THE UNIVERSITY
OF ILLINOIS
LIBRARY
PUBLICATIONS
OF
FIELD MUSEUM OF NATURAL
HISTORY
v
S' NATURAL
HISTORY
THE LIBRARY OF THE
MAR 151939
UNIVERSITY OF ILLINOIS
CHICAGO, U.S.A.
1936-1938 !
PRINTED IN THE UNITED STATES OF AMERICA
BY FIELD MUSEUM PRESS
BOTANICAL SERIES
FIELD MUSEUM OF NATURAL HISTORY
FOUNDED BY MARSHALL FIELD, 1893
VOLUME XIII
FLORA OF PERU
PART II
BY
J. FRANCIS MACBRIDE
ASSISTANT CURATOR OF THE HERBARIUM, DEPARTMENT OF BOTANY
B. E. DAHLGREN
CURATOR, DEPARTMENT OF BOTANY
EDITOR
PUBLICATION 357
CHICAGO, U.S.A.
JUNE 10, 1936
PRINTED IN THE UNITED STATES OF AMERICA
BY FIELD MUSEUM PRESS
FB
v, \$r
FLORA OF PERU
J. FRANCIS MACBRIDE
40. PI PER AGE AE. Pepper Family
By William Trelease
Herbs, shrubs, or small trees; leaves usually alternate, some-
, times opposite or verticillate, entire; stipules none or adnate to the
petiole; flowers minute, usually green, perfect or unisexual, in spikes
i or racemes, these pedunculate, terminal or opposite the leaves,
sometimes axillary or several on a common peduncle, the flowers
usually densely crowded in the spike, sometimes annulately or
spirally arranged; bracts usually peltate, sessile or stipitate, some-
times adnate or concave; perianth none; stamens usually 2-6, the
filaments free, the anthers terminal, erect, 2-celled; ovary sessile
or stipitate, 1-celled, 1-ovulate; fruit baccate, very small.
Stigma 1; plants herbaceous, usually small, most often epiphytic.
1. Peperomia.
Stigmas usually 3; plants large, usually woody and terrestrial.
Spikes in an axillary umbel 2. Pothomorphe.
Spikes opposite the leaves.
Spikes solitary 3. Piper.
Spikes often accompanied by a collateral spike.
4. Pleiostachyopiper.
1. PEPEROMIA R. & P.
Annual or perennial herbs, epiphytic or terrestrial, usually very
fleshy; leaves alternate, opposite, or verticillate, without stipules;
flowers scattered or densely crowded upon the rachis, often partly
immersed in pits; stamens 2. A vast genus, with many species in
almost all parts of tropical America.
A. Plants acaulescent.
Leaves distinctly peltate.
Leaves scarcely 1 cm. in diameter; spikes single.
Spikes scarcely 10 cm. long.
Berries much wrinkled; stylopodium thick.
Berries ellipsoid P. parvifolia.
3
1 067037
4 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Berries subovoid P. verruculosa.
Berries not wrinkled.
Corm small P. minuta.
Corm large P. macrorhiza.
Spikes distinctly longer; stylopodium elongate.
Spikes 20 mm. long : P. Seleri.
Spikes 60-80 mm. long P. Hillii.
Leaves 1-2 cm. in diameter; stylopodium elongate.
Spikes scarcely over 10 mm. long P. umbilicata.
Spikes 20-60 mm. long P-..peruviana.
Leaves 2-4 cm. in diameter.
Spikes 15-30 mm. long P. falsa.
Spikes 50-60 mm. long.
Leaves from a corm P. scutellaefolia.
Leaves from a rhizome P. rupiseda.
Spikes 90-100 mm. long P. polycephala.
Leaves not peltate, 2-4 cm. wide.
Spikes single, elongate P. Sprucei.
Spikes paniculate, short P. villipetiola.
A. Plants with leafy stems.
B. Leaves alternate but sometimes congested upward.
C. Leaves distinctly peltate.
Spikes paniculate; leaves 10-15 cm. long P. tarapotana.
Spikes not paniculate.
Leaves 10-15 cm. long.
Plants glabrous.
Spikes from axils of reduced leaves P. dependens.
Spikes subtended by ordinary leaves P. variegata.
Plants hairy P. Schunkeana.
Leaves 6-10 cm. long.
Leaves glabrous P. rugata.
Leaves ciliate P. majalis.
Leaves commonly 4-6 cm. long.
Plants glabrous.
Leaves round-ovate . .' P. deficiens.
Leaves more elongate P. puberulaeformis.
FLORA OF PERU 5
.Plants crisp-pubescent P. cuprea.
Plants subvillous P. arbor igaudens.
Leaves 2.5-4 cm. long; plants pubescent P. flavescens.
C. Leaves at most obscurely subpeltate at the base.
D. Spikes paniculate.
Leaves 10-15 cm. long, pinnately nerved.
Spikes very short; glabrous P. Lechleriana.
Spikes elongate.
Plants pubescent.
Leaves ovate P. spectabilis.
Leaves elliptic P. striata.
Plants glabrous.
Leaves elliptic-oblanceolate P. balsapuertana.
Leaves oblanceolate.
Panicle branched P. arcualispica.
Panicle simple . P. uncatispica.
Leaves lanceolate P. acutifolia.
Leaves 6-10 cm. long, pinnately nerved, glabrous.
P. Weberbaueri.
Leaves 4-6 cm. long.
Leaves round, cordate, submultiple-nerved.
Plants appressed-villous ; panicle ample P. secunda.
Plants subglabrescent; panicle with a few branches.
P. Macbndeana.
Leaves obovate, acute-based, glabrous.
Leaves palmately nerved P. arborised a.
Leaves pinnately nerved P. obliqua.
Leaves 2.5-4 cm. long, cleaver-shaped, vertical rather than hori-
zontal P. dolabriformis.
Leaves 1-2.5 cm. long.
Leaves compressed from the side.
Leaves glabrous P. nualis.
Leaves ciliate on the keel.
Spikes 20 mm. long .P. lepadiphylk i.
Spikes 40 mm. long P. naviculaefolia.
6 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves broader than thick P. macrolhyrsa.
D. Spikes umbellate or solitary, attached at the end of the
peduncle.
Leaves 2.5-4 cm. long.
Plant glabrous P. foliiflora.
Plant pubescent P. Haenkeana.
Leaves scarcely 2.5 cm. long.
Leaves cordate; spikes several P. phyllaniha.
Leaves not cordate; spikes solitary.
Leaves 5-7-nerved P. Fortunati.
Leaves 3-nerved P. umbelliformis.
D. Spikes neither paniculate nor umbellate nor attached to the
end of the peduncle.
* E. Leaves 10-15 cm. long.
Plants villous; leaves ovate P. yapasana.
Plants crisp-pubescent; leaves elliptic, narrowly acuminate.
P. pubiramea.
Plants glabrous.
Leaves ovate, cordate or round-based.
Stem very short; nerves palmate P. curtipes.
Stem elongate.
Nerves palmate P. pinedoana.
Nerves multiple.
Leaves with nearly simple midrib P. itayana,.
Leaves nerved from the lower third.
P. pseudorhynchophoro,.
Leaves nerved from the lower two-thirds.
P. pergamentacea.
Leaves elliptic, acute-based, pinnately nerved.
Leaves half as broad as long.
Ovary sharp-beaked P. quimiriana.
Ovary not beaked P. exiliramea.
Leaves more elongate . P. tenuiramea.
Leaves lanceolate, pinnately nerved.
Spikes 2 mm. thick P. binispica.
Spikes 5 mm. thick P. huacachiana.
FLORA OF PERU 7
Leaves obovate-palmate, pinnately nerved.
Spikes 2 mm. thick.
Leaves scarcely emarginulate P. cruciata.
Leaves emarginulate ; P. Fieldiana.
Spikes 5-10 mm. thick P. emarginata.
E. Leaves 6-10 cm. long.
F. Leaves palmately nerved.
Leaves broad.
Leaves velvety throughout P. pubinervosa.
Leaves fugaciously pubescent P. rhombiformis.
Leaves ciliate and locally puberulent, blunt-acuminate.
P. Poeppigii.
Leaves glabrous.
Leaves acute at the base.
Leaves prominently cross- veined P. enckeaefolia.
Leaves not prominently cross- veined .... P. chartacea.
Leaves rounded at the base or cordulate.
Leaves pale-areolate P. areolata.
Leaves white between the nerves.
Leaves orbicular P. procumbens.
Leaves subelliptic P. sarcostachya.
Leaves purple beneath P. monostachya.
Leaves uniformly rosy P. rubrifolia.
Leaves elongate, acute-based.
Stem winged P. alata.
Stem not winged P. non-alata.
F. Leaves pinnate- or multiple-nerved.
G. Leaf base acute.
Plants glabrous.
Leaves clustered at the top of the stem, acuminate.
P. racemifolia.
Leaves scattered.
Spikes not paired.
Leaves subobovate P. acuminata .
Leaves ovate P. purpurea.
Leaves subrhombic-elliptic P. basellaefolia.
8 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves lanceolate .P. bermudezana.
Spikes paired.
Leaves obovate P. eleganti/olia.
Leaves ovate-elliptic P. granulatilimba.
Leaves lance-ovate P. santiagoana.
Leaves elliptic P. elegans.
Leaves rhombic-ovate P. mercedana.
Leaves and petioles ciliate.
Leaves elliptic P. ciliatifolia.
Leaves broadly lanceolate.
Spikes 150 mm. long P. quatrometralis.
Spikes shorter.
Plant subrepent P. semisupina.
Plant erect P. blepharipus.
Plants locally puberulent P. limbuchiana.
Plants crisp-pubescent P. enenyasensis.
G. Leaf-base auriculate P. pilosa.
G. Leaves cordate, submultiple-nerved.
Leaves about 5 cm. wide.
Plants somewhat pubescent ...:.... P. juniniana.
Plants glabrous P. erylhroslachya .
Leaves wider, glabrous.
Stem short and few-leaved P. curticaidis.
Stem elongate.
Leaves subtruncate at the base P. ainana.
Leaves cordate.
Peduncle 3 cm. long P. longepedunculata.
Peduncle 7-10 cm. long P. Opiziana.
G. Leaf base rounded P. santiagoana.
E. Leaves 4-6 cm. long.
F. Leaves pinnate- or multiple-nerved.
Leaf base acute or subacute.
Leaves scarcely 2 cm. wide.
Stems compactly branched.
Leaves papillate P. crystalline.
Leaves not papillate P. metallica.
FLORA OF PERU 9
Stem slender and elongate , .P. pseud o-salicifolia.
Leaves distinctly wider.
Berries slender-beaked.
Plants pubescent.
Leaves acute P. syluatica.
Leaves obtuse P. teresitensis.
Plants glabrous.
Leaves elliptic-obovate.
Spikes 50 mm. long P. emarginulala.
Spikes longer P. deodorata.
Leaves round-obovate P. haematolepis.
Leaves oblanceolate P. parvipynctulala.
Leaves rhombic-elliptic ........ P. elhplico-rhombea.
Berries mucronate P. rhombilimba.
Berries Scutulate-truncate.
Plants tomentulose P. pertomenlella.
Plants transiently pubescent; leaves ovate.
Leaves drying coriaceous P. coloniac.
Leaves drying thin P. pachiteana.
Plants somewhat puberulent P. cilialifolia.
Plants somewhat ciliate and granular.
Leaves lance-elliptic '. . . . .P. scutalei 'folia.
Leaves elliptic P. flavescentifolia.
Leaves subovate P. granulalifolia.
Leaves cordate, glabrous.
Leaves becoming subtriangular P. crotalophora.
Leaves round-ovate.
Stem slender and fragile . P. fragilissima.
Stem stronger P. huacapistanana.
F. Leaves palmately nerved.
Leaves cordate.
Plants puberulent P, Grisari.
Plants glabrous.
Spikes solitary.
Leaves subacuminate P. gibba.
Leaves obtuse P. rufescens.
10 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Spikes paired P. currucaeform.is.
Leaves rounded at the base, glabrous.
Leaves obtuse P. alismaefolia.
Leaves subacute.
Blade narrowed to the petiole, subacuminate.
P. rufescentifolia.
Blade crossing the petiole P. verediana.
Leaves acute at the base.
Plants glabrous.
Leaves elongate-lanceolate, long-attenuate to apex.
P. salicifolia.
Leaves lance-elliptic.
Peduncle about 1 cm. long.
Spikes short P. Vidaliana,
Spikes 90 mm. long P. exiguispica.
Spikes 100-150 mm. long P. spiculata.
Spikes 150-200 mm. long P. oxyphylla.
Peduncle 2 cm. long P. montana.
Peduncle 2-3 cm. long P. obruenda.
Leaves elliptic.
Peduncle 5 cm. long P. variculata.
Peduncle 1 cm. long P. puberulibacca.
Plants velvety P. dasystachya.
Plants crisp-pubescent.
Spikes with pseudo-pedicels P. crusculibacca.
Spikes without pseudo-pedicels P. leuconeura.
Plants villous P. tricolor.
E. Leaves 2.5-4 cm. long.
H. Leaves palmately nerved.
Leaf base acute.
Leaves 5-7-nerved.
Leaves at most ciliate or ciliate-lined.
Leaves drying thick.
Spikes short P. hamiUonianifoUa.
Spikes long P. induratifolia.
FLORA OF PERU 11
Leaves drying rather thin.
Spikes 70 mm. long P. megalepis.
Spikes about 40 mm. long.
Leaves suborbicular P. furcata.
Leaves subelliptic P. microlepis.
Leaves subobovate P. debilipes.
Leaves ovate P. Obex.
Leaves crisp-pubescent.
Leaves subovate-elliptic P. famelica.
Leaves round-ovate P. nigricans.
Leaves 3-nerved.
Plants glabrous.
Stem angled P. alatiscapa.
Stem not angled.
Spikes about 50 mm. long.
Leaves oblanceolate P. fortipes.
Leaves lance-oblong P. carrapana.
Spikes long P. chanchamayana.
Plants subpuberulent P. caducifolia.
Plants crisp-pubescent.
Leaves elliptic-subobovate P. hyporhoda.
Leaves lance-ovate P. trinervis.
Leaf base obtuse or cordulate.
Plants appressed-pubescent P. carpapatana.
Plants crisp-pubescent P. chalhuapuquiana.
Plants subpuberulent P. Maransara.
Plants glabrous.
Plant erect, pellucid P. pellucida.
Plant repent P. pseudo-furcata.
H. Leaves pinnately nerved.
Plants glabrous P. ventricosicarpa.
Plants villous P. yananoensis.
Plants transiently silky P. percalvescens.
E. Leaves 1-2.5 cm. long.
Leaves pinnately nerved, acute-based.
12 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Plants glabrous.
Leaves obovate ,...-. P. parva.
Leaves elliptic-obovate P. roqueana.
Leaves oblanceolate . P. cusilluyocana.
Plants subvillous P. pilifera.
Leaves palmately nerved.
Leaves acute-based.
Plants glabrous.
Leaves lanceolate.
Leaves acute P. aerea.
Leaves obtuse P. mansericheana.
Leaves subrhombic P. rhombeifolia,
Plants crisp-pubescent P. pumila.
Plants subvillous . . P. Drapeta.
Leaves cordate or round-based.
Plants glabrous P. antoniana.
Plants somewhat crisp- villous P. pontina.
Plants crisp-pubescent P. scandens.
E. Leaves scarcely 1 cm. long, if somewhat longer the blades
narrow.
I. Leaves nearly as broad as long.
Leaves subreniform.
Plants crisp-pubescent, very delicate, repent . . P. scandens.
Plants soft-hirtellous P. perhispidula.
Leaves round-ovate P. longipila.
Leaves orbicular.
Leaves varying into elliptic.
Plants glabrous P. multiformis.
Plants finely subvillous.
Peduncle 5-10 mm. long P. pleiomorpha.
Peduncle very short P. pichisemis.
Plants crisp-pubescent.
Peduncle very short P. polymorpha.
Peduncle 5 mm. long P. vana.
Leaves uniform.
FLORA OF PERU 13
Plants glabrous : P. fuscispica.
Plants somewhat pilose P. andinacea.
Plants crisp-pubescent P. rotundifolia.
Leaves round-obcordate.
Plants puberulent P. cordulilimba.
Plants glabrous P. obcordatifolia.
Leaves obliquely elliptic P. pruinosi folia.
I. Leaves subelongate-ovate.
Leaves scarcely more than 1-nerved.
Plants transiently silky P. subsericata.
Plants at most obscurely puberulent.
Leaves very variable P. heterophylla.
Leaves uniform P. playapampana.
Leaves 5-nerved P. modicilimba.
Leaves obscurely pinnately nerved.
Leaves half as broad as long P. non-hispidula.
Leaves narrower P. plicatifolia.
B. Leaves opposite or whorled.
J. Leaves 6-10 cm. or more long, glabrous.
Leaves 3-nerved P. heterostachya.
Leaves 5-nerved P. tetragona.
Leaves 7-9-nerved.
Leaves lance-elliptic P. septemnervis.
Leaves elliptic P. undeninerma.
Leaves oblanceolate P. deceptrix.
J. Leaves 4-6 cm. long.
Plants glabrous.
Drying purple.
Leaves narrowly lanceolate P. rubeo,.
Leaves lance-elliptic or ovate P. rubens.
Not drying purple.
Leaves 5-nerved P. puteolata.
Leaves 7-nerved P. san-roqueana.
Plants locally puberulent.
Leaves lance-elliptic P. Pavoniana.
Leaves subovate-elliptic P. mishuyacana.
14 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Plants generally puberulent P. inaequilatera.
Plants with the stem pubescent.
leaves glabrous P. exilamenta.
Leaves pubescent above on the nerves, acuminate.
P. pubescentinervis.
Leaves generally pubescent.
Upper leaves reduced P. pampalcana.
Upper leaves not smaller P. casapiana.
J. Leaves 2.5-4 cm. long.
Leaves commonly opposite.
Leaves papillate-puberulent, acute at the base.
P. ollantaitambona.
Leaves pubescent.
Upper leaves reduced P. confertispica.
Upper leaves scarcely smaller.
Plant crisp-pubescent P. apurimacana.
Plant subvillous-tomentose P. cotoneasterifolia.
Plant hirsute P. Gayi.
Leaves mostly whorled.
Plants glabrous.
Leaves drying thin and green P. profissa.
Leaves drying firm.
Leaves purple P. rubescens.
Leaves bronzed, finely wrinkled, obtuse-acuminate.
P. aster ophylla,
Leaves scarcely purpling or bronzed.
Leaves revolute and hard P. sderophylla.
Leaves scarcely revolute.
Leaves 3-nerved P. flabilis.
Leaves 5-nerved P. ficta.
Plants more or less puberulent.
Leaves lance-ovate P. flavamenta.
Leaves broadly lanceolate P. glareosa.
Leaves narrowly lanceolate P. tetrica.
Leaves obovate to subspatulate P. inaequali folia.
Plants crisp-pubescent.
FLORA OF PERU 15
Leaves rhombic-spa tulate P. Mathewsiana.
Leaves elliptic P. praeruptorum.
Stem subvillous P. subvillicaulis.
J. Leaves 1-2.5 cm. long.
Leaves opposite.
Plants glabrous.
Leaves not papillate P. Herrerae.
Leaves minutely papillate P. gucayana.
Plants hairy P. villicaulis.
Leaves commonly whorled.
K. Leaves widened upwards.
Leaves uniform, glabrous.
Spikes short, apiculate P. obcordata.
Spikes not apiculate.
Spikes short P. quadricoma.
Spikes 25 mm. long P. estrellana.
Spikes 50 mm. long P. san-buenaventurana.
Spikes 60-80 mm. long.
Leaves coriaceous, revolute P. palcana.
Leaves scarcely revolute, granular, emarginulate.
P. pillahuatana .
Spikes over 100 mm. long.
Leaves 12 cm. long P. santa-cruzana.
Leaves becoming twice as long P. Pakipski.
Leaves partly subspatulate.
Plants glabrous.
Petiole scarcely 2 mm. long; leaves emarginate.
P. subdichotoma.
Petiole longer, winged P. huallagana.
Plants velvety P. anisophylla.
Plants hirsute P. Klotzschiana.
K. Leaves not widened upwards.
Leaves lanceolate.
Leaves not rhombic.
Plants tomentulose P. variifolia.
Plants somewhat puberulent.
16 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves acute P. subcalvescens.
Leaves obtuse P. pongoana.
Leaves rhombic P. rhombea.
Leaves elliptic.
Leaves subrhombic P. puberulipes.
Leaves not rhombic.
Plants glabrous or slightly papillate.
Leaves 1-1.5 cm. long.
Leaves plicate P. cushiana.
Leaves flat : ...... P. rhodophylla.
Leaves 1.5-2.5 cm. long.
Spikes solitary . P. concam.
Spikes numerous P. congesta.
Plants puberulent.
Leaves scarcely 1 cm. wide P. huanucoana.
Leaves 1.5 cm. wide P. viracochana.
Plants soft-hairy P. calvescens.
Plants stiff-hirsute P. crinigera.
Plants villous.
Leaves acute P. congestispica.
Leaves obtuse P. caniana.
Leaves lance-elliptic P. guayrapurana.
Leaves lance-oblong P. lanosa.
Leaves subrhombic-ovate P. Smithii.
J. Leaves scarcely over 1 cm. long, narrow if slightly longer.
Leaves opposite, orbicular P. cyclophylla.
Leaves mostly whorled.
L. Leaves round-elliptic or subovate-elliptic.
Leaves glabrous.
Spikes 60 mm. long P. Ppucu-ppucu.
Spikes twice as long.
Leaves not waxen P. marcapatana.
Leaves drying waxen-translucent P. career-
Leaves pubescent.
Stem exfoliating, glabrous P. scabiosa.
Stem transiently hirtellous P. Diehliana.
Stem crisp-hairy P. roseo.
FLORA OF PERU 17
Stem yellow-tomentulose P. chrysotricha.
L. Leaves round, at most slightly puberulent.
Peduncle 3-4 cm. long P. trichopus.
Peduncle 1-2 cm. long; leaves rounded at the end.
Spikes 10 mm. long, 2 mm. thick P. subandina.
Spikes 10-15 mm. long, 1 mm. thick P. andicola.
Peduncle 1 cm. long.
Leaves round at the end P. filiformis.
Leaves subtruncate P. subandina.
Peduncle very short P. Claudii.
L. Leaves obovate.
Plants glabrous P. muscigaudens.
Plants somewhat puberulent P. pseudo-galapagensis.
L. Leaves subelongate-obovate.
Leaves scarcely rhombic or rugose, at most slightly puberulent.
Leaves rounded at the apex.
Leaves 3 or 4 at a node.
Leaves obscurely pinnate-nerved, acute at base.
P. dendroides.
Leaves 1-nerved P. Pearcei.
Leaves about 5 at a node, 1-nerved P. Gilberti.
Leaves truncate-emarginulate P. bracteispica.
Leaves subrhombic, rugulose P. rugatifolia.
L. Leaves elliptic.
Leaves subrhombic.
Spikes short; somewhat pubescent.
Pubescence subvillous P. huantana.
Pubescence short.
Leaves little longer than broad, obtuse-acuminate.
P. nigro-oculata.
Leaves more elongate.
Peduncle short P. rhombeo-elliptica.
Peduncle 2 cm. long P. quadratifolia.
Spikes subelongate.
Plants repent-assurgent P. quaerata.
Plants erect.
18 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Plants at most papillate P. milmfolia.
Plants somewhat crisp-hirtellous P. quaesita.
Leaves scarcely subrhombic.
Plants glabrous.
Stem very thick at the base; leaves elliptic.
P. crassulaecaulis.
Stem not greatly thickened.
Leaves round-elliptic, subobtuse at the base.
P. quispicanchiana.
Leaves twice as long as broad P. arctebaccata.
Plants papillate.
Papillae not bristly P. micromamillata.
Papillae bristly P. naevifolia.
Plants puberulent.
Leaves short-elliptic P. chillonensis.
Leaves more elongated.
Leaves obscurely pinnately nerved.
Stem very slender P. brachyiula.
Stem 2-3 mm. thick P. ceapanana.
Leaves scarcely more than 1-nerved.
Stem 5 mm. thick below P. distractiflora.
Stem slender P. sangabanensis.
L. Leaves oblong or linear.
Upper leaves elongate.
Lower leaves barely 5 mm. long P. atocongona.
Leaves larger.
Spikes and peduncles short.
Stem very leafy P. foliata.
Internodes long P. seposila.
Spikes 60-150 mm. long P. longispica.
Upper leaves scarcely elongated.
Leaves subconduplicate P. trullaefolia.
Leaves not deeply concave.
Stem slender.
Peduncle 5 mm. long P. dendromorphis.
Peduncle twice as long P. agapatensis.
Stem 4-5 mm. thick P. arcuatispica.
FLORA OF PERU 19
Peperomia acuminata R. & P. Fl. 1: 32. pi. 51. 1798. Piper
acuminatum Pers. Syn. 1: 35. 1805, non P. acuminatum L. 1753.
Piper nemorosum Vahl, Enum. 1: 341. 1805. Peperomia Ruiziana
C. DC. Candollea 1: 337. 1923.
A rather large, glabrous, ascending herb; leaves alternate, oblong-
obovate, acuminate, acute-based, 6-9 cm. long, 2.5-3.5 cm. wide,
pinnately nerved; petiole 1 cm. long; spikes terminal, 4 mm. thick,
100 mm. long or more; peduncle short.
Huanuco: Muna, Pillao, and Macosa, Ruiz & Pavon, type.
San Martin: San Roque, Williams 7428.
Peperomia acutifolia C. DC. Journ. Bot. 4: 142. 1866.
A glabrous, moderate-sized herb; leaves alternate, lanceolate,
acutely protracted, long-cuneate, 15 cm. long, 3-4 cm. wide, pin-
nately nerved; petiole 3 cm. long or more; spikes 2 or several in a
terminal panicle, filiform, 80 mm. long; peduncle 10 mm. long;
bracts round-peltate; berries narrowly oblong, obliquely sharp-
scutulate, with central stigma. Neg. 8311.
San Martin: Tarapoto, Spruce 4094, type. San Roque, 1,350-
1,500 meters, Williams 7428. "Afasi quihua."
Peperomia aerea Trelease, sp. nov.
Herba gracilis rhizomatosa repens glabra, ramis brevibus ad-
scendentibus, caule 1 mm. crasso; folia alterna lanceolata sub-
acuminata basi acutiuscula 12-30 mm. longa 6-12 mm. lata coriacea
opaca 3-nervia vel obscure 5-nervia anguste revoluta, subtus con-
coloria granulosa, petiolo 3 mm. longo; spicae terminales 50 mm.
longae 1 mm. crassae, floribus densis vel subzonulatis, pedunculo
10 mm. longo; bracteae rotundo-peltatae.
Junin: Chanchamayo Valley, 1,800 meters, Carlos Schunke 494
(herb. Field Mus. No. 571,552, type), 187.
Peperomia agapatensis C. DC. in DC. Prodr. 16, pt. 1:
455. 1869.
A puberulent, suffrutescent, erect, branching herb; leaves
commonly 4 at a node, oblong or somewhat enlarged upward,
obtuse, subacute at the base, 8-15 mm. long, 3-4 mm. wide, ob-
scurely pinnate-nerved, yellowish beneath; petiole scarcely 1 mm.
long; spikes terminal, 50 mm. long, 1 mm. thick; peduncle 10-15
mm. long; bracts round-peltate; berries round, pointed; stigma
apical. Neg. 8304.
20 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Type from Agapata, Lechler 1934. Junin: La Merced, Schunke
Hacienda, Killip & Smith 24576.
Peperomia ainana Trelease, sp. nov.
Herba modice alta simplex assurgens glabra, caule 2-4 mm.
crasso; folia alterna ovata acuminata, basi rotundata vel sub-
truncata atque ad petiolum acute contracta, 8-10 cm. longa 5-7
cm. lata, multiplinervia nervis basilaribus utroque latere 2-3, majori-
bus ut costa 1-2-ramosis, statu sicco tenuia viridia, petiolo 3-8
cm. longo gracili; spicae sympodiales 40 mm. longae 2 mm. crassae,
pedunculo aequilongo; bracteae rotundo-peltatae; ovarium globosum
mucronatum, stigmate obliquo.
Ayacucho: Aina, near Rio Apurimac, 750-1,000 meters, Killip
& Smith 22766 (type in U. S. Nat. Herb.; dupl. in herb. Field Mus.).
Junin: La Merced, Killip & Smith 23725.
A terrestrial plant; flower spikes white.
Peperomia alata R. & P. Fl. 1: 31. pi. 48. 1798. Piper alatum
Vahl, Enum. 1:342. 1805.
A moderate-sized, glabrous, assurgent herb; stem alately angled
below the petiole margins; leaves alternate, lance-elliptic, sub-
acuminate, 4-7 cm. long, 2-2.5 cm. wide, acute-based, 5-nerved
with the inner nerves somewhat confluent; petiole 5-10 mm. long,
clasping-decurrent; spikes terminal and from the upper axils, 70
mm. long, 2 mm. thick; peduncle 10 mm. long; bracts round-peltate.
Huanuco: Posuso, Ruiz & Pavon, type; at 600 meters, on mossy
rocks, 4687, 4680. Ayacucho: Aina, Killip & Smith 22830, 22769.
Junin: Balsapuerto, Killip & Smith 28428. San Nicolas to
Azupizu, Killip & Smith 24088. Puerto Bermudez, Killip &
Smith 26465.
Peperomia alatiscapa Trelease, sp. nov.
Planta epiphytica mediocris subsimplex glabra; folia alterna vel
suprema ternata elliptico-subobovata obtuse acuminata basi acuta
2.5-4 cm. longa 1.5-2.5 cm. lata 3-5-nervia, costa obscure ramosa,
in sicco opaco-olivacea et opaca, statu vivo obscure rosea, petiolo
10 mm. longo in costulas caulinas decurrente; spicae terminales 70
mm. longae 2 mm. crassae densiflorae, pedunculo 10 mm. longo;
bracteae rotundo-peltatae.
Junin: Enefias, Pichis Trail, 1,600-1,900 meters, Killip & Smith
25623 (type in U. S. Nat. Herb.).
FLORA OF PERU 21
Peperomia alismaefolia Presl, Epim. Bot. 222. 1845.
A rhizomatous herb; stem short, toothed from fallen leaf bases;
leaves few, alternate, elliptic-ovate, obtuse at both ends, 4-6 cm.
long, 2.5-4 cm. wide, 5-nerved, pubescent above and ciliate; petiole
4-5 cm. long, glabrous; spikes from between the leaves, 20 mm.
long, 2 mm. thick; bracts round-peltate, ciliate; ovary subglobose,
umbilicate; stigma apical.
Type from mountains of Peru.
Peperomia andicola Dahlst. Svensk. Vet. Akad. Handl. 33,
pt. 2: 182. pi. 7. 1900.
A delicate, repent, short-branched herb, at least transiently
puberulent; stem filiform; leaves commonly 3 or 4 at a node, round-
obovate, subtruncate, cuneate, 5 mm. wide, 3-nerved; petiole very
short; spikes terminal, 10-15 mm. long, 1 mm. thick; peduncle of
the same length; bracts round-peltate; berries ovoid, pointed, with
pseudo-cupule; stigma apical.
Junin: Tarma, Philippi, type. Loreto: Yapas, 1,350-1,600
meters, Killip & Smith 25462(1).
Peperomia andinacea C. DC. Journ. Bot. 4: 134. 1866.
A delicate, repent herb, glabrous except for a few hairs on the
upper leaf surface; leaves alternate, orbicular, 2-3 mm. in diameter,
obscurely 3-nerved; petiole 1-2 mm. long; spikes terminal, 10 mm.
long, 1 mm. thick; peduncle 2 mm. long.
Puno: Sangaban, Lechler 2410, type.
Peperomia anisophylla C. DC. Bot. Jahrb. 40: 266. 1908.
An erect, succulent herb; stem 2-3 mm. thick, with short inter-
nodes; leaves about 3 at a node, reflexed, obovate, subtruncate or
very obtuse, subcuneate at the base, 10-17 mm. long, 5-10 mm.
wide, velvety on both sides, 5-nerved; petiole very short; spikes
terminal and from the upper axils, 45 mm. long, 1 mm. thick;
peduncle scarcely 5 mm. long; bracts round-peltate; ovary ovoid;
stigma subapical.
Huanuco: Between Matucana and Arinia, 2,300 meters, Weber-
bauer 72, type.
Peperomia antoniana Trelease, sp. nov.
Herba parvula subrepens glabra terrestris; folia alterna rotundo-
ovata obtusissima basi cordata 15-20 mm. longa et lata 5-nervia
in sicco viridia et pellucida, petiolo 5-20 mm. longo; spicae in sicco
22 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
filiformes, 10-15 mm. longae 3 mm. crassae; bracteae rotundo-
peltatae.
Loreto: San Antonio, Rio Itaya, 110 meters, Killip & Smith
29498 (type in U. S. Nat. Herb.).
Peperomia apurimacana Trelease, sp. nov.
Plata epiphytica alta succulenta ramosa, caule 2-4 mm. crasso
crispo-floccoso ; folia opposita subrhombeo-elliptica utrinque acuta,
foliis inferioribus reductis et obovato-truncatis, 3 cm. longa et 1
cm. lata vel usque ad 5 cm. longa atque 3 cm. lata, trinervia,
subpersistente crispo-pubescentia, in sicco tenuia, petiolo 5-10 mm.
vel ad 15 mm. longo crispo-pubescente; spicae terminales et axillares
filiformes 30-40 mm. longae sublaxiflorae floribus pseudopedicellis
brevibus stipatis, pedunculo 10-15 mm. longo translucente minute
pubescente; bracteae rotundo-peltatae.
Ayacucho: Carrapa, 1,500 meters, Killip & Smith 22355.
Peperomia arborigaudens Trelease, sp. nov.
Planta epiphytica repenti-pendula, caule gracili elongato breviter
villoso; folia alterna ovata acuminata brevissime et obscure peltata,
basi rotundata vel subtruncata, 4-5.5 cm. longa 3-4 cm. lata,
5-7-nervia, statu juvenili subtus tomentulosa serius sparse pilosa
et glanduloso-granulosa, in sicco tenuia atque plus minusve rosea,
petiolo 3-5 mm. longo laxe villoso; spicae terminales graciles 90
mm. longae, pedunculo filiformi 2 cm. longo sub villoso; bracteae
minutae rotundo-peltatae.
Loreto: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25509 (type in U. S. Nat. Herb.).
Peperomia arboriseda C. DC. Bot. Jahrb. 40: 261. 1908.
A glabrous, spreading epiphyte; stem slender, sublignescent;
leaves alternate, lanceolate or the lower suborbicular, acute, round-
based, 4 cm. long, 2 cm. wide, 7-nerved, the inner nerves sub-
confluent below, ciliate; petiole 5 mm., or less than 15 mm., long;
spikes 3 on a slender, terminal, 2-bracted stalk, slender; peduncle
5 mm. long; bracts round-peltate; berries oblong, obliquely scutulate;
stigma central.
Junin: La Merced, Chanchamayo Valley, in forest, 1,000 meters,
Weberbauer 1917, type.
Peperomia arboriseda var. moyobambana C. DC. Bot. Jahrb.
40: 261. 1908.
FLORA OF PERU 23
Internodes 5 cm. long; leaves 6 cm. long, 2.5 cm. wide, with
densely ciliate nerves; scales 15 mm. long subtending the spikes.
San Martin: Moyobamba, Weberbauer 1+691, type.
Peperomia arctebaccata Trelease, sp. nov.
Herba modice parva arete radicans caespitosa dichotoma glabra,
caule 2 mm. crasso firmo plus minus ve furfuraceo-exfoliante; folia
opposita vel ternata vel quaternata elliptica obtuse submucronata
vix 10 mm. longa et 5 mm. lata opaca crassissima revoluta, subtus
in sicco fusca, nervis non manifestis, petiolo 1-2 mm. longo; spicae
terminales 30-50 mm. longae 2 mm. crassae densiflorae, floribus in
alveolis oblongis profundis insertis, pedunculo crasso 5-10 mm.
longo; bracteae minutae rotundo-peltatae; baccae numerosissimae
pseudocupula brevi instructae, stigmate terminali, stylo brevi.
Junin: Chanchamayo Valley, 1,000 meters, Carlos Schunke 471
(herb. Field Mus. No. 571,529, type), A89, A86, 386, 372.
Peperomia arcuatispica Trelease, sp. nov.
Herba majuscula glabra stolonifera, caule 5 mm. crasso; folia
alterna oblanceolata acuta basi cuneata 16-18 cm. longa 6 cm. lata
ubique pinnatinervia, in sicco tenuia viridia, petiolo 1 cm. longo
lamina decurrente alato; spicae paniculatae, panicula foliis vix
longiore, ramis circa 3 basi spicis 1 vel 2 et 2-4 cm. longis atque
curvis basilaribus onustis apice spicas plures gerentibus, spicis 30
mm. longis 1 mm. crassis, pedunculo lere nullo; bracteae rotundo-
peltatae.
Junin: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
25992 (type in U. S. Nat. Herb.).
Peperomia areolata Trelease, sp. nov.
Herba glabra rhizomatosa, caulibus brevibus crassis paucifoliis;
folia reniformi-ovata vel rotundo-ovata apice rotundata vel sub-
acuta vel submucronata, basi breviter aperte cordata, 6-8 cm.
longa, 7-8 cm. lata 7-nervia in sicco venulose pallido-areolata,
petiolo 8-12 cm. longo; inflorescentia non visa.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28660
(type in U. S. Nat. Herb.). Huallaga, Weberbauer 6807.
Peperomia artatiflora Trelease, sp. nov.
Herba modice alta parvifolia succulenta, caule velutino inferne
ad 4-5 mm. crasso in sicco lucido-stramineo ; folia 3-4-nata linearia
ad subobovato-elliptica obtusa vel subacuta basi acutiuscula 6-15
24 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
mm. longa 2-3 mm. lata, utrinque papilloso-subpuberula, 1-nervia,
petiolo 1 mm. longo; spicae terminales et ex axillis superioribus,
40 mm. longae 1 mm. crassae densiflorae, pedunculo 10 mm. longo
glabrato; bracteae rotundo-peltatae brunneae.
Junin: Palca, Stevens 47 (type).
Peperomia asterophylla Trelease, sp. nov.
Herba mediocris stolonifera suberecta caespitosa, caule 2-3 mm.
crasso ad nodos velutino; folia ad nodum 3-5 elliptica obtuse acumi-
nata basi acuta 2-3 cm. longa 1.5-2.5 cm. lata obscure trinervia,
in sicco opaca et minute rugosa glabra vel fere glabra, petiolo 3-4 mm.
longo brunneo- velutino; spicae axillares et terminales 40-80 mm.
longae 2 mm. crassae, pedunculo 10-15 mm. longo glabrato; bracteae
rotundo-peltatae.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28909 (type
in U. S. Nat. Herb.). Sapote Yacu, Santa Rosa, 200 meters,
Williams 4901 . San Antonio, Alto Rio Itaya, 145 meters, Williams
3479.
Peperomia atocongona Trelease, sp. nov.
Herba modice parvula suberecta caespitosa, caule 1-3 mm.
crasso plus minusve ramoso minute puberulo; folia ad nodum 3-6
elliptica vel subobovata obtusa basi acuta 4-6 mm. longa 2-3 mm.
lata, superioribus ad 10-20 mm. longis et 5 mm. latis, subtus obscure
pinnatinervia atque granulosa, petiolo vix 1 mm. longo; spicae
terminales et ex axillis superioribus 15 mm. longae 1 mm. crassae
densiflorae, pedunculo 5 mm. longo; bracteae rotundo-peltatae.
Lima: Atocongo, 250-500 meters, F . W. Pennell 14752 (type in
herb. Acad. Nat. Sci. Phila.).
Peperomia balsapuertana Trelease, sp. nov.
Herba epiphytica majuscula fere acaulescens glabra; folia alterna
elliptico-oblanceolata obtusa ad acute subacuminata basi cuneata
17-25 cm. longa 7-10 cm. lata ubique pinnatinervia, in sicco tenuia,
petiolo 5-6 cm. longo; inflorescentia non visa.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28632
(type in U. S. Nat. Herb.).
Peperomia basellaefolia HBK. Nov. Gen. & Sp. 1: 66. 1815.
Piper basellaefolium Poir. Encycl. Suppl. 4: 468. 1816.
A glabrous, assurgent, succulent herb; stem 3 mm. thick; leaves
alternate, subelliptic, acuminate, acute at the base, 3-5 cm. long,
FLORA OF PERU 25
3-4 cm. wide, ciliate, pinnately few-nerved; petiole 5-10 mm. long;
spikes terminal, when young 25 mm. long, 2 mm. thick, the peduncle
1 cm. long.
Type from Peru, without definite locality, Mathews 1838.
Ecuador.
Peperomia bermudezana Trelease, sp. nov.
Herba glabra repenti-assurgens arboricola, caule 2-3 mm. crasso;
folia alterna elliptico-suboblonga acuminata basi acuta 7-9 cm.
longa 3-4 cm. lata trinervia vel obscure 5-nervia costa obscure
ramosa, in sicco tenuia opaca, subtus plus minusve pallida, petiolo
gracili 1-2 cm. longo; spicae sympodiales interdum in ramis brevibus
2-bracteatis insertae, filiformes, 70 mm. longae, pedunculo 5-10
mm. longo; bracteae rotundo-peltatae.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26451
(type in U. S. Nat. Herb.).
Peperomia binispica Trelease, sp. nov.
Herba mediocris subsimplex glabra, caule 2 vel ad basin 5 mm.
crasso; folia alterna lanceolata 15 cm. longa 4 cm. lata vel majora,
utrinque acuta, apice interdum obtusa, pinnatinervia, petiolo vix
10 mm. longo vel fere nullo; spicae axillares vulgo binae et pedunculos
graciles ad 10 cm. longos terminantes, 100-140 mm. longae 2 mm.
crassae densiflorae, pedunculo 15-25 mm. longo filiformi; bracteae
rotundo-peltatae; baccae rotundato-ovoideae oblique rostro crasso
terminatae, stigmate ad basin rostri inserto.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 358
(herb. Field Mus. No. 571,412, type), A84-
Peperomia blepharipus Trelease, sp. nov.
Planta modice alta epiphytica erecta subsimplex, petiolis crispo-
ciliatis et lineis decurrentibus exceptis glabra, caule 2-3 mm. crasso;
folia alterna lanceolata utrinque acuta 8-10 cm. longa 2-3.5 cm.
lata fere e basi 3-5-nervia costa superne obscure ramosa, in sicco
tenuia opaca, petiolo 5-10 vel ad 30 mm. longo amplexicauli decur-
rente; spicae terminales vel sympodiales 60 mm. longae 2 mm.
crassae, pedunculo 2 cm. longo; bracteae rotundo-peltatae.
Loreto: Yurimaguas, 135 meters, Killip & Smith 28030 (type
in U. S. Nat. Herb.).
Peperomia blepharipus var. binispica Trelease, var. nov.
Planta gracilior, spicis binis ramulum gracili-bracteatum termi-
nantibus 60 mm. longis 1 mm. crassis.
26 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Loreto: San Antonio, Alto Rio Itaya, 145 meters, Williams
3435 (type in herb. Field Mus., No. 618,141).
Peperomia blepharipus var. iquitosensis Trelease, var. nov.
Planta validior, caule 2-4 mm. crasso; folia elliptica vel sub-
ovata 7-9 cm. longa 3-5.5 cm. lata; spicae solitariae.
Loreto: Iquitos, 100 meters, Killip & Smith 27462.
Peperomia brachyiula Trelease, sp. nov.
Herba erecta modice parva dense caespitosa breviter ramosa,
caule gracili 1-2 mm. crasso papilloso-velutino; folia ad nodum
4-6 patentia elliptica vel rotundato-elliptica obtusa vel subretusa
basi plus minusve acuta parva, 6-9 mm. longa, 2-4 mm. lata, sup-
rema magis rotundata, modice tenuia, obscure pinnatinervia, primo
utrinque dense puberula, petiolo vix 2 mm. longo puberulo; spicae
ramos breves dense foliatos terminantes vix 35 mm. longae 1 mm.
crassae laxiflorae, pedunculo 5 mm. longo ; bracteae rotundo-peltatae ;
ovaria inter costulas anastomosantes immersa; baccae rotundato-
ovoideae oblique mucronulato-truncatae; stigma anterius.
Lima: Matucana, steep southern shale slope, in large clumps,
2,400 meters, Macbride 129 (herb. Field Mus. No. 516,662, type).
Peperomia bracteispica Trelease, sp. nov.
Herba parva repenti-assurgens glabra epiphytica, caule 1 mm.
crasso; folia vulgo ternata obovata truncato-emarginulata basi
cuneata 9-11 mm. longa 5-6 mm. lata obscurissime trinervia,
petiolo filiformi 3-4 mm. longo; spicae terminales vix 40 mm. longae
et 1 mm. crassae, floribus sublaxe dispositis, pedunculo 5 mm.
longo folio unico 15 mm. supra verticillos supremos inserto fulcrato;
bracteae rotundo-peltatae; rhachis pseudopedicellis onusta.
Huanuco: Huacachi, near Muna, 1,950 meters, 41 71 a (type in
herb. Field Mus. No. 551,173).
Peperomia caducifolia Trelease, sp. nov.
Herba alta repenti-assurgens, caulibus carnosissimis ramosis
puberulis, internodiis brevibus; folia alterna, foliis ramorum steri-
lium parvis et suborbicularibus, ceteris ellipticis obtuse acuminatis
basi subacutis 3 cm. longis 1.5-2 cm. latis, superioribus minoribus,
caducis, trinerviis, minute puberulis, petiolo 5-10 mm. longo glab-
rescente; spicae plures terminales et ex axillis nudis approximatis
brevem pseudopedunculum efformantibus nascentes 30 mm. longae,
floribus laxiuscule insertis, pedunculo 5 mm. longo; bracteae
rotundo-peltatae.
FLORA OF PERU 27
Ayacucho: Carrapa, 3,000 meters, Killip & Smith 22270 (type
in U. S. Nat. Herb.).
Peperomia calvescens Trelease, sp. nov.
Herba minuscula stolonifera succulenta, caulibus plus minusve
caespitosis erectis subgracilibus 2 mm. crassis primum sparse mol-
liter pilosis cito epidermi furfuraceo-exfoliante glabrescentibus vel
ad nodos persistente pilosis; folia 3-4-nata lanceolato-elliptica
utrinque acuta trinervia parva 16 mm. longa 8 mm. lata supra ad
nervos plus minusve persistente crispo-pubescentia, subtus glabra
et epidermi exfoliato albida, petiolo brevi 2 mm, longo glabro;
spicae terminales 60 mm. longae 1 mm. crassae, floribus laxiuscule
insertis, pedunculo filiformi 2 mm. longo glabro; bracteae rotundo-
peltatae; ovarium impressum ovoideum, stigmate subapicali, rhachi
pseudopedicellis brevibus onusta.
Huanuco: Cueva Grande near Posuso, 1,050 meters, sunny clay
banks, Macbride 4800 (herb. Field Mus. No. 535,834, type).
Peperomia caniana Trelease, sp. nov.
Herba modice parva repenti-assurgens, caule 2-4 mm. crasso
crispo-villoso; folia opposita orbicularia, superiora late ovata,
utrinque obtusa vel rotundata, inferiora submucronata, minuscula,
12-25 mm. longa 12-20 mm. lata, trinervia arctiuscule adpresso-
pilosa vel supra glabrescentia, petiolo vix 5 mm. longo crispo-piloso ;
spicae terminales subumbellatae 60 mm. longae 1 mm. crassae laxi-
florae, pedunculo 1.5-2 cm. longo laxe piloso; bracteae rotundo-
peltatae; baccae in rhachi immersae globulosae ovoideae obtusae,
stigmate subapicali.
Huanuco: Cani, near Mito, 2,550 meters, Macbride 3415 (herb.
Field Mus. No. 534,486, type).
Peperomia carpapatana Trelease, sp. nov.
Herba majuscula ramosa, caule 2-5 mm. crasso praesertim
superne dense luteo-intertexto-piloso; folia alterna rotundo-ovata
apice rotundata et subabrupte obtuso-acuminata, basi rotundata
vel abrupte subacuta, 2-4 cm. longa 2-3.5 cm. lata, obscure 3-5-
nervia subtus adpresso-pilosa et concolori-granulosa in sicco opaca,
petiolo 5-10 mm. longo adpresso-piloso ; spicae ad axillas aggregatae
graciles 50-150 mm. longae, floribus sublaxe insertis, pedunculo
gracili 10-15 mm. longo plus minusve adpresso-piloso; bracteae
rotundo-peltatae; baccae parvae globosae apice mucronatae, stigmate
subapicali.
28 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Junin: Carpapata, 2,700-3,200 meters, Killip & Smith 24438
(U. S. Nat. Herb., type), 24393.
Peperomia carrapana Trelease, sp. nov.
Planta parva epiphytica rhizomatoso-adscendens glabra, caule
1-2 mm. crasso; folia alterna lanceolata vel lanceolato-oblonga sub-
obtusa vel attenuata, basi subacuta, 2-4 cm. longa, 1-1.5 cm. lata,
obscure trinervia, in sicco opaca tenuia aliquanto crispo-revoluta,
petiolo 5 mm. longo; spicae terminales 60 mm. longae 2 mm. crassae,
floribus laxiuscule insertis, pedunculo 5 mm. longo; bracteae rotundo-
peltatae.
Ayacucho; Carrapa, 1,500 meters, Killip & Smith 23207 (U. S.
Nat. Herb., type).
Peperomia casapiana C. DC. Journ. Bot. 4: 146. 1866.
A pilose herb; leaves 3 at a node, ovate, acuminate, rounded at
the base, 5 cm. long, 3 cm. wide, 5-nerved, drying pellucid; petiole
8 mm. long; spikes subumbellately verticillate from the axils of the
reduced upper leaves, 50 mm. long, 1 mm. thick; peduncle 15 mm.
long; bracts round-peltate; berries ovoid, mucronate; stigmas sub-
apical.
Huanuco: Casapi, Mathews 1689, type.
Peperomia ceapanana Trelease, sp. nov.
Herba erecta plus minusve caespitosa breviter ramosa, caule
2-3 mm. crasso dense papilloso-puberulo; folia ad nodum circa 4
adscendentia elliptica vel elliptico-obovata obtusa basi acuta parva
10 mm. longa 4 mm. lata subtenuia obscure pinnatinervia glabra,
subtus glanduloso-granulosa, petiolo 1 mm. longo papilloso-puberulo;
inflorescentia non visa.
Cuzco: Hacienda Ceapana, Paucartambo Valley, 3,450 meters,
F. L. Herrera 1468 (U. S. Nat. Herb., type).
Peperomia cerea Trelease, sp. nov.
Herba modice alta succulenta glabra e rupibus pendula, caule
valido 5 mm. crasso, superne graciliore; folia opposita vel superne ad
nodum 3-5 rotundato-elliptica utrinque rotundata circa 15 mm. longa
et 10 mm. lata, superiora magis oblonga et minora, vix 10 mm.
longa et 7 mm. lata, margine tenui plus minusve revoluto, in sicco
cerino-translucentia, 1-nervia, petiolo brevi 1-3 mm. longo; spicae
terminales 100 mm. longae et ultra 3-4 mm. crassae obscure rubrae,
floribus laxiuscule insertis, pedunculo crasso 15 mm. longo; bracteae
FLORA OF PERU 29
rotundo-peltatae ; ovaria in rhachi immersa acuminata, stigmate
subapicali.
Huanuco: Mito, 2,700 meters, Macbride 3330 (type in herb.
Field Mus. No. 534,400).
Peperomia chachapoyasensis C. DC. in DC. Prodr. 16, pt.
1: 449. 1869. P. umbellata Miq. Lond. Journ. Bot. 4: 427. 1845,
non Kunth.
A delicate, suberect, glabrous herb, the stem almost filiform;
leaves alternately clustered at the apex, round-ovate, very obtuse,
rounded at the base or obscurely cordulate, 2-2.5 cm. long, 2 cm.
wide, 3- or somewhat 5-nerved; petiole 5 mm. long; spikes umbellate
at the ends of slender, terminal or axillary, 2-bracted stalks 3-4 cm.
long, scarcely 15 mm. long and 1 mm. thick; peduncle 5 mm. long;
bracts round-peltate; berries round-ovoid, mucronate; stigma sub-
apical. Neg. 8394.
Amazonas: Seringal Chachapoyas, Mathews 3230, type; 1862.
Peperomia chalhuapuquiana Trelease, sp. nov.
Herba modice parvula erecta divaricate multiramosa, caule 2-3
mm. crasso crispo-pubescente; folia alterna ovata vel elliptico-ovata
obtuse subacuminata basi vulgo rotundata 2-4 cm. longa 1.5-2.5
cm. lata ciliata, in sicco subcoriacea, subtus pallidiora minute nigro-
glandulosa, petiolo 2-3 vel ad 10 mm. longo glabrato; spicae termin-
ales et ex axillis superioribus 100 mm. longae 2 mm. crassae, floribus
laxiuscule vel verticillatim insertis, pedunculo gracili vix 10 mm.
longo glabro; bracteae rotundo-peltatae; baccae subglobosae mucro-
natae in sicco nigrae, stigmate obliquo.
Junin: Hacienda Chalhuapuquio, Stevens 215, type.
Peperomia chanchamayana Trelease, sp. nov.
Herba epiphytica modice parva rhizomatosa simplex glabra,
caule 2 mm. crasso; folia alterna ovata vel lanceolata utrinque
acuta 2.5-3.5 cm. longa 1-1.5 cm. lata trinervia vel obscure 5-nervia
subtus pallidiora coriacea aliquanto crispo-revoluta, petiolo 1 cm.
longo amplexicauli ; spicae terminales 60-90 mm. longae 1 mm.
crassae densiflorae, pseudopedicellis brevissimis, pedunculo 5 mm.
longo; bracteae rotundo-peltatae; baccae subglobosae oblique apicu-
latae, stigmate subapicali.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 371
(herb. Field Mus. No. 571,425, type); at 1,500 meters, Schunke
1554.
30 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia chartacea Trelease, sp. nov.
Herba caespitosa epiphytica modice alta glabra, caule 2-3 mm.
crasso nodoso; folia alterna elliptico-subobovata acuta vel submu-
cronata basi cuneata 6-11 cm. longa 3-6 cm. lata essentialiter pal-
matim 7-nervia vel nervis minoribus plus minusve circa basin
confluentibus, in sicco firmo-papyracea, petiolo 5-15 mm. longo;
spicae solitariae et terminates 20 mm. longae 2 mm. crassae,
pedunculo gracili aequilongo; bracteae rotundo-peltatae.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26636
(type in U. S. Nat. Herb.).
Peperomia chillonensis Trelease, sp. nov.
Herba modice parva subsimplex(?) griseo-puberula ; folia ad
nodum 8 et ultra elliptica vel subobovata obtusa basi subacuta opaca
obscure 3-5-nervia subrevoluta, 10 mm. longa 5 mm. lata, petiolo
1 mm. longo; spicae axillares vel ramulos brevissimos terminantes 20
mm. longae 1 mm. crassae, floribus laxiuscule insertis; bracteae ro-
tundo-peltatae; ovarium ovoideum, stigmate subapicali.
Lima: Along Rio Chillon, above Obrajillo, 2,800-3,200 meters,
open rocky banks, F. W. Pennell 14413 (herb. Field Mus. No.
558,460, type).
"A fleshy herb with greenish yellow spikes."
Peperomia chrysotricha Miq. Syst. 163. 1843.
A rather small, erect, succulent herb, yellow-puberulent or tomen-
tulose; leaves commonly 4 at a node, subreflexed, round-obovate or
elliptic, rounded at both ends, 6-10 mm. long, 3-6 mm. wide, cori-
aceous, 1- or obscurely 3-nerved; petiole very short; spikes terminal
and from the upper axils, 20-40 mm. long; peduncle 5-10 mm. long;
bracts round-peltate, pale-margined; berries globose: stigma apical.
Type from Peru, Dombey 933.
Peperomia ciliatifolia Trelease, sp. nov.
Herba pendula arboricola foliis sparse pilosis et rigido-ciliatis ex-
ceptis glabra, caule gracili 2 mm. crasso; folia alterna elliptica acute
acuminata basi rotundata vel subacuta 11 cm. longa 5 cm. lata infra
medium pinnatim paucinervia in sicco tenuia translucentia, petiolo
vix 10 mm. longo; spicae sympdiales geminatae graciles 150 mm.
longae, pedunculo vix 5 mm. longo; bracteae rotundo-peltatae;
baccae oblongae vel subconicae scutulo acuto stigma gerente oblique
truncatae.
FLORA OF PERU 31
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28684-
(type in U. S. Nat. Herb.). Yurimaguas, Killip & Smith 27982.
Mishuyacu, near Iquitos, 100 meters, Klug 58.
Peperomia ciliatifolia var. eciliatifolia Trelease, var. nov.
Planta glabra vel sparse obscure ciliata, aliter ut in forma typica.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28723 (type in
U. S. Nat. Herb.).
Peperomia ciliatifolia var. iquitosana Trelease, var. nov.
Magis glabrescens, spicis filiformibus.
Loreto: Iquitos, Killip & Smith 27167 (type in U. S. Nat. Herb.).
Peperomia ciliatifolia var. santarosana Trelease, var. nov.
Folia minora, 6 cm. longa 3 cm. lata.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28988 (type in
U. S. Nat. Herb.).
Peperomia Claudii C. DC. Ann. Cons. Jard. Bot. Geneve 2:
285. 1898.
A small, cespitose, spreading epiphyte; stem filiform, puberulent;
leaves 3 at a node, round-obovate, acute-based, 6 mm. long, 5 mm.
wide, obscurely 3-nerved, glabrous; spikes terminal, 10 mm. long,
1 mm. thick; peduncle very short; bracts round-peltate; berries ovoid,
pointed; stigma apical.
Type collected in Peru by Gay.
Peperomia coloniae Trelease, sp. nov.
Herba suffruticosa repenti-assurgens transiente sparse molliter
pubescens; folia alternalanceolataplus minusve acuminatabasi obtusa
5-6 cm. longa 2.5-3 cm. lata coriacea opaca obscure pinnatinervia,
subtus flavescentia, petiolo 10 mm. longo; spicae terminales, solitariae
vel geminatae et pedunculos breves bibracteatos terminantes, breves.
Junin: Colonia Perene", 680 meters, Killip & Smith 25053 (type
in U. S. Nat. Herb.).
Peperomia concava R. & P. Fl. 1: 31. pi. 46. 1798. Piper
concavum Vahl, Enum. 1: 350. 1805.
A moderately small, cespitose, somewhat branched, succulent
herb; stem 2-3 mm. thick; leaves 2-4 at a node, elliptic, rounded at
the apex, acutely subsessile, 2.5 cm. long, 1.5 cm. wide, concave,
with 5 red nerves; spikes terminal, 60 mm. long, 3-4 mm. thick.
Huanuco : Chinchao and Cochero, Ruiz & Pavon, type.
32 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia confertispica Trelease, sp. nov.
Herba modice parva procumbenti-assurgens obscure viridis,
caule 2-3 mm. crasso, internodiis inferioribus longis sordide molliter
hirsutis; folia alterna, superne in caulibus erectis opposita, rotundata
vel late ovata, apice rotundata vel obtusa, basi rotundata, modice
parva, 1.5-3.5 cm. longa, 1.5-3.5 cm. lata, 3-5-nervia, subtus palli-
diora, utrinque sparse praesertim ad nervos adpresso-pilosa, petiolo
1-1.5 cm. longo dense piloso; spicae solitariae vel plures in axillis
superioribus, juveniles vix 15 mm. longae et 1 mm. crassae densi-
florae, pedunculo gracili vix 5 mm. longo piloso; bracteae rotundo-
peltatae majusculae flavae.
Huanuco: Mito, 2,700 meters, Macbride 1466 (herb. Field Mus.
No. 518,142, type). "Plant dark green, sprawling on the deep
humus floor of a stream thicket."
Peperomia confertispica var. erecta Trelease, var. nov.
Planta stolonifera erecta, caulibus crassioribus 5 mm. crassis in
sicco plus minusve quadrangularibus; folia elliptica vel obovata vix
3 cm. longa et 2 cm. lata, superiora 2 cm. longa 1.5 cm. lata, subtus
flavescentia, in sicco plus minusve supra rugosa vel granulosa,
petiolo vix 1 cm. longo; spicae juveniles vix 25 mm. longae, pedunculo
ad 5 mm. longo; bracteae primo flavescentes, brunnescentes.
Huanuco: Mito, 2,700 meters, Macbride 1915 (herb. Field Mus.
No. 518,410). "Stems and upper leaves beneath dark red. Bracts of
inflorescence pale yellowish at the tip. Leaves dark green, glossy
above, the lower pale beneath."
Peperomia congesta HBK. Nov. Gen. & Sp. 1: 68. pi. 14.
1815.
A rather large, erect, glabrous herb; stem 5 mm. thick below,
with short internodes; leaves 5 to 7 at a node, round-ovate, short-
acuminate, acute at the base, 1.5-2.5 cm. long, 1-2 cm wide, 1-nerved ;
petiole 5 mm. long; spikes numerous in the axils of congested whorls
at the end of the stem so as to appear subpaniculate, 15-20 mm.
long, 1 mm. thick; peduncle 5 mm. long.
Piura : Rio Chuchancanao to Huancabamba, Bonpland, type.
Peperomia congestispica Trelease, sp. nov.
Herba mediocris assurgens plus minusve ramosa in sicco nigres-
cens, caule 1-4 mm. crasso dense molliter f errugineo-hirsuto ; folia ad
nodum circa 5 elliptica vel subobovata subacuta basi acuta 12-20
mm. longa 6-12 mm. lata obscure trinervia breviter hirtella, supra
FLORA OF PERU 33
glabrescentia, petiolo 2-4 mm. longo f errugineo-hirtello ; spicae
terminates et in axillis superioribus aggregatae, juveniles 10 mm.
tan turn longae et 1 mm. crassae breviter pedunculatae; bracteae
rotundo-peltatae.
Junin: Huacapistana, Stevens 62, type.
Peperomia cordulilimba C. DC. Bot. Jahrb. 40: 261. 1908.
A small epiphyte; stem slender, velvety; leaves alternate, obovate,
emarginulate, acute-based, 7 mm. in diameter, 3-nerved, puberulent
beneath; petiole barely 5 mm. long, puberulent; spikes terminal,
filiform, 15 mm. long; peduncle very short; bracts round-peltate;
ovary obovoid; stigma oblique.
Puno: Near Tambo Frilluma, road from Sandia to Chunchus-
mayo, 1,000 meters, Weberbauer 1197, type.
Peperomia cotoneasterifolia Trelease, sp. nov.
Herba arboricola modice alta plus minusve ramosa, caule 2-3
mm. crasso primo dense adpresso-villoso; folia opposita elliptico-
subovata utrinque abrupte subacuta 2-2.5 cm. longa 1.5-2 cm. lata
3- vel obscure 5-nervia supra glabrata, subtus pallidiora tomentosa,
petiolo 5 mm. longo adpresso-piloso ; spicae axillares 60 mm. longae
2 mm. crassae, pedunculo 10 mm. longo; bracteae rotundo-peltatae;
baccae ovoideae acutiusculae, stigmate apicali.
Junin: Carpapata, 2,700-3,200 meters, Killip & Smith 24390
(type in U. S. Nat. Herb.).
Peperomia crassulaecaulis Trelease, sp. nov.
Planta glabra succulenta suffrutescens, caulibus 3-4 mm. crassis
erectis superne sparse ramosis sed e basi ad 2 cm. crassa numerosis
flavescentibus, internodiis brevibus; folia ad nodum 6 elliptica emar-
ginulata basi acuta 10 mm. longa 4-5 mm. lata in sicco dura subre-
voluta opaca obscure nervata, subtus minute granulosa, petiolo 2
mm. longo; spicae terminales 140 mm. longa 4 mm. crassae, floribus
laxiuscule insertis, pedunculo 15 mm. longo; bracteae rotundo-
peltatae; flores in alveolis immersae; ovarium ovoideum, stigmate
obliquo.
Junin: Carpapata, 2,400 meters, Killip & Smith 24355 (type in
U. S. Nat. Herb.; dupl. in herb. Field Mus.).
Peperomia crinigera Trelease, sp. nov.
Herba modice alta plus minusve ramosa assurgens, caule 2-3
mm. crasso rigide crispo-piloso ; folia ad nodum circa 5 elliptico-
34 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
subobovata obtusa basi subacuta 10-16 mm. longa 6-8 mm. lata
obscure trinervia crispo-hispida, petiolo 3 mm. longo crispo-pilosa ;
spicae terminales et ex axillis superioribus 60 mm. longae 1 mm.
crassae, floribus laxiuscule insertis, pedunculo 1-2 cm. longo rigide
crispo-piloso ; bracteae rotundo-peltatae.
Junin: Palca, Stevens 45, type. Carpapata, Killip & Smith
24449.
Peperomia crotalophora Trelease, sp. nov.
Herba modice alta procumbens glabra in sicco viridis, caule
subgracili 2-3 mm. crasso, internodiis elongatis; folia alterna rotun-
dato-ovata ad subtriangulari-ovata acuta vel interdum obtusa vel
breviter acuminata mediocria 5.5-6.5 cm. longa 4.5 cm. lata
multiplinervia, nervis basilaribus 4, alteris 2 e quarto inferiore
nascentibus, petiolo 3 cm. vel ad 10 cm. longo; spicae terrrinales
solitariae pedunculos unibracteatos 2 cm. longos terminantes, 50-70
mm. longae 1 mm. crassae, floribus arete verticillatis, verticillis
antherarum magnarum dense aggregatis valde in statu juvenili
conspicuis, pedunculo 15 mm. longo; bracteae rotundo-peltatae
obscurae in pseudoverticillo circa 12.
Junin: Hacienda Schunke, La Merced, 1,200 meters, Macbride
5644 (type in herb. Field Mus. No. 536,682), terrestrial, growing on
forest floor.
Peperomia cruciata Trelease, sp. nov.
Herba glabra rhizomatoso-assurgens; folia alterna obovato-sub-
spathulata obtusa basi longicuneata 6-11 cm. longa 3-4.5 cm. lata
pinnatinervia, in sicco subtenuia, petiolo 2-4 cm. longo; spicae ter-
minales 110 mm. longae 2 mm. crassae, pedunculo 8-9 cm. longo
1-bracteato; bracteae spicae rotundo-peltatae minutae; baccae
oblongae graciliter rostratae, stigmate anteriore.
Junin: Chanchamayo Valley, 800 meters, Carlos Schunke 343
(type in herb. Field Mus. No. 571,399).
Peperomia crusculibacca Trelease, sp. nov.
Herba modice alta assurgens, caule 2-4 mm. crasso crispo-
pubescente; folia alterna elliptica subacuminata basi acuta 4-6 cm.
longa 2-3 cm. lata 3- vel obscure 5-nervia glabra subtus nigro-
glandulosa, petiolo gracili 10-15 mm. longo glabrato; spicae
(terminales? et) axillares filiformes 100 mm. longae plus minusve
laxiflorae, pseudopedicellis gracilibus, pedunculo 10 mm. longo gla-
FLORA OF PERU 35
brato; bracteae rotundo-peltatae; baccae subglobosae oblique mucro-
natae stigmate subapicali.
Junin: Huacapistana, Stevens 90, type; Killip & Smith 24305.
Peperomia crystallina R. & P. Fl. 1: 30. 1798. Piper crys-
tallinum Vahl, Enum. 341. 1805.
A glabrous, branching, anise-scented succulent with 4-sided,
pellucid stem; leaves alternate, oblong, papillate above, impressed-
punctulate beneath; spikes slender, loosely flowered.
Lima: Lima, Ruiz & Pavon, type.
A very obscure species, scarcely comparable with anything known
except P. metallica.
Peperomia cuprea Trelease, sp. nov.
Herba mediocris repens, caule 2 mm. crasso obscure evanescente
plus minusve crispo-pubescente; folia alterna ovata subacuminata
basi rotundata 3.5-4.5 cm. longa 2-3.5 cm. lata, 3-5 mm. supra
basin peltata, opaca coriacea obscure multiplinervia anguste revoluta
subtus cuprea sericeo-ciliata, petiolo 1-2.5 cm. longo molliter evan-
escenti-piloso ; spicae axillares 55 mm. longae 3 mm. crasse arete
zonatiflorae, pedunculo 2 cm. longo glabro; bracteae parvae rotundo-
peltatae in pseudoverticillo circa 20.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 354
(herb. Field Mus. No. 571,408, type), 1500.
Peperomia cuprea var. cordulifolia Trelease, var. nov.
- Herba mediocris repens, caule 1 mm. crasso evanescentissime plus
minusve crispo-sericeo ; folia alterna ovata subacuminata breviter
cordata 3.5-5.5 cm. longa 2.5-3.5 cm. lata circa 5 mm. supra basin
peltata, opaca coriacea subtus brunnescentia obscure multiplinervia
anguste revoluta ciliata, petiolo 1-5 cm. longo sparse minute piloso;
spicae ramulos graciles 1-bracteatos terminantes statu juvenili
50 mm. longae 2 mm. crassae dense zonatiflorae, pedunculo 10-15
mm. supra folium reductum longo; bracteae spicae rotundo-
peltatae parvae in pseudoverticillo circa 20.
Junin: Chanchamayo Valley, 1,500 meters, Carlos Schunke
379 (herb. Field Mus. No. 571,433, type), 1466; Killip & Smith
24839. Enefias, Killip & Smith 25672.
Peperomia currucaeformis Trelease, sp. nov.
Herba terrestris repens glabra; folia late ovata vel deltoideo-ovata
obtusa basi truncato-cordata 5-6 cm. longa 4-6 cm. lata 7-nervia,
36 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
nervis intimis lateralibus con costa confluentibus, in sicco opaco-
olivacea granulosa, petiolo 2-3 cm. longo; spicae ad apicem caulis
sympodialis 4 cm. longi divergente geminatae et sursum curvatae
60-70 mm. longae 3 mm. crassae, pedunculo 15-20 mm. longo;
bracteae rotundo-peltatae; baccae ellipsoideae rostratae, stigmate
ad basin rostri inserto.
Junin: Schunke Hacienda, La Merced, 1,400-1,700 meters, Killip
& Smith 24542 (type in U. S. Nat. Herb.).
Peperomia curticaulis Trelease, sp. nov.
Herba glabra, caule brevi crasso paucifoliato ; folia alterna late
ovata obtusa, basi rotundata vel brevi ter cordata, 7-10 cm. longa
7-9 cm. lata 7-nervia, costa ut quoque nervis lateralibus ramosis,
petiolo 7 cm. longo; spicae terminales 120 mm. longae 3 mm. crassae,
floribus subdistantibus, pedunculo 1-2 cm. longo; bracteae rotundo-
peltatae; ovarium ovoideum submucronatum, stigmate obliquo.
Junin: Cahuapanas, Rio Pichis, 30 meters, Killip & Smith 26773
(type in U. S. Nat. Herb.).
Peperomia curtipes Trelease, sp. nov.
Herba subacaulescens glabra; folia alterna rotundata subacu-
minata basi sinu fere clauso cordata 8-13 cm. longa 8-12 cm. lata
palmatim 9-nervia vel costa interdum superne obscure ramosa, in
sicco fusca, petiolo 15-20 cm. longo; spicae 50 mm. longae 2 mm.
crassae vel ma j ores, pedunculo ut videtur elosgato; bracteae rotundo-
peltatae; ovarium globosum, stigmate subapicali.
Junin: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
26019 (type in U. S. Nat. Herb.). East of Quimiri Bridge, 900
meters, Killip & Smith 30679.
Peperomia curtipes var. contracta, Trelease, var. nov.
Caulis foliatus brevissimus, foliis congestis, petiolo vix 15 cm.
longo; spicae 100 mm. longae 3 mm. crassae vel ma j ores, laxiflorae,
pedunculo 15 cm. longo; bracteae rotundo-peltatae; ovarium rotunda-
turn mucronatum, stigmate obliquo.
Junin: La Merced, Schunke Hacienda, 1,400-1,700 meters, Killip
& Smith 24726 (type in U. S. Nat. Herb.).
Peperomia cushiana Trelease, sp. nov.
Herba subparva caespitosa plus minusve furcata succulenta
rubescens glabra, in sicco fusca, caule vix 2 mm. crasso, internodiis
brevibus plus minusve furfuraceo-exfoliantibus; folia vulgo ternata
elliptica utrinque acuta parva 10-15 mm. longa 5-7 mm. lata carno-
FLORA OF PERU 37
sissima in sicco plicata 1-nervia, petiolo brevissimo 1-2 mm. longo;
spicae terminates et ex axillis superioribus 60 mm. longae 1.5 mm.
crassae densiflorae, pedunculo 10-17 mm. longo; bracteae rotundo-
peltatae; ovarium immersum ovoideo-acuminatum, stigmate fere
apicali.
Huanuco: Cushi, 1,500 meters, Macbride 4816 (type in herb.
Field Mus. No. 535,850).
Peperomia cusilluyocana Trelease, sp. nov.
Herba stolonifera modice parva plus minusve ramosa glabra,
caule 2-3 mm. crasso; folia alterna oblanceolata obtusa vel subacu-
minata basi cuneata circa 2.5 cm. longa et 1.5 cm. lata e dimidio
inferiore submultiplinervia, ramulis costae 4, firma olivacea, petiolo
vix 5 mm. longo amplexicauli ; spicae terminales 30 mm. longae 1
mm. crassae densiflorae, pedunculo filiformi 10-15 mm. longo; brac-
teae rotundo-peltatae.
Cuzco: Pillahuata, Cerro de Cusilluyoc, 2,400 meters, Pennell
14016 (herb. Field Mus. No. 558,254, type).
Peperomia cyclophylla Miq. in Seem. Bot. Voy. Herald 198.
1856; Dahlst. Svensk. Vet. Akad. Handl. 33, pt. 2: pi. 5.
A small, repent, more or less crisp-pilose herb; stem filiform;
leaves opposite, orbicular, scarcely 10 mm. in diameter; petiole
almost suppressed; spikes terminal, 30-65 mm. long, 1 mm. thick;
peduncle 1 cm. long, 2-bracted at the middle; bracts of the spike
round-peltate; berries ovoid, pointed; stigma apical. Neg. 8306.
San Martin: Tarapoto, Spruce 4987, type. Junin: Puerto Lira,
Killip & Smith 26830. Rio Pinedo, Killip & Smith 23614. Loreto:
Puerto Arturo, Killip & Smith 27248, 27723.
Peperomia dasystachya Miq. Lond. Journ. Bot. 4: 416. 1845.
A moderate-sized, erect, short-velvety herb; stem 2 mm. thick;
leaves alternate, elliptic or subovate, blunt-acuminate, subacute at
the base 3-5 cm. long, 1-3 cm. wide, or sometimes 8 cm. long and 3.5
cm. wide, 5-nerved, subappressed-pubescent but the upper side
glabrate in age; petiole barely 10 mm. long; spikes terminal, 60 mm.
long, 2 mm. thick, velvety; peduncle scarcely 1 cm. long; bracts
round-peltate; stigma subapical. Neg. 8376.
Amazonas: Chachapoyas, Mathews 3229, type.
Peperomia debilipes Trelease, sp. nov.
Planta epiphytica parva rhizomatoso-assurgens glabra, caule
2-3 mm. crasso; folia alterna oblanceolato-obovata obtusa basi
38 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
cuneata 2-3 cm. longa 1-2 cm. lata 5-nervia, in sicco opaca olivacea,
in vivo subtus rubro-venosa, petiolo gracillimo 10-15 mm. longo;
spicae terminates 45 mm. longae 3 mm. crassae, pedunculo filiformi
2 cm. longo, pseudopedicellis brevibus; bracteae rotundo-peltatae.
Junin: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25545 (type in U. S. Nat. Herb.).
Peperomia debilipes var. dimorpha Trelease, var. nov.
Herba parva repens arboricola paullo crispo-pubescens vel glab-
rescens; folia alterna, ramulorum sterilium rotundo-elliptica et 5-10
mm. longa, cetera subovato-elliptica et usque ad 25 mm. longa atque
15 mm. lata, obscure 3-nervia, petiolo 5 mm. longo.
Junin: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25591 (type in U. S. Nat. Herb.).
Peperomia deceptrix Trelease, sp. nov.
Planta glabra epiphytica, ut videtur repens et radicans ex nodis
4-foliatis florifera; folia oblanceolata acuta basi cuneata 11-12 cm.
longa 3-3.5 cm. lata 7-9-nervia in sicco opaca viridia et subtenuiter
papyracea, petiolo 4-9 cm. longo; spicae (delapsae) ut videtur ad
apicem ramuli 5-6 cm. longi solitariae, juveniles 40 mm. longae 1 mm.
crassae, pedunculo filiformi 2 cm. longo; bracteae rotundo-peltatae.
Loreto: San Antonio, 110 meters, Killip & Smith 29325 (type in
U. S. Nat. Herb.).
Peperomia deficiens Trelease, sp. nov.
Herba modice alta simplex glabra, caulibus gracilibus; folia
alterna subdeltoideo-ovata basi rotundata vel subtruncata et dis-
tincte peltata vel breviter cordata et epeltata 5-5.5 cm. longa 3.5-4.5
cm. lata circa 7-nervia, in sicco tenuia subtus brunneo-granulosa,
petiolo gracili 2-4 cm. longo; inflorescentia non visa.
San Martin: Alto Rio Huallaga, 360-900 meters, Williams
5758a (herb. Field Mus. No. 624,781, type).
Peperomia dendroides Trelease, sp. nov.
Herba parva stolonifera epiphytica superne dense ramosa, caule
gracili 1-2 mm. crasso primo puberulo; folia vulgo 3-4-nata elliptico-
obovata vel oblongo-obovata obtusa basi acuta parva 6-9 mm.
longa 3-4 mm. lata cito glabrescentia valde indistincte pinnatinervia,
petiolo 1-2 mm. longo primo velutino; spicae terminales vix 25 mm.
longae et 1 mm. crassae, floribus laxiuscule insertis, pedunculo vix
5 mm. longo sulcato glabrato; bracteae rotundo-peltatae; baccae
FLORA OF PERU 39
semi-immersae rotundo-ovoideae rubrae scutulate subtruncatae,
stigmate anteriore.
Junin: Chahua, 2,700 meters, Macbride 3640 (herb. Field Mus.
No. 534,704, type). Mito, 1784.
Peperomia dendromorphis Trelease, sp. nov.
Herba parvula rhizomatosa, caulibus erectis superne ramosis
gracilibus 1-2 mm. crassis puberulis; folia inferne opposita vel vulgo
ad nodum 3-5 lanceolato-oblonga vel elliptico-subobovata obtusa
basi acuta 8-10 mm. longa 3-4 mm. lata transiente puberula obscure
pinnatinervia, petiolo 1 mm. longo; spicae terminales filiformes 20-
30 mm. longae in sicco fuscae, pedunculo 5-10 mm. longo; bracteae
rotundo-peltatae.
