Q
11
U563
CRLSSI
SMITHSONIAN
INSTITUTION
MUSEUM
OF
NATURAL
HISTORY
UNITED STATES NATIONAL MUSEUM BULLETIN 280
A Revision of the Peoriinae
and Anerastiinae (Auctorum)
of America North of Mexico
(Lepidoptera: Pyrahdae)
JAY C. SHAFFER
Visiting Research Associate
Smithsonian Institution
SMITHSONIAN INSTITUTION PRESS
WASHINGTON, D.C.
1968
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Contents
Page
Introduction 1
Materials and Methods 2
Acknowledgments 4
Review of the Literature 5
Classification 6
Subfamily Peoriinae 8
Genus Peoria Ragonot 9
P. longipalpella (Ragonot) 13
P. bipartitella Ragonot 14
P. tetradella (Zeller) 16
P. opacella (Hulst) 18
P.floridella Shaffer 20
P. rostrella (Ragonot) 21
P. gemmatella (Hulst) 22
P. roseotinctella (Ragonot) 24
P. johnstoni Shaffer 26
P. santaritella (Dyar) 27
P. holoponerella (Dyar) 28
P. approximella (Walker) 29
P. luteicostella ( Ragonot) 36
Genus Anacostia Shaffer 37
A. tribulella Shaffer 38
Genus Arivaca Shaffer 39
The Pimella Group 40
A. pimella (Dyar) 41
A. linella Shaffer 42
The OstreeUa Group 42
A. ostreella ( Ragonot) 43
A. poohella 44
The Albidella Group 45
A. albidella (Hulst) 46
A. artella Shaffer 46
The Albicostella Group 47
A. albicostella (Grossbeck) 47
Genus Atascosa Hulst 48
A. glareosella (Zeller) 49
Genus Homosassa Hulst 50
H. ella Hulst 51
H. platella Shaffer 53
H. incudella Shaffer 54
Genus Reynosa Shaffer 54
R. floscella (Hulst) 55
Genus Goya Ragonot 56
G. stictella (Hampson) 56
V
VI CONTENTS
Page
Subfamily Phycitinae 57
Genus Anerastia Iliibner 58
A. lolella Hiibner 59
Genus Coenochroa Ragonot 60
C . californiella Ragonot 61
C. illihella (Hulst) 65
C. bipundella (Barnes and McDunnough) 67
Genus Bandera Ragonot 68
B. binolella (Zeller) 69
B. cwpidinella Hulst 71
B. virginella Dyar 73
Genus Wakulla Shaffer 74
W. carneella (Barnes and McDunnough) 75
Genus Tampa Ragonot 75
T. dimediatella Ragonot 76
Genus Barberia Dyar 78
B. affinitella Dyar 78
Genus Ragonotia Grote 79
R. dotalis (Hulst) 80
Genus Martia Ragonot 83
M. arizonella Ragonot 84
Synonymical List of Species Considered 85
Unplaced Genera and Species 87
Literature Cited 90
Explanation of Figures 93
Introduction
This study began as a revision of the North American Anerastiinae,
largely in the sense of the McDunnough (1939) checklist. In keeping
with the original goals of the investigation, this paper treats all species
formerly placed under that subfamily name, although Anerastia and
several other genera are herein transferred to the Phycitinae, and
Peoria is taken as the nominal genus of the subfamily. The transferred
genera have not previously been treated in detail and so are discussed
in this paper. The following introductory sections deal with the litera-
tm-e and history of the classification of all the genera formerly placed
in the Anerastiinae.
The species of moths which previously constituted the subfamily
Anerastiinae are a mixture of various pyralid elements. Members of
this worldwide group are generally uncommon, rather inconspicuous
insects of little or no economic importance, and perhaps largely for
these reasons have received relatively little attention from taxonomists.
Much of the work which has been done on the North American
Anerastiinae appeared during the last 15 years of the 19th centmy
and is superficial by current standards. The only world monograph of
the group is that of Ragonot (1901). Because existing works on the
anerastiines have not been based on the detailed examination of
species which is so necessary to a natiu^al taxonomic system, a number
of areas remain as major problems.
Identification of species has been difiicult, even for specialists, be-
cause the descriptions and illustrations are inadequate. Most of the
original descriptions occupy but a few lines of text and are almost
useless for identifying the species. Illustrations of the adults are nearly
absent from the literature with the exception of the Ragonot revision,
a rare publication not easily available to most workers. The genitalia,
one of the most taxonomically useful structures, are nowhere illus-
trated for North American species and are described in only a few
instances. In addition most past studies were made from only one or a
few specimens so that no account of individual and geographic varia-
tion was possible. Thus, reliable identifications can be made only by
comparison with the type specimens. This is difficult because the
types are scattered in various museums throughout the United States
and Europe.
Placement of species into genera has been on the basis of usually
one or a few superficial characters of wings, antennae, and palpi,
1
2 U.S. NATIONAL MUSEUM BULLETIN 280
without emphasis on the sum of these characters and without examina-
tion of the genitaUa. As a result most species have been misplaced,
often associated in Old World genera to which they have no close
aflBnities.
The Anerastiinae and Phycitinae have long been considered closel}''
related within the Pyralidae and were often collectively recognized as
the family Phycitidae. The two groups show close similarity in wing
venation, notably the loss of vein R5 in both groups. They also have
similar palpi and the frenulum reduced to a single bristle in the females.
Separation of the Anerastiinae from the Phycitinae has been made
on the basis of a single reduction character, loss or extreme reduction
of the tongue, hence the supposed monophyletic nature of the group
has been highly questionable. The results of the present investigation
indicate that in North America eight genera properly belong with the
phycitines and seven genera, forming a natural group, are quite dis-
tinctly separate from the Phycitinae. The latter of these two groups
has been given subfamily status as the Peoriinae, a subfamily name
proposed by Hulst.
The present study is preliminary; essentially all that is known of
the group is based on studies of the adults; the immature stages and
the biology of the living insects are almost wholly unknown. Food
plant records are available for only two species of peoriines, both grass
feeders. Peoria bipartitella is reported to feed on Panicum lanuginosura,
and Peoria tetradella on Elymus canadensis as a borer in the stem. The
life history oi Anerastia lotella, a Holarctic species, is well described in
the literature, notably by Buckler (1901). The species feeds on a
number of grasses including Ammophila arenaria, Festuca ovina, Secale
cereale, and Aira species. It is hoped that this paper will encourage
taxonomic and biological studies on the Peoriinae throughout their
worldwide range.
Materials and Methods
Specimens examined. — The U.S. National Museum and the
Canadian National collection have provided most of the specimens
used in this study. A complete listing of individuals and institutions
from which material was borrowed is given below along with the
abbreviations used in the text.
ABK Collection of Alexander B. Klots, New York, N.Y.
AMNH American Museum of Natural History, New York, N.Y.
ANS Academy of Natural Sciences, Philadelphia, Pa.
BM Collection of Bryant Mather, Jackson, Miss.
BMNH British Museum (Natural History), London, England
BPI State Bureau of Plant Industry, Harrisburg, Pa.
CM Carnegie Museum, Pittsburgh, Pa.
CNC Canadian National Collection, Ottawa, Ontario
REVISION OF PEORIINAE AND ANERASTIINAE 3
CNHM Chicago Natural History Museum, Chicago, 111.
CPK Collection of Charles P. Kimball, Sarasota, Fla.
CU Cornell University, Ithaca, N.Y.
INHS Illinois State Natural History Survey, Urbaua, 111.
JCS Collection of the author, Shamokin Dam, Pa.
JGF Collection of John G. Franclemont, Ithaca, N.Y.
KSU Kansas State University, ISIanhattan, Kans.
LACM Los Angeles County Museum, Los Angeles, Calif.
MOG Collection of IMurray O. Glenn, Henry, HI.
UCB University of California, Berkeley, Calif.
UK University of Kansas, Lawrence, Kans.
UM University of Minnesota, St. Paul, Minn.
USNM U.S. National Museum, Washington, D.C.
Techniques. — Preparatoiy to examining the genitalia the abdomen
was broken away from the thorax and macerated in cold 10 percent
potassimn hydroxide (KOH) solution for from 4 to 8 hours, then
dissected in 50 percent alcohol. The abdomen was carefully torn open
with jewelers' forceps along the entire right plem-al area, enabling the
cuticle to be mounted flat with the outer surface upward. The genitalia
were separated by carefully tearing along the membrane between
segments eight and nine. After the removal of loose scales and KOH,
the male genitalia were usually stained with mercmochrome (0.3
percent aqueous solution), or occasionally with chlorazol black or
acid fuchsin to improve rendition of various structures. The aedeagus
was removed, and the valves were spread and held open by a small
rectangular section of microscope slide placed over the genitaha. The
genitalia were held in this position in 95 percent alcohol until hardened,
then transferred successively to clove oU and xylene for about 15
minutes each, prior to mounting in Canada balsam on a microscope
slide.
Female genitalia were put through the same series of chemicals as
those of the males, but were usually stained with chlorazol black or
mercm-ochrome.
In order to allow observation from any desired angle the genitalia
of at least one specimen of each sex of each species were transferred
from xylene into small (5 x 10 mm) genitalia vials, along with a drop
of Cargille's type A microscope immersion oU; the males were left
unspread. These specimens were eventually washed in xylene and
moimted in balsam; before mounting they proved very useful in
preparing descriptions because it was possible to study the relation-
ships of the various structures.
Attempts to spread the male genitalia of the Peoriinae in the
manner described above caused considerable distortion; thus the
method had to be modified. If one will visualize the external male
genitaha as the aedeagus and a surrounding cyhnder, and then the
cylinder as severed lengthwise, the genitalia can be "unrolled" and
4 U.S. NATIONAL MUSEUM BULLETIN 2 80
flattened. In practice this was achieved by carefully separating the
right valve from the vinculum and tearing the diaphi-agma so that
the juxta remams adjacent to the right valve. The method was highly
satisfactory as all the genitalic structures then lay flat in one plane.
The genitalia were held flat, run through the alcohol, clove oil, xylene
series, and mounted as explained above. Illustrations of the peoriine
species and several other species were made with the male genitalia
prepared in this manner.
Terms. — The Comstock-Needham system has been used in naming
the wing veins (fig. 50). I have followed Kornerup and Wanscher
(1962) for color terminology, viewing both the color plates and the
specimen by incandescent illumination. Physiographic distribution
terminology follows Lobeck (1948). Unless otherwise indicated, the
phrase "North America" is taken to mean America north of Mexico.
Acknowledgments
The investigation was carried out under the dhection of Prof.
John G. Franclemont, to whom I wish to express my gratitude for
his kind advice during the course of the project and for his helpful
criticisms of the manuscript. A special note of appreciation is due
Mr. Paul E. S. Whalley, who checked several type specimens in the
British Museum (Natural Histor37^) against my descriptions and photo-
graphs; to Dr. Pien-e E. L. Viette, who made available many of the
Ragonot types in the Paris Museum; to Dr. Alexander B. Klots for
his numerous helpful suggestions; to Dr. J. F. Gates Clarke, who sug-
gested and encouraged my work on this interesting problem, and to
Dr. H. E. Moore, Jr. and Dr. W. D. Duckworth, who offered useful
comments on the manuscript.
I wish to thank the Allied Chemical Corporation for a contribution
toward the cost of preparing the manuscript and for a grant which
enabled me to spend one month collecting specimens at the Archbold
Biological Station, Lake Placid, Fla. I am also indebted to the pub-
lisher of the Goode Base Map Series, Department of Geography,
The University of Chicago, for granting permission to reproduce
copyrighted material used in several of the plates. Maps 1 through 6
and Map 12 are based on Goode Base Map No. 202, copyi'ight by
the University of Chicago.
Finally, the investigation would not have been possible without the
kind cooperation of numerous individuals and institutions who loaned
specimens for my studies. I extend my appreciation to Dr. George W.
Byers, University of Kansas; Mr. Harry K. Clench, Carnegie Museum;
Mr. Hugh B. Cunningham, Illinois Natural History Survey; Dr.
Donald R. Davis, U.S. National Museum; Dr. Howard E. Evans,
REVISION OF PEORIINAE AND ANERASTIINAE 5
Museum of Comparative Zoology, Harvard University; Dr. John G.
Franclemont, personal collection; Mr. Murray O. Glenn, personal
collection; the late Dr. Harold J. Grant, Jr., Philadelphia Academy
of Natural Sciences; Mr. Charles P. Kimball, personal collection;
Mr. Norman Marston, Kansas State University; Mr. Lloyd M.
Martin, Los Angeles County Museum; Mr. Bryant Mather, personal
collection; Dr. Eugene Munroe, Canadian National collection; Dr.
L. L. Pechuman, Cornell University; Dr. Jerry Powell, University of
California, Berkeley; Dr. Frederick H. Rindge, American Museum of
Natural History; Mr. George B. Sleesman, Pennsylvania State Biu-eau
of Plant Industry; Dr. Frederick W. Stehr (now at Michigan State
University), University of Minnesota; Mr. Paul E. S. Whalley,
British Museum (Natural History); and Mr. Alex Wyatt, Chicago
Natural History Museum.
Review of the Literature
The genus Anerastia Hiibner [1826] originally comprised four
species; three of these were transferred to Hypochalcia Hiibner by
Zeller (1848), and the one remaining species, lotella Hiibner [1810-
1813], was assumed to be the type of the genus although it was not
explicitly designated as such until 1886 by Moore. In the original
publication of lotella, Hiibner gives only a watercolor figure, no
description. Genitalia of both sexes were figured and described briefly
by Pierce (1938).
Walker (1866) originally described approodmella, the same species
later being named haematica by Zeller (1872), and roseatella by
Packard (1873). ZeUer's paper included descriptions of three other
new species of North American Anerastiinae.
In 1886 Ragonot established the family Phycitidae, based on
Phycita Curtis, and divided it into the subfamilies Phycitinae and
Anerastiinae based upon the "absence, or nearly so, of the tongue"
in the latter group. He states his intention to prepare a monograph
of the Phycitidae and Galleridae of the world and gives a review of
previous works on these insects.
The publication of Hulst's "Descriptions of New Pyralidae" (1886)
prompted Ragonot to publish new species of his own in advance of
the projected monograph. The appearance of the Hulst papers at
this time was in some respects unfortunate, for in the next several
years both men published many new specific and generic names
without seeing each other's types, thereby creating many synonyms.
During 1887-89 Ragonot published 16 species and 15 genera, and
Hulst published 7 species and 4 genera listed as new for the North
American Anerastiinae.
6 U.S. NATIONAL MUSEUM BULLETIN 280
Grote (1888) regarded "... the Phycidae or Phyciinae as a
sub-family of the Pyrahdae; the M. Ragonot's Anerastinae [sic] as
merely a tribal division of the sub-family."
The first paper summarizing knowledge of the Anerastiinae was
Hulst's (1890) "Revision of American Phycitidae." Therein he
divided the family into the subfamilies Phycitinae and Peoriinae, the
latter group largely corresponding to Ragonot's Anerastiinae, but
excluding Anerastia and two other genera. The division was rather
auspiciously made on the basis of the male genitalia, but unfortunately
his assignment of genera into the two groups suggests that he neglected
careful examination of these structures for most species. Indeed,
genitalia are discussed for only 4 of the 19 peoriine genera included
in the paper. The revision treats 30 species in 19 genera of Peoriinae,
with 7 of the species and 4 of the genera being described as new.
The great world monograph of Ragonot (1901) was completed by
Sir George F. Hampson after the author's death in 1895 and contains
descriptions of 31 species and illustrations in color of 30 species
(lotella Hiibner not being illustrated) of North American Anerastiinae
placed in 17 genera. Three species and one genus were described as
new. A number of the specific names have since been synonymized,
and several of the genera have been found not to be North American.
Hampson's (1918) classification of the subfamily termed the group
Hypsotropinae based on Hypsotropa Zeller. He considered dignella
Hiibner to be the type oi Anerastia, but Ragonot (1901) had already
designated lotella Hiibner as such. His paper covers the entire group
and ascribes one new species to the North American fauna.
Sixteen new specific names have been added to the literature since
the publication of Ragonot's monograph, these mostly in occasional
papers by Dyar (1904, 1904a, 1906, 1908, 1923), Barnes and McDun-
nough (1913), Hampson (1918, 1930), and Grossbeck (1917). The
most recent checklist for North America (McDunnough, 1939)
gives 23 genera and 51 species, plus 1 genus and 10 species as junior
synonyms. Heinrich's "Revision of the New World Phycitinae"
(1956) transfers a number of genera across the subfamily lines.
Classification
As has been indicated previously, available information on the
Anerastiinae is minimal, so that the present study is only a step
toward a more reliable taxonomic system such as exists for many
better known groups of insects. Nevertheless, morphological studies
of adult specimens indicate that radical changes in the classification
of the Anerastiinae are needed.
REVISION OF PEORIINAE AND ANERASTIINAE 7
I have accepted the male genitalia as being the most reliable
morphological indicators of natural relationships, especially at the
generic level. They offer a relatively large number of stable characters.
Female genitalia are in many cases too similar to be of much value
in separating species, and even genera. Characters of wings, antennae,
and palpi previously used to define genera often exhibit more intra-
generic than intergeneric variation and consequently are by them-
selves extremely unreliable indicators of generic lines.
Numerous examples of wing venation variability could be given,
and the following will illustrate the lack of reliabiHty of venational
characters: In the genus Peoria the presence or absence in the fore-
wing of vein M3 does not correlate with any other morphological
feature investigated. This vein was present in about one-half of the
specimens of P. santaritella, and one specimen possessed M3 only on
the right wing. Aberrations in the wing venation are very common
within the subfamily, and the usefulness of any particular venational
feature must be evaluated separately.
Orientation of the palpi may vary with the sex, and in living
specimens the position they assume may be different from that
found in dead specimens after desiccation. Unfortunately, most species
have been described from very small series of specimens, so the
variation has not always been apparent. Thus the use of this character
in assigning species to genera has proved to be extremely unreliable.
The definition of the Anerastiinae has not generally been regarded
as very satisfactory. Although most of the species have a distinctive
habitus, this and the reduced tongue are not, in the absence of other
substantial correlated characters, sufficient to define the group.
Heinrich (1956), following Ragonot's division of the Phycitidae, felt
that ". . . in the main the subfamilies [Phycitinae and Anerastiinae]
themselves appear to be natural entities, although their definition
leaves much to be desired."
On the basis of the genitalia, the North American Anerastiinae
divide into two distinctly separate groups. One of these groups is
composed of eight phycitine genera, including Anerastia, which seem
to be related to diverse areas of that subfamily, and the remaining
seven genera comprise a natural group, the Peoriinae.
The genitalia of both sexes in the peoriines differ distinctly from
those in the phycitines. Females all have the ovipositor greatly com-
pressed, probably as an adaptation for a particular mode of oviposi-
tion. In the males the uncus is so different from its usual appearance
in the phycitines as to be nearly unrecognizable in the North American
species. Indeed, the assumption that these terminal parts represent
uncus is based on an examination of several Old World peoriine
8 U.S. NATIONAL MUSEUM BULLETIN 280
genera in which the uncus is similar, in varying degrees, to that of
the phycitines.
A number of other features are generally, but not exclusively,
characteristic of the Peoriinae. In all species the males have two to
five (possibly more) basal antennal segments fused to some degree,
the exact number often varying within a species. Longitudinal wing
markings are more common than transverse ones, and the ocelli
are well developed in all the species examined.
Inasmuch as the present state of our knowledge is so unsatisfactory,
the author hopes to undertake studies of the group on a world basis
to better deal with the problem. The interpretation of just how
closely the Phycitinae and the Peoriinae are related will depend in
part on how homologies between the male genitalic structures of the
two groups are interpreted, and in part on the significance of the
compressed ovipositor. A better understanding of this problem will
require examination of the genitalic structures of all the species,
and perhaps studies of their embryology and soft-part anatomy as
weU. Life history studies would be extremely valuable and should
help to elucidate the functional significance of the compressed
ovipositor.
Subfamily Peoriinae * Hulst
Hulst, 1890, p. 102.
Diagnosis. — Similar to Phycitinae; distinguished by spicate
modifications of uncus in males and by highly compressed ovipositor
of females.
Description. — Labial palpi weU developed, maxillary palpi mod-
erate to very small; male antennae with basal segments of shaft
variously fused; tongue reduced; ocelli well developed.
Forewing maculation predominantly longitudinal; transverse bands,
when present, usually expressed as dots, more rarely solid; 10 or 11
veins; Ri free from ceU; R3+4 stalked; Kg absent; Mi free from cell;
Cu2 free from ceU. Hind wing with 6 or 7 veins; Rs closely approximate
to or stalked with Sc-f Ri beyond cell; frenulum of female simple.
Male genitalia with uncus bearing spicate processes. Gnathos
various, usually bearing medial process. Transtilla absent (except
in Goya). Juxta well developed, various. Vinculum well developed.
Eighth abdominal segment without special hair tufts.
Female genitalia with ovipositor strongly compressed. Eighth
abdominal segment compressed, rarely well expanded anteriorly.
Apophyses well developed, approximately equal in length. Ductus
1 Based upon North American species.
REVISION OF PEORIINAE AND ANERASTIINAE 9
bursae moderately short. Bursa well developed, rarely armed. Ductus
seminalis from or near posterior end of bursa.
Key to Genera of Peoriinae Based Upon the Male Genitalia
Within the Peoriinae intrageneric variation of alar, palpal, and
antennal features seriously limits their usefulness in separating genera.
In the absence of other distinguishing features, keys to genera must
rely on characters of the male genitalia. Whenever slide preparation is
impractical, reasonably certain determinations can often be made by
comparing external features of the specimens to descriptions and
photographs of adults. Genitalia slides should be prepared if correct
identification is critical. It is suggested that the indicated figures be
consulted when using the keys.
1. Each spicate process (of uncus) basally branched into two long perpendicular
tapering spines of nearly equal length; spicate processes connected by medial
bandlike process (fig. 51) 2
Spicate processes unbranched, or one branch at least twice length of other;
medial process absent or not bandlike 3
2. Medial process of uncus bearing prominent posteromedial protuberance; juxta
with pair of setaceous tubercles (fig. 118) Anacostia
INIedial process of uncus unarmed, or with minute cusps only; juxta without
pair of setaceous tubercles (figs. 105-117) Peoria
3. Spicate processes unbranched, blunt, elbowed in middle; gnathos tapering to
single sharp apical spine (fig. 126) Atascosa
Spicate processes various, not angled; gnathos with medial process not a single
sharp spine 4
4. Each spicate process with rhomboidal lateral pad; juxta elliptical; aedeagus
strongly tapered (fig. 131) Goya
Rhomboidal pad absent; juxta rectangular or shield shaped; aedeagus not
strongly tapered 5
5. Each spicate process a reciu-ved stout hook shorter than gnathos arm, tapering
to sharp point; gnathos apically tricuspidate (fig. 130) Reynosa
Spicate processes not as above, at least as long as gnathos arm; gnathos not
apically tricuspidate 6
6. Apical process of gnathos digitate; spicate processes each with long lateral and
much shorter caudal branches (figs. 119-125) Arivaca
Gnathos without digitate apical process; spicate processes unbranched (figs.
127-129) Homosassa
Peoria Ragonot
Aurora Ragonot, 1887, p. 18; 1889, p. 117. — Hulst, 1890, pp. 209-210. — Smith,
1891, p. 84. — Ragonot, 1901, p. 337. — Hulst, 1902, p. 437. — Barnes and
McDunnough, 1917, p. 149. — Kampson, 1918, p. 106. — Forbes, 1923,
p. 638. — McDunnough, 1939, p. 35. [New synonymy. Type: Aurora
longipalpella Ragonot, 1887. IMonobasic.]
Peoria Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p. 213. — Smith,
1891, p. 85. — Ragonot, 1901, p. 386. — Hulst, 1902, p. 439. — Barnes and
10 U.S. NATIONAL MUSEUM BULLETIN 280
McDunnough, 1917, p. 149. — Forbes, 1923, p. 638. — McDunnough, 1939,
p. 36. [Type: Anerastia haematica Zeller, 1872. Original designation.]
Statina Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p. 216. — Smith,
1891, p. 85. — Ragonot, 1901, p. 415. — Hulst, 1902, p. 440. — Barnes and
McDunnough, 1917, p. 150. — Hampson, 1918, p. 59. — McDunnough, 1939,
p. 36. [New synonymy. Type: Statina roseotinctella Ragonot, 1887, p. 19.
Monobasic]
Calera Ragonot, 1888, p. 50. — Smith, 1891, p. 85. — Ragonot, 1901, p. 417.
— Hulst, 1890, p. 217, — Barnes and McDunnough, 1917, p. 150. — Hamp-
son, 1918, p. 59. — Hulst, 1902, p. 441. — Forbes, 1923, p. 639. — McDun-
nough, 1939, p. 36. [New synonymy. Type: Calera pundilimbella R-agonot,
1888, p. 50. Monobasic]
Altoona Hulst, 1888, p. 116; 1890, pp. 206-207. — Smith, 1891, p. 84. — Hulst,
1902, p. 438. [New synonymy. Type: Anerastia opacella Hulst, 1887.
Original designation.]
Cayuga Hulst, 1888, p. 116; 1890, pp. 208-209. — Smith, 1891, p. 84. — Hulst,
1902, p. 438. [New synonymy. Type: Spermatophthora gemmatella Hulst,
1887. Monobasic]
Volusia Hulst, 1890, p. 206 [not Robineau-Desvoidy, 1830, p. 674, in Diptera,
Ortalidae; not Adams, 1861, p. 306, in Gastropoda]. — Smith, 1891, p. 84.
[New synonymy. Type: Volusia roseopennella Hulst, 1890. Monobasic]
Wekiva Hulst, 1890, p. 215. — Smith, 1891, p. 85. — Hulst, 1902, p. 440. [New
synonymy. Type: Wekiva nodosella Hulst, 1890. Monobasic]
Osceola Hulst, in Smith, 1891, p. 85 [not Baird and Girard, 1853, p. 133 p. 133, in
Reptilia]. [Nomen nudum. New synonymy. Type: Chipeta perlepidella
Hulst, 1892. Monobasic]
Chipeta Hulst, 1892, p. 62. — Barnes and McDunnough, 1917, p. 150. — McDun-
nough, 1939, p. 36. [New synonymy. Type: Chipeta perlepidella Hulst.
1892. Monobasic]
Chipota [sic] — Hulst, 1902, p. 441. [Misspelling for Chipeta.]
Trivolusia Dyar, 1902 [1903], p. 438. [New synonymy. Type: Volusia roseopen-
nella Hulst, 1890. Monobasic]
Ollia Dyar, 1904, pp. 107-108. — Barnes and McDunnough, 1917, p. 149. — Mc-
Dunnough, 1939, p. 36. [New synonymy. Type: Ollia santaritella Dyar,
1904. Monobasic]
Diagnosis. — Members of this rather variable genus may be recog-
nized by the characteristic uncus with its bandUlie medial ])rocess and
paired spicate processes, bifm-cate with each arm round, slender,
sharply pointed, and oriented at about 90 degrees to the other (fig. 51).
Description. — Labial palpi porrect (obliquely ascending in lutei-
costella); tongue rudimentary; antennae with base compressed, male
shaft with basal segments partly fused, female with shaft filiform,
finely cUiate.
Forewings with 10 or 11 veins; Ri from cell well before upper outer
angle; R2 from cell near base of or stalked with R3+4; R3+4 always
stalked, from just before the angle; Mi from the angle; M2 stalked or
fused with M3; M3 sometimes stalked with Cui; Cui at or near lower
outer angle of cell; Cu2 from just before the angle. Hindwings with
6 or 7 veins; Sc and Rs closely approximate or stalked; Mi from upper
REVISION OF PEORIINAE AND ANERASTIINAE 11
outer angle of cell; M2 absent; M3 stalked with Cui for at least one-
thii'd of its length, sometimes fused; Cui from lower outer angle; Cu2
from just before the angle.
Male genitalia with uncus bearing bandlike medial process; spicate
processes each terminating in slender, sharply pointed ventrad and
caudad arms. Gnathos arms broad and flat. Vinculum well developed,
broadly rounded. Aedeagus somewhat flattened, three to four times as
long as broad; vesica, when armed, bearing one or two cornuti.
Female genitalia with posterior and anterior apophyses of about
equal length, well developed. Ductus bursae moderately short. Bursa
unarmed. Ductus seminalis from posterior end of bursa, broadened at
base.
Discussion. — The genus includes 13 known species and is the
largest in the American fauna of the subfamily. Affinities vary within
the genus, and the species have been grouped to show relationships
in so far as this is possible with a linear arrangement.
There is enough variation of characters within many of the species
to render difficidt the construction and use of keys. To aid in the identi-
fication of specimens, a chart summarizing the characters found to be
of greatest value in separating species is included in addition to the
key.
In a few cases the male genitalia are quite distinctive of the species,
but for the most part the differences are rather subtle. The form of
the gnathos, number of cornuti in the vesica, and shape of the valvae
(not summarized in table, see figures) are the most useful characters
of the male genitalia.
Navasota hebetella may belong to Peoria and has been included in
the key (see Unplaced Genera and Species, p. 87).
In accordance with the recommendation of the Code (Article 24),
I have chosen the name Peoria for this genus rather than the lesser
known Aurora which has page precedence.
Explanation of Table 1
1. Fore wing, vein R2
x=free from cell
0= stalked with R3f4
2. Forewing, vein M2
x= stalked with M3
*= fused with M3 (apparently absent)
3. Forewing, vein Cui
x=free from cell
0= stalked with M3
4. Forewing, region anterior to cell
x= abruptly pale, with few or no darker scales
0= not abruptly pale, Sc and Rs sometimes white traced
285-934—68 2
12
U.S. NATIONAL MUSEUM BULLETIN 280
10.
11.
12.
Forewlng
x= ground darker posterior to cell, at least at base
0= ground not darker posterior to cell
Forewing
x=soine or all veins white traced
o= without white tracing on veins
Forewing, transverse posterior line
x= present
* = represented by dots
o= absent
Labial palpi
p=porrect
a= ascending
Maxillary palpi
x= reaching frons or nearly so
o= small, not approaching frons
Male antennae
1= laminate
S= serrate
ss=subserrate
Gnathos of male genitalia
a = with apical process
l=with pair of subapical processes
o== without special processes
Aedeagus, number of cornuti in vesica
The use of two symbols separated by a comma indicates two characters appear-
ing with about equal frequency. A symbol in parentheses indicates a character
appearing less frequently than its alternative.
Table 1. — Summary of characters found to be the most useful in distinguishing
species of Peoria*
Species
1
2
3
4
5
6
7
8
9
10
11
12
longipalpella
X
X
X
p
X
ss
a
1
bipartitella
*
X
o(x)
x(o)
x(o)
o
p
X
1
a
1
tetradella
x,o
X
X
X
X
o
p
X
ss
a
1
opacella
x,o
X
X
X
X
o
p
X
1
a
1
floridella
x,o
X
X
X
X
o
p
o
1
a
1
rostrella
o
X
X
o
o
X
p
X
ss
a
1
gemmatella
X
X
X
o
o
X
o
p
o
1
a,o
2
roseotinctella
*
X,0
o
X
*
p
o
1
a(o)
2
johnstoni
o
*
X
o
o
X
o
p
o
ss
o
2
santaritella
X
*,x
X
o
X
o
p
s
a
1
holoponerella
X
X
X
X
o
o
o
p
s
a
1
approximella
o
*
X
X
o
o
p
o
1
1
luteicostella
o
*
X
X
o
a
o
1
1
1
*See explanation on preceding pages.
REVISION OF PEORIINAE AND ANERASTIINAE 13
Key to the Species of Peoria Based Upon External Features
1. R2 stalked with R3+4; M J absent 2
II2 not stalked with R3+4; if stalked, then M2 present 7
2. M3 stalked with Cui; hindwing M3 absent 3
Ms not stalked with Cui, sometimes from point; hindwing M3 usually
present 4
3. Transverse posterior indicated by line of dots (fig. 10) roseotinctella
Transverse posterior absent; terminal line of dots present . . N. hebetella
4. Forewing abruptly pale anterior to cell, with no more than a few scattered
nonwhite scales 5
Forewing not abruptly pale anterior to cell 6
5. Forewing pale posterior to A2; palpi porrect (fig. 14) approximella
Forewing not pale posterior to Aj; palpi ascending (fig. 12) , . luteicostella
6. Forewing darker posterior to cell; maxillary palpi approaching frons; (fig.
13) bipartitella
Forewing not darker posterior to cell; maxillary palpi not approaching
frons; (fig. 1) johnstoni
7. Forewing with transverse bands (fig. 9) longipalpella
Transverse bands absent 8
8. Forewing light buff, abruptly pale anterior to cell (figs. 3, 4) . . holoponerella
Forewing not abruptly pale anterior to ceU 9
9. Veins traced with white 11
Veins not traced with white 10
10. Ground white anterior to cubitus; cubitus bordered posteriorly with red;
maxillary palpi not reaching frons (fig. 5) santaritella
Ground a mixture of brown and white scales; usually lighter anterior to
cubitus, at least near base; maxillary palpi reaching frons (fig.
11) opacella
11. Forewing darker posterior to cell, at least near base 12
Forewing not darker posterior to cell 13
12. Aa traced with white, bordered with brown; maxillary palpi not reaching
frons (fig. 8) floridella
A2 not traced with white; maxillary palpi reaching frons (fig. 11). . . opacella
13. Cubitus and lower outer angle of cell broadly traced with white (fig.
6) tetradella
Notasabove;ScandRstracedwith white; ground reddish brown . . . , 14
14. R2 free from cell (fig. 7) gemmatella
R2 stalked with Rj+4 (fig. 2) rostrella
Peoria longipalpella (Ragonot), new combination
Figures 9, 51, 70, 105, 144
Aurora longipalpella Ragonot, 1887, p. 18. — Hulst, 1890, p. 210. — Smith, 1891,
p. 84. — Ragonot, 1901, pp. 337-338. — Hulst, 1902, p. 437. — Barnes and
McDunnough, 1917, p. 149. — Hampson, 1918, p. 106. — Forbes, 1923, p,
638. — McDunnough, 1939, p. 35. — KimbaU, 1965, p. 250.
Diagnosis. — This is the only known species of Peoria in wliich the
forewings are marked with unbroken transverse bands, although
transverse rows of dots occur in a few other members of the genus.
14 U.S. NATIONAL MUSEUM BULLETIN 280
Description. — Frons brown; labial palpi with basal segments
white, second and third segments brown on outer sides, white ventrally
and on inner sides; maxillary palpi reaching frons, brown; antennae
brown, male subserrate; occiput laterally, patagia, and tegulae brown,
vertex and occiput light brown dorsally.
Forewings with ground white anterior to cubitus, brown posterior
to cell; cell sprinkled with brown, with orange red anterior to cell;
transverse anterior and posterior bands brown; ground brown on
underside, bands darker; with 11 veins; R2 stalked with R3+4; M2+3
stalked. Hindwings light brown; 7 veins; Sc and Rs stalked; M3
stalked with Cui.
Male genitalia with gnathos bearing apical digitate projection.
Juxta U-shaped. Valvae with costa sparsely setose, terminating in
sharp projecting tooth; sacculus densely pubescent. Aedeagus with
vesica bearing a single serrate cornutus.
Female genitalia with ovipositor tip moderately setose, caudal
margin with numerous fine setae, sparsely setose laterally. Apophyses
curved at base, posterior with base pointed and slightly flattened;
anterior with base thickened.
Type. — In the British Museum (Natural History).
Type data. — Lectotype female, hereby designated, "667; U.S.A.;
n, gen. R. longipalpella Rag"; genitaha slide No. 707, J. Shaffer,
Jan. 4, 1967. The specimen is glued together between the meso- and
metathorax. Type data given with the original description consists of:
"9 22 mill." Ragonot (1901) in a presumed reference to the type
reports: "Amerique boreale, 11 octobre. Une 9. Mus brit."
Specimens examined. — 2cf, 2 9.
Distribution (Map 8). — Recorded from North Carolina and
Virginia.
UNITED STATES: North Carolina, Polk Co., Tryon, 1 9 , no date (Fiske),
[USNM]; Wake Co., Raleigh, 1 9 , "M. Aug. 09," [USNM].
Virginia: Arlington Co., Arlington, 1 cT, Aug. 28, 1951 (J. G. Fran demon t),
[USNM].
Locality unknown: IcT, no date, Sweadner collection [CM].
Discussion. — Kimball reports the species from Siesta Key, Fla.,
May 13, 1946, but I have not seen the specimen.
Peoria bipartitella Ragonot
Figures 13, 71, 106, 145
Peoria bipartitella Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p. 214. —
Smith, 1891, p. 85. — Ragonot, 1901, p. 386. — Hulst, 1902, p. 439. —
Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36. —
KimbaU, 1965, p. 251.
REVISION OF PEORimAE AND ANERASTIINAE 15
Volusia roseopennella Hulst, 1890, p. 206. — Smith, 1891, p. 84. — Rindge, 1955,
p. 170. [New synonymy.]
Tolima roseopennella (Hulst). — Ragonot, 1901, p. 340. — Barnes and McDun-
nough, 1917, p. 149. — McDunnough, 1939, p. 35. — Kimball, 1965, p. 250.
Trivolusia roseopennella Hulst, 1902, p. 438. — Grossbeck, 1917, p. 134.
Commotria roseopennella (Hulst). — Hampson, 1918, p. 108.
Hypsotropa bipartitella (Ragonot). — Hampson, 1918, p. 78.
Peoria bipunctella Ragonot. — Forbes, 1923, p. 639. [Not Ragonot.]
Diagnosis. — The presence of a well-developed digitate medial
process on the gnathos serves to delimit this species from among
those of the genus with similar wing venation.
Description. — Frons conical, varying b^o^vn to Ught pink; labial
palpi mth basal segments white, outer sides of second and third
segments varying brown to light pink, inner sides with white-tipped
scales; maxillary palpi approaching frons, brownish white; antennae
light brown, laminate in male; occiput broAvn, vertex white behind
antennae, patagia and tegulae varying grayish red to brown, legs
grayish red to brown on outer sides, white on inner sides.
Forewings mth extremely variable maculation; area anterior to
cell red, brown, white, or mixtures of scales of these colors; cell usually
red or white, brown in some (Mississippi) specimens, often a mixture
of these three colors; area posterior to cell usually red, sometimes
brown; veins traced with white, tracing variable, sometimes absent;
10 veins, R2 stalked with R3+4; M2+3 fused, from lower outer angle.
Hindmngs light brown, darker in apical area; 7 veins; M3 and Cur
long stalked.
Male genitalia with gnathos bearing apical digitate process. Juxta
V-shaped. Vinculum with anterior margin flat and flanged. Valvae
mth costa sparsely setose, bearing blunt apical tooth; sacculus with
fine hairs. Aedeagus with vesica bearing single subserrate cornutus.
Female genitaha with ovipositor finely setose on caudal margin,
moderately setose laterally. Apophyses curved, posterior curving
upward at base and tapering to point, anterior shghtly thickened
at base.
