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UNITED STATES 
DEPARTMENT OF AGRICULTURE 



/< 7 



Washington, D.C. 



February 1967 



HYMENOPTERA OF 

AMERICA NORTH OF MEXICO 

SYNOPTIC CATALOG 

(Agriculture Monograph No. 2) 

SECOND SUPPLEMENT 

Prepared by the staff and collaborators of the Hymenoptera Unit, 
Insect Identification and Parasite Introduction Research Branch, 
Entomology Research Division, Agricultural Research Service, 

under the direction of 
KARL V. KROMBEIN and B. D. BURKS 



^ nj 
= a 

= CD 

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UNITED STATES 

GOVERNMENT PRINTING OFFICE 

WASHINGTON: 1967 



For sale by the Superintendent of Documents, U. S. Government Printing Office 
Washington, D.C. 20402 Price $2.75 



AUTHORSHIP 

This publication was prepared by the staff and collaborators 
of the Hymenoptera Unit whose individual responsibilities were 
as follows: Introductory section and general editorial super- 
vision by Karl V. KromlDein * and B. D. Burks ; Symphyta and 
Chalcidoidea by B. D. Burks ; Braconidae and Pelecinoidea by C. 
F. W. Muesebeck; Ichneumonidae, Evanioidea, and Trigonalidae 
by L. M. Walkley ; Cynipoidea by the late L. H. Weld ; Proctotru- 
poidea by C. F. W. Muesebeck and L. Masner; Formicidae by M. 
R. Smith ; and Aculeata by Karl V. Krombein. 

* Now Chairman, Department of Entomology, Smithsonian Institution. 



HYMENOPTERA OF AMERICA NORTH OF MEXICO 



ACKNOWLEDGMENT 



We are indebted to a number of our colleagues for furnishing 
information which has been included in this supplement. Their 
names and the groups in which they furnished information are as 
follows : 



H. W. Allen (Ichneumonidae) 

J. de Beaumont (Sphecidae) 

G. Becker (Ichneumonidae) 

R. M Bohart (Vespoid and Sphecoid 

wasps) 
W. F. Buren (Formicidae) 
J. C. Downey (Ichneumonidae) 
L. E. Eighme (Sphecidae) 
H. E. Evans (Vespoid and Sphecoid 

wasps) 
W. E. Ferguson (Mutillidae) 
J. E. Gillaspy (Sphecidae) 
J. D. Haddock (Mutillidae) 
C. P. Haskins (Formicidae) 
G. Heinrich (Ichneumonidae) 
W. W. Judd (Biology) 
P. B. Kannowski (Formicidae) 
R. L, King (Formicidae) 
F. E. Kurczewski (Sphecidae) 
W. R. M. Mason (Ichneumonidae) 



A. Menke (Sphecidae) 
C. D. F. Miller (Chalcidoidea) 
H. E. Milliron (Symphyta, Chalci- 
doidea, and Apidae) 
T. B. Mitchell (Apoidea) 
S. Momoi (Ichneumonidae) 

F. D. Parker (host-parasite records) 
0. Peck (Chalcidoidea) 

J. F. Perkins (Ichneumonidae) 
T. C. Schneirla (Formicidae) 
H. A. Scullen (Sphecidae) 
L. K. Smith (Ichneumonidae) 
R. R. Snelling (Colletidae) 
R. W. Taylor (Formicidae) 
P. H. Timberlake (Apoidea) 
H, K. and M, Townes (Ichneumoni- 
dae) 

G. S. Walley (Ichneumonidae) 
N. A. Weber (Formicidae) 

E. 0. Wilson (Formicidae) 



CONTENTS 

Page 

Introduction 4 

Order Hymenoptera 6 

Suborder Symphyta, by B. D. Burks 6 

Superfamily Megalodontoidea 6 

Superfamily Tenthredinoidea 14 

Superfamily Siricoidea 26 

Superfamily Cephoidea 27 

Suborder Apocrita 27 

Superfamily Ichneumonoidea 27 

Family Braconidae, by C. F. W. Muesebeck 27 

Family Ichneumonidae, by L. M. Walkley 60 

Superfamily Chalcidoidea, by B. D. Burks 213 

Superfamily Cynipoidea, by L. H. Weld 282 

Superfamily Evanioidea, by L. M. Walkley 284 

Superfamily Pelecinoidea, by C. F. W. Muesebeck 285 

Superfamily Proctotrupoidea, by C. F. W. Muesebeck and 

L. Masner 285 

Superfamily Bethyloidea, by Karl V. Krombein 305 

Family Trigonalidae, by L. M. Walkley 322 

Superfamily Scolioidea, by Karl V. Krombein 322 

Family Formicidae, by M, R. Smith 343 

Superfamily Vespoidea, by Karl V. Krombein 374 

Family Pompilidae, by Karl V. Krombein 378 

Superfamily Sphecoidea, by Karl V. Krombein 386 

Superfamily Apoidea, by Karl V. Krombein 422 

Index to new names proposed in the Supplement 521 

Index 521 



4 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Hymenoptera of America North of 
Mexico-Synoptic Catalog 

(Agriculture Monograph No. 2) 
SECOND SUPPLEMENT 

INTRODUCTION 

The "Hymenoptera of America North of Mexico — Synoptic 
Catalog" was published on July 11, 1951. It contained references 
to the literature published through 1949, and to a few papers that 
appeared during 1950. The First Supplement to this catalog was 
published on October 8, 1958. It covered the literature that ap- 
peared from 1950-1951 through 1956. The Second Supplement, 
which is presented herewith, includes the literature published 
from 1957 through 1963, and in some sections, most of the papers 
that appeared during 1964. 

In general, this Second Supplement to the "Hymenoptera of 
America North of Mexico — Synoptic Catalog" preserves the form 
of the First Supplement. New supergeneric, generic, subgeneric, 
and trivial names are starred. Abbreviations and symbols agree 
with those used in the Catalog and First Supplement. 

Additions to distribution, hosts, or prey, or corrections to 
entries in the Catalog or First Supplement are made following the 
citation to the generic, subgeneric, or trivial name as it appears in 
the Catalog or First Supplement. Additional references to papers 
on biology, distribution, or taxonomy are made in the same way. 
In the center headings the first page reference is to the Catalog, 
followed by a reference to the First Supplement, if the latter is 
necessary. 

Generic transfers are shown as follows: 

Genus CEPHALGIA Panzer, pp. 9-10, Sup. 1, p. 8 
apicalis (Westwood). Transf. to Acantholyda subg. Acantholyda A. Costa. 

ACANTHOLYDA subgenus ACANTHOLYDA A. Costa 
( = Acan/hofyda A. Costa of Catalog, in part, pp. 7—9) 

apicalis (Westwood). Transf. from Cephalcia Panzer, p. 9 (teste Middle- 
kauff, 1958, loc. cit. supra, p. 77). 

When a species is transferred to a genus or subgenus different 
from the one in which it appears in the Catalog or First Supple- 
ment, all synonyms and other information under it are also trans- 
ferred, unless indicated otherwise. 

New synonymy is shown: 

modesta MacGillivray. Syn. of pini Rohwer. 



INTRODUCTION 5 

pini Rohwer. 

Acantholyda modesta MacGillivray, 1923 (syn. by Middlekauff, 1958, loc. 
cit. supra, p. 85). 

New status is shown as : 

johnsoni johnsoni Viereck, changed status. Reduced to subsp. rank (teste 
Schuster, 1958, loc. cit. supra, p. 46). 

Generic or subgeneric transfer and a change in status may be 
combined : 

SPHAEROPTHALMA subgenus PHOTOPSIS Blake 
( = Pho/ops/$ Blake of Catalog, in part, pp. 753—756) 

amphion (Fox), Transf. to Sphaeropthalnia subg. Photopsioides Schuster. 

♦SPHAEROPTHALMA subgenus PHOTOPSIOIDES Schuster 

amphion amphion (Fox), changed status. Transf. from Photopsis Blake, 
p. 753 and reduced to subsp. rank (teste Schuster, 1958, loc. cit. 
supra, p. 38). 

In the Aculeata, subgeneric assignment may be indicated by 
a citation to the subgeneric name in parentheses following the 
species entry: 

robustior Cockerell (JEumelissodes) 

In many entries here, the compiler of a particular section has 
not agreed with the taxonomic placement or nomenclature used in 
a publication by some other worker. This divergence of opinion is 
expressed by citing the publication under the placement or usage 
here considered to be preferable, followed by an indication of the 
different placement or nomenclature in parentheses. For example: 

Genus EUPELMELLA Masi, p. 512 
Nomenclature: Peck, 1963, loc. cit. supra, p. 492 (Macroneura) . 

In this Second Supplement the citations to the authorities for 
some changes, additions, or deletions have been shortened in order 
to save space. The references for new genera, subgenera, species, 
subspecies, and varieties are cited in full throughout, but fre- 
quently cited works may be referred to only by author, year, and 
page following the first reference. For instance, in the Ichneu- 
monidae, Townes and Townes, 1960. U. S. Natl. Mus. Bui. 216 
(Pt. 2) : i-iv + 1-676 is, after its first citation, referred to as 
Townes and Townes, 1960, followed by particular page references. 
This has been done when this work is the authority for generic 
transfers, deletions, changes in status, or synonymy. References 
to discussions of biology, distribution, or taxonomy from this work 
are cited in the same way. 

Elsewhere in this supplement frequently cited works may be 
cited as loc. cit. supra, where such usage will save space without 
producing ambiguity of reference. For instance, in the Symphyta, 
Middlekauff, 1958. Calif. Univ. Pubs., Ent. 14: 51-174 is cited 
completely under the Family PAMPHILIIDAE. Subsequently it is 
referred to as Middlekauff, 1958, loc. cit. supra, with the page 
reference, where it is the source for generic transfers, changes in 
status, deletions, synonymy, or discussions of biology, distribu- 



6 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

tion, or taxonomy. 

In this supplement, some changes, deletions, or additions are 
made on the authority of the individual who prepared a particular 
section. These entries include no citation to the literature or to 
personal communications from some other worker. For instance, 
in the Symphyta, Genus SIREX Linnaeus, p. 83 of the catalog, 
the information that areolatus (Cresson) is "Adventive in Eastern 
States, emerging from lumber" is teste B. D. Burks, although this 
last phrase does not appear in the text. 

Order HYMENOPTERA 

Taxonomy: Richards, 1956. Hymenoptera, Introduction and Keys to 
Families. Handbooks Ident. Brit. Ins., Roy. Ent. Soc, London 6, 
pt. 1, 94 pp., 197 text figs., 11 pis. — Berland, 1958. Atlas des 
Hymenopteres de France. I, Tenthredes, Parasites, Porte-aiguillon 
(Bethyloides, pars), 155 pp., 14 pis. II, Porte-aiguillon (Bethyl- 
oides (fin), Scolioides, Formicoides, Pompiloides, Vespoides, 
Sphecoides, Apoides), 184 pp., 14 pis. Editions N. Boubee & Cie. — 
Kerrich, 1960. Soc. Brit. Ent. Trans. 14: 1-18 (review systematics 

Biology: Fulmek, 1957. Naturhist. Mus. Wien Ann. 61: 110-227 (aphid 
parasites and predators). — Michener, 1961. Roy. Ent. Soc, London 
Symp. 1: 43-56 (aspects of social polymorphism). — Krombein, 
1962. Ent. Soc. Wash. Proc. 64: 11-19 (parasitism of several 
wasps and bees by acarid mites). — Flanders, 1962. Canad. Ent. 
94: 1133-1147 (host-parasite interactions). — Evans, 1963. Wasp 
Farm, 178 pp., 25 figs, (popular account of the biology of some 
wasps). 

Morphology: Short, 1959. Roy. Ent. Soc, London Trans. Ill: 175-203, 
11 figs, (abdominal musculature). — Daly, 1963. Ent. Soc Amer. 
Ann. 56: 295-306 (thoracic musculature). 

Evolution: Lanham, 1951. Ent. Soc. Amer. Ann. (wing venation of 
Clistogastra). — Brandley, 1958 (1956). Tenth Internatl. Congr. 
Ent. Proc 1: 265-269.— Evans, 1958 (1956). Tenth Internatl. 
Congr. Ent. Proc. 2: 449-457 (social life in wasps). — Evans, 1962. 
Evolution 16: 468-483 (prey-carrying in wasps). 

Suborder SYMPHYTA 

Superfamily MEGALODONTOIDEA 
Family XYELIDAE 

Genus XYELA Dalman, pp. 5—6 

*Xyelatana Benson, 1938. Roy. Ent. Soc, London, Proc, Ser. B, 7: 35. 
(Syn. by Burdick, 1961, loc. cit. infra, p. 322.) 
Tvpe: Xyela longula Dalman. Orig. desig. 
Revision: Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 285-353. 

alberta (Curran). Mont., Wyo. Host: Pinus contorta. 

alni Rohwer. Syn. of minor Norton. 

*alpigena (Strobl). Que., Ont., N. Y., Md., 111., Colo.; Europe. Host: Pinus 

sfvouiis 
Pinicola alpigena Strobl, 1895. Wien. Ent. Zeit, 14: 277. 9, $. 
Xyela brunneiceps Rohwer, 1913 (syn. by Benson, 1962. Brit. Mus. (Nat. 

Hist.) Bui. Ent. 12: 385). 



XYELIDAE 7 

*Xyela middlekauffi. Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 343. $. 
(Syn. by Benson, 1962, loc. cit. supra, p. 385.) 

bakeri Konow. Eastern, Que. south to Fla. and E. Tex.; Ont., Mich,; Western, 
Alta, south to N. Mex., B. C. south to Calif. Hosts: Pinus palustris, 
P. ponderosa, P. sabiniana, P. virginiana. 

Xyela salicis Rohwer, 1913 (syn. by Burdick, 1961, loc cit. supra, p. 325). 

Xyela intrabilis MacGillivray, 1923 (syn. by Burdick, 1961, loc. cit. supra, 
p. 325). 

brunneiceps Rohwer. Syn. of alpigena (Strobl). 

californica Rohwer. 

Taxonomy: Burdick 1961, loc. cit. supra, p. 340. 

*cheloma Burdick. Idaho, Nev., B. C, Wash., Oreg., Calif. Host: Prob. Pinus 

Xyela cheloma Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 336. 9, 5- 

*concava Burdick. Nev., Calif. Host: Pinus monophylla. 

Xyela concava Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 342. 9, $. 

♦deserti Burdick. Calif. Host: Pinus monophylla. 

Xyela deserti Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 347. $, $. 

dissimilis Rohwer. Syn. of minor Norton. 

errans Rohwer. Syn. of minor Norton. 

intrabilis MacGillivray. Syn. of bakeri Konow. 

♦linsleyi Burdick. Idaho, Wash., Calif. Host: 1 Pinus ponderosa. 

Xyela linsleyi Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 344. ?, $. 

'^lunata Burdick. Calif. Hosts: Pinus sabiniana, prob. P. coulteri. 

Xyela lunata Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 334. 9, 5. 

luteopicta Cockerell. Syn. of minor Norton. 

^middlekauffi Burdick. Syn. of alpigena (Strobl). 

minor Norton. Eastern, Que. and Ont. south to E. Tex., Western, Alta. south 

to N. Mex., B. C. south to Calif. Hosts: Pinus coulteri, P. muricata, 

P. ponderosa, P. sabiniana, P. virginiana. 
Xyela luteopicta Cockerell, 1902 (syn. by Burdick, 1961, loc. cit. supra, p. 

330). 
Xyela alni Rohwer, 1913 (syn. by Burdick, 1961, loc. cit. supra, p. 330). 
Xyela dissimilis Rohwer, 1913 (syn. by Burdick, 1961, loc. cit. supra, p. 

330). 
Xyela errans Rohwer, 1913 (syn. by Burdick, 1961, loc. cit. supra, p. 330). 
Xyela nevadensis Rohwer, 1913 (syn. by Burdick, 1961, loc. cit. supra, p. 

330). 
Xyela winnemanae Rohwer, 1913 (syn. by Burdick, 1961, loc. cit. supra, 

p. 330). 
Morphology: Daly, 1963. Ent. Soc. Amer. Ann. 56: 299. 

nevadensis Rohwer. Syn. of minor Norton. 

'*=ob8cura (Strobl). Newfoundland and Ont. south to Fla. and E. Tex.; 

N.W.T. south to Alta., and Mont.; B, C. south to Calif.; Holarctic. 

Hosts: Pinus palustris, P. banksiana, P. ponderosa, P. virginiana. 
Pinicola julU var. obscura Strobl, 1895. Wien. Ent. Zeit. 14 : 277. 9 . 
Xyela pini Rohwer, 1913 (syn. by Benson, 1962, loc. cit. supra, p. 385). 

pini Rohwer. Syn. of obscura (Strobl). 

Xyela slossonae Rohwer, 1913 (syn. by Burdick, 1961, loc. cit. supra, p. 

327). 
Xyela similis Rohwer, 1913 (syn. by Burdick, 1961, loc. cit. supra, p. 327). 

♦priceae Burdick. Calif. Host: Pinus coulteri. 

Xyela priceae Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 339. 9, $. 

*radiatae Burdick. Calif. Host: Prob. Pinus radiata. 

Xyela radiatae Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 337. 9, $. 



8 HYMENOPTERA OF AMERICA NORTH OP MEXICO 

salicis Rohwer. Syn. of bakeri Konow. 

*serrata Burdick. Calif. Host: Pinus rmiricata. 

Xyela serrata Burdick, 1961. Calif. Univ. Pubs. Ent. 17: 341. $. 

similis Rohwer. Syn. of obscura (Strobl). 

slossonae Rohwer. Syn. of obscura (Strobl). 

*styrax Burdick. Md., Va., Ga. Host: Pinus virginiana. 

Xyela styrax Burdick, 1961. Calif. Univ. Pubs., Ent. 17: 345. 9,5. 

winnemanae Rohwer. Syn. of minor Norton. 

Genus PLERONEURA Konow, p. 6 

Taxonomy: Burdick, 1959. Kansas Ent. Soc. Jour. 32: 121 {Pleuro- 
neura ( 1 ) ) . 

Genus XYELECIA Ross, p. 7 
nearctica Ross. Wash. 

Taxonomy: Smith, 1964. Pan-Pacific Ent. 40: 54-56. $. 

Genus MEGAXYELA Ashmead, p. 7 
langstoni Ross. S. C, Ga., Fla. 

Family PAMPHILIiDAE 

Taxonomy, Morphology, Phylogeny: Middlekauff, 1958. Calif. Univ. 
Pubs., Ent. 14: 51-73». 

Genus ACANTHOLYDA A. Costa, pp. 7-9, Sup. 1, p. 8 

Delete Itycorsia Konow from syn. 

Revision: Middlekauff, 1958, loc. cit. supra, pp. 73-132. 

ACANTHOLYDA subgenus ACANTHOLYDA A. Costa 

(=:Acanfho//da A. Costa of Catalog, in part, pp. 7—9) 

abdominalis Norton. Syn. of tesselata (Klug). Transf. montivaga Cresson to 
syn. of veriicalis (Cresson). 

apicalis (Westwood). N. C, Fla., Miss. Host: Pinus taeda. Transf. from 
Cephalcia Panzer, p. 9 {teste Middlekauff, 1958, loc. cit. supra, 
p. 77). 

atripes (Cresson). N. J., Pa., Ga. 

erythrocephala (Linnaeus). Ont. Hosts: Pinus spp. 

marginiventris (Cresson). Mass., N. J., Va., N. C. 

modesta MacGillivray. Syn. of pini Rohwer. 

pini Rohwer. Que., Ont., N. Y., Pa., Md., Va., N. C, Ga., Mich. Host: Pinus 
resinosa. 
Acantholyda modesta MacGillivray, 1923 (syn. by Middlekauff, 1958, 
loc. cit. supra, p. 85). 

poeppigii (Brischke and Zaddach). Pa. 

*runcinata Middlekauff. Tex., Colo., N. M., Ariz. 

Acantholyda (.Acantholyda) runcinata Middlekauff, 1958. Calif. Univ. 
Pubs., Ent. 14: 89. ?, $. 

tesselata (Klug). Que., Conn., Ont., N. Y., N. J., Md., Va., N. C, Mich., Wis., 
Minn. Hosts: Pinus banksiana, P. resinosa. Transf. rufiventris 
Cresson to syn. of verticalis (Cresson). 
Lyda abdominalis Norton 1862 (syn. by Cresson, 1880. Amer. Ent. Soc. 
Trans. 8: 64). 



PAMPHILIDAE 9 

ACANTHOLYDA subgenus ITYCORSIA Konow 
( = Acanf/io/yda A. Costa of Catalog, in port, pp. 7—9) 

*aequorea Middlekauff. Calif. 

Acantholyda (Itycorsia) aequorea Middlekauff, 1958. Calif. Univ. Pubs., 
Ent. 14: 93. ?, $. 

albomarginata (Cresson). Mich., Mont., Alaska, B. C, Wash., Oreg., 
Calif. Hosts: Prob. Pinus rmiricata, P. ponderosa. 

Itycorsia marginiventris Kincaid, 1900 (not Cresson). Misident. 

Itycorsia kincaidi Rohwer, 1910 (syn. by Middlekauff, 1958, loc. cit. supra, 
p. 95). 

angulata (MacGillivray). N. B., Que., Maine, Mass., Ont., Pa., Mich., Minn. 
Hosts: Pinus austriaca, P. rigida, P. densiflora, P. banksiana, P. 
strobus. 

atrata (Cresson). Idaho, B. C, Wash., Oreg., Calif. Hosts: Tsuga hetero- 
phylla, Abies concolor. 
Itycorsia balata MacGillivray, 1923 (syn. by Middlekauff, 1958, loc. cit. 
supra, p. 98). 

*aurigera Middlekauff. Calif. ■ 

Acantholyda ^Itycorsia) aurigera Middlekauff, 1958. Calif. Univ. Pubs., 
Ent. 14: 100. ?. 

balanata MacGillivray. Alaska, B. C, Calif. Hosts: Pseudotsuga menziesii, 
Picea sitchensis, Picea sp., Abies sp., Tsuga sp. 

balata MacGillivray. Syn. of atrata (Cresson). 
ballista MacGillivray. Syn. of terminalis (Cresson). 

bicolorata (Norton). Maine. Delete nigripcs Cresson from syn. 

brunnicans (Norton). Mont., Wyo., Nev., B. C, Oreg., Calif. Hosts: Prob. 

Pinus contorta, P. sabiniana. P. ponderosa, P. radiata. Delete 

olyntpia MacGillivray from syn. 
Lyda discolor Cresson, 1880 (syn. by Middlekauff, 1958, loc. cit. supra, p. 

103). 
Lyda morrisoni Cresson, 1880 (syn. by Middlekauff, 1958, loc. cit. supra, 

p. 103). 
Lyda nevadensis Cresson, 1880 (syn. by Middlekauff, 1958, loc. cit. supra, 

p. 103). 

brunniceps (Cresson). N. B., Que., Mass., Ont., Minn. 

bucephala (Cresson). Mont., B. C, Wash., Oreg., Calif. Hosts: Pseudotsuga 
menziesii, prob. Abies concolor. 
Lyda olympia MacGillivray, 1893 (syn. by Middlekauff, 1958, loc. cit. 
supra, p. 107) . 

*burkei Middlekauff. Calif. Host: Pinus radiata. 

Acantholyda {Itycorsia) burkei Middlekauff, 1958. Calif. Univ. Pubs., 
Ent. 14: 108. $, $. 

chicoutimiensis (Huard). Que., Ont. Hosts: Picea mariana, ? Pinus 
banksiana. 
Nomenclature: Comeau, 1957. Nat. Canad. 84: 29. 

circumcincta (Klug). N. B., Que., Pa. Hosts: Pinus banksiana, prob. other 
sp. of Pinus. 

*crocina Middlekauff. Idaho, Calif. Hosts: Possibly Pinus muricata, P. 
sabiniana. 
Acantholyda ^Itycorsia) crocina Middlekauff, 1958. Calif. Univ. Pubs., 
Ent. 14: 113. ?, $. 

*depressa Middlekauff. Ariz. 

Acantholyda ^Itycorsia') depressa Middlekauff, 1958. Calif. Univ. Pubs., 
Ent. 14: 115. $. 

discolor Cresson. Syn. of brunnicans (Norton). 



10 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

kincaidi Rohwer. Syn. of albomarginata (Cresson). 

luteomaculata (Cresson). N. B., Que., Maine, Vt., Mass., Ont., Pa., Ohio, 
Mich., Wis., Minn. Hosts: Pinus strobus, prob. P. sylvestris, P. 
banksiana. 

luteopicta Rohwer. Syn. of verticalis (Cresson). 

maculiventris (Norton). Labrador, Que. and Maine south to N. C, west to 
B. C. and Calif. Hosts: Abies balsamea, Picea glauca, possibly 
Picea mariana, Pinus strobus. Delete chicoutimiensis (Huard) 
from syn. 

montivaga Cresson. Syn. of verticalis (Cresson). 

morrisoni Cresson. Syn. of brunnicans (Norton). Delete nevadensis Cresson 
from syn. 

nevadensis Cresson. Syn. of brunnicans (Norton). 

nigripes (Cresson). Nev., Oreg., Calif. 

nigrita (Cresson). Calif. 

ochrooera (Norton). Que., Ont., Mich., Wis. 

olympia MacGillivray. Syn. of bucephala (Cresson). 

ruficeps (Harrington). Que., Mass., Ont., N. Y., Mich., Minn., B. C. Hosts: 
Prob. Pinus resinosa, P. contorta, P. banksiana. 

rufiventris Cresson. Syn. of verticalis (Cresson). 

terminalis (Cresson). Colo., Wash., Oreg., Calif. 

Itycorsia ballista MacGillivray, 1923 (syn. by Middlekauff, 1958, loc. cit. 
supra, p. 125). 

*thalictra Middlekauff. Tex., N. Mex. Host: ? Pinus edulis. 

Acantholyda (Itycorsia) thalictra Middlekauff, 1958. Calif. Univ. Pubs., 
Ent. 14: 126. ?. 

verticalis (Cresson). Minn., Nebr., Mont., Wyo., Colo., Alta., Nev., B. C, 

Wash., Oreg., Calif. Hosts: Pinus radiata, prob. P. contorta, P. 

ponderosa, P. monticola. 
Lyda similaris Cresson, 1880 (questionably syn. in Catalog, p. 9). 
Lyda montivaga Cresson, 1880 (syn. by Middlekauff, 1958, loc. cit. supra, 

p. 127). 
Lyda rufiventris Cresson, 1880 (syn. by Middlekauff, 1958, loc. cit. supra, 

p. 127). 
Itycorsia luteopicta Rohwer, 1910 (syn. by Middlekauff, 1958, loc. cit. 

supra, p. 127). 

zappei (Rohwer). Que., N. H., Mass., Ont., N. J., Md., Mich. Hosts: Pinus 
austriaca, P. strobus, P. rigida. 

Unplaced Species of Acantholyda 
variegata Norton. Delete bucephala Cresson from syn. 

Genus CEPHALGIA Panzer, pp. 9-10, Sup. 1, p. 8 
Revision: Middlekauff, 1958, loc. cit. supra, pp. 132-149. 
apicalis (Westwood). Transf. to Acantholyda subg. Acantholyda A. Costa. 

*californica Middlekauff. Mont., ? Wyo., Calif. Hosts: prob. Pinus murray- 
ana, P. ponderosa. 
Cephalcia calif ornica Middlekauff, 1958. Calif. Univ. Pubs., Ent. 14: 
135. 9. 

canadensis (Norton). Transf. to Unplaced species of Pamphiliidae. 

criddlei (MacGillivray). Syn. of fascipennis (Cresson). 

dissipator (MacGillivray). Syn. of fascipennis (Cresson). 
distincta MacGillivray. Syn. of provancheri (Huard). 



PAMPHILIDAE 11 



fascipennis (Cresson). N. B., N. S., Que., Maine, Ont., Man. Hosts: Picea 
pungens, ? P. glaiica, Pinus rigida. 
Cephaleia criddlei MacGillivray, 1912 (syn. by Middlekauff, 1958, loc. cit. 

supra, p. 137). 
Cephaleia dissipator MacGillivray, 1923 (syn. by Middlekauff, 1958, loc. 
cit. supra, p. 137). 

frontalis (Westwood). Ont., N. Y., N. J., Pa., N. C, Ga., Mich. 

Caenolyda onekania MacGillivray, 1923 (syn. by Middlekauff, 1958, loc. 
cit. supra, p. 137). 

fulviceps (Rohwer). Que., N. H., Conn., Ont., Ark. Hosts: Pinus banksiana, 
P. resinosa. 

hopkinsi (Rohwer). Host: Prob. Pinus ponderosa. 

jenseni MacGillivray. Syn. of semidea (Cresson). 

marginata Middlekauff. Conn., W. Va. Hosts: Pinus resinosa, Picea sp. 

mathematica (Kirby). Syn of nigropectus (Cresson). 

♦nigra Middlekauff. N. B., N. S., Maine, Mich. 

Cephaleia nigra Middlekauff, 1958. Calif. Univ. Pubs., Ent. 14: 143. 9. 

nigropectus (Cresson). Newfoundland south to N. C, west to Alaska and 
Calif. Hosts: Picea glauca, prob. other spruces. 

Pumphilius mathematicus Kirby, 1882 (syn. by Middlekauff, 1958, loc. cit. 
supra, p. 144) . 

Caphaleia punctata Rohwer, 1908 (syn. by Middlekauff, 1958, loc. cit. 
supra, p. 144). 

nortoni Rohwer. Syn. of semidea (Cresson). 

onekama (MacGillivray). Syn. of frontalis (Westwood). 

ochreipes (Cresson). Transf. to Pamphilius Latreille. 

provancheri (Huard). N. H., N. Y., Ont. Trans, from Pamphilius, p. 11 

(teste Middlekauff, 1958, loc. cit. supra, p. 146). 
Cephaleia distincta MacGillivray, 1912 (syn. by Middlekauff, 1958, loc. 
cit. supra, p. 146). 

punctata (Rohwer). Syn, of nigropectus (Cresson). 

*rossi Middlekauff. Newfoundland, N. B., Que., Mass., Ont., Mich. Host: 

^ hies l)(lls(Xt7lCQ,w 

Cephaleia rossi Middlekauff, 1958. Calif. Univ. Pubs., Ent. 14: 147. 9. 

semidea (Cresson). N. B., Que., Conn., Ont., N. Y., Mich. Host: ? Picea sp. 
Caenolyda nortoni Rohwer, 1910 (syn. by Middlekauff, 1958, loc. cit. 

supra, p. 148). 
Cephaleia jenseni MacGillivray, 1912 (syn. by Middlekauff, 1958, loc. cit. 

supra, p. 148). 

Genus NEUROTOMA Konow, p. 10, Sup. 1, p. 8 

Revision: Middlekauff, 1958, loc. cit. supra, pp. 149-158. 

crataegi Middlekauff. N. Y., Mich. 

fasciata (Norton). N. H., south to Fla., west to Mich., 111., and Mo. Host: 
Primus pennsylvanica. 

inconspicua (Norton). Mass., N. Y., and Pa., west to B. C, Mont., Kans., 
Ark. Hosts: Prunus spp. 

*willi Middlekauff. Pa. 

Neurotoma tvilli Middlekauff, 1958. Calif. Univ. Pubs., Ent. 14: 158. $. 

Genus PAMPHILIUS Latreille, pp. 10-12 
*?Pseudocephaleia Zirngiebl, 1937. Embrik Strand 60. Geburtst. 
Festschr. (Riga) 3: 339. (Syn. questionably teste Benson, 1945. 
Roy. Ent. Soc, London Proc, ser. B, 14: 31.) 

Type: Pseudocephaleia hrachycerus Zirngiebl. Orig. desig. 



12 HYMENOPTERA OF AMERICA NORTH OP MEXICO 

*PamphiUus subg. Onycholyda Takeuchi, 1938. Tenthredo 2: 218. (Syn. 
teste Benson, 1945, loc. cit. supra, p. 31.) 

Type: Pamphilius sulphureipes Kirby. Orig. desig. 
Revision: Middlekauff, 1964. Calif. Univ. Pubs., Ent. 38: 1-80. 

amplectus (Fabricius). N. J. south to Fla. west to 111. and Tex.; delete Colo, 
from dist. Host: Rubus sp. Delete cavifrons Cresson from syn. 
and transf. to syn. of muhisignatus (Norton). 
Lyda plagiata Klug (syn. teste Middlekauff, 1964, loc. cit. supra, p. 14). 

burquei (Provancher) . Ont., Va. 

cinctus Harrington. Que., N. Y., Mich., Wis. 

dentatus MacGillivray. Syn. of luteicornis (Norton). 

excavatus (Norton). Que. south to Va. west to Mich., Minn., Ohio; B. C. 
Host: Cornus stolonifera. 

fletcheri MacGillivray. Syn. of luteicornis (Norton). 

fortuitus MacGillivray. Syn. of *sitkensis (Kincaid). 

fulvifrons Rohwer. Syn. of *sitkensis (Kincaid). 

greenei Rohwer. Syn. of ocreatus (Say). 

harringtonii (Provancher). Syn. of luteicornis (Norton). 

*infuscatus Middlekauff. Maine, Alta., B. C. Host: Populus tremuloides. 
Pamphilius infuscatus Middlekauff, 1964. Calif. Univ. Pubs., Ent. 38 : 24. 
?, 5. 

luteicornis (Norton). N. B. and N. S. south to Fla. west to Man. and 111., 

Alta. Hosts: Rubus spp. 
Lyda harringtonii Provancher, 1886 (syn. teste Middlekauff, 1964, loc. 

cit. supra, p. 25). 
Pamphilius ocellatus Rohwer, 1910 (syn. teste Middlekauff, 1964, loc. cit. 

supra, p. 25). 
Pamphilius ruhi Rohwer, 1910 (syn. teste Middlekauff, 1964, loc. cit. 

supra, p. 25). 
Pamphilius dentatus MacGillivray, 1912 (syn. teste Middlekauff, 1964, 

loc. cit. supra, p. 26). 
Pamphilius fletcheri MacGillivray, 1912 (syn. teste Middlekauff, 1964, 

loc. cit. supra, p. 26). 
Biology: Fletcher, 1900. Rpt. Ent. Bot. Dom. Expt. Farm Ottawa, Sess. 

