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FLORA 
MALESIANA 


SERIES I - SPERMATOPHYTA 


Flowering Plants 
Vol. 6, part 6 


INDEX TO REVISED FAMILIES 


Aceracede,: +. 4: 3,592 Elatinaceae 4: 203 Pentaphragmataceae. 4: 517 
Actinidiaceae s.str. . 4: 37  lEpacridaceae . 6: 422 Pentaphylacaceae. 53-121 
Aizoaceae ees ia 207n subricaceae. : 6: 469  Philydraceae 4: 5 
Alismataceae . . . 5:317 + #Erythroxylaceae 5: 543 Phytolaccaceae 4: 229 
Amaranthaceae . 4: 69,593  Ficoidaceae 4: 267 __Pittosporaceae 5: 345 
Ancistrocladaceae . 4: 8 _ Flacourtiaceae 5: 1  Plumbaginaceae 4: 107 
Aponogetonaceae 4: 11, 7: 213 Flagellariaceae 4: 245  Podostemaceae te G5 
Basellaceae - 5:300 Geraniaceae 6: 445, Polemoniaceae . . 4: 195 
Batidaceae 5: 414 Gnetaceae. 4: 336 Pontederiaceae aS oS 
Betulaceae 5: 207  Gonystylaceae 4: 349 Portulacaceae TWA 
Bixaceae s. str. 4: 239 Goodeniaceae 5: 335  Primulacea 6: 173 
Burmanniaceae . 4: 13,592 Haemodoraceae 5: 111 Proteaceae 5: 147 
Burseraceae 5: 209 Haloragaceae 7: 239 Punicaceae 4: 226 
Butomaceae 5: 118 Hamamelidaceae . 5: 363 Restionaceae . 5: 416 
Byblidaceae 7: 135 Hippocrateaceae . 6: 389 Rhizophoraceae i TS ae 
Callitrichaceae 4: 251 Hydrocaryaceae 4: 43 - Salicaceac,. (..°) Sieceesemn 
Campanulaceae 6: 107  Hydrocharitaceae. 5: 381 Salvadoraceae. «.) eaaaees 
Cannabinaceae 4: 223 Hydrophyllaceae . 4: 207 Sarcospermaceae. . 4: 32 
Capparidaceae 6: 61  Icacinaceae 7: 21 Saururaceae :..’ sean 
Caprifoliaceae . 4: 175,598  Juglandaceae . 6: 143 Scyphostegiaceae. . 5: 297 
Cardiopteridaceae 7: 93  Juncaceae . 4:210 Simaroubaceae . . 6: 193 
Celastraceae . . 6: 227, 389  #Juncaginaceae. 4: 57  Sonneratiaceae . 4: 280, 513 
Centrolepidaceae §: 421 Lemnaceae 7: 219 Sparganiaceae <) (eases 
Ceratophyllaceae 4: 41 Loganiaceae 6: 293°. Sphenocleaceae’ | eae, 
Chenopodiaceae . 4: 99,594 Lophopyxidaceae 7: 89 Stackhousiaceae < )-yAewa 
Clethraceae 7: 139 Malpighiaceae 5: 125 Staphyleaceae « “EL Geeeas 
Cochlospermaceae 4: 61 Martyniaceae . 4: 216, Stylidiaceae’.. 2 ieee 
Combretaceae 4: 533 Molluginaceae 4: 267 | Styracacede . -. -eaaaeeas 
Connaraceae . . . 5:495 Moringaceae . 4: 45 Thymelaeaceae 4: 349, 6: 1 
Convolvulaceae . 4: 338,599 Myoporaceae . 4: 265. -Trigonidceae.. 4. eee 
Corynocarpaceae 4: 262 Myricaceae 4: 277 -’Turneraceae .° 3 Seer 
Crassulaceae . 4: 197 Najadaceae 6: 157 Typhaceae . eA ae SSSA 
Datiscaceae 4: 382 Nyctaginaceae : 62450 # Umbelliferae .  ~ 41133595 
Dichapetalaceae . 5: 305... Nyssaceae.... . |... . y42029- ‘Walerianaceae’:) 2 aeeenaeecee 
Dilleniaceae 4: 141 Ochnaceae 1) Meee Pa eba Oat AVG laceae ids Sen coe 
Dioscoreaceae 4° 293 « Oxalidaceae wo” Sm feasts 151 Xyridaceae . . 4: 366, 598 
Dipsacaceae 4: 290 Papaveraceae. . . 5:114 # Zygophyllaceae, . . 4: 64 
Droseraceae A: 37) -Pedaliaceae 2 0-0, ee 42 216 


TAXONOMICAL REVISIONS 


REPUBLIK INDONESIA 
REPUBLIC OF INDONESIA 
LEMBAGA ILMU PENGETAHUAN INDONESIA (L.LP.L.) / 
INDONESIAN INSTITUTE OF SCIENCES 


FLORA MALESIANA 


BEING 
AN ILLUSTRATED SYSTEMATIC ACCOUNT OF THE MALESIAN FLORA | 
INCLUDING KEYS FORDETERMINATION | DIAGNOSTIC DESCRIPTIONS | 
REFERENCES TO THE LITERATURE|SYNONYMY]/AND DISTRIBUTION | 
AND NOTES ON THE 'ECOLOGY OF 
ITS WILD AND COMMONLY CULTIVATED PLANTS 


PUBLISHED 
UNDER THE AUSPICES OF LEMBAGA BIOLOGI NASIONAL 
BOTANIC GARDENS OF INDONESIA / BOGOR / JAVA AND 
OF THE RIJSKSHERBARIUM / LEYDEN / NETHERLANDS 


PREPARED 
ON AN INTERNATIONAL CO-OPERATIVE BASIS UNDER THE SUPERVISION OF 
SEVERAL DIRECTORS OF BOTANIC GARDENS / KEEPERS OF HERBARIA 
AND VARIOUS PROMINENT BOTANISTS 


FOR: THE PROMOTION. OF 
BOTANICAL SCIENCE AND THE CULTURAL ADVANCEMENT OF 
THE PEOPLES OF SOUTH-EASTERN ASIA TO 
THE SOUTHWEST PACIFIC REGION 


SERIES I 


VOLUME 6 
SPERMATOPHYTA 


GENERAL EDITOR: 


Dr CrG. G. IVAN STEENIS 
DIRECTOR OF THE FOUNDATION ‘FLORA MALESIANA’ 


PUBLISHED BY 
WOLTERS-NOORDHOFF PUBLISHING, GRONINGEN, 
THE NETHERLANDS 
PRINTED IN THE NETHERLANDS 
1960-1972 


COPYRIGHT 1972 


All rights reserved, including the right to reproduce 
this book or parts thereof in any form 


ISBN 9001 31822 3 


Printed in the Netherlands 


Title-page 
Contents . 
Dedication 


Abbreviations and signs. 


CONTENTS 


TAXONOMICAL REVISIONS 


in alphabetical sequence 


Campoanulaceae by B. Moeliono & P. Tuyn . 


Capparidaceae by M. Jacobs . 
Celastraceae-I by Ding Hou . 
Celastraceae-II by Ding Hou . 
Epacridaceae by H. Sleumer . 
Ericaceae by H. Sleumer 

Geraniaceae by R. C. Carolin. 
Juglandaceae by M. Jacobs 
Loganiaceae by P. W. Leenhouts . 
Najadaceae by W. J. J. O. de Wilde . 
Nyctaginaceae by J. F. Stemmerik 
Primulaceae by P. A. J. Bentvelzen 
Simaroubaceae by H. P. Nooteboom. 


Staphyleaceae by B. L. van der Linden . 


Thymelaeaceae by Ding Hou . 


Addenda, corrigenda, et emendanda by C. G. G. J. van Steenis and collaborators . 


ADDENDA 
to volumes 4, 5 and 6 


INDEX 


Index to scientific plant names by M. J. van Steenis-Kruseman 


Page 
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(5) 
(7) 

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915 


985 


Dedicated to the memory of 
ELMER DREW MERRILL 


DEDICATION 


The completion of the sixth volume of this Flora gives me the privilege to dedicate this to the 
memory of ELMER DREW MERRILL, a man who has achieved more for the knowledge of the 
Malesian flora than any other individual botanist. 

It is neither my intention to give nor is it the proper place for a full biography of this most 
distinguished American scientist, as it would for the greater part be duplication of his own 
‘Autobiographical’ (1953), the scholarly essay by RossBins (1958), and the vivid life sketch by 
ScCHULTES (1957), which together give the story of his life, his ambitions, his personality, his 
immense drive, his multiple interests, his capacity for establishing botanical periodicals as well as 
successfully filling the posts of Dean of a Faculty of Agriculture, director of the Bureau of 
Science at Manila, director of the New York Botanical Gardens, and administrator of Botanical 
Collections of Harvard University. 

It is my purpose to review MERRILL’s aims and vision, ambitions and achievements in the 
light of his time, to explain the value of his pioneer works for Indo-Malesian botany, how he used 
opportunities and had to bow to unforeseen events and circumstances which in no mean way 
influenced his career. Naturally MERRILL’s personality pervades the story, that of a straight- 
forward, righteous person, unbiassed in scientific matters, appreciating any progress in bio- 
logical science. It is of course especially his great achievements with regard to the knowledge of 
the Malesian flora which are the main theme and I will try to elucidate several aspects which he 
pursued. 

A glance through his immense bibliography, containing some 550 entries, among which ten 
very large books, reveals his fantastic productivity, largely centered on the flora of Malesia, 
East Asia and the Pacific. 

Scanning the herbarium one becomes aware of the fact that during his lifetime he must have 
pre-identified, named or definitely identified over half a million specimens from the East, 
including the Philippines, Sumatra, Borneo, Amboyna, New Guinea, Melanesia, Micronesia, 
southern China (including Hainan), Indo-China and Burma, made possible by his unequalled 
knowledge of forms, his cast-iron memory and his zealous devotion. 

It is with awe that one observes such a great achievement and then one wonders how one man 
could find time to do all this. The answer is given by SCHULTES who wrote that ‘his reaction to 
added work was to lengthen the day’. 

As Rossins wrote, ‘he was a man in a hurry who saw clearly a program of research which 
absorbed him, but which was greater than any individual could complete within the limits of a 
single life-time. The demands of this program possessed him. It led him to make various in- 
novations and modifications of convential procedures in herbarium methods with increased 
efficiency in the use of this, for him, essential tool and to advocate briefer citations, one-name 
periodicals, and other means of economizing time and effort. It induced him, at least in part, 
to make quick decisions without long considerations of pros and cons and to act at times without 
regard to the feelings of others.’ 

‘In spite of his absorption with his speciality MERRILL was not a recluse. Nothing pleased him 
more than to light his pipe and sit down to talk with a group of gardeners or with a student or 
his colleagues, mainly, of course, about plants. He played an active part in many organizations 
and valued the associations he made in them. He joined the Masonic order and eventually 
became a thirty-third degree Mason. He enjoyed an evening cocktail in his later years, and loved 
to have visitors and dinner guests. Though he had no hobbies outside his profession, he was 
interested in sports, especially baseball, football, and tennis. The ‘Autobiographical’ accounts of 
his adventures in collecting in the Philippines and in China reveal some of the human aspects of 
his character.’ 


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FLORA MALESIANA 


Such human aspects became evident to us very shortly after the war as he sent food parcels 
to his German colleagues and distributed amongst other botanical material on loan or for gift to 
colleagues in other freed countries envelopes with pounds of tobacco labelled with dry humour 
Nicotiana tabacum L. in parcels marked as ‘botanical specimens for scientific study only’. 

‘His energy was boundless. It was his habit, at the University of California, to arrive at the 
herbarium at 6:00 a.m. and work on shipments of plants until he left for the Dean’s office at 
9:00 a.m.; he continued at noon after office hours, or on sundays and holidays’ (RosBINs, 1958). 

As will appear later, his drive was probably born from the challenge he had to face in the 
Philippines in his early days and which activated his innate energy: it became a life-long habit. 

MERRILL was born in East Auburn, Maine, October 15, 1876. His parents were of limited 
means and belonged to the industrious people of New England. One of his ancestors was 
NATHANIEL MERRILL who immigrated in 1635 and was of Huguenot descent (originally DE 
MERLE). In his family there was also English and Scottish blood, a ‘melting pot’ feature not 
unusual in the United States. As a young boy he was interested in natural history, in birds, rocks, 
minerals, fungi, and local woods; before reaching highschool age he became interested in col- 
lecting and naming plants, but he went in for engineering in 1894 in Maine State College at 
Orono which in his senior year became the University of Maine. He took the general science 
course, but remained interested in biological work, particularly in the classification of plants 
and worked under Prof. F. L. HARVeEy especially on cryptogams; he attended, however, only two 
semesters on botany. His private herbarium contained some 2000 specimens. In 1898 he got 
his B.Sc. and became assistant in natural history. While working for his M.Sc. he accepted a 
post as assistant-agrostologist in the U.S. Department of Agriculture, Washington, as assistant 
to F. LAMSON-SCRIBNER, then the leading authority on North American grasses, composing 
amongst others a Manual of the Grasses of Alaska. 

In 1902, following the establishment of the sovereignty of the United States over the Phil- 
ippines, his chief accepted a post as director of the newly established Department of Agriculture 
in Manila. LAMSON-SCRIBNER was Obviously so very satisfied with MERRILL’s person and capacity 
that he asked him to become botanist in his department. MERRILL, however, was completely 
satisfied with being an agrostologist in his own country and refused twice, arguing that he knew 
nothing about tropical plants from the East. Finally ScRIBNER convinced him that ‘nobody in 
the United States knew anything about the Philippine flora and that he had as good a chance 
as any one’. He accepted on Monday afternoon, February 20, 1902, and had to agree to be 
ready to sail from New York at 1:00 p.m. February 22! He nearly missed the boat because of a 
heavy sleet storm delaying the train from Washington D.C. to New York, but he made it. He, 
least of all, could not anticipate that he would remain for twenty-two years in the Philippines. 
Slightly over two months later he reached Manila where the new personnel had to start work ina 
vacant building, ‘without a chair or table, much less a botanical publication or a botanical 
specimen’. 

With a huge program before him MERRILL started energetically: one month after arrival he 
made a six weeks trip, partly under military guard, and during his first years he spent approxi- 
mately one half of his time in the field; in the next twenty-two years he explored in almost all 
parts of the extensive archipelago. 

In September 1902 he paid a two-months visit to the Herbarium Bogoriense, taking with him 
his botanical specimens, for no trustworthy work could be done in Manila in the complete 
absence of authentically named material. Here he became acquainted with the literature of 
Malesia of which he acquired an excellent working knowledge. He himself said that it was of 
infinite value for building up the botanical library at Manila. He wrote also an extensive report 


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Dedication 


on this visit explaining the methodologies of collecting, forest plotting, numbering of trees in 
forest reserves, herbarium methods and techniques, efc. used in Java, which served as a model for 
the work in the Philippines. 

At that time the responsibility for taxonomical research work in the whole of Malesia vir- 
tually hinged on two systematists, RIDLEY in Malaya and VALETON at Bogor who collaborated 
with Koorpers, the organiser of the collecting; to these MERRILL was now added as a third. 

In comparison with Malaya and Java, work in the Philippines had the great disadvantage 
of having a much less solid basis; this consisted only of BLANCO’s, FERNANDEZ-VILLAR’s and 
VIDAL’s early works, and at Manila there were no voucher specimens of these authors’ collec- 
tions, nor of the vital collections of CUMING and HAENKE for comparison. 

This vacuum led him to realize that he had to start from scratch and had to build up a col- 
lection and library, to interpret plants from older works, enabling the description of new plants 
and accounts of collections, that it would be compulsory to ask assistance from foreign botanical 
specialists in various groups, that he had to stimulate interest in the botany of the Philippines by 
distributing duplicates on a large scale and by bringing Philippine plants into ‘circulation’, and 
that he had to bring into being a publication medium for scientific results and for the correlation 
of Philippine botany with that of adjacent countries. 

This clear vision of the items of the enormous amount of pioneer work ahead would have 
discouraged any average scientist, but to MERRILL it meant a great challenge, to create a centre 
from scratch. 

It became almost a one man show. In the course of twenty years he had never a permanent 
collaborator, except two honorary collaborators, viz E. B. COPELAND for the Pteridophytes and 
Oakes Ames for the Orchidaceae. Another honorary American collaborator, Miss JANET PER- 
KINS, started a series of publications under the name ‘Fragmenta Florae Philippinae’; she 
settled at Berlin where she worked in conjunction with OTro WARBURG who had made large 
Philippine collections himself. These collections, together with those of MERRILL and Capt. 
AHERN formed her basic material, but after 3 fascicles (1904-1905) this promising series was 
abandoned. Another American botanist who later joined the Bureau of Science at Manila and 
was employed for three years (1908-1911) was CHARLES BUDD ROBINSON, a critical, promising 
botanist. He returned in 1912 for another period, but unfortunately met a premature death in 
1913 on the island of Amboyna while re-collecting Rumphian plants. 

MERRILL of course worked in close collaboration with the members of the staff of the Forestry 
Institute. They were mostly collectors describing forest composition and timbers, such as H. N. 
WHITFORD, H. M. CurrAn, and others. An exception was F. W. Foxwortuy who actually 
served for some years as botanist at the Bureau of Science and who took care of a revision of 
Philippine Dipterocarpaceae, the most important timber family. 

An American botanist of great impact on the development of Philippine botany was ADOLPH 
DANIEL EDWARD ELMER (1870-1942). He was originally in the employ of the Bureau of Science in 
Manila (1904-1905), but settled later as a professional free-lance plant collector and collected 
over 20.000 numbers in the Philippines in a large number of sets. ELMER published lavishly in a 
series of his own, the ‘Leaflets of Philippine Botany’, ten volumes, together covering 3936 pages 
in print, among which descriptions of over 1500 new species. What MERRILL’s relation to ELMER 
was and whether he appreciated him or not are not clear, as he does not mention him in his 
‘Autobiographical’. At the time of his stay in Leyden, where I had the privilege to have him as 
our guest, Oct-Nov. 1951, I omitted to unearth this. I can hardly believe that ELMER’s super- 
ficial uncorrelated descriptions can have been very welcome. In Ficus MERRILL reduced 31 of the 
70 described species by ELMER before 1923 and of the total of 85 novelties of ELMER’s Ficus only 


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FLORA MALESIANA 


13 are still accepted in CORNER’s monograph. MERRILL had to tolerate ELMER’s work because 
the latter was an entirely independent man whose virtue lies mainly in his profuse collecting. 

Why MerriLL never had a larger permanent staff of at least 2-3 qualified botanists is another 
question left unanswered. He could easily have claimed staff for the Phanerogams as besides his 
collecting work, he had curatorial and organisational tasks. Since 1912 he also had an associate 
professorship in the University of the Philippines and served as head of the Department on a 
half-time basis and without additional compensation (sic), “which until 1918 seriously interfered 
with productive work in systematical botany’. To make botany popular and to frame a textbook 
of systematic botany for both residents and students, he had already prepared a ‘Flora of Manila’ 
(1912), still a very useful book, covering some thousand species. During the academic year his 
teaching duties never occupied less than eighteen hours per week, and during certain semesters 
even thirty-six. It is amusing to read in his ‘Autobiographical’ how he got rid of it, early in 1919, 
‘his full time being for the first time in many years available for what he most desired to do’. 
But after two weeks he was appointed as director of the Bureau of Science, a post neither sollicited 
nor desired, but which he could not refuse under the circumstances, it being an order. 

In passing I remark that of the Filipino students he taught not one specialized in plant system- 
atics as far as I know. Those with a biological tendency probably all went in for more applied 
branches, such as pharmacy, entomology, agriculture, forestry, fisheries, etc., so that when MER- 
RILL left the Bureau of Science in 1923 he had no immediate successor as botanist. It was five 
years later when Dr. E. QUISUMBING was engaged as such. He had received his primary education 
at the College of Agriculture, University of the Philippines and at Chicago University. He worked 
together with MERRILL, then Dean of the Faculty of Agriculture, University of California at 
Berkeley, for two years, 1926-1928, from which a joint paper “New Philippine Plants’ emanated; 
from this it appears also that MERRILL was quite capable of making botanical drawings. The 
main thing was that MERRILL induced him in these two years to start revisional work for the 
initiation of a ‘Flora of the Philippines’ and along this line QuUISUMBING elaborated the first 
sample, Philippine Piperaceae, while at Berkeley, under MERRILL’s supervision, as a final coach- 
ing and a start towards this new goal. 

Notwithstanding the time-consuming handicaps connected with education and administration 
MERRILL poured all his available energy into the botanical aims set forth above. He had by 
necessity been induced to take on administrative duties in the Philippines and had shown his 
capacity to meet this challenge. This experience served him later enabling him to fulfill other 
administrative duties in California, New York and Harvard in a successful way. That he could 
pursue his own botanical work simultaneously in free time and leisure hours is due to the fact 
that his heart was in botany and that he regarded work in plant systematics and floristics as his 
‘safety valve’ ‘when he could immediately forget his administrative problems’. The taking of 
vacations was rather foreign to him and in the entire period in the Philippines, from 1902 to 
1923, he only took vacation in 1905, while in other years he used his annual month’s leave for 
making collecting trips, mainly in China. 

He built up a collection, personally, with various famous skilled Filipino collectors (RAMos, 
EDANO, SULIT and many others) from all parts of the Philippines (the Bureau of Science = B.S. 
series), acquiring huge collections from private collectors (ELMER, Mrs. CLEMENS, WENZEL, 
LOHER, VANOVERBERGH, efc.) and from the Forestry Service officers (the F.B. series). Besides 
this he acquired large collections from surrounding regions, Guam, China, Indo-China, Amboy- 
na, and in addition very large sets of duplicates in exchange for the material which he widely 
distributed ‘on a free exchange basis’, that is, the liberal but in the long run profitable principle 
of sending out duplicates as many as each institution has available, thus not on a precise | : 1 


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Dedication 


specimen basis. He estimated that in 1922 he had sent out more than 500.000 duplicates, the 
reference collection at Manila amounting to c. 275.000 mounted specimens, containing repre- 
sentatives of practically all Philippine species, in the form of types, isotypes, fragments of 
types, material critically compared with originals, photographs of types and sketches. 

The library, similarly built up from scratch, was after a decade admittedly ranked on the same 
level with those at Bogor and Calcutta and far larger than the one at Singapore. 

He had also to provide for a publication medium and was instrumental in founding the 
Philippine Journal of Science, Botany Supplement, which was filled mostly by his own contri- 
butions but also served for revisions prepared by his associates, Philippine and foreign. 

Interpreting the older works he found a great necessity, both for botanical and nomenclatural 
reasons. He started with the work of BLANco, for which purpose he had special collections made 
in Blanco’s classical areas, from which emanated his ‘Species Blancoanae’ (1918), followed by a 
similar key work on ‘Rumphius’ Herbarium Amboinense’ (1917), both unrivalled for their 
critical standard. He tried to correlate both interpretations with material from the locus classicus. 
Only first-class botanists with a very wide knowledge of plants are capable of composing such 
works. In 1921 he published an evaluation of BURMAN’s ‘Flora Indica’, but this was not sustained 
by a study of the types at Geneva; unfortunately BURMAN’s herbarium is dispersed through the 
general Herbarium by HocHREUTINER. For his “Commentary on Loureiro’s Flora Cochinchi- 
nensis’ (1935) he had at his disposal earlier collections made by Mrs. CLEMENs in the locus 
classicus near Hué; he also tried to locate types of LouUREIRO in the Herbarium of the British 
Museum, London. At the end of his life he was working on a very large work on the location and 
evaluation of the RoxBURGH plants, the MSS for which are left unfinished (a complete copy has 
been deposited in the Arnold Arboretum, a less complete one at Leyden). We would have 
preferred that he had started this work earlier instead of his immense evaluation of the works of 
RAFINESQUE which occupied him for several years onwards of 1940. Admittedly the location and 
evaluation of the ROXBURGH types could only be performed by prolonged visits to European 
herbaria, which were impossible during the war and which, moreover, MERRILL could not make 
while he was Administrator of the Harvard Collections. All these works are extremely important 
for typification and nomenclature, often of very common tropical plants as many errors or 
omissions were made in their former interpretation, if indeed any was ever made. 

MERRILL contributed lavishly to describing new species — some 4000, of which 3000 from the 
Philippines — and several new genera of plants. They were partly published in a series of New 
and Noteworthy Philippine Plants (18 numbers, 1904-1922), partly in accounts of particular 
collections from certain islands, districts or mountains. It is quite certain that only a fraction of 
these will stand the test of time. To understand this we must consider his vacuum position and 
the dilemma with which he was faced, either to do critical-botanical work or to do the best he 
could in determining plants by reading descriptions, comparisons with available material and if 
nothing fitted to describe the plant as new. Critical-botanical work of course goes slowly, at a 
rate of some 20-100 accepted species a year depending on the group and the botanist’s capacities 
and zeal; it also requires that one has access to a large library and can borrow types and authentic 
material. The absence of the latter facility is the greatest handicap for monographic work in the 
tropics, as the largely European-centred ‘type herbaria’ are not prepared, and rightly so, to make 
large loans to the tropics, an affair which is too risky in several respects. This necessitates that 
workers in the tropics, after having prepared preliminary MSS, must pay prolonged visits to 
these institutions for checking type and authentic material and establish identities and names. 
And although MERRILL gradually assembled at Manila a huge herbarium and photographs or 
fragments of types from European herbaria, and attached figures, descriptions, notes, pencil 


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FLORA MALESIANA 


sketches, carbon leaf impressions, etc. to the sheets, framing in this way a new tool of informa- 
tion, a sort of combined plant file and library, he had little of this during the first decade of his 
work at Manila. At that time he had to be content to study descriptions, an art which he fully 
mastered, an art which is gradually becoming extinct among contemporary botanists who adhere 
to the examination of type specimens. Moreover, the incoming Philippine material was over- 
whelming, all completely unnamed. To cope with this he asked the co-operation of all available 
specialists in the world to elaborate certain groups. The bulk he had to name himself and this 
could be done only in a superficial way, thus it amounted to mere floristic name-giving and was 
not truly systematic, comparative, either regional or semi-monographic. Besides all the already 
mentioned handicaps to work of lasting value, the Philippine flora offers an additional com- 
plication in its specific variability, which is in part certainly a consequence of the archipelagic 
conditions due to fragmentation of once continuous populations before the block-sinking 
disrupted these in past geologic time. With scanty material a botanist will under such circum- 
stances be inclined to describe more species than there really are. It must be admitted, however, 
that the specialists to whom he entrusted the work on as many groups as possible were faced by 
exactly the same difficulty and their work suffers from the same evil. By working on this level 
MERRILL certainly created problems for others to solve. I have not scanned families on that 
point, but the six sedges he described as new are now all in the synonymy. 

He himself was of course fully aware that there were far less species in the Philippine flora than 
listed at the time of his enumeration (1923), and being honest and wise to a high degree, he 
frankly admitted during a discussion on the virtue of different policies for unravelling the botan- 
ical treasures of Malesia, that ‘many of my new species, and even new genera, were optimistically 
proposed’, adding that he had seen no other way to create a botanical basis. 

Of course nowadays, half a century later, ‘collection description’ is an obsolete procedure, 
detested as an inferior sort of botany creating unnecessary duplication and spending valuable 
time in a useless way. Nowadays it is clear that thorough work is needed and this can be done 
because in general collections have accumulated sufficiently to provide a solid basis, which we 
hope is about as good as, or slightly better, than the basis collections which were used for the 
compilation of the ‘Flora of British India’. Fortunately collections are becoming more ample 
every year due to enthousiasts in Malaya, Sarawak, Sabah, the Philippines, and Lae, to whom 
we cannot be thankful enough. In MERRILL’s ‘period’ the time was simply not ripe for launching 
comprehensive botanical works. To have a fair judgement of his pioneer work the commentary 
just given should be considered. 

Several facets of his work were intended to cope with the urgencies of the ‘vacuum position’, 
e.g. the building up of collections, the library, a publication medium, the nomenclatural evalua- 
tion of old works, the description of supposed novelties, the framing of enumerations, as pre- 
cursors to Floras, and bibliographies. Having a vast knowledge of the literature he was in the 
latter extremely successful, in accuracy and completeness. His enumerations of the floras of 
Borneo, Hainan, Banguey I., Guam, and some others are basic for later work, especially that 
of Borneo. 

After the passing of Taft’s Law (1917) it was American policy to prepare the Philippines for 
independence and MERRILL foresaw that his future career would not allow him to compile his 
ultimate goal, a ‘Flora of the Philippines’ for which his ‘Flora of Manila’ had served as a model — 
actually in 1922 although he was director of the Bureau of Science he was on a year-contract, 
without pension and could be dismissed with a year’s salary as bonus (sic) — so he prepared his 
‘Enumeration of Philippine Flowering Plants’, to synthesize what was achieved and leave a basis. 
This is still a most useful work, without which we would be much the poorer. The introductory 


(12) 


Dedication 


essays are a masterpiece of work, the enumeration is complete, and the bibliography is still a 
reliable source of information. Of equal value are his bibliographies of Borneo, the Pacific, and 
East Asia. 

MERRILL’s ability of mastering and analyzing literature entailed of course changes in name- 
giving according to the principle of priority and typification. As a good botanist of course he 
favoured stability of nomenclature, which cannot always be realized, however, particularly not 
at species level. He listed especially, in his works on LouREIRO and RAFINESQUE, the cases where 
conservation of generic names was necessary. 

A corollary of his bibliographic interest was establishing new journals; he was instrumental in 
founding the ‘Philippine Journal of Science’ and many others, later pointing out the virtue of 
one-name periodicals of which he founded several, a procedure later widely accepted (Blumea, 
Willdenowia, Persoonia, Adansonia, e/c.). 

Shortly after the war he also started a promising offset reprinting of rare early works, the 
desirability of which probably emanated from his study of the rare works of RAFINESQUE. 
He told me that with a subscription of 150 copies this could be done without loss at a very low 
cost on a non-profit basis. It is a pity that this was not continued by somebody for the benefit of 
botany, as there was and is a need for such inexpensive reprints. Before the war I had already 
tried to raise the interest of Chronica Botanica for such a purpose and later made similar pro- 
posals to the I.A.P.T., but found no sympathy for the idea. The result is that large firms 
nowadays produce such offset prints at prohibitive prices on a profit basis, a most undesirable 
situation for botany in general and the Philippines especially as the reprints of the ‘Flora Manila’ 
and the ‘Enumeration’ fall entirely beyond the financial capacity of most Philippine residents 
for whom they are intended. That a low-price large sale is still possible is apparent from the 
excellent offset reprinting in Formosa for educational purposes. 

Another corollary which proved to be of great scientific interest was his careful recording of 
vernacular plant names, an intricate affair because of the many dozens of native languages in the 
Philippines. He examined these names together with linguists and found that several of them 
were corrupted Spanish names; thus he contributed to both linguistic and botanic science. 
He found also that many early plant introductions had never spread and still retained station in 
the coastal hinterland of Cagayan Bay where the Spanish galleons had anchored. This, combined 
with the vernacular names, brought him to a consideration of early post-Columbian transtropical 
transport of cultivated plants and aliens. In this, he found that a clear distinction must be made 
between the galleon routes of the Portuguese from Brazil to Goa and Macao and that of the 
Spanish from Acapulco in West Mexico to Manila via Guam. Both galleon routes proved very 
important indeed, as they persisted for centuries. In this way he was able to unearth the origin 
of a number of plants on which there was no unanimity of opinion, including such important 
crops as maize, and of sweet potato and tobacco in the New Guinea highlands. Later he was 
able to check this in the herbarium by studying the lists of the early BANKs & SOLANDER col- 
lections made during the first of Cook’s voyages in the Pacific. On this subject he published a 
book which, notwithstanding acid criticisms on ‘certain diffusionists’, is a magnificent source on 
the subject of ethnobotany affecting all tropical botany. In passing I remark that a similar study 
should still be made on the ComMMeErRSON collections, as a second proof for his theory. 

I do not know of course what MERRILL’s reaction would have been to a dedication of a volume 
of our Flora to him. In comparison with the great many honours fallen to him, honorary docto- 
rates and honorary memberships, memberships of academies of science, the gold medal of the 
French Ministry of Agriculture, the gold medal of the Linnean Society of London, the Geoffrey 
St. Hilaire medal, Officer of the Netherlands Order of Oranje Nassau, and so forth, our dedication 


(13) 


FLORA MALESIANA 


is only a very small one. Still, I feel certain that it would have ranked very high with him, 
because it lies precisely in the field of botany in which he spent probably the best years of his life, 
and which intrigued him most of all. 

He had never any intention to join in our work by personal contributions. Apart from the 
fact that the Flora started when he was already well in his seventies, he had used his capacities 
primarily in making tools for botanical workers, such as his critical evaluations of basic early 
works (BLANCO, RUMPHIUS, BURMAN, LOuREIRO, and in his last years ROXBURGH), enumerations 
of the flora of local areas (the Philippines, Borneo, Hainan, Guam, Banguey I.), the elaboration 
of large single collections from various parts of the East (Burma, Sumatra, NE. Borneo, New 
Guinea, together with Miss Dr. L. M. Perry) and in compiling valuable bibliographies (the 
Pacific, East Asia, the Philippines, Borneo). Through his immense knowledge of plants and books 
he could produce such works at a speed at which normally large teams of workers would have 
been necessary. 

By necessity there is a certain restriction of purpose and objects in making such tools, several 
were also called ‘bibliographical enumerations’, indicating that they were not critical and that 
he was not responsible for the correctness of the names. To expect or request this is of course 
completely unfair and unjust; they were intended as tools, and this purpose was accomplished. 

There was a similar restriction in his systematical work, as he confined his revisions mostly to 
limited areas, many in the Philippines (Syzygium, grasses, Leguminosae, etc.), Borneo or New 
Guinea. Nevertheless it was an immense achievement, but at a certain level, as good as it could 
be under the circumstances, but largely floristic and not critical. He did not contribute essays on 
theoretical systematics. 

Quoting RosBIns, ‘it has been said that MERRILL seldom went far below the surface, and that 
he was content in most cases to classify the plants with which he dealt’. Ropsins defended 
MERRILL in commenting: ‘This type of research was proper for the region he studied and was 
the only procedure which permitted him to do what he did in his lifetime. It is exactly this 
characteristic ability to deal superficially with extraordinarily large numbers of plants that makes 
so apt the epithet ‘the American Linnaeus’ which has been applied to him. Actually, however, 
MERRILL did go below the surface. He recognized that classification was prerequisite for other 
investigations and the magnitude of the job he set for himself left little time to pursue anything 
else. But his studies of the floristic and faunistic relationships of the Philippines to other Malesian 
areas, of the significance of vernacular plant names, and of the origin of cultivated plants are 
examples of ‘below the surface’ investigations carried out by MERRILL.’ I may add his excellent 
synthesis of the distribution of the Dipterocarpaceae which induced him to make a rough outline 
of the phytogeography of Malesia, and especially its relation to the Formosan flora, in the light 
of correlating biological distribution with geological history. 

To be fair, we should always keep in mind that during the period in which he achieved his 
great contributions to Malesian botany, that is 1902-1923, he started without a predecessor from 
absolute scratch, without personnel, without a book or collection, in an almost unexplored very 
rich archipelago covered largely by primary forest. Later too he had to work under scientific 
vacuum conditions in that he had almost no colleagues around him in Manila, there were hardly 
any botanists in the whole of Malesia — except for VALETON and J. J. SmirH in Bogor and 
RIDLEY in Singapore — while the floras of the islands surrounding the Philippines (Borneo, Ce- 
lebes, the Moluccas and New Guinea) were botanically only known in the most fragmentary way. 

If he had aimed at a critical systematical study of the Philippine flora in 1902, he would have 
had to wait for half a century for publication and could never have composed the major tools 
which now belong to our standard bibliographical equipment. 


(14) 


Dedication 


Even in the Netherlands Indies, which had an infinitely better botanical basis, TREUB had at 
the turn of the century concluded that a composition of a comprehensive Flora would be entirely 
premature because of the primitive state of exploration and publication and that only the framing 
of the very local ‘Flore de Buitenzorg’ (from Batavia to the peak of Mt Gedeh, W. Java) could 
be realized as a cautious approach to later projects of large size. Even for this very restricted 
Flora the volume on Phanerogams never appeared, except for the Orchids. And that was for 
Java, botanically the best known island in the whole archipelago with a proportionally poor 
flora compared with the true Malesian element in the Larger Sunda islands. Even a dozen years 
later BACKER spent three years in the field to get an overall coverage of the Javanese flora! 

This illustrates the desperate position MERRILL had to face, a position that contemporary 
botanists working on the Malesian flora do not always realize. I have sometimes traced arro- 
gance in the rejection by a few contemporary, ‘angry’ youngsters of MERRILL’s floristic methods 
and premature publication of novelties. It also is for their education that I have in some detail 
accounted for MERRILL’s work and life, projected on the background of the state of Malesian 
botany in the first decades of this century, with full exposure of the then prevailing conditions 
in the hope that they will reach a better understanding of the level at which MERRILL had to 
work by necessity. I have also pointed out what our present-day knowledge owes to his collecting 
drive and to the pioneer works, bibliographies and other tools which are in our constant use 
and which he had created from scratch. Possibly they may ask themselves what they would have 
achieved had they stood in MERRILL’s shoes in 1902. 

Naturally it is a blessing to be able to work now in well-equipped centres provided with ample 
facilities, under social security and pension conditions, at leisure on a regional-monographic 
basis. But let us remember that we harvest what others have sown during the past seven decades. 

Among those who paved the way MERRILL was the outstanding figure, a man of boundless 
energy and vision, a great organisator and a great botanist. It is for these reasons that we dedicate 
this volume with due respect to the memory of this prominent American scientist. 


BIOGRAPHICAL SOURCES 


BREMEKAMP, C. E. B.: Levensbericht, E. D. Merrill. Jaarboek Kon. Ned. Ak. Wet. (1955/56) 1-7. 

BurKILL, I. H.: Prof. E. D. Merrill. Nature 177 (1956) 687-688. 

—: Elmer Drew Merrill. Proc. Linn. Soc. Lond. 168 (1957) 51-56. 

Ewan, J.: E. D. Merrill. J. Wash. Ac. Sci. 46 (1956) 267-268. 

Howarp, R. A.: Elmer Drew Merrill, 1876-1956. J. Arn. Arb. 37 (1956) 197-216, port. 

Lam, H. J.: A Dutch tribute to Merrill’s work. Natuurwet. Tijdschr. Ned. Ind. 102 (1946) 153- 
154. 

LANJouw, J.: Elmer D. Merrill, 1876-1956. Taxon 5 (1956) 21. 

MERRILL, E. D.: Autobiographical: Early years, the Philippines, California. Asa Gray Bull. n.s. 2 
(1953) 335-370, 3 ports. 

QUISUMBING, E.: Elmer Drew Merrill. Philip. J. Sc. 85 (1956) 181-188. 

RicketTT, H. W.: Elmer Drew Merrill, 1876-1956. J. N.Y. Bot. Gard. 6 (1956) 84, 90. 

RossBins, W. J.: Elmer Drew Merrill (1876-1956). Yearb. Amer. Philos. Soc. (1956) 117-119. 

—: Elmer Drew Merrill, 1876-1956. A Biographical Memoir. Biograph. Memoirs 32 (1958) 
273-333. 

ROLLINs, R. C.: Elmer Drew Merrill, Administrator and Botanist. Science, n.s. 123 (1956) 831- 
832. 


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FLORA MALESIANA 


ScHuLtTes, R. E.: Elmer Drew Merrill. An Appreciation. Taxon 6 (1957) 89-101, 2 ports. and a 
caricature. 

VeRDOORN, F. (ed.): Merrilleana. A selection of the general writings of Elmer Drew Merrill, 
Sc.D., LL.D. Chronica Botanica 10 (1946) 129-393. 

WaLkKeR, E. H.: Dr. Elmer D. Merrill, 1876-1956. Pacific Sci. 11 (1957) 135-136. 


C. G. G. J. VAN STEENIS 


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ABBREVIATIONS AND SIGNS 


acc. = according 

Ak. Bis. = Aklan Bisaya (Philip. language) 
Alf. Cel. = Alfurese Celebes (language) 
alt. = altitude 

Anat. = Anatomy 

Ap. = Apayao (Philip. language) 

app. = appendix, appendices 

appr. = approximate 


Apr. = April 
Arch. = Archipelago 
atl. = atlas 


auct. div. = auctores diversi; various authors 

auct(t). mal. = auctores malayenses; authors 
dealing with Malesian flora 

auct(t). plur. = auctores plures; several authors 

Aug. = August 

Bag. = Bagobo (Philip. language) 

basionym = original name of the type specimen; 
its epithet remains permanently attached to the 
taxon which is typified by it provided it is of the 
same rank 

Bg. = Buginese (language) 

Bik. = Bikol (Philip. language) 

Bil. = Bila-an (Philip. language) 

Bill. = Billiton 

Bis. = Bisdya (Philip. language) 

Bon. = Bontok (Philip. language) 

Born. = Borneo 

Bt = Bukit; mountain 

Bug. = Buginese (language) 

Buk. = Bukidnon (Philip. language) 

. = circiter; about 

C. Bis. = Cebu Bisaya (Philip. language) 

cf. = confer; compare 

Chab. = Chabecano (Philip. language) 

citations = see references 

cm = centimetre 

c.n. = see comb. nov. 

comb. nov. = combinatio nova; new combination 

c.S. = cum suis; with collaborators 

cum fig. = including the figure 

cur. = curante; edited by 

D (after a vernacular name) = Dutch 

Daj. = Dyak (language) 

Dec. = December 

D.E.I. = Dutch East Indies 

descr. added behind a reference = means that this 
contains a valid description 

diam. = diameter 

Distr. (as an item) = Distribution 

Distr. (with a geographical name) = District 

ditto = the same, see do 

Div. = Division, or Divide 

div. = diversus (masc.); various 

do = ditto (Ital.); the same 

Dum. = Dumagat (Philip. language) 

dupl. = duplicate 

E = east (after degrees: eastern longitude) 

E (after a vernacular name) = English 

Ecol. = Ecology 

ed. = edited; edition; editor 

e.g. = exempli gratia; for example 

elab. = elaboravit; revised 

em(end). = emendavit; emended 


em(erg). ed. = emergency edition 

Engl. = English 

etc., &c. = et cetera; and (the) other things 

ex auctt. = ex auctores; according to authors 

excl. = exclusus (masc.); excluding, exclusive of 

ex descr. = known to the author only from the 
description 

f. (before a plant name) = forma; form 

f. (after a personal name) = filius; the son 

f. (in citations) = figure 

fam. = family 

Feb(r). = February 

fide = according to 

fig. = figure 

fl. = flore, floret (floruit) ; (with) flower, flowering 

For. Serv. = Forest Service 

Sr. = fructu, fructescit; (with) fruit, fruiting 

Fr. (after a vernacular name) = French 

G. = Gunung (Malay); mountain 

Gad. = Gaddang (Philip. language) 

gen. = genus; genus 

genus delendum = genus to be rejected 

Germ. = German 

geront. = Old World 

haud = not, not at all 

holotype = the specimen on which the original 
description was actually based or so designated 
by the original author 

homonym = a name which duplicates the name of 
an earlier described taxon (of the same rank) but 
which is based on a different type species or type 
specimen; all later homonyms are nomencla- 
turally illegitimate, unless conserved 

I. = Island 

ib (id). = ibidem; the same, in the same place 

Ibn. = Ibanag (Philip. language) 

ic. = icon, icones; plate, plates 

ic. inedit. = icon ineditum, icones inedita; inedited 
plate(s) 

id. = idem; the same 

i.e. = id est; that is 

If. = Ifugao (Philip. language) 

Ig. = Igorot (Philip. language) 

Ilg. = Llongot (Philip. language) 

Ik. = Lloko (Philip. language) 

in adnot. = in adnotatione; in note, in annotation 

incl. = inclusus (masc.); including, inclusive(ly) 

indet. = indetermined 

Indr. = Indragiri (in Central Sumatra) 

inedit. = ineditus (masc.); inedited 

in herb. = in herbario; in the herbarium 

in litt. = in litteris; communicated by letter 

in sched. = in schedula; on a herbarium sheet 

in sicc. = in sicco; in a dried state 

in syn. = in synonymis; in synonymy 

Is. = Islands 

Is. (after a vernacular name) = 
language) 

Ism. = Isamal (Philip. language) 

isotype = a duplicate of the holotype; in arboreous 
plants isotypes have often been collected from a 
single tree, shrub, or liana from which the 
holotype was also derived 

Iv. = Ivatan (Philip. language) 


Isindi (Philip. 


(17) 


FLORA MALESIANA 


J(av). = Javanese (language) 

Jan. = January 

Jr = Junior 

Klg. = Kalinga (Philip. language) 

Kul. = Kulaman (Philip. language) 

Kuy. = Kuyonon (Philip. language) 

Lamp. = Lampong Districts (in S. Sumatra) 

Lan. = Lanao (Philip. language) 

lang. = language 

l.c. = loco citato; compare reference 

lectotype = the specimen selected a posteriori 
from the authentic elements on which the taxon 
was based when no holotype was designated or 
when the holotype is lost 

livr. = livraison, part 

(heea— e-x(plurs) 

m = metre 

M = Malay (language) 

Mag. = Magindando (Philip. language) 

Mak. = Makassar, Macassar (in SW. Celebes) 

Mal. = Malay(an) 

Mal. Pen. = Malay Peninsula 

Mand. = Mandaya (Philip. language) 

Mang. = Mangyan (Philip. language) 

Mar. = March 

Mbo = Manobo (Philip. language) 

Md. = Madurese (language) 

Minangk. = Minangkabau (a Sumatran language) 

min. part. = pro minore parte; for the smaller part 

mm = millimetre 

Mng. = Mangguangan (Philip. language) 

Morph. = Morphology 

ms(c), MS(S) = manuscript(s) 

Mt(s) = Mount(ains) 

n. = numero; number 

N = north (after degrees: northern latitude); or 
New (e.g. in N. Guinea) 

NE. = northeast 

nec = not 

neerl, = Netherlands, Netherlands edition 

Neg. = Negrito (Philip. language) 

N.E.I. = Netherlands East Indies 

neotype = the specimen designated to serve as 
nomenclatural type when no authentic speci- 
mens have existed or when they have been lost; 
a neotype retains its status as the new type as 
long as no authentic elements are recovered and 
as long as it can be shown to be satisfactory in 
accordance with the original description or 
figure of the taxon 

N.G. = New Guinea 

N.I. = Netherlands Indies 

no = numero; number 

nom. = nomen; name (only) = nomen nudum 


nom. al. = nomen aliorum; name used by other 
authors 

nom. alt(ern). = nomen alternativum; alternative 
name 

nom. cons(ery). = nomen conservandum, nomina 


conservanda; generic name(s) conserved by the 
International Rules of Botanical Nomenclature 

nom. fam. cons. = nomen familiarum conservan- 
dum; conserved family name 

nom. gen. cons. = see nomen conservandum 

nom. gen. cons. prop. = nomen genericum conser- 


(18) 


vandum propositum; generic name proposed for 
conservation 


nom. illeg(it). = nomen illegitimum; illegitimate 
name 
nom. leg(it). = nomen legitimum; legitimate name 


nom. nov. = nomen novum; new name 

nom. nud. = nomen nudum; name published with- 
out description and without reference to pre- 
vious publications 

nom. rej(ic). = nomen rejiciendum; name rejected 
by the International Rules of Botanical No- 
menclature 

nom. seminudum = a name which is provided with 
some unessential notes or details which cannot 
be considered to represent a sufficient descrip- 
tion which is, according to the International 
Rules of Botanical Nomenclature, compulsory 
for valid publication of the name of a taxon 

nom. subnudum = nomen seminudum 

nom. superfl. = a name superfluous when it was 
published; in most cases it is a name based on 
the same type as an other earlier specific name 

non followed by author’s name and year, not 
placed in parentheses, and put at the end of a 
citation = means that this author has published 
the same name mentioned in the citation in- 
dependently. These names (combinations) are 
therefore homonyms. 
Compare p. 111 under Wahlenbergia lines 6 & 7 
where there appear to be three different genera 
all called Lightfootia by three different authors, 
and belonging to three different families. The 
same can happen to taxa of lower ranks, for 
example species; compare p. 1294 lines 7 & 8 
under Lobelia heyniana, where it appears that 
there are two different species of Lobelia named 
L. decurrens, viz by ROTH and by CAVANILLES 
in which the latter antedates the former 

(non followed by abbreviation of author’s name) 
before a reference (citation) headed by an other 
author’s name = means that the second author 
has misinterpreted the taxon of the first author. 
Compare for example p. 1265 under Lobelia 
alsinoides lines 7 & 8 the synonym name L. 
filiformis; CAVANILLES misapplied in his de- 
scription and figure the name LAMARCK had 
given to another species through an erroneous 
identification. The sense CAVANILLES gave to the 
Lamarckian plant name does not invalidate the 
latter: CAVANILLES’s use of the name also does 
not represent a proper synonym; his name has 
no status and its mention serves only to indicate 
the identity of his text and plate 

non al. = non aliorum; not of other authors 

non vidi = not seen by the author 

noy. = nova (femin.); new (species, variety, e/c.) 

Nov. = November 

n.s. = new series 

Nn. Sp. = nova species; new species 

n. (sp.) prov. = nomen (specificum) provisorium; 
provisional new (specific) name 

n.v. = non vidi; not seen 

NW. = northwest 

Oct. = October 

op. cit. = opere citato; in the work cited 


Abbreviations and signs 


p. = pagina; page 

P. = Pulau, Pulu (in Malay); Island 

Pal(emb). = Palembang 

Pamp. = Pampangan (Philip. language) 

Pang. = Pangasinan (Philip. language) 

paratype = a specimen cited with the original 
description other than the holotype 

part. alt. = for the other part 

P. Bis. = Panay Bisaya (Philip. language) 

P.[. = Philippine Islands 

pl. = plate 

plurim, = plurimus; most 

p.p. = pro parte; partly 

pr. max. p. = pro maxima parte; for the greater 
part 

pro = as far as is concerned 

prob. = probabiliter; probably 

prop. = propositus; proposed 

Prov. = Province 

pr.p. = pro parte; partly 

pt = part 

quae est = which is 

quoad basionym, syn., specimina, etc. = as far as 
the basionym, synonym((s), specimen(s), efc. are 
concerned 

references = see for abbreviations the list in vol. 5, 
pp. cxly-clxv 

Res. = Residency 

resp. = respective(ly) 

S = south (after degrees: southern latitude) 

S (after a vernacular name) = Sundanese (lan- 
guage) 

Sbl. = Sambali (Philip. language) 

SE. = southeast 

sec. = secus; according to 

sect. = sectio; section 

sens. ampl. (ampliss.) = sensu amplo (amplissimo) ; 
in a wider sense, in the widest sense 

sens. lat. = sensu lato; in a wide sense 

sens. str. (strictiss.) = sensu stricto (strictissimo) ; 
in the narrow sense, in the narrowest sense 

Sept. = September 

seq., Seqq. = sequens, sequentia; the following 

ser. = series 

s.l. = sensu lato; in a wide sense 

S.-L. Bis. = Samar-Leyte Bisaya (Philip. language) 

Sml. = Samal (Philip. language) 

5.n. = sine numero; (specimen) without the col- 
lector’s number 

Sp. = Spanish (language) 

sp(ec). = species; species 

specim. = specimen(s) 

sphalm. = sphalmate; by error, erroneous 

Spp. = species; species (plural) 

Sr — Senior 

5.5. = see sens. str. 

ssp. = subspecies; subspecies 

Susirs — SEG sens. str: 

Stat. nov. = status nova; proposed in a new rank 

Sub. = Subanum (Philip. language) 

subg(en). = subgenus; subgenus 

subsect. = subsectio; subsection 

subsp. = subspecies; subspecies 

Sul. = Sulu (Philip. language) 

Sum. E.C. = Sumatra East Coast 


Sum. W.C. = Sumatra West Coast 

Suppl. = Supplement 

SW. = southwest 

syn. = synonymum; synonym 

synonyms = the names of taxa which have been 
referred to an earlier described taxon of the 
same rank and with which they have been united 
on taxonomical grounds or which are bound 
together nomenclaturally 

syntypes = the specimens used by the original 
author when no holotype was designed or more 
specimens were simultaneously designated as 
type 

t. = tabula; plate 

Tag. = Tagalog (Philip. language) 

Tagb. = Tagbanua (Philip. language) 

Tagk. = Tagaka-olo (Philip. language) 

Tapan. = Tapanuli (in NW. Sumatra) 

taxon = each entity throughout the hierarchic 
ranks of the plant kingdom which can be 
described and discriminated from other taxa of 
the same rank 

Taxon. = Taxonomy 

Tg = Tandjung (Malay); cape 

Ting. = Tinggian (Philip. language) 

Tir. = Tirurai (Philip. language) 

transl. = translated 

type = each taxon above the rank of a species is 
typified by a type belonging to a lower rank, for 
instance a family by a genus, a genus in its turn 
by a species; a species or infraspecific taxon is 
typified by a specimen. The name of a taxon is 
nomenclaturally permanently attached to its 
type; from this it cannot be inferred that the 
type always represents botanically the most 
typical or average structure found in the cir- 
cumscription of the taxon 

type specimen = the specimen or other element to 
which the name of a species or infraspecific 
taxon is (nomenclaturally) permanently at- 
tached; botanically a type specimen is a random 
specimen on which the name was based by de- 
scription. Therefore, it does not need to repre- 
sent the average or most typical representative 
of a population. See holotype, isotype, lectotype, 
syntype, paratype, and neotype 

typ. excl. = typo excluso; type exluded 

typ. incl. = typo incluso; type included 

typus = see type and type specimen 

var. = varietas; variety 


var. nov. = varietas nova; new variety 
Vern. = Vernacular 
vide = see 


viz = videlicet; namely 

vol. = volume 

W = west (after degrees: western longitude) 
Yak. = Yakan (Philip. language) 


@ = diameter 

3 = male (flower, etc.) 

2 = female (flower, etc.) 

3, 9 = bisexual (flower) 

) (2) = dioecious with unisexual flowers 
©) = monoecious with unisexual flowers 


(19) 


FLORA MALESIANA 


(3%) = polygamous 


< = less than (size, number, efc.) 
(2°) = polygamous x 2/5 = 2/5 of natural size 
oo = many montana = means that the epithet montana is 
~ = more than (in size, number, etc.) 


that of a hybrid 


(20) 


ADDENDA, CORRIGENDA ET EMENDANDA 


As was done in the preceding volumes, it seemed useful to correct some errors which have crept into the 
text of volumes 4, 5 & 6 as well as to add some additional data, new records, and new species or other 
taxa which came to our knowledge and are worth recording. 

Additions of the Amaranthaceae | owe to Dr. R. C. BAKHUIZEN VAN DEN BRINK f. and Mr. J. F. 
VELDKAMP, of the Alismataceae and Hydrocharitaceae to Dr. C. DEN HARTOG, of the Celastraceae and 
Thymelaeaceae p.p. to Dr. DinG Hou, of the Malpighiaceae to Dr. M. Jacoss, of the Burseraceae p.p. 
to Dr. C. KALKMAN, of the Caprifoliaceae to Dr. J. H. KERN, of the Burseraceae p.p., Connaraceae, 
Dichapetalaceae, Goodeniaceae and Loganiaceae to Dr. P. W. LEENHOUTS, of the Gnetaceae to Dr. F. 
MARKGRAF, Of the Simaroubaceae to Mr. H. P. Nootesoom, of the Convolvulaceae to Dr. S. J. VAN 
OosTsTROoM, of the Thymelaeaceae p.p. to Mr. H. K. Airy SHAW, of the Ericaceae and Flacourtiaceae 
to Dr. H. SLEUMER. 

Printing errors have only been corrected if they might give rise to confusion. 

Volume and page number are separated by a colon. Page numbers provided with either a or b denote 
respectively the left and right columns of a page. 


Aceraceae 4556, cultivated, a recent introduction; 
Morobe Distr., Bulolo, NGF 7384). All 
4: 3-4, Acer laurinum Hassk. identifications by SAUER, 1971. 
592ab Add to Distr.: Rare in Borneo and the 4: 79b Change the name of species 8 into: 
few localities at unusual lowland altitude; 8. Amaranthus hybridus L. ssp. incurvatus 
now also found on Mt Kinabalu at c. (GREN. & GODR.) BRENAN var. panicula- 
1000 m (SAN 38438); probably also in the tus (L.) MANsr. See MANSFELD, Die 
mountains of N. Thailand, as E. MURRAY Kulturpfl. Beih. 2 (1959) 54, and BRENAN, 
merged (in sched.) A. garrettii CRAIB, Watsonia 4 (1961) 268. 
Kew Bull. (1920) 301, with it. 4: 91  Emend the key to the species of Al/ter- 
nanthera as follows, second entry of fork 
Aizoaceae under 3: 
3. Three outer tepals in their lower 
4: 269a Glinus lotoides L. 1/31/59 distinctly 3-nerved, their bases 
Add to synonymy: Holosteum hirsutum at least indurate. Anthers 5. Pseudo- 
L. Sp. Pl. (1753) 88. Holotype from staminodes well-developed, wider than 
India. Cf. STEEN. Blumea 13 (1965) 167. long to ligular, lobed or fimbriate. 
4: 274a Trianthema triquetra ROTTL. ex WILLD. Hairs dentate. 
Add to Distr.: Philippines (Mindanao). 3a. Leaves mucronate, often coloured. 
Cf. STEEN. Blumea 12 (1964) 320. Anthers linear, °/4-1 mm long; 
pseudo-staminodes ligular, apex 
Alismataceae (DEN HARTOG) fimbriate, as long as to longer than 
the stamens. Pistil bottle-shaped, 
5: 327 Bottomline, replace ‘6. S. sagittifolia ssp. apex not emarginate. Apparently 
leucopetala’ by: 6. S. trifolia. never setting seed in Malesia. 
5: 332b Replace the name Sagittaria sagittifolia 3. A. ficoidea 
ssp. leucopetala (MiQ.) Hartoc by: 3a. Leaves acute, not mucronate (in 
6. Sagittaria trifolia LINNE, Sp. Pl. 2 Malesia). Anthers reniform to 
(1753) 933, and add the subspecific name ellipsoid, !/3—?/5 mm long; pseudo- 
to its synonymy. There is no change in the staminodes wider than long, with 
text. 3-4 + triangular lobes, reaching 
5: 333b In text line 3 from bottom, replace ‘S. the base of the anthers or shorter. 
sagittifolia ssp. leucopetala’ by: S. trifolia. Pistil broadly obcordate. Seeds 
5: 334b Add to the Notes: This taxon cannot be usually developed, (broadly) obcor- 
maintained as a subspecies of S. sagitti- date, narrowly winged. 
folia, as the sepals in the mature 9 flowers 3a. A. paronychioides 
are reflexed. For this reason it belongs to 4: 93a, Alternanthera ficoidea (L.) R.BR. ex 
another species group within the genus. 594b R. & S. 1819, non A. ficoides P. BEAUVY. 
Fl. Oware & Benin 1818. As these are 
Amaranthaceae homonyms (Code Art. 75) this specific 
(BAKHUIZEN VAN DEN BRINK, name is illegitimate. 
VAN STEENIS & VELDKAMP) We are, however, not certain of its full 
synonymy and for this reason tentatively 
4: 79b Amaranthus dubius Marv. refrain from making new combinations 
Add to Distr.: Central Java (Mt Lawu, both for the species and the variety. It 
Temanggung: LORZING 346), Lesser may be that Telanthera manillensis WALP. 
Sunda Is. (Alor: JAAG 413), New Guinea 1843 contains the oldest basionym. 


(West: Star Mts, Sibil Valley, KALKMAN To avoid this difficulty we accept tenta- 


FLORA MALESIANA 


[ser. I, vol. 68 


Fig. 1. Alternanthera bettzickiana (REGEL) NICHOLS. /. Flower, abaxial, without bract and bracteoles 


< 12, 2. ditto, without abaxial tepals, » 


12, 3. staminal tube, « 24, 4. young fruit, x 24. — A. parony- 


chioides St. Hit. var. paronychioides. 5. Flower, abaxial, without bract, « 12, 6. ditto, without bracteoleS 


and abaxial tepals, 


4: 93b 


12, 7. staminal tube, x 


tively specific status for this taxon in 
Flora Malesiana: 

Alternanthera bettzickiana (REGEL) NI- 
CHOLS. Ill. Dict. Gard. 1 (1884) 59 
(‘bettzichiana’); Voss in Vilmorin’s Blu- 
mengartn. ed. 3 (by SIEBERT & Voss), 
1 (1895) 69 (non vidi, ex Ind. Kew.); 
ASCHERS. & GRAEBN. Synopsis 5, 1 
(1914) 365. — Telanthera bettzickiana 
REGEL, Gartenflora 11 (1862) 178, descr.; 
Ind. Sem. Hort. Petrop. (1862) 28 (non 
vidi). — Fig. 1. 

VELDKAMP (Blumea 19, 1971, 169) 
assumes the addition by NICHOLSON 
“Brasilia, 1862’ to be a very indirect 
reference to REGEL’s basionym with 
which I can agree. VELDKAMP, /. c., also 
indicated that var. versicolor (REGEL) 
BACKER, sometimes regarded as a sepa- 
rate species, must be regarded as a 
synonym of A. bettzickiana. 

Add the following species: 

3a. Alternanthera paronychioides St. Hiv. 
Voy. Brésil 2, 2 (1833) 43; DuLTA & 
Mitra, Ind. Forester 87 (1961) 304, f. 2; 
PEDERSEN, Darwiniana 14 (1967) 437; 
VELDKAMP, Blumea 19 (1971) 167, f. 6-8. 


24, 8. young fruit (J-4 Cayosa 75, 5-8 CAyosa 104). 


var. paronychioides — Fig. 1. 

Prostrate, branched annual. Branches 
up to 70 cm, rooting at the nodes, angular 
and villose at apex, becoming terete and 
glabrous at base, indument only persis- 
tent at the nodes. Hairs dentate. Leaves 
spathulate-oblong to -lanceolate, up to 
7 by 2 cm, acute, never mucronate, base 
cuneate, narrowed into the petiole, 
moderately appressed-pubescent to glab- 
rous, usually tufted under the inflores- 
cences. Spikes sessile, apical on short 
axillary branches, subglobose to shortly 
cylindric, up to 2 by 1 cm. Bracts, 
bracteoles scarious, white. Bracts ovate- 
oblong, 21/4-3 by 11/4-11/2 mm, acu- 
minate, glabrous, -+_ convex, 2/3—3/4 
times as long as tepals, longer than brac- 
teoles. Bracteoles ovate-oblong, 13/4—21/2 
by 3/4 mm, acute to acuminate, glabrous, 
+ falcately folded along midrib. Tepals 
oblong to lanceolate, acute to mucronu- 
late, in lower half 3-nerved and sparsely 
strigose, upper half stiff, but not coria- 
ceous. Adaxial tepal 3—4 by 4/s—11/4 mm, 
flat; abaxials 31/44 by 1-11/4 mm, flat; 
laterals 21/4—31/4 by 1—11/4 mm, folded 


September 1972] 


Addenda, corrigenda et emendanda 


917 


4: 964, 
594b 


5: 415a 


5: 207b 


along midrib. Stamens 5, all fertile (in 
the material seen); filaments at base 
united into a c. '/4 mm high cup, free 
parts c. 1/2 mm long, filiform; anthers 
reniform to ellipsoid, 4/3—!/4 mm long, 
yellow; pseudo-staminodes wider than 
long, with 3-4 + triangular lobes, reach- 
ing the base of the anther or shorter. 
Pistil broadly obcordate with a short, 
stout style in the notch; stigma capitate, 
papillose. Utricle broadly obcordate to 
obcordate, up to 13/4 by 11/4 mm, 
narrowly winged, brown; the plants seem 
to fructify during drying; so young fruits 
are usually present, contrary to the situa- 
tion in A. ficoidea ssp. bettzickiana, where 
they have never been observed in Malesia. 

Distr. Native of tropical America, 
introduced as an alien in other countries, 
including Europe and _ Indo-Malesia, 
India, Thailand; in Malesia: West-Java, 
Philippines (Luzon: Manila, Quezon 
City; Guimaras I.; Mindanao). 

Ecol. Disturbed places, railway tracks, 
banks of rivers and lakes, up to 250 m. 

Vern. & Uses. Simsim (Bis.), used for 
feeding hogs (Mindanao). 

Note. By BACKER confused with A. 
ficoidea ssp. bettzickiana and in habit 
difficult to discern from it, but distinctly 
different in the structure of the filaments, 
pseudo-staminodes, and anthers. 
Gomphrena celosioides MART. 

Add to references: STEEN. Nova Guinea, 
Bot. 23 (1965) 495. 

The spreading of this weed continues 
steadily eastwards and has now reached 
Timor, Papua, and Micronesia. It was 
also found in Goodenough I., off NE. 
Papua, along an old military road 
(Brass 24432). 


Batidaceae 


Batis argillicola VAN ROYEN. 

Add to Distr.: It also occurs in Northern 
Queensland. Cf. S. T. BLAKE, Proc. R. 
Soc. Queensl. for 1961, 73 (1963) 61. 
It occurs there in Burke Distr., near 
Karumba, at the mouth of the Norman 
R., on clay-pans adjacent to the shore, in 
exactly the same habitat as near Merauke. 


Betulaceae 


The basionym of Alnus maritima 
(Marsu.) NuTTALL is Betula-Alnus mari- 
tima MARSHALL, Arb. Am. (1785) 20. 
Though not noted, I had not seen this 
work and depended on _ authority. 
Dr. BAKHUIZEN VAN DEN BRINK Saw it at 
Paris, in 1959, and reported that the only 
description or distinction mentioned by 
MARSHALL is ‘leaves long and narrow’. 
This, I believe, cannot be accepted even 


5: 208 


5) 210 


Se li 


5: 214a 


Se021Sa 


5: 218a 


5: 220 


as a specific diagnosis, and the name 
I regard as an invalid nomen seminudum. 
The proper name for this species is: 
Alnus japonica (THUNB.) STEUD. 

Legend fig. 1: Change Alnus maritima 
(MarsH.) Nutt. into Alnus japonica 
(THUNB.) STEUD. 


Burseraceae 


(LEENHOUTS, Dacryodes and Santiria in 
co-operation with KALKMAN) 


Add to Dispersal, 2nd paragraph: In 

Java, fruits of Canarium are occasionally 

eaten by bats (see VAN DER PiL, Acta 

Bot. Neerl. 6, 1957, 299). 

Add to Wood anatomy: BurGEss, Tim- 

bers of Sabah (1966) 60-70, f. 11. 

Insert after the section on 

anatomy: 

Phytochemistry. See HEGNAUER, Che- 

motaxonomie 3 (1964) 310-318, 647, 669. 

Add to Morphology, 3rd paragraph: See 

for a more detailed discussion on the 

stipules of the Burseraceae and some 
related families WEBERLING & LEEN- 

HoutTs, Abh. Ak. Wiss. Lit. Mainz M.-N. 

Kl. 1965, n. 10 (1966) 495-584. 

Protium javanicum Buro. f. 

Add to description under Branches, 

before ‘spines’: sometimes branched. 

Add to Distr.: Flores. 

Protium macgregorii 

LEENH. 

In description delete after 

(2 unknown). Insert after 

3 flowers. 

Add to Distr.: Also in Normanby I. 

Add to Ecol.: also in swamps. Change 

highest altitude into 1100 m. 

Garuga floribunda DECNE. 

Add to description, Ist sentence: decid- 

uous. 

Add to Distr.: Malay Peninsula. 

Dacryodes VAHL. 

Add to Distr.: According to NoRMAND, 

Comptes Rendus [Ve Réun. A.E.T.F.A.T. 

(1962) 291, the number of African species 

is about 10. 

Replace the Key to the species by the 

following one: 

1. Leaf-bearing branchlets c. 2 cm thick. 
Leaves 6-8-jugate; petiole c. 20 cm 
long; leaflets 20-60 cm long, with 
18-38 pairs of nerves. Inflorescence 
35-120 cm long, the main branches up 
to 55 cm long 5. D. kingii 

1. Leaf-bearing branchlets up to 11/2 cm 
thick. Leaves rarely more than 
4-jugate; petiole mostly up to 15 cm 
long; leaflets mostly less than 20, 
rarely up to c. 40 cm long, with 
usually less than 15, rarely up to 
25 pairs of nerves. Inflorescences 


Wood 


(Be Me “BAi) 


‘Flowers’: 
SPistiles aan 


918 


FLORA MALESIANA 


[ser. I, vol. 68 


mostly shorter, their main branches 
up to 30 (2) or 40 (4) cm long. 

2. Lowermost pair of leaflets more or 
less stipule-like,. often caducous. 
Inflorescences 10-140 cm long, their 
main branches up to 30 (2) or 40 (3) 
Cmiblong.)) ue ete a) laxa 

2. Lowermost pair of leaflets not much 
different from the others, not cadu- 
cous. Inflorescences mostly less than 
30, rarely up to 60 cm long, their 
main branches up to c. 25 cm long. 

3. Indumentum at least partly consist- 
ing of minute, stellate hairs. 
16. D. nervosa 
3. Indumentum consisting of simple 
hairs only. 
4. Nodes of leaf-rachis distinctly 
swollen. 

5. Leaflets when dried greenish 
above. Inflorescences mostly axil- 
lary, together sometimes pseudo- 
terminal, rarely terminal. Calyx 
sparsely hairy, corolla (sub)glab- 
rous. Fruits distinctly bulging 
on one side . . 1. D. rugosa 

5. Leaflets when dried brown above. 
Inflorescences terminal. Flowers 
densely tomentose. Fruits not 


bulging . . 7. D. rubiginosa 
4. Nodes of leaf-rachis not or hardly 
swollen. 


6. Inflorescences terminal (vegeta- 
tive terminal bud absent), more- 
over sometimes in the upper 
leaf-axils. 

7. Pith of branchlets without vascu- 
lar strands. 

8. Inflorescences densely woolly; 
calyx slightly pubescent, corol- 
la (sub)glabrous. 

2. D. costata 

8. Inflorescences densely minutely 


tomentose, including the 

flowers . . 6. D. incurvata 

7. Pith of branchlets with vascular 
strands. 


9. Leaf-bearing branchlets !/4-!/2 
cm thick, pith with some to 
many vascular strands. Petiole 
3-15 cm long, pith with some 
to many vascular strands; 
leaflets glabrous, with 6-18 
pairs of nerves. Peduncle 0-6 
cm long. 

Fruits 11/2-3 by 3/4-13/4 cm. 
6. D. incurvata 

9. Leaf-bearing branchlets c. 1/2 
cm thick, pith with some vascu- 
lar strands. Petiole 3-9 cm 
long, pith with some vascular 
strands; leaflets beneath on 
midrib and nerves densely 
pubescent, with 7-13 pairs of 
nerves. Inflorescences branched 


from the base. 
Fruits 2—21/2 by 1-11/4 cm. 
7. D. rubiginosa 


9. Leaf-bearing branchlets c. 1 cm 


thick, pith with many vascular 
strands. Petiole 9-15 cm long, 
pith with many vascular 
strands; leaflets glabrous, with 
13-18 pairs of nerves. Peduncle 
3/42 cm. Fruits 4-43/4 by 
221\/2cem . . 8. D. elmeri 


6. Inflorescences axillary (vegetative 


terminal bud present), sometimes 
together pseudoterminal. 


10. Leaflets pubescent beneath. 


11. Leaves 1-4-jugate. Inflores- 
cences short-peduncled. 

3. D. puberula 

11. Leaves 7—9-jugate. Inflores- 
cences long-peduncled. 

9. D. rostrata f. pubescens 


10. Leaflets glabrous or only hairy 


on the midrib beneath. 

12. Pith of branchlets with some 
to many, pith of petiole with 
several vascular strands. 

13. Branchlets densely minutely 
villous. Leaves 3-4-jugate; 
petiole strongly flattened 
at base, 9-15 cm long; leaf- 
lets 12-22 cm long, brown 
above when dry, shortly 
acuminate, with 13-18 pairs 
of nerves, reticulation incon- 
spicuous above. Inflores- 
cences 17-45 cm long, pe- 
duncle 3/4-2 cm. Stamens 
unknown. Fruit 4-4/4 by 
2-2!/2cm  . 8. D. elmeri 

13. Branchlets glabrous except 
the tip. Leaves 2—10-jugate; 
petiole terete to strongly 
flattened at base, 3-26 cm 
long; leaflets 3-25 cm long, 
brownish to greenish above 
when dry, mostly rather long 
and slender acuminate, with 
5-20 pairs of nerves, reti- 
culation manifest above. In- 
florescences 5-35 cm long, 
peduncle 0-15 cm. Stamens 
adnate to the disk. Fruit 
13/44 by 3/4-21/4 cm. 

9. D. rostrata 

13. Branchlets glabrous except 
the tip. Leaves 3-4-jugate; 
petiole strongly flattened at 
base, 41/2-10 cm long; leaf- 
lets 5121/2 cm long, green- 
ish above when dry, shortly 
acuminate, with 10-13 pairs 
of nerves, reticulation mani- 
fest above. Inflorescences 
71/2-10 cm long, peduncle 
31/2-5 cm. Stamens free 


September 1972] 


2216 


fe 220 
222b 


mn 


5: 224a 


5: 2246 


Sie225a 


Addenda, corrigenda et emendanda 


919 


from the disk. Fruit un- 
known 10. D. crassipes 
12. Pith of branchlets without, 


pith of petiole with few 
vascular strands. 
14. Petiole terete to slightly 


flattened at base. Wall of 
fruit kernel thick and hard. 
12. D. expansa 
14. Petiole strongly flattened to 
channelled at base. Wall of 
fruit kernel very thin. 

15. Leaflets 3-6 times as long 
as wide; leaves 3—15-jugate, 
the petiole 2—20 cm long, 
leaflets 6-42 cm long. 

14. D. longifolia 

15. Leaflets up to 3 times as 
long as wide; leaves up to 
5-jugate, the petiole up to 
8 cm long, leaflets up to 
17 cm long. 

16. Base of leaflets often 
oblique. Peduncle 0-4 cm 
long. Calyx 3 mm high. 
Fruit 21/4-31/2 by 11/4-2 


cm 11. D. macrocarpa 
16. Base of leaflets not 
oblique. Peduncle 1/2-1 


cm long. Calyx smaller. 
Fruit 16-18 by 9-11 mm. 
15. D. breviracemosa 
Dacryodes rugosa (BL.) H. J. LAM. 
Change in description: Branchlets ...; 
pith without or with many peripheral 
vascular strands ... Petioles terete to 
distinctly flattened at base. 
Add to Ecol.: Also in secondary forests. 
Dacryodes costata (BENN.) H. J. LAM. 
Change in description: Leaves ...., 
glabrous to densely pubescent ...; 
nerves up to 17 or 18 pairs. Fruits 
sometimes ellipsoid. 
Add to Ecol.: Also in secondary forests. 
Dacryodes laxa (BENN.) H. J. LAM. 
Description: add to Ist sentence: small 
buttresses sometimes present. Branchlets 
. .; pith sometimes with a closed cylinder 
of many small vascular strands. 
Dacryodes kingii (ENGL.) KALKMAN. 
Change in description, Ist sentence ‘12 
m’ into: 20 m. Branchlets ...; pith with 
some peripherally arranged to several, 
partly peripheral, partly scattered small 
vascular strands. Petiole up to 28 cm. 
Leaflets ...; apex up to 2 cm blunt- 
acuminate. Fruits oblong or ovoid, ..., 
more or less oblique. 
Dacryodes incurvata (ENGL.) H. J. LAM. 
Add to literature: ANDERSON, Gard. Bull. 
Sing. 20 (1963) 164. 
Description: Change greatest height of 
tree into 40 m. Leaves 1-—5-jugate. 
3 Panicles may be up to 30 cm long. 
Fruits ovoid to ellipsoid, up to 31/2 cm 


5: 226a 


5: 226b 


22a 


S22 1D 
5: 228a 


5: 2285 


5: 229a 


53 2296 


long, said to be orange when ripe. 

Add to Ecol.: SAN 25326 and 29004 are 
collected at 1400 and 1500 m altitude 
resp. Fi. Jan.Aug. 

Add: Uses. Fruits edible. 

Dacryodes rostrata (BL.) H. J. LAM. 
Description: Change greatest height of 
tree into 45 m. Branchlets exceptionally 
up to 15mm &. Petioles terete to strongly 
flattened at base, . . . Leaflets up to 25 cm 
long. Sometimes all nerves looped and 
joined near the margin. 

Add after Fruits: yellow to purple when 
ripe. 

Add to Uses: In Brunei cultivated by the 
Kedayans for the fruits, the pulp of 
which is eaten. 

Dacryodes macrocarpa (K1NG) H. J. LAM. 
Change in description, 3rd line, ‘glabrous’ 
into: hairy at the tip only. 

Line 4 from top, add after ‘Fruits ovoid’: 
to ellipsoid. 

var. macrocarpa. 

Add to literature: ANDERSON, Gard. 
Bull. Sing. 20 (1963) 164. 

Ecol.: Change highest altitude into 1400 
m. 

var. kostermansii (KALKM.) KALKMAN. 
Add to Distr.: N. Borneo. 

Dacryodes expansa (RIDL.) H. J. LAM. 
Add to literature: SMyTHIES, Common 
Sarawak Trees (1965) t. 8. 

Change in description line 4: Petioles 
21/2-91/2 cm. Change in line 9 ‘not 
arching’ into: arching or not. Change 
dimensions of fruits into: 31/2—-S by 2-31/2 
by 21/2-3 cm; add: rosy apple-red when 
ripe. 

Change Uses: Only the pulp of the 
fruits is eaten. 

13. Dacryodes papuana Husson. 

This turned out to be synonymous with 
Scutinanthe  brevisepala_ LEENH.; cf. 
LeEENH. Blumea 12 (1964) 19. 

Dacryodes longifolia (KING) H. J. LAM. 
Description: Add after tree: 10 m by 
30 cm. Line 2, after ‘glabrous’ add: apart 
from the tip. Add to fruits: red when ripe. 
Add to Distr.: Philippines (Mindanao). 
var. longifolia. 

Change in description: 4-15-jugate. 
Add to Distr.: Philippines. 

In Ecol. change as follows: Fr. Nov. 
(Mal. Pen.), May (Philippines). 

Add after 15. Dacryodes breviracemosa: 


16. Dacryodes nervosa (H. J. LAm) 
LEENH. Blumea 12 (1964) 19. — Santiria 
nervosa H. J. LAM, Ann. Jard. Bot. 
Btzg 42 (1932) 206, t. 11 f. 65; Bull. Jard. 
BoteBize sil, 1251932) S87sk. 6 t 29; 
KALKMAN, Blumea 7 (1954) 539; LEENH. 
Fl. Mal. I, 5 (1956) 233. 

Tree, 12-30 m by 20-100 cm, with up to 
21/2 m high buttresses which are 11/2 m 


920 


FLORA MALESIANA 


[ser. I, vol. 66 


spreading. Branchlets 4-8 mm thick, the 
tips ferrugineous-tomentose; terminal 
bud 1/2-1 cm long; pith without vascular 
strands. Leaves 1—4-jugate. Petioles 4—9 
(-14) cm, slightly to strongly flattened 
at the base; pith with few vascular strands. 
Leaflets oblong to ovate (to suborbicular), 
51/2-17 (-22) by 3—71/2 (-101/2) cm, upper 
surface greenish when dried, lower 
surface pubescent to glabrous, indumen- 
tum partly or entirely consisting of 
minute stellate hairs; base broadly 
cuneate to rounded; apex subabruptly, 
bluntly acuminate; nervation § rather 
prominent beneath; nerves 10-15 (—18) 
per side, more or less curved, mostly not 
distinctly looped and joined except 
towards base and apex; reticulations 
minute, hardly or not conspicuous above. 
Panicles axillary, often on short, leafless 
lateral shoots with a terminal bud, 
narrow, 11/2—51/2 (—20) cm long, ferrugi- 
neous-tomentose; peduncle up to 10 cm 
long. Flowers c. 2 mm long, sessile to 
shortly pedicelled, glabrous or stellate- 
tomentose and glabrescent. Calyx 11/2 
mm. Petals outside glabrous or hairy. 
Stamens free from the disk. Disk thick- 
annular or (2) cupular, radially furrowed 
and with undulate rim. Pistil in 3 flowers 
moderately reduced. /nfructescences up 
to c. 5 cm long, ferrugineous-tomentose, 
with a few fruits. Fruits ellipsoid, 
immature ones c. 11/2 by 0,9 cm. 