Cuzco: Marcapata, 3,000 meters, Weberbauer 7800 (herb. Field
Mus. No. 605,350, type).
Peperomia deodorata Trelease, sp. nov.
Herba modice parva rhizomatoso-assurgens glabra carnosa; folia
alterna elliptica vel subobovata obtusa vel emarginulata basi cuneata
4-7 cm. longa 2-3 cm. lata pinnatinervia opaca, petiolo 1-2 cm. longo;
spicae 1-2 pedunculum terminalem 1-bracteatum terminantes vel
ex axillis superioribus nascentes 120 mm. longae 2 mm. crassae,
pedunculo 15 mm. longo; bracteae rotundo-peltatae majusculae;
baccae ellipsoideae, rostro brevi crasso, stigmate anteriore.
Junin: Chanchamayo Valley, 1,500-1,800 meters, Carlos Schunke
381 (herb. Field Mus. No. 571,435, type), 551, 1569, 1493, 1584.
In its dried state this plant lacks the usual fragrance of P. mag-
noliaefolia and its allies.
Peperomia dependens R. & P. Fl. 1 : 33. pi. 48. 1798. Piper
dependens Vahl, Enum. 1: 338. 1805.
A rather large, glabrous herb; leaves alternate, ovate, acuminate,
cordate, supeltate at the very base, 15-16 cm. long, 9-13-nerved,
drying pellucid; petiole 10 cm. long; spikes paired in the axils of
reduced upper leaves, 80 mm. long, 3 mm. wide; peduncle 2 cm.
long; bracts ovate-peltate; berries ovoid; stigmas subapical. Neg.
8379.
Huanuco: Posuso, Pavon 36; Ruiz, type.
Peperomia Diehliana Trelease, sp. nov.
Herba parva fragilis succulenta ramosa, caule 1-2 mm. crasso
primo plus minusve hirtello; folia ad nodum circa 3 rotundo-elliptica
40 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
paullo apiculata vel reflexo-emarginulata basi subobtusa 6 mm. longa
5 mm. lata glabra, petiolo 1-2 mm. longo sparse hirtello; inflores-
centia non visa.
Type in herb. Field Mus. (No. 630,026), probably collected in
the Department of Cuzco, Adolfo Diehl 2526.
Similar to P. rosea.
Peperomia distractiflora Trelease, sp. nov.
Herba modice parva, caule 1-5 mm. crasso puberulo sed
glabrescente et propter exfoliationem lucido; folia ad nodum 3-5
elliptica vel superiora oblonga utrinque subobtusa 10 mm. longa
3-4 mm. lata, supra obscure papillosa, subtus pallidiora glanduloso-
granulosa, petiolo 1 mm. longo; spicae terminales et ex axillis
superioribus 30-50 mm. longae 1 mm. crassae in statu adulto
laxiflorae, pedunculo 5 mm. longo glabro; bracteae rotundo-peltatae
virides; ovarium immersum ovoideum, stigmate obliquo.
Junin: Hacienda Chalhuapuquio, Stevens 214, type.
Peperomia dolabriformis HBK. Nov. Gen. & Sp. 1: 60. pi. 4..
1815. Piper dolabriforme Poir. Encycl. Suppl. 4: 464. 1816.
A rather tall, simple, glabrescent, succulent herb; stem 5 mm.
thick; leaves alternate, compressed from the sides so as to stand
vertically and not horizontally, spatulate or dolabriform, more or
less obliquely submucronate, grooved on the upper edge, about 3
cm. long and 1.5 cm. wide, cuneately subsessile, coriaceous; spikes
10-60 mm. long, 1 mm. thick, in a panicle long-stalked above the
leaves; bracts round-peltate.
Piura: Huancabamba, Bonpland, type. Without locality, Weber-
bauer 6061 .
Peperomia dolabriformis var. velutina Trelease, var. nov.
Herba, caule dense velutino; folia 2.5 cm. longa 1 cm. lata;
spicae 30 mm. longae 1 mm. crassae.
Cajamarca: Cajamarca, 500-600 meters, Weberbauer 3797, type.
Peperomia Drapeta Trelease, sp. nov.
Herba modice parva diffusa arboricola, caule in statu sicco
anguloso subgracili 1-2 mm. crasso, ramis dense intertextis, sterilibus
ad apices crispo-villosis; folia alterna rotunda vel ovata et subacum-
inata, longiora basi subacuta, parva 1-3.5 cm. longa 1.5-2 cm. lata,
in sicco subopaca, 3-5-nervia, supra crispo-pubescentia vel nervis
exceptis glabrescentia, petiolo 1 cm. longo adpresso-piloso ; spicae
FLORA OF PERU 41
terminales et axillares 30-50 mm. longae 1 mm. crassae laxiflorae,
pedunculo 5 mm. longo sparse molliter piloso vel glabrato; bracteae
rotundo-peltatae majusculae; baccae rotundato-ovoideae mucrona-
tae, stigmate subapicali.
Huanuco: Huacachi near Muna, 1,950 meters, Macbride 4122
(herb. Field Mus. No. 535,193, type).
Peperomia elegans C. DC. in DC. Prodr. 16, pt. 1: 430. 1869.
A glabrous, spreading herb; stem slender; leaves alternate, lance-
elliptic, acute at both ends or gradually acuminate, 4-8 cm. long,
2.5-5 cm. wide, pinnately nerved; petiole 3-7 cm. long; spikes termi-
nal or sympodial, paired from a slender stalk 4 cm. long, 30-40 mm.
long, 2 mm. thick; peduncle 1-2 cm. long; bracts round-peltate;
berries oblong, with a slender beak; stigma anterior at the base of
the beak. Neg. 8319.
San Martin: Rio Mayo, Tarapoto, Spruce 4871, type. Huanuco:
Mountains of Huanuco, Haenke (?; herb. Prague, under the name P.
reptans Opiz).
Peperomia elegantifolia Trelease, sp. nov.
Herba mediocris glabra silvicola; folia alterna lanceolata vel
typice lanceolato-obovata subacuta basi cuneata 9-10 cm. longa
3-4.5 cm. lata e dimidio inferiore pinnatinervia, petiolo vix 1 cm.
longo; spicae in axillis binae et graciliter pedunculatae, pedunculo 3
cm. longo, vel pedunculo longiore et apice folio reducto onusto, 40-80
mm. longae, pedunculo ultimo 5-10 mm. longo; bracteae subelliptico-
peltatae.
San Martin: San Roque, 1,350-1,500 meters, Williams 7781 (herb.
Field Mus. No. 629,533, type). Loreto: Pumayacu, 600-1,200
meters, King 3180. "Sacha-chullcu."
Peperomia elliptico-rhombea C. DC. in DC. Prodr. 16,
pt. 1: 407. 1869.
A moderate-sized, suberect herb, from crisp-pubescent gla-
brescent; leaves alternate, subrhombic-elliptic or ovate, acute or
subobtuse at the base, 3-4 cm. long, 2-3 cm. wide, submultiple-
nerved from the lower half; petiole 10-15 mm. long; spikes terminal
and from the upper axils, 1 or 2 on a short, 1-bracted stalk, slender,
over 100 mm. long; peduncle 1 cm. long; bracts round-peltate;
berries round-ovoid, obliquely beaked; stigma at the base of
the beak.
42 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Type collected in Peru, without definite locality, Mathews 1835.
Huanuco: Casapi, Mathews 1689. San Martin: Tarapoto, Spruce
4095 (?).
Peperomia emarginata R. & P. Fl. 1: 33. pi. 49. 1798. Piper
emarginatum Vahl, Enum. 1: 339. 1805. Peperomia oblusifolia var.
emarginata Dahlst. Svensk. Vet. Akad. Handl. 33, pt. 2: 66.
pi 2. 1900.
A rather large, repent-assurgent, glabrous herb; stem 3-5 mm.
thick; leaves alternate, oblanceolate-spatulate, emarginate, cuneate,
12 cm. long, 5 cm. wide, pinnately nerved; petiole 2.5 cm. long;
spikes 1 or 2 on terminal or axillary stalks 4 cm. long, 120 mm. long,
10 mm. thick; peduncle 5 cm. long; bracts round-peltate; ovary
ovoid, obliquely beaked; stigma anterior.
Huanuco: Posuso, Ruiz 77, type.
Peperomia emarginulata C. DC. in DC. Prodr. 16, pt. 1: 433.
1869. P. obtusifolia Auct. in part as to Peru.
A moderately small, erect, glabrous herb; stem angular; leaves
alternate, obovate, the typically somewhat narrowed apex emarginu-
late, cuneate at the base, 3-3.5 cm. long, 1.5-2 cm. wide, pinnately
nerved below; petiole 5-10 mm. long; spikes paired on a slender
terminal stalk 2 cm. long, 50 mm. long, 2 mm. thick; peduncle 10-15
mm. long; bracts round-peltate.
Type collected in Peru by Pa von. Huanuco: Casapi, Mathews
1687.
Peperomia (?) enckeaefolia Trelease, sp. nov.
Herba plus minusve caespitosa glabra, interdum suffruticosa, 15
cm. tantum alta, caule 2 mm. crasso, internodiis brevibus; folia
alterna ovata subacuminata basi rotundata ad petiolum angustata
5-6 cm. longa 3.5-4 cm. lata 5-nervia, nervis ut venis transversis
conspicuis in statu adulto impressis, in sicco firmo-papyracea viridia
lucida, nervis serius albescentibus, margine superne tenui albescente,
petiolo 10 mm. longo; inflorescentia non visa.
Loreto: San Lorenzo, Rio Maranon Valley, 150 meters, Killip
& Smith 29201 (type in U. S. Nat. Herb.).
Peperomia enenyasensis Trelease, sp. nov.
Herba majuscula gracilis pendula epiphytica ubique minute
crispo-pubescens vel caulibus vetustioribus et pagina superiore
foliorum glabrescentibus; folia alterna subovata acute acuminata
FLORA OF PERU 43
basi acuta vel subobtusa 7-10 cm. longa 4-5 cm. lata pinnatinervia
in sicco tenuia fusca, petiolo 1.5-4 cm. longo; spicae geminatae
ramulum brevem gracili-bracteatum sympodialem terminantes,
adultae 150 mm. longae cum baccis 8 mm. diam., pedunculo 2 cm.
longo; bracteae rotundo-peltatae; baccae pallidae anguste oblongae
scutulo acuto oblique truncatae, stigmate centrali.
Junin: Enenas, Pichis Trail, 1,600-1,900 meters, Killip & Smith
25654 (type in U. S. Nat. Herb.).
Peperomia erythrocorma Trelease, sp. nov.
Herba modice parva repens tantum ciliata, caule subgracili 2-3
mm. crasso; folia alterna reniformia vel rotundo-ovata ad ovata
interdum abrupte breviacuminata, basi truncate subcordata vel
rotundata et breviter attenuata, mediocria 3-6 cm. longa 3.5-5 cm.
lata, in sicco opaca, subpalmatim 7-9-nervia transiente ciliata,
petiolo 2-2.5 cm. longo; inflorescentia non visa.
Huanuco: Posuso, 600 meters, Macbride 4678 (herb. Field Mus.).
Peperomia erythrostachya Trelease, sp. nov.
Herba majuscula glabra pendula epiphytica, caule modice
gracili 2-3 mm. crasso in sicco rubro-brunneo et sublucido; folia
alterna ovata vel elongato-ovata acuta vel subacuminata basi
rotundata vel subcordata majuscula 6-9 cm. longa 4 cm. lata, in
sicco opaca coriacea multiplinervia, ramulis costae 3-5x2 paribus
1-2 superioribus exceptis basilaribus, petiolo 2.5-6 cm. longo et
ultra; spicae axillares longae graciles 200 mm. longae 2 mm. crassae
obscure rubrae densiflorae, pedunculo 2 cm. longo medio bracteato;
bracteae rotundo-peltatae majusculae.
Huanuco: Posuso, 600 meters, Macbride 4667 (herb. Field Mus.
No. 535,741, type).
Peperomia estrellana Trelease, sp. nov.
Herba subparva glabra rhizomatosa epiphytica, ramis erectis
foliatis; folia ad nodum 3-5, foliis stolonum subspathulatis emargin-
ulatis basi cuneatis opace cinereo-viridibus obscure 3-nerviis, petiolo
2 mm. longo; spicae terminales 20 mm. longae 2 mm. crassae, pedun-
culo 15 mm. longo; bracteae rotundo-peltatae; baccae ovoideae
acutatae pseudocupula onustae, stigmate apicali.
Ayacucho: Estrella, 500 meters, Killip & Smith 23064 (type in
U. S. Nat. Herb.). Junin: Peren< Bridge, 700 meters, Killip &
Smith 25281.
44 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia exiguispica Trelease, sp. nov.
Herba gracilis subsimplex majuscula glabra arboricola, caule
2-3 mm. crasso; folia alterna lanceolato-elliptica utrinque subacuta
4-5 cm. longa 2-3 cm. lata obscure 5-nervia subtus glanduloso-
granulosa in sicco opaca, petiolo 10 mm. longo; spicae gracillimae
90 mm. longae, pedunculo 10 mm. longo; bracteae rotundo-peltatae.
Loreto: Puerto Arturo, 155-210 meters, Williams 5831 (herb.
Field Mus. No. 622,538, type).
Peperomia exilamenta Trelease, sp. nov.
Herba majuscula epiphytica vel terrestris, caule in sicco 3-4 mm.
crasso profunde angulato sparsissime molliter setuloso; folia vulgo
ternata late lanceolata vel elliptico-lanceolata obtuse acuminata
basi acuta 5-6 cm. longa 2-2.5 cm. lata 3-nervia, nervis supra im-
pressis subtus prominentibus, glabra supra impresso-punctulata
revoluta valde rigida, petiolo 5 mm. longo puberulo; spicae terminales
et ex axillis superioribus 120 mm. longae 1 mm. crassae, pedunculo
10-25 mm. longo; bracteae rotundo-peltatae; baccae ovoideae
acutatae pseudocupula onustae, stigmate apicali.
San Martin: Tarapoto, 360-900 meters, Williams -5722 (herb.
Field Mus. No. 622,513, type), 6258.
Peperomia exiliramea Trelease, sp. nov.
Herba majuscula glabra epiphytica, caule 3 mm. crasso; folia
alterna elliptica acute acuminata basi acuta 11-12 cm. longa 5-6
cm. lata e dimidio inferiore graciliter pinnatinervia, in sicco tenuia,
petiolo 1-2 cm. longo; spicae solitariae ramulos breves aphyllos
sympodiales basi anguste bibracteatos aequilongos terminantes 150
mm. longae 2 mm. crassae vel majores; bracteae rotundo-peltatae;
ovarium rotundatum acute subscutulatum.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28768 (type in
U. S. Nat. Herb.).
Peperomia falsa A. W. Hill, Ann. Bot. 21: 151. pi. 15. 1907.
A small, glabrous, acaulescent herb; leaves from a corm rooting
from the sides and base; leaves round-peltate, 1-3 cm. in diameter,
subcoriaceous; petiole 3-5 cm. long; spikes 15-30 mm. long; peduncle
elongate; bracts round-ovate; berries globose-ellipsoid, reticulate;
pseudopodium thick and cylindric; stigma apical.
Puno: Pucara, 3,700 meters, among rocks, Weberbauer 451, type.
FLORA OF PERU 45
Peperomia famelica Trelease, sp. nov.
Herba depauperata terrestris plus minusve caespitosa, caule plus
minusve crispo-pubescente; folia alterna subovato-elliptica utrinque
subacuta 2.5-5 cm. longa 1.5-2.5 cm. lata 5-nervia in vivo supra
obscure viridia, nervis albis, subtus rosea, petiolo 5 mm. longo;
spicae graciles 60 mm. longae pseudopedicellis brevibus onustae,
pedunculo 10 mm. longo; bracteae rotundo-peltatae.
Junin: East of Quimiri Bridge, near La Merced, 800-1,300
meters, Killip & Smith 23961 (type in U. S. Nat. Herb.).
Peperomia ficta Trelease, sp. nov.
Herba valida caespitoso-repens succulenta epiphytica, caule 2-4
mm. crasso glabro in sicco sulcato; folia vulgo ternata rotundo-
elliptica vel subovata vel subobovata obtuse subacuminata basi
acuta 3-4 cm. longa 2-3 cm. lata circa 5-nervia glabra in sicco dura
revoluta opaca, petiolo 5 mm. longo brunneo-puberulo ; spicae
terminales, juveniles 30 mm. longae 2 mm. crassae, pedunculo 15 mm.
longo; bracteae rotundo-peltatae.
Junin: East of Quimiri Bridge, La Merced, 800-1,300 meters,
Killip & Smith 23989 (type in U. S. Nat. Herb.).
Peperomia Fieldiana Trelease, sp. nov.
Herba modice parva repens glabra, caule crassiusculo 3-5 mm.
crasso, internodiis brevibus, ramis adscendentibus brevibus; folia
alterna elliptica vel obovato-elliptica apice rotundato emarginata
basi longicuneata majuscula 10-13 cm. longa 4.5-6.5 cm. lata in
sicco subtenuia pinnatinervia, costae ramulis circa 5x2, petiolo
15-25 mm. longo alato; spicae solitariae vel ad nodos ramuli lateralis
1-foliati vix 25 mm. longi geminatae vix 110 mm. longae et 2 mm.
crassae densiflorae, pedunculis 2.5-3 cm. longis; bracteae rotundo-
peltatae parvae in pseudoverticillo circa 12; baccae oblongae breviter
mucronatae, stigmate anteriore.
Huanuco: Posuso, 600 meters, Macbride 4684 (herb. Field Mus.
No. 535,758, type). Junin: Rio Perene", Killip & Smith 25167.
Peperomia filiformis R. & P. Fl. 1: 31. pi. 47. 1798. Piper
brachystachyum Vahl, Enum. 1: 354. 1805. Troxirum filiforme Raf.
Sylv. Tellur. 85. 1838.
A transiently puberulent or quickly glabrate, repent herb; stem
filiform; leaves mostly 4 at a node, obovate, rounded at the apex,
cuneate at the base, 7 mm. long, 5 mm. wide, 3-nerved; petiole
46 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
very short; spikes terminal, 10 mm. long, 1 mm. thick, the peduncle
of equal length; bracts ovate-peltate; ovary ovoid, the stigma apical.
Huanuco: Chinchao, Ruiz & Pawn, type. Pampayaco, Poeppig.
Junin: Carpapata, 2,700-3,200 meters, Kittip & Smith 24392
Puno : Tabina, Lechler 2089. Sangaban, Lechler 2409.
Peperomia fissicola Trelease, sp. nov.
Herba modice alta glabra e fissuris rupium cito erecta, caule 3 mm.
crasso firmo; folia alterna ad apicem caulis aggregata recurva
dolabriformia utrinque acutata 15 mm. longa 5 mm. lata, petiolo vix
5 mm. longo; spicae paniculatae prolongationem longiusculam
gracilem paucifoliatam caulis terminantes 50 mm. longae 1 mm.
crassae, pedunculis 5 mm. longis; bracteae rotundo-peltatae.
Huanuco: Huanuco, 2,100 meters, Macbride 3255 (herb. Field
Mus. No. 534,325, type).
Peperomia flabilis Trelease, sp. nov.
Herba modice parva plus minusve furcata glabra, caule 2-3 mm.
crasso; folia ad nodum 3-4 obtuse acuminata basi acuta 2-3 cm.
longa 1-1.5 cm. lata 3-nervia opaca, subtus minute granulosa palli-
diora, petiolo 3-6 mm. longo gracili; spicae terminales et ex axillis
superioribus filiformes 100 mm. longae, pedunculo gracillimo 10
mm. longo; bracteae rotundo-peltatae.
Loreto: Victoria on the Amazon River, Williams 2988 (herb.
Field Mus. No. 608,872, type).
Peperomia flavamenta Trelease, sp. nov.
Herba modice parva repenti-caespitosa succulenta fere glabra,
caule 1-2 mm. crasso in sicco sulcato; folia 2-3-nata lanceolato-
ovata acuminata basi acuta 2-5 cm. longa 1-2 cm. lata 3- vel obscure
5-nervia, petiolo 3-5 mm. longo puberulo; spicae flavae terminales
filiformes 40 mm. longae, pedunculo 10 mm. longo; bracteae rotundo-
peltatae.
Loreto: Mishuyacu, near Iquitos, 100 meters, G. Klug 987 (herb.
Field Mus. No. 630,005, type).
Peperomia flavescens C. DC. in DC. Prodr. 16, pt. 1:
436. 1869.
A spreading herb, glabrous except for the loosely villous petioles
and lower leaf surface; stem 2-3 mm. thick; leaves alternate, ovate,
subacute, peltate close to the subcordate base, 3.5 cm. long, 2-3
cm. wide, 5-7-nerved, coriaceous and yellow; petiole 2 cm. long;
FLORA OF PERU 47
spikes subterminal, 80 mm. long, 3 mm. thick or larger; peduncle
2-3 cm. long; bracts round-peltate; berries ovoid, stout-beaked;
stigma anterior. Neg. 8527.
Type collected in Peru by Pavon.
Peperomia flavescens var. Lechleri Trelease, var. nov.
Planta pubescentia plus minusve persistente induta; folia 2.5 cm.
longa, 2.5 cm. lata.
Puno: Sangari, Lechler 2691 in part (type in herb. Kew.).
Peperomia flavescens var. (?) Markhami Trelease, var. nov.
Crispo-pubescens, foliis glabrescentibus 3.5 cm. longis 2 cm. latis,
costa lata pallida.
Puno: Tambopata, "carpeting the ground in parts of the forests,"
in 1910, Clements R. Markham 4 (type in herb. Kew.; detached
leaves only).
Peperomia flavescentifolia Trelease, sp. nov.
Herba gracilis pendula epiphytica essentialiter glaber; folia
elliptica subacuminata basi subacuta 3-5 cm. longa 2-3 cm. lata
obscure pinnatinervia subtus flavescentia granulosa, in sicco valde
coriacea, petiolo vix 5 mm. longo; spicae ad apicem ramuli brevis
sympodialis geminatae 100-150 mm. longae graciles, pedunculo
15 mm. longo; bracteae rotundo-peltatae; baccae pallidae conico-
oblongae acute oblique truncatae; stigma ad scutulum insertum.
Loreto: Iquitos, 100 meters, Killip & Smith 271 42 (type in U. S.
Nat. Herb.). Mishuyacu, near Iquitos, Klug 99 (lower leaves
lanceolate, 7 cm. long, 2-3 cm. wide).
Peperomia foliata Trelease, sp. nov.
Herba modice alta gracilis dense foliata, caule subquadrangulari
puberulo; folia vulgo ad nodum 6-8 inf erne reflexa et breviora lineari-
oblanceolata obtusa vel subacuta basi subacuta 10-15 mm. longa
3 mm. lata pinnatinervia, supra ad costam subpuberula, subtus
pallidiora granulosa, petiolo 1 mm. longo; spicae terminales et ex
axillis superioribus, juveniles 25 mm. longae 1 mm. crassae densi-
florae, pedunculo vix 3 mm. longo; bracteae rotundo-peltatae.
Junin: Huacapistana, Stevens 65 (type), 178.
Peperomia foliiflora R. & P. Fl. 1: 30. pi. 45. 1798. Phyllo-
bryon Pavoni Miq. Syst. 50. 1843; 111. Piper, pi. 2. Piper foliiflorum
Vahl, Enum. 1: 356. 1805.
48 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
A rather small, stoloniferous, erect, glabrous herb; stem 2-3
mm. thick; leaves alternately clustered at the top of the stem, ovate,
acuminate, cordate, 3^4 cm. long, 2-3 cm. wide, 9-nerved with the
longer nerves forked; petiole 2-4 cm. long, winged; spikes several,
umbellate from the apex of the peduncle; bracts round-peltate; berries
ellipsoid, 6 mm. long, 1 mm. wide, flat, distinctly wrinkled, nearly
sessile, mucronate; stigma apical. Neg. 8381.
Huanuco: Chinchao, Pawn 36, type. Jauja: Rio Masamerich,
1,500 meters, Weberbauer 6665. Junin: La Merced, Killip & Smith
24656.
The specific name appears as foliflora on the original plate.
Peperomia fortipes Trelease, sp. nov.
Herba parva repenti-assurgens glabra foliosa, caule 2 mm. crasso;
folia alterna oblanceolata obtusa basi cuneata 2-3 cm. longa 1-1.5
cm. lata obscure 3-nervia in sicco opaco-olivacea, petiolo 5 mm.
longo; spicae 40-50 mm. longae 2 mm. crassae densiflorae, pedunculo
15 mm. longo 1 mm. crasso; bracteae rotund o-peltatae.
Ayacucho: Aina, 750-1,000 meters, Killip & Smith 22589 (type
in U. S. Nat. Herb.).
Peperomia Fortunati Trelease, sp. nov. P. Mandoni Auct.
Herba modice longicaulis delicata glabra pellucida, ramulis
floriferis in sicco gracillimis; folia ad apices ramorum alterne con-
gesta subrhombeo-ovata vel elliptica acute vel obtusissime acuminata
basi acuta vel obtusa 13-20 cm. longa 8-10 cm. lata delicate 5-7-
nervia, costa obscure ramosa, petiolo 2 mm. longo; spicae solitariae
ad basin laminae adnatae 5 mm. longae fere sessiles; bracteae sub-
cucullatae distichae; baccae ellipsoideae breviter stipitatae, stylo
gracili, stigmate apicali.
Cuzco : Valley of Rio Apurimac, Fortunato L. Herrera, February,
1929 (herb. Field Mus. No. 589,470, type). Without locality,
Weberbauer 5684.
Peperomia fragilissima Trelease, sp. nov.
Herba carnosa fragilissima, caule gracili; folia alterna rotundo-
ovata obtusissima basi cordata 2-4 cm. longa 2-3.5 cm. lata multi-
plinervia, nervis longioribus furcatis, petiolo 2-3 mm. longo; spicae
axillares et terminales gracillimae 40-50 mm. longae, pedunculo
20-25 mm. longo.
Loreto(?): Yurimaguas, May, 1855, Spruce (type in herb. Kew.).
FLORA OF PERU 49
Peperomia furcata Opiz in Presl, Rel. Haenk. 164. 1831.
A small, repent herb, at most with ciliate leaves; leaves alternate,
round-ovate, subacute at both ends or round-based, 5-7-nerved with
the inner nerves confluent at the base; petiole 5-10 mm. long;
spikes terminal or paired on a sympodial common stalk, 40 mm. long,
2 mm. thick; peduncle 1-2 cm. long.
Type from mountains of Peru, Haenke.
Peperomia fuscispica C. DC. Bot. Jahrb. 40: 260. 1908.
A small, glabrous epiphyte; stem filiform; leaves alternate, round
or subobovate, acute-based, 9 mm. in diameter, 3- or less evidently
5-nerved; petiole 3 mm. long; spikes terminal, 20 mm. long, 2mm.
thick; peduncle 3 mm. long; bracts round- or elliptic-peltate; ovary
subovoid; stigma oblique.
Huanuco: Monzon to Rio Huallaga, Weberbauer 3627, type.
Peperomia galoides HBK. var. aromatica C. DC. Bot Jahrb.
40:266. 1908.
Aromatic; leaves oblong, somewhat black-punctulate.
Ancash: Caraz, Weberbauer 3227, type. "Tunacongona."
The typical form of the species has a wide distribution in tropical
America.
Peperomia Gayi C. DC. Ann. Cons. Jard. Geneve 2: 283. 1898.
A moderately small, simple, very dingy-hirsute herb; stem slender;
leaves opposite, round in cross section below, elliptic upward, short-
acuminate, acute-based, 2-4 cm. long, 1.5-2.5 cm. wide, 5-nerved,
black-punctulate beneath; petiole 1 cm. long; spikes terminal and
from the upper axils, 50 mm. long, 1 mm. thick; peduncle 1 cm. long;
bracts round-peltate; ovary obovoid; stigma oblique.
Based on material collected in Peru, Gay 2181, 2310, 2681.
Peperomia gibba C. DC. Verh. Bot. Ver. Brandenb. 47:
116. 1905.
A moderate-sized, spreading, glabrous herb; leaves alternate,
round-ovate, slightly mucronate, shallowly and openly cordate,
5-6 cm. long, 3.5 cm. wide, 7-nerved; petiole 3-4 cm. long; spikes
axillary and terminal, slender, 75 mm. long; peduncle 3-4 cm. long;
bracts elliptic-peltate; berries oblong, obliquely swollen at the base,
subtruncate; stigma subapical.
Loreto: Pongo de Cainarachi, Ule 6322, type.
50 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia Gilberti Trelease, sp. nov.
Herba minor stolonifero-erecta divaricato-ramosa, caule gracili
1-2 mm. crasso puberulo serius furfuraceo-exfoliante; folia ad nodum
circa 5 suboblonga, inferiora spathulato-revoluta, obtusa basi sub-
acuta parva 8-10 mm. longa 3-4 mm. lata in sicco fusca opaca non
manifeste nervata, supra puberula, petiolo vix 2 mm. longo puberulo;
spicae ramos terminantes 25 mm. longae 1 mm. crassae, floribus
laxiuscule dispositis, pedunculo vix 5 mm. longo subpuberulo;
bracteae rotundo-peltatae; baccae globosae submucronulatae, stig-
mate obliquo.
Cuzco: Ollantaitambo, 3,000 meters, Cook & Gilbert 740 (type in
U. S. Nat. Herb.).
Peperomia glabrirhachis Trelease, sp. nov.
Herba modice parva stolonifero-caespitosa, caule 1-2 mm. crasso
anguloso sparse retrorso-pubescente; folia ad nodum circa 4 sub-
rhombeo-elliptica vel obovata obtusa basi acuta 10 mm. longa 5-6
mm. lata 1-nervia revoluta, subtus sparse pubescentia, utrinque
papilloso-rugosa, petiolo vix 1 mm. longo; spicae terminales 15
mm. longae 1 mm. crassae glabrae, pedunculo filiformi velutino
spicam aequante; bracteae rotundo-peltatae.
Junin: Hacienda Chalhuapuquio, Stevens 211 (type). Huaca-
pistana, Stevens 64, 179. Enenas, Killip & Smith 25653.
Peperomia glareosa Trelease, sp. nov.
Herba majuscula caespitosa assurgens, caule 1 mm. vel prope
basin ad 4-5 mm. crasso in sicco sulcato puberulo granuloso; folia
ad nodum 2-4 late lanceolata obtusa basi acuta in sicco firma opaca,
subtus granulosa, 3-nervia, superiora 1.5 cm. longa 1 cm. lata, folia
ramulorum sterilium 3.5 cm. longa 1.5 cm. lata, petiolo 2-3 mm.
longo subpuberulo-granuloso; spicae terminales et axillares 55 mm.
longae 2 mm. crassae, pedunculo 15 mm. longo granuloso; bracteae
rotundo-peltatae; baccae ovoideae ad stylum attenuatae pseudo-
cupula onustae, stigmate apicali.
Loreto: Rio Santiago, 200 meters, Ynes Mexia 6236 (type in
herb. Univ. Calif.).
Peperomia granulatifolia Trelease, sp. nov.
Herba epiphytica majuscula pendula glabra, caulibus gracil-
limis; folia alterna elliptica acuminata basi acuta 4-7 cm. longa
2-3.5 cm. lata in sicco coriacea obscure pinnatinervia, petiolo 5
FLORA OF PERU 51
mm. longo; spicae terminales et sympodiales solitariae vel ad apicem
ramuli brevis 2-bracteati geminatae 100 mm. longae 1 mm. crassae
vel ultra, pedunculo 5-10 mm. longo.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26390 (type
in U. S. Nat. Herb.).
Peperomia granulatilimba Trelease, sp. nov.
Herba epiphytica fere glabra pendula, caulibus gracilibus 2 mm.
crassis; folia alterna subovato-elliptica acute acuminata basi rotun-
data 6 cm. longa 3 cm. lata, vel elliptico-oblonga basi subacuta
atque 10 cm. longa et 4 cm. lata, pinnatim paucinervia pallido-
granulosa sparse pilosa longiciliata, petiolo 5-10 mm. longo; spicae
3 caulem brevem sympodialem terminantes 100 mm. longae 2 mm.
crassae, pedunculo vix 5 mm. longo; bracteae rotundo-peltatae;
baccae oblongae vel conice productae, oblique acuto-scutulatae,
stigmate centrali.
Loreto: Yurimaguas, Lower Rio Huallaga, 155-210 meters,
Llewelyn Williams 4617 (herb. Field Mus. No. 629,584, type).
La Victoria, Williams 2945, 2802, 3084. Pichis Trail, Killip &
Smith 26215. Pumayacu, 600-1,200 meters, on tree, Klug 8200.
Peperomia Grisari C. DC. Bull. Herb. Boiss. 6: 516. 1898.
A slender, repent-assurgent herb, the petiole and the nerves
beneath coarsely puberulent; leaves alternate, round-ovate, acute,
shallow-cordate with the margin crossing the petiole, 7 cm. long,
6 cm. wide, 9-nerved, paler beneath; petiole 4 cm. long; spikes
terminal, paired on a stalk 2 cm. long, 50 mm. long, 2 mm. thick;
peduncle 15-20 mm. long.
Type collected in Ecuador or Peru, Grisar.
Peperomia guayrapurana Trelease, sp. nov.
Herba gracilis, caule, petiolis et foliis subtus rigido-hirtellis;
folia ad nodum 3-4 elliptica utrinque acuta 1.5-2 cm. longa 1 cm.
lata 3-nervia, petiolo vix 5 mm. longo; spicae terminales 20 mm.
longae 1 mm. crassae, pedunculo 1 cm. longo.
San Martin: Monte Guayrapurana, near Tarapoto, Spruce 4010
(type in herb. Kew.).
Peperomia gucayana Trelease, sp. nov.
Herba erecta vix ramosa firma, caule 2-3 mm. crasso minute
fusco-papilloso; folia plerumque opposita late obovata utrinque
acuta vix 2 cm. longa et 1.5 cm. lata 3-nervia, costa plus minusve
52 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
ramosa, in sicco tenuia et saepe conduplicata, petiolo 2-4 mm.
longo minute papillose; spicae axillares 60-80 mm. longae 1 mm.
crassae, floribus subdense dispositis, pedunculo 10 mm. longo;
bracteae rotundo-peltatae ; baccae ovoideae crasse submucronatae,
stigmate obliquo.
Cuzco: Near Gucay, Urubamba Valley, 3,000 meters, Fortunato
L. Herrera 1464 (type in U. S. Nat. Herb.).
Peperomia haematolepis Trelease, sp. nov.
Herba majuscula (repenti-assurgens?) succulenta glabra, caule
3-5 mm. crasso furfuraceo-exfoliante; folia alterna orbicularia vel
rotundo-subobovata emarginulata basi rotundata vel subacuta 4.5
cm. longa 3-4 cm. lata pinnatim paucinervia, coriacea opaca, petiolo
5-10 mm. longo; spicae terminales, juveniles 15-20 mm. longae 3
mm. crassae, pedunculo 2-2.5 cm. longo; bracteae rotundo-peltatae
rubro-aurantiacae.
Junin: Hacienda Chalhuapuquio, Stevens 212 (type).
Peperomia Haenkeana Opiz in Presl, Rel. Haenk. 162. 1831.
A rather small, erect herb, the stem, petioles, and nerves beneath
short- villous or crisp-pubescent; leaves alternately clustered at the
top of the stem, round-ovate, acuminate, cordate with overlapping
lobes, 4-6 cm. long, 3-5 cm. wide, 7-9-nerved with some nerves
forked; petiole 2-3 cm. long; spikes several, umbellate at the end
of the peduncle, nearly sessile, barely 10 mm. long, flattened and
flowered from the edge; bracts round-peltate.
Type from mountains of Peru, Haenke.
Peperomia hamiltonianifolia Trelease, sp. nov.
Herba parva stolonifero-erecta glabra, caule 2 mm. crasso; folia
alterna pauca elliptica utrinque subacuta 2-2.5 cm. longa 1 cm.
lata in sicco crassa subtus 5-nervia, petiolo 3-5 mm. longo; spicae
terminales et axillares 25 mm. longae 1 mm. crassae, pedunculo 5
mm. longo; bracteae rotundo-peltatae comparate magnae.
Type collected in Peru by Gay (herb. Paris).
Peperomia Herrerae Trelease, sp. nov.
Herba assurgens plus minusve ramosa modice parva glabra,
caule 2-3 mm. crasso; folia opposita elliptica utrinque acuta 2-2.5
cm. longa 1-1.5 cm. lata delicate 3-nervia, costa plus minusve
ramosa, in sicco tenuia viridia, petiolo 2 mm. longo; spicae termi-
nales et axillares subabundantes 40-50 mm. longae 2 mm. crassae,
FLORA OF PERU 53
floribus sublaxe dispositis, pedunculo gracili 15 mm. longo; bracteae
rotundo-peltatae; baccae rotundato-ovoideae mucronatae; stigma
subapicale plus minusve lobatum.
Cuzco: Hacienda Churu, Paucartambo, 3,500 meters, Fortunate
L. Herrera, February, 1924 (type in U. S. Nat. Herb.).
Peperomia heterophylla Miq. Lond. Journ. Bot. 4: 415. 1845,
A small, repent-assurgent epiphyte, glabrous except that the
filiform stems are at first puberulent; leaves alternate, round-
obovate below and 5 mm. in diameter, becoming obovate in the
middle and lanceolate and 2 cm. long upward, with the narrowed
apex subemarginulate, acute-based, 1-nerved, punctulate beneath;
petiole very short; spikes curved, 50 mm. long, 1 mm. thick; bracts
round-peltate.
Huanuco: Casapi, Mathews 1686, type.
Peperomia heterostachya A. Dietr. Syn. PL 1: 717. 1831.
Peperomia tristachya Opiz in Presl, Rel. Haenk. 103. pi. 30. 1831,
non P. tristachya HBK. 1815. Piper heterostachyon D. Dietr. Syn.
PI. 1: 111. 1839.
A glabrous, succulent, branching herb: leaves opposite below,
3 or 4 at a node above, lance-ovate, long-acuminate, acute-based,
7 cm. long, 3 cm. wide, 3-nerved; petiole 3 mm. long; spikes terminal
and from the uppermost axils, elongate.
Huanuco: Huanuco mountains, Haenke, type.
Peperomia Hillii Trelease, nom. nov. Peperomia Gaudichaudii
A. W. Hill, Ann. Bot. 21: 154. 1907, non P. Gaudichaudii Miq. 1843.
A small, glabrous, acaulescent herb; leaves from a small, rooted
corm, round or subdeltoid, peltate rather below the middle, 5-10
mm. long, 6-8 mm. wide; petiole 2-8 cm. long; spikes 60-80 mm.
long; peduncle equaling the petiole; bracts elliptic-peltate; berries
round-ovoid, slightly reticulate. Neg. 8345.
Lima: Lima and San Lorenzo, Gaudichaud 150. Atocongo,
clefts of limestone rocks, 250-500 meters, Pennell 14750. Chile.
Peperomia huacachiana Trelease, sp. nov.
Herba majuscula assurgens glabra, caule crassiusculo 3-6 mm.
crasso, internodiis brevibus; folia alterna elliptica vel lanceolato-
elliptica acuta vel acuminata basin versus sensim acuta, in sicco
subtenuia opaca, subtus pallidiora, pinnatinervia ramis costae circa
8-10x2, petiolo 1-2 cm. longo alato; spicae solitariae terminales
54 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
vel oppositifoliae ramulum crassum 1-bracteatum vel 1-foliatum
1.5-3 cm. longum terminantes, magnae 200 mm. longae 5 mm.
crassae floribus sublaxe insertis; bracteae rotundo-peltatae modice
magnae diametro suo remotae, floribus in alveolis lenticularibus
immersis; baccae ellipsoideo-oblongae fuscae suboblique truncatae,
stigmate in disco anteriore.
Huanuco: Huacachi near Muna, 1,950 meters, Macbride 3899
(herb. Field Mus. No. 534,962, type). Cani near Mito, 2,550
meters, 341 6.
Peperomia huacapistanana Trelease, sp. nov.
Herba mediocris glabra repens epiphytica, ramis fructiferis '
brevibus erectis, foliis reductis, caule 2-3 mm. crasso; folia alterna '
orbicularia vel ovata vel subreniformia apice rotundata basi sub-
truncata 3.5-4 cm. longa 2.5-3.5 cm. lata 7-nervia, nervis interiori- I
bus interdum confluentibus, in sicco coriacea supra obscure viridia
subtus pallidiora, petiolo 1.5-2 cm. longo; spicae solitariae vel
geminatae vel 4 et ultra et paniculatae, foliis ramulorum fertilium
bracteiformibus, 25 mm. longae 2 mm. crassae densiflorae, pedunculo
filiformi 10 mm. longo; bracteae rotundo-peltatae.
Junin: Huacapistana, Stevens 89 (type). Junin: Enefias, Killip
& Smith 25655, 25736. San Nicolas, Killip & Smith 26064. Hua-
nuco: Posuso, 4678.
Peperomia huallagana Trelease, sp. nov.
Herba epiphytica modice parva erecta plus minusve dichotoma
glabra, caule 2-3 mm. crasso; folia ad nodum vulgo 3-4 obovata
ad obovato-oblonga in eodem verticillo, emarginulata basi cuneata
12-20 mm. longa 6-10 mm. lata obscure 3-nervia, costa indistincte
ramosa, petiolo 2-5 mm. longo alato; spicae terminales parvae 30
mm. longae 1 mm. crassae densiflorae, pedunculo 1 cm. longo;
bracteae rotundo-peltatae, floribus inter costas anastomosantes
rhachis immersis.
Huanuco: Rio Huallaga Canyon below Rio Santo Domingo,
1,200 meters, Macbride 4264 (herb. Field Mus. No. 535,345, type).
Peperomia huallagana var. compacta Trelease, var. nov.
Compacte ramosa brevis; spicae 40 mm. longae 2 mm. crassae,
pedunculo duplo breviore.
Junin: Chanchamayo Valley, Carlos Schunke 1774 (herb. Field
Mus. No. 625,560, type).
FLORA OF PERU 55
Peperomia huantana Trelease, sp. nov.
Herba subparva repens corticicola, caule 1 mm. crasso sparse
crispo-subvilloso; folia ad nodum 2-4 rhombeo-obovata ad sub-
elliptica obtusa basi acuta 10-15 mm. longa 7-10 mm. lata subtus
sparse subvillosa et pallidiora, in sicco tenuia opaca, petiolo 2-3
mm. longo; spicae terminales (?) filiformes 60 mm. longae, pedunculo
20 mm. longo; bracteae rotundo-peltatae; baccae ovoideae acutatae
pseudocupula onustae, stigmate apicali.
Ayacucho: Aina, 750-1,000 meters, Killip & Smith 22728 (type
in U. S. Nat.Herb.), 22780. Junin: Yapas, Killip & Smith 25510(1}.
Peperomia huantana var. enenyasana Trelease, var. nov.
Pubescentia longior; folia magis lanceolata 20 mm. longa 10
mm. lata.
Junin: Enenas, Pichis Trail, 1,600-1,900 meters, Killip & Smith
25727 (type in U. S. Nat. Herb.).
Peperomia huanucoana Trelease, sp. nov.
Herba modice parva assurgens ramosa, caule gracili 1-2 mm.
crasso dense papilloso-puberulo; folia ad nodum vulgo 3-4 patentia
vel reflexa ovato-elliptica vel oblongo-elliptica utrinque obtusa sub-
sessilia parva 10-20 mm. longa 5-6 mm. lata utrinque papilloso-
puberula subtenuia anguste 3-nervia, costa plus minusve" ramosa;
spicae terminales et axillares 40 mm. longae 1 mm. crassae laxi-
florae, pedunculo vix 10 mm. longo papilloso-puberulo; bracteae
rotundo-peltatae; ovarium impressum ovoideum, stigmate sub-
apicali.
Huanuco: Huanuco, 2,100 meters, on ledges of rock cliffs, Mac-
bride 3256 (herb. Field Mus. No. 534,326, type).
On stolons the foliage is very different: leaves overlapping,
round-elliptic, thick, revolute, and scarcely more than 5 mm. long
and 4 mm. wide.
Peperomia hyporhoda Trelease, sp. nov.
Herba simplex mediocris rhizomatoso-assurgens, caule 2-3 mm.
crasso plus minusve crispo-pubescente; folia alterna subelliptica
vel inferiora minora et obovata obtusa vel subacuminata basi
acuta 2.5-4 cm. longa 2-3 cm. lata 3- vel minus distincte 5-nervia,
in sicco granuloso-punctulata, in vivo subtus rubra sed in sicco
brunnescentia, petiolo 1 cm. longo crispo-pubescente; spicae 2
apicales 100 mm. longae 3 mm. crassae pseudopedicellis onustae,
pedunculo 10-15 mm. longo; bracteae rotundo-peltatae.
56 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Junin: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25479 (type in U. S. Nat. Herb.).
Peperomia inaequalifolia R. & P. Fl. 1: 30. pi. 46. 1798.
Piper inaequalifolium Vahl, Enum. 1 : 355. 1805. Piper aromaticum
Willd. Enum. PI. Suppl. 3. 1813.
A rather large, aromatic, subcespitose herb, or suffrutescent;
leaves 4 at a node, oblong-spatulate, obtuse, cuneate at the base,
5 cm. long, 1.5 cm. wide, reduced below to scarcely 3.5 cm. long
and 1 cm. wide, 3-nerved; petiole 5 mm. long; spikes terminal and
axillary, elongate.
Type material from Lima and Huanuco, Pavon. Without
locality, Mathews 124, 933.
Peperomia inaequilatera Trelease, sp. nov.
Herba modice parva (repens?) assurgens dense puberula, caule
subgracili 2-3 mm. crasso, internodiis brevibus; folia ad nodum
vulgo ternata elliptico-oblonga utrinque subacuta vel obtusa parva
2 cm. longa 1 cm. lata, foliis ramulorum sterilium majoribus 4.5
cm. longis 2 cm. latis vel multo majoribus 7 cm. longis et 2.5 cm.
latis atque rhombeo-oblanceolatis, uno latere ad petiolum 3 mm.
profundius quam altero decurrente, in sicco tenuia pellucida, 5-
nervia costa plus minusve ramosa, petiolo brevi vix 5 mm. longo;
spicae terminales 60 mm. longae 1 mm. crassae, floribus laxe sub-
annulatis, pedunculo 1 cm. longo ut rhachi puberulo; bracteae
rotundo-peltatae; ovarium ovoideum, stigmate subapicali.
Huanuco: Piedra Grande near Rio Santo Domingo, 1,500 meters,
Macbride 3858 (herb. Field Mus. No. 534,921, type).
Peperomia induratifolia Trelease, sp. nov.
Herba parva essentialiter glabra repens vel stolonifera in sicco
nigrescens, caule 1-2 mm. crasso, internodiis brevibus; folia alterna
vel suprema ternata lanceolata acuminata basi acuta 20-30 mm.
longa 5-12 mm. lata opaca vix plus quam 1-nervia, petiolo 5-10
mm. longo; spicae ut videtur terminales filiformes 80-90 mm.
longae, pedunculo 10 mm. longo; bracteae rotundo-peltatae.
Junin: Chanchamayo Valley, 1,500 meters, Carlos Schunke 1582
(herb. Field Mus. No. 627,436, type).
Peperomia itayana Trelease, sp. nov.
Herba subparva brevicaulis glabra; folia pauca rotundo-ovata
obtusissima basi brevi ter cordata 7-9 cm. longa 6-8 cm. lata 7-
FLORA OF PERU 57
nervia, costa nervisque lateralibus 4-ramosis, in sicco tenuia viridia,
petiolo 8-10 mm. longo (et ultra?); inflorescentia non visa.
Loreto: Soledad, Rio Itaya, 110 meters, Killip & Smith 29753
(type in U. S. Nat. Herb.), 29618. San Antonio, Killip & Smith
29364, younger and smaller. Santa Rosa, Killip & Smith 28724-
Peperomia juniniana Trelease, sp. nov.
Herba mediocris ramosa, caule 1-3 mm. crasso transiente crispo-
villoso; folia alterna ovata acuminata basi breviter cordata basi
intra sinum breviter cuneata, 4-10 cm. longa 3-5 cm. lata multi-
plinervia nervis lateralibus furcatis, costa ramosa, in sicco tenuia,
subtus pallida minute nigro-punctata, nervis hirtellis, petiolo 1-4
cm. longo crispo-hirtello; spicae axillares, juveniles 65 mm. longae
2 mm. crassae densiflorae, pedunculo 1 cm. longo hirtello; bracteae
rotundo-peltatae.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 357
(herb. Field Mus. No. 571,411, type); Killip & Smith 24809.
Peperomia Klotzschiana Miq. Syst. 145. 1843.
A moderate-sized, hirsute herb; stem 2-3 mm. thick, branching
upward; leaves 4-6 at a node, oblanceolate-subspatulate, subacute
at both ends, 20-25 mm. long, 6-10 mm. wide, 3-nerved; petiole
5 mm. long; spikes numerous in the axils of the congested and
reduced upper leaves, 10 mm. long, 1 mm. thick; peduncle very
short; bracts round-peltate, pale-margined.
Loreto (?) : Mainas, Poeppig, type.
Peperomia lanosa Trelease, sp. nov.
Herba modice alta vix valida stolonifero-erecta, caule 1-3 mm.
crasso, internodiis longiusculis dense longipilosis; folia inferiora
opposita obovato-cuneata 2 cm. longa 1.5 cm. lata longe (1 cm.)
petiolata, superiora vulgo ad nodum 3-5 spathulata vel suboblonga
et brevipetiolata, 3 cm. longa et 1 cm. lata vel 2 cm. tantum longa
et 6 mm. lata, 3-nervia, utrinque vel supra tantum ad nervos ad-
presso-pilosa, petiolo dense piloso; spicae terminales et axillares
40-70 mm. longae 1 mm. crassae laxiflorae, pedunculo 1.5-2 cm.
longo piloso; bracteae rotundo-peltatae; baccae ovoideae mucronatae,
stigmate oblique apicali.
Huanuco: Cani near Mito, 2,550 meters, Macbride 3450 (herb.
Field Mus. No. 534,525, type).
58 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia Lechleriana Trelease, sp. nov.
Herba majuscula glabra; folia alterna subelliptico-lanceolata
utrinque acuta 14-19 cm. longa 5.5-6.5 cm. lata pinnatinervia,
petiolo 2 cm. longo; spicae anguste paniculatae caudiculum scapi-
formem terminantes 10 mm. longae 1 mm. crassae, pedunculo
brevissimo; bracteae rotundo-peltatae; baccae ellipsoideae oblique
truncatae, stigmate centrali.
Puno: Sangaban, Lechler 2375 (herb. Kew.).
Peperomia lepadiphylla Trelease, sp. nov.
Herba alpina intertexta fere glabra, caule mediocri 2-3 mm.
crasso usque ad apicem foliato; folia alterna crassissima compressa
V-sulcata oblique ovata acuta basi rotundata 10 mm. longa 6 mm.
lata enervia ad costam plus minusve puberula, petiolo filiformi
vix 5 mm. longo; spicae terminales 20 mm. longae 3 mm. crassae
densiflorae, pedunculo nullo; bracteae rotundo-peltatae; ovarium
ovoideum, stigmate apicali.
Lima: Rio Blanco, 4,500 meters, Macbride 3055 (herb. Field
Mus. No. 534,125, type).
Peperomia leuconeura Trelease, sp. nov.
Herba modice parva assurgens, caule subgracili 2 mm. crasso
crispo-pubescente; folia alterna rhombeo-elliptica utrinque sub-
acuminata modice parva 4 cm. longa 2.5 cm. lata 3- vel obscure
5-nervia, supra ad nervos crispo-pubescentia, subtus nigro-punctu-
lata et inter nervos albidos rubescentia, petiolo gracili 15 mm.
longo glabrato; spicae axillares et terminales, juveniles 25 mm.
longae 1 mm. crassae densiflorae, pedunculo brevissimo glabrato;
bracteae rotundo-peltatae pallido-marginatae.
Junin: Hacienda Schunke, La Merced, 1,200 meters, Macbride
5628 (herb. Field Mus. No. 536,667, type).
Peperomia limaensis Trelease, sp. nov.
Herba parva compacte ramosa in sicco fusca, caule 1-2 mm.
crasso, internodiis brevibus dense puberulis; folia ad nodum vulgo
4 obovata utrinque rotundata parva 5 mm. longa 3 mm. lata, supra
versus apicem leviter puberula, in sicco opaca non manifeste nervata,
petiolo vix 1 mm. longo glabrato; spicae terminales 50 mm. longae 1
mm. crassae densiflorae, pedunculo brevissimo vix 2 mm. longo;
baccae impressae ellipsoideae obtusae, stigmate subapicali.
Lima: San Geronimo, Lima, 150 meters, Macbride 5920 (herb.
Field Mus., type).
FLORA OF PERU 59
Peperomia longepedunculata Opiz in Presl, Rel. Haenk.
164. 1831.
A rather large but slender-stemmed, glabrous herb; leaves alter-
nate, ovate, acute or subacuminate, openly cordate, 9-13 cm. long,
7-9 cm. wide, multiple-nerved with about 6 nearly basal branches of
the midrib and 2 or 3 upward, the longer nerves forking; petiole
10-15 cm. long; spikes sympodial, 40 mm. long, 2-3 mm. thick;
peduncle slender, 3 cm. long.
Huanuco: Huanuco mountains, Haenke in 1790, type.
Peperomia longipila C. DC. Verh. Bot. Ver. Brandenb. 47:
115. 1905.
A small, suberect, long-pilose or villous herb; stem slender;
leaves alternate, round-ovate or elliptic, 11 mm. long, 8 mm. wide,
3-nerved ; petiole 5-10 mm. long; spikes extremely slender and loosely
flowered, scarcely 10 mm. long; peduncle 10 mm. long, glabrous;
bracts round-peltate; berries ellipsoid, scarcely stipitate, with a style;
stigma apical.
Loreto: On cliffs, Pongo de Cainarachi, Ule 6407, type.
Peperomia longispica Trelease, sp. nov.
Herba modice parva assurgens ramosa, caule gracili 1-2 mm.
crasso vel ad basin 4-5 mm. crasso evanescente papillose, furfuraceo-
exfoliante: folia vulgo ad nodum 4-6 primo patentia obovato-oblonga
vel oblonga apice rotundata basi subacuta subsessilia parva 9-15
mm. longa 4 mm. lata supra prope apicem puberula, subtenuia,
obscure pinnatinervia; spicae terminales et axillares elongatae 60-
150 mm. longae 1 mm. crassae laxiflorae, pedunculo vix 1 cm. longo
glabro; bracteae rotundo-peltatae: baccae globosae oblique mucron-
ulatae, e pseudopedicellis brevibus, stigmate subapicali.
Huanuco: Muna, 2,100 meters, Macbride 3928 (herb. Field Mus.
No. 534,990, type). Junin: Huacapistana, Killip & Smith 24140,
24188, 24147. Carpapata, Killip & Smith 24342. Carrapa, Killip
& Smith 22380, 22426.
Peperomia Macbrideana Trelease, sp. nov.
Herba majuscula assurgens, caule crassiusculo 3-6 mm. crasso,
internodiis brevibus glabratis; folia alterna vel interdum subopposita
rotundo-elliptica vel orbicularia utrinque rotundata mediocria 4-6
cm. longa 3.5-5 cm. lata palmatim 5-nervia, nervis ramosis, subtus
ad nervos ferrugineo-villosa, petiolo subgracili 3-8 cm. longo,
60 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
foliorum superiorum breviore; spicae ad apicem ramuli pilosi
spicatim congestae, juveniles vix 5 mm. longae; bracteae rotundo-
peltatae.
Huanuco: Mima, Huacachi, 1,950 meters, Macbride 4144 (herb.
Field Mus. No. 535,218, type).
Peperomia macrorhiza HBK. Nov. Gen. & Sp. 1: 72. 1815.
A small, glabrous, acaulescent herb; leaves from a (rather large?)
corm, round, peltate, 6 mm. in diameter; petiole 5 cm. long; spikes as
yet small; bracts round-peltate.
Cajamarca: Cajamarca, Humboldt, type.
Peperomia macrothyrsa Miq. Lond. Journ. Bot. 4: 419. 1845.
A glabrous, succulent, mountain herb, stoloniferous-assurgent,
sometimes dwarfed; stem rather thick, naked below; leaves clustered
alternately at the top of the stem, subelliptic, rather obtuse, acute-
based, 20 mm. long, 10 mm. wide; petiole 1 cm. long; spikes on a
prolonged, somewhat branched shoot, nearly sessile in the axils of
gradually reduced leaves, 40 mm. long, 1 mm. thick; bracts
round-peltate; berries round-ovoid, mucronate; stigma apical.
Neg. 8385.
Amazonas: Chachapoyas, Mathews 3228, type.
Peperomia majalis Trelease, sp. nov.
Herba mediocris arboricola foliis ciliatis exceptis glabra, caule 4
mm. crasso, internodiis brevibus; folia alterna elliptica vel typice
ovata subacuta basi rotundata vel breviter cordata prope basin
peltata, 6-10 cm. longa 3.5-6.5 cm. lata graciliter multiplinervia, in
sicco flavescenti-brunnea et coriacea, petiolo 2-4 cm. longo; spicae
terminales ad apicem ramuli 1-bracteati geminatae 40 mm. longae
2 mm. crassae, pedunculo 1 cm. longo; bracteae rotundo-peltatae.
Junin: Dos de Mayo, Pichis Trail, 1,700-1,900 meters, Killip &
Smith 25886 (type in U. S. Nat. Herb.).
Peperomia mansericheana Trelease, sp. nov.
Herba parva repenti-assurgens arboricola, caule filiformi puber-
ulo; folia alterna lanceolata obtusa basi acuta 18-28 mm. longa 6-7
mm. lata glabra, in sicco tenuia 1-3-nervia, petiolo 2 mm. longo;
spicae terminales filiformes 40 mm. longae, pedunculo 10 mm. longo;
bracteae rotundo-peltatae.
Loreto: Mouth of Rio Santiago, Pongo de Manseriche, 210
meters, Ynes Mexia 6196 (type in herb. Univ. Calif.).
FLORA OF PERU 61
Peperomia Maransara Trelease, sp. nov.
Herba procumbens, caulibus gracilibus 1-2 mm. crassis obscure
adpresso-puberulis; folia acuminata basi rotundata vel subtruncato-
cordulata modice parva 2.5-5 cm. longa 2-3.5 cm. lata in sicco
viridia firma ciliata 5-nervia supra ad nervos primo adpresso-pu-
bescentia, petiolo 2-5 cm. longo glabro; spicae ramulos puberulos
3-4 cm. longos prope medium 1-bracteatos terminantes 30 mm.
longae 2 mm. crassae densiflorae; bracteae rotundo-peltatae; baccae
fuscae oblongae, rostro fere aequilongo setaceo; stigma ad basin
rostri anterius.
Cuzco: San Miguel, Urubamba Valley, 1,800 meters, Cook &
Gilbert 960 (type in U. S. Nat. Herb.).
Peperomia marcapatana Trelease, sp. nov.
Herba mediocris repenti-assurgens plus minusve ramosa glabra,
caule 3-4 mm. crasso basi plus minusve lignescente; folia ad nodum
4-6 rotundo-elliptica emarginulata basi obtusa 10-15 mm. longa
6-10 mm. lata 1-nervia opaca subrevoluta, subtus granulosa, petiolo
3-5 mm. longo; spicae terminales 120 mm. longae 3 mm. crassae
laxiflorae, pedunculo 15-20 mm. longo; bracteae rotundo-peltatae,
floribus in alveolis insertis.
Cuzco: Marcapata, Prov. Quispicanchi, 3,000 meters, Weberbauer
7801 (herb. Field Mus. No. 605,155, type).
Peperomia Marshalliana Trelease, sp. nov.
Herba delicata glabra repens, caule filiformi sed firmo; folia
alterna obovata apice rotundata vel subtruncata basi acuta parva
10-20 mm. longa 7-12 mm. lata in sicco subcoriacea 5-nervia, petiolo
brevi vix 2 mm. longo; spicae terminales ramulos breves compacti-
foliatos terminantes parvae 10 mm. longae 1 mm. crassae densiflorae,
pedunculo spicae aequilongo; bracteae rotundo-peltatae; baccae sub-
globosae, stigmate obliquo.
Huanuco: Posuso, 600 meters, associated with P. scandens,
Macbride 4710 (herb. Field Mus. No. 551,172, type). Junin: Chan-
chamayo Valley, Carlos Schunke 460. San Martin: San Roque,
1,350-1,500 meters, Williams 7382.
Peperomia Mathewsiana Miq. Lond. Journ. Bot. 4: 423. 1845.
Erect, suffrutescent, pubescent; leaves 3-7 at a node, rhombic-
spatulate, obtusely narrowed at the apex, cuneate at the base, 3 cm.
62 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
long, pellucid, 3-nerved; petiole 3-5 mm. long; spikes terminal and
axillary, elongate; peduncle 5-7 mm. long; bracts round-peltate.
Huanuco: "Zacapota," Casapi, Mathews 1688, type.
Peperomia megalepis Trelease, sp. nov.
Herba modice parva repenti-assurgens glabra arboricola, caule
subgracili 2-3 mm. crasso, internodiis brevibus; folia alterna rhom-
beo-elliptica vel obovata subacuta parva 1.5-3.5 cm. longa 1-2 cm.
lata, in sicco opaca 5-nervia, petiolo 5-10 mm. longo; spicae ter-
minales et axillares 70 mm. longae 2 mm. crassae sublaxiflorae,
pedunculo fere 2 cm. longo; bracteae magnae 1 mm. latae rotundo-
peltatae.
Huanuco: Trail to Tambo de Vaca, Muna, 2,400 meters, Macbride
4275 (herb. Field Mus. No. 535,357, type).
Peperomia mercedana C. DC. Bot. Jahrb. 40: 299. 1908.
A rather large, glabrous herb; leaves alternate, subrhombic-
obovate, blunt-acuminate, cuneate at the base, 8-9 cm. long, 3.5-4
cm. wide, pinnately nerved; petiole barely 1 cm. long; spikes solitary
in the axils on a slender stalk, or 2 or 4 on a similar terminal stalk
3-4 cm. long; bracts round-peltate; ovary obovoid, obliquely beaked;
stigma anterior.
Junin: La Merced, Weberbauer 1882, type.
Peperomia mercedana var. carpapatana Trelease, var. nov.
Folia subrhombeo-obovata acutiuscule acuminata basi longi-
cuneata 11 cm. longa 5 cm. lata, petiolo 1-2 cm. longo.
Junin : Carpapata above Huacapistana, 2,700-3,200 meters, Killip
& Smith 24485 (type in U. S. Nat. Herb.).
Peperomia metallica Lind. & Rod. 111. Hort. 39: 79.
pi. 557. 1892.
A much-branched, succulent, glabrous herb with short, pellucid,
pinkish internodes; leaves lanceolate or lance-ovate, the narrowed
apex obtuse, rather obtuse-based, only 2 cm. long and 1 cm. wide,
or 6 cm. long and 2 cm. wide, pinnately nerved, bronzed, with pale
center above, drying opaque; petiole barely 5 mm. long; spikes
terminal, slender, 25-40 mm. long, rather loosely flowered; bracts
round-peltate; ovary ovoid, submucronate; stigma oblique.
Type collected in Peru.
A very beautiful plant, now in cultivation, its exact home not
recorded.
FLORA OF PERU 63
Peperomia microlepis Trelease, sp. nov.
Herba subparva suberecta glabra, caule 2-3 mm. crasso; folia
alterna subrhombeo-elliptica vel obovata emarginulato-acuminata
basi acuta 3-4 cm. longa 2 cm. lata 5-7-nervia in sicco subfirma,
subtus pallida, petiolo 5 mm. longo; spicae terminales 40 mm. longae
1 mm. crassae sublaxiflorae, pedunculo filiformi 15 mm. longo;
bracteae rotundo-peltatae crispae.
Junin: Huacapistana, Stevens 91 (type).
Peperomia micromamillata Trelease, sp. nov.
Herba subparva caespitosa succulenta in sicco fusca, caule 1-3
mm. crasso, internodiis brevibus primo velutinis cito furfuraceo-
exfoliantibus; folia congesta ad nodum circa 6 elliptica utrinque
subobtusa parva 8 mm. longa 3 mm. lata carnosissima opaca 1-nervia
subtus microscopice mamillata apice ciliato-barbata, petiolo brevis-
simo vix 1 mm. longo; spicae (terminales?) 60 mm. longae 1 mm.
crassae laxiflorae, pedunculo vix 10 mm. longo; bracteae rotundo-
peltatae; ovaria ovoidea, stigmate subapicali.
Huanuco: Huacachi near Muna, 1,950 meters, Macbride 3872
(herb. Field Mus. No. 534,935, type). Piedra Grande, 1,500
meters, 3687.
Peperomia milvifolia Trelease, sp. nov.
Herba subparva suberecta in statu juvenili papillosa, caule 2
mm. crasso; folia ad nodum vulgo 4 rhombeo-quadrata vel milvi-
formia subacuta basi cuneata 10-11 mm. longa 8 mm. lata 3-nervia
in sicco modice tenuia viridia, petiolo 1 mm. longo; spicae terminales
et ex axillis superioribus graciles ad 100 mm. longae, pseudopedicellis
longis filiformibus; bracteae rotundo-peltatae.
Type from Santa Cruz, 1,800-2,100 meters, Pearce, December,
1864 (herb. Kew.). There is some doubt as to whether the locality
is Peruvian or Bolivian.
Peperomia minuta A. W. Hill, Ann. Bot. 21: 159. pi. 15. 1907.
A small, glabrous, acaulescent herb resembling P. parvifolia;
leaves 2-3.5 mm. in diameter; berries subellipsoid, minutely punctu-
late-roughened ; stylopodium short-conic-cylindric.
Ancash: Ocros to Chonta, Weberbauer 2776, type. Cuzco: Pau-
cartambo Valley, 3,500 meters, H err era 1387.
64 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia mishuyacana Trelease, sp. nov.
Herba majuscula repens glabra vel ad nodos parce puberula,
caule 2-8 mm. crasso; folia opposita vel ternata ovata subobtuse
acuminata basi acuta 5-6 cm. longa 2.5-3 cm. lata 3-5-nervia in
sicco subtenuia opaca, subtus olivacea minute granulosa, petiolo
5 mm. longo; inflorescentia non visa.
Loreto: Mishuyacu, 100 meters, G. King 582 (herb. Field Mus.
No. 630,002, type).
Peperomia modicilimba C. DC. Bot. Jahrb. 40: 258. 1908.
A rather small, stoloniferous-ascending, often branched, glabrous
herb; leaves alternate, round-ovate or elliptic-ovate, acuminate,
acute-based, 1.5-3 cm. long, 1-1.5 cm. wide, 5-nerved; petiole 1 cm.
long; spikes terminal, 80 mm. long, 1 mm. thick; peduncle 15-30
mm. long; bracts round-peltate; ovary obovoid, contracted at the
base, apiculate; stigma subapical.
Puno: Tambo Frilluma, on rocks, 1,000 meters, Weberbauer
1211, type.
Peperomia monostachya R. & P. Fl. 1: 33. 1798. Piper
monostachyum Vahl, Enum. 1: 343. 1805.
A rather large, glabrous, assurgent herb; leaves alternate, ovate
or subobovate, obtuse at both ends, 5-7-nerved, purple beneath and
lined by the nerves; spikes solitary.
Huanuco: Chinchao, Ruiz & Pavon, type.
An obscure species, apparently closely related to P. procumbens.
Peperomia montana C. DC. in DC. Prodr. 16, pt. 1: 426. 1869.
A small, repent-assurgent or erect, glabrous herb; stem slender,
simple; leaves alternate, lanceolate-oblanceolate or obovate, acute
or obtuse at both ends, 2.5-6.5 cm. long, 1.5-2.5 cm. wide, 5-nerved
with the inner nerves somewhat confluent; petiole 5-10 mm. long;
spikes terminal, 70 mm. long, 1 mm. thick; peduncle 15-20 mm.
long; bracts round-peltate; ovary ovoid, somewhat beaked; stigma
oblique. Neg. 8532.
Type from subandine Peru, Poeppig D1519.
Peperomia multiformis Trelease, sp. nov.
Herba delicata repenti-assurgens, caule filiformi crispo-pubes-
cente vel subvilloso; folia alterna, stolonum orbicularia et 5 mm.
lata, ramulorum adscendentium obovata et vix 10 mm. longa vel
FLORA OF PERU 65
elliptica, utrinque plus minusve acuta, glabrata vel subtus crispo-
pubescentia et glanduloso-granulosa, petiolo vix 5 mm. longo; spicae
terminales filiformes 40-70 mm. longae, pedunculo vix 1 cm. longo.
San Martin: Tarapoto, Spruce 4011 (type in herb. Kew.).
Peperomia muscigaudens C. DC. Bot. Jahrb. 40: 267. 1908.
A small, glabrous epiphyte; stem filiform; leaves 3 or 4 at a node,
obovate, slightly retuse, 8 mm. long, 6 mm. wide, 5-nerved; spikes
terminal, 20 mm. long, 1 mm. thick; peduncle 1 cm. long; bracts
round-peltate; berries ovoid, pointed, striatulate; stigma apical.
Puno: Sandia, in bamboo thickets, 2,500-3,000 meters, Weber-
bauer 736, type.
Peperomia naevifolia Trelease, sp. nov.
Herba subparva stolonifera terrestris, caulibus plus minusve
ramosis erectis in sicco angulosis; folia ad nodum 3-4 lanceolato-
elliptica vel subobovata obtusa basi acuta 10-12 mm. longa 5-10
mm. lata, subtus granulosa et papillis apice setiformibus conspersa,
obscure circa 3-nervia, petiolo 1-2 mm. longo; spicae terminales(?)
20 mm. longae 1 mm. crassae, pedunculo aequilongo; bracteae
rotundo-peltatae, rhachi velutina.
Junin: Huacapistana, 1,800 meters, Killip & Smith 24278 (type
in U. S. Nat. Herb.).
Peperomia naviculaefolia Trelease, sp. nov.
Herba alpina parva vix ramosa glabra, caule 5 mm. crasso inferne
dense foliato, scapo remote bracteato terminate; folia alterna
crassissima profunde V-sulcata vel cymbiformia, oblique lanceolato-
elliptica, 15 mm. longa 7 mm. lata, utrinque acuta, enervia vel
obscurissime 3-nervia granulosa, petiolo gracili 5 mm. longo; spicae
plures ad apicem scapi paniculatae 40 mm. longae 2 mm. crassae
laxiflorae, pedunculo 5 mm. longo; bracteae rotundo-peltatae;
ovarium ovoideum, stigmate apicali.
Junin: Huariaca, 2,850 meters, Macbride 3253 (herb. Field Mus.
No. 534,323, type).
Peperomia nigricans Trelease, sp. nov.
Herba mediocris plus minusve ramosa erecta, caule 3-4 mm.
crasso crispo-pubescente; folia alterna rotundo-ovata utrinque
abrupte subacuta 3.5-4 cm. longa 3-4 cm. lata graciliter 5-nervia,
costa obscure ramosa, in sicco fusca tenuia opaca, subtus nigro-
66 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
granulosa et praesertim ad nervos crispo-pubescentia, petiolo 1-3
mm. longo crispo-pubescente; inflorescentia non visa.
Junin: La Merced, 700 meters, Killip & Smith 23674 (type in
U. S. Nat. Herb.).
Peperomia nigro-oculata Trelease, sp. nov.
Herba parva repenti-assurgens, caule gracili plus minusve piloso ;
folia vulgo quaternata rotundo-rhombea obtuse acuminata basi
acuta 8 mm. longa 6-8 mm. lata obscure 1-3-nervia, subtus sparse
puberula, utrinque punctulis centre nigris vel papillis conspersa,
petiolo 1 mm. longo paullo piloso; spicae terminates, juveniles 10
mm. longae 1 mm. crassae, rhachi velutina, pedunculo 15 mm. longo
sparse piloso; bracteae rotundo-peltatae.
Amazonas: Chachapoyas, Mathews 3231 (type). Junin: Por-
venir, Killip & Smith 25931. Enefias, Killip & Smith 25653 (?).
Yapas, Killip & Smith 25510(1], 25517(1}. Huanuco: Piedra
Grande, 1,500 meters, on mossy trees, 3859.
Peperomia nivalis Miq. Syst. 112. 1843; 111. Piper, pi. 9; Arch.
Soc. Phys. Nat. 23: pi. 4.
A small, succulent, glabrous herb; stem rather thick below and
naked; leaves alternate above, dolabriform, submucronate, cuneate
at the base, only 10-20 mm. long and 5 mm. wide, grooved at the
top; petiole nearly suppressed; spikes several in a narrow panicle
on a slender terminal stalk 1-2 cm. long, scarcely 10 mm. long and
1 mm. thick; peduncle very short; bracts round-peltate; berries
ovoid, apiculate; stigma apical.
Type collected at some unknown locality in the high mountains
near the limit of snow, Philippi. Huanuco: Tomaiquichua near
Ambo, 2,550 meters, 21+12. Without locality, Weberbauer
1721, 2416.
Peperomia non-alata Trelease, sp. nov.
Herba mediocris simplex erecta glabra, caule 3 mm. crasso non
alato; folia alterna lanceolata apice longiattenuata basi acuta 7-9
cm. longa 2-2.5 cm. lata 5-nervia, costa obscure ramosa, in sicco
obscure viridia subopaca, petiolo 5 mm. longo; spicae terminales et
ex axillis superioribus graciles 90 mm. longae, pedunculo 10 mm.
longo; bracteae rotundo-peltatae.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
7708 (herb. Field Mus. No. 629,575, type). "Sacha-chullcu."
FLORA OF PERU 67
Peperomia non-hispidula Trelease, sp. nov.
Herba delicata prostrata glabra, caule gracili; folia alterna sub-
rhombea utrinque subacuta minuta 10 mm. longa 7 mm. lata sub-
pinnatim circa 7-nervia, petiolo brevissimo 1-2 mm. longo; spicae
ex axillis superioribus solitariae filiformes minutae vix 5 mm. longae
laxe pauciflorae; bracteae rotundo-subpeltatae; ovarium lageniforme
sublonge attenuatum, stigmate apicali; baccae subrotundatae
utrinque contractae subalveolatae.
Lima: Lurin, 60 meters, Macbride 5967 (herb. Field Mus. No.
536,971, type).
Peperomia obcordata Presl, Epim. Bot. 223. 1849.
A rather small, glabrous herb; leaves 4 at a node, obcordate,
ciliate-emarginulate, acute-based, 12 mm. long, 6 mm. wide, 1-
nerved; petiole very short; spikes terminal, 25 mm. long, apiculate;
peduncle of equal length; bracts round-peltate; berries round-
ovoid, with pseudocupule; stigma apical.
Type from mountains of Peru.
Peperomia obcordatifolia Trelease, sp. nov.
Herba parva glabra repenti-assurgens, caule filiformi; folia
alterna apice caulis congesta obovata plus minusve emarginulata
basi acuta 10 mm. longa 8 mm. lata 3-nervia, petiolo brevissimo;
spicae terminales 15 mm. longae 1 mm. crassae, pedunculo vix 10
mm. longo; bracteae rotundo-peltatae; baccae rotundo-ovoideae
oblique apicatae, stigmate subapicali.
San Martin: Cerro de Campana, Spruce (type in herb. Kew.).
Peperomia Obex Trelease, sp. nov.
Herba majuscula plus minusve ramosa stolonifero-assurgens
glabra, caule 2-3 mm. crasso interdum versus basin crassissimo; folia
alterna lanceolato-ovata acuminata basi acuta 3-5 cm. longa 1.5-2
cm. lata conspicue 5-nervia et nigro-punctata, in sicco tenuia pur-
purascentia, petiolo 5-10 mm. longo; spicae terminales et ex axillis
superioribus 25 mm. longae breviter pedunculatae.
Huanuco: Muna, Macbride 3992 (herb. Field Mus. No. 535,056,
type).
Peperomia obliqua R. & P. Fl. 1: 32. pi. 51. 1798. Piper
obliquum Pers. Syn. 1: 35. 1805.
A moderately large, ascending, glabrous herb; leaves alternate,
subrhombic-oblanceolate, acuminate, cuneate at the base, 5-6 cm.
68 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
long and 2-3 cm. wide, pinnately nerved; petiole barely 1 cm.
long; spikes 3 or 4 each on short, terminal or sympodial stalks,
2-3 mm. thick, 50 mm. long; peduncle 1 cm. long.
Huanuco: Cochero, Posuso, and Pillao, Ruiz & Pavdn, type.
Department unknown: Callcan, Pearce in 1868 (Kew).
Peperomia obruenda Trelease, sp. nov.
Herba subparva erecta glabra, caule 1-2 mm. crasso; folia ob-
ovato-oblanceolata obtusa ad acuminata basi cuneata 3.5-4.5 cm.
longa 1.5-2 cm. lata pinnatim fere e basi circa 7-nervia, in sicco
chartacea, subtus pallidiora, petiolo gracili 10 mm. longo; spicae
terminales et ex axillis superioribus 60 mm. longae 2 mm. crassae
densiflorae, pedunculo filiformi 2-3 cm. longo; bracteae rotundo-
peltatae undulatae.
Junin: Hacienda Chalhuapuquio, Stevens 202 (type).
Peperomia ollantaitambona Trelease, sp. nov.
Herba subparva adscendens succulenta, caule 2-3 mm. crasso,
internodiis brevibus dense papilloso-puberulis; folia plerumque
opposita elliptica obtusa vel abrupte obtuse breviacuminata basi
acuta 1.5-3 cm. longa 1-2 cm. lata delicate 5-nervia, costa obscure
ramosa, in sicco tenuia pellucida, petiolo 5 mm. longo; spicae ter-
minales et ex axillis supremis 25 mm. longae 1 mm. crassae laxiflorae,
pedunculo 1 cm. longo; bracteae rotundo-peltatae.
Cuzco: Ollantaitambo, 2,300-3,100 meters, F. W. Pennell 13661
(type in herb. Acad. Nat. Sci. Phila.).
Peperomia Opiziana A. Dietr. Sp. PI. 1: 162. 1831. Peperomia
cordifolia Opiz in Presl, Rel. Haenk. 164. 1831. Piper Opizianum
D. Dietr. Encycl. 1: 122. 1841.
A slender, glabrous, spreading herb; leaves alternate, round-
ovate, abruptly short-acuminate, openly shallow-cordate, 5.5-7.5
cm. long, 4.5-6.5 cm. wide, 7-nerved with the costa and upper nerves
1-branched upward; petiole 2-5 cm. long; spikes sympodial, 40 mm.
long, 2 mm. thick; peduncle slender, 7-10 cm. long; bracts round-
peltate; berries round-ovoid, mucronate; stigma oblique.
