 
Type Species: Penottus jalorensis Distant =Monanthia monticollis Walker. 

*Penottus bunus Drake and Ruhoff 

Penottus bunus Drake and Ruhoff 1961b, p. 179, fig. 20 [Borneo]. 
Type: Holotype 9, macropterous; Sandakan, Borneo; Drake Coll. 
(USNM). 
Distribution: Greater Sunda Islands (Borneo). 
Host Plant: Unrecorded. 

*Penottus monticollis (Walker) 

Monanthia monticollis Walker 1873a, p. 196 [Sarawak]. — Distant 1902b, 

p. 356. 
Penottus jalorensis Distant 1903c, p. 254, pi. 14 [Malaya]. — Blote 1945, 

p. 92 [Sumatra]. 
Diplocysta nubila Drake 1927a, p. 55 [Singapore]; 1930e, p. 167 

[Negros]. — -Drake and Poor 1937a, p. 11. 
Cetiocysta nubilia [sic]: Drake and Poor 1939c, p. 205. 
Penottus monticollis: Drake and Ruhoff 1960a, p. 75, pi. 8; 1962c, p. 133. 
Type: Sex unknown; Sarawak; British Mus. 

Distribution: Greater Sunda Islands (Sarawak; Sumatra); Federation 
of Malaya; Singapore; Philippine Islands (Negros). 
Host Plant: Unrecorded. 
Note : Only hemelytron remains of type specimen. 

*Penottus nimius (Drake) 

Diplocysta nimia Drake 1927a, p. 54 [Negros]. — Drake and Poor 1937a, 

p. 11 [Mindanao; Samar]. 
Cetiocysta nimia: Drake and Poor 1939c, p. 205. 
Penottus nimius: Drake and Ruhoff 1960c, p. 29. 



322 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Type: Holotype cf, macropterous; Curenos Mountains, Negros, 
Philippine Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Mindanao; Negros; Samar). 
Host Plant: Unrecorded. 
*Penottus opiparus (Drake) 

Diplocysta opipara Drake 1927c, p. 18 [Luzon]. 
Cetiocysta opipara: Drake and Poor 1939c, p. 205. 
Penottus opiparus: Drake and Runoff 1960c, p. 29. 
Type: Holotype o 71 , macropterous; Bagnio, Luzon, Philippine Islands; 
Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 
*Penottus oresbius Drake and Runoff 

Penottus oresbius Drake and Ruhoff 1961b, p. 178, figs. 19 a, b [Borneo; 
Malaya]. 
Type: Holotype 9, macropterous; Malacca, Southern Malay Peninsula; 
Drake Coll. (USNM). 
Distribution: Federation of Malaya; Greater Sunda Islands (Borneo). 
Host Plant: Unrecorded. 
*Penottus tibetanus Drake and Maa 

Penottus tibetanus Drake and Maa 1954, p. 115 [Tibet]. 
Type: Holotype d% macropterous; Tibet; Drake Coll. (USNM). 
Distribution: Tibet. 
Host Plant: Unrecorded. 
*Penottus verdicus Drake and Maa 

Penottus verdicus Drake and Maa 1953, p. 88 [Taiwan]. 
Type: Holotype o 71 , macropterous; Formosa; Drake Coll. (USNM). 
Distribution: Taiwan. 
Host Plant: Unrecorded. 

Genus PERBRINCKEA Drake 

Perbrinckea Drake 1956f, p. 427.— Drake and Ruhoff 1960a, p. 75. 
Type Species: Perbrinckea brincki Drake. 
*Perbrinckea brincki Drake Plate 16 

Perbrinckea brincki Drake 1956f, p. 427, fig. 1 [Basutoland]. 
Type: Holotype c 71 ; Mount Morosi, Basutoland, South Africa; Lund 
Zool. Inst. 
Distribution: Basutoland. 
Host Plant: Unrecorded. 

Genus PERISSONEMIA Drake and Poor 

Perissonemia Drake and Poor 1937a, p. 2. — Drake and Ruhoff 1960a, 
p. 75. 
Type Species: Perissonemia torquata Drake and Poor. 



LACEBUGS OF THE WORLD 323 

*Perissonemia absita Drake and Ruhoff 

Perissonemia absita Drake and Ruhoff 1961b, p. 137, fig. 11 [Mindanao]. 

Type: Holotype cf, macropterous; Butuan, Mindanao, Philippines; 
Drake Coll. (USNM). 

Distribution: Philippine Islands (Mindanao). 

Host Plant: Unrecorded. 

*Perissonemia bimaculata (Distant) 

Teleonemia bimaculata Distant 1909c, p. 166, pi. 10, figs. 6, 6a [Borneo]. 

Cromerus bimaculatus: Drake 1953b, p. 92. 

Perissonemia bimaculata: Drake and Ruhoff 1961c, p. 145. 
Type: Holotype o 71 ; Kuching, Borneo; British Mus. 
Distribution: Greater Sunda Islands (Borneo); India. 
Host Plant: Unrecorded. 

*Perissonemia borneenis (Distant) 

Teleonemia borneenis Distant 1909c, p. 166, pi. 10, figs. 1, la [Borneo]. 
Perissonemia (Ulonemia) borneensis [sic]: Drake and Poor 1937a, p. 4 

[Singapore]. 
Perissonemia borneensis [sic]: Drake and Poor 1937b, p. 401 [Mindanao; 

Sibuyan].— Drake 1938b, p. 195 [China]. 
Perissonemia borneenis: Drake 1960, p. 356 [New Guinea]. 
Type: Sex unknown; Kuching, Borneo; British Mus. 
Distribution: Greater Sunda Islands (Borneo), Singapore; China; 
Philippine Islands (Luzon; Mindanao; Sibuyan); New Guinea (Northeast). 
Host Plant: Unrecorded. 

*Perissonemia delagoana Drake 

Perissonemia delagoana Drake 1954b, p. 9 [Mozambique]. 
Type: Holotype cf; Dalagoa Bay, Mozambique; British Mus. 
Distribution: Mozambique. 
Host Plant: Unrecorded. 

*Perissonemia ecmeles Drake and Mohanasundarum Plate 25 

Perissonemia ecmeles Drake and Mohanasundarum 1961, p. 110, fig. 2 
[India; Ficus], 
Type: Holotype o", macropterous; Coimbatore, India; Drake Coll. 
(USNM). 
Distribution: India. 
Host Plant: Ficus sp. 

*Perissonemia gressitti Drake and Poor 

Perissonemia gressitti Drake and Poor 1936a, p. 440 [Hainan]. 

Type: Holotype cf, macropterous; Nodoa, Hainan Island; Drake Coll. 
(USNM). 

Distribution: China (Hainan Island). 

Host Plant: Unrecorded. 



324 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Perissonemia illustris Drake and Poor 

Perissonemia {Ulonemia) illustris Drake and Poor 1937a, p. 4 [Luzon]. 
Perissonemia illustrus [sic]: Drake and Poor 1937b, p. 401. 
Type: Holotype o 71 , macropterous; Imugin, Nueva Vizcaya Province, 
Luzon, Philippine Islands; USNM. 
Distribution: Philippine Islands (Luzon); Singapore. 
Host Plant: Unrecorded. 
*Perissonemia kietana Drake and Runoff 

Perissonemia kietana Drake and Ruhoff 1961b, p. 136 [Bougainville]. 
Type: Holotype o 71 , macropterous; Kieta, Solomon Islands; Drake Coll. 
(USNM). 
Distribution: Solomon Islands (Bougainville). 
Host Plant: Unrecorded. 
*Perissonemia nigerrima (Schouteden) 

Teleonemia nigerrima Schouteden 1923, p. 104 [Congo]. 
Perissonemia nigerrima: Drake and Ruhoff 1960c, p. 29. 
Type: Sex unknown; Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo. 
Host Plant: Unrecorded. 
*Perissonemia occasa Drake 

Perissonemia occasa Drake 1942a, p. 2 [Japan]. — Drake and Ruhoff 

1961b, p. 135, fig. 10. 
Perissonemia {Ulonemia) occasa: Takeya 1951a, p. 18. 
Type: Holotype o 71 , macropterous; Japan; Drake Coll. (USNM). 
Distribution: Japan; China. 
Host Plant: Unrecorded. 
*Perissonemia onerosa Drake and Poor 

Perissonemia onerosa Drake and Poor 1939c, p. 204 [India; sandal]. 
Type: Holotype 9, macropterous; Vellore, Madras, India; Drake Coll. 
(USNM). 
Distribution: India. 

Host Plant: Taken on "unspiked sandal." 
*Perissonemia pagnana Drake 

Perissonemia pagnana Drake 1954b, p. 8 [Uganda]. 
Type: Holotype 9, macropterous; Kampala, Uganda; Drake Coll. 
(USNM). 
Distribution: Uganda. 
Host Plant: Unrecorded. 
*Perissonemia sandakana Drake and Ruhoff 

Perissonemia sandakana Drake and Ruhoff 1961b, p. 134, fig. 9 [Borneo]. 
Type: Holotype o 71 , macropterous; Sandakan, Borneo; Drake Coll. 
(USNM). 
Distribution: Greater Sunda Islands (Borneo). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 325 

*Perissonemia sierrana Drake 

Perissonemia sierrana Drake 1954b, p. 9 [Sierra Leone; "Bomamagbai"]. 
Type: Holotype cf; Njala, Sierra Leone; British Mus. 
Distribution: Sierra Leone. 
Host Plant: Taken on "Bomamagbai." 

*Perissonemia sodalis Drake 

Perissonemia sodalis Drake 1948e, p. 151 [Queensland]. — Drake and 
Davis I960, fig. 65. 
Type: Holotype cf 1 , macropterous; Redlynch, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

*Perissonemia torquata Drake and Poor 

Perissonemia torquata Drake and Poor 1937a, p. 2, pl. 1 [Mindanao]; 
1937b, p. 401.— Drake 1960, p. 356 [Solomon; New Guinea]. 
Type: Holotype o 71 , macropterous; Surigao, Mindanao, Philippine Is- 
lands; Drake Coll. (USNM). 

Distribution: Philippine Islands (Mindanao); New Guinea; Solomon 
Islands. 
Host Plant: Unrecorded. 

*Perissonemia vegata Drake and Poor 

Perissonemia {Ulonemia) vegata Drake and Poor 1937b, p. 401 [Borneo]. 

Type: Holotype 9, macropterous; Sandakan, Borneo; Drake Coll. 
(USNM). 

Distribution: Greater Sunda Islands (Borneo). 

Host Plant: Unrecorded. 

Genus PHAENOTROPIS Horvath 

Monosteira (Phaenotropis) Horvath 1906a, p. 106. — Oshanin 1908, p. 456; 

1912, p. 46. 
Phaenotropis: Drake 1957a, p. 415. — Drake and Runoff 1960a, p. 76. 
Type Species: Monanthia {Monosteira) parvula Signoret. 

*Phaenotropis cleopatra (Horvath) 

Monosteira cleopatra Horvath 1905b, p. 572 [Egypt]; 1911c, p. 109. 

Monosteira {Phaenotropis) cleopatra: Horvath 1906a, p. 107. 

Monostira [sic] {Phaenotropis) cleopatra: Priesner and Alfieri 1953, p. 66 

[Panicum turgidum]. 
Phaenotropis cleopatra: Drake 1957a, p. 415 [Aden]. 
Type: Sex unknown; Egypt; Vienna Mus. 
Distribution: Egypt; Aden Protectorate; India; Israel. 
Host Plants: Panicum turgidum; Tephrosia sp. 



326 U.S. NATIONAL MUSEUM BULLETIN 243 

*Phaenotropis parvula (Signoret) 

Monanthia {Monosteira) parvula Signoret 1865, p. 117 [France]. 
Monanthia parvula: Walker 1873a, p. 190.— Puton 1879c, p. 124.— Rey 

1893, p. 97 [Dorycnium suffruticosum]. 
Monosteira (Phaenotropis) parvula: Horvath 1906a, p. 107. 
Monosteira parvula: Stichel 1926, p. 116; 1935, p. 349; 1938a, p. 408; 
1960a, p. 344, fig. 182 [Sardinia]; 1960b, p. 401; 1960c, p. 140.— 
Blote 1945, p. 92. 
Monostira [sic] parvula: Guide 1938, p. 321. — Gomez-Menor 1949, 

p. 161, fig. [Spain]; 1955b, p. 249. 
Phaenotropis parvula: Drake 1957a, p. 415. 
Type: Sex unknown; France; Vienna Mus. 
Distribution: France; Spain; Sardinia; Senegal. 
Host Plant: Dorycnium suffruticosum. 

Genus PHAEOCHILA Drake and Hambleton 

Phaeochila Drake and Hambleton 1945, p. 358. — Drake and RuhofF 
1960a, p. 75. 
Type Species: Amblystira hirta Monte. 

*Phaeochila hirta (Monte) 

Amblystira hirta Monte 1940e, p. 284, fig. 5 [Brazil]. 
Phaeochila hirta: Drake and Hambleton 1945, p. 359 [Peru]. 
Type: Holotype 9; Catanduva, Sao Paulo, Brazil; Monte Coll. (Mus. 
Nacional). 
Distribution: Brazil; Peru. 
Host Plant: Unrecorded. 

Genus PHYMACYSTA Monte 

Phymacysta Monte 1942d, p. 106. — Hurd 1946, p. 476. — Drake and 
RuhofF 1960a, p. 77. 
Type Species: Leptostyla tumida Champion. 

*Phymacysta magnifica (Drake) 

Gelchossa magnifica Drake 1922c, p. 373 [Brazil]. 
Leptopharsa magnifica: Drake 1930a, p. 1; 1931b, p. 511; 1935, p. 18 
[Paraguay]. — Drake and Hambleton 1938a, p. 63 [Chuquiragua 
glabra].— -Monte 1939b, p. 74; 1941e, p. 115 [Argentina]. 
Phymacysta magnifica: Monte 1942d, p. 107. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil; Paraguay; Argentina. 
Host Plant: Chuquiragua glabra. 



LACEEUGS OF THE WORLD 327 

*Phymacysta mcelfreshi (Drake) 

Leptostyla mcelfreshi Drake 1918a, p. 176 [Haiti]. 
Gelehossa mcelfreshi: Drake and Bruner 1924a, p. 149. 
Phymacysta mcelfreshi: Hurd 1946, p. 476. 
Type: Holotype cf, macropterous; Port-au-Prince, Haiti; Drake Coll. 
(USNM). 
Distribution: Haiti. 
Host Plant: Unrecorded. 

*Phymacysta praestans (Drake) (emendation) 

Gelehossa praestantis Drake 1927b, p. 116 [Mexico]. 
Phymacysta praestantis: Monte 1942d, p. 107. 
Type: Holotype 9, macropterous; Montana de Sumidero, Chiapas, 
Mexico; Drake Coll. (USNM). 
Distribution: Mexico. 
Host Plant: Unrecorded. 

*Phymacysta tumida (Champion) 

Leptostyla tumida Champion 1897, p. 14, pi. 1, figs. 17, 17a [Guate- 
mala]. — Van Duzee 1907, p. 19 [Jamaica]. 
Leptostyla malpigheae Drake 1922b, p. 38 [Cuba; Malpighia urens]. 
Gelehossa malpigheae: Drake and Bruner 1924a, p. 149. 
Gelehossa tumida: Drake and Bruner 1924a, p. 150 [Haiti; Trinidad; 

cherry]. — Gowdy 1926, p. 35. 
Leptopharsa walcotti Drake 1928a, p. 55. 
Leptopharsa cub ana Drake 1928 a, p. 56. 

Leptopharsa tumida: Monte 1939b, p. 75 [Brazil; Venezuela; Adeno- 
calymma bracteatum]; 1940d, p. 105 [Malpighia punicifolia]; 1941e, 
p. 119. 
Phymacysta tumida: Monte 1942d, p. 106, fig. 2.- — Drake and Hambleton 
1945, p. 362 [Peru; Ecuador; Panama].— Hurd 1946, p. 477 [Mal- 
pighia glabra]. — Drake and Cobben 1960, p. 85, fig. 85 [Tex.; Mexico; 
Aruba; Curacao; Bonaire; St. Eustatius]. — -Drake and Runoff 
1962a, p. 156. 
Phymacysta malpigheae: Monte 1942d, p. 107. 
Phymacysta cubana: Monte 1942d, p. 107. 
Phymacysta walcotti: Monte 1942d, p. 107. 

Leptopharsa malpigheae: Bruner, Scaramuzza, and Otero 1945, p. 105. 
Phymacysta malpighae [sic]: Drake and Hambleton 1945, p. 362. 
Type: Sex unknown; Pantaleon, Guatemala; British Mus. 
Distribution: Brazil; Ecuador; Venezuela; Peru; Panama; Guatemala; 
Mexico; U.S. (Tex.); Jamaica; Haiti; Trinidad; Cuba; Netherlands 
Antilles (Aruba; Bonaire; Curagao); Leeward Islands (St. Eustatius). 

Host Plants: Adenocalymma bracteatum; Malpighia glabra; Malpighia punici- 
folia; Malpighia urens; cherry. 



328 U.S. NATIONAL MUSEUM BULLETIN 243 

Phymacysta vesiculosa (Champion) 

Leptostyla vesiculosa Champion 1897, p. 13, pi. 1, figs. 15, 15a [Panama]. 

Phymacysta vesiculosa: Monte 1942d, p. 107. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

Genus PHYSATOCHEILA Fieber 

Monanthia (Physatocheila) Fieber 1844, p. 80. 

Physatochila: Stal 1873, pp. 120, 129; 1874, p. 56.— Lethierry and 
Severin 1896, p. 21.— Oshanin 1908, p. 447; 1912, p. 45.— Banks 
1910, p. 57.— Osborn and Drake 1916a, p. 242 (key) .— Stichel 1935, 
p. 349; 1938a, p. 408; 1938b, p. 454.— Guide 1938, pp. 245, 305 
(key). — Singer 1952, p. 54. 
Phyllochisme Kirkaldy 1904, p. 280.— Drake and Runoff 1960a, p. 76. 
Physatocheila: Horvath 1906a, pp. 15, 94 (key). — Muzik 1907, pp. 51, 
61.— Van Duzee 1916, p. 25; 1917b, p. 219.— Osborn and Drake 
1917a, p. 155 (key).— Stichel 1926, pp. 105, 114 (key); 1960a, p. 332; 
1960b, p. 399; 1960c, p. 138.— de Seabra 1931, pp. 408, 432.— China 
1943, p. 247; 1952, p. 50 (key).— Hurd 1946, p. 451.— Kiritshenko 
1951, pp. 242, 252 (key).— Drake and Ruhoff 1960a, p. 77. 
Type Species: Acanthia quadrimaculata Wolff= Acant hia costata Fabricius. 
*Physatocheila aglaia Drake and Ruhoff 

Physatocheila aglaia Drake and Ruhoff 1961b, p. 131, fig. 8 [Burma]. 
Type: Holotype cf, macropterous; Mandalay, Burma; Drake Coll. 
(USNM). 
Distribution: Burma. 
Host Plant: Unrecorded. 

*Physatocheila arbicola Drake 

Physatocheila arbicola Drake 1958a, p. 102 [Angola]. 
Type: Holotype c? ; Cazombo, Angola; Cent. Afr. Mus. 
Distribution: Angola. 
Host Plant: Unrecorded. 

*Physatocheila biseriata Hacker 

Physatochila biseriata Hacker 1929, p. 327, pi. 33, fig. 5 [Queensland]. 

Type: Sex unknown; Brookfield, Queensland, Australia; Queensland 
Mus. 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

*Physatocheila brevirostris Osborn and Drake 

Physatochila brevirostris Osborn and Drake 1916a, p. 243 [Ohio; N.Y.]. 

Physatocheila brevirostris: Osborn and Drake 1917a, p. 155. — Parshley 

1917b, p. 56 [Mass.]; 1922a, p. 236 [Conn.]; 1923b, p. 706— Barber 



LACEBUGS OF THE WORLD 329 

1922a, p. 17.— McAtee 1923, p. 145 [Md.; Va.].— Blatchley 1926, 
p. 484 [N.J.].— Drake 1928b, p. 102.— Hurd 1946, p. 452 [Que.; 
Pa.; 111.].— Bailey 1951, p. 24.— Ash 1954, p. 185 [Ind.]. 
Type: Holotype 9, macropterous; Chillicothe, Ross Co., Ohio, U.S.; 
Ohio State Univ. 

Distribution: U.S. (Mass.; Conn.; N.Y.; Pa.; N.J.; Md.; Va.; Ohio; 
111.; Ind.; Iowa); Canada (Que.). 
Host Plant: Unrecorded. 

*Physatocheila chatterjeei Drake and Poor 

Physatocheila chatterjeei Drake and Poor 1936b, p. 142 [India]. 

Type: Holotype 9, macropterous; Bodyar, Chakrata Division, United 
Province; Drake Coll. (USNM). 

Distribution: India. 

Host Plant: Unrecorded. 

*Physatocheila civatis Drake 

Physatocheila civatis Drake 1942a, p. 10 [New South Wales]; 1961b, p. 
107 [Queensland]. 
Type: Holotype o", macropterous; Mt. Wilson, New South Wales, 
Australia; Drake Coll. (USNM). 
Distribution: Australia (New South Wales; Queensland). 
Host Plant: Unrecorded. 
*Physatocheila confinis Horvath 

Tingis {Physatocheila) quadrimaculata (not Wolff): Kolenati 1856, p. 427 

(in part) [Caucasus]. 
Physatocheila confinis Horvath 1905b, p. 570 [Crimea; Armenia; Hun- 
gary; Yugoslavia; Turkey; Greece]; 1906a, p. 95 [Syria; Py r us com- 
munis].— Stichel 1926, p. 114; 1960a, p. 333 [France].— Kiritshenko 
1951, p. 252. — Novak and Wagner 1951, p. 71 [Qiiercus pubescens; 
Ostrya carpinifolia]. — Hoberlandt 1955, p. 94 [Transcaucasus; Qiiercus 
ilex; Quercus cocci/era]. 
Physatochila confinis: Stichel 1938a, p. 408 [Austria; Albania]. — Gomez- 
Menor 1954, p. 390. 
Type: Unknown. 

Distribution: France; Austria; Hungary; Yugoslavia; Albania; Greece; 
Turkey; Syria; U.S.S.R. (Caucasus; Transcaucasus; Crimea; Armenia). 
Host Plants: Ostrya carpinifolia; Pyrus communis; Qiiercus cocci/era; Quercus 
ilex; Quercus pubescens. 
Physatocheila confinis var. putoni Horvath 

Monanthia dumetorum var. Puton 1881, p. 123. 
Physatocheila confinis var. putoni Horvath 1911b, p. 584 [Syria]. 
Type: Holotype 9; Damascus, Syria; Hungarian Mus. 
Distribution: Syria. 
Host Plant: Unrecorded. 

6S1-552— 64 22 



330 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Physatocheila costata (Fabricius) 

Acanthia costata Fabricius 1794, p. 77. 

Tingis costata: Fabricius 1803, p. 125.— Latrielle 1804, p. 253. 

Acanthia quadrimaculata Wolff 1804, p. 132, pi. 13, fig. 127. 

Tingis 4-maculata: Fallen 1829, p. 144.— Herrich-Schaeffer 1835, p. 58. 

Tingis corticea Herrich-Schaeffer 1830, heft 18, tab. 22 [Germany]. 

Monanthia 4-maculata: Burmeister 1835, p. 261. — Thomson 1871, p. 
398.— Jakovlev 1893, p. 294. 

Monanthia quadrimaculata: Herrich-Schaeffer 1838, p. 58, pi. 125, fig. a. — 
Mayr 1858, p. 571.— Fieber 1861, p. 124.— Frey-Gessner 1865, p. 232 
[Switzerland; Chrysanthemum]. — Garbiglietti 1869, p. 273. — Gredler 
1870, p. 75.— Jakovlev 1875b, p. 262; 1880b, p. 108.— Reiber and 
Puton 1876, p. 69.— Hiieber 1893, p. 345.— Jensen-Haarup 1912, 
pp. 155, 156, fig. 99 [Denmark]. 

Monanthia {Physatocheila) quadrimaculata: Fieber 1844, p. 81, pi. 7, figs. 
1-3 [Czechoslovakia; Austria; Italy; Sweden]. — Herrich-Schaeffer 
1850, p. 154. 

Monanthia costata: Scholz 1847, p. 120. — Mayr 1858, p. 570. — Frey- 
Gessner 1865, p. 232.— Garbiglietti 1869, p. 273.— Gredler 1870, p. 
75.— Puton 1874a, p. 227.— Ferrari 1885, p. 411.— d'Antessanty 
1890, p. 33. 

Monanthia {Tropidocheila) quadrimaculata: Sahlberg 1848, p. 132 [Finland]. 

Monanthia {Physatocheila) 4-maculata: Flor 1860, p. 350 [France; 
Caucasus; Transcaucasus; Estonia; Latvia; Arnebia cornuta]. 

Laccometopus costata: Stal 1868, p. 92. 

Eurycera costata: Stal 1873, p. 129. 

Physatochila quadrimaculata: Stal 1873, p. 129; 1874, p. 57. — Reuter 
1882b, p. 120 [Alnus].— Oshanin 1908, p. 448 [Belgium; Hungary].— 
Schumacher 1914, p. 258 (in part) [Quercus]. — Scholte 1935, p. 78, 
fig. 22 [Netherlands].— Borner 1935, p. 78.— Guide 1938, p. 308, fig. 
[Alnus incana]. 

Monanthia 4-macalata [sic]: Siebke 1874, p. 23 [Norway]. 

Physatochila 4-maculata: Sahlberg 1878, p. 21. — Lindberg 1927, p. 20; 
1932a, p. 43 [Spain]. 

Monanthia {Physatochila) quadrimaculata: Puton 1879c, p. 120. 

Copium costatum: Horvath 1906a, p. 94. 

Physatocheila quadrimaculata: Horvath 1906a, pp. 94, 95 [Siberia; Alnus 
glutinosa]. — Reichensperger 1920, p. 62. — Sahlberg 1920, p. 84 
[Populus tremula].— Stichel 1926, p. 114, fig. 204; 1960a, p. 164 
[Poland; Yugoslavia; Rumania; Betula]. — Hoberlandt 1942, p. 126; 
1943b, p. 124.— Blote 1945, p. 89.— Kiritshenko 1951, p. 252 — 
China 1952, p. 49, figs. 1 b, d [Alnus montana]. — Stusak 1957b, p. 
140.— Bator 1953a, p. 326, pi. 3, fig. 4.— Cobben 1958a, p. 10, figs. 
11, 12, 14. 



LACEBUGS OF THE WORLD 331 

Paracopium costatum: Horvath 1929, p. 322. 

Physatocheila costata: Drake and RuhofF 1960a, p. 77, pi. 9. 

Type: Sex unknown; "Europa boreali"; Kiel Coll. (Copenhagen Mus.). 

Distribution: Spain; France; Belgium; Netherlands; Germany; Czech- 
oslovakia; Switzerland; Austria; Hungary; Poland; Italy; Yugoslavia; 
Rumania; Norway; Sweden; Finland; Denmark; Turkey; U.S.S.R. 
(Estonia; Latvia; Siberia; Caucasus; Transcaucasus). 

Host Plants: Alnus glutinosa; Alnus montana; Alnus incana; Alnus sp.; 
Arnebia cornuta; Betula sp.; Chrysanthemum sp.; Populus tremula; Quercus sp. 

Note: Species not found in England [China]. 

*Physatocheila costata subsp. smreczynskii China 

Tingis quadrimaculata (not Wolff): Curtis 1839, tab. 741 [England; 

apple]. 
Monanthia quadrimaculata: Douglas and Scott 1865, p. 247. — Saunders 

1875, p. 249. 
Monanthia {Physatochila) quadrimaculata: Saunders 1892, p. 135, pi. 12, 

fig. 9. 
Physatochila quadrimaculata: Oshanin 1908, p. 448 (in part); 1912, p. 45 

(in part).— Schumacher 1914, p. 258 (in part).— Butler 1923, p. 213, 

fig. (in part). 
Physatocheila quadrimaculata: China 1936, p. 271, fig. lb; 1943, p. 247. 
Physatocheila smreczynskii China 1952, p. 49, fig. lc [Poland; Austria; 

Primus padus; Sorbus]. — Cobben 1958a, p. 10, figs, lla-c [Netherlands; 

Sorbus aucuparia]. — Forster 1959, p. 75.- — Southwood and Leston 1959, 

p. 150, fig. 53. — Stichel 1960a, p. 334 [Sorbus intermedia; Pyrus com- 
munis; Pyrus malus]. 
Physatochila smreczynskii: Gomez-Menor 1955a, p. 260 [Spain]; 1955b, 

p. 249. 
Physatocheila smreczynsky [sic]: Gomez-Menor 1956a, p. 112. 
Physatocheila quadrimaculata subsp. smreczynskii: Wagner 1960, pp. 83-92, 

4 pis. 
Type: Holotype d"; New Forest, Hants, England; British Mus. 
Distribution: England; Spain; Poland; Austria; Netherlands; Germany. 
Host Plants: Crataegus sp.; Prunus padus; Pyrus communis; Pyrus malus; 
Sorbus aucuparia; Sorbus intermedia; Sorbus sp. 

*Physatocheila delicatula Horvath 

Physatocheila delicatula Horvath 1903a, p. 78 [Algeria]; 1906a, p. 95. — 

Drake and Ruhoff 1961c, p. 145. 
Physatocheila chanceli Bergevin 1920, p. 412, fig. 1. 
Type: Holotype cf; Mont Edough, Algeria; deposition unknown. 
Distribution: Algeria. 
Host Plant: Unrecorded. 



332 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Physatocheila distinguenda (Jakovlev) 

Monanthia (Physatochila) distinguenda Jakovlev 1880a, p. 139 [Russia]. 
Physatocheila distinguenda: Horvath 1906a, p. 96. — Kiritshenko 1951, p. 
252.— Stichel 1960a, p. 335 [Turkestan]. 
Type: Holotype 9, macropterous; Sarepta, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 

*Physatocheila dryadis Drake and Poor 

Physatocheila dryadis Drake and Poor 1936b, p. 143 [India; Quercus 
dilatata].— Singh 1953, p. 119. 
Type: Holotype d\ macropterous; Ramgarh, Naini Tal, United Prov- 
inces; Drake Coll. (USNM). 
Distribution: India. 
Host Plant: Quercus dilatata. 

*Physatocheila dumetorum (Herrich-Schaeffer) 

Monanthia dumetorum Herrich-Schaeffer 1838, p. 57, pi. 124, figs. 391, 
a, b, d, e — Scholz 1847, p. 120.— Mayr 1858, p. 571 [Hungary].— 
Fieber 1861, p. 125 [Mespilus oxyacantha]. — Douglas and Scott 1865, 
p. 246. — Frey-Gessner 1865, p. 232 [Switzerland; Juniperus]. — • 
Garbiglietti 1869, p. 273 [Italy].— Jakovlev 1876b, p. 67 [Russia].— 
Horvath 1874b, p. 432.— Saunders 1875, p. 249.— Reiber and Puton 
1876, p. 69.— Bolivar and Chicote 1879, p. 166 [Spain].— Dubois 
1888, p. 121.— d'Antessanty 1890, p. 33.— Hiieber 1893, p. 346 
[Prunus padus]. — Jensen-Haarup 1912, pp. 155, 157 [Denmark]. — 
Butler 1923, p. 214 [Wales]. 

Tingis oxyacanthae Curtis 1839, pi. 16, tab. 741 [England; whitethorns]. 

Monanthia {Physatocheila) dumetorum: Fieber 1844, p. 82, pi. 7, figs. 4-6 
[Czechoslovakia; Germany; Austria]. — Herrich-Schaeffer 1850, p. 
154.— Saunders 1892, p. 135. 

Monanthia quadrimaculata (not Wolff): Costa 1847a, p. 22; 1847c, p. 
162.— Vollenhoven 1878, p. 276, pi. 9, figs. 8, 8a [Netherlands]. 

Physatochila dumetorum: Stal 1874, p. 57. — Reuter 1891a, p. 26 [Greece]; 
1908, p. 88 [Crataegus].— Oshanin 1908, p. 447 [Belgium; Portugal; 
Crimea].— Schumacher 1914, p. 258.— Lindberg 1932a, p. 43 
[Morocco]; 1948, p. 59 [Cyprus; Amygdalus; Acer obtusifolium]. — 
Scholte 1935, p. 78— Borner 1935, p. 78.— Stichel 1938a, p. 408 
[Turkey]; 1938b, p. 455— Blote 1945, p. 88 [Tunisia; Algeria].— 
Mancini 1949b, p. 37; 1953b, p. 186; 1953c, p. 22; 1953d, p. 16 
[Sicily]. — Priesner and Alfieri 1953, p. 65. — Gomez-Menor 1954, p. 
388, fig. 8 [Pinus]; 1955a, p. 261, fig. 36; 1955b, p. 249; 1956a, p. 115, 
fig. 9. 

Monanthia (Physatochila) dumetorum: Puton 1879c, p. 120 [France]. 



LACEBUGS OF THE WORLD 333 

Physatocheila dumetorwn: Horvath 1906a, p. 95 [Yugoslavia; Bulgaria; 
Egypt; Crataegus oxycantha; Crataegus monogyna; Primus domestical 
Primus spinosa]; 1906d, p. 1; 1916, p. 9 [Albania].— Reichensperger 
1920, p. 62. — Stichel 1926, p. 114 [Pyrus malus; Pyrus communis]; 
1960a, p. 332, fig. 159 [Poland; Acer creticum].—de Seabra 1931, pp. 
433, 444, figs. 504(1-5), 505.— China 1936, p. 271, fig. la.— Hober- 
landt 1942, p. 126; 1943b, p. 124; 1955, p. 94 [Israel].— Kiritshenko 
1951, p. 252— Stusak 1957b, p. 140; 1961a, p. 82, fig. 10; pi. 1, 
fig. a. — Cobben 1958a, p. 12, fig. 13. — Southwood and Leston 
1959, p. 150, pi. 21, fig. 5; pi. 25, fig. 1; fig. 51. 
Type: Unknown. 

Distribution: Portugal; Spain; France; Netherlands; Belgium; Wales; 
England; Denmark; Germany; Czechoslovakia; Poland; Austria; Switzer- 
land; Hungary; Italy; Sicily; Yugoslavia; Greece; Bulgaria; Albania; 
Cyprus; Turkey; U.S.S.R. (Crimea); Israel; Egypt; Algeria; Morocco; 
Tunisia. 

Host Plants: Acer creticum; Acer obtusifolium; Amygdalus sp.; Crataegus 
monogyna; Crataegus oxycantha; Crataegus sp.; Juniperus sp.; Mespilus oxyacantha; 
Pinus sp.; Primus domestica; Prunus padus; Prunus spinosa; Pyrus communis; 
Pyrus malus. 
Note: Life history study [Butler]. Eggs [Stusak]. 

*Physatocheila enalla Drake and Ruhoff 

Physatocheila enalla Drake and Ruhoff 1961b, p. 132 [Burma]. 

Type: Holotype 9, macropterous; Mandalay, Burma; Drake Coll. 
(USNM). 

Distribution: Burma. 

Host Plant: Unrecorded. 

*Physatocheila enodis Drake 

Physatochila enodis Drake 1948a, p. 3 [China], 

Type: Holotype cf , macropterous; Tsinleong, East Kwantungs, China; 
Drake Coll. (USNM). 

Distribution: China. 

Host Plant: Unrecorded. 

*Physatocheila exolasca Drake 

Physatocheila exolasca Drake 1954b, p. 2 [India]. 

Type: Holotype o", macropterous; Dehra Dun, India; Drake Coll. 
(USNM). 

Distribution: India. 

Host Plant: Unrecorded. 



334 U.S. NATIONAL MUSEUM BULLETIN 243 

Physatocheila fasciata (Fieber) 

Monanthia (Physatocheila) fasciata Fieber 1844, p. 84, pi. 7, figs. 13-16 

[Ostindien]. — Herrich-Schaeffer 1850, p. 154. 
Monanthia fasciata: Stal 1873, p. 134 [India orientalis]. — Distant 

1903b, p. 145 [India], 
Physatocheila fasciata: Drake 1937b, p. 592 [China]. 
Type: "Ostindien"; sex and deposition of type unknown. 
Distribution: China; India. 
Host Plant: Unrecorded. 
Physatocheila ferruginea Bergroth 

Physatocheila ferruginea Bergroth 1912, p. 146 [Guinea]. 
Type: Holotype 9; Assini, Guinea; Helsin. Mus. 
Distribution: Guinea. 
Host Plant: Unrecorded. 
*Physatocheila fulgoris Drake 

Physatocheila fulgoris Drake 1937b, p. 592 [China]. 
Type: Holotype d\ macropterous; Wong-sa-shue, S. Kiangsi, South 
China; Drake Coll. (USNM). 
Distribution: China. 
Host Plant: Unrecorded. 
Physatocheila gibba (Fieber) 

Monanthia (Physatocheila) gibba Fieber 1844, p. 83, pi. 7, figs. 7-12 

[Ostindien].— Herrich-Schaeffer 1850, p. 154. 
Monanthia gibba: Stal 1873, p. 134. — Lethierry and Severin 1896, p. 25. 
Physatocheila gibba: Drake and Ruhoff 1960c, p. 29. 
Type: "Ostindien"; sex and deposition of type unknown. 
Distribution: Uncertain. 
Host Plant: Unrecorded. 
*Physatocheila harwoodi China 

Physatocheila harwoodi China 1936, p. 271, fig. 1c [England; maples]; 
1943, p. 247.— Stichel 1938b, p. 455 [Germany; Acer]; 1960a, p. 
335 [Acer platanoides; Acer pseudoplatanus]. — Southwood and Leston 
1959, p. 151, fig. 52. 
Type: Sex unknown; Witchamton, Dorsetshire, England; British Mus. 
Distribution: England; Germany. 
Host Plants: Acer platanoides; Acer pseudoplatanus; Acer sp. 
Physatocheila irregularis Montrouzier 

Physatocheila irregularis Montrouzier 1861, p. 68 [Lifu]. — Stal 1873, 

p. 134.— Drake and Ruhoff 1961c, p. 145, fig. 1. 
Monanthia irregularis: Walker 1873a, p. 196. 
Type: Holotype cf , macropterous; Lifu, Loyalty Islands; Vienna Mus, 
Distribution: Loyalty Islands (Lifu). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 335 

*Physatocheila katbergana Drake 

Physatochelia [sic] katbergana Drake 1953d, p. 212 [Cape Province]. 
Type: Holotype o*; Katberg, Cape Province, South Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

*Physatocheila lautana Drake and Ruhoff 

Physatochelia lautana Drake and Ruhoff 1961b, p. 133 [Laut]. 

Type: Holotype d% macropterous; Laut Island, Macassar Strait, near 
Borneo; Drake Coll. (USNM). 

Distribution: Laut Island. 

Host Plant: Unrecorded. 

*Physatocheila lenis Drake and Poor 

Physatochelia lenis Drake and Poor 1939c, p. 204 [India]. 

Type: Holotype 9 3 macropterous; Chakrata, United Provinces; Drake 
CoU. (USNM). 

Distribution: India. 

Host Plant: Unrecorded. 

*Physatocheila major Osborn and Drake 

Physatochelia major Osborn and Drake 1917a, pp. 155, 158, fig. Id 
[111.].— McAtee 1923, p. 145 [Md.; Va.; D.C.].— Blatchley 1926, 
p. 484.— Froeschner 1944, p. 669.— Hurd 1946, p. 452 [Ind.]. 
Type: Holotype 9, macropterous; Urbana, Illinois, U.S.; Drake Coll. 
(USNM). 
Distribution: U.S. (111.; Ind.; Md.; Va.; D.C). 
Host Plant: Unrecorded. 
*Physatocheila marginata (Distant) 

Teleonemia marginata Distant 1909a, p. 121 [India]; 1910a, p. 123, 

fig. 64. 
Physatochila marginata: Bergroth 1921, p. 105. — Drake and Poor 1937b, 

p. 399 [Luzon]. 
Cysteochlla (Parada) marginata: Horvath 1925a, p. 3. 
Type: Sex unknown; Margherita, Assam; British Mus. 
Distribution: India; Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

*PhysatocheiIa muluana Drake 

Physatochelia muluana Drake 1954g, p. 661, fig. 3 [Ethiopia]. 
Type: Holotype 9; Mulu, Ethiopia; British Mus. 
Distribution: Ethiopia. 
Host Plant: Unrecorded. 



336 U.S. NATIONAL MUSEUM BULLETIN 243 

Physatocheila municeps Horvath 

Physatocheila municeps Horvath 1903a, p. 78 [Algeria]; 1906a, p. 95. 
Type: Holotype cf; Oued-Harris, Algeria; deposition unknown. 
Distribution: Algeria. 
Host Plant: Unrecorded. 

*Physatocheila objicis Drake 

Physatocheila objicis Drake 1942a, p. 10 [Victoria]; 1961b, p. 108 [New 
South Wales; Queensland]. 
Type: Holotype o 71 , macropterous; Kista, Victoria, Australia; Drake 
Coll. (USNM). 
Distribution: Australia (Victoria; Queensland; New South Wales). 
Host Plant: Unrecorded. 

*Physatocheila orientis Drake 

Physatocheila orientis Drake 1942a, p. 9 [Japan]. — Takeya 1953a, p. 2 

[Manchuria]; 1953d, p. 173, pi. 6, fig. 2. 
Physatochila orientis: Takeya 1951a, p. 20. 
Type: Holotype 9, macropterous; Sapporo, Hokkaido, Japan; Drake 
Coll. (USNM). 
Distribution: Japan; China (Manchuria). 
Host Plant: Unrecorded. 

*Physatocheila plexa (Say) 

Tingis plexus Say 1832, p. 27.— Fitch 1857, p. 794.— Le Conte 1859, 

p. 349. 
Physatochila plexa: Stal 1873, p. 129 [111.].— Provancher 1886, p. 160 
[Canada].— Uhler 1886, p. 22.— Winner 1904, p. 202 [Pa.].— Van 
Duzee 1905, p. 549 [N.Y.]; 1912, p. 323.— Torre-Bueno 1910, 
p. 31.— Parshley 1914, p. 145 [Maine]. 
Physatocheila plexa: Smith 1910, p. 149 [N.J.].— Parshley 1916a, p. 165 
[R.I.]; 1917b, p. 56 [Mass.]; 1923b, p. 705, fig. 164 [Conn.].— Van 
Duzee 1917b, p. 219 [Ind.; W. Va.; Mich.; Que.; Ont.].— Osborn 
and Drake 1917a, p. 156 [Nebr.; Kans.; Iowa; Wis.]. — Barber 
1922a, p. 17 [willow].— McAtee 1923, p. 145 [Kalmia latifolia].— 
Torre-Bueno 1926a, p. 54; 1929, p. 311.— Blatchley 1926, p. 484, 
pi. 4, fig. 4 [Va.; D.C.; oak; hickory].— Drake 1928b, p. 102 — 
Froeschner 1944, p. 669.— Proctor 1946, p. 75.— Bailey 1951, p. 25 
[N.H.]. 
Physatocheila parshleyi Osborn and Drake 1917a, pp. 155, 156, figs. 
1 a-c (in part). 
Type : Presumably lost. 

Distribution: U.S. (Maine; N.H.; Mass.; R.I.; Conn.; N.Y.; N.J.; 
Pa.; Md.; D.C.; Va.; W. Va.; Ind.; 111.; Mich.; Wis.; Nebr.; Iowa; Kans.); 
Canada (Ont.; Que.; N.S.). 
Host Plants: Kalmia latifolia; hickory; willow; oak. 



LACEBUGS OF THE WORLD 337 

*Physatocheila pulchella Lindberg 

Physatochila pulchella Lindberg 1932a, p. 43, pi. 2, fig. 6 [Morocco]. 
Type: Sex unknown; Morocco; Helsin. Mus. 
Distribution: Morocco. 
Host Plant: Unrecorded. 

*Physatocheila ruris Drake 

Physatocheila ruris Drake 1942a, p. 11 [China]. 

Type: Holotype 9, macropterous; Krvangtung, Kan-lin San, South 
China; Drake Coll. (USNM). 

Distribution: China. 

Host Plant: Unrecorded. 

*PhysatocheiIa scotti Drake 

Physatocheila scotti Drake 1954g, p. 660, fig. 2 [Ethiopia]. 
Type: Holotype o"; Doukam, Ethiopia; British Mus. 
Distribution : Ethiopia. 
Host Plant: Unrecorded. 

*Physatocheila senegalensis Drake 

Physatocheila senegalensis Drake 1954a, p. 234 [Senegal]. 
Type: Holotype o"; Bambery, Senegal; Inst. Fr. Afr. N. 
Distribution: Senegal. 
Host Plant: Unrecorded. 

*Physatocheila stricta Bergevin 

Physatocheila stricta Bergevin 1929, p. 114, fig. 1 [Algeria]. 
Type: Holotype 9; Tamanrasset, Algeria; Paris Mus. 
Distribution: Algeria. 
Host Plant: Unrecorded. 

♦Physatocheila suttoni Drake 

Physatochila irregularis (not Montrouzier) Hacker 1929, p. 328, pi. 33, 

fig. 6 [Queensland]. 
Physatocheila suttoni Drake 1947c, p. 121. 
Type: Sex unknown; Stanthorpe, Queensland, Australia; Queensland 
Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

♦Physatocheila thomasi Drake 

Physatocheila thomasi Drake 1956b, p. Ill [Kenya]. 
Type: Holotype o 71 , macropterous; Limuru, Kenya Colony; Drake 
Coll. (USNM). 
Distribution: Kenya. 
Host Plant: Unrecorded. 



338 U.S. NATIONAL MUSEUM BULLETIN 243 

*Physatocheila uniseriata Hacker 

Physatochila uniseriata Hacker 1929, p. 327, pi. 33, fig. 7 [South Australia; 
Western Australia; New South Wales]. 
Type: Sex unknown; Mount Lofty Ranges, South Australia; So. Austr. 
Mus. 

Distribution: Australia (South Australia; Western Australia; New 
South Wales). 

Host Plant: Unrecorded. 
*Physatocheila variegata Parshley 

Physatocheila variegata Parshley 1916a, p. 166 [N.Y.; W.Va.; Mass., 
Conn.; pine]; 1917b, p. 56 [Maine]; 1919a, p. 24 [B.C.]; 1923b, 
p. 705. — Barber 1922a, p. 17 [N.J.; hickory; willow]. — Downes 
1925, p. 18, fig. 9 [Salix lasiandra].— Blatchley 1926, p. 485 [Ind.].— 
Froeschner 1944, p. 669 [Mo.].— Hurd 1946, p. 452 [111.].— Proctor 
1946, p. 75.— Bailey 1951, p. 26 [N.H.; Va.]. 
Physatochila plexa (not Say): Osborn and Drake 1916a, p. 242 [Iowa; 

Mich.; Ont.]. — Downes 1927a, p. 10 [alder; cottonwood]. 
Physatocheila plexa var. variegata: Drake 1928b, p. 102. 
Type: Holotype o 71 ; Gowanda, New York, U.S.; deposition unknown. 
Distribution: U.S. (Maine; N.H.; Mass.; Conn.; N.Y.; N.J.; Va.; 
W. Va.; 111.; Ind.; Mo.; Iowa; Mich.); Canada (Ont.). 

Host Plants: Salix lasiandra; hickory; pine; cottonwood; alder. 
*Physatocheila variegata var. ornata Van Duzee 

Physatochila plexa (not Say): Van Duzee 1914, p. 11. 

Physatocheila ornata Van Duzee 1917a, p. 259 [Calif.]. 

Physatocheila par shleyi Osborn and Drake 1917a, p. 156 (in part) [Oreg.]. 

Physatocheila plexa: Osborn and Drake 1917a, p. 156 (in part) [Idaho]. 

Physatocheila variegata var. ornata: Hurd 1946, p. 452. 
Type: Holotype 9; Lake County, California, U.S.; Cal. Acad. 
Distribution: U.S. (Calif.; Oreg.; Idaho). 
Host Plant: Unrecorded. 
*Physatocheila veteris Drake 

Physatocheila veteris Drake 1942a, p. 10 [Japan]. 
Physatochila veteris: Takeya 1951a, p. 20. 
Type: Holotype o 71 , macropterous; Sapporo, Hokkaido, Japan; Drake 
Coll. (USNM). 
Distribution: Japan. 
Host Plant: Unrecorded. 
*Physatocheila weenenana Drake 

Physatocheila weenenana Drake 1953d, p. 212 [Natal; Kenya; Cape 
Province]. 
Type: Holotype d"; Weenen, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal; Cape Province); Kenya. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 339 

Physatocheila witmeri Mancini 

Physatochila dumetorum subsp. witmeri Mancini 1960a, p. 105 [Lebanon]. 
Physatocheila witmeri: Stichel 1960a, p. 332. 
Type: Sex unknown; Hasrun, Lebanon; Museo Civico Storia Naturale 
Milano, Milan, Italy. 
Distribution: Lebanon. 
Host Plant: Unrecorded. 

Genus PLACOTINGIS Drake 

Placotingis Drake 1960, p. 369. 
Type Species: Placotingis merga Drake. 

Placotingis merga Drake 

Placotingis merga Drake 1960, p. 370, fig. 21 [New Guinea]. 
Type: Holotype cf ; Mt. Wilhelm, Northeast New Guinea; Bishop Mus. 
Distribution: New Guinea (Northeast). 
Host Plant: Unrecorded. 

Placotingis par Drake 

Placotingis par Drake 1960, p. 370, fig. 22 [New Guinea]. 

Type: Holotype cf; Wisselmeren, Entarotadi, Netherlands New Guinea; 
Bishop Mus. 

Distribution: New Guinea (Netherlands). 

Host Plant: Unrecorded. 

Genus PLANIBYRSA Drake and Poor 

Planibyrsa Drake and Poor 1937c, p. 164.— Monte 1939b, p. 77; 1941e, 
p. 127.— Drake and Runoff 1960a, p. 78. 
Type Species: Leptobyrsa splendida Drake. 

*Planibyrsa elegantula (Drake) 

Leptobyrsa elegantula Drake 1922c, p. 373, fig. 2b [Brazil]; 1931b, p. 511 
[Peru]; 1936b, p. 700 [Argentina].— Monte 1937a, p. 32, fig. 11 
[Bignoniaceae]. 
Planibyrsa elegantula: Drake and Poor 1937c, p. 165.- — Drake and 
Hambleton 1938a, p. 65.— Kormilev 1955a, p. 66 [Bolivia]. 
Type: Holotype o 71 ; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil; Argentina; Peru; Bolivia. 
Host Plant: Bignoniaceae. 
*Planibyrsa montei Drake and Hambleton Plate 40 

Leptobyrsa splendida (not Drake): Monte 1937a, p. 32, fig. 2 [Brazil; 

Serjania], 
Planibyrsa montei Drake and Hambleton 1938a, p. 65. — Monte 1938b, 
p. 129.— Singh 1953, p. 119. 



340 U.S. NATIONAL MUSEUM BULLETIN 243 

Type: Holotype cf ; Belo Horizonte, Minas Gerais, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Serjania sp. 

* Planibyrsa sodalis (Drake and Bondar) 

Leptobyrsa sodalis Drake and Bondar 1932, p. 91 [Brazil; Arrabidaea]. — 

Costa Lima 1936, p. 128. 
Planibyrsa sodalis: Drake and Poor 1937c, p. 165. — Silva 1956, p. 46. 
Type: Holotype d% macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Arrabidaea sp. 

*Planibyrsa splendida (Drake) 

Leptobyrsa splendida Drake 1922c, p. 374, fig. 2a [Brazil]. 

Planibyrsa spendida [sic]: Drake and Poor 1937c, p. 164. 

Planibyrsa splendida: Monte 1940e, p. 287.— Silva 1956, p. 48. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

Genus PLATYTINGIS Drake 

Platytingis Drake 1925b, p. 107.— Drake and Runoff 1960a, p. 78. 
Type Species: Platytingis pediades Drake. 

*Platytingis pediades Drake 

Platytingis pediades Drake 1925b, p. 108, fig. [Malagasy]. 
Type: Holotype d\ macropterous; Imamonto, Madagascar; Drake 
Coll. (USNM). 
Distribution: Malagasy Republic; Tanganyika. 
Host Plant: Unrecorded. 

Genus PLEROCHILA Drake 

Plerochila Drake 1954c, p. 69.— Drake and Ruhoff 1960a, p. 78. 
Type Species: Teleonemia australis Distant. 

*Plerochila australis (Distant) 

Teleonemia australis Distant 1904, p. 432, pi. 8, fig. 13 [Cape Province; 

01 ea europea]. 
Cysteochila (Parada) australis: Horvath 1925a, p. 3. 
Plerochila australis: Drake 1954c, p. 69 [Mozambique; Transvaal]; 
1954g, p. 663 [Ethiopia].— Orian 1956, p. 647 [Mauritius]. 
Type: Sex unknown; Capetown, South Africa; British Mus. 
Distribution: South Africa (Cape Province; Transvaal); Mozambique; 
Ethiopia; Mascarene Islands (Mauritius). 
Host Plant: Olea europea. 



LACEBUGS OF THE WORLD 341 

*Plerochila horvathi (Schouteden) 

Cysteochila horvathi Schouteden 1907, p. 286 [Mauritius]. 
Cysteochila (Parada) horvathi: Horvath 1925a, p. 3. 

Plerochila horvathi: Drake 1954c, p. 69 [Kenya; Tanganyika]; 1957e, 
p. 401 [Reunion; Congo; South-West Africa]. — Orian 1956, p. 
647.— Mamet 1957, p. 57. 
Pleurochila [sic] horvathi: Drake and Mamet 1956, p. 301 [Olea europea; 
Jasminum], 
Type: Sex unknown; Mauritius Island; Cent. Afr. Mus. 
Distribution: Mascarene Islands (Reunion; Mauritius); Tanganyika; 
Kenya; Congo; South-West Africa. 
Host Plants: Jasminum sp.; Olea europea. 

*Plerochila rutshurica Schouteden 

Plerochila rutshurica Schouteden 1954, p. 140 [Congo; Ruanda; Olea]. 

Type: Unknown. 

Distribution: Congo; Ruanda-Urundi. 

Host Plant: Olea sp. 
*Plerochila zululandana Drake 

Plerochila (nom. nud.) zululandana Drake 1953d, p. 213 [Zululand]. 

Type: Holotype d 1 ; Eshowe, Zululand; British Mus. 

Distribution: South Africa (Zululand). 

Host Plant: Unrecorded. 

Genus PLESEOBYRSA Drake and Poor 

Pleseobyrsa Drake and Poor 1937c, p. 165.— Hurd 1946, p. 470.— Drake 
and Runoff 1960a, p. 79. 
Type Species: Pleseobyrsa boliviana Drake and Poor. 

*Pleseobyrsa ablusa Drake and Hambleton 

Pleseobyrsa ablusa Drake and Hambleton 1946b, p. 124 [Peru]. 
Type: Holotype o 71 , macropterous; Changos, Peru; Drake Coll. (USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 

*Pleseobyrsa atratarsis Drake and Hambleton 

Pleseobyrsa atratarsis Drake and Hambleton 1944b, p. 127, fig. 1 
[Brazil].— Drake and Davis 1960, fig. 49. 
Type: Holotype 9, macropterous; Pocas de Caldas, Minas Gerais, 
Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 



342 U.S. NATIONAL MUSEUM BULLETIN 243 

Pleseobyrsa bicincta Monte 

Pleseobyrsa bicincta Monte 1946b, p. 283 [Brazil]. 

Type: Holotype 9; Palmerias, Monte Alegre, Mato Grosso, Brazil; 
Monte Coll. (Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Pleseobyrsa boliviana Drake and Poor 

Pleseobyrsa boliviana Drake and Poor 1937c, p. 165 [Bolivia]. — Monte 
1938b, p. 129. 
Type: Holotype cf, macropterous; Cochabamba, Bolivia; Drake Coll. 
(USNM). 
Distribution : Bolivia. 
Host Plant: Avocado. 

*Pleseobyrsa chiriquensis (Champion) 

Leptobyrsa chiriquensis Champion 1897, p. 27, pi. 2, figs. 16, 16a 

[Panama]. 
Pleseobyrsa chiriquensis: Drake and Poor 1937c, p. 165. — Monte 1942a, 
p. 97 [Venezuela]; 1947b, p. 432 [Colombia; Persea americana], — 
Hurd 1946, p. 471 [Costa Rica]. 
Type: Sex unknown; Panama; British Mus. 
Distribution: Panama; Colombia; Venezuela; Costa Rica. 
Host Plant: Persea americana. 

Pleseobyrsa lichyi Monte 

Pleseobyrsa lichyi Monte 1945a, p. 251 [Venezuela]. 

Type: Holotype cf; Carreteras Maracay-Ocumare, Aragua, Venezuela; 
Monte Coll. (Mus. Nacional). 

Distribution: Venezuela. 

Host Plant: Unrecorded. 

Pleseobyrsa nigriceps (Champion) 

Leptobyrsa nigriceps Champion 1897, p. 27, pi. 2, fig. 17 [Panama; 

Guatemala]. 
Pleseobyrsa nigriceps: Drake and Poor 1937c, p. 165. 
Type: Sex unknown; Cerro Zunil, Guatemala; British Mus. 
Distribution: Panama; Guatemala. 
Host Plant: Unrecorded. 

*Pleseobyrsa peruana Drake 

Pleseobyrsa peruana Drake 1939d, p. 529 [Peru]. 
Type: Holotype <?, macropterous; Peru; Drake Coll. (USNM). 
Distribution: Peru. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 343 

*Pleseobyrsa plicata (Champion) 

Leptobyrsa plicata Champion 1897, p. 26, pi. 2, fig. 15 [Panama]. — 

Swezey 1945, p. 372 [Costa Rica]. 
Pleseobyrsa plicata: Drake and Poor 1937c, p. 165. — Monte 1940e, 

p. 286 [Venezuela]. 
Pleseobyrsa parana Drake and Hambleton 1944a, p. 95 [Brazil]. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Panama; Venezuela; Brazil; Costa Rica. 
Host Plant: Unrecorded. 

Note: Intercepted on Cattleya dowiana aurea from Costa Rica at port-of- 
entry, Washington, D.C. [Swezey]. 

Genus PLIOBYRSA Drake and Hambleton 

Pliobyrsa Drake and Hambleton 1946b, p. 123. — Drake and Ruhoff 
1960a, p. 79. 
Type Species: Leptopharsa inflexa Drake and Hambleton. 

*Pliobyrsa adversa (Drake and Hambleton) 

Leptopharsa adversa Drake and Hambleton 1938a, p. 60, fig. 10 [Brazil; 

Ocotea].— Monte 1941e, p. 108. 
Pleseobyrsa adversa: Monte 1940e, p. 286. 
Pliobyrsa adversa: Drake and Hambleton 1946b, p. 123. 
Type: Holotype d\ macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Ocotea sp. 

*Pliobyrsa inflexa (Drake and Hambleton) 

Leptopharsa inflexa Drake and Hambleton 1938b, p. 54, fig. 1 [Brazil; 

Ocotea lanata]. — Monte 1941 e, p. 113. 
Pliobyrsa inflata [sic]: Drake and Hambleton 1946b, p. 123. 
Type: Holotype cf, macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution : Brazil. 
Host Plant: Ocotea lanata. 

*Pliobyrsa lateris Drake and Hambleton 

Pliobyrsa lateris Drake and Hambleton 1946b, p. 124 [Guatemala]. 

Type: Holotype o", macropterous; Mazatenango, Guatemala; Drake Coll. 
(USNM). 

Distribution: Guatemala. 

Host Plant: Unrecorded. 



344 U.S. NATIONAL MUSEUM BULLETIN 243 

*Pliobyrsa mollinediae (Drake and Hambleton) 

Leptobyrsa mollinediae Drake and Hambleton 1934, p. 445 [Brazil; 

Mollinedia]. — Costa Lima 1936, p. 127. 
Pleseobyrsa mollinediae: Drake and Poor 1937c, p. 165. — Monte 1939b, 

p. 77; 1941e, p. 128. 
Pliobyrsa mollinediae: Drake and Hambleton 1946b, p. 123. — Drake and 
Davis 1960, fig. 63. 
Type: Holotype c? , macropterous; Vicosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Mollinedia sp. 

*Pliobyrsa pulcherrima Monte 

Pliobyrsa pulcherrima Monte 1947b, p. 430 [Brazil]. 

Type: Holotype o 71 ; Parque Nacional, Serra des Orgaos, Teresopolis, 
Estado do Rio, Brazil; Monte Coll. (Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

Pliobyrsa translucida (Champion) 

Leptobyrsa translucida Champion 1897, p. 26, pi. 2, figs. 14, 14a [Guate- 
mala]. 
Stephanitis translucida: Hurd 1946, p. 482. 
Pliobyrsa translucida: Drake and Hambleton 1946b, p. 123. 
Type: Holotype macropterous; sex unknown; San Geronimo, Guatemala; 
British Mus. 
Distribution: Guatemala. 
Host Plant: Unrecorded. 

Note: Champion states type apterous; his figure shows unique specimen 
with long elytra and no hind pair of wings. 

Genus POGONOSTYLA Drake 

Pogonostyla Drake 1953d, p. 221.— Drake and Ruhoff 1960a, p. 79. 
Type Species: Pogonostyla intonsa Drake. 
*Pogonostyla afra Drake and Smithers 

Pogonostyla afra Drake and Smithers 1958, p. 315 [Natal]. 
Type: Holotype d\ macropterous; Sweetwater, Natal, South Africa; 
Drake Coll. (USNM). 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 
Pogonostyla crispa Drake 

Pogonostyla crispa Drake 1958c, p. 327 [Malagasy]. 
Type: Holotype o 71 ; Perinet, Madagascar; Magadascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 345 

*Pogonostyla discrega Drake and Smithers 

Pogonostyla discrega Drake and Smithers 1958, p. 313, figs. 1 a, b [So. 
Rhodesia]. 
Type: Holotype o", macropterous; Penhalonga, Southern Rhodesia; 
Drake Coll. (USNM). 
Distribution: Southern Rhodesia. 
Host Plant: Unrecorded. 

*Pogonostyla herana Drake 

Pogonostyla herana Drake 1958c, p. 328 [Malagasy]. 
Type: Holotype cf; Tongobory, Madagascar; Madagascar Sci. Inst. 
Distribution: Malagasy Republic. 
Host Plant: Unrecorded. 
*Pogonostyla intonsa Drake 

Pogonostyla intonsa Drake 1953d, p. 222 [Natal]. 
Type: Holotype cf; Verulan, Natal, South Africa; British Mus. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 
*Pogonostyla natalicola (Distant) 

Leptostyla natalicola Distant 1902a, p. 240, pi. 15, fig. 15 [Natal]. 

Pogonostyla natalicola: Drake 1953d, p. 222. 
Type: Sex unknown; Natal, South Africa; British Mus. 
Distribution: South Africa (Natal). 
Host Plant: Unrecorded. 

Genus PONTANUS Distant 

Pontanus Distant 1902b, p. 354.— Drake 1956f, p. 425.— Drake and 

Ruhoff 1960a, p. 79. 
Teratochila Drake and Poor 1936b, p. 147.— Drake and Ruhoff 1960a, 
p. 84. 
Type Specdes: Monanthia gibbifera Walker. 
*Pontanus accedens (Drake) (emendation) 

Teratocheila [sic] accedentis Drake 1947c, p. 120 [Australia]. 
T"Pe: Holotype 9, macropterous; Aldgate, Australia; Drake Coll. 
(USNM). 
Distribution: Australia. 
Host Plant: Unrecorded. 
* Pontanus cafer (Distant) 

Phyllontochila cafer Distant 1902a, p. 241, pi. 15, fig. 17 [Cape Province]. 
Tingis cafer: Blote 1945, p. 86. 
Pontanus cafer: Drake 1956f, p. 426. 
Type: Sex unknown; Cape Colony, South Africa; So. Afr. Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

681-552—64 23 



346 U.S. NATIONAL MUSEUM BULLETIN 243 

Pontanus gibbiferus (Walker) 

Monanthia gibbifera Walker 1873a, p. 197 [Australia]. — Lethierry and 

Severin 1896, p. 25. 
Pontanus gibbiferus: Distant 1902b, p. 354. 
Type: Sex unknown; Australia; British Mus. 
Distribution : Australia. 
Host Plant: Unrecorded. 

*Pontanus puerilis (Drake and Poor) 

Teratochila puerilis Drake and Poor 1936b, p. 147 [India; teak]. — Singh 
1953, p. 119. 
Type: Holotype 9, macropterous; Rahatgaon, Hoshangabad, Central 
Provinces; Drake Coll. (USNM). 
Distribution: India. 
Host Plant: Teak. 