Paper 8 a: 180-181 (multisignata) . — Walden, 1913. Conn. Agr. 

Expt. Sta. Ann. Rpt. 36: 236-240 (dentatus). 

muhisignatus (Norton). Ont. west to B. C. south to Kans., and N. Mex. 

Delete melliventris Cresson from syn. and transf. to syn. of 

pacificus (Norton). 
Lyda cavifrons Cresson, 1865 (syn. teste Middlekauff, 1964, loc. cit. 

supra, p. 30). 
*Bactroceros pugnax Rohwer, 1908. N. Y. Ent. Soc. Jour. 16: 103. ?. 

Preocc. (Syn. teste Middlekauff, 1964, loc. cit. supra, p. 30.) 
Anoplolyda saeva Rohwer, 1910, n. name for pugnax Rohwer. 

nigritibialis Rohwer. Que. south to N. C. west to Alta. and Colo. 

Pamphilius transversus MacGillivray, 1912 (syn. teste Middlekauff, 1964, 
loc. cit. supra, p. 33). 

occWatMs Rohwer. Syn. of luteicornis (Norton). 

ochreipes (Cresson). Transf. from Cephalcia Panzer teste Middlekauff, 1964, 
loc. cit. supra, p. 38. Maine and Ont. south to N. C. west to 
Alaska and B. C. Host: Adults taken on Cornus and Corylus. 
Lyda perplexa Cresson, 1880 (syn. teste Middlekauff, 1964, loc. cit. supra, 
p. 38). 

ocreatus (Say). Maine south to N. C. west to Man. and 111. Host: Corylus 
rostrata. 



PAMPHILIDAE 13 

Pamphilius greenci Rohwer, 1913 (syn. teste Middlekauff, 1964, loc. cit. 
supra, p. 35). 

pacificus (Norton). B. C. south to Calif, east to Colo. Host: Rosa calif omica. 
Lyda melliventris Ci'esson, 1880 (syn. teste Middlekauff, 1964, loc. eit. 

supra, p. 40). 
Biology: Middlekauff, 1964, loc. cit. supra, pp. 41-42. 

palachei (Ashmead). 

Taxonomy: Middlekauff, 1964, loc. cit. supra, p. 43. 

pallimaculus (Norton). Que. and Maine south to Ga. west to B. C. and Colo. 
Host: Collected on rose. 

persicum MacGillivray. Que. south to Fla. west to Minn., Nebr., and Tex. 
Pamphilius unalatus MacGillivray, 1920 (syn. teste Middlekauff, loc. cit. 

supra, p. 46). 
Biology and Taxonomy: Middlekauff, 1964, loc. cit. supra, pp. 47-49. 

*phyllisae Middlekauff. Que. south to N. C. west to Mich, and Wis. Host: 
Collected on Cornus. 
Pamphilius phyllisae Middlekauff, 1964. Calif. Univ. Pubs., Ent. 38: 46. 
9, $. 

plagiatus (Klug). Syn. of amplectus (Fabricius). 

provancheri (Huard). Transf. to Cephalcia Panzer. 
Nomenclature: Comeau, 1957. Nat. Canad. 84: 29. 

pullatus (Cresson). Newfoundland south to Va. west to 111. and Mo. Host: 
Viburnum dentatum. 
Taxonomy: Middlekauff, 1964, loc. cit. supra, p. 52 ($). 

quebccensis (Provancher) . Que. and Ont. south to N. C. west to Mich. 
Taxonomy: Middlekauff, 1964, loc. cit. supra, pp. 53-54 {$). 

rileyi (Cresson). Conn, south to W. Va. west to Mich., Iowa, Mo. Delete Ga. 
from dist. Host: Ovipositing on Amelanchier. 
Pamphilius subcavifrons Rohwer, 1910 (syn. teste Middlekauff, 1964, 
loc. cit. supra, p. 55). 

ruhi Rohwer. Syn. of luteicornis (Norton). 

rufocinctus (Cresson). Syn. of rufofasciatus (Norton). 

rufofasciatus (Norton). Newfoundland, P. E. I., Que., and Maine south to 
Ga., M^est to Alta. and Colo. 
Lyda rufocincta Cresson, 1880 (syn. teste Middlekauff, 1964, loc. cit. 
supra, p. 56). 

saevus (Rohwer). N. name for *pugnax Rohwer, syn. of multisignatus 
(Norton). 

scriptus (Say.) Transf. to Unplaced Species of Pamphiliidae. 

semicinctus (Norton). Que. and Maine south to N. C. west to Ind. and 
Tenn. Hosts: Amelanchier canadensis, Amelanchier sp., ? Pyrus 
vialus, Primus virginiana. 
Taxonomy: Middlekauff, 1964, loc. cit. supra, p. 59 ($). 

*sitkensis (Kincaid). Alaska south to Calif., Idaho, Nev. Hosts: Rubiis spp. 
Bactroceros sitkensis Kincaid, 1900. Wash. Acad. Sci. Proc. 2: 344. 9. 
Pamphilius fulvifrons Rohwer, 1910 (syn. teste Middlekauff, 1964, loc. 

cit. supra, p. 61). 
Pamphilius fortuitus MacGillivray, 1923 (syn teste Middlekauff, 1964, 

loc. cit. supra, p. 61). 
Biology: Middlekauff, 1964, loc. cit. supra, pp. 62-64. 

subcavifrons Rohwer. Syn. of rileyi (Cresson). 

transversus MacGillivray. Syn. of nigritibialis Rohwer. 

unalatus MacGillivray. Syn. of persicum MacGillivray. 



14 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

"'uniunguis Middlekauff. N. C, Minn., Man. 

Pamphilius uniunguis Middlekauff, 1964. Calif, Univ. Pubs., Ent. 38: 

65. 5. 

*vernalis Middlekauff. Nebr., Man. 

Pamphilius vernalis Middlekauff, 1964. Calif. Univ. Pubs., Ent. 38: 

66. ?. 

Unplaced Species of Pamphiliidae 

Insert at end of Family Pamphiliidae, p. 12. 

Lyda canadensis Norton. Transf. from Cephalcia Panzer, p. 9 (vide 
Middlekauff, 1964, loc. cit. supra, p. 68). 

Tarpa scripta Say. Transf. from Pamphilius Latreille, p. 12 {vide Middle- 
kauff, 1964, loc. cit. supra, p. 68). 

Superfamily TENTHREDINOIDEA 
Family ARGIDAE 

Taxonomy: Benson, 1963 (1962). Ann. Mag. Nat. Hist., ser. 13, 5: 633- 
635. 

Genus ARGE Schrank, pp. 13-14 

pectoralis (Leach). 

Biology: Schwarz, 1909. Ent. Soc. Wash. Proc. 11: 106-108. 

Genus NEOPTILIA Ashmead, p. 15 
malvacearum (Cockerell). Tex., Ariz. Host: Althaea rosea. 

* Genus SCHIZOCERELLA Forsius 
( = Sofus Malaise of Catalog, p. 16) 

Schizocerella Forsius, 1927. Not. Ent. 7: 19. 

Type: Schizocerella axillaris Forsius. Orig. desig. 
Taxonomy: Benson, 1963, loc. cit. supra, p. 631. 

pilicornis (Holmgren). Introd. into Australia from Amer. 

* Schizocerella srapularis Forsius, 1927. Not. Ent. 7: 19 (syn. by 
Benson, 1963, loc. cit. supra, p. 631). 

Family CIMBICIDAE 

C=Clavellariidae), Townes, 1962. U. S. Natl. Mus. Bui. 216, p. 67. 

Family DIPRIONIDAE 

Genus ZADIPRION Rohwer, p. 19 

rohweri (Middleton). Nev. 

Genus NEODIPRION Rohwer, pp. 19-21, Sup. 1, p. 9 

abbotii (Leach). Host: Pinus elliottii var. densa. 

edwardsii (Norton). Host: Pinus radiata. 

Lophyrus edtvardsii Norton, change date to 1869. 

excitans Rohwer. Southeastern, west to 111. and Tex. 

Biology: Hetrick, 1959. Fla. Ent. 42: 159-162.— Fatzinger, 1964. Jour. 
Econ. Ent. 57: 412. 

fulviceps (Cresson). 

Morphology: Daly, 1963. Ent. Soc. Amer. Ann. 56: 303. 

hypomelas (Rohwer). Colo. 



TENTHREDINIDAE 15 

lecontei (Fitch). 

Biology: Benjamin, 1955. U. S. Dept. Agr. Tech. Bui. 1118, 57 pp. 

*merkeli Ross, Fla. Host: Pinus elliotti var. densa. 

ISeodiprion merkeli Ross, 1961. Ent. Soc. Amer. Ann. 54: 452. ?, $. 

nanulus nanuliis Schedl. Hosts: Pinus echinata, P. rigida. 

Biology: McCormick and Andresen, 1961. Amer. Mus. Novit. 2032, 6 pp. 

uigroscutum Middleton. Mich. 

pratti banksianae Rohwer. 

Biology: Ghent, 1958. Canad. Jour. Zool. 36: 175-183. — Moens and 
Atwood, 1963. Canad. Ent. 95: 779-782. 

pratti pratti (Dyar). N. J. south to N. C. west to Ind. and Ky. Hosts: Pinus 
echinata, P, rigida, P. virginiana. 
Biology: Morris, Schroecler, and Bobb, 1963. Pine Sawfly in Va., Va. 
Div. For., 42 pp.— Bobb, 1963. Jour. Econ. Ent. 56: 618-621. 

sertifer ( Geoff roy). 

Biology: Griffiths, 1959. Canad. Ent. 91: 500-511.— Wallace and Sullivan, 

1963. Canad. Ent. 95: 1051-1066. 
Physiology: Marek, 1963. ceskol. Spolec. Zool. Vest. 27: 115-124. 

swainci Middleton. Mich., Minn. 

Biology: Ghent and Wallace, 1958. Forest Sci. 4: 264-272.— Franklin, 
1959. Ent. News 70: 62. 

taedae linearis Ross. La. Host: Pinus echinata. 

Biology: Coyne, 1959. U. S. Dept. Agr. For. Pest Leaflet 34, 4 pp. 

*warreni Ross. Ark. Host: Pinus echinata. 

ISeodiprion warreni Ross, 1961. Ent. Soc. Amer. Ann. 54: 451. ?, 3. 

DIPRION subgenus DIPRION Schrank, p. 21 

similis (Hartig). 

Biology: Casida, Coppel, and Watanabe, 1963. Jour. Econ. Ent. 56: 
18-24. 

DIPRION subgenus GILPINIA Benson, p. 21 

hercyniae ( Hartig ) . 

Biology: Martineau, 1963. Canad. Ent. 95: 317. 

Family TENTHREDINIDAE 

Subfamily SELANDRIINAE 
Taxonomy: Malaise, 1963. Ent. Tidskr. 84: 159-215 (genera of world). 

Genus HEMITAXONUS Ashmead, p. 22 

dubitatus (Norton). 

Biology: Krombein, 1960. Ent. News 71: 30. 

ANEUGMENUS subgenus ANEUGMENUS Hartig, p. 23 

*padi (Linnaeus). Oreg., widespread in Canada; Europe, Asia Minor, 

Transcaucasus. Hosts : Ferns. 
Tenthredo padi Linnaeus, 1761. Fauna Suec, ed. 2, p. 390. [$.] 
Tenthredo stramineipes Klug (syn. by Malaise and Benson, 1934, Arkiv 

for Zool. 26 A (20) :6). 
Selandria urbis Ross (syn. by Benson, 1962. Brit. Mus. (Nat. Hist.) 

Bui. Ent. 12: 387). 

stramineipes (Klug). Syn. of padi (Linnaeus). 
urbis Ross. Syn. of padi (Linnaeus). 

♦Tribe HEPTAMELINI 
Place in Selandriinae, after genus Aneugmenus Hartig, p. 24. 



16 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Genus HEPTAMELUS Holiday 

Heptamelus Haliday, 1855. Nat. Hist. Rev. 2 (Proc.) : 60. 

Type: Melicerta ochroleuca Stephens. By monotypy. 

*ochroleucus (Stephens). B. C; N. Eurasia. Hosts: Ferns. 

Melicerta ochroleuca Stephens, 1835. Illust. Brit. Ent. Mand., v. 7, p. 95. 
Distribution: Benson, 1962, loc. cit. supra, p. 387. 

Genus SELANDRIA Leach, p. 24 

♦flavens (Klug). Northeast, and Northcent. N. Amer.; Holarctic. Host: 

Carex sp. 
Tenthredo (,Allantus) flavens Klug, 1814. Mag. Ges. Naturf. Fr. Berlin 

8: 48. 9, $. 
Selandridea vanduzeei Rohwer, 1911 (syn. by Benson, 1962, loc. cit. 

supra, p. 387). 

vanduzeei (Rohwer). Syn. of flavens (Klug). 

*wuestneii Konow. Man.; Europe. Host: Carex sp. 

Selandria tviistneii Konow, 1885. Wein. Ent. Zeit. 4: 122. 9, $ 
Distribution: Benson, 1962, loc. cit. supra, p. 387. 

DOLERUS subgenus DOLERUS Panzer, pp. 24-27 

*aspep Zaddach. Conn., Mass., Mich., S. Dak.; Europe, Siberia. Hosts: 

Grasses and sedges. 
Dolerus asper Zaddach, 1859. Prog. Fried. Coll. Konigsb., p. 21. $. 
Dolerus tectus MacGillivray, 1914 (syn. by Benson, 1962, loc. cit. supra, 

p. 389). 

tectus MacGillivray. Syn. of asper Zaddach. 

Genus CALIROA O. Costa, pp. 28-29 
aniygdalina Rohwer. Md., Va., Tenn., Mo. 
lineata MacGillivray. Pa., Md., Va., N. C. 

Genus HETERARTHRUS Stephens, pp. 29-30 

nemoratus ( Fallen ) . 

Taxonomy: Lindquist, 1959. Canad. Ent. 91: 625-627 (larva). 

Genus MESSA Leach, p. 30 
alaskana (Kincaid). Syn. of wuestneii (Konow). 

*wuestneii (Konow). N. W. T., Alaska; Circumpolar. Host: Leafminer on 

Salix sp. 
Fenusa tviistneii Konow, 1894. Wien. Ent. Ztg. 13: 91. 9, $. 
Fenusa alaskana Kincaid, 1900 (syn. by Benson, 1959. Roy. Ent. Soc, 

London, Proc. (B) 28: 90). 
Taxonomy: Benson, 1959, loc. cit. supra, pp. 90-91. 

Genus PROFENUSA MacGillivray, pp. 30-31 

alumna (MacGillivray). Syn. of thomsoni (Konow). 

"'platani Burks. Calif. Host: Platanus racemosa. 

Profenusa platanae (!) Burks, 1957. Ent. News 68: 207. 9, $. 

*thomsoni (Konow). Ont., Wis.; Holarctic. Host: Leafminer on Betula sp. 
Fenusa thomsoni Konow, 1886. Wien. Ent. Ztg. 5: 270. 9. 
Messa alumna MacGillivray, 1923 (syn. by Benson, 1959, loc. cit. supra, 

p. 91). 
Taxonomy: Lindquist, 1959. Canad. Ent. 91: 625-627 (larva, alumna). — 

Benson, 1959, loc. cit. supra, p. 91. 

Genus FENUSA Leach, p. 31 

pusilla (Lepeletier). Wis., Wash., Oreg., Calif. 

Taxonomy: Lindquist, 1959, loc. cit. supra, pp. 625-627 (larva). 



TENTHREDINIDAE 17 

Genus NEFUSA Ross, p. 31 
anita (MacGillivray). Host: Viola sp. 

Genus HOPLOCAMPA Hartig, pp. 33-34 
*brevis (Klug). Ont., Conn., R. I., N. Y.; introd. from Europe. Host: Pear. 
Tenthredo (Allantus) hrevis Klug, 1814. Gesell. Naturf. Freunde Berlin 
Mag. 8: 53. ?. 

testudinea (Klug). N. J. 

Genus CRATEROCERCUS Rohwer, p. 35 

albidovariatus (Norton). Ark. 

Genus NEOPAREOPHORA MacGillivray, p. 36 

litura (Klug). 

Taxonomy: Nielson, 1958. Canad. Ent. 90: 229-234. 

Genus PRISTIPHORA Latreille, pp. 37-39 

Taxonomy and Distribution: Benson, 1962. Brit. Mus. (Nat. Hist). Bui. 
Ent. 12: 394-396 (Holarctic spp.). 

*abbreviata (Hartig). Oreg., Wash., Calif.; Europe. Host: Pear [California 

pear-slug]. 
Nematus abbreviatus Hartig, 1837. Fam. Blatt. und Holzwespen, p. 

205. $. 
Gymnonychus calif ornicus Marlatt, 1896 (syn. by Benson, 1962, loc. cit. 

supra, p. 394). 

*aphania Wong and Ross. N. Y. 

Pristiphora aphanta Wong and Ross, 1960. Canad. Ent. 92: 196. 9. 

*aquilegiae (Vollenhoven) . Ont.; Europe. Hosts: Aquilegia spp. 

ISematus aquilegiae Vollenhoven, 1866. Tidskr. v. Ent. 11: 202. ?. 
Taxonomy: Lindqvist, 1962. Not. Ent. 42: 105. 

bivittata (Norton). 

Taxonomy: Nielson, 1958. Canad. Ent. 90: 229-234. 

*borea (Konow). Labrador, Que., Man., N.W.T.; Circumpolar. 

Lygaeoneniauts boreus Konow, 1904. Ztg. syst. Hym. Dipt, 4: 253. 9, 3- 
Taxonomy: Benson, 1962, loc. cit. supra, p. 395. 

*cadma Wong and Ross. Sask. 

Pristiphora catlma Wong and Ross. 1960. Canad. Ent. 92: 198. 9. 

californica (Marlatt). Syn. of abbreviata (Hartig). 

*carolinensis Marlatt. N. C. 

Pristiphora carolinensis Marlatt, 1896. U, S. Dept. Agr. Div. Ent. Tech. 
Ser. 3:120. $. 

*coactula (Ruthe). Man., Colo, Alaska; Circumpolar. Host: Vaccinium. 
Nematus coactulus Ruthe, 1859. Stett. Ent. Ztg. 20: 307. 9. 
Nematus Winnipeg Norton, 1867 (syn. by Benson, 1962, loc. cit. supra, 
p. 395). 

*elaphita Wong and Ross. Sask. 

Pristiphora elaphita Wong and Ross, 1960. Canad. Ent. 92: 194. 9. 

erichsonii (Hartig). 

Biology: Graham, 1956. For. Sci. 2: 132-160.— Buckner, 1959, Canad. 
Ent. 91: 535.— Drooz, 1960. U. S. Dept. Agr. Tech. Bui. 1212, 
52 pp.— Ives, 1960. Canad. Ent. 92: 668-674.— Turnock, 1960. 
Canad. Ent. 92: 500-516.— Nairn, et al, 1961. Ent. Soc. Manitoba 
Proc. 17: 31.— Ives, 1963. Canad. Ent. 95: 887-892.— Pschorn- 
Walcher, 1963. Ztschr. Angew. Ent. 51: 208. 
Morphology: Wong, 1963. Canad. Ent. 95: 897-921. 

*hucksena Wong and Ross. B. C. 

Pristiphora husksena Wong and Ross, 1960. Canad. Ent. 92: 194. 9. 



18 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

idiota (Norton). 

Taxonomy: Nielson, 1958, loc. cit. supra, pp. 229-234. 

*Iativentris (Thomson). Man., N.W.T.; Circumpolar. 

ISeniatus lativentris Thomson, 1871. Hyn. Scand., v. 1, p. 99. ?, $. 

Distribution: Benson, 1962, loc. cit. supra, p. 395. 

*leechi Wong and Ross. Mont., B. C, Wash. Host: Larix occidentalis. 
Pristiphora leechi Wong and Ross, 1960. Canad. Ent. 92: 193. $, $. 

*leucostoma (Lindqvist). Man., N.W.T.; Lapland. Host: Salix sp. 

Lygaeoneniatus ( Lygaeophora) leucostonia Lindqvist, 1952. Not. Ent. 

32: 108. 9, $. 
Taxonomy: Benson, 1962, loc. cit. supra, p. 394. 

pallipes Lepeletier. Valid name for rufipes of Catalog; refs. and syn. under 
rufipes Lepeletier. 
Nomenclature: Benson, 1962, loc. cit. supra, p. 394. 

*paloma Wong and Ross. 111. 

Pristiphora paloma Wong and Ross, 1960. Canad. Ent. 92: 196. 9. 

*parbcta Wong and Ross. Man. 

Pristiphora parbcta Wong and Ross, 1960. Canad. Ent. 92: 198. $. 

*pseudocoactuIa (Lindqvist). Man.; prob. Circumpolar. 

Lygaeonematus (Lygaeoneniatus) pseudocoactulus Lindqvist, 1952. Not. 

Ent. 32: 115. 9, $. 
Taxonomy: Benson, 1962, loc. cit. supra, p. 395. 

*reuteri (Lindqvist). Man.; prob. Circumpolar. Host: Salix sp. 

* Lygaeonematus arcticola Enslin, 1916. Deut. Ent. Ztschr. Beih., p. 

514. 9, S- Preocc. 
Lygaeonematus (^Lygaeophora) reuteri Lindqvist, 1960. Not. Ent. 40: 

33. 9, $. 
Taxonomy: Benson, 1962, loc. cit. supra, p. 395. 

rufipes Lepeletier, of Catalog. Valid name, pallipes Lepeletier. 

*serrula Wong and Ross. Calif. 

Pristiphora serrula Wong and Ross, 1960. Canad. Ent. 92: 196. 9. 

*staudingeri (Ruthe). Man.; Circumpolar. Host: Salix sp. 

Nematus slaudingeri Ruthe, 1859. Stett. Ent. Ztg. 20: 306. 9, $. 
Distribution: Benson, 1962, loc. cit. supra, p. 394. 

*valvangula Wong and Ross. ? N. Y. 

Pristiphora valvangula Wong and Ross, 1960. Canad. Ent. 92: 194. $. 

*venatta Wong and Ross. Idaho. 

Pristiphora venatta Wong and Ross, 1960. Canad. Ent. 196. 9. 

Winnipeg (Norton). Syn. of coaclula (Ruthe). 

Genus PACHYNEMATUS Konow, pp. 39-41 

Taxonomy and Distribution: Benson, 1962. Brit. Mus. (Nat. Hist.) 
Bui. Ent. 12: 405-406 (Holarctic spp.).— Benson, 1963. Roy. Ent. 
Soc. London Proc, B. 32: 162-164 {rumicis group). 

clibrichellus (Cameron). Host: Car ex. 

*clitellatus (Lepeletier). Labrador, Que.; Europe. Host: Grasses. 
Nematus clitellatus Lepeletier, 1823. Monog. Tenthred., p. 62. 9. 

corticosus MacGillivray. Syn. of vagus (Fabricius). 

diaphanus (Eversman). Syn. of kirbyi (Dahlbom). 

*excisus (Thomson). Greenland; Arctic Europe. 

ISematus excisus Thomson, 1862. Svenska Vetensk. Akad. Ofvers. Forh. 
19: 628. 9. 

extensicornis (Norton). Northern U. S., Canada; Holarctic. 

*Pachynematus foveolatus Konow^, 1903. Zeit. Hym. Dipt. 3: 379. (Syn. 
by Benson, 1962, loc. cit. supra, p. 406). 



TENTHREDINIDAE 19 

*freyi Lindqvist. N. W. T., Yukon; Circumpolar. 

Pachynematus freyi Lindqvist, 1949. Not. Ent, 28: 81. 9. 

*kirbyi (Dahlbom). Holarctic. Host: Carex. 

ISematus kirbyi Dahlbom, 1835, Clavis Hym. Syst., p. 40. $. 
ISematus diaphanus Eversman, 1847 (syn. by Benson, 1958. Roy. Ent. 
Soc. Lond., Handb. Ident. Brit. Ins. 6: 237). 

moerens (Foerster). Host: Gramineae. 

*nevadensis Marlatt. Nev. 

Pachynematus nevadensis Marlatt, 1896. U. S. Dept. Agr. Div. Ent. 
Tech. Ser. 3: 110. $. 

*obductus (Hartig). Northern U. S., Canada; Holarctic. 

ISematus obductus Hartig, 1837. Fam. Blatt. und Holzwesp., p. 201. 9, $. 
ISematus palUventris Cresson, 1880 (syn. by Benson, 1962, loc. cit. supra, 
p. 405). 

palUventris (Cresson). Syn. of obductus (Hartig). 

*parvilabris (Thomson). N. W. T.; prob. Circumpolar. 

ISematus parvilahris Thomson, 1862. Svenska Vetensk. Akad. Ofvers. 

Forh. 19: 638. 9, $. . 
Taxonomy: Benson, 1962, loc. cit. supra, p. 405. 

*rumiois (Linnaeus). Man., Alaska: Circumboreal. Host: Rumex. 
Tenthredo rumicis Linnaeus 1758. Syst. Nat., ed. 10, v. 1, p. 559. 

smithae Ross. N. H.; prob. Circumboreal. 

*vagus (Fabricius). Transcontinental in Northern U. S. and Canada; 

Holarctic. Hosts: Carex, Salix. 
Tenthredo vaga Fabricius, 1781. Spec. Ins., v. 1, p. 415. 
Pachynematus corticosus MacGillivray, 1901 (syn. by Benson, 1962, loc. 

cit. supra, p. 405). 

Genus CROESUS Leach, p. 42 

Taxonomy: Benson, 1963. Ent. Tidskr. 84: 25. 
varus (Villaret). Host: Alnus. 

NEMATUS subgenus PTERONIDEA Rohwer, pp. 42-46 

*boreaIis Marlatt. Greenland. 

ISematus borealis Marlatt in Fox, 1892. Acad. Nat. Sci. Phila. Proc. 
44: 133. $. 

*crassu8 (Fallen). N. W. T. ; Eurasia. Hosts: Betula, Populus, Salix, Rumex. 
Tenthredo crassa Fallen, 1808. Svenska Vetensk. Akad. Handl. 29: 106. 
Distribution: Benson, 1962, loc. cit. supra, p. 404. 

fulvicrus Provancher 

ISematus salicis Ashmead, change to Messa salicis. 

*ISematus salicicola Dalla Torre, 1894. Cat. Hym., v. 1, p. 252. N. name 
for salicis Ashm. 

*jugicoIa Thomson. Man.; N. Europe. Host: Salix sp. 

ISematus jugicola Thomson, 1871. Hym. Scand., v. 1, p. 146. 9. 

mendicus Walsh. Syn. of oligospilus Foerster. 

*oligospilus Foerster. Transcont. in N. Amer.; Holarctic. Host: Salix sp. 
Nematus oligospilus Foerster, 1854. Naturh. Ver. Preuss. Rheinl. Verb. 

11: 284. 9. 
ISematus mendicus Walsh, 1866 (syn. by Benson, 1962, loc. cit. supra, 
p. 405). 

*reticulatus Holmgren. Labrador, N. H., Man., N. W. T., Alaska; Circum- 
polar. Host : Vaccinium. 

ISematus reticulatus Holmgren, 1883. Ent. Tidskr. 4: 143. 9. 

Pontania glinka Kincaid, 1900 (syn. by Benson, 1962, loc. cit. supra, p. 
404). 



20 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Pontania popofiana Kincaid, 1900 (syn. by Benson, 1962, loc. cit. supra, 
p. 404). 

salicisodoratus Dyar. Wis. 

ventralis Say. 

Transf. ISematus latus Cresson to Unplaced Nematinae, p. 54. 
Morphology: Judd, 1954. Ent. Soc. Ont. Ann. Rpt. 85: 32-39. 

*villosus Thomson. N. W. T.; N. Europe. 

ISematus villosus Thomson, 1862. Svenska Vetensk. Akad. Ofvers. Forh. 

19: 629. 9. 
Distribution: Benson, 1962, loc. cit. supra, p. 404. 

Genus PONTANIA O. Costa 
IzzNemafus subgenus Pontania O. Costa of Catalog, pp. 46-48) 

Taxonomy: Benson, 1960a. Brit. Mus. (Nat. Hist.) Bui. Ent. 8: 367- 
384.— Benson, 1960b. Ent. Mo. Mag. 96: 60.— Benson, 1962, loc. 
cit. supra, pp. 403-404 (Holarctic spp.). 

acuminata (Marlatt). Transf. to Phyllocolpa Benson. 

agamus (Rohwer). Transf. to Phyllocolpa Benson. 

amentivorus (Rohwer). Transf. to Amauronematus Konow. 

arctica (MacGillivray) . Alaska, Yukon, N. W. T., Man.; Lapland, Switzer- 
land. Host: Leaf galls on Salix reticulata. Transf. from Euura 
(teste Benson, 1960a, loc. cit. supra, p. 377). 

Euura abortiva MacGillivray, 1919 (syn. by Benson, 1960a, loc. cit. supra, 
p. 377). 

Pontania delicatula MacGillivray, 1919 (syn. by Benson, 1960a, loc. cit. 
supra, p. 377). 

Pontania deminuta MacGillivray, 1919 (syn. by Benson, 1960a, loc. cit. 
supra, p. 377). 

*arctophilae Benson. Man. Host: Ovip. on Salix arctophila. 

Pontania arctophilae Benson, 1960. Brit. Mus. (Nat. Hist.) Bui. Ent. 8: 
373. 9. 

*beckettae Benson. Man. Hosts: Galls on Salix planifolia, S. discolor, S. 
glaiica, S. reticulata. 
Pontania beckettae Benson, 1960. Brit. Mus. (Nat. Hist.) Bui. Ent. 8: 
378. ?. 

bozemani (Cooley). Transf. to Phyllocolpa Benson. 

bruneri Marlatt. Transf. from Euura Newman (teste Benson, 1960a, loc. 
cit. supra, p. 382). 

calif ornica Marlatt. Transf. from Euura Newman (teste Benson, 1960a, loc. 
cit. supra, p. 382). 

*crassipes (Thomson). Alaska, Yukon, N. W. T., Man.; N. Europe, Switzer- 
land. Hosts: Galls on Salix herbacea, S. polaris, S. arctica, S. 
retusa, S. arbuscula, S. myrsinites, S. lapponum. 

Nematus crassipes Thomson, 1871. Hym. Scand. 1: 162. 9. 

Pontania ora Kincaid, 1920 (syn. by Benson, 1960a, loc. cit. supra, 
p. 379). 

Pontania atrata MacGillivray, 1919 (syn. by Benson, 1960a, loc. cit. 
supra, p. 379). 

Pontania lorata MacGillivray, 1919 (syn. by Benson, 1960a, loc. cit. 
supra, p. 379). 

cressoni (Marlatt). Transf. to Phyllocolpa Benson. 
delicatulus (MacGillivray). Syn. of arctica (MacGillivray). 
deminutus (MacGillivray). Syn. of arctica (MacGillivray). 
destrictus (MacGillivray). Syn. of Phyllocolpa excavata (Marlatt). 



TENTHREDINIDAE 21 

^dolichura (Thomson). Man.; N. Europe, Austria, Switzerland. Host: Galls 
on Salix plani folia; many other spp. of Salix recorded in Pale- 
arctic lit. 

Nematus dolichurus Thomson, 1871. Hym. Scand. 1: 164. ?, 5. 

Distribution: Benson, 1962, loc. cit. supra, p. 404. 

glinka (Kincaid). Syn. of Nematus subg. Pteronidea reticulatus Holmgren. 

hoppingi (Ross). Transf. from Euura Newman {teste Benson, 1960a, loc. 
cit. supra, p. 382). 

hospes (Walsh). Transf. from Euura Newman {teste Benson, 1960a, loc. cit. 
supra, p. 382). 

hyalinus (Norton). Syn. of proxima (Lepeletier). 

leavitti (Rohwer). Transf. to Phyllocolpa Benson. 

leucostoma (Rohwer). Transf. to Phyllocolpa Benson. 

marianus (Ross). Transf. to Phyllocolpa Benson. 

marlatti (Ross). Transf. to Phyllocolpa Benson. 

ntegacephalus (Rohwer). Syn. of Phyllocolpa excavata (Marlatt). 

melanosonia (Rohwer). Transf. to Phyllocolpa Benson. 

tnellinus Cresson. Transf. to Phyllocolpa Benson. 

*myrtillifoliae Benson. Man. Host: on Salix myrtillifolia. 

Pontania myrtilUfoliae Benson, 1960. Brit. Mus. (Nat. Hist.) Bui. Ent. 
8: 372. 9. 

nevadensis Cresson. Transf. to Phyllocolpa Benson. 

pallicornis (Norton). Transf. to Phyllocolpa Benson. 

parvus Cresson. Transf. to Phyllocolpa Benson. 

pectoralis (Marlatt). Transf. to Phyllocolpa Benson. 

pepii (Ross). Transf. to Phyllocolpa Benson. 

popofianus (Kincaid). Syn. of Nematus subg. Pteronidea reticulatus Holm- 
gren. 

popuellus (Ross). Transf. to Phyllocolpa Benson. 
populi (Marlatt). Transf. to Phyllocolpa Benson. 

*proxinia (Lepeletier). Northeastern; Europe. Host: Salix fragilis. 

