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FIELD MUSEUM OF NATURAL
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S' NATURAL
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THE LIBRARY OF THE
MAR 151939
UNIVERSITY OF ILLINOIS
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1936-1938 !
PRINTED IN THE UNITED STATES OF AMERICA
BY FIELD MUSEUM PRESS
BOTANICAL SERIES
FIELD MUSEUM OF NATURAL HISTORY
FOUNDED BY MARSHALL FIELD, 1893
VOLUME XIII
FLORA OF PERU
PART II
BY
J. FRANCIS MACBRIDE
ASSISTANT CURATOR OF THE HERBARIUM, DEPARTMENT OF BOTANY
B. E. DAHLGREN
CURATOR, DEPARTMENT OF BOTANY
EDITOR
PUBLICATION 357
CHICAGO, U.S.A.
JUNE 10, 1936
PRINTED IN THE UNITED STATES OF AMERICA
BY FIELD MUSEUM PRESS
FB
v, \$r
FLORA OF PERU
J. FRANCIS MACBRIDE
40. PI PER AGE AE. Pepper Family
By William Trelease
Herbs, shrubs, or small trees; leaves usually alternate, some-
, times opposite or verticillate, entire; stipules none or adnate to the
petiole; flowers minute, usually green, perfect or unisexual, in spikes
i or racemes, these pedunculate, terminal or opposite the leaves,
sometimes axillary or several on a common peduncle, the flowers
usually densely crowded in the spike, sometimes annulately or
spirally arranged; bracts usually peltate, sessile or stipitate, some-
times adnate or concave; perianth none; stamens usually 2-6, the
filaments free, the anthers terminal, erect, 2-celled; ovary sessile
or stipitate, 1-celled, 1-ovulate; fruit baccate, very small.
Stigma 1; plants herbaceous, usually small, most often epiphytic.
1. Peperomia.
Stigmas usually 3; plants large, usually woody and terrestrial.
Spikes in an axillary umbel 2. Pothomorphe.
Spikes opposite the leaves.
Spikes solitary 3. Piper.
Spikes often accompanied by a collateral spike.
4. Pleiostachyopiper.
1. PEPEROMIA R. & P.
Annual or perennial herbs, epiphytic or terrestrial, usually very
fleshy; leaves alternate, opposite, or verticillate, without stipules;
flowers scattered or densely crowded upon the rachis, often partly
immersed in pits; stamens 2. A vast genus, with many species in
almost all parts of tropical America.
A. Plants acaulescent.
Leaves distinctly peltate.
Leaves scarcely 1 cm. in diameter; spikes single.
Spikes scarcely 10 cm. long.
Berries much wrinkled; stylopodium thick.
Berries ellipsoid P. parvifolia.
3
1 067037
4 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Berries subovoid P. verruculosa.
Berries not wrinkled.
Corm small P. minuta.
Corm large P. macrorhiza.
Spikes distinctly longer; stylopodium elongate.
Spikes 20 mm. long : P. Seleri.
Spikes 60-80 mm. long P. Hillii.
Leaves 1-2 cm. in diameter; stylopodium elongate.
Spikes scarcely over 10 mm. long P. umbilicata.
Spikes 20-60 mm. long P-..peruviana.
Leaves 2-4 cm. in diameter.
Spikes 15-30 mm. long P. falsa.
Spikes 50-60 mm. long.
Leaves from a corm P. scutellaefolia.
Leaves from a rhizome P. rupiseda.
Spikes 90-100 mm. long P. polycephala.
Leaves not peltate, 2-4 cm. wide.
Spikes single, elongate P. Sprucei.
Spikes paniculate, short P. villipetiola.
A. Plants with leafy stems.
B. Leaves alternate but sometimes congested upward.
C. Leaves distinctly peltate.
Spikes paniculate; leaves 10-15 cm. long P. tarapotana.
Spikes not paniculate.
Leaves 10-15 cm. long.
Plants glabrous.
Spikes from axils of reduced leaves P. dependens.
Spikes subtended by ordinary leaves P. variegata.
Plants hairy P. Schunkeana.
Leaves 6-10 cm. long.
Leaves glabrous P. rugata.
Leaves ciliate P. majalis.
Leaves commonly 4-6 cm. long.
Plants glabrous.
Leaves round-ovate . .' P. deficiens.
Leaves more elongate P. puberulaeformis.
FLORA OF PERU 5
.Plants crisp-pubescent P. cuprea.
Plants subvillous P. arbor igaudens.
Leaves 2.5-4 cm. long; plants pubescent P. flavescens.
C. Leaves at most obscurely subpeltate at the base.
D. Spikes paniculate.
Leaves 10-15 cm. long, pinnately nerved.
Spikes very short; glabrous P. Lechleriana.
Spikes elongate.
Plants pubescent.
Leaves ovate P. spectabilis.
Leaves elliptic P. striata.
Plants glabrous.
Leaves elliptic-oblanceolate P. balsapuertana.
Leaves oblanceolate.
Panicle branched P. arcualispica.
Panicle simple . P. uncatispica.
Leaves lanceolate P. acutifolia.
Leaves 6-10 cm. long, pinnately nerved, glabrous.
P. Weberbaueri.
Leaves 4-6 cm. long.
Leaves round, cordate, submultiple-nerved.
Plants appressed-villous ; panicle ample P. secunda.
Plants subglabrescent; panicle with a few branches.
P. Macbndeana.
Leaves obovate, acute-based, glabrous.
Leaves palmately nerved P. arborised a.
Leaves pinnately nerved P. obliqua.
Leaves 2.5-4 cm. long, cleaver-shaped, vertical rather than hori-
zontal P. dolabriformis.
Leaves 1-2.5 cm. long.
Leaves compressed from the side.
Leaves glabrous P. nualis.
Leaves ciliate on the keel.
Spikes 20 mm. long .P. lepadiphylk i.
Spikes 40 mm. long P. naviculaefolia.
6 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves broader than thick P. macrolhyrsa.
D. Spikes umbellate or solitary, attached at the end of the
peduncle.
Leaves 2.5-4 cm. long.
Plant glabrous P. foliiflora.
Plant pubescent P. Haenkeana.
Leaves scarcely 2.5 cm. long.
Leaves cordate; spikes several P. phyllaniha.
Leaves not cordate; spikes solitary.
Leaves 5-7-nerved P. Fortunati.
Leaves 3-nerved P. umbelliformis.
D. Spikes neither paniculate nor umbellate nor attached to the
end of the peduncle.
* E. Leaves 10-15 cm. long.
Plants villous; leaves ovate P. yapasana.
Plants crisp-pubescent; leaves elliptic, narrowly acuminate.
P. pubiramea.
Plants glabrous.
Leaves ovate, cordate or round-based.
Stem very short; nerves palmate P. curtipes.
Stem elongate.
Nerves palmate P. pinedoana.
Nerves multiple.
Leaves with nearly simple midrib P. itayana,.
Leaves nerved from the lower third.
P. pseudorhynchophoro,.
Leaves nerved from the lower two-thirds.
P. pergamentacea.
Leaves elliptic, acute-based, pinnately nerved.
Leaves half as broad as long.
Ovary sharp-beaked P. quimiriana.
Ovary not beaked P. exiliramea.
Leaves more elongate . P. tenuiramea.
Leaves lanceolate, pinnately nerved.
Spikes 2 mm. thick P. binispica.
Spikes 5 mm. thick P. huacachiana.
FLORA OF PERU 7
Leaves obovate-palmate, pinnately nerved.
Spikes 2 mm. thick.
Leaves scarcely emarginulate P. cruciata.
Leaves emarginulate ; P. Fieldiana.
Spikes 5-10 mm. thick P. emarginata.
E. Leaves 6-10 cm. long.
F. Leaves palmately nerved.
Leaves broad.
Leaves velvety throughout P. pubinervosa.
Leaves fugaciously pubescent P. rhombiformis.
Leaves ciliate and locally puberulent, blunt-acuminate.
P. Poeppigii.
Leaves glabrous.
Leaves acute at the base.
Leaves prominently cross- veined P. enckeaefolia.
Leaves not prominently cross- veined .... P. chartacea.
Leaves rounded at the base or cordulate.
Leaves pale-areolate P. areolata.
Leaves white between the nerves.
Leaves orbicular P. procumbens.
Leaves subelliptic P. sarcostachya.
Leaves purple beneath P. monostachya.
Leaves uniformly rosy P. rubrifolia.
Leaves elongate, acute-based.
Stem winged P. alata.
Stem not winged P. non-alata.
F. Leaves pinnate- or multiple-nerved.
G. Leaf base acute.
Plants glabrous.
Leaves clustered at the top of the stem, acuminate.
P. racemifolia.
Leaves scattered.
Spikes not paired.
Leaves subobovate P. acuminata .
Leaves ovate P. purpurea.
Leaves subrhombic-elliptic P. basellaefolia.
8 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves lanceolate .P. bermudezana.
Spikes paired.
Leaves obovate P. eleganti/olia.
Leaves ovate-elliptic P. granulatilimba.
Leaves lance-ovate P. santiagoana.
Leaves elliptic P. elegans.
Leaves rhombic-ovate P. mercedana.
Leaves and petioles ciliate.
Leaves elliptic P. ciliatifolia.
Leaves broadly lanceolate.
Spikes 150 mm. long P. quatrometralis.
Spikes shorter.
Plant subrepent P. semisupina.
Plant erect P. blepharipus.
Plants locally puberulent P. limbuchiana.
Plants crisp-pubescent P. enenyasensis.
G. Leaf-base auriculate P. pilosa.
G. Leaves cordate, submultiple-nerved.
Leaves about 5 cm. wide.
Plants somewhat pubescent ...:.... P. juniniana.
Plants glabrous P. erylhroslachya .
Leaves wider, glabrous.
Stem short and few-leaved P. curticaidis.
Stem elongate.
Leaves subtruncate at the base P. ainana.
Leaves cordate.
Peduncle 3 cm. long P. longepedunculata.
Peduncle 7-10 cm. long P. Opiziana.
G. Leaf base rounded P. santiagoana.
E. Leaves 4-6 cm. long.
F. Leaves pinnate- or multiple-nerved.
Leaf base acute or subacute.
Leaves scarcely 2 cm. wide.
Stems compactly branched.
Leaves papillate P. crystalline.
Leaves not papillate P. metallica.
FLORA OF PERU 9
Stem slender and elongate , .P. pseud o-salicifolia.
Leaves distinctly wider.
Berries slender-beaked.
Plants pubescent.
Leaves acute P. syluatica.
Leaves obtuse P. teresitensis.
Plants glabrous.
Leaves elliptic-obovate.
Spikes 50 mm. long P. emarginulala.
Spikes longer P. deodorata.
Leaves round-obovate P. haematolepis.
Leaves oblanceolate P. parvipynctulala.
Leaves rhombic-elliptic ........ P. elhplico-rhombea.
Berries mucronate P. rhombilimba.
Berries Scutulate-truncate.
Plants tomentulose P. pertomenlella.
Plants transiently pubescent; leaves ovate.
Leaves drying coriaceous P. coloniac.
Leaves drying thin P. pachiteana.
Plants somewhat puberulent P. cilialifolia.
Plants somewhat ciliate and granular.
Leaves lance-elliptic '. . . . .P. scutalei 'folia.
Leaves elliptic P. flavescentifolia.
Leaves subovate P. granulalifolia.
Leaves cordate, glabrous.
Leaves becoming subtriangular P. crotalophora.
Leaves round-ovate.
Stem slender and fragile . P. fragilissima.
Stem stronger P. huacapistanana.
F. Leaves palmately nerved.
Leaves cordate.
Plants puberulent P, Grisari.
Plants glabrous.
Spikes solitary.
Leaves subacuminate P. gibba.
Leaves obtuse P. rufescens.
10 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Spikes paired P. currucaeform.is.
Leaves rounded at the base, glabrous.
Leaves obtuse P. alismaefolia.
Leaves subacute.
Blade narrowed to the petiole, subacuminate.
P. rufescentifolia.
Blade crossing the petiole P. verediana.
Leaves acute at the base.
Plants glabrous.
Leaves elongate-lanceolate, long-attenuate to apex.
P. salicifolia.
Leaves lance-elliptic.
Peduncle about 1 cm. long.
Spikes short P. Vidaliana,
Spikes 90 mm. long P. exiguispica.
Spikes 100-150 mm. long P. spiculata.
Spikes 150-200 mm. long P. oxyphylla.
Peduncle 2 cm. long P. montana.
Peduncle 2-3 cm. long P. obruenda.
Leaves elliptic.
Peduncle 5 cm. long P. variculata.
Peduncle 1 cm. long P. puberulibacca.
Plants velvety P. dasystachya.
Plants crisp-pubescent.
Spikes with pseudo-pedicels P. crusculibacca.
Spikes without pseudo-pedicels P. leuconeura.
Plants villous P. tricolor.
E. Leaves 2.5-4 cm. long.
H. Leaves palmately nerved.
Leaf base acute.
Leaves 5-7-nerved.
Leaves at most ciliate or ciliate-lined.
Leaves drying thick.
Spikes short P. hamiUonianifoUa.
Spikes long P. induratifolia.
FLORA OF PERU 11
Leaves drying rather thin.
Spikes 70 mm. long P. megalepis.
Spikes about 40 mm. long.
Leaves suborbicular P. furcata.
Leaves subelliptic P. microlepis.
Leaves subobovate P. debilipes.
Leaves ovate P. Obex.
Leaves crisp-pubescent.
Leaves subovate-elliptic P. famelica.
Leaves round-ovate P. nigricans.
Leaves 3-nerved.
Plants glabrous.
Stem angled P. alatiscapa.
Stem not angled.
Spikes about 50 mm. long.
Leaves oblanceolate P. fortipes.
Leaves lance-oblong P. carrapana.
Spikes long P. chanchamayana.
Plants subpuberulent P. caducifolia.
Plants crisp-pubescent.
Leaves elliptic-subobovate P. hyporhoda.
Leaves lance-ovate P. trinervis.
Leaf base obtuse or cordulate.
Plants appressed-pubescent P. carpapatana.
Plants crisp-pubescent P. chalhuapuquiana.
Plants subpuberulent P. Maransara.
Plants glabrous.
Plant erect, pellucid P. pellucida.
Plant repent P. pseudo-furcata.
H. Leaves pinnately nerved.
Plants glabrous P. ventricosicarpa.
Plants villous P. yananoensis.
Plants transiently silky P. percalvescens.
E. Leaves 1-2.5 cm. long.
Leaves pinnately nerved, acute-based.
12 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Plants glabrous.
Leaves obovate ,...-. P. parva.
Leaves elliptic-obovate P. roqueana.
Leaves oblanceolate . P. cusilluyocana.
Plants subvillous P. pilifera.
Leaves palmately nerved.
Leaves acute-based.
Plants glabrous.
Leaves lanceolate.
Leaves acute P. aerea.
Leaves obtuse P. mansericheana.
Leaves subrhombic P. rhombeifolia,
Plants crisp-pubescent P. pumila.
Plants subvillous . . P. Drapeta.
Leaves cordate or round-based.
Plants glabrous P. antoniana.
Plants somewhat crisp- villous P. pontina.
Plants crisp-pubescent P. scandens.
E. Leaves scarcely 1 cm. long, if somewhat longer the blades
narrow.
I. Leaves nearly as broad as long.
Leaves subreniform.
Plants crisp-pubescent, very delicate, repent . . P. scandens.
Plants soft-hirtellous P. perhispidula.
Leaves round-ovate P. longipila.
Leaves orbicular.
Leaves varying into elliptic.
Plants glabrous P. multiformis.
Plants finely subvillous.
Peduncle 5-10 mm. long P. pleiomorpha.
Peduncle very short P. pichisemis.
Plants crisp-pubescent.
Peduncle very short P. polymorpha.
Peduncle 5 mm. long P. vana.
Leaves uniform.
FLORA OF PERU 13
Plants glabrous : P. fuscispica.
Plants somewhat pilose P. andinacea.
Plants crisp-pubescent P. rotundifolia.
Leaves round-obcordate.
Plants puberulent P. cordulilimba.
Plants glabrous P. obcordatifolia.
Leaves obliquely elliptic P. pruinosi folia.
I. Leaves subelongate-ovate.
Leaves scarcely more than 1-nerved.
Plants transiently silky P. subsericata.
Plants at most obscurely puberulent.
Leaves very variable P. heterophylla.
Leaves uniform P. playapampana.
Leaves 5-nerved P. modicilimba.
Leaves obscurely pinnately nerved.
Leaves half as broad as long P. non-hispidula.
Leaves narrower P. plicatifolia.
B. Leaves opposite or whorled.
J. Leaves 6-10 cm. or more long, glabrous.
Leaves 3-nerved P. heterostachya.
Leaves 5-nerved P. tetragona.
Leaves 7-9-nerved.
Leaves lance-elliptic P. septemnervis.
Leaves elliptic P. undeninerma.
Leaves oblanceolate P. deceptrix.
J. Leaves 4-6 cm. long.
Plants glabrous.
Drying purple.
Leaves narrowly lanceolate P. rubeo,.
Leaves lance-elliptic or ovate P. rubens.
Not drying purple.
Leaves 5-nerved P. puteolata.
Leaves 7-nerved P. san-roqueana.
Plants locally puberulent.
Leaves lance-elliptic P. Pavoniana.
Leaves subovate-elliptic P. mishuyacana.
14 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Plants generally puberulent P. inaequilatera.
Plants with the stem pubescent.
leaves glabrous P. exilamenta.
Leaves pubescent above on the nerves, acuminate.
P. pubescentinervis.
Leaves generally pubescent.
Upper leaves reduced P. pampalcana.
Upper leaves not smaller P. casapiana.
J. Leaves 2.5-4 cm. long.
Leaves commonly opposite.
Leaves papillate-puberulent, acute at the base.
P. ollantaitambona.
Leaves pubescent.
Upper leaves reduced P. confertispica.
Upper leaves scarcely smaller.
Plant crisp-pubescent P. apurimacana.
Plant subvillous-tomentose P. cotoneasterifolia.
Plant hirsute P. Gayi.
Leaves mostly whorled.
Plants glabrous.
Leaves drying thin and green P. profissa.
Leaves drying firm.
Leaves purple P. rubescens.
Leaves bronzed, finely wrinkled, obtuse-acuminate.
P. aster ophylla,
Leaves scarcely purpling or bronzed.
Leaves revolute and hard P. sderophylla.