Distr. Malesia: Sumatra, Banka, 
Malay Peninsula, and Borneo. 

Ecol. Primary and secondary forests, 
up to 60 (-750) m. F/. Febr.—March, 
June, Oct. 

Vern. Bantan burung, kedondong tund- 
juk, sisip baniéng, Sum., asam-asam, 
Banka, kedudong, Mal. Pen., engai, 
mendjelih, selada, Borneo. 

Note. Originally, KALKMAN already 
inclined towards inclusion of this species 
in the genus Dacryodes, as appears from 
his identification labels, but finally he 
decided to leave it in Santiria by lack of 
evidence. Only when fruiting material 
became available the generic identity 
became sufficiently clear. 

Santiria Bu. 
Replace the Key to the species by the 
following one: 


1. Petiolules 3-3/4 cm long.6. S. ridleyi 
1. Petiolules up to 3 cm long. 

2. Bract-like cataphylls between the 
leaves present. Leaves (60—)80—135 
cm long . . 10. S. megaphylla 

2. No bract-like cataphylls between the 
leaves. Leaves rarely more than 
60 (up to 85) cm long. 

3. Anthers adnate (mostly also visible 
under the fruit). 


4. Pith of branchlets with vascular 


strands. Flowers 4-10 mm long, 
calyx 3-7 mm high, ¢ flower with 
6 fertile stamens. Stigma on fruit 
c. 90° excentric . 11. S. griffithii 
. Pith of branchlets without vascu- 
lar strands. Flowers 2-3 mm long, 
calyx 1-21/2 mm high, ¢ flower 
with 3 fertile stamens and some- 
times up to 3 staminodes. Stigma 

on fruit less than 90° excentric. 
12. S. rubiginosa 


3. Anthers basi- to dorsifixed. 
5. Calyx at anthesis 1!/2-3 mm high. 


6. Pith of branchlets with rather 
many vascular strands. 
2. S. mollis 
6. Pith of branchlets without vascu- 
lar strands. 

7. Terminal bud 2-3 cm _ long. 
Flowers 4-4!/2 mm long, calyx 
21/2-3 mm high. 

3. S. grandiflora 

7. Terminal bud up to 2 cm long. 
Flowers 2-4 mm, calyx 1-2 mm 
high. 

8. Terminal bud 1-2 cm long. 
Stigma on fruit less than 90° 
excentric . 5. S. oblongifolia 

8. Terminal bud 14/2-1 cm iong. 
Stigma on fruit usually more 
then 90° excentric. 

9. S. apiculata 


5. Calyx at anthesis up to 11/2 mm 


high. 

9. Pith of branchlets with vascular 
strands. 

10. Flowers (3) 3-4 mm, calyx 
11/2-21/2 mm high. 2. S. mollis 

10. Flowers 2 mm, calyx 1/2—3/4 
mmhigh . . 4. S. laevigata 

9. Pith of branchlets without vascu- 
lar strands. 

11. Mature leaves beneath hairy at 
least on midrib and nerves, 
mostly also on the veins. 

12. Calyx in anthesis 1/2-1 mm 
high. Stigma on fruits less 
than 90° excentric. 

1. S. tomentosa 

12. Calyx in anthesis 1-11/2 mm 
high. Stigma on fruits 90° or 
more excentric. 

9. S. apiculata 

11. Mature leaves beneath glab- 
rous or only hairy on midrib. 

13. Calyx during anthesis 1/2—3/4 


mm high. 
14. Stigma on the fruits up to 90° 
excentric . 4. S. laevigata 


14. Stigma on fruits more than 
90° excentric . 7. S. conferta 

13. Calyx during anthesis 1-2 mm 
high. 

15. Terminal bud 1-2 cm long. 


September 1972] 


Addenda, corrigenda et emendanda 


O20 


So231la 


Si 23 
5223240 


: 232b 
2334 


nn 


530233b 


5: 234b 


s256a 


5: 236b 


Stigma on fruit less than 90° 
excentric. 
5. S. oblongifolia 
15. Terminal bud !/2-1 cm long. 
Stigma on fruit usually more 
than 90° excentric. 
9. S. apiculata 
Santiria tomentosa BL. 
Add to literature: ANDERSON, 
Bull. Sing. 20 (1963) 165. 
In description, line 4, add after ‘pubes- 
cent’: to tomentose. Add under petioles, 
after ‘flattened’: or channelled. 
Add to Distr.: Philippines (Mindanao). 
Add to Ecol.: also in secondary forests. 
Santiria grandiflora KALKMAN. 
Add to Distr.: several new collections 
from Brunei. 
Santiria laevigata BL. 
Add to literature: ANDERSON, Gard. Bull. 
Sing. 20 (1963) 164. 
Add to description: Branchlets thin-hairy 
when young. Leaflets beneath sometimes 
sparsely hairy on midrib and nerves; 
base not rarely somewhat oblique; apex 
acutely to bluntly short-acuminate. 


Gard. 


Add to Distr.: Philippines (Mindanao). . 


Santiria oblongifolia Bu. 

Add to description: Buttresses up to 2m 
high. Petioles sometimes narrowly grooved 
at base. Fruits red when ripe. 

Santiria conferta BENN. 

Change in description: Tree, 4-35 m by 
up to 70 cm @&, withup to 1!/2 m high 
buttresses. Infructescences to 30 cm long. 
Fruits to 13/4 cm long, stigma lateral 
to near the pedicel. 

Add to Distr.: N. Borneo, at 1500-1800 
m alt. 

Delete 8. Santiria nervosa H. J. Lam. 
Santiria apiculata BENN. 

In description add at end of sentence on 
calyx: to sepals less than 1/2 connate. 
Petals inside glabrous or sparsely hairy. 
Add under fruits: red when ripe. 

Insert before the Key to the varieties: 
Note. A great part of the material can 
easily be subdivided into the following 
three varieties; however, some specimens 
show characters of more than one variety. 
var. apiculata. 

Change Ecol. highest alt. into 1500 m. 
var. rubra (RIDL.) KALKMAN. 

Add. to Ecol.: Primary and secondary 
forests up to 1600 m. 

Santiria megaphylla KALKMAN. 

In description, add at end of Ist sentence: 
by 45 cm @. Change in line 2 ‘11/2’ into: 
1, and in line 4 ‘5S’ into: 8. 

Line 8, after ‘part’ add: to all looped and 
joined. Delete after panicles “(3 un- 
known)’, the same after flowers. Insert 
before ‘Infructescences’: Pistillode in 
3 flowers minute. 

Add to Distr.: Brunei. 


5: 236a 
5: 236b 
59 2376 
5: 246 
58 WA 
5: 247a 
5: 2476 
5: 2476, 
567ab 


SeeZoil 


Add to Ecol.: In primary Dipterocarp 
forest on damp to swampy, shallow clay 
soil, 0-150 m. 

Santiria griffithti (HOOK. f.) ENGL. 

Add to synonymy: Amoora aphanamixis 
Auct. non R. & S.: Mia. Sum. (1861) 196. 
Change in description: Leaves excep- 
tionally to 15-jugate. Petioles sometimes 
strongly flattened at base. Leaflets 
exceptionally also hairy on midrib above 
and on veins beneath. 

Calyx (in vivo) olive to red. Petals (id.) 
yellowish-white. 

Change in Ecol. highest alt. into 700 m. 
Add to Notes: The collections For. 
Dept. Sarawak 12745 and 15613 repre- 
sent a strongly pubescent form with 
flattened petioles. 

Santiria rubiginosa BL. 

Change in description: Tree not always 
buttressed. Branchlets mostly glabrous. 
Add to Distr., under New Guinea: 
Vogelkop Peninsula. 

var. rubiginosa. 

Add to literature: ANDERSON, Gard. Bull. 
Sing. 20 (1963) 164. 

Scutinanthe THw. 

Line 2 from bottom, delete ‘pilose’. 
Key: A further difference between the 
two species is in the fruits, these being 
densely pubescent in S. brunnea, glabrous 
in S. brevisepala. 

Scutinanthe brunnea THW. 

Add to literature: WyYATT-SMITH & 
KOcHUMMEN, Mal. For. Rec. 17, rev. ed. 
(1965) 348. 

In description, add: Sometimes the 
leaves are fully glabrous. Change: 
Fruits finally glabrescent, yellow. 

Add to Ecol.: In Sarawak at c. 800 m, in 
Sabah at c. 1200 m alt. 

Scutinanthe brevisepala LEENH. 

Add to literature and synonymy: LEENH. 
Blumea 12 (1964) 19. Dacryodes 
papuana Husson, Blumea 7 (1952) 167, 
f. 1; Leenu. FI. Mal. I, 5 (1956) 228. 
Add to description: Tree up to 35 m 
high, 43-55 cm @, sometimes buttressed 
up to 3 m. Branchlets not always con- 
spicuously lenticellate. Leaflets lanceolate 
to broad-elliptic or subovate, up to 10 cm 
wide, coriaceous or pergamentaceous to 
chartaceous, the base broadly cuneate to 
subcordate, mostly slightly oblique. 
Inflorescences axillary on short axillary 
shoots the vegetative terminal bud of 
which usually develops later on. Ovary 
(2-)3-celled. Mature fruits — slightly 
oblique, ovoid to ellipsoid, constricted or 
shortly stalked at the base, pointed at the 
apex, 21/4-3 by 1!/2-13/4 cm, glabrous. 
Seed 1. 

Canarium STICKM. 

Add to Distr.: The genus can be sub- 
divided into 3 subgenera: subg. Canarium, 


922 


FLORA MALESIANA 


Sie2525 
SS, 


comprising the sections Canarium and 

Pimela; subg. Africanarium LEENH. nov. 

stat. (Canarium sect. Africanarium LEENH. 

Blumea 13, 1966, 396), monotypic, 

W. Africa; subg. Canariellum. 

In Key to the species, add to couplet 5, 

2nd lead: (in C. album sometimes 

papillose, then stamens connate halfway 
up or more, pistil pilose, and fruits white 
when ripe). 

Replace couplets 14 to 19 incl. by the 

following: 

14. Stipules fugacious, present only in 
the terminal bud, even the scars 
nearly invisible. 

15. Leaflets equal-sided at base. Pith of 
branchlets always with central 
vascular strands. Fruit very peculiar 
(see fig. 21k, in vol. 5), 4-41/2 by 
21/9-23/4 by c. 11/2 cm. 

48. C. cestracion 

15. Leaflets oblique at base. Pith of 
branchlets mostly without central 
vascular strands. Fruit ovoid to 
spindle-shaped, round in  cross- 
section, 21/2-31/2 by 11/2-2 cm. 

53. C. album 

14. Stipules persistent to caducous, scars 
well visible. 

16. Infructescences (sub)spicate, often 
with many fruits; fruits ovoid to 
subglobose, rather small (9-14 by 
4-11 mm) 51. C. asperum 

16. Infructescences racemoid to thyr- 
soid, mostly with few fruits; fruits 
mostly relatively longer and always 
bigger. 

17. Vascular strands in pith of branch- 
lets all peripherally arranged; 
twigs long remaining densely hairy 
(rarely, only one of these charac- 
ters holds good). 

18. Indumentum pilose. Flowers long 
and slender, corolla more than 
two times as long as the calyx; 
filaments in 2 flowers nearly 
completely connate. Fruits usual- 
ly prismatic, blunt 3-angular in 
cross-section, at apex mostly 
truncate and ‘shouldered’, (sub)- 
glabrous. 

33. C. pilosum ssp. pilosum 

18. Indumentum tomentose to velve- 
ty. Corolla less than two times 
as long as the calyx; filaments in 
© flowers free. Fruits ellipsoid, 
usually velvety. 

49. C. vrieseanum 

17. Vascular strands in pith of branch- 
lets only partly peripherally ar- 
ranged; twigs soon glabrescent. 

19. Stipules inserted on the petiole 
up to 3 cm from its base. Fruits 
fusiform, 4 by 2 cm. 

52. C. vitiense 


[ser. I, vol. 6® 


19. Stipules inserted on the twig at 
the base of the petiole. Fruits 
ellipsoid, up to 31/2 by 2 cm. 

19A. Leaflets rounded at base. 

Fruits 3-31/2 by 114/2-2 cm, 

pyrene smooth, sterile cells 
moderately reduced. 

44. C. macadamii 

19A. Leaflets cuneate at base. Fruits 

2 by 3/4 cm, pyrene irregularly 

grooved, sterile cells nearly 
completely reduced. 

45. C. chinare 


5: 254, Replace couplets 44 to 50 incl. by the 
255 following: 


44. Inflorescences axillary. 
45. Stipules rather caducous, roundish, 
+ herbaceous 17. C. luzonicum 
45. Stipules persistent, linguiform, stiff- 
coriaceous 18. C. ovatum 
44. Inflorescences terminal (lower 
branches often in the upper leaf 
axils). 
46. ¢ Flowers with 3 stamens. 

47. Vascular strands in pith of branch- 
lets all peripheral. Stipules cadu- 
cous, c.2mm @. 

5. C. caudatum f. caudatum 

47. Vascular strands in pith of branch- 
lets partly central. Stipules sub- 
persistent, c. 15 by 10 mm. 

54. C. reniforme 
46. 3 Flowers with 6 stamens. 

48. Stipules (rather) persistent, attach- 
ed on the petiole 1/2-11/2 cm from 
its base . : . «) 10. €. lamii 

48. Stipules caducous, mostly attach- 
ed on the twig at or partly, excep- 
tionally fully, on the base of the 
petiole. 

49. Stipules oblong, 1-5 by !/2-12/4 
cm; scar linear, c. 1/2 cm long. 

14. C. vulgare 

49. Stipules reniformous, much smal- 

ler; scar elliptic to drop-shaped, 

1—2 mm long. 

50. 3 Inflorescences very lax, bran- 

ches long and patent. J Flowers 

13 mm long. 6. C. divergens 

50. 3 Inflorescences not very lax. 

3 Flowers c. 8 mm long. 

50A. Veins and reticulations more 

or less prominulous and 

well-visible on the under 

surface of the leaflet; nerves 

11-15 pairs, mutual distance 

along the midrib usually less 

than 1 cm 1. C. littorale 

50A. Veins and reticulations nearly 

invisible in dried specimens; 

nerves 5-10-15 pairs, mutual 

distance along midrib 1-11/e 

cm . 4. C. patentinervium 


5: 255 Replace 63 Ist lead and couplet 64 by the 


following: 


September 1972] 


Addenda, corrigenda et emendanda 


a 250d 


5: 258a 


5: 258b 


5: 259a 


Be 2590 


5: 260a 


5: 260b 


63. Filaments at least nearly halfway 
connate. 

64. Branchlets long remaining densely 
ferruginous-woolly. Leaves up to 
8-jugate; nerves 14-17 pairs. Fruit- 
ing calyx c.5mm @. 40. C. kipella 
Branchlets glabrous or puberulous 
at the tip only. Leaves up to 
6-jugate; nerves mostly less than 
14 pairs. Fruiting calyx 8-11 mm @. 
64A. Leaflets lanceolate. Pith of 
branchlets always with central 
vascular strands. 

39. C. intermedium 
Leaflets (oblong to) broad-ellip- 
tic (to suborbicular). Pith of 
branchlets mostly without central 
vascular strands . 55. C. pimela 
Canarium littorale Bv. 
Add to literature: LEENH. Blumea 9 
(1959) 337; Back. & BAKH. f. Fl. Java 2 
(1965) 115; Meer, Bot. Bull. Herb. 
Sandakan 11 (1968) plate between p. 111 
and 112 (seedling). 
Add in Notes to the area of f. pruinosum 
(ENGL.) LEENH.: Brunei and Sabah. 
Canarium latistipulatum RIDL. 
Add to literature: LEENH. Blumea 9 


64. 


64A. 


(1959) 341. 
Add to description: Fruits  spindle- 
shaped, -+ round in_ cross-section, 


7 by 24/4 cm, glabrous; pyrene smooth, 
rounded triangular in cross-section, the 
lids intruded, lids c. 3-4 mm thick. 
Seeds 2, sterile cell moderately reduced. 
Canarium perlisanum LEENH. 

Add to literature: LEENH. Blumea 9 
(1959) 342. 

Canarium patentinervium MIQ. 

Add to literature: LEENH. Blumea 9 
(1959) 342, f. 15. 

Add to description: 
buttressed. /nfructescences 
glabrescent. 

Add to Ecol.: also in secondary and 
swamp forests. 


Tree sometimes 
sometimes 


Canarium caudatum KING. 

Add to literature: LEENH. Blumea 9 
(1959) 343. 

Add to Notes sub f. auriculiferum 


LEENH.: also known from the Malay 
Peninsula. 

Canarium divergens ENGL. 

Add to literature: LEENH. Blumea 9 
(1959) 346. 

Canarium kinabaluense LEENH. 

Add to literature: LEENH. Blumea 9 
(1959) 346. 

Add to description: Jnflorescences termi- 
nal, laxly thyrsoid, c. 4-6 cm long, 
few-flowered, minutely tomentose, gla- 
brescent. Flowers (2) 1 cm long. Calyx 
5 mm, minutely tomentose. Stamens 6, 
glabrous, inserted on the rim of the disk. 
Disk glabrous, adnate to the receptacle. 


Si 


5; 2615; 


an 


26la 


567a 


: 262a 
: 262b 


: 263a 


923 
Pistil glabrous. 
Add to Distr.: East Borneo. 
Add to Ecol.: down to 450 m. 
Canarium maluense LAUT. 
Add to literature: LEENH. Blumea 9 


(1959) 347. 

ssp. maluense. 

Add to description, Ist sentence: up to 
60 m high, with up to 11/2 m high 
buttresses. 

Add to Distr.: Louisiade Arch. 

Add: Uses. Timber for construction. 
Canarium megacarpum LEENH. 
Add to literature: LEENH. 
(1959) 351. 

Canarium lamii LEENH. 

Add to literature: LEENH. Blumea 9 
(1959) 351. 

The following changes should be made in 
the description: Tree up to 42 m by 66 
cm, sometimes with buttresses and stilt- 
roots. Leaves 3-4-jugate. Stipules sub- 
persistent or more or less caducous, 
inserted 1/2-11/2 cm from the base of 
the petiole. Leaflets up to 22 by 10 cm, 
base rounded to cordate; nerves 10—15 
pairs. Inflorescences (2) remotely spicate, 
densely tomentose. Flowers: old 2 known. 
Calyx 7 mm high, the lobes 11/2 mm; 
outside tomentose, inside appressed 
short-hairy, densest near base and 
margin. Corolla: petals 71/2 by 5 mm, 
outside densely appressed short-hairy 
in the upper half mainly along the midrib, 
inside glabrous, in vivo orange to red. 
Staminodes 6, inserted on the disk, c. 
31/24 mm long, glabrous. Disk adnate 
to the hollowed receptacle, free rim 
1/5-3/4 mm _ high, fleshy, glabrous. Pistil 
glabrous; ovary 3 mm; style 11/4 mm, 
thick; stigma globular, 1 mm @. 
Infructescences with 1 or few fruits, 
rusty tomentose; calyx to 2!/2 cm @, 
densely rusty tomentose when young. 
Add: Ecol. Primary and _ secondary 
forests up to c. 1250 m. 

Canarium sylvestre GAERTN. 

Add to literature: LEENH. Blumea 9 
(1959) 352. 

Add to description, Ist sentence: some- 
times with stiltroots. 

Add to Ecol.: also in secondary forests. 
Canarium piloso-sylvestre LEENH. 

Add to literature: LEENH. Blumea 9 
(1959) 353, f. 16. 

Add to description: Acumen of leaflets 
blunt to acute. Calyx 21/24 mm high, 
outside sometimes slightly pubescent 
towards the apex. Disk 3—6-lobed. 

Add to Ecol.: In seasonally inundated 
primary forest. 

Canarium salomonense B. L. BURTT. 
Add to literature: LEENH. Blumea 9 
(1959) 353. 

Change in description: Tree up to c. 


Blumea 9 


924 FLORA MALESIANA [ser. I, vol. 6° 
40 m by 80 cm, mostly with buttresses. LANESSAN, PI. Utiles Col. Frang. (1886) 
Canarium vulgare LEENH. 309, nom. nud. 

Add to literature: LEENH. Blumea 9 Add to description: Pith of branchlets 
(1959) 358; Back. & BAKH. f. Fl. Java 2 dark- to light-brown, either with peri- 
(1965) 115. pherally arranged, or with scattered 

5: 265a Add to Distr.: New Guinea. vascular strands. 

5: 265b Add to Ecol.: Alt. up to 1200 m. Add to Distr.: Borneo, Sabah (Beaufort 

5: 2665 Canarium indicum L. Distr., Ulu Lumat, SAN 44543). 

Add to literature: LEENH. Blumea 9 Add to Ecol.: Primary forest. 
(1959) 359; Back. & BAKH. f. Fl. Java 2 Canarium apertum H. J. LAM. 
(1965) 115. Add to literature: LEENH. Blumea 9 

5: 269b Canarium kaniense LAUT. (1959) 386. 

Add to literature: LEENH. Blumea 9 : 2756 Incertae sedis: The systematic position of 
(1959) 362. C. pseudodecumanum and C. decumanum 

5: 270b Canarium luzonicum (BL.) A. GRAY. remains uncertain, even with growing 
Add to literature: LEENH. Blumea 9 knowledge. The blastogeny is in full 
(1959) 363. accordance with sect. Canarium, the 
Add to Distr.: Mindanao. ontogeny of the stipules of C. decumanum 

5: 27la Canarium ovatum ENGL. with sect. Pimela, however. 

Add to literature: LEENH. Blumea 9 : 276a Canarium pseudodecumanum HOcnrR. 
(1959) 364. Add to literature: LEENH. Blumea 9 

5: 2716 Canarium odontophyllum Mia. (1959) 388: Meter, Bot. Bull. Herb. 
Add to literature: LEENH. Blumea 9 Sandakan 11 (1968) plate between p. 111 
(1959) 365. and 112 (seedling). 

5: 272a Add to Uses: In Sarawak grown for its : 277a Canarium decumanum GAERTN. 
fruits. Add to literature: DouGLas & BAAS 
Canarium denticulatum BL. BECKING, Bull. Bot. Gard. Btzg III, 17 
Add to literature: LEENH. Blumea 9 (1947) 295-296, t. 11; LEENH. Blumea 9 
(1959) 367; BAck. & BAKH. f. FI. Java 2 (1959) 389: BAcK. & BAKH. f. FI. Java 2 
(1965) 115. (1965) 114; Meer, Bot. Bull. Herb. 

5: 274a ssp. kostermansii LEENH. Sandakan 10 (1968) plate between p. 138 
Add to Distr.: Sabah. and 139. 

Canarium karoense H. J. LAM. Add to Distr. under New Guinea: 
Add to literature: LEENH. Blumea 9 Japen I. 

(1959) 370. : 278a Canarium oleosum (LAMK) ENGL. 
Canarium megalanthum MERR. Add to literature: LEENH. Blumea 9 
Add to literature: LEENH. Blumea 9 (1959) 391. 

(1959) 370, f. 17; Meer, Bot. Bull. Add to description, Ist sentence: Tree 
Herb. Sandakan 11 (1968) plate between up to 50 m high. 

p. 111 and 112 (seedling). : 27865 Add to Distr. under Lesser Sunda Is.: 
Add to description: Sometimes with up Sumbawa. 

to 1 m high buttresses. Stipules sometimes : 279a Canarium balsamiferum WILLD. 
caducous, sometimes inserted on the base Add to literature: LEENH. Blumea 9 
of the petiole or even slightly on the twig. (1959) 392. 

Leaflets often chartaceous, base some- : 279b Add to Distr.: Lesser Sunda Is. (Sum- 
times nearly equal-sided, margin some- bawa). 

times serrulate near the apex. : 280a Canarium trigonum H. J. LAM. 

5: 274b Canarium pseudopatentinervium H. J. Add to literature: LEENH. Blumea 9 

LAM. (1959) 393. 

Add to literature: LEENH. Blumea 9 Canarium euryphyllum PERK. 

(1959) 385. Add to literature: LEENH. Blumea 9 

Add to description: Buttresses sometimes (1959) 394. 

present. Leaflets: apex shortly blunt- to : 280b Add to description: Corolla reported 

acute-acuminate; nerves not to distinctly to be yellowish red. 

looped and joined. 3 Inflorescences with : 28la, Canarium kostermansii LEENH. 

up to 10 cm long lower branches, these 567b Add to literature: LEENH. Blumea 9 

as well as the main axis laxly set with (1959) 398, f. 23. 

subsessile glomerules of flowers. 3 Flowers Add to description: Leaflets: acumen 

3-4 mm long pedicelled. Disk in 3 acute to bluntish; nerves 20-25 pairs. 

flowers cushion-shaped, c. 1 mm high, Add: Ecol. Primary forest. 

densely hairy, without rudimentary : 2816 Canarium pilosum BENN. 

pistil. Add to literature: LEENH. Blumea 9 
5: 275a Canarium grandifolium (RIDL.) H. J. LAM. (1959) 398. 


Add to literature: LEENH. Blumea 9 
(1959) 386, f. 21, non BAILLON ex DE 


: 282a 


ssp. borneensis LEENH. 
Add to Distr.: Sarawak (G. LAMBIR). 


September 1972] 


Addenda, corrigenda et emendanda 


5: 282b 


5: 283a 


5: 2836 


5: 284b 


5: 285a 


Si 285b 


5: 286a 


5: 286b 


5: 287a 


5: 289a 


5: 289b 


Add to Ecol.: from the lowland up to c. 
1500 m. 
Canarium merrillii H. J. LAM. 


Add to literature: LEENH. Blumea 9 
(1959) 402. 

Canarium gracile ENGL. 

Add to literature: LEENH. Blumea 9 


(1959) 396, f. 22. 

Add to description: The flowers are said 
to be purplish, the fruits first red, when 
ripe black. 

Canarium dichotomum (BL.) Mia. 

Add to literature: LEENH. Blumea 9 
(1959) 423. 

Add to description: Leaflets sometimes 
beneath rather densely appressed short- 
hairy; acumen sometimes slender and 
acute. 2 Inflorescences up to 35 cm long. 
Add to Ecol.: Sometimes also in secon- 
dary forests. Alt. up to 1000 m. 
Canarium fusco-calycinum RIDL. 

Add to literature: LEENH. Blumea 9 
(1959) 424. 

Add to description: Nerves geniculate or 
looping near the margin. 

Add to Ecol.: Primary lowland Diptero- 
carp forest at 100 m. 

Canarium australianum F.v.M. 

Add to literature: LEENH. Blumea 9 
(1959) 421; SpecHT, Rec. Am. Austr. 
Exp. Arnhem Land 3 (1958) 460. 

Add to description: Leaflets sometimes 
oblong. 

Add to Distr. under Australia: 
northern part of W. Australia. 
Canarium intermedium H. J. LAM. 
Add to literature: LEENH. Blumea 9 
(1959) 419, f. 29. 

Canarium kipella (BL.) Mia. 

Add to literature: LEENH. Blumea 9 
(1959) 419; Back. & BAKH. f. FI. Java 2 
(1965) 115. 

Canarium pseudosumatranum LEENH. 
Add to literature: LEENH. Blumea 9 
(1959) 431. 

Change in description: Leaves 6—9-juga- 
te; leaflets 7-20 by 3-6 cm. /nflorescences 
20-30 cm long. 

Canarium sumatranum BOERL. & KoorD. 
Add to literature: LEENH. Blumea 9 
(1959) 432, f. 31. 

Add to description: Leaflets up to 23 cm 
long, rough above; nerves up to more 
than 30 pairs. 2 Inflorescences \ike the 
3 ones. 2 Flowers 6 mm long, the calyx 
21/2 mm. 

Canarium hirsutum WILLD. 

Add to literature: LEENH. Blumea 9 
(1959) 424, f. 30; Back. & BAKH. f. 
Fl. Java 2 (1965) 115. 


the 


Canarium hirsutum WILLD. var. hirsu- 
tum. 

Add to Distr.: Solomon Is. 

Canarium macadamii LEENH. — Fig. 2. 


Add to literature: LEENH. Blumea 9 


Fig. 2. Canarium macadamii LEENH. a. Staminodes 
and disk of & flower from outisde, b. & c. stamens 
and disk of 3 flower from outisde (6) and on 
longitudinal section (c: mind the tiny pistillode), 


all » 


5: 290a 


5: 2906 


6 (a NGF 21682, b-c MACADAM 206). 


(1959) 448. 

Add to description: Tree 30-40 by 
3/,-1 m. Leaflets 10-20 cm _ long. 
2 Inflorescences 8-10 cm. ¥% Flowers 


7 mm; calyx 4 mm; petals outside 
sparsely hairy; staminodes confluent 
with the disk; disk glabrous, protruded 
into 6 tongue-shaped, fleshy, 2 mm long 
lobes alternating with and nearly equal- 
ling staminodes. Infructescences 8-15 cm 
long; calyx 5-7 mm @. 

Add to Ecol.: Alt. 1100-1500 m. FI. 
Febr. 

Add: Uses. Seeds edible. 

Canarium chinare GRUTTERINK & H. J. 


LAM. 
Add to literature: LEENH. Blumea 9 
(1959) 449. 


Canarium rigidum (BL.) Mia. 

Add to literature and synonymy: LEENH. 
Blumea 9 (1959) 434, f. 32. — C. poly- 
phyllum K. Scu. in K. Sch. & Hollr. 
Fl. Kais. Wilh. Land (1889) 63; LEENH. 
Fl. Mal. I, 5 (1956) 290, f. 20 d; Blumea 9 
(1959) 434, f. 33. 

Add to description: Tree up to 30 m by 
40-45 cm. 2 Inflorescences 8-9 cm long, 
the partial inflorescences c. 2 cm long, 
with 1 or 2 flowers. 2 Flowers 71/2 mm 
long, the calyx 4!/2 mm high, outside 
densely appressed pubescent; staminodes 
free outside the disk, 2 mm long; disk 
faintly lobed, c. 1 mm high, long-hairy 
towards the margin; pistil glabrous. 
Add: Uses. Reported to be used for 
buildings proas. 

Canarium polyphyllum K. SCH. 

With new collections that became avail- 
able it became more and more clear that 
the present species and C. rigidum repre- 
sented only extremes as to hairiness, 
leaf ratio, and sculpture of fruit kernel, 
but could not well be demarcated. Hence, 
C. polyphyllum had to be reduced to 
C. rigidum. 

Canarium cestracion LEENH. 

Add to literature: LEENH. Blumea 9 
(1959) 436. 

Change in description: Vascular strands 
in pith of branchlets may also be mainly 
peripherally arranged. Infructescences 


926 


FLORA MALESIANA 


[ser. I, vol. 6% 


Se 29a 


55 29 0D 


293a 


5: 295b 


5: 296a 


5: 2965 


5: 2966 


3-20 cm long. Fruits, if 1-seeded, much 
flattened in cross-section. 

Add to Distr.: A second collection 
(NGF 25654) from Morobe Distr., at 
30 m alt. 

Canarium vrieseanum ENGL. 

Add to literature: LEENH. Blumea 9 
(1959) 437. 

Canarium acutifolium (DC.) MERR. 

Add to literature: LEENH. Blumea 9 
(1959) 445. 

var. acutifolium. 

Change in description: Tree 13-45 m. 
Leaflets up to 40 by 15 cm. Fruiting calyx 
up to 74/2 mm @. Fruits up to 171/2 by 
121/2 (1-seeded) or 171/2 (2-seeded) mm. 
Seeds exceptionally 2. 

Add to Distr.: New Britain. 

Canarium asperum BENTH. 

Add to literature: LEENH. Blumea 9 
(1959) 439; Back. & BAKH. /. Fl. Java 2 
(1965) 115. 

SSP. aSperum var. asperum. 

Add to Uses: On wood see SCHNEIDER, 
Bull. Bur. For. Philip. 14 (1916) 129. 
Replace the heading of 52. Canarium 
schlechteri Laut. by the following: 

52. Canarium vitiense A. Gray, U.S. 
Expl. Exp. Bot. 1 (1854) 373; LEENH. 
Blumea 9 (1959) 451; Blumea 13 (1965) 
166. — C. samoense ENGL. in DC. Mon. 
Phan. 4 (1883) 134; LEENH. Blumea 9 
(1959) 452. — C. schlechteri LAvuT. Bot. 
Jahrb. 56 (1920) 328; LEENH. FI. Mal. 
I, 5 (1956) 296, f. 20 e; Blumea 9 (1959) 
444. — C. smithii LEENH. Bish. Mus. 
Bull. 216 (1955) 12, f. 6; Blumea 9 (1959) 
450. — C. hacciferum LEENH. Bish. Mus. 
Bull. 216 (1955) 19, f. 9. Add to de- 
scription: 2 Inflorescences racemose, 6—9 
cm long, tomentose. 2 Flowers 1 cm long, 
less slender than ¢ ones, subsessile; 
calyx 4 mm; staminodes 3 mm, connate 
for less than 1 mm; disk annular, low, 
long-ciliate; pistil 4 mm, glabrous, ovary 
tapering into the style. 

Replace Distr. by: Malesia: eastern half 
of New Guinea, Louisiade Arch., Admi- 
ralty Is., Bismarck Arch., Solomons, 
Fiji, Samoa, and Tonga. 

After species 52 the following species 
should be added: 


53. Canarium album (Lour.) RAEUSCH. 
Nomencl. ed. 3 (1797) 287; Hance, J. 
Bot. 9 (1871) 38; GUILLAUMIN, Bull. Soc. 
Bot. Fr. 55 (1908) 617, t. 19 f. 1; HAYATA, 
J. Coll. Sc. Imp. Univ. Tokyo 30 (1911) 
52; Merr. Int. Rumph. (1917) 304; 
WALKER, Imp. Trees Ryukyu (1954) 148, 
f. 82; LEENH. Blumea 9 (1959) 402, f. 24; 
BARANOV, Quart. J. Taiw. Mus. 20 (1967) 
367, cum fig.; non BLCo, FI. Filip. (1837) 
793 (= C. luzonicum A. GRAY). — ? C. 
sinense Cana Rumen. Herb. Amb. 2 


(1741) 154. — Pimela alba Lour. FI. 
Coch. (1790) 408. 

Tree, up to 30m by 1!/2 m &. Branch- 
lets 5-6 mm @&, the young parts fulvous 
tomentose, older parts sometimes gnarly 
by the strongly prominent scars of leaves 
and inflorescences; pith with a peripheral 
cylinder of small vascular strands, rarely 
moreover some strands in the central part. 
Leaves 3—6-jugate. Stipules present in the 
bud only, inserted on the twig next to 
the petiole, even the scar usually nearly 
invisible. Leaflets usually distinctly sinu- 
ous, especially the basal pairs, lanceolate 
or elliptic (to ovate), 61/2-14 by 2-5!/s 
cm, chartaceous to coriaceous, glabrous 
or with some scattered bristles on the 
nerves below, lower side often minutely 
verrucose; base oblique, cuneate to 
rounded; margin entire; apex tapering to 
subabruptly acuminate, acumen up to 
2 cm long, blunt; nerves 12-16 pairs, 
angle to midrib 65-75”, straight to faintly 
curved in the broader, strongly curved 
in the narrower side of the leaflet, more 
or less distinctly looped and joined near 
the margin; intercalary veins sometimes 
distinctly developed; reticulation dense. 
Inflorescences axillary, tomentellous to 
glabrous, 3 thyrsoid, 15-30 cm long, 
many-flowered, 2 racemoid, 3-6 cm 
long, with up to 12 flowers. Flowers 
sparsely tomentose to glabrous, 3 
51/2-8 mm long, 2 c. 7 mm. Calyx 21/2-3 
mm, in 2 flowers subtruncate. Stamens 6, 
glabrous, the filaments more than halfway 
(in 2 flowers up to nearly completely) 
connate. Disk 3 globular to cylindrical, 
1-11/2 mm high, slightly 6-lobed, solid 
or with a central canal, the upper side 
with some bristles; in 2 flowers annular, 
faintly 3-lobed, 1 mm high, thick and 
fleshy, pilose on the inner surface. Pistil 
densely short-pilose, in flowers minute 
or none. Infructescences up to 15 cm long 
with up to 6 fruits; calyx flat, 3-lobed, 
1/5 cm @, the lobes recurved. Fruits 
ovoid to spindle-shaped, round in 
cross-section, 21/2-31/2 by 11/2-2 cm, 
glabrous, in vivo white when ripe; 
pericarp rather thick; pyrene acuminate, 
rounded (to 6-angular) in cross-section, 
with a distinct groove between the blunt 
angle-ribs and the lids, the latter with a 
faint median rib, surface furthermore 
slightly undulated; lids 11/2-2 (3) mm 
thick. Seeds 1 or 2, the sterile cell(s) 
slightly reduced, round in cross-section. 

Distr. Annam (from about 16° N 
northwards), Tonkin, southern China (up 
to about 27° N), and Hainan; as thespecies 
is much cultivated, mainly in the same 
region, it is sometimes difficult to decide 
whether in some part of the area it is 
wild or naturalized, or even planted. In 


September 1972] 


Addenda, corrigenda et emendanda 927 


Malesia: Sumatra, East Coast (Medan, 
planted in and naturalized near a park: 
LORZING 16519, 17240). 

Ecol. In light to dense forests on dry to 
moist soils, usually at medium altitude 
(400-1200 m). 

Uses. Especially in SE. China common- 
ly planted for ornament and as a fruit tree. 
The fruits, of which the pulp as well as 
the seeds are eaten, prepared in several 
ways, are highly esteemed by the Chinese. 
Furthermore, the wood and the resin are 
sometimes used. 

Note. C.albumbelongs to sect. Pimela, 
to the relationship of C. pilosum, and 
seems to be nearest to C. pimela. 


54. Canarium reniforme KOCHUMMEN & 
Wuitmore, Gard. Bull. Sing. 24 (1969) 2. 

Tree, up to 18 m by 30cm @. Branchlets 
5-7 mm @, long remaining fulvous- 
tomentellous; pith with many vascular 
strands, partly peripheral. Leaves 2-4-ju- 
gate. Stipules attached on the base of the 
petiole, mostly partly on the twig, oblong 
to reniform, 15-25 by c. 10 mm, rather 
stiff, persistent. Leaflets ovate to ovate- 
oblong or elliptic, 31/2-16 by 11/2-10 cm, 
stiff-coriaceous, beneath thinly puberu- 
lous, further glabrous; base of laterals 
slightly oblique, cuneate to rounded, 
margin entire, apex rather abruptly, 
bluntly to acutely acuminate; nerves 
7-14 per side, angle to midrib c. 60-70°, 
nearly straight to curved, looped and 
joined near the margin, veins and vein- 
lets much more slender, but well-visible 
on both sides. Inflorescences terminal, c. 
25 cm long. Flowers: 2 unknown. Calyx 
5 mm high, outside puberulous, inside 
glabrous. Petals outside hairy in the 
central part. Stamens 3, adnate to the 
disk. Disk solid, globular, tapering into 
a ‘style’, glabrous. J/nfructescence c. 
10 cm long, glabrous; calyx flat, orbic- 
ular, 1-11/2 cm ©, with inside an annu- 
lar, not-lobed, glabrous disk. Fruits ovoid 
to ellipsoid, 5—51/2 by c. 23/4 cm, in 
cross-section blunt-triangular; pyrene 
rather smooth; lids 3-4 mm thick. Seed 1, 
the fertile cell orbicular in cross-section, 
the sterile ones nearly completely redu- 
ced. 

Distr. Malesia: the Malay Peninsula 
(Perak). 

Ecol. Understorey of primary hill- 
forest at c. 300 m alt. Fr. June. 

Note. C. reniforme belongs to sect. 
Canarium and is closely allied to C. 
patentinervium and C. caudatum. With 
the latter it shares the number of 3 
stamens, with the former the general 
habit, apart from the peculiar stipules. 


55. Canarium pimela LEENH. Blumea 9 
(1959) 406, f. 25. — ? C. sinense Tsjacana 


RumpuH. Herb. Amb. 2 (1741) 154. — 
Pimela nigra Lour. Fl. Coch. (1790) 
407. — C. pimela Koen. Ann. Bot. 1 
(1805) 361, t. 7, f. 1, nom. illeg.; HANCE, 
J. Bot. 9 (1871) 38; Fors. & HEMSL. 
J. Linn. Soc. Bot. 23 (1886) 113; Merr. 
Int. Rumph. (1917) 304; Merr. & CHUN, 
Sunyatsenia 2 (1935) 253; Merr. Comm. 
Lour. (1935) 227; non BL. Bijdr. (1826) 
1162 (= C. kipella), nec SPAN. Hook. 
Comp. Bot. Mag. 1 (1835) 346 (= C. 
oleosum), nec Bico, FI. Filip. (1845) 545 
(= C. asperum). 

Tree, up to 30m by 11/2m @. Branch- 
lets 7-10 mm 2, glabrous; pith with a 
peripheral cylinder of vascular strands 
and sometimes a few in the central part. 
Leaves 4-6-jugate, glabrous. No stipules. 
Leaflets oblique, often distinctly falcate, 
broad-elliptic (to ovate or suborbicular, 
rarely oblong), 6-17 by 2—71/2 cm, charta- 
ceous to coriaceous; base acute, often 
decurrent; margin entire; apex rather 
abruptly acuminate, acumen _ short, 
broad, and blunt; nerves (8—) 11 (—15) 
pairs, angle to midrib 70—75°, straight 
to faintly curved, looped and + joined 
near the margin; veins and veinlets 
coarsely reticulate. Inflorescences axillary, 
glabrous, laxly thyrsoid (¢) to race- 
moid (2), 15-40 cm long, ¢ many-, 
© few-flowered. Flowers (sub)glabrous, 


3S 7 mm long, slender, 2 9 mm long. 


Calyx in 3 flowers 21/2 mm, distinctly 
lobed, in 2 flowers 31/24 mm, subtrun- 
cate. Corolla in 2 buds characteristic- 
ally conical. Stamens 6, glabrous (except 
2 rows of bristles on the anthers in 2 
flowers), in dj flowers nearly halfway, in 
2 flowers slightly more than halfway 
connate. Disk annular, 1/2-1 mm high, 
fimbriate, in 3 flowers thick-fleshy with 
a narrow central canal, in 2 flowers thin, 
slightly 6-lobed. Pistil glabrous, in ¢ 
flowers absent. Infructescences 8-35 cm 
long, lax, with 1-4 long-stalked fruits; 
calyx nearly flat, faintly triangular to 
suborbicular, 8-10 mm @. Fruits 
narrowly ovoid, 34 by 11/4-2 cm, round 
to slightly triangular in cross-section; 
pericarp thin; pyrene smooth or with 
a faint median rib on the lids; lids c. 
3 mm thick. Seeds 1 or 2; fertile cell(s) 
usually with a distinct adaxial rib, sterile 
cells moderately reduced. 

Distr. SE. China (from about 27° N 
southwards), Hainan, and Indo-China; 
as the species is much cultivated, espe- 
cially in southern China and Tonkin, it 
is not well possible to draw the natural 
limits of its present area of distribution. 
In Malesia: Borneo (Sarawak, near 
Belaga, JAcoss 5239). 

Ecol. Indense to open forests, usually at 
medium altitude; in Malesia in primary 


928 


FLORA MALESIANA 


[ser. I, vol. 68 


358296 


6: 


6: 


121b 


122 


hill-forest below 500 m. Fi. Aug. 

Uses. Planted as an ornamental and 
especially as a fruit tree. The fruits are 
highly esteemed among the Chinese; they 
are comfited or pickled. Only the pulp 
is eaten. The wood and resin are some- 
times used, but are of no great value. 

Notes. C. pimela belongs to sect. Pime- 
la, to the relationship of C. pilosum, with 
C. album and C. parvum LEENH. (N. Viet- 
nam) as its possibly nearest allies. 

For the complicated nomenclature of 
the present species see LEENH. /.c. 

Add to Excluded genera: 
Nothoprotium Mia. Sum. (1861) 527 = 
Pentaspadon (Anacardiaceae); reduction 
already made by MARCHAND, Reéy. 
Anacard. (1869) 90, 183. 


Campanulaceae 


Sphenoclea zeylanica GAERTN. 

Add. to Distr.: Northern Territory of 
Australia. Cf. BLAKE, Austr. J. Bot. 2 
(1954) 137. 

Codonopsis lancifolia (ROXB.) MOELIONO 
ssp. lancifolia. 

Add to synonymy: Canarina moluccana 
Roxs. [Hort. Beng. 87] Fl. Ind. ed. 
Carey 2 (1832) 173; ed. Clarke (1872) 
298. Cf. STEEN. Nova Guinea, Bot. 12 
(1963) 191. 

No specimens or drawing could be 
traced, but the brief description is clear. 
ROXBURGH had this species twice in his 
Flora, but the Moluccan one had 6-mer- 
ous flowers and was therefore arranged 
in another Linnean class. 

In his monograph of Canarina, HED- 
BERG erroneously dismissed the Rox- 
BURGH name as a nomen nudum (Svensk 
Bot. Tidskr. 55, 1961, 19). 

Add to the species of Lobelia: 


Lobelia donanensis VAN ROYEN, Kew Bull. 
20 (1966) 305, f. 1. 

Small herb, witha glabrous, up to 12mm 
long stem. Leaves alternate, 11/2-21/2 
cm long; limb ovate or ovate-elliptic, 
7-12 by 5-7 mm, rounded or broadly 
cuneate at base, wavy along margins 
and with distinct, pear-shaped glands, 
obtuse or rounded at apex, coriaceous 
when alive, glabrous on either side, pilose 
along margin; petiole up to 11 mm, 
pilose in apical part only. Flowers choco- 
late-coloured, axillary and _ terminal; 
pedicels up to 6 mm, glabrous. Calyx 
21/23 mm long; lobes lanceolate-linear, 
1—11/2 mm long, usually with 3 glands on 
the teeth along the margin, pilose on 
outside. Corolla 5-6 mm long; tube 
2-21/2 mm, pilose on outside, dorsally 
split to 1 mm from the base; free part of 
all lobes c. 3 by 3 mm, acutely acuminate; 


Gemlssy 
Geas7 
6: 141 


ventral lobes on inside with dark purple 
papillae and near the throat also with 
3 or 4 longitudinal crests. Stamens c. 
3 mm, entirely connate; filaments glab- 
rous; anterior anthers 11/2 mm long, the 
posterior ones c. 1 mm; all connectives 
papillate, not barbate at tips. Ovary glob- 
ose, c. | mm @, glabrous; style glab- 
rous; stigmas dark purple, papillate 
along the margins. Capsule globose, 2!/2- 
3 mm @, glabrous, thin-walled. Seeds 
obovoid, c. 0.7 mm long, subtriangular 
in cross-section, smooth. 

Distr. Malesia: East New Guinea 
(Milne Bay Distr., Maneau Range, 
Mt Donana), one collection. 

Ecol. Between dead moss in open 
grassland on limestone, 2250 m. Fi. fr. 
Aug. 

Notes. According to the author allied 
to L. archboldiana, but differing in the 
gland-bearing leaf margin, the corolla 
lobes which are papillose inside and with 
longitudinal crests near the tube, the 
papillose connective, and glabrous ovary. 

At Leiden we have no material of either 
species and these data are not checked. 
Lobelia borneensis (HEMSL.) MOELIONO. 
Add to Distr.: Also in Flores (Lesser 
Sunda Is.). Cf. STEEN. Blumea 15 
(1967) 153. 

Replace Phyllocharis DieELs, 1917, non 
FéE, 1824, by: Ruthiella STEEN. Blumea 13 
(1965) 127, and the species names of the 
four species of this Papuan genus on 
p. 137-139 by: 

1. Ruthiella oblongifolia (DrieELS) STEEN. 
Ee 

2. Ruthiella schlechteri (DIELS) STEEN. /.c. 
3. Ruthiella subcordata (MERR. & PERRY) 
STEEN. /.c. 

4. Ruthiella saxicola (VAN ROYEN) STEEN. 
hes 

Bottom line: Replace Legousia speculum- 
veneris (L.) Fiscu. by: Specularia specu- 
lum-veneris (L.) CARUEL (1888). 

The generic name Specularia HEIST. ex 
Fase. Enum. Pl. Hort. Helmst. (1759) 
151, nom. valid.; ed. 2 (1763) 225, descr., 


has. distinct priority over Legousia 
DuRAND, Fl. Bourg. 1 (1782) 37; 
2 (1782) 26. 


Caprifoliaceae (VAN STEENIS & KERN) 


4: 175 


4: 194 


In this family we have omitted to mention 
any cultivated species, as none were 
known outside scientific botanic gardens, 
as far as collections were made. It was 
overlooked that in the former century 
KorTHALs had collected one which was 
described by MIQUEL as a new species 
from Java. Add to: 


Formerly cultivated 


September 1972] 


4: 178a 


4: 181 


4: 182 


Weigelia coraeensis THUNB. Trans. Linn. 

Soc. 2 (1794) 331; Hara, En. Sperm. 

Jap. 2 (1952) 63; STEEN. Blumea 13 

(1965) 167. — Weigelia fallax Mia. 

Fl. Ind. Bat. 2 (1856) 128. — Diervilla 

fallax (Miq.) BorrL. Handl. FI. Ned. 

Ind. 2 (1891) 6. 

Note. Native in Kyushyu I., Japan, but 
rare in cultivation. This may well have 
been an original import by the East 
India Company, via the Dutch Settle- 
ment in Deshima, in VON SIEBOLD’s time. 
In the Catalogue of TEYSMANN & BINNEN- 
DIJK (1866) two Diervillas are mention- 
ed to be cultivated, but this was appar- 
ently unsuccessful and the genus is ab- 
sent in the Catalogue of the Bogor Bota- 
nic Gardens of 1930. 

Lonicera malayana HENDERSON. 

Of this extremely rare species two new 

collections have come in, both collected 

on the Selangor/Pahang Gentinh new 

access road, on steep hillside, 1000 m 

(FRI 3882 & 4519 WuitmMore, ff. and fr. 

respectively). 

Add to description: Big woody climber, 

stem 2!/2 cm @, reaching to 20 m up a 

tree; leafy branches scattered all the way 

up. Fruit globular, c. S-8 mm @, black, 
juicy. Seeds 1-4, c.4 mm long, + convex, 
surface wrinkled. 

Note. In both specimens the condensed 
inflorescence is terminal, without axillary 
stalks. 

Line 35 from bottom. Delete from the 

species of which the flowers are unknown 

V. amplificatum and V. clemensae. 

Replace lines 5-26 and complete the key 

for flowering specimens as follows: 

11. Corolla not distinctly tubular, tube 
less than 3 times as long as lobes, the 
latter often more than 1 mm long. 

14. Inflorescence shortly pyramidal, 
paniculate. Filaments 1-3 mm long. 
Tube of corolla 1/4-1 mm long. 

15. Leaves entire, glandular-pitted 
beneath in the axils of the primary 
and secondary side-nerves. 

14. V. clemensae 

15. Leaves crenate-dentate. 

16. Leaves thinly coriaceous. Corolla 
2-2!/2 mm long. Filaments in- 
serted at the base of the corolla. 
(Between 500 and 1500 m altitude, 
sometimes up to 2300 m.) 

10. V. lutescens 

16. Leaves manifestly coriaceous. 
Corolla 3 mm long. Filaments 
adnate to the corolla !/2-1 mm 
above the base. Without fruits 
hardly distinguishable from V. 
lutescens. (Altitude at least 2300 
m.) . . . 21. V. junghuhnii 

14. Inflorescence corymbiform. Fila- 
ments at least 6 mm long. Tube of 


Addenda, corrigenda et emendanda 929 


the corolla usually exceeding 1 mm. 
17. Filaments in bud with inflexed top, 
6 (-7) mm long. 

18. Corolla broadly tubular, obovoid 
in bud, tube about 21/2 mm, lobes 
about 1!/2 mm. Leaves obtuse or 
shortly and bluntly acuminate. 

3. V. glaberrimum 

18. Corolla shortly tubular-turbinate, 
globular in bud, tube about 2 mm, 
lobes 1!/2-2 mm. Leaves gradu- 
ally long-acuminate. 

4. V. platyphyllum 
17. Filaments serpentine in bud, (7-) 
8-10 mm long. 

19. Underside of leaves with distinct 
glandular pits at the base on both 
sides of the midrib and often 
smaller ones in higher nerve-axils. 
Corolla turbinate, tube 21/2-3 
mm, lobes 11/2-2 mm. 

9. V. vernicosum 

19. Leaves without glandular pits. 
Corolla rotate-cupular, tube 1 
(-11/2) mm. 

20. Leaves up to 26 by 14 cm, 
glabrous to softly villous be- 
neath. Corolla lobes 11/2 mm 
long . . 12. V. amplificatum 

20. Leaves 12-17 by 8-9 mm, 
hispidulous beneath. Corolla 
lobes 2—2!/2 mm long. 

8. V. hispidulum 


4: 1885 12. Viburnum amplificatum Kern, Reinw. 


1951) 1508. 8 BIS Mailed: 4: (1951) 
188; Sarawak Mus. J. 9 (1960) 679, 
f. 1. — Descr. emend. — Fig. 3. 

Small tree up to 15m, sometimes shrub- 
like. Leaves more or less coriaceous, dull, 
dark olivaceous above, brownish green 
or brown beneath, glabrous above, 
glabrous to softly villous with simple, 
forked, and stellate hairs beneath, neither 
glandular-pitted at the base nor bearded 
in the nerve-axils, elliptic-oblong, ovate 
or obovate, up to 26 by 14 cm; apex 
abruptly short-acuminate (acumen rather 
blunt, 1/2-1 cm); base cuneate to some- 
what rounded, slightly decurrent on the 
petiole; margin entire, sometimes remote- 
ly and obscurely undulate; nervation 
indistinct above, prominent beneath; 
primary nerves 4~7 on either side of the 
midrib, anastomosing; petioles 2-4 cm. 
Inflorescence umbellate, corymbiform, 
up to 20 cm across; axes stellately pubes- 
cent to subglabrous; peduncle stout, up 
to 10 cm; primary rays up to 8, up to 
10 cm. Flowers small, c. 4 mm wide. 
Calyx-limb distinctly 5-lobed; lobes 
triangular, glandular-ciliolate, c. 1 mm. 
Corolla globular in bud, rotate-cupulate, 
glabrous, white; tube 1 mm, lobes ovate, 
c. 11/2 mm. Stamens inserted at the base 


FLORA MALESIANA 


[ser. I, vol. 68 


Fig. 3. Viburnum amplificatum KERN. a. Flower 
bud, x5, b. stamen in bud, 13, c. expanding 


flower, 


x 61/2, d. calyx, e. glands on margin of 


calyx lobe, f. part of corolla with stamen, x5, 
g. anther, x 13, A. fruit, 2, i. ditto, in cross sec- 


6: 230 


tion (a—i KOSTERMANS 13864). 


of the corolla, much exserted; filaments 
serpentine in bud, 7-8 mm; anthers 
oblong, 1 mm. Ovary cylindrical, glabrous, 
11/2 mm. Drupe oblong, sometimes very 
slightly dilated upwards, much flattened, 
with a distinct groove on both sides, at 
first green, twining black, 15-16 by 6-7 
mm. Endocarp with a broad longitudinal 
groove on the dorsal side, the incurved 
edges forming a deep, broad, in cross- 
section bilobed furrow on the ventral 
side. 

Distr. Malesia: N. and E. Borneo. 

Ecol. Primary forests, up to 600 m. 


Celastraceae (DING Hou) 


Add before Taxonomy: 

Palynology. An important study of 
the pollen of seven genera of Malesian 
Celastraceae was published by DING 
Hou, Blumea 17 (1969) 97-112, 1 fig., 
8 pl., of which the main results with 
relation to affinity and generic distinc- 
tion are the following: 

(i) The distinction of Kokoona and 
Lophopetalum is corroborated by the 
difference in pollen types. 

(ii) In Lophopetalum at least four pollen 
subtypes can be distinguished. 


(iii) Pollen of Sarawakodendron, de- 


6: 


25ie 


232 


: 2416 


: 243a 


: 256 


260, 
264 


: 2665 


: 2685 


31391 


: 392 


39392 


scribed below, shows a great resemblance 
to that of the related genera Kokoona and 
Xylonymus. 

(iv) Pollen of Hedraianthera and Bras- 
siantha resemble that of the three latter 
genera but shows also resemblance to 
that of the African genus Salacighia. 
Unfortunately the numbering of the 
genera is erroneous; in the sequence of 
the first key it must be: 

9. Bhesa, 1. Celastrus, 2. Maytenus, 
3. Xylonymus, 12. Perrottetia, 8. Micro- 
tropis, 4. Euonymus, 5. Glyptopetalum, 
6. Kokoona, 7. Lophopetalum, 10. Cas- 
sine, 11. Pleurostylia. 

Maytenus emarginata (WILLD.) DING 
Hou. 

Add to Distr.: Lesser Sunda Is. (Flores). 
Maytenus diversifolia (MaAxImM.) DING 
Hou. 

Add to Distr.: Flores. 


: 248ab Euonymus cochinchinensis PIERRE. 


Add to Distr.: Burma, Formosa, and 
Flores. 
Glyptopetalum Taw. 
Add to Distr. (in China); Kweichow. Cf. 
Dinc Hou, Blumea 12 (1963) 59. 
BALAN MENON (Mal. For. 27, 1964, 
18-21) confirmed the generic difference 
between Lophopetalum and Kokoona on 
their wood anatomy. 
Lophopetalum WIGHT ex ARN. 
Add to Distr.: A new species of the 
genus, the first from Australia, has been 
discovered by Mr. ByRNEs in the Kimber- 
ley District; it is related to the West 
Malesian species. 
Lophopetalum floribundum WIGHT. 
Add to Distr.: Two additional collections 
from Johore (FRI 8843) and Pahang 
(FRI 8147). 
Lophopetalum macranthum (LoEs.) DING 
Hou. 
Add to Distr.: A good flowering speci- 
men collected from East New Guinea 
(Morobe Distr.: NGF 37402). 
The new genus Sarawakodendron (re- 
cently found in Borneo) keys out for 
flowering material to Salacia. 
For fruiting material Sarawakodendron 
keys out at fork 8, which can be improved 
as follows: 
8c. Fruits spindle-shaped, 3-angled, 3-cel- 
led, 3-valved. Seeds with a caruncle- 
like aril surrounded by many 
filamentous, fringed appendages. 
3a. Sarawakodendron 


3a. SARAWAKODENDRON 


Dinc Hou, Blumea 15 (1967) 141. — 
Fig. 4. 

Small tree, containing kautchuk, Leaves 
alternate. Stipules small, caducous. Jn- 
florescences solitary, axillary, simple, pe- 


September 1972] Addenda, corrigenda et emendanda 931 


Fig. 4. Sarawakodendron filamentosum DiNG Hou. a. Habit, x 2/3, b. inflorescence, <4, c. flower, 
<2, d. flower with petals removed, 3, e. stamen, x 8, f. longitudinal section of flower shown in d, x3, 
g. longitudinal section of ovary, <8, A. cross section of ovary with position of stamens indicated, x 8, 
i. fruit x 2/3, 7. dehiscent fruit with one valve showing scars of seed attachments, x 2/3, k. one fruit valve 
with attached descendent seeds,  2/s, /. seed, <1, m. basal part of seed seen from beneath, x 2, 7. co- 
tyledons, <2 (a-h DinG Hou 333, i JUGAH ANAK Kupt S 24897, j-n JUGAH ANAK Kupi S 24898). 


932 


6: 397b 


6: 404 


6: 410a 


6: 4105 


6: 4135 


FLORA MALESIANA 


[ser. I, vol. 68 


duncled, few-flowered, the axis of the 
racemes densely set with imbricate, decus- 
sate, persistent bracts. Flowers bisexual, 
solitary, pedicelled. Sepals 5, imbricate. 
Petals ditto. Disk extra-staminal, fleshy, 
flattened. Stamens 3, extrorse, transver- 
sely dehiscing. Ovary partly immersed 
in the disk, 3-celled, with 3 stigmatic 
lobes; no style; ovules c. 8 in each cell. 
Capsule ellipsoid, pointed to both ends, 
hard, 3-gonous, with 3 thick valves, 
dehiscing to the base. Seeds descending, 
narrowly elliptic, with a fleshy orbicular 
aril provided with numerous filiform, 
fringy appendages. 

Distr. Malesia: 
Monotypic. 

Note. Allied to Salacia, Lophopetalum, 
Kokoona, and Polycardia of Madagascar. 


Borneo: Sarawak. 


1. Sarawakodendron filamentosa DING 
Hou, Blumea 15 (1967) 141, f. 1. — 
Fig. 4. 

Tree, 7-12 m, 10-15cm @, the vege- 
tative parts containing yellow kautchuk 
particles and resinous threads. Stipules 
1/> mm, -+ erose. Leaves oblong to 
lanceolate, acuminate, 10-25 by 4-10 
cm, subentire; nerves 6—7 pairs; petiole 
2/3-1 cm. Inflorescences 1-2/2 cm; 
pedicels articulated near base, 11/2—-2 cm. 
Sepals 11/2-2 by 1 mm, semi-orbicular. 
Petals c. 5 mm @, pale-orange. Disk c. 
4 mm @&. Stamens reflexed at anthesis. 
Capsule 6-81/2 by 2-3 cm. Seeds 2-21/ 
by 1/2 cm, albuminous; aril 5-7 mm @, 
the chalazal filaments 11/2-2 cm long; 
embryo narrow-lanceolate, 18 by 4 mm; 
cotyledons foliaceous. 

Distr. Malesia: Borneo (Sarawak). 

Ecol. Understorey tree in lowland 
kérangas forest. 

Note. Its position exactly links the 
families Celastraceae and Hippocrateaceae 
in their former circumscription. 
Siphonodon peltatus DING Hou. 

Add to Distr.: A second collection is 
HARTLEY 13179 with flowers in anthesis. 
Salacia L. 

Line 1 literature: replace (1767) by (1771). 
Add to synonymy: Annulodiscus TARDIEU 
Biot, Bull. Soc. Bot. Fr. 95 (1948) 264; 
FI. Gén. L.-C. Suppl. 1 (1948) 812. 
Salacia sororia Mia. 

Add to Distr.: Now also found in the 
Solomons (Guadalcanal, BSIP 9152). 
Salacia forsteniana Mia. 

Add to Distr.: Now also found in the 
Solomons (Wagnia & SE. New Georgia 
Is.: BSIP 5436, 5985). 

Salacia grandiflora Kurz. 

Add to synonymy following NG, 
Blumea 18 (1970) 412: Maba hierniana 
K. & G. J. As. Soc. Beng. 74, ii (1905) 
203. — Diospyros hierniana (K. & G.) 


6: 415a 


6: 419a 


4: 595a 


4: 533 


4: 569a 


4: 584b 


BAKH. Gard. Bull. S. S. 7 (1933) 173. 
Salacia verrucosa WIGHT. 

Add to synonymy: Annulodiscus nigri- 
cans TARDIEU BLOT, Bull. Soc. Bot. Fr. 
95 (1948) 264; Fl. Gén. I.-C. Suppl. 1 
(1948) 812. 

Salacia chinensis L. 

Line 1 literature: replace (1767) by (1771). 


Chenopodiaceae 


Chenopodium pumilio R. Br. Prod. (1810) 
407; BLAck, FI. S. Austr. 2 (1948) 289; 
AELLEN in Hegi, Ill. Fl. Mitt.-Eur. 3, 
2 (pt 2) (1960) 578, 597, f. 255 D-E; 
STEEN. Blumea 15 (1967) 154. 

Small, soft, prostrate aromatic herb, 
with green flowers. Easily distinct from 
C. carinatum R. Br. by narrow, not 
crested perianth segments, the nut being 
discernible between them. 

Distr. New Zealand and Australia, 
apparently recently introduced in Male- 
sia: New Guinea: Morobe Distr. (B.G.D. 
Leron Cattle Station: E. E. HENtry NGF 
16668, Aug. 1, 1963). 

Ecol. Bare patch in browsed grassland, 
at 150 m. 


Combretaceae 


Due to the interest and activity of the 
Division of Botany, Lae, many additions 
must be made to the Combretaceae of 
New Guinea, two papers on which ap- 
peared by M. J. E. Coope, in the 
‘Manual of the Forest Trees of Papua and 
New Guinea’, Port Moresby 1964, n. 1, 
45 pp., 25 pl., and in a much revised 
edition of this, 1969, 86 pp., 32 pl., 
8 maps. Herein one named and three 
unnamed new New Guinean species are 
distinguished, besides some of the 
Solomons and New Ireland and full 
keys are given. A formal treatment will 
soon appear. : 
It has also been found that within 
Terminalia occur at least four distinct 
types of germination, a character which 
may add to the systematy of the genus. 
Coope’s paper also adds considerably 
to our understanding of the ecology of 
the species and provided numerous field 
notes. The reader is referred to COODE’s 
paper for details. 
Terminalia crassifolia EXELL. 
Add to Distr.: This Papuan endemic now 
also recorded from Australia: Northern 
Territory, Fletcher Creek, Wearyan R.; 
cf. Muelleria 2 (1971) 135. 
Add to Insufficiently known species: 


Terminalia macrantha Rojo, Blumea 17 
(1969) 93, f. 1. 
Tree, c. 18-20 m, 40cm @. Indumentum 


September 1972] 


5: 495 


5: 496 


of simple, brown, sericeous hairs. Young 
branchlets c. 1-2!/2 cm @, glabrescent. 
Leaves densely crowded at the very ends 
of branchlets, chartaceous, rather laxly 
hairy, above shiny, verruculose, glabres- 
cent except on the main nerves, beneath 
persistently hairy, obovate-oblong, 12-31 
by 5-12 cm, generally widest at about 3/4 
of the length, top rounded or sometimes 
emarginate, base narrowly cuneate or 
sometimes decurrent, glandless; midrib 
prominent on both sides, densely hairy; 
nerves on both sides rather widely spaced, 
hairy, the upper 3 or 4 pairs arcuating 
towards the top but not anastomosing at 
the margin, connected by thin veins in a 
more or less scalariform pattern with 
some interconnections between them, 
domatia none; petiole 8-24 mm, densely 
to sparsely hairy, without glands. Spike 
axillary, c. 13-25 cm, lower c. 7-8 cm 
flowerless; densely hairy. Bracts + 
obovate-oblong to + linear, c. 3 mm. 
Flowers sessile, densely hairy, greenish 
yellow, fragrant. Part of the flowers, 
scattered in the inflorescence, with rudi- 
mentary style and somewhat smaller. In 
the bisexual ones, the lower receptacle 
(ovary) 3-6 mm; upper receptacle funnel- 
shaped, 2-3 by 31/2 mm. Calyx lobes 
deltoid, 2 mm, sparsely hairy inside. 
Filaments 9-11 mm, glabrous; anthers 
2/3 mm. Disk sparsely set with rather 
lax hairs. Style simple, terete, 9-10 mm, 
glabrous. Ovules 2, pendulous. 

Distr. Malesia: Philippines (Samar: 
Mt Calbiga, Wright), once collected 
(PNH 6409). 

Ecol. On top of flat ridge, 300 m, lo- 
cally common. 

Vern. Bongoran, S. L. Bis. 

Note. For lack of fruit its affinity can- 
not well be established. There is some re- 
semblance to 7. zollingeri, which has 
coriaceous leaves, more nerves, a smaller 
upper receptacle, and shorter filaments, 
and also with T. darlingii, which has also 
more nerves, subopposite glands at the 
base or top of the lamina, larger bracts, 
a shorter lower receptacle, and longer 
filaments. 


Connaraceae (LEENHOUTS) 


Add to Ecology: Cf. H. G. BAKER, Bot. 
Gaz. 123 (1962) 206-211, on heterostyly 
and pollination; he also confirms the seed 
dispersal by birds. 

Add to Morphology, paragraph on the 
arilloid: Cf. CORNER, Phytomorphology 
3 (1953) 471, for a different opinion as to 
the morphological nature of the aril-like 
structures. 

After Morphology add: 
Phytochemistry. Cf. HEGNAUER, Che- 


Addenda, corrigenda et emendanda 


: 499a 


= 505 


: 506a 


: 507b 


933 


motaxonomie 3 (1964) 545-546, 673. 
Add to Taxonomy, paragraph on affinity 
with other families: On embryological 
grounds, MAURITZON, Act. Un. Lund N.S. 
35, n. 2 (1939) 13 & 39, points to a 
possible relationship with the Cunonia- 
ceae; GUTZ-WILLER, Bot. Jahrb. 81 
(1961) 38-39, gives a not convincing 
argumentation for the inclusion of the 
family in the Sapindales; HUTCHINSON, 
Gen. Pl. 1 (1964) 162, derives the family 
from the Dilleniaceae; HEGNAUER, Che- 
motaxonomie 3 (1964) 546, finds some 
phytochemical support for a position 
near the Leguminosae; LEINFELLNER, 
Oest. Bot. Z. 118 (1970) 542-559, from 
a study of the gynoecium, also con- 
cluded to a closer relationship with the 
Leguminosae. 

Cnestis palala (LoUR.) MERR. 

Add to literature: VipAL, Fl. Camb. 
Laos & Vietn. 2 (1962) 13, t. 1 f. 1-9; 
Corner, Life of Plants (1964) t. 23. 
Add to Distr. of ssp. diffusa: Erroneously 
cited by VIDAL, /. c., from Sumatra, the 
Malay Peninsula, and Borneo. 

Agelaea macrophylla (ZOLL.) LEENH. 
Add to synonymy: Myristica laurina 
(non BL.) Hocure. Candollea 6 (1936) 459. 
Agelaea trinervis (LLANOS) MERR. 

Add to literature: VIDAL, Fl. Camb. 
Laos & Vietn. 2 (1962) 18, t. 1 f. 10-20. 
Add to Uses: In Vietnam, an oil from 
the seeds is used for lighting (VIDAL, /.c.). 
Agelaea borneensis (HooK. f.) MERR. 
Add to Ecol. as exceptional highest 
altitude: (-1300) m. 

Agelaea insignis (SCHELLENB.) LEENH. 
Change description as follows: after 
Branchlets, ‘probably’ should be omitted. 
Leaflets, Sth line, add after tomentose: 
above glabrescent. Include before Fruits: 
Infructescences solitary or fascicled, 
cymose, up to 5 cm long, fulvous- 
velutinous. Add at the end of the de- 
scription of the fruits: via greenish-choco- 
late to red. 

Roureopsis PLANCH. 

Change the date of publication of 
B. & H. Gen. Pl. 1 (sub Taeniochlaena) 
into: 1862. 

Roureopsis asplenifolia SCHELLENB. 

Add to literature: CorNER, Life of Plants 


(1964) t. 23. 
Insert after 1. Roureopsis asplenifolia 
SCHELLENB.: 
la. Roureopsis stenopetala (GRIFF.) 
SCHELLENB. Kew Bull. (1927) 375; 
Cras, Fl. Siam. En. 1 (1928) 362; 


SCHELLENB. Pfl. R. Heft 103 (1938) 110; 
VIDAL, Fl. Camb. Laos & Vietn. 2 (1962) 


23, t. 2 f. 1-6. — Cnestis stenopetala 
GriFF. Notul. 4 (1854) 433, t. 611 f. 2 
(‘steriopetala’). — R. incurva PIERRE, 


Fl. Coch. 5 (1898) t. 379 A. 


934 


FLORA MALESIANA 


[ser. I, vol. 6% 


5: 508a 


S13 S10) 


5: 509a 


5: 5095 


De a10 


Liana. Branchlets fulyvous-tomentose 
when young, glabrescent. Leaves 5—10-ju- 
gate, leaflets mostly opposite or nearly so, 
the petiole and the rachis tomentose. 
Leaflets subsessile, the lateral ones 
(lower- and uppermost excepted) distinct- 
ly asymetric, 11/2-3 by 1/4—-1!/2 cm, basal 
ones broad-ovate, up to 13/4 by 11/4 cm, 
terminal leaflet elliptic or oblong, 2—31/a 
by 1!/4-12/4 cm, all stiff-chartaceous to 
subcoriaceous, midrib puberulous at 
both sides, furthermore glabrous; base 
rounded to (terminal) cuneate; apex 
blunt to rounded, emarginate; nerves 
4-5 pairs, the lowermost at the broader 
acroscopic side ascending, all looped and 
joined, veins about as strongly developed 
as the nerves. IJnflorescences racemose, 
umbelliform, with up to c. 6 flowers, 
c. 11/2-2 cm long, bracts tomentellous, 
furthermore glabrous; rachis c. 4 mm, 
pedicels slender, c. | cm long or more. 
Flowers 5-merous. Sepals linear-lanceo- 
late, 3-5 by 1 mm, blunt, glabrous but 
for the tomentose outer side of the tip. 
Petals linear, c. 8 mm long, puberulous 
at the tip. Stamens confluent at base. 
Fruits 1 (?) per flower, c. 11/2 cm long, 
glabrous. 

Distr. Burma, Thailand, Cambodia, 
Laos, and Malesia: Malay Peninsula 
(Ranong, Phangnga). 

Ecol. Primary and secondary forests 
at low altitude. F/. Febr. 

Note. Nearest to R. asplenifolia 
SCHELLENB. as already mentioned under 
that species (5: 5075). The main differ- 
ences are the usually smaller number of 
slightly bigger and stiffer leaflets which 
are not hairy along the margin and have 
the base mostly rounded, the much longer 
pedicels, and especially the long and 
narrow sepals in the present species. 
Roureopsis emarginata (JACK) MERR. 
Add to literature: VipAL, Fl. Camb. 
Laos & Vietn. 2 (1962) 24, f.2 f. 7 & 8. 
Add to Distr.: Laos. 

Sect. Taeniochlaena LEENH. 

Change the year of publication of B. & H. 
Gen. Pl. 1 into: 1862. 

Roureopsis acutipetala (M1Q.) LEENH. 
Add to literature: VipAL, Fl. Camb. 
Laos & Vietn. 2 (1962) 26, t. 2 f. 9-11. 
Line 4 literature: change 1865 into: 1862. 
ssp. borneensis (SCHELLENB.) LEENH. 
Add to Distr.: According to VIDAL, /.c., 
also in S. Vietnam. 

Rourea AUBL. 

The first citation for Jaundea should be 
changed as follows: Jaundea GILG in 
E. & P. Nat. Pfl. Fam. 3, 3 (1894) 388. 
Add to Sect. Palliatus: Cf. LEENH. in 
Steen. Pac. Pl. Areas 1 (1963) 278, map 
7h 

Rourea oligophlebia MERR. 


5: 5l4a 


5 ibd 


5: 5155 


Silda 


SHAS TB 


525192 


3115204 


Add to literature: VmDAL, FI. Camb. 
Laos & Vietn. 2 (1962) 31, t. 3 f. 1-4. 
Rourea minor (GAERTN.) LEENH. 

Add to literature: VipAL, Fl. Camb. 
Laos & Vietn. 2 (1962) 34, t. 4 f. 1-10; 
W. R. Sykes, Fl. Niue (1970) 70, f. 4. 
Add to literature sub Rourea acropetala 
PIERRE: VIDAL, Fl. Camb. Laos & Vietn. 
2 (1962) 32, t. 3 f. 5-7. 

Add to synonymy, after Santaloides 
cordatum: Santaloides ovale SCHELLENB. 
Bot. Jahrb. 59 (1924) Beibl. . 131, p. 29. 
Add to synonymy, after Santaloides 
luzoniensis SCHELLENB.: Connaropsis ru- 
bescens RIDL. J. Bot. 62 (1924) 295; cf. 
LEENH. Blumea 12 (1963) 20. 

Add to synonymy, after Santaloides 
elmeri SCHELLENB.: Rourea ovale LEENH. 
Fl. Mal. I, 5 (1958) 520; cf. LEENH. 
Blumea 12 (1963) 21. 

Add to Distr.: Solomon Is., Tonga, 
Niue I. 

VIDAL, /.c., distinguished the three groups 
cited as subspecies as follows: ssp. 
microphylla (Hook. & ARN.) VIDAL for 
the small-leaved category, ssp. minor for 
the intermediates, and ssp. monadelpha 
(RoxsB.) VIDAL for the group with few, 
relatively large leaflets. I had refrained 
from giving such a subdivision as these 
groups are neither morphologically, nor 
geographically sharply delimited. Fur- 
thermore, ViDAL excluded R. acropetala 
PIERRE from the synonymy. That species 
should mainly differ from R. minor by 
the reduction of the terminal leaflet and 
by the distinctly mucronate apex of the 
leaflets. 

Add to the 4th paragraph, on more or 
less intermediate forms: R. ovale (Bor- 
neo). 

Rourea mimosoides (VAHL) PLANCH. 
Add to literature: VIDAL, Fl. Camb. 
Laos & Vietn. 2 (1962) 41, t. 3 f. 8-12. 
Add to Distr.: Vietnam. 

Add to Ecol.: peat-swamp. 


f. mimosoides. Add to literature: ANDER- 


son, Gard. Bull. Sing. 20 (1963) 172. 
Omit under Distr.: Sumatra excepted. 
Rourea prainiana TALBOT. 

Sometimes a big climber. 

Add to Distr.: Ceylon (acc. to TALBOT). 
Add after Malay Peninsula: also Pahang 
(G. Benom). 

Under Incertae sedis: Drop R. ovale 
(SCHELLENB.) LEENH. as ripe fruits 
showed that this is identical with R. minor. 
Add under Incertae sedis: 


Rourea pinnata (MERR.) VELDKAMP, 
Blumea 15 (1967) 543. — Sarcotheca 
pinnata Mere. J. Str. Br. R. As. Soc. 
86 (1922) 314; KNutnH, Pfl. R. IV, 130 
(1930) 426. 

Liana. Leaves 


Twigs subglabrous. 


September 1972] 


Addenda, corrigenda et emendanda 


o55 


5: 521a 


52 5216 
23a 


5: 523b 


5: 5246 


39/526 


3-4-jugate; lateral petiolules 4-5 (-7) mm 
long. Leaflets elliptic to oblong, 12-27 
by 4!/2-11 cm, thin chartaceous, shining 
above, rather dull beneath, minutely 
hairy on midrib and nerves beneath; 
base equilateral, in lateral leaflets round- 
ed (to subcordate), in the terminal leaf- 
let broadly cuneate; apex tapering acu- 
minate, acumen short, broad, and blunt; 
nerves 10-12(-15) per side, pinnate, 
ascending, slightly curved, only the 
uppermost ones joined, intermediate 
veins often nearly as strongly developed 
as the nerves, veins and veinlets scalari- 
form, beneath much better visible 
than above. /nflorescences axillary, soli- 
tary or 2 collateral ones together, up to 
7 cm long, sparsely branched, rather 
densely minutely hairy. Calyx 21/2-23/4 
mm high, outside densely appressed 
hairy, margin densely ciliolate, inside 
glabrous. Coro/la (not yet fully developed) 
33/4 mm long. Ovary and lower half of 
the style sparsely woolly. Fruit unknown. 

Distr. Malesia: North Borneo (once 
collected at Batu Lima near Sandakan). 

Ecol. On dry forested slopes at low 
altitudes. Fl. Oct. 

Note. As the fruit is still unknown 
it is not well possible to include this 
species into one of the subgenera or 
sections. The key in Fl. Mal. I, 5 (1958) 
513, leads to R. minor. It is distinctly 
different from that species, however, in 
the following characters: greater number 
of nerves, nervation open and — by the 
intermediate veins — seemingly very 
dense, venation distinctly scalariform 
(in R. minor mostly laxly reticulate), 
inflorescences puberulous, calyx outside 
densely hairy. 

Ellipanthus tomentosus KURZ. 

