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CONTRIBUTIONS OF THE

ROYAL ONTARIO MUSEUM OF PALAEONTOLOGY

NO.4: DEVONIAN BRYOZOA FROM FORTIN AND
MALBAY TOWNSHIPS, GASPE COUNTY,
QUEBEC

BY Madeleine A. Fritz

APRIL, 1940

ana vir TaQ4 Moss ADSOYRSG WAIMOVEG
YTWUOD seas BRENWOT YAS Mo
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99 .A onivieboM Yaa

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Published” by auvhorirty™of the*Boardctor! Trustees
Royal Ontario hiuseum

Torontc, Canada

Price, ls: cents

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Tung sf) to vilnetisue va Sedetidu®

ferwauoraein® Laval

DEVONIAN BRYOZOA FROM FORTIN AND MALBAY TOWNSHIPS

GASPE COUNTY, QUEBEC

By Madeleine A. Fritz
Assistant Director of Invertebrate Palaeontology

INTRODUCTION

The bryozoan faunas of Gaspé have received very
little attention in the past. Aside fyom a few brief
references to these organisms by Clarke and a recent
paper by the writer” no attempt has been made to con-
sider these stratigraphically important fossils.

Fortunately the Quebec Bureau of Mines in their
recent surveys in Gaspé are making an effort to in-

clude in their fossil collections examples from ali’.

the bryozoa-bearing strata met with in the course of
their work. From the materiel which is being thus as-
sembled there is ample evidence of a prolific bryozoan
fauna although the preservation of the various forms
represented is, on the whole, not good. The rocks,
which consist chiefly of sandstones and crystalline
limestones, are so hard that in many instances it is
practically impossible to remove the specimens fram
their matrix or to make them available for study. Fur-
thermore, the sandstone species are represented for
the most part only by casts and molds. This condition
of preservation leads to considerable uncertainty with
regard to the determination of specific characteris-
tics. In spite of these difficulties, however, a dis-

tinctive fauna is being gradually recognized.

PRESENT INVESTIGATION

The material which forms the subject of this paper
was collected by Dr. H.W. McGerrigle of the Bureau of
Mines, Quebec. It was obtained from Fortin and Malbay
townships which are located towards the southeastern
part of Gaspé county, at the eastern end of Gaspé

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peninsula. Fossil localities represented include:

F7-M-59 Fortin Tp.; Range II; near head of small
brook one mihe west of the Fortin-Malbay
line. | ;

F9-M-3S9 Fortin Tp.; Range III; debris on top of

@ivide between Malbay river and the Big
Fork of the Malbay river; one-third mile
east of Fortin Centre line.

F13-M-S9 Fortin Tp.; Range VIII; close to R. VII-
VIII line; about two miles west of For-
tin-Malbay line.

Fl1-M-39 Fortin Tp.; Range VII, close to R. VII-
VIII line, about one mile and a quarter

| west of Fortin-Mal bay line... |
. F21B-M-39 Malbay Tp.; Range IV south; on Otter
(Pass) areens one-half mile north of Por-
« ottige rivers.
. F25-M-39 Malbay Tp.; Range IV, l, 000 feet west of
- Range III- -IV line on Portage river.

From the small collection obtained at these loca-
lities the following apa ria Ue. been pee 2eG

A nestom opora giegeee eh Fritz 1956
Anastomopora sp. indet
| Fenestrellina fFfortinensis De. SD.
Fenesfrellina gaspiensis Fritz 1938
Fistdiphragma jones! De. Sp.
Fisfolipora sp. indet
3 Lioclema Gepcnmg iy ns SP-.

DISCUSSION OF FAUNA

Of these above mentioned species Anostomopora gue-
- ecens/s and Fenesfre/lina gaspiensis are known only
from the Gaspé peninsula. The formerS was described
Originally from Four Mile creek, near Causapscal, Que-
bec, in the Matapedia valley of western Gaspé; the
latter* from the east branch of Sonneau brook, Galt
township, on the north side of York river valley in
eastern Gaspe. The strata from both of these locali-
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ties are considered to be of Gaspe sandstone age” The

finding of these two species in association in the -

Fortin-Malbay district assists in establishing rela-
tionships between Gaspe sandstone strata in several
areas throughout the peninsula.