Types. — P. bipartitella, in the Museiun National d'Histoire
Naturelle; V. roseopennella, in the American Museum of Natural
History.
Type data. — P. bipartitella, lectotype female, hereby designated,
North CaroHna, Morrison; genitaha slide No. 534, J. Shaffer, May 14,
1965; V. roseopennella, lectotype male, hereby designated, Florida,
March; genitaha shde No. 3220, Carl Heinrich, June 14, 1946; the
original description reports "Volusia County, Fla.," no date.
Recorded host. — Poaceae: Panicum lanuginosum Ell. from slide
label in USNM, specimens taken July 8, 1950, Kill Devil Hills, N.C
Specimens EXAMINED. — 34 cT, 19 $.
16 U.S. NATIONAL MUSEUM BULLETIN 280
Distribution (Map 8). — Chiefly the Atlantic and Gulf Coastal
Plain, recorded from Mississippi to Massachusetts, also from north
central Illinois.
Fernald (1950) reports the food plant as occurring in "Sandy open
soil, thin woods, etc., Fla. to Tex., n., rather locally, to s. R.I."
UNITED STATES: Florida: Alachua Co., Gainesville, 2cf, July 8, 1927
(J. Speed Rogers) [CU, USNM]; Icf, July 1927 (J. Speed Rogers) [CU]; High-
lands Co., Lake Placid, 1?, Mar. 6, 1945 (J. G. Needham) [CU]; Manatee Co.,
Myakka City, 1 ? , Feb. 17, 1945 (J. G. Needham) [CU]; Sarasota Co., Siesta
Key, IcT, Feb. 28, 1951 (C. P. Kimball) [CNC].
Illinois: Putnam Co., 1 9 , Aug. 29, 1961 (M. O. Glenn) [MOG]; 1 9 , Aug. 1,
1964; 1 9 , Aug. 29, 1964.
Massachusetts: Barnstable Co., Barnstable, 1 cT, July 10, 1949 (C. P. Kimball)
[CNC]; Icf, 2 9 , July 11, 1949 (C. P. Kimball) [CNC, CPK]; 2 d^, July 14, 1949
(C. P. Kimball) [CNC]; Icf, July 15, 1949 (C. P. Kimball) [CPK]; West Barn-
stable, 2 cf, 19, July 16, 1949 (C. P. Kimball) [CNC, CPK]; Barnstable, Icf,
July 11, 1950 (C. P. Kimball) [CPK]; Icf, July 28, 1950; 19, Aug. 7, 1952;
1 cf , July 4, 1958; Dukes Co., Martha's Vineyard, 1 9 , Aug. 1, 1926 (F. M. Jones)
[CNC]; 1 cf, Aug. 8, 1941 (F. M. Jones) [USNM]; 1 9 , Aug. 4, 1944 (F. M. Jones)
[ANS]; Id", Aug. 6, 1944; Id^, Aug. 6, 1946 (F. M. Jones) [CNC]; Icf, Aug. 4
(F. M. Jones) [USNM]; 19, Aug. 21 (F. M. Jones) [USNM]; Vineyard Haven,
Icf, July 17 (F. M. Jones) [ANS]; Plymouth Co., Agric. Exp. Sta., East Wareham,
Icf, July 11, 1962 [CPK].
Mississippi: Forrest Co., Camp Shelby, 6 cf , 3 9, Sept. 1-15, 1944 (C. D.
Michener) [AMNH]; 2 cf, Sept. 16-30, 1944.
New Jersey: Bergen Co., Oakland, 1 cf , Aug. 5, 1948 (C. P. Kimball) [CPK];
1 (?, Aug. 13, 1948 (C. P. Kimball) [CNC]; 1 9 , Aug. 14, 1948; Burlington Co.,
Whitesbog, 1 9, July 12, 1939 (E. P. Darlington) [ANS]: 1 cf, Aug. 2, 1940.
North Carolina: Macon Co., Horse Cove, Highlands, 1 cf, Aug. 3, 1957
(C. J. Curen) [CNC]; Polk Co., Tryon, 1 cT, Aug. 18, 1903 (Fiske) [USNM];
1 9 , no date.
Virginia: Nasemond Co., Holland, 1 9, Aug. 11, 1944 (O. Buchholz) [ANS].
Peoria tetradella (Zeller), new combination
Figures 6, 72, 107, 146
Anerastia tetradella Zeller, 1872, pp. 552-553.
Saluria tetradella (Zeller). — Ragonot, 1889, p. 117; 1901, pp. 362-363. — Hulst,
1902, p. 439. — Barnes and McDunnough, 1917, p. 149. — Hampson, 1918,
p. 102. — McDunnough, 1939, p. 36.
Altoona tetradella (Zeller). — Hulst, 1890, p. 207. — Smith, 1891, p. 84.
Diagnosis. — The broad white band on the cubitus and lower outer
angle of the cell marks this species as distinct from other members of
the genus.
Description. — Frons brown; labial palpi with basal segments white,
second and third segments gray on outer sides, white on inner sides
and ventrally; maxillary palpi reaching frons, light brown; antennae
light brown, male subserrate; occiput laterally, patagia, and tegulae
REVISION OF PEORIINAE AND ANERASTIINAE 17
light brown, occiput dorsally and vertex white, legs white on inner
sides, outer sides clothed with white-tipped brown scales.
Forewings v\dth white band anterior to cell tapering to point at
apex, sprinkled with reddish-brown scales, bordered posteriorly by
dark brown band of varying width, often extending to Ai fold, fre-
quently scattered with reddish-brown scales; reddish brown posterior
to Ai fold, A2 white traced and bordered with dark brown scales;
veins traced with white, very prominently so at lower outer angle of
cell; 11 veins; R.2 from cell close to base of or stalked with R3+4; M2+3
stalked. Hind wings light brown, darker toward apex; with 7 veins.
Male genitalia with gnathos bearing apical digitate projection.
Juxta V-shaped. Vinculum flanged and thickened at apex. Valvae
pointed apically; costa sparsely setose; sacculus with fine hairs. Aedea-
gus with vesica bearing single serrate cornutus.
Female genitalia \\dth ovipositor bearing numerous fine setae on
caudal margin, tip moderately setose. Apophyses curved, posterior
tapering to point.
Type. — In the British Museum (Natural History) (from the Zeller
collection) .
Type data. — In the original description Zeller notes: "^^aterland:
Texas (Boll, BeLfrage). Mehrere cf und 9 fing Belfrage in der ersten
Halfte des June, 2 9 am 8. und 9. Juli."
Lectotype male, hereby designated, labeled: "10/6, Bosque Co
Texas, Tetradella Z. Texas [green label in Zeller's handwriting], Zell.
Coll. 1884, 6" genitaha sUde 6-II-1967 J. Shaffer No. 728."
Lectoparatypes : One male and four females, all labeled Bosque
County, Tex. and bearing Zeller's personal type label; male dated 11/6
(June 11), two females dated 7/6, other two females dated 8/7 and 9/7.
Recorded host. — Poaceae: Elymus canadensis L., borer in stem.
From specimen label, three males, Lafayette, Ind.
The host occurs throughout most of the United States and southern
Canada. Hitchcock (1951) notes: "River banks, open ground, and
sandy soil, Quebec to southern Alaska, south to North Carolina,
Missouri, Texas, Arizona, and northern California."
Specimens EXAMINED. — 22 cf , 41 9.
Distribution (Map 6). — California to Texas, northeastward to
Ontario.
UNITED STATES: California: Modoc Co., Canby, 2 &, July 20, 1927
[CU]; county unknown, "Mad R. ab. Maple Cr.," 1 c?, July 28-29, 1927 [CU].
Illinois: Putnam Co., 1 ? , Aug. 6, 1958 (M. O. Glenn) [MOG]; 1 cf, June 21,
1961; 1 &, July 10, 1961; 1 9 , July 12, 1961; 1 9 , July 17, 1961; 1 ? , June 24,
19G2; 1 ?, July 5, 1962; 1 d^, June 23, 1963; 1 ? , July 2, 1964.
Indiana: Tippecanoe Co., Lafayette, 3 cT, 1916 (C. N. Ainslie) [USNM].
Kansas: Thomas Co., (3150 ft.), 1 cf , no date (F. X. Williams) [UK].
18 U.S. NATIONAL MUSEUM BULLETIN 280
Texas: Blanco Co., 1 9 , May [USNM]; 1 ? , June [AMNH]; Id',!?, July
[AMNH]; 1 9, August [AMNH]; 2 9 , no date, [USNM]; Bosque Co., 1 c?,
"11/6" (Zeller collection) [BM]; Burnet Co., 2 cf, no date (F. G. Schaupp)
[USNM]; Collin Co., Piano, 4 9 , July (E. S. Tucker) [USNM]; Dallas Co., 1 9 ,
June [AMNH]; Kerr Co., Kerrville, 1 9, May 30, 1906 (F. C. Pratt) [USNM];
3 9 , May 31, 1906; 1 cT, 2 9 , June 1, 1906; 1 9 , Apr. 11, 1907; 1 9 , no date;
La SaUe Co., CotuUa, 1 9 , May 12, 1906 (Crawford and Pratt) [USNM]; Medina
Co., Sabinal River opposite Hondo, 1 cf , July 1, 1917 [CU]; county unknown,
"Black Jack Spgs.," 1 9 , no date [USNM]; locality unknown, 5 9 [USNM];
3 d', 5 9 [INHS].
CANADA: Ontario: Port Colborne, 1 cT, June 24, 1934 (J. J. de Gryse)
[CNC].
Peoria opacelUi (Hulst), new combination
Figures 11, 73, 108, 147
Anerastia opacella Hulst, 1887, p. 138. — Rindge, 1955, p. 168.
Altoona opacella Hulst, 1887, p. 116; 1890, p. 207. — Smith, 1891, p. 84. — Hulst,
1902, p. 438.
Saluria dichroeella Ragonot, 1889, pp. 113, 117. — Barnes and McDunnough,
1917, p. 149. — McDunnough, 1939, p. 36. [New synonymy.]
Tolima opacella (Hulst). — Ragonot, 1889, p. 117; 1901, p. 341. — Barnes and
McDunnough, 1917, p. 149. — McDunnough, 1939, p. 35.
Altoona dichroeella (Ragonot). — Hulst, 1890, p. 207.
Altoona dichroeella [sic]. — Smith, 1891, p. 84.
Saluria dichroeella Hampson in Ragonot, 1901, p. 363; 1918, p. 101. [New
synonymy, objective.]
Saluria dichroella [sic]. — Hulst, 1902, p. 439.
Commotria opacella (Hulst). — Hampson, 1918, p. 108.
Diagnosis. — Tliis species is distinguished from ail ottier members
of tlie genus with similar venation by the combination of the forewing
being basally darker posterior to the cell and the absence of white
tracing on forewing vein A2.
Description. — Frons obUque, scales brown with white tips; labial
palpi with basal segments white, second and third segments clothed
with white-tipped brown scales dorsally and on outer sides, with light
brown to white scales ventrally and on inner sides; maxUlary palpi
almost reaching frons; antennae brown, laminate in male; occiput
behind eye, patagia, and tegulae brown, vertex behind antennae and
dorsum of occiput light brown; legs w4th tarsi clothed with white-
tipped brown scales.
Forewings brown, darker posterior to cubitus, especially basal half;
aU veins except anals variously traced wdth white; areas between
veins a mixture of white and brown scales; usually solid brown poste-
rior to cubitus, becoming lighter distally. Hindwings light brown,
slightly darker in apical region. Venation as in tetradella.
Male genitalia with gnathos bearing apical digitate projection.
Juxta V-shaped. Vinculum with anterior edge flattened and flanged.
Valvae with costa sparsely setose, sometimes with short apical tooth;
REVISION OF PEORIJNAK AND ANERASTIINAE 19
sacculiis with numerous fine hairs. Aedeagus with vesica bearing a
single serrate cornutus.
Female genitalia with ovipositor tip and caudal margin moderately-
setose, sparsely setose laterally. Posterior apophyses curved on basal
third, base flattened, tear-shaped, tapering to point; anterior slightly
curved.
Types. — A. opacella, in the American Museum of Natural History
(lectotype), in the U.S. National Museum (lectoparatype) ; S. di-
chroeella, in the Museum National d'Histoire Naturelle.
Type data. — ^4. opacella, lectotype male, hereby designated,
Blanco Co., central Texas, collection G. D. Hulst, genitalia slide
No. 3221, Carl Heinrich, June 14, 1946.
Lectoparatype: Texas, one female, Fernald collection; USNM
40080; genitaha slide No. 1107, Carl Heinrich, Apr. 5, 1938.
S. dichroeella, lectotype male, hereby designated, Texas, October,
genitaha slide No. 533, J. Shaffer, May 14, 1965.
Specimens examined, — 47 d^, 46 9 .
Distribution (Map 7). — Arizona, New Mexico, southwestern
Texas to Brownsville.
UNITED STATES: Arizona: Cochise Co., Cochise Stronghold, 1 9, Aug. 30,
1958 (P. Opler) [UCB]; Ramsay Canyon, Huachuca Mts., 1 9 , July 10-15, 1941
(A. B. Klots) [AMNH]; Southwestern Res. Sta., 5 mi. west Portal (5400 ft.),
1 d^, July 7, 1956 (Cazier and Ordway) [AMNH]; 1 9 , July 9, 1956; 1 9 , July 20,
1957 (M. Statham) [AMNH]; 1 d", Aug. 2, 1956 (C. and M. Cazier) [AMNH];
1 cf, July 26-Aug. 3, 1959 (A. B. Klots) [ABK]; Cococino Co., 6/3 mi. EESE.
Flagstaflf (6500 ft.), 1 cf, Aug. 7, 19G4 (J. G. Franclemont) [JGF]; Vail Lake Rd.,
dVz mi. SE. Flagstaflf (6500 ft.), 1 9, July 11, 1961 (J. G. Franclemont) [JGF];
1 9, July 18, 1961 (R. W. Hodges) [JGF]; Pima Co., Baboquivari Mts., 4 9 ,
July 1-15, 1923 (O. C. Poling) [USNM]; 13 c?, 4 9 , July 15-30, 1923; 1 9,
July 27-31, 1923; 3 9 , Aug. 1-4, 1923; 7 9 , Aug. 1-15, 1923; 3 9 , Aug. 1.5-30,
1923; 2 9 , Aug. 24-31, 1923; 1 9 , Sept. 1-15, 1923; 2 9 , July 1-15, 1924; 5 9 ,
Aug. 15-30, 1924; 1 9, Sept. 1.5-30, 1924; 2 9 , no date, (F. H. Snow) [UK];
1 9 , no date (F. H. Snow) [USNM]; Pinal Co., Desert Arboretum, Superior,
1 c?, July 18-21, 1941 (A. B. Klots) [AMNH]; Santa Cruz Co., Santa Rita Mts.,
Madera Canyon (4880 ft.), 1 c?, July 9, 1959 (J. G. Franclemont) [JGF]; 1 cf ,
Aug. 23, 1959.
New Mexico: Eddy Co., White City, 2 c?, May 15, 1950 (E. C. Johnston)
[CNC]; 1 cT, May 16, 1950; 1 d^. May 17, 1950; 1 9 , July 23, 1959 (A. B. Klots)
[ABK].
Texas: Brewster Co., Alpine, 2 d^. May 22, 1950 (E. C.Johnston) [CNC];
Cameron Co., Brownsville, Icf, no date [USNM]; San Benito, 4d', March 16-23
[USNM]; 1 c?, March 24-30; 1 d^, 1 9 , July 16-23; 1 9 , July 24-31; 1 d^, no date;
Hidalgo Co., Mercedes, 2 d^, Aug. 31, 1958 (H. Smalzried) [AMNH]; Jeff Davis
Co., Ft. Davis, 2 0?, May 20, 1950 (E. C. Johnston) [CNC]; Limpia Canyon, 8 d"
1 9 , May 20, 1950 (E. C. Johnston) [CNC]; 1 d^, June 4, 1950.
Discussion. — The species shows close afRnities to tetradella and
floridella, particularly to the latter.
20 U-S- NATIONAL MUSEUM BULLETIN 2 80
Wing maculation in opacella is rather variable, especially with
respect to the degree of white tracing on the veins. The variation is
apparent within local populations and does not appear to be geographic.
Peoria floridelUif new species
Figures 8, 55, 74, 109
Diagnosis. — The species is rather similar to opacella in most re-
spects, but is easily distinguished by the white trace on forewing vein
Description. — Frons conical, scales solid light brown; labial palpi
with basal segments white, second segments reddish brown on outer
sides, third segments brown on outer sides, both white on inner sides;
maxillaiy palpi moderately small, not approaching frons; antennae
light brown, male laminate; occiput behind eye, patagia, and tegulae
light brown, vertex behind antennae bro^vnish white; legs white,
tarsi light brown.
Forewings reddish orange anterior to cell; ground terra cotta;
costa and all other veins traced with white, trace of A2 bordered with
dark bro\vn, cubitus bordered posteriorly with dark brown line.
Hindwings light brown, darker toward apex. Venation as in tetradella.
Male genitalia with gnathos bearing apical digitate projection.
Juxta V-shaped. Valvae with costa sparsely setose, bearing poorly de-
veloped apical tooth; sacculus with numerous fine hairs. Aedeagus
with vesica bearing a single serrate cornutus.
Female unknown.
Types. — In the Canadian National collection (holotype); in the
American Museum of Natural History (one paratype), in the Carnegie
Museum (two paratypes), in the collection of Charles P. Kimball (one
paratype).
Type data. — Holotype, male, Volusia County, Fla., Aug. 2, 1956,
H. A. Denmark; C.N.C. Type No. 9439; genitalia slide No. 412, J.
ShaflFer, Mar. 15, 1965.
Paratypes: One male. Summer Haven, St. Johns County, Fla.,
May 30, 1950 (Fred H. Rindge) [AMNH]; one male, Pellicer Cr.,
13 mi. north of Bunnell, Flagler County, Fla., Apr. 11, 1954 (J.
Bauer), Carnegie Museum Ace. 17023; one male, Apr. 22, 1954,
genitalia slide No, 521, J. Shaffer, May 2, 1965; one male, same data
as holotype, abdomen lost [CPK].
Other specimens examined. — 15 cf .
Distribution (Map 10). — Known only from the east coast of Florida.
UNITED STATES: Florida: Indian River Co., Vero Beach, 13 d', April
1941 (J. R. Malloch), USNM; 1 cf, May 1941; 1 cf, Nov. 15-Dec. 31, 1941.
Discussion. — This form has its closest affinities to opacella, and
differs mainly in size and in that the valvae are rounded and lack the
REVISION OF PEORIINAE AND ANERASTIINAE 21
pointed apex of that species. The most obvious distinctions are those
of size, Jloridella having an alar expanse of 25 mm to 28 mm in the
specimens examined, contrasted with about 18 mm to 22 mm for
opacella, and maculation, the forewing veins all showing a prominent
white trace in the former species. In opacella the white trace is less
distinct and essentially absent on A2. The maxillary palpi are moder-
ately short in the Florida species, not nearly attaining the frons as in
opacella, and the frons itself is clothed with white-tipped brown scales
in the latter species rather than the solid brown scales of Jloridella.
Peoria rostrella (Ragonot), new combination
Figures 2, 75, 110, 148
Saluria rostrella Ragonot, 1887, p. 18; 1889, p. 117. — Hulst, 1890, p. 211. —
Smith, 1891, p. 85. — Ragonot, 1901, p. 363. — Hulst, 1902, p. 439. —
Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36.
Diagnosis. — The combination of reddish-brown forewings, veins
traced with white, R2 well stalked with R3+4, and M2 well stalked with
M3 identify this from within the genus.
Description. — Frons conical, reddish brown; labial palpi with
basal segments white, second and third segments brown on outer
sides, light brown on inner sides; maxillary palpi cylindrical, reaching
frons, light brown; antennae light reddish brown, subserrate in male,
cilia about one-half as long as segment width; occiput, patagia, and
tegulae reddish brown, vertex somewhat lighter.
Forewings reddish brown; all veins traced with white, Ai traced on
distal third; 11 veins; R2 well stalked with R3+4; M2 well stalked with
M3. Hindwings with 7 veins; Sc and Rs stalked; M3 stalked with Cui
for about one-half its length.
Male genitalia with gnathos bearing well-developed medial digitate
process. Juxta U-shaped. Valvae with costa unarmed. Aedeagus with
vesica bearing a single small serrate cornutus.
Type. — In the Museum National d'Histoire Naturelle.
Type data. — Lectotype male, hereby designated, genitalia slide
No. 552, J. Shaffer, July 28, 19G5. The type locality is given in the
original description as California.
Other specimens examined. — 4 cf, 1 9-
Distribution (Map 6) .—Washington southward to central
California.
UNITED STATES: California: Sonoma Co., Forestville, 1 c?", July 10, 1935
(E. C. J.) [CNC]; Tuolumne Co., Twain Harte, 1 ? , Aug. 18, 1960 (M. Lundgren)
[UCB]; 1 cP, July 20, 1961.
Washington: Grant Co., Dry Falls, 1 cf , June 30, 1949 (E. C. Johnston)
[CNC]; county unknown, Berne, 1 d", July 12, 1942 (E. C. Johnston) [CNC].
22 U.S. NATIONAL MUSEUM BULLETIN 280
Discussion. — The forewing color and maculation is very similar
to that of P. johnstoni.
Peoria gemmatella (Hulst), new combination
Figures 7, 76, 111, 149
Spermatophthora gemmatella Hulst, 1887, p. 134. — Rindge, 1955, p. 163.
Cayugag emmalella Hulst, 1888, p. 116; 1890, p. 209. — Smith, 1891, p. 84.— Hulst.
1902, p. 438.
Poujadia gemmatella (Hulst). — Ragonot, 1889, p. 117.
Cayuga bistriatella Hulst, 1890, p. 209. — Smith, 1891, p. 84. — Rindge, 1955, p.
158. [New synonymy.]
Pectinigeria gemmatella (Hulst). — Ragonot, 1901, pp. 356-357. — Barnes and Mc-
Dunnough, 1917, p. 149. — Forbes, 1923, p. 638. — McDunnough, 1939, p.
35.
Peclinigera [sic] bistriatella (Hulst). — Dyar, 1908, pp. 117-118.
Pectinigera [sic] pamponerella Dyar, 1908, p. 117. [New synonymy.]
Pectinigeria bistriatella (Hulst). — Barnes and McDunnough, 1917, p. 149. — Mc-
Dunnough, 1939, p. 35. — Kimball, 1965, p. 250.
Pectinigeria pamponerella Barnes and McDunnough, 1917, p. 149. — McDunnough,
1939, p. 35.
Prophtasia bistriatella (Hulst.) — Hampson, 1918, p. 106.
Saluria gemmatella (Hulst). — Hampson, 1918, p. 100.
Diagnosis. — This species is distinguished within the genus by the
stalked veins M2+3 coupled with the presence of two (subequal)
cornuti in the vesica of the male.
Description. — Frons conical, orange red; labial palpi with basal
segments light reddish brown to white, second and third segments
coral red on outer sides, light brown ventrally; maxillary palpi small,
inconspicuous; antennae with scape coral red anteriorly and on inner
side, white posteriorly and on outer side, shaft brown, red toward
base, laminate in male; lateral regions of occiput, patagia, and tegulae
orange red, vertex white behind antennae.
Forewings brownish red; Sc, Rj, and Eg white traced; Eg, E4, and Mi
black traced; A2 white traced; M3 and cubitals traced mth varying
mixtures of brown, pink, and white; Ai orange traced; cell variable,
often with more brown or orange than other areas; 11 veins; E2 free
from cell; M2+3 stalked. Hindwings light brown, darker in apical area;
7 veins; M3 and Cui stalked.
Male genitalia with gnathos bearing short apical digitate projection,
somethnes very short or absent. Juxta V-shaped. Valvae with costa
bearing blunt tooth at apex, sparsely setose; sacculus minutely pubes-
cent. Aedeagus with vesica bearing pair of subequal cornuti, smaller
one serrate and more heavily sclerotized.
Female genitalia with ovipositor tip moderately setose, caudal
margin finely setose. Posterior apophyses with base somewhat curved,
spear-shaped; anterior very gradually thickened toward base.
REVISION OF PEORIENAE AND ANERASTIINAE 23
Types. — S. gemmatella and C. histriatella, in the American Museum
of Natural History; P. pamponeralla, in the U.S. National Museum.
Type data. — S. gemmatella, lectotype male, hereby designated,
Illinois, collection G. D. Hulst, genitalia slide No. 3223, Carl Hein-
rich, June 14, 1946; histriatella, lectotype male, hereby designated,
Colorado, collection G. D. Hulst, genitalia slide No. 3224, Carl Hein-
rich, June 14, 1946; in the original description Hulst gives as locality
and date: "Taken at San Bernardino, S. Cal., the last week in June";
pamponerella, lectotype male, hereby designated. Chimney Gulch,
Golden, Colo., Aug. 12, 1907, Oslar; USNM 11854; genitalia slide No.
565, J. Shaffer, Nov. 23, 1965; lectoparatype: one male, same data as
lectotype except no genitalia slide, F. Haimbach coll., Lot No. 72,
Brackenredge Clemens Memorial collection, in the Academy of Natu-
ral Sciences, Philadelphia.
Specimens EXAMINED. — 47cr,10 9
Distribution (Map 6). — Colorado east to Iowa, Illinois, southern
Ontario, New Jersey, northward along coast to Maine.
UNITED STATES: Connecticut: New Haven Co., East River, 1?, July 13,
1909 (Charles R. Ely) [USNM]; 1 c?, Aug. 12, 1909; 1 cf, August 1911; 1 cT, July
10, 1912.
Illinois: Cook Co., Arlington Heights, Icf, July 18, 1934 (A. L. McElhose)
[CNHM]; 1 9, Aug. 15, 1936; Chicago, 1 9, Aug. 5, 1916 (E. Beer) [CNHM];
Macon Co., Decatur, 1 c?, Aug. 1, 1890 (W. Barnes) [USNM]; Hancock Co.
Webster, 1 cf, Aug. 12, 1883 [USNM].
Indiana: Lake Co., Hessville, 1 9, July 15, 1905 (A. Kwiat) [CNHM]; 1 cf,
Aug. 1, 1908 (E. Beer) [CNHM].
Iowa: Story Co., Ames, 1 &, June 27, 1912 (G. C. Decker) [USNM]; 1 cf,
July 17, 1932; 1 9, August [USNM]; Woodbury Co., Sioux City, 1 cT, June
25, 1936 (C. N. Ainslie) [UM].
Maine: York Co., Kennebunk, 1 c?, Aug. 11, 1901 (G. H. Clapp) [CM];
1 d", Aug. 22, 1901.
Massachusetts: Nantucket Co., Nantucket, 1 cf, July 21, 1945 (C. P.
Kimball) [CPK]; Barnstable Co., Barnstable, 1 &, Aug. 19, 1952 (C. P. Kimball)
[CPK]; Woods Hole, 3 cf, August 1917 (W. T. M. Forbes) [CU]; Plymouth
Co., Agric. Exp. Sta., East Wareham, 1 cf, Aug. 7, 1964 [CPK]; 1 cf, Aug. 22,
1964.
New Jersey: Burlington, Brown's Mills, 1 cT, August 31, (F. H. Benjamin)
[CU]; New Libson, 1 d', June 12, 1942 (E. P. Darlington) [ANS]; 1 &, Aug. 4,
1942; Whitesbog, 1 cf, June 22, 1940 (E. P. Darlington) [ANS]; 1 cf, Aug. 5
1940; 1 d", Sept. 19, 1936; Ocean Co., Lakehurst, 1 9, Aug. 30, 192? (F. M.
Schott) [USNM]; Lakehurst, Wrangle Brook Rd., 2 cf, June 27, 1954 (J. G.
Franclemont) [JGF]; 2 d", June 27, 1955; 1 cf , Aug. 9, 1955.
New York: Suffolk Co., Orient, 1 d", July 16, 1932 (Roy Latham) [CU];
1 c?, Apr. 21, 1935 [AMNH]; 1 9 , June 14, 193?, [CU]; 1 9 , July 3, 1945; 1 9 ,
Sept. 4, 1945; 1 cT, Sept. 16, 1945; 1 d", July 7, 1947 [AMNH]; 2 d, Aug. 17,
1947; 1 cf , no date [CNC].
CANADA: Ontario: Kent Co., Chatham Lab., 1 d, Aug. 28, 1932 [CNC];
1 &, July 12, 1934; 1 c? , July 18, 1934; 1 cf, July 20 .1935.
24 U.S. NATIONAL MUSEUM BULLETIN 280
Locality unknown: Edgebrook (Prob. Cook Co., Illinois), 1 cT, July 8, 1911
(A. Kwiat) [USNM].
2 &, no date, [INHS]; 2 cT, 1 9 , no date [USNM].
Peoria roseotlnctelUi (Ragonot), new combination
FiGTJKEs 10, 61, 77, 112, 150
Statina roseotinctella Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p. 216
— Smith, 1891, p. 85. — Ragonot, 1901, p. 416. — Hulst, 1902, p. 440. —
Barnes and McDunnough, 1917, p. 150. — Grossbeck, 1917, p. 134. —
Hampson, 1918, p. 60. — McDunnough, 1939, p. 36. — Kimball, 1965,
p. 251.
Calera punctilimbella Ragonot, 1888, p. 50. — Hulst, 1890, p. 217. — Smith,
1891, p. 85. — Ragonot, 1901, pp. 417-418. — Hulst, 1902, p. 441. — Barnes
and McDunnough, 1917, p. 150. — Hampson, 1918, p. 59. — Forbes, 1923,
p. 639. — McDunnough, 1939, p. 36. [New synonymy.]
Statina bifasciella Hampson in Ragonot, 1901, pp. 416-417. — Barnes and
McDunnough, 1917, p. 150. — Hampson, 1918, p. 60. — McDunnough
1939, p. 36. [New synonymy.]
Diagnosis. — Tlie transverse posterior line of dots on tlie forewing
veins delimits this species within the genus.
Description. — Frons conical, deep red, often brown laterally;
labial palpi with basal segments white, second and third segments
deep red on outer sides, ventral third lighter, dorsal third often
brown; maxillary palpi moderately small; antennae with scape deep
red on inner side, white on outer side, shaft light brown, pink near
base, laminate in male; occiput, patagia, tegulae, vertex, and dor-
sum of thorax deep red; legs pink on outer sides, light brown on
inner sides, tarsi often brown.
Forewings orange red, mixed with white anterior to cell ; Sc and Rs
sometimes white traced, Rs often bordered posteriorly by a broad
line of scattered black scales extending from base to apex; transverse
posterior indicated by black spots on Rs, Mi, M3, Cui, Cu2, and A2;
terminal line indicated by dark spots on M3, Cui, Cu2, and Ai fold;
terminal spots visible on lower surface of wing; 10 veins; R2 stalked
with R3+4, M2 fused with M3; M3 stalked with Cui. Hindwings with
6 veins ; M3 and Cui completely fused.
Male genitaUa with medial process of uncus bearing small apical
bulge. Gnathos weakly sclerotized, without apical or lateral processes,
occasionally with very short apical process. Juxta U-shaped. Valvae
with costa sparsely setose, terminating in a dorsally directed hook;
sacculus densely pubescent. Aedeagus with vesica bearing equal pair
of finely serrate cornuti.
Female genitalia with ovipositor moderately setose laterally, caudal
margin with numerous fine setae. Posterior apophyses straight, base
flat and spearlike; anterior curved, tapering gradually to slightly
thickened base.
REVISION OF PEORIINAE AND ANERASTIINAE 25
Types. — S. roseotinctella, in the Museum National d'Histoire
Naturelle; C. ijunctilimhella, in the Zoologisches Museum der Hum-
bolt- Universitat, Berlin; S. hijasciella, in the British Museum (Natural
History) .
Type data. — S. roseotinctella, lectotj^pe male, hereby designated,
Florida, genitalia slide No. 539, J. Shaffer, June 20, 1965.
C. punctilimbella, lectotype female, hereby designated, Carolina;
genitalia slide No. 818.. J. Shaffer, Mar. 20, 1967.
S. hijasciella, lectotype female, hereby designated, labeled as follows:
"11/9; Type; Belfrage Texas 1869; Stainton Coll. 93—134.; Statina
bifasciella type 9 Hmpsn; 9 genitalia slide 1-4-1967 J. Shaffer No.
702."
Specimens examined. — 57 cf, 25 9 .
Distribution (Map 6). — Kansas and eastern Texas to Florida,
southern Florida north to New Jerse}'-.
UNITED STATES: Alabama: Macon Co., LaPlace near Tuskeege, 4 c?,
June 9, 1917 [CU].
District of Columbia: Washington, 19, June 1902 (Aug. Busck) [USNM].
Florida: Alachua Co., Gainesville, 1 ? , June 2, 1927 (J. Speed Rogers)
[CU]; 1 &, June 3, 1927; 19, June 29, 1927; 1 d", July 7, 1927; 13 &, July 8,
1927; 3 c?, July 10, 1927; 36^, July 1927; 19, Apr. 24, 1952 (O. Peck) [CNC];
1 cf , Sept. 3, 1956 (H. A. Denmark) [CPK]; no locality, Icf , Sept. 13, 1956 (H. A.
Denmark) [CPK]; Dade Co., Homestead, 1 9 , Aug. 7, 1963 (D. O. Wolfenbarger)
[CPK]; Id^, Mar. 10, 1964; Ic?, 19, Sept. 29, 1964; Princeton, 19, Apr. 4,
1952 (J. R. Vockeroth) [CNC]; Escambia Co., Pensacola, Icf, May 22, 1961
(Shirley Holls) [CPK]; Hernando Co., Weeki Wachee Springs, Icf, May 28, 1960
(J. F. May) [CPK]; Highlands Co., Archbold Biol. Sta., Id', June 27, 1964 (Jay
C. Shaflfer) [JCS]; HUlsborough Co., Stemper, Id', Aug. 19, 1912 (G. Krautwurm)
[CM]; Id', Sept. 1 1912; Id^, Sept. 6, 1912; Id', Oct. 3, 1912; Ic?, Oct. 6, 1912;
2d', Oct. 30, 1912; Manatee Co., Oneco, Id', Aug. 3, 1953 (Paula Dillman)
[CPK]; Id', Mar. 28, 1957 (J. G. Franclemont) [JGF]; Orange Co., Orlando,
Id', June 29, 1927 (C. C. McBride) [CU]; Id", July 24, 1927; Winter Park,
1 9 , July 4, 1942 (H. T. Fernaid) [USNM]; Volusia Co., Cassadaga, Id', Oct. 18,
1963 (S. V. Fuller) [CPK]; Wakulla Co., Panacea, 2d', Aug. 11, 1926 (C. O.
Handley) [USNM].
Georgia: Bryan Co., Clyde, Id', Sept. 11-12, 1931 (Bradley and Knorr)
[CU].
Kansas: Riley Co., Manhattan, 1 9 , June 11, 1933 (H. L. Nonamaker) [KSU].
Louisiana: Vernon Ph., 1 9 , August (G. Coverdale) (USNM].
Mississippi: Forrest Co., Camp Shelby, 3d', 29, Oct. 1-15, 1944 (CD.
Michener) [AMNH]; Hinds Co., Clinton, 1 9 , June 12, 1960 (Bryant Mather)
[BM].
New Jersey: Ocean Co., Lakehurst, Wrangle Brook Rd., Id", June 30, 1965
(Jay C. Shaffer) [JCS].
North Carolina: Polk Co., Tryon, Id', Aug. 8, 1904 (Fiske) [USNM]; 2d',
Aug. 9, 1904; Id', Aug. 10, 1904; Id', Aug. 11, 1904; Id', no date.
South Carolina: Oconee Co., Cherry Hill Rec. Area, Route 107 (2000 ft.),
Id", Aug. 22, 1958 (J. G. Franclemont) [JGF].
26 U.S. NATIONAL MUSEUM BULLETIN 2 80
Tennessee: Knox Co., Knoxville, 1 9 , June 26, 1916 (G. G. Ainslie), [USNM].
Texas: Blanco Co., 1 9 , no date [USNM]; Liberty Co., Devers, 1 cf, 2 9, June
21, 1917 [CU]; Nueces Co., Corpus Christi, 69, Sept. 25-Oct. 15, 1943 (W. M.
Gordon) [CU].
Peoria johnstonif new species
Figures 1, 78, 113, 151
Diagnosis. — Among the three other species of Peoria with similar
venation, johnsioni differs from approximella and luteicostella in
lacking a well-developed white band anterior to the cell, and from
bipartitella in having the forewings a rather uniform brownish orange.
Either the presence of two cornuti on the vesica of the aedeagus or
the absence of medial or lateral processes on the gnathos will de-
limit males of johnstoni from the above three species.
Description. — Frons conical, reddish brown; labial palpi with
basal segments white, second and third segments reddish brown
dorsally and on outer sides, lighter ventrally and on inner sides;
maxillary palpi small; antennae light brown, scape light pink an-
teriorly, male subserrate; occiput, patagia, and tegulae reddish
brown, vertex somewhat lighter.
Forewings brownish orange; Sc, radials, cubitals, M3, and A2
white traced, costa traced with narrow inconspicuous white line;
brownish orange on lower surface. Venation as in P. bipartitella.
Male genitalia Avith medial process of uncus bearing small ventral
bulge at apex. Gnathos weakly sclerotized, without apical or lateral
processes. Juxta U-shaped. Valvae with costa sparsely setose; sacculus
densely setose. Aedeagus with vesica bearing pair of ovate subserrate
cornuti.
Female genitalia mth ovipositor moderately setose along caudal
margin. Posterior apophyses straight, base flat and shaped like
parallelogram with short side horizontal; anterior slightly curved,
tapering gradually to thickened base.
Type. — In the Canadian National collection.
Type data. — Holotype, male. Fort Davis, Jeff Davis County,
Tex., May 20, 1950, E. C. Johnston; C.N.C. Type No. 9440; genitalia
slide No. 591, J. Shaffer, Mar. 24, 1966.
Paratypes: Five males, same data as holotype, no genitalia slides
prepared.
Specimens examined. — 16 cf, 15 9 •
Distribution (Map 7). — Mexican Highland Province; southern
Arizona and New Mexico to southwestern Texas.
UNITED STATES: Arizona: Pima Co., Baboquivari Mts., 39, Sept. 1-15,
1923 (O. C. Poling) [USNM]; I9, Oct. 1-15, 1923; 19, Apr. 15-30, 1924; 3cf
29, Oct. 1-15, 1924; Icf, 2 9, Oct. 15-30, 1924; Pinal Co., Oracle, Id^, June 5,
REVISION OF PEORIINAE AND ANERASTIINAE 27
1935 (Grace II. and John L. Speny) [AMNII]; Santa Cruz Co., Pena Blanca
(3950 ft.), Icf, May 31, 1963 (J. G. Franclemont) [JGF]; county unknown
(southeastern Arizona), Huachuca Mts., Icf, June 1, 1935 (J. A. Comstock)
[CNC]; 1 9 , May 28, 1935 (Grace H., and John L. Sperry) [CNC]; 1 ? , June 1,
1935 [AMNH].