Add to literature: VmpAL, FI. Camb. 
Laos & Vietn. 2 (1962) 46, t. 4 f. 11-14. 
Add to first sentence of description: up 
to 40 m high, sometimes buttressed. 

ssp. tomentosus. Add to literature: 
ANDERSON, Gard. Bull. Sing. 20 (1963) 
172. 

var. tomentosus. 

Add to Uses: According to VIDAL, /.c., 
the wood is soft and not durable. 
Ellipanthus beccarii PYeRRE. 

Add to literature: Meter, Bot. Bull. 
Herb. Sandakan 10 (1968) pl. between 
p. 104 and 105. 

Add to Ecol.: peat swamp forests. 

Key to the species. 

Lead 10(b), add to length of petals: 
(—5) mm. 

Lead 16(a), insert between ‘dry’ and 
‘blackish-verrucose’: mostly. 

Connarus euphlebius MERR. 

var. bullatus LEENH. Replace ‘in the press” 
by: 106. 


Sisto, 


SIC Sys ys 17) 


5: 533b 


5: 534a 


5: 535a 


329'535D 


5: 536a 


5: 538a 


5: 5386 


5: 539a 
5: 5395 


ssp. moluccanus LEENH. Replace ‘in the 
press’ by: 106. 

Connarus villosus JACK. 

Add to Distr.: N. Borneo (P. Gaya). 
Add to Ecol.: Altitude up to 500 m. 
Connarus culionensis MERR. 

Add to literature: VIpAL, Fl. Camb. 
Laos & Vietn. 2 (1962) 54, t. 5 f. 13 & 14. 
Add to Distr.: Sarawak, Sabah, S. Viet- 
nam (VIDAL). 

var. culionensis. 

Add to Distr.: According to VIDAL, /.c., 
also in S. Vietnam. 

var. stellatus (MERR.) LEENH. 

Delete in description the sentence 
‘Fruit unknown.’ (The fruit is not differ- 
ent from that of the type variety.) 
Connarus odoratus Hook. f. 

Add to Note: The delimitation of 
C. villosus, culionensis, and odoratus has 
to be studied anew when more material 
will be available. Especially in northern 
Borneo, where the three species overlap, 
the identification is often uncertain. 
Connarus paniculatus RoxB. 

Add to literature: VmaAL, FI. Camb. 
Laos & Vietn. 2 (1962) 55, t. 6 f. 1-10; 
ANON. Ic. Roxb. (1970) t. 28. 

Add to Distr.: Burma. 

Add to Notes: VIDAL, /.c., separates the 
species into two varieties: var. panicu- 
latus and var. hainanensis (MERR.) 
VipaL. Among the Indo-Chinese mate- 
rial of the former of these he distinguishes 
3 formae. According to him, the material 
of the Malay Peninsula belongs to var. 
paniculatus. 

Connarus semidecandrus JACK. 

Add to literature: Vmar, FI. Camb. 
Laos & Vietn. 2 (1962) 51, t. 5 f. 1-12; 
ANDERSON, Gard. Bull. Sing. 20 (1963) 
172. 

Add to  Distr.: Sumbawa, Flores; 
according to Hosokawa, J. Jap. Bot. 
13 (1937) 275, also Truk I. 

VIDAL, /.c., distinguishes among the 
Indo-Chinese material, here all included 
in group 8, 3 forms, exclusively on leaf 
characters. 

Connarus cochinchinensis (BAILL.) PIERRE. 
Add to literature: VmaL, FI. Camb. 
Laos & Vietn. 2 (1962) 58, t. 6 f. 11-18. 
Connarus lamii LEENH. 

Replace ‘in the press’ by: 106. 

Connarus monocarpus L. 

Add to literature: CORNER, Life of 
Plants (1964) t. 23. 

Description of the fruit, 3rd line, read: 
in the latter case usually distinctly shortly 
stipitate. 

Connarus lucens SCHELLENB. 

Start description with: Liana, up to 10 m. 
Add to Distr.: Now collected in some 
parts of Sarawak. 

Add to Ecol.: Clayey river bank, peri- 


936 


FLORA MALESIANA 


[ser. I, vol. 68 


5: 540a 


5: 5406 


5: 5406 


5: 54la 


4: 390 


odically flooded, up to 250 m. 

Connarus winkleri SCHELLENB. ssp. philip- 
pinensis LEENH. 

Replace ‘in the press’ by: 106. 

Connarus schumannianus GILG. 

The head should be changed as follows: 


18. Connarus conchocarpus F. v. M. 
Fragm. 5 (1866) 105; SCHELLENB. Pfl. 
R. Heft 103 (1938) 228. 


ssp. schumannianus (GILG) LEENH., nov. 
stat. — C. schumannianus GiLG in K. 
Sch. & Laut. Fl. Schutzgeb. (1900) 341; 
SCHELLENB. Pfl. R. Heft 103 (1938) 253; 
LEENH. FI. Mal. I, 5 (1958) 540, f. 11 j. 
Add to Notes: Ssp. conchocarpus from 
NE. Queensland differs from ssp. schu- 
mannianus in the following points: 
branches more densely lenticellate; leaves 
more often 2-jugate; leaflets thicker, 
mostly at base more acute and _ less 
peltate, midrib often above flat rather 
than sunken, nervation and venation 
more coarse and more prominent be- 
neath; inflorescences apparently usually 
axillary; fruits hardly curved, smaller 
(21/4 by 13/4 cm). 

The relationship is doubtless with 
C. pickeringii A. GRAY (Fiji, Solomon Is.) 
and with C. salomoniensis. 

Connarus salomoniensis SCHELLENB. 

Add to description of Petals, after 
‘21/2 by 11/2 mm’: or linear-lanceolate, 
5 by 1 mm. 

Replace the Ist paragraph of the Notes 
by: 

The present species is doubtless related 
to C. conchocarpus and to C. pickeringii 
A. GRaAy from Fiji and the eastern 
Solomon Is. (San Christobal, Vanikoro 
I.). From the former it is distinctly 
different in its flowers and fruits, but 
hardly so in the leaves. C. pickeringii 
differs constantly by the on both surfaces 
hairy petals and the smaller fruits (up to 
c. 3 by 21/2 cm); moreover, the leaflets 
of the latter species are as a whole 
narrower (up to 71/2 cm) and hardly, if 
all, ovate. I should not be surprised, 
however, if these differences should fade 
away when more material from the 
Solomon Is. comes available. The only 
reason that I not yet reduce C. salomon- 
iensis to C. pickeringii is that the charac- 
ter ‘glabrous petals’ respectively ‘hairy 
petals’ seems as a whole to be important 
in this genus. 

If C. salomoniensis and C. pickeringii 
should be combined, no reason would be 
left to keep C. peekelii upright as a species. 


Convolvulaceae (VAN OosTSTROOM) 


In Key line 18 from top read instead of 
“6. Styles 2, free or united near the base’: 


4: 392 


4: 398 


6. Styles 2 or 1, in the latter case 2-bran- 
ched with the branches distinctly visible. 

Replace forks 7 and 8 bij the following: 

7. Style 1, with 2 branches. 

7a. Flowers in a long racemose inflores- 
cence, each flower inserted on a 
large, elliptic, white bract. Corolla 
to 11/2 cm long. Ovary glabrous; 
stigmas horse-shoe-shaped. 
5a. Neuropeltopsis 
Flowers in an umbelliform cyme, 
not inserted on the bracts. Corolla 
3-5 cm long. Ovary hairy; stigmas 
globose-peltate 4. Bonamia 
7. Styles 2, free. 

8. Each style forked and with 2 filiform 
or slightly clavate stigmas. Corolla- 
limb nearly entire. Small herbaceous 
PlantSiemery | eee 3. Evolvulus 

8. Styles not forked; stigmas capitate 
or kidney-shaped. Corolla-limb dis- 
tinctly lobed. 

8a. Large woody twiners. Bracts much 
enlarged in fruit, elliptic, scarious. 
Stigmas kidney-shaped. 
5. Neuropeltis 
8a. Low perennials. Bracts unaltered 
in fruit. Stigmas capitate. 
3a. Cressa 
Line 1 from top: the correct name for 
1. Subgenus Grammica is: 1. Subgenus 
Cuscuta. 
Insert above 4. Bonamia: 


3a. CRESSA 


LINNE, Sp. PI. ed. 1 (1753) 223; Gen. PI. 
ed. 5 (1754) 104; R. Br. Prod. 1 (1810) 
489; Bru. Fl. Austr. 4 (1869) 437; 
F. M. BatLey, Queensl. FI. 4 (1901) 
1074. — Fig. 5. 

Low, much-branched, grey-pilose, pe- 
rennial plants, often suffrutescent at the 
base. Leaves small, sessile, entire. Flowers 
small, shortly pedicelled or subsessile in 
the upper leaf-axils, in terminal clusters or 
spikes; bracteoles 2, small, at the base of 
the calyx. Sepals 5, subequal, obovate, 
imbricate, about as long as the corolla- 
tube, not enlarged in fruit. Corolla 
regular, funnel-shaped, white or rose; 
lobes 5, spreading or reflexed, about as 
long as the tube, pilose outside. Stamens 
5, exserted; filaments adnate to the 
corolla-tube, filiform, glabrous; pollen 
globular, smooth. Ovary hairy, 2-celled, 
4-ovuled; styles 2, exserted, free, filiform, 
each with a capitate stigma. Capsule 
exceeding the calyx, opening by valves, 
usually 1-seeded; seed smooth, glabrous. 

Distr. A genus with a small number 
of closely related species, considered by 
some authors as a single one (for example 
Cuotsy and BENTHAM), occurring in the 
warm temperate and tropical regions of 
both hemispheres in sandy, maritime 


Ta. 


September 1972] 


Addenda, corrigenda et emendanda 


ox" 


Fig. 5. Cressa cretica L. a. Habit, 
x4, c. calyx, supported by 2 bracteoles, 


or saline circumstances; in Malesia: 
Lesser Sunda Is. (Timor). 
Ecol. Generally in sandy, maritime 


or saline terrain. 


1. Cressa cretica LINNE, Sp. Pl. (1753) 
223; Cuotsy in DC. Prod. 9 (1845) 440; 
Bru. Fl. Austr. 4 (1869) 437; F. M. 
BAILEY, Queensl. Fl. 4 (1901) 1074. — 
C. australis R. Br. Prod. (1810) 490. — 


< 2/3, b. flower, 


x4, 
x6 (a-d CINATTI 339, Port. Timor). 


An erect or diffuse, much-branched 
perennial, sometimes almost woody at 
the base, c. 20 cm, strigose appressed- 
hairy all over. Leaves subsessile to shortly 
petioled, elliptic-oblong or slightly ob- 
long-obovate, acutish, 5-9 by 3-4 mm. 
Bracteoles ovate-oblong. Flowers sessile, 
in terminal heads. Sepals broadly obova- 
te, obtuse, ciliate, 3!/2 mm long. Corolla 
shortly exceeding the calyx, hairy out- 
side, 5!/2-6 mm long. Anthers oblong, 
c. 14/3-11/2 mm long. 

Distr. As the genus; in Malesia: 
Lesser Sunda Is. (Portuguese Timor: 
Batugadé, lagoon of Bé-Malai, CrNnaAtTTI 
339), once collected. 


4: 402 


Ecol. At 100 m from the sea in a pure 
stand in the shade of the mangrove tree 
Avicennia marina. Fl. Aug. 1962. 

Note. The Timor material exactly 
matches specimens from tropical Austra- 
lia, which form BENTHAM referred to 
C. cretica sens. lat., observing that the 
flowers of tropical Australian specimens 
are larger than those in the European 
typical form. In the latter the bracteoles 
are mostly linear, the sepals c. 3 mm long, 
the corolla c. 4 mm, the anthers 1 mm. 
Insert above 6. Porana: 


5a. NEUROPELTOPSIS 


Ooststr. Blumea 12 (1964) 365, f. 1. — 
Fig. 6. 

Differs from Neuropeltis by the longer 
and less dense racemose inflorescences, 
the already at flowering time large, white 
bracts, the broader glabrous wings at 
both sides of the hairy midpetaline areas 
of the corolla, the filaments almost 
wholly adnate to the corolla, and the 
style with 2 short branches, each bearing 
a horse-shoe-shaped stigma. 

Distr. Monotypic, endemic in Malesia. 


1. Neuropeltopsis alba Ooststr. Blumea 
12 (1964) 365, f. 1. — Fig. 6. 

A woody twiner. Young branches 
tomentellous, adult ones glabrescent or 
glabrous. Leaves elliptic, c. 8-12 by 4-7 
cm, rounded at the base or very shortly 
attenuate into the petiole, abruptly 
acuminate and mucronulate at the apex, 
glabrous or nearly so above, rather 
densely pilose beneath; lateral nerves 
5-6 on either side of the midrib; petiole 
c. 2-3 cm long. Jnflorescences from the 
leaf-axils, exceeding the leaves, c. 18-20 
cm long; pedicels 2-8 mm long. Bracts 
white, elliptic to broadly elliptic, at the 
base very shortly attenuate into the 
petiole, abruptly acuminate and mucro- 
nulate at the apex, bearing the 
flower a _ little below their centre, 
c. 4 by 2-21/2 cm, shortly pilose, 
mainly on the midrib and at the margins. 
Sepals c. 3 mm long, two outer ones 
elliptic, obtuse, three inner ones broader 
than long, c. 4 mm broad. Corolla funnel- 
shaped, c. 12-13 mm long, 5-lobed, the 
lobes a little shorter than the glabrous 
tube, broadly ovate, the midpetaline 
areas pilose outside, the glabrous wings 
irregularly dentate. Filaments pilose at 
the base. Ovary glabrous; style 1, with 
2 short branches; stigmas horse-shoe- 
shaped, papillose. 

Distr. Malesia: E. Borneo (Sangkuli- 
rang Distr., Karangan R., NW. of 
Sangkulirang, KosTERMANS 13588). 

Ecol. On sandstone, at c. 20 m alti- 
tude. F/. Aug. 


FLORA MALESIANA [ser. I, vol. 6® 


Za 

y 
Z 
ZA 
Z 


AK 


Fig. 6. Neuropeltopsis alba OoststR. a. Flowering branch, 2/3, b1~®. sepals 1-5, x6, c. corolla with 
stamens, ™ 3, d. pistil, « 6 (a-d KOSTERMANS 13588). 


September 1972] 


4: 431 


4: 435a 


4: 440 


4: 441 


4: 444a 


4: 4455 


4: 446b 


Addenda, corrigenda et emendanda 


939 


Jacquemontia CHOISY. 
Line 5 from bottom, after ‘lanceolate’ 
add: to linear. 
Jacquemontia browniana OOSTSTR. 
Line 2 from top, after ‘with’ add: greyish, 
pale brown or. 
Line 4 from top, after ‘lanceolate’ add: 
to linear. 
Line 4 & 5 from top, alter the sizes as 
follows: 21/2-7 cm by 2-10 mm. 
Merremia DENNSTEDT. 
Change in the Key forks 13 & 14 (lines 
20-22 & 51 from top) as follows: 
13. Sepals partly or all attenuate-acu- 
minate towards the acute apex. 
13a. Leaves entire. Sepals 12-15 mm 
long, the outer ones verruculose 
and sparsely hairy. Ovary hairy. 
6a. M. aniseiifolia 
13a. Leaves generally dentate near the 
base. Longest sepals to 7 mm, 
rarely to 10 mm long, smooth and 
glabrous. Ovary glabrous. 
6. M. tridentata 
13. Sepals obtuse or rounded at apex. 
14. Sepals 10 mm long or mostly 
shorter. 
15. Twining or prostrate herbs .. . ec. 
14. Sepals longer than 10 mm. 

20. Sepals at most 12 mm long... efc. 
Line 1 from top: the correct name for 
1. Section Eu-Merremia is: 1. Section 
Merremia. 

Merremia emarginata (BuRM. f.) HALLIER 


As already stated in my monograph of 
the genus Evolvulus (Thesis Utrecht, 
1934, 245; Med. Bot. Mus. Herb. 
Utrecht 14, 1934, 245) Evolvulus gange- 
ticus)(.) LL: Sp! Pl. ed. 2 (1762) 391 
(= Convolvulus gangeticus L. Amoen. 
Acad. 4, 1756, 306) belongs to the Menis- 
permaceous Cocculus hirsutus (L.) DIELS 
(= Cocculus villosus DC.), such in 
agreement with HALLIER f. (Bull. Herb. 
Boiss. 6, 1898, 720, 723). VERDCOURT 
(in Hubbard & Milne-Redhead, ed., FI. 
Trop. East Afr., Convolv., 1963, 55) who 
could study the type in the Linnean 
Herbarium, also arrived at this conclu- 
sion. The statement by CUFODONTIS 
(Bull. Jard. Bot. Brux. 31, Suppl., 1961, 
743) that the correct name for M. 
emarginata should be M. gangetica (L.) 
CuF. (also based on Convolvulus gange- 
ticus L.) is therefore untenable. 

Merremia tridentata (L.) HALLIER f. 
Line 4 from top: the correct name is: 
ssp. hastata Ooststr. Blumea 3 (1939) 
317, f. 2-0, because the synonym 
Convolyulus hastatus Drsr. in LAMK, 
Enc. 3 (1789) 542, non Forsk. 1775, is 
nom. illeg. This is in accordance with 
Art. 72, note, of the International Code. 
Insert before 7. Merremia quinquefolia: 


4: 447a 


4: 451b 


4: 452b 


4: 457b 


4: 458 
4: 459 


6a. Merremia aniseiifolia Ooststr. Blu- 
mea 12 (1964) 363, f. 1. — Fig. 7. 

A herbaceous twiner (or prostrate 7). 
Stems slender, sparsely pilose, glabres- 
cent. Leaves sparsely pilose, lanceolate 
to linear-lanceolate or sometimes oblong, 
(21/2-) 5-7 cm by 6-10 mm, rounded at 
the base, acute and mucronulate at the 
apex, entire, adpressed-pilose at the 
margins, otherwise sparsely pilose to 
glabrous; lateral nerves 4-6 on either 
side of the midrib; petiole 3-5 mm long. 
Flowers in 1-flowered cymes. Peduncles 
axillary, 2-4 (-6) cm long, slender, 
sparsely pilose to glabrous; pedicels 
thickened towards their top, minutely 
warty, 6-10 mm long. Bracts minute, 
subulate. Flower-buds conical, acute. 
Sepals equal in length or the inner ones 
a little shorter, 12-15 mm long, the two 
outer ones rather thick, ovate-lanceolate 
to narrowly ovate, attenuate or acumi- 
nate towards the acute apex, minutely 
warty and sparsely short-pilose, the 
three inner ones membranaceous, oblong, 
cuspidate, smooth and glabrous or only 
the third sepal warty at the base. Corolla 
funnel-shaped, probably c. 2-21!/2 cm 
long, glabrous, yellow. Stamens included; 
filaments inserted c. 21/2 mm above the 
corolla-base, 6-7 mm long, shortly hairy 
at the base. Ovary pilose; style included, 
c. 8-10 mm long, glabrous. 

Distr. Malesia: West 
(BW 9511 Mort). 

Ecol. In grassland, at c. 600 m altitude. 
The correct reference for species 8 is: 
8. Merremia quinata (R. Br.) OosTsTR. 
J. Arn. Arb. 29 (1948) 417, not Nova 
Guinea n.s. 5: 22. This combination 
antedates that made by Kerr, FI. Siam. 
En. 3, 2 (1954) 106. 

Merremia borneensis MERR. 

Add to Distr.: E. Borneo (KOSTERMANS 
10035). 

Merremia peltata (L.) MERR. 

Line 15 from bottom, after ‘petioles’ add: 
rarely on the whole surface. 

Operculina riedeliana (OLIV.) OOSTSTR. 
A specimen from the Malay Peninsula 
(Negri Sembilan), taken by RIDLEY 
(Fl. Mal. Pen. 2, 1923, 459) for Merremia 
crispatula PRAIN, belongs to O. riedelia- 
na. RIDLEY misquoted both collecting 
site and collector. Mr. H. M. BuRKILL 
(in litt.) was so kind to indicate the right 
collecting site as Bukit Dusun Paya and 
the collector as ALvmns (7. 1181). See my 
paper in Blumea 3 (1939) 368, line 1 from 
top. 

Line 12 from bottom, replace ‘J. ochro- 
leuca’ by: I. ochracea. 

Line 13 from bottom, replace 
congesta’ by: 6. I. acuminata. 
Bottomline, replace ‘3la. J. * sloteri’ by: 


New Guinea 


One 


940 


FLORA MALESIANA 


[ser. I, vol. 66 


Fig. 7. Merremia aniseiifolia Ooststr. a. Flowering stem, 
(a-c MOLL BW 9511). 


4: 460 


4: 461 


4: 462 


3la. I. x multifida. 

Line 9 from top, replace ‘32. J. digitata’ 
by: 32. I. mauritiana. 

Line 21 from bottom, replace ‘18. /. 
maxima by: 18. I. sepiaria. 

Line 7 from top, replace ‘14. J. gracilis’ 
by: 14. I. littoralis. 

Line 13 from bottom, replace ‘35. J. 
riparia’ by: 35. 1. rubens. 

Line 2 from top, replace ‘6. /. congesta’ 
by: 6. I. acuminata. 


x 2/3, b1-5, sepals 1-5, x3, c. pistil, x4 


4: 464a Ipomoea plebeia R. Br. 


4: 464 


Add to Distr.: Also in tropical Africa 
(ssp. africana MEEUSE) and India (ssp. 
indica VERDC.). The specimens from 
Malesia and Australia belong to ssp. 
plebeia. See VERDCOURT in Hubbard & 
Milne-Redhead (ed.), Fl. Trop. East 
Afr., Convolv. (1963) 94. 

Line 14 from bottom: the correct name 
for 2. Section Pharbitis is: 2. Section 
Ipomoea. 


September 1972] Addenda, corrigenda et emendanda 


94] 


4: 465a The correct name for species 4 is: 


ns 


: 465b 


: 470a 


: 472a, 
: 56la 


: 472b, 
: 561b 


: 472b 


: 473b 


: 475b 


: 483a 


: 4845 


4. Ipomoea purpurea Rotu, Bot. Abh. 
(1787) 27, because the synonym Con- 
volvulus purpureus L. Sp. Pl. ed. 2, 1 
(1762) 219 is nom. illeg. This is in ac- 
cordance with Art. 72, note, of the Inter- 
national Code. 

The correct name for species 6. [pomoea 
congesta R. BR. is: 

6. Ipomoea acuminata (VAHL) R. & SCH. 
Syst. 4 (1819) 228, based on Convolvulus 
acuminatus VAHL, Symb. Bot. 3 (1794) 
26. I. acuminata Ruiz & Pav. FI. 
Peruv. 2 (1799) 11 & pl. 120 f. b is not 
valid, because it was published as a 
superfluous name for /. angulata ORTE- 
GA, Hort. Matr. Dec. 7 (1798) 83. 

The correct name for species 14. [pomoea 
gracilis R. BR. is: 

14. Ipomoea littoralis BL. Bijdr. (1826) 
713. I. gracilis R. BR. appears to be a 
distinct species, apparently rare and local, 
and confined to the north coast of 
Australia (STONE, Micronesica 1, 1964, 
126; FosBerG, ibid. 2, 1966, 151-152). 
The correct name for species 16. [pomoea 
ochroleuca SPANOGHE is: 

16. Ipomoea ochracea (LINDL.) G. Don, 
Gen. Syst. 4 (1837) 270, based on Convol- 
vulus ochraceus LINDL. in Edwards, Bot. 
Reg. 13 (1827) t. 1060. 

Add to Distr.: Also in tropical Africa 
and perhaps in tropical America. See 
VERDCOURT in Hubbard & Milne- 
Redhead (ed.), Fl. Trop. East Afr., 
Convolv. (1963) 115-116. 

The correct name for species 18 is: 

18. Ipomoea sepiaria KOEN. ex ROxs. 
Fl. Ind. ed. Carey & Wall. 2 (1824) 90. 
I. maxima (L. f.) DON ex SWEET (Con- 
volvulus maximus L. f.) is a distinct 
species. For the typification of the latter, 
see VERDCOURT, Kew Bull. 15 (1961) 7. 
Add to Distr.: Also in tropical Africa. 
The correct reference for species 20 is: 
20. Ipomoea pes-caprae (L.) R. Br. in 
Tuckey, Narr. Exp. Zaire (March 1818) 
477, non (L.) Sweet, Hort. Suburb. 
Lond. (July 1818) 35. See STEARN, Taxon 
10 (1961) 237-238. 

The correct name for the hybrid 31a is: 
31a. Ipomoea = multifida (RAFIN.) SHIN- 
NERS, Sida 2 (1966) 265. 

The correct name for species 32 is: 

32. Ipomoea mauritiana JAcQ. Collect. 
4 (1791) 216; Pl. Rar. Hort. Schoenbr. 
2 (1797) 39, t. 200. J. digitata L. is a 
distinct species, endemic in the West 
Indies, the name of which was long used 
for J. mauritiana. 

The correct name for species 35. /po- 
moea riparia G. DON is: 

35. Ipomoea rubens CHotsy, Mém. Soc. 
Phys. Genéve 6 (1834) 463. See VERD- 
COURT, Webbia 13 (1958) 324 and do 


4: 487a 


4: 488a 


4: 495 
4: 502a 


4: 510a 


: 2646, 
2 Spiho) 


nas 


S205 


SRO307 


S30307D 


5: 309a 


in Hubbard & Milne-Redhead (ed.), FI. 
Trop. East Afr., Convolv. (1963) 134. 
Ipomoea tuba (SCHLECHTEND.) G. Don. 
Add to Distr.: Thailand (Pur 4326). 
Line 24 from top: the correct name for 
Ipomoea dasysperma JACQ. Eclog. Pl. 1 
(Aug. 1816) 132, t. 89 is: 

Ipomoea tuberculata KeR-GAWL. in Ed- 
wards, Bot. Reg. 1 (Febr. 1816) t. 86. See 
VERDCOURT in Hubbard & Milne-Red- 
head (ed.), Fl. Trop. East Afr., Convolv. 
(1963) 123. 

Line 16 from bottom, replace ‘17. A. 
capitata’ by: 17. A. capitiformis. 

The correct name for species 17. Argyreia 
capitata (VAHL) CuHotsy (1833), based on 
Convolvulus capitatus VAHL (1794), non 
Desv. 1792, nec Cav. 1793, is: 

17. Argyreia capitiformis (PoiR.) Oost- 
STR. nov. comb., based on Convolvulus 
capitiformis Pork. in LAMK, Encycl. 
Suppl. 3 (1814) 469. 

Argyreia congesta OOSTSTR. 

Add to description: Fruit ellipsoid, 
10-12 mm long, red (MUKMIN AMIR 
in herb. Sandakan, n. 35633). 


Corynocarpaceae 


Corynocarpus cribbianus (F. M. BAILEY) 
L. S. SMITH. 

Add to Distr.: This species is also found 
in several of the Solomon Is. 


Dichapetalaceae (LEENHOUTS) 


Dichapetalum THOU. 

Change the year of publication of 
THouars into 1806. Add to literature: 
HuTCHINSON, Gen. Pl. 1 (1964) 216. 
Add to genus diagnosis: The fruits of at 
least some species are apparently 
dehiscent (D. papuanum, D. helferianum, 
D. gelonioides, cf. KANNLAL & Das, FI. 
Assam 1, 1937, 246), exposing the orange- 
coloured to scarlet, thin mesocarp. 
Include after Wood anatomy: 
Phytochemistry. See HEGNAUER, Che- 
motaxonomie 4 (1966) 14-17. 

Insert in the Key after fork 8 2nd lead: 
8A. Inflorescences scorpioid. 

16. D. scorpioideum 
8A. Inflorescences not scorpioid. 

Insert in the Key after fork 15 2nd lead: 
15A. Fruits 3 cm long. 

17. D. grandifolium 
15A. Fruits up to c. 11/2 cm long. 
Dichapetalum timoriense (DC.) BOERL. 
Omit the questionmark before D. 
peekelii KRAUSE. 

Add to Distr.: New Britain, Solomon Is. 
Add to Ecol.: Stems sometimes hollow 
and inhabited by ants. 

Notes, 3rd paragraph: From the addi- 
tional notes in PEEKEL’s MS flora of the 


942 


FLORA MALESIANA 


5: 3096 


5: 310a 


5: 310b 


Se Silas) 
52 3124 


S125 


Bismarck Archipelago it is clear that 
D. peekelii is synonymous with D. 
timoriense. 

Dichapetalum papuanum (BEcc.) BOERL. 
Omit from synonymy: D. grandifolium 
RIDL. 

Omit the headline of ssp. papuanum. 

The entry on D. papuanum ssp. borneense 
LEENH. should be replaced by the follow- 
ing: 


17. Dichapetalum grandifolium RIDL. Kew 
Bull. (1930) 373; LeENH. Blumea 12 
(1963) 21. D. papuanum (BECC.) 
BOERL. ssp. borneense LEENH. Reinw. 4 
(1956) 81; Fl. Mal. I, 5 (1957) 310. 

Dioecious liana. Branchlets glabrous, 
purple-brown, later on greyish. Petioles 
'lo-11/2 cm long, glabrous. Leaves 
elliptic to oblong, 13-25 by 6-12 cm, 
chartaceous, glabrous; glands few, small, 
scattered all over the lower surface of the 
leaf; base acute, slightly decurrent; 
apex more or less abruptly acuminate, 
acumen short and broad, blunt to acute, 
mucronulate; nerves 7-10 pairs, curved, 
most of them distinctly looped and 
joined. Inflorescences (Q unknown) 1-4 
cm long, distinctly stalked, repeatedly 
branched, with several flowers. Flowers 
(3) 5'/2 mm long. Petals obovate, 
halfway incised, glabrous. Disk lobes c. 
1/5-3/4 mm, 2-lobed, glabrous. Pistil- 
lode densely tomentose. Infructescences 
small, short-stalked, with 1-2 fruits. 
Fruits 2—3-lobed, c. 3 cm long, 214/23 
cm wide, smooth, shortly and densely 
fulvous-tomentose, with distinct, narrow 
sutures. 

Distr. Malesia: Borneo (Sarawak and 
North Borneo). 

Ecol. In primary forests at low alt. Fi. 
June, July, Nov., fr. Nov. 

Note. Possibly nearest allied to D. 
papuanum which it distinctly resembles 
especially vegetatively and in the flowers 
(apart from the larger dimensions). 
Well characterized by the exceptionally 
large fruits. 

Dichapetalum gelonioides (ROxB.) ENGL. 
Add to literature: KANJILAL & Das, 
FI. Assam 1 (1937) 245, and sub Chailletia 
sumatrana MiqQ.: TALBOT, For. FI. 
Bombay Pres. 1 (1909) 253, f. 151. 

ssp. tuberculatum LEENH. 

Add to Distr.: Sumatra. 

ssp. pilosum LEENH. 

Add to Distr.: Malay Peninsula (Selan- 
gor). 

Add to Notes: The fruits are sometimes 
reported as being poisonous. 
Dichapetalum tricapsulare (BLCO) MERR. 
After the entry on D. glabrum Exo. add: 
non D. glabrum (VAHL) PRANCE (1968), 
nom. illeg. 


5 


[ser. I, vol. 66 


: 313a 


: 313b 


: 314ab Dichapetalum 


: 3145 


oloa 


e) SSD 


Dichapetalum griffithii (HooK f.) ENGL. 
Change in description the minimum 
number of nerves into 8. 

Dichapetalum setosum LEENH. 

Add to Ecol.: Altitude up to 360 m. 
Dichapetalum steenisii LEENH. 

Add to description: Sometimes a shrub. 
Leaf base acute to blunt. 

Add to Distr.: E. Borneo (ssp. steenisit). 
longipetalum (TURCZ.) 
ENGL. 

Add to literature: CHUN & CHANG, 
Fl. Hainan 2 (1965) 203, f. 407, and sub 
D. hainanense ENGL.: CHUN & How, 
Act. Phytotax. Sin. 7 (1958) 16. 

Change in description: Branchlets some- 
times nearly terete. Leaves ovate- to 
obovate-elliptic to -oblong, above some- 
times glabrous. 

Line 15 from top, replace ‘1 (—2) —lobed’ 
by: 1 (3) -lobed. 

Add to Distr.: S. China, SE. Thailand. 
Dichapetalum helferianum (KURZ) PIERRE. 
Add to Distr. after Malay Peninsula: 
Peninsular Thailand. 

Dichapetalum laurocerasus (Hook. f.) 
ENGL. 

Change in description: Leaves up to 
15 cm long, thin-coriaceous to charta- 
ceous; acumen short to rather long; 
nervation mostly inconspicuous. 
Dichapetalum sessiliflorum LEENH. 
Replace the description by the follow- 
ing: Dioecious liana, shrub, or up to 
c. 5 m high treelet. Branches densely 
fulvous-tomentose to -velutinous when 
young, glabrescent, greyish to purplish 
brown. Leaves elliptic to lanceolate, 
9-28 by 3-11 cm, pergamentaceous to 
chartaceous, above glabrous to hairy on 
midrib and nerves, beneath sparsely to 
densely appressed-pilose at least on 
midrib and nerves; glands few, on the 
lower side, mainly near the base; base 
acute to rounded; margin minutely 
crenulate to entire; apex acuminate, 
acumen short, broad, and blunt to long, 
slender, and acute; nerves 7—16 pairs, 
slightly to strongly curved, at least the 
upper ones distinctly looped and joined. 
Flowers (only 3 buds known) axillary, 
1 or 2, subsessile, or in short-stalked, 
few- to several-flowered glomerules, in 
vivo pale pink. Calyx densely ferruginous- 
tomentose. Petals ovate, slightly emar- 
ginate, outside (margin excepted) and 
inside at the base long-pilose. Disk 
annular, adnate to the stamens. Pistillode 
patently stiff-pilose. Fruits solitary, short- 
stalked, triangular-ovoid, c. 3 by 21/2-3 
cm, smooth to tuberculate, densely and 
shortly ferruginous-tomentose, glabres- 
cent, without sutures, in vivo orange when 
ripe, 3 (-1) -celled; stones free, woody, 
strongly corrugated. 


September 1972] 


Addenda, corrigenda et emendanda 


943 


5: 3l6a 


4: 379a 


Add to Distr.: Vogelkop Peninsula, New 
Britain, Solomon Is. (Three Sisters L., 
BSIP 17224). 

Change in Ecol. altitude: from sea-level 
to 1800 m. Add: Fi. Febr., Oct.-Nov., 


fr. March-April, Oct.-Nov. 


Add: Uses. The leaves are medicinally 
used (‘chewed and the extract spat onto 
wounds to relieve soreness’; FRODIN, 
New Britain); the fruits are edible. 
Dichapetalum tenerum LEENH. 
Add to Ecol.: Fr. July-Aug. 
Add after 15. Dichapetalum 
LEENH. : 


tenerum 


16. Dichapetalum scorpioideum LEENH. 
Blumea 13 (1965) 162. 

Dioecious (?) liana. Branchlets densely 
fulvous-tomentose, glabrescent, purple- 
brown, sparsely lenticellate. Leaves c. 
1 cm long petioled, blade elliptic, 16-20 
by 9-11 cm, chartaceous, when young 
fulvous-hairy mainly on midrib and 
nerves, later subglabrous, lower surface 
with scattered small glands mainly in the 
basal part; base rounded, slightly atten- 
uate; apex rather gradually, shortly, 
broadly, and bluntly acuminate; nerves 
6-8 per side, curved, only the upper 2 or 
3 more or less distinctly looped and join- 
ed near the margin. J/nflorescences 
shortly (2!/2-5 mm) and thickly pedun- 
cled, with 2 spirally recurved, c. 11/2 cm 
long, densely and shortly fulvous-hairy 
cincinni which are densely covered on 
one side with two rows of alternating 
flowers, on the other side with the bracts. 
Flowers seen in bud only, 3 unknown. 
Petals rather deeply bifid, outside 
sparsely appressed-hairy. Disk lobes low, 
broad, slightly bi- to trifid, long woolly- 
ciliate. Ovary densely fulvous woolly, 
2-celled; style 1, cylindrical, short, with 
2 spreading stigmas. Fruits about 
semi-elliptic, flattened, 20 by 13 by 9 mm, 
densely fulvous-velvety; pericarp with 
broad suture; 1-seeded. 

Distr. Solomon Is. (Treasury Group, 
Mono I.). 

Ecol. Secondary forest 
sandstone slope. Fi. fr. April. 

Note. The relationships of this species 
are uncertain. The kind of inflores- 
cence, its most distinctive character, is 
unique among the Asiatic and Pacific spe- 
cies, but is known from some African 
species. 


on rocky 


Droseraceae 


Drosera petiolaris R. Br. 

Of this rare species a second collection was 
made in the Western District, T.N.G., in 
an open grass-sedge plain, growing on 
sand over clay, rosettes over 14 cm @, 


locally common c. 1 mile south of 
Morehead Patrol Post, along track to 
Tonda, Aug. 8, 1967, R. PULLEN 7139. 


4: 379ab Drosera spathulata LABILL. 


6: 423 
6: 426b 


6: 469 


6: 474 


6: 474 


6: 480 


6: 661 


6: 662 
6: 663a 


This species has for the second time been 
found at low altitude in Malesia, viz 
in Sarawak, near Telok Asam, Bako 
National Park, 120 m, by Prof. Purse- 
GLOVE, on mud by sides of streams. Bako 
National Park is situated on generally 
poor, podsolized soils carrying heath 
forest. The size of the flower dissected is 
slightly smaller than described in FI. 
Mal.: sepals 2 mm, petals pink, nearly 
2 mm; but otherwise the specimen shows 
no deviations. 


Epacridaceae 


Line 8 from top, replace ‘R. Br.’ by 
‘(RA BR) I SPRee 

Styphelia malayana (JACK) Spr. 

Add to synonymy: Leucopogon ophirensis 
GriFF. J. As. Soc. Beng. 23, ii (1854) 638. 


Ericaceae (SLEUMER) 


Line 13 from bottom replace ‘ Wirtgenia’ 
by: Andresia. 

In Key to the genera (line 9) replace 
‘2. Wirtgenia’ by: 2. Andresia. 
Rhododendron L. 

It has appeared that there are still new 
species of Rhododendron, amongst others 
from Borneo and New Guinea. I am 
planning to give a supplement in future 
but not of other Ericaceous genera. 


Line 2 from top replace ‘3. Subg. 
Pentanthera sect. Pentanthera by: 3. 
SuBG. ANTHODENDRON sect. Antho- 
dendron. 


Line 16 from top, omit ‘(BL.)’. 

Line 18 from top, after ( Vireya’), insert: 
non RAFIN. 1814. 

Line 18 from top, omit ‘(BL.)’. 

Replace the infrageneric epithet 3. Sub- 
genus Pentanthera by: 3. Subgenus An- 
thodendron (RcuB.) ENDL. ex WILS. & 
REHDER, Monogr. Azaleas (1921) 115. 
Replace the infrageneric epithet 1. Sec- 
tion Pentanthera by: 1. Section Anthoden- 
dron (RcHB.) ENDL. Gen. (1839) 759. 

In Key to the species line 8 read: 
283. R. macrosepalum. 

Replace the name 283. Rhododendron 
linearifolium S. & Z., non Por. in LAMK, 
Encycl. 6 (1804) 267, by: 283. Rhodo- 
dendron macrosepalum Maxim. 


Flacourtiaceae (SLEUMER) 


Scolopia SCHREB. 

I am engaged in a new revision of this 
genus. It has appeared that the new 
record of S. kermodei C.E.C. FISCHER 


944 FLORA MALESIANA [ser. I, vol. 68 
(hitherto known from Burma and Anda- finely reticulate, crenate, 6-11 by 4-8 cm, 
mans) from Malaya (cf. Blumea 17, with 1 or 2 small glands at base near the 
1969, 270), has proved to belong to an apex of the petiole, the latter slender, 
undescribed species. 1-1!/2 cm. Racemes 4—8-flowered, rachis 

5: 14a Paropsia vareciformis (GRIFF.) MAST. (5 mm or less) and pedicels (11/2-2 mm) 
Add to synonymy: Alsodeia chrysodasys puberulent. Perianth segments 4, sub- 
Mia. Fl. Ind. Bat. Suppl. 1 (1861) 390; orbicular-ovate, puberulent outside, ci- 
Ann. Mus. Bot. Lugd.-Bat. 5 (1869) 215. liate, c. 2 mm. Disk shortly 8-lobed. 
Cf. Jacoss, Blumea 15 (1967) 137; 3 Flowers: stamens c. 25, exserted; 
SLeum. Bull. Jard. Bot. Brux. 40 (1970) filaments unequal, 2-2!/2 mm; ovary 
67, f. 5 (distr.). rudimentary, 0.7 mm. 2 Flowers: ovary 

5: 35b, Scaphocalyx spathacea RIDL. ovoid, attenuate at apex, glabrous, 2-3 

565a Add to Distr.: E. Borneo (Berao). mm; stigmas 2, sessile. Fruit not known. 

5: 39a Trichadenia philippinensis MERR. Distr. Malesia: Philippines (Palawan), 
Add to synonymy: The plant distributed twice found. 
under the name Neotrewia arborea ELM. Ecol. On limestone hill. 
nomen in sched., which was referred by Note. X. palawanense is distinguished 
MERRILL to Neotrewia cumingii (M.A.) from X. luzonense by the broadly ovate, 
P. & H. (cf. MerRR. En. Philip. 2, 1923, at base truncate, rounded or slightly 
437), has appeared to belong to 7. cordate leaves, which bear 2 basal pairs 
philippinensis. of lateral nerves. 

5: 51 Line 17 from bottom replace ‘Sect. 5: 80 Under Fig. 35d read 3 instead of 9. 
Pythagorea’ by: SECT. BLACKWELLIA. 5: 95a Casearia grewiaefolia VENT. 

5: 52 Line 14 from bottom replace ‘Subgenus Replace the epithet var. deglabrata 
Pythagorea (LoUR.) SLEUM.’ by: Sub- K. & V. by: 
genus Blackwellia (LAMK) WARB. var. gelonioides (BL.) SLEUM. comb. noy. 
Line 8 from bottom replace ‘Section 5: 985 Casearia pallida CRA‘. 

Pythagorea (Lour.) O.K. *by: Section Add to Distr.: Malay Peninsula (Se- 
Blackwellia BENTH. langor). 

5: 65b Bennettia papuana GiLG is reduced to 5: 105 Under Excluded: Dovyalis macrodendron 
Blumeodendron papuanum P. & H. GILG, which was already tentatively 
(Euphorbiaceae). Cf. Airy SHAW, Kew removed from Flacourtiaceae, has ap- 
Bull. 16 (1963) 349. peared to belong to Suregada ROTTL. (Ge- 

5: 65b, Hemiscolopia trimera (BOERL.) SLOOT. lonium) of the Euphorbiaceae. Cf. 

566a Add to Distr.: Peninsular Thailand STEEN. Nova Guinea, Bot. n. 12 (1963) 
(Nakon Srithamarat). 190. 
5: 685 Add after 3. Xylosma luzonense (PRESL) 
CLOs, etc.: 
Gnetaceae (MARKGRAF, Zurich) 
3a. Xylosma palawanense MENDOZA, Phi- 
lip. J. Sc. 93 (1964) 514, f. 2. 4: 337 Add before Vern.: 

Shrub or tree, up to 7 m, glabrous, Palynology. ERDTMAN (Bot. Notis. 
whether or not armed. Leaves broadly 1954, 80) has found that the subdivisions 
ovate, apex subacuminate, base truncate, of the genus have separate pollen types. 
rounded or slightly cordate, coriaceous, 4: 338 Replace the keys by the following: 


KEY TO MALE PLANTS 


1. Trees or shrubs, only occasionally and partly climbing. Leaves thin, yellowish when dried. 
Inflorescence yellowish; collars flat, almost always conspicuously distant from each other. 


Sect. Gnetum subsect. Eugnemones. 


NN 


neither velvety nor whitish 


Trees. Sterile 2 flowers ovate, long-beaked; beak finely velvety, whitish. 2. G. costatum 
. Trees or shrubs. Sterile 2 flowers sas pe (only in var. griffithii beaked), the tip 


1. G. gnemon 


3. Shrub. Inflorescence simple, slender, its | axis scarcely Ven mm , thick, flower clusters up to 


2mm @ 


var. tenerum 


3. Trees or shrubs. Inflorescence simple or ‘branched, thick, its a axis dn mm \ thick, flower clusters 


up to5mm @. 


4. Trees. Inflorescences almost always branched, all collars distant 


var. gnemon 


4. Shrubs. Inflorescences always simple, often only the lowermost collars distant (inflores- 


cence unknown in var. gracile). 


5. All collars distant. Sterile 2 flowers beaked (oblong, glabrous) 


var. griffithii 


5. Collars at least partly contiguous. Sterile 9 flowers shortly acuminate. 
6. Only the uppermost collars contiguous. Sterile 2 flowers globose with short tip. 


var. brunonianum 


September 1972] Addenda, corrigenda et emendanda 945 


6. All collars contiguous or only the two lowermost distant. Sterile Y flowers tapering. 
var. ovalifolium 
1. Lianas. Leaves brown or black when dry, coriaceous (thin only in 6. G. neglectum and 3. 
G. tenuifolium). Inflorescence not yellowish, its collars always fairly approaching each other 
(the axis never visible between them), their edges bent upward!. Sect. Cylindrostachys 
MARKGR. 
7. Collars of inflorescence dish-like, the flowers coming out freely. 
8. Leaves with numerous spicular cells parallel to the secondary nerves, therefore silky above 
when dry. Stamen with one microsporangium only . . . . . 1. G. gnemonoides 
8. Leaves with few or no spicular cells. Stamen with two microsporangia. 
9. Leaves thin, green when dry, large, elliptic. Inflorescence simple (rarely once branched), 
slender (3 mm broad), spike itself 2cmlong .... . . . . 3. G. tenuifolium 
9. Leaves coriaceous. Inflorescence always branched. 

10. Leaves small, obovate and tailed, distinctly reticulate, black when dry, striate above by 
spicular cells. Inflorescence once branched, slender (3 mm broad), spike itself 1-1+/2 cm 
long) = . . 5. G.arboreum 

10. Leaves often large and broad, not striate, "mostly black when dry and inconspicuously 
reticulate. Inflorescence branched several times, thicker (4 mm broad), spike itself up 


to 4 cm long : . . 4. G. latifolium 
11. Leaves elliptic, brown when dry, distinctly reticulate, secondary nerves distinctly 

joined : . . var. funiculare 
11. Dry leaves nigrescent, “inconspicuously reticulate, secondary nerves ending open. 
12 seaves almost! orbiculanm ents) lesen ee Ge ee. oo Vaseelaxiintescens 
12. Leaves elliptic. 

13. Leaves small, up to9 cm. Spike short,11/2cmlong . . . . . . . var. minus 

13. Leaves large, up to 20 cm. Spike 2-4cmlong . .. . . . ._ var. latifolium 


7. Collars of inflorescence cylindric, keeping the flowers enclosed. 
14. Inflorescence branched, mostly large, with often very long stalks. Exserted part of the 
stamen longer than the perianth. 
15. Dried leaves black, with narrowed base, relatively rich in fibres. Inflorescence 15—20 cm 
long, catkins twice the length of their stalks. Sterile 2 flowers obliquely conical. 
9. G. ridleyi 
15. Dried leaves brown, with rounded base, without conspicuous fibres. Inflorescence 30 cm 
long (in var. abbreviatum 4-6 cm), catkins as long as their stalks. Sterile 2 flowers ovate, 
straight . ire) eh ee oe al Galeptostachyum 
16. Catkins at most 34 mm broad and 4 cm long. 
17. Leaves large, up to 30 by 12 cm. Inflorescence 30 cm long, catkins 3-4 cm long. 
var. leptostachyum 
17. Leaves small, not more than 12 by 6 cm. Inflorescence 4—6 cm long, catkins 1!/2cm Jong. 
var. abbreviatum 
16. Catkins at least 4mm broad and upto6cmlong . . . . var. robustum 
14. Inflorescence unbranched (once branched in 6. G. neglectum), often cauline. Exserted part 
of the stamen shorter than the perianth. 
18. Leaves thin, tapering at both ends. Inflorescence slender (3 mm). 6. G.neglectum 
18. Leaves coriaceous. Inflorescence thick (4-5 mm). 
19. Inflorescences 6 cm long, drooping (unknown in 8. G. k/ossii from Borneo, but its 
rough @ one is of this type). Flowers numerous, imbedded in many hairs. 7.G.cuspidatum 
19. Inflorescence short, mostly erect (2-3 cm). 
20. Inflorescence very thick (7 mm). Leaves large. 
21. Leaves oblong-obovate, somewhat silky above by spicular cells. Flowers immersed 
between few hairs. S02) 22 10 Gi loerzineh 
21. Leaves elliptic, not silky. "Flowers. immersed ‘between numerous long hairs. 
13. G. macrostachyum 
20. Inflorescence moderately thick (4 mm), shortly stalked, mostly erect. Leaves small, 
up to 15 cm long. 
22. Leaves firm (not fleshy), with distinct nervation, not glaucous, not cuneate. In- 
florescence 3 cmlong . . oy 2G: tla 
22. Leaves fleshy, with indistinct nervation, “more or ‘less glaucous. Inflorescence 11/2—2 
cm long. 
23. Leaf base mostly acute, leaves often lanceolate. Collars of inflorescence with angular 
lower edge. Fruit short-acuminate . . . . . . . . . 14. G.microcarpum 


(1) 3 Inflorescence unknown in 10a. G. raya MARKGR., 1la. G.globosum MARKGR., and 7a. 
G. acutum MARKGR. 


946 FLORA MALESIANA [ser. I, vol. 68 


23. Leaf base mostly rounded. Collars of inflorescence with vaulted lower edge. Fruit 
loMe-ACUMIMALE ps GE 8 ee ae Gage, 6 ee 


KEY TO FEMALE PLANTS 


1. Trees and shrubs, only occasionally or partly climbing. Leaves thin, yellowish when dry. 
Inflorescence yellowish, collars flat. Fruit almost velvety. Sect. Gnetum subsect. Eugnemones. 


2. Tree. Flowers ovate, long-beaked; beak finely velvety, whitish . . . . 2. G.costatum 
2. Tree or shrub. Flowers globose, shortly lipped: ae in var. hac tii beaked), the tip not 
velvety nor whitish . . ; . . . 1. G. gnemon 


3. Collars of inflorescence remote. 
4. Flowers globose. Fruit ovate, obtuse. 
. Tree. Inflorescence mostly branched, all its internodes long (*/2-1 cm). Fruit large 
(2 cm long) Poti . . var. gnemon 
5. Shrub. Inflorescence simple, ‘at least its two lowermost ‘internodes long, all others short 
and hidden. Fruit small (1 cm long), inserted on a thickened rachis. var. brunonianum 
4. Flowers oblong, beaked (unkown in var. gracile). Fruit acute. 
6. Fruit oblong. Axis of inflorescence thick (1 mm); internodes 1/2 cm long. var. gracile 
6. Fruit ovate. Axis of inflorescence slender (1/2 mm); internodes 11/2 cm long. var. tenerum 
3. Collars of inflorescence contiguous. Inflorescence short. Flowers acute. 
7. Flowers globose, beaked. Fruit globose .. . a ges. ome. voRentirchin 
7. Flowers ovate, acuminate. Fruit ovate, long- acute i . . var. oyalifolium 
. Lianas. Leaves coriaceous, brown or black when dry (thin only in 6. G. neglectum and 3. 
G. tenuifolium). Inflorescence not yellowish. Collars dish-like. Fruit smooth or warty, not 
velvety. Sect. Cylindrostachys MARKGR. 
8. Inflorescence branched (unknown in 5. G. arboreum, but the 3 one branched, small). 
9. Leaves obovate-cuneate, tailed, small, distinctly nerved below, densely striate by spicular 
cells above. Fruit long- stalked ew . . 5. G.arboreum 
9. Leaves broadest in or below the middle, mostly large, ‘not densely striate above. 
10. Leaves brown when dry, secondary nerves distinctly joining. 
11. Nerves all remote, arcuate. Inflorescence rich and spreading, often 30 cm long, in var. 
abbreviatum much shorter. Flowers globose, shortly tipped embedded in dense hairs. 


Fruit sessile. . . 16. G. leptostachyum 
12. Leaves large, up to 30 by I2¢ cm. ‘Infructescence 20-40 cm long, its internodes 8-12 mm 
long. 


13. Infructescence 20-25 cm long. Fruit ellipsoidal, 2 by 11/2 cm. Internodes 12 mm long. 
var. leptostachyum 
13. Infructescence 30-40 cm long. Fruit parade eis 1.8 2 1.3 cm. Internodes 8 


mmlong . . . . var. robustum 
12. Leaves small, not 1 more than 12 by 6 cm. Infructescence 10 cm 1 long, its internodes 5 
mm long. Fruit 21/2 by 11/2 cm. . . . var, abbreviatum 


11. Secondary nerves at the leaf base approaching each other, all with a straight lower part. 
Inflorescence not so rich, 15 cm abs Flowers obliquely beaked, conical, embedded in 
few hairs. Fruit stalked aebets . . 4. G. latifolium var. funiculare 

10. Leaves black when dry, secondary nerves ’ indistinctly joining. 
14. Leaves with conspicuous spicular cells, though not striate. Flowers embedded in 


numerous hairs. Fruit very large (6 cm), obtusely turbinate, sessile . . 9. G. ridleyi 
14. Leaves without conspicuous spicular cells. Flowers not embedded in numerous hairs. 
Fruit elliptic, up to 21/2 cm long, stalked. . . ‘ . . 4 G. latifolium 


15. Leaves almost or quite orbicular. Fruit oblong- ‘obovate, rather long-stalked. 
var. laxifrutescens 
15. Leaves elliptic. Fruit stalk thick. 
16. Leaves small (not longer than 9 cm). Fruit ovate, small (11/2 cm long) var. minus 
16. Leaves large. Fruit large, 2-21/2 cm, broadly ovate, long- or short-stalked. 
var. latifolium 
8. Inflorescence simple (exceptionally once branched in 6. G. neglectum). 
17. Leaves thin, tapering at both ends. Secondary nerves straight and broken. 
18. Secondary nerves remote but not extremely so. Spike short (4 cm). Collars contiguous. 
Fruit 2 cm long, longitudinally furrowed, acute, with a long, slender stalk. 
3. G. tenuifolium 
18. Secondary nerves extremely remote (up to 3 cm). Spike long, 8 cm. Collars not contigu- 
ous. Fruit small, brownish yellow, sessile, smooth, obtuse, 11/2 cm long. 6. G. neglectum 
17. Leaves coriaceous, secondary nerves bent, not broken. 
19. Leaves silky above by numerous parallel spicular cells. Flowers obtuse. Fruit obtuse, 
large, Warty.). 30. kw wl os we ww mtn vk ce 


September 1972] 


4: 340 
4: 343a 


4: 343a 


4: 344a 


Addenda, corrigenda et emendanda 


947 


20. Tertiary nervation of leaves indistinct. Fruit smooth. 


21. Leaves elliptic, 15 by 7 cm. Fruit ellipsoidal, 5 by 3 cm 


. 10a. G. raya 


21. Leaves lanceolate, 8-9 by 2!/2 cm. Fruit globose, 41/2 ‘cm @. (la. G. globosum 
20. Tertiary nervation of leaves distinct below. Fruit verrucose, S-6 by 2-3 cm. 


11. G. gnemonoides 


19. Leaves not silky by spicular cells. Flowers acuminate. Most inflorescences cauline. 
22. Leaves fleshy, with indistinct nervation, more or less glaucous. 

23. Flowers ovate. Fruit more or less obtuse, yellow, 2 cm long. 14. G. microcarpum 

23. Flowers oblong-conical. Fruit long-acuminate, 21/2 cm long, pink. 15. G. oxycarpum 
22. Leaves firm, not fleshy, not glaucous, with distinct nervation. 

24. Leaves obovate-cuneate, striate above by spicular cells. Flowers obtuse, not embedded 


in thick hair masses. Fruit obtuse, 4 cm long 


10. G. loerzingii 


24. Leaves elliptic, not striate above. Flowers embedded i in thick hair masses. 


25. Leaves broad-elliptic. Fruit large, acute, rough 


8. G. klossii 


25. Leaves twice as long as broad. Fruit not rough. 
26. Collars contiguous, their hair tufts enormous. Fruit small, up to 2 cm long, almost 


globose 


13. G. macrostachyum 


26. Collars remote, hair tufts large, but not enormous. 
27. Leaves large. Inflorescence elongate, fruiting 9-15 cm long. Fruit 2!/2-31/2cm long. 


28. Fruit shining, broad-ovate, obtuse, 21/2 by 11/2 cm 


. 7. G. cuspidatum 


28. Fruit opaque, fleshy, acute, slightly bent upward, 31/2 by 1.8 cm. 7a. G. acutum 


27. Leaves small. Inflorescence short. Fruit small, elliptic, 


Line 13 from top 
Gnemonomorphi’ by: 

Gnetum neglectum BL. 
Add to literature: MARKGR. Blumea 10 
(1960) 431; ibid. 19 (1971) 108. 

Add to description: ¢ Inflorescence often 
cauliflorous, branched once, its branches 
slender, 2-4 cm by 1 mm; catkins 3-5 cm 
by 3 mm. Collars cylindric-infundibuli- 
form, 3 mm high. ¢ Flowers numerous, 
obconical, 2 mm high; stamen bilocular, 
cells white, splitting on their top. Sterile 
2 flowers 4-6, ellipsoidal, short-acute, 
11/2 mm long; involucre chartaceous; 
ovule obliquely ovoid, gradually acumin- 
ate, 1 mm high. 
Add to Distr.: 

Sarawak, S_ coll. 
28459). 

Gnetum cuspidatum BL. — Fig. 8. 

Add to literature: MOLESWORTH ALLEN, 
Mal. Nat. J. 18 (1964) 168-169, 3 photogr. 
(finely illustrated description of 3 in- 
florescence). 


replace ‘Section 
Section Gnetum. 


Borneo (Brunei, S 5752; 
202; Sabah, SAN 


7a. Gnetum acutum MARKGR. 0m. 
nov. — G. acutatum MARKGR. Blumea 13 
(1966) 404, non Mia. 1860. 

Large liana, 25 m by 2!/2cm @; stems 
sulcate and densely lenticellate; twigs 
terete, smooth. Leaves glabrous, elliptic, 
acuminate, 11-17 by 6-7/2 cm, firmly 
chartaceous; nerves 9 pairs, arcuate, 
united before the margin, prominent 
beneath; petiole 12 mm. 2 /nflorescence 
caulifiorous, not branched, in fruiting 
state 9 cm, rachis 5 mm @; peduncle 
8 mm; collars initially infundibuliform, 
later 1 cm wide. 3 Flowers unknown. 
© Flowers immersed in a dense hair-cush- 
ion, cream flushed with pink, acute- 


4: 344a 


4: 3445 


11/2 by 0.8 cm. 
12. G. diminutum 


ovoid, apex upcurved; external involucre 
fleshy, 4 by 2 mm, !/2 mm thick, median 
one conical, thin, 2 by 1 mm, internal 
one chartaceous, ampullaceous, 11/2 by 
0.8 mm, lengthened into a thin apical 
tube 2 mm exserted, fid at apex. Fruit 
acute-ovoid, slightly curved, 31/2 by 
13/4 cm; external involucre fleshy, 1 mm 
thick, median one hard, glossy and fibrous, 
internal one chartaceous, glossy; nucellus 
with embryo 21/2 by 3/4 cm, acute-ovoid. 

Distr. Malesia: West Borneo: Sara- 
wak (Rejang delta, ASHTON S 17804). 

Ecol. Alluvial forest. 

Vern. Layah, Iban. 

Note. Allied to G. cuspidatum by 
the simple, cauliflorous inflorescence with 
thick axis and densely hairy pulvini, 
fruit-size and leaf-shape. Differs by the 
acute fruit with dull, fleshy external 
involucre; also by more acute @ flowers, 
farther exserted micropylar tube, and 
obviously thinner leaves. 

Gnetum ridleyi GAMBLE ex MARKGR. 
Add to literature: Blumea 12 (1963) 78. 
Add to description: 3 Inflorescence 15 cm 
(or longer), widely divaricate-branched; 
fertile parts 3-5 cm, twice as long as 
peduncle; collars numerous, 2 by 4 mm, 
cylindric. Flowers interspersed with 
numerous hairs; ¢ flowers obconical, 
much narrowed to base, 11/2 by 3/4 mm; 
sporophyll thin, 2 mm exserted; sporan- 
gia 2, ovoid. Sterile 2° flowers 6, obliquely 
conical, 11/2 by 1 mm, external and inter- 
nal involucre strongly fibrous (E. 
Malay Peninsula, P. Tioman, KADIM & 
Noor 644). 


10a. Gnetum raya Markar. Blumea 14 
(1966) 284. 


{ser. I, vol. 68 


Fig. 8. Gnetum cuspidatum BL. Abundant setting of seed, in Sarawak, near Bintulu; * '/10 (DING Hou 321). 


September 1972] 


Addenda, corrigenda et emendanda 


949 


4: 3445 


Large, glabrous liana, 20 m; twigs 
terete, smooth, lenticellate below the 
thicker nodes. Leaves elliptic, coriaceous, 
at base narrowed into petiole 1 cm, at 
apex short-acuminate, up to 15 by 7 cm, 
with parallel fibres visible on upper sur- 


face, grey-shining; nerves 6-8 pairs, 
arcuate-joining 1 cm from margin, 
prominent beneath, veins indistinctly 


reticulate. ¢ Unknown. Fruiting 2 in- 
florescences often several at one node, 
3-4 cm by 4-5 mm, collars dish-like, 
6-8 mm ©. Fruit immersed into cushions 
of short hairs, ellipsoidal, obtuse, 5 by 
3 cm, attenuated at base into a hollow 
pseudostipe 5-10 by 8 mm, opaque, finely 
punctulate, grey-green, inside with a sour 
smell. Outer involucre fleshy, 2 mm thick, 
fibrous at the inner surface, middle one 
coriaceous, longitudinally sulcate, inner 
one chartaceous. Seed (unripe) oblong, 
30 by 8 mm. 

Distr. Malesia: Central Borneo (Up- 
per Kapuas, Bt Raya, Sibu, S 23801); 
Sarawak (Kuching, near Matang, 500 m, 
mixed Dipterocarp forest, S 25646). 

Ecol. On sandy clay, 200 m. 

Vern. Akar téngang, Iban. 

Notes. Allied to the Sumatran 
G. loerzingii MARKGR., with which it 
shares a large-fruited group with fibrous 
leaves and fruits (G. klossii MErRR., 
G. ridleyi GAMBLE, G. gnemonoides 
BRONGN., G. cuspidatum BL.). The 
peculiar smell of the seed is also found 
in G. loerzingii. This species differs by 
obovate-cuneate leaves and by a rough 
fruit surface. 


lla. Gnetum globosum MarkGr. Blu- 
mea 19 (1971) 108. 

Liana with terete, smooth twigs. 
Leaves coriaceous, light brown when dry, 
lanceolate, 8-9 by 2-2!/2 cm; nerves 
straight, at an angle of 30° with the 
midrib, arcuate-joining !/2 cm before the 
margin, hidden among numerous parallel 
fibres on either surface; petiole !/2 mm. 
3 Inflorescence unknown. Fruiting ° 
inflorescence axillary, simple, peduncle 
7 mm, rachis 13-23 by 3 mm, with 6-8 
mm long joints; collars dish-like, 1 by 
4 mm. Fruit globose, smooth, grey-green, 
41/2 cm, contrasted at base; outer invo- 
lucre rather fleshy, 2 mm thick, fibrous 
inside; middle one 1/2 cm thick, outside 
fibrous, inside smooth; inner one 
chartaceous, outside sparsely fibrous, 
inside smooth. Seed globular, 21/2cm ©. 


Distr. Malesia: Malay Peninsula 
(Pahang: Ulu Sat, FRI 15262), one 
collection. 

Ecol. Riverside forest, 120 m. Fr. 


July. 
Note. Approaching G. ridleyi GAMBLE 


4: 346b 


52335 


from Pahang by its rather large fruit and 
fibrous leaves; G. ridleyi differs by much 
larger and less fibrous leaves, by ramified 
inflorescences and larger, spindle-shaped 
fruits. G. gnemonoides BRONGN., of 
the same group, has lenticellate fruits 
and leaves that beneath are not fibrous 
but reticulate. 

Gnetum leptostachyum BL. 

Add the following variety: 


var. abbreviatum MARKGR. Reinwardtia 
1 (1952) 462. 

Leaves up to 12 by 6 cm, coriaceous, 
beneath distinctly reticulate-veined. g 
Inflorescence 4-6 cm, branched. ¢ Cat- 
kins 11/2 cm by 3 mm. Infructescence up 
to 10 cm (internodes 5 mm). Fruit large, 
21/2 by 11/2 cm. 

Distr. Malesia: N. Borneo, Mt 
Kinabalu, 1200-1800 m (CLEMENs 32990, 
type; 32276, 32475, 32488, 32601, 
32698, 32991). 

Note. Some inadequate specimens I 
have identified formerly as G. diminutum 
MarkGr.; the branched inflorescences, 
however, prove them to belong to G. 
leptostachyum, \t is a mountain variety 
with reduced size of leaves and inflores- 
cences, combining the narrow, short 3 
catkins of the lowland variety /eptosta- 
chyum with the short-jointed, large- 
fruited 2 catkins of the lowland variety 
robustum. 


Goodeniaceae (LEENHOUTS) 


Add to family diagnosis: Leaves some- 
times verticillate. 

Under Distribution the number of 
Malesian species of Scaevola, apart from 
the littoral S. taccada, should be changed 
into 5. 

Add to Pollination: See also CAROLIN, 
Proc. Linn. Soc. N.S.W. 85 (1960) 
197-207. G. W. GILLETT, in a letter 
dated 1-9-1964, on Hawaiian Scaevola 
informed me: ‘Incidentally the S. 
taccada populations around here are 
strictly inbreeders, the indusium being 
filled with pollen before the corolla opens, 
after which the indusium closes over the 
pollen-filled stigma so that there is no 
chance that foreign pollen could ever 
make contact with the stigmatic surface. 
I have checked this in a very large num- 
ber of flowers in widely separated 
populations. Our Hawaiian endemics do 
not behave this way.’ 

Add to Anatomy: CARLQuisT, Ann. Mo. 
Bot. Gard. 56 (1970) 358-390 (moreover 
a very interesting general paper). 

Add after Anatomy: 

Phytochemistry. See HEGNAUER, Che- 
motaxonomie 4 (1966) 212-215. 


950 


357506 


5: 336a 


5: 336 


5: 337b 


Add to the introduction to the family: 


FLORA MALESIANA 


[ser. I, vol. 66 


Morphology. CAROLIN, Proc. Linn. Soc. 
N.S.W. 84 (1959) 242-255, on the anat- 
omy and especially vasculation of the 
flowers. CAROLIN, ibid. 91 (1966) 58-83, 
on the morphology of fruit and seed as 
well as on the systematics of the family 
in general. 

Cytology. See PEACOCK, Proc. Linn. Soc. 
N.S.W. 88 (1963) 8-27. 

Velleia J. E. SMITH. 

Add to literature: CAROLIN, Proc. Linn. 
Soc. N.S.W. 92 (1967) 51. 

Velleia spathulata R. BROWN. 

Add to literature: CAROLIN, Proc. Linn. 
Soc. N.S.W. 92 (1967) 51. 

Description, line 3, replace ‘herbaceous’ 
by: fleshy. 

Goodenia J. E. SMITH. 

Distr., change to: three species known 
from outside Australia/Tasmania. 

Add the following key: 


KEY TO THE SPECIES 


1. Plant 20-60 cm high. Leaves linear- 
lanceolate, up to 22 cm long. Inflores- 
cences many-flowered. 

3. G. purpurascens 

1. Plant up to 10 cm high. Leaves ovate 
or obovate, up to 6!/2 cm long. 
Flowers solitary. 

2. Plant glabrous. Leaves coarsely 
dentate 1. G. koningsbergeri 
2. Plant densely hairy. Leaves entire. 
2. G. pumilio 
After Goodenia koningsbergeri add: 


2. Goodenia pumilio R. Brown, Prod. 
(1810) 579; BANKs & SOLAND. Bot. Cook’s 
Voy. 2 (1901) t. 177 f. A; Krause, Pfi. 
R. Heft 54 (1912) 93; RipsDALE, Trans. 
Papua N.G. Sci. Soc. 9 (1968) 18. — 
Fig. 9. 

Erect or prostrate, fairly densely stel- 
late-hairy, annual or perennial herb. 
Leaves radical-rosulate; (broad-)ovate 
to spathulate, up to 6 by 3 cm but usually 
much smaller, herbaceous, attenuate at 
base; margin entire; apex blunt. Flowers 
solitary, axillary, 4-5 mm long, hairy; 
pedicel c. 1 cm long; bracteoles 0. 
Corolla c. 3 mm long, dark reddish. 
Capsules (ovoid-)oblong, 3-4 mm long. 
Seeds many, orbicular, small. 

Distr. Australia (Northern Terr., 
Queensland) and Malesia: New Guinea 
(Papua, Western Distr., near Weam, 
RIDSDALE & GALORE NGF 33733; 
Bula village, PULLEN 7009). 

Ecol. Savannah grassland, c. 10-35 
m. Fi. fr. Aug. 

Note. We owe the identification of 
this and the following species to Prof. 
R. C. CAROLIN, Sydney. 


Fig. 9. Goodenia pumilio R.Br. a. Habit, 2, b-c. 


x10, d. corolla, from above, x10, e. 


style and stigma, 20, f. calyx with fruit, «10, g. 


seeds, 8 (a—g PULLEN 7009). 


3. Goodenia purpurascens R. BROWN, 
Prod. (1810) 578; Krause, Pfl. R. Heft 
54 (1912) 91; F. M. BAILEY, Compr. Cat. 
Queensl. PI. (1913) f. 254. 

Erect, sparsely hairy to subglabrous, 
annual or perennial herb. Leaves mainly 
radical rosulate; linear-lanceolate, up to 
22 by 1/4-13/4 cm (the few cauline ones 
much shorter), somewhat fleshy, sessile 
with a slightly narrowed base; margin 
subentire, with a few minute teeth; 
apex acute. /nflorescences terminal, up to 
25 by c. 5 cm, thyrsoid, repeatedly laxly 
branched and many-flowered. Flowers 
10-15 mm long, thin-hairy to nearly 
glabrous; pedicel c. °/4 cm _ long; 
bracteoles 0. Corolla c. 3/4-11/4 cm 
long, pink to purple. Capsules ovoid, 
3-4 mm long. Seeds many, lenticular, 
minute. 

Distr. Australia (Northern Terr., 


September 1972] 


an 


: 3385 


: 3385 


: 3395 
= 839 


Addenda, corrigenda et emendanda 


eI 


Queensland) and Malesia: New Guinea 
(Papua, Western Distr., Wassi Kussa R., 
Henty & KATIK NGF 38744). 

Ecol. Savannah, in shallow 

alt. a few metres. F/. July. 
Calogyne pilosa R. BROWN. 
Add to Distr.: Now known from several 
localities in both eastern and western 
parts of New Guinea. 
Add to Ecology: Also collected in marshy 
places. Change highest altitude to 250 m. 
Lechenaultia filiformis R. BROWN. 
Add to Ist sentence of description: 
sometimes woody at base, hence appar- 
ently perennial, up to c. 50 cm high 
(NGF 38778). 
Line 1, replace ‘650’ by : 850. 
Scaevola L. Add: nom. cons. 
Description line 1, after ‘opposite’ add: 
or whorled. 
Distr., last line: the number of Malesian 
species “1° should be changed into 3. 
Replace the Key to the species by the 
following one: 
1. Leaves opposite, not tufted. Scrambl- 
ing or climbing shrubs. Flowers yellow. 
3. S. oppositifolia 
1. Leaves either spirally arranged, or 
verticillate, usually tufted at the ends 
of the branches. Erect shrubs. Flowers 
not yellow. 
2. Leaves in whorls of four. 
S. verticillata 
2. Leaves spirally arranged. 

3. Calyx-lobes usually 2!/2 mm or 
more. Flowers 2-21/2 cm _ long. 
Plant from the beach 1. S. taccada 

3. Calyx-lobes 1 mm long or calyx not 
distinctly lobed. Flowers 1—-11/2 cm 
long. Mountain plants. 

4. Leaves petioled, 31/2—-20 by 11/2-8 
cm, flat. Flowers arranged in 
inflorescences, 1 cm long; calyx 
distinctly lobed . 2. S. micrantha 

4. Leaves subsessile, 21!/2-41/2 by 
1/5—-3/4 cm, with revolute margins. 
Flowers solitary, 11/2 cm long; ca- 
lyx not distinctly lobed. New Guinea 
(Fl. Mal. 5: 567) S. pauciflora 


water ; 


5: 339a Replace the name Scaevola sericea VAHL 


by: 


1. Scaevola taccada (GAERTN.) ROXB. 
Hort. Beng. (1814) 15, based upon 
Lobelia taccada GAERTN. Fruct. 1 (1788) 
119, t. 25 f. 5. The vernacular name 
taccada, used as specific epithet, must be 
accepted as an indirect reference from 
ROXBURGH to GAERTNER’S name. 

This is strengthened by the direct 
reference of both to Béla-Modagam 
RHEEDE, Hort. Malab. 4 (1673) 119, t. 59, 
as well as by the reference in Roxs. FI. 
Ind. 2 (1824) 146 to GAERTNER. See H. 
St. JOHN, Taxon 9 (1960) 200-208; 


an 


: 340a 


: 341la 
: 341b 


: 342a 


: 342b 


FosBerG, Taxon 10 (1961) 225-226. 
The synonym Scaevola lobelia Murr. 
Syst. Veg. ed. 13 (1774) 178 is illegitimate 
by the citation of Lobelia plumierii 
L. Sp. Pl. (1753) 929 as a synonym. 
Moreover, Scaevola plumierii as well as 
S. taccada are included, hence it has 
partly to be excluded from the Malesian 
flora. 
Line 3 from top replace ‘12-26 by 5-10’ 
by: 39 by 13. 
Add to Distr.: E. Africa (Kenya). 
Add to Ist paragraph: For dispersal see 
also G. L. Lesko & R. B. WALKER, 
Ecology 50 (1969) 730. According to these 
authors the fruits, which remain viable 
for a long time, float in sea-water but 
germinate in fresh water only, that 
means after having been drifted ashore 
and after rain. 
Scaevola micrantha PRESL. 
Add to Ist sentence of description: or 
treelet to 10 m by 5 cm. 
Add to Distr. after Borneo: Mt Meliau, 
Mt Tavau. Add at the end of Distr.: 
Cited from Botel Tobago (near Taiwan) 
by HatusimA, Mem. Fac. Agr. Kagosh. 
Un. 7 (1970) 327. 
After Scaevola micrantha PResL add: 


Scaevola verticillata LEENH. Blumea 12 
(1964) 317, f. 1. — Fig. 10. 

Shrub. Branchlets + terete, woolly 
tomentose; leaf-axils provided with a 
tuft of c. 11/2 cm long, isabelline, 
sericeous hairs, in older leaves forming 
a kind of papyraceous ligule. Leaves 
whorled in fours, + tufted at the end of 
the branches; petiole c. 3/4-1 cm long, 
grooved above, densely woolly; blade 
obovate-oblong, 5!/2-7 by 21!/2-3 cm, 
chartaceous, apparently convex with 
recurved margins, fairly densely shortly 
tomentose above, glabrescent, densely 
woolly tomentose beneath; base cuneate, 
slightly attenuate; margin minutely 
serrate towards the apex; apex acute; 
midrib slightly sunken above, prominent 
beneath; nerves c. 10-12 pairs, promi- 
nulous above, invisible beneath. /nflores- 
cences condensed, few-flowered, peduncle 
short; bracts relatively large. Flowers 14 
mm long. Calyx lobes different, dorsal 
one narrowly deltoid, acute, 2!/2 mm, 
the others ovate, rounded, ventral one 
1 mm, laterals 3/4 mm, all outside (as 
well as the ovary) fairly densely, inside 
sparsely woolly. Corolla tube inside 
densely woolly, more sparsely so in the 
basal 1/4 of the lobes, outdise densely 
woolly-tomentose but for the basal 
half of the tube; membranous margins 
of the lobes broad and crispy in the 
upper half, in the lower half narrowed 
and with some long and stiff marginal 


952 


FLORA 


MALESIANA 


[ser. I, vol. 68 


Fig. 10. Scaevola verticillata LEENH. a. Flower, 4, b. calyx, <4 (note the three different kinds of lobes!), 
c. & d. anther from in- and outside resp., < 8, e. indusium, = 8 (showing the spreading hairs at its base) 
(a-e W. MEWER SAN 28818). 


5: 3426 


5: 344a 


5: 369 


35) 370 


hairs and 2 or 3 lengthened teeth crowned 
with a brush-like bundle of hairs. Style 
with a few woolly hairs in the basal half 
and a collar of long, stiff, spreading hairs 
just under the indusium. Fruits unknown. 

Distr. Malesia: Borneo (Mt Tambuyo- 
kon near Mt Kinabalu). 

Ecol. Subalpine vegetation, on ser- 
pentine, 2500 m. Fi. July. 

Note. Nearest allied to S. micrantha 
and S. pauciflora. Especially charac- 
terized by the whorled leaves and the 
heterosepalous calyx. 

Scaevola oppositifolia R. BROWN. 

Line 21 from bottom, add after ‘acute’: 
or sometimes rounded, truncate, or even 
subcordate. 

Line 7 from bottom, add after ‘Style 
glabrous’: to sparsely pilose. 

Add to Distr.: Ferguson I., Normanby I., 
and Goodenough I. 


Hamamelidaceae 


Distylium Step. & Zucc. 

The genus Distylium was also recorded 
from Central America with 2-3 spp. 
Recently ENpREss (Bot. Jahrb. 89, 1969, 
355) has referred these American species 
to a separate genus Molinadendron 
ENpRESS, which he even says stands very 
remote and does not belong in the same 
tribe. The arguments for this seem to me, 
however, to be of more trivial nature and 
I do not agree with their inflation to 
generic rank. 

Sycopsis OLIVER. 

Recently ENpREss (Bot. Jahrb. 90, 1970, 
30) has from the genus Sycopsis split off 


323710 


55/381 


5: 393a 


5: 402a 


4 Sino-Malesian spp. to represent a 
separate genus Distyliopsis ENDRESS, 
which the author maintains close to 
Sycopsis. To this also belongs the single 
Malesian species S. dunnii HEMSL. The 
arguments on which this distinction is 
based are partly due to a theoretical 
explanation of inflorescences, partly 
certainly not of generic rank (degree of 
connation of sepals) and partly of no 
value whatsoever (almost replacing 
ranges). I am not prepared to accept this 
as a distinct genus. 

Sycopsis dunnii HEMSL. 

Add to Distr.: Now also found in North 
Borneo (Mt Kinabalu, Mesilau R., 
RSNB 7022) (add to fig. 5). 


Hydrocaryaceae, see Trapaceae 
Hydrocharitaceae (DEN HARTOG) 


A monograph has appeared by DEN 
HartoG, The Seagrasses of the World, 
Verh. Kon. Ned. Akad. Wet. A’dam, afd. 
Natuurk. ser. 2, 591 (1970) 1-275, 63 fig., 
31 pl., and furthermore world maps of 
distribution of seagrasses in Pacific 
Plant Areas 2 (Blumea, Suppl. 5) (1966) 
208-219. Both contain a wealth of new 
data of various kind. 

Blyxa novoguineensis HARTOG. 

Add to Distr.: It has probably also been 
found in the Philippines (Luzon) by 
Jacoss, but unfortunately the specimen 
is sterile. 

The authority of Enhalus acoroides is: 
(L. f.) Royre Ill. (1840) 453. This 
publication antedates STEUDEL’s Nom. 


September 1972] 


5: 408 
5: 410a 


5: 4lla 


5: 412a 


6: 153a 


4: 211b 


4°) 215b 


Bot. ed. 2, 1 (1840) 554 by a few months. 
In Key, line 7, replace ‘2. H. minor’ by: 
2. H. ovata. 