An undetermined species of Arastomopora occurs at

locality Fel-M-59. It is represented only by a single -

cast which does not warrant definite specific determi-
nation. It bears a marked resemblance to the Gasped
sandstone species A. guebecensis but the horizon at
which it occurs is, according to McGerrigle, approxi-
mately one thousand feet (stratigraphically) down in
- the Grande Gréve limestone from the Gaspé sandstone
contact. The striking similarity between the form in
question and A. guebecensis might provide evidence to
substantiate the belief that the Grande Gréve lime-
stone and overlying Gaspe& sandstone are closer in age
than is generally admitted.
- ‘fwo other species ferestrellina fortinensis and
Fistuliphragma_jonesi represent new species from the
Grande Gréve limestone. Descriptions of these species
appear later in this paper. ferestrelling fortinensis
is abundant and well preserved at localities F13-M-39
and Fll-M-39. At the former locality it represents a
good horizon marker. fistuliphkragma jonesi is of rare
occurrence at locality Fl1-M-39. It is of particular
significance in this fauna in that the genus fistuli-
pkragma, which is typically developed in the Hamilton
formation of Michigan, has not been reported hitherto
from a point in North America as far east as Gaspe.
Occurring in association with Fistulipkragma jonesi is
an undetermined species of féstulipora. Though frag-
mentary remains of this species are not uncommon the
state of preservation permits only a generic refereme.
" The age of the remaining species Lioclema mcgerri -
glei , herein described from locality F25-M-39, is at-
tended with some difficulty. The Devonian representa-
tives of this genus are characteristic of the Helder-
‘erg and Hamilton formations. Since the strata in
which this species is found are definitely not Hamil-
ton I would incline to the belief that a Lower Devoni-
_ €@n horizon below the Grande Greve is here represented.

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DESCRIPTICN OF NEW SPECIES

Order Cyclostomata
Family Fistuliporidae
Genus Fistulephragma Bassler

According to Bassler® this genus differs from the
typical Féstudipora in that hemiphragms or semidia-
phragms are developed in the zooecial tubes.

Fistuliphragma jonesi n. sp.
Plate I, Figures 2,3,4,7.

General features.--This species is founded upon a
single fragment of a zoarium which forms a thin in-
erustation over a portion of crinoid column. In its
length the specimen measures 7 mm. The observed thick-
ness does not exceed 1 mm. Surface features are not
distinguishable since the specimen is longitudinally
fractured through the centre and reveals only a natu-
ral vertical section of the tubes. |

Tangential Section.--The zooecia are small elon-
gate-ovate structures. On an average four appear in
the space of ] m. They are separated by vesicular
tissue which about equals the width of a zooeciun.
Occasionally, however, broader areas of vesicular tis-
sue are seen which may constitute inconspicuous macu-
lae. A fairly well-marked lunarium is present, the
ends of which project into the zooecia.

Vertical Section .--The zooecia curve gradually
from their point of origin to the surface. From 2 to
8 rows of well rounded vesicles separate the zooecial
tubes which are, for the most part, filled with clay.
Hemiphragms are present in certain zooecia and are ab-
sent in others. Their absence may be due to poor pre-
servation or to the fact that they were not developed.
When present there are from two to four in each zooe-
cium. They appear on either side of the tubes and are
Situated at distances apart of over a tube diameter.

Remarks +-- Fistutiphragma jonesi differs from the
Other species included in this genus in its incrusting
habit of growth and small dimensions. The genotype

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F. spinulifera (Rominger) is a stout, monticulose,
branching form while F. seffordi. (winchell) is jueaia
than the present form and grows in lamellate layers.
Hololype.--Number 20374, Royal Ontario Musewn of
Palaeontology.
Occurrence.--Rare in Grande Gr2ve limestone, loca-
pre | F1l1-M-39.