New Mexico: Eddy Co., White City, Icf, May 15, 1950 (E. C. Johnston)
[CNC].
Texas: Brewster Co., Alpine, 49, May 22, 1950 (E. C. Johnston) [CNC];
Jeff Davis Co., Limpia Canyon, 8 cf, May 20, 1950 (E. C. Johnston) [CNC].
Peoria santaritella (Dyar), new combination
Figures 5, 79, 114, 152
Ollia santaritella Dyar, 1904, p. 108. — Barnes and McDunnough, 1917, p. 149. —
McDunnough, 1939, p. 36.
Diagnosis. — Ttie forewings are white anterior to ttie cubitus, red
between the cubitus and the Ai fold, and light pink posterior to the
fold; the combination of these features serves to delimit the species
within the genus.
Description. — Frons conical, blood red; labial palpi with basal
segments white, second and third segments red on outer sides, white
ventrally and on inner sides; maxillary palpi small; antennae with
scape red, shaft brown, male serrate and fasciculate; occiput deep red
behind eye, vertex posterior to antennae and dorsal area of occiput
white, patagia and tegulae pmk; legs white, tarsi light brown, forelegs
pink on inner sides.
Forewings white anterior to cubitus, a few scattered red scales
anterior to cell, costal margin bordered with red near base; red
between cubitus and Ai fold, bounded distally by M3 and Cu2; light
pink posterior to red line; underside with red showing through faint-
ly; 10 or 11 veins; K2 free from cell; M2+3 stalked for about half its
length or completely fused, both conditions about equally common,
sometimes (rarely) difTering on right and left wings of same specimen.
Hindwings with M3 and Cui stalked.
Male genitalia with gnathos bearing short apical digitate projection.
Juxta V-shaped. Valvae with costa terminating in a blunt tooth,
cucculus not projecting beyond costa; sacculus with fine hairs. Aedeag-
us with vesica bearing single serrate cornutus.
Female genitalia with ovipositor tip sparsely setose, caudal margin
with numerous fine setae. Posterior apophyses with base tapering to
point, not flattened or broadened, curved near base; anterior tapering
gradually to slightly thickened base.
Type. — In the U.S. National Museum.
Type data. — Holotype male, Santa Kita Mountains, Arizona,
'7. 6," E. A. Schwarz; USNM 7893 ; genitalia slide No. 569, J. ShaflFer,
285-934—68 3
28 U.S. NATIONAL MUSEUM BULLETIN 280
Nov. 23, 1965. In the original description Dyar gives the date of
capture as June 7.
Specimens examined.— 4 cf, 21 9 .
Distribution (Map 7). — Northern extension of Sierra Madre
Occidental into southeastern Arizona.
UNITED STATES: Arizona: Cochise Co., Paradise, Chiricahua Mts., 19,
July 3, 1954 (Cazier and Gertsch) [AMNH]; Southwestern Res. Sta. (5400 ft.),
5 mi. west Portal, 1 9 , May 28, 1956 (Cazier and Ordway) [AMNH]; 1 9 , May
25, 1958; 49, June 11, 1958 (W. J., and J. W. Gertsch) [AMNH]; 29, June 12,
1958; 19, June 16, 1958 (M. A. Cazier) [AMNH]; 19, May 27, 1960 (Carl W.
Kirkwood) [CPK]; 19, June 2, 1960; Id", May 10, 1961 (Gertsch and Cazier)
[AMNH]; Pima Co., Santa Rita Mts., Madera Canyon (4400 ft.), 19, June 12,
1963 (J. G. Franclemont) [JGF]; Santa Cruz Co., Santa Rita Mts., Madera
Canyon (4800 ft.), 1 9 , June 19, 1960 (David A. Wallesz) [CU]; (5800 ft.), 1 9 ,
June 24, 1960; Madera Canyon (4800 ft.), Icf, May 26, 1963 (J. G. Franclemont)
[JGF]; Icf , 19, June 19, 1963; 1 9 , June 13, 1960; 1 9 , June 3, 1963; (5600 ft.),
Icf , June 11, 1963; 1 9 , June 18, 1963; Pena Blanca (3950 ft.), 1 9 , June 7, 1963
(J. G. Franclemont) [JGF]; county unknown, Huachuca Mts., 19, June 2, 1935
(Grace H., and John L. Sperry) [AMNH].
Peoria holoponerella (Dyar), new combination
Figures 3, 4, 80, 115, 153
Ollia holoponerella Dyar, 1908, p. 117. — Barnes and McDunnough, 1917, p.
149. — McDunnough, 1939, p. 36.
Diagnosis. — The light buff forewing with its conspicuous white
band anterior to the cell delimits this species within Peoria. A similar
white band marks the forewings of approximella and luteicostella, but
in both of these species the forewings are marked with red and lack
vein M2.
Description. — Frons conical, brown; labial palpi with basal
segments white, second and third segments brown on outer sides,
light brown to white ventrally and on inner sides; maxillary palpi
small; antennae silver, male shaft serrate and fasciculate; occiput
behind eye, patagia, and tegulae light brown, vertex and dorsum of
occiput white; legs white, tarsi brown.
Forewings with white band anterior to cell, bordered anteriorly
near base with dark brown, posteriorly by black line; ground light
buff; veins traced with black scales. Hindwings light brown. Venation
as in gemmatella.
Male genitalia with gnathos bearing medial digitate projection.
Juxta V-shaped. Valvae with costa unarmed, bearing scattered setae;
sacculus with fine hairs, Aedeagus with vesica bearing single round
serrate cornutus.
Female genitalia with ovipositor slender, caudal margin rather
heavily setose, bare laterally, Apophyses straight, posterior hooked
upward and coming to point at base; anterior with thickened base.
REVISION OF PEORIINAE AND ANERASTIINAE 29
Type. — In the U.S. National Museum.
Type data. — Holotype male, San Bernardino Ranch, Cochise
County, Ariz., 3750 ft., August, F. H. Snow; USNM 11855; genitalia
slide No. 568, J. Shaffer, Nov. 23, 1965.
Specimens examined. — 18 cf, 2 9.
Distribution (Map 7). — Southwestern United States,
UNITED STATES: Arizona: Cochise Co., Douglas, 2 d', August (F. H.
Snow) [UK]; San Bernardino Ranch (3750 ft.), 5 cf , 1 ? , August (F. H. Snow)
[UK].
California: Inyo Co., Furnace Creek, Death Valley, 3 cf, Apr. 21, 1942
(George WUlett) [LACM]; Triangle Springs, Death VaUey, 1 ?, Apr. 14-15,
1942 (George Willett) [LACM]; 1 cT, Apr. 17, 1942.
New Mexico: Sandoval Co., Jamez Springs (6200 ft.), 4 cf, July 16, 1950
(T. Cohn, P. Boone, M. Cazier) [AMNH].
Texas: Jeff Davis Co., Limpia Canyon, 2 cf , May 20, 1950 (E. C. Johnston)
[CNC].
Utah: Locality unknown, 1 cf , July 1900 (Poling) [USNM].
Discussion. — The five specimens from Death Valley entirely lack
brown or black markings on the forewings.
Peoria approximella (Walker)
Figures 14, 59, 81, 116, 154
Eurhodope approximella Walker, 1866, p. 1722.
Anerastia haematica Zeller, 1872, pp. 555-556. — Grote, 1879, p. 12, pi. 2, fig. 14
Nephopteryx roseatella Packard, 1873, p. 270.
Peoria haematica (Zeller). — Ragonot, 1889, p. 117. — Hulst, 1890, pp. 213-214.
— Smith, 1891, p. 85.
Peoria approximella (Walker). — Ragonot, 1901, p. 386. — Hulst, 1902, p. 439.
— Barnes and McDunnough, 1917, p. 149. — Grossbeck, 1917, p. 134. —
Forbes, 1923, p. 638. — McDunnough, 1939, p. 36. — Kimball, 1965, p. 251.
Hypsotropa approximella (Walker). — Hampson, 1918, p. 77.
Hypsotropa cremoricosta Hampson, 1918, p. 74. — McDunnough, 1939, p. 36.
[New synonymy.]
Diagnosis. — The presence of both a prominent white band anterior
to the cell and a fainter one posterior to vein A2 marks this species
as distinct within the genus. The gnathos is also unique in that it
bears a pair of short lateral finger-like processes.
Description. — Frons conical, blood red to orange red; labial palpi
with basal segments varying white to light pink, second and third
segments somewhat rough scaled, blood red to orange red dorsaUy
and on outer sides, usually paler ventrally and on inner sides ; maxillary
palpi small; antenna with scape red on anterior and inner sides, light
pink to white on posterior and outer sides, shaft light brown, pink
toward base, laminate in male; occiput laterally, patagia, and tegulae
orange red, vertex light brown to white.
Forewings with broad white band anterior to cell, narrowing to
point just short of apex, bordered anteriorly with black near wing
30 U.S. NATIONAL MUSEUM BULLETIN 280
base, bordered posteriorly with black, fading into blood red triangular
area extending to anal vein; brownish white posterior to anal vein;
anal and costal bands joined near wing base; fringe white. Venation
as in bipartitella.
Male genitalia with gnathos weakly sclerotized, bearing pair of short,
lateral, subapical projections. Juxta scoop-shaped. Valvae with costa
terminating in free spine not attaining apex of bluntly rounded cuc-
culus; sacculus minutely pubescent. Aedeagus with vesica unarmed.
Female genitalia with ovipositor tip moderately setose, caudal mar-
gin with numerous fine setae, sparsely setose laterally. Apophyses uni-
form, curved at base; posterior tapering to point at base, base not
flattened.
Types. — E. approximella, A. haematica, and H. cremoricosta in the
British Museum (Natural History) ; A^. roseatella, in the Museum of
Comparative Zoology (Harvard University).
Type data. — E. approximella, Walker reports: "North America.
From Mr. Carter's collection." Type examined by Mr. Paul Whalley.
A. haematica, Zeller in his description noted: ". . . New York und
aus Massachusetts . . . ." Type examined by Mr. Paul Whalley.
A^. roseatella, two specimens, both from Massachusetts, and both
Type No, 14282, one from Dorchester, the other hereby designated as
lectotype and labeled male genitalia slide No. 540, J. Shaffer, June 24,
1965. In his description Packard indicated that he had two males and
noted: "Length of fore wing .37 of an inch. Dorchester, Mass. (F. G.
Sanborn)."
H. cremoricosta, lectotype male, hereby designated, labeled as fol-
lows: "Type; Colorado Springs, Colorado. 97-278 June-Aug 1896;
Hypsotropa cremoricosta type cf . Hmpsn.; cf genitalia slide 1-4-1967
J. Shaffer No. 703."
Specimens examined. — 562 cf , 302 9 .
Distribution (Map 5). — Southern Canada west to Alberta, south-
ward in mountains to Utah and New Mexico, east to Tennessee and
Georgia, north to Nova Scotia; absent from Florida and the Gulf
Coastal Plain.
UNITED STATES: Colorado: El Paso Co., Rock Creek Canyon, Colorado
Springs, 2 c?, Aug. 10, 1957 (Margot May) [CPK]; 1 d^, Aug. 13, 1957; 4 cf , Aug.
14, 1957; 1 cf , Aug. 23, 1957; 2 cf, June 27, 1960; 1 cf , June 28, 1960; 1 d^, July
15, 1960; 1 c?, July 9, 1961; 1 d", July 19, 1961; 6 cT, June 30, 1963; 2 cf, July 1,
1963; 1 cf, July 3, 1963; 2 <f, July 14, 1963; 1 cf, July 17, 1963; 1 d', July 18,
1963; 1 cf , Aug. 11, 1963; 1 c?, June 27, 1964; 3 cf , June 28, 1964; vie. Colorado
Springs, 1 9 , July 7, 1939 [AMNH]; Jefferson Co., Chimney Gulch, Golden, 1 cf ,
Sept. 3, 1908 (Oslar) [ANS]; Larimer Co., Estes Park, 1 cf , Aug. 29, 1936 (Grace
H. and John L. Sperry) [AMNH]; Teller Co., Big Springs Ranch, Florissant, 1 cf,
July 13, 1960 (Thomas C. Emmel) [LACM]; 1 cf , July 21, 1960; 2 d', July 23,
1960; 2 cf , July 26, 1960; 1 d", July 29, 1960; 1 cf , July 30, 1960; 1 cf, July 31,
1960; 2 d", Aug. 2, 1960; 1 d', Aug. 3, 1960, 1 d', Aug. 4, 1960; 1 d', Aug. 8, 1960;
REVISION OF PEORIINAE AND ANERASTIINAE 31
1 cT, Aug. II, 1960; 1 d', Aug. 12, 1960; 1 d", Aug. 14, 1960; locality unknown, 1
cf , no date [AMNII].
Connecticut: Windham Co., South Shore, Killingly Pd., 1 ?, July 25, 1930
(A. B. Klots) [AKB]; Putnam, 1 d, Aug. 14, 1954 (A. B. Klots) [ABK]; 1 d,
July 1, 1960; 1 ?, July 20-25, 1961; 1 9, July 25-30, 1961.
District of Columbia: 1 cf, Aug. 17, 1899 (August Busck) [USNM]; 2 ?,
July 1901; 2 9, June 1902; 1 9, June 1907; 1 9, June.
Georgia: Charlton Co., 1 9 , June 14, 1946 (Otto Buchholz) [ANS].
Illinois: Cook Co., Arlington Heights, 1 cf, June 20, 1930 (A. L. McElhose)
[CNHM]; 1 9 , June 23, 1930; I 9 , June 24, 1930; I 9 , July 29, 1930; Chicago,
3 cf, June 1900 [USNM]; I 9, July 14, 1904; 1 cf, July 8, 1906 (W. J. Gerhard)
[CNHM]; I cf, June 1912 (A. Kwiat) [CNHM]; Jackson Co., Murphysboro,
1 9, Aug. 10, 1932 (W. J. Gerhard) [CNHM]; McHenry Co., Algonquin, 1 cf,
July 6, 1903 (W. A. Nason) [INHS]; 1 9, July 16, 1904; I cf, June 8, 1906;
McLean Co., Normal, 1 9, June 19, 1884 [INHS]; Macon Co., Decatur, I cf
June 8-15 (Barnes) [USNM]; Putnam Co., 1 cf. May 31, 1934 (M. O. Glenn)
[MOG]; I d, June 21, 1954; I cf, Sept. 5, 1955; 1 cf, June 8, 1959; 1 9, July
19, 1962; 1 9 , June 25, 1964; 1 9 • July 15, 1964; Magnolia, 1 cf , June 29, 1932
(Murray O. Glenn) [USNM]; locality unknown, 2 cf, no date [USNM].
Indiana: Lake Co., Hessville, 1 cf. May 14, 1905 (A. Kwiat) [CNHM];
1 cf, June 17, 1905; 1 cf , June 17, 1912.
Iowa: Johnson Co., Iowa City, 1 9, July 12, 1898 (Wickham) [AMNH];
1 9 , July 27, 1898; Story Co., Ames, I cf , July 9, 1915 (G. G. Ainslie) [USNM];
Woodbury Co., Sioux City, I 9 , July 23, 1923 (C. N. Ainslie) [UM]; locality
unknown, 1 cf , no date (C. P. Gillette) [USNM].
Kansas: Pottawatomie Co., Onaga, 1 cf, June 17, 1901 (Crevecoeur) [KS];
1 cf, July 7, 1901; Riley Co., I cf, June 22 (F. Marlatt) [KS]; 1 9, July; 1 cf,
July 28.
Kentucky: Rockcastle Co., 1 cf, June 24, 1955 (R. Beebe) [CNC].
Maine: Franklin Co., Rangeley, 1 cf, June 23, 1938 (V. H. dos Passos)
[AMNH]; 1 cf , July 9, 1938; Knox Co., Rockport, 1 cf , July 10, 1941 (Morgan
Hebard) [ANS]; Penobscot, Orono, 2 9 , no date, [CU]; York Co., Kennebunk-
port, 2 cf, 1 9, July (G. H. Clapp) [CM]; 3 cf, 1 9, August.
Maryland: Montgomery Co., Plummer Isl., I 9, May 1903 (August Busck)
[USNM]; 1 cf, June 1903; 1 9, Aug. 2, 1905 (H. S. Barber) [USNM]; 2 9,
Aug. 8, 1905 (Barber and Schwarz) [USNM]; Prince Georges Co., Beltsville,
1 cf , June 10, 1939 (L. J. Bottimer) [ANS].
Massachusetts: Barnstable Co., Woods Hole, 1 9, August 1917 (W. T. M.
Forbes) [CU]; Barnstable, 1 cf , July 5, 1949 (C. P. Kimball) [CPK]; 1 cf , July 6,
1949; 4 cf , July 7, 1949; 1, June 24, 1958; 1 cf , Aug. 1, 1958; 1 9 , June 18, 1962;
West Barnstable, 3 cf , July 15, 1949 (C. P. Kimball) [CPK]; I cf , July 16, 1949;
Dukes Co., Martha's Vineyard, 1 cf, June 17, 1931 (F. M. Jones) [ANS]; 1 9,
July 22 [CPK]; Essex Co., Magnolia, 1 cf , June 29, 1900 (Holland) [CM]; Hamp-
den Co., Chicopee, 1 9 , no date (F. Knab) [USNM]; Nantucket Co., Nantucket,
1 cf, July 9, 1941 (C. P. Kimball) [CNC]; Plymouth Co., Agric. Exp. Sta.,
East Wareham, 2 cf , Aug. 7, 1961 [CPK]; 1 cf , July 3, 1962; 1 cf , July 19, 1962;
1 cf, Aug. 17, 1964; 1 cf , Aug. 18, 1964; I cf , Aug. 20, 1964; Suffolk Co., Arling-
ton, 1 cf, June 25, 1920 (C. S. Anderson) [BPI]; Boston, 1 9 , no date [AMNH];
Worcester Co., Paxton, 2 cf, July 3, 1945 (W. T. M. Forbes) [CU]; locality
unknown, 1 cf , June 27, 1868 (BM, ex Zeller coll.].
Michigan: Kalamazoo Co., Gull Lake Biol. Sta., 1 9 , July 21, 1959 (Roland
L. Fischer) [CNC].
32 U.S. NATIONAL MUSEUM BULLETIN 280
Minnesota: Cass Co., Icf, June 29, 1937 (L. W. Orr), [UM]; Icf, June 30,
1937; Ic?, July 2, 1937; Id", July 11, 1937; Cass Lake, Id', June 18, 1934 (A. A.
Granovsky) [UM]; 19, July 17, 1936 (R. H. Daggy) [UM]; Id, Aug. 3, 1936;
1 d, Aug. 14, 1936; Hennepin Co., Saint Anthony Park, 1 9 , July 12, 1900 [UM];
Olmsted Co., 1 9 , no date [UM]; Polk Co., Crookston, Icf, July 18, 1935 (D. G.
Denning) [UM]; Id, June 27, 1937; Id, July 4, 1937; Id, July 23, 1939 (A. W.
Buzicky) [UM]; Crookston, Red Lake River, Id, July 9, 1935 (D. G. Denning)
[UM]; Pope Co., Sedan, Id, July 7, 1927 (D. Denning) [UM]; Ramsey Co., St.
Paul, Id, July 21, 1927 (Carl T. Schmidt) [UM]; county unknown, Itasca Park,
LaSalle Valley, Id, June 21, 1940 (C. E. Mickel) [UM]; Id, June 23, 1940;
Id, June 24, 1940; Id, July 7, 1940; 2d, 19, July 9, 1940; Id, July 12, 1940;
Id, July 14, 1940.
Missouri: Clinton Co., Lathrop, 19, June 22, 1955 (E. C. Becker) [CNC];
city of St. Louis, 5d, 59, June 1904 (August Busck) [USNM]; Id, July 1904;
Icf, 19, Aug. 6, 1906 (McElhose) [CNHM].
Nebraska: Sioux Co., Canon region north of Harrison, 4d, July 19, 1917
(R. A. Leussler) [USNM, AMNH].
New Hampshire: Coos Co., Randolph, 19, Aug. 16, 1936 (Grace H. and
John L. Sperry) [AMNH]: Grafton Co., Franconia, 2c?, no date [AMNH];
Rockingham Co., Hampton, Id, June 27, 1913 (S. Albert Shaw) [USNM];
1 9 , July 18, 1914.
New Jersey: Bergen Co., Bergenfield, 1 9 , July 3 (F. M. Schott) [AMNH];
Oakland, 19, July 25, 1947 (C. P. Kimball) [CPK, CNC]; 19, July 26, 1947;
19, July 27, 1947; Id', Aug. 8, 1948; Id', Aug. 9, 1948; Aug. 10, 1948; Id',
Aug. 12, 1948; Id', Aug. 13, 1948; Ramsey, Id', 39, July 12, 1935 [AMNH];
Id', July 29, 1935 [AMNH]; Burlington Co., New Lisbon, Id', July 14, 1930
(E. P. Darlington) [ANS]; Id', July 1, 1932; 19, July 19, 1932; Id', July 29,
1931; Id', July 20, 1942; Essex Co., Montclair, Id', June 15 (W. D. Kearfott)
[USNM]; Id', July 10; Id', July 18; Newark, Icf, June 13, 1898 [USNM];
Middlesex Co., Dayton, 2d', June [USNM]; New Brunswick, Id', June 11
[AMNH]; Id, June 1933 [AMNH]; Monmouth Co., Como, 19, July 1919
[AMNH]; Morris Co., Mendham, Id', 19, June 20, 1934 [AMNH]; Ocean Co.,
Wrangle Brook Road, Lakehurst, Id*, June 17, 1955 (J. G. Franclemont) [JGF];
Union Co., Elizabeth, 19, "8-13" [CNHM]; county unknown, 1 9 , no date
[AMNH].
New Mexico: McKinley Co., McGaflFey, Zuni Mts. (7500 ft.). Id', July 22,
1962 (E. and I. Munroe) [CNC]; 2d', July 23, 1962; Sandoyal Co., Frijoles
Canyon, Bandelier Nat. Mon. (6050 ft.). Id', July 18, 1962 (E. and I. Munroe)
[CNC]; county unknown. Id*, June 30, 1935 (Grace H. and John L. Sperry)
[AMNH].
New York: Cattaraugus Co., Otto, 1 9 , July 17, 1882 [CU]; Clinton Co.,
Peru, 2d', 1 9 , June 21, 1915 [CU]; 1 9 , July 1, 1916; Id', July 2, 1916 (Mix and
Everett) [CU]; Delaware Co., 4d', Aug. 3, 1962 (Robert SUberglied) [JCS]
4d', 9 , Aug. 5, 1962; Erie Co., East Aurora, Id', July 20, 1912 (E. Wild) [CU]
North Collins, 1 d', 2 9 , June 27, 1939 (W. T. M. Forbes) [CU]; 1 d, June 28-29^
1939; Sardinia, Id', July 24, 1940 (L. R. Rupert) [CU]; 19, July 27, 1940
Jefferson Co., Picton Island, Clayton, 1 d, June 22, 1958 (B. Heineman) [AMNH]
Lewis Co., 1 d, June (W. W. Hill) [UK]; Monroe Co., 1 9 , July 11, 1945 [CNC]
1, June 27, 1947 [CNC]; 1 9 , July 9, 1947 [CNC]; 1 9 , July 11, 1947 [CNC]; 1 d
July 21, 1947 [CPK]; 19, June 28, 1948 [CNC]; 19, June 29, 1948 [CNC]
Id', July 1, 1948 [CPK]; Rochester, Id', June 26, 1932 [ABK]; New York City
West Farms, 1 9 , no date (J. Angus) [AMNH]; Orange Co., Tuxedo, 1 9 , June 28;
1928 [AMNH]; 1 9 , July 10, 1928 [AMNH]; Oswego Co., Minetto, Id, June 19,
REVISION OF PEORIESTAE AND ANERASTIINAE 33
1938 (W. T. M. Forbes) [CU]; 2c?', 29, June 21, 1938; Sd', 99, June 22, 1938;
2^", 89, June 24, 1938; 19, Aug. 22, 1938; Otsego Co., Oneonta, 19, July 4,
1935 (H. K. Townes) [CU]; 1 9 , July 5, 1935; Saint Lawrence Co., Oswegatchie,
ScT, Aug. 3, 1926 (B. K. Smith) [CU]; Suffolk Co., Flanders, IcT, Sept. 1, 1946
(Roy Latham) [CU]; Huntington, 19, July 25, 1925 (F. M. Schott) [USNM];
Montauk, 19, Sept. 8, 1946 (Roy Latham) [CU]; Orient, 19, July 28, 1931
(Roy Latham) [CU]; Icf, July 6, 1932; Icf, Aug. 3, 1932; 19, Aug. 4, 1932;
1 o^, Nov. 6, 1932; 1 cf , June 22, 1934; 1 cT, June 22, 1946 (Roy Latham) [AMNH];
Id', June 21, 1947; 19, June 23, 1947; Id', July 21, 1947; Id', June 12, 194?;
Shelter Island, 19, Sept. 14, 1946 (Roy Latham) [CU]; Southold, 19, July 1,
1934 (Roy Latham) [CU]; Id', July 24, 1946; Tompkins Co., Ithaca, 2d, July 4,
1882 [CU]; 3d', 69 , July 8, 1882 [CU]; Id', July 14, 1885 (E. H. Sargent) [CU];
1 9 , June 13. 1893 [CU]; 1 9 , July 7, 1893 [CU]; 1 d, June 1913 [CU]; 1 9 , July 2,
1914 [CU]; 1 9 , June 26, 1915 [CU]; 1 d, June 15, 1919 [CU]; 1 d, June 16, 1919
[CU]; 1 9 , June 21, 1919 [CU]; 2 9 , June 21, 1922 [CU]; 1 d, June 29, 1922 [CU];
29, June 30, 1922 [CU]; Id', 19, July 2, 1922 [CU]; 19, July 4, 1944 [CU];
Id', July 7, 1922 [CU]; Id', July 2, 1925 [CU]; Id', July 2, 1925 [CU]; 19,
July 9, 1925 [CU]; 1 9 , July 25, 1926 (Bolton K. Smith) [CU]; 1 9 , July 5, 1928
[AMNH]; Id, 19, June 16, 1929 [CU]; Id', 19, July 15 [CU]; 29, June 16,
1939 [CU]; Six Mile Creek, Ithaca, Id', June 14, 1940 (J. G. Franclemont) [CU];
1 d, June 4, 1957; Id', 19, June 14, 1957; 1 9 , July 6, 1957; McLean Bogs Res.,
1 d, July 4, 1924 [CU]; 2 9 , July 22, 1924; 1 9 , July 28, 1924; 1 d, July 6, 1946
(J. G. Franclemont) [CU]; 19, June 30, 1954 (J. G. Franclemont) [JGF]; 19,
July 27, 1954; Id', July 28, 1954; Varna, Id', May 6, 1946 [CNC]; Id', June 17,
1946 [CNC]; Id', June 22, 1946 (E. G. Munroe) [CNC]; Id', June 25, 1949
(Travassos and Rabella) [CNC]; West Chester Co., Katonah, Id', July [ANS].
North Carolina: Polk Co., Tryon, Id', no date (Fiske) [USNM]; Robeson
Co., Maxton, Id, May 23, 1944 (A. B. Klots) [AMNH]; Transylvania Co.,
Brevard, 19, June 13, 1942 (W. J. Westfall, Jr.) [CU]; 29, July 19, 1942; 2d',
July 23, 1942; 1 9 , July 31, 1942.
Ohio: Athens Co., Athens, 1 9 , June 27, 1931 (W. C. Stehr) [UM]; Hamilton
Co., Cincinnati, Id', July 5, 1904 (Annette F. Braun) [ANS].
Pennsylvania: Adams Co., Arendtsville, 1 d', 1 9 , July 9, 1921 (S. W. Frost)
[CU]; Allegheny Co., 2 9 , July 1900 [CM]; 1 d, July 26 [CM]; Oak Station, 1 9 ,
Aug. 2, 1908 (Fred MarloflF) [CNHM]; 1 d, June 23, 1909; 1 d, June 4, 1911 (Fred
Marloff) [UK]; 1 d, June 14, 1911 (Fred Marloff) [CNHM]; 1 d, June 15, 1911
(Fred Marloff) [UK]; 1 9, June 24, 1911 (Fred Marloff) [CNHM]; Pittsburgh, 2
d, June 13, 1905 (Henry Engel) [CM]; 1 9 , June 19, 1905 (Henry Engel) [CM]; 1
d, June 23, 1905 (Henry Engel) [USNM]; 1 9, July 9, 1905; 1 9, July 9, 1905
(Henry Engel) [CM]; 1 d, June 4, 1906 (Henry Engel) [CM]; 1 9 , June 17, 1906;
1 d', 1 9 , June 18, 1906; 1 d, June 20, 1906; 1 9 , June 26, 1906; 1 9 , June 27,
1906; 1 d, June 29, 1906; 1 d, July 1, 1906; 1 d, July 6, 1906; 1 9 , July 8, 1906
(Henry Engel) [USNM]; 1 d, July 10, 1906; 1 9 [CM]; 2d', July 15, 1906 (Henry
Engel) [CM]; 1 9 , Aug. 3, 1906; 2 9 , July 15, 1907 (Hugo Kahl) [CM]; 1 d, July
16, 1907; 1 d, July 19, 1907; 1 d, June 23, 1908 (Henry Engel) [CM]; 1 9 , June
24, 1908 (F. W. Friday) [LACM]; 1 d', 1 9, June 27, 1909; 1 9, July 15, 1909
(Hugo Kahl) [CM]; 2 d', 1 9, July 8, 1910 (Henry Engel) [CM]; 1 d, June 13,
1911 (Hugo Kahl) [CM]; 1 9, June 15, 1911; 2 d, June 17, 1911; 1 9, Aug. 7,
1911; 1 d, July 12, 1912; 2 d, July 2, 1914 [CM]; 1 d, July 22, 1914 [CM]; 1 d,
July 24, 1916 (Henry Engel) [CM]; 1 d, July 28, 1917 [CM]; 1 d, July 31, 1917
[CM]; 1 9 , Aug. 17, 1917 [CM]; 1 d, June 6, (Henry Engel) [CM]; 1 d, June 17;
1 d, June 30; 1 d, July 3; 3 9 , July 11; 1 9 , July 12; 1 d', 1 9 , July 14; 1 9 ,
July 15; 1 d, July 20; 1 d, July 24; 1 d', 1 9 , July 25; 1 9 , August 2; 1 d, Au-
34 U.S. NATIONAL MUSEUM BULLETIN 280
gust 3; Sharpsburg, 3 d", June 15-21 (Sweadner) [CM]; 2 cf, June 22-30; 1
9, July 8-14; 6 cf, 1 9, July 15-21; 1 cf, August 15-21; Beaver Co., New
Brighton, 1 9, July 2, 1902 (H. D. Merrick) [USNM]; 1 d', May 31, 1903; 1 cf,
July 11, 1904; 1 9 , June 26, 1905 (H. D. Merrick) [UK]; 1 9 , July 20, 1905 (H.
D. Merrick) [USNM]; 1 d", June 13, 1906 (H. D. Merrick) [UK]; 1 cf, Julyl3,
1906 (H. D. Merrick) [USNM]; Berks Co., Sinking Spring, 1 9 , June 29, 1952 (H.
C. Moyer) [CNC]; Blair Co., Tyrone, 1 9, July 25, 1917 (J. G. Sanders) [BPI];
Bucks Co., Langhorne, 1 9 , June 21, 1924 (F. Haimbach) [ANS], Butler Co.,
Butler, 5 &, June L5, 1945 (Preston) [CM]; 4 cf, June 17, 1945; 1 cT, June 20,
1945; 2 cf, June 22, 1945; 1 c?", June 29, 1945; 7 d^, 1 9 , June 1945; 1 d", July 8,
1945; 1 d^, 1 9 , July 15, 1945; 1 &, July 16, 1945; 2 d^, July 17, 1945; 4 d", July
25, 1945; 5 c^, 4 9 , July 1945; 1 c^, Aug. 7, 1945; 1 9 , Aug. 17, 1945; 2 d^, 1 9 ,
August 1945; 1 d", 2 9 , July 1-7 (Sweadner) [CM]; Cumberland Co., Carlisle, 1
d^, June 2.5, 1931 (C. C. Hill) [USNM]; Dauphin Co., Rockville, 2 9 , July 22, 1917
[BPI]; 1 6", June 28, 1918 [BPI]; Fayette Co., Ohiopyle, 1 &, Aug. 10, 1905 [CM];
Forest Co., Kelletville, 1 d", July [CM]; Lycoming Co., Barbours, 1 d', July 8,
1921 (J. C. Bradley) [CU]; Monroe Co., Delaware Water Gap, 1 d", no date
[AMNH]; Montgomery Co., Homebrook, Lower Merion Tp., 2 9, June 23, 1916
[ANS]; 2 9 , July 11, 1916 [ANS, CNC]; 1 9 , July 18, 1916 [ANS]; 1 9 , July 19,
1916 [ANS]; 3 9 , July 22, 1916 [ANS, CNC]; 2 9 , July 23, 1916 [ANS]; 1 d", Aug.
2, 1916 [ANS]; 1 9 , Aug. 3, 1916 [ANS]; 1 9 , Aug. 7, 1916 [ANS]; 1 9 , July 4,
1917 [ANS]; 1 9 , July 8, 1917 [ANS]; 1 9 , July 23, 1917 [ANS]; 1 9 , Aug. 9, 1917
[ANS]; 1 9 , June 10, 1920 [ANS]; 1 9 , June 28, 1921 [ANS]; 1 d", 1 9 , July 7,
1921 [ANS]; Spring Mount, 1 d', June 15 (H. A. Wenzel) [ANS]; Philadelphia
Co., Philadelphia, 1 d', June 16, 1909 (F. Weigand) [CNC]; 1 9, June 23 (F.
Haimbach) [ANS]; Roxborough, 1 d', June 22, 1913 [CNC]; 1 d^, June 24, 1913
[CNC]; 1 d, May 26, 1916 (F. Haimbach) [ANS]; Tioga Co., Blackwell, 1 d",
July 1, 1921 (J. C. Bradley) [CU]; Washington Co., Finleyville, 2 d', June 8-14
[CM]; 1 9, June 24, [CM]; 1 d', August 15-21 [CM]; 1 d", Sept. 22-30, [CM];
Westmoreland Co., Jeannette, 1 d', June 15 (H. G. Klages) [CM]; 1 d", June 19;
1 d", June; 1 d', 3 9 , July 8; 2 9 , July 19; 1 d", 1 9 , July 25; 1 9 , July 29; 1 9 ,
July 30; 5 9 , Aug. 6, 1904; 1 d", Sept. 19.
South Carolina: Anderson Co., Anderson, 2d', June 5, 1917 [CU].
South Dakota: Davison Co., Mitchell, 1 d", Sept. 14, 1945 (E. C. Johnston)
[CNC]; Union Co., Elk Point, 1 9 , no date (C. N. Ainslie) [USNM]; county
unknown. Cedar Pass, Bad Lands, 1 d", Aug. 13, 1940 (C. E. Mickel) [UM].
Tennessee: Davidson Co., Nashville, 1 d" "6/3" (Ainslie) [USNM]; Grundy
Co., Monteagle, 1 9, June 30, 1930 (Richards) [CU]; Knox Co., Knoxville,
1 9, June 28, 1916 (G. G. Ainslie) [USNM]; Morgan Co., Burrville, 1 d", June
27, 1955 (A. K. Wyatt) [CNHM].
Texas: Culberson Co., Guadalupe Mts., 2 mi. north Pine Springs (5700 ft.),
1 9 , July 19, 1963 (Harry Clench) [CM].
Utah: Tooele Co., Loop Camps, 13 mi. southwest Grantsville (7400 ft.),
1 &, July 19, 1958 (F., P., B., and J. Rindge) [AMNH]; 1 d^, July 6, 1960.
Vermont: Rutland Co., Brandon, 1 9 , July 15, 1962 (J. C. Shaffer) [JCS].
Virginia: Giles Co., Mountain Lake, 1 d", July 8, 1938 (L. J. and M. J.
Milne) [CNC].
West Virginia: Greenbrier Co., White Sulphur Springs, 2 d", July 21, 1930
(J. G. Needham) [CU].
Wyoming: Carbon Co., Bottle Creek Camp. 7 mi. southwest Encampment
(8800 ft.), 1 d^, Aug. 8, 1959 (F., P., and B. Rindge) [AMNH]; 1 d^, Aug. 9,
1959; 2 d", Aug. 11, 19.59; Crook Co., 5 mi. west Alva, 3d', July 8, 1949 [AMNH;
Reuter Canyon Camp, 5 mi. north Sundance (6100 ft.), 2 d*, July 12, 1959
REVISION OF PEORIINAE AND ANERASTIINAE 35
(F., P., and B. Rindge) [AIMNH]; 1 d', July 13, 1959; 1 &, July 14, 1959;
(5900 ft.), 1 (f, July 11, 1962; Park Co., Codj-, 5 c?, no date [CM]; Lake Creek
Camp, 13 mi. southwest Cooke City, Mont. (6900 ft.), 1 d", July 25, 1959 (F.,
P., and B. Rindge) [AMNH],
CANADA: Alberta: Calgary, 1 d", July 24, 1905 (F. H. WooUey-Dod)
[CNC]; Horseshoe Canyon, Drumheller (2750 ft.), 1 cT, July 17, 1960 (D. F.
Hardwick) [CNC]; Lethbridge, 1 d, July 30, 1921 (H. L. Seamans) [CNC];
1 d, June 27, 1956 (E. E. Sterns) [CNC]; 1 d, July 7, 1956; Dominion Range
Sta. Manyberries, 1 &, July 23, 1951 (D. F. Hardwick) [CNC]; 1 cf, Aug. 2,
1951; 2 cf, Aug. 3, 1951; 1 d, Aug. 4, 1941.
Manitoba: Aweme, 1 d, June 23, 1904 (Criddle) [USNM]; 1 d, July 1,
1905; 3 d, July 2, 1921 (N. Criddle) [CNC]; 1 c^, July 6, 1921; 1 d,
1922; 1 d, July 15, 1923; Brandon, 2 cf, 1 9, July 20, 1958 (R. B. Madge)
[CNC]; 1 cf, July 31, 1958 (R. L. Hurley) [CNC]; Cartwright, 1 d, July 4, 1904
(E. F. Heath) [USNM]; 1 d, July 26, 1905; 2 d, July 29, 1905; 1 d, no date;
Red Rock Lake, Whiteshell For. R., 1 ?, July 10, 1954 (C. D. Bird) [AMNH];
locality unknown, 1 cT, July 15 [USNM].
Nova Scotia: Annapolis, 1 cf , June 20, 1946 (McDunnough and D. Ferguson)
[CNC]; 1 d, July 5, 1946; 1 d, Aug. 1, 1946; 1 d, Aug. 13, 1946; Petite Riviere,
2^, July 21, 1935 (J. McDunnough) [CNC]; Truro, 1 9 , July 9, 1913 (R. Mathe-
son) [CU]; 1 d, JulOy 1, 1913; 1 9, Aug. 3, 1913.