Leaves scarcely revolute.
Leaves 3-nerved P. flabilis.
Leaves 5-nerved P. ficta.
Plants more or less puberulent.
Leaves lance-ovate P. flavamenta.
Leaves broadly lanceolate P. glareosa.
Leaves narrowly lanceolate P. tetrica.
Leaves obovate to subspatulate P. inaequali folia.
Plants crisp-pubescent.
FLORA OF PERU 15
Leaves rhombic-spa tulate P. Mathewsiana.
Leaves elliptic P. praeruptorum.
Stem subvillous P. subvillicaulis.
J. Leaves 1-2.5 cm. long.
Leaves opposite.
Plants glabrous.
Leaves not papillate P. Herrerae.
Leaves minutely papillate P. gucayana.
Plants hairy P. villicaulis.
Leaves commonly whorled.
K. Leaves widened upwards.
Leaves uniform, glabrous.
Spikes short, apiculate P. obcordata.
Spikes not apiculate.
Spikes short P. quadricoma.
Spikes 25 mm. long P. estrellana.
Spikes 50 mm. long P. san-buenaventurana.
Spikes 60-80 mm. long.
Leaves coriaceous, revolute P. palcana.
Leaves scarcely revolute, granular, emarginulate.
P. pillahuatana .
Spikes over 100 mm. long.
Leaves 12 cm. long P. santa-cruzana.
Leaves becoming twice as long P. Pakipski.
Leaves partly subspatulate.
Plants glabrous.
Petiole scarcely 2 mm. long; leaves emarginate.
P. subdichotoma.
Petiole longer, winged P. huallagana.
Plants velvety P. anisophylla.
Plants hirsute P. Klotzschiana.
K. Leaves not widened upwards.
Leaves lanceolate.
Leaves not rhombic.
Plants tomentulose P. variifolia.
Plants somewhat puberulent.
16 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Leaves acute P. subcalvescens.
Leaves obtuse P. pongoana.
Leaves rhombic P. rhombea.
Leaves elliptic.
Leaves subrhombic P. puberulipes.
Leaves not rhombic.
Plants glabrous or slightly papillate.
Leaves 1-1.5 cm. long.
Leaves plicate P. cushiana.
Leaves flat : ...... P. rhodophylla.
Leaves 1.5-2.5 cm. long.
Spikes solitary . P. concam.
Spikes numerous P. congesta.
Plants puberulent.
Leaves scarcely 1 cm. wide P. huanucoana.
Leaves 1.5 cm. wide P. viracochana.
Plants soft-hairy P. calvescens.
Plants stiff-hirsute P. crinigera.
Plants villous.
Leaves acute P. congestispica.
Leaves obtuse P. caniana.
Leaves lance-elliptic P. guayrapurana.
Leaves lance-oblong P. lanosa.
Leaves subrhombic-ovate P. Smithii.
J. Leaves scarcely over 1 cm. long, narrow if slightly longer.
Leaves opposite, orbicular P. cyclophylla.
Leaves mostly whorled.
L. Leaves round-elliptic or subovate-elliptic.
Leaves glabrous.
Spikes 60 mm. long P. Ppucu-ppucu.
Spikes twice as long.
Leaves not waxen P. marcapatana.
Leaves drying waxen-translucent P. career-
Leaves pubescent.
Stem exfoliating, glabrous P. scabiosa.
Stem transiently hirtellous P. Diehliana.
Stem crisp-hairy P. roseo.
FLORA OF PERU 17
Stem yellow-tomentulose P. chrysotricha.
L. Leaves round, at most slightly puberulent.
Peduncle 3-4 cm. long P. trichopus.
Peduncle 1-2 cm. long; leaves rounded at the end.
Spikes 10 mm. long, 2 mm. thick P. subandina.
Spikes 10-15 mm. long, 1 mm. thick P. andicola.
Peduncle 1 cm. long.
Leaves round at the end P. filiformis.
Leaves subtruncate P. subandina.
Peduncle very short P. Claudii.
L. Leaves obovate.
Plants glabrous P. muscigaudens.
Plants somewhat puberulent P. pseudo-galapagensis.
L. Leaves subelongate-obovate.
Leaves scarcely rhombic or rugose, at most slightly puberulent.
Leaves rounded at the apex.
Leaves 3 or 4 at a node.
Leaves obscurely pinnate-nerved, acute at base.
P. dendroides.
Leaves 1-nerved P. Pearcei.
Leaves about 5 at a node, 1-nerved P. Gilberti.
Leaves truncate-emarginulate P. bracteispica.
Leaves subrhombic, rugulose P. rugatifolia.
L. Leaves elliptic.
Leaves subrhombic.
Spikes short; somewhat pubescent.
Pubescence subvillous P. huantana.
Pubescence short.
Leaves little longer than broad, obtuse-acuminate.
P. nigro-oculata.
Leaves more elongate.
Peduncle short P. rhombeo-elliptica.
Peduncle 2 cm. long P. quadratifolia.
Spikes subelongate.
Plants repent-assurgent P. quaerata.
Plants erect.
18 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Plants at most papillate P. milmfolia.
Plants somewhat crisp-hirtellous P. quaesita.
Leaves scarcely subrhombic.
Plants glabrous.
Stem very thick at the base; leaves elliptic.
P. crassulaecaulis.
Stem not greatly thickened.
Leaves round-elliptic, subobtuse at the base.
P. quispicanchiana.
Leaves twice as long as broad P. arctebaccata.
Plants papillate.
Papillae not bristly P. micromamillata.
Papillae bristly P. naevifolia.
Plants puberulent.
Leaves short-elliptic P. chillonensis.
Leaves more elongated.
Leaves obscurely pinnately nerved.
Stem very slender P. brachyiula.
Stem 2-3 mm. thick P. ceapanana.
Leaves scarcely more than 1-nerved.
Stem 5 mm. thick below P. distractiflora.
Stem slender P. sangabanensis.
L. Leaves oblong or linear.
Upper leaves elongate.
Lower leaves barely 5 mm. long P. atocongona.
Leaves larger.
Spikes and peduncles short.
Stem very leafy P. foliata.
Internodes long P. seposila.
Spikes 60-150 mm. long P. longispica.
Upper leaves scarcely elongated.
Leaves subconduplicate P. trullaefolia.
Leaves not deeply concave.
Stem slender.
Peduncle 5 mm. long P. dendromorphis.
Peduncle twice as long P. agapatensis.
Stem 4-5 mm. thick P. arcuatispica.
FLORA OF PERU 19
Peperomia acuminata R. & P. Fl. 1: 32. pi. 51. 1798. Piper
acuminatum Pers. Syn. 1: 35. 1805, non P. acuminatum L. 1753.
Piper nemorosum Vahl, Enum. 1: 341. 1805. Peperomia Ruiziana
C. DC. Candollea 1: 337. 1923.
A rather large, glabrous, ascending herb; leaves alternate, oblong-
obovate, acuminate, acute-based, 6-9 cm. long, 2.5-3.5 cm. wide,
pinnately nerved; petiole 1 cm. long; spikes terminal, 4 mm. thick,
100 mm. long or more; peduncle short.
Huanuco: Muna, Pillao, and Macosa, Ruiz & Pavon, type.
San Martin: San Roque, Williams 7428.
Peperomia acutifolia C. DC. Journ. Bot. 4: 142. 1866.
A glabrous, moderate-sized herb; leaves alternate, lanceolate,
acutely protracted, long-cuneate, 15 cm. long, 3-4 cm. wide, pin-
nately nerved; petiole 3 cm. long or more; spikes 2 or several in a
terminal panicle, filiform, 80 mm. long; peduncle 10 mm. long;
bracts round-peltate; berries narrowly oblong, obliquely sharp-
scutulate, with central stigma. Neg. 8311.
San Martin: Tarapoto, Spruce 4094, type. San Roque, 1,350-
1,500 meters, Williams 7428. "Afasi quihua."
Peperomia aerea Trelease, sp. nov.
Herba gracilis rhizomatosa repens glabra, ramis brevibus ad-
scendentibus, caule 1 mm. crasso; folia alterna lanceolata sub-
acuminata basi acutiuscula 12-30 mm. longa 6-12 mm. lata coriacea
opaca 3-nervia vel obscure 5-nervia anguste revoluta, subtus con-
coloria granulosa, petiolo 3 mm. longo; spicae terminales 50 mm.
longae 1 mm. crassae, floribus densis vel subzonulatis, pedunculo
10 mm. longo; bracteae rotundo-peltatae.
Junin: Chanchamayo Valley, 1,800 meters, Carlos Schunke 494
(herb. Field Mus. No. 571,552, type), 187.
Peperomia agapatensis C. DC. in DC. Prodr. 16, pt. 1:
455. 1869.
A puberulent, suffrutescent, erect, branching herb; leaves
commonly 4 at a node, oblong or somewhat enlarged upward,
obtuse, subacute at the base, 8-15 mm. long, 3-4 mm. wide, ob-
scurely pinnate-nerved, yellowish beneath; petiole scarcely 1 mm.
long; spikes terminal, 50 mm. long, 1 mm. thick; peduncle 10-15
mm. long; bracts round-peltate; berries round, pointed; stigma
apical. Neg. 8304.
20 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Type from Agapata, Lechler 1934. Junin: La Merced, Schunke
Hacienda, Killip & Smith 24576.
Peperomia ainana Trelease, sp. nov.
Herba modice alta simplex assurgens glabra, caule 2-4 mm.
crasso; folia alterna ovata acuminata, basi rotundata vel sub-
truncata atque ad petiolum acute contracta, 8-10 cm. longa 5-7
cm. lata, multiplinervia nervis basilaribus utroque latere 2-3, majori-
bus ut costa 1-2-ramosis, statu sicco tenuia viridia, petiolo 3-8
cm. longo gracili; spicae sympodiales 40 mm. longae 2 mm. crassae,
pedunculo aequilongo; bracteae rotundo-peltatae; ovarium globosum
mucronatum, stigmate obliquo.
Ayacucho: Aina, near Rio Apurimac, 750-1,000 meters, Killip
& Smith 22766 (type in U. S. Nat. Herb.; dupl. in herb. Field Mus.).
Junin: La Merced, Killip & Smith 23725.
A terrestrial plant; flower spikes white.
Peperomia alata R. & P. Fl. 1: 31. pi. 48. 1798. Piper alatum
Vahl, Enum. 1:342. 1805.
A moderate-sized, glabrous, assurgent herb; stem alately angled
below the petiole margins; leaves alternate, lance-elliptic, sub-
acuminate, 4-7 cm. long, 2-2.5 cm. wide, acute-based, 5-nerved
with the inner nerves somewhat confluent; petiole 5-10 mm. long,
clasping-decurrent; spikes terminal and from the upper axils, 70
mm. long, 2 mm. thick; peduncle 10 mm. long; bracts round-peltate.
Huanuco: Posuso, Ruiz & Pavon, type; at 600 meters, on mossy
rocks, 4687, 4680. Ayacucho: Aina, Killip & Smith 22830, 22769.
Junin: Balsapuerto, Killip & Smith 28428. San Nicolas to
Azupizu, Killip & Smith 24088. Puerto Bermudez, Killip &
Smith 26465.
Peperomia alatiscapa Trelease, sp. nov.
Planta epiphytica mediocris subsimplex glabra; folia alterna vel
suprema ternata elliptico-subobovata obtuse acuminata basi acuta
2.5-4 cm. longa 1.5-2.5 cm. lata 3-5-nervia, costa obscure ramosa,
in sicco opaco-olivacea et opaca, statu vivo obscure rosea, petiolo
10 mm. longo in costulas caulinas decurrente; spicae terminales 70
mm. longae 2 mm. crassae densiflorae, pedunculo 10 mm. longo;
bracteae rotundo-peltatae.
Junin: Enefias, Pichis Trail, 1,600-1,900 meters, Killip & Smith
25623 (type in U. S. Nat. Herb.).
FLORA OF PERU 21
Peperomia alismaefolia Presl, Epim. Bot. 222. 1845.
A rhizomatous herb; stem short, toothed from fallen leaf bases;
leaves few, alternate, elliptic-ovate, obtuse at both ends, 4-6 cm.
long, 2.5-4 cm. wide, 5-nerved, pubescent above and ciliate; petiole
4-5 cm. long, glabrous; spikes from between the leaves, 20 mm.
long, 2 mm. thick; bracts round-peltate, ciliate; ovary subglobose,
umbilicate; stigma apical.
Type from mountains of Peru.
Peperomia andicola Dahlst. Svensk. Vet. Akad. Handl. 33,
pt. 2: 182. pi. 7. 1900.
A delicate, repent, short-branched herb, at least transiently
puberulent; stem filiform; leaves commonly 3 or 4 at a node, round-
obovate, subtruncate, cuneate, 5 mm. wide, 3-nerved; petiole very
short; spikes terminal, 10-15 mm. long, 1 mm. thick; peduncle of
the same length; bracts round-peltate; berries ovoid, pointed, with
pseudo-cupule; stigma apical.
Junin: Tarma, Philippi, type. Loreto: Yapas, 1,350-1,600
meters, Killip & Smith 25462(1).
Peperomia andinacea C. DC. Journ. Bot. 4: 134. 1866.
A delicate, repent herb, glabrous except for a few hairs on the
upper leaf surface; leaves alternate, orbicular, 2-3 mm. in diameter,
obscurely 3-nerved; petiole 1-2 mm. long; spikes terminal, 10 mm.
long, 1 mm. thick; peduncle 2 mm. long.
Puno: Sangaban, Lechler 2410, type.
Peperomia anisophylla C. DC. Bot. Jahrb. 40: 266. 1908.
An erect, succulent herb; stem 2-3 mm. thick, with short inter-
nodes; leaves about 3 at a node, reflexed, obovate, subtruncate or
very obtuse, subcuneate at the base, 10-17 mm. long, 5-10 mm.
wide, velvety on both sides, 5-nerved; petiole very short; spikes
terminal and from the upper axils, 45 mm. long, 1 mm. thick;
peduncle scarcely 5 mm. long; bracts round-peltate; ovary ovoid;
stigma subapical.
Huanuco: Between Matucana and Arinia, 2,300 meters, Weber-
bauer 72, type.
Peperomia antoniana Trelease, sp. nov.
Herba parvula subrepens glabra terrestris; folia alterna rotundo-
ovata obtusissima basi cordata 15-20 mm. longa et lata 5-nervia
in sicco viridia et pellucida, petiolo 5-20 mm. longo; spicae in sicco
22 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
filiformes, 10-15 mm. longae 3 mm. crassae; bracteae rotundo-
peltatae.
Loreto: San Antonio, Rio Itaya, 110 meters, Killip & Smith
29498 (type in U. S. Nat. Herb.).
Peperomia apurimacana Trelease, sp. nov.
Plata epiphytica alta succulenta ramosa, caule 2-4 mm. crasso
crispo-floccoso ; folia opposita subrhombeo-elliptica utrinque acuta,
foliis inferioribus reductis et obovato-truncatis, 3 cm. longa et 1
cm. lata vel usque ad 5 cm. longa atque 3 cm. lata, trinervia,
subpersistente crispo-pubescentia, in sicco tenuia, petiolo 5-10 mm.
vel ad 15 mm. longo crispo-pubescente; spicae terminales et axillares
filiformes 30-40 mm. longae sublaxiflorae floribus pseudopedicellis
brevibus stipatis, pedunculo 10-15 mm. longo translucente minute
pubescente; bracteae rotundo-peltatae.
Ayacucho: Carrapa, 1,500 meters, Killip & Smith 22355.
Peperomia arborigaudens Trelease, sp. nov.
Planta epiphytica repenti-pendula, caule gracili elongato breviter
villoso; folia alterna ovata acuminata brevissime et obscure peltata,
basi rotundata vel subtruncata, 4-5.5 cm. longa 3-4 cm. lata,
5-7-nervia, statu juvenili subtus tomentulosa serius sparse pilosa
et glanduloso-granulosa, in sicco tenuia atque plus minusve rosea,
petiolo 3-5 mm. longo laxe villoso; spicae terminales graciles 90
mm. longae, pedunculo filiformi 2 cm. longo sub villoso; bracteae
minutae rotundo-peltatae.
Loreto: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25509 (type in U. S. Nat. Herb.).
Peperomia arboriseda C. DC. Bot. Jahrb. 40: 261. 1908.
A glabrous, spreading epiphyte; stem slender, sublignescent;
leaves alternate, lanceolate or the lower suborbicular, acute, round-
based, 4 cm. long, 2 cm. wide, 7-nerved, the inner nerves sub-
confluent below, ciliate; petiole 5 mm., or less than 15 mm., long;
spikes 3 on a slender, terminal, 2-bracted stalk, slender; peduncle
5 mm. long; bracts round-peltate; berries oblong, obliquely scutulate;
stigma central.
Junin: La Merced, Chanchamayo Valley, in forest, 1,000 meters,
Weberbauer 1917, type.
Peperomia arboriseda var. moyobambana C. DC. Bot. Jahrb.
40: 261. 1908.
FLORA OF PERU 23
Internodes 5 cm. long; leaves 6 cm. long, 2.5 cm. wide, with
densely ciliate nerves; scales 15 mm. long subtending the spikes.
San Martin: Moyobamba, Weberbauer 1+691, type.
Peperomia arctebaccata Trelease, sp. nov.
Herba modice parva arete radicans caespitosa dichotoma glabra,
caule 2 mm. crasso firmo plus minus ve furfuraceo-exfoliante; folia
opposita vel ternata vel quaternata elliptica obtuse submucronata
vix 10 mm. longa et 5 mm. lata opaca crassissima revoluta, subtus
in sicco fusca, nervis non manifestis, petiolo 1-2 mm. longo; spicae
terminales 30-50 mm. longae 2 mm. crassae densiflorae, floribus in
alveolis oblongis profundis insertis, pedunculo crasso 5-10 mm.
longo; bracteae minutae rotundo-peltatae; baccae numerosissimae
pseudocupula brevi instructae, stigmate terminali, stylo brevi.
Junin: Chanchamayo Valley, 1,000 meters, Carlos Schunke 471
(herb. Field Mus. No. 571,529, type), A89, A86, 386, 372.
Peperomia arcuatispica Trelease, sp. nov.