Order Trepostomata
Fanily Batostomellidae
Genus / oclema Ulrich

errr ee ren meres te eh et eR ec AOR NI

| | _ Lioclema megerriglei n. sp.
erate J. Figures: 6.9).

General Features.--The zoarium is ramose. Only a
single branch is present in the collection. It is
round in cross section the diameter measuring 4 mn.
The length of. the branch is approximately 30 m. The
surface is not well preserved but would appear to be
smooth.

Tangential Section.--The zooecia are round to oval
in outline. They are separated by mumerous, rather
thick-walled mesopores of varying sizes. An average
of three zooecia appear in the space of one millimetre
measuring longitudinally. Acanthopores occur but they
are not. conspicuous and they do not tend to greatly
inflect the walls. Their presence is indicated merely
by a few fairly large mural granules. Certain of the
mesopores appear to be closed at the surface which fact
probably accounts for their varying sizes.

| Vertical Section.--The zonecia curve gradually
from the axial to the peripheral region. The walls,
which are thin, maintain a uniform thickness through-
Qqut their entire length. In the peripheral region an
Occasional diaphragm may be detected in the zooecia.
The numerous mesopores, however, contain diaphragms in
abundance, which gives to these structures the appear-
ance of regular vesicular tissue.

Remarks.--The available material jeee wat shod
very clear relationships between this form and any
Other previously described species of Lioclema. It

: 6

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elias sf? .nbsget fovedy}teq edt of Inlee ef mot
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has been thought desirabie, however, to record the
above features with the hope that further material
will be found that may permit a closer gat ae with
the known species.

Holotype. --Number 20375, Royal Ontario Museum of
Palaeontology. ;

Occurrence .--Rare in Lower Devonian(?), locality
F25-M-39. ce, Come

Order Cryptostomata
Family Fenestrellinidae
Genus Ferestrellina Bassler

Fenestrellina fortinensis n. sp.
Frate |, Figures 1,5,8<

General features.--The form of the zoarium sug-
gests a flabellate expansion but since identification
has been made from fragments only the exact size of
the colony is not known. The largest specimen observ-
ed measures 30 m. by 20 mm.

Qbverse.--Unfortunately only the reverse side is
exposed on the various specimens representing -this
species. The following characters of the obverse side
have been determined, however, by means of thin sec-
tions. The zooecia are round. They are arranged, as
a rule, in two rows though three rows may occur prior
to the bifurcation of a branch. A somewhat zigzag
median keel separates the rows. Three to four zooecia
occur in the length of a fenestrule which they tend to
slightly inflect. The dissepiments are stout struc-
tures. Three commonly appear in the space of 3 m.
measuring longitudinally. They are broadest at the
point of junction with a branch.

Reverse.--In the mature portion of the colony the
branches show a parallel arrangement. They are round
and fairly stout measuring 0.7 mm. in width an average
of six or seven apnearing in the space of 5 m. The
fenestrules are round to oval in the immature part of
the zoarium. They tend to elongate, however, as colo-
nial growth continues. Reaching maturity they become
‘quite irregular in size and shape ranging from a width

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a sotwents A .fenerd a To solssouySid edt.
eeniy twoh of somiT emer ade eeseyages. Loeat we Ls
of hoes yed? dota’ olutésamet » to dignel Ads ats
-tgues 2v0ds ews eiaueiqsueth off .topl tat ti
~om €° to goes: ud? cl tasqoa Yincrmoe south + Vagal
aot @e deeheosd sta yet? .¢identbutignol gd taoem
ia?) _domend « ditiw sottoryt to: Fag
eft wholoo add to nortsog etvtom edé a aphandedinis

bawos aha yedl .idewogrenas Lellioteg s wore ac
gat re ue cihie of «i 7.0 gatavesem fnpys eirist,
ant 6 to sooqs ond at gettneqqs anyes ‘to aie |

to seg ph Pil ea? ¢) Lev of Baset ete gels ¢
-oloy gn Ahvaved .singtote od Raed yamt .owkeser.
anonoe yard yt iawtem gtidsea .aeuntdeos dtworg o

of from 0.5 to 0.75 mm. to a length of from 0.75 mm.
to over 1.5 m.