Ontario: Blackburn, Ic?, June 18, 1941 (J. McDunnough) [CNC]; Black
Sturgeon Lake, Id, July 23, 1963, [CNC]; Chatham, 19, June 27, 1931 (D. A.
Arnott) [CNC]; Id, July 11, 1931 [CNC]; Constance Bay, 2d, July 9, 1934
(W. J. B.) [CNC]; Grand Bend, Id^, July 17, 1939 (T. N. Freeman) [CNC];
Marmore, Id', June 13, 1952 (J. R. Vockeroth) [CNC]; 1 d^, June 14, 1952; 3d',
2 9, June 15, 1952; Id', June 17, 1952; Id', June 19, 1952; 2d', 3 9 , June 20, 1952;
8d', 1 9 , June 21, 1952; 1 d, June 22, 1952; 1 d', 2 9 , July 3, 1952; 1 d, July 4,1952;
Id', July 21, 1952; Mer Bleue, Ic^, June 12, 1937 (G. A. Hobbs) [CNC]; Mer
Bleue, Hawthorne, Id', June 15, 1937 (E. G. Lester) [CNC]; Normandale, Id',
July 7, 1956 (Freeman and Lewis) [CNC]; Ottawa, Id', June 30, 1907 (C. H.
Young) [CNC]; Ottawa, Hunt Club, 2d, July 8, 1937 (G. S. Walley) [CNC];
Palmer Rapids, 1 9 , July 25, 1935 (F. A. Urquhart) [CNC]; Port Colborne, 1 9 ,
June 17, 1934 (J. J. de Gryse) [CNC]; Id', June 20, 1934; Sault Ste. Marie, Id',
July 17, 1957 [CNC]; Simcoe, 2d', June 9, 1939 (T. N. Freeman) [CNC, LACM];
1 d', 1 9 , June 19, 1939; 1 d, June 20, 1939 (T. N. Freeman) [CNC]; 1 d, July 27,
1939; Id', June 5, 1949; 1 9 , July 5, 1949; 4d', July 8, 1949; Strathroy, 2d', 1 9 ,
July 5, 1926 (H. F. Hudson) [CNC, LACM]; Id', July 11, 1929 (H. F. Hudson)
[CNC]; Teeswater, Id', July 18, 1949 (D. F. Hardwick) [CNC]; Id, 19, July
20, 1949; Id', July 29, 1949; 2d', Aug. 6, 1949; 1 9 , Aug. 9, 1949; Toronto, Id',
June 1930 (H. S. Parish) [CU]; Id', July 1930; Trenton, 2d', June 18, 1901
(Evans) [CNC]; Id', June 16, 1904; 1 9 , June 23, 1907; Id', June 22, 1908; Id',
June 14, 1911; Icf, July 16. 1911; Id', June 27, 1912; 1 9 , July 6, 1912; Id', July
12, 1912; Wanbamie, Parry Sound, 1 d, July 12, 1915 (H. S. Parish) [CU]; Wiarton
3 9 , July 26, 1949 (D. F. Hardwick) [CNC].
Quebec: Alcove, 19, July 4, 1935 (T. N. Freeman) [CNC]; Duchesnay, 19,
July 27, 1943 (R. Lambert) [CNC]; Id', July 14, 1947; Granby, Id', June 22,
1938 (P. E. Mercier) [CNC]; Harrington Lake, Gatineau Park, 1 d, June 11, 1954
(J. E. H. Martin) [CNC]; Id', June 13, 1954 (H. J. Huckel) [CNC]; Id', June 15,
1954 (Martin) [CNC]; 29, June 16, 1954 (Huckel) [CNC]; 2d', June 18, 1954
(Martin) [CNC]; 2d', 2 9 , June 20, 1954; 3d', 1 9 , June 21, 1954; 4 d', 2 9 , June
24, 1954; Id', June 25, 1954; Knowlton, Id', July 7, 1929 (L. J. MUne) [CNC];
19, July 11, 1929; Lac Mondor, Ste. Flore, Id', July 13, 1951 (E. G. Munroe)
36 U.S. NATIONAL MUSEUM BULLETIN 280
[CNC]; L. a la Torrue, 1 ? , July 16, 1947 [CNC]; Icf, 2 9 , July 17, 1947 (E. G.
Monroe) [CNC]; Meach Lake, Id', June 23, 1941 (G. A. Hobbs) [CNC]; Icf,
June 25, 1941 (G. A. Hobbs) [CNC]; Norway Bay, 1?, July 9, 1937 (E. G.
Lester) [CNC]; 1 ? , July 15, 1937 (F. A. Urquhart) [CNC]; Ste. Anne de Bellevue,
IcT, June 29, 1947 (E. G. Monroe) [CNC]; Icf', July 3, 1947; Id', 29, July 8,
1947; St. Annes, 2^^, 3 9 , July 5, 1947 (E. G. Munroe) [CNC]; 1 cf , 9 9 , July 6,
1947: 7 9 , July 8, 1947; 3 9 , Aug. 2, 1947.
Saskatchewan: Christopher Lake, 1 cf , July 3, 1939 (A. R. Brooks) [CNC];
Id^, July 5, 1939; Indian Head, Id^, July 2, 1924 (J. J. deGryse) [CNC]; Id',
July 29, 1924; Saskatoon, Id', June 30, 1923 (Kenneth M. King) [CNC]; Id',
July 7, 1923; Id", July 17, 1923; Id', July 19, 1923; Willow Bunch. Id', July 27,
1955 (C. D. Miller) [CNC].
Discussion. — This is tlie most common and widely distributed
member of the subfamily, the species being common from mid-June
to early August throughout the northeastern United States. Western
specimens tend to be larger and more uniform in maculation than
those from the eastern and central states.
Peoria luteicostella (Ragonot), new combination
Figures 12, 82, 117, 155
Hypsotropa luteicostella Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p-
212, — Smith, 1891, p. 85. — Ragonot, 1901, p. 376. — Hulst, 1902, p. 439. —
Barnes and McDunnough, 1917, p. 149. — Grossbeck, 1917, p. 134. — Hamp-
son, 1918, p. 70. — McDunnough, 1939, p. 36. — Kimball, 1965, p. 251.
Wekiva nodosella Hulst, 1890, p. 215. — Smith, 1891, p. 85. — Hulst, 1902,
p. 440. — Grossbeck, 1917, p. 134. — Rindge, 1955, p. 167.
Osceola perlepidellus Smith, 1891, p. 85. [Nomen nudum. New synonymy.]
Chipeta perlepidella Hulst, 1892, p. 62. — Barnes and McDunnough, 1917, p.
150. — McDunnough, 1939, p. 36. — Rindge, 1955, p. 169. — KimbaU, 1965,
p. 251. [New synonymy.]
Chipota [sic] perlepidella Hulst, 1902, p. 441.
Bandera perlepidella (Hulst). — Hampson, 1918, p. 90.
Diagnosis. — This species resembles approximella in venation and
maculation, but lacks the brownish white area posterior to the anal
vein. The obliquely ascending palpi of both sexes and the postero-
lateral gnathos hooks of the male are unique to this species.
Description. — Frons wine red; labial palpi obliquely ascending,
wine red on outer sides, light pink on inner sides, third segment and
dorsal half of second segment sometimes black on outer sides; maxillary
palpi small, wine red; antennae with base wine red anteriorly, white
posteriorly, shaft brown, wine red toward base, male laminate; outer
sides of occiput, patagia, and tegulae wine red, vertex behind antennae
and dorsal area of occiput white; legs white on outer sides, wine red
on inner sides, tarsi black on inner sides.
Forewings with broad white band anterior to cell, narrowing to
point at apex, bordered anteriorly near base with wine red to black,
entire band sprinkled with wine red scales; white band bordered
REVISION OF PEORIINAE AND ANERASTIINAE 37
abruptly behind with black, black blending into posterior mne red
area; outer fringe wine red to black, Hindwing light brown, darker
in apical area. Venation as in P. bipartitella.
Male genitalia with medial process of uncus bearing minute central
tooth, rarely several. Gnathos broadening posteriorly into a pair of
large flat anteriorly directed hooks. Juxta V-shaped. Vinculum Avith
dorsal margin flanged at apex. Valvae with costa terminating in
strong slender spine reaching or surpassing the blunt membranous
tip of cucculus; sacculus with numerous fine rather long hairs. Aedea-
gus with vesica bearing small subserrate cornutus.
Female genitalia with ovipositor moderately setose laterally,
numerous fine setae on caudal margin. Posterior apophyses tapering
to point at base, not flattened, posterior half strongly curved; an-
terior with base slightly thickened. Eighth abdominal segment
unusually large.
Types. — H. luteicostella, in the Museum National d'Histoire
Naturelle; W. nodosella, in the U.S. National Museum (lectotype),
in the American Museum of Natural History (lectoparatype) ; C.
perlepidella, in the American Museum of Natural History.
Type data. — H. luteicostella, lectotype male, hereby designated,
Florida; genitalia slide No. 535, J. Shaffer, June 20, 1965,
W. nodosella, lectotype male, hereby designated, Florida, Fernald
collection; USNM 40081; genitalia slide No. 570, J. C. S., Nov. 23,
1965.
Lectoparatype: Florida; abdomen, antennae, and labial palpi lost.
In the original description of the species Hulst gives April as the date
of capture.
C. perlepidella, lectotype female, hereby designated, Florida,
collection G. D. Hulst; genitalia slide No. 3228, Carl Heinrich, June
14, 1946.
Specimens examined. — 28 cT, 2 9 •
Distribution (Map 10). — Southern Georgia and Florida.
UNITED STATES: Florida; Alachua Co., GainesvUle, Sc?, 19, July 8,
1927 (J. Speed Rogers) [CU]; !&, July 1927 (J. Speed Rogers) [CU]; Highlands
Co., Archbold Biol. Sta., Icf, June 17, 1964 (Jay C. Shaffer) [JCS]; 2 6^, June 29,
1964; 3d^, July 5, 1964; IcT, July 12, 1964; Hillsborough Co., Stemper, Id",
Aug. 13, 1912 (G. Krautworm) [CM]; Volusia Co., Cassadaga, Id', Oct. 12, 1962
(S. V. Fuller) [CPK]; Wakulla Co., Panacea, ISd', 1?, Aug. 11, 1926 (C. O.
Handley) [USNM].
Georgia: County unknown, Billy's Island, Okefenoke Swamp, 2cf, June 1912
[CU].
Anacostia, new genus
Type. — Anacostia tribulella, new species.
Diagnosis. — The male genitalia are similar to those of Peoria, but
differ mainly in the following ways : In Anacostia the spicate processes
38 U.S. NATIONAL MUSEUM BULLETIN 280
are three-branched rather than two-branched, the medial band of the
uncus has a central pentagonal process, and the juxta bears a pair of
lateral knobs, each with a small cluster of setae.
Description. — Labial palpi porrect, about 3% times as long as eye
diameter; maxillary palpi moderate, not reaching frons; male antennae
with basal segments of shaft fused, shaft serrate and fasciculate, female
unknown; oceUi well developed.
Forewing with 11 veins; Ri from cell relatively near to R2+4; R2
relatively long stalked with R3+4; M2+3 stalked for about one-half
their length or less; Cui from near M2+3. Hindwing with 7 veins;
Sc and Rs stalked for abovit one-half their length; M3 and Cui stalked
for about one-third their length.
Male genitalia with medial band of uncus very broadly rounded,
bearing a well-developed, ventraUy directed, apical, pentagonal
process; spicate processes three-branched. Gnathos broad, flat, thin,
broadened at base; sharp pointed apicaUy, but without special process.
Juxta V-shaped, bearing pair of setose knobs. Vinculum subquadrate,
anterior corners weU rounded. Valvae relatively slender; costa sparsely
setose, with smaU apical tooth at tip of valve; sacculus densely
pubescent. Aedeagus with vesica unarmed.
Female unknown.
Discussion. — The genus is apparently closely related to Peoria,
but differences in the male genitalia are too great to allow inclusion
of the one species in that genus.
Anacostia trihulella, new species
Figures 25, 83, 118
Diagnosis. — This is the only known species in the genus.
Description. — Labial palpi with all segments dark brown to
black; antennae brown, shaft with tuft of black scales on basal seg-
ments; head and thorax black; legs black on outer sides, brown
on inner sides; abdomen dorsally brown, becoming golden brown
anteriorly, ventrally black.
Forewing ground dark brown to black; area anterior to cell white
sprinkled mth black scales; discal spot, transverse anterior, transverse
posterior, and subterminal lines black, not conspicuous against
ground. Hindwing broAvn, darker toward apex; terminal line white,
brown bordered on both sides.
Genitalia as described for the genus.
Types. — In the Academy of Natural Sciences (holotype); in the
Cornell University collection (two paratypes).
Type data.- — Holotype, male, Holland Va. (Nansemond Co.),
Aug. 1, 1945, O. Buchholz; A.N.S. Type No. 7817; genitalia slide
No. 595, J. Shaffer, Apr. 24, 1966.
REVISION OF PEORIINAE AND ANERASTIINAE 39
Paratypes: Two males, both Clyde (Bryan Co.), Georgia, Sept.
11-12, 1931, Bradley and Knorr; C.U. Type No. 4404; one male
genitalia slide No. 439, J. Shaffer, Mar. 25, 1965.
Other specimens examined. — None.
Distribution (Map 4). — Probably the Atlantic Coastal Plain;
known only from Georgia and Virginia,
Arivaca, new genus
Type. — Poujadia pimella Dyar, 1906.
Diagnosis. — Males of this genus may be recognized by the com-
bination of (a) the gnathos bearing a bulbous or padlike medial
process, and (b) each spicate process of the uncus having a short
posterior and a much longer anterior sharp-pointed spine (figs.
119-125).
Description. — Frons conical, one-half to three-quarter times as
long as eye diameter; labial palpi porrect; maxiUary palpi very small;
tongue poorly developed; antennae with scape compressed, male
shaft with basal segments fused, female shaft filiform, scaled dorsally
and laterally, finely ciliate ventrally, somewhat compressed; eye
diameter about 0.2 mm gi-eater in male than in female; oceUi normal.
Forewings with 10 or 11 veins; Ri from well before upper outer
angle of cell; II3+4 stalked; Mi from the angle; M2+3 stalked or fused,
from lower outer angle; Cui from just before the angle; Cu2 from
well before the angle. Hind^^dngs with 6 or 7 veins; Sc and Rs approxi-
mate or connate on basal half; Mi from upper outer angle of cell;
M2 absent; Ms+Cui free, stalked, or fused, from lower outer angle;
Cu2 from before the angle.
Male genitalia ^\^th spicate processes of uncus flat, each bearing
short sharp posterior and long tapering anterior thorns. Gnathos
arms straplike, bearing anteromedial projection; medial process
bulbous or padlike. Vinculum broadly rounded. Aedeagus flattened,
vesica bearing pair of cornuti.
Female genitaha \\dth ductus biu-sae flattened, rather long, well
sclerotized. Bursa smaU, hghtly sclerotized. Ductus seminahs from
posterior half of ductus bursae.
Discussion. — The seven species of Arivaca faU into foiu- rather
distinct subunits herein distinguished as species groups rather than
as genera or subgenera. Because the genus is apparently best devel-
loped in the unstudied region south of the United States, it is possible
that intermediate forms exist between the species groups, and it is
felt best not to give the groups nomenclatural status.
40 U.S. NATIONAL MUSEUM BULLETIN 2 80
Key to the Species Groups of Arivaca Based Upon the Maculation
1. Cubitus with prominent white streak; white band anterior to cell (figs. 19,
20) ostreella group
White trace of cubitus not prominent, absent if white band present anterior
to cell 2
2. M2+3 stalked; forewing not appearing lighter anterior to cell (figs. 15, 16).
pimella group
M2+3 fused; forewing somewhat lighter anterior to cell 3
3. Forewing ground uniform reddish brown, prominent white band anterior
to cell (Florida) albicostella group
Forewing gray, mixture of black and white scales, radius and subcosta white
traced (west of Mississippi River) albidella group
The Pimella Group
Diagnosis. — This group may be distinguished by the male valvae,
in which the costa ends in a short triangular tooth well before the
rounded tip of the cucullus. A similar projecting membranous cucullus
exists in albicostella, but the costa terminates in a stout spine in that
species.
Description. — ^Frons light brown to brown; labial palpi with basal
segment white, second and third segments clothed with white-tipped
brown scales dorsally and on outer sides, white ventrally and on inner
sides; vertex Ught brown anteriorly, white posterior to antennae,
occiput, patagia, and tegulae brown laterally, lighter dorsally; legs
clothed with white-tipped brown scales on outer sides, white on inner
sides.
Forewing ground Ught orange brown, veins white traced and
bordered -with brown, Ai and A2 traced for most of their length, A3
traced at base; 11 veins; M2+.3 stalked. Hindwing hght brown, fringe
white; 7 veins; M3 usually free from lower outer angle of cell.
Male genitaUa with uncus moderately well sclerotized between
spicate processes. Juxta rounded, with caudal V-shaped notch. Valvae
with costa sparsely setose, apically bearing short triangular tooth;
cucullus with rounded membranous tip projecting well beyond costa;
sacculus with patch of stiff setae. Aedeagus with cornuti serrate,
small, subequal.
Female genitalia with base of posterior apophyses flat, shaped Uke
parallelogram. Bursa unarmed.
Discussion. — The two included species may be separated by the
condition of vein R2, which is stalked with R3+4 in linella but free
in pimella, but this character is usually variable elsewhere in the sub-
family so that positive identification should rest with examination of
the male genitalia.
REVISION OF PEORIESTAE AND ANERASTIINAE 41
Key to Species of the Pimella Group Based Upon the Male Genitaha
Posterior margin of medial process of uncus fiat, usually bearing pair of short,
sharp lateral teeth; apical process of gnathos padlike, not strongly projecting
(fig. 120) linella
Posterior margin of medial process of uncus rounded, unarmed; apical process of
gnathos bulbous, strongly projecting dorsally (fig. 119) pimella
Arivaca pimella (Dyar), new combination
Figures 15, 50, 52, 84, 119, 156
Poujadia pimella Dyar, 1906, p. 31. — Barnes and McDunnough, 1917, p. 149.
— McDunnough, 1939, p. 36.
Diagnosis. — The broadly rounded unarmed medial process of the
uncus and the strongly projected medial process of the gnathos each
separate this species from linella.
Description. — Antennae white, male shaft laminate, cilia about
one-fifth as long as segment width.
Forewing sprinkled with brown and white anterior to cell; cell
brownish white; R2 free from cell. Hindwing light brown in female,
somewhat darker in male.
Male genitalia with medial process of uncus rounded, unarmed.
Gnathos with apical process strongly projected dorsally.
Type. — In the U.S. National Museum.
Type data. — Lectotype male, hereby designated, Babaquivera
Mts., Pima County, Ariz.; USNM 9012; genitalia shde No. 567, J.
Shaffer, Nov. 23, 1965.
Specimens examined. — 45 cf , 36 9 •
Distribution (Map 9). — Southern Arizona.
UNITED STATES: Arizona: Cochise Co., Cochise Stronghold, 1 9, Aug.
30, 1958 (P. Opler) [UCB]; Paradise, 1 cf , 1 9 , September (Poling) [CM]; Chiri-
cahua, 1 mi. west portal, 1 cf , 1 9 , Sept. 10, 1950 (Cohn, Boone, Cazier) [AMNH];
Ramsay Canyon, Huachuca Mts., 1 d', 1 9, July 10-15, 1941 (A. B. Klots)
[AMNH]; San Bernardino Ranch (3750 ft.), 1 9, August (F. Snow) [UK];
Southwestern Res. Sta., 5 mi. west Portal, 1 cf, Sept. 6, 1959 (J. R. Bowers)
[UCB]; 1 d^, Aug. 25, 1962 (M. Statham) [AMNH]; 1 9, Sept. 2, 1959 (Carl W.
Kirkwood) [CPK]; 1 9, Sept. 17, 1959; 1 9, Sept. 20, 1959; Gila Co., vicinity
Globe, 1 c?, Aug. 4-5, 1937 (A. B. Klots) [AMNH]; Pima Co., "Baboquavaria
Mts.," 4 c?, July 15-30, 1903 (O. C. Poling) [USNM]; 2 9 , July 1-15, 1923;
2 d", July 15-30, 1923; 1 d^, 29, Aug. 1-4, 1923; 1 9, Aug. 15-30, 1923; 1 c?,
1 9, Aug. 24-31, 1923; 4c?, 49, Sept. 1-15, 1923; 1 d", Oct. 1-15, 1923; 1 d^,
July 15-30, 1924; 7 d^, 1 9, Sept. 1-15, 1924; 1 cT, Oct. 1-15, 1924; Brown
Canyon, Baboquivari Mts., 1 d^, Sept. 5, 1953 (Lloyd M. Martin) [LACM];
1 d', Sept. 6, 1953; Madera Canyon, Santa Rita Mts., 1 9, Aug. 25, 1946 (J. A.
Comstock, Lloyd M. Martin) [LACM]; 1 d', Aug. 2, 1947; 3 9, Sept. 7, 1948
(Lloyd Martin) [LACM]; 1 &, Sept. 8, 1948; 1 9, Aug. 15, 1949 (C. W. Kirk-
wood) [LACM]; 1 9, Aug. 17, 1949; 1 9, Sept. 11, 1950; 1 d^, 1 9 , Sept. 13,
1950; 1 9, Sept. 15, 1950; 1 d", Sept. 5-12, 1951 (William Hammer) [CNC];
1 9 , Aug. 8, 1952 (C. W. Kirkwood) [LACM]; 1 d^, Aug. 14, 1952; 2 9 , Aug. 30,
42 U.S. NATIONAL MUSEUM BULLETIN 280
1952 (Lloyd Martin) [LACM]; 6 cf , 1 9, Aug. 19, 1953 (Robert J. Ford) [CNC];
1 9, Aug. 16, 1953; 1 9, Aug. 18, 1953; 1 9, Aug. 21, 1953; 1 9, Sept. 2, 1953
(Lloyd Martin) (LACM]; 1 cf, Sept. 8, 1953; 2 9, Aug. 24-Sept. 2, 1957 (W. A.
Hammer) [LACM]; locality unknown: "So. Arizona," 3 c?, 1 9, August 1-15
(Poling) [USNM].
Discussion. — In the original description Dyar lists two males, one
of these is in the National IVEuseum, the fate of the other is unknown.
Arivaca linelkif new species
Figures 16, 56, 85, 120, 157
Diagnosis. — The flat, moderately well sclerotized medial process
of the uncus distinguishes this species within the genus. The stalked
condition of R2 separates both sexes from pimella.
Description.— Antennae light brown, sublaminate in male, cilia
about one-third as long as segment width, female with shaft white.
Forewings brown anterior to cell; R2 stalked with R3+4.
Male genitalia with mediodistal margin of uncus irregular, usually
bearing pair of small sharp teeth laterally. Gnathos with apical
process padlike, not strongly projected dorsally.
Types. — In the Canadian National collection.
Type data. — Holotype male, Colfax County, N. Mex., Sangre de
Cristo Mts., Cimarron Canyon (7900 ft.), July 13, 1962, E. and I.
Munroe; C.N.C. Type No. 9442; genitalia slide No. 274, J. Shaffer,
Sept. 15, 1964.
Paratypes: Four males, same data and locality as holotype.
Specimens examined. — 18 cf , 3 9 •
Distribution (Map 9).
UNITED STATES: Arizona: Apache Co., 3 mi. west Eagar (7100 ft.),
Pinon-Juniper life zone, 1 cf , July 13, 1962 (E. and I. Munroe) [CNC]; Greer,
White Mts. (8500 ft.), 1 d', Aug. 2, 1962 (E. and I. Munroe) [CNC]; 1 9 , Aug. 4,
1962; 5 cT, Aug. 6, 1962; 1 9, Aug. 9, 1962; 1 cf, Aug. 11, 1962.
Colorado: "Rock Creek Canyon," Icf, July 19, 1961 (Margot May) [CPK].
New Mexico: Colfax Co., Cimarron Canyon, Sangre De Cristo Mts. (7900
ft.), 2 d", July 14, 1962 (E. and I. Munroe) [CNC]; McKinley Co., McGaffey,
Zuni Mts. (7500 ft.), 1 <?, July 21, 1962 (E. and I. Munroe) [CNC]; 1 cf , July 25,
1962; Sandoyal Co., Frijoles Canyon, Bandelier Nat. Mon. (6050 ft.), 4 cf , July
17, 1962 (E. and I. Munroe) [CNC]; 1 9, July 18, 1962; Horseshoe Springs
Camp, 2 mi. west La Cueva (7900 ft.), 1 cf , July 29, 1961 (F., P., and J.
Rindge) [ABK].
The Ostreella Group
Diagnosis. — The group is easily recognized by the light orange
forewings with the costal margin and cubitus both prominently
marked with white. The tapering pointed valvae distinguish males
of the group.
REVISION OF PEORIINAE AND ANERASTIESTAE 43
Description. — Frons about tkree-foiirths as long as diameter of
eye, orange laterally, lighter dorsaUy; labial palpi Avith basal segment
white, second and third segments orange brown on outer sides, white
on inner sides; antennae white, finely pubescent, bearing tuft of scales
on basal segments of shaft; occiput and vertex white behind antennae,
occiput brown laterally, patagia and tegulae Hght orange.
Forewing vdih broad white band anterior to cell, costa margined
basally mth dark brown; white band bordered posteriorly with
narrow dark brown band dividing distally to trace K4 and Mi; promi-
nent white band tracing cubitus and M2, broadest near lower outer
angle of cell; Cui and Cu2 traced with mixtm-e of brown and white;
R2 from close to R.3+4, sometimes short stalked. Hind^\-ing with 7 veins;
M3 and Cui from point or short stalked.
Male genitaha Avith medial process of uncus well sclerotized,
rounded, bearing pair of caudomesally directed spines. Gnathos mth
medial process strongly projected. Valvae tapering to point.
Female genitaha mth eighth abdominal segment heavily sclerotized,
tapering posteriorly. Anterior apophyses strongly divergent; posterior
parallel, pointed at base.
Key to the Species of the Ostreella Group Based Upon External
Characters
M2+3 forked; white band extending from cell to costal margin .... ostreella
M2+3 fused; white band extending half way from cell to costal margin, bordered
anteriorly with light orange poohella
Arivaca ostreella (Ragonot), new corabination
Figures 19, 53, 86, 121, 158
Saluria ostreella Ragonot, 1887, p. 18; 1889, p. 117. — Smith, 1891, p. 85.—
Ragonot, 1901,p. 362. — Hulst, 1902, p. 438. — Barnes and McDunnough,
1917, p. 149. — Hampson, 1918, p. 102. — McDunnough, 1939, p. 36.
Saluria ostrella [sic] Hulst, 1890, p. 211.
Peoria discostrigella Dyar, 1904, p. 115. [New synonymy.]
Diagnosis. — ■ The stalked condition of M2+3 distinguishes both
sexes from poohella. Males are easily recognized by the pectinate
antennae.
Description. — Labial palpi about 4)2 times as long as diameter of
eye; male antennae pectinate.
Fore^ving with cell brownish orange; Ai faintly traced on distal
third; A2 and A3 basally traced with mixture of white and brown,
A2 usually broadly so; ground yellow or Ught orange posterior to cell;
11 veins; M2+3 stalked. Hindmng light brown in both sexes.
Male genitaha with uncus bearing heavy mediodorsal triangular
plate, sharp pointed caudally. Juxta round, with deep V-shaped
285-934—68 4
44 U.S. NATIONAL MUSEUM BULLETIN 2 80
caudal incision. Valvae tapering to rounded tip; costa vnth small
medial tooth. Aedeagus with vesica bearing one smaU serrate and
one large pectinate cornutus.
Female genitalia with bursa unarmed.
Types. — S. ostreella, in the British Museum (Natural History) ;
P. discostrigeUa, in the U.S. National Museum.
Type data. — S. ostreella, lectotype female, hereby designated,
labeled as follows: "Type; ARIZONA. Morrison.; Paravicine Coll.
B.M. 1937-383.; Anerastia ostreella Rag. type; Saluria ostreella
Ragonot; 9 genitaha slide 1-4-1967 J. Shaffer No. 704." P. discos-
trigeUa, lectotype female, hereby designated, Roswell, New Mexico,
August 22, Cockerell; USNM 7933; genitaha shde No. 566, J. C. S.,
Nov. 23, 1965. In the original description Dyar reports: "Three 99,
Roswell, New Mexico, August 22 (T. D. A. Cockerell), Tucson,
Arizona, July 21 (E. A. Schwarz)."
Specimens examined. — 6 cf, 49 9-
Distribution (Map 9). — ^Southern Arizona to BrownsviUe, Tex.
UNITED STATES: Arizona: Cochise Co., Southwestern Research Sta.,
5 mi. west Portal (5400 ft.), 9 ? , July 15, 1955 (W. J. Gertsch) [AMNH]; 1 9 ,
July 26, 1955; 1 cf, June 12, 1958; Pima Co., Redington, 3 9 , no date [USNM];
"Tuson," 1 9 , "21. 7." (E. A. Schwarz) [USNM].
New Mexico: Chaves Co., Roswell, 1 9 , August (Cockerell) [USNM]; Eddy
Co., White City, 3 9 , May 14, 1950 (E. C. Johnston) [CNC]; 1 9 , May 16, 1950;
1 cT, 2 9 , July 23, 1959, (A. B. Klots) [ABK]; Hidalgo Co., Lordsburg, 3 9 , Aug. 4,
1937 [AMNH]; Luna Co., 10 mi. east Deming, 3 d', 29, July 12, 1917 [CU];
14 9, July 8-15 [USNM].
Texas: Brewster Co., Alpme, Davis Mts., 1 9 , Sept. 12, 1958 (R. R. McElvare)
[CNHM]; Big Bend, 1 cf, Apr. 15-30, 1926 (O. C. Poling) [USNM]; 1 9 , May
1-15, 1926; Cameron Co., San Benito, 29, July 8-15 [USNM]; 9, August
[USNM]; Reves Co., 35 mi. northwest Pecos, 4 9 , June 15, 1937 (George Willett)
[LACM].
Arivaca poolzella, new species
Figures 20, 57, 87, 122, 159
Diagnosis. — The fused condition of M24.3 distinguishes this species
from ostreella, as does the light orange trace on the costal margin.
Description. — Labial palpi 3K to 5)^ times as long as eye diameter;
antennae sublaminate in male.
Foremng with white costal band extending half way from cell to
costa, bordered anteriorly with light orange; cell orange; Ai sometimes
faintly traced near outer margin; A2 and A3 near base traced with
white and bordered A\dth dark brown; ground lighter orange posterior
to Ai fold; 10 veins; M2+3 fused. Hingwing dark brovna in male, light
brown in female.
REVISION OF PEORIINAE AND ANERASTIESTAE 45
Male genitalia with juxta rectangular; distally concave, proximally
convex. Valvae tapering to point at apex of costa. Aedeagus with
cornuti elliptical, serrate.
Female genitalia mth bursa bearing near its base a cluster of about
20 stout, sharp-pointed, inwardly directed spines.
Types. — In the collection of John G. Franclemont, Cornell
University.
Type data. — Holotype male, Madera Canyon, 5600 ft., Santa Rita
Mts., Santa Cruz County, Ariz., June 24, 1963, J. G. Franclemont;
genitalia slide No. 714, J. Shaffer, Jan. 16, 1967.
Paratypes : Three males, two females, same data as holotype except
as follows: two males, June 29, 1963, one ^\ith genitalia slide No. 715,
J. Shaffer, Jan. 16, 1967; one male, June 28, 1963, genitalia slide
No. 724, J. Shaffer, Jan. 27, 1967; one female, June 28, 1963, genitalia
slide No. 716, J. Shaffer, Jan. 16, 1967.
Other SPECIMENS EXAMINED. — 7 cf , 15 9-
Distribution (Map 9). — Southern Arizona to Brownsville, Tex.
UNITED STATES: Arizona: Cochise Co., Southwestern Res. Sta., 5 mi.
west Portal (5400 ft.), 1 9 , May 15, 1956 (M. Statham) [AMNH]; 1 ? , June 29,
1956 (Cazier and Ordway) [AMNH]; 1 ? , July 5, 1956; 1 d^, July 9, 1956, Ic?,
1 ? , July 11, 1958 (M. A. Cazier) [AMNH]; Pima Co., Baboquivari Mts., Ic?,
July 1-15, 1923 (O. C. Poling) [USNM]; 1 9 , July 1-15, 1924; 1 9 , Aug. 15-30,
1924; Redington, 1 9 , no date [USNM]; Santa Cruz Co., Madera Canyon,
Santa Rita Mts. (5800 ft.), Ic?, 1 9, June 22, 1960 (David A. Wallesz) [CU];
2 9, June 24, 1960; (4800 ft.) 1 9, June 18, 1963 (J. G. Franclemont) [JGF];
2 9 , June 29, 1963; 1 cf , 1 9 , July 1, 1963.
The Albidella Group
Diagnosis. — Males of the group are delimited by the broadly
rounded apex of the valvae, the cucullus not exceeding the costa.
Description. — Frons of white-tipped brown scales; labial palpi
about twice length of head, basal segment light brown, of white-tipped
brown scales dorsally and on outer sides, white ventrally and on inner
sides; antennae light brown, finely ciliate ventrally, tuft of scales on
basal segments of shaft.
Fore-wing uniformly gray, a mixtiu-e of dark brown and white scales,
Rs white traced; 10 veins; R2 stalked with R3+4; M2+3 fused. Hind\\-ing
light brown; 7 veins, but M3 sometimes completely fused with Cuj.
Male genitalia with medial part of uncus poorly sclerotized, irregu-
larly rounded. Apical process of gnathos well projected dorsally. Juxta
subquadrate, anterior margin rounded, posterior wdth broad V-shaped
notch. Valvae with costa setose, blunt tooth at apex and another just
under half distance to base; tip rounded; sacculus finely setose.
Aedeagus with cornuti equal or nearly so, serrate.
Female genitalia as in the Pimella Group.
46 U.S. NATIONAL MUSEUM BULLETIN 280
Key to the Species of the Albidella Group
Forewing appearing lighter posterior to Aj fold; posterior cusp of spicate process a
small triangular tooth about one-eighth length of anterior cusp (fig. 123)
albidella
Forewing not appearing lighter posterior to Ai fold; posterior cusp of spicate
process a sharp thorn about one-fourth length of anterior cusp (fig. 124) . artella
Arivaca albidella (Hulst), new combination
Figures 17, 88, 123, 160
Peoria albidella Hulst, 1900, pp. 175, 439. — Barnes and McDunnough, 1917,
p. 149.— McDunnough, 1939, p. 36.
Diagnosis. — Distinguishing features are given in the above key.
Description. — Forewings Hght tan to grayish brown, appearing
Ughter posterior to Ai fold.
Male genitaha with posterior cusp of spicate process poorly de-
veloped, usually rounded, sometimes sharp pointed, usually no more
than one-tenth length of anterior process.
Type. — In the U.S. National Museum.
Type data. — Lectotype male, hereby designated, Death Valley,
"April 91 K."; USNM 4809; genitalia shde No. 571, J. Shaffer,
Nov. 23, 1965. In the original description Hulst notes: "Death Valley,
Cal. ; taken by Mr. Koebele, in April. The type number of the
National Museum is 4709."
Specimens EXAMINED. — 7 d^, 1 9.
Distribution (Map 9). — Southwestern United States.
UNITED STATES: New Mexico: Dona Ana Co., Mesquite, near MesiUa
Park,2 2 cT, July 12, 1917 [CU].
Texas: Reeves Co., Pecos, 4 cf , 1 ?, May 18, 1950 (E. C. Johnston) [CNC];
1 cf , June 2, 1950.
Arivaca artella, new species
Figures 18, 89, 124, 161
Diagnosis. — Distinguishing features are given in the above key.
Description. — Male genitalia with posterior cusp of spicate process
well developed as a sharp-pointed thorn about one-fourth as long as
anterior cusp. Apical process of gnathos strongly projected, somewhat
T-shaped in lateral view.
Type. — In the Cornell University collection.
Type data. — Holotype male, Mesquite near MesiHa Park, N. Mex.,
July 12, 1917; Cornell University lot No. 542, sublot No. 46; lot No.
551, sublot No. 916; CU Type No. 4406; genitalia slide No. 366, J.
Shaffer, Feb. 12, 1965.
2 Spelled Mesille Park on specimen labels.
REVISION OF PEORIINAE AND ANERASTIINAE 47
Paratypes: Three males, same date and locality as holotype, Cornell
University lot No. 542, sublot No. 46.
Other specimens examined. — 14 cT, 3 9 .
Distribution (Map 9).
UNITED STATES: Arizona: Apache Co., St. Johns, 3 cf, July 26, 1937
(A. B. Klots) [AMNH].
New Mexico: Dona Ana Co., Mesquite near Mesilla Park, 10 cf, July 12,
1917 (from same lot as type series) [CU]; Hidalgo Co., Cienaga Ranch, near
Rodeo, 1 9 , July 12, 1948 (C. and P. Vaurie) [AMNH]; Sandoval Co., Jemez
Springs (6200 ft.), 1 9, July 16, 1950 (Cohn, Boone, Cazier) [AMNH]; Sulphur
Dam, 5 mi. north Jemez Springs (6300 ft.), 1 cf, July 17, 1950 (Cohn, Boone,
Cazier) [AMNH]; county unknown, 1 9 , no date [AMNH].
The Alhicostella Group
Arivaca alhicostella (Grossbeck), new combination
Figures 21, 60. 90, 125, 162
Calera (?) albicosleUa Grossbeck, 1917, pp. 134-135. — Klots, 1942, p. 419.
Peoria alhicostella (Grossbeck). — McDunnough, 1939, p. 36. — Kimball, 1965,
p. 251.
Diagnosis. — The presence of a white costal band coupled with the
absence of white tracing on the cubitus distinguishes this species wdthin
the genus. In no other species of Arivaca does the male costa terminate
in a subapical spine.
Description. — Frons brown; labial palpi about 2^ times as long as
eye diameter, basal segments light brown, second and third segments
clothed with light brown tipped brown scales on outer sides, light
brown on inner sides and ventrally; antennae with scape brown, shaft
light brown, laminate, basal segments bearing tuft of scales, cilia
slightly less than half as long as segment width; frons and vertex light
brown dorsally, vertex brownish white posterior to antennae, occiput
brownish white dorsally, brown laterally, patagia and tegulae brown;
legs clothed with white tipped brown scales on outer sides, white on
inner sides.
Forewing reddish brown, often sprinjiled wdth light brown and dark
brown scales; light brown anterior to cell, sometimes sprinkled rather
heavily ^vith darker scales. Hindmngs light bro^vn. Venation as in the
Albidella Group.