Type from Peru, Haenke in 1790.
Very near P. longepedunculata.
Peperomia oxyphylla C. DC. Bot. Jahrb. 40: 260. 1908.
A glabrous, slender-stemmed herb ; leaves alternate, lance-elliptic,
acute at both ends, 4-7 cm. long, 1.5-2 cm. wide, 5-nerved with the
FLORA OF PERU 69
inner nerves somewhat confluent below; petiole 5 mm. long; spikes
axillary and terminal, subfiliform, 150-200 mm. long; peduncle 1 cm.
long; bracts round-peltate; berries round, apiculate; stigma subapical.
Junin: La Merced, Chanchamayo Valley, 1,000 meters, Weber-
bauer 1854, type.
Peperomia pachiteana Trelease, sp. nov.
Herba repens vel pendula epiphytica ramis evanescente minute
crispo-pubescentibus exceptis glabra; folia alterna ovata acuminata
basi rotundata vel fere truncata 5-7 cm. longa 3-3.5 cm. lata obscure
pinnatinervia, in sicco tenuia opaca fusca, petiolo 1 cm. longo;
spicae ramulum brevem 2-bracteatum terminantes geminatae 25 mm.
longae 1 mm. crassae breviter pedunculatae.
Huanuco: Puerto Lira, on Rio Pachitea, 300 meters, Killip &
Smith 26831 (type in U. S. Nat. Herb.).
Peperomia Pakipski C. DC. Bot. Jahrb. 40: 258. 1908.
A spreading, glabrous epiphyte, the stem 2 mm. thick; leaves 4
or 5 at a node, oblong-obovate, emarginulate, acute-based, 10-20
mm. long, 12 mm. wide, 3-5-nerved; petiole 5 mm. long; spikes
terminal, 130 mm. long, 2 mm. thick; bracts round-peltate; berries
oblong-ovoid, with a style; stigma apical.
Puno: Between Sandia and Cuyocuyo, 2,000 meters, Weberbauer
869, type.
Peperomia palcana C. DC. Bot. Jahrb. 40: 266. 1908.
An erect, glabrous, branching, succulent herb; stem 4 mm. thick,
with approximate leaf scars below; leaves 4 or 5 at a node, oblong-
obovate, emarginulate, acute-based, 12 mm. long, 7 mm. wide,
1-nerved, the petiole very short; spikes terminal, 70 mm. long, 2
mm. thick; peduncle 10 mm. long; bracts round-peltate; berries
oblong (with pseudocupule?) ; stigma apical.
Junin: Below Palca, on cliffs, 2,500 meters, Weberbauer
1754, type.
Peperomia pampalcana Trelease, sp. nov.
Herba alta furcata vel divaricato-ramosa succulenta, caule 3-5
mm. crasso sparse rigido-villoso; folia opposita vel rare ternata
elliptica vel rotundo-ovata obtuse breviacuminata basi rotundata
vel subacuta 4 cm. longa 3.5 cm. lata, superiora 1.5 cm. longa 1 cm.
lata vel ad ramulos ultimos floriferos 8 mm. longa et 5 mm. lata, 3-
vel longiora 5-nervia in sicco firma brunnea, subtus flavescentia laxe
70 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
villosa, petiolo 1 cm. longo vel in foliis reductis multo breviore
subvilloso; spicae in axillis foliorum reductorum solitariae et quoque
terminales in ramulis brevibus insertis et ut videtur subpaniculatae,
40-60 mm. longae 3 mm. crassae, pedunculo 10-15 mm. longo
sparse piloso; bracteae rotundo-peltatae; baccae ovoideae acutatae
pseudocupula onustae, stigmate apicali.
Ayacucho: Pampalca, 3,200 meters, Killip & Smith 23282 (type
in U. S. Nat. Herb.).
Peperomia parva Trelease, sp. nov.
Herba parva epiphytica erecta glabra; folia alterna obovata
apice rotundata basi acuta 15 mm. longa 10-12 mm. lata pinnati-
nervia in sicco coriacea opaca, petiolo 5 mm. longo; spicae terminales
30 mm. longae 2 mm. crassae, pedunculo duplo breviore.
Junin: Dos de Mayo, Pichis Trail, 1,700-1,900 meters, Killip &
Smith 25881 (type in U. S. Nat. Herb.).
Peperomia parvifolia C. DC. Journ. Bot. 4: 133. 1866; Dahlst.
Svensk. Vet. Akad. Handl. 33, pt. 2: pi. 1 ; Hill, Ann. Bot. 20: pi. 29.
A small, glabrous, acaulescent herb; leaves from a subglobose
corm rooting from the bottom, round or subdeltoid, 5-7 mm. long,
peltate below the middle; petiole 15-25 mm. long; spikes 4-5 mm.
long, closely flowered; peduncle 20 mm. long; bracts subelliptic-
peltate; berries subellipsoid, verrucose; stylopodium short-conic;
stigma apical.
Type from Peru, Pavon, doubtless at very high elevations. High
mountains of Bolivia.
Peperomia parvipunctulata Trelease, sp. nov.
Herba modice parva suffrutescens ramosa glabra, caule geniculato
3 mm. crasso, internodiis brevibus; folia alterna oblanceolata utrinque
acuta 5-7 cm. longa 2-3 cm. lata e dimidio inferiore pinnatinervia
minute granulosa, in sicco tenuia opaca, petiolo 5-10 mm. longo;
spicae terminales et ex axillis superioribus 50 mm. longae 1 mm.
crassae, pedunculo 5-10 mm. longo; bracteae rotundo-peltatae.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
7145 (herb. Field Mus. No. 623,573, type).
Peperomia Pavoniana C. DC. in DC. Prodr. 16, pt. 1:
460. 1869.
An herb with puberulent nodes and leaf nerves; leaves 2-4 at
a node, lance-elliptic, 6 cm. long, 3-5-nerved; petiole 15 mm. long;
FLORA OF PERU 71
spikes axillary, filiform; peduncle 8-10 mm. long; berries globose,
hirtellous.
Type from Peru, Pavdn.
Peperomia Pearcei Trelease, sp. nov.
Herba gracilis ramosa glabra; folia ad nodum circa 3 obovata
apice rotundata basi cuneata 8 mm. longa 5 mm. lata 1-nervia,
pedunculo brevi; spicae terminales 30-40 mm. longae 1 mm. crassae,
pedunculo 1 cm. longo; bracteae rotundo-peltatae; baccae ovoideae
acutatae pseudocupula onustae, stigmate apicali.
Huanuco: Muna, in 1863, Pearce 166 (type in herb. Kew.).
Peperomia pellucida (L.) HBK. Nov. Gen. & Sp. 1: 64. 1815.
Piper pellucidum L. Sp. PL 30. 1753; Hort. Cliff, pi. 4; Philip. Journ.
Sci. 43: 218. /. 117. Micropiper pellucidum Miq. Comm. Phyt. 54.
1840. Piper continuum Haw. Succ. Rev. 198. 1821. Peperomia
concinna A. Dietr. Sp. PI. 1: 164. 1831. Peperomia pellucida var.
pygmaea Kunth, Syn. 1: 117. P. pellucida var. minor Van Heurck &
Muell. Obs. Bot. 115. 1870.
A pellucid, erect, more or less branched, succulent, annual herb ;
leaves round-ovate, obtusely short-acuminate, rounded, truncate,
or subcordate at the base, 1.5-2.5 cm. long, 1-2.5 cm. wide, or even
larger, 3-5-nerved, thin, often pale or whitened beneath; petiole 5
mm. to as much as 20 or 30 mm. long; spikes terminal, axillary, and
sympodial, filiform, 20-30 mm. long, loosely flowered; petiole 1 cm.
long; bracts round-peltate; berries round-ovoid, mucronate, longi-
tudinally ridged; stigma apical.
Loreto: Puerto Arturo, 200 meters, Williams 5128. Iquitos,
abandoned land, Williams 7925; Killip & Smith 27497. Santa Rosa,
Kittip & Smith 28869. La Victoria, Williams 2730. Rio Itaya,
Williams 243. Mishuyacu, in clearing, Klug 1210. Fortaleza, Wil-
liams 4460. Lower Rio Nanay, Williams 275. A species of cosmo-
politan distribution in the tropics. "Sacha-yuyu."
Peperomia percalvescens Trelease, sp. nov.
Herba pendula hie inde transiente paullo crispo-sericea, caule
firmo 2-3 mm. crasso; folia alterna ovata subobtusa basi rotundata
vel obscure cordulata 4 cm. longa 2 cm. lata pinnatinervia in sicco
coriacea opaca, subtus brunnea, petiolo 5-10 mm. longo; spicae
solitariae terminales vel ramulum decidue 2-bracteatum 1 cm. longum
terminantes graciles 150 mm. longae, in statu fructifero 5 mm.
72 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
crassae, pedunculo 10 mm. longo; bracteae rotundo-peltatae; baccae
oblongae oblique acuto-scutulatae, stigmate centrali.
San Martin: Cumbasa, Tarapoto, 360-900 meters, Llewelyn
Williams 5758 (herb. Field Mus. No. 624,780, type).
Peperomia pereneana Trelease, sp. nov.
Herba succulentissima procumbens terrestris plus minusve sparse
transiente molliter pubescens, caule 3-5 mm. crasso; folia alterna
ovata acuminata basi rotundata vel obscure cordulata 4.5-6 cm.
longa 2.5-3.5 cm. lata obscure pinnatinervia, in sicco coriacea opaca,
subtus flavescentia, petiolo 1 cm. longo; spicae terminates et sym-
podiales solitariae vel geminatae et ramulos longibracteatos termi-
nantes, in statu juvenili parvae.
Junin: Colonia Perene", 680 meters, Killip & Smith 25413 (type
in U. S. Nat. Herb.).
Peperomia pergamentacea Trelease, sp. nov.
Herba majuscula repens glabra, caule 3 mm. crasso, internodiis
elongatis; folia alterna ovata acute acuminata basi breviter cordata
vel rotundata, interdum abrupte acuta, 10-15 cm. longa 6-12 cm.
lata submultiplinervia, 3 paribus nervorum lateralium prope basin
nascentibus, pare altero versus medium laminae subpergamentaceae
nascente, petiolo 5-15 cm. longo; inflorescentia non visa.
Junin: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
26063 (type in U. S. Nat. Herb.).
Peperomia perhispidula C. DC. Bot. Jahrb. 40: 257. 1908.
P. hispidula var. perhispidula C. DC. Candollea 1: 335, 392. 1925.
A small, suberect herb; stem slender, hispidulous-canescent;
leaves alternate, subreniform-ovate, 13 mm. wide, 11 mm. long,
densely hispidulous; petiole scarcely 5 mm. long; spikes sympodial,
10-15 mm. long, 1 mm. thick, the peduncle as long; bracts round-
peltate; berries ellipsoid, stipitate, with a style; stigma apical.
Junin: Above Huacapistana, road to Palca, moist shaded cliffs,
2,000 meters, Weberbauer 2014, type. Pariahuanca, 2,400 meters,
Weberbauer 6592. Huacachi, 1,950 meters, 4152.
Peperomia perhispidula var. glabrescens Trelease, var. nov.
In statu juvenili subconspicue hispidula, serius fere glabra.
Cuzco: Below Pillahuata, Rio Yanamayo, 2,000-2,300 meters,
Pennell 14076 (type in herb. Acad. Nat. Sci. Phila.).
FLORA OF PERU 73
Peperomia pertomentella Trelease, sp. nov.
Herba majuscula diffusa terrestris sparse ramosa ferrugineo-
tomentulosa; folia alterna ovata plus minusve acuminata basi
rotundata vel subcordulata 4-6 cm. longa 2.5-4 cm. lata obscure
pinnatinervia, in sicco brunnea coriacea, petiolo 1-2 cm. longo;
spicae sympodiales 1 vel 2 ramulum bibracteatum 1 cm. longum
terminantes 90-100 mm. longae 1 mm. crassae, pedunculo 10-15
mm. longo; baccae oblongae apice angustatae scutulatae, stigmate
centrali.
San Martin: Rio Mayo, Tarapoto, 360-900 meters, Llewelyn
Williams 6202 (herb. Field Mus. No. 623,588, type), 6204.
Peperomia peruviana (Miq.) Dahlst. Svensk. Vet. Akad.
Handl. 33, pt. 2: 32. 1900; Hill, Ann. Bot. 20: pi. 29. Tildenia
peruviana Miq. Nov. Act. Nat. Cur. 19: Suppl. 483. 1843.
A small, glabrous, acaulescent herb; leaves from a corm rooting
from the sides as well as from the bottom; leaves round, 1-2 cm. in
diameter, peltate at the center, thin; petiole 2-5 cm. long; spikes 2-6
cm. long; peduncle elongate; bracts round-ovate, subacute; berries
ellipsoid, minutely reticulate; stylopodium elongate-conic; stigma
apical.
Puno: Lake Titicaca, Gaudichaud 150, type. Araranca, rock
ledges on open slope, 4,200 meters, Pennell 13451. Huanuco: Chas-
qui, on mossy rocks, 3305. Lima: Matucana, 2,400 meters, 2894.
Rio Blanco, Killip & Smith 21539. Loreto: Santa Rosa, Killip
& Smith 28869.
Peperomia peruviana var. major A. W. Hill, Ann. Bot. 21:
151. 1907.
Leaves 30-35 mm. in diameter, thin-coriaceous, peltate at the
middle; petiole 8-14 cm. long; spikes 80 mm. long; peduncle 10 cm.
long; berries ovoid-conic.
Cuzco: Ollantaitambo, Urubamba, at 2,500 meters, Hill
183, type.
Peperomia phyllantha Opiz in Presl, Rel. Haenk. 162. 1831.
A small, erect, somewhat branched herb, glabrous except for a
few soft thick hairs on the upper leaf surface; leaves few, alternate
at the end of the stem, round-ovate, acuminate, cordate, 2-2.5 cm.
long, 1.5-2 cm. wide, 5-7-nerved with some nerves branching;
petiole 1-2 cm. long; spikes umbellately clustered at the end of the
74 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
peduncle, 10 mm. long, 1 mm. thick, nearly sessile, distichously
flowered; bracts round-peltate; berries subglobose, mucronate;
stigma apical.
Type from mountains of Peru, Haenke.
Peperomia pichisensis Trelease, sp. nov.
Herba parva repens epiphytica, ramis erectis, caule gracili
minute molliter villoso; folia alterna rotundata, rotundo-obovata
vel elliptica obtusa vel longiora subacuta basi acuta 5-14 mm.
longa 5-7 mm. lata supra crispo-subvillosa obscure 3-nervia, petiolo
3-5 mm. longo glabro; spicae terminales 30-40 mm. longae 1 mm.
crassae, pedunculo brevissimo; bracteae rotundo-peltatae.
Junin: Dos de Mayo, Pichis Trail, 1,700-1,900 meters, Killip
& Smith 25829 (type in U. S. Nat. Herb.).
Peperomia pilifera Trelease, sp. nov.
Herba mediocris simplex suberecta, caule 1-2 mm. crasso plus
minusve laxe villoso; folia alterna, inferiora obovata, superiora sub-
lanceolata, utrinque subacuta 15-30 mm. longa 6-15 mm. lata
minute pubescentia 3-5-nervia, petiolo 3-10 mm. longo laxe piloso;
spicae terminales et axillares filiformes 20-30 mm. longae, pedunculo
gracillimo 10 mm. longo subvilloso; bracteae rotundo-peltatae;
baccae rotundato-ovoideae mucronatae, stigmate subapicali.
Huanuco: Cani, 2,550 meters, Macbride 3460 (herb. Field Mus.
No. 534,535, type).
Peperomia pillahuatana Trelease, sp. nov.
Herba mediocris assurgens erecta glabra, caule 4-5 mm. crasso
repetite furcato furfuraceo-exfoliante; folia opposita vel ad nodos
superiores 3-4 obovata emarginulata basi cuneata 8-20 mm. longa
8-14 mm. lata opaca coriacea, subtus granulosa, petiolo 3-10 mm.
longo; spicae terminales 80 mm. longae 2-3 mm. crassae, pedunculo
10-15 mm. longo; bracteae rotundo-peltatae; ovaria in alveolis
inserta.
Cuzco: Near Rio Yanamayo, below Pillahuata, 2,000-2,300
meters, F. W. Pennell 14074 (herb. Field Mus. No. 558,291, type).
Peperomia pilosa R. & P. Fl. 1: 32. pi. 50. 1798. Piper pilosum
Vahl, Enum. 1: 338. 1805.
An ascending herb, glabrous except for a few conspicuous soft
hairs on the upper leaf surface; stem 2 mm. thick; leaves alternate,
oblong, acuminate, inequilaterally cordate-auricled, 5-8 cm. long,
FLORA OF PERU 75
2 cm. wide, pinnately nerved, black-dotted beneath; petiole 1 cm.
long; spikes terminal, 40 mm. long, 2 mm. thick; peduncle barely
1 cm. long; bracts round-peltate; ovary mucronate; stigma anterior
at base of the mucro (?).
Huanuco: Posuso, Ruiz & Pawn, type.
Peperomia pinedoana Trelease, sp. nov.
Herba majuscula glabra, caule 2-5 mm. crasso; folia alterna late
ovata acuminata basi subtruncato-rotundata sed ad petiolum abrupte
decurrentia 10-14 cm. longa 7-10 cm. lata palmatim 9-nervia, nervis
lateralibus tantum ramosis, in sicco tenuia viridia, petiolo 5-11 cm.
longo; spicae sympodiales 50 mm. longae 3 mm. crassae, pedunculo
gracili 4 cm. longo; bracteae rotundo-peltatae; ovarium rotundatum,
stigmate subapicali.
Junin: Rio Pinedo north of La Merced, 700-900 meters, Killip
& Smith 23620 (type in U. S. Nat. Herb.).
Peperomia playapampana Trelease, sp. nov.
Herba parva glabrata, caule filiformi, stolonifero-assurgens plus
minusve caespitosa; folia alterna oblonga vel oblongo-ovata apice
paullo attenuate obtusa vel obscure emarginulata basi rotundata
parva 8-10 mm. longa 3-5 mm. lata 1-nervia utrinque plus minusve
transiente pilis paucis mollibus praesertim versus apicem conspersa,
subtus pallido-granulosa, petiolo brevissimo 1 mm. longo; spicae
terminales parvae 20 mm. longae 1 mm. crassae, adultae laxiflorae,
pedunculo filiformi 5-10 mm. longo; bracteae rotundo-peltatae;
baccae sessiles ovoideae submucronatae, stigmate apicali.
Huanuco: Playapampa, 2,700 meters, Macbride 4496 (herb.
Field Mus. No. 535,582, type).
Peperomia pleiomorpha Trelease, sp. nov.
Herba parva stolonifero-assurgens epiphytica, caule 1 mm. crasso
laxe minute piloso; folia alterna stolonum rotundo-obovata minuta,
ramorum floriferorum gradatim majora et elliptico-ovata vel lanceo-
lata 10-15 mm. longa 3-10 mm. lata obtusa basi subacuta minute
adpresso-pubescentia, vel majora tantum ciliata, obscure 3-5-nervia,
petiolo 3-5 mm. longo laxe pubescente; spicae 30 mm. longae 1 mm.
crassae, floribus densiuscule insertis, pedunculo filiformi 5-10 mm.
longo plus minusve pubescente; bracteae rotundo-peltatae.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26426 (type
in U. S. Nat. Herb.).
76 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia plicatifolia Trelease, sp. nov.
Herba subparva glabra stolonifero-assurgens, caule gracili; folia
alterna lanceolata vel lanceolato-oblonga utrinque subacuta 15 mm.
longa 4-5 mm. lata e parte inferiore pinnatinervia vulgo plicata
coriacea flavescentia, petiolo brevissimo; spicae terminates 30 mm.
longae 1 mm. crassae, pedunculo 10 mm. longo; bracteae rotundo-
peltatae; baccae piriformes curvae stipitatae; stigma subapicale.
Puno: Sangaban, Lechler 2231 (type in herb. Kew.).
Peperomia plurispica Trelease, sp. nov.
Herba mediocris stolonifera assurgens, caule 2-3 mm. crasso vel
ad basin multo crassiore crispo-piloso; folia ad nodum vulgo 4-5
elliptico-oblonga utrinque subacuta subparva 15-30 mm. longa
5-10 mm. lata supra glabrata, subtus ad nervos crispo-pilosa et plus
minusve conspicue nigro-punctulata, petiolo brevi 2-3 mm. longo
crispo-piloso; spicae terminales et axillares filiformes 100 mm.
longae et ultra 1 mm. crassae laxiflorae, pedunculo 2.5 cm. longo
adpresso-villoso; bracteae rotundo-peltatae majusculae; ovaria im-
mersa ovoidea, stigmate subapicali.
Huanuco: Cani near Mito, 2,550 meters, Macbride 3414 (herb.
Field Mus. No. 534,485, type).
Peperomia Poeppigii Miq. Syst. 84. 1843.
Glabrous except for the leaves, which are ciliate and obscurely
puberulent on the nerves beneath, rooting from the nodes; leaves
alternate, succulent, elliptic, blunt-acuminate, obtuse at the base,
9 cm. long, 6.5 cm. wide, 9-nerved from near the base, with cross
venation; petiole 1 cm. long, ciliate, hirtellous; spikes axillary and
terminal; bracts round-peltate; ovary ovoid, with apical stigma.
Loreto (?): Mainas, Poeppig, type.
Peperomia polycephala Trelease, sp. nov.
Herba parva glabra acaulis; folia ex apice rhizomae crassae
compacte ramosae nascentia orbicularia ad centrum peltata 2-4 cm.
lata in sicco tenuia, petiolo 5 cm. longo et ultra; spicae filiformes
90-100 mm. longae, pedunculo gracillimo petiolum subaequantes;
bracteae rotundo-peltatae; baccae ellipsoideo-ovoideae, stylopodio
gracili, stigmate apicali.
Cuzco: Gucay, Urubamba Valley, 3,000 meters, Fortunato L.
Herrera 1460, type.
FLORA OF PERU 77
Peperoniia polymorpha Trelease, sp. nov.
Herba parva rhizomatoso-erecta arboricola, caule gracili crispo-
pubescente; folia alterna rotundo-obovata ad elliptica obtusa basi
acuta 6-8 mm. longa 3-6 mm. lata supra primum crispo-pubescentia,
obscure 3-nervia, petiolo 3 mm. longo glabro; spicae terminales 30
mm. longae, pedunculo brevissimo glabro; bracteae majusculae
rotundo-peltatae.
Ayacucho: Aina, 700-1,000 meters, Killip & Smith 22774 (type
in U. S. Nat. Herb.).
Peperoniia pongoana Trelease, sp. nov.
Herba modice parva erecta ramosa, petiolis et caulibus transiente
puberulis exceptis glabra, caule 1-2 mm. crasso in sicco sulcato;
folia ad nodum 3-4 lanceolata obtusa basi acuta 15-25 mm. longa
5-10 mm. lata 3-nervia, subtus pallida subpapillosa, petiolo 2-5
mm. longo; spicae terminales et ex axillis superioribus filiformes 60
mm. longae, pedunculo 5 mm. longo; bracteae ellipticae peltatae.
Loreto: Pongo de Manseriche, near mouth of Rio Santiago, 400
meters, Ynes Mexia 6210a (type in herb. Univ. Calif.); also at 210
meters, 6196a.
Peperomia pontina Trelease, sp. nov.
Herba repens intertexta epiphytica, caulibus gracilibus, caule,
petiolis et nervis folii subtus plus minusve crispo-villosis; folia
alterna rotundo-cordata obtuse subacuminata 10-25 mm. longa
10-25 mm. lata 7-nervia viridia et purpurascentia, petiolo 5-10
mm. longo; spicae axillares(?) 20 mm. longae 1 mm. crassae, pedun-
culo filiformi 10-15 mm. longo.
Junin: Near Peren Bridge, Paucartambo Valley, 700 meters,
Killip & Smith 25366 (type in U. S. Nat. Herb.; dupl. in herb.
Field Mus.).
Peperoniia Ppucu-ppucu Trelease, Journ. Wash. Acad. Sci.
16: 206. 1926.
A moderately small, glabrous herb, rooting from many nodes;
leaves crowded, about 3 at a node, round-elliptic, rounded at both
ends, often emarginulate, drying opaque and without evident
nerves, 10 mm. long and wide, the petiole 2 mm. long; spikes termi-
nal, 6 mm. long, 2 mm. thick, rather densely flowered, with anas-
tomosing ridges, the peduncle 5 mm. long; bracts round-peltate,
rather large; berries ovoid-attenuate, with pseudocupule; stigma
apical.
78 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Cuzco: Ollantaitambo, Urubamba Valley, 2,800 meters, Herrera
802, type, 3357. "Ppucu-ppucu."
Peperomia praeruptorum Trelease, sp. nov.
Herba majuscula plus minusve ramosa suberecta rupicola, caule
1-3 mm. crasso sparse crispo-piloso ; folia ad nodum 2-4 elliptica
utrinque acuta 2-4 cm. longa 1-1.5 cm. lata 5-nervia subtus palli-
diora ad nervos sublaxe pubescentia, petiolo 5-15 mm. longo crispo-
pubescente; spicae terminales et axillares 80-100 mm. longae 1 mm.
crassae, pseudopedicellis brevibus, pedunculo filiformi 1-2 cm.
longo; bracteae rotundo-peltatae; baccae rotundato-ovoideae acuta-
tae, stigmate apicali.
Cuzco: Rio Yanamayo below Pillahuata, 2,200-2,400 meters,
F. W. Pennell 14033 (herb. Field Mus. No. 558,268, type).
Peperomia procumbens C. DC. Journ. Bot. 4: 141. 1866.
A glabrous, spreading herb; stem 3-5 cm. thick; leaves alternate,
orbicular or round-ovate, very obtuse, more or less shallow-cordate,
4-7 cm. long, 4-8 cm. wide, 5-nerved, white-banded between the
purple nerves; petiole 5-10 cm. long or more; spikes 1 or 2 on a slender
stalk equaling the petiole, 50-70 mm. long, 4 mm. thick; peduncle
2 cm. long.
Type collected in Peru by Pa von. Huanuco: Posuso, Pearce
585. Loreto: Yurimaguas, Spruce, May, 1855. San Martin:
Tarapoto, Spruce 4279, 4279a.
Peperomia profissa Trelease, sp. nov.
Herba mediocris glabra, caule 2 mm. crasso (inferne crassiore?) ;
folia vulgo ternata lanceolata acuminata basi subacuta 4 cm. longa
1.5 cm. lata 5-nervia, in sicco tenuia viridia, subtus pallidiora
minute glanduloso-granulosa, petiolo gracili 10 mm. longo; spicae
terminales 100 mm. longae 3-4 mm. crassae densiflorae, petiolo
crassiusculo 15 mm. longo; bracteae rotundo-peltatae; baccae ob-
longo-ovoideae acutatae, pseudocupula basilari, stigmate oblique
apicali.
Junin: Hacienda Chalhuapuquio, Stevens 193 (type).
Peperomia pruinosifolia Trelease, sp. nov.
Herba parva succulentissima foliosa rupicola, caule 3-4 mm.
crasso, internodiis brevibus in statu adulto plus minusve pruinosis;
folia alterna sed congesta oblique ovata vel elliptica submucronata
FLORA OF PERU 79
basi acuta 8 mm. longa 5 mm. lata minute glandulosa et in statu
adulto valde pruinosa, interdum plicata, opaca, nervis non
manifestos, petiolo 3 mm. longo; inflorescentia non visa.
Lima: Rio Blanco, 3,000-3,500 meters, Kittip & Smith 21742
(type in U. S. Nat. Herb.).
Peperomia pseudo-furcata C. DC. in DC. Prodr. 16, pt. 1:
399. 1869.
A slender-stemmed, repent, glabrous herb; leaves alternate,
round-ovate, acute, slightly cordate, 3.5-4.5 cm. long, 2.5-3.5 cm.
wide, 7- or obscurely 9-nerved; petiole 2-3 cm. long; spikes sympo-
dial, 90 mm. long, 2 mm. thick; peduncle 15 mm. long; bracts round-
peltate; berries globose, apiculate, deeply pitted; stigma oblique.
Neg. 8334.
San Martin : Tarapoto, Spruce 4009, type.
Peperomia pseudo-galapagensis Trelease, sp. nov.
Herba parva ramosa, caule gracili 1-2 mm. crasso in sicco fusco
anguloso superne puberulo, inferne furfuraceo-exfoliante; folia ad
nodum 3-6, inferiora reflexa, breviter elliptica ad oblonga obtusa
interdum emarginulata basi subacuta parva 5-7 mm. longa 2-3 mm.
lata subtus granulosa obscurissime pinnatinervia, petiolo brevissimo
vix 1 mm. longo; spicae terminales et ex axillis superioribus parvae
15 mm. longae 1 mm. crassae densiflorae, pedunculo vix 5 mm. longo
glabro; bracteae rotundo-peltatae.
Lima: Lima, Wilkes Expl. Exped. (type in Gray Herb.). San
Geronimo, 5920.
Peperomia pseudorhynchophora C. DC. in DC. Prodr. 16,
pt. 1:425. 1869.
A rather large, repent-ascending, glabrous herb; leaves alter-
nate, ovate, somewhat acuminate, subtruncately shallow-cordate,
12-15 cm. long, 8-10 cm. wide, multiple-nerved from the lower
third; petiole 8 cm. long; spikes axillary, 1 (or 2?) at the end of a
short-bracted stalk, 90 mm. long, 3 mm. thick; peduncle 2 cm. long;
bracts round-peltate; berries ovoid, with a short beak; stigma ante-
rior at the base of the beak. Neg. 8529.
Huanuco: Cochero, Poeppig in 1830, type.
Peperomia pseudo-salicifolia Trelease, sp. nov.
Herba subparva subsimplex caespitosa erecta glabra, caule modice
gracili 2-3 mm. crasso; folia alterna, inferiora caduca, oblanceolata
80 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
subacuta vel obtusa basi cuneata 4.5-6 cm. longa 1-1.2 cm. lata
pinnatinervia, ramis delicatis costae circa 4x2, petiolo vix ullo,
lamina ad basin decurrente, petiolo ipso lineari-decurrente; spicae
circa 4 terminates et ex axillis superioribus ut videtur propter folia
fulcrantia decidua false paniculatae 50-70 mm. longae 1 mm.
crassae laxiflorae; bracteae rotundo-peltatae; baccae non visae.
Huanuco: Rio Huallaga Canyon below Santo Domingo, 1,200
meters, Macbride 4254 (herb. Field Mus. No. 535,334, type).
Peperomia puberulaeformis Trelease, sp. nov.
Herba mediocris glabra epiphytica arboricola, caule brevi e
rhizomate crasso erecto; folia pauca elliptica obtusa ut videtur prope
basin rotundata peltata 6 cm. longa 4 cm. lata in sicco viridia
tenuissima translucentia circa 7-nervia, petiolo 3 cm. longo; spicae
150 mm. longae 2 mm. crassae succulentae, pedunculo 2-4 cm.
longo; bracteae rotundo-peltatae; ovarium ovoideum infra stigma
subcapitatum angustatum.
Ayacucho: Kimpitiriki, 400 meters, Killip & Smith 22981 (type
in U. S. Nat. Herb.).
Peperomia puberulibacca C. DC. Bot. Jahrb. 40: 260. 1908.
A moderate-sized, glabrous, somewhat branching herb; leaves
alternate, subelliptic, acute at both ends, 3-5 cm. long, 1.5-2 cm.
wide, 5-nerved, black-punctulate; petiole barely 10 mm. long; spikes
terminal, filiform, 90 mm. long; peduncle scarcely 10 mm. long;
bracts round-peltate; berries densely puberulent, globose, stipitate,
mucronate; stigma subapical.
Junin: Janangu Mountains east of Huacapistana, 2,100 meters,
Weberbauer 2111, type.
Peperomia puberulibacca var. atricha Trelease, var. nov.
A typo foliis subtus pilosulis non nisi foliis glabris differt; baccae
non visae.
Huanuco: Muna, 2,100 meters, Macbride 4009 (type in herb.
Field Mus. No. 535,073).
Peperomia puberulipes Trelease, sp. nov.
Herba modice parva diffusa essentialiter glabra, caule subgracili
2-3 mm. crasso in sicco sulcato; folia ad nodum vulgo 3-4 rhombea
obtusa basi acuta parva 2-2.5 cm. longa 1-1.5 cm. lata, in sicco plus
minusve revoluta 3- vel obscure 5-nervia, petiolo brevissimo 2 mm.
longo microscopice ut costa subtus puberulo; spicae terminales
FLORA OF PERU 81
100 mm. longae 2 mm. crassae densiflorae, pedunculo spicae aequi-
crasso 3.5 cm. longo; bracteae rotundo-peltatae; antherae oblongae
0.5 mm. longae.
Huanuco: Piedra Grande near Rio Santo Domingo, 1,500 meters,
Macbride 3856 (herb. Field Mus. No. 534,919, type).
Peperomia pubescentinervis Trelease, sp. nov.
Herba majuscula sed delicata ramosa, caule 1-2 mm. superne
crasso transiente sparse molliter piloso; folia vulgo ternata lance-
olato-ovata acute acuminata vulgo basi acuta 4-7 cm. longa 2-3
cm. lata 5-nervia, nervis basi 5 mm. longe confluentibus, in sicco
tenuia viridia lucida, supra ad nervos pubescentia, subtus pallidiora
plus minusve punctata; spicae terminales et axillares filiformes 60
mm. longae sublaxiflorae, pedunculo delicatissimo 15-20 mm. longo;
bracteae rotundo-peltatae.
Junin: Hacienda Chalhuapuquio, Stevens 127 (type).
Peperomia pubinervosa Trelease, sp. nov.
Herba repens ubique velutina, caule 3 mm. crasso, inter nodiis
breviusculis; folia orbicularia abrupte brevissime acuminata basi
cordata sinu clause 9 mm. longa 8 mm. lata palmatim 9-11-nervia,
in sicco tenuia viridia, petiolo 3-12 cm. longo; inflorescentia non visa.
Junin: Near Peren Bridge, Rio Paucartambo Valley, 700 meters,
Killip & Smith 25769 (type in U. S. Nat. Herb.), 25269.
Peperomia pubiramea Trelease, sp. nov.
Herba majuscula pendula epiphytica, internodiis subgracilibus
longis microscopice crispo-puberulis; folia alterna elliptica anguste
acuminata basi subacuta 12-15 cm. longa 5-6 cm. lata e dimidio
inferiore pinnatinervia subtus sparsissime minute pilosa, in sicco
tenuia pellucida, petiolo 10-15 mm. longo; spicae geminatae ramulos
anguste bibracteatos sympodiales terminantes 150 mm. longae 2-3
mm. crassae, pedunculo 15 mm. longo; bracteae rotundo-peltatae;
baccae oblongae acute oblique scutulatae, stigmate in scutulo
inserto.
Junin: Near Perene Bridge, Rio Paucartambo Valley, 700
meters, Killip & Smith 25360 (type in U. S. Nat. Herb.).
Peperomia pumila Opiz in Presl, Rel. Haenk. 163. pi. 30. 1831.
A small, erect, crisp-pubescent herb; leaves alternate, elliptic-
ovate, 2-2.5 cm. long, 1.5-2 cm. wide, 3-nerved, black-punctulate;
petiole 5-10 mm. long; spikes terminal and from the uppermost
82 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
axils, 40-50 mm. long, 1 mm. thick; peduncle 10-15 mm. long,
rather loosely flowered ; bracts round-peltate.
Hudnuco: Huanuco mountains, Haenke in 1790, type. "Song-
air," Lechler 2691 in part, at Kew.
Peperomia purpurea R. & P. Fl. 1: 32. pi. 49. 1798. Piper
purpureum Vahl, Enum. 1 : 343. 1805.
A rather large, glabrous herb; stem flexuous, 3-4 mm. thick,
with short internodes, branching; leaves alternate, ovate, acumi-
nately narrowed at both ends, 4-7 cm. long, 2-4.5 cm. wide, sub-
multiple-nerved; petiole 1 cm. long; spikes sympodial, 100 mm.
long, 2-3 mm. thick, or larger; peduncle short; bracts round-peltate.
Huanuco: Chinchao, Ruiz & Pawn, type.
Peperomia puteolata Trelease, sp. nov.
Herba alta glabra, caule 3 mm. crasso in sicco valde angulato;
folia vulgo ternata lanceolata vel ovata acuminata basi acuta 5-6
cm. longa 2-2.5 cm. lata 5-7-nervia, nervis supra profunde impressis
subtus prominentibus, in sicco dura, supra impresso-punctulata,
subtus concolori-glanduloso-granulosa, petiolo vix 5 mm. longo;
spicae terminales 140 mm. longae 2-3 mm. crassae densiflorae,
pseudopedicellis columnaribus, pedunculo 10 mm. longo; bracteae
rotundo-peltatae.
Junin: Huacapistana, Stevens 66 (type); Kittip & Smith 24155,
24190.
Peperomia quadratifolia Trelease, sp. nov.
Herba parva repens epiphytica ad nodos transiente plus minusve
pilosa, caule filiformi radicante; folia ad nodum circa 4 rhombeo-
quadrata obtuse subacuminata basi acuta, in sicco opaco-viridia
1-nervia; spicae terminales filiformes 40 mm. longae, pedunculo 20
mm. longo; bracteae rotundo-peltatae, rhachi glabra sublaxiflora;
baccae ovoideae acutatae pseudocupula onustae; stigma apicale.
Junin: Schunke Hacienda, 1,400-1,700 meters, Killip & Smith
24816 (type in U. S. Nat. Herb.).
Peperomia quadricoma Trelease, sp. nov.
Herba subparva repetite furcata glabra, caule 1-2 mm. crasso;
folia ad nodum vulgo 3-4 elliptico-ovata subemarginulata basi
acuta 12-15 mm. longa 7 mm. lata plus minusve 3-nervia, costa
obscure ramosa, in sicco tenuia fusca opaca, petiolo vix 5 mm.
FLORA OF PERU 83
longo; spicae terminales, juveniles 15 mm. longae 1 mm. crassae,
pedunculo gracili aequilongo; bracteae rotundo-peltatae.
Junin: Chanchamayo Valley, 1,800 meters, Carlos Schunke 370
(herb. Field Mus. No. 571,424, type), A85.
Peperomia quaerata Trelease, sp. nov.
Herba modice parva repens vel assurgens epiphytica, caule 1-2
mm. crasso sparse pallido-villoso; folia ad nodum 3-5 rhombea
obtusa vel subacuta basi cuneata 10-15 mm. longa 10 mm. lata
subtus pallido-villosa, in sicco tenuia opaca, petiolo 1-2 mm. longo
villoso; spicae terminales 80 mm. longae 1 mm. crassae, pedunculo
20 mm. longo; bracteae rotundo-peltatae; flores in alveola elongata
insertae.
Junin: Rio Pinedo, La Merced, 700-900 meters, Killip & Smith
23653 (type in U. S. Nat. Herb.).
Peperomia quaesita Trelease, sp. nov.
Herba modice parva erecta sparse ramosa epiphytica, caule 1-2
mm. crasso plus minusve crispo-hirtello ; folia ad nodum 3-6 rhombea
vel inferiora deltoidea subacuta basi cuneata 10-15 mm. longa 5
mm. lata, in sicco tenuia opaca, glabra, petiolo vix 1 mm. longo
glabrato; spicae terminales, juveniles 15 mm. longae 1 mm. crassae,
pedunculo 2 mm. longo; bracteae rotundo-peltatae.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26676
(type in U. S. Nat. Herb.).
Peperomia quatrometralis Trelease, sp. nov.
Herba pendula epiphytica usque ad 4 m. longa foliis ciliatis
exceptis glabra, caule gracili; folia alterna lanceolato-oblonga vel
elliptica acuta basi obscure cordulata 5-7 cm. longa 2-2.5 cm.
lata in sicco opaca fusca obscure pinnatinervia, rare subtus pilosa,
petiolo 5 mm. longo; spicae terminales et sympodiales ramulos
bibracteatos 2 cm. longos terminantes, pedunculo vix 5 mm. longo;
bracteae rotundo-peltatae; baccae oblongae truncatae, scutulo
lateraliter acutato, stigmate apicali.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26330 (type
in U. S. Nat. Herb.).
Peperomia quimiriana Trelease, sp. nov.
Herba majuscula repenti-assurgens glabra, caule 3-5 mm. crasso;
folia alterna elliptica apice abrupte acuta vel rotundata basi acuta
vel in foliis infimis cuneata 10-12 cm. longa 5-7 cm. lata e 2 tertiis
84 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
inferioribus graciliter pinnatinervia, petiolo 2-6 cm. longo; spicae
solitariae ramulos breves bibracteatos sympodiales terminantes 110
mm. longae 3 mm. crassae, pedunculo 2-4 cm. longo; bracteae
parvae rotundo-peltatae; baccae ellipsoideae, rostro gracili, stig-
mate ad basin rostri inserto.
Junin: East of Quimiri Bridge, La Merced, 800-1,300 meters,
Killip & Smith 2386 '4 (type in U. S. Nat. Herb.). La Merced, Kittip
& Smith 23730.
Peperomia quispicanchiana Trelease, sp. nov.
Herba erecta rhizomatosa plus minusve ramosa glabra in sicco
pallida, caule 3 mm. crasso, internodiis inferioribus elongatis;
folia ad nodum 4-6 rotundo-elliptica obtusa vel inflexo-emarginulata
basi subobtusa 6-10 mm. longa 5-8 mm. lata revoluta, in sicco
dura opaca nervis non manifestis, petiolo 3 mm. longo; spicae
terminales 35 mm. longae 3 mm. crassae, pedunculo vix 10 mm.
longo; bracteae rotundo-peltatae, rhachi pubescente, floribus in
alveolis insertis.
Cuzco: Marcapata, Quispicanchi, Weberbauer 7802 (herb. Field
Mus. No. 605,156, type).
Peperomia racemifolia Trelease, sp. nov.
Herba simplex erecta glabra, caule 2-3 mm. crasso inferne 20-30
cm. longe aphyllo; folia alterna subpauca ad 2-3 cm. supremi caulis
congesta oblanceolata obtuse acuminata basi cuneata 6-8 cm. longa
2 cm. lata ubique pinnatinervia, petiolo gracili 2 cm. longo; spicae
terminales 100 mm. longae 1 mm. crassae, floribus subdistantibus,
pedunculo filiformi 3 cm. longo; bracteae rotundo-peltatae, rhachi
obscure subpuberula.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26657
(type in U. S. Nat. Herb.).
Peperomia rhodophylla Trelease, sp. nov.
Herba parva succulentissima petrophila opaca sed vix plus quam
minutissime papillosa, caule 2-4 mm. crasso; folia ad nodum circa
4 subovato-elliptica obtusissima basi subacuta 16 mm. longa 8
mm. lata vix nervosa, petiolo 2-3 mm. longo; spicae terminales,
juveniles 10 mm. longae 1 mm. crassae densiflorae, pedunculo
aequilongo filiformi ; bracteae rotundo-peltatae.
Cuzco: Ollantaitambo, 2,900-3,100 meters, F. W. Pennell 13654
(herb. Field Mus. No. 558,031, type), 13661 (of very different
appearance).
FLORA OF PERU 85
Peperomia rhombea R. & P. Fl. 1: 31. pi. 46. 1798; Dahlst.
Svensk. Vet. Akad. Handl. 33, pt. 2: pi. 3. Piper rhombeum Vahl,
Enum. 1: 353. 1805. Peperomia reflexa var. peruviana Miq. Syst.
174. 1843.
A cespitose, moderate-sized herb, sparsely rusty-hirsute; stem
2-3 mm. thick, somewhat branched; leaves 4 at a node, rhombic-
lanceolate, gradually acuminate at both ends, 15-20 mm. long r
5-10 mm. wide, leathery, 5-nerved, drying purplish; petiole very
short; spikes terminal, 20-50 mm. long, 1-2 mm. thick; peduncle 2
cm. long; bracts round-peltate; berries ovoid, pointed, with pseudo-
cupule; stigma apical.
Huanuco: Pillao and Chachahuassi, Ruiz & Pawn, type. Ya-
nano, 1,800 meters, on mossy tree branch, 3767. Junin: La Merced,
600 meters, shaded tree trunk, 5474. Chanchamayo Valley, 1,000
meters, Schunke A82, 445a. Without locality: Haenke; Poeppig
in 1829.
Peperomia rhombeifolia Trelease, sp. nov.
Herba parva rhizomatosa caespitosa epiphytica glabra minute
papillosa, caule 3 mm. crasso; folia alterna congesta rhombeo-
lanceolata 2.5 cm. longa 1 cm. lata opaca obscure 5-nervia, petiolo
5 mm. longo; spicae axillares in sicco nigrescentes.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28866 (type in
U. S. Nat. Herb.).
Peperomia rhombeo-elliptica Trelease, sp. nov.
Herba delicata repens essentialiter glabra arboricola pendula;
folia ad nodum 2-5 elliptica vel rhombea utrinque aequaliter rotun-
data basi magis acuta parva 9-12 mm. longa 6 mm. lata vel infima
magis rotundata et 3 mm. tantum longa tenuia obscure 3-nervia,
costa superne pauciramosa, petiolo brevissimo 1 mm. longo puberulo;
spicae terminales 15 mm. longae 2 mm. crassae densiflorae, pedunculo
5-10 mm. longo; bracteae rotundo-peltatae glandulosae.
Huanuco: Huacachi near Muna, 1,950 meters, Macbride 4171
(herb. Field Mus. No. 535,246, type). Junin: Yapas, Killip &
Smith 25510(1}.
Peperomia rhombiformis Trelease, sp. nov.
Herba modice alta silvicola ad nodos fugaciter crispo-pubescens,
basibus petiolorum ciliatis; folia alterna rhombice rotundo-ovata vel
obovata breviter acuminata basi cuneata 5-8 cm. longa 4-6 cm. lata
86 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
7-9-nervia, paribus 1-2 nervorum interiorum cum costa 1-2 cm.
longe confluentibus, supra minute nigro-granulosa, subtus grosse
granulosa, petiolo 1-2 cm. longo; inflorescentia non visa.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
6996 (herb. Field Mus. No. 629,618, type).
Peperomia rhombilimba Trelease, sp. nov.
Herba mediocris assurgens, caule 2-3 mm. crasso sparse crispo-
pubescente; folia alterna rhombea vel inferiora rotundo-obovata
utrinque acuta 5-6 cm. longa 3.5 cm. lata subpinnatim 7-nervia ad
costam plus minusve crispo-pubescentia, petiolo gracili 5-10 mm.
longo; spicae terminales filiformes fere 150 mm. longae pseudopedi-
cellis onustis, pedunculo 15-20 mm. longo; bracteae rotundo-
peltatae; baccae rotundatae mucronatae, stigmate oblique in
mucrone inserto.
Ayacucho: Aina, 750-1,000 meters, Killip & Smith 22564 (type
in U. S. Nat. Herb.).
Peperomia roqueana Trelease, sp. nov.
Herba compacta parva glabra repenti-assurgens vel suberecta;
folia alterna elliptico-obovata basi acuta apice truncata vel emargin-
ulata 15-20 mm. longa 10-15 mm. lata opaca obscure pinnatinervia
petiolo 5 mm. longo; spicae terminales et axillares 15 mm. longae
2 mm. crassae, pedunculo filiformi 20-30 mm. longo; bracteae
rotundo-peltatae; baccae rotundato-ovoideae oblique apicatae
stigmate obliquo.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn William*.
7635 (herb. Field Mus. No. 629,588, type), 7272, 7382.
Peperomia rosea Trelease, sp. nov.
Herba subparva diffusa rubro-tincta arboricola, caule subgracil
2 mm. crasso sordido-hirsuto vel crispo-piloso; folia ad nodum vulgo
3-5 rotundata vel rotundo-elliptica revoluta, subtus et supra prope
basin laxe hirsuta, parva 9-10 mm. longa 5-10 mm. lata coriacea,
nervis non manifestis, petiolo vix 1 mm. longo; spicae terminales et
axillares 80 mm. longae 1 mm. crassae subdensiflorae, pedunculo 2
cm. longo sublaxe piloso; ovaria in rhachi immersa; baccae ovoideae
acutatae pseud ocupulis onustae, stigmate apicali.
Huanuco: Muna, 2,100 meters, Macbride 3927 (herb. Field Mus.
No. 534,989, type).
FLORA OF PERU 87
Peperomia rotundifolia (L.) Dahlst. Svensk. Vet. Akad. Handl.
33, pt. 2: 99. pi. 11. 1800; Ref. Bot. 1: pi. 13. Piper rotundifolium
L. Sp. PI. 30. 1753. Piper nummularifolium Sw. Prodr. 16. 1788.
Peperomia nummularifolia HBK. Nov. Gen. & Sp. 1: 66. 1815.
Acrocarpidium nummularifolium Miq. Syst. 51. 1843. Peperomia
prostrata Hort. Williams ex Gard. Chron. 1879: 716. /. 102; 111.
Hort.26:113.
A very delicate, commonly epiphytic, repent herb; stem filiform,
subcrisply pubescent; leaves alternate, orbicular or slightly obovate,
scarcely 1 cm. in diameter, more or less pubescent, glandular-granular
beneath; petiole 2 mm. long; spikes terminal, 15-25 mm. long, 1 mm.
wide; peduncle barely 10 mm. long, bracteate at the middle; bracts
of the spike round-peltate; berries round-ovoid, apiculate, granular;
stigma oblique.
Ayacucho: Kimpitiriki, Killip & Smith 22894. Junin: La
Merced, 1,200 meters, 5625; Killip & Smith 25410, 23957, 23619,
24603, 24679, 24703; Schunke A87, 221,444- Puerto Bermudez to
Cahuapanas, Killip & Smith 26705. Puerto Bermudez, Killip & Smith
26588. San Nicolas, Killip & Smith 25984. Perene" Bridge, Killip
& Smith 25248. Loreto : Puerto Arturo, Williams 5137. Mishuyacu,
Klug 1198. Santa Rosa, Killip & Smith 28789, 28957. Yuri-
maguas, Killip & Smith 27992, 28038, 28067, 28256. Soledad,
Killip & Smith 29807. Widespread, in scarcely differentiate forms,
from the Antilles into South America, and northward through
Central America.
Humboldt, Bonpland and Kunth seem to have tried to differ-
entiate the obovate and orbicular-leaved forms under the specific
names applied to the West Indian plant by Linnaeus in 1753 and
Swartz in 1788, and report the continental plant under both names.
The similar but opposite-leaved Acrocarpidium rotundifolium of
Miquel has added to the confusion.
Peperomia rubea Trelease, sp. nov.
Herba repens glabra rubescens terrestris, ramis erectis foliosis;
folia ad nodum 2-3 anguste lanceolata versus apicem anguste
rotundatum sensim angustata, basi acutissima 4-5.5 cm. longa
1 cm. lata vel ultra, 3-nervia, nervis supra impressis subtus promi-
nentibus, petiolo 5 mm. longo; inflorescentia non visa.
Loreto: Nanay to Altura, Llewelyn Williams 837 (herb. Field
Mus. No. 617,142, type). "Lancetilla del monte."
88 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia ruhens Trelease, sp. nov.
Herba majuscula subsimplex assurgens glabra, in sicco pur-
purascens, caule 3 mm. crasso; folia ternata lanceolato-elliptica
acute acuminata basi acuta 4-5 cm. longa 1.5-2 cm. lata 3-nervia,
nervis supra impressis subtus prominentibus, petiolo gracili 1 cm.
longo; spicae terminates et ex axillis superioribus 100 mm. longae
2 mm. crassae et ultra, pedunculo 1 cm. longo; bracteae rotundo-
peltatae.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 353
(herb. Field Mus. No. 571,407, type).
Peperomia rubescens C. DC. Bot. Jahrb. 40: 265. 1908.
A glabrous herb; stem slender; leaves 2 or 3 at a node, lance-
olate, subacute at both ends, faintly nerved, barely 3 cm. long
and 1.5 cm. wide; petiole very short; spikes terminal, 60 mm. long,
2 mm. thick, peduncle scarcely 5 mm. long; bracts round-peltate;
berries ovoid, mucronulate, with pseudocupule; stigma apical.
Junin: La Merced, Chanchamayo Valley, 1,000 meters, Weber-
bauer 1944, type.
Peperomia rubrifolia Trelease, sp. nov.
Herba modice alta subsimplex diffusa glabra, caule 2-3 mm.
crasso; folia alterna elliptica vel subovata subacuta basi rotundata
vel latiora basi subcordata 7-10 cm. longa 4-8 cm. lata 9-nervia,
pare intimo nervorum ad costam aliquid adnato, nervis et pagina
inferiore in sicco rubris, petiolo 3-5 cm. longo; spicae sympodiales,
juveniles 20 mm. longae 2 mm. crassae, pedunculo duplo breviore;
bracteae rotundo-peltatae.
Junin: Schunke Hacienda, 1,400-1,700 meters, Killip & Smith
24664 (type in U. S. Nat. Herb.). Porvenir, Killip & Smith 25937.
Enefias, Killip & Smith 25770.
Peperomia rubrispica Trelease, sp. nov.
Herba longiuscula pendula glabra arboricola, caule subgracili 2
mm. crasso, internodiis elongatis; folia alterna ovata acuta vel
acute protracta, interdum basi subpeltata, majuscula 5-12 cm.
longa 3-5.5 cm. lata coriacea e dimidio inferiore pinnatinervia,
ramis delicatis costae circa 3x2, petiolo 6-7 cm. longo; spicae
terminates solitariae vel geminatae ramulum 5-15 mm. longum
terminantes rubrae elongatae 160 mm. longae 2 mm. crassae densi-
florae, pedunculo 1-2 cm. longo; bracteae rotundo-peltatae magnae
plus minusve imbricatae.
FLORA OF PERU 89
Huanuco: Posuso, 600 meters, Macbride J+667 (herb. Field
Mus., type).
Peperomia rufescens C. DC. in DC. Prodr. 16, pt. 1:
416. 1869.
A decumbent, glabrous herb; stem 2-3 mm. thick; leaves alter-
nate, round-ovate or elliptic, obtuse, cordate, 4-6 cm. long, 3.5-4
cm. wide, 7-nerved; petiole 3-5 cm. long; spikes axillary, succulent
and when pressed 10 mm. wide; peduncle 2 cm. long; bracts round-
peltate; berries round, wrinkled; stigma apical.
Type collected in Peru, Pawn 277, as P. clavata sp. nov.
Loreto (?) : Mainas, Poeppig 2322.
Peperomia rufescentifolia Trelease, sp. nov.
Herba gracilis plus minusve ramosa erecta glabra; folia alterna
ovata subacuminata basi rotundata abrupte in petiolum angustata
3.5-5 cm. longa 2.5-4 cm. lata 5-nervia purpurascentia subtus
glanduloso-granulosa, petiolo 1-2 cm. longo; spicae axillares et
terminales filiformes 120 mm. longae sublaxiflorae, pedunculo 5
mm. longo; bracteae rotundo-peltatae; baccae rotundato-ovoideae
mucronatae, stigmate obliquo.
San Martin: Lamas near Tarapoto, 840 meters, Llewelyn Williams
6487 (herb. Field Mus. No. 622,968, type).
Peperomia rugata Trelease, sp. nov.
Herba glabra repens majuscula, caule 3-4 mm. crasso; folia
alterna ovata vel subdeltoideo-ovata acuminata basi obtusa vel
subcordulata, prope basin peltata, 5-9 cm. longa 3-7 cm. lata
obscure submultiplinervia in sicco coriacea flavescentia, supra
minute albido-pustulata, subtus rugulosa, petiolo 1.5-4 cm. longo;
spicae 120 mm. longae 2 mm. crassae, pedunculo 2-3 cm. longo
ramulum brevem 1-bracteatum terminante; bracteae rotundo-
peltatae.
Junin: East of Quimiri Bridge, near La Merced, 800-1,300
meters, Killip & Smith 23823 (type in U. S. Nat. Herb.).
Peperomia rugatifolia Trelease, sp. nov.
Herba parva repens glabra epiphytica, caule 1-2 mm. crasso;
folia ad nodum 3-5 subrhombeo-obovata obtusa basi acuta 10-15
mm. longa 8 mm. lata, in sicco firma lucida supra rugosa, obscure
3-nervia, petiolo vix ullo; spicae terminales (?) filiformes 30 mm.
90 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
longae laxiflorae, pedunculo 5-10 mm. longo; bracteae rotundo-
peltatae.
Junin: Santa Rosa, Pichis Trail, 625-900 meters, Killip &
Smith 26183 (type in U. S. Nat. Herb.).
Peperomia rugatifolia var. exilispica Trelease, var. nov.
Herba repens epiphytica, caule gracili crispo-pubescente; folia
ad nodum vulgo 4 rugosa; spicae 40-70 mm. longae vix 0.5 mm.
crassae.
Junin: Chanchamayo Valley, 1,500 meters, Carlos Schunke 1561
(herb. Field Mus. No. 627,239, type), 177.
Peperomia rupiseda C. DC. ex Hill, Ann. Bot. 21: 148, 157.
pi. 5. 1907; Bot. Jahrb. 40: 256. 1908.
A small, "rhizomatous," acaulescent, glabrous herb; leaves
orbicular, peltate in the middle, about 25 mm. in diameter; petiole
and peduncle elongate; spikes 60 mm. long; bracts obovate-peltate;
berries subglobose, rugulose; stylopodium short; stigma apical.
Lima: Between Matucana and Tambo de Viso, 2,370-2,650
meters, Weberbauer 145, type.
Peperomia salicifolia C. DC. Ann. Cons. Jard. Geneve 2:
281. 1898.
A glabrous herb; stem slender; leaves alternate, linear-lanceolate,
long-attenuate rather than acuminate, more abruptly acute at the
base, 5-6 cm. long, 1-1.5 cm. wide, 5-nerved; petiole 5 mm. long;
spikes about 4, subumbellately clustered close to the uppermost
leaves, 40-50 mm. long, 1 mm. thick; peduncle barely 5 mm. long;
berries subovoid with oblique stigma.
Type collected in Peru, Gay 847.
Peperomia salmonicolor Trelease, sp. nov.
Herba delicata repens epiphytica, saltern ramis obscurissime
pilosulis; folia vulgo ternata suborbicularia apice rotundata basi
subacuta 4-6 mm. lata in sicco subrosea opaca, nervis non mani-
festis, petiolo 1-2 mm. longo; spicae terminales 10 mm. longae 2
mm. crassae rubescentes densiflorae, pedunculo filiformi 15 mm.
longo; bracteae rotundo-peltatae.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
7329 (herb. Field Mus. No. 624,523, type).
FLORA OF PERU 91
Peperomia salmonicolor var. viridis Trelease, var. nov.
Folia minus rubescentia usque ad 10 mm. longa et 6 mm. lata;
spicae ad 15 mm. longae, pedunculo ad 20 mm.; baccae rotundato-
ovoideae acutatae pseudocupula onustae, stigmate apicali.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
7258 (herb. Field Mus. No. 629,593, type).
Peperomia san-buenaventurana Trelease, sp. nov.
Herba modice alta assurgens glabra, caule 2-5 mm. crasso ad
nodos plus minusve incrassato; folia ad nodum 3-6 elliptica vel
subobovata apice rotundata basi acuta 10-15 mm. longa 5-8 mm.
lata subpinnatinervia, petiolo 1-2 mm. longo; spicae ramulos graciles
paucif olios terminantes 50 mm. longae 1 mm. crassae laxiflorae,
pedunculo 10 mm. longo; bracteae rotundo-peltatae; ovarium
rotundato-ovoideum apiculatum, stigmate apicali.
Lima: San Buenaventura, 2,700 meters, F. W. Pennell 14566
(type in herb. Acad. Nat. Sci. Phila.).
Peperomia sangabanensis Trelease, sp. nov.
Herba delicata diffusa, caule filiformi puberulo; folia ad nodum
2-6 lanceolata utrinque acuta vel obovata et apice rotundata 7-8
mm. longa 3-4 mm. lata 3-nervia; inflorescentia non visa.
Puno: Sangaban, Lechler 2378 (type in herb. Kew.).
Peperomia san-roqueana Trelease, sp. nov.
Herba repens glabra arboricola, caule elongato 2-3 mm. crasso;
folia opposita elliptica vel lanceolato-elliptica obtusa basi acuta
4.5-5.5 cm. longa 2-2.5 cm. lata circa 7-nervia subtus minute
granulosa in sicco opaca, petiolo 5-10 mm. longo; inflorescentia
non visa.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
7260 (herb. Field Mus. No. 629,783, type).
Peperomia santa-cruzana Trelease, sp. nov.
Herba suberecta statu juvenili papillosa, caule 2 mm. crasso,
internodiis modice brevibus; folia ad nodum 3-4 rhombeo-obovata
subacuta basi cuneata 12 mm. longa 8 mm. lata 3-nervia in sicco
subtenuia viridia, petiolo brevissimo; spicae terminales et ex axillis
superioribus 40-100 mm. longae 2 mm. crassae, pseudopedicellis
longis filiformibus, pedunculo 15-20 mm. longo; bracteae rotundo-
peltatae.
92 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peru(?): Santa Cruz, 1,800-2,100 meters, Pearce, December,
1864 (type in herb. Kew.).
Peperomia santiagoana Trelease, sp. nov.
Herba majuscula repenti-pendula glabra, caule 2-3 mm. crasso;
folia alterna lanceolato-ovata sensim subacuta basi rotundato-acuta
6-8.5 cm. longa 2.5-3.5 cm. lata multiplinervia, costa ramis 2
fere basilaribus et uno medio utroque latere onusta, in sicco opaca
olivacea, petiolo 1-2 cm. longo; spicae geminatae ramulum 1-brac-
teatum terminale vix 2 cm. longum terminantes 80 mm. longae (et
ultra?) 2 mm. crassae; bracteae rotundo-peltatae; ovarium oblique
scutulatum.
Loreto: Rio Santiago, 200 meters, Ynes Mexia 6146 (type in
herb. Univ. Calif.).
Peperomia sarcostachya Trelease, sp. nov.
Herba brevicaulis comparate magna glabra, caule 3 mm. crasso;
folia pauca alterna rotundo-ovata obtusissima basi cordata 9-10
cm. longa 6.5-8 cm. lata 5- vel obscure 7-nervia, in sicco tenuia,
supra fusca inter nervos ma j ores pallido-vittata, petiolo 2-7 cm.
longo; spicae solitariae laterales carnosae obtusae in sicco 35 mm.
longae 7 mm. crassae, pedunculo erecto gracili 2 cm. longo; bracteae
rotundo-peltatae; ovarium immersum subglobosum, stigmate fere
apicali.
Loreto: Santa Rosa, 135 meters, Kittip & Smith 28889 (type in
U.S. Nat. Herb.).
Peperomia sarcostachya var. repens Trelease, sp. nov.
Herba late repens plus minusve radicans, internodiis ut petiolis
magis elongatis; folia 4.5-10 cm. longa 3-6.5 cm. lata pallido-zonata
et in statu juvenili albo-venulosa; spicae clavatae carnosae in sicco
10 mm. latae, pedunculo 3.5 cm. longo.
Junin: Peren Bridge, Paucartambo Valley, 700 meters, Killip
& Smith 25307 (type in U. S. Nat. Herb.).
Peperomia scabiosa Trelease, sp. nov.
Herba subparva repens furcata epiphytica, caule gracili 1 mm.
crasso glabro inferne ad 3 mm. crasso epidermide exfoliante argenteo-
furfuraceo; folia ad nodum vulgo 3-4 orbicularia vel rotundo-
elliptica vel obovata vulgo emarginulata basi subacuta parva 7-10
mm. longa 6-9 mm. lata subtus brunneo-granulosa et evanescente
plus minusve pubescentia, 3-nervia costa prope medium ramosa,
FLORA OF PERU 93
petiolo 1 mm. longo transiente pubescente; spicae terminates 70
mm. longae 1 mm. crassae laxiflorae, pedunculo vix 1 cm. longo
glabro; bracteae rotundo-peltatae; baccae oblongo-ovoideae pseudo-
cupula onustae oblique acutatae, stigmate apicali.
Huanuco: Piedra Grande near Rio Santo Domingo, 1,500 meters,
Macbride 3857 (herb. Field Mus. No. 534,920, type).
Peperomia scandens R. & P. Fl. 1: 32. pi 51. 1798. Piper
scandens Vahl, Enum. 1: 346. 1805. Acrocarpidium scandens Miq.
Syst. 59. 1843.
A delicate, repent, crisp-pubescent herb, commonly arboricolous;
stem filiform; leaves alternate, round-ovate, subacute, rounded or
truncate at the base or subcordate, 1-2 cm. long and wide, 3-5-
nerved; petiole 5-15 mm. long; spikes terminal on a 1-bracted
filiform stalk 1 cm. long, 10 mm. long, 1 mm. thick, the peduncle of
equal length; bracts round-peltate; berries oblong, slender-bracted;
stigma anterior at the base of the beak.
Huanuco: Chinchao, Cochero, Posuso, and Mocora, Ruiz &
Pavon, type. Posuso, Ruiz; common on tree trunks, logs, and rocks,
600 meters, 4715. Ayacucho: Near Kimpitiriki, Killip & Smith
22866, 22895. Junin: Above San Ramon, 1,300-1,700 meters,
Schunke ASS, 367. Rio Pinedo, 700-900 meters, Killip & Smith
23616. Puerto Bermudez, Killip & Smith 16445. Puerto Yessup,
Killip & Smith 26253, 23613. La Merced, Killip & Smith 23724,
23939. Peren Bridge, Killip & Smith 24963, 25282. Loreto:
Soledad, Killip & Smith 29592, 29725. Puerto Arturo, Killip &
Smith 27772. Balsapuerto, Killip & Smith 28670, 28674. Santa
Rosa, Killip & Smith 28836. San Martin: Tarapoto, Spruce 3952,
the type of var. fertilior C. DC. in DC. Prodr. 16, pt. 1: 434. 1869.
Like P. rotundifolia, a widespread aggregate of interblending forms,
reaching from the Antilles into South and Central America.
Peperomia scandens var. longispica Trelease, var. nov.
Folia longiora, 10-20 mm. longa 12-25 mm. lata; spicae 15-30
mm. longae.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28382
(type in U. S. Nat. Herb.).
The leaves are more like the large ones figured by Ruiz and
Pavon, which are as much as 30 mm. long and 35 mm. wide, but their
specimens are actually small-leaved.
94 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia Schunkeana Trelease, sp. nov.
"Frutex," ramis 5 mm. crassis dense fulvo-villosis, pilis gracilibus;
folia alterna ovata obscure breviter acuminata basi concave trun-
cata 14-15 cm. longa 11-12 cm. lata fere 2 cm. supra basin peltata e
dimidio inferiore multiplinervia, nervis circa 11, utrinque intertexto-
sericea, petiolo circa 7 cm. longo sericeo-villoso; spicae (terminales?)
23 mm. longae 4 mm. crassae fulvo-pilosae densiflorae, pedunculo 2
cm. longo fulvo-villoso; bracteae rotundo-peltatae; baccae ellipsoideae
rostratae; stigma ad basin rostri insertum.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 342
(herb. Field Mus. No. 571,398, type), 56; Killip & Smith 24526.
Peperomia sclerophylla Trelease, sp. nov.
Herba stolonifera assurgens succulenta glabra, caule 2-4 mm.
crasso in sicco sulcato; folia opposita vel ternata lanceolato-ovata
vel elliptico-ovata obtuse acuminata basi acuta 2-3 cm. longa
1-1.5 cm. lata 3-nervia in sicco dura revoluta impresso-punctulata,
petiolo 2-3 mm. longo; spicae terminales 80 mm. longae 2 mm.
crassae, pedunculo gracili 20 mm. longo; bracteae rotundo-peltatae;
baccae ovoideae acutatae pseudocupula onustae, stigmate apicali.
San Martin: Near Tarapoto, 360-900 meters, Llewelyn Williams
6879 (herb. Field Mus. No. 624,519, type).
Peperomia scutaleifolia Trelease, sp. nov.
Herba majuscula arboricola repens, ramis foliatis divergentibus,
foliis et petiolis obscure ciliatis exceptis glabra; folia alterna elliptica
vel lanceolata 4-7 cm. longa 2-5 cm. lata acuminata basi acuta ob-
scure pinnatinervia in sicco brunnea coriacea opaca, petiolo 5-10
mm. longo; spicae sympodiales elongatae; baccae oblongae oblique
acute scutulatae, stigmate centrali.
Junin: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25557 (type in U. S. Nat. Herb.).
Peperomia scutellaefolia R. & P. Fl. 29. pi. 44- 1798. Piper
scutellaefolium Vahl, Enum. 1: 356. 1805. Piper scutellatum Pers.
Syn. 1: 34. 1805.
A small, glabrous, acaulescent herb; leaves from a rather large
corm rooting from the sides and bottom, the blades round-ovate,
peltate toward the base, 30 mm. long, 20 mm. wide; petiole 3-7 cm.
long; spikes rather thick, 50-60 mm. long; bracts round-ovate;
peduncle scarcely equaling the petiole.
Arequipa: Ruiz & Pavon, type.
FLORA OF PERU 95
Peperomia secunda R. & P. Fl. 1: 30. pi. 45. 1798; Miq. 111.
Piper, pi. 22. Piper racemosum Vahl, Enum. 1 : 337. 1805.
A short and thick-stemmed, simple herb, appressed-villous
throughout; leaves congested, alternate, round-ovate, subacuminate,
very shallowly cordate, 4-6 cm. in diameter, submultiple-nerved with
about 6 basal nerves, the costa as well as the longer laterals again
branched; petiole 3-9 cm. long; spikes many, secund on the branches
of an ample, stout-stemmed, terminal panicle, scarcely 10 mm. long
and short-peduncled ; bracts round-peltate; berries ovoid, pointed;
stigma apical. Neg. 8389.
Huanuco: Pillao and Chachahuassi, Ruiz & Pavdn, type.
Peperomia Seleri C. DC. Ann. Cons. Jard. Geneve 21:
247. 1920.
A small, glabrous, acaulescent herb; leaves apparently from a
corm, ovate, peltate below the middle, about 5 mm. in diameter;
petiole 3 cm. long; spikes 20 mm. long; peduncle equaling the petiole;
bracts ovate; ovary suboblong, with a short cylindric stylopodium;
stigma apical.
Lima: Cerro de Amazonas near Lima, on stones, Seler 262, type.
Cerro de San Cristobal, on rocks, Seler 250.
Peperomia semielongata Trelease, sp. nov.
Herba mediocris repens glabra, caule subgracili 2 mm. crasso
in sicco plus minusve quadrangulato; folia alterna ovata obtuse
subacuminata basi truncata vel breviter cordata 4.5-6 cm. longa
3-3.5 cm. lata in sicco viridia multiplinervia, nervis basilaribus 2
paribus, pare altero ramorum e tertio inferiore costae nascente,
petiolo gracili 2.5-3 vel ad 10 cm. longo; spicae terminales 50 mm.
longae 2 mm. crassae dense subverticillatim densiflorae, pedunculo
3 cm. longo medio bracteato; bracteae rotundo-peltatae parvae
fuscae.
Junin: Hacienda Schunke, La Merced, Macbride 5644 (herb.
Field Mus., type); Killip & Smith 24683.
Peperomia semisupina Trelease, sp. nov.
Herba majuscula semidecumbens essentialiter glabra in sicco
fusca, caule 4 mm. crasso, internodiis elongatis; folia alterna late
lanceolata anguste longiacuminata basi rotundata ad acuta ma-
juscula 9-10 mm. longa 3-4 mm. lata e dimidio inferiore sub-
pinnatinervia, ramis delicatis costae circa 4x2, subtus pallidiora
pallido-granulosa, ciliata, petiolo 10-15 mm. longo ciliato; spicae
96 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
oppositifoliae caulem caduci-bracteatum vix 15 mm. longum ter-
minantes 50 mm. longae baccis inclusis 5 mm. crassae densiflorae,
pedunculo filiformi 2 cm. longo; bracteae rotundo-peltatae magnae;
baccae rubrae oblongae oblique acuminato-truncatae, stigmate
subapicali.
Junin: Hacienda Schunke, La Merced, 1,200 meters, Macbride
5765 (herb. Field Mus. No. 536,795, type); Killip & Smith 24575;
Schunke 185, 638.
Peperomia seposita Trelease, sp. nov.
Herba majuscula adscendenti-ramosa foliosa, caule 3 mm.
crasso et ultra subquadrangulari puberulo; folia ad nodum circa 6
oblonga obtusa basi sensim acuta 12-25 mm. longa 2-5 mm. lata
obscure pinnatinervia, supra fusca costa puberula, subtus flaves-
centia glabra minute granulosa, petiolo 1 mm. longo plus minusve
puberulo; spicae terminales et ex axillis superioribius, juvenes 15-
20 mm. longae 1 mm. crassae, pedunculo 1-2 mm. longo subpuberulo;
bracteae rotundo-peltatae rubro-brunneae.
Junin: Hacienda Chalhuapuquio, Stevens 213 (type).
Peperomia septemnervis R & P. Fl. 1: 31. pi. 47. 1798. Piper
septemnerve Vahl, Enum. 1: 349. 1805.
A glabrous, suberect herb, the nodes purplish; leaves 2 or 3 at a
node, lance-elliptic, acuminate, acute-based, 5-8 cm. long, 2-3 cm.
wide, 7-nerved; petiole 5 mm. long; spikes curved, 80-100 mm. long,
4 mm. thick; peduncle 10 mm. long; bracts round-peltate; ovary
ovoid (with pseudocupule?) ; stigma apical.
Huanuco: Posuso, Ruiz & Pawn, type. Piedra Grande, 1,500
meters, 3860. Junin: Yapas, 1,350-1,600 meters, Killip &
Smith 3860.
Peperomia Smithii Trelease, sp. nov.
Herba glabra repenti-assurgens epiphytica, caule 2 mm. crasso,
internodiis longiusculis; folia ad nodum 2-4 late rhombeo-ovata
obtusa vel obtuse acuminata basi subacuta 1.5-2 cm. longa 1-1.5
cm. lata in sicco subtenuia opaca, obscure 3-5-nervia, petiolo 3 mm.
longo; spicae terminales filiformes 70 mm. longae, pedunculo 15 mm.
longo; bracteae rotundo-peltatae.
Loreto: Yurimaguas to Balsapuerto, 135-150 meters, Killip &
Smith 28115 (type in U. S. Nat. Herb.).
FLORA OF PERU 97
Peperomia spectabilis Miq. Lond. Journ. Bot. 4: 417. 1845.
A rather large, hard-stemmed herb, finely crisp- or appressed^
pubescent throughout; leaves alternate or suboppositely congested,
ovate, subacuminate, rounded at the base, 10-13 cm. long, 7-8
cm. wide, pinnately nerved; petiole 3 cm. long; spikes clustered in
the upper axils and paniculately whorled upward, filiform, 100 mm.
long; peduncle 10 mm. long; bracts round-peltate; ovary suboblong,
acutely scutulate; stigma subcentral.
Type collected in Peru, Mathews 1635.
Peperomia spiculata Trelease, sp. nov.
Herba majuscula gracilis glabra nigrescens, caule 1-2 vel inferne
4 mm. crasso alato; folia alterna lanceolata anguste acuminata
basi acuta 5-5.5 cm. longa 1.5-2 cm. lata 5-nervia, in sicco fusca
opaca, petiolo 5 mm. longo amplexicauli-decurrente; spicae termi-
nales et axillares filiformes 100-150 mm. longae subdensiflorae,
pedunculo gracili 10 mm. longo; bracteae rotundo-peltatae; baccae
rotundato-ovoideae mucronatae, stigmate obliquo in mucrone
inserto.
Junin: Chanchamayo Valley, Carlos Schunke 352 (herb. Field
Mus. No. 571,406, type), A88.
Peperomia spiculata var. elliptica Trelease, var. nov.
Folia elliptica 6-6.5 cm. longa 3 cm. lata 7-nervia.
Junin: Chanchamayo Valley, Carlos Schunke 388 (herb. Field
Mus. No. 571,442, type).
Peperomia Sprucei C. DC. Journ. Bot. 4: 133. 1866.
A small, glabrous, acaulescent herb; leaves few, from a corm (?),
elliptic or rounded, obtuse at both ends, 3.5 cm. long, 2.5 cm. wide,
5-nerved, coriaceous, paler beneath; petiole 2-10 cm. long; spikes
filiform, 40-60 mm. long; peduncle 3 cm. long or more; bracts
round-peltate; berries ovoid, with a short style; stigma apical.
Neg. 8391.
San Martin: Tarapoto, Spruce 4931, type.
Peperomia striata R. & P. Fl. 1: 32. pi. 52. 1798. Piper
striatum Vahl, Enum. 1: 343. 1805.
A rather large, simple, erect herb, soft-hairy throughout, the
stem striate; leaves elliptic, acute at both ends or short-acuminate,
6-8 cm. wide, 12-16 cm. long, pinnately nerved; petiole 2-5 cm.
98 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
long; spikes several in a terminal panicle, 70 mm. long, 2-3 mm.
thick; peduncle 10 mm. long; bracts round-peltate; ovary ovoid,
mucronulate, with an anterior stigma.
Huanuco: Cochero and Chinchao, Ruiz & Pavon, type. Cochero,
Dombey; Poeppig.
Peperomia subandina Trelease, sp. nov.
Herba parva repens foliosa glabra, caule 1 mm. crasso; folia
quaternata obovata subtruncata basi cuneata 5 mm. longa 6 mm.
lata in sicco pallida opaca, petiolo 1 mm. longo; spicae terminales
20 mm. longae 1 mm. crassae, pedunculo filiformi 10 mm. longo;
bracteae rotundo-peltatae crenulatae, rhachi hirtella.
Junin: Pichis Trail, 400 meters, Kittip & Smith 26218 (type in
U. S. Nat. Herb.).
Peperomia subcalvescens Trelease, sp. nov.
Herba majuscula delicata plus minusve ramosa, caule 1-2 mm.
crasso, internodiis elongatis glabris; folia ad nodum 4-6 elliptico-
subobovata acuta vel anguste acuminata basi acuta 3-nervia in
sicco tenuia viridia, supra ad nervos paullo pubescentia, subtus
pallidiora glabra, petiolo gracili 4-5 mm. longo; spicae subnumerosae
terminales et ex axillis superioribus filiformes 40 mm. longae sub-
laxiflorae, pedunculo 5-10 mm. longo; bracteae rotundo-peltatae.
Junin: Huacapistana, Stevens 84 (type); Killip & Smith 24321,
24248.
Peperomia subdichotoma Trelease, sp. nov.