Herba majuscula glabra stolonifera, caule 5 mm. crasso; folia
alterna oblanceolata acuta basi cuneata 16-18 cm. longa 6 cm. lata
ubique pinnatinervia, in sicco tenuia viridia, petiolo 1 cm. longo
lamina decurrente alato; spicae paniculatae, panicula foliis vix
longiore, ramis circa 3 basi spicis 1 vel 2 et 2-4 cm. longis atque
curvis basilaribus onustis apice spicas plures gerentibus, spicis 30
mm. longis 1 mm. crassis, pedunculo lere nullo; bracteae rotundo-
peltatae.
Junin: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
25992 (type in U. S. Nat. Herb.).
Peperomia areolata Trelease, sp. nov.
Herba glabra rhizomatosa, caulibus brevibus crassis paucifoliis;
folia reniformi-ovata vel rotundo-ovata apice rotundata vel sub-
acuta vel submucronata, basi breviter aperte cordata, 6-8 cm.
longa, 7-8 cm. lata 7-nervia in sicco venulose pallido-areolata,
petiolo 8-12 cm. longo; inflorescentia non visa.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28660
(type in U. S. Nat. Herb.). Huallaga, Weberbauer 6807.
Peperomia artatiflora Trelease, sp. nov.
Herba modice alta parvifolia succulenta, caule velutino inferne
ad 4-5 mm. crasso in sicco lucido-stramineo ; folia 3-4-nata linearia
ad subobovato-elliptica obtusa vel subacuta basi acutiuscula 6-15
24 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
mm. longa 2-3 mm. lata, utrinque papilloso-subpuberula, 1-nervia,
petiolo 1 mm. longo; spicae terminales et ex axillis superioribus,
40 mm. longae 1 mm. crassae densiflorae, pedunculo 10 mm. longo
glabrato; bracteae rotundo-peltatae brunneae.
Junin: Palca, Stevens 47 (type).
Peperomia asterophylla Trelease, sp. nov.
Herba mediocris stolonifera suberecta caespitosa, caule 2-3 mm.
crasso ad nodos velutino; folia ad nodum 3-5 elliptica obtuse acumi-
nata basi acuta 2-3 cm. longa 1.5-2.5 cm. lata obscure trinervia,
in sicco opaca et minute rugosa glabra vel fere glabra, petiolo 3-4 mm.
longo brunneo- velutino; spicae axillares et terminales 40-80 mm.
longae 2 mm. crassae, pedunculo 10-15 mm. longo glabrato; bracteae
rotundo-peltatae.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28909 (type
in U. S. Nat. Herb.). Sapote Yacu, Santa Rosa, 200 meters,
Williams 4901 . San Antonio, Alto Rio Itaya, 145 meters, Williams
3479.
Peperomia atocongona Trelease, sp. nov.
Herba modice parvula suberecta caespitosa, caule 1-3 mm.
crasso plus minusve ramoso minute puberulo; folia ad nodum 3-6
elliptica vel subobovata obtusa basi acuta 4-6 mm. longa 2-3 mm.
lata, superioribus ad 10-20 mm. longis et 5 mm. latis, subtus obscure
pinnatinervia atque granulosa, petiolo vix 1 mm. longo; spicae
terminales et ex axillis superioribus 15 mm. longae 1 mm. crassae
densiflorae, pedunculo 5 mm. longo; bracteae rotundo-peltatae.
Lima: Atocongo, 250-500 meters, F . W. Pennell 14752 (type in
herb. Acad. Nat. Sci. Phila.).
Peperomia balsapuertana Trelease, sp. nov.
Herba epiphytica majuscula fere acaulescens glabra; folia alterna
elliptico-oblanceolata obtusa ad acute subacuminata basi cuneata
17-25 cm. longa 7-10 cm. lata ubique pinnatinervia, in sicco tenuia,
petiolo 5-6 cm. longo; inflorescentia non visa.
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28632
(type in U. S. Nat. Herb.).
Peperomia basellaefolia HBK. Nov. Gen. & Sp. 1: 66. 1815.
Piper basellaefolium Poir. Encycl. Suppl. 4: 468. 1816.
A glabrous, assurgent, succulent herb; stem 3 mm. thick; leaves
alternate, subelliptic, acuminate, acute at the base, 3-5 cm. long,
FLORA OF PERU 25
3-4 cm. wide, ciliate, pinnately few-nerved; petiole 5-10 mm. long;
spikes terminal, when young 25 mm. long, 2 mm. thick, the peduncle
1 cm. long.
Type from Peru, without definite locality, Mathews 1838.
Ecuador.
Peperomia bermudezana Trelease, sp. nov.
Herba glabra repenti-assurgens arboricola, caule 2-3 mm. crasso;
folia alterna elliptico-suboblonga acuminata basi acuta 7-9 cm.
longa 3-4 cm. lata trinervia vel obscure 5-nervia costa obscure
ramosa, in sicco tenuia opaca, subtus plus minusve pallida, petiolo
gracili 1-2 cm. longo; spicae sympodiales interdum in ramis brevibus
2-bracteatis insertae, filiformes, 70 mm. longae, pedunculo 5-10
mm. longo; bracteae rotundo-peltatae.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26451
(type in U. S. Nat. Herb.).
Peperomia binispica Trelease, sp. nov.
Herba mediocris subsimplex glabra, caule 2 vel ad basin 5 mm.
crasso; folia alterna lanceolata 15 cm. longa 4 cm. lata vel majora,
utrinque acuta, apice interdum obtusa, pinnatinervia, petiolo vix
10 mm. longo vel fere nullo; spicae axillares vulgo binae et pedunculos
graciles ad 10 cm. longos terminantes, 100-140 mm. longae 2 mm.
crassae densiflorae, pedunculo 15-25 mm. longo filiformi; bracteae
rotundo-peltatae; baccae rotundato-ovoideae oblique rostro crasso
terminatae, stigmate ad basin rostri inserto.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 358
(herb. Field Mus. No. 571,412, type), A84-
Peperomia blepharipus Trelease, sp. nov.
Planta modice alta epiphytica erecta subsimplex, petiolis crispo-
ciliatis et lineis decurrentibus exceptis glabra, caule 2-3 mm. crasso;
folia alterna lanceolata utrinque acuta 8-10 cm. longa 2-3.5 cm.
lata fere e basi 3-5-nervia costa superne obscure ramosa, in sicco
tenuia opaca, petiolo 5-10 vel ad 30 mm. longo amplexicauli decur-
rente; spicae terminales vel sympodiales 60 mm. longae 2 mm.
crassae, pedunculo 2 cm. longo; bracteae rotundo-peltatae.
Loreto: Yurimaguas, 135 meters, Killip & Smith 28030 (type
in U. S. Nat. Herb.).
Peperomia blepharipus var. binispica Trelease, var. nov.
Planta gracilior, spicis binis ramulum gracili-bracteatum termi-
nantibus 60 mm. longis 1 mm. crassis.
26 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Loreto: San Antonio, Alto Rio Itaya, 145 meters, Williams
3435 (type in herb. Field Mus., No. 618,141).
Peperomia blepharipus var. iquitosensis Trelease, var. nov.
Planta validior, caule 2-4 mm. crasso; folia elliptica vel sub-
ovata 7-9 cm. longa 3-5.5 cm. lata; spicae solitariae.
Loreto: Iquitos, 100 meters, Killip & Smith 27462.
Peperomia brachyiula Trelease, sp. nov.
Herba erecta modice parva dense caespitosa breviter ramosa,
caule gracili 1-2 mm. crasso papilloso-velutino; folia ad nodum
4-6 patentia elliptica vel rotundato-elliptica obtusa vel subretusa
basi plus minusve acuta parva, 6-9 mm. longa, 2-4 mm. lata, sup-
rema magis rotundata, modice tenuia, obscure pinnatinervia, primo
utrinque dense puberula, petiolo vix 2 mm. longo puberulo; spicae
ramos breves dense foliatos terminantes vix 35 mm. longae 1 mm.
crassae laxiflorae, pedunculo 5 mm. longo ; bracteae rotundo-peltatae ;
ovaria inter costulas anastomosantes immersa; baccae rotundato-
ovoideae oblique mucronulato-truncatae; stigma anterius.
Lima: Matucana, steep southern shale slope, in large clumps,
2,400 meters, Macbride 129 (herb. Field Mus. No. 516,662, type).
Peperomia bracteispica Trelease, sp. nov.
Herba parva repenti-assurgens glabra epiphytica, caule 1 mm.
crasso; folia vulgo ternata obovata truncato-emarginulata basi
cuneata 9-11 mm. longa 5-6 mm. lata obscurissime trinervia,
petiolo filiformi 3-4 mm. longo; spicae terminales vix 40 mm. longae
et 1 mm. crassae, floribus sublaxe dispositis, pedunculo 5 mm.
longo folio unico 15 mm. supra verticillos supremos inserto fulcrato;
bracteae rotundo-peltatae; rhachis pseudopedicellis onusta.
Huanuco: Huacachi, near Muna, 1,950 meters, 41 71 a (type in
herb. Field Mus. No. 551,173).
Peperomia caducifolia Trelease, sp. nov.
Herba alta repenti-assurgens, caulibus carnosissimis ramosis
puberulis, internodiis brevibus; folia alterna, foliis ramorum steri-
lium parvis et suborbicularibus, ceteris ellipticis obtuse acuminatis
basi subacutis 3 cm. longis 1.5-2 cm. latis, superioribus minoribus,
caducis, trinerviis, minute puberulis, petiolo 5-10 mm. longo glab-
rescente; spicae plures terminales et ex axillis nudis approximatis
brevem pseudopedunculum efformantibus nascentes 30 mm. longae,
floribus laxiuscule insertis, pedunculo 5 mm. longo; bracteae
rotundo-peltatae.
FLORA OF PERU 27
Ayacucho: Carrapa, 3,000 meters, Killip & Smith 22270 (type
in U. S. Nat. Herb.).
Peperomia calvescens Trelease, sp. nov.
Herba minuscula stolonifera succulenta, caulibus plus minusve
caespitosis erectis subgracilibus 2 mm. crassis primum sparse mol-
liter pilosis cito epidermi furfuraceo-exfoliante glabrescentibus vel
ad nodos persistente pilosis; folia 3-4-nata lanceolato-elliptica
utrinque acuta trinervia parva 16 mm. longa 8 mm. lata supra ad
nervos plus minusve persistente crispo-pubescentia, subtus glabra
et epidermi exfoliato albida, petiolo brevi 2 mm, longo glabro;
spicae terminales 60 mm. longae 1 mm. crassae, floribus laxiuscule
insertis, pedunculo filiformi 2 mm. longo glabro; bracteae rotundo-
peltatae; ovarium impressum ovoideum, stigmate subapicali, rhachi
pseudopedicellis brevibus onusta.
Huanuco: Cueva Grande near Posuso, 1,050 meters, sunny clay
banks, Macbride 4800 (herb. Field Mus. No. 535,834, type).
Peperomia caniana Trelease, sp. nov.
Herba modice parva repenti-assurgens, caule 2-4 mm. crasso
crispo-villoso; folia opposita orbicularia, superiora late ovata,
utrinque obtusa vel rotundata, inferiora submucronata, minuscula,
12-25 mm. longa 12-20 mm. lata, trinervia arctiuscule adpresso-
pilosa vel supra glabrescentia, petiolo vix 5 mm. longo crispo-piloso ;
spicae terminales subumbellatae 60 mm. longae 1 mm. crassae laxi-
florae, pedunculo 1.5-2 cm. longo laxe piloso; bracteae rotundo-
peltatae; baccae in rhachi immersae globulosae ovoideae obtusae,
stigmate subapicali.
Huanuco: Cani, near Mito, 2,550 meters, Macbride 3415 (herb.
Field Mus. No. 534,486, type).
Peperomia carpapatana Trelease, sp. nov.
Herba majuscula ramosa, caule 2-5 mm. crasso praesertim
superne dense luteo-intertexto-piloso; folia alterna rotundo-ovata
apice rotundata et subabrupte obtuso-acuminata, basi rotundata
vel abrupte subacuta, 2-4 cm. longa 2-3.5 cm. lata, obscure 3-5-
nervia subtus adpresso-pilosa et concolori-granulosa in sicco opaca,
petiolo 5-10 mm. longo adpresso-piloso ; spicae ad axillas aggregatae
graciles 50-150 mm. longae, floribus sublaxe insertis, pedunculo
gracili 10-15 mm. longo plus minusve adpresso-piloso; bracteae
rotundo-peltatae; baccae parvae globosae apice mucronatae, stigmate
subapicali.
28 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Junin: Carpapata, 2,700-3,200 meters, Killip & Smith 24438
(U. S. Nat. Herb., type), 24393.
Peperomia carrapana Trelease, sp. nov.
Planta parva epiphytica rhizomatoso-adscendens glabra, caule
1-2 mm. crasso; folia alterna lanceolata vel lanceolato-oblonga sub-
obtusa vel attenuata, basi subacuta, 2-4 cm. longa, 1-1.5 cm. lata,
obscure trinervia, in sicco opaca tenuia aliquanto crispo-revoluta,
petiolo 5 mm. longo; spicae terminales 60 mm. longae 2 mm. crassae,
floribus laxiuscule insertis, pedunculo 5 mm. longo; bracteae rotundo-
peltatae.
Ayacucho; Carrapa, 1,500 meters, Killip & Smith 23207 (U. S.
Nat. Herb., type).
Peperomia casapiana C. DC. Journ. Bot. 4: 146. 1866.
A pilose herb; leaves 3 at a node, ovate, acuminate, rounded at
the base, 5 cm. long, 3 cm. wide, 5-nerved, drying pellucid; petiole
8 mm. long; spikes subumbellately verticillate from the axils of the
reduced upper leaves, 50 mm. long, 1 mm. thick; peduncle 15 mm.
long; bracts round-peltate; berries ovoid, mucronate; stigmas sub-
apical.
Huanuco: Casapi, Mathews 1689, type.
Peperomia ceapanana Trelease, sp. nov.
Herba erecta plus minusve caespitosa breviter ramosa, caule
2-3 mm. crasso dense papilloso-puberulo; folia ad nodum circa 4
adscendentia elliptica vel elliptico-obovata obtusa basi acuta parva
10 mm. longa 4 mm. lata subtenuia obscure pinnatinervia glabra,
subtus glanduloso-granulosa, petiolo 1 mm. longo papilloso-puberulo;
inflorescentia non visa.
Cuzco: Hacienda Ceapana, Paucartambo Valley, 3,450 meters,
F. L. Herrera 1468 (U. S. Nat. Herb., type).
Peperomia cerea Trelease, sp. nov.
Herba modice alta succulenta glabra e rupibus pendula, caule
valido 5 mm. crasso, superne graciliore; folia opposita vel superne ad
nodum 3-5 rotundato-elliptica utrinque rotundata circa 15 mm. longa
et 10 mm. lata, superiora magis oblonga et minora, vix 10 mm.
longa et 7 mm. lata, margine tenui plus minusve revoluto, in sicco
cerino-translucentia, 1-nervia, petiolo brevi 1-3 mm. longo; spicae
terminales 100 mm. longae et ultra 3-4 mm. crassae obscure rubrae,
floribus laxiuscule insertis, pedunculo crasso 15 mm. longo; bracteae
FLORA OF PERU 29
rotundo-peltatae ; ovaria in rhachi immersa acuminata, stigmate
subapicali.
Huanuco: Mito, 2,700 meters, Macbride 3330 (type in herb.
Field Mus. No. 534,400).
Peperomia chachapoyasensis C. DC. in DC. Prodr. 16, pt.
1: 449. 1869. P. umbellata Miq. Lond. Journ. Bot. 4: 427. 1845,
non Kunth.
A delicate, suberect, glabrous herb, the stem almost filiform;
leaves alternately clustered at the apex, round-ovate, very obtuse,
rounded at the base or obscurely cordulate, 2-2.5 cm. long, 2 cm.
wide, 3- or somewhat 5-nerved; petiole 5 mm. long; spikes umbellate
at the ends of slender, terminal or axillary, 2-bracted stalks 3-4 cm.
long, scarcely 15 mm. long and 1 mm. thick; peduncle 5 mm. long;
bracts round-peltate; berries round-ovoid, mucronate; stigma sub-
apical. Neg. 8394.
Amazonas: Seringal Chachapoyas, Mathews 3230, type; 1862.
Peperomia chalhuapuquiana Trelease, sp. nov.
Herba modice parvula erecta divaricate multiramosa, caule 2-3
mm. crasso crispo-pubescente; folia alterna ovata vel elliptico-ovata
obtuse subacuminata basi vulgo rotundata 2-4 cm. longa 1.5-2.5
cm. lata ciliata, in sicco subcoriacea, subtus pallidiora minute nigro-
glandulosa, petiolo 2-3 vel ad 10 mm. longo glabrato; spicae termin-
ales et ex axillis superioribus 100 mm. longae 2 mm. crassae, floribus
laxiuscule vel verticillatim insertis, pedunculo gracili vix 10 mm.
longo glabro; bracteae rotundo-peltatae; baccae subglobosae mucro-
natae in sicco nigrae, stigmate obliquo.
Junin: Hacienda Chalhuapuquio, Stevens 215, type.
Peperomia chanchamayana Trelease, sp. nov.
Herba epiphytica modice parva rhizomatosa simplex glabra,
caule 2 mm. crasso; folia alterna ovata vel lanceolata utrinque
acuta 2.5-3.5 cm. longa 1-1.5 cm. lata trinervia vel obscure 5-nervia
subtus pallidiora coriacea aliquanto crispo-revoluta, petiolo 1 cm.
longo amplexicauli ; spicae terminales 60-90 mm. longae 1 mm.
crassae densiflorae, pseudopedicellis brevissimis, pedunculo 5 mm.
longo; bracteae rotundo-peltatae; baccae subglobosae oblique apicu-
latae, stigmate subapicali.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 371
(herb. Field Mus. No. 571,425, type); at 1,500 meters, Schunke
1554.
30 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia chartacea Trelease, sp. nov.
Herba caespitosa epiphytica modice alta glabra, caule 2-3 mm.
crasso nodoso; folia alterna elliptico-subobovata acuta vel submu-
cronata basi cuneata 6-11 cm. longa 3-6 cm. lata essentialiter pal-
matim 7-nervia vel nervis minoribus plus minusve circa basin
confluentibus, in sicco firmo-papyracea, petiolo 5-15 mm. longo;
spicae solitariae et terminates 20 mm. longae 2 mm. crassae,
pedunculo gracili aequilongo; bracteae rotundo-peltatae.