Male genitalia with medial part of uncus weakly sclerotized, round-
ed; spicate process with posterior thorn directed dorsally, then re-
flexed 90° near base and caudally directed (fig. 125). Juxta rounded,
posterior margin concave. Valvae with costa sparsely setose, produced
as free spine before rounded membranous apex of cucuUus; sacculus
finely setose. Aedeagus with cornuti elliptical, serrate.
Female genitalia as in the Pimella Group.
48 U.S. NATIONAL MUSEUM BULLETIN 280
Type. — In the American Museum of Natural History.
Type data. — Lectotype female, designated by Klots (1942), Fort
Myers, Fla., Apr. 23, 1912; genitalia slide No. 586, J. Shaffer, Nov. 28,
1965.
Lectoparatype female, Everglades, Fla., Apr. 7, 1912.
Specimens examined. — 10 cf, 3 9-
Distribution (Map 10). — Known only from southern Florida.
UNITED STATES: Florida; Broward Co., Ft. Lauderdale, ? , Mar. 27, 1928
(D. M. Bates) [CPK]; Collier Co., Everglades, 4 d^, AprU 1-7 [USNM]; 3 cT,
April 8-15 [USNM]; Dade Co., Paradise Key, Everglades National Park, ?,
Apr. 2, 1952 (G. S. Walley) [CNC]; cf, Apr. 3, 1952; Sarasota Co., 9 , May 13,
1946 (C. P. Kimball) [CPK]; Siesta Key, cf, Mar. 31, 1952 (C. P. Kimball)
[CPK]; &, Mar. 30, 1954 (C. P. Kimball) [CPK].
Atascosa Hulst
Atascosa Hulst, 1890, p. 210. — Smith, 1891, p. 85. — Hulst, 1902, p. 438. [Type:
Atascosa bicolorella Hulst, 1890. Original designation.]
Diagnosis. — The flat, elbowed, bluntly pointed spicate processes
of the uncus (fig. 126) are diagnostic of the genus.
Description. — Frons conical; labial palpi upturned with third
segment decumbent in males, porrect in females; maxillary palpi
reaching frons, cylindrical tuft of long slender scales, sometimes ex-
panded and fanlike; antennae sublaminate, ciliate in males, filiform,
finely ciliate in females; ocelli weU developed.
Forewing mth 11 veins; Ri free from cell; R2 from near to or stalked
with R3+4; M2+3 stalked for about half its length. Hindwing with 7
veins; Sc and Rs stalked; Mi from upper outer angle of cell; M2
absent; M3 stalked with Cui, from lower outer angle; Cu2 from just
before the angle.
Male genitalia with spicate processes in form of a pair of flat lateral
arms, ventroposteriorly directed from base, then elbowed and distal
half ventrally directed; arms bluntly pointed. Gnathos with apical
process a small postero ventrally directed thorn ; arms straplike. Juxta
subquadrate, shieldlike. Vinculum V-shaped, somewhat flattened
anteriorly. Valvae with costa bearing subapical spine; sacculus with
numerous stout setae; cucculus rounded, membranous, sm-passing
costa. Aedeagus flattened, vesica unarmed.
Female genitalia A\'ith ovipositor moderately setose, caudal margin
finely setose. Apophyses fairly straight, of about equal length; well
developed; base of posterior shaped like a parallelogram; anterior
curved downward near base. Bursa unarmed. Ductus seminalis from
posterior end of bursa.
REVISION OF PEORIENAE AND ANERASTIINAE 49
Atascosa glareosella (Zeller)
FiC4URES 26, 54, 91, 126, 163
Anerastia glareosella Zeller, 1872, pp. 553-554.
Saluria glareosella (Zeller). — Ragonot, 1889, p. 117. — Hulst, 1890, pp. 211-
212. — Smith, 1891, p. 85. — Hampson, 1918, pp. 95-96.
Atascosa bicolorella Hulst, 1890, pp. 210, 227. — Smith, 1891, p. 85. — Rindge,
1955, p. 158.
Maricopa albocostella Hulst, 1900, p. 176 [described in Phycitinae]; 1902, p.
438. — Rindge, 1955, p. 157. — Heinrich, 1956, p. 316 [transferred to Aner-
astiinae]. [New synonymy.]
Poiijadia glareosella (Zeller). — Ragonot, 1901, pp. 345-346. — Barnes and
McDunnough, 1917, p. 149. — Forbes, 1923, p. 638. — McDunnough,
1939, p. 36.
Atascosa glareosella (Zeller). — Hulst, 1902, p. 438.
Valdiva albocostella (Hulst). — Barnes and McDunnough, 1917, p. 148. — Mc-
Dunnough, 1939, p. 34.
Diagnosis. — This is the only known species in the genus.
Description. — Frons dark brown, with red posterodorsal patch;
labial palpi with basal segments white, second and third segments
dark brown on outer sides, ventrally white sprinkled with red; anten-
nae with scapes white on outer sides, mixture of red and brown on
inner sides, shaft light brown; vertex and occiput brown, lighter
dorsally; patagia, tegulae, a,nd dorsum of thorax dark brown, usually
sprinkled with red.
Forewing with white costal band extending posterior to middle of
cell, narrowing to point at apex, sprinkled with grayish red, grayish-
red basal dash; white band bordered posteriorly by dark brown band,
mixture of grayish red and dark brown distal to cell; posterior of
grayish red and dark brown distal to cell; posterior third of wing
grayish red sprinkled with brown; transverse posterior band dark
brown; discoceUular traced with dark brown, forming discal spot;
fringe brown, with three white lines. Hindwing Hght brown, darker
toward apex.
Genitalia as described for the genus.
Types. — A. glareosella, in the British Museum (Natural History),
(from Zeller collection); A. hicolorella and M. albocostella, in the
American Museum of Natural History.
Type data. — A. glareosella, lectotype female, hereby designated,
labeled as follows: "Type; 15/8; Bosque Co. Texas; Zell. Coll. 1884.;
Anerastia glareosella Z. N.A.I. 553 [green label in Zeller's handwrit-
ing]; 9 genitalia slide 1-4-1967 J. Shaffer No. 709."
A. hicolorella, lectotype male, hereby designated, Blanco County,
central Texas, August, collection G. D. Hulst; genitalia slide No.
3222, Carl Hemrich, June 14, 1947.
50 U.S. NATIONAL MUSEUM BULLETIN 280
M. albocostella, lectotype male, hereby designated, Anglesea, N.J.,
August 21?, collection G. D. Hulst, genitalia slide No. 3219, Carl
Heinrich, June 14, 1946. Lectoparatype male, Anglesea, N.J., August
21?, collection G. D. Hulst.
Specimens examined. — 44 cf, 18 9.
Distribution (Map 12). — Atlantic and Gulf Coastal Plain, Texas
to Florida, north to Massachusetts.
UNITED STATES: Colorado: El Paso Co., Colorado Springs, Fountain
Valley School, 1 cf , July 7, 1935 [AMNH] (see discussion section).
Connecticut: New Haven Co., East River, 1 cT, Aug. 12, 1907 (Charles R.
Ely) [USNM]; 1 c?, Aug. 16, 1907; 2 c?, July 19, 1908; 2cf, July 20, 1908; 1 cf,
July 22, 1908; 1 d', July 24, 1908; 2 cr', 1 ? , July 27, 1908; 1 cf , July 28, 1908;
1 cf, July 30, 1908; 1 cT, Aug. 1, 1908; 1 9, Aug. 2, 1908; 1 ?, Aug. 7, 1909;
1 cf, Aug. 10, 1909; 1 d^, Aug. 12, 1909; 1 d^, 1 ? , July 1910; 1 9 , July 19, 1912;
1 d", Aug. 1, 190?.
Florida: Dade Co., Homestead, 1 cf, Aug. 7, 1963 (D. O. Wolfenbarger)
[CPK].
Massachusetts: Barnstable Co., Barnstable, 1 cf, July4, 1949 (C. P. Kimball)
[CU]; 1 d, July 10, 1949 (C. P. Kimball) [CU]; 2 d, July 10, 1949 (C. P. Kimball)
[CPK]; 1 9, July 12, 1949; 1 d, Aug. 1, 1952; West Barnstable, 3 d, July 15,
1949 (C. P. Kimball) [CPK]; 1 d, July 16, 1949 (C. P. Kimball) [CNC]; 2 d
4 9, [CU]; 1 d, July 17, 1949 (C. P. Kimball) [CU]; 1 &, July 18, 1949 (C. P.
Kimball) [CPK]; 1 9 , July 20, 1949; Dukes Co., Martha's Vineyard, 1 d, Aug. 11,
1946 (F. M. Jones) [CNC]; 1 d, July 18 (F. M. Jones) [CPK].
New Jersey: Bergen Co., Oakland, 1 d, July 26, 1947 (C. P. Kimball) [CPK];
1 d, July 27, 1947 (C. P. Kimball) [CU]; 1 9, Aug. 4, 1948 (C. P. Kimball)
[CPK]; 1 cf, Aug. 9, 1948 (C. P. Kimball) [CU]; 1 d, Aug. 13, 1948; 1 d, Aug.
14, 1948; 1 cf , Aug. 24, 1948; Burlington Co., New Lisbon, 1 d, July 31, 1931
(E. P. DarUngton) [ANS]; 1 9 , July 14, 1933; 1 cT, Aug. 10, 1939; 1 9 , July 31,
1942; 1 cf , Aug. 4, 1942; Morris Co., Mendham, 1 d, Aug. 3, 1935 (G. H. Tate)
[AMNH]; Ocean Co., Lakehurst, Wrangle Brook Road, 1 9, Aug. 30, 1956
(J. G. Franclemont) [JGF].
New York: Suffolk Co., Riverhead, 1 cf, July 1, 1933 (Roy Latham) [CU].
North Carolina: Polk Co., Tryon, 1 9, Aug. 11, 1904 (Fiske) [USNM];
1 9 , Aug. 14, 1904.
Discussion. — The Colorado specimen has wang venation and male
genitalia typical of glareosella, but differs in having uniform brownish-
orange forewings, unmarked except by a faint discal spot, and in being
slightly larger than typical members of the species. The specimen may
or may not be glareosella.
Homosassa Hulst
H(mosassa Hulst, 1890, p. 214. — Smith, 1891, p. 85. — Hulst, 1902, p. 440.
[Type: Ephestia ella Hulst, 1887. Monobasic]
Diagnosis. — The long, straight, unbranched ventrolaterally di-
rected spicate processes of the uncus are diagnostic of the genus.
REVISION OF PEORIINAE AND ANERASTIINAE 51
Description. — Labial palpi of males obliquely ascending w^th
third segment somewhat decumbent, two to 2}^ times as long as eye
diameter, females porrect, three times eye diameter; maxillary palpi
small; tongue rudimentary; antennae sublaminate and ciliate in
males, filiform and finely ciliate in females; oceUi normally developed.
Foremng ^nth 10 or 11 veins; R2 free from cell or stalked ^nth
K3+4; M2+3 stalked or fused, from lower outer angle of ceU; Cui from
just before the angle. Hindwing with 7 veins; Sc and Rs stalked; Mi
from upper outer angle; M2 lost; M3 stalked with Cuj, from lower
outer angle of ceU; Cu2 from just before the angle.
Male genitalia \vith spicate processes of uncus straight, unbranched,
ventrolaterally directed, tapering to broad triangular base.
Juxta shield-shaped. Valvae subrectangular vnth costa projecting,
bluntly roimded and flattened apically; sacculus rather densely setose.
Aedeagus with thickened ring on posterior end.
Female genitalia with ovipositor strongly compressed. Apophyses
rather straight, about equal in length. Bursa unarmed. Ductus
seminalis from posterior end of bursa.
Key to the Species of Homosassa
1. M2+3 fused (fig. 58) ella
M2+3 stalked (fig. 54) 2
2. Male with uncus bearing well sclerotized ring (fig. 129) incudella
Male with uncus not bearing sclerotized ring (fig. 128) platella
Homosassa ella Hulst
Figures 27, 58, 92, 127, 164
Ephestia ella Hulst, 1887. — Rindge, 1955, p. 161.
Anerastia ella (Hulst). — Ragonot, 1889, p. 117; 1901, p. 400. — Barnes and
McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36. — Kimball, 1965,
p. 251.
Homosassa ella Hulst, 1890, pp. 214, 227. — Smith, 1891, p. 85. — Hulst, 1902
p. 440. — Grossbeck, 1917, p. 134.
Rhinaphe ella (Hulst). — Hampson, 1918, p. 86.
Diagnosis. — The loss of vein M2 distinguishes this species from
the other two members of the genus.
Description. — Frons conical, dark brown; labial palpi ^nth basal
segments white, second and third segments dark brown dorsally,
white ventrally and on inner sides; antennae with scape dark brown
anteriorly, white posteriorly, shaft light brown; vertex, occiput,
patagia, tegulae, and dorsum of thorax brown to dark brown; legs
brown, lighter on inner sides, tarsal scales white tipped.
ForcAnng \nth white band anterior to cell, tapering to point at
apex; proximal half of band margined anteriorly with dark brown line
broadened at base; white band usually sprinkled ^^^th red; ground
52 U.S. NATIONAL MUSEUM BULLETIN 280
uniform dark brown posterior to white band, sometimes with reddish
hue posterior to Ai fold; 10 veins; Ri from well before upper outer
angle of cell ; R2 free from cell or stalked mth R3+4 ; M2+3 fused. Hind-
wing brown or Ught brown, darker toward apex.
Male genitalia with uncus bearing well sclerotized ring, open
anteriorly and forming pair of ventrally directed sharp-pointed spines.
Gnathos (fig. 127) pi-shaped, transverse portion upturned and sharp,
pointed laterally. Aedeagus with vesica bearing single sharply serrate
inconspicuous cornutus.
Female genitalia as described for the genus.
Types. — In the American Museum of Natural History (lectotype) ;
in the U.S. National Museum (lectoparatype).
Type data. — Lectotype male, hereby designated, Florida, collection
G. D. Hulst; genitalia slide No. 3225, Carl Hemrich, June 14, 1946.
Lectoparatype: Male, Fernald collection, USNM 40079; genitalia
slide No. 572, J. Shaffer, Nov. 23, 1965.
In the original description Hulst lists three males and two females
from Florida.
Specimens examined. — 90 cf, 42 9.
Distribution (Map 11). — Atlantic and Gulf Coastal Plain, eastern
Texas to Florida, north to North Carolina.
UNITED STATES: Alabama: Mobile Co., Theodore, 1 d', June 12, 1917,
[CU].
Florida: Alachua Co., Gainesville, 1 cT, 29, July 8, 1927 (J. Speed Rogers)
[CU]; 1 cf, July 10, 1927; 1 cf, Apr. 22, 1952 (O. Peck) [CNC]; Charlotte Co.,
Punta Gorda, 2 9 , Apr. 12, 1952 (G. S. Walley) [CNC]; Dade Co., Florida City,
2 d', Apr. 17, 1947 (Otto Buchholz) [ANS]; 1 cf, Apr. 20, 1947; Homestead,
1 &, Feb. 24,1959 (D. O.Wolfenbarger) [CPK]; 1 d^, Apr. 15, 1959; 1 &, Apr. 16,
1959; 2 cf, July 17, 1959; 1 cf , Feb. 22, 1955; 1 c?, July 29, 1958; Royal Palm
Hammock (also called Paradise Key, or Royal Palm State Park), Everglades
National Park,ld^, Apr. 11 (F. M. Jones) [ANS]; 1 cf, 1930; 1 d'. Mar. 15, 1938
(E. P. Darlington) [ANS]; 1 cT, Mar. 17, 1938; 1 &, Mar. 20, 1938; 2 cf, 2 9
Apr. 2, 1952 (G. S. Walley) [CNC]; 1 9 , Apr. 3, 1952; Flager Co., Pellicer Crk.
13 mi. north Bunnell, 1 9 , May 8, 1954 (J. Bauer) [CM]; 1 cf, Apr. 10, 1954
1 9 , Apr. 24, 1954; Highlands Co., Archbold Biol. Sta., 1 9 , Apr. 27, 1947
(J. G. Needham) [CU]; 1 d^, Apr. 28, 1947; 2 9, May 14, 1947; 2 9, June 17
1964 (Jay C. Shaffer) [JCS]; 1 d^, 3 9 , June 18, 1964; 1 d', 1 9 , June 20, 1964
2 d", 1 9 , June 22, 1964; 2 d", June 25, 1964; 2 d", 1 9 , June 26, 1964; 2 d"
June 28, 1964; 1 9 , June 29, 1964; 1 9 , July 2, 1964; 1 d^, July 3, 1964; 1 d^
July 5, 1964; 1 d', July 11, 1964; 1 9, July 12, 1964; 1 d", July 13, 1964
Hillsborough Co., Stemper, 1 d", Aug. 19, 1912 (G. Krautwurm) [CM]; 2 d"
1 9, Sept. 1, 1912; 2 d^, Sept. 4, 1912; 2 d^, Sept. 5, 1912; 2 d", Sept. 6, 1912
1 d', Sept. 9, 1912; 1 d^, 1 9 , Sept. 13, 1912; 1 9 , Oct. 1, 1912; 1 d", Oct. 8, 1912
1 d', Oct. 9, 1912; 1 d", June 4 [CM]; 1 9 , June 19 [CM]; Lake Co., Leesburg
1 d', May 14, 1961 (C. H. Curran) [AMNH]; Manatee Co., Gulf Coast Exp
Sta., Bradenton, 1 d". May 11, 1955 (E. G. Keisheimer) [CPK]; Oneco, 1 9
May 5, 1953 (Paula Dillman) [CPK]; 1 6", May 8, 1953; 1 9, May 19, 1953
1 d". May 25, 1953; 1 9 , June 6, 1953; 1 d", 1 9 , June 7, 1953; 1 9 , June 10,
REVISION OF PEORIINAE AND ANERASTIINAE 53
1953; 1 cf , July 15, 1953; 1 &, Oct. 15, 1953; 1 &, June 1, 1954; 1 9 , June 9,
1954; Pinellas Co., Dunedin, 1 &, Apr. 22, 1925 (W. S. Blatchley) [ANS]; 2 cT,
May 24, 1925; Sarasota Co., 1 d", May 7, 1946 (C. P. Kimball) [CPK]; 1 ? ,
May 12, 1946; 1 c?, May 19, 1946; Siesta Key, 1 9 , May 21, 1956 (C. P. Kimball)
[CPK]; St. Johns Co., Hastings, 6 cT, 2 ? , April [USNM]; 5 cT, 1 9, May
[USNM]; 3 d', 1 9, June [USNM]; 1 cT, Sept. 30 [USNM]; county unknown,
"Altmont," 1 9 , Sept. 21, 1924 (F. R. Colo) [CU]; locality unknown, 1 cf ,
no date [USNM].
Georgia: Charlton Co., 1 c?", June 7, 1946 (Otto Buchholz) [ANS
Id', June 9, 1946; 2d', June 10, 1946; 1 d', June 11, 1946.
Mississippi: Jackson Co., Biloxi, 1 d', June 13, 1917 [CU].
North Carolina: Brunswick Co., Leland, 1 9 , June 17, 1946 (Otto Buchholz)
[ANS]; 2 d", June 20, 1946; 1 9 , Sept. 2, 1946; Robeson Co., Maxton, 1 d',
May 1-15, 1943 (A. B. Klots) [AMNH].
Texas: Burnet Co., 1 d", no date (F. G. Schaupp) [USNM]; Nueces Co.
Corpus Christie, 1 d', Mar. 26, 1943 (W. M. Gordon) [CU].
Discussion. — In many specimens the distal one-half of each spine
of the uncus ring is bent anteroventraUy (fig. 127), and the anteriorly
directed spine of the gnathos is posterior (fig. 127a) in relation to that
of the type (fig. 1276). Specimens of both types have been collected
at the Archbold Biological Station and do not represent geographical
varieties. In view of the small differences between the two types, they
are held to be conspecific.
Homosassa platella, new species
Figures 28, 93, 128
Diagnosis. — The broad flat transtUla and lack of a sclerotized
ring on the uncus each will distinguish this species.
Description. — Maculation similar to that of ella. Forewing with
11 veins; M2+3 stalked.
Male genitalia with uncus lacking sclerotized ring.
Gnathos expanded into pair of large, flat, medially fused, posteriorly
bicuspitate or tricuspitate plates. Juxta four-sided with medial V-
shaped heavy sclerotization. Aedeagus with vesica bearing pair of
weakly sclerotized sharply serrate cornuti.
Female unknown.
Types. — In the U.S. National Museum.
Type data. — Holotype, male, Hastings, Fla. (Saint Johns Co.),
March, collection W. D. Kearfott; USNM 69385; genitalia slide
No. 596, J. Shaffer, Apr. 26, 1966.
Paratypes: Three males, Hastings, Fla., collection W. D. Kearfott;
two dated March, one male genitalia slide No. 103, J. C. S., Dec. 30,
1963; one dated April, male genitalia slide No. 493, J. Shaffer, Apr. 13,
1965.
Other specimens examined. — 4 cT.
Distribution (Map 11). — Known only from Florida.
54 U.S. NATIONAL MUSEUM BULLETIN 280
UNITED STATES: Florida: Collier Co., Everglades, 1 cf, Apr. 6, 1912
[AMNH]; Dade Co., Homestead, 1 d', Apr. 28, 1952 (J. R. Vockeroth) [CNC];
Pinellas Co., Dunedin, 1 d', Feb. 23, 1921 (W. S. Blatchley) [ANS]; Saint Johns
Co., Hastings, 1 cf , May [AMNH].
Homosassa incudelkif new species
Figures 29, 94, 129, 165
Diagnosis. — The sclerotized ring of the uncus bears a pair of two-
pointed processes rather than a simple process as in ella, or with the
ring absent as in platella. Female specimens may be distinguished from
those of ella by the presence of a dorsal invagination on the eighth
abdominal segment.
Description. — Maculation similar to that of ella. Forewing with
white costal band sewn with dark brown scales; 11 veins; M2+3 stalked.
Male genitalia with uncus similar to that of ella, but anterior
processes of ring each anvil-shaped with sharp-pointed anterior and
posterior processes. Gnathos with pair of subapical thickened proc-
esses, each bearing anterior and posterior spines. Aedeagus with
vesica unarmed.
Female genitalia with dorsal invagination on eighth abdominal
segment, and smaller invagination dorsally on the membrane between
segments eight and nine.
Types. — In the Canadian National collection.
Type data. — Holotype, male, Lake Murray, Love and Carter
Counties, Oklahoma, May 20, 1950, W. J. Eeinthal, "339/50";
C.N.C. Type No. 9441; genitalia slide No. 546, J. Shaffer, July 15,
1965. A mass of glue supports the specimen on the pin from beneath,
and I have added a small amount of glue beneath the right wings to
support them.
Paratype: Female, data as given for the holotype; genitalia sHde
No. 548, J. Shaffer, July 16, 1965.
Other specimens examined. — None.
Distribution (Map 11).
Reynosa-) new genus
Type. — Atascosa Jloscella Hulst, 1890.
Diagnosis. — The short, stout, sharp-pointed spicate processes of the
uncus are diagnostic of the genus.
Description. — Frons conical; labial palpi porrect, about twice as
long as eye diameter in males, about 2J^ times in females; maxillary
palpi reaching frons or nearly so, spreading; tongue not exposed
between palpi; antennae filiform to subserrate in both sexes, com-
pressed, ciliate ventrally, male shaft with two basal segments fused;
ocelli present.
REVISION OF PEORIINAE AND ANERASTIINAE 55
Forewing with 11 veins; R2 usually free from cell, sometimes stalked
with R3+4; Ml from upper outer angle of cell; M2+3 stalked about one-
half their length, from lower outer angle; Cui from just before the
angle. Hind wing with 7 veins; Sc and Rs well stalked; Mi from upper
outer angle of cell; M3 and Cui long stalked, from lower outer angle;
Cui from cell very near to M3 and Cui.
Male genitaHa with uncus bearing pair of short apical lobes; spicate
processes a pair of short, stout, sharp-pointed anteriorly directed,
ventral hooks. Gnathos bearing small, sharp-pointed, posteriorly
directed apical and pair of subapical hooks. Juxta subquadrate, an-
terior margin convex, posterior concave. Vinculum well developed,
rounded. Valvae with cucullus membranous and projecting beyond
costa; sacculus rather densely pilose. Aedeagus untapered, vesica
unarmed.
Female genitalia with ovipositor compressed. Apophyses rather
straight, anterior and posterior about equal in length. Ductus bursae
short. Bursa unarmed, with long slender neck. Ductus seminalis from
posterior end of bursa, slender.
Reynosa floscella (Hulst), new combination
Figures 22, 95, 130, 166.
Atascosa floscella Hulst, 1890, pp. 210-211. — Smith, 1891, p. 80. — Hulst, 1902,
p. 438. — Rindge, 1955, p. 162.
Poujadia floscella (Hulst). — Ragonot, 1901, p. 346. — Barnes and McDunnough
1917, p. 149. — McDunnough, 1939, p. 36.
Saluria floscella (Hulst). — Hampson, 1918, p. 96.
Diagnosis. — This is the only known species in the genus.
Description. — Labial palpi Avith basal segments white, second and
third segments brown to reddish brown on outer sides, white on inner
sides; frons, occiput, patagia, and tegulae brown.
Forewing with ground light brown; costal area and ceU white,
sprinkled with reddish-brown scales; prominent dark brown orbicular
spot and line extending posteriorly from spot to inner margin forming
second spot on Ai fold; dark brown transverse posterior beginning
mth prominent spot between Mi and Cui, continuing to inner margin;
terminal line of dark brown dots. Hindwing light brown on both sides.
Genitalia as described for the genus.
Types. — In the U.S. National Museum (lectotype) ; in the American
Museum of Natural History (lectoparatype) .
Type data. — Lectotype female, hereby designated, Blanco County,
central Texas; Fernald collection; USNM 40078; genitalia slide
No. 594, J. Shaffer, Apr. 23, 1966.
Lectoparatype: Male?, Blanco County, central Texas, collection
G. D. Hulst; abdomen lost.
56 U.S. NATIONAL MUSEUM BULLETEST 280
Specimens examined. — 10 cf, 12 9.
Distribution (Map 12). — Known only from Gulf Coastal Plain
of Texas.
UNITED STATES: Texas: Cameron Co., Brownsville, 1 9, May 4, 1904
(H. S. Barber) [USNM]; 1 ? , no date [USNM]; 1 d", Mar. 19, 1937 (T. N.
Freeman) [CNC]; 2 d^, Mar. 22, 1937; San Benito, 1 9, June 16-23 [USNM]
1 9, July 8-15 [USNM]; 1 9, July 24-31 [USNM]; 1 9, August [USNM]
1 9 , Aug. 1-7 [USNM]; 1 cf , 1 9 , Sept. 8-15 [USNM]; 1 cf , Mar 16-23 [USNM]
Fort Bend Co., Richmond, Brazos River, 1 cf, June 22, 1917 [CU]; Nueces
Co., Corpus Christie, 1 cf, 2 9, Apr. 15, 1943 (W. M. Gordon) [CU]; 1 d'
May 22, 1943; 1 9, June 4, 1943; 1 cf, June 20, 1943; 1 cf , 1 9, Sept. 25-Oct.
15, 1943.
Goya Ragonot
Goya Ragonot 1888, p. 43. [Type: Goya albivenella Ragonot, 1888. Designated
by Hampson in Ragonot 1901, p. 349.]
Diagnosis. — The unusual male genitalia make this genus quite
distinctive within the subfamily; the presence of a transtilla and
lateral subrhomboid processes of the uncus are both unique to Goya.
Description. — Labial palpi about twice as long as eye diameter;
tongue rudimentary; antennae with basal segments fused in male;
eye large; oceUi present.
Forewing with 1 1 veins ; R2 free from cell near R3+4 or stalked with
the latter; Mi from upper outer angle of cell; Mz+z stalked, from lower
outer angle. Hindwing with 7 veins; Sc and Rs well stalked; Mi from
upper outer angle of cell; M3 and Cui stalked, from lower outer angle;
Cu2 from cell very near to M3 and Cui.
Male genitalia with uncus in form of pair of triangular plates,
each joined on its anterior angle to a large subtriangular process
which distally ciu-ves medially in tapering to a blunt point. Transtilla
present, incomplete. Gnathos with medial process subquadrate in
ventral view. Juxta rounded. Vinculum round to subtruncate. Valvae
rectangular, apex rounded, inner half somewhat concave. Aedeagus
short, compressed, with vesica unarmed.
Female genitalia with apophyses slender, rather straight. Bursa
unarmed. Ductus seminalis from posterior end of bursa, slender.
Goya stictella (Hampson), new combination
Figures 23, 96, 131
Saluria stictella Hampson, 1918, pp. 96-97.
Diagnosis. — This species is easily distinguished from albivenella
by the presence of dark brown spots on the forewing and by the
absence of well-marked white tracing on the forewing veins.
Description. — Labial palpi porrect, basal segments white, second
and third segments light brown on outer sides, third decumbent;
REVISION OF PEORIENAE AND ANERASTIINAE 57
antennae sublaminate in male, light brown, basal segments bearing
tuft of scales.
Forewing with M2+3 stalked for about one-half their length ; ground
light brown to grayish red, predominantly white anterior to cell;
maculation variable; transverse anterior and median lines marked
by dark brown spots in cell; Ai with two prominent dark brown spots,
one just distal to transverse anterior in the cell, other on the mod-
erately well-developed transverse posterior; terminal line of dark
brown spots. Hindwing with M3 and Cui long stalked.
Male genitalia with triangular plates of uncus each bearing on its
terminal angle a posteromedially directed thorn. Gnathos with medial
process bearing short sharp anterior and posterior spines on each
lateral margin. Juxta broader than long. Aedeagus tapering strongly
posteriorly.
Female genitalia as described for the genus.
Type. — In the British Museum (Natural History).
Type data. — Lee to type male, hereby designated, labeled as fol-
lows: "Type; Bahamas. M.C. Andros. 11. I. 1902 J.L. Bonhote. 1902-
278; Saluria stictella Type cf. Hmpsn,; Pyralidae Brit. Mus. Slide
No. 10904."
Specimens EXAMINED. — 5 cf , 2 9.
Distribution (Map 4). — In the United States the species is known
from Arkansas, Mississippi, and west central Florida.
UNITED STATES: Arkansas: Washington Co., Devel's Den State Park,
1 d^, May 30, 1966 (R. W. Hodges) [USNM].
Florida: Manatee Co., Oneco, 1 <f,l ? , June 13, 1954 (Paula Dillman)
[USNM]; 1 9 , Aug. 3, 1953; Sarasota Co., Siesta Key, 1 cf , Apr. 2, 1952 (Charles
P. KimbaU) [CNC].
Mississippi: Harrison Co., Biloxi, 2 d', June 13, 1917 [CU].
Discussion. — This is the only species of Goya known to occur in
the United States; the type locality of albivenella is Argentina.
Subfamily Phycitinae Ragonot
The following seven genera which were previously placed in the
Anerastiinae and a new genus, Wakulla, established for Bandera
carneella, are transferred to the Phycitinae.
The relationships within the Phycitinae are clear for only a few of
the eight genera. Anerastia and Coenochroa show affinities with each
other but none with any particular group in the Phycitinae. Barberia is
closely related to Anderida, and Bandera to Aiiagasta. The other four
genera have typical phycitine genitalia, although their exact place-
ment within the subfamily is uncertain. Of these four, Ragonotia and
Martia are closely related and have typical phycitine wing maculation.
58 U.S. NATIONAL MUSEUM BULLETIN 280
Anerastia Iliibner
Anerastia Hubner [1816H1826], p. 3G7. — Ragonot, 1901, pp. 394r-397. — Spuler,
1904, p. 200 — McDunnough, 1939, p. 36. [Type: Tinea lotella Hubner
[1810]-[June 20, 1813.] Designated by Moore, 1886, p. 357.]
Prinanerastia Hampson, 1918, p. 80. [Type: Tinea lotella Hubner. Original
designation.]
Diagnosis. — -Anerastia and two other North American genera
possess a toothed frons. In Coenochroa and Martia the teeth are borne
at the end of a well-developed protuberance; in Anerastia they form a
ring on an otherwise smooth frons.
Description. — -Frons rounded, fine projecting serrations forming a
transverse elhptical ring about one-half as wide as frons, vesture short;
labial palpi deflexed, about 3)^ times eye length in male, about foiu-
times in female; maxillary palpi rudimentary, minute; tongue re-
duced; antennae compressed, male shaft with basal segments fused,
sublaminate, ciha about one-fifth segment width, female filiform, cilia
minute, each segment bearing on its distal haK several longer ciha,
each about one-half segment width; ocelli well developed.
Fore wings mth 10 veins; Ri from well before upper outer angle of
cell; R2 free, just before the angle; R3 stalked for about one-half its
length onto R4; Mi from below the angle; M2+3 fused, well separated
from Cui; Cu2 from well before Cui at angle. Hindwing with 7 veins;
Sc and Rs approximate beyond cell, then diverging; IVIa and Cui
stalked for over one-half their length, from lower outer angle of cell;
Cu2 from just below the angle.
Male genitaha with uncus bilobed, membranous medially. Gnathos
complex, medial process stout, subrectangular, posteriorly bearing
very short truncate lateral arms, mediodorsal process, and strong
anteroventrally directed hook, dorsoanteriorly with large curved
flat hook between uncus lobes. Transtilla crescent-shaped, poorly
developed. Juxta eUiptical, anterior margin sclerotized. Vinculum
broadly rounded. Valvae tapering on distal thu-d to bluntly pointed
apex of costa, anterior-facing crescent-shaped ridge near base of taper-
ing portion. Aedeagus Avith vesica bearing two large cornuti, one
lance-shaped, the other deeply bicuspidate.
Female genitaha with ovipositor well sclerotized, tapering to bluntly
pointed tip. Dorsum of eighth abdominal segment deeply incised
medially. Apophyses well developed, anterior very shghtly shorter
than posterior. Ductus biu"sae moderate. Bursa unarmed. Ductus
seminahs from posterior end of biu-sa.
Discussion. — This Old World genus is represented in North
America by lotella, a species common in Europe.
REVISION OF PEORimAE AND ANERASTIINAB 59
Anerastia lotella Hiibner
Figures 34, 48, 64, 97, 132, 167
Tinea lotella Hubner [1810]-[June 20, 1813].
Anerastia lotella Hubner [1826], p. 367. — Buckler, 1901, pp. 203-206, pi. 156,
fig. 4. — Ragonot, 1901, pp. 397-398. — Spuler, 1904, p. 200, pi. 82,
fig. 2. — Beirne, 1952, pp. 74-75, pi. 6, fig. 1.
Prinanerastia lotella (Hubner). — Hampson, 1918. p. 80.
Anerastia lobelia [sic] (Hubner). — McDunnough, 1939, p. 36.
Diagnosis. — This is the only North American species in the genus.
Description. — Labial palpi on outer sides varying bro\vn to light
brown, light orange, or orange; antennae orange browTi; frons light
brown, darker laterally, vertex and occiput brownish white dorsally,
occiput laterally, patagia, tegulae, and dorsum of thorax brown to
orange brown; legs brown to orange brown on outer sides, brownish
white on inner sides.
Forewing varying brown to orange; veins white traced, sprinkled
with brown; A2 broadly sprinkled with brown; Ai fold lightly white
traced on outer half. Hindwings brown, fringe light brown.
Genitalia as described for the genus.
Type. — Lost.
Type locality. — Not given, but presumably Europe.
Recorded hosts. — Poaceae : Ammophila arenaria (L.) Link
(European beach grass), larvae feeding on roots near junction \vith
stem; also on Festuca ovina L., Secale cereale L., (Rye) to which they
are sometimes a serious pest, and Aira species, (Buckler).
Specimens examined. — 152 c?', 45 9 .
Distribution (Map 1). — Alaska (Fort Yukon) south to Washing-
ton, east to Minnesota.
UNITED STATES: Alaska: near Fort Yukon, 1 cT [USNM].
Minnesota: Cass Co., Cass Lake, 1 cf June 18, 1934 (A. A. Granovsky) [UM]
Icf, June 27, 1934; 2<^, July 27, 1936 (R. H. Daggy) [UM]; 1 cf , Aug. 2, 1936
Cass Co., 1 d^, June 24, 1937 (L. W. Orr) [UM]; 1 d', June 26, 1937; 2 c?, June 30
1937; Icf, July 1, 1937 (P. M. Schroeder), 1 cT, July 11, 1937 (L. W. Orr); Id'
Aug. 11, 1937; Kittson Co., HaUock, Ic?', Aug. 9, 1935 (D. G. Denning) [UM]
Id", June 20, 1936; Sd, June 21, 1936; 2d, JiUy 7, 1937; Polk Co., Crookston
1 d, July 20, 1935 (D. G. Denning) [UM]; Id, July 21, 1935; 2 cf, July 22, 1935
Id, July 27, 1935; 2d, June 21, 1937; 1 9, June 23, 1937; 3^^, June 27, 1937
1 d, June 26, 1937; 1 d, June 26, 1941; 1 d, June 27, 1941.
Washington: WaUa WaUa Co., Wallula, 1 d, July 30, 1948 (William C. Cook)
[CNC].
CANADA: Alberta: Banflf, Id, June 29, 1922 (C. B. D. Garrett) [CNC];
Dominion Range Sta. Manyberries, 1 d, July 10, 1951 (D. F. Hardwick) [CNC];
Scandia, Id, July 9, 1956 (E. E. Sterns) [CNC]; Id, July 10, 1956.
Manitoba: Aweme, Id, July 26, 1914 (N. Criddle) [CNC]; 3d, July 6, 1920;
Id, 19, July 17, 1920; Id, 4 9, June 28, 1921; Id, 19, July 1, 1921; 19,
(P. N. Vroom); 1 cf , 6 9 , July 2, 1921 (N. Criddle) [CNC]; 3d^, 4 9 , July 6, 1921;
3 9, July 26, 1921; 1 d, July 28, 1921; 3 9. Aug. 3, 1921 (P. N. Vroom) [CNC];
285-934—68 5
60 U.S. NATIONAL MUSEUM BULLETIN 280
1 cf, Aug. 10, 1921 (N. Criddle) [CNC]; 1 ^, Aug. 25, 1921; 3 cf', 3 9, June 13, 1922;
2 9, June 23, 1922; 2 cf , June 27, 1922; 19, July 19, 1922; 1 cf, July 22, 1922;
2cf, Sept. 15, 1923; 1 c?, July 11, 1925; 1 cf, Aug. 10, 1925; 19, Aug. 21, 1925;
3d', 2 9, July 2, 1926; 19, Aug. 11, 1926; 7 <^, July 20, 1928; Brandon, 1 rf",
June 27, 1958 (R. B. Madge) [CNC]; 1 cf', July 6, 1958 (R. L. Hurley) [CNC];
1 9 , July 7, 1958 (R. B. Madge) [CNC]; 1 d^ July 8, 1958; 5 d^, 2 9 , July 17, 1958;
(R. L. Hurley) [CNC]; 2 9, July 20, 1957 (R. B. Madge) [CNC]; Id', July 31,
1958 (R. L. Hurley) [CNC]; 19, Aug. 7, 1958 (R. B. Madge) [CNC]; Glenboro,
38 d', "Spruce-sand community," June 17, 1958 (R. L. Hurley) [CNC]; Ninette,
Id', "Maple-Elm floodplain community," June 17, 1958; Wabowden, Id", Aug.