Genus PSEUDACYSTA Blatchley 

Pseudacysta Blatchley 1926, p. 497.— Hurd 1946, p. 459.— Drake and 
Ruhoff 1960a, p. 80. 
Type Species: Acysta perseae Heidemann. 

*Pseudacysta perseae (Heidemann) 

Acysta perseae Heidemann 1908, p. 103, pi. 4, figs, a-c [Fla.; La.; Persea 

carolinensis; Persea gratissima; Camphor a officinalis]. — Barber 1914, p. 

507.— Van Duzee 1917b, p. 223.— Moznette 1922, p. 17, figs. 12, 13. 

Pseudacysta persea [sic]: Blatchley 1926, p. 497, fig. 120. — Gibson and 

Carrillo 1959, p. 19. 
Pseudacysta perseae: Hurd 1946, p. 459 [Tex.; Mexico]. 
Type: Holotype cf, macropterous; Miami, Florida, U.S.; USNM. 
Distribution: U.S. (Fla.; La.; Tex.); Mexico. 

Host Plants: Comphora officinalis; Persea carolinensis; Persea gratissima. 
Note: Life history study [Heidemann]. 

Genus PSILOBYRSA Drake and Hambleton 

Psilobyrsa Drake and Hambleton 1935, p. 148. — Drake and Ruhoff 

1960a, p. 80. 
Psylobyrsa [sic]: Monte 1941e, p. 129. 
Type Species: Psilobyrsa aechemeae Drake and Hambleton. 
*Psilobyrsa aechemeae Drake and Hambleton 

Psilobyrsa aechemeae Drake and Hambleton 1935, p. 149 [Brazil; Aechmea]. 
Type: Holotype c? , macropterous; S. Vicente, Sao Paulo, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Aechmea sp. 



LACEBUGS OF THE WORLD 347 

*Psilobyrsa vriesiae Drake and Hambleton 

Psilobyrsa vriesiae Drake and Hambleton 1935, p. 149, fig. 4 [Brazil; 
Vriesia hoehneana]. 
Type: Holotype cf, macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Vriesia hoehneana. 

Genus RADINACANTHA Hacker 

Radinacantha Hacker 1929, p. 330.— Drake and Runoff 1960a, p. 80. 
Type Species: Radinacantha reticulata Hacker. 

*Radinacantha prudens Drake and Poor (emendation) 

Radinacantha prudentis Drake and Poor 1937a, p. 15 [Malagasy]. 

Type: Sex unknown (abdomen broken); Ambalamadakana, Madagas- 
car; Drake Coll. (USNM). 

Distribution: Malagasy Republic. 

Host Plant: Unrecorded. 

*Radinacantha reticulata Hacker 

Radinacantha reticulata Hacker 1929, p. 330, pi. 34, fig. 11 [Queensland; 
Bribie] . 
Type: Sex unknown; Samsonvale, Queensland, Australia; Queensland 
Mus. 
Distribution: Australia (Queensland; Bribie Island). 
Host Plant: Unrecorded. 

*Radinacantha tasmanica Hacker 

Radinacantha tasmanica Hacker 1929, p. 331, pi. 34, fig. 12 [Tasmania]* 
Type: Sex unknown; Strahan, Tasmania, Australia; So. Austr. Mus- 
Distribution : Australia (Tasmania; New South Wales). 
Host Plant: Unrecorded. 

Genus RECAREDUS Distant 

Recaredus Distant 1909b, p. 361; 1910a, p. 104.— Drake 1950a, p. 166. — 
Drake and Ruhoff 1960a, p. 29. 
Type Species: Recaredus rex Distant. 

*Recaredus rex Distant 

Recaredus rex Distant 1909b, p. 361 [West Bengal] ; 1910a, p. 104, fig. 51. 

Type: Sex unknown; Paresnath, West Bengal, India; Indian Mus. 

Distribution: India. 

Host Plant: Unrecorded. 

Note: Taken at light. This genus belongs to subfamily Tinginae, 
tribe Tingini, not Cantacaderinae, and is here so transferred. 



348 U.S. NATIONAL MUSEUM BULLETIN 243 

Genus RENAUDEA Drake 

Renaudea Drake 1958c, p. 332.— Drake and Ruhoff 1960a, p. 80. 
Type Species: Renaudea pauliani Drake. 
Renaudea pauliani Drake 

Renaudea pauliani Drake 1958c, p. 333, fig. 2 [Comoro]. 
Type: Holotype 9; Fomboni, Moheli, lies Comores; Madagascar Sci. 
Inst. 
Distribution: Comoro Islands (Moheli). 
Host Plant: Unrecorded. 

Genus SABESTENA Drake 

Sabestena Drake 1944a, p. 67. — Drake and Ruhoff 1960a, p. 25. 
Type Species: Sabestena africana Drake. 
*Sabestena africana Drake 

Sabestena africana Drake 1944a, p. 67, fig. 6 [Kenya]; 1956a, p. 6 
[Congo; Ethiopia]. — Schouteden 1957c, p. 312. 
Type: Holotype 9, macropterous; Naburu, Kenya; Drake Coll. 
(USNM). 
Distribution: Kenya; Congo; Ethiopia. 
Host Plant: Unrecorded. 
*Sabestena alfierii Drake and Ruhoff 

Ceratinoderma sp. Priesner and Alfieri 1953, p. 66 [Egypt; Gymnocarpos 

decandrum]. 
Sabestena alfierii Drake and Ruhoff 1961b, p. 166. 
Type: Holotype 9, macropterous; Wadi Nouega, Egypt; Drake Coll. 
(USNM). 
Distribution: Egypt. 
Host Plant: Gymnocarpos decandrum. 

Genus SANAZARIUS Distant 

Sanazarius Distant 1904, p. 431.— Drake and Ruhoff 1960a, p. 81. 
Type Species: Sanazarius cuneatus Distant. 
*Sanazarius biseriatus Drake 

Sanazarius biseriatus Drake 1953d, p. 217 [Cape Province]. 
Type: Holotype d\ brachypterous; Swellendam, Cape Province, South 
Africa; British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 
*Sanazarius cuneatus Distant 

Sanazarius cuneatus Distant 1904, p. 431, pi. 8, fig. 11 [Cape Province]. 
Type: Sex unknown; Grahamstown, Cape Colony, South Africa; 
British Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 349 

*Sanazarius productus Distant 

Sanazarius productus Distant 1911a, p. 43, fig. [Cape Province]. — Drake 
1956f, p. 426 [Protea]. 
Type: Sex unknown; Grahamstown, Cape Colony, South Africa; So. 

Afr. Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Protea sp. 

Note: Figure p. 44 incorrectly labeled by Distant as "Sanazarius inflatus" 
instead of "S. productus.'''' 

Genus SCYMNOTINGIS Drake 

Scymnotingis Drake 1960, p. 354. 
Type Species: Scymnotingis sigillata Drake. 

Scymnotingis sigillata Drake 

Scymnotingis sigillata Drake 1960, p. 355, fig. 10 [New Guinea]. 
Type: Holotype 9; Daulo Pass, Asaro-Chimbu Divide, Northeast New 
Guinea; Bishop Mus. 
Distribution: New Guinea (Northeast). 
Host Plant: Unrecorded. 

Genus SINUESSA Horvath 

Phyllontochila [sic] (Sinuessa) Horvath 1910, p. 63. 
Phyllontocheila (Sinuessa): Horvath 1911a, pp. 329, 332 (key). 
Sinuessa: Drake 1957c, p. 32.— Drake and Ruhoff 1960a, p. 81. 
Type Species: Phyllontochila (Sinuessa) subinermis Horvath. 

*Sinuessa colens (Drake) (emendation) 

Belenus colentis Drake 1953d, p. 218 [Cape Province]. 
Sinuessa calentis [sic]: Drake 1957c, p. 32. 
Type: Holotype cf; Queenstown, Cape Province, South Africa; British 
Mus. 
Distribution: South Africa (Cape Province). 
Host Plant: Unrecorded. 

*Sinuessa nairobia Drake 

Sinuessa nairobia Drake 1957c, p. 34 [Kenya; Transvaal; Congo]. 

Type: Holotype 9, macropterous; Nairobi, Kenya Colony; Drake Coll. 
(USNM). 

Distribution: Kenya; Congo; South Africa (Transvaal). 

Host Plant: Unrecorded. 



350 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Sinuessa subinermis (Horvath) 

Phyllontochila {Sinuessa) subinermis Horvath 1910, p. 64 [Tanganyika]. 
Phyllontocheila (Sinuessa) subinermis: Horvath 1911a, pp. 329, 332. 
Phyllontocheila subinermis: Schouteden 1916a, p. 272 [Eritrea]. — Mancini 
1939b, p. 162; 1954a, p. 143; 1956b, p. 78.— Drake 1954g, p. 663 
[Ethiopia]; 1957a, p. 416 [Aden]. 
Sinuessa subinermis: Drake 1957c, p. 32. 

Ammianus (Sinuessa) subinermis: Drake 1958a, p. 106 [Angola; Uganda; 
Kenya; So. Rhodesia]. 
Type: Sex unknown; Kibongoto, Kilimandjaro; Stockholm Mus. 
Distribution: Tanganyika; Kenya; Southern Rhodesia; Uganda; Angola; 
Aden Protectorate; Eritrea; Ethiopia. 
Host Plant: Unrecorded. 

*Sinuessa waelbroecki (Schouteden) 

Phyllontocheila waelbroecki Schouteden (in Bergroth and Schouteden 
1905, p. 381, fig. 1) [Congo]; 1916b, p. 292; 1923, p. 102.— Drake 
1956f, p. 424 [Transvaal]. 
Phyllontochila (Sinuessa) waelbroecki: Horvath 1910, p. 63. 
Phyllontocheila (Sinuessa) waelbroecki: Horvath 1911b, pp. 329, 332. 
Sinuessa waelbroecki: Drake 1957c, p. 32; 1958b, p. 28 [Kenya]. 
Ammianus (Sinuessa) waelbroecki: Drake 1958a, p. 106 [Angola]. 
Type: Sex unknown; Kinchassa, Belgian Congo; Cent. Afr. Mus. 
Distribution: Congo; Kenya; Angola; South Africa (Transvaal). 
Host Plant: Unrecorded. 

Genus SPHAERISTA Kiritshenko 

Sphaerista Kiritshenko 1951, pp. 240, 245.— Drake and Ruhoff 1960a, 
p. 81. 
Type Species: Orthostira paradoxa Jakovlev. 
*Sphaerista paradoxa (Jakovlev) 

Orthostira paradoxa Jakovlev 1880a, p. 128 [Russia]; 1893, p. 293. — 

Lethierry and Severin 1896, p. 7. 
Acalypta paradoxa: Horvath 1906a, p. 34 [Siberia]. — Oshanin 1908, 

p. 412; 1912, p. 42. 
Sphaerista paradoxa: Kiritshenko 1951, p. 245, fig. 324. 
Type: Holotype d% brachypterous; Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Siberia). 
Host Plant: Unrecorded. 

Genus SPHAEROCYSTA Stal 

Sphaerocysta Stal 1873, pp. 120, 128.— Lethierry and Severin 1896, 
p. 15.— Drake 1928a, p. 42.— Monte 1939b, p. 78; 1941e, p. 129. — 
Drake and Ruhoff 1960a, p. 82. 
Type Species: Tingis globifera Stal. 



LACEBUGS OF THE WORLD 351 

Sphaerocysta angulata Monte 

Sphaerocysta angulata Monte 1941a, p. 373, fig. 1 [Brazil; Bignoniaceae]. 

Type: Holotype cf ; Chavantes, Sao Paulo, Brazil; Monte Coll. (Mus. 
Nacional). 

Distribution: Brazil. 

Host Plant: Bignoniaceae. 

*Sphaerocysta biseriata Drake 

Sphaerocysta inflata var. biseriata Drake 1928a, p. 43 [Brazil]. 

Sphaerocysta biseriata: Drake and Hambleton 1934, p. 438. 
Type: Holotype o" ; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Sphaerocysta brasiliensis Monte 

Sphaerocysta brasiliensis Monte 1938c, p. 64, fig. [Brazil]; 1939a, p. 516 
fig. [redescribed] . 
Type: Holotype c? ; Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Sphaerocysta egregia Drake 

Sphaerocysta egregia Drake 1928a, p. 44 [Brazil]. 
Type: Holotype cf ; Corumba, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Sphaerocysta fumosa Drake 

Sphaerocysta fumosa Drake 1928a, p. 44 [Brazil]. 
Type: Holotype 9, macropterous; Para, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

♦Sphaerocysta globifera (Stal) 

Tingis ? globifera Stal 1858, p. 65 [Brazil].— Walker 1873a, p. 181. 
Sphaerocysta globifera: Stal 1873, p. 128.— Drake 1929, p. 35; 1930b, 
p. 25; 1935, p. 11.— Drake and Hambleton 1934, p. 438 
[Bignoniaceae]. — Costa Lima 1936, p. 130. — Drake and Poor 1937d, 
p. 305.— Silva 1956, p. 48, figs. 18, 18a. 
Sphaerocysta globiffera [sic]: Drake 1928a, p. 43. 
Type: Sex unknown; Brazil; Stockholm Mus. 
Distribution: Brazil; Paraguay. 
Host Plants: Bignoniaceae; hollyhocks. 



352 U.S. NATIONAL MUSEUM BULLETIN 243 

*Sphaerocysta inflata (Stal) 

Tingis? inflata Stal 1858, p. 64 [Brazil].— Walker 1873a, p. 181. 
Sphaerocysta inflata: Stal 1873, p. 128.— Drake 1930a, p. 1; 1935, p. 11 
[Paraguay]; 1936b, p. 699 [Argentina]. — Drake and Hambleton 
1934, p. 438 [Bignoniaceae]. — Costa Lima 1936, p. 130. — Drake and 
Poor 1937d, p. 305, pi. 36, fig. 5; 1939b, p. 96.— Monte 1938b, p. 
131; 1939b, p. 78 [Bolivia].— Kormilev 1955a, p. 67.— Drake and 
Davis 1960, fig. 69. 
Type: Holotype 9; Brazil; Stockholm Mus. 
Distribution: Brazil; Argentina; Paraguay; Bolivia. 
Host Plant: Bignoniaceae. 
Sphaerocysta maculata Monte 

Sphaerocysta maculata Monte 1942a, p. 95, fig. 2 [Brazil]. 
Type: Holotype cf; Palmeiras, Impaussu, Sao Paulo, Brazil; Monte 
Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
* Sphaerocysta nosella Drake and Hambleton 

Sphaerocysta nosella Drake and Hambleton 1945, p. 358 [Ecuador]. 
Type: Holotype cf, macropterous; Vinces, Ecuador; Drake Coll. 
(USNM). 
Distribution: Ecuador. 
Host Plant: Unrecorded. 
*Sphaerocysta paris Drake 

Sphaerocysta paris Drake 1939d, p. 529 [Brazil]. — Silva 1956, p. 48. — 

Drake and Ruhoff 1961c, p. 145. 
Sphaerocysta romani Drake 1947, p. 2. 
Type: Holotype 9; Bahia, Brazil; Hungarian Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Sphaerocysta propria Drake and Poor 

Sphaerocysta propria Drake and Poor 1939a, p. 31 [Brazil]. 
Type: Holotype 9, macropterous; Rio Grande do Sul, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Sphaerocysta stali Drake 

Sphaerocysta globifera (not Stal): Champion 1898b, pi. 2, figs. 11, 11a 

(fig. only). 
Sphaerocysta stali Drake 1928a, p. 43 [Brazil]. — Drake and Bondar 
1932, p. 87.— Silva 1956, p. 51. 
Type: Holotype cf; Rio de Janeiro, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 353 

Genus STENOCYSTA Champion 

Stenocysta Champion 1897, p. 28. — Hurd 1946, p. 473. — Drake and 
Ruhoff 1960a, p. 82. 
Type Species: Stenocysta pilosa Champion. 

Stenocysta pilosa Champion 

Stenocysta pilosa Champion 1897, p. 29, pi. 2, figs. 18, 18 a, b [PanamaJ. 
Type: Holotype 9; Bugaba, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

Genus STEPHANITIS Stal 

Stephanitis Stal 1873, pp. 119, 123; 1874, p. 53— Horvath 1906a, 
pp. 14, 54 (key); 1912b, pp. 319-339 (key).— Oshanin 1908, p. 424; 
1912, p. 43.— Van Duzee 1917b, p. 216.— Stichel 1926, p. 110; 
1935, p. 349; 1938a, p. 406; 1960a, p. 300; 1960b, p. 394; 1960c, 
p. 133.— de Seabre 1931, pp. 407, 419.— Borner 1935, pp. 73, 76 
(key).— Guide 1938, pp. 243, 277 (key).— China 1943, p. 246 — 
Blote 1945, p. 83.— Hurd 1946, p. 48 1 .—Gonzalez 1948, p. 50 — 
Drake 1948b, pp. 45-56 (catalog).— Kiritshenko 1951, pp. 242, 
247.— Takeya 1951a, p. 7.— Drake and Maa 1953, pp. 99-101 
(catalog). — Gomez-Menor 1954, p. 369. — Drake and Ruhoff 1960a, 
p. 82. 

Tingis (not Fabricius): Laporte 1833, p. 48. — Drake and Ruhoff 
1960a, p. 86. 

Tingis (not Fabricius): Lethierry and Severin 1896, p. 12. — Drake 
and Ruhoff 1960a, p. 86. 

Cadamustus Distant 1903a, p. 47; 1903b, p. 132; 1910a, p. 108.— Drake 
and Ruhoff 1960a, p. 41. 

Maecenas Kirkaldy 1904, p. 280.— Drake and Ruhoff 1960a, p. 67. 

Calliphanes Horvath 1906b, p. 34.— Drake and Ruhoff 1960a, p. 41. 

Mokanna Distant 1910a, p. 111. — Drake and Ruhoff 1960a, p. 68. 
Type Species: Acanthia pyri Fabricius. 

Subgenus STEPHANITIS (STEPHANITIS) Stal 

Type Species: Acanthia pyri Fabricius. 
*Stephanitis agaica Drake 

Stephanitis agaica Drake 1960, p. 374 [New Guinea]. 

Type: Holotype cT; Wisselmeren, Enarotadi, Netherlands New Guinea; 
Bishop Mus. 

Distribution: New Guinea (Northeast; Netherlands). 

Host Plant: Unrecorded. 



354 U.S. NATIONAL MUSEUM BULLETIN 243 

*Stephanitis ambigua Horvath 

Tingis pyrioides (not Scott): Matsumura 1905, p. 33. 
Stephanitis ambigua Horvath 1912b, pp. 321, 328, fig. 2 [Japan]. — 
Esaki and Takeya 1931, p. 54, figs, lc, 2c; pi. 4, fig. 3; pi. 5, fig. 3; 
pi. 6, fig. 3. — Takeya 1932, p. 8 [Korea; Lindera obtusiloba]; 1951a, 
p. 9 [Taiwan]; 1953d, p. 168 [Lindera glauca].— Drake 1938b, p. 197 
[China]. 
Type: Sex unknown; Japan; Hungarian Mus. 
Distribution: Japan; China; Korea; Taiwan. 
Host Plants: Lindera glauca; Lindera obtusiloba. 

Stephanitis amboinae Drake and Poor 

Stephanitis amboinae Drake and Poor 1941, p. 163 [Amboina]. 
Type: Holotype 9; Amboina Island; Cal. Acad. 
Distribution: Moluccas (Amboina Island). 
Host Plant: Unrecorded. 

*Stephanitis assamana Drake and Maa 

Stephanitis assamana Drake and Maa 1954, p. 117 [India]. 

Type: Holotype 9; Cha Che, Delai Valley, Mishmi Hills, Assam, India; 
British Mus. 

Distribution: India. 

Host Plant: Unrecorded. 

Stephanitis assamana subsp. eremnoa Drake and Ruhoff 

Stephanitis assamana subsp. marginata Drake and Maa 1954, p. 118 

[India]. 
Stephanitis assamana subsp. eremnoa Drake and Ruhoff 1960c, p. 36. 
Type: Holotype 9; Cha Che, Delai Valley, Mishmi Hills, Assam, India; 
British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 

*Stephanitis aucta Drake 

Stephanitis aucta Drake 1942a, p. 20 [Brazil], 
Type: Holotype 9, macropterous; Chapoda, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Stephanitis blatchleyi Drake 

Stephanitis blatchleyi Drake 1925c, p. 37 [Fla.]. 

Leptobyrsa blatchleyi: Blatchley 1926, p. 470. 
Type: Holotype o", macropterous; Dunedin, Florida, U.S.; Drake Coll. 
(USNM). 
Distribution: U.S. (Fla.). 
Host Plant: Unrecorded. 
Note: Wrongly transferred by Blatchley to genus Leptobyrsa. 



LACEBUGS OF THE WORLD 355 

*Stephanitis caucasica Kiritshenko 

Stephanitis caucasica Kiritshenko 1939, p. 10 (nom. nud.); 1951, p. 247, 
fig. 325 [Caucasus; Transcaucasus; Rhododendron]. — Stichel 1960a, p. 
304, fig. 107. 
Type: Sex unknown; Transcaucasia, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Caucasus; Transcaucasus). 
Host Plant: Rhododendron sp. 

*Stephanitis chlorophana (Fieber) 

Tingis chlorophana Fieber 1861, p. 129 [Portugal]. — Walker 1873a, 

p. 179.— Chicote 1880, p. 189 [Spain]. 
Stephanitis chlorophana: Horvath 1905b, p. 568; 1906a, p. 56; 1912b, p. 
333.— de Seabra 1931, p. 421, figs. 492 (2-6), 494 [Viburnum tinus].— 
Gomez-Menor 1954, p. 381, fig. 4.— Stichel 1960a, p. 303, fig. 104. 
Type: Portugal; sex and deposition of type unknown. 
Distribution: Portugal; Spain. 
Host Plant: Viburnum tinus. 
*Stephanitis colocasiae Horvath 

Stephanitis colocasiae Horvath 1912b, pp. 320, 326 [New Guinea; Colo- 
casia esculenta]. — Drake 1960, p. 372. 
Type: Sex unknown; Friedrich-Wilhelmshafen, New Guinea; Hungarian 
Mus. 
Distribution: New Guinea (Northeast). 
Host Plant: Colocasia esculenta. 
Stephanitis depressa Blote 

Stephanitis depressa Blote 1945, p. 83 [Sumatra]. 
Type: Sex unknown; Gunung Teleman, Sumatra; Leiden Zool. Inst. 
Distribution: Greater Sunda Islands (Sumatra). 
Host Plant: Unrecorded. 
*Stephanitis farameae Drake and Hambleton 

Stephanitis farameae Drake and Hambleton 1935, p. 151 [Brazil; Faramea 
montevidensis]. — Monte 1939b, p. 78; 1941e, p. 132. 
Type: Holotype cf, macropterous; Parque do Estado, Sao Paulo, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Faramea montevidensis. 
*Stephanitis fasciicarina Takeya 

Stephanitis fasciicarina Takeya 1931, p. 70, pi. 7, fig. 2; pi. 8, fig. 6 
[Japan; Machilus thunbergii; Cinnamomum camphora]; 1951a, p. 10; 
1953d, p. 168— Drake and Poor 1937b, p. 403. 
Stephanitis kyushuana Drake 1948b, p. 52. 
Type: Holotype cf ; Akama, Kyushu, Japan; Kyushu Univ. 
Distribution: Japan. 
Host Plants: Cinnamomum camphora; Machilus thunbergii. 



356 U.S. NATIONAL MUSEUM BULLETIN 2 43 

*Stephanitis gallarum Horvath 

Stephanitis gallarum Horvath 1906b, p. 33 [India; Machilus gamblei]; 
1912b, p. 330.— Distant 1910a, p. 108.— Drake and Poor 1936b, p. 
148.— Drake 1937a, p. 387; 1938b, p. 197 [China].— Singh 1953, p. 
119.— Drake and Maa 1953, p. 100. 
Stephanitis distinctissima Esaki and Takeya 1933, p. 1, figs. 1, 2 [Taiwan; 
Machilus pseudo-longifolia]. — Takeya 1951a, p. 9 [Machilus longifolia]. 
Type: Sex unknown; Kurseong, Bengal, India; Hungarian Mus. 
Distribution: India; China; Taiwan. 

Host Plants: Alnus nepalensis; Machilus gamblei; Machilus longifolia; 
Machilus pseudo-longifolia. 

*Stephanitis gressitti Drake 

Stephanitis gressitti Drake 1948b, p. 50 [China]. 
Type: Holotype 9, macropterous; Szechwan, north of Chengtu, West 
China; Drake Coll. (USNM). 
Distribution: China. 
Host Plant: Unrecorded. 
*Stephanitis hikosana Drake 

Stephanitis hikosana Drake 1948b, p. 53 [Japan]. — Takeya 1951a, p. 10 
[Rhododendron hymenanthes var. heptamerum]; 1953d, p. 171. 
Type: Holotype cf; Hikosan (Buzen), Kyushu, Japan; Kyushu Univ. 
Distribution: Japan. 

Host Plant: Rhododendron hymenanthes var. heptamerum. 
*Stephanitis hydrangeae Drake and Maa 

Stephanitis hydrangeae Drake and Maa 1955, p. 8 [Taiwan; Hydrangea 
integra]. 
Type: Holotype cf, macropterous; Arisan, Formosa; Drake Coll. 
(USNM). 
Distribution: Taiwan. 
Host Plant: Hydrangea integra. 
*Stephanitis laudata Drake and Poor 

Stephanitis laudata Drake and Poor (in Drake and Maa 1953, p. 98) 
[China; Hainan; Taiwan]. 
Type: Holotype d\ macropterous; Ta Han, Hainan Island, China; 
Drake Coll. (USNM). 
Distribution: China (Hainan Island); Taiwan. 
Host Plant: Unrecorded. 
*Stephanitis ligyra Drake Figure la 

Stephanitis ligyra Drake 1960, p. 372, fig. 23 [New Guinea]. 
Type: Holotype 9; Okaitadi, Wisselmeren, Netherlands New Guinea; 
Bishop Mus. 
Distribution: New Guinea (Netherlands). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 357 

*Stephanitis luzonana Drake 

Stephanitis luzonana Drake 1948b, p. 50 [Luzon]. 
Type: Holotype cf, macropterous; Mt. Makiling, Luzon, Philippine 
Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

*Stephanitis matsumurae Horvath 

Stephanitis matsumurae Horvath 1912b, pp. 322, 332 [Japan], 
Type: Sex unknown; Sapporo, Japan; Hungarian Mus. 
Distribution: Japan. 
Host Plant: Unrecorded. 

Stephanitis mitrata (Stal) 

Tingis mitrata Stal 1858, p. 64 [Brazil].— Walker 1873a, p. 181.— 

Lethierry and Severin 1896, p. 12. 
Stephanitis mitrata: Stal 1873, p. 123. — Champion 1898b, p. 58, pi. 2, 
figs. 3, 3a. — Drake and Poor 1937d, p. 309. — -Drake and Hambleton 
1938a, p. 66 [Merostachys].— Monte 1941e, p. 133. 
Calliphanes mitratus: Horvath 1906b, p. 34. 
Type: Holotype 9; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Merostachys sp. 

*Stephanitis nashi Esaki and Takeya 

Tingis pyri (not Fabricius): Matsumura 1908, p. 147, fig. 

Stephanitis ambigua (not Horvath): Matsumura 1917, p. 440, fig.; 1930, 

p. 158, pi. 14, fig. 11. 
Stephanitis pyri (not Fabricius): Drake 1923a, p. 104. 
Stephanitis nashi Esaki and Takeya 1931, p. 54, pi. 4, fig. 1; pi. 5, fig. 1; 
pi. 6, fig. 1; figs, la, 2a [Japan; Korea; Pyrus; Prunus; Malus fioribunda; 
Chaenomeles cathayensis; Cormus tschonoskii; Kerria japonica; Crataegus; 
Malus].— Takeya 1932, p. 8; 1951a, p. 9 [Crataegus cuneata]; 1953d, 
p. 168.— Esaki 1932, p. 1635, fig.; 1954, p. 236, fig. 609.— Kato 
1933, pi. 31, fig. 3. — Saito 1933, p. 6 [Pyrus ussuriensis]. — Drake and 
Hsiung 1936, p. 288 [China] .—Drake and Poor 1937b, p. 403 — 
Drake 1937b, p. 594; 1938b, p. 197.— Fukui 1938, pi. 50, fig. 35 — 
Shinji 1938, p. 316.— Maa 1957, pp. 128, 133. 
Type: Holotype cf; Fukuoka, Kyushu, Japan; Kyushu Univ. 
Distribution: China; Japan; Korea; Taiwan. 

Host Plants: Chaenomeles cathayensis; Cormus tschonoskii; Crataegus cuneata; 
Crataegus sp.; Kerria japonica; Malus fioribunda; Malus sp.; Prunus sp.; Pyrus 
ussuriensis; Pyrus sp. 



358 U.S. NATIONAL MUSEUM BULLETIN 243 

Stephanitis nashi var. suigensis Saito 

Stephanitis nashi var. suigensis Saito 1933, p. 5 [Korea; Potentilla 
freyniana], — Takeya 1951a, p. 10. 
Type: Korea; sex and deposition of type unknown. 
Distribution: Korea. 
Host Plant: Potentilla freyniana. 

*Stephanitis oberti (Kolenati) 

Tingis pyri (not Fabricius): Fallen 1807, p. 39 [Sweden; Carduus lanceo- 
latus]; 1829, p. 149 [Myrica gale].— Sahlberg 1848, p. 136 [Finland]. 
Campylosteira (Dictyonota) oberti Kolenati 1856, p. 433 [Estonia]. 
Tingis oberti: Flor 1860, p. 369 [Pinus sylvestris]. — Jakovlev 1875b, p. 
262; 1880b, p. 102 (in part) [Caucasus]; 1902b, p. 276— Hiieber 
1893, p. 326 [Austria].— Lethierry and Severin 1896, p. 12. 
Stephanitis oberti: Stal 1873, p. 123. — Reuter 1882b, p. 118 [Myrtillus 
niger; Ledum palustre]. — Horvath 1906a, p. 56 [Siberia; Netherlands; 
Germany; Vaccinium myrtillus; Vaccinium vitis-idaea]; 1912b, pp. 322, 332 
[Rhododendron] .— Lindberg 1919, p. 42.— Sahlberg 1920, p. 83 [Calluna 
vulgaris; Myrtillus uliginosus]. — Stichel 1926, p. 110; 1928a, p. 206; 
1938a, p. 406 [Czechoslovakia; Albania; Norway; Denmark]; 1960a, 
p. 303 [Yugoslavia; Vaccinium uliginosum]. — Scholte 1935, p. 60. — 
Guide 1938, p. 280, fig.— Kiritshenko 1951, p. 248.— Cobben 1958a, 
p. 5, figs. 5-10. 
Type: Dorpat, Livonia; sex and deposition of type unknown. 
Distribution: Netherlands; Germany; Austria; Czechoslovakia; Albania; 
Yugoslavia; Denmark; Norway; Sweden; Finland; U.S.S.R. (Estonia; 
Caucasus; Siberia). 

Host Plants: Calluna vulgaris; Carduus lanceolatus; Ledum palustre; Myrica 
gale; Myrtillus niger; Myrtillus uliginosus; Pinus sylvestris; Rhododendron sp.; 
Vaccinium myrtillus; Vaccinium uliginosum; Vaccinium vitis-idaea. 

Stephanitis oberti var. biseriata Sahlberg 

Stephanitis oberti var. biseriata Sahlberg 1920, p. 83 [Finland]. — Stichel 
1960a, p. 303. 
Type: Sex unknown; Finland; Helsin. Mus. 
Distribution: Finland. 
Host Plant: Unrecorded. 

*Stephanitis olyrae Drake and Hambleton 

Stephanitis olyrae Drake and Hambleton 1935, p. 150, fig. 5 [Brazil; 
Olyra micrantha].— Monte 1939b, p. 78; 1941e, p. 133. 
Type: Holotype cf, macropterous; Ponte Nova, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Olyra micrantha. 



LACEBUGS OF THE WORLD 359 

*Stephanitis parana Drake and Hambleton 

Stephanitis parana Drake and Hambleton 1944b, p. 128 [Brazil]. 
Type: Holotype o", macropterous; Para, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Stephanitis princeps (Distant) 

Mokanna princeps Distant 1910a, p. 112, fig. 55 [India]. 
Stephanitis princeps: Horvath 1912b, pp. 321, 329. 
Type: Sex unknown; Trichinopoly, India; British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 
*Stephanitis propinqua Horvath 

Stephanitis propinqua Horvath 1912b, pp. 322, 331 [Japan]. — Takeya 
1932, p. 9, pi. 1, fig. 1 [Korea; Rhododendron]. 
Type: Sex unknown; Kagoshima, Japan; Hungarian Mus. 
Distribution: Japan; Korea. 
Host Plant: Rhododendron sp. 
*Stephanitis pyri (Fabricius) 

Acanthia pyri Fabricius 1775, p. 696 [Pyrus]; 1781, p. 338; 1787, p. 280; 

1794, p. 78.— Rossi 1790, p. 225.— Walckenaer 1802, p. 338. 
Cimex appendiceus Fourcroy 1785, p. 212.— Villers 1789, p. 488, pi. 3, 

%• 19. 
Cimex pyri: Gmelin 1790, p. 2127. 

Tingis pyri: Fabricius 1803, p. 126.— Latrielle 1804, p. 254; 1829, p. 
201.— Le Peletier and Serville 1828, p. 653.— Laporte 1833, p. 48.— 
Burmeister 1835, p. 259 [Germany; Portugal]. — Spinola 1837, 
p. 166.— Herrich-Schaeffer 1838, p. 69, pi. 126, fig.; pi. 130, fig. d; 
1850, p. 162.— Blanchard 1840, p. 112, pi. 2, fig. 7.— Westwood 
1840a, p. 478 [England; pear].— Amyot and Serville 1843, p. 297 
[France].— Guerin-Meneville 1844, p. 349, pi. 56, fig. 13.— Fieber 
1844, p. 102, pi. 8, figs. 34-36 [Czechoslovakia; Austria; Hungary; 
Italy; Sweden]; 1861, p. 129.— Costa 1847a, p. 21; 1847c, p. 161.— 
Mayr 1858, p. 572.— Frey-Gessner 1865, p. 233 [Switzerland].— 
Garbiglietti 1869, p. 275— Gredler 1870, p. 76.— Walker 1873a, 
p. 179.— Horvath 1874b, p. 432— Ferrari 1874, p. 171; 1878, p. 66; 
1885, p. 411.— Reiber and Puton 1876, p. 69.— Bolivar and Chicote 
1879, p. 166 [Spain].— Puton 1879c, p. 105; 1881, p. 123 [Syria].— 
Jakovlev 1880b, p. 102 [Caucasus].— Dubois 1888, p. 120.— d'Antes- 
santy 1890, p. 31.— Reuter 1891, p. 26 [Greece].— Hueber 1893, 
p. 325 [Sorbus aria; Vaccinium myrtillus]. — Champion 1898b, p. 58.— 
Reichensperger 1920, p. 61— Gautier 1925, p. 321; 1927a, p. 12; 
1927b, p. 26. 

Tingis marginata Lamarck 1816, p. 504. 

Tingis appendicea: Le Peletier and Serville 1828, p. 653. 



360 U.S. NATIONAL MUSEUM BULLETIN 243 

Stephanitis pyri: Stal 1873, p. 123— Horvath 1906a, pp. 54, 55 
[Turkestan; Armenia; Turkey; Pyrus malus; Castanea sativa]; 1906d, 
p. 1; 1912b, pp. 320, 327 [Prunus lusitanica]; 1916, p. 9 [Albania].— 
Muzik 1907, p. 57.— Blanchard 1926, p. 362.— Stichel 1926, p. 110 
[Pyrus aria]; 1938a, p. 406 [Belgium; Rumania]; 1960a, p. 301, 
fig. 98 [Sicily; Sardinia; Poland; Sorbus torminalis; Pyrus aucuparia; 
Prunus mahaleb; Prunus armeniaca; Cotoneaster integerrima; Crataegus 
crus-galli; Robina pseudoacacia; Ulmus effusa; Ulmus campestris; Populus; 
Tilia; Ligustrum; Cormus; Ribes nigrum; Juglans regia]. — Esaki and 
Takeya 1931, p. 53, figs, lb, 2b; pi. 4, fig. 2; pi. 5, fig. 2; pi. 6, 
fig. 2.— de Seabra 1931, p. 420, figs. 492(1), 493.— Silvestri 1934, 
p. 257, fig. 221.— Balachowsky and Mesnil 1935, p. 277, fig. 212 — 
Scholte 1935, p. 60 [Netherlands].— Vidal 1937, p. 197 [Morocco].— 
Lindberg 1948, p. 58 [Bulgaria; Yugoslavia; Prunus cerasus; Amygdalus; 
Rosa; Cyprus] .— Mancini 1949b, p. 36; 1952, p. 12; 1953b, p. 186; 
1953d, p. 16. — Novak and Wagner 1951, p. 70 [Crataegus pyracantha; 
Cydonia vulgaris; Prunus marasca; Prunus per ska; Prunus avium]. — 
Kiritshenko 1951, p. 247, fig. 327.— Poisson 1951, p. 1798, fig. 1591 — 
Gomez-Menor 1954, p. 371, figs. 1, 2 [Crataegus]; 1955a, p. 255, 
figs. 32, 33a-d; 1955b, p. 249 [Malus communis]. — Hoberlandt 1955, 
p. 98 [Iraq; Israel; Quercus ilex; Quercus cocci/era]. — Carayon 1960, 
p. 111.— Drake and Ruhoff 1962c, p. 133. 
Stephanitis piri [sic]: Borner 1935, p. 76. 
Stephanitis oschanini Vasiliev 1935, p. 151. 
Phyllontocheila schoutedeni (not Distant): Gomez-Menor 1956a, fig. 14 

(fig. only). 
Maecenas pyri: Drake and Ruhoff 1960a, p. 67. 
Type: Holotype 9, macropterous; locality unknown; Kiel Coll. (Copen- 
hagen Mus.). 

Distribution: Portugal; Spain; France; Belgium; Netherlands; Eng- 
land; Germany; Austria; Switzerland; Czechoslovakia; Hungary; Italy; 
Sicily; Sardinia; Yugoslavia; Albania; Rumania; Bulgaria; Poland; 
Sweden; Greece; Cyprus; Turkey; Iran; Iraq; Syria; Israel; U.S.S.R. 
(Caucasus; Armenia; Turkestan); Morocco. 

Host Plants: Amygdalus sp.; Castanea sativa; Chaenomeles japonica; 
Cormus sp; Cotoneaster integerrima; Crataegus crus-galli; Crataegus pyracantha; 
Crataegus sp.; Cydonia vulgaris; Juglans regia; Malus communis; Ligustrum sp.; 
Populus sp.; Prunus armeniaca; Prunus avium; Prunus cerasus; Prunus lusitanica; 
Prunus mahaleb; Prunus marasca; Prunus persica; Pyrus aria; Pyrus aucuparia; 
Pyrus communius; Pyrus malus; Pyrus sp.; Quercus cocci/era; Quercus ilex; Ribes 
nigrum; Rosa sp.; Sorbus aria; Sorbus torminalis; Robina pseudoacacia; Tilia 
sp.; Ulmus campestris; Ulmus effusa; Vaccinium myrtillus. 

Note: Predators and parasites [Silvestri; Gautier; Carayon]. Notes 
on hibernation [Gautier 1925, 1927a]. 



LACEBUGS OF THE WORLD 361 

*Stephanitis pyri var. sareptana Horvath 

Stephanitis pyri var. sareptana Horvath 1912b, pp. 320, 327 [Russia]. — 
Stichel 1960a, p. 302. 
Type: Sarepta, Russia; sex and deposition of type unknown. 
Distribution: U.S.S.R. 
Host Plant: Unrecorded. 
*Stephanitis pyrioides (Scott) 

Tingis pyrioides Scott 1874, pp. 291, 440 [Japan].— Uhler 1896, p. 265. 
Stephanitis azaleae Horvath 1905b, p. 568 [Netherlands; azalea]; 1906a, 
pp. 55, 56; 1912b, p. 333 [D.C.].— Stichel 1926, p. 110; 1928, p. 
206.— Scholte 1935, p. 60. 
Stephanitis pyrioides: Oshanin 1908, p. 425. — Dickerson and Weiss 
1917, p. 101, pi. 9 [N.J.; Pa.; Rhododendron hinodegeri; amoena; ledifolia 
alba; benigeri; yodogawa; kaempheri; pontica; mollis; indica; shirogeri; 
hatsugeri; shibori; amurasaki; schilippenbachii]. — Weiss 1918, p. 8, 
figs. 7-9.— Barber and Weiss 1922, p. 10, fig. 4.— Parshley 1922a, 
p. 236 [Conn.]; 1923b, p. 704, pi. 17, fig. 8.— McAtee 1923, p. 
149.— Drake 1923a, p. 104; 1928b, p. 101 [N.Y.; Kalmia latifolia]; 
1961b, p. 112 [New South Wales].— Blatchley 1926, p. 468, pi. 4, 
fig. 8. — Blanchard 1926, p. 361, fig. [Argentina]. — Esaki and Takeya 

1931, p. 53, figs. Id, 2d; pi. 4, fig. 4; pi. 5, fig. 4; pi. 6, fig. 4.— Esaki 

1932, p. 1635, fig.; 1954, p. 236, fig. 610.— Takeya 1932, p. 10 
[Korea]; 1951a, p. 12 [Taiwan; Pieris ovalifolia]; 1953c, pp. 8-12; 
1953d, p. 168.— White 1933, p. 631.— Kato 1933, pi. 31, fig. 2 .— 
Saito 1933, p. 6 [Rhododendron mucronatum; yedoense poukhanense]. — 
Shinji 1938, p. 316.— Froeschner 1944, p. 668 [Mo.].— Bailey 1950, 
p. 144; 1951, p. 56 [Mass.; R.I.; Rhododendron calendulaceum]; 1959, 
p. 65.— Gomez-Menor 1954, p. 377, fig. 3 [Morocco].— Maa 1957, 
pp. 128, 133.— Cobben 1958a, p. 9, fig. 10.— Stichel 1960a, p. 302 
[England].— Drake and Ruhoff 1926b, p. 250. 

Stephanitis pyriodes [sic]: Barber 1922a, p. 16. 

Stephanitis oberti (not Kolenati): Lindlinger 1927, p. 36 [Germany]. 

Stephanitis pyroides [sic]: Drake 1937b, p. 594 [China]. — Bosq 1937, 

p. 131. 
Stephanitis pyri (not Fabricius): Gomez-Menor 1956a, fig. 7 (fig. only). 
Type: Sex unknown; Japan; British Mus. 

Distribution: Japan; China; Korea; Taiwan; Germany; Netherlands; 
England; Argentina; Morocco; U.S. (N.Y.; Conn.; Mass.; R.I.; N.J.; 
Pa.; D.C; Fla.; Mo.); Australia (New South Wales). 

Host Plants: Kalmia latifolia; Pieris ovalifolia; Rhododendron species 
(including azalea) especially: amoena; amurasaki; benigeri; calendulaceum; 
hinodegeri; hatsugeri; indica; kaempheri; ledifolia alba; mollis; mucronatum; 
obtusum amoenum; pontica; shirogeri; shibori; schilippenbachii; yedoense pouk- 
hanense; yodogawa. 
Note: Life history studies [Bailey; Dickerson and Weiss]. 

6S1-552— 64 24 



362 U.S. NATIONAL MUSEUM BULLETIN 243 

*Stephanitis queenslandensis Hacker 

Stephanitis queenslandensis Hacker 1927, p. 28, pi. 7, fig. 8 [Queensland; 
Stephania hernandifolia]; 1928, p. 183. 
Type: Sex unknown; Queensland, Australia; Queensland Mus. 
Distribution: Australia (Queensland). 
Host Plant: Stephania hernandifolia. 
*Stephanitis querca Bergroth 

Stephanitis quercus Bergroth 1924, p. 83 [Luzon; oak]. — Drake 1930e, 

p. 168. 
Stephanitis querca: Drake and Poor 1937a, p. 17. 

Type: Sex unknown; Luzon, Philippine Islands; Helsin. Mus. 

Distribution: Philippine Islands (Luzon; Mindanao). 

Host Plant: Oak. 
*Stephanitis rhododendri Horvath 

Stephanitis rhododendri Horvath 1905b, p. 567 [Netherlands; Rhodo- 
dendron}; 1906a, p. 55; 1912b, pp. 321, 329.— Distant 1910c, p. 395, 
fig. [England].— Theobald 1911, p. 206; 1913, p. 297, figs. 40-43 — 
Steyer 1915, p. 434 [Germany]. — Van den Broeck and Schenk 
1915, p. 170.— Green 1916, p. 207.— Marchal 1917, p. 93 [France].— 
Barber and Weiss 1922, p. 11, figs. 5-7.— Lounsbury 1923, p. 548 
[Cape Province].— Wilson 1925, p. 52 [Switzerland].— Stichel 1928, 
p. 206; 1960a, p. 300, fig. 94 [Scotland; Denmark; Sweden; 
Finland].— Schmidt 1928, p. 205.— Scholte 1935, p. 60, fig. 13 — 
Balachowsky and Mesnil 1936, p. 1532, fig. 1225.— Proctor 1946, 
p. 74.— Van der Bruel 1947, p. 191, fig. 1 [Belgium].— Bailey 1951, 
p. 58 [N.H.; R.I.; Kalmia angustifolia]; 1959, p. 65.— Cobben 1958a, 
p. 5, figs. 5-8 [Rhododendron Van der Hoop; Mme. Mason; Mme. Carvalho; 
parsons gloriosium; Mr. R.S. Holford; america; caratacus; Cunningham'' s 
White; Antoon van Welie; fastuosum; cynthia; everestianum; Peter Koster; 
hollandia; Dr. V. H. Rutgers; Mme. de Bruin]. — Southwood and Leston 
1959, p. 146, pi. 18, fig. 5. 
Leptobyrsa species nova? Wirtner 1904, p. 202. 

Leptobyrsa explanata Heidemann 1908, p. 105, pi. 4, figs, e, f [Fla.; Mass.; 
D.C.; N.C.; Md.; Pa.; N.Y.; W. Va.; Ohio; Kalmia latifolia; Rhodo- 
dendron maximum]. — Felt 1910, p. 72, fig. 1. — Banks 1910, p. 56. — 
Torre-Bueno 1910, p. 31.— Smith 1910, p. 148 [N.J.].— Parshley 1914, 
p. 145 [Maine].— Crosby and Hadley 1915, pp. 409-415, pi. 23, figs. 
1-6; pi. 22 [Conn.].— Osborn and Drake 1916a, p. 240, pi. 7. 
Leptobyrsa rhododendri: Parshley 1917a, p. 15; 1917b, p. 55; 1923b, p. 
703.— Dickerson 1917, pp. 105-112, pi. 8.— Weiss 1918, p. 5, figs. 
4-6.— McAtee 1923, p. 149.— Downes 1925, p. 17, fig. 8 [B.C.]; 
1927a, p. 10.— Blatchley 1926, p. 469, fig. 105 [Va.].— Drake 1928b, 
p. 102.— Ascot 1930, p. 238.— Johnson 1936, p. 342, figs. 2-4; pis. 
13-15. 



LACE BUGS OF THE WORLD 363 

Type: Sex unknown; Boskoop, Netherlands; Hungarian Mus. 

Distribution: Netherlands; Belgium; France; Germany; Switzerland; 
Finland; Sweden; Denmark; Scotland; England; South Africa (Cape 
Province); Canada (Nfld.; B.C.); U.S. (Maine; N.H.; Mass.; Conn.; R.I.; 
N.Y.; N.J.; Pa.; Ohio; Md.; D.C.; Va.; W. Va.; N.C.; S. C; Fla.; Oreg.; 
Wash.; Vt); New Zealand. 

Host Plants: Kalmia angustifolia; Kalmia latifolia; Pieris floribunda; Pieris 
japonica; Rhododendron species, those reported: america; Antoon van Welie; 
caractacus; Cunningham's White; cynthia; Dr. V.H. Rutgers; everestianum; 
Jastuosum; hollandia; maximum; Mme. Mason; Mme. Carvalho; Mr. R.S. 
Holford; Mme. de Bruin; parsons gloriosum; Peter Koster; Van der Hoop. 

Note: Life history studies [Heidemann; Crosby and Hadley; Dickerson; 
Johnson; Wilson; Felt; Bruel; Balachowsky and Mesnil]. Found on 120 
species of Rhododendron [Van den Broek and Schenk]. 

♦Stephanitis steeleae Drake and Maa 

Stephanitis steeleae Drake and Maa 1954, p. 116 [India]. 
Type: Holotype cf, macropterous; Cha Che, Delai Valley, Mishmi Hills, 
Assam, India; British Mus. 
Distribution: India. 
Host Plant: Unrecorded. 

*Stephanitis subfasciata Horvath 

Stephanitis subfasciata Horvath 1912b, pp. 320, 325 [China; Taiwan]. — • 
Esaki 1926, p. 163.— Drake and Poor 1937b, p. 404 [Java].— Drake 
1948b, p. 56 [India]; 1956d, p. 115 [Burma; Cocculus; Hernandia; 
Peleliu; Argaut]. — Maa 1957, pp. 129, 132 [Cocculus trilobus]. 
Stephanitis (Menodora) subfasciata: Wu 1935, p. 449. 
Type: Unknown. 

Distribution: China; Taiwan; Burma; India; Palau Islands (Peleliu; 
Argaut); Greater Sunda Islands (Java). 

Host Plants: Cocculus trilobus; Cocculus sp.; Hernandia sp. 
♦Stephanitis svensoni Drake 

Stephanitis svensoni Drake 1948b, p. 51 [Japan]. — Takeya 1953d, p. 169, 

fig. la. 
Stephanitis shirozui Takeya 1951b, p. 60, fig. 2 [Illicium religiosum]. 
Type: Holotype o", macropterous; Shimabara Pen., Japan; Drake Coll. 
(USNM). 
Distribution: Japan. 
Host Plant: Illicium religiosum. 
♦Stephanitis tabidula Horvath 

Stephanitis tabidula Horvath 1912b, pp. 323, 333 [Japan]. 
Type: Sex unknown; Kanagawa, Japan; Hungarian Mus. 
Distribution: Japan. 
Host Plant: Unrecorded. 



364 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Stephanitis takeyai Drake and Maa Plate 48 

Tingis globulifera Matsumura 1905, p. 36, pi. 19, fig. 16 [Japan]. 
Stephanitis globulifera: Matsumura 1908, p. 148, fig.; 1930, p. 161, pi. 14, 
fig. 16; 1931, p. 1203, fig. 7— Horvath 1912b, pp. 322, 330 [Cin- 
namomum camphora].- — Drake 1923a, p. 104. — Kato 1933, pi. 31, fig. 
4. — Takeya 1930b, p. 72; 1951a, p. 11 [Lindera citriodora; Styrax 
iaponica; Pierus ovalifolia]; 1953d, p. 168, fig. lb [Lyonia neziki; Lindera 
obtusiloba; Lindera umbellata; Salix; Aperula citriodora; Pierus japonica]. — 
Drake and Poor 1937b, p. 403.— Bailey 1950, pp. 143-145 [Conn.]; 
1951, p. 53; 1959, p. 65.— Warner 1956, p. 6 [N.J.]. 
Stephanitis takeyai Drake and Maa 1955, p. 10 [India]. 
Type: Unknown. 

Distribution: Japan; India; U.S. [Conn.; N.J.]. 

Host Plants: Andromedea sp.; Aperula citriodora; Cinnamomum camphora; 
Lindera sericea; Lindera citriodora; Lindera obtusiloba; Lindera umbellata; Lyonia 
neziki; Pierus japonica; Pierus ovalifolia; Salix sp.; Styrax japonica. 

*Stephanitis typica (Distant) Plate 47 

Cadamustus typicus Distant 1903a, p. 47 [Ceylon]; 1903b, p. 132, fig. 95 

[Hedychium; Musa], 
Stephanitis typicus: Distant 1910a, p. 108 [India; Luzon]. — Drake 1926e, 
p. 335; 1933, p. 1016; 1937b, p. 594; 1938b, p. 197 [Hong Kong].— 
Nagaraj and Menon 1956, pp. 161-165.— Mathen 1960, pp. 8-24, 
figs. 1-6, pis. 1-2, tables 1-6 [Artocarpus integrifolia]. — Shanta, 
Menon, and Pillai 1960, p. 58-62. 
Stephanitis typica: Horvath 1912b, pp. 320, 325 [Java; Musa paradisiaca 
sapientum]; 1912c, p. 341; 1926, p. 329 [Sumatra].— Bergroth 1914a, 
p. 361 [Taiwan].— Fletcher 1920, p. 263.— Drake 1923a, p. 104; 
1960, p. 373 [Korea; Papua].— Esaki 1926, p. 163.— Takeya 1931, p. 
70 [Alpinia nutans]; 1951a, p. 8 [Japan; Zingiber mioga]; 1952, pp. 39-46, 
fig. 1 [Pakistan; Malaya; Alpinia japonica; Alpinia speciosa; Zingiber 
kawangoii]. — Hoffman 1935, p. 643, pi. 47 [China; Elettaria cardamo- 
mum; coconut; camphor]. — Takahashi 1936, p. 295. — Drake and Poor 
1936a, p. 443 [Hainan]; 1937a, p. 16; 1937b, p. 404 [Mindanao; 
Annona muricata]. — Shirozu 1939, p. 205 [Alpinia chinensis]. — Maa 1957, 
pp. 127, 133. 
Stephanitis Indiana Drake 1948b, p. 51. 
Type: Sex unknown; Pundalu-oya, Ceylon; British Mus. 
Distribution: Ceylon; India; Pakistan; Hong Kong; China (Hainan 
Island); Japan; Korea; Taiwan; New Guinea (Papua); Greater Sunda 
Islands (Sumatra; Java); Federation of Malaya; Philippine Islands (Luzon; 
Mindanao; Leyte). 

Host Plants: Alpinia chinensis; Alpinia nutans; Alpinia japonica; Alpinia speciosa; 
Annona muricata; Artocarpus integrifolia; Elettaria cardamomum; Hedychium sp.; 



LACEBUGS OF THE WORLD 365 

Musa paradisciaca sapientum; Musa sp.; Zingiber kawagoii; Zingiber mioga; 
camphor; coconut. 

Note: (1) Life history studies [Hoffman; Mathen]; (2) possible vector of 
wilt (root) disease of coconut palm [Nagaraj and Menon; Mathen; Shanta, 
Menon and Pillai]; (3) Indiana here placed in synonymy. (New synonymy.) 

*Stephanitis veridica Drake 

Stephanitis veridica Drake 1948b, p. 54 [Taiwan]. 
Type: Holotype o 71 , macropterous; Tainansliu, Formosa; Drake Coll. 
(USNM). 
Distribution: Taiwan. 
Host Plant: Unrecorded. 

*Stephanitis yasumatsui Takeya 

Stephanitis yasumatsui Takey a 1951b, p. 58, fig. 1 [Japan]. 
Type: Holotype o"; Sobo-san, Prov. Bungo, Japan; Kyushu Univ. 
Distribution: Japan. 
Host Plant: Unrecorded. 

*Stephanitis yunnana Drake and Maa 

Stephanitis yunnana Drake and Maa 1955, p. 9 [China]. 
Type: Holotype 9, macropterous; Yunnan, China; Drake Coll. (USNM). 
Distribution: China. 
Host Plant: Unrecorded. 

Subgenus STEPHANITIS (MENODORA) Horvath 

Stephanitis (Menodora) Horvath 1912b, pp. 319, 324.— Takeya 1951a, 
p. 7. — Drake and Maa 1953, p. 99 (catalog). — Drake and Ruhoff 
1960a, p. 83. 
Type Species: Stephanitis (Menodora) formosa Horvath. 

*Stephanitis (Menodora) charieis Drake and Mohanasundarum 

Stephanitis (Menodora) charieis Drake and Mohanasundarum 1961, 
p. Ill, fig. 3 [India; Artocarpus integrifolia]. 
Type: Holotype o 71 , macropterous; Coimbatore, India; Drake Coll. 
(USNM). 
Distribution: India. 
Host Plant: Artocarpus integrifolia. 

*Stephanitis (Menodora) formosa Horvath 

Stephanitis (Menodora) formosa Horvath 1912b, pp. 319, 324, fig. 1 
[Taiwan]. 

Stephanitis formosa: Esaki 1926, p. 163. 
Type: Sex unknown; Takao, Formosa; Hungarian Mus. 
Distribution : Taiwan. 
Host Plant: Unrecorded. 



366 U.S. NATIONAL MUSEUM BULLETIN 2 43 

*Stephanitis (Menodora) kardia Drake and RuhofT 

Stephanitis {Menodora) kardia Drake and RuhofT 1960c, p. 36, fig. 5 a, b 
[Singapore]. 
Type: Holotype cf, macropterous; Singapore; Drake Coll. (USNM). 
Distribution : Singapore. 
Host Plant: Unrecorded. 

*Stephanitis (Menodora) sondaica Horvath 

Stephanitis {Menodora) sondaica Horvath 1912b, p. 338 [Java; Musa 

sapientum]. — Takeya 1951a, p. 8 [Taiwan]. 
Stephanitis sondaica: Horvath 1912c, p. 341. 
Type: Sex unknown; Muara Antjol near Batavia, Java; Hungarian 
Mus. 

Distribution: Greater Sunda Islands (Java); Taiwan. 
Host Plant: Musa paradisiaca sapientum. 

Subgenus STEPHANITIS (NORBA) Horvath 

Stephanitis {Norba) Horvath 1912b, pp. 323, 334.— Takeya 1951a, 
p. 12.— Drake and Maa 1953, p. 100 (catalog) .—Drake and Runoff 
1960a, p. 83. 
Type Species: Stephanitis {Norba) mendica Horvath. 

*Stephanitis (Norba) aperta Horvath 

Stephanitis {Norba) aperta Horvath 1912b, pp. 323, 335 [Japan]. — Takeya 
1931, p. 75, pi. 7, fig. 4; pi. 9, fig. 1; 1951a, p. 13 [Taiwan; Cin- 
namomum camphor a; Machilus thunbergii]. 
Stephanitis aperta: Maa 1957, pp. 127, 132. 
Type: Sex unknown; Sakuna, Japan; Hungarian Mus. 
Distribution: Japan; Taiwan; China. 
Host Plants: Cinnamomum camphor a; Machilus thunbergii. 
*Stephanitis (Norba) astralis Drake and Poor 

Stephanitis astralis Drake and Poor 1941, p. 164 [Amboina]. 
Type: Holotype o 71 ; Amboina Island; Cal. Acad. 
Distribution: Moluccas (Amboina Island). 
Host Plant: Unrecorded. 

*Stephanitis (Norba) chinensis Drake 

Stephanitis {Norba) chinensis Drake 1948b, pp. 46, 48, 55, figs. 3, 4 
[China; tea]. 
Type: Holotype o 71 , macropterous; Meitanhsieh, Kweichow Province, 
China; Drake Coll. (USNM). 
Distribution: China. 
Host Plant: Tea. 



LACEBUGS OF THE WORLD 367 

*Stephanitis (Norba) esakii Takeya 

Stephanitis {Norba) esakii Takeya 1931, p. 72, pi. 8, figs. 7-10 [Taiwan; 

Mach ilus thunb erg ii] . 
Stephanitis (Norba) shintenana Drake 1948b, pp. 46, 48, 56. 
Stephanitis esakii: Maa 1957, pp. 125, 132 [Machilus; Tetradenia doli- 
chocarpa]. 
Type: Holotype cf; Musha, Formosa; Kyushu Univ. 
Distribution: Taiwan. 
Host Plants: Machilus thunbergii; Machilus sp.; Tetradenia dolichocarpa. 

♦Stephanitis (Norba) exigua Horvath 

Stephanitis (Norba) exigua Horvath 1912b, pp. 323, 336 [Okinawa; Japan; 
Lycopodiaceae]. — Takeya 1953d, p. 168 [Alachilus thunbergii]. 

Stephanitis (Norba) vitrea Takeya 1931, p. 74, pi. 7, fig. 3; pi. 8, figs. 11, 12. 

Stephanitis exigua: Drake 1937b, p. 594 [China]. 
Type: Unknown. 

Distribution: Ryukyu Islands (Okinawa); Japan; China. 
Host Plants: Machilus thunbergii; Lycopodiaceae. 

♦Stephanitis (Norba) laratana Drake 

Stephanitis (Norba) laratana Drake 1948b, pp. 46, 55 [Larat]. 
Type: Holotype cf, macropterous; Larat; Drake Coll. (USNM). 
Distribution: Tanimbar Islands (Larat). 
Host Plant: Unrecorded. 

*Stephanitis (Norba) macaona Drake 

Stephanitis (Norba) macaona Drake 1948b, pp. 46, 47, 55 [China]. 
Type: Holotype cf, macropterous; Macao, Kwangtung Province, China; 
Drake Coll. (USNM). 
Distribution: China. 
Host Plant: Unrecorded. 

♦Stephanitis (Norba) mendica Horvath 

Stephanitis (Norba) mendica Horvath 1912b, pp. 323, 334 [Japan]. — 
Takeya 1931, p. 77, pi. 9, figs. 2-6 [Cinnamomum pedunculatum]. 
Type: Sex unknown; Japan; Hungarian Mus. 
Distribution: Japan. 
Host Plant: Cinnamomum pedunculatum. 

♦Stephanitis (Norba) nitor Drake and Poor (emendation) 

Stephanitis nitoris Drake and Poor 1937a, p. 17 [Luzon; Negros]; 1937b, 
p. 403 [Uvaria rufa]. 
Type: Holotype d\ macropterous; Mount Maquiling, Laguna Province, 
Luzon, Philippine Islands; USNM. 

Distribution: Philippine Islands (Luzon; Negros); Greater Sunda 
Islands (Borneo). 
Host Plant: Uvaria rufa. 



368 U.S. NATIONAL MUSEUM BULLETIN 243 

*Stephanitis (Norba) outouana Drake and Maa 

Stephanitis {Norba) outouana Drake and Maa 1953, p. 97 [China]. 
Type: Holotype o 71 , macropterous; Outou, Kienyang Hsien, Fukien, 
China; Drake Coll. (USNM). 
Distribution: China. 
Host Plant: Unrecorded. 

*Stephanitis (Norba) pagana Drake and Maa 

Stephanitis {Norba) pagana Drake and Maa 1953, p. 98 [China]. 

Type: Holotype o 71 , macropterous; Tsinleong Shan, E. Kwangtung, 
China; Drake Coll. (USNM). 

Distribution: China. 

Host Plant: Unrecorded. 

*Stephanitis (Norba) sordida Distant 

Stephanitis sordidus Distant 1909a, p. 114 [Burma]; 1910a, p. 109. 

Stephanitis sordida: Horvath 1912b, p. 338. 

Stephanitis {Stephanitis) sordida: Drake and Maa 1953, p. 101. 

Stephanitis {Norba) sordida: Drake and Maa 1954, p. 116. 
Type: Sex unknown; Tenasserim, Merqui, Burma; British Mus. 
Distribution: Burma. 
Host Plant: Unrecorded. 

*Stephanitis (Norba) suffusa (Distant) 

Cadamustus suffusus Distant 1903a, p. 47 [Ceylon]; 1903b, p. 133 [Burma]. 

Stephanitis suffusa: Horvath 1912b, p. 338. 

Stephanitis {Norba) bankana Drake 1948b, pp. 46, 47, 54 [Bangka; China; 
Per sea gratissima]. 

Stephanitis {Stephanitis) suffusa: Drake and Maa 1953, p. 101. 

Stephanitis {Norba) suffusa: Drake and Maa 1954, p. 116. 
Type: Sex unknown; Matale, Ceylon; British Mus. 
Distribution: Ceylon; Burma; China; Bangka Island. 
Host Plant: Per sea gratissima. 

Subgenus STEPHANITIS (OMOPLAX) Horvath 

Stephanitis {Omoplax) Horvath 1912b, pp. 323, 336.— Takeya 1951a, p. 
14.— Drake and Maa 1953, p. 100 (catalog) .—Drake and Ruhoff 
1960a, p. 83. 
Type Species: Stephanitis {Omoplax) desecta Horvath. 