IKematus proximus Lepeletier, 1823. Monog. Tenthred., p. 67. 9. 

Messa hyalina Norton, 1864 (syn. by Benson, 1960a loc. cit. supra, p. 
382). 
quadrifasciatus (MacGillivray) . Syn. of Amauronematus fallax (Lepeletier). 

resinicola Marlatt. Transf. from Euura (teste Benson, 1960a, loc. cit. supra, 
p. 382). 

robustus (Marlatt). Transf. to Phyllocolpa Benson. 

salicispisum (Thomas). Transf. from Euura Newman (teste Benson, 1960a, 
loc. cit. supra, p. 382). 

terminalis (Marlatt). Transf. to Phyllocolpa Benson. 

tundra (Kincaid). Transf. to Phyllocolpa Benson. 

♦Genus PHYLLOCOLPA Benson 

Phyllocolpa Benson, 1960b. Ent. Mo. Mag. 96: 60. 

Type: Nematus leucapsis Tischbein. Orig. desig. (Equals Sect. 
A of Pontania in Benson, 1960a. Brit. Mus. (Nat. Hist.) 
Bui. Ent. 8: 380-381.) 

Place in Nematini after Nematus subg. Pontania 0. Costa, p. 48. 



22 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*aculiserra Lindqvist. Man.; Holarctic. 

Pontania acutiserra Lindqvist, 1949 (1948). Not. Ent. 28: 66. ?, $. 
Taxonomy: Benson, 1960b, loc. cit. supra, p. 60. 
Distribution: Benson, 1962, loc. cit. supra, p. 403. 

acuminata (Marlatt). Transf. from Nematus subg, Pontania 0. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

agama (Rohwer). Transf. from Nematus subg. Pontania 0. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

borealis (Marlatt). N. Y. Transf. from Euura Newman (teste Benson, 1960b, 
loc. cit. supra, p. 60). 

bozemani (Cooley). Transf. from Nematus subg. Pontania 0. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

*coriacea (Benson). Alta.; Holarctic. 

Nematus (Pontania) coriaceus Benson, 1953. Ent. Mo. Mag. 89: 150. 

Taxonomy: Benson, 1960b, loc. cit. supra, p. 60. 

crcssoni (Marlatt). Transf from Nematus subg. Pontania 0. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

excavata Marlatt. Colo., Calif., Alaska; Holarctic. 

Pontania megacephala Rohwer, 1908 (syn. by Benson, 1960a, loc. cit. 

supra, p. 380). 
Pontania destricta MacGillivray, 1923 (syn. by Benson, 1960a, loc. cit. 

supra, p. 380). 
Taxonomy: Benson, 1960b, loc. cit. supra, p. 60. 

leavitti (Rohwer). Transf. from Nematus subg. Pontania O. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

leucostoma (Rohwer). Transf. from Nematus subg. Pontania 0. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

mariana (Ross). Transf. from Nematus subg. Pontania 0. Costa (teste Ben- 
son, 1960b, loc. cit. supra, p. 60). 

marlatti (Ross). Transf. from Nematus subg. Pontania 0. Costa (teste Ben- 
son, 1960b, loc. cit. supra, p. 60). 

melanosoma (Rohwer). Transf. from Nematus subg. Pontania O. Costa 
(teste Benson, 1960b, loc. cit. supra, p. 60). 

mellina (Cresson). Transf. from Nematus subg. Pontania O. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

nevadensis (Cresson). Transf. from Nematus subg. Pontania O. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

pallicomis (Norton). Transf. from Nematus subg. Pontania O. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

parva (Cresson). Transf. from Nematus subg. Pontania 0. Costa (teste Ben- 
son, 1960b, loc. cit. supra, p. 60). 

pectoralis (Marlatt). Transf. from Nematus subg. Pontania O. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

peninsularis Kincaid. Transf. from Amauronematus Konow (teste Benson, 
1960a, loc. cit. supra, p. 382; Benson 1960b, loc. cit. supra, p. 60). 

pepii (Ross). Transf. from Nematus subg. Pontania 0. Costa (teste Benson, 
1960b, loc. cit. supra, p. 60). 

popuella (Ross). Transf. from Nematus subg. Pontania 0. Costa (teste Ben- 
son, 1960b, loc. cit. supra, p. 60). 

populi (Marlatt). Transf. from Nematus subg. Pontania O. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

robusta (Marlatt). Transf. from Nematus subg. Pontania O. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 



TENTHREDINIDAE 23 

terminalis (Marlatt). Transf. from Nematus subg. Pontania 0. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

tundra (Kincaid). Transf. from Nematus subg. Pontania O. Costa (teste 
Benson, 1960b, loc. cit. supra, p. 60). 

Genus EUURA Newman, pp. 48-51 

Taxonomy and Distribution: Benson, 1962, loc. cit. supra, pp. 402-403. 

abortiva MacGillivray. Syn. of Pontania arctica (MacGillivray). 

arctica MacGillivray. Transf. to Pontania O. Costa. 

*atra (Jurine). Que., Ont., Alta.; Europe. Hosts: Feeding in stems of 
Salix spp. 
Pteronus ater Jurine, 1807. Nouv. Meth. Class. Hym., pi. 6, fig. 6b. 9. 
Euura nigra Provancher (syn. by Benson, 1962, loc. cit. supra, p. 403). 

horealis (Marlatt). Transf. to Phyllocolpa Benson. 

bruneri (Marlatt). Transf. to Pontania 0. Costa. 

californica (Marlatt). Transf. to Pontania O. Costa. 

hoppingi Ross. Transf. to Pontania O. Coata. 

hospes (Walsh). Transf. to Pontania O. Costa. 

insularis Kincaid. Syn. of mucronata (Hartig). 

♦lanatae Malaise. Man., Alta.; Lapland. Hosts: Feed inside buds of Salix 
calcicola and S. sp. 
Euura lanatae Malaise, 1920. Ent. Tidskr. 40; 105. 9, $. 
Biology: Benson, 1962, loc. cit. supra, p. 403. 

*mucronata (Hartig). Man., Alaska; Circumboreal. Hosts: Salix spp., feed- 
ing inside buds. 

Nematus (Cryptocampus) mucronatus Hartig, 1837. Fam. der Blatt. 
und Holzwesp., p. 223. 9, $. 

Euura insularis Kincaid (syn. by Benson, 1962, loc. cit. supra, p. 402). 

nigra Provancher. Syn. of atra (Jurine). 
ora Kincaid. Syn. of Pontania crassipes (Thomson). 
resinicola (Marlatt). Transf. to Pontania 0. Costa. 
salicispisum (Thomas). Transf. to Pontania 0. Costa. 

Genus AMAURONEMATUS Konow, pp. 51-53 

Taxonomy and Distribution: Benson, 1962, loc. cit. supra, pp. 396-402 
(Holarctic spp.). 

*abnormis (Holmgren). N. W. T., Alaska; Circumpolar. 

Nematus abnormis Holmgren, 1883. Ent. Tidskr. 4: 148. 9. 
Amauronematus aulatus MacGillivray, 1919 (syn. by Benson, 1962, loc. 
cit. supra, p. 401). 

amentivorus (Rohwer). Trans, from Nematus subg. Pontania O. Costa, (teste 
Benson, 1960, loc. cit. supra, p. 381). 

*amentorum (Foerster). Greenland; Circumboreal. Host: Catkins of Salix 

spp. 
Nematus amentorum Foerster, 1854. Naturh. Ver. Preuss. Rheinl. Verb. 

11: 337. 9. 
Nematus suavis Ruthe, 1859 (syn. by Benson, 1962, loc. cit. supra, p. 396). 

♦arcticola (Dalla Torre). N. W. T., Alaska; Arctic Europe and Siberia. 

Nematus arcticus Thomson, 1871. Hym. Scand., v. 1, p. 134. 9. Preocc. 
Nematus arcticola Dalla Torre. 1894. Cat. Hym., v. 1, p. 207. N. name. 
Amauronematus arcticola Enslin, 1915. Deut. Ent. Ztschr. Beih., p. 389. 

N. name. 
Taxonomy: Benson, 1962, loc. cit. supra, p. 398. 

atrata (MacGillivray). Syn. of Pontania crassipes (Thomson). 



24 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

aulatus MacGillivray. Syn. of abnormis (Holmgren). 
cogitatus MacGillivray. Syn. of stordalensis (Strand). 

*faUax (Lepeletier). Nova Scotia, Ont., Man., Alta., N. W. T., Yukon, 

Alaska; Circumboreal. Host: Salix sp. 
Nematus fallax Lepeletier, 1823. Monog. Tenthred., p. 64. 9, $. 
Pontania quadrifasciuta MacGillivray, 1919 (syn. by Benson, 1962, loc. 

cit. supra, p. 399) . 

*groenIandicus Malaise. Greenland, N. W. T., Alaska; Circumboreal. Host: 
Prob. Salix sp. 
Amauronematus groenlandicus Malaise, 1933. Skr. om Svalb. og Ischavet 
53: 3. ?. 

*hebes Konow. N. W. T. ; prob. Circumpolar. 

Amauronematus hebes Konow, 1907. Akad. Nauk. Fiz. — Mat. Otd. Zap. 
18(3): 17. 9. 

*helleni Lindqvist. Que., Man., N. W. T.; Scandinavia. 

Amauronematus helleni Lindqvist, 1941. Not. Ent. 21: 65. ?, $. 

*hislrio (Lepeletier). Northeastern, west to Colo.; Circumboreal. Host: 
Salix sp. 
Nematus histrio Lepeletier, 1823. Monog. Tenthred., p. 63. 9. 
Nematus luteotergum Norton, 1861 (syn. by Benson, 1962, loc. cit. supra, 
p. 400). 

*hulteni Malaise. Man., N. W. T., Yukon; Siberia. 

Amauronematus variabilis var. hulteni Malaise, 1931. Arkiv for Zool. 
23 A(8): 43. 9, $. 

islandica (Kincaid). Alaska. Erroneously placed as a syn. of peninsularis 
Kincaid in Catalog {teste Benson, 1960a, loc. cit. supra, p. 382). 

*itelmena (Malaise). Man.; Siberia. Host: Salix alexensis. 

Pontopristia itelmena Malaise, 1931. Arkiv for Zool. 23 A (8): 54. 9. 

*Ieptocephalus (Thomson). Man.; Arctic Eurasia. 

Nematus leptocephalus Thomson, 1862. Svenska Vetensk. Akad. 
Forhandl. Of vers. 19: 632. 9. 

*leucolaenus (Zaddach). Ont., Man., Sask., Alta., N. W. T., Yukon, Alaska. 
ISematus leucolaenus Zaddach, 1882. Phys. Okon. Ges. Konigsb. Schr. 

23: 196. 9. 
Pontania unga Kincaid 1900 (syn. by Benson, 1962, loc. cit. supra, p. 402). 

*longicauda (Hellen). Man., B. C; Lapland. 

Nematus ^Amauronematus) longicauda Hellen, 1948. Not. Ent. 27: 
113. 9. 

lorata (MacGillivray). Syn. of Pontania crassipes (Thomson). 

luteotergum (Norton). Syn. of histrio (Lepeletier). 

*microphyes (Foerster). Ont.; Europe. Host: Salix sp. 

Nematus microphyes Foerster, 1854. Naturh. Ver. Preuss. Rheinl. Verh. 
11: 344. 9. 

neglectus (Kirby). Ont., 111., Man., N. W. T., Colo., Yukon, B. C; Lapland, 
Siberia. Host: Salix sp. Transf. from Unplaced Nematinae, p. 
54 (teste Benson 1962, loc. cit. supra, p. 400). 

*nitidlpleuris Malaise. Greenland, Man.; Circumpolar. 

Amauronematus nitidipleuris Malaise, 1931. Arkiv for Zool. 23 A (8) : 
45. 9. 

orbitalis Marlatt. Syn. of viduatus (Zetterstedt) . 

peninsularis (Kincaid). Transf. to Phyllocolpa Benson. Delete islandica 
Kincaid from syn. 

*polaris (Holmgren). N. W. T., Alta. Alaska; Circumboreal. Host: Salix sp. 
Nematus polaris Holmgren, 1883. Ent. Tidskr. 4: 143. 9. 



TENTHREDINIDAE 25 

*pravus Konow. Alaska; N, Eurasia. Host: Salix sp. 

Amauronematus pravus Konow, 1896. Termes. Fuzetek 18: 176. $. 

*sagmarius Konow. Ont., Man., Alta., N. W. T., Alaska; Circumboreal. Host: 
Salix sp. 
Amauronematus sagmarius Konow, 1896. Termes. Fuzetek 18: 177. 5, 5- 

*seinilacteus (Zaddach). B. C; N. and Cent. Europe. Host: Salix sp. 

Nematus semilacteus Zaddach, 1883. Phys. Okon. Ges. Konigsb. Schr, 
24: 148. $. 

*8tordalensis (Strand). N. W. T., Alaska. 

Nematus stordalensis Strand, 1905. Second Norw. Arctic Exped. "Fram" 

Rpt., no. 3, p. 7. 9, $. 
*Nematus marginifer Strand, 1905, loc. cit. supra, p. 8. $. (Syn. by 

Benson, 1962, loc. cit. supra, p. 400.) 
Amauronematus cogitatus MacGillivray, 1919 (syn. by Benson, 1962, loc 

cit. supra, p. 400) . 
Amauronematus varianus MacGillivray, 1919 (syn. by Benson, 1962, loc 
cit. supra, p. 400) . 

suavis (Ruthe). Syn. of amentorum (Foerster). 

*tillbergi Malaise. N. W. T., Alaska; Arctic Europe and Siberia. Hostx 
Salix sp. 
Amauronematus tillbergi Malaise, 1920. Ent. Tidskr. 40: 125. 9. 

unga (Kincaid). Syn. of leucolaenus (Zaddach). 

*variator (Ruthe). Man., N. W. T., Yukon; Circumboreal. Host: Salix sp 
Nematus variator Ruthe, 1859. Stett. Ent. Ztg. 20: 308. ?. 

varianus MacGillivray. Syn. of stordalensis (Strand). 

*viduatus (Zetterstedt) . Ont., Sask., Oreg.; Europe. 

Tenthredo viduata Zetterstedt, 1838. Ins. Lap., v. 1, p. 351. 9. 
Amauronematus orbitalis Marlatt, 1896 (syn. by Benson, 1962, loc. cih. 
supra, p. 402). 

Unplaced Nematinae, pp. 54—55 

*Nematus desmodioides Walsh, 1866. Ent. Soc. Philad. Proc. 6: 257. 9, $. 

[Has been considered the same as N. inquilinus Walsh, 1866; the 
latter is also unplaced.] 

*Nematus latus Cresson, 1880. Amer. Ent. Soc. Trans. 8: 4. ?. 
Nomenclature: Catalog, p. 43, under ventralis Say. 

Nematus neglectus Kirby. Transf. to Amauronematus Konow. 

Pontania salicum Cockerell in Aldrich, 1905. Smiths. Misc. Coll., v. 46, p. 
163. Nam. nud. 

EMPRIA subgenus EMPRIA Lepeletier, pp. 55-56 

^candidata (Fallen). N. Que. and N. W. T. west to Alta. and Yukon; 
Circumpolar. Host: Betula sp. 
Tenthredo candidata Fallen, 1807. Svenska Vetensk. Akad. Handl. 28: 

105. 
Distribution: Benson, 1962, loc. cit. supra, p. 390. 

Genus ALLANTUS Panzer, pp. 58-59 

basalis (Klug). Host: Rose. 

Genus ARDIS Konow, p. 66 

sulcata (Cameron). Delete from Catalog; a misidentification {teste D. R. 
Smith, 1965. Pan-Pacific Ent. 41: 9). 

*brunniventris (Hartig). Holarctic; Ont. south to N. C. west to B. C. and 

Calif. 
Monophadnus brunniventris Hartig, 1837. Fam. Blattw. und Holzw., p. 

274. 9. 
Biology: Middlekauff, 1958. Pan-Pacific Ent. 43: 202 (sulcata Cameron). 



26 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus LAGIUM Konow, p. 69 
atro^-iolaceum (Norton). Wis., Kans. 
tardum (Norton). Kans. 

Genus TENTHREDO Linnaeus, pp. 71-78 

♦colon Klug. Labrador, Que., Ont.; Holarctic. Hosts: Polyphagus. 

Tenthredo (Allantus) colon Klug, 1814. Mag. Gesell. Naturf. Freunde 

Berlin 8: 182. 9, $. 
Tenthredo nigricollis Kirby, 1882 (syn. by Benson, 1962, loc. cit. supra, 

p. 392). 
Tenthredo jocosa Provancher, 1882 (syn. by Benson, 1962, loc. cit. supra, 

p. 392). 
Tenthredo livida of Catalog, not Linnaeus, p. 75 (syn. by Benson, 1962, 

loc. cit. supra, p. 392). 

*alaskana (Enslin). Syn. of devia (Konow). 

*devia (Konow). Alaska; Holarctic. 

Allantus devius Konow, 1900. Ent. Nachr. 26: 124. 9. 

Allantus heraclei Kincaid, 1900 (syn. by Benson, 1959. Roy. Ent. Soc. 

Lond. Proc. (B) 28: 98). Preocc. 
* Allantus alaskanus Enslin, 1910. Rev. Russe d'Ent. 10: 371 (syn. by 

Benson, 1959, loc. cit. supra, p. 98). N. name for heraclei Kincaid. 

jocosa Provancher. Syn. of colon Klug. 

livida of Catalog, not Linnaeus. Syn. of colon Klug. 

nigricollis Kirby. Syn. of colon Klug. 

Unrecognized Species of Tenthredo Linnaeus, p. 78 
heraclei Kincaid. Syn. of devia (Konow). 

Genus MACROPHYA Dahlbom, pp. 79-82 
bifasciata (Say), corrected spelling of bifasciatus of Catalog. 

Superfamily SIRICOIDEA 
Family SIRICIDAE 

Genus SIREX Linnaeus, p. 83 

Biology: Flanders, 1962. Canad. Ent. 94: 1133-1147. 

areolatus (Cresson). Adventive in Eastern States, emerging from lumber. 

behrensii ( Cresson ) . 

Morphology: Daly, 1963. Ent. Soc. Amer. Ann. 56: 301. 

calif ornicus (Ashmead). 

Taxonomy: Benson, 1963. Ent. Mo. Mag. 98: 252 (subsp. of juvencus 
Linnaeus). 

edwardsii Brulle. Ark. 

juvencus (Linnaeus). 

Taxonomy: Benson, 1963, loc. cit. supra, p. 252 (subsp.) 

longicauda Middlekauff. Colo., Idaho, Utah, Oreg.; adventive in Eastern 
States, emerging from lumber. 
Taxonomy: Middlekauff, 1962. Pan-Pac. Ent. 38: 31-32. $. 

varipes Walker. Adventive in Eastern States, and Britain, emerging from 
lumber. 
Taxonomy: Benson, 1963, loc. cit. supra, p. 252 (hybrid of juvencus 
juvencus Linnaeus x juvencus calif ornicus Ashmead). 

Genus UROCERUS Geoffrey, p. 84 
californicus Norton. Adventive in Eastern States, emerg^g from lumber. 



ORUSSIDAE 27 

cressoni Norton. Minn. 

Genus XERIS A. Costa, p. 84, Sup. 1, p. 17 
morrisoni morrisoni (Cresson). Ariz. 
spectrum townesi Maa. Idaho, Ariz. 

Genus TREMEX Jurine, pp. 84-85 

Nomen Nudum in Tremex Jurine 

*Tremex hospes Cockerell, 1889. Colo. Biol. Assn. Rpt. 10: [3]. Nom. nud. 

Family ORUSSIDAE 

Catalog: Guiglia, 1957. Mus. Civ. Stor. Nat. Genova Ann. 69: 48-59 
(supplement). 

Superfamily CEPHOIDEA 
Family CEPHIDAE 

Genus JANUS Stephens, p. 88 
bimaculatus (Norton). Wis., Minn. 
integer (Norton). Wash. 

Suborder APOCRITA 

Superfamily ICHNEUMONOIDEA 
Family BRACONIDAE 

Taxonomy: Hellen, 1958. Soc. Fauna Flora Fennica, Fauna Fennica 4: 
3-37.— Beardsley, 1961. Haw. Ent. Soc. Proc. 17: 333-366.— 
Baltazar, 1962. Pacific Insects 4 (4) : 750-771.— Marsh, 1963. 
Ent. Soc. Amer. Ann. 56: 522-527. 

Morphology: Riegel, 1948. Ent. Soc. Amer. Ann. 41: 439-449 (wings). — 
Iwata, 1959. Kontyu 27: 231-238.— Oeser, 1961. Berlin Zool. Mus. 
Mitt. 37: 70-71. 

Larval taxonomy and anatomy: Short, 1952. Roy Ent. Soc. London 
Trans. 103: 67-84. 

Subfamily APHIDIINAE 

This group has been given family rank by some authors, including Quilis, 
1931 (Eos 7: 25, 27), Stelfox, 1941 (Roy. Irish Acad. Proc. 46 (B) : 126-127), 
Stary, 1960 (Acta Soc. Ent. Cechosl. 57: 238-252) and Mackauer, 1961 
(Beitr. z. Ent. 11: 792-803). 

Taxonomy: Stary, 1958. Acta Faun. Ent. Mus. Natl. Pragae 3: 53-96 + 
12 pis.— Stary, 1960. Acta Soc. Ent. Cechosl. 57: 238-252.— 
Mackauer. 1961. Beitr. z. Ent. 11: 96-154, 792-803. 
Biology: Mackauer, 1961. Ztschr. f. Parasitenk. 20: 576-591. — Stary, 
1962. Acta Soc. Ent. Cechosl. 59: 42-58. 

Genus EPHEDRUS Haliday, p. 91 

Embryology: Ivanova-Kasas, 1956. Ent. Obozr. 35: 245-261. 
Taxonomy: Stelfox, 1941. Roy. Irish Acad. Proc. 46: 125-142.— Stary, 
1958. Acta Faun. Ent. Mus. Natl. Pragae 3: 64-85. 

californicus Baker. B. C. 

Ephedrus neradensis Baker. Remove to syn. under persicae Froggatt. 
Taxonomy: Mackauer, 1963. Canad. Ent. 95: 923. 



28 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

incompletus (Provancher). N. B. Host: Macrosiphum agrimoniella (Ckll.). 

nevadensis Baker. Remove from syn. under californicus Baker and place 
as syn. of persicae Froggatt, teste Mackauer, 1963. Ztschr. f. 
Angew. Ent. 52: 343-354; also Mackauer, 1963. Canad. Ent. 95: 
924. 

nitidus Gahan. Syn. of persicae Froggatt, teste Mackauer, 1963. Ztschr. f. 
Angew. Ent. 52: 343-354. 

*persicae Froggatt. Distribution world-wide. Hosts: Myzus persicae (Sulz.), 
M. cerasi (F.), M. momonis Mats., M. mumecola Mats., M. ligustri 
(Mosley), Brachycaudus persicacola (Boisd.), B. cardui (L.), B. 
amygdalinus (Schout.), iS. klugkisti (Born.), B. helichrysi (Kalt.), 
B. lychnidis (L.), Brevicoryne brassicae (L.), Dysaphis planta- 
ginea (Pass.), D. crataegi (Kalt.), D. devecta (Wlkr.), Macro- 
sipfuim euphorbiae (Thos.), Allocotaphis quaestionis (Born.), 
Toxoptera aurantii (Fonsc), Aphis fabae Scop., A. idaei v. d. Goot, 
A. rubrorum (Born.), A. craccivora Koch, A. medicaginis Koch, 
A. umbrella (Born.), Longiunguis pyrarius (Pass.), Phorodon 
humuli (Schrank), Rhopalosiphum padi (L.), R. maidis Fitch, 
Roepkea marchali (Born.) 

Ephedras persicae Froggatt, 1904. Agr. Gaz. 15: 611-612. 9. 

Ephedras nevadensis Baker, 1909. (Syn. by Mackauer, 1963. Ztschr. 
f. Angew. Ent. 52: 352.) 

Ephedras nitidus Gahan, 1917. (Syn. by Mackauer, 1963, loc. cit. supra, 
p. 352.) 

"Ephedras vidali Quilis, 1931. Eos 7: 72. ?. (Syn. by Mackauer, 1963, 
loc. cit. supra, p. 352.) 

"Ephedras palchellas Stelfox, 1941. Roy. Irish Acad. (B) 46: 133. 9. 
(Syn. by Mackauer, 1963, loc. cit. supra, p. 352.) 

"Ephedras interstitialis Watanabe, 1941. Insecta Matsumurana 15: 139. 
9. (Syn. by Mackauer, 1963, loc. cit. supra, p. 352.) 

"Ephedras imprcssus Granger, 1949. Inst. Sci. Madagascar Mem. (A) 
2: 412. 9, 5. (Syn. by Mackauer, 1963, loc. cit. supra, p. 352.) 

"Ephedras (E.) holmani Stary, 1958. Acta Faun. Ent. Mus. Natl. 
Pragae 3(34): 63, 68-79. 9, $. (Syn. by Mackauer, 1963, loc. 
cit. supra, p. 352.) 

Taxonomy: Mackauer, 1963. Ztschr. f. Angew. Ent. 52: 343-354. 

Genus PRAON Holiday, pp. 91-92 

Taxonomy: Mackauer, 1959. Beitr. z. Ent. 9: 810-865 (revision of 
European species). — Mackauer, 1961. Beitr. z. Ent. 11: 794. 

abjectum ( Haliday ) . Change from ahjectas. 

aguti C. F. Smith. N. B.; Calif. Hosts: Macrosiphum frigidicola (G. and P.), 
Myzus persicae (Sulz.). 
Biology: Sekhar, 1957. Ent. Soc. Amer. Ann. 50: 370-375.— Sekhar, 1959. 
Currect Sci. 28: 333-335.— Sekhar, 1960. Canad. Jour. Zool. 38: 
593-603. 

alaskense Ashmead. Change from alaskensis. 

americanum (Ashmead). Change from americanus. 

nanus Baker. 

Taxonomy: Mackauer, 1963. Canad. Ent. 95: 925-926. 

occidentale Baker. Change from occidentalis. 

Taxonomy: Mackauer, 1963. Canad. Ent. 95: 926-927. 
Biology: Schlinger, 1960. Jour. Econ. Ent. 53: 151-154. 

palitans Muesebeck. Ariz., Calif., Colo., Kans., Nebr., Nev., N. Mex., Utah. 

Hosts: Therioaphis maculata (Buckt.), T. riehmi (Borner). 
Praon exoletam palitans Muesebeck. Mackauer, 1959. Beitr, z. Ent. 9: 

832, 870. 
Propagation: Finney, Putler and Dawson, 1960. Jour. Econ. Ent. 53: 

657. 



BRACONIDAE 29 

Colonization and establishment: Van den Bosch, et. al, 1959. Jour Econ. 
Ent. 52: 136-141.— Barnes, 1960. Jour. Econ. Ent. 53: 1094-1096. 

Biology: Van den Bosch, et al, 1959. Jour. Econ. Ent. 52: 142-154.— 
Schlinger and Hall, 1959. Jour. Econ. Ent. 52: 154-157.— 
Schlinger and Hall, 1960. Ent. Soc. Amer. Ann. 53: 144-159. 

Taxonomy: Mackauer, 1959. Beitr. z. Ent. 9: 829-833, 870. 

pequodorum Viereck. Remove from syn. under simulans (Provancher). Re- 
places simulans of Catalog and authors, not Provancher (teste 
M. J. P. Mackauer, in Hit.). U. S. and south. Canada. Hosts: 
A certho siphon pisum (Harris) ; Macrosiphum euphorbiae 
(Thomas). 

simulans (Provancher). Ont.; Md. Host: Dactynotus rudbeckiae (Fitch). 
Not simulans (Provancher) of authors and Catalog, which is 
really pequodorum Viereck (teste Mackauer, in litt.) . Remove 
pequodorum Viereck from syn. and treat as valid sp. 

unicum C. F. Smith. Change from unicus. Calif. Hosts: Aphis rumicis L., 
Macrosiphum ambrosiae (Thomas), M. rosae (L.), Myzus per- 
sicae (Sn]z.) , Phorodon menthae (Buckt.). 
Biology: Schlinger and Hall, 1960. Ent. Soc. Amer. Ann. 53: 411. — 
Schlinger, 1960. Jour. Econ. Ent. 53: 151-154. 

virginiense Ashmead. Change from virginiensis. 

Genus MONOCTONUS Holiday, p. 92 

*crepidis (Haliday). Que., N. B. 

Aphidius crepidis Haliday, 1834. Ent. Mag. 2: 94. 9, $. 

*Monoctonus paludum Marshall, 1896. In Andre, Spec. Hym. Eur. Alg. 

5: 548. 2. 
Taxonomy: Stary, 1959. Acta Soc. Ent. Cechosl. 56: 246-247. 
Biology: Griffiths, 1960. Bui. Ent. Res. 51: 303-319.— Griffiths, 1961. 

Bui. Ent. Res. 52: 147-161. 

*Genus EUAPHIDIUS Mackauer 

Aphidius subg. Euaphidius Mackauer, 1961. Beitr. z. Ent. 11: 110. 

Type: Aphidius pterocommae Ashmead. Orig. desig. 
Euaphidius Mackauer, changed status. Mackauer, 1961, Beitr. z. Ent. 
11: 801. 

Insert in Aphidiinae following Monoctonus Haliday, p. 92. 

pterocommae (Ashmead). N. B., Mich., Iowa, Utah. Hosts: Pterocomma 
bicoior (Oestl.), Clavigerus populifoliae (Fitch). Transf. from 
Aphidius Nees, p. 94, teste Mackauer, 1961. Beitr. z. Ent. 11: 110. 

Aphidius gregarius Marshall, 1896. In. Andre, Spec. Hym. Eur. Alg. 
5: 569. Syn. by Mackauer, 1961, loc. cit. supra, p. 110. 

Aphidius pterocommae Marshall, 1896. In Andre, Spec. Hym. Eur. Alg. 
5: 578. Syn. by Mackauer, 1961, loc. cit. supra, p. 110. 

Genus XENOSTIGMUS C. F. Smith 

IzzAphidius subgenus Xenos/igmus C. F. Smith, p. 92) 

Raised to generic rank (teste Stary, 1960. Acta Soc. Ent. cechosl. 
57: 238, 247, 251). 

Taxonomy: Mackauer, 1961. Beitr. z. Ent. 11: 801. 

bifasriatus (Ashmead). Calif. 

Taxonomy: Stary, 1960, loc. cit. supra, p. 251. 

APHIDIUS subgenus PROTAPHIDIUS Ashmead, pp. 92-93 

Protaphidius Ashmead (= Coelonotus Foerster, preocc.) is not known 
to occur in North America, teste Stary, 1958. Acta Faun. Ent. Mus. Natl. 
Pragae 3: 90. All species in Catalog are transf. to Pauesia Quills. 



30 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Genus PAUESIA Quilis 

Pauesia Quilis, 1931. Eos 7: 67. 

Type: Pauesia alhuferensis Quilis. Orig. desig. 

Place in Aphidiinae after Xenostigmus C. F. Smith, p. 92. 

Aphidius subg. Protaphidius of Catalog and authors. (Syn. by Stary, 
through Paraphidius Stary, 1961. Acta Ent. Mus. Natl. Pragae 
34: 22). All species listed in Catalog under Aphidius subg. 
Protaphidius, pp. 92-93, are to be transf. to Pauesia and all are 
to be cited as n. combs., except californicus (Ashmead) which was 
transf. to Pauesia by Stary, 1961; all adjectival names are to 
have feminine endings and all author names are to be in paren- 
theses. 

Aphidius subg. Paraphidius Stary, 1958. Acta Faun. Ent. Mus. Natl. 
Pragae 3: 91. 

Type : Aphidius californicus A?hmead. Monob. and orig. desig, 

Paraphidius Stary, 1960. Acta Faun. Ent. Mus. Natl. Pragae 6: 5. 
(Changed status). 

Taxonomy: Stary, 1960. Acta Faun. Ent. Mus. Natl. Pragae 6: 5-44 (re- 
vision of European species). 

calif ornica (Ashmead). Va. ; Ont. 

*ponderosae (Muesebeck). B. C; Calif. Host: Cinara sp. 

Aphidius (Protaphidius) ponderosae Muesebeck, 1958. Ent. News 69: 
141. ?, $. 

Genus APHIDIUS Nees, pp. 92-96 

*lncuhus Schrank, 1802. Fauna Boica, v. 2, p. 173. 

Type: Ichneumon aphidum Linnaeus. By monotypy. 

Incubus Schrank suppressed, and Aphidius Nees placed on Official List 
of Generic Names with Aphidius avenue Haliday as type, Opinion 284, In- 
ternatl. Comm. Zool. Nomencl., October, 1954. 

The groups treated in the Catalog as subgenera of Aphidius are now 
considered valid genera, teste Stary, 1960. Acta Faun. Ent. Mus. Natl. 
Pragae 6: 6.— Stary, 1960. Acta Soc. Ent. Cechosl. 57: 238. 

Embryology: Ivanova-Kasas, 1956. Ent. Obozr. 35: 245-261. 

*aliu8 Muesebeck. Calif. Host: Macrosiphum rosae (L.). 

Aphidius (Aphidius) alius Muesebeck, 1958. Ent. News 69: 142. 
Biology: Schlinger and Hall, 1960. Ent. Soc. Amer. Ann. 53: 405. — 
Schlinger, 1960. Jour. Econ. Ent. 53: 151-154. 

coloratus Baker. 

Taxonomy: Mackauer, 1963. Canad. Ent. 95: 928-930. 

confusus Ashmead. Hosts: Macrosiphum nidbeckiae (Fitch), M. rosae (L.). 
Biology: Schlinger and Hall, 1960. Ent. Soc. Amer. Ann. 53: 405.— 
Schlinger, 1960. Jour. Econ. Ent. 53: 151-154. 

delicatus Baker. 