Herba mediocris stolonifera erecta pseudo-furcata glabra epi-
phytica, caule 2-3 mm. crasso; folia ad nodum vulgo 3-4 obovata
emarginata basi acuta vel in foliis longioribus cuneata 17-18 mm.
longa 6-10 mm. lata obscure 3-nervia, costa indistincte ramosa,
petiolo vix 2 mm. longo; spicae terminales vix 40 mm. longae et 2
mm. crassae densiflorae, pedunculo circa 1 cm. longo; bracteae
rotundo-peltatae primo superficiales, serius cum ovariis profunde in
fissuris longitudinalibus rhachis immersae.
Huanuco: Muna, 2,100 meters, Macbride 4008 (herb. Field Mus.
No. 535,072, type).
Peperomia subsericata Trelease, sp. nov.
Herba parva stolonifero-caespitosa epiphytica, caule filiformi
laxe evanescente minute piloso; folia alterna lanceolata utrinque
acuta vel inferiora obtusa et elliptica, 10 mm. longa 4 mm. lata
FLORA OF PERU 99
glabra opaca obscurissime 3-nervia, petiolo gracillimo 2 mm. longo;
spicae terminales filiformes 30 mm. longae densiflorae, pedunculo
gracillimo 5 mm. longo; bracteae rotundo-peltatae.
Junin: Chanchamayo Valley, 1,500-1,800 meters, Carlos Schunke
(herb. Field Mus. No. 571,537, type), A81, 366, 369, 380, 545a, 549;
Killip & Smith 24815. Yapas, Killip & Smith 25547, 25513.
Peperomia subvillicaulis Trelease, sp. nov.
Herba mediocris repenti-assurgens rupicola, caule 2 mm. crasso
laxe subvilloso; folia opposita vel ternata obovata et obtusa, vel in
ramis sterilibus typice subobovato-elliptica et utrinque obtusa,
1.5-3.5 cm. longa 1-2 cm. lata, 3- vel obscure 5-nervia, prope basin
plus minusve molliter pilosa vel glabrescentia, petiolo 5-10 mm.
longo molliter piloso; spicae terminales et axillares 30-40 mm.
longae 1 mm. crassae pseudopedicellis brevibus onustae, pedunculo
filiformi 5-15 mm. longo glabrato; bracteae rotundo-peltatae;
baccae ovoideae acutatae, stigmate apicali.
Cuzco: Pillahuata, 2,200-2,400 meters, F. W. Pennell 13990
(herb. Field Mus. No. 558,232, type).
Peperomia sylvatica C. DC. in DC. Prodr. 16, pt. 1: 435. 1869.
A suberect herb, the stem 3 mm. thick, loosely subvillous; leaves
alternate, subrhombic-ovate or elliptic, acute at both ends, submul-
tiple-nerved from the lower half, black-punctulate; petiole 15 mm.
long; spikes terminal and axillary, 110 mm. long, 2 mm. thick;
peduncle 15-20 mm. long; bracts round-peltate; ovary ovoid, sub-
mucronate; stigma apical.
Huanuco: Near Cochero, Poeppig in 1839, type.
Peperomia tarapotana C. DC. Journ. Bot. 4: 137. 1866.
A glabrous, stoloniferous herb; leaves alternate, ovate, subacu-
minate, peltate above the slightly retuse or deeply cordate base,
12-15 cm. long, 11 cm. wide, about 10-nerved; petiole as long as the
blade; spikes panicled, 3-5 at the end of an axillary 4-winged stalk
as long as the petiole, 20-30 mm. long; peduncle short; bracts round-
peltate; berries ovoid, pointed; stigma apical. Neg. 8343.
San Martin: Tarapoto, Spruce 4570, type.
Peperomia tenuiramea C. DC. Bot. Jahrb. 40: 263. 1908.
A glabrous, rather large epiphyte, the stem 3 mm. thick; leaves
alternate, lance-elliptic, acute at both ends, 15-20 cm. long, 5.5-7.5
cm. wide, pinnately nerved; petiole 1 cm. long; spikes few on a short,
100 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
bracted, sympodial or terminal stalk, 150 mm. long, 2 mm. thick;
peduncle 1 cm. long; bracts round-peltate; berries glandular, ellip-
soid, beaked; stigma anterior.
Huanuco: Between Monzon and Rio Huallaga, 600-700 meters,
Weberbauer 3656, type.
Peperomia teresitensis Trelease, sp. nov.
Herba compacta mediocris foliosa, caule ad 5 mm. crasso griseo-
tomentoso; folia alterna elliptico-obovata vulgo obtusa basi acuta
3.5-6.5 cm. longa 2-3.5 cm. lata, subtus 5-nervia, costa superne
pauciramosa, papyracea, supra obscure viridia glabrata, subtus palli-
diora albida ad nervos subvillosa, petiolo ad 2 cm. longo subtomen-
toso; spicae apicales et comparate numerosae ex axillis superioribus
40 mm. longae 1 mm. crassae subdensiflorae, pedunculo vix ullo;
bracteae rotundo-peltatae concavae.
Junin: Teresita, Stevens 88 (type).
Peperomia tetragona R. & P. Fl. 1: 31. pi. 47. 1798. Piper
tetragonum Vahl, Enum. 1: 349. 1805.
A rather coarse, succulent, repent, glabrous herb, the stem stri-
atulate; leaves 2 or 3 at a node, elliptic, acute or acuminate, acute-
based, 6-10 cm. long, 2.5-4.5 cm. wide, 5-nerved; petiole 5 mm.
long; spikes 2 at the end of a reduced-leaved stalk 2 cm. long, curved,
70-100 mm. long, 4 mm, thick; peduncle 2 cm. long; bracts round-
peltate; berries ovoid, with pseudocupule (?); stigma apical.
Huanuco: Chinchao, Ruiz & Pavon, type. San Martin: San
Roque, epiphytic, 1,400 meters, Williams 7198, 7710.
Peperomia tetrica Trelease, sp. nov.
Herba modice parva late repens, ramis paucis adscendentibus,
nodis et petiolis puberulis exceptis glabra, caule 2 mm. crasso; folia
opposita vel ternata lanceolata obtuse acuminata 4 cm. longa 1 cm.
lata 3-nervia in sicco opaca, subtus minute punctulata, petiolo circa
3 mm. longo; inflorescentia non visa.
Loreto: Balsapuerto, 350-550 meters, Killip & Smith 28525
(type in U. S. Nat. Herb.).
Peperomia timbuchiana Trelease, sp. nov.
Herba gracilis diffusa majuscula paullo ramosa, caule minute
evanescente puberulo; folia alterna elliptica vel sublanceolata acute
acuminata basi acuta vel rotundato-acuta, in sicco coriacea, subtus
brunnea, obscure pinnatinervia ; spicae 1-2 ramulum brevem termin-
FLORA OF PERU 101
alem 2-bracteatum terminantes 100 mm. longae, pedunculo filiformi
10 mm. longo; baccae oblongae scutulatae oblique apicatae, stigmate
apicali.
Loreto: Timbuchi, Rio Nanay, Llewelyn Williams 1163 (herb.
Field Mus. No. 608,106, type).
Peperomia trichopus Trelease, sp. nov.
Herba delicatissima epiphytica repens, caule filiformi glabro;
folia ad nodum vulgo 5 obovato-cuneata apice rotundata parva vix
5 mm. longa et 4 mm. lata 3-nervia, supra sparse adpresso-pubes-
centia, petiolis filiformibus in verticillo interdum inaequalibus 2-5
mm. longis; spicae terminales 30 mm. longae 1 mm. crassae laxi-
florae, pedunculo filiformi 3-4 cm. longo; bracteae rotundo-peltatae;
baccae rubro-brunneae oblongae curvae stipite brevi sed distincto
onustae subconice apiculatae, stigmate terminali.
Huanuco: Piedra Grande near Rio Santo Domingo, 1,500 meters,
Macbride 3855 (herb. Field Mus. No. 534,918, type).
Peperomia tricolor Trelease, sp. nov.
Herba majuscula repenti-assurgens epiphytica, caule 3 mm.
crasso laxe pallido-villoso; folia alterna subrhombeo-ovata obtuse
acutata basi acuta 4-5 cm. longa 2.5-3 cm. lata 5-nervia, supra
viridia nervis albis, subtus rubra nigro-granulosa glabra, petiolo
1 cm. longo crispo-pubescente; spicae terminales 100 mm. longae 2
mm. crassae, pedunculo 10 mm. longo crispo-pubescente; bracteae
rotundo-peltatae.
Junin: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
25973 (type in U. S. Nat. Herb.). San Nicolas to Azupizu, Killip &
Smith 26110.
Peperomia trinervis R. & P. Fl. 1: 32. pi 50. 1798; Miq. 111.
Piper, pi 8. Piper trinerve Vahl, Enum. 1: 342. 1805.
A more or less persistently crisp-pubescent, black-punctulate
herb; leaves alternate, ovate, acuminate, acute-based, 3-4 cm. long,
1.5-2 cm. wide, 3-nerved; petiole 5 mm. or in the lower leaves 15
mm. long; spikes terminal and from the upper axils, 100 mm. long,
2 mm. thick; peduncle 10 mm. long; bracts round-peltate; berries
subglobose, mucronate; stigma oblique. Neg. 8393.
Huanuco: Chinchao, Ruiz & Pavdn, type. Loreto: Without
locality, Weberbauer 4504- Puerto Arturo, Killip & Smith 2789.
Santa Rosa, Killip & Smith 28873. Yurimaguas, Killip & Smith
27690. San Martin: Tarapoto, Spruce 4310.
102 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia trullaefolia Trelease, sp. nov.
Herba majuscula laxe sparse ramosa, caule crassiusculo versus
basin 5 mm. crasso et ultra puberulo; folia ad nodum vulgo circa 6
oblonga profunde concava vel conduplicata parva 10 mm. longa 3
mm. lata lutescentia granulosa carnosa obtusa, nervis obscuris, gla-
brata, petiolo brevi 1 mm. longo velutino; spicae terminales ad apices
ramulorum brevium axillarium deciduo-foliatorum, 50 mm. longae
1 mm. crassae lutescentes laxiflorae, pedunculo vix 1 cm. longo
glabrato; bracteae rotundo-peltatae; baccae ovoideae mucronatae,
stigmate obliquo.
Cuzco: Ollantaitambo, 3,000 meters, Cook & Gilbert 401 (type in
U. S. Nat. Herb.).
Peperomia umbelliformis C. DC. Bot. Jahrb. 40: 257. 1908.
A rather small, umbellately branched, glabrous herb, the stem
slender; leaves alternate (?), lanceolate, acute at both ends (?), 14
mm. long, 6 mm. wide, 3-nerved; petiole suppressed; spike solitary
at the base of a leaf terminating each scaly floriferous branch, very
short and subsessile; bracts round-peltate; ovary oblong-ellipsoid,
with a style; stigma apical.
Lima: Monte Amancaes near Lima, on cliffs, 200-800 meters,
Weberbauer 1631, type; 1684.
Peperomia umbilicata R. & P. Fl. 1: 30. pi. 45. 1798; Hill,
Ann. Bot. 21: 150. 1907. Piper umbilicatum Vahl, Enum. 1: 356.
1805. Tildenia pemviana Miq. Nov. Act. Nat. Cur. 19: Suppl. 1:
483. 1843.
A small, glabrous, acaulescent herb; leaves from a corm rooting
from the sides as well as the bottom, round, sometimes subacute,
peltate, 10-20 mm. in diameter; petiole 4-6 cm. long; spikes 6-9
mm. long; peduncle elongate; bracts round-ovate; berries with small
stylopodium; stigma apical. Neg. 8395.
Type material from Lima, Chancai, and Huanuco, growing on
rocks, Ruiz & Pavdn. Without locality, Haenke.
Peperomia uncatispica Trelease, sp. nov.
Herba majuscula glabra, caule breviusculo crassiusculo, inter-
nodiis brevibus; folia alterna oblanceolata sensim acutata basi
cuneata 14-16 cm. longa 4-6 cm. lata ubique pinnatinervia, petiolo
2 cm. longo; spicae in paniculam simplicem ramulum lateralem
terminantem dispositae ad nodos paniculae solitariae uncatae graciles
70 mm. longae, pedunculo 5 mm. longo; bracteae rotundo-peltatae.
FLORA OF PERU 103
Junin: Schunke Hacienda above San Ramon, 1,400-1,700 meters,
Killip & Smith 24578 (type in U. S. Nat. Herb.).
Peperomia undeninervia C. DC. in DC. Prodr. 16, pt. 1: 441.
1869.
A glabrous, rather large, suberect herb; stem not sulcate; leaves
2, or upward 3, at a node, elliptic, subacuminate, 6-10 cm. long,
2.5-4 cm. wide, 11-nerved; spikes terminal and from the upper axils,
gently curved, 70-100 mm. long, 4 mm. thick; peduncle 10 mm. long;
bracts subovate-peltate; ovary ovoid; stigma subapical. Neg. 8533.
Type collected in Peru, Pavon. Junin: Colonia Perene, Killip
& Smith 25404.
As now understood, this is scarcely separable from P. septem-
nervis.
Peperomia vana Trelease, sp. nov.
Herba parva repens plus minusve crispo-pubescens, stolonibus
elongatis filiformibus foliis rotundatis alternis 5 mm. latis onustis,
P. rotundifoliae similis, ramulis floriferis brevibus erectis, foliis
ellipticis vel lanceolato-ellipticis 10-11 mm. longis 5 mm. latis;
spicae graciles 25 mm. longae, pedunculo 5 mm. longo; bracteae
rotundo-peltatae.
Junin: Schunke Hacienda above San Ramon, 1,400-1,700 meters,
Killip & Smith 24614 (type in U. S. Nat. Herb.), 24838.
Peperomia variculata Trelease, sp. nov.
Herba glabra repens epiphytica, caule gracili; folia alterna late
elliptica subabrupte obtuso-acuminata basi acuta 5-nervia, in sicco
fusca tenuia, supra nigro-puncticulata, petiolo gracili 5-10 mm.
longo; spicae terminales et ex axillis superioribus, juvenes 25 mm.
longae 1 mm. crassae.
Junin: Cahuapanas, 340 meters, Killip & Smith 26792 (type in
U. S. Nat. Herb.).
Peperomia variegata R. & P. Fl. 1: 33. pi. 52. 1798. Piper
variegatum Pers. Syn. 1: 35. 1805.
A stoloniferous, glabrous herb; leaves ovate, alternate, abruptly
subacuminate, peltate above the slightly retuse base, 12-15 cm.
long, 10 cm. wide, about 7-nerved, with the upper laterals above the
others; petiole 6-8 cm. long; spikes 1 or 2, on a terminal stalk 3 cm.
long, 150 mm. long, 5 mm. thick; peduncle 1.5-2 cm. long; bracts
round-peltate; ovary obliquely scutulate, with central stigma.
Huanuco: Chachahuassi, Ruiz & Pavon, type.
104 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia variifolia C. DC. in DC. Prodr. 16, pt. 1: 462. 1869.
A moderately small, crisp-tomentulose herb; stem 2 mm. thick,
striate; leaves characteristically 5 at a node, elliptic or suboblong,
obtuse, acute-based, 20 mm. long, 10 mm. wide, or the upper 10 mm.
long and 4 mm. wide, 3-nerved; petiole 1 mm. long; spikes from the
upper axils, scarcely 10 mm. long; peduncle very short; bracts round-
peltate; stigma oblique. Neg. 25254.
Type from Peru, Poeppig.
Peperomia ventricosicarpa Trelease, sp. nov.
Herba subparva erecta plus minusve ramosa glabra mollis aliquid
roseo-tincta, caule 2-3 mm. crasso; folia alterna ovata obtuse acu-
minata basi cordata 3.5 cm. longa 2 cm. lata 5-nervia, costa superne
2-ramosa, in sicco tenuia pellucida, petiolo 1-2 cm. longo; spicae
terminales graciles laxiflorae 40-50 mm. longae, pedunculo 1 cm.
longo; bracteae rotundo-peltatae; baccae substipitatae rotundo-
lageniformes, stigmate apicali.
Loreto: Soledad, Rio Itaya, 110 meters, Kittip & Smith 29800
(type in U. S. Nat. Herb.).
Peperomia verediana Trelease, sp. nov.
Herba mediocris assurgens glabra, caule 2-3 mm. crasso; folia
alterna ovata sensim subacuta basi rotundata et margine minute
cordulata, 5.5-6.5 cm. longa 3.5-4 cm. lata 5-nervia, petiolo 2 cm.
longo; inflorescentia non visa.
Loreto: Caballo-cocha, Llewelyn Williams 2139 (herb. Field Mus.
No. 608,592, type).
Peperomia verruculosa Dahlst. ex Hill, Ann. Bot. 20: 406.
pi. 29. 1906; 21: 149. 1907.
A small, glabrous, acaulescent herb closely resembling P. parvi-
folia but with more ovoid, verruculose berries.
Cuzco: Near Sacsaihuaman, Cuzco, 3,700 meters, Hill 182, type;
Herrera 2197a. Junin: Between Tarma and Oroya, 4,000 meters,
Weberbauer 2544- Lima: Rio Blanco, 4,500 meters, rocky summits,
2979. Lima-Oroya Railway, 4,400 meters, Weberbauer 305.
Peperomia Vidaliana Trelease, sp. nov.
Herba glabra roseo-tincta, caule gracili; folia alterna lanceolato-
elliptica utrinque acuta 5.5 cm. longa 1-2 cm. lata, inferiora reducta,
5-nervia, petiolo 5 mm. longo; spicae subterminales, juvenes 10 mm.
longae 1 mm. crassae, pedunculo brevi.
FLORA OF PERU 105
Type from Ecuador or Peru, Vidal-Senege in 1876-77 (type in
herb. Paris).
Peperomia villicaulis C. DC. Bot. Jahrb. 40: 264. 1908.
A rather small herb, densely villous throughout; leaves opposite,
ovate, obtuse at both ends, 22 mm. long, 16 mm. wide, 5-nerved,
the petiole very short; spikes axillary, 35 mm. long, 1 mm. thick;
peduncle 5 mm. long; bracts round-peltate; ovary obovoid; stigma
oblique.
Junin: Below Palca, on cliffs, 2,400-2,700 meters, Weberbauer
17U, type.
Peperomia villipetiola C. DC. in Lee. Notul. Syst. 3: 43. 1914.
A small, acaulescent herb with villous axes; leaves round-elliptic,
very obtuse, cordate, about 3 cm. long, 2.5 cm. wide, 7-nerved;
petiole 2 cm. long; spikes scarcely 5 mm. long, 1 mm. thick; panicled
at the top of a scape surpassing the leaves; bracts round-peltate.
Type from Ecuador or Peru, Vidal-Senege in 1876-77.
Peperomia viracochana Trelease, sp. nov.
Herba subparva caespitosa assurgens, caule graciusculo 1-2 mm.
crasso cinereo-puberulo; folia ad nodum 2-4 elliptica utrinque sub-
acuta parva 2-2.5 cm. longa 1.5 cm. lata in sicco subtenuia conspicue
venulosa 3-nervia, costa ramosa, utrinque subpersistente puberula,
petiolo vix 5 mm. longo plus minusve puberulo; spicae terminales
vel in verticillis axillaribus et foliis delapsis false paniculatae 30 mm.
longae 1 mm. crassae laxiflorae, pedunculo 1 cm. longo puberulo;
bracteae rotundo-peltatae; baccae rotundatae submucronatae, stig-
mate obliquo.
Cuzco: Temple of Viracocha near Tinta, 3,500 meters, Cook &
Gilbert 218 (type in U. S. Nat. Herb.).
Peperomia Weberbaueri C. DC. Bot. Jahrb. 40: 255. 1908.
A rather large, suberect, glabrous herb; stem 5 mm. thick; leaves
alternate, elliptic-oblong, subacute at both ends, 8-10 cm. long, 2-3
cm. wide, 7-nerved; petiole 2-4 cm. long; spikes 2-5 at the end of a
slender terminal stalk 6-8 cm. long and few-bracted at the top, 20-
100 mm. long, 2 mm. thick; peduncle 15-30 mm. long; bracts round-
peltate; ovary round, subsessile; stigma terminal.
Amazonas: Near Chachapoyas, on shaded cliffs, 1,600 meters,
banks of Rio Utcubamba, Weberbauer 4306, type.
106 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia yananoensis Trelease, sp. nov.
Herba parvula longivillosa subcaespitosa, caule subgracili 2-3
mm. crasso; folia alterna rotundata vel rotundo-ovata vel inferiora
latiora quam longa obtusa basi subacuta 1.5-3 cm. longa 1.5-2.5
cm. lata submultiplinervia, nervis basilaribus 1-2, nervo uno utroque
latere e medio nascente, utrinque villosa, petiolo 5-10 mm. longo
villoso; spicae axillares (folia fulcrantes saepe delapsa) 25 mm.
longae 1 mm. crassae sublaxiflorae glabrae, pedunculo brevi 5 mm.
longo glabrato; bracteae rotundo-peltatae.
Huanuco: Yanano, 1,800 meters, Macbride 3741 (herb. Field
Mus. No. 534,803, type).
Peperomia yananoensis var. caniana Trelease, var. nov.
Folia minora; spicae filiformes, pedunculo gracillimo.
Huanuco: Cani near Mito, 2,550 meters, Macbride 3460 (herb.
Field Mus., type).
Peperomia yapasana Trelease, sp. nov.
Herba majuscula epiphytica P. Schunkeanae similis, sed folia
non peltata, caule 5 mm. crasso et ultra brevi ter intertexto- villoso;
folia alterna ovata acuta basi rotundata 16-20 cm. longa 12-15 cm.
lata e 2 tertiis inferioribus submultiplinervia, in sicco tenuia brunnea,
pagina inferiore in vivo rubra breviter adpresso-subhirsuta vel sericea,
supra glabrescentia, petiolo 10-15 cm. longo molliter piloso; spicae
oppositifoliae 200 mm. longae 5 mm. crassae, pedunculo 5 cm. longo
piloso; bracteae rotundo-peltatae.
Junin: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25478 (type in U. S. Nat. Herb.).
2. POTHOMORPHE Miq.
Shrubs or large herbs; leaves large and broad, cordate at the
base, thin, often peftate; spikes very dense, long and slender, sub-
umbellate at the end of an axiTrary peduncle, the flowers sessile,
perfect; stamens 2 or 3. A small genus, but widely distributed in
tropical America.
Leaves not peltate P. Dombeyana.
Leaves peltate P. peltata.
Pothomorphe Dombeyana Miq. Syst. 211. 1843; 111. Piper.
pi. 26. Piper peltatum R. & P. non L. Piper umbellatum Auct. as
to Peru.
FLORA OF PERU 107
Plants suffrutescent; leaves round-reniform or openly cordate,
acuminate, becoming 20 cm. wide, somewhat velvety or crisp-
pubescent and ciliate, glandular-granular, multiple-nerved, with
about 14 basal or nearly basal nerves and one pair from somewhat
above the base, the petiole 6-10 cm. long, winged, somewhat hairy;
spikes umbellate on an axillary stalk 4 cm. long, 90 mm. long and
3 mm. thick or larger, the peduncle 1 cm. long.
Huanuco: Type from Cochero, Dcmbey, Chinchao, Ruiz &
Pavon 1-98. Amazonas: San Isidro, Tessmann 4999. Junin: Chan-
chamayo Valley, 1,500 meters, Schunke 1241, 1238. Loreto: San
Lorenzo, Killip & Smith 29205. Without locality: Ruiz; Haenke;
Poeppig 1389, 57; Mathews 1701.
Pothomorphe peltata (L.) Miq. Comm. Phyt. 37. 1839.
Piper peltatum L. Sp. PI. 30. 1753. Heckeria peltata Kunth, Linnaea
13: 565. 1839.
Scarcely different from the preceding except that the leaves are
peltate far above the base; it is found sometimes in much smaller-
leaved forms.
Loreto: Caballo-cocha, in forest, Williams 2103. Pebas, Williams
1650. Near Iquitos, Williams 1346, 8189. Mishuyacu, 100 meters,
Klug 1422, 295. La Victoria, Williams 3101, 2567. Pinto-cocha,
edge of forest, Williams 831. Paraiso, Alto Itaya, 145 meters,
Williams 3205. Santa Rosa, in clearing, 200 meters, Williams 4947.
San Martin: Rio Mayo, Tarapoto, 400-900 meters, in forest,
Williams 6243. Widely distributed in tropical America. "Santa
Maria."
3. PIPER L.
Shrubs or small trees, or the plants sometimes almost wholly
herbaceous, rarely scandent; leaves alternate, the stipules adnate
to the base of the petiole, or connate into one stipule opposite the
leaf, sometimes reduced to a dilation at the base of the petiole;
flowers densely crowded in the spike, subtended by peltate or rarely
concave bracts. A vast group, probably with a greater number of
species than any other genus of tropical American plants. Some
of the Old World species furnish such important products as black
pepper, but the American species have few economic applications,
and none of importance.
A. Leaves palmately nerved.
Leaves commonly about half as broad as long.
Leaves cordate at the base; petiole winged P. sororium.
108 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves not cordate at the base; petiole exceptionally winged.
Leaf base unguiculately inrolled ; leaves glabrate.
P. unguiculatum.
Leaf base not unguiculate.
Leaves elliptic.
Leaves glabrous P. Herrerae.
Leaves pubescent P. intonsum.
Leaves ovate; stigmas on a disk P. reticulatum.
Leaves more elongate.
Leaves elliptic P. heptandrum.
Leaves both lanceolate and elliptic P. Sellertianum.
Leaves lanceolate P. paralaevigatum.
A. Leaves pinnately or multiple nerved.
B. Midrib subequally branched throughout.
C. Leaves inequilateral and petiole winged.
Leaves about half as broad as long, obtuse at the base.
Leaves about 12 cm. long, not mottled, cordulate at the base.
P. tuber culat.um.
Leaves about 15 cm. long, not cordulate at the base.
Leaves brown-lepidote-mottled P. nanayanum.
Leaves not mottled P. excessum.
Leaves more elongate.
Leaf base obtuse, at least on one side.
Leaves 10-12 cm. long P. yurimaguasanum.
Leaves becoming 16 cm. long P. sericeonervosum.
Leaves distinctly larger.
Petiole suppressed on one side P. claudicans.
Petiole developed on both sides.
Leaves 20 cm. long and 8 cm. wide P. iquitosense.
Leaves 25 cm. long and 10 cm. wide P. pebasense.
Leaf base appreciably acute.
Leaves subelliptic.
Leaves 10-15 cm. long P. san-roqueanum.
Leaves becoming 20 cm. long P. fortalezanum.
Leaves narrowly lanceolate P. serpavillanum.
FLORA OF PERU 109
. Leaves equilateral or else petiole not winged.
D. Leaves scarcely over 10 cm. long.
Leaves about half as broad as long.
Leaves scarcely 3 cm. wide; petiole often winged.
P. phthinotrichon.
Leaves 4-5 cm. wide; petiole not winged P. daedalum.
Leaves more elongate.
Leaves rugulose, pubescent P. stupposum.
Leaves not rugulose, at most puberulent.
Peduncle 5 mm. long P. mediocre.
Peduncle 10 mm. long .P. recreoense.
D. Leaves commonly 10-15 cm. long.
E. Leaves about half as broad as long.
Stigmas sessile.
Berries angular P. asterocarpum.
Berries not angular.
Plants not glandular-granular P. multimammosum.
Plants glandular-granular P. heterophyllum.
Stigmas on a style.
Spikes scarcely 20 mm. long.
Leaves glabrous P. Peltobryon.
Leaves pilosulous P. brevispicatum.
Spikes about 30 mm. long.
Style stout P. dichotomum.
Style slender P. neglectum.
Spikes about 50 mm. long; style slender P. tenuistylum.
E. Leaves more elongate.
Leaves at most transiently or obscurely puberulent.
Stigmas on a style.
Spikes 5 mm. thick P. illautum.
Spikes slender P. stenostachys.
Stigmas sessile.
Spikes short and thick.
Berries wrinkled.
Leaves oblanceolate P. obovatum.
Leaves lanceolate. . P. rugosibaccum.
110 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Berries smooth.
Leaves scarcely 4 cm. wide.
Nerves impressed P. gilvibaccum.
Nerves not impressed.
Leaves glabrous P. oblongatilimbum.
Leaves somewhat puberulent. . .P. lanceolatum.
Leaves 4-5 cm. wide, sometimes not elongated.
P. sangorianum.
Leaves 5-6 cm. wide, drying black. .P. epunctatum.
Spikes moderately elongate.
Leaf base nearly equilateral.
Leaf base obscurely bicalloused ... P. huacachianum.
Leaf base not bicalloused.
Leaves 5-6 cm. wide.
Leaves not glandular-punctulate.
P. san-ramonense.
Leaves pale-punctulate P. dispansum.
Leaves black-punctulate P. Mocco-mocco.
Leaves scarcely 5 cm. wide.
Longer nerves 10 X 2 P. pernigricans.
Longer nerves more numerous. . . .P. curvatum.
Leaf base inequilateral.
Leaves black-punctulate P. Durvilleanum.
Leaves not glandular-punctulate P. expolitum.
Leaves more or less pubescent beneath only, the spikes short
and thick.
Twigs puberulent P. timbuchianum.
Twigs crisp-pubescent.
Nerves puberulent beneath P. sapoto-yacuense.
Nerves crisp-pubescent beneath.
Leaves lepidote-dotted P. perpurgatum.
Leaves not lepidote. P. ascendentispicum.
Twigs and nerves stiff- villous P. gratum.
Twigs and nerves slightly strigose P. rugosilimbum.
Leaves rough or pubescent on both sides.
Leaves granular-scabrous above P. Mathewsii.
FLORA OF PERU 111
Leaves crisp-pubescent.
Spikes slender.
Leaves soft-hairy . . _ P. calamistratum.
Leaves bearded P. setigerum.
Spikes 4 mm. thick P. satisfactum.
Spikes 7 mm. thick P. musteum.
D. Leaves becoming over 15 cm. long.
F. Leaves about half as broad as long.
Stigmas on a style.
Leaves not cordate at the base P. barbulantherum.
Leaves cordate at the base.
Petiole very short P. cordatum.
Petiole longer than the sinus P. magnificum.
Stigmas sessile.
Spikes short and thick.
Leaves subauricled-cordulate P. interitum.
Leaves scarcely cordulate P. asclepiadifolium.
Spikes elongate.
Leaves about 9 cm. wide, glabrous, black-dotted.
Spikes spreading P. heterophyllum.
Spikes ascending P. punctatum.
Leaves distinctly larger.
Leaves at most obscurely puberulent ... P. roqueanum.
Leaves more or less stellately pubescent.
Leaves shallowly cordate at the base.
P. asterotrichum.
Leaves obliquely subtruncate at the base.
P. tridentipilum.
F. Leaves more elongate.
Stigmas on a style.
Spikes short and thick.
Leaves cordulate at the base.
Leaves glabrous P. conispicum.
Leaves pubescent P. sapotoanum.
Leaves hirsute P. circumspectantis.
Leaves not cordulate at the base, hairy.
112 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves 4 cm. wide, narrowly lanceolate ... P. cirratum.
Leaves 6-8 cm. wide, lance-elliptic P. soledadense.
Spikes elongate but thick; leaf base acute; leaves glabrous.
Leaves oblanceolate-oblong P. costatum.
Leaves elliptic-subobovate.
Peduncle 1 cm. long; spikes 50 mm. long.
P. pseudo-munchanum.
Peduncle 2 cm. long; spikes longer P. longifolium.
Peduncle longer P. pseudo-curvatum.
Stigmas sessile.
G. Spikes elongated.
Leaves glabrous, fully 10 cm. wide.
Peduncle elongate P. casapiense.
Peduncle short and curved P. tocacheanum.
Leaves more or less pubescent.
Leaves about 10 cm. wide.
Nerves 12 on each side P. cahuapanasense.
Nerves 8 on each side P. marcescentispicum.
Leaves 6 cm. wide.
Nerves subvillous beneath P. indecorum.
Nerves hispid-hirsute P. san-nicolasense.
G. Spikes shorter and relatively thicker.
Leaves lance-oblong.
Leaves glabrous.
Leaf base acute.
Leaves 8 cm. wide P. maranyonense.
Leaves 5-6 cm. wide P. yessupianum.
Leaf base obtuse P. faecatum.
Leaves crisp-hairy beneath P. Klugianum.
Leaves soft-hirsute, rugulose P. dasyoura.
Leaves yellow- villous P. leticianum.
Leaves elliptic-subovate.
Leaves glabrous.
Spikes blunt P. bermudezense.
Spikes pointed.
FLORA OF PERU 113
Berries truncate P. itayanum.
Berries pointed P. ratisportanum.
Leaves pubescent beneath, about 10 cm. wide, not
cordate.
Pubescence crisp P. balsapuertanum.
Pubescence tomentose P. mishuyacuense.
G. Spikes comparatively slender.
Leaves glabrous: black-punctulate.
Peduncle 5 mm. long-; spikes cuspidate . . P. restrictum.
Peduncle twice as long.
Nerves straight P. fagijolium.
Nerves upcurved.
Leaves elliptic-ovate P. positum.
Leaves elliptic-oblong.
Nerves nearly horizontal P. phegophyllum.
Nerves ascending P. auristellanum.
Leaves more or less villous.
Leaves glabrous above; spikes 100 mm. long.
P. monzonense.
Leaves pubescent on both sides; spikes shorter.
Leaves 5 cm. wide P. Spruceanum.
Leaves 8-9 cm. wide P. affictum.
B. Midrib without strong ascending branches upcurved.
H. Leaves scarcely over 10 cm. long.
I. Leaves about half as broad as long.
Stigmas on a style; leaf base appreciably acute; leaves glabrate.
Leaves bicalloused at the base P. callosum.
Leaves not calloused at the base.
Leaves glabrous P. nasutum.
Leaves transiently puberulent P. tristigmum.
Stigmas sessile; leaf base appreciably obtuse.
Leaves glabrous above.
Leaves glabrous beneath.
Leaves scarcely cordate at the base . . P. pseudobarbatum.
Leaves cordate at the base.
114 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Spikes thick P. llatanum.
Spikes slender P. Carpunya.
Leaves pubescent beneath at least on the nerves, cordulate.
Spikes slender.
Leaves elliptic .P. monostigmum.
Leaves ovate P. Mohomoho.
Spikes 5 mm. thick P. stomachicum.
Leaves pubescent on both sides.
Plant scrambling.
Leaves scabrous P. cinereoramulurn.
Leaves not scabrous P. silvioberrans.
Plant not scrambling.
Leaves scarcely 5 cm. long P. redactum.
Leaves larger.
Twigs and leaves scabrous.
Petiole and peduncle short P. obesum.
Petiole and peduncle 10-15 mm. long P. Opizii.
Twigs gray-pubescent. P. leucojustum.
Twigs silky.
Leaves cordate at the base P. anonymum.
Leaves merely cordulate at the base . P. tenebricosum.
I. Leaves more elongate.
Leaves glabrous above between the nerves. .
Leaves glabrous beneath.
Spikes elongate P. comasense.
Spikes short P. arduum.
Leaves pubescent beneath, at least on the nerves.
Plant climbing P. adreptum.
Plant straggling P. strigillosicaule.
Plant neither climbing nor scrambling.
Leaves 1 cm. wide P. Dennisii.
Leaves 3-4 cm. wide.
Twigs glabrous P. Lehmannianum.
Twigs transiently puberulent P. delatum.
Leaves rough or pubescent on both sides.
Spikes short and thick.
FLORA OF PERU 115
Plant climbing P. acreanum.
Plant not climbing.
Petiole broadly winged P. ciniflonis.
Petiole not winged P. apurimacanum.
Spikes moderately elongate.
Twigs glabrous P. pontis.
Twigs somewhat scabrous P. leucophaeum.
Twigs appressed-pubescent.
Leaves not scabrous, gray P. posusanum.
Leaves scabrous.
Nerves appressed-pubescent beneath . . P. masiseanum.
Nerves hirsute beneath P. paucartamboanum.
Twigs crisp-hispid.
Twigs not granular P. brachytrichum.
Twigs pale-granular P. subflavicans.
Twigs shaggy-hirsute.
Leaves white-granular. P. margaritatum.
Leaves hairy-scabrous P. incomptum.
Leaves soft-pubescent P. stupposum.
Twigs soft-hirsute P. gracile.
H. Leaves commonly 10-15 cm. long.
J. Leaves about half as broad as long.
K. Leaf base appreciably acute.
Nerves not deeply impressed.
Leaves glabrous above between the nerves.
Leaves glabrous beneath.
Stigmas sessile.
Leaves hard and thick P. edurum.
Leaves thin.
Leaves granular, not venose.
Leaves 12 cm. long P. factum.
Leaves 15 cm. long P. heterophyllum.
Leaves venose P. nuncupatum.
Stigmas on a style.
Leaves bicalloused at the base P. callosum.
Leaves not calloused at the base P. Peltobryon.
116 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves puberulent beneath on the nerves .... P. resaturru
Leaves crisp-pubescent beneath on the nerves.
P. quimirianum.
Leaves strigose beneath on the nerves.
P. breveslrigillosum.
Leaves pubescent on both sides P. brevispicatum.
Nerves deeply impressed.
Nerves numerous.
Plant glabrous, or the petiole subpuberulent . P excultum.
Plant pubescent, at least locally P. lineatum.
Nerves few; leaves coriaceous P. carpapatanum.
K. Leaf base appreciably obtuse.
L. Leaves glabrous above.
M. Leaves glabrous beneath; stigmas sessile.
Spikes short and thick.
Leaf base rounded P. ecallosum.
Leaf base cordulate P. mikaniaefolium.
Spikes elongate.
Leaves cordate at the base P. cocherense.
Leaves cordulate at the base.
Berries round P. arrectispicum.
Berries triquetrous.
Leaves 6 cm. wide P. demoratum.
Leaves larger P. planipes.
Leaves neither cordate nor cordulate at the base.
Leaf base equilateral.
Twigs hollow P. excavatum.
Twigs solid.
Leaves 5-6 cm. wide.
Spikes rather thick.
Spikes 80 mm. long P. Churumayu.
Spikes 40 mm. long P. tumidicondyli.
Spikes slender P. laurinum.
Leaves broader P. Macbrideanum.
Leaf base shorter on one side.
FLORA OF PERU 117
Spikes moderate.
Plant not black-granular P. urubambanum.
Plant black-glandular P. conditum.
Spikes very thick P. sandianum*
M. Leaves pubescent beneath, at least on the nerves
Leaves bullulate-rugose; peduncle long. . . .P. areolatum*
Leaves coarsely subrugose.
Twigs glabrous.
Bracts yellow P. chromatolepis.
Bracts pale and inconspicuous .... P. achromatolepis.
Twigs puberulent P. santiaganum.
Twigs velvety, pale-granular P. paganicum.
Leaves not rugose.
Leaves cordulate at the base.
Twigs glabrous.
Nerves appressed-pubescent beneath . P. tabinense,
Nerves tomentulose P. coactaepilum.
Twigs transiently hispid P. huallaganum.
Leaves not cordulate at the base.
Leaf base equilateral.
Leaves elliptic P. albogranulatum.
Leaves ovate P. munyanum.
Leaf base shorter on one side. . . P. machupicchuense.
L. Leaves rough or pubescent on both sides.
Spikes short and thick; stigmas on a style . P. lagenaebaccum.
Spikes elongate; stigmas sessile.
Leaves not scabrous.
Leaves lanceolate, bullulate P. verruculaefdium.
Leaves broadly ovate.
Leaves rugose, villous P. suffruticosum.
Leaves not rugose.
Leaves not cordulate at the base.
P. huacapistanum.
Leaves cordulate or cordate at the base.
Leaves round-ovate P. lamasense.
Leaves elliptic P. pichisense.
118 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves scabrous.
Leaves becoming rugose.
Leaves obliquely cordulate at the base.
Internodes short P. rubiginosum.
Internodes long P. caballo-cochanum.
Leaves not cordulate at the base.
Spikes 80-100 mm. long.
Nerves subhirsute beneath; leaves elliptic.
P. malifolium.
Nerves short-pubescent .P. ucayalianum.
Spikes twice as long P. mundum.
Leaves scarcely rugose.
Leaf base equilateral.
Twigs velvety-pilose P. glabrius.
Twigs upcurved-hairy P. patens.
Leaf base oblique.
Leaves subvillous .P. pseudo-mollicomum.
Leaves closely pubescent, narrow-acuminate.
P. amnigaudens.
J. Leaves more elongate.
N. Leaf base appreciably acute.
Nerves deeply impressed P. subflavispicum.
Nerves not deeply impressed.
Stigmas on a style; spikes short; leaves narrow.
Leaves glabrous P. stenostachys.
Leaves somewhat pubescent P. Claudii.
Stigmas sessile.
O. Leaves glabrous above between the nerves.
P. Leaves glabrous beneath.
Spikes short.
Leaves scarcely punctulate; spikes cuspidate.
P. recensitum.
Leaves black-punctulate.
Leaves bicalloused at the base.
P. pseud o-callosum.
Leaves not bicalloused.
FLORA OF PERU 119
Spikes cuspidate P. arreptum.
Spikes not cuspidate.
Petiole very short P. reductipes.
Petiole 5-10 mm. long.
Nerves 4x2 P. morator.
Nerves 3x2 P. incitum.
Spikes elongated.
Spikes U-shaped P. Opizianum.
Spikes nearly straight.
Leaves narrowly lanceolate.
Leaves short-pointed ....... P. cumbasonum.
Leaves long-acuminate P. ostii.
Leaves lance-elliptic.
Petiole very short P. apus.
Petiole 5-10 mm. long.
Leaves 3-4 cm. wide P. temptum.
Leaves wider.
Leaves glandular-granular . P. gilvescens.
Leaves scarcely granular . P. veredianum.
P. Leaves pubescent beneath, at least on the nerves.
Spikes rather short.
Leaves narrowly lanceolate P. mansericheanum.
Leaves lance-elliptic P. monostigmum.
Spikes elongate; leaves lance-elliptic.
Leaves slightly rugose P. seminitidulum.
Leaves scarcely rugose.
Nerves puberulent beneath . P. chanchamayanum.
Nerves appressed-pubescent beneath.
Twigs granular.
Twigs crisp-pubescent P. portalense.
Twigs strigillose P. strigillosum.
Twigs glabrous P. pleninervosum.
Twigs not granular, glabrous P. subtextum.
Nerves crisp-pubescent beneath.
Leaves lance-ovate P. paraguassuanum.
Leaves lance-elliptic P. dendropse.
120 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Nerves subvillous beneath ; leaves lanceolate.
P. kimpitirikianum.
O. Leaves rough or pubescent on both sides.
Spikes strongly curved P. ollantaitambanum.
Spikes nearly straight.
Leaves scarcely scabrous.
Leaves lepidote; spikes pointed, 70 mm. long.
P. desultorium.
Leaves soft-hairy; spikes about 60 mm. long.
P. amphitrichum.
Leaves scabrous.
Leaves bullulate-lacunose.
Pubescence yellow P. crocatum.
Pubescence pale P. pseudo-matico.
Leaves rugose.
Leaves 4 cm. wide, lance-ovate, acute only.
P. ictum.
Leaves 5 cm. wide, lanceolate, acuminate.
P. occultum.
Leaves neither rugose nor bullulate.
Nerves appressed-pubescent beneath.
P. longegerminatum.
Nerves spreading-pubescent beneath.
Twigs gray-pubescent P. perenensc.
Twigs hispid.
Petiole 10 mm. long P. asperifolium.
Petiole half as long.
Leaves shorter on one side.
P. porveniricola .
Leaves equilateral.
Petiole hirsute P. subpatens.
Petiole scabrous P. pervulgatum.
Nerves subvillous beneath P. mercedense.
N. Leaf base appreciably obtuse.
Nerves deeply impressed.
Nerves 8 or 9 on each side P. lanceaefolium .
FLORA OF PERU 121
Nerves half as many; leaves lanceolate, long-attenuate.
P. callcanense.
Nerves not lineately impressed.
Leaves glabrous.
Spikes short; plant black-punctulate.
Twigs glabrous P. semicrndum.
Twigs hirtellous P. paludosum.
Spikes elongate P. oblongatifolium.
Leaves more or less pubescent beneath, at least upon the
nerves.
Nerves merely puberulent; leaves dimorphic.
P. rubribaccum.
Nerves appressed-pubescent beneath.
Twigs pale-granular.
Leaves broadly lanceolate.
Nerves 3-4 X 2 P. papillipetiolatum.
Nerves 5-6 X 2 P. indianonum.
Leaves lance-oblong P. Lehmannianum.
Twigs scarcely granular P. semicordulatum.
Twigs velvety P. fulvescenticaulc,
Twigs gray-pubescent P. griseocaule.
Nerves crisp-pubescent beneath; leaves acuminate.
P. aleyreanum.
Nerves velvety beneath; leaves bullulate.
P. santa-ritanum.
Nerves hirsute beneath P. dimetrale.
Leaves rough or pubescent on both sides.
Leaves scarcely scabrous.
Spikes curved.
Twigs glabrous, granular.
Spikes short and thick; stigmas on a style.
P. lagenaebaccum.
Spikes longer; stigmas not on a style.
P. granulosum.
Twigs gray-pubescent P. griseum.
Twigs velvety P. Kanehiranum.
122 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Twigs hirsute-villous.
Leaves lanceolate P. mellibractum.
Leaves lance-ovate P. velutinum.
Leaves lance-elliptic P. praeclarum.
Spikes nearly straight.
Twigs crisp-pubescent P. pseudo-flexuosum.
Twigs villous.
Peduncle short P. puniceum.
Peduncle 25 mm. long P. heterocarpum,
Leaves scabrous.
Spikes nearly straight.
Leaves not rugose.
Twigs subtomentose ........ P. longeacuminatum.
Twigs strigillose P. cispontinum.
Twigs hispid.
Leaves scabrous in spots P. scabiosifolium.
Leaves uniformly scabrous.
Leaves 4-5 cm. wide P. oculatispicum.
Leaves 6-7 cm. wide.
Midrib stiff -bristly .P. setulosum.
Midrib hirsute.
Peduncle slender P. rasile.
Peduncle short and thick P. celer.
Twigs subvillous.
Leaves 4-5 cm. wide P. lanosicaule.
Leaves 6-8 cm. wide.
Peduncle 10 mm. long P. Schunkeanum.
Peduncle 5 mm. long. .P. pseudo-mollicomum.
Leaves rugose P. Uleanum.
Spikes curved.
Leaves not rugose.
Plants climbing P. volubile.
Plants not climbing.
Twigs quickly glabrous.
Twigs pale-granular.
FLORA OF PERU 123
Leaves cordulate at the base.
P. camphoriferum.
Leaves not cordulate at the base.
P. glaberrimicaule.
Twigs not granular; leaves cordulate at the
base P. cocherense.
Twigs at first crisp-pubescent, granular.
P. subflavicans.
Twigs soft-hairy.
Twigs granular P. chalhuapuquianum.
Twigs not granular P. moyobambanum.
Leaves somewhat rugose, cordulate.
Leaves subvillous beneath P. mellibracteum.
Leaves strigillose beneath.
Twigs not scabrous P. granulosum.
Twigs transiently scabrous. . . P. asperrimicaule.
Leaves bullulate, cordulate P. elongatum.
H. Leaves characteristically becoming over 15 cm. long.
Q. Leaf base appreciably acute.
Stigmas on a style.
Leaves glabrous, glandular-punctulate, sharp-acuminate.
P. propinquum.
Leaves puberulent beneath on the nerves P. anastylum.
Leaves stellate-pubescent beneath P. jaculifolium.
Leaves crisp-sub villous; spikes large P. pubinervulum.
Stigmas sessile.
Leaves glabrous.
Leaves drying glossy-bronzed P. politaereum.
Leaves drying green.
Leaves black-glandular-punctulate P. peruvianum.
Leaves scarcely punctulate.
Petiole 10-15 mm. long.
Leaves subovate-elliptic P. nridilimbum.
Leaves elliptic P. duke.
Petiole scarcely 5 mm. long.
Spikes very long P. soricicaudatum.
124 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Spikes rather short.
Spikes curved P. reductipes.
Spikes nearly straight ..P. subsessile.
Leaves somewhat puberulent or pubescent beneath.
Leaves about half as broad as long P. huanucanum.
Leaves more elongate.
Twigs pale-granular P. contamanum.
Twigs scarcely granular.
Leaf base equilateral P. relaxatum.
Leaf base shorter on one side.
Nerves puberulent P. huantanum.
Nerves crisp-pubescent P. fuscispicum.
Nerves substrigose P. pullispicum.
Leaves rough or pubescent on both sides.
Spikes nearly straight.
Leaves scarcely rugose.
Twigs glabrescent and granular .... P. verruculaecaule.
Twigs sparsely subvillous P. offensum.
Twigs hirsute.
Spikes 4 mm. thick P. afflictum.
Spikes 2 mm. thick.
Leaves sparsely hairy beneath P. aegrum.
Leaves glabrous between the nerves.
P. leucophaeocaule.
Leaves becoming rugose P. paraisense.
Spikes large and curved.
Leaves scabrous P. dasyoura.
Leaves merely lepidote P. granuligerum.
Q. Leaf base appreciably obtuse.
R. Stigmas on a style.
Spikes short and thick.
Leaves about half as broad as long.
Leaves barely 10 cm. wide, cordate at the base.
Leaves glabrescent above.
Nodes, only, hairy P. moense.
Axes hairy P. xapurayense.
FLORA OF PERU 125
Leaves villous throughout P. dasypodum.
Leaves wider, subtruncate at the base.
P. constipatispicum.
Leaves more elongate.
Leaves cordate at the base P. villosispicum.
Leaves not cordate at the base P. tortuosivenosum.
Spikes elongate.
Leaves auricled, elongate.
Leaves glabrous : P. longestamineum.
Leaves puberulent P. longifilamentosum.
Leaves somewhat cordate.
Leaves glabrous.
Leaves 10-18 cm. wide P. Sumpi.
Leaves 10 cm. wide P. Endlicherianum.
Leaves stellately crisp-pubescent.
Spikes slender P. formosum.
Spikes stout P. mite.
Leaves subvillous.
Petiole 2 cm. long P. Pavoni.
Petiole short P. glabribaccum.
Leaves neither cordate nor auricled.
Leaves glabrous.
Twigs glabrous P. Hookerianum.
Twigs hirtellous.
Spikes 100 mm. long; peduncle slender.
P. ovantherum.
Spikes much longer; peduncle thick P. apodum.
Leaves pubescent beneath.
Twigs glabrous P. sciaphilum.
Twigs subpuberulent P. immite.
Twigs stellate-pubescent P. sidipilum.
Twigs crisp-pubescent.
Spikes 10 mm. thick P. crassamentum.
Spikes 5 mm. thick P. attenuatamentum.
Twigs subvillous.
Leaves about half as broad as long . . P. cassinoides.
126 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves more elongate.
Spikes scarcely 50 mm. long. . .P. transpontinum.
Spikes over 100 mm. long, 5 mm. thick.
P. resalutatum.
Twigs cobwebby.
Leaves stellate-pubescent beneath.
Spikes rather slender P. brevipedunculum.
Spikes 5-7 mm. thick P. ornithorhynchum.
Leaves villous beneath.
Leaves 10 cm. wide P. carrapanum.
Leaves broader P. araneatum.
Leaves pubescent on both sides.
Leaves rugose. . :- P. trichostylum.
Leaves not rugose P. trichostylopse.
R. Stigmas sessile.
S. Leaves about half as broad as long, or auricled if longer.
Leaves peltate, glabrous . P. subscutatum.
Leaves not peltate.
T. Leaves glabrous.
Leaves round-subtruncate at the base.
P. pauHanifolium.
Leaves cordulate at the base.
Leaves narrowed to the base. ... P. parvicordulatum.
Leaves round-based P. subcordulatum.
Leaves equilaterally cordate at the base . P. cocker ense.
Leaves very oblique at the base.
Leaves 16 cm. long; spikes slender P. Restio.
Leaves larger.
Leaves obliquely cordate at the base, paler
beneath P. azupizuanum.
Leaves auricled on one side.
Leaves pale beneath P. pallididorsum.
Leaves not pale beneath P. obliquum.
Leaves auricled, with lateral sinus.
Leaves round-obovate P. calvibaccum.
Leaves round-elliptic P. procerum.
FLORA OF PERU 127
T. Leaves puberulent or pubescent beneath, at least on
the nerves.
Leaves round-subtruncate at the base.
Nerves puberulent beneath.
Leaves 18 cm. long.
Leaves round-ovate P. abditum.
Leaves elliptic-ovate P. tingens.
Leaves lance-ovate P. yananoanum.
Leaves larger P. pseudo-churumayu.
Nerves crisp-pubescent beneath . . P. albopapillatum.
Nerves tomentulose beneath P. scapispicum.
Nerves villous beneath ; ...... P. parcepilosum.
Leaves truncate-subcordate at the base.
Leaves bullulate-alveolate.
Spikes sjyaight, 100 mm. long P. alveolatum.
Spikes curved, much longer P. perareolatum.
Leaves not bullulate P. euryphyllum.
Leaves equilaterally cordate at the base.
P. franciscanum.
Leaves inequilaterally shallow-cordate at the base.
P. riojanum.
Leaves very oblique at the base.
Leaves auricled, with basal sinus.
Petiole shorter on one side.
Twigs glabrous.
Nerves puberulent beneath.
Spikes constricted spirally . P. ligatispicum.
Spikes not constricted.
Leaves not pentagonal . . P. obovatifolium.
Leaves somewhat pentagonal, bronzed.
P. cupreatum.
Nerves velvety beneath.
Leaves round-ovate P. pubibaccum.
Leaves elongate-ovate . P. velutininervosum.
Twigs cobwebby P. acutulum.
Petiole equilong on both sides.
128 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Nerves merely puberulent P. obliquum.
Nerves subtomentose P. controversum.
Nerves villous beneath P. calvescentinerve.
Leaves auricled, with lateral sinus.
Nerves puberulent beneath.
Twigs glabrous.
Leaves elliptic P. celsum.
Leaves obovate P. sanguineispicum.
Twigs puberulent.
Leaves subobovate P. cinereonervosum.
Leaves elliptic P. capillistipes.
Nerves striate-hairy beneath .... P. striatinervosum.
Nerves crisp-pubescent beneath . P. ferruginispicum.
Nerves villous beneath.
Leaves broad P. confusionis.
Leaves narrow.
Leaves subrugose-pouched, subobtuse.
P. marsupiifemm.
Leaves not rugose, acute, lanceolate.
P. escaleranum.
T. Leaves rough or pubescent on both sides.
Leaves neither cordate nor auricled at the base.
Leaves inequilateral at the base.
Twigs scabrous P. scabricaule.
Twigs loosely subvillous P. loretoanum.
Twigs gray-hairy
Leaves not rugose P. canescenticaule.
Leaves becoming rugose P. serotinum.
Leaves equilateral at the base P. late-ovatum.
Leaves deeply cordate at the base.
Leaves obscurely rugulose P. indiciflexum.
Leaves bullulate P. acutifolium.
Leaves auricled at the base, red-villous.
Leaves elliptic-obovate P. pervillosum.
Leaves oblong-obovate P. strigosum. \
FLORA OF PERU 129
S. Leaves more elongate.
Leaves glabrous.
Leaves elliptic-oblong P. secundum.
Leaves lanceolate P. oblongatum.
Leaves pubescent beneath, at least on the nerves.
Leaves equilateral at the base.
Spikes long; leaves not cordulate at the base.
P. Brongniartii.
Spikes short; leaves cordulate at the base.
Spikes not cuspidate P. mitifolium.
Spikes cuspidate P. rhinostachyum.
Leaves inequilateral at the base.
Leaf base subacute.
Spikes 65 mm. long P. subandinum.
Spikes 100 mm. long P. ucayalianum.
Leaf base cordulate.
Leaves lance-oblong.
Peduncle elongate P. pothophyllum.
Peduncle short P. puberulescens.
Leaves lanceolate P. scalariforme.
Leaves oblanceolate P. tenuicuspe.
Leaf base cordate P. sagittifer.
Leaf base auricled P. escaleranum.
Leaves rough or pubescent on both sides.
Leaves inequilateral at the base.
Leaves neither cordate nor cordulate at the base.
Leaves rugose.
Petiole suppressed on one side .... P. immutatum.
Petiole evident P. diver sipilum.
Leaves not rugose.
Leaves soft-pubescent above. . . .P. chrysoneurum.
Leaves very granular-roughened.
Spikes curved P. scabridulicaule.
Spikes not strongly curved P. tardum.
Leaves cordulate at the base.
Spikes straight or slightly curved.
130 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves not rugose P. Lechlerianum.
Leaves rugose-bullulate P. verbascifolium.
Spikes strongly curved.
Leaves neither rugose nor bullulate.
P. elongatifolium.
Leaves somewhat rugose and scabrid.
Bracts inconspicuous P. Stuebelii.
Bracts conspicuous, pale P. Stevensi.
Leaves bullulate-lacunose P. elongatum.
Leaves equilaterally cordate at the base.
Spikes rather short P. ulceratum-
Spikes long and curved.
Leaves scarcely scabrous, elliptic, acuminate.
P. barbicuspe.
Leaves scabrous, lance-oblong, acuminate.
P. martinense.
Piper abditum Trelease, sp. nov.
Frutex 2-metralis nervis subtus griseo-puberulis exceptis glaber,
internodiis gracilibus modice elongatis; folia rotundato-ovata breviter
acuminata basi rotundata, uno latere paullo breviore, 15-18 cm.
longa 10-13 cm. lata e dimidio inferiore pinnatinervia, nervis 5x2,
petiolo 10-30+2 mm. longo, longioribus caduce alatis; spicae juve-
niles parvae graciles.
Loreto: Yurimaguas, 135 meters, Killip & Smith 27659 (type
in U. S. Nat. Herb.).
Piper achromatolepis Trelease, sp. nov.
Frutex nervis subtus puberulis exceptis glaber, internodiis flori-
feris brevibus crassis; folia late ovata breviter acuminata basi
aequilaterali obscure cordulata 14-16 cm. longa 9-10 cm. lata e 2
tertiis inferioribus pinnatinervia, nervis circa 6x2 ut venis longiori-
bus impressis, petiolo 1-2 cm. longo concavo fragiliter alato; spicae
120 mm. longae 3 mm. crassae; pedunculus 15 mm. longus 2 mm.
crassus; bracteae inconspicuae subpeltatae.
Loreto: Fortaleza, Yurimaguas, 155-210 meters, Williams 4244
(herb. Field Mus. No. 624,649, type). Iquitos, Killip & Smith
27254- "Cordoncillo."
FLORA OF PERU 131
Piper acreanum C. DC. Notizbl. Bot. Gart. Berlin 7: 486. 1917.
A climbing shrub, rooting from the nodes, somewhat crisp-
pubescent except for the upper side of the leaves; leaves elliptic,
subacuminate, subequilaterally acute at the base or distinctly
obtuse or even cordulate, 10-11 cm. long, 4-5 cm. wide, pinnately
nerved from the lower half, the nerves 4-5x2; petiole 10 mm.
long; spikes about 15 mm. long, 6 mm. thick, blunt; peduncle
slender, 10 mm. long; berries globose; stigmas 3, linear sessile.
Type from Seringal Auristella, Rio Acre, Brazil, Ule 9294.
Loreto(?): Rio Purus, Ule.
Piper acutifolium R. & P. Fl. 1: 38. pi 64- 1798. Steffensia
acutifolia Kunth, Linnaea 13: 650. 1839. Artanthe acutifolia Miq.
Syst. 428. 1844; 111. Piper, pi. 78.
A shrub; twigs yellow- villous; leaves lance-ovate, gradually
acute, inequilaterally cordate at the base with the longer side
auriculately cordate, bullulate, hairy on both sides and becoming
scabrous above from the bases of fallen hairs, 21-28 cm. long, 8-13
cm. wide, subpinnately nerved from the lower half, the nerves about
5+7; petiole barely 10 mm. long, hairy; spikes 3-4 mm. thick,
80 mm. long, straight; peduncle 15 mm. long, thinly hairy; bracts
rounded-subpeltate, honey-colored with a yellow ciliate border.
Huanuco: Cochero and Posuso, Ruiz & Pavon, type. Cordillera
valleys, Haenke 64.
Piper acutifolium var. Stuebelii Trelease, var. nov.
Folia late ovata basi fere aequilaterali cordata 25-26 cm. longa
15-19 cm. lata conspicue bullulata supra glabrescentia, subtus ut
petiolus crassus 4 cm. longus intertexto-villosa.
Amazonas: Near Chachapoyas, Stuebel 69 (herb. Berol., type),
21 (herb. Berol.).
Piper acutulum Trelease, sp. nov.
Arbor 6-8-metralis, internodiis floriferis modice gracilibus et
elongatis araneoso-villosis; folia elliptico-ovata acuta basi inaequi-
latera, latere longiore late auriculato et petiolo longiore, 33 cm.
longa 17 cm. lata e dimidio inferiore subpinnatinervia, nervis 5-6x2
superne puberulis, in sicco tenuia subtus lucida, petiolo 55+5 mm.
longo alato ut caule et basi costae subvilloso; spicae 400 mm. longae
5 mm. crassae, pedunculo 2 cm. longo plus minusve piloso, bracteae
rotundo-subpeltatae; baccae depresso-globosae glabrae; stigmata 3
filiformia sessilia.
132 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Junin: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
26014 (type in U. S. Nat. Herb.).
Piper adreptum Trelease, sp. nov.
Frutex gracilis scandens essentialiter glaber, internodiis elongatis;
folia lanceolata acuta vel anguste acuminata basi acuta 5.5-8 cm.
longa 1.5-3 cm. lata e dimidio inferiore pinnatinervia, nervis circa
5x2 inferne sensim approximatis, interdum subtus adpresso-sericeis,
petiolo 5 mm. longo exalato; spicae 100 mm. longae 5 mm. crassae
rectae mucronatae, pedunculo 3-5 mm. longo; bracteae rotundo-
subpeltatae ciliolatae; baccae breviter cylindraceae truncatae;
stigmata minuta sessilia.
Loreto: Mishuyacu, 100 meters, G. Klug 1394 (herb. Field Mus.
No. 627,627, type).
Piper aegruni Trelease, sp. nov.
Frutex 3-5-metralis, internodiis floriferis gracilibus subelongatis
hirsutis; folia elliptica acuminata basi fere aequilateraliter acuta,
18-20 cm. longa 7.5-8 cm. lata, e dimidio inferiore pinnatinervia,
nervis 5x2 supra minute scabro-pubescentibus, subtus ad nervos,
inter nervos sparsius, hirsuta, petiolo 10-15 mm. longo hirsute;
spicae juveniles 90 mm. longae 2 mm. crassae, pedunculo gracili
10 mm. longo plus minusve piloso.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26478
(type in U. S. Nat. Herb.).
Piper affictum Trelease, sp. nov.
Frutex 2-3-metralis ubique laxe longe crispo-pilosus, internodiis
floriferis breviusculis crassiusculis; folia elongata elliptica acuminata
basi inaequali-acuta 20-25 cm. longa 8-9 cm. lata e 2 tertiis inferi-
oribus pinnatinervia, nervis elevatis circa 6x2, petiolo 15-20 mm.
longo exalato; spicae 40 mm. longae 3 mm. crassae, pedunculo gracili
15 mm. longo; bracteae rotundo- vel triangulari-subpeltatae cilio-
latae; ovarium teres, stigmatibus 3 sessilibus.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28581
(type in U. S. Nat. Herb.), 28563.
Piper afflictum Trelease, sp. nov.
Frutex, ramis, petiolo atque nervis subtus subvilloso-hirsutis;
folia lanceolato-elliptica acuminata basi subaequilaterali acuta, 14-19 I
cm. longa 5-9 cm. lata e dimidio inferiore pinnatinervia, nervis
5-6x2 supra dense albido-scabris, petiolo 10 mm. longo; spicae 100-
FLORA OF PERU 133
130 mm. longae 3-4 mm. crassae, pedunculo 10 mm. longo; bracteae
rotundo-subpeltatae.
Loreto: Mainas, Poeppig in 1830 (type in herb. Vienna). La
Victoria, edge of forest, Williams 2699. Iquitos, Killip & Smith
27469, 27067. Balsapuerto, Killip & Smith 28426. San Antonio,
Killip & Smith 29293. "Cordoncillo."
Piper albogranulatum Trelease, sp. nov.
Frutex 2-metralis, internodiis subgracilibus et elongatis puberulis;
folia elliptico-suboblonga abrupte acuto-acuminata basin obtusum
versus angustata 11-14 cm. longa 5-7 cm. lata e 2 tertiis inferioribus
pinnatinervia, nervis 6x2, in sicco tenuia, subtus microscopice albo-
puncticulata et ad nervos puberula, petiolo 1 cm. longo puberulo
inferne alato; inflorescentiae non visae.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28883 (type in
U. S. Nat. Herb.).
Piper albopapillatum Trelease, sp. nov.
Frutex 3-metralis nervis subtus paullo crispo-puberulis exceptis
glaber, internodiis gracilibus elongatis in sicco fuscis et microscopice
albo-puncticulatis; folia ovata anguste acuminata basi obtusa,
margine altero latere abrupte subacuto et longiore, 20 cm. longa 10-12
cm. lata e dimidio inferiore vel magis inferne submultiplinervia,
nervis 5+6, petiolo 10+5 mm. longo vel in foliis inferioribus longiore
vaginante albo-puncticulato; spicae adhuc parvae et juveniles.
Junin: Schunke Hacienda, San Ramon, 1,400-1,700 meters,
Killip & Smith 24616 (type in U. S. Nat. Herb.).
Piper aleyreanum C. DC. Notizbl. Bot. Gart. Berlin 6:
488. 1917.
A shrub with subretrorsely soft-hairy twigs; leaves elliptic,
acuminate, slightly inequilaterally rounded at the base, 14-15 cm.
long, 6 cm. wide, pinnately nerved from the lower half, the nerves
about 4x2, thinly crisp-pubescent beneath with the nerves soft-
hairy; petiole about 5+2 mm. long; young spikes 30 mm. long, 2 mm.
thick, with a short peduncle; bracts triangular-subpeltate.
Brazil: Monte Aleyre, Seringal Sao Francisco, Alto Rio Acre,
Ule 9274, type. Doubtless occurring also in adjacent Peru.
Piper alveolatum Opiz in Presl, Rel. Haenk. 155. 1830. Artanthe
alveolata Miq. Syst. 406. 1844.
A soft-pubescent shrub; leaves ovate, acuminate, shallow-
cordate, 27 cm. long, 19 cm. wide, submultiple-nerved from the
134 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
lower half, the nerves about 6x2, bullulate-lacunose, nearly or quite
glabrous above; petiole 4-5 cm. long, pale-granular; spikes straight,
110 mm. long, 4 mm. thick; peduncle slender, 2 cm. long; bracts
yellow-puberulent.
Huanuco: Huanuco mountains, Haenke, type. Cuzco: Rio
Pillahuata, in forest, 2,400 meters, Pennell 14014-
Piper amnigaudens Trelease, sp. nov.
Frutex arborescens 4-metralis spec. P. pseudo-mollicomum similis
nodosus, internodiis supremis brevibus subgracilibus dense hispido-
pubescentibus serius scabridis; folia lanceolato-elliptica anguste
acuminata basi rotundata, uno latere plus minusve breviore, 12-15
cm. longa 4.5-6.5 cm. lata e dimidio inferiore pinnatinervia, nervis
circa 6x2, supra minute granuloso-scabra, nervis subtus minute
excurvo-pubescentibus, petiolo circa 5+3 mm. longo; spicae 100 mm.
longae 2-3 mm. crassae, pedunculo gracili 10 mm. longo subscabrido;
bracteae rotundo-subpeltatae ciliatae; baccae rotundatae, stigma-
tibus 3 gracilibus sessilibus.
Loreto: Steep bank of Rio Santiago, 200 meters, Ynes Mexia
6332 (type in herb. Univ. Calif.).
Piper amphitrichum Trelease, sp. nov.
Frutex 3-metralis nodosus, internodiis floriferis brevibus modice^
crassis crispo-hirsutis; folia lanceolato-elliptica vel subrhombea acute
acuminata basi inaequilaterali acuta, 10-14 cm. longa 3.5-5 cm.
lata e 2 tertiis inferioribus pinnatinervia, nervis circa 6x2 utrinque
rigide adpresso-pilosis, nervis subtus excurvo-hirsutis, petiolo 3+3
mm. longo exalato hirsute; spicae 60 mm. longae 3 mm. crassae,
pedunculo 10 mm. longo plus minusve crispo-piloso; bracteae sub-
triangulari-subpeltatae.
Junin: Colonia Perene", 680 meters, Killip & Smith 24938 (type
in U. S. Nat. Herb.).
Piper anastylum Trelease, sp. nov.
Frutex vix 2-metralis, ramis gracilibus plus minusve geniculatis,
petiolo et nervis subtus obscure puberulis exceptis glaber; folia
variabilia lanceolata vel lanceolato-ovata usque ad late ovata
anguste acuminata basi inaequilaterali acuta, 12-15 cm. longa 5-7
cm. lata e 3 quartis inferioribus pinnatinervia, nervis 7x2, in sicco
tenuia et plus minusve lucida, petiolo 10+3 mm. longo exalato;
spicae 30-40 mm. longae 3 mm. crassae, pedunculo adscendente 5
FLORA OF PERU 135
mm. longo; bracteae crescentico-subpeltatae; baccae globosae,
stylo gracili, stigmatibus 3 patentibus.
Junin: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
26015 (type in U. S. Nat. Herb.).
Piper anonymum Trelease, sp. nov.
Frutex(?), ramis longisericeis; folia ovata subacuta basi cordata
9 cm. longa 4 cm. lata e 2 tertiis inferioribus pinnatinervia, nervis
circa 5x2, supra basibus pilis longis persistentibus scabra, subtus
adpresso-sericea, petiolo 5-10 mm. longo sericeo; spicae 80 mm.
longae 3 mm. crassae acuminatae, pedunculo 5-10 mm. longo
griseo-hispido.
"Peruvia subandina," Poeppig 73 (herb. Paris, type).
Piper apodum Trelease, sp. nov.
Arbor gracilis 5-metralis nodosa, internodiis floriferis elongatis
crassiusculis sparsissime vel ad nodos densius hirtellis; folia ovato-
lanceolata vel late ovata sensim acuminata basi subaequaliter
rotundata, magna 16-35 cm. longa 8-23 cm. lata, e tertio infimo
submultiplinervia, nervis 5-7x2, glabra, petiolo 2-5 cm. longo plus
minusve hirtello anguste alato; spicae oppositifoliae rectae magnae
200 mm. longae et 8 mm. crassae vel majores, pedunculo crassissimo
brevi 8 mm. longo 5 mm. crasso velutino; bracteae rotundo-sub-
peltatae ferrugineo-ciliatae, floribus sessilibus perfectis; stigmata 3
recurva essentialiter sessilia.
Huanuco: Muna, 2,100 meters, Macbride 3987 (herb. Field
Mus. No. 535,049, type).
Piper apurimacanum Trelease, sp. nov.
Frutex 1.5 m. altus gracilis nodosus, internodiis floriferis brevibus
gracilibus pilis arete adpressis canescentibus; folia subrhombeo-
lanceolata vel lanceolato-elliptica anguste falcato-acuminata basi
inaequilaterali acuta 9-10 cm. longa 2.5-4 cm. lata e dimidio in-
feriore pinnatinervia, nervis circa 5x2, supra minute adpresso-
pubescentia vel subscabrida in statu adulto glabrata et opaca,
nervis subtus breviter adpresso-pubescentibus, petiolo vix 2+2
mm. longo ut caule pubescente; spicae curvae 30 mm. longae 3
mm. crassae, pedunculo filiformi 10 mm. longo ut caule pubescente;
bracteae subtriangulari-subpeltatae albae; baccae subglobosae, stig-
matibus 3 minutis sessilibus.
Ayacucho: Rio Apurimac near Kimpitiriki, 400 meters, Killip
& Smith 22926 (type in U. S. Nat. Herb.).
136 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper apus Trelease, sp. nov.
Frutex 3-4-metralis nodosus glaber, internodiis floriferis brevibus
gracilibus; folia lanceolata longiacuminata basi inaequilaterali acuta
15 cm. longa 5-6 cm. lata e dimidio inferiore pinnatinervia, nervis
5+6, in sicco tenuia subtus viridia pallidiora, petiolo latere longiore
nullo altero 5 mm. longo, lamina decurrente, non vaginante; spicae
juveniles 20 mm. longae 2 mm. crassae, pedunculo 5 mm. longo;
bracteae rotundo-subpeltatae.
Junin: Santa Rosa, Pichis Trail, 625-900 meters, Killip &
Smith 26192 (type in U. S. Nat. Herb.).
Piper araneatum Trelease, sp. nov.
Arbor 4-5-metralis, internodiis floriferis elongatis gracillimis
laxe subaraneoso-villosis; folia ovata acute attenuata basi rotundata
abrupte ad petiolum contracta, 18-26 cm. longa 8-15 cm. lata e
dimidio inferiore submultiplinervia, nervis 5-6x2, supra albo-
puncticulata, subtus sparse stellato-pubescentia, nervis subhirsutis,
petiolo 10-20+2 mm. longo vaginante pallido-villoso ; spicae in
anthesi 80 mm. longae 5 mm. crassae, pedunculo gracili breviter
villoso 15 mm. longo recurvo; bracteae rotundo-subpeltatae pilosae.
Junin: Colonia Perene 1 , 680 meters, Killip & Smith 25242 (type
in U. S. Nat. Herb.).
Piper arduum Trelease, sp. nov.
Frutex gracilis glaber obscure nigro-puncticulatus, internodiis
floriferis gracilibus breviusculis; folia sublate lanceolata sensim acute
subacuminata basi acuta e dimidio inferiore pinnatinervia, nervis
superioribus cum costa venis circa 3 obscuris reticulatis transversis
connexis, petiolo gracili 10 mm. longo exalato; spicae 12 mm. longae
4 mm. crassae obtusae, pedunculo 5 mm. longo recurvo; bracteae
rotundo-subpeltatae anguste marginatae; baccae globosae mucro-
natae, stigmatibus parvis sessilibus.
Loreto: Puerto Arturo, Yurimaguas, 155-200 meters, in forest,
Llewelyn Williams 5174 (herb. Field Mus. No. 618,063, type).
Piper areolatum (Miq.) C. DC. in DC. Prodr. 16, pt. 1: 303.
1869. Artanthe areolata Miq. Lond. Journ. Bot. 4: 454. 1866.
A shrub, loosely villous except for the glabrous upper leaf surface;
leaves ovate, subacuminate, obtuse at the base, 14 cm. long, 7.5
cm. wide, pinnately nerved from the lower half, the nerves about
FLORA OF PERU 137
5x2, bullulate; petiole 2-3 cm. long, winged; spikes flexuous,
125 mm. long, 4 mm. thick, slender; peduncle slender, 5 cm. long.
Neg. 8401.
Amazonas: Sierra de Chachapoyas, Mathews 3227, type.
Piper arrectispicum Trelease, sp. nov.
Frutex nodosus glaber, internodiis floriferis modice gracilibus
brevibus in sicco luteis; folia elliptica acuminata basi acuta, in-
feriora ovata et cordulata, 13 cm. longa 6-7 cm. lata e tertio in-
feriore submultiplinervia, nervis 4-5x2, coriacea revoluta subtus
pallidiora, petiolo 6 mm. longo exalato vel in foliis inferioribus
duplo longiore et alato; spicae fructiferae 65 mm. longae 3-4 mm.
crassae propter pedunculum 7 mm. longum primo curvum serius
rectum suberectae; bracteae crescenticae; baccae globosae vel sub-
truncatae, stigmatibus 3 minutis sessilibus.
San Martin: San Roque, in forest, 1,350-1,500 meters, Llewelyn
Williams 6993 (herb. Field Mus. No. 629,617, type). "Cordoncillo."
Piper arreptum Trelease, sp. nov.
Arbor 3-5-metralis gracilis glaber, internodiis floriferis brevibus
gracilibus; folia lanceolato-oblonga vel elliptico-oblonga acute longi-
acuminata basi inaequilaterali acuta 12-15 cm. longa 4-4.5 cm.
lata e dimidio inferiore pinnatinervia, nervis 5x2, in sicco tenuia
viridia, petiolo 5-10 mm. longo exalato; spicae 30 mm. longae 4
mm. crassae caudatae, pedunculo 5 mm. longo; bracteae triangulari-
subpeltatae ciliolatae; baccae rubro-brunneae globosae obscure
mucronatae, stigmatibus 3 sessilibus.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26487
(type in U. S. Nat. Herb.).
Piper ascendentispicum Trelease, sp. nov.
Frutex 1.5 m. altus, internodiis floriferis brevibus gracilibus
evanescente paullo crispo-pubescentibus; folia elliptica vel oblonga
abrupte acuminata basi subaequilaterali acuta vel in foliis latioribus
obtusa 8-12 cm. longa 4-6 cm. lata ubique pinnatinervia, nervis
longioribus 6X2, subtus ad nervos sparse crispo-pubescentia, petiolo
5-10 mm. longo; spicae floriferae 12 mm. longae 4 mm. crassae non
cuspidatae, pedunculo adscendente 4 mm. longo 2 mm. crasso;
bracteae crescenticae.
Junin: Cahuapanas, 340 meters, Killip & Smith 26724 (type in
U. S. Nat. Herb.).
138 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper asclepiadifolium Trelease, sp. nov.
Frutex 2-3-metralis glabratus, internodiis floriferis brevibus
fistulosis sordido-puberulis; folia late elliptica abrupte breviacumi-
nata basi fere aequilaterali rotundata 15-17 cm. longa 8-9 cm.
lata ubique pinnatinervia, nervis 8-10x2, subtus ad nervos cinereo-
puberula et inter nervos obscure pallido-granulosa, petiolo 5 mm.
longo exalato; inflorescentia non visa.
Loreto: Iquitos, Killip & Smith 27016 (U. S. Nat. Herb.).
Piper asperifolium R. & P. Fl. 1: 37. pi. 56. 1798. Steffensia
asperifolia Kunth, Linnaea 13: 644. 1839. Artanthe asperifolia Miq.
Syst. 441. 1844.
A hispid or crisp-pubescent shrub; leaves elliptic or sublan-
ceolate, acuminate, subacute at the base, 9-14 cm. long, 4-6 cm.
wide, pinnately nerved from the lower half, the nerves about 5+2,
closely short-scabrous above, paler and dark-punctulate beneath
with spreading-hairy nerves; petiole 10 mm. long; spikes 80 mm.
long, 3 mm. thick; peduncle 10-15 mm. long; bracts transversely
subpeltate, ciliate. Neg. 8402.
Huanuco: Type from Pillao (Ruiz in Herb. Willdenow, No.
674, and No. 16 in general herbarium, at Berlin; Pawn, at Kew).
Piper asperrimicaule Trelease, sp. nov.