*Stephanitis (Omoplax) desecta Horvath 

Stephanitis {Omoplax) desecta Horvath 1912b, pp. 323, 337, fig. 3 [Japan].— 
Drake 1956d, p. 116, fig. 8. 
Type: Sex unknown; Ogasawara, Japan; Hungarian Mus. 
Distribution: Japan. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 369 

Genus STYMNONOTUS Reuter 

Stymnonotus Reuter 1887, p. 103.— Drake and RuhofF 1960a, p. 83. 
Type Species: Stymnonotus apicalis Reuter. 
Stymnonotus apicalis Reuter 

Stymnonotus apicalis Reuter 1887, p. 104 [Malagasy]. — Lethierry and 
Severin 1896, p. 13. 
Type: Holotype o" ; Nossibe, Madagascar; Frankfort Mus. 
Distribution : Malagasy Republic. 
Host Plant: Unrecorded. 

Genus TANYBYRSA Drake 

Tanybyrsa Drake 1942a, p. 21.— Drake and Ruhoff 1960a, p. 83. 
Type Species: Compseuta secundus Hacker. 
*Tanybyrsa ampliata (Hacker) 

Compseuta ampliatus Hacker 1927, p. 26, pi. 7, fig. 5 [Queensland]. 
Tanybyrsa ampliata: Drake 1942a, p. 21; 1961b, p. 112. 
Type: Sex unknown; Southport, Queensland, Australia; Queensland 
Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Tanybyrsa cumberi Drake 

Tanybyrsa cumberi Drake 1959, p. 67, fig. 1 [New Zealand]. 

Type: Holotype cf; Arapae, Te Kuiti-Tawaro Range, Waitomo County, 
New Zealand; Entomology Division, Dept. Scientific and Industrial Re- 
search, Nelson, New Zealand. 

Distribution: New Zealand. 

Host Plant: Unrecorded. 

*Tanybyrsa secunda (Hacker) 

Compseuta secundus Hacker 1927, p. 27, pi. 7, fig. 6 [Queensland; New 

South Wales]. 
Tanybyrsa secunda: Drake 1942a, p. 21. 
Type: Sex unknown; Australia; Queensland Mus. 
Distribution: Australia (Queensland; New South Wales). 
Host Plant: Unrecorded. 

Genus TANYTINGIS Drake 

Tanytingis Drake 1939b, p. 205.— Drake and Ruhoff 1960a, p. 83. 
Type Species: Tanytingis takahashii Drake. 



370 U.S. NATIONAL MUSEUM BULLETIN 243 

*Tanytingis assamana Drake and Lutz 

Tanytingis assamana Drake and Lutz 1953, p. 105 [India]. 

Type: Holotype cf, macropterous; Shillong, Assam, India; Drake Coll. 
(USNM). 

Distribution: India. 

Host Plant: Unrecorded. 

*Tanytingis takahashii Drake 

Tanytingis takahashii Drake 1939b, p. 206 [Taiwan]. — Takeya 1951a, 
p. 17. 
Type: Holotype cf; Buta, Taihoku-shu, Formosa; Kyushu Univ. 
Distribution: Taiwan. 
Host Plant: Unrecorded. 

Genus TELEONEMIA Costa 

Teleonemia Costa 1864, p. 144.— Stal 1873, pp. 122, 131.— Uhler 
1886, p. 22.— Lethierry and Severin 1896, p. 22.— Champion 1898a, 
p. 34 (key).— Banks 1910, p. 57.— Van Duzee 1916, p. 26; 1917b, 
p. 221— Drake 1918b, pp. 323-332 (key) .— Horvath 1925b, p. 
219.— Blatchley 1926, p. 487 (key).— Monte 1939b, p. 79; 1941e, 
p. 133.— Hurd 1946, p. 447.— Drake and Ruhoff 1960a, p. 84. 

Tingis (Amaurosterphus) Stal 1868, p. 92. 

Teleonemia (Amaurosterphus): Stal 1873, p. 131. — Drake and Runoff 
1960a, p. 35. 

Tingis (Americia) Stal 1873, p. 131.— Drake and Ruhoff 1960a, p. 36. 
Type Species: Teleonemia funerea Costa. 

*Teleonemia abdita Drake 

Teleonemia abdita Drake 1939d, p. 527 [Brazil]. 
Type: Holotype cT; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Teleonemia absimilis Drake and Hambleton 

Teleonemia absimilis Drake and Hambleton 1944b, p. 122 [Colombia]. 

Type: Holotype 9, macropterous; Villa Vicenzio, Colombia; Drake Coll. 
(USNM). 

Distribution: Colombia. 

Host Plant: Unrecorded. 

*Teleonemia aemula Monte 

Teleonemia aemula Monte 1942b, p. 137, fig. 2 [Brazil]. 

Type: Holotype 9; Nova Teutonia, Santa Catarina, Brazil; Monte Coll. 
(Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 371 

*Teleonemia altilis Drake and Hambleton 

Teleonemia altilis Drake and Hambleton 1944b, p. 122 [Bolivia]. 

Type: Holotype 9, macropterous; Las Juntas, Bolivia; Drake Coll. 
(USNM). 

Distribution: Bolivia. 

Host Plant: Unrecorded. 

Teleonemia amazonica Horvath 

Teleonemia {Americia) amazonica Horvath 1925b, p. 220 [Brazil]. 
Type: Holotype 9; Manaos, Brazil; Hungarian Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

Teleonemia angustata Monte 

Teleonemia angustata Monte 1943d, p. 268 [Brazil]. 
Type: Holotype o 71 ; Belo Horizon te, Minas Gerais, Brazil; Monte Coll. 
(Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Teleonemia annae (Kirkaldy) 

Americia annae Kirkaldy 1905, p. 216 [Peru]. 
Teleonemia annae: Drake and Hambleton 1938b, p. 52 [Brazil]. 
Type: Holotype 9, macropterous; Marcapata, Peru; Drake Coll. 
(USNM). 
Distribution: Peru; Brazil. 
Host Plant: Unrecorded. 

*Teleonemia argentinensis Drake and Poor 

Teleonemia argentinensis Drake and Poor 1942, p. 300 [Argentina]. 

Type: Holotype o 71 ; Estacion Experimental de Loreto, Misiones, Argen- 
tina; Argentina Mus. 

Distribution : Argentina. 

Host Plant: Unrecorded. 

*Teleonemia aterrima Stal 

Teleonemia aterrima Stal 1873, p. 131 [Colombia]. — Champion 1898b, 
p. 62, pi. 3, fig. 3 [Brazil].— Drake 1922c, p. 356 [Peru]; 1930b, 
p. 25.— Drake and Poor 1937d, p. 303, pi. 36, fig. 15.— Drake and 
Hambleton 1938b, p. 52— Blote 1945, p. 89.— Silva 1956, p. 51, 
fig. 19. 
Type: Holotype cf ; Bogota, Nova Granada; Stockholm Mus. 
Distribution: Colombia; Brazil; Peru. 
Host Plant: Unrecorded. 



372 U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Teleonemia atrata Champion 

Teleonemia atrata Champion 1898a, p. 38, pi. 3, fig. 7 [Panama]. — 
Osborn and Drake 1915b, p. 536 [Guatemala].— Hurd 1946, p. 448 
[Brazil]. 
Type: Holotype 9; Bugaba, Panama; British Mus. 
Distribution: Panama; Guatemala; Brazil. 
Host Plant: Unrecorded. 

Teleonemia atriflava Monte 

Teleonemia atriflava Monte 1943b, p. 204 [Argentina]. 

Type: Holotype 9; El Dorado, Misiones, Argentina; Monte Coll. (Mus. 
Nacional) . 

Distribution: Argentina. 

Host Plant: Unrecorded. 

*Teleonemia bahiana Drake 

Teleonemia bahiana Drake 1942a, p. 1 [Brazil]; 1947a, p. 1. — Silva 
1956, p. 51. 
Type: Holotype 9, macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Teleonemia barberi Drake 

Teleonemia barberi Drake 1918b, pp. 325, 328 [Ariz.; Tex.; Chilopsis]. 
Type: Holotype 9, macropterous; Huachuca Mountains, Arizona, 
U.S.; USNM. 
Distribution: U.S. (Ariz.; Tex.). 
Host Plant: Chilopsis sp. 

*Teleonemia belfragii Stal 

Teleonemia belfragii Stal 1873, p. 132 [Tex.]. — Champion 1898b, p. 62, 
pi. 3, fig. 8.— Van Duzee 1909, p. 173 [Fla.].— Drake 1926b, p. 
376.— Hurd 1946, p. 448 [Miss.; Ala.]. 
Teleonemia belfragei [sic]: Lethierry and Severin 1896, p. 22. — Drake 

1918b, p. 331 [Callicarpa americana].— Blatchley 1926, p. 490. 
Teleonemia belfragei [sic]: Barber 1914, p. 507. 
Type: Holotype 9; Texas, U.S.; Stockholm Mus. 
Distribution: U.S. (Tex.; Miss.; Ala.; Fla.). 
Host Plant: Callicarpa americana. 

Teleonemia bierigi Monte 

Teleonemia bierigi Monte 1943d, p. 269 [Costa Rica]. 
Type: Holotype cf; La Perla, Costa Rica; Monte Coll. (Mus. Nacional). 
Distribution: Costa Rica. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 373 

*Teleonemia bifasciata Champion 

Teleonemia bifasciata Champion 1898a, p. 38, pi. 3, figs. 8, 8a [Guatemala; 
Panama]. — Drake and Bruner 1924a, p. 145 [Grenada]. — Drake and 
Hambleton 1940, p. 534 [Brazil].— Hurd 1946, p. 448 [Lantana]. 
Type: Sex unknown; Bugaba, Panama; British Mus. 
Distribution: Brazil; Panama; Guatemala; Windward Islands 
(Grenada) . 
Host Plant: Lantana sp. 

*Teleonemia boliviana Drake 

Teleonemia boliviana Drake 1939d, p. 528 [Bolivia; Peru]. 
Type: Holotype d"; San Antonio, Bolivia; Stockholm Mus. 
Distribution: Bolivia; Peru. 
Host Plant: Unrecorded. 

Teleonemia bondari Monte 

Teleonemia bondari Monte 1943d, p. 270 [Brazil]. 

Type: Holotype cf; Belo Horizon te, Minas Gerais, Brazil; Monte Coll. 
(Mus. Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Teleonemia bosqi Monte 

Teleonemia bosqi Monte 1943b, p. 202, fig. [Argentina]. 

Type: Holotype o"; El Dorado, Misiones, Argentina; Monte Coll. (Mus. 
Nacional). 

Distribution: Argentina. 

Host Plant: Unrecorded. 

♦Teleonemia brevipennis Champion 

Teleonemia brevipennis Champion 1898b, p. 63, pi. 3, fig. 9 [Brazil]. — 
Drake 1922c, p. 357 [Peru]; 1929, p. 35; 1930a, p. 1.— Drake and 
Hambleton 1934, p. 438 [Cassia]; 1938a, p. 45 [Vernonia polyanthes]. — 
Costa Lima 1936, p. 130.— Monte 1938d, p. 390 [Argentina]; 1939b, 
p. 79 [Venezuela]; 1940c, p. 191; 1940d, p. 101.— Drake and Poor 
1939b, p. 96 [Buddleia].— Silva 1956, p. 51, fig. 20 [Ipomoea fistulosa]. 
Teleonemia brevicornis [sic]: Drake 1935, p. 10 [Paraguay]. 
Type: Holotype 9; Amazons, Brazil; British Mus. 
Distribution: Brazil; Argentina; Peru; Paraguay; Venezuela. 
Host Plants: Buddleia sp.; Cassia sp.; Ipomoea fistulosa; Vernonia polyanthes. 

*Teleonemia carmelana (Berg) 

Leptostyla carmelana Berg 1892, p. 99 [Uruguay]. 
Cantacader chilensis Reed 1900, p. 180 [Chile]. 
Teleonemia {Cantacader) chiliensis [sic]: Drake 1922b, p. 50. 
Teleonemia chilensis: Drake 1922c, p. 358 [Brazil; Argentina]; 1935, p. 10 
[Peru; Paraguay]; 1936b, p. 699. 



374 U.S. NATIONAL MUSEUM BULLETIN 243 

Teleonemia jensoni Bergroth 1922, p. 150. 

Teleonemia carmelana: Drake 1935, p. 11; 1939c, p. 332 [Bolivia]. — Drake 
and Poor 1938b, p. 107.— Monte 1938d, p. 390; 1941e, p. 135 [Lan- 
tana]; 1947b, p. 432 [Lippiajuncea; Rhaphithamnus spinosus]. 
Type: Sex unknown; Uruguay; La Plata Mus. 

Distribution: Uruguay; Paraguay; Peru; Argentina; Chile; Brazil; 
Bolivia. 
Host Plants: Lantana sp.; Lippiajuncea; Rhaphithamnus spinosus. 

*Teleonemia chacoana Drake 

Teleonemia chacoana Drake 1942a, p. 1 [Paraguay]. 

Type: Holotype 9, macropterous; Chaco, Paraguay; Drake Coll. 
(USNM). 

Distribution: Paraguay. 

Host Plant: Unrecorded. 

Teleonemia chapadiana Drake 

Teleonemia chapadiana Drake 1922c, p. 356 [Brazil]. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Teleonemia consors Drake 

Teleonemia consors Drake 1918b, pp. 324, 327 [Ariz.]. 
Type: Holotype 9, macropterous; Bonita, Post Creek Can., Arizona, 
U.S.; Drake Coll. (USNM). 
Distribution: U.S. (Ariz.). 
Host Plant: Unrecorded. 

Teleonemia crassispinosa Monte 

Teleonemia crassispinosa Monte 1946b, p. 285 [Brazil]. — Silva 1956, 
p. 54. 
Type: Holotype cf; Agua Preta, Baia, Brazil; Monte Coll. (Mus. 
Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Teleonemia cylindricornis Champion 

Teleonemia cylindricornis Champion 1898a, p. 41, pi. 3, figs. 14, 14a 

[Guatemala; British Honduras]. — Van Duzee 1907, p. 22 [Jamaica]. 

—Drake 1925c, p. 38 [Miss.; 111.].— Blatchley 1926, p. 488 — 

Froeschner 1944, p. 669. 

Type: Sex unknown; San Geronimo, Vera Paz, Guatemala; British 

Mus. 

Distribution: Jamaica; British Honduras; Guatemala; Mexico; U.S. 
(Miss.; 111.). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 375 

*Teleonemia dulcis Drake 

Teleonemia dulcis Drake 1939d, p. 525 [Brazil]. 

Type: Holotype 9, macropterous; Belem, Para, Brazil; Drake Coll. 
(USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Teleonemia elata Drake 

Teleonemia chilensis var. elata Drake 1935, p. 10 [Brazil; Paraguay]. — 
Drake and Poor 1938b, p. 107 [Chile; Peru].— Monte 1939b, p. 79 
[Lantana]. 
Teleonemia elata: Monte 1942b, p. 138. 
Type: Holotype cf ; Rio Grande do Sul, Brazil; Vienna Mus. 
Distribution: Brazil; Paraguay; Chile; Peru. 
Host Plant: Lantana sp. 

Teleonemia elevata (Fabricius) 

Aradus elevatus Fabricius 1803, p. 120. 

Tingis {Tropidocheila) elevata: Stal 1868, p. 91. 

Tingis elevata: Walker 1873a, p. 181 [South America]. 

Monanthia elevata: Walker 1873a, p. 191. 

Teleonemia elevata: Stal 1873, p. 132. 
Type: Lectotype 9, macropterous; "Americae meridionalis"; Copen- 
hagen Mus. 
Distribution: Uncertain, possibly South America. 
Host Plant: Unrecorded. 

*Teleonemia forticornis Champion 

Teleonemia forticornis Champion 1898a, p. 36, pi. 3, fig. 5 [Panama]. — 
Drake and Hambleton 1938b, p. 52 [Brazil; Peru; Argentina]. — 
Drake and Poor 1939b, p. 95. — Monte 1939b, p. 79 [Ipomoea batatas]. 
Type: Holotype o 71 ; Bugaba, Panama; British Mus. 
Distribution: Panama; Brazil; Peru; Argentina. 
Host Plant: Ipomoea batatas. 

*Teleonemia funerea Costa 

Teleonemia junerea Costa 1864, p. 145, pi. 2, fig. 5. — Stal 1873, p. 132. — 

Drake and Hambleton 1938b, p. 52 [Brazil]. 
Monanthia junerea: Walker 1873a, p. 197. 
Type: Unknown. 
Distribution: Brazil. 
Host Plant: Unrecorded. 
Note: Type specimen apparently lost [Drake and Hambleton]. 



376 U.S. NATIONAL MUSEUM BULLETIN 24 3 

Teleonemia granulosa Monte 

Teleonemia granulosa Monte 1942b, p. 139 [Argentina]. 

Type: Holotype o"; San Javier, Misiones, Argentina; Monte Coll. 
(Mus. Nacional). 

Distribution : Argentina. 

Host Plant: Unrecorded. 

*Teleonemia guyanensis Drake and Carvalho 

Teleonemia guyanensis Drake and Carvalho 1944, p. 41 [British Guiana]. 

Type: Holotype 9, macropterous; Mallali, British Guiana; Drake Coll. 
(USNM). 

Distribution: British Guiana. 

Host Plant: Unrecorded. 

*Teleonemia hasemani Drake 

Teleonemia hasemani Drake 1922c, p. 357 [Brazil]. 
Type: Holotype 9; Sao Antonio de Guapore, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Teleonemia huachucae Drake 

Teleonemia huachucae Drake 1941b, p. 140 [Ariz.]. 
Type: Holotype cf; Huachuca Mountains, Arizona, U.S.; Cal. Acad. 
Distribution: U.S. (Ariz.). 
Host Plant: Anisacanthus thurberi. 

*Teleonemia inops Drake and Hambleton 

Teleonemia inops Drake and Hambleton 1944b, p. 122 [Honduras]. 

Type: Holotype cT, macropterous; La Ceiba, Honduras; Drake Coll. 
(USNM). 

Distribution: Honduras. 

Host Plant: Unrecorded. 
Teleonemia inornata Monte 

Teleonemia inornata Monte 1941a, p. 377 [Bolivia], 

Type: Holotype o 71 ; Caranavi, Coroico, Nor Yungas, Bolivia; Monte 
Coll. (Mus. Nacional). 

Distribution: Bolivia. 

Host Plant: Unrecorded. 

Teleonemia Jamaicans Hurd (nom. nud.) 

Teleonemia Jamaicans Hurd 1946, p. 448. 
*Teleonemia jubata Drake and Hambleton 

Teleonemia jubata Drake and Hambleton 1939, p. 153 [Brazil]. 
Type: Holotype 9, macropterous; Belo Horizon te, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 377 

*Teleonemia jucunda Drake 

Teleonemia jucunda Drake 1939d, p. 526 [Brazil; French Guiana; British 
Guiana]; 1947a, p. 1.— Silva 1956, p. 54. 
Type: Holotype o 71 , macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil; French Guiana; British Guiana. 
Host Plant: Unrecorded. 

*Teleonemia leitei Drake and Hambleton 

Teleonemia leitei Drake and Hambleton 1939, p. 153 [Brazil; Pithecoc- 
tenium echinatum]. 
Type: Holotype o", macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Pithecoctenium echinatum. 

*Teleonemia limbata (Stal) 

Tingis {Americia) limbata Stal 1873, p. 131 [Brazil; Colombia]. 
Lasiacantha {Americia) limbata: Lethierry and Severin 1896, p. 19. 
Teleonemia limbata: Champion 1898b, p. 62, pi. 3, fig. 10. — -Drake 1922c, 
p. 356; 1930a, p. 1; 1935, p. 10 [Paraguay]; 1936b, p. 699 [Argen- 
tina]. — Drake and Poor 1937d, p. 302.— Drake and Hambleton 
1938b, p. 52.— Monte 1939b, p. 79 [Lantana camara]; 1940c, p. 191. 
Type: Unknown. 

Distribution: Brazil; Paraguay; Argentina; Colombia; Venezuela. 
Host Plant: Lantana camara. 

Teleonemia longicornis Champion 

Teleonemia longicornis Champion 1898b, p. 62, pi. 3, fig. 7 [Brazil]. — ■ 
Drake 1930, p. 1. 
Type: Holotype cf; Amazons; Oxford Mus. 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Teleonemia luctuosa (Stal) 

Laccometopus luctuosus Stal 1858, p. 65 [Brazil], 
Tingis {Amaurosterphus) luctuosus: Stal 1868, p. 92. 
Monanthia luctuosa: Walker 1873a, p. 193. 

Teleonemia luctuosa: Stal 1873, p. 132.— Champion 1898b, p. 62, pi. 3, 
fig. 5.— Drake 1935, p. 9 [Paraguay].— Drake and Poor 1937d, p. 
303.— Drake and Hambleton 1938b, p. 52. 
Type: Holotype o"; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil; Paraguay. 
Host Plant: Unrecorded. 



681-552 — 64 25 



378 U.S. NATIONAL MUSEUM BULLETIN 243 

Teleonemia lustrabilis Drake 

Teleonemia lustrabilis Drake 1953c, p. 151 [Dominican Republic]. 
Type: Holotype 9; Constanza, Dominican Republic; MGZ. 
Distribution: Dominican Republic. 
Host Plant: Unrecorded. 

*Teleonemia lutzi Drake 

Teleonemia lutzi Drake 1941b, p. 139 [Paraguay]. 
Type: Holotype d\ macropterous; Horqueta, Paraguay; Drake Coll. 
(USNM). 
Distribution: Paraguay; Brazil. 
Host Plant: Unrecorded. 

*Teleonemia mera Drake and Hambleton 

Teleonemia mera Drake and Hambleton 1942b, p. 76 [Brazil]. 

Type: Holotype o 71 , macropterous; Santarem, Brazil; Drake Coll. 
(USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Teleonemia molinae Drake 

Teleonemia molinae Drake 1940b, p. 243 [Paraguay]. 
Type: Holotype cf, macropterous; Horqueta, Paraguay; Drake Coll. 
(USNM). 
Distribution : Paraguay. 
Host Plant: Unrecorded. 

*Teleonemia monile Van Duzee 

Teleonemia monile Van Duzee 1918, p. 279 [Calif.]. 
Type: Holotype o 71 ; Lundy, Mono County, California, U.S.; Cal. Acad. 
Distribution: U.S. (Calif.). 
Host Plant: Unrecorded. 

*Teleonemia montivaga Drake 

Teleonemia montivaga Drake 1920, p. 52 [Calif.]. — Hurd 1946, p. 448 
[Penstemon]. 
Type: Holotype 9, macropterous; Mount Diablo, California, U.S.; 
Drake Coll. (USNM). 
Distribution: U.S. (Calif.). 
Host Plant: Penstemon sp. 

*Teleonemia morio (Stal) 

Tropidocheila morio Stal 1855b, p. 187 [Brazil], 
Laccometopus morio: Stal 1858, p. 65. 
Tingis (Amaurosterphus) morio: Stal 1868, p. 92. 
Monanthia morio: Walker 1873a, p. 193. 

Teleonemia {Amaurosterphus) morio: Stal 1873, p. 131. — Horvath 1925b, 
p. 219. 



LACE BUGS OF THE WORLD 379 

Teleonemia morio: Champion 1898b, p. 62, pi. 3, fig. 2. — Drake 1922c, 
p. 356; 1930b, p. 25.— Drake and Hambleton 1934, p. 438 [Annona 
squamosa]. — Costa Lima 1936, p. 130. — Drake and Poor 1937d, p. 
302 [Paraguay].— Bondar 1936, p. 51, figs, a, b [Annona cherimolia]. — 
Silva 1956, p. 56, fig. 22. 
Teleonemia moria [sic]: Monte 1938b, p. 131 [Annona reticulata]. 
Type: Sex unknown; Brazil; Stockholm Mus. 
Distribution: Brazil; Paraguay. 

Host Plants: Annona cherimola; Annona reticulata; Annona squamosa. 
Note: Bondar states this species attacks exclusively the new bark of 
branches and twigs. 
Teleonemia multimaculata Monte 

Teleonemia multimaculata Monte 1940e, p. 298 [Brazil]. 
Type: Holotype 9; Fazenda do Bosque, Cordeiro, Limeira, Sao Paulo, 
Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Teleonemia nigrina Champion 

Teleonemia nigrina Champion 1898a, p. 41, pi. 3, figs. 13, 13 a, b [Tex.; 
Mexico; Guatemala].— Uhler 1904, p. 362 [N. Mex.].— Barber 
1906, p. 281; 1922a, p. 17; 1922b, p. 23.— Van Duzee 1914, p. 11 
[Calif.; Rhus laurina]; 1917a, p. 261.— Drake 1918b, pp. 324, 325 
[Ariz.; Kans.; Utah; Mo.; Ark.; Ga.; N.C.; S.C.; Verbena; Sphaeralcea 
angustifolia; Adenostegia pilosa; Adenostegia filifolia; Helenium tenuifolium; 
sugar beets]; 1938a, p. 70.— Blatchley 1926, p. 488, fig. 115.— Hixson 
1942, p. 605, fig. 1 [snapdragon]. — Froeschner 1944, p. 669 [Plantago 
aristata]. — Hurd 1946, p. 448 [Eriogonum]. 
Teleonemia elongata Uhler 1886, p. 22 (nom. nud.). — Smith 1910, 
p. 149.— Van Duzee 1917b, p. 222. 
Type: Sex unknown; Duenas, Guatemala; British Mus. 
Distribution: Mexico; Guatemala; U.S. (Calif.; Ariz.; N. Mex.; Tex.; 
Okla.; Utah; Kans.; Iowa; Mo.; Ark.; Ga.; S.C.; N.C.; Va.; N.J.). 

Host Plants: Adenostegia filifolia; Adenostegia pilosa; Eriogonum sp.; 
Helenium tenuifolium; Plantago aristata; Rhus laurina; Sphaeralcea angustifolia; 
Verbena sp.; sugar beets; snapdragon. 
*Teleonemia notata Champion 

Teleonemia notata Champion 1898a, p. 40, pi. 3, figs. 11, 11 a, b [Mexico; 
Guatemala; Panama]. — Perkins and Swezey 1924, p. 52 [Lantana]. — 
Monte 1942a, p. 97 [Cuba]. — Bruner, Scaramuzza and Otero 1945, 
p. 69 [Duranta repens]. — Hurd 1946, p. 448 [Adenostegia filifolia; 
Adenostegia pilosa]. 



380 U.S. NATIONAL MUSEUM BULLETIN 2 43 

Type: Sex unknown; El Tumbador, Guatemala; British Mus. 

Distribution: Guatemala; Mexico; Panama; Cuba. 

Host Plants: Adenostegia filifolia; Adenostegia pilosa; Duranta repens; 
Lantana sp. 
*Teleonemia novicia Drake 

Teleonemia novicia Drake 1920, p. 53 [Calif.].— Hurd 1946, p. 448 [Ariz.]. 

Type: Holotype 9, macropterous; Oriocle, California, U.S.; Drake Coll. 
(USNM). 

Distribution: U.S. (Calif.; Ariz.). 

Host Plant: Unrecorded. 

Teleonemia ochracea Champion 

Teleonemia ochracea Champion 1898a, p. 36, pi. 3, fig. 3 [Panama]. 
Type: Holotype cf; Volcan de Chiriqui, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

*Teleonemia paraguayana Drake 

Teleonemia par aguayana Drake 1942a, p. 2 [Paraguay]. 
Type: Holotype 9, macropterous; Horqueta, Paraguay; Drake Coll. 
(USNM). 
Distribution : Paraguay. 
Host Plant: Unrecorded. 

*Teleonemia patagonica Drake 

Teleonemia patagonica Drake 1948J , p. 429 [Argentina]. 
Type: Holotype 9, macropterous; Rio Santa, Patagonia, Argentina; 
Drake Coll. (USNM). 
Distribution : Argentina. 
Host Plant: Unrecorded. 

*Teleonemia picta Champion 

Teleonemia picta Champion 1898a, p. 42, pi. 3, figs. 16, 16a [Panama]. 
Type: Sex unknown; Caldera, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 
*Teleonemia pilicornis Champion 

Teleonemia pilicornis Champion 1898a, p. 37, pi. 3, fig. 6 [Guatemala]. 
Type: Holotype o 71 ; Zapote, Guatemala; British Mus. 
Distribution: Guatemala. 
Host Plant: Unrecorded. 
*Teleonemia prolixa (Stal) 

Laccometopus prolixus Stal 1858, p. 65 [Brazil]. 

Monanthia (Tropidochila) sacchari (not Fabricius): Stal 1862, p. 325. 

Tingis (Amaurosterphus) prolixa: Stal 1868, p. 92. 

Monanthia prolixa: Walker 1873a, p. 193. 



LACEBUGS OF THE WORLD 381 

Teleonemia prolixa: Stal 1873, p. 132. — Berg 1884, p. 103 [Argentina]. — 
Champion 1898a, p. 39, pi. 3, figs. 9a, b, 10 [Mexico; Guatemala; 
Panama]; 1898b, p. 62, pi. 3, fig. 6.— Van Duzee 1907, p. 22 [Jamaica; 
British Guiana]. — Pennington 1921, p. 20. — Drake and Bruner 1924a, 
p. 145 [Trinidad].— Drake 1929, p. 35 [Surinam; Ecuador]; 1930a, 
p. 1; 1931b, p. 510 [Peru]; 1932, p. 100 [Barro Colorado]; 1935, p. 9 
[Paraguay]; 1936b, p. 699.— Drake and Hambleton 1934, p. 438; 

1945, p. 357; 1946b, p. 122.— Drake and Poor 1937d, p. 304 [Co- 
lombia]; 1938b, p. 105 [Cinchona].— Monte 1938d, p. 390; 1939b, p. 80 
[Lantana camera]; 1940c, p. 191; 1940d, p. 101 [Venezuela].— Ramos 

1946, p. 25 [Mona].— Silva 1956, p. 58, fig. 23.— Gibson and Carrillo 
1959, p. 19. 

Teleonemia prollixa [sic]: Gowdy 1926, p. 35. 

Type: Holotype c 71 ; Rio de Janeiro, Brazil; Stockholm Mus. 

Distribution: Venezuela; Surinam; British Guiana; Brazil; Paraguay; 
Argentina; Bolivia; Peru; Ecuador; Colombia; Panama; Canal Zone 
(Barro Colorado); Guatemala; Mexico; Trinidad; Jamaica; Mona Island. 

Host Plants: Acacia riparia; Cinchona sp.; Lantana camara. 

*Teleonemia prunellae Drake and Hambleton 

Teleonemia pninellae Drake and Hambleton 1946b, p. 122 [Guatemala; 
Prunella vulgaris]. 
Type: Holotype o 71 , macropterous; Guatemala City, Guatemala; Drake 
Coll. (USNM). 
Distribution: Guatemala. 
Host Plant: Prunella vulgaris. 
*Teleonemia quecha Monte 

Teleonemia quecha Monte 1943a, p. 106, fig. 2 [Peru]. — Drake and 
Hambleton 1944b, p. 121 [Colombia]; 1945, p. 357. 
Type: Holotype 9; Peru; Monte Coll. (Mus. Nacional). 
Distribution: Peru; Colombia; Brazil. 
Host Plant: Unrecorded. 
*Teleonemia rugosa Champion 

Teleonemia rugosa Champion 1898a, p. 37, pi. 3, fig. 4 [Guatemala; 
Panama]. — Drake 1928e, p. 2 [Honduras]. 
Type: Sex unknown; Panzos, Vera Paz, Guatemala; British Mus. 
Distribution: Guatemala; Honduras; Panama; Bolivia. 
Host Plant: Unrecorded. 
*Teleonemia ruthae Monte 

Teleonemia ruthae Monte 1942b, p. 136, fig. 1 [Brazil]. 
Type: Holotype o 71 ; Guaruja, Santos, Sao Paulo, Brazil; Monte Coll. 
(Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 



382 U.S. NATIONAL MUSEUM BULLETIN 243 

*Teleonemia sacchari (Fabricius) 

Acanthia sacchari Fabricius 1794, p. 77. 

Tingis sacchari: Fabricius 1803, p. 126. — Guerin-Meneville 1857, p. 409 

[Cuba; sugar]. 
Monanthia sacchari: Herrich-Schaeffer 1840, p. 85, pi. 173, fig. 533 

[Brazil; Mexico].— Walker 1873a, p. 191. 
Monanthia (Tropidocheila) sacchari: Fieber 1844, p. 76, pi. 6, figs. 22-25 
[Martinique].— Herrich-Schaeffer 1850, p. 152.— Stal 1858, p. 62. 
Tingis (Tropidocheila) sacchari: Stal 1868, p. 92. 

Teleonemia sacchari: Stal 1873, p. 132 [St. Bartholemy]. — Distant 
1888, p. lxxxiii — Uhler 1894a, p. 202 [Grenada]; 1894b, p. 278 — 
Champion 1898b, p. 62 [St. Vincent].— Van Duzee 1907, p. 22 
[Jamaica].— Barber 1914, p. 507 [Fla.]; 1939, p. 371 [Antigua; 
St. Croix; St. Thomas].— Drake 1918b, p. 330; 1926a, p. 86 [Lantana 
camara]; 1931b, p. 510; 1932, p. 100 [Panama].— Wolcott 1923, 
p. 247 [Puerto Rico; Verbesina]. — Blatchley 1926, p. 490. — Monte 
1941e, p. 139 [Calif.].— Blote 1945, p. 89 [Trinidad].— Bruner, 
Scaramuzza, and Otero 1945, p. 98 [Lantana involucrata]. — Box 
1953, p. 37.— Drake and Cobben 1960, p. 79, fig. 79c [St. Eustatius; 
Saba; St. Martin; Lantana canescens]. 
Type: Holotype o", macropterous; "Americae meridionalis Insulis"; 
Copenhagen Mus. 

Distribution: Brazil; Panama; Mexico; U.S. (Calif.; Fla.); Trinidad; 
Cuba; Jamaica; Puerto Rico; Windward Islands (Grenada; Martinique; 
St. Vincent); Virgin Islands (St. Croix; St. Thomas); Leeward Islands 
(Antigua; St. Martin; Saba; St. Eustatius; St. Bartholomy). 

Host Plants: Lantana camara; Lantana canescens; Lantana involucrata; 
Verbesina sp.; sugar. 

*Teleonemia sandersi Drake and Hambleton 

Teleonemia sandersi Drake and Hambleton 1944b, p. 123 [Canal Zone]. — 
Hurd 1946, p. 448 [Honduras]. 
Type: Holotype o", macropterous; Panama Canal Zone; Drake Coll. 
(USNM). 
Distribution: Panama (Canal Zone); Honduras. 
Host Plant: Unrecorded. 

*Teleonemia schildi Drake 

Teleonemia schildi Drake 1940b, p. 242 [Costa Rica]. 
Type: Holotype 9, macropterous; Suiza tur'bla, Costa Rica; Drake 
Coll. (USNM). 
Distribution: Costa Rica. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 383 

*Teleonemia schwarzi Drake 

Teleonemia schwarzi Drake 1918b, pp. 324, 326 [Calif.; Ariz.]; 1941b, 
p. 141 [Beloperone calif ornica], — Hurd 1946, p. 448 [Hymenoclea salsola]. 

Teleonemia sororcula Van Duzee 1923, p. 142 [Baja Calif.]. 
Type: Holotype 9, macropterous; San Diego, California, U.S.; USNM. 
Distribution: U.S. (Calif.; Ariz.); Mexico (Baja Calif.). 
Host Plants: Beloperone calif ornica; Hymenoclea salsola. 

*Teleonemia scrupulosa Stal 

Teleonemia scrupulosa Stal 1873, p. 132 [Brazil; Colombia]. — Distant 
1888, p. lxxxiii.— Champion 1898a, p. 40, pi. 3, figs. 12, 12a [Mexico; 
Guatemala; Panama; Grenada; St. Vincent]. — Barber 1906, p. 281 
[Tex.]; 1914, p. 507 [Fla.].— Van Duzee 1907, p. 22 [Jamaica].— 
Drake 1918b, p. 329 [Haiti; Callirhoe involucrata]; 1926a, p. 86 [Cuba]; 
1926b, p. 376; 1930b, p. 25; 1931b, p. 510; 1935, p. 10 [Paraguay]; 
1956d, p. 108, fig. 3 [Ponape].— Drake and Burner 1924a, p. 145 
[Trinidad].— Blatchley 1926, p. 489, fig. 114 [ebony].— Drake and 
Hambleton 1934, p. 438; 1944b, p. 123; 1945, p. 357.— Costa Lima 
1936, p. 130.— Drake and Poor 1937d, p. 304; 1943, p. 192, fig. 1 — 
Monte 1938b, p. 131; 1940c, p. 191; 1942d, p. 109; 1944b, p. 459 — 
Drake and Frick 1939, p. 199, fig. 1 [Peru; British Guiana; French 
Guiana; Venezuela].— Currie and Fyfe 1939, p. 259.— Khan 1945, 
p. 149. — Bruner, Scaramuzza, and Otero 1945, p. 98. — Cashmore 
and Campbell 1946, p. 26.— Zimmerman 1948, p. 121, figs. 44, 
45. — Fullaway 1951, p. 208 [Xanthium; Lantana montevidensis]; 1958, 
p. 550.— Roonwal 1952, p. 3, fig. 1.— Singh 1953, p. 119.— van der 
Vecht 1953, p. 170 [Java].— Maehler 1955, p. 377 [Myoporum 
sandwicense]. — Silva 1956, p. 59, figs. 24, 25 [Lantana brasiliensis]. — 
Orian 1956, p. 647 [Mauritius].— Drake and Cobben 1960, p. 73, 
figs. 79a, 81a [Aruba; Curacao; Klein Bonaire; Bonaire; Lantana 
canescens; Lippia alba]. — Stusak 1961, p. 77, fig. 5a. 

Teleonemia bifasciata (not Champion): Kirkaldy 1905, p. 216. 

Teleonemia lantanae Distant 1907, p. 60 [Hawaii]. — Kirkaldy 1907, p. 
154 [Lantana camera]; 1908d, p. 190.— Swezey 1924, p. 75, fig. 2 — 
Perkins and Swezey 1924, p. 52.— Simmonds 1929, p. 36, figs. 1-4 
[Fiji].— Drake 1929, p. 35 [Costa Rica].— Fyfe 1935, p. 35; 1937, p. 
182, pi. 1, figs. 1, 2 [Australia].— Van Duzee 1936, p. 225.— Beeson 
and Chatterjee 1940, pp. 42, 48 [Lantana aculeata; India]. 

Teleonemia vanduzeei Drake 1919c, p. 24.— Drake and Bruner 1924a, p. 
144.— Blatchley 1926, p. 489.— Gowdy 1926, p. 35.— Krauss 1953, 
p. 123. 
Type: Unknown. 

Distribution: Peru; Brazil; Paraguay; British Guiana; French Guiana; 
Venezuela; Colombia; U.S. (Tex.; Fla.; Hawaii); Cuba; Jamaica; Trin- 



384 U.S. NATIONAL MUSEUM BULLETIN 24 3 

idad; Haiti; Windward Islands (Grenada; St. Vincent); Netherlands 
Antilles (Aruba; Curagao; Klein Bonaire; Bonaire); Costa Rica; Panama; 
Mexico; Guatemala; Fiji Islands; Greater Sunda Islands (Java); Senyavin 
Islands (Ponape); Australia; India; Mascarene Islands (Mauritius). 

Host Plants: Callirhoe invulucrata; Lantana aculeata; Lantana brasiliensis; 
Lantana camara; Lantana canescens; Lantana montevidensis ; Lippia alba; Myoporum 
sandwicense; Xanthium sp.; ebony. 

Note: Life history studies [Khan; Roonwal; Simmonds; Fyfe]. 

*Teleonemia scrupulosa var. haytiensis Drake 

Teleonemia haytiensis Drake 1920, p. 53 [Haiti]. — Drake and Bruner 

1924a, p. 145; 1924b, p. 155 [Cuba]. 
Teleonemia scrupulosa var. haytiensis: Drake and Frick 1939, p. 201. 
Type: Holotype 9, macropterous; Port au Prince, Haiti; Drake Coll. 
(USNM). 
Distribution: Haiti; Cuba. 
Host Plant: Unrecorded. 

Teleonemia sidae (Fabricius) 

Acanthia sidae Fabricius 1794, p. 77. 

Tingis sidae: Fabricius 1803, p. 126. — Fieber 1844, p. 108. — Walker 

1873a, p. 180 [West Indies]. 
Tingis (Tropidocheila) sidae: Stal 1868, p. 92. 
Monanthia sidae: Walker 1873a, p. 121. 
Teleonemia sidae: Stal 1873, p. 132. 
Tropidocheila sidae: Uhler 1886, p. 22. 
Type: Holotype cf, macropterous; "Americae meridionalis Insulis 
Dom."; Kiel Mus. (Copenhagen Mus.). 
Distribution: Uncertain, possibly West Indies. 
Host Plant: Unrecorded. 

Teleonemia simillima Monte 

Teleonemia simillima Monte 1941a, p. 376, fig. 3 [Brazil]. 

Type: Holotype o*; Puerto America, Rio Putumayo; Monte Coll. (Mus. 
Nacional). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Teleonemia simulans Drake 

Teleonemia simulans Drake 1922c, p. 358 [Argentina]; 19281, p. 72. — 
Drake and Poor 1938b, p. 106.— Monte 1940e, p. 298. 
Type: Holotype 9, macropterous; "El Gran Chaco, 'Bords du Rio 
Tapenaga, Colonia Florencia, S.A.' "; Drake Coll. (USNM). 
Distribution: Argentina. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 385 

*Teleonemia syssita Drake and Cobben 

Teleonemia syssita Drake and Cobben 1960, pp. 67, 75, figs. 79d, 80, 81b 
[St. Eustatius; Saba; St. Martin; Lantana canescens}. 
Type: Holotype cf, macropterous; Cul de Sac, St. Martin; Drake Coll. 
(USNM). 
Distribution: Leeward Islands (St. Eustatius; Saba; St. Martin). 
Host Plant: Lantana canescens. 

*Teleonemia tellus Drake and Hambleton (emendation) 

Teleonemia telluris Drake and Hambleton 1939, p. 154 [Brazil]. 

Type: Holotype cf, macropterous; Chapada, Mato Grosso, Brazil; 
Drake Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Teleonemia teres Drake (emendation) 

Teleonemia teretis Drake 1940b, p. 242 [Brazil]. 
Type: Holotype 9, macropterous; Chapada, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Teleonemia triangularis (Blanchard) 

Tingis triangularis Blanchard 1846, p. 219, pi. 29, fig. 10 [Bolivia]. — 
Stal 1873, p. 134. — Lethierry and Severin 1896, p. 26. — Champion 
1898a, p. 43. 
Laccometopus albilaterus Stal 1858, p. 65 [Brazil]. 
Tingis (Americia) albilatera: Stal 1873, p. 131. 
Monanthia albilatera: Walker 1873a, p. 193. 

Lasiacantha {Americia) albilatera: Lethierry and Severin 1896, p. 18. 
Teleonemia triangularis: Champion 1898b, p. 61. — Drake 1922c, p. 359; 
1935, p. 10 [Paraguay].— Drake and Poor 1937d, p. 302, pi. 36, 
fig. 14.— Drake and Hambleton 1938b, p. 53.— Monte 1940c, p. 190; 
1941e, p. 141 [Argentina].— Silva 1956, p. 64, fig. 26. 
Type: Holotype 9; Province de Chiquitos, Bolivia; Paris Mus. 
Distribution: Bolivia; Paraguay; Argentina; Brazil. 
Host Plant: Unrecorded. 

Teleonemia tricolor (Mayr) 

Monanthia {Gargaphia) tricolor Mayr 1865, p. 442 [Venezuela]. — Walker 
1873a, p. 192. 

Monanthia lanceolata Walker 1873a, p. 194 [Brazil].— Distant 1902b, 
p. 357. 

Gargaphia tricolor: Stal 1873, p. 125. 

Teleonemia albomarginata Champion 1898a, p. 43, pi. 3, figs. 18, 18a 
[Panama].— Drake 1922c, p. 359 [Guatemala]; 1929, p. 35 [Suri- 
nam]; 1931c, p. 226 [Paraguay; Colombia]; 1932, p. 100 [Barro 



386 U.S. NATIONAL MUSEUM BULLETIN 243 

Colorado]. — Drake and Bruner 1924a, p. 145 [Trinidad]. — Drake 
and Hambleton 1934, p. 438.— Drake and Poor 1939b, p. 95 [Ar- 
gentina].— Monte 1940c, p. 190; 1940e, p. 298; 1943a, p. 107; 
1944b, p. 454; 1947a, p. 233. 
Americia albomarginata: Kirkaldy 1905, p. 216 [Peru]. 
Teleonemia {Americia) albomarginata: Horvath 1925b, p. 219. 
Teleonemia spectabilis Drake 1931c, p. 226; 1935, p. 10. — Drake and 

Bondar 1932, p. 87. 
Teleonemia dispersa Drake 1931c, p. 227, fig. 1 [Ecuador]. 
Teleonemia lanceolata: Drake and Hambleton 1938b, p. 52, pi. 9, fig. b 
[Cucurbita moschata]; 1944b, p. 121; 1945, p. 357.— Drake and Poor 
1942, p. 299.— Drake 1948j, p. 430.— Silva 1956, p. 54, fig. 21 
[Sicana odor if era; Sechium edule]. 
Teleonemia tricolor: Drake and Ruhoff 1962c, p. 133. 
Type: Sex unknown; Venezuela; Vienna Mus. 

Distribution: Brazil; Venezuela; Colombia; Surinam; Ecuador; 
Paraguay; Peru; Argentina; Panama; Canal Zone (Barro Colorado); 
Guatemala; Trinidad. 
Host Plants: Cucurbita moschata; Sechium edule; Sicana odorifera. 

*Teleonemia validicornis Stal 

Teleonemia validicornis Stal 1873, p. 132 [Colombia]. — Champion 
1898b, p. 62, pi. 3, fig. 4.— Van Duzee 1901, p. 348 [British Guiana].— 
Drake 1929, p. 35 [Surinam]; 1936b, p. 699 [Argentina].— Drake and 
Bondar 1932, p. 87 [Brazil; Macherium oblongifolium subglabrum]. — 
Costa Lima 1936, p. 130.— Drake and Poor 1937d, p. 303 [French 
Guiana].— Drake and Hambleton 1938b, p. 52.— Monte 1939b, 
p. 80 [Jacaranda paucifoliata]; 1940c, p. 191.— Hurd 1946, p. 448 
[Panama; Macherium oblongifolium].- — Silva 1956, p. 64, fig. 27.— Drake 
and Cobben 1960, p. 73, figs. 78, 79b [Venezuela; Curacao; Lantana 
camara] . 
Type: Holotype c"; Bogota, Colombia; Stockholm Mus. 
Distribution: Colombia; Surinam; French Guiana; British Guiana; 
Brazil; Argentina; Venezuela; Panama; Netherlands Antilles (Curacao). 
Host Plants: Jacaranda paucifoliata; Lantana camara; Macherium oblongi- 
folium; Macherium oblongifolium v. subglabrum. 

♦Teleonemia variegata Champion 

Teleonemia variegata Champion 1898a, p. 42, pi. 3, figs. 15, 15a [Mexico; 
Guatemala].— Barber 1910, p. 38 [Ariz.].— Drake 1918b, p. 328 — 
Perkins and Swezey 1924, p. 52 [Lantana]. — Hurd 1946, p. 449 
[Honduras]. 
Type: Sex unknown; Capetillo, Guatemala; British Mus. 
Distribution: Honduras; Guatemala; Mexico; U.S. (Ariz.). 
Host Plant: Lantana sp. 



LACEBUGS OF THE WORLD 387 

*Teleonemia veneris Drake 

Teleonemia veneris Drake 1939d, p. 527 [Brazil]. 

Type: Holotype 9, macropterous; Belem, Para, Brazil; Drake Coll. 
(USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Teleonemia vidua Van Duzee 

Teleonemia vidua Van Duzee 1918, p. 278 [Calif.]. 

Type: Holotype 9; Keen Camp, San Jacinto Mountains, California, 
U.S.; Cal. Acad. 

Distribution: U.S. (Calif.). 

Host Plant: Unrecorded. 

*Teleonemia vulgata Drake and Hambleton 

Teleonemia vulgata Drake and Hambleton 1940, p. 533 [Brazil]. 

Type: Holotype cf, macropterous; Nictheroy, Est. do Rio, Brazil; 
Drake Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Teleonemia vulsa Drake and Hambleton 

Teleonemia vulsa Drake and Hambleton 1944b, p. 123 [Brazil]. 

Type: Holotype cf, macropterous; Chapada, Brazil; Drake Coll. 
(USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

Genus TIGAVA Stal 

Tigava Stal 1858, p. 63; 1873, pp. 121, 130.— Lethierry and Severin 
1896, p. 22.— Monte 1939b, p. 80; 1941e, p. 143; 1944a, p. 158 — 
Hurd 1946, p. 449.— Drake and Ruhoff 1960a, p. 84. 
Type Species: Tigava praecellens Stal. 

*Tigava bombacis Drake and Poor 

Tigava convexicollis (not Champion): Drake and Hambleton 1935, 
p. 143 [Brazil; Bombax; Chorisia speciosa]. — Monte 1937a, p. 35, 
fig. 5 [Chorisia]. 
Tigava bombacis Drake and Poor 1938a, p. 32.— Monte 1940c, p. 193 — 
Kormilev 1955a, p. 63 [Argentina]. 
Type: Holotype cf , macropterous; Vigosa, Brazil; Drake Coll. (USNM). 
Distribution: Brazil; Argentina. 
Host Plants: Bombax sp.; Chorisia speciosa; Chorisia sp. 



388 U.S. NATIONAL MUSEUM BULLETIN 243 

*Tigava brevicollis Monte 

Tigava brevicollis Monte 1944a, p. 157, fig. 1 [Brazil; Ficus]. 
Type: Holotype cf; Rio de Janeiro, Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Ficus sp. 
*Tigava ceibae Drake and Poor 

Tigava seibae Drake and Poor 1938a, p. 28 [Brazil; Ceiba pentandra]. 

Tigava ceibae (emended): Silva 1956, p. 64, fig. 28. 
Type: Holotype 9, macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil; Paraguay. 
Host Plant: Ceiba pentandra. 

*Tigava convexicollis Champion 

Tigava convexicollis Champion 1897, p. 33, pi. 2, figs. 29, 29a [Panama]. 
Type: Sex unknown; Volcan de Chiriqui, Panama; British Mus. 
Distribution: Panama. 
Host Plant: Unrecorded. 

*Tigava corumbiana Drake 

Tigava corumbiana Drake 1942a, p. 12 [Brazil]. 

Type: Holotype o", macropterous; Corumba, Matto Grosso, Brazil; 
Drake Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Tigava ferruginea Monte 

Tigava jerruginea Monte 1940b, p. 378 [Brazil; Bombax]. 
Type: Holotype cf ; Brazil; Monte Coll. (Mus. Nacional). 
Distribution: Brazil. 
Host Plant: Bombax sp. 

*Tigava gracilis Monte 

Tigava gracilis Monte 1940b, p. 379 [Brazil; Papilionaceae] ; 1941e, p. 
144 [Leguminosae]. 
Type: Holotype o"; Lapinha, Lagoa Santa, Brazil; Monte Coll. (Mus. 
Nacional). 
Distribution: Brazil. 

Host Plants: Leguminosae; Papilionaceae. 
*Tigava graminis Drake and Poor 

Tigava graminis Drake and Poor 1938a, p. 33 [Brazil; Gramineae]. — 
Silva 1956, p. 67. 
Type: Holotype 9, macropterous; Bahia, Brazil; Drake Coll. (USNM) 
Distribution: Brazil. 
Host Plant: Gramineae. 



LACEBUGS OF THE WORLD 389 

*Tigava hambletoni Drake 

Tigava hambletoni Drake 1948d, p. 16 [Peru]. 
Type: Holotype cf, macropterous; Tingo Maria, Peru; USNM. 
Distribution: Peru. 
Host Plant: Unrecorded. 
*Tigava notabilis Drake 

Tigava notabilis Drake 1922c, p. 364, pi. 39, fig. 9 [Brazil]; 1930a, 
p. 1. 
Type: Holotype o 71 ; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil; Bolivia. 
Host Plant: Unrecorded. 
*Tigava praecellens Stal 

Tigava praecellens Stal 1858, p. 63 [Brazil]; 1873, p. 130. — Walker 
1873b, p. 4.— Champion 1898b, p. 61, pi. 3, fig. 1.— Drake and 
Poor 1937d, p. 305 [Bolivia].— Drake and Hambleton 1938a, 
p. 45.— Monte 1939b, p. 81 [Leguminosae] ; 1940c, p. 193 
[Papilionaceae.]. 
Tigava pracellens [sic]: Drake 1931b, p. 510. 
Type: Holotype 9; Rio de Janeiro, Brazil; Stockholm Mus. 
Distribution: Brazil; Bolivia. 
Host Plants: Leguminosae; Papilionaceae. 
*Tigava pulchella Champion 

Tigava pulchella Champion 1897, p. 32, pi. 2, fig. 26 [Mexico]. — Drake 
and Bruner 1924a, p. 146 [Solanum torvutn]. 
Type: Sex unknown; Vera Cruz, Mexico; British Mus. 
Distribution: Mexico; Guatemala; Honduras; Cuba. 
Host Plants: Solanum torvum; avocado. 
*Tigava semota Drake 

Tigava semota Drake 1931a, p. 405 [Brazil]; 1935, p. 13 [Paraguay]. 
Type: Holotype cf, macropterous; Chapada, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil; Paraguay. 
Host Plant: Unrecorded. 
Tigava spatula Monte 

Tigava spatula Monte 1945a, p. 250 [Brazil]. 
Type: Holotype cf; Rio Verde, Goias, Brazil; Monte Coll. (Mus. 
Nacional). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Tigava tingoana Drake 

Tigava tingoana Drake 1948d, p. 17 [Peru]. 
Type: Holotype c?, macropterous; Tingo Maria, Peru; USNM. 
Distribution: Peru. 
Host Plant: Unrecorded. 



390 U.S. NATIONAL MUSEUM BULLETIN 243 

Genus TIGAVARIA Drake 

Tigavaria Drake 1945, p. 99.— Drake and Ruhoff 1960a, p. 84. 
Type Species: Tigava unicarinata Hacker. 

*Tigavaria unicarinata (Hacker) 

Tigava unicarinata Hacker 1929, p. 325, pi. 32, fig. 2 [Queensland]. 
Tigavaria unicarinata: Drake 1945, p. 99. 
Type: Sex unknown; Gold Creek, Queensland, Australia, Queensland 
Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Genus TINGIS Fabricius 

Tingis Fabricius 1803, p. 124— Stal 1874, p. 54— Horvath 1906a, pp. 
14, 61 (key).— Muzik 1907, pp. 50, 58 (key).— Oshanin 1908, p. 429; 
1912, p. 44.— Schumacher 1914, p. 257.— Stichel 1926, pp. 104, 110; 
1935, p. 349; 1938a, p. 406; 1960a, p. 307; 1960b, p. 396; 1960c, p. 
135.— Blatchley 1926, p. 487.— de Seabra 1931, pp. 407, 422 (key).— 
Borner 1935, p. 76.— Guide 1938, pp. 244, 287 (key).— Monte 1939b, 
p. 81; 1941e, p. 146.— China 1943, p. 247— Blote 1945, p.85— Hurd 
1946, p. 463.— Gonzalez 1948, p. 50.— Kiritshenko 1951, pp. 242,248 
(key). — Priesner and Alfieri 1953, p. 64 (key). — Drake and Ruhoff 
1960a, p. 85. 

Phyllontocheila Fieber 1844, p. 59 (in part).— Distant 1903b, p. 135. — 
Drake and Ruhoff 1960a, p. 76. 

Platychilae Fieber 1861, p. 119 (in part; without species). — Drake and 
Ruhoff 1960a, p. 78. 

Monanthia (Platychila) Puton 1879c, p. 107. — Drake and Ruhoff 1960a, 
p. 78. 

Phyllontochila: Lethierry and Severin 1896, p. 17 (in part). 
Type Species: Cimex cardui Linnaeus. 

Subgenus TINGIS (TINGIS) Fabricius 

Tingis (Tingis) Fabricius 1803, p. 124. — Horvath 1906a, p. 71. — Drake 
and Ruhoff 1960a, p. 85. 
Type Species: Cimex cardui Linnaeus. 

*Tingis aemula Drake 

Tingis aemula Drake 1947c, p. 115 [South Australia]. 

Type: Holotype 9, macropterous; Oldea, South Australia, Australia; 
Drake Coll. (USNM). 

Distribution: Australia (South Australia). 

Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 391 

*Tingis americana Drake 

Tingis americana Drake 1922c, p. 366, pi. 39, fig. 11 [Brazil]; 1926d, 
pp. 83, 84; 1930a, p. 1; 1931b, p. 511; 1935a, p. 11 [Paraguay].— 
Monte 1937a, p. 34, fig. 15 [Tecoma]; 1937c p. 72.— Drake and 
Hambleton 1938a, p. 64. — Kormilev 1955a, p. 64 [Argentina]. 
Type: Holotype 9 ; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil; Paraguay; Argentina. 
Host Plant: Tecoma sp. 

*Tingis ampliata (Herrich-Schaeffer) 

Monanthia ampliata Herrich-Schaeffer 1838, p. 62, pi. 127, fig. 397a 
[Czechoslovakia; Austria]. — Mayr 1858, p. 569. — Fieber 1861, p. 120 
[Ukraine]. — Douglas and Scott 1865, p. 252. — Garbiglietti 1869, 
p. 272.— Walker 1873a, p. 183.— Saunders 1875, p. 248.— Reiber 
and Puton 1876, p. 69.— Dubois 1888, p. 121.— d'Antessanty 1890, 
p. 32. — Hiieber 1893, p. 329 [Switzerland; Germany]. — Butler 
1923, p. 209. 
Tingis ampliata: Curtis 1839, tab. 741 [England].— Stal 1874, p. 54 — 
Sahlberg 1878, p. 21— Horvath 1906a, pp. 63, 71 [Hungary; 
Caucasus; Cirsium]. — Reuter 1908, p. 88 [Pinus].— Scholte 1935, 
p. 62, fig. 15 [Netherlands]. — Reclaire 1940, p. 109 [Cirsium arvense]. — 
Cobben 1948, p. 82.— Kiritshenko 1951, p. 249.— Singer 1952, p. 52 
[Cirsium palustre]. — Bator 1953, p. 325, pi. 3, fig. 5. — Southwood and 
Scudder 1956, p. 93, figs. 1, 2a, 3-5, 8-13a— Stusak 1957b, p. 140, 
fig. 15a. — Southwood and Leston 1959, p. 148, pi. 21, fig. 8; figs. 
41, 46.- — Stichel 1960a, p. 311 [Poland; Turkestan; Cirsium lanceo- 
latum] . 
Monanthia {Phyllontocheild) ampliata: Fieber 1844, p. 59, pi. 5, figs. 10-11 

[Verbascum; Italy]. — Herrich-Schaeffer 1850, p. 155. 
Monanthia (Platychila) ampliata: Puton 1879c, p. 108 [France; Carduus]. — 

Saunders 1892, p. 132, pi. 12, fig. 5. 
Phyllonthocheila [sic] ampliata: Reichensperger 1920, p. 62. 
Type: Unknown. 

Distribution: England; Netherlands; France; Germany; Czechoslovakia; 
Poland; Austria; Switzerland; Italy; Hungary; U.S.S.R. (Caucasus; 
Turkestan; Ukraine). 

Host Plants: Carduus sp.; Cirsium arvense; Cirsium lanceolatum; Cirsium 
palustre; Cirsium sp.; Pinus sp.; Verbascum sp. 

Note: Life history studies [Butler; Southwood and Scudder; Stusak]. 
Predators [Southwood and Leston]. 

*Tingis angustata (Herrich-Schaeffer) 

Monanthia angustata Herrich-Schaeffer 1838, p. 61, pi. 127, fig. 397b 
[Germany].— Scholz 1847, p. 119.— Fieber 1861, p. 121 [Ukraine].— 
Garbiglietti 1869, p. 272.— Gredler 1870, p. 75.— Walker 1873a, 



392 U.S. NATIONAL MUSEUM BULLETIN 243 

p. 184.— Chicote 1880, p. 189.— Horvath 1881b, p. xxxiv.— d'Antes- 
santy 1890, p. 32.— Hiieber 1893, p. 333.— Butler 1923, p. 212 
[Algeria]. 
Monanthia {Phyllontocheila) angustata: Fieber 1844, p. 62, pi. 5, fig. 16-18 

[Czechoslovakia; Austria]. — Herrich-Schaeffer 1850, p. 155. 
Catopolatus parallelus Costa 1847b, p. 19 [Italy]; 1847d, p. 255.— Fieber 

1861, p. 383.— Stichel 1960d, p. 119. 
Catoplatus variolosus Costa 1847b, p. 20; 1847d, p. 256, pi. 3, fig. 2 — 

Fieber 1861, p. 383.— Stichel 1960d, p. 119. 
Monanthia brachycera Fieber 1861, p. 121 [Yugoslavia]. 
Monanthia par allela: Garbiglietti 1869, p. 273.— Walker 1873a, p. 187. — 

Bolivar and Chicote 1879, p. 166 [Spain]. 

Monanthia variolosa: Garbiglietti 1869, p. 273.— Walker 1873a, p. 187. 

Tingis angustata: Stal 1874, p. 54.— Horvath 1906a, pp. 63, 73; 1906d, 

p. 2; 1916, p. 9.— Reuter 1908, p. 88.— Stichel 1926, p. 112; 1960a, 

p. 313, fig. 117 [Sicily; Sardinia; Pyrus aria]. — Hoberlandt 1943b, 

pp. 121, 123; 1955, p. 89.— Lindberg 1948, p. 59 [Greece; Cyprus; 

Sorbus cretica]. — Kiritshenko 1951, p. 250. — Mancini 1953b, p. 186; 

1953d, p. 16. — Seidenstiicker 1954, p. 235. — Southwood and Leston 

1959, p. 149. 

Monanthia {Platychila) angustata: Puton 1879c, p. 110 [France; Corsica; 

England].— Saunders 1892, p. 134, pi. 12, fig. 6. 
Monanthia angustata var. sympathica Horvath 1881a, p. 41 [Hungary], 
Monanthia fieberi Puton 1886b, p. 33.— Hiieber 1893, p. 333. 
Phyllontochila angustata: Reuter 1891a, p. 26 [Albania]. 
Phyllontocheila parallela: Horvath 1901, p. 475 [Turkey]. 
Monanthia {Platychila) taurica J akovlev 1903b, p. 193 [Crimea]. 
Type: Sex unknown; Regensburg, Germany; Munich Mus. 
Distribution: England; Spain; France; Corsica; Italy; Sicily; Sardinia; 
Austria; Czechoslovakia; Germany; Hungary; Albania; Greece; Yugo- 
slavia; Cyprus; Turkey; U.S.S.R. (Crimea; Ukraine); Algeria. 

Host Plants: Pyrus aria; Sorbus cretica. 
Tingis angustata var. diminuta Horvath 

Tingis angustata var. diminuta Horvath 1906a, p. 74 [Spain]. 
Type: Sex unknown; Albarracin, Spain; Helsin. Mus. 
Distribution: Spain. 
Host Plant: Unrecorded. 
*Tingis auriculata (Costa) 

Monanthia cardui (not Linnaeus): Herrich-Schaeffer 1838, p. 61, pi. 127, 

fig. b. 
Monanthia {Phyllontocheila) sinuata Fieber 1844, p. 60, pi. 5, figs. 12-15 

[Czechoslovakia; Austria; Sicily; Statice]. 
Catoplatus auriculatus Costa 1847b, p. 20 [Italy]; 1847d, p. 256, pi. 3, 
fig. 3.— Stichel 1960d, p. 119. 



LACEBUGS OF THE WORLD 393 

Monanthia sinnata: Mayr 1858, p. 570 [Hungary]. — Fieber 1861, p. 120 

[France; Ukraine; Corsica]. — Jakovlev 1869, p. Ill [Russia]; 1880b, 

p. 104 [Caucasus].— Garbiglietti 1869, p. 272.— Walker 1873a, p. 

183.— Horvath 1874b, p. 432. 