Taxonomy: Mackauer, 1963. Canad. Ent. 95: 928. 
ferruginosus Baker, changed status. 

Taxonomy: Mackauer, 1963. Canad. Ent. 95: 930-931. 

*matricariae Haliday. Holarctic. Hosts: Aphis fahae Scop., .4. gossypii 
Glov., A. helianthi Monell, A. nasturtii Kalt., A. rumicis L., A. 
umbrella (Borner), Amiraphis crataegifoliae (Fitch), Aula- 
corthiim solani (Kalt.), Brarhycaudus amygdalinus (Schout.), B. 
cardui (L.), B. helichrysi (Kalt.), B. lychnidis (L.), Calaphis sp., 
CapitopJwrus sp., Dactynottis rudbeckine (Fitch), Galiobmm 
langei Borner, Liosomaphis berberidis (Kalt.), Macrosiphoniella 
abrotani (Wlkr.), Macrosiphum rosae (L.), Metopolophium dir- 
hodum. (Wlkr.), Myzus ajugae Schout., M. certus (Wlkr.), M. 
ornatus Laing, Nasonovia ribis-nigri (Mosl.), Neomyzus circum- 
flexus (Buckt.), Ovatus sp.. Phorodon humuli (Schrank), 



BRACONIDAE 31 

Rhopalosiphum padi (L.), Sappaphis mali (Ferr.), S. plantaginea 

(Pass.), Schizaphis graminum (Rond.). 
*Aphidius cirsii Haliday, 1834. Ent. Mag. 2: 101. 9. Preocc. (Syn. by 

Schlinger and Mackauer, 1963. Ent. Soc. Amer. Ann. 56: 648.) 
Aphidiua matricariae Haliday, 1834. Ent. Mag. 2: 103. 9, $. 
*Aphidiu8 arundinis Haliday, 1834. Ent. Mag. 2: 104. ?. (Syn. by 

Schlinger and Mackauer, 1963, loc. cit. supra, p. 648.) 
Aphidius phorodontis Ashmead, 1889 (1888). U. S. Natl. Mus. Proc. 11: 

658, 662. 9, $. (Syn. by Schlinger and Mackauer, 1963, loc. cit. 

supra, p. 648.) 
*Aphidius polygoni Marshall, 1896. In Andre, Spec. Hym. Eur. Alg., v. 

5, p. 602. 9. (Syn. by Schlinger and Mackauer, 1963, loc. cit. 

supra, p. 648.) 

* Aphidius lychnidis Marshall, 1896. In Andre, Spec. Hym. Eur. Alg., v. 

5, p. 607. $. (Syn. by Schlinger and Mackauer, 1963, loc. cit. 
supra, p. 648.) 

* Aphidius valentinus Quilis, 1931. Eos 7: 46. 9. (Syn. by Schlinger 

and Mackauer, 1963, loc. cit. supra, p. 648.) 

* Aphidius affinis Quilis, 1931. Eos. 7: 48. 9, 5- (Syn. by Schlinger 

and Mackauer, 1963, loc. cit. supra, p. 648.) 

* Aphidius renominatus Hincks, 1943. Ent. Monthly Mag. 79: 44. N. 

name for A. cirsii Haliday, preocc. (Syn. by Schlinger and 

Mackauer, 1963, loc. cit. supra, p. 648.) 
Aphidius nigriteleus C. F. Smith, 1944. Aphidiinae of No. Amer., pp. 51, 

61. 9, $. (Syn. by Schlinger and Mackauer, 1963, loc. cit. supra, 

pp. 648-649.) 
Taxonomy, biology, distribution, hosts: Schlinger and Mackauer, 1963. 

Ent. Soc. Amer. Ann. 56: 648-653. 

nigripes Ashmead. N. B., N. C, Kans., Minn., Calif. Hosts: Macrosiphum 
avenae (F.), Aphis nasturtii Kalt., Dactynotus ambrosiae 
(Thomas). 
Biology: Schlinger, 1960. Jour. Econ. Ent. 53: 151-154. 

nigriteleus C. F. Smith. Syn. of matricariae Haliday. 

ohioensis C. F. Smith. N. B. Host: Dactynotus rudbeckiae (Fitch). 

pisivorus C. F. Smith. Syn. of pulcher Baker. 

phorodontis Ashmead. Syn. of matricariae Haliday. 

polygonaphis (Fitch). N. B. 

pulcher Baker. North Amer. 

Aphidius pisivorus C. F. Smith. (Syn. by Mackauer, 1963, Canad. Ent. 

95: 931-932.) 
Taxonomy: Mackauer, 1963, loc. cit. supra, pp. 931-932. 

ribis Haliday. B. C. 

Biology: Schlinger and Hall, 1960. Ent. Soc. Amer. Ann. 53: 407. 

rosae Haliday. Delete from Catalog. I have seen no authentically identified 
specimens of this species from No. America. The records given 
under rosae in the Catalog apparently apply to nigripes Ashmead. 

*salicis Haliday. Europe; N. B., Ont., Calif. Host: Cavariella aegopodii 
(Scop.) 
Aphidius salicis Haliday, 1834. Ent. Mag. 2: 102. 9, $. 

*6mithi Sharma and Subba Rao. India; Calif., Wash., Idaho, Colo., Kans., 
Wis., Hawaii. Host: Acyrthosiphon pisum (Harris). 

Aphidius (Aphidius) smithi Sharma and Subba Rao, 1959 (1958). Indian 
Jour. Ent. 20: 183. 9, $. 

Biology: Hagen and Schlinger, 1960. Calif. Agr. 14: 5. — Wiackowski, 
1962. Polskie Pismo Ent. 32: 253-310. 

Genus LYSAPHIDUS C. F, Smith 
{ = Aphidius subgenus Lysaphidus C. F. Smith, p. 95) 

Raised to generic rank (teste Stary, 1960. Acta Soc. Ent. Cechosl. 57: 



32 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

238, 248, 250). Author names of the contained species are placed in paren- 
theses. 

Taxonomy: Narayanan, et al., 1962, Beitr. z. Ent. 12: 676-677. 

adelocarinus (C. F, Smith). N. B., Calif. Hosts: Aphis helianthi Monell, 
CapitopJiorus shepherdiae G. and B. 

rosaphidis (C. F. Smith). N. B. 

Genus LYSIPHLEBUS Foerster 
i^Aphidius subgenus Lysiphlebus Foerster, pp. 95—96) 

Raised to generic rank (teste Mackauer, 1960. Portici Lab. Ent. Agr. 
Bol. 18: 230-236.— Stary, 1960. Acta Soc. Ent. cechosl. 57: 238, 248, 250). 
Place in parentheses author names of knowltoni and utahensis and remove 
parentheses from author of flavidus. 

Taxonomy: Mackauer, 1960, loc. cit. supra, pp. 230-236. — Mackauer, 
1960. Beitr. z. Ent. 10: 582-623.— Stary, 1961. Polskie Pismo Ent. 
31: 97-103. 

chrysoaphidis C. F. Smith. Syn. of testaceipes (Cresson). 

*distinctus Mackauer. Nev. Host: Microsiphum acophorum S. & K. 

Lysiphlebus (^Lysiphlebus) distinctus Mackauer, 1960. Portici Lab. Ent. 
Agr. Bol. 18: 236. 9, $. 

fuscicornis (Ashmead). Transf. from Diaeretus Foerster, p. 97, teste 
Mackauer, 1960. Portici Lab. Ent. Agr. Bol. 18: 240. 

salicaphls (Fitch). Transf. from Diaeretus Foerster, p. 97, teste Mackauer, 
1960. Portici Lab. Ent. Agr. Bol. 18: 243. 

testaceipes (Cresson). South. Canada, all Continental U. S.; Mexico, West 
Indies, Central and S. America, Hawaii. Hosts: Aphis fabae 
Scop., A. pseudohederae Theob., A. spiraecola Patch, Rhopalo- 
siphum maidis (Fitch). 
Aphidius (Lysiphlebus) chrysoaphidis C. F. Smith, 1944. Aphidiinae 
of No. Amer., pp. 77, 78. 9, $. Syn. by Mackauer, 1960. Portici 
Lab. Ent. Agr. Bol. 18: 245. 
Biology: Schlinger and Hall, 1960. Ent. Soc. Amer. Ann. 53: 407-409. 

TRIOXYS subgenus TRIOXYS Holiday, pp. 96-97 

utilis Muesebeck. Ariz., Calif., Nev., N. Mex., Utah; Mexico, Iraq, Iran, 

Israel, Turkey. Hosts: Therioaphis maculata (Buckt.), T. riehmi 

(Borner) . 
Propagation: Finney, Putler, and Dawson, 1960. Jour. Econ. Ent. 53: 

658. 
Distribution and importance: Van den Bosch, 1957. Jour. Econ. Ent. 

50: 353-356. 
Colonization and establishment: Van den Bosch, et al., 1959. Jour. Econ. 

Ent. 52: 136-141.— Barnes, 1960. Jour. Econ. Ent. 53: 1094-1096. 
Biology: Van den Bosch, et al, 1957. Jour. Econ. Ent. 52: 142-154. — 

Schlinger and Hall, 1959. Jour. Econ. Ent. 52: 154-157.— 

Schlinger and Hall, 1961. Ent. Soc. Amer. Ann. 54: 34-45.— Hall, 

Schlinger and Van den Bosch, 1962. Ent. Soc. Amer. Ann. 55: 

566-568. 
Taxonomy: Mackauer, 1959. Beitr, z. Ent. 9: 168, 872.— Hall, Schlinger 

and Van den Bosch, 1962. Ent. Soc. Amer. Ann. 55 : 566-588. 

TRIOXYS subgenus ACANTHOCAUDUS C. F. Smith, p. 97 

♦schlingeri Muesebeck. B. C. (near Lytton). Host: Macrosiphum ambrosiae 
(Thomas). 
Trioxys (Acanthocaudus) schlingeri Muesebeck, 1958. Ent. News 69: 
144. 2, $. 

Genus DIAERETUS Foerster, p. 97 

Delete from Catalog. No North American species known. Transf. rapae 
(M'Intosh) to Diaeretiella Stary, with chenopodiaphidis (Ashmead) as syn. 



BRACONIDAE 33 

under rapae; transf. fuscicornis (Ashmead) and salicaphis (Fitch) to 
Lysiphlebus Foerster. 

♦Genus DIAERETIELLA Stary 

Diaeretus of authors, not Foerster. 

Diaeretiella Stary, 1960. Acta Soc. Ent. cechosl, 57: 242, 248. 

Type: Aphidius rapae M'Intosh. Orig. desig. 
Insert in Aphidiinae after Lysiphlebus Foerster, p. 96. 
Taxonomy: Stary, 1961. Acta Ent. Mus. Natl. Pragae 34: 383-397.— 
Mackauer, 1961. Beitr. z. Ent. 11: 801. 

rapae (M'Intosh). Distribution virtually world-wide. Hosts: Cuemavaca 
noxius (Moi'dv.), Aphis acanthi Schrank, A. rumicis L., A. 
nasturtii Kalt., Calaphis betulaecolens (Fitch), Euceraphis^ 
betulae (Koch), Hayhurstia atriplicis (L.), Hyadaphis foeniculi 
(Pass.), Macrosiphiim euphorbiae (Thos.), Rhopalosiphum maidis 
(Fitch), Myzaphis beibienkoi Narzyk., Schizaphis scirpi (Kittle). 
Lipolexis chenopodiaphidis Ashmead. (Syn. by Stary, 1961. Acta Ent. 

Mus. Natl. Pragae 34: 385.) 
*Diaeretus nipponensis Vlereck, 1911. U. S. Natl. Mus. Proc. 40: 182. 
$,9. (Syn. by Stary, 1961, loc. cit. supra, p. 385). 

* Diaeretus napus Quilis, 1931. Eos 7: 71. 2. (Syn. by Stary, 1961, loc. 

cit. supra, p. 385.) 

* Aphidius affinis Quilis, 1931. Eos 7: 48. 9, $. (Syn. by Stary, 1961, 

loc. cit. supra, p. 385.) 
*Diaeretus plesiorapae Blanchard, 1940. Rev. Chil. Hist. Nat. 44: 45. 

9, $. (Syn. by Stary, 1961, loc. cit. supra, p. 385.) 
*Diaeretus aphidum Mukerji and Chaterjee, 1950. Roy. Ent. Soc, London 

Proc. (B) 19: 4. 9, $. (Syn. by Stary, 1961, loc. cit. supra, p. 

385.) 

♦Genus BOREOGALBA Mackauer 

Boreogalba Mackauer, 1962. Canad. Ent. 94: 1107. 

Type: Boreogalba gladifer Mackauer. By monotypy and 
orig. desig. 

Insert in Aphidiinae after Diaeretus Foerster, p. 97. 

♦gladifer Mackauer. Que. (Great Whale River). 

Boreogalba gladifer Mackauer, 1962. Canad. Ent. 94: 1108. 9. 

Subfamily EUPHORINAE 
Tribe METEORINI 

Genus METEORUS Haliday, pp. 98-101 

acronyctae Muesebeck. Host: Halisodota ingens Hy. Edw. 

angustipennis Muesebeck. W. Va. 

argyrotaeniae Johansen. Wis. 

autographae Muesebeck. Host: Autoplusia egena (Gn.). 

bakeri Cook and Davis. Wis. Host: Dasychira plagiata (Wlkr.). 

campestris Viereck. Maine. 

♦clinophthalmicus Shenefelt. Wis. (Nekoosa), 

Meteorus clinophthalmicus Shenefelt, 1961. Ent. Soc. Wash. Proc. 63: 
276. 9, $. 

dimidiatus (Cresson). Hosts: Epizeuxis sp., Platynota stultana Wlsm. 

fumipennis Muesebeck. Ariz. 

hypophlaei Cushman. Idaho. 

indagator (Riley). Host: Dioryctria auranticella (Grote). 



34 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

laphygmae Viereck. N. Mex.; Costa Rica, Columbia, Venezuela. Host: 
Prodenia eridania (Cram.). 

levis Muesebeck. Calif. 

maximus Muesebeck. S. C, Ga. 

niveitarsis (Cresson). Maine, Mass., Wis., Oreg., Alaska. Host: Eulype 
hastata (L.). 

*piiiifolii Mason. Calif., B. C, Alta. Host: Recurvaria milleri Busck. 
Meteorus pinifolii Mason, 1960. Canad. Ent. 92: 143. 9, $. 

polittLs (Provancher). Mich., Minn. 

proximus (Cress.). Minn. 

tauricornis (Provancher). Md., Fla. 

tersus Musebeck. Host: Amorbia essigana Busck. 

Biology: Fleschner, Ricker, and Johnson, 1957. Calif. Avocado Soc. 
Ybk. 41: 115. 

tibialis Muesebeck. Ont. Host: Hyperplatys aspersa (Say). 

trachynotus Viereck. N. S., Maine, Tenn., Mich., Wis., Minn., Iowa, Colo. 
Distribution apparently includes all U. S. Hosts: Dioryctria 
reniculella (Grote), Choristoneura pinus Freem., Archips rosanus 
(L.), Epismus argutamis (Clem.). 
Biology: McGugan, 1955. Canad. Ent. 87: 178-187. 

versicolor ( Wesmael ) . 

Biology: Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, p. 114.— 
Dowden, 1962. U. S. Dept. Agr., Agr. Handb. 226, p. 60. 

*Tribe EUPHORINI 
Biology: Muesebeck, 1963. Ent. Soc. Wash. Proc. 65: 306 (host relations). 

Genus ARiDELUS Marshall, p. 98 
melanderi (Brues). N. Y., Tenn. 
nigrithorax Muesebeck. S. C. 

Genus PERILITUS Nees, p. 101 

Taxonomy and host relations: Smith, 1953. Ohio Jour. Sci. 53 (3): 
173-178. 

coccinellae (Schrank). Hosts: Coccinella novemnotata Hbst., C. septempunc- 
tata L., C. quinquepunctata L., Adonia variegata (Goeze), Halyzia 
quattuordecimguttata Balduf, Coleomegilla maculata lengi Timb. 
Biology: Stich, 1929. Ztschr. f. wiss. Insektenbiol. 24: 91-95.— Ibrahim, 
1955. Soc. Ent. d'Egypte Bui. 39: 414-416.— Hudon, 1959. Canad. 
Ent. 91:63-64.— Walker, 1961. Ent. Monthly Mag. 97: 240-244. 

*Genus ROPALOPHORUS Haliday 

Ropalophorus Haliday, 1840. In Westwood, Introd. Mod. Classif. Ins. 2, 
Synopsis p. 61. 

Type: Microctonus clavicornis Wesmael. By monotypy and 
orig. desig. 

Insert in Euphorinae following Perilitus Nees, p. 101. 

*wisconsinensis Shenefelt. Wis. (Nekoosa). 

Ropalophorus wisconsinensis Shenefelt, 1960. Ent. Soc. Amer. Ann. 53: 
543. 9. 

*Genus STREBLOCERA Westwood 

Streblocera Westwood, 1833. Phil. Mag. and Jour. Sci. (3) 3: 342. 

Type: Streblocera fulviceps Westwood. By monotypy. 
*Eutanyceru8 Foerster, 1862. Naturh. Ver. Rheinlande Verb. 19: 251. 

Type : Eutanyceru* halidayanus Foerster. By monotypy. 



BRACONIDAE 35 

*Lecythodella Enderlein, 1912. Arch. f. Naturgesch. 78A (2) : 41. 
Type: Lecythodella garleppi Enderlein. By monotypy. 

Place in Euphorinae after Perilitus Nees, p. 101. 

Taxonomy: Muesebeck, 1936. U. S. Dept. Agr. Misc. Pub. 241, pp. 13-14. 
— De Saeger, 1942. Rev. de Zool. et de Bot. Africaines 35: 
376-385.— Watanabe, 1942. Insecta Matsumurana 16: 1-12.— De 
Saeger, 1946. Exploration du Pare National Albert, fas. 50, pp. 
144-178. — Granger, 1949. Inst. Sci. Madagascar, Mem., Ser. A., 
Biol. Anim., v. 2, pp. 318-320.— Walley, 1963. Canad. Ent. 95: 
999-1001. 

*pulvillicornis Walley. Que. (Gatineau Park). 

Streblocera pulvillicornis Walley, 1963. Canad. Ent. 95: 999. 9. 

Genus MICROCTONUS Wesmael, p. 102 

Taxonomy and host relations: Smith, 1953. Ohio Jour. Sci. 53 (3): 
173-178. 

aethiops (Nees). N. J., Fla., Ariz. Hosts: Hypera nteles (F.), H. nigrirostris 
(F.), H. postica (GylL), Sitona crinita Hbst., S. hispidula (F.), S. 
humeralis Steph., S. spp. 
Biology: Loan and Holdaway, 1961. Canad. Ent. 92: 1057-1079.— Coles 
and Putler, 1963. Jour. Econ. Ent. 56: 609-610. 

carabivonis Muesebeck. Md., Tenn., Mich., Iowa, Idaho. 

*invictus Muesebeck. Calif., Ariz. Host: Trigonoscuta s^. 

Microctonus invictus Muesebeck, 1961. Brooklyn Ent. Soc. Bull. 56: 59. 

9, $. 

*pacIiylobii Muesebeck. La., Fla. Host: Pachylobius picivorus (Germar). 

Microctonus pachylohii Muesebeck, 1961. Brooklyn Ent. Soc. Bull. 56: 
58. 2, 5. 

*8itonae Mason. Ont. Host: Sitona scissifrons Say. 

Microctonus sitonae Mason, 1960. Canad. Ent. 92: 436. 9, $. 
Biology: Loan, 1963. Ent. Soc. Amer. Ann. 56: 605-611. 

vittatae Muesebeck. Apparently distributed over most of U. S. and south. 
Canada; also in Hawaii, Japan and Korea. Hosts: Phyllotreta 
striolata discedens Weise, P. s. lineolata Chitt., P. bipustulata 
(F.). 

Taxonomy: Watanabe, 1955. Mushi 29 (8) : 51-55. 

Biology: Nagasawa, 1943. Mushi 15: 103-107. — Nagasawa, 1943. Shoku- 
butsu oyobi Dobutsu 11: 460-466 (In Japanese). — Nagasawa, 
1947. Seibutsu 2: 48-52.— Nagasawa, 1948. Bui. Takarazuka In- 
sectarium 58, 18 pp. (In Japanese). — Smith and Peterson, 1950. 
Jour. Econ. Ent. 43: 581-585.— Smith, 1952. Univ. Calif. Pub. 
Ent. 9: 315-344.— Smith, 1953. Ohio Jour. Sci. 53 (3) : 173-178. 

Genus WESMAELIA Foerster, p. 102 

pendula Foerster. Calif., Alaska. Host: Nabis sp. 

Biology : Muesebeck, 1963. Ent. Soc. Wash. Proc. 65 : 306. 

Genus MYIOCEPHALUS Marshall, p. 102 

boops (Wesmael). Maine, Mich., Minn., Tex., N. Mex., Calif. Distribution 
apparently includes all U. S. 

Genus SYNTRETUS Foerster, pp. 102-103 

Biology: Cole, 1959. Ent. Monthly Mag. 95: 18-21. First record of host 
association. Hosts are adults of Ichneumonidae. 

Genus EUPHORIELLA Ashmead, p. 103 

sommermanae Muesebeck. 

Biology: Sommerman, 1956. Ent. Soc. Wash. Proc. 58 (3) : 149-152 



36 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus EUPHORIANA Gahan, p. 103 
uniformis Gahan. Iowa, Utah, Calif. Host: Lygus elisus Van Duzee. 

*Genus LEIOPHRON Nees 
i — Euphorus Nees, pp. 103—104) 

Leiophron Nees, 1818. Nova Acta Acad. Leop. Carol. 9 : 103. No species. 
Leiophron Nees, Haliday, 1833. Ent. Mag. 1 : 263. Three species. 

Type: Leiophron apicalis (Curtis) Haliday. Desig. by Vier- 
eck, 1914. 

Replaces Euphorus Nees, which falls as a syn. of Leiophron (teste 
Muesebeck, 1958. U.S. Natl. Mus. Proc. 107: 410-412). 

pallipes (Curtis). Sask. Hosts: Adelphocoris lineolatus (Goeze), Chlamy- 
datus sp., Lygus lineolaris (Beauv.), P lag io gnat lius medicagus 
Arrand. 

tuberculatus (Muesebeck). N. H. 

Subfamily COSMOPHORINAE 

Placement of this group in the cyclostomine section of the Braconidae 
seems not to be tenable. The clypeus, while impressed, is not circularly emar- 
ginate as in cyclostomines, and in habits and larval structure the species 
resemble the Euphorinae. For the present the group is treated here as a 
distinct subfamily, but it is removed from the cyclostomine section of the 
family and placed after the Euphorinae. It may eventually need to be rein- 
corporated in that subfamily. 

Taxonomy: Hedqvist, 1955. Ent. Tidskr. 76: 92-98.— Watanabe, 1960. 
Insecta Matsumurana 23: 55. 

Biology: Hedqvist, 1955, loc. cit. supra, p. 92. 

Genus COSMOPHORUS Ratzeburg, Sup. 1, p. 36 

pityophthori Rohwer. Colo., Idaho, Wash. Host: Pityophthorus nitidulus 
(Mann.). 

Subfamily MACROCENTRINAE 

Genus MACROCENTRUS Curtis, pp. 104-107 

Taxonomy: Eady and Clark, 1964. Ent. Gaz. 15: 97-127 (revision of the 
European spp.). 

aegeriae Rohwer. Syn. of marginator (Nees). 

affinis Muesebeck. Minn., Idaho. 

ancylivorus Rohwer. Host: Sparganothis sulfurana (F.). 

Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, pp. 15-24.— Put- 
nam, 1963. Canad. Ent. 95: 1022-1023. 

atratus Muesebeck. N. C, Fla., Iowa. 

cerasivoranae Vereck. Hosts: Archips argyrospilus (Wlkr.), A. infumatanus 
(Zell.). 

clypeatus Muesebeck. N. H. 

crambi (Ashmead). N. Y., Md., La., Ark. 

cuniculus Walley. Mont. 

delicatus Cresson. Host: Tetralopha scortealis (Led.). 

Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, pp. 19, 24-25. 

exartemae Walley. Host: Exartema valdanum McD. 

gifuensis Ashmead. Delete; not known to occur in No. Amer. Records in 
Catalog apply to grandii Goidanich. 



BRACONIDAE 37 

*grandii Goidanich. Europe, Japan; No. Amer. Replaces gifuensis Ashmead 
of Catalog {teste Eady and Clark, 1964. Ent. Gaz. 15: 121-122). 

Macrocentrus grandii Goidanich, 1937. 1st. Ent. Univ. Bologna Bol. 9: 
196. 9. 

Biology: Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, p. 113 (as 
gifuensis Ashmead). 

impressus Muesebeck. Fla. 

incompletus Muesebeck. N. Mex., Ariz. 

instabilis Muesebeck. 

Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, p. 26. 

iridescens French. Host: Harpipteryx xylostella (L.). 

*marginator (Nees). All U. S. except southern tier of States; also N. S. to 
B. C. in Canada; Europe. Hosts: As given under aegeriae in 
Catalog and 1st Sup.; also, in Europe, Aegeria culiciformis Esp., 
A. fomnicaeformis Esp., A. flaviventris Staud., A. specif ormis 
Schiff., A. espiformis (L.), Epinotia sordidana Hiibn., E. ca- 
prana (F.) and Lathronympha striga7ia (F.). 

Bracon marginator Nees, 1812 (1811). Gesell. Naturf. Freunde Berlin 
Mag. 5: 14. 2. 

Macrocentrus aegeriae Rohwer. (Syn. by Eady and Clark, 1964. Ent. 
Gaz. 15: 113.) 

nigridorsis Viereck. Utah, Oreg., Calif. Hosts: Choristoneura conflictana 
(Wlkr.), Sciaphila duplex Wlsm., Sparganothis sp. 

reticulatus Muesebeck. N. C, Fla. 

texanus Muesebeck. Kans., Ariz.; Mexico. 

utilis Muesebeck. Fla. 

Subfamily HELCONINAE 

Genus HELCONIDEA Viereck, p. 108 

ligator (Say). Host: Agrilus anxius Gory. 

Genus HELCON Nees, p. 108 

*Edyia Cameron, 1905. Ann, Mag. Nat. Hist. (7), 16: 159. (Syn. by 
Baltazar, 1961. Philippine Jour. Sci. 90 (3) : 392.) 

Type: Edyia annulicornis Cameron. By monotypy. 

Genus METEORIDEA Ashmead, p. 109 
*Benama Nixon, 1941. Bui. Ent. Res. 32: 98. (Syn. by Shenefelt and 
Muesebeck, 1957. Ent. Soc. Wash. Proc. 59: 130.) 

Type: Benama hutsoni Nixon. By monotypy and orig. desig. 
Taxonomy: Shenefelt and Muesebeck, 1957. Ent. Soc. Wash. Proc. 59: 
129-134. 

*compressiveniris Shenefelt and Muesebeck. Wis. (Douglas Co.). 

Meteoridea compressiventris Shenefelt and Muesebeck, 1957. Ent. Soc. 
Wash. Proc. 59: 131. 9. 

Subfamily BLACINAE 
Tribe BLACINI 

Genus BLACUS Nees, p. 110 

Miocolus Foerster, 1862. (Syn. by Muesebeck, 1964. Ent. Soc. Wash. 
Proc. 66: 88.) 

In line with current literature Ganychorus Haliday and Goniocormus 
Foerster are kept as synonyms of Blacus Nees, but it seems likely that when 
a critical study of the group is made both will be recognized as distinct 
genera. 

Biology: Arnaud, 1952. Ent. News 63: 149-150. 



38 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

♦Genus PYGOSTOLUS Holiday 

Pygostolus Haliday, 1833. Ent. Mag. 1 : 263. 

Type : Cry plus stricticus Fabricius. By monotypy. 
Insert in Blacinae after Syrrhizus Foerster, p. 110. 

*falcatu8 (Nees). Man. Host: Sitona cylindricollis Fahr.; in Europe, Sitona 
lineatus (L.), S. humeralis Steph., S. hispidulus (F.). 
Leiophron falcatus Nees, 1834, Hym. Ichn. Aff. Monogr., v. 1, p. 44. .9 
Biology: Loan and Holdaway, 1961. Bui. Ent. Res. 62: 473-488.— Loan, 
1961. Jour. Econ. Ent. 54: 1026-1031.— Loan, 1963. Ent. Soc. 
Amer. Ann. 56: 605-606. 

Genus ORGILUS Holiday, p. 1 11 

dioryctriae Rohwer. Host: Dioryctria auranticella (Grote). 

gelechiae (Ashmead). N. S., P. E. I., Que., Ont. Hosts: Lithocolletis sp., 
Gracilaria sp. on Cornus canadensis. 

obscurator (Nees). Que., Ont., Mass., R. I., N. Y., N. J., W. Va., Mich. Hosts 
(in Europe): Semasia hypericana (Hbn.), Exoteleia dodecella 
(L.). 
Biology: Watson and Arthur, 1959. Canad. Ent. 91: 481.— Miller, 1960. 
Jour. Econ. Ent. 53: 318.— Juillet, 1960. Canad. Ent. 92: 342-346. 
— Leius, 1960. Canad. Ent. 92: 371-375.— Juillet, 1960. Canad. 
Ent. 92: 701-704.— Dowden, 1962. U. S. Dept. Agr., Agr. Handb. 
226, p. 62. 

Genus EUBADIZON Nees, pp. 111-112 

'''definituin Muesebeck. Oreg., Calif. Hosts: Conophthorus ponderosae Hopk., 
Ernobius sp. 
Eubadizon definitum Muesebeck, 1957. Brooklyn Ent. Soc. Bui. 52: 53. 
9, $. 

*electuin Muesebeck. Oreg., Calif.; B. C. Host: Phymatodes sp. 

Eubadizon electum Muesebeck, 1957. Brooklyn Ent. Soc. Bui. 52: 54. 9, 

*emobu Muesebeck. Calif., Oreg. Host: Ernobius sp. 

Eubadizon ernobii Muesebeck, 1957. Brooklyn Ent. Soc. Bui. 52: 51. 9, 

*extensor (Linnaeus). Europe, U. S., South. Canada, Japan. Hosts: Choris- 
toneura muriana (Hbn.), Depressaria heracleana (L.), Epinotia 
nigricana (H-S.), E. p^isillana Peyer., Spilonota lariciana Hein., 
Zeiraphera griseana CRhn.) , Z. rufimatrana (H.-S.). 

Ichneumon extensor Linnaeus, 1758. Syst. Nat., 10th Ed., p. 564. 

Eubadizon pleurale Cresson, 1872. N. syn. (based on comparison of type 
with material from Europe and Japan by C. F. W. Muesebeck). 

Biology: Capek, 1956. Biologia 11: 602.— Allen, 1962. U. S. Dept. Agr. 
Tech. Bui. 1265, pp. 27-29. 

gracile Provancher. N. H., Mass., Wis., Minn., Oreg. Hosts: Acleris variana 
(Fern.), Eucordylea ducharmei Freem., Evagora starki (Freem.), 
Recurvaria apicitripunctella (Clem.). 
Biology: SchaflFner, 1959. U. S. Dept. Agr. Misc. Pub. No. 767, p. 57.— 
Stark, 1961. Canad. Jour. Zool. 39: 895. 

"'salicicola Muesebeck. Calif. Host : Dorytomus sp. on willow. 

Eubadizon salicicola Muesebeck, 1957. Brooklyn Ent. Soc. Bui. 52: 55. 

9, $. 

Genus STANTONIA Ashmeod, p. 1 1 2 
carpocapsae Muesebeck. Tex. 

Genus MICROTYPUS Rotzeburg, p. 113 
dioiyctriae Rohwer. Host: Dioryctria auranticella (Grote). 



BRACONIDAE 39 

Tribe TRIASPIDINI 

Genus UROSIGALPHUS Ashmead, pp. 113-114 

barberi Crawford. Conn., N. Y,, Ala., Fla. 

f orbesi Martin. Miss. Host : Curculio sp. on Quercus falcata. 

robustus Ashmead. Ark. 

Genus ALIOLUS Say, Sup. 1, pp. 22-24 

collaris (Brues). Ark. Host: Conotrachelus nenuphar (Hbst.). 

crabilli Martin. Ark. 

curculionis (Fitch). Ont. Host: Listronotus oregonensis (Lee), Tyloderma 
foveolatum Say. 
Biology: Armstrong, 1958. Canad. Ent. 90: 16-17. 

marginatus Martin. Tenn., Mich. 

rufus (Riley). Ont. Hosts: Conotrachelus nenuphar (Hbst.), Tyloderma fo- 
veolatum Say. 

trilobatus (Say). Md., Va., Mich, 

TRIASPIS subgenus TRIASPIS Holiday, Sup. 1, p. 24 

kurtogaster Martin. Wis. B-Ost: Conotrachelus nenuphar (Wost.). 

*xanthochila Martin. Ont. (Belleville). 

Triaspis (T.) xanthochila Martin, 1958. Canad. Ent. 90: 191. 9. 

Subfamily AGATHIDINAE 

Genus CREMNOPS Foerster, pp. 115-116 

Revision: Marsh, 1961. Ent. Soc. Amer. Ann. 54: 851-861. 

ashmeadi (Morrison). Md., Ind., Alta. 

californicus (Morrison). N. Mex., Ariz., Colo., Wyo., Utah, Nov., Oreg., 
Wash. 

comstocki (Morrison). Kans., Nebr., Idaho. 

crassifemur (Muesebeck). Calif. 

haematodes (Brulle). Fla., Ariz., Calif. 

melanoptera Ashmead. Idaho, Nev., Wash. 

montrealensis (Morrison). Sask., N. C, Ala., S. Dak., Kans., Tex. 

nigrosternum (Morrison). N. Y., N. J., Md., D. C, S. C, Mich, Ark., Kans., 
Colo., N. Mex., Utah, Ariz., Calif. 