Frutex 3-metralis nodosus, internodiis floriferis brevibus gracili-
bus primo scabro-pubescentibus serius glabris et pallido-granulosis;
folia lanceolata subacuminata attenuata basi inaequilaterali cor-
dulata 10-12 cm. longa 4 cm. lata e dimidio inferiore pinnatinervia,
nervis 4+5, subrugosa, supra albo-granuloso-scabra, subtus ad
nervos strigillosa, petiolo 5 mm. longo scabrido; spicae arcuatae
80 mm. longae 3 mm. crassae obtusae; pedunculo 5 mm. longo
punctulato; bracteae rotundo-subpeltatae.
Loreto: Yurimaguas, 135 meters, Killip & Smith 27955 (type
in U. S. Nat. Herb.).
Piper asterocarpum Trelease, sp. nov.
Frutex (?) glaber non glanduloso-punctulatus, ramulis gracilibus;
folia lanceolato-ovata subacuminata basi acuta 12 cm. longa 5 cm.
lata fere ubique pinnatinervia in sicco tenuia, petiolo 10 mm. longo
exalato; spicae 90 mm. longae 4-5 mm. crassae, pedunculo 10 mm.
longo; baccae depresso-globosae profunde 4-sulcatae submucronatae,
stigmatibus parvis in mucrone insertis.
Type without data, from Peru (?), in Herb. Endlicher at Vienna.
FLORA OF PERU 139
Piper asterotrichum C. DC. Verb. Bot. Ver. Brandenb. 47:
111. 1905.
A glandular-granular shrub, tufted-pubescent except for the
upper leaf surface; leaves 22-25 cm. long, 11-15 cm. wide; abruptly
short-acuminate, nearly equilaterally shallow-cordate; petiole 2-3
cm. long; spikes becoming 50 mm. long, 10 mm. thick; peduncle
scarcely 10 mm. long; stigmas sessile.
Loreto: Iquitos, Ule 6247, type.
Piper asterotrichum var. yessupense Trelease, var. nov.
Arbor 4-6-metralis sparse transiente barbato-pubescens; folia
lata ovata vix cordata 24 cm. longa 16 cm. lata; spicae juveniles
20 mm. longae 4 mm. crassae luteae, pedunculo 10 mm. longo.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26271 (type
in U. S. Nat. Herb.).
Piper attenuatamentum Trelease, sp. nov.
Arbor 4-5-metralis, internodiis floriferis brevibus gracilibus
sparse transiente crispo-pilosis; folia ovata anguste acuminata basi
rotundata 26 cm. longa 13 cm. lata e dimidio inferiore submulti-
plinervia, nervis 5-6x2 subtus crispo-pilosis, petiolo 2 cm. longo
alato laxe crispo-piloso; spicae fructiferae 100-120 mm. longae 5
mm. crassae, pedunculo compresso 7 mm. longo subpiloso; bracteae
crescentico-subpeltatae; baccae globosae, stylo gracili, stigmatibus
3 recurvis.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26513
(type in U. S. Nat. Herb.).
Piper auristellanum C. DC. Notizbl. Bot. Gart. Berlin, 7:
488. 1917. Piper nigropunctatum var. franciscoanum C. DC. loc. cit.
A glabrous, very black-granular shrub; leaves elongate-elliptic,
subacuminate, slightly inequilaterally acute at the base, 20 cm.
long, 7 cm. wide, pinnately nerved nearly throughout; petiole
10+5 mm. long; young spikes 20 mm. long, 2 mm. thick; peduncle
10 mm. long.
Brazil: Seringal Auristella, Rio Acre, Ule 9276, type. Probably
extending into adjacent Peru.
Piper azupizuanum Trelease, sp. nov.
Arbor 5-6-metralis glabra, internodiis modice elongatis; folia
obliqua ovata subacuta basi inaequilaterali cordata sinu aperto 27
cm. longa 14 cm. lata e 3 quartis inferioribus subpinnatinervia,
140 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
nervis 7x2, in sicco subcoriacea opaca, petiolo 6.5+1.5 cm. longo
anguste alato; spicae 350 mm. longae 5 mm. crassae, pedunculo
gracili 3 cm. longo; bracteae rotundo-subpeltatae brunneo-ciliatae;
baccae paullo cum rhachi elongatae, stigmatibus 3 filiformibus
sessilibus.
Junin: San Nicolas to Azupizu, Pichis Trail, 650-900 meters,
Killip & Smith 26094 (type in U. S. Nat. Herb.).
Piper balsapuertanum Trelease, sp. nov.
Frutex 2-3-metralis, internodiis floriferis modice brevibus sed
crassis glabratis; folia elliptico-oblonga subacuminata basi acuta,
altero latere breviore, 24-27 cm. longa 11-12 cm. lata ubique pin-
natinervia, nervis circa 15x2, subtus praesertim ad nervos sparse
crispo-pubescentia, petiolo 10+10 mm. longo exalato; spicae juve-
niles 85 mm. longae 6 mm. crassae mucronatae, pedunculo 20 mm.
longo 2 mm. crasso stellato-pubescente; bracteae magnae triangulari-
subpeltatae ciliatae flavidae; ovarium paullo acutatum, stigmatibus
3 sessilibus.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28394
(type in U. S. Nat. Herb.).
Piper barbicuspe Trelease, sp. nov.
Arbor 3.5 m. alta nodosa in sicco fuscescens, internodiis floriferis
breviusculis crassiusculis tomentulosis; folia elliptica acuminata,
acumine in cuspem barbatum 50 mm. longum 1 mm. latum pro-
tracto, basi inaequilaterali breviter cordata 27 cm. longa 9.5 cm.
lata e dimidio inferiore pinnatinervia, nervis 7-8x2, in sicco tenuia,
supra ad nervos basinque sparse pilosa, in statu adulto ut videtur
subtus magis persistente pilosa, petiolo crasso vix 1 cm. longo;
spicae U-formes vix 150 mm. longae et 3 mm. crassae, apice barbatae,
pedunculo 10-15 mm. longo velutino; bracteae triangulari-sub-
peltatae ciliatae; stigmata 3 digitata subsessilia.
Loreto: Mishuyacu, 100 meters, in clearing, G. King 253 (herb.
Field Mus. No. 629,989, type.)
Piper barbulantherum Trelease, sp. nov.
Frutex, ramis transiente crispo-villosis; folia elliptica abrupte
subacuminata basi acuta 17 cm. longa 9 cm. lata ubique pinnati-
nervia glanduloso-granulosa, nervis subtus subadpresso-pilosis, peti-
olo 2 cm. longo crispo-pubescente; spicae fructiferae 150 mm. longae
15 mm. crassae leviter curvae, pedunculo 15 mm. longo 3 mm.
FLORA OF PERU 141
crasso curvo crispo-pubescente; baccae obovoideae, stylo crasso,
stigmatibus magnis latis patentibus.
Piura: Cordillera east of Huancabamba, 3,000 meters, Weber-
bauer 6093 (herb. Berol., type).
Piper bermudezense Trelease, sp. nov.
Frutex 2-3-metralis glanduloso-granulosa, ramis foliisque glabris,
internodiis floriferis breviusculis gracilibus; folia elliptico-subovata
acuminata basi inaequilaterali saltern uno latere subacuta, 27 cm.
longa 11 cm. lata ubique pinnatinervia, nervis circa 15x2, petiolo
5+5 mm. longo exalato; spicae arcuatae obtusae 50 mm. longae
10 mm. latae, pedunculo 15 mm. longo 2 mm. crasso; bracteae
triangulari-subpeltatae ciliatae; baccae tetragonae fulvo-velutinae,
stigmatibus 3 sessilibus.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26424
(type in U. S. Nat. Herb.), 26674-
Piper brachytrichum Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis subgracilibus elongatis
pallide subcrispo-hispidis; folia ovato-lanceolata acute acuminata
basi inaequilaterali bicordulata 10 cm. longa 4 cm. lata supra pilulis
brevibus scaberrima serius lepidota, subtus pallidiora nervis molliter
pubescentibus, petiolo 5+2 mm. longo exalato breviter hirsute;
spicae juveniles 60 mm. longae 3 mm. crassae, pedunculo gracili
10 mm. longo piloso.
Ayacucho: Estrella, 500 meters, Killip & Smith 22652 (type in
U. S. Nat. Herb.).
Piper brevestrigillosum Trelease, sp. nov.
Frutex (fistulosus?) nodosus 2-3-metralis, internodiis floriferis
subgracilibus modice elongatis glabris; folia elliptica vel sublanceolata
acuminata basi subaequilaterali acuta 12-16 cm. longa 5.5-8 cm.
lata e dimidio inferiore pinnatinervia, nervis circa 6x2, supra
granuloso-asperata, subtus pallidiora nervis breviter patenti-strigosis,
petiolo 10 mm. longo infra medium vaginante; spicae (normales?)
curvae 75 mm. longae et ultra 3 mm. crassae, pedunculo 5 mm.
longo glabrato; bracteae rotundo-subpeltatae.
Ayacucho: Carrapa, 1,500 meters, Killip & Smith 22375 (type
in U. S. Nat. Herb.).
Piper brevipedunculum Trelease, sp. nov.
Arbor 3-5-metralis, trunco 5 cm. diam., internodiis floriferis
gracilibus brevibus plus minusve araneosis; folia ovata acuminata
142 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
ad petiolum abrupte contracta, 15-24 cm. longa 9-15 cm. lata e
dimidio inferiore submultiplinervia, nervis 5-7x2, subtus praesertim
ad nervos stellate- velutina, petiolo 2 cm. longo vaginante velutino;
spicae floriferae 80 mm. longae 4 mm. crassae, pedunculo vix 5 mm.
longo crasso velutino; bracteae rotundo-subpeltatae ciliolatae; ova-
rium teres, stylo brevi, stigmatibus 3 recurvis.
Ayacucho: Estrella, 500 meters, Kittip & Smith 22677 (type in
U. S. Nat. Herb.).
Piper brevispicatum Opiz in Presl, Rel. Haenk. 151.
pi. 28. 1830.
A shrub, glabrous except that the leaves are minutely pilose on
both faces; leaves subrhombic-elliptic, equally subacuminate-pointed
at both ends, 12 cm. long, 5.5 cm. wide, pinnately nerved nearly
throughout; petiole 10 mm. long, the blade decurrent upon it; young
spikes scarcely 5 mm. long, 1 mm. thick, short-stalked.
Type locality unknown.
A very doubtful species, apparently of the Peltobryon group but
unique in its velvety foliage.
Piper Brongniartii (Miq.) C. DC. in DC. Prodr. 16, pt. 1:
327. 1869. Artanthe Brongniartii Miq. Syst. 486. 1844.
A shrub, glabrous except for the puberulent nerves beneath;
leaves elliptic, acuminate, almost equilaterally obtuse at the base,
17-22 cm. long, 7-15 cm. wide, pinnately nerved from the lower
half, the nerves about 5+2, glossy; petiole scarcely 10 mm. long;
spikes 100 mm. long, 2-3 mm. thick, flexuous; peduncle 10 mm. long.
Huanuco; Cochero, Dombey 927 or 907, type.
Piper caballo-cochanum Trelease, sp. nov.
Frutex, internodiis floriferis gracilibus elongatis hispidis; folia
elliptico-ovata anguste acuminata basi inaequilaterali subcordulata
14-16 cm. longa 6-7.5 cm. lata e dimidio inferiore pinnatinervia,
nervis 6x2, in statu adulto plus minusve rugosa, supra dense granu-
loso-scabra, subtus punctulata nervis incurvo-hispidis, petiolo 10+2
mm. longo; spicae 100 mm. longae 2 mm. crassae, pedunculo 10 mm.
longo hispido.
Loreto: Caballo-cocha, in forest, Llewelyn Williams 2309 (herb.
Field Mus. No. 608,282, type). "Cordoncillo."
Piper cahuapanasense Trelease, sp. nov.
Frutex 2-4-metralis nodosus, internodiis floriferis breviusculis
subelongatis sparse persistente crispo-barbatis ; folia sublanceolato-
FLORA OF PERU 143
elliptica acute acuminata basi fere aequilaterali acuta 20-25 cm.
longa 8-9 cm. lata ubique pinnatinervia, nervis circa 12x2, supra
ad costam paullo longihirsuta, subtus nigro-punctulata ad nervos
longihirsuta, petiolo 2 cm. longo ut caule costaque barbato exalato;
spicae 100 mm. longae 2 mm. crassae, pedunculo 15 mm. longo sparse
barbato; bracteae rotundo-subpeltatae; baccae subglobosae interdum
ad apicem depressae mucronatae; stigmata 3 brevia in mucrone
inserta.
Junin: Cahuapanas, 340 meters, Killip & Smith 26791 (type in
U. S. Nat. Herb.), 26758.
Piper calamistratum Trelease, sp. nov.
Frutex parvus (?), internodiis gracilibus breviusculis subcrispe
hirtis; folia lanceolato-elliptica acuminata basi inaequilaterali sub-
acuta 12-14 cm. longa 3.5-4.5 cm. lata undique pinnatinervia, nervis
longis circa 8x2, nervis pluribus reticulatis brevibus superne adjectis,
utrinque laxe crispo-pilosa, subtus ad nervos subvillosa, petiolo 5-10
mm. longo vix alato crispo-piloso; spicae 25 mm. longae 2 mm. crassae
acutatae, pedunculo filiformi 10 mm. longo; bracteae rotundo-sub-
peltatae; ovarium oblongo-ovoideum, apice attenuate stigmatibus
3 terminato.
Loreto: Mishuyacu, in forest, 100 meters, G. Klug 206 (herb.
Field Mus. No. 629,945, type).
Piper callcanense Trelease, sp. nov.
Frutex(?), ramulis crispo-hispidis asperis; folia lanceolata longi-
attentuata basi rotundata 14 cm. longa 4 cm. lata e dimidio inferiore
pinnatinervia, nervis 4-5x2 subtus valde prominentibus et subad-
presso-pilosis, in sicco viridia rugosa, supra granuloso-scabra, petiolo
5 mm. longo; spicae valde curvae 60 mm. longae 3 mm. crassae,
pedunculo 10 mm. longo.
Callcan, 2,400 meters, Pearce, November, 1864 (type in herb.
Kew.).
Piper callosum R. & P. Fl. 1 : 34. pi. 53. 1798. Schilleria callosa
Kunth, Linnaea 13: 689. 1839. Peltobryon callosum Miq. Syst.
369. 1844. Peltobryon Poeppigii Klotzsch in Miq. Syst. 370. 1844.
Piper Poeppigii C. DC. in DC. Prodr. 16, pt. 1: 260. 1869. Piper
callosum var. franciscoanum C. DC. Notizbl. Bot. Gart. Berlin 7:
487. 1917.
A glabrous, slender-twigged shrub; leaves broadly elliptic,
acuminate, the acute base bicallously inrolled, 8-12 cm. long, 4-5
144 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
cm. wide, pinnately nerved from the lower half, the nerves 3-4x2;
petiole 1 cm. long; spikes 2-5 mm. thick, 10-20 mm. long, blunt;
peduncle 5-10 mm. long; bracts inflexed-subpeltate; berries round,
with a style; stigmas 3.
Huanuco: Pillao, Ruiz & Pavon, type. Cochero, Poeppig 1783.
Ayacucho: Aina, Killip & Smith 22715. Junin: La Merced, 600
meters, 5419. Colonia Perene", 680 meters, Killip & Smith 24922,
25095. Rio Pinedo, Killip & Smith 23617. Rio Paucartambo
Valley, Killip & Smith 25399.
Piper calvescentinerve Trelease, sp. nov.
Arbor 5-6-metralis, internodiis floriferis brevibus modice crassis
breviter brunnescenti-villosis; folia oblongo-obovata obtuse subacum-
inata basi inaequilaterali cordata sinu aperto, latere longiore duplo
petiolo breviore, 29-35 cm. longa 13-16 cm. lata e 2 tertiis inferiori-
bus pinnatinervia, nervis circa 6+7, subtus lucida costa breviter
villosa, nervis glabrescentibus, petiolo 5 cm. longo alato breviter
villoso; spicae juveniles 40 mm. longae 2 mm. crassae pendulae,
pedunculo piloso 15 mm. longo; bracteae rotundo-subpeltatae.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26248 (type
in U. S. Nat. Herb.).
Piper calvibaccum Trelease, sp. nov.
Arbor glabra, internodiis floriferis modice gracilibus elongatis
nigrescentibus; folia rotundo-obovata apice rotundata, basi inaequi-
lateralia, sinu laterali fere clause, lobulo longiore rotundato petiolo
longiore, 25 cm. longa 18 cm. lata e 2 tertiis inferioribus subpinna-
tinervia, nervis 6x2, nervis secundariis in auricula dispositis, in
sicco tenuia, petiolo 5 cm. longo alato; spicae 300 mm. longae et
ultra, 5 mm. crassae, pedunculo subgracili 10-20 mm. longo; bracteae
rotundo-subpeltatae pubescentes; baccae globosae, stigmatibus 3
filiformibus sessilibus.
Junin: Chanchamayo Valley, 1,500 meters, Carlos Schunke 1804
(herb. Field Mus. No. 625,549, type) ; at 1,200 meters, Schunke 413.
Piper camphoriferum C. DC. Ann. Cons. Jard. Geneve 21:
235. 1923.
A camphor-scented shrub ; twigs transiently striolate-puberulent,
pale-granular; leaves lanceolate or lance-oblong, very long-attenuate,
the narrowed base cordulate, 15-17 cm. long, 3-4.5 cm. wide, pin-
nately nerved from the lower half, the nerves about 6x2, microscop-
ically granular-roughened, appressed-pubescent beneath toward the
FLORA OF PERU 145
ends of the nerves; petiole scarcely 5 mm. long, slightly longer on
one side; spikes 100 mm. long, 3-4 mm. thick, curved ; bracts lunulate-
subpeltate; berries somewhat quadrate; stigmas 3, sessile. Neg.
24981.
Huanuco: Chihuangola, Gilg 37, type.
Piper canescenticaule Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis gracilibus breviusculis
griseis pilis rigidis incurvis pilosis; folia subelliptico-ovata acuminata
basi inaequilateralia, latere longiore acuto, 16-20 cm. longa 8-10
cm. lata e dimidio inferiore pinnatinervia, nervis 6-7x2, supra
minute scabro-pubescentia et minute aspero-granulosa, subtus ad
nervos subadpresso-strigosa, petiolo 10+5 mm. longo cinereo-piloso
exalato; spicae 100 mm. longae 3 mm. crassae, pedunculo 5 mm.
longo scabrido; bracteae rotundo-subpeltatae.
Loreto: Puerto Arturo, 135 meters, Killip & Smith 27726 (type
in U. S. Nat. Herb.).
Piper canescenticaule var. expansum Trelease, var. nov.
Folia late elliptico-subovata acuminata basi inaequilaterali
subtruncata 22 cm. longa 11 cm. lata.
Loreto: Puerto Arturo, 135 meters, Killip & Smith 27717 (type
in U. S. Nat. Herb.).
Piper capillistipes Trelease, sp. nov.
Frutex 1-2-metralis, internodiis floriferis brevibus subgracilibus
microscopice cinereo-puberulis; folia oblanceolata apicem obtusum
versus sensim angustata, basi inaequilateralia sinu laterali fere
clauso, latere longiore rotundato-auriculato, 29-33 cm. longa 13-15
cm. lata e dimidio inferiore vel paullo ultra subpinnatinervia, nervis
circa 6x2, petiolo 3-5 cm. longo ut pagina inferiore laminae griseo;
spicae brunneae, juveniles 20-25 mm. longae 2 mm. crassae anguste
acutatae, pedunculo 20-35 mm. longo 1 mm. crasso microscopice
cinereo-puberulo ; bracteae rotundo-subpeltatae.
Loreto: Soledad, Rio Itaya, 110 meters, Killip & Smith 29607
(type in U. S.Nat. Herb.).
Piper carpapatanum Trelease, sp. nov.
Frutex nodosus glaber 3-metralis, internodiis floriferis subgracili-
bus brevibus; folia elliptico-subovata acuminata basi inaequilaterali
acuta et ad petiolum decurrentia 10-13 cm. longa 4-6 cm. lata
e dimidio inferiore pinnatinervia, nervis impressis 3-4x2, petiolo
146 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
10 mm. longo; spicae fructiferae 100 mm. longae 5 mm. crassae,
pedunculo crasso 7 mm. longo; bracteae rotundo-subpeltatae ;
baccae cubicae, stigmatibus 3 parvis sessilibus.
Junin: Carpapata above Huacapistana, 2,700-3,200 meters,
Killip & Smith 24403 (type in U. S. Nat. Herb.).
Piper carrapanum Trelease, sp. nov.
Arbor 4-6-metralis, internodiis floriferis modice brevibus et
crassis ferrugineo-araneosis; folia ovata acuta vel subacuminata
basi paullo inaequilateralia uno latere vel utroque latere ad petiolum
acute angustata, 20-22 cm. longa 9-10 cm. lata e tertio inferiore
submultiplinervia, nervis 5-6x2, subtus praesertim ad nervos
stellate- villosa, petiolo 20+3 mm. longo vaginante crispo-subvilloso;
spicae juveniles 30 mm. longae 5 mm. crassae acutatae plus minusve
pilosae.
Ayacucho: Carrapa, 1,200 meters, Killip & Smith 22485 (type
in U. S. Nat. Herb.).
Piper carrapanum var. lancifolium Trelease, var. nov.
Frutex 3-metralis, ramis pilis crassioribus villosis; folia lanceolata
16-17 cm. longa 7 cm. lata; spicae juveniles obtusiores.
Junin: Enenas, Pichis Trail, 1,600-1,900 meters, Killip & Smith
25758 (type in U. S. Nat. Herb.).
Piper Carpunya R. & P. Fl. 1: 37. pi. 63. 1798. Schilleria
Carpunya Kunth, Linnaea 13: 693. 1839. Ottonia Carpunya Miq.
Syst. 547. 1844.
A glabrous shrub; leaves ovate, acuminate, cordate at the base,
9 cm. long, 5 cm. wide, pinnately nerved from the lower half, the
nerves 3x2, drying firm and revolute; petiole 10 mm. long; spikes
2-3 mm. thick, 60 mm. long; peduncle 5 mm. long; bracts incurved;
berries round; stigmas 3, sessile.
Huanuco : Cochero and Chinchao, Ruiz & Pavon, type. Cochero,
Poeppig 1354.
Piper casapiense (Miq.) C. DC. in DC. Prodr. 16, pt. 1: 367.
1869. Artanthe casapiensis Miq. Lond. Journ. Bot. 4: 463. 1866.
A glabrous, glandular-punctulate shrub; leaves elliptic-sub-
oblong, slightly acuminate, subacute at the base with one side
characteristically much shorter, 29-33 cm. long, 10-12 cm. wide,
pinnately nerved throughout; petiole 3+1-3 cm. long; spikes
FLORA OF PERU 147
150 mm. long, 4 mm. thick; peduncle 3 cm. long; bracts triangular-
subpeltate; berries round; stigmas sessile.
Huanuco: Casapi Mathews 1722 bis, type. Junin: La Merced, in
forest, 600 meters, 5517. Chanchamayo Valley, 1,000-1,500 meters,
Schunke 341, 468, 447, 332. Loreto: Balsapuerto, Kittip & Smith
28425.
Piper cassinoides Opiz in Presl, Rel. Haenk. 153. 1830.
Artanlhe cassinoides Miq. Syst. 490. 1844.
A shrub, glabrous except that the petioles and the nerves beneath
are crisp-pubescent; leaves round-ovate, short-acuminate, subtrun-
cate at the base but abruptly contracted to the petiole, 19 cm. long,
13 cm. wide, submultiple-nerved from below the middle, the nerves
about 5x2; petiole scarcely 1 cm. long, winged; young spikes 35 mm.
long, 5 mm. thick, cuspidate; peduncle 10 mm. long.
Huanuco: Huanuco mountains, Haenke, type.
Piper celer Trelease, sp. nov.
Frutex nodosus, internodiis floriferis brevibus et crassis hirsutis;
folia elliptica acuminata basi subacuta 15-16 cm. longa 6 cm. lata e
dimidio inferiore pinnatinervia, nervis 5X2, granuloso-scabra, nervis
subtus hirsutis, petiolo 10 mm. longo hispido; spicae 85 mm. longae
4 mm. crassae, pedunculo crasso 5 mm. longo hispido.
Loreto: Rio Itaya, Llewelyn Williams 132 (herb. Field Mus. No.
604,312, type). Iquitos, Kittip & Smith 27059.
Piper celsum Trelease, sp. nov.
Arbor 5-7-metralis, nervis subtus obscure puberulis exceptis gla-
bra; folia late elliptica obtusa basi inaequilateralia, sinu laterali clauso
lobo majore petiolo longiore, 35 cm. longa et 23 cm. lata vel ultra,
e 2 tertiis inferioribus subpinnatinervia, nervis 6x2, nervis supple-
mentariis in auricula majore dispositis, in sicco tenuia plus minusve
lucida, petiolo 5 cm. (et ultra?) longo alato; spicae 400 mm. longae
3 mm. crassae, pedunculo 1 cm. longo; bracteae crescentico-sub-
peltatae; baccae rotundae glabrae, stigmatibus sessilibus.
Junin: East of Quimiri Bridge, La Merced, 800-1,300 meters,
Kittip & Smith 23907 (type in U. S. Nat. Herb.).
Piper chalhuapuquianum Trelease, sp. nov.
Frutex (?), internodiis fioriferis breviusculis subgracilibus transi-
ente subvillosis serius concolori-granulosis; folia lanceolata acuminata
basi inaequilaterali rotundata vel subcordulata 15 cm. longa 4.5-5
148 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
cm. lata e dimidio inferiore pinnatinervia, nervis 6-7x2, in sicco
tenuia viridia, supra minute lepidoto-asperata, subtus ad nervos
molliter incurvo-pilosa, petiolo 3-5+2 mm. longo incurvo-piloso ;
spicae oppositifoliae 100 mm. longae 3 mm. crassae valde curvae,
pedunculo gracili 10-15 mm. longo molliter hirsute; bracteae sub-
triangulari-subpeltatae pallidae ciliatae; baccae oblongae truncatae
glabrae, stigmatibus 3 minutis sessilibus.
Junin: Hacienda Chalhuapuquio, Stevens 104 (type), 106, 144-
Piper chanchamayanum Trelease, sp. nov.
Frutex nervis subtus crispo-puberulis exceptis glaber, internodiis
floriferis brevibus subgracilibus; folia lanceolato-elliptica acuminata
basi plus minusve inaequilaterali acuta 13-15 cm. longa 5-6 cm.
lata e tertiis 2 inferioribus pinnatinervia, nervis circa 5x2, in sicco
viridia subfirma, supra lucida paullo rugosa, petiolo 6+5 cm. longo
exalato; spicae 100 mm. longae 3 mm. crassae obtusae, pedunculo
crasso 5 mm. longo; bracteae zonatae triangulari-subpeltatae cilio-
latae; baccae quadratae vel cum rhachi elongatae, stigmatibus 3
minutis sessilibus.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 1595
(herb. Field Mus. No. 627,279, type). Ayacucho: Estrella, 500
meters, Killip & Smith 22634.
Piper chromatolepis Trelease, sp. nov.
Frutex plus minusve nodosus nervis subtus adpresso-pubescent-
ibus exceptis glaber, internodiis floriferis brevibus et subcrassis;
folia elliptico-ovata subabrupte breviacuminata basi aequilaterali
cordulata 13-15 cm. longa 7-9 cm. lata e 2 tertiis inferioribus pin-
natinervia, nervis circa 7x2 ut venis impressis, lucida, subtus palli-
diora, petiolo 3 mm. longo caduce alato; spicae 130-150 mm. longae
5 mm. crassae subacutae, pedunculo 10 mm. longo 3 mm. crasso;
bracteae luteae triangulari-subpeltatae ciliatae; ovarium subteres,
stigmatibus 3 filiformibus sessilibus.
Loreto: South side of lower Rio Nanay, in forest, Llewelyn Wil-
liams 536 (herb. Field Mus. No. 604,014, type).
Piper chrysoneurum Trelease, sp. nov.
Frutex nodosus, internodiis floriferis gracilibus brevibus sparse
breviter villosis; folia lanceolata longiacuminata basi inaequilateralia,
latere breviore rotundato, 16-19 cm. longa 6-7 cm. lata e dimidio
inferiore pinnatinervia, nervis 6x2, supra sparse molliter pubescentia,
nervis aurantiacis vel aureis subtus pilosis, petiolo 5+2 mm. longo
FLORA OF PERU 149
inferne vaginante hirsute; spicae juveniles 40 mm. longae 2 mm.
crassae breviter pedunculatae; bracteae rotundo-subpeltatae.
Loreto: Alto Rio Itaya, 145 meters, Llewelyn Williams 3207
(herb. Field Mus., type).
Piper Churumayu R. & P. Fl. 1: 35. pi. 58. 1798. Schilleria
Churumayu Kunth, Linnaea 13: 696. 1839. Artanthe Churumayu
Miq. Syst. 481. 1844.
A glabrous shrub; leaves subovate-lanceolate, acuminate, rounded
at the base, 10-14 cm. long, 4-5 cm. wide, pinnately nerved from the
lower half, the nerves about 4X2; petiole 10 mm. long; spikes 5 mm.
thick, 80 mm. long, straight; peduncle 5 mm. long.
Huanuco: Forests of Huassahuassi ; Ruiz & Pavon, type; Dombey.
"Churumayu."
Piper cinereonervosum Trelease, sp. nov.
Arbor 3-4-metralis, internodiis superioribus ut videtur elongatis
ut petiolo atque nervis subtus microscopice cinereo-puberulis; folia
elliptico-obovata sensim obtusa basi inaequaliter cordata, sinu
laterali, uno latere rotundato altero auriculato, 38 cm. longa 22 cm.
lata e dimidio inferiore vel paullo altius subpinnatinervia, nervis
circa 7x2, pluribus brevioribus in auricula dispersis, petiolo 4 cm.
longo alato; inflorescentiae non visae.
Loreto: San Antonio, 110 meters, Killip & Smith 29380 (type in
U. S. Nat. Herb.).
Piper cinereoramulum Trelease, sp. nov.
Frutex subscandens 3-metralis, ramis geniculatis griseis, inter-
nodiis superioribus brevibus gracilibus sericeis; folia lanceolato-
ovata acute acuminata basi rotundata uno latere paullo breviore,
9-11 cm. longa 3.5-5.5 cm. lata e dimidio inferiore pinnatinervia,
nervis circa 5x2, supra minute granulosa scabra, subtus ad nervos
pallido-sericea, petiolo 5-10-J-3 mm. longo canescente; spicae 65
mm. longae 2 mm. crassae, pedunculo gracili 5 mm. longo; bracteae
rotundo-subpeltatae.
Loreto: Mouth of Rio Santiago, 200 meters, Ynes Mexia 6117
(type in herb. Univ. Calif.).
Piper ciniflonis Trelease, sp. nov.
Frutex parvus, internodiis floriferis brevibus gracilibus crispe
cinereo-strigillosis; folia late lanceolata subacuminata basi breviter
cordata uno latere interdum breviore, 9-11 cm. longa 4-5 cm. latae
150 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
dimidio inferiore pinnatinervia, nervis circa 5x2, griseo-strigillosa,
petiolo 12-15+3 mm. longo late alato cinereo; spicae juveniles 10
mm. longae 1 mm. crassae, pedunculo duplo breviore cinereo; brac-
teae rotundo-subpeltatae, centro fusco, margine angusto cinereo.
Junin: Santa Rosa, Pichis Trail, 625-900 meters, Killip & Smith
26166 (type in U. S. Nat. Herb.).
Piper circumspectantis Trelease, sp. nov.
Frutex metralis nodosus, internodiis floriferis brevibus modice
gracilibus laxe subcrispe hirsutis; folia elliptico-oblonga vel sub-
oblanceolata acuta basi subaequilaterali cordulata 12-16 cm. longa
5-7 cm. lata fere ubique submultiplinervia, nervis longioribus 6x2,
in sicco tenuia, subtus pallidiora nervis hirsutis, petiolo 5 mm. longo
concavo hirsute; spicae 10 mm. longae 5 mm. crassae, staminibus et
stylis exsertis, pedunculo 5 mm. longo hirsute; bracteae subcrescen-
ticae; stigmata 3 brevia.
Loreto: Mishuyacu, 100 meters, in forest, G. Klug 507 (herb.
Field Mus. No. 629,968, type).
Piper cirratum Trelease, sp. nov.
Frutex, internodiis floriferis gracilibus breviusculis rigide sub-
crispo-villosis; folia lanceolata sensim acuta basi inaequaliter
acuta 16 cm. longa 4 cm. lata ubique pinnatinervia, nervis majoribus
circa 5x2, brevioribus superne reticulatis, sparse utrinque crispo-
pilosa, petiolo fere nullo crispo-piloso alato; spicae fructiferae 20
mm. longae 8 mm. crassae, pedunculo sursum curvato gracili 5 mm.
longo crispo-piloso; baccae globosae; bracteae rotundo-subpeltatae,
stigmatibus 3 brevistylosis.
Loreto: La Victoria on the Amazon River, edge of forest, Llewelyn
Williams 2969 (herb. Field Mus. No. 608,159, type).
Piper cispontinum Trelease, sp. nov.
Frutex nodosus 3-4-metralis, internodiis floriferis breviusculis
gracilibus incurvo-hirsutis; folia lanceolato-elliptica anguste falcato-
acuminata basi inaequilaterali angusta obtusa vel cordulata 11-16
cm. longa 4-6 cm. lata e 2 tertiis inferioribus pinnatinervia, nervis
circa 6x2, supra adpresso-scabra, subtus substrigosa, petiolo vix
3+3 mm. longo hispido; spicae juveniles 55 mm. longae 3 mm.
crassae, pedunculo 7 mm. longo scabrido; bracteae rotundo-sub-
peltatae.
Junin: Near Perene" Bridge, Rio Paucartambo, 700 meters, Killip
& Smith 25284 (type in U. S. Nat. Herb.).
FLORA OF PERU 151
Piper claudicans Trelease, sp. nov.
Arbor gracilis 6-metralis nodosa, internodiis floriferis brevibus
cito crassis tuberculatis luteo-subtomentosis; folia elliptico-sub-
oblonga sensim acutata basi rotundata, latere majore sessili altero
15 mm. breviore, 22-20 cm. longa 7-10 cm. lata ubique pinnatinervia,
in sicco tenuia subtus molliter ferrugineo-pilosa, petiolo alato; spicae
100 mm. longae 2 mm. crassae, pedunculo 10 mm. longo flavido-
pubescente; bracteae rotundo-subpeltatae ciliatae; baccae cum
rhachi elongatae; stigmata 3 minuta sessilia.
Loreto: Iquitos, 100 meters, Killip & Smith 26938 (type in U. S.
Nat. Herb.).
Piper Claudii C. DC. Ann. Cons. & Jard. Geneve 2: 264. 1898.
A shrub, the branches and the nerves beneath somewhat crisp-
pubescent; leaves lanceolate, acute at both ends, 10-12 cm. long,
2.5-3 cm. wide, pinnately nerved from the lower half, the nerves
about 5X2; petiole 5 mm. long; spikes 5 (becoming 10) mm. thick,
15-25 mm. long; bracts spatulate; ovary with a style.
Type from Peru, Gay in 1839-1840 in herb. Paris.
Piper coactaepilum Trelease, sp. nov.
Fruticulus nervis subtus breviter tomentosis exceptis glaber
nodosus, internodiis floriferis modice brevibus et gracilibus nigrescen-
tibus; folia ovata subacuminata basi angustata cordulata 13 cm.
longa 6 cm. lata e dimidio inferiore submultiplinervia, nervis 5x2,
in sicco tenuia lucida, petiolo 15 mm. longo infra medium vaginante;
spicae 110 mm. longae 3 mm. crassae acutatae, pedunculo 10-15
mm. longo; bracteae rotundo- vel triangulari-subpeltatae ciliatae;
stigmata minuta sessilia.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26310 (type in
U. S. Nat. Herb.).
Piper cocherense Trelease, sp. nov. P. Carpunya Auct. ex parte.
Ottonia Lessertiana Miq. Syst. 546. 1844; 111. Piper, pi. 92. P.Lesser-
tianum C. DC. in DC. Prodr. 16, pt. 1: 258. 1869, non C. DC. Journ.
Bot. 4: 164. 1866.
Frutex glaber ramulis gracilibus; folia ovata subacuta basi
obtusa vel impresso-subcordata 13-15 cm. longa 7 cm. lata e dimidio
inferiore submultiplinervia, nervis circa 5x2, in sicco viridia coriacea
revoluta, petiolo brevi; spicae 100 mm. longae 2 mm. crassae, pedun-
culo 10 mm. longo; bracteae cucullatae; baccae late obovoideae
breviter mucronatae; stigmata sessilia.
Huanuco: Cochero, Poeppig D1354 (type in herb. Vienna).
152 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper comasense Trelease, sp. nov.
Frutex compactus nodosus glaber vel fugaciter puberulus,
internodiis floriferis brevibus crassis; folia lanceolato-elliptica
utrinque subacuta 7 cm. longa 3 cm. lata e dimidio inferiore pinna-
tinervia, paribus 2 nervorum subbasilaribus, in sicco firma fragilia
saepe plicata, petiolo vix 3 mm. longo exalato; spicae 70 mm. longae
4 mm. crassae leviter curvae, pedunculo crassiusculo 10 mm. longo;
bracteae crescentico-subpeltatae; baccae rotundatae vel ellipsoideae,
stigmatibus 3 parvis sessilibus.
Junin: Rio de Comas, 2,500-2,600 meters, Weberbauer 6615 (herb.
Field Mus. No. 628,196, type).
Piper conditum Trelease, sp. nov.
Frutex glaber nodosus 1-2-metralis, ramis floriferis gracilibus
breviusculis; folia elliptico-subovata acute acuminata basi inaequi-
lateralia latere longiore acuto, breviore subtruncato, 14 cm. longa 6-7
cm. lata e dimidio inferiore pinnatinervia, nervis 5+4, in sicco tenuia
subtus minute nigro-glandulosa, petiolo 7+3 mm. longo evaginante;
spicae juveniles 35 mm. longae 2 mm. crassae obtusae; pedunculo
10 mm. longo; bracteae rotundo-triangulari-subpeltatae ciliatae.
Loreto: Yurimaguas, 135 meters, Killip & Smith 28208 (type in
U. S. Nat. Herb.). Yurimaguas to Balsapuerto, Killip & Smith
28090, 28370.
Piper conditum var. angustum Trelease, var. nov.
Folia lanceolato-oblonga vel elliptico-oblonga 13-16 cm. longa
5-6 cm. lata in sicco firmiora; spicae juveniles vix 10 mm. longae
et 1 mm. crassae.
Loreto: Yurimaguas, Killip & Smith 29105 (type in U. S. Nat.
Herb.).
Piper confusionis Trelease. sp. nov.
Arbor 3-4-metralis, ramulis dense rubro-villosis; folia obovata
acuminata basi inaequilateralia sinu laterali, lobo ma j ore rotundato
petiolum rubro-villosum superante, 33 cm. longa et 20 cm. lata vel
ultra, e dimidio inferiore subpinnatinervia, nervis 5x2, nervis supple-
mentariis in auricula conspersis, supra glabra subtus ad nervos
dense ferrugineo-villosa, in sicco tenuia, petiolo 4 cm. longo alato;
spicae juveniles 55 mm. longae 3 mm. crassae, pedunculo rubro-
villoso 2 cm. longo; bracteae rotundo-subpeltatae glabrae.
Loreto: Yurimaguas to Balsapuerto, 135-150 meters, Killip &
Smith 28231 (type in U. S. Nat. Herb.). Mainas, Poeppig in 1831.
FLORA OF PERU 153
Junin: Enenas, 1,600-1,900 meters, Killip & Smith 25734. With-
out locality, Poeppig 25(1). Brazil: Seringal Sao Francisco, Ule
9293.
Piper conispicum Trelease, sp. nov.
Arbor humilis glabra, internodiis floriferis subgracilibus brevibus;
folia lanceolata vel elliptica acute acuminata basi cordulata, lobo
altero majore, 15-18 cm. longa 6-8 cm. lata undique pinnatinervia,
nervis 8-10x2, petiolo vix 5 mm. longo latus subacutum longius folii
aequante late alato; spicae 11 mm. longae 6 mm. latae initio anthesis
late conicae, pedunculo vix 4 mm. longo; ovarium obconicum in
stylum gracilem attenuatum; stigmata 3 minuta patentia.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28767 (type in
U. S. Nat. Herb.).
Piper constipatispicum Trelease, sp. nov.
Frutex vel arbor 3-4-metralis, internodiis floriferis graciusculis
elongatis interdum fistulosis araneosis; folia ovata acuminata basi
plus minusve inaequaliter rotundata vel subretrorsa, margine acute
ad petiolum protracto, 19-22 cm. longa 10-13 cm. lata vel usque ad
28 cm. longa atque 15 cm. lata, e dimidio inferiore submultiplinervia,
nervis 5x2, in sicco tenuia, subtus pallidiora stellato-tomentosa ad
nervos subvillosa, petiolo 15-20+5 mm. longo infra medium va-
ginante subvilloso; spicae juveniles 4-6 mm. longae 3 mm. crassae,
pedunculo conico 6 mm. longo pubescente; bracteae rotundo-sub-
peltatae hirtellae.
Loreto: San Antonio, 110 meters, Killip & Smith 29415 (type in
U. S. Nat. Herb.), 29442, 29302.
Piper constipatispicum var. lanosum Trelease, var. nov.
Arbor 4-5-metralis, ramulis, petiolis, pedunculis et pagina
inferiore folii stellatim albido-lanosis; spicae juveniles 10 mm.
longae 3 mm. crassae acutatae.
Loreto: Iquitos, 110 meters, Killip & Smith 27467 (type in U. S.
Nat. Herb.). Soledad, Killip & Smith 29802.
Piper constipatispicum var. tonsum Trelease, var. nov.
Frutex 3-4-metralis, ramulis tantum tomentulosis, aliter ut in
var. lanosum.
Loreto: Soledad, 110 meters, Killip & Smith 29802 (type in
U. S, Nat. Herb.).
154 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper contamanum Trelease, sp. nov.
Frutex nodosus 3-metralis nervis folii subtus minute puberulis
exceptis glaber, internodiis floriferis brevibus gracilibus pallido-
granulosis; folia lanceolato-elliptica subacuminata basi inaequaliter
acuta 13-18 cm. longa 4-7 cm. lata e 2 tertiis inferioribus pinna-
tinervia, nervis 5x2, petiolo 7+3 mm. longo non vaginante; spicae
juveniles 30 mm. longae 2 mm. crassae, pedunculo brevi; bracteae
parvae albae triangulari-subpeltatae.
Loreto: Contamana on Rio Ucayali, 150 meters, Killip & Smith
26880 (type in U. S. Nat. Herb.).
Piper controversum Steud. Nom. Bot. ed. 2. 2: 340. 1841.
Piper caracasanum Opiz in Presl, Rel. Haenk. 155. 1830, non P.
caracasanum Bredem.
A shrub, subtomentose-subvillous except for the glabrescent upper
surface of the leaves; leaves broadly ovate, short-acuminate, ob-
liquely cordate with the larger side equaling the petiole, 24-30 cm.
long, 14-17 cm. wide, subpinnately nerved from below the upper
third, the nerves about 6x2; petiole 4 cm. long, scarcely shorter on
one side; inflorescence unknown.
Huanuco: Huanuco, Haenke, at Berlin; Haenke 71, at Vienna,
types.
Piper cordatum C. DC. Journ. Bot. 4: 213. 1866.
A shrub with twigs, leaf margin, and nerves beneath somewhat
crisp-puberulent; leaves broadly elliptic, acute, shallowly cordate,
25-28 cm. long, 11-14 cm. wide, pinnately nerved throughout;
petiole barely 2 cm. long, dilated at the base, sheathing to the middle;
spikes becoming 80 mm. long, 10 mm. thick; peduncle thick, scarcely
10 mm. long; bracts large, brown, subpeltate; berries obovoid, with a
style; stigmas short. Neg. 7229.
San Martin: Monte Pingullu, Tarapoto, Spruce 4847, type.
Loreto: Cerro de Ponasa, 1,200 meters, Ule 6828.
Piper correptum Trelease, sp. nov.
Frutex 3-metralis nodosus, internodiis floriferis brevibus crassius-
culis cito glabrescentibus; folia lanceolata longiacuminata basi
paullo inaequilateralia, latere breviore minute subcordulato, 13-14
cm. longa 4 cm. lata e dimidio pinnatinervia, nervis circa 6x2,
subtus ad nervos adpresso-pubescentia, petiolo 5 mm. longo plus
minusve crispo-pubescente ; spicae 80 mm. longae 3 mm. crassae,
FLORA OF PERU 155
pedunculo vix 10 mm. longo glabrescente; bracteae parvae rotundo-
peltatae ciliolatae; baccae subglobosae, stigmatibus sessilibus.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26524
(type in U. S. Nat. Herb.).
Piper costatum C. DC. Bot. Jahrb. 40: 243. 1908.
A shrub, glabrous except that the leaves are crisp-ciliate and
obscurely puberulent on the nerves beneath; twigs costate; leaves
narrowly oblanceolate, acute at both ends, 20-28 cm. long, 5-7 cm.
wide, pinnately nerved throughout; petiole very short, with the
blade decurrent on it; spikes gently curved, 50 mm. long, 5-6 mm.
thick; peduncle 10-15 mm. long; berries ovoid, with a slender style;
stigmas 3, spreading.
Junin: Chanchamayo valley, Weberbauer 1857, type.
Piper crassamentum Trelease, sp. nov.
Frutex 3-metralis nodosus, internodiis floriferis crassis breviusculis
crispe ferrugineo-pilosis; folia ovata acute acuminata basi rotundata
23 cm. longa 11 cm. lata e tertio inferiore submultiplinervia, nervis
5x2, subtus ferrugineo-pubescentia, nervis breviter subvillosis,
petiolo 15 mm. longo crispe ferrugineo-piloso alato; spicae fructiferae
100-140 mm. longae 10 mm. crassae, pedunculo 10 mm. longo 4 mm.
crasso ferrugineo-piloso; bracteae triangulari-subpeltatae; baccae
rotundatae, stylo gracili, stigmatibus 3 linearibus.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26354 (type
in U. S. Nat. Herb.).
Piper crocatum R. & P. Fl. 1: 35. pi. 55. 1798. Steffensia
crocata Kunth, Linnaea 13: 652. 1839. Artanthe crocata Miq. Syst.
429. 1844.
A yellow- villous-subhirsute shrub; leaves elliptic, acuminate,
subacute at the base, 9-12 cm. long, 4-5 cm. wide, pinnately nerved
from the lower half, the nerves 4-5x2, bullulate-lacunose, hirsutely
scabrid above, villous-hirsute beneath; petiole 10 mm. long; spikes
90-110 mm. long, 5 mm. thick, nearly straight; peduncle 5 mm. long;
bracts subtriangular-subpeltate, ciliate; berries subglobose; stigmas
3, rather broad, sessile. Neg. 8408.
Huanuco: Chachahuassi, Ruiz, at Berlin, and as No. 657 in Herb.
Willdenow.
Piper cumbasonum C. DC. Verh. Bot. Ver. Brandenb. 47:
108. 1905.
156 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
A glabrous, obscurely punctulate shrub, scarcely separable from
P. politum except for its dull leaves.
Loreto: San Pedro, Rio Cumbaso, Ule 6826, type. San
Martin: Tarapoto, Williams 5793. San Roque, Williams 7639.
"Cordoncillo."
Piper cupreatum Trelease, sp. nov.
Arbor gracilis 3-4-metralis nervis subtus obscure puberulis
exceptis glabra, internodiis superioribus gracilibus nigrescentibus;
folia subpentagone elliptico-obovata paullo obtuso-acuminata basi
oblique aperte cordata 23-26 cm. longa 12-15 cm. lata e 2 tertiis
inferioribus subpinnatinervia, nervis circa 7+8, in sicco tenuia
lucida pubescentia, petiolo 30+5 mm. longo alato; spicae juveniles 20
mm. tantum longae 2 mm. crassae, pedunculo contorto duplo
breviore; bracteae rotundo-subpeltatae.
Loreto: Balsapuerto, 350-550 meters, Killip & Smith 28509
(type in U. S, Nat. Herb.). Santa Rosa, Killip & Smith 28784.
Yurimaguas to Balsapuerto, Killip & Smith 28274-
Piper cupreatum var. longum Trelease, var. nov.
Arbor gracilis 4-5-metralis; folia elongato-ovata 30 cm. longa
15 cm. lata; petiolo 6+1 cm. longo; spicae juveniles erectae 40 mm.
longae 2 mm. crassae, pedunculo duplo breviore.
Loreto: Iquitos, 100 meters, Killip & Smith 27486 (type in U. S.
Nat. Herb.).
Piper curvatum R. & P. Fl. 1: 36. pi. 54- 1798. Artanthe curvata
Miq. Syst. 522. 1844.
An obscure, glabrous, somewhat glandular-punctulate shrub;
leaves elliptic-oblong, acute at both ends, 12-13 cm. long, 4 cm. wide,
pinnately nerved throughout; petiole 10-15 mm. long; spikes becom-
ing strongly curved and 80 mm. long, 3-4 mm. thick; peduncle 10
mm. long.
Huanuco: Cochero and Chinchao, Ruiz & Pawn, type.
Unknown except for the original description and illustration,
unless the Peruvian P. colubrinum, with obliquely excised leaf
base, be the same.
Piper daedalum Trelease, sp. nov.
Frutex humilis plus minusve nodosus, internodiis floriferis gracili-
bus subelongatis; folia lanceolato-elliptica acuminata basi sub-
inaequilaterali acuta vel in foliis latioribus subobtusa 9-10 cm. longa
4-4.5 cm. lata undique pinnatinervia vel nervis e tertio superiore
FLORA OF PERU 157
obscuris, inferioribus circa 5X2, in sicco subopaca subtus glanduloso-
granulosa, petiolo 10+3 vel 15 mm. longo exalato; spicae fructiferae
20 mm. longae 5 mm. crassae caduce apicatae, pedunculo filiformi;
bracteae triangulari-subpeltatae ferrugineo-ciliatae; baccae globosae,
stigmatibus 3 mimitis, stylo distincto.
Loreto: Lower Rio Huallaga, 155-210 meters, Llewelyn Williams
3861 (herb. Field Mus., type).
Piper dasyoura (Miq.) C. DC. in DC. Prodr. 16, pt. 1: 269.
1869. Artanthe dasyoura Miq. Lond. Journ. Bot. 4: 452. 1866.
A shrub with subretrorsely hirsute- villous twigs; leaves lance-
elliptic, subacuminate, acute-based, 16-20 cm. long, 6-8 cm. wide,
pinnately nerved throughout, finely rugulose, soft-pubescent; petiole
10 mm. long; spikes 65 mm. long, 5 mm. thick, gently curved;
peduncle 10 mm. long, hirtellous; bracts crescentic-subpeltate,
ciliate; berries round; stigmas 3, sessile.
Type collected in Peru, Mathews 1716.
Piper dasypodum (Miq.) C. DC. in DC. Prodr. 16, pt. 1:
304. 1869. Artanthe dasypoda Miq. Lond. Journ. Bot. 4: 458. 1866.
A shrub, fuscous-villous throughout; leaves elliptic-subovate,
slender-acuminate, nearly equilaterally cordulate, 14-18 cm. long,
6-8 cm. wide, pinnately nerved from the lower half, the nerves
about 5x2; petiole 10 mm. long; young spikes 15 mm. long, 8
mm. thick, subcuspidate; peduncle barely 5 mm. long. Neg. 8407.
Type from Peru, Mathews 1 702, type.
Piper delatum Trelease, sp. nov.
Frutex nodosus 3-4-metralis, internodiis floriferis gracilibus
brevibus evanescente puberulis albido-granulosis; folia anguste
lanceolata acute acuminata basi inaequaliter acuta 10 cm. longa
3.5 cm. lata e 2 tertiis inferioribus pinnatinervia, nervis 5-6x2,
subtus canescentia, nervis dense adpresso-pubescentibus, inter nervos
sparsius pubescentia, petiolo 3+2 mm. longo cinereo-pubescente;
spicae 75-85 mm. longae 2-3 mm. crassae, pedunculo vix 5 mm.
longo glabrato; bracteae parvae rotundo-subpeltatae ciliolatae;
baccae subcylindraceae truncatae; stigmata 3 minuta sessilia.
Loreto: Puerto Arturo, 135 meters, Killip & Smith 27713 (type
in U. S. Nat. Herb.).
Piper demoratum Trelease, sp. nov.
Frutex glaber nodosus, internodiis brevibus graciliusculis ; folia
ovata vel lanceolato-ovata subacuminata basi aequilateralia, basi
158 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
angusta subacuta vel subcordulata, 12-13 cm. longa 6 cm. lata e
dimidio inferiore submultiplinervia, nervis 5x2, coriacea, petiolo
10 mm. longo exalato; spicae 60 mm. longae 3 mm. crassae, pedunculo
5 mm. longo; bracteae triangulari-subpeltatae ciliolatae; baccae plus
minusve triquetrae; stigmata 3 lata sessilia adpressa.
Junin: Chanchamayo Valley, 1,500 meters, Carlos Schunke 279
(herb. Field Mus. No. 571,335, type).
Piper dendropse Trelease, sp. nov.
Arbor 3-5-metralis nodosa nervis folii subtus minute crispo-
pubescentibus exceptis glabra, internodiis floriferis brevibus gracili-
usculis; folia lanceolato-elliptica subacuminata basi inaequaliter
acuta 12-17 cm. longa 4-6 cm. lata e 2 tertiis inferioribus pinnati-
nervia, nervis circa 6x2, in sicco tenuia sed firma, petiolo 6-f4
mm. longo exalato; spicae 90-100 mm. longae 3 mm. crassae mucro-
natae, pedunculo vix 5 mm. longo; bracteae triangulari-subpeltatae
ciliolatae; baccae depressae rotundatae vel cum rhachi elongatae;
stigmata 3 minuta sessilia.
Junin: La Merced, 700 meters, Killip & Smith 2S488 (type in
U. S. Nat. Herb.).
Piper Dennisii Trelease, sp. nov.
Frutex 2-metralis multiramosus nodosus, internodiis floriferis
brevibus gracillimis evanescente intertexto-sericeis; folia anguste
lanceolata attenuato-acuta basi subaequaliter acuta 8 cm. longa
1 cm. lata e tertio inferiore pinnatinervia, nervis 4x2 breviter
araneosis, petiolo 4+1 mm. longo paullo pubescente exalato; spicae
40 mm. longae 2 mm. crassae acutatae, pedunculo 5 mm. longo
minute cinereo-pubescente; bracteae rotundo-subpeltatae ciliolatae;
baccae teretes, stigmatibus sessilibus.
Loreto: Rio Maranon Valley, 150 meters, W. J. Dennis (Killip
6 Smith 29137; type in U. S. Nat. Herb.).
Piper desultorium Trelease, sp. nov.
Arbor 3-4-metralis, internodiis floriferis brevibus gracilibus in-
curvo- vel adpresso-hirsutis; folia oblongo-lanceolata acuminata basi
plus minusve inaequaliter acuta 11-15 cm. longa 5-7 cm. lata e
dimidio inferiore pinnatinervia, nervis 5-6x2, in sicco tenuia caerule-
scentia, supra lepidoto-asperata sed vix granulosa, subtus ad nervos
subadpresso-strigillosa, petiolo 8+2 mm. longo hispido exalato;
spicae 70 mm. longae 3-4 mm. crassae caudatae, pedunculo 5-6
mm. longo scabrido-villoso ; bracteae rotundo-subpeltatae; baccae
teretes, stigmatibus sessilibus.
FLORA OF PERU 159
Junin: Schunke Hacienda, La Merced, 1,400-1,700 meters,
Killip & Smith 24654 (type in U. S. Nat. Herb.). Chanchamayo
Valley, 1,500 meters, Schunke 252 (herb. Field Mus.).
Piper dichotomum R. & P. Fl. 1: 35. pi. 60. 1798. Peltobryon
dichotomum Klotzsch in Miq. Syst. 372. 1844.
A glabrous shrub, but with the young parts minutely papillate;
leaves elliptic, acuminate, acute-based, 15 cm. long, 6 cm. wide,
pinnately nerved from below about the upper third; petiole 10
mm. long; spikes 30 mm. long, 8-10 mm. thick; peduncle 10 mm.
long; bracts subpeltate; berries round, with a style; stigmas 3, short.
Huanuco: Chinchao, Macora, Mesapata, Ruiz & Pawn, type.
Piper dimetrale C. DC. Bot. Jahrb. 40: 245. 1908.
A shrub, somewhat soft-hairy when young; leaves lance-ovate,
acuminate, slightly inequilaterally rounded at the base, 14 cm.
long, 6 cm. wide, pinnately nerved from somewhat more than the
lower half, the nerves 5-6x2, becoming rugose, hirsute beneath;
petiole 8+2 mm. long, hirsute; spikes 70 mm. long, 2 mm. wide;
peduncle 15-20 mm. long.
Puno: Sillacunca, at 1,500 meters, in thicket, Weberbauer
1281, type.
Piper dispansum Trelease, sp. nov.
Frutex glaber 2-3-metralis, ramis patentibus plus minusve
nodosis et geniculatis, internodiis floriferis subgracilibus brevibus;
folia elliptico-oblonga breviter acuminata basi fere aequaliter acuta
15 cm. longa 5-6 cm. lata fere ubique pinnatinervia, nervis circa
6x2, intermediis adjectis, pallido-granulosa, petiolo 1 cm. longo
exalato; spicae juveniles vix 6 mm. longae 1 mm. crassae, pedunculo
brevissimo.
Junin: Puerto Bermudez, 375 meters, in thicket, Killip & Smith
26434 (herb. Field Mus. No. 607,559, type). Loreto: Yurimaguas,
Killip & Smith 27684, 27612. Puerto Arturo, Killip & Smith 27882.
Piper diversipilum Trelease, sp. nov.
Subarborescens ubique laxe hirtella et longivillosa, internodiis
floriferis breviusculis crassiusculis; folia inaequaliter vel subfalcate
lanceolato-oblonga sensim longiacuminata, basi inaequaliter obtusa,
modice parva 18-20 cm. longa 6-6.5 cm. lata e dimidio inferiore
pinnatinervia, nervis circa 6x2, in statu adulto plus minusve grosse
bullata vel rugosa, petiolo brevissimo vix 3+3 mm. longo exalato;
160 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
spicae oppositifoliae fere rectae 80 mm. longae 3 mm. crassae,
pedunculo breviusculo 10-15 mm. longo; bracteae rotundo-sub-
peltatae ciliatae, floribus sessilibus perfectis; stigmata 3 sessilia
brevia; baccae globosae glabrae.
Huanuco: Posuso, 600 meters, Macbride 4666 (herb. Field Mus.
No. 535,740, type).
Piper dulce Trelease, sp. nov.
Frutex glaber plus minusve nodosus 3^4-metralis, internodiis
superioribus brevibus gracilibus; folia elliptica anguste acuminata
basi fere aequaliter subacuta e dimidio inferiore pinnatinervia,
nervis 5-6x2, in sicco tenuia, petiolo 10 mm. longo tantum ad
basin vaginante; spicae juveniles 15 mm. tantum longae 2 mm.
crassae, pedunculo brevi crasso; bracteae rotundo-subpeltatae.
Junin: La Merced, 700 meters, Kittip & Smith 23705 (type in
U. S. Nat. Herb.).
Piper Durvilleanum (Miq.) Trelease, comb. nov. Artanthe
Durvilleana Miq. Syst. 507. 1844. P. anonaefolium var. rigidulum
C. DC. in DC. Prodr. 16, pt. 1: 299. 1869. P. colubrinum Auct.
as to Peru.
A glabrous, glandular-granular shrub; leaves elliptic-oblong,
short-acuminate, acute-based with one side concavely excised, 15
cm. long, 6 cm. wide, pinnately nerved throughout; petiole 10-20
mm. long; spikes curved, 60 mm. long, 4 mm. thick, pointed ; peduncle
15 mm. long; bracts subtriangular-subpeltate; berries round; stigmas
sessile.
Huanuco: Cochero, Poeppig 3113, type.
Piper ecallosum Trelease, sp. nov.
Frutex glaber; folia ovata acuminata basi subaequaliter obtusa
12 cm. longa 6 cm. lata e dimidio inferiore pinnatinervia, nervis
circa 4x2, subcoriacea, subtus granulosa, nervis subimpressis,
petiolo 8+2 mm. longo exalato; spicae juveniles 20 mm. longae 5
mm. crassae, pedunculo brevissimo.
Junin: Pangoa, Mathews 1147 (type in herb. Vienna).
Piper edurum Trelease, sp. nov.
Frutex 2-metralis nodosus glaber, internodiis floriferis brevibus
modice gracilibus; folia elliptico-ovata acute subacuminata basi
infima subaequaliter acuta 14 cm. longa 6-7 cm. lata e dimidio
inferiore submultiplinervia, nervis 4-5x2, in sicco crassa dura,
FLORA OF PERU 161
petiolo 10-15 mm. longo exalato; spicae 35-50 mm. longae 3-4
mm. crassae, pedunculo 5 mm. longo; bracteae crescenticae; baccae
rotimdae in sicco conico-subtruncatae, stigmatibus 3 sessilibus.
Ayacucho: Aina, 750-1,000 meters, Killip & Smith 22618 (type
in U. S. Nat. Herb.), 22566. Junin: Huacapistana, Killip & Smith
24325.
Piper elongatifolium Trelease, sp. nov. P. angustifolium Auct.
ex parte. P. elongatum Auct.
Frutex nodosus, internodiis floriferis brevibus crassiusculis tran-
siente hirtellis dense pallido-granulosis; folia lanceolato-oblonga vel
elliptico-oblonga acuminata basi breviter cordulata 13-19 cm. longa
4-6.5 cm. lata e dimidio inferiore pinnatinervia, nervis 5-6x2,
supra lepidota, subtus microscopice granuloso-punctulata nervis
subadpresso-hirsutis glabrescentibus, petiolo 5 mm. longo sub-
hispido exalato; spicae oppositifoliae valde curvae 120 mm. longae
3 mm. crassae, pedunculo 10 mm. longo plus minusve hirtello;
bracteae rotundae vel triangulari-subpeltatae ciliolatae, floribus
sessilibus perfectis; baccae minutae obconicae, stigmatibus 3 minutis
sessilibus.
Junin: La Merced, sunny slopes, 600 meters, Macbride 5287
(herb. Field Mus. No. 536,312, type). Rio Perene", Killip & Smith
25113, 25121*.
Piper elongatum Vahl, Enum. 1: 312. 1805. P. angustifolium
R. & P. Fl. 1: 38. pi. 57. 1798; Kramer, Soc. Appl. Pharmacog. 189.
/. 65; Mansfield in Squibbs Atlas, 368. /; non P. angustifolium
Lam. 1791. P. purpurascens D. Dietr. Syn. PI. 1: 119. 1839. Stef-
fensia elongata Kunth, Linnaea 13: 637. 1839; Gaudichaud, Voy.
Bonite pi. 95. Artanthe elongata Miq. Syst. 434. 1844.
A resinous-aromatic shrub about 2 meters tall, very nodose;
flowering internodes short and moderately stout, sparsely hamate-
hirsute, glabrescent and somewhat pale-granular in age; leaves
lance-oblong, long-attenuate rather than acuminate, slightly un-
equally cordulate at the base with the sinus sometimes closed by
overlapping of the shorter lobe, 15-22 cm. long, 3.5-5 cm. wide,
pinnately nerved from the lower half or two-thirds, the nerves
8-9x2, very bullulate, scabrous above, densely upcurved-hirsute
on the nerves and veins beneath; petiole very short, 5-6 mm. long,
upcurved-hirsute, not winged; spikes strongly curved, 100 mm.
long, 3 mm. thick, or larger; peduncle slender, 10 mm. long, sparsely
hirtellous; bracts conspicuous, round- or triangular-subpeltate, the
162 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
tawny margin rusty-ciliate; stigmas 3, sessile; berries small, obconic.
Neg. 8400 (P. angustifoUum).
Huanuco: Huanuco, Ruiz 122, 401; Macbride 3498. Cuzco;
Machupicchu, 2,000-2,400 meters, Herrera 3191. Also in Bolivia.
"Matico."
The narrow-leaved "matico" of pharmacy, from the eastern
slope of Peru and Bolivia. The confusion attending its scientific
name is discussed in Bull. Torrey Bot. Club 56: 535.
Piper elongatum var. Mocco-mocco Trelease, var. nov.
Frutex nodosus, internodiis brevibus caduce molliter pallido-
villosis; folia conspicue bullulata supra glabra; spicae juvenes 40
mm. longae 3 mm. crassae rectae, apice sterili curvato, pedunculo
5 mm. longo velutino.
Cuzco: Santa Ana Valley, Santa Rita, 2,300 meters, Herrera
1551, type. "Mocco-mocco."
Piper elongatum var. pampayacusum Trelease, var. nov.
Frutex nodosus, internodiis breviusculis; folia scabrido-bullulata
subtus molliter subadpresso-pilosa; spicae curvae vel serius fere
rectae 60 mm. longae 3 mm. crassae apice gracili sterili, pedunculo
5 mm. longo sparse velutino.
Huanuco: Pampayacu, at mouth of Rio Chinchao, 1,100 meters,
Macbride 5047 (herb. Field Mus., type).
Piper elongatum var. salviaefolium (Miq.) Trelease, comb,
nov. Artanthe salviaefolia Miq. Syst. 430. 1844. P. salviaefolium
C. DC. in DC. Prodr. 16, pt. 1: 270. 1869.
A "matico" shrub with the general characters of the type but
with broadly lanceolate leaves 10-15 cm. long and 2-5 cm. wide.
Huanuco: Huanuco, Dombey 934, 71; Gilg 39; Haenke in 1790^
Kanehira 101. Without locality: Pavon at Paris; Mathews 66,
Piper Endlicherianum (Miq.) Trelease, comb. nov. Arta
Endlicheriana Miq. Lond. Journ. Bot. 4: 441. 1866.
A glabrous shrub with pale-granular axes; leaves ovate, acumi-
nate, shallow-cordate, 20 cm. long, 10 cm. wide, pinnately nerved
from the lower third, the nerves about 5x2; petiole 15 mm. long,
narrow, winged; spikes 110 mm. long, 3 mm. thick; peduncle 10
mm. long.
Huanuco: Casapi, Mathews 1698 (corrected from 305), type.
FLORA OF PERU 163
Very likely a glabrous form of P. formosum, to which de Candolle
reduced it.
Piper enenyasense Trelease, sp. nov.
Frutex 2-3-metralis, internodiis floriferis brevibus gracilibus
dense albido-villosis; folia lanceolata acuminata basi interdum in-
aequilaterali obtusa 9-15 cm. longa 3-5 cm. lata supra basibus
pilorum mollium fugacium minute granulosa, subtus pallidiora
nervis venisque pallido-villosis, petiolo 5 mm. vel 3+2 mm. longo
villoso; spicae juveniles 10 mm. longae 2 mm. crassae, pedunculo
pallido-villoso fere aequilongo.
Junin: Enenas, Pichis Trail, 1,600-1,900 meters, Killip & Smith
25646 (type in U. S. Nat. Herb.).
Piper epunctatum Trelease, sp. nov.
Frutex glaber 2-metralis in sicco nigrescens, internodiis floriferis
longis gracilibus; folia lanceolata ad elliptico-subovata anguste
acuminata basi paullo inaequilaterali acuta vel in foliis majoribus
aequilaterali-obtusa 15 cm. longa 4.5-6.5 cm. lata ubique pinnati-
nervia, nervis circa 10x2, in sicco tenuia lucida, petiolo 5-20 mm.
longo tenuiter alato; spicae juveniles 4 mm. tantum longae et 3
mm. latae cuspidatae, pedunculo aequilongo; bracteae rotundo-
subpeltatae pubescenti-marginatae.
Loreto: Mishuyacu near Iquitos, 100 meters, in forest, G. Klug
1271 (herb. Field Mus. No. 627,689, type).
Piper escaleranum C. DC. Verh. Bot. Ver. Brandenb. 47:
115. 1905.
A shrub with the young parts and the nerves beneath thinly
villous-hirsute; leaves lanceolate, acute, inequilaterally cordate
with the longer side covering the petiole, 20-21 cm. long, 6.5-8
cm. wide, subpinnately nerved from the lower half, the nerves
I about 5+6; petiole short, winged; young spikes 15-20 mm. long,
7*^2 mm. thick, pointed; peduncle filiform, 20-30 mm. long. , .
Loreto: Cerro de Escalera, 1,200 meters, Ule 6818 (type), 6827.
Piper euryphyllum Trelease, sp. nov.
Arbor 3-metralis nervis folii subtus minute adpresso-pubescenti-
bus exceptis glabra, internodiis floriferis comparate brevibus et
crassis; folia rotundato-ovata breviter acuminata basi truncato-
cordata, uno latere paullo breviore, 28 cm. longa 18-20 cm. lata e
dimidio inferiore pinnatinervia, nervis 7-8x2, in sicco crasso-papy-
164 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
racea viridescentia, petiolo 20+5 mm. longo basin versus vaginante
ut costa transiente pubescente; spicae 150 mm. longae 5 mm. crassae,
pedunculo 15 mm. longo; bracteae crescenticae inconspicuae; baccae
pallide brunneae depressae cum rhachi paullo elongatae, stigmatibus
sessilibus.
Junin: East of Quimiri Bridge, near La Merced, 800-1,300
meters, Killip & Smith 29891 (type in U. S. Nat. Herb.).
Piper excavatum R. & P. Fl. 1: 34. pi. 54- 1798. Artanthe
excavata Miq. Syst. 534. 1844.
A glabrous shrub, the twigs fistulous; leaves elliptic or subovate-
elliptic, acuminate, subacute at the base, 11-15 cm. long, 4.5-7
cm. wide, pinnately nerved from the lower half, the nerves about
5X2; petiole 10 mm. long; spikes about 70 mm. long, 4 mm. thick,
straight, blunt.
Huanuco: Forests of Cochero, Ruiz & Pawn, type. An obscure
species.
Piper excessum Trelease, sp. nov.
Frutex glaber; folia subovato-elliptica acuminata basi obliqua,
uno latere cuneatim ad petiolum decurrente, 17 cm. longa 8 cm.
lata undique pinnatinervia, nervis 8-10x2, petiolo 5+10 mm. longo
alato; spicae 80-110 mm. longae 3 mm. crassae, pedunculo 10
mm. longo; bracteae triangulari-subpeltatae ciliatae; baccae cum
rhachi elongatae depressae; stigmata 3 parva sessilia.
San Martin: Tocache, Huallaga, Poeppig addenda 154 in 1830
(type in herb. Vienna).
Piper excultum Trelease, sp. nov.
Frutex nodosus margine petioli paullo puberulo excepto glaber,
internodiis floriferis gracilibus breviusculis; folia elliptico-ovata
acuminata basi inaequilateralia, latere longiore subacuto, 10-12
cm. longa 3.5-6 cm. lata e dimidio inferiore pinnatinervia, nervis
circa 7x2 supra impressis subtus elevatis, in sicco firma lucida non
granulosa, petiolo vix 8+2 mm. longo exalato; spicae modice curvae
100 mm. longae 2 mm. crassae, pedunculo 5-10 mm. longo; bracteae
parvae rotundo-subpeltatae ciliolatae.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
7461 (herb. Field Mus. No. 623,031, type).
Piper expolitum Trelease, sp. nov.
Frutex 3-metralis nodosus glaber, internodiis floriferis brevius-
culis crassis fistulosis; folia elliptica vel subovata vel oblonga abrupte
FLORA OF PERU 165
breviacuminata basi plus minusve inaequilaterali acuta 12-14 cm.
longa 5-6 cm. lata e 2 tertiis inferioribus pinnatinervia, nervis circa
5x2, in sicco tenuia flavido-lucida, petiolo circa 10+3 mm. longo
exalato; spicae plus minusve arcuatae 50 mm. longae 4 mm. crassae
cuspidatae, pedunculo 10 mm. longo; bracteae rotundo-subpeltatae,
margine lato firmo.
Loreto: Iquitos, Llewelyn Williams 3590 (herb. Field Mus. No.
618,287, type); Killip & Smith 26897, 27228. Pebas, in forest,
Williams 1944- "Cordoncillo."
Piper faecatum Trelease, sp. nov.
Frutex glaber, internodiis floriferis subgracilibus brevibus; folia
elliptico-suboblonga acuminata basi rotundata, latere majore inter-
dum subacuto, 23-26 cm. longa 8-10 cm. lata undique pinnati-
nervia, nervis circa 12x2, obscure glanduloso-granulosa, petiolo
vix 5 mm. longo exalato; spicae juveniles 35-40 mm. longae 4-5
mm. crassae acutatae, pedunculo 10 mm. longo; bracteae trans-
versae subpeltatae.
Loreto: Pebas on the Amazon River, Llewelyn Williams 1593
(herb. Field Mus. No. 613,181, type). "Cordoncillo."
Piper factum Trelease, sp. nov.
Frutex gracilis glaber 2-metralis, internodiis floriferis primo
brevibus serius longis gracilibus; folia elliptica longiacuminata basi
inaequaliter acuta 10-12 cm. longa 5-6 cm. lata e dimidio inferiore
pinnatinervia, nervis 4+5, in sicco tenuia viridia subtus concolori-
granulosa, petiolo 8 mm. vel 10+3 mm. longo non vaginante;
spicae juveniles 8 mm. longae 1 mm. crassae acutae; bracteae
rotundo-subpeltatae.
Loreto: Yurimaguas, 135 meters, Killip & Smith 27588 (type
in U. S. Nat. Herb.).
Piper fagifolium Trelease, sp. nov.
Frutex essentialiter glaber plus minusve nodosus; folia lance-
olata acute acuminata basi inaequilaterali acuta 17-20 cm. longa
6-7.5 cm. lata undique pinnatinervia, nervis prominentibus glabratis
circa 16x2, subtus dense glanduloso-granulosa, petiolo 10+3 mm.
longo exalato; spicae 80 mm. longae 3 mm. crassae, pedunculo
gracili 10 mm. longo; bracteae rotundo-subpeltatae glabrae; baccae
subteretes vulgo apice depressae; stigmata 3 rotundata sessilia.
Loreto: Santa Rosa, in pasture, 155-210 meters, Llewelyn Williams
4782 (herb. Field Mus. No. 629,670, type) ; Killip & Smith 28743.
166 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper ferruginispicum Trelease, sp. nov.
Frutex metralis nervis folii subtus obscure minute crispo-pubes-
centibus exceptis glaber, internodiis floriferis modice crassis ut
videtur elongatis; folia obovato-elliptica obtusa basi inaequilatera-
liter cordata, sinu basilari, auricula maxima rotundata petiolo
longiore, 32 cm. longa 18 cm. lata e dimidio inferiore subpinnati-
nervia, nervis circa 6x2, nervis supplementariis in auricula
dispositis, in sicco tenuia, petiolo 5 cm. longo alato; spicae
rubro-brunneae 100 mm. longae 4 mm. crassae, pedunculo 40 mm.
longo; bracteae subcrescenticae.
Loreto: Mishuyacu near Iquitos, 100 meters, in forest, G. King
1421 (herb. Field Mus. No. 627,683, type).
Piper formosum (Miq.) C. DC. in DC. Prodr. 16, pt. 1: 286.
1869. Artanthe formosa Miq. Syst. 386. 1844; 111. Piper, pi. 69.
A shrub, somewhat tomentulose-velvety except for the glabrous
upper surface of the leaves, these ovate, acuminate, cordate, 13-15
cm. wide, 27 cm. long, submultiple-nerved from the lower half,
the nerves 6-7x2; petiole 3 cm. long, winged; young spikes 40
mm. long, 2-3 mm. thick, on a short peduncle.
Huanuco: Casapi, Mathews 1697, type. Junin: Chanchamayo
Valley, 1,200-1,800 meters, Schunke 93, 284, 286, 299, 411, 488.
Above San Ramon, Killip & Smith 24628. La Merced, Killip
& Smith 23784. Huacapistana, Killip & Smith 29187. Yapas,
Killip & Smith 25450.
Piper fortalezanum Trelease, sp. nov.
Frutex glaber, internodiis floriferis modice gracilibus atque
elongatis; folia elliptica sensim subacuminata basi fere aequila-
teraliter acuta vel uno latere breviora, 17-20 cm. longa 6.5-7.5
cm. lata undique pinnatinervia, nervis circa 8-10x2, in sicco tenuia
non lepidota, petiolo 10 mm. vel 5+5 mm. longo alato; spicae 80
mm. longae 3 mm. crassae, pedunculo 10 mm. longo; bracteae
rotundo-subpeltatae ciliatae; baccae depresso-ellipsoideae, stigmati-
bus 3 parvis sessilibus.
Loreto: Fortaleza, in forest, 155-210 meters, Llewelyn Williams
4351 (herb. Field Mus. No. 629,586, type). Puerto Yessup, Killip
& Smith 26281.
Piper franciscanum Trelease, sp. nov.
Frutex petiolo et nervis folii subtus crispo-puberulis exceptis
glaber; folia rotundato-ovata subacuminata basi truncato-cordulata
FLORA OF PERU < 167
17 cm. longa 11 cm. lata e dimidio inferiore vel paullo altius
submultiplinervia, nervis 5x2, petiolo 2 cm. longo, alls latis
fragilibus.
Brazil: Seringal Sao Francisco, Rio Acre, Ule 9278 (type), 9283.
Probably also in adjacent Peru.
Piper fulvescenticaule Trelease, sp. nov.
Frutex 2-3-metralis nodosus, internodiis superioribus brevibus
gracilibus griseo-velutinis serius rubescentibus; folia oblanceolata
caudata, basi angustata rotundata vel cordulata, uno latere breviore,
11-16 cm. longa 4-6 cm. lata e dimidio inferiore pinnatinervia,
nervis circa 6x2, subtus adpresso-pubescentia, petiolo vix 2+2
mm. longo; spicae 60 mm. longae 2 mm. crassae apicatae; bracteae
rotundo-subpeltatae.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 285? '4
(type in U. S. Nat. Herb.).
Piper fuscispicum Trelease, sp. nov.
Frutex 1.5 m. altus, internodiis floriferis modice gracilibus
breviusculis crispo-pubescentibus vel tomentulosis; folia lanceolata
acuminata basi acuta uno latere vulgo breviore, 19-22 cm. longa
7-8 cm. lata e dimidio inferiore pinnatinervia, nervis circa 6x2,
subtus plus minusve crispo-pubescentia in statu adulto rugosa;
petiolo 15+3 mm. vel 18 mm. longo tomentuloso; spicae 90 mm.
longae 3-4 mm. latae obtusae, primo brunneo-rubrae serius brun-
neae, pedunculo 5 mm. longo glabrato; bracteae rotundo-sub-
peltatae.
Loreto: Pongo de Manseriche, 280 meters, Ynes Mexia 6352a
(type in herb. Univ. Calif.).