Junin: Puerto Bermudez, 375 meters, Killip & Smith 26636
(type in U. S. Nat. Herb.).
Peperomia chillonensis Trelease, sp. nov.
Herba modice parva subsimplex(?) griseo-puberula ; folia ad
nodum 8 et ultra elliptica vel subobovata obtusa basi subacuta opaca
obscure 3-5-nervia subrevoluta, 10 mm. longa 5 mm. lata, petiolo
1 mm. longo; spicae axillares vel ramulos brevissimos terminantes 20
mm. longae 1 mm. crassae, floribus laxiuscule insertis; bracteae ro-
tundo-peltatae; ovarium ovoideum, stigmate subapicali.
Lima: Along Rio Chillon, above Obrajillo, 2,800-3,200 meters,
open rocky banks, F. W. Pennell 14413 (herb. Field Mus. No.
558,460, type).
"A fleshy herb with greenish yellow spikes."
Peperomia chrysotricha Miq. Syst. 163. 1843.
A rather small, erect, succulent herb, yellow-puberulent or tomen-
tulose; leaves commonly 4 at a node, subreflexed, round-obovate or
elliptic, rounded at both ends, 6-10 mm. long, 3-6 mm. wide, cori-
aceous, 1- or obscurely 3-nerved; petiole very short; spikes terminal
and from the upper axils, 20-40 mm. long; peduncle 5-10 mm. long;
bracts round-peltate, pale-margined; berries globose: stigma apical.
Type from Peru, Dombey 933.
Peperomia ciliatifolia Trelease, sp. nov.
Herba pendula arboricola foliis sparse pilosis et rigido-ciliatis ex-
ceptis glabra, caule gracili 2 mm. crasso; folia alterna elliptica acute
acuminata basi rotundata vel subacuta 11 cm. longa 5 cm. lata infra
medium pinnatim paucinervia in sicco tenuia translucentia, petiolo
vix 10 mm. longo; spicae sympdiales geminatae graciles 150 mm.
longae, pedunculo vix 5 mm. longo; bracteae rotundo-peltatae;
baccae oblongae vel subconicae scutulo acuto stigma gerente oblique
truncatae.
FLORA OF PERU 31
Loreto: Balsapuerto, 150-350 meters, Killip & Smith 28684-
(type in U. S. Nat. Herb.). Yurimaguas, Killip & Smith 27982.
Mishuyacu, near Iquitos, 100 meters, Klug 58.
Peperomia ciliatifolia var. eciliatifolia Trelease, var. nov.
Planta glabra vel sparse obscure ciliata, aliter ut in forma typica.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28723 (type in
U. S. Nat. Herb.).
Peperomia ciliatifolia var. iquitosana Trelease, var. nov.
Magis glabrescens, spicis filiformibus.
Loreto: Iquitos, Killip & Smith 27167 (type in U. S. Nat. Herb.).
Peperomia ciliatifolia var. santarosana Trelease, var. nov.
Folia minora, 6 cm. longa 3 cm. lata.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28988 (type in
U. S. Nat. Herb.).
Peperomia Claudii C. DC. Ann. Cons. Jard. Bot. Geneve 2:
285. 1898.
A small, cespitose, spreading epiphyte; stem filiform, puberulent;
leaves 3 at a node, round-obovate, acute-based, 6 mm. long, 5 mm.
wide, obscurely 3-nerved, glabrous; spikes terminal, 10 mm. long,
1 mm. thick; peduncle very short; bracts round-peltate; berries ovoid,
pointed; stigma apical.
Type collected in Peru by Gay.
Peperomia coloniae Trelease, sp. nov.
Herba suffruticosa repenti-assurgens transiente sparse molliter
pubescens; folia alternalanceolataplus minusve acuminatabasi obtusa
5-6 cm. longa 2.5-3 cm. lata coriacea opaca obscure pinnatinervia,
subtus flavescentia, petiolo 10 mm. longo; spicae terminales, solitariae
vel geminatae et pedunculos breves bibracteatos terminantes, breves.
Junin: Colonia Perene", 680 meters, Killip & Smith 25053 (type
in U. S. Nat. Herb.).
Peperomia concava R. & P. Fl. 1: 31. pi. 46. 1798. Piper
concavum Vahl, Enum. 1: 350. 1805.
A moderately small, cespitose, somewhat branched, succulent
herb; stem 2-3 mm. thick; leaves 2-4 at a node, elliptic, rounded at
the apex, acutely subsessile, 2.5 cm. long, 1.5 cm. wide, concave,
with 5 red nerves; spikes terminal, 60 mm. long, 3-4 mm. thick.
Huanuco : Chinchao and Cochero, Ruiz & Pavon, type.
32 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia confertispica Trelease, sp. nov.
Herba modice parva procumbenti-assurgens obscure viridis,
caule 2-3 mm. crasso, internodiis inferioribus longis sordide molliter
hirsutis; folia alterna, superne in caulibus erectis opposita, rotundata
vel late ovata, apice rotundata vel obtusa, basi rotundata, modice
parva, 1.5-3.5 cm. longa, 1.5-3.5 cm. lata, 3-5-nervia, subtus palli-
diora, utrinque sparse praesertim ad nervos adpresso-pilosa, petiolo
1-1.5 cm. longo dense piloso; spicae solitariae vel plures in axillis
superioribus, juveniles vix 15 mm. longae et 1 mm. crassae densi-
florae, pedunculo gracili vix 5 mm. longo piloso; bracteae rotundo-
peltatae majusculae flavae.
Huanuco: Mito, 2,700 meters, Macbride 1466 (herb. Field Mus.
No. 518,142, type). "Plant dark green, sprawling on the deep
humus floor of a stream thicket."
Peperomia confertispica var. erecta Trelease, var. nov.
Planta stolonifera erecta, caulibus crassioribus 5 mm. crassis in
sicco plus minusve quadrangularibus; folia elliptica vel obovata vix
3 cm. longa et 2 cm. lata, superiora 2 cm. longa 1.5 cm. lata, subtus
flavescentia, in sicco plus minusve supra rugosa vel granulosa,
petiolo vix 1 cm. longo; spicae juveniles vix 25 mm. longae, pedunculo
ad 5 mm. longo; bracteae primo flavescentes, brunnescentes.
Huanuco: Mito, 2,700 meters, Macbride 1915 (herb. Field Mus.
No. 518,410). "Stems and upper leaves beneath dark red. Bracts of
inflorescence pale yellowish at the tip. Leaves dark green, glossy
above, the lower pale beneath."
Peperomia congesta HBK. Nov. Gen. & Sp. 1: 68. pi. 14.
1815.
A rather large, erect, glabrous herb; stem 5 mm. thick below,
with short internodes; leaves 5 to 7 at a node, round-ovate, short-
acuminate, acute at the base, 1.5-2.5 cm. long, 1-2 cm wide, 1-nerved ;
petiole 5 mm. long; spikes numerous in the axils of congested whorls
at the end of the stem so as to appear subpaniculate, 15-20 mm.
long, 1 mm. thick; peduncle 5 mm. long.
Piura : Rio Chuchancanao to Huancabamba, Bonpland, type.
Peperomia congestispica Trelease, sp. nov.
Herba mediocris assurgens plus minusve ramosa in sicco nigres-
cens, caule 1-4 mm. crasso dense molliter f errugineo-hirsuto ; folia ad
nodum circa 5 elliptica vel subobovata subacuta basi acuta 12-20
mm. longa 6-12 mm. lata obscure trinervia breviter hirtella, supra
FLORA OF PERU 33
glabrescentia, petiolo 2-4 mm. longo f errugineo-hirtello ; spicae
terminates et in axillis superioribus aggregatae, juveniles 10 mm.
tan turn longae et 1 mm. crassae breviter pedunculatae; bracteae
rotundo-peltatae.
Junin: Huacapistana, Stevens 62, type.
Peperomia cordulilimba C. DC. Bot. Jahrb. 40: 261. 1908.
A small epiphyte; stem slender, velvety; leaves alternate, obovate,
emarginulate, acute-based, 7 mm. in diameter, 3-nerved, puberulent
beneath; petiole barely 5 mm. long, puberulent; spikes terminal,
filiform, 15 mm. long; peduncle very short; bracts round-peltate;
ovary obovoid; stigma oblique.
Puno: Near Tambo Frilluma, road from Sandia to Chunchus-
mayo, 1,000 meters, Weberbauer 1197, type.
Peperomia cotoneasterifolia Trelease, sp. nov.
Herba arboricola modice alta plus minusve ramosa, caule 2-3
mm. crasso primo dense adpresso-villoso; folia opposita elliptico-
subovata utrinque abrupte subacuta 2-2.5 cm. longa 1.5-2 cm. lata
3- vel obscure 5-nervia supra glabrata, subtus pallidiora tomentosa,
petiolo 5 mm. longo adpresso-piloso ; spicae axillares 60 mm. longae
2 mm. crassae, pedunculo 10 mm. longo; bracteae rotundo-peltatae;
baccae ovoideae acutiusculae, stigmate apicali.
Junin: Carpapata, 2,700-3,200 meters, Killip & Smith 24390
(type in U. S. Nat. Herb.).
Peperomia crassulaecaulis Trelease, sp. nov.
Planta glabra succulenta suffrutescens, caulibus 3-4 mm. crassis
erectis superne sparse ramosis sed e basi ad 2 cm. crassa numerosis
flavescentibus, internodiis brevibus; folia ad nodum 6 elliptica emar-
ginulata basi acuta 10 mm. longa 4-5 mm. lata in sicco dura subre-
voluta opaca obscure nervata, subtus minute granulosa, petiolo 2
mm. longo; spicae terminales 140 mm. longa 4 mm. crassae, floribus
laxiuscule insertis, pedunculo 15 mm. longo; bracteae rotundo-
peltatae; flores in alveolis immersae; ovarium ovoideum, stigmate
obliquo.
Junin: Carpapata, 2,400 meters, Killip & Smith 24355 (type in
U. S. Nat. Herb.; dupl. in herb. Field Mus.).
Peperomia crinigera Trelease, sp. nov.
Herba modice alta plus minusve ramosa assurgens, caule 2-3
mm. crasso rigide crispo-piloso ; folia ad nodum circa 5 elliptico-
34 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
subobovata obtusa basi subacuta 10-16 mm. longa 6-8 mm. lata
obscure trinervia crispo-hispida, petiolo 3 mm. longo crispo-pilosa ;
spicae terminales et ex axillis superioribus 60 mm. longae 1 mm.
crassae, floribus laxiuscule insertis, pedunculo 1-2 cm. longo rigide
crispo-piloso ; bracteae rotundo-peltatae.
Junin: Palca, Stevens 45, type. Carpapata, Killip & Smith
24449.
Peperomia crotalophora Trelease, sp. nov.
Herba modice alta procumbens glabra in sicco viridis, caule
subgracili 2-3 mm. crasso, internodiis elongatis; folia alterna rotun-
dato-ovata ad subtriangulari-ovata acuta vel interdum obtusa vel
breviter acuminata mediocria 5.5-6.5 cm. longa 4.5 cm. lata
multiplinervia, nervis basilaribus 4, alteris 2 e quarto inferiore
nascentibus, petiolo 3 cm. vel ad 10 cm. longo; spicae terrrinales
solitariae pedunculos unibracteatos 2 cm. longos terminantes, 50-70
mm. longae 1 mm. crassae, floribus arete verticillatis, verticillis
antherarum magnarum dense aggregatis valde in statu juvenili
conspicuis, pedunculo 15 mm. longo; bracteae rotundo-peltatae
obscurae in pseudoverticillo circa 12.
Junin: Hacienda Schunke, La Merced, 1,200 meters, Macbride
5644 (type in herb. Field Mus. No. 536,682), terrestrial, growing on
forest floor.
Peperomia cruciata Trelease, sp. nov.
Herba glabra rhizomatoso-assurgens; folia alterna obovato-sub-
spathulata obtusa basi longicuneata 6-11 cm. longa 3-4.5 cm. lata
pinnatinervia, in sicco subtenuia, petiolo 2-4 cm. longo; spicae ter-
minales 110 mm. longae 2 mm. crassae, pedunculo 8-9 cm. longo
1-bracteato; bracteae spicae rotundo-peltatae minutae; baccae
oblongae graciliter rostratae, stigmate anteriore.
Junin: Chanchamayo Valley, 800 meters, Carlos Schunke 343
(type in herb. Field Mus. No. 571,399).
Peperomia crusculibacca Trelease, sp. nov.
Herba modice alta assurgens, caule 2-4 mm. crasso crispo-
pubescente; folia alterna elliptica subacuminata basi acuta 4-6 cm.
longa 2-3 cm. lata 3- vel obscure 5-nervia glabra subtus nigro-
glandulosa, petiolo gracili 10-15 mm. longo glabrato; spicae
(terminales? et) axillares filiformes 100 mm. longae plus minusve
laxiflorae, pseudopedicellis gracilibus, pedunculo 10 mm. longo gla-
FLORA OF PERU 35
brato; bracteae rotundo-peltatae; baccae subglobosae oblique mucro-
natae stigmate subapicali.
Junin: Huacapistana, Stevens 90, type; Killip & Smith 24305.
Peperomia crystallina R. & P. Fl. 1: 30. 1798. Piper crys-
tallinum Vahl, Enum. 341. 1805.
A glabrous, branching, anise-scented succulent with 4-sided,
pellucid stem; leaves alternate, oblong, papillate above, impressed-
punctulate beneath; spikes slender, loosely flowered.
Lima: Lima, Ruiz & Pavon, type.
A very obscure species, scarcely comparable with anything known
except P. metallica.
Peperomia cuprea Trelease, sp. nov.
Herba mediocris repens, caule 2 mm. crasso obscure evanescente
plus minusve crispo-pubescente; folia alterna ovata subacuminata
basi rotundata 3.5-4.5 cm. longa 2-3.5 cm. lata, 3-5 mm. supra
basin peltata, opaca coriacea obscure multiplinervia anguste revoluta
subtus cuprea sericeo-ciliata, petiolo 1-2.5 cm. longo molliter evan-
escenti-piloso ; spicae axillares 55 mm. longae 3 mm. crasse arete
zonatiflorae, pedunculo 2 cm. longo glabro; bracteae parvae rotundo-
peltatae in pseudoverticillo circa 20.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 354
(herb. Field Mus. No. 571,408, type), 1500.
Peperomia cuprea var. cordulifolia Trelease, var. nov.
- Herba mediocris repens, caule 1 mm. crasso evanescentissime plus
minusve crispo-sericeo ; folia alterna ovata subacuminata breviter
cordata 3.5-5.5 cm. longa 2.5-3.5 cm. lata circa 5 mm. supra basin
peltata, opaca coriacea subtus brunnescentia obscure multiplinervia
anguste revoluta ciliata, petiolo 1-5 cm. longo sparse minute piloso;
spicae ramulos graciles 1-bracteatos terminantes statu juvenili
50 mm. longae 2 mm. crassae dense zonatiflorae, pedunculo 10-15
mm. supra folium reductum longo; bracteae spicae rotundo-
peltatae parvae in pseudoverticillo circa 20.
Junin: Chanchamayo Valley, 1,500 meters, Carlos Schunke
379 (herb. Field Mus. No. 571,433, type), 1466; Killip & Smith
24839. Enefias, Killip & Smith 25672.
Peperomia currucaeformis Trelease, sp. nov.
Herba terrestris repens glabra; folia late ovata vel deltoideo-ovata
obtusa basi truncato-cordata 5-6 cm. longa 4-6 cm. lata 7-nervia,
36 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
nervis intimis lateralibus con costa confluentibus, in sicco opaco-
olivacea granulosa, petiolo 2-3 cm. longo; spicae ad apicem caulis
sympodialis 4 cm. longi divergente geminatae et sursum curvatae
60-70 mm. longae 3 mm. crassae, pedunculo 15-20 mm. longo;
bracteae rotundo-peltatae; baccae ellipsoideae rostratae, stigmate
ad basin rostri inserto.
Junin: Schunke Hacienda, La Merced, 1,400-1,700 meters, Killip
& Smith 24542 (type in U. S. Nat. Herb.).
Peperomia curticaulis Trelease, sp. nov.
Herba glabra, caule brevi crasso paucifoliato ; folia alterna late
ovata obtusa, basi rotundata vel brevi ter cordata, 7-10 cm. longa
7-9 cm. lata 7-nervia, costa ut quoque nervis lateralibus ramosis,
petiolo 7 cm. longo; spicae terminales 120 mm. longae 3 mm. crassae,
floribus subdistantibus, pedunculo 1-2 cm. longo; bracteae rotundo-
peltatae; ovarium ovoideum submucronatum, stigmate obliquo.
Junin: Cahuapanas, Rio Pichis, 30 meters, Killip & Smith 26773
(type in U. S. Nat. Herb.).
Peperomia curtipes Trelease, sp. nov.
Herba subacaulescens glabra; folia alterna rotundata subacu-
minata basi sinu fere clauso cordata 8-13 cm. longa 8-12 cm. lata
palmatim 9-nervia vel costa interdum superne obscure ramosa, in
sicco fusca, petiolo 15-20 cm. longo; spicae 50 mm. longae 2 mm.
crassae vel ma j ores, pedunculo ut videtur elosgato; bracteae rotundo-
peltatae; ovarium globosum, stigmate subapicali.
Junin: San Nicolas, Pichis Trail, 1,100 meters, Killip & Smith
26019 (type in U. S. Nat. Herb.). East of Quimiri Bridge, 900
meters, Killip & Smith 30679.
Peperomia curtipes var. contracta, Trelease, var. nov.
Caulis foliatus brevissimus, foliis congestis, petiolo vix 15 cm.
longo; spicae 100 mm. longae 3 mm. crassae vel ma j ores, laxiflorae,
pedunculo 15 cm. longo; bracteae rotundo-peltatae; ovarium rotunda-
turn mucronatum, stigmate obliquo.
Junin: La Merced, Schunke Hacienda, 1,400-1,700 meters, Killip
& Smith 24726 (type in U. S. Nat. Herb.).
Peperomia cushiana Trelease, sp. nov.
Herba subparva caespitosa plus minusve furcata succulenta
rubescens glabra, in sicco fusca, caule vix 2 mm. crasso, internodiis
brevibus plus minusve furfuraceo-exfoliantibus; folia vulgo ternata
elliptica utrinque acuta parva 10-15 mm. longa 5-7 mm. lata carno-
FLORA OF PERU 37
sissima in sicco plicata 1-nervia, petiolo brevissimo 1-2 mm. longo;
spicae terminates et ex axillis superioribus 60 mm. longae 1.5 mm.
crassae densiflorae, pedunculo 10-17 mm. longo; bracteae rotundo-
peltatae; ovarium immersum ovoideo-acuminatum, stigmate fere
apicali.