♦shenefelti Marsh. Nev., Calif. 

Cremnops shenefelti Marsh, 1961. Ent. Soc. Amer. Ann. 54: 857. 5> $• 

virginiensis (Morrison). Maine, Kans., Colo., Tex., Ariz., Wash. 

washingtonensis (Shenefelt). Nebr., Utah, Nev., Idaho, Oreg., Calif. 

Genus EARINUS Wesmael, p. 116 

zeirapherae Walley. Oreg. Hosts: Zeiraphera griseana (Hbn.), Rhyacionia 
adana Heinr. 

Genus AGATHIRSIA Westwood, pp. 116-117 
cressoni Muesebeck and Walkley. Colo. 
testacea Muesebeck. Calif.; Mexico. 

Genus CRASSOMICRODUS Ashmead, p. 117 
apicipennis Muesebeck. Mo., Iowa, Wyo., Calif. 



40 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

divisus (Cresson). Fla., Ariz. 

fulvescens (Cresson). Calif.; Mexico. 

medius (Cresson). Tex., Ariz. 

*inuesebecki Marsh. Calif. 

Crassomicrodus muesebecki Marsh, 1960. Pan-Pacific Ent. 36: 153. ?, $. 

nigrithorax Muesebeck. Calif.; Mexico. 
pallens (Cresson). Fla. 

Genus AGATHIS Latreille, pp. 117-121 
abdominalis (Muesebeck). Mich., Ariz. 

acrobasidis (Cushman). Va. Hosts: Laspeyresia ingens Heinr., Rhyacionia 
frustrana {Comst.) , Choristoneura fumiferana (Clem.). 

agilis (Cresson). Maine, Vt. Hosts: Archips cerasivoranus (Fitch), Spilon- 
ota ocellana (D. and S.). 

annulipes (Cresson). Wyo., Utah, Idaho, Ariz., Calif. Hosts: Paralobesia 
viteana (Clem.) , Enarmonia interstinctana (Clem.). 

atripes Cresson. Ariz. 

bakeri (Muesebeck). Mich., Minn. 

binominata Muesebeck. Que. Hosts: Petrova comstockiana (Fern.), Phalonia 
rutilana (Hbn.), Recurvaria coniferella Kft. 

brevicomis (Muesebeck). Idaho. 

buttricki (Viereck). Fla. 

calcarata (Cresson). Minn. Hosts: Exartema permandanum Clem., E. serico- 
ranum Wlsm., Sparganothis sulfurana (F.). 

coleophorae (Roh.). Idaho. Host: Coleophora laricella (Hbn.). 

difficilis (Muesebeck). N. C, Ark. 

diversa (Muesebeck). Mich. 

Biology: Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, pp. 72-73.— 
Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, pp. 30-31. 

fcstivo lyfuGScbGck 

Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, pp. 31-34. 

gibbosa (Say). Host: Mompha eloisella Clem. 

laticincta (Cresson). Wis., Oreg. Hosts: Acleris variana (Fern.). Archips 
cerasivoranus (Fitch). 

nigripes Cresson. N. Mex., Calif. 

pumila (Ratzeburg). N. S., P. E. I., N. B., Mich., Wis. 

Biology: Dowden, 1962. U. S. Dept. Agr., Agr. Handb. 226, p. 47. 

reticulata (Muesebeck). Mo. 

rufofemorata (Muesebeck). Nebr. 

simillima (Cresson). Host: Papaipema nebris (Gn.). 

spiracularis (Muesebeck). Fla., Calif. Hosts: Cogia cutis (Skin.), Erynnis 
gesta H.-S., E. zarucco Luc, Systasea pulverulenta (Feld.). 

stigmatera (Cresson). La. 

Subfamily MICROGASTERINAE 

Genus DIRRHOPE Foerster, p. 122 
americana Muesebeck. Mich., Tex. 



BRACONIDAE 41 

*Genus MUESEBECKIA Mason 

Muesebeckia Mason, 1957. Canad. Ent. 89: 355. 

Type: Muesebeckia eximius Mason. By monotypy and orig. 
desig. 

Insert in Microgasterinae following Paroligoneurus Muesebeck, p. 122. 

*eximius Mason. Ont., Que., N. C. 

Muesebeckia eximius Mason, 1957. Canad. Ent. 98: 355. 5, $. 

Genus APANTELES Foerster, pp. 123-135 

alaskensis (Ashmead). Calif. 

alticola (Ashmead). N. H., Idaho, Calif. 

argynnidis Riley. Tex. Host: Erynnis funeralis (Scud, and Burg.), 

autographae Muesebeck. Que., Va, Hosts: Autoplusia egena (Gn.), Pseuda- 
letia unipuncta {Kaw.) , Rachiphisia ou (Gn.). 

banks! Viereck. N. Y., Mich. 

bedelliae Viereck. Hawaii. 

bushnelli Muesebeck. Fla. Hosts: Dioryctria abietella (D. and S.), D. clario- 
ralis (Wlkr.), 

cacoeciae Riley. Hosts: Archips cerasivoranus (Fitch), Exartema valdanum 
McD. 

calif omicus Muesebeck. Host: Eucosma bobana Kearf. 

Biology: Telford, 1961. Jour. Econ. Ent. 54: 347-355.— Stark, 1961. 
Canad. Jour. Zool. 39: 894. 

canarsiae Ashmead. Ont. Hosts: Acrobasis betulella Hulst, Argyrotaenia 
velutinana (Wlkr.) , Dioryctria renicidella (Grote). 

carpatus (Say). Hosts: Tinea despecta Meyr., Oecia maculata Wlsm. 

*Urogaster solitarius Ashmead, 1901. Ent. Soc. London Trans., 1900, p. 

287. 9. Preocc. Syn. by Muesebeck, 1958. U. S. Natl. Mus. Proc. 

107: 431. 
*Apanteles piceoventris Muesebeck, 1921. U. S. Natl. Mus. Proc. 58: 

515. N. name for solitarius (Ashmead). Syn. by Muesebeck, 1958, 

loc. cit. supra, p. 431. 
Biology: Key and Common, 1959. Austral. Jour. Zool. 7: 64. 

cinctiformis Viereck. Ind., Wis. 

cingiliae Muesebeck. Hosts: Enypia venata (Grote), E. griseata Grossb. 

clavatns (Provancher) . Wis., Ark. Host: Etiella zinckenella (Treit.). 

cockerelli Muesebeck. Mich., Mo., Nebr., Calif. 

conanchetorum Viereck. N. Y., D. C, W. Va., Mich., Wis., Ark. Host: Lox- 
ostege mancalis (Led.). 

congregatus ( Say) . 

Biology: Lawson, 1959. Ent. Soc. Amer. Ann. 52: 743-744. 

consimilis Viereck. Pa., Ohio, Mich. 

cyaniridis Riley. 111. Hosts: Philotes battatoides bemardino B. and McD., 
Incisalia henrici (G. and R.). 

edwardsii Riley. Minn., Iowa. 

electrae Viereck. Tex. Host: Automeris io (F.), 

empretiae Viereck. Mass. Host: Lithacodes fasciola (H.-S.). 

Biology: Schaffner, 1959. U. S. Dept. Agr. Misc. Pub. 767, p. 60. 

ensiger (Say). Ala., Colo. 

*enypiae Mason. B. C. Hosts: Enypia griseata Grossb., E. venata (Grote). 
Apanteles enypiae Mason, 1959. Canad. Ent. 91: 47. 9, $. 



42 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

epiblemae Muesebeck. Fla., Calif. 

epinotiae Viereck. Hosts: Carpocapsa pomonella (L.), Keiferia lycopersi- 
cella (Busck), Platynota stultana Wlsm., Stegasta basqueella 
(Chamb.), Suleivia helianthana (Riley). 
Biology: Satterthwaite, 1948. Jour. Econ. Ent. 41: 730.— Allen, 1962. 
U. S. Dept. Agr. Tech. Bui. 1265, pp. 34-35. 

etiellae Viereck. Colo., Ariz., Calif. 

feltiae Viereck. Mich., Utah, Ariz., Calif. 

flaviconchae Riley. Utah, Wash. 

Biology and taxonomy : Muesebeck, 1947. Pan-Pacific Ent. 23 : 21-22. 

forbesi Viereck. Ariz. 

fumiferanae Viereck. Hosts: Dioryctria reniculella (Grote), Thymelicus lin- 
eola (Ochs.). 
Biology: McGugan, 1955. Canad. Ent. 87: 178-187.— Miller, 1959. Can- 
ad. Ent. 91: 457-477.— McGugan and Blais,. 1959. Canad. Ent. 91: 
774.— Lewis, 1960. Canad. Ent. 92: 881-891. 

*galleriae Wilkinson. France, Argentina, Mauritius; N. C, S. C, Ga. Host: 
Gallerin mellonella (L.). 
Apanteles galleriae Wilkinson, 1932. Stylops 1: 139. 9, $. 

gillettei Baker. Ariz. 

glomeratns (Linnaeus). New Zealand. 

Distribution: Todd, 1962. New Zealand Jour. Sci. 5: 1-7. 
Biology: Matsuzawa, 1955. Kagawa-Ken Agr. Co. Tech. Bui. 7, p. 60. — 
Clausen, 1956. U S. Dept. Agr. Tech. Bui. 1139, p. 77.— Blunck 
and Wilbert, 1959. Internatl. Congr. Crop Prot. Proc. 4, v. 1, p. 
1017. — Moiseeva, 1960. Leningrad. Vsesoiuzn. Inst. Rast. Trudy, 
14: 51. 
griffini Viereck. Okla., S. Dak. 

harrisinae Muesebeck. Calif. Host: Harrisina americana (Guer.) . 

Biology: Schaffner, 1959. U. S. Dept. Agr. Misc. Pub. 767, p. 60.— Clau- 
sen, 1961. Hilgardia 31: 613-638. 

harti Viereck. Mich. 

hesperidivorus Viereck. Fla., Ariz. Host: Hylephila phyleus (Drury). 

hyphantriae Riley. Europe. 

Biology: Tadic, 1958 (1956). Tenth Intern. Congr. Ent. Proc. 4: 
859-861. 

koebelei Riley. Host: Euphydryas chalcedona (D. and H.). 

lacteicolor Viereck. 

Biology: Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, p. 78.— Dow- 
den, 1962. U. S. Dept. Agr., Agr. Handb. 226, p. 48. 

*Iaricellae Mason N. B., Ont., Wis. Hosts: Argyresthia laricella Kit., Coleo- 
phora laricella (Hbn.). 
Apanteles laricellae Mason, 1959. Canad. Ent. 91: 45. 9, $. 

laspeyresiae Viereck. Hosts: Laspeyresia miscitata Heinr., Hedulia injectiva 

Heinr. 
longicomis (Provancher) . 

Biology: Wood and Neilson, 1960. Canad. Ent. 92: 760. 

marginiventris (Cresson). Calif., Hawaii. Hosts: Ethmia colonella Wlsm., 
Heliothis virescens (F.), Hymenia recurvalis (F.), Scotorythra 
caryopsis Meyr., Spodoptera exempta (Wlkr.), S. mauritia acronyc- 
toides (Gn.). 

medicaginis Muesebeck. 

Biology: Allen, 1958. Hilgardia 27: 515-541.— Allen and Smith, 1958. 
Hilgardia 28: 1-42.— Stern and Bowen, 1963. Ent. Soc. Amer. 
Ann. 56: 358-372. 



BRACONIDAE 43 



megathymi Riley. N. C. Host : Megathymus ursus Poling. 

melanoscelus (Ratzeburg). 

Biology: Clausen, 1956. U. S, Dept. Agr. Tech. Bui. 1139, p. 79.— Dow- 
den, 1962. U. S. Dept. Agr., Agr. Handb. 226, p. 48. 

miantonomoi Viereck. Mich. 

militaris (Walsh). N. B., N. Mex., Ariz. Apparently distributed over all U. 
S. and south. Canada; also in Mexico and Hawaii. Hosts: ChoH- 
zagrotis auxiliaris (Grote) , Laphygma exigua (Hbn.). 

neomexicanus Muesebeck. Mich., Idaho. 

nephoptericis (Packard). N. Y., Ind., Iowa, Calif. Host: Vitula edmandsii 
serratilineella Rag. 

niger Muesebeck. Va., Mich., Minn. 

nitens Muesebeck. Colo., Wyo., Calif. Host: Pseudaletia unipuncta (Haw.). 

omigis Weed. Que. Hosts: Bucculatrix ainsliella Murtf., Lithocolletis robin- 
iella Clem., Tischeria raalif oliella Clem. 

paleacritae Riley. Hosts: Enypia venata (Grote), E. packardata Tayl. 

papaipemae Muesebeck. Ohio. Host: Arzaina obliquaWlkr . 

paralechiae Muesebeck. N. H., Pa., Ohio, Tenn., Wis., Calif. Host: Argy- 
resthia sp. 
Biology: Schaffner, 1959. U. S. Dept. Agr. Misc. Pub. 767, p. 61. 

paranthrenidis Muesebeck. Pa., Fla., Miss.; Mexico. Host: Paranthrene dolli 
Neum., Prionoxystus sp. in red oak. 

petrovae Walley. S. C, Wis., Minn,, Calif. Hosts: Barbara colfaxiana 
Kearf., Rhyacionia frustrana (Comst.). 

plathypenae Muesebeck. N. Y., Ind., Idaho, Wash. Hosts: Anagrapha falcif- 
era (Kirhy) , Autographa precationis {Gn.) , Rachiplusia ou (Gn.). 

plesius Viereck. Wis. 

polychrosidis Viereck. Fla., Kans., Wis., Minn. Hosts: Gracilaria superbi- 
frontella Clem., Archips rosanus (L.)., Argyrotaenia citrana 
(Fern.). 

pseudoglossae Muesebeck. Mich., Minn. 

pyralidis Muesebeck. D. C, N. C, Ohio, Ark. 

*salaiicus Mason. Oreg. Host: Lithocolletis gaultheriella Wlsm. 
Apanteles salalicus Mason, 1959. Canad. Ent. 91: 42. 9. 

*salicifolieIIae Mason. Ont. Host: Lithocolletis salicif oliella Chamb. 
Apanteles salicif oliellae Mason, 1959. Canad. Ent. 91: 43. $, $. 

schaffneri Muesebeck. N. J., Tex. Hosts: Euclea delphinii Bdv., Parasa sp. 
Biology: Schaifner, 1959. U. S. Dept. Agr. Misc. Pub. 767, p. 62. 

scitulus Riley. Ky., 111. 

scutellaris Muesebeck. Ariz.; Bulgaria. 

Taxonomy: Beardsley, 1961. Hawaii. Ent. Soc. Proc. 17: 341, 346. 

eolitarius (Ratzeburg). 

Biology: Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, pp. 79-80.— 
Dowden, 1962. U. S. Dept. Agr. Agr. Handb. 226, p. 49. 

*starki Mason. Alta, B. C, Utah, Idaho. Hosts: Evagora starki (Freem.), 
Recurvaria milleri Busck. 
Apanteles starki Mason, 1960. Canad. Ent. 92: 140. 9, $. 

theclae Riley. Colo. Host: Plebe jus icarioides (Bdv.). 

Biology: Downey, 1962. Ent. Soc. Amer. Ann. 55: 367-373. 

tischeriae Viereck. Wis., Mo. Hosts: Paromix geminatella (Pack.), ArgyrO' 
taenia velutinana (Wlkr.). 



44 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

yakutatensis Ashmead. Idaho, Calif. 

'^zizaniae Muesebeck. D. C, Del. Host: Diatraea sp. in wild rice. 
Apanteles zizaniae Muesebeck, 1957. Ent. News 68: 22. $, $. 

Genus MICROGASTER Latreille, pp. 135-137 

auripes Provancher. Mich., Nebr., N. B. Hosts: Farontia diffusa (Wlkr.), 
Pseudaletia unipuncta (Haw.). 

brittoni Viereck. Mich. 

calliptera Say. Iowa, Nebr. 

canadensis Muesebeck, N. S. Hosts: Archips argyrospilus (Wlkr.), Stilpno- 
tia salicis (L.). 

comptanae Viereck. Va., Iowa. 

congregatiformis Viereck. Mich. 

ecdytolophae Muesebeck. Nebr. Hosts: Carpocapsa pomonella (L.), Walshia 
amorphella Clem. 
Biology: Brandhorst, 1962. Ent. Soc. Amer. Ann. 55: 477.— Allen, 1962. 
U. S. Dept. Agr. Tech. Bui. 1265, p. 36. 

epagoges Gahan. Hosts: Acleris chalybeana (Fern.), Anacampsis rhoifruc- 
tella (Clem.), Sparganothis acerivorana MacKay. 
Biology: Schaflfner, 1959. U. S. Dept. Agr. Misc. Pub. 767, p. 62. 

facetosa Weed. Host: Argyrotaenia velutinana (Wlkr.). 

gelechiae Riley. Host: Gnorimoschema gallaeasterella (Kell.). 
Biology: Judd, 1962. Amer. Micros. Soc. Trans. 81: 204. 

hamedi Muesebeck. Mich. 

iridescens Cresson. Host: Sylepta gordialis (Gn.). 

leechi Walley. Mass., Md., Mich., Oreg. 

pantographae Muesebeck. Iowa. 

peronea Walley. D. C, Wash. Hosts: Archips argyrospilus (Wlkr.), Peron- 
ea logiana (h.), Sparganothis pettitana (Rob.). 

phthorimaeae Muesebeck. Hosts: Udea ruhigalis (Gn.), Platynota stultana 
(Wlsm.). 

schizurae Muesebeck. Mich., Ark. 

♦scotica Marshall. Europe; Mich. 

Microgaster scotica Marshall, 1885. Ent. Soc. London Trans. 1885, p. 251. 

$. 
*tiro Reinhard. Europe; Newfoundland, P. E. I. Host: Cnephasia virgua- 
reana Treit. 
Microgaster tiro Reinhard, 1880. Deut. Ent. Ztschr. 24: 356, 357. ?, $. 

zonaria Say. Pa., Fla., Iowa, Wis. 

Genus MICROPLITIS Foerster, pp. 137-139 

alaskensis Ashmead. Ariz. Hosts: Rachiplusia ou (Gn.), Trichoplusia ni 
(Hbn.), Pseudaletia unipuncta (Haw.). 

autographae Muesebeck. Ariz. 

crenulatus (Provancher). Mich. 

croceipes (Cresson). Ariz. 

feltiae Muesebeck. Calif. 

gortynae Riley. Pa. 

kewleyi Muesebeck. Calif. Host: Agr otis ipsilon {Uutn.). 

longicauda Muesebeck. Nev., Calif. 



BRACONIDAE 45 



montanus Muesebeck. Nev. 

perplexus Muesebeck. Mich. 

plutellae Muesebeck. Ont., N. Dak. 

Biology: Harcourt, 1960. Canad. Ent. 92: 420-421. 

scutellatus Muesebeck. Idaho. 

stigmaticus Muesebeck. Calif. 

striatus Muesebeck. Mich. 

varicolor Viereck. N. B. Host: Syngrapha epigaea (Grote). 
Biology: Wood and Neilson, 1960. Canad. Ent. 92: 761. 

Subfamily CARDIOCHILINAE 

Genus CARDIOCHILES Nees, pp. 139-141 
apicalis (Cresson). D. C, S. C, Fla., Ariz. 
insculptus Mao. Fla. 
levis Mao. Calif. 
magnus Mao. Iowa. 
minutus (Cresson). Md. 
pullus Mao. Iowa, Calif. 
rubicundus Mao. Ariz. 
rubidus Mao. Idaho. 
libiator (Say). Fla., Mo. 
truncus Mao. Wyo. 
viator (Say). N. Y., Mich., Wyo., N. Mex. 

Subfamily ICHNEUTINAE 

Genus ICHNEUTIDEA Ashmead, pp. 141-142 
proteroptoides Viereck. Miss. 

Subfamily CHELONINAE 

Genus PHANEROTOMA Wesmael, pp. 142-143 

*Sulydus Du Buysson, 1897. Soc. Ent. de France Ann. 66: 354. Syn. by 
Fischer, 1963. Beitr. z. Ent. 13: 195. 

Type : Sulydus marshalli Du Buysson. By monotypy. 

erythrocephala Rohwer. Ga., Wis., Mont., Calif. Hosts: Laspeyresia piper- 
ana (Kearf.), Helulia injectiva Heinr. 

fasciata Provancher. Hosts: Laspeyresia ingens Heinr., L. anaranjada 
Miller. 
Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, pp. 40-41. 

laspeyresiae Rohwer. Mich. Hosts: Laspeyresia piperana (Kearf.), L. misci- 
tata Heinr., Helulia injectiva Heinr. 

longicauda Walley. N. Y., Va., Tenn., Wis., S. Dak., Tex. 

*planifron8 (Nees). Europe, North Africa, West and Central Asia, Siberia, 
Korea, Japan; Calif. Hosts: Etiella zinckenella (Treit.), Grapho- 
litha molesta (Busck), G. glycinivorella Mats., Laspeyresia strobi- 
lella. (L.), Margaronia pyloalis (Wlkr.), Phycita diaphana 
(Stgr.). 
Sigalphus planifrons Nees, 1816 (1813). Gesell. Naturf. Freunde Berlin 
Mag. 7: 259. 



46 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Taxonomy : Watanabe, 1940. Insecta Matsumurana 14 : 88-89. 
Biology: Parker, 1951. U. S. Dept. Agr. Tech. Bui. 1036, pp. 12-16.— 
Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, p. 121. 

recurvariae Gush. Wis. 

thapsina Walley. Ariz., Calif. 

tibialis (Haldeman). Ont., Maine, Del., Ala., 111., Mich., Wis., Minn. Host: 
Acrobasis betulella Hulst. 

Genus ASCOGASTER Wesmael, p. 143 

flaviceps Ashmead. N. Y. 

quadridentata Wesmael. Wis., Colo., Calif. Hosts: Laspeyresia nigricana 
(Steph.), Spilonota ocellana (D. and S.). 
Biology: Boyce, 1936. Canad. Ent. 68: 241-246.— Allen, 1962. U. S. 
Dept. Agr. Tech. Bui. 1265, pp. 37-40. 

CHELONUS subgenus CHELONUS Panzer, pp. 143-144 

annulipes Wesmael. Del., N. Y. Host: Diatraea grandiosella Dyar (in lab.). 
Biology: Bradley and Arbuthnot, 1938. Ent. Soc. Amer. Ann. 31: 
259-269. — Harris, 1950. Iowa Yearbook of Agr. for 1949, pp. 
143-159.— Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, p. 91. 

aughei (La Munyan). Delete "Locality not given." Nebr. (North Platte). 

*inanitus (Linnaeus). Europe; Calif. Host: Etiella zinckenella (Treit.). 
Cynips inanitus Linnaeus, 1767. Syst. Nat., 12th Ed., p. 919. 
Biology: Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, pp. 91-92. 

*muesebecki McComb. Calif., Alaska, Ariz., Colo., Nev., N. Mex., Oreg., 
S. D.. Utah, Wash. 
Chelonus (Chelonus) muesebecki McComb, 1962. Ent. Soc. Wash. Proc. 
64: 19. 9, $. 

nebraskensis (La Munyan.). Delete "Locality not given." Nebr. (North 
Platte) . 

*phaloniae Mason. Man. Host: Phalonia hospes (Wlsm.). 

Chelonus phaloniae Mason, 1959. Canad. Ent. 91: 49. 9, 5. 

CHELONUS subgenus MICROCHELONUS Szepligeti, pp. 145-146 

egregicolor Viereck. N. H., Mass. 

Biology: Schaffner, 1959. U. S. Dept. Agr. Misc. Pub. 767, p. 64. 

phthorimaeae Gahan. Idaho; Mexico, Bermuda. Host: Gnorimoschema gud- 
manella (Wlsm.). 

Subfamily DACNUSINAE 

Taxonomy: Nixon, 1943. Ent. Monthly Mag. 79: 23-34, 159-168; 1944. 
Idem 80: 88-108, 140-151, 193-200, 249-255; 1945. Idem 81: 
189-204, 217-220; 1946. Idem 82: 279-300; 1948. Idem 84; 
207-224, 1949. Idem 85: 289-298; 1954. Idetn 90: 257-290.— Rie- 
gel, 1947. The American Species of Dacnusinae, Urbana, Illinois, 
8 pp.— Docavo, 1955. Graellsia 13: 1-34.— Griffiths, 1964. Beitr. z. 
Ent. 14: 823-914 (Dacnusini). 

Biology: Griffiths, 1956. Ent. Monthly Mag. 92: 25-30.— Griffiths, 1962. 
Ent. Record and Jour. Variation 74: 183-184.— Griffiths, 1964. 
Beitr. z. Ent. 14: 823-914. 

Genus CHOREBUS Holiday, p. 146 

Liposcia Foerster, 1862. Transf. from syn. here to syn. under Pachysema 
Foerster. 

aquaticus Muesebeck. Minn. 



BRACONIDAE 47 

*Genus PACHYSEMA Foerster 

Pachysema Foerster, 1862. Naturh. Ver. Rheinlande Verh. 19: 274. 

Type: Dacnusa macrospila Haliday. By monotypy and orig. 
desig. 
Liposcia Foerster, 1862. Transf. from syn. under Chorebus Haliday, p. 
146. 

Place in Dacnusinae after Chorebus Haliday. 

As yet no North Amer. species have been recorded. 

Taxonomy: Nixon, 1964. Ent. Monthly Mag. 90: 257-274. 

Subfamily ALYSIINAE 

Taxonomy and Biology: Griffiths, 1962. Ent. Rec. and Jour. Variation 
74: 184.— Griffiths, 1964. Beitr. z. Ent. 14: 823-914. 

Genus APHAERETA Foerster, p. 148 

Taxonomy: McComb, 1960. Ent. News 71: 175-178. 

auripes (Provancher) . Syn. of pallipes (Say). 

delosa Viereck. Syn. of pallipes (Say). 

*excavata McComb. Ga. (Somerville). Host: ■Sarcop^a^'a sp. 

Aphaereta excavata McComb, 1960. Ent. News 71: 176. ?, $. 

*inasoni McComb. Ont. (Stratford), Host: Cordllur a STp. 

Aphaereta masoni McComb, 1960. Ent. News 71: 177. 9, 5. 

pallipes (Say). Ont.; Hawaii. Hosts: Musca autumnalis DeG., M. domestica 

L. 
Trichesia auripes Provancher (syn. by McComb, 1958. Ent. Soc. Wash. 

Proc. 60: 223). 
Aphaereta delosa Viereck (syn. by McComb, 1958, loc. cit. supra, p. 223). 
Aphaereta subtricarinata Viereck (syn. by McComb, 1958, loc. cit. supra, 

p. 223). 
Aphaereta pegomyiae Brues (syn. by McComb, 1958, loc. cit. supra, p. 

223). 
Aphaereta sarcophagae Gahan (syn. by McComb, 1958, loc. cit. supra, p. 

223). 
Taxonomy: McComb, 1958. Ent. Soc. Wash. Proc. 60: 223-224.— 

McComb, 1960. Ent News 71: 175-178. 
Biology: Salkeld, 1959. Canad. Ent. 91: 93-97. 

pegomyiae Brues. Syn of pallipes (Say). 

sarcophagae Gahan. Syn. of pallipes (Say). 

subtricarinata Viereck. Syn. of pallipes (Say). 

Genus IDIASTA Foerster, p. 150 

Taxonomy: Konigsmann, 1960. Beitr. z. Ent. 10: 624-654. 

Genus MESOCRINA Foerster, p. 150 
Taxonomy: Konigsmann, 1959. Beitr. z. Ent. 9: 609-619. 

Genus PHAENOCARPA Foerster, p. 152 
picinervis (Haliday). Md. Kans., Idaho. 

Subfamily OPIINAE 

Taxonomy: Fischer, 1959. Acta Ent. Mus. Natl. Pragae 33: 241-263.— 
Fischer, 1963. Beitr. z. Ent. 13: 194-222, 662-747.— Fischer, 1964. 
Polskie Pismo Ent. 34: 197-530. 

Genus ADEMON Holiday, p. 152 

*Giardinaia Destefani Perez, 1902. Zool. Jahrb. Syst. 15: 631. N. syn. 
{teste C. F. W. Muesebeck, from literature). 



48 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Type : Giardinaia urinator Destef ani Perez. By monotypy. 
Biology: Destefani Perez, loc. cit. supra, pp. 625-634. — Rimsky-Korsa- 
kov, 1933. Handb. biol. Arbeitsmethoden, Abt. 9, Tail 7, p. 255. 

niger (Ashmead). Wis., Minn. 

Genus GNAPTODON Holiday, p. 153 

nepticulae (Rohwer). N. Y., Md. 

Genus OPIUS Wesmael, pp. 153-157 

*Tolhia Cameron, 1907. Linn. Soc. London Trans. (2) 12: 84. Syn. by 
Baltazar, 1961. Philippine Jour. Sci. 90(3) : 392. 

Type : Tolbia scaevolae Cameron. By monotypy. 
*Hexaulax Cameron, 1910. Soc. Ent. 25: 25. Syn. by Fischer, 1964. 
Polskie Pismo Ent. 34: 209. 

Type : Hexaulax ruficeps Cameron. By monotypy. 
*Eristernaulax Viereck, 1913. U. S. Natl. Mus. Proc. 46: 362. Syn. by 
Fischer, 1963. Polskie Pismo Ent. 33: 272. 

Type: Erisiemaulax leucotaenia Viereck. By monotypy and 

orig. desig. 

Taxonomy. Fischer, 1957. Deut. Ent. Ztschr., N. F. 4: 332-358.— Fischer, 

1958. Mus. Civ. Stor. Nat. Geneva Ann. 70: 33-70, 245-304.— 

Fischer, 1958. Hist. Nat. Mus. Nat. Hungarici 50: 241-260.— 

Fischer, 1958. Naturh. Mus. Wien. Ann. 62: 210-219.— Fischer, 

1958. Acta Ent. Mus. Natl. Pragae 32: 295-316.— Fischer, 1958. 
Beitr. z. Ent. 8: 189-212.— Fischer, 1959. Munchner Ent. Gesell. 
Mitt. 49: 1-35.— Fischer, 1959. Mus. Civ. Stor. Nat. Genova Ann. 
71: 145-161.— Fischer, 1959. Opuscula Ent. 24: 55-78.— Fischer, 

1959. Polskie Pismo Ent. 29: 519-542.— Fischer, 1959. Hist. Nat. 
Mus. Nat. Hungarici 51: 419-423.— Fischer, 1960. Ann. Zool. 19: 
33-112.— Fischer, 1962. Naturh. Mus. Wien Ann. 65: 117-131.— 
Fischer, 1963. Acta Ent. Mus. Natl. Pragae 35: 197-242— Fischer, 
1963. Polskie Pismo Ent. 33: 253-409. 

aberrans Viereck. Ont., S. C, Wis. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 464-466. 

*aciurae Fischer. Fla. Host: My olej a limata (Coq.). 

Opius aciurae Fisher, 1964. Polskie Pismo Ent. 34: 272. 9, $. 

*adductus Fischer. Calif. Host: Agromyza sp. 

Opius adductus Fischer, 1964. Polskie Pismo Ent. 34: 413. ?, $. 

*alachuaiius Fischer. Fla. (Gainesville). 

Opius alachuanus Fischer, 1964. Polskie Pismo Ent. 34: 379. ?. 

*aIconanus Fischer. Mich. (Alcona Co.). 

Opius alconanus Fischer, 196i. Polskie Pismo Ent. 34 : 416. $. 

alloeus Muesebeck. Pa., Mich., Wis. 

'''ainensis Fischer. N. Y., Kans., Iowa. 

Opius amensis Fischer, 1964. Polskie Pismo Ent. 34: 309. 9, $. 

amplus (Ashmead). Newfoundland, Que., Alta., B. C. Host: Agromyza albU 
tdvszs IM^isT 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 351-352. 

*angustistriatus Fischer. S. C, N. Y. 

Opius angustistriatus Fischer, 1964. Polskie Pismo Ent. 34: 353. ?. 

*apertu8 Fischer. Mich. (Midland Co.). 

Opius apertus Fischer, 1964. Polskie Pismo Ent. 34: 417. $. 

apicalis Gahan. N. Y., Md., Conn., Wis., Kans., Calif. Host : Liriomyza sp. 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 372-373. 

*aquaticu8 Muesebeck, n. name. Eeplaces hydrelliae Muesebeck, preocc. 



BRACONIDAE 49 

*aureliae Fischer. Europe; Colo., Wash, 

Opius aureliae Fischer, 1957. Deut. Ent. Ztschr., N. F., 4: 343. 9. 
Taxonomy and distribution: Fischer, 1964. Polskie Pismo Ent. 34: 
355-356. 

baldufi Muesebeck. Mich., Host: Replace Rhagoletis altemata (Fall.) with 
R. basiola (O. S.). 
Biology: Balduf, 1959. 111. Biol. Monogr. 26: 32-40. 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 357-358. 

'''basicastaneus Fischer. S. C. (Pickens Co.). 

Opius basicastaneus Fischer, 1964. Polskie Pismo Ent. 34: 501. $. 

♦berberidis Fischer. Wash., N. Y. Hosts: Rhagoletis berberis Curran, R. 
tabellaria (Fitch). 
Opius berberidis Fischer, 1964. Polskie Pismo Ent. 34: 358. 9, $. 