The correct name of 2. Halophila minor 
(ZoLL.) HartoG is: 2. Halophila ovata 
GAupD. in Freyc. Voy. Bot. (1827) t. 40, 
f. 1. In the latter published text of the 
same work, /.c. 430, GAUDICHAUD added 
as a synonym Caulinia ovalis R.BR., 
which would make his own name illegit- 
imate; but his earlier published plate is 
validly published. Cf. HARTOG (1970) 251. 
Halophila decipiens var. pubescens HAR- 
TOG is no longer recognized. Cf. HARTOG 
(1970) 254. 

Halophila beccarii ASCHERS. 

Add to Distr.: Found abundantly in the 
Malay Peninsula. Cf. HARTOG (1970) 262. 


Juglandaceae 


Engelhardia spicata LECHEN. ex BL. 
Replace the name var. colebrookeana 
(LINDL. ex WALL.) O.K. by: 

var. integra (KURZ) MANNING, comb. nov. 
— E. villosa Kurz var. integra KURZ, 
For. Fl. Burma 2 (1877) 492. 


Juncaceae 


Juncus effusus L. 

Add to Distr.: Malay Peninsula (Pahang, 
Cameron Highlands, G. Batu Brinchang, 
2 clumps along roadside, c. 1 mile from 
summit, c. 2000 m, J. SINCLAIR 9956 
(SING), Nov. 4, 1958). 

Widely distributed in Malesia. In the 
Malayan specimen the inflorescence is up 
to 18 cm, due to elongated branchings. 
The fact that this species was found so late 
in Malaya and along a roadside in the 
newly opened Cameron Highlands is no 
proof at all that it is not native. In these 
forested mountains Juncus could original- 
ly equally well have been very scarce 
because of its heliophilous ecology, but 
extended after man artificially opened 
the forest. This happened also in Came- 
ron Highlands with Juncus prismatocar- 
pus R.Br., first collected in 1958. Cf 
KeRN, Gard. Bull. Sing. 17 (1958) 
91-92. The like happens frequently; for 
example on Mt Diéng, in Java, Gentiana 
quadrifaria is a common weed on paths, 
though doubtless native and_ scarce 
before this mountain was deforested. 
Add to Excluded: 

Luzula silvatica (HuDSs.) GAUDIN, Agrost. 
Helvet. 2 (1811) 240; Bucn. Pfl. R. Heft 
25 (1906) 55. — Juncodes silvaticum 
(Hups.) O.K. Rev. Gen. PI. 2 (1891) 725. 

This was recorded by O. KUNTZE from 
Central Java: Diéng Mt (O. KUNTZE 
5715b, Bucn. /.c.). Though I have not 
seen the specimen there is presumably not 


Addenda, corrigenda et emendanda 


6: 296 


953 


the slightest doubt about BUCHENAU’s 
identification. Dr. BAssetTr MAGUIRE 
wrote me (26-2-60) that is it not in the 
New York Bot. Gard. Herbarium with 
KUNTZE’s collection. The species is not 
known to occur outside Europe. It is 
also strange that it is a b-number, which 
might suggest that it was mixed with 
something else. Could it have happened 
that KUNTZE brought along old drying 
paper which he had used in Europe and 
in this way transported a European 
herbarium specimen to Java? This has 
occurred before, and by miracle also 
with a plant from Mt Diéng, viz Luronium 
natans (L.) RAFIN., cf. Fl. Mal. I, 5 
(1957) 3346. This mislocation could be 
proved by means of using diatoms adher- 
ing to the plant as tracers (cf. STEEN. 
Taxon 5, 1956, 157-158) by VAN DER 
WeRrFFE (Blumea 7, 1954, 599-601). If we 
had KUNTZE’s specimen we _ could 
probably apply this method with success. 
Another solution to the mystification 
could be that a European collection got 
mixed with Javanese collections after 
KuNTZE had returned to Europe; a 
similar thing happened with RIDLEyY’s 
Javanese collections which contain a few 
European Carices. 

It is said that L. silvatica is sometimes 
dispersed outside its natural habitat with 
grass-seed in Holland, but this cannot 
have happened in KUNTZE’s time to Java. 
It certainly does and did not occur in Java. 


Loganiaceae (LEENHOUTS) 


Add to Phytochemistry: Cf. HEGNAUER, 
Chemotaxonomie 3 (1964) 307-310, 647. 
Insert before Delimitation and subdivi- 
sion: 

Cytology. Cf. GaADELLA, Act. 
Neerl. 11 (1962) 51-S5. 
Palynology. Cf. PUNT & LEENHOUTS, 
Grana Palynologica 7 (1967) 469-516. 
Add to the discussion on the position of 
the Buddlejeae: W. TROLL, Jahrb. 1965 
Akad. Wissensch. Lit. Mainz (1966) 128, 
on arguments derived from the morphol- 
ogy of the inflorescence, also concludes 
to a closer relationship with the Loga- 
niaceae. 

Add to (4) Peltanthera: Revision: 
LEEUWENBERG, Act. Bot. Neerl. 16 (1967) 
143-146. 

Add to (8) Gomphostigma: Revision: 
VERDOORN, FI. S. Afr. 26 (1963) 168-171. 
Add to (10) Emorya: See: NORMAN & 


Bot. 


Moore, Southwestern Naturalist 13 
(1968) 137-142. 
Add to (11) Adenoplusia and (12) 


Adenoplea: Both reduced to Buddleja by 
LEEUWENBERG, Act. Bot. Neer]. 16 (1967) 
143. 


954 


FLORA MALESIANA 


(ser. I, vol. 6® 


6:5 297 


6: 300 


6: 302 


6: 303a 


6: 304b 


6: 308a 


6: 3085 


6: 3lla 


6: 315a 


6: 3166 


6: 317b 


6: 318a 
6: 320a 


Add to (13) Bonyunia: 4 or 5 spp. 6: 320b Fagraea ridleyi K. & G. 


Revision: LEEUWENBERG, Act. Bot. Neer]. 
18 (1969) 152-158. 

Add to (16) Usteria: Revision: LEEUWEN- 
BERG, Act. Bot. Neerl. 12 (1963) 112-118. 
Insert before E. Strychneae: D’. Ploco- 
spermeae. 

(18a) Plocosperma. Monotypic; Central 
America. Revision: LEEUWENBERG, Act. 
Bot. Neerl. 16 (1967) 56-61. 

Add to Desfontainea: Revised and placed 
in a separate tribe Desfontaineae by 
LEEUWENBERG, Act. Bot. Neer]. 18 (1969) 
669-679. 

Add to Retzia: Revised and placed in a 
separate tribe Retzieae by LEEUWENBERG, 
Act. Bot. Neer]. 13 (1964) 333-339. 
Add to Dispersal, 1stjsentence, after ‘bats’: 
and by Viverridae (‘luwaks’), cf. DOCT. v. 
LEEUWEN, Trop. Natuur 21 (1932) 142. 
Add to Anat.: See SminGH, Gard. Bull. 
Sing. 22 (1967) 193-212, for sclereids; 
BurGEss, Timbers of Sabah (1966) 385- 
388, for wood-anatomy. 

Add to 22, 2nd lead: (Confusion is 
possible with specimens of F. auriculata 
ssp. borneensis with very faint auricles 
and 1-3-flowered inflorescences.) 
Fagraea elliptica Roxs. 

Add to literature: BACK. & BAKH. f. 
Fl. Java 2 (1965) 211. 

Fagraea fragrans Rox. 

Add to literature: Troup, Silvic. Ind. 
Trees 2 (1921) 676; MITCHELL, Mal. 
For. 27 (1964) 127; BAck. & BAKH. /. 
FI. Java 2 (1965) 211; SmytTHiEs, Common 
Sarawak Trees (1965) 87; KENG, Ma- 
layan Seed Plants (1969) f. 140-141. 
Add to Distr. 2nd paragraph: MIQUEL’s 
citation from the Moluccas is apparently 
based upon De Fretes 5742 from Ambon 
U) 


Insert under Uses before ‘A decoction. .’: 
see SCHNEIDER, Bull. Bur. For. Philip. 14 
(1916) 202, f. 71. 

Fagraea racemosa JACK ex WALL. 

Add to literature: ANDERSON, Gard. 
Bull. Sing. 20 (1963) 192; Back. & BAKH. 
f. Fl. Java 2 (1965) 211. 

Fagraea ceilanica THUNB. 

Add to literature: BACK. & BAKH. f. FI. 
Java 2 (1965) 211. 

Add (sub F. litoralis Bu.): Doct. v. 
LEEUWEN, Trop. Natuur 21 (1932) 142; 
ANDERSON, Gard. Bull. Sing. 20 (1963) 
192. 

Add to Distr.: Bougainville (NGF 31290, 
‘ternatana’), Solomon Is. (New Georgia: 
WHITMORE R.S.S. 6361, ‘ternatana’; 
Guadalcanal: R.S.S. 86, ‘/itoralis’, R.S.S. 
98, ‘ternatana’). 

3rd paragraph, ‘F. oblonga’, 
distribution: Borneo (Sarawak). 
Fagraea acuminatissima MERR. 
Add to Ecol.: also on coastal rocks. 


add to 


6: 323a 


6: 324a 


6: 327b 


6: 328a 
6: 328b 


6: 332b 


6: 333a 
6: 335a 


6: 335b 


6: 336 


6: 337b 


6: 340a 


6: 343 


6: 345 
6: 346 


Add to description, Ist sentence: 
treelet (5 m). 

Add to Ecol. 3rd line, after ‘sandstone’: 
and limestone. 

Fagraea blumei G. Don. 

Add to literature: BAck. & BAKH. /. 
Fl. Java 2 (1965) 211. 

ssp. blumei. 

Add to Ecol., after altitude: 
Kinabalu up to 3000 m. 
Fagraea fastigiata BL. 

Add to literature: BACK. & BAKH. f. FI. 
Java 2 (1965) 211. 

Fagraea auriculata JACK. 

Add to literature: BACK. & BAKH. f. FI. 
Java 2 (1965) 211. 

Add to Distr.: Sumbawa, Flores. 

ssp. auriculata. 

Add to Distr.: Sumbawa, Flores. 

ssp. borneensis (SCHEFF.) LEENH. 

Read Inflorescences: (1—)3-7— flowered. 
Fagraea gracilipes A. GRAY. 

Add to literature: PARHAM, Pl. Fiji Is. 
(1964) 176, f. 64. 

Fagraea gracilipes A. GRAY. 

Add to Distr. sub Solomon Is.: Choiseul. 
Fagraea berteriana A. GRAY ex BTH. 
Add to literature (sub F. schlechteri GILG 
& BENED.): GUILL., THORNE & VIROT, 
Un. Iowa Stud. Nat. Hist. 20, 7 (1965) f. 
15 (p. 55). 

Ecol.: Change highest altitude into 1700 
m. Add: Flowers open in the morning, 
drop off in mid-afternoon. 

Buddleja Houst. ex LINNE. 

Replace ‘Spelled Buddleia by most au- 
thors’ by: Incorrectly spelled Buddleia by 
most authors; cf. Int. Code of Bot. 
Nomencl. (1961) Appendix II, Nomina 
Familiarum conservanda, sub Buddle- 
jJaceae. 

Add to synonymy: Toxina Norona, 
Verh. Bat. Gen. 5 (1791) 4, nom. nud. 
Buddleja asiatica Lour. 

Add to literature: BAcK. & BAKH. f,. 
Fl. Java 2 (1965) 212. 

Buddleja davidii FRANCH. 

Add to literature: BACK. & BAKH. f. 
Fl. Java 2 (1965) 212. 

Gelsemium JuSSIEU. 

Add to literature: DUNCAN & DE JONG, 
Sida 1 (1964) 346-357; ORNDuFF, J. Arn. 
Arb. 51 (1970) 1-17. 

Add to description: Seeds not winged in 
G. rankinii. 

Strychnos L. 

Add to literature: KRUKOFF, Mem. N.Y. 
Bot. Gard. 12, 2 (1965) 1-94; LEEUWEN- 
BERG, Med. Landb. Hogesch. Wagenin- 
gen 69 (1969) 1-316. 

Add to Distr.: Cf. VAN BALGooy, 
Blumea Suppl. 5 (1966) 256, map 141. 
Add to Taxonomy: A new subdivision 
has been proposed by LEEUWENBERG, /.c., 


or 


on Mt 


September 1972] 


6: 347 


Addenda, corrigenda et emendanda 955 


who distinguishes between 12 sections. 

As far as the Malesian species are con- 

cerned, these are the following: 

Sect. Strychnos: S. angustiflora, S. 

ignatii, S. kerrii, S. lucida, and S. nux- 

vomica. 

Sect. Penicillatae: S. axillaris, S. ridleyi, 

and possibly S. me/lanocarpa. 

Sect. Brevitubae: S. flavescens, S. 

luzonensis, and S. vanprukii. 

Sect. Lanigerae: S. borneensis, S. curtisit, 

S. lanata, S. lanceolaris, S. ledermannii, 

S. maingayi, S. minor, S. oleifolia, S. 

ovata, S. polytrichantha, S._ thorelii, 

S. villosa, and possibly S. rufa. 

Key to the species: 

Replace couplet 3, as follows: 

3. Petiole 2-4 mm. Inflorescences few- 
flowered. Style glabrous. Fruits 2—21/2 
cm @, pedicels only slightly thickened. 

3. S. lucida 

3. Petiole 4-11 mm. Inflorescences many- 
flowered. Style thin-woolly. Fruits 3—6 
cm @, pedicels thickened. 

3A. Leaves elliptic-oblong, 2—3 times as 
long as wide, shining green when 
dry. Calyx glabrous; corolla 14-15 
mm long, inside woolly at the 
mouth; anthers 1 mm. Fruit thick- 
walled. Paired tendrils frequent 
(always a liana) 23. S. kerrii 
3A. Leaves broad-ovate to elliptic, c. 
11/4-11/2 times as long as wide, dull 
green when dry. Calyx outside 
pubescent; corolla 10-12 mm long, 
inside woolly in the lower half of the 
tube; anthers 13/4 mm. Fruit 
thin-walled. No tendrils (mostly a 
tree) . . . . 2. S.nux-vomica 
Insert between 6 and 7: 
6A. Twigs sharply quadrangular, branch- 
es often rounded quandrangular. 
Dried leaves beneath mostly yellow- 
ish brown to copper-red. Anthers 
glabrous 12. S. vanprukii 

6A. Twigs and branches terete. Dried 
leaves beneath either dark brown or 
greenish. Anthers hairy. 

Add to 8, 2nd lead, at end of Ist sentence: 

(exceptionally glabrous). 

Replace couplet 12, as follows: 

12. Corolla 9-10 mm long. 

12A. Leaves elliptic, c. 21/2 times as 
long as wide, base cuneate, apex 
mostly distinctly acuminate. Calyx 
outside glabrous, sepals rounded 

to nearly truncate; corolla inside 
densely woolly from slightly above 

the base to halfway the lobes; 
pistil fairly densely hairy from 
about halfway the ovary upwards. 

13. S. borneensis 

12A. Leaves mostly ovate to broad- 
ovate, 1-2 times as long as wide, 

base (obtuse to) rounded to 


cordate, apex not to. shortly 
acuminate. Calyx outside puber- 
ulous, sepals acute; corolla 
inside woolly in the mouth and on 
the lower part of the lobes; pistil 
glabrous . 22. S. angustiflora 
12. Corolla up to 61/2 mm long. 
Replace in 14, Ist lead, ‘S. colubrina’ by: 
S. minor. 
Add to 14, 2nd lead, after ‘Anthers 
subsessile’: or filaments short. 

15, 2nd lead, 2nd sentence, read: 
Leaves 3—5—plinerved. 
Insert after the key to the species the 
following: 
With the key as published originally in 
the Flora Malesiana, only flowering 
material could be identified. Four species 
the flowers of which were known too 
incompletely or not at all (S. melanocarpa, 
S. quandrangularis, S. rufa, and S. 
thorelii) could then not be included. Out 
of these four, one (S. quadrangularis, now 
S. vanprukii) could be added in the above 
Addenda to the key as in the mean time 
the flowers became known. But fruiting 
and sterile material, more than half the 
collections coming in, could still not be 
named. For that reason I have compiled 
the following synoptic key to all Malesian 
species, making use of all main characters. 
This is a multiple entry key, which 
means that one can start with every 
character, and can make use of all 
characters available in the material. 
The numbers after each lead represent 
the species showing that character state: 
species in which that character is still 
unknown are included with a question 
mark. The best way to work with this 
key is first to try whether the specimen 
to be identified shows one or more ‘rare’ 
characters, to note down all numbers of 
species that may show that character 
(including those with a question mark), 
and, with each following character, to 
narrow down this series of possibilities. 
Finally, this will lead to one or a few 
species; comparison with the descrip- 
tions and with material will mostly be 
conclusive. If identification in this way 
does not lead to any species, you may try 
what species shows the smallest number 
of differences: it may be that your 
specimen represents a species not included 
in the key, but it is also possible that it 
shows a character state not yet known 
from one of the species. The numbers of 
the species are in accordance with the 
following enumeration. 

1. Twigs 

aliglabrousAds 3a4+54) oe okON EI. 

122 1S 14S ANG ANSA22° 235-24. 

b. hairy (mostly slightly and early 
glabrescent): 2. 3. 5. 6. 14. 16. 17. 18. 


956 


FLORA MALESIANA 


[ser. I, vol. 68 


OS 


co) 


LOX 2OR215 22% 
Thorns or spines 
presents 24.35 lsa22. 


. absent: all species. 


Leaves, shape (irrespective of place 
of greatest width) 
rhomboid: 18. 


. suborbicular (about as long as wide): 


DeSe WE Sy22: 

elliptic (+ 11/2 times as long as 
Wide) cele sn4. Sami ecrelile toes: 
14. 16. 18. 20. 22. 23. 24. 

oblong (+ 21/2 times as long as 
Wide) aula TOs ss OMAOM 1213: 
[AS Gi Sen ORO 23 245 
lanceolate (3 or more times as long 
as wide): 1. 4. 8.9. 11. 12. 15. 16. 18. 
Di 2D 

Leaves, place of greatest width 
about the middle: 1. 2. 3. 4.5. 6.7.8. 
LOMAS Ae Sel Ga ison 20! 
Mle pss 38). 

distinctly below the middle: 1. 2. 3. 
SHON OO MOA Gy 17. 18! 19: 
2ORAII22F 23824" 

distinctly above the middle: 11. 
Length of leaves 

Up toMlOicnil 2s S44 onGs 08. 110: 
eee al Saati Saal Oates 9) 20) 
De 22 23924) 

1O=Sveme le 2 Ay SaGass 9) 10) 11: 
P25 AG eS20221. 22823) 24: 
15-20 cm: 1. 6. 10. 11. 12. 18. 20. 
more than 20 cm: 1. 11. 

Width of leaves 

up to 5 cm: all species. 
S—OlemiPile2esn4s 52608. 10s 12: 
1ANIGI1 8520222573424" 

more than 10 cm: 1. 2. 11. 12. 
Leaves 

glabrous: 1. 2. 3. 4. 5. 6. 7. 8. 9. 10. 
LD AS 4a Sreliowlica ol 22. 
24. 


. not fully glabrous: 6. 17. 18. 19. 20. 
Dit, 


Lower side of leaf 

warty(-papillose): 3. 11. 18. 20. 
SMOODNMl AS Ones oO lO ndiie 
Ll SAS 6s lalSealOs 21) 22) 
2504) 

Leaf base 

anpilarswle 234445: 6s. 9, LON 11 
17.3: 14 AS Ges MOND 14 22) 23. 
24. 

rounded: tine as a4 aS NOne SH lO iie 
2A aGs 1 Sel OMORe 1 22" 23) 
24. 

(sub)cordate: 2. 3. 6. 11. 18. 20. 21. 
aby 


Leaf apex 

acuminate: 1.2.4.5. 6.7.8.9. 10. 11. 
IDRIS 14 iSaOmiia dss 19420) 2i- 
DO OBO A 

not acuminate: 1. 2. 3. 5. 10. 11. 12. 
GPUS 922923: 


11. 


Leaf apex or acumen 


ay acute: 14223574: 5: GalOmiieae rs: 


14. 
. terminal (either on a main branch 


LAS TSS Lor 17s l8 19 A2ORele 22503" 
24. 

blunt to rounded: 1. 2. 3. 5. 7. 8. 9. 
LOM 22. 

emarginate: 3. 18. 


. Number of main nerves 


32.1..2. 324.5 -9R1E eS eS. 
16.17.18. 19:20) 219228235245 
52:3. 526.758) 9 gM es. 
16:1718:.20. 21.2258: 

Te Mdia: 


. Petiole, length 


up tor4 mmia3, Sale 2 Se Ges: 
20. 24. 

4-71/2 mm: 1. 2. 4. 5.6. 7. 8. 9. 10. 
10512513) 14 1S AG ae Oe oe 
223i 

more than 71/2 mm: 1. 2. 5. 6. 9. 10. 
11512516; 18: 19921 eas 
Inflorescences 


or on a short axillary shoot with at 
least one pair of leaves; sometimes 
with a pair of basal branches in the 
upper leaf axils): 1. 2. 3.4. 5. 6.7. 8. 
Salil, 14: 167; ls2iee2 28h eAs 


. axillary, sometimes ramiflorous or 


together pseudoterminal: 1. 4. 5. 6. 
78.10) 115 12: ABA IS GateealS: 
20. 21. 24. 

Inflorescences, length 

up tos cmitle 2345557 cement. 
NUSy5 MIC a IMR iss, TIS P10), Ail P22. 228), 
24. 

5=10' cms 1.54.75 2168 8) SOM 2. 
1B), Moy 9/5 Il, 728), 

more than 10 cm: 6. 11. 


. Calyx outside 


hairy? 1. 2: 35:4: SaGas,Oniiealeelio: 
16M SP20R21 R22 

glabrous: 4. 6. 7. 10. 11. 12. 13. 14. 
18. 19. 21?. 23. 24. 

Calyx inside 

hairy 74: 6:12. 139195 202222 
glabrous: 1. 2. 3. 4. 5. 7. 8.9. 10. 11. 
12) 13.9145 IS oy se ZO ree) 
Dye), OEE 

Sepals 

acutes 12 2532556: 102 aleater 
18519920?) 217522824" 


. blunt, rounded, or truncate: 1. 2. 4. 


SH6D 78h 9 AO ey 12S ealeree eels 
18.202:212. 23:24, 

Corolla, total length 

up to 5mm: 4. 5. 6. 11. 12. 14. 15. 16. 
17. 18. 19. 20. 21?. 24. 


2 S=10imms 7. 899. LOPE Aeie. 


22» 

10 mm or more; 1. 2. 3. 21?. 22. 23. 
Corolla tube 

longer than limb: 1. 2. 3. 18. 20?. 217. 
wey 


. equal to limb: 11. 13. 14. 15. 16. 18. 


September 1972] 


Addenda, corrigenda et emendanda 


o57 


one 


22. 


19; 200; 217.122: 24: 


. shorter than limb: 4. 5. 6. 7. 8. 9. 10. 


27. 2025217. 


. Mature corolla outside 


hairy: 5. 6. 9. 15. 16, 18. 20?. 217. 
fully glabrous: 1. 2. 3. 4. 5. 7. 8. 10. 
ie 2a 47 182195 2009 20?. 
22. 23. 24. 

Corolla inside 

glabrous: 3. 16. 20?. 21?. 

with a whorl of bristle-like hairs in 
the mouth or up to halfway on the 
lobes: 17. 18. 19. 20?. 21?. 24?. 
woolly in the tube, sometimes ex- 
tending to the lobes: 1. 2. 3. 6. 8. 9. 
TOM MR1S 32.07.21): 

only woolly about the mouth: 4. 5. 
214s 116.202, 219222. 235247: 
woolly on the lobes, sometimes from 
the mouth upwards: 7. 11. 12. 15. 16. 
19. 20?. 217. 247. 


. Insertion of stamens 


inthe mouth: 1)..2) 3.4. 5s 6. 7.8. 9: 
LOM IOS 1AM Selo 202752 7. 
22N23) 

in the tube: 12. 14. 18. 19. 20?. 21?. 
2223. 24: 

Filament, length 

up to 1/2 mm: 1. 2. 3. 5.6. 11. 14. 15. 
16. 17. 18. 19. 20?. 21?. 23. 24. 
ol foomment. sa SnonlOwli. Wa lst 
LSS U8s 202. 217: 

11/2-21/2 mm: 4. 5. 7. 9. 12. 13. 20?. 
PAE 

21/2 mm or more: 4. 8. 12. 20?. 217. 
DD: 


. Anther 


OVvater 4) SO Ul 12s US16. Wie US: 
195202, 21. 24: 

elliptic? 25 5, 9F 1 1213145207: 
DIED 3: 

oblong: 1 37457510! MA12? 132207: 
P22 623: 

lanceolate: 4. 7. 8. 10. 11. 13. 207. 
21?. 


. Anther, length 


up to 1 mm: 4. 5. 6. 7. 11. 12. 14. 15. 
GHG UseOe 2075217. 23" 24: 
more than 1 mm: 1. 2. 3. 5. 8. 9. 10. 
MINIS 202 217h 22) 23% 


. Anther 


hairy: 4. 5. 6.8. 10. 11. 13.15. 16. 17. 
18. 19. 20?. 217. 24. 

elabrous:le2:3. 457. cool 2 13: 
1451162202221. 22.23. 


. Pistil, length 


up to 4mm: 4. 5.6. 11. 12. 14. 15. 16. 
17. 18. 19. 20?. 21?. 24. 
more than 4 mm: 1. 2. 3. 7. 8. 9. 10. 
AIS 2.024219 22523" 


. Ovary 


Hain yA KOM erOwo Ons 135 15. 
17. 20?. 21?. 24. 

Slabrouse le 20344. SaGai2. 14516. 
18. 19. 20?. 212. 22.§23. 


: 347a 


: 349b 


: 350a 


: 350b 
>) SI 


2 shlsye 


30. Style 

ay hairy 2 AS NGe Tash. 1113: 
(4. 15206217 72075:217, 23. 247. 

b: @elabrouss1) 3943'5.6)-11..12. 16. 18. 
19. 20?. 21?. 22. 247. 

31. Pedicels in fruit 

a. much thickened: 1. 2. 7?. 9?. 107. 13. 
14?. 15. 17?. 19?. 20. 23. 247. 

b. only slightly thickened: 3. 4. 5. 6. 7?. 
SiO? sOP aie 2er4? Gh 722 18: 
19?. 21. 22. 24?. 

32. Fruits 

a: Vglobulan?1) 253545556777; 897. 107. 
PQS: 4 eS Pom? Lee 9 2: 
20. 22. 23. 24. 

b. ellipsoid or ovoid: 1. 5. 6. 7?. 9?. 10?. 
LAP AUS Pa aS 2 

33. Fruits, diameter or length 

a.) upito 24/5 emir 3h4: 5165772977; 102. 
IE 2S 1421S oaks. 197. 
Pile Q2ADA 

b. more than 21/2 cm: 1. 2. 7?. 8. 97. 
ODA SP ate 2a 20) 212. 
23: 

34. Fruits 

a. thick-walled: 1. 7?. 9?. 10?. 14?. 15?. 
172 ASP 23H 24? 

b. thin-walled: 2. 3. 4. 5. 6. 7?. 8. 92. 
LO? UID. 1S 4 ispalGs 1772 1S: 
19?. 20. 21. 22. 24?. 

35. Number of seeds 

aot 16) 4S SG ia oe el Orentem 2. 
13% L416 Ba99821. 225 247 

bs more thani219243562 775 8. 9% 107: 
LTS PAA LS Oi Oe 2022 223: 
24?. 

1. Strychnos ignatii BERG. 

Add to literature: BACK. & BAKH. /. FI. 
Java 2 (1965) 210; TrrEL-ROUDET, Logan. 
Cambodge, Laos & Vietnam (1970) 59, 
t. 4 f. 6-8. 

Change S. philippinensis BLco into S. 
philippensis BLco. 

2. Strychnos nux-vomica L. 

Add to literature: Troup, Silvic. Ind. 
Trees 2 (1921) 673; PETELoT, PI. Médic. 
Cambodge, Laos & Viet-Nam 2 (1953) 
169; BAcK. & BAKH. f. Fl. Java 2 (1965) 
210; TirEL-ROUDET, Logan. Cambodge, 
Laos & Vietnam (1970) 71. 

Add to Distr.: Burma. 

3. Strychnos lucida R.Br. 

Add to literature: BACK. & BAKH. f. FI. 
Java 2 (1965) 210. 

4. Strychnos maingayi CLARKE. 

5. Strychnos ovata HILL. 

Add to literature: TrREL-ROUDET, Logan. 
Cambodge, Laos & Vietnam (1970) 55, 
tshiail=se 

Add to synonymy: Strychnos sp. MERR. 
Pl. Elm. Born. (1929) 252. 

Add to description: Young twigs some- 
times sparsely minutely hairy. Leaves up 
to 11 cm long. 

Insert after the paragraph on Ecology: 


958 FLORA MALESIANA [ser. I, vol. 66 
Uses. In Sarawak, Ukit nomads pre- Distr. S. and NE. India, Thailand, 
pare one of the elements of their dart poi- Laos, Vietnam, and Malesia: Malay 
son from this species. Peninsula (Perak, Selangor), Borneo 

6: 3515 6. Strychnos villosa HILL. (Sarawak, Central E. Borneo). 

Add to literature: BACK. & BAKH. f. FI. Ecol. Dense jungle, from the lowland 
Java 2 (1965) 209. to above 1000 m. F/. Aug., Nov. fr. Sept. 

6: 353a 7. Strychnos curtisii K. & G. Uses. The bark of the roots is used for 
8. Strychnos polytrichantha GILG. arrow-poison. See BURKILL, Dict. (1935) 

6: 353b 9. Strychnos oleifolia HILL. 2099. 

6: 355a 10. Strychnos lanata HILL. Vern. Akar ipoh, Mal. Pen. 

The correct name for species I1 is: 13. Strychnos borneensis LEENH. 

11. Strychnos minor DeNNst. Schluess. 6: 357a 14. Strychnos ledermannii GILG & BENED. 
Hort. Malab. (1818) 33; Busser & Add to description: Leaves sometimes 
PuiLtcox, Taxon 20 (1971) 537-543. — S. with 3 main nerves. 

colubrina Auct. non L.: Hitt, Kew Bull. 6: 3575 15. Strychnos lanceolaris Miaq. 

(1917) 157 et seg.; LEENH. FI. Mal. I, 6 16. Strychnos flavescens K. & G. 

(1962) 355; BAcK. & BAKH. f. FI. Java 2 Add to description: Leaves in a specimen 
(1965) 209; TireEL-RoubeT, Logan. Cam- from Borneo (Sarawak For. Dept. 
bodge, Laos & Vietnam (1970) 50, t. 13460) 5-plinerved, the outer nerves 
INE Sh. diverging at, the inner distinctly above 
Add to description: Leaves sometimes the base. Corolla of the same specimen 
not acuminate at apex; lower side mostly woolly inside all over the inner surface of 
smooth to rarely minutely papillose. the lobes; stamens with short filaments. 

6: 356a Add to Distr.: Lesser Sunda Is. (Flores). 6: 358a Add to Distr.: Borneo (Sarawak, Sabah). 

6: 356b Replace species 12 as follows: Add to Ecol.: Altitude up to 1350 m. F7/. 
12. Strychnos vanprukii Crais, Kew Bull. from March onwards, fr. Febr., Sept. 
(1911) 421; Hitt, Kew Bull. (1917) 139; 17. Strychnos luzonensis ELM. 
in Craib, Fl. Siam. En. 3 (1951) 62; 18. Strychnos axillaris COLEBR. 
TiREL-RoubeET, Logan. Cambodge, Laos Add to literature: BACK. & BAKH. f. FI. 
& Vietnam (1970) 48, t. 2 f. 1-7. — S. Java 2 (1965) 209; TireEL-RoubeT, Logan. 
aenea Hitt, Kew Bull. (1917) 138, cum Cambodge, Laos & Vietnam (1970) 37, 
fig., incl. also var. acuminata. — S. t. 12 f. 1-3. 
quadrangularis Hitt, Kew Bull. (1917) 6: 359a Add to Distr.: Lesser Sunda Is. (Flores: 
205; LEENH. FI. Mal. I, 6 (1962) 356. — S. KOSTERMANS 22156). 
maingayi CLARKE ssp. borneensis LEENH. 6: 3605 19. Strychnos ridleyi K. & G. 

Blumea 14 (1966) 230. 6: 36la 20. Strychnos rufa CLARKE. 

Climbing shrub or liana, provided with 21. Strychnos thorelii PIERRE ex Dop. 
double tendrils. Twigs slender, mostly Add to literature: TrREL-RoubDET, Logan. 
sharply quandrangular, glabrous; branch- Cambodge, Laos & Vietnam (1970) 82, 
es rounded quadrangular. Leaves 71+/2- iia UPA I Bh. 

20 by 21/2-11 cm, ratio c. 2-3, widest 6: 361b 22. Strychnos angustiflora Bru. J. Proc. 


about or sometimes below the middle, 
thin-chartaceous to papyraceous, be- 
neath copper-red to yellowish brown or 
sometimes green when dried, smooth and 
glabrous; base cuneate to rounded; apex 
short- to caudate-acuminate, acute; 
strongly 3—S-plinerved; petiole 4-5 mm. 
Inflorescences axillary, 11/2-6 cm long, 
lax, few- to many-flowered, puberulous or 
glabrous. Flowers 5-merous. Sepals c. 
1 mm, acute or blunt, outside glabrous, 
inside glabrous or nearly so. Corolla 3-4 
mm long, the tube 1/4-11/4 mm, outside 
glabrous, inside woolly about the mouth 
up to halfway the lobes and papillose 
hairy at the tips. Stamens inserted at or 
slightly below the mouth, filament 1-3 mm 
long, anther ovate, elliptic, or oblong, 
1/5-3/,4 mm long, glabrous. Pistil 3/4— 
23/4 mm long, glabrous. Pedicels in fruit 
hardly thickened. Fruits globular, c. 2 
cm @, thin-walled, said to be white. 
Seeds 1, semiglobular, c. 1 em @ and 8 
mm thick, glabrous. 


Linn. Soc. Bot. 1 (1856) 102; Fl. Hong- 
kong (1861) 232; Hit, Kew Bull. (1917) 
182 cum fig.; MERR. Lingn. Sc. J. 5 (1927) 
148; Herktots, Hongkong Natur. 4 
(1934) 108, f. 3; TrreL-RoubDeET, Logan. 
Cambodge, Laos & Vietnam (1970) 62, t. 
5 f. 1-5. — §. nux-vomica Auct. non L.: 
LEENH. FI. Mal. I, 6 (1962) 349 pro 
specim. Philipp. 

Liana, provided with simple tendrils 
and spines. Twigs puberulous, early 
glabrescent. Leaves 3-12 by 11/2-7 cm, 
ratio 1-2, widest below, sometimes about 
the middle, chartaceous to thin-coria- 
ceous, smooth and glabrous; base rounded 
to cordate, exceptionally obtuse; apex 
not or tapering short-acuminate, acute to 
blunt, mucronate; 3—5-plinerved; petiole 
4-10 mm. Inflorescences terminal on 
short axillary shoots, rather lax and 
fairly many-flowered, 2-4 cm_ long. 
Flowers 5-merous. Calyx outside puber- 
ulous, inside glabrous, sepals acute, 1—-11/» 
mm long. Corolla c. 1 cm long, the tube 


September 1972] 


Addenda, corrigenda et emendanda 


about as long as the lobes, outside 
papillose but not hairy, inside woolly in 
the mouth and on the lower part of the 
lobes. Stamens inserted about the mouth, 
filament 3-4 mm long, anther oblong, 
13/4-2 mm long, glabrous, blunt. Pistil c. 
1 cm long, glabrous. Pedicels in fruit only 
slightly thickened. Fruits globular, 2-4cm 
@®, thin-walled. Seeds 1 or 2 (rarely 
more), disk-shaped, c. 11/2-1?/4 cm @, 
sericeous. 

Distr. China (Kwantung), Hainan, 
Hong-Kong, Thailand, S. Vietnam, and 
Malesia: Philippines (Mindoro, Oriental 
Prov., Puerto Galera Bay, once collected). 

Ecol. Collected on the shore above 
tide level. Fr. April. 

Note. S. angustiflora is rather closely al- 
lied to S. nux-vomica and, whereas the dif- 
ferences in the flowers are clear, vegetativ- 
ely and in fruit they look much alike. Asa 
whole the leaves are broader, more ovate, 
and at base deeper cordate than in S. nux- 
vomica, but less so than in the also 
closely allied S. nux-blanda. It was Dr. 
N. G. Bisset, London, who, on account 
of the alkaloids found in the seeds, first 
expressed his doubt as to the identifica- 
tion of the Philippine specimen. 


23. Strychnos kerrii HitL, Kew Bull. 
(1925) 426; in Craib, Fl. Siam. En. 3 
(1951) 58; TrrEL-ROoUDET, Logan. Cam- 
bodge, Laos & Vietnam (1970) 78, t. 5 f. 
6-9. 

Liana, provided with double tendrils. 
Twigs glabrous, sparsely lenticellate. 
Leaves 7!/2-14 by 4-5 cm, ratio c. 2-3, 
widest about or sometimes below the 
middle, papyraceous to  chartaceous, 
smooth and glabrous; base cuneate to 
rounded; apex acute to abruptly acute- 
acuminate; 3 (rarely 5) main nerves; 
petiole 5-10 mm. /nflorescences terminal, 
dense and many-flowered, 4-6 cm long. 
Flowers 5-merous. Calyx glabrous on 
both sides, sepals rounded, | mm long. 
Corolla 10-15 mm long, the tube 4-5 
times as long as the limb, outside glab- 
rous, inside densely woolly in the mouth. 
Stamens inserted in or slightly below the 
mouth, filament very short, anther elliptic 
or oblong, 1-1/2 mm long, glabrous. 
Pistil c. 12-13 mm long, ovary glabrous, 
style hairy in the lower half. Pedicels in 
fruit strongly thickened. Fruits globular, 
3-5 cm @, the wall c. 1/2 cm thick. 
Seeds unknown. 

Distr. Assam, Burma, Thailand, Indo- 
China, and Malesia: probably in the 
Malay Peninsula. 

Ecol. Lowland rain-forest. 

Note. Dr. N. G. Bisset, London, in- 
formed me that he had seen sterile 
material of this species collected in the 


6: 


: 363a 


365 


: 365b 


: 366a 


: 3665 


: 367a 


Salta 


S/2a 


—3575a 


ons 


= 35a 


1a 


Malay Peninsula. 

24. Strychnos melanocarpa GILG & BE- 
NED. 

Gardneria ovata WALL. 

Add to literature: BACK. & BAKH. f. FI. 
Java 2 (1965) 210. 

Neuburgia BL. 

Add to Notes: Some species of Psychotria 
(Rubiaceae-Coffeoideae) show a_ great 
resemblance to Neuburgia. They differ 
mainly by the connate stipules not adnate 
to the petioles, by the cupular 5-toothed 
calyx, the glabrous anthers, and the 
triangular style with a truncate stigma; 
the ovary is for the main part superior but 
the fruits are inferior. 

Neuburgia corynocarpa (A. GRAY) LEENH. 
Add to Uses: In NE. New Guinea, 
Finisterre Mts, the wood was formerly 
used for making bowls and plates. 

Add to Notes: For an opinion as to 
specific delimitation in the Pacific 
contrary to the one expressed here see 
A. C. SMITH, Pac. Sci. 23 (1969) 387. 
Neuburgia sarcantha (GILG & BENED.) 
LEENH. 

Add to description: Leaves from 7 by 
51/2 cm, nerves 5-8 pairs. 

Neuburgia kochii (VAL.) LEENH. 

Add to description, Ist sentence: ap- 
parently sometimes a creeper (NGF 
14779). Add to leaves, entry on the base: 
rarely rounded to cordate. 

Neuburgia rumphiana LEENH. 

Add to Distr.: Terr. of New Guinea 
(Sepik Distr.). 

Neuburgia celebica (KOORD.) LEENH. 
Add to Distr.: New Britain, Solomon Is. 
Geniostoma rupestre FORST. 

Add to literature: BACK. & BAKH. /f. FI. 
Java 2 (1965) 207. 

Add to Notes, at the end of the Ist 
paragraph: An example of such a local 
subdivision has been given by BACKER & 
BAKHUIZEN f., J/.c., who distinguish 
among the material from Java between 3 
“microspecies’. 

Geniostoma arfakense KAN. & HAT. 
Add to description, under fruits: purple 
when ripe. 

Cynoctonum GMEL. 

The correct name and citation for the 
genus is Mitreola LINNE, Opera Varia 
(1758) 214, validated by indirect refer- 
ence to Gen. PI. ed. 1 (1737) 377. 
Cynoctonum mitreola (L.) BRITT. 

The correct name is: Mitreola petiolata 
(GMEL.) Torr. & GRAY, FI. N. Am. 2 
(1841) 45. 

Add to literature: BACK. & BAKH. f. FI. 
Java 2 (1965) 208; TrReEL-ROUDET, 
Logan. Cambodge, Laos & Vietnam 
(1970) 132, t. 9. 

Add to Distr.: W. Africa (possibly in- 
troduced, cf. Herne, Kew Bull. 17, 1963, 


6: 378a 


6: 378b 


6: 380b 
6: 381b 


6: 382a 


6: 383a 


6: 384a 


FLORA MALESIANA 


[ser. I, vol. 66 


17D) 
Cynoctonum sphaerocarpum LEENH. 
The name should be corrected as follows: 
Mitreola sphaerocarpa (LEENH.) LEENH., 
nov. comb. 
Add to Note, distribution of C. pedicel- 
latum: Assam (Khasi Hills). A third 
species of the same Asian relationship is 
Mitreola reticulata TirEL, Adansonia II, 
9 (1969) 119; Logan. Cambodge, Laos & 
Vietnam (1970) 136, t. 10, only known 
from N. Vietnam. 
Spigelia anthelmia L. 
Add to literature: BACK. & BAKH. f/f. FI. 
Java 2 (1965) 207. 
Add to Distr.: Introduced in Java 1845, 
cf. Back. Trop. Natuur Jubileumuitg. 
(1936) 54. 
Mitrasacme elata R.BR. 
Add to Distr.: Flores. 
var. brevicalyx LEENH. 
Add to Distr.: Flores, SE. New Guinea. 
Add to Ecol.: from c. 30 m onwards. 
Mitrasacme pygmaea R.BR. 
Add to literature: BACK. & BAKH. /. FI. 
Java 2 (1965) 208; TiREL-ROUDET, Logan. 
Cambodge, Laos & Vietnam (1970) 148, 
t. 11 f. 10-16. 
(sub M. nudicaulis Bru.): De Voocp, 
Trop. Natuur 30 (1941) 103, f. 4. 
var. pygmaea. 
Add to Distr.: Sumbawa. Drop: Mt 
Merapi, Bali, Lombok, Timor. 
Insert after 2. Mitrasacme pygmaea 
R.BR.: 
2a. Mitrasacme erophila Le&ENH. Bull. 
Jard. Bot. Brux. 32 (1962) 446, f. 72; 
TIREL-ROUDET, Logan. Cambodge, Laos 
& Vietnam (1970) 143, t. 11 f. 1-5. — M. 
pygmaea R.Br. var. pygmaea LEENH. FI. 
Mal. I, 6 (1962) 383 p.p., incl. f. 43. 
Erect, annual herb, up to 15 cm high, 
simple or branched at base. Leaves 
rosulate, (oblong-)lanceolate, 3-6 by 
3/4—2 mm, herbaceous, puberulous above, 
glabrous beneath, base cuneate, apex 
acute, 1-nerved. Jnflorescences laxly 
umbellate, few-flowered; peduncle 4-11 
cm long, terete, puberulous at base, with 
some scattered pairsof scale-like, 1—11/2 
mm long empty bracts; pedicels !/2—-21/e 
cm long. Ca/yx campanulate, 2 mm long, 
glabrous or puberulous, the lobes trian- 
gular, acute. Corolla widely infundibu- 
liform, 31/2-4 mm long, white, hairy in 
the throat, the lobes c. 11/2 mm long, 
oblong or obovate, rounded. Stamens 
inserted in the tube at c. 2/3 of the height; 
filament 1/2 mm long; anther orbicular, 
1/7 mm &, cleft at base, dehiscing all 
around, ciliate along the margin. Pistil 
2'/4 mm high, styles nearly completely 
connate, stigma deeply bilobed. Capsule 
globular, 11/2-2 mm @, styles short, free. 
Seed angular-ovoid, brown, reticulate. 


6: 385a 


6: 386a 


6: 386b 


Sea27b 


4: 45a 


5: 345 


Distr. Assam and Malesia: Central 
Java (Merbabu-Merapi), Lesser Sunda Is. 
(Bali, Lombok, Alor, Timor). 

Ecol. Grassland, Eucalyptus-forest 
etc., locally often common, from 650- 
2000 m, on Bali, Mt Agung, near solfata- 
ra, even at 3150 mm. Fi. fr. March-June. 

Note. Originally, when revising the 
genus Mitrasacme for the Flora Malesia- 
na, I did not recognize this material as 
different from M. pygmaea var. pygmaea, 
even though later onI described the pres- 
ent species as new from Assam. For 
differences with M. pygmaea see there. 
Mitrasacme indica WIGHT. 

Add to literature: BACK. & Baku. f. FI. 
Java 2 (1965) 208; TiREL-ROUDET, Logan. 
Cambodge, Laos & Vietnam (1970) 145. 
Add to synonymy: Non RIDSDALE, 
Trans. Papua N. G. Sci. Soc. 9 (1968) 18 
(=Lindernia subulata R.BR.). 
Mitrasacme saxatilis BACK. 

Add to literature: BACK. & BAKH. f. FI. 
Java 2 (1965) 208. 

Mitrasacme neglecta LEENH. 

Add to literature and synonymy: BACK. 
& BAKH. f. Fl. Java 2 (1965) 209. — 
Evolvulus sp. RIDSDALE, Trans. Papua 
N. G. Sci. Soc. 9 (1968) 17. 

Add to Distr.: SE. New Guinea. 
Mitrasacme bogoriensis LEENH. 

Add to literature: BACK. & BAKH. f. FI. 
Java 2 (1965) 209. 

Add to Excluded: 

Logania dentata (ELM.) HAYATA, J. Coll. 
Sc. Imp. Un. Tokyo 25, art. 19 (1908) 
162, t. 28. — Nertera dentata ELM. Leafl. 
Philip. Bot. 1 (1906) 15 = Hemiphragma 
heterophylla WA. (Scrophulariaceae). 
Cf. Merr. in Hayata, /c.; HALL. f/f. 
Beih. Bot. Centralbl. 39, 2 (1923) 161. 


Malpighiaceae (JACOBS) 


Aspidopterys elliptica (BL.) Juss. 

Add to Distr.: Lesser Sunda Is. (Bali, 
KOSTERMANS 272, distributed as Tristel- 
lateia; Flores, cf. STEEN. Blumea 15, 
1967, 153). 


Moringaceae 


Moringa oleifera LAMK. 
Add to synonymy: M. domestica HAM. ex 
HENSCHEL, Clavis Rumph. (1833) 44. 


Pittosporaceae 


In the revision of Pittosporum only 3 
endemic species were known from New 
Guinea which stands in contrast with the 
large number of endemic species in New 
Caledonia and also the much higher 
number in Australia. 

It has appeared, however, that further 


September 1972] 


5: 348 


5: 348 


exploration has yielded more novelties in 

the last decade, partly described by Dr. 

BAKKER, partly by Dr. ScHopbe. But I 

hesitate to share the view of the latter that 

East New Guinea would represent a 

‘centre of diversity’ for the genus. It may 

later appear that some novelties are 

possibly marginal extremes of other spe- 
cies and I feel that hybridization is also 
not excluded. | cannot recognize two of 

SCHODDE’s taxa at specific level. 

In the Key to the species (flowering 

material), replace the first entry of fork 5 

by: 

5. Flowers always axillary, solitary. 

5a. Leaves mostly small, obovate, c. 
2-4(-81/2) by 1-3(-4) cm, abruptly 
shortly acute-acuminate, coriaceous. 
Reticulation conspicuous, prom- 
inent on both sides. Twigs glab- 
rous. Flowers 12 mm. Ovary glab- 
rous. Fruit 12 by 8 mm. 

5. P. berberidoides 

Sa. Leaves small, obovate to spathulate, 
3-25 by 2-10 mm, glabrous, herb- 
aceous; apex broadly cuneate, with 
a blunt tip; nerves not prominent, 
few. Twigs covered by a persistent 
indument of short, thickish ferrug- 
ineous hairs. Flowers 8 mm long. 
Ovary with a few long hairs at the 
base . .  . P. inopinatum 

In the Key to the species (dry fruiting 

material), replace forks 5 and 6 by: 

5. Nerves and larger veins distinctly 
prominent on both leaf surfaces. 
Leaves obovate, + pointed, c. 2-4 by 
1-3 cm. 

5a. Main nerves c. 10-15 pairs, close. 
Leaf apex suddenly acute-acumina- 
tely contracted. Young twigs glab- 
rous 5. P. berberidoides 

5a. Main nerves 4-6 pairs. Leaves with 
rather bluntish apex, in pseudo- 
verticils. Young twigs densely fer- 
rugineous-hairy 5b. P. pumilum 

5. Nerves and larger veins not prominent 
above. 

6. Fruits longer than broad (not includ- 

ing the stipe and apical mucro). 

6a. Twigs densely persistently set with 
dark-brown thickish hairs. Leaves 
small, 3-25 by 2-10 mm, spathula- 
te, herbaceous, spirally arranged, 
with blunt tip; nerves 3-4 pairs, 
rather indistinct above (neither 
impressed nor prominent). Fruit 
valves thickish. 5a. P. inopinatum 

6a. Otherwise. 

7. Remains unaltered. 

7. Remains unaltered. 

6. Fruits as long as broad or slightly 

broader than long. 
7a. Fruits globular, 11-13 mm @, 
with thick valves. Leaves not in 


Addenda, corrigenda et emendanda 


961 


pseudo-verticils, very thick, small, 
+ bullate, 6-14 by 3-7 mm, with 
3-4 pairs of nerves which are im- 
pressed above. 
6a. P. pullifolium var. globosa 
7a. Leaves much larger with many 
more pairs of nerves which are not 
impressed above. 


5: 349a Pittosporum sinuatum BL. 


5: 349b 


5: 350b 


33-3516 


D324 


5: 353a 


Add to description: A specimen from 
East New Guinea (Brass 30801) has 
entirely glabrous, slender ovaries in pre- 
dominantly ¢ flowers; all other material 
has pubescent ovaries. 

Add after 1. Pittosporum sinuatum the 
following variety: 


la. var. efuniculare STEEN. var. nov. — P. 
tenuivalve SCHODDE, Blumea 15 (1967) 
AVipaiee: 

Funiculus brevipes. Typus ROBBINS 888. 

Falls within the variable P. sinuatum, 
differs obviously only by the short fu- 
nicles. 

Distr. Malesia: East New Guinea 
(Madang Distr., Kubor Range: Ros- 
BINS 888, 1121), not seen. 

Ecol. Montane forest, 2100-2300 m. 

Note. I agree with Dr. SCHODDE’s 
tentative suggestion that this represents 
perhaps only a mountain form of P. 
sinuatum. 

Pittosporum ramiflorum (ZoLL. & Mor.) 
ZOLL. ex MiqQ. 

Add to Distr.: Lesser Sunda Is. (Sum- 
bawa). Cf. BAKKER, Blumea 11 (1962) 
426. 

Line 6 from top, add after P. ramiflorum 
f. macrocarpum: Another specimen, 
again from NW. New Guinea, Vogelkop 
Peninsula, was collected (BW 6880). It 
can easily be distinguished from the 
equally large-fruited Philippine P. resini- 
ferum by the absence of the large resini- 
ferous cavities in the fruit valves. Cf. 
BAKKER, Blumea 11 (1962) 426. 
Pittosporum resiniferum HEMSL. 

Omit in key and description that fruit is 
compressed; this is only so in immature 
ones and caused by pressure. It is really 
very broad-ellipsoid, tending to globular; 
occasionally it is 3-valved. 

Add to Distr.: Solomon Is. (Bougainvil- 
le: SCHODDE 3767), climbing tree, in fr., 
800 m. Hitherto only known from N. 
Borneo and Philippines. 

Pittosporum berberidoides BURKILL. 

Add to description: Tree up to 14 m, 
stem to 20 cm @. Leaves up to 12 by 
41/5 cm; petiole to 2 cm. Cf. also BAKKER, 
Nova Guinea n.s. 9 (1958) 339; Blumea 
11 (4962) 426. 

Add to Distr.: Now found in many other 
places (Mt Wilhelm; Chimbu; Sepik- 
Wangi Divide; Goroka, efc.) and as low 


962 


FLORA MALESIANA 


[ser. I, vol. 66 


Fig. 11. Pittosporum inopinatum BAKKER. a. Habit, nat. size, 6. flower, <3, c. petal, «5, d. style, 7, 
f. fruit, nat. size, g. seeds, <2 (a-g ROBBINS 829), 


5: 353a 


down as 2400 m. 
Add after 5. Pittosporum berberidoides 
BURKILL: 


5a. Pittosporum inopinatum BAKKER, No- 
va Guinea n.s. 9 (1958) 339, f. 1. — 
Fig. 11. 

Much-branched shrub or dwarf tree, 
11/2-21/2 m. Twigs covered with a 
persistent indument of short, thickish 
ferrugineous hairs; internodes up to 
c. 11/2 cm. Leaves spirally arranged, 
herbaceous, obovate to  spathulate, 
entire, 3-25 by 2-10 mm, glabrous, 
apex broadly cuneate with a blunt tip, 
base decurrent, cuneate; nerves 3-4 pairs, 
indistinct above, not prominent on either 
surface; petiole c. 1-2 mm below the 
decurrent leaf base. Flowers solitary, 
axillary, pendent, c. 8 mm. Pedicels 
ferrugineous-hairy, c. 4 mm. Bracts 
triangular, very narrow, acute, c. 2 mm. 
Sepals free, narrow-oblong, acute, c. 2 
by ?/4-1 mm, yellowish green with a 
purple top. Petals free though cohering, 
ligulate, narrowing towards the rounded 
tip, c. 7-8 by 2 mm, pink or purplish 
especially at the top. Stamens sagittiform, 
filaments narrowing towards the anthers, 
c. 21/2 mm; anthers c. 1 by 1/2 mm. 
Ovary depressed-ellipsoid, glabrous, c. 
3 by 2 mm; style glabrous, c. 1 mm; 
stigma capitate. Fruit 2-valved, ellipsoid, 
c. 11/2 by 1 cm; valves coriaceous, 
orange-yellow to brown, rather thin; 
placentas slightly raised; funicles c. 10-12 


on each placenta, inserted up to ec. 2/3 et 
the length of the valves, in ripe seeds up 
to c. 3 mm long, very dark violet. Seeds 
c. 4-6 on each placenta, irregular- 
globose, dark violet, c.4 mm @. 

Distr. Malesia: East New Guinea 
(Mt Otto, 3 collections; Mt Piora, NGF 
16535). 

Ecol. Mountain forests of Podocarpus- 
Libocedrus, 2550-2860 m, said to be 
common. Fi. fr. Sept. 


5b. Pittosporum pumilum SCHODDE, Blu- 
mea 15 (1967) 406, f. 1. 

Slender shrub, 1 m. Twigs densely 
ferrugineous-hairy; internodes with a 
few tiny cataphylls. Leaves mostly in 
pseudo-verticils, spathulate, with rather 
rounded apex, glabrous, thinly coria- 
ceous, 11/2-41/2 by 1/2-2 cm; nerves 4-6 
pairs, on both surfaces prominent; base 
attenuate, a petiole hardly discernible. 
Fruit solitary, pseudo-terminal, later 
subterminal, on a rather stout glabrescent 
peduncle 5-8 mm long, ellipsoid, when 
young shortly stipitate and rostellate, 
21/2-3 by 11/2 cm, glabrescent, red 
(drying orange-yellow); valves outside 
rugose, inside without transverse ribs and 
set with funicles all along their length. 
Seeds c. 8, semi-reniform, 5mm, on rather 
long funicles, reddish brown, drying 
black. 

Distr. Malesia: East New Guinea 
(Morobe Distr.: Mt Shungol), one col- 
lection. 


September 1972] 


Addenda, corrigenda et emendanda 


963 


5: 3536 


5: 354a 


5: 3554 


5: 356b 


5: 3605 


Ecol. Lower mossy forest, 2100 m. Fr. 
Dec; 

Note. According to SCHODDE most 
related to P. sinuatum, but to my opin- 
ion most allied to P. berberidoides 
through the remarkably prominent veins. 
Pittosporum pullifolium BURKILL. 

Add to description: A specimen with 
young fruit had the _ infructescences 
axillary on the twigs instead of terminal. 
Cf. BAKKER, Blumea 11 (1962) 426. 

Add after 6. Pittosporum pullifolium the 
following variety: 


6a. var. globosum STEEN. var. nov. — P. 
nubicola SCHODDE, Blumea 15 (1967) 411, 
f. 3-4. 

Differt ab species: Capsula_ globosa, 
I]-Il3 mm 2. 

Shrub 1-2 m. Leaves thick, bullate by 
recurved margins, c. 10 by 5-6 mm; 
midrib and c. 3 pairs of nerves impressed 
above. Infructescence of c. 5 fruits apical; 
capsules globose, 11-13 mm @. 

Distr. Malesia: East New Guinea 
(Kubor Range: Mt Kinkain), one collec- 
tion. 

Ecol. Border of alpine shrubbery 
adjacent to peaty grassland, at 3600 m. 
Fr. July. 

Note. As SCHODDE already remarked 
close to P. pullifolium, of which the alpine 
dwarfed forms have similarly small 
leaves, and in fact grew to near 100 m 
from this form which differs by the globo- 
se fruit. 

Pittosporum pentandrum (BLANCO) MERR. 
Add to Distr.: N. Borneo. Cf. BAKKER, 
Blumea 11 (1962) 426. 

Pittosporum moluccanum (LAMK) Mia. 
Add to synonymy: Vareca moluccana 
Roxs. FI. Ind. ed. Carey 1 (1832) 647. 

In Fl. Mal. I, 5 (1954) 33, this was 
excluded from Hydnocarpus and _ tenta- 
tively assigned to Rinorea. ROXBURGH’S 
type (BM) was kindly identified by Mr. 
H. K. Atry SHAW (in /itt. to Dr. JACoBs, 
cf. Blumea 15, 1967, 138). No change of 
epithet is necessary. 

Add to Excluded: 

Pittosporum serrulatum JACK ex ROXB. 
Fl. Ind. 2 (1824) 401; ex Grirr. Calc. J. 
Nat. Hist. 4 (1843) 195; cf. Merr. J. Arn. 
Arb. 33 (1952) 240= Rinorea lanceolata 
(Roxs.) O.K. (Violaceae). 


Podostemaceae 


Add to Distr.: The knowledge of the 
distribution of the family in Indo- 
Australia is still expanding. A new spe- 
cies, of Indotristicha, has been found in 
Malaya. TUYAMA & Hara (J. Jap. Bot. 
39, 1964, 185-188) mapped Asian local- 
ities, which comprise amongst others 


localities in the East Himalaya, the 

Chinese mainland in Fukien, Kwantung, 

Kanton (TUYAMA & Hara, l/c. 185; 

CHAO, Contr. Inst. Bot. Nat. Peiping 6, 

1948) and Hainan (CHUN, FI. Hainanica, 

1964, 373), Thailand (VAN ROoyEN, 

Blumea 10, 1960, 141; Dansk Bot. Ark. 

23, 1965, 185) and the Ryu Kyu Is. Prob- 

ably the range of the family extends by 

several genera all over SE. and E. Asia 

(tropical to warm-temperate), but the 

small plants are evasive to collectors. 

Recently, an undoubted member has also 

been found by Mr. Byrnes in the Kim- 

berley District, NW. Australia; this is 
distinctly different from Torrenticola 
from Queensland. It bears resemblance to 

Indotristicha malayana, but has only 2 

stamens. Miss ASHTON, Melbourne, has 

tentatively referred it to Tristicha trifaria 

(Bory ex WILLD.) SPRENG., a species 

known from the tropics of America and 

Africa. Replace the Key to the genera 

into the following: 

1. Leafy (flowering) stems very short, 
hardly 10 mm long, with scattered, 
imbricate, 3—7-segmented leaves. Spa- 
thella oval, nippled, usually dehiscing 
irregularly. Tepals 2, narrow. Stamen 
1. Stigma 2 avis, like Gladopus 

1. Leafy (flowering) stems 25-100 mm, 
often branched. Leaves distichous, 
laterally compressed, the lower entire, 
the upper 3(-4)-dentate. Spathella oval, 
tipped, irregularly circumscissile-de- 
hiscent. Tepals 2, narrow. Stamen |. 
Stigmas 2. . . 2. Torrenticola 

1. Leafy shoots up to 20 mm, rarely 
branched, with imbricate triangular 
leaves in 3 ranks, those of 2 ranks 11/2 
by !/2 mm, those of the 3rd rank 1 by 
3/4 mm. No spathella. Tepals 3, half- 
way connate. Stamens 3. Ovary 9- 
veined. Stigmas 3. 3. Indotristicha 

Cladopus H. MOLL. 

Add to Distr.: SE. Asia (Siam, Hainan, 

Kwantung, Kanton, Fukien) and Ryu 

Kyu Is. Possibly monotypic. 

Cladopus nymani H. MOLL. 

Add to Distr.: Possibly all the names and 

localities in SE. & E. Asia of Cladopus 

refer only to one species; see VAN ROYEN, 

Dansk Bot. Ark. 23 (1965) 185. 

Add after 2. Torrenticola etc.: 


3. INDOTRISTICHA 


VAN RoyeEN, Acta Bot. Neerl. 8 (1959) 
474; Baku. f. Taxon 18 (1969) 598. — 
Dalzellia (non WiGHT) ENGL. Nat. Pf. 
Fam. Nachtr. 3 (1908) 135-136; ibid. ed. 
2, 18a (1930) 33, f. 24. 

See for the main characters in the key. 

Distr. Species 2, one in India (W. 
Ghats and S. Canara to Travancore), one 
in Malaya. 


FLORA MALESIANA 


[ser. I, vol. 66 


Fig. 12. Indotristicha malayana DRANSF. & WHITMORE. a. Habit, sterile, < 3, b. ditto, fertile, = 3, c. tepals, 


<7, d-g, four stages in the development from flower to fruit, » 14. 


1. Indotristicha malayana DRANSF. & 
WHITMORE, Blumea 18 (1970) 154, pl. 1, 
f. 1. — Fig. 12. 

Rhizome creeping, flattened, thalloid, 
closely adpressed to the substratum, 
1/2 mm wide, occasionally to 1 mm, of 
indefinite length, branching irregularly. 
Leafy shoots rarely flowering, borne 
irregularly, usually closely, to 2 cm long, 
rarely branching, usually curved, pale 
green and attenuate in shade, tinged red 
and shorter in full sun, irridescent. Stem 
triangular in section. Leaves sessile in 
very close whorls of 3 unequal ranks, 
the bases touching, increasing in size to 
about two thirds of the length from the 
rhizome, and thence decreasing slightly; 
two ranks long triangular to 11/2 by 
'/o mm, outline slightly asymmetric 
recurved, convex towards the third rank 
which is broadly triangular to 1 by 0.7 
mm. Also with flower bearing leafy 
shoots, often clustered, with smaller 
rather distant leaves. Flowers protogyn- 
ous, lateral, solitary; peduncle slender, 
stiff, erect, to 5 mm long at anthesis, 
subtended by two unequal, concave, 
sometimes mucronate, chartaceous im- 
bricate bracts to 1 by 1/2 mm. Tepals 3, 
valvate, obovate, 1 by 0.8 mm, slightly 
concave, chartaceous, translucent. Sta- 
mens 3, at first hidden by tepals, later at 
anthesis longly exserted to 1 mm on stiff, 
erect, pink filaments, later shrivelling and 
becoming thread-like and twisted ; anthers 
ovate-oblong, 0.4 mm long, cream. Ovary 
pale straw-coloured, obovoid, becoming 
narrowly obovoid by anthesis to 1.6 by 
1 mm with 9 rib-like veins; styles short, 
pink, 3, flattened in a vertical plane, with 
a deeply divided outer margin; placenta- 
tion free central, ovules numerous, tiny, 
cylindrical. Capsule narrowly obovoid, 


4: 258b 


6: 180a 


2 by 1.3 mm, light brown, crowned by 
remains of styles. 

Distr. Malesia: Malay Peninsula 
(Pahang, Trengganu), at least 3 localities. 


Pontederiaceae 


Monochoria hastata (L.) SOLMS. 

Add to Distr.: Now also recorded from 
Australia: Northern Territory, Litch- 
field Homestead. Cf. Muelleria 2 (1971) 
134. 


Primulaceae 


Lysimachia capillipes HEMSL. 

Add to Distr.: East New Guinea (Moro- 
be Distr.: Bulolo, NGF 30727, Wau, 
TGH 11596, NGF 35800; Eastern Highl. 
Distr.: NGF 27158), 100-1700 m. Fi. fr. 
Jan.-July. 

Note. The Papuan material is homo- 
geneous and is tentatively best classified 
with the Chinese-Luzon distributed L. 
capillipes. It must be remarked that the 
anthers have no permanent apical pore; 
they seem to dehisce with an apical slit- 
like pore which later extends downwards 
to a completely lengthwise splitting of the 
anther-cells. The calyx is narrow in all 
specimens (in contrast with that of L. 
laxa BAUDO) but its length seems to vary 
in proportion to the length of the capsule. 
It seems to be not always exceeding the 
capsule, but this could not well be 
checked as most capsules were dehisced 
and the more or less flattened valves are 
of course much longer than the undehisc- 
ed capsule. The Papuan specimens are 
more-stemmed but rather erect. More 
and better preserved material is desirable 
for a definite conclusion. — L. R6st, 
Oct. 1971. 


September 1972] 


Addenda, corrigenda et emendanda 


965 


6: 1855 


6: 192 


SAD 
5: 192a 


Sie 4b 


5: 444 


Lysimachia decurrens Forsv. f. 

Add to Distr. and map (fig. 10): SW. 
Central Celebes (Latimodjong Ra., KJELL- 
BERG 1425). Cf. BENTVELZEN, Blumea 13 
(1965) 140. 

Add to the excluded names: 

Hottonia sessiliflora VAHL, Symb. 2 
(1791) 36, the type of which was describ- 
ed from ‘India orientalis’ (C), from 
BURMAN’s herbarium; it belongs accord- 
ing to VAN DER MEWDEN (Blumea 17, 
1969, 311) to a species of Limnophila 
(Scrophulariaceae). The epithet is al- 
ready occupied in that genus. 


Proteaceae (SLEUMER) 


Helicia rufescens PRAIN. 

Add to Distr.: Borneo (Brunei). 
Heliciopsis rufidula SLEUM. 

Add to Distr.: Borneo (Sarawak). 


Rhizophoraceae 


Range maps of many species were given 
by Dinc Hou in Sreen. Pacific Plant 
Areas | (1963) maps 2-10, 20, 23. 
Distribution past and present. The second 
paragraph refers to the peculiar distribu- 
tion of mangrove species, Rhizophoraceae 
and otherwise, of which no species is in 
common to those in the Atlantic and In- 
dian Oceans. As a matter of fact this 
holds also for marine Phanerogams. On 
the other hand it has appeared that at 
least of the genus Rhizophora the three 
species of the Caribbean occur also on the 
west coast of Africa and that at least two 
of these also occur on the Pacific coast 
of tropical America. Cf. Dinc Hou, 
Blumea 10 (1960) 625-634. This means 
that there must have been in the past an 
open seaway between the Americas and 
this is corroborated by the geology. 
Africa seems to have been a distinct bar- 
rier between the Indian and Atlantic 
Oceans. 

I have discussed these major phytogzo- 
graphical features in 1962 in a pader 
(Proc. Kon. Akad. Wet. A’dam ser. C, 
65: 164-169) in which it was also shown 
that the marine Rhizophoraceae seem to 
have originated in the Indo-Malesian 
tropics where all 4 genera are centred, the 
Americas having only 3 out of 7 species 
of the one genus Rhizophora. 

As to the African barrier, it seemed to 
me that whereas even today species of the 
shore plants of the pes-caprae and 
Barringtonia formations cannot round the 
Cape of Good Hope because of temper- 
ature conditions and oceanic circulation 
system, the temperature must have been 
the chief factor that the dispersal of more 
megatherm mangrove species — of 


5: 445 


5: 448 


which dispersal is far more restricted by 
the viviparous drifting embryos which 
can only disperse in non-turbulent water 
and near-shore shallow muddy coasts — 
could not be effective even under the 
more ameliorated Tertiary climatic con- 
ditions. 

Mr. MuLLER (Review Palaeobot. 
Palyn. 6, 1968, 281-282) correctly stated 
that there was a northern open seaway, 
the Tethys, between the Indian and At- 
lantic Oceans and that this must have 
had an impact on diffusion of seaborne 
dispersed marine Phanerogams. He 
derives that the replacement of Atlantic 
and Indian Ocean species — if we pre- 
sume that the Tethys was an effective 
dispersal route indeed (including suitable 
ecological shore conditions!) — must be 
ascribed to the assumption that these 
Phanerogams had not yet evolved (or 
were at least not yet occurring in the 
Indian Ocean — y.St.) in the Early 
Tertiary and that when they came there 
the Tethys had by that time lost its 
capacity for their dispersal. That the 
Tethys must have had this function 
seems to be proved by the recent find of 
fossil Nypa pollen in West Africa (Upper 
Cretaceous to Upper Eocene), a genus 
which obviously got extinct there and in 
the Caribbean simultaneously for un- 
known reasons. 

Unfortunately we have no clear view 
of the course of the sea current regime 
at the time of the Tethys Sea. 

In addition we should mention that Rhi- 
zophoraceous pollen is only found as early 
as the Upper Eocene, but not (yet) in Paleo- 
cene or Upper Cretaceous sediments Cf. 
MULLER, Biol. Rev. 45 (1970) 434, f. 5. 

One important conclusion can, I be- 
lieve, not be doubted, except by super- 
stitious diffusionists, namely that dispers- 
al of both mangroves and marine sea- 
grasses can only take place at short 
distances and their occurrence is testi- 
mony of near-landmasses in the past, in 
contrast to most beach and Barringtonia 
formation species which are indeed 
frequently equipped to perform long- 
distance waterborne dispersal. 

Add to Taxonomy: According to Mr. 
MULLER (in Jitt.) the pollen of Carallia is 
similar to that of marine genera (Rhizo- 
phora, Bruguiera) and does not warrant a 
separation into a distinct family. 
Rhizophora L. 

Add to Distr.: DING Hou has revised the 
entire genus Rhizophora after the account 
in Fl. Mal. was written. Though this has 
brought no changes in taxonomy or 
distribution, some ranges in the West 
Pacific appear to be more extensive: 
R. mucronata LAMK extends to Tonga, 


966 


FLORA MALESIANA 


[ser. I, vol. 68 


5: 453b 


5 457 


5: 46la 


5: 464a 


5: 468b 


5: 474 


5: 477a 


5: 477a 


5: 480 


5: 483 


R. apiculata Bui. to the Solomons and 
New Hebrides, R. stylosa GRIFF. is also 
in Micronesia (Guam & Marshalls). Cf. 
DinGc Hou, Blumea 10 (1960) 625-634, 
map. 
The proper references of Rhizophora 
mucronata seem to be: Porr. in LAMK, 
Tabl. Encycl. Méth. Bot. (text) 2 (1794) 
517; LAMK, Tabl. 1 (1797) t. 396 f. 2; 
Porr. in LAMK, Encycl. 6 (1804) 189; DC. 
etc. 
Bruguiera LAMK. 
The date of publication of the genus is: 
1797. 
Bruguiera gymnorrhiza (L.) LAMK. 
In references line 2 the year of publication 
is: 1798. SAVIGNY seems to be responsible 
for the treatment of the genus in LaA- 
MARCK’S work. 
Bruguiera exaristata DING Hou. 
Add to synonymy: B. eriopetala var. 
exsetata VALETON, Bull. Dép. Agr. Ind. 
Néerl. 10 (1907) 38 (type: Kocu s.n., L). 
Bruguiera hainesii C. G. ROGERS. 
Add to Distr.: Also found in the Sundri- 
bans. Cf. S. K. MUKERJEE, Bull. Bot. 
Surv. India 8 (1966) 357. 
Anisophyllea R.BR. ex SABINE. 
Add to description of embryo: This is 
solid and indeed consists of an almost 
undifferentiated hypocotyl which con- 
tains the reserve food, similar as in several 
other tropical tree genera e.g. Barring- 
tonia and Bertholletia. From this ‘pre- 
adapted’ structure one can imagine the 
origin of the further differentiated em- 
bryogeny of the marine species. 

Anisophyllea beccariana BAILLON. 

Add to Distr.: Central East Sumatra 

(Tenajan R., SOEPADMO 37). 

Anisophyllea ferruginea DING Hou. 

Add to description: Mature leaves gla- 

brous. Inflorescences up to 4cm. Fruits 

1(-2)-seeded, 8 by 4 cm, glabrous; 
pericarp 8-10 mm @, exocarp soft. Seeds 
very hard, solid and large, 51/2 by 2 cm. 

The two additional specimens (ASHTON 

BRUN 580, S 7867) have mature in- 

florescences and fruit. In mature state the 

rusty tomentum disappears. 

Combretocarpus Hook. f. and C. rotun- 

datus (MIQ.) DANSER. 

Add to Distr.: Malay Peninsula (Johore), 

in peat swamp forest of Ayer Hitam 

South For. Res., occupying c. 30% of 

the big trees in the Reserve. Cf. F.S.P. 

Na, Mal. For. 29 (1966) 32-33, 1 fig. 

Carallia Roxs. 

Replace in the Key forks 5 & 6 into the 

following: 

5. Petals persistent. Calyx lobes hairy 
inside or at least at base or margin. 
Seed obovoid or ellipsoid. Embryo 
curved. 

6. Leaf margin distinctly fimbriate. 


5: 484a 


5: 485a 


Disk slightly crenulate. Fruit more 

than twice as long as wide, c. 15 by 

6mm 6. C. suffruticosa 
6. Leaf margin entire or almost so, at 
most very short-dentate or crenulate. 

Disk distinctly lobed halfway. Fruit 

less than twice as long as wide. 
6a. Cymes only in the axils of the upper 
pair of leaves of each shoot. Stip- 
ules up to 16 mm long. Calyx 
lobes 3-4 mm. Petals 3-4 mm. 
Seeds c. 9 by 6 mm. 5. C. borneensis 
Cymes in the axils of successive 
leaf pairs of each shoot. Stipules 
c. 5 mm long. Calyx lobes c. 11/2 
mm long. Petals c. 21/4 by 1 mm. 
Seeds c. 8 by 3 mm. 5a. C. longipes 
5. Petals caducous. Calyx lobes glabrous. 

Seed reniform or curved. Embryo 

curved. 

Carallia eugenioidea KING. 

Add to Distr.: Indo-China (Vietnam). 
Cf. VU VON CUONG, FI. Camb. Laos & 
Vietn. 4 (1965) 172. 

Add the following species: 

5a. Carallia longipes DING Hou, Nova 
Guinea, Bot. n. 4 (1960) 21-23, f. 4. — 
Fig. 13. 

Shrub, 3-5 m. Leaves chartaceous to 
thin-coriaceous, elliptic-oblong or ellip- 
tic, sometimes ovate, 31/2-61/2 by 11/2- 
31/2 cm, base cuneate rarely rounded, 
apex acute, margin obscurely serrulate in 
upper half; nerves 5—7 pairs, + elevated 
on both surfaces, veins reticulate, + 
elevated above, visible or obscure be- 
neath; petiole 5-10 mm; 2-4 small, with 
conical to subulate appendages at the 
base on each side, persistent but breaking 
off easily. Stipules c. 5 mm. Inflorescences 
up to 7 cm long, axillary, solitary, on 
several nodes along the young branches, 
usually few-flowered cincinnal cymes; 
internodes distinct, up to 11/2 cm, very 
rarely with a few short internodes between 
the long ones; peduncles 1!/2-4 cm. 
Flowers sessile, not resinous, slightly 
obovoid-oblong, c. 6 by 3 mm. Calyx 
lobes 5, ovate, c. 1.6 by 1 mm, acuminate, 
sparsely puberulous on the margins and 
towards the base inside. Petals persistent, 
obovate-spathulate, c. 21/4 by 1 mm, 
unguiculate, lamina short-fimbriate. Fi/- 
aments 31/2-41/2 mm, the lower 2/5 
united into a tube, free parts fleshy; 
anthers small, c. 0.4 by !/4 mm, obtuse. 
Disk fleshy, distinctly 10-lobed. Ovary 
l-celled, 10-ovuled; style cylindric, c. 
31/2 mm, slightly and gradually thickened 
towards the base; stigma obscurely 
capitate. Fruits dark-red, broadly el- 
lipsoid or subglobose, 13-15 by 10-13 
mm. Seed 1, oblong-obovoid, slightly 
ribbed, c. 8 by 3 mm, reddish to dark 
brown. Embryo straight. 


6a. 


September 1972] 


Addenda, corrigenda et emendanda 


Fig. 13. Carallia longipes DING Hou. a. Habit, 2/3, b. apex of branchlet showing appendages outside 


the base of the petioles and stipule, 


<2, c. inside view of stipule showing colleters, «7, d. node of 


branchlet, <3, e. flower, <3, f. longitudinal section of flower (style removed), «7, g. petal, «13, A. 


stamen, x 13, 7. style and stigma, <7, /. cross-section of ovary, <7, k. yGung fruit, 


5: 4885 


5: 491b 


SPIE eee Se 


(a & e-k BW 4980, b-d & | BW 7371). 


Distr. Malesia: West New Guinea 
(Vogelkop Peninsula). 

Ecol. Lowland forest, on limestone, 
275 m. 

Note. Closely related to C. papuana 
DING Hou. 

Gynotroches axillaris Bu. 

Omit under Distr. ‘Australia’; cf. DING 
Hou & STEEN. Pac. Pl. Areas 1 (1963) 
284, map 20. 

Though the distribution was correctly 
mentioned under the genus, Australia 
was by error mentioned in the distribution 
of the species, which induced Miss 
BURBIDGE to enter this generic record in 
the Dict. Austr. Pl. Gen. (1963) 141. 
It is of course not excluded that this may 
be found in future in the N. Queensland 
rain-forest, as it is abundant in New 
Guinea and the Solomons, but at present 
there is no proof for this. 

Pellacalyx saccardianus SCORT. 
Add to Distr.: Also in Peninsular Thai- 
land (near Trang, RFD 35161). 


Sarcospermataceae 


4: 3465 Sarcosperma uittienti H. J. LAM. 


S297. 


Add to Distr.: In addition to the 2 spe- 
cimens mentioned by H. J. LAM & 
VAROSSIEAU, Blumea 3 (1938) 194: 
Sumatra (Eastcoast, Wampa Valley, 
550 m, GALOENGI 432; Karo country, 
Biang Valley, 800 m, LORZING 14457) and 
Malay Peninsula (Selangor, Ulu Gom- 
bak, 600 m, SF 34191 Mow. Nur, FRI 
1950 FRANCIS NG). 


Scyphostegiaceae 


Scyphostegia STAPF and S. borneensis 
STAPF, — Fig. 14. 