Huanuco: Cushi, 1,500 meters, Macbride 4816 (type in herb.
Field Mus. No. 535,850).
Peperomia cusilluyocana Trelease, sp. nov.
Herba stolonifera modice parva plus minusve ramosa glabra,
caule 2-3 mm. crasso; folia alterna oblanceolata obtusa vel subacu-
minata basi cuneata circa 2.5 cm. longa et 1.5 cm. lata e dimidio
inferiore submultiplinervia, ramulis costae 4, firma olivacea, petiolo
vix 5 mm. longo amplexicauli ; spicae terminales 30 mm. longae 1
mm. crassae densiflorae, pedunculo filiformi 10-15 mm. longo; brac-
teae rotundo-peltatae.
Cuzco: Pillahuata, Cerro de Cusilluyoc, 2,400 meters, Pennell
14016 (herb. Field Mus. No. 558,254, type).
Peperomia cyclophylla Miq. in Seem. Bot. Voy. Herald 198.
1856; Dahlst. Svensk. Vet. Akad. Handl. 33, pt. 2: pi. 5.
A small, repent, more or less crisp-pilose herb; stem filiform;
leaves opposite, orbicular, scarcely 10 mm. in diameter; petiole
almost suppressed; spikes terminal, 30-65 mm. long, 1 mm. thick;
peduncle 1 cm. long, 2-bracted at the middle; bracts of the spike
round-peltate; berries ovoid, pointed; stigma apical. Neg. 8306.
San Martin: Tarapoto, Spruce 4987, type. Junin: Puerto Lira,
Killip & Smith 26830. Rio Pinedo, Killip & Smith 23614. Loreto:
Puerto Arturo, Killip & Smith 27248, 27723.
Peperomia dasystachya Miq. Lond. Journ. Bot. 4: 416. 1845.
A moderate-sized, erect, short-velvety herb; stem 2 mm. thick;
leaves alternate, elliptic or subovate, blunt-acuminate, subacute at
the base 3-5 cm. long, 1-3 cm. wide, or sometimes 8 cm. long and 3.5
cm. wide, 5-nerved, subappressed-pubescent but the upper side
glabrate in age; petiole barely 10 mm. long; spikes terminal, 60 mm.
long, 2 mm. thick, velvety; peduncle scarcely 1 cm. long; bracts
round-peltate; stigma subapical. Neg. 8376.
Amazonas: Chachapoyas, Mathews 3229, type.
Peperomia debilipes Trelease, sp. nov.
Planta epiphytica parva rhizomatoso-assurgens glabra, caule
2-3 mm. crasso; folia alterna oblanceolato-obovata obtusa basi
38 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
cuneata 2-3 cm. longa 1-2 cm. lata 5-nervia, in sicco opaca olivacea,
in vivo subtus rubro-venosa, petiolo gracillimo 10-15 mm. longo;
spicae terminates 45 mm. longae 3 mm. crassae, pedunculo filiformi
2 cm. longo, pseudopedicellis brevibus; bracteae rotundo-peltatae.
Junin: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25545 (type in U. S. Nat. Herb.).
Peperomia debilipes var. dimorpha Trelease, var. nov.
Herba parva repens arboricola paullo crispo-pubescens vel glab-
rescens; folia alterna, ramulorum sterilium rotundo-elliptica et 5-10
mm. longa, cetera subovato-elliptica et usque ad 25 mm. longa atque
15 mm. lata, obscure 3-nervia, petiolo 5 mm. longo.
Junin: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25591 (type in U. S. Nat. Herb.).
Peperomia deceptrix Trelease, sp. nov.
Planta glabra epiphytica, ut videtur repens et radicans ex nodis
4-foliatis florifera; folia oblanceolata acuta basi cuneata 11-12 cm.
longa 3-3.5 cm. lata 7-9-nervia in sicco opaca viridia et subtenuiter
papyracea, petiolo 4-9 cm. longo; spicae (delapsae) ut videtur ad
apicem ramuli 5-6 cm. longi solitariae, juveniles 40 mm. longae 1 mm.
crassae, pedunculo filiformi 2 cm. longo; bracteae rotundo-peltatae.
Loreto: San Antonio, 110 meters, Killip & Smith 29325 (type in
U. S. Nat. Herb.).
Peperomia deficiens Trelease, sp. nov.
Herba modice alta simplex glabra, caulibus gracilibus; folia
alterna subdeltoideo-ovata basi rotundata vel subtruncata et dis-
tincte peltata vel breviter cordata et epeltata 5-5.5 cm. longa 3.5-4.5
cm. lata circa 7-nervia, in sicco tenuia subtus brunneo-granulosa,
petiolo gracili 2-4 cm. longo; inflorescentia non visa.
San Martin: Alto Rio Huallaga, 360-900 meters, Williams
5758a (herb. Field Mus. No. 624,781, type).
Peperomia dendroides Trelease, sp. nov.
Herba parva stolonifera epiphytica superne dense ramosa, caule
gracili 1-2 mm. crasso primo puberulo; folia vulgo 3-4-nata elliptico-
obovata vel oblongo-obovata obtusa basi acuta parva 6-9 mm.
longa 3-4 mm. lata cito glabrescentia valde indistincte pinnatinervia,
petiolo 1-2 mm. longo primo velutino; spicae terminales vix 25 mm.
longae et 1 mm. crassae, floribus laxiuscule insertis, pedunculo vix
5 mm. longo sulcato glabrato; bracteae rotundo-peltatae; baccae
FLORA OF PERU 39
semi-immersae rotundo-ovoideae rubrae scutulate subtruncatae,
stigmate anteriore.
Junin: Chahua, 2,700 meters, Macbride 3640 (herb. Field Mus.
No. 534,704, type). Mito, 1784.
Peperomia dendromorphis Trelease, sp. nov.
Herba parvula rhizomatosa, caulibus erectis superne ramosis
gracilibus 1-2 mm. crassis puberulis; folia inferne opposita vel vulgo
ad nodum 3-5 lanceolato-oblonga vel elliptico-subobovata obtusa
basi acuta 8-10 mm. longa 3-4 mm. lata transiente puberula obscure
pinnatinervia, petiolo 1 mm. longo; spicae terminales filiformes 20-
30 mm. longae in sicco fuscae, pedunculo 5-10 mm. longo; bracteae
rotundo-peltatae.
Cuzco: Marcapata, 3,000 meters, Weberbauer 7800 (herb. Field
Mus. No. 605,350, type).
Peperomia deodorata Trelease, sp. nov.
Herba modice parva rhizomatoso-assurgens glabra carnosa; folia
alterna elliptica vel subobovata obtusa vel emarginulata basi cuneata
4-7 cm. longa 2-3 cm. lata pinnatinervia opaca, petiolo 1-2 cm. longo;
spicae 1-2 pedunculum terminalem 1-bracteatum terminantes vel
ex axillis superioribus nascentes 120 mm. longae 2 mm. crassae,
pedunculo 15 mm. longo; bracteae rotundo-peltatae majusculae;
baccae ellipsoideae, rostro brevi crasso, stigmate anteriore.
Junin: Chanchamayo Valley, 1,500-1,800 meters, Carlos Schunke
381 (herb. Field Mus. No. 571,435, type), 551, 1569, 1493, 1584.
In its dried state this plant lacks the usual fragrance of P. mag-
noliaefolia and its allies.
Peperomia dependens R. & P. Fl. 1 : 33. pi. 48. 1798. Piper
dependens Vahl, Enum. 1: 338. 1805.
A rather large, glabrous herb; leaves alternate, ovate, acuminate,
cordate, supeltate at the very base, 15-16 cm. long, 9-13-nerved,
drying pellucid; petiole 10 cm. long; spikes paired in the axils of
reduced upper leaves, 80 mm. long, 3 mm. wide; peduncle 2 cm.
long; bracts ovate-peltate; berries ovoid; stigmas subapical. Neg.
8379.
Huanuco: Posuso, Pavon 36; Ruiz, type.
Peperomia Diehliana Trelease, sp. nov.
Herba parva fragilis succulenta ramosa, caule 1-2 mm. crasso
primo plus minusve hirtello; folia ad nodum circa 3 rotundo-elliptica
40 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
paullo apiculata vel reflexo-emarginulata basi subobtusa 6 mm. longa
5 mm. lata glabra, petiolo 1-2 mm. longo sparse hirtello; inflores-
centia non visa.
Type in herb. Field Mus. (No. 630,026), probably collected in
the Department of Cuzco, Adolfo Diehl 2526.
Similar to P. rosea.
Peperomia distractiflora Trelease, sp. nov.
Herba modice parva, caule 1-5 mm. crasso puberulo sed
glabrescente et propter exfoliationem lucido; folia ad nodum 3-5
elliptica vel superiora oblonga utrinque subobtusa 10 mm. longa
3-4 mm. lata, supra obscure papillosa, subtus pallidiora glanduloso-
granulosa, petiolo 1 mm. longo; spicae terminales et ex axillis
superioribus 30-50 mm. longae 1 mm. crassae in statu adulto
laxiflorae, pedunculo 5 mm. longo glabro; bracteae rotundo-peltatae
virides; ovarium immersum ovoideum, stigmate obliquo.
Junin: Hacienda Chalhuapuquio, Stevens 214, type.
Peperomia dolabriformis HBK. Nov. Gen. & Sp. 1: 60. pi. 4..
1815. Piper dolabriforme Poir. Encycl. Suppl. 4: 464. 1816.
A rather tall, simple, glabrescent, succulent herb; stem 5 mm.
thick; leaves alternate, compressed from the sides so as to stand
vertically and not horizontally, spatulate or dolabriform, more or
less obliquely submucronate, grooved on the upper edge, about 3
cm. long and 1.5 cm. wide, cuneately subsessile, coriaceous; spikes
10-60 mm. long, 1 mm. thick, in a panicle long-stalked above the
leaves; bracts round-peltate.
Piura: Huancabamba, Bonpland, type. Without locality, Weber-
bauer 6061 .
Peperomia dolabriformis var. velutina Trelease, var. nov.
Herba, caule dense velutino; folia 2.5 cm. longa 1 cm. lata;
spicae 30 mm. longae 1 mm. crassae.
Cajamarca: Cajamarca, 500-600 meters, Weberbauer 3797, type.
Peperomia Drapeta Trelease, sp. nov.
Herba modice parva diffusa arboricola, caule in statu sicco
anguloso subgracili 1-2 mm. crasso, ramis dense intertextis, sterilibus
ad apices crispo-villosis; folia alterna rotunda vel ovata et subacum-
inata, longiora basi subacuta, parva 1-3.5 cm. longa 1.5-2 cm. lata,
in sicco subopaca, 3-5-nervia, supra crispo-pubescentia vel nervis
exceptis glabrescentia, petiolo 1 cm. longo adpresso-piloso ; spicae
FLORA OF PERU 41
terminales et axillares 30-50 mm. longae 1 mm. crassae laxiflorae,
pedunculo 5 mm. longo sparse molliter piloso vel glabrato; bracteae
rotundo-peltatae majusculae; baccae rotundato-ovoideae mucrona-
tae, stigmate subapicali.
Huanuco: Huacachi near Muna, 1,950 meters, Macbride 4122
(herb. Field Mus. No. 535,193, type).
Peperomia elegans C. DC. in DC. Prodr. 16, pt. 1: 430. 1869.
A glabrous, spreading herb; stem slender; leaves alternate, lance-
elliptic, acute at both ends or gradually acuminate, 4-8 cm. long,
2.5-5 cm. wide, pinnately nerved; petiole 3-7 cm. long; spikes termi-
nal or sympodial, paired from a slender stalk 4 cm. long, 30-40 mm.
long, 2 mm. thick; peduncle 1-2 cm. long; bracts round-peltate;
berries oblong, with a slender beak; stigma anterior at the base of
the beak. Neg. 8319.
San Martin: Rio Mayo, Tarapoto, Spruce 4871, type. Huanuco:
Mountains of Huanuco, Haenke (?; herb. Prague, under the name P.
reptans Opiz).
Peperomia elegantifolia Trelease, sp. nov.
Herba mediocris glabra silvicola; folia alterna lanceolata vel
typice lanceolato-obovata subacuta basi cuneata 9-10 cm. longa
3-4.5 cm. lata e dimidio inferiore pinnatinervia, petiolo vix 1 cm.
longo; spicae in axillis binae et graciliter pedunculatae, pedunculo 3
cm. longo, vel pedunculo longiore et apice folio reducto onusto, 40-80
mm. longae, pedunculo ultimo 5-10 mm. longo; bracteae subelliptico-
peltatae.
San Martin: San Roque, 1,350-1,500 meters, Williams 7781 (herb.
Field Mus. No. 629,533, type). Loreto: Pumayacu, 600-1,200
meters, King 3180. "Sacha-chullcu."
Peperomia elliptico-rhombea C. DC. in DC. Prodr. 16,
pt. 1: 407. 1869.
A moderate-sized, suberect herb, from crisp-pubescent gla-
brescent; leaves alternate, subrhombic-elliptic or ovate, acute or
subobtuse at the base, 3-4 cm. long, 2-3 cm. wide, submultiple-
nerved from the lower half; petiole 10-15 mm. long; spikes terminal
and from the upper axils, 1 or 2 on a short, 1-bracted stalk, slender,
over 100 mm. long; peduncle 1 cm. long; bracts round-peltate;
berries round-ovoid, obliquely beaked; stigma at the base of
the beak.
42 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Type collected in Peru, without definite locality, Mathews 1835.
Huanuco: Casapi, Mathews 1689. San Martin: Tarapoto, Spruce
4095 (?).
Peperomia emarginata R. & P. Fl. 1: 33. pi. 49. 1798. Piper
emarginatum Vahl, Enum. 1: 339. 1805. Peperomia oblusifolia var.
emarginata Dahlst. Svensk. Vet. Akad. Handl. 33, pt. 2: 66.
pi 2. 1900.
A rather large, repent-assurgent, glabrous herb; stem 3-5 mm.
thick; leaves alternate, oblanceolate-spatulate, emarginate, cuneate,
12 cm. long, 5 cm. wide, pinnately nerved; petiole 2.5 cm. long;
spikes 1 or 2 on terminal or axillary stalks 4 cm. long, 120 mm. long,
10 mm. thick; peduncle 5 cm. long; bracts round-peltate; ovary
ovoid, obliquely beaked; stigma anterior.
Huanuco: Posuso, Ruiz 77, type.
Peperomia emarginulata C. DC. in DC. Prodr. 16, pt. 1: 433.
1869. P. obtusifolia Auct. in part as to Peru.
A moderately small, erect, glabrous herb; stem angular; leaves
alternate, obovate, the typically somewhat narrowed apex emarginu-
late, cuneate at the base, 3-3.5 cm. long, 1.5-2 cm. wide, pinnately
nerved below; petiole 5-10 mm. long; spikes paired on a slender
terminal stalk 2 cm. long, 50 mm. long, 2 mm. thick; peduncle 10-15
mm. long; bracts round-peltate.
Type collected in Peru by Pa von. Huanuco: Casapi, Mathews
1687.
Peperomia (?) enckeaefolia Trelease, sp. nov.
Herba plus minusve caespitosa glabra, interdum suffruticosa, 15
cm. tantum alta, caule 2 mm. crasso, internodiis brevibus; folia
alterna ovata subacuminata basi rotundata ad petiolum angustata
5-6 cm. longa 3.5-4 cm. lata 5-nervia, nervis ut venis transversis
conspicuis in statu adulto impressis, in sicco firmo-papyracea viridia
lucida, nervis serius albescentibus, margine superne tenui albescente,
petiolo 10 mm. longo; inflorescentia non visa.
Loreto: San Lorenzo, Rio Maranon Valley, 150 meters, Killip
& Smith 29201 (type in U. S. Nat. Herb.).
Peperomia enenyasensis Trelease, sp. nov.
Herba majuscula gracilis pendula epiphytica ubique minute
crispo-pubescens vel caulibus vetustioribus et pagina superiore
foliorum glabrescentibus; folia alterna subovata acute acuminata
FLORA OF PERU 43
basi acuta vel subobtusa 7-10 cm. longa 4-5 cm. lata pinnatinervia
in sicco tenuia fusca, petiolo 1.5-4 cm. longo; spicae geminatae
ramulum brevem gracili-bracteatum sympodialem terminantes,
adultae 150 mm. longae cum baccis 8 mm. diam., pedunculo 2 cm.
longo; bracteae rotundo-peltatae; baccae pallidae anguste oblongae
scutulo acuto oblique truncatae, stigmate centrali.
Junin: Enenas, Pichis Trail, 1,600-1,900 meters, Killip & Smith
25654 (type in U. S. Nat. Herb.).
Peperomia erythrocorma Trelease, sp. nov.
Herba modice parva repens tantum ciliata, caule subgracili 2-3
mm. crasso; folia alterna reniformia vel rotundo-ovata ad ovata
interdum abrupte breviacuminata, basi truncate subcordata vel
rotundata et breviter attenuata, mediocria 3-6 cm. longa 3.5-5 cm.
lata, in sicco opaca, subpalmatim 7-9-nervia transiente ciliata,
petiolo 2-2.5 cm. longo; inflorescentia non visa.
Huanuco: Posuso, 600 meters, Macbride 4678 (herb. Field Mus.).
Peperomia erythrostachya Trelease, sp. nov.
Herba majuscula glabra pendula epiphytica, caule modice
gracili 2-3 mm. crasso in sicco rubro-brunneo et sublucido; folia
alterna ovata vel elongato-ovata acuta vel subacuminata basi
rotundata vel subcordata majuscula 6-9 cm. longa 4 cm. lata, in
sicco opaca coriacea multiplinervia, ramulis costae 3-5x2 paribus
1-2 superioribus exceptis basilaribus, petiolo 2.5-6 cm. longo et
ultra; spicae axillares longae graciles 200 mm. longae 2 mm. crassae
obscure rubrae densiflorae, pedunculo 2 cm. longo medio bracteato;
bracteae rotundo-peltatae majusculae.
Huanuco: Posuso, 600 meters, Macbride 4667 (herb. Field Mus.
No. 535,741, type).