♦bidentis Fischer. Fla. (Lake Placid). 

Opius bidentis Fischer, 1964. Polskie Pismo Ent. 34: 360. 9, $. 

*blantoni Fischer. Alaska, Wash., Oreg. 

Opius blantoni Fischer, 1964. Polskie Pismo Ent. 34: 338. 9. 

brunneiventris Cresson. 

Taxonomy : Fischer, 1964. Polskie Pismo Ent. 34 : 274-276. 

*brunnescens Fischer. N. Y. 

Opius brunnescens Fisher, 1964. Polskie Pismo Ent. 34: 340, 2, $. 

canaliculatus Gahan. 111., Minn., Man. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 277-279. 

*carinu8 Fischer. N. C. (Pink Beds). 

Opius carinus Fischer, 1964. Polskie Pismo Ent. 34: 253. $. 

*castaneigaster Fischer, N. Y., N. C, S. C, Ohio, Tenn., Wis. 

Opius castaneigaster Fischer, 1964. Polskie Pismo Ent. 34: 419, 9, $. 

*chapmani Fischer. Mass. (So. Hadley). 

Opius chapmani Fischer, 1964. Polskie Pismo Ent. 34: 311. $. 

cincticornis Gahan. N. Y., N. J., N. C, Mich., Newfoundland, Que., Sask. 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 226-228. 

cinctus Provancher. Mich., Utah, N. S., Ont., Yukon. 

Taxonomy : Fischer, 1964. Polskie Pismo Ent. 34 : 255-258. 

*cinereifactus Fischer. N. Y. 

Opius cinereifactus Fischer, 1964. Polskie Pismo Ent. 34: 503. ?, $. 

*clellanvillensis Fischer. S. C. (MeClellanville). 

Opius clellanvillensis Fischer, 1964. Polskie Pismo Ent. 34: 454. 5. 

♦clevelandensis Fischer. N. Y., S. C. 

Opius clevelandensis Fischer, 1964. Polskie Pismo Ent. 34: 421. 9, $. 

*coIeogaster Fischer. Fla. 

Opius coleogaster Fischer, 1964. Polskie Pismo Ent. 34: 399. 9, S. 

*columbiacu8 Fischer. N. Y., N. C, S. C, Ariz. 

Opius columbiacus Fischer, 1964. Polskie Pismo Ent. 34: 466. 9, $. 

*comatus Wesmael. Europe; N. Y., Md., S, C. 

Opius comatus Wesmael, 1835. Nouv. Mem. Acad. Sci. Bruxelles 9: 145. 

9 $ . 
Taxonomy: 'Fischer, 1964. Polskie Pismo Ent. 34: 229-230. 

*confundens Fischer. Tex., La., Fla., S. C. 

Opius confundens Fischer, 1964. Polskie Pismo Ent. 34: 381. 9, $. 

♦convergitalis Fischer. N. C. (Pink Beds). 

Opius convergitalis Fischer, 1964. Polskie Pismo Ent. 34: 468. 9. 

coriaceus Gahan. D. C, Del., N. C, S. C, Ga. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 383-385. 



50 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*crabtreeanus Fischer. N. C, N. Y. 

Opius crabtreeanus Fischer, 1964. Polskie Pismo Ent. 34: 470. $. 

*crandalli Fischer. Ariz. (Mt. Lemon). 

Opius crandalli Fischer, 1964. Polskie Pismo Ent. 34: 312. ?. 

""curtiarticulatus Fischer. Sask., Mass., N. Y. 

Opius curtiarticulatus Fischer, 1964. Polskie Pismo Ent. 34: 315. ?. 

^dimensus Fischer. N. Y. (Ithaca). 

Opius dimensus Fischer, 1964. Polskie Pismo Ent. 34: 362. 9, 

dimidiatus (Ashmead). Host: Liriomyza munda Yrick. 

Taxonomy: Fischei% 1964. Polskie Pismo Ent. 34: 448-450. 

downesi Gahan, Mich. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 364-365. 

*dreisbachi Fischer. Mich., N. Y. 

Opius dreishachi Fischer, 1964. Polskie Pismo Ent. 34: 385. ?, $. 

^euwattacooanus Fischer. Md., S. C, Mich. 

Opius euwattacooanus Fischer, 1964. Polskie Pismo Ent. 34: 505. 9, $. 

*extiratus Fischer. Mass., N. Y., Md. 

Opius extiratus Fischer, 1964. Polskie Pismo Ent. 34: 473. $. 

Tarmingdalensis Fischer. N. Y. (Farmingdale). 

Opius fanningdalensis Fischer, 1964. Polskie Pismo Ent. 34: 507. 9. 

*farmingda!icus Fischer. N. Y. (Farmingdale). 

Opius farmingdalicus Fischer, 1964. Polskie Pismo Ent. 34: 475. $. 

ferrugineus Gahan. N. J. 

flaviceps Gahan. Que., R. I., N. Y., N. J., N. C. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 423-425. 

foersteri Gahan. Md., Va., Colo. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 365-367. 

*foutsi Fischer. Md. (Cabin John). 

Opius foutsi Fischer, 1964. Polskie Pismo Ent. 34: 401. 9. 

*frequens Fischer. Me., N. Y., Mich., Minn., Oreg., Wash., Que., Ont. Man. 
Host: Rhagoletis fausta (O. S.). 
Opius freguens Fischer, 1964. Polskie Pismo Ent. 34: 279. 9, $. 

fulvicollis Thomson. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 237-239. 

gahani Muesebeck. Pa., Wis., Minn. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 281-283. 

*gahanoides Fischer. 111., Mich. 

Opius gahanoides Fischer, 1964. Polskie Pismo Ent. 34: 283. 9. 

*gainesvillensis Fischer. Fla., N. C, Md. 

Opius gainesvillensis Fischer, 1964. Polskie Pismo Ent. 34: 402. 9, $. 

*gIasgowi Fischer. N. Y., S. Dak. Host: Rhagoletis cingulata (Loew). 
Opius glasgowi Fischer, 1964. Polskie Pismo Ent. 34: 286. 5. 

*globiformis Fischer. Md., N. Y., Mich., Wis., Ont. 

Opius glohiforniis Fischer, 1964. Polskie Pismo Ent. 34: 497. 9, 5. 

*gIobigaster Fischer. N. Y., Pa., Va., N. C. 

Opius globigaster Fischer, 1964. Polskie Pismo Ent. 34: 387. 9, $. 

gracillariae Gahan. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 425-426. 

*grafi Fischer. Md., Va., N. C, S. C. 

Opius graft. Fischer, 1964. Polskie Pismo Ent. 34: 389. 9, S. 

*gravicornis Fischer. Que. (Wakefield). 

Opius gravicornis Fischer, 1964. Polskie Pismo Ent. 34: 230. 5. 



BRACONroAE 51 



*hancockanus Fischer. N. Y. (Hancock). 

Opius hancockanus Fischer, 196A. Polskie Pismo Ent. 34: 426. 9. 

'^hardmanni Fischer. Calif., Oreg., Wash. 

Opius hardmanni Fischer, 1964. Polskie Pismo Ent. 34: 211. 5, $. 

*hartlcyi Fischer. Alaska. 

Opius hardeyi Fischer, 1964. Polskie Pismo Ent. 34: 258. $. 

*hoflfmanni Fischer. Mich., Ga. 

Opius hoffmanni Fischer, 1964. Polskie Pismo Ent. 34: 391. ?, $. 

^hoUisterensis Fischer. Idaho (Hollister). 

Opius hollisterensis Fischer, 1964. Polskie Pismo Ent. 34: 328. $. 

hydrelliae Muesebeck. Preocc. Replaced by aquaticus Muesebeck. 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 393-395. 

*igneus Fischer. Fla., S. C; Cuba. 

Opius igneus Fischer, 1964. Polskie Pismo Ent. 34: 477. 9, $. 

*ignifer Fischer. N. Y. (Oneonta). 

Opius ignifer Fischer, 1964. Polskie Pismo Ent. 34: 232. ?. 

*interinissus Fischer. S. C. 

Opius intermissus Fischer, 1964. Polskie Pismo Ent. 34: 428. ?, $. 

*irregulari8 Wesmael. Europe; Wash., Oreg., N. C, N. Y. 

Opius irregularis Wesmael, 1835. Nouv. Mem. Acad. Sci. Bruxelles 9: 

132. $ $. 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 336-338. 

*isolatae Fischer. Calif., Wash. Host: Agromyza isolata Mall. 

Opius isolatae Fischer, 1964. Polskie Pismo Ent. 34: 478. $, ?. 

*ithaceiisis Fischer. N. Y. (Ithaca). 

Opius ithacensis Fischer, 1964. Polskie Pismo Ent. 34: 239. 9. 

*jacobi Fischer. B. C, Que., Alaska, Wash., Oreg., Calif. 

Opius jacobi Fischer, 1964. Polskie Pismo Ent. 34: 329. 9, $. 

*juglandis Muesebeck. Ariz. Host: Rhagoletis juglandis (Cress.). 

Opius juglandis Muesebeck, 1961. Brooklyn Ent. Soc. Bui. 56: 57. 9, $. 

*juniperi Fischer. Ariz., Man. 

Opius juniper! Fischer, 1964. Polskie Pismo Ent. 34: 288. 9, $. 

'''karesuandensis Fischer. Sweden; Alaska, Yukon. 

Opius karesuandensis Fischer, 1964. Polskie Pismo Ent. 34: 241. $, 9. 

*kinleyensis Fischer. Alaska (Mt. McKinley). 

Opius kinleyensis Fischer, 1964. Polskie Pismo Ent. 34: 317. $, 9. 

*krombeini Fischer. Va. (Arlington). 

Opius krombeini Fischer, 1964. Polskie Pismo Ent. 34: 367. 9. 

kukakensis (Ashmead). Oreg. 

*IatitemporaHs Fischer. N. Y. (Ithaca). 

Opius latitemporalis Fischer, 1964. Polskie Pismo Ent. 34: 509. $. 

lectoidcs Gahan. N. S., Ont. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 289-291. 

lectus Gahan. N. B., Ont. Mich., Calif. Host: Rhagoletis fausta (0. S.). 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 291-293. 

*leevingensis Fischer. Calif. (Leevining). 

Opius leevingensis Fischer, 1964. Polskie Pismo Ent. 34: 369. $. 

lemnaphilae Muesebeck. 

Biology: Scotland, 1939. Ent. Soc. Amer. Ann. 32: 716-717.— Scotland, 

1940. N. Y. Ent. Soc. Jour. 48: 323-325. 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 430-431. 



52 HYMENOPTERA. OF AMERICA NORTH OP MEXICO 

*lemonensis Fischer. Ariz.; Mexico. 

Opius lemonensis Fischer, 1964. Polskie Pismo Ent. 34: 319. 5, $. 

*Iiriomyzae Fischer. Calif. (Santa Cruz). Host: Liriomyza subpusilla 
(Frost). 
Opius firiomyzae Fischer, 1964. Polskie Pismo Ent. 34 : 260. $. 

*macrocerus Thomson. Europe; Mich. 

Opius macrocerus Thomson, 1895. Opusc. Ent. fasc. 20, p. 2186. ?, $. 
* Opius hians Stelfox, 1949. Ent. Monthly Mag. 85: 306. 9, $. 
Taxonomy and distribution: Fischer, 1964. Polskie Pismo Ent. 34: 
321-323. 

*maIivorelIae Fischer. Iowa, S. Dak., W, Va., Mass. Host: Coleophora mali- 
vorella Riley. 
Opius malivorellae Fischer, 1964. Polskie Pismo Ent. 34: 213. 9, $. 

mandibularis Gahan. R. I., Md., Ga., Ariz. Host: Phytobia maculosa (Mall.). 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 215-217. 

*masoni Fischer. Ont., Md., N. Y., S. C. 

Opius inasoni Fischer, 1964. Polskie Pismo Ent. 34: 480. 9, $. 

melleus Gahan. N. J., Ohio. Host: Rhagoletis suavis (Loew). 

^middlckauffi Fischer. Calif. (Berkeley). Host: Pegomya hyoscyami 
( Panz.) . 
Opius middle kauffi Fischer, 1964, Polskie Pismo Ent. 34: 244. 9, $. 

montanus (Ashmead). 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 511-513. 

*myakken8is Fischer. Fla. (Myakka State Park). 

Opius myakkensis Fischer, 1964. Polskie Pismo Ent. 34: 404. $. 

'^nigritellae Fischer. Calif. (Berkeley). Host: Near Phytomyza nigritella 
(Zett.). 
Opius nigritellae Fischer, 1964. Polskie Pismo Ent. 34: 342. ?. 

*nitiduIator (Nees). Europe; Calif., Wyo., Colo., Ont., Alta., Sask., B. C. 
Host: Pegomya hyoscyami (Panz.). 
Bracon nitidulator Nees, 1834. Hym. Ichn. Aff. Monog. v. 1, p. 56. $. 
*Opius vittatus Ruschka, 1915. Ztschr. f. Angew. Ent. 2: 395. 9, $. 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 246-248. 

*northcarolinensis Fischer. N. Y., Md., N. C, S. C, Mich. 

Opius northcarolinensis Fischer, 1964. Polskie Pismo Ent. 34: 513. 2, $. 

*notaulicus Fischer. Que., Mass., R. I. 

Opius notaulicus Fischer, 1964. Polskie Pismo Ent. 34: 217. 9, $. 

*nudiscutum Fischer. Calif., Tenn., Sask. Hosts: Liriomyza subpusilla 
(Frost), Agromyza sp. 
Opius nudiscutum Fischer, 1964. Polskie Pismo Ent. 34: 373. ?, 3. 

*ordinariu8 Fischer. Calif., Oreg., Wash. Hosts: Agroinyza sp., Liriomyza 
sp. 
Opius ordinarius Fischer, 1964. Polskie Pismo Ent. 34: 450. 9, $. 

*omatigaster Fischer. Calif. (Laurel). Host: A grofnyza s'p. 

Opius ornatigaster Fischer, 1964. Polskie Pismo Ent. 34: 515. 9. 

*ornatus Fischer. N. Y., N. J., Md., Va., N. C, S. C. 

Opius ornatus Fischer, 1964. Polskie Pismo Ent. 34: 262. 9, $. 

oscinidis (Ashmead). Md., Va., Mich. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 234-236. 

otiosus Gahan. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 499-500. 

*^pallidipalpis Fischer. N. C, S. C. 

Opius pallidipalpis Fischer, 1964. Polskie Pismo Ent. 34: 482. 9, $. 



BRACONIDAE 53 

*paradisi*acus Fischer. Fla. (Paradise Key). 

Opius paradisiacus Fischer, 1964. Polskie Pismo Ent. 34: 456. $. 

*paralleIiformis Fischer. N. Y. (Ithaca). 

Opius paralleliformis Fischer, 1964. Polskie Pismo Ent. 34: 452. $. 

'''paratakomanus Fischer. Md. (Takoma Park). 

Opius paratakomanus Fischer, 1964. Polskie Pismo Ent. 34: 484. ?. 

"parawattacooanus Fischer. S. C. (Wattacoo). 

Opius parawaitacooanus Fischer, 1964. Polskie Pismo Ent. 34: 517. 9. 

*parkeranu8 Fischer. Ariz. (Parker Creek). 

Opius parkeranus Fischer, 1964. Polskie Pismo Ent. 34: 458. ?. 

*parkercreekensis Fischer. Ariz., Calif., S. C. 

Opius parkercreekensis Fischer, 1964. Polskie Pismo Ent. 34: 486. 9. 

*parvifossa Fischer. N. Y., Mich., Tenn., Wis., Kans., N. B., Que., Ont. 
Opius parvifossa Fischer, 1964. Polskie Pismo Ent. 34: 294. S, 9. 

*pauper Fischer. Wash., Oreg., Calif. 

Opius pauper Fischer, 1964. Polskie Pismo Ent. 34: 519. 9, $. 

pegomyiae Gahan. Syn. of spinaciae Thomson. 

*perininutus Fischer. Calif. (Santa Cruz). 

Opius perminutus Fischer, 1964. Polskie Pismo Ent. 34: 332. 9, $. 

*perpygmaeus Fischer. Md., S. C. 

Opius perpygmaeus Fischer, 1964. Polskie Pismo Ent. 34: 521. 9, $. 

^pilosinotum Fischer. N. Y. (Ithaca). 

Opius pilosinotum Fischer, 1964. Polskie Pismo Ent. 34: 522. 9. 

*piIosus Fischer. Calif. (Yosemite). 

Opius pilosus Fischer, 1964. Polskie Pismo Ent. 34: 333. 9. 

*pinkbedsanus Fischer. N. C, N. Y. 

Opius pinkbedsanus Fischer, 1964. Polskie Pismo Ent. 34: 488. 9, $. 

*pisgahensis Fischer. N. Y., N. C. 

Opius pisgahensis Fischer, 1964. Polskie Pismo Ent. 34: 323. $. 

*poIyzonius Wesmael. Europe; N. Y., Md., Wis., Ont., Sask., B. C. Hosts: 
Agromyza beatica Grif., A. hendeli Grif., A. phragmitidis Hend. 

Opius polyzonius Wesmael, 1835. Nouv. Mem. Acad. Sci. Bruxelles 9: 
136. 9, $. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 248-250. 

*porteri Fischer. Fla. (Dade Co.), 

Opius porteri Fischer, 1964. Polskie Pismo Ent. 34 : 431. 9. 

^prolongatus Fischer. S. C. (Greenville). 

Opius profongam* Fischer, 1964. Polskie Pismo Ent. 34: 459. 9. 

provancheri Dalla Torre. Pa., Kans., Iowa, Colo. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 296-298. 

'''pseudocolumbiacus Fischer. N. C. (Mt. Pisgah). 

Opius pseudocolumbiacus Fischer, 1964. Polskie Pismo Ent. 34: 489. $. 

*punctipes Fischer. Fla. (Gainesville). 

Opius punciipes Fischer, 1964. Polskie Pismo Ent. 34: 405. 9. 

♦pusilloides Fischer. N. Y., Md., N. C, S. C, Kans. 

Opius pusilloides Fischer, 1964. Polskie Pismo Ent. 34: 433. 9, $. 

*rainierensis Fischer. Wash. (Mt. Rainier). 

Opius rainierensis Fischer, 1964. Polskie Pismo Ent. 34: 244. 9. 

*relativus Fischer. S. C. 

Opius relativus Fischer, 1964. Polskie Pismo Ent. 34: 435. 9, S. 

*re8irictus Fischer. S. C, Md. 

Opius restrictus Fischer, 1964. Polskie Pismo Ent. 34: 407. ,9 $. 



54 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*resupinns Fischer. Calif. (San Bernardino Co.). 

Opius resupmu* Fischer, 1964. Polskie Pismo Ent. 34 : 409. 9. 

richmondi Gahan. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 298-300. 

*robsonensis Fischer. B. C, Wash. 

Opius robsonensis Fischer, 1964. Polskie Pismo Ent. 34: 300. 9, $. 

rosicola Muesebeck. Sask. Host: Replace Rhagoletis altemata (Fall.) with 
R. basiola (O. S.). 
Biology: Balduf, 1959. 111. Biol. Monogr. 26: 32-40. 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 303-305. 

♦rudifomiis Fischer. Europe; Ont. 

Opius rudiformis Fischer, 1958. Beitr. z. Ent. 8: 201. 9. 
Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 395-396. 

*ruficolor Fischer. Fla. 

Opius ruficolor Fischer, 1964. Polskie Pismo Ent. 34: 492. ?. 

*rufipleurum Fischer. N. Y., S. C. 

Opius Tufipleurum Fischer, 1964. Polskie Pismo Ent. 34: 264. 9, $. 

rufocinctus Gahan. Md., N. C, S. C, Kans. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 376-378. 

*sabroskyi Fischer. Va. (Fairfax Co.). 

Opius sabroskyi Fischer, 1964. Polskie Pismo Ent. 34: 396. 9, $. 

*salmonensis Fischer. B. C. (Shuswap Lake). 

Opius salmonensis Fischer, 1964. Polskie Pismo Ent. 34: 438. 5. 

sanguineus (Ashmead). Ariz. 

Biology: Cazier, 1962. Pan-Pacific Ent. 38: 184-186. 

*8everini Fischer. N. Y., Mich., S. Dak., Iowa, Colo., Utah, Alta., Que. 
Opius severini Fischer, 1964. Polskie Pismo Ent. 34: 223. 9, $. 

*shuleri Fischer. S. C. (McClellanville). 

Opius shuleri Fischer, 1964. Polskie Pismo Ent. 34: 524, $. 

*sierraanchaensis Fischer. Ariz. (Sierra Ancha). 

Opius sierraanchaensis Fischer, 1964. Polskie Pismo Ent. 34: 346. $. 

'*'similarius Fischer. N. Y. 

Opius similarius Fischer, 1964. Polskie Pismo Ent. 34: 267. 9, $. 

*sniithi Fischer. D. C, Va., S. C. 

Opius smithi Fischer, 1964. Polskie Pismo Ent. 34: 410. 9, $. 

*southcaroIinensi8 Fischer. S. C. (Greenville). 

Opius southcarolinensis Fischer, 1964. Polskie Pismo Ent. 34: 526. $. 

spinaciae Thomson. 

Opius pegomyiae Gahan. Syn. according to Max Fischer (in Ktt.). 

*8tenopectus Fischer. S. C. (McClellanville). 

Opius stenopectus Fischer, 1964. Polskie Pismo Ent. 34: 461. S. 

striativentris Gahan. Mich., Kans. 

Taxonomy : Fischer, 1964. Polskie Pismo Ent. 34 : 440-442. 

succineus Gahan. Pa., Md., D. C, Ont., Sask., Alta. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 269-271. 

♦suffixus Fischer. N. Y., Mich., Wis. 

Opius suffixus Fischer, 1964. Polskie Pismo Ent. 34: 493. $, 

suturalis Gahan. Tex. Host: LzWowyza ptcfeWa (Thoms.). 

*tabellariae Fischer. N. Y. (Geneva). Host: Rhagoletis tabellaria (Fitch). 
Opius tabellariae Fischer, 1964. Polskie Pismo Ent. 34: 305. 9, $. 

^tablerockensis Fischer. S. C. 

Opius tablerockensit Fischer, 1964. Polskie Pismo Ent. 34: 250. 9, ^. 



BRACONIDAE 55 

♦tangens Fischer. Fla. (Gainesville). 

Opius tangens Fischer, 1964. Polskie Pismo Ent. 34: 442. 9. 

♦testaceifactus Fischer. Fla., S. C. 

Opius testaceifactus Fischer, 1964. Polskie Pismo Ent. 34: 220. 9, $. 

tibialis (Ashmead). Mass., N. Y., N. J., N. C, S. C, Calif. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 495-497. 

*troyensis Fischer. N. Y. (Troy). 

Opius troyensis Fischer, 1964. Polskie Pismo Ent. 34: 325. ?. 

atahensis Gahan. Mass., Va., Fla., Minn., Que. 

Taxonomy: Fischer, 1964. Polskie Pismo Ent. 34: 222-223. 

*vockerothi Fischer. Que., B. C, Wash. 

Opius vockerothi Fischer, 1964. Polskie Pismo Ent. 34: 348. 9, $. 

*wallcyi Fischer. Fla. (Chattachoochee). 

Opius walleyi Fischer, 1964. Poliskie Pismo Ent. 34: 444. $. 

*weemsi Fischer. N. C. (Mt. Pisgah). 

Opius weenisi Fischer, 1964. Polskie Pismo Ent. 34: 446. 9, $. 

*westerlyanus Fischer. R. I. (Westerly). 

Opius westerlyanus Fischer, 1964. Polskie Pismo Ent. 34: 528. 9. 

Subfamily BRACONINAE 

Taxonomy: Telenga, 1936. Faune URSS, v. 5, No. 2 (1) (keys to gen- 
era), pp. 24-27 (Russian), 307-309 (German).— Tobias, 1957. 
Zool. Zhur. 36: 1338-1354. 

Genus VIPIO Lotreille, pp. 157-158 
belfragei (Cresson). Ga., Ind., Mo. Host: Sphenophthorus cariosus (Oliv.). 
moneilemae Gahan. Calif. 
sanguineus (Provancher). Transf. to Iphiaulax Latreille, p. 159. 

Genus IPHIAULAX Foerster, pp. 158-159 

erythrus (Viereck). Tex., N. Mex. 

manteri Nettleton. Ga. Host: Oiketicus abbottii Grote. 

perepicus Viereck. Tex., N. Mex. 

sanguineus (Provancher), n. comb. Transf. from Vipio, 158 {teste C. F. W. 
Muesebeck, after examination of type). Ariz. 

Genus ATANYCOLIMORPHA Viereck, p. 159 
dissitus (Cresson). Oreg. 

Genus CYANOPTERUS Holiday, p. 159 
laevis (Provancher). Pa., Minn. 

Genus COELOIDES Wesmael, pp. 160-161 

brunneri Viereck. 

Coeloides scolyti Cushman (syn. by Muesebeck, in Ryan, 1962. Ent. Soc. 

Amer. Ann. 55: 403.) 
Biology: Ryan, 1962. Ent. Soc. Amer. Ann. 55: 403-409.— Ryan and 

Rudinsky, 1962. Canad. Ent. 94: 748-763. 
Embryology: Ryan, 1963. Ent. Soc. Amer. Ann. 56: 639-648. 

dendroctoni Cushman. 

Biology: Reid, 1963, Canad. Ent. 95: 233-235. 

pissodis (Ashmead). N. S., N. B., Ont. Hosts: Ips calligraphus (Germ.), /. 
grandicollis (Eichh.), Pissodes approximatus Hopk. 

scolyti Cushman. Syn. of brunneri Viereck. 



56 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus ATANYCOLUS Foerster, pp. 161-163 

hicoriae Shenefelt. Mich. 

rugosiventris (Ashmead). Fla. 

Genus BRACON Fabricius, pp. 163-169 

*acrobasidis Muesebeck. Fla., N. J. Hosts: Acrobasis caryae Grote, A. caryi- 
vorella Rag., A. comptoniella Hulst. 
Bracon acrobasidis Muesebeck, 1963. Ent, News 74: 158. 9, $. 

*agathymi Muesebeck. Ariz., Texas. Hosts: Agathymus neumoegini (Edw.), 
A. aryxna (Dyar), A. mcalpinei (Freem.). 
Bracon agathymi Muesebeck, 1963. Ent. News 74: 157. 9, $. 

americanus (Ashmead). Colo., Ariz. 

analcidis Ashmead. Host: Sphenophthorus parvulus Gyll. 

*bruchivorus Muesebeck. Calif. Hosts: "Bruchids," Acanthoscelides s^. 
Bracon hruchivorus Muesebeck, 1963. Ent. News 74: 162. 9, 5- 

caulicola (Gahan). Host: Suleima helianthana (Riley). 

Biology: Wester, 1956. Ent. Soc. Wash. Proc. 58: 283-284. 

cephi (Gahan). 

Biology: Nelson and Farstad, 1953. Canad. Ent. 85: 103-107.— Holmes, 
et al., 1963. Canad. Ent. 95: 113-126. 

cushmani (Muesebeck). Host: Rhyacionia frustrana (Comst.). 

erucarum (Cushman). Okla., Wye, Mont. 

furtivus Fyles. 

Biology: Miller, 1963. Ohio Jour. Sci. 63: 73. 

gastroideae Ashmead. S. Dak. 

gelechiae Ashmead. Hosts: Anacampsis fragariella Busck, Anthophila pcv- 
riana (Clerck), Argyrotaenia citrana (Fern.), Dichomeris mar- 
ginella. (F.), Dioryctria disclusa Heinr., D. reniculella (Grote), 
Keiferia glochinella (Zell.), Pterophorus periscelidactylus Fitch, 
Pyrausta signatalis (Wlkr.), Thymelicus lineola (Grote). 

Biology: Poos, 1928. Ent. Soc. Wash. Proc. 30: 149.— Gupta, 1951. Na- 
ture 168: 793.— Narayanan and Rao, 1955. Beitr. z. Ent. 5: 36-60. 
—Wester, 1956. Ent. Soc. Wash. Proc. 58: 284. 

Anatomy: Narayanan and Rao, 1955. Beitr. z. Ent. 5: 286-293 (9 repro- 
ductive system). 

gemmaecola (Cushman). N. J., Ohio, Ark., Tex. 

hebetor Say. Host: Laetilia coccidivora (Comst.). 

Taxonomy: Lai, 1946. Indian Jour. Ent. 8: 85-88. — Puttarudriah and 

Channa Basavanna, 1956. Bui. Ent. Res. 47: 183-191. 
Biology: Kanungo, 1955. Current Sci. 24: 59. — Grosch, La Chance and 
Sullivan, 1955. Ent. Soc. Amer. Ann. 48: 415-416.— Grosch, 1959. 
Ent. Soc. Amer. Ann. 52: 94-98.— Clark, 1963. Ent. Soc. Amer. 
Ann. 56: 616-619. 

*gossypii Muesebeck. S. C, La., Ga., Miss., Tex., Fla., Ark. Host: ? Anthon- 
omus grandis Boh. 
Bracon gossypii Muesebeck, 1963. Ent. News 74: 161. 9, $. 

hobomok (Viereck). Mich., Iowa. 

hyslopi (Viereck). Host: Platyptilia williamsii Grin. 

*jani Muesebeck. Minn. (Cloquet). Host: Janus abb reviatus (Say). 
Bracon jani Muesebeck, 1963. Ent. News 74: 164. 9, $. 

juncicola Ashmead. B. C. 

lissosTdstcF IVTuGScbGck 

Biology: Somsen'and Luginbill, 1956. U. S. Dept. Agr. Tech. Bui. 1153, 
7 pp. 



BRACONIDAE 57 

mellitor Say. Ont. Hosts: Coeliodes inaequalis (Say), Conotrachelus nenu- 
phar (Hbst.), Cylas formicarius elegantulus (Sum.), Ostrinia nu- 
bilalis (Hbn.), Rliyacionia adana Heinr., R. fnistrana (Comst.), 
Rhynchites bicolor (F.), Smicraulax tuberculatus Pierce, Spar- 
ganothis sulfurana (F.), Tachypterellus quadrigibbus (Say), Ty- 
loderma foveolatum (Say). 

nevadensis Ashmead. Ariz. 

oenotherae (Muesebeck). Minn., Nebr. 

papaipemae (Gahan). Mich., Minn. 

*piger Wesmael. Europe; Calif. Host: Etiella zinckenella {Treit.). 

Bracon piger Wesmael, 1838. Brussels Acad. Sci. Mem. 11: 48. $. 
Biology: Parker, 1951. U. S. Dept. Agr. Tech. Bui. 1036, pp. 8-11.— 
Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, p. 87. 

pini (Muesebeck) N. S., N. B., Va., Colo., Calif. Host: Pissodes terminalis 
Hopping. 

platynotae (Cushman). Host: Platynota stultana (Wlsm.). 

politiventris (Cushman). Hosts: Archips rosanus (L.), Dioryctria reniculella 
(Grote). 

punctatus (Muesebeck). Md., Mich. "Rost : Faustinus cubae (Boh.). 

quinnipiacorum (Viereck). S. Dak. 

*radicis Shenefelt. Wis. Host: Hylobius radicis Buch. 

Bracon radicis Shenefelt, 1960. Canad. Ent. 92: 872. 9, $. 

rhjacioniae (Muesebeck). Colo. Host: Laspeyresia youngana (Kearf.). 

*rosaceani (Muesebeck). Oreg. (Gresham). Host: Archips resaceanus 
(Harris). 
Bracon rosaceani Muesebeck, 1963. Ent. News 74: 160. ?, $. 

rudbeckiae (Muesebeck). Ark. 

sanninoideae (Gahan). Wis. Host: Alcathoe caudata (Harr,). 

sesiae (Muesebeck). Colo. 

tenuiceps (Muesebeck). Ohio, Mich., Idaho. 

thurberiphagae (Muesebeck). 

Biology: Bennett, 1960. Bui. Ent. Res. 50: 747-749. 

tychii (Muesebeck). Idaho. 

xanthonotus Ashmead. Hosts: Malacosoma disstria Hbn.. M. pluviale recen- 
seo Dyar, Sabulodes caberatae Gn. 

Subfamily SPATHIINAE 
Taxonomy: Nixon, 1943. Roy. Ent. Soc, London Trans. 93: 173-456. 

Genus SPATHIUS Nees, pp. 169-170 

brachyums Ashmead. Mass. 

brunneri Viereck. Utah, Idaho, Oreg., Calif. Host: Pseudohylesinus nebu- 
losus (Lee). 

californicus Ashmead. Idaho. 

canadensis Ashmead. Pa., Ohio, Ky. 

Morphology: Snodgrass, 1941. Smithsn. Misc. Collect. 99: 35-36 ($ genita- 
lia). 
Biology: Kaston, 1937. Conn. Agr. Expt. Sta. Bui. 396, pp. 351-353.— 
Kaston, 1939. Conn. Agr. Expt. Sta. Bui. 420, pp. 33-35. 

pallidus Ashmead. Tex. 

simiilimus Ashmead. Miss., Wis. 



58 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Subfamily ROGADINAE 

Genus ROGAS Nees, pp. 170-172 

abdominalis (Cresson). D. C. 

fumipennis (Cresson). N. H., N. Y., Wis. 

granulatus DeGant. Mich. 

graphicus Cresson. S. Dak., Wyo. 

hyphantriae Gahan. 

Biology: Bogavac, 1955. Plant Prot. (Belgrade), No. 32, pp. 17-30.— 
Tadic, 1959. Internatl. Congr. Crop Prot. Proc. 4, v. 1, p. 874. 