Monanthia unicolor Garbiglietti 1869, p. 274 [Sardinia], — Walker 1873a, 

p. 188. 
Monanthia (Platychila) auriculata: Puton 1879c, p. 109 [Stachys recta]. 
Monanthia auriculata: Chicote 1880, p. 189 [Spain]. — Puton 1881, p. 123 
[Syria].— d'Antessanty 1890, p. 32.— Hiieber 1893, p. 332.— Mancini 
1949b, p. 36; 1953b, p. 186; 1953d, p. 16. 
Phyllontochila auriculata: Lethierry and Severin 1896, p. 16 [Algeria]. 
Phyllontocheila auriculata: Horvath 1901, p. 475 [Turkey]. 
Tingis auriculata: Horvath 1905b, p. 568; 1906a, pp. 62, 71 [Rumania; 
Armenia; Turkestan]; 1906d, p. 2; 1916, p. 9 [Albania]. — Oshanin 
1908, p. 431 [Switzerland; Portugal; Bulgaria; Yugoslavia; Greece]. — • 
Stichel 1926, p. Ill [Daucus carota; Caucalis daucoides; Germany]; 
1938a, p. 406 [Morocco]; 1960a, p. 310 [Torilis arvensis]. — -de Seabra 
1931, p. 426, figs. 495(3), 497.— Lindberg 1932a, p. 41; 1948, p. 59 
[Cyprus].— Vidal 1937, p. 197.— Hoberlandt 1943b, p. 120; 1955, 
p. 89 [Israel].— Kir itshenko 1951, p. 249.— Seidenstiicker 1954, p. 
235.— Gomez-Menor 1955b, p. 248.— Stusak 1961a, p. 84, figs. 11, 
18.— Drake and Ruhoff 1962a, p. 156. 
Monanthia ? necopina Drake 1919a, p. 420 [Md.]; 1926d, p. 83. 
Tingis necopina: McAtee 1923, p. 145. — Blatchley 1926, p. 487. 
Type: Sex unknown; Naples, Italy; Naples Mus. 

Distribution: Spain; Portugal; France; Corsica; Sardinia; Sicily; Italy; 
Switzerland; Czechoslovakia; Germany; Austria; Hungary; Albania; Yugo- 
slavia; Rumania; Bulgaria; Greece; Turkey, Cyprus; Israel; Syria; U.S.S.R. 
(Armenia; Ukraine; Turkestan; Caucasus); Algeria; Morocco; U.S. (Md.). 
Host Plants: Caucalis daucoides; Daucus carota; Stachys recta; Stachys sp.; 
Statice sp. ; Torilus arvensis. 

Note: Study of eggs [Stusak]. 
*Tingis auriculata var. dauci Horvath 

Tingis auriculata var. dauci Horvath 1905b, p. 568 [Hungary; Rumania; 
Spain; Algeria; Daucus carota]; 1906a, p. 71. — Mancini 1953c, p. 22 
[Italy]. — Stichel 1960a, p. 310 [Sardinia; Czechoslovakia; Iraq; 
Caucalis daucoides]. 
Type: Unknown. 

Distribution: Italy; Hungary; Rumania; Spain; Iraq; Sardinia; Czecho- 
slovakia; Algeria. 
Host Plants: Caucalis daucoides; Daucus carota. 



681-552—64 26 



394 U.S. NATIONAL MUSEUM BULLETIN 243 

Tingis brevicornis (Horvath) 

Phyllontocheila brevicornis Horvath 1902a, p. 595 [Armenia]. 
Tingis brevicornis: Horvath 1906a, p. 78 [Turkmen; Turkestan]. — 
Oshanin 1908, p. 436 [Transcaucasus] . 
Type: Holotype cf; Armenia, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Armenia; Turkestan; Turkmen; Transcau- 
casus). 
Host Plant: Unrecorded. 

*Tingis buddleiae Drake 

Tingis buddleiae Drake 1930e, p. 168 [Luzon; Buddleia asiatica]. — Drake 

and Poor 1937b, p. 402.— Drake and Ruhoff 1962c, p. 133. 
Tingis himalayae Drake 1948g, p. 173 [India]. 
Type: Holotype d", macropterous; Los Banos, Philippine Islands; 
USNM. 
Distribution: Philippine Islands (Luzon); China; India. 
Host Plants: Buddleia asiatica; Vitex trifolia. 

*Tingis capillata Kiritshenko 

Tingis capillata Kiritshenko 1914, p. 194 [Turkestan], 
Type: Sex unknown; Turkestan, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 

*Tingis cardui (Linnaeus) Figures \b, c; 6c, d 

Cimex sp. 660 Linnaeus 1746, p. 207 [Carduus], 

Cimex cardui Linnaeus 1758, p. 443; 1761, p. 247; 1767, p. 718. — 
Houttuyn 1765, p. 343.— de Geer 1773, p. 309, pi. 16, figs. 1-6 — 
Miiller 1774, p. 483.— Schrank 1781, p. 266; 1782, p. 276.— Villers 
1789, p. 486.— Gmelin 1790, p. 2127. 

Acanthia cardui: Fabricius 1775, p. 696; 1781, p. 338; 1787, p. 280; 
1794, p. 77— Panzer 1796, heft 5, tab. 24.— Cederhielm 1798, p. 
270.— Wolff 1801, p. 45, pi. 5, fig. 42.— Schrank 1801, p. 65 — 
Walckenaer 1802, p. 338.— Meixner 1915, p. 270. 

Cimex minutulus Goeze 1778, p. 277. 

Acanthia clavicornis (not Linnaeus): Panzer 1793, heft 3, tab. 24. 

Tingis cardui: Fabricius 1803, p. 125.— Latrielle 1804, p. 252; 1810, 
p. 433.— Fallen 1807, p. 36 [Sweden]; 1829, p. 143.— Le Peletier 
and Serville 1828, p. 653.— Zetterstedt 1828, p. 480; 1840, p. 269 
[Lapland].— Herrich-Schaeffer 1835, p. 58.— Curtis 1839, tab. 
741 [Scotland; England].— Westwood 1840b, p. 120.— Stal 1874, 
p. 54.— Sahlberg 1878, p. 21 [Siberia]; 1920, p. 83.— Reuter 1882b, 
p. 119 [Cirsium lanceolatum]. — Horvath 1906a, pp. 63, 72 [Canary 
Islands; Algeria; Turkey; Armenia; Causasus; Marrubium vulgare]; 
1906d, p. 2.— Oshanin 1908, p. 432 [Morocco; Turkestan].— Lindberg 
1919, p. 42; 1932a, p. 41 [Galactites tomentosa]; 1934, p. 29 [China]; 



LACEBUGS OF THE WORLD 395 

1936b, p. 30.— Stichel 1926, p. 112, fig. 301; 1938a, p. 406 [Albania; 
Yugoslavia]; 1960a, p. 312, fig. 116 [Sicily, Sardinia, Bulgaria, 
Silybum].—de Seabra 1931, p. 426, fig. 498.— Scholte 1935, p. 62, 
fig. 16— Borner 1935, p. 77, fig. 116.— Mancini 1935, p. 9; 1952, 
p. 12; 1953b, p. 186; 1953d, p. 16.— Guide 1938, p. 294, fig.— 
Hoberlandt 1942, p. 125; 1943b, p. 123; 1955, p. 89.— Blote 1945, 
p. 86 [Belgium]. — Cobben 1948, p. 82, fig. 3 [Cirsium vulgare]. — 
Novak and Wagner 1951, p. 70.— Kiritshenko 1951, p. 249.— Bator 
1953, p. 325, pi. 3, fig. 5.— Strawinski 1953, p. 380.— Gomez-Menor 
1955b, p. 248; 1956b, p. 82, fig.— Southwood and Scudder 1956, 
pp. 93, 102, figs. 2b, 6, 7, 13b [Cirsium palustre]. — Stusak 1957b. 
pp. 139, 140, figs. 9, 12a, 15b; 1958, p. 368, fig. 15.— Southwood and 
Leston 1959, p. 149, pi. 5, fig. 11; pi. 18, fig. 6; pi. 21, fig. 11.— Drake 
and Davis 1960, figs. 1, 3, 4, 7-10, 12, 16, 17. 

Monanthia cardui: Burmeister 1835, p. 260. — Herrich-Schaffer 1838, p. 
61, pi. 127, fig. A.— Amyot and Serville 1843, p. 298 [France].— Sahl- 
bergl848, p. 131 [Finland].— Mayr 1858, p. 570 [Austria; Ukraine; 
Hungary; Ligustrum vulgare]. — Fieber 1861, p. 120. — Douglas and 
Scott 1865, p. 251.— Garbiglietti 1869, p. 272 [Italy].— Gredler 
1870, p. 75.— Thomson 1871, p. 398— Horvath 1874b, p. 432; 
1892b, p. 131.— Siebke 1874, p. 22 [Norway].— Saunders 1875, 
p. 248; 1892, p. 133.— Reiber and Puton 1876, p. 69.— Vollenhoven 
1878, p. 272, pi. 9, fig. 5 [Netherlands].— Ferrari 1878, p. 85 — 
Bolivar and Chicote 1879, p. 166 [Spain; Portugal]. — Puton 1886a. 
p. 5 [Tunisia].— Dubois 1888, p. 121.— d'Antessanty 1890, p. 32 — 
Hiieber 1893, p. 330 [Czechoslovakia].— Mason 1898a, p. 140 — 
Kirkaldy 1908e, p. 59 [Carduus crispus]. — Jensen-Haarup 1912, pp. 
155, 156, fig. 98b.— Butler 1923, p. 210 [Wales; Ireland; Madeira; 
Pinus austriaca; Pinus sylvestris; Serratula]. 

Monanthia (Phyllontocheila) cardui: Fieber 1844, p. 61, pi. 5, figs. 1-8 
[Carduus nutans; Carduus acanthoides]. — Herrich-Schaeffer 1850, p. 
155. — Flor 1860, p. 345 [Estonia; Latvia; Denmark; Poland; Iran; 
Germany]. 

Catoplatus cardui: Costa 1847a, p. 23; 1847c, p. 163. 

Monanthia cardni [sic]: Frey-Gessner 1865, p. 231 [Switzerland]. 

Monanthia (Platychila) cardui: Puton 1879c, p. 108 [Corsica]. — Mason 
1898b, p. 210 [Carduus lanceolatus]. 

Phyllontochila cardui: Reuter 1891a, p. 26 [Greece]. 

Phyllonthocheila [sic] cardui: Reichensperger 1920, p. 62. 

Tingis montana Lindberg 1932a, p. 42, pi. 2, fig. 5 (fig. only). — Drake 
and Runoff 1962c, p. 134. 
Type: Unknown. 

Distribution: Ireland; Scotland; Wales; England; Portugal; Spain; 
France; Belgium; Netherlands, Germany; Czechoslovakia; Switzerland; 



396 U.S. NATIONAL MUSEUM BULLETIN 243 

Austria; Hungary; Italy; Corsica; Sardinia; Sicily; Yugoslavia; Albania; 
Bulgaria; Greece; Poland; Denmark; Norway; Sweden; Finland; (Lap- 
land); U.S.S.R. (Estonia; Latvia; Ukraine; Caucasus; Armenia; Turkestan; 
Siberia); Turkey; Iran; Tunisia; Algeria; Morocco; Canary Islands; 
Madeira Island; China. 

Host Plants: Carduus acanthoides ; Carduas crispus; Carduus lanceolatum; 
Carduus nutans; Carduus sp.; Cirsium lanceolatum; Cirsium palustre; Cirsium 
vulgar e; Galactites tomentosa; Marrubium vulgar e; Pinus austriaca; Pinus sylvestris; 
Serratula sp.; Silybum sp. 

Note: Life history studies [Southwood and Scudder; Stusak]. 
*Tingis cardui var. cognata (Fieber) 

Monanthia cognata Fieber 1861, p. 121 [Corsica]. — Garbiglietti 1869, 

p. 272. 
Monanthia (Platychila) cardui var. cognata: Puton 1879c, p. 109 [France]. 
Tingis cardui var. cognata: Horvath 1906a, p. 73 [Italy; Algeria]. — 
Oshanin 1908, p. 433 [Russia].— Lindberg 1932a, p. 41 [Spain].— 
Hoberlandt 1943b, pp. 121, 123 [Czechoslovakia].— Blote 1945, p. 86 
[Tunisia]. — Mancini 1952, p. 13 [Turkey; Iran; Caucasus; Turke- 
stan]. 
Tingis auriculata var. cognata: Vidal 1937, p. 197 [Morocco]. 
Type: Sex unknown; Corsica; Vienna Mus. 

Distribution: Spain; France; Corsica; Italy; Czechoslovakia; Turkey; 
Iran; U.S.S.R. (Caucasus; Turkestan); Tunisia; Morocco; Algeria. 
Host Plant: Unrecorded. 

*Tingis cardui var. maderensis (Reuter) 

Monanthia (^Platychila) cardui var. maderensis Reuter 1890b, p. 262 

[Madeira]. 
Tingis cardui var. maderensis: Horvath 1906a, p. 73. — China 1938, 
p. 20.— Lindberg 1960a, p. 13 [La Palma].— Stichel 1960a, p. 312. 
Type: Sex unknown; Madeira; Berlin Mus. 
Distribution: Madeira; Canary Islands (La Palma). 
Host Plant: Unrecorded. 

*Tingis colombiana Drake 

Tingis colombiana Drake 1929, p. 35, fig. 1 [Colombia]. — Drake and 
Hambleton 1938a, p. 63 [Brazil].— Monte 1938b, p. 127 [Bignoni- 
aceae]. — Drake and Davis 1960, fig. 47. 
Type: Holotype o 71 ; Sabanilla, Colombia; Hamburg Mus. 
Distribution: Colombia; Brazil. 
Host Plant: Bisrnoniaceae. 



LACEBUGS OF THE WORLD 397 

*Tingis comosa (Takeya) 

Dictyonota comosa Takeya 1931, p. 66, pi. 7, fig. 1; pi. 8, figs. 1-3 

[Japan] . — Drake and Hsiung 1936, p. 288. 
Tingis modosa Drake 1937b, p. 593 [China]. 

Tingis comosa: Drake 1948a, p. 2. — Takeya 1951a, p. 15; 1953a, p. 
2. — Drake and Maa 1953, p. 91 [Artemesia]. 
Type: Holotype o 71 ; Hikosan, Prov. Buzen, Kyushu, Japan; Kyushu 
Univ. 
Distribution: Japan; China; India. 
Host Plant: Artemesia sp. 

*Tingis consaepta Drake and Poor 

Tingis consaepta Drake and Poor 1939c, p. 205 [India]. 
Type: Holotype 9, macropterous; Ayur, North Salem, Madras, India; 
Drake Coll. (USNM). 
Distribution: India. 
Host Plant: Unrecorded. 
*Tingis coomani Drake 

Tingis coomani Drake 1947d, p. 227, fig. [Viet-Nam]; 1947h, no. 70, 
(color fig.). 
Type: Holotype o 71 ; Tonkin, Hoabinh, Indochina; Heude Mus. 
Distribution: Viet-Nam. 
Host Plant: Unrecorded. 
*Tingis corumbiana Drake 

Tingis corumbiana Drake 1926d, pp. 83, 84 [Brazil]. 
Tingis corubiana [sic]: Drake 1930a, p. 1. 
Type: Holotype o 71 ; Corumba, Matto Grosso, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Tingis crispata (Herrich-Schaeffer) 

Derephysia crispata Herrich-Schaeffer 1838, p. 72, pi. 128, figs. 399, a-c 

[Hungary]. 
Monanthia (Phyllontocheila) crispata: Fieber 1844, p. 66, pi. 5, figs. 

28-30.— Herrich-Schaeffer 1850, p. 154. 
Monanthia crispata: Fieber 1861, p. 384.— Garbiglietti 1869, p. 273 

[Italy].— Horvath 1874b, p. 432.— Hiieber 1893, p. 336. 
Phyllantocheila [sic] crispata: Costa 1863, p. 6. 
Monanthia (Platychila) crispata: Puton 1879c, p. 110 [France]. 
Lasiotropis crispata: Lethierry and Severin 1896, p. 18. 
Tingis crispata: Horvath 1906a, pp. 64, 75 [Caucasus; Bulgaria; 
Rumania; Austria; Artemisia vulgaris]; 1906d, p. 2. — Oshanin 1908, 
p. 434 [Turkestan].— Stichel 1926, p. 112 [Germany]; 1960a, p. 313, 
fig. 118 [Switzerland; Artemisia campestris; Artemisia absinthium]. — 



398 U.S. NATIONAL MUSEUM BULLETIN 243 

Ross and Hedicke 1927, p. 92.— Hoberlandt 1943b, pp. 121, 123 
[Czechoslovakia]. — Blote 1945, p. 86 [Tunisia]. — Mancini 1949b, 
p. 37.— Kiritshenko 1951, p. 250.— Putshkov 1960, p. 303. 
Type: Sex unknown; Hungary; Munich Mus. 

Distribution: France; Germany; Czechoslovakia; Austria; Switzerland; 
Italy; Hungary; Rumania; Bulgaria; U.S.S.R. (Caucasus; Turkestan); 
Tunisia. 
Host Plants: Artemisia absinthium; Artemisia campestris; Artemisia vulgaris. 
Note: Life history study [Putshkov]. 
Tingis crispata var. addita Horvath 

Tingis crispata var. addita Horvath 1911b, p. 584 [Rumania]. 
Type: Holotype 9; Comana, Rumania; Hungarian Mus. 
Distribution: Rumania. 
Host Plant: Unrecorded. 
Tingis demissa Horvath 

Tingis demissa Horvath 1906a, p. 78 [Turkey]. — Hoberlandt 1955, p. 
90.— Stichel 1960a, p. 315 [Cirsium]. 
Type: Holotype 9; Konia, Turkey; deposition unknown. 
Distribution: Turkey. 
Host Plant: Cirsium sp. 
*Tingis deserticola Horvath 

Tingis deserticola Horvath 1906a, p. 71 [Turkestan]. 
Type: Sex unknown; Tshu, Turkestan, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 
*Tingis drakei Hacker 

Tingis drakei Hacker 1929, p. 328, pi. 33, fig. 8 [Queensland]. — Drake 
1961b, p. 110.— Drake and RuhofF 1962b, p. 250 [Lord Howe]. 
Type: Sex unknown; South Pine River, Australia; Queensland Mus. 
Distribution: Australia (Queensland; Lord Howe Island). 
Host Plant: Unrecorded. 
*Tingis elongata (Fieber) 

Monanthia elongata Fieber 1861, p. 121 [Yugoslavia]. — Walker 1873a, p. 

184.— Chicote 1880, p. 189 [Spain]. 
Monanthia par allela (not Costa): Puton 1879c, p. 110. 
Phyllontochila elongata: Lethierry and Severin 1896, p. 17 [Algeria]. 
Phyllontocheila fallax Horvath 1902a, p. 596. 
Phyllontocheila prolixa Horvath 1902a, p. 597. 

Tingis elongata: Horvath 1906a, p. 79, pi. 1, fig. 8.— Vidal 1937, p. 197 
[Morocco]. — Lindberg 1948, p. 59 [Juniperus joetidissima; Cyprus]. — 
Stichel 1960a, p. 315.— Drake and RuhofF 1962a, p. 156. 
Type: Sex unknown; Serbia; Vienna Mus. 
Distribution: Spain; Yugoslavia; Cyprus; Algeria; Morocco. 
Host Plant: Juniperus Joetidissima. 



LACEBUGS OF THE WORLD 399 

tTingis florissantensis Cockerell 

Tingis florissantensis Cockerell 1914, p. 719, fig. 4 [Colo.; Miocene].— 
Drake and RuhofT 1960a, p. 11. 
Type: Sex undeterminable; fossil; Miocene shales; Florissant, Colorado, 
U.S.; deposition unknown. 
Distribution: Fossil. 

Tingis foleyi Bergevin 

Tingis foleyi Bergevin 1929, p. 115, fig. 2 [Algeria]. 
Type: Holotype 9; l'Oued Ilaman, Algeria; Paris Mus. 
Distribution: Algeria. 
Host Plant: Unrecorded. 

Tingis fuentei Horvath 

Tingis fuentei Horvath 1906a, p. 76 [Spain].— Wagner 1958, p. 240 
[France; Centaur ea solstitialis]. — Stichel 1960a, p. 314. 
Type: Holotype 9; Fuencaliente, Spain; Hungarian Mus. 
Distribution: Spain; France. 
Host Plant: Centaur ea solstitialis. 

*Tingis grisea Germar 

Tingis grisea Germar 1835, fasc. 15, tab. 13 [Germany]. — Herrich- 
Schaeffer 1835, p. 58.— Horvath 1906a, pp. 65, 77 [Yugoslavia].— 
Oshanin 1908, p. 435 [Turkestan; Transcaucasus].— Stichel 1926, 
p. 112 [Centaurea rhenana]; 1960a, p. 314 [Morocco]. — Hoberlandt 
1942, p. 125; 1943a, p. 118; 1943b, p. 120; 1955, p. 90 [Turkey; 
Israel].— Stusak 1957a, p. 23, figs. 6-8; 1959b, p. 181, figs. 1-8; 
1961a, p. 77, fig. 5g.— Putshkov 1960, p. 304. 
Monanthia grisea: Herrich-Schaeffer 1838, pp. 52, 60, pi. 122d [Czecho- 
slovakia].— Scholz 1847, p. 119.— Mayr 1858, p. 570 [Rumania; 
Hungary].— Fieber 1861, pp. 120, 383.— Walker 1873a, p. 183 
[England].— Horvath 1874b, p. 432.— Jakovlev 1876b, p. 67 [Russia]; 
1880b, p. 106 [Caucasus].— Pu ton 1874a, p. 226; 1881, p. 123 
[Syria].— Rey 1893, p. 97.— Hiieber 1893, p. 336. 
Monanthia {Phyllontocheila) grisea: Fieber 1844, p. 64, pi. 5, figs. 25-27 

[Austria; Centaurea paniculata]. — Herrich-Schaeffer 1850, p. 154. 
Monanthia pallida Garbiglietti 1869, p. 273 [Italy].— Walker 1873a, 

p. 187. 
Lasiotropis grisea: Stal 1874, p. 55. 

Monanthia (Platychila) grisea: Puton 1879c, p. 110 [France]. 
Type: Germany; sex and deposition of type unknown. 
Distribution: England; France; Germany; Czechoslovakia; Austria; 
Italy; Hungary; Yugoslavia; Rumania; Turkey; Syria; Israel; U.S.S.R. 
(Caucasus; Transcaucasus; Turkestan); Morocco. 



400 U.S. NATIONAL MUSEUM BULLETIN 2 43 

Host Plants: Centaurea paniculata; Centaur ea rhenana. 

Note: Life history studies [Putshkov; Stusak 1959b]. Study of eggs 
[Stusak 1957a; 1961a]. 
*Tingis hackeri Drake 

Tingis hackeri Drake 1947c, p. 118 [Queensland]. 

Type: Holotype c 71 , macropterous; National Park, Queensland, Aus- 
tralia; Drake Coll. (USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

*Tingis hurdae Drake 

Tingis hurdae Drake 1947c, p. 113, fig. 25 [Queensland]; 1961b, p. 110. 

Type: Holotype o 71 , macropterous; Queensland, Australia; Drake Coll. 
(USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

*Tingis impensa Drake 

Tingis impensa Drake 1947c, p. 112, fig. 24a [Tasmania]. 
Type: Holotype 9, macropterous; Tasmania, Australia; Drake Coll. 
(USNM). 
Distribution: Australia (Tasmania). 
Host Plant: Unrecorded. 

Tingis laetabilis (Horvath) 

Phyllontocheila laetabilis Horvath 1903a, p. 77 [Algeria]. 
Tingis laetabilis: Horvath 1906a, p. 75. 
Type: Holotype o 71 ; Ain-Sefra, Algeria; Hungarian Mus. 
Distribution: Algeria. 
Host Plant: Unrecorded. 

*Tingis lanigera (Puton) 

Monanthia lanigera Puton 1886a, p. 5 [Algeria]. 
Monanthia {Platychila) lanigera: Puton 1886a, p. 16. 
Phyllontochila lanigera: Lethierry and Severin 1896, p. 17 [Tunisia]. 
Tingis lanigera: Horvath 1906a, p. 75. — Lindberg 1932a, p. 41 [Morocco; 
Populus nigra; Ephedra']. — Vidal 1937, p. 197. 
Type: Sex unknown; Philippeville, Algeria; Paris Mus. 
Distribution: Algeria; Tunisia; Morocco. 
Host Plants: Ephedra sp.; Populus nigra. 

*Tingis lasiocera Matsumura 

Lasiotropis grisea (not Germar): Matsumura 1905, p. 39. 
Tingis lasiocera Matsumura 1908, p. 148, fig.; 1930, p. 163, pi. 14, fig. 20. 
Tingis lasicera [sic]: Matsumura 1931, p. 1203, fig. 
Tingis laciocera [sic]: Takeya 1933, p. 37, pi. 3, fig. 4, text fig. 4 [Japan; 
Korea; Serratura atriplicifolia]; 1951a, p. 16. 



LACEBUGS OF THE WORLD 401 

Type: Unknown. 
Distribution: Japan; Korea. 
Host Plant: Serratura atriplicifolia. 

Tingis leptochila Horvath 

Tingis leptochila Horvath 1906a, p. 74 [Turkestan]. 
Type: Sex unknown; Turkestan, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 
Tingis neotropicalis Monte 

Tingis neotropicalis Monte 1940e, p. 283 [Brazil; Byrsonima]. — Silva 
1956, p. 67. 
Type: Holotype c? 1 ; Agua Preta, Baia, Brazil; Monte Coll. (Mus. 
Nacional). 
Distribution: Brazil. 
Host Plant: Byrsonima sp. 
Tingis nyogana Drake 

Tingis nyogana Drake 1955c, p. 106 [Cameroun]. 
Type: Holotype 9; Case du Nyong, French Cameroons; Copenhagen 
Mus. 
Distribution: Cameroun. 
Host Plant: Unrecorded. 
fTingis obscura Heer 

Tingis obscura Heer 1853, p. 74, pi. 13, fig. 15 [Yugoslavia]. — Giebel 
1856, p. 363.— Scudder 1891, p. 449 [Horizon: Mayencian 
(Tertiary)].— Drake and Ruhoff 1960a, p. 11. 
Type: Sex undeterminable; fossil; Radoboj, Croatia; deposition unknown. 
Distribution: Fossil. 
*Tingis perkensi Drake 

Tingis perkensi Drake 1947c, p. 117 [Queensland]. 
Type: Holotype o 71 , macropterous; National Park, Queensland, Aus- 
tralia; Drake Coll. (USNM). 

Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Tingis populi Takeya 

Tingis populi Takeya 1932, p. 10, pi. 1, fig. 2 [Korea; Japan; Taiwan; 
Populus; Salix]; 1951a, p. 16 [Populus tremula; Populus nigra]; 1953a, 
p. 2— Saito 1933, p. 7 [Salix koreensis].— Drake and Maa 1953, p. 91 — 
Maa 1957, pp. 117, 131. 
Nobarnus hqffmani Drake 1938b, p. 195 [China]. 
Type: Holotype o 71 ; Taikyu, Korea; Kyushu Univ. 
Distribution: Japan; China; Korea; Taiwan. 

Host Plants: Populus nigra; Populus tremula; Populus sp.; Salix koreensis; 
Salix sp. 



402 U.S. NATIONAL MUSEUM BULLETIN 243 

*Tingis rotundicollis (Jakovlev) 

Monanthia {Platychila) rotundicollis Jakovlev 1883, p. 105 [Dagestan]. 
Phyllontochila rotundicollis: Lethierry and Severin 1896, p. 17 [Caucasus]. 
Tingis rotundicollis: Horvath 1906a, p. 76 [Yugoslavia; Greece; Bul- 
garia; Rumania; Turkey]; 1916, p. 9. — Stichel 1926, p. 112; 1938a, 
p. 406 [Albania]; 1960a, p. 314.— Lindberg 1948, p. 59 [Cyprus].— 
Kiritshenko 1951, p. 250.— Mancini 1952, p. 13 [Italy]; 1953b, p. 
186; 1953d, p. 16; 1956a, p. 192.— Seidenstiicker 1954, p. 235 — 
Hoberlandt 1955, p. 90 [Transcaucasus]. 
Type: Holotype 9, macropterous; Derbent, Dagestan, Russia; Lenin- 
grad Inst. 

Distribution: Italy; Greece; Cyprus; Bulgaria; Yugoslavia; Rumania; 
Albania; Turkey; U.S.S.R. (Caucasus; Transcaucasus; Dagestan). 
Host Plant: Unrecorded. 

*Tingis silvacata Drake 

Tingis silvacata Drake 1926d, p. 83 [Brazil].— Monte 1938b, p. 127 
[Davilla rugosa]. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plant: Davilla rugosa. 

Tingis similis (Douglas and Scott) 

Monanthia similis Douglas and Scott 1869, p. 259. — Walker 1873a, p. 

187 [England]. 
Monanthia ampliata var. similis: Saunders 1875, p. 248. 
Monanthia {Platychila) ampliata var. similis: Saunders 1892, p. 132. 
Tingis similis: Horvath 1906a, p. 72. — Stichel 1938a, p. 406 [Austria]; 
1960a, p. 311 [Czechoslovakia]. 
Type: Unknown. 

Distribution: England; Austria; Czechoslovakia. 
Host Plant: Unrecorded. 

Note: Douglas and Scott state "Two specimens taken by Mr. Wollaston, 
but the place and date of capture are not recorded." 

Tingis sinuaticollis (Jakovlev) 

Monanthia {Platychila) sinuaticollis Jakovlev 1883, p. 103 [Dagestan]. 
Tingis sinuaticollis: Horvath 1906a, p. 71. — Kiritshenko 1951, p. 249. — 
Stichel 1960a, p. 311. 
Type: Holotype 9, macropterous; Derbent, Dagestan, Russia; Leningrad 
Inst. 
Distribution: U.S.S.R. (Dagestan). 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 403 

*Tingis strictula (Puton) 

Monanthia (Platychila) strictula Puton 1878, p. lxvi [Algeria]. 

Monanthia striculata [sic]: Chicote 1880, p. 189 [Spain]. 

Lasiotropis strictula: Lethierry and Severin 1896, p. 18. 

Tingis strictula: Horvath 1906a, p. 77. — Gomez-Menor 1955b, p. 248. — 

Stichel 1960a, p. 315.— Drake and Runoff 1962c, p. 134. 
Tingis montana Lindberg 1932a, p. 42, pi. 2, fig. 5 [Spain]. — Stichel 
1960a, p. 315. 
Type: Sex unknown; Oran, Algeria; Vienna Mus. 
Distribution: Algeria; Spain. 
Host Plant: Unrecorded. 

Tingis stupidula Horvath 

Tingis stupidula Horvath 1906a, p. 77 [Syria]. 
Type: Holotype 9; Baalbeck, Syria; Hungarian Mus. 
Distribution: Syria. 
Host Plant: Unrecorded. 

Tingis suavis (Horvath) 

Phyllontocheila suavis Horvath 1902a, p. 595 [Turkmen]. 
Monanthia (Lasiotropis) longipennis Jakovlev 1903a, p. 3 [Transcapia]. 
Tingis suavis: Horvath 1906a, p. 78. 
Type: Sex unknown; Beush-berma, Turcomania, Russia; Hungarian 
Mus. 

Distribution: U.S.S.R. (Turkmen; Transcapia). 
Host Plant: Unrecorded. 

Tingis tecomae Monte 

Tingis tecomae Monte 1940b, p. 377 [Brazil; Tecoma]. 
Type : Holotype o 71 ; Belo Horizonte, Brazil ; Monte Coll. (Mus. Nacional) . 
Distribution: Brazil. 
Host Plant: Tecoma sp. 

*Tingis teretis Drake 

Tingis teretis Drake 1947c, p. 119 [South Australia]. 

Type: Holotype cf, macropterous; Oldea, South Australia, Australia; 
Drake Coll. (USNM). 

Distribution: Australia (South Australia) . 

Host Plant: Unrecorded. 

*Tingis tonkinana Drake 

Tingis tonkinana Drake 1947d, p. 228 [Viet-Nam]. 
Type: Holotype o"; Tonkin, Hoabinh, Indochina; Heude Mus. 
Distribution: Viet-Nam. 
Host Plant: Unrecorded. 



404 U.S. NATIONAL MUSEUM BULLETIN 243 

Tingis toxopeusi Drake 

Tingis toxopeusi Drake 1960, p. 362, fig. 14 [New Guinea]. 
Type: Holotype 9; Neth. Ind. American, New Guinea Expedition 1939, 
top camp, 2,100 m.; Leiden Mus. 
Distribution: New Guinea. 
Host Plant: Unrecorded. 

*Tingis veteris Drake 

Phyllontocheila ampliata (not Herrich-Schaeffer) : Matsumura 1905, p- 

33, pi. 19, fig. 11; 1908, p. 148, fig.; 1930, p. 158, pi. 14, fig. 11. 
Tingis ampliata (not Herrich-Schaeffer): Matsumura 1931, p. 1203, 
fig. — Takeya 1951a, p. 16 (in part) [Japan; Siberia; Cirsium; 
Cardiacs]. — Esaki 1954, p. 237, figs. 6, 13. 
Tingis (Tropidocheila) veteris Drake 1942a, p. 13 [China]. 
Tingis veteris: Drake 1947d, p. 228 [Taiwan]. — Takeya 1951a, p. 17. 
Type: Holotype d% macropterous; Kianzu Province, China; Drake 
Coll. (USNM). 
Distribution: China; Japan; Taiwan; U.S.S.R. (Siberia). 
Host Plants: Carduus sp.; Cirsium sp. 

tTingis sp. Berendt 

Tingis sp. Berendt 1845, p. 55 [fossil, amber]. — Scudder 1891, p. 449 
[Horizon: Ligurian]. — Drake and Ruhoff 1960a, p. 11. 
Type: Sex undeterminable; fossil in Prussian amber; deposition unknown. 
Distribution: Fossil. 

tTingis sp. Hope 

Tingis sp. Hope 1847, p. 252 [France]. — Scudder 1891, p. 449 [Horizon: 
Ligurian]. — Drake and Ruhoff 1960a, p. 11. 
Type: Sex undeterminable; fossil; Aix, France; deposition unknown. 
Distribution: Fossil. 

tTingis sp. Scudder 

Tingis sp. Scudder 1881, p. 292 [Colo.]; 1891, p. 449 [Horizon: Oligo- 
cene]. — Drake and Ruhoff 1960a, p. 11. 
Type: Sex undeterminable; fossil; Florissant, Colorado, U.S.; deposition 
unknown. 
Distribution: Fossil. 

tTingis sp. Serres 

Tingis sp. Serres 1829, p. 227 [France]. — Scudder 1891, p. 449 [Horizon: 

Ligurian (Tertiary)]. — Drake and Ruhoff 1960a, p. 11. 
Phyllontocheila sp. Scudder 1890, p. 357. 
Type: Sex undeterminable; fossil; Aix, France; deposition unknown. 
Distribution: Fossil. 



LACEBUGS OF THE WORLD 405 

Subgenus TINGIS (CAENOTINGIS) Drake 

Tingis (Caenotingis) Drake 1928h, p. 283. — Drake and Ruhoff 1960a, 
p. 85. 
Type Species: Tingis {Caenotingis) beesoni Drake. 
*Tingis (Caenotingis) beesoni Drake 

Tingis (Caenotingis) beesoni Drake 1928h, p. 283 [Burma; Gmelina 

arboreal. — Singh 1953, p. 119. 
Tingis beesoni: Drake and Poor 1936b, p. 146. — Mathur 1955, p. 248, 
figs. 1-6. 
Type: Holotype o 71 , macropterous; North Toungoo, Burma; Drake Coll. 
(USNM). 
Distribution: Burma. 
Host Plant: Gmelina arbor ea. 

Subgenus TINGIS (LASIOTROPIS) Stal 

Lasiotropis Stal 1874, p. 55. — Lethierry and Severin 1896, p. 18.— 

Blote 1945, p. 87. 
Tingis {Lasiotropis): Horvath 1906a, p. 69. — Oshanin 1908, p. 429; 
1912, p. 44.— Stichel 1926, pp. 110, 111 (key); 1935, p. 349; 1960a, 
p. 307.— de Seabra 1931, pp. 423, 425— Borner 1935, p. 77 — 
Guide 1938, pp. 286, 287, 288 (key).— China 1943, p. 247.— 
Priesner and Alfieri 1953, p. 64.— Drake and Ruhoff 1960a, p. 85. 
Type Species: Monanthia (Platychila) trichonota Puton. 
Tingis (Lasiotropis) aetheria Drake and Ruhoff 

Tingis (Lasiotropis) wollastoni China 1938, p. 20, fig. 3 [Madeira]. 
Tingis aetheria Drake and Ruhoff 1960c, p. 32. 
Type: Sex unknown; Dezerta Grande, Madeira; British Mus. 
Distribution: Madeira Island. 
Host Plant: Unrecorded. 
*Tingis (Lasiotropis) ajugarum (Frey-Gessner) 

Monanthia ajugarum Frey-Gessner 1872, p. 22, pi. 1, fig. 4 [Switzer- 
land].— d'Antessanty 1890, p. 32.— Horvath 1892b, p. 131. 
Monanthia (Platychila) ragusana (not Fieber): Puton 1879c, p. 112 

[France; Ajuga chamaepytis; Ajuga genevensis]. 
Tingis (Lasiotropis) ajugarum: Horvath 1906a, p. 70 [Italy; Yugo- 
slavia; Algeria].— Stichel 1926, p. Ill; 1960a, p. 390 [Greece].— 
Ramade 1960, p. 221 [Ajuga iva]. 
Tingis ajugarum: Reuter 1908, p. 88. — Lindberg 1932a, p. 41 [Mo- 
rocco].— Mancini 1949b, p. 36 [Sicily]; 1953d, p. 16.— Novak and 
Wagner 1951, p. 70. 
Type: Switzerland; sex and deposition of type unknown. 
Distribution: Switzerland; France; Italy; Sicily; Greece; Yugoslavia; 
Algeria; Morocco. 
Host Plants: Ajuga chamaepytis; Ajuga genevensis; Ajuga iva. 



406 U.S. NATIONAL MUSEUM BULLETIN 243 

*Tingis (Lasiotropis) beieri Drake 

Tingis beieri Drake 1935, p. 11 [Paraguay; Brazil]. — Monte 1937a, 
p. 33, fig. 14 [Cordia]; 1939b, p. 81 [Croton] .—Drake and Hambleton 
1938a, p. 64.— Singh 1953, p. 119.— Kormilev 1955a, p. 64 [Argen- 
tina; Bolivia]. 
Type: Holotype o 71 ; S. Bernardino, Paraguay; Vienna Mus. 
Distribution: Paraguay; Argentina; Brazil; Bolivia. 
Host Plants: Cordia sp.; Croton sp. 

*Tingis (Lasiotropis) ciliaris (Puton) 

Monanthia (Platychila) ciliaris Puton 1879a, p. lix [Caucasus]. 
Monanthia {Platychila) balassogloi J akovlev 1880b, p. 105. 
Tingis (Lasiotropis) ciliaris: Horvath 1906a, p. 70 [Syria; Hungary; 
Greece; Turkey]. — Oshanin 1908, p. 430 [Crimea]. — Kiritshenko 
1951, p. 249— Seidenstiicker 1954, p. 235.— Hoberlandt 1955, 
p. 89 [Transcaucasus]. — Stichel 1960a, p. 309 [Bulgaria]. 
Type: Sex unknown; Caucasus, Russia; Paris Mus. 
Distribution: Bulgaria; Hungary; Greece; Syria; Turkey; U.S.S.R. 
(Caucasus; Crimea; Transcaucasus). 

Host Plant: Unrecorded. 
Tingis (Lasiotropis) denudata Horvath 

Tingis denudata Horvath 1906a, p. 75 [Algeria; Egypt]. 
Tingis (Lasiotropis) denudata: Priesner and Alfieri 1953, p. 64. 
Type: Unknown. 
Distribution: Algeria; Egypt. 
Host Plant: Unrecorded. 
*Tingis (Lasiotropis) hellenica (Puton) 

Monanthia hellenica Puton 1877, p. lxviii [Greece]. — Horvath 1889, 

p. 329 [Syria]. 
Phyllontochila hellencia: Reuter 1891a, p. 26. 

Monanthia (Lasiotropis) corniculata J akovlev 1903a, p. 3 [Crimea]. 

Tingis (Lasiotropis) hellenica: Horvath 1906a, p. 69 [Israel; Turkey; 

Crete; Phlomis fruticosa]. — Kiritshenko 1951, p. 249. — Seidenstiicker 

1954, p. 235.— Hoberlandt 1955, p. 89.— Stichel 1960a, p. 308. 

Tingis hellenica: Lindberg 1948, p. 58 [Cyprus; Pistacia; Ceratonia]. 

Type: Sex unknown; Corfu, Greece; Paris Mus. 

Distribution: Greece; Crete; Cyprus; Turkey; Syria; Israel; U.S.S.R. 
(Crimea). 

Host Plants: Ceratonia sp.; Phlomis fruticosa; Pistacia sp. 
*Tingis (Lasiotropis) paranana Drake 

Tingis par anana Drake 1954d, p. 75 [Argentina], 
Type: Holotype & ; Parana, Entre Rois, Argentina; Hungarian Mus. 
Distribution: Argentina. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 407 

*Tingis (Lasiotropis) ragusana (Fieber) 

Monanthia ragusana Fieber 1861, p. 121 [Yugoslavia]. — Garbiglietti 
1896, p. 272.— Walker 1873a, p. 184.— Puton 1874a, p. 226.— Ferrari 
1874, p. 170 [Italy]; 1878, p. 66.— Horvath 1881b, p. xxxiv [Hun- 
gary]; 1889, p. 329.— Rey 1893, p. 97.— Hiieber 1893, p. 336 [Austria]. 
Lasiotropis ragusana: Stal 1874, p. 55. 
Monanthia ovatula Jakovlev 1877, p. 92 [Iran]. 

Tingis (Lasiotropis) ragusana: Horvath 1906a, p. 69, pi. 1, fig. 9 [Cau- 
casus; Rumania; Greece; Germany; France; Stachys italica]. — Stichel 
1926, p. Ill [Stachys]; 1960a, p. 308 [Poland; Malta; Verbascum phlo- 
moides]. — Gonzalez 1948, p. 50 [Spain]. — Kiritshenko 1951, p. 249. — 
Hoberlandt 1952, p. 112 [Cyprus]; 1955, p. 88 [Transcaucasus].— 
Seidenstiicker 1954, p. 235 [Turkey]. 
Tingis ragusana: Novak and Wagner 1951, p. 70 [Stachys salviaefolia]. 
Type: Sex unknown; Dalmatia; Vienna Mus. 

Distribution: Spain; France; Germany; Austria; Italy; Malta; Yugo- 
slavia; Rumania; Greece; Cyprus; Hungary; Poland; Turkey; Iran; 
U.S.S.R. (Caucasus; Transcaucasus). 

Host Plants: Stachys italica; Stachys salviaefolia; Stachys sp.; Verbascum 
phlomoides. 

*Tingis (Lasiotropis) reticulata Herrich-Schaeffer 

Tingis reticulata Herrich-Schaeffer 1835, p. 58. — Horvath 1916, p. 9 
[Albania]; 1918, p. 334.— Scholte 1935, p. 61, fig. 14 [Netherlands].— 
Stichel 1938a, p. 406 [Poland].— Poisson 1938, p. 589 [Psamma].— 
Mancini 1953b, p. 186.— Bator 1953, p. 325, pi. 2, fig. 7— Gomez- 
Menor 1955b, p. 248— Stusak 1957a, p. 24, figs. 9-13; 1961a, p. 85, 
figs. 1, 5b. — Cobben 1958b, p. 16. — Southwood and Leston 1959, 
p. 148, fig. 47. 

Monanthia reticulata: Herrich-Schaeffer 1836, p. 72, pi. 72, fig. 288 
[Germany].— Douglas and Scott 1865, p. 250.— Horvath 1874b, 
p. 432.— Ferrari 1874, p. 170; 1878, p. 66.— Saunders 1875, p. 247. 

Derephysia reticulata: Spinola 1837, p. 166 [Italy; France]. 

Tingis costata (not Fabricius): Schilling 1838, p. 105. 

Monanthia (Phyllontocheila) ciliata Fieber 1844, p. 67, pi. 5, figs. 31-33 
[Czechoslovakia; Austria; Senecio jacobaea; Verbascum thapsus]. 

Monanthia ciliata: Scholz 1847, p. 119. — Mayr 1858, p. 570 [Hungary; 
Ukraine; Greece].— Fieber 1861, p. 122.— Frey-Gessner 1865, p. 231 
[Switzerland].— Garbiglietti 1869, p. 273.— Gredler 1870, p. 75.— 
Walker 1873a, p. 184.— Reiber and Puton 1876, p. 69.— Vollenhoven 
1879, p. 230, pi. 12, fig. e— Chicote 1880, p. 189 [Spain] .—Jakovlev 
1880b, p. 104.— Dubois 1888, p. 121.— d'Antessanty 1890, p. 32.— 
Hiieber 1893, p. 334.— Jensen-Haarup 1912, p. 155, fig. 98a [Den- 
mark].— Butler 1923, p. 212 [Crimea; Ajuga]. 



408 U.S. NATIONAL MUSEUM BULLETIN 243 

Monanthia (Phyllontocheila) reticulata: Herrich-Schaeffer 1850, p. 154. 
Campylosteira (Derephysia) reticulata: Kolenati 1856, p. 431 [Caucasus; 

Transcaucasus; Cirsium; Verbascum]. 
Lasiotropis ciliata: Stal 1874, p. 55. 
Monanthia (Platychila) ciliata: Puton 1879c, p. 113 [Ajuga reptans]. — 

Saunders 1892, p. 133, pi. 12, fig. 7 [England]. 
Platychila ciliata: Reuter 1885, p. 44. 

Tingis (Lasiotropis) reticulata: Horvath 1906a, p. 70 [Ajuga genevensis] . — 
Oshanin 1908, p. 430 [Belgium; Yugoslavia; Rumania]. — Stichel 
1926, p. Ill, fig. 300 [Erica vulgaris]; 1960a, p. 309, fig. 113 [Sweden; 
Bulgaria; Calluna vulgaris; Salvia verticillata]. — Guide 1938, p. 291, 
fig— Hoberlandt 1942, p. 125; 1943a, p. 118; 1943b, p. 123.— 
Kiritshenko 1951, p. 249.— Stehlik 1952, p. 205. 
Phyllonthocheila [sic] ciliata: Reichensperger 1920, p. 62. 
Lasiotropis reticulata: Blote 1945, p. 87 [Tunisia]. 
Type: Unknown. 

Distribution: England; France; Netherlands; Belgium; Spain; Italy; 
Switzerland; Czechoslovakia; Germany; Denmark; Sweden; Poland; 
Austria; Hungary; Albania; Bulgaria; Yugoslavia; Rumania; Greece; 
U.S.S.R. (Crimea; Ukraine; Caucasus; Transcaucasus); Tunisia. 

Host Plants: Ajuga genevensis; Ajuga reptans; Ajuga sp.; Calluna vulgaris; 
Cirsium sp.; Erica vulgaris; Erica sp.; Psamma sp.; Salvia verticillata; Senecio 
jacobaea; Verbascum thapsus; Verbascum sp. 

Note: Description of species appears in Herrich-Schaeffer 1836. Eggs 
[Stusak]. 
*Tingis (Lasiotropis) rotundipennis Horvath 

Tingis rotundipennis Horvath 1911c, p. 108 [Egypt]. 

Tingis (Lasiotropis) rotundipennis: Priesner and Alfieri 1953, p. 64 
[Echinops spinosa]. 
Type: Lectotype 9, macropterous; Materie, Egypt; USNM. 
Distribution: Egypt. 
Host Plant: Echinops spinosa. 
Note: Lectotype here designated. 

*Tingis (Lasiotropis) shaowuana Drake and Maa 

Tingis (Lasiotropis) shaowuana Drake and Maa 1953, p. 91 [China]. 

Type: Holotype 9, macropterous; Shaowu, Fukien, China; Drake Coll. 
(USNM). 

Distribution: China. 

Host Plant: Unrecorded. 

*Tingis (Lasiotropis) trichonota (Puton) 

Monanthia clavicornis (not Linnaeus) : Burmeister 1835, p. 260 [Portugal]. 
Monanthia (Platychila) trichonota Puton 1874a, p. 216 [France]. 
Lasiotropis trichonota: Stal 1874, p. 55. 



LACE BUGS OF THE WORLD 409 

Monanthia (Platychila) trichonota: Puton 1879c, p. 112 [Phlomis lychnitis], 
Tingis (Lasiotropis) trichonota: Horvath 1906a, p. 69. — Oshanin 1908, 
p. 429 [Greece].— de Seabra 1931, pp. 425, 444, figs. 495(2), 496 — 
Gonzalez 1948, p. 50 [Spain].— Seidenstucker 1954, p. 235 [Tur- 
key].— Stichel 1960a, p. 208. 
Tingis trichonota: Stichel 1938a, p. 406 [Switzerland]. 
Type: Sex unknown; Avignon, France; Paris Mus. 
Distribution: Portugal; Spain; France; Switzerland; Greece; Turkey. 
Host Plant: Phlomis lychnitis. 

Subgenus TINGIS (TROPIDOCHEILA) Fieber 

Monanthia (Tropidocheila) Fieber 1844, p. 72. 

Monanthia (Tropidochila): Puton 1879c, p. 114. 

Tropidochila: Lethierry and Severin 1896, p. 19. — Blote 1945, p. 87. 

Tingis {Tropidocheila): Horvath 1906a, p. 79— Muzik 1907, p. 59 — 

Stichel 1926, pp. Ill, 113; 1960a, p. 316.— de Seabra 1931, p. 427 — 

Drake and Ruhoff 1960a, p. 86. 
Tingis (Tropidochila): Oshanin 1908, p. 437; 1912, p. 44— Borner 

1935, p. 77.— Stichel 1935, p. 349.— Guide 1938, pp. 288, 296 (key).— 

Priesner and Alfieri 1953, p. 64. 
Type Species: Monanthia (Tropidocheila) stachydis Fieber. 

::: Tingis (Tropidocheila) abundans Drake and Hambleton 
Tingis abundans Drake and Hambleton 1945, p. 357 [Peru]. 

Type: Holotype o", macropterous; Pucallpa, Peru; Drake Coll. 
(USNM). 

Distribution: Peru. 

Host Plant: Unrecorded. 

*Tingis (Tropidocheila) acris Drake 

Tingis (Tropidocheila) acris Drake 1947c, p. 117 [Queensland]. 

Type: Holotype cf, macropterous; Benakin, Queensland, Australia; 
Drake Coll. (USNM). 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

*Tingis (Tropidocheila) aegyptiaca Priesner 

Tingis (Tropidochila) aegyptiaca Priesner 1951, p. 138, fig. [Egypt]. — 
Priesner and Alfieri 1953, p. 65. 
Type: Holotype 9; Arabian Desert, Wadi Nouega, Egypt; Alfieri Coll. 
(USNM). 

Distribution: Egypt. 
Host Plant: Unrecorded. 



6S1-552— G4 27 



41 (J U.S. NATIONAL MUSEUM BULLETIN 24 3 

*Tingis (Tropidocheila) bodenheimeri Lindberg 

Tingis bodenheimeri Lindberg 1930, p. 69, fig. 1 [Israel]. 
Tingis (Lasiotropis) bodenheimeri: Seidenstiicker 1954, p. 235 [Turkey]. 
Tingis (Tropidocheila) bodenheimeri: Stichel 1960a, p. 319 [Syria]. 
Type: Sex unknown; Jerusalem, Palestine; Helsin. Mus. 
Distribution: Turkey; Israel; Syria. 
Host Plant: Unrecorded. 
Tingis (Tropidocheila) cappadocica Horvath 

Tingis (Tropidocheila) cappadocica Horvath 1906a, p. 82 [Turkey]. — 
Hoberlandt 1955, p. 91.— Stichel 1960a, p. 319. 
Type: Holotype 9; Kaisarie, Asia Minor; Vienna Mus. 
Distribution: Turkey. 
Host Plant: Unrecorded. 
* Tingis (Tropidocheila) caucasica (Jakovlev) 

Monanthia (Tropidochila) caucasica Jakovlev 1880a, pp. 136, 137 [Dages- 
tan]: 1880b, p. 107. 
Tingis (Tropidocheila) caucasica: Horvath 1906a, p. 83 [Hungary; 

Yugoslavia; Crimea]. — Stichel 1960a, p. 320. 
Tingis (Tropidochila) caucasica: Kiritshenko 1951, p. 250. — Seiden- 
stiicker 1954, p. 235 [Turkey]. 
Tingis caucasica: Stichel 1938a, p. 406 [France; Albania]. — Putshkov 

1960, p. 304 [Ajuga]. 
Tingis (Tropidocheila) caucasia [sic]: Hoberlandt 1955, p. 91. 
Type: Holotype 9, macropterous; Derbent, Dagestan, Russia; Leningrad 
Inst. 

Distribution: France; Hungary: Yugoslavia; Albania; Turkey; U.S.S.R. 
(Dagestan; Crimea). 

Host Plant: Ajuga sp. 
*Tingis (Tropidocheila) gamboana Drake and Hambleton 

Tingis gamboana Drake and Hambleton 1945, p. 357 [Canal Zone; elm]. 
Tingis (Tropidocheila) gamboana: Hurd 1946, p. 465. 
Type: Holotype 9, macropterous; Gamboa, Panama Canal Zone; Drake 
Coll. (USNM).' 
Distribution: Panama (Canal Zone). 
Host Plant: Elm. 
*Tingis (Tropidocheila) geniculata (Fieber) 

Monanthia (Tropidocheila) geniculata Fieber 1844, p. 75, pi. 6, figs. 19-21 

[Hungary; Austria]. — Herrich-Schaeffer 1850, p. 152. 
Monanthia geniculata: Mayr 1858, p. 570. — Fieber 1861, p. 124 
[Ukraine].— Garbiglietti 1869, p. 273 [Italy].— Walker 1873a, p. 
185.— Horvath 1874b, p. 432.— Ferrari 1874, p. 170; 1878, p. 66 — 
Bolivar and Chicote 1879, p. 166 [Spain].— Jakovlev 1880b, p. 106 
[Caucasus]. — Puton 1886a, p. 5 [Tunisia]. — d'Antessanty 1890, p. 
32.— Hueber 1893, p. 339. 



LACEBUGS OF THE WORLD 411 

Tingis geniculata: Stal 1874, p. 54. — Reuter 1908, p. 88 [Abies alba; 
Juniperus communis; Populus]. — Novak and Wagner 1951, p. 71. — 
Mancini 1953d, p. 16 [Sicily]. 
Monanthia (Tropidochila) geniculata: Puton 1879c, p. 115. 
Tropidochila geniculata: Reuter 1891a, p. 26 [Greece]. — Lethierry and 

Severin 1896, p. 19 [Algeria].— Vidal 1937, p. 197 [Morocco]. 
Tingis (Tropidocheila) geniculata: Horvath 1906a, pp. 68, 83, pi. 1, 
fig. 7 [France; Germany; Yugoslavia; Rumania; Bulgaria; Turkey]. — 
Muzik 1907, p. 60 [Czechoslovakia].— Stichel 1926, p. 113; 1960a, 
p. 321, fig. 121 [Sardinia; Poland].— Hoberlandt 1943b, p. 120; 
1955, p. 91 [Transcaucasus]. 
Tingis (Tropidochila) geniculata: Borner 1935, p. 77, fig. 120b. — 
Kiritshenko 1951, p. 250.— Gomez-Menor 1955b, p. 248. 
Type: Unknown. 

Distribution: Spain; France; Germany; Czechoslovakia; Austria; 
Hungary; Italy; Sicily; Sardinia; Yugoslavia; Rumania; Poland; Bulgaria; 
Greece; Turkey; U.S.S.R. (Ukraine; Caucasus; Transcaucasus); Tunisia; 
Algeria; Morocco. 

Host Plants: Abies alba; Juniperus communis; Populus sp. 

*Tingis (Tropidocheila) granadensis Horvath 

Tingis (Tropidocheila) granadensis Horvath 1960a, p. 80 [Spain]. — 

Stichel 1960a, p. 317. 
Tingis granadensis: Lindberg 1932a, p. 42. — Wagner 1958, p. 240 
[France; Phlomis herba-venti]. 
Type: Holotype 9; Granada, Spain; Helsin. Mus. 
Distribution: Spain; France. 
Host Plant: Phlomis herba-venti. 

*Tingis (Tropidocheila) griseola (Puton) 

Monanthia (Tropidochila) geniculata var. griseola Puton 1879c, p. 116 

[France; Corsica; Sardinia; Algeria]. — Horvath 1892b, p. 131. 
Tropidochila geniculata var. griseola: Reuter 1891a, p. 26 [Greece]. 
Tingis (Tropidocheila) griseola: Horvath 1906a, p. 84 [Yugoslavia; 
Italy; Majorca].— Moroder Sala 1920, p. 12 [Spain].— Stichel 1926, 
p. 113; 1960a, p. 321. 
Tingis griseola: Novak and Wagner 1951, p. 71 [Juniperus phoenicea}. — 
Mancini 1952, p. 13 [Sicily]. 
Type: Sex unknown; France; Paris Mus. 

Distribution: Spain; France; Corsica; Balearic Islands (Majorca): 
Sicily; Italy; Sardinia; Greece; Yugoslavia; Algeria. 
Host Plant: Juniperus phoenicea. 



681-552—64 28 



412 U.S. NATIONAL MUSEUM BULLETIN 2 43 

Tingis (Tropidocheila) griseola var. miscella (Horvath) 

Phyllontocheila miscella Horvath 1902a, p. 598 [Sardinia; Sicily; Algeria]. 
Tingis {Tropidocheila) griseola var. miscella: Horvath 1906a, p. 84. — 
Stichel 1926, p. 113; 1960a, p. 321.— Hoberlandt 1955, p. 91 [Turkey]. 
Tingis griseola var. miscella: Novak and Wagner 1951, p. 71 [Yugosla- 
via].— Mancini 1953c, p. 22 [Italy]. 
Type: Unknown. 

Distribution: Italy; Sardinia; Sicily; Yugoslavia; Turkey; Algeria. 
Host Plant: Unrecorded. 

*Tingis (Tropidocheila) helvina (Jakovlev) 

Monanthia {Platychila) helvina Jakovlev 1876a, p. Ill [Siberia]. 

Phyllontochila helvina: Lethierry and Severin 1896, p. 17. 

Tingis {Tropidocheila) helvina: Horvath 1906a, p. 85. 
Type: Holotype 9, macropterous; Ussuri, Russia; Leningrad Inst. 
Distribution: U.S.S.R. (Siberia). 
Host Plant: Unrecorded. 

*Tingis (Tropidocheila) insularis (Horvath) 

Phyllontocheila insularis Horvath 1902a, p. 599 [Madeira]. 

Tingis {Tropidocheila) insularis: Horvath 1906a, p. 84. — China 1938, 

p. 20. 
Tingis {Tropidochila) insularis: Gomez-Menor 1955b, p. 248 [Spain]. 
Type: Holotype o 71 ; Madeira; Vienna Mus. 
Distribution: Madeira; Spain. 
Host Plant: Unrecorded. 

Tingis (Tropidocheila) juvenca (Horvath) 

Phyllontocheila juvenca Horvath 1902a, p. 598 [Algeria]. 

Tingis {Tropidocheila) juvenca: Horvath 1906a, p. 83 [Spain]. — Stichel 

1960a, p. 320. 
Tingis {Tropidochila) juvenca: Gomez-Menor 1955b, p. 248. 
Type: Holotype 9; Algeria; Hungarian Mus. 
Distribution: Algeria; Spain. 
Host Plant: Unrecorded. 

Tingis (Tropidocheila) kirinana Drake 

Tingis {Tropidocheila) kirinana Drake 1948a, p. 1, fig. 1 [China]. 
Type: Holotype 9; Kao-lin-tze, Kirin Province, Manchuria; Heude Mus. 
Distribution: China. 
Host Plant: Unrecorded. 

*Tingis (Tropidocheila) liturata (Fieber) 

Monanthia {Tropidocheila) liturata Fieber 1844, p. 74, pi. 6, figs. 16-18 

[Spain].— Herrich-Schaeffer 1850, p. 152. 
Monanthia liturata: Fieber 1861, p. 123. — Walker 1873a, p. 185. — 
Bolivar and Chicote 1879, p. 166. 



LACEBUGS OF THE WORLD 413 

Tingis liturata: Stal 1874, p. 54.— Lindberg 1932a, p. 42. 
Monanthia (Tropidochila) liturata: Puton 1879c, p. 116 [France]. 
Tropidochila liturata: Lethierry and Severin 1896, p. 19 [Algeria]. — 

Vidal 1937, p. 197.— Blote 1945, p. 87 [Egypt]. 
Tingis (Tropidocheila) liturata: Horvath 1906a, p. 84 [Portugal; Sicily; 

Tunisia; Mauritania]. — Stichel 1960a, p. 322, fig. 124 [Libya]. 
Tingis (Tropidochila) liturata: Oshanin 1908, p. 441 [Morocco]. — 
Priesner and Alfieri 1953, p. 65. 
Type: Andalusia; sex and deposition of type unknown. 
Distribution: Portugal; Spain; France; Sicily; Italy; Algeria; Tunisia; 
Mauritania; Morocco; Libya; Egypt. 
Host Plant: Unrecorded. 

*Tingis (Tropidocheila) marrubii Vallot 

Tingis marrubii Vallot 1829, p. 98 [Alarrubium vulgare]. 
Monanthia kiesenwetteri Mulsant and Rey 1852a, p. 135 [France]; 1852b, 
p. 154.— Fieber 1861, p. 383.— Walker 1873a, p. 187.— Bolivar and 
Chicote 1879, p. 166.— Jakovlev 1893, p. 294 [Russia]. 
Monanthia villosa Costa 1855a, p. 11; 1855b, p. 233. — Puton 1874a, 

p. 277.— Stichel 1960d, p. 120. 
Lasiotropis kiesenwetteri: Stal 1874, p. 56. 

Lasiotropis kiesenvetteri [sic]: Sahlberg 1878, p. 21 [Italy; Spain; Siberia]. 
Monanthia (Tropidochila) kiesenwetteri: Puton 1879c, p. 115 [Carduus; 

Marrubium]. 
Tropidochila kiesenwetteri: Lethierry and Severin 1896, p. 19 [Cauca- 
sus].— Vidal 1937, p. 197.— Blote 1945, p. 87 [Greece]. 
Tingis (Tropidocheila) kiesenwetteri: Horvath 1906a, p. 81 [Hungary; 
Rumania; Turkey; Turkestan; Algeria]. — Moroder Sala 1920, 
p. 12.— Stichel 1926, p. 113; 1960a, p. 318 [Bulgaria].— Hoberlandt 
1955, p. 90. 
Tingis kiesenwetteri: Horvath 1906d, p. 2. — Lindberg 1932a, p. 42 
[Morocco] .—Novak and Wagner 1951, p. 71 [Yugoslavia]. — 
Mancini 1960b, p. 114. 
Tingis (Tropidochila) kiesenwetteri: Guide 1938, p. 299 [Germany]. — 

Gomez-Menor 1955b, p. 248. 
Tingis (Tropidocheila) marrubii: Drake and Runoff 1960c, p. 32. 
Type: Unknown. 

Distribution: Spain; France; Germany; Italy; Hungary; Rumania: 
Yugoslavia; Bulgaria; Greece; Turkey; U.S.S.R. (Caucasus; Turkestan; 
Siberia); Algeria; Morocco. 
Host Plants: Carduus sp.; Marrubium vulgare; Marrubium sp. 



414 U.S. NATIONAL MUSEUM BULLETIN 243 

*Tingis (Tropidocheila) muiri Drake 

Tingis {Tropidocheila) muiri Drake 1947c, p. 116 [Queensland]. 
Type: Holotype cf, macropterous; Coolangata, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Tingis (Tropidocheila) oliveirai Drake and Hambleton 
Tingis oliveirai Drake and Hambleton 1938a, p. 64 [Brazil]. 
Type: Holotype o 71 , macropterous; Curvello, Minas Gerais, Brazil; 
Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 
*Tingis (Tropidocheila) pauperata (Puton) 

Monanthia kiesenwetteri var. pauperata Puton 1879b, p. 297 [Caucasus]. 
Monanthia (Tropidochila) kiesenwetteri var. pauperata: Puton 1879c, p. 115. 
Monanthia (Tropidochila) angustipennis Jakovlev 1880b, p. 107. 
Tingis (Tropidocheila) pauperata: Horvath 1906a, p. 81 [Armenia; 
Spain; France; Albania; Rumania]. — Stichel 1960a, p. 318 [Italy; 
Yugoslavia; Turkey; Turkestan]. 
Tingis (Tropidochila) pauperata: Kiritshenko 1913b, p. 482; 1951, p. 