Peperomia estrellana Trelease, sp. nov.
Herba subparva glabra rhizomatosa epiphytica, ramis erectis
foliatis; folia ad nodum 3-5, foliis stolonum subspathulatis emargin-
ulatis basi cuneatis opace cinereo-viridibus obscure 3-nerviis, petiolo
2 mm. longo; spicae terminales 20 mm. longae 2 mm. crassae, pedun-
culo 15 mm. longo; bracteae rotundo-peltatae; baccae ovoideae
acutatae pseudocupula onustae, stigmate apicali.
Ayacucho: Estrella, 500 meters, Killip & Smith 23064 (type in
U. S. Nat. Herb.). Junin: Peren< Bridge, 700 meters, Killip &
Smith 25281.
44 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia exiguispica Trelease, sp. nov.
Herba gracilis subsimplex majuscula glabra arboricola, caule
2-3 mm. crasso; folia alterna lanceolato-elliptica utrinque subacuta
4-5 cm. longa 2-3 cm. lata obscure 5-nervia subtus glanduloso-
granulosa in sicco opaca, petiolo 10 mm. longo; spicae gracillimae
90 mm. longae, pedunculo 10 mm. longo; bracteae rotundo-peltatae.
Loreto: Puerto Arturo, 155-210 meters, Williams 5831 (herb.
Field Mus. No. 622,538, type).
Peperomia exilamenta Trelease, sp. nov.
Herba majuscula epiphytica vel terrestris, caule in sicco 3-4 mm.
crasso profunde angulato sparsissime molliter setuloso; folia vulgo
ternata late lanceolata vel elliptico-lanceolata obtuse acuminata
basi acuta 5-6 cm. longa 2-2.5 cm. lata 3-nervia, nervis supra im-
pressis subtus prominentibus, glabra supra impresso-punctulata
revoluta valde rigida, petiolo 5 mm. longo puberulo; spicae terminales
et ex axillis superioribus 120 mm. longae 1 mm. crassae, pedunculo
10-25 mm. longo; bracteae rotundo-peltatae; baccae ovoideae
acutatae pseudocupula onustae, stigmate apicali.
San Martin: Tarapoto, 360-900 meters, Williams -5722 (herb.
Field Mus. No. 622,513, type), 6258.
Peperomia exiliramea Trelease, sp. nov.
Herba majuscula glabra epiphytica, caule 3 mm. crasso; folia
alterna elliptica acute acuminata basi acuta 11-12 cm. longa 5-6
cm. lata e dimidio inferiore graciliter pinnatinervia, in sicco tenuia,
petiolo 1-2 cm. longo; spicae solitariae ramulos breves aphyllos
sympodiales basi anguste bibracteatos aequilongos terminantes 150
mm. longae 2 mm. crassae vel majores; bracteae rotundo-peltatae;
ovarium rotundatum acute subscutulatum.
Loreto: Santa Rosa, 135 meters, Killip & Smith 28768 (type in
U. S. Nat. Herb.).
Peperomia falsa A. W. Hill, Ann. Bot. 21: 151. pi. 15. 1907.
A small, glabrous, acaulescent herb; leaves from a corm rooting
from the sides and base; leaves round-peltate, 1-3 cm. in diameter,
subcoriaceous; petiole 3-5 cm. long; spikes 15-30 mm. long; peduncle
elongate; bracts round-ovate; berries globose-ellipsoid, reticulate;
pseudopodium thick and cylindric; stigma apical.
Puno: Pucara, 3,700 meters, among rocks, Weberbauer 451, type.
FLORA OF PERU 45
Peperomia famelica Trelease, sp. nov.
Herba depauperata terrestris plus minusve caespitosa, caule plus
minusve crispo-pubescente; folia alterna subovato-elliptica utrinque
subacuta 2.5-5 cm. longa 1.5-2.5 cm. lata 5-nervia in vivo supra
obscure viridia, nervis albis, subtus rosea, petiolo 5 mm. longo;
spicae graciles 60 mm. longae pseudopedicellis brevibus onustae,
pedunculo 10 mm. longo; bracteae rotundo-peltatae.
Junin: East of Quimiri Bridge, near La Merced, 800-1,300
meters, Killip & Smith 23961 (type in U. S. Nat. Herb.).
Peperomia ficta Trelease, sp. nov.
Herba valida caespitoso-repens succulenta epiphytica, caule 2-4
mm. crasso glabro in sicco sulcato; folia vulgo ternata rotundo-
elliptica vel subovata vel subobovata obtuse subacuminata basi
acuta 3-4 cm. longa 2-3 cm. lata circa 5-nervia glabra in sicco dura
revoluta opaca, petiolo 5 mm. longo brunneo-puberulo ; spicae
terminales, juveniles 30 mm. longae 2 mm. crassae, pedunculo 15 mm.
longo; bracteae rotundo-peltatae.
Junin: East of Quimiri Bridge, La Merced, 800-1,300 meters,
Killip & Smith 23989 (type in U. S. Nat. Herb.).
Peperomia Fieldiana Trelease, sp. nov.
Herba modice parva repens glabra, caule crassiusculo 3-5 mm.
crasso, internodiis brevibus, ramis adscendentibus brevibus; folia
alterna elliptica vel obovato-elliptica apice rotundato emarginata
basi longicuneata majuscula 10-13 cm. longa 4.5-6.5 cm. lata in
sicco subtenuia pinnatinervia, costae ramulis circa 5x2, petiolo
15-25 mm. longo alato; spicae solitariae vel ad nodos ramuli lateralis
1-foliati vix 25 mm. longi geminatae vix 110 mm. longae et 2 mm.
crassae densiflorae, pedunculis 2.5-3 cm. longis; bracteae rotundo-
peltatae parvae in pseudoverticillo circa 12; baccae oblongae breviter
mucronatae, stigmate anteriore.
Huanuco: Posuso, 600 meters, Macbride 4684 (herb. Field Mus.
No. 535,758, type). Junin: Rio Perene", Killip & Smith 25167.
Peperomia filiformis R. & P. Fl. 1: 31. pi. 47. 1798. Piper
brachystachyum Vahl, Enum. 1: 354. 1805. Troxirum filiforme Raf.
Sylv. Tellur. 85. 1838.
A transiently puberulent or quickly glabrate, repent herb; stem
filiform; leaves mostly 4 at a node, obovate, rounded at the apex,
cuneate at the base, 7 mm. long, 5 mm. wide, 3-nerved; petiole
46 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
very short; spikes terminal, 10 mm. long, 1 mm. thick, the peduncle
of equal length; bracts ovate-peltate; ovary ovoid, the stigma apical.
Huanuco: Chinchao, Ruiz & Pawn, type. Pampayaco, Poeppig.
Junin: Carpapata, 2,700-3,200 meters, Kittip & Smith 24392
Puno : Tabina, Lechler 2089. Sangaban, Lechler 2409.
Peperomia fissicola Trelease, sp. nov.
Herba modice alta glabra e fissuris rupium cito erecta, caule 3 mm.
crasso firmo; folia alterna ad apicem caulis aggregata recurva
dolabriformia utrinque acutata 15 mm. longa 5 mm. lata, petiolo vix
5 mm. longo; spicae paniculatae prolongationem longiusculam
gracilem paucifoliatam caulis terminantes 50 mm. longae 1 mm.
crassae, pedunculis 5 mm. longis; bracteae rotundo-peltatae.
Huanuco: Huanuco, 2,100 meters, Macbride 3255 (herb. Field
Mus. No. 534,325, type).
Peperomia flabilis Trelease, sp. nov.
Herba modice parva plus minusve furcata glabra, caule 2-3 mm.
crasso; folia ad nodum 3-4 obtuse acuminata basi acuta 2-3 cm.
longa 1-1.5 cm. lata 3-nervia opaca, subtus minute granulosa palli-
diora, petiolo 3-6 mm. longo gracili; spicae terminales et ex axillis
superioribus filiformes 100 mm. longae, pedunculo gracillimo 10
mm. longo; bracteae rotundo-peltatae.
Loreto: Victoria on the Amazon River, Williams 2988 (herb.
Field Mus. No. 608,872, type).
Peperomia flavamenta Trelease, sp. nov.
Herba modice parva repenti-caespitosa succulenta fere glabra,
caule 1-2 mm. crasso in sicco sulcato; folia 2-3-nata lanceolato-
ovata acuminata basi acuta 2-5 cm. longa 1-2 cm. lata 3- vel obscure
5-nervia, petiolo 3-5 mm. longo puberulo; spicae flavae terminales
filiformes 40 mm. longae, pedunculo 10 mm. longo; bracteae rotundo-
peltatae.
Loreto: Mishuyacu, near Iquitos, 100 meters, G. Klug 987 (herb.
Field Mus. No. 630,005, type).
Peperomia flavescens C. DC. in DC. Prodr. 16, pt. 1:
436. 1869.
A spreading herb, glabrous except for the loosely villous petioles
and lower leaf surface; stem 2-3 mm. thick; leaves alternate, ovate,
subacute, peltate close to the subcordate base, 3.5 cm. long, 2-3
cm. wide, 5-7-nerved, coriaceous and yellow; petiole 2 cm. long;
FLORA OF PERU 47
spikes subterminal, 80 mm. long, 3 mm. thick or larger; peduncle
2-3 cm. long; bracts round-peltate; berries ovoid, stout-beaked;
stigma anterior. Neg. 8527.
Type collected in Peru by Pavon.
Peperomia flavescens var. Lechleri Trelease, var. nov.
Planta pubescentia plus minusve persistente induta; folia 2.5 cm.
longa, 2.5 cm. lata.
Puno: Sangari, Lechler 2691 in part (type in herb. Kew.).
Peperomia flavescens var. (?) Markhami Trelease, var. nov.
Crispo-pubescens, foliis glabrescentibus 3.5 cm. longis 2 cm. latis,
costa lata pallida.
Puno: Tambopata, "carpeting the ground in parts of the forests,"
in 1910, Clements R. Markham 4 (type in herb. Kew.; detached
leaves only).
Peperomia flavescentifolia Trelease, sp. nov.
Herba gracilis pendula epiphytica essentialiter glaber; folia
elliptica subacuminata basi subacuta 3-5 cm. longa 2-3 cm. lata
obscure pinnatinervia subtus flavescentia granulosa, in sicco valde
coriacea, petiolo vix 5 mm. longo; spicae ad apicem ramuli brevis
sympodialis geminatae 100-150 mm. longae graciles, pedunculo
15 mm. longo; bracteae rotundo-peltatae; baccae pallidae conico-
oblongae acute oblique truncatae; stigma ad scutulum insertum.
Loreto: Iquitos, 100 meters, Killip & Smith 271 42 (type in U. S.
Nat. Herb.). Mishuyacu, near Iquitos, Klug 99 (lower leaves
lanceolate, 7 cm. long, 2-3 cm. wide).
Peperomia foliata Trelease, sp. nov.
Herba modice alta gracilis dense foliata, caule subquadrangulari
puberulo; folia vulgo ad nodum 6-8 inf erne reflexa et breviora lineari-
oblanceolata obtusa vel subacuta basi subacuta 10-15 mm. longa
3 mm. lata pinnatinervia, supra ad costam subpuberula, subtus
pallidiora granulosa, petiolo 1 mm. longo; spicae terminales et ex
axillis superioribus, juveniles 25 mm. longae 1 mm. crassae densi-
florae, pedunculo vix 3 mm. longo; bracteae rotundo-peltatae.
Junin: Huacapistana, Stevens 65 (type), 178.
Peperomia foliiflora R. & P. Fl. 1: 30. pi. 45. 1798. Phyllo-
bryon Pavoni Miq. Syst. 50. 1843; 111. Piper, pi. 2. Piper foliiflorum
Vahl, Enum. 1: 356. 1805.
48 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
A rather small, stoloniferous, erect, glabrous herb; stem 2-3
mm. thick; leaves alternately clustered at the top of the stem, ovate,
acuminate, cordate, 3^4 cm. long, 2-3 cm. wide, 9-nerved with the
longer nerves forked; petiole 2-4 cm. long, winged; spikes several,
umbellate from the apex of the peduncle; bracts round-peltate; berries
ellipsoid, 6 mm. long, 1 mm. wide, flat, distinctly wrinkled, nearly
sessile, mucronate; stigma apical. Neg. 8381.
Huanuco: Chinchao, Pawn 36, type. Jauja: Rio Masamerich,
1,500 meters, Weberbauer 6665. Junin: La Merced, Killip & Smith
24656.
The specific name appears as foliflora on the original plate.
Peperomia fortipes Trelease, sp. nov.
Herba parva repenti-assurgens glabra foliosa, caule 2 mm. crasso;
folia alterna oblanceolata obtusa basi cuneata 2-3 cm. longa 1-1.5
cm. lata obscure 3-nervia in sicco opaco-olivacea, petiolo 5 mm.
longo; spicae 40-50 mm. longae 2 mm. crassae densiflorae, pedunculo
15 mm. longo 1 mm. crasso; bracteae rotund o-peltatae.
Ayacucho: Aina, 750-1,000 meters, Killip & Smith 22589 (type
in U. S. Nat. Herb.).
Peperomia Fortunati Trelease, sp. nov. P. Mandoni Auct.
Herba modice longicaulis delicata glabra pellucida, ramulis
floriferis in sicco gracillimis; folia ad apices ramorum alterne con-
gesta subrhombeo-ovata vel elliptica acute vel obtusissime acuminata
basi acuta vel obtusa 13-20 cm. longa 8-10 cm. lata delicate 5-7-
nervia, costa obscure ramosa, petiolo 2 mm. longo; spicae solitariae
ad basin laminae adnatae 5 mm. longae fere sessiles; bracteae sub-
cucullatae distichae; baccae ellipsoideae breviter stipitatae, stylo
gracili, stigmate apicali.
Cuzco : Valley of Rio Apurimac, Fortunato L. Herrera, February,
1929 (herb. Field Mus. No. 589,470, type). Without locality,
Weberbauer 5684.
Peperomia fragilissima Trelease, sp. nov.
Herba carnosa fragilissima, caule gracili; folia alterna rotundo-
ovata obtusissima basi cordata 2-4 cm. longa 2-3.5 cm. lata multi-
plinervia, nervis longioribus furcatis, petiolo 2-3 mm. longo; spicae
axillares et terminales gracillimae 40-50 mm. longae, pedunculo
20-25 mm. longo.
Loreto(?): Yurimaguas, May, 1855, Spruce (type in herb. Kew.).
FLORA OF PERU 49
Peperomia furcata Opiz in Presl, Rel. Haenk. 164. 1831.
A small, repent herb, at most with ciliate leaves; leaves alternate,
round-ovate, subacute at both ends or round-based, 5-7-nerved with
the inner nerves confluent at the base; petiole 5-10 mm. long;
spikes terminal or paired on a sympodial common stalk, 40 mm. long,
2 mm. thick; peduncle 1-2 cm. long.
Type from mountains of Peru, Haenke.
Peperomia fuscispica C. DC. Bot. Jahrb. 40: 260. 1908.
A small, glabrous epiphyte; stem filiform; leaves alternate, round
or subobovate, acute-based, 9 mm. in diameter, 3- or less evidently
5-nerved; petiole 3 mm. long; spikes terminal, 20 mm. long, 2mm.
thick; peduncle 3 mm. long; bracts round- or elliptic-peltate; ovary
subovoid; stigma oblique.
Huanuco: Monzon to Rio Huallaga, Weberbauer 3627, type.
Peperomia galoides HBK. var. aromatica C. DC. Bot Jahrb.
40:266. 1908.
Aromatic; leaves oblong, somewhat black-punctulate.
Ancash: Caraz, Weberbauer 3227, type. "Tunacongona."
The typical form of the species has a wide distribution in tropical
America.
Peperomia Gayi C. DC. Ann. Cons. Jard. Geneve 2: 283. 1898.
A moderately small, simple, very dingy-hirsute herb; stem slender;
leaves opposite, round in cross section below, elliptic upward, short-
acuminate, acute-based, 2-4 cm. long, 1.5-2.5 cm. wide, 5-nerved,
black-punctulate beneath; petiole 1 cm. long; spikes terminal and
from the upper axils, 50 mm. long, 1 mm. thick; peduncle 1 cm. long;
bracts round-peltate; ovary obovoid; stigma oblique.
Based on material collected in Peru, Gay 2181, 2310, 2681.
Peperomia gibba C. DC. Verh. Bot. Ver. Brandenb. 47:
116. 1905.
A moderate-sized, spreading, glabrous herb; leaves alternate,
round-ovate, slightly mucronate, shallowly and openly cordate,
5-6 cm. long, 3.5 cm. wide, 7-nerved; petiole 3-4 cm. long; spikes
axillary and terminal, slender, 75 mm. long; peduncle 3-4 cm. long;
bracts elliptic-peltate; berries oblong, obliquely swollen at the base,
subtruncate; stigma subapical.
Loreto: Pongo de Cainarachi, Ule 6322, type.
50 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia Gilberti Trelease, sp. nov.
Herba minor stolonifero-erecta divaricato-ramosa, caule gracili
1-2 mm. crasso puberulo serius furfuraceo-exfoliante; folia ad nodum
circa 5 suboblonga, inferiora spathulato-revoluta, obtusa basi sub-
acuta parva 8-10 mm. longa 3-4 mm. lata in sicco fusca opaca non
manifeste nervata, supra puberula, petiolo vix 2 mm. longo puberulo;
spicae ramos terminantes 25 mm. longae 1 mm. crassae, floribus
laxiuscule dispositis, pedunculo vix 5 mm. longo subpuberulo;
bracteae rotundo-peltatae; baccae globosae submucronulatae, stig-
mate obliquo.
Cuzco: Ollantaitambo, 3,000 meters, Cook & Gilbert 740 (type in
U. S. Nat. Herb.).
Peperomia glabrirhachis Trelease, sp. nov.
Herba modice parva stolonifero-caespitosa, caule 1-2 mm. crasso
anguloso sparse retrorso-pubescente; folia ad nodum circa 4 sub-
rhombeo-elliptica vel obovata obtusa basi acuta 10 mm. longa 5-6
mm. lata 1-nervia revoluta, subtus sparse pubescentia, utrinque
papilloso-rugosa, petiolo vix 1 mm. longo; spicae terminales 15
mm. longae 1 mm. crassae glabrae, pedunculo filiformi velutino
spicam aequante; bracteae rotundo-peltatae.