""indicus Cameron. Alaska (Sitka), not "Sitka on the Ganges" as given with 
orig. descr. {teste Mather, 1957. Zool. Soc. Calcutta Proc, Moc- 
ker jee Mem. vol., p. 350). 
Rhogas indicus Cameron, 1910. Wien. Ent. Ztg. 29: 3. $. 

laphygmae Viereck. Fla. ; Cuba, Nicaragua. 

mandibularis Cresson. 111., Kans. 

molestus Cresson. Wyo. 

perplexus Gahan. Host: Trichoplusia ni (Hbn.). 

Genus PELECYSTOMA Wesmael, p. 172 
discoideum (Cresson). N. Y., Pa., Md. 

Genus PETALODES Wesmael, p. 172 
palmatus Walley. Wis., Minn., Calif. 
ufei Walley. Nev. 

Genus YELICONES Cameron, p. 173 

*Pectenopius Fischer, 1961. Naturhist. Mus. Wien. Ann. 64:156. N. 
syn., teste C. F. W. Muesebeck, from literature. 

Type: Pectenopius paradoxus Fischer. By monotypy and orig. 
desig. 

Genus BUCCULATRIPLEX Viereck, p. 173 

bucculatricis (Ashmead). 111. Host: Bucculatrix ainsliella Murt. 
Biology: Schaffner, 1959. U. S. Dept. Agr. Misc. Pub. 767, p. 65. 

Genus ONCOPHANES Foerster, p. 173 

americanus (Weed). Wash. Host: Endothemia hebesana (Wlkr.). 

Biology: Zeller and Schuh, 1944. Oreg. Agr. Exp. Sta. Bui. 418, p. 48.— 
Schaffner, 1959. U. S. Dept. Agr. Misc. Pub. 767, p. 66.— Bennett, 
1961. Tenn. Acad. Sci. Jour. 36: 353. 

bctulsc ]\Tu.GS€b6ck« Wis. 

Biology: Schaffner, 1959. U. S. Dept. Agr. Misc. Pub. 767, p. 66. 

Genus PHANOMERIS Foerster, pp. 174-175 

metalli Muesebeck. Maine, Mich., Wis. Host: Cameraria bethunella Chamb. 

phyllotomae Muesebeck. 

Biology: Dowden, 1962. U. S. Dept. Agr., Agr. Handb. 226, p. 63. 

Genus XENARCHA Foerster, Sup. 1, p. 35 
hopkinsi (Ashmead). Ark. 

♦Subfamily HORMIINAE 
Insert before Tribe Pambolini, p. 175. 



BRACONIDAE 59 

Taxonomy: Hedqvist, 1963. Ent. Tidskr. 84: 30-61 (key to genera of the 
World). 

Tribe Pambolini and Tribe Hormiini, p. 1 75 

Delete both headings and combine component groups under Subfamily 
Hormiinae. 

Genus HORMIUS Nees, pp. 175-176 

dispar (Brues). Md., 111., Mo., Tex. 

*Hormius micropterus Hedqvist, 1963. Ent. Tidskr. 84: 50-51. 9. N. 
syn. {teste C. F. W. Muesebeck, from literature). 

^Genus MONITORIELLA Hedqvist 

Monitoriella Hedqvist, 1963. Ent. Tidskr. 84: 44. 

Type : Monitoriella elongata Hedqvist. Orig. desig. 

Insert in Hormiinae after Pambolus Haliday, p. 175. 

*elongata Hedqvist. Tex. ; Mexico. From galls on Philodendron. 

Monitoriella elongata Hedqvist, 1963. Ent. Tidskr. 84: 46. 9, $. 

*Genus CANTHAROCTONUS Viereck 

Cantharoctonua Viereck, 1912. U. S. Natl. Mus. Proc. 42: 617. 

Type: Cantharoctonus stramineus Viereck. By monotypy and 
orig. desig. 

Insert in Hormiinae after Parahonnius Nixon, p. 176. 

*brunneus Hedqvist. Fla. (Sebring). 

Cantharoctonus brunneus Hedqvist, 1963. Ent. Tidskr. 84: 38. $. 

Subfamily DORYCTINAE 

Taxonomy: Nixon, 1943. Bui. Ent. Res. 34: 257-267.— Granger, 1949. 
Inst. Sci. Madagascar Mem., Ser. A. Biol. Anim. 2: 94-145. — Par- 
rott, 1955. Roy. Soc. N. Zealand Trans. 82: 1157-1163.— Tobias, 
1961. Zool. Zhur. 40: 529-535. 

Genus DORYCTES Holiday, pp. 176-177 
apacheus Viereck. Mo. 
atripes (Provancher) . Pa., D. C, Wis. 
fartus (Provancher). N. Y., Minn. 
incertus Ashmead. Mass. 

pacificus (Provancher). Host : Paraiima com'coZa Fisher. 
rufiventris (Brulle). Ont. Host: Xylotrechus colonus (F.). 

Genus DORYCTINUS Roman, P. 178 
secnndus Muesebeck and Walkley. Ariz. 

Genus POLYSTENOIDES Muesebeck, P. 178 
lignicola Muesebeck. Md. 

Genus GLYPTOCOLASTES Ashmead, p. 178 

bruchivorus Crawford. Ariz.; Peru, Canal Zone. Host: Dendrobiella quadri- 
spinosa (Lee). 

Genus RUSSELLELLA Muesebeck and Walkley, p. 178 
prosopidis (Muesebeck). Ariz. 



60 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus HETEROSPILUS Holiday, pp. 178-180 

koebelei (Ashmead). Hosts: Prodoxus aenescens Riley, P. marginatus Riley, 
Tegeticula 7naculata (Riley). 

melleus (Ashmead). Delete "W. Va. (Morgantown.) " Fla., Md. Hosts: Gno- 
rimoscheme gallaeasterella Kell., Mompha sexnotella Chamb. 

prosopidis Viereck. Okla., Ariz., Calif.; Mexico. 

Genus PIOSCELUS Muesebeck and Walkley, p. 180 
wichitus (Viereck). Md. 

Genus ALLORHOGAS Gahan, p. 180 

bruesi (Muesebeck and Walkley). Transf. from Miocolus Foerster, p. 182 
(teste Muesebeck, 1964. Ent. Soc. Wash. Proc. 66: 88). 

Genus DENDROSOTER Wesmael, p. 181 

scaber Muesebeck. Oreg. Host: CylindrocoptuTnis longulus (hec.) . 

scolytivorus (Viereck and Rohwer). Idaho, Ariz. Host: Ips oregonis 
(Eichh.). 

sulcatus Muesebeck. Host: Ips grandicollis (Eichh.). 

Genus ODONTOBRACON Cameron, pp. 181-182 
grandis Ashmead. Tex., Calif. 

Genus Miocolus Foerster, p. 1 82 

Transf. to Blacinae and place as syn. of Blacus Nees, p. 110; transf. 
bruesi Muesebeck and Walkley to AUorhogas Gahan, p. 180, and place both 
minimus (Cresson) and utilis (Cresson) as synonyms of Monolexis fuscicornis 

Foerster, p. 182 (teste Muesebeck, 1964. Ent. Soc. Wash. Proc. 66: 88). 

Genus MONOLEXIS Foerster, p. 182 

fuscicornis Foerster. 

Anisopelma minimus Cresson (syn. teste Muesebeck, 1964. Ent. Soc. 

Wash. Proc. 66: 88). 
Anisopelma utilis Cresson (syn. teste Muesebeck, 1964. Ent. Soc. Wash. 

Proc. 66: 88). 

Genus ACRISIS Foerster, p. 182 
americana (Ashmead). Pa., Mich. 

Family ICHNEUMONIDAE 

This second supplement of the Hymenoptera Catalog brings many 
changes, some of them complex, in the nomenclature of the Ichneumonidae. 
These changes have resulted from revisionary work in several subfamilies, 
and from a treatise of the type-species of the Foerster genera which were 
established originally without included species. In several instances, where 
the conclusions of two competent systematists differed and no specimens of 
the species involved were available, or no definitive characters given, the 
course followed was the one involving the least change. 

The pagination for species in "Hymenoptera of Beulah, New Mexico," by 
H. L. Viereck, 1903 (February), Amer. Ent. Soc. Trans. 29: 43 (bottom) -100, 
differs in the separate (published March 20). In the latter, the article begins 
at the top of page 43 and ends on page 99. The description of a given species 
usually occurs one page earlier in the separate. The pagination of species 
listed in the Catalog is that of the separate except in the Second Supplement, 
where a few were changed before the reason was discovered. 

The following revisions, when they occur in the text, are cited by author 
and date only because of frequency of occurrence, except that the complete 
reference is given for ail new species and subspecies. 



ICHNEUMONIDAE 61 

Revisions: Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 1]: 
MX, 1-318 (Metopiinae).— Townes and Townes (et al.) 1960. U. 
S. Natl. Mus. Bui. 216 (Pt. 2) : I-VII, 1-673 (Ephialtinae, Xoridin- 
ae, Acaenitinae) . — Townes, Townes, and Gupta, 1961. A Reclas- 
sification of the Indo-Australian Ichneumonidae [Amer. Ent. Inst. 
Mem. 1] pp. I-IV, 1-552 (supergen. & gen. keys; syn.). — Heinrich, 
1961 (1960) -1962. Canad. Ent. Sup. 15 [vol. 92], pt. I, pp. 1-87; 
Sup. 18 [vol. 92], pt. II, pp. 89-205; Sup. 21 [vol. 93], pt. Ill, pp. 
207-368; Sup. 23, pt. IV, pp. 370-505; Sup. 26, pt. V, pp. 507-671 
Sup. 27, pt. VI, pp. 675-802; Sup. 29, pt. VII, pp. 803-886 (Ichneu- 
moninae. — Townes and Townes, 1962. U. S. Natl. Mus. Bui. 216 
(Pt. 3): I-IV, 1-602 ((Mesostenini)=Cryptini).— Townes and 
Gupta, 1962. Amer. Ent. Inst. Mem. 2: 1-305 (Hemigasterini). — 
Dasch, 1964, Amer. Ent. Inst. Mem. 3: 1-304 (Diplazoninae). — 
Peck, 1964. Canad. Ent. Mem. 35, pt. VIII of Heinrich Synopsis 
Nearc. Ichneumoninae, pp. 891-925. — Townes, Momoi, and Townes, 
1965. A catalogue and Reclassification of the Eastern Palearctic 
Ichneumonidae. Amer. Ent. Inst. Mem. 5: 1-661. [This publica- 
tion appeared after the ms. for this supplement was completed: it 
is cited only when obligatory.] 

Keys: Constantineanu, 1959. Fauna Republicii Populare Romine, 
Ins., V. 9, fasc. 4, pp. [107]-109.— Perkins, 1959. Roy. Ent. Soc, 
London Handbook 7 (Pt. 2 (ai)): 1-18 (keys to subfamilies). — 
Short, 1959. U. S. Natl. Mus. Proc. 110 (3419) : 391-511 (keys to 
larvae).— Constantineanu, 1961. Beit. z. Ent. 11 (7/8): 685-732 
(keys to subfamilies). 

Taxonomy and/or Nomenclature: Perkins, 1962. Brit. Mus. (Nat. Hist.) 
Bui., Ent. 11 (8) : [385]-483 (types of Foerster genera). 

Subfamily PIMPLINAE 

Revision: Townes and Townes (et al) 1960. U. S. Natl. Mus. Bui. 216 
(Pt. 2) : 3-676 (as Ephialtinae, Xoridinae, Acaenitinae). 

Tribe EPHIALTINI 
[zzPimplini of Catalog) 

♦Genus EXERISTES Foerster 

Exeristes Foerster, 1868. Naturh. Ver. Rheinlande Verh., v. 25, p. 164. 

Type: (Pimpla roboralor Gravenhorst) = roborator (Fabri- 
cius). Inch by Schmiedeknecht, 1888. 

Place before Scambus Hartig, p. 185. 

comstockii (Cresson). S. C. Hosts: Dioryctria clarioralis (Wlkr.) , Laspeyre- 
sia caryana (Fitch). Transf. from Calliephialtes Ashmead (teste 
Townes and Townes, 1960, p. 11). 
Biology: Arthur, 1963. Canad. Ent. 95: 1079-1085 (life history, imm. 
stages). 

Genus SCAMBUS Hartig, pp. 185-187, Sup. 1, pp. 36-37 

Ateleophadnus Cameron. Place as valid subg. of Scambus. 
Pseudopoenienia Kiss. Place as syn. of subg. Ateleophadnus. 
Endrotnopoda Hellen. Place as valid subg. of Scambus. 
Revision: Walley, 1960. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 2) : 14-79. 

SCAMBUS subgenus ENDROMOPODA Hellen 

Remove from syn. of Scambus Hartig (teste Walley 1960. In Townes 
and Townes Op. cit. supra, pp. 17, 18). 

detritus (Holmgren). Host: Cephus cinctus Nort. 

Taxonomy and/or Biology: Salt 1931. Bui. Ent. Res. 22 (4): 519-526 
figs.— Holmes 1953. Canad. Ent. 85: 339. 



62 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

♦productus Walley. Alaska, Yukon, N. W. T., N. S., south to Calif. (Mts.), 
Colo., and R. I. 
Scambus (Endromopoda) productus Walley, 1960. In Townes and 
Townes, U. S. Natl. Bus. Bui. 216 (Pt. 2) : 19, 23. 9, $. 

*rube8cens Walley. Ont. to N. Y. west to Sask., Wyo., and Kansas. 

Scambus (Endromopoda) rubescens Walley 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 18, 19. S, 9. 

SCAMBUS subgenus ATELEOPHADNUS Cameron 

Pseudopoemenia Kiss. (Syn. by Walley, 1960. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 2) : 17, 18, 27.) 

*deceptor Walley. Que. to B. C, south to R. I., N. Y., Mich., Colo., Wash. 

Scambus (Ateleophadnus) deceptor Walley, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) 28, 35. $, 9. 

*granuIosus Walley. N. W. T. to N. S. south to Calif., N. Mex., S. C. Host: 
Mompha eloisella (Clem.). 
Scam.bus (Ateleophadnus) granulosus Walley, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 28, 31. $, 9. 

pterophori (Ashmead). Alaska, Yukon. 

SCAMBUS subgenus SCAMBUS Hartig 

alboricta (Cresson). Transf. To Acropimpla Townes. 

aplopappi (Ashmead). Idaho. Hosts: Stomopteryx nigrella (Chamb.), 
Periploca ceanothiella (Cosens). 

atrocoxalis (Ashmead). Yukon, N. W. T., Newfoundland, Alta.; Sweden. 
Male unknown {teste Walley, 1960. In Townes and Townes, p. 
53). 

btcoloripes (Ashmead). Syn. of brevicomis (Gravenhorst). 

*brevicornis (Gravenhorst). Alaska, Yukon to Newfoundland, south to Calif., 
Utah, Colo., Mich., N. Y., N. J. Hosts: Anthophila pariana 
(Clerck) ; Choristoneura rosaceana (Harris) ; Laspeyresia nigri- 
cana (Steph.) ; Laspeyresia sp.; Polychrosis viteana (Clem.) Spi- 
lonota occellana (D. & S). 
Epiurus bicoloripes Ashmead. (Syn. by Walley, 1960. In Townes and 
Townes, p. 55.). 

^canadensis Walley. Alaska, Yukon, N. W. T., B. C. to N. B. and N. S., south 
to Oreg. and Maine, Colo. Hosts: Choristoneura conflictana 
(Wlkr.) ; Glypta fumiferanae (Vier.). 
Scambus (Scambus) canadensis Walley, 1960. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 2) : 38, 39, 58. $, 9. 

*decorus Walley. N. S. to B. C, south to N. H. and Wash., Alaska. Hosts: 
Amorbia humerosa (Clem.); Choristoneura fumiferana (Clem.); 
Coleophora pruniella Clem.; C. salmani Heinr. Chrysopora sti- 
pella var. naeviferella (Dup.) ; Gracillaria sp. ; Paraclemensia ac- 
erifoliella (Clem.) ; Phalonia rutilana (Hbn.) ; Spilonota ocellana 
(D. & S.). 
Scambus (Scambus) decorus Walley, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 37, 40, 75. $, 9. 

*dioryctriae Walley. Alta., Sask. Host: Dioryctria reniculella (Grote). 

Scambus (Scambus) dioryctriae Walley, 1960. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 2) : 38, 40, 61. $, 9. 

ephialtoides Cushman. B. C, Oreg. Host: "anthomyid larva." 

Taxonomy: Walley, 1960. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 2), pp. 42-43, 46 (under occidentalis) . 

euurae (Ashmead). A subsp. of vesicarius (Ratzeburg). 



ICHNEUMONIDAE 63 

hispae (Harris). Alaska, Yukon, N. W. T. Hosts: Acrobasis betulella Hist.; 
A, comptoniella Hist.; A. nibrifasciella Pack.; Anacampsis inno- 
ciiella (Zell.) ; Anisota nibicunda (F.) ; Archips rosanus (L.) ; 
Choristoneufa pinus (Free.) ; Dioryctria reniculella (Grt.) ; Epi- 
simus argutamis (Clem.); Exartema connectum (McD.) ; Hedia 
variegana (Hbn.) ; Laspeyresia molesta Busck; Lexis bicolor 
(Grt.) ; Malacosoma disstria (Hbn.) ; Metallus rohweri MacG.; 
Psilocorsis sp. ; Spilonota ocellana (D. & S.) ; Tischeria sp.; Arth- 
rochlamys bebbianae Brown ; Xenochalepjis dorsalis Thnbg. 
Pimpla (Epiurus) nigrifrons Viereck. Remove from syn., a valid species. 
Epiurus innoniinaius Viereck. Place as syn. of transgressus (Holmgren). 
Biology: Arthur, 1963. Canad. Ent. 95 (10): 1085-1086 (life history, 
imm. stages). 
*imparis Walley. B. C, Alta., Wash., Calif., Utah, Colo. Hosts: Recurvaria 
milleri Busck; Zelleria haimbachi Busck. 
Scambus (Scambus) imparls Walley, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 37, 38, 51. $, 9. 
lithocolletidis (Ashmead). "It seems to belong in the nominate subgenus." 

(Walley, 1960. In Townes and Townes, p. 15.) 
*Iongicorpus longicorpus Walley. ^ Alaska, Alta., Sask, Ont., Que., N. H. 
Hosts: Dioryctria reniculella (Grote) ; ?Laspeyresia yoiingana 
(Kft.), IHylemyia anthracina (Czerny). 
Scambus (Scambus) longicorpus longicorpus Walley, 1960. In Townes 
and Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 37, 38, 43, 44. $, 
9. 
*longicorpus occidenlalis Walley. B. C, Wash., Calif., Mont., Colo. 

Scambus (Scambus) longicorpus occidentalis Walley, 1960. In Townes 
and To%vnes, U. S. Natl. Mus. Bui. 216 (Pt. 2): 43, 46. $, ?. 
nigrifrons (Viereck). N. B., P. E. I., N. S., Mass., Mich. Host: Hemero- 
campa leucostigma (A. & S.) ; "Orgyia" sp. ; tussock moth. 
Removed from syn. of hispae Harris. A valid species {teste Walley, 
1960. In Townes and Townes, pp. 37, 40, 74). 
*nucum (Ratzeburg). Alaska, N. W. T., Que., N. S., N. H., B. C. Host: 
Spilonota ocellana (D. & S.). 
Pimpla nucuni Ratzeburg, 1844. Ichn. d. Forstins. v. 1, p. 115. $. 

*portentus Walley. B. C, Wash., Colo. Host: ILaspeyresia sp. on Picea en- 
gehnanni. 
Scambus (Scambus) portentus Walley, 1960. In Townes and Townes, U. 
S. Natl. Mus. Bui. 216 (Pt. 2) : 37, 38, 40. $, ?. 
*subtilis Walley. Oreg., Calif., Nev. 

Scambus (Scambus) subtilis Walley, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 39, 47. ?. 

lecumseh Viereck. N. S. to Alaska south to N. C. and Calif. Hosts: Acleris 
sp. on Cornns; Archips argyrospilus (Wlkr.) ; Choristoneura 
conflictana (Wlkr.) ; C. fumiferana (Clem.) ; Croesia albicomana 
Clem.); Endothenia hebesana (Hbn.); Exotelia dodecella (L.) ; 
Lyvinaecia phragmitclla (Staint.) ; Malacosoma pluviale (Dyar) ; 
Phalonia rutilana (Hbn.); Recurvaria sp. ; Rhyacionia buoliana 
(Schiff.) ; Sparganothis sulplnireana Clem.; Spilonota ocellana 
(D. & S.) ; Trichotaphe levisella (Fyles). 
Biology: Arthur, 1963. Canad. Ent. 95 (10): 1085-1086 (life history, 
imm. stages). 

*tenebrosus Walley. Wash., Ariz., Colo. 

Scambus (Scambus) tenebrosus Walley, 1960. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 2) : 38, 39, 60. $, ?. 

transgressus (Holmgren). B. C, Wash., Oreg. Hosts: Acleris variana 
(Clem.) ; Anthophila pariana (Clem.), Archijis rosanus (L.), Cho- 
ristoneura fumiferana (Clem.), Grapholitha packardi (Zell.); ne- 
matine stem gall on Salix. 
Epiurus innominatus Viereck. Removed from syn. of hispae (Harris) 
(teste Walley, 1960. In Townes and Townes, p. 65). 



64 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

vesicarius euurae (Ashmead). Changed status. Alaska to Calif., Ariz., Colo., 
east to N. W. T., Ont. Que., Minn. Hosts: Pontania sp., P. 
calif ornica Marl., P. ?desmodiodes Walsh; P. pacifica Marl., Euura 
sp., Nematus sp. 
Scambus euurae (Ashmead). Reduced to subsp. rank (teste Walley, 
1960. In Townes and Townes, p. 50). 

♦vesicarius vesicarius (Ratzeburg). Alaska. 

Pimpla vesicaria Ratzeburg, 1844. Ichn. d. Forstins., v. 1, p. 115. $,9. 

*SCAMBUS subgenus ERYTHROSCAMBUS Walley 

Scambus subg. Lissoscambus Walley, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 17, 25. 

Type: Scambus (Lissoscambus) arizonensis Walley. Orig. 
desig. and by monotypy. 

*arizonensis Walley. Colo., Ariz., Mexico. 

Scambus (Lissoscambus) arizonensis Walley, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2): 25. $, ?. 

♦SCAMBUS subkerus ERYTHROSCAMBUS Walley 

Scambus subg. Erythroscambus Walley, 1960. In Townes and Townes, 
1960. U. S. Natl. Mus. Bui. 216 (Pt. 2) : 17, 77. 

Type : Pimpla hirticauda Provancher. Orig. desig. 

hirticauda (Provancher). B. C, Calif., Ariz. Hosts: Gracillaria sp. on Qu- 
ercus garryana; Malacosoma pluviale (Dyar.). 

Genus ALOPHOSTERNUM Cushman, p. 187 

foliicola Cushman. W. Va., Calif. Hosts: Xenochalepus dorsalis (Thnbg.), 
Paraclemensia acerifoliella (Fitch), "Cameraria aceriella?" (in 
Maine). 

Genus CALLIEPHIALTES Ashmead, pp. 187-188, Sup. 1, p. 37 
comstockii (Cresson). Transf . to Exeristes Foerster. 

ferrugineus Cushman. Hosts: Hypsipyla grandella (Zell.) and Pectinophora 
gossypiella (Saund.) m Puerto Rico. 

grapholithae (Cresson). Ont. Hosts: Acrohasis caryae (Grt.) ; A. hebescella 
Hulst; A. rubrifasciella Pack.; Conotrachelus nenuphar (Herbst); 
Stagmatophora ceanothiella Cos.; Thyridopteryx ephemeraeformis 
(Haw.); Phylloxera gall; ICarmenta texana; moth on Fagus 
nuts; shucks of Gary a nuts. 

notandus (Cresson). Transit. Zone. YioB.t : Epihlema desertana ZeW. 

nubilipennis (Viereck). Transf. to Dolichomitus as subsp. of terebrans Ratz. 

nucicola Cushman. Transf. to Apistephialtes Seyrig. 

petulca (Cresson). Transf. to Apistephialtes Seyrig. 

stultor (Davis). Transf. to Apistephialtes Seyrig as syn. of variatipes (Pro- 
vancher). 

thurberiae Cushman. Host: Gall of Disholcaspis sp. on Quercus oblongifolia. 

variatipes (Provancher). Transf . to Apistephialtes Seyrig. 

♦Genus APISTEPHIALTES Seyrig 

*Apistes Seyrig, 1927. Eos 3(2) : 221. Preocc. 

Type: (Apistes perversus Seyrig) = ascaniae (Rudow). By 
monotypy. 
* Apistephialtes Seyrig, 1928. Eos 4 (3-4) : 380. New name. 
Place after Calliephialtes Ashmead, p. 188 of Catalog; Sup. 1, p. 37. 



ICHNEUMONIDAE 65 

*arcticus (Roman). Alaska, N. W, T.; Siberia. 

Ephialtes arcticus Roman, 1914. Acad. Imp. Sci. St. Petersbourg Mem. 
ser. 8, cl. Phys. Math. 29 (7) : 6. ?. 
*coracinus Townes. N. Y., N. J., Md., N. C, Tex. 

Apistephialies coracinus Townes, 1960. In To^vnes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 88, 89, 99. $, 9. 
*dentatus Townes. B. C. and Alta. to Calif., Colo. Hosts: Enoclerus 
sphegeus (F.), Thanashnus nigriventris (LeC). 
Apistephialies deniatus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 88, 97. $, 9. 
*niasoni Townes. Alaska, Yukon, N. W. T.. Alta., Man., Ont. 

Apistephialtes masoni Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 89, 93. 9. 
nucicola (Cushman). B. C, Wash., Ariz., N. Mex., Colo. Hosts: Andricus 
galls; "Cynips maculipennis'" ; Disholcaspis eldoradensis (Beut.) ; 
cynipid gall on oak; nuts of Corylus; Pontania gall on salix; 
Gnorhnoschema bacharisella Busck; Periploca ceanothiella (Co- 
sens) ; Proteoteras arizonae Kft. 
Calliephialtes nucicola Cushman (transf. by Townes and Townes, 1960, 
pp. 88, 89, 95). 
petulcus (Cresson). Tex., Mex. 

Calliephialtes petulca (Cresson) (transf. by Townes and Townes, 1960, 
pp. 88,89,97). 

tenuiventris (Holmgren). Hosts: Melanophila consputa Lee, Xiphydria sp.; 

sawfly in bark. 
Pimpla tenuiventris (Holmgren) (transf. by Townes and Townes, 1960, 

pp. 88, 89). 
Ephialtes pusio Walsh (transf. and syn. by Townes and Townes, 1960, p. 

89). 

variatipes (Provancher) . Ont. to N. J. west to Minn. Host: Cariarsia ulmiar- 

rosorella (Clem.). 
Calliephialtes variatipes (Provancher) (transf. by Townes and Townes, 

1960, pp. 88, 89, 91). 
Calliephialtes stultor (Davis) (transf. and syn. by Townes and Townes, 

1960, p. 92). 

Genus EPHIALTES Gravenhorst, Sup. 1, p. 37 
IzzPimpla of Catalog, pp. 188-189) 

Closterocerus Hartig. Transf. to syn. of Dolichoniitus Smith. 
Dolichomitus Smith. Valid genus. 

Diclosterocerus Viereck. Transf. to syn. of Dolichomitus Smith. 
Exeristeroidea Viereck. Place as syn. of Dolichomitus Smith. 

brevis Morley. Alta. to Que. Hosts: Megachile inermis Prov., M. nivalis 
Friese, M. sp. 

cephalotes (Holmgren). Transf . to Dolichomitus Smith. 

*decumbens (Townes), n. comb. B. C, Calif., Colo., Iowa. 

Pimpla decumbens Townes, 1960. hi Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2): 102. $, 9. 

dolichosoma Viereck. Transf. to Dolichomitus Smith. 

gigas Walsh. Transf. to Dolichomitus Smith as syn. of messor perlongus 

(Cresson). 

imperator (Kriechbaumer) . Transf. to Dolichomitus Smith. 
irritator (Fabricius). Transf. to Dolichomitus Smith. 

macer Cresson. Host : "N. brevicomis" in Pinus pondcrosa. 

Taxonomy (identity of species) : Townes and Townes, 1960, pp. 104-105 
(under Pimpla) . 
perlongus (Cresson). Transf. to Dolichomitus Smith as subsp. of messor 

(Gravenhorst). 
pterelas (Say). Transf. to Dolichomitus Smith. 



66 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

pusio Walsh. Transf. to Apistephialtes Seyrig as syn. of tenuiventris (Holm- 
gren). 

pygmaeus (Walsh). Transf. to Dolichomitus Smith. 

*spatulata (Townes) n. comb. Maine and N. S. to N. C, west to B. C. and 
Calif. Hosts: Ancistrocerus lineativentris Cameron; Rygchium 
sp. 
Pimpla spatulata Townes, 1960. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 2) : 102, 106. $, 9. 

tenuiventris Holmgren. Transf. to Apistephialtes Seyrig. 

♦Genus ANASTELGIS Townes 

Anastelgis Townes, 1960. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 2) : 10, 109. 

Type: Anastelgis terminalis Townes. Orig. desig. 

Place after Ephialtes Gravenhorst, Sup. 1, p. 37 ( = Pimpla of Catalog, 
p. 189). 

*terminalis Townes. Wash., Calif. 

Anastelgis terminalis Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 109, 110. $, 9. 

Genus DOLICHOMITUS Smith 

Closterocerus Hartig. (Preocc.) Transf. from syn. of {Pimpla of Cata- 
log) = Ephialtes Grav., p. 188. 

Dolichomitus Smith. Transf. from syn. of {Pimpla of Catalog) = 
Ephialtes Grav., p. 188. Valid genus. 

Disclosterocerus Viereck. Transf. from syn. of {Pimpla of Catalog) = 
Ephialtes Grav., p. 188. 

Exeristoidea Viereck. Transf. from syn. of {Pimpla of Catalog) = 
Ephialtes Grav., p. 188. 

Place in Ephiahini on p. 189 after Anastelgis Townes {teste Townes and 
Townes, 1960, p. 111). 

*aciculatus (Hellen). Yukon, B. C, Que., Europe. Host (in Europe): Tetro- 
pium gahrieli Weise in Larix europea. 
Ephialtes aciculatus Hellen, 1915. Acta Soc. Fauna et Flora Fennica 40: 
23,25. 9. 

*buccatus Townes. B. C, Oreg., Colo. 

Dolichomitus buccatus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 115, 141. 9. 

*californicus Townes. B. C. to Calif., Idaho, Ariz., Utah, Colo., N. Mex. 
Hosts: Oeme costata Lee, ISemanotus nicolas (White). 
Dolichomitus californicus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 113, 116, 146. $, 9. 

cephalotes (Holmgren). Que., Sask. 

Pimpla cephalotes (Holmgren) (transf. by Townes and Townes, 1960, 
pp. 115, 127). 

*cuspidatus Townes. Calif. Host: Monocha'tnus maculosus Hald. 

Dolichomitus cuspidatus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 113, 114, 120. $, 9. 

dolichosoma (Viereck). Ga., Miss., Kans., Colo. 

Pimpla dolichosoma (Viereck) (transf. by Townes and Townes, 1960, pp. 
113, 114, 116). 

*flexilis Townes. Yukon, N. W. T., Man. 

Dolichomitus flexilis Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 115, 142. 9. 

*foxleei Townes. B. C, Wash., Oreg., Calif., Alta., Idaho, Mont., Colo., Ont., 
Que., N. S. Maine. Host: Acanthocinus sp. 



ICHNEUMONIDAE 67 

Dolichomitus foxleei Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 116, 145. ?. 

imperator (Kriechbaumer) . Transcont. in Hudsonian, Canad. and Transit. 
Zones. Hosts: Monochamus sp.; ? borers in conifers. 
Pimpla imperator (Kriechbaumer) (transf. by Townes and Townes, 
1960, pp. 114,154). 

irritator (Fabricius). Hosts: Anoplodera proxima (Say) , Megacylleyie cai'yae 
(Gahan). Elaphidionoides incertus (Newm.), E. parellelus 
(Newm.), E. villosus (F.), Elaphidion sp. in Quercus, Graphisurus 
sp., Melandrya striata Say, Ohrium i-ufuluvi Gahan, Pachyta 
lamed (L.), Pissodes approximatus Hopk., Stenocorus lineatus 
(Oliv.), Ramosia tipuliformis (Clerck), Saperda discoidea (F.), 
Synchroa punctata Newm., Xylotrechus annosus (Say). 
Pimpla irritator (Fabricius) (transf. by Townes and Townes, 1960, pp. 
113, 114,137). 

niessor perlongus (Cresson), changed status. Hosts: Saperda calcarata Say, 
S. concolor Lee, S. moesta Lee. Dolichomitus messor niessor 
(Grav.) is European. 

Pimpla perlongus (Cresson) (transf. and reduced to subsp. by Townes 
and Townes, 1960, pp. 132, 134). 

Pimpla gigas (Walsh) (syn. by Townes and Townes, 1960, p. 134). 

*messor sparsus Townes. Yukon to Calif., east to Wyo., Colo., N. Mex. 
Hosts: Buprestis confluenta Say, Saperda moesta LeC, S. in. tu- 
lari F. & J., S. sp., Xylocrius agassizi (LeC). 
Dolichomitus messor sparsus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2): 132, 133. $, 9. 

*populneus (Ratzeburg). Ont., Sask., Colo., Wash.; Europe. Hosts: wood 
borer in Populus sp.; Saperda populnea (L.) in Germany. 
Ephialtes populneus Ratzeburg, 1848. Die Ichn. d. Forstins., v. 2, p. 100. 

pterelas (Say). Europe. 