Piper gilvescens Trelease, sp. nov.
Frutex 2-metralis glaber glanduloso-granulosus, internodiis flori-
feris brevibus gracilibus; folia lanceolato-elliptica vel ovato-ellip-
tica anguste falcato-acuminata, basi subaequilateraliter acuta et ad
petiolum decurrentia, 14-16 cm. longa 6-7 cm. lata e 2 tertiis inferi-
oribus pinnatinervia, nervis circa 5x2, in sicco tenuia viridia,
petiolo 10 mm. longo; spicae erectae 75 mm. longae 3 mm. crassae
mucronatae in sicco rubescentes, pedunculo 8 mm. longo; bracteae
rotundo-subpeltatae parvae.
Junin: Cahuapanas, 340 meters, Killip & Smith 26784 (type in
U. S. Nat. Herb.).
168 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper gilvibaccum Trelease, sp. nov.
Frutex glaber, internodiis floriferis subgracilibus brevibus; folia
lanceolata subfalcato-acuminata basi acuta 8-13 cm. longa 3-4 cm.
lata ubique pinnatinervia, nervis profunde impressis circa 8x2, in
sicco firma sublucida, petiolo 5 mm. longo late alato; spicae 20-35
mm. longae 5 mm. crassae, pedunculo 3 mm. longo; baccae depresso-
globosae rubrae; stigmata 3 parva sessilia.
Loreto: La Victoria on the Amazon River, in forest, Llewelyn
Williams 2754 (herb. Field Mus. No. 616,894, type).
Piper glaberrimicaule Trelease, sp. nov.
Frutex nodosus 3-4-metralis, internodiis floriferis brevibus graci-
libus glabris in statu adulto copiose pallido-granulosis; folia lanceo-
lata interdum falcata acute acuminata basi inaequilateralia saltern
uno latere rotundata, 12-14 cm. longa 4-5 cm. lata e dimidio inferi-
ore pinnatinervia, nervis circa 5x2, in sicco tenuia viridia subrugosa
supra scabrida, subtus ad nervos incurvo-pubescentia, petiolo circa
3+3 mm. longo non vaginante; spicae U-formes 80 mm. longae 3
mm. crassae obtusae, pedunculo 5 mm. longo glabro; bracteae
rotundo-subpeltatae.
Ayacucho: Estrella, 500 meters, Killip & Smith 22624 (type in
U. S. Nat. Herb.).
Piper glabribaccum Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis modice brevibus et graci-
libus fulvo-villosis pallido-granulosis; folia ovata sensim acuminata
basi breviter cordata majuscula 20-33 cm. longa 11-13 cm. lata e
dimidio inferiore submultiplinervia, nervis circa 5x2, subtus ad nervos
et supra transiente versus basin araneoso-villosa, petiolo 1 cm. longo
fulvo-villoso alato; spicae oppositifoliae 110 mm. longae 7 mm.
crassae, pedunculo 1 cm. longo brunneo-villoso ; bracteae rotundo-
subpeltatae villosae, floribus sessilibus perfectis; ovarium glabrum
obovoideum in stylum aequilongum angustatum; stigmata 3
patentia.
Junin: La Merced, 600 meters, edge of forest, Macbride 5335
(herb. Field Mus. No. 536,362, type).
Piper glabrius (Miq.) Trelease, sp. nov. Artanthe Poeppigii var.
glabrior Miq. in herb. Kew.
Ramuli laxe longe velutino-pilosi pallido-granulosi ; folia elliptica
vel ovata acuminata basi fere aequilateraliter obtusa 14-17 cm.
longa 6-8 cm. lata e dimidio inferiore pinnatinervia, nervis 5x2,
FLORA OF PERU 169
supra granuloso-scabra, subtus ad nervos molliter patenti-pilosa,
petiolo 10-15 mm. longo; spicae 100 mm. longae 2-3 mm. crassae,
pedunculo 10 mm. longo.
Type from Peru, Mathews 1705 (type in herb. Kew.).
Piper gracile R. & P. Fl. 1: 35. pi. 59. 1798. Artanthe gracilis
Miq. Syst. 448. 1844.
A slender, hirsute shrub; leaves lanceolate, acuminate, acute at
the base, 8-10 cm. long, 3-4 cm. wide, pinnately nerved from below
the upper third, rugose, becoming scabrous above from the bases of
fallen hairs; petiole 10 mm. long; spikes 40-60 mm. long, 3-4 mm.
thick; peduncle 10 mm. long; bracts triangular-subpeltate; berries
round; stigmas 3, linear, sessile.
Huanuco: Chinchao, Ruiz & Pawn, type. Amazonas: Chacha-
poyas, Mathews 1695 (sometimes labeled as from San Carlos and
Casapi) .
Piper granulosum R. & P. Fl. 1: 38. pi. 64. 1798. Artanthe
granulosa Miq. Syst. 435. 1844.
A shrub; twigs very transiently puberulous or glabrous, pale-
granular; leaves lance-oblong, more or less acuminate, the relatively
broad base cordulate and slightly shorter on one side, 10-15 cm.
long, 3-4 cm. wide, rugose, granular-scabrous above, slightly punc-
tulate beneath, with upcurved-pubescent nerves; petiole barely
3+2 mm. long; spikes curved, 75 mm. long, 3 mm. thick; peduncle
10 mm. long, glabrous; bracts subpeltate, orange-bordered; berries
terete, or angular by pressure; stigmas sessile.
Huanuco: Pillao, Ruiz & Pavon, type. Junin: La Merced,
Killip & Smith 23376, 23377, 23681. San Martin, Tarapoto, edge
of river, Williams 5416. "Cordoncillo."
Piper granuligerum Trelease, sp. nov.
Frutex nodosus, internodiis floriferis gracilibus breviusculis in
sicco purpurascentibus sparse transiente molliter pilosis plus minusve
lepidoto-punctatis; folia elliptico-lanceolata sensim anguste acumin-
ata basi subinaequilaterali subacuta 18-20 cm. longa 7 cm. lata e
dimidio inferiore pinnatinervia, nervis circa 7x2, transiente molliter
pubescentia serius minute lepidoto-granulosa ciliata, nervis subtus
sparse subvillosis, petiolo 10 mm. longo ciliato; spicae curvae vel
fere rectae 130 mm. longae 3-4 mm. latae obtusae, pedunculo vix
10 mm. longo glabrato; bracteae rotundo- vel triangulari-subpeltatae
ciliatae.
170 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Loreto: San Antonio, Alto Rio Itaya, 145 meters, Llewelyn
Williams 3459 (herb. Field Mus. No. 622,875, type). "Cordoncillo."
Piper gratum Trelease, sp. nov.
Frutex 1-metralis, internodiis floriferis modice brevibus gracilibus
plus minusve sparse rigido-villosis; folia lanceolata vel lanceolato-
elliptica acuminata basi subinaequilaterali rotundata vel cordulata
12-14 cm. longa 4-6 cm. lata undique vel in foliis latissimis tantum e
dimidio inferiore pinnatinervia, nervis 5-6x2, subtus praesertim ad
nervos laxe rigido-villosa, petiolo 10 mm. longo piloso alato; spicae
15 mm. longae 3 mm. crassae, pedunculo refracto 5 mm. longo
glabrato; bracteae subcucullatae; ovarium mucronatum, stigmati-
bus ad mucronem insertis.
Loreto: Soledad, 110 meters, Killip & Smith 29785 (type in
U. S. Nat. Herb.).
Piper gratum var. ingratum Trelease, var. nov.
Folia anguste lanceolata ad subelliptica 12-14 cm. longa 4-5 cm.
lata; spicae pendulae in statu florifero 15-20 mm. longae 2 mm.
crassae; ovaria conica ut in speciebus subgeneris Enckea protrusa.
Loreto: Mishuyacu near Iquitos, 100 meters, G. Klug 206 (type
in U. S. Nat. Herb.).
Piper griseocaule Trelease, sp. nov.
Frutex gracilis 1.5 m. altus nodosus, ramis viridibus laevibus
dense microscopice adpresse griseo-puberulis; folia oblonga subcau-
data basi paullo inaequaliter obtusa mediocria 15-17 cm. longa 4.5-
5.5 cm. lata e tertio vel dimidio inferiore pinnatinervia, nervis circa
4x2, lucida subtus ad nervos plus minusve pubescentia, petiolo
brevi 5+2 mm. longo microscopice adpresso-puberulo basi alato;
spicae oppositifoliae fere rectae 70-90 mm. longae 3 mm. crassae,
pedunculo 4-5 mm. longo obscure canescente; bracteae rotundo-
subpeltatae ciliolatae, floribus sessilibus perfectis; stigmata 3 minuta
sessilia; baccae parvae breviter cylindraceae apice concavae.
Junin: La Merced, 1,200 meters, forested slope, Macbride 5723
(herb. Field Mus. No. 536,755, type), 5595.
Piper griseovenosum Trelease, sp. nov.
Arbor 4-5-metralis, ramulis crispe brunneo-villosis; folia obovata
obtusa basi inaequilateralia, latere longiore auriculate producto et
sinum lateralem claudente petiolo villoso longiore, 32 cm. longa
FLORA OF PERU 171
18 cm. lata e dimidio inferiore subpinnatinervia, nervis circa 5x2, in
sicco tenuia, supra glabra, costa et nervorum basibus subtus villosis,
nervis superne ut venis griseo-puberulis, petiolo 8 cm. longo alato;
spicae juveniles 60 mm. longae 3 mm. crassae; bracteae rotundo-
subpeltatae glabrae.
Loreto: Puerto Arturo, 135 meters, Killip & Smith 27803 (type
in U. S. Nat. Herb.). Santa Rosa, Killip & Smith 28915.
Piper griseum C. DC. in DC. Prodr. 16, pt. 1: 281. 1869.
A gray-pubescent shrub; twigs reflexed-appressed-pubescent;
leaves somewhat inequilaterally lanceolate, acuminate, subcordu-
late at the narrowed base, 11-12 cm. long, 4-4.5 cm. wide, pinnately
nerved from below the upper third, the nerves 5x2, appressed-
canescent on both sides; petiole scarcely 5 mm. long; spikes curved,
70 mm. long, 2 mm. thick; peduncle 15-20 mm. long, subhispid;
bracts round-subpeltate, ciliolate.
Huanuco: Cochero, Poeppig 1737, D1737, type.
Piper heptandrum (Miq.) C. DC. in DC. Prodr. 16, pt. 1:
244. 1869. Enckea heptandra Miq. Syst. 350. 1844; 111. Piper, pi. 64.
A glabrous shrub; leaves elliptic-suboblong, acuminate, acute-
based, 20 cm. long, 9 cm. wide, 5- or obscurely 7-nerved; petiole 2
cm. long; spikes 50-80 mm. long, 3 mm. thick; peduncle 10-15 mm.
long; bracts spatulate; stamens (abnormally?) 7 instead of 6; ovary
ovoid; stigmas 3, sessile. Neg. 8415.
Type from mountains of Peru, Ruiz. Without locality, Ruiz &
Pavon 10-20, 10-19.
Piper heptandrum var. crassispicum Trelease, var. nov.
Spicae 40-50 mm. longae 5 mm. crassae; baccae subpuberulae.
Type from Peru, Pavon. Loreto: Ucayali River, Tessmann 3165.
Piper Herrerae Trelease, sp. nov.
Frutex gracilis glaber, internodiis superioribus gracillimis brevi-
bus; folia elliptica vel lanceolato-elliptica longiacuminata basi acuta
9-10 cm. longa 3-4 cm. lata 3-nervia opaca firmo-papyracea plus
minus ve venosa, petiolo 5 mm. longo; spicae semimaturae 30 mm.
longae 4 mm. crassae, pedunculo 5 mm. longo; ovarium ovoideum,
stigmatibus brevibus sessilibus.
Cuzco: Capana, 3,700 meters, January, 1926, FortunatoL. Herrera
(herb. N. Y. Bot. Gard., type).
172 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper heterocarpum Trelease, sp. nov.
Frutex cinereo-villosus 2-metralis nodosus in sicco nigrescens,
internodiis floriferis brevibus modice gracilibus; folia lanceolato-
elliptica acuminata basi inaequilaterali obtusa, uno latere vel inter-
dum ambobus cordulatis, 15-16 cm. longa 5.5-6 cm. lata e dimidio
inferiore pinnatinervia, nervis 4-f5, petiolo 5+5 mm. longo; spicae
55 mm. longae 4 mm. crassae, pedunculo filiformi 25 mm. longo;
bracteae crescentico-subpeltatae; baccae granulosae rotundatae vel
varie polygonae propter pressum submucronato-truncatae, stigmati-
bus sessilibus.
Loreto: Yurimaguas to Balsapuerto, 135-150 meters, Killip &
Smith 28327 (type in U. S. Nat. Herb.).
Piper heterophyllum R. & P. Fl. 1: 34. pi. 56. 1798. Schilleria
heterophylla Kunth, Linnaea 13: 688. 1839. Artanthe heterophylla
Miq. Syst. 502. 1844.
A glabrous, glandular-granular shrub; leaves elliptic, obscurely
short-acuminate, subacute at the base, 11-16 cm. long, 5.5-9 cm.
wide, pinnately nerved throughout; petiole 2 cm. long; spikes
curved, 60-75 mm. long (or figured as twice as long), 3 mm.
thick; peduncle 10 mm. long; bracts roundish-subpeltate, with pale
ciliate border; stigmas sessile. Neg. 8414.
Huanuco: Chacahuassi, Pillao, Chinchao, and Muna, Ruiz &
Pavon, type.
Piper Hookerianum (Miq.) C. DC. Prodr. 16, pt. 1: 272. 1869.
Artanthe Hookeriana Miq. Lond. Journ. Bot. 4: 446. 1866.
A glabrous shrub; leaves ovate, acuminate, slightly inequilaterally
rounded at the base, 16 cm. long, 10 cm. wide, pinnately nerved
from somewhat less than the lower half, the nerves 5-6x2; petiole
20+2 mm. long, winged; spikes 25 mm. long, 5 mm. thick; peduncle
10 mm. long; bracts half-round, subpeltate, ciliolate; berries
depressed-globose, with a short thick style; stigmas spreading.
Amazonas: Chachapoyas, Mathews 3222, type.
Piper huacachianum Trelease, sp. nov.
Frutex 2.5 m. altus nodosus glaber; folia lanceolata utrinque
acute protracta 12-17 cm. longa 3-4.5 cm. lata undique pinnatinervia,
nervis circa 12x2, petiolo 10-15 mm. longo gracili exalato; spicae
plus minusve curvae in statu fructifero 40-50 mm. longae 5 mm.
crassae apicatae, pedunculo 7 mm. longo; bracteae rotundo-subpel-
tatae; baccae globosae, stigmatibus obscuris sessilibus.
FLORA OF PERU 173
Huanuco: Huacachi near Mufia, 1,950 meters, Macbride 4167
(herb. Field Mus. No. 535,230, type).
Piper huacapistanum Trelease, sp. nov.
Frutex, internodiis floriferis gracilibus breviusculis dense hirtellis;
folia ovata acute acuminata basi rotundata sed basi infima acute
contracta, 15-16 cm. longa 8-9 cm. lata e dimidio inferiore multipli-
nervia, nervis 5x2, in sicco tenuia viridia, supra circa basin plus
minusve velutina, subtus praesertim ad nervos crispo-velutina,
petiolo 15 mm. longo hirtello alato; spicae oppositifoliae, juveniles
40 mm. longae 3 mm. crassae, pedunculo vix 5 mm. longo velutino;
bracteae rotundo-subpeltatae ciliolatae.
Junin: Huacapistana, Stevens 77 (type), 79.
Piper huallaganum Trelease, sp. nov.
Frutex laxe ramosus metralis plus minusve nodosus, internodiis
floriferis gracilibus elongatis granuloso-asperatis et primum ad-
presso-hispidis; folia elliptico-ovata acuminata basi inaequali rotun-
data vel cordulata, subparva 10-15 cm. longa 5-7 cm. lata e dimidio
inferiore pinnatinervia, nervis circa 5x2, in sicco tenuia opaca,
subtus ad nervos adpresso-hispida, petiolo 10+2 mm. longo adpresso-
hispido basi alato; spicae oppositifoliae 70-90 mm. longae 3 mm.
crassae, pedunculo gracili 15 mm. longo adpresso-hispido; bracteae
triangulari-subpeltatae glabrae, floribus sessilibus perfectis ; stigmata
3 minuta sessilia.
Huanuco: Huallaga Canyon below Rio Santo Domingo, 1,200
meters, dense forest, Macbride 4235 (herb. Field Mus. No. 535,314,
type).
Piper huanucanum Trelease, sp. nov. P. aequale Auct.
Frutex (?) glaber vel transiente papilloso-puberulus; folia elliptica
acuminata basi acuta 10-20 cm. longa 6-9 cm. lata subpinnati-
nervia, nervis basilaribus 2-3 paribus, pare altero bene infra medium
inserto, pare alterno bene supra medium inserto, petiolo vix 10 mm.
longo; spicae juveniles 60 mm. longae 3 mm. crassae, pedunculo
10 mm. longo.
Huanuco: Huanuco mountains, Haenke (in herb. Prague, Vienna,
and Berlin).
Piper huantanum Trelease, sp. nov.
Arbor 3-5-metralis nodosa nervis folii subtus puberulis exceptis
glabra, internodiis floriferis longis gracilibus; folia elliptica vel
174 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
elliptico-oblonga acuminata basi inaequilaterali acuta 15-16 cm.
longa 6-7 cm. lata e dimidio inferiore pinnatinervia, nervis 5x2,
petiolo 10+3-4 mm. longo basi tantum vaginante; spicae 100 mm.
longae 4 mm. crassae, pedunculo 5 mm. longo; bracteae rotundo-
subpeltatae ciliolatae; baccae cylindraceae, stigmatibus parvis
sessilibus.
Ayacucho: Estrella, between Huanta and Rio Apurimac, 500
meters, Kittip & Smith 23074 (type in U. S. Nat. Herb.). Junin:
Rio Perene", Kittip & Smith 25193.
Piper huantanum var. elongatum Trelease, var. nov.
Magis nodosus, internodiis brevioribus; folia lanceolato-elliptica
vel oblonga 20 cm. longa 7 cm. lata, petiolo 10+5 mm. longo primo
alato; spicae juveniles 30 mm. longae 2 mm. crassae obtusae flaves-
centi-albae, pedunculo 10 mm. longo.
Junin: Near Perene* Bridge, Rio Paucartambo Valley, 700 meters,
Kittip & Smith 25304 (type in U. S. Nat. Herb.).
Piper ictum Trelease, sp. nov.
Frutex(?), ramis crispo-subtomentosis; folia rugoso-lacunosa
lanceolato-ovata sensim acuta basi acuta 12 cm. longa 4 cm. lata e 2
tertiis inferioribus pinnatinervia, nervis circa 6x2, supra molliter
pilosa et scabra, subtus dense flavo-pilosa, petiolo 1 cm. longo;
spicae 70 mm. longae 5 mm. crassae obtusae, pedunculo 10 mm. longo
hispido; bracteae triangulari-subpeltatae ciliatae; baccae globosae,
stigmatibus 3 sessilibus.
Type collected in Peru, without definite locality, Mathews 1711
or 1712 (herb. Kew.), 1716 (herb. Berol.).
Piper illautum Trelease, sp. nov.
Frutex(?); folia oblanceolata acuta basi cuneata 15 cm. longa
5 cm. lata ubique pinnatinervia tantum supra glabra, petiolo vix
10 mm. longo; spicae curvae 40 mm. longae 5 mm. crassae; baccae
subglobosae, stylo brevi; stigmata 3 paten tia.
Known only from a fragmentary specimen in Herb. Paris,
ascribed to Peru, Gay 1003.
Piper immite Trelease, sp. nov.
Frutex 3-5-metralis nodosus, internodiis floriferis brevibus graci-
libus glabris vel obscure evanescente puberulis; folia ovata acumin-
ata basi inaequilaterali rotundata et paullo ad petiolum angustata,
12-16 cm. longa 6-10 cm. lata e dimidio inferiore submultiplinervia,
FLORA OF PERU . : 175
nervis circa 5x2, in sicco firme papyracea, subtus ad nervos incon-
spicue breviter incurvo-pilosa, inter nervos sparse substellato-
pubescentia vel glabrata, petiolo 10-15-1-2 mm. longo varie vaginante
glabrato; spicae 80-140 mm. longae 3 mm. crassae, pedunculo vix
5 mm. longo subpuberulo; bracteae triangulari-subpeltatae; baccae
teretes, stylo gracili; stigmata 3 gracilia.
Junin: La Merced, 700 meters, Killip & Smith 23524 (type in
U. S. Nat. Herb.), 23437, 23466, 24074-
Piper immutatum Trelease, sp. nov.
Frutex subfistulosus 3-metralis, internodiis floriferis crassiusculis
brevibus dense rubescenti-villosis; folia elliptica plus minusve acumin-
ata basi inaequilaterali obtusa latere longiore omnino ad petiolum
adnato, 20 cm. longa 8-10 cm. lata e dimidio inferiore pinnatinervia,
nervis 5-6x2, in sicco tenuia brunnescentia, supra sparse molliter
pilosa, subtus ad nervos pilis rigidioribus conspersa, petiolo 3 mm.
uno latere longo villoso; spicae 80 mm. longae 3-4 mm. crassae,
pedunculo 3 mm. longo villoso; bracteae rotundo-subpeltatae
ciliolatae.
Loreto: Balsapuerto, 350-550 meters, Killip & Smith 28536
(type in U. S. Nat. Herb.).
Piper incitum Trelease, sp. nov.
Frutex nodosus glaber 2-3-metralis, internodiis floriferis gracilibus
brevibus; folia lanceolata vel elliptica anguste acuminata basi
aequilaterali acuta 9-12 cm. longa 3-5 cm. lata e dimidio inferiore
pinnatinervia, nervis 3-4x2 infimis prope basin nascentibus, in
sicco caerulescenti-viridia, petiolo 5-10 mm. longo exalato; spicae
fructiferae 50 mm. longae 5 mm. crassae obtusae, pedunculo gracili
10 mm. longo; bracteae crescenticae; baccae subglobosae truncatae,
stigmatibus sessilibus.
Junin: Huacapistana, 1,800 meters, Killip & Smith 24304 (type
in U. S. Nat. Herb.).
Piper incomptum Trelease, sp. nov.
Arbor, internodiis floriferis gracilibus brevibus albido-villosis
serius glabrescentibus; folia lanceolata acuminata basi subinae-
quilaterali acuta 9-11 cm. longa 3-4.5 cm. lata e 2 tertiis inferioribus
pinnatinervia, nervis 6x2, supra pilis basi bulbosis asperata, subtus
ad nervos albido-villosa; petiolo 3 mm. longo albido- villoso; spicae
55 mm. longae 3 mm. crassae, pedunculo 10 mm. longo scabrido.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 1793
(herb. Field Mus. No. 625,555, type).
176 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper indecorum Kunth, Linnaea 13: 722. 1839. Schilleria
indecora Kunth, I.e. Artanthe polyneura Miq. Syst. 504. 1844.
P. maynasianum C. DC. in DC. Prodr. 16, pt. 1: 262. 1869.
A shrub, drying yellowish; internodes rather thick and short,
glabrescent; leaves elliptic-lanceolate, acuminate, acute-based, 14-19
cm. long, 6-7 cm. wide, pinnately nerved throughout, subvillous on
the nerves beneath; petiole scarcely 10 mm. long, subvillous; spikes
100 mm. long, 4 mm. thick or larger, gently curved; peduncle 10
mm. long, glabrescent; bracts subtriangular-subpeltate; berries short-
cylindric, depressed at the top, with a short mucro; stigmas 3, sessile.
Huanuco: Cochero, Poeppig 2353B, 1463, 143. Junin: Pangoa,
Matthews 11 44'
Piper indianonum Trelease, sp. nov.
Frutex 5-metralis, internodiis brevibus subgracilibus glabris
granulosis; folia late lanceolata vel lanceolato-elliptica sublonge
acuminata basi subacuta, uno latere magis rotundato, 12-15 cm.
longa 4-5.5 cm. lata e dimidio inferiore vel paullo altius pinnati-
nervia, nervis 5x2, subtus adpresso-pubescentia, petiolo 5 mm. longo;
spicae strictae 90-100 mm. longae 2 mm. crassae, pedunculo 10 mm.
longo; bracteae rotundo-subpeltatae; baccae depresso-ellipsoideae,
stigmatibus 3 sessilibus.
Loreto: Fundo Indiano, 110 meters, Ynes Mexia 6396 (type in
herb. Univ. Calif.), also No. 6441 with less granular twigs.
Piper indiciflexum Trelease, sp. nov.
Arbor 4-5-metralis, internodiis crassis longis glabris in sicco
nigrescentibus; folia rotundo-ovata breviter acuminata basi sub-
oblique cordata 26 cm. longa 20 cm. lata e dimidio inferiore submul-
tiplinervia, nervis circa 7x2 e latere proximo ramosis, in sicco tenuia,
obscure rugulosa, supra sparse molliter brevipubescentia, subtus
densius crispo-pubescentia, petiolo 5 cm. longo glabro tantum prope
basin vaginante; spicae 60 mm. longae 6 mm. crassae curvae, petiolo
2 cm. longo glabro; bracteae triangulari-subpeltatae; baccae rotun-
datae, stigmatibus sessilibus.
Junin: Huacapistana, 1,800-2,400 meters, Killip & Smith 24507
(type in U. S. Nat. Herb.).
Piper interitum Trelease, sp. nov.
Arbor 3-5-metralis nodosa glabra, internodiis modice gracilibus
longis; folia late elliptica acuminata basi inaequilaterali subauricu-
lata 17 cm. longa 9-10 cm. lata ubique pinnatinervia, nervis circa
FLORA OF PERU 177
10x2, petiolo circa 15 mm. longo alis latis obscurato; spicae pendulae
25 mm. longae 12 mm. crassae, pedunculo 5 mm. longo; bracteae
rotundo- vel triangulari-subpeltatae; baccae globosae submucrona-
tae, stigmatibus 3.
Junin: Santa Rosa, Pichis Trail, 625-900 meters, Killip & Smith
26150 (type in U. S. Nat. Herb.).
Piper intonsum Trelease, sp. nov.
Frutex 4-metralis, internodiis floriferis breviusculis gracilibus in
sicco nigrescentibus persistente densiuscule griseo-velutinis; folia
elliptica subabrupte protracto-acuminata submucronatae vulgo basi
acuta minora 10-12 cm. longa 5-7 cm. lata palmatim 5-6-nervia,
supra prope basin et ad nervos atque subtus ubique minute crispo-
pubescentia, petiolo breviusculo 10 mm. longo tantum prope basin
alato sordide crispo-pubescente; spicae oppositifoliae modice graciles
et elongatae, 60 mm. longae 2-4 mm. crassae, pedunculo vix 10 mm.
longo plus minusve crispo-pubescente vel velutino; bracteae con-
cavae glabratae, floribus sessilibus perfectis; stigmata 3 conniventia;
baccae ovoideae subattenuatae.
Junin: La Merced, sandy valley floor, 600 meters, Macbride 5445
(herb. Field Mus. No. 536,479, type). San Martin: Rumizapa,
Tarapoto, 360-900 meters, Williams 6829. "Cordoncillo."
Piper iquitosense Trelease, sp. nov.
Frutex nodosus nervis folii subtus velutinis exceptis glaber,
internodiis floriferis brevibus subgracilibus in statu adulto tubercu-
latis; folia suboblongo-elliptica acute acuminata basi rotundata, uno
latere multo breviore, 20 cm. longa 7-8 cm. lata ubique pinnati-
nervia, petiolo 5-10+10 mm. longo alato; spicae 120 mm. longae
3 mm. crassae, pedunculo gracili 10 mm. longo; bracteae triangulari-
subpeltatae brunnescentes ciliatae; baccae depresso-ellipsoideae,
stigmatibus 3 minutis sessilibus.
Loreto: Iquitos, Llewelyn Williams 11+32 (herb. Field Mus. No.
617,448, type). Punchana near Iquitos, Williams 1332. Iquitos,
Killip & Smith 26899. Yurimaguas, Killip & Smith 28181.
Junin: Puerto Yessup, Killip & Smith 26275. "Cordoncillo. "
Piper itayanum Trelease, sp. nov.
Frutex glaber minute nigro-granulosus, internodiis floriferis brevi-
usculis gracilibus; folia lanceolato-ovata breviter acuminata saltern
basi infima acuta, uno latere breviora, 24-29 cm. longa 10-11 cm.
178 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
lata ubique pinnatinervia, nervis 15x2, petiolo vix 5+3 mm. longo
exalato; spicae 60-80 mm. longae 10 mm. crassae, pedunculo 10
mm. longo; bracteae triangulari-subpeltatae; baccae subteretes vel
vulgo acute 3-4-gonae; stigmata 3 sessilia.
Loreto: Paraiso, Alto Rio Itaya, in forest, 145 meters, Llewelyn
Williams 3320 (herb. Field Mus. No. 623,464, type). San Antonio,
Williams 3509. Soledad, Killip & Smith 29617. Yurimaguas to
Balsapuerto, Killip & Smith 28130. Iquitos, Killip & Smith 27245.
Puerto Bermudez, Killip & Smith 26453. Junin: Puerto Yessup,
400 meters, in forest, Killip & Smith 26260.
Piper itayanum var. antoniense Trelease, var. nov.
Folia subovato-elliptica 25-30 cm. longa 12-14 cm. lata; spicae
80-140 mm. longae 10 mm. crassae.
Loreto: San Antonio, 110 meters, Killip & Smith 29450 (type in
U. S. Nat. Herb.). Iquitos, Killip & Smith 27324, the spikes
mucronate.
Piper itayanum var. pichisense Trelease, var. nov.
Folia late elliptica 20-22 cm. longa 9-11 cm. lata; spicae 50-100
mm. longae 10 mm. crassae.
Junin: Cahuapanas, Rio Pichis, 340 meters, Killip & Smith 26738
(type in U. S. Nat. Herb.), 26725, 26750. Puerto Yessup, Killip
& Smith 26260.
Piper itayanum var. san-juanum Trelease, var. nov.
Folia magis elliptica 22-25 cm. longa 11 cm. lata, latere majore
basi magis obtuso, petiolo 2 cm. longo; spicae juveniles 95 mm.
longae 7 mm. crassae.
Loreto: San Juan, Iquitos, 120 meters, Llewelyn Wittiams 3720
(herb. Field Mus. No. 618,346, type).
Piper jaculifolium Trelease, sp. nov.
Frutex 3-metralis, pagina inferiore folii excepta glaber, inter-
nodiis gracilibus elongatis; folia lanceolata 21 cm. longa 7 cm. lata,
vel subovata et 10 cm. lata, protracto-acuminata, basi fere aequila-
terali acuta e quarto inferiore submultiplinervia, nervis circa 5x2,
subtus stellato-pubescentia, petiolo 15-20 mm. longo alato; spicae
juveniles 30 mm. longae 2-3 mm. crassae, pedunculo 10 mm. longo
paullo pubescente; bracteae triangulari-subpeltatae.
Junin: La Merced, 700 meters, Killip & Smith 24060 (type in
U. S. Nat. Herb.).
FLORA OF PERU 179
Piper Kanehiranum Trelease, sp. nov.
Frutex nodosus, internodiis floriferis brevibus crassiusculis vel-
utinis; folia lanceolata acuta basi cordata 12 cm. longa 4 cm. lata e
dimidio inferiore pinnatinervia, nervis circa 5x2, supra punctulata,
subtus ad nervos adpresso-pubescentia, in sicco tenuia conduplicata,
statu adulto minute rugosa, petiolo vix 5 mm. longo; spicae curvae
50 mm. longae 2-3 mm. crassae, pedunculo 5 mm. longo.
Huanuco: Pampayacu, Ryozo Kanehira 14 (herb. Field Mus. No.
580,066, type).
The leaves are said to be used medicinally.
Piper kimpitirikianum Trelease, sp. nov.
Frutex nodosus 3-4-metralis, internodiis floriferis gracilibus
brevibus crispe breviter griseo-villosis; folia lanceolata longe anguste
acuminata basi inaequilaterali acuta 10-13 cm. longa 4-5 cm. lata e
dimidio inferiore pinnatinervia, nervis circa 5x2, subtus ad nervos
breviter villosa, in sicco tenuia, petiolo 3+2 cm. longo breviter vill-
oso; spicae 95-120 mm. longae 2 mm. crassae obtusae, pedunculo
7 mm. longo plus minusve piloso; bracteae inconspicuae; baccae
rotundatae, stigmatibus 3 minutis sessilibus.
Ayacucho: Near Kimpitiriki, Rio Apurimac, 400 meters, Killip
& Smith 22847 (type in U. S. Nat. Herb.).
Piper Klugianum Trelease, sp. nov.
Frutex 1.5 m. altus, internodiis floriferis gracilibus modice brevi-
bus crispo-pilosis; folia lanceolata subacuminata basi fere aequilate-
rali acuta 20 cm. longa 5.5-7 cm. lata ubique pinnatinervia, nervis
10-12x2, vix nigro-punctulata (plus minusve transiente?), subtus
crispo-pilosa, in sicco tenuia, petiolo 15-20 mm. longo ad medium
vel altius alato crispo-piloso; spicae juveniles 25 mm. longae 3 mm.
crassae, pedunculo crasso 5 mm. longo glabrato; bracteae triangulari-
subpeltatae; ovarium ovoideum, stigmatibus 3 rotundatis sessilibus.
Loreto: Mishuyacu near Iquitos, 100 meters, in forest, G. Klug
1493 (herb. Field Mus. No. 627,496, type), 146.
The spikes are said to be red-brown.
Piper lagenaebaccum Trelease, sp. nov.
Frutex vel arbor 3-5-metralis, internodiis floriferis gracilibus
elongatis, primo pilis rigidulis albidis villosis; folia elliptica vel subo-
vato-lanceolata subacuminata basi obtusa, uno latere auriculato et
petiolum velante, 10-11 cm. longa 3-5 cm. lata e 2 tertiis inferioribus
180 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
vel fere undique pinnatinervia, nervis validis circa 5x2, supra sparse
subtus ad nervos densius rigido-villosa, rigido-ciliata, petiolo vix 5
mm. longo villoso; spicae pendulae 20 mm. longae 8 mm. crassae,
pedunculo 10 mm. longo pallido- villoso; bracteae crescenticae; baccae
lagenaeformes longiattenuatae, stigmatibus 3 apice styli insertis.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26249 (type
in U. S. Nat. Herb.), 26244- Santa Rosa, Killip & Smith
26175. Puerto Bermudez, Killip & Smith 26570. Loreto: Pebas,
Williams 1626.
Piper lamasense Trelease, sp. nov.
Frutex, internodiis floriferis brevibus gracilibus sparse pallido-
villosis; folia rotundo-elliptica vel subovata acuminata basi inaequila-
terali cordulata vel subcordata 12-15 cm. longa 6-9 cm. lata e dimi-
dio inferiore pinnatinervia, nervis 6x2, paullo rugosa, supra sparse
molliter pubescentia, subtus ad nervos crispe fulvo-pubescentia,
petiolo 10 mm. longo exalato, dense fulvo-pubescente; spicae 90 mm.
longae 3 mm. crassae, pedunculo gracili 10-15 mm. longo breviter
villoso; bracteae rotundo-subpeltatae ciliolatae; baccae subteretes
depresso-truncatae, stigmatibus 3 parvis sessilibus.
San Martin : Lamas near Tarapoto, 840 meters, Llewelyn Williams
6401 (herb. Field Mus. No. 623,619, type). "Cordoncillo."
Piper lanceaefolium HBK. Nov. Gen. & Sp. 1: 49. 1815.
Schilleria lanceae/olia Kunth, Linnaea 13: 703. 1839. Artanthe
lanceaefolia Miq. Syst. 433. 1844; 111. Piper, pi. 79.
A shrub with silky twigs; leaves lance-elliptic, acuminate, slightly
inequilaterally rounded at the base or cordulate, 15-17 cm. long,
4.5-6 cm. wide, pinnately nerved from the lower half, the nerves
about 8x2, deeply impressed above and salient beneath; petiole
5-|-2 mm. long, silky; spikes curved, 50 mm. long, 3 mm. thick;
peduncle 10 mm. long; berries triquetrous; stigmas 3, sessile.
Cajamarca: Venta, Jae"n de Bracamoros, Humbolclt in Herb.
Willdenow, No. 661. Loreto: Santa Rosa, Killip & Smith 28994-
Mainas, Poeppig 2289. Upper Amazon, Mathews 1704 or 1714-
Piper lanceolatum R. & P. Fl. 1: 36. pi. 61. 1798. Steffensia
evonymifolia Kunth, Linnaea 13 : 624. 1839. Artanthe Brunoniana
Miq. Syst. 509. 1844; 111. Piper, pi. 87.
A glabrous or obscurely and transiently puberulent shrub;
leaves lanceolate, gradually acute, acute-based, 8-13 cm. long, 2-3.5
cm. wide, pinnately nerved throughout; petiole 5-10 mm. long;
FLORA OF PERU 181
spikes 15-20 mm. long, 3 mm. thick, blunt; peduncle 5 mm. long;
bracts rounded-subpeltate; berries round; stigmas 3, small, sessile.
Huanuco: Chacahuassi, Ruiz & Pavon, type. Loreto(?):
Mainas(?), Poeppig 1434, 219.
Piper lanosicaule Trelease, sp. nov.
Frutex 3-metralis nodosus, internodiis floriferis gracillimis brevi-
bus pallide breviter lanoso-villosis; folia lanceolata anguste acuminata
basi plus minusve inaequilaterali et angustata obtusa, 9-13 cm.
longa 2.5-5 cm. lata e dimidio inferiore pinnatinervia supra pilis
minutis arete adpressis scabrida, subtus ad nervos incurvo-hirsuta,
petiolo 3+3 mm. longo sublanoso exalato; spicae 50 mm. longae 3
mm. crassae, pedunculo vix 10 mm. longo plus minusve piloso;
bracteae rotundo-peltatae.
Junin: Schunke Hacienda, San Ramon, 1,400-1,700 meters,
Killip & Smith 24723 (type in U. S. Nat. Herb.). Enenas, Killip &
Smith 25646.
Piper late-ovatum Trelease, sp. nov.
Frutex humilis nodosus ubique tomentulosus, costa folii subtus
villosa, internodiis floriferis brevibus modice gracilibus; folia late
ovata subabrupte acuminata basi rotundata vel subemarginata 20-
24 cm. longa 13-17 cm. lata e 2 tertiis inferioribus submultiplinervia,
nervis circa 7x2, in sicco tenuia brunnescentia, petiolo 2 cm. longo
exalato; spicae acutae, juveniles 60 mm. longae 3 mm. crassae,
pedunculo 5 mm. longo bicarinato; bracteae rotundo-subpeltatae
ciliolatae.
San Martin: San Roque, in forest, 1,350-1,500 meters, Llewelyn
Williams 7083 (herb. Field Mus. No. 623,091, type).
Piper laurinum R. & S. Syst. 1: 291. 1817. P. laurifolium
HBK. Nov. Gen. & Sp. 1: 47. 1815, non P. laurifolium Mill. 1758.
Schilleria laurifolia Kunth, Linnaea 13: 691. 1839. Artanthe lauri-
folia Miq. Syst. 520. 1844; 111. Piper, pi. 89.
A glabrous shrub; leaves elliptic or subovate, acuminate, rounded
at the base, 9.5-13 cm. long, 4-7 cm. wide, pinnately nerved from
the lower half or more, the nerves 4-5x2, drying coriaceous, glossy
above, paler beneath; petiole 10 mm. long; spikes 100 mm. long, 3
mm. thick, flexuous; peduncle 5 mm. long; berries subtriquetrous,
obpyramidal, somewhat pointed ; stigmas sessile.
La Libertad: Type collected between Chala and Trujillo, Hum-
boldt as No. 654 in Herb. Willdenow. Trujillo, Bonpland 161.
182 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper Lechlerianum C. DC. in DC. Prodr. 16, pt. 1: 269. 1869.
A shrub with loosely villous twigs; leaves ovate, acuminate,
inequilaterally cordulate, 20 cm. long, 8 cm. wide, pinnately nerved
from the lower half, the nerves about 6x2, white-granular-scabrous
above, loosely hirsute-villous beneath; petiole about 5+5 mm. long;
spikes 120 mm. long, 3 mm. thick; peduncle 10 mm. long.
Puno: Sangaban, Lechler 2438, type.
Piper Lehmannianum (Miq.) C. DC. in DC. Prodr. 16, pt. 1:
368. 1869. Artanthe Lehmanniana Miq. Lond. Journ. Bot. 4: 446.
1866. P. salicariaefolium var. Lehmannianum C. DC. op. cit. 317.
A slender shrub; twigs glabrous, pale-granular, zig-zag; leaves
lance-oblong or elliptic-oblong, acuminate, subobtuse at the base,
10-12 cm. long, 3-3.5 cm. wide, pinnately nerved from the lower half,
the nerves 4-5x2, appressed-pubescent beneath; petiole 5 mm.
long; spikes 2-3 mm. thick, 70 mm. long; peduncle 5-10 mm. long.
Huanuco: Casapi, Mathews 1694, type. Junin: Puerto Yessup,
Killip & Smith 26297. Loreto: Soledad, Killip & Smith 29551.
San Martin: Tarapoto, Williams 6742. "Cordoncillo."
Piper leticianum C. DC. Verh. Bot. Ver. Brandenb. 47:
104. 1905.
A yellow- villous shrub; leaves lance-oblong, subacute, acute-
based, 22 cm. long, 7 cm. wide, pinnately nerved throughout; petiole
2 cm. long; spikes curved, 20 mm. long, 5 mm. thick; peduncle 5
mm. long, curved; bracts crescentic-subpeltate, fringed; stigmas
sessile.
Loreto: Leticia, Vie 6216, type. La Victoria, in forest, Williams
3018.
Piper leticianum var. Tessmannii Trelease, var. nov.
Folia supra glabrescentia basi inaequilateralia, uno latere rotun-
dato; spicae 30 mm. longae.
Loreto: Mouth of Rio Santiago, Rio Maranon, Tessmann 3946
(type in herb. Berol.).
Piper leucofustum Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis brevibus gracilibus per-
sistente pilis arete retrorso-adpressis albidis; folia ovata obtuse vel
emarginate subacuminata basi inaequilaterali cordulata 9-10 cm.
longa 4-5 cm. lata e dimidio inferiore pinnatinervia, nervis 6x2,
FLORA OF PERU 183
basibus incrassatis pilorum caducorum mollium albido-scabra, subtus
ad nervos venasque conspicue molliter albido-pubescentia, petiolo
5+5 mm. longo adpresso-piloso basi vaginante; spicae 90 mm. longae
3 mm. crassae acutae, pedunculo vix 10 mm. longo retrorse albido-
piloso; bracteae parvae rotundo-subpeltatae ciliolatae.
Junin: La Merced, 70 meters, Killip & Smith 23736 (type in
U. S. Nat. Herb.).
Piper leucophaeocaule Trelease, sp. nov.
Frutex, internodiis gracilibus elongatis rigido-pilosis serius
scabris; folia elliptica acuminata basi inaequilaterali subobtusa
16-18 cm. longa 7-8 cm. lata e dimidio inferiore pinnatinervia,
nervis 6x2, utrinque dense breviter scabra, subtus ad nervos hispida,
petiolo 10+2 mm. longo; spicae juveniles graciles mucronatae,
pedunculo griseo.
Loreto: Lower Rio Nanay, Llewelyn Williams 301 (herb. Field
Mus. No. 603,748, type). Iquitos, Killip & Smith 27037, 27469.
Junin: Schunke Hacienda, San Ramon, Killip & Smith 24629.
Piper leucophaeum Trelease, sp. nov.
Frutex gracilis subgriseus, internodiis floriferis brevibus gracilibus
scabris; folia lanceolata vel subovata sensim acuminata basi obtusa
uno latere paullo minore 8-9 cm. longa 3 cm. lata e dimidio inferiore
pinnatinervia, nervis 4x2, supra minute pubescentia serius granulosa
et lepidota, subtus ad nervos breviter strigosa, petiolo 3+2 mm.
longo exalato; spicae juveniles 35 mm. longae 2 mm. crassae acutatae,
pedunculo 3 mm. longo scabrido; bracteae rotundo-subpeltatae
ciliolatae.
San Martin: Tarapoto, in forest, 750 meters, Llewelyn Williams
6086 (herb. Field Mus. No. 618,084, type).
Piper ligatispicum Trelease, sp. nov.
Arbor 4-5-metralis foliis subtus obscure puberulis exceptis
glabra, internodiis floriferis subgracilibus longis; folia subelliptica
obtuse breviter acuminata basi inaequilaterali cordata, latere longiore
petiolo duplo breviore, 30 cm. longa 17 cm. lata e dimidio inferiore
vel paullo altius subpinnatinervia, nervis circa 7+8, subtus versus
apicem basinque obscure puberula, petiolo 5+1 cm. longo alato;
spicae 400 mm. longae 5 mm. crassae vel in statu fructifero majores,
ad pseudoverticilla bractearum spiraliter ligatae, pedunculo 2 cm.
longo; bracteae rotundo-subpeltatae brunneae ciliatae; baccae
rotundatae vel subcubicae brunneae glabrae, stigmatibus sessilibus.
184 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Loreto: Yurimaguas, 135 meters, Killip & Smith 28183 (type in
U. S. Nat. Herb.).
Piper lineatum R. & P. Fl. 1: 35. pi. 60. 1798. Schilleria lineata
Kunth, Linnaea 13: 704. 1839. Artanthe lineata Miq. Syst. 459.
1844; 111. Piper, pi. 80. P. limatum Auct., by error.
A nodose shrub ; axes and nerves beneath subvillous, more or less
glabrescent in age; leaves lance-elliptic, acuminate, inequilaterally
subacute at the base, 9-11 cm. long, 3-4 cm. wide, pinnately nerved
from the lower half, the nerves, 5-6x2, lineately impressed above
and salient beneath, chartaceous, rugose, granular-roughened;
petiole 10 mm. long; spikes curved, about 100 mm. long, 3 mm. thick;
peduncle 10-15 mm. long; bracts triangular-subpeltate. Neg. 8417.
Huanuco: Chinchao, Rio Huanuco, and Panao, Ruiz & Pawn,
type. Huancabamba, Esposto 335a, 336. Huanuco Mountains,
Haenke. Chinanugula, Gilg 38. Cuzco : Santa Ana valley, Herrera
986. San Martin: San Roque, in the forest, Williams 7121.
"Cordoncillo."
Piper lineatum var. cordulatum Trelease, var. nov.
Folia basi rotundata et cordulata.
Type collected in Peru, Mathews 1691 (type in herb. Kew.).
Junin: Perene", Nordenskjold & Rospigliosi 147.
Piper lineatum var. leucolepidum Trelease, var. nov.
Cito glabrata; folia obscure versus basin granulosa 14 cm. longa
5 cm. lata; spicae 90-100 mm. longae, pedunculo 15 mm. longo;
bracteae pallidae.
Junin: Carrapa, 1,000 meters, Killip & Smith 22495 (type in
U. S. Nat. Herb.), 22451 . Ayacucho : Aina, Killip & Smith 22565.
Piper lineatum var. hirtipetiolatum Trelease, var. nov.
Frutex metralis valde nodosus, internodiis modice brevibus
gracilibus retrorse albido-hispidis pallido-granulosis, folia lanceolato-
elliptica attenuata basi plus minusve inaequali acuta 13 cm. longa
4.5-5 cm. lata e dimidio inferiore pinnatinervia, nervis profunde
impressis 5-6x2, obscurissime rugosa sublucida, supra conspicue
albido-granulosa, subtus laxe crispo-pubescentia nervis hirsutis,
petiolo 5-10 mm. longo subadpresso-hirsuto basi alato; spicae
oppositifoliae rectae vel serius arcuatae 80 mm. longae 3 mm. crassae,
pedunculo 10 mm. longo retrorso-hispido ; bracteae rotundo-sub-
peltatae ciliatae, floribus sessilibus perfectis.
FLORA OF PERU 185
Huanuco: Huacachi near Mima, 1,950 meters, Macbride 4182
(herb. Field Mus. No. 535,259, type).
Piper llatanum Trelease, sp. nov.
Frutex 2-4-metralis nodosus glaber, internodiis floriferis brevibus
crassis microscopice glanduloso-granulosis; folia late ovata subacuta
basi lata brevissime cordata parva 7-10 cm. longa 4-8 cm. lata e
parte inferiore 1-1.5 cm. longa submultiplinervia, nervis circa 3x2,
opaca, subtus dense brunneo-puncticulata, petiolo 10-12 mm. longo
circum caulem in lineam ciliatam protracto exalato; spicae erectae
pallido-flavae 60 mm. longae 4 mm. crassae, pedunculo 8 mm. longo;
bracteae rotundo-subpeltatae ciliolatae.
Huanuco: Llata, Rio Maranon Canyon, 2,100 meters, Macbride
& Feather stone 2302 (herb. Field Mus. No. 518,785, type).
Piper longeacuminatum Trelease, sp. nov.
Frutex nodosus, internodiis superioribus modice brevibus et
gracilibus molliter pilosis; folia ovata anguste longiacuminata basi
subaequilaterali obtusa 15 cm. longa 6 cm. lata e dimidio inferiore
pinnatinervia, nervis 5x2, in sicco modice firma brunnea, supra
minute granuloso-asperata, subtus ferrugineo-subtomentosa, petiolo
1 cm. longo ferrugineo-tomentoso exalato; inflorescentia non visa.
Junin: Hacienda Schunke, 1,300-1,700 meters, Carlos Schunke
A90 (type in U. S. Nat. Herb.).
Piper longegerminatum Trelease, sp. nov.
Frutex nodosus in sicco fuscus, ramis longis gracilibus, internodiis
floriferis gracilibus brevibus retrorso-subcanescentibus; folia lan-
ceolata acuta basi subaequilaterali subacuta 12 cm. longa 3-5
cm. lata e dimidio inferiore pinnatinervia, nervis 5x2, tenuia,
supra minute griseo-scabrida, subtus ad nervos strigosa, petiolo
5 mm. longo griseo-scabrido ; spicae 35 mm. longae 2 mm. crassae
mucronatae, pedunculo 3 mm. longo cinereo; bracteae parvae
rotundo-peltatae ciliolatae.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28839 (type in
U. S. Nat. Herb.). Balsapuerto, Killip & Smith 28439.
Piper longestamineum Trelease, sp. nov.
Arbor glabra 5-6-metralis, internodiis fioriferis gracilibus varie
elongatis; folia protracta elliptica obtusa basi inaequilateralia, sinu
laterali clause, latere longiore auriculato et petiolo longiore, 30 cm.
186 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
longa 12 cm. lata e dimidio inferiore subpinnatinervia, nervis 5-6x2,
in sicco tenuia sublucida, petiolo 4 cm. longo et ultra alato; spicae
200 mm. longae 4 mm. crassae, pedunculo gracili recurvo 3 cm. longo;
bracteae rotundo-subpeltatae ciliolatae; filamenta longa gracilia,
antheris ellipsoideis; stigmata 3 filiformia.
Junin: San Nicolas, 1,100 meters, Killip & Smith 26011 (type in
U. S. Nat. Herb.).
Piper longifilamentosum Trelease, sp. nov.
Arbor 4-5-metralis nervis folii subtus griseo-puberulis exceptis
glabra, internodiis floriferis longis gracilibus; folia elliptico-sub-
obovata obtusa basi inaequilateralia, sinu laterali, lobo longiore
protracto et petiolo longiore, 37 cm. longa 16 cm. lata e dimidio
inferiore pinnatinervia, nervis circa 6x2, in sicco tenuia et plus
minusve lucida, petiolo 6-7 mm. longo; spicae 110 mm. longae 5
mm. crassae, pedunculo gracili 3 cm. longo; bracteae triangular! -
subpeltatae; filamenta gracilia longiexserta; baccae globosae, stylo
gracili, stigmatibus 3 linearibus.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26452
(type in U. S. Nat. Herb.).
Piper longif olium R. & P. Fl. 1 : 37. 1798. Schilleria longifolia
Kunth, Linneaea 13: 725. 1839. Peltobryon longif olium Miq. Syst.
373. 1844.
A glabrous, more or less glandular-granular shrub ; leaves elliptic-
subobovate, abruptly short-acuminate, almost equilaterally long-
acute at the base, 32-41 cm. long, 13-18 cm. wide, pinnately nerved
throughout; petiole 3-4 cm. long, winged by the decurrent blade;
spikes gently curved, 110 mm. long, 10 mm. thick; peduncle 15 mm.
long; berries round, with an elongate style; stigmas small. Not
at all corresponding to the illustration of R. & P. loc. cit. pi. 61.
Huanuco: Forests of Pillao, Ruiz & Pavon, type; Ruiz 152, and
No. 641 in Herb. Willdenow. Loreto (?): Huallaga, Mathews 1712.
Junin: Schunke Hacienda, San Ramon, Killip & Smith 24552.
Piper loretoanum Trelease, sp. nov.
Frutex 3-4-metralis plus minusve nodosus, internodiis floriferis
brevibus modice gracilibus plus minusve sparse incurvo-subvillosis ;
folia subovato-elliptica anguste acuminata basi inaequilaterali ob-
tusa latere breviore subcordulato, 15-17 cm. longa 6.5-7 cm. lata
e dimidio inferiore pinnatinervia, nervis 5x2, in statu adulto rugosa,
supra conspicue granuloso-scabra, subtus ad nervos subvillosa et
FLORA OF PERU 187
inter nervos molliter pilosa, petiolo 3+3 mm. longo rigide subvilloso;
spicae juveniles 60 mm. longae 2 mm. crassae, pedunculo vix 5 mm.
longo subvilloso.
Loreto: Yurimaguas, 135 meters, Killip & Smith 27555 (type
in U. S. Nat. Herb.). Lower Rio Huallaga, Williams 4785.
Piper Macbrideanum Trelease, sp. nov.
Frutex 2.5 m. altus, r'amis divaricatis cavis glabris; folia elliptico-
oblonga ad subovata longiuscule acuminata basi subacuta vel
rotundata et acute contracta, mediocria 6-16 cm. longa 5.5-9 cm.
lata e 2 tertiis inferioribus pinnatinervia, nervis majoribus valde
sursum curvatis 4x2, supremis ex proximis distantibus, in sicco
subtenuia papyracea, petiolo 5-10 vel ad 30 mm. longo alato; spicae
oppositifoliae 60-90 mm. longae 2-3 mm. crassae, pedunculo gracili
5-10 mm. longo; bracteae rotundo-subpeltatae ciliolatae incon-
spicuae, floribus sessilibus perfectis; stigmata 3 minuta sessilia.
Junin: La Merced, on forest slope, 600 meters, Macbride 5516
(herb. Field Mus. No. 536,556, type).
Piper machupicchuense Trelease, sp. nov.
Frutex nervis folii subtus obscure adpresso-pubescentibus exceptis
glaber, in sicco griseo-viridis, internodiis floriferis gracilibus elongatis;
folia ovata vel ovato-oblonga plus minusve acuminata basi rotundata
et interdum uno latere breviora, 14 cm. longa 5-7 cm. lata vel usque
ad 9 cm. lata e dimidio inferiore multiplinervia, nervis 4-5x2, in
sicco tenuia, petiolo 15 mm. longo semi-alato; spicae 60-100 mm.
longae 3 mm. crassae, pedunculo 5-10 mm. longo; bracteae conspicuae
triangulari-subpeltatae ciliatae; ovarium suboblongo-truncatum,
stigmatibus 3 filiformibus vix connatis.
Cuzco: San Miguel, Valle de Machupicchu, 2,300 meters,
Fortunato L. Herrera 2006 (herb. Field Mus. No. 580,236, type).
Piper magnificum (Linden) Gentil, PI. Serr. Jard. Brux. 152.
1907, nomen. Artanthe magnified Linden, Cat. 1876; 111. Hort. 23:
pi. 239. 1876. A. decurrens Fourn. 111. Hort. 23: 58. 1876, non
A. decurrens Miq.
A soft-wooded shrub, glabrous except for the puberulent nerves
beneath and the obscure striation of petiole and stem ; leaves elliptic,
ranging to oblong and ovate, subacuminate, shallow-cordate, 15-22
cm. long, 7-11 cm. wide, pinnately nerved throughout, purple
beneath; petiole 1-5 cm. long, broadly winged; spikes 15 mm. long,
6 mm. thick, cuspidate; peduncle becoming equally long; bracts
188 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
rounded-subpeltate; berries round, with a slender style; stigmas
short, spreading.
Loreto: Balsapuerto, Killip & Smith 28391*, 28420, 28598.
Piper malifolium Trelease, sp. nov.
Frutex compactus 2-metralis valde nodosus, internodiis floriferis
brevibus gracilibus persistente adpresso-hispidis; folia elliptica
acuminata basi plus minusve inaequali obtusa 15-16 cm. longa 6-7.5
cm. lata e dimidio inferiore pinnatinervia, nervis circa 5x2, scabra,
subtus ad nervos patenti-hirsuta, in statu adulto rugosa, petiolo
10-15 mm. longo hirsute basi alato; spicae oppositifoliae 90 mm.
longae 2-3 mm. crassae, pedunculo 5 mm. longo adpresso-hispido ;
bracteae rotundo-subpeltatae dense ciliolatae, floribus sessilibus
perfectis.
Huanuco: Posuso, 600 meters, Macbride 4620 (herb. Field Mus.
No. 535,697, type).
Piper malifolium var. mercedanum Trelease, var. nov.
Arbor gracilis 5-metralis nodosa, internodiis floriferis subelongatis
gracilibus hirtellis; folia subelliptica acuminata basi subinaequali
obtusa 14-15 cm. longa 6-7 cm. lata e dimidio inferiore pinnatinervia,
nervis 5-6x2, scabra, subtus ad nervos villosa, venis in statu adulto
paullo impressis, petiolo 10-15 mm. longo hirsute basi alato; spicae
oppositifoliae 80-100 mm. longae 2-3 mm. crassae, pedunculo 10
mm. longo hirtello; bracteae rotundo-subpeltatae ciliolatae, floribus
sessilibus perfectis; baccae breviter cylindraceae truncatae; stigmata
3 minuta sessilia.
Junin: La Merced, 600 meters, Macbride 5288 (herb. Field Mus.
No. 536,313, type).
Piper mansericheanum Trelease, sp. nov.
Frutex, ramis transiente crispo-pubescentibus; folia anguste
lanceolata sensim acuminata basi acuta 13 cm. longa 1.5-2.5 cm.
lata e dimidio inferiore pinnatinervia, nervis circa 3x2 utrinque
minute glanduloso-granulosa, subtus pallidiora ad nervos adpresso-
pubescentia, petiolo 10 mm. longo; spicae 50 mm. longae 2 mm.
crassae mucronatae, pedunculo gracili 10-15 mm. longo; baccae
parvae teretes, stigmatibus sessilibus.
Loreto: Pongo de Manseriche, Tessmann 3883 (type in herb.
Berol.).
FLORA OF PERU 189
Piper maranyonense Trelease, sp. nov.
Frutex glaber; folia lanceolato-oblonga acuta basi subaequilaterali
acuta 22 cm. longa 8 cm. lata ubique pinnatinervia, nervis circa
12x2, plus minusve indistincte glanduloso-granulosa, petiolo vix
10 mm. longo; spicae 60 mm. longae 6 mm. crassae curvae obtusae,
pedunculo 15 mm. longo; bracteae triangulari-subpeltatae; baccae
rotundatae, stigmatibus 3 latis sessilibus.
Loreto: Mouth of Rio Santiago, upper Maranon, 160 meters,
Tessmann 4056 (type in herb. Berol.).
Piper marcescentispicum Trelease, sp. nov.
Frutex vel arbor 3-5-metralis, internodiis floriferis brevibus
crassiusculis primo sparse crispo-pilosis glabrescentibus; folia elliptica
subacuminata basi fere aequilaterali acuta 16-20 cm. longa 6-8 cm.
lata fere ubique pinnatinervia, nervis circa 6-8x2, in sicco tenuia
viridia, subtus ad nervos incurvo-hirsuta, petiolo 10 mm. longo
exalato; spicae pendulae 40 mm. longae 2 mm. crassae acutae,
pedunculo gracili 10 mm. longo glabrato; bracteae rotundo-sub-
peltatae, margine angusto lacerate.
Junin: Pichis Trail, 625-900 meters, Kittip & Smith 26182 (type
in U. S. Nat. Herb.).
Piper margaritatum Trelease, sp. nov.
Frutex, internodiis floriferis brevibus crassiusculis hirsutis;
folia late lanceolata vel lanceolato-ovata longiacuminata basi plus
minusve inaequilaterali cordulata 10-12 cm. longa 4-5 cm. lata e
dimidio inferiore pinnatinervia, nervis 4-5x2, supra grosse lepidoto-
asperata, subtus ad nervos hispida, petiolo 5 mm. longo; spicae
juveniles 45 mm. longae 2 mm. crassae, pedunculo sericeo
8 mm. longo.
San Martin: San Roque, in forest, 1,350-1,500 meters, Llewelyn
Williams 7742 (herb. Field Mus. No. 623,556, type). "Cordoncillo."
Piper marsupiiferum Trelease, sp. nov.
Frutex metralis, internodiis floriferis comparate brevibus et
gracilibus breviter villosis; folia anguste triangularia vel lanceolato-
oblonga subobtusa basi inaequilateralia, sinu lateral!, latere longiore
in auriculam petiolo fere duplo longiorem producto, 30 cm. longa
8-9 cm. lata e 2 tertiis inferioribus subpinnatinervia, nervis 4-5x2,
nervis supplementariis in auricula dispersis, in sicco tenuia, subtus
ad nervos breviter villosa, areolis inter venas majores crescentiis
bursiformibus completis, petiolo 3-4 cm. longo anguste alato patenti-
190 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
piloso; spicae juveniles 25 mm. longae 2 mm. crassae, pedunculo
gracili aequilongo piloso; bracteae rotundo-subpeltatae.
Loreto: Soledad, Rio Itaya, 110 meters, Killip & Smith 29606
(herb. Field Mus. No. 688,345, type).
Piper martinense Trelease, sp. nov.
Frutex nodosus, internodiis floriferis subgracilibus brevibus
breviter adpresso-villosis; folia lanceolato-oblonga sensim acuminata
basi subaequilaterali breviter cordata 25 cm. longa 6.5 cm. lata e
dimidio inferiore pinnatinervia, nervis 6-f7, supra leviter rugosa et
basibus pilorum caducorum scabrida, subtus sparse pubescentia ad
nervos subadpresso-villosa, petiolo 5 mm. longo adpresso-villoso ;
spicae uncatae 60 mm. longae 3 mm. crassae acutatae, pedunculo
10-15 mm. longo laxe piloso; bracteae rotundo-subpeltatae ciliatae.
San Martin: Lamas, near Tarapoto, 840 meters, Llewelyn Wil-
liams 6407 (herb. Field Mus. No. 623,618, type).
Piper masiseanum Trelease, sp. nov.
Frutex vel arbuscula 3-4-metralis, internodiis gracilibus cito
elongatis molliter adpresse retrorso-pilosis; folia subovata acuta
basi inaequilaterali bicordulata 10 cm. longa 4 cm. lata e dimidio
inferiore pinnatinervia, nervis 5+6, supra albide granuloso-scabra,
subtus ad nervos pallide adpresso-pubescentia, petiolo 3+3 mm.
longo basi tantum vaginante pallide adpresso-piloso ; spicae 90 mm.
longae 3 mm. crassae obtusae, pedunculo 10 mm. longo hispido;
bracteae rotundo-subpeltatae ciliolatae.
Loreto: Masisea, 275 meters, Killip & Smith 26846 (type in
U. S. Nat. Herb.).
Piper Mathewsii C. DC. in DC. Prodr. 16, pt. 1: 261. 1869.
A shrub with densely appressed-yellow-hirsute twigs; leaves
lanceolate, acute at both ends, 15 cm. long, 4.5 cm. wide, pinnately
nerved throughout, granular-scabrous and loosely hairy above,
densely appressed-hairy beneath; petiole 5-10 mm. long; spikes 40
mm. long, 3 mm. thick; peduncle 5-10 mm. long; bracts triangular-
subpeltate, yellow; berries subquadrangular; stigmas 3, sessile.
Amazonas: Chachapoyas, Mathews 3224, type.
Piper mediocre C. DC. Verh. Bot. Ver. Brandenb. 47: 104.
1905.
A delicate shrub resembling P. phthinotrichon but with narrower,
more lanceolate leaves, 7 cm. long and 2 cm. wide, more or less
evanescently crisp-puberulent.
FLORA OF PERU 191
Loreto: Iquitos, Ule 6245, type. Above Iquitos, Tessmann 3987.
San Antonio, Killip & Smith 29428. Mishuyacu, Klug 1491, 467,
501 ; Killip & Smith 29965. Soledad, Killip & Smith 29563.
Piper melendezense Trelease, sp. nov.
Frutex nodosus diffusus vel scandens, internodiis floriferis brevibus
gracilibus minute crispo-pubescentibus in statu adulto plus minusve
granulosis; folia elliptico-lanceolata acute acuminata basi angustata
cordulata, uno latere paullo breviore, 8-11 cm. longa 3-5 cm. lata e
dimidio inferiore pinnatinervia, nervis 5x2, supra minute granulosa,
subtus pallida granulosa ad nervos adpresso-pubescentia, petiolo
55+1-2 mm. longo; spicae 100 mm. longae 2-3 mm. crassae, pedun-
culo gracili vix 10 mm. longo glabrato; bracteae transverse subpel-
tatae ciliolatae; baccae depressae, stigmatibus sessilibus.
Loreto: Puerto Mele"ndez, below Pongo de Manseriche, 200
meters, Ynes Mexia 6382 (type in herb. Univ. Calif.).
Piper mellibracteum Trelease, sp. nov.
Frutex 1.5 m. altus valde nodosus, internodiis floriferis brevibus
crassis evanescente subdense hamato-hirsuto-villosis pallide granu-
losis; folia lanceolato-oblonga longiattenuata basi inaequaliter cor-
dulata, sinu clause, 15-16 cm. longa 3.5-4 cm. lata e dimidio inferiore
pinnatinervia, nervis circa 8x2, paullo rugosa, supra basibus pilis
longis adpressis scabra et ad costam plus minusve barbata, subtus ad
nervos subadpresso-villosa, petiolo brevissimo 5 mm. longo adpresso-
hirsuto exalato; spicae oppositifoliae valde curvatae 90 mm. longae
3-4 mm. crassae, pedunculo 8 mm. longo sparse hirtello; bracteae
rotundo-subpeltatae centre lutescente, margine pallido ciliato,
floribus sessilibus perfectis; stigmata 3 minuta sessilia; baccae
parvae cylindraceae.
Huanuco: Rio Huallaga Canyon below Rio Santo Domingo,
1,200 meters, Macbride 4234 (herb. Field Mus. No. 535,313, type).
Piper mercedense Trelease, sp. nov.
Frutex 2-3-metralis, internodiis floriferis longiusculis modice
gracilibus pallidis sparse breviter villosis glabrescentibus; folia
lanceolato-elliptica acute falcato-acuminata basi inaequaliterali
acuta, latere breviore plus minusve rotundato, 15-16 cm. longa 5-6
cm. lata e 2 tertiis inferioribus pinnatinervia, nervis circa 7x2,
supra scabro-hispida et lepidoto-granulosa, subtus ad nervos albido-
villosa, petiolo 3+5 mm. longo exalato; spicae floriferae 50 mm.
192 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
longae 2 mm. latae mucronatae, pedunculo gracili 15 mm. longo
glabro vel sparse hirtello; bracteae triangulari-subpeltatae ciliatae.
Junin: La Merced, 700 meters, Killip & Smith 23731 (type in
U. S. Nat. Herb.).
Piper mikaniaefolium Trelease, sp. nov.
Frutex nodosus glaber (scandens?), ramis fuscis, internodiis
superioribus brevibus gracilibus; folia elliptico-ovata vel lanceolato-
ovata acuminata basi cordata 11-14 cm. longa 4.5-6.5 cm. lata e
dimidio inferiore submultiplinervia, nervis 6x2, in sicco viridia,
subtus minute nigro-granulosa, petiolo 15-25 mm. longo infra
medium vaginante; spicae 20 mm. longae 4 mm. crassae, pedunculo
5 mm. longo recurvo; bracteae crescentico-subpeltatae; baccae
teretes conice prolongatae, stigmatibus minutis.
Loreto: Yurimaguas to Balsapuerto, 135-150 meters, Killip &
Smith 30691 (type in U. S. Nat. Herb.).
Piper mishuyacuense Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis comparate brevibus
gracilibus dense luteo-tomentulosis; folia elliptica plus minusve
falcata breviter acuminata basi subacuta, uno latere paullo breviore,
16-20 cm. longa 7-8.5 cm. lata ubique pinnatinervia, nervis circa
12x2, subtus praesertim ad nervos minute tomentulosa, petiolo
10+2 mm. longo tomentuloso; spicae juvenes 35 mm. longae 6 mm.
crassae, pedunculo crasso vix 10 mm. longo glabrato; baccae sub-
teretes, stigmatibus 3 sessilibus.
Loreto: Mishuyacu near Iquitos, 100 meters, Killip & Smith
29903 (type in U. S. Nat. Herb.). Balsapuerto, Killip & Smith
28611.
Piper mite R. & P. Fl. 1: 37. pi. 58. 1798. Artanthe mitis Miq.
Syst. 407. 1844.
A shrub with glabrous twigs; leaves ovate, acuminate, subtrun-
cate at the base, 20 cm. long, 11 cm. wide, submultiple-nerved from
the lower half, the nerves about 7x2, stellate-pubescent beneath;
petiole 2-3 cm. long, soft-pubescent; spikes 225 mm. long, 8 mm.
thick; peduncle 25 mm. long, thick, pubescent; bracts triangular-
subpeltate, ciliate; ovary round, with a short style; stigmas narrow.
Huanuco: Cochero, Ruiz & Pavon, type.
Piper mitifolium Trelease, sp. nov.
Frutex nodosus 2-3-metralis nervis folii subtus puberulis exceptis
glaber, internodiis floriferis brevibus crassis; folia elliptica obtuse
FLORA OF PERU 193
breviter acuminata basi cordulata 16-22 cm. longa 7-11 cm. lata
e 2 tertiis inferioribus submultiplinervia, nervis 6-8x2, lucida
viridia plus minusve rugosa, petiolo 2 cm. longo alato; spicae 25
mm. longae 4 mm. crassae, fructiferae 40 mm. longae 10 mm. crassae,
obscure apiculatae, pedunculo vix 10 mm. longo; bracteae triangulari-
subpeltatae vel crescentico-subpeltatae ciliatae; baccae globosae,
stigmatibus 3 sessilibus.
Junin: La Merced, 700 meters, in forest, Macbride 5325
(herb. Field Mus. No. 536,350, type). Hacienda Chalhuapuquio,
Stevens 204.
Piper Mocco-mocco Trelease, sp. nov.
Frutex glaber glanduloso-granulosus; folia elongato-elliptica
brevissime obtuso-acuminata basi fere aequilaterali acuta 15 cm.
longa 6 cm. lata ubique pinnatinervia, nervis circa 6x2, petiolo 2
cm. longo exalato; spicae leviter curvae 60 mm. longae 5 mm.
crassae cuspidatae, pedunculo brevi; bracteae triangulari-subpeltatae
ciliatae; baccae rotundatae, stigmatibus 3 parvis sessilibus.
Cuzco: Hacienda Potrero, Santa Ana, 1,250 meters, Fortunato
L. Herrera 954 (type in herb. Berol.). "Mocco-mocco."
Piper Mocco-mocco var. angustum Trelease, var. nov.
A forma typica foliis oblongis 18 cm. longis 5.5 cm. latis, pedun-
culo 10 mm. longo differt.
Cuzco: San Miguel, Urubamba Valley, 1,800 meters, Cook &
Gilbert 1123 (type in U. S. Nat. Herb.), 1124.
Piper moense C. DC. Notizbl. Bot. Gart. Berlin 6: 491. 1917.
A shrub with loosely white-villous nodes; leaves elliptic-sub-
acuminate, slightly inequilaterally cordulate, 15-16 cm. long, 7-9
cm. wide, pinnately nerved from below the upper third, the nerves
about 6+7, microscopically granular above, dark-punctulate beneath
and paler, with upcurved-pubescent nerves; petiole about 10 mm.
long; spikes 20 mm. long, 10 mm. thick, cuspidate; peduncle barely
5 mm. long; bracts round-subpeltate ; berries round, with an elongate
slender style; stigmas small.
Brazil: Monte Mo, Rio Acre, Ule 9286, type. Probably extend-
ing into Peru.
Scarcely more than a glabrescent form of P. xapuryense.
Piper Mohomoho C. DC. Bot. Jahrb. 40: 251. 1908.
A shrub, crisp-puberulent or the petiole and leaf base cobwebby ;
leaves ovate, acuminate, cordulate, 7.5-8 cm. long, 4-5 cm. wide,
194 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
pinnately nerved from below the middle, the nerves 3x2; petiole
10 mm. long; spikes 60 mm. long, 2-3 mm. thick; peduncle 10 mm.
long. Neg. 10995.
Cajamarca: San Pablo, 2,400-2,700 meters, in thicket, Weber-
bauer 3839, type. "Mohomoho."
Piper monostigmum C. DC. Notizbl. Bot. Gart. Berlin 7:
486. 1917.
A shrub, the young parts and the nerves beneath puberulent;
leaves elliptic, acuminate, acute-based, 11-12 cm. long, 4.5-5 cm.
wide, pinnately nerved from the lower half, the nerves 4-5x2;
petiole 5-15 mm. long; spikes 30 mm. long, 3 mm. thick; peduncle
5 mm. long; bracts round-subpeltate, ciliate; stigma 1, sessile.
Brazil: Seringal Auristella, Rio Acre, Vie 9273, type. Seringal
Sao Francisco, Ule 9274- Probably in adjacent Peru.
Piper monzonense C. DC. Bot. Jahrb. 40: 248. 1908.
A shrub, stiffly white-villous except for the upper surface of the
leaves, or the branches glabrescent; twigs pale- warty; leaves lanceo-
late or elliptic-oblong, acuminate, acute-based, 20 cm. long, 6 cm.
wide, pinnately nerved throughout; petiole 2 cm. long; spikes 100
mm. long, from filiform becoming 5 mm. thick; peduncle 10 mm. long;
stigmas sessile. Neg. 10998.
Huanuco: Monzon, in forest, 1,000 meters, Weberbauer 3429, type.
Piper morator Trelease, sp. nov.
Frutex vel arbor 3-5-metralis nodosa, internodiis floriferis brevi-
bus gracilibus; folia lanceolato-ovata acuminata basi subaequi-
laterali acuta 10-12 cm. longa 4-5 cm. lata e dimidio inferiore
pinnatinervia, nervis plus minusve sursum curvatis 4x2, petiolo 5
mm. longo infra medium vaginante; spicae juvenes 15 mm. longae
2 mm. crassae, pedunculo brevi; bracteae rotundo-subpeltatae
anguste alatae.
Junin: San Ramon, 900-1,300 meters, Kittip & Smith 24743
(type in U. S. Nat. Herb.).
Piper moyobambanum Trelease, sp. nov.
Frutex, ramis molliter subadpresso-pilosis; folia lanceolato-
oblonga vel elliptico-oblonga subacuminata basi inaequilaterali
cordulata 13 cm. longa 4 cm. lata e dimidio inferiore pinnatinervia,
nervis circa 6+7, supra granuloso-scabrida, subtus ad nervos adpres-
FLORA OF PERU 195
so-pubescentia, petiolo brevissimo; spicae uncatae 110 mm. longae
3 mm. crassae, pedunculo glabrato; bracteae subdeltoideae flavae.
San Martin: Moyobamba, Mathews (type). Amazonas: Cha-
chapoyas, Mathews 3225. Without locality, Mathews 1693.
Piper multimammosum Trelease, sp. nov.
Frutex 1-2-metralis, internodiis floriferis breviusculis gracilibus
crispo-puberulis; folia subelliptica anguste acuminata basi subae-
quilaterali acuta 9-12 cm. longa 4.5-5.5 cm. lata ubique pinnati-
nervia, nervis circa 10x2, subtus ad nervos crispo-puberula, petiolo
5-15 mm. longo puberulo interdum alato; spicae pendulae 30-35
mm. longae 5 mm. crassae, pedunculo 5-10 mm. longo puberulo;
bracteae inconspicuae concavae; baccae congestae mammaeformes,
stigmatibus minutis mucronem terminantibus.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26645
(type in U. S. Nat. Herb.). Santa Rosa, 135 meters, Killip &
Smith 28848. Puerto Arturo, Killip & Smith 27786. Yurimaguas,
Killip & Smith 27621.
Piper munchanum Trelease, var. latutn Trelease, var. nov.
Frutex minute glanduloso-granulosus glaber vel ad nervos folii
subtus obscure puberulus; folia late elliptica 22-30 cm. longa 15-19
cm. lata breviter acuminata basi rotundata, uno latere 2-3 mm.
breviore; spicae juvenes 25 mm. longae 5 mm. crassae.
San Martin: San Roque, 1,350-1,500 meters, in forest, Llewelyn
Williams 6954 (herb. Field Mus. No. 623,002, type), 6961.
Piper mundum Trelease, sp. nov.
Frutex subvalidus, internodiis floriferis modice crassis brevibus
pilis fragilibus incurvo-subvillosis; folia elliptica breviter acuminata
basi paullo inaequilaterali cordulata 13-16 cm. longa 6.5-8.5 cm.
lata e dimidio inferiore pinnatinervia, nervis ut venis impressis circa
7x2, supra minute aspero-granulosa, subtus ad nervos ut videtur
sordide subvillosa, petiolo 7+3 mm. longo subvilloso exalato; spicae
150 mm. longae 4 mm. crassae acutatae, pedunculo 5 mm. longo
griseo-piloso; bracteae rotundo-subpeltatae ciliatae; baccae teretes,
stigmatibus sessilibus.
Loreto: Santa Rosa, Yurimaguas, in forest, 155-210 meters,
Llewelyn Williams 4758 (herb. Field Mus. No. 623,762, type).
Piper munyanum Trelease, sp. nov.
Frutex 1.5 m. altus nodosus, internodiis floriferis modice elongatis
et gracilibus glabris sparsissime pallido-granulosis; folia rotundo-
196 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
ovata vel elliptico-ovata acuminata basi rotundata 5-16 cm. longa
4.5-10 cm. lata e tertio inferiore submultiplinervia, nervis circa
4x2, subtus pallidiora ad nervos sparse villosa ad nervulos hirtella,
petiolo 2-3 cm. longo sparse dorso villoso alato; spicae oppositifoliae
60-80 mm. longae 4 mm. crassae, pedunculo subgracili 15 mm.
longo sparse villoso; bracteae rotundo-subpeltatae vel triangulari-
subpeltatae ciliatae, floribus sessilibus perfectis; ovarium glabrum
in stylum brevem attenuatum, stigmatibus 3 paten tibus.