Junin: Hacienda Chalhuapuquio, Stevens 211 (type). Huaca-
pistana, Stevens 64, 179. Enenas, Killip & Smith 25653.
Peperomia glareosa Trelease, sp. nov.
Herba majuscula caespitosa assurgens, caule 1 mm. vel prope
basin ad 4-5 mm. crasso in sicco sulcato puberulo granuloso; folia
ad nodum 2-4 late lanceolata obtusa basi acuta in sicco firma opaca,
subtus granulosa, 3-nervia, superiora 1.5 cm. longa 1 cm. lata, folia
ramulorum sterilium 3.5 cm. longa 1.5 cm. lata, petiolo 2-3 mm.
longo subpuberulo-granuloso; spicae terminales et axillares 55 mm.
longae 2 mm. crassae, pedunculo 15 mm. longo granuloso; bracteae
rotundo-peltatae; baccae ovoideae ad stylum attenuatae pseudo-
cupula onustae, stigmate apicali.
Loreto: Rio Santiago, 200 meters, Ynes Mexia 6236 (type in
herb. Univ. Calif.).
Peperomia granulatifolia Trelease, sp. nov.
Herba epiphytica majuscula pendula glabra, caulibus gracil-
limis; folia alterna elliptica acuminata basi acuta 4-7 cm. longa
2-3.5 cm. lata in sicco coriacea obscure pinnatinervia, petiolo 5
FLORA OF PERU 51
mm. longo; spicae terminales et sympodiales solitariae vel ad apicem
ramuli brevis 2-bracteati geminatae 100 mm. longae 1 mm. crassae
vel ultra, pedunculo 5-10 mm. longo.
Junin: Puerto Yessup, 400 meters, Killip & Smith 26390 (type
in U. S. Nat. Herb.).
Peperomia granulatilimba Trelease, sp. nov.
Herba epiphytica fere glabra pendula, caulibus gracilibus 2 mm.
crassis; folia alterna subovato-elliptica acute acuminata basi rotun-
data 6 cm. longa 3 cm. lata, vel elliptico-oblonga basi subacuta
atque 10 cm. longa et 4 cm. lata, pinnatim paucinervia pallido-
granulosa sparse pilosa longiciliata, petiolo 5-10 mm. longo; spicae
3 caulem brevem sympodialem terminantes 100 mm. longae 2 mm.
crassae, pedunculo vix 5 mm. longo; bracteae rotundo-peltatae;
baccae oblongae vel conice productae, oblique acuto-scutulatae,
stigmate centrali.
Loreto: Yurimaguas, Lower Rio Huallaga, 155-210 meters,
Llewelyn Williams 4617 (herb. Field Mus. No. 629,584, type).
La Victoria, Williams 2945, 2802, 3084. Pichis Trail, Killip &
Smith 26215. Pumayacu, 600-1,200 meters, on tree, Klug 8200.
Peperomia Grisari C. DC. Bull. Herb. Boiss. 6: 516. 1898.
A slender, repent-assurgent herb, the petiole and the nerves
beneath coarsely puberulent; leaves alternate, round-ovate, acute,
shallow-cordate with the margin crossing the petiole, 7 cm. long,
6 cm. wide, 9-nerved, paler beneath; petiole 4 cm. long; spikes
terminal, paired on a stalk 2 cm. long, 50 mm. long, 2 mm. thick;
peduncle 15-20 mm. long.
Type collected in Ecuador or Peru, Grisar.
Peperomia guayrapurana Trelease, sp. nov.
Herba gracilis, caule, petiolis et foliis subtus rigido-hirtellis;
folia ad nodum 3-4 elliptica utrinque acuta 1.5-2 cm. longa 1 cm.
lata 3-nervia, petiolo vix 5 mm. longo; spicae terminales 20 mm.
longae 1 mm. crassae, pedunculo 1 cm. longo.
San Martin: Monte Guayrapurana, near Tarapoto, Spruce 4010
(type in herb. Kew.).
Peperomia gucayana Trelease, sp. nov.
Herba erecta vix ramosa firma, caule 2-3 mm. crasso minute
fusco-papilloso; folia plerumque opposita late obovata utrinque
acuta vix 2 cm. longa et 1.5 cm. lata 3-nervia, costa plus minusve
52 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
ramosa, in sicco tenuia et saepe conduplicata, petiolo 2-4 mm.
longo minute papillose; spicae axillares 60-80 mm. longae 1 mm.
crassae, floribus subdense dispositis, pedunculo 10 mm. longo;
bracteae rotundo-peltatae ; baccae ovoideae crasse submucronatae,
stigmate obliquo.
Cuzco: Near Gucay, Urubamba Valley, 3,000 meters, Fortunato
L. Herrera 1464 (type in U. S. Nat. Herb.).
Peperomia haematolepis Trelease, sp. nov.
Herba majuscula (repenti-assurgens?) succulenta glabra, caule
3-5 mm. crasso furfuraceo-exfoliante; folia alterna orbicularia vel
rotundo-subobovata emarginulata basi rotundata vel subacuta 4.5
cm. longa 3-4 cm. lata pinnatim paucinervia, coriacea opaca, petiolo
5-10 mm. longo; spicae terminales, juveniles 15-20 mm. longae 3
mm. crassae, pedunculo 2-2.5 cm. longo; bracteae rotundo-peltatae
rubro-aurantiacae.
Junin: Hacienda Chalhuapuquio, Stevens 212 (type).
Peperomia Haenkeana Opiz in Presl, Rel. Haenk. 162. 1831.
A rather small, erect herb, the stem, petioles, and nerves beneath
short- villous or crisp-pubescent; leaves alternately clustered at the
top of the stem, round-ovate, acuminate, cordate with overlapping
lobes, 4-6 cm. long, 3-5 cm. wide, 7-9-nerved with some nerves
forked; petiole 2-3 cm. long; spikes several, umbellate at the end
of the peduncle, nearly sessile, barely 10 mm. long, flattened and
flowered from the edge; bracts round-peltate.
Type from mountains of Peru, Haenke.
Peperomia hamiltonianifolia Trelease, sp. nov.
Herba parva stolonifero-erecta glabra, caule 2 mm. crasso; folia
alterna pauca elliptica utrinque subacuta 2-2.5 cm. longa 1 cm.
lata in sicco crassa subtus 5-nervia, petiolo 3-5 mm. longo; spicae
terminales et axillares 25 mm. longae 1 mm. crassae, pedunculo 5
mm. longo; bracteae rotundo-peltatae comparate magnae.
Type collected in Peru by Gay (herb. Paris).
Peperomia Herrerae Trelease, sp. nov.
Herba assurgens plus minusve ramosa modice parva glabra,
caule 2-3 mm. crasso; folia opposita elliptica utrinque acuta 2-2.5
cm. longa 1-1.5 cm. lata delicate 3-nervia, costa plus minusve
ramosa, in sicco tenuia viridia, petiolo 2 mm. longo; spicae termi-
nales et axillares subabundantes 40-50 mm. longae 2 mm. crassae,
FLORA OF PERU 53
floribus sublaxe dispositis, pedunculo gracili 15 mm. longo; bracteae
rotundo-peltatae; baccae rotundato-ovoideae mucronatae; stigma
subapicale plus minusve lobatum.
Cuzco: Hacienda Churu, Paucartambo, 3,500 meters, Fortunate
L. Herrera, February, 1924 (type in U. S. Nat. Herb.).
Peperomia heterophylla Miq. Lond. Journ. Bot. 4: 415. 1845,
A small, repent-assurgent epiphyte, glabrous except that the
filiform stems are at first puberulent; leaves alternate, round-
obovate below and 5 mm. in diameter, becoming obovate in the
middle and lanceolate and 2 cm. long upward, with the narrowed
apex subemarginulate, acute-based, 1-nerved, punctulate beneath;
petiole very short; spikes curved, 50 mm. long, 1 mm. thick; bracts
round-peltate.
Huanuco: Casapi, Mathews 1686, type.
Peperomia heterostachya A. Dietr. Syn. PL 1: 717. 1831.
Peperomia tristachya Opiz in Presl, Rel. Haenk. 103. pi. 30. 1831,
non P. tristachya HBK. 1815. Piper heterostachyon D. Dietr. Syn.
PI. 1: 111. 1839.
A glabrous, succulent, branching herb: leaves opposite below,
3 or 4 at a node above, lance-ovate, long-acuminate, acute-based,
7 cm. long, 3 cm. wide, 3-nerved; petiole 3 mm. long; spikes terminal
and from the uppermost axils, elongate.
Huanuco: Huanuco mountains, Haenke, type.
Peperomia Hillii Trelease, nom. nov. Peperomia Gaudichaudii
A. W. Hill, Ann. Bot. 21: 154. 1907, non P. Gaudichaudii Miq. 1843.
A small, glabrous, acaulescent herb; leaves from a small, rooted
corm, round or subdeltoid, peltate rather below the middle, 5-10
mm. long, 6-8 mm. wide; petiole 2-8 cm. long; spikes 60-80 mm.
long; peduncle equaling the petiole; bracts elliptic-peltate; berries
round-ovoid, slightly reticulate. Neg. 8345.
Lima: Lima and San Lorenzo, Gaudichaud 150. Atocongo,
clefts of limestone rocks, 250-500 meters, Pennell 14750. Chile.
Peperomia huacachiana Trelease, sp. nov.
Herba majuscula assurgens glabra, caule crassiusculo 3-6 mm.
crasso, internodiis brevibus; folia alterna elliptica vel lanceolato-
elliptica acuta vel acuminata basin versus sensim acuta, in sicco
subtenuia opaca, subtus pallidiora, pinnatinervia ramis costae circa
8-10x2, petiolo 1-2 cm. longo alato; spicae solitariae terminales
54 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
vel oppositifoliae ramulum crassum 1-bracteatum vel 1-foliatum
1.5-3 cm. longum terminantes, magnae 200 mm. longae 5 mm.
crassae floribus sublaxe insertis; bracteae rotundo-peltatae modice
magnae diametro suo remotae, floribus in alveolis lenticularibus
immersis; baccae ellipsoideo-oblongae fuscae suboblique truncatae,
stigmate in disco anteriore.
Huanuco: Huacachi near Muna, 1,950 meters, Macbride 3899
(herb. Field Mus. No. 534,962, type). Cani near Mito, 2,550
meters, 341 6.
Peperomia huacapistanana Trelease, sp. nov.
Herba mediocris glabra repens epiphytica, ramis fructiferis '
brevibus erectis, foliis reductis, caule 2-3 mm. crasso; folia alterna '
orbicularia vel ovata vel subreniformia apice rotundata basi sub-
truncata 3.5-4 cm. longa 2.5-3.5 cm. lata 7-nervia, nervis interiori- I
bus interdum confluentibus, in sicco coriacea supra obscure viridia
subtus pallidiora, petiolo 1.5-2 cm. longo; spicae solitariae vel
geminatae vel 4 et ultra et paniculatae, foliis ramulorum fertilium
bracteiformibus, 25 mm. longae 2 mm. crassae densiflorae, pedunculo
filiformi 10 mm. longo; bracteae rotundo-peltatae.
Junin: Huacapistana, Stevens 89 (type). Junin: Enefias, Killip
& Smith 25655, 25736. San Nicolas, Killip & Smith 26064. Hua-
nuco: Posuso, 4678.
Peperomia huallagana Trelease, sp. nov.
Herba epiphytica modice parva erecta plus minusve dichotoma
glabra, caule 2-3 mm. crasso; folia ad nodum vulgo 3-4 obovata
ad obovato-oblonga in eodem verticillo, emarginulata basi cuneata
12-20 mm. longa 6-10 mm. lata obscure 3-nervia, costa indistincte
ramosa, petiolo 2-5 mm. longo alato; spicae terminales parvae 30
mm. longae 1 mm. crassae densiflorae, pedunculo 1 cm. longo;
bracteae rotundo-peltatae, floribus inter costas anastomosantes
rhachis immersis.
Huanuco: Rio Huallaga Canyon below Rio Santo Domingo,
1,200 meters, Macbride 4264 (herb. Field Mus. No. 535,345, type).
Peperomia huallagana var. compacta Trelease, var. nov.
Compacte ramosa brevis; spicae 40 mm. longae 2 mm. crassae,
pedunculo duplo breviore.
Junin: Chanchamayo Valley, Carlos Schunke 1774 (herb. Field
Mus. No. 625,560, type).
FLORA OF PERU 55
Peperomia huantana Trelease, sp. nov.
Herba subparva repens corticicola, caule 1 mm. crasso sparse
crispo-subvilloso; folia ad nodum 2-4 rhombeo-obovata ad sub-
elliptica obtusa basi acuta 10-15 mm. longa 7-10 mm. lata subtus
sparse subvillosa et pallidiora, in sicco tenuia opaca, petiolo 2-3
mm. longo; spicae terminales (?) filiformes 60 mm. longae, pedunculo
20 mm. longo; bracteae rotundo-peltatae; baccae ovoideae acutatae
pseudocupula onustae, stigmate apicali.
Ayacucho: Aina, 750-1,000 meters, Killip & Smith 22728 (type
in U. S. Nat.Herb.), 22780. Junin: Yapas, Killip & Smith 25510(1}.
Peperomia huantana var. enenyasana Trelease, var. nov.
Pubescentia longior; folia magis lanceolata 20 mm. longa 10
mm. lata.
Junin: Enenas, Pichis Trail, 1,600-1,900 meters, Killip & Smith
25727 (type in U. S. Nat. Herb.).
Peperomia huanucoana Trelease, sp. nov.
Herba modice parva assurgens ramosa, caule gracili 1-2 mm.
crasso dense papilloso-puberulo; folia ad nodum vulgo 3-4 patentia
vel reflexa ovato-elliptica vel oblongo-elliptica utrinque obtusa sub-
sessilia parva 10-20 mm. longa 5-6 mm. lata utrinque papilloso-
puberula subtenuia anguste 3-nervia, costa plus minusve" ramosa;
spicae terminales et axillares 40 mm. longae 1 mm. crassae laxi-
florae, pedunculo vix 10 mm. longo papilloso-puberulo; bracteae
rotundo-peltatae; ovarium impressum ovoideum, stigmate sub-
apicali.
Huanuco: Huanuco, 2,100 meters, on ledges of rock cliffs, Mac-
bride 3256 (herb. Field Mus. No. 534,326, type).
On stolons the foliage is very different: leaves overlapping,
round-elliptic, thick, revolute, and scarcely more than 5 mm. long
and 4 mm. wide.
Peperomia hyporhoda Trelease, sp. nov.
Herba simplex mediocris rhizomatoso-assurgens, caule 2-3 mm.
crasso plus minusve crispo-pubescente; folia alterna subelliptica
vel inferiora minora et obovata obtusa vel subacuminata basi
acuta 2.5-4 cm. longa 2-3 cm. lata 3- vel minus distincte 5-nervia,
in sicco granuloso-punctulata, in vivo subtus rubra sed in sicco
brunnescentia, petiolo 1 cm. longo crispo-pubescente; spicae 2
apicales 100 mm. longae 3 mm. crassae pseudopedicellis onustae,
pedunculo 10-15 mm. longo; bracteae rotundo-peltatae.
56 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Junin: Yapas, Pichis Trail, 1,350-1,600 meters, Killip & Smith
25479 (type in U. S. Nat. Herb.).
Peperomia inaequalifolia R. & P. Fl. 1: 30. pi. 46. 1798.
Piper inaequalifolium Vahl, Enum. 1 : 355. 1805. Piper aromaticum
Willd. Enum. PI. Suppl. 3. 1813.
A rather large, aromatic, subcespitose herb, or suffrutescent;
leaves 4 at a node, oblong-spatulate, obtuse, cuneate at the base,
5 cm. long, 1.5 cm. wide, reduced below to scarcely 3.5 cm. long
and 1 cm. wide, 3-nerved; petiole 5 mm. long; spikes terminal and
axillary, elongate.
Type material from Lima and Huanuco, Pavon. Without
locality, Mathews 124, 933.
Peperomia inaequilatera Trelease, sp. nov.
Herba modice parva (repens?) assurgens dense puberula, caule
subgracili 2-3 mm. crasso, internodiis brevibus; folia ad nodum
vulgo ternata elliptico-oblonga utrinque subacuta vel obtusa parva
2 cm. longa 1 cm. lata, foliis ramulorum sterilium majoribus 4.5
cm. longis 2 cm. latis vel multo majoribus 7 cm. longis et 2.5 cm.
latis atque rhombeo-oblanceolatis, uno latere ad petiolum 3 mm.
profundius quam altero decurrente, in sicco tenuia pellucida, 5-
nervia costa plus minusve ramosa, petiolo brevi vix 5 mm. longo;
spicae terminales 60 mm. longae 1 mm. crassae, floribus laxe sub-
annulatis, pedunculo 1 cm. longo ut rhachi puberulo; bracteae
rotundo-peltatae; ovarium ovoideum, stigmate subapicali.
Huanuco: Piedra Grande near Rio Santo Domingo, 1,500 meters,
Macbride 3858 (herb. Field Mus. No. 534,921, type).
Peperomia induratifolia Trelease, sp. nov.
Herba parva essentialiter glabra repens vel stolonifera in sicco
nigrescens, caule 1-2 mm. crasso, internodiis brevibus; folia alterna
vel suprema ternata lanceolata acuminata basi acuta 20-30 mm.
longa 5-12 mm. lata opaca vix plus quam 1-nervia, petiolo 5-10
mm. longo; spicae ut videtur terminales filiformes 80-90 mm.
longae, pedunculo 10 mm. longo; bracteae rotundo-peltatae.
Junin: Chanchamayo Valley, 1,500 meters, Carlos Schunke 1582
(herb. Field Mus. No. 627,436, type).
Peperomia itayana Trelease, sp. nov.
Herba subparva brevicaulis glabra; folia pauca rotundo-ovata
obtusissima basi brevi ter cordata 7-9 cm. longa 6-8 cm. lata 7-
FLORA OF PERU 57
nervia, costa nervisque lateralibus 4-ramosis, in sicco tenuia viridia,
petiolo 8-10 mm. longo (et ultra?); inflorescentia non visa.
Loreto: Soledad, Rio Itaya, 110 meters, Killip & Smith 29753
(type in U. S. Nat. Herb.), 29618. San Antonio, Killip & Smith
29364, younger and smaller. Santa Rosa, Killip & Smith 28724-
Peperomia juniniana Trelease, sp. nov.