One of the most striking features of the reverse
. side is the presence on the brenches of numerous fine,
well-marked, irregularly disposed tubercles of varying
sizes. These markings give to the surface of the
branch a distinctly granular appearance. The dissepi-
ments, which are strongly developed structures, are
similarly ornamented.

Remarks ---F fortinensis is the most characteris-
tic svecies at locality F13-M-39 where it is abundant.
It occurs, also, at locality Fl1-M-39. This form may
be distinguished from all the other Gaspe species thus
far studied by the finely tuberculated reverse surface.
This feature is not unique among the fenestrellinids
but the svecies in which this particular characteris-
tic has been observed differ from the present form in
other essential details. |

Cotypes.--Number 20373, Royal Ontario Museum of
Palaeontology.

Occurrence ---Grande Gréve Limestone. Common at
locality F13-M-39.

SUMMARY

_ From the foregoing study of a small collection of
bryozoa obtained by H.W. McGerrigle of the Quebec Bu-
reau of Mines from Fortin and Malbay townshins in
Gaspe’ county, in the southeastern part of Gaspe penin-
sula, three new species f¢stuliphkragma_jonesi, Liocle-
ma mc errig leis and Fenestrellina fortirnenrsis have
been described. In addition, the investigation has
provided evidence which may assist in correlating
Gaspe sandstone strata in several areas throughout the
peninsula.

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~s\aor \ ,s2arc\ eenewcjiete antoegs wan ood? .alon
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sad aolingtiesvat st Reaeere ai ebudiieeeh aeed
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= mh

REFERENCES

Ciarke, J.ti., EVCS; NeY.S.M., Mem. 9, pt. I, pp.
215-242.

Britz, «hs, 1956; Bulls. Amer, Pal., vok. 24, App.
82A, pp. 1-14.

Brite. M.A. ; 1958; Bul MepeAmerer Mal 4p Velswes, App.
S2a, pa GS, pi. L, fice 5.

Peita, M.A.BiaAg9S6; Bodie.’ Ame tv, Pal) Mods! 24, App.
BZA 2 ps) Tea pliod, fess: i5,4.

Bindie, E.M., 1956;Bullis. Amer. Pals, yol. 24, p.4l.

Jones, I.W., 1957; York River Area; Quebec Bureau
of Mimes, p. 22.

Kindle, E.M., 1958;Bulls. Amer. Pal.,vol. 24, p.44.

Bassler, R.S., 1934; Jour. Wash. Acad. Sci., vol.
24, no. 9, p.- 407. |

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EXPLANATION OF PLATES

Fig. l. Fenestrellina fortinensis n.- Sip. Tangen-
tial. secrion, ‘cel luliferous, side ;. X35:

Figs ee Fistalipkragma joresé nh. sp. Tangentiai
section showing maculum and zooecia with lunaria; X35.

Fig. 3. Fistulipkragma jonesi nu. sp. Vertical
section showing hemiphragms and vesicular tissue; X25.

Fig. 4. Fistulipkragma jonesi 2. sp. Vertical
section showing hemivhragms and vesicular tissue; X25.

Fig. 5. Aerestrellinag fortinensis un. sp. MTangen-
tial section, non-celluliferous side showing granules;
X55.

Fig. 6. Léoclema megerriglel n. sp. Vertical sec-
tion showing a zooecium with few diaphragms also
mesopores with numerous irregular diaphragms; X35.

Fig. 7. Fistulipkragma 1) pee ne sp. Vertical
section; X50.

Fig. 8. fenestrellina fortinensis n. sp. Fragment
of zoarium, non-celluliferous side; X. |

Fig. 9. Lioclema megerriglet n. sp. Tangential

section showing zooecia,irregular mesopores, and acan-
thopores; X35.

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