Pimpla pterelas (Say) (transf. by Townes and Townes, 1960, p. 150). 

pygniaeus (Walsh). N. H. to N. C. west to B. C. and Calif. Host: IPlectrura 
spinicauda Mann. 
Pimpla pygmaeus (Walsh) (transf. by Townes and Townes, 1960, pp. 
113,115, 149). 

*sericeus (Hartig). Transcont. in Canad. Zone; Europe. Hosts: buprestid 
borer in Picea, ITetropium cinnamojyterum Kby. 
Closterocerus sericeus Hartig, 1847. Ber. Naturw. Ver. Harzes f. Jahre 
1846/7, col. 18. $. 

*taeniatus Townes. Calif., Ariz. Hosts: Chrysobothris edwardsii Horn. 

Dolichomitus taeniatus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 113, 114, 118. $, 9. 

terebrans nubilipennis (Viereck), changed status. Hosts: Pissodes duhius 
Rand., P. engelmanni Hopk,, P. fasciatus LeC, P. fraseri 
Hopk., P. rotundatus LeC, Pissodes spp., Scolytus sp. Typical 
terebrans (Ratz.) occurs in Europe. 
Calliephialtes nubilipennis (Viereck) (transf. and reduced to subsp. by 
Townes and Townes, 1960, p. 122). 

*tuberculalus tuberculatus (Fourcroy). Alaska, B. C to Que. and N. B., 
south to N. Y. and Oreg. Europe. Hosts: Hylobius pinieola 
(Coup.); Monochamus sp., M. scutellatus (Say). Another subsp. 
occurs in Japan. 
Ichneumon tuberculatus Fourcroy, 1785. Ent. Parisiensis, p. 385. 9. 

*vitticrus Townes. Que. and Ont. south to Pa., west to Man., Wis., Kans. 

Dolichomitus vitticrus Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 114, 116. $, 9. 



68 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Genus ACROPIMPLA Townes 

*Selenaspis Roman, 1910. Ent. Tidskr. 31 (2-3) : 191. Preocc. 

Type: Hemipimpla alboscutellaris Szepligeti. Orig. desig. 
Acropimpla Townes, 1960. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 2): 9, 159. 

Type: Hemipimpla alboscutellaris Szepligeti. By autotypy. 

Townes designated Charitopimpla leucosioma Cameron as type-species of 
Acropimpla but said "the unavailable name Selenaspis applies to 
the same zoological group as Acropimpla." Art. 67 (i), Internatl. 
Code of Zool. Nomenclature, p. 65 says ". . . Type-fixation for 
either applies also to the other, despite any statement to the con- 
trary." 

Place on p. 189 before Iseropus Foerster. 

alboricta (Cresson). 

Scambus alboricta (Cresson) (transf. by Townes and Townes, 1960, pp. 
161, 162). 

*pronexus Townes. N. B., Que., Ont., N. H., N. Y., Wyo., B. C, N. M. 
Hosts: Glypta fimiiferanae (Vier.), Choristoneura fumiferana 
(Clem.) prob. secondary. 
Acropimpla pronexus Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 161. $, ?. 

Genus ISEROPUS Foerster, pp. 189-190, Sup. 1, p. 37 

*Cnemopimpla Cameron, 1903. Ent. 36: 159 (syn. teste Townes, Townes, and 
Gupta, 1961, p. 19). 

Type: Cnemopimpla pilosa Cameron. By monotypy. 

califomiensis Cushman. Wash., Colo. Hosts: Malacosoma sp., M. fragile 
(Stretch). 

coelebs (Walsh). Hosts: Dasychira plagiata (Wlkr.) ; Hemerocampa sp., 
Malacosoma cUsstria Hbn., Metallus rohweri MacG., Olene sp. 

stercorator orgyiae (Ashmead), changed status. Transcont. in Canad. and 
Transit. Zones. Hosts: Choristoneura rosaceana (Harr.), CteJi- 
ucha virginica (Charp.), Halisidota sp., Hemerocampa sp., H. leu- 
costignia (J. E. Sm.), H. vetusta (Bdvl.), Malacosoma sp., M. 
disstria Hbn., Simyra henrici (Grt.), "pine tussock". 
Iseropus orgyiae (Ashmead) (reduced to subsp. rank by Townes and 

Townes, 1960, p. 168). 
Iseropus stercorator brunneifrons [sic] (Viereck) (syn. by Townes and 
Townes, 1960, p. 168). Change to original spelling: bruneifrons. 

Genus TROMATOBIA Foersfer, p. 190 

In type citation delete "Desig. by Viereck, 1914" and add: Desig. by 

Ashmead, 1900. 
*Austropimpla Brethes, 1913. Anal. Mus. Nac. Hist. Nat. Buenos Aires 

24: 40. 

*Type: Austropimpla huebrichi Brethes. Orig. desig. 

Syn. by DeSantis, 1956. La Plata Univ. Nac. Inst. Mus, Notas (Zool.) 
18 (165) : 305. 

lateralis (Cresson). Syn. of notator (Fabricius). 

*notator (Fabricius). Fla., N. C. 

Pimpfa nofaf or Fabricius, 1804. Systema piezatorum, p. 115. 9. 
Tromatobia lateralis (Cresson) (syn. by Townes and Townes, 1960, p. 

186). 
*Ephialte8 cressoni Dewitz, 1881. Berliner Ent. Ztschr. 25: 205. 9. 
(Syn. by Townes and Townes, 1960, p. 186.) 

*ovivora ovivora (Boheman). Transcont. in Huds., Canad., Transit, and U. 
Austral. Zones. Hosts: Araneus diadema L., A. solitaria 
(Emrtn.), Argiope avara Thor., A. argentata (F.), Cyrtophora 
califomiensis Keys. 



ICHNEUMONIDAE 69 

Pimpla ovivora Boheman, 1821. Svenska Vetensk.-Akad. Handl. (ser. 3) 

9: 336. $, 9. 
Tromatobia ovivora scriptifrons (Cresson) (syn. by Townes and Townes, 

1960, p. 179). 
*Pimpla (Iseropus) viduiformis Viereck, 1912. U. S. Natl. Mus. Proc. 43 

(1942): 591. $. (See Catalog, p. 190, in discussion of ovivora 

scriptifrons (Cresson).) 
Biology: Nielsen, 1923. Ent. Medd. 14: 194-195. 

ovivora rufopectus (Cresson), changed status. Ont., Mich., Kans., Tex. 
Hosts: Egg sacs of: Argiope trifasciata (Forskal), "Epeira" sp. 
Tromatobia rufopectus (Cresson) (reduced to subsp. rank to Townes 
and Townes, 1960, p. 183). 

ovivora scriptifrons (Cresson). Syn. of ovivora ovivora (Boheman). 

rufopectus (Cresson). A subsp. of ovivora (Boheman). 

variabilis (Holmgren). 

*Polysphinctus forsiusi Hellen, 1915. Acta Soc. Fauna Flora Fennica 40: 
45. $, $. (Syn., teste Townes and Townes, 1960, p. 176.) 

zonata (Davis). 

Taxonomy: Townes and Townes, 1960, pp. 175, 185. 

Genus ZAGLYPTUS Foerster, p. 191 

*arizonicus Townes. Ariz. (Parker Creek, Sierra Ancha). 

Zaglyptus arizonicus Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 189, 192. ?. 

*pictilis Townes. Fla., Ala., Ga., ?Cuba. 

Zaglyptus pictilis Townes, 1960. In TowTies and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 189, 194. $, 9. 

varipes inconipletus (Cresson). Transcont. in Huds. Zone. 

Genus CLISTOPYGA Gravenhorst, p. 192 

*Hymenomacropyga Uchida, 1941. Ins. Matsumurana 15 (3) : 116. 

Type: Hymenomacropyga latifrontalis Uchida. Orig. desig. 
and by monotypy. 
Transf. from tribe Polysphinctini, p. 192 (teste Townes and Townes, 

1960, p. 195). 
Biology: Nielsen, 1929, Ent. Meddel. 16: 366-368. 

*alutaria Townes. Ont., Del., N. C, Fla., Wis. 

Clistopyga alutaria Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 196, 200. $, 9. 

atraia Cushman. Host: ICalopus angustus LeC. in Pinus murrayana (in gal- 
lery of). 

canadensis Provancher. Canad. Zone, Atlantic to Rocky Mts. Hosts: tLepto- 
chillis rufinodus (Cress).; IMalacosoma disstria Hbn.; ?lep. bud- 
worm on Picea. 

maculifrons Cushman. Mass., N. Y., N. C, Ohio, Mich., Wis., Minn. 

manni Cushman. B. C, Calif., N. Mex. Hosts: Agelenopsis sp.; ?salticid 
spider. 

nigrifrons Cushman. B.C. Host: spider nest. 

pulchripicta Ashmead. Syn. of recurva (Say). 

recurva (Say). 

Clistopyga pulchripicta Ashmead (syn. by Townes and Townes, 1960, p. 
204). 

Genus Pseudorh/ssa Merrill, p. 191 
Transf. to Theroniini, p. 202. 



70 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus De/omer/sfa Foerster, p. 191 
Transf. to Theroniini, p. 202. 

Genus PERITHOUS Holmgren, p. 191 

divinator (Rossi). Que., Conn., Pa., Mich., Wis. Hosts: Pemphredonini in 
stem of Rosa, Pemphredon lethifer Shuckard. 
Biology: Thomas, 1964. Mich. Acad. Sci. Arts Letters, Papers 49: 
199-201. 

mediator neomexicanus (Viereck), changed status. Va., Oreg., Calif. Host: 
Pemphredon errans Roh. P. mediator mediator (Fab.) is Euro- 
pean. 
Perithous pleuralis Cresson of Catalog, in part. P. neomexicana removed 
from syn. and made subsp. (teste Townes and Townes, 1960, p. 
213) ; change p. 87 to p. 88 in cit. of neomexicana. 

mediator pleuralis (Cresson), changed status. Alaska, Sask. and east of 
Rocky Mts. Host: Pemphredon harbecki Roh. (?). 
Perithous pleuralis Cresson (reduced to subsp. rank by Townes and 
Townes, 1960, p. 211). N. B., neomexicana Viereck removed from 
syn. and placed as subsp. of mediator. 
Biology: Krombein, 1960. Ent. News 71: 31. 

pleuralis (Cresson). A subsp. of mediator (Fabricius) ; (neomexicana Vier- 
eck removed from syn. and made subsp. of mediator (Fabri- 
cius) . 

Tribe POLYSPHINCTINI 

Biology: Nielsen, Ent. Meddel. v. 14, pp. 137-205 (1923); v. 16, pp. 

152-155 (1928) ; pp. 366-368 (1929) ; v. 19, pp. 192-215 (1935) ; v. 

20, pp. 25-28 (1937). 
Taxonomy: Townes and Townes, 1960. U. S. Natl. Mus. Bui. 216 (Pt. 

2) : 216-279 (keys to genera and species inch). 

Genus Clistopyga Gravenhorst, p. 1 92 
Transf. to Ephialtini and place after Zaglyptus Foerster, p. 191. 

Genus ACRODACTYLA Haliday, p. 192 

Colpomeria Holmgren. Change type desig to: (Colpomeria laevigata 
Holmgren) = quadrisculpta Gravenhorst, (teste Townes and 
Townes, 1960, p. 230). 

*degener (Haliday). Transcont. in Huds., Canad., and Transit. Zones; Eu- 
rope. 

Cryptus (Acrodactyla) degener Haliday, 1838. Ann. Nat. Hist. 2(8): 
117. $, 9. 

Symphylus hadrodactylus Foerster, 1871. Naturh. Ver. Rheinlande Verh. 
28: 106. $, 9. 

Taxonomy: Townes and Townes, 1960, pp. 231, 233. 

*jubata Townes. Yukon, Wash., Colo. 

Acrodactyla jubata Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2): 231. 9. 

*ocelIata Townes. Que. to N. C, Mich. Host: Linyphia phrygiana C.L.K. 

Acrodactyla ocellata Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 231, 232. S, 9. 

*quadrisculpta (Gravenhorst). Transcont. in Huds., Canad. and Transit. 

Zones; Eurasia. Host: Tetragnatha lahoriosaYientz. 
Ichneumon quadrisculptus Gravenhorst, 1820. Mem. Accad. Sci. Torino 

24: 378. $. 
Colpomeria laevigata Holmgren. 1859. Ofvers. Svenska Vetensk. Akad. 

Forh. 16: 127. 



ICHNEUMONIDAE 71 

Colpomeria kincaidii (Ashmead) (syn. by Townes and Townes, 1960, p. 
235). 

Genus Colpomeria Holmgren, p. 192 

A syn. of Acrodactyla Haliday, p. 192. 

kincaidii (Ashmead). Syn. of Acrodactvla quadrlsculpta (Gravenhorst) , p. 
192. 

*Genus ERUGA Townes 

Eruga Townes, 1960. hi Townes and Townes, U. S. Natl. Mus. Bui. 216 
(Pt. 2) : 220, 238. 

Type: Eruga lineata Townes. Orig. desig. 

Place in Polysphinctini after Acrodactyla Haliday, p. 192. 

*atrata Townes. Oreg., Ariz., Mex. 

Eruga atrata Townes, 1960. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 2): p. 238. $, $. 

*lineata Townes. Que., Maine, N. Y., N. C, Minn., B. C, Wash., Oreg., 
Calif., Ariz. 
Eruga lineata Townes, 1960. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 2) : 238, 239. $, 9. 

*rufa Townes. Md., S. C. 

Eruga rufa Townes, 1960. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 2) : 238, 240. $, ?. 

Genus Laufeia Tosquinet, p. 192 
Syn. of Dreisbachia Townes. 

*Genus DREISBACHIA Townes 

Laufeia Tosquinet, 1903. (Preocc. by Simon, 1889.) 

Dreisbachia Townes, 1962. Ent. Soc. Wash. Proc. 64 (1) : 38. N. name. 

*navajo (Townes). Ariz.; Honduras. Host: ?Pisa?iW?? a sp. 

Laufeia navajo Townes, 1960. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 2): 221, 223, $, ?. (Transf. by Townes, 1962, loc. 
cit. supra, p. 38.) 

slossonae (Davis). Transcont. in Canad. Zone. (Transf. by Townes, 1962, loc. 
cit. supra, p. 38.) 

Genus SCHIZOPYGA Gravenhorst, pp. 192-193 

frigida Cresson. Europe. 

*Schizopyga atra Kriechbaumer, 1887. Ent. Nachr. 14: 87. $,9. (Syn. 
by Townes and Townes, 1960, p. 225.) 

pulchra Walley. Ont. to N. C. west to B. C. 

♦Genus PIOGASTER Perkins 

Piogaster Perkins, 1958. Entomologist 91: 263. 

Type : *Piogaster rugosa Perkins. Orig. desig. 

Place in Polysphinctini before Polysphlncta Gravenhorst, p. 193. 

*niaculata Townes. Calif. (Crane Flat, Yosemite Park) . 

Piogaster maculata Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 241. ?. 

Genus POLYSPHINCTA Gravenhorst, p. 193, Sup. 1, p. 37 

albipes Cresson. 

Taxonomy: Hubbard, 1885. Ins. Affecting the Orange, p. 153. 



72 HYMENOPTERA OF AMERICA NORTH OF MEXICO 



burgessii Cresson. 

Taxonomy : Townes and Townes, 1960, pp. 246, 247. 

*eboripes Townes. N. Y. to S. C, Kans. 

Polysphincta eboripes Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 246, 254. $, 9. 

koebelei Howard. Calif., Wash. Host: Epeira foliata (Four.) or E. undata 
(Oliv.). 

limata Cresson. Transcont. in Transit. Zone. Remove bruneti Provancher, a 
valid subsp., and elongata Cushman as its syn., from syn. of li- 
mata Cresson, and place as subsp. of tuberose (Gravenhorst). 

pallipes Holmgren. Delete from Catalog; in s. str. Palearc. subsp. (teste 
Townes and Townes, 1960, p. 259). 
Polysphincta (Zatypota) strigis Howard. Transf. to *Sinarachna Townes 
as valid subsp. of pallipes (Holmgren). 

tuberosa bruneti Provancher, changed status. Transcont. in Huds. and Canad. 
Zones. P. tuberosa tuberosa (Gravenhorst) occurs in Europe. 
Polysphincta limata Cresson of Catalog in part. Remove bruneti with its 
syn. elongata Cushman, and place as subsp. of tuberosa (Graven- 
horst) (teste Townes and Townes, 1960, p. 251). 

*Genus SINARACHNA Townes 

Sinarachna Townes, 1960. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 2) : 219,258. 

Type: Polysphincta pallipes Holmgren. Orig. desig. 

Place in Polysphinctini, p. 193, after Polysphincta Gravenhorst. 

'^anomala (Holmgren). N. S. to Fla., west to Alaska and Calif.; Mexico, 

Europe, Palestine. Hosts: Dictyna sub lata (Hentz). 
Polysphincta anomala Holmgren, 1860. Svenska Vetensk.-Akad. Handl. 

(N. F.) V. 3 (Pt. 10): 34. 9. 
Polysphincta dictynae Howard (transf. with its syn. minuta (Davis) 

from Zatypota Foerster, p. 194, by Townes and Townes, 1960, p. 

261). 

pallipes strigis (Howard), changed status. Transcont. in Transit., Canad., 
and Huds. Zones. iS. pallipes pallipes (Holmgren) occurs in Eu- 
rope. Host: Epeira foliata (Fourc). 
Polysphincta pallipes Holmgren (transf. to Sinarachna Townes, and 
strigis made a subsp. of pallipes, teste Townes and Townes, 1960, 
p. 260). 

Genus Hymenoepimecis Viereck, p. 193 

Remove from Catalog. No nearctic species. 
tviltii (Cresson). Transf. to *Acrotaphus Townes. 

*Genus ACROTAPHUS Townes 

Acrotaphus Townes, 1960. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 2) : 219, 256. 

Type: Epimecis tviltii Cresson. Orig. desig. 

Place in Polysphinctini on page 193 before Oxyrrhexis Foerster. 

wiltii (Cresson). L. Austral. Zone. Transf. from Hymenoepimecis Viereck 
(teste Townes and Townes, 1960, pp. 256, 267). 

Genus OXYRRHEXIS Foerster, p, 193 

In type citation after "Included" add : and designated. 

carbonator texana (Cresson). 

Taxonomy: Cushman, 1920. U. S. Natl. Mus. Proc. 58: 22. — Townes and 

Townes, 1960. U. S. Natl. Mus. Bui. 216 (Pt. 2): 242, 243-45. 
(cocoon descr.). 



ICHNEUMONIDAE 73 

Genus ZATYPOTA Foerster, p. 1 94 

*Lycorinopsi8 Haupt, 1954. Deut. Ent. Ztschr. (N.F.) 1 (H, 1/11) : 110. 
(Syn. by Townes and Townes, 1960, p. 262.) 

Type: Lycorinopsis rhombifer Haupt. Orig. desig. 

alborhombarta (Davis). Ariz., Mex. 

*arizonicus Townes. Ariz. (Santa Rita Mts.). 

Zatypota arizonicus Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 264, 271. $. 

bohemani (Holmgren). B.C. 

*cing:ulata Townes. Ont., N. B., N. S. south to S. C, Mich., Wis., Colo. 

Zatypota cingulata Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2): 264, 270. $. 

*crassipes Townes. S. C. (Greenville). 

Zatypota crassipes Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 265, 278. 9. 

dictynae (Howard). Transf. to *Smarachna Townes as syn. of anomala 
(Holmgren). 

*exilis Townes. N. H., Que. 

Zatypota exilis Townes, 1960. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 2) : 264, 269. 9. 

*favosa Townes. Calif. (Fish Camp). 

Zatypota favosa Townes, 1960. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 2) : 264, 269. ?. 

*luteipes Townes. Ont., Maine, Mass., N. C. 

Zatypota luteipes Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 265, 277. $, 9. 

nigriceps (Walsh). 

IPolysphincta nigrita Walsh. Questionable syn. (teste Townes and 
Townes, 1960, p. 273, 274). 

parva (Cresson). Que., Conn., N. C, Mich., Wis. 

*patellata Townes. B. C. to Calif. 

Zatypota patellata Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 264, 268. $, 9. 

percontatoria (Miiller). Huds. Zone. 

*Polysphincta gracilis Holmgren, 1860. Svenska Vetensk.-Akad. Handl. 

(N.F.) 3 (Pt. 10) : 32. $, 9. (Syn. by Townes and Townes, 1960, 

p. 275.) 
* Polysphincta scutellaris Holmgren, 1860, Op. cit. supra: 33. $. (Syn. by 

Townes and Townes, 1960, p. 275.) 

*wal!eyi Townes. Que., Ont., Maine, Mass., Mich. 

Zatypota tvalleyi Townes, 1960. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 2): 264, 272. $, 9. 

Tribe PIMPLINI 
(izEphialtini of Catalog) 

Genus PIMPLA Fabricius, Sup. 1, p. 38 
( = Coccygom/mus Saussure of Catalog, pp. 194—196) 

^Habropimpla Cameron, 1900. Manchester Lit. Phil. Soc. Mem. Proc. 44. 
Pt. 5(15): 96. 

Type; Habropimpla bilineata Cameron. By monotypy. 

*Phytodiaetoides Morley, 1913. Fauna Brit. India Hym. v. 3, pp. 216, 
221. 

Type: (Phytodiaetoides megaera Morley) =/7aripa/pi« (Cam- 
eron) . Orig. desig. and by monotypy. 



74 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Liotheronia Enderlein, 1919. Transf. from syn. of Theronia Holmgren, 

Cat. p. 202 {teste Townes and Townes, 1960, p. 311). 
*Dihyboplax Enderlein, 1919. Gesell. Naturf. Freunde Sitzber, p. 148. 
Type: Dihyboplax flavipenuis Enderlein. By orig. desig. 

*ISeogabunia Brethes, 1927. Ent. Mitt., 16(4) : 322. 

Type: ISeogabunia paulistana Brethes. By monotypy. 

*Pimpla subg. Oxypimpla Noskiewicz and Chudoba, 1951. Polskie Pismo 
Ent. 21 (1-4) : 42. 

Type: Pimpla turionellae Linnaeus. By monotypy. 

Generic syn. teste Townes and Townes, 1960, p. 311. 
aequalis Provancher. Host: Polychrosis vitiana (Clem.). 
aethiops Curtis. Remove from Catalog. Spm. listed from Greenland misdet. 

*annulipes Brulle. N. S. to Fla. west to Ont. and Tex. Hosts: Ac7'obasis 
caryae (Horn), Carpocapsa pomonella (L.), Grapholitha molesta 
(Busck), Gretchena bolliana (Sling.), Laspeyresia nigricana 
(Steph.), Rhyacionia buoliana (Schiff.), Salebria tenebrosella 
(Hulst.), Synanthedon pictipes (G. & R.), Thamnosphecia pyri 
(Harr.). 

Pimpla aunuUpcs Brulle, 1946. hi Lepeletier, Hist. Nat. Ins. Hym., v.4, 
p. 102. 9. 

CoccygomUnus inflaius (Townes) (syn. by Townes and Townes, 1960, p. 
346). 

aquilonia aquilonia Cresson, changed status. Newfoundland to Tenn. and 

N. C. west to Wis., Minn., Alta., Alaska. Hosts: Coleophora 

salmani Heinr., Lambdina fiscellaria (Guen.), Psilocoisis faginella 

Chamb. 

Itoplectis exareolatus Ashmead. Remove from syn. and place as valid 

subsp. {teste Townes and Townes, 1960, p. 324). 
Pimpla yakutatensis Ashmead. Remove and place as syn. of subsp. exa- 
reolatus (Ashmead). 

aquilonia exareolata (Ashmead). Alaska, B. C. to Sask., south to Calif., N. 
Mex. (Mts.), and Colo. 
Pimpla yakutatensis Ashmead. Transf. fi'om syn. of aquilonia Cresson 
{teste Townes and Townes, 1960, p. 324). 

""dimidiata (Townes) , n. comb. Tex. (Hempstead). 

Coccygomimus dimidiatus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 313, 350. $. 

ellopiae Harrington. N. Mex., Mex. Host: Eucaterva variaria Grote. 

*hesperus (Townes) n. comb. Alaska, Oreg., Calif., Ariz., Nev., N. Mex., 
Utah, Colo., Neb., S. Dak. Host: Rhyacionia frustrana biishnelli 
(Busck), Anarsia lineatella Zell., Udea rubigalis (Guen.). 
Coccygomimus hesperus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 314, 315, 338. $, 9. 

longigenalis (Cushman). A subsp. of sodalis Ruthe. 

maura Cresson. Hosts: Hemerocainpa leucostigma (J. E. Sm.), Orgyia sp. 

nuda Townes. Ont., Que., N. S. to Va. west to Mich, and Minn. 

pedalis Cresson. Alaska, N.W.T., N. Mex., Mex. Hosts: Acronicta americana 
(Harr.), A. sp., Ctenucha virginica (Charp.), Dasychira plagiata 
(Wlkr.). Epargyreus clarus (Cram.), Halisidota caryae (Harr.), 
Lambdina fiscellaria somniaria (Hulst), Leurania linita Guen., 
Orgyia antiqua badia Hy. Edw., Sciaphyla duplex (Wish.), Xylo- 
trechus sp. 
Biology: Finlayson, 1960. Canad. Ent. 92 (12) : 925. 

punicipes Cresson. Tex., Mexico, South America, and Hawaii. 

*Pimpla feralis Cresson, 1873. Acad. Nat. Sci. Phila. Proc. 25: 399. B- 



ICHNEUMONIDAE 75 

sanguinipes erjthropus Viereck. B. C. to Calif., Idaho, Utah, An/., N. Mex., 
Mexico. Hosts: Carpocap&a pomonella (L.), Diacrisia virginica 
(F.), Hemerocampa vetusta (BdvL), Hemileuca sp., Malacosoma 
sp., M. fragile (Stretch), Prochoerodes forficaria catenulata 
Grote, Tholeria reversalis (Gnen.) , Zojjhodia convohdella (Hbn.). 
Coccygoiuinius sanguinipes (Cresson) of Catalog, in part. P. erythropus 
Viereck removed from syn. and made subsp. of sanguinipes Cres- 
son {teste Townes and Townes, 1960, p. 336). 

sanguinipes sanguinipes Cresson, changed status. Tex., Mexico. Host: Ala- 
bavia argillacea (Hbn.). 
Pimpla (Pinipla) erythropiis Viereck. Removed from syn. and made 
subsp. of sanguinipes. 

sodalis longigenalis (Cushman) , changed status. Calif ., Colo. 

Coccygomimus longigenalis (Cushman). Reduced to subsp. rank {teste 
Townes and Townes, 1960, pp. 328, 330). 

sodalis sodalis Ruthe. Changed status. Alaska, Yukon, N.W.T., Man., Que., 
Iceland, Greenland, Europe. 
Coccygomimus sodalis (Ruthe) reduced to subsp. rank {teste Townes and 
Townes, 1960, p. 328). 

*stri<'klandi (Townes), n. comb. Alta., Sask., Ont., Que., Mich. 

Coccygomimus siricklandi Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2): 314, 315, 340. $, $. 

lenuicornis Cresson. Alta., B. C. Host: Seiniothisa sp. 

*varians (Townes) n. comb. Rocky Mts. to Pacific Ocean in Canad. and 
Transit. Zones. Host: Carpocapsa pomonella (L.). 
Coccygomimus varians Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 314, 333. ?5, 9. 

Genus APECHTHIS Foersfer, Sup. 1, p. 38 

(=:£phia/fes of Catalog, p. 196) 

Apechtis Thomson. Add: "Emend." after bibliographic citation in Cata- 
log. Delete type citation. 
*Taiwatheronia Sonan, 1936. Nat. Hist. Soc. Formosa, Trans. 26: 256. 
N. syn. 

Type: (Taiwatheronia mahasenae Sonan) ^taiwana Uchida. 
By orig. desig. and monotypy. 

*Parapechthis Blanchard, 1936. Anal. Soc. Cient. Argentina 122: 404. 
N. syn. 

Type : Parapechthis bazani Blanchard. By orig. desig. and 
monotypy. 

Taxonomy: Townes, 1957, Proc. Ent. Soc. Wash. 59 (3) : 118-119; 
Townes and Townes, 1960, p. 302 {Ephialtes Schrank). 

annulicornis (Cresson). Que., Ont., Alta., Nebr., Wis. Hosts: Choristonexira 
fumiferana (Clem.), C. rosaceana (Harr.). 

componoius (Davis). Idaho, Calif. Host: Hemerocampa pseudotsugata McD. 

Ontario (Cresson). Ariz., Calif. 

pacificus Cushman. B. C, Oreg. 

picticornis (Cresson). Hosts: Choristoneura conflictana Wlkr., C. rosaceana 
(Harr.), Sciaphila sp.; geometrid; noctuid. 
*Apechihis ornatulus Walley, 1929. Canad. Ent. 61(8) : 190. $. (Syn. by 
Townes, 1940. Ent. Soc. Amer. Ann. 33(2): 313; Townes and 
Townes, 1960, p. 803.) 

Genus ITOPLECTIS Foerster, pp. 196-198, Sup. 1, p. 38 

Change type citation to: Ichneumon scanicus Villers. Desig. by Viereck, 
1914 {teste Perkins, 1962, p. 433). 



76 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Exeristesoide8 Uchida, 1928. Jour. Fac. Agr. Hokkaido Imp. Univ. 25 
(Pt. 1) : 37, 51. (Syn. by Townes, 1957. Ent. Soc. Wash. Proc. 59: 
107.) 

Type: (Pimpla spectabilis Ma.tsumur a) =:alternans specta- 
bills (Matsumura). Orig. desig. and by monotypy. 

behrensii (Cresson). "Western Calif, from Marin Co. southwards". 

clavicornis (Thomson). Delete from Catalog, not in Nearctic region. Species 
considered this, is ?fustiger Townes (teste Townes and Townes, 
1960, p. 299). 

conquisitor (Say). B. C. Hosts: Anisota rubicunda F., Choristoneura conflic- 
tana Wlkr., Dasychira plagiata Wlkr. ; Dioryctria reniculella 
(Grt.), Epermenia cicutaella Kft., Exyra semicrocea (Gn.), Hya- 
loscotes pithopoera (Dyar), Nematocampa limbata, (Haw.), Onci- 
deres cingidata (Say), Siniyra henrici (Grt.), Sparganothis flavi- 
basana Fern., S. sp. on Lonicera, Tortrix allenina Fern., T. cock- 
erellana Kft., T. pallorana Rbnsn., Urbanus proteus (L.), Hypera 
postica (GyW.) . 
Biology: Leius, 1961. Canad. Ent. 93 (9): 771-780.— Arthur, 1963. 
Canad. Ent. 95 (10) : 1086-1090 (also imm. stages). 

curticauda brevacus Townes, changed status. Canad. Zone, Atlantic Ocean to 
Mont. Removed from syn. of curticauda (Kriechbaumer) and 
placed as valid subsp. (teste Townes and Townes, 1960, p. 301). 

curticauda curticauda (Kriechbaumer), changed status. Delete from Catalog, 
p. 198. This subsp. is European. Remove brevacus Townes from 
syn. as a valid subsp. 

evetriae Viereck. Transcont. in Canad. Zone. Hosts: Barbara mappana 
Free., B. sp., Choristoneura fumiferana (Clem.). 

*fustiger Townes. Transcont. in Transit. Zone. Host: "Neviroptera". 

Itoplectis fustiger Townes, 1960. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 ,Pt. 2): 282, 298. $, 9. 

quadricingulatus (Provancher) . Hosts: Croesia albicomana (Clem.); Dioryc- 
tria reniculella (Grt.), Euceratia castella Wlsm., Galleria mellon- 
ella F., Mnlacosoma sp., M. disstria Hbn., Nepytia phantasmaria 
(Stkr.), Platyptilia sp., Taniva albolineana (Kft.). 

*vesca Townes. Alaska to Calif., Ariz., N. Mex., Colo., Maine, P.E.I. N. B., 
N. S. Hosts: Acleris variana (Fern.), Colephora laricella 
(Hbn.), Recurvaria milleri Busck. 
Itoplectis vesca Townes, 1960. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 2) : 282, 297. S, 5. 

viduata (Gravenhorst) . B. C, Alta., N.W.T., Sask., Wyo. Hosts: Ancylis 
coniptana (Frol.), Malacosoma disstria Hbn. 

Tribe NEOXORIDINI 
{zzPoemeniini of Catalog) 

Genus Diacrifus Foerster, p. 1 98 

Remove from (Poemeniini of Catalog) = Neoxoridini, Sup. 1, p. 38, and 
place provisionally in Microcleptinae (= Plectiscinae of auctt.), on 
p. 342. 

Genus POEMENIA Holmgren, Cat. p. 199, Sup. 1, p. 39 

Revision: Habeck and Townes, 1960. In Townes and Townes, U. S. Natl, 
Mus. Bui. 216 (Pt. 2) : 376-387. 

albipes (Cresson). Alta, 

americana americana (Cresson), changed status, N. S., Ont., N. H. Host: 
Passaloecus mandibularis (Cress.). 



ICHNEUMONIDAE 77 

Poemenia americana (Cresson). Reduced to subsp. rank {teste Habeck 
and Townes, 1960. In Townes and Townes, p. 384). 

♦americana nebulosa Habeck and Townes. B. C. to Calif, east to Colo.; Ont. 
Hosts: Laspeyresia toreuta (Grt.), Parativiia conicola Fahr. 
Poemenia americana nebulosa Habeck and Townes, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216: 382 (key), 383. $,9. 

*pacifica Habeck and Townes. Wash, to Calif. Host: Galls of Disholcaspis 
truckeensis (Ashm.). 
Poemenia pacifica Habeck and Townes, 1960. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (key), 385. $, 9. 

thoracica (Cresson). B. C. to Calif, east to Sask. and Wyo. 

Poemenia vancouveriensis