Herba mediocris ramosa, caule 1-3 mm. crasso transiente crispo-
villoso; folia alterna ovata acuminata basi breviter cordata basi
intra sinum breviter cuneata, 4-10 cm. longa 3-5 cm. lata multi-
plinervia nervis lateralibus furcatis, costa ramosa, in sicco tenuia,
subtus pallida minute nigro-punctata, nervis hirtellis, petiolo 1-4
cm. longo crispo-hirtello; spicae axillares, juveniles 65 mm. longae
2 mm. crassae densiflorae, pedunculo 1 cm. longo hirtello; bracteae
rotundo-peltatae.
Junin: Chanchamayo Valley, 1,200 meters, Carlos Schunke 357
(herb. Field Mus. No. 571,411, type); Killip & Smith 24809.
Peperomia Klotzschiana Miq. Syst. 145. 1843.
A moderate-sized, hirsute herb; stem 2-3 mm. thick, branching
upward; leaves 4-6 at a node, oblanceolate-subspatulate, subacute
at both ends, 20-25 mm. long, 6-10 mm. wide, 3-nerved; petiole
5 mm. long; spikes numerous in the axils of the congested and
reduced upper leaves, 10 mm. long, 1 mm. thick; peduncle very
short; bracts round-peltate, pale-margined.
Loreto (?) : Mainas, Poeppig, type.
Peperomia lanosa Trelease, sp. nov.
Herba modice alta vix valida stolonifero-erecta, caule 1-3 mm.
crasso, internodiis longiusculis dense longipilosis; folia inferiora
opposita obovato-cuneata 2 cm. longa 1.5 cm. lata longe (1 cm.)
petiolata, superiora vulgo ad nodum 3-5 spathulata vel suboblonga
et brevipetiolata, 3 cm. longa et 1 cm. lata vel 2 cm. tantum longa
et 6 mm. lata, 3-nervia, utrinque vel supra tantum ad nervos ad-
presso-pilosa, petiolo dense piloso; spicae terminales et axillares
40-70 mm. longae 1 mm. crassae laxiflorae, pedunculo 1.5-2 cm.
longo piloso; bracteae rotundo-peltatae; baccae ovoideae mucronatae,
stigmate oblique apicali.
Huanuco: Cani near Mito, 2,550 meters, Macbride 3450 (herb.
Field Mus. No. 534,525, type).
58 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia Lechleriana Trelease, sp. nov.
Herba majuscula glabra; folia alterna subelliptico-lanceolata
utrinque acuta 14-19 cm. longa 5.5-6.5 cm. lata pinnatinervia,
petiolo 2 cm. longo; spicae anguste paniculatae caudiculum scapi-
formem terminantes 10 mm. longae 1 mm. crassae, pedunculo
brevissimo; bracteae rotundo-peltatae; baccae ellipsoideae oblique
truncatae, stigmate centrali.
Puno: Sangaban, Lechler 2375 (herb. Kew.).
Peperomia lepadiphylla Trelease, sp. nov.
Herba alpina intertexta fere glabra, caule mediocri 2-3 mm.
crasso usque ad apicem foliato; folia alterna crassissima compressa
V-sulcata oblique ovata acuta basi rotundata 10 mm. longa 6 mm.
lata enervia ad costam plus minusve puberula, petiolo filiformi
vix 5 mm. longo; spicae terminales 20 mm. longae 3 mm. crassae
densiflorae, pedunculo nullo; bracteae rotundo-peltatae; ovarium
ovoideum, stigmate apicali.
Lima: Rio Blanco, 4,500 meters, Macbride 3055 (herb. Field
Mus. No. 534,125, type).
Peperomia leuconeura Trelease, sp. nov.
Herba modice parva assurgens, caule subgracili 2 mm. crasso
crispo-pubescente; folia alterna rhombeo-elliptica utrinque sub-
acuminata modice parva 4 cm. longa 2.5 cm. lata 3- vel obscure
5-nervia, supra ad nervos crispo-pubescentia, subtus nigro-punctu-
lata et inter nervos albidos rubescentia, petiolo gracili 15 mm.
longo glabrato; spicae axillares et terminales, juveniles 25 mm.
longae 1 mm. crassae densiflorae, pedunculo brevissimo glabrato;
bracteae rotundo-peltatae pallido-marginatae.
Junin: Hacienda Schunke, La Merced, 1,200 meters, Macbride
5628 (herb. Field Mus. No. 536,667, type).
Peperomia limaensis Trelease, sp. nov.
Herba parva compacte ramosa in sicco fusca, caule 1-2 mm.
crasso, internodiis brevibus dense puberulis; folia ad nodum vulgo
4 obovata utrinque rotundata parva 5 mm. longa 3 mm. lata, supra
versus apicem leviter puberula, in sicco opaca non manifeste nervata,
petiolo vix 1 mm. longo glabrato; spicae terminales 50 mm. longae 1
mm. crassae densiflorae, pedunculo brevissimo vix 2 mm. longo;
baccae impressae ellipsoideae obtusae, stigmate subapicali.
Lima: San Geronimo, Lima, 150 meters, Macbride 5920 (herb.
Field Mus., type).
FLORA OF PERU 59
Peperomia longepedunculata Opiz in Presl, Rel. Haenk.
164. 1831.
A rather large but slender-stemmed, glabrous herb; leaves alter-
nate, ovate, acute or subacuminate, openly cordate, 9-13 cm. long,
7-9 cm. wide, multiple-nerved with about 6 nearly basal branches of
the midrib and 2 or 3 upward, the longer nerves forking; petiole
10-15 cm. long; spikes sympodial, 40 mm. long, 2-3 mm. thick;
peduncle slender, 3 cm. long.
Huanuco: Huanuco mountains, Haenke in 1790, type.
Peperomia longipila C. DC. Verh. Bot. Ver. Brandenb. 47:
115. 1905.
A small, suberect, long-pilose or villous herb; stem slender;
leaves alternate, round-ovate or elliptic, 11 mm. long, 8 mm. wide,
3-nerved ; petiole 5-10 mm. long; spikes extremely slender and loosely
flowered, scarcely 10 mm. long; peduncle 10 mm. long, glabrous;
bracts round-peltate; berries ellipsoid, scarcely stipitate, with a style;
stigma apical.
Loreto: On cliffs, Pongo de Cainarachi, Ule 6407, type.
Peperomia longispica Trelease, sp. nov.
Herba modice parva assurgens ramosa, caule gracili 1-2 mm.
crasso vel ad basin 4-5 mm. crasso evanescente papillose, furfuraceo-
exfoliante: folia vulgo ad nodum 4-6 primo patentia obovato-oblonga
vel oblonga apice rotundata basi subacuta subsessilia parva 9-15
mm. longa 4 mm. lata supra prope apicem puberula, subtenuia,
obscure pinnatinervia; spicae terminales et axillares elongatae 60-
150 mm. longae 1 mm. crassae laxiflorae, pedunculo vix 1 cm. longo
glabro; bracteae rotundo-peltatae: baccae globosae oblique mucron-
ulatae, e pseudopedicellis brevibus, stigmate subapicali.
Huanuco: Muna, 2,100 meters, Macbride 3928 (herb. Field Mus.
No. 534,990, type). Junin: Huacapistana, Killip & Smith 24140,
24188, 24147. Carpapata, Killip & Smith 24342. Carrapa, Killip
& Smith 22380, 22426.
Peperomia Macbrideana Trelease, sp. nov.
Herba majuscula assurgens, caule crassiusculo 3-6 mm. crasso,
internodiis brevibus glabratis; folia alterna vel interdum subopposita
rotundo-elliptica vel orbicularia utrinque rotundata mediocria 4-6
cm. longa 3.5-5 cm. lata palmatim 5-nervia, nervis ramosis, subtus
ad nervos ferrugineo-villosa, petiolo subgracili 3-8 cm. longo,
60 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
foliorum superiorum breviore; spicae ad apicem ramuli pilosi
spicatim congestae, juveniles vix 5 mm. longae; bracteae rotundo-
peltatae.
Huanuco: Mima, Huacachi, 1,950 meters, Macbride 4144 (herb.
Field Mus. No. 535,218, type).
Peperomia macrorhiza HBK. Nov. Gen. & Sp. 1: 72. 1815.
A small, glabrous, acaulescent herb; leaves from a (rather large?)
corm, round, peltate, 6 mm. in diameter; petiole 5 cm. long; spikes as
yet small; bracts round-peltate.
Cajamarca: Cajamarca, Humboldt, type.
Peperomia macrothyrsa Miq. Lond. Journ. Bot. 4: 419. 1845.
A glabrous, succulent, mountain herb, stoloniferous-assurgent,
sometimes dwarfed; stem rather thick, naked below; leaves clustered
alternately at the top of the stem, subelliptic, rather obtuse, acute-
based, 20 mm. long, 10 mm. wide; petiole 1 cm. long; spikes on a
prolonged, somewhat branched shoot, nearly sessile in the axils of
gradually reduced leaves, 40 mm. long, 1 mm. thick; bracts
round-peltate; berries round-ovoid, mucronate; stigma apical.
Neg. 8385.
Amazonas: Chachapoyas, Mathews 3228, type.
Peperomia majalis Trelease, sp. nov.
Herba mediocris arboricola foliis ciliatis exceptis glabra, caule 4
mm. crasso, internodiis brevibus; folia alterna elliptica vel typice
ovata subacuta basi rotundata vel breviter cordata prope basin
peltata, 6-10 cm. longa 3.5-6.5 cm. lata graciliter multiplinervia, in
sicco flavescenti-brunnea et coriacea, petiolo 2-4 cm. longo; spicae
terminales ad apicem ramuli 1-bracteati geminatae 40 mm. longae
2 mm. crassae, pedunculo 1 cm. longo; bracteae rotundo-peltatae.
Junin: Dos de Mayo, Pichis Trail, 1,700-1,900 meters, Killip &
Smith 25886 (type in U. S. Nat. Herb.).
Peperomia mansericheana Trelease, sp. nov.
Herba parva repenti-assurgens arboricola, caule filiformi puber-
ulo; folia alterna lanceolata obtusa basi acuta 18-28 mm. longa 6-7
mm. lata glabra, in sicco tenuia 1-3-nervia, petiolo 2 mm. longo;
spicae terminales filiformes 40 mm. longae, pedunculo 10 mm. longo;
bracteae rotundo-peltatae.
Loreto: Mouth of Rio Santiago, Pongo de Manseriche, 210
meters, Ynes Mexia 6196 (type in herb. Univ. Calif.).
FLORA OF PERU 61
Peperomia Maransara Trelease, sp. nov.
Herba procumbens, caulibus gracilibus 1-2 mm. crassis obscure
adpresso-puberulis; folia acuminata basi rotundata vel subtruncato-
cordulata modice parva 2.5-5 cm. longa 2-3.5 cm. lata in sicco
viridia firma ciliata 5-nervia supra ad nervos primo adpresso-pu-
bescentia, petiolo 2-5 cm. longo glabro; spicae ramulos puberulos
3-4 cm. longos prope medium 1-bracteatos terminantes 30 mm.
longae 2 mm. crassae densiflorae; bracteae rotundo-peltatae; baccae
fuscae oblongae, rostro fere aequilongo setaceo; stigma ad basin
rostri anterius.
Cuzco: San Miguel, Urubamba Valley, 1,800 meters, Cook &
Gilbert 960 (type in U. S. Nat. Herb.).
Peperomia marcapatana Trelease, sp. nov.
Herba mediocris repenti-assurgens plus minusve ramosa glabra,
caule 3-4 mm. crasso basi plus minusve lignescente; folia ad nodum
4-6 rotundo-elliptica emarginulata basi obtusa 10-15 mm. longa
6-10 mm. lata 1-nervia opaca subrevoluta, subtus granulosa, petiolo
3-5 mm. longo; spicae terminales 120 mm. longae 3 mm. crassae
laxiflorae, pedunculo 15-20 mm. longo; bracteae rotundo-peltatae,
floribus in alveolis insertis.
Cuzco: Marcapata, Prov. Quispicanchi, 3,000 meters, Weberbauer
7801 (herb. Field Mus. No. 605,155, type).
Peperomia Marshalliana Trelease, sp. nov.
Herba delicata glabra repens, caule filiformi sed firmo; folia
alterna obovata apice rotundata vel subtruncata basi acuta parva
10-20 mm. longa 7-12 mm. lata in sicco subcoriacea 5-nervia, petiolo
brevi vix 2 mm. longo; spicae terminales ramulos breves compacti-
foliatos terminantes parvae 10 mm. longae 1 mm. crassae densiflorae,
pedunculo spicae aequilongo; bracteae rotundo-peltatae; baccae sub-
globosae, stigmate obliquo.
Huanuco: Posuso, 600 meters, associated with P. scandens,
Macbride 4710 (herb. Field Mus. No. 551,172, type). Junin: Chan-
chamayo Valley, Carlos Schunke 460. San Martin: San Roque,
1,350-1,500 meters, Williams 7382.
Peperomia Mathewsiana Miq. Lond. Journ. Bot. 4: 423. 1845.
Erect, suffrutescent, pubescent; leaves 3-7 at a node, rhombic-
spatulate, obtusely narrowed at the apex, cuneate at the base, 3 cm.
62 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
long, pellucid, 3-nerved; petiole 3-5 mm. long; spikes terminal and
axillary, elongate; peduncle 5-7 mm. long; bracts round-peltate.
Huanuco: "Zacapota," Casapi, Mathews 1688, type.
Peperomia megalepis Trelease, sp. nov.
Herba modice parva repenti-assurgens glabra arboricola, caule
subgracili 2-3 mm. crasso, internodiis brevibus; folia alterna rhom-
beo-elliptica vel obovata subacuta parva 1.5-3.5 cm. longa 1-2 cm.
lata, in sicco opaca 5-nervia, petiolo 5-10 mm. longo; spicae ter-
minales et axillares 70 mm. longae 2 mm. crassae sublaxiflorae,
pedunculo fere 2 cm. longo; bracteae magnae 1 mm. latae rotundo-
peltatae.
Huanuco: Trail to Tambo de Vaca, Muna, 2,400 meters, Macbride
4275 (herb. Field Mus. No. 535,357, type).
Peperomia mercedana C. DC. Bot. Jahrb. 40: 299. 1908.
A rather large, glabrous herb; leaves alternate, subrhombic-
obovate, blunt-acuminate, cuneate at the base, 8-9 cm. long, 3.5-4
cm. wide, pinnately nerved; petiole barely 1 cm. long; spikes solitary
in the axils on a slender stalk, or 2 or 4 on a similar terminal stalk
3-4 cm. long; bracts round-peltate; ovary obovoid, obliquely beaked;
stigma anterior.
Junin: La Merced, Weberbauer 1882, type.
Peperomia mercedana var. carpapatana Trelease, var. nov.
Folia subrhombeo-obovata acutiuscule acuminata basi longi-
cuneata 11 cm. longa 5 cm. lata, petiolo 1-2 cm. longo.
Junin : Carpapata above Huacapistana, 2,700-3,200 meters, Killip
& Smith 24485 (type in U. S. Nat. Herb.).
Peperomia metallica Lind. & Rod. 111. Hort. 39: 79.
pi. 557. 1892.
A much-branched, succulent, glabrous herb with short, pellucid,
pinkish internodes; leaves lanceolate or lance-ovate, the narrowed
apex obtuse, rather obtuse-based, only 2 cm. long and 1 cm. wide,
or 6 cm. long and 2 cm. wide, pinnately nerved, bronzed, with pale
center above, drying opaque; petiole barely 5 mm. long; spikes
terminal, slender, 25-40 mm. long, rather loosely flowered; bracts
round-peltate; ovary ovoid, submucronate; stigma oblique.
Type collected in Peru.
A very beautiful plant, now in cultivation, its exact home not
recorded.
FLORA OF PERU 63
Peperomia microlepis Trelease, sp. nov.
Herba subparva suberecta glabra, caule 2-3 mm. crasso; folia
alterna subrhombeo-elliptica vel obovata emarginulato-acuminata
basi acuta 3-4 cm. longa 2 cm. lata 5-7-nervia in sicco subfirma,
subtus pallida, petiolo 5 mm. longo; spicae terminales 40 mm. longae
1 mm. crassae sublaxiflorae, pedunculo filiformi 15 mm. longo;
bracteae rotundo-peltatae crispae.
Junin: Huacapistana, Stevens 91 (type).
Peperomia micromamillata Trelease, sp. nov.
Herba subparva caespitosa succulenta in sicco fusca, caule 1-3
mm. crasso, internodiis brevibus primo velutinis cito furfuraceo-
exfoliantibus; folia congesta ad nodum circa 6 elliptica utrinque
subobtusa parva 8 mm. longa 3 mm. lata carnosissima opaca 1-nervia
subtus microscopice mamillata apice ciliato-barbata, petiolo brevis-
simo vix 1 mm. longo; spicae (terminales?) 60 mm. longae 1 mm.
crassae laxiflorae, pedunculo vix 10 mm. longo; bracteae rotundo-
peltatae; ovaria ovoidea, stigmate subapicali.
Huanuco: Huacachi near Muna, 1,950 meters, Macbride 3872
(herb. Field Mus. No. 534,935, type). Piedra Grande, 1,500
meters, 3687.
Peperomia milvifolia Trelease, sp. nov.
Herba subparva suberecta in statu juvenili papillosa, caule 2
mm. crasso; folia ad nodum vulgo 4 rhombeo-quadrata vel milvi-
formia subacuta basi cuneata 10-11 mm. longa 8 mm. lata 3-nervia
in sicco modice tenuia viridia, petiolo 1 mm. longo; spicae terminales
et ex axillis superioribus graciles ad 100 mm. longae, pseudopedicellis
longis filiformibus; bracteae rotundo-peltatae.
Type from Santa Cruz, 1,800-2,100 meters, Pearce, December,
1864 (herb. Kew.). There is some doubt as to whether the locality
is Peruvian or Bolivian.
Peperomia minuta A. W. Hill, Ann. Bot. 21: 159. pi. 15. 1907.
A small, glabrous, acaulescent herb resembling P. parvifolia;
leaves 2-3.5 mm. in diameter; berries subellipsoid, minutely punctu-
late-roughened ; stylopodium short-conic-cylindric.
Ancash: Ocros to Chonta, Weberbauer 2776, type. Cuzco: Pau-
cartambo Valley, 3,500 meters, H err era 1387.
64 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIII
Peperomia mishuyacana Trelease, sp. nov.