250.— Gomez-Menor 1955b, p. 248. 
Tingis pauperata: Horvath 1916, p. 9. — Mancini 1953b, p. 186. — 

Putshkov 1960, p. 312 [Phlomis pungens]. 
Tropidochila pauperata: Vidal 1937, p. 197 [Morocco]. 
Type: Sex unknown; Caucasus, Russia; Paris Mus. 
Distribution: Italy; France; Spain; Yugoslavia; Rumania; Albania; 
Turkey; U.S.S.R. (Turkestan; Caucasus; Armenia); Morocco. 

Host Plant: Phlomis pungens. 
*Tingis (Tropidocheila) pilosa Hummel 

Tingis pilosa Hummel 1825, p. 69 [Russia]. — Reuter 1885, p. 44. — 
Lindberg 1932a, p. 42 [Morocco]; 1934, p. 29 [China].— Scholte 
1935, p. 62, fig. 17 [Netherlands].— Stichel 1938a, p. 406 [Portugal; 
Yugoslavia; Bulgaria; Poland]. — Bator 1953, p. 326, pi. 4, fig. 4. 
Monanthia angusticollis Herrich-SchaefFer 1836, p. 72, fig. 289 [Ger- 
many].— d' An tessanty 1890, p. 32.— Hiieber 1893, p. 340. 
Tingis car dui (not Linnaeus): Schilling 1838, p. 105. 
Derephysia angusticollis: Herrich-Schaeffer 1838, p. 71. 
Monanthia (Tropidocheila) pilosa: Fieber 1844, p. 79, pi. 6, figs. 36-37 

[Czechoslovakia; Austria]. 
Monanthia (Tropidocheila) angusticollis: Herrich-Schaeffer 1850, p 152. 
Monanthia pilosa: Mayr 1858, p. 571 [Hungary]. — Fieber 1861, p. 122 
[France]. — Frey-Gessner 1865, p. 231 [Switzerland; Stachys sylvatica; 
Galeopsis tetrahit].— Garbiglietti 1869, p. 273 [Italy].— Gredler 1870, 
p. 75.— Horvath 1874a, p. 432.— Reiber and Puton 1876, p. 69. 



LACEBUGS OF THE WORLD 415 

Lasiotropis pilosa: Stal 1874, p. 55. 

Monanthia (Tropidochila) angusticollis: Puton 1879c, p. 114 [Leonurus 

cardiaca; Lappa minor; Ballota nigra]. 
Monanthia {Tropidochila) pilosa: Jakovlev 1880a, p. 136. 
Tropidochila angusticollis: Reuter 1885, p. 44; 1891b, p. 184. 
Tingis (Tropidocheila) pilosa: Horvath 1906a, pp. 66, 79 [Siberia; 
Turkestan; Caucasus; Turkey]. — Stichel 1926, p. 113 [Humulus; 
Crepis tectorum; Senecio]; 1960a, p. 316 [Greece]. — Hoberlandt 1943a, 
p. 119; 1943b, p. 124; 1955, p. 91 [Transcaucasus; Turkmen]. 
Tingis (Tropidochila) pilosa: Oshanin 1908, p. 437 [Belgium; Ruma- 
nia].— Wu 1935, p. 450.— Guide 1938, p. 297, fig.— Gonzalez 1948, 
p. 50 [Spain].— Kiritshenko 1951, p. 250. 
Phyllonthocheila [sic] angusticollis: Reichensperger 1920, p. 62. 
Type: Russia; sex and deposition of type unknown. 

Distribution: Portugal; Spain; France; Netherlands; Belgium; Ger- 
many; Poland; Czechoslovakia; Switzerland; Austria; Italy; Hungary; 
Rumania; Yugoslavia; Bulgaria; Greece; Turkey; U.S.S.R. (Caucasus; 
Transcaucasus; Turkestan; Turkmen; Siberia); Morocco; China. 

Host Plants: Ballota nigra; Crepis tectorum; Galeopsis tetrahit; Lappa minor; 
Leonurus cardiaca; Humulus sp; Senecio sp.; Stachys sylvatica. 
*Tingis (Tropidocheila) pilosa var. amplicosta (Montandon) 
Lasiotropis amplicosta Montandon 1897, p. 100 [Algeria]. 
Tingis (Tropidocheila) pilosa var. amplicosta: Horvath 1906a, p. 80. — de 
Seabra 1931, p. 427, figs. 499, 495 (4-6) [Portugal; Ballota nigra; 
Galeopsis tetrahit; Lappa minor; Leonurus cardiaca; Stachys sylvatica]. — 
Stichel 1960a, p. 317. 
Type: Sex unknown; Tenient-el-Haad, Algeria; Bucharest Mus. 
Distribution: Algeria; Portugal. 

Host Plants: Ballota nigra; Galeopsis tetrahit; Lappa minor; Leonurus 
cardiaca; Stachys sylvatica. 
*Tingis (Tropidocheila) pilosa var. antennalis (Puton) 

Monanthia kiesenwetteri var. antennalis Puton 1879b, p. 297 [Russia]. 
Monanthia (Tropidochila) kiesenwetteri vox. antennalis: Puton 1879c, p. 115. 
Monanthia (Tropidochila) tenuicornis Jakovlev 1880a, p. 136. 
Phyllontocheila antennalis: Horvath 1898b, p. 279. 
Tingis (Tropidocheila) pilosa var. antennalis: Horvath 1906a, p. 80 

[Turkestan; Siberia; Mongolia]. — Stichel 1960a, p. 317. 
Tingis (Tropidochila) pilosa var. antennalis: Wu 1935, p. 450. 
Type: Sex unknown; Sarepta, Russia; Paris Mus. 
Distribution: Mongolia Republic; U.S.S.R. (Turkestan; Siberia). 
Host Plant: Unrecorded. 



416 U.S. NATIONAL MUSEUM BULLETIN 2 43 

*Tingis (Tropidocheila) pusilla (Jakovlev) 

Monanthia pusilla Jakovlev 1874a, pp. 10, 33 [Russia]. 

Tingis {Tropidocheila) pusilla: Horvath 1906a, p. 83. — Seidenstucker 

1954, p. 235 [Turkey].— Stichel 1960a, p. 319. 
Tingis (Tropidochila) pusilla: Kiritshenko 1951, p. 250. 
Type: Holotype 9; Russia; Leningrad Inst. 
Distribution: U.S.S.R.; Turkey. 
Host Plant: Unrecorded. 

Tingis (Tropidocheila) reuteri Horvath 

Tingis (Tropidocheila) reuteri Horvath 1906a, p. 81 [Turkmen]. — 
Hoberlandt 1955, p. 91 [Turkey] .—Stichel 1960a, p. 318. 
Type: Holotype o"; Ashkhabad, Turkmen, Russia; Helsin. Mus. 
Distribution: U.S.S.R. (Turkmen); Turkey. 
Host Plant: Unrecorded. 

*Tingis (Tropidocheila) saueri Drake and Hambleton 
Tingis saueri Drake and Hambleton 1939, p. 156 [Brazil]. 

Type: Holotype o 71 , macropterous; Recife, Pernambuco, Brazil; Drake 
Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

Tingis (Tropidocheila) stachydis (Fieber) 

Monanthia maculata Herrich-Schaeffer 1838, pp. 51, 56, pi. 123, figs. 

389a-c [Germany].— Ferrari 1884, p. 475 [Tunisia].— Puton 1886a, 

p. 12.— d'Antessanty 1890, p. 32.— Rey 1893, p. 97.— Hiieber 1893, 

p. 339. 
Monanthia (Tropidocheila) stachydis Fieber 1844, p. 73, pi. 6, figs. 13-15 

[Austria; Czechoslovakia; Stachys recta]. 
Monanthia (Tropidocheila) maculata: Herrich-Schaeffer 1850, p. 152. 
Monanthia stachydis: Fieber 1861, p. 123. — Frey-Gessner 1865, p. 232 

[Switzerland].— Garbiglietti 1869, p. 273 [Italy].— Walker 1873a, 

p. 185.— Horvath 1874b, p. 432.— Jakovlev 1876b, p. 67 [Russia].— 

Reiber and Puton 1876, p. 69. — Bolivar and Chicote 1879, p. 166 

[Spain]. 
Tingis maculata: Walker 1873a, p. 179.— Horvath 1874b, p. 432 

[Hungary]. — Putshkov 1960, p. 304 [Sideritis]. 
Tingis stachydis: Stal 1874, p. 54. — Drake and Ruhoff 1962c, p. 135. 
Monanthia stahidis [sic]: Ferrari 1878, p. 66. 

Monanthia (Tropidochila) maculata: Puton 1879c, p. 116 [France]. 
Tingis (Tropidocheila) maculata: Horvath 1906a, pp. 69, 84. — Stichel 

1926, p. 113 [Teucrium chamaedrys]; 1960a, p. 322 [Sicily; Poland; 

Stachys annua]. 
Phyllonthocheila [sic] maculata: Reichensperger 1920, p. 62. 



LACEBUGS OF THE WORLD 417 

Tingis {Tropidochila) maculata: Bonier 1935, p. 77, fig. 120a. — Kirit- 
shenko 1951, p. 251. 
Type: Sex unknown; Regensburg, Germany; Munich Mus. 
Distribution: Spain; France; Sicily; Italy; Switzerland; Germany; 
Czechoslovakia; Poland; Austria; Hungary; U.S.S.R.; Tunisia (?). 

Host Plants: Sideritis sp.; Stachys annua; Stachys recta; Stachys sp.; Tsucrium 
chamaedrys. 

Note: See Horvath 1906a re Tunisia distribution. 
*Tingis (Tropidocheila) torpida (Horvath) 

Tropidochila geniculata var. Reuter 1900, p. 187. 

Phyllontocheila geniculata var. torpida Horvath 1902a, p. 597 [Algeria; 

Tunisia; Morocco]. 
Tingis (Tropidocheila) torpida: Horvath 1906a, p. 83 [Spain; Mauri- 
tania].— Moroder Sala 1920, p. 12.— Stichel 1960a, p. 320 [France]. 
Tropidochila torpida: Blote 1945, p. 88 [Sardinia]. 
Type: Unknown. 

Distribution: Spain; Sardinia; Morocco; Tunisia; Algeria; Mauritania. 
Host Plant: Unrecorded. 
Tingis (Tropidocheila) valida (Puton) 

Monanthia (Lasiotropis) valida Puton 1878, p. Ixviii [Syria]. 

Tingis (Tropidocheila) valida: Horvath 1906a, p. 79 [Turkey; Lebanon; 

Iran]. — Stichel 1960a, p. 316 [Turkestan]. 
Tingis (Tingis) valida: Hoberlandt 1955, p. 90. 
Type: Sex unknown; Syria; Vienna Mus. 

Distribution: Syria; Lebanon; Iran; Turkey; U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 
Tingis (Tropidocheila) valida var. biseriata (Horvath) 

Phyllontocheila valida var. biseriata Horvath 1902a, p. 600 [Turkestan]. 
Tingis (Tropidocheila) valida var. biseriata: Horvath 1906a, p. 79. — 
Stichel 1960a, p. 316. 
Type: Holotype 9; Varzaminor, Turkestan, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 
Tingis (Tropidocheila) valida var. discessa (Horvath) 

Phyllontocheila valida var. discessa Horvath 1902a, p. 600 [Turkestan]. 
Tingis (Tropidocheila) valida var. discessa: Horvath 1906a, p. 79. — 
Stichel 1960a, p. 316. 
Type: Holotype 9: Varzaminor, Turkestan, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Turkestan). 
Host Plant: Unrecorded. 

Genus TRACHYPEPLUS Horvath 

Trachypeplus Horvath 1926, p. 329.— Drake and Ruhoff 1960a, p. 86. 
Type Species: Trachypeplus Jacob soni Horvath. 



418 U.S. NATIONAL MUSEUM BULLETIN 243 

*Trachypeplus bakeri Drake 

Trachypeplus bakeri Drake 1927d, p. 308 [Mindanao]. — Drake and 
Poor 1937a, p. 12. 
Type: Holotype cf , macropterous; Iligan, Mindanao, Philippine Islands; 
Drake Coll. (USNM). 
Distribution: Philippine Islands (Mindanao). 
Host Plant: Unrecorded. 

^Trachypeplus chinensis Drake and Poor 

Trachypeplus chinensis Drake and Poor 1936a, p. 440 [Hainan; Hong 
Kong].— Drake 1937b, p. 593; 1938b, p. 195; 1947d, p. 229, fig. 
[Viet-Nam]; 1947g, no. 69, fig. (color). 
Type: Holotype cf , macropterous; Liamui, Hainan Island; Drake Coll. 
(USNM). 
Distribution: China (Hainan Island); Viet-Nam; Hong Kong. 
Host Plant: Unrecorded. 

"'Trachypeplus guinaicus Drake 

Trachypeplus guinaicus Drake 1960, p. 350 [New Guinea]. 
Type: Holotype 9; Asama, Morobe, Northeast New Guinea; Drake Coll. 
(USNM). 
Distribution: New Guinea (Northeast). 
Host Plant: Unrecorded. 

*Trachypeplus idoneus Drake and Ruhoff 

Trachypeplus idoneus Drake and Ruhoff 1962d, p. 491 [Sumatra]. 

Type: Holotype 9, macropterous; Meclan, Sumatra; Drake Coll. 
(USNM). 

Distribution: Greater Sunda Islands (Sumatra). 

Host Plant: Unrecorded. 

*Trachypeplus jacobsoni Horvath 

Trachypeplus jacobsoni Horvath 1926, p. 330, fig. 3 [Sumatra; Mallotus 
philippinensis']. — Drake and Poor 1936b, p. 144 [India]. — Blote 1945, 
p. 92.— Singh 1953, p. 119. 
Type: Fort de Kock, Sumatra; sex and deposition of type unknown. 
Distribution: Greater Sunda Islands (Sumatra); India. 
Host Plant: Mallotus phi lippinensis. 

*Trachypeplus malloti Drake and Poor 

Trachypeplus malloti Drake and Poor 1936b, p. 144 [India; Mallotus 
philippinensis]. 
Type: Holotype cf, macropterous; Dehra Dun, India; Drake Coll. 
(USNM). 
Distribution: India. 
Host Plant: Mallotus philippinensis. 



LACEBUGS OF THE WORLD 419 

Genus UHLERITES Drake 

Uhlerites Drake 1927a, p. 56; 1948b, p. 56.— Drake and Ruhoff 1960a, 
p. 87. 
Type Species: Phyllontocheila debile Uhler. 

*Uhlerites debilis (Uhler) 

Phyllontochila debile Uhler 1896, p. 265 [Japan]. — Matsumura 1931, 

p. 1202, fig. 
Stephanitis (Norba) x-nigrum Lindberg 1927, p. 15, pi. 1, fig. 3 [Siberia]. 
Uhlerites debile: Drake 1927a, p. 56; 1948b, p. 56 [China].— Takeya 

1951a, p. 14 [Taiwan]; 1953d, p. 173. 
Uhlerites debilis: Takeya 1931, p. 78, pi. 7, fig. 5; pi. 9, fig. 7 [Quercus 
serrata; Quercus glandulifera; Quercus dentata]; 1932, p. 10 [Korea]. — 
Esaki 1932, p. 1636, fig.; 1954, p. 237, fig. 612.— Saito 1933, p. 7 
[Quercus acutissima]. — Kato 1933, pi. 31, fig. 1. — Fukui 1938, pi. 50, 
tab. 36. 
Type: Holotype 9, macropterous; Japan; USNM. 
Distribution: Japan; China; Taiwan; Korea; U.S.S.R. (Siberia). 
Host Plants: Quercus acutissima; Quercus dentata; Quercus glandulifera; 
Quercus serrata. 
*Uhlerites latius Takeya 

Uhlerites latiorus Takeya 1931, p. 80, pi. 9, figs. 8-10 [Japan]. 
Uhlerites latius (emended): Drake 1948b, p. 56 [China]. — Takeya 
1951a, p. 14. 
Type: Sex unknown; Honghu, Pro v. Harima, Japan; Kyushu Univ. 
Distribution: Japan; China. 
Host Plant: Unrecorded. 

Genus ULOCYSTA Drake and Hambleton 

Ulocysta Drake and Hambleton 1945, p. 364. — Drake and Ruhoff 
1960a, p. 87. 
Type Species: Ulocysta praestabilis Drake and Hambleton. 

"Ulocysta praestabilis Drake and Hambleton 

Ulocysta praestabilis Drake and Hambleton 1945, p. 365 [Colombia]. 

Type: Holotype 9, macropterous; San Jose, Colombia; Drake Coll. 
(USNM). 

Distribution: Colombia. 

Host Plant: Unrecorded. 

Genus ULONEMIA Drake and Poor 

Perissonemia (Ulonemia) Drake and Poor 1937a, p. 3. 
Ulonemia: Drake 1942b, p. 359.— Drake and Ruhoff 1960a, p. 87. 
Type Species: Perissonemia (Ulonemia) dignata Drake and Poor. 



420 U.S. NATIONAL MUSEUM BULLETIN 243 

Ulonemia aptata Drake 

Ulonemia aptata Drake 1960, p. 351, fig. 8 [New Guinea]. 
Type: Holotype 9; Nondugl, Northeast New Guinea; Bishop Mus. 
Distribution: New Guinea (Northeast). 
Host Plant: Unrecorded. 
*Ulonemia assamensis (Distant) 

Teleonemia assamensis Distant 1903a, p. 49 [India]; 1903b, p. 143, fig. 

106.— Drake 1937b, p. 592 [China]. 
Perissonemia assamensis: Drake and Poor 1936a, p. 439 [Hainan; 

Taiwan]. 
Perissonemia {Ulonemia) assamensis: Takeya 1951a, p. 18. 
Ulonemia assamensis: Drake and Ruhoff 1960c, p. 29. 
Type: Sex unknown; Margherita, Assam, India; British Mus. 
Distribution: India; Taiwan; China (Hainan Island); Greater Sunda 
Islands (Borneo). 

Host Plant: Unrecorded. 
*Ulonemia concava Drake 

Ulonemia concava Drake 1942b, p. 359 [Queensland]. 
Type: Holotype cf , macropterous; Cedar Creek, Queensland, Australia; 
Drake Coll. (USNM). 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 
*Ulonemia decoris Drake 

Ulonemia decoris Drake 1942b, p. 360 [Queensland; South Australia]. 
Type: Holotype d\ macropterous; Mount Glorious, Queensland, 
Australia; Drake Coll. (USNM). 
Distribution: Australia (Queensland; South Australia). 
Host Plant: Unrecorded. 
*Ulonemia dignata (Drake and Poor) 

Perissonemia {Ulonemia) dignata Drake and Poor 1937a, p. 3 [Luzon; 

Mindanao]. 
Ulonemia dignata: Drake and Ruhoff 1960c, p. 29. 
Type: Holotype cf, macropterous; Baguio, Benguet Subprovince, 
Luzon, Philippine Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon; Mindanao). 
Host Plant: Unrecorded. 
*Ulonemia electa (Drake and Poor) 

Perissonemia {Ulonemia) electa Drake and Poor 1937a, p. 5 [Luzon]. 
Ulonemia electa: Drake and Ruhoff 1960c, p. 29. 
Type: Holotype cf, macropterous; Baguio, Benguet Subprovince, Luzon, 
Philippine Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 



LACE BUGS OF THE WORLD 421 

*Ulonemia leai Drake 

Ulonemia leai Drake 1942b, p. 360 [Australia]. 
Type: Holotype 9, macropterous; Cairns District, Australia; Drake Coll. 
(USNM). 
Distribution : Australia. 
Host Plant: Unrecorded. 

*Ulonemia malaccae (Drake) 

Perissonemia malaccae Drake 1942a, p. 3 [Malaya]. 
Ulonemia malaccae: Drake and Runoff 1960c, p. 29. 
Type: Holotype 9, macropterous; Malacca, Perak, Federated Maiay 
States; Drake Coll. (USNM). 
Distribution: Federation of Malaya. 
Host Plant: Unrecorded. 

^ Ulonemia mjobergi (Horvath) 

Tingis (Tingis) mjobergi Horvath 1925a, p. 5 [Australia]. 
Tingis myobergi [sic]: Drake 1947c, p. 114, fig. 24b. 
Ulonemia mjobergi: Drake and Ruhoff 1960c, p. 29; 1961b, p. 139, fig. 
12.— Drake 1961b, p. 112 [New South Wales]. 
Type: Sex unknown; Australia occidentalis; Stockholm Mus. 
Distribution: Australia (New South Wales; Western Australia). 
Host Plant: Unrecorded. 

*Ulonemia plesia Drake and Ruhoff 

Ulonemia plesia Drake and Ruhoff 1961b, p. 138 [Western Australia]. 

Type: Holotype 9, macropterous; Swan River, Western Australia, 
Australia; Drake Coll. (USNM). 

Distribution: Australia (Western Australia). 

Host Plant: Unrecorded. 

Genus ULOTINGIS Drake and Hambleton 

Ulotingis Drake and Hambleton 1935, p. 144. — Monte 1939b, p. 82; 
1941e, p. 148.— Drake and Ruhoff 1960a, p. 87. 
Type Species: Acysta brasiliensis Drake. 

*Ulotingis brasiliensis (Drake) 

Acysta brasiliensis Drake 1922b, p. 42 [Brazil]. 

Ulotingis brasiliensis: Drake and Hambleton 1935, p. 145; 1938a, 
p. 64 [Psidium guajava]. — Silva 1956, p. 67. 
Type: Holotype cf, macropterous; Para, Brazil; USNM. 
Distribution: Brazil. 
Host Plant: Psidium guajava. 



422 U.S. NATIONAL MUSEUM BULLETIN 243 

*Ulotingis decor Drake and Hambleton 

Ulotingis decor Drake and Hambleton 1935, p. 145 [Brazil; Myrtaceae]. 

Type: HoJotype cf, macropterous; Santo Amaro, Sao Paulo, Brazil; 
Drake Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Myrtaceae. 

*Ulotingis nitor Drake and Hambleton (emendation) 

Ulotingis nitoris Drake and Hambleton 1935, p. 145, fig. 2 [Brazil; 
Myrtaceae] . 
Type: Holotype d\ macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Myrtaceae. 

*Ulotingis uniseriata (Drake) 

Acysta brasiliensis var. uniseriata Drake 1922c, p. 368, pi. 39, fig. 10 

[Brazil]. 
Acysta uniseriata: Drake and Bondar 1932, p. 91 [Byrsonima sericea]. — 
Costa Lima 1936, p. 125. — Monte 1937a, p. 35, fig. 13 [Byrsonima 
verbascifolia] . 
Ulotingis uniseriata: Drake and Hambleton 1935, p. 145; 1938a, p. 65. — 
Silva 1956, p. 68, fig. 29. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plants: Byrsonima sericea; Byrsonima verbascifolia. 

Genus URENTIUS Distant 

Urentius Distant 1903b, p. 134.— Bergroth 1911, p. 186; 1914b, p. 183 — 
Priesner and Alfieri 1953, p. 63 (key) .— Stichel 1960a, p. 298; 1960b. 
p. 394; 1960c, p. 133.— Drake and Ruhoff 1960a, p. 88. 
Ayrerus Distant 1903b, p. 140— Horvath 1909b, p. 632.— Drake and 

Ruhoff 1960a, p. 38. 
Prionostirina Schumacher 1913, p. 457. — Drake and Ruhoff 1960a, p. 79. 
Type Species: Urentius echinus Distant = Tingis hystricellus Richter. 

*Urentius chobauti Horvath 

Urentius chobauti Horvath 1907, p. 304 [France; Spain; Cistus albidus]. — 
Oshanin 1908, p. 457; 1912, p. 43.— Stichel 1935, p. 349; 1938a, 
p. 406; 1960a, p. 298. 
Type: Unknown. 
Distribution: France; Spain. 
Host Plant: Cistus albidus. 



LACEBUGS OF THE WORLD 423 

*Urentius euonymus Distant 

Urentius euonymus Distant 1909a, p. 116 [Ceylon]; 1910a, p. 114, fig. 

57.— Drake and Ruhoff 1962a, p. 156; 1962c, p. 133. 
Prionostirina nana Schumacher 1913, p. 458 [Israel]. 
Urentius nanus: Bergroth 1914b, p. 183. — Hoberlandt 1955, p. 88 

[Turkey; Syria].— Stichel 1960a, p. 299. 
Urentius hoggari Bergevin 1928, p. 336, fig. 1 [Algeria; Helianthemum 

lippii}.— Stichel 1960a, p. 299. 
Urentius maculatus Drake 1933, p. 1015 [India]. 

Urentius abutilinus Priesner and Alfieri 1953, p. 63 [Egypt; Abutilon 
muticum]. — Stichel 1960a, p. 299 [Abutilon pannosum]. 
Type: Sex unknown; Batticaloa, Ceylon; British Mus. 
Distribution: Ceylon; India; Israel; Syria; Turkey; Egypt; Sudan; 
Algeria. 

Host Plants: Abutilon muticum; Abutilon pannosum; Abutilon sp.; Cajanus 
indicus; Helianthemum lippii; Sida cordifolia. 
Urentius euphorbiae Menon and Hakk (nom. nud.) 

Urentius euphorbiae Menon and Hakk 1959b, p. 393. 
* Urentius hystricellus (Richter) 

Tingis hystricellus Richter 1869, p. 84, fig. 55 [Ceylon].— Green 1882, p. 

161. 
Ayrerus hystricellus: Distant 1903b, p. 141, fig. 104 [Solanum melongena]. 
Urentius echinus Distant 1903b, p. 134, fig. 97. — Maxwell-Lefroy 1909, 
p. 693 [India].— Fletcher 1920, p. 263.— Singh 1953, p. 120.— Drake 
1954a, p. 232 [Niger].— Patel and Kulkarny 1955, pp. 86-96, pis. 
1, 2. — Thontadarya and Channa Basavanna 1959, p. 289. — Drake 
and Davis 1960, fig. 32. 
Urentius olivaceus Distant 1909a, p. 115; 1910a, p. 112. 
Urentius sentis Distant 1909a, p. 115; 1910a, p. 113, fig. 56. 
Urentius aegyptiacus Bergevin 1930, p. 18, fig. [Egypt]. — Priesner and 

Alfieri 1953, p. 63.— Stichel 1960a, p. 299. 
Urentius hystricellus: Drake and Ruhoff 1960a, p. 89; 1962a, p. 156; 
1962c, p. 133. 
Type: Holotype 9; Ceylon; deposition unknown. 

Distribution: Ceylon; India; Niger; Egypt; Kenya; Sudan; Senegal; 
Uganda; Southern Rhodesia. 
Host Plant: Solanum melongena. 

Note: (1) Taken at port-of-entry, Washington, D.C. on leaves of egg- 
plant; (2) notes on egg laying [Thontadarya and Channa Basavanna]; 
(3) life history study [Patel and Kulkarny]. 
Urentius indicus Menon and Hakk (nom. nud.) 

Urentius indicus Menon and Hakk 1959b, p. 393. 
Urentius pusaensis Menon and Hakk (nom. nud.) 

Urentius pusaensis Menon and Hakk 1959b, p. 393. 



424 U.S. NATIONAL MUSEUM BULLETIN 243 

Urentius sarinae Hacker 

Urentius sarinae Hacker 1929, p. 325, pi. 32, fig. 3 [Queensland]. 
Type: Sex unknown; Sarina, Queensland, Australia; Queensland Mus. 
Distribution: Australia (Queensland). 
Host Plant: Unrecorded. 

Urentius sidae Menon and Hakk (nom. nud.) 

Urentius sidae Menon and Hakk 1959b, p. 393. 

*Urentius vepris Drake 

Urentius vepris Drake 1945, p. 100 [Transvaal; Hermannia micropetela]. 
Type: Holotype o 71 ; Commando Nek, Pretoria, Transvaal; British Mus. 
Distribution : South Africa (Transvaal) . 
Host Plant: Hermannia micropetela. 

Urentius zizyphifolius Menon and Hakk (nom. nud.) 

Urentius zizyphifolius Menon and Hakk 1959b, p. 393. 

Genus VATIGA Drake and Hambleton 

Vatiga Drake and Hambleton 1946a, p. 10. — Hurd 1946, p. 466. — 
Drake and Ruhoff 1960a, p. 88. 
Type Species: Vatiga vicosana Drake and Hambleton. 

* Vatiga cassiae (Drake and Hambleton) 

Tigava cassiae Drake and Hambleton 1934, p. 440 [Brazil; Cassia]. — ■ 

Costa Lima 1936, p. 130.— Monte 1941e, p. 144. 
Vatiga cassiae: Drake and Hambleton 1946a, p. 10. 
Type: Holotype d% macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Cassia sp. 

* Vatiga celeb rata (Drake) 

Leptopharsa celebratis Drake 1928a, p. 53 [Brazil]. — Monte 1941e, p. 109. 
Vatiga celebratis: Drake and Ruhoff 1960c, p. 29. 
Type: Holotype cf, macropterous; Rio Grande do Sul, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

* Vatiga illudens (Drake) Figure 3 

Atheas nigricornis (not Champion): Van Duzee 1907, p. 22. 

Leptopharsa illudens Drake 1922c, p. 370 [Puerto Rico; Jamaica; Domin- 
ican Republic]; 1930b, p. 25 [Brazil] .— Wolcott 1923, p. 246 [Mani- 
hot]. — Drake and Bruner 1924a, p. 148 [Haiti; Cuba; Manihot 
utilissima].— Barber 1939, p. 370.— Monte 1941e, p. 113; 1942d, p. 
108. — Bruner, Scaramuzza, and Otero 1945, p. 109 [Manihot 
esculenta]. — Gomez-Menor 1956a, p. 112. 



LACEBUGS OF THE WORLD 425 

Atheas pallidas Barber 1923, p. 6. 

Vatiga illudens: Drake and Hambleton 1946a, p. 10. — Silva 1956, p. 69 
[Manihot dulcis]. — Drake and Cobben 1960, p. 85, figs. 84a, b [St. 
Eustatius]. 
Type: Holotype 9, macropterous; Mandeville, Jamaica; Drake Coll. 
(USNM). 

Distribution: Cuba; Haiti; Dominican Republic; Jamaica; Puerto Rico; 
Leeward Islands (St. Eustatius); Brazil. 

Host Plants: Manihot dulcis; Manihot esculenta; Manihot utilissima; 
Manihot sp. 

* Vatiga illudens var. varianta (Drake) 

Leptopharsa illudens var. variantis Drake 1930b, p. 25 [Brazil]. — Monte 

1940b, p. 378; 1941e, p. 113 [Manihot dulcis]. 
Vatiga illudens var. variantis: Drake and Hambleton 1946a, p. 10. — 
Silva 1956, p. 70. 
Type: Holotype cf , macropterous; Bahia, Brazil; Drake Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Manihot dulcis. 

* Vatiga lonchocarpa (Drake and Hambleton) 

Tigava lonchocarpa Drake and Hambleton 1944b, p. 125 [Brazil; Loncho- 

carpus] . 
Vatiga lonchocarpa: Drake and Hambleton 1946a, p. 10. 
Type: Holotype 9, macropterous; Vicpsa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil. 
Host Plant: Lonchocarpus sp. 

* Vatiga longula (Drake) 

Leptopharsa longula Drake 1922c, p. 371, pi. 39, fig. 4 [Brazil]. — Costa 
Lima 1936, p. 129.— Monte 1937a, p. 35; 1939b, p. 74 [Bauhinia; 
Manihot utilissima]; 1941e, p. 114. 
Vatiga longula: Drake and Hambleton 1946a, p. 10. 
Type: Holotype 9; Chapada, Brazil; Carnegie Mus. 
Distribution: Brazil. 
Host Plants: Bauhinia sp.; Manihot utilissima. 

* Vatiga manihotae (Drake) 

Leptopharsa manihotae Drake 1922c, p. 371 [Trinidad; Manihot]. — Drake 
and Bruner 1924a, p. 148.— Drake and Hambleton 1934, p. 445 
[Brazil; Manihot utilissima]. — Costa Lima 1936, p. 129. — Monte 
1937a, p. 35, fig. 4; 1939b, p. 74 [Cuba]; 1941e, p. 115. 
Vatiga manihotae: Drake and Hambleton 1946a, p. 10. 
Type: Holotype 9, macropterous; Trinidad, BWI; USNM. 
Distribution: Trinidad; Cuba; Brazil; Peru; Paraguay; Argentina. 
Host Plants: Manihot utilissima; Manihot sp. 



426 U.S. NATIONAL MUSEUM BULLETIN 243 

* Vatiga pauxilla (Drake and Poor) 

Leptopharsa pauxilla Drake and Poor 1939a, p. 32 [Argentina]. 
Vatiga pauxilla: Drake and Ruhoff 1960c, p. 29. 
Type: Holotype 9, macropterous; Empedrado, Corrientes, Argentina; 
Drake Coll. (USNM). 
Distribution: Argentina. 
Host Plant: Unrecorded. 

* Vatiga sesoris (Drake and Hambleton) 

Tigava sesoris Drake and Hambleton 1942b, p. 77 [Brazil]. 

Vatiga sesoris: Drake and Hambleton 1946a, p. 10. 
Type: Holotype o", macropterous; Sao Paulo, Brazil; Drake Coll. 
(USNM). 
Distribution: Brazil. 
Host Plant: Unrecorded. 

*Vatiga variana Drake and Hambleton 

Vatiga variana Drake and Hambleton 1946a, p. 11 [Brazil]. 

Type: Holotype 9, macropterous; Rio Grande do Sul, Brazil; Drake 
Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

*Vatiga vicosana Drake and Hambleton 

Vatiga vicosana Drake and Hambleton 1946a, p. 10 [Brazil]. 

Type: Holotype o*, macropterous; Vigosa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 

Distribution: Brazil. 

Host Plant: Unrecorded. 

Genus XENOTINGIS Drake 

Xenotingis Drake 1923a, p. 105; 1927a, p. 57.— Drake and Ruhoff 
1960a, p. 88. 
Type Species: Xenotingis horni Drake. 
*Xenotingis bakeri Drake 

Xenotingis bakeri Drake 1927a, p. 57 [Luzon]. 
Type: Holotype d% macropterous; Los Bafios, Phillippine Islands; 
Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 
Xenotingis horni Drake 

Xenotingis horni Drake 1923a, p. 105, fig. 1 [Taiwan]; 1927a, pp. 57, 
58.— Esaki 1926, p. 163.— Takeya 1951a, p. 21. 
Type: Holotype 9; Kosempo, Formosa, Japan; Horn Coll. 
Distribution: Taiwan. 
Host Plant: Unrecorded. 



LACE BUGS OF THE WORLD 427 

*Xenotingis luzonana Drake 

Xenotingis luzonana Drake 1954f, p. 8 [Luzon]. 
Type: Holotype 9, macropterous; Mount Makiling, Luzon, Philippine 
Islands; Drake Coll. (USNM). 
Distribution: Philippine Islands (Luzon). 
Host Plant: Unrecorded. 

*Xenotingis malkini Drake 

Xenotingis malkini Drake 1948e, p. 149 [New Guinea]; I960, p. 378. 

Type: Holotype d\ macropterous; Hollandia, Dutch New Guinea; 
USNM. 

Distribution: New Guinea (Netherlands). 

Host Plant: Unrecorded. 

*Xenotingis papuana Drake 

Xenotingis papuana Drake 1954f, p. 9 [Papua]; 1960, p. 379, fig. 27 
[New Guinea]. 
Type: Holotype cf; Kokoda, Papua; British Mus. 
Distribution: New Guinea (Papua; Northeast). 
Host Plant: Unrecorded. 
Note: Taken at light [Drake I960]. 

Genus XYNOTINGIS Drake 

Xynotingis Drake 1948a, p. 8.— Drake and Ruhoff 1960a, p. 88. 
Type Species: Xynotingis hoytona Drake. 

*Xynotingis hoytona Drake 

Xynotingis hoytona Drake 1948a, p. 8, fig. 3 [Japan]. — Takeya 1951a, 
p. 19; 1953d, p. 173, pi. 6, fig. 3.— Drake and Maa 1953, p. 95. 
Type: Holotype 9, macropterous; Japan; Drake Coll. (USNM). 
Distribution: Japan. 
Host Plant: Unrecorded. 

Genus ZATINGIS Drake 

Zatingis Drake 1928a, p. 44.— Drake and Ruhoff 1960a, p. 88. 
Type Species: Zatingis extraria Drake. 

*Zatingis extraria Drake 

Zatingis extraria Drake 1928a, p. 45 [Paraguay]; 1935, p. 16. 

Type: Holotype o 71 , macropterous; S. Bernardino, Paraguay; Drake 
Coll. (USNM). 

Distribution: Paraguay. 

Host Plant: Unrecorded. 

Genus ZEIRATINGIS Drake and Ruhoff 

Zeiratingis Drake and Ruhoff 1961b, p. 173. 
Type Species: Z e ^ ra ^ n S^ s pwosa Drake and Ruhoff. 

681-552—64^—29 



428 U.S. NATIONAL MUSEUM BULLETIN 243 

*Zeiratingis dissita Drake and Ruhoff 

Zeiratingis dissita Drake and Ruhoff 1961b, p. 174 [Borneo]. 

Type: Holotype d", macropterous; Sandakan, Borneo; Drake Coll. 
(USNM). 

Distribution: Greater Sunda Islands (Borneo). 

Host Plant: Unrecorded. 

*Zeiratingis peirosa Drake and Ruhoff 

Zjeiratingis peirosa Drake and Ruhoff 1961b, p. 173, figs. 18a, b [Viet- 
Nam]. 
Type: Holotype 9, macropterous; Tonkin, Viet-Nam; Drake Coll. 
(USNM). 
Distribution: Viet-Nam. 
Host Plant: Unrecorded. 

Genus ZELOTINGIS Drake and HambLtton 

^elotingis Drake and Hambleton 1946a, p. 9. — Drake and Ruhoff 
1960a, p. 89. 
Type Species: Stenocysta aspidospermae Drake and Hambleton. 

*Zelotingis aspidospermae (Drake and Hambleton) 

Stenocysta aspidospermae Drake and Hambleton 1934, p. 444 [Brazil; 
Aspidosperma melanocalyx]. — Costa Lima 1936, p. 130. — Monte 1937a, 
p. 33; 1939b, p. 78; 1941e, p. 132. 
Zjlotingis aspidospermae: Drake and Hambleton 1946a, p. 10 [Paraguay]. 
Type: Holotype 9, macropterous; Vicpsa, Minas Gerais, Brazil; Drake 
Coll. (USNM). 
Distribution: Brazil; Paraguay. 
Host Plant: Aspidosperma melanocalyx. 

Genus ZOROTINGIS Drake and Ruhoff 

£orotingis Drake and Ruhoff 1961b, p. 168. 
Type Species: ^orotingis scitula Drake and Ruhoff. 

*Zorotingis scitula Drake and Ruhoff 

Zjarotingis scitula Drake and Ruhoff 1961b, p. 169, fig. 15 [Mindanao]. 

Type: Holotype 9, macropterous; Davao, Mindanao, Philippines; Drake 
Coll. (USNM). 

Distribution: Philippine Islands (Mindanao). 

Host Plant: Unrecorded. 

Tribe Ypsotingini, new tribe 
Type Genus: Tpsotingis Drake. 



LACEBUGS OF THE WORLD 429 

Genus BISKRIA Puton 

Dictyonota {Biskria) Puton 1874c, p. 440. 

Biskria: Lethierry and Severin 1896, p. 10. — Horvath 1906a, pp. 13, 
34 (key).— Oshanin 1908, p. 413; 1912, p. 43.— Kiritshenko 1951, 
pp. 240, 245 (key).— Drake and Ruhoff 1960a, p. 40.— Stichel 1960a, 
p. 282 (key); 1960b, p. 392; 1960c, p. 131. 
Type Species: Dictyonota {Biskria) gracilicomis Puton. 

*Biskria gracilicomis (Puton) 

Dictyonota {Biskria) gracilicomis Puton 1874c, p. 440 [Algeria]. — Lethierry 

and Puton 1876, p. 32. 
Biskria gracilicomis: Horvath 1906a, p. 35. — Stichel 1960a, p. 282. 
Type: Sex unknown; Biskria, Algeria; Paris Mus. 
Distribution: Algeria; French Somaliland (Dijibouti). 
Host Plant: Unrecorded. 

Biskria hispanica Gomez-Menor 

Biskria hispanica Gomez-Menor 1955b, p. 250, figs. 1-2 [Spain]. — 
Stichel 1960a, p. 283, figs. 66, 67. 
Type: Sex unknown; El Escorial, Madrid, Spain; Inst. Ent. Madrid. 
Distribution: Spain. 
Host Plant: Unrecorded. 

Biskria horvathi Kiritshenko 

Biscria [sic] horvathi Kiritshenko 1913a, p. 413 [Turkestan]. 

Type: Holotype o 71 , macropterous; Termez, Turkestan, Russia; Lenin- 
grad Inst. 

Distribution: U.S.S.R. (Turkestan). 

Host Plant: Unrecorded. 

Biskria lepida Horvath 

Biskria lepida Horvath 1905b, p. 562 [Tunisia]; 1906a, p. 35.— Stichel 
1960a, p. 283. 
Type: Holotype 9; Feriana, Tunisia; Hungarian Mus. 
Distribution: Tunisia. 
Host Plant: Unrecorded. 

Biskria sareptana (Jakovlev) 

Dictyonota sareptana Jakovlev 1876b, p. 67, pi. 2, fig. 7 [Russia]. 
Scraulia sareptana: Lethierry and Severin 1896, p. 8. 
Biskria sareptana: Horvath 1906a, p. 35. — Kiritshenko 1951, p. 245. — 
Stichel 1960a, p. 283. 
Type: Holotype o 71 , macropterous; Sarepta, Russia; Leningrad Inst. 
Distribution: U.S.S.R. 
Host Plant: Unrecorded. 



430 U.S. NATIONAL MUSEUM BULLETIN 243 

Biskria sareptana var. adelpha Horvath 

Biskria sareptana var. adelpha Horvath 1905b, p. 563 [Russia]; 1906a, 
p. 35.— Kiritshenko 1951, p. 245.— Stichel 1960a, p. 283. 
Type: Sex unknown; Theodosia, Tauria, Russia; Hungarian Mus. 
Distribution: U.S.S.R. (Tauria). 
Host Plant: Unrecorded. 

Genus CHOROTINGIS Drake 

Chorotingis Drake 1961b, p. 111. 
Type Species: Chorotingis indigena Drake. 

Chorotingis indigena Drake Plate 51 

Chorotingis indigena Drake 1961b, p. Ill, pi. 6 [Queensland]. 

Type: Holotype cf; Saint George's Sound, Queensland, Australia; 
Austr. Mus. 

Distribution: Australia (Queensland). 

Host Plant: Unrecorded. 

Genus DEREPHYSIA Spinola 

Derephysia Spinola 1837, p. 166.— Fieber 1844, p. 99; 1861, p. 128. — • 
Douglas and Scott 1865, p. 253.— Stal 1874, p. 48.— Vollenhoven 
1878, p. 283.— Lethierry and Severin 1896, p. 10.— Horvath 1906a, 
pp. 13, 45 (key).— Oshanin 1908, p. 419; 1912, p. 43.— Stichel 1926, 
pp. 104, 108 (key); 1935, p. 348; 1938a, p. 404; 1960a, p. 290 (key); 
1960b, p. 393; 1960c, p. 132— Borner 1935, pp. 73, 76 (key).— China 
1943, p. 246.— Kiritshenko 1951, pp. 241, 246 (key).— Drake and 
Ruhoff 1960a, p. 50. 
Type Species: Tingis foliacea Fallen. 

Derephysia brevicornis Reuter 

Derephysia brevicornis Reuter 1888, p. 224 [Greece]; 1891a, p. 25. — 
Horvath 1906a, p. 46.— Stichel 1960a, p. 290. 
Type: Sex unknown; Greece; Helsin. Mus. 
Distribution: Greece. 
Host Plant: Unrecorded. 

*Derephysia crista ta (Panzer) 

Tingis cristata Panzer 1806, heft 99, tab. 19 [Germany]. — Lamarck 
1816, p. 504.— Le Peletier and Serville 1828, p. 653— Brulle 1835, 
p. 338, pi. 25, fig. 5 [France].— Burmeister 1835, p. 259— Herrich- 
Schaeffer 1835, p. 58; 1838, p. 70, pi. 130, figs, i-1— Blanchard 1840, 
p. 112. 
Galeatus cristata: Westwood 1840b, p. 121 [England]. 
Derephysia cristata: Fieber 1844, p. 100, pi. 8, figs. 28-33 [Czechoslovakia; 
Austria]; 1861, p. 128.— Scholz 1847, p. 121.— Herrich-Schaeffer 
1850, p. 161.— Jakovlev 1874a, p. 10 [Russia].— Puton 1879c, p. 



LACEBUGS OF THE WORLD 431 

104.— Reuter 1891b, p. 184— Hiieber 1893, p. 319.— Horvath 
1906a, pp. 46, 47 [Hungary; Siberia; Artemisia campestris]. — Muzik 
1907, p. 56.— Stichel 1926, p. 109; 1938a, p. 406 [Poland]; 1960a, 
p. 292, fig. 84.— Scholte 1935, p. 31 [Netherlands].— Guide 1938, 
p. 269.— Hoberlandt 1942, p. 125.— Kiritshenko 1951, p. 246. 
Tingis {Derephysia) cristata: Flor 1860, p. 362 [Estonia; Latvia]. 

Type: Sex unknown; Mannheim, Germany; Berlin Mus. 

Distribution: Germany; Czechoslovakia; Austria; Hungary; Poland; 
Netherlands; France; England; U.S.S.R. (Estonia; Latvia; Siberia). 

Host Plant: Artemisia campestris. 
*Derephysia foliacea (Fallen) 

Tingis foliacea Fallen 1807, p. 39 {Chrysanthemum leucanthemum]; 1829, 
p. 149.— Herrich-Schaeffer 1830, heft 118, tab. 18 [Germany]; 
1835, p. 58; 1838, p. 70, pi. 129, fig. d; pi. 130, figs, m, n. 
Derephysia foliacea: Spinola 1837, p. 116. — Fieber 1844, p. 99, pi. 8, 
figs. 23-27 [Sweden; Czechoslovakia; Italy; Austria; Artemisia 
campestris]; 1861, p. 128 [Statice armeria]. — Sahlberg 1848, p. 135 
[Finland].— Herrich-Schaeffer 1850, p. 161.— Costa 1855a, p. 11; 
1855b, p. 233.— Mayr 1858, p. 572.— Frey-Gessner 1865, p. 232 
[Switzerland]. — Douglas and Scott 1865, p. 254, pi. 9, fig. 4. — 
Garbiglietti 1869, p. 275.— Gredler 1870, p. 76.— Jakovlev 1874a, 
p. 10 [Russia]; 1880b, p. 101; 1893, p. 294.— Siebke 1874, p. 23 
[Norway].— Saunders 1875, p. 250; 1892, p. 130, pi. 12, fig. 4 [ivy; 
moss]. — Reiber and Puton 1876, p. 68. — Vollenhoven 1878, p. 284, 
pi. 10, figs. 4, 4a [Netherlands]. — Puton 1879c, p. 104. — Reuter 
1882b, p. 117.— d'Antessanty 1890, p. 31.— Hiieber 1893, p. 317. — 
Horvath 1906a, pp. 45, 46 [Turkey; Turkestan; Siberia; Thymus 
serpyllus]; 1906d, p. 2.— Muzik 1907, p. 56.— Oshanin 1908, p. 419 
[Algeria]. — Jensen-Haarup 1912, p. 154, fig. 97 [Denmark]. — 
Schumacher 1914, p. 257.— Lindberg 1919, p. 42; 1927, p. 15 — 
Reichensperger 1920, p. 61.— Sahlberg 1920, p. 82.— Butler 1923, 
p. 208 [ash; Chenopodium; Wales; Scotland; Ireland]. — Stichel 1926, 
p. 108, fig. 299 [Echium; Quercus]; 1938a, p. 406 [Poland]; 1960a, p. 
291, fig. 82 [Bulgaria; Armeria vulgaris; Echium vulgar e; Artemisia 
vulgaris; Hedera helix; Lonicera; Vaccinium myrtillus; Juncus; Carex]. — de 
Seabra 1931, p. 418, figs. 490, 491 [Portugal].— Borner 1935, p. 76, 
fig. 114.— Scholte 1935, p. 30, fig. 5 [Plantago media].— Guide 1938, 
p. 268, fig. [Thymus].— Hoberlandt 1942, p. 125; 1943a, p. 118; 
1943b, p. 123; 1955, p. 87.— Blote 1945, p. 82 [Belgium; Tunisia; 
Yugoslavia]. — Kiritshenko 1951, p. 246. — Novak and Wagner 1951, 
p. 70 [Cydoma vulgaris] .— Stehlik 1952, p. 205.— Mancini 1953b, 
p. 186 [Albania]; 1953d, p. 16.— Stusak 1957b, pp. 136, 140, figs. 5, 
13c [Climacium dendroides]; 1961a, p. 79, figs. 6, 16. — Southwood and 
Leston 1959, p. 146, pi. 18, fig. 3. 



432 U.S. NATIONAL MUSEUM BULLETIN 2 43 

Campylosteira {Derephysia) foliacea: Kolenati 1856, p. 432 [Artemisia; 

Caucasus] . 
Tingis {Derephysia) foliacea: Flor 1860, p. 364 [France; England; 
Estonia; Latvia]. 
Type: Unknown. 

Distribution: England; Wales; Scotland; Ireland; Belgium; Nether- 
lands; France; Portugal; Italy; Switzerland; Czechoslovakia; Germany; 
Austria; Poland; Sweden; Norway; Denmark; Finland; U.S.S.R. (Estonia; 
Latvia; Caucasus; Turkestan; Siberia); Albania; Bulgaria; Yugoslavia; 
Turkey; Algeria; Tunisia. 

Host Plants: Armeria vulgaris; Artemisia campestris; Artemisia vulgaris; 
Artemisia sp.; Car ex sp.; Chenopodium sp.; Chrysanthemum leucanthemum; 
Climacium dendroides; Cydonia vulgaris; Echium vulgar e; Echium sp.; Hedera 
helix; Juncus sp.; Lonicera sp.; Plantago media; Quercus sp.; Statice ameria; 
Thymus serpyllus; Thymus sp.; Vaccinium myrtillus; moss; ivy; ash. 
Note: Eggs [Stusak]. 

Derephysia foliacea var. biroi Horvath 

Derephysia foliacea var. biroi Horvath 1896, p. 326 [Yugoslavia]; 1906a s 
p. 47 [Hungary]. 
Type: Holotype 9; Croatia; Hungarian Mus. 
Distribution: Yugoslavia; Hungary. 
Host Plant: Unrecorded. 

*Derephysia gardneri Drake and Poor 

Derephysia gardneri Drake and Poor 1936b, p. 148 [India]. 

Type: Holotype 9, macropterous; Chakrata District, Bodyar, India; 
Drake Coll. (USNM). 

Distribution: India. 

Host Plant: Unrecorded. 

Derephysia lugens Horvath 

Derephysia lugens Horvath 1902a, p. 593 [Yugoslavia]; 1906a, p. 47 
[Bulgaria].— Stichel 1938a, p. 406 [Albania]; 1960a, p. 291 — 
Mancini 1953b, p. 186. 

Type: Holotype o 71 ; Vranja, Serbia; Hungarian Mus. 

Distribution: Yugoslavia; Albania; Bulgaria. 

Host Plant: Unrecorded. 

Derephysia nigricosta Horvath 

Derephysia nigricosta Horvath 1905a, p. 272 [Spain]; 1906a, p. 47 — 

Ribaut 1937, p. 247 [France].— Stichel 1960a, p. 292. 
Derephysia nigricostata [sic]: Gonzalez 1948, p. 50. 
Type: Sex unknown; Spain; Hungarian Mus. 
Distribution: Spain; Portugal; France. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 433 

Derephysia rectinervis Puton 

Derephysia rectinervis Puton {in Puton and Lethierry 1887, p. 304) 
[Algeria].— Horvath 1906a, p. 47.— Stichel 1960a, p. 291. 
Type: Sex unknown; Mecheria, Algeria; Paris Mus. 
Distribution: Algeria. 
Host Plant: Unrecorded. 
Derephysia sinuatocollis Puton 

Derephysia joliacea var. sinuatocollis Puton 1879c, p. 104 [France]. — 

Chicote 1880, p. 189 [Spain]. 
Derephysia sinuatocollis: Horvath 1906a, p. 46. — Stichel 1960a, p. 290 
[Italy]. 
Type: Sex unknown; Hautes-Pyrenees, France; Paris Mus. 
Distribution: France; Spain; Italy. 
Host Plant: Unrecorded. 

Genus DICTYONOTA Curtis 

Dictyonota Curtis 1827, tab. 154.— Laporte 1833, p. 50.— Fieber 1844, 
p. 91; 1861, p. 126.— Douglas and Scott 1865, p. 255.— Garbiglietti 
1869, p. 274.— Walker 1873a, p. 177.— Stal 1874, p. 49.— Horvath 
1874b, p. 432; 1906a, pp. 13, 36 (key).— Oshanin 1908, p. 413; 1912, 
p. 43.— Schumacher 1914, p. 257.— Parshley 1923b, p. 699.— Stichel 
1926, pp. 103, 107 (key); 1935, p. 348; 1938a, p. 404; 1960a, p. 284 
(key); 1960b, p. 392; 1960c, p. 131.— de Seabra 1931, pp. 406, 413 
(key).— Scholte 1935, p. 21 (key) .— Borner 1935, pp. 73, 75 (key).— 
Guide 1938, p. 262.— China 1943, p. 246.— Blote 1945, p. 81.— Hurd 
1946, p. 461.— Kiritshenko 1951, pp. 240, 245 (key) .—Singer 1952, 
p. 49.— Mancini 1953d, p. 15; 1954b, p. 1 1 .— Gomez-Menor 1955b, 
p. 248.— Drake and Ruhoff 1960a, p. 51. 

Scraulia Stal 1874, p. 50. — Lethierry and Severin 1896, p. 8 (in part). — 
Kirkaldy 1900, p. 241.— Drake and RuhofF 1960a, p. 81. 
Type Species: Dictyonota strichnocera Fieber. 

Subgenus DICTYONOTA (DICTYONOTA) Curtis 

Dictyonota Curtis 1827, tab. 154. 

Scraulia Stal 1874, p. 50. 

Dictyonota {Dictyonota): Horvath 1906a, p. 36. — Oshanin 1908, p. 413; 
1912, p. 43.— Drake and Ruhoff 1960a, p. 51. 
Type Species: Dictyonota strichnocera Fieber. 
*Dictyonota albipennis von Baerensprung 

Dictyonota albipennis von Baerensprung 1858, p. 207, pi. 2, fig. 12 
[Italy].— Fieber 1861, p. 127.— Puton 1874a, p. 227; 1879c, p. 102 
(in part) [France; Corsica]; 1886a, p. 5 [Tunisia]. — Chicote 1880, 
p. 189 [Spain].— Horvath 1906a, p. 39.— Lindberg 1932a, p. 41 
[Morocco; Genista anglica]. 



434 U.S. NATIONAL MUSEUM BULLETIN 243 

Scraulia albipennis: Lethierry and Severin 1896, p. 8 (in part). 
Type: Sex unknown; Piedmont, Italy; Berlin Mus. 
Distribution: Italy; Corsica; France; Spain; Morocco; Tunisia. 
Host Plant: Genista anglica. 

*Dictyonota fuliginosa Costa 

Dyctionota [sic] fuliginosa Costa 1855a, p. 10, pi. 6, fig. 5; 1855b, p. 

232.— Stichel 1960d, p. 120. 
Dictyonota fuliginosa: Costa 1863, pi. 1, fig. 6. — Garbiglietti 1869, p. 
275 [Italy].— Pu ton 1874a, p. 227; 1879c, p. 101 [Corsica].— Saunders 

1875, p. 251; 1892, p. 129, pi. 12, fig. 3 [broom].— Reiber and Puton 

1876, p. 68.— Ferrari 1878, p. 84.— Chicote 1880, p. 189 [Spain].— 
Hiieber 1893, p. 316— Horvath 1906a, p. 40 [Belgium; Netherlands; 
Spartium scopariuni]; 1906d, p. 2. — Schumacher 1914, p. 257 [Saro- 
thamus].— Reichensperger 1920, p. 61. — Butler 1923, p. 207. — Stichel 
1926, p. 108; 1960a, p. 286, fig. 73 [Czechoslovakia].— de Seabra 
1931, p. 416, figs. 486(1), 488 [Portugal].— Scholte 1935, p. 23, fig. 
2.— Gonzalez 1948, p. 49.— Seidenstiicker 1954, p. 235 [Turkey].— 
Massee 1954, p. 260. — Southwood and Leston 1959, p. 145. — - 
Scudder 1960, p. 22 [B.C.]. 

Dictyonota fieberi Fieber 1861, p. 127 [France]. — Frey-Gessner 1865, 
p. 232 [Switzerland].— Douglas and Scott 1868, p. 245, pi. 2, fig. 2 
[England].— Walker 1873a, p. 177 [Germany]. 
Scraulia fieberi: Stal 1874, p. 50. 
Type: Unknown. 

Distribution: England; Netherlands; Belgium; France; Spain; Portugal; 
Corsica; Italy; Switzerland; Czechoslovakia; Germany; Turkey; Canada 
(B.C.). 
Host Plants: Sarothamus sp.; Spartium scoparium; broom. 
Note: Dictyonota fieberi was a manuscript name of Forster used by Fieber 
(1861) thus giving Fieber the credit for the species. Species recently 
introduced in Canada. 

*Dictyonota marmorea von Baerensprung 

Dictyonota marmorea von Baerensprung 1858, p. 206, pi. 2, fig. 11 
[Spain].— Fieber 1861, p. 127.— Puton 1874a, p. 227; 1879c, p. 102 
[France; Corsica]. — Ferrari 1878, p. 84. — Horvath 1906a, p. 40. — 
Lindberg 1932a, p. 41 [Morocco]. 
Scraulia marmorea: Stal 1874, p. 50. 
Type: Sex unknown; Andalusia; Berlin Mus. 
Distribution: Spain; France; Corsica; Morocco. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 435 

*Dictyonota mitoris Drake and Hsiung 

Dictyonota mitoris Drake and Hsiung 1936, p. 287 [China; ash]. 

Type: Holotype cf, macropterous; Tientsin, China; Drake Coll. 
(USNM). 

Distribution: China. 

Host Plant: Chinese ash. 

*Dictyonota pakistana Drake and Maldonado 

Dictyonota pakistana Drake and Maldonado 1959, p. 25, fig. 1 [Paki- 
stan]. — Drake and Davis 1960, fig. 35. 
Type: Holotype o 71 , macropterous; Lahore, West Pakistan; Drake Coll. 
(USNM). 
Distribution: Pakistan. 
Host Plant: Unrecorded. 
Note: Taken at light [Drake and Maldonado]. 

*Dictyonota pulchella Costa 

Dictyonota pulchella Costa 1863, p. 9, pi. 1, fig. 7 [Italy]. — Horvath 
1906a, p. 39 [Spain; Algeria; Mauritania; Genista scorpius]; 1906d, 
p. 3.— Oshanin 1908, p. 414 [Morocco; Tunisia].— Stichel 1926, 
p. 108; 1960a, p. 284, fig. 70 [Bulgaria]; 1960d, p. 122.— de Seabra 
1931, p. 415, figs. 486 (3-5), 487 [Ulex].— Ribaut 1932, p. 576.— 
Guide 1938, p. 263 [Genista]. 
Dictyonota aubei Signoret 1865, p. 118 [France]. 
Scraulia aubei: Stal 1874, p. 50. 

Dictyonota albipennis (not von Baerensprung) : Ferrari 1878, p. 84. — 
Puton 1879c, p. 102 (in part).— Rey 1893, p. 97. 
Type: Sex unknown; Italy; Naples Mus. 

Distribution: Italy; France; Spain; Portugal; Bulgaria; Algeria; 
Morocco; Mauritania; Tunisia. 
Host Plants: Genista scorpius; Genista sp.; Ulex sp. 

*Dictyonota strichnocera Fieber 

Dictyonota eryngii (not Latrielle): Curtis 1827, tab. 154, section iv 
[England; Ulex]. 

Tingis crassicornis (not Fallen): Herrich-Schaeffer 1835, p. 57. 

Dictyonota crassicornis (not Fallen): Herrich-Schaeffer 1838, p. 74, 
pi. 129, fig. B. 

Dictyonota strichnocera Fieber 1844, p. 95, pi. 8, figs. 4-7 [Czechoslovakia; 
Austria; Yugoslavia]; 1861, p. 127 [Germany]. — Herrich-Schaeffer 
1850, p. 157.— Mayr 1858, p. 571 [Hungary].— Costa 1863, p. 8, 
pi. 1, fig. 5 [Italy]. — Douglas and Scott 1865, p. 256 [moss]. — 
Frey-Gessner 1865, p. 232 [Switzerland]. — Gredler 1870, p. 76. — 
Jakovlev 1876b, p. 67 [Russia].— Saunders 1875, p. 251; 1892, p. 129, 
pi. 12, fig. 2 [broom].— Vollenhoven 1878, p. 269, pi. 22, fig. 8 
[Netherlands].— Ferrari 1878, p. 84.— Puton 1879c, p. 102 [France].— 



436 U.S. NATIONAL MUSEUM BULLETIN 243 

Chicote 1880, p. 189 [Spain].— Rey 1893, p. 97.— Hiieber 1893, 
p. 316.— Kirkaldy 1900, p. 241— Horvath 1906a, p. 40 [Belgium; 
Rumania; Spartium scoparium]; 1906d, p. 241. — Muzik 1907, p. 55. — 
Jensen-Haarup 1912, p. 153 [Denmark] .—Butler 1922, p. 179; 
1923, p. 206, fig.— Stichel 1926, p. 108; 1938a, p. 404 [Poland]; 
1960a, p. 285 [Bulgaria; Sarothamus scoparius; Pinus silvestris; Larix 
decidua].— Scholte 1935, p. 23, fig. 1 .— Hoberlandt 1943, p. 118 — 
Kiritshenko 1951, p. 245— Stusak 1957b, p. 140; 1961a, p. 77, 
fig. 5H — Southwood and Leston 1959, p. 145, pi. 18, fig. 4; pi. 21, 
figs. 4; pi. 45, fig. 5; fig. 45. 
Scraulia strichnocera: Stal 1874, p. 50. 
Dictyonota idonea Jakovlev 1903c, p. 291 [Ukraine]. 
Type: Unknown. 

Distribution: England; Belgium; Netherlands; France; Spain; Italy; 
Switzerland; Czechoslovakia; Germany; Yugoslavia; Bulgaria; Denmark; 
Poland; Hungary; Austria; Rumania; U.S.S.R. (Ukraine). 

Host Plants: Larix decidua; Pinus silvestris; Sarothamus scoparius; Spartium 
scoparium; Ulex sp.; broom; moss. 

Note: Life history study [Butler 1922]; eggs [Butler 1923]; eggs and im- 
mature stages [Stusak]. 
Dictyonota teydensis Lindberg 

Dictyonota teydensis Lindberg 1936b, p. 29, pi. 1, fig. 4 [Canary Islands]. 
Type: Sex unknown; Canary Islands; Helsin. Mus. 
Distribution: Canary Islands. 
Host Plant: Unrecorded. 

Subgenus DICTYONOTA (ALCLETHA) Kirkaldy 

Dictyonota (not Curtis): Stal 1874, p. 49. — Lethierry and Severin 1896, 

p. 7. 
Alcletha Kirkaldy 1900, p. 241. 
Dictyonota {Alcletha) : Horvath 1960a, p. 40 (key). — Oshanin 1908, 

p. 415; 1912, p. 43.— de Seabra 1931, pp. 414, 416 (key) .— Borner 

1935, p. 75 (key).— Stichel 1935, p. 348; 1960a, p. 286.— China 

1943, p. 246.— Drake and Ruhoff 1960a, p. 52. 
Type Species: Acanthia tricornis Schrank. 

*Dictyonota (Alcletha) aethiops Horvath 

Dictyonota truncaticollis ? (not Costa): Puton 1879c, p. 100 [France; 
Corsica]. 

Dictyonota aethiops Horvath 1905b, p. 563 [Algeria]. — Novak and Wag- 
ner 1951, p. 70 [Yugoslavia]. 

Dictyonota {Alcletha) aethiops: Horvath 1906a, p. 41 [Spain]. — Guide 
1938, pp. 263, 266.— Stichel 1906a, p. 286 [Sardinia]. 

Alcletha aethiops: Vidal 1937, p. 197 [Morocco]. 



LACEBUGS OF THE WORLD 437 

Type: Unknown. 

Distribution: Spain; France; Corsica; Sardinia; Yugoslavia; Algeria; 
Morocco. 
Host Plant: Unrecorded. 