7, 1949 (J. B. Wallis) [CNC].
Northwest territories: Bathurst Inlet, 1 9, July 28, 1951 (W. I. Campbell)
[CNC]; Hay River, 2d^, July 30, 1951 (P. R. Ehrlich) [CNC]; Id", July 31, 1951.
Quebec: ForestviUe, Id", July 11, 1950 (R. deRuette) [CNC].
Saskatchewan: Attons Lake, Cut Knife, Id', June 23, 1940 (A. R. Brooks)
[CNC]; Od', July 11, 1940; 2d^, 19, July 12, 1940; Rutland, 5d^, July 15, 1940
(A. R. Brooks) [CNC]; 1 d', Aug. 2, 1940; Saskatoon, 1 d', June 26, 1923 (Kenneth
M. King) [CNC]; 1 d^, July 2, 1923; 2d', July 7, 1923; 1 d^, July 21, 1923; Was-
kesiu Lake, 19, July 11, 1939 (A. R. Brooks) [CNC]; 2d', July 13, 1939; 2d',
19, July 15, 1939; 1 d^, July 18, 1939; 2d', 19, July 21, 1939.
Coenochroa Ragonot
Coenochroa Ragonot, 1887, p. 20; 1889, p. 117, — Hulst, 1890, p. 217. — Smith,
1891, p. 85. — Ragonot, 1901, pp. 418-419. — Hulst, 1902, p. 441. — Barnes
and McDunnough, 1917, p. 150. — Hampson, 1918, p. 58. — McDunnough,
1939, p. 36. [Type: Coenochroa californiella Ragonot, 1887. Original
designation.]
Petaluma Hulst, 1888, p. 116; 1890, pp. 215-216. — Smith, 1891, p. 85. — Hulst,
1902, p. 440. [Type: Anerastia ilUhella Hulst, 1887. Original designation.]
Alamosa Hampson in Ragonot, 1901, p. 369. — Barnes and McDunnough, 1917,
p. 149. — Hampson, 1918, p. 65. — McDunnough, 1939, p. 36. [New synon-
ymy.] [Ty^Q-. Alamosa piperatella Hampson in Ragonot, 1901. Monobasic]
Diagnosis. — Tlie genus may be easily recognized by the combina-
tion of a toothed protuberant frons and longitudinal wing pattern.
Martia has a somewhat similar frons, but the wing pattern is predom-
inantly transverse. The wing maculation of Anerastia is similar to
that of Coenochroa, and the frons is toothed, but not protuberant.
The bifid male uncus is unique to Coenochroa.
Description. — Frons with prominent anterior protuberance ter-
minating in dorsoposteriorly sloping closed rim of irregular teeth sur-
rounding central beak (figs. 44-47); labial palpi porrect or deflexed;
maxillary palpi rudimentary, naked, usually hidden by labials; tongue
reduced, usually hidden by palpi; antennae compressed, each segment
bearing pair of perpendicidarly projecting ciha ventrally near distal
end, male shaft scaled dorsally, ventrally and laterally bearing dense
fine recurved ciha, female shaft scaled dorsally and laterally, ventrally
with fine sparse recurved ciha; oceUi small, covered with scales.
REVISION OF PEORIINAE AND ANERASTIINAE 61
Forewings Avith 10 veins; vein Cu2 from before lower outer angle of
cell; Cui and M3 from angle, stalked; M2 absent; Mi from upper outer
angle of cell; R2+4 stalked, from just before the upper outer angle; Ri
from well before angle. Hindwing with 6 or 7 veins; vein Cu2 from be-
fore lower outer angle of cell; Cui and M3 from angle, stalked for at
least one-half their length, sometimes fused; M2 absent; Mi from just
below upper outer angle of cell; Rs and Sc from upper angle, stalked
for at least one-half their length.
Male genitaha with uncus bifid, setose ventrally, terminating in
minute ventrally directed hook. Gnathos with medial process U-
shaped. Transtilla absent. Juxta membranous, margin sclerotized
anteriorly. Vinculum variable, U-shaped or V-shaped, anteriorly
rounded or bluntly pointed. Valvae broadly rounded, variable in out-
Une, setose distally. Aedeagus tapering posteriorly, vesica armed.
Female genitaha with ovipositor lobes triangular, moderately setose.
Apophyses rather straight and uniform. Ductus bursae moderately
broad. Bursa slender and unarmed. Ductus seminaHs leaving near mid-
point of ductus bursa.
Key to the Species of Coenochroa Based Upon External Structure
1. Central beak of frons protruding beyond rim (figs. 45, 46); costa paler than
cell; R2 usually first to branch from stalked R2+4 illibella
Central beak of frons not protruding beyond rim; costa not pale; R^ usually
first to branch from stalked R2+4 2
2. Central beak of frons with irregular tip, just attaining end of rim (fig. 47) ;
Atlantic and Gulf Coastal Plain bipunctella
Central beak of frons pointed and small, not attaining end of rim (fig. 44) ;
Texas and Western United States californiella
Coenochroa californiella Ragonot
Figures 41, 44, 65, 98, 133, 168
Coenochroa californiella Ragonot, 1887, p. 20; 1889, p. 117. — Hulst, 1890, p. 217.—
Smith, 1891, p. 85. — Ragonot, 1901, p. 420. — Hulst, 1902, p. 441. —
Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 59. — McDun-
nough, 1939, p. 36.
Coenochroa inspergella Ragonot, 1887, p. 20; 1889, p. 117. — Hulst, 1890, p.
216. — Smith, 1891, p. 85. — Ragonot, 1901, p. 419. — Hulst, 1902, p. 440.—
Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 59. — McDun-
nough, 1939, p. 36. [New synonymy.]
Diagnosis. — This is the only member of the genus in which the
central beak of the frons does not attain the rim. The genitaha are
identical to those of illibella, a species which lacks the discal spot
and has a distinctive pale costa.
Description. — Frons with protuberance cylindrical, ventrally
about as long as wide, rim slanting about 45°, teeth projecting for-
ward, central beak a very small cone not reaching rim, vesture light
62 U.S. NATIONAL MUSEUM BULLETLNT 280
brown dorsally, white ventrally; labial palpi deflexed, basal segments
white, second and third uniform light yellow, occasionally brown
dorsally; antennae light brown, finely ciliate; patagia, tegulae, vertex,
occiput, and legs varying light yellow to brown.
Fore wing radius varying from about 8 mm to 12 mm, average of
9.5 mm; ground color light yellow to orange yellow; veins usually
traced with white, often sprinkled with black scales; dark discal spot at
lower outer angle of cell; vein R2 usually first to leave stalked R2+4.
Male genitalia with aedeagus slender, slightly thicker anteriorly,
vesica with a single slender conutus.
Female genitalia with ductus seminalis slender, not thickened
basally.
Types. — C. californiella and C. inspergella, in the Museum National
d'Histoire Naturelle.
Type data. — C. californiella, lectotype male, hereby designated,
Walsingham, genitalia slide No. 536, J. Shaffer, June 20, 1965. In
the original description the type locality is given as California.
C. inspergella, lectotype male, hereby designated, Arizona, Morrison,
1881, genitalia sHde No. 537, J. Shaffer, June 20, 1965.
Specimens examined. — 260 cf, 133 9 .
Distribution (Map 2). — British Columbia south to California,
east to Kansas and Texas.
UNITED STATES: Arizona: Cochise Co., Carr Canyon, Huachuca Mts.,
2 cf , June 3, 1952 (M. Cazier, W. Gertsch, R. Schrammel) [AMNH]; Ft. Grant,
Pinaleno Mts., 1 9, July 15-19, 1917 (C. U. Biol. Expend.) [CU]; Huachuca
Mts., 1 cf, May 1-7 [USNM]; 1 9, August 8-15; Montezume Pass, Huachuca
Mts. (6500 ft.), 1 cf, Sept. 7, 1950 (T. Cohn, P. Boone, M. Cazier) [AMNH];
Paradise, 1 cf, March [USNM]; Paradise, 1 9, July (Poling) [CM]; 1 <f, 3 9,
August; Portal, 1 9 , June 1, 1952 (M. Cazier, W. Gertsch, R. Schrammel)
[AMNH]; San Bernardino Ranch (3750 ft.), 1 9, August (F. H. Snow) [UK];
Southwestern Research Sta., 5 mi. west of Portal (5400 ft.), 1 cf , Apr. 10, 1956
(Cazier, Ordway) [AMNH]; 1 d', Apr. 22, 1956; 1 cT, Apr. 23, 1956; 1 cf, Apr.
25, 1956; 2 cf , May 16, 1956 (M. Statham) ; 1 cf , 2 9 , May 18, 1956; same locality,
1 9, July 28, 1959 (E. G. Linsley) [UCB]; same locality, 1 9, July 26-Aug. 3,
1959 (A. B. Klots) [ABK]; same locality, 1 9, Nov. 1, 1959 (J. R. Powers)
[UCB]; 1 cf, Nov. 2, 1959; same locaHty, 1 cf, May 16, 1960 (Carl W. Kirkwood)
[CPK]; 1 d', May 18, 1960; 1 9 , May 25, 1960; 1 cT, May 18, 1960; 1 9 , May 25
1960; 1 cf, May 28, 1960; 1 cf. May 29, 1960; 1 d, Apr. 10, 1961; 1 d, Apr. 12,
1961; 1 cf, Apr. 13, 1961; 1 d, Apr. 17, 1961; same locality, 1 9, Apr. 23, 1961
(M. A. Cazier) [AMNH]; 1 d, Apr. 25, 1961; 1 9, May 2, 1961; same locality,
1 d^, 2 9, May 10, 1961 (Gertsch & Cazier) [AMNH]; 1 cf, 1 9 , May 15, 1961
(M. Cazier); 1 cf, May 27, 1961 (M. Statham); same locality, 1 d, Apr. 10,
1962 (Carl W. Kirkwood) [CNC]; 4 d, Apr. 15, 1962; 1 d, Apr. 16, 1962; 1
d, Apr. 18, 1962; 1 d, Apr. 19, 1962; 2 cT, Apr. 20, 1962; 1 cf, Apr. 21, 1962;
2 cf, Apr. 22, 1962; 1 d, Apr. 23, 1962; 1 d, Apr. 27, 1962; same locality, 1 d,
Aug. 23, 1962 (M. Statham) [AMNH]; 1 d, Aug. 30, 1962; Coconino Co., Flag-
stafif, 2 cf , no date ( H. S. Barber) [USNM]; Gila Co., vicinity Globe, 3 9 , Aug.
4-5, 1937 (A. B. Klots) [AMNH]; Globe, 1 d, May 18, 1950 (E. C. Johnston)
REVISION OF PEORIINAE AND ANERASTIINAE 63
[CNC]; San Carlos, 1 d", May 12-13 (1918 (J. C. Bradley) [CU]; no locality,
1 9, no date (O. C. Poling) [USNM]; Maricopa Co., New River, 1 cf, May
7, 1950 (E. C. Johnston) [CNC]; Tempe, 1 cf, Apr. 19, 1920 (E. V. Walter and
H. L. Arnold) [USNM]; 1 d', Apr. 26, 1920; 1 9, May 3, 1920; 1 9, July 26,
1920; 1 9, Sept. 27, 1920 (E. V. Walter); Pima Co., Baboquivari Mts. (5-7000
ft.), 1 (^, July 15-30, 1923 (O. C. Poling) [USNM]; 1 9, July 27-31, 1923;
1 cT, 1 9, Aug. 1-4, 1923; 3 9, Aug. 1-15, 1923; 3 d^, 7 9 , Aug. 15-30, 1923;
1 9, Sept. 1-15, 1924; 1 d", Sept. 15-30, 1924; 11 cf, 12 9, Oct. 1-15, 1924;
9 cf, 1 9, Oct. 15-30, 1924; 1 d, Nov. 1-15, 1924; Baboquivari Mts., 3 cf,
2 9 , Apr. 23, 1938 (J. A. Comstock) [LACM]; 4 cf , 5 9 , Apr. 24, 1928;
14 cT, 1 9, Apr. 25, 1938; 1 cT, 2 9, Apr. 26, 1938; Baboquivari Mts., 1 9 , no
date (F. H. Snow) [UK]; Madera Canyon, Santa Rita Mts. (4400 ft.), 2 d,
2 9 , May 26, 1963 (J. G. Franclemont) [JGF]; 1 9 , June 2, 1963; 1 9 , June 4,
1963; Pinal Co., Oracle, 1 9, May 19, 1933 (Grace H. and John L. Sperry)
[AMNH]; 1 cf , June 4, 1935, [USNM]; [CUC]; 1 9 , [AMNH]; 3 9 , June 5, 1935
[AMNH]; Boyce Thompson Arboretum, Superior, 1 9 , Aug. 1, 1937 (A. B. Klots)
[AMNH]; Santa Curz Co., Madera Canyon, Santa Rita Mts., 1 cf, Aug. 15, 1949
(Lloyd M. Martin) [LCAM]; (5800 ft.), 1 9, June 22, 1955; Madera Canyon,
Santa Rita Mts., 1 &, Aug. 24^Sept. 2, 1957 (William A. Hammer) [LACM];
same locality (4880 ft.), 1 &, Apr, 14, 1963 (J. G. Franclemont) [JGF]; 1 cf ,
May 20, 1963; 1 &, May 21, 1963; 1 9, May 28, 1963; 1 9, June 1, 1963;
Santa Rita Mts., 4 9 , Aug. 22, 1946 (J. A. Comstock and Lloyd M. Martin)
[LACM]; 2 9 , Aug. 24, 1946; 1 9 , Aug. 27, 1946; 1 9 , Aug. 29, 1946; Nogales,
1 9 , May 30, 1899 [USNM]; Nogales, 2 d^, 3 9 , May 24-30 [USNM]; 1 d^, 2 9 ,
June 1-7; 1 &, July 8-15; Pena Blanca (3950 ft.), 1 9, June 7, 1963 (J. G.
Franclemont) [JGF]; Yavapai Co., 10 mi. east of Congress, 1 9, Aug. 22-23,
1927; Mayer, 3 d', 1 9, May 21, 1959 (M. O. Glenn) [MOG]; Prescott, 6 &,
May 6, 1950 (E. C. Johnston) [CNC]; no county, "Catal Spgs," 2 9 , (E. A.
Schwarz) [USNM]; Fish Creek, Tonto Nat. For., 5 d', 3 9 , May 9-10, 1918
(J. G. Bradley) [CU]; Todd's Lodge, Oak Creek Canyon, 1 d, June 12, 1941
(Grace H. and John L. Sperry) [CNC]; Santa Rita Mts., 1 d", 1 9 , May 21, 1898
(E. A. Schwarz) [USNM]; 1 9 , May 26, 1898; 1 d", June 8, 1898; 1 9 , June 13
1898; 1 9 , June 14, 1898; Santa Rita Mts. (5-8000 ft.), 1 d, June (F. H. Snow)
[UK]; no locality, 1 d, no date (Morrison), labeled "Coll. Ragonot 95-85"
[BM]; no locality, 1 d", 1881 (Morrison) [USNM]; no locality, 1 d, August (O.
Poling) [USNM]; 6 d, September (Poling) [CM]; 1 d, no date, from Fernald
collection [USNM]; 1 d^, no date, from C. V. Riley collection [USNM]; 1 d', 1 9 ,
no date [USNM].
California: Inyo Co., Bishop, 2 d", June 14, 1937 (E. C. Johnston) [CNC];
Lassen Co., Ravendale, 1 9, June 20, 1959 (G. I. Stage) [ABK]; Modoc Co.,
Canby, 3 d, July 16, 1936 (E. C. Johnston) [CNC]; Orange Co., Fullerton,
2 9, Nov, 14, 1962 (C. A. Toschi) [UCB]; Riverside Co., Rancho La Sierra,
Arlington, 1 d, Aug. 26, 1949 (A. H. Rindge) [AMNH]; 1 d, Aug. 23, 1952;
IdyUwild, 1 d, May 13, 1937 (H. Little) [AMNH]; Lake Hemet, 6 d, June 9,
1937 (E. C. Johnston) [CNC]; Riverside, 1 &, May 4, 1926 (Grace H. and John
L. Sperry) [CPK]; 1 d, Sept. 28, 1931 (C. H. Dammers) [USNM]; 1 d, 1931;
1 9, May 4, 1934 (Grace H. and John L. Sperry) [CPK]; 2 d, May 23, 1935
(Grace H, and John L. Sperry) [AMNH], [CNC]; 1 d^, 1 9 [AMNH]; 1 d",
July 12, 1935 (Grace H. and John L. Sperry) [AMNH]; 1 d, Apr. 29, 1936
[CNC]; 1 d, Apr. 27, 1937 (H. Buckwalter) [AMNH]; 3 d, June 1, 1939 [AMNH];
San Bernardino Co., Barton Flats, 2 d', June 27, 1946 [AMNH]; Upper Santa
Ana River, 1 9 , June 2, 1946 [AMNH]; 1 d, July 10, 1946; 1 d, July 18, 1946;
64 U.S. NATIONAL MUSEUM BULLETIN 280
1 cf, Aug. 18, 1948 (Melander) ; San Diego Co., Mt. Palomar, 3 &, July 18, 1963
(J. Powell) [UCB]; Mt. Palomar St. Pk., 1 &, July 12, 1953 (W. J. and J. W.
Gertsch) [AMNH]; San Diego, 1 &, June 16, 1912 (George H. Field) [ASNM];
Jidian, 2 9 , no date [USNM]; Siskiyou Co., Indian Butte, 1 cf, July 16, 1936
(E. C. Johnston) [CNC]; county unknown, Hathaway Creek, San Bernardino
Mts., 2 cf , Aug. 2, 1940 (J. A. Comstock and C. Henne) [LACM].
Idaho: Canyon Co., Parma, 1 cf, July 6, 1951 (A. J. Walz) [AMNH].
Nevada: Clark Co., Kyle Canyon, Charleston Mts., 1 ?, Apr. 26, 1950 (E.
C, Johnston) [CNC]; Wheeler's Spring, Charleston Mts., 2 cf. May 14, 1934
[CPK].
New Mexico: Bernalillo Co., Albuquerque, 1 9 , July 21, 1902 (Oslar) [USNM];
1 9 , July 24, 1902; 1 cf , 9 , no date [UK]; Hidalgo Co., Rodeo, 1 9 , Nov. 7,
1959 (J. Pv.. Powers) [UCB]; county unknown, "so. N. Mex.," 1 cT, 1 9, Aug.
23-30 (Poling), Rothschild Bequest B.M. 1939-1 [BM].
Texas: Blanco Co., 1 9 , no date, G. D. Hulst [AMNH]; Brewster Co., Alpine,
1 9, May 22, 1950 (E. C. Johnston) [CNC]; Burnet Co., 2 cf, 2 9 , no date
(F. G. Schaupp) [USNM]; Carmeron Co., Brownsville, 2 9, May 31, 1904
(H. S. Barber) [USNM]; 1 9, June 3, 1904; 1 d", June [UK]; 2 cf, June [ANS];
Hidalgo Co., Mercedes, 1 9, Aug. 31, 1958 (H. Smalzried) [AMNH]; 1 c?, no
date [CPK]; Jeff Davis Co., Limpia Canyon, 8 cf , 2 9 , May 20, 1950 (E. C.
Johnston) [CNC]; Randall Co., Palo Duro Canyon St. Pk., 3 cf", May 5, 1961
(Lloyd M. Martin, Robert H. Reid, William A. Rees, Robert J. Ford) [LACM];
3 cf, May 6, 1961; 3 <?, May 8, 1961; 1 9, May 9, 1961; 3 cf, 1 9, May 11,
1961; 1 9, May 12, 1961.
Utah: Utah Co., Vineyard, 2 d', July 7, 1917 (Tom Spalding) [ANS]; 1 cf ,
July 11, 1917; 1 d^, Aug. 6, 1917.
Washington: Adams Co., Othello, 1 cf, June 23, 1959 (R. F. Harwood)
[USNM]; 1 6", July 9, 1959; Chelan Co., First Creek, 2 cf, July 2, 1949 (E. C.
Johnston) [CNC]; Grant Co., Dry Falls, 1 &, June 30, 1949 (E. C. Johnston)
[CNC]; Quincy, 1 &, July 5, 1959 (C.S. Crawford) [USNM]; Okanogan Co.,
Black Canyon, 4 c?, July 1, 1949 (E. C. Johnston) [CNC]; Walla Walla Co.,
Walla Walla, 1 cf, June 27, 1935 (H. P. Lanchester) [USNM]; Whiteman Co.,
Snake River, opposite Clarkston, 1 cf. May 29, 1931 (J. F. Clarke) [USNM];
Yakima Co., Yakima, 1 cf, June 8, 1931 (Fred Dauy) [USNM]; 1 cf, Sept. 11,
1955 (A. I. Good) [AMNH].
CANADA: British Columbia: Kamloops, 1 9, June 28, 1937 (J. K. Jacob)
[CNC]; Oliver (1500 ft.), 1 &, June 5, 1953 (D. F. Hardwick) [CNC]; 7 cf,
June 16, 1953; 13 &, June 17, 1953; (2500 ft.), 3 cf, June 26, 1953; (1000 ft.),
1 cT, July 9, 1953 (J. E. H. Martin) [CNC]; (2500 ft.), 2 d^, July 10, 1953 (D. F.
Hardwick) [CNC]; Osoyoos, 1 d^. May 19, 1923 (C. B. Garrett) [CNC]; (1200 ft.),
1 9 , July 22, 1953 (D. F. Hardwick) [CNC].
Discussion. — The degree of black scaling on the forewings is
quite variable and ranges from a very marked broad tracing of the
veins to an almost total absence of black scales. The type specimen of
californiella is very sparsely set with black scales and that of insper-
gella rather heavily so, thus it is not surprising that they were
regarded as representing separate species. Wing venation, frons, and
genitalia are essentially identical in the two holotypes.
Small specunens may easily be mistaken for bipunctella.
REVISION OF PEORIINAE AND ANERASTIINAE 65
Coenochroa illibella (Hulst)
Figures 39, 40, 45, 67, 134, 169
Anerastia illihella Hulst, 1887, p. 138. — Rindge, 1955, p. 164.
Coenochroa puricostella Ragonot, 1887, p. 20.
Coenochroa illibella (Hulst). — Ragonot, 1889, p. 117; 1901, p. 419.— Barnes
and McDunnough, 1917, p. 150. — Hampson, 1918, p. 58. — McDunnough,
1939, p. 36.
Alamosa piperatella Hampson in Ragonot, 1901, pp. 369-370. — Barnes and
McDunnough, 1917, p. 149. — Hampson, 1918, p. 65. — McDunnough, 1939,
p. 36. [New synonomy.]
Diagnosis. — The species may be recognized by either the pale
costal region of the fo^e\^dng, the compressed and protruding central
beak of the frons, the absence of a discal spot, or the fact that K.2 is
almost always the first vein to leave the stalked K.2-f-4.
Description. — Frons with protuberance cyhndrical, ventraUy
about as long as wide, rim slanting at about 45°, teeth projecting
forward, central beak strongly compressed, protruding well beyond
rim, vestm-e hght browTi dorsaUy, white ventrally; labial palpi some-
what deflexed, basal segments white, second and third brown on
outer sides and dorsally, light brown to white ventrally; antennae as
in calif orniella; patagia, tegulae, vertex, occiput, and legs as in cali-
forniella.
Forewing radius averaging about 8.8 mm; area anterior to cell
white, sometimes variously sprinkled with black scales; region posterior
to radius quite variable, usually grayish orange sprinkled with
black scales predominantly on veins; area between veins sometimes
white, veins never traced with white; vein R2 first to branch from
R2+4. Hindwing with fringe white.
Genitalia as in calijorniella.
Types. — A. illihella, in the American Museimi of Natural History;
C. 'puricostella, in the Museum National d'Histoire Natiu-eUe; A.
'piperatella, in the British Museum (Natiu-al History).
Type data. — A, illihella, lectotype female, hereby designated,
Blanco Co., central Texas, November, collection G. D. Hulst, geni-
talia slide No. 3227, Carl Heinrich, June 14, 1946. In the original
description Hulst reports only that he had two males and one female
from Texas.
C. puricostella, lectotype male, hereby designated, Arizona, Morison,
genitaUa shde No. 538, J. Shaffer, June 20, 1965.
A. piperatella, lectotype male, hereby designated, labeled as follows:
"Syntype; Colorado Ameriq. sept.; CoU. Ragonot, 95 — 85.; Alamosa
piperatella Rag. Colorado ex Coll. Rag; cf genitaha shde 1-4-1967
J. Shaffer No. 706."
Specimens examined. — 104 cf , 56 9 .
66 U.S. NATIONAL MUSEUM BULLETIN 280
Distribution (Map 2). — Ontario west to Alberta and Washington,
south to California and Texas.
UNITED STATES: Arizona: County unknown, "Br't Angel," 1 &, no date
[USNM].
California: Inyo Co., 7 mi. north Parcher's Camp, 4 cf , June 30, 1961 (J.
Powell) [UCB]; Mono Co., Leevining, 2 cf, July 19, 1938 (E. C. Johnston)
[CNC]; 1 &, July 20, 1938; 1 mi. southwest Tom's Place, 1 cf , 2 ? , Aug. 10, 1963
(C. A. Toschi) [UCB]; 1 cf , Aug. 13, 1963 (M. J. Tauber and C. A. Toschi).
Colorado: Adams Co., Watkins, 8 cf, July 4-5, 1927 [CU]; Boulder Co.,
Boulder, 1 cf, June- August, 1896 [BMNH]; Chaffee Co., Salida, 1 cf, July 9, 1937
[AMNH]; Denver Co., Denver, 1 cf, no date [USNM]; 2 &, (Oslar) ; Prowers
Co., Lamar, 2 cT, Sept. 24, 1945 (E. C. Johnston) [CNC]; county unknown, 1 cf,
no date (Oslar) [USNM]; 1 cf , no date [AMNH]; 6 &, no date [USNM].
Kansas: Clark Co. (1962 ft.), 1 ?, June (F. H. Snow) [ANS]; 1 cf [UK];
2 $ [USNM].
New Mexico: Eddy Co., Artesia, 2 cf , May 13, 1950 (E. C. Johnston) [CNC]
Sandoyal Co., Frijoles Canyon, Bandelier Nat. Mon., 1 cf, July 9-11, 1957;
(A. B. Klots) [AMNH]; (6050 ft.), 4 &, July 17, 1962 (E. and I. Munroe) [CNC];
Otero Co., Mescalero, 1 cT, May 12, 1950 (E. C. Johnston) [CNC]; Dona Ana Co.,
Mesilla, 1 $ , July 30 (Cockerell) [BM]; Mesilla Park, 1 c?, 1 ? , May 8 (Cockerell)
[USNM]; 3 c?, 3 ? , May 13; 2 d", (3800 ft.) [CNC]; 1 d", May 22 (Cockerell)
[CNC]; 1 d [USNM]; 1 9 , June 8; 1 ? , July 8; 1 9 , Aug. 29; 1 ? , no date;
1 ? [CNC]; Mesquite near Mesille Park, 1 9, July 12, 1917 [CNC]; 5 d [CU];
Grant Co., Silver City, 1 cf. May 25, 1913 (J. B. Wallis) [CNC]; 1 9, May 27
1913; 1 9 , July 25, 1913; 4 cf , 3 9 , July 26, 1913; 2 cf , July 27, 1913; 7 cT, 1 9 ,
July 28, 1913; county unknown, southern New Mexico, 2 9 , Aug. 23-30 (Poling)
[BM].
North Dakota: Ransom Co., 1 mi. southeast McLeod, 1 cf , July 19, 1963
(J. R. Powers) [UCB].
South Dakota: Davison Co., Mitchell, 1 cf, Sept. 14, 1945 (E. C. Johnston)
[CNC].
Texas: Reeves Co., Pecos, 2 d', 17 9, May 18, 1950 (E. C. Johnston) [CNC];
1 cf , 8 9 , June 2, 1950; county unknown, 1 9 , no date, labeled, "Fernald Col-
lection," "U.S.N.M. Type No. 40077," "Petaluma ilUbella Hulst, Type,"
"Aneraslia illibella Hulst, Type" [USNM].
Utah: Garfield Co., 11 mi. southeast Panguitch (7200 ft.), 4 d, July 11, 1960
(F., P., and B. Rindge) [AMNH]; 3 &, July 12, 1960; 5 d, July 13, 1960; Tooele
Co., Stockton, 1 9 , Aug. 8, 1904 (Tom Spalding) [USNM]; 1 9 , no date; Utah
Co., Vineyard, 1 d, Aug. 6, 1917 (Tom Spalding) [ANS]; 1 9, July 16, 1917.
Washington: Grant Co., Quincy, 1 d, July 1, 1959 (C. S. Crawford) [USNM];
1 c?, July 5, 1959.
CANADA: Alberta: Medicine Hat, 1 d, July 15, 1956 (E. E. Sterns) [CNC];
Scandia, 1 cf , July 6, 1956 (E. E. Sterns) [CNC].
Ontario: Marmora, 1 9, Aug. 16, 1952 (J. F. McAlpine) [CNC]; Point Pelee
1 d,l 9 , June 29, 1927 (F. P. Ide) [CNC].
Discussion. — The original description of illibella lists two males
and one female from Texas. The disposition of the male types is not
known, and two female specimens from Texas bear Hulst's "Type"
label. Available evidence is insuflScient to determine which, if either,
of these two females was included in the original series, and I have
REVISION OF PEORIINAE AND ANERASTIESTAE 67
designated as lectotype the specimen in the American Museum of
Natural History. The remaining female is deposited in the U.S.
National Museum, full data being given above.
The specimens from Pecos, Tex. are atypical in that the frons beak
is unusually well developed and the rim is flared anteriorly so that
the teeth point somewhat anterolaterally. Most of the Pecos speci-
mens are only sparsely set with black scales on the forewings, but
otherwise are typical of the species.
Coenochroa bipunctella (Barnes and McDnnnoiigh), new combination
Figures 42, 47, 66, 135, 170
Alamosa bipunctella Barnes and McDunnough, 1913, p. 184; 1917, p. 149. —
Grossbeck, 1917, p. 134. — Hampson, 1918, p. 65. — McDunnough, 1939,
p. 36. — Kimball, 1965, p. 251.
Diagnosis. — This is the only member of the genus having numer-
ous small cornuti on the vesica of the male rather than a single one.
The ductus seminalis of females is thickened where it leaves the
ductus bursa, not slender as in other species. Otherwise, specimens
could be confused with small examples of calijorniella, which has
similar wing venation and maculation, but the two species are largely
or wholly allopatric.
Description. — Frons wdth protuberance cylindrical, ventrally
about as long as wide, rim slanting at about 45°, teeth projecting
forward, central beak conical Avith irregular tip even with end of rim,
vesture light brown to browTi dorsally, white to light brown ventrally;
labial palpi usually porrect, basal segments white, second and third
brown dorsally and on outer sides, light brown ventrally; antennae as
in californiella; patagia and tegulae brown, vertex and occiput light
brown dorsally, occiput brown dorsolaterally, white below middle of
eye; legs brown on outer sides, white on inner sides.
Forewing radius averaging about 6.9 mm; cell, discocellular, radials,
medials, cubitals, and anal veins each traced with a broad white band
sprinkled vnih. black scales; area anterior to cell varying yellow to
brown and sprinkled \^dth black scales; ground color yellow to yellow
brown; prominent black discal spot at lower outer angle of cell; brown
on underside; vein R4 first to branch from stalked E2+4. Hindwing
bordered with double brown line; fringe white; ground color light
broAvn, darker in apical area; Cui and M3 stalked for at least one-half
length, sometimes fused.
Male genitalia mth vinculum V-shaped, pointed anteriorly.
Aedeagus club-shaped vesica bearing numerous small cornuti which
diminish in size anteriorly.
Female genitalia with ductus bursae rather short. Ductus seminalis
thickened basally.
68 U.S. NATIONAL MUSEUM BULLETIN 280
Type. — In the U.S. National Museum.
Type data. — One female, Fort Myers, Fla., Apr. 16-23, Barnes
collection; USNM 69383; genitalia slide No. 573, J. Shaffer, Nov. 23,
1965.
Specimens examined. — 63 cf , 3 9 .
Distribution (Map 2). — Atlantic and Gidf Coastal Plain, recorded
from Mississippi to New Jersey.
UNITED STATES: Florida: Alachua Co., GainesvUle, 1 9, June 2, 1927
(J. Speed Rogers) [CU]; 1 cf, June 29, 1927; 1 d", July 7, 1927; 8 cf , July 8, 1927;
1 cf , 1 9 , July 10, 1927; 1 & [CNC]; 5 d", July 1927 (J. Speed Rogers) [CU] 1 d,
Apr. 20, 1952 (O. Peck) [CNC]; 1 &, Apr. 22, 1952; Dade Co., South Campus,
Richmond, 2 cf, Nov. 27, 1951 (F. G. Butcher) [CNC]; Indian River Co., Vero
Beach, 1 cf , no date (J. R. Malloch) [USNM]; Orange Co., Orlando, 1 cf , June
14-18, 1927 (C. C. McBride) [CU]; Pasco Co., Elfers, 1 9, Apr. 17, 1952 (G. S.
Walley) [CNC]; 6 d, Apr. 18, 1952 (J. R. Vockeroth) [CNC]; Polk Co., Lake
Alfred, 1 cf, June 30, 1928 (L. J. Bottimer) [USNM]; Sarasota Co., 1 cf , May 5,
1946, (C. P. Kimball) [CPK]; Siesta Key, 1 d, Feb. 23, 1951 (C. P. Kimball)
[CNC]; Ic?, Feb. 27, 1951; 1 cf, Dec. 24, 1941; 2 cT, Feb. 13, 1952; 1 cf, Feb. 23,
1952; 3 cf , Mar. 1, 1952; 1 &, Mar. 26, 1952; 2 cf , Apr. 2, 1952; 1 d, Nov. 6, 1952;
1 &, Nov. 15, 1952; 1 cT, Apr. 11, 1953; 1 d, Apr. 24, 1953; 1 d, Feb. 13, 1954;
1 &, Mar. 14, 1954; 1 &, Feb. 9, 1955; 1 d, Feb. 18, 1955; 1 d, Apr. 29, 1956;
1 &, Oct. 24, 1956; 1 9 , Jan. 23, 1957; Seminole Co., Altamont, 1 cf , Sept. 21,
1924 (F. R. Cole) [USNM].
Mississippi: Forrest Co., Camp Shelby, 1 cf , Sept. 1-15, 1944 (C. D. Michener)
[AMNH].
New Jersey: Ocean Co., Lakehurst, 1 cf, July 24, 1939 (E. P. Darlington)
[ANS].
North Carolina: Robeson Co., Maxton, 1 d, May 11, 1944 (A. B. Klots)
[AMNH]; 2 cf , May 23, 1944; 1 d, Oct. 4, 1944.
Discussion. — The species has been poorly collected outside of
Florida, as is generally true for Coastal Plain groups, and is probably
more common there than the few scattered records indicate. It may
be sympatric with calijorniella in the Coastal Plain section of Texas.
Bandera Ragonot
Figure 68
Bandera Ragonot, 1887, p. 19. — Hulst, 1890, p. 202 [listed in Phycitinae].—
Smith, 1891, p. 84 [listed in Phycitinae].— Ragonot, 1889, p. 117; 1901, p. 409.
— Hulst, 1902, p. 440. — Barnes and McDunnough, 1917, p. 150. — Hampson
1918, p. 89. — McDunnough, 1939, p. 36. — Heinrich, 1956, pp. 1, 315, 316.
[Type: Anerastia binotella Zeller, 1872. Original designation.]
Nasutes Hampson, 1930, p. 53 [described in Phycitinae]. — McDunnough, 1939,
p. 28. — Heinrich, 1956, pp. 315-316 [listed as synonym of Bandera]. [Type:
Nasutes venata Hampson, 1930, Monobasic]
Diagnosis. — The absence of ocelli and the presence of a tongue,
reduced but exposed between the palpi, serve to distinguish this
genus from the other North American Phycitinae.
REVISION OF PEORIINAE AND ANERASTIINAE 69
Description. — Frons conical; labial palpi porrect, outer sides of
second and third segments clothed with white-tipped brown scales,
basal segments clothed with broad white scales; maxillary palpi
cylindrical, almost reaching frons; tongue exposed betw^een palpi;
antennae filiform, compressed, ciliate ventrally; ocelli absent; legs
brown to fuscous laterally, tarsi brown and speckled with white.
Forewings with a brown-speckled white band anterior to cell;
veins traced wdth white and margined on both sides with a sprinkling
of brown scales; 9 veins; Ri from well before outer angle of cell; II2
from cell; R44.5 fused; Mj from upper outer angle; M2+3 fused and from
the angle; Cui from before the angle; Cu2 from well before the lower
outer angle of cell. Hindwing light brown above and below, slightly
darker toward apex; 7 veins; Sc and Rs long stalked; Cui stalked with
fused M24.3; Cu2 from well before the lower outer angle of cell.
Male genitalia \vith uncus broadly triangulate, slightly tapering,
\vith apex broadly rounded, laterally and dorsocaudally setose.
Gnathos arms terminating in an anterior ventral hook; apical process
a simple posteriorly directed hook. Juxta U-shaped, wdth small
lateral knobs. Vincidum stout. Valvae distally rounded; costa tubular,
slightly tapering distally. Vesica unarmed.
Female genitalia \\'ith ovipositor tapering distally, about as long as
basal width. Apophyses slender, well sclerotized; posterior about
five or six times as long as their basal separation; anterior about two
to four times as long as their basal separation. Ductus bursae long,
weakly sclerotized. Bursa small; \vith a group of three or four small
barlike signa near posterior end. Ductus seminalis threadlike; from
near or at posterior end of bursa near signa.
Discussion. — The genus shows affinities to a number of other
phycitine genera and is probably closest to Anagasta Heinrich. The
great preponderance of males in collections suggests that females
are far less readily attracted to light.
Key to the Species of Bandera Based Upon the Maculation
1. Space between vein A2 and cell of forewing a pale yellow field sprinkled with
brown or with browTi scales on fold of Ai virginella
Space between vein A2 and cell of forewing a solid yellow field, not sprinkled
with brown ^
2. Basal area of costa yellow cupidinella
Entire length of costa white, sprinkled with brown scales .... binotella
Bandera binotella (Zeller)
Figures 31, 68, 99, 136, 171
Anerastia binotella Zeller, 1872, pp. 554-555.
Bandera subluteella Ragonot, 1887, p. 19; 1889, p. 117. — Smith, 1891, p. 84
[listed in Phycitinae]. — Ragonot, 1901, pp. 410-411. — Hulst, 1902, p.
70 U.S. NATIONAL MUSEUM BULLETIN 2 80
440. — Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 90. —
McDunnough, 1939, p. 36. [New synonymy.]