At my suggestion Dr. W. A. VAN HEEL 
made a detailed anatomical-morphologi- 
cal investigation because Dr. J. Hut- 
CHINSON in the 2nd edition of his ‘The 
Families of Flowering Plants’ 1 (1959) 
326-329, f. 187 a—b, did still accept a 
disharmony between the 3 and ° flowers, 


968 FLORA MALESIANA [ser. I, vol. 68 


Fig. 14. Scyphostegia borneensis STAPF. a. Ls. 
of the ovule at the stage of the first formation of 
nuclear endosperm, the funicle shows a con- 
striction just below the meristem (indicated by a 
cross) of the later formed arilloid, x20, b. l.s. of 
young seed, with the arilloid developed above the 
constriction, embryo still small, 3, c. fully 
mature, dehisced fruit, the fleshy pericarp valves 
each with a part of the stigma on top, the bunch of 
dark, erect seeds attached to the base of the fruit, 
at base each surrounded by the (pale) arilloid, 
x 4/3 (after VAN HEEL). 


the latter being accepted as a fleshy 
receptacle containing numerous achenes, 
thus remaining at variance with SwAmMy 
(Proc. Nat. Inst. India 19, 2, 1953, 
127-142) and myself (Fl. Mal. I, 5, 1957, 
297-299), who interpreted the fruits 
(‘achenes’) of HUTCHINSON as ovules. 
Furthermore, HUTCHINSON placed the 
family in Celastrales, near Siphonodon, 
probably because of the similarity of the 


6: 193 


thickened stigma. 

VAN HEEL has found (Blumea 15, 1967, 
107-125, 13 fig.) that the reproductive 
units are doubtless seeds, that Hut- 
CHINSON’S ‘receptacle’ is a pericarp, and 
that his assumed ‘tepals’ in the 2 flower 
are really an aril(loid). Besides, the 
ovule shows some remarkable char- 
acters, namely a pedestal funicle with a 
constriction, a protuberance on the 
chalazal side, and a 5-lobed exostome. 

In my description of the fruit I had 
already observed that in later stages it 
was breaking up at the mouth. Dr. W. 
MEIER, to whom we are extremely 
obliged for the very complete pickled 
material, succeeded in finding in Sabah 
the final fruiting stage in which the 
pericarp is split to the base into 9-12 
valves; it is really a fleshy capsule, but is 
probably swallowed by animals in im- 
mature state as happens with so many 
fleshy fruits in Malesia. Fig. 14. 

Our suggestion of affinity with Fla- 
courtiaceae seems to be not unreasonable 
to VAN HEEL, from the morphological 
viewpoint the only discrepancy being that 
the placentation in the latter family is as 
far as known lateral-parietal and never 
basal as in Scyphostegia. But the pla- 
centation in Tamaricaceae, also listed in 
Parietales, seems intermediate, according 
to VAN HEEL, who will pursue further 
studies in Flacourtiaceae for checking 
this. 

It is curious to find that HUTCHINSON 
even in his latest work ‘Evolution and 
Phylogeny of Flowering Plants’ (1969: 
360, f. 310 A, B) maintains his view that 
the ovary of Scyphostegia represents a 
disk concealing free carpels, completely 
neglecting the detailed morphological 
work by VAN HEEL which I had personal- 
ly brought to his attention in 1967. A 
remarkable case of prejudice leading to 
quixotic stubbornness which as far as I 
know finds no parallel in botanical 
science. 


Simaroubaceae (NOOTEBOOM) 


Line 5 from top, replace ‘Samadera’ by: 
Quassia. 

Line 6 from top, after ‘Irvingia’ add: 
Allantospermum. 

Line 2 from bottom, replace ‘AUBL’ by: 
L. 

Line 6 from bottom, after ‘Klainedoxa 
(erron. Klaineodoxa)y add: Allantosper- 
mum. 

Add to the phytochemistry: 

H. P. Nootesoom (Blumea 14, 1966, 
309-315) made an additional study on 
phenolic compounds in the family of 13 
Malesian species of 9 genera. This con- 


September 1972] 


6: 196 


6: 196 
6: 198 


6: 201b 
6: 2065 
6: 218a 
6: 220a 


firms the suggestions that Simaroubaceae 
are fairly closely related to Rutaceae. 

The genus /rvingia seems to fit well 
with Simaroubaceae. 

But Suriana deviates distinctly from all 
others which would sustain the findings of 
GUTZWILLER (Bot. Jahrb. 81, 1961, 1-49) 
and others who regard this as a separate 
family. 

Change in the Key to the genera: 

1. Leaves simple. 

2. Leaf-buds enclosed by caducous 
intra-petiolar stipules. Carpels con- 
nate. Stamens 10, without an adaxial 
scale. Plant not bitter. 

2a. Stipules leaving conspicuous an- 

nular scars. Ovary 2-celled. Fruit 
drupaceous . ... 9. Irvingia 

2a. Stipular scars obscure. Ovary 5- 

celled. Fruit a capsule. 
10. Allantospermum 

2. Stipules absent. Carpels free or con- 
nate. Stamens 5, or 10 and then with 
an adaxial scale. 

Be SEO HE 
In Key fork 4 second lead line 2, replace 
‘without’ by: with. 

Quassia L. 
Add to references: NOOTEBOOM, Blumea 

11 (15 Dec. 1962) 509-528, gave a new 

subdivision of Quassia. 

In caption fig. 4 replace ‘Q. harmandiana’ 
by: Eurycoma harmandiana. 

Eurycoma harmandiana PIERRE. 

Add: Distr. Fig. 4. 

Lines 10 & 11 from top, replace “(K. & 
V.) Koorb.” by: (non K. & V.) Koorb. 
Ailanthus fordii NOOTEBOOM, FI. Mal. I, 6 
(1962) 220. — Fig. 15. 

In my world revision of the genus 
Ailanthus in Fl. Mal. (/.c. 215-220, f. 
17-18) I described a new species, known 
only from one old collection from 
Hong Kong made by Forpb, a. 1884- 
1886, and named this A. fordii, represent- 
ed in the Kew Herbarium and that of the 
British Museum (Nat. Hist.), London. 
Forp collected this in SE. Hong Kong I. 
‘near Cape d’Aguilar, as a small but 
conspicuous emergent tree from the 
shrubberies, the bare trunk being sur- 
mounted with foliage like a palm, the 
leaves being c. 40 cm long.’ 

Thanks to the kind intermediary of 
Mr. D. R. W. ALEXANDER, Director of 
Urban Services, where the Hong Kong 
Herbarium is housed, the Forester of the 
Urban Services Department, Mr. Lau 
YUNG-SuM, who acts as a collector of 
plants and seeds, has found that in the 
original locality which is still fully intact, 
viz on the Cape d’Aguilar headland, 10 
trees could be located. In Nov. 1968 
only one tree was in flower and that 
proved to be male. But its identification 


Addenda, corrigenda et emendanda 969 


Fig. 15. Ailanthus fordii NOOTEBOOM. a. Fruit, 
b-d. germlings, the pericarp sometimes remaining 
at the base of the hypocotyl, sometimes elevated 
with the cotyledons, plumule central and germina- 
tion distinctly epigeal, seed germinated imme- 
diately after receipt, b—d after 2 weeks, e. germling 
with first leaves, 2 months later; all in greenhouse 
at Leyden. All x 3/7. 


is fully certain, as the species differs from 
all others in both flowers and in fruit 
structure and could be easily checked on 
a duplicate forwarded to me and de- 
posited in the Rijksherbarium. 

Mr. ALEXANDER added that ‘the 
flowering tree is c. 30 feet tall, growing 
between granite rocks in a steep ravine 
where it faces the westerly afternoon sun 
but is sheltered from the strong north- 
east wind. All ten trees have the same 
characteristics of a grey trunk, with 
surprisingly large bole at the _ base, 
tapering sharply upwards. Most of the 
trees are forked about two feet from the 
ground and continue upwards with two 
main stems. The wood appears brittle; 
the lower trunk carries numerous scars 
and callouses where the wood has grown 
over the sites of the broken-off branches. 
It appears likely that the tree is slow- 
growing. It is possible that more trees 
may be found on this headland which is 
about 2 km long and 1 km broad, rising 
at its highest point to 325 m. There are 
numerous ravines with big granite boul- 
ders. The undergrowth in parts is very 
thick and it is by no means easy to tra- 
verse the headland looking for Ailan- 
thus. 

In examining the Hong Kong Herba- 
rium I found 4 Ailanthus sheets of A. 
fordii, all wrongly assigned to A. mala- 
barica. 


FLORA MALESIANA 


[ser. I, vol. 66 


It is interesting to enumerate the 
specimens now known, and their locali- 
ties, herbarium specimens as well as 
living trees: 

Hong Kong: Cape d’ Aguilar, 1884-1886 
Cu. Forpb s.n. (K, BM); ditto, Y.S. Lau 
(Mr. LAU YUNG-SuM) 1438, Nov. 1968, 
tree 30 ft, in rocky ravine. Mt Gough, a 
spur of the Peak area, south, leg. W.J.T. 
(UTCHER), 24 Oct. 1905, Gard. Dept. 
Hong Kong Herb. No. 666, in flower; 
ditto, No. 667 leg. Mr. Lo (Quai), 16 Jan. 
in fruit; beside the Peak Tram track near 
its upper terminus, c. 400 m, a well-grown 
tree c. 10 m high (ALEXANDER, in /itt.). 

New Territories: Sha Tin, leg.? 24 
Jan. 1905, Bot. & For. Dept. No. 4715, 
in fruit; in Jan._March 1969 16 living 
trees located on hillsides (ALEXANDER, 
in litt.). 

Cultivated: In N.B.G. (New Botanic 
Gardens) 93, leg. H. C. TANG, Gard. 
Dept. No. 12830, in flower, tall decidu- 
ous tree, bark grey, smooth, 16 March 
1949. Mr. ALEXANDER reported that this 
cultivated tree is still alive. It is 12 m 
high, with a single trunk bare of branches 
for its first 5 m. It shows no thickened 
stem-base. It flowered Nov. 1968 and is 
male. 

Notes. It is of course a pleasure to 
observe that this unique endemic of 
Hong Kong is still present in native 
vegetation and at least in its /Jocus 
classicus in such rough terrain that it will 
probably remain there for a long time to 
come. A Nature Reserve is, however, 
badly needed for it and we hope that the 
Hong Kong Government will be sym- 
pathetic towards this. Because the other 
places are partially residential areas, no 
tree could be found any more on Mt 
Gough and only one at another place on 
the Peak. Also at Sha Tin there is con- 
siderable development and though still 
17 trees could be discovered this coastal 
area, adjacent to the big inlet in the east 
central part of the N. T., is a threatened 
area for native trees. 

Early March 1969 Mr. ALEXANDER 
sent a parcel of fruits to the Rijksher- 
barium and these were distributed to 
various botanical gardens. Germination 
was at Leyden perfect and provided the 
material for the figure here reproduced. 
Fruiting is obviously in Jan.-Febr. 

The fruits collected from the single 
tree in the Hong Kong Botanic Garden, 
which can therefore not be male, as the 
herbarium suggested. Hitherto the genus 
was said to be dioecious. A local study 
of the flower morphology and biology 
seems desirable to solve this discrepancy. 


6: 220b Add at base: 


Doubtful 


Ailanthus esquirolii LEVEILLE, Fl. Kouy- 
Tcheou (1914-15) 404, nomen; Le Monde 
des Plantes 17 (1915) 23, descr. — The 
diagnosis is too short for identification; 
the specimen could not be located and 
probably does not belong to Ailanthus 
according to REHDeER (J. Arn. Arb. 14, 
1933, 227). 

Add above Excluded: 


10. ALLANTOS PERMUM 


FORMAN, Kew Bull. 19 (1965) 517, t. 1; 
WEBERLING & LEENHOUTS, Abh. Akad. 
Wiss. Lit. Mainz, M.-N. KI. 2. 10 (1965) 
544; Nootesoom, Adansonia 7 (1967) 
161-168. — Cleistanthopsis CAPURON, 
Adansonia 5 (1965) 213, t. 1. — Fig. 16. 

Trees, glabrous. Leaves simple, pen- 
ninerved, entire; beneath the upper 
epidermis a hypodermal layer which 
contains mucilage. Stipules intra-petiolar, 
soon caducous. Inflorescence a panicle. 
Flowers bisexual. Sepals 5, imbricate, 
connate towards the base. Petals 5, free, 
imbricate. Stamens 10, free, sigmoid- 
folded in bud. Disk intrastaminal, 10- 
lobed. Ovary 5-celled with 1 pendent 
anatropous-epitropous ovule centrally 
attached near the apex of each cell; 
style terminal, sigmoid-folded in bud. 
Fruit a_ septicidal and incompletely 
loculicidal capsule, the valve-halves 
becoming slightly twisted after dehiscen- 
ce, detaching, leaving a central columella; 
the (often aborted) seeds are attached to 
an enlarged placenta towards the apex of 
the columella. In A. multicaule the young 
fruit is + drupaceous. Seeds shining and 
waxy, cotyledons planoconvex, radicle 
pointing upwards; plumule very small. 

Distr. 2 spp., 1 in Malesia (Malaya 
and Borneo) and | in Madagascar (A. 
multicaule (CAPURON) Noor., /.c. 163). 

Notes. This genus has a remarkable 
distribution and an equally interesting 
taxonomy. It was only recently in- 
dependently described by CAPURON who 
arranged it in /rvingioideae and by For- 
MAN who removed I/rvingioideae from 
Simaroubaceae and Ixonanthoideae from 
Linaceae, joining them as distinct sub- 
families into a new family /xonanthaceae. 
Allantospermum he arranged in subfamily 
Ixonanthoideae. 

Independently I came to the same 
conclusion as CAPURON, viz that A/lan- 
tospermum belongs to Simaroubaceae- 
Irvingioideae. The morphological char- 
acters are in favour of this and this 
disposition is also sustained on chemo- 
taxonomical grounds; see NOOTEBOOM, 
Adansonia 7 (1967) 161-168. 


September 1972] Addenda, corrigenda et emendanda 971 


Fig. 16. Allantospermum borneense FORMAN. a. Habit, x 2/3, b. flower, 4, c. ditto, floral parts removed 

except ovary, <8, d. ditto, lengthwise section, e. fruit, x 2/3, f-e, fruit valves, x 2/3, h. seed, » 2/3, 7. 

columella, (7. placental outgrowth, 2. sterile ovule), + x2. — ssp. rostratum NOoTEBOOM, j. Columella, 
x2, k. young fruit, x 2/3 (a-d S 3364, e-h S 15166, i S 15162, 7 SAN 36068, k SAN 36015). 


The controversy about the systematical 
affinity of A//antospermum has also been 
tried to solve by means of anatomical and 
palynological comparison. 

Wood Anatomy. Rojo (Adansonia 8, 
1968, 73-83) examined its wood anatomy 
and found this distinctly different from 
that in Jrvingioideae and possibly more 
approaching, but still different from 
Ixonanthes and Ochthocosmus (Ixonan- 


thaceae sens. str.). Wood anatomically 
Irvingioideae constitute a distinct group 
within Simaroubaceae. For its wood anat- 
omy Allantospermum could be another 
group of the same rank as the other 
distinct wood anatomical types /rvingioi- 
deae, Kirkioideae, Picramnioideae and 
Alvaradoideae (cf. WEBBER, Am. J. 
Bot. 23, 1936, 577-587). 

Leaf Anatomy. METCALFE, LESCOT & 


FLORA MALESIANA 


[ser. I, vol. 6® 


LosBreEAU (Adansonia 8, 1968, 337-351) 
reported the differences between Al/lanto- 
spermum and Irvingioideae, mainly being 
the absence of mucilage cavities in the 
former and their presence in the latter 
and other Simaroubaceae. Remarkable, 
however, is the common presence of 
cristarque cells in both Allantospermum, 
Klainedoxa and Irvingia. The absence of 
mucilage canals is a negative character 
shared with [xonanthaceae sens. str. 

Palynology. In her comparison with 
Irvingia, LOBREAU, /.c., stresses differ- 
ences more than the undoubted similarities 
and is apparently unaware that the 
general Simaroubaceae pollen type is 
close to that of A/lantospermum. 

In comparing with /xonanthaceae she 
does the opposite and stresses similarities 
more than the differences and strangely 
enough does not comment on the striking 
difference in exine structure between 
Allantospermum and Ixonanthes. Her 
conclusions rest therefore only on a 
comparison with Ochthocosmus. Here she 
states that in both genera the polar 
triangle is small, but omits to mention 
that this is only true for A. multicaule. In 
A. borneense the polar triangle is large. 
The endexinous thickenings which form 
the other support for her thesis are in all 
types concerned weakly developed and 
should not carry weight. 

LopreAu’s idea that Allantospermum 
pollen is more similar to /xonanthaceae 
pollen than to Irvingiaceae would there- 
fore appear to be weakly founded. The 
striking similarity of Allantospermum 
pollen to other Simaroubaceous pollen 
and the undoubted resemblance to 
Irvingia pollen support, in my opinion, 
NOoTEBOOM’s idea to include A//anto- 
spermum in Simaroubaceae subfam. Ir- 
vingioideae. — J. MULLER. 

Concluding, the evidence is first that 
the genus stands apart in various aspects. 
Second that the morphological, chemo- 
taxonomical and palynological characters 
all point to inclusion with Irvingioideae 
in Simaroubaceae, that the balance in leaf 
anatomy is 50: 50, and that the wood 
anatomy would be in favour of inclusion 
in [xonanthaceae sens. str. 


1. Allantospermum borneense FORMAN, 
Kew Bull. 19 (1965) 517, t. 1; NooTe- 
BooM, Adansonia 7 (1967) 162, pl. 1 f. 
a—b. — Fig. 16 a-i. 

Tree, up to 30m by 50cm @ ; buttresses 
short to steep, up to 75 cm high, some- 
times with warts. Stipules only present in 
leaf-buds, narrowly elliptic-oblong, point- 
ed, c. 1 cm (ForMAN, /.c.). Leaves glossy 
above, dull beneath, elliptic with cuneate 
to broadly rounded base and acuminate, 


blunt apex, 7-16 by 3-8 cm; nerves 5-10 
pairs, arching upwards and meeting in a 
not very conspicuous intramarginal vein; 
veins + transverse, netted; midrib and 
nerves prominent, veins slightly promi- 
nent on both surfaces; petiole dark-col- 
oured, grooved above, c. 1 cm. Panicles 
lax, up to 7 cm. Bracts early caducous, 
leaving an obvious scar. Pedicels 7-9 mm. 
Sepals boat-shaped, reflexed at anthesis, 
3-4 by 2-3 mm, caducous when the fruit 
matures. Petals elliptic to obovate, 4-5 
by 21/2-3 mm, membranous, veined, 
reflexed at anthesis, caducous. Stamens 
up to 6 mm long, with versatile, c. 1 mm 
long latrorse-introrse anthers. Disk c. 
11/2 mm @ and !/2 mm thick. Ovary 
5-lobed, c. 11/2 by 2 mm;; style filiform, 
c. 3{-4) mm; stigma capitellate. Capsule 
broadly ellipsoidal, 5-lobed, 21/2-5 cm 
long, abruptly (up to 3 mm) acuminate 
at apex. Seeds cylindrical, often slightly 
curved, 2—2!/5 by 4-6 mm. 

Distr. Malesia: Malaya (Penang, 
Perak, Trengganu), Borneo (Sarawak, 
Brunei). 

Ecol. Obviously not rare in primary 
lowland rain-forest. 

Notes. In my paper (/.c. 164) I wrote: 
‘I never saw a lobed structure as depicted 
in Forman’s paper’. In fact, the placental 
outgrowths are somewhat lobed, and the 
lowermost part sometimes tears from the 
columella (probably due to desiccation in 
the herbarium). In some cases the seeds 
leave some material when tearing off 
from the placenta. 

A. multicaule from Madagascar differs 
from A. borneense in the following char- 
acters: Tree with many trunks. Stipules 
narrowly lanceolate, acute, 2—2!/2 mm 
long. Leaves (narrowly) ovate, with 
rounded base and long-acuminate, acute 
apex, 31/2-10 by 11/2-31/2 cm; nerves 
3-5 pairs. Fruit c. 2 by 1.7—2 cm. Seeds 
10-14 mm long, possessing a_horse- 
shoe-shaped arillodium directly above the 
hilum. 


ssp. rostratum NOOTEBOOM, ssp. nov. — 
Fig. 16 j-k. 

Inflorescentia praesertim basi race- 
mosae_ plusminus condensae. Capsula 
apice longiter acuminato. — Typus: SAN 
36068 (L). 

Distr. Malesia: Sabah (Sandakan, 
Lahad Datu and P. Sakar, SAN A 4162, 
17478, 26030, 36015, 36068, 37479, 
37931, 38777). 

Note. Different from the type sub- 
species in having a shorter, more con- 
densed_ inflorescence predominantly 
branched towards the base and up to 
2 cm long; acumen on the fruit 6-8 mm 
long. 


September 1972] 


Addenda, corrigenda et emendanda 973 


4: 281 


4: 


282 


Sonneratiaceae 


Sonneratia L. f. 

It has appeared that S. caseolaris (L.) 
ENGL. can grow in fully fresh water 
without any connection with brackish 
water. It has been found along the shore 
of the Sentani Lake, at 75 m altitude, on 
sandy clay or peaty soil, and also along 
the Obehfareh R., near Hollandia, in the 
North of West New Guinea. Other plants 
occurring along the lake-shore are Crinum 
asiaticum L. and Pongamia pinnata (L.) 
Merr. which are otherwise largely con- 
fined to the sea-shore. This lake contains 
also several marine animals. It is accepted 
that it was, even in historical time, a 
lagoon in open connection with the sea 
which has become an inland lake by the 
rapid upheaval to which the northcoast 
of New Guinea is subjected. It is most 
curious that the species has been able to 
maintain itself. It produces the normal 
aerophores. 

Shortly afterwards we received material 
of the same species from East New 
Guinea, again from the northcoast, 
where Mr. J. S. WOMERSLEY collected it at 
150 m altitude, in Morobe Distr., 10 
miles from Lae, as a 9 m tall tree along 
the inner margin of a sago swamp, near 
Oomsis on Wau road (NGF 15307), Aug. 
15, 1962. 

Though it can be cultivated in fresh 
water, e.g. at Bogor Botanic Gardens, 
at 250 m alt., this is the first time it was 
found in fresh water under purely natural 
conditions. Obviously, if conditions are 
favourable and upheaval rapid, marine 
plants may maintain themselves for some 
time. The trees in question are not thick 
enough to have survived since the be- 
ginning of the upheaval which started 
some 4-5 centuries ago and must date 
from later generations. Also at Bogor 
regeneration takes place under fresh- 
water conditions. Cf. STEEN. Nova 
Guinea, Bot. n. 12 (1963) 189. 

A distribution map of the genus is 
published in Pac. Pl. Areas 2 (1966) 
248-249, 

The species of Sonneratia can mostly 
be distinguished also in the sterile state, 
except for S. alba and S. caseolaris. 
Mr. J. MULLER has found a most easy 
differentiating character in the leaf tip 
which bears a thickened pitted gland 
(? hydathode) peculiar to S. caseolaris; 
in S. alba instead the leaf margin at and 
near the ending of the midrib is thinner 
and finely recurved without such thicken- 
ed tip. This character is possibly not 
absolute, but in any case most helpful. 
Bic lize 
Add the following new paragraphs: 


Anatomy. It is to be regretted that Dr. 
C. A. STACE in his work on epidermal 
characters on mangroves (New Phyto- 
logist 66, 1966, 304-318) omitted to 
examine Sonneratia. Dr. DinG Hou 
found that the stomata of Sonneratia 


® : a 


Fig. 17. Leaf tips of Sonneratia. — a—d. S. caseolaris 
(L.) ENGL., <3. — e-g. S. alba J.SM., <3. The 
difference is not completely exclusive and some 
intermediates occur (a Koorpers 4442, b6 NGF 
5026, c BW 3198, d Koorpers 14219, e KARSTEN 
s.n., f COERT 1456, g BORSSUM WAALKES 272). 


974 FLORA MALESIANA [ser. I, vol. 6 


Fig. 18. A. Sonneratia ovata, B. S. alba * ovata, C. S. alba, D. S. alba x caseolaris, E. S. caseolaris, 
all x 1/2 (A MULLER 5, Labuan, B MULLER 8, Brunei, C MULLER s.n., 28-12-63, Labuan, D MULLER 11, 
Brunei, E MULLER s.n., 30—5—64, Brunei). 


agree in structure very well with those of 
Rhizophoraceae. 

He found, however, that they differ in 
the species, those of 8. alba and SS. ovata 
being much larger than those in S. 
caseolaris. Moreover, the two former 
species have a peculiar type of glands (?) 
in the leaves, of the size of stomata and 
equalling the latter in number. They are 
surrounded by a fairly large number of 
radially arranged cells. In section there 
appears to be a large sac-like hollow 
(cell or intercellular space ?) below them. 
These glands or whatever they may be 
are not found in S. caseolaris. 

Hybridisation & Chromosomes. 
J. Mutter & Mrs. S. Y. Hovu-Liu 
(Blumea 14, 1966, 337-343) have together 
worked on the identity, the pollen and the 
chromosomes of pickled material of 
Sonneratia collected by MULLER and by 
Mr. J. P. VAN NIEL, in Seria, Brunei. This 
was initiated through Mr. MULLER who 
in his pollen studies had found discrep- 
ancies in the pollen when in Brunei. 

It has been shown that in material 
collected by MULLER S. alba may form 
hybrids with S. caseolaris and with S. 
ovata, the characters of which have been 
tabulated by Mutter (/.c. tab. 1) as to 
morphological characters, amongst them 
the fruits (fig. 18). This is the first time 
that the occurrence of hybrids in man- 
grove trees was reported (it was recently 
suggested by BretTeLreR in Acta Bot. 
Neerl. 18, 1969, 434-444, that Rhizo- 


Phora harrisonii is also a hybrid). 

Mr. MULLER says that the hybridisa- 
tion is due to the rather unusual situation 
that the the three parent species grow side 
by side in the Brunei River estuary; 
usually they are ecologically separated, 
S. alba being the more marine, S. ovata 
the least marine, while S. caseolaris is 
ecologically intermediate. The very nar- 
row mangrove belt along the rather steep 
banks of the Brunei R. has telescoped the 
various ecological zones and increased 
chances of contact. Another factor pos- 
sibly promoting hybridisation may have 
been the scarcity of S. ovata which thus 
stands a large chance of being cross- 
fertilized with S. alba pollen. This is also 
suggested by the obvious concentration 
of alba * ovata hybrids around the 
single observed locality of S. ovata. 

The reduced fertility of the hybrids 
indicates that the three are good species, 
but the sterility barrier between S. ovata 
and S. alba appears lower as compared 
with that between S. alba and S. caseo- 
laris. 

Mrs. Hou-Liu recorded her experience 
with the chromosomes which appeared 
very difficult to count; all species have 
i1— ile 

MuLLteR & VAN STEENIS (North 


* Queensl. Natur. 35, 1968, n. 147, 6-8, 1 


map) revised the genus Sonneratia for 
Australia and mapped its localities in 
Northern Australia and Queensland 
where only S. alba and S. caseolaris 


September 1972] Addenda, corrigenda et emendanda 975 


occur, but also indubitable hybrids be- 
tweenthem,as proved by fruit anomalies 
and pollen sterility. 

Palynology. J. MULLER (Pollen et 
Spores 11, 1969, 223-298, 15 fig., 15 pl.) 
has made an extensive examination of 
pollen of Sonneratia and compared it 
with that of Duabanga, and various 
Lythraceae. In S. alba and S. caseolaris 
he found a distinct intraspecific varia- 
bility whichis geographical and which is 
assumed to be genotypic. These were 
charted. A key was given to the 5 species 
on pollen characters. 

Fossil pollen. J. MULLER (in Cran- 
well, ed., Ancient Pacific Floras, Univ. 
Hawaii Press, 1964, 33-42, 2 fig., 1 pl.) 
studied fossil pollen of Borneo. In NW. 
Borneo pollen of the S. caseolaris type 
is oldest and found onwards the transition 
between Oligocene and Miocene; the S. 
alba type is younger, from the start of the 
Pliocene. This is not reflected in the pre- 
sent range, S. alba is the wider distributed 
species. S. ovata was not charted; this 


Fig. 19. Fruit and stamen details of Duabanga. 
a-a’. D. grandiflora (RoxB. ex DC.) WALP., b-b’. 
D. taylorii JAYAWEERA, c-c’. D. moluccana BL. 
Fruits 1/2, stamens x2!/2 (after GEESINK). 


4: 283a 


4: 288 


is said to be closest related to S. alba. 

MULLER refers also to papers of Indian 
palaeobotanists who have recorded 
Pliocene fossil wood of Sonneratia-like 
structure and a flower and fruit from the 
Eocene. 

A tentative picture of the evolution of 
Sonneratia pollen was presented by 
GERMERAAD, HOPPING & MULLER (Rev. 
Palaeobot. Polynol. 6, 1968, 189-348) in 
which is discussed a Lythraceous Tertiary 
pollen type, Florschuetzia trilobata, which 
is presumed to be ancestral to Sonneratia, 
and pointed to the remarkable resem- 
blance of the latter to the pollen of the 
Lythraceous genus Lagerstroemia. 

Affinity. Pollen types closely com- 
parable to those of Sonneratia are found 
in Duabanga and furthermore in various 
Lythraceae (MULLER, /.c. 1969, 291-292), 
notably the New World subtribe Dipluso- 
dontinae (Diplusodon, Lafoénsia) and the 
Old World subtribe Lagerstroemiineae). 
Palynologically there would be no objec- 
tion to merge Sonneratiaceae with 
Lythraceae. 

Sonneratia acida L. f. var. mucronata MiqQ. 
was reduced here to S. caseolaris (L.) 
ENGL. MIQUEL’s variety was based on two 
sheets, one of HORSFIELD and one of 
REINWARDT. The first is in U and appears 
to belong to S. alba J. Situ. I could not 
find at U or L the REINWARDT sheet 
ticketed by MIQUEL. 

Duabanga Bucu.-Ham. — Fig. 19. 

Add under the genus the following para- 
graphs: 

Blastogeny. Seedlings have been de- 
scribed and depicted by Troup (Silv. Ind. 
Trees 2, 1921, f. 228) and JAYAWEERA & 


Howard (Baileya 10, 1962, 8-13, 2 fig.). 

Hybridisation & Chromosomes. 
JAYAWEERA (J. Arn. Arb. 48, 1967, 89- 
100, 8 fig., 1 tab.) has found that four 
large trees growing in the Botanic Garden 
at Peradeniya and raised from seed 
received from the Botanic Gardens, 
Bogor about 1853, are distinctly different 
from the two known species. He described 
this as a new species, D. taylorii JAYA- 
WEERA. Fig. 19 b—b’. 

It struck us that in a number of char- 
acters the new species is intermediate 
between the continental SE. Asian spe- 
cies D. grandiflora and the Malesian D. 
moluccana, which have replacing ranges. 
At our suggestion Mr. GEESINK has 
studied the new species and satisfactorily 
shown that it must be a primary hybrid 
between the two known species. See his 
study in Blumea 18 (1970) 453-456, | fig. 

Naturally this hybrid could not have 
been found in the wild, because of the 
replacing ranges of the parents. However, 
in the Botanic Gardens at Bogor, where 
cultivated species are grown in systematic 
arrangement, the two species grow side 
by side in the same compartment parcel 
VII. D, so that hybridisation can easily 
occur. See TEYSMANN & BINNENDIJK, Cat. 
Hort. Bog. (1866) 241 and Dakkus, Cat. 
Bot. Gard. (1927) 119. 

A further communication from Dr. 
JAYAWEERA iS most interesting, viz that 
the seed of D. x taylorii is for 95% fer- 
tile, from which it should consequently be 
deduced that D. grandiflora and D. 
moluccana are rather subspecies (replac- 


976 


FLORA MALESIANA 


[ser. I, vol. 66 


ing races) of one species only than species 
in their own right. 

A further study of chromosomes seems 
desirable; this has only been observed for 
D. grandiflora by J. L. THOMAS (Baileya 
10, 1962, 13), n=24. 


Sphenocleaceae, see Campanulaceae 


4: 530b 


4: 53la 


4: 532a, 
5: 564a 


42 352, 
353 


Stylidiaceae 


Stylidium inconspicuum SLOOT. 

Add to literature: ERICKSON in Specht, 
Rec. Am.-Austr. Exp. Arnhem Land 3 
(1958) 312; Triggerplants (1958) 197-198. 
Add to Distr.: North Australia, Arnhem 
Land (Groote Eylandt: Hemple Bay). 
Note. Mrs. ERICKSON refers this with 
some doubt to the species which was 
first described from Java. In our expe- 
rience Malesian representatives of genera 
which are characteristic of Australia, 
especially herbaceous ones, are almost 
never endemic in Malesia but almost 
always also present in Australia (as for 
example in Goodeniaceae, Amaranthaceae, 
etc.); the existence in Australia of this 
hitherto endemic species of Java would 
be no great surprise. 

Stylidium javanicum SLoot. 

Add to literature: ERICKSON, Trigger- 
plants (1958) 196. 

Note. At some time it was assumed that 
also this Malesian endemic (Indramaju; 
Sumba) occurred in NW. Australia, but 
on closer examination Mrs. ERICKSON 
and Mr. WiILuLis found the Australian 
specimens allied but specifically different 
and have described them as S. fluminense 
ERICKSON & WILLIS, Vict. Natur. 83 
(1966) 108, pl. 2, f. 1-6. 

Stylidium pedunculatum R.BR. 


Add to literature: ERICKSON, Trigger- 
plants (1958) 190, pl. 55, f. 5-9. 
Note. I have reduced to this S. erickso- 
nae WILLIS, Vict. Natur. 73 (1956) 43; 
ERICKSON, Triggerplants (1958) 189, pl. 
55, f. 10-17. — syn. S. androsaceum 
O. SCHWARZ in Fedde, Rep. 24(1927) 105, 
non LINDL., nec DC. 

Both Mrs. ERICKSON and Mr. WILLIS 
believe these two species to be different 


49 


50a 


Thymelaeaceae 


and distinguish them as follows: 

1. Leaves shortly ciliate on the margins 
only, terminating into a long hair-like 
point. Peduncles 2-6 cm. Small petals 
notched, corolla throat quite naked. 

S. pedunculatum 

1. Leaves bristly-hairy all over, blunt to 
acuminate but not hair-pointed. Pe- 
duncles 6-11 cm. Smaller petals entire, 
corolla throat with 4 obscure ap- 
pendavesimn sna S. ericksonae 
I must remark that the Aru specimens 

are almost glabrous and that I am not 
particularly impressed by the amount of 
hairs on the leaves. A very hairy specimen 
of the North. Terr., viz S. T. BLAKE 
16371, was by him identified as S. pe- 
dunculatum, but must then belong to 
S. ericksonae, but the smaller petals are 
not notched as far as I can observe. And 
also I am not impressed by the notching 
of the smaller petals if I see the plate; it 
may be that two taxa are involved but the 
scant material at present available does 
seem to indicate the desirability of future 
research on delimitation and status. 


Styracaceae 


Bruinsmia BOERL. & KoorD. 

Change in the description: Leaves almost 
entire to coarsely crenate. Calyx truncate 
or 5-lobed. Corolla dull greenish or white. 
Ovary 3-5(-6)-locular. Fruit oblong to 
globular. Pericarps pulpy or drupaceous. 
Seeds l-co per cell. Cf. STEEN. Bot. 
Jahrb. 86 (1967) 390. 

Change last line on Distr. as follows: 
Distr. Two species, one in Malesia, the 
other one in Burma, Assam, and Thai- 
land. 

Bruinsmia styracoides BOERL. & KoorD. 
Add to Distr.: Philippines, Mindanao 
(PNH 36146). Cf. STEEN. Philip. J. Sc. 88 
(1959) 121. 

By this discovery also this genus can be 
removed from the few which cross Ma- 
kassar Straits without intermediary 
stations either in the Philippines or Lesser 
Sunda Islands. Also Bromheadia and 
Pericopsis are now removed from this 
list, published by me in Bull. Jard. Bot. 
Btzg III, 12 (1932) 259. 


Gonystyloideae (AiRY SHAW) 


Gonystylus TEYSM. & BINN. 


Replace the Key to the species by the following: 


REVISED KEY TO THE SPECIES 


1. Leaves very long, 40-50 cm, venation bullately impressed above, very prominent below. 
Inflorescence robust, with a very thick rachis, up to 5mm @. Flowers large, with c. 40 disk- 
lobes and 80 stamens. Style robust, with large stigma, and 4—7 small clavate ‘parastyles’ around 


September 1972] Addenda, » corrigenda et emendanda 977 


the base. . . 2. G. areolatus 

1. Leaves shorter, "3-40(-43) ¢ cm, ‘venation rarely bullately impressed (cf. 26. G. nervosus). Inflo- 
rescence much less robust. Flowers smaller, disk-lobes and stamens 10-40. Style slender; 
stigma small; ‘parastyles’ absent (rarely present: cf. 6. G. xylocarpus and 18. G. micranthus), 
but sometimes represented by 3-4 small rounded humps. 

2. Inflorescence-branches elongating almost indefinitely, forming long crowded cicatricose 
racemes with a few flowers at the apex at the time of flowering. Leaves 10-29 by 5-14 cm. 
Disk-lobes (where known) 15-16, densely retrorse-setulose. Capsule + lanceolate, 3- or 
6-ribbed (Sect. Auxanthus Airy SHAW). 

3. Leaves up to 29 cm long, with a dull, ‘shagreened’ surface, the lower surface always bearing 
minute adpressed hairs, lying parallel to the nerves, often difficult to see (W. Indonesian 


Borneo; SW. Sarawak) . . . . 1. G. augescens 
3. Leaves up to 20 cm long, with a very smooth ‘and somewhat shining surface, the lower side 
entirely devoid of minute hairs (NE. Sarawak; Brunei) . . . . . 25. G.lucidulus 


2. Inflorescence-branches not elongating, bearing fascicles or short irregular racemes of 
flowers on short nodulose side-branches. Disk-lobes glabrous or occasionally setulose. 
Capsule + globose, not or sometimes weakly ribbed. 

4. Leaves gradually narrowed at the apex into a relatively long slender acumen, cuneate at 
the base, up to 20 by 51/2 cm, almost glabrous, drying chestnut-brown. Inflorescence 
ferrugineous-tomentellous, branches shortly racemiform. Flowers truncate at the base. 

11. G. acuminatus 

4. Leaves not gradually narrowed into a long acumen, usually rather suddenly narrowed into 

a relatively short acumen or cusp, sometimes rounded or even retuse. 
5. Pedicels 2-3 cm. Leaves usually large (up to 40 by 15 cm). 

6. Midrib distinctly raised above; leaves up to 43 by 12cm, drying olivaceous-brown, almost 
glabrous; petiole 11/2-2 cm. Inflorescence rather robust, to 17 cm; disk-lobes 30-35, 
glabrous) “29 207): pcan Oe Hol. er i) 2G. See 22s Gyeostalis 

6. Midrib impressed above. 

7. Petioles usually or sometimes exceeding 2 cm. 
8. Leaves, especially midrib and nerves, ochraceous-tomentellous beneath; petioles and 
inflorescence-rachis strongly angled, tawny-tomentellous; disk-lobes 40-45. 
28. G. spectabilis 
8. Leaves glabrous beneath; petioles and inflorescence-axis terete or less strongly angled, 
cinereous-pubescent; disk-lobes 20-40. 
9. Petiole up to 21/2 cm; leaves drying some shade of ochraceous brown. Inflorescence 


elongate, to 22 cm : . . 5. G.macrophyllus 
9. Petiole 3-4 cm, 4-5 mm @; ': leaves drying pale greenish above, with a narrow purple 
margin. Inflorescence to 11 cm = RUB Ae NEF ee... 27. Ganopilis 


7. Petioles not exceeding 2 cm. 
10. Leaves glabrous or almost so. 
11. Leaves drying a dark purplish-leaden colour; base of calyx narrowing gradually 


into pedicel; disk-lobes 20-30 . . . b pasher oer ure calophyllus 
11. Leaves drying a light brownish colour; base of calyx broadly truncate, passing 
abruptly into the pedicel; disk-lobesc.50 . . . . . . 20. G. calophylloides 


10. Leaves thinly pilose or pubescent beneath, at least on the midrib. 
12. Leaves drying a light green with a narrow dark brown margin; disk-lobes c. 40. 
4. G. reticulatus 
12. Leaves greyish-green above, ochraceous-brown beneath; disk-lobes c. 30. 
21. G. consanguincus 
5. Pedicels !/2-2 cm. Leaves small or medium. 

13. Calyx-segments strongly reflexed or revolute at anthesis; disk-lobes 7-12. Inflorescence 
usually considerably branched. Flowers small. Leaves small, up to 13 by 5 cm, dis- 
tinctly shagreened. 

14. Disk-lobes tomentellous throughout and setulose within. Leaves elliptic or almost 
rhomboid, glabrous or almost so, usually ochraceous when dry; nerves rather steeply 
ascending : . . 14. G. forbesii 

14. Disk-lobes glabrous. “Leaves elliptic to oblong but 1 never subrhomboid, drying brown- 
ish or greenish; nerves rather widely spreading. 

15. Young parts and inflorescence densely fulvo-velutinous. Leaves usually -- pubescent 

below, glossy above (even when dry), rather variable in shape, often cuneate at base. 

15. G. velutinus 

15. Young parts and inflorescence tomentellous. Leaves glabrous or almost so below, 
dull above when dry, regularly elliptic-oblong, mostly rounded at base. 

16. G. maingayi 

13. Calyx-segments not or scarcely reflexed at anthesis. 


978 FLORA MALESIANA [ser. I, vol. 6% 


16. Disk-lobes 10-12, very slender, pustulate toward apex. Flowers very small (4-6 mm). 
3 filiform, clavate parastyles present. Leaves drying a dull purplish-leaden colour, 
up: to5iby Gicmiy i. at . bee fos solseiGeimicranthus 

16. Disk-lobes 13-40. Flowers larger. 

17. Leaves drying a purplish-leaden colour; midrib flat or slightly raised above. 
12. G. confusus 
17. Leaves not drying purplish-leaden. 
18. Disk-lobes retrorse-setulose within. 

19. Leaves drying greyish-green above with a narrow brown border, pinkish-brown 
below, chartaceous; nervation lax, the primary nerves being distinctly differentiated 
from the secondaries: disk=lobest20=22.5) (ae tee . . 9. G. keithii 

19. Leaves drying brownish or ochraceous, or greenish without a brown border, 
+ coriaceous; venation dense, without distinctly differentiated primary nerves. 

20. Leaves drying chestnut-brown throughout; disk-lobes 25-30. 10. G. brunnescens 

20. Leaves drying ochraceous or greenish; disk-lobes 35-40 . . 23. G. decipiens 

18. Disk-lobes glabrous. 

21. Midrib distinctly prominent above. Sepals narrowly triangular-lanceolate. Leaves 
yellow-ochraceous when dry : . . . 7. G. stenosepalus 

21. Midrib flat or channelled above. Sepals ovate- deltoid. 

22. Leaves small, 4-15 by 2-7 cm, coriaceous, often + conduplicate, drying dull 
purplish-red below and chestnut above; nervation relatively inconspicuous. 
19. G. bancanus 

22. Leaves various, but not conduplicate, nor drying as above; nervation more 

conspicuous. 

23. Indumentum of inflorescence tomentose or tomentellous, 7.e. with short 
spreading hairs, usually + fulvous. 
24. Leaf-surface not dull and ‘shagreened’ (though not shining); pedicels 11/2-21/2 
cm; calyx 5-8 mm long; disk-lobes c.30 . . . . . 21. S. consanguineus 
24. Leaf-surface dull and ‘shagreened’, slightly glaucescent below; pedicels 0.8—2 
cm; calyx 5-6 mm long; disk-lobes 20-30. 

25. Leaves long and narrow, up to 27 by 7 cm, at least 4 times as long as broad, 
glabrous; inflorescence up to 28 cmlong . . . . . 24. G.glaucescens 

25. Leaves less elongate, up to 17 by 81/2 cm, only 2-3 times as long as broad, 
often tomentellous below; inflorescence up to 12 cm long. 13. G. affinis 

23. Indumentum of inflorescence thinly adpressedly grey-pubescent. 
26. Leaves large, glabrous, up to 34 by 12 cm. 

27. Leaves coriaceous, drying pale green above with a narrow brown edge (as 
in 9. G. keithii and 4. G. reticulatus); nerves not bullately impressed; petiole 
robust, terete, up to4cmlong . . ohn | 2iatGenobilis 

27. Leaves chartaceous, not drying pale green with a brown edge; primary nerves 
+ bullately impressed; petiole much less robust, up to 2 cm long. 

26. G. nervosus 
26. Leaves small or medium, 9-24 by 3-71/2 cm, glabrous or shortly adpressed- 
pubescent below. 

28. Leaves rigidly coriaceous, broadly elliptic, up to 17 by 9 cm; inflorescence 
robust, up to 18 cm; disk-lobes 35-40; parastyles 2—3; fruit massive, pericarp 
up to 2 cm thick, almost StOUYs sane . . 6. G.xylocarpus 

28. Leaves thinly to firmly chartaceous; inflorescence rather slender; disk-lobes 
25-30; parastyles absent; fruit with moderately thick and woody pericarp. 

29. Leaves 12-24 by 41/2-71/2 cm; inflorescence 10-20 cm. 8. G. borneénsis 
29. Leaves 9-11 by 3-4 cm; inflorescence 2-3 cm, few-flowered. 17. G. pendulus 


4: 35Sa Gonystylus xylocarpus Airy SHAW. parastyles 2-3, flattened-clavate, 11/2 
Add to literature: Fl. Mal. I, 4 (1953) 355; mm. Fruit dehiscing by 4 valves. 
Kew Bull. 17 (1964) 450. Add to Ecol.: Dominant tree in kerangas 
Add to description: Tree up to 36 m. on flat ridge, 30-150 m altitude. FI. 
Inflorescence robust, up to 18 cm long. March, fr. March, June, July. 
Pedicels up to 1.8 cm, grey-sericeous. Add to Vern.: Ramin batu, garu mélitan, 
Calyx 7 mm long, 7-8 mm @, thickened Sarawak. 
and truncate at the base, very shortly Replace the Note by: Noteworthy as 
adpressed-sericeous; segments + deltoid, one of the very few species of the genus in 
somewhat obtuse and very shortly re- which ‘parastyles’ are developed. 
curved at the apex. Disk-lobes 35-40, 4: 359a Gonystylus maingayi Hook. f. 
narrowly subulate, 3-4 mm long, glab- Add to literature: Airy SHAW, FI. Mal. 


rous, epustulate. Style pilose below; I, 4 (1953) 359; Kew Bull. 17 (1964) 456. 


September 1972] 


4: 359b 


4: 36la 


4: 361b 


Add to Distr.: Borneo (Sarawak, Brunei, 
Sabah). 

Add to Ecol.: Common dominant tree in 
primary peat-swamps, up to 15 m. F7. 
Febr., Oct.-Nov., fr. Febr., April. 

Add: Uses. Wood used for planks and 
boards for domestic buildings (Sabah). 
Add to Vern.: Ramin batu, Sarawak, 
bidaru, Sabah (from Brunei informant). 
Add to Notes: This species apparently 
occupies a similar peat-swamp habitat to 
19. G. bancanus, but is evidently far more 
local. The fruit develops into a curiously 
asymmetrical, bean-like form, with one 
of the three valves almost abortive. 
Gonystylus micranthus AiRY SHAW. 

Add to literature: Fl. Mal. I, 4 (1953) 
361; Kew Bull. 17 (1964) 457. 

Add to description: Tree to 15-20 m. 
Leaves with drip-tip up to 2!/2 cm. 
Calyx ovoid, up to 6 mm. Disk-lobes 
10-12, very slender, pustulate towards 
apex, glabrous. Style surrounded at the 
base by 3 conspicuous, filiform, clavate- 
capitate parastyles, 11/2 mm long. 

Add to Distr.: S. Indonesian Borneo. 
Add to Ecol.: Primary lowland forest or 
marshy forest, on sandy loam soil, 20-90 
m. Fl. Febr., March, July, Aug. 

Add to Notes: The fruit of this very 
distinct species is still a desideratum. 
After 19. Gonystylus bancanus (MiIQ.) 
Kurz add the following species: 


20. Gonystylus calophylloides Airy SHAW, 
Kew Bull. 17 (1964) 448. 

Small tree to 6 m. Leaves oblong or 
slightly oblanceolate-oblong, 30-37 by 
11-12 cm, slightly cordate at base, round- 
ed and abruptly shortly caudate at apex 
(cauda 1.2-1.8 cm by 2-4 mm), chartaceo- 
coriaceous, glabrous, pale ochraceous- 
brownish (greenish tinged above) when 
dry; midrib moderately robust, very 
prominent and subcylindric beneath, flat 
or scarcely prominulous above; nerves 
very slender, c. 35 pairs, distinctly 
prominulous on both surfaces; petiole 
1-11/2 by 4 mm, very rugose. Jnflores- 
cence very abbreviated, 21/2 cm, few- 
flowered, sericeous. Pedicels 1.2—2.5 cm, 
densely ochraceous-sericeous. Calyx 
broadly ovoid, truncate at base, 7-8 cm 
@: sepals broadly deltoid, 7-8 by 4-6 
mm, slightly recurved and sometimes 
with a slight thickening at the apex, 
dorsally ochraceous-sericeous. Disk-lobes 
c. 50, robust, laterally flattened, 6 mm 
long, glabrous, epustulate. Style elongate, 
long-hairy, surrounded at the base by 4-5 
small obtuse ‘parastyles’. Fruit subglob- 
ose, 31/2 cm @%, apparently 5-valved, 
rugulose. obscurely puberulous, subtend- 
ed by the 10 mm long sepals and borne 
on a striate pedicel 2!/2 cm by 3-4 mm. 


Addenda, corrigenda et emendanda 979 


Seed apparently solitary, large. 

Distr. Malesia: Borneo (NE. Sara- 
wak). 

Ecol. Banks of rocky stream at 210 m. 
Fi. fr. July. 

Vern. Ramin, Sarawak. 

Note. Closely related to 3. G. calophyl- 
lus of SW. Sarawak, differing in the pale 
ochraceous-brown colour on drying, and 
in the broadly ovoid shape of the calyx, 
the truncate base of which passes abrupt- 
ly into the pedicel. 


21. Gonystylus consanguineus Airy SHAW, 
Kew Bull. 17 (1964) 454. 

Tree, 16-40 m. Leaves elliptic to 
oblong, 10-25 by 5-9 cm, broadly 
cuneate or rarely rounded at base, rather 
abruptly caudate-acuminate at apex 
(cauda 1/2-21/2 cm), chartaceous to 
coriaceous, mostly grey-green above 
when dry, ochraceous-brown beneath, 
upper surface dull but scarcely ‘shagreen- 
ed’ and glabrous or very sparsely pilose 
towards the base, lower surface also dull, 
very shortly tomentellous or spreading- 
puberulous, at least along the midrib; 
midrib moderate, cylindric-prominent 
beneath, not deeply impressed above; 
primary nerves 15—25 pairs, together with 
the numerous minor nerves conspicuous- 
ly prominulous, especially beneath; 
petiole 7-12 by 1-2 mm, shortly fulvo- 
tomentellous or rarely glabrescent. Jn- 
florescence 10-13 cm, sparingly and very 
shortly branched, sparsely fulvo-to- 
mentellous. Pedicels 11/2—21/2 cm, dense- 
ly fulvo-tomentellous. Sepals 5-8 by 
3-4 mm, externally densely fulvo- 
tomentellous. Disk-lobes c. 30, glabrous, 
epustulate, irregularly connate. Style 
glabrous. 

Distr. Malesia: Borneo (W. & E. 
Indonesian Borneo, Sarawak, Sabah). 

Ecol. Very varied: primary forest or 
disturbed forest on black or brown soil, 
once over sandstone, up to 60 m (Sabah); 
primary lowland Dipterocarp forest 
(Sarawak); loam soil and coral limestone, 
or sand and limestone, at 50-400 m (E. 
Borneo); old secondary forest, in bog on 
clay submerged during Westmonsoon, at 
40 m (W. Borneo; identity not quite 
certain). F/. June, Nov., fr. July, Sept., 
Dec. 

Vern. Bidaru, Sabah, ramin bukit, 
Sarawak, ngalin, njoelir (nyulir), tempé- 
éng, E. Borneo, mélingkat pépah, W. 
Borneo (?). 

Note. This species resembles 8. G. 
borneénsis in its conspicuous raised 
parallel nervation and large flowers, and 
13. G. affinis RADLK. in its more stiffly 
coriaceous leaves and subtomentellous 
indumentum, but the inflorescence and 


FLORA MALESIANA 


[ser. I, vol. 68 


flowers are larger than in either. It seems 
to be remarkably indifferent in its ecology. 


22. Gonystylus costalis Airy SHAW, Kew 
Bull. 23 (1969) 269. 

Small tree, 41/2 m. Leaves large, oblong- 
elliptic, 25-421/2 by 8-12 cm, base shortly 
cuneate or subrotundate, apex subrotun- 
date and shertly (11/2 cm) apiculate- 
caudate, margin conspicuously revolute, 
coriaceous, glabrous or with a few lax 
hairs beneath near the base of the midrib, 
brownish when dry, or greenish above, 
dull, under a lens densely minutely 
puncticulate above, very minutely granu- 
lose below; midrib moderately robust, 
prominent and 2—2!/2 mm thick beneath, 
broadly (2 mm) but shallowly elevate 
above and bordered on each side by a 
groove which is often indistinct or obso- 
lete on account of the oblique nerve-bases 
traversing it; principal nerves c. 30 pairs, 
scarcely distinguishable from the minor 
ones, prominulous on both surfaces, 
occasionally subbullately impressed, con- 
spicuously anastomosing near the margin 
and conspicuously abruptly decurving 
below into the midrib; petiole 11/2-2 cm 
by 4-6 mm, rugulose. /nflorescence robust 
up to 17 cm, with an occasional subbasal 
branch to 7 cm, rachis compressed, very 
shortly fulvous-tomentellous. Pedicels up 
to 2.3 cm, tomentellous. Calyx 7-9 mm 
long, externally sericeous. Disk-lobes 
30-35, subulate, glabrous, epustulate. 
Style 7-8 mm, glabrous. 

Distr. Malesia: Borneo (Central 
Sarawak). 

Ecol. Presumably evergreen rain-for- 
est, on ridge on sandy clay soil at 210 m. 
Fl. Oct. 

Note. Readily distinguished from all 
other species except 7. G. stenosepalus by 
the raised midrib on the upper surface of 
the leaves. From that species it differs in 
the much greater size of the leaves, the 
broader elevation of the midrib, the 
denser arrangement and _ occasionally 
bullate impression of the nerves, which 
are more or less decurrent on to the 
midrib, and by the much less conspicuous 
glandular puncticulation. 


23. Gonystylus decipiens AIRY SHAW, Kew 
Bull. 17 (1964) 454. 

Tree, 25-30 m. Leaves elliptic or almost 
oblong or oblanceolate, 15-20 by 6—7!/2 
cm, cuneate or somewhat rounded at 
base, shortly caudate-acuminate at apex, 
cauda obtuse, chartaceo-coriaceous, en- 
tirely glabrous except for the midrib, not 
or scarcely shining, greenish or fuscous 
above when dry, subochraceous below; 
midrib moderate, prominent and rather 
long-adpressed-pilose below, narrowly 


impressed and glabrous above; nerves 
densely parallel and prominulous, the 
primaries hardly distinct from the re- 
mainder; petiole 11-14 cm by 2-21/2 mm, 
striate, sparsely long-pilose or glabres- 
cent. Inflorescence 10-15 cm, rather many- 
flowered, with numerous abbreviated 
branches, very shortly grey-subsericeous. 
Pedicels 8-12 mm, sericeous. Calyx 5-7 
mm long and wide, sepals subobtuse. 
Disk-lobes 35-40, subulate, 3-4 mm, 
sparsely retrorse-setulose, epustulate. 
Style glabrous, surrounded at the base by 
4-5 very small subglobose parastyles. 

Distr. Malesia: Borneo (Central Sa- 
rawak). 

Ecol. Primary rain-forest on sand- 
stone, below 500 m. F/. Sept. 

Notes. Among the rather few species 
with setulose disk-lobes, this is perhaps 
nearest to 9. G. keithii, from which it is at 
once distinguished by the dense promi- 
nent venation with scarcely differentiated 
primary nerves, and by the quite different 
colour assumed by the leaves on drying, 
especially lacking the narrow brown 
margin of G. keithii. From the somewhat 
similar 8. G. borneénsis and 21. G. 
consanguineus it differs in the setulose 
disk-lobes and adpressedly long-pilose, 
rather than shortly tomentellous, midrib 
beneath. 


24. Gonystylus glaucescens Airy SHAW, 
Kew Bull. 17 (1964) 451. 

Tree, 10 m; branchlets conspicuously 
elevate-lenticellate. Leaves oblong-oblan- 
ceolate, 21-27 by 5-7 cm, + rounded- 
cuneate at base, narrowed and shortly 
(5-10 mm) acuminate-caudate at apex 
(acumen obtuse), margin distinctly thick- 
ened, rigidly coriaceous, quite glabrous, 
greenish and dull-shagreened above when 
dry, or scarcely shining, greyish-ochra- 
ceous beneath and very dull, as though 
glaucescent; midrib moderately robust, 
prominent and subcylindric beneath, 
slightly impressed or flat above; nerves 
numerous, widely spreading, the smaller 
veins clearly reticulate above but lax and 
rather indistinct below; petiole 1—-11/2 cm 
by 3-4 mm, rugose, glabrous. Jnflores- 
cence terminal, elongate, robust, 28 cm 
long, rachis 3-5 mm thick, conspicuously 
fuscous-lenticellate, in the fruiting stage 
finely fulvous-puberulous, branches ab- 
breviated, nodose, densely fulvous-pu- 
bescent. Flowers unknown, but (from the 
fruits) sepals 5-6 by 2-4 mm, subobtuse, 
ochraceous-sericeous; disk-lobes 25-30, 
glabrous, epustulate. Fruit (immature) 
obtusely tetragonous-globose, 3!/2-4 cm 
@, clearly 4-valved, pedicel 11/2-2 cm by 
3-4 mm, fulvous-puberulous. 

Distr. Malesia: E. Indonesian Borneo. 


Septem ber 1972] 


Addenda, corrigenda et emendanda 981 


Ecol. Sandstone ridge at 400 m alt. 
Fr. Sept. 

Note. The long, narrow, _ stiffly 
coriaceous leaves, more than 4 times as 
long as broad, with their dull, slightly 
shagreened, almost glaucescent surface, 
and the robust, elongate inflorescence, 
are highly distinctive. 


25. Gonystylus (§ Auxanthus) lucidulus 
Airy SHAW, Kew Bull. 17 (1964) 447. 

Tree to 36 m. Leaves oblong, elliptic- 
oblong or slightly lanceolate-oblong, 
10-20 by 5—7%/4 cm, base rounded (rarely 
slightly cuneate or very slightly cordate), 
apex narrowed or rounded, shortly 
abruptly caudate (cauda 5-15 mm), 
subacute or obtuse, moderately coria- 
ceous, greenish above when dry, grey- 
brown to purplish beneath, very smooth 
on both surfaces (except for the nerves) and 
manifestly somewhat shining, quite glab- 
rous, under a lens + clearly dark- 
puncticulate; midrib moderately robust, 
almost cylindric beneath, deeply impres- 
sed above; nerves slender, spreading, 
sharply prominulous on both surfaces; 
petiole 1.2-1.8 cm by 2-3 mm, grooved 
above, sparsely fulvous-puberulous. /n- 
florescence very similar to that of G. 
augescens, but the rachis often thicker 
and more nodose, and the bracts much 
broader, suborbicular, 7-8 mm broad, 
dorsally carinate. Immature fruit asym- 
metrically lanceolate-ovoid, 3.5 by 1.7 by 
1.5 cm, obliquely acuminate or sub- 
rostrate, 3-ridged, the ridges often marked 
with a slender groove, and alternating 
with 3 humps between the ridges towards 
the apex, very finely + stellate-ochra- 
ceous-puberulous, subtended by the 
persistent 5-6 mm long densely sericeous 
sepals and borne ona5 mm long densely 
grey-sericeous pedicel. 

Distr. Malesia: Borneo (NE. Sara- 
wak; Brunei). 

Ecol. Primary lowland Dipterocarp 
forest, 30-270 m, on yellow sandy clay. 
Fl. April-June, young fr. June. 

Vern. Ramin, Sarawak. 

Note. Closely related to 1. G. augescens, 
of SW. Sarawak, and adjacent W. 
Indonesian Borneo, but differing in the 
smaller and narrower, smooth and 
shining leaves, and in the total lack of the 
minute adpressed hairs, lying parallel to 
the nerves on the undersurface, which are 
always found (by careful searching!) in 
that species. 


26. Gonystylus nervosus Airy SHAW, Kew 
Bull. 17 (1964) 452. 

Small tree, 4-5 m, Leaves elongate- 
oblanceolate, rarely oblong, 17-34 by 
5-9 cm, cuneate at base, narrowed to 


subrotundate at apex and abruptly 
narrowly caudate, cauda 1-4 cm, acute, 
chartaceous, glabrous, not shining, ob- 
scurely grey-brown above when dry, 
similar or subcastaneous below; midrib 
relatively slender, prominent below, 
impressed above; primary nerves 25-30 
pairs, strictly parallel, sharply raised 
beneath, subbullately impressed above, 
conspicuously arcuate-anastomosing to- 
ward the margin; petiole 1-11/2 cm by 
2-4 mm, fulvous-pubescent at first, 
finally glabrescent. /nflorescence 2-4 cm, 
few-flowered, grey-sericeous. Flowers on- 
ly known in bud stage: disk-lobes about 
30, glabrous, epustulate; style glabrous. 
Fruit ellipsoid, 31/2 cm, 3-valved, subtend- 
ed by 5-6 mm long sepals and borne on 
a 2 cm long peduncle. Seeds 2, flattened 
hemi-ellipsoid, 2.2 by 1.6 cm, shining, 
light chestnut. 

Distr. Malesia: Borneo (SW. & NE. 
Sarawak). 

Ecol. In primary rain-forest, appar- 
ently always on or near limestone hills, 
60-270 m. F/. June, July, fr. June, August. 

Note. The elongate leaves, with their 
somewhat bullately impressed main 
nerves, recall those of 2. G. areolatus, but 
G. nervosus is probably most closely 
related to 8. G. borneénsis. The latter 
species, however, seems almost always to 
occur on more or less acid soils (basalt, 
kerangas, efc.); I have only seen one 
collection from limestone. 


27. Gonystylus nobilis Airy SHAW, Kew 
Bull. 23 (1969) 271. 

Tree to 24 m; branches robust, almost 
1 cm @. Leaves large, elliptic to sub- 
oblong, 23-34 by 9-12 cm, base broadly 
cuneate to rounded, apex narrowed to 
somewhat rounded, very shortly, broadly 
and not abruptly acuminate (acumen 
acute, | cm), firmly coriaceous, quite 
glabrous, dull and grey-green above when 
dry, with a very narrow purplish margin, 
smooth beneath, not shining, distinctly 
pale when dry; midrib robust, subcy- 
lindrically prominent beneath, deeply and 
narrowly impressed above, primary 
nerves c. 20 pairs, lax, conspicuously 
anastomosing near the margin, obscurely 
prominulous above, sharply prominent 
beneath; petiole very robust, 3-4 cm by 
4-5 mm, terete, sparsely adpressed- 
pubescent. Inflorescence 11 cm, simple, 
rachis terete, 3-4 mm @, adpressed- 
pubescent, flower-nodules very short. 
Fruit transversely subellipsoid, 61/2 by 
51/2 cm, conspicuously obtusely 4-ridged, 
strongly rugulose. 

Distr. Malesia: Borneo (W. Central 
Sarawak). 

Ecol. Primary lowland Dipterocarp 


982 


FLORA MALESIANA 


[ser. I, vol. 68 


6: 


forest, altitude and soil unknown. 

Vern. Ramin. Sarawak. 

Note. Notable for its large glabrous 
leaves and especially for the remarkably 
long, robust, terete petiole. The nervation 
and the colouring of the leaves on drying 
somewhat recalls 9. G. keithii on a larger 
scale. 


28. Gonystylus spectabilis AIRY SHAW, 
Kew Bull. 23 (1969) 269. 

Tree, 24 m; branches robust, ridged, 
tomentellous. Leaves broadly elliptic- 
oblong, up to 32 by 14!/2 cm, slightly but 
distinctly cordate (rarely rounded) at 
base, rounded and shortly (to 2 cm) 
abruptly cuspidate-caudate, chartaceous 
to thinly coriaceous, glabrous above, 
finely velutinous beneath with short 
white hairs, bright green (especially be- 
low) when dry, slightly shining above; 
midrib strong, very prominent below, 
3-4 mm thick, densely fulvo-tomen- 
tellous, narrowly impressed above; main 
nerves (scarcely distinguishable from the 
minor ones) 15-20 pairs, slender, sharply 
prominulous below, scarcely so above; 
minor nerves very numerous; petiole 
robust, 11/2-21/2 cm by 45 mm, 
longitudinally rugose, often almost tetra- 
gonous, densely tomentellous. /nflores- 
cence robust, terminal, to 22 cm, little 
branched; rachis compressed, up to 6 mm 
thick, strongly grooved, fulvo-tomentel- 
lous, branches to 7 cm, widely spreading. 
Pedicels to 2!/2 cm by 21/2 mm, tomentel- 
lous. Calyx 1-1.2 cm, truncate at base, 
externally tomentellous. Disk-lobes 40- 
45, narrowly subulate, glabrous, epustul- 
ate. Style 8-9 mm, glabrous. 

Distr. Malesia: Borneo 
Sarawak). 

Ecol. Presumably rain-forest, on ridge 
at 195 m. Fi. Oct. 

Note. Related to 4. G. reticulatus, 3. 
G. calophyllus and 20. G. calophylloides, 
but differing in its dense fulvous-tomen- 
tellous indumentum and in its petioles 
sometimes reaching 21/2 cm; further from 
G. reticulatus in the absence of a narrow 
brown border to the leaves when dry, and 
from G. calophyllus in the 40-45 rather 
than 20-30 disk-lobes. 


(Central 


Aquilarioideae & Thymelaeoideae (DiING Hov) 


9a Aquilaria malaccensis LAMK. 


23 


Add to synonymy: A. agallocha Roxs.: 
cf. DinG Hou, Blumea 12 (1964) 286. 
Enkleia GRirF. 
Add to references: NEVLING, J. Arn. Arb. 
42 (1961) 373-396, 9 fig., map. 

He gave an extensive, detailed treat- 
ment of the morphology and anatomy, 
including a systematical revision. He 


27b 


35a 


added one new species from SE. Asia. 
See the remark below on the generic 
difference with Linostoma. 
E. malaccensis seems to be restricted to 
Malesia. 
Linostoma WALL. ex ENDL. 
Add to references: NEVLING, J. Arn. Arb. 
42 (1961) 295-320. 
NEVLING does not distinguish sections 
within the genus to which I agree. 
Furthermore he remarked correctly 
that the generic difference between 
Linostoma and Enkleia tends to disappear 
as the main differential character : stamens 
in One or two series respectively is rather 
breaking down in one species which is 
almost intermediate in this respect. 
Linostoma pauciflorum GRIFF. 
Add to Distr.: NEVLING, who first 
hesitated to accept Psilaea dalbergioides 
Mia. from Sumatra as a synonym (1/.c.), 
later traced its type in CAL and found 
(J. Arn. Arb. 43, 1962, 221) it correctly 
reduced by DinG Hou. Thus there are 2 
collections from Sumatra: mainland, pr. 
Sibolga and Simalur I. 
Linostoma longiflorum HAL. f. is by 
NEVLING (J. Arn. Arb. 42, 1961, 313) 
reduced to L. pauciflorum; he showed that 
the size and shape of the bracts is variable, 
but he found no such graded variability 
in the flower length. 
Line 14 from top, replace ‘375’ by: 357. 


Trapaceae 


The family name Hydrocaryaceae must 
give way to the now conserved name 
Trapaceae. 


Typhaceae 


Typha L. 

Miss B. G. BricGcs and Dr. L. A. S. 
JOHNSON (Contr. N.S.W. Nat. Herb. 4, 
1968, 57-69, 2 fig., 1 tab.) have published 
a detail account of Typha in Australia in 
which they also examined Malesian 
material. They maintain that what has 
been called in Malesia and Australia 7. 
angustifolia L. sens. lat. is not that species 
which to them is confined to the northern 
hemisphere; however, they provide for 
this no further arguments. The Malesian 
and Australian material they bring to two 
species, 7. domingensis PERS. 1807 and 7. 
orientalis PR. 1852 (type from the 
Philippines). On the basis of the Austra- 
lian material it was established that two 
taxa are concerned, which differ in 
chromosome number, JT. domingensis 
2n=—30 and T. orientalis 2n=60. They 
differ also in minute other characters. 
The authors state that they are often 
found in mixed stands but do not hybrid- 


September 1972] 


Addenda, corrigenda et emendanda 


983 


ize; this is not so astonishing, as hybrids 

between 7. angustifolia and T. latifolia 

(2n also 30) are in Europe also very rare. 
The two taxa, which both occur in 

Malesia, can be distinguished by some 

minute morphological details. To my 

view they are merely cyto-subspecies. 

For those who want to do further work 

on this in Malesia I have copied the key 

given: 

1. Bracts in the 2 inflorescence numerous, 
broadly spathulate (usually 4—8 cells 
across the lamina). Stigmas linear. 
Mature 2 spikes !/2-2 cm @, length 
6-20(—30) times @, cinnamon-brown 
(due to the numerous pale ends of the 
bracts interspersed among the darker 
stigmas and the carpodia). 3 and & 
spikes separated by (4/2-)2-5 cm. 
Sheaths of the upper leaves not auric- 
ulate or only the uppermost 1-2 


leaves distinctly auriculate. 
T. domingensis 
1. Bracts in the 2 inflorescence few, or 
sometimes apparently absent, narrow- 
ly spathulate (usually 3-4 cells across 
the lamina). Stigmas narrow-obovate. 


Mature & spikes 1-3 cm @, length 
5—10(-18) times @, chestnut-brown 
(due to the great predominance of the 
brown stigmas at the surface). J and & 
spikes contiguous or separated by up 
to 2(-6) cm. Sheaths of the 2-4 
uppermost leaves usually distinctly 
auriculate T. orientalis 


4: 243a Typha angustifolia L. 


4: 244b 


4: 


117 


Add to synonymy: 7. angustata BoRY & 
CHAUBARD, Exp. Sc. Morée 2, | (1832) 
3385; HENDERSON, Mal. Wild Flow., 
Monoc. (1954) 211 f. 126. 

Add to Distr.: Malay Peninsula: Kuala 
Selangor; Kedah: Kuala Muda; Lang- 
kawi: Tasek Bayang Bunting. 

Add to Vern.: Bulrush, E, banat, M. 
Note. By a curious oversight this was 
omitted from RIDLEY’s Materials and his 
Flora. Near Kuala Selangor it is found in 
ditches along the roadside in association 
with Acrostichum aureum and Pluchea, 
both typical for brackish water. 


Umbelliferae 


Trachymene RUDGE. 


Fig. 20. Trachymene tripartita HOOGL. a. Habit, 1, b. leaf-blade, < 10, c. leaf-sheath, = 10, d. involucral 
bract, x10, e. & f. flower with only one petal still present, = 15, g. fruit, x 15 (a-g HOOGLAND & PULLEN 
5965). 


984 


4: 118 


FLORA MALESIANA 


Many new collections of the genus have 
come in from the Papuan highlands and a 
revised account has become most desir- 
able. A newly described species is to be 
added: 


Trachymene tripartita HooGL. Blumea 
Suppl. 4 (1958) 231. — Fig. 20. 
Glabrous perennial. Stem little branch- 
ed, bearing rosettes. Leaves: sheath 4-8 
by 2 mm, attenuate into petiole; petiole 
7-30 mm, exceeding the blade; blade 
3-partite, 5-8 by 5-8 mm, broad-cuneate 
at the base, lobes almost equal, obtuse, 
mucronate. Umbel 1 from a rosette, 
peduncle + as long as the leaves or + 
shorter, in fruit twice as long, striate or 
subsulcate; involucral bracts 5—7, linear- 
lanceolate, 3-4 mm by 3/4 mm, obtuse, 
subappressed. Flowers 5—10, in 1(—2) rows. 


[ser. I, vol. 68 


5: 556b 


Pedicels in fl. 1-2 mm, in fr. to 7 mm, 
incurved. Calyx lobes minute. Petals 
rounded, c. 1 by 0.8 mm. Filaments 
linear, widened to base, 0.4 mm; anther- 
cells 0.3 by 0.15 mm. Ovary 0.7 by 1 mm; 
style c. 0.4 mm. Mericarps 2 by 1.7 mm, 
equal; carpophore undivided, 4-apicul- 
ate, sulcate. 

Distr. Malesia: East New Guinea 
(Mt Wilhelm), open rocky slopes and 
tussock grasslands, 3600-3750 m. 

Note. Closest related to 7. novo- 
guineensis (DOMIN) Buw., clearly char- 
acterized by the 3-partite leaves with 
undivided lobes. 


Daucus glochidiatus (LABILL.) FISCHER. 
Add to Distr.: East New Guinea: 
Foramburo, 2500 m, 28 Oct. 1960, E. 
BORGMANN 340. 


INDEX TO SCIENTIFIC PLANT NAMES 


compiled by 


M. J. VAN STEENIS- KRUSEMAN 


Families and higher taxa have been entered under their name. 
Names of families which have been revised in volumes 4, 5, and 6 have been entered and are printed 
in bold type, so that as far as this is concerned this index is complete for all preceding volumes as well. 
Suprageneric epithets have been entered under the family name to which they belong preceded by the 
indication of their rank (subfamilies, tribes, efc.). 
Infrageneric epithets have been entered immediately under the generic name to which they belong 
preceded by the indication of their rank (subgenera, sections, series, e/c.). 
Infraspecific epithets have been entered under the specific name to which they belong preceded by the 
indication of their rank (subspecies, variety, forma, efc.). 
New names and new combinations have been printed in bold type, synonyms in italics. 
‘Map’ printed behind a page number denotes that a map of the concerned taxon is present on that 


page. 


An asterisk behind a page number denotes the presence of a figure of the concerned taxon. 

Page numbers in bold type denote main treatment. 

Some minor printing errors in plant names have been corrected. 

Of synonyms with a double authority, the latter has not always been cited in full. See for example 
under Dortmann(i)a Adans. which was accepted by O. Kuntze to replace Lobelia. The full authority can 


easily be derived from the text. 


Acer garrettii Craib 915 
laurinum Hassk. 915 
Aceraceae 4: 3—4, 592; 6: 915 
Actinidiaceae s. str. 4: 37-39 
Adenanthera triphysa Dennst. 
219 
Adenoplea Radlk. 295, 296, 337, 
953 
madagascariensis Eastw. 340 
Adenoplusia Radlk. 295, 296, 
337, 953 
Adnaria /anceolata (Bl.) O.K. 
761 
Aegiphila viburnifolia Juss. 248 
Aegle marmelos (L.) Correa 69 
Aeschrion Velloso 212 
excelsa (Sw.) O.K. 193 
Aétoxylon 3, 5, 6 
Agallochum Rumph. ex Lamk 6 
coinamense Rumph. 9 
malaccense O.K. 9 
malaicense Rumph. 9 
secundarium Rumph. 9 
Agapetes D.Don 469, 471, 473, 
474, 746, 878, 879 map 
sect. Pseudagapetes Airy Shaw 
878 


subg. Dimorphanthera 
(F.v.M.) Drude 885 
acuminatissima (Miq.) 
Niedenzu 761 
alberti-eduardi Sleum. 879, 
882*, 883 
amblyornidis Becc. 892 
beccariana Koord. 904 
brassii Sleum. 879, 881, 882* 
brevicuspis Sleum. 884 
carrii Sleum. 879, 881, 882* 


coriacea (Bl.) G.Don 828 
costata C.H.Wright 879,882*, 
883 
cuneifolia (Bl.) G.Don 874 
elliptica (Bl.) G.Don 873 
filicicola Sleum. 883 
floribunda (Bl.) G.Don 828, 
872 
forbesii F.v.M. 904 
griffithii (non Clarke in 
Hook.) K. & G. 753 
helenae (F.v.M.) F.v.M. ex 
Sleum. 878, 879, 882* 
lanceolata (Bl\.) Niedenzu 761 
laurifolia (Bl.) G.Don 872 
leptantha (Miq.) Niedenzu 764 
lucida (B\.) G.Don 814 
meliphagidum Becc. 906 
micrantha Ridl. 753 
microphylla Jungh. 814 
moorhousiana Lane-Poole 892 
moorhousiana F.v.M. 893 
myrtoidea = myrtoides 812 
myrtoides (Bl.) G.Don 812 
myzomelae Becc. 913 
obtusata Sleum. 884 
parviflora Ridl. 753 
perakensis Ridl. 753 
polyantha (‘Miq.’) Niedenzu 
872 
prainiana Koord. 911 
pubescens Ridl. 753 
rubrocalyx Sleum. 879, 881 
var. pilicalyx Sleum. 881 
var. rubrocalyx 881 
sclerophylla 879, 880, 882* 
scortechinii (K. & G.) Sleum. 
878, 879, 880*, 882* 


stenantha (Schltr) Sleum. 879, 
884* 
varingiaefolia (Bl.) G.Don 815 
viridiflora (Schltr) Sleum. 879, 
883 
vitis-idaea Sleum. 879, 881, 
882* 
vonroemeri Koord. 911 
vulgaris Jungh. 815 
wrayi Ridl. 753 
wrightiana Koord. 914 
Agapetes (non D.Don) Koord. 
885 
Agelaea borneensis (Hook. f.) 
Merr. 933 
insignis (Schellenb.) Leenh. 
933 
macrophylla (Zoll.) Leenh. 
933 
trinervis (Llanos) Merr. 933 
Aglaia sapindina (F.v.M.) 
Harms 237 
Aglaiopsis glaucescens Mig. 237 
miquelii Merr. 237 
Aidia cochinchinensis Lour. 309 
Ailanthopsis  poilanei Gagnep. 
226 
Ailanthus Desf. 193, 194, 195, 
196, 215, 216, 969 
altissima (Mill.) Swingle 193, 
215,216; 2075. 220 
blancoi Merr. 218 
cacodendron L’Herit. 220 
calycina Pierre 218 
erythrocarpa Carr. 220 
esquirolii Léveillé 970 
excelsa Roxb. 216, 217*, 219 
fauveliana Pierre 219 


986 


FLORA MALESIANA 


[ser. I, vol. 66 


fordii Nooteboom 216, 217*, 
220, 969* 
giraldii Dode 220 
glandulosa Desf. 220 
gracilis Salisb. 211 
grandis Prain 218 
imberbiflora F.v.M. 219 
integrifolia Lamk 193, 206*, 
216, 217*, 218 map, 220 
ssp. calycina (Pierre) 
Nooteboom 218 map 
ssp. integrifolia 218 map 
var. B Lamk 219 
Kurzii Prain 219 
macrophylla Hort. 220 
matirei Gagnep. 220 
malabarica DC. 218, 219, 969 
var. mollis K. & V. 219 pro 
parte 
malabarica (non DC.) F.-Vill. 
218 
var. mollis K. & V. 218 pro 
parte 
mascula Hort. 220 
moluccana DC. 218 
var. mollis (non K. & V.) 
Koord. 218 
peekelii Melch. 218 
var. glabrata C.T.White 218 
peregrina F.A.Barkl. 220 
Philippinensis Merr. 219 
pongelion Gmel. 218, 220 
procera Salisb. 220 
punctata F.v.M. 220 
rhodoptera F.v.M. 220 
rubra Hort. 220 
scripta Gagnep. 220 
sutchuensis Dode 220 
triphysa (Dennst.) Alston 193, 
217*, 219 map, 220 
vilmoriniana Dode 220 
Ailantus = Ailanthus 215 
Aizoaceae 4: 267-275; 6: 384, 
450, 915 
Albonia Buchoz 215 
peregrina Buchoz 220 
Alfaroa 143, 146, 148 
Alismataceae 5: 317-334; 6: 158, 
915 
Allanthospermum Forman 968, 
969, 970, 971*, 972 
borneense Forman 971*, 972 
ssp. rostratum Nooteboom 
971*, 972 
multicaule (Capuron) 
Nooteboom 970, 972* 
Allospondias lakonensis (Pierre) 
Stapf 226 
Alnus japonica (Thunb.) Steud. 
917 
maritima (Marsh.) Nuttall 917 
Aloe 31 
Alphonsea arborea (Blco) Merr. 
421 
Alsodeia chrysodasys Mig. 944 


glabra Burgersd. 235 
Alternanthera Forsk. 915 
bettzichiana = bettzickiana 
916 
bettzickiana (Regel) Nichols. 
916* 
ficoidea (L.) R.Br. ex R. & S. 
915 
ssp. bettzickiana (Nichols.) 
Backer 917 
var. versicolor (Regel) 
Backer 916 
ficoides P.Beauv. 915 
paronychioides St.Hill 915, 
916 
var. paronychioides 916* 
Alytostylis Hook. f. 95 
Alyxia spanogheana Mig. 336 
tetragona R.Br. 336 
Amaracarpus 387 
Amaranthaceae 4: 69-98, 593; 5: 
554; 6: 450, 915-917 
Amaranthus dubius Mart. 915 
hybridus L. 
ssp. incurvatus (Gren. & 
Godr.) Brenan 
var. paniculatus (L.) 
Mansf. 915 
Amaroria A.Gray 221 
soulameoides A.Gray 221 
Amentiferae 143 
Amoora aphanamixis Auct. non 
Re 8S. 921 
Amphicalyx heterophyllus (Bl\.) 
Hassk. 738 
latifolius Bl. ex Hassk. 738 
pilosa Bl. 729 
Amyxa cf. Airy Shaw 3, 5, 6, 47 
pluricornis (Radlk.) Domke 
47* 
Anacardiaceae 194, 
226, 291, 928 
Anacolosa frutescens BI. 420 
Anacyclodon Jungh. 424, 425 
pungens Jungh. 431 
Anagallis L. 173, 174, 175, 176 
subg. Anagallis 176 
subg. Centunculus (L.) 
P.Taylor 176 
subg. Jirasekia (Schmidt) 
P. Taylor 176 
arvensis L. 176 
nana Schinz 176 
pumila Sw. 176* 
var. pumila 176 
Anasser Juss. 369 
laniti Blco 373 
moluccana Lamk 373 
Anassera rumphii Span. 373 
Ancistrocladaceae 4: 8-10; 5: 
553 
Andrachne cavalerieri Léveillé 
179 
Andresia Sleum. 669, 943 
malayana (Scort. in Hook. f.) 