*Dictyonota (Alcletha) aridula Jakovlev 

Dictyonota aridula Jakovlev 1902a, p. 66 [Crimea]. — Drake and Runoff 

1962d, p. 489. 
Dictyonota tricornis (not Schrank): Horvath 1905b, p. 563 [Hungary]. 
Dictyonota {Alcletha) aridula: Horvath 1906a, p. 42 [Greece]. — Kirit- 
shenko 1951, p. 246. — Seidenstiicker 1954, p. 235 [Turkey]. — Hober- 
landt 1955, p. 87.— Stichel 1960a, p. 288. 
Dictyonota maroccana Ribuat 1939, p. 186 [Morocco]. 
Dictyonota (Alcletha) maroccana: Drake and Ruhoff 1962c, p. 141. 
Type: Holotype 9, macropterous; Eupatoria, Crimea, Russia; Lenin- 
grad Inst. 
Distribution: U.S.S.R. (Crimea); Turkey; Greece; Hungary; Morocco. 
Host Plant: Unrecorded. 

*Dictyonota (Alcletha) lugubris Fieber 

Dictyonota lugubris Fieber 1861, p. 126 [Yugoslavia]. 

Dictyonota (Alcletha) lugubris: Horvath 1906a, p. 41, pi. 1, fig. 6. — Stichel 
1960a, p. 286. 
Type: Sex unknown; Serbia; Vienna Mus. 
Distribution : Yugoslavia. 
Host Plant: Unrecorded. 

*Dictyonota (Alcletha) pusana Drake and Maa 

Dictyonota (Alcletha) pusana Drake and Maa 1955, p. 6 [India]. 
Dictyonota pusana: Lindberg 1958a, p. 78 [Borneo; Cape Verde Islands]. 
Type: Holotype 9; Pusa, Bengal, India; British Mus. 
Distribution: India; Greater Sundra Islands (Borneo); Cape Verde 
Islands (Sao Nicolao). 
Host Plant: Unrecorded. 
Note: Taken on "potatoes in store" [Drake and Maa]. 

*Dictyonota (Alcletha) tricornis (Schrank) 
Acanthia tricornis Schrank 1801, p. 67. 
Tingis crassicornis Fallen 1807, p. 38 [Sweden]; 1829, p. 147. — Zetterstedt 

1828, p. 480 [Lapland]; 1840, p. 269.— Brulle 1835, p. 339, pi. 25, 

fig. 6. 
Tingis erythrophthalma Germar and Kaulfuss 1817, fasc. 3, tab. 25. — 

Schilling 1838, p. 106. 
Dictyonota crassicornis: Curtis 1827, pi. 4, tab. 154 [England]. — Fieber 

1844, p. 92, pi. 7, figs. 42-47 [Austria; Czechoslovakia; Italy; Artemisia 

campestris]; 1861, p. 127 [Thymus]. — Sahlberg 1848, p. 134 [Finland]. — 



438 U.S. NATIONAL MUSEUM BULLETIN 243 

Herrich-Schaeffer 1850, p. 157.— Flor 1860, p. 358 [Estonia; 
Latvia]. — Douglas and Scott 1865, p. 255, pi. 9, fig. 5. — Frey- 
Gessner 1865, p. 232 [Switzerland]. — Jakovlev 1869, p. Ill [Russia]; 
1880b, p. 101; 1893, p. 294.— Siebke 1874, p. 23 [Norway].— StM 
1874, p. 49.— Saunders 1875, p. 250; 1892, p. 128, pi. 12, fig. 1 
[Ireland].— Vollenhoven 1878, p. 268, pi. 9, figs. 4, 4a [Netherlands].— 
Puton 1879c, p. 100.— Bolivar and Chicote 1879, p. 166 [Spain].— 
Rey 1893, p. 97.— Hiieber 1893, p. 313. 

Tingis pilicornis Herrich-Schaeffer 1830, heft 118, tab. 17 [Germany]; 
1835, p. 57. 

Piesma marginatum (not Wolff): Burmeister 1835, p. 258 [Hungary]. 

Dictyesthes (monomial) Amyot 1846, p. 181. 

Dictyonota pilicornis: Herrich-Schaeffer 1838, p. 74, pi. 129, fig. 401. — 
Fieber 1844, p. 95, pi. 8, figs. 8-9. 

Tingis mar ginata: Blanchard 1840, p. 112 [France]. 

Dictyonota erythrophthalma: Fieber 1844, p. 94, pi. 8, figs. 1-3 [Senecio 
jacobaea]; 1861, p. 127. — Herrich-Schaeffer 1850, p. 158. 

Dyctinota [sic] mar ginata: Costa 1847a, p. 21; 1847c, p. 161. 

Campylosteira (Derephysia) pilicornis: Kolenati 1856, p. 432 (no. 215) 
[Caucasus]. 

Dictyonota truncaticollis Costa 1863, p. 7, pi. 1, fig. 4. — Stal 1874, p. 49. — 
Puton 1886a, p. 4 [Tunisia].— Stichel 1960d, p. 122. 

Monanthia lurida Garbiglietti 1869, p. 274. 

Dictyonota dictyesthes Garbiglietti 1869, p. 275. — Drake and Ruhoff 
1960a, p. 7. 

Dictyonota erythrocephala Garbiglietti 1869, p. 275. 

Dictyonota cr assicornis var. erythrophthalma: Puton 1879c, p. 100. 

Alcletha tricornis: Kirkaldy 1900, p. 241. 

Dictyonota {Alcletha) tricornis: Plorvath 1906a, p. 41 [Iran; Turkey; 
Siberia; Turkestan; Armenia; Mentha; Artemisia]. — Butler 1923, p. 205 
[Algeria; Wales; Cytisus scoparius]. — de Seabra 1931, p. 416, figs. 
486(2), 489 [Portugal].— Guide 1938, pp. 262, 265, fig.— Kiritshenko 
1951, p. 245.— Stehlik 1952, p. 205.— Hoberlandt 1955, p. 87.— 
Stichel 1960a, p. 287, fig. 76 [Balearic; Crete; Greece; Bulgaria; 
Sardinia; Sicily; Belgium; Hieracium: Chrysanthemum; Echium vulgare]. 

Dictyonota tricornis: Horvath 1906d, p. 1; 1916, p. 9 [Albania]. — Muzik 
1907, p. 55.— Reuter 1908, p. 89.— Jensen-Haarup 1912, p. 153, 
fig. 96 [Denmark].— Sahlberg 1920, p. 81.— Stichel 1926, p. 108, 
fig. 298 [Verbascum; Echium]; 1938a, p. 404 [Poland].— Scholte 1935, 
p. 23, fig. 3. — Novak and Wagner 1951, p. 70 [Yugoslavia]. — Bator 
1953, p. 325, pi. 2, fig. 3.— Strawinski 1953, p. 380.— Southwood 
and Leston 1959, p. 145, pi. 21, fig. 9; fig. 44.— Drake and Ruhoff 
1962c, p. 141. 



LACEBUGS OF THE WORLD 439 

Dictyonota tricornis var. americana Parshley 1916a, p. 164 [Maine]; 1917b, 
p. 53; 1923a, p. 103; 1923b, p. 699 [N.S.].— Blatchley 1926, p. 472.— 
Proctor 1946, p. 74.— Bailey 1951, p. 30.— Lindberg 1958, p. 78 
[Cape Verde; Heliotr opium erosum]. 
Dictyonota ifranensis Vidal 1951, p. 63 [Morocco]. 
Type: Unknown. 

Distribution: England; Ireland; Wales; Netherlands; Belgium; France; 
Portugal; Spain; Italy; Sardinia; Sicily; Balearic Islands; Switzerland; 
Czechoslovakia; Austria; Germany; Poland; Denmark; Sweden; Norway; 
(Lapland); Finland; U.S.S.R. (Estonia; Latvia; Caucasus; Siberia; 
Turkestan; Armenia); Hungary; Albania; Greece; Crete; Bulgaria; 
Yugoslavia; Turkey; Iran; Algeria; Tunisia; Morocco; Cape Verde Islands 
(Sao Nicolao); U.S. (Maine); Canada (N.S.; N.B.). 

Host Plants: Artemisia campestris; Artemisia sp.; Chrysanthemum sp.; 
Cytisus scoparius; Echium vulgare; Echium sp.; Heliotropium erosum; Hieracium 
sp.; Mentha sp.; Sedum acre; Senecio jacobaea; Thymus sp.; Verbascum sp. 

Note: (1) Myrmecophile, nymphs and adults found in nests of Myrmica 
rubra [Butler]; Lasius niger [Sahlberg]; (2) introduced into Canada and U.S.; 
(3) see Drake and Ruhoff 1960a, p. 7 re monomial system. 

Dictyonota (Alcletha) tricornis var. cicur Horvath 

Dictyonota tricornis var. cicur Horvath 1905b, p. 563 [Hungary]. — Dobsik 

1951, p. 40. 
Dictyonota (Alcletha) tricornis var. cicur: Horvath 1906a, p. 42. — Stichel 
1960a, p. 287 [Czechoslovakia]. 
Type: Holotype o 71 ; Kecskemet, Hungary; Hungarian Mus. 
Distribution: Hungary; Czechoslovakia. 
Host Plant: Unrecorded. 

Subgenus DICTYONOTA (ELINA) Ferrari 

Dictyonota (Elina) Ferrari 1878, p. 84. — Horvath 1906a, p. 44. — 
Oshanin 1908, p. 417; 1912, p. 43.— Stichel 1935, p. 348; 1960a, 
p. 288.— Drake and Ruhoff 1960a, p. 52. 

Elina: Lethierry and Severin 1896, p. 9. 
Type Species: Dictyonota beckeri Jakovlev. 

*Dictyonota (Elina) beckeri Jakovlev 

Dictyonota beckeri Jakovlev 1871, p. 25 [Russia]. 

Dictyonota (Elina) beckeri: Ferrari 1878, p. 84. — Puton 1895, p. 86. — 
Horvath 1906a, p. 44 [Rumania]. — Kiritshenko 1951, p. 246. 
Type: Holotype 9, brachypterous; Astrakhan, Russia; Leningrad Inst. 
Distribution: U.S.S.R.; Rumania. 
Host Plant: Unrecorded. 



440 U.S. NATIONAL MUSEUM BULLETIN 243 

Dictyonota (Elina) fuentei Puton 

Dictyonota {Elina) fuentei Puton 1895, pp. 86, 87 [Spain]. — Horvath 
1906a, p. 44.— Gomez-Menor 1955b, p. 256, fig. 4.— Stichel 1960a, 
p. 288, fig. 78. 
Type: Sex unknown; Pozuelo de Calatrava, Prov. de Ciudad Real, 
Spain; Paris Mus. 
Distribution: Spain. 
Host Plant: Unrecorded. 

*Dictyonota (Elina) henschi Puton 

Dictyonota putoni (not Stal) Ferrari 1878, p. 85. 

Dictyonota {Elina) henschi Puton 1892, p. 72 [Hungary]; 1895, p. 87. — 
Horvath 1906a, p. 44.— Guide 1938, p. 266.— Kiritshenko 1951, 
p. 246. 
Dictyonota {Elina) putoni: Puton 1895, p. 87. 
Dictyonota {Elina) eupatoriae Jakovlev 1902a, p. 67 [Russia]. 
Dictyonota henschi: Horvath 1905b, p. 565 [Italy]. — Stichel 1926, p. 108; 
1938a, p. 404 [Turkey]. 
Type: Sex unknown; Budapest, Hungary; Paris Mus. 
Distribution: Hungary; Italy; Turkey; U.S.S.R. (Crimea). 
Host Plant: Unrecorded. 

Dictyonota (Elina) iberica Horvath 

Dictyonota {Elina) iberica Horvath 1905b, p. 564 [Spain]; 1906a, p. 44. — 
Stichel 1960a, p. 288. 
Type: Sex unknown; Sierra de Espuna, Spain; Hungarian Mus. 
Distribution: Spain. 
Host Plant: Unrecorded. 

Dictyonota (Elina) marqueti Puton 

Dictyonota {Elina) marqueti Puton 1879b, p. 297 [France]; 1879c, p. 103; 
1895, p. 86.— Horvath 1906a, p. 45.— Stichel 1960a, p. 289. 
Type: Sex unknown; Banyuls-sur-Mer, France; Paris Mus. 
Distribution: France. 
Host Plant: Unrecorded. 

Dictyonota (Elina) nevadensis Gomez-Menor 

Dictyonota {Elina) nevadensis Gomez-Menor 1955b, p. 254, fig. 3 
[Spain].— Stichel 1960a, p. 289, fig. 80. 
Type: Sex unknown; Puerto de la Ragua, Sierra Nevada, Granada, 
Spain; Inst. Ent. Madrid. 
Distribution: Spain. 
Host Plant: Unrecorded. 



LACEBUGS OF THE WORLD 441 

Dictyonota (Elina) sicardi Puton 

Dictyonota {Elina) sicardi Puton 1894, p. 115 [Tunisia]; 1895, p. 87. — 
Horvath 1906a, p. 44. 
Type: Sex unknown; Teboursouk, Tunisia; Paris Mus. 
Distribution: Tunisia. 
Host Plant: Unrecorded. 

Subgenus DICTYONOTA (KALAMA) Puton 

Dictyonota (Kalama) Puton (in Lethierry and Puton 1876, p. 34). — 
Horvath 1906a, p. 42.— Oshanin 1908, p. 416; 1912, p. 43.— Drake 
and Ruhoff 1960a, p. 53. 

Kalama: Lethierry and Severin 1896, p. 10. 
Type Species: Kalama coquereli Puton. 
Dictyonota (Kalama) brevicornis Ferrari 

Dictyonota (Kalama) putonii var. brevicornis Ferrari 1884, p. 474 [Tunisia]. 

Dictyonota (Kalama) brevicornis: Horvath 1906a, p. 43. 
Type: Sex unknown; Tunisia; Genova Mus. 
Distribution: Tunisia. 
Host Plant: Unrecorded. 
Dictyonota (Kalama) coquereli (Puton) 

Kalama coquereli Puton (in Lethierry and Puton 1876, p. 34) [Algeria]. 

Dictyonota (Kalama) coquereli: Horvath 1906a, p. 44. 
Type: Sex unknown; Algeria; Paris Mus. 
Distribution: Algeria. 
Host Plant: Unrecorded. 
*Dictyonota (Kalama) putonii Stal 

Dictyonota putonii Stal 1874, p. 50 [Algeria]. 

Dictyonota putoni [sic]: Puton 1886a, p. 12 [Tunisia]. 

Kalama putoni [sic]: Lethierry and Severin 1896, p. 10. 

Dictyonota (Kalama) putonii: Horvath 1906a, p. 43. 
Type: Sex unknown; Bone, Algeria; Stockholm Mus. 
Distribution: Algeria; Tunisia. 
Host Plant: Unrecorded. 
Dictyonota (Kalama) reuteri Horvath 

Dictyonota (Kalama) reuteri Horvath 1906a, p. 42 [Syria]. 
Type: Sex unknown; Kaifa, Syria; Helsin. Mus. 
Distribution: Syria. 
Host Plant: Unrecorded. 
*Dictyonota (Kalama) theryi (Montandon) 

Kalama theryi Montandon 1897, p. 99 [Algeria]. 

Dictyonota (Kalama) theryi: Horvath 1906a, p. 43. 
Type: Sex unknown; Algeria; Bucharest Mus. 
Distribution: Algeria. 
Host Plant: Unrecorded. 



442 U.S. NATIONAL MUSEUM BULLETIN 2 43 

Genus DICTYOTINGIS Drake 

Dictyotingis Drake 1942a, p. 8.— Drake and Runoff 1960a, p. 53. 
Type Species: Dictyotingis gibberis Drake. 
*Dictyotingis gibberis Drake 

Dictyotingis gibberis Drake 1942a, p. 8 [India]. — Drake and Davis 1960, 
fig. 37. 
Type: Holotype d\ macropterous; Madura, India; Drake Coll. 
(USNM). 
Distribution: India. 
Host Plant: Unrecorded. 
*Dictyotingis monticula Drake 

Dictyotingis monticula Drake 1956h, p. 21 [India]. 
Type: Holotype 9, macropterous; Naga Hills, Assam; Drake Coll. 
(USNM). 
Distribution: India.. 
Host Plant: Unrecorded. 

Genus EUAULANA Drake 

Euaulana Drake 1945, p. 96.— Drake and Ruhoff 1960a, p. 56. 
Type Species: Euaulana j err itincta Drake. 
*Euaulana ferritincta Drake 

Euaulana ferritincta Drake 1945, p. 96 [Queensland; Tasmania; Victoria]. 
Type: Holotype d", macropterous; Burleigh, Queensland, Australia; 
Drake Coll. (USNM). 

Distribution: Australia (Queensland; Tasmania; Victoria; New South 
Wales). 

Host Plant: Unrecorded. 
*Euaulana tasmaniae Drake 

Euaulana tasmaniae Drake 1945, p. 97 [Tasmania]; 1954c, p. 72; 1961b, 
p. Ill [New South Wales; Banksia robur minor]. — Drake and Ruhoff 
1962b, p. 250 [South Australia; Banksia]. 
Type: Holotype 9, macropterous; Hobart, Tasmania, Australia; Drake 
Coll. (USNM). 

Distribution: Australia (Queensland; Tasmania; New South Wales; 
South Australia). 
Host Plants: Cones of Banksia robur minor; Banksia sp. 

Genus YPSOTINGIS Drake 

Tpsotingis Drake 1947d, p. 229; 1958d, pp. 149-153.— Drake and 
Ruhoff 1960a, p. 88. 
Type Species: Tpsotingis sideris Drake. 
*Ypsotingis bakeri Drake 

Tpsotingis bakeri Drake 1958d, p. 149, pi. 45, figs, a, b [Borneo]. — 
Drake and Davis 1960, fig. 36. 



LACEBUGS OF THE WORLD 443 

Type: Holotype d% macropterous; Sandakan, Borneo; Drake Coll. 
(USNM). 

Distribution: Greater Sunda Islands (Borneo). 

Host Plant: Unrecorded. 
*Ypsotingis bornea Drake 

Ypsotingis bornea Drake 1958d, p. 152 [Borneo]. 

Type: Holotype 9, macropterous; Sandakan, Borneo; Drake Coll. 
(USNM). 

Distribution: Greater Sunda Islands (Borneo). 

Host Plant: Unrecorded. 
*Ypsotingis luzonana Drake Plate 52 

Ypsotingis luzonana Drake 1958d, p. 150, pi. 46, figs, a, b [Luzon]. 

Type: Holotype cf , macropterous; Mount Makiling, Luzon, Philippine 
Islands; Drake Coll. (USNM). 

Distribution: Philippine Islands (Luzon). 

Host Plant: Unrecorded. 
Ypsotingis sideris Drake 

Ypsotingis sideris Drake 1947d, p. 230, fig. [Viet-Nam]; 1947e, p. 67, 
fig. (color); 1948b, p. 45, fig. 1; 1958d, p. 149. 

Type: Holotype 9; Tonkin, Indochina; Mus. Heude. 

Distribution: Viet-Nam. 

Host Plant: LInrecorded. 
Ypsotingis vicinitas Drake (emendation) 

Ypsotingis vicinatis Drake 1948h, p. 74, fig. [Soemba]; 1958d, p. 153 
[Java]. 

Type: Holotype 9; Kananggar, Soemba Island, Netherlands East Indies; 
Leiden Mus. 

Distribution: Lesser Sunda Islands (Soemba); Greater Sunda Islands 
(Java). 

Host Plant: Unrecorded. 

Subfamily Vianaidinae Kormilev 

Yianaididae Kormilev 1955b, p. 466. — Drake and Runoff 1960a, p. 6 

(as family). 
Vianaidanae Drake and Davis 1960, pp. 8, 82, 84 (as subfamily). 
Type Genus: Vianaida Kormilev = Anommatocoris China. 
[Note: According to the rules, Vianaidinae must be retained as the 
proper subfamily name.] 

Genus ANOMMATOCORIS China 

Anommatocoris China 1945, p 126. — Drake and Davis 1960, pp. 84, 88. 
Vianaida Kormilev 1955b, p. 468. 
Type Species: Anommatocoris minutissimus China. 



444 U.S. NATIONAL MUSEUM BULLETIN 243 

* Anommatocoris coleopteratus (Kormilev) Plate 55 

Vianaida coleopterata Kormilev 1955b, p. 472, figs. 1-5 [Argentina; 

Gleditsia triacanthos]. 
Anommatocoris coleoptratus [sic]: Drake and Davis 1960, pp. 88, 89, figs. 5, 
21-23, 26 a-b, 27, 72 a-c, 73. 
Type: Holotype 9, brachypterous; Tigre, Rio Jujan, Buenos Aires 
Prov., Argentina; Drake Coll. (USNM). 
Distribution: Argentina. 

Host Plant: Roots of Gleditsia triacanthos ("imported tree called 'Acacia 
negra' in Argentina"). 

Note: Myrmecophile; found in nest of Acromyrmex lundi (Guerin) [Kor- 
milev]. 

*Anommatocoris minutissimus China Plates 54, 56c 

Anommatocoris minutissimus China 1945, pp. 127, 128, figs. 1 a-d [Trini- 
dad]. — Drake and Davis 1960, pp. 88, 90, figs. 74 a-c. — Drake and 
Froeschner 1962, fig. 3. 
Type: Sex unknown; Trinidad, British West Indies; British Mus. 
Distribution: Trinidad. 
Host Plant: Unrecorded. 

Note: Found in soil of cacao experimental plantation [China]; myrme- 
cophile, associated with undetermined species of ant [Drake and Davis]. 

*Anommatocoris zeteki Drake and Froeschner Plate 56 

Anommatocoris zeteki Drake and Froeschner 1962, p. 8, figs. 1, 2 [Canal 
Zone]. 
Type: Holotype d\ brachypterous; Barro Colorado Island, Panama 
Canal Zone; USNM. 
Distribution: Panama (Canal Zone). 
Host Plant: Unrecorded. 
Note: Myrmecophile [Drake and Froeschner]. 

Genus THAUMAMANNIA Drake and Davis 

Thaumamannia Drake and Davis 1960, p. 90. 
Type Species: Thaumamannia manni Drake and Davis. 

*Thaumamannia manni Drake and Davis Plate 53 

Thaumamannia manni Drake and Davis 1960, pp. 88, 92, figs. 75 a-c 
[Bolivia]. 
Type: Holotype 9, brachypterous; Santa Cruz, Bolivia; USNM. 
Distribution: Bolivia. 
Host Plant: Unrecorded. 
Note: Myrmecophile; found in nest of undetermined species of ant 

[Drake and Davis]. 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 1 




Alloeoderes davao Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 2 




^ .'■ 








> 



k, Li^c^> 



Phatnoma pacifica Kirkald) 



. NATIONAL MUSEUM 



BULLETIN 243, PLATE 3 





& 



/ 



U 




t Phatnoma baltica Drake (fossil in amber) 



l ? S \ 



U S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 4 




-,-, 



Zetekella pulla Drake and Plaumann 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 5 



. 




%*& 



1 



Ceratocader armatus (Hacker' 



US NATIONAL MUSEUM 



BULLETIN 243. PLATE 6 




Cantacader tener Bergroth 



U.S. NATIONAL MUSEUM 



BULLETIN 243 PLATE 





pocH 



> v iXBM^ 









Litadea delicatida China 



S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 




Litadea delicatula China, dorsal and ventral views of last nymphal instar 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 9 




Cysteochila poecilia Drake and Ruhoff 



U.S NATIONAL MUSEUM 



BULLETIN 243, PLATE 10 





K"^ 












/ TO 





Ambotingis senta (Drake and Hambleton) 



US. NATIONAL MUSEUM 



BULLETIN 243, PLATE It 




-Letter f 







R.H. 



Eteoneus peroronus Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 12 







«| 



k 



\A 



r 






m 





rr.. 



R.H. 



Alloeocysta approba Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 13 




it r r-tt-te ft 'cr#f 




H. 



Epimixia vittata Horvath 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 14 




J 



Agramma onar Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 15 




At ?H 



B 




Lullius spinifemur Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 16 




Perbrinckea brincki Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 17 



; 




Leaf down-curl on Dichapctalum cymosum by feeding ofOnymochila dichapetali (Horvaih) 




Onymochila dichapetali (Horvath) 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 18 




lH, Li$rO/ 



Copium clavicorne (Linnaeus) 



U.S. NATIONAL MUSEUM 



BULLETIN 243 PLATE 19 




^-W In, L /?0- 

Copium clavicome (Linnaeus), last nymphal instar 



U.S NATIONAL MUSEUM 



BULLETIN 243. PLATE 20 




T ^ 

Floral cecidia of Copium clavicorne, one side removed to expose inmate 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 21 




hi . Li 5-A 



Copium teucrii (Host) 



U.S. NATIONAL MUSEUM 



. • ■- • - . \ ; .- 



^ : 



%- 



t 



*;^ 



v'N 



BULLETIN 243, PLATE 




-<$V 



* 




ki. / r> a- 



Copium teucrii (Host), last nymphal instar 
Floral cecidia of Copium teucrii, one side removed to expose inmate 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 23 







i 



- m 




Paracopium hamadryas (Drake) 



US NATIONAL MUSEUM 



BULLETIN 243. PLATE 24 




In. Li 



Ku 



Floral cecidium of Paracopium hamadryas, one side removed to show the seven inmates 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 25 



S 




■ 

! 

v I 
k 




\cl 



/ ,.[J 



ETpS 



Perissonemia ecmeles Drake and Mohanasundarum 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 26 





B 



Dicrotingis digitalis (Drake) 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 27 




XCC-r**' 



5 



H. 



Codotingis evanis Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 28 




*,. 







Diplocysta trilobata Drake and Poor 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 29 






V;:\| ; V lw\ 









r ,( dim 



hen 



f 

f 






t 












Hesesidemus otiosus Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 30 




Diconocoris capusi (Horvath) 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 31 




Diconocoris capusi (Horvath), last nymphal instar 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 32 




mm 



x 



i \ r f ! 






r r 



■r f l 



( 



7 

-r ^' 



i \f 



( r r 



( 

r 






/" ^ — 




- / J- ; t r r I 




n 



H. 



Australotingis franzeni Hacker 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 33 








WELLS 











ira^fe 



A c 



-"?> 



?0r 5- H - 



Australotingis franzeni Hacker, last nymphal instai 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 34 




PjH 



Dictyla picturata (Distant) 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 35 



' ) 



y 






/ . 






,^/i 












. 



!EpS 



t Dictyla veterna (Scudder) (fossil in stone) 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 









s, 



j 



/. 



' V 






■--',1 '-'S I, ' 



< 



% 



-S~z£ 



m 




Ammianus alberti (Schouteden) 



U.S. NATIONAL MUSEUM 



ULLETIN 243. PLATE 37 




Ammianui alberti (Schouteden), last nymphal instar 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 33 




Ammianus dilatatus (Guerin-Meneville) 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 39 




U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 40 








* f f 



I 



{ 






L/ 



- 



< / ' 

r 



,-•- St 






* C SS( 









/ r 



r 



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/ 



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7 /- 



( 



rfirf) r<r ( 

r ( W$rfc n<X) r 

' r - - r \ ■ ■' > J / ( r A ) 









Planibyrsa montei Drake and Hambleton 



Ij.H. 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 41 




Nobarnit i rignatus (Distant) 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 42 




Inoma multispinosa Hacker 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 43 







Hyalochiton colpochilus (Horvath) 



U.S NATIONAL MUSEUM 



BULLETIN 243, PLATE 44 




/ 



jjf—fjl^^ 




r 



^\ 




KBV- 



Galeatus spinifrons (Fallen) 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 45 




Galeatus sinuatus (Herrich-Schaeffer), brachypterous 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 46 




Galeatus maculatus (Herrich-Schaeffer), macropterous 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 47 




/ 



J 



HP EpS 



Stephanilis typica (Distant) 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 48 




lu. L/s-c*/ 



Stephanitis takeyai Drake and Maa 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 49 












mwM 



r ,r 



g.H. 



Holophygdon melanesica Kirkaldy 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 50 




Gargaphia sororia Hussey 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 51 




Chorotingis indigena Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 52 




Ypsotingis luzonana Drake 



U.S. NATIONAL MUSEUM 



BULLETIN 243. PLATE 53 






Thaumamannia manni Drake and Davis 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 54 




■P^ 





EHF 



Anommatocoris minutissimus Chiru 



U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 55 




U.S. NATIONAL MUSEUM 



BULLETIN 243, PLATE 56 




a, b, Anommatocoris zeteki Drake and Froeschner, and c, A. minutissimus China 



Species Wrongly Classified as Tingidae 

A number of species belonging to other families of Hemiptera have been 
wrongly described as Tingidae, or later mistakenly included as such in the 
literature. For convenience in finding and correcting these errors, the 
technical names of such species are entered below in alphabetical sequence. 
The correct family status is given in the second column opposite the trivial 
name. The asterisk (*) indicates the name of the author who corrected 
the error in the literature. The generic and specific names are also included 
in the index. 

Agrammodes costatus Uhler. — Van Duzee 191 6, p. 26; 1917b, p. Piesmatidae 

223.— Banks 1910, p. 55. 

Copium guttatum Thunberg 1825, p. 10. — *Stal 1870b, p. 157. Coreidae 

Copium maculatum Thunberg 1825, p. 8. — *Stal 1870b, p. 157. Coreidae 

Copium serratum Thunberg 1825, p. g. — Fallen 1829, p. 153. — Coreidae 

Burmeister 1835, p. 330. 

Cymus basicornis Motschulsky. — Distant 1903b, p. 145. — *Bergroth Lygaeidae 

1921, P- 99- 
Lasiocera Laporte 1833, p. 50. — *Fieber 1844, p. 21. Reduviidae 

Maynea maynei Schouteden. — Mayne and Ghesquiere 1934, p. 20. Aradidae 
Metatropis aurita Breddin. — Zool. Rec. 1907, p. 380. Berytidae 

Opisthochasis albocostata Berg 1884, p. 100. — Pennington 1921, p. Miridae 

20. — *Drake and Poor 1938a, p. 103, fig. 1. 
Phyllogasterotinginae Menon and Hakk 1959a, p. 392 [nom. Coreidae 

nud.]. — Menon, Beri and Singh 1959, p. 286. 
Phyllogasterotingis acheranthi Menon and Hakk 1959a, p. 392 [gen., Coreidae 

sp., nom. nud.]. — Menon, Beri and Singh 1959, p. 286. 
Phyllotingis arida Walker 1873b, p. 3. — Lethierry and Severin Aradidae 

1896, p. 5. — *Champion 1898b, p. 5. 
\Tingiopsis reticulata Bekker-Migdisova 1953, p. 461, fig. [fossil]. — Cercopidae 

*Evans 1957, p. 289, fig. 6e. 
Tingis alata (Fabricius). — Fabricius 1803, p. 125. Aradidae 

Tingis virescens (Fabricius) . — Fabricius 1803, p. 127. — Fieber 1844, Lygaeidae 

p. 109. 

445 



681-552—64- 



-30 



Types and Their Locations 

An endeavor has been made to record, whenever possible, pertinent 
data relative to the type (holotype or lectotype) of all species. These 
data include locality (as published in original description without change 
in geographic name), form (pterygopolymorphic), sex (male or female), 
and present deposition of the type specimen. When such information 
was not included in the original description or subsequent publications, 
the data on locality, wing development, and sex were obtained insofar as 
possible from the type specimens and labels thereon. When the type 
specimen is apparently "lost" or no information available on sex, or type 
locality, the term "unknown" is used. 

As heretofore recorded in the pages, tingid types are lodged in museums, 
other institutions, and private collections in many countries. The full name 
and address of the museum or institution (including private collections) 
to which the abbreviations refer are as follows: 



Amer. Mus. 

Argentina Mus. 

Austr. Mus. 
Berlin Mus. 

Bishop Mus. 
British Mus. 

Brussels Mus. 

Bucharest Mus. 
Cal. Acad. 

Canada. Coll. 

Carnegie Mus. 
Cent. Afr. Mus. 

Copenhagen Mus. 

Cornell Univ. 
Coryndon Mus. 
Drake Coll.(USNM) 

446 



American Museum Natural History, New York 

City, New York 
Museo Argentina de Ciencias Naturales, Buenos 

Aires, Argentina 
Australian Museum, Sydney, Australia 
Humboldt-Universitat zu Berlin Zoologisches 

Museum, Berlin, Germany 
Bernice P. Bishop Museum, Honolulu, Hawaii 
British Museum (Natural History), London, 

England 
Institut Royal des Sciences naturelles de Bel- 

gique, Brussels, Belgium 
Muzel de Istorie Naturala, Bucharest, Rumania 
California Academy of Sciences, San Francisco, 

California 
National Collection, Entomological Branch, 

Ottawa, Canada 
Carnegie Museum, Pittsburgh, Pennsylvania 
Musee Royal de l'Afrique Centrale, Tervuren, 

Belgium (formerly Musee du Congo Beige) 
Universitets Zoologiske Museum, Copenhagen, 

Denmark 
Cornell University, Ithaca, New York 
Coryndon Museum, Nairobi, Kenya 
Carl J. Drake Collection (United States Na- 
tional Museum) 



LACE BUGS OF THE WORLD 



447 



Ent. Agr. Brazil 

Florida Univ. 
Frankfort Mus. 

Genova Mus. 

Hamburg Mus. 

Hawaii. Sugar Plant. Assn. 

Helsin. Mus. 
Heude Mus. 
Horn Coll. 

Hungarian Mus. 
Indian Mus. 
Inst. Ent. Madrid 

Inst. Fr. Afr. N. 

Keil Coll. 
Kyushu Univ. 
La Plata Mus. 

Leiden Mus. 

Leningrad Inst. 

Lingnan Surv. 

Lund Zool. Inst. 

Lyon Mus. 

Madagascar Sci. Inst. 

Mancini Coll. 
MCZ 

Michigan Univ. 

Monte Coll. (Mus. Nacional) 

Moscow Univ. 
Munich Mus. 



Servico de Entomologia Agricola do Instituto 

Biologico de Defensa Agricola, Rio de 

Janeiro, Brazil 
University of Florida, Gainesville, Florida 
Senckenbergische Naturforschende Gesellschaft, 

Frankfort a.M., Germany 
Museo Civico di Storia Naturale, Genova, 

Italy 
Zoologische Staatsinstitut und Zoologische 

Museum, Hamburg, Germany 
Hawaiian Sugar Planters Association, Hono- 
lulu, Hawaii 
Kansaiimuseo, Helsinki, Finland 
Museum Heude, Shanghai, China 
Deutsches Entomologisches Institut, Berlin, 

Germany 
Magyar Nemzeti Museum, Budapest, Hungary 
Zoological Survey of India, Calcutta, India 
Instituto Espanol de Entomologica, Madrid, 

Spain 
Institut Francais d'Afrique Noire, Dakar, 

Senegal 
In Copenhagen Museum 

Kyushu Imperial University, Fukuoka, Japan 
Instituto del Museo de la Universidad Nacional 

de La Plata, La Plata, Argentina 
Rijksmuseum van Natuurlijke Historie, Leiden, 

Netherlands 
Zoological Institute of Academy of Sciences, 

Leningrad, U.S.S.R. 
Lingnan Natural History Survey, Canton, 

China 
Zoological Institute, Lund University, Lund, 

Sweden 
Museum des Sciences Naturalles des Lyon, 

Rhone, France 
Institut de Recherche Scientifique de Mada- 
gascar, Tenanarive, Malagasy Republic 
C. Mancini Collection in Genova Museum 
Museum of Comparative Zoology, Harvard 

University, Cambridge, Massachusetts 
Museum of Zoology, University of Michigan, 

Ann Arbor, Michigan 
O. Monte Collection (Museum Nacional, Rio 

de Janeiro, Brazil) 
University of Moscow, Moscow, U.S.S.R. 
Zoologische Sammlung des Bayerischen Staates, 

Munchen, West Germany 



448 



U.S. NATIONAL MUSEUM BULLETIN 2 43 



Naples Mus. 

Ohio State Univ. 
Oxford Mus. 
Pares Nat. Inst. 

Paris Mus. 

Prague Mus. 
Queensland Mus. 
Roubal Coll. 

Seidenstiicker Coll. 

Snow Mus. 
So. Afr. Mus. 

So. Austr. Mus. 
Stockholm Mus. 
USNM 

Vidal Coll. 
Vienna Mus. 
Wagner Colh 



Musee Zoologique l'Universite de Naples, 

Naples, Italy 
Ohio State University, Columbus, Ohio 
Oxford Museum, Oxford, England 
Institut des Pares Nationaux du Congo Beige, 

Brussels, Belgium 
Museum National d'Histoire Naturelle, Paris, 

France 
Narodni Museum, Praha, Czechoslovakia 
Queensland Museum, Brisbane, Australia 
Jean Roubal, U.2, baterie 6, Praha, Czecho- 
slovakia 
Gustav Seidenstiicker, Eichstatt, Bayern Romer 

Strasse 21, AOK, Germany 
University of Kansas, Lawrence, Kansas 
South African Museum, Capetown, South 

Africa 
South Australian Museum, Adelaide, Australia 
Naturhistoriska Rijsmuseet, Stockholm, Sweden 
United States National Museum, Washington, 

D.C. 
J. P. Vidal Collection in Paris Museum 
Naturhistorisches Museum, Wien, Austria 
Edward Wagner, Mooreyhe 103, Hamburg, 

Germany 



Host Plants 



Most species of lacebugs are restricted to a single species of host plants 
or to a relatively small number of closely related plants. A few typical 
exceptions include the cotton or castor bean lacebug, Corythucha gossypii 
(Fabricius), the chrysanthemum lacebug, Corythucha marmorata (Uhler), 
the banana or coconut lacebug, Stephanitis typica (Distant), and the almond 
lacebug, Monosteira unicostata (Mulsant and Rey). 

Since the technical and common names of host plants have been ex- 
tracted directly from the literature, the determination of host plants as 
well as that of the tingid species concerned should be accredited to the 
authority as cited in the text. Except for corrections in spelling, the 
scientific names of plants are listed herein as printed in the literature, with- 
out nomenclatorial modernization or changes. The technical names of 
host plants are arranged alphabetically below the respective family names, 
which too are placed in alphabetical order. 



ACANTHACEAE 

Acanthus sp. 

Naochila kivuensis (Schouteden) 
Anisacanthus thurberi 

Teleonemia huachucae Drake 
Barleria cristata 

Habrochila darthula (Kirkaldy) 
Barleria strigosa 

Habrochila darthula (Kirkaldy) 
Bathysa stipulata 

Gargaphia patria (Drake and Hamble- 
ton) 
Beloperone californica 

Teleonemia schwarzi Drake 
Strobilanthes sp. 

Hegesidemus eliyanus Distant 

ACERACEAE 

Acer sp. 

Physatocheila harwoodi China 
Acer creticum 

Physatocheila dumetorum (Herrich- 
Schaeffer) 
Acer obtusifolium 

Physatocheila dumetorum (Herrich- 
Schaeffer) 
Acer pennsylvanicum 

Corythucha pallipes Parshley 



Acer platanoides 

Physatocheila harwoodi China 
Acer pseudoplatanus 

Physatocheila harwoodi China 
Acer saccharinum 

Corythucha pallipes Parshley 
Acer saccharum 

Corythucha pallipes Parshley 
Acer spicatum 
Corythucha pallipes Parshley 
Aesculaceae 
Aesculus glabra 

Corythucha aesculi Osborn and Drake 
Aesculus hippocastanum 

Corythucha aesculi Osborn and Drake 
Aesculus octandra 
Corythucha aesculi Osborn and Drake 
Agaricaceae 
Pholiota mutabilis 
Acalypta parvula (Fallen) 
Amarantaceae 
Achyranthes aspera 

Corythaica carinata Uhler 
Anacardiaceae 
Pistacia sp. 
Elasmotropis testacea (Herrich-Schaef- 
fer) 

449 



450 



U.S. NATIONAL MUSEUM BULLETIN 243 



Elasmotropis testacea var. vicina Hor- 
vath 

Tingis (Lasiotropis) hellenica (Puton) 
Pseudospondias microcarpa 

Cysteochila ghesquierei Schouteden 
Rhus laurina 

Teleonemia nigrina Champion 
Rhus toxicodendron 

Corythucha padi Drake 

Annonaceae 

Gargaphia concursa Drake 

Leptopharsa mira Drake and Hamble- 
ton 
Annona cherimola 

Teleonemia morio (Stal) 
Annona chrysophylla 

Haedus pallens Schouteden 
Annona coriacea 

Idiostyla anonae (Drake and Hamble- 
ton) 
Annona diversifolia 

Corythucha gossypii (Fabricius) 
Annona muricata 

Corythucha gossypii (Fabricius) 

Haedus pallens Schouteden 

Stephanitis typica (Distant) 
Annona reticulata 

Teleonemia morio (Stal) 
Annona squamosa 

Corythucha gossypii (Fabricius) 

Teleonemia morio (Stal) 
Rollinia silvatica 

Idiostyla rolliniae (Drake and Hamble- 
ton) 
Uvaria rufa 

Stephanitis {Norba) nitor Drake and Poor 
Apocynaceae 

Paseala arnoldi Schouteden 
Aspidosperma sp. 

Liotingis affinata Drake and Hamble- 
ton 

Liotingis aspidospermae Drake and 
Hambleton 

Liotingis evidens Drake 

Liotingis immaculata (Drake and Ham- 
bleton) 



Aspidosperma melanocalyx 
^elotingis aspidospermae (Drake and 
Hambleton) 
Aspidosperma warmingii 

Leptopharsa sobrina Monte 
Condylocarpon rauwolfiae 
Leptopharsa valida Drake and Hamble- 
ton 
Forsteronia sp. 
Leptopharsa forsteroniae Drake and 
Hambleton 
Nerium oleander 
Monosteira unicostata var. buccata Hor- 
vath 

Aquifoliaceae 
Ilex sp. 

Leptoypha elliptica McAtee 
Leptoypha ilicis Drake 
Araceae 
Colocasia esculenta 

Stephanitis colocasiae Horvath 
Raphidophora merrillii 
Cottothucha oceanae Drake and Poor 
Holophygdon melanesica var. fusca Drake 
and Poor 

Araliaceae 
Hedera helix 

Derephysia foliacea (Fallen) 

Aristolochiaceae 
Aristolochia sp. 
Gargaphia flexuosa (Stal) 

Berberidaceae 
Podophyllum peltatum 
Corythucha pallida Osborn and Drake 

Betulaceae 

Alnus sp. 

Corythucha heidemanni Drake 
Corythucha pergandei Heidemann 
Physatocheila costata (Fabricius) 

Alnus acuminata 
Atheas nigricornis Champion 
Corythucha decepta Drake 

Alnus crispa 
Corythucha heidemanni Drake 



LACEBUGS OF THE WORLD 



451 



Alnus glutinosa 

Corythucha pergandei Heidemann 

Monosteira unicostata (Mulsant and 
Rey) 

Physatocheila costata (Fabricius) 
Alnus incana 

Corythucha bellula Gibson 

Corythucha pergandei Heidemann 

Physatocheila costata (Fabricius) 
Alnus montana 

Physatocheila costata (Fabricius) 
Alnus nepalensis 

Stephanitis gallarum Horvath 
Alnus rugosa 

Corythucha pergandei Heidemann 
Betula sp. 

Physatocheila costata (Fabricius) 
Betula alba 

Corythucha pallipes Parshley 
Betula lenta 

Corythucha pallipes Parshley 

Corythucha pergandei Heidemann 
Betula lutea 

Corythucha heidemanni Drake 

Corythucha pallipes Parshley 

Corythucha pergandei Heidemann 
Betula nigra 

Corythucha pergandei Heidemann 
Betula papyrifera 

Corythucha pallipes Parshley 
Betula populifolia 

Corythucha pergandei Heidemann 
Carpinus caroliniana 

Corythucha lowyri Drake 
Corylus sp. 

Corythucha bellula Gibson 

Corythucha coryli Osborn and Drake 
Corylus americana 

Corythucha coryli Osborn and Drake 

Corythucha hewitti Drake 

Corythucha pergandei Heidemann 
Corylus californica 

Corythucha hewitti Drake 
Corylus rostrata 

Corythucha scitula Drake 
Ostrya carpinifolia 

Physatocheila confinis Horvath 



Ostrya virginiana 
Corythucha coryli Osborn and Drake 
Corythucha pallipes Parshley 

BlGNONIACEAE 

Acanthocheila hollandi Drake 

Amblystira silvicola Drake 

Dicysta arnica Drake and Hambleton 

Dicysta sagillata Drake 

Planibyrsa elegantula (Drake) 

Sphaerocysta angulata Monte 

Sphaerocysta globifera (Stal) 

Sphaerocysta inflata (Stal) 

Tingis colombiana Drake 
Adenocalymma sp. 

Dicysta lauta Drake and Hambleton 

Dicysta smithi Drake 

Dicysta vitrea Champion 
Adenocalymma bracteatum 

Phymacysta tumida (Champion) 
Anemopaegma prostratum 

Acanthocheila tumida Drake 
Arrabidaea sp. 

Planibyrsa sodalis (Drake and Bondar) 
Bignonia exoleta 

Acanthocheila visenda Drake and Ham- 
bleton 
Chilopsis sp. 

Teleonemia barberi Drake 
Jacaranda paucifoliata 

Teleonemia validicornis Stal 
Mansoa glaziovii 

Dicysta vitrea Champion 
Petastoma formosum 

Dicysta vitrea Champion 
Petastoma samydoides 

Dicysta vitrea Champion 
Pithecoctenium echinatum 

Teleonemia leitei Drake and Hamble- 
ton 
Tecoma sp. 

Tingis americana Drake 

Tingis tecomae Monte 

BOMBACACEAE 

Bombax sp. 

Tigava bombacis Drake and Poor 
Tigava ferruginea Monte 



452 



U.S. NATIONAL MUSEUM BULLETIN 24 3 



Bombax munguba 

Eocader vegrandis Drake and Hamble- 
ton 
Ceiba pentandra 

Tigava ceibae Drake and Poor 
Chorisia sp. 

Tigava bombacis Drake and Poor 
Chorisia speciosa 

Gargaphia lunulata (Mayr) 

Tigava bombacis Drake and Poor 

BORAGINACEAE 

Acysta praeclara Drake and Hambleton 
Anchusa arvensis 

Dictyla echii (Schrank) 
Anchusa officinalis 

Dictyla echii (Schrank) 

Octacysta echii (Fabricius) 
Arnebia cornuta 

Physatocheila costata (Fabricius) 
Cochranea anchusaefolia 

Dictyla parmata (Distant) 
Cordia sp. 

Compseuta cordiae Drake 

Dictyla cheriana (Drake) 

Dictyla figurata (Drake) 

Dictyla loricata (Distant) 

Dictyla monotropida (Stal) 

Dictyla parmata (Distant) 

Naochila arete Drake and Mohana- 
sundarum 

Tingis (Lasiotropis) beieri Drake 
Cordia abyssinica 

Compseuta cordiae Drake 

Dictyla abyssinica (Drake) 

Horvathula uniseriata (Horvath) 
Cordia alba 

Dictyla alia Drake and Cobben 
Cordia alliodora 

Dictyla monotropida (Stal) 
Cordia chrysocarpa 

Dictyla leroyi (Schouteden) 

Horvathula uniseriata (Horvath) 
Cordia corymbosa 

Dictyla loricata (Distant) 

Dictyla parmata (Distant) 
Cordia curassavica 

Dictyla parmata (Distant) 



Cordia cylindristachya 

Dictyla parmata (Distant) 
Cordia gerascanthus 

Dictyla monotropida (Stal) 
Cordia millenii 

Horvathula uniseriata (Horvath) 
Cordia myxa 

Compseuta lejroyi Distant 

Dictyla sauteri (Drake) 
Cordia serrata 

Dictyla haitiensis (Drake and Poor) 
Cordia tomentosa 

Dictyla monotropida (Stal) 
Cordia verbenacea 

Dictyla parmata (Distant) 
Cynoglossum sp. 

Dictyla echii (Schrank) 

Dictyla nassata (Puton) 

Dictyla triconula (Seidenstiicker) 

Octacysta echii (Fabricius) 
Cynoglossum creticum 

Dictyla putoni var. pulla (Horvath) 
Cynoglossum officinale 

Dictyla echii (Schrank) 

Dictyla triconula (Seidenstiicker) 
Echinospermum lappula 

Dictyla platyoma (Fieber) 
Echium sp. 

Derephysia foliacea (Fallen) 

Dictyla echii (Schrank) 

Dictyla putoni (Montandon) 

Dictyonota (Alcletha) tricornis (Schrank) 

Octacysta echii (Fabricius) 
Echium altissimum 

Dictyla echii (Schrank) 

Dictyla putoni var. pulla (Horvath) 
Echium candicans 

Dictyla indigena (Wollaston) 
Echium creticum 

Dictyla echii (Schrank) 
Echium hypertropicum 

Dictyla indigena (Wollaston) 
Echium italicum 

Dictyla echii (Schrank) 

Octacysta echii (Fabricius) 
Echium plantagineum 

Dictyla nassata (Puton) 



LACEBUGS OF THE WORLD 



453 



Echium pustulatum 

Dictyla echii (Schrank) 
Echium rubrum 
Dictyla echii (Schrank) 
Octacysta echii (Fabricius) 
Echium sericeum 

Dictyla nassata (Puton) 
Echium stenosiphon 

Dictyla indigena (Wollaston) 
Echium vulgare 

Acalypta gracilis (Fieber) 

Derephysia joliacea (Fallen) 

Dictyla echii (Schrank) 

Dictyonota (Alcletha) tricornis (Schrank) 

Octacysta echii (Fabricius) 
Ehretia acuminata 

Dictyla evidens (Drake) 
Ehretia dicksonii 

Dictyla evidens (Drake) 
Ehretia elliptica 

Dictyla ehrethiae (Gibson) 
Ehretia laevis 

Naochila sufflata (Drake and Poor) 
Ehretia taiwaniana 

Dictyla evidens (Drake) 
Ehretia thyrsiflora 

Dictyla evidens (Drake) 
Heliotropium erosum 

Dictyonota {Alcletha) tricornis (Schrank) 
Lappula sp. 

Leptopharsa clitoriae (Heidemann) 
Lycopsis arvensis 

Dictyla echii (Schrank) 
Myosotis sp. 

Dictyla convergens (Herrich-Schaeffer) 

Dictyla lupuli (Herrich-Schaeffer) 

Dictyla platyoma (Fieber) 
Myosotis cespitosa 

Dictyla convergens (Herrich-Schaeffer) 
Myosotis lingulata 

Dictyla convergens (Herrich-Schaeffer) 
Myosotis palustris 

Dictyla convergens (Herrich-Schaeffer) 

Dictyla lupuli (Herrich-Schaeffer) 

Dictyla platyoma (Fieber) 
Onosma sp. 

Dictyla nassata (Puton) 



Onosma echioides 

Campylosteira verna (Fallen) 
Onosma visianii 

Dictyla putoni var. pulla (Horvath) 
Pulmonaria sp. 

Dictyla echii (Schrank) 
Symphytum sp. 

Dictyla echii (Schrank) 

Dictyla humuli (Fabricius) 

Dictyla platyoma (Fieber) 
Symphytum officinale 

Dictyla humuli (Fabricius) 
Tournefortia sp. 

Dictyla loricata (Distant) 
Tournefortia sibirica 

Dictyla montandoni (Horvath) 

Bromeliaceae 
Aechmea sp. 
Ambycysta gibbifera (Picado) 
Psilobyrsa aechemeae Drake and Ham- 
bleton 
Vriesia hoehneana 
Psilobyrsa vriesiae Drake and Hamble- 
ton 

Capparidaceae 
Atamisquea emarginata 

Corythucha gossypii (Fabricius) 
Capparis cynophallophora 

Corythucha gossypii (Fabricius) 
Capparis flexuosa 
Corythucha gossypii (Fabricius) 

Gaprifoliaceae 
Lonicera sp. 

Derephysia joliacea (Fallen) 
Viburnum tinus 

Stephanitis chlorophana (Fieber) 

Caricageae 
Carica papaya 
Corythucha gossypii (Fabricius) 

Caryophyllaceae 
Gymnocarpos decandrum 

Sabestena alfierii Drake and Ruhoff 
Herniaria sp. 

Acalypta parvula (Fallen) 
Herniaria glabra 

Galeatus maculatus (Herrich-Schaeffer) 



454 



U.S. NATIONAL MUSEUM BULLETIN 243 



Casuarinaceae 
Casuarina sp. 

Epimixia veteris Drake 
Casuarina equisetifolia 

Eocader bouclei (Bruner) 

Chenopodiaceae 
Chenopodium sp. 

Derephysia foliacea (Fallen) 
Halogeton sativus 

Dictyla nassata (Puton) 
Salsola pestifer 

Corythaica venusta (Champion) 

ClSTACEAE 

Cistus albidus 

Urentius chobauti Horvath 
Helianthemum lippii 

Urentius euonymus Distant 

Climaciaceae 
Climacium americanum 

Acalypta lillianis Torre-Bueno 
Climacium dendroides 

Acalypta parvula (Fallen) 

Derephysia foliacea (Fallen) 

Compositae 
Ambrosia sp. 

Gargaphia iridescens Champion 
Ambrosia artemisiifolia 

Corythucha morrilli Osborn and Drake 
Ambrosia tenuifolia 

Gargaphia subpilosa Berg 
Ambrosia trifida 

Corythucha marmorata (Uhler) 
Amphiachyris sp. 

Gargaphia solani Heidemann 
Antennaria sp. 

Leptocysta sexnebulosa (Stal) 
Antennaria dioica 

Galeatus spinifrons (Fallen) 
Artemisia sp. 

Acalypta gracilis (Fieber) 

Derephysia foliacea (Fallen) 

Dictyonota {Alcletha) tricornis (Schrank) 

Galeatus scrophicus Saunders 

Tingis comosa (Takeya) 
Artemisia absinthium 

Tingis crispata (Herrich-Schaeffer) 



Artemisia campestris 

Acalypta gracilis (Fieber) 

Acalypta marginata (Wolff) 

Derephysia cristata (Panzer) 

Derephysia foliacea (Fallen) 

Dictyonota {Alcletha) tricornis (Schrank) 

Galeatus spinifrons (Fallen) 

Tingis crispata (Herrich-Schaeffer) 
Artemisia dracunculoides 

Corythucha morrilli Osborn and Drake 
Artemisia vulgaris 

Derephysia foliacea (Fallen) 

Galeatus spinifrons (Fallen) 

Tingis crispata (Herrich-Schaeffer) 
Artemisia vulgaris v. indica 

Galeatus spinifrons (Fallen) 
Aster sp. 

Corythucha marmorata (Uhler) 
Aster macrophyllus 

Galeatus spinifrons (Fallen) 
Baccharis sp. 

Corythucha baccharides Drake 
Baccharis halimifolia 

Corythucha baccharides Drake 
Baccharis mesoneura 

Leptobyrsa baccharidis Drake and Ham- 
bleton 

Leptobyrsa steini (Stal) 
Baccharis pilularis 

Corythucha morrilli Osborn and Drake 
Baccharis pingraea 

Coleopterodes brunnea Drake and Poor 

Coleopterodes liliputiana (Signoret) 
Balsamorhiza sagittata 

Corythucha immaculata Osborn and 
Drake 
Bidens pilosa v. radiata 

Leptopharsa machalana var. vinnula 
Drake and Hambleton 
Cacalia thunbergii 

Cochlochila lewisi (Scott) 
Carduus sp. 

Corythucha distincta var. spinata Osborn 
and Drake 

Tingis ampliata (Herrich-Schaeffer) 

Tingis cardui (Linnaeus) 

Tingis veteris Drake 

Tingis (Tropidocheila) marrubii Vallot 



LACEBUGS OF THE WORLD 



455 



Carduus acanthoides 

Tingis cardui (Linnaeus) 
Carduus crispus 

Tingis cardui (Linnaeus) 
Carduus lanceolatus 

Corythucha distincta Osborn and Drake 

Stephanitis oberti (Kolenati) 
Tingis cardui (Linnaeus) 
Carduus nutans 

Tingis cardui (Linnaeus) 
Carthamus tinctorius 

Cochlochila bullita (Stal) 
Cassinia leptophylla 

Cyperobia carectorum Bergroth 
Centaurea paniculata 

Tingis grisea Germar 
Centaurea rhenana 

Tingis grisea Germar 
Centaurea solstitialis 

Tingis juentei Horvath 
Chrysanthemum sp. 

Catoplatus fabricii (Stal) 

Corythucha marmorata (Uhler) 

Dictyonota (Alcletha) tricornis (Schrank) 

Galeatus scitulus Drake and Maa 

Galeatus scrophicus Saunders 

Physatocheila costata (Fabricius) 
Chrysanthemum indicum 

Galeatus spinifrons (Fallen) 
Chrysanthemum leucanthemum 

Catoplatus fabricii (Stal) 

Derephysia joliacea (Fallen) 
Chuquiragua glabra 

Phymacysta magnifica (Drake) 
Cirsium sp. 

Tingis ampliata (Herrich-Schaeffer) 

Tingis demised Horvath 

Tingis veteris Drake 

Tingis (Lasiotropis) reticulata Herrich- 
Schaeffer 
Cirsium arvense 

Tingis ampliata (Herrich-Schaeffer) 
Cirsium lanceolatum 

Tingis ampliata (Herrich-Schaeffer) 

Tingis cardui (Linnaeus) 
Cirsium palustre 

Tingis ampliata (Herrich-Schaeffer) 

Tingis cardui (Linnaeus) 



Cirsium vulgare 

Tingis cardui (Linnaeus) 
Cnicus sp. 

Corythucha distincta Osborn and Drake 
Crepis tectorum 

Tingis ( Tropidocheila) pilosa Hummel 
Dahlia sp. 

Corythucha gossypii (Fabricius) 
Dahlia parryi 

Gargaphia condensa Gibson 
Dahlia spinosa 

Gargaphia opacula Uhler 
Echinops sp. 

Elasmotropis testacea (Herrich-Schaef- 
fer) 

Elasmotropis testacea var. platydera Hor- 
vath 
Echinops sphaerocephala 

Corythucha marmorata (Uhler) 

Elasmotropis testacea (Herrich-Schaef- 
fer) 
Echinops spinosa 

Galeatus scrophicus Saunders 

Tingis {Lasiotropis) rotundipennis Hor- 
vath 
Egletes prostrata 

Corythucha morrilli Osborn and Drake 
Erigeron canadensis 

Corythucha morrilli Osborn and Drake 
Eupatorium sp. 

Galeatus spinifrons (Fallen) 
Eupatorium intermedium 

Leptobyrsa baccharidis Drake and Ham- 
bleton 

Leptobyrsa steini (Stal) 
Galactites tomentosa 

Tingis cardui (Linnaeus) 
Helenium tenuifolium 

Teleonemia nigrina Champion 
Helianthus sp. 

Corythucha marmorata (Uhler) 

Corythucha morrilli Osborn and Drake 
Helianthus annuus 

Corythucha morrilli Osborn and Drake 

Gargaphia subpilosa Berg 
Helianthus tuberosus 

Corythucha marmorata (Uhler) 



456 



U.S. NATIONAL MUSEUM BULLETIN 24 3 



Helichrysum angustifolium 

Copium teucrii (Host) 
Helichrysum arenarium 

Galeatus spinifrons (Fallen) 
Helichrysum italicum 

Copium intermedium (Rey) 
Hieracium sp. 

Dictyonota (Alcletha) tricornis (Schrank) 

Galeatus maculatus (Herrich-Schaeffer) 

Galeatus spinifrons (Fallen) 

Lasiacantha capucina (Germar) 
Hieracium murorum 

Acalypta nigrina (Fallen) 
Hieracium pilosella 

Acalypta marginata (Wolff) 

Galeatus maculatus (Herrich-Schaeffer) 

Galeatus sinuatus (Herrich-Schaeffer) 
Hymenoclea salsola 

Teleonemia schwarzi Drake 
Lagascea mollis 

Corythucha morrilli Osborn and Drake 
Lappa minor 

Tingis {Tropidocheila) pilosa Hummel 

Tingis { Tropidocheila) pilosa var. ampli- 
costa (Montandon) 
Launaea spinosa 

Galeatus scrophicus Saunders 
Ligularia sibirica 

Cochlochila lewisi (Scott) 
Mikania sp. 

Leptocysta sexnebulosa (Stal) 
Mikania auricularis 

Gargaphia subpilosa Berg 
Moquinia sp. 

Leptobyrsa tersa Drake and Hambleton 
Moquinia polymorpha 

Leptobyrsa tersa Drake and Hambleton 
Onopordon sp. 

Catoplatus carthusianus (Goeze) 
Petasites japonica 

Cochlochila lewisi (Scott) 
Pluchea indica 

Corythucha morrilli Osborn and Drake 
Rudbeckia serotina 

Corythucha marmorata (Uhler) 
Senecio sp. 

Tingis {Tropidocheila) pilosa Hummel 



Senecio jacobaea 

Dictyonota {Alcletha) tricornis (Schrank) 

Oncochila scapularis (Fieber) 

Tingis {Lasiotropis) reticulata Herrich- 
Schaeffer 
Serratula sp. 

Tingis cardui (Linnaeus) 
Serratula atriplicifolia 

Tingis lasiocera Matsumura 
Silybum sp. 

Tingis cardui (Linnaeus) 
Solidago sp. 

Corythucha marmorata (Uhler) 

Melanorhopala clavata (Stal) 
Solidago sempervirens 

Corythucha marmorata (Uhler) 
Symphyopappus reticulatus 

Leptobyrsa ardua Drake 

Leptobyrsa steini (Stal) 
Synedrella nodiflora 

Corythucha morrilli Osborn and Drake 
Tanacetum sp. 

Corythucha marmorata (Uhler) 
Verbesina sp. 

Teleonemia sacchari (Fabricius) 
Verbesina encelioides 

Corythucha morrilli Osborn and Drake 
Vernonia sp. 

Ammianus alberti subsp. tricarinatus 
(Schouteden) 

Corythucha agalma Drake and Cobben 

Leptocysta sexnebulosa (Stal) 

Nectocader gounellei (Drake) 

Phatnoma annulipes Champion 
Vernonia conferta 

Ammianus alberti (Schouteden) 
Vernonia polyanthes 

Phatnoma vernoniae Drake and Hamble- 
ton 

Teleonemia brevipennis Champion 
Wedelia jacquini 

Corythucha morrilli Osborn and Drake 
Xanthium sp. 

Corythucha morrilli Osborn and Drake 

Teleonemia scrupulosa Stal 
Xanthium canadense 

Corythucha morrilli Osborn and Drake 



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Zinnia elegans 
Gargaphia torresi Costa Lima 

CONVOLVULACEAE 

Ipomoea sp. 

Gargaphia deceptiva (Drake and Bruner) 

Gargaphia torresi Costa Lima 
Ipomoea batatas 

Corythucha morrilli Osborn and Drake 

Leptocysta sexnebulosa (Stal) 

Teleonemia jorticornis Champion 
Ipomoea bonariensis 

Gargaphia subpilosa Berg 
Ipomoea fistulosa 

Teleonemia brevipennis Champion 
Crassulaceae 
Sedum acre 

Dictyonota (Alcletha) tricornis (Schrank) 

CUCURBITACEAE 

Cucurbita moschata 

Teleonemia tricolor (Mayr) 
Lagenaria leucantha 

Corythucha gossypii (Fabricius) 
Sechium edule 

Teleonemia tricolor (Mayr) 
Sicana odorifera 

Teleonemia tricolor (Mayr) 

CUPRESSACEAE 

Cupressus sp. 
Elasmotropis testacea var. vicina Horvath 

Cyperaceae 

Agramma confusum (Puton) 
Carex sp. 

Acalypta platycheila (Fieber) 

Agramma laetum (Fallen) 

Agramma ruficorne (Germar) 

Derephysia foliacea (Fallen) 
Eriophorum sp. 

Agramma tropidopterum Flor 
Eriophorum angustifolium 

Agramma femorale Thomson 
Rhynchospora alba 

Agramma femorale Thomson 

Agramma laetum (Fallen) 
Schoenus albus 

Agramma femorale Thomson 

Agramma laetum (Fallen) 



Scirpus holoschoenus 
Agramma atricapillum (Spinola) 
Agramma dubium var. imbecillum (Hor- 
vath) 
Scleria myricocarpa 
Leptopharsa pensa Drake and Hamble- 
ton 

DlCHAPETALACEAE 

Dichapetalum cymosum 
Onymochila dichapetali (Horvath) 

DlLLENIACEAE 

Davilla rugosa 

Tingis silvacata Drake 

Ericaceae 
Andromedea sp. 

Stephanitis takeyai Drake and Maa 
Azalea see Rhododendron 
Calluna sp. 