Huanuco: Muna, 2,400 meters, in forest, Macbride 4272 (herb.
Field Mus. No. 535,354, type).
Piper musteum Trelease, sp. nov.
Frutex nodosus pilosus 1.5-2 m. altus, internodiis floriferis
brevibus crassiusculis primo dense rigido-villosis glabrescentibus ;
folia elliptica acuminata basi fere aequilaterali acuta 15 cm. longa
6 cm. lata ubique pinnatinervia, nervis circa 10x2, utrinque sparse
crispo-pilosa, subtus ad nervos rigido-villosa, petiolo 1-2 cm. longo
pilosissimo; spicae 60 mm. longae 7 mm. crassae obtusae, pedunculo
1 cm. longo plus minusve pubescente; bracteae triangulari-sub-
peltatae ciliatae; baccae breviter cylindraceae granulosae interdum
apice depressae mucronatae, stigmatibus 3.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28982 (type in
U. S. Nat. Herb.).
Piper nanayanum Trelease, sp. nov.
Arbuscula glabra, internodiis floriferis comparate brevibus et
crassis; folia subovato-elliptica subacuminata basi cordulata, uno
latere distincte breviore, 15-16 cm. longa 7-7.5 cm. lata ubique
pinnatinervia, nervis circa 12x2, in sicco tenuia, supra obscuriora
et albo- vel purpureo-lepidoto-maculata, petiolo 5+5 mm. longo
alato; spicae 100 mm. longae 2 mm. vel fructiferae 4 mm. crassae,
pedunculo 10 mm. longo; bracteae rotundo-subpeltatae vel trian-
gulari-subpeltatae ciliatae; baccae subglobosae, stigmatibus 3
minutis sessilibus.
Loreto : South bank of Rio Amazonas, Nanay, Llewelyn Williams
516 (herb. Field Mus. No. 603,936, type).
Piper nasutum Trelease, sp. nov.
Planta suffrutescens glabra, internodiis floriferis brevibus graci-
libus nigrescentibus plus minusve papillosis; folia elliptica vel sub-
lanceolata anguste acuminata basi paullo inaequilateralia, latere
breviore subobtuso, 7-10 cm. longa 3-4.5 cm. lata e 2 tertiis inferiori-
FLORA OF PERU 197
bus pinnatinervia, nervis 4+5, in sicco opaca brunnescentia, subtus
microscopice nigro-punctulata, petiolo gracili 10 mm. longo exalato;
spicae fructiferae 15-20 mm. longae 5 mm. crassae, juvenes longi-
acuminatae, pedunculo paten te filiform! 10 mm. longo; bracteae cres-
centicae;baccae globosae, stigmatibus brevibus stylum terminantibus.
Loreto: Yurimaguas, edge of forest, 155-210 meters, Llewelyn
Williams 3861 (herb. Field Mus. No. 623,840, type).
Piper neglectum Trelease, sp. nov.
Frutex glaber; folia elliptico-lanceolata vel lanceolato-ovata
subabrupte acuminata basi acuta vel in foliis latioribus obtusa 9-14
cm. longa 4-6 cm. lata ubique pinnatinervia vulgo glanduloso-
granulosa, petiolo 1 cm. longo et exalato vel in foliis latioribus 2-3
cm. longo et alato; spicae leviter curvae 30 mm. longae 5-8 mm.
crassae obtusae, pedunculo 5-10 mm. longo; baccae globosae, stylo
gracili, stigmatibus 3 patentibus.
San Martin: Tocache, Rio Huallaga, Poeppig D1949 (type in
herb. Vienna); also in 1832 and 1833. Huanuco: Cochero, Poeppig
in 1829.
Piper nuncupatum Trelease, sp. nov.
Frutex nodosus glaber 3-4-metralis, internodiis superioribus
brevibus gracilibus; folia lanceolato-elliptica falcato-acuminata basi
fere aequilaterali acuta 12-14 cm. longa 5-7 cm. lata e tertio inferiore
pinnatinervia, nervis 3-4x2, in sicco tenuia venulosa, petiolo 5 mm.
longo; inflorescentia non visa.
Junin: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25577 (type in U. S. Nat. Herb.).
Piper obesum Vahl, Enum. 1: 321. 1804. P. scabrum R. & P.
Fl. 1: 34. pi. 55. 1798, non P. scabrum Lam. Artanthe obesa Miq.
Syst. 470. 1844.
A scabrous shrub; leaves subelliptic, acuminate, subacute at the
base, 8 cm. long, 4 cm. wide, rugose; petiole scarcely 10 mm. long;
spikes 25 mm. long, 5 mm. thick; peduncle 5 mm. long.
Huanuco: Huassahuassi, Ruiz & Pavon, type.
Piper obliquum R. & P. Fl. 1: 37. pi. 63. 1798. Steffensia
obliqua Kunth, Linnaea 13: 663. 1839. Artanthe magnifica Miq.
Syst. 391. 1844; 111. Piper, pi. 71. P. Ruizianum, D. Dietr. Syn. PL
1:118.1839.
198 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
A shrub, glabrous or glabrate except for the typically puberulent
nerves beneath; leaves broadly ovate, subacute, obliquely cordate
with the larger side round-auricled and equaling the petiole, 29-39
cm. long, 14-26 cm. wide, subpinnately nerved from below the upper
third, the nerves about 7+9; petiole 4-6 cm. long, equally long
on both sides; spikes 300-500 mm. long, 5 mm. thick; peduncle
3-4 cm. long.
Huanuco: Cochero, Ruiz & Pavon, type; Dombey; Poeppig.
Loreto(?): Mainas, Poeppig 13. Without locality: Mathews 1699.
Piper oblongatifolium Trelease, sp. nov.
Frutex nodosus glaber 3-metralis, internodiis floriferis brevibus
gracilibus prope nodos minute granulosis; folia oblonga anguste
acuminata basi paullo inaequilateralia, latere breviore obtuso, 12-15
cm. longa 4-5.5 cm. lata e dimidio inferiore pinnatinervia, nervis
5x2, in sicco tenuia, subtus paullo pallidiora, petiolo 5+2 mm. longo
ut basi costae papillose non vaginante; spicae juvenes 60 mm. longae
2 mm. crassae, pedunculo 10-15 longo; bracteae triangulari-sub-
peltatae.
Loreto: Valley of Rio Maranon, 150 meters, Killip & Smith
29153 (type in U. S. Nat. Herb.).
Piper oblongatilimbum Trelease, sp. nov.
Frutex glaber 1-2-metralis, internodiis floriferis gracilibus modice
elongatis; folia lanceolato-oblonga acuminata basi inaequilaterali
obtusa vel basi ipsa subacuta 12-15 cm. longa 4 cm. lata e 2 tertiis
inferioribus pinnatinervia, nervis 4-5x2, in sicco tenuia, petiolo vix
5 mm. longo vaginante; spicae 20 mm. longae 5 mm. crassae caudatae
nigrescentes, pedunculo vix 5 mm. longo; baccae globosae, stig-
matibus sessilibus.
Loreto: Yurimaguas to Balsapuerto, 135-150 meters, Killip &
Smith 28237 (type in U. S. Nat. Herb.).
Piper oblongatum Opiz in Presl, Rel. Haenk. 153. 1830.
Carpunya oblongata Presl, Epim. Bot. 229. 1849. Artanthe oblongata
Miq. Syst. 535. 1844.
A glabrous shrub; leaves lanceolate or lance-ovate, gradually
subacuminate, inequilaterally rounded at the base, 15-16 cm. long,
5-7 cm. wide, pinnately nerved from below the middle, the nerves
4x2, coriaceous; petiole 10 mm. long; spikes 125 mm. long, 2 mm.
thick; bracts subpeltate, brown-hairy; stigmas sessile, filiform.
Huanuco: Huanuco mountains, Haenke, type.
FLORA OF PERU 199
Piper obovatifolium Trelease, sp. nov.
Arbor 3-5-metralis nervis folii subtus obscure puberulis exceptis
glabra, internodiis floriferis subelongatis crassis; folia elliptico-
obovata obtusa basi inaequilaterali cordata, uno latere subauriculate
prolongato, 30-33 cm. longa 18-21 cm. lata e 2 tertiis inferioribus
subpinnatinervia, nervis 6+7, in sicco tenuia lucida, petiolo 5-7+
1.5-2 cm. longo firme crispo-alato; spicae juvenes 2 mm. crassae,
pedunculo recurvo 2 cm. longo; bracteae rotundo-subpeltatae.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28980 (type in
U. S. Nat. Herb.). Yurimaguas to Balsapuerto, 135-150 meters,
Killip & Smith 28096.
Piper obovatum R. & P. Fl. 1: 36. pi. 53. 1798. Schilleria
obovata Kunth, Linnaea 13: 686. 1839. Artanthe obovata Miq. Syst.
508. 1844.
A glabrous or locally obscurely puberulent shrub; leaves oblan-
ceolate, acuminate, acute-based, 11-12 cm. long, 3-4 cm. wide, pin-
nately nerved throughout; petiole 10-15 mm. long; spikes in fruit
30 mm. long, 10 mm. thick, gently curved; peduncle scarcely 10
mm. long, recurved; bracts triangular-subpeltate; berries subglobose,
wrinkled ; stigmas sessile. Neg. 8419.
Huanuco: Cochero and Chinchao, Ruiz & Pavon, type.
The Ruiz specimen in the Willdenow Herbarium, with young
spikes, is accompanied by detached long spikes (as figured) ; but the
Ruiz specimen in the general herbarium at Berlin, though in full
fruit, possesses the short, thick spikes here described.
Peperomia occultum Trelease, sp. nov.
Frutex folia P. oculatispico similis nodosus, internodiis brevibus
modice crassis flavescenti-hispidis; folia lanceolata acuminata basi
paullo inaequilaterali uno latere basis angustatae obtusa 12 cm.
longa 5 cm. lata e dimidio inferiore vel paullo altius pinnatinervia,
nervis circa 6+5, plus minusve rugosa supra breviter scabrida et
serius minute granuloso-asperata nervis hispidis, subtus ad nervos
hispida, petiolo 8-10+2 mm. longo flavescenti-hispido; spicae 90
mm. longae 3 mm. crassae, pedunculo 6 mm. longo scabro; bracteae
rotundo-subpeltatae.
Loreto: Near Iquitos, 120 meters, Llewelyn Williams 1406 (herb.
Field Mus. No. 617,357, type). "Cordoncillo."
200 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper oculatispicum Trelease, sp. nov.
Frutex plus minusve nodosus, internodiis floriferis modice brevi-
bus et gracilibus flavescenti-hispidis; folia lanceolata acuminata basi
subinaequilaterali obtusa 11-14 cm. longa 4-5 cm. lata e dimidio
inferiore pinnatinervia, nervis 6x2, granuloso-scabra, subtus ad
nervos ferrugineo-hispida, petiolo 5+2 mm. longo f errugineo-hispido ;
spicae 75 mm. longae 3 mm. crassae, pedunculo 5 mm. longo scabro;
bracteae rotundo-subpeltatae, centre brunneo, margine pallido.
Loreto: Iquitos, 120 meters, Llewelyn Williams 7954 (herb. Field
Mus. No. 629,718, type). "Cordoncillo."
Piper offensum Trelease, sp. nov.
Frutex 2-3-metralis, internodiis superioribus subgracilibus brevi-
bus subvilloso-hirsutis ; folia elliptica acuminata basi acuta 17 cm.
longa 7-8.5 cm. lata e dimidio inferiore vel paullo altius pinnatinervia,
nervis 5-6x2, granuloso-scabra, subtus ad nervos longihirsuta,
petiolo vix 10 mm. longo piloso; spicae juvenes parvae.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28426
(type in U. S. Nat. Herb.).
Piper ollantaitambanum Trelease, sp. nov.
Frutex nodosus, internodiis floriferis modice gracilibus breviter
adpresse griseo-pubescentibus; folia late lanceolata subacuminata
basi inaequilaterali acuta vel vulgo uno latere rotundata 10-11 cm.
longa 3.5-4 cm. lata e dimidio inferiore pinnatinervia, nervis circa
6+5, subrugosa, supra viridia minute scabra lepidota, subtus
praesertim ad nervos puberula et breviter adpresso-pubescentia,
petiolo 7-10+2 mm. longo breviter adpresso-canescente; spicae
oppositifoliae fere rectae vel vulgo conspicue curvae 70-90 mm.
longae 4 mm. crassae, pedunculo 10 mm. longo breviter canescente;
bracteae rotundo-subpeltatae brunnescenti-tomentulosae.
Cuzco: Ollantaitambo, 3,000 meters, Cook & Gilbert 558 (type in
U. S. Nat. Herb.).
Piper Opizianum Fuernr. Flora, 12: 228. 1829. P. arcuatum
Opiz in Presl, Rel. Haenk. 151. 1831, non Blume. P. falcatum
A. Dietr. Syn. 1: 137. 1831.
A shrub; young parts sparsely crisp-pubescent; twigs granular;
leaves lanceolate, acutely attenuate at both ends, 15-16 cm. long,
5-7 cm. wide, pinnately nerved from the lower half, the nerves
FLORA OF PERU 201
5X2, ciliate; petiole 15 mm. long, not winged; spikes U-shaped, 100
mm. long, 2 mm. thick; peduncle 10 mm. long.
Type from mountains of Peru, Haenke in herb. Prague.
Piper Opizii (Kunth) Steud. Nom. Bot. ed. 2. 342. 1840. Steffen-
sia Opizii Kunth, Linnaea 13: 641. 1839. Artanthe Opizii Miq. Syst.
440. 1844. P. asperifolium Opiz in Presl, Rel. Haenk. 153. 1831,
non R. & P.
A nodose shrub; internodes short, slender, retrorsely scabrid,
granular in age; leaves elliptic-ovate, short-acuminate, shallowly
cordate or cordate-truncate, 8-11 cm. long, 5-6 cm. wide, pinnately
nerved from the lower half, the nerves 4-5x2, very scabrous above,
with hispid nerves beneath; petiole 10-15 mm. long, scabrous, not
winged; spikes 40 mm. long, 3-5 mm. thick, blunt; peduncle 15 mm.
long, scabrid; bracts transversely subpeltate; stigmas 3, sessile.
Huanuco: Huanuco mountains, Haenke.
Piper ornithorhynchum Trelease, sp. nov.
Frutex vel arbuscula 3-5-metralis, internodiis floriferis modice
gracilibus et elongatis ferrugineo-araneoso-villosis; folia rotundo-
ovata acuminata basi truncata vel breviter concava abrupte in
petiolum contracta 20-23 cm. longa 14-16 cm. lata e tertio inferiore
submultiplinervia, nervis 6-7x2, subtus sparse stellato-pubescentia
ad nervos ferrugineo-araneosa, petiolo 2-4 cm. longo vaginante
subvilloso; spicae 100 mm. longae 5-7 mm. crassae curvae, apice
sterili, ferrugineo-tomentosae, pedunculo 7-12 mm. longo 3 mm.
crasso ferrugineo-pubescente; bracteae rotundo-subpeltatae pilosae;
stigmata majuscula stylum distinctum terminantia.
Ayacucho: Carrapa, 1,500 meters, Killip & Smith 22334 (type
in U. S. Nat. Herb.), 23221.
Piper ostii Trelease, sp. nov.
Frutex humilis dense ramosus glaber subnodosus, internodiis
superioribus brevibus gracillimis; folia lanceolata sensim longissime
acuminata basi aequilaterali acuta, uno latere interdum distincte
breviore, 11-12 cm. longa 2-3.5 cm. lata e tertio inferiore pinnati-
nervia, nervis gracilibus 4x2, petiolo 10 mm. vel 10+2-3 mm. longo
gracili; spicae 30-40 mm. longae 2 mm. crassae, pedunculo filiformi
10 mm. longo.
Loreto: Mouth of Rio Santiago, 200 meters, Ynes Mexia 6152
(type in herb. Univ. Calif.).
202 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper ovantherum C. DC. Notizbl. Bot. Gart. Berlin 7:
491. 1917.
A shrub, glabrous except that the upper part of the peduncle is
retrorsely hirtellous; leaves ovate, acuminate, subtruncate at the
base but abruptly narrowed to the petiole, with one side slightly
shorter, 12-15 cm. wide, 21-23 cm. long, pinnately nerved from the
lower half, the nerves 5x2; petiole 20+5 mm. long, winged; spikes
80-110 mm. long, 3-5 mm. thick; peduncle 10 mm. long; bracts
triangular-subpeltate, ciliate; berries round, with a nearly equilong
style; stigmas 3, slender, recurving.
Brazil: Seringal Sao Francisco, Rio Acre, Ule 9279, type; 9281.
Loreto: Rio Maranon, Tessmann 3862.
Piper paganicum Trelease, sp. nov.
Frutex 2-3-metralis nodosus, internodiis floriferis brevibus
gracilibus transiente hirtellis plus minusve pallido-granulosis; folia
subovato-elliptica acuminata basi rotundata, uno latere paullo
breviore, 10-12 cm. longa 5.5-6 cm. lata e dimidio inferiore pinnati-
nervia, nervis 5X2, in sicco viridia lucida subrugosa, subtus ad nervos
adpresso-pubescentia, petiolo 10 vel 8+2 mm. longo inferne vaginante
subhirtello; spicae 45-65 mm. longae 3 mm. crassae, pedunculo 10
mm. longo glabrato; bracteae rotundo-subpeltatae ciliolatae.
Loreto: Iquitos, 100 meters, Killip & Smith 27432 (type in U. S.
Nat. Herb.).
Piper pallididorsum Trelease, sp. nov.
Frutex glaber 2-metralis, internodiis floriferis mediocribus; folia
subelliptica obtuso-subacuminata basi inaequilaterali cordata, latere
majore subauriculate producto et petiolum aequante, 20 cm. longa
15 cm. lata, e dimidio inferiore vel paullo altius subpinnatinervia,
nervis circa 6+7, subfirma, supra lucida, subtus opaca pallida,
petiolo 5+1 cm. longo alato; spicae 360 mm. longae (normales?) 7
mm. crassae versus basin steriles, pedunculo 15-20 mm. longo
recurvo; bracteae lunulato-subpeltatae ciliatae; baccae cubicae
glabrae, stigmatibus 3 sessilibus.
Junin: Schunke Hacienda, La Merced, 1,300 meters, Macbride
5801 (herb. Field Mus. No. 536,830, type); inflorescences pendent;
leaves whitish beneath when fresh.
Piper paludosum C. DC. in DC. Prodr. 16, pt. 1: 283. 1869.
A black-glandular-granular shrub, glabrous except for the
slightly velvety twigs; leaves elliptic-subovate, acuminate, slightly
FLORA OF PERU 203
inequilateral at the base and cordulate, 11 cm. long, 4.5 cm. wide,
pinnately nerved from below the upper third, the nerves 3-4x2,
the petiole 5 mm. long; young spikes 15 mm. long, 2 mm. thick,
cuspidate; peduncle filiform, 15 mm. long; bracts transversely
subpeltate, ciliate.
Loreto: Mainas, Yurimaguas, Poeppig, type.
Piper papillipetiolatum Trelease, sp. nov.
Frutex gracilis metralis nervis folii subtus obscure adpresso-
pubescentibus exceptis glaber, caulibus ex parte praesertim prope
nodos pallido-granulosis; folia lanceolata acute falcato-acuminata
basi subaequilaterali obtusa vel uno latere paullo longiore, 11-12
cm. longa 4-5 cm. lata infra medium pinnatinervia, nervis 4x2, in
sicco tenuia, petiolo 5-10 mm. longo papillose; inflorescentia
non visa.
Loreto: Iquitos, 100 meters, Kittip & Smith 27407 (type in U. S.
Nat. Herb.).
Piper paraguassuanum C. DC. Notizbl. Bot. Gart. Berlin 7:
489. 1917.
A transiently crisp-pubescent shrub; leaves glabrous above,
lance-ovate, gradually acute, acute-based, 15 cm. long, 5.5 cm. wide,
pinnately nerved from the lower half, the nerves 4-5x2; petiole 10
mm. long; spikes 65-95 mm. long, 2 mm. thick; peduncle 15 mm.
long; bracts smoky- tomentulose; berries compressed from the side;
stigmas 3, sessile.
Brazil: Paraguassu, Rio Acre, Vie 9284, type. Probably also
in adjacent Peru.
Piper paraisense Trelease, sp. nov.
Frutex nodosus, internodiis brevibus crassis transiente crispo-
pilosis; folia lanceolato-elliptica 19 cm. longa 6-6.5 cm. lata acute
acuminata basi inaequilaterali acuta e dimidio inferiore pinnatinervia,
nervis 6+7, in sicco firma obscure rugosa, supra basibus pilorum
longorum caducorum grosse granuloso-asperata, subtus ad nervos
rigido-subvillosa, petiolo 6+4 mm. longo laxe longipiloso; inflorescen-
tia non visa.
Loreto: Paraiso, Alto Rio Itaya, 145 meters, Llewelyn Williams
3207 (herb. Field Mus. No. 618,571, type).
Piper paralaevigatum Trelease, sp. nov. P. laevigatum Auct.;
P. tenue Auct.
204 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Frutex glaber vel in statu juveni obscure hinc inde papilloso-
puberulus; folia lanceolata anguste falcato-acuminata e medio fere
cuneatim attenuata ad petiolum, 21-24 cm. longa 7-9.5 cm. lata 3-
vel obscure 5-nervia, in sicco viridia nervis albidis, petiolo 10-15 mm.
longo vix alato; spicae oppositifoliae graciles 90 mm. longae, pedun-
culo 15 mm. longo; bracteae subcucullatae; baccae ellipsoideo-
oblongae apice contractae, stigmatibus 3 brevibus crassis.
Junin(?): Mainas Alto, Poeppig 1 (type in herb. Paris).
Piper paranapuranum Trelease, sp. nov.
Frutex 2-metralis diffusus, internodiis superioribus brevibus
subgracilibus villosis; folia oblongo-lanceolata vel elliptico-lanceo-
lata acuminata basi obtusa, uno latere paullo breviore, 11-15 cm.
longa 3-5 cm. lata e dimidio inferiore pinnatinervia, nervis 4-5x2,
supra granuloso-scabra, subtus pallidiora praesertim ad nervos
sparse hirsuto-subvillosa, petiolo 5-10 mm. longo subvilloso; spicae
90 mm. longae 3 mm. crassae, pedunculo 5 mm. longo hirsuto;
bracteae rotundo-subpeltatae ciliolatae.
Loreto: Rio Parana-pura near Yurimaguas, 180 meters, Ynes
Mexia 6075 (herb. Univ. Calif., type).
Piper parcepilosum Trelease, sp. nov.
Frutex 3-4-metralis, internodiis floriferis brevibus subgracilibus
glabris vel ad nodos parce pilosis; folia late ovata acuminata basi
rotundata acute ad petiolum contracta 20-23 cm. longa 10-15 cm.
lata e tertio inferiore pinnatinervia, in sicco tenuia sed firma, subtus
ad nervos sparse breviter villosa, petiolo 25 mm. longo infra medium
alato sparse molliter piloso; spicae 90 mm. longae 4 mm. crassae,
pedunculo gracili 10 mm. longo glabrescente; bracteae rotundo-
subpeltatae, centro fusco, margine brunneo crispo; stigmata sessilia.
Junin: Carpapata, 2,700-3,200 meters, Killip & Smith 24385
(type in U. S. Nat. Herb.).
Piper parvicordulatum Trelease, sp. nov.
Frutex plus minusve nodosus, internodiis gracilibus elongatis;
folia subelliptico-ovata acuminata basi rotundata vel abrupte
angustata et supra petiolum cordulata 14-18 cm. longa 8-12 cm.
lata e dimidio inferiore vel paullo altius submultiplinervia, nervis
longioribus 6-7x2, in sicco tenuia opaca, subtus pallidiora, petiolo
gracili 2 cm. longo non vaginante; spicae juvenes 50 mm. longae
3 mm. crassae, pedunculo 10 mm. longo; bracteae crescentico-
subpeltatae; stigmata 3 filiformia sessilia.
FLORA OF PERU 205
Junin: Chanchamayo Valley, 1,800 meters, Carlos Schunke 319
(herb. Field Mus. No. 571,375, type).
Piper patens Opiz in Presl, Rel. Haenk. 153. 1831.
A shrub with gray-hispid or hirsute twigs; leaves elliptic, acumin-
ate, obtuse at the base, 14 cm. long, 7 cm. wide, pinnately nerved
from somewhat more than the lower half, the nerves 4-5x2, hairy,
and with the upper side scabrous from the bases of fallen hairs;
petiole 5 mm. long; spikes as yet very young and small.
Huanuco: Huanuco mountains, Haenke in 1790, type.
Piper patentifolium Trelease, sp. nov.
Herba(?), internodiis floriferis gracilibus breviusculis hispide
griseo-hirsutis; folia elliptica vel elliptico-ovata et basi rotundata,
vel elliptico-subobovata et basi acuta, acute acuminata 15-16 cm.
longa 7-8.5 cm. lata e dimidio inferiore pinnatinervia, nervis circa
6x2, tenuia sed rugosa, supra dense granuloso-scabra, subtus ad
nervos adscendenti-hirsuta, petiolo 10 vel 5+5 mm. longo hirsute;
spicae oppositifoliae 100 mm. longae 3 mm. crassae acutatae, pedun-
culo vix 5 mm. longo plus minusve hispido; bracteae parvae rotundo-
subpeltatae ciliolatae.
Junin: Hacienda Chalhuapuquio, Stevens 136 (type), 148.
Piper paucartamboanum Trelease, sp. nov.
Frutex gracilis diffusus griseus metralis nodosus, internodiis
floriferis brevibus gracilibus strigoso-pilosis; folia lanceolata acute
acuminata basi inaequilateralia, uno vel ambobus lateribus rotun-
datis, 6.5-10 cm. longa 2.5-3.5 cm. lata, supra scabrida, subtus ad
nervos venasque prominentes hirsuta, petiolo 5-10 mm. longo inferne
vaginante; spicae 50 mm. longae 3 mm. crassae, pedunculo 5 mm.
longo hispido; bracteae majusculae triangulari-subpeltatae; baccae
oblongae truncatae, stigmatibus parvis sessilibus.
Junin : Near Perene Bridge, Rio Paucartambo Valley, 700 meters,
Killip & Smith 25388 (type in U. S. Nat. Herb.).
Piper paulianifolium Trelease, nom. nov. Artanthe Haenke-
ana Miq. Linnaea 20: 145. 1847, non P. Haenkeanum Opiz. P.
paulianum Auct.
A shrub, glabrous; leaves elliptic-subovate, acuminate, rounded
at the base, with one side shorter, 22 cm. long, 12 cm. wide, sub-
multiple-nerved from the lower half, the nerves about 6x2, minutely
cellular-papillate beneath; spikes elongate, 4 mm. thick, the peduncle
206 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
5 mm. long; bracts subtriangular-subpeltate, the broad yellow margin
ciliate; berries rounded; stigmas 3, sessile.
Type collected in Peru(?) by Haenke.
Piper Pavoni (Miq.) C. DC. in DC. Prodr. 16, pt. 1: 294. 1869.
Artanthe Pavoni Miq. Lond. Journ. Bot. 4: 45. 1866.
A shrub, more or less persistently yellow-subvillous or crisp-
hairy except for the nearly glabrous upper side of the leaves; leaves
broadly ovate, acuminate, subtruncate at the base or shallowly
cordate, 13-30 cm. long, 9-17 cm. wide, submultiple-nerved from the
lower third, the nerves about 5x2; petiole 2-3 cm. long, narrowly
winged; spikes 90-100 mm. long, 3-5 mm. thick; peduncle 10 mm.
long; bracts triangular-subpeltate; ovary round, with a deciduous
style; stigmas spreading. Neg. 11014.
Type from Quebrada of Panuahuanuca, Mathews 793. Peru-
vian Andes, Dombey 936. Junin: Huacapistana, Stevens 179(1}.
Piper pebasense Trelease, sp. nov.
Arbuscula glabra nodosa, internodiis floriferis brevibus modice
gracilibus; folia late elliptico-oblanceolata sensim acute acuminata
basi rotundata 26 cm. longa 10 cm. lata ubique pinnatinervia, in
sicco tenuia brunnescentia, petiolo 5+15 mm. longo alato; spicae
150 mm. longae et ultra 3 mm. crassae, pedunculo crasso 5 mm.
longo; bracteae triangulari-subpeltatae ferrugineo-ciliatae; ovarium
subglobosum, stigmatibus 3 parvis sessilibus.
Loreto : Pebas on the Amazon River, in forest, Llewelyn Williams
1746 (herb. Field Mus. No. 608,248, type).
Piper Peltobryon C. DC. in DC. Prodr. 16, pt. 1: 259. 1869.
Peltobryon attenuatum Klotzsch in Miq. Syst. 371. 1844, non Piper
attenuatum Auct. pi.
A glabrous, glandular-granular shrub; leaves elliptic, equally
acute at both ends, 5-6 cm. wide, 13 cm. long, pinnately nerved from
below the upper third, the petiole 10 mm. long; spikes about 15 mm.
long, 10 mm. thick; peduncle barely 10 mm. long; berries round, with
a rather slender style; stigma short.
Type from Peru, Meyen 150.
Piper perareolatum C. DC. Bot. Jahrb. 40: 246. 1908.
A resin-scented, yellowish-villous shrub or small tree, 3-6 meters
high; leaves round-ovate, acuminate, subequilaterally subtruncate
at the base or cordate, with the margin abruptly contracted to the
FLORA OF PERU 207
petiole, 18-21 cm. long, 15-16 cm. wide, submultiple-nerved from the
lower third, the nerves about 5x2, bullulate-lacunose, glabrous
above, the petiole 3 cm. long; spikes curved, 5-8 cm. thick, 250-300
mm. long, or larger; peduncle thick, 2 cm. long, pale-granular.
Neg. 11016.
Junin: Huacapistana, Weberbauer 2195, type. Huanuco: Muna,
2,400 meters, 4340.
Piper perenense Trelease, sp. nov.
Frutex subcanescens 3-4-metralis, internodiis floriferis brevibus
gracilibus subcrispe pubescentibus; folia lanceolata vel elliptica
anguste falcato-acuminata basi subaequilaterali acuta 14-16 cm.
longa 5.5-6.5 cm. lata e dimidio inferiore pinnatinervia, nervis 5-6x2,
supra basibus pilorum fugacium granuloso-scabra, subtus ad nervos
breviter hirsuta, petiolo vix 10 mm. longo exalato; spicae curvae,
juvenes 20 mm. longae 2 mm. crassae, pedunculo scabrido 3 mm.
longo; bracteae rotundo-subpeltatae.
Junin: Colonia Perene", 600 meters, Killip & Smith 25108 (type
in U. S. Nat. Herb.). Puerto Bermudez, Killip & Smith 26442.
Piper perenense var. cupulifolium Trelease, var. nov.
Margine folii basi in cupulam conicam 5 mm. profundam connato.
Junin: La Merced, 700 meters, Killip & Smith 23698 (U. S. Nat.
Herb., type), growing with the normal form of the species.
Piper pernigricans Trelease, sp. nov.
Frutex 3-metralis glaber glanduloso-granulosus nigrescens, inter-
nodiis floriferis gracilibus brevibus; folia lanceolata vel suboblonga
sensim acuminata basi aequilaterali acuta 12-15 cm. longa 3.5-4
cm. lata ubique pinnatinervia, nervis circa 10x2, nervis e dimidio
inferiore magis elongatis, in sicco lucida, petiolo 10 mm. longo
inferne alato; spicae juvenes 20 mm. longae 1 mm. crassae subrectae.
Loreto: Mishuyacu, 100 meters, Killip & Smith 29976 (type in
U.S. Nat. Herb.).
Piper perpunctulatum Trelease, sp. nov.
Frutex glaber 3-metralis, internodiis floriferis gracilibus brevibus;
folia elliptico-lanceolata acuminata basi inaequilateralia, latere
longiore subabrupte acuto, 14 cm. longa 7-8 cm. lata e dimidio in-
feriore submultiplinervia, nervis 4+5, in sicco tenuia subtus minute
glanduloso-punctata, petiolo 7+3 mm. longo non vaginante; spicae
208 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
60 mm. longae 3 mm. crassae, pedunculo 10 mm. longo; bracteae
rotundo-subpeltatae ciliolatae.
Loreto: Yurimaguas to Balsapuerto, 135-150 meters, Killip &
Smith 28090 (type in U. S. Nat. Herb.), 2833. Yurimaguas, Killip
& Smith 28203.
Piper perpurgatum Trelease, sp. nov.
Frutex metralis, internodiis floriferis brevibus subgracilibus primo
crispo-pubescentibus glabrescentibus; folia elliptico-lanceolata acu-
minata basi acuta 10-14 cm. longa 3-5 cm. lata ubique pinnatinervia,
nervis circa 10x2, supra lepidoto-punctulata, subtus ad nervos
crispo-pubescentia, petiolo 5-10 mm. longo, pubescente, longioribus
late alatis; spicae patentes 15 mm. longae 4 mm. crassae, pedunculo
5 mm. longo pubescente; bracteae inconspicuae; ovarium subteres
depressum vel mucrone protruso terminatum, stigmatibus 3 in
mucrone insertis.
Loreto: San Antonio, Rio Itaya, 110 meters, Killip & Smith
29455 (type in U. S. Nat. Herb.), 29414, 29461. Soledad, Killip
6 Smith 29707.
Piper peruvianum (Miq.) C. DC. in DC. Prodr. 16, pt. 1: 283.
1869. Artanthe peruviana Miq. Lond. Journ. Bot. 4: 461. 1866.
P. pseudo-peruvianum C. DC. op. cit. 282. A. peruviana var.
laevigata Miq. in herb.
A glandular-granular, glabrous shrub; leaves lance-elliptic, acu-
minate, acute-based, 16-19 cm. long, 8-10 cm. wide, pinnately
nerved from the lower half, the nerves 4-5x2; petiole 10-15 mm.
long; spikes 100-120 mm. long, 3 mm. thick; peduncle 10-15 mm.
long; bracts triangular-subpeltate; berries quadrate; stigmas sessile.
Junin: Pangoa, Mathews 1145 (type), 1147. Mainas, Poeppig
1037. Huanuco: Casapi, Mathews (1713), 1703 in Herb. Bentham.
Loreto: Santa Rosa, Killip & Smith 28783, Balsapuerto, Killip &
Smith 28385.
Piper pervillosum Trelease, sp. nov.
Frutex undique spicis exceptis rubescenti-villosus, internodiis
floriferis comparate gracilibus ut videtur elongatis; folia elliptico-
obovata obtusa basi inaequilaterali cordata, sinu laterali, lobo majore
auriculate producto, 29 cm. longa 12 cm. lata e dimidio inferiore
subpinnatinervia, nervis 5x2, nervis supplementariis in auricula
dispersis, petiolo 4 cm. longo alato; spicae 90 mm. longae et ultra
FLORA OF PERU 209
4 mm. crassae, pedunculo gracili 3 cm. longo et ultra; bracteae rotun-
do-subpeltatae.
San Martin: San Roque, in forest, 1,350-1,500 meters, Llewelyn
Williams 6963 (herb. Field Mus. No. 622,979, type).
Piper pervulgatum Trelease, sp. nov.
Frutex 2-3-metralis, internodiis floriferis brevibus subgracilibus
adpresso-hispidis; folia subelliptica anguste acute acuminata basi
inaequilaterali acuta 11-15 cm. longa 4.5-6 cm. lata e dimidio in-
feriore pinnatinervia, nervis 5-6x2, supra minute sed conspicue
granuloso-scabra, subtus ad nervos incurvo-hispida, petiolo 5+2-3
mm. longo hispido exalato; spicae 90 mm. longae 3 mm. crassae,
pedunculo 6 mm. longo scabro; bracteae parvae rotundo-subpeltatae
ferrugineo-ciliolatae.
Loreto: Mishuyacu near Iquitos, in clearing, 100 meters, G. King
468 (herb. Field Mus. No. 629,836, type). San Antonio, Rio Itaya,
110 meters, Killip & Smith 29312. Pebas on the Amazon River,
in forest, Williams 1889, 1594. "Cordoncillo."
Piper phegophyllum Trelease, sp. nov.
Frutex vel arbuscula 3-5-metralis glabra plus minusve nodosa
gracilis, internodiis floriferis gracilibus brevibus; folia elongato-
elliptica acuminata basi fere aequilaterali acuta 15-20 cm. longa 6.5-
8 cm. lata ubique pinnatinervia nervis gracilibus prominentibus
circa 16x2, subtus dense nigro-punctulata, petiolo vix 10 mm. longo
exalato; spicae 50-60 mm. longae 2 mm. crassae, pedunculo gracili
7 mm. longo; bracteae rotundo-subpeltatae laceratae glabrae.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26479
(type in U. S. Nat. Herb.).
Piper phthinotrichon (Miq.) C. DC. in DC. Prodr. 16, pt. 1:
298. 1869. Artanthe phthinotricha Miq. Syst. 527. 1844; 111. Piper.
pi. 90.
A delicate, glabrous or transiently puberulent, glandular-granular
shrub; leaves elliptic-lanceolate, sharp-acuminate, subacute at the
base, 6 cm. long, 2-3 cm. wide, pinnately nerved throughout; petiole
5 mm. long, winged; spikes 15 mm. long, 3 mm. thick; peduncle 10
mm. long, filiform; bracts cucullate; berries globose; stigmas 3,
sessile.
Loreto: Mainas, Poeppig 2289 (type), 2283. Yurimaguas, Killip
6 Smith 27619. Lower Rio Huallaga, Killip & Smith 29260.
210 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Ayacucho: El Puquio, Poeppig D1736. Huanuco: Cochero, Poeppig.
Junin: Chancfcamayo Valley, Schunke 404-
Piper phthinotrichon var. san-roqueanum Trelease, var. nov.
Frutex humilis nodosus glaber granulosus, internodiis floriferis
brevissimis gracilibus; folia lanceolata vel lanceolato-ovata sub-
acuminata basi subabrupte acuta 4.5-5.5 cm. longa 2 cm. lata e
dimidio inferiore pinnatinervia, nervis longioribus 4x2 sursum con
brevioribus junctis, petiolo vix 5 mm. longo caduce alato; spicae
fructiferae 20 mm. longae 5 mm. crassae breviter caudatae, pedunculo
vix 5 mm. longo in statu fructifero refracto; bracteae crescenticae;
baccae globosae, stigmatibus fere sessilibus vel in mucrone brevi
insertis.
San Martin: San Roque, in forest, 1,350-1,500 meters, Llewelyn
Williams 7218 (herb. Field Mus. No. 624,599, type).
Piper pichisense Trelease, sp. nov.
Frutex 2-metralis, internodiis floriferis brevibus crassiusculis
dense flavide incurvo-pilosis; folia elliptica abrupte acuminata basi
fere aequilaterali cordata 14-16 cm. longa 7-8 cm. lata e dimidio
inferiore submultiplinervia, nervis 5-6x2, in sicco tenuia, supra
pilis crispis flavidis pilosa, subtus ad nervos sparse hirsuto-villosa,
petiolo 5-10 mm. longo flavo-barbato; spicae juveniles 50 mm. longae
4 mm. crassae, pedunculo 15-20 mm. longo plus minusve crispo-
hirtello; bracteae triangulari-subpeltatae ciliolatae.
Junin: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
26012 (type in U. S. Nat. Herb.).
Piper planipes Trelease, sp. nov.
Frutex glaber 3-metralis, internodiis floriferis modice gracilibus
et elongatis; folia ovata acuminata basi rotundata vel subtruncata
supra petiolum minute cordulata 15 cm. longa 8-10 cm. lata e
dimidio inferiore submultiplinervia, nervis 4x2, in sicco caerules-
centia, petiolo 10-15 mm. longo supra late sulcato; spicae 60 mm.
longae 3 mm. crassae, pedunculo gracili 15 mm. longo; baccae
obpyramidales apice subacutae; stigmata 3 minuta sessilia.
Junin: Colonia Perene", 680 meters, Killip & Smith 24928 (type
in U. S. Nat. Herb.).
Piper planipes var. amplum Trelease, var. nov.
Frutex 3-4-metralis; folia interdum basi breviter cordata 17-22
cm. longa 10 cm. lata, petiolo 2-4 cm. longo basi infima carnoso-
FLORA OF PERU 211
verrucoso; spicae 25-30 mm. longae 4 mm. crassae, pedunculo 7 mm.
longo; bracteae crescenticae; baccae obconicae apice plus minusve
apicatae, stigmatibus minutis sessilibus.
Junin: Schunke Hacienda, La Merced, 1,400-1,700 meters,
Killip & Smith 24721 (type in U. S. Nat. Herb.).
Piper pleninervosum Trelease, sp. nov.
Frutex nodosus 2-3-metralis nervis folii subtus adpresso-pub-
escentibus exceptis glaber, ramulis petiolisque papillosis vel granu-
losis; folia lanceolata acuminata basi subacuta vel uno latere paullo
breviora et obtusa 9-12 cm. longa 3-4 cm. lata e 2 tertiis inferioribus
pinnatinervia, nervis 5x2, petiolo vix 5 mm. longo non vaginante;
spicae 90 mm. longae 2-3 mm. crassae, pedunculo 5-10 mm. longo;
bracteae rotundo-subpeltatae ciliolatae; baccae teretes, stigmatibus
minutis sessilibus.
Loreto: Puerto Arturo, 135 meters, Killip & Smith 27875 (type
in U. S. Nat. Herb.).
Piper politaereum Trelease, sp. nov.
Frutex vel arbuscula glabra nodosa 3-5-metralis, internodiis
floriferis breviusculis gracilibus; folia elliptica vel paullo subovata
anguste acuminata basi inaequilaterali acuta 12-16 cm. longa 5-8
cm. lata e dimidio inferiore pinnatinervia, nervis 5x2, in sicco
tenuia fusca, subtus minute granulosa lucida, petiolo 5+1-4 mm.
longo exalato; spicae juvenes 35 mm. longae 2 mm. crassae, pedunculo
5 mm. longo; bracteae parvae rotundo-subpeltatae ciliolatae.
Junin: Schunke Hacienda, San Ramon, 1,400-1,700 meters,
Killip & Smith 24545 (type in U. S. Nat. Herb.), 24553. La
Merced, Killip & Smith 23905.
Piper politaereum var. elongatum Trelease, var. nov.
Frutex 3-metralis; folia lanceolato-elliptica 22 cm. longa 7 cm.
lata, subtus ad nervos obscure adpresso-pubescentia, nervis superi-
oribus separatis vel ab aliis remotis.
Loreto: Soledad, 110 meters, Killip & Smith 29587 (type in
U. S. Nat. Herb.).
Piper politaereum var. mucronatum Trelease, var. nov.
Frutex 3-4-metralis glaber; folia elliptica 22 cm. longa 8-9 cm.
lata; spicae juvenes 55 mm. longae 2 mm. crassae cuspidatae,
pedunculo gracili 6-8 mm. longo.
Loreto: San Antonio, Rio Itaya, 110 meters, Killip & Smith
29460 (type in U. S. Nat. Herb.).
212 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper politum C. DC. Ann. Cons. Jard. Geneve 21: 237. 1920.
A shrub, glabrous except for obscure puberulence of the twigs;
leaves lance-oblong, short-acuminate, almost equilaterally acute at
the base, 12-13 cm. long, 3.5-4 cm. wide, pinnately nerved from
below the upper third, the nerves 4-5x2, glossy; petiole 10-15 mm.
long; spikes gently curved, 50 mm. long, 3 mm. thick; peduncle
5-10 mm. long; bracts long, black with a narrow pale margin;
berries depressed-globose; stigmas 3, sessile. Neg. 25041.
San Martin: Tarapoto, Spruce 4137, type. Lore to: Mishuyacu,
Killip & Smith 29976.
Piper politum var. pallidibracteum Trelease, var. nov.
Folia 13 cm. longa 3.5 cm. lata; spicae 6.5 mm. longae 3 mm.
crassae, pedunculo 10 mm. longo; bracteae pallidae.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
7639 (herb. Field Mus. No. 629,703, type). "Cordoncillo."
Piper ponganum Trelease, sp. nov.
Frutex humilis diffusus vix frutescens, internodiis floriferis 3-4
mm. crassis rigide fulvo-villosis; folia elliptica acuminata basi
cordata 15 cm. longa 7-8 cm. lata e dimidio inferiore pinnatinervia,
nervis 5-7x2, subtus ad nervos laxe villosa, petiolo 1-2 cm. longo
vaginante plus minusve villoso; spicae 25-30 mm. longae 5 mm.
crassae cuspide puberulo 10 mm. longo 1 mm. crasso terminatae,
pedunculo crasso 5 mm. longo villoso; bracteae crescenticae ciliatae;
ovarium obconicum, stylo longo filiformi, stigmatibus 3 recurvis.
Loreto: Pongo de Manseriche, 200 meters, Ynes Mexia 6123
(herb. Univ. Calif., type).
Piper pontis Trelease, sp. nov.
Arbuscula 4-5-metralis, internodiis floriferis gracilibus modice
brevibus fere glabris; folia lanceolata et sensim acuminata vel
subovato-elliptica et subabrupte acuminata basi subaequilaterali
obtusa vel uno latere majora et acuta 8.5-12 cm. longa 4-5 cm. lata
e dimidio inferiore pinnatinervia, nervis 4x2, supra microscopice
granuloso-asperata sed in statu adulto laevia, viridia plus minusve
rugosa, subtus ad nervos obscure adpresso-pubescentia, petiolo 10
mm. longo exalato; spicae juvenes 30 mm. longae 2 mm. crassae
acutatae, pedunculo 5 mm. longo; bracteae rotundo-subpeltatae.
Junin : Near Perene Bridge, Rio Paucartambo Valley, 700 meters,
Killip & Smith 25294 (type in U. S. Nat. Herb.).
FLORA OF PERU 213
Piper portalense Trelease, sp. nov.
Arbuscula 3-5-metralis nodosa, internodiis floriferis brevibus
gracilibus transiente plus minusve crispo-pubescentibus et pallido-
granulosa; folia lanceolato-elliptica anguste acuminata basi inaequi-
lateralia, latere breviore rotundato, 12-16 cm. longa 4-6 cm. lata e
dimidio inferiore pinnatinervia, nervis 5x2, subtus pallidiora ad
nervos griseos adpresso-pubescentia, petiolo 8+2 mm. longo crispo-
pubescente non vaginante; spicae 100 mm. longae 2-3 mm. crassae,
pedunculo 8 mm. longo glabrato; bracteae rotundo-subpeltatae
ciliatae; baccae teretes, stigmatibus sessilibus.
Loreto: Portal, lower Rio Huallaga, 135 meters, Killip & Smith
29258 (type in U. S. Nat. Herb.).
Piper porveniricola Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis brevibus gracilibus sub-
tomentoso-hispidis ; folia lanceolata sensim longiacuta basi in-
aequilaterali acuta 15 cm. longa 5 cm. lata fere ubique pinnatinervia,
nervis longioribus 6x2, venosa sed tenuia, supra granuloso-scabra,
subtus ad nervos adscendenti-strigosa, petiolo 5+5 mm. longo;
spicae juveniles 15 mm. longae 2 mm. crassae obtusae, pedunculo
vix 5 mm. longo; bracteae rotundo-subpeltatae.
Junin: Porvenir, Pichis Trail, 1,500-1,900 meters, Killip &
Smith 2594.0 (type in U. S. Nat. Herb.).
Piper positum Trelease, sp. nov.
Frutex 3-metralis nodosus glaber, internodiis floriferis brevibus
crassis fistulosis; folia elliptica subacuminata basi subaequilaterali
acuta 16-17 cm. longa 7-8 cm. lata ubique pinnatinervia, nervis
circa 8x2, in sicco firmo-papyracea sublucida subtus nigro-punctu-
lata, petiolo 10-15 mm. longo exalato; spicae leviter curvae, floriferae
40 mm. longae 2 mm. crassae, pedunculo gracili 10 mm. longo;
bracteae rotundo-subpeltatae ciliatae, stigmatibus sessilibus.
Loreto: Portal, Rio Huallaga, 135 meters, Killip & Smith 29256
(type in U. S. Nat. Herb.).
Piper posusanum Trelease, sp. nov.
Frutex compactus 1.5 m. altus nodosus, internodiis floriferis
brevibus gracilibus pilis brevibus retrorsis adpressis canescentibus;
folia lanceolata subacuminate attenuata basi subaequali subacuta
parva 6-8 cm. longa 1.5-2 cm. lata e dimidio inferiore pinnatinervia,
nervis 4-5x2, utrinque pilis brevibus adpressis canescentia sed vix
214 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
asperata, petiolo brevissimo vix 3 mm. longo lit nervis subtus plus
minusve incurvo-hirsuta; spicae oppositifoliae valde curvatae 70
mm. longae 3 mm. crassae, pedunculo vix 10 mm. longo adpresso-
canescente; bracteae minutae rotundo-subpeltatae griseae, floribus
sessilibus perfectis; stigmata 2-3 minuta sessilia; baccae rubescentes
parvae cum rhachi subelongatae.
Huanuco: Posuso, 600 meters, Macbride 4609 (herb. Field Mus.
No. 535,686, type).
Piper pothophyllum Trelease, sp. nov.
Frutex, internodiis floriferis modice gracilibus et longis dense
crispo-pubescentibus; folia oblongo-lanceolata vel elliptico-lanceolata
sensim acuta basi obliqua, latere longiore cordulato, 25-27 cm. longa
6-9 cm. lata ubique pinnatinervia, nervis circa 20x2, subtus palli-
diora ad nervos pubescentia, petiolo 4 cm. longo alato crispo-puber-
ulo; spicae rubrae 50 mm. longae 3 mm. crassae acutatae, pedunculo
gracili 4 cm. longo puberulo; bracteae transversae glabrae; ovarium
angulatum, stigmatibus 3 subsessilibus.
Loreto: Victoria on the Amazon River, in forest, Llewelyn Wil-
liams 2998 (herb. Field Mus. No. 608,343, type). "Cordoncillo."
Piper praeclarum Trelease, sp. nov.
Frutex vix 1.5 m. altus pilis brevibus intertextis ubique villosus;
folia lanceolato-elliptica sensim acuminata basi inaequali obtusa
13-15 cm. longa 5-6 cm. lata e dimidio inferiore pinnatinervia, nervis
circa 5x2, petiolo brevissimo 3+3 mm. longo exalato; spicae oppo-
sitifoliae valde curvae 60 mm. longae 3 mm. crassae, pedunculo
elongate 15-20 mm. longo; bracteae triangulari- vel lunulato-
subpeltatae ciliatae, floribus sessilibus perfectis; stigmata 3 brevia
sessilia.
Junin: Hacienda Schunke, La Merced, 1,200 meters, in forest,
Macbride 5623 (herb. Field Mus. No. 536,662, type).
Piper procerum Trelease, sp. nov.
Arbor glabra 10-13-metralis; folia rotundato-elliptica obtusa
basi inaequilateralia, sinu laterali clauso, latere majore petiolo
aequilongo, 31 cm. longa 24 cm. lata e 3 quartis inferioribus sub-
pinnatinervia, nervis 4+5 pluribus supplementariis in auricula
dispersis, petiolo 8 cm. longo alato basi verrucoso; spicae 200 mm.
longae 3 mm. crassae, pedunculo 15 mm. longo; bracteae rotundo-
subpeltatae ciliatae; baccae teretes apice depressae; stigmata 3
sessilia.
FLORA OF PERU . 215
Junin: Schunke Hacienda, La Merced, 1,400-1,700 meters,
Killip & Smith 24636 (type in U. S. Nat. Herb.).
Piper propinquum C. DC. Journ. Bot. 4: 167. 1866.
A glabrous, glandular-granular shrub ; leaves lance-elliptic, sharp-
acuminate, acute-based, 15-17 cm. long, 6-7 cm. wide, submultiple-
nerved from the lower half, the nerves 4x2; petiole 10 mm. long;
spikes 90 mm. long, 3-4 mm. thick, the peduncle 5-10 mm.
long; bracts crescentic-subpeltate, yellow-ciliate; berries globose,
with a slender style; stigmas spreading. Negs. 7231, 8421.
San Martin: Tarapoto, Spruce 4032, type. San Roque, 1,500
meters, Williams 7229. Junin: Pangoa, Mathews 1147. Yapas,
Killip & Smith 25577. Chanchamayo Valley, Schunke 488.
Piper propinquum var. brevistylum Trelease, var. nov.
Stylo brevi crasso, stigmatibus latis differt.
Huanuco: Cochero, Poeppig 1037 (type in herb. Vienna). Toca-
che, Poeppig in 1830. Without locality; Poeppig 1337(1).
Piper propinquum var. grande Trelease, var. nov.
Subarborescens; folia 20-25 cm. longa 9-14 cm. lata; spicae
juvenes curvae 20 mm. longae 3 mm. crassae, bracteis conspicue
brunneo-marginatis, pedunculo 1 cm. longo.
Loreto: Yurimaguas to Balsapuerto, 135-150 meters, Killip &
Smith 28109 (type in U. S. Nat. Herb.). Santa Rosa, Killip
& Smith 28734. Puerto Arturo, Killip & Smith 27757.
Piper pseudobarbatum C. DC. Bot. Jahrb. 40: 249. 1908.
A glabrous, compact shrub; leaves ovate, subacute, rounded at
the base or subcordulate, 6-9 cm. long, 3.5-5 cm. wide, pinnately
nerved from the lower half, the nerves 3x2; petiole 5 mm. long;
spikes 30-40 mm. long, 3 mm. thick; peduncle 5 mm. long; bracts
round-subpeltate, ciliolate; stigmas sessile. Neg. 11025.
Ancash: Puccha Valley in thickets along river, 2,700 meters,
Weberbauer 3738, type.
Piper pseudo-callosum Trelease, sp. nov.
Frutex 2.5 m. altus subnodosus, internodiis floriferis longis
gracilibus glabris; folia lanceolato-oblonga caudata basi subinaequali
acuta et bicallosa 14-16 cm. longa 5-6 cm. lata e dimidio inferiore
pinnatinervia, nervis circa 6x2, rugosa, subtus ad nervos prope
marginem adpresso-pubescentia, petiolo gracili 1-2.5 cm. longo
216 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
glabro, petiolis longioribus ad medium alatis; spicae oppositifoliae
comparate crassis et brevibus 35-40 mm. longae 5 mm. crassae,
pedunculo gracili 15 mm. longo glabro; bracteae cucullato-inflexae
glabrae, floribus sessilibus perfectis; stigmata 2-3 brevia sessilia;
baccae breviter cylindraceae vel pressu rhombicae.
Huanuco: Huacachi near Muna, 1,950 meters, in forest, Mac-
bride 4103 (herb. Field Mus. No. 535,172, type). Trail to Tambo de
Vaca, 2,400 meters, in forest, 4333.
Piper pseudo-churumayu (Kunth) C. DC. in DC. Prodr. 16,
pt. 1: 288. 1869. Steffensia pseudo-churumayu Kunth, Linnaea 13:
662. 1839. Artanthe pseudo-churumayu Miq. Syst. 407. 1844; 111.
Piper, pi. 75.
A shrub, glabrous except for the appressed-pubescent or puberu-
lent nerves beneath; leaves ovate, acuminate, rounded at the base
with one side slightly longer and abruptly decurrent to the petiole,
26 cm. long, 16 cm. wide, pinnately nerved from somewhat more
than the lower half, the nerves 5-6x2; petiole 20+2-5 mm. long,
winged; spikes 140 mm. long, 3 mm. thick; peduncle 25 mm.
long; stigmas sessile. Neg. 11026.
Type from subandine Peru, Poeppig 35. Huanuco: Cochero,
Poeppig 166, DlSOl.Loreto: Mainas, Yurimaguas, Poeppig.
Ucayali River, Tessmann 3209. Also in Brazil and Venezuela.
Piper pseudo-curvatum Trelease, sp. nov. P. munchanum var.
magnifolium C. DC. in DC. Prodr. 16, pt. 1: 259. 1869.
Frutex glaber vel ad nervos paginae inferioris folii obscure puber-
ulus; folia elliptico-subobovata abrupte obtuso-acuminata basi
cuneata 40 cm. longa 22 cm. lata undique pinnatinervia, petiolo
crassissimo lamina decurrente alato; spicae 50-200 mm. longae 10-15
mm. crassae leviter curvae, pedunculo 30 mm. longo 5 mm. crasso;
baccae rotundatae, stylo breviusculo, stigmatibus subparvis.
Huanuco: Cochero, Poeppig 1312, type. San Martin: Tarapoto,
Spruce 4829.
Piper pseudo-flexuosum Trelease, sp. nov.
Frutex crispo-hirtellus; folia lanceolato-ovata acuminata basi
inaequilaterali cordulata 13 cm. longa 6 cm. lata e dimidio inferiore
pinnatinervia, nervis 5x2, utrinque pubescentia, petiolo 10+3 mm.
longo; inflorescentia non visa.
Type from subandine Peru, Poeppig (herb. Vienna, type;
herb. Berol.).
FLORA OF PERU 217
Piper pseudo-matico Trelease, sp. nov.
Frutex 3-metralis conspicue nodosus, internodiis floriferis modice
elongatis et crassis primo dense retrorse villoso-hirsutis, serius glabres-
centibus et lenticellis parvis conspersis; folia elliptica vel elliptico-
obovata sensim longiuscule acuminata basi inaequaliter obtusa 12-
15 cm. longa 5-6 cm. lata e dimidio inferiore pinnatinervia, nervis
valde prominentibus 6-7x2, rugosissima granuloso-scabra, subtus ad
nervos conspicue hirsuta, petiolo 10 mm. vel 10+3 mm. longo in-
curvo-hirsuto exalato; spicae oppositifoliae fere rectae vel arcuatae
70 mm. longae 3 mm. crassae in statu florifero, spicae fructiferae
rectae 90 mm. longae 4 mm. crassae, pedunculo brevi 5-6 mm. longo
plus minusve hamato-hirsuto ; bracteae rotundo-subpeltatae ciliatae,
floribus perfectis sessilibus; stigmata 3 brevia sessilia; baccae nigrae
breviter cylindraceae glabrae.
Huanuco: Mito, 2,700 meters, in canyon along a stream, Macbride
& Featherstone 1488 (herb. Field Mus. No. 517,987, type), 3327.
Junin: Carpapata, Killip & Smith 24460. "Matico."
Piper pseudo-mollicomum C. DC. in DC. Prodr. 16, pt. 1:
278. 1869. P. Poeppigii Steud. Norn. Bot. ed. 2. 2: 342. 1840.
Steffensia Poeppigii Kunth, Linnaea 13: 646. 1839. Artanthe
Poeppigii Miq. Syst. 437. 1844.
A shrub with retrorsely appressed-hairy twigs; leaves elliptic-
subovate, acuminate, rounded at the base with one side shorter, 12-
14 cm. long, 5.5-7 cm. wide, pinnately nerved from somewhat more
than the lower half, the nerves 5-6x2, soft-hairy but scabrous above
from persistent hair bases; petiole 10+2 mm. long, hirsute;
spikes 100 mm. long, 2-3 mm. thick; peduncle 10 mm. long, hispid.
Neg. 8422.
Loreto: Mainas, Yurimaguas, Poeppig 2290 (type), D2290. Rio
Blanco, Tessmann 3011 .Huanuco : Cochero, Dombey 928. With-
out locality, Gay in 1839-1840; Mathews 1693, 1145 bis, 1146, 1705.
Piper pseudo-mollicomum var. dichotomum C. DC. in
DC. Prodr. 16, pt. 1: 279. 1869.
A young, less pubescent form with oblanceolate leaves 9.5 cm.
long and 3.5 cm. wide.
Type from subandine Peru, Poeppig.
Piper pseudo-munchanum Trelease, sp. nov. P. mun-
chanum Auct.
Frutex glaber; folia lanceolato-elliptica utrinque acuta 28 cm.
longa 12 cm. lata ubique pinnatinervia, petiolo 2 cm. longo alato;
218 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
spicae 50 mm. longae 10 mm. crassae, pedunculo 10 mm. longo
curvo; baccae obovoideae, stylo longo, stigmatibus parvis patentibus.
San Martin: Tarapoto, Spruce 4341, type.
Piper puberulescens Trelease, sp. nov.
Frutex 3-metralis nervis folii subtus subpuberulis exceptis glaber;
folia late lanceolata subacuta basi breviter subtruncato-cordata et
aequilateralia vel distincte obliqua 25-30 cm. longa 9-12 cm. lata
undique pinnatinervia, nervis e dimidio inferiore magis protractis,
petiolo ad 11 cm. longo membranaceo-alato ; spicae juvenes 15 mm.
longo 2 mm. crassae, pedunculo recurvo aequilongo.
Loreto: Yurimaguas, 135 meters, Killip & Smith 29057 (type in
U. S. Nat. Herb.).
Piper pubibaccum C. DC. Bot. Jahrb. 40: 251. 1908.
A shrub, 3.5 meters high, glabrous except for the hirtellous nerves
beneath; leaves obliquely round-ovate, abruptly short-acuminate,
somewhat shorter and rounded on one side, the longer side round-
auricled with the auricle about equaling the petiole, 22 cm. long,
18 cm. wide, subpinnately nerved from below the upper third, the
nerves 5+7-8; petiole 5+1 cm. long, winged; bracts crescentic-
subpeltate; berries round, puberulent. Neg. 11029.
Huanuco: Between Monzon and Rio Huallaga, 600-700 meters,
in forest, Weberbauer 3581, type. Junin: La Merced, in forest, 600
meters, 5518.
Piper pubinervulum C. DC. Ann. Cons. Jard. Geneve 21:
240. 1923.
A shrub, glandular-granular, more or less persistently crisp-sub-
villous except for the upper leaf surface; leaves elliptic or suboblong,
abruptly subacuminate, acute-based, 16 cm. long, 6-8 cm. wide,
pinnately nerved from below the upper third, the nerves about 6x2;
petiole 10 mm. long; spikes 140 mm. long, 10 mm. thick; peduncle
15-20 mm. long; bracts round-subpeltate ; berries round, with a short
thick style; stigmas broad and large, spreading. Neg. 11032.
Piura: Huancabamba, Weberbauer 6093, type.
Piper pullispicum Trelease, sp. nov.
Frutex subhumilis, internodiis superioribus gracilibus sub-
elongatis crispo-pubescentibus; folia lanceolata ad elliptico-acuminata
basi acuta uno latere paullo breviora 16-25 cm. longa 6-11 cm. lata
e dimidio inferiore pinnatinervia, nervis 5-6x2, in sicco viridia
FLORA OF PERU 219
lucida, subtus ad nervos substrigosa, petiolo 15-30 mm. longo
crispo-pubescente; spicae 80 mm. longae 3-4 mm. crassae, primo
obscure rubrae, serius brunnescentes, pedunculo 10-15 mm. longo
sparse pubescente; bracteae rotundo-subpeltatae ciliolatae.
Loreto: Pongo de Manseriche, 280 meters, Ynes Mexia 6352
(type in herb. Univ. Calif.).
Piper punctatum R. & P. Fl. 1: 36. pi. 62. 1798. Artanthe
punctata Miq. Syst. 522. 1844.
A glabrous, black-punctulate shrub scarcely distinguishable from
P. heterophyllum except that its spikes are straight and nearly erect.
Huanuco: Iscutuna and Pillao, Ruiz & Pavon, type.
Piper punctulatifolium Trelease, sp. nov.
Frutex glaber valde nigro-punctulatus, internodiis superioribus
gracilibus elongatis; folia elliptica anguste acuminata basi acuta
18-19 cm. longa 8-11 cm. lata, superiora basi inaequilateralia, in-
feriora basi aequilateralia, e dimidio inferiore pinnatinervia, nervis
6x2, in sicco tenuia subtus lucida, petiolo 10+5 mm. longo superne
exalato, foliorum latiorum 30-40 mm. longo et fere ad apicem alato;
inflorescentia non visa.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28783 (type in
U. S. Nat. Herb.).
Piper puniceum Trelease, sp. nov.
Frutex vel arbuscula nodosa, in statu juvenili ubique dense
rubescenti-sericeo-villosa; folia oblique lanceolato-elliptica anguste
acuminata basi rotundata 14-15 cm. longa 6 cm. lata e dimidio
inferiore pinnatinervia, nervis 6x2, supra inter pilos dense fusco-
granulosa, petiolo 3+3 mm. longo; spicae juvenes 20 mm. longae
2 mm. crassae; bracteae rotundo-subpeltatae.
Junin: Perene, 1,500 meters, Carlos Schunke 236 (herb. Field
Mus. No. 627,350, type), 196.
Piper quimirianum Trelease, sp. nov.
Frutex nodosus ramosus 2-metralis, internodiis floriferis brevibus
cito crassioribus transiente crispo-pubescentibus; folia elliptica vel
elliptico-subobovata acuminata basi subaequilaterali sensim acutata
10-12 cm. longa 5-6 cm. lata e 2 tertiis inferioribus pinnatinervia,
nervis 5x2, in sicco firmo-papyracea caeruleo-viridia, subtus ad
nervos subobscure adpresso-pubescentia, petiolo 3+2 mm. longo
220 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
exalato; spicae juvenes 40 mm. longae 2 mm. crassae, pedunculo
crasso 5 mm. longo glabro; bracteae rotundo-subpeltatae fimbriatae.
Junin: East of Quimiri Bridge, La Merced, 800-1,300 meters,
Killip & Smith 23904 (type in U. S. Nat. Herb.).
Piper rasile Trelease, sp. nov.
Frutex 3-4-metralis, internodiis floriferis subgracilibus elongatis
incurvo-hispidis; folia elliptica acuminata basi obtusa uno latere
longiora et interdum acuta 14-15 cm. longa 5 cm. lata e 2 tertiis
inferioribus pinnatinervia, nervis 6x2, granuloso-scabra, subtus ad
nervos incurvo-hirsuta, petiolo 10+2 mm. longo hispido; spicae 90
mm. longae 3 mm. crassae, pedunculo gracili 10 mm. longo scabrido.
Junin : Puerto Bermudez, 375 meters, Killip & Smith 26404 (type
in U. S. Nat. Herb.).
Piper ratisportanum Trelease, sp. nov.
Frutex glaber glanduloso-granulosus 3-4-metralis, internodiis
floriferis modice crassis et elongatis; folia subovato-elliptica breviter
acuminata basi paullo inaequilateralia basi infima acuta 30 cm.
longa 15 cm. lata ubique pinnatinervia, nervis circa 16x2, petiolo
5+5 mm. longo exalato; spicae juveniles 85 mm. longae 6 mm.
crassae mucronatae, pedunculo 10 mm. longo; bracteae rotundo-
subpeltatae ciliatae; ovarium submucronatum, stigmatibus 3 in
mucrone insertis.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28570
(type in U. S. Nat. Herb.).
Piper recensitum Trelease, sp. nov.
Frutex vel arbuscula glabra, internodiis floriferis brevibus
gracilibus; folia subovato-elliptica anguste acuminata basi fere
aequilaterali subcuneata 11-14 cm. longa 4-5 cm. lata e dimidio
inferiore pinnatinervia, nervis circa 6x2, in sicco tenuia viridia,
petiolo vix 5 mm. longo exalato; spicae floriferae 20 mm. longae 2
mm. crassae, pedunculo 3-4 mm. longo; bracteae rotundo-sub-
peltatae ciliolatae.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28810 (type in
U. S. Nat. Herb.), 28728, 28748.
Piper recreoense Trelease, sp. nov.
Frutex humilis glaber plus minusve conspicue nigro-punctulatus,
internodiis floriferis gracilibus modice elongatis; folia anguste lanceo-
lata utrinque sensim acutata 9-10 cm. longa 1.5-2 cm. lata ubique
FLORA OF PERU 221
pinnatinervia, nervis 10-12x2, in sicco tenuia viridia, petiolo gracili
5 mm. longo exalato; spicae 15 mm. longae 4 mm. crassae primo
mucronatae, pedunculo refracto filiform! 10 mm. longo; bracteae
inconspicuae subpeltatae; baccae subglobosae, stigmatibus 3 majus-
culis fere sessilibus.
Loreto: Recreo, Yurimaguas, 155-210 meters, Llewelyn Williams
3928 (herb. Field Mus. No. 622,942, type).
Piper redactum Trelease, sp. nov.
Frutex humilis, ramulis tomentosis; folia acuta vel subacuminate
acutata basi cordata 5 cm. longa 2.5-3 cm. lata e dimidio inferiore
pinnatinervia, nervis 2-3x2, supra minute pubescentia papillosa,
subtus dense crispo-pubescentia, petiolo 5 mm. longo; spicae 30
mm. longae 5 mm. crassae, pedunculo 10 mm. longo retrorso-hirsuto;
bracteae rotundo-subpeltatae nigrae, margine angusto brunneo;
stigmata 3 linearia sessilia.
Amazonas: Chachapoyas, Mathews in 1836 (type in herb. Kew.).
Piper reductipes Trelease, sp. nov.
Frutex glaber nigro-granulosus P. subsessili similis, internodiis
floriferis brevibus gracilibus; folia lanceolato-elliptica anguste
acuminata basi inaequilaterali acuta, margine ad basin petioli
breviusculi decurrente, 16 cm. longa 5.5-6.5 cm. lata e dimidio
inferiore pinnatinervia, nervis 6x2; spicae juvenes 25 mm. longae
2 mm. crassae arcuatae, pedunculo 5 mm. longo; bracteae parvae
subpeltatae; baccae teretes, stigmatibus sessilibus.
Loreto: La Victoria on the Amazon River, edge of forest, Llewelyn
Williams 2700 (herb. Field Mus. No. 617,253, type), 3143.
"Cordoncillo."
Piper reductipes var. amplum Trelease, var. nov.
Folia elliptica 22 cm. longa 8 cm. lata infra tertium supremum
pinnatinervia, nervis 7-8x2, petiolo 2+3 vel 3+7 mm. longo.
Loreto: La Victoria on the Amazon River, in forest, Llewelyn
William.s 2971 (herb. Field Mus. No. 608,157, type). "Cordoncillo."
Piper reductipes var. stenostachys Trelease, var. nov.
Petiolo ut pedunculo filiformi 10 mm. longo; spicis vix 35 mm.
longis et 2 mm. crassis.
Loreto: La Victoria on the Amazon River, in forest, Llewelyn
Williams 2578 (herb. Field Mus. No. 617,343, type). "Cordoncillo."
222 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper relaxatum Trelease, sp. nov.
Frutex nodosus 2-3-metralis glaber vel ad nervos folii subtus
obscure puberulus, internodiis floriferis brevibus gracilibus; folia
lanceolato-elliptica anguste acuminata basi acuta 15-16 cm. longa
6-7 cm. lata e dimidio inferiore pinnatinervia, nervis 5X2, in sicco
tenuia subopaca, petiolo 15 mm. longo exalato; spicae 90 mm. longae
3 mm. crassae subacutae, pedunculo 8-10 mm. longo; bracteae
parvae rotundo-subpeltatae ciliatae.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28385
(type in U. S. Nat. Herb.).
Piper resalutatum Trelease, sp. nov.
Arbor 5-6-metralis, internodiis floriferis brevibus subgracilibus
sordide villosis; folia ovata vel inaequilateraliter suboblonga acumin-
ata latere breviore basi subacuta 20-27 cm. longa 9-13 cm. lata e
dimidio inferiore submultiplinervia, nervis 6-7x2, subtus sparse
molliter pubescentia ad nervos breviter villosa, petiolo 10+3-20+5
mm. longo alato breviter villoso; spicae 140 mm. longae 5 mm.
crassae, pedunculo 10 mm. longo ferrugineo-pubescente; bracteae
rotundo- vel crescentico-subpeltatae ciliolatae; baccae rotundatae,
stylo gracili, stigmatibus 3 gracilibus.
Junin: San Nicolas, Pichis Trail, 110 meters, Killip & Smith
26024 (type in U. S. Nat. Herb.).
Piper resatum Trelease, sp. nov.
Frutex nodosus 3-4-metralis nervis folii subtus minute puberulis
exceptis glaber, internodiis floriferis gracilibus breviusculis ; folia
late elliptica vel subovata acuminata basi aequilaterali acuta 13-16
cm. longa 7-9 cm. lata e dimidio inferiore pinnatinervia, nervis
6x2, in sicco tenuia opaca, subtus paullo pallidiora, petiolo 10
mm. longo non vaginante; spicae juvenes 60 mm. longae 2 mm.
crassae acutatae, pedunculo 5 mm. longo; bracteae rotundo-sub-
peltatae ciliolatae.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28451
(type in U. S. Nat. Herb.).
Piper Restio Trelease, sp. nov.
Arbor gracilis 7-metralis glabra, internodiis floriferis breviusculis
gracilibus nigrescentibus plus minusve nigro-punctulatis; folia ro-
tundo-elliptica vel oblongo-elliptica obscure obtuso-acuminata basi
inaequaliter aperte cordata 16 cm. longa 9-12 cm. lata e 2 tertiis
inferioribus submultiplinervia, nervis circa 6+5, petiolo 4-7 cm.
FLORA OF PERU 223
longo ad laminam alato; spicae oppositifoliae 350 mm. longae et
3 mm. crassae vel ultra, pedunculo 15-20 mm. longo; bracteae
triangulari-subpeltatae ciliatae; baccae subquadrangulares cum
rhachi elongatae glabrae, stigmatibus 3 sessilibus.
Huanuco: Posuso, 600 meters, on gravel flat, Macbride 4689
(herb. Field Mus. No. 535,763, type).
Piper restrictum Trelease, sp. nov.
Frutex 3-4-metralis glaber plus minusve nodosus, internodiis
floriferis gracilibus subelongatis; folia elliptica sensim acuminata
basi subacuta, uno latere paullo breviore, 15-19 cm. longa 7-8 cm.
lata undique pinnatinervia, nervis circa 10x2, in sicco firmo-papy-
racea, petiolo 5+3 mm. longo exalato; spicae 80 mm. longae 3 mm.
crassae anguste cuspidatae, pedunculo 5 mm. longo; baccae prop-
ter pressum angulosae depresso-truncatae, stigmatibus 3 minutis
sessilibus.
Junin: Cahuapanas, 340 meters, Killip & Smith 26797 (type
in U. S. Nat. Herb.), 26737.
Piper reticulatum L. Sp. PI. 29. 1753. P. smilacifolium
HBK. Nov. Gen. & Sp. 1: 56. 1815. Enckea smilacifolia Kunth,
Linnaea 13: 605. 1839. Artanthe Ruiziana Miq. Lond. Journ. Bot.
4: 444. 1866. E. reticulata Miq. Syst. 365. 1844. P. tarapotianum
C. DC. in DC. Prodr. 16, pt. 1: 246. 1869. P. pangoense C. DC.
op. cit. 368.
A glabrous or very evanescently and locally puberulent shrub;
leaves ovate, acuminate, rounded or subtruncate at the base, 11-16
cm. long, 6-14 cm. wide, or even much larger, 7-nerved; petiole
5-10 mm. long; spikes slender but becoming 80 mm. long, 3 mm.
thick; peduncle 10 mm. long; bracts spatulate; berries globose,
papillate, with an apical disk bearing the broad sessile stigmas.
Loreto: Puerto Arturo, 200 meters, Williams 5096. Junin:
La Merced, Killip & Smith 23737. Puerto Yessup, Killip & Smith
26353. San Martin: Tarapoto, Williams 6037. As here accepted,
a scarcely divisible species, ranging through the West Indies and
from Brazil northwestward into Central America.
Piper reticulatum var. santa-rosanum Trelease, var. nov.
Differt foliis deltoideo-ovatis 9-nerviis basi retrorso-truncatis
21-29 cm. longis 18-25 cm. latis.
Loreto: Santa Rosa, 150-210 meters, in forest, Llewelyn Williams
4793 (herb. Field Mus. No. 629,672-673, type).
224 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper rhinostachyum Trelease, sp. nov.
Frutex 3-5-metralis nodosus nervis folii subtus sparse velutinis
exceptis glaber, internodiis floriferis brevibus crassiusculis; folia
subovato-elliptica breviter obtuso-acuminata basi truncato-sub-
cordata 22 cm. longa 12 cm. lata e 2 tertiis inferioribus pinnati-
nervia, nervis 8-10x2 intermediis adjectis, in sicco tenuia viridia
lucida, petiolo 2 cm. longo fragili-alato ; spicae fructiferae 40 mm.
longae 5 mm. crassae, ante anthesin 20 mm. longae 3 mm. crassae
cum rostro gracili 4 mm. longo, pedunculo crasso vix 10 mm. longo;
bracteae rotundo-subpeltatae; baccae rotundatae, stigmatibus
sessilibus.
Piper riojanum Trelease, sp. nov.
Frutex pagina superiore folii excepta transiente puberulus, inter-
nodiis floriferis subgracilibus elongatis; folia ovata vel subobovata
acute acuminata basi paullo inaequilateraliter cordata 18-22 cm.
longa 10-14 cm. lata e dimidio inferiore pinnatinervia, nervis circa
6x2, petiolo 15+2 mm. longo transiente alato; spicae 150 mm.
longae 4 mm. crassae, pedunculo 10 mm. longo; bracteae crescentico-
subpeltatae; baccae breviter cylindraceae vel cubicae puberulae,
stigmatibus 3 minutis sessilibus.
Amazonas: Rioja, Moyobamba, 2,700-3,300 meters, Llewelyn
Williams 7600 (herb. Field Mus. No. 629,721, type). "Cordoncillo."
Piper roqueanum Trelease, sp. nov.
Frutex glaber nigro-punctulatus, nodis plus minusve carnoso-
verrucosis, internodiis crassiusculis; folia elliptica vel subovata basi
obtusa vel uno latere breviora 22-30 cm. longa 15-19 cm. lata
undique pinnatinervia, nervis circa 16x2, petiolo 5+5 mm. longo;
spicae juveniles 25 mm. longae 5 mm. crassae, pedunculo vix 10
mm. longo.
San Martin: San Roque, in forest, 1,350-1,500 meters, Llewelyn
Williams 6961 (herb. Field Mus. No. 623,003, type).
Piper rubiginosum Trelease, sp. nov.
Frutex 3-4-metralis nodosus, internodiis floriferis brevibus crassis
ferrugineo-tomentulosis; folia lanceolata vel elliptico-subovata acumi-
nata basi inaequilaterali rotundata vel subcordulata 12-16 cm.
longa 4.5-8 cm. lata e dimidio inferiore pinnatinervia, nervis 5-6x2,
in statu adulto subrugosa, supra minute granuloso-asperata, subtus
molliter ferrugineo-hirsuta vel breviter villosa, petiolo 5+2 mm.
longo exalato velutine ferrugineo-tomentuloso; spicae 110 mm.
FLORA OF PERU 225
longae 3-4 mm. crassae, pedunculo 5 mm. longo tomentuloso;
bracteae rotundo-subpeltatae, margine crispato.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26292 (type
in U. S. Nat. Herb.).