1O5SE220) 


Sleum. 669* 
Andreya 295, 296 
Andromeda 472 
ovalifolia Wall. 675 
Androsace Tourn. 173, 175, 186 
rotundifolia Hardw. 192 
saxifragifolia Bunge 191, 192 
tonkinensis Bonati 382, 383 
umbellata Merr. 191, 192 
Anhella tristis 873 
Anisophyllea R.Br. ex Sabine 
966 
beccariana Baill 966 
ferruginea Ding Hou 966 
grandis (Bth.) Burk. 361 
Annamocarya Chev. 143 
Annonaceae 421 
Annulodiscus Tardieu Blot 932 
nigricans Tardieu Blot 932 
Anthocleista 293, 295 
Anthodendron Rchb. 661 
Antirrhoea hexasperma (Roxb.) 
Merr. 38 
Antonia Pohl 295, 296 
griffithii Wight 341 
Antoniaceae Hutch. 296 
Apocynaceae 93, 294, 297, 315, 
318, 363, 368, 369 
tribe Tabernaemontanineae 
297 
Apodostigma 227, 390 
Aponogetonaceae 4: 11-12; 5S: 
553 
Aquilaria Lamk 2, 3, 4, 5, 6, 31 
sect. Agallochum Hall. f. 6 
sect. Amphinoman Hall. f. 6 
sect. Brachythalamus Hall. f. 
39 
sect. Gyrinops Hall. f. 39 
sect. Gyrinopsis Hall. f. 6 
sect. Lachnolepsis Hall. f. 39 
acuminata Quis. 11 
agallocha Roxb. 982 
apiculata Merr. 2, 8, 11 
bancana Miq. 15 
beccariana v. Tiegh. 7*, 8, 13, 
41 
borneensis Gilg 9 
brachyantha (Merr.) Hall. f. 
7* 2810812 
citrinaecarpa (Elm.) Hall. f. 8, 
10 
cumingiana (Decne) Ridl. 1, 
4.6. 7*. 85 la sks 
var. parviflora Airy Shaw 13 
filaria (Oken) Merr. 8, 11 
map, 12 
grandifolia Domke 13 
hirta Ridl. 7*, 8, 12 
macrophylla Miq. 15 
malaccensis Lamk 7*, 8, 9, 
982 
microcarpa Baill. 7*, 8, 9 
moluccana Hall. f. 39 
moszkowskii Gilg 12 


Sept. 1972] 


Index to scientific names 


987 


ovata Cav. 9 
parvifolia (Quis.) Quis. 8, 12 
pentandra Blco 15 
podocarpus Hall. f. 42 
rostrata Ridl. 8, 13 
secundaria DC. 9 
tomentosa Gilg 11 
urdanetensis (Elm.) Hall. f. 6, 
8, 10 
versteegii Hall. f. 42 
Aquilariella v.Tiegh. 6 
borneensis v.Tiegh. 9 
malaccensis v.Tiegh. 9 
microcarpa v.Tiegh. 9 
Araliaceae 194 
Araucaria 356 
Arbor coeli sive Caju langit 
Rumph. 218 
Arbutus 472 
coriacea Bl. 828 
Arctostaphylos 472 
Ardisia ochracea Elm. 260 
Argyreia capitata (Vahl) Choisy 
941 
capitiformis (Poir.) Ooststr. 
941 
congesta Ooststr. 941 
Arthrosolen 4 
Aruba Aubl. 202 
Arytera litoralis Bl. 226 
Asperula 294 
Aspidopterys elliptica (BI.) Juss. 
960 
Astrephia chinensis Dufr. 455 
Astroloma R.Br. 424 
Azalea (non L. sensu Salisb.) 
Desv. 661 
subg. Pentanthera (G.Don) 
K.Koch 661 
subg. Tsutsutsi (G.Don) 
K.Koch 661 
brookeana (Low ex Lindl.) 
O.K. 655 
citrina Hassk. 579 
indica L. 663 
indica (non L.) Sims 665 
var. angustifolia Bl. 663 
var. floribunda Bl. 663 
var. spathulata Bl. 663 
var. variegata Bl. 663 
Jasminiflora (Hook.) O.K. 565 
javanica (Bl.) O.K. 624 
lamponga (Miq.) O.K. 544 
ledifolia Hook. 662 
malayana (Jack) O.K. 532 
mollis Bl. 661 
moulmainensis (Hook.) O.K. 
666 
mucronata Bl. 662 
multicolor (Miq.) O.K. 621 
retusa (Bl.) O.K. 482 
rosmarinifolia Burm. f. 662 
sinensis Lodd. 661 
subsessilis (Rendle) Copel. f. 
664 


teysmannii (Miq.) O.K. 626 
tubiflora Bl. ex DC. 532 


Banksia Forst. 44 
musculiformis Gaertn. 367, 
368 
Basellaceae 5: 300-304; 6: 450 
Batidaceae 5: 414-415; 6: 917 
Batis argillicola v.Royen 917 
Béla-Moédagam Rheede 951 
Bennettia papuana Gilg 944 
Bernadina Baudo 177, 184 
laurina Baudo 
var. stenia Baudo 184 
mauritiensis Baudo 184 
parviflora Baudo 185 
Bertuchia Dennst. 299, 301 | 
Betula-Alnus maritima Marsh. 
917 
Betulaceae 5: 207-208; 6: 143, 
472, 917 
Bhesa Ham. ex Arn. 227, 228, 
230, 231, 232, 280, 283 map, 
390, 391, 930 
archboldiana (Merr. & Perry) 
Ding Hou 281%, 282, 284 
indica (Bedd.) Ding Hou 
281*, 282, 283 
moja Ham. ex Arn. 283 
paniculata Arn. 229, 231, 280, 
281*, 282, 283 
robusta (Roxb.) Ding Hou 
280, 281*, 282, 283 
Biporeia Petit-Thouars 199 
Bischofia javanica Bl. 53 
Bixaceae s. str. 4: 239-241 
Blumeodendron papuanum P. & 
H. 944 
Blyxa novoguineensis Hartog 
952 
Boerhaavia = Boerhavia 452 
Boerhavea = Boerhavia 454 
Boerhavia L. 450, 451, 452 
sect. Adenophorae Heimer! 
452 
acutifolia S.Moore 454 
chinensis (L.) Aschers. & 
Schweinf. 450, 452, 453%, 
454, 455 map 
diandra L. 454 
diffusa L. 453*, 454 
f. B paniculata (Rich.) O.K. 
454 
f. y repens (L.) O.K. 454 
var. acutifolia DC. 454 
var. obtusifolia DC. 454 
var. pubescens DC. 454 
erecta (non L.) Burm. f. 454 
erecta L. 452, 453*, 454 
glabrata Bl. 454 
glutinosa Vahl 454 
helenae R. & S. 455 
hirsuta L. 454 
mutabilis R.Br. 454 
var. pubescens (R.Br.) 


Choisy 454 
procumbens Banks ex Roxb. 
454 
pubescens R.Br. 454 
repanda Willd. 455 
repens L. 454 
var. diffusa (L.) Boiss. 454 
var. procumbens (Roxb.) 
Hook. 454 
scandens L. 
var. chinensis (L.) O.K. 455 
tetrandra Forst. 454 
Bonamia Thou. 936 
Bonyunia Schomb. 295, 296, 954 
Boscia 61 
Bougainvillea Comm. ex Juss. 
450, 451, 455 
sect. Tricycla 456 
< buttiana Holtt. & Stand. 
456 
glabra Choisy 451, 457 
peruviana Humb. & Bonpl. 
456, 457 
spectabilis Willd. 456, 457 
var. glabra (Choisy) Hook. 
457 
Brachythalamus Gilg 39 
caudatus Gilg 42 
podocarpus Gilg 42 
versteegii Gilg 42 
Brassiantha A.C.Smith 227, 230, 
389, 390, 391, 392, 393, 394, 
930 
pentamera A.C.Smith 393*, 
394 
Bromheadia Lindl. 976 
Brossaea bandongensis (Zoll. ex 
Miq.) O.K. 695 
fragrantissima (non Wall.) 
O.K. 688 
leucocarpa Bl. 692 
nummularioides (D.Don) 
O.K. 684 
a. normalis O.K. 684 
B glauca O.K. 684 
Brownlowia tersa (L.) Kosterm. 
203 
Brucea J.F.Mill. 193, 194, 196, 
209 
acuminata Li 212 
amarissima Desy. ex Gomes 
211 
dubia Steud. 214 
glabrata Decne 211 
gracilis DC. 211 
javanica (L.) Merr. 195, 209, 
210*, 211 map, 212 
luzonensis Vidal 212 
macrobotrys Merr. 212 
membranacea Merr. 212 
mollis Wall. ex-Kurz 193, 
210*, 211, 212 map 
quercifolia Seem. 212 
stenophylla Merr. 212 
sumatrana Roxb. 211 


988 


FLORA MALESIANA 


[ser. I, vol. 68 


sumatrensis Spreng. 211 
Bruguiera Lamk 965, 966 
eriopetala W. & A. ex Arn. 
var. exsetata Val. 966 
exaristata Ding Hou 966 
gymnorrhiza (L.) Lamk 966 
hainesii C.G.Rogers 966 
Bruinsmia Boerl. & Koord. 976 
styracoides Boerl. & Koord. 
976 
Buddleia, see Buddleja 336 
Buddleja Houst. ex L. 293, 294, 
295, 296, 297, 336, 953, 954 
subg. Buddleja 336, 337 
sect. Alternifolia Marquand 
339 
subg. Nicodemia Leenh. 336, 
337, 340 
acuminatissima Bl). 337 
amentacea Kranzl. 339 
arfakensis Kaneh. & Hatus. 
339 
asiatica Lour. 293, 336, 337, 
338*, 339%: 954 
var. brevispica Val. 337 
var. densiflora (Bl.) K. & V. 


337 

var. salicina (Bl.) K. & V. 
337 

var. sundaica (Bl.) K. & V. 
S57 


brasiliensis Jacq. f. 340 
crispa Bth. 337 
curviflora H. & A. 337 
davidii Franchet 336, 340, 954 
densiflora Bl. 339 
diversifolia Vahl 340 
gynandra Marquand 336 
indica Lamk 340 
japonica Linden 337 
lindleyana Fortune 337 
madagascariensis Lamk 337, 
340 
neemda Buch.Ham. ex Roxb. 
337 
var. philippinensis Cham. & 
Schlechtend. 337 
officinalis Maxim. 336 
otophylla Hassk. 340 
paniculata Wall. 337 
salicina Lamk 337 
sundaica Bl. 339 
venenifera Makino 337 
virgata L. f. 339 
Buddlejaceae 295, 954 
Buginvillaea = Bougainvillea 
455 
Buginvillia racemosa Blco 460 
Bulbophyllum 109, 113 
Burmanniaceae 4: 13-26, 592; 5: 
553 
Burseraceae 5: 209-296, 567; 6: 
194, 195, 917-928 
Busbeckea Endl. 69 
Butomaceae 5: 118-120, 566 


Cactaceae 450 
Cadaba Forsk. 61, 62, 93, 97 
map 
sect. Cadaba 93 
sect. Eu-Cadaba Endl. 93 
capparoides DC. 62, 93, 94*, 
95*, 97 map 
Cadellia 194 
Callitrichaceae 4: 251-252 
Calogyne pilosa R.Br. 951 
Calpidia Thouars 457, 459 
brubobiana (Endl.) Heimerl 
461 
cauliflora (Scheff.) Heimer] 
462 
corniculata Heimer! 464 
cuspidata Heimer! 462 
excelsa Heimer] 461 
grandifolia Heimer! 463 
lauterbachii Warb. ex Heimer] 
463 
longirostris Heimer! 463 
miilleriana (Warb.) Heimer] 
462 
pancheriana 459 
rostrata Heimerl 463 
spathiphylla Heimer! 463 
Camotain Blco 355 
Campanopsis (R.Br.) O.K. 111 
marginata O.K. 116 
var. rigida O.K. 116 
Campanula 107, 108, 109, 112 
sect. Campanopsis R.Br. 111 
agrestis Wall. 115 
carnosa Wall. 111 
ceylanica Seba 129 
circaeoides Fr.Schmidt ex 
Mig. 111 
dehiscens Roxb. ex Wall. 115 
gracilis Forst. 115 
gracilis (non Forst.) Sims 118 
indica Dietr. 115 
lancifolia Roxb. 120 
lavandulaefolia Reinw. ex BI. 
ii 15) 
littoralis Labill. 115 
marginata Thunb. 115 
medium L. 141 
quadrifida R.Br. 115 
rapunculoides L. 141 
rotundifolia 110 
sieberi Dietr. 115 
vincaeflora Vent. 118 
Campanulaceae 107-141, 928 
subfam. Campanuloideae 108 
tribe Campanuleae Bth. 109 
tribe Pentaphragmeae 
Schonl. 109 
tribe Sphenocleae Schon. 
09 


subfam. Lobelioideae 108, 122 
tribe Lobelieae Bth. 109, 
140 
Campanulopsis Zoll. & Mor. 111 
cyanea Zoll. & Morr. 115 


Campanumoea BI. 109, 118, 119 
axillaris Oliv. 120 
celebica Bl. 120, 121 
cordata Miq. 120 
Japonica Maxim. 120 
javanica Bl. 119 
var. japonica (Maxim.) Ma- 
kino 120 
lancifolia Merr. 119, 121 
maximowiczii Honda 120 
truncata Diels 120 
Campylostemon Welw. 389, 
390 
Canarina moluccana Roxb. 928 
Canarium Stickm. 917, 921 
subg. Africanarium Leenh. 922 
sect. Africanarium Leenh. 
922 
subg. Canariellum 922 
subg. Canarium 921 
sect. Canarium 922, 924, 
927 
sect. Pimela 922, 924, 927, 
928 
acutifolium (DC.) Merr. 926 
var. acutifolium 926 
album (Lour.) Raeusch. 922, 
926, 928 
apertum H.J.Lam 924 
asperum Bth. 922, 926, 927 
ssp. asperum 
var. asperum 926 
australianum F.v.M. 925 
bacciferum Leenh. 926 
balsamiferum Willd. 924 
caudatum King 923, 927 
f. auriculiferum Leenh. 923 
f. caudatum 922 
cestracion Leenh. 922, 925 
chinare Grutterink & HJ. 
Lam 922, 925 
decumanum Gaertn. 924 
denticulatum Bl. 924 
ssp. kostermansil 
924 
dichotomum (BI.) Miq. 925 
divergens Engl. 922, 923 
euryphyllum Perk. 924 
fusco-calycinum Ridl. 925 
gracile Engl. 925 
grandifolium (Ridl.) H.J.Lam 
924 
hirsutum Willd. 925 
var. hirsutum 925 
indicum L. 924 
intermedium H.J.Lam 923, 
925 
kaniense Laut. 924 
karoense H.J.Lam 924 
kinabaluense Leenh. 923 
kipella (BI.) Mig. 923, 925, 
927 
kostermansii Leenh. 924 
lamii Leenh. 922, 923 
latistipulatum Ridl. 923 


Leenh. 


Sept. 1972] 


Index to scientific names 


989 


littorale Bl. 922, 923 
f. pruinosum (Engl.) Leenh. 
923 
luzonicum (BI.) A.Gray 922, 
924, 926 
macadamii Leenh. 922, 925* 
maluense Laut. 923 
ssp. maluense 923 
megacarpum Leenh. 923 
megalanthum Merr. 924 
merrillii H.J.Lam 925 
odontophyllum Migq. 924 
oleosum (Lamk) Engl. 924, 
927 
ovatum Engl. 922, 924 
parvum Leenh. 928 
patentinervium Miq. 922, 923, 
927 
perlisanum Leenh. 923 
piloso-sylvestre Leenh. 923 
pilosum Benn. 924, 927, 928 
ssp. borneensis Leenh. 924 
ssp. pilosum 922 
pimela Koen. 927 
pimela Leenh. 923, 927 
polyphyllum K.Sch. 925 
pseudodecumanum Hochr. 
924 
pseudopatentinervium 
H.J.Lam 924 
pseudosumatranum Leenh. 
925 
reniforme Kochummen & 
Whitmore 922, 927 
rigidum (BI.) Mig. 925 
salomonense B.L.Burtt 923 
samoense Engl. 926 
schlechteri Laut. 926 
sinense Cana Rumph. 926 
sinense Tsjacana Rumph. 927 
smithii Leenh. 926 
sumatranum Boerl. & Koord. 
925 
sylvestre Gaertn. 923 
trigonum H.J.Lam 924 
vitiense A.Gray 922, 926 
vrieseanum Engl. 922, 926 
vulgare Leenh. 922, 924 
Cankrienia de Vriese 186 
chrysantha de Vriese 190 
farinosa Zoll. 190 


Cannabinaceae 4: 222-223 


Capparidaceae 61-105 
subfam. Capparidoideae 62 
subfam. Cleomoideae 62 
Capparidales 63 
Capparis Tourn. ex L. 61, 62, 63, 
69, 73 map, 89, 91 
sect. Busbeckea (Endl.) B. & 
H. 69, 70, 92 
sect. Capparis 70 
subsect. Corymbosae DC. 
70 


subsect. Pedicellares DC. 70 
subsect. Seriales DC. 70 


sect. Eucapparis Plum. ex DC. 
70 
sect. Monostichocalyx Radlk. 
70, 86 
acuminata Willd. 84 
acuta (sphalm.) Koord. 92 
affinis Merr. 79 
andamanica King 78 
aurantioides Pres] 87 
baducca (non L.) Blco 90 
billardierii DC. 86 
borneensis Merr. 83 
brachybotrya Hall. f. 71, 73 
f. angustifolia (Hall. f.) Ja- 
cobs 75* 
f. brachybotrya 75 
var. angustifolia Hall. f. 75 
brachyscias Turez. 83 
brevispina DC. 87 
breynia Jacq. 70 
buwaldae Jacobs 71, 85 
callophylla BI. 71, 81, 82 map 
callophylla (non Bl.) Mig. 77 
callosa Bl. 86 
canescens Banks ex DC. 92 
cantoniensis Lour. 70, 71, 76 
map, 77, 81, 93 
carandas Burm. f. 93 
carolinensis Kaneh. 82 
celebica Miq. 76 
cerasifolia A.Gray 83, 84 
copelandii Elm. 77, 78 
cordifolia Lamk 89 
corymbosa Lamk 81 
crassifolia (sphalm.) A.Gray 
84 
crassifolia (sphalm.) C.Muell. 
83, 84 
cucurbitina King 71, 85 
cumingii Merr. & Rolfe 81 
dahlii Gilg & K.Sch. 82 
dasypetala Turcz. 84 
dealbata (non DC.) Back. 83 
dealbata DC. 87 
diffusa Ridl. 71, 81 
discolor (non Donn.Smith) 
Standl. 93 
elliptica Span. 77 
emarginata Pres] 79 
erycibe Hall. f. 70, 71, 74*, 75 
erythrodasys Miq. 87 
finlaysoniana Wall. ex Hook. 
f. & Th. 86 
flexuosa Bl. 86 
floribunda Wight 70, 71, 78 
map 
f. floribunda 78 map 
f. induta Jacobs 70, 78 map 
foetida Bl. 84 
forsteniana Miq. 86 
galeata Fresen. 90 
grandiflora Wall. ex Hook. f. 
& Th. 84 
hasseltiana Miq. 76 
horrida L. f. 87 


var. B erythrodasys Miq. 87 
horrida (non L. f.) Mig. 84 
var. a Miq. 84 
ilocana Merr. 79 
korthalsiana Miq. 86 
kunstleri King 73 
lanceolaris DC. 70, 71, 76, 77 
map, 78, 79, 82 
larutensis King 70, 89 
lasiopoda Turcz. 83 
linearis (non Jacq.) Blco 87 
littoralis Merr. 79 
lobbiana Turcz. 70, 71, 79, 
80* 
loheri Merr. 79 
longestipitata Heine 70, 71, 77 
longipes Merr. 73, 93 
longipes (non Merr.) Stand. 
93 
lucida (Banks ex DC.) Bth. 62, 
70, 92 map 
luzonensis Turcz. 78, 79 
var. ampla Merr. 78, 79 
mariana Jacq. 89 
micracantha (non DC.) Blco 
87 
micracantha DC. 62, 70, 71, 
75, 85, 86, 89 
ssp. korthalsiana (Miq.) Ja- 
cobs 85, 86 map 
ssp. micracantha 85, 86 map 
var. callosa (Bl.) Hall. f. 86 
micrantha (sphalm.) (non A. 
Rich.) Spreng. 85 
mitchellii Lindl. 92 
mucronata Elmer 81 
myrioneura Hall. f. 83, 86 
var. latifolia Hall. f. 86, 87 
nemorosa (non Jacq.) Blco 87 
nigricans Span. 83 
nobilis (non Bth.) F.v.M. 92 
nummularia DC. 91 
oblongata Merr. 77, 78 
octandra Jacq. 78 
odorata Blco 86 
oligostema Hayata 78 
ovalifolia Zipp. ex Miq. 86, 87 
oxyphylla Miq. 84 
palawanensis Merr. 79 
paniculata Ridl. 75, 76 
perakensis (Scort. ex King) 
Rid]. 83 
platyacantha Turcz. 77 
pubiflora DC. 62, 70, 71, 82, 
83*, 84 map, 85, 93 
var. moluccana Miq. 83 
var. perakensis Scort. 83 
var. sumatrana Miq. 83 
pumila Champ. 76 
pyrifolia Lamk 70, 71, 84 
map, 89 
quiniflora DC. 70, 71, 73 map, 
81, 88*, 89 
renominata Jacobs 93 
retusella Thw. 79 


990 


FLORA MALESIANA 


[ser. I, vol. 66 


richii A.Gray 89 
robusta Heine 81 
roxburghii (non DC.) Span. 77 
roydsiaefolia Kurz 86 
rufescens Turcz. 87 
salaccensis Bl. 76, 93 
var. celebica Miq. 76, 77 
sandwichiana DC. 90 
scortechinii King 62, 70, 71 
sepiaria L. 71, 76, 79, 81 map, 
92 
var. acuta Vidal 79 
var. B glabrata DC. 79 
var. grandifolia Kurz & 
Prain 79 
var. retusella Thw. 79 
var. trichopetala Val. 79 
var. vulgaris Hook. f. & Th. 
79 
spinosa L. 62, 70, 89, 91, 92 
var. mariana (Jacq.) K.Sch. 
& Laut. 70, 89, 90*, 91* 
var. nummularia (DC.) F. 
M.Bailey 91 
subacuta Miq. 79, 92 
subcordata Span. 89 
subspinosa Roxb. 77 
tomentosa Lamk 81 
torricellensis Laut. 77 
trapeziflora Span. 89 
trichopetala Val. 79 
trinervia Hook. f. & Th. 70, 
71, 73 map 
turczaninowli Elmer 81 
tylophylla Spreng. 81 
umbellata R.Br. ex DC. 79, 81 
venosa Merr. 86 
versicolor Griff. 93 
viminea (non Hook. f. & Th.) 
Fern.-Vill. 87 
viridis Elmer 77, 78 
zeylanica (non L.) DC. 84 
zeylanica L. 70, 71, 72*, 86, 87 
map 
zeylanica (non L.) Roxb. 87 
zippeliana Miq. 71, 78, 82, 86 
map 
var. novobritannica Laut. 82 
var. novohibernica Laut. 82 
Caprifoliaceae 4: 175-194, 598: 
6: 928-930 
Capura L. 28 
Capusia Lecomte 394 
Carallia Roxb. 965, 966 
borneensis Oliv. 966 
eugenioidea King 966 
longipes Ding Hou 966, 967* 
papuana Ding Hou 967 
suffruticosa Rid]. 966 
Cardiocarpus Reinw. 221 
amarus Reinw. 221 
Cardiophora Bth. 221 
hindsii Bth. 221 
Carices 953 
Carissa carandas L. 93 


grandis Bert. ex Guillemin 335 
Carya 143 
illinoensis 
143 
Caryo juglans Kirchh. 143 
Caryophyllaceae 450 
Caryospermum BI. 230, 288 
alpestre O.K. 290 
arborescens F.v.M. 291 
moluccanum Bl. 291 
Philippinense Vidal 291 
serrulatum Migq. 290 
Casearia grewiaefolia Vent. 944 
var. deglabrata K. & V. 944 
var. gelonioides (BI.) Sleum. 
9 


(Wang) K.Koch 


pallida Craib 944 
velutina BI. 38 
Cassine L. 231, 232, 284, 391, 
392, 930 
subg. Elaeodendron Loes. 285 
sect. Cassine Loes. 285 
sect. Elaeodendron Loes. 
285 
australe (Vent.) O.K. 287 
discolor Wall. in Roxb. 275 
elliptica O.K. 286 
fortunei O.K. 252 
glauca (Rottb.) O.K. 228, 284, 
286 
var. cochinchinensis Pierre 
285*, 286 map 
viburnifolia (Juss.) Ding Hou 
228, 284, 285*, 286 map 
Cassiope 472 
Casuarina 152, 221 
junghuhniana Miq. 153, 187 
Casuarinaceae 143 
Catanthera F.v.M. 914 
Catha Forsk. 229 
edulis Forsk. 230 
fasciculata Tul. 51 
montana G.Don 241 
vitiensis A.Gray 242 
Caulinia Willd. 153, 164 
indica Willd. 164, 166 
ovalis R.Br. 953 
Cedrota guianensis Blco 461 
Celastraceae 49, 227-291, 389- 
421, 930-932 
tribe Hippocrateaceae Hook. 
f. 389 
Celastrales 53, 390, 968 
Celastrineae R.Br. 389 
Celastrus L. 228, 229, 230, 231, 
232, 233, 238, 240, 391, 392, 
930 
sect. Eucelastrus W. & A. 233 
sect. Gymnosporia W. & A. 
238 
subg. Celastrus 234 
ser. Axillares Rehd. & Wils. 
234 
ser. Paniculati 
Wils. 234 


Rehd. & 


alpestris Bl. 290 
apoensis Elmer 236 
australis Harv. & F.v.M. 235 
axillaris Ridl. 236 
bivalvis Jack 276, 277 
championii (non Bth.) King 
236 
dispermus 230 
diversifolius Hemsl. 242 
emarginatus Willd. 241, 242 
franchetiana Loes. 236 
glaucus Vahl 286 
hindsii Bth. 234, 235, 236 
jackianus Steud. 237 
lucida Wall. 237 
lucidus L. 237 
malayensis Ridl. 236 
marianensis Koidz. 236 
micrantha Roxb., nomen 237 
monospermoides Loes. 234, 
235, 236, 237, 243 
monospermus Roxb. 234 
montana = montanus 241 
montanus Roth 241, 242 
montanus Roxb. 242 
multiflorus Roxb., nomen 235 
novoguineensis Merr. & Perry 
233722345255 
nutans Roxb., nomen 235 
obtusifolia Roth 237 
oppositus Wall. in Roxb. 288 
orbiculatus 228 
paniculatus Willd. 229, 231, 
234, 235 
ssp. multiflorus Ding Hou 
235 
ssp. paniculatus Ding Hou 
235 
ssp. serratus Ding Hou 235 
var. balansae Loes. 235 
var. poilanei Tardieu 235 
var. venulosoides Kanj. & 
Das 235 
papuana Warb. 235 
pauciflora Wall. 237 
polybotrys Turcz. 235 
racemosa Turcz. 235, 290 
racemulosa Franch. 236 
racemulosus Hassk. 236 
repandus Bl. 237 
robustus Roxb. 283 
scandens L. 228, 230 
semiarillata Turez. 241 
senegalensis Lam. 242 
stylosus Wall. 234, 235*, 237, 
(sphalm. Willd.) 420 
ssp. glaber Ding Hou 237 
ssp. stylosus Ding Hou 237 
subspicatus Hook. 235 
tonkinensis Pitard 236 
trigyna Roxb. 237, 238 
wallichii G.Don 237 
Centrolepidaceae 5: 421-427 
Centropogon lucyanus Schonl. 
108 


Sept. 1972] 


Index to scientific names 


Centrospermae v.Wettst. 174, 
450, 451 
Centunculus L. 174, 175, 176 
indicus Royle 176 
pentandrus R.Br. 176 
pumilus O.K. 176 
tenellus Duby 176 
Ceodes J. & G.Forst. 457, 459 
brunoniana Skottsb. 461 
corniculata Merr. & Perry 464 
corniculata (non Barg.-Petr.) 
Merr. & Perry 462 
excelsa Skottsb. 461 
longirostris Merr. & Perry 463 
umbellata = umbellifera 460 
umbellifera J. & G.Forst. 460, 
(sphalm. Skottsb.) 461 
urocarpa Merr. & Perry 463 
Cephaelis 336 
Cephalostigma DC. 
12 113 
erectum (Roth) Vatke 113 
hirsuta Edgew. 113 
hookeri Clarke 114 
paniculatum DC. 114 
paniculatum (non DC.) Hos- 
seus 114 
schimperi Hochst. ex Rich. 
113 


LOO 10 1, 


Ceratophyllaceae 4: 41-42 
Ceratopteris thalictroides 
Brongn. 171 
Cerbera 368 
musculiformis Lamk 367, 368 
Cerium Lour. 177 
spicatum Lour. 185 
Cervicina Delile 111 
gracilis Britt. 116 
Chailletia sumatrana Miq. 942 
Chamaedaphne 472 
Cheiloclinium 227, 389 
Cheilotheca Hook. f. 670 
Cheilotheca (non Hook. f.) Prain 
669 
malayana Scort. 669 
Chenopodiaceae 4: 99-106, 594; 
6: 450, 932 
Chenopodium carinatum R.Br. 
932 
pumilio R.Br. 932 
Chilianthus 296 
Cladopus H.MOIl. 963 
nymani H.MOll. 963 
Cleidion spiciflorum (Burm. f.) 
Merr. 203 
Cleisocratera Korth. 387 
Cleistanthopsis Capuron 970 
Cleome L. 61, 62, 63, 99, 100 
sect. Cleomes (DC.) Schult. 
100 
subsect. Pedicellaria (DC.) 
Schult. 100 
subsect. Siliquaria (Forsk.) 
Schult. 100 
sect. Corynandra(e) (Schrad.) 


Schult. 100 

sect. Gymnogonia R.Br. 100 

sect. Gynandropses (DC.) 
Schult. 100 

sect. Polanisiae (DC.) Schult. 
100 


sect. Ranmanissa (Endl.) Gri- 
seb. 100 
sect. Rutidosperma [Jtis 100 
sect. Tarenaya (Raf.) IItis 100 
aculeata L. 100, 104, 105* 
acutifolia Elmer 103 
affinis (non DC.) Spreng. 101 
alliacea Blco 101 
alliodora Blico 101 
aspera Koen. ex DC. 
105* 
blumeana D.Dietr. 101 
blumeana Schult. 101 
chelidonii (non L. f.) Burk. 103 
chelidonii L. f. 62, 100, 102, 
103*, 104 
ciliata Schum. & Thonn. 104 
gigantea (non L.) Blco 101 
gynandra L. 62, 100, 101 
houtteana Schlechtend. 102 
hulletii King 104 
icosandra L. 103 
natalensis 62 
pentaphylla L. 101 
rutidosperma DC. 98*, 100, 
104, 105* 
sandwicensis A.Gray 102 
speciosa Raf. 62, 100, 101 
f. alba 101 
speciosissima Deppe ex Lindl. 
101 
spinosa Jacq. 62, 100, 102 
triphylla L. 101 
viscosa L. 62, 100, 103*, 104 
f. deglabrata (Back.) Jacobs 
104 
f. viscosa 103 
Cnestis palala (Lour.) Merr. 933 
ssp. diffusa (Blco) Andreas 
933 
stenopetala Griff. 933 
steriopetala = stenopetala 933 
Cocculus hirsutus (L.) Diels 939 
villosus DC. 939 
Cochlospermaceae 4: 61-63 
Codonopsis Wall. 107, 109, 110, 
118, 119* 
albiflora Griff. 120 
celebica (Bl.) Miq. 119, 120, 
121 
cordata Hassk. 120 
cordifolia Komarov 120 
javanica (Bl.) Hook. f. 108, 
110, 119, 120* 
lancifolia (Roxb.) Moeliono 
110, 120, 928 
ssp. celebica (BI.) Moeliono 
121 
ssp lancifolia 121, 928 


100, 


99] 


leucocarpa Miq. 120, 121 
parviflora 119* 
purpurea Wall. in Roxb. 119 
truncata Wall. ex DC. 120 
viridis 119 
Coinochlamys 296 
Combretaceae 4: 533-589; 5: 
564; 6: 932-933 
Combretocarpus Hook. f. 966 
rotundatus (Miq.) Danser 966 
Commicarpus Standl. 452 
chinensis Heimerl 455 
plumbaginea 453 
Comocladia serrata Blco 419, 
420 
Compositae 108, 109 
subfam. Cichorieae 108 
Connaraceae 5: 495-541; 6: 
933-936 
Connaropsis rubescens Rid\. 934 
Connarus cochinchinensis 
(Baill.) Pierre 935 
conchocarpus F.v.M. 936 
ssp. conchocarpus 936 
ssp. schumannianus (Gilg) 
Leenh. 936 
culionensis Merr. 935 
var. culionensis 935 
var. stellatus (Merr.) Leenh. 
935 
euphlebius Merr. 935 
ssp. euphlebius 
var. bullatus Leenh. 935 
ssp. moluccanus Leenh. 935 
lamii Leenh. 935 
lucens Schellenb. 935 
monocarpus L. 935 
odoratus Hook. f. 935 
paniculatus Roxb. 935 
var. hainensis (Merr.) Vidal 
935 
var. paniculatus 935 
peekelii Schellenb. 936 
pickeringii A.Gray 936 
salomoniensis Schellenb. 936 
schumannianus Gilg 936 
semidecandrus Jack 935 
villosus Jack 935 
winkleri Schellenb. 
ssp. philippinensis Leenh. 
936 


Conopharyngia 294 
Contortae 294, 297 
Convolvulaceae 4: 388-512, 599: 
5: 558; 6: 390, 936-941 
Convolvulus acuminatus Vahl 
941 
capitatus Vahl 941 
capitiformis Poir. 941 
gangeticus L. 939 
hastatus Desr., non Forsk. 939 
maximus L. f. 941 
ochraceus Lindl. 941 
purpureus L. 941 
Cordia clitoria Bleo 464 


992 


Coris 174 
Cortex filarius Rumph. 11, 12 
foetidus Rumph. 12 
Cortusa 174 
matthioli L. 174 
Corynocarpaceae 4: 262-264; 5: 
557; 6: 941 
Corynocarpus cribbianus (F.M. 
Bailey) L.S.Smith 941 
Costera J.J.S. 469, 470, 471, 473, 
474, 740, 742 map, 746 
borneensis J.J.S. 741, 744 
cyclophylla (Airy Shaw) J.J.S. 
& Airy Shaw 741, 742, 743* 
elegans J.J.S. ex Dunselman 
745 
endertii J.J.S. 741, 745 
lanaensis (Merr.) Airy Shaw 
741, 745 
loheri (Merr.) Airy Shaw 741, 
744* 
lucida (Merr.) Airy Shaw 741, 
742 
ovalifolia J.J.S. 741, 745 
sumatrana J.J.S. 741, 744 
tetramera Sleum. 741, 
744* 
Couthovia A.Gray 296, 363, 365 
alata A.C.Smith 366 
astyla Gilg & Bened. 365, 366 
brachyura Gilg & Bened. 365 
brassii S.Moore 365 
calophylla Gilg & Bened. 367 
celebica Koord. 366, 367 
collina A.C.Smith 365, 366 
corynocarpa A.Gray 365 
densiflora K.Sch. 365 
kochii Val. 366 
leucocarpa Merr. & Perry 365 
macrocarpa A.C.Smith 366 
macroloba A.C.Smith 366 
macrophylla Merr. & Perry 
367 
neo-ebudica Guill. 366 
novo-britannica Kaneh. & Ha- 
tus. 365 
novo-caledonica Gilg & Bened. 
366 
nymanii Gilg & Bened. 365 
pachyantha A.C.Smith 366 
pachypoda Gilg & Bened. 365 
rhynchocarpa Gilg & Bened. 
365, 366 
sarcantha (non Gilg & Bened.) 
Cammerl. 367 
sarcantha Gilg & Bened. 366 
seemanni A.Gray 365 
terminalioides Gilg & Bened. 
365 
toua Kaneh., 367 
undulatifolia Kaneh. & Hatus. 
366 
urophylla Gilg & Bened. 366 
yunzaingensis Merr. & Perry 
365 


742, 


FLORA MALESIANA 


[ser. I, vol. 66 


Covilhamia Korth. 95 
ovata Korth. 97 
Coxia Endl. 177 
Crassulaceae 4: 197-202; 6: 173, 
195 
Crataeva L. 63 
Nirvala Ham. 68 
octandra Blco 78 
religiosa Forst. f. 65 
Crateriphytum Scheff. ex Koord. 
296, 363, 365 
moluccanum Scheff. ex Boerl. 
367, 368 
Crateva L. 61, 63, 69 
A Corner 68 
adansonii DC. 68 
axillaris Presl 66 
B Corner 65 
hansemannii K.Sch. 65 
hygrophila Kurz 65, 69 
lophosperma Kurz 68, 69 
macrocarpa Kurz 65, 66 
magna (Lour.) DC. 68 
marmelos (L.) Correa 69 
membranifolia Miq. 65 
nurvala Ham. 62, 68 
var. nurvala 64*, 65*, 68* 
odora Ham. 66 
f. axillaris (Presl) Jacobs 
65*, 66, 67* 
religiosa (non Forst. f.) Blco 
68 
religiosa (non Forst. f.) Bl. 68 
var. nurvala (Ham.) Hook. 
f. & Th. 68 
religiosa Forst. f. 65*, 66, 67, 
69 
speciosa Volkens 65 
tapia (non L.) Bl. 66 
tumulorum Miq. 66 
unilocularis Ham. 69 
Cressa L. 936 
australis R.Br. 937 
cretica L. 937* 
Crinum asiaticum L. 973 
Cruciferae 62, 63 
Cuervea 390 
Cunoniaceae 49, 933 
Cupania spinosa Blco 241, 242 
Cuscuta L. 
subg. Cuscuta 936 
subg. Grammica (Lour.) Yun- 
cker 936 
Cyathodes Lab. 424, 433 
colensoi (Hook. f.) Hook. f. 
428, 431 
laurina R.Br. ex Drude 437 
Cyclamen 173, 175 
persicum L. 175 
Cyclocodon Griff. 118, 119 
adnatus Griff. 120 
distans Griff. 119 
lancifolium Kurz 120 
truncatum Hook. f. & Th. 120 
Cymothoe Airy Shaw 740 


cyclophylla Airy Shaw 742 
Cynoctonum J.F.Gmel. 293, 295, 
296, 297, 375, 959 
mitreola (L.) Britt. 293, 374*, 
375, 959 
var. intermedia Hochr. 377 
var. lilacina (Back.) Bakh. f. 
375 
var. orthocarpa Hochr. 375 
paniculatum (Wall.) B.L.Rob. 
375 
pedicellatum (Bth.) B.L.Rob. 
377, 960 
petiolatum Gmel. 375 
sphaerocarpum Leenh. 374*, 
375, 377, 960 
Cyrtophyllum Reinw. ex Bl. 299, 
303 
caudatum (Ridl.) Ridl. 307 
fragrans (Roxb.) A.DC, 307 
giganteum (Ridl.) Ridl. 307 
lanceolatum DC. 307, 309 
peregrinum Reinw. ex BI. 307 
speciosum Bl. 303 
var. montanum Ridl. 303 
wallichii (Bth.) Ridl. 307 


Dacryodes Vahl 917 
breviracemosa Kalkman 919 
costata (Benn.) H.J.Lam 918, 

919 
crassipes Kalkman 919 
elmeri H.J.Lam 918 
expansa (Ridl.) H.J.Lam 919 
incurvata (Engl.) H.J.Lam 
918, 919 
kingii (Engl.) Kalkman 917, 
919 
laxa (Benn.) H.J.Lam 918, 919 
longifolia (King) H.J.Lam 919 
var. longifolia 919 
macrocarpa (King) H.J.Lam 
919 
var. kostermansii (Kalk- 
man) Kalkman 919 
var. macrocarpa 919 
nervosa (H.J.Lam) Leenh. 
918, 919 
papuana Husson 919, 921 
puberula (Benn.) H.J.Lam 
918 
rostrata (Bl.) H.J.Lam 918, 
919 
f. pubescens 918 
rubiginosa H.J. Lam 918 
rugosa (BI.) H.J.Lam 918, 919 

Dais (non L.) auct. 15 
coccinia Gaudich. 18, 21 
dubiosa Bl. 20 
dubiosa (non BI.) Decne 17 
laurifolia (non Jacq.) Blco 18 
octandra L. 17, 18 

Dalrympelea Roxb. 51 
pomifera Roxb. 58 

Dalrympelia = Dalrympelea 51, 


Sept. 1972] 


58 
Dalzellia (non Wight) Engl. 963 
Daphne L. 2, 3, 4, 5, 6, 34, 35 
sect. Daphnanthoides Gilg 37 
sect. Eriosolena (BI.) Meisn. 
S18); Big) 
aquilaria Blco 31, 35 
arisanensis Hayata 38 
cannabina (non Lour.) Schau- 
er 33 
cannabina (non Lour.) Wall. 
37 
composita (L. f.) Gilg 2, 36*, 
37, 38 
var. montana Hochr. 37 
f. macrophylla Hochr. 37 
decandra Bl. 38 
foetida (non L.) Blco 32 
indica (non L.) Blco 32 
indica L. 35 
javanica Thunb. 38 
kiusiana Miq. 38 
luzonica C.B.Rob. 2, 36*, 37, 
38 
montana Meisn. 38 
odora Thunb. 38 
papyracea Wall. ex Steud. 37 
pendula Sm. 38 
var. y concolor Meisn. 38 
var. B montana Meisn. 38 
Phaetida 32 
sp. Steen 35 
Daphnobryon Meisn. 43 
ericoides Meisn. 44 
Datiscaceae 4: 382-387 
Datura 253 
Daucus glochidiatus 
Fischer 984 
Decaisnella O.K. 6 
cumingiana O.K. 15 
Decaspora laurina O.K. 437 
Decatoca F.v.M. 423, 424, 434, 
436 map 
spenceri F.v.M. 435* 
Deltaria 3 
Denhamia Meisn. 240, 243 
parvifolia L.S.Smith 240 
pittosporoides F.v.M. 230, 
240 
Dentella 253 
erecta Roth ex R. & S. 113 
perotifolia Willd. ex R. & S. 
tS 
Desfontainea Kunth 295, 297, 
954 
Dichapetalaceae 5: 
567; 6: 941-943 
Dichapetalum Thou. 238, 941 
gelonioides (Roxb.) Engl. 941, 
942 


(Labill.) 


305-316, 


ssp. pilosum Leenh. 942 
ssp. tuberculatum Leenh. 
9 


glabrum (Vahl) Prance 942 
grandifolium Ridl. 941, 942 


Index to scientific names 


griffithii (Hook. f.) Engl. 942 
hainanense Eng}. 942 
helferianum (Kurz) 
941, 942 
laurocerasus (Hook. f.) Engl. 
942 
longipetalum (Turcz.) Engl. 
942 
papuanum (Becc.) Boerl. 941, 
942 
ssp. borneense Leenh. 942 
ssp. papuanum 942 
peekelii Krause 941, 942 
scorpioideum Leenh. 941, 943 
sessiliflorum Leenh. 942 
setosum Leenh. 942 
steenisii Leenh. 942 
ssp. steenisii 942 
tenerum Leenh. 943 
timoriense (DC.) Boer]. 941, 
942 
tricapsulare (Blco) Merr. 942 
Diervilla fallax (Miq.) Boerl. 
929 
Dillenia 49 
Dilleniaceae 4: 141-174; 5: 557; 
6: 933 
Dimorphanthera F.v.M. 469, 
470, 471, 473, 474, 746, 885, 
886 map 
sect. Brachychone Schltr 890 
sect. Cyclosiphon Wernh. 890 
sect. Dimorphanthera Sleum. 
886, 890 
subsect. Brachychone 
(Schltr) Sleum. 890 
sect. Pteridosiphon Wernh. 
886, 889 
sect. Trochilanthe Schltr 887, 
896 
subsect. Trochilanthe 
(Schltr) Sleum. 896 
alba J.J.S. 904 
albiflora Schltr 887, 892 
alpina J.J.S. 888, 905 
var. alpina 905 
var. pubigera Sleum. 905 
alpivaga Sleum. 886, 889, 912 
amblyornidis (Becc.) F.v.M. 
892 
var. amblyornidis 887, 892, 
893*, 896 
var. moorhousiana (F.v.M.) 
Sleum. 887, 893, 894* 
amoena Sleum. 886, 887, 896 
anchorifera J.J.S. 887, 888, 
902 
apoana (Merr.) Schltr 887, 
895 
arfakensis J.J.S. 892 
beccariana (Koord.) J.J.S. 
888, 904 
brachyantha Sleum. 887, 899 
brassii Sleum. 887, 888, 897, 
898* 


Pierre 


993 


breviflos Sleum. 886, 891 
brevipes Schltr 887, 888, 900 
calodon Sleum. 887, 889, 910 
chlorocarpa Sleum. 895 
clemensiae Sleum. 888, 902, 
903 * 
collinsii Sleum. 908 
var. collinsii 888, 908 
var. montis-wilhelmi Sleum. 
888, 908, 909* 
cornuta J.J.S. 887, 899 
var. cornuta 899 
var. tenuiflora Sleum. 899 
crassifolia Sleum. 888, 901 
darmandvillei J.J.S. 914 
declinata Sleum. 886, 890 
dekockii J.J.S. 887, 893 
var. chlorocarpa (Sleum.) 
Sleum. 894, 895 
var. dekockii 893, 894 
var. pubiflora Sleum. 887, 
894, 895 
denticulifera Sleum. 905 
var. denticulifera 888, 889, 
905 
var. pubens Sleum. 888, 905 
dielsiana J.J.S. 891 
doctorsii J.J.S. 887, 888, 906 
dryophila Sleum. 889, 914 
var. dryophila 914 
var. trichoclada Sleum. 914 
elegantissima K.Sch. 888, 908 
eymae Sleum. 887, 898 
forbesii (F.v.M.) F.v.M. 888, 
904 
forbesii [non (F.v.M.) F.v.M.] 
Warb. 908 
gracilis Sleum. 887, 900 
hirsutiflora Sleum. 887, 897 
intermedia (non J.J.S.) Diels 
891 
intermedia J.J.S. 886, 892 
kalkmanii Sleum. 888, 908 
kaniensis Schltr 908 
kempteriana Schltr 886, 891 
lancifolia Sleum. 889, 910 
latifolia Schltr 887, 897 
leucostoma Sleum. 888, 889, 
905 
longifolia Kaneh. & Hatus. 
888, 907 
macleaniaefolia = macleanii- 
folia 889 
macleaniifolia Wernh. 886, 
889 
magnifica Sleum. 886, 891 
megacalyx Sleum. 889, 910 
meliphagidium = meliphagi- 
dum 906 
meliphagidum (Becc.) F.v.M. 
888, 906 
microphylla Sleum. 889, 913 
militaris J.J.S. 889, 910 
mindanaensis Merr. 887, 892 
moluccana J.J.S. 896 


994 


FLORA MALESIANA 


[ser. I, vol. 66 


moorhousiana F.v.M, 893 
moorhousiana (non F.v.M. s. 
str.) Schltr 892 
myzomelae (Becc.) J.J.S. 889, 
913 
nigropunctata Sleum. 888, 901 
obovata J.J.S. 736, 911, 912 
obtusifolia Sleum. 889, 913 
ovatifolia Sleum. 888, 907 
parviflora J.J.S. 889, 912 
parvifolia J.J.S. 889, 912 
peekelii Sleum. 888, 907 
prainiana (Koord.) J.J.S. 889, 
911* 
profusa J.J.S. 891 
pulchra J.J.S. 887, 896 
racemosa Schltr 887, 898 
robbinsii Sleum. 888, 902 
splendens Sleum. 888, 904 
steinii Sleum. 887, 895 
thibaudifolia Sleum. 888, 901 
torricellensis Schltr 887, 899 
tridens J.J.S. 886, 890 
umbellata Wernh. 886, 890 
vaccinioides Sleum. 889, 912 
velutina Schltr 887, 896 
vestita Sleum. 887, 900 
vonroemeri (Koord.) J.J.S. 
889, 911 
wollastonii Wernh. 887, 898 
womersleyi Sleum. 888, 906 
wrightiana (Koord.) J.J.S. 
889, 914 
Dioscoreaceae 4: 293-335; 5: 
557 
Diosma serrata Blco 235 
Diospyros hierniana (K. & G.) 
Bakh. 932 
Diotocranus 297 
Diplomorpha Meisn. 28 
Diplusodon Pohl 975 
Diplycosia Bl. 469, 471, 473, 
474, 677, 696 map, 912 
abscondita Sleum. 697, 703 
acuminata Becc. 700, 722, 
132% 
amboinensis Becc. 699, 716, 
132% 
aperta J.J.S. 697, 706 
apiculifera J.J.S. 701, 731 
apoensis Elm. 699, 716 
atjehensis Sleum. 697, 701, 
707 
aurea Sleum. 697, 704 
haclay(a)ensis Elm. 740 
barbigera Sleum. 697, 703 
bartlettii Merr. 738 
brachyantha Sleum. 698, 699, 
701, 713 
var. brachyantha 713 
var. parvula Sleum. 713 
breviflora Ridl. 676 
calelanensis Elm. 720 
capitata Sleum. 699, 702, 721 
carrii Sleum. 697, 702 


caryophylloides J.J.S. 699, 
721 
var. caryophylloides 721 
var. longipes Sleum. 721 
caudatifolia Sleum. 700, 729 
celebensis J.J.S. 700, 730 
chrysothrix Stapf 697, 702 
ciliolata Hook. f. 698, 708, 
(Bis 
cinnabarina Sleum. 700, 724, 
(32% 
cinnamomifolia Stapf 702, 
735*, 736 
clementium Sleum. 697, 704, 
105% 
commutata Sleum. 701, 734 
consobrina Becc. 700, 724 
cordifolia Ridl. 723 
coriifolia Sleuam. 734 
crassiramea Sleum. 699, 701, 
719 
crenulata Sleum. 700, 723 
edulis Schltr 699, 701, 735 
elliptica Rid]. 700, 723, 732*, 
778 
endertii J.J.S. 740 
ensifolia Merr. 702, 735*, 737 
erythrina (Hook.) K. & G. 815 
fasciculiflora Merr. 720 
filipes Sleum. 698, 708, 732* 
fimbriata Sleum. 699, 722 
glabra Merr. 720 
glauciflora Sleum. 698. 712 
gracilipes J.J.S. 700, 730 
haemantha Sleum. 700, 730 
heterophylla Bl. 702, 738 
var. heterophylla 738 
var. latifolia (Bl.) Sleum. 
701, 738, 739*, 740* 
var. obovata J.J.S. 738 
heterophylla (non BI. s. str.) 
K. & G. 740 
hirsuta Sleum. 698, 709 
kalmiifolia Sleum. 699, 717 
kemulensis J.J.S. 701, 731 
kerintjensis J.J.S. 740 
var. elliptica J.J.S. 740 
kinabaluensis Stapf 699, 701, 
MAT 35% 
kingii Merr. 723 
kjellbergii J.J.S. 698, 709 
kosteri Sleum. 698, 710 
kostermansii Sleum. 700, 724 
lamii J.J.S. 699, 719 
lancifolia Ridl. 700, 728 
var. calvescens Sleum. 729 
var. lancifolia 728 
latifolia Bl. 738 
latifolia (non B.) Hall. f. 738 
lavandulifolia Sleum. 700, 727 
ledermannii Schltr 698, 700, 
713 
lilianae J.J.S. 697, 706 
loheri Merr. 697, 706 
lorentzii Koord. 701, 736 


lucida Merr. 742 
luzonica (A.Gray) Merr. 699, 
701, 720 
var. calelanensis 
Sleum. 720 
var. luzonica 720 
var. merrittii (Merr.) Sleum. 
720, 721 
var. pubens Sleum.720 
lysolepis Sleum. 701, 734 
macrophylla Becc. 740 
memecyloides (non 
Merr. 717 
memecyloides Stapf 700, 729 
merrittii Merr. 721 
microphylla Becc. 700, 723, 
(32 
microphylla (non Becc.) Clar- 
ke 723 
minutiflora Sleum. 701, 731 
morobeensis Sleum. 698, 713 
var. morobeensis Sleum. 
698, 713 
var. ovatifolia Sleum. 714 
mundula (F.v.M.) Schltr 684 
muscicola Sleum. 699, 716 
myrtillus Stapf 700, 728 
opaca C.B.Robins. 721 
orophila Sleum. 699, 702, 720 
parvifolia Merr. 698, 711 
pendens Sleum. 698, 699, 714 
penduliflora Stapf 697, 701, 
707, 735* 
piceifolia Sleum. 700, 726 
pilosa Bl. 700, 729 
pinifolia Stapf 700, 725*, 
726", Que 
pittosporifolia J.J.S. 732 
var. pittosporifolia 701, 732 
var. punctiloba Sleum. 699, 
733 
pokapindjangensis J.J.S. 706 
pseudorufescens Sleum. 700, 
711 
var. elliptifolia Sleum. 698, 
700, 711 
var. pseudorufescens 698, 
711 
pubivertex Sleum. 697, 707 
punctulata Stapf 701, 736 
retusa Sleum. 701, 733 
rosea Sleum. 698, 709, 710* 
rosmarinifolia Sleum. 701, 
Wil Bye 
rubella Sleum. 700, 728 
rubidiflora J.J.S. 701, 733 
rufa (non Stapf) Merr. 704 
rufa Stapf 697, 702 
rufescens Schltr 698, 708 
rupicola Sleum. 698, 714, 732* 
sagittanthera J.J.S. 700, 722 
salicifolia Sleum. 702, 737 
sanguinolenta Sleum. 701, 733 
saurauioides J.J.S. 697, 704 
scabrida Becc. 699, 721 


(Elm.) 


Stapf) 


Sept. 1972] 


scandens Merr. 720 
schramii Sleum. 697, 703 
schultzei Schltr 699, 714 
setiloba Sleum. 700, 724 
setosa J.J.S. 698, 710 
soror Becc. 698, 699, 712 
var. nuda Sleum. 701, 712 
var. soror 712 
sphenophylla Sleum. 
132 e133 
stellaris Sleum. 699, 715 
stenophylla Sleum. 700, 727, 
132* 
subglobularis Sleum. 699, 715 
sumatrensis Merr. 699, 718* 
tetramera Sleum. 697, 698, 
TA lat 
triangulanthera J.J.S. 698, 708 
trinervia Elm. 699, 719 
var. trinervia 719 
var. urdanetensis 
Sleum. 719 
undata (non J.J.S.) H.J.Lam 
706 
undata J.J.S. 699, 715 
urceolata (non Stapf) Ridl. 
var. 740 
urceolata (non Stapf) Sleum. 
734 
urceolata Stapf 701, 734, 735* 
urdanetensis Elm. 719 
varians Sleum. 699, 715 
viridiflora Sleum. 702, 737 
var. megalantha Sleum. 738 
var. viridiflora 737 
Dipsacaceae 4: 290-292; 5: 557 
Dipterocarpaceae 154 
Disiphon Schltr 785 
papuanum Schltr 786 
Distyliopsis Endress 952 
Distylium Sieb. & Zucc. 952 
Dodecatheon 174 
Dodonaea 152 
Dolianthus C.H.Wright 387 
Dortmanna Adans. 121 
Dortmannia (sphalm.) O.K. = 
Dortmanna Adans. 121 
alsinoides O.K. 127 
campanuloides O.K. 131 
chinensis O.K. 131 
colorata O.K. 124 
erecta O.K. 124 
griffithii O.K. 127 
haenkeana O.K. 136 
inconspicua O.K. 129 
leschenaultiana O.K. 123 
nicotianaefolia O.K. 124 
pyramidalis O.K. 124 
radicans O.K. 131 
reinwardtiana O.K. 129 
rosea O.K. 124 
subcuneata O.K. 128 
succulenta O.K. 128 
trialata O.K. 129 
trigona O.K. 127 


701, 


(Elm.) 


Index to scientific names 


var. affinis O.K. 128 
var. microcarpa O.K. 127 
var. terminalis O.K. 127 
zeylanica O.K. 128, 129 
Dovyalis macrodendron Gilg 944 
Drapetes Banks ex Lamk 2, 4, 5, 
6, 43 
sect. Daphnobryon Boer). 43 
dieffenbachii Hook. 43 
ericoides Hook. f. 2, 43, 44* 
map 
muscosus Lamk 43 
tasmanicus Hook. f. 43 
Drimyspermum Reinw. 15, 16 
ambiguum Meisn. 17 
blumei Decne 20 
blumei (non Decne) Hassk. 17 
burmanni Decne 17, 18 
cauliflorum Thw. 20 
coccineum Becc. 18, 21 
cumingti Meisn. 21 
laurifolium Decne 17 
laurifolium (non Decne) 
Hassk. 20 
longifolium Miq. 17 
macrocarpum Scheff. 22, 23 
perrottetianum Decne 18 
phaleria Meisn. 20 
revolutum T. & B. 21 
urens Reinw. 20 
urens (non Reinw.) Scheff. 18 
Drosera petiolaris R.Br. 943 
spathulata Labill. 943 
umbellata Lour. 191, 192 
Droseraceae 4: 377-381; 5: 557; 
6: 943 
Drymispermum = 
mum 15, 16 
Duabanga Buch.-Ham. 975* 
grandiflora (Roxb. ex DC.) 
Walp. 975*, 976 
moluccana BI. 975* 
< taylorii Jayaweera 975* 
Durio 70 


Drimysper- 


Dysoxylum dasyphyllum Mia. 
218 
quercifolium (Seem.) A.C. 


Smith 212 


Ebelingia Rchb. 297 
brownii Steud. 2)7 
paucijuga O.K. 208 
Edgeworthia 3 
Elaeocarpaceae 420, 914 
Elaeocarpus oxypyren K. & V. 
420 
Elaeodendron Jacq. f. ex Jacq. 
228, 284, 285 
australe Vent. 287 
ellipticum Decne 286, 287 
var. glauca Pierre 287 
var. montana (Thw.) Pierre 
287 
var. roxburghii (W. & A.) 
Pierre 287 


995 


fortunei Turez. 252, 253 
glaucum (Rottb.) Pers. 286 
var. macrocarpa K. & V. 
286 
horizontale Turcz. 241 
Javanicum Turez. 252, 253 
microcarpum White & Francis 
288 
mindanaense Merr. 288 
pauciflorum Tulasne 287 
roxburghii W. & A. 286, 287 
subrotundum King 228, 286 
viburnifolium Merr. 286 
Elaeodendrum = Elaeodendron 
284 
Elatinaceae 4: 203-206 
Eleocharis 165 
Ellipanthus beccarii Pierre 935 
tomentosus Kurz 935 
ssp. tomentosus 935 
var. tomentosus 935 
Emorya Torr. 295, 296, 953 
Enchysia Presl 140 
Engelhardia Lechen. ex BI. 143, 
145, 146 map, 147, 148 


sect. Engelhardia 145, 147, 
148 

sect. Oreomunnea_ (Oerst.) 
C.DC. 148 

sect. Psilocarpeae Nagel 
emend. Leroy 145, 148 

sect. Prerilema  (Reinw.) 
C.DC. 148 

sect. Trichotocarpeae Nagel 
148 


aceriflora Bl. 152 
acerifolia (sphalm.) Hook. f. 
152 
apoensis Elm. ex Nagel 148, 
151 
chrysolepis Hance 154 
colebrookeana Lindl. ex Wall. 
153 
esquirolii Lév. 153 
fenzelti Merr. 154 
formosana Hayata 143, 154 
lepidota Schltr 148 
mollis Hu 148 
nucifera (Ludwig) Madler 147 
nudiflora Hook. f. 150 
var. crenata Hook. f. ex 
Gamble 150 
palembanica Miq. 150 
parvifolia C.DC. 146, 150 
permicrophylla Elm. 150 
philippinensis C.DC. 152 
polystachya Radlk. 143, 154 
pterococca O.K. 154 
var. a roxburghiana O.K. 
154 
rigida BI. 144*, 145, 148, 149* 
roxburghiana Lindl. ex Wall. 
152 
roxburghiana Wall. 148, 149*, 
154 


996 


FLORA MALESIANA 


[ser. I, vol. 6® 


selanica Bl. 154 
serrata Bl. 144*, 146, 148, 149, 
150* 
sp. Corner 151 
spicata Lechen. ex BI. 143, 
144*, 145, 146, 147*, 148, 
151%) 152*) 153815451953 
var. a genuina K. & V. 151 
var. B aceriflora K. & V. 
Syl 
var. y colebrookiana K. & 
V. 151, 154 
var. colebrookeana (Lindl. 
ex Wall.) O.K. 151, 153, 
953 
var. formosana Hayata 154 
var. integra (Kurz) Man- 
ning 953 
var. spicata 153 
subsimplicifolia Merr. 148 
villosa Kurz 153 
var. integra Kurz 153, 953 
wallichiana (Lindl. ex Wall.) 
C.DC. 154 
B chrysolepis C.DC. 154 
zambalensis Elm. 148, 149 
Engelhardtia = Engelhardia 145 
Englerodaphne Gilg 79 
Enhalus acoroides (L. f.) Royle 
952 
Enkleia Griff. 2, 4, 5, 6, 23, 982 
coriacea Hall. f. 25 
malaccensis Griff. 24*, 25, 982 
malayana Griff. 25 
paniculata (Merr.) Hall. f.24*, 
25 
riouwensis Hall. f. 25 
zippeliana Hall. f. 25, 26 
Epacridaceae 50, 422 map-444, 
943 
subfam. Epacrideae 423 
tribe Epacrideae 423 
subfam. Styphelieae 423 
tribe Styphelieae 423 
Epacris 423 
Epigynium coriaceum (Bl.) KI. 
828 
ellipticum (Bl.) K1. 873 
floribundum (Bl.) K1. 872 
laurifolium (Bl.) K1. 872 
lucidum (Bl.) KI. 814 
varingiaefolium (Bl.) Kl. 815 
Ericaceae 49, 423, 424, 469-914, 
943 
subfam. Arbutoideae 473, 474 
tribe Andromedeae 474 
tribe Gaultherieae 474 
tribe Pyroleae 474 
subfam. Ericoideae 473 
subfam. Monotropoideae 469, 
473 
subfam. Rhododendroideae 
473 
tribe Rhododendreae 474 


subfam. Vaccinoideae 472, 
474 
Ericales 473, 669 
Eriosolena Bl. 35 
affinis Zoll. 38 
composita v.Tiegh. 38 
montana Bl. 38 
var. @ macrophylla Hassk. 
38 
var. & minor Hassk. 38 
pendula Bl. ex Lecomte 38 
viridiflora Zoll. & Mor. 33 
Erodium L’Heérit. 445, 448 
cicutarium (L.) L’Heérit. ex 
Ait. 445, 448 
Ervatamia sphaerocarpa 
Burk. 347 
Erycibe 390 
Erythroxylaceae 5: 543-552 
Eucommiaceae 229 
Euonymus Tournef. ex L. 227, 
D282 955 OMS eRe S252 43. 
245, 253, 254, 256, 264, 274, 
391, 392, 930 
subg. Euonymus 247 
subg. Kalonymus Beck 247 
acuminifolius Blakelock 247, 
251 
var. borneensis 
251 
alatus {non (Thunb.) Sieb.] 
Elm. 249 
alatus Sieb. 230 
atropurpureus Jacq. 229, 230 
bancanus Miq. 249 
benguetensis Merr. 248, 253 
castaneifolius Ridl. 246*, 247, 
251 
cochinchinensis 
248, 286, 930 
congolensis Wilczek 245 
coriaceus Ridl. 249 
decora W.W.Sm. 245 
elmeri Merr. 249 
europaeus L. 229, 230 
f. genuinus Back. 248 
fengii Chun & How 256 
fimbriatus Baill. ex Laness. 
267 
fortunei (Turez.) Hand.-Mazz. 
DID e253 
var. radicans (Miq.) Rehd. 
DD, 
fungosus Ohwi 252, 253 
glaber Roxb. 247, 250 
glandulosus (Merr.) Ding Hou 
248, 251 
grandiflorus Wall. 264 
horsfieldii Turcz. 249 
impressus Blakelock 247, 250 
japonicus (Thunb.) L. f. 252, 
253 
japonicus Thunb. 246*, 248, 
252 map 
var. radicans Miq. 252 


(Bl.) 


Blakelock 


Pierre) 247, 


javanicus Bl. 231, 245, 247, 
248, 250 
f. genuinus 248 
var. conocarpus Hassk, 249 
var. coriaceus 248 
var. elmeri 248 
var. genuina K. & V. 248 
var. genuinus K. & V. 248 
var. horsfieldii Turcz. 248 
var. sphaerocarpus Hassk. 
248, 249 
var. talungensis Pierre 248 
var. timorensis 248 
latifolia (L.) Mill. 245, 246 
lichiangensis W.W.Sm. 245 
linearifolia Franch. 245 
marivelensis Elm. 258 
micropetalus Ridl. 249 
moluccensis Blakelock 248, 
254 
moultoni Ridl. 251 
nana M.B. 243 
oliganthus Merr. 248 
omeiensis Fang 245 
pahangensis Ridl. 248 
philippinensis Merr. 248 
platyclinis Ohwi 243 
radicans (Miq.) Sieb. ex Miq. 
25D) 
recurvans Miq. 247, 250 
rufulus Rid]. 250 
sumatranus Miq. 248 
timorensis (non Zipp. ex 
Span.) Turez. 248 
timorensis Zipp. ex Span. 248, 
249 
tonkinensis Loes. 247 
viburnifolius Merr. 248, 286 
wrayi King 247, 250, 258 
yunnanensis Franch. 245 
Eupatorium inulifolium 145 
Euphorbiaceae 4, 203, 420, 944 
Eurycoma Jack 193, 196, 203 
apiculata Benn. 205, 206 
dubia Elmer 206 
eglandulosa Merr. 206 
harmandiana Pierre 205, 206, 
969 
latifolia (sphalm.) Rid]. 205 


longifolia Jack 204*, 205, 
206* map 
ssp. eglandulosa (Merr.) 


Nooteboom 206 map 
ssp. longifolia 206 map 
var. cochinchinensis Pierre 

205 
var. merguensis Pierre 205 

merguensis Planch. 205 
tavoayana Wall. 205 
Euscaphis Sieb. & Zucc. 49, 51 
Evodia 226 
meliaefolia (Hance) Bth. 206 
parviflora Craib 56 
triphylla DC. 214 
Evolvulus L. 936 


Sept. 1972] 


Index to scientific names 


997 


gangeticus (L.) L. 939 

sp. Ridsdale 960 
Evonymus Auct. 245 
Exobasidium 172, 471, 659, 747 


Fagaceae 143, 145, 472 
Fagaceae-Castaneae 143 
Fagara 207, 209 
piperita (non L.) Naves 208 
Fagraea Thunb. 293, 294, 295, 
297, 298*, 299, 301*, 303 map, 
312, 318 
sect. Corymbosae Bth. 315 
sect. Cyrtophyllum (Reinw.) 
BI. 300, 303, 309 
sect. Eufagraea Bl. 315 
sect. Fagraea 300, 315 
sect. Fagraeae verae Bl. 315 
sect. Parviflorae Bth. 303 
sect. Pseudocorymbosae Soler. 
315 
sect. Pseudoracemosae Soler. 
311 
sect. Racemosae Bth. 300, 309 
subg. Cyrtophyllum (Bl.) Miq. 
303 
subg. Eufagraea Miq. 315 
acuminatissima Merr. 302, 
319, 320, 954 
affinis S.Moore 335 
alteniana F.v.M., nomen 335 
amabilis S.Moore 332 
amboinensis Bl. 316, 318 
ampla S.Moore 333 
angiensis Kaneh. & Hatus. 
Siveslseslo* 
annulata Hiern 300, 302, 303, 
318, 333, 334, 335 
anthocleistifolia Gilg & Bened. 
326 
appendiculata Bl. 311 
archboldiana Merr. & Perry 
317, 318 
aurantiodora §.Moore 304 
auricularia = auriculata 327 
auriculata Jack 300, 301*, 
302, 326, 329*, 954 
ssp. auriculata 328, 954 
ssp. borneensis (Scheff.) 
Leenh. 328, 329, 954 
ssp. parviflora Leenh. 303, 
328 
berteriana A.Gray 301, 302, 
S116. 3175 31.9, 331,,333;335, 
954 
birmanica Gandoger 317 
blumei G.Don 293, 299, 300, 
SOI 302, 315; 3116; 317, 318, 
320, 322*, 324, 325, 954 
ssp. blumei 323, 324, 954 
ssp. plumeriaeflora (DC.) 
Leenh. 323, 325, 326 
blumei Steud. 320 
blumii = blumei 329 
bodenii Wernh. 302, 333, 335, 


336 
borneensis Scheff. 327, 328 
bracteosa Cammerl. 328 
calcarea M.R.Hend. 303, 332 
calophylloides Gilg & Bened. 
335 
cambagei Domin 300, 332, 333 
cardinalicarpa Elm. 336 
carnosa Jack 302, 316, 331 
carstensensis Wernh. 302, 326, 
327* 
caudata Ridl. 307, 309 
ceilanica Thunb. 301, 302, 
BOS NS1SeS16s) 31828319. 
320, 323, 954 
f. khasiana 303 
var. brevituba Clarke 315 
celebica Bl. 316, 318 
chinensis Merr. 317 
coarctata Bl. 311 
var. ligustrina (Bl.) Miq. 311 
cochinchinensis (Lour.) 
A.Chev. 307, 309 
congesta Bl. 316 
congestiflora Elm. 311 
cordifolia Bl. 311 
coromandelina Wight 316 
crassifolia Bl. 316 
crassifolia Wall., non Bl. 320 
crassipes Bth. 311 
crenulata Maing. ex C.B.Clar- 
ke 293, 299, 300, 301, 324* 
cuernosensis Elm. 322, 323 
currant Merr. 327, 328, 329 
curtisii K. & G. 303, 331, 332 
cuspidata Bl. 311 
cymosa Merr. 311 
dasyantha Gilg & Bened. 335 
dolichopoda Gilg & Bened. 
332) 
dubia Wall. 336 
elata Merr. & Perry 300, 332 
elliptica Roxb. 293, 300, 301, 
303, 304*, 309, 315, 954 
epiphytica Elm. 327, 328 
eucalyptifolia Cammerl. 311, 
SSS 15 
euneura Scheff. 327, 328 
eymae Back. 301*, 303, 333 
fagraeacea Druce 332 
fastigiata Bl. 301*, 323, 324, 
325, 954 
fastigiata (non Bl.) Cammerl. 
325, 328 
fastigiata (non BI.) Ridl. 324 
flavidula Ridl. 331 
forstenii Bl. 316, 318 
forstenii (non Bl.) Koord. 325 
fragrans Roxb. 293, 300, 301, 
303, 304, 305*, 306*, 307*, 
308*, 309, 323, 954 
fuscescens Bl. 316 
galilai Gilg & Bened. 335 
gardeniaeflora Wernh. 317, 
318 


gardenioides Ridl. 300, 302, 
331, 332 
ssp. borneensis Leenh. 332 
ssp. gardenioides 331 

garneri Thw. 317 

gigantea Ridl. 307, 309 

gitingensis Elm. 302, 317, 334, 
335 

gracilipes A.Gray 300, 301, 
303, 309, 332, 334, 335, 954 

gracilis Cammerl. 311 

grandifolia Merr. 311 

grandis Panch. & Sébert 335 

heterophylla Bl. 316 

imperialis Miq. 327, 328, 329* 

intermedia Ridl. 322, 323 

involucrata Merr. 301, 328, 
3297755054 sole oor 
var. Jlongipetiolata 

329, 330 

Jackii Elm. 327, 328 

jJasminodora Gilg & Bened. 
333 

Javanensis Bakh. f. 304 

javanica (Reinw. ex) Bl. 327, 
328 

khasiana Bth. 316 

kimanga = kimangu 303 

kimangu Bl. 303 

ksid Gilg & Bened. 335 

kusaiana Hosok. 335 

lanceolata Bl. 316 

lanceolata (non Bl.) Hend. 331 

lanceolata Schnizl. 309 

lanceolata Wall., non Bl. 307 

latifolia Mig. 311 

ledermannii Gilg & Bened. 
335 

leschenaultii B\. 320, 323 

ligustrina Bl. 311, 312*, 315 
var. brachystachya Bth. 311 
var. disparifolia Bl. 311 

litoralis Bl. 316, 317, 318, 323, 
954 
var. amboinensis Bl. 316 
var. forstenii (Bl.) Mig. 316 
var. moluccana Miq,. 316 

littoralis = litoralis 316 

loheri Merr. 317, 318 

longicuspis Gandoger 319 

longiflora Merr. 302, 326 

lutea Cammerl. 317, 318 

lutea (non Cammerl.) Merr. 
322.1323 

macgregorii Merr. 326 

macrodendron Gilg & Bened. 
31 

macroscypha Baker 302, 328, 
329*, 330, 331 

macroscypha (non Baker) Hei- 
ne 329 

maingayi C.B.Clarke 311, 315 

malabarica Bl. 316 

malabarica Wight, non Bl. 316 

malayana Mart. 311, 315 


Merr. 


998 


melanochlora Gilg & Bened. 
333, 335 

minor (non BI.) Heine 311 

minor Reinw. ex BI. 316, 318 

monantha Migq. 331 


336 


morindaefolia (Reinw.) BI. 
293, 311 
var. robusta BI. 311 
morindifolia = morindaefolia 
311 


muelleri Bth. 300, 332, 333 
negrosensis Elm. 322, 323 
nonok Elm. 327, 328 
novae-guineae Cammerl. 335 
nov. sp. aff. longiflora 325 
oblonga K. & G. 317, 318, 954 
obovata Bl., non Wall. 320, 
323, 324 
obovata (non Wall.) King 319, 
320, 323 
obovata (non Wall.) Miq. 321, 
323 
var. brevicalyx Bakh. f. 322, 
323 
var. latifolia Mig. 321, 323 
obovata Wall. 315, 317 
var. brevicalyx Bakh. f. 316 
var. gardneri Clarke 315 
var. latifolia Miq. 315 
var. papuana F.M.Bailey 
316, 335 
obovato-javana Bl. 293, 320, 
323 
var. bebeak Bl. 320, 323 
var. latifolia Mig. 320, 323 
obtusifolia Merr. & Perry 332 
oxyphylla Mig. 316, 318 
pachyclados K.Sch. 336 
pachypoda Gilg & Bened. 335 
papuana Merr. & Perry 333 
pauciflora (K. & G.) Ridl. 311, 
315 
peekelii Gilg & Bened. 335 
pendula Merr. 310*, 311, 315 
peregrina (Reinw.) Bl. 307 
picrophloea Bl. 303 
plumeriaeflora DC. 320, 323 
plumeriaeflora (non DC.) 
Koord. 325 
plumeriaefolia = plumeriaeflo- 
ra 320, 325 
pluvialis S.Moore 335 
prainii Gandoger 317 
pseudoelliptica Kaneh. & Ha- 
tus. 304 
pusilliflora Bakh. f. 304 
pyriformis 8.Moore 317, 318 
racemosa Jack ex Wall. 293, 
S005. 302, 3107 Siiee3i12=, 
313*, 314*, 316, 954 
var. coarctata (Bl.) Miq. 311 
var. grandis Wall. ex A.DC. 
311 


FLORA MALESIANA 


[ser. I, vol. 6& 


var. pauciflora K. & G. 311 

rahmatii Merr. 317 

resinosa Leenh. 302, 331 

ridleyi Gandoger (non K. & 
G.) 307 

ridleyi K. & G. 303, 316, 320, 
321*, 954 

robusta Bl. 311 

rodatzii K.Sch. & Laut. 311 

rosenstromii C.T.White 335 

rostrata Bl. 316 

rotundifolia Ridl. 331 

sair Gilg & Bened. 335 

salomonensis Gilg & Bened. 
335 

salticola Leenh. 
834453354550 

samoensis Gilg & Bened. 335 

sasakii Hayata 317, 318 

schlechteri Gilg & Bened. 335, 
954 

scholaris Blco 311 

sororia J.J.S. 307 

sp. Endert 307 

sp. Kaneh. & Hatus. 332 

sparei M.R.Hend. 317, 318 

spatiosa S.Moore 322, 323 

speciosa (Bl.) Bl. 303 

speciosa (non Bl.) Ridl. 307 

spicata Baker 310*, 311, 315 

splendens Bl. 316 

stenophylla Becc. 311, 315 

suaveolens Cammer]. 333 

subreticulata Bl. 311 

sumatrana Miq. 304 

tacapala Leenh. 301*, 323, 
325, 326 
ssp. ceramensis Leenh. 325, 

326 

ssp. gracilis Leenh. 325 
ssp. tacapala 325 

ternatana (non Miq.) Holth. 
& Lam 324 

ternatana Miq. 317, 318 

tetragona Span. 336 

teysmannii Cammerl. 
BIS 

thwaitesii F.v.M. 311 

truncata Bl. 302, 316, 320, 
325 

tubulosa Bl. 302, 320 

umbelliflora Gilg & Bened. 
301, 303, 309, 333 

uniflora (non Merr.) Heine 329 

uniflora Merr. 331 

vaginata K. & G. 322, 323 

valida Miq. 304 

vitiensis Gilg & Bened., non 
Seem. 335 

vitiensis Seem. 336 

volubilis Wall. 311 

wallichiana Bth. 307, 309 

woodiana F.v.M. 301, 325, 
326 

zeylanica = ceilanica 315 


802% 333; 


311, 


Fagus 49 

Feroniella pubescens Tanaka 208 

Flacourtiaceae 5: 1-106, 565; 6: 
4, 943-944, 968 

Flagellariaceae 4: 245-250; 5: 
557 

Flemingia Hunter 299, 301, 303 
fragrans Hunter 307, 309 

Florschuetzia trilobata 975 

Fli(e)ggea serrata Miq. 236 

Forsellesia 227 

Fructus musculiformis Rumph. 
367, 368 

Fumariaceae 62, 63 

Funis toaccae Rumph. 318 


Gaertnera Lamk 387 
Galium 253, 294 
Gardneria Wall. 295, 296, 297, 
361 
fagraeacea F.v.M. 332, 333 
ovata Wall. 362*, 363, 959 
wallichii Wight ex Wall. 363 
Garuga floribunda Decne 917 
Gaultheria Kalm ex L. 470, 471, 
472, 473, 474, 677, 678 map, 
696, 697 
sect. Amphicalyx Endl. 696 
abbreviata J.J.S. 678, 682*, 
683 
acroleia Sleum. 678, 681, 682* 
arfakana Sleum. 679, 689 
atjehensis J.J.S. 678, 680*, 681 
bandongensis Zoll. 694, 695 
barbulata Sleum. 678, 682 
benguetensis Copel. f. 679 
berberidifolia Sleum. 679, 686, 
687* 
blumei F.v.M. 778 
borneensis Stapf 678, 679, 
680* 
calyculata Wernh. 691 
celebica J.J.S. 679, 686 
var. celebica 686 
var. petiolata J.J.S. 686 
ciliolata (Hook. f.) F.v.M. 708 
crenulata Kurz 691 
crenulata (non Kurz) J.J.S. 
694 
cumingiana (non Vidal) Merr. 
695 
cumingiana Vidal 694 
dialypetala Sleum. 678, 680*, 
681 
fragrantissima (non Wall.) K. 
& G. 690 
fragrantissima 
Mor. 688 
var. punctata (Bl.) J.J.S. 688 
fragrantissima Wall. 688 
var. papuana J.J.S. 691 
gracilescens Sleum. 679, 687 
heterophylla (Bl.) Endl. ex 
Hassk. 738 
hirta Ridl. 679, 691 


(non Wall.) 