Bandera binotella (Zeller). — Ragonot, 1889, p. 117. — Hulst, 1890, p. 202.—
Smith, 1891, p. 84. — Ragonot, 1901, pp. 409-410. — Hulst, 1902, p. 440. —
Dyar, 1908c, p. 116. — Barnes and McDunnough, 1917, p. 150. — Hampson,
1918, pp. 89-90. — McDunnough, 1939, p. 36.
Diagnosis. — The presence of a transverse posterior line of dots and
a transverse anterior dot on vein A2 of the forewing, a truncate
vinculum, and an incomplete transtilla each by itself distinguish this
species from other members of the genus.
Description. — Frons about as long as diameter of eye, light brown;
vertex, occiput, tegula, and prothorax white to light brown dorsally,
gray brown laterally; abdomen light creamy brown dorsally, brown
ventraUy.
ForeAving yellow between veins; discal cell, area between cell and
vein A2, and area posterior to A2 solid yellow; transverse posterior
variously developed as a row of dots on veins, best represented by a
dark spot near end of vein A2; a dark spot on vein A2 halfway between
tp line and wing base; light brown to brown on underside.
Male genitalia Avith transtilla incomplete. Lateral knobs of juxta
well developed, setose. Vinculum rectangular, broadly truncate ante-
riorly. Aedeagus tapering posteriorly, subtruncate anteriorly. Eighth
abdominal segment of male bearing lateral hair tufts.
Female genitalia with ovipositor sparsely setose. Ductus bursae
broad.
Types. — A. binotella, in the British Museum (Natural History),
(from Zeller collection); B. suhluteella, in the Museum National
d'Histoire Naturelle.
Type data. — A. binotella, in the original description Zeller reports
"Vaterland: Texas. Belfrage fing das Exemplar am 12. Juli." Type
examined by Mr. Paul Whalley.
B. subluteella, lectotype male, hereby designated, labeled as follows:
"TYPE; WALSM; 1901 coll. E. L. Ragonot Museum Paris; Bandera
subluteella Rag. type orig. pi. XL f. 21; cf genitalia slide July-28-
1965 J. Shaffer No. 553." The specimen also bears a small handwritten
label inscribed AAdth "Col9," probably intended to be "Colo" (Colo-
rado). Ragonot may have interpreted the inscription as an abbrevia-
tion for California since that state is given as the type locality. I
have seen no specimens of binotella from California although the species
is found in Washington and therefore might be expected to occur in
northern California.
Specimens examined. — 71 cf, 8 9.
Distribution (Map 4). — Washington southeastward to Arizona
and Texas; eastern population on Atlantic Coastal Plain, New Jersey
to Massachusetts.
REVISION OF PEORIINAE AND ANERASTIINAE 71
UNITED STATES: Arizona: Apache Co., Greer, White Mts. (8500 ft.),
1 cT, Aug. 6, 1962 (E. and I, Munroe) [CNC].
Colorado: Rocky Mountain National Park, 9 cf , Aug. 15, 1937 (A. B. lOots)
[AMNH].
Connecticut: New Haven Co., East River, 1 cf , July 10, 1909 (Charles R.
Ely) [USNM]; 1 d^, July 30, 1910; 1 c?, Aug. 13, 1910; 1 d^, Aug. 29, 1910;
1 d^, August 1910; 1 9, July 1911; Windham Co., Putnam, 1 d', Aug. 9,
1940 (A. B. Klots) [AMNH]; 1 ? , August 1942; 1 d^, Aug. 16, 1960 [ABK].
Massachusetts: Barnstable Co., Barnstable, 1 9, July 4, 1949 (C. P. Kim-
ball) [CU]; same locality, 1 d", Sept. 11, 1949 (C. P. Kimball) [CPK]; 2 d^,
Aug. 5, 1950; 1 d", Sept. 12, 1952; Dukes Co., Martha's Vineyard, 1 9, July 20
(F. M. Jones) [CPK]; Worcester Co., Worcester, 1 d', July 11, 1933 [CU].
Montana: Roosevelt Co., Poplar, 1 d", July 15, 1921 (H. G. Dyar) [USNM].
New Jersey: Bergen Co., Oakland, 1 9 , July 26, 1947 (C. P. Kimball) [CNC];
3 d^, Aug. 4, 1948; 3 d^, Aug. 6, 1948; 1 9 , Aug. 8, 1948 [CU]; 1 &, Aug. 8, 1948
[CNC]; 3 &, Aug. 9, 1948 [CNC, CPK]; 2 d', Aug. 10, 1948 [CPK]; 3 d^, Aug. 13,
1948 [CNC, CPK]; 2 d^, Aug. 14, 1948 [CPK]; 1 d^, Aug. 22, 1948 [CPK]; 1 d", 1
9 , Aug. 24, 1948 [CNC]; 1 d", August 1948; "oM Beach," 1 d, "7.2" (F. Haim-
bach) [USNM].
New Mexico: Colfax Co., Cimarron Canyon, Sangre de Cristo Mts. (7900 ft.),
1 d, July 7, 1962 (E, and I. Munroe) [CNC]; 1 d, July 13, 1962; Lincoln Co.,
Cedar Creek Camp, 2 mi. north of Ruidoso (7000 ft.), 1 d, June 30, 1961 (F., P.,
and J. Rindge) [AMNH]; 6 d, July 29, 1962 (E. and I. Munroe) [CNC]; Mc-
Kinley Co., McGaffey, Zuni Mts. (7500 ft.), 2 d, July 20, 1962 (E. and I. Munroe)
[CNC]; 4 d, July 21, 1962; 2 &, July 22, 1962; 2 d, July 24, 1962; Sandoyal Co.,
Frijoles Canyon, Bandelier Nat. Mon. (6000 ft.), 1 cf , July 17, 1962 (E. and I.
Munroe) [CNC].
Texas: HUdalgo Co., Mercedes, 1 d, Aug. 31, 1958 (H. Smalzried) [AMNH],
Washington: Chelan Co., Leavenworth, 2 d", July 3, 1949 (E. C. Johnston)
[CNC]; Whitman Co., Pullman, 1 d, Aug. 6, 1898 (C. V. Piper) [USNM], from
the paratype series of Bandera virginella, No. 11851 [USNM], genitalia slide, Carl
Heinrich No. 6, Dec. 20, 1932.
CANADA: British Columbia: Oliver, 1 9, Aug. 2, 1953 (D. F. Hardwick)
[CNC]; 1 d, Aug. 10, 1953; 1 d, Aug. 11, 1953; 1 d, Sept. 14, 1953.
Alberta: Dominion Range Sta., Manyberries, 1 d, Aug. 14, 1951 (D. F, Hard-
wick) [CNC].
Discussion. — Tlie presumed habitat differences and the wide geo-
graphic separation between eastern and western populations indicate
that they may well represent distinct biological species, but the
absence of clear-cut morphological differences between them dictates
that better criteria must be found if a new species name is to be given
to the eastern population. Eastern specimens differ in having the frons,
vertex, occiput, tegula, and prothorax covered with brown scales.
The forewing ground color is reddish brown rather than yellow, and
the transverse posterior line of dots is usually better developed.
Bandera cupidinella Hulst
Figures 32, 137, 172
Bandera cupidinella Hulst, 1888, p. 118. — Ragonot, 1889, p. 117. — Hulet, 1890,
p. 203. — Smith, 1891, p. 84. — Ragonot, 1901, p. 410. — Hulst, 1902
72 U.S. NATIONAL MUSEUM BULLETIN 2 80
p. 440. — Dyar, 1908c, p. 116. — Barnes and McDunnough, 1917, p. 150.
— Hampson, 1918, p. 90. — McDunnough, 1939, p. 36. — Rindge, 1955,
p. 160. — Heinrich, 1956, pp. 315-316.
Anerastia conspersella Ragonot, 1901, p. 404. — Barnes and McDunnough, 1917,
p. 149. — McDunnough, 1939, p. 36.
Rhinaphe conspersella (Ragonot). — Hampson, 1918, p. 86.
Nasutes venata Hampson, 1930, p. 54. — McDunnough, 1939, p. 28 [listed in
Phycitinae]. — Heinrich, 1956, pp. 315-316 [Hsted as a synonym of
cupidinelld\.
Diagnosis. — This species may be recognized either by the absence
of hair brushes in the male, or by the presence of a basal yellow margin
on the costa of forewing of both sexes.
Description. — Frons about as long as diameter of eye, light brown;
lateral scales of labial palpi with brown restricted to a subterminal
band; vertex, occiput, and thorax light yellow brown dorsally, occiput
and prothorax gray brown laterally; abdomen varying light brown to
brown.
Forewing with costa abruptly yellow marginally at base to one-fifth
distance to apex; yellow to orange yellow between veins; discal cell,
area between cell and vein A2, and area posterior to vein A2 solid
yellow with no brown scales; fringe on outer margin of alternating
white and brown bands; brown beneath.
Male genitalia with transtilla complete, weakly sclerotized medially.
Lateral knobs of juxta poorly developed. Vinculum bluntly rounded
anteriorly. Aedeagus tapering very slightly posteriorly, bluntly
rounded anteriorly. Eighth abdominal segment without lateral hair
tufts.
Female genitalia with ovipositor setose. Ductus bursae broad.
Types. — B. cupidinella, in the U.S. National Museum; N. venata
and A. conspersella, in the British Museum (Natural History).
Type data. — B. cupidinella, lectotype male, hereby designated,
Colorado, Denver (Fernald collection), USNM 40074, genitalia slide
No. 575, J. Shaffer, Nov. 23, 1964.
N. venata, in the original description Hampson reports: Colorado,
Boulder, 1 cf ; Cockerell; alar expanse 20 mm. Type examined by
Mr. Paul Whalley.
A. conspersella, lectotype male, hereby designated, Colorado, Akron,
"11/9/83"; genitalia slide No. 705, J. Shaffer, Jan. 4, 1967.
Specimens examined. — 36 cf, 2 9.
Distribution (Map 4).
UNITED STATES: Colorado: Boulder Co., Boulder, 2 cf , September 5 (Cocke-
rell) [USNM]; Denver Co., Denver, 2 cT, Sept. 1, 1904; 1 d', Sept. 3, 1904; 9 d^,
no date; Denver, 4 cf , no date [USNM]; Prowers Co., Lamar, 4 cf, 1 9 , Sept. 24,
1945 (B. C. Johnston) [CNC]; Pueblo Co., Pueblo, 1 cf , September 1899 [AMNH];
no locaUty, 11 d" [USNM].
REVISION OF PEORIINAE AND ANERASTIESTAE 73
New Mexico: Hot Springs (7000 ft.), 1 cf, September [USNM]; no locality
(7000 ft.), 1 9 , 1927 (H. S. Parish) [CU].
Texas: Jeff Davis Co., Ft. Davis (oOOO ft.), 1 d^, 1927 (IT. S. Parish) [CU].
Discussion. — Two males in the American Museum of Natural
History are labeled "Type" from Hot Springs, N. Mex., 7000 ft.,
August, September. A male genitalia slide of the August specimen
was prepared by Carl Heinrich, No. 3226, June 14, 1946. The type
locality is given as Colorado and these specimens must be regarded
as another example of Hulst pseud oty pes. The specimen in the U.S.
National Museum labeled "Type" is from Colorado and is assumed to
be genuine.
Bandera virginella Dyar
FiGXJBES 33, 138, 173
Bandera virginella Dyar, 1908c, pp. 116-117. — Barnes and McDunnough, 1917,
p. 150. — McDunnough, 1939, p. 36.
Diagnosis. — In the other members of this genus the discal cell
and the area between the cell and vein A2 are scaled with solid yellow,
but in virginella these areas are pale yeUow sprinkled with brown
scales. The male genitalia are identical with those of cupidinella, but
the eighth abdominal segment bears lateral hair tufts.
Description. — Frons about two-thirds as long as diameter of eye,
dirty white dorsally and light brown ventrally; tongue somewhat less
well developed than on other members of the genus; vertex, occiput,
and thorax light yellow brown dorsally, occiput and prothorax gray
brown laterally; abdomen varying light brown to brown.
Forewings with areas between veins pale yellowish white, variously
sprinkled with brown scales; fold of Ai traced with brown scales;
brown beneath.
Male genitalia as in B. cupidinella. Eighth abdominal segment with
hair tufts similar to those of B. binotella, but with fewer scales and
therefore smaller.
Female genitalia with ovipositor sparsely setose. Ductus bursae
slender.
Type. — In the U.S. National Museum.
Type data. — Holotype: Washington, Pullman, male, July 24, 1898;
C. V. Piper; Washington Exp. Sta. No. 463; USNM 11851; male
genitaha slide No. 5, Carl Heinrich, Dec. 20, 1932.
Paratypes: Five specimens, Washington, Pullman, males; C. V.
Piper; USNM 11851. Dates are July 8, 1898, Aug. 6, 1898, Aug. 10,
1898, July 31, 1899, Aug. 3, 1899. The specimen collected Aug. 6, 1898
bears genitalia slide label C.H. No. 6, Dec. 20, 1932 and belongs to
B. binotella.
Other specimens examined. — 51 cf , 7 9 .
74 U.S. NATIONAL MUSEUM BULLETIN 280
Distribution (Map 4). — New Mexico to California, north to
Alberta and British Columbia.
UNITED STATES: Akizona: Cochise Co., Southwestern Res. Sta., Chiri-
cahua Mts., 1 d', Apr. 11, 1962 (Carl W. Kirkwood) [CNC]; 1 cf, Apr. IS, 1962;
Yavapai Co., Prescott, 5 cf , May 6, 1950 (E. C. Johnston) [CNC].
California: Riverside Co., Kenworthy, 6 cf, June 9, 1937 (E. C. Johnston)
[CNC]; Lake Hemet, 1 cf , June 9, 1937 (E. C. Johnston) [CNC]; San Bernardino
Co., 12 mi. southeast of Ivanpan, 1 cf, May 1, 1956 (J. Powell) [UCB]; Upper
Santa Ana River. 1 d", Aug. 12, 1948 (Grace H. and John L. Sperry) [AMNH];
Siskiyou Co., Indian Butte, 4 cf , July 16, 1936 (E. C. Johnston) [CNC]; Ventura
Co., Camp Ozena, Upper Cuyama, 4 d, June 13, 1963 (C. W. Kirkwood) [CPK];
3 cf , June 14, 1963; 1 cf, June 18, 1963, 1 cf , June 19, 1963; 2 cf , June 24, 1963;
1 cf , June 27, 1963.
Colorado: Rock Creek Canyon, 1 cf, Sept, 26, 1957 (Margot May) [CNC];
1 cf , Sept.28, 1957.
New Mexico: Eddy Co., White City, 1 cf , May 15, 1950 (E. C. Johnston)
[CNC].
Washington: Walla Walla Co., Walla Walla, 1 9, Apr. 9, 1931 (D. R. Bran-
non) [USNM]; 1 cf, Apr. 21, 1931; 3 cf, Apr. 18, 1935 (H. P. Lanchaster) ;
Whitman Co., Pullman, 1 cf, June 19, 1930 (J. F. Clarke) [USNM]; 1 cf , 1 9 ,
Aug. 3, 1932; 2 cf, July 3, 1935; Yakima Co., Satus Creek, 2 cf, Aug. 19, 1949
(E. C. Johnston) [CNC]; 1 cf, Sept. 16, 1949; Tieton, 1 cf, May 12, 1931 (Fred
Deari) [USNM].
CANADA: Alberta: Medicine Hat, 1 cf , June 24, 1945 (K. Bowman) [CNC].
British Columbia: Shingle Cr. Road, Keremeos, 1 cf , June 8, 1935 (A. N.
Gartrell) [CNC]; 3 9 , July 15, 1935; 2 9 , July 22, 1935; Ohver (1000 ft.), 1 cf ,
Aug. 10, 1953 (D. F. Hardwick) [CNC]; 1 cf, Aug. 11, 1953.
No data given except S. Diego, 1 cf , July 19, 1924 [USNM].
Wakulla, new genus
Type. — Bandera carneeUa Barnes and McDunnough, 1913.
Diagnosis. — Among the phycitine genera exhibiting a reduced
tongue this is the only genus in which the males have both hair tufts
on the eighth abdominal segment and a spatulate uncus.
Description. — Frons conical, about two-thirds as long as eye
diameter; labial palpi slightly ascending and extending somewhat
beyond frons in male, female ascending, reaching beyond vertex;
maxillary palpi well developed, cylindrical; tongue reduced, visible
between bases of palpi, scaled at base; antennae filiform in both sexes,
compressed, ciliate ventrally, cilia short, about one-sixth as long as
segment width; ocelli minute.
Forewing and hindwing venation as in Bandera.
Male genitalia with uncus spatulate, setose dorsally and laterally.
Gnathos arms each terminating in anterior hook; apical process bi-
furcate. Transtilla incomplete. Juxta U-shaped, surrounding aedeagus
ventrally and laterally. Vinculum stout, tapering, broadly truncate
anteriorly, length and width about equal. Valvae with costa tubular,
REVISION OF PEORIINAE AND ANERASTIINAE 75
terminating in short free spine just before apex of cuculliis; sacculus
inflated on basal one-fourth of valve. Aedeagus stout, slightly tapering
distally, vesica unarmed. Eighth abdominal segment of male with
hair tufts.
Female genitaUa with ovipositor tapering distally, sparsely setose,
about 13^ times as long as broad. Apophyses well sclerotized, posterior
about four to five times as long as basal separation, anterior slightly
less than twice as long as basal separation. Ductus bursae long. Bursa
small, bearing three small barlike signae. Ductus seminalis leaving
bursa near posterior end, just posterior to signae, threadlike, well
sclerotized.
Wakulla carneella (Barnes and McDunnough), new combination
Figures 43, 100, 139, 174
Bandera carneella Barnes and McDunnough, 1913, p. 184; 1917, p. 150. — Gross-
beck, 1917, p. 134. — McDunnough, 1939, p. 36. — Kimball, 1965, p. 251.
Diagnosis. — This is the only known species in the genus.
Description. — Labial palpi with basal segments white, second and
third segments Hght brown or reddish brown on outer sides, white or
nearly so on inner sides; maxillary palpi reddish brown; antennae with
scape brown, shaft light brown; frons, vertex, occiput, patagia, and
tegulae reddish brown.
Forewing reddish brown, sprinkled with dark brown scales; two
small dark brown spots at distal end of cell; male with tuft of black
scales at base of costa. Hindwing light brown.
GenitaUa as described for the genus.
Type. — In the U.S. National Museum.
Type data. — Holotype male, Everglades Fla., Apr. 8-15, Barnes
collection, "Photograph pi. 1 No. 5"; USNM 69384; genitalia slide
No. 574, J. Shaffer, Nov. 23, 1965.
Specimens examined. — 3cf, 1 9.
Distribution (Map 3). — Known only from Florida.
UNITED STATES: Florida; Manatee Co., Gulf Coast Exp. Sta., Bradenton,
Id', Aug. 5, 1955 (E. G. Kelsheimer) [CPK]; Id', Mar. 28, 1956; Sarasota Co.,
Siesta Key, Icf, Feb. 13, 1951 (C. P. Kimball) [CPK]; 1?, Nov. 9, 1952.
Tampa Ragortot
Tampa Ragonot, 1887, pp. 19-20; 1889, p. 117. — Hulst, 1890, p. 203 [listed in
Phycitinae]. — Smith, 1891, p. 84 [listed in Phycitinae]. — Ragonot, 1901,
p. 411. — Hulst, 1902, p. 440. — Barnes and McDunnough, 1917, p. 150. —
Forbes, 1923, p. 637. — Hampson, 1918, p. 62. — McDunnough, 1939, p. 36.
[Type: Tampa dimediatella, 1887. Monobasic]
285-934—68-
76 U.S. NATIONAL MUSEUM BULLETIN 2 80
Diagnosis. — The presence of only two radial veins in the forewing
distinguishes Tampa among the phycitine genera showing a reduced
tongue.
Description. — Frons conical; labial palpi ascending, often porrect
in females, about three times eye diameter in males, 3}^ times in
females; maxillary palpi three-segmented, cylindrical, reaching frons
or nearly so; tongue reduced, usually visible between palpi; antennae
filiform in both sexes, somewhat thicker in male, female scaled dor-
sally and laterally, ventrally with short appressed cilia, male scaled
dorsally, ventrally wdth cilia about one-half segment wddth, hooked
at tips; ocelli absent.
Forewing with 8 veins; Ri from near upper outer angle of cell,
R2+4 fused, from the angle; Mi from about one-third from top of cell;
M2+3 fused, from lower outer angle; Cui from just before the angle;
Cu2 from w^ell before the angle. Hindwing with 7 veins; Sc and Rs
long stalked: Mi from upper outer angle; M2+3 fused, long stalked
with Cui, from lower outer angle; Cu2 from just before the angle.
Male genitalia with uncus tapering, broadly and bluntly rounded
apically; rather densely setose dorsally and laterally. Gnathos wdth
medial process small, simple, shallowly notched apically. Transtilla
complete, w^eakly sclerotized. Juxta crescent shaped, with pair of
lateral setose tubercles. Vinculum V-shaped, apex bluntly rounded.
Valve narrow, sides nearly parallel, apex rounded. Vesica unarmed.
Female genitalia with ovipositor tapering distally, about twice as
long as wide. Apophyses slender, posterior about six or seven times
as long as their basal separation; anterior about 2}^ times as long as
their basal separation, diverging. Ductus bursae long, slender. Bursa
small, with a group of five to seven parallel barlike signae extending
anteriorly from ductus seminalis near posterior end. Ductus seminalis
from posterior end of bursa, tapering, then slender.
Tampa dimediatella Ragonot
Figures 35, 69, 101, 140, 175
Tampa dimediatella Ragonot, 1887, p. 20; 1889, p. 117. — Hulst, 1890, p. 203. —
Smith, 1891, p. 84. — Ragonot, 1901, p. 411. — Hulst, 1902, p. 440. —
Barnes and McDunnough, 1917, p. 150. — Grossbeck, 1917, p. 134. —
Hampson, 1918, p. 63. — Forbes, 1923, p. 367. — McDunnough, 1939,
p. 36. — Kimball, 1965, p. 251.
Diagnosis. — This is the only known species in the genus.
Description. — Labial palpi with basal segments and ventral and
inner sides of second segments white, third segments and outer sides
of second dark brown to black; maxillary palpi, frons, vertex, sides
of occiput dark brown to black; dorsum of occiput, patagia, and
REVISION OF PEORIINAE AND ANERASTIENAE 77
tegulae somewhat lighter; antennae brown; legs white on inner sides,
outer sides varying light brown to dark brown.
Fore wings light brown anterior to cell, sprinkled with dark brown
scales ; cell a white band extending to wing apex, sprinkled with a few
dark brown scales; discal spot small, black; cubitus traced with black
sometimes extending from cell to Ai fold; wings grayish-red posterior
to black trace; A2 traced with black. Underside brown to grayish
brown, cell hghter. Hindwings light brown above and below, darker
apicaUy.
Genitalia as described for the genus.
Type.— In the Museum National d'Histoire Naturelle.
Type data. — Lectotype male, hereby disignated. Archer, Florida,
*'23/3 82": collection C. V. Kiley; male genitalia sHde No. 550, J.
Shaffer, July 28, 1965.
Specimens EXAMINED. — 180 cT, 39 9.
Distribution (Map 1). — Atlantic and GuK Coastal Plain, eastern
Texas to Florida, northward into Vhginia.
UNITED STATES: Florida: Alachua Co., Archer, 1 c?, "23/3 82" [USNM];
Gainesville, 7 d^, 2 9 , July 8, 1927 (J. Speed Rogers) [CU]; 1 ? , July 10, 1927;
Charlotte Co., Punta Gorda, 1 ?, Apr. 11, 1952 (J. R. Vockeroth) [CNC]; 1 cT,
1 9, Apr. 12, 1952 (G. S. Walley); Dade Co., Princeton, 2 cf, 1 9, Apr. 4,
1952 (J. R. Vockeroth) [CNC]; South Campus, Richmond, 1 &, Nov. 27, 1951
(F. G. Butcher) [CNC]; Highlands Co., Lake Placid, Archbold Biol. Sta., 1 9,
Apr. 2, 1945 (J. G. Needham) [CNC]; Archbold Biol. Sta., 3 d^, June 17, 1964
(Jay C. Shaffer) [JCS]; 8 d^, 2 9 , June 18, 1964; 2 c?, June 20, 1964; 3 d', June 22
1964; 1 d", June 23, 1964; 1 d", 1 9 , June 25, 1964; 2 d', June 26, 1964; 1 9
June 27, 1964; 3 d^, 5 9 , June 28, 1964; 5 d", 2 9 , June 29, 1964; 3 d", July 1
1964; 2 d^, July 2, 1964; 12 d^, 2 9 , July 3, 1964; 4 d^, July 4, 1964; 20 d', 7 9 ,
July 5, 1964; 5 d^, July 6, 1964; 1 cf 1 9 , July 7, 1964; 1 d", July 11, 1964; 36 d",
3 9 , July 12, 1964; 6 &, July 13, 1964; 1 6", Dec. 21, 1964 (S. W. Frost)[ CPK];
Manatee Co., Oneco, 1 d", April 1954 (Paula Dillman) [CPK]; Monroe Co.,
Key Largo, 2 d", Nov. 6, 1964 (Mrs. Spencer Kemp) [CPK]; 2 d, Nov. 16, 1964;
Pasco Co., Elfers, 1 d, Apr. 10, 1952 (J. R. Vockeroth) [CNC]; Pinellas Co., St.
Petersburg, 1 d, September [USNM]: Sarasota Co., Siesta Key, 1 d, Apr. 16,
1953 (Charles P. Kimball) [CPK]; 1 d, Mar. 28, 1954; 1 &, Apr. 1, 1954; 1 &,
Apr. 27, 1956; 4 d, Oct. 24, 1956; 1 d, Jan. 28, 1957; 2 9, May 21, 1957; St.
Johns Co., Hastings, 1 &, Aug. 1, 1900 [USNM].
Georgia: Bryan Co., Clyde, 1 d, Sept. 11-12, 1931 (Bradley and Knorr)
ICU].
Louisiana: Natchitoches Parish, 2 d, August (G. CoverdaJe) [USNM];
Vernon Parish, 1 d^, 1 9 , July (G. Coverdale) [USNM].
Texas: Collin Co., Piano, 1 9, August (E. S. Tucker) [USNM]; Fort Bend
Co., Richmond, Brazos River, 19 d^, 4 9 , June 22, 1917 [CU]; 1 d" [CNC];
Wharton Co., Wharton, 6 d', 1 9, June 24, 1917 [CU]; Victoria Co., Victoria,
1 d, June 24, 1917 [CU]; Victoria, 1 d, no date [USNM].
Virginia: Dinwiddle Co., Dinwiddie, 1 d, June 1, 1917 [CU].
78 U.S. NATIONAL MUSEUM BULLETIN 280
Barberia Dyar
Barberia Dyar, 1905, p. 39. — Barnes and McDunnough, 1917, p. 149 — Hamp-
son, 1918, p. 131. — McDunnough, 1939, p. 36. [Type: Barberia affinitella
Dyar, 1905. Monobasic]
Diagnosis. — The distinctive uncus, basally triangular and with a
narrow terminal portion, distinguishes the genus from all other New
World phycitines with similar wing venation.
Description. — Frons oblique; labial palpi upturned, slender;
maxillary palpi well developed, reaching frons; tongue exposed between
palpi; antennae filiform, compressed; ocelli small, appearing as dark-
ened spots.
Forewings with 9 veins; II2 from near upper outer angle of cell;
R3H-4 fused, separate from R2 at base; Mi from angle; M2+3 fused,
from lower outer angle; Cui from just before angle; Cuj from before
angle. Hindwing with 7 veins; Sc and Rs long stalked; Mi from upper
outer angle of cell; M2+3 fused, short stalked with Cui, from lower
outer angle; Cu2 from before the angle.
Male genitalia with uncus triangular on basal two-thirds, distal
third narrow, distal half heavily setose dorsally and laterally. Gnathos
arms terminating in anterior ventral hook; medial process a flattened
posteriorly directed hook, cur^^ed dorsally at tip. Transtilla incomplete,
a pair of flat plates about 3^ times as long as wide. Juxta U-shaped,
arms sharp pointed. Vinculum truncate. Valvae with apex obliquely
rounded. Aedeagus straight, vesica unarmed.
Female genitalia with ovipositor tip broadly rounded. Apophyses
straight, broadened basaUy; posterior parallel, anterior slightly longer,
diverging. Ductus bursa narrow, leaving bursa about one-fifth from
posterior end. Ductus seminalis from posterior end of bursa. Bursa
well developed; signum a long narrow sclerotized infolding about one-
fourth from anterior end of bursa.
Discussion. — The male genitalia are quite similar to Heinrich's
illustration of Anderida, but males of Barberia differ in the truncate
vinculum and lack of hair tufts on the eighth abdominal segment.
The two genera are less similar with respect to female genitalia and
wing venation.
Barberia affinitella Dyar
Figures 24, 102, 141, 176
Barberia affinitella Dyar, 1905, p. 39. — Barnes and McDunnough, 1917, p. 149.
— Hampson, 1918, p. 131.— McDunnough, 1939, p. 36.
Diagnosis. — This is the only known species in the genus.
Description. — Labial palpi with basal segments and inner sides
of second and third white, outer sides brown ; maxillary palpi brown ;
REVISION OF PEORIINAE AND ANERASTITNAE 79
antennae brown scaled on all sides; frons, occiput, patagia, and tegulae
brown, lighter dorsally.
Forewings with brown-sprinkled white band anterior to cell, reaching
just short of apex, ground brownish orange, lighter posterior to cell.
Hindwing light brown above and below, slightly darker toward apex.
Genitalia as described for the genus.
Types. — In the U.S. National Museum.
Type data. — Lectotype, hereby designated, male, Brownsville,
Tex., Los Borregos, June 6, 1904, H. S. Barber; USNM 8196; genitalia
sUde No. 576, J. Shaffer, Nov. 23, 1965. Labeled in Dyar's handwrit-
ing: "Barberia qffiniteUa Type Dyar."
Lectoparatypes: Brownsville, Tex., Los Borregos, male, two females,
June 5, 1904; two males, June 6, 1904, H. S. Barber; USNM 8196.
The original description gives June 5, 1904 as the date for all six
specimens.
Other SPECIMENS EXAMINED. — 23 cf, 4 9.
Distribution (Map 3).
UNITED STATES: Arizona: Cochise Co., Paradise, 3 cf , August, September,
October (Poling) [CN]; Southwestern Res. Sta., 5 mi. west Portal, 1 cf, Apr. 27,
1961 (M. A. Cazier) [AMNH]; Santa Cruz Co., Madera Canyon (4880 ft.),
Santa Rita Mts., 1 cT, Sept. 24, 1959 (R. W. Hodges) [JGF]; county unknown,
5 cf, August (0. C. Poling) [USNM].
Texas: Cameron Co., Brownsville, Los Borregos, 4 cf, 4 9 , June 5, 1904
(H. S. Barber) [USNM]; 1 cf , June 6, 1904; Brownsville, 4 d', June 1904 (H. S.
Barber) [USNM]; 1 cf , March 27-28 (F. H. Benjamin) [USNM]; San Benito,
1 d", March 24-30 [USNM]; 1 cf , June 24-30; JeflF Davis Co., Fort Davis, 1 cf ,
May 20, 1950 (E. C. Johnston) [CNC].
Ragonotia Grote
Ciris Ragonot, 1887, p. 17 [not Koch, 1848, p. 85 in Arachnida; not Grote, 1863,
p. 65, in Agaristidae]. [Type: Ciris discigerella Ragonot, 1887. Original
designation.]
Ragonotia Grote, 1888, p. 75. — Ragonot, 1889, p. 117. — Hulst, 1890, p. 204. —
Smith, 1891, p. 84. — Ragonot, 1901, p. 329. — Hulst, 1902, p. 437. — Barnes
and McDunnough, 1917, p. 149. — Hampson, 1918, pp. 123-124. — McDun-
nough, 1939, p. 35. [Type: Ciris discigerella Ragonot, 1887.]
Psammia Hampson, 1930, p. 71. — McDunnough, 1939, p. 31. — Heinrich, 1956,
p. 315. [New synonymy. Type: Psammia flavipicia Hampson, 1930. Mono-
basic]
Diagnosis. — Among the phycitine genera with reduced tongue and
prominent transverse Mang markings, the presence of vein M2 in the
hindwing distinguishes Ragonotia from Laetilia and Martia.
Description. — Frons rounded, smooth ; labial palpi decurved, about
3^ to 4 times eye length; maxillary palpi greatly reduced, usually not
easily discernible; tongue greatly reduced; antennae filiform, male
wdth ventral side of segments convex, cUia about twice segment width.
80 U.S. NATIONAL MUSEUM BULLETIN 2 80
forming two bands, female with cilia about half segment width; ocelli
well developed.
Forewings with 11 veins; R2 free from cell or short stalked with
R3+4; 1^3+4 stalked; Mi from about one-third distance from top of cell;
M2-1.3 short stalked, from lower outer angle of cell; Cui from before
the angle. Hindwing mth 8 veins; Sc and Rs approximate for short
distance beyond cell, then diverging; Mi from upper outer angle; M2
stalked with M3 for about two-thirds length; Cui stalked with M2+3
for about one-third length, from lower outer angle; Cu2 from just
before angle.
Male genitalia with uncus triangular, apex rounded, dorsal surface
uniformly and rather densely covered with short cilia. Gnathos with
medial process simple, tip hooked dorsally. Transtilla incomplete.
Juxta bell-shaped. Vinculum rounded. Valvae rounded, densely pu-
bescent on inner side; costa tubular; sacculus with clasper bearing
scattered setae. Aedeagus with vesica bearing a single large sharp-
pointed cornutus and a patch of numerous short, sharp spines.
Female genitalia mth dorsal margin of ovipositor rather densely
covered with long hairs. Apophyses rather straight, anterior slightly
longer and stouter than posterior. Bursa small, very weakly sclerotized,
unarmed.
Ragonotia dotalis (Hulst)
Figures 38, 62, 103, 142, 177
Anerastia dotalis Hulst, 1886, p. 164. — Rindge, 1955, p. 161.
Ciris discigerella Ragonot, 1887, p. 17.
Anoristia olivella Hulst, 1888, p. 117. — Rindge, 1955, p. 168.
Ragonotia saganella Hulst, 1890, p. 205. — Smith, 1891, p. 84. — Hulst, 1902,
p. 437. — Rindge, 1955, p. 170. [New synonymy.]
Ragonotia dotalis (Hulst). — Ragonot, 1889, p. 117. — Hulst, 1890, p. 205. —
Smith, 1891, p. 84.— Ragonot, 1901, p. 329.— Hulst, 1902, p. 437.— Barnes
and McDunnough, 1917, p. 149. — Hampson, 1918, p. 124. — McDunnough,
1939, p. 35.
Megasis indianella Dyar, 1923, pp. 28-29 [described in Phycitinae].
Psammia flavipicta Hampson, 1930, pp. 71-72. — McDunnough, 1939, p. 31. —
Heinrich, 1956, p. 315.— Kimball, 1965, p. 251. [New synonymy.]
Diagnosis. — This is the only known species in the genus.
Description. — Labial palpi white, outer sides of second and third
segments variously sprinkled with subapically brown banded white
scales; antennae white, brown band on center of each segment; frons
white with scattered brown scales, vertex and dorsum of occiput
white, often with scattered brown scales, occiput laterally of alternat-
ing black and white bands; legs white, sprinkled with brown on outer
sides, proximal end of each tarsal segment black banded.
Forewings with ground white, finely sown with dark brown scales,
variable in number, producing a light brown to gray color; basal spot
REVISION OF PEORIINAE AND ANERASTIINAE 81
light orange, usually indistinctly bordered with black distally; trans-
verse anterior light orange, partially bordered internally with black,
externally with narrow white band, then black blending graduallj^
into ground color; discal spot light orange, usually with white center,
spot bordered above and below with black; transverse posterior
light orange, bordered internally with narrow white band, then
black, veins traced with black on tp band, tracing extending to the
black terminal line.
Hindwings light brown, apical area and terminal Une brown;
fringe white, brown band near its base; base of cubitus fringed, A2,
A3, and inner margin heavily fringed.
Genitaha as described for the genus.
Types. — A. dotalis, in the American Museum of Natural History
(lectotype); C. discigerella, in the Museum National d'Histoire
Naturelle; A. olivella and R. saganella, in the American Museum of
Natural History; M. indianella, m the U.S. National Museum; P.
flavipicta, in the British Museum (Natural History).
Type data. — A. dotalis, lectotype female, hereby designated.
Phoenix, Ariz,, June (abdomen intact); discigerella, lectotype female,
hereby designated, Arizona, genitalia slide No. 555, J. Shaffer, Aug. 11,
1965; olivella, lectotype female, hereby designated, southern California,
genitaha shde No. 587, J. Shaffer, Nov. 28, 1965, in the origmal
description the type locality is given as Needles, Calif.; saganella,
lectotype female, hereby designated, Colorado, May, genitalia slide
No. 3217, Carl Heinrich, June 14, 1946, in the original description
Hulst reports "From T.D.A. Cockerell, of West Cliff, Col., where
the unique specimen was taken at light. May 24th."; indianella, one
female, Indian Wells, Calif., May 8, 1921, USNM 25840, genitaha
slide No. 1103, Carl Heinrich, Apr. 2, 1938; flavipicta, lectotype
male, hereby designated, labeled as follows: "Type; Florida; 99-49.;
1945/206; Psammia flavipicta type cT. Hmpsn.; cf genitaha shde
1-4-1967 J. Shaffer No. 708." The vahdity of Florida as the type
locality must be questioned as the species has not otherwise been
recorded east of western Texas.
Specimens EXAMINED. — 75 cf , 38 9.
Distribution (Map 1). — Southwestern United States, southern
California eastward into western Texas.
UNITED STATES: Arizona: Cochise Co., Paradise, 1 9 [USNM]; San
Bernardino Ranch (3750 ft.), 2 cf , August (F. H. Snow) [USNM]; 2 ? , [UK];
Maricopa Co., Phoenix, 3 d^, 4 ? , March (Kunze) [USNM]; 1 d^, April; 1 ? ,
June; 1 9 , September; 1 cT, no date; 1 9 , June [USNM]; 1 cf, June [USNM];
labeled "Ragonotia dotalis Type Hulst," "Type No. 40073 U.S.N.M.," Male
genitalia slide No. 1100, Carl Heinrich, Apr. 2, 1938; 1 9, AprU [AMNH]; 1 cT,
May [AMNH]; 1 9 , June [AMNH]; 1 9 , no date [AMNH]; Tempe, 1 cf , Mar. 8,
1920 (E. V. Walter, H. L. Arnold) [USNM]; Pima Co., Santa Rita Mts., 1 9 ,
82 U-S. NATIONAL MUSEUM BULLETIN 280
May 1-7 [USNM]; Yavapai Co., 1 &, no date [USNM]; Yuma Co., Indian Wash
near Martinez Lake, 1 d", June 12, 1961 (C. A. Toschi) [UCB]; Palomas, 2 d',
Aug. 8, 1917 [CU]; Wellton, 2 d', Aug. 9, 1917 [CU]; Yuma, 1 ? , May [AMNH];
no county, southern Arizona, 1 cf, no date [USNM]; 1 $ , Sept. 8-15 [USNM].