Acalypta gracilis (Fieber) 

Galeatus spinifrons (Fallen) 
Calluna vulgaris 

Acalypta marginata (Wolff) 

Acalypta nigrina (Fallen) 

Acalypta parvula (Fallen) 

Galeatus spinifrons (Fallen) 

Stephanitis oberti (Kolenati) 

Tingis (Lasiotropis) reticulata Herrich- 
Schaeffer 
Chamaedaphne sp. 

Corythucha ciliata (Say) 
Erica sp. 

Tingis {Lasiotropis) reticulata Herrich- 
Schaeffer 
Erica vulgaris 

Galeatus maculatus (Herrich-Schaeffer) 

Tingis {Lasiotropis) reticulata Herrich- 
Schaeffer 
Kalmia angustifolia 

Stephanitis rhododendri Horvath 
Kalmia latifolia 

Physatocheila plexa (Say) 

Stephanitis pyrioides (Scott) 

Stephanitis rhododendri Horvath 
Ledum palustre 

Stephanitis oberti (Kolenati) 
Lyonia neziki 

Stephanitis takeyai Drake and Maa 



458 



U.S. NATIONAL MUSEUM BULLETIN 24 3 



Myrtillus niger 

Stephanitis oberti (Kolenati) 
Myrtillus uliginosus 

Stephanitis oberti (Kolenati) 
Pierus floribunda 

Stephanitis rhododendri Horvath 
Pierus japonica 

Stephanitis rhododendri Horvath 

Stephanitis takeyai Drake and Maa 
Pieris ovalifolia 

Stephanitis pyrioides (Scott) 

Stephanitis takeyai Drake and Maa 
Rhododendron sp. 

Stephanitis caucasica Kiritshenko 

Stephanitis oberti (Kolenati) 

Stephanitis propinqua Horvath 

Stephanitis pyrioides (Scott) 

Stephanitis rhododendri Horvath 
Rhododendron america 

Stephanitis rhododendri Horvath 
Rhododendron amoena 

Stephanitis pyrioides (Scott) 
Rhododendron amurasaki 

Stephanitis pyrioides (Scott) 
Rhododendron Antoon van Welie 

Stephanitis rhododendri Horvath 
Rhododendron benigeri 

Stephanitis pyrioides (Scott) 
Rhododendron calenulaceum 

Stephanitis pyrioides (Scott) 
Rhododendron caractacus 

Stephanitis rhododendri Horvath 
Rhododendron Cunningham's White 

Stephanitis rhododendri Horvath 
Rhododendron cynthia 

Stephanitis rhododendri Horvath 
Rhododendron Dr. V. H. Rutgers 

Stephanitis rhododendri Horvath 
Rhododendron everestianum 

Stephanitis rhododendri Horvath 
Rhododendron fastuosum 

Stephanitis rhododendri Horvath 
Rhododendron hatsugeri 

Stephanitis pyrioides (Scott) 
Rhododendron hinodegeri 

Stephanitis pyrioides (Scott) 
Rhododendron hollandia 

Stephanitis rhododendri Horvath 



Rhododendron hymenanthes var. hep- 
tamerum 
Stephanitis hikosana Drake 
Rhododendron indica 

Stephanitis pyrioides (Scott) 
Rhododendron kaempheri 
Stephanitis pyrioides (Scott) 
Rhododendron ledifolium alba 

Stephanitis pyrioides (Scott) 
Rhododendron maximum 

Stephanitis rhododendri Horvath 

Rhododendron Mme. de Bruin 

Stephanitis rhododendri Horvath 

Rhododendron Mme. Carvalho 

Stephanitis rhododendri Horvath 

Rhododendron Mme. Mason 

Stephanitis rhododendri Horvath 
Rhododendron mollis 

Stephanitis pyrioides (Scott) 
Rhododendron Mr. R. S. Holford 

Stephanitis rhododendri Horvath 
Rhododendron mucrunatum 

Stephanitis pyrioides (Scott) 
Rhododendron obtusum amoenum 

Stephanitis pyrioides (Scott) 
Rhododendron parsons gloriosum 

Stephanitis rhododendri Horvath 
Rhododendron Peter Koster 

Stephanitis rhododendri Horvath 
Rhododendron ponticum 

Stephanitis pyrioides (Scott) 
Rhododendron schilippenbachii 

Stephanitis pyrioides (Scott) 
Rhododendron shibori 

Stephanitis pyrioides (Scott) 
Rhododendron shirogeri 

Stephanitis pyrioides (Scott) 
Rhododendron Van der Hoop 
Stephanitis rhododendri Horvath 
Rhododendron yedoense pokhenense 

Stephanitis pyrioides (Scott) 
Rhododendron yodogawa 

Stephanitis pyrioides (Scott) 
Vaccinium sp. 

Leptoypha ilicis Drake 
Vaccinium myrtillus 
Derephysia foliacea (Fallen) 



LACEBUGS OF THE WORLD 



459 



Stephanitis oberti (Kolenati) 

Stephanitis pyri (Fabricius) 
Vaccinium uliginosum 

Stephanitis oberti (Kolenati) 
Vaccinium vitis-idaea 

Stephanitis oberti (Kolenati) 
Xolisma ferruginea 

Leptoypha mcateei Drake 

Erythroxylaceae 
Erythroxylon sp. 

Corycera rugulosa Drake 

Corycera separata Drake and Hamble- 
ton 
Erythroxylon deciduum 

Leptopharsa decens Drake and Hamble- 
ton 

EUPHORBIACEAE 

Corythucha socia Monte 

Gargaphia patricia (Stal) 
Acalypha sp. 

Gargaphia penningtoni Drake 
Cnidoscolus sp. 

Gargaphia lunulata (Mayr) 
Croton sp. 

Gargaphia concursa Drake 

Gargaphia costalimai Monte 

Gargaphia torresi Costa Lima 

Haedus decellei Schouteden 

Tingis (Lasiotropis) beieri Drake 
Croton antisyphiliticus 

Corycera spissa Drake 
Croton flavens 

Leptopharsa ruris Drake 
Croton lobatus 

Gargaphia penningtoni Drake 
Croton pungens 

Gargaphia mirabilis Monte 
Croton urucurana 

Gargaphia crotonae Drake and Hamble- 
ton 

Gargaphia decoris Drake 
Dalechampia ficifolia 

Gargaphia differitas Drake 
Euphorbia cyparissias 

Oncochila simplex (Herrich-Schaeffer) 
Euphorbia gerardiana 

Oncochila scapularis (Fieber) 

Oncochila simplex (Herrich-Schaeffer) 



Euphorbia heterophylla 

Gargaphia lunulata (Mayr) 
Glochidion sp. 

Phatnoma veridica Drake and Maa 
Hevea sp. 

Cysteochila biseriata Schouteden 

Cysteochila lueboensis Schouteden 
Hevea brasiliensis 

Leptopharsa heveae Drake and Poor 
Jatropha sp. 

Corythucha gossypii (Fabricius) 
Jatropha gossypifolia 

Corythucha gossypii (Fabricius) 
Jatropha multifida 

Corythucha gossypii (Fabricius) 
Jatropha urens 

Gargaphia niginervis Stal 
Mallotus philippinensis 

Trachypeplus Jacob soni Horvath 

Trachypeplus malloti Drake and Poor 
Manihot sp. 

Atheas cearanus Monte 

Vatiga illudens (Drake) 

Vatiga manihotae (Drake) 
Manihot dulcis 

Vatiga illudens (Drake) 

Vatiga illudens var. varianta (Drake) 
Manihot esculenta 

Vatiga illudens (Drake) 
Manihot utilissima 

Gargaphia lunulata (Mayr) 

Vatiga illudens (Drake) 

Vatiga longula (Drake) 

Vatiga manihotae (Drake) 
Ricinus communis 

Corythaica cyathicollis (Costa) 

Corythucha gossypii (Fabricius) 

Corythucha spinosa (Duges) 

Gargaphia lunulata (Mayr) 
Fagaceae 
Castanea americana 

Corythucha arcuata (Say) 
Castanea sativa 

Stephanitis pyri (Fabricius) 
Fagus grandifolia 

Corythucha pallipes Parshley 



460 



U.S. NATIONAL MUSEUM BULLETIN 243 



Quercus sp. 

Acalypta hellenica Reuter 

Corthuchay arcuata (Say) 

Corythucha cydoniae (Fitch) 

Derephysia foliacea (Fallen) 

Physatocheila costata (Fabricius) 
Quercus acuminata 

Corythucha arcuata (Say) 
Quercus acutissima 

Uhlerites debilis (Uhler) 
Quercus alba 

Corythucha arcuata (Say) 
Quercus coccifera 

Monosteira lobulifera Reuter 

Physatocheila confinis Horvath 

Stephanitis pyri (Fabricius) 
Quercus dentata 

Uhlerites debilis (Uhler) 
Quercus dilatata 

Physatocheila dryadis Drake and Poor 
Quercus glandulifera 

Uhlerites debilis (Uhler) 
Quercus ilex 

Monosteira lobulifera Reuter 

Physatocheila confinis Horvath 

Stephanitis pyri (Fabricius) 
Quercus macrocarpa 

Corythucha arcuata (Say) 
Quercus muehlenbergii 

Corythucha arcuata (Say) 
Quercus prinoides 

Corythucha arcuata (Say) 
Quercus prinus 

Corythucha arcuata (Say) 
Quercus pubescens 

Physatocheila confinis Horvath 
Quercus rubra 

Corythucha arcuata (Say) 
Quercus serrata 

Uhlerites debilis (Uhler) 
Quercus suber 

Catoplatus mamorensis Bergevin 
Flacourtiaceae 
Olmediella betschleriana 

Leptopharsa dapsilis Drake and Ham- 
bleton 



Geraniaceae 
Erodium sp. 

Acalypta marginata (Wolff) 

Acalypta nigrina (Fallen) 

Acalypta parvula (Fallen) 
Erodium circutarium 

Acalypta gracilis (Fieber) 
Gnetaceae 
Ephedra sp. 

Tingis lanigera (Puton) 

GOODENIACEAE 

Scaevola koenigii 

Paracopium summervillei (Hacker) 
Gramineae 

Agramma atricapillum var. pallens (Hor- 
vath) 

Arusha horvathi Drake 

Leptodictya (Hanuala) plana Heidemann 

Leptopharsa lineata (Champion) 

Tigava graminis Drake and Poor 
Andropogon sp. 

Hesperotingis antennata Parshley 
Bambusa sp. 

Corythucha spinosa (Duges) 

Leptodictya {Hanuala) approximata (Stal) 

Leptodictya {Hanuala) venezolana Monte 

Nyctotingis obsorni Drake 
Bambusa vulgaris 

Leptodictya {Hanuala) bambusae Drake 
Cenchrus glaucus 

Bako malayanus (Drake) 
Cynodon dactylon 

Bako malayanus (Drake) 
Dendrocalamus strictus 

Leptodictya {Hanuala) bambusae Drake 
Ichnanthus leiocarpus 

Aepycysta undosa Drake and Bondar 
Merostachys sp. 

Stephanitis mitrata (Stal) 
Olyra sp. 

Leptodictya {Hanuala) austrina Drake 
and Hambleton 

Leptodictya {Hanuala) comes Drake and 
Hambleton 

Leptodictya {Hanuala) madra Drake and 
Hambleton 

Leptodictya {Hanuala) nota Drake and 
Hambleton 



LACEBUGS OF THE WORLD 



461 



Leptodictya (Hanuala) olyrae Drake 

Leptodictya {Hanuala) solita Drake and 
Hambleton 
Olyra latifolia 

Leptodictya {Hanuala) olyrae Drake 
Olyra micrantha 

Stephanitis olyrae Drake and Hamble- 
ton 
Oryza sativa 

Bako malayanus (Drake) 
Panicum lindheimeri 

Corythaica bellula Torre-Bueno 
Panicum maximum 

Leptodictya {Hanuala) dola Drake and 
Hambleton 
Panicum repens 

Bako malayanus (Drake) 
Panicum turgidum 

Phaenotropis cleopatra (Horvath) 
Paspalum sp. 

Leptodictya {Hanuala) paspalii Drake 
and Hambleton 
Phyllostachys castilloni 

Leptodictya {Hanuala) litigiosa Monte 
Psamma sp. 

Tingis {Lasiotropis) reticulata Herrich- 
Schaeffer 
Saccharum officinarum 

Agramma jormosanum (Matsumura) 

Leptodictya {Hanuala) bambusae Drake 
Urochloa reptans 

Aconchus urbanus (Horvath) 
Zea mays 

Corythucha salicata Gibson 
Hamamelidaceae 
Hamamelis sp. 

Leptoypha costata Parshley 
Hernandiaceae 
Hernandia sp. 

Stephanitis subjasciata Horvath 

Hydrophyllaceae 
Eriodictyon californicum 
Corythucha eriodictyonae Osborn and 
Drake 

Hypericaceae 
Hypericum sp. 
Catoplatus fabricii (Stal) 
681-552—64 31 



JUGLANDACEAE 

Carya sp. 

Corythucha caryae Bailey 
Carya illinoensis 

Corythucha juglandis (Fitch) 
Carya olivaeformis 

Corythucha juglandis (Fitch) 
Carya ovata 

Corythucha caryae Bailey 

Corythucha ciliata (Say) 
Hicoria alba 

Leptoypha costata Parshley 
Juglans cinerea 

Corythucha juglandis (Fitch) 
Juglans nigra 

Corythucha juglandis (Fitch) 
Juglans regia 

Stephanitis pyri (Fabricius) 
Juglans sieboldiana 

Corythucha juglandis (Fitch) 

JUNCACEAE 

Juncus sp. 

Agramma atricapillum (Spinola) 

Agramma carinatum (Distant) 

Agramma confusum (Puton) 

Agramma jallax (Horvath) 

Agramma laetum (Fallen) 

Agramma melanoscele (Horvath) 

Agramma rujicorne (Germar) 

Agramma tropidopterum Flor 

Derephysia joliacea (Fallen) 
Juncus acutus 

Agramma atricapillum (Spinola) 
Juncus gerardi 

Agramma confusum (Puton) 
Juncus maritimus 

Agramma atricapillum (Spinola) 

Agramma laetum (Fallen) 
Luzula campestris 

Agramma laetum (Fallen) 
Labiatae 

Lasiacantha altimitrata (Takeya) 
Ajuga sp. 

Lasiacantha capucina (Germar) 

Tingis {Lasiotropis) reticulata Herrich- 
Schaeffer 

Tingis { Tropidocheila) caucasica ( Jakov- 
lev) 



462 



U.S. NATIONAL MUSEUM BULLETIN 243 



Ajuga chamaepytis 

Tingis (Lasiotropis) ajugarum (Frey- 
Gessner) 
Ajuga genevensis 

Tingis (Lasiotropis) ajugarum (Frey- 
Gessner) 

Tingis (Lasiotropis) reticulata Herrich- 
Schaeffer 
Ajuga iva 

Tingis (Lasiotropis) ajugarum (Frey- 
Gessner) 
Ajuga reptans 

Tingis (Lasiotropis) reticulata Herrich- 
Schaeffer 
Ballota nigra 

Tingis (Tropidocheila) pilosa Hummel 

Tingis (Tropidocheila) pilosa var. am- 
plicosta (Montandon) 
Betonica officinalis 

Hyalochiton colpochilus (Horvath) 
Coleus sp. 

Cochlochila bullita (Stal) 
Galeopsis tetrahit 

Tingis (Tropidocheila) pilosa Hummel 

Tingis (Tropidocheila) pilosa var. am- 
plicosta (Montandon) 
Hoslundia sp. 

Dictyla sjostedti (Horvath) 
Hoslundia oppositifolia 

Naochila boxiana (Drake) 
Keiskea japonica 

Copium japonicum Esaki 
Leonurus cardiaca 

Tingis (Tropidocheila) pilosa Hummel 

Tingis (Tropidocheila) pilosa var. am- 
plicosta (Montandon) 
Marrubium sp. 

Tingis (Tropidocheila) marrubii Vallot 
Marrubium vulgare 

Tingis cardui (Linnaeus) 

Tingis (Tropidocheila) marrubii Vallot 
Mentha sp. 

Cochlochila bullita (Stal) 

Dictyonota (Alcletha) tricornis (Schrank) 
Mosla punctata 

Copium japonicum Esaki 
Ocium basilicum 

Cochlochila bullita (Stal) 



Ocium canum 

Cochlochila bullita (Stal) 
Ocium kilimandscharicum 

Cochlochila bullita (Stal) 
Ocium sanctum 

Cochlochila bullita (Stal) 
Phlomis sp. 

Hyalochiton multiseriatus (Reuter) 
Phlomis fruticosa 

Tingis (Lasiotropis) hellenica (Puton) 
Phlomis herba-venti 

Tingis (Tropidocheila) granadensis Hor- 
vath 
Phlomis lynchitis 

Hyalochiton colpochilus (Horvath) 

Tingis (Lasiotropis) trichonota (Puton) 
Phlomis pungens 

Tingis ( Tropidocheila) pauper ata (Puton) 
Phlomis tuberosa 

Galeatus sinuatis (Herrich-Schaeffer) 
Prunella vulgaris 

Teleonemia prunellae Drake and Ham- 
bleton 
Salvia azurea 

Gargaphia solani Heidemann 
Salvia officinalis 

Cochlochila bullita (Stal) 
Salvia pitcheri 

Gargaphia solani Heidemann 
Salvia verticillata 

Tingis (Lasiotropis) reticulata Herrich- 
Schaeffer 
Satureja montana 

Lasiacantha gracilis (Herrich-Schaef- 
fer) 
Sideritis sp. 

Tingis ( Tropidocheila) stachydis (Fieber) 
Stachys sp. 

Tingis auriculata (Costa) 

Tingis (Lasiotropis) ragusana (Fieber) 

Tingis ( Tropidocheila) stachydis (Fieber) 
Stachys annua 

Tingis ( Tropidocheila) stachydis (Fieber) 
Stachys italica 

Tingis (Lasiotropis) ragusana (Fieber) 
Stachys recta 

Tingis auriculata (Costa) 

Tingis ( Tropidocheila) stachydis (Fieber) 



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463 



Stachys salviaefolia 

Tingis {Lasiotropis) raguasana (Fieber) 
Stachys sylvatica 

Tingis {Tropidocheila) pilosa Hummel 
Tingis {Tropidocheila) pilosa var. am- 
plicosta (Montandon) 
Teucrium sp. 
Copium clavicorne (Linnaeus) 
Copium horvathi Wagner 
Copium teucrii (Host) 
Hyalochiton komaroffii (Jakovlev) 
Teucrium canum 

Copium clavicorne (Linnaeus) 
Teucrium capitatum 
Copium teucrii (Host) 
Teucrium chamaedrys 
Copium clavicorne (Linnaeus) 
Copium magnicorne (Rey) 
Copium teucrii (Host) 
Tingis { Tropidocheila) stachydis (Fieber) 
Teucrium flavum 

Copium clavicorne (Linnaeus) 
Teucrium marum 

Copium clavicorne (Linnaeus) 
Teucrium montanum 
Copium clavicorne (Linnaeus) 
Copium magnicorne (Rey) 
Copium teucrii (Host) 
Teucrium polium 
Copium bernardi Wagner 
Copium teucrii (Host) 
Teucrium scordium 

Copium clavicorne (Linnaeus) 
Teucrium scorodonium 
Copium clavicorne (Linnaeus) 
Copium reyi Wagner 
Teucrium supinum 

Copium teucrii (Host) 
Thymus sp. 
Acalypta gracilis (Fieber) 
Acalypta marginata (Wolff) 
Acalypta parvula (Fallen) 
Derephysia foliacea (Fallen) 
Dictyonota (Alcletha) tricornis (Schrank) 
Lasiacantha gracilis (Herrich-Schaeffer) 
Oncochila simplex (Herrich-Schaeffer) 
Thymus montanus 
Lasiacantha capucina (Germar) 



Thymus serpyllus 
Acalypta gracilis (Fieber) 
Acalypta nigrina (Fallen) 
Acalypta parvula (Fallen) 
Derephysia foliacea (Fallen) 
Lasiacantha capucina (Germar) 
Lauraceae 
Aperula citriodora 

Stephanitis takeyai Drake and Maa 
Camphora officinalis 

Pseudacysta perseae (Heidemann) 
Cinnamomum camphora 
Stephanitis fasciicarina Takeya 
Stephanitis takeyai Drake and Maa 
Stephanitis (Norba) aperta Horvath 
Cinnamomum pedunculatum 

Stephanitis {Norba) mendica Horvath 
Lindera citriodora 

Stephanitis takeyai Drake and Maa 
Lindera glauca 

Stephanitis ambigua Horvath 
Lindera obtusiloba 
Stephanitis ambigua Horvath 
Stephanitis takeyai Drake and Maa 
Lindera sericea 

Stephanitis takeyai Drake and Maa 
Lindera umbellata 

Stephanitis takeyai Drake and Maa 
Machilus sp. 

Stephanitis {Norba) esakii Takeya 
Machilus gamblei 

Stephanitis gallarum Horvath 
Machilus longifolia 

Stephanitis gallarum Horvath 
Machilus pseudo-longifolia 

Stephanitis gallarum Horvath 
Machilus thunbergii 
Stephanitis fasciicarina Takeya 
Stephanitis {Norba) aperta Horvath 
Stephanitis {Norba) esakii Takeya 
Stephanitis {Norba) exigua Horvath 
Nectandra sp. 
Acysta nectandrae Drake and Hamble- 
ton 
Ocotea sp. 
Pliobyrsa adversa (Drake and Hamble- 
ton) 



464 



U.S. NATIONAL MUSEUM BULLETIN 24 3 



Ocotea lanata 

Pliobyrsa infiexa (Drake and Hamble- 
ton) 
Ocotea pretiosa 

Acysta ocoteae Drake and Hambleton 

Leptopharsa ocoteae Drake and Ham- 
bleton 
Persea americana 

Pleseobyrsa chiriquensis (Champion) 
Persea carolinensis 

Pseudacysta perseae (Heidemann) 
Persea gratissima 

Pseudacysta perseae (Heidemann) 

Stephanitis {Norba) suffusa (Distant) 
Tetrandenia dolichocarpa 

Stephanitis {Norba) esakii Takeya 
Leguminosae 
Atheas fuscipes Champion 
Leptopharsa probala Drake and Ham- 
bleton 

Tigava gracilis Monte 

Tigava praecellens Stal 
Acacia sp. 

Nethersia setosa (Hacker) 
Acacia cavenia 

Coleopterodes liliputiana (Signoret) 
Acacia riparia 

Teleonemia prolixa (Stal) 
Aeschynomene sp. 

Atheas paganus Drake 
Aeschynomene falcata 

Atheas ornatipes Drake and Hambleton 
Aeschynomene nivea 

Atheas tristis Van Duzee 
Amorpha fruticosa 

Gargaphia amorphae (Walsh) 

Leptopharsa oblonga (Say) 
Arachis hypogaea 

Corythucha gossypii (Fabricius) 
Astragalus sp. 

Galeatus scrophicus Saunders 
Baptisia tinctoria 

Leptopharsa heidemanni (Osborn and 
Drake) 
Bauhinia sp. 

Vatiga longula (Drake) 
Bauhinia purpurea 

Cysteochila delineatus (Distant) 



Bauhinia variegata 

Cysteochila ablusa Drake 
Cajanus indicus 

Corythucha gossypii (Fabricius) 

Gargaphia penningtoni Drake 

Urentius euonymus Distant 
Canavalia ensiformis 

Campylotingis bondari (Drake) 

Corythucha gossypii (Fabricius) 

Gargaphia lunulata (Mayr) 

Gargaphia opima Drake 
Canavalia obtusifolia 

Gargaphia torresi Costa Lima 
Cassia sp. 

Campylotingis mollis Drake and Bondar 

Gargaphia solani Heidemann 

Leptopharsa mira Drake and Hamble- 
ton 

Leptopharsa principis Drake and Ham- 
bleton 

Teleonemia brevipennis Champion 

Vatiga cassiae (Drake and Hambleton) 
Cassia emarginata 

Corythucha gossypii (Fabricius) 
Cassia fistula 

Gargaphia lunulata (Mayr) 
Cassia imperialis 

Gargaphia lunulata (Mayr) 
Ceratonia sp. 

Elasmotropis testacea var. vicina Hor- 
vath 

Tingis (Lasiotropis) hellenica (Puton) 
Ceratonia siliqua 

Elasmotropis testacea (Herrich-Schaef- 
fer) 
Cercis canadensis 

Gargaphia tiliae Walsh 
Clitoria mariana 

Leptopharsa clitoriae (Heidemann) 
Cytisus scoparius 

Dictyonota (Alcletha) tricornis (Schrank) 
Dahlstedtia pinnata 

Corycera alboater Drake and Hamble- 
ton 
Dalbergia sp. 

Leptopharsa tenuatis Drake 



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465 



Derris elliptica 

Amblystira fuscitarsis Champion 

Amblystira opaca Champion 
Desmcdium sp. 

Atheas austroriparius Heidemann 

Atheas insignis Heidemann 

Atheas mimeticus Heidemann 

Leptopharsa machalana Drake and 
Hambleton 

Leptopharsa machalana var. vinnula 
Drake and Hambleton 
Dolichos lablab 

Campylotingis bondari (Drake) 

Gargaphia lunulata (Mayr) 
Dorycnium suffruticosum 

Phaenotropis parvula (Signoret) 
Erythrina berteroana 

Corythucha gossypii (Fabricius) 
Falcata comosa 

Leptopharsa oblonga (Say) 
Ferreirea spectabilis 

Campylotingis carvalhoi Drake and 
Hambleton 

Campylotingis machaerii Drake and 
Hambleton 
Genista sp. 

Catoplatus jabricii (Stal) 

Dictyonota pulchella Costa 
Genista anglica 

Dictyonota albipennis Baerensprung 
Genista scorpius 

Dictyonota pulchella Costa 
Genista tinctoria 

Catoplatus jabricii (Stal) 
Gleditsia triacanthos 

Anommatocoris coleopteratus (Kormilev) 
Glycine max 

Gargaphia lunulata (Mayr) 
Hosackia crassifolia 

Corythucha juscigera (Stal) 
Hymenaea stigonocarpa 

Amblystira peltogyne Drake and Ham- 
bleton 
Ichthyomethia grandifolia 

Leptopharsa tenuis (Champion) 
Ichthyomethia piscipula 

Corythucha gossypii (Fabricius) 



Indigofera sp. 

Monosteira minutula Montandon 
Kuhnistera sp. 

Leptopharsa oblonga (Say) 
Lathyrus nuttallii 

Corythucha distincta Osborn and Drake 
Lespedeza sp. 

Leptopharsa clitoriae (Heidemann) 
Lonchocarpus sp. 

Vatiga lonchocarpa (Drake and Ham- 
bleton) 
Lonchocarpus sericeus 

Amblystira fuscitarsis Champion 
Machaerium sp. 

Atheas laetantis Drake and Hambleton 

Campylotingis bondari (Drake) 

Campylotingis jansoni (Drake) 

Campylotingis lenatis Drake 

Campylotingis machaerii Drake and 
Hambleton 

Campylotingis prudens Drake and Ham- 
bleton 

Campylotingis tantilla Drake 

Corycera machaerii Drake and Ham- 
bleton 

Leptopharsa prudens Drake and Ham- 
bleton 
Machaerium angustifolium 

Atheas fiavipes Champion 

Atheas laetantis Drake and Hambleton 
Machaerium nictitans 

Leptopharsa machaerii Drake and Ham- 
bleton 
Machaerium oblongifolium 

Teleonemia validicornis Stal 
Machaerium oblongifolium v. sub- 
glabrum 

Teleonemia validicornis Stal 
Machaerium pedicillatum 

Leptopharsa ogloblini Drake 
Machaerium stipitatum 

Leptopharsa gracilenta (Champion) 
Machaerium villosum 

Campylotingis prudens Drake and Ham- 
bleton 
Medicago sp. 

Galeatus spinifrons (Fallen) 



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U.S. NATIONAL MUSEUM BULLETIN 243 



Medicago lupulina 

Catoplatus fabricii (Stal) 
Meibomia sp. 

Leptopharsa clitoriae (Heidemann) 
Meibomia adscendens 

Gargaphia lunulata (Mayr) 
Myrocarpus fastigiatus 

Acysta myrocarpi Drake and Poor 

Campylotingis clavata Drake and Ham- 
bleton 
Oxytropis sp. 

Acalypta lillianis Torre-Bueno 
Parosela citriodora 

Atheas nigricornis Champion 
Parosela emoryi 

Gargaphia insularis Van Duzee 
Peltogyne sp. 

Amblystira peltogyne Drake and Ham- 
bleton 
Petalostemon sp. 

Leptopharsa oblonga (Say) 
Petalostemon purpureum 

Atheas mimeticus Heidemann 
Phaseolus sp. 

Gargaphia penningtoni Drake 

Gargaphia torresi Costa Lima 
Phaseolus limensis 

Corythucha gossypii (Fabricius) 
Phaseolus lunatus 

Campylotingis bondari (Drake) 

Gargaphia lunulata (Mayr) 

Gargaphia torresi Costa Lima 
Phaseolus vulgaris 

Campylotingis bondari (Drake) 

Gargaphia lunulata (Mayr) 

Gargaphia subpilosa Berg 

Gargaphia torresi Costa Lima 
Robina pseudoacacia 

Stephanitis pyri (Fabricius) 
Sarothaumus sp. 

Dictyonota fuliginosa Costa 
Sarothaumus scoparius 

Dictyonota strichnocera Fieber 
Schrankia sp. 

Atheas austroriparius Heidemann 
Spartium sp. 

Catoplatus fabricii (Stal) 



Spartium junceum 

Aclypta nigrina (Fallen) 
Spartium scoparium 

Dictyonota fuliginosa Costa 

Dictyonota strichnocera Fieber 

Monosteira unicostata (Mulsant and 
Rey) 
Stylosanthes biflora 

Atheas insignis Heidemann 
Swartzia sp. 

Leptopharsa mira Drake and Hamble- 
ton 
Tamarindus indica 

Cysteochila endeca Drake 
Tephrosia sp. 

Pkaenotropis cleopatra (Horvath) 
Ulex sp. 

Dictyonota pulchella Costa 

Dictyonota strichnocera Fieber 
Vigna sinensis 

Campylotingis bondari (Drake) 
Zornia diphylla 

Gargaphia lunulata (Mayr) 

LlLIACEAE 

Yucca arborescens 
Corythucha caelata Uhler 

LOGANIACEAE 

Buddleia sp. 

Teleonemia brevipennis Champion 
Buddleia asiatica 

Tingis buddleiae Drake 

Lycopodiaceae 

Stephanitis {Norba) exigua Horvath 

Magnoliaceae 
Illicium religiosum 

Stephanitis svensoni Drake 
Liriodendron sp. 

Melanorhopala infuscata Parshley 

Malpighiaceae 
Byrsonima sp. 

Tingis neotropicalis Monte 
Byrsonima sericea 

Ulotingis uniseriata (Drake) 
Byrsonima verbascifolia 

Ulotingis uniseriata (Drake) 
Malpighia glabra 

Phymacysta tumida (Champion) 



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467 



Malpighia punicifolia 

Phymacysta tumida (Champion) 
Malpighia urens 

Phymacysta tumida (Champion) 
Stigmaphyllon sp. 
Gargaphia lunulata (Mayr) 
Malvaceae 

Compseuta picta Schouteden 

Macrocorytha rhomboptera (Fieber) 
Abelmoschus esculentus 

Gargaphia lunulata (Mayr) 
Abutilon sp. 

Corythucha hispida Uhler 

Gargaphia penningtoni Drake 

Urentius euonymous Distant 
Abutilon muticum 

Urentius euonymus Distant 
Abutilon pannosum 

Urentius euonymus Distant 
Abutilon umbellatum 

Corythucha championi Drake and Cob- 
ben 
Althaea rosea 

Corythaica carinata Uhler 
Ayenia magna 

Corythucha championi Drake and Cob- 
ben 
Callirhoe involucrata 

Teleonemia scrupulosa Stal 
Gossypium sp. 

Corythaica costata Gibson 

Corythaica monacha (Stal) 

Corythucha gossypii (Fabricius) 

Gargaphia subpilosa Berg 

Gargaphia torresi Costa Lima 
Gossypium arboreum 

Gargaphia lunulata (Mayr) 
Gossypium herbaceum 

Gargaphia solani Heidemann 
Hibiscus sp. 

Corythucha gossypii (Fabricius) 
Hibiscus esculentus 

Corythucha gossypii (Fabricius) 

Gargaphia acmonis Drake and Hamble- 
ton 

Gargaphia lunulata (Mayr) 
Hibiscus rosa-sinensis 

Corythucha gossypii (Fabricius) 



Hibiscus syriacus 

Gargaphia torresi Costa Lima 
Malva sp. 

Gargaphia iridescens Champion 
Malvaviscus arboreus 
Leptopharsa rumiana Drake and Ham- 
bleton 
Malvaviscus drummondii 
Calotingis knighti Drake 
Sida sp. 
Gargaphia penningtoni Drake 
Gargaphia torresi Costa Lima 
Haedus sidae (Drake and Poor) 
Sida acuta 

Gargaphia penningtoni Drake 
Sida cordifolia 
Corythaica monacha (Stal) 
Urentius euonymus Distant 
Sida glomerata 

Corythaica monacha (Stal) 
Sida procumbens 

Corythaica carinata Uhler 
Sida rhombifolia 

Corythaica monacha (Stal) 
Sida ulmifolia 

Corythaica monacha (Stal) 
Sphaeralcea sp. 

Corythucha sphaeralceae Drake 
Sphaeralcea angustifolia 

Teleonemia nigrina Champion 
Sphaeralcea miniata 

Corythaica cucullata (Berg) 
Urena lobata 
Gargaphia lunulata (Mayr) 
Haedus vicarius (Drake) 
Leptopharsa machalana var. vinnula 
Drake and Hambleton 
Urena lobata v. sinuata 
Haedus vicarius (Drake) 
Wissadula sp. 
Gargaphia penningtoni Drake 
Gargaphia subpilosa Berg 

Melastomataceae 
Miconia sp. 
Leptopharsa cognata Drake and Ham- 
bleton 
Leptopharsa miconiae Drake and Ham- 
bleton 



468 



U.S. NATIONAL MUSEUM BULLETIN 243 



Menispermaceae 
Cissampelos glaberrima 

Gargaphia fiexuosa (Stal) 
Cissampelos tamoides 

Gargaphia flexuosa (Stal) 
Cocculus sp. 

Stephanitis subjasciata Horvath 
Cocculus trilobus 

Stephanitis subjasciata Horvath 
Stephania hernandifolia 

Stephanitis queenslandensis Hacker 

MONIMIACEAE 

Mollinedia sp. 

Pliobyrsa mollinediae (Drake and Ham- 
bleton) 
Siparuna guianensis 

Leptopharsa distincornis Drake 

MORACEAE 

Artocarpus incisa 

Nesocypselas dicysta Kirkaldy 
Artocarpus integrifolia 

Alloiothucha artocarpi (Horvath) 

Leptopharsa angustata (Champion) 

Leptopharsa artocarpi Drake and Ham- 
bleton 

Leptopharsa vittipennis (Stal) 

Stephanitis typica (Distant) 

Stephanitis (Menodora) chariesis Drake 
and Mohanasundarum 
Broussonetia papyrifera 

Corythucha ciliata (Say) 
Chlorophora tinctoria 

Leptopharsa difficilis Drake and Ham- 
bleton 
Ficus sp. 

Naochila kivuensis (Schouteden) 

Perissonemia ecmeles Drake and Moha- 
nasundarum 

Tigava brevicollis Monte 
Ficus elastica 

Leptopharsa fici Drake and Hambleton 
Ficus subtriplinervia 

Leptopharsa fici Drake and Hambleton 
Humulus sp. 

Dictyla humuli (Fabricius) 

Tingis (Tropidocheila) pilosa Hummel 
Morus rubra 

Corythucha pallida Osborn and Drake 



Musaceae 
Musa sp. 

Stephanitis typica (Distant) 
Musa paradisiaca 

Corythucha gossypii (Fabricius) 
Musa paradisiaca v. sapientum 
Stephanitis typica (Distant) 
Stephanitis {Menodora) sondaica Hor- 
vath 

Myoporaceae 
Myoporum sandwicense 
Teleonemia scrupulosa Stal 

Myricaceae 
Myrica gale 

Stephanitis oberti (Kolenati) 

Myrtaceae 

Ulotingis decor Drake and Hambleton 

Ulotingis nitor Drake and Hambleton 
Eugenia rhombea 

Hybopharsa colubra (Van Duzee) 
Pimenta pimenta 

Hybopharsa colubra (Van Duzee) 
Psidium sp. 

Gargaphia lunulata (Mayr) 
Psidium guajava 

Ulotingis brasiliensis (Drake) 
Nyctaginaceae 
Pisonia sp. 

Acanthocheila armigera (Stal) 
Pisonia aculeata 

Acanthocheila sigillata Drake and Bru- 
ner 
Pisonia domingensis 

Acanthocheila spinicosta Van Duzee 
Pisonia tomentosa 

Acanthocheila amigera (Stal) 
Torrubia fragrans 

Acanthocheila spinicosta Van Duzee 
Ochnaceae 
Ouratea sp. 

Acanthocheila armigera (Stal) 
Oleaceae 
Adelia acuminata 

Leptoypha mutica (Say) 
Chionanthus virginica 

Leptoypha mutica (Say) 



LACEBUGS OF THE WORLD 



469 



Fraxinus sp. 
Corythucha ciliata (Say) 
Leptoypha mutica (Say) 
Fraxinus americana 

Leptoypha mutica (Say) 
Fraxinus berlandierana 

Leptoypha barberi Drake and Runoff 
Fraxinus caroliniana 

Leptoypha costata Parshley 
Fraxinus mandshurica v. japonica 
Leptoypha wuorentausi (Lindberg) 
Fraxinus oregona 

Leptoypha minor McAtee 
Fraxinus pubinervis 

Leptoypha wuorentausi (Lindberg) 
Fraxinus rhynchophylla 

Leptoypha wuorentausi (Lindberg) 
Fraxinus velutina 

Leptoypha minor McAtee 
Jasminum sp. 

Plerochila horvathi (Schouteden) 
Jasminum dichotomum 

Froggattia hargreavesi Drake 
Jasmimum pubescens 

Corythauma ayyari (Drake) 
Ligustrum sp. 

Stephanitis pyri (Fabricius) 
Notelaea longifolia 
Froggattia olivinia Froggatt 

Olea sp. 

Catoplatus dilatatus (Jakovlev) 
Plerochila rutshurica Schouteden 

Olea chrysophylla 
Cysteochila pollens Horvath 

Olea europaea 
Froggattia olivinia Froggatt 
Olastrida oleae Schouteden 
Plerochila australis (Distant) 
Plerochila horvathi (Schouteden) 

Olea verrucosa 
Cysteochila sordida (Stal) 

Osmanthus americana 
Leptoypha mcateei Drake 

Orchidaceae 

(Intercepted at port-of-entry) 

Cattleya dowiana var. aurea 
Pleseobyrsa plicata (Champion) 



Cattleya schroederae 
Phatnoma barberi Drake 
Phatnoma varians Drake 
Epidendrum cochleatum 
Phatnoma guatemalana Drake 
Palmae 
Roystonea regia 

Corythucha gossypii (Fabricius) 
Thrinax wendlandiana 
Allotingis binotata (Drake and Bruner) 

Papilionaceae 
Campylotingis carvalhoi Drake and 

Hambleton 
Tigava gracilis Monte 
Tigava praecellens Stal 

Passifloraceae 
Passiflora caerulea 
Corythaica cyathicollis (Costa) 
Gargaphia lunulata (Mayr) 
Passiflora foetida 

Corythaica carinata Uhler 
Passiflora violacea 
Gargaphia lunulata (Mayr) 

Phytolaccaceae 
Phytolacca acinosa 
Cochlochila lewisi (Scott) 

Pemaceae 
Abies alba 

Tingis {Tropidocheila) geniculata (Fie- 
ber) 
Juniperus sp. 

Physatocheila dumetorum (Herrich- 
Schaeffer) 
Juniperus communis 

Tingis {Tropidocheila) geniculata (Fie- 
ber) 
Juniperus foetidissima 

Tingis elongata (Fieber) 
Juniperus phoenicea 
Copium intermedium (Rey) 
Tingis {Tropidocheila) griseola (Puton) 
Larix decidua 

Dictyonota strichnocera Fieber 
Pinus sp. 
Physatocheila dumetorum (Herrich- 

Schaeffer) 
Tingis ampliata (Herrich-Schaeffer) 



470 



U.S. NATIONAL MUSEUM BULLETIN 243 



Pinus austriaca 

Tingis cardui (Linnaeus) 
Pinus halepensis 

Monosteira unicostata (Mulsant and 
Rey) 
Pinus monophylla 

Dictyla labeculata (Uhler) 
Pinus sylvestris 

Dictyonota strichnocera Fieber 

Stephanitis oberti (Kolenati) 

Tingis cardui (Linnaeus) 

PlPERACEAE 

Piper nigrum 

Diconocoris distanti Drake 
Plantaginaceae 
Plantago sp. 

Galeatus scrophicus Saunders 
Plantago aristata 

Teleonemia nigrina Champion 
Plantago cynops 

Acalypta gracilis (Fieber) 
Plantago maritima 

Agramma confusum (Puton) 
Plantago media 

Derephysia Joliacea (Fallen) 

Platanaceae 
Platanus sp. 

Corythucha conjraterna Gibson 
Platanus occidentalis 

Corythucha ciliata (Say) 

Plumbaginaceae 
Armeria vulgaris 

Derephysia Joliacea (Fallen) 
Statice sp. 

Agramma nigrum Fieber 

Tingis auriculata (Costa) 
Statice armeria 

Derephysia Joliacea (Fallen) 

Galeatus maculatus (Herrich-Schaeffer) 

POLYGALACEAE 

Bredemeyera sp. 

Amblystira pallipes (St°l) 

POLYGONACEAE 

Eriogonum sp. 

Corythaica venusta (Champion) 
Teleonemia nigrina Champion 



POLYPORACEAE 

Coriolus versicolor 

Acalypta musci (Schrank) 

Acalypta parvula (Fallen) 
Polyporus hirsutus 

Acalypta parvula (Fallen) 
Polyporus unicolor 

Acalypta musci (Schrank) 
Trametes gibbosa 

Acalypta musci (Schrank) 

Acalypta parvula (Fallen) 

POLYTRICHACEAE 

Polytrichum sp. 

Acalypta lillianis Torre-Bueno 
Proteaceae 
Banksia robur minor 

Euaulana tasmaniae Drake 
Protea sp. 

Sanazarius productus Distant 
Roupala sp. 

Amblystira pensa Drake and Hamble- 
ton 

Corycera gibbosa Monte 
Rhamnaceae 
Ceanothus sp. 

Corythucha obliqua Osborn and Drake 

Leptoypha minor McAtee 

Melanorhopala infuscata Parshley 
Ceanothus americanus 

Gargaphia angulata Heidemann 

Melanorhopala infuscata Parshley 
Ceanothus cordulatus 

Corythucha obliqua Osborn and Drake 
Ceanothus sanguineus 

Corythucha obliqua Osborn and Drake 
Zizyphus jujuba 

Monosteira minutula Montandon 
Zizyphus spina-christi 

Monosteira minutula Montandon 
Rosaceae 
Amelanchier sp. 

Corythucha cydoniae (Fitch) 
Amelanchier canadensis 

Corythucha cydoniae (Fitch) 
Amelanchier intermedia 

Corythucha cydoniae (Fitch) 

Corythucha juglandis (Fitch) 



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471 



Amygdalus sp. 

Physatocheila dumetorum (Herrich- 
Schaeffer) 

Stephanitis pyri (Fabricius) 
Amygdalus communis 

Monosteira unicostata (Mulsant and 
Rey) 
Cerasus sp. 

Monosteira unicostata (Mulsant and 
Rey) 
Chaenomeles sp. 

Corythucha cydoniae (Fitch) 
Chaenomeles cathayensis 

Stephanitis nashi Esaki and Takeya 
Chaenomeles japonica 

Stephanitis pyri (Fabricius) 
Cormus sp. 

Stephanitis pyri (Fabricius) 
Cormus tschonoskii 

Stephanitis nashi Esaki and Takeya 
Cotoneaster hupehensis 

Corythucha cydoniae (Fitch) 
Cotoneaster integerrima 

Stephanitis pyri (Fabricius) 
Crataegus sp. 

Corythucha bellula Gibson 

Corythucha brunnea Gibson 

Corythucha cydoniae (Fitch) 

Physatocheila cost at a subsp. smreczyn- 
skii China 

Physatocheila dumetorum (Herrich- 
Schaeffer) 

Stephanitis nashi Esaki and Takeya 

Stephanitis pyri (Fabricius) 
Crataegus albicans 

Corythucha bellula Gibson 
Crataegus coccinea 

Corythucha cydoniae (Fitch) 
Crataegus cordata 

Corythucha cydoniae (Fitch) 
Crataegus crus-galli 

Corythucha cydoniae (Fitch) 

Stephanitis pyri (Fabricius) 
Crataegus cuneata 

Stephanitis nashi Esaki and Takeya 
Crataegus monogyna 

Monosteira unicostata (Mulsant and 
Rey) 



Physatocheila dumetorum (Herrich- 
Schaeffer) 
Crataegus neonuvialis 

Corythucha bellula Gibson 
Crataegus oxyacantha 

Physatocheila dumetorum (Herrich- 
Schaeffer) 
Crataegus parvifolia 

Corythucha cydoniae (Fitch) 
Crataegus pruninosa 

Corythucha bellula Gibson 
Crataegus punctata 

Corythucha bellula Gibson 
Crataegus pyracantha 

Stephanitis pyri (Fabricius) 
Crataegus tomentosa 

Corythucha cydoniae (Fitch) 
Cydonia vulgaris 

Corythucha cydoniae (Fitch) 

Derephysia foliacea (Fallen) 

Monosteira unicostata (Mulsant and 
Rey) 

Stephanitis pyri (Fabricius) 
Dryas sp. 

Acalypta lillianis Torre -Bueno 
Heteromeles arbutifolia 

Corythucha incurvata Uhler 
Kerria japonica 

Stephanitis nashi Esaki and Takeya 
Malus sp. 

Corythucha cydoniae (Fitch) 

Stephanitis nashi Esaki and Takeya 
Malus communis 

Stephanitis pyri (Fabricius) 
Malus noribunda 

Stephanitis nashi Esaki and Takeya 
Malus manschurica 

Leptoypha capitata (Jakovlev) 
Mespilus oxyacantha 

Physatocheila dumetorum (Herrich- 
Schaeffer) 
Persica vulgaris 

Monosteira unicostata var. buccata Hor- 
vath 
Potentilla sp. 

Catoplatus nigriceps Horvath 

Galeatus maculalus (Herrich-Schaeffer) 



472 



U.S. NATIONAL MUSEUM BULLETIN 243 



Potentilla freyiana 

Stephanitis nashi var. suigensis Saito 
Prunus sp. 

Monosteira unicostata (Mulsant and 
Rey) 

Stephanitis nashi Esaki and Takeya 
Prunus americana 

Corythucha pergandei Heidemann 
Prunus amygdalus 

Monosteira unicostata (Mulsant and 
Rey) 
Prunus armeniaca 

Stephanitis pyri (Fabricius) 
Prunus avium 

Stephanitis pyri (Fabricius) 
Prunus cerasus 

Gargaphia subpilosa Berg 

Monosteira unicostata (Mulsant and 
Rey) 

Stephanitis pyri (Fabricius) 
Prunus demissa 

Corythucha padi Drake 
Prunus domestia 

Physatocheila dumetorum (Herrich- 
Schaeffer) 
Prunus lusitanica 

Stephanitis pyri (Fabricius) 
Prunus mahaleb 

Stephanitis pyri (Fabricius) 
Prunus marasca 

Stephanitis pyri (Fabricius) 
Prunus padus 

Physatocheila dumetorum (Herrich- 
Schaeffer) 

Physatocheila costata subsp. smreczynskii 
China 
Prunus persica 

Corythucha associata Osborn and Drake 

Corythucha gossypii (Fabricius) 

Gargaphia subpilosa Berg 

Monosteira unicostata (Mulsant and 
Rey) 

Stephanitis pyri (Fabricius) 
Prunus serotina 

Corythucha associata Osborn and Drake 

Corythucha pruni Osborn and Drake 

Gargaphia tiliae (Walsh) 



Prunus spinosa 

Physatocheila dumetorum (Herrich- 
Schaeffer) 
Pyrus sp. 

Stephanitis nashi Esaki and Takeya 

Stephanitis pyri (Fabricius) 
Pyrus americana 

Corythucha cydoniae (Fitch) 

Corythucha pallipes Parshley 
Pyrus aria 

Stephanitis pyri (Fabricius) 

Tingis angustata (Herrich-Schaeffer) 
Pyrus aucuparia 

Corythucha cydoniae (Fitch) 

Stephanitis pyri (Fabricius) 
Pyrus communis 

Gargaphia lunulata (Mayr) 

Gargaphia subpilosa Berg 

Monosteira unicostata (Mulsant and 
Rey) 

Physatocheila confinis Horvath 

Physatocheila dumetorum (Herrich- 
Schaeffer) 

Physatocheila costata subsp. smreczynskii 
China 

Stephanitis pyri (Fabricius) 
Pyrus malus 

Corythucha arcuata (Say) 

Corythucha salicata Gibson 

Monosteira unicostata (Mulsant and 
Rey) 

Physatocheila dumetorum (Herrich- 
Schaeffer) 

Physatocheila costata subsp. smreczynskii 
China 

Stephanitis pyri (Fabricius) 
Pyrus mamorensis 

Monosteira unicostata (Mulsant and 
Rey) 
Pyrus melanocarpa 

Corythucha cydoniae (Fitch) 
Pyrus prunifolia 

Corythucha pergandei Heidemann 
Pyrus ussuriensis 

Leptoypha capitata (Jakovlev) 

hanitis nashi Esaki and Takeya 



LACEBUGS OF THE WORLD 



473 



Pyracantha coccinea 

Corythucha cydoniae (Fitch) 
Rosa sp. 

Gargaphia lunulata (Mayr) 

Stephanitis pyri (Fabricius) 
Rubus sp. 

Corythucha juglandis (Fitch) 
Sorbus sp. 

Corythucha heidemanni Drake 

Corythucha juglandis (Fitch) 

Physatocheila costata subsp. smreczynskii 
China 
Sorbus americana 

Corythucha juglandis (Fitch) 
Sorbus aria 

Stephanitis pyri (Fabricius) 
Sorbus aucuparia 

Physatocheila costata subsp. smreczynskii 
China 
Sorbus cretica 

Tingis angustata (Herrich-Schaeffer) 
Sorbus intermedia 

Physatocheila costata subsp. smreczynskii 
China 
Sorbus torminalis 

Stephanitis pyri (Fabricius) 
Vauquelinia californica 
Corythucha sagillata Drake 

RUBIACEAE 

Acanthotingis apicicornis Monte 
Leptopharsa distinconis Drake 
Leptopharsa divisa (Champion) 
Leptopharsa fiava Monte 
Leptopharsa furculata (Champion) 

Cephaelis hastisepala 
Leptopharsa fortis Drake and Hamble- 
ton 

Cephalanthus sp. 
Corythucha cydoniae (Fitch) 
Corythucha floridana Heidemann 

Cephalanthus occidentalis 
Corythucha cydoniae (Fitch) 
Corythucha floridana Heidemann 

Cinchona sp. 

Teleonemia prolixa (Stal) 

Diodia conferta 
Corythaica umbrosa (Monte) 



Faramea sp. 
Leptopharsa farameae Drake and Hara- 
bleton 
Faramea montevidensis 
Stephanitis farameae Drake and Ham- 
bleton 
Galium sp. 

Lasiacantha capucina (Germar) 
Guettarda viburnoides 

Pachycysta hambletoni Drake and Poor 
Morinda sp. 

Dulinius conchatus Distant 
Posoqueria acutifolia 
Leptopharsa posoqueriae Drake and 
Hambleton 
Psychotria sp. 
Leptopharsa fortis Drake and Hamble- 
ton 
Leptopharsa psycho triae Drake and 
Hambleton 
Psychotria hancorniaefolia 
Leptopharsa ignota Drake and Hamble- 
ton 
Psychotria suterella 

Leptopharsa hyaloptera (Stal) 
Richardia brasiliensis 
Corythaica monacha (Stal) 
Corythaica umbrosa (Monte) 
Rudgea sp. 
Leptopharsa perbona Drake 
Leptopharsa rudgeae Drake and Ham- 
bleton 
Rudgea blanche tiana 
Leptopharsa nota Drake and Hamble- 
ton 
Rudgea villiflora 

Leptopharsa furcata (Stal) 
Sickingia tinctoria 
Gargaphia patria (Drake and Hamble- 
ton) 

RUTACEAE 

Citrus aurantium 

Leptobyrsa decora Drake 
Citrus medica 

Corythucha gossypii (Fabricius) 
Zanthoxylum martinicense 

Corythucha gossypii (Fabricius) 



474 



U.S. NATIONAL MUSEUM BULLETIN 243 



Salicaceae 
Populus sp. 

Corythucha mollicula Osborn and Drake 

Monosteira unicostata (Mulsant and 
Rey) 

Stephanitis pyri (Fabricius) 

Tingis populi Takeya 

Tingis (Tropidocheila) geniculata (Fie- 
ber) 
Populus alba 

Monosteira unicostata (Mulsant and 
Rey) 
Populus balsamifera 

Corythucha elegans Drake 
Populus candicans 

Leptoypha minor McAtee 
Populus grandidentata 

Corythucha elegans Drake 
Populus nigra 

Monosteira unicostata (Mulsant and 
Rey) 

Tingis lanigera (Puton) 

Tingis populi Takeya 
Populus tremula 

Monosteira unicostata (Mulsant and 
Rey) 

Physatocheila costata (Fabricius) 

Tingis populi Takeya 
Populus tremuloides 

Corythucha elegans Drake 
Salix sp. 

Corythucha elegans Drake 

Corythucha mollicula Osborn and Drake 

Corythucha pallipes Parshley 

Monosteira lobulifera Reuter 

Monosteira unicostata (Mulsant and 
Rey) 

Stephanitis takeyai Drake and Maa 

Tingis populi Takeya 
Salix discolor 

Corythucha mollicula Osborn and Drake 
Salix hookeriana 

Corythucha mollicula Osborn and Drake 
Salix koreensis 

Tingis populi Takeya 
Salix lasiandra 

Corythucha salicata Gibson 

Physatocheila variegata Parshley 



Salix purpurea 

Dictyla salicorum (Baba) 
Salix scouleriana 

Corythucha salicata Gibson 
SalLx sieboldiana 

Corythucha pallipes Parshley 
Sapindaceae 
Serjania sp. 

Amblystira pallipes (Stal) 

Amblystira silvicola Drake 

Gargaphia obliqua Stal 

Gargaphia serjaniae Drake and Hara- 
bleton 

Planibyrsa montei Drake and Hamble- 
ton 

Sapotaceae 
Lucuma sp. 
Aristobyrsa latipennis (Champion) 

Saxifragaceae 
Hydrangea integra 

Stephanitis hydrangeae Drake and Maa 
Ribes sp. 

Corythucha mollicula Osborn and Drake 
Ribes nigrum 

Stephanitis pyri (Fabricius) 
Ribes oxyacanthoides 

Corythucha bellula Gibson 

SCROPHULARIACEAE 

Adenostegia filifolia 

Teleonemia nigrina Champion 

Teleonemia notata Champion 
Adenostegia pilosa 

Teleonemia nigrina Champion 

Teleonemia notata Champion 
Penstemon sp. 

Teleonemia montivaga Drake 
Verbascum sp. 

Dictyonota (Alcletha) tricornis (Schrank) 

Tingis ampliata (Herrich-Schaeffer) 

Tingis (Lasiotropis) reticulata Herrich- 
Schaeffer 
Verbascum phlomoides 

Tingis (Lasiotropis) ragusana (Fieber) 
Verbascum thapsus 

Tingis (Lasiotropis) reticulata Herrich- 
Schaeffer 



LACEBUGS OF THE WORLD 



475 



SOLANACEAE 

Brunfelsia sp. 

Gargaphia brunjelsiae Monte 

Gargaphia munda (Stal) 
Brunfelsia hopeana 

Gargaphia brunjelsiae Monte 

Gargaphia munda (Stal) 
Lycopersicum esculentum 

Corythaica costata Gibson 

Corythaica cyathicollis (Costa) 

Gargaphia solani Heidemann 
Nicotiana tabacum 

Acanthocheila armigera (Stal) 
Solanum sp. 

Corythucha juscomaculata (Stal) 

Corythucha globigera Breddin 

Corythucha nobilis Drake and Bondar 

Gargaphia angulata Heidemann 

Gargaphia crotonae Drake and Hamble- 
ton 

Gargaphia decoris Drake 

Gargaphia iridescens Champion 

Gargaphia munda (Stal) 

Gargaphia nigrinervis Stal 

Gargaphia solani Heidemann 
Solanum argillicolum 

Corythaica cyathicollis (Costa) 
Solanum auriculatum (?) 

Gargaphia decoris Drake 
Solanum balbisii 

Corythaica cyathicollis (Costa) 
Solanum bonariense 

Corythaica cyathicollis (Costa) 
Solanum capsicastrum 

Gargaphia oreades Drake 
Solanum carolinense 

Gargaphia solani Heidemann 
Solanum concinnum 

Gargaphia formosa (Stal) 
Solanum dulcamara 

Galeatus spinifrons (Fallen) 
Solanum elaeagnifolium 

Corythaica cyathicollis (Costa) 

Gargaphia solani Heidemann 
Solanum gilo 

Corythaica cyathicollis (Costa) 
Solanum grandiflorum 

Corythaica cyathicollis (Costa) 



Solanum hindsianum 
Gargaphia gentilis Van Duzee 

Solanum hirtum 
Corythaica cyathicollis (Costa) 

Solanum juripeba 
Corythaica cyathicollis (Costa) 

Solanum lycopersicum 
Corythaica cyathicollis (Costa) 
Corythucha gossypii (Fabricius) 

Solanum melongena 
Corythaica carinata Uhler 
Corythaica cyathicollis (Costa) 
Corythucha gossypii (Fabricius) 
Gargaphia opacula Uhler 
Gargaphia solani Heidemann 
Urentius hystricellus (Richter) 

Solanum nigrum 
Gargaphia oreades Drake 

Solanum nigrum americanum 
Corythaica cyathicollis (Costa) 

Solanum paniculatum 
Corythaica cyathicollis (Costa) 

Solanum pulverulentum 
Corythaica cyathicollis (Costa) 

Solanum quitoense 
Corythaica cyathicollis (Costa) 

Solanum racemifiorum 
Corythaica cyathicollis (Costa) 

Solanum racemosum 
Corythaica cyathicollis (Costa) 

Solanum sisymbrifolium 
Corythaica cyathicollis (Costa) 

Solanum tabacifolium 
Corythucha cyathicollis (Costa) 

Solanum torvum 
Corythaica cyathicollis (Costa) 
Corythucha gossypii (Fabricius) 
Corythucha spinosa (Duges) 
Tigava pulchella Champion 

Solanum tuberosum 
Corythaica cyathicollis (Costa) 
Gargaphia solani Heidemann 
Gargaphia subpilosa Berg 

Solanum variabile 
Corythaica cyathicollis (Costa) 

Solanum verbascifolium 
Dictyla rasilis (Drake and Maa) 



476 



U.S. NATIONAL MUSEUM BULLETIN 243 



Staphyleaceae 
Staphylea trifolia 

Corythucha bulbosa Osborn and Drake 
Sterculiaceae 
Hermannia micropetela 

Urentius vepris Drake 

Styracaceae 
Styrax japonica 

Stephanitis takeyai Drake and Maa 
Thymeleaceae 
Pimelea sp. 

Oncophysa vesiculata (Stal) 
Tiliaceae 
Corchorus hirsutus 

Corythaica carinata Uhler 
Tilia sp. 

Gargaphia tiliae (Walsh) 

Stephanitis pyri (Fabricius) 
Tilia americana 

Corythucha juglandis (Fitch) 

Corythucha pallida Osborn and Drake 

Gargaphia tiliae (Walsh) 
Tilia pubescens 

Gargaphia tiliae (Walsh) 
Triumfetta sp. 

Gargaphia torresi Costa Lima 
Triumfetta fiavescens 

Compseuta montandoni Distant 
Triumfetta grossulariaefolia 

Corythucha spinosa (Duges) 
Triumfetta rhombeaefolia 

Ammianus schoutedeni Distant 
Triumfetta rhomboidea 

Corythucha juscomaculata (Stal) 
Triumfetta semitriloba 

Corythucha juscomaculata (Stal) 

TURNERACEAE 

Piriqueta ovata 
Corythaica carinata Uhler 
Typhaceae 
Typha latifolia 
Agramma atricapillum (Spinola) 
Agramma intermedium (Wagner) 
Ulmaceae 
Celtis brasiliensis 
Atheas placentis Drake and Poor 
Corythucha pellucida Drake and Ham- 
bleton 



Celtis crassifolia 

Corythucha pergandei Heidemann 
Celtis ferruginea 

Corythucha pellucida Drake and Ham- 
bleton 
Celtis mississippiensis 

Corythucha celtidis var. mississippiensis 
Drake 
Celtis occidentalis 

Corythucha celtidis Osborn and Drake 
Ulmus sp. 

Corythucha pergandei Heidemann 
Ulmus americana 

Corythucha heidemanni Drake 

Corythucha ulmi Osborn and Drake 
Ulmus campestris 

Stephanitis pyri (Fabricius) 
Ulmus effusa 

Stephanitis pyri (Fabricius) 
Ulmus fulva 

Corythucha heidemanni Drake 

Corythucha ulmi Osborn and Drake 
Ulmus glabra camperdownii 

Corythucha ulmi Osborn and Drake 
Ulmus thomasii 

Corythucha ulmi Osborn and Drake 

Umbelliferae 
Caucalis daucoides 

Tingis auriculata (Costa) 

Tingis auriculata var. dauci Horvath 
Daucus carota 

Tingis auriculata (Costa) 

Tingis auriculata var. dauci Horvath 
Eryngium sp. 

Catoplatus carthusianus (Goeze) 

Catoplatus carthusianus var. albidus 
(Herrich-Schaeffer) 

Catoplatus horvathi (Puton) 
Eryngium amethystinum 

Catoplatus carthusianus (Goeze) 
Eryngium campestre 

Catoplatus carthusianus (Goeze) 

Catoplatus horvathi (Puton) 

Catoplatus nigriceps Horvath 
Eryngium maritinum 

Catoplatus carthusianus (Goeze) 



LACEBUGS OF THE WORLD 



477 



Seseli glaucum 

Catoplatus carthusianus (Goeze) 

Catoplatus carthusianus var. albidus 
(Herrich-Schaeffer) 
Torilis arvensis 

Tingis auriculata (Costa) 
Urticaceae 

Corythucha clara Drake and Hambleton 
Boehmeria cylindrica v. drummondiana 

Leptopharsa machalana var. vinnula 
Drake and Hambleton 
Boehmeria spicata 

Cysteochila fieberi (Scott) 
Phyllostylon rhamnoides 

Dicrotingis digitalis (Drake) 
Pipturus sp. 

Phatnoma veridica Drake and Maa 
Vacciniaceae 
Myrtillus uliginosus 

Stephanitis oberti (Kolenati) 

Verbenaceae 
Callicarpa americana 

Teleonemia belfragii Stal 
Clerodendron sp. 

Paracopium hamadryas (Drake) 
Clerodendron buchholzi 

Paracopium hamadryas (Drake) 
Clerodendron giletii 

Paracopium bequaerti (Schouteden) 
Clerodendron micans 

Paracopium dauphinicum Drake 
Clerodendron phlomoides 

Paracopium cingalense (Walker) 
Clerodendron subreniforme 

Paracopium hamadryas (Drake) 
Duranta repens 

Teleonemia notata Champion 
Gmelina arborea 

Tingis (Caenotingis) beesoni Drake 
Gmelina leichardtii 

Lasiacantha compta (Drake) 
Kalaharia (Clerodendron) spinescens 

Paracopium stolidum (Horvath) 
Lantana sp. 