Sept. 1972] 


intermedia J.J.S. 679, 690 
itoana Hayata 679, 680 
kemiriensis Sleum. 678, 682*, 
683 
latifolia (B\.) Endl. ex Hassk. 
738 
latifola Endl. ex Zoll. 738 
laxiflora Diels 694 
leiotheca Sleum. 692 
leucocarpa BI. 470, 677, 679, 
690, 692, 693* 
f. glabra J.J.S. 692 
f. pubescens J.J.S. 695 
var. hirta Val. ex J.J.S. 679, 
692, 695 
var. leucocarpa 692, 694* 
f. cumingiana (Vidal) 
Sleum. 692, 694 
f. leucocarpa 692 
f. melanocarpa J.J.S. ex 
Amsh. 692, 695 
f. scandens Hochr. 692, 


694 
var. melanocarpa J.J.S. ex 
Steen. 695 
var. melanocephala Steen. 
695 


var. papuana Becc. 689 
var. psilocarpa (Copel.) 
Sleum. 692, 695 
var. seminuda J.J.S. 694 
losirensis Sleum. 678, 682%, 
683 
luzonica A.Gray 720 
malayana Airy Shaw 679, 
682*, 690 
mundula F.v.M. 470, 678, 
680*, 684, 778 
var. mundula 684, 685%, 
686 
var. setifolia 
Sleum. 684, 686 
var. tanythrix 
Sleum. 684, 686 
notabilis Anthony 681 
novaguineensis J.J.S. 679, 
680*, 695 
var. novaguineensis 695 
var. pascua Sleum. 695 
nummularioides D.Don 677, 
678, 680*, 684 
pernettyoides Sleum. 
680*, 695 
pilosa (Bl.) Zoll. 729 
psilocarpa Copel. f. 695 
pullei J.J.S. 679, 691 
var. leiotheca 
Sleum. 692 
var. pullei 691 
punctata BI. 677, 679, 688, 
689*, 694 
repens Bl., non Raf. 684 
solitaria Sleum. 678, 680*, 681 
sp. Vidal 782 
tanythrix Sleum. 686 


(Sleum.) 
(Sleum.) 


679, 


(Sleum.) 


Index to scientific names 


var. setifolia Sleum. 686 
trichophylla (non Royle in 
DC.) Hassk. 684 
viridiflora Sleum. 679, 691 
yunnanensis (Franch.) Rehd. 
694 
(Gay) Lussacia 
761 
Gaylussacia 
Mor. 873 
lanceolata Bl. 761 
Gelonium Roxb. ex Willd. 944 
Gelsemium Juss. 293, 294, 295, 
296, 297, 343, 954 
elegans (Gardn. & Champ.) 
Bth. 343, 344* 
rankinii 954 
sempervirens 343 
sumatrana (Bl.) Boerl. 343 
Geniostoma J.R. & G.Forst. 
293, 295, 296, 297, 369 
acuminatissima Merr., non 
Gilg & Bened. 373 
acuminatissimum Gilg & Be- 
ned. 371 
acuminatum Wall. 373 
acutifolium Hiern 372 
antherotrichum Gilg & Bened. 
Sil 
arboreum O.K. 371 
var. laevigatum(BI.)O.K. 371 
archboldianum Merr. & Perry 
373 
arfakense Kaneh. & Hatus. 
371, 373, 959 
australianum F.v.M. 371 
avene Val. 371, 372 
batanense Mertr. 371 
brevipes Merr. 371 
caulocarpum K.Sch. 371 
celebicum Val. 371 
crassifolium Bth. 373 
cumingianum Bth. 371, 372 
dallmannense Kaneh. & Ha- 
tus. 371 
dasyneurum Gilg & Bened. 
371 
fasciculata Quis. & Merr. 371 
gilgii Merr. & Perry 373 
haemospermum Steud. 293, 
Sil. ove 
var. angustifolia Bl. 371 
var. elongata BI. 371 
var. erosa Bl. 371 
var. laevigata Bl. 371 
var. rugulosa Bl. 371 
lanceolatum (non Bojer) Z. & 
M. ex Mig. 371, 372 
lancilimbum Merr. 371 
lasiostemon Bl. 371 
var. moluccanum Bl. 371 
laxa Elm. 371 
longipes Merr. 371 
micranthum DC. 371 
mindanaense Elm. 371 


Gaylussacia 


elliptica Zoll. & 


999 
miquelianum K. & V. ex Val. 
293, 371 
moluccanum Val. 371 
montanum Zoll. & Mor. 371, 
372 
nigrescens (Blco) Merr. 371, 
B75 
oblongifolium K. & V. ex Val. 
293, 371 
obtusum Merr. & Perry 372 
oleifolium S.Moore 373 
pachyphyllum Merr. 371 
philippinense Merr. 371, 373 
psychotrioides Gilg & Bened. 
371 
pulgarense Elm. 371, 372, 373 
pullei Cammerl. 371, 373 
ramosii Merr. 371 
randianum Merr. 
36973715 372 
reticulata Bl. 371 
rupestre Forst. 293, 369%, 370, 
371, 373, 959 
schlechteri Gilg & Bened. 371 
stenophyllum Gilg & Bened., 
non Merr. 373 
stenophyllum Merr. 371 
weinlandii K.Sch. 371, 372 
Gentiana quadrifaria 953 
Gentianaceae 294 
Gentianales 297 
Geraniaceae 195, 445-449 
Geraniales 195 
Geranium L. 445 
sect. Australiensia 445 
sect. Chilensia 445 
sect. Columbinum 445 
sect. Striata 445 
affine (non W. & A.) Britten 
447 
ardjunense Zoll. & Mor. 447 
australe (non Nees) Allan 448 
carolinianum 445 
clemensiae Knuth 446 
dissectum (non L.) Hemsl. 446 
var. glabratum Hook. f. 448 
glabratum (Hook. f.) Small ex 
Hanks & Small 448 
homeanum Turcz. 446, 448 
macrorrhizum L. 445 
microphyllum Hook. f. 446 
monticola Ridl. 446 
nepalense (non Sweet) Backer 
448 
nepalense (non Sweet) Doct.v. 
Leeuwen 447 
nepalense Sweet 448 
papuanum Ridl. 446 
var. alpestris Ridl. 446 
parviflorum Willd. 448 
philonothum DC. 446 
pilosum [non (Sol.) Forst.] 
F.v.M. 446 
potentilloides L’Hérit. ex DC. 
446, 447* 


& Perry 


1000 


FLORA MALESIANA 


var. ardjunense (Zoll. & 
Mor.) Carolin 447 
var. potentilloides 447 
sarawaketense Knuth 446 
Girocarpus (sphalm.) = Gyro- 
carpus 152 
Glaphyria nitida Jack 814 
Glaux 173 
Glinus lotoides L. 915 
Glochidion 420 
Glosocomia D.Don 118 
Glyptopetalum Thw. 228, 232, 
252, 254, 264, 391, 392, 421, 
930 
acuminatissimum Merr. 256, 
257 
calocarpum 256 
calyptratum 256 
euonymoides Merr. 256 
euphlebium (Merr.) 
256, 257, 258 
fengii (Chun & How) Ding 
Hou 256 
glandulosum Merr. 251 


Merr. 


loheri Merr. 256, 257 
marivelense (Elm.) 
255*, 256, 258, 421 
var. euphlebium Merr. 258 
palawanense Merr. 256, 257 
quadrangulare Prain ex King 
DIAS 259%, LIOVLOI ZO 
remotinervium Merr. 258 
reticulatum Merr. 258 
scortechinii King 248 
zeylanicum Thw. 256, 257 
var. brevipedicellatum Ding 
Hou 255*, 256, 257 
var. zeylanicum 258 
Gnetaceae 4: 336-347; 6: 944— 
949 


Merr. 


Gnetum L. 


sect. Cylindrostachys Mef 
945, 946 
sect. Gnemomorphi Mgf 947 
sect. Gnetum 944, 946, 947 
subsect. Eugnemones Megf 
944, 946 
acutatum Mgf 947 
acutum Mef 945, 947 
arboreum Foxw. 945, 946 
costatum K.Sch. 944, 946 
cuspidatum BI. 945, 947, 948*, 
949 
diminutum Mef 945, 947, 949 
globosum Mef 945, 947, 949 
gnemon L. 944, 946 
var. brunonianum (Griff.) 
Mef 944, 946 
var. gnemon 944, 946 
var. gracile Mgf 944, 946 
var. griffithii (Parl.) Mef 
944, 946 
var. ovalifolium (Poir.) BI. 
945, 946 
var. tenerum Mef 944, 946 


[ser. I, vol. 66 


gnemonoides Brongn. 945, 
946, 947, 949 
klossit Merr. 945, 947, 949 
latifolium BI. 945, 946 
var. funiculare (BI.) Mef 
945, 946 
var. latifolium 945, 946 
var. laxifrutescens (Elm.) 


Mef 945, 946 
var. minus (Foxw.) Megf 
945, 946 


leptostachyum BI. 945,946,949 
var. abbreviatum Mef 945, 
946, 949 
var. leptostachyum 945, 946 
var. robustum Megf 945, 
946, 949 
loerzingii Mgf 945, 947, 949 
macrostachyum Hook. f. 945, 
947 
microcarpum BI. 945, 947 
neglectum BI. 945, 946, 947 
oxycarpum Ridl. 946, 947 
raya Mef 945, 947 
ridleyi Gamble ex Mgf 945, 
946, 947, 949 
tenuifolium Ridl. 945, 946 


Gnidia 3, 4, 30 


oppositifolia (non L.) Blco 48 
Philippinica Meisn. 48 


lucidulus Airy Shaw 977, 981 

macrophyllus (Miq.) Airy 
Shaw 15, 977 

maingayi Hook. f. 977, 978 

micranthus Airy Shaw 977, 
978, 979 

nervosus Airy Shaw 977, 978, 
981 

nobilis Airy Shaw 977, 978, 
981 

pendulus Airy Shaw 978 

reticulatus (Elm.) Merr. 977, 
978, 982 

spectabilis Airy Shaw 977, 982 

stenosepalus Airy Shaw 978, 
980 

velutinus Airy Shaw 977 

xylocarpus Airy Shaw 977, 
978 


Goodenia J.E.Smith 950 


koningsbergeri (Back.) Back. 
ex Bold. 950 

pumilio R.Br. 950* 

purpurascens R.Br. 950 


Goodeniaceae 5: 335-344; 6: 
108, 109, 949-952 

Guioa koelreuteria (Blco) Merr. 
226 

Guttiferae 294, 336 

Gymnosporia Hook. f. 229, 238, 


Gomphostigma Turcz. 295, 296, 
953 
Gomphrena celosioides Mart. 
917 
Gonus Lour. 209 
amarissimus Lour. 211 
Gonystylaceae, see Thymelaea- 
ceae 
Gonystylus Teysm. & Binn. 1, 3, 
5, 6, 976-982 
sect. Auxanthus Airy Shaw 
977, 981 
acuminatus Airy Shaw 977 
affinis Radlk. 978, 979 
areolatus Domke ex Airy 
Shaw 977, 981 
augescens Ridl. 977, 981 
bancanus (Miq.) Kurz 2, 5, 
15, 978, 979 
borneénsis (v.Tiegh.) Gilg 
978, 979, 980, 981 
brunnescens Airy Shaw 978 
calophylloides Airy Shaw 
977, 979, 982 
calophyllus Gilg 977, 979, 982 
confusus Airy Shaw 978 
consanguineus Airy Shaw 
977, 978, 979, 980 
costalis Airy Shaw 977, 980 
decipiens Airy Shaw 978, 980 
forbesii Gilg 977 
glaucescens Airy Shaw 978, 
980 
keithii Airy Shaw 987, 980, 
982 


240, 241 
ambigua Vidal 241, 242 
curtisii King 240 
diversifolia Maxim. 241, 242 
emarginata Thw. 241 
inermis Merr. & Perry 241 
montana Bth. 241 
var. /ittoralis Back. 241 
var. parvifolia Pitard 242 
neglecta Wall. ex Laws. 237, 
420 
nitida Merr. 242 
senegalensis (Lamk) Loes. 241 
spinosa (Blco) Merr. & Rolfe 
241 
var. parva Merr. & Rolfe 
241 
trigyna 238 


Gynandropsis DC. 99, 100 


affinis Bl. 101 

gynandra Briq. 101 

pentaphylla (nor. DC.) Blco 
101 

pentaphylla DC. 101 

speciosa DC. 101 


Gynotroches axillaris Bl. 967 
Gyrinops Gaertn. 2, 3, 4, 5, 39, 


41 map 

caudata (Gilg) Domke 39, 
40*, 42 

decipiens Ding Hou 4, 39, 40*, 
41 

ledermannii Domke 39, 41 

ledermannii (non Domke) 
Merr. & Perry 42 


Sept. 1972] 


moluccana (Miq.) Baill. 4, 39, 
40*, 41 

podocarpus (Gilg) Domke 39, 
40*, 42 

salicifolia Ridl. 39, 40*, 41 

sp. Hall. f. 42 

versteegii (Gilg) Domke 2, 39, 
40*, 42 

walla Gaertn. 39 

walla (non Gaertn.) Koord. 42 

Gyrinopsis Decne 6, 12 

acuminata Merr. 11 

brachyantha Merr. 10, 11, 12 

brachyantha (non Merr. 1912) 
Merr. 11 

citrinaecarpa Elm. 10 

cumingiana Decne 15 
var. pubescens Elm. 15 

decemcostata Hall. f. 15 

grandifolia Quis. 13 

parvifolia Quis. 12 

pubifolia Quis. 15 

salicifolia Quis. 41 

urdanetense Elm. 10 

Gyrocarpus pendulus Blco 152 


Haematosperma arborescens 
Hassk. 371 
Haemodoraceae 5: 111-113 
Haemospermum Reinw. 369 
arboreum Reinw. 371, 372 
Halophila 171 
beccarii Aschers. 953 
decipiens Ostenf. 
var. pubescens Hartog 953 
minor (Zoll.) Hartog 953 
ovata Gaud. 953 
Hamamelidaceae 5: 363-379; 6: 
145, 195, 472, 952 
Hannoa Planch. 198, 202 
Harrisonia R.Br. 193, 195, 196, 
207 
bennettii Benn. 208 
var. multijuga F.-Vill. 208 
var. paucijuga F.-Vill. 208 


brownii A.Juss. 207, 208* 
map 

citrinaecarpa Elm. 208 

paucijuga Oliv. 208 

perforata (Blco) Merr. 207, 


208*, 209 map 
Hasskarlia Meisn. 51 
Hebonga Radlk. 215 
mollis Radlk. 219 
obliqua Radlk. 219 
siamensis Radlk. ex Craib 219 
Hedraianthera F.v.M. 390, 393, 
394, 930 
porphyropetala F.v.M. 393 
Hedyotis 379 
Heimerlia Skottsb. 457, 459 
brunoniana Skottsb. 461 
Heimerliodendron Skottsb. 457, 
459 
brunonianum Skottsb. 461 


Index to scientific names 
Helicia rufescens Prain 965 
Heliciopsis rufidula Sleum. 965 
Helobieae 157 
Hemiphragma heterophylla 

Wall. 960 
Hemiscolopia 
Sloot. 944 

Herbaceae Hutch. 62, 451 
Hermicium (sphalm.) = Hermi- 
dium 
Hermidium alipes S.Wats. 450 
Hexaspora C.T.White 240 
pubescens C.T.White 229 
Heynea Roxb. 226 
Hibbertia 49 
Hippobroma G.Don 139, 140 
longiflora G.Don 140 
Hippocratea 231, 389, 397 
arborea Roxb. 420 
beccarii Tuyn 402 
bojeri Tul. 420 
brachystachys Ridl. 400 
cassinoides DC. 402 
cumingti Laws. 399, 400 
ellipticarpa Merr. 402 
ferruginea King 402 
glaga Korth. 402 
grahami Wight 402 
hasseltiana Miq. 398 
indica (non Willd.) Span. 402 
indica Willd. 401, 402 
f. timorensis Miq. 401 
var. evonymoides Bl. 401 
lawsonii Elm. 400, 417 


trimera (Boerl.) 


macrantha auct. non Korth. 
400 

macrantha (non Korth.) King 
400 


macrantha Korth. 398 
macrantha (non Korth.) Rolfe 
399 
maingayi Laws. 262 
maingayi (non Laws.) Vidal 
269 
megalocarpa Merr. 402 
nigricaulis Ridl. 400 
obtusa Ridl. 406 
obtusifolia (non 
Koord.-Schum. 400 
obtusifolia (non Roxb.) Merr. 
406 
pauciflora DC. 400 
f. minor Miq. 400 
f. novoguineensis Miq. 400 
rigida Span. 400 
salacioides Korth. 402, 404 
sogerensis Baker f. 398 
sp. Burk. & Haniff 406, 407 
timorensis Span. 421 
trichopetala Merr. 399 
trilobulata Ridl. 398 
volubilis (non L.) Blco 401, 
402 
zippeliana Miq. 402 
Hippocrateaceae 228, 229, 230, 


Roxb.) 


1001 
231, 389, 390, 932 
Hippocraticeae Juss. 389 
Hiptage /awsonii Elm. 417 
Holosteum hirsutum L. 915 
Homalanthus giganteus 152 
Homalium Jacq. 


subg. Blackwellia (Lamk) 
Warb. 944 

sect. Blackwellia Bth. 944 

subg. Pythagorea  (Lour.) 
Sleum. 944 

sect. Pythagorea (Lour.) 
O.K. 944 


Hornera Jungh. 48 
glomerata Jungh. 48 
umbellata Jungh. 48 
Hottonia sessiliflora Vahl 965 
Huertea 49 
Hydnocarpus 963 
Hydrilla verticillata 164 
Hydrocaryaceae 4: 43-44; 6: 
982; see Trapaceae 
Hydrocharitaceae 5: 
5695163) 15Se 1752 
Hydrophyllaceae 4: 207-209 
Hymenanthes Bl. 656 
Hypericineae 49 
Hypsagine Jack ex Burk. 421 
Hyptiandra Hook. f. 202 


381-413, 


laera Copel. f. 740 
lanaensis (Merr.) Copel. f. 745 
loheri (Merr.) Copel. f. 741 
lucida (Merr.) Copel. f. 742 
Ignatia amara L. f. 347, 349 
Ignatiana philippinica Lour. 347, 
349 
Ilex racemosa Oliv. 290 
Indotristicha v.Royen 963 
malayana Dransf. & Whitmo- 
re 963, 964* 
Inocarpus 201, 226 
Ipomoea sect. Ipomoea 940 
sect. Pharbitis (Choisy) Gri- 
seb. 940 
acuminata Ruiz & Pay. 941 
acuminata (Vahl) R. & Sch. 
939, 940, 941 
angulata Ortega 941 
congesta R.Br. 939, 940, 941 
dasysperma Jacq. 941 
digitata L. 940, 941 
gracilis R.Br. 940, 941 
littoralis Bl. 940, 941 
mauritiana Jacq. 940, 941 
maxima (L. f.) Don ex Sweet 
940, 941 
< multifida (Raf.) 
940, 941 
ochracea (Lindl.) G.Don 939, 
941 
ochroleuca 
939, 941 
pes-caprae (L.) R.Br. 941 
plebeia R.Br. 940 


Shinners 


(sphalm.) Span. 


1002 


FLORA MALESIANA 


[ser. I, vol. 68 


ssp. africana Meeuse 940 
ssp. indica Verdc. 940 
ssp. plebeia 940 
purpurea Roth 941 
riparia G.Don 940, 941 
rubens Choisy 940, 941 
sepiaria Koen. ex Roxb. 940, 
941 
< sloteri 939 
tuba (Schlechtend.) 
941 
tuberculata Ker-Gawl 941 
Irvingella v.Tiegh. 223 
harmandiana v.Tiegh. 225 
malayana v.Tiegh. 225 
oliveri v.Tiegh. 225 
Irvingia Hook. f. 193, 194, 195, 
196, 201, 223, 969, 972 
harmandiana Pierre ex Lecom- 
te 225 
longipedicellata Gagnep. 225 
malayana Oliv. ex Benn. 223, 
224*, 225*, 226 map 
oliveri Pierre 225 
Irvingiaceae 195, 972 
Iso Kuroggi Kaempf. 253 
Tsolobus DC. 121 
caespitosus Hassk. 130 
campanuloides DC. 130 
kerii DC. 130 
radicans DC. 130 
roxburghianus DC. 130 
Tsotoma Lindl. 108, 139, 140, 141 
longiflora Presl 140 
runcinata Hassk. 140 
Itea 280 
Ixonanthaceae 970, 972 
Ixonanthes Jack 971, 972 


G.Don 


Jacquemontia Choisy 939 
browniana Ooststr. 939 

Jasione 108 

Jaundea Gilg 934 

Johnia Roxb. 404 
sumatrana Jack 421 

Juglandaceae 143-154, 953 

Juglandicarya Reid & Chandler 
143 

Juglans 143 
plerococca = pterococca 154 
pterococca Roxb. 152, 154 
regia 145 
rupestris Engelm. 143 
sieboldiana 143 

Juncaceae 4: 210-215; 5: 557; 6: 
953 

Juncaginaceae 4: 56-57; 5: 554 

Juncodes — silvaticum  (Huds.) 
O.K. 953 

Juncus effusus L. 953 
prismatocarpus R.Br. 953 


Kaernbachia Schltr 49, 51 
brachypetala Schltr 59 
pentandra Schltr 59 


Kalmia 472 
Karin-Njoti Rheede 199 
Kelleria Endl. 43 
papuana Domke 44 
Kentia Steud. 299, 301, 309 
morindaefolia Steud. 311, 315 
Kippistia Miers 389 
Klainedoxa 194, 195, 972 
Kokoona Thw. 228, 231, 232, 
258, 260 map, 264, 389, 391, 
392, 930, 932 
coriacea King 260, 261 
lanceolata Ridl. 261 
littoralis Laws. 231, 260, 261 
luzoniensis Merr. 262, 402 
ochracea (Elm.) Merr. 259%, 
260 
ovatolanceolata 
259*, 260, 261 
reflexa (Laws.) Ding Hou 231, 
259*, 260, 262 
scortechinii King 261 
scortechinii (non King) Steen. 
261 
sessilis Ding Hou 260, 261 
zeylanica 260 
Kokoonia = Kokoona 260, 402 
Kuhlia H.B.K. 301 
Kuhlia Reinw. ex BL, 
H.B.K. 299, 301, 309 
morindaefolia Reinw. ex BI. 
Sia sis 
Kurrimia Wall. ex Arn. 230, 280 
archboldiana Merr. & Perry 
284 
bipartita Laws. 283 
calophylla Wall. 283 
gracilis Vidal 284 
indica Gamble 283 
luzonica Vidal 282 
maingayi Laws. 283 
minor Rid]. 282 
paniculata Wall. ex Arn. 282 
pulcherrima (non Wall.) Baker 
f. 420 
pulcherrima Wall. ex Laws. 
283 
robusta Kurz 282, 283 
var. roxburghii Pierre 283 
var. thorelii Pierre 283 
Kurrimia Wall. ex Meisn. 280 
Kurrimiaceae 230 


Ruidly228, 


non 


Labordia Gaud. 295, 296, 370 
Lachnolepis Miq. 39 
moluccana Mig. 39 
Lachnopylis 296 
Lafoénsia Vand. 975 
Lagansa alba Rumph. 103 
rubra Rumph. 101 
Lagerstroemia L. 975 
Lasianthus furcatus (Miq.) Bre- 
mek. 373 
Lasiolepis Benn. 207 
bennettii Planch. 208 


var. @ paucijuga (Benn.) 
Planch. 208 
var. 2B multijuga (Benn.) 
Planch. 208 
paucijuga Benn. 208 
Lasiosiphon 4 
scandens Endl. 25 
Launaea pinnatifida 223 
Lauraceae 194 
Laurentia Mich. ex Adans. 109, 
110, 139, 141 
sect. Isotoma (R.Br.) Endl. 
107, 140 
subg. Jsotoma Peterm. 140 
gasparrinii (Tineo) Strobl. 140 
longiflora (L.) Peterm. 108, 
138*, 140 
var. runcinata (Hassk.) 
Wimmer 140 
Lechenaultia _ filiformis 
951 
Ledum 472 
Leea 52 
Legousia Durand 928 
speculum-veneris (L.) Fisch. 
141, 928 
Leguminosae 216, 226, 933 
Lemnopsis Zoll. 171 
Leptopteris Bl. 343 
sumatrana Bl. 343 
Leucopogon R.Br. 424, 425 
sect. Anacyclodon (Jungh.) 
Miq. 425 
sect. Stypheliopsis Miq. 425 
abnormis Sond. 433 
acuminatus R.Br. 432 
acuminatus (non R.Br.) Du- 
perrey 433 
colensoi Hook. f. 428 
hookeri Sond. 428 
javanicus de Vriese 431 
lancifolius Hook. f. 433 
malayanus Jack 426 
var. o Kurz 426 
var. moluccanus (non 
Scheff.) Kurz 426, 433 
moluccanus Scheff. 433 
obovatus Fawe. 429, 432 
obtusatus Hook. f. 428 
ophirensis Griff. 943 
papuanus C.H.Wright 443 
Philippinensis (Merr.) Hoso- 
kawa 429 
suaveolens Hook. f. 428 
Leucosmia Bth. 15 
Leucothoé 472 
Libocedrus 333 
Lightfootia L’Hérit., non Sw. 
109, 111; 1123918 
gracilis Miq. 115 
var. lavandulaefolia Miq. 
ib iey 
Lightfootia Sw. 113 
Ligia Fasano 28 
Lignosae 62 


R.Br. 


Sept. 1972] 


Index to scientific names 


Lignum colubrinum _ timorense 
Rumph. 350 
Limnophila sp. 965 
Limonia pubescens 
Hook. f. 208 
Linaceae 970 
subfam. Ixonanthoideae 970 
tribe Erythroxyleae 195 
Linaria 294 
Lindernia subulata R.Br. 960 
Linociera sp. 280 
vitiensis A.C.Smith 336 
Linostoma Wall. ex Endl. 2, 4, 
5, 26, 982 
sect. Eulinostoma Meisn. 27 
sect. Psilaea (Miq.) Hall. f. 27 
subg. Linostoma Kurz 23 
subg. Nectandra {(non Berg.) 
Roxb.] Kurz 27 
leucodipterum Hall. f. 27 
longiflorum Hall. f. 2, 26*, 27, 
982 
pauciflorum Griff. 2, 26*, 27, 
982 
scandens Kurz 25 
Lissanthe R.Br. 424 
montana R.Br. 429 
Lobelia L. 107, 108, 109, 110, 
MOM 1 122)) 129591355137, 
140, 928 
sect. /sotoma R.Br. 140 
affinis Wall. ex G.Don 128, 
129 
var. lobbiana Clarke 128 
alata Labill. 136 
aligera Haines 129 
alsinoides Lamk 107, 122, 123, 
12659127amap; 128; 130, 
1527 
anceps L. f. 129 
anceps Thunb., non L. f. 136 
angulata Forst. 108, 110, 123, 
1SS51347°1135* 
archboldiana (Merr. & Perry) 
Moeliono 123, 131*, 928 
arenarioides DC. 129 
arfakensis Gibbs 134 
aromatica Moon ex Wight, 
nomen 123 
barbata Warb. 128 
beddomeana Wimmer 124 
hegonifolia Wall. 133 
bialata Merr. 129, 130 
borneensis (Hemsl.) Moeliono 
122, 133 map, 928 
brachyantha Merr. & Perry 
Ay I1si: 182* 
caespitosa Bl. 130 
campanuloides Thunb. 130 
camptodon Wimmer 124 
chevalieri Danguy 127 
chinensis (non Lour.) Hance 
127 
chinensis Lour. 108, 123, 130* 
cliffortiana L. 136 


Wall. ex 


colorata Wall. 123, 126 
conferta Merr. & Perry 123, 
185" 
cordigera Cav. 136 
decurrens Roth 129 
dichotoma Migq. 129 
var. aligera Wimmer 129 
var. pilosella Wimmer 129 
donanensis v.Royen 928 
doniana Skottsb. 124 
dopatrioides Kurz 127, 128 
dubia de Vriese 115 
epilobioides Wimmer 124 
var. sarasinorum Wimmer 
124 
erecta Hook. f. & Th. 123 
erinoides (non L.) Thunb. 130 
erinus L. 136 
erinus (non L.) Thunb. 130 
eryliae Fischer 124 
eurostos Voigt 123 
excelsa Lesch. ex Roxb. 123 
filiformis (non Lamk) Cav. 
126 
var. /uzoniensis Pers. 126 
fossarum Wimmer 124 
graminea Lamk 136 
griffithii Hook. f. & Th. 127 
var. dopatrioides 127 
var. genuina 127 
haenkeana DC. 136 
hainanensis Wimmer 127 
hederacea Cham. 134 
heyneana = heyniana 
127, 129 
heyniana R. & S. 123, 129, 
130 map, 132*, 176 
horsfieldiana Mig. 134 
hosseusii Wimmer 127 
var. villosa 127 
inconspicua Rich. 129 
inflata L. 108 
javanica Thunb. 133 
/aurentia L. 140 
laxiflora H.B.K. 136 
var. nelsonii McVaugh 136 
leucanthera Kerr 124 
linnaeoides (Hook. f.). Petrie 
134, 135 
var. brevipilis 134 
littoralis R.Cunn. ex A.Cunn. 
134 
lobbiana Hook. f. & Th. 128 
longifiora L. 140 
longifolia DC. 136 
luzoniensis (Pers.) Merr. 127, 
128 
micrantha Hook. 129 
microcarpa Clarke 127 
montana Reinw. ex BI. 107, 
108, 122, 132*, 133 map 
nelsonii Fernald 136 
nicotianaefolia Roth ex R. & 
SHO, (NO 22.9123: 124". 
125* map, 126 


126, 


1003 
var. bibarbata Wimmer 123 
var. macrostemon Skottsb. 
123 
var. mollis Elm. 123 
var. nicotianaefolia 
mer 123 
var. trichandra Clarke 123 
nummularia Lamk 133 
obliqua Ham. ex D.Don 133 
palustris Kerr 124 
paradoxa Wimmer 134, 135 
philippensis Skottsb. 124 
plumierii L. 951 
pratiana Gaudich. 133 
pumila Burm. f. 136 


Wim- 


purpurescens Wall., nomen 
123 

pyramidalis Wall. 123 
var. B 123 


radicans (non Thunb.) Hos- 
seus 127 
radicans Thunb. 130 
reinwardtiana DC. 129 
reniformis Cham. 134 
robusta Wall. ex Voigt 123 
rosea Wall. 123 
rotundifolia Banks & Sol. ex 
Hook. f. 134 
rugulosa Graham 134 
sebae DC. 129 
seguinii Léveillé & Van 124 
sinaloae 140 
sp: Gril 127 
sp. Steen. 126 
stimulans Ham. ex D.Don 123 
stipularis Roth 127, 128 
subcuneata Miq. 128 
subincisa Wall. ex DC. 129 
subracemosa Miq. 129 
var. rigidior 129 
succulenta Bl. 128, 129 
f. glabra 128 
var. lobbiana 128 
sumatrana Merr. 122, 126* 
syphilitica L. 136 
taccada Gaertn. 951 
terminalis Clarke 127, 128 
tetragona Bl. 136 
thorelii Wimmer 127 
trialata Ham. ex D.Don 126, 
127, 129 
var. lamiifolia 129 
triangulata Roxb., nomen 127 
trichandra Wight 123 
trigona (non Roxb.) Hook. f. 
& Th. 129 
trigona Roxb., nomen 126 
wallichiana Hook. f. & Th. 
123 
wallichii Steud. 123 
zeylanica L. 122, 123, 128, 129 
var. affinis 128 
zeylanica (non L.) Moon 129 
var. aligera Haines 129 
var. parviflora Danguy 129 


1004 


FLORA MALESIANA 


[ser. I, vol. 68 


var. walkeri Clarke 129 
Lobeliaceae 108, 109 
Locandi Adans. 199 
Locandia O.K. 201 

glandulifera Pierre 201 

indica O.K. 201 

madagascariensis O.K. 201 

mekongensis Pierre 201 

merguensis Pierre 201 

pendula Pierre 201 
Loeseneriella A.C.Smith 262, 

390, 391, 397 

cumingii (Laws.) Ding Hou 

397, 399 

macrantha (Korth.) 

Smith 397, 398 map 

pauciflora (DC.) A.C.Smith 

399*, 400, 417 

sogerensis (Baker 

Smith 397, 398 

zippeliana A.C.Smith 402 
Logania 295, 296 
dentata (Elm.) Hayata 960 
Loganiaceae 298-387, 953-960 
subfam. Buddlejoideae 294 
subfam. Loganioideae 294 
tribe Antonieae 294, 295, 296, 
297 
tribe Buddlejeae 294, 295, 296, 
297, 953 
tribe Desfontainea Leeuwenb. 
954 
tribe Euloganieae 295 
subtribe Buddlejeae 295 
tribe Gelsemieae 294, 295, 
296, 297 
tribe Loganieae 295, 296 
tribe Plocospermeae 954 
tribe Potalieae 294, 295, 296, 
297 
tribe Retzieae Leeuwenb. 954 
tribe Spigelieae 295, 296, 297 
tribe Strychneae 294, 295, 296 
Loganiales 294, 297 
Lonicera malayana Hend. 929 
Lophopetalum Wight ex Arn. 
DDD Bae 22930 Me Sila 32. 
260, 262, 266 map, 391, 392, 
930, 932 
subg. Solenospermum (Zoll.) 
Val. 262 
beccarianum Pierre 263*, 264, 
266 
celastroides Laws. 269 
celebicus Koord. 270 
coriacea Ridl. 261 
curtisii King 268, 269 
dubium Laws. 261 
fimbriatum (non Wight) F.- 
Vill. 269 
fimbriatum Wight 267, 268 
floribundum Wight 229, 265, 
266, 930 
fuscescens Kurz 269 
glabrum Ding Hou 265, 266 


A.C. 


ED ALC: 


grandiflorum Arn. 264 

havilandii Ridl. 266 

intermedium Ridl. 270 

javanicum (Zoll.) Turez. 228, 
265, 269 

javanum = javanicum 269 

ledermannii (Loes.) Ding Hou 


265, 271 
littoralis Ridl. 261 
macranthum (Loes.) Ding 


Hou 263*, 265, 268, 930 
maingayi Ridl. 261 
micranthum Loes. 265, 268 
multinervium Ridl. 227, 228, 

232655 2702 Tlie 
oblongifolium King 270 
oblongum King 269 
pachyphyllum King 265, 267 
pallidum Laws. 231, 263%, 

264, 265, 268 

var. curtisii (King) Ridl. 268 
paucinervium Merr. 270 
penduculatum Ridl. 250 
reflexum (non Laws.) King 

266 
reflexum Laws. 262 
rigidum Ridl. 228, 263*, 265, 

267 
scortechinii King 266 
sessilifolium Ridl. 228, 262, 

264, 265 
sp. Thorenaar 269*, 270 
subobovatum King 265, 272 
subsessile Ridl. 267 
torricellense Loes. 265, 271 
toxicum Loher 230, 270 
wallichii Kurz 269 
wightianum Arn. 231, 264, 

265, 267 

var. macrocarpum 

267 
winkleri Loes. 267 
Lubinia Vent. 177, 184 
lineariloba Pax 184 
lubinoides Pax 184 
mauritiana Spreng. 184 
spathulata Vent. 184 
Lunasia 221 
Luronium natans (L.) Raf. 953 
Lussa Rumph. 209 
radja Rumph. 211 
(Gay)Lussacia = Gaylussacia 
761 
Luzula silvatica (Huds.) Gaudin 
953 
Lyonia Nutt. 469, 472, 474, 674 
ovalifolia (Wall.) Drude 673*, 
674*, 675 map 
Lysimachia Tourn. ex L. 173, 
(WEE IB 17/7/ 
subg. Lysimachia (£u-lysima- 
chia) 177 
sect. Alternifoliae Knuth 
180 
sect. Apodanthera 177 


Pierre 


subsect. Ramosae 177 
ser. Evalves 177 
ser. Valvatae 177 
sect. Nummularia 177, 183 
ser. Debiles 177, 183 
ser. Japonicae 177, 183 
subg. Palladia (Moench) H.- 
M. 177, 184 
acroadenia Maxim. 185, 186 
ardisioides Masam. 178, 179 
capillipes Hemsl. 177, 178, 
179, 180 map. 181, 182, 964 
chapaensis Merr. 181 
consobrina Hance 185 
cuspidata BI. 180, 181 
var. glabra Mor. 181 
var. glabrescens Knuth 181 
var. hispida Knuth 181 
cuspidata (non BI.) Hand.- 
Mazz. 179 
debilis Wall. 182 
var. minor Baudo 182 
var. vulgaris Baudo 182 
decurrens Forst. f. 177, 178, 
184, 185 map, 965 
var. acroadenia Makino 185 
var. recurvata Mats. 185 
deltoides Wight 183 
floribunda Zoll. & Mor. 180, 
182 
fragrans Hayata 178, 179 
garrettii Fletch. 178 
glaucophylla Hook. & Arn. 
184, 185 
japonica Thunb. 178, 181, 182 
var. japonica 177, 182, 183 
map 
var. minutissima Masam. 
183 
var. papuana S.Moore 177, 
183 map 
var. subsessilis Hara 182 
var. thunbergiana Hara 182 
javanica Bl. 185 
keiskeana Miq. 185, 186 
klattiana Hance 181 
laxa Baudo: 17 eigeceio =: 
180 map, 182, 964 
lineariloba Hook. & Arn. 184 
lobelioides Wall. 186 
lubinoides Sieb. & Zucc. 184 
maculata R.Br. 182 
mauritiana Lamk 177, 178, 
183, 184 map 
montana (Reinw.) Bakh. f. 
177, 178, 180, 181 map 
var. montana 181 
var. platyphylla 
Bentvelzen 181 
microphylla Merr. 182, 183 
minutissima (Masam.) Ma- 
sam. 182, 183 
multiflora Wall. ex Duby 185 
nebeliana Gilg 184 
obovata Ham. ex Hook. f. 186 


(Merr.) 


Sept. 1972] 


Index to scientific names 


peduncularis Wall. ex Kurz 
177, 178, 182 map 

pierrei Petitmengin 182 

platyphylla Merr. 181 

procumbens Baudo 183 

ramosa Wall. ex Duby 180, 
181, 182 
var. grandiflora Franch. 180 
var. typica Knuth 178, 180 
var. zeylanica Hook. f. 180 

rapensis F.B.H.Brown 185 

recurvata  (Mats.) Masam. 
185, 186 

rubida Koidz. 184 

siamensis Bonato 182, 183 

sikokiana Miq. 177, 178, 179* 
map, 180 

simulans Hemsl. 178 

sinica Merr. 185 

solanoides H.-M. 174 

spathulata Schouw 184 

spatulata = spathulata 184 

suborbicularis Went 182, 183 

trichopoda Franch. 181 

uliginosa Bl. 181, 182 

uliginosa (non Bl.) Klatt 181 

vulgaris L. 174 

Lythraceae 198, 975 

subtribe Diplusodontinae 975 

subtribe Lagerstroemiineae 
975 


Maba hierniana K. & G. 932 

Macanea arborea Blco 421 

Macgregorianthus Merr. 23 
paniculatus Merr. 25 

Maerua 61 

Maesa membranifolia Mez 237 
perakensis Rid]. 290 
ramentacea (Roxb.) A.DC. 

237 

Magnolia 228 

Magnoliaceae 228 

Malpighiaceae 5: 125-145, 566; 
6: 960 

Malpighiales 195 

Mangifera L. 231 
glauca Rottb. 286 

Mannia Hook. f. 202 

Manotes asiatica Gagnep. 205 

Manungala Blico 199 
pendula Blco 201 

Mastixiodendron — pachyclados 
(K.Sch.) Melch. 336 

Mauduita Comm. ex DC. 199 
penduliflora Comm. ex DC. 

200 


Maurocenia sect. Triceros O.K. 
51 
pomifera O.K. 58 
zollingeri O.K. 56 

Maytenus Molina 227, 228, 229, 
2305 231-,)232/9238, 240) 243; 
391, 392, 930 
acuminatus 228 


bilocularis (F.v.M.) Loes. 243 

boaria Molina 240 

crassa Ding Hou 240, 242 

cupularis Ding Hou 239*, 240 
243 


curtisii (King) Ding Hou 
239*, 240 
dispermus 230 
diversifolia (Maxim.) Ding 


Hou 231, 232, 240, 241, 242, 
391, 930 
emarginata (Willd.) Ding Hou 
229, 239*, 240, 241, 930 
ilicifolia Mart. 229 
magnifolia 228 
nitida Mart. 242 
rapakir Loes. 242 
senegalensis (Lamk) Exell 240, 
241 
spinosa (Gris.) Lourt. & O’- 
Donell 229 
vitiensis (A.Gray) Ding Hou 
242 
Mazus pumilus (Burm. f.) Steen. 
136 
Medicia Gardn. & Champ. 343 
elegans Gardn. & Champ. 343 
Melastomataceae 914 
Meliaceae 194, 212, 220 
Melichrus R.Br. 424 
Menyanthes trifoliata 294 
Merremia Dennstedt 939 
sect. Eu-Merremia 939 
sect. Merremia 939 
aniselifolia Ooststr. 939, 940* 
borneensis Merr. 939 
crispatula Prain 939 
emarginata (Burm. f.) Hall. f. 
939 
gangetica (L..) Cufod. 939 
peltata (L.) Merr. 939 
quinata (R.Br.) Ooststr. 939 
tridentata (L.) Hall. 939 
ssp. hastata Ooststr. 939 
Messerschmidia argentea 197 
Micropyxis Duby 176 
pumila Duby 176 
tenella Wight 176 
Microsemma 3 
Microtropia Rchb. 272 
Microtropis Wall. ex Meisn. 
DI TEDISS 2295230) 2325272. 
275 map, 391, 392, 930 
basilanensis Merr. & Freem. 
279 
bicolor Merr. & Freem. 276 
bivalvis (non Wall.) Koord. 
277 
bivalvis (Jack) Wall. 272, 274, 
276 
borneensis Metr. 
279 
chartacea Merr. & Freem. 279 
coriacea Wall. ex Ettingsh. 
280, 412 


& Freem. 


1005 
curranii Merr. 273, 274, 278 
var. obovata Merr. & 
Freem. 278 
var. zambalesensis Merr. & 
Freem. 278 
discolor (Wall.) Wall. 272, 
274, 275 


elliptica King 273*, 274, 277 
fasciculata Quis. & Merr. 279 
filiformis King 232, 276, 277, 
391 
javanica Merr. & Freem. 277 
kinabaluensis Merr. & Freem. 
274, 277 
var. acuminata 277, 278 
lanceolata Boerl. & Koord. 
280 
longifolia Wall. 274 
longirostris Merr. & Freem. 
277 
ophirensis Ridl. 277 
ovata Merr. & Freem. 274, 
275, 279 
pachyphylla Merr. & Freem. 
274 
pauciflora Boerl. ex Merr. & 
Freem. 276 
peduncularis Rid). 276, 277 
Philippinensis Merr. 279 
platyphylla Merr. 274, 275, 
279 
var. ellipticifolia Merr. & 
Freem. 279 
ramiflora (non Wight) Thw. 
279 
rigida Ridl. 275, 278 
rostrata Merr. 279 
rubra Elm. ex Merr. & Freem. 
279 
sterrophylla Merr. & Freem. 
277, 278 
suborbiculata Merr. & Freem. 
279 
sumatrana Merr. 275, 278 
tenuis Symington 272, 274, 
275, 276 
tetrameris Ding Hou 
DBF DTA 215 
valida Ridl. 274, 276 
vinculans Boerl. & Koord. 277 
wallichiana Wight ex Thw. 
215, 279 
zeylanica Merr. & Freem. 279 
Mirabilis L. 450, 451 
jalapa L. 450, 451 
Mirabilis Rumph. 451 
longiflora (non L.) Blco 451 
Mischopleura Wernh. 914 


272, 


Mitrasacme Labill. 293, 295, 
296, 297, 375, 378 
albomarginata Leenh. 379, 


385* 
alsinoides R.Br. 379, 384 
alsinoides (non R.Br.) Clarke 
384 


FLORA MALESIANA 


[ser. I, vol. 66 


var. indica (Wight) Hara 


384 
alsinoides (non R.Br.) Merr. 
382 
bogoriensis Leenh. 379, 386*, 
960 
capillaris Wall. 382, 383, 384 
chinensis Griseb. 382 
connata R.Br. 381 
crystallina Griff. 384 
elata R.Br. 293, 379, 380, 381* 
map, 960 
var. brevicalyx Leenh. 380*, 
381, 960 
var. elata 380*, 381 
erophila Leenh. 380, 384, 960 
exserta F.v.M. 382 
indica Wight 379, 384 map, 
386, 960 
longiflora F.v.M. ex Bth. 380, 
381 
malaccensis Wight 382, 383 
micrantha Domin 382, 383, 
384 
neglecta Leenh. 379, 386, 960 
nudicaulis (non Bl.) Bth. 382, 
383, 960 
nudicaulis Reinw. ex Bl. 380, 
381, 382 
paludosa R.Br. 385 
polymorpha R.Br. 379, 384 
polymorpha (non R.Br.) Clar- 
ke 382, 383, 384 
var. grandiflora Hemsl. 382 
var. parishii Clarke 382 
pusilla Dalz. 384, 385 
pygmaea R.Br. 379, 382, 960 
var. grandiflora (Hemsl.) 
Leenh. 383 
var. malaccensis (Wight) 
Hara 380, 382*, 383, 384 
var. parishii (Clarke) 
Leenh. 383 
var. pygmaea 380, 382*, 
383*, 384, 960 
saxatilis Back. 379, 385, 960 
setosa Hance 382 
setosa (non Hance) Masam. 
382 
tenuiflora Bth. 379 
trinervis Span. 380, 381, 382, 
383 
Mitrasacmopsis Jovet 297 
Mitreola Boehm. 375 
Mitreola L. 296, 375, 959 
inconspicua Zoll. & Mor. 375 
oldenlandioides (Wall.) G.Don 
YS, SMT) 
paniculata Wall. ex G.Don 
375 
var. lilacina Back. 375, 377 
perriert Jovet 377 
petiolata (Gmel.) Torr. & 
Gray 375, 959 
reticulata Tirel 960 


sphaerocarpa (Leenh.) Leenh. 
960 
turgida Jovet 377 
Mitreola Schaeff. 375 
Modagam Rheede 315 
Molinadendron Endress 952 
Mollugo nudicaulis Lamk 384 
Monocelastrus Wang & Tang 
231233234 
monosperma (Roxb.) Wang. & 
Tang 234 
virens Wang & Tang 234 
Monochoria hastata (L.) Solms 
964 
Monopsis simplex (L.) Wimmer 
129 
Monotropa L. 472 
humilis D.Don 670 
uniflora L. 670 
Monotropaceae 473 
Monotropanthum Andres 670 
Monotropastrum Andres 469, 
470, 474, 670 
humile (D.Don) Hara 670, 
671* 
Moringa domestica Ham. ex 
Henschel 960 
oleifera Lamk 960 
Moringaceae 4: 45-46; 5: 554; 
6: 62, 960 
Mostuea 294, 295, 296 
Moya Griseb. 238 
spinosa Griseb. 229 
Muenteria Walp. 212 
Myoporaceae 4: 265-266 
Myricaceae 4: 276-279; 6: 143, 
146 
Myristica Jaurina (non BI.) 
Hochr. 933 
Myristicaceae 194 
Myrsinaceae 173, 174, 237 
Mystroxylon 229, 284 


Najadaceae 157-171 
Najadales 157 
Najadeae 171 
Najas L. 157, 158 
subg. Caulinia Aschers. ex 
Rendle 161, 164 
sect. Caulinia A. Braun 161, 
164 
subg. Eunajas Aschers. ex 
Rendle 162 
sect. Eunajas A.Braun 162 
sect. Najas 161, 162 
bengalensis Horn af Rantzien 
169 
brevistyla Rendle 167 
browniana Rendle 158, 160*, 
161, 163 map, 165 
celebica Koord. 167, 168 
falciculata A.Braun 165, 166 
falciculata’ (non A.Braun) 
Coert 168 
foveolata A.Braun ex Rendle 


166 
gracillima (A.Braun) Magnus 
165, 166 
graminea Del. 158, 161, 169, 
170 map 
var. angustifolia Rendle 169 
var. delilei Magnus 169 
var. graminea 158, 160*, 
170 
var. minor Rendle 169 
var. robusta de Wilde 170 
var. (B) ‘enuifolia (R.Br.) 
A.Braun 167, 168, 169 
graminea (non Del.) Mig. 168 
graminea (non Del.) Ridl. 169 
hagerupi Horn af Rantzien 
165 
horrida 170 
indica (Willd.) Cham. 158, 
160*, 161, 164, 166, 167 
map, 170 
var. gracillima A.Braun ex 
Engelm. 165 
var. macrodictya A.Braun 
166 
var. rigida A.Braun 166 
intermedia Gorski 162 
kingii Rendle 166 
kurziana Rendle 160*, 161, 
163 map, 165 
lacerata Rendle 166 
lobata Blco 166 
major All. 162 
B angustifolia A.Braun 162 
¢ intermedia A.Braun 162 
malesiana de Wilde 158, 160*, 
161, 169 map 
marina L. 158, 159, 160, 161, 
162, 170 
var. B 162 
var. y 162 
var. angustifolia Rendle 162 
var. intermedia Rendle 162 
var. marina 160*, 162, 163 
map 
var. sumatrana de Wilde 
160*, 162, 163 map 
var. zollingeri Rendle 158, 
160*, 162, 163* 
minor All. 161, 164 
var. indica A.Braun 166 
minor (non All.) Hook. f. 166 
obyvoluta Blco 171 
oguraensis Miki 164 
palustris Blco 166 
pseudograminea W.Koch 167, 
168 
seminuda Griff. ex Voigt 169 
tenuifolia (non R.Br.) auct. 
168 
tenuifolia R.Br. 158, 161, 167, 
168 
ssp. pseudograminea (W. 
Koch) de Wilde 167, 168 
var. celebica (Koord.) de 


Sept. 1972] 


Index to scientific names 


Wilde 158, 160*, 168 


var. pseudograminea 
159*, 160*, 168 map 
ssp. tenuifolia 167, 168 
tenuifolia (non R.Br.) Miq. 
166 
Naumburgia Moench 174, 175, 
iL 7/7 
Nectandra (non Berg.) Roxb. 26 
Neerija dichotoma Roxb. 286, 
287 
Neojunghuhnia Koord. 785 
insignis Koord. 787 
Neotrewia arborea Elm. 944 
cumingii (M.A.) P. & H. 944 
Nephelium 283 
Nertera dentata Elm. 960 
Neuburgia BI. 293, 295, 296, 
297, 363, 959 
celebica (Koord.)  lLeenh. 
364*, 365, 367, 368, 959 
corynocarpa (A.Gray) Leenh. 
364*, 365, 366, 959 
kochii (Val.) Leenh. 365, 366, 
959 
moluccana (Boerl.) Leenh. 
364*, 365, 367, 368*, 369 
musculiformis (Lamk) Miq. 
367, 368 
rumphiana Leenh. 364*, 365, 
367, 959 
sarcantha (Gilg & Bened.) 
Leenh. 363, 365, 366, 959 
sumatrana (Miq.) Boerl. 369 
tuberculata Bl. 365, 367, 368 
tubiflora Bl. 365, 367, 368 
Neuropeltis Wall. 936, 937 
Neuropeltopsis Ooststr. 936, 937 
alba Ooststr. 937, 938* 
Nicodemia Tenore 296, 336, 340 
diversifolia Tenore 340 
madagascariensis R.N. Parker 
340 
philippinensis Elm. 341 
Nima Ham. ex Juss. 212 
Niota Lamk 199 
commersonii Pers. 200 
globosa Blico 203 
lamarckiana Bl}. 200 
lucida Wall. 200 
pentapetala Poir. 199 
polyandra Buch.Ham. ex W. 
& A. 203 
tetrapetala Poir. 200 
Norrisia Gardn. 293, 295, 296, 
297, 341 
maior Soler. 341, 342* 
malaccensis Gardn. 341, 342*, 
373 
var. major (Soler.) Ridl. 342 
var. malaccense Ridl. 341 
var. pubescens K. & G. 341 
malaccensis (non  Gardn.) 
Merr. 341 
Philippinensis Elm. 341 


Nothocnestis Miq. 280 
sumatrana Miq. 283 
Nothofagus 49 
Nothoprotium Miq. 928 
Niarvala Rheede 68 
Nuxia 294, 295, 296 
Nyctaginaceae 450-468 
Nypa 965 
Nyssaceae 4: 29-31 


Ochranthe Lindl. 49, 51 
Ochthocosmus Bth. 971, 972 
Odyendyea (Pierre) Engl. 198, 
202 
Olacaceae 420 
Oldenlandia brachyphylla Merr. 
384 
Oleaceae 280, 296, 297, 336 
Olus album Rumph. 464 
album insulare Rumph. 464 
Operculina riedeliana (Oliv.) 
Ooststr. 939 
Ophiorhiza L. 375 
mitreola L. 375 
mungos L. 375 
Ophispermum Lour. 6 
Orchipeda sumatrana Miq. 369 
Oreamunoa = Oreomunnea 148 
Oreomunnea Oerst. 145, 148 
Orescia Reinw. 177 
montana Reinw. 181 
Orobanchaceae 294 
Orobanche 294 
Oxybaphus 451 


Pachistima 228 
Paliurus dubius Blco 208 
_ perforatus Blco 208 
Palmerella 140 
Papaveraceae 5: 114-117; 6: 62, 
63 
Paphia Seem. 878 
helenae (F.v.M.) Schltr 879 
stenantha Schltr 884 
viridiflora Schltr 883 
Paracelastrus Miq. 272, 277 
bivalvis Miq. 276 
wallichianus F.N.Williams 279 
Parietales 968 
Parophiorrhiza Clarke 375 
Paropsia vareciformis (Griff.) 
Mast. 944 
Passerina javanica Thunb. 48 
Pedaliaceae 4: 216-221; 5: 557 
Pedicellaria Schrank 99 
pentaphylla Schrank 101 
Pelargonium L’Herit. 445, 448, 
449 
<asperum Ehrh. ex Willd. 
449 
crispum 449 
graveolens Thunb. 449 
radens H.E.Moore 449 
Pellacalyx saccardianus Scort. 
967 


1007 


Pelletiera 173 

Peltanthera Bth. 294, 295, 296, 
953 

Pemphis acidula Forst. 198 


Pentaceras australis Hook. f. 
220 

Pentachondra javanica Zoll. 431 

Pentaphragma 107, 108, 109, 
110 


Pentaphragmataceae 4: 517-528 
Pentaphylacaceae 5: 121-124, 
566 
Pentapterygium Klotzsch 878 
scortechinii K. & G. 879 
Pentaspadon Hook. f. 928 
Peracarpa Hook. f. & Th. 107, 
109, 110 
carnosa (Wall. in 
Hook. f. &. Th. 110 
var. circaeoides 111 
circaeoides Feer 111 
luzonica Rolfe 111 
Pericopsis Thw. 976 
Pernettya 472 
repens (BI.) Zoll. 684 
Pernettyopsis K. & G. 469, 474, 
675 
breviflora (Ridl.) Ridl. 676 
malayana K. & G. 676* 
subglabra K. & G. 676 
Perriera 194 
Perrottetia” HUBiK., non’ DC. 
228-229) 1230.0 231g 252-0240: 
288, 390, 391, 392, 930 


Roxb.) 


alpestris {non (BI.) Loes.] 
Koord. 291 

alpestris (BI.) Loes. 229, 290, 
291 map 


ssp. alpestris 289*, 290 
ssp. moluccana (BI.) Ding 
Hou 289*, 290, 291 
ssp. philippinensis (Vidal) 
Ding Hou 289*, 290, 291 
var. philippinensis Stapf 291 
arborescens (F.v.M.) Loes. 
29] 
arisanensis Hayata 290 
caudata Ridl. 291 
grandifolia Rid]. 291 
lauterbachiana Loes. 291 
f. B macrophylla Loes. 291 
moluccana (Bl.) Loes. 291 
nervosa Ridl. 291 
philippinensis Loes. 291 
racemosa (Oliv.) Loes. 290 
schlechteri Loes. 291 
traumatophylla Merr. & Perry 
291 
Phaleria’ Jack.2; 3,.5,'15; 19* 
ambigua Hook. f. 17 
amboinensis Mertr. 21 
axillaris Elm. 23 
blumei (non Bth.) Hemsl. 18 
calantha Gilg 22, 23 
capitata Jack 2, 14*, 16, 20*, 


1008 


FLORA MALESIANA 


[ser. I, vol. 66 


21 map 
cauliflora Bedd. 20 
coccinea (Gaudich.) 
16, 21 
cumingit F.-Vill. 20 
dubiosa Zoll. 20 
elegans Perry 14*, 16, 17 
laurifolia Hook. f. 2, 17, 18 
var. javanica Val. 17 
longifolia Boer|. 17 
macrocarpa (Scheff.) Boerl. 2, 
144516522" 
neumanni F.v.M. 2 
nisidai Kaneh. 16, 19 
octandra (L.) Baill. 2, 14*, 16, 
17, 18 map 
var. /aurifolia Warb. ex von 
Malm 17 
octandra [non (L.) 
K.Sch. & Hollr. 22 
papuana Warb. ex K.Sch. & 
Laut. 22 
parvifolia Back. 17 
pentecostalis Léandri 4, 16 
perrottetiana (Decne) F.-Vill. 
5, 16, 18 map, 19 
platyphylla Mertr. 21 
revoluta Boerl. 21 
sogerensis S.Moore 16, 21 
sp. Gilg 22 
splendida Val. 18 
subcaudata Merr. & Perry 21 
urens Koord. 20 
vriesii Baill. 21 
wichmannii Val. 22, 23 
zippelti Baill. 21 
Phanrangia Tardieu 231 
Philagonia Bl. 226 
Phylidraceae 4: 5-7 
Phyllocharis Diels 107, 109, 110, 
137, 928 
lamiifolia Wimmer 139 
oblongifolia Diels 137, 139* 
saxicola v.Royen 137, 139*, 
schlechteri Diels 137, 139%, 
subcordata Merr. & Perry 137, 
139*, 
Phyteuma 108 
Phytolaccaceae 4: 228-232; 5: 
557, 6: 109, 451 
Picraena Lindl. 212 
Picramnia 194 
Picrasma Bl. 193, 195, 196, 212 
andamanica Kurz ex Benn. 
214 
denhamii Seem, 214 
javanica Bl. 213*, 214 map 
nepalensis Benn. 214 
philippinensis Elm. 214 
quassioides (D.Don) 
193, 194, 214 
Picroderma laotica Thorel ex 
Gagnep. 226 
Picrolemma pseudocoffea Ducke 
194 


F.v.M. 


Baill. ] 


Benn. 


Picrophloeus Bl. 299, 303 
javanensis Bl). 303, 304 
Picroxylon Warb. 203 
siamense Warb. 205 
Piddingtonia DC. 121 
cyanocarpa Hassk. 132 
montana Mig. 132 
nummularia DC. 134 
palliardii Lehm. 134 
patens Miq. 132 
Pieris (non D.Don 1834) Clarke 
472, 674 
ovalifolia (Wall.) D.Don 675 
Pierreodendron Engl. 198, 202 
Pimela alba Lour. 926 
nigra Lour. 927 
Pimelea Banks & Soland. ex 
Gaertn. 2, 3, 4, 5, 6, 43, 44 
subg. Thecanthes (Wikstr.) 
Gilg 46 
brevituba Fawc. 46 
concreta F.v.M. 45*, 46* map 
cornucopiae Vahl 45*, 46 map 
philippinensis C.B.Rob. 46 
sp. Dammerm. 46 
Piper 468 
Pirola = Pyrola 
Pisonia Plum. ex L. 450, 451, 
457, 462 map 
aculeata L. 450, 457, 459, 460, 
463*, 467 map 
alba Span. 464, 466* 
‘anisophylla Hassk. 467 
artensis (Montr.) Barg.-Petr. 
459, 460 
aruensis Barg.-Petr. 461 
beccariana Barg.-Petr. 463 
brunoniana Endl. 460 
cauliflora Scheff. 460, 462 map 
corniculata Barg.-Petr. 460, 
464 
diandra Pulle 458*, 459, 460, 
463 
excelsa Bl. 460 
excelsa (non BI.) Corner 459, 
464, 465 
fragrans 459 
gammillii Merr. 461 
grandifolia Warb. 463 
grandis R.Br. 197, 223, 459, 
460, 464, 465* map, 466* 
var. sylvestris (T. & B.) 
Heimer! 464 
inermis (non Jacq.) Forst. ex 
Seem. 464, 466, 467 
inermis Jacq. 466, 468 
limonella B\. 467 
lineatipilum C.DC. 468 
longirostris Teysm. & Binn. 
459, 460, 463*, 464 map 
macrophylla (Bojer) Choisy 
464 
major Baill. 462 
membranacea K.Sch. & Hollr. 
468 


micrantha Val. 463 
mitis L. 467 
mooriana F.v.M. 461 
morindifolia R.Br. ex Wight 
464 
miilleriana Warb. 459, 460, 
462 map, 464 
nishimura Koidz. 461 
procera Bertero ex Guill. 464, 
467 
rostrata Warb. 463 
spathiphylla K.Sch. & Laut. 
463 
sylvestris Teysm. & Binn. 464, 
467 
triandra Barg.-Petr. 463 
umbelliflora (Forst.) Seem. 
460, 461* map 
villosa Poir. 467 
Pittosporaceae 5: 345-362; 6: 
373, 960-963 
Pittosporales 63 
Pittosporum Banks ex Gaertn. 
152, 960 
berberidoides Burkill 961, 963 
ferrugineum 12 
var. filarium DC. 11 
filarium Oken 11, 12 
inopinatum Bakker 961, 962* 
moluccanum (Lamk) Miq. 
373, 963 
nubicola Schodde 963 
pentandrum (Blco) Merr. 15, 
963 
pullifolium Burkill 963 
var. globosum Steen. 961, 
963 
pumilum Schodde 961, 962 
ramiflorum (Z. & M.) Zoll. ex 
Mid. 961 
f. macrocarpum Bakker 961 
resiniferum Hemsl. 961 
serrulatum Jack ex Roxb. 963 
sinuatum BI. 961, 963 
var. efuniculare Steen. 961 
tenuivalye Schodde 961 
Pityopus Small 670 
Platycarya 143 
strobilacea 143 
Platycodon grandiflorum (Jacq.) 
DC. 108, 141 
Pleurostylia W. & A. 227, 228, 
232, 287, 391, 392, 930 
heynei W. & A. 288 
opposita (Wall.) Alston 287*, 
288 


wightii W. & A. 288 
var. neocaledonica Loes. 288 

Pleurostylis = Pleurostylia 288 
Plocosperma Bth. 954 
Plumbaginaceae 4: 107-112; 6: 

173, 174, 451 
Plumbaginales 451 
Plumbago 451 
Podandrogyne 100 


Sept. 1972] 


Podocarpus 333 
Podostemaceae 4: 65-68; 6: 
963-964 
Polanisia (non Raf.) DC. 62, 99, 
100 
angulata DC. 102 
chelidonii DC. 102, 103 
dodecandra (L.) DC. 100 
icosandra W. & A. 103 
f. deglabrata Back. 104 
f. typica Back. 103 
viscosa DC. 103, 104 
f. typica Back. 103 
var. @ Back. 103 
var. deglabrata Back. 104 
Polanisia Raf. 62, 99, 100 
graveolens Raf. 99, 100 
Polemoniaceae 4: 195-196 
Polycardia Juss. 932 
Polygalales 63 
Polypremum 293, 295, 296 
Pongamia pinnata (L.) Merr. 
973 
Pongelion Adans. 215, 216 
sect. Ailanthus Pierre 215 
sect. Euailanthus Engl. 215 
sect. Eupongelion Rumph. ex 
Pierre 215 
cacodendron Farwell 220 
calycinum Pierre 218 
excelsum Pierre 219 
fauvelianum Pierre 219 
grandis v.Tiegh. 218 
imberbiflora Pierre 219 
malabaricum Pierre 219 
wightii v.Tiegh. 219, 220 
Pongelion Rheede 219 
Pongelium glandulosum 
220 
vilmorinianum v.Tiegh. 220 
Pontederiaceae 4: 255-261; 5: 
557; 6: 964 
Posoqueria longiflora Aubl. 349 
Potalia 295 
Potaliaceae Hutch. 297 
Potamogetonaceae 157, 171 
Pragmatropa Pierre 245 
Pragmotessara Pierre 245 
japonica Pierre 252 
Pratia Gaudich. 107, 109, 119, 
APM 22. 135 
angulata (Forst.) Hook. f. 122, 
134, 135 
var. arenaria 134 
archboldiana Merr. & Perry 
131 
arenaria Hook. f. 134 
arenosa = arenaria 134 


Pierre 


begoniaefolia = _ begonifolia 
134 

begonifolia (non Lindl.) Hos- 
seus 128 


begonifolia Lindl. 134 
begoniifolia = begonifolia 134 
borneensis Hemsl. 133 


Index to scientific names 


var. grandiflora Stapf 133 
conferta Wimmer 135 
hederacea G.Don 134 

var. elliptica 134 
linnaeoides Hook. f. 134 
montana Hassk. 132 

f. variegata Hochr. 132 


var. cyanocarpa Wimmer 
132 
nummularia A.Br. & Aschers. 
134, 135 


ovata Elm. 128 

papuana §.Moore 134, 135 

podenzanae S.Moore 134 

radicans G.Don 131 

reniformis Kanitz 134 

repens Gaudich. 134 

serpyllacea Presl. 134 

thunbergii G.Don 131 

torricellensis K.Sch. & Laut. 
128 

wollastonii S.Moore 134, 135 

zeylanica Hassk. 134 

Primula Tourn. ex L. 173, 174, 

175, 186, 188 

sect. Callianthae Pax 191 

sect. Candelabra Balf. f. 186, 
191 

sect. Cankrienia (de Vriese) 
Pax 186 

sect. Proliferae Pax 186, 188, 
191 

auricula L. 175 

cortusioides L. 174 

elatior (L.) Hill 173 

farinosa L. 186, 189 

helodoxa Balf. f. 190, 191 

hirsuta All. 188 

hortensis 188 

imperialis Jungh. 189, 190, 191 
var. gracilis Pax 189 

kewensis W.Wats. 175 

khasiana Balf. f. 190, 191 

kuhlii Bl. 190 

malacoides Franch. 192 

minutiflora Forrest 191 

mollis Hook. 174 

obconica Hance 174, 192 

praenitens Ker-Gawl 175, 188, 
192 

prolifera Wall. 173, 186, 187*, 
189*, 190* map 

saxifragifolia O.K. 191 

sieboldii Morren 174 

sinensis Lindl., non Lour. 175, 
188, 192 

smithiana Craib 190, 191 

sumatrana Merr. 190, 191 

umbellata (Lour.) Bentv. 186, 
189, 191*, 192 map 

veris L. 173, 174 
var. elatior L. 192 

Primulaceae 173-192, 964 
subfam. Androsaceae 174 
subfam. Corideae 174 


1009 


subfam. Cyclamineae 174 
subfam. Lysimachieae 174 
subfam. Samoleae 174 
Pristimera grahamii A.C.Smith 
402 
indica A.C.Smith 401 
Proteaceae 5: 147-206, 566; 6: 
368, 965 
Protium javanicum Burm. f. 917 
macgregorii (F.M.Bailey) 
Leenh. 917 
Pseudais Decne 15 
coccinea Decne 21 
Pseudogardneria Racib. 296 
Pseudospigelia 296 
Psilaea Miq. 27 
dalbergioides Miq. 27, 982 
Psychotria L. 387, 959 
Pteridophyta 171 
Pterilema Reinw. 145 
aceriflorum Reinw. 152 
Pteris wallichiana Agardh 187* 
Pterocarya 143 
Punicaceae 4: 226-227 
Pyrola Tourn. ex L. 469, 472, 
474, 671 
Japonica Klenze ex Alefeld 
673 
ssp. coreana Andres 673 
Japonica (non Klenze ex Ale- 
feld) Steen. 671 
sumatrana Andres 671, 672* 
Pyrolaceae 472, 473 
Pyroleae 472 
Pyrospermum Miq. 280 
calophyllum Miq. 282 
Pyrus granulosa Bertol. 878 


Quassia L. 968, 969 
harmandiana (Pierre) 
969 


Noot. 


Randia cochinchinensis (Lour.) 
Merr. 309 

Rapuntium Miller 121 
affine Pres] 128 
alsinoides Pres] 127 
angulatum Pres] 134 
arenarioides Pres] 129 
caespitosum Presl 130 
campanuloides Pres 130 
chinense Pres| 130 
coloratum Pres! 123 
haenkeanum Pres| 136 
leschenaultianum Pres! 123 
longiflorum Miller 140 
longifolium Presl 136 
nicotianaefolium Pres| 123 
nummularium Pres| 134 
pyramidale Pres} 123 
radicans Pres| 130 
reinwardtianum Presl 129 
reniforme Pres] 134 
roseum Pres| 123 
succulentum Presl 128 


FLORA MALESIANA 


[ser. I, vol. 68 


1010 
trialatum Pres] 129 
wallichianum Presl 123 
zeylanicum Presl 128 
Reissantia Hallé 390, 391, 400 
cassinoides (DC.) Ding Hou 
402 

ferruginea (King) Ding Hou 
402 

grahamii (Wight) Ding Hou 
402, 403* 

indica (Willd.) Hallé 401 

Resedaceae 62, 63 

Restionaceae 5: 416-420, 569 

Retzia Thunb. 295, 297, 954 

Rex amaroris Rumph. 221 

Rhamnaceae 49, 280, 284 

Rhamphocarya Kuang 143 

Rhesa = Bhesa 282, 283 

Rhizophora L. 965 
apiculata Bl. 966 
harrisonii Leechm. 974 
mucronata Lamk 965, 966 
stylosa Griff. 966 

Rhizophoraceae 5: 429-493; 6: 
361, 965-967, 974 

Rhododendron L. 469, 470, 471, 
472, 473, 474, 475 map 
subg. Anthodendron (Rchb.) 

Endl. ex Wils. & Rehd. 661, 
943 
sect. Anthodendron (Rchb.) 
Endl. 661, 943 
subg. Anthodendron (non 
Wils. & Rehd.) Sleum. 661, 
662 
sect. Anthodendron Sleum. 
662 
subg. Azaleastrum Planch. 
480, 665 
sect. Choniastrum Franch. 
480, 665, 666 map 
subg. Eurhododendron (Endl.) 
Koehne 656 
sect. Eurhododendron Endl. 
657 
subg. Hymenanthes  (BI.) 
Koch 480, 656 
sect. Hymenanthes 480, 657 
subsect. Irrorata Sleum. 
480, 657 map 
subg. Hymenanthus = Hyme- 
nanthes 656 
subg. Lepidorrhodion Koehne 
480 


subg. Pentanthera (G.Don) 
Pojarkova 480, 661, 943 
sect. Pentanthera 480, 661, 
943 
subg. Pseudovireya Clarke 480 
subg. Rhododendron 479, 
480, 943 
sect. Discovireya Sleum. 480 
sect. Eurhododendron Endl. 
480 


sect. Hadranthe Schltr 497, 
501, 547 

sect. Hapalanthe Schltr 606 

sect. Linnaeopsis Schltr 568 

sect. Schistanthe Schltr 501, 
606 

sect. Vireya (Bl.) Copel. f. 
475, 477*, 478*, 479, 480, 
943 
subsect. Albovireya 

Sleum. 479, 537, 538 


map 
subsect. Astrovireya 
Sleum. 580 


subsect. Euvireya Copel. 
1, ss, ASS SPS De 
567 map, 606 
ser. Buxifolia Sleum. 
538, 568, 580 
ser. Citrina Sleum. 568, 
579 
ser. Javanica Sleum. 
536, 568, 606 
ser. Linnaeoidea 
Sleum. 567, 568 
ser. Saxifragoidea 
Sleum. 567, 574 
ser. Stenophylla Sleum. 
568, 576 
ser. Taxifolia Sleum. 
568, 575 
subsect. Leiovireya 
Copel. f. 606 
subsect. Linearanthera 
Copel. f. 580 
subsect. Linnaeopsis 
(Schltr) Sleum. 568 
subsect. Malayovireya 
Sleum. 477*, 478*, 479, 
527 map 
subsect. Malesia Copel. f. 
580 
subsect. Phaeovireya 
Sleum. 478*, 479, 501, 
511, 527 map 
subsect. Pseudovireya 
(Clarke) Sleum. 476, 
478*, 479, 480 map, 
527, 536 
subsect. Schizovireya 
Sleum. 606 
subsect. Siphonovireya 
Sleum. 476, 479, 497, 
527 map 
subsect. Solenovireya 
Copel. f. 476, 479, 527, 
547 map 
sect. Zygomorphanthe 
Schltr 480, 501, 537, 576, 
580, 606 
ser. Malayanum Copel. 
ii SPT) 
subg. Tsutsutsi (G.Don) Po- 
jarkova 480, 661 
sect. Tsutsia — Tsutsutsi 


662 

sect. Tsutsugi = Tsutsutsi 
662 

sect. Tsutsusi = Tsutsutsi 
662 


sect. Tsutsutsi 480, 657 map, 
662 

subg. Vireya (Bl.) Clarke 480 

abietifolium Sleum. 581, 590, 
591 

acrocline Sleum. 652 

acrophilum Merr. & Quis. 
581, 595 

acuminatum Hook. f. 528, 
534* 

adinophyllum Merr. 481, 490* 

aequabile J.J.S. 538, 539 

agathodaemonis J.J.S. 497, 
499, 505 

agathodaemonis (non J.J.S. 
1913) J.J.S. 1936 500 

album BI. 538, 539, 626 

album (non BI.) Ridl. 539 

album (non BI.) Zoll. 540 

alternans Sleum. 581, 592 

alticolum Sleum. 582, 597 

amabile Sleum. 548, 559 

anagalliflorum Wernh. 568 

andersonii Rid. 536 

angiense J.J.S. 652 

angulatum J.J.S. 607, 616 

apoanum Stein 528, 531 

araiophyllum Balf. f. & W.W. 
Sm. 661 

archboldianum Sleum. 548, 
555 

arenicolum Sleum. 539, 546 

arfakianum Becc. 581, 609, 
610, 648 

armitii F.M.Bailey 548, 557 

asparagoides Wernh. 643 

asperrimum Sleum. 509 

asperum J.J.S. 476, 478, 517, 
Ryle 

astrapiae Foerster 515 

atjehense Sleum. 472, 657, 
658*, 659* 

atropurpureum Sleum. 570, 
582, 602*, 603 

aurigeranum Sleum. 609, 642 

baenitzianum Laut. 608, 627 

bagobonum Copel. f. 580, 586 

banghamiorum (J.J.S.) Sleum. 
581, 596 

basirotundatum J.J.S. 626 

beccarii Sleum. 607, 622, 626 

beyerinckianum Koord. 477*, 
478, 511 
var. longipetiolatum J.J.S. 