California: Imperial Co., Dixieland, 1 cf. Mar. 1-15, 1922 (O. C. Poling),
[USNM]; 3 cf", 2 ? , spring 1922; Travertine Rock, 1 cf , Apr. 10, 1954 (Stange,
Menke, Cline, and Thomas) [LACM]; Riverside Co., Coxcomb Mts., 5 d^, 1 ? ,
Feb. 28, 1937 (Lloyd M. Martin) [LACM]; (J. A. Comstock); 1 c?, 3 9 , Mar. 1
1937 (Lloyd M. Martin); Dis Palmas Springs, 1 9, Mar. 17, 1940 (George
Willett) [LACM]; Indian Wells, 1 &, 2 9, Mar. 8, 1940 (J. C. von Bloeker)
[LACM]; Indio, 1 cf, Mar. 23, 1955 (W. R. M. Mason) [CNC]; La Quinta, 1 &,
Mar. 5, 1955 (J. E. H. Martin) [CNC]; Palm Desert, 1 d^, Mar. 15, 1954 (A. H.
& S. K. Rindge) [AMNH]; Thermal, 5 d^, 1 9 , Aug. 17-18, 1927 [CU]; 1 &,
July 10, 1956 (M. Wasbauer) [UCB]; Thousand Palms, 1 &, Feb. 24, 1955
(D. F. Hardwick) [CNC]; Feb. 24, 1955 (W. R. Richards); (W. R. M. Mason)
Feb. 28, 1955 (D. F. Hardwick); 1 9 , Mar. 1, 1955 (W. R. M. Mason); Id", 19,
Mar. 2, 1955 (D. F. Hardwick); 1 d", Mar. 5, 1955 (W. R. M. Mason); 4 d",
Mar. 7, 1955; 2 d", 1 9, Mar. 8, 1955 (J. E. H. Martin); 1 &, Mar. 10, 1955
(W. R. M. Mason); 2 d", Mar. 11, 1955 (J. E. H. Martin); 1 &, Mar. 13, 1955;
2 d', Mar. 18, 1955; 1 <?, Mar. 23, 1955 (W. R. M. Mason); 1 9 , Mar. 24, 1955
(J. E. Martin); Mar. 28, 1955 (W. R. Richards); 1 d", Apr. 16, 1955 (W. R. M.
Mason); San Bernardino Co., Baker, 1 d", July 30, 1954 (L. Martin & Rees)
[LACM]; Yermo, Mojave Desert, 1 6", Apr. 10, 1936 (J. A. Comstock) [USNM];
near Yermo, 5 d", [LACM]; Yermo, 3 d", Mar. 18, 1940 (G. Beevor) [LACM];
San Diego Co., Borrego, 1 9 , Mar. 22, 1940 (Grace H. & John L. Sperry) [CNC];
Borrego, 1 9 , May 3, 1956 (J. Powell) [UCB]; Apr. 21, 1960 (J. F. Lawrence).
New Mexico: Dona Ana Co., Mesilla Park (3800 ft.), 2 d", July 8 (Cockerell)
[USNM]; Mesilla, 1 d", no date (C. N. Ainslie) [USNM]; Mesquite near Mesilla
Park, 1 d', July 12, 1917 [CU]; Hidalgo Co., Lordsburg, 2 9 , May 9, 1950 (E. C.
Johnston) [CNC]; Luna Co., 10 mi. east of Deming, 1 d", July 12, 1917 [CU].
Texas: Reeves Co., Pecos, 2 9, May 18, 1950 (E. C. Johnston) [CNC];
Brewster Co., Marathon, 2 d", May 23, 1950 (E. C. Johnston) [CNC].
Utah: No locality, 1 9 , Henry Edwards collection. No. 14158 [AMNH].
Discussion. — Tliere is uncertainty regarding the identity of tlie
type specimens of dotalis and saganella. In the American Museum of
Natural History is a series of two females and one male, all labeled
Phoenix, Ariz., June, "Anerastia dotalis Type Hulst" (Hulst's hand-
writing); the male specimen also bears a slide preparation label,
genitalia slide No. 3216, Carl Heinrich, June 14, 1946. One of the
females has the tip of its abdomen broken off. A female specimen
in the U.S. National Museum also bears Hulst's type label and the
data Phoenix, Ariz., May. In his original description of dotalis,
Hulst reported only that he had one female from Arizona. In the
absence of any evidence indicating which specimen Hulst based
his original description upon, I have chosen as the lectotype the more
nearly intact of the two female specimens in the type series of the
American Museum of Natural History. The other three "types" are
thus presumed to be pseudotypes.
REVISION OF PEORIESTAE AND ANERASTIINAE 83
In describing saganella, Hulst indicated clearly that his new species
was very different from dotalis and stated that the two were "hardly
congeneric." Hulst's type of saganella, on the other hand, is ob-
viously a specimen of dotalis, thus the validity of the type and the
accuracy of the description are open to question.
Martia Ragonot
Martia Ragonot, 1887, p. 18; 1889, p. 117. — Hulst, 1890, p. 209. — Smith,
1891, p. 84. — Ragonot, 1901, p. 367. — Hulst, 1902, p. 439. — Barnes and
McDunnough, 1917, p. 149. — Hampson, 1918, p. 118. — McDunnough,
1939, p. 36. [Type: Martia arizonella Ragonot, 1887. Monobasic]
Urula Hulst, 1900, p. 175; 1902, p. 437. [Type: Urula incongruella Hulst, 1900.
Monobasic]
Diagnosis. — The genus is easily recognized by the combination
of protuberant frons and prominent transverse lines on the forewings.
Description. — Frons produced into obliquely truncate cone,
its rim incised ventrally, produced and finely serrate laterally, deeply
incised dorsally forming shallow trough rimmed with fine teeth; labial
palpi porrect or somewhat decurved, about 3J^ times eye length in
male, and about four times in female; maxillary palpi cylindrical, al-
most reaching frons; tongue rudimentary; antennae subserrate in male,
each segment with two ventral bands of cUia, cilia about 2% times as
long as segment wddth, filiform in female, ciha not in bands, shorter
than segment width; ocelli small, conical.
Forewings with 11 veins; R2 free from cell near R3+4; R3+4 stalked,
from upper outer angle of cell; Mi arising about one-third from top
of cell; M2+3 short stalked or from point at lower outer angle of cell;
Cui from just before the angle. Hindwing with 7 veins; Sc and Rs
approximate for short distance beyond cell; Mi from upper outer
angle of cell; M2+3 fused, short stalked A\dth Cui, from lower outer
angle.
Male genitalia mth uncus triangular, rounded apically, finely ciliate
dorsally and laterally. Gnathos with medial process simple, extending
caudad, tip sharp, curving dorsad. Transtilla a pair of weakly sclero-
tized irregular plates. Juxta scoop-shaped. Vinculum broader than
long. Costa tubular on basal four-fifths, tapering. Aedeagus slender,
vesica unarmed.
Female genitalia with ovipositor rather densely setose, tip broadly
rounded. Posterior apophyses curved posteroventrally on posterior
fourth; anterior straight, directed an tero ventrally at angle of about
45° to posterior. Ductus bursae short. Bursa unarmed. Ductus semi-
nalis leaving bursa about one-fourth distance from posterior end.
84 U.S. NATIONAL MUSEUM BULLETIN 280
Martin arizonclla Ragonot
Figures 37, 49, 63, 104, 143, 178
Martia arizonella Ragonot, 1887, p. 18; 1889, p. 117. — Hulst, 1890, p. 209. —
Smith, 1891, p. 84. — Ragonot, 1901, p. 367. — Hulst, 1902, p. 439. —
Barnes and McDunnough, 1917, p. 149. — Hampson, 1918, p. 118. — Mc-
Dunnough, 1939, p. 36.
Urulaincongruella Hulst, 1900, pp. 175-176; 1902, p. 437. — Rindge, 1955, p. 164.
Diagnosis. — This is the only known species in the genus.
Description. — Frons close scaled, brown; labial palpi with basal
segments light brown, second and third segments clothed with white-
tipped brown scales on outer sides, lighter on inner sides; maxillary
palpi of white to white-tipped brown scales; antennae white, each
segment brown banded; vertex brown, lighter behind antennae; occi-
put, patagia, and tegulae varying white to brown; legs clothed with
light brown to white-tipped brown scales on outer sides, distal end of
each tibial segment white banded.
Forewings with transverse anterior, transverse posterior, discal
spot, area between Cu2 and A2, and basal area posterior to A2 golden
yellow; transverse anterior interrupted in cell by dark brown area
which gradually narrows and extends posteriorly to A2 as a wedge
between the ta line and the prominent narrow white line bordering it
distally. Transverse anterior bordered inside by white anterior to cubi-
tus, by brown band posterior to cell, band extending to costa at wing
base; wing anterior to cell and between transverse lines white with
sprinkling of brown scales; area surrounding discal spot brown, ex-
tending posteriorly to Cu2, often enclosing an irregular white area at
lower outer angle of cell; terminal line dark brown, bordered internally
by white wedge on anterior third of wing. Underside light brown,
darker in apical area; transverse posterior faintly traced. Hindwings
light brown above and below, fringe white with darker band near its
inner side.
Genitalia as described for the genus.
Types. — M. arizonella, in the Museum National d'Histoire Natu-
relle; U. incongruella, in the U.S. National Museum (lectotype), in the
American Museum of Natural History (two lectoparatypes) .
Type data. — M. arizonella, type not examined, the type locality is
given in the original description as Arizona; U. incongruella, lectotype
male, hereby designated, Argus Alts., April, 91K, USNM 4708, male
genitalia slide No. 1121, Carl Heinrich, Feb. 27, 1941 [USNM]; lecto-
paratypes, two females, Phoenix, Ariz., June 4, June 5, the former with
female genitalia on slide No. 3218, Carl Heinrich, June 14, 1946, the
latter has the abdomen lost [AMNH]. In his original description Hulst
gives the following data: "Argus Mountains, Cal.; taken by Mr. Koe-
bele, in April. Phoenix, Ariz.; taken early in June."
REVISION OF PEORIINAE AND ANERASTIESTAE
85
Specimens EXAMINED. — 83 6^,43 9.
Distribution (Map 3).
UNITED STATES: Arizona: Coconino Co., Tuba City, 2 cf, 5 ?, Aug. 12,
1948 (C. and P. Vaurie) [AMNH]; Maricopa Co., Phoenix, 1 9, June 4, 1897,
(bears label: "Urula incongruella Type Hulst") [USNM]; Phoenix, 2 9 , Sept. 11,
1904 [USNM]; 1 9 , Sept. 13, 1904 [USNM]; Pima Co., 30 mi. east of Quijotoa, 1
9 , Aug. 28-29, 1927 [CU]; Yuma Co., Dome, 1 cf, July 21, 1924 (E. P. Van Duzee)
[USNM]; Indian Wash near Martinez Lake, 1 cf, 2 9, June 12, 1961 (C. A.
Toschi) [UCB]; Palomas, 3 d", Aug. 8, 1917 [CU]; no locality, 1 9 , July 1-7
[USNM]; 2 9 , June 8-15 [USNM].
California: Kern Co., Shafter, Icf, 19, Aug. 17, 1955 (J. Powell) [UCB];
Riverside Co., Coachella, 1 9 , May 2, 1918 (J. C. Bradley) [CU]; Indio, 3d^, 7 9 ,
May 1, 1918 (J. C. Bradley) [CU]; Palm Springs, Icf, May 6, 1921 (Karl B. Cool-
idge) [USNM]; 1 cf, Aug. 20, 1955 (A. H. Rindge) [AMNH]; Riverside, Icf,
Sept. 8, 1940 (H. Buckwalter) [CNC]; Thermal, 19, June 17, 1956 (M. Was-
bauer) [UCB]; San Bernardino Co., 10 mi. north of Earp. 2^^, 19, Apr. 22, 1960
(J. R. Powers) [UCB]; Loma Linda, 1 cf , Sept. 1-7 [USNM]; Mojave Desert,
Yermo, 1 d^, Apr. 10, 1936 (J. A. Comstock) [USNM] ;Needles, 33d^, 9 9 , Apr. 24,
1950 (E. C. Johnston) [CNC]; Twentynine Palms, 12 0", 29, Apr. 20, 1950
(E. C. Johnston) [CNC]; 7cf, Apr. 21, 1950; San Diego Co., 6 mi. east of Banner,
19, July 13, 1963 (J. Powell) [UCB]; county unknown, LaPuerta, Icf, 19,
July 1911 (Wright and Field) [USNM]; LaPuerta VaUey, 3cf, July 11 (George H.
Field) [USNM].
New Mexico: Dona Ana Co., Mesquite near Mesilla Park, Icf, July 12, 1917
[CNC]; 6cf , 1 9 [CU]; Otero Co., Alamogordo, 1 9 , May 10, 1950 (E. C. Johnston)
[CNC]; 25 mi. west of Tularosa, 1 9 , July 1, 1940 (L. J. Lipovsky) [UK].
Synonymical List of Species Considered
North American Peoriinae
Peoria Ragonot
Aurora Ragonot
Statina Ragonot
Calera Ragonot
Altoona Hulst
Cayuga Hulst
Volusia Hulst
Wekiva Hulst
Osceola Hulst
Chipeta Hvilst
Trivolusia Dyar
Ollia Dyar
1. longipalpella (Ragonot)
2. bipartitella Ragonot
roseopennella (Hulst)
3. tetradella (Zeller)
4. opacella (Hulst)
dichroeella (Ragonot)
dichroella (Hampson)
5. floridella Shaffer
6. rostrella (Ragonot)
7. gemmatella (Hulst)
bistriatella (Hulst)
pamponerella (Dyar)
8. roseotinctella (Ragonot)
punctilimbella (Ragonot)
bifascieUa (Hampson)
9. johnstoni Shaffer
10. santaritella (Dyar)
11. holoponerella (Dyar)
12. approximella (Walker)
haematica (Zeller)
roseatella (Packard)
cremoricosta (Hampson)
13. luteicostella (Ragonot)
nodosella (Hulst)
perlepidella (Hulst)
Anacostia Shaffer
14. tribulella Shaffer
86
U.S. NATIONAL MUSEUM BULLETIN 280
Arivaca Shaffer
The Pimella Group
15. pimella (Dyar)
16. linella Shaffer
The Ostreella Group
17. ostreella (Ragonot)
discostrigella (Dyar)
18. poohella Shaffer
The Albidella Group
19. albidella (Hulst)
20. artella Shaffer
The Aibicostella Group
21. aibicostella (Grossbeck)
Atascosa Hulst
22. glareosella (Zeller)
bicolorella Hulst
albocostella (Hulst)
Homosassa Hulst
23. ella Hulst
24. platella Shaffer
25. incudella Shaffer
Reynosa Shaffer
26. floscella (Hulst)
Goya Ragonot
27. stictella (Hampson)
Genera Transferred to Phycitinae
Anerastia Hiibner
Prinanerastia Hampson
1. lotella Hiibner
Coenochroa Ragonot
Petaluma Hulst
Alamosa Hampson
2. californiella Ragonot
inspergella Ragonot
3. illibella (Hulst)
puricostella Ragonot
piperatella (Hampson)
4. bipunctella (Barnes and
McDunnough)
Bandera Ragonot
Nasutes Hampson
5. binotella (Zeller)
subluteella Ragonot
6. cupidinella Hulst
conspersella (Ragonot)
venata (Hampson)
7. virginella Dyar
Wakulla Shaffer
8. carneella (Barnes and McDunnough)
Tampa Ragonot
9. dimediatella Ragonot
Barberia Dyar
10. afEnitella Dyar
Ragonotia Grote
Ciris Ragonot
Psammia Hampson
11. dotalis (Hulst)
discigerella ( Ragonot)
olivella (Hulst)
saganella Hulst
indianella (Dyar)
flavipicta (Hampson)
Martia Ragonot
Urula Hulst
12. arizonella Ragonot
incongruella (Hulst)
REVISION OF PEORIINAE AND ANERASTIINAE 87
Unplaced Genera and Species
Navasota Ragonot
Figure 30
Navasota Ragonot, 1887, p. 18; 1889, p. 117. — Hulst, 1890, p. 212. — Smith,
1891, p. 85. — Ragonot, 1901, p. 369. — Hulst, 1902, p. 439. — Barnes
and McDunnough, 1917, p. 149. — Hampson, 1918, pp. 65-66. — Mc-
Dunnough, 1939, p. 36. [Type: Navasota hebetella Ragonot, 1887, p. 18.
Monobasic. References as given for the genus.]
Description. — The original descriptions of the genus and species
are as follows :
Navasota, gen. nov. — Antennae thick, pubescent, a very flattened pad of
scales in sinus. Palpi oblique, ascending, 3rd joint horizontal. Fore-wings with
ten veins, 10 from stem of 8 and 9, 4 and 5 stemmed, 3 wanting. Hind wings
with seven veins, 8 stalked, 4 and 5 on long stem, 3 wanting, 2 before angle.
86. N. hebetella. — 16 mill. — Fore-wings reddish-ochreous, paler on inner
margin. Costa with a broad whitish streak not reaching the apex and finely
streaked with reddish-ochreous. Hind- wings pale yellowish.
Type. — In the Museum National d'Histoire Naturelle.
Type data. — A specimen label gives: "ex. coll. Bellfr. coll. Riley."
Discussion. — Navasota probably belongs in the Peoriinae, but its
exact placement is uncertain as the abdomen of the type is lost. The
species may belong to Peoria, but apparently is not conspecific with
any of the 13 species listed in that genus. No type locality is given
in the original description, but Hampson (in Ragonot, 1901) in a
probable reference to the type reports: "Texas. Un cf, octobre.
Coll. Ragonot." All of Belfrage's southwestern material was collected
in Texas.
Uinta Hulst
Uinta Hulst, 1888. p. 116. — McDunnough, 1939, p. 35. Type: Uinta oreadella
Hulst, 1888, p. 116. — Hulst, 1890, p. 221-222 [listed as a crambid]. — Mc-
Dunnough, 1939, p. 35. — Rindge, 1955, p. 168. [Monobasic]
Description. — The original descriptions of the genus and species
are as follows:
Uinta, n. gen.
Fore wings 9 veins: 4 and 9 wanting, 3 and 5 separate; hind wings 7 veins;
3 and 5 separate, 2 far from angle, 7 and 8 separate to base; labial palpi hori-
zontal, heavy, long; maxillary palpi strong, heavily scaled on end; antennae
of cf bent above base, pubescent; ocelli present; tongue wanting; legs shorter
and more heavy than usual. Type: oreadella, Hulst.
U. oreadella, n. sp.
Palpi fuscous gray, as is also the head; thorax blackish gray; abdomen
blackish gray, becoming fuscous and yellowish anally, each segment except
basal edged narrowly with gray; fore wings dark fuscous, darker at base;
basal line not evident; outer line broad, blackish, a marginal broad line also
blackish; at base of center of middle field a lengthened black point; hind wings
88 U.S. NATIONAL MUSEUM BULLETIN 2 SO
fuscous, shining, margin darker; beneath even fuscous on all wings, margins
darker. — Colorado."
Type. — In the American Museum of Natural History.
Type data. — Texas, June; G. D. Hulst collection, Kutgers.
Discussion. — The species probably belongs to the Crambinae as
Hulst (1890) later indicated, but cannot be placed to subfamily with
certainty as the hindwings and abdomen have been lost. The locality
given on the specimen (Texas) and that published in that original
description (Colorado) do not agree.
Tolitna cincaidella Dyar
Figure 36
Tolima cincaidella Dyar, 1904, pp. 115-116. — Barnes and McDunnough, 1917,
p. 149. — McDunnough, 1939, p. 35.
Description. — The original description of the species is as follows:
Fore wings with 11 veins, 2 well before angle of cell, 3 before angle, 4 and 5
very shortly stalked, 8 and 9 stalked, 10 connate with the base of the stalk,
11 from the cell. Hind wings with 7 veins, 2 from the angle of the cell, 3 and 4
long stalked, 5 absent, 8 distinct. Tongue moderate, apparently about twice
the length of the head; palpi very long, porrect and drooping; antennae simple,
lengthily ciliate.
Pale ocherous whitish, lines paler, obscure, defined by dusky edgings toward
the median space. Inner line produced a little on costa, preceded by a very
faint, russet patch on the inner margua. Discal dots blackish, separate. Outer
line drawn inward subcostally, else even, a slight dentation indicated in the
dusky border. The wing is all very pallid. Hind wing whitish, translucent.
Expanse 27 mm.
Three specimens, all with the abdomens broken, but apparently males,
Rock Spring, Wyoming (T. Kincaid).
Type: No. 7934, U.S. National Museum.
Type. — In the U.S. National Museum.
Type data. — No locality given on specimens, USNM 7934; printed
tag bears number "37049." The three specimens bear identical labels.
Discussion. — The species does not belong in the Peoriinae, but
otherwise placement to subfamily is uncertain.
Statina gaudiella Hulst
Statina gaudiella Hulst, 1890, pp. 216-217. — Smith, 1891, p. 85. — Ragonot,
1901, pp. 415-416. — Hulst, 1902, p. 440. — Dyar, 1904c, p. 228. — Barnes
and McDunnough, 1917, p. 150. — Hampson, 1918, p. 60. — McDunnough,
1939, p. 36. — Rindge, 1955, p. 163.
Description. — The original description of the species is as foUows:
2. S. gaudiella n.sp. — Expands 13 mm. Palpi and antennae ochreous fuscous;
abdomen ochreous. Fore wings squamose; ground color ochreous, with a faint
violet fuscous tinge, this sprinkled with lengthened black scales, much more
REVISION OF PEORIINAE AND ANERASTIESTAE 89
thickly about basal line and within outer line forming indeterminate blackish
bands; lines indistinct, indeterminate, not evident except by the blackish
scaling; costa on outer field darker, with black scales; hind wings yellowish
fuscous, darker on veins.
Differs from typical Statina in that cells of both wings are very short.
Type. — In the American Museum of Natural History.
Type data. — Central Texas, Blanco County, September, collection
G. D. Hulst.
Discussion. — The holotype consists of meso- and metathorax, the
right pair of wings, and the left hindwing. On the basis of wing
venation gaudiella probably belongs in the Peoriinae, but exact
placement is impossible owing to the poor condition of the holotype.
Ollia parvella Dyar
Ollia parvella Dyar, 1906, p. 31. — Barnes and McDunnough, 1917, p. 149. —
McDunnough, 1939, p. 36. — Kimball, 1965, p. 250.
Description. — The original description of the species is as foUows :
Costal half of fore wing white with slight darker lines on the veing toward
apex. Inner half pale ocherous, shading to gray next to white part. Hind wing
whitish. Expanse, 12 mm.
Six 9 , BrownsviUe, Tex., May 31 to June 9, 1904 (H. S. Barber).
Type. — In the U.S. National Museum.
Type data. — Holotype: Texas, Brownsville, female, June 3?, 1904;
H. S. Barber; USNM 9103; female genitalia sHde No. 10, Carl Hein-
rich, Dec. 20, 1932.
Discussion, — In the absence of male specimens placement is
uncertain. The species probably belongs in the Phycitinae and may
require a new genus when males can be examined. Ollia is a synonym
of Peoria.
Altoona ardiferella Hiilst, Aurora nigromaculella Hulst, and Parramatta
placidella Barnes and McDunnough
These three names belong in the Phycitinae and should probably
be referred to Laetilia Kagonot, but placement within the subfamily
should await a thorough study of Laetilia and allied genera.
Altoona ardiferella Hulst
Altoona ardiferella Hulst, 1888, p. 116; 1890, p. 208. — Smith, 1891, p. 84. —
Barnes and McDunnough, 1918, p. 176. — Rindge, 1955, p. 157.
Zophodia ardiferella (Hulst). — Ragonot, 1889, p. 116.
Tolima ardiferella (Hulst). — Ragonot, 1901, pp. 506-507.
Saluria ardiferella Hulst, 1902, p. 439. — Hampson, 1918, p. 99.
Pectinigera [sic] ardiferella (Hulst). — Dyar, 1904b, p. 159.
Pedinigeria ardiferella (Hulst). — Barnes and McDunnough, 1917, p. 149. —
McDunnough, 1939, p. 35.
90 U.S. NATIONAL MUSEUM BULLETIN 280
Type. — Possibly lost. A badly damaged specimen in the American
Museum of Natural History bears a Hulst type label and "Col., July,
Collection G D Hulst," female genitalia slide No. 3229, Carl Heinrich,
June 14, 1946. The original description gives the type locality as
Texas. See discussion by Barnes and McDunnough (1918).
Host. — Dyar (1904b) reports specimens bred from Orthezia annae
(Coccidae) on Atriplex canescens (Chenopodiaceae).
Aurora nigrotnaculella Hulst
Aurora nigramaculella Hulst, 1900 1901, 224: 1902, p. 438. — Rindge, 1955, p. 167.
Zohpodia nigromaculella (Hulst). — Dyar, 1904c, p. 228.
Saluria nigromaculella (Hulst), — Hampson, 1918, p. 100.
Type. — Holotype, in the U.S. National Museum.
Type data. — Arizona, Santa Rita Mts., June 8, 1898, E. A.
Schwarz; USNM 5185; female genitalia slide 2269, Carl Heinrich,
Sept. 11, 1934. A specimen in the American Museum of Natural
History bears a cotype label and "Hot Springs, N. Mex. 7000 ft. alt.
Sept., Collection G D Hulst"; abdomen lost. The original description
mentions only the Arizona specimen.
Discussion. — Dyar (1904b) and McDunnough (1939) consider this
to be a synonym of ardiferella Hulst.
Parramatta placidella Barnes and McDunnough
Parramatta placidella Barnes and McDunnough, 1918, p. 177, pi. 24, fig. 17. —
McDunnough, 1939, p. 35.
Types. — In the U.S. National Museum.
Type data. — Lectotype, hereby designated, male, Olancha, Inyo
County, Calif., June 24-30, Barnes collection, male genitalia sUde
No. 1127, Carl Heinrich, Feb. 6, 1942: labeled "Parramatta placidella
Type B & McD."
Lectoparatypes: Olancha, Inyo County, Calif., four females, all
labeled "Parramatta placidella Paratype B & McD," Barnes collection;
three specimens dated June 8-15, the fourth dated June 16-23 and
labeled female genitalia sUde No. 1128, Carl Heinrich, Feb. 6, 1942.
On all five specimens Inyo is misspelled on the label as "Inya."
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REVISION OF PEORIINAE AND ANERASTIINAE 91
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Explanation of Figures
The figures have not been reproduced to any fixed scale, and in
general smaller specimens are presented in greater enlargement than
larger ones.
Components of the male genitalia are enlarged to the same degree
for each species unless otherwise indicated. Sketches of the aedeagus,
gnathos, or transtilla have been added in addition to or in place of
photographs where these did not render the structures with sufBcient
clarity. Male genitalia of Coenochroa, Martia, Ragonotia, and all of
the Peoriinae have been dissected, opened, and flattened as described
on page 3.
Female genitalia of the Peoriinae are illustrated in lateral view,
those of phycitine specimens in dorsal view.
94
U.S. NATIONAL MUSEUM BULLETIN 280
Map I. — Distribution of species
^Jnerastia lotella QRagonotia dotalis
® Tampa dimediatella
Map 2. — Distribution of Coenochroa species
-^C californiella j^C. illibella
%C. bipunctella
REVISION OF PEORIINAE AND ANERASTIINAE
95
Map 3. — Distribution of species
^Barheria afinitella ■^Marita arizonella
Wakulla carnedla ^
Map 4. — Distribution of species
^Bandera binoUlla %B. cupidinella
B. virginella ^ -j^Anacostia tribulella
i^Goya stictella
' Open five-pointed star within black circle.
96
U.S. NATIONAL MUSEUM BULLETIN 2 80
Map 5.— Distribution of Peoria approximella
Map 6. — Distribution of Peoria species
•P. gevimatella Qp, roseotinctella
P. rostrella *
> Open five-pointed star within black circle.
■jIV-P. tetradella
REVISION OF PEORIINAE AND ANERASTIINAE
97
Map 7. — Distribution of Peoria species
%P. holoponerella ^P. johnstoni
DP. opacella Q)P. santaritella
Map 8. — Distribution of Peoria species
#P. hipartitella P. longipalpella '■
Map 9. — Distribution of Arivaca species
A. albidella ' ^A. arUlla
UA. poohella %A. linella
if A. ostreella QA. pimella
* Open five-pointed star within black circle.
Map 10. — Distribution of species
O Peoria floridella %P. luteicostella
if Arivaca albicostella
98
U.S. NATIONAL MUSEUM BULLETIN 28
Map 11. — Distribution of Homosassa species
•^- e^l'^ H. incudella i
*//. platella
Map 12. — Distribution of species
% Atascosa glareosella Reynosa fioscella '
* Open five-pointed star within black circle.
REVISIOX OF PEORIIXAE AND ANERASTIINAE
99
Figures 1-6. — Wing Expanse of Adults: 1, Peoria johnstoni, 19mm; 2, P. rostrella, holo-
type, approximately 24 mm; 3, P. holoponerella, 24 mm; 4, P. holoponerella, 28 mm;
5, P. santaritella, 25 mm; 6, P. tetradella, approximately 24 mm.
100
U.S. NATIONAL MUSEUM BULLETIN 2 80
Figures 7-12.— Wing Expanse of Adults: 7, Peoria gemmatella, 26 mm; 8, P. floridella,
holotype, 26 mm; 9, P. longipdpella, 23 mm; 10, P. roseotinctella, 19 mm; 11, P. opacella,
23 mm; 12, P. luteicostella, 22 mm.
REVISION OF PEORIINAE AND ANERASTIINAE
101
Figures 13-18.— Wing Expanse of Adults: 13, Peoria hipartitella, 17 mm; 14, P. approxl-
mella, 19 mm; 15, Arivaca pimella, holotype, 25 mm; 16, A. linella, 25 mm; 17, A. al-
hidella, 24 mm; 18, A. artella, 25 mm.
102
U.S. NATIONAL MUSEUM BULLETIN 2 80
Figures 19-24.— Wing Expanse of Adults: 19, Arivaca ostreella, 28 mm; 20, A. poohella,
holotype, 27 mm; 21, A. albicostella, 18 mm; 22, Reynosafloscella, 13 mm; 23, Goya stictella,
17 mm; 24, Barberia affinitella, 14 mm.
REVISION OF PEORIINAE AND ANERASTIINAE 103
Figures 25-30. — Wing Expanse of Adults: 25, Anacostia tribulella, holotype, 23 mm;
26, A. glareosella, 22 mm; 27, Homosassa ella, 17 mm; 28, H. platella, holotype, 17 mm;
29, H. incudella, para type, 21 mm; 30, Navasota herhetella, holotype, approximately 16 mm.
104
U.S. NATIONAL MUSEUM BULLETIN 2 80
Figures 3 1-36. — Wing Expanse of Adults : 31, Bandera bmote/la, 23 mm ; ^^2, B. cupidinella
22 mm; 33, B. virginella, 24 mm; 34, Anerastia lotella, 29 mm; 35, Tampa dimediatella,
17 mm; 36, Tolinia cincaidella, holotype, 28 mm.
REVISIOX OF PEORIINAE AND ANERASTIINAE
105
Figures 37— ^}i. — Wing Expanse of Adults: 37, Mania arizonella, 17 mm; 38, Ragonotia
dotalis, 29 mm; 39, Coenochroa illibella, 30 mm; 40, C. illibella, 22 mm; 41, C. californiella,
20 mm; 42, C. bipunctella, 18 mm; 43, Wakulla carneella, holotype, approximately 11 mm.
106
TJ.S. NATIONAL MUSEUM BULLETIN 2 80
46a
46b
47 b
48b
Figures 44-48.— Adult crania: 44-48, lateral view; 44^-48^, anterior view; 44Z--48^
dorsal view. 44-44^-, Coenochroa californiella; 45-45^, C. illibella, typical; 46-46^ C.
illibella, specimen from series taken at Pecos, Tex.; 47-47^-, C. bipunctella; 48-48^, Aner-
astia lotella.
REVISION OF PEORIINAE AND ANERASTIINAE
107
Sc+ R|
49
51
Figures 49-51. — 49, Mania arizonella, dorsal view of adult cranium; 50, Arivaca pimella,
wing venation; 51, Peoria longipalfella, flattened male genitalia. jx=juxta, sp=spicate
process of uncus, tg=tegumen, mp^medial process of uncus, gn=gnathos, va^valva,
vin^vinculum.
108
U.S. NATIONAL MUSEUM BULLETIN 280
Figures 52-62.— Wing venation: 52, Arivaca pimella; 53, A. ostreella; 54, Atascosa glareo-
sella; 55, Peoria floridella; 56, Arivaca linella; 57, A. poohella; 58, Homosassa ella; 59,
Peoria approximella; 60, Arivaca alhicostella; 61, Peoria roseotinctella; 62, Ragonotia dotalis.
REVISION OF PEORIINAE AND ANERASTIINAE
109
Figures 63-82.— Wing venation and male antennae: 63, Martia arizonella; 64, Anerastia
lotella; 65, Coenochroa californiella; 66, C. bipunctella; 67, C. illihella; 68, Bandera binotella;
69, Tampa dimediatella; 70, Peoria longipalpella; 71, P. bipartitella; 72, P. tetradella; 73,
P. opacella; 74, P. floridella; 75, P. rostrella; 76, P. gemviatella; 77, P. roseotinctella; 78,
P. johnstoni; 79, P. santaritella; 80, P. holoponerella; 81, P. approximella; 82, P. luteicostella.
110
U.S. NATIONAL MUSEUM BULLETIN 2 80
Figures 83-104.— Male antennae: 83, Anacostia tribulella; 84, Arivaca fimella; 85, A.
linella; 86, A. ostreella; 87, A. poohella; 88, A. albidella; 89, A. artella; 90, A. alhicostella;
91, Atascosa glareosella; 92, Homosassa ella; 93, H. platella; 94, H. incudella, holotype;
95, Reynosa floscella; 96, Goya stictella; 97, Anerastia lotella; 98, Coenochroa californiella;
99, Bandera binotella; 100, fVakulla carneella; 101, Tampa dimediatella; 102, Barberia
affinitella; 103, Ragonotia dotalis; 104, Martia arizonella.
REVISION OF PEORIINAE AND ANERASTIINAE
111
106
Figures 105-112. — Male genitalia: 105, Peoria longipalpella; 106, P. bipartitella; 107,
P. tetradella; 108, P. opacella; 109, P.floridella, holotype; 110, P. rostrella, vesica everted;
111, P. gemmatella, vesica everted; 112, P. roseotinctella.
112
U.S. NATIONAL MUSEUM BULLETIN 2 80
116
118
120
Figures 113-120. — Male genitalia: 113, Peoria johnstoni; 114, P. santaritella; 115, P.
holoponerella, vesica everted; 116, P. approximella; 117, P. hileicostella; 118, Anacostia
trihulella, para type; 119, Arivaca pimella; 120, A. linella.
REVISION OF PEORIINAE AND ANERASTIINAE
113
123
126a
125
Figures 121-126. — Male genitalia: 121, Arivaca ostreella, vesica everted; 122, A. poohella,
paratype, vesica everted; 123, A. albidella; 124, A. artella, paratype; 125, A. alhicostella;
126, Atascosa glareosella; 126a, gnathos.
114
U.S. NATIONAL MUSEUM BULLETIN 2 80
127a
127b
130a
129a
I3la
I3lb
Figures 127-131.— Genitalia: 127, Homosassa ella, male; 127«, gnathos; 127^', gnathos
similar to that of lectotype; 128, H. platella, paratype male; 128a, gnathos; 129, H.
incudella, holotype male; 129a, gnathos; 130, Reynosa floscella, male; 130a, gnathos;
131, Goya stictella, male; 131a, transtilla; 131^, female genitalia.
REVISION OF PEORIIXAE AND ANERASTIINAE
115
I35a
Figures 132-135— Male genitalia: 132, Anerastia lotella, vesica everted; 133, Coenochroa
californiella; 133a, gnathos; 134, C. illibella; 135, C. hipunctella; 135a, gnathos.
116
U.S. NATIONAL MUSEUM BULLETIN 280
137a
138a
139
139 a
139b
139c
Figures 136-139.— Male genitalia: 136, B. binotella; 136a, tufts and sclerotizations of 8th
abdominal segment; 137, B. cupidineila; 137a, sclerotizations of 8th abdom. seg.; 138,
B. virginella; 138a, tufts and sclerotizations of 8th abdom. seg.; 139, Wakulla carneella;
139a, gnathos; 139^, elements of transtilla; I39c, tufts and sclerotizations of 8th abdom.
seg.
REVISION OF PEORIINAE AND ANERASTIINAE
117
I40a
140b
Figures 140-143. — Male genitalia: 140, Tampa dimediatella; 140a, gnathos; 140^, trans-
tilla; 141, Barberia affinitella; 142, Ragonoiia dotalis; 143, Martia arizonella.
118
U.S. NATIONAL MUSEUM BULLETIN 280
Figures 144-148.— Female genitalia: 144, Peoria longipalpella; 145, P. bipariitella;
146, P. tetradella; 147, P. opacella; 148, P. rostrella.
REVISION OF PEORIINAE AND ANERASTIINAE
119
Figures 149-153. — Female genitalia: 149, Peoria gemmatella; 150, P. roseotinctella; 151,
P. johnsioni; 152, P. santaritella; 153, P. holoponerella.
120
U.S. NATIONAL MUSEUM BULLETIN 2 80
Figures 154-159. — Female genitalia: 154, Peoria approximella; 155, P. luteicosiella; 156
Arivaca piviella; 157, A. linella; 158, A. oitreella; 159, A. poohella.
REVISION OF PEORIINAE AND ANERASTIINAE
121
Figures 160-165. — Female genitalia: 160, Arivaca albidella; 161, A. artella; 162, A.
albicostella; 163, Atascosa glareosella; 164, Homosassa ella; 165, H. incudella, paratype.
122
U.S. NATIONAL MUSEUM BULLETIN 280
Figures 166-170.— Female genitalia: 166, Reynosa floscella; 167, Jnerastia lotella; 168,
Coenochroa californiella; 169, C. illihella; 170, C, bipunctella.
REVISION OF PEORIINAE AND ANERASTIINAE
123
171
Figures 171-175— Female genitalia: 171, Bandera binolella; 172, B. cupidinella; 173,
B. virginella; 174, Wakulla carneella; 175, Tampa dimediatella.
124
U.S. NATIONAL MUSEUM BULLETIN 2 80
I'/v:^
k
176
Figures 176-178.— Female genitalia: 176, Barberia affinitella; 177, Ragonotia dotalis;
178, Martia arizonella.
U.S. GOVERNMENT PRINTING OFFICE; 1968
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