Compseuta lefroyi Distant 

Corythaica venusta (Champion) 



Corythauma ayyari (Drake) 

Habrochila africana Drake 

Haedus pictus (Distant) 

Naochila sufflata (Drake and Poor) 

Phatnoma takasago Takeya 

Teleonemia bifasciata Champion 

Teleonemia carmelana (Berg) 

Teleonemia elata Drake 

Teleonemia notata Champion 

Teleonemia variegata Champion 
Lantana aculeata 

Teleonemia scrupulosa Stal 
Lantana brasiliensis 

Teleonemia scrupulosa Stal 
Lantana camara 

Corythucha spinosa (Duges) 

Teleonemia limbata (Stal) 

Teleonemia prolixa (Stal) 

Teleonemia sacchari (Fabricius) 

Teleonemia scrupulosa (Stal) 

Teleonemia validicornis Stal 
Lantana canescens 

Teleonemia sacchari (Fabricius) 

Teleonemia scrupulosa Stal 

Teleonemia syssita Drake and Cobben 
Lantana involucrata 

Teleonemia sacchari (Fabricius) 
Lantana montevidensis 

Teleonemia scrupulosa Stal 
Lippia alba 

Teleonemia scrupulosa Stal 
Lippia juncea 

Teleonemia carmelana (Berg) 
Premna sp. 

Eteoneus esakii Drake 
Premna gaudichaudii 
Berotingis guamensis (Drake) 
Berotingis yapensis (Drake) 
Rhaphithamnus spinosus 

Teleonemia carmelana (Berg) 
Tectona grandis 

Cochlochila nilgiriensis (Distant) 
Verbena sp. 

Teleonemia nigrina Champion 
Vitex heterophylla 
Cromerus kalshoveni Drake 



681-552—64- 



-32 



478 



U.S. NATIONAL MUSEUM BULLETIN 243 



Vitex pubescens 

Ammianus ravanus (Kirkaldy) 
Vitex ruganda 

Dasytingis rudis Drake and Poor 
Vitex trifolia 

Ammianus ravanus (Kirkaldy) 

Tingis buddleiae Drake 

Vitaceae 
Cissus japonica 
Cysteochila chiniana Drake 

ZlNGIBERACEAE 

Alpina chinensis 
Stephanitis typica (Distant) 



Alpina japonica 

Stephanitis typica (Distant) 
Alpina nutans 

Stephanitis typica (Distant) 
Alpina speciosa 

Stephanitis typica (Distant) 
Elettaria cardamomum 

Stephanitis typica (Distant) 
Hedychium sp. 

Stephanitis typica (Distant) 
Zingiber kawagoii 

Stephanitis typica (Distant) 
Zingiber mioga 

Stephanitis typica (Distant) 



Host Plants Cited Only as Common Names 



Alder 
Corythucha padi Drake 
Corythucha salicata Gibson 
Leptopharsa clitoriae (Heidemann) 
Physatocheila variegata Parshley 
Ash 
Derephysia foliacea (Fall6n) 
Dictyonota mitoris Drake and Hsiung 
Leptoypha drakei McAtee 
Avocado 
Pleseobyrsa boliviana Drake and Poor 
Tigava pulchella Champion 
Bamboo 
Leptodictya {Hanuala) comes Drake and 

Hambleton 
Leptodictya {Hanuala) cretata Champion 
Leptodictya {Hanuala) decor Drake and 

Hambleton 
Leptodictya {Hanuala) ecuadoris Drake 

and Hambleton 
Leptodictya {Hanuala) fraterna Monte 
Leptodictya {Hanuala) fuscocincta (Stal) 
Leptodictya {Hanuala) laidis Drake and 

Hambleton 
Leptodictya {Hanuala) litigiosa Monte 
Leptodictya {Hanuala) lucida Drake and 

Hambleton 
Leptodictya {Hanuala) nigrosis Drake 

and Hambleton 
Leptodictya ochropa (Stal) 
Leptodictya {Hanuala) parilis Drake and 

Hambleton 
Leptodictya {Hanuala) simulans Heide- 
mann 
Leptodictya {Hanuala) tabida (Herrich- 
Schaeffer) 
Bean 

Gargaphia angulata Heidemann 
Bean, kidney 

Leptodictya {Hanuala) colombiana Drake 
Broom 
Dictyonota fuliginosa Costa 
Dictyonota strichnocera Fieber 



Cabbage 

Corythaica cyathicollis (Costa) 
Cacao 

Amblystira machalana Drake 

Ammianus alberti (Schouteden) 

Dichocysta pictipes Champion 

Habrochila placida Horvath 

Phatnoma marmorata Champion 
Camphor 

Stephanitis typica (Distant) 
Chaparral 

Corythucha fuscigera (S t al) 
Cherry 

Corythucha cerasi Drake 

Phymacysta tumida (Champion) 
Coconut 

Stephanitis typica (Distant) 
Coffee 

Habrochila ghesquierei Schouteden 

Habrochila placida Horvath 

Leptopharsa flava Monte 
Cotton 

Dictyla monotropidia (Stal) 
Cottonwood 

Physatocheila variegata Parshley 
Ebony 

Corythucha morrilli Osborn and Drake 

Teleonemia scrupulosa Stal 
Elm 

Tingis {Tropidocheila) gamboana Drake 
and Hambleton 
Grapefruit 

Corythucha gossypii (Fabricius) 
Grass 

Leptodictya {Hanuala) madelinae Drake 
Hazelnut 

Corythucha padi Drake 
Heath 

Agramma laetum (Fallen) 
Heliotrope 

Cochlochila bullita (Stal) 

479 



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U.S. NATIONAL MUSEUM BULLETIN 243 



Hickory 
Physatocheila plexa (Say) 
Physatocheila variegata Parshley 
Hollyhock 
Corythucha distincta Osborn and Drake 
Gargaphia iridescens Champion 
Gargaphia solani Heidemann 
Sphaerocysta globifera (Stal) 
Hops 

Acalypta barberi Drake 
Ivy 

Derephysia foliacea (Fallen) 
Jasmine 

Cochlochila exolenta Drake 
Lemon 

Corythucha gossypii (Fabricius) 
Loquat 

Corythucha cydoniae (Fitch) 
Maize 
Gargaphia torresi Costa Lima 
Leptodictya {Hanuala) bambusae Drake 
Leptodictya {Hanuala) tabida (Herrich- 
Schaeffer) 
Mango 

Corythucha gossypii (Fabricius) 
Mangle 

Leptoypha morrisoni Drake 
Maple 

Corythucha arcuata (Say) 
Millet 

Kapiriella polita (Drake) 
Moss 
Acalypta barberi Drake 
Acalypta brunnea (Germar) 
Acalypta carinata (Panzer) 
Acalypta cooleyi Drake 
Acalypta duryi Drake 
Acalypta gracilis (Fieber) 
Acalypta lillianis Torre-Bueno 
Acalypta marginata (Wolff) 
Acalypta mniophila Drake and Ruhoff 
Acalypta musci (Schrank) 
Acalypta musci var. ditata (Puton) 
Acalypta nigrina (Fallen) 
Acalypta nyctalis Drake 
Acalypta parvula (Fallen) 
Acalypta platycheila (Fieber) 
Acalypta pulchr a StuSak 



Acalypta saundersi (Downes) 
Acalypta thomsonii Stal 
Agramma laetum (Fallen) 
Agramma ruficorne (Germar) 
Campylosteira ciliata Fieber 
Campylosteira falleni Fieber 
Campylosteira verna (Fallen) 
Derephysia foliacea (Fallen) 
Dictyonota strichnocera Fieber 
Esocampylia hackeri Drake 
Gonycentrum tindalei (Hacker) 
Lasiacantha capucina (Germar) 
Nettle, sand 

Gargaphia iridescens Champion 
Oak 
Corythucha cerasi Drake 
Corythucha floridana Heidemann 
Melanorhopala clavata (Stal) 
Physatocheila plexa (Say) 
Stephanitis querca Bergroth 
Orange 

Corythucha gossypii (Fabricius) 
Papaya 

Aconchus urbanus (Horvath) 
Pea 

Gargaphia subpilosa Berg 
Pear 

Corythucha cydoniae (Fitch) 
Pepper 
Diconocoris greeni (Kirby) 
Diconocoris hewetti (Distant) 
Nesocypselas piperica Drake 
Pine 

Physatocheila variegata Parshley 
Pineapple 
Phatnoma marmorata Champion 
Phatnoma trinidadana Drake 
Polony 

Leptopharsa milleri Drake 
Poplar 

Corythucha salicata Gibson 
Rose 

Corythucha arcuata (Say) 
Sedge 

Cyperobia carectorum Bergroth 
Snapdragon 

Teleonemia nigrina Champion 



LACEBUGS OF THE WORLD 



481 



Soursop 

Corythucha gossypii (Fabricius) 
Sugarcane 

Abdastartus sacchari Drake 

Leptodictya (Hanuala) tabida (Herrich- 
Schaeffer) 

Teleonemia sacchari (Fabricius) 
Sugar beets 

Teleonemia nigrina Champion 
Sunflower 

Galeatus scrophicus Saunders 
Sweet potato 

Gargaphia torresi Costa Lima 
Sycamore 

Corythucha salicata Gibson 
Tea 

Habrochila chinensis Drake 

Stephanitis {Norba) chinensis Drake 



Teak 

Amblystira machalana Drake 

Pontanus puerilis (Drake and Poor) 
Thistle 

Gargaphia bimaculata Parshley 
Tobacco 

Corythaica cyathicollis (Costa) 

Malala bulliens Distant 
Tomato 

Corythaica monacha (Stal) 
Walnut 

Corythucha padi Drake 
Willow 

Gargaphia opacula Uhler 

Physatocheila plexa (Say) 
Yautia 

Corythucha gossypii (Fabricius) 



Literature Cited 

The authors have seen the original publication of every title and reference 
listed in the bibliography, except two and these are so indicated. As cus- 
tomarily cited over the years, the dates of publications for a number of im- 
portant papers are not always in agreement with the actual date of issue. 
For example, certain works of Herrich-Schaeffer, Costa, Stal, and Blanchard, 
as cited in the literature, do not at all times bear the same date or even the 
correct date of publication. The authority for and reference to the litera- 
ture of such changes are cited [in brackets] just beneath the publication 
concerned. 

Abbott, Cyril E. 

1935. Notes on the oviposition and hatching of Corythucha marmorata 

Uhler. Bull. Brooklyn Ent. Soc, vol. 30, No. 1, p. 13. 
Alfieri, A. : see Priesner and Alfieri 
Amyot, Charles Jean Baptiste 

1845-1847. Entomologie franchise. Rhynchotes. Methode monony- 
mique. Ann. Soc. Ent. France, ser. 2 (1845, vol. 3, pp. 369— 
492; 1846, vol. 4, pp. 73-192, 359-452; 1847, vol. 5, pp. 143- 
238, 453-524). 
Amyot, Charles Jean B., and Ser ville, Jean Guillaumi Audinet 

1843. Histoire naturelle des insectes. Hemipteres. Paris, Ixvii 
+675 pp. Tingitides, pp. 295-303. 
Andre, E. 

1876. Euricera clavicornis-Canthophorus maculipes. Feuille des Jeunes 

Naturalistes, No. 73, pp. 34-35. 
1878. Voyage d'un naturaliste. Deux kilometers en six heures. 
Feuille des Jeunes Naturalistes, No. 87, pp. 25-27. 
d'Antessanty, Gabriel 

1890. Catologue des hemipteres-heteropteres de l'Aube. Mem. 
Soc. Acad. l'Aube, vol. 54, pp. 1-53. 
d'Araujo e Silva, Aristoteles G. 

1936. Alguns insectos com os seus respectivos hospedeiros. Rio de 

Janeiro, 30 pp. Tingidae p. 7. 

DE ArAUJO MENDONgA, FLORIANO 

1949. Notas sobre Gargaphia lanulata (Mayer, 1865) {Gargaphia 
lasciva Gibson). Boletim da Secretaria da Agricultura, 
Industria e Comercio do Estado da Bahia, Servico de Divul- 
gagao, vol. 45, No. 3, pp. 183-187, 15 figs. 
482 



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Ascot, H. W. 

1930. Rhododendron bug (Leptobyrsa rhododendri) . The Gardeners' 
Chronicle, Sept. 20, 1930, p. 238. 
Ash, Charles R. 

1954. Tingoidea (Hemiptera) of Tippecanoe County, Indiana. Proc. 
Indiana Acad. Sci., vol. 63, p. 185, 1953. 
Ashmead, William H. 

1886. On two new Hemiptera-Heteroptera. Canadian Ent.,vol. 18, 

No. 1, pp. 18-20. 

1887. Hemipterological contributions (No. 1). Ent. Amer., vol. 3, 

pp. 155-156. 
Bab a, Shosuke 

1925. "Dosokwai-Jiho," Nagano Agric. Exp. Station, vol. viii, No. 1, 

p. 3. [Publication not seen.] 
Bachofen-echt, Adolf 

1949. Der Bernstein und seine Einschlusse. Springer- Verlag. Vienna. 

204 pp., 188 figs. Tingidae fig. 160. 

von Baerensprung, Felix 

1858. Neue und seltene Rhynchoten der europaischen Fauna. Ber- 
liner Ent. Zeitschr., vol. 2, No. 2, pp. 188-208, 1 pi. 

Bailey, Norman S. [see also Feldman and Bailey] 

1950. An Asiatic tingid new to North America (Heteroptera) . Psyche, 

vol. 57, No. 4, pp. 143-145. 

1951. The Tingoidea of New England and their biology. Ent. Amer., 

vol. 31, pp. 1-140, 6 figs. 
1959. Additions to the bioecology of the New England Tingidae and 
Piesmidae (Heteroptera). Psyche, vol. 66, No. 4, pp. 63-69. 
Baker, Carl F.: see Gillette and Baker 
Balachowsky, Alfred, and Mesnil, L. 

1935-1936. Les Insectes nuisibles aux plantes cultivees leurs moeurs 
leur destruction, vol. 1 [1935], 1137 pp., 931 figs.; vol. 2 
[1936], pp. 1138-1921, figs. 932-1369. 
Banks, Nathan 

1909. Dates of Guerin's Iconographie regne animal. In Notes and 

News, Ent. News, vol. 20, No. 9, pp. 396-397. 

1910. Catalogue of the Nearctic Hemiptera-Heteroptera. Amer. 

Ent. Soc, Philadelphia, 103 pp. Tingitidae pp. 55-57. 
Barber, Harry Gardner 

1906. Hemiptera from southwestern Texas. Brooklyn Inst. Arts and 

Sci. Bull. 1, No. 9, pp. 255-289. 
1910. Some Mexican Hemiptera-Heteroptera new to the fauna of 

the United States. Journ. New York Ent. Soc, vol. 18, No. 1, 

pp. 34-39. 



484 U.S. NATIONAL MUSEUM BULLETIN 243 

1914. Insects of Florida. 11. Hemiptera. Bull. Amer. Mus. Nat. Hist., 
vol. 33, art. 31, pp. 495-535. 

1922a. List of the superfamily Tingidoidea of New Jersey with synon- 
ymy and food plants. State of New Jersey Dept. Agric., Bur. 
Statistics and Inspection, circ. 54, pp. 16-17. 

1922b. Systematic treatment of the superfamily Tingidoideae of New 
Jersey. State of New Jersey Dept. Agric, Bur. Statistics 
and Inspection, circ. 54, pp. 18-24. 

1923. A preliminary report on the Hemiptera-Heteroptera of Porto 
Rico collected by the American Museum of Natural History. 
Amer. Mus. Nov., No. 75, pp. 1-13. 

1925. Hemiptera-Heteroptera from the Williams Galapagos Expe- 
dition. Zoologica, vol. 5, No. 21, pp. 241-254, 4 figs. 

1934. The Norwegian Zoological Expedition to the Galapagos 
Islands 1925, conducted by Alf Wollebaek. XI. Hemiptera- 
Heteroptera. Meddelelser fra det Zoologiske Mus., Oslo, No. 
42, pp. 281-289. 

1939. Insects of Porto Rico and the Virgin Islands — Hemiptera- 

Heteroptera (excepting the Miridae and Coxidae). 
Scientific Survey of Porto Rico and the Virgin Islands. 
New York Acad. Sci., vol. 14, No. 3, pp. 263-441, 36 figs. 
1954. A report on the Hemiptera-Heteroptera from the Bimini Islands, 
Bahamas, British West Indies. Amer. Mus. Nov., No. 1682, 
pp. 1-18, 3 figs. 
Barber, Harry G., and Weiss, Harry B. 

1922. The lace bugs of New Jersey. State of New Jersey Dept. Agric, 
Bur. Statistics and Inspection, circ. 54, pp. 3-15, 8 figs. 
Bator, Adelheid 

1953. Die heteropteren Nordtirols. Beitrage zur Entomologie, vol. 
3, No. 3, pp. 323-333, 3 pis. 
Beeson, Cyril Frederick, and Chatterjee, N. G. 

1940. Possibilities of control of lantana {Lantana aculeata Linn.) by 

indigenous insect pests. Indian Forest Records, new ser., vol. 
6, No. 3, pp. 41-84, 1939. 

Behr, L. 

1952. Uber die Bliitengalle des Teucrium chamaedrys L. Berichte der 

Deutschen Botanishchen Gesellschaft, vol. 65, No. 9, pp. 
326-330, 6 figs. 
Bekker-Migdisova, E. E. 

1953. Paleozoiskie ravnokrylyie voprosy filogenii otryada. Do- 

klady Akad. Nauk S.S.S.R., Moscow, new ser., vol. 90, pp. 
461-464, 2 figs. 



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Berendt, Georg Carl [see also Germar and Berendt] 

1845. Der Bernstein und die in ihm befindlichen Pflanzenreste der 

Vorwelt. In Die im Bernstein befindlichen Organischen reste 

der Vorwelt, vol. 1, No. 1, pp. 1-60. 
Berg, Carlos 

1879a. Hemiptera Argentina. Anal. Soc. Cient. Argentina, vol. 7, 

pp. 41-47. [Section that included Tingitidae; this publica- 
tion was issued in parts then reprinted as an entire volume.] 
1879b. Hemiptera Argentina enumeravit speciesque novas descripsit. 

Bonariae, Hamburg. 316 pp. [Reprint.] Tingitidae pp. 

135-137. 
1884. Addenda et emendanda ad Hemiptera Argentina. Anal. Soc. 

Cient. Argentina, 213 pp. Tingitidae pp. 99-104. 
1891-1892. Nova Hemiptera faunarum Argentinae et Uruguayensis. 

Anal. Soc. Cient. Argentina, vol. 32, No. 4, pp. 164-175; No. 

5, pp. 231-243; No. 6, pp. 276-287; [1892] vol. 33, No. 1, 

pp. 1-11; No. 2, pp. 43-50; No. 3, pp. 65-72; No. 4, pp. 

98-104; No. 5, pp. 151-165; vol. 34, No. 2, pp. 82-96; No. 

5, pp. 193-205. [Reprint 1892, 112 pp. Tingitidae pp. 

99-102.] 
Bergevtn, Ernest 

1920. Description d'une espece nouvelle de Physatocheila d'Algerie 

[Hemip. Tingitidae]. Ann. Soc. Ent. France, vol. 88, pp. 

412-413, 1 fig. 
1922. Description d'une nouvelle espece de Catoplatus (Hemiptere 

Tingitidae) du Maroc Occidental. Bull. Soc. Sci. Nat. 

Maroc, vol. 2, Nos. 5-6, pp. 108-109, 2 figs. 

1928. Description d'une nouvelle espece d'Urentius (Hemiptere 

Tingitidae) du Hoggar. Bull. Soc. Hist. Nat. Afrique du 
Nord, vol. 19, pp. 336-338, 1 fig. 

1929. Description de deux especes nouvelles d'hemipteres Tingitidae 

provenant de la mission du Hoggar. Bull. Soc. d'Hist. Nat. 
Afrique du Nord, vol. 20, pp. 114-116, 2 figs. 

1930. Note sur trois especes d'hemipteres recueillis en Egypte et 

description d'une nouvelle espece d'Urentius (Hemiptere 
Tingitidae) et d'une nouvelle variete d'Ommatissus binotatus 
Fieb. (Homoptere Cixiidae). Bull. Soc. Roy. Ent. Egypte, 
new ser., fasc. 1, pp. 17-20, 1 fig. 
Bergroth, Ernst Evald 

1892. Notes synonymiques. Rev. Ent, vol. 11, pp. 262-264. 

1894. TingidaetresMadagascariensis. Rev. Ent., vol. 13, pp. 167-168. 

1898. Eine neue Tingide. Wiener Ent. Zeitung, vol. 17, No. 1, p. 9. 

1903. Rhynchota Aethiopica. Ann. Soc. Ent. Belgique, vol. 47, 
pp. 287-297. 



486 U.S. NATIONAL MUSEUM BULLETIN 243 

1908. Neue Hemiptera aus Siid-Abyssinien. Rev. Russe Ent., Nos. 
2-3, pp. 106-110, 1907. 

1911. On some recently described Hemiptera, chiefly from India. 

Ann. Soc. Ent. Belgique, vol. 55, pp. 184-189. 

1912. Tingidae duae novae Africanae. Ann. Soc. Ent. Belgique, vol. 

56, pp. 145-146. 
1914a. H. Sauter's Formosa-Ausbeute: Hemiptera Heteroptera I. 

Ent. Mitt., vol. 3, Nos. 10-12, pp. 353-364. 
1914b. Zwei neue palaarktische hemipteren, nebst synonymischen 

Mitteilungen. Wiener Ent. Zeitung, vol. 33, Nos. 5-6, pp. 

177-184. 
1919. Die Erscheinungsdata zweier hemipterologischen Werke. Ent. 

Mitt., vol. 8, Nos. 10-12, pp. 188-191. [Contains validation 

of publication dates of papers by Herrich-Schaeffer.] 

1921. On the types of the exotic Hemiptera Heteroptera described 

by V. Motschulsky. Rev. Russe Ent., vol. 17, pp. 96-109, 
1917. 

1922. On some Neotropical Tingidae (Hem.). Ann. Soc. Ent. 

Belgique, vol. 62, pp. 149-152. 
1924. Some Heteroptera from the alpine region of central Luzon. 

Ann. Soc. Ent. Belgique, vol. 64, pp. 82-84. 
1927. Hemiptera Heteroptera from New Zealand. Trans. New 
Zealand Inst., vol. 57, pp. 671-684. 
Bergroth, Ernst, and Schouteden, Henri 

1905. Note sur les hemipteres recueillis a Kinchassa par M. Wael- 
broeck. Ann. Soc. Ent. Belgique, vol. 49. pp. 379-389, 1 fig. 
Beri, Y. P. : see Menon, Beri, and Singh 
Berry, Paul A. 

1959. Entomologia Economica de El Salvador. Servicio Cooperativo 
Agricola Salvadoreno Americano Ministerio de Agricultura y 
Ganaderia Santa Tecla, El Salvador. Boletin Tecnico 24, 
255 pp., 187 figs. 

BlANCHI, F. A. 

1955a. Corythucha morrilli. In Minutes of the meeting, April 12, 1954, 

Hawaiian Entomological Society. Proc. Hawaiian Ent. Soc, 

vol. 15, No. 3, p. 378. 
1955b. Corythucha morrilli. In Minutes of the meeting, August 9, 1954, 

Hawaiian Entomological Society. Proc. Hawaiian Ent. Soc, 

vol. 15, No. 3, p. 383. 
Blanchard, Emile 

1840. Histoire naturelle des animaux articules, annelides, crustaces, 

arachnides, myriapodes et insectes., vol. 3, 672 pp., many 

plates. Tingis, pp. 112-113. 



LACEBUGS OF THE WORLD 487 

1846. Insectes du voyage dans l'Amerique Meridionale par Alcide 
d'Orbigny. Paris, 1 837-1 846, 222 pp., 32 pis. [See Sherborn 
and Woodward 1901, for the dates of this publication.] 
Blanchard, Everard 

1926. Sobre un Tingido nuevo para la fauna Argentina. Physis, 
vol. 8, pp. 361-363, 1 fig. 
Blatchley, Willis Stanley 

1926. Heteroptera or true bugs of eastern North America, with 
especial references to the faunas of Indiana and Florida. 
Nature Publ. Co., Indianapolis, 1116 pp., 215 text figs., 
12 pis. Tingididae pp. 448-501, figs. 96-121. 
1928. Notes on the Heteroptera of eastern North America with 
descriptions of new species, I. Journ. New York Ent. Soc, 
vol. 36, No. 1, pp. 1-23. 
Blote, Hendrick Coenaad 

1945. Catalogue of the Berytidae, Piesmidae and Tingidae in the 
Rijksmuseum van Natuurlijke Historic Zoologische Me- 
dedeelingen, vol. 25, pp. 72-92, 1 fig. 
Bolivar, Ignacio, and Chicote, Cesar 

1879. Enumeration de los hemipteros observados en Espana y 
Portugal. Ann. Soc. Espafiola Hist. Nat., vol. 8, pp. 147-186. 
Bondar, Gregorio [see also Drake and Bondar] 

1928. Amarellecimento das folhas do feijao causado pelo Tingitideo. 
Correio Agricola, vol. 6, Nos. 11-12, pp. 244-247, 3 figs. 

1936. Tingitideo nocivo as Anonaceas frutiferas no Brasil. O Campo, 

January, p. 51, 2 figs. 
Borner, Carl 

1935. Ordnung. Schnabelkerfe, Rhynchota (Hemiptera). In Die 

Tierwelt Mitteleuropas von P. Brohmer, P. Ehrmann, G. 

Ulmer, band 4, teil 1, insekten lief 3, pp. 1-113, 181 figs. 
Boso_, Juan M. 

1937. Lista preliminar de los hemipteros (heteropteros), especial- 

men te relacionados con la agricultura nacional. Revista 
Soc. Ent. Argentina, vol. 9, pp. 111-134. 
Box, Harold E. 

1953. List of sugar-cane insects. Commonwealth Institute of Ento- 
mology, 101 pp. 
Breddin, Gustav 

1901. Neue neotropische Wanzen und Zirpen. Soc. Ent., vol. 16, 
No. 11, pp. 82-84. 

1907. Rhynchotographische Beitrage. Wiener Ent. Zeitung, vol. 26, 
pp. 93-97. 



488 U.S. NATIONAL MUSEUM BULLETIN 243 

Bremond, P. 

1938. Le Faux-Tigre des arbres fruitiers {Monostira unicostata Muls.) 
au Maroc. Rev. Pathologie Vegetale et d'Entomologie 
Agricole de France, vol. 25, No. 4, pp. 294-307, 3 pis., 2 figs. 
Britton, Wilton Everett 

1920. Check-list of the insects of Connecticut. State Geol. Nat. 
Hist. Surv., Connecticut, Bull. 31, 397 pp. Tingidae p. 77. 
Broek, M. van den, and Schenk, P. J. 

1915. Dierlijke en Plantaardige Parasieten. Vol. 1. Ziekten en 
Beschadigingen der Tuinbouwgewassen ten dienste van 
tuinbouwscholen en -cursussen en de practijk. Groningen, 
J. B. Wolters. 382 pp. 
Bruel, W. E. van den 

1947. A propos du comportement en Belgique de Stephanitis rhodo- 
dendri Horv. (Tingidae). Bull. Ann. Soc. Ent. Belgique, 
vol. 84, pp. 191-197, 1 fig. 
Brulle, Auguste 

1835. Orthopteres et hemipteres. In Histoire naturelle des in- 
sectes (Audouin et Brulle) leur classification et la description 
des especes par M. A. Brulle. Paris. Vol. 4, 415 pp., pis. 
Bruner, Stephen C. [see also Drake and Bruner] 

1940. A new tingitid from Cuba (Hemiptera). Mem. Soc. Cubana 
Hist. Nat., vol. 14, No. 3, pp. 245-247, 1 fig. 
Bruner, Stephen; Scaramuzza, L. C; and Otero, A. R. 

1945. Catalogo de los insectos que atacan a las plantas economicas 
de Cuba. Estacion Experimental Agronomica Santiago de 
las Vegas, Provincia de la Habana. Bol. 63, pp. 1-246, 12 pis. 
Buchanan- White, Francis 

1877. Laccometopus clavicornis L. and its relation to Teucrium chamaedrys. 

Entomologists' Monthly Mag., vol. 13, p. 283. 
1879. Description of new Hemiptera. (1). Linnean Soc. Journ. 
Zool., vol. 14, pp. 482-489. 
Bueno: see Torre -Bueno 

BURMEISTER, HERMANN 

1835. Handbuch der Entomologie. Berlin. Vol. 2, 400 pp. Mem- 
branacei pp. 249-262. 
Butler, Arthur Gardiner 

1877. Lepidoptera, Orthoptera, and Hemiptera. In Account of the 
zoological collection made during the visit of H.M.S. Peterel 
to the Galapagos Islands, by Albert Giinther. Proc. Scien- 
tific Meetings Zool. Soc. London, pp. 86-91. 



LACEBUGS OF THE WORLD 489 

Butler, Edward A. 

1922. A contribution towards the life-history of Dictyonota strichnocera 

Fieb. Entomologists' Monthly Mag., vol. 58, No. 92, pp. 
179-182. 

1923. A biology of the British Hemiptera-Heteroptera. London. 

682 pp., many figs. Tingidae pp. 196-216, 3 figs. 
Byers, George W. 

1959. Types of Tingidae described by Torre-Bueno (Hemiptera). 

Ent. News, vol. 70, No. 7, pp. 191-193. 
Campbell, T. G.: see Cashmore and Campbell 
Carayon, Jacques [see also Monod and Carayon] 

1960. Stethocomis Jrappai n. sp. Miride predateur du Tingide du 

cafeier, Dulinius unicolor (Sign.), a Madagascar. Journ. 
d' Agriculture Tropicale et de Botanique Appliquee, vol. 7, 
Nos. 1-3, pp. 110-120, 5 figs. 
Carrillo, Jose Luie: see Gibson and Carrillo 
Carvalho, Jose C: see Drake and Carvalho 
Cashmore, A. B., and Campbell, T. G. 

1946. The weeds problem in Australia: A review. Journ. Council 
for Scientific and Industrial Research., vol. 19, No. 1, 
pp. 16-31, 2 figs. 
Casdenau, Francois Louis Nompar de Caumont de Laporte, Comte de: 

see Laporte 
Cederhielm, Johann 

1798. Faunae ingricae prodromus exhibens methoicam descriptionem 
insectorum agri petropolensis praemissa mammalium, avium, 
amphibiorum et piscium enumeratione. 348 pp., 3 pis. 
Ryngota pp. 263-286. 
Champion, George Charles 

1897. Rhynchota. In Godman and Salvin, Biologia Centrali- 

Americana, vol. 2, pp. 1-32. 
1898a. Rhynchota. In Godman and Salvin, Biologia Centrali- 
Americana, vol. 2, pp. 33-193, 22 pis. Tingitidae pp. 33-48. 
1898b. Notes on American and other Tingitidae, with descriptions of 
two new genera and four new species. Trans. Ent. Soc. 
London, vol. 1, pp. 55-64, 2 pis. 

1924. The insects of Galapagos Islands. Entomologists' Monthly 

Mag., vol. 60, pp. 259-260. 
Channa Basavanna, G. P.: see Thontadarya and Channa Basavanna 
Chatterjee, N. C: see Beeson and Chatterjee 
Chicote, Cesar [see also Bolivar and Chicote] 

1880. Adiciones a la enumeration de los hemipteros observados en 

Espafia y Portugal. Anal. Soc. Espanola Hist. Nat., vol. 9, 

pp. 185-203. 



490 U.S. NATIONAL MUSEUM BULLETIN 243 

China, William E. 

1924. The Hemiptera-Heteroptera of Rodriguez, together with the 
description of a new species of Cicada from that island. 
Ann. Mag. Nat. Hist., ser. 9, vol. 14, pp. 427-453, 4 figs. 

1926. Synonymic notes on Hemiptera. Entomologist, vol. 59, pp. 
227-228. 

1930. Heteroptera. In Insects of Samoa and other Samoan Terres- 
trial Artropoda, part II, Hemiptera, fasc. 3, pp. 81-162, 28 
figs. British Museum (Natural History). 

1936. A new species of the genus Physatocheila Fieb. (Heteroptera: 
Tingitidae) from Dorsetshire. Entomologists' Monthly Mag., 
vol. 72, pp. 270-272, 1 fig. 

1938. Die Arthropod enfauna von Madeira nach den Ergebnissen der 
Reise von Prof. Dr. O. Lundblad Juli-August 1935. 3. 
Terrestrial Hemiptera. Arkiv for Zoologi, Svenska Veten- 
skapsakademien, vol. 30, No. 2, pp. 1-68, 23 text figs.; 1 pi. 

1941. Genotype fixations in Hemiptera Heteroptera. Proc. Roy. 
Ent. Soc. London (B), vol. 10, part 7, p. 130. 

1943. The generic names of the British Hemiptera-Heteroptera with 
a check list of the British species. In The generic names of 
British insects prepared by the Committee on Generic Nomen- 
clature of Royal Entomological Society of London, part 8, 
pp. 211-342. 

1945. A completely blind bug of the family Lygaeidae (Hemiptera- 
Heteroptera). Proc. Roy. Ent. Soc. London, vol. 14, parts 
9-10, pp. 126-128, 1 fig. 

1952. On the identity of Physatocheila quadrimaculata Wolff (Heter- 
optera: Tingidae). Entomologists' Monthly Mag., vol. 88, 
pp. 49-50, 1 fig. 
China, William, and Miller, N.C.E. 

1955. Check-list of family and subfamily names in Hemiptera- 
Heteroptera. Ann. Mag. Nat. Hist., ser. 12, vol. 8, pp. 
257-267. 

1959. Check-list and keys to the families and subfamilies of the 
Hemiptera-Heteroptera. Bull. British Museum (Nat. Hist.), 
Entomology, vol. 8, No. 1, pp. 1-45. 
Chittenden, Frank H. 

1900. A new tingitid on bean. In Some insects injurious to garden 
crops, a series of articles dealing with insects of this class. 
U.S. Dept. Agric, Bull. 23, new ser., pp. 32-33, 1 fig. 



LACEBUGS OF THE WORLD 491 

Cobben, Rene 1 [see also Drake and Cobben] 

1948. Wantsennieuws uit midden Limburg. Natuurhistorisch 

Maandblad, vol. 37, Nos. 11-12, pp. 81-82, 1 fig. 
1958a. Biotaxonomische Einzelheiten iiber Niederlandische Wanzen 

(Hemiptera-Heteroptera). Tijdschrift voor Entomologie, 

deel 101, No. 1, pp. 1-46, 40 figs. 
1958b. Einige Bemerkenswerten Wanzenfunde aus den Niederlanden 

(Hem.-Heteroptera). Natuurhistorisch Maandblad, vol. 47, 

pp. 15-21. 

COCKERELL, THEODORE D. A. 

1893. The entomology of the mid-Alpine zone of Custer County, 

Colorado. Trans. Amer. Ent. Soc., vol. 20, pp. 305-370. 
1914. New and little-known insects from the Miocene of Florissant, 

Colorado. Journ. Geology, vol. 22, No. 7, pp. 714-724, 

11 figs. 
1921. Fossil arthropods in the British Museum. Ann. Mag. Nat. 

Hist., ser. 9, vol. 8, pp. 541-545, 5 figs. 
Comstock, Anna Botsford: see Comstock and Comstock 
Comstock, John Henry 

1880. The hawthorn Tingis (Corythucha arcuata, Say, var.), order 

Hemiptera; family Tingidae. U.S. Dept. Agric, Rep. of 

the Entomologist for 1879, pp. 221-222. 
Comstock, John Henry, and Comstock, Anna Botsford 

1895. A manual for the study of insects. Comstock Publ. Co., Ithaca, 

N.Y. 701 pp. 
Costa, Achille 

1847a. Cimicum regni neapolitani centurae. Napoli, Centuria 1, 

76 pp., pi. 
1847b. Cimicum regni neapolitani centuria secunda. 43 pp., 2 pis. 
1847c.*Cimicum regni neapolitani centuria. Atti del Reale Istitute 

dTcoraggiamento alle Scienze di Napoli, vol. 7, pp. 143-216. 
1847d.*Cimicum regni neapolitani centuria secunda, decas prima, 

secunda, tertia, quatra, et quinta. Atti del Reale Istitute 

dTcoraggiamento alle Scienze di Napoli, vol. 7, pp. 239-279. 
1855a. Cimicum regni neapolitani centuria tertia et quartae frag- 

mentum. 77 pp. 
1855b.*Cimicum regni neapolitani. Centuria tertia et quartae frag- 

mentum. Atti del Reale Istitute d'Icoraggiamento alle 

Scienze Naturali di Napoli, vol. 8, pp. 225-300. 



*Cimicum regni neapolitani was originally published in various parts. 
Atti del Reale is a reprint of these using different pagination. See Stichel 
1960d for dates of these publications. 



492 U.S. NATIONAL MUSEUM BULLETIN 243 

1863. Additamenta as centurias cimicum regni neapolitani. 41 pp., 

3 pis. (color). 

1864. Annuario del Museo Zoologico della R. Universita di Napoli. 

Napoli, vol. 2, pp. 1-176, 4 pis. (color), 1862. 
de Costa Lima, Angelo M. 

1922. Nota sobre os insectos que atacam o algodoeiro no Brazil. 

Chacaras e Quintaes, vol. 25, No. 2, pp. 110-112, 6 figs. 
1930. Supplemento ao 2 Catalogo systematico dos insectos que 
vivem nas plantas do Brasil e Ensaio de Bibliographia Ento- 
mologica Brasileira. O Campo, vol. 1, No. 8, pp. 84-91. 
1936. Terceiro catalogo dos insectos que vivem nas plantas do Brasil. 
Rio de Janeiro. 460 pp. Tingidae pp. 124-131. 
Cotton, Richard T. 

1917. The eggplant lace-bug in Porto Rico. Journ. Dept. Agric. 
Porto Rico, vol. 1, No. 3, pp. 170-173. 
Crosby, Cyrus R., and Hadley, C. H. 

1915. The rhododendron lace-bug, Leptobyrsa explanta Heidemann. 
Journ. Econ. Ent., vol. 8, No. 4, pp. 409-414, 2 pis. 
Currie, G. A., and Fyfe, R. V. 

1939. The lantana bug in Australia. Progress report. Journ. 
Council for Scientific and Industrial Research, vol. 12, 
No. 3, pp. 259-263. 
Curtis, John 

1823-1840. British entomology; being illustrations and descriptions 
of the genera of insects found in Great Britain and Ireland. 
London. 16 vols. (193 nos.), 770 pis. (color). 
1833. Characters of some undescribed genera and species, indicated 
in the "Guide to an arrangement of British insects." Ent. 
Mag., vol. 1, pp. 186-199. 
d'Antessanty: see Antessanty 

d'Araujo e Silva: see Araujo e Silva, Aristoteles G. 
Davis, Norman T. : see Drake and Davis 
de Araujo Mendonca: see Araujo Mendonqa, Floriano 
de Costa Lima: see Costa Lima 
De Geer, Charles 

1773. Memoires pour servir a Phistoire des insectes. Vol. 3, Stock- 
holm. 696 pp., 43 pis. 
de Seabra: see Seabra 
de Serres: see Serres 
de Walsch: see Walsch 

Dickerson, Edgar L., 

1917. Notes on Leptobyrsa rhododendri Horv. Journ. New York Ent. 
Soc, vol. 25, No. 2, pp. 105-112, 1 pi. 



LACEBUGS OF THE WORLD 



493 



1917. 



1918. 



Dickerson, Edgar L., and Weiss, Harry B. [see also Weiss and Dickerson] 
1916. Notes on Leptoypha mutica Say (Hemip.). Ent. News, vol. 27 

pp. 308-310, 1 pi. 
The azalea lace-bug, Stephanitis pyrioides Scott (Tingitidae, 

Hemiptera). Ent. News, vol. 28, pp. 101-105, pi. 1. 
Corythncha spinulosa Gibson, a new lace-bug on wild cherry 

(Hem. Horn.). Ent. News, vol. 29, pp. 121-125, 1 pi. 
Distant, William Lucas 

1888. Enumeration of the van Volxem collection of Rhynchota in 

the Brussels Museum, part 3. Ann. Soc. Ent. Belgique, 

vol. 32, pp. lxxviii-lxxxiii. 
1902a. XII. Rhynchotal miscellanea. Ann. South African Mus., 

vol. 2, No. 9, pp. 237-254, 1 pi. 
1902b. Rhynchotal notes. XIII. Heteroptera: Families Tingididae, 

Phymatidae and Aradidae. Ann. Mag. Nat. Hist., ser. 7, 

vol. 9, pp. 353-362. 
1903a. Contributions to a knowledge of the Rhynchota. Ann. Soc. 

Ent. Belgique, vol. 47, pp. 43-65. 
1903b. The fauna of British India including Ceylon and Burma, 

Rhynchota, vol. II (Heteroptera). Taylor and Francis, 

London. [1903 pp. 1-242; 1904 pp. 243-503]. Tingitidae 

pp. 122-145. 
1903c. Report on the Rhynchota. Part I, Heteroptera. Fasc. of 

Malayan Zoology, vol. 1, No. 2, pp. 219-273, 2 pis. 
1904. On the South African Tingididae and other heteropterous 

Rhynchota. Trans. South African Philos. Soc, vol. 14, No. 

5, pp. 425-436, 1 pi. 
1907. Description of a new species of Tingididae from Honolulu. 

Etomologist, vol. 40, pp. 60-61. 
1908a. Description of a new tingid from Congo. Ann. Soc. Ent. 

Belgique, vol. 52, p. 220. 
1908b. XV. Rhynchota Malayana, part 1. Rec. Indian Mus., vol. 2, 

part 2, No. 15, pp. 127-151, 2 pis. 
1909a. New Oriental Tingididae. Ann. Soc. Ent. Belgique, vol. 53, 

pp. 113-123. 
1909b. Rhynchota (Heteroptera) from British India. Ann. Soc. 

Ent. Belgique, vol. 53, pp. 360-376. 
1909c. XL Rhynchota Malayana, part 2. Rec. Indian Mus., vol. 3, 

part 2, No. 2, pp. 163-181, 2 pis. 
1910a. The fauna of British India, including Ceylon and Burma. 

Rhynchota, vol. V (Heteroptera: Appendix). Taylor and 

Francis, London. 362 pp., 214 figs. Tingidae pp. 100-126. 
1910b. Rhynchota Philippinensia, part 1. Philippine Journ. Sci., 

vol. 5, No. 1, pp. 57-66, 1 pi. 

681-552—64 33 



494 U.S. NATIONAL MUSEUM BULLETIN 243 

1910c. An introduced pest to rhododendrons. Zoologist, ser. 4, 

vol. 14, No. 166, pp. 395-396, 2 figs. 
1911a. On some South African Rhynchota in the South African Mu- 
seum. Ann. South African Mus., vol. 10, No. 2, pp. 39-49, 

9 figs. 
1911b. Rhynchota Indica (Heteroptera). Entomologist, vol. 44, pp. 

269-271. 
1913. The Percy Sladen Trust Expedition to the Indian Ocean in 

1905, vol. 5, Rhynchota, part 1: suborder Heteroptera. 

Trans. Linnean Soc. London, vol. 16, No. 2, pp. 139-191, 

3 pis. 
1920. Rhynchota from New Caledonia. Ann. Mag. Nat. Hist., ser. 9, 

vol. 6, pp. 143-164. 
Dobs'ik, B. 

1951. Pfispevek k poznani pIostic Csr. [Contribution a la connais- 

sance des he terop teres de Csr. IV]. Entomologicke Listy 

(Folia entomologica), vol. 13, Nos. 1-4, pp. 36-40, 1950. 
Douglas, John W. 

1877. The economy of Laccometopus clavicornis, Lin. Entomologists' 

Monthly Mag., vol. 13, pp. 236-237. 
Douglas, John William, and Scott, John 

1863. Descriptions of two new species. Entomologist's Ann. for 1863. 

London, pp. 143-146, 2 figs. 
1865. The British Hemiptera. Vol. 1, Hemiptera-Heteroptera, 627 

pp., 21 pis. Tingitidae pp. 243-264. 

1868. British Hemiptera: additions and corrections. Entomologists' 

Monthly Mag., vol. 4, pp. 238-246, 1867-1868. 

1869. British Hemiptera: additions and corrections. Entomologists' 

Monthly Mag., vol. 5, pp. 259-268, 1868-1869. 
Downes, W. 

1925. Lace bugs of British Columbia. Proc. Ent. Soc. British Colum- 
bia, No. 22, pp. 12-19, 1 pi. 
1927a. A preliminary list of the Heteroptera and Homoptera of British 

Columbia. Proc. Ent. Soc. British Columbia, No. 23, pp. 

1-22. 
1927b. A new species of Drakella (Heteroptera-Tingitidae). Canadian 

Ent., vol. 59, p. 60. 
Drake, Carl J. [see also Osborn and Drake] 

1916. A new tingid from Tennessee. Ohio Journ. Sci., vol. 16, No. 7, 

pp. 326-328, 1 fig. 
1917a. New and noteworthy Tingidae from the United States. Ohio 

Journ. Sci., vol. 17, No. 6, pp. 213-216. 
1917b. Key to the nearctic species of Gargaphia with the description of 

a new species (Hem.-Het.). Ent. News, vol. 28, pp. 227-228. 



LACEBUGS OF THE WORLD 495 

1917c. The North American species of Monanthia (Tingidae). Bull. 

Brooklyn Ent. Soc, vol. 12, No.3, pp. 49-52, 1 fig. 
1918a. Two new tingids from the West Indies (Hem.-Het.). Ohio 

Journ. Sci., vol. 18, No. 5, pp. 175-177. 
1918b. The North American species of Teleonemia occurring north of 

Mexico. Ohio Journ. Sci., vol. 18, No. 8, pp. 323-332. 
1918c. A new corn insect from California (Heteroptera). Journ. 

Econ. Ent., vol. 11, No. 4, p. 385. 
1918d. Notes on North American Tingidae (Hem.-Het.). Bull. 

Brooklyn Ent. Soc, vol. 13, pp. 86-88. 
1919a. On some North American Tingidae (Hemip.). Ohio Journ. 

Sci., vol. 19, No. 7, pp. 417-421. 
1919b. On some Tingidae new to the fauna of Canada (Hemip.). 

Canadian Ent., vol. 51, pp. 159-160. 
1919c. An undescribed Teleonemia from Florida and Jamaica (Hemip.). 

Florida Buggist, vol. 3, p. 24. 
1920. Descriptions of new North American Tingidae. Ohio Journ. 

Sci., vol. 20, No. 3, pp. 49-54. 
1921a. A new species oiPachycysta (Hem.-Het.). Bull. Mus. Hist. Nat., 

Paris., vol. 5, pp. 344-345, 1 fig. 
1921b. Notes on some American Tingidae, with descriptions of new 

species. Florida Ent., vol. 4, pp. 49-54, 1 pi. 
1922a. The genus Dicysta Champion (Hemiptera). Ann. Carnegie 

Mus., vol. 13, Nos. 3-4, pp. 269-273, 1 fig. 
1922b. On some North and South American Tingidae (Hemip.). 

Florida Ent., vol. 5, No. 3, pp. 37-43, 48-50, 1 fig. 
1922c. Neotropical Tingitidae with descriptions of three new genera 

and thirty-two new species and varieties (Hemiptera). Mem. 

Carnegie Mus., vol. 9, No. 2, pp. 351-378, 2 figs., 1 pi. 
1922d. Heteroptera in the vicinity of Cranberry Lake. Tech. publ. 

No. 16, New York State College of Forestry, Syracuse Univ., 

vol. 22, No. 5, pp. 54-86, figs. 22-36. 
1922e. Contribution toward the life history of Galeatus peckhami Ash- 
mead. Tech. publ. No. 16, New York State College of 

Forestry, Syracuse Univ., vol. 22, No. 5, pp. 105-110, 1 pi. 
1922f. The life history of the birch tingitid, Corythucha pallipes Parshley. 

Tech. publ. No. 16, New York State College of Forestry, 

Syracuse Univ., vol. 22, No. 5, pp. 111-116, 1 pi. 
1923a. Some Tingitidae from Japan (Hemip.). Ohio Journ. Sci., 

vol. 23, No. 2, pp. 102-106, 1 fig. 
1923b. Two new species of Cantacaderia (Hemip. -Tingitidae). Bull. 

Brooklyn Ent. Soc, vol. 18, No. 3, pp. 81-84, 1 fig. 
1924. A new species of Acanthocheila from Bolivia (Hemiptera-Tingi- 

tidae). Bull. Brooklyn Ent. Soc, vol. 19, No. 3, p. 94. 



496 U.S. NATIONAL MUSEUM BULLETIN 243 

1925a. An undescribed gall-making hemipteron (Tingitidae) from 

Africa. Amer. Mus. Nov., No. 158, pp. 1-2, 1 fig. 
1925b. A new genus and new species of Tingitidae from Madagascar 

(Homopt.). Ent. Mitt., vol. 14, No. 2, pp. 107-109, 1 fig. 
1 925c. Concerning some Tingitidae from the Gulf States (Heteroptera) . 

Florida Ent., vol. 9, pp. 36-39. 
1926a. Notes on some Tingitidae from Cuba (Hemiptera). Psyche, 

vol. 33, No. 3, pp. 86-88. 
1926b. The North American Tingitidae (Heteroptera) described by 

Stal. Ann. Carnegie Mus., vol. 16, Nos. 3-4, pp. 375-380, 

lpl. 
1926c. An undescribed tingitid from Arizona (Hemiptera). Bull. 

Brooklyn Ent. Soc, vol. 21, No. 3, pp. 126-127. 
1926d. The South American species of the genus Tingis Fabricius 

(Hemiptera). Ann. Carnegie Mus., vol. 17, No. 1, pp. 83- 

85. 
1926e. On some Tingitidae from Java (Heteroptera). Treubia, vol. 8, 

Nos. 3-4, pp. 334-335. 
1927a. Tingitidae from the Far East (Hemiptera). Philippine Journ. 

Sci., vol. 32, No. 1, pp. 53-59. 
1927b. Two undescribed tingitids from Mexico. Bull. Brooklyn Ent. 

Soc, vol. 22, No. 2, pp. 116-117. 
1927c. A new species of Diplocysta from the Philippines (Hemip. - 

Tingitidae). Pan-Pacific Ent., vol. 4, No. 1, p. 18. 
1927d. Tingitidae from the Far East and Madagascar (Hemiptera). 

Philippine Journ. Sci., vol. 34, No. 3, pp. 307-312. 
1928a. New and little known Neotropical Tingitidae. Iowa State 

Coll. Journ. Sci., vol. 3, No. 1, pp. 41-56. 
1928b. A list of the insects of New York, families Piesmidae and 

Tingitidae. Cornell Univ. Agric. Exp. Sta., Mem. 101, pp. 

99-103. 
1928c. A synopsis of the American species of Acalypta (Hemip. - 

Tingitidae). Bull. Brooklyn Ent. Soc, vol. 23, No. 1, pp. 

1-9, 2 figs. 
1928d. Synonymical notes on tingitid genera with the descriptions of 

two new species from Haiti (Hemip.). Proc Biol. Soc. 

Washington, vol. 41, pp. 21-24. 
1928c Some Tingitidae (Heteroptera) from Honduras. Occ Pap, 

Mus. Zool., Univ. Michigan, No. 190, pp. 1-5, 1 fig. 
1928f. Four undescribed tingitids from United States. Florida Ent., 

vol. 12, pp. 3-5. 
1928g. A synopsis of the species of Pachycysta (Hemip: Tingitidae). 

Ent. News, vol. 39, pp. 184-185. 



LACEBUGS OF THE WORLD 497 

1928h. A new subgenus and species of Tingis from Burma (Hemip.). 

Indian Forest Records, vol. 8, No. 4, pp. 283-284. 
1928i. Concerning some Tingitidae from Argentina with descriptions 

of five new species. Physis (Rev. Soc. Argentina Cienc. 

Nat.), vol. 9, pp. 72-76. 
1929. Some Tingitoidea from Central and South America. Bull. 

Brooklyn Ent. Soc, vol. 24, No. 1, pp. 35-37, 1 fig. 
1930a. Some Tingitidae (Hemiptera) from Brazil. Amer. Mus. Nov., 

No. 398, pp. 1-3. 
1930b. Some Tingitidae from Brazil (Hemiptera). Bull. Brooklyn 

Ent. Soc, vol. 25, No. 1, pp. 25-26. 
1930c A new sugar-cane tingitid from Java and Sumatra (Hemiptera). 

Pan-Pacific Ent., vol. 7, No. 1, pp. 15-16. 
1930d. Notes on American Tingitidae (Hemiptera). Bull. Brooklyn 

Ent. Soc, vol. 25, No. 5, pp. 268-272. 
1930e. Concerning some Tingitidae from the Philippines (Hemiptera), 

with new species. Proc Ent. Soc. Washington, vol. 32, 

No. 9, pp. 165-168. 
1931a. Two new species of Tigava from South America, (Hemiptera, 

Tingitidae). Proc. Hawaiian Ent. Soc, vol. 7, No. 3, pp. 

405-406. 
1931b. The Cornell University Entomological Expedition to South 

America 1919 and 1920, scientific results. No. 5. Hemip- 

tera-Tingitidae. Ann. Ent. Soc. America, vol. 24, No. 3, 

pp. 510-514. 
1931c Neotropical Tingitidae (Hemiptera). Ann. Mag. Nat. Hist., 

ser. 10, vol. 8, pp. 225-227, 1 fig. 
1931d. An undescribed tingitid from Borneo and Strait Settlements 

(Hemiptera). Pan-Pacific Ent., vol. 8, No. 2, p. 96. 
1931c Concerning the genus Leptodictya Stal (Hemiptera, Tingiti- 
dae). Bol. Mus. Nac, Rio de Janeiro, vol. 7, No. 2, pp. 

119-122. 

1932. Notes on some American Tingitidae (Hemiptera). Psyche, 

vol. 39, No. 4, pp. 100-102. 

1933. On some Tingitidae from South India including two new 

species (Hemiptera). Journ. Bombay Nat. Hist. Soc, vol. 
36, No. 4, pp. 1015-1016. 

1934a. An undescribed Acalypta from New York (Tingitidae: Hemip- 
tera). Bull. Brooklyn Ent. Soc, vol. 29, No. 5, p. 196. 

1934b. Un tingide nouveau provenant de Borneo et de Malacca. 
La Terre et La Vie, vol. 4, pp. 107-108, 1 fig. [Translation 
of 1931b.] 



498 U.S. NATIONAL MUSEUM BULLETIN 243 

1935. American Tingitoidea (Hemiptera) in the Natural History 

Museum of Vienna. Sondar-Abdruck aus "Konowia," vol. 

14, No. l,pp. 9-20, 2 figs. 
1936a. An undescribed tingitid from South India. Journ. Bombay 

Nat. Hist. Soc., vol. 39, No. 1, p. 145. 
1936b. Some Tingitoidea from Argentina (Hemiptera). Travaux de 

lTnstitut Zoologique de l'Academie des Sciences de l'URSS, 

vol. 3, pp. 699-701, 1 fig. 
1937a. Some Tingitidae from China, East Indies and India. Lingnan 

Sci. Journ., vol. 16, No. 3, pp. 385-388, 3 figs. 
1937b. Tingitidae from South China (Hemiptera). Lingnan Sci. 

Journ., vol. 16, No. 4, pp. 591-594, 1 fig. 
1938a. Mexican Tingitidae (Hemiptera). Pan-Pacific Ent., vol. 14, 

No. 2, pp. 70-72. 
1938b. Chinese Tingitidae (Hemiptera). Lingnan Sci. Journ., vol. 

17, No. 2, pp. 195-197. 
1939a. Three new species of Tingitidae (Hemiptera) from Australia. 

Pan-Pacific Ent., vol. 15, No. 2, pp. 87-88. 
1939b. A new genus and species of tingitid (Hemiptera) from Formosa. 

Trans. Nat. Hist. Soc. Formosa, vol. 29, No. 191, pp. 205-206. 
1939c. Chilean Tingitoidea (Hemiptera). Revista Ent., Rio de 

Janeiro, vol. 10, No. 2, pp. 330-334. 
1939d. Seven new South American Tingitidae (Hemiptera). Revista 

Ent., Rio de Janeiro, vol. 10, No. 3, pp. 525-530. 
1939e. A new tingitid from Palau Islands (Hemiptera). Mushi, vol. 

12, pp. 102-103. 
1939f. Two new tingitids (Hempitera) from Panama. Psyche, vol. 

46, Nos. 2-3, pp. 68-69. 
1940a. An undescribed Corythucha (Tingitidae-Hemip.) from Colorado. 

Ent. News, vol. 51, p. 172. 
1940b. New American Teleonemia (Hemip.: Tingitidae). Revista 

Chilena Hist. Nat., vol. 44, pp. 242-243. 
1941a. New American Tingitidae (Hemiptera). Journ. Washington 

Acad. Sci., vol. 31, No. 4, pp. 141-145. 
1941b. Three new American Tingitidae (Hemiptera). Pan-Pacific 

Ent., vol. 17, No. 3, pp. 139-141. 
1942a. New Tingitidae (Hemiptera). Iowa State Coll. Journ. Sci., 

vol. 17, No. 1, pp. 1-21. 
1942b. New Australian Tingitidae (Hemiptera). Journ. Washington 

Acad. Sci., vol. 32, No. 12, pp. 359-364. 
1943a. A list of species of Monanthia Lep. and Serv. of the Western 

Hemisphere, including description of a new species (Hemip- 
tera: Tingidae). Proc. Ent. Soc. Washington, vol. 45, No. 6, 

pp. 141-142. 



LACEBUGS OF THE WORLD 499 

1943b. Concerning the genus Lepturga Stal (Hemiptera, Tingitidae). 

Revista Chilena Hist. Nat., vol. 45, pp. 175-177. 
1944a. A new genus and ten new species of serenthiines (Hemiptera: 

Tingitidae). Proc. Ent. Soc. Washington, vol. 46, No. 3, 

pp. 67-76, 2 figs. 
1944b. Concerning the American cantacaderinids (Hemiptera: Tingi- 

dae). Bol. Ent. Venezolana, vol. 3, No. 3, pp. 139-142, 1 fig. 

1945. New Tingidae (Hemiptera). Bull. Southern California Acad. 

Sri., vol. 44, No. 3, pp. 96-100. 

1946. A new tingid from Yap Island (Hemiptera). Mushi, vol. 17, 

No. 6, pp. 27-28. 

1947a. Wissenschaftliche Ergebnisse der schwedischen entomologi- 
schen Reisen des Herrn Dr. A. Roman 1914-15 und 1923-24 
in Amazonas. 17. Tingididae. Arkiv for Zoologi, vol. 39B, 
No. 3, pp. 1-2. 

1947b. A new genus and two new species of Tingidae (Hemiptera). 
Bol. Mus. Nac, Rio de Janeiro, vol. 81, pp. 1-4, 1 fig. 

1947c. Australian Tingidae (Hemiptera). Bull. Southern California 
Acad. Sci., vol. 46, No. 3, pp. 111-121, 3 figs. 

1947d. Tingidae (Hemiptera) from the Orient and South Pacific. 
Musee Heude, Notes d'Entomologie Chinoise, vol. 11, No. 7, 
pp. 225-231, 4 figs. 

1 947e. Tpsotingis sideria Drake. Musee Heude, Fichier Entomologique 
Chinois, No. 67, fig. (color). 

1947f. Cantacader japanicus Drake. Musee Heude, Fichier Entomologi- 
que Chinois, No. 68, fig. (color). 

1947g. Trachypeplas chinensis Drake. Musee Heude, Fichier Entomo- 
logique Chinois, No. 69, fig. (color). 

1947h. Tingis coomani Drake. Musee Heude, Fichier Entomologique 
Chinois, No. 70, fig. (color). 

1948a. Some Tingidae (Hemiptera) from China, Japan and India. 
Musee Heude, Notes d'Entomologie Chinoise, vol. 12, No. 1, 
pp. 1-9, 3 figs. 

1948b. New species of Stephanitis Stal including a list of species of the 
world (Hemiptera). Musee Heude, Notes d'Entomologie 
Chinoise, vol. 12, No. 6, pp. 45-56, 4 figs. 

1948c. New American Tingidae (Hemiptera). Bol. Ent. Venezolana, 
vol. 7, Nos. 1-2, pp. 20-25. 

1948d. Five new American Tingidae (Hemiptera). Bol. Ent. Vene- 
zolana, vol. 7, Nos. 1-2, pp. 15-19. 

1948e. New genera and species of Tingidae (Hemiptera). Proc. Biol. 
Soc. Washington, vol. 61, pp. 149-156. 

1948f. The genus Compseuta Stal (Hemiptera, Tingidae). Proc. Ent. 
Soc. Washington, vol. 50, No. 8, pp. 197-204, fig. 



500 U.S. NATIONAL MUSEUM BULLETIN 243 

1948g. New Tingidae from the Orient and other regions (Hemiptera: 

Heteroptera) . Musee Heude, Notes d'Entomologie Chinoise, 

vol. 12, No. 15, pp. 173-178. 
1948h. New and little known Piesmidae and Tingidae (Hemiptera). 

Zoologische Mededeelingen, vol. 30, No. 4, pp. 73-76, 1 fig. 
1948L The genus Cochlochila Stal (Hemiptera: Tingidae). Musee 

Heude, Notes d'Entomologie Chinoise, vol. 12, No. 16, pp. 

179-181. 
1948j. American Tingidae (Hemiptera). Revista Ent., Rio de 

Janeiro, vol. 19, No. 3, pp. 429-436. 
1950a. Concerning the Cantacaderinae of the world (Hemiptera: 

Tingidae). Arthropoda, vol. 1, Nos. 2-4, pp. 153-166, 3 figs. 
1950b. A new tingid from the Canal Zone (Hemiptera). Proc. Ent. 

Soc. Washington, vol. 52, No. 6, pp. 299-300. 

1951. New genera and species of Tingidae (Hemiptera) in the Hun- 

garian National Museum. Ann. Hist. Nat. Mus. Nat. 
Hungarici, vol. 1, No. 1, pp. 165-178. 

1952. The genus Parada Horvath (Hemiptera: Tingidae). Psyche, 

vol. 59, No. 4, pp. 143-147. 
1953a. New Neogaean Tingidae (Hemiptera). Great Basin Nat., vol. 

13, Nos. 1-2, pp. 13-16. 
1953b. Synonymic data and description of new genera and species 

of Tingidae (Hemiptera). Great Basin Nat., vol. 13, Nos. 

3-4, pp. 91-99. 
1953c. Two new Tingidae (Hemiptera). Psyche, vol. 60, No. 4, 

pp. 151-154. 
1953d. A new genus and new species of Tingidae (Hemiptera). Proc. 

Biol. Soc. Washington, vol. 66, pp. 211-224. 
1954a. New Senegalese Tingidae (Hemiptera). Bull. Inst. Francais 

d'Afrique Noire, vol. 16, No. 1, pp. 232-237. 
1954b. A miscellany of new Tingidae (Hemiptera). Proc. Biol. Soc. 

Washington, vol. 67, pp. 1-15. 
1954c. New genera and species of Tingidae from the Old World 

(Hemiptera). Philippine Journ. Sci., vol. 83, No. 1, pp. 

69-73. 
1954d. Four new species of American Tingidae (Hemiptera). Proc. 

Ent. Soc. Washington, vol. 56, No. 2, pp. 75-78. 
1954e. Some tingids from the Belgian Congo (Hemiptera: Tingidae). 

Bull. Inst. Roy. Sci. Nat. Belgique, vol. 30, No. 17, pp. 1-10, 

lfig. 
1954f. Tingidae: descriptions and synonymic data (Hemiptera). 

Great Basin Nat., vol. 14, Nos. 1-2, pp. 1-10. 



LACEBUGS OF THE WORLD 501 

1954g. New and little-known Hemiptera-Heteroptera, Tingidae, from 
Ethiopia. Ann. Mag. Nat. Hist, ser. 12, vol. 7, pp. 657-655, 
4 figs. 
1955a. Three new Tingidae (Hemiptera). Bull. Southern California 

Acad. Sci., vol. 54, No. 2, pp. 78-81, 1 fig. 
1955b. Angolan Tingidae (Hemiptera). Publicacoes Culturais da 

Companhia de Diamantes de Angola, separata 27, pp. 83-90, 

2 figs. 
1955c. Some Tingidae from the French Cameroons (Hemiptera). 

Proc. Biol. Soc. Washington, vol. 68, pp. 105-108. 
1955d. New Tingidae (Hemiptera) from the Belgian Congo. Bull. 

Inst. Roy. Sci. Nat. Belgique, vol. 31, No. 85, pp. 1-5, 1 fig. 
1955e. A new genus and species of African lacebug (Tingidae: Hemip- 
tera). Rev. Zool. Bot. Africaine, vol. 52, Nos. 3-4, pp. 280-