511 

bloembergenii Sleum. 608, 633 

bodenii Wernh. 497 

brachygynum Copel. f. 609, 
610, 650 

brachypodarium Sleum. 549, 


Sept. 1972] 


Index to scientific names 


561, 562* 
brassii Sleum. 470, 582, 600 
brevipes Sleum. 609, 642 
brevitubum J.J.S. 607, 622 
breynii Planch. 663 
brookeanum Low ex Lindl. 
475, 537, 610, 655 
var. brookeanum 655, 656 
var. cladotrichum Sleum. 
655, 656 
var. extraneum Sleum. 655, 
656 
var. gracile (Low ex Lindl.) 
Sleum. 655, 656 
brookeanum (non Low ex 
Lindl.) Stapf 653 
bryophilum Sleum. 477*, 509 
bullifolium Sleum. 522 
burmanni G.Don 662 
buruense J.J.S. 608, 632 
buxifolium Low ex Hook. f. 
582, 603 
var. buxifolium 603, 604 
var. robustum Sleum. 591, 
604, 612 
caespitosum Sleum. 568, 569*, 
ai /i be 
calceolarioides Wernh. 634 
callichilioides Wernh. 628 
var. minor Wernh. 628 
calodon Ridl. 591 
calosanthes Sleum. 582, 605 
candidapiculatum Wernh, 572 
carrii Sleum. 548, 557 
carringtoniae (non F.v.M.) 
Lane-Poole 500 
carringtoniae F.v.M. 548, 552, 
558 
var. maius J.J.S. 554 
carstensense Wernh. 548, 553 
caryophyllum Hayata 664 
celebicum (Bl.) DC. 606, 615 
cernuum Sleum. 538, 545 
cerochitum Balf. f. & Forr. 
661 
chamaepitys Sleum. 548, 552 
chevalieri Dop 475 
christi Foerster 606, 612, 614 
var. loniceroides Schltr 612 
christianae Sleum. 471, 608, 
637 
chrysopeplon Sleum. 512 
ciliilobum Sleum. 482, 494 
cinchoniflorum Sleum. 497, 
498* 
cinerascens Sleum. 540, 561 
citrinum (Hassk.) Hassk. 579 
var. albiflorum Miq. 579 
var. citrinum 579 
var. discoloratum Sleum. 
580 
citrinum (non Hassk.) Miq. 
580 
clementis Merr. 625 
coelorum Wernh. 568, 573 


coenenii J.J.S. 652 
commonae Foerster 581, 587, 
588*, 589* 
commutatum Sleum. 608, 629 
comparabile Sleum. 608, 636 
comptum C.H.Wright 543 
var. comptum 538, 543, 
544* 
var. trichodes 538, 544 
coniferum Wernh. 543 
convexum Sleum. 651 
copelandii Merr. 567 
coriifolium Sleum. 612 
cornu-bovis Sleum. 580, 585 
correoides J.J.S. 538, 543 
corruscum Ridl. 659 
crassifolium Stapf 607, 626 
crassineryium Ridl. 626 
cruttwellii Sleum. 548, 554, 
555+ 
culminicolum F.v.M. 580, 
581, 608, 610, 651 
var. angiense (J.J.S.) Sleum. 
476, 524, 651, 652 
var. culminicolum 597, 651, 
652 
var. nubicola (Wernh.) 
Sleum. 580, 581, 651, 652 
cuneifolium (non Stapf) 
Rendle 495 
cuneifolium (non Stapf) Ridl. 
496 
cuneifolium Stapf 496 
var. subspathulatum (non 
Ridl.) Merr. 496 
var. subspathulatum Rid. 
586 
curranit Merr. 583 
curviflorum J.J.S. 606, 613 
cuspidellum Sleum. 608, 610, 
629 
cyatheicolum Sleum. 520 
cyrtophyllum Wernh. 481, 491 
dasylepis Schltr 512 
delicatulum Sleum. 501, 507 
var. delicatulum 507 
var. lanceolatoides Sleum. 
509 
devrieseanum Koord. 515 
ssp. astrapiae Foerster 515 
devrieseanum vel aff. Stonor 
505 
devriesianum = devrieseanum 
515 
dianthosmum Sleum. 477*, 
526 
dielsianum Schltr 510, 511 
var. dielsianum 510 
var. stylotrichum Sleum. 
510 
discolor Warb. 615 
disterigmoides Sleum. 568, 
569*, 573 
doctersii J.J.S. 643 
dubium K. & G. 659 


1011 


durionifolium Becc. 528, 529, 


531 
durionifolium (non Becc.) 
Stapf 531 
edanoi Merr. & Quis. 549, 564 
elegans Ridl. 591 
elongatum BI. 565 
englerianum Koord. 608, 609, 
628 
ericoides (non Low ex Hook. 
f.) Burtt 496 
ericoides Low ex Hook. f. 481, 
491 
var. ericoides 491* 
var. silvicolum Sleum. 492 
erosipetalum J.J.S. 481, 485 
extrorsum J.J.S. 501, 504 
eymae Sleum. 501, 503 
fallacinum Sleum. 528, 530*, 
531 
filamentosum Wernh. 560 
flavoviride J.J.S. 608, 637 
fortunans J.J.S. 528, 535 
franssenianum J.J.S. 614 
frey-wysslingii J.J.S. 580, 582 
fuchsii Sleum. 609, 648 
fuchsioides Schltr 486 
fuchsioides aff. Stonor 575 
fuscum Bl. 532 
galioides J.J.S. 586 
gardenia Schltr 514 
gaultheriifolium J.J.S. 477%, 
481, 484 
var. expositum Sleum. 485 
var. gaultheriifolium 484 
gibbsiae J.J.S. 652 
gilliardii Sleum. 522 
giulianettii Laut. 538, 546 
glabrifilum J.J.S. 635 
glabriflorum J.J.S. 606, 614 
goodenoughii Sleum. 549, 561 
gorumense Schltr 634 
gracile (non Low ex Lindl.) 
Becc. 629 
gracile Low ex Lindl. 656 
gracilentum F.v.M. 568, 570* 
gregarium Sleum. 651 
habbemae Koord. 497 
habbemai = habbemae 497 
haematophthalmum Sleum. 
522, 524 
hameliiflorum Wernh. 481, 
486 
hansemannii Warb. 633 
hatamense Becc. 580, 584, 
606, 608 
hatamense (non Becc.) Sleum. 
652 
helwigii (non Warb.) 
Koord. 499 
hellwigii Warb. 501, 504 
helodes Sleum. 581, 594 
herzogii Warb. 470, 497, 500* 
himantodes Sleum. 528 
var. himantodes 528 


1012 


FLORA MALESIANA 


var. lavandulifolium Sleum. 
529 
hirtolepidotum J.J.S. 607, 618 
hooglandii Sleum. 501, 507, 
576* 
hybridogenum Sleum. 528, 
537 
impositum J.J.S. 609, 649 
impressopunctatum J.J.S. 607, 
617 
incommodum Sleum. 497, 500 
inconspicuum J.J.S. 470, 
477*, 538, 582, 598 
inconspicuum aff. Stonor 569 
indicum (L.) Sweet 662, 663 
f. variegatum (BI.) DC. 663 
intranervatum Sleum. 610, 
654 
inundatum Sleum. 497, 499 
invasorium Sleum. 481, 488 
jasminifiorum (non Hook.) 
F.-Vill. 531 
jasminiflorum Hook. 476, 478, 
537, 548, 549, 565 
var. copelandii (Merr.) 
Sleum. 566, 567 
var. heusseri (J.J.S.) Sleum. 
565, 566 
var. jasminiflorum 566 
var. maculatum Ridl. 566 
var. oblongifolium Sleum. 
566, 567 
var. punctatum Ridl. 528, 
537, 566 
jasminiflorum (non Hook.) 
Koord. 579 
jasminiflorum (non Hook.) 
Merr. 567 
jasminiflorum (non Hook.) 
Ridl. 563, 566 
Jasminiflorum (non Hook.) 
Sarasin 560 
javanicum (BI.) Benn. 471, 
478*, 537, 607, 610, 624* 
var. citrinum Hassk. 624 
var. gymnocarpum Hochr. 
624 
var. javanicum 625, 626 
var. schadenbergii (Warb.) 
Sleum. 625 
var. teysmannii (Miq.) K. 
& G. 528, 537, 625 
var. tubiflorum Hook. 640 
jJavanicum (non Benn.) Clarke 
620 
jJavanicum (non Benn.) F.-Vill. 
641 
jJavanicum (non Benn.) Koord. 
615 
javanicum (non Benn.) Malm 
650 
jJavanicum (non Benn.) Steen. 
621 
keditii Sleum. 606, 612 
kemulense J.J.S. 607, 623 


keysseri Foerster 651 
kjellbergii J.J.S. 630 
klossii Rid]. 666 map, 667 
kochii Stein 609, 641 
konori Becc. 477*, 478*, 515, 
SUG7, 51873 S19N652 
korthalsii Miq. 657, 660 
lacteum Stapf 551 
laetum J.J.S. 476, 478, 517, 
521, 608, 609, 638* 
laetum (non J.J.S. 1914) J.J.S. 
1936 643 
lagunculicarpum J.J.S. 538, 
542 
lami J.J.S. 582, 599 
lampongum Mig. 538, 544 
lanceolatum Ridl. 609, 610, 
647 
langbianense Chey. ex Dop 
659 
laoticum Dop 667 
lauterbachianum Foerster 633 
ledifolium (Hook.) G.Don 662 
leptanthum F.v.M. 478, 512, 
Sls 
leptobrachion Sleum. 608, 631 
leptomorphum Sleum. 581, 
593 
leptopeplum Balf. f. & Forr. 
661 
leucobotrys Rid]. 666 
leucogigas Sleum. 477*, 478, 
608, 632 
leytense Merr. 609, 649 
var. leytense 649 
var. loheri (Copel.  f.) 
Sleum. 649 
lindaueanum Koord. 481, 486 
var. bantaengense J.J.S. 486 
var. cyclopicum Sleum. 486 
var. latifolium J.J.S. 486 
var. lindaueanum 486 
var. psilacrum Sleum. 486 
lineare Merr. 528, 535 
linearifolium S. & Z., non 
Poir. 943 
var. macrosepalum 
(Maxim.) Makino 662, 
663 
linnaeoides Schltr 568 
lobbii Veitch 640 
loboense Copel. f. 607, 619 
lochae F.v.M. 475, 636 
loerzingii J.J.S. 607, 616 
loheri Copel. f. 649 
lompohense J.J.S. 608, 632 
var. grandifolium J.J.S. 633 
longiflorum Lindl. 475, 478, 
537, 566, 608, 609, 640 
var. bancanum Sleum. 641 
var. heusseri J.J.S. 566 
var. longiflorum 640 
loranthiflorum Sleum. 549, 
564 
lowei = lowii 654 


[ser. I, vol. 68 


lowii Hook. f. 610, 654 


lowii (non Hook. f.) F.v.M. 
628 
lukiangense Franch. 661 
luraluense Sleum. 580, 608, 
636 
lussoniense Rendle 583 
luteosquamatum Sleum. 582, 
598, 599 
maboroense Schltr 627 
macgregoriae F.v.M. 478, 522, 
608, 609, 610, 633, 634*, 
652 
var. glabrifilum (J.J.S.) 
Sleum. 634, 635 
var. macgregoriae 634, 635 
var. mayrii (J.J.S.) Sleum. 
634, 635 
macrosepalum Maxim. 662, 
663, 943 
macrosiphon Sleum. 548, 552 
magnificum Sleum. 515 
maius (J.J.S.) Sleum. 548, 554 
malayanum Jack 477*, 478*, 
528, 532, 586,193 p97, 017 
var. axillare J.J.S. 533 
f. axillare 532, 533 
f. latifolium Sleum. 532, 
533 
f. ovatum Sleum. 532, 
533 
var. infrapilosum Sleum. 
532, 533 
var. malayanum 528, 532 
var. pilosifilum Sleum. 528, 
532, 539 
var. pubens Sleum. 528, 
5325559 
malayanum (non Jack) Koord. 
540 
malindangense Merr. 496 
maxwellii Gibbs 609, 644 
mayrii J.J.S. 635 
megalostigma F.v.M. 628 
melantherum Schltr 520 
meliphagidum J.J.S. 482, 493 
microphyllum J.J.S. 568, 571 
mindanaense Merr. 610, 650 
minimifolium Wernh. 483, 571 
mjoébergii Merr. 529 
molle (Bl.) G.Don 661 
mollianum Koord. 610, 652 
moszkowskii Schltr 643 
moulmainense Hook. 666 
map, 667*, 668 
moultonii Ridl. 607, 623 
mucronatum (BI.) G.Don 662 
multicolor Miq. 607, 621 
var. curtisii Henslow 621 
multicolor (non Miq.) S.Moo- 
re 580 
multinervium 
549, 550* 
murudense Merr. 626 
murudense (non Merr.) J.J.S. 


Sleum. 547, 


Sept. 1972] 


Index to scientific names 


618 
muscicola J.J.S. 568, 572 
myrsinites Sleum. 576, 578 
nanophyton Sleum. 492 
var. nanophyton 481, 492 
var. petrophilum Sleum. 
482, 493 
natalicium Sleum. 547, 549 
neriifolium Schltr 501, 506 
nervulosum Sleum. 646 
var. exuberans Sleum. 610, 
646 
var. nervulosum 609, 646* 
nieuwenhuisii J.J.S. 580, 586 
ningyuenense Hand.-Mazz. 
659 
nitens Sleum. 593 
nodosum C.H.Wright 651 
nortoniae Merr. 528, 533 
nubicola Wernh. 652 
nummatum J.J.S. 481, 484 
obscurinervium Merr. 656 
obscurum Sleum. 528, 535 
oliganthum Sleum. 548, 556 
ombrochares Balf. f. & Ward 
661 
opulentum Sleum. 519 
oranum J.J.S. 643 
orbiculatum Rid]. 549, 563, 
608, 609 
oreadum Wernh. 548, 560 
oreites Sleum. 481, 485 
var. chlorops Sleum. 485 
var. oreites 485 
orion Ridl. 490 
var. aurantiacum Ridl. 640 
oxycoccoides Sleum. 568, 
369s 57/3 
oxyphyllum Franch. 666 
pachycarpon Sleum. 606, 614 
pachystigma Sleum. 607, 616 
papuanum Becc. 581, 594, 
5955 
papuanum (non Becc.) C.H. 
Wright 546 
partitum J.J.S. 647 
parvulum Sleum. 568, 569*, 
573 
pauciflorum K. & G. 581, 591 
var. calodon (Ridl.) Sleum. 
591 
var. pauciflorum 591 
pennivenium Balf. f. & Forr. 
660 
perakense K. & G. 481, 488, 
536 
perplexum Sleum. 607, 626 
phaeochitum F.v.M. 511 
Phaeochitum (non F.v.M.) 
Wright 526 
phaeochristum Sleum. 476, 
478, 523 
phaeopeplum Sleum. 476, 
478, 517, 519, 644 
phaeops Sleum. 525 


planecostatum Sleum. 580, 
586 

pleianthum Sleum. 548, 556 

pneumonanthum Sleum. 549, 
563 

podocarpoides Schltr 579 

polyanthemum Sleum. 609, 
641 

poremense J.J.S. 610, 652 

porphyranthes Sleum. 582, 
599 

prainianum Koord. 501, 505 

proliferum Sleum. 538, 540 

protandrum Sleum. 497, 498 

psammogenes Sleum. 581, 
591 

pseudobuxifolium Sleum. 581, 
587 

pseudomurudense Sleum. 607, 
618 

pseudonitens Sleum. 581, 593 

pseudotrichanthum Sleum. 
548, 559 

psilanthum Sleum. 511 

pubigermen J.J.S. 580, 583, 
597 
var. banghamiorum J.J.S. 

596 

pubitubum Sleum. 548, 558 

pudorinum Sleum. 538, 545 

pulleanum J.J.S. 483 
var. maiusculum Sleum. 

481, 484 

var. pulleanum 481, 483 

purpureiflorum J.J.S. 576, 578 

pusillum J.J.S. 568, 572 

pyrrhophorum Sleum. 527, 
582, 597 

quadrasianum Vidal 482, 494 

banahaoense Copel. f. 495 

davaoense Copel. f. 495 

halconense Copel. f. 495 

lutea H.J.Lam 496 

marivelesense Copel. f. 

496 

monodii H.J.Lam 496 

negrosense Copel. f. 495 

pulogense Copel. f. 495 

f. pulogense (non Copel. f.) 
H.J.Lam 496 

f. rubra H.J.Lam 496 

var. angustissimum Sleum. 
494, 496 

var. borneense J.J.S. 494, 
497 

var. cuneifolium (Stapf) 
Copel. f. 494, 496, 478* 

var. davaoense (Copel. f.) 
Sleum. 494, 495 

var. intermedium Merr. p.p. 
495 

var. intermedium Merr. s. 
str. 494, 496 

var. malindangense (Merr.) 
Copel. f. 494, 496 


1013 


var. marivelesense (Copel. 
f.) Sleum. 494, 496 
var. quadrasianum 494, 495 
var. rosmarinifolium (Vi- 
dal) Copel. f. 494, 495 
var. selebicum J.J.S. 494, 
496 
var. villosum J.J.S. 494, 
496 
radians J.J.S. 548, 560 
var. minahasae Sleum. 560 
var. radians 560 
rappardii Sleum. 519, 525 
rarelepidotum J.J.S. 620 
var. ootrichum Sleum. 609, 
621 
var. rarelepidotum 607, 620 
rarum Schltr 501, 507, 508* 
renschianum Sleum. 610, 650 
retivenium Sleum. 607, 610, 
653 
retrorsipilum Sleum. 549, 565 
retusum (BIl.) Benn. 470, 471, 
477*, 481, 820* 
f. angustifolia Miq. 482 
var. epilosum Sleum. 483 
var. macranthum Hochr. 
482 
var. retusum 482 
var. trichostylum Sleum. 
483 
retusum (non Benn.) F.-Vill. 
494 
retusum (non Benn.) Steen. 
566 
retusum (non Benn.) Wernh. 
486 
revolutum Sleum. 501, 503 
rhodochroum Sleum. 521 
rhodoleucum Sleum. 548, 558 
rhodopus Sleum. 608, 627 
rhodosalpinx Sleum. 548, 551 
rhodostomum Sleum. 581, 
592 
ripense Makino 663 
ripleyi Merr. 581, 596 
var. basitrichum Sleum. 
581, 596, 597 
var. cryptogonium Sleum. 
527, 596, 597 
var. ripleyi 596 
robinsonti Rid]. 607, 620, 621 
rosendahlii Sleum. 608, 635 
rosmarinifolium Vidal 495 
rubellum Sleum. 526 
rubrobracteatum Sleum. 582, 
605 
rubropilosum Hayata 664, 665 
rugosum Low ex Hook. f. 590, 
606, 611, 612 
var. coriifolium (Sleum.) 
Sleum. 581, 590, 612 
var. rugosum 611 
ruttenti J.J.S. 547, 550 
saavedranum Diels 512 


salicifolium Becc., non BI. 
609, 646 
salicifolium Bl. 621 
sarasinorum Warb. 625 
saruwagedicum Foerster 481, 
487* 
var. alpinum Sleum. 487 
saxifragoides J.J.S. 574*, 575, 
589* 
sayeri Sleum. 606, 615 
scabridibracteum Sleum. 608, 
609, 639* 
scrabrum G.Don 662 
scarlatinum Sleum. 582, 601 
schadenbergii (non Warb.) 
Merr. 619, 641 
schadenbergti Warb. 625 
schizostigma Sleum. 482, 493 
schlechteri Laut. 608, 631, 632 
schoddei Sleum. 523 
schultzei Schltr 512 
scortechinii K. & G. 481, 490, 
536 
seimundii J.J.S. 481, 490 
seranense = seranicum 619 
seranicum J.J.S. 607, 617, 619 
sessilifolium J.J.S. 607, 622 
sheilae Sleum. 581, 590 
siamense Diels 666 
simsii Planch. 662, 663, 665 
simulans Sleum. 582, 600 
sinense (Lodd.) Sweet 661 
solitarium Sleum. 523 
spathulatum Ridl. 481, 487, 
536 
spectabile Merr. 625 
spondylophyllum F.v.M. 520 
stapfianum Hemsl. ex Prain 
548, 551 
stelligerum Sleum. 501, 506 
stenaulum Balf. f. & Forr. 666 
stenophyllum Hook. f. 576*, 
Sif 
var. angustifolium J.J.S. 577 
stolleanum Schltr 510 
stonori Sleum. 587 
stresemannii J.J.S. 610, 653 
suaveolens Sleum. 563 
subcordatum Becc. 608, 630 
subcrenulatum Sleum. 582, 
605 
subpacificum Sleum. 549, 565 
subsessile Rendle 662, 663, 
664*, 665* 
var. baucoense Copel. f. 
664, 665 
subuliferum Sleum. 582, 601 
subulosum Sleum. 576, 578 
sumatranum Merr. 609, 647 
superbum Hort. ex Lavallée 
505 
superbum Sleum. 501, 505 
syringoideum Sleum. 548, 549, 
553 
taxifolium Merr. 575 


FLORA MALESIANA 


[ser. I, vol. 68 


taxoides J.J.S. 481, 483 

teysmannii (non Miq.) Hend. 
620 

teysmannii Miq. 626 

thaumasianthum Sleum. 515 

torricellense Schltr 635 

toverenae F.v.M. 515 

toxopei J.J.S. 610, 655 

trichanthum Sleum. 559 

triumphans Yersin & Cheval. 
475 

truncicolum Sleum. 525 

tuba Sleum. 548, 557 

tuberculiferum J.J.S. 509 

tubiflorum Low ex Lindl. 640 

tubiflorum (non Bl.) Mor. 540 

tubiflorum Reinw. in Bl. 532 

tubulosum Reinw. 615 

uliginosum J.J.S. 638 

ultimum Wernh. 582, 600 

undulaticalyx J.J.S. 648 

vanderbiltianum Merr. 481, 
488* 

vandeursenii Sleum. 580, 581, 
585 

vanvuurenii J.J.S. 471, 608, 
630 

variolosum Becc. 536 
var. andersonii 

Sleum. 528, 536 

var. variolosum 528, 536 

velutinum Becc. 611 

versteegiil J.J.S. 539, 546 

verticillatum (non Low ex 
Lindl.) Becc. 567 


(RidI.) 


verticillatum (non Low) 
Koord. 560 
verticillatum (non Low ex 


Lindl.) Low ex Hook. f. 604 
verticillatum Low ex Lindl. 
606, 610 
f. velutinum (Becc.) Sleum. 
606, 607, 611 
f. verticillatum 609, 610 
verticillatum (non Low ex 
Lindl.) Vidal 583 
vidalii Rolfe 580, 583, 584* 
villosulum J.J.S. 606, 613 
vinicolor Sleum. 528, 529 
vinkii Sleum. 482, 493 
vitis-idaea Sleum. 581, 592 
vonroemeri Koord. 633 
warianum Schltr 514 
wentianum Koord. 608, 610, 
628 
wentianum all. Stonor 612 
westlandii Hems]. 666 
whiteheadii Rendle 583 
wilhelminae Hochr. 528, 536 
williamsii Merr. ex Copel. f. 
607, 619 
wollastonii Wernh. 628 
womersleyi Sleum. 603, 568, 
569* 
wrayi K. & G. 657, 659, 660* 


var. elliptica Ridl. 659 
var. minor Ridl. 659 
wrightianum Koord. 478, 582, 
604 
var. cyclopense J.J.S. 604 
var. insulare Sleum. 604 
var. ovalifolium J.J.S. 600, 
604 
var. piliferum J.J.S. 594 
var. wrightianum 604 
xanthopetalum Merr. 607, 617 
yelliotii Warb. 538, 542, 599 
yelliottii = yelliotii 542 
zippelti Bl. 579 
zoelleri Warb. 476, 478, 519, 
609, 635, 639, 643, 644*, 
645*, 652 
zollingeri J.J.S. 538, 540, 541 * 
var. latifolium J.J.S. 540 
zollingerianum = zollingeri 
540 
Rhoeadales 62 
Rhoipteleaceae 143 
Rhus cacodendron Ehrh. 220 
javanica L. 211 
cf. linguata W.Slis 291 
vernicifera DC. 220 
Rigiolepis Hook. f. 754 
bigibba (J.J.S.) J.J.S. 761 
borneensis Hook. f. 756 
caudatifolia (Merr.) J.J.S. 765 
endertii J.J.S. 758 
filiformis J.J.S. 758 
korthalsii J.J.S. 763 
lanceifolia Ridl. 755 
lanceolata (Bl.) J.J.S. 761, 762 
f. elliptica J.J.S. 762 
f. marapiensis J.J.S. 762 
f. sumatrana (Miq.) J.J.S. 
762 
lancifolia = lanceifolia 755 
leptantha (Miq.) J.J.S. 764 
f. elliptica J.J.S. 764 
lobbii Ridl. 755 
macrophylla J.J.S. 757 
moultonii (Merr.) J.J.S. 763 
salicifolia J.J.S. 757 
sulcata (Ridl.) J.J.S. 758 
uniflora (J.J.S.) J.J.S. 757 
Rinorea 963 
glabra O.K. 235 
lanceolata (Roxb.) O.K. 237, 
963 
Rockia Heimer] 457, 459 
Rosaceae 472, 878 
Rosales 49 
Rourea Aubl. 934 
subg. Palliatus Leenh. 
sect. Palliatus 934 
acropetala Pierre 934 
mimosoides (Vahl) Planch. 
934 
f. mimosoides 934 
minor (Gaertn.) Leenh. 934, 
935 


Sept. 1972] 


Index to scientific names 


ssp. microphylla (Hook. & 


Arn.) Vidal 934 
ssp. minor 934 
ssp. monadelpha 
Vidal 934 
oligophlebia Merr. 934 
ovale (Schellenb.) Leenh. 934 
pinnata (Merr.) Veldkamp 
934 
prainiana Talbot 934 
Roureopsis Planch. 933 
sect. Taeniochlaena Leenh. 
934 
acutipetala (Miq.) Leenh. 934 
ssp. borneensis (Schellenb.) 
Leenh. 934 
asplenifolia Schellenb. 933, 
934 
emarginata (Jack) Merr. 934 
incurva Pierre 933 
stenopetala (Griff.) Schellenb. 
933 
Roydsia Roxb. 95, 97 
floribunda Planch. ex Hook. f. 
97 
parviflora (non Griff.) King 
99 


(Roxb.) 


philippinensis Turcz. 97 
scortechinii King 99 

Rubiaceae 107, 195, 294, 297, 
309, 336, 349, 373, 379, 387, 
401 
subfam. Coffeoideae 959 
tribe Hedyotideae 297 
tribe Psychotrieae 387 

Rubiales 294, 297 

Ruppia 171 

Ruppiaceae 171 

Rutaceae 69, 194, 195, 206, 214, 
220, 221, 226, 969 

Ruthiella Steen. 928 
oblongifolia (Diels) Steen. 928 
saxicola (v.Royen) Steen. 928 
schlechteri (Diels) Steen. 928 
subcordata (Merr. & Perry) 

Steen. 928 


Sagittaria sagittifolia L. 
ssp. /eucopetala (Miq.) Har- 
tog 915 
trifolia L.915 
Salacia L. 231, 390, 391, 392, 404 
map, 421, 930, 932 
alternifolia Scort. 421 
amentacea Ridl. 408 
amplifolia Merr. ex Chen & 
How 412, 413 
hartletti Ridl. 420 
beccarii Ridl. 417 
blepharophora Ding Hou 405, 
408 
buddinghii Scheff. 412 
campanuloidea King 408 
castaneifolia Ridl. 406, 413 
celebica Bl. 412 


cerasiformis Teysm. & Binn. 
| 


chinensis L. 392, 406, 419 
map, 421, 932 

coromandeliana 
Binn. 421 

cymosa Elm. 405, 407, 408, 
416* 

diandra Miq. 410 
f. lanceolata Miq. 411 

disepala (C.T.White) 
Hou 420 

erythrocarpa K.Sch. 404, 405, 
411 

euphlebia Merr. 406, 418 

evonymiflora Zipp. ex Mid. 
419 

exsculpta Korth. 406, 417 

flavescens Kurz 412 
var. dumosa King 412 

forsteniana Miq. 404, 405, 410, 
932 

grandiflora Kurz 406, 413, 932 
var. longifolia (Hook. f.) 

King 413 

griffithi Laws. 400 

integrifolia Merr. 415 

intermedia Ding Hou 
411* 

jJavanensis Bl. 400 

kalahiensis Korth. 406, 420 

kamputensis Pierre 412 

klossii Ridl. 408 

korthalsiana Miq. 405, 406, 
407 map, 420 

kraemeri Ding Hou 410 

lanceolata Teysm. & Binn. 
408 

latifolia Wall. ex Laws. 419 

laurifolia Stapf 405, 406, 417 

lawsoni King 412 

ledermannii (Loes. ex Harms) 
Ding Hou 405, 410 

leucoclada Ridl. 405, 406, 415 

litseifolia Ridl. 415 

littoralis Back. 419 

lobbii Laws. 417 

longifolia Hook. f. ex Laws. 
413 

longipedicellata) Ding Hou 
406, 413 

macrocarpa Korth. 412 

macrophylla BI. 405, 412 map, 
416* 
var. angustifolia Miq. 412 

maingayi Laws. 406, 416%, 
417 

marginata Ding Hou 405, 406, 
413 

megasperma Ridl. 417 

melitocarpa Bl. 408 

minutiflora Rid]. 420 

naumanni Engl. 419 

nitidissima Merr. 406, 418 

oblonga (non Wall.) Rendle 


Teysm. & 


Ding 


405, 


412 
oblonga Wall. 420 
oblongifolia Bl. 405, 408 
f. latior Miq. 408 
ovalis (non Korth.) Koord.- 
Schum. 419 
ovalis Korth. 406, 415, 416* 
ovalis Laws., non Korth. 412 
papuana (Loes.) Ding Hou 
405, 408, 409* 
patens Decne 419, 420 
perakensis King 400 
Philippinensis Merr. 406 
polyantha Korth. 414 
prinoides (Willd.) DC. 419, 
420, 421 
var. macrophylla (Bl.) King 
412, 419 
var. timorensis Span. 419 
radula (non G.Don) Dietr. ex 
Hassk. 406, 407 
radula G.Don 407 
roxburghii (non Wall.) Vidal 
415 
roxburghii Wall. ex Laws. 420 
rubra Laws. 417, 418 
scortechinii King 413, 414 
sinensis (non L.) Bleco 406 
sinensis Gmelin 418, 420 
socia Craib 419 
sororia Mig. 405, 409*, 420, 
932 
sp. 280 
subalternifolia Merr. & Perry 
405, 407, 416* 
subscandens Elm. 420 
triplinervis Llanos 421 
venosa Ding Hou 406, 417 
verrucosa Wight 406, 414%, 
932 
vimeria Wall. 392, 406, 418, 
421 
viridis Craib 408 
wenzelii Merr. 405, 411, 416* 
wrayi King 418 
Salacicratea Loes. 404 
australis Loes. 420 
brassti A.C.Smith 409 
diandra (Miq.) A.C.Smith 410 
glandulosa A.C.Smith 405, 409 
kraemeri (non Loes.) Kaneh. 
419, 420 
kraemeri Loes. 410 
ledermannii Loes. 410 
papuana Loes. 408 
sarasinorum Harms 410 
sororia A.C.Smith 409 
Salacighia Loes. 930 
Salicaceae 5: 107-110 
Salvadoraceae 4: 224-225 
Samadera Gaertn. 193, 194, 198, 
199, 226, 968 
brevipetala Scheff. 201 
glandulifera Presl 201 
indica Gaertn. 199 


1016 


FLORA MALESIANA 


[ser. I, vol. 66 


var. brevipetala_ (Scheff.) 
Back. 199 
var. papuana Laut. 199 
madagascariensis Juss. 200 
mekongensis Engl. 201 
pentapetala G.Don 201 
tetrapetala Wall. 200 
Samandura L. ex Baill. 199 
indica Baill. 201 
madagascariensis Perr. de la 
Bathie 201 
mekongensis Pierre 201 
Sambucus 51 
Samolus 173, 174 
valerandi L. 173, 192 
Sanango G.S.Bunting & J.A. 
Duke 295, 296 
Santaloides ovale Schellenb. 934 
Santiria Bl. 917, 920 
apiculata Benn. 920, 921 
var. apiculata 921 
var. rubra (Ridl.) Kalkman 
921 
conferta Benn. 920, 921 
grandiflora Kalkman 920, 921 
griffithii (Hook. f.) Engl. 920, 
921 
laevigata Bl. 920, 921 
megaphylla Kalkman 920, 921 
mollis Engl. 920 
nervosa H.J.Lam 919, 921 
oblongifolia Bl. 920, 921 
ridleyi H.J.Lam 920 
rubiginosa BI. 920, 921 
var. rubiginosa 921 
tomentosa BI. 920 
Sapindaceae 49, 194, 226 
Sapindales 49, 53, 933 
Sapindus saponaria L. 226 
Sapotaceae 229 
Saprosma 387 
Sarawakodendron 
930 
filamentosa Ding Hou 931*, 
932 
Sarcocaulon 445 ’ 
rigidum Schinz 445 
Sarcosperma uittienii H.J.Lam 
967 
Sarcospermataceae 4: 32-34; 6: 
967 
Sarcotheca pinnata Merr. 934 
Saururaceae 4: 47-48 
Saxifragaceae 173, 174, 280, 472 
Scaevola L. 949, 951 
lobelia Murr. 951 
micrantha Pres! 951, 952 
oppositifolia R.Br. 951, 952 
pauciflora Leenh. 951, 952 
plumierii 223 
sericea Vahl 951 
taccada (Gaertn.) Roxb. 949, 
951 


Ding Hou 


verticillata Leenh. 951, 952* 
Scaphocalyx spathacea Ridl. 944 


Schmidelia conferta Blco 282 
Schmidellia = Schmidelia 282 
Scolopia Schreb. 943 
kermodei C.E.C.Fischer 943 
Scopolia (non Jacq.) L. f. 35, 37 
composita L. f. 38 
Scrophulariaceae 136, 294, 295, 
296, 297, 960, 965 
Scrophulariales 297 
Scutinanthe Thw. 921 
brevisepala Leenh. 919, 921 
brunnea Thw. 921 
Scyphostegia Stapf 967, 968* 
borneensis Stapf 967, 968* 
Scyphostegiaceae 5: 297-299; 6: 
967 
Scyphostrychnos 296 
Selenocera Zipp. ex Span. 375 
secundiflora Zipp. ex Span. 
375 
Sericolea Schltr 914 
Shorea selanica BI. 154 
Simaba Aubl. 198, 202 
Simarouba 194, 198 
cedron Planch. 194 
glauca 194 
Simaroubaceae 193-226, 968— 
972 
subfam. Alvaradoideae 195, 
971 
subfam. Irvingioideae 193, 
194, 195, 970, 971, 972 
subfam. Kirkioideae 195, 971 
subfam. Picramnioideae 195, 
971 
subfam. Simarouboideae 195 
subfam. Surianioideae 193, 
194 
tribe Simaroubeae 198 
subtribe Eurycominae 205 
subtribe Simaroubinae 195, 
205 
Simarubopsis Engl. 202 
Simirestis 390 
Siphonodon Griff. 227, 228, 229, 
230, 231, 390, 391, 392, 394, 
968 
australe Bth. 230 
celastrineus Griff. 229, 231, 
394, 395*, 396* map 
var. acuminatissima Merr. 
395 
var. integrifolia Tardieu 412 
var. subglobosa Merr. 395 
peltatus Ding Hou 395*, 397, 
932 
pyriformis Merr. 395, 396 
var. parvifolius Merr. 395 
Sirioides Rumph. 355 
alter Rumph. 355 
Solanaceae 297 
Solenopsis 140 
Solenospermum Zoll. 231, 262, 
264 
apiculatum Ridl. 272 


aquatile Ridl. 270, 271 
Javanicum Zoll. 269 
ledermannii Loes. 271 
littorale Loes. 261 
macranthum Loes. 264, 268 
micranthum Loes. 268 
oblongifolium Loes. 270 
pallidum Loes. 268 
paucinervium Loes. 270 
torricellense Loes. 271 
var. opacum Loes. 271 
toxicum Loes. 270 
Solmsia 3 
Sonneratia L.f. 973 
acida L. f. 
var. mucronata Miq. 975 
alba J.Smith 973*, 974*, 975 


alba * ovata 974* 
caseolaris) (.) Engel )973*, 
974*, 975 


ovata Backer 974*, 975 
Sonneratiaceae 4: 280-289; 5: 
557; 6: 973 
Soulamea Lamk 193, 196, 221 
amara Lamk 193, 197, 221, 
222*, 223 map 
soulameoides (A.Gray) Noote- 
boom 221 
terminalioides Baker 221 
Sparganiaceae 4: 233-234 
Specularia Heist. ex Fabr. 928 
speculum-veneris (L.) Caruel 
928 
Speirema Hook. f. & Th. 121 
montanum Hook. f. & Th. 132 
Sphaerodiscus Nakai 245 
cochinchinensis Nakai 248 
Sphenoclea 107, 108, 109, 110 
zeylanica Gaertn. 928 
Sphenocleaceae 4: 27-28. See 
Campanulaceae 
Spigelia L. 295, 296, 297, 377 
anthelmia L. 376*, 378, 960 
Spilanthes urens 223 
Spinifex 221 
Spiraeaceae 195 
Stackhousiaceae 4: 35-36 
Staphylea 49, 52 
Staphyleaceae 49-59, 214, 230 
Steironema Raf. 177 
Stellera L. 28 
chamaejasme L. 28 
passerina L. 28 
Stenanthera R.Br. 424 
Stenocarpus moorei F.v.M. 428 
Stixis Lour. 61, 63, 95, 97 map 
fasciculata var. borneensis 
Heine 97 
floribunda (Planch. ex Hook. 
f.) Pierre 97 
obtusifolia (Hook. f. & Th.) 
Pierre 97 
ovata (Korth.) Hall. f. 97 
ssp. fasciculata (King) Ja- 
cobs 97 map 


Sept: 1972] 


ssp. ovata (Korth.) Hall. 
f. 96*, 97 map 

philippinensis (Turcz.) Merr. 
95, 97 map 

scortechinii (King) Jacobs 97 
map, 99 

suaveolens (Roxb.) Pierre 95, 
97 


Struthiola 4 


Strychnos L. 205, 293, 294, 295, 

296, 297, 343, 361, 954 

sect. Booneae Duvign. 346 

sect. Breviflorae Progel 345 

sect. Brevitubae A.W.Hill 345, 
346, 955 

sect. Floribundae Duvign. 346 

sect. Intermediae Progel 345 

sect. Lanigerae A.W.Hill 345, 
346, 955 

sect. Ligustroides Duvign. 346 

sect. Longiflorae Progel 345 

sect. Penicillatae A.W.Hill 
34529346, 352*, 357,955 

sect. Sambae Duvign. 346 

sect. Strychnos 955 

sect. Tubiflorae A.W.Hill 345, 
346 

acuminata Wall. ex DC. 355 

aenea A.W.Hill 958 
var. acuminata A.W.Hill 
958 

andamanensis A.W.Hill 351 

angustiflora Bth. 345, 955, 
958, 959 

arborea A.W.Hill 358, 360 

armata A.W.Hill 358, 359 

axillaris Colebr. 346, 347, 351, 
352%, 958.) 360)" 3615955, 
958 

balansae A.W.Hill 347, 349 

bancroftiana F.M.Bailey 355, 
356 

barbata A.W.Hill 355 

beccariti Gilg 347, 349 

borneensis Leenh. 346, 347, 
355, 356, 357*, 955, 958 

celebica Koord. 355 

cenabrei Merr. 358 

chloropetala A.W.Hill 358, 
359 

cinnamophylla Gilg & Bened. 
355 

colubrina Auct. non L. 346, 
BAPE SOs 55941 3945.11355, 
356, 358, 361, 955, 958 

confertiflora Merr. & Chun 
351 

curtisii King & Gamble 345, 
346, 351, 353, 355, 955, 958 

cuspidata A.W.Hill 347, 349 

dubia A.W.Hill 355 

flavescens K. & G. 346, 347, 
357, 955, 958 

forbesii A.W.Hill 354*, 355, 
356 


grandis Wall. 361 

hirsutiflora A.W.Hill 351 

horsfieldiana (non Miq.) 
Koord.-Schum. 351 

horsfieldiana Miq. 358, 359 

hypogyna C.B.Clarke 355 

ignatii Bergius 297, 345, 346, 
347, 348*, 351, 361, 955, 957 

ignatii (non Berg.) Vidal 355 

impressinervis A.W.Hill 351, 
358, 359 

kawbet A.W.Hill 358, 359 

kerrii Hill 955, 959 

kerstingti Gilg & K.Sch. 355 

krabiensis A.W.Hill 347, 349 

lanata A.W.Hill 345, 346, 355, 
955, 958 

lanceolaris Miq. 346, 347, 357, 
955, 958 

laurina DC. 350 

laurina (non DC.) Heine 356 

laurina (non DC.) K. & V. 351 

laurina Wall. ex DC. 355 
var. thorelii A.W.Hill 355 

ledermannii Gilg & Bened. 
346, 347, 357, 955, 958 

leuconeura Gilg & Bened. 355 

ligustrina Bl. 350 

lucida R.Br. 293, 346, 349, 
350 map, 355, 955, 957 

luzonensis Elm. 346, 347, 355, 
358, 955, 958 

maingayi C.B.Clarke 346, 
350, 351, 955, 957, 958 
ssp. borneensis Leenh. 958 
var. fructuosa Clarke 347, 

350 

malaccensis Bth. 358, 360 

melanocarpa Gilg & Bened. 
361, 955, 959 

merrillii A.W.Hill 355, 356 

minahassae Koord. ex Boerl. 
355 

minor Dennst. 955, 958 

monosperma Miq. 358 

mucronata A.W.Hill 358 

multiflora Bth. 355, 356 

muricata Kosteletzky 350 

myriantha Gilg & Bened. 355 

nux-blanda A.W.Hill 349, 959 

nux-vomica Auct. non L. 958 

nux-vomica L. 297, 345, 346, 
348*, 349, 955, 957 
f. depauperata Miq. 349 

oleifolia A.W.Hill 346, 347, 
353, 355, 955, 958 

oophylla Gilg & Bened. 358, 
360 

ovalifolia Wall. 347, 349, 351 

ovata A.W.Hill 346, 351, 
352 SS gS OW, 

ovata (non Hill) Merr. 358 

palembanica Miq. 358, 359 

panayensis A.W.Hill 351 

penicillata A.W.Hill 358, 360, 


Stylidium 


1017 


361 
philippensis Blco 957 
philippinensis Blco 347, 349, 

957 
pilgeriana Gilg 358 
plumosa A.W.Hill 358, 359 
polytoma Gilg & Bened. 358, 

360 
polytoma (non Gilg & Bened.) 

Kaneh. & Hatus. 355 
polytrichantha Gilg 345, 346, 

353, 357, 955, 958 
potatorum (non L. f.) F.-Vill. 

355 

var. multiflora Vidal 355 
pseudotieuté A.W.Hill 347 
psilosperma F.v.M., 358, 360 
pubescens C.B.Clarke 358, 360 

var. scortechinii K. & G. 

358 
pycnoneura Gilg & Bened. 355 
quadrangularis A.W.Hill 346, 

347, 356, 955, 958 
quintuplinervis A.W.Hill 358, 

359 
ridleyi K. & G. 346, 347, 360, 

955, 958 
robinsonti A.W.Hill 358, 360 
roborans A.W.Hill 350 
rufa C.B.Clarke 358, 361, 955, 

958 
schmidtii Gilg 358, 359 
scortechinii A.W.Hill 358, 

359 
septemnervis C.B.Clarke 355 

var. imberbis A.W.Hill 355 
silvicola A.W.Hill 355 
similis A.W.Hiull 355 
sp. Hill 358 
sp. Merr. 957 
sp. 1 Hill 351 
sp. a & B Koord.-Schum. 351 
spireana Dop 349 
tesseroidea A.W.Hill 358, 359 
thorelii Pierre ex Dop 361, 

955, 958 
tieute Leschen. 347, 349 
vanprukii Craib 357, 955, 958 
villosa A.W.Hill 346, 351, 355, 

955, 958 
viridiflora A.W.Hill 358 
wenzelii Merr. 358, 360 


Stylidiaceae 4: 529-532; 5: 564; 


6: 108, 109, 976 

androsaceum O. 

Schwarz 976 

ericksonae Willis 976 

fluminense Erickson & Willis 
976 

inconspicuum Sloot. 976 

javanicum Sloot. 976 

pedunculatum R.Br. 976 


Styphelia J.E.Smith 423, 424 


subg. Cyathodes (Lab.) Drude 
425, 426 map, 433 


1018 


FLORA MALESIANA 


[ser. I, vol. 68 


subg. Leucopogon (R.Br.) 
Drude 423, 425, 426 map 
sect. Styphelia 423 

‘abnormis (Sond.) J.J.S. 425, 
433 

abscondita J.J.S. 425 

acuminata (R.Br.) Spr. 423, 
425, 432 

arfakensis Kaneh. & Hatus. 
443 

brassii Sleum. 425, 433, 434* 

brevistyla Moore 434 

carstensensis Wernh. 441 

celebica J.J.S. 439 

cornifolia Rudge 437 

culminis Wernh. 440 

cuspidata (R.Br.) Spr. 432 

cymbulae (Lab.) Spr. 427 

dekockii J.J.S. 440 

forbesii Sleum. 425, 432 

gjellerupti J.J.S. 438 

hookeri (Sond.) J.J.S. 429 

javanica (de Vriese) J.J.S. 423, 
425, 430*, 431*, 432 

juniperina (Forst.) Pers. 434 

lamii J.J.S. 443 

lancifolia J.5.S. 433 

learmonthiana Gibbs 439 

leptospermoides (R.Br.) Spr. 
433 

malaica Jack 426 

malayana (Jack) Spr. 
425, 426, 943 


423, 


var. malayana 425, 426, 
427* 
var. novoguineensis 425, 


427, 428*, 429* 
moluccana J.J.S. 433 
montana F.v.M. 429 

var. hookeri 429 
nesophila (DC.) Sleum. 432 
nubicola Wernh. 442 
nutans J.J.S. 441 
var. arfakensis J.J.S. 441 
obovata J.J.S. 432 
obovata Malm 429 
obtusifolia J.J.S. 429 
var. hypoleuca J.J.S. 429 
oxycedrus Lab. 434 
papuana (C.H.Wright) Koord. 
443 
Philippinensis Merr. 429 
pungens Koord. 431 
rapae Sleum. 434 
spicata J.J.S. 429 
suaveolens (Hook. f.) Warb. 
423, 425, 428, 429* 
trilocularis J.J.S. 429 
var. guinquelocularis J.J.S. 

429 
trochocarpoides F.v.M. 437 
vandewateri Wernh. 429 
vannouhuysii J.J.S. 440 
wetarensis J.J.S. 432 

Styracaceae 4: 49-56; 6: 976 


Sulamea = Soulamea 221 

Suregada Rottl. 944 

Suriana L. 193, 194, 195, 196, 
969 
maritima L. 

map, 223 

Surianaceae 195, 196 

Sycopsis Oliver 952 
dunnii Hemsl. 952 


193 aL Gaal 97 


Tabernaemontana corymbosa 
Roxb. 315 
Taeniochlaena Hook. f. 933 
Talauma 228 
Tamaricaceae 968 
Tamaricales 63 
Tapiscia 49 
Tayotum nigrescens Blco 373 
Telanthera bettzickiana Regel 
916 
manillensis Walp. 915 
Terminalia L. 932 
catappa 325 
crassifolia Exell 922 
darlingii Merr. 933 
macrantha Rojo 932 
polyantha Pr. 48 
zollingeri Exell 933 
Ternstroemia megacarpa Elm. 
336 
penangiana Choisy 336 
Tetramyxis Gagnep. 226 
Theaceae 472 
Thecanthes Wikstr. 44 
cornucopiae Wikstr. 46 
Thibaudia coriacea (Bl.) BI. 828 
cuneifolia Bl. 847 
elliptica Bl. 873 
floribunda Bl. 872 
laurifolia Bl. 871 
lucida Bl. 814 
myrtoides Bl. 812 
rosea Jungh. 872 
singalensis Korth. ex Boerl. 
762 
varingiaefolia Bl. 815 
Thylachium lucidum Banks ex 
DC. 92 
Thymelaea 28 
Thymelaeaceae 4: 349-365 (Go- 
nystyloideae); 6: 1-48, 976 
subfam. Aquilarioideae 1, 3, 
982 
tribe Microsemmatidae 3 
subfam. Gilgiodaphnoideae 3 
subfam. Gonystyloideae 1, 2, 
3, 976-982 
subfam. Thymelaeoideae 3, 
982 
Thymelaeales 451 
Tiliaceae 4, 203 
Timeroyea Montrouz. 457 
Tonsella chinensis (L.) Spreng. 
419 
prinoides Willd. 418 


Toona sinensis (Juss.) Roem. 
220 
Torrenticola Domin 963 
Tovariaceae 62 
Toxicodendron altissimum Mill. 
220 
Toxina Norona 954 
Trachymene Rudge 983 
novoguineensis (Domin) Buw. 
984 
tripartita Hoogl. 983*, 984 
Trapaceae 98? 
Trianthema triquetra Rottl. ex 
Willd. 915 
Tricalysia tinagaoensis Elm. 23 
Triceros Lour. 51 
cochinchinensis (non Lour.) 
Mor. 56 
Trichadenia 39 
philippinensis Merr. 944 
Trichilia 194 
connaroides (W. & A.) Bent- 
velzen 
f. connaroides 226 
f. glabra Bentvelzen 226 
Trientalis 174 
Trigoniaceae 4: 58-60 
Tripterygium Hook. f. 229, 230, 
231 
wilfordii Hook. f. 229, 230 
Triscaphis Gagnep. 49, 212, 214 
kerrii Gagnep. 214 
Tristellateia Thouars 960 
Tristicha trifaria (Bory ex 
Willd.) Spreng. 963 
Triumfetta 223 
Trochisandra Bedd. 280 
indica Bedd. 283 
Trochocarpa R.Br. 423, 424, 
434, 436 map 
subg. Pseudocyathodes 
Sleum. 437, 443 
subg. Trochocarpa 436, 437 
arfakensis (Kaneh. & Hatus.) 
Sleum. 443, 444* 
bellendenkerensis Domin 437 
celebica (J.J.S.) Steen. 439 
dekockii (J.J.S.) H.J.Lam 
435*, 440* 
dispersa Sleum. 442 
gjellerupii (J.J.S.) H.J.Lam 
438, 439* 
lamii H.J.Lam 443 
laurina (R.Br. ex Drude) R. 
Br. 423, 437, 438* 
learmonthiana (Gibbs) H.J. 
Lam 439 


nubicola  (Wernh.) Sleum. 
442, 443 

nutans (J.J.S.) H.J.Lam 435*, 
441*, 442*, 443 


papuana (Wright) Sleum. 443 
vannouhuysii H.J.Lam 441 
Tubiflorae 294, 297 
Turneraceae 4: 235-238 


Sept. 1972] 


Index to scientific names 


Turpinia Vent. 49, 51, 56 map 
arguta 55 
borneensis (Merr. & Perry) v. 
d.Linden 50*, 53, 54,55, 56 
brachypetala (Schltr) v.d. 
Linden 52, 54, 59 
cochinchinensis (Lour.) Merr. 
58 
formosana Nakai 55 
glaberrima Merr. 56 
gracilis Nakai 56 
grandis v.d.Linden 52, 53, 54, 
55, 58 
indochinensis Merr. 55 
laxiflora Ridl. 53, 54, 55, 57 
lucida Nakai 58 
malabarica Gamble 59 
montana (BI.) Kurz 50*, 52, 
994, 55256 
a@ genuina Kurz 55 
f. arborescens Hochr. 56 
f. scandens Hochr. 56 
var. borneensis Merr. & 
Perry 56 
nepalensis 52, 55, 56, 57, 58, 
59 
nitida Merr. & Perry 52, 54, 
58 
ovalifolia Elm. 50*, 53, 54, 58 
pachyphylla Merr. 58 
papuana Harms 59 
papuana Merr. & Perry 59 
parva K. & V. 56 
parviflora Craib 53, 56 
pentandra (Schltr) v.d.Linden 
52, 54, 55, 59 
pomifera (Roxb.) DC. 50*, 
52, 54, 58 
var. sphaerocarpa (non 
Hassk.) King 57, 58 
robusta Craib 58 
simplicifolia Merr. 52, 53, 54, 
55 
sp. Merr. 58 
sphaerocarpa Hassk. 50*, 52, 
53; 54, 57, 58, 59 
var. pubescens v.d.Linden 
57 
stipulacea v.d.Linden 50%, 
SIP 52953254: 55 
trifoliata Ridl. 58 
unifoliata Merr. & Chun 53, 
5 


versteeghii Merr. 59 


Typha L. 982 


angustata Bory & Chaubard 
983 

angustifolia L. s.]. 982, 983 

domingensis Pers. 982, 983 

latifolia L. 983 

orientalis Pr. 982, 983 


Typhaceae 4: 242-244; 6: 982 


Umbelliferae 4: 113-140, 595; 5: 
555; 6: 194, 983-984 


1019 


Urophyllum streptopodium 
Wall. 373 

Usteria 295, 296, 954 

Utania G.Don 299, 301, 309 
morindaefolia G.Don 311 


Vacciniaceae 473 
Vaccinium L. 469, 470, 471, 472, 
473, 474, 741, 746, 878 
sect. Bracteata Nakai 747, 
791, 792 map 
sect. Euepigynium Schltr 791 
sect. Galeopetalum (J.J.S.) 
Sleum. 747, 748 map, 753 
sect. Neojunghuhniana 
(Koord.) Sleum. 747, 753 
map, 785 
sect. Nesococcus Copel. f. 791 
sect. Oarianthe Schltr 747, 
765 map 
sect. Pachyanthum Sieum. 
747, 748 map 
sect. Rigiolepis (Hook. f.) 
J.J.S. 747, 753 map, 754 
subg. Galeopetalum J.J.S. 753 
absconditum J.J.S. 767, 775 
acrobracteatum K.Sch. 470, 
793, 796, 800, 868* 
acuminatissimum (non Miq.) 
Merr. 756 
acuminatissimum Miq. 755, 
761 
f. acuminatissimum 761 
f. ellipticum (J.J.S.) Sleum. 
761, 762 
f. javanicum Miq. 762 
f. leptantha = leptanthum 
f. leptanthum (Miq.) Vuyck 
764 
f. marapiense (J.J.S.) 
Sleum. 761, 762 
f. sumatranum Miq. 762 
var. singalense J.J.S. 762 
acuminatissimum (non Mig. s. 
str.) Ridl. 762 
acutissimum F.v.M. 831 
var. acutissimum 795, 831 
var. pilosistylum Sleum. 
796, 831 
acutissimum (non 
Warb. 830 
adenanthum Sleum. 868 
adenopodum Sleum. 755, 765 
adenotrichum Sleum. 796, 846 
adenurum C.E.C.Fischer 840 
agusanense Elm. 793, 807 
aitapense Sleum. 795, 800, 833 
albicans Sleum. 795, 832 
var. albicans 797, 832 
var. pseudopsammogenes 
Sleum. 832, 833 
var. pubens Sleum. 832, 
833 
alvarezii Merr. 798, 800, 857 
var. alvarezii 857 


F.v.M.) 


var. moisense Copel. f. 858 
ambivalens Sleum. 796, 799, 
801, 847 
amblyandrum F.v.M. 768, 783 
var. amblyandrum 766, 783, 
784 
var. maiusculum Sleum. 783 
var. pungens Sleum. 783, 
784* 
ambyandrum = amblyandrum 
783 
amphoterum Sleum. 793, 797, 
798, 848 
amplexicaule J.J.S. 794, 799, 
861 
amplifolium F.v.M. 748, 751 
var. amplifolium 751, 752, 
753 
var. giganteum Sleum. 751, 
752 
var. oblongum Sleum. 751, 
752 
var. stabilipes Sleum. 751, 
752 
ampullaceum Sleum. 792, 804 
andersonii Sleum. 755, 763 
angiense Kaneh. & Hatus. 
798, 800, 856 
angulatum J.J.S. 798, 799, 856 
angustilimbum Merr. 870 
apiculatum Sleum. 800, 869 
apoanum Merr. 895 
apophysatum Sleum. 797, 850 
appendiculatum Schltr 794, 
826, 861 
ardisiflora = ardisiiflorum 843 
ardisiiflorum Ridl. 843 
ardisioides Ridl. 843 
ardisioides Wernh., non Ridl. 
863 
arenarium Sleum. 784 
artum J.J.S. 798, 854 
atrescens Sleum. 753 
aucupis Sleum. 794, 822 
auriculifolium Sleum. 792, 
805 
bancanum (non Mig. s.str.) 
Clarke 842 
bancanum (non Mig. s:.str.) 
K. & G. 842 
bancanum (non Miq.) Koord. 
847 
bancanum Midq. 796, 803, 840, 
841* 
var. bancanum 841, 842%, 
843 
var. kemulense J.J.S. ex 
Sleum. 841, 843 
var. kunstleri (K. & G.) 
Sleum. 841, 842 
var. tenuinervium 
841, 842, 843 
bancanum (non Mig.) Ridl. 
854 
banksii Merr. 797, 849 


JS. 


1020 


FLORA MALESIANA 


[ser [fyolie® 


banksii (non Merr. 1906) 
Merr. 850 
barandanum Vidal 793, 806 
var. barandanum 807 
var. cagayanense Copel. f. 
807 


var. hutchinsonii (Merr.) 
Copel. f. 807 
barbatum J.J.S. 768, 781 
bartlettii Merr. 800, 867 
benguetense Vidal 797, 801, 
875 
besagiense J.J.S. 796, 838 
bigibbum J.J.S. 754, 761 
blepharocalyx (non Schltr) 
Lane-Poole 876 
blepharocalyx Schltr 801, 876 
blumeanum Niedenzu 872 
bodenii Wernh. 766, 767, 774 
borneense (non W.W.Sm.) 
Anderson 763 
borneense W.W.Sm. 754, 756 
var. borneense 756 
var. poianum (J.J.S.) 
Sleum. 756 
brachycladum Sleum. 766, 770 
brachygyne J.J.S. 794, 824 
brachytrichum Sleum.796, 847 
bracteatum Thunb. 747, 791, 
793, 801 
brassii Sleum. 799, 864 
var. brassii 864, 866 
var. madarum Sleum. 866 
breviflos Ridl. 842 
brevipedunculatum J.J.S. 799, 
863 
buxifolium Hook. f. 839 
buxoides J.J.S. 777 
calelanum Elm. 895 
camiguinense Merr. 795, 836 
capillatum Sleum. 794, 799, 
866 
capillipes Sleum. 754, 757 
cardiophorum Sleum. 786, 
789 
carneolum Sleum. 831 
var. carneolum 795, 831 
var. nesopbilum Sleum. 
795, 832 
caudatifolium Merr. 765 
caudatum Warb. 797, 801, 
875 
cavendishioides Sleum. 798, 
800, 801, 858 
centrocelebicum Sleum. 792, 
803 
var. centrocelebicum 803 
var. maius Sleum. 803 
ceramense Sleum. 767, 775 
cercidifolium J.J.S. 794, 827 
chionostomum Sleum. 776 
ciliatipetalum J.J.S. 777 
claoxylon J.J.S. 798, 859 
clementis Merr. 795, 796, 836* 
coelorum Wernh. 768, 784 


collivagum Sleum. 772 
contractum Sleum. 794, 821 
convallariiflorum J.J.S. 794, 
826, 861 
convexifolium J.J.S. 766, 772 
cordifolium Stapf 794, $27 
coriaceum Hook. f. 792, 795, 
796, 838 
coriaceum (Bl.) Miq. 828 
cornigerum Sleum. 796, 837 
corymbiferum Miq. 878 
costerifolium Sleum. 796, 801, 
845 
costeroides Merr. 742 
crassiflorum J.J.S. 767, 777 
crassiflorum (non J.J.S. 1912 
& 1914) J.J.S. 774 
crassifolium Andr. 746 
crassistylum Sleum. 799, 863 
crenatifolium Sleum. 797, 849 
cruentum Sleum. 798, 831, 
857, 877 
cryptodon Sleum. 776 
culminicolum Wernh. 766, 
767, 774 
cumingianum Vidal 797, 798, 
850 
var. cumingianum 850, 851 
var. igorotorum Copel. f. 
851 
var. marivelesense Copel. 
f. 851 
var. pyriforme (Merr.) Co- 
pel. f. 851 
var. tayabasense Copel. f. 
850 
cuneifolium (BI.) Mig. 796, 
847 
var. acutum Miq. 847 
cyclopense J.J.S. 773 
f. cyclopense 767, 773 
f. glabrum Sleum. 768, 773 
var. arfakense J.J.S. 773 
cyrtodon Miq. 872, 873 
daphniphyllum Schltr 794, 
826, 861 
debilescens Sleum. 797, 852 
decorum Ridl. 852 
decumbens J.J.S. 767, 776 
dempoense Fawc. 810 
dempoense (non Sp.Moore) de 
Voogd 811 
densifolium J.J.S. 766, 769 
dialypetalum J.J.S. 752*, 753 
dictyoneuron Sleum. 798, 800, 
858 
var. dictyoneuron 798, 858 
var. koébrense Sleum. 859 
var. oreophilum Sleum. 859 
dipladenium Sleum. 755, 763 
disterigmoides Sleum. 777 
dominans Sleum. 471, 793, 
809 
dubiosum J.J.S. 747, 793, 808 
eburneum Ridl. 843 


elegans Elm. 795, 829 

ellipticum (Bl.) Miq. 815, 873 
var. macrocalyx J.J.S. 873 

elliptifolium Merr. 795, 835 

endertii (J.J.S.) Masam. 758 

endertii J.J.S. 795, 835 

epiphythicum Merr. 793, 806 

erythrinum Hook. 815 

euanthum Bl. 872 

eugenoides Sp.Moore 811 

evanidinervium Sleum. 768, 
784 

eymae Sleum. 786, 790, 791* 

fastigiatum Sp.Moore 872 

filiforme (J.J.S.) Sleum. 754, 
758 

filipes Schltr 796, 844 

finisterrae Schltr 766, 769 

fissiflorum Sleum. 747, 748 

flagellatifolium Copel. f. 755, 
765 

floribundum (BI.) Miq. 872 

forbesii Fawe. 810 

foxworthii Copel. f. 837, 847 

fraternum Sleum. 793, 798, 
859 

gitingense Elm. 792, 803 

gjellerupii J.J.S. 798, 801, 871 

glabrescens K. & G. 792, 804 

glandellatum Sleum. 794, 813 

globosum J.J.S. 768, 778, 781 
var. latifolium J.J.S. 779 

goodenoughii Sleum. 795, 829 

gracile J.J.S. 797, 799, 851 

gracilipes Sleum. 801, 874 

gracillimum J.J.S. 801, 877 

grandibracteatum Schltr 793, 
809 

habbemae Koord. 798, 855 
var. habbemae 855 
var. parvifolium J.J.S. 855 
var. pluriglandulosum J.J.S. 

798, 855 

haemanthum Sleum. 769 

haematochroum Sleum. 766, 
767, 770 

halconense Merr. 798, 799, 
860 

hasseltii Miq. 854 
var. sabuletorum Ridl. 854 

hatamense Becc. 766, 770 

hatamense (non Becc.) Koord. 
782 

helenae F.v.M. 879 

hellwigianum Sleum. 795, 830 

henrici Sleum. 754, 759 

hispidulissimum Wernh. 786, 
789 

hooglandii Sleum. 793, 799, 
862 

horizontale Sleum. 796, 839, 
840* 

hosei Merr. 873 

hutchinsonii Merr. 807 

igneum J.J.S. 767, 775 


Sept. 1972] 


Index to scientific names 


1021 


igorotorumCopel.f. ex Elm.851 
ilocanum Merr. 810 
imbricans J.J.S. 786, 789 
inconspicuum J.J.S. 768, 781 
indutum Vidal 792, 793, 804 
ingens Sleum. 747, 748, 749* 
insigne (Koord.) J.J.S. 786, 
787 
irigaense Merr. 798, 801, 859 
jagori Warb. 800, 870 
Javanicum Hook. 873 
kemulense Sleum. 754, 758 
keysseri Schltr ex Diels 748, 
750* 
var. acutatum Sleum. 751 
var. keysseri 750 
kjellbergii J.J.S. 791, 793, 802 
korinchense Ridl. 794, 811 
var. korinchense 792, 811* 
var. losirense Sleum. 812 
korthalsii (J.J.S.) Masam. 763 
korthalsii Mig. 795, 828 
kostermansii Sleum. 786 
kunstleri K. & G. 842 
kunstleri (non K. & G.) Ridl. 
843 
lageniforme (non J.J.S.) Ka- 
neh. & Hatus. 868 
lageniforme J.J.S. 799, 800, 
862 
lanaense Merr. 745 
lanceifolium (Ridl.) Sleum. 
754, 755 
lanceolatum (B\.) J.J.S. 761 
f. ellipticum Sleum. 762 
f. marapiense (J.J.S.) 
Sleum. 762 
f. sumatrana (Mig.) Sleum. 
762 
lancifolium = lanceifolium 
ies) 
latissimum J.J.S. 794, 796, 821 
laurifolium (BI.) Mig. 471, 
746, 801, 871*, 878 
var. arborescens O.K. 873 
var. ellipticum (Bl.) Sleum. 
872, 873 
var. glanduligerum Sleum. 
798, 872, 874 
var. laurifolium 872, 873, 
874 
var. pensile (Sp.Moore) 
Sleum. 872, 873 
var. robustum 
Sleum. 872, 873 
var. sarawakense (Merr.) 
Sleum. 872, 873 
var. trichodes Sleum. 872, 
873 
ledermannii Schitr 773 
leptanthum Mig. 755, 764 
f. ellipticum (J.J.S.) Sleum. 
764 
f. leptanthum 764 
f. malayanum Sleum. 764 


(Ridl.) 


var.  ellipticum  (J.J.S.) 
Masam. 764 
leptocladum Sleum. 799, 861 
leptomorphum Sleum. 799, 
866 
leptospermoides J.J.S. 779, 
780 
f. glabrum J.J.S. 767, 768, 
780 
f. leptospermoides 767, 768, 
779*, 780* 
ligustrifolium J.J.S. 800, 870 
littoreum Miq. 747, 797, 854 
lobbii (Ridl.) Sleum. 754, 755 
loheri Merr. 741 
longepedicellatum Sleum. 799, 
863 
longibracteatum Ridl. 804 
longilingua Sleum. 753 
longipes Sleum. 753 
longiporum (non Schltr) Diels 
876 
longiporum Schltr 795, 831 
longisepalum J.J.S. 786, 788 
loranthifolium Ridl. 797, 853 
lorentzii Koord. 782 
f. lorentzii 768, 782 
fi, puberultiny Jasna 768; 
782 
lucidum (BI.) Mig. 746, 794, 
814* 
f. epiphyticum O.K. 814 
f. terrestre O.K. 814 
var. lucidum 799, 800, 814 
var. micranthum Hochr. 814 
var. orientale Hochr. 820 
var. pumilum J.J.S. 867 
var. roseitinctum Sleum. 
815 
var. typicum Hochr. 814 
luridum Sleum. 777 
luzoniense Vidal 796, 843 
macbainii F.v.M. 747, 748 
macgillivrayi Seem. 791 
macgregorii Merr. 804 
macrophyllum (J.J.S.) Sleum. 
757 
malacca = malaccense 801 
malaccense Wight 801 
var. bancanum Miq. 801 
var. celebense J.J.S. 802 
malacothrix Sleum. 786, 787 
marginellum Sleum. 778 
mearnsii Elm. 870 
medinilloides Elm. 745 
megalophyes Sleum. 747, 794, 
825 
megaphyllum Sleum. 754, 757 
var. adenophorum Sleum. 
757 
var. megaphyllum 757 
micrantherum Stapf 843 
micranthum = micrantherum 
843 
microphyllum Reinw. ex BI. 


742, 766, 768, 778 
microphyllum (non Reinw. ex 
Bl.) F.-Vill. 812, 850 
microphyllum (non Bl.) K. & 
G23 


microphyllum (non BI.) J.J.S. 
775 
mindorense Rendle 778 
minimiflorum Sleum. 754, 759 
minusculum Sleum. 797, 852 
minuticalcaratum J.J.S. 794, 
797, 822 
f. capillatum Sleum. 866 
f. glabrum J.J.S. 822, 823* 
f. latifolium J.J.S. 822 
f. minuticalcaratum 822, 
823 
miquelii Boerl. 470, 794, 810* 
var. atjehense Sleum. 811 
var. miquelii 793, 810 
mjoebergii J.J.S. 795, 834 
molle J.J.S. 796, 844 
var. molle 844, 845 
var. mollissimum (Sleum.) 
Sleum. 792, 845 
mollissimum Sleum. 845 
monanthum Ridl. 754, 756 
montis-ericae Sleum. 793, 809 
moultonii Merr. 755, 763 
muriculatum J.J.S. 795, 829 
var. albidum J.J.S. 829 
myrianthum Sleum. 765 
myrsinoides Schltr 766, 772 
myrtoides (Bl.) Miq. 746, 747, 
793-0 1944 S12 813754821 
var. B 810 
var. celebicum J.J.S. 812 
nitens Sleum. 800, 869 
nitidum Andr. 814 
nyctericoides =  nycteroides 
823 
nycteroides Wernh. 823 
oblongum Wright 752 
obovalifolium Sleum. 824 
obversum Miq. 815 
oranjense J.J.S. 777, 778 
var. marginellum (Sleum.) 
Sleum. 767, 778 
var. oranjense 767, 777, 778 
oreites Sleum. 786, 788 
oreomyrtus Sleum. 767, 777 
otophyllum Sleum. 794, 827, 
861 
pachydermum Stapf 795, 834* 
palawanense Merr. 795, 837 
var. foxworthii (Copel. f.) 
Sleum. 837 
var. palawanense 837 
paludicolum Sleum. 794, 814 
papuanum J.J.S. 868 
paradisearum Becc. 794, 824* 
parvibaccatum J.J.S. 811 
parvulifolium (non F.v.M.) 
Diels 769 
parvulifolium F.v.M. 766, 771 


1022 


FLORA MALESIANA 


(ser. I, vol. 68 


pauciflorum Fletch. 753 
pensile Sp. Moore 873 
perakense Ridl. 852 
perrigidum Elm. 798, 855 
philippinense Warb. 793, 808 
phillyreoides Sleum. 799, 861 
piceifolium Wernh. 767, 774 
pilosiflorum J.J.S. 768 
pilosilobum J.J.S. 474, 799, 
867 
piperifolium Sleum. 755, 762 
piroliflorum J.J.S. 826 
platyphyllum Merr. 793, 810 
polvanthum O.K. 872 
var. bicolor O.K. 872 
var. viridiflorum O.K. 872 
profusum J.J.S. 824 
prostratum Sleum. 767, 768, 
785 
psammogenes Sleum. 795, 
830 
pseudocaudatum Sleum. 797, 
801, 874 
psittacobium Sleum. 824 
pubicarpum Ridl. 801 
pugionifolium Sleum. 767, 
774 
pullei J.J.S. 766, 768, 782 
pyriforme Merr. 851 
quinquefidum J.J.S. 794, 823 
var. oranjense J.J.S. 824 
var. quinquefidum 823, 824 
rariflorum Schltr 773 
retevenium = retivenium 856 
reticulato-venosum Sleum. 
799, 866 
retivenium Sleum. 798, 856 
retusifolium J.J.S. 799, 864 
ridleyi Sleum. 753 
rigidifolium Sleum. 796, 799, 
846 
rizalense Merr. 810 
robustum Ridl. 873 
rollinsoni Hook. 814 
roseiflorum J.J.S. 798, 799, 
860 
rubroviolaceum Sleum. 801, 
877 
sabuletorum Ridl. 854 
sabuletrum = sabuletorum 854 
salicifolium (J.J.S.) Masam. 
757 
sanquineum Schltr 768, 781 
sarawakense Merr. 873 
scandens Schltr 797, 853 
schimperi Koord. 815, 820 
schlechterianum Sleum. 785, 
786, 787 
schoddei Sleum. 799, 862 
schultzei Schltr 767, 773 
sclerophyllum Sleum. 799, 867 
scortechinii K. & G. 793, 807 
scyphocalyx Sleum. 766, 771 
sessiliflorum Schltr 769 
simulans Sleum. 795, 833 


var. leptopodum Sleum: 834 
var. simulans 833 
sororium J.J.S. 767, 776 
sorsogonense Elm. 8C4 
sp. Koord. 774, 815 
sp. Merr. 844 
sp. Rappard 874 
sp. Stapf 859, 861 
sp. Vidal 812, 850, 875 
spaniotrichum Sleum. 786, 
790 
sparsicapillum Sleum. 766, 
769 
sparsum Sleum. 801, 876 
stabilipes Sleum. 752 
stapfianum Sleum. 793, 796, 
838, 839 
var. minus Sleum. 839 
var. stapfianum 839 
steinii Sleum. 801, 876 
stellae-montis Sleum. 799, 
800, 862 
stenanthum Sleum. 796, 843 
stenolobum Schltr 794, 825 
striicaule Sleum. 471, 876 
var. adenodes Sleum. 798, 
877 
var. pubiflorum Sleum. 796, 
877 
var. striicaule 801, 876, 877 
subobovatum Fletcher 852 
subulisepalum J.J.S. 786, 790 
sulcatum Ridl. 754, 758 
suluense Copel. f. 840 
summifaucis Sleum. 796, 845 
sumatranum Jack 877 
sylvaticum Elm. 800, 870 
taxifolium Sleum. 766, 772 
tenerellum Sleum. 754, 760 
tentaculatum J.J.S. 795, 828 
tenuipes Merr. 796, 844 
teysmannii {non (Bl.) Miq.] 
Koord. 873 
teysmannil Miq. 872 
teysmannii (non Miq.) Rid. 
852 
thibaudifolium Wernh. 793, 
806 
tiariforme J.J.S. 868 
timonioides Wernh. 786, 788 
timorense Fawc. 792, 793, 
796, 797, 802 
var. denticulatum Fawc. 802 
tomicipes J.J.S. 796, 800, 869 
torricellense Schltr 868 
trichocarpum Sleum. 792, 805 
tubiflorum J.J.S. 797, 851 
turbinatum Merr. 857 
turfosum Sleum. 799, 801, 874 
uniflorum J.J.S. 754, 757 
urnigerum Sleum. 779 
uroglossum Sleum. 754, 759, 
760* 
urophyllum Merr. 753 
varingiaefolium (BI.) Miq. 


470, 471, 746, 794, 802, 815, 
816*, 817*, 818, 819*, 820* 
f. parvifolia Miq. 815 
f. sublanceolata Miq. 815 
var. angustifolium O.K. 815 
var. calcaratum Sleum. 793, 
815, 820 
var. erythrinum (Hook.) 
O.K. 815 
var. orientale 
Sleum. 815, 820 
var. pilosiusculum Hochr. 
815 
var. racemosum Hochr. 815 
var. typicum Hochr. 815 
var. varingiaefolium 815, 
816, 821 
varingiaefolium [non (BI.) 
Miq.] Miq. 845 
varingiaefolium [non (BI.) 
Miq.] Vidal 812 
varingifolium = varingiaefo- 
lium 815 
versteegil Koord. 766, 771 
vidalii (non Merr. & Rolfe) 
H.J.Lam & Holth. 850 
vidalii Merr. & Rolfe 797, 848 
villarit Vidal 812 
villosiflorum J.J.S. 766, 768 
viridiflorum J.J.S. 795, 833 
viscifolium K. & G. 800, 852, 
853 
var. bicalcaratum Sleum. 
853 
var. minus K. & G. 852 
var. viscifolium 797, 853 
vonroemeri Koord. 768, 780 
warburgii Sleum. 794, 821 
whiteanum Sleum. 767, 775, 
778 
whitfordii Merr. 768, 782 
wisselianum Sleum. 792, 805 
wollastonii Wernh. 768, 785 
wondiwoiense J.J.S. 767, 772 
woodianum Copel. f. 797, 850 
wrayi Ridl. 843 
xerampelinum Sleum. 799, 
864, 865* 
zollingeri Miq. 872 


(Hochr.) 


Valeriana chinensis L. 455 
Valerianaceae 4: 253-254 
Valli-Modagam Rheede 326, 328 
Vareca moluccana Roxb. 963 
Velleia J.E.Smith 950 


spathulata R.Br. 950 


Ventilago 280 


dichotoma (Blco) Merr. 284 


Vernonia arborea 152 
Vertifolia rubra Rumph. 291 
Viburnum amplificatum Kern 


929, 930* 

clemensae Kern 929 
glaberrimum Merr. 929 
hispidulum Kern 929 
junghuhnii Miq. 929 


Sept. 1972] 


lutescens BI. 929 

platyphyllum Merr. 929 

vernicosum Gibbs 929 
Vieillardia Brongn. & Gris 457 
Viereya = Vireya 480 
Violaceae 237, 963 
Violales 63 
Vireya Bl. 480 

alba Bl. 539 

celebica Bl. 615 

javanica Bl. 624 

retusa Bl. 482 

tubiflora Bl. 532 
Vitmannia Vahl 199 

elliptica Vahl 199 

lucida Steud. 201 

polyandra Steud. 203 
Voacanga sp. 369 


Wahlenbergia Schrad. ex Roth 
1072 109) 110; WI 125 3: 
128 
agrestis DC. 115 
bivalvis Merr. 117, 118 
candollei Tuyn 114 
confusa Merr. & Perry 107, 

mise i4. 115* 
consimilis Lothian 118 
dehiscens (Roxb.) DC. 115 
erecta (Roth ex R. & S.) Tuyn 

113, 114* 
eurycarpa Domin 115 
gloriosa Lothian 118 
gracilenta Lothian 116, 118 
gracilis (Schrad.) DC. 115, 

116, 118 

var. hirsuta Jungh. 115 

var. vincaeflora DC. 118 
gracilis E.Mey., nomen 117 
hirsuta Steud., nomen 113 
hirsuta (Edgew.) Tuyn 113 
hookeri (Clarke) Tuyn 107, 

113, 114* 
indica DC. 115 
lavandulaefolia DC. 115 
marginata (Thunb.) DC. 107, 

MMOs iss 10S; 1l6*, 117* 

map 

var. grandiflora Tuyn 113, 

118 
subvar. trichogyna 
(Stearn) Tuyn 118 


multicaulis Bth. 115 

paniculata (Thunb.) DC. 114 

perotifolia W. & A. 113 

quadrifida (R.Br.) DC. 115, 
118 


sieberi DC. 115 
simplicicaulis de Vriese 115 
trichogyna Stearn 118 
vincaeflora (Vent.) Decne 118 
Weigelia coraeensis Thunb. 929 
fallax Mig. 929 
Wendlandia 152 
Wickstroemia = 
28 
Wikstroemia Endl. 2, 3, 4, 5, 28 
subg. Chamaejasme (Amman) 
Domke 28 
subg. (Eu)wikstroemia 28 
sect. Euwikstroemia Meisn. 


Wikstroemia 


acuminata Merr. 31 

amplifolia (Schltr) Domke 35 

androsaemifolia Decne 2, 29*, 
30) 325338 

androsaemifolia Hand.-Mazz. 
34 

angustissima Mertr. 30 

aurantiaca 4 

australis Endl. 28 

brachyantha Merr. 2, 29*, 30, 
31 

calva Back. 32, 33 

candolleana Meisn. 33 

candolleana (non 
Ridl. 32 

candollet = candolleana 32 

chamaejasme (L.) Domke 28 

clementis Merr. 31 

crassifolia Merr. ex Domke 31 

fenicis Merr. 33 

indica (L.) C.A.Mey. 2, 3, 4, 
30, 34 
var. viridiflora Hook. f. 35 

junghuhniana = junghuhnii 32 

Junghuhnii (non Miq.) K. & V. 
32 

Junghuhnii Migq. 32, 33, 34 

lanceolata Merr. 30 

linearifolia Elm. 35 

longifolia Lecomte 33 

meyeniana Warb. 30, 33 

nutans Champ. 33 


Meisn.) 


Index to scientific names 


1023 
ovata C.A.Mey. ex Meisn. 30, 
31, 32 map 
ovata (non C.A.Mey.) Vidal 
35 
pachyphylla Merr. 35 
polyantha Merr. 30, 32 
pulgarensis Elm. 35 
ridleyi Gamble 30, 34 
ridleyi (non Gamble) Gibbs 32 
spanoghii Decne 33, 34 
subcoriacea Merr. 35 
tenuiramis Miq. 29*, 30, 31 
venosa Merr. & Perry 30, 33 
viridiflora Meisn. 35 
Willughbeia auriculata Spreng. 
327, 328 
elliptica Spreng. 303 
fragrans Spreng. 307 
racemosa Spreng. 311 
volubilis Spreng. 311 
Wimmeria Schlechtend. 228, 
229, 231 
Wirtgenia Andres 
474, 669, 943 
malayana (Scort. in Hook. f.) 
Andres 669 
Wrightia laniti (Blco) Merr. 373 


Xanthophyllum subglobosum 
Elm. 395 
var. longifolium Elm. 395 
Xanthostemon brassii Merr. & 
Perry 428 
Xanthoxylum = 
56 
Xolisma Raf. 674 
ovalifolia (Wall.) Rehd. 675 
Xylonymus Kalkman 231, 232, 
243, 391, 392, 930 
versteeghii Kalkman 
245 
Xylosma luzonense (Presl) Clos 
944 
palawanense Mendoza 944 
Xyridaceae 4: 366-376, 598; 5: 
557 


Zannichellia 157 
Zannichelliaceae 157 
Zanthoxylum 207, 209 
montanum BI. 56 
serrulatum Bl. 56 
Zygophyllaceae 4: 64 


469, 470, 


Zanthoxylum 


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Volume 1. Cyclo 1 collector , Collec ?: 0. pp. cl i 639 at 
: oe (in preparation) ‘ 
Volume 3. ian plant geo; | : “(In pr eparation) ak 
Volume 4: General chapters and revisions. 1948-1954 . pp. ccix + 63] 
Volume 5. Bibliography, specific delimitation, & re visions. 1955-1958. pp. 
cccxlii + 596 | 
Volume 6. Systematic revisions. 6 parts, 1960-1972. pp. 20 +- 1023. 
Volume 7. Systematic revisions. Part 1. 1971. pp. 263 


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