Piper rubribaccum Trelease, sp. nov.
Frutex compactus 2-metralis nodosus, internodiis subgracilibus
elongatis glabris praesertim ad nodos plus minusve pallido-granu-
losis; folia lanceolato-oblonga vel elliptico-suboblonga acuminata
basi inaequaliter rotundata vel latere longiore cordulata 11-14
cm. longa 3.5-6 cm. lata e dimidio vel 2 tertiis inferioribus pinnati-
nervia, nervis circa 5x2, supra lucida, subtus ad nervos minute
puberula, petiolo 5-5+2 vel 10 mm. longo glabro exalato; spicae
oppositifoliae fere rectae 80 mm. longae 4 mm. crassae, pedunculo
5 mm. longo glabro; bracteae inconspicuae transverso-subpeltatae
ciliolatae, floribus sessilibus perfectis; stigmata 2-3 minuta sessilia;
baccae rotundo-subquadrangulares primo rubescentes.
Huanuco: Posuso, 600 meters, Macbride 4565 (herb. Field Mus.
No. 535,642, type).
Piper rugosibaccum Trelease, nom. nov. P. lanceolatum var.
mucronatum C. DC. Verb. Bot. Ver. Brandenb. 47: 106. 1905.
A more or less obscurely crisp-puberulent, granular shrub;
leaves lanceolate, acuminate, acute-based, 14-15 cm. long, 4-5
cm. wide, pinnately nerved throughout; petiole 10 mm. long; spikes
30 mm. long, 5 mm. thick, gently curved, cuspidate; peduncle 10
mm. long; berries globose, wrinkled; stigmas sessile.
Loreto: Iquitos, Ule 6246, type.
Piper rugosilimbum Trelease, sp. nov.
Frutex humilis nodosus, internodiis brevibus transiente sparse
pilosis; folia lanceolata acuta vel subobtusa basi acuta et aequila-
teralia vel uno latere (abnormaliter?) multo breviore 9-14 cm. longa
3.5-5 cm. lata et ultra e 2 tertiis inferioribus pinnatinervia, nervis
4-5x2 ut venis majoribus supra profunde impressis et subtus
prominentibus, obscure strigosa firma viridia, petiolo vix 10 mm.
longo primo alato; spicae in vivo luteae erectae 15 mm. longae
(et ultra?) 3 mm. crassae, pedunculo 7 mm. longo glabro; bracteae
crescenticae ; ovarium teres conice acutatum, stigmatibus 3 bre-
vibus latis.
Loreto: Mishuyacu near Iquitos, 100 meters, in forest, G. Klug
193 (herb. Field Mus. No. 630,030, type).
226 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper(?) sagittifer Trelease, sp. nov.
Suffrutex, internodiis breviusculis inferne crassis et fistulosis plus
minusve dense pilosis; folia oblonga vel triangulari-lanceolata vel
sagittata obtusa basi oblique cordata 20 cm. longa et 9 cm. lata
vel ultra ubique submultiplinervia, nervis 6-8x2, subtus ad nervos
subadpresso-pilosa, petiolo 4-6 cm. longo alato; inflorescentia
non visa.
Loreto : Pebas on the Amazon River, in forest, Llewelyn Williams
1605 (herb. Field Mus. No. 608,181, type), 1945.
Piper sandianum C. DC. Bot. Jahrb. 40: 247. 1908.
A glabrous shrub, or the nerves crisp-pubescent above; leaves
ovate, acuminate, obtuse at the base with one side distinctly shorter,
15-16 cm. long, 9-11 cm. wide, pinnately nerved from the lower
half, the nerves about 5x2; petiole 10+5 mm. long; spikes very
thick, 130 mm. long, 20 mm. thick; peduncle 5 mm. long; bracts
transversely subpeltate, ciliolate; berries round; stigmas sessile.
Neg. 11038.
Puno: Sandia, 2,400 meters, Weberbauer 727, type.
Piper sangorianum C. DC. Candollea 1: 140. 1923; 2: 187. 1925.
A glandular-punctulate shrub with gray-puberulent or velvety
twigs and nerves beneath; leaves elliptic or elongate-elliptic, acute
at both ends, 9-15 cm. long, 4-5.5 cm. wide, pinnately nerved
throughout; petiole 5-10 mm. long; spikes 15 mm. long, 3 mm.
thick; peduncle 5 mm. long; ovary round; stigmas sessile.
Puno: Sangari, Lechler 2527, type.
Piper sanguineispicum Trelease, sp. nov.
Arbor 5-6-metralis nervis folii subtus griseo-pubescentibus ex-
ceptis glabra, internodiis floriferis modice brevibus et gracilibus;
folia obovata basi oblique auriculata, sinu lateral! aperto, latere
longiore petiolum bene excedente, 30-32 cm. longa 13-16 cm. lata
e 2 tertiis inferioribus subpinnatinervia, nervis circa 5X2, nervis
supplementariis in auricula dispositis, petiolo 4-5 mm. longo alato;
spicae juvenes 50 mm. longae 3 mm. crassae acutatae in vivo obscure
rubrae, pedunculo gracili 3 cm. longo; bracteae rotundo-subpeltatae
glabrae.
Loreto: Iquitos, 100 meters, Killip & Smith 27135 (type in
U. S. Nat. Herb.), 27325.
FLORA OF PERU 227
Piper san-nicolasense Trelease, sp. nov.
Frutex 3-metralis nodosus, internodiis floriferis brevibus modice
gracilibus transiente rigide crispo-villosis; folia elliptico-lanceolata
acute acuminata basi subaequilaterali acuta 16-18 cm. longa 6-7
cm. lata ubique pinnatinervia, nervis circa 12x2, supra glabrescentia,
subtus ad nervos rigide pilosa, petiolo 10 mm. longo plus minusve
incurvo-piloso exalato; spicae 70 mm. longae 2-3 mm. crassae,
pedunculo gracili 10 mm. longo sparse piloso; bracteae rotundo-
subpeltatae ciliolatae; ovarium acutatum, stigmatibus 3 sessilibus.
Loreto: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
26009 (type in U. S. Nat. Herb.).
Piper san-ramonense Trelease, sp. nov.
Frutex gracilis P. dispanso similis, foliis non nigro-punctulatis,
nervis subtus grosse granulosis; spicae juvenes 7 mm. longae 1
mm. crassae.
Junin: Schunke Hacienda, San Ramon, 1,400-1,700 meters,
Killip & Smith 24585 (type in U. S. Nat. Herb.).
Piper san-roqueanum Trelease, sp. nov.
Frutex nodosus glaber, internodiis floriferis brevibus cito cras-
siusculis; folia subelliptica utrinque acuta, uno latere breviore,
10-15 cm. longa 3-4.5 cm. lata undique pinnatinervia, nervis 8x2,
in sicco firma pallida, petiolo 5+5 mm. longo alato; spicae fili-
formes 100 mm. longae, pedunculo 1 cm. longo; bracteae pallido-
zonatae rotundo-subpeltatae parvae ciliolatae; baccae truncatae
cum rhachi elongatae, stigmatibus 3 minutis sessilibus.
San Martin: San Roque, in forest, 1,350-1,500 meters, Llewelyn
Williams 7402 (herb. Field Mus. No. 623,525, type). Loreto:
Iquitos, Killip & Smith 27306.
Piper santa-ritanum Trelease, sp. nov.
Frutex nodosus, internodiis gracilibus brevissimis transiente
pallide sericeo-villosis; folia lanceolata sensim attenuata basi in-
aequilaterali cordulata 11-14 cm. longa 3-4 cm. lata e dimidio
inferiore pinnatinervia, nervis circa 6x2, valde bullulato-lacunosa,
supra glabra, subtus molliter velutina, petiolo vix 5 mm. longo
molliter piloso; spicae rectae, juvenes 40 mm. longae 3 mm. crassae,
pedunculo brevissimo velutino.
Cuzco: Santa Rita, Santa Ana Valley, 2,300 meters, Fortunato
L. Herrera 1551 (type).
228 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper santiaganum Trelease, sp. nov.
Frutex, petiolis et nervis folii subtus velutino-puberulis; folia
ovata subacuminata basi subcordulata 15 cm. longa 8-9 cm. lata
e dimidio inferiore vel paullo altius pinnatinervia, nervis 5x2,
petiolo 2 cm. longo infra medium vaginante; spicae juveniles 60
mm. longae 3 mm. crassae mucronatae, pedunculo 10 mm. longo.
Loreto: Mouth of Rio Santiago, 160 meters, G. Tessmann 3971
(type in herb. Berol.). Also Seringal Sao Francisco, Rio Acre,
Brazil, Vie 9283 (herb. Kew.).
Piper sapotoanum Trelease, sp. nov.
Frutex, internodiis floriferis mediocribus aliquantum transiente
crispo-pubescentibus vel laxe subtomentosis; folia elliptica vel sub-
rhombea acuminata basi inaequilaterali cordulata 16-19 cm. longa
7-8 cm. lata undique subpinnatinervia, nervis circa 8x2, aliis brevi-
bus basilaribus in auriculis dispositis, subtus sparse pilosa, petiolo
10-15 mm. longo subtomentoso vix alato; spicae 10 mm. longae
5 mm. crassae deflexae breviter mucronatae, pedunculo 5 mm.
longo tomentoso; bracteae rotundo-subpeltatae; stigmata 3 stylum
distinctum terminantia.
Loreto: Sapoto Yacu, Santa Rosa, 160-210 meters, Lleivelyn
Williams 4816 (herb. Field Mus. No. 627,172, type).
Piper sapoto-yacuense Trelease, sp. nov.
Frutex humilis, internodiis floriferis gracilibus breviusculis
crispo-pubescentibus; folia lanceolato-elliptica acuminata basi acuta
9-14 cm. longa 4-6 cm. lata undique pinnatinervia, nervis 8-9x2,
plus minusve conspicue nigro-granulosa, subtus praesertim ad nervos
puberula, petiolo 10 mm. longo gracili exalato nonnihil griseo-
pubescente; spicae fructiferae 35 mm. longae 5 mm. crassae acutatae,
pedunculo crasso 5 mm. longo griseo-pubescente; bracteae triangu-
lari-subpeltatae; baccae subglobosae plus minusve rugosae, stig-
matibus 3 sessilibus.
Loreto: Sapoto Yacu, Santa Rosa, in forest, 155-210 meters,
Llewelyn Williams 4923 (herb. Field Mus. No. 624,945, type).
Puerto Arturo, in forest, 155-210 meters, Williams 5027. Paraiso,
145 meters, Williams 3326 (fruit mucronate).
Piper satisfactum Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis brevibus crassiusculis laxe
rigido-villosis; folia lanceolato-elliptica acuminata basi subaequila-
FLORA OF PERU 229
terali acuta 14 cm. longa 5-6 cm. lata undique pinnatinervia, nervis
majoribus circa 4X2, utrinque subsparse rigido-villosa, petiole 5
mm. longo vel in foliis longioribus 10 mm. longo atque late alato,
piloso; spicae pendulae 40 mm. longae 4 mm. crassae rubescentes,
pedunculo filiformi 7 mm. longo glabrato; bracteae triangulari-
subpeltatae ciliatae; baccae rotundatae subtruncatae, stigmatibus
3 brevibus sessilibus.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26245 (type
in U. S. Nat. Herb.).
Piper scabiosifolium Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis modice gracilibus et longia
incurvo-pilosis vel molliter adpresso-pilosis; folia elliptico-lanceolata
vel subovata anguste acuminata basi inaequilateralia, uno vel
ambobus lateribus rotundatis, 12-14 cm. longa 4-5.5 cm. lata e 2
tertiis inferioribus pinnatinervia, nervis circa 7x2, supra scabra vel
hinc inde glabra, subtus ad nervos molliter hirsuta, petiolo 2+3
mm. longo exalato molliter piloso; spicae 25 mm. longae 3 mm.
crassae acutatae, pedunculo 10 mm. longo glabro; bracteae rotundo-
subpeltatae ciliolatae.
Loreto: Puerto Arturo, 135 meters, Killip & Smith 27730 (type
in U. S. Nat. Herb.).
Piper scabricaule Trelease, sp. nov.
Frutex 3-metralis, internodiis superioribus gracilibus subelongatis
minute sed manifeste scabridis; folia rotundo-elliptica vel sub-
obovato-elliptica subabrupte acuto-acuminata basi rotundata, uno
latere breviore, 19-21 cm. longa 10-11 cm. lata e dimidio inferiore
pinnatinervia, nervis 6x2, supra subgrosse albide granuloso-scabrida,
subtus ad nervos indistincte adpresso-pubescentia, petiolo 10+5
mm. longo exalato; inflorescentia non visa.
Junin: Near Perene Bridge, Rio Paucartambo, 700 meters,
Killip & Smith 25299 (type in U. S. Nat. Herb.).
Piper scabridulicaule Trelease, sp. nov.
Frutex nodosus 3-metralis, internodiis superioribus gracilibus
crassiusculis ochroleucis dense breviter scabridis; folia lanceolato-
elliptica anguste acuminata basi inaequilateralia, uno vel ambobus
lateribus vulgo obtusis, e dimidio inferiore pinnatinervia, nervis
5-6x2, supra granuloso-scabra, subtus ad nervos dense scabra;
spicae flexuosae vel arcuatae 110 mm. longae 2 mm. crassae, pedun-
culo 5 mm. longo scabrido; bracteae rotundo-subpeltatae ciliolatae.
230 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Loreto: Mouth of Rio Santiago, Ynes Mexia 6100 (type in
herb. Univ. Calif.).
Piper scalariforme Trelease, sp. nov.
Frutex in sicco fuscescens, internodiis floriferis brevibus modice
gracilibus puberulis vel obscure transiente sericeis; folia lanceolata
vel lanceolato-oblonga sensim subacuminata basi inaequilaterali
rotundata vel cordulata 16-20 cm. longa 5.5-7 cm. lata e dimidio
inferiore pinnatinervia, nervis 5x2 venis scalariformibus transversis
connexis, in sicco tenuia lucida, subtus pallidiora ad nervos ad-
presso-puberula, petiolo 10 mm. vel 5-7+3 mm. longo subpuberulo
alato; spicae 85 mm. longae 3 mm. crassae, pedunculo 5 mm. longo
glabrato; bracteae parvae rotundo-subpeltatae; baccae teretes, stig-
matibus sessilibus.
Loreto: Caballo-cocha on the Amazon River, in forest, Llewelyn
Williams 2057 (herb. Field Mus. No. 613,146, type), 2056. Mishu-
yacu, 100 meters, Killip & Smith 29882.
Piper scapispicum Trelease, sp. nov.
Frutex fistulosus(?) metralis, internodiis floriferis gracilibus
elongatis laxe intertexto-villosis; folia ovata sensim acuminata basi
subtruncata 22-24 cm. longa 11-13 cm. lata e dimidio inferiore
submultiplinervia, nervis 5-6x2, in sicco tenuia brunnescentia,
subtus ferrugineo-tomentulosa ad nervos crispo-pilosa, petiolo 15
mm. longo crispo-piloso primo alato; spicae juvenes 25 mm. longae
3 mm. crassae, pedunculo fere aequilongo tomentuloso; bracteae
subclavatae.
Loreto: Mishuyacu near Iquitos, 100 meters, in forest, G. Klug
1420 (herb. Field Mus. No. 627,684, type).
Piper Schunkeanum Trelease, sp. nov.
Frutex, internodiis floriferis mediocribus griseo-villoso-hirsutis;
folia subelliptica longiuscule anguste acuminata basi paullo in-
aequilaterali obtusa 15-16 cm. longa 6.5-7.5 cm. lata e dimidio
inferiore pinnatinervia, nervis circa 6x2, supra glabra sed granuloso-
asperata, subtus praesertim ad nervos molliter adscendenti-pubes-
centia, petiolo vix 10 mm. longo molliter brunneo-pubescente basi
alato; spicae 110 mm. longae 3 mm. crassae, pedunculo vix 10
mm. longo hirtello; bracteae parvae rotundo-subpeltatae ciliolatae.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 275
(herb. Field Mus. No. 571,331, type).
1
FLORA OF PERU 231
Piper sciaphilum C. DC. Bot. Jahrb. 40: 242. 1908.
A shrub, glabrous except for the obscurely puberulent nerves
beneath; leaves ovate, acuminate, rounded at the base with one
side slightly longer and abruptly narrowed to the petiole, 15-28
cm. long, 7-17 cm. wide, submultiple-nerved from the lower half
or less, the nerves 4-5x2; petiole 20+2 mm. long, scarcely winged;
spikes 130 mm. long, 3 mm. thick; peduncle 15 mm. long; bracts
transversely subpeltate, ciliate; berries round, with a very short,
tapering style; stigmas spreading. Neg. 11040.
Huanuco: Mountains west of Monzon, 1,800 meters, in forest,
Weberbauer 3557, type.
Piper secundum R. & P. Fl. 1: 36. pi. 62. 1798. P. hetero-
mallum Pers. Syn. PI. 1: 33. 1805 (fide Kunth). Steffensia secunda
Kunth, Linnaea 13 : 626. 1839. Artanthe secunda Miq. Syst. 516. 1844.
A glabrous, slightly papillate shrub; leaves lance-oblong or
elliptic-oblong, acuminate, acute-based, 12-16 cm. long, 6-7 cm.
wide, pinnately nerved from below the upper third, the nerves 5x2;
petiole barely 10 mm. long; spikes secund, straight, 100 mm. long,
3 mm. thick; peduncle 10 mm. long. Neg. 11041.
Huanuco: Pillao, Muna, Cochero, and Chinchao, Ruiz &
Pavon, type.
Piper Sellertianum Trelease, nom. nov. Enckea Lessertiana
Miq. Syst. 349. 1844; 111. Piper, pi. 63, nee P. Lessertianum C. DC.
Journ. Bot. 4: 164. 1866, nee P. Lessertianum C. DC. in DC. Prodr.
16, pt. 1: 258. 1869, nee Artanthe Lessertiana Miq. Syst. 405, nee
Carpunya Lessertiana Presl, Epim. 229, nee Chavica Lessertiana
Miq. Syst. 270, nee Ottonia Lessertiana Miq. Syst. 516; 111. Piper.
pi. 92, nee Pothomorphe Lessertiana Miq. Syst. 210; 111. Piper, pi. 26.
A glabrous shrub, nodose; flowering internodes slender and
short; leaves acuminate, dimorphic, narrowly lanceolate, acute at
the base and 13-15 cm. long, 3-4 cm. wide, or broadly elliptic or
elliptic-subovate, subacute to round at the base, and 18-19 cm.
long, 8-10 cm. thick, 5- or sometimes 7-nerved with connecting
cross veins, drying thin, green and sometimes with whitened nerves;
petiole 10-15 mm. long, not winged; spikes slender, 50-70 mm. long;
peduncle 10 mm. long; bracts subcucullate; ovary oblong-conic,
becoming ovoid, scarcely constricted; stigmas 3, sessile.
Loreto: Type from Amazon River (perhaps in Brazil), Poeppig.
Parana Pura, Yurimaguas, in forest, Williams, 4605, 4593. Lower
232 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Rio Nanay, in forest, Williams 484- Yurimaguas, Killip & Smith
27616. Puerto Arturo, Killip & Smith 27872. Santa Rosa, Killip
& Smith 28834, 28880.
Piper Sellertianum var. tenuiforme Trelease, var. nov. P.
tenue Auct.
Frutex nodosus glaber P. Sellertiano similis; folia lanceolata
acuminata basi acuta 13-15 cm. longa 4-5 cm. lata 5-nervia, petiolo
10 mm. longo versus basin vaginante; spicae graciles 70 mm. longae,
pedunculo 10 mm. longo; bracteae subcucullatae; baccae ovoideae
non constrictae, stigmatibus 3 sessilibus.
Huanuco: Cochero, Poeppig (type).
Piper semicordulatum Trelease, sp. nov.
Frutex nerviis folii subtus aliquantum adpresso-pubescentibus
exceptis glaber, internodiis gracilibus modice brevibus; folia elliptica
vel oblanceolato-obovata anguste acuto-acuminata basi inaequila-
teralia et obtusa, latere breviore breviter truncato-cordulato, 15 cm.
longa 5.5-6.5 cm. lata e tertio vel dimidio inferiore pinnatinervia,
nervis circa 5x2 ut venis majoribus supra impressis subtus promin-
entibus, petiolo 8-10+2 mm. longo exalato; spicae juvenes 50 mm.
longae 2 mm. crassae, pedunculo 5 mm. longo; bracteae parvae
rotundo-subpeltatae.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 238
(herb. Field Mus. No. 627,313, type).
Piper semicrudum Trelease, sp. nov.
Frutex nodosus glaber vix 2-metralis, internodiis floriferis brevi-
bus gracillimis; folia lanceolata acuminata basi aequilaterali cordato-
truncata vel basi inaequilateralia uno latere in petiolum acute decur-
rente, 10-14 cm. longa 3-5 cm. lata e dimidio inferiore pinnatinervia,
nervis circa 5X2, in sicco tenuia subtus pallidiora sparsiuscule nigro-
glandulosa, petiolo 10+2 vel 15 mm. longo exalato; spicae fructiferae
20 mm. longae 8 mm. crassae breviter mucronatae, pedunculo patente
filiformi 5 mm. longo; bracteae crescenticae; baccae globosae breviter
mucronatae, stigmatibus in mucrone insertis.
Loreto : Balsapuerto, 350-550 meters, Killip & Smith 28524
(type in U. S. Nat. Herb.).
Piper seminitidulum Trelease, sp. nov.
Frutex compactus vel arbuscula nodosa, internodiis floriferis
modice brevibus et gracilibus glabris; folia lanceolato-oblonga sub-
acuta basi inaequilaterali acuta 11-13 cm. longa 3.5-5.5 cm. lata e 2
FLORA OF PERU 233
tertiis inferioribus pinnatinervia, nervis circa 4x2, supra lucida, sub-
tus ad nervos minute puberula, petiolo 7+3 mm. longo glabro exalato;
spicae oppositifoliae 100 mm. longae 3 mm. crassae, pedunculo
brevissimo 3 mm. longo glabro; bracteae parvae rotundo- vel trans-
verso-subpeltatae ciliolatae; flores sessiles perfectae; baccae quad-
rangulares saepe cum rhachi elongatae, stigmatibus 3 brevibus
sessilibus.
Junin: La Merced, 600 meters, Macbride 5289 (herb. Field Mus.
No. 536,314, type). Chanchamayo Valley, Schunke 528, 1594;
Killip & Smith 24093. Colonia Perene", Killip & Smith 24964.
Piper sericeonervosum Trelease, sp. nov.
Arbuscula aliquantum nodosa nerviis folii subtus sericeis exceptis
glabra, internodiis floriferis brevibus gracilibus; folia lanceolato-
elliptica sensim acuminata basi rotundata, uno latere multo breviore,
11-16 cm. longa 4.5-6 cm. lata ubique pinnatinervia, petiolo 3-5+
10-12 mm. longo alato; spicae 90 mm. longae 3 mm. crassae, pedun-
culo gracili 10 mm. longo; bracteae rotundo-subpeltatae ciliatae;
baccae rotundo-truncatae, stigmatibus 3 minutis sessilibus.
Loreto: La Victoria on the Amazon River, Llewelyn Williams
2928 (herb. Field Mus. No. 608,548, type).
Piper serotinum Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis breviusculis gracilibus
submolliter incurvo-griseo-pubescentibus; folia elliptica acuminata
basi obtusa, uno latere paullo breviore, 18 cm. longa 8-9 cm. lata e
dimidio inferiore pinnatinervia, nervis 6x2, in statu adulto subru-
gosa, supra grosse granuloso-scabra, subtus ad nervos breviter pilosa,
petiolo 10-15+3 mm. longo subsericeo.
Junin: Perene Bridge, Rio Paucartambo, 700 meters, Killip &
Smith 25344 (type in U. S. Nat. Herb.).
Piper serpavillanum Trelease, sp. nov.
Frutex glaber; folia lanceolata sensim longiacuta vel subacu-
minata basi acuta, uno latere paullo breviora, 18 cm. longa 4-5 cm.
lata undique pinnatinervia, nervis circa 10-12x2, petiolo 10+5 mm.
longo alato; spicae 80 mm. longae 3 mm. crassae, pedunculo 5-10
mm. longo; baccae truncato-subcubicae, stigmatibus 3 parvis
sessilibus.
Type from Serpa Villa, Para, Brazil, Martius (type in
herb. Munich). Loreto: Caballo-cocha on the Amazon River,
Williams 2120.
234 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper setigerum Trelease, sp. nov.
Frutex gracilis erectus metralis nodosus, internodiis floriferis
brevibus gracilibus barbatis; folia lanceolato-elliptica acuminata basi
fere aequilaterali obtusa 11-14 cm. longa 3.5-5 cm. lata ubique pin-
natinervia, nervis circa 6x2, in sicco tenuia viridia utrinque sed
praesertim subtus ad nervos sparse barbata, petiolo 3-10 mm. longo
late alato; spicae pendulae 25 mm. longae 2 mm. crassae, pedunculo
filiformi 8 mm. longo barbato; bracteae rotundo-subpeltatae ciliatae;
ovarium globosum, stigmatibus sessilibus.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26510
(type in U. S. Nat. Herb.).
Piper setulosum Trelease, sp. nov.
Frutex 3-metralis, internodiis superioribus gracilibus cito elonga-
tis incurvo-hispidis vel hirsutis; folia elliptica vel sublanceolata vel
subovata aliquantum acuminata basi rotundata, uno latere saepe
breviore, 11-16 cm. longa 6-7 cm. lata e dimidio inferiore pinnati-
nervia, nervis 6x2, supra subgrosse granuloso-scabra, subtus ad
nervos incurvo-hirtella, petiolo 5+5 mm. longo exalato hispido;
infiorescentia non visa.
Loreto: Yurimaguas, 135 meters, Killip & Smith 27568 (type in
U. S. Nat. Herb.).
Piper sidipilum Trelease, sp. nov.
Arbuscula 3-5-metralis, internodiis floriferis longis gracilibus sub-
persistente inconspicue stellato-pubescentibus; folia ovata falcato-
acuminata basi rotundata acute in petiolum contracta 20-22 cm.
longa 9-13 cm. lata e dimidio inferiore submultiplinervia, nervis
5x2, in sicco tenuia viridia, subtus praesertim ad nervos inconspicue
stellato-pubescentia, petiolo 2-4 cm. longo vaginante puberulo;
spicae 110 mm. longae 3 mm. crassae, pedunculo 10 mm. longo
puberulo; bracteae triangulari-subpeltatae; baccae teretes, stylo
gracili, stigmatibus 3 filiformibus.
Junin: Rio Perene, 600 meters, Killip & Smith 25160 (type in
U. S. Nat. Herb.). Colonia Perene", Killip & Smith 24965.
Piper silvioberrans Trelease, sp. nov. P. silvivagum var. peru-
vianum C. DC. Notizbl. Bot. Gart. Berlin 6: 489. 1917.
Frutex reclinatus nodosus, internodiis floriferis brevibus gracilibus
retrorse canescentibus in statu adulto velutinis; folia elliptica pro-
tracto-acuminata basi angusta inaequilaterali rotundata vel sub-
cordulata 10-11 cm. longa 4-5.5 cm. lata e dimidio inferiore pinnati-
FLORA OF PERU 235
nervia, nervis circa 5x2, supra minute granulosa et serius lepidota,
subtus ad nervos pallido-strigosa, petiolo vix 5 mm. longo canescente;
spicae 120 mm. longae 3 mm. crassae acutatae; bracteae rotundo-
subpeltatae ciliolatae; baccae teretes subpuberulae, stigmatibus 3
parvis sessilibus.
Loreto: Yurimaguas to Balsapuerto, 135-150 meters, Killip &
Smith 28119 (type in U. S. Nat. Herb.). Brazil: Seringal Auristella,
Rio Acre, Ule 9290.
Piper sinuclausum Trelease, sp. nov.
Subarborescens nodosus, internodiis floriferis brevibus crassius-
culis primo dense hamate hirsuto-villosis, serius glabrescentibus et
pallido-granulosis; folia lanceolata longiattenuata basi inaequilate-
rali subprofunde (7 mm.) auriculato-cordulata, auricula breviore
longiorem late imbricante, 18 cm. longa 6-7 cm. lata e dimidio in-
feriore pinnatinervia, nervis circa 6x2, in statu adulto rugosa, supra
scabra et pilosa, subtus ad nervos hirsuta, petiolo brevi vix 10 mm.
longo lamina occulto incurvo-hirsuto exalato; spicae oppositi-
foliae fere rectae 70 mm. longae 5 mm. crassae, pedunculo brevi 5
mm. longo hamato-hirsuto; bracteae subtriangulari-subpeltatae, mar-
gine angustissimo ciliato, floribus sessilibus perfectis; stigmata 3
brevia sessilia; baccae parvae oblongae subteretes.
Huanuco: Maria del Valle, 2,100 meters, Macbride 3564 (herb.
Field Mus. No. 534,640, type). Cuzco: Machupicchu, 2,400 meters,
F. L. Herrera 2007. "Matico."
Piper soledadense Trelease, sp. nov.
Frutex 2-4-metralis, internodiis floriferis subgracilibus elongatis
persistente crispe fulvo-villosis; folia elliptico-lanceolata vel ovato-
lanceolata acute acuminata basi subobliqua obtusa vel acuta 12-19
cm. longa 5.5-7.5 cm. lata undique pinnatinervia, nervis circa
12X 2, praesertim subtus sparse molliter stellato-pubescentia, subtus
ad nervos crispo-villosa, petiolo vix 10 mm. longo rigide crispo-
villoso exalato; spicae 20-25 mm. longae 3 mm. crassae, pedunculo
7-10 mm. longo rigido-villoso; bracteae triangulari-subpeltatae
ciliatae; stigmata 3 in stylo vel mucrone brevi inserta.
Loreto: Soledad, Rio Itaya, 110 meters, Killip & Smith 29769
(type in U. S. Nat. Herb.), 29598, 29711. Iquitos, Killip & Smith
27101, 27134- Between Yurimaguas and Balsapuerto, Killip &
Smith 281 07.
236 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper soledadense var. august um Trelease, var. nov.
Folia lanceolato-oblonga 14-16 cm. longa 3.5-4.5 cm. lata;
spicae fructiferae curvae 15 mm. longae 5 mm. crassae; baccae glo-
bosae magnae dense insertae, stigmatibus sessilibus.
Loreto: San Antonio, Rio Itaya, 110 meters, Killip & Smith
29350 (type in U. S. Nat. Herb.).
Piper soricicaudatum Trelease, sp. nov.
Frutex glaber 3-4-metralis, internodiis floriferis brevibus modice
gracilibus; folia lanceolato-oblonga anguste longiacuminata basi
subaequilaterali acuta 17 cm. longa 5 cm. lata e dimidio inferiore
pinnatinervia, nervis circa 6x2, in sicco tenuia, petiolo 5 mm.
longo exalato; spicae 100-140 mm. longae 3 mm. crassae sinuatae,
pedunculo 10 mm. longo; bracteae zonulatae parvae rotundo-sub-
peltatae ciliatae; ovarium teres, stigmatibus 3 minutis sessilibus.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26661
(type in U. S. Nat. Herb.).
Piper sororium (Miq.) C. DC. in DC. Prodr. 16, pt. 1: 295.
1869. Artanthe sororia Miq. Lond. Journ. Bot. 4: 443. 1866. P. un-
deninervium C. DC. loc. cit.
A shrub, glabrous or transiently puberulent near the bases of the
leaves, these ovate, acuminate, truncately or even deeply cordate,
11-17 cm. long, 7-15 cm. wide, nerved; petiole 3-6 cm. long, winged;
spikes slender, curved, equaling the leaves; peduncle 5 mm. long;
bracts triangular-subpeltate; berries ovoid; stigmas sessile.
Huanuco: Chinchao, Ruiz & Pavon, type. Loreto (?): Mainas,
Poeppig (herb. Leningrad). San Martin: Rumizapa, 400-900
meters, Williams 6827.
Piper Spruceanum C. DC. Journ. Bot. 4: 216. 1866.
A shrub, villous with long, pale or yellow hairs throughout;
leaves lanceolate or suboblanceolate, subacuminate, acute-based,
18 cm. long, 5 cm. wide, pinnately nerved throughout; petiole 1 cm.
long, margined by the decurrent blade; spikes 35 mm. long, 3 mm.
thick; peduncle 15 mm. long; bracts inflexed-subpeltate ; berries
round, granular; stigmas 3, broad, sessile.
San Martin: Tarapoto, Spruce 4072, type.
Piper stenostachys C. DC. Ann. Cons. Jard. Geneve 2:
254. 1898.
A glabrous shrub; twigs dark-glandular; leaves lance-oblong,
acute at both ends, 14 cm. long, 5 cm. wide, pinnately nerved
FLORA OF PERU 237
nearly throughout, the nerves about 12x2; petiole 5 mm. long;
spikes curved, filiform, 50 mm. long; bracts triangular-subpeltate,
ciliate; ovary glabrous; stigmas on a short style.
Type from Peru, Gay in herb. Paris.
Piper Stevensi Trelease, sp. nov.
Frutex, ramulis plus minusve caduce subvillosis; folia lanceolata
acuminata basi inaequilaterali cordulata 15-20 cm. longa 5.5 cm.
lata e dimidio inferiore pinnatinervia, nervis 6-8x2, supra paullo
scabrida, subtus ad nervos laxe pilosa, in statu adulto rugosa, petiolo
vix 10 mm. longo piloso; spicae valde curvatae 100 mm. longae 2-3
mm. crassae, pedunculo gracili 10-15 mm. longo hirtello; bracteae
triangulari-subpeltatae ciliolatae pallidae; baccae rotundatae, stig-
matibus 3 filiformibus sessilibus.
Junin: Huacapistana, F. L. Stevens 85, type.
Piper stomachicum C. DC. Bot. Jahrb. 40: 250. 1908.
A medicinal shrub, yellowish-crisp-pilose; leaves ovate, subacute,
cordulate, 8-9 cm. long, 4-5 cm. wide, pinnately nerved from below
the middle, the nerves 3x2, glabrous above between the nerves;
petiole 5-10 mm. long; spikes 3-5 mm. thick, 30-40 mm. long; pe-
duncle 10-15 mm. long; berries round ; stigmas 3, sessile. Neg. 11052.
Ancash: Caraz, in thicket, 3,200 meters, Weberbauer 3028,
type. Amazonas: Chachapoyas, Mathews 3223.
Piper striatinervosum Trelease, sp. nov.
Arbuscula 4-5-metralis petiolis et nervis folii subtus obscure
adpresse striato-pubescentibus exceptis glabra, internodiis superi-
oribus comparate gracilibus subelongatis; folia elliptica obtusa basi
inaequilateralia, sinu laterali fere clauso, latere majore auriculato
et petiolo longiore, 32 cm. longa 19 cm. lata e dimidio inferiore vel
altius subpinnatinervia, nervis circa 6+7, in sicco tenuia lucida,
petiolo 3-5 mm. longo alato; inflorescentia non visa.
Loreto: Puerto Arturo, 135 meters, Killip & Smith 27910 (type
in U. S. Nat. Herb.).
Piper strigillosicaule Trelease, sp. nov.
Frutex debilis 3-4-metralis, internodiis floriferis brevibus graci-
libus pallidis sparse retrorso-strigillosis ; folia elliptica ad lanceolata
vel oblanceolata acuminata basi inaequilateralia et anguste rotundata
vel cordulata 8-10 cm. longa 2-4 cm. lata e dimidio inferiore
pinnatineryia, nervis 4-5x2, subtus ad nervos strigillosa, petiolo
238 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
4+1 mm. longo strigoso; spicae 100 mm. longae 2 mm. crassae,
pedunculo 8 mm. longo glabrato; bracteae parvae griseae rotundo-
subpeltatae.
Loreto: Yurimaguas, 135 meters, Killip & Smith 27615 (type in
U. S. Nat. Herb.).
Piper strigillosum Trelease, sp. nov.
Frutex 3-metralis, internodiis superioribus brevibus gracilibus
evanescente adpresso-pubescentibus serius albide granuloso-asperatis;
folia anguste lanceolata vel subelliptica acute acuminata basi in-
aequilaterali acuta 12-15 cm. longa 3-5.5 cm. lata e dimidio inferiore
pinnatinervia, nervis minute pallido-strigillosis 5x2, in sicco tenuia,
petiolo 5+3 mm. longo ut costa griseo-strigilloso ; spicae juvenes
15 mm. longae 2 mm. crassae.
Loreto: Puerto Arturo, 135 meters, Killip & Smith 27813 (type
in U. S. Nat. Herb.).
Piper strigosum Trelease, sp. nov.
Frutex 3-4-metralis, internodiis longis crassiusculis adultis rubro-
pilosis; folia oblongo-obovata breviter acuminata basi inaequila-
teralia, uno latere semicirculari altero in auriculam subtruncatam
petiolo aequilongam prolongato, 33 cm. longa 17 cm. lata e dimidio
inferiore vel paullo altius subpinnatinervia, nervis circa 7x2, in
sicco tenuia, utrinque subtus ad nervos densius longipilosa, petiolo
6 cm. longo dense rubro-piloso; inflorescentia non visa.
Loreto: Balsapuerto, 350-550 meters, Killip & Smith 28538
(type in U. S. Nat. Herb.).
Piper Stuebelii Trelease, sp. nov.
Frutex, internodiis primo molliter crispo-pilosis vel subtomentosis
serius aliquanto pallido-granulosis ; folia elliptico-lanceolata acu-
minata basi inaequilaterali cordulata 16-20 cm. longa 6-7 cm. lata
e dimidio inferiore pinnatinervia, nervis circa 6+7, granuloso-scabra,
subtus ad nervos adpresso-pubescentia, petiolo 5 mm. longo sub-
crispe pubescente; spicae valde curvae 120 mm. longae 3 mm.
crassae, pedunculo 5-10 mm. longo hispido; bracteae inconspicuae
rotundo-subpeltatae; baccae teretes.
Loreto(?): Rio Huallaga, Stuebel 68 (type in herb. Berol.).
San Martin: San Roque, Williams 7056. Tarapoto, Williams 5622.
"Cordoncillo."
FLORA OF PERU 239
Piper stupposum A. Dietr. Sp. PI. 1: 645. 1831. P. rugosum
Opiz in Presl, Rel. Haenk. 1: 152. pi. 27. 1830, non P. rugosum Lam.
1791. Artanthe stupposa Miq. Syst. 430. 1844.
A densely yellow-hairy shrub ; leaves lance-elliptic, subacuminate,
subobtuse at the base, 7-10 cm. long, 3.5 cm. wide, pinnately nerved
nearly throughout, bullulate-lacunose, not scabrous; petiole 5 mm.
long; spikes gently curved, 40 mm. long, 3 mm. thick, blunt; peduncle
10 mm. long; berries round; stigmas 3, sessile.
Type from mountains of Peru, Haenke in 1790.
Piper subandinum C. DC. in DC. Prodr. 16, pt. 1: 275. 1869.
A shrub, the twigs and petioles hirtellous with intermingled
long soft hairs; leaves elliptic, acuminate, acute at the base with one
side slightly shorter, 22 cm. long, 9 cm. wide, pinnately nerved from
the lower half, the nerves 4x2, of which the lower 6 are congested
toward the base, appressed-pubescent beneath on the nerves; petiole
4-f4 mm. long; spikes 65 mm. long, 4 mm. thick; peduncle 5 mm.
long, at first hirtellous; bracts round-subpeltate, small, lead-colored,
ciliolate.
Huanuco: Cochero, Poeppig in 1830, type.
Piper subcordulatum Trelease, sp. nov.
Frutex glaber 2-metralis, internodiis floriferis breviusculis graci-
libus; folia ovata acuminata basi rotundata vel cordulata vel cor-
data 15-22 cm. longa 9-15 cm. lata e 2 tertiis inferioribus submulti-
plinervia, nervis circa 7x2 intermediis interpositis, petiolo 15-30
mm. longo; spicae 90 mm. longae 3 mm. crassae, pedunculo vix
10 mm. longo; bracteae crescenticae; stigmata fere sessilia.
Loreto: Soledad, Rio Itaya, 110 meters, Killip & Smith 29794
(type in U. S. Nat. Herb.).
Piper subflavicans Trelease, sp. nov.
Frutex in sicco flavescens, ramulis transiente paullo crispo-
pubescentibus conspicue pallido-granulosis; folia lanceolata sub-
acuminata basi cordata 7-12 cm. longa 3-5 cm. lata e dimidio infe-
riore pinnatinervia, nervis circa 5x2, supra granuloso-asperata,
subtus sparse adpresso-pubescentia, petiolo vix 5 mm. longo alato;
spicae leviter curvae 75 mm. longae 4 mm. crassae mucronatae,
pedunculo 20 mm. longo; bracteae subtriangulari-subpeltatae cili-
atae; baccae teretes; stigmata 3 sessilia.
Type collected in Peru by Haenke (herb. Munich).
240 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper subflavispicum C. DC. Bot. Jahrb. 40: 244. 1908.
A shrub, the glabrescent twigs somewhat pilose at the nodes and
pale-granular; leaves elliptic or lanceolate, acuminate, subacute at
the base, 11-12 cm. long, 4-5 cm. wide, pinnately nerved from the
lower half, the nerves about 5X2, deeply impressed above, salient
and obscurely appressed-pubescent beneath; petiole 5 mm. long,
granular, crisp-ciliate; spikes 30 mm. long, 3 mm. thick, straight,
pointed; peduncle barely 10 mm. long; bracts subpeltate, with a
narrow yellow margin. Neg. 11054.
Junin: Mountains west of Huacapistana, in thicket, 2,000-
2,400 meters, Weberbauer 2156, type; Stevens 82.
Piper subpatens Trelease, sp. nov.
Frutex 1.5 m. altus conspicue nodosus, internodiis floriferis
brevibus crassis dense hirsutis et persistente hispidis; folia lanceolato-
elliptica acuminata basi angustata typice subacuta 12-14 cm. longa
4-5 cm. lata e dimidio inferiore pinnatinervia, nervis circa 4x2,
scabra, subtus praesertim ad nervos hirsuta; spicae oppositifoliae
longae patentes 110-120 mm. longae 2-3 mm. crassae, pedunculo
minus quam 5 mm. longo breviter hirsute; bracteae rotundo-
subpeltatae ciliolatae, floribus sessilibus perfectis; stigmata 3 minuta
sessilia; baccae breviter cylindraceae.
Huanuco: Posuso, 600 meters, Macbride 4633 (herb. Field Mus.
No. 535,708, type).
Piper subscutatum (Miq.) C. DC. in DC. Prodr. 16, pt. 1:
321. 1869. Artanthe scutata Miq. Lond. Journ. Bot. 4: 444. 1866,
non Piper scutatum Auct.
A glabrous shrub; leaves ovate, slender-acuminate, peltate above
the round or subcordate base, 26 cm. long, 20 cm. wide, submultiple-
nerved from the lower half, the nerves 6-8x2; petiole 7-9 cm. long,
becoming fleshy-warty, winged; spikes 110 mm. long, 3 mm. thick,
or becoming 225 mm. long and 6 mm. thick; bracts triangular-
subpeltate.
Type from Peru, Mathews 1700. San Martin (?): Spruce 4848.
Loreto: Soledad, 110 meters, Killip & Smith 29774, 29771.
Piper subsessile Trelease, sp. nov.
Frutex vel arbuscula glabra, internodiis floriferis brevibus graci-
libus; folia lanceolato-elliptica acuminata basi acuta et secus peti-
olum in alam decurrentem protracta 15-18 cm. longa 5-7 cm. lata e
2 tertiis inferioribus pinnatinervia, nervis longioribus circa 6x2,
FLORA OF PERU 241
brevioribus superne dispositis, in sicco tenuia viridia subtus minute
nigro-granulosa, petiolo brevissimo; spicae rubescentes 40 mm.
longae 3 mm. crassae obtusae, pedunculo filiformi 10 mm. longo;
bracteae triangulari- vel crescentico-subpeltatae ciliolatae; baccae
rotundatae submucronatae, stigmatibus minutis in mucrone insertis.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28747 (type in
U. S. Nat. Herb.), 28818. Yurimaguas, Killip & Smith 28031.
El Recreo, lower Rio Huallaga, Williams 3921 , 3971 . "Cordoncillo."
Piper subtextum Trelease, sp. nov.
Frutex 3-4-metralis nerviis folii subtus adpresso-pubescentibus
exceptis glaber, internodiis floriferis brevibus modice crassis; folia
lanceolato-oblonga vel elliptico-oblonga anguste acuminata basi
aliquanto inaequilaterali acuta, vel latere breviore obtuso, 15 cm.
longa 5 cm. lata e dimidio inferiore pinnatinervia, nervis circa 6x2,
in sicco tenuia, petiolo 5+2 mm. longo exalato; spicae primo curvae
90 mm. longae 3 mm. crassae, pedunculo 5 mm. longo; bracteae
parvae rotundo-subpeltatae ciliolatae; baccae pressu subangulatae,
stigmatibus minutis sessilibus.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26265 (type
in U. S. Nat. Herb.), 26278.
Piper suffruticosum Trelease, sp. nov.
Suffruticosa metralis ubique crispe flavo-villosa ; folia rotundo-
ovata vel elliptico-ovata acute acuminata basi cordata 13-15 cm.
longa 8-10 cm. lata e dimidio inferiore pinnatinervia, nervis 3x2, in
sicco tenuia rubescentia in statu adulto bullulata, petiolo 5 mm.
longo; inflorescentia non visa.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26586
(type in U. S. Nat. Herb.).
Piper Sumpi Trelease, sp. nov.
Frutex glaber, internodiis floriferis modice gracilibus elongatis;
folia late ovata breviter acuminata basi cordata 20-28 cm. longa
11-18 cm. lata e dimidio inferiore multiplinervia, nervis 6-7x2, in
sicco viridia, petiolo 2-4 cm. longo infra medium alato; spicae oppo-
sitifoliae 130 mm. longae 6 mm. crassae mucronatae, pedunculo
15-20 mm. longo; bracteae rotundae vel semirotundae subpeltatae
brunneae ciliatae; baccae subglobosae, stylo brevi crasso, stigma-
tibus 3 filiformibus.
Cuzco: San Miguel, Urubamba Valley, 2,340 meters, Cook &
Gilbert 967 (type in U. S. Nat. Herb.). "Sumpi."
242 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Piper tabinense Trelease, sp. nov. P. Hookerianum Auct.
Frutex mediocris nervis folii subtus exceptis glaber; folia ovata
acuminata basi rotundata 14 cm. longa 8 cm. lata e tertio inferiore
pinnatinervia, nervis circa 4x2, subtus ad nervos subadpresso-pube-
scentia vel glabrescentia, coriacea sublucida, petiolo 1 cm. longo;
spicae 50 mm. longae 5 mm. crassae obtusae, pedunculo vix
2 mm. longo.
Puno: Tabina, Lechler 2081, type.
Piper tardurn Trelease, sp. nov.
Frutex 3-metralis, internodiis floriferis modice brevibus et graci-
libus hamate hirsuto-hispidis; folia elliptica vel subovata acute
acuminata basi rotundata, uno latere breviore, 16 cm. longa 6-7 cm.
lata e dimidio inferiore pinnatinervia, nervis 5X2, supra valde
granuloso-asperata, subtus laxe subtomentulosa ad nervos incurvo-
pilosa, petiolo 10+2 mm. longo hirsute; spicae 100 mm. longae 3
mm. crassae, pedunculo vix 10 mm. longo molliter piloso.
Junin: Colonia Perene\ 680 meters, Killip & Smith 25025 (type
in U. S. Nat. Herb.).
Piper temptum Trelease, sp. nov.
Frutex gracilis glaber, ramulis pallido-granulosis; folia lanceolata
vel elliptico-lanceolata acuminata basi acuta 11 cm. longa 3-4 cm.
lata e dimidio inferiore pinnatinervia, nervis 4x 2, subtus punctulata,
petiolo 10 mm. longo; spicae fere rectae 80 mm. longae 3 mm. crassae.
Type collected in Peru, Mathews 1713 (herb. Kew.).
Piper tenebricosum Trelease, sp. nov.
Frutex in sicco fuscus, internodiis floriferis gracilibus brevibus
sparse sericeis; folia subelliptica ad oblanceolata vel obovata acu-
minata basi obliqua angustata et obtusa 8-10 cm. longa 3.5-5 cm.
lata e dimidio inferiore pinnatinervia, nervis 5-6x2, supra minute
granuloso-scabrida, subtus ad nervos adpresso-pubescentia, petiolo
vix 5 mm. longo pubescente; spicae 80 mm. longae 2-3 mm. crassae
acutatae, pedunculo 8 mm. longo griseo-pubescente; bracteae
rotundo-subpeltatae ciliolatae.
Loreto: Fortaleza, lower Rio Huallaga, 155-210 meters, Llewelyn
Williams 4201 (herb. Field Mus. No. 623,585, type). "Cordoncillo."
Piper tenuicuspe (Miq.) C. DC. in DC. Prodr. 16, pt. 1: 304.
1869. Artanthe tenuicuspis Miq. Lond. Journ. Bot. 4: 457. 1866.
FLORA OF PERU 243
A generally pale-villous shrub; leaves broadly oblanceolate,
slender-acuminate, gradually narrowed to the inequilateral sub-
cordulate base, 24 cm. long, 9.5 cm. wide, pinnately nerved from
the lower half, the nerves 4+5, glabrous above; petiole barely 3+3
mm. long; spikes 110 mm. long, 3 mm. thick; peduncle 5 mm. long,
hirtellous.
Type from Peru, Mathews 1704-
Piper tenuicuspe var. nervosius Trelease, var. nov.
Folia 6-7 cm. lata basi fere aequilaterali distincte cordulata.
Huanuco: Cochero, Poeppig 1463 (type in herb. Vienna).
Piper tenuistylum C. DC. in DC. Prodr. 16, pt. 1: 260. 1869.
Peltobryon Mathewsii Miq. Lond. Journ. Bot. 4: 441. 1866, non
Piper MathewsiiC. DC.
A glabrous shrub, or with the petiole and nerves puberulent
beneath; leaves elliptic or subovate, acuminate, acute-based, 14 cm.
long, 6 cm. wide, pinnately nerved throughout; petiole 10-15 mm.
long; spikes 55 mm. long, 5 mm. thick; peduncle scarcely 10 mm.
long; bracts inflexed-subpeltate; berries globose, with a slender
style; stigmas spreading.
Type from Peru, Mathews 1710.
Piper Tessmanni Trelease, sp. nov.
Frutex, ramulis subtomentosis; folia elliptica obscure acuminata
basi subaequilaterali acuta 24 cm. longa 10 cm. lata undique pin-
natinervia, nervis circa 10x2, supra fusco-granulosa, subtus bar-
bato-pubescentia, petiolo 2-3 cm. longo; spicae curvae 50 mm. longae
10 mm. crassae, mucronatae, pedunculo 10 mm. longo; bracteae
rotundo-subpeltatae ciliatae, stigmatibus 3 brevibus, stylo longo.
Loreto: Mouth of Rio Santiago, Tessmann 4197 (type in
herb. Berol.).
Piper timbuchianum Trelease, sp. nov.
Frutex nodosus, internodiis floriferis brevibus gracilibus, infe-
rioribus crassioribus transiente pubescentibus ; folia lanceolata
longiacuta basi rotundata vel typice subabrupte acuta 7-11 cm.
longa 2-3 cm. lata ubique pinnatineriva, nervis 6x2, subtus ad nervos
crispo-pubescentia, petiolo 5 mm. longo in foliis majoribus alato
crispo-pubescente; spicae juvenes 8 mm. longae 2 mm. crassae non
mucronatae, pedunculo paten te filiformi 7 mm. longo pubescente;
bracteae rotundo-subpeltatae anguste albo-marginatae.
244 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Loreto: Timbuchi, Rio Nanay, Llewelyn Williams 1036 (herb.
Field Mus. No. 617,129, type).
Piper tingens Trelease, sp. nov.
Suffrutex 2-metralis nervis folii subtus griseo-puberulis exceptis
glaber, internodiis superioribus brevibus comparate gracilibus; folia
elliptica subacuminata basi angustata obscure cordulata, uno latere
paullo breviore, 15-18 cm. longa 8-11 cm. lata e dimidio inferiore
pinnatinervia, nervis 6x2, in sicco tenuia et in statu adulto sub-
rugosa, petiolo 2 cm. longo alato; spicae 120 mm. longae 3 mm.
crassae, pedunculo gracili 10 mm. longo; bracteae rotundo- vel
crescentico-subpeltatae ciliatae.
Loreto: Yurimaguas, 180 meters, Ynes Mexia 6072 (type in herb.
Univ. Calif.).
Piper tocacheanum C. DC. in DC. Prodr. 16, pt. 1: 264. 1869.
A glabrous, glandular-punctulate shrub; leaves elliptic or lance-
elliptic, subacuminate, rather abruptly acute at the base, 26-30 cm.
long, 10-15 cm. wide, pinnately nerved throughout; petiole rather
short, winged by the decurrent blade; spikes 110 mm. long, 5 mm.
thick; peduncle 10 mm. long, curved; berries round; stigmas sessile.
San Martin: Tocache, Poeppig 1918, D1918, type.
Piper tortuosivenosum Trelease, sp. nov.
Frutex metralis, internodiis modice crassis, superioribus brevi-
bus sordido-villosis; folia lanceolata vel ovata acuta vel subacumin-
ata basi rotundata et fere sessilia vel inferiora basi subcordulato-
truncata et 1 cm. longe petiolata, petiolo vaginante, 18-22 cm.
longa 7-10 cm. lata e dimidio inferiore pinnatinervia, nervis 4-5x2,
supra et subtus at ad nervos sparse crispo-villosa obscure viridia et
tortuose venosa, subtus conspicue nigro-punctulata ; spicae 35 mm.
longae 5 mm. crassae obtusae, pedunculo 5 mm. longo laxe rigido-
villoso; bracteae rotundo- vel crescentico-subpeltatae pallido-margin-
atae; stigmata apice styli inserta.
Loreto: Mouth of Rio Santiago, 210 meters, Ynes Mexia 6199
(type in herb. Univ. Calif.).
Piper transpontinum Trelease, sp. nov.
Frutex vel arbuscula 3-5-metralis nodosa, internodiis floriferis
brevibus graciusculis laxe rigido-villosis; folia lanceolato-ovata acute
acutata basi plus minusve obliqua truncata vel breviter subcordata
16 cm. longa 5-8 cm. lata e tertio inferiore submultiplinervia, nervis
FLORA OF PERU 245
circa 4x2, in sicco tenuia sublucida, subtus ad nervos rigido-villosa
et inter nervos stellato-pilosa, petiolo 10-15 mm. longo villoso
vaginante; spicae juvenes dense brunneo-tomentosae typice
cuspidatae.
Junin: Perene Bridge, Paucartambo Valley, 700 meters, Killip
& Smith 25352 (type in U. S. Nat. Herb.), 25273.
Piper transpontinum var. ecordulatum Trelease, var. nov.
Frutex 3-4-metralis; folia vix cordata 18 cm. longa 8 cm. lata;
spicae floriferae 60 mm. longae 5 mm. crassae, pedunculo 10 mm.
longo crispo-piloso ; bracteae rotundo-subpeltatae pilosae; stigmata
3 gracilia, stylo fere aequilongo.
Junin: Perene, 680 meters, Killip & Smith 25055 (type in U. S.
Nat. Herb.).
Piper trichostylopse Trelease, sp. nov.
Frutex 2-metralis, internodiis floriferis modice brevibus et
crassis transiente sparsissime villosis vel hirtellis; folia ovata vel
lanceolato-ovata sensim acuminata basi cordulate subtruncata vel
rotundata vel subacuta 16-25 cm. longa 7-15 cm. lata e dimidio
inferiore submultiplinervia, nervis 5-6x2, supra glabra vel prope
basin paullo pubescentia, subtus ad nervos ferrugineo-villosa et
inter nervos crispo-pubescentia, petiolo 2-2.5 cm. longo villoso
glabrescente alato; spicae oppositifoliae 100 mm. longae 4 mm.
crassae, pedunculo crasso 10 mm. longo transiente plus minus ve
ferrugineo- villoso; bracteae rotundo- vel triangulari-subpeltatae
ciliatae, floribus sessilibus perfectis; ovarium glabrum obovoideum
in stylum aequilongum angustatum, stigmatibus 3 patentibus.
Huanuco: Posuso, 600 meters, Macbride 4646 (herb. Field Mus.
No. 535,721, type), 4664- Junin: La Merced, 5336.
Piper trichostylopse var. latum Trelease, var. nov.
Magis glabrata, foliis majoribus 29 cm. longis 18 cm. latis.
Huanuco: Cushi, 1,500 meters, Macbride 4849 (herb. Field Mus.
No. 535,889, type).
Piper trichostylum C. DC. Bot. Jahrb. 40: 244. 1908.
A shrub, matted-villous throughout except for the short-hairy
upper surface of the leaves; leaves ovate, acuminate, almost equilat-
erally truncate at the base with the blade abruptly contracted to
the petiole, 21 cm. long, 13 cm. wide, submultiple-nerved from below
the middle, the nerves 5x2, very rugose; petiole 2 cm. long, winged;
246 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
spikes 150-180 mm. long, 5 mm. thick; peduncle rather thick, 2 cm.
long; bracts large, round-subpeltate, yellow-hairy; berries round,
with a rather short, thick style; stigmas outcurved. Neg. 11069.
Puno: Sandia, at 2,800-3,000 meters, in bamboo thickets,
Weberbauer 728, type.
Piper tridentipilum C. DC. Notizbl. Bot. Gart. Berlin 7:
490. 1917.
A shrub, stellately crisp-subtomentose except for the upper leaf
surface; leaves elliptic-ovate, scarcely acuminate, subtruncate at the
base with one side slightly shorter and concave, 24 cm. long, 12 cm.
wide, pinnately nerved throughout; petiole 15+5 mm. long; spikes
slightly curved, 50 mm. long, 5 mm. thick; peduncle 2 cm. long;
bracts triangular-subpeltate, ciliate; berries round; stigmas short
and broad, sessile.
Type from Seringal Sao Francisco, Alto Rio Acre, Brazil, Ule
9277. Probably also in adjacent Peru.
Piper tristigmum Trelease, sp. nov.
Frutex glaber vel ad novellos et folii nervos subtus obscure
puberulus; folia elliptica anguste acuminata basi fere aequilaterali
acuta 11-12 cm. longa 4.5-5 cm. lata e 2 tertiis inferioribus pinnati-
nervia, nervis longioribus circa 6x2, in sicco firma viridia albo-
punctulata sed laevia, petiolo 5 mm. longo anguste alato; spicae
25-35 mm. longae 5 mm. crassae caudatae, pedunculo 5 mm. longo;
bracteae concavae; baccae globosae majusculae, stylo distincto,
stigmatibus 3 rotundatis.
Loreto: Caballo-cocha on the Amazon River, Llewelyn Williams
2029 (herb. Field Mus. No. 613,314, type).
Piper tuberculatum Jacq. Icon. Rar. 2: 2. pi. 210. 1786.
Steffensia tuberculata Kunth, Linnaea 13: 614. 1839. Artanthe
tuberculata Miq. Syst. 497. 1844; 111. Piper, pi. 86.
A slender- twigged, nodose shrub, glabrous except for the more or
less transiently puberulent axes and nerves beneath; leaves elliptic,
rather abruptly acute, rounded at the base or cordulate with one
side distinctly shorter, 9-14 cm. long, 5-7 cm. wide, pinnately nerved
throughout, drying thin; petiole 5+5 mm. long, winged throughout;
spikes 100 mm. long, 3-4 mm. thick or larger; peduncle 10-15 mm.
long; bracts triangular-subpeltate; berries depressed-ellipsoid; stig-
mas 3, small, broad, sessile.
FLORA OF PERU 247
Loreto: Puerto Arturo, in pasture, 200 meters, Williams 5184.
Masisea, Killip & Smith 26862. Santa Rosa, in forest, Williams
4805. San Martin : Tarapoto, edge of river, Williams 5442. Widely
distributed, from Brazil to Mexico. "Cordoncillo."
Piper tumidicondyli Trelease, sp. nov.
Frutex glaber, internodiis brevibus, nodis incrassatis; folia lucida
ovata subacuminata basi rotundata, uno latere paullo breviore,
10-13 cm. longa 4-6 cm. lata e dimidio inferiore submultiplinervia,
nervis circa 5x2, in sicco coriacea anguste revoluta, petiolo 10 mm.
longo; spicae pendulae curvae 40 mm. longae 5 mm. crassae breviter
cuspidatae, pedunculo 10 mm. longo; bracteae rotundo-subpeltatae,
margine lato brunneo.
Loreto: Rancho Indiano, Iquitos, 110 meters, Ynes Mexia 6431
(type in herb. Univ. Calif.).
Piper ucayalianum Trelease, sp. nov.
Frutex, internodiis graciusculis superioribus brevibus sparse
hispidis et serius minute pallido-granulosis; folia lanceolato-ovata
sensim acuminata basi paullo inaequilaterali rotundata 12-16 cm.
longa 5-7.5 cm. lata e dimidio inferiore pinnatinervia, nervis 5X2,
in sicco ex parte rugosa, supra minute granulosa et aliquanto lepidota,
subtus ad nervos strigosa, petiolo 8+2 mm. longo hispido; spicae
110 mm. longae 2 mm. crassae, pedunculo vix 10 mm. longo sparse
scabro-hispido ; bracteae transverso-subpeltatae ciliolatae.
Loreto: Lower Rio Ucayali, Tessmann 3011 (type in herb.
Stockholm).
Piper udisilvestre C. DC. Notizbl. Bot. Gart. Berlin 7: 485.
1917. P. longifolium var. pilosum C. DC. op. cit. 492.
A somewhat dark-punctulate shrub, glabrous except for the some-
what puberulent nerves beneath; leaves elliptic-subobovate, acute
at both ends, 35 cm. long, 18-20 cm. wide, pinnately nerved through-
out; petiole 2 cm. long; spikes in fruit 70-90 mm. long, 20 mm.
thick; peduncle thick, 15 mm. long; berries obconic-obovoid, with a
long slender style; stigmas small.
Loreto : Pampas de Ponasa, Ule. Type from Seringal Sao Fran-
cisco, Alto Rio Acre, Brazil, Ule 9296.
Piper ulceratum Trelease, sp. nov.
Frutex, internodiis superioribus modice brevibus et crassis
flavido-villosis; folia elliptico-obovata acuminata basi breviter
248 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
cordata 22 cm. longa 9 cm. lata e tertio vel dimidio inferiore pin-
natinervia, nervis 6-7x2, bullulata, supra sparse stellato-pilosa,
subtus villosa, petiolo crasso vix 10 mm. longo villoso; inflorescentia
non visa.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
6962 (herb. Field Mus. No. 622,978, type).
Piper ulceratum var. angustifolium Trelease, var. nov.
A typo differt foliis lanceolato-ellipticis vel lanceolatis 16-20
cm. longis 6-8 cm. latis, basi angustata obscure cordulatis; spicae
curvae caudatae 30 mm. longae 3 mm. crassae, pedunculo filiformi
40 mm. longo reflexo hirsute; bracteae crescenticae; baccae rotun-
datae vel cum rhachi subelongatae paullo mucronatae, stigmatibus
3 sessilibus.
San Martin: San Roque, 1,350-1,500 meters, Llewelyn Williams
6982 (herb. Field Mus. No. 629,612, type).
Piper Uleanum Trelease, sp. nov.
Frutex rigide hirsuto-villosus; folia subovato-elliptica acuminata
basi aequilaterali et sublata paullo cordulata 12 cm. longa 5 cm. lata
e dimidio inferiore pinnatinervia, nervis 5x2, rugosa, supra granuloso-
scabra, subtus minute punctulata et ad nervos incurvo-hirsuta,
petiolo vix 10 mm. longo; spicae 80 mm. longae 2 mm. crassae
rectae, pedunculo 10-15 mm. longo sparse hirsute; bracteae sub-
peltatae parvae, margine ciliato plumbeo-tincto.
Type from Seringal Auristella, Rio Acre, Brazil, Ule 9288 (type
in herb. Berol.), and probably extending into Peru.
Piper unguiculatum R. & P. Fl. 1: 34. pi 57. 1798. P. glauces-
cens Jacq. Eclog. 112. pi. 76. 1811. Gonistum unguiculatum Raf.
Sylv. Tellur. 85. 1838. Enckea unguiculata Kunth, Linnaea 13:
592. 1839. E. glaucescens Kunth, Linnaea 13: 594. 1839.
A slender shrub, glabrous or the axes and leaf bases transiently
and sparsely velvety; leaves ovate, blunt-acuminate, obtuse or
subtruncate at the base with the margin commonly inrolled next the
petiole, 5-9 cm. long, 3-4 cm. wide, 5-7-nerved; spikes 20-40 mm.
long, 3 mm. thick; peduncle 15 mm. long; bracts spatulate; berries
ovoid ; stigmas sessile.
Huanuco: Posuso and Chinchao, Ruiz & Pavon, type. Intro-
duced into cultivation at Madrid by Ruiz and Pavon, and at present
known only as cultivated in botanical gardens.
FLORA OF PERU 249
Piper urubambanum Trelease, sp. nov.
Frutex glaber, ramulis gracilibus; folia ovata acuminata basi
aequaliter rotundata, uno latere paullo breviore, 12-14 cm. longa
4.5-7 cm. lata e dimidio inferiore multiplinervia, nervis circa 4x2,
petiolo gracili 15-18+2 mm. longo alato; spicae oppositifoliae 60-70
mm. longae 5 mm. crassae, pedunculo gracili 10 mm. longo; bracteae
triangulari-subpeltatae, margine flavescente ciliato; baccae sub-
globosae, stigmatibus 3 filiformibus sessilibus deciduis.
Cuzco: San Miguel, Urubamba Valley, 1,800 meters, CooA; &
Gilbert 1122 (type in U. S. Nat. Herb.).
Piper velutininervosum Trelease, sp. nov.
Arbuscula 4-5-metralis petiolis et folii nervis subtus velutinis
exceptis glabra, internodiis superioribus gracilibus elongatis; folia
subovata subacuta basi inaequilaterali cordata, sinu aperto, latere
longiore auriculato, 28 cm. longa 14 cm. lata e 2 tertiis inferioribus
subpinnatinervia, nervis 5X2, in sicco tenuia lucida brunnescentia,
petiolo 4-6 cm. longo, uno latere 5 mm. breviore alato; spicae juvenes
parvae breviter pedunculatae.
Junin: East of Quimiri Bridge, La Merced, 800-1,300 meters,
Kittip & Smith 24006 (type in U. S. Nat. Herb.).
Piper velutinum HBK. Nov. Gen. & Sp. 1: 52. 1815. Schilleria
velutina Kunth, Linnaea 13: 701. 1839. Artanthe velutina Miq. Syst.
471. 1844.
A white-hirsute- villous shrub; leaves lance-ovate or oblong,
acuminate, inequilaterally obtuse at the base, 14-15 cm. long, 4.5-5
cm. wide, pinnately nerved from the lower half, the nerves about
5x2; petiole 5-10 mm. long; spikes curved, 60 mm. long, 2 mm.
thick; peduncle 10 mm. long; berries depressed-cylindric ; stigmas
sessile.
Cajamarca: Zaulaca and San Felipe, Jaen de Bracamoros,
Humboldt at Paris, and in herb. Willdenow as A 7 o. 667.
Piper verbascifolium (Miq.) C. DC. in DC. Prodr. 16, pt. 1:
258. 1869, as to Peru only. Artanthe verbascifolia Miq. Syst. 431.
1844. P. acutifolium var. subverbascifolium C. DC. Bot. Jahrb.
40: 247. 1908.
A soft-hairy shrub; leaves sublanceolate-elliptic, gradually
pointed, inequilateral at the base and cordate-auricled with lateral
sinus, 16-20 cm. long, 5-7 cm. wide, pinnately nerved from the lower
half, the nerves 7+8, glabrescent above, bullulate; petiole scarcely
250 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
5 mm. long; spikes nearly straight, becoming 90 mm. long, 5 mm.
thick; peduncle 5 mm. long; bracts roundish-subpeltate, ciliolate;
berries rounded-sub triquetrous; stigmas 3, linear, sessile. Neg. 7233.
Type from Peru, Dombey 932; Gay and Grisar. Amazonas:
Mathews in 1836. Huanuco: Casapi, Mathews, 1715. Huanuco,
Mathews 1795 (with cuspidate spikes). Muna, Pearce in 1863.
Huassihuassi, Dombey 504. Junin: Tarma, Weberbauer 2163.
Loreto: Pongo de Manseriche, Tessmann 4779.
Piper veredianum Trelease, sp. nov.
Frutex nodosus glaber glanduloso-punctulatus; folia lanceolata
acute acuminata basi inaequilaterali longe acuta 14-16 cm. longa
5-6 cm. lata e 2 tertiis inferioribus pinnatinervia, nervis circa 5x2,
petiolo 1 cm. longo, lamina basi decurrente; spicae 80 mm. longae
3 mm. crassae, pedunculo 10 mm. longo; bracteae triangulari-
subpeltatae ciliatae; baccae teretes vel 3-4-gonae truncatae, stig-
matibus 3 sessilibus.
Loreto: Caballo-cocha on the Amazon River, Llewelyn Williams
2106 (herb. Field Mus. No. 608,810, type). "Cordoncillo."
Piper verruculaecaule Trelease, sp. nov.
Frutex 3-4-metralis, internodiis superioribus gracilibus brevius-
culis evanescente brevipilosis serius albide granuloso-asperatis;
folia lanceolato-elliptica vel subobovato-elliptica acuminata basi
subinaequilaterali acuta 15-20 cm. longa 6-10 cm. lata, supra minute
granuloso-asperata, e dimidio inferiore pinnatinervia, nervis 6x2,
subtus ad nervos incurvo-hispida, petiolo 10-15+3 mm. longo
exalato; spicae juveniles 30 mm. longae 2 mm. crassae.
Ayacucho: Near Kimpitiriki, Apurimac Valley, 400 meters,
Kittip & Smith 23025 (type in U. S. Nat. Herb.; dupl. in herb.
Field Mus.).
Piper verruculaefolium Trelease, sp. nov.
Frutex 1.5 m. altus, internodiis floriferis brevibus gracilibus
incurvo-hirsutis in statu adulto granulosis; folia lanceolata anguste
acuminata basi inaequilaterali obtusa vel subcordulata 9-12 cm.
longa 3-4 cm. lata e dimidio inferiore pinnatinervia, nervis 4-5x2,
supra rugoso-verruculosa et primum scabra, subtus ad nervos
hirsuta, petiolo vix 5+3 mm. longo exalato hirsuto; spicae 70 mm.
longae 3 mm. crassae cuspidatae, pedunculo 6 mm. longo primo
granuloso; bracteae rotundo-subpeltatae ciliolatae.
FLORA OF PERU 251
Junin: La Merced, 700 meters, Killip & Smith 24089 (type in
U. S. Nat. Herb.).
Piper villosispicum Trelease, sp. nov.
Arbuscula sordide villosa 4-6-metralis, internodiis floriferis
breviusculis gracilibus; folia oblonga anguste acuminata basi in-
aequilaterali truncato-cordulata 16 cm. longa 6 cm. lata e dimidio
inferiore submultiplinervia, nervis 5X2 inferne approximatis, in
sicco chartacea lucida, petiolo 5 mm. longo vaginante; spicae 40-50
mm. longae 5-6 mm. crassae, pedunculo crasso 10 mm. longo;
bracteae dense sordido-villosae; baccae rotundatae, stigmatibus 3
apice styli insertis.
Junin: Santa Rosa, Pichis Trail, 625-900 meters, Killip & Smith
26178 (type in U. S. Nat. Herb.). Schunke Hacienda, La Merced,
Killip & Smith 24533.
Piper virgulticola Trelease, sp. nov.
Frutex 5-metralis nodosus, internodiis superioribus gracilibus
brevibus ochroleucis dense hispidis; folia lanceolato-elliptica acute
acuminata basi inaequilateralia, saltern latere breviore basi rotun-
dato, 15-16 cm. longa 5-6.5 cm. lata e dimidio inferiore pinnatinervia,
nervis circa 6x2, supra albo-granulosa et breviter scabra, subtus ad
nervos incurvo-pubescentia, petiolo 5-10+2 mm. longo dense hispido;
spicae 70 mm. longae 2-3 mm. crassae, pedunculo 5 mm. longo his-
pido; bracteae rotundo-subpeltatae ciliolatae.
Loreto: In thicket, Fundo Indiano, Iquitos, 110 meters, Ynes
Mexia 6430 (type in herb. Univ. Calif.).
Piper viridilimbum Trelease, sp. nov.
Frutex glaber 3-metralis, internodiis floriferis gracilibus brevius-
culis; folia lanceolata acuminata basi acuta, uno latere paullo
breviore, 16 cm. longa 6.5-7.5 cm. lata e dimidio inferiore sub-
multiplinervia, nervis 4-5x2, in sicco tenuia viridia, subtus pallidiora,
petiolo 12-15+2 mm. longo infra medium vaginante; spicae juvenes
10 mm. longae 1 mm. crassae, pedunculo duplo breviore.
Ayacucho: Kimpitiriki, Rio Apurimac Valley, 400 meters, Killip
& Smith 22990 (type in U. S. Nat. Herb.).
Piper volubile C. DC. Bot. Jahrb. 40: 245. 1908.
A climbing shrub; twigs subretrorsely appressed-gray-hirsute ;
leaves lanceolate, slender-acuminate, obtuse or inequilaterally
subacute at the base, 11 cm. long, 4 cm. wide, pinnately nerved from
252 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
the lower half, the nerves 4-5x2, coarsely white-granular-scabrous
on both sides with the nerves beneath subappressed-hairy ; petiole
scarcely 5 mm. long; spikes gently curved, 100 mm. long, 3 mm.
thick, mucronate; bracts subpeltate; berries minute, subterete;
stigmas sessile. Neg. 11079.
Huanuco: Monzon, at 900-100 meters, in forest, Weberbauer
3444, type.
Piper xapurayense C. DC. Notizbl. Bot. Gart. Berlin 6:
492. 1917.
A shrub with axes and nerves beneath stiffly subvillous; leaves
elliptic, subacuminate, almost equilaterally cordulate, 9-16 cm.
long, 4-7 cm. wide, pinnately nerved from the lower half, the nerves
4-6x2, glabrescent above; petiole 10 mm. long, winged; spikes 15
mm. long, 10 mm. thick, cuspidate; berries round, with a long style;
stigmas short.
Type from Alto Zapury, Seringal Sao Francisco, Rio Acre,
Brazil, Ule 9285, 9286. Doubtless also in Peru.
Piper yananoanum Trelease, sp. nov.
Arborescens 10-metralis nodosa, internodiis floriferis elongatis
gracilibus ad nodos paullo pilosulis minute pallido-granulosis; folia
lanceolata ad ovata acuminata basi subacuta 12-18 cm. longa
4.5-9.5 cm. lata e dimidio inferiore submultiplinervia, nervis 4-5x2,
subtus ad nervos pilosula, petiolo 1.5 cm. longo velutino alato;
spicae oppositifoliae 150 mm. longae 3-5 mm. crassae, pedunculo
vix 10 mm. longo velutino; bracteae triangulari-subpeltatae ciliatae,
floribus sessilibus perfectis; baccae obconicae glabrae, stigmatibus 3
patentibus subsessilibus.
Huanuco: Yanano, 1,800 meters, Macbride 3796 (herb. Field
Mus. No. 534,858, type).
Piper yessupianum Trelease, sp. nov.
Frutex glaber 1-2-metralis, internodiis floriferis breviusculis
crassiusculis; folia lanceolato-oblonga sensim acutata basi subaequi-
laterali acuta 16-20 cm. longa (et ultra?) 4.5-6 cm. lata ubique
pinnatinervia, nervis 8-10x2, in sicco tenuia fusca sublucida,
petiolo in foliis superioribus nullo, lamina ad caulem decurrente, in
foliis inferioribus 2 cm. longo sursum e basi alato; spicae 35 mm.
longae 5 mm. crassae, pedunculo gracili 15 mm. longo; bracteae
concavae glabrae; stigmata sessilia.
FLORA OF PERU 253
Junin: Puerto Yessup, 400 meters, Killip & Smith 26373 (type
in U. S. Nat. Herb.).
Piper yurimaguasanum Trelease, sp. nov.
Frutex 3-6-metralis nodosus, internodiis floriferis brevibus
gracilibus primo puberulis; folia lanceolato-elliptica vel oblonga
atque acuta, basi inaequilaterali rotundata ubique pinnatinervia,
subtus ad nervos aliquanto crispo-puberula, petiolo 3+8 mm. longo
alato; spicae 80 mm. longae 2 mm. crassae, pedunculo gracili 10 mm.
longo paullo puberulo; bracteae rotundo- vel triangulari-subpeltatae
ciliatae; baccae paullo cum rhachi elongatae, stigmatibus 3 sessilibus.
Loreto: Iquitos, 100 meters, Killip & Smith 272? '4 (type in U. S.
Nat. Herb.), 29045. Yurimaguas, Killip & Smith 27628, 29045.
San Martin: Tarapoto, Williams 5442. Lower Rio Huallaga, Wil-
liams 5184.
4. PLEIOSTACHYOPIPER Killip, Smith & Trelease, gen. nov.
Frutices, foliis ut in sectione Enckea generis Piper palmatinerviis;
flores perfectae, in spicas globosas solitarias et pedunculum ebractea-
tum oppositifolium terminantes vel typice cum pedunculo collateral!
1-bracteato spica solitaria globosa terminato dispositae, perfectae;
ovarium in stylum aequilongum attenuatum.
Type species, Pleiostachyopiper nudilimbum (C. DC.) Killip,
Smith & Trelease.
Pleiostachyopiper nudilimbum (C. DC.) Killip, Smith &
Trelease, comb. nov. Piper nudilimbum C. DC. Verh. Bot. Ver.
Brandenb. 47: 113. 1905.
A glabrous shrub with slender, green branches; leaves elliptic,
varying to lance-ovate, gradually acute, 11-16 cm. long, 5-8 cm.
wide, 5-nerved, drying firm and green, the petiole 10-15 mm. long;
spikes almost globose, 10 mm. long and 10 mm. broad, the peduncle
about as long as the spike, that of the collateral spike supported by a
stalk 1 or even 2 cm. long with a narrow bract at the top; bracts
clavate-subpeltate ; berries ovoid, with an equally long style; stigmas
3, broad, connivent.
Junin: Puerto Bermudez, Killip & Smith 2644$ > Loreto:
Soledad, Killip & Smith 29562, 29731. San Martin: Chorrillo,
Tessmann 30383. Type from Jurua Miry, Amazonas, Brazil; also
Monte Mo, Vie 9281 ; Seringal Sao Francisco, Ule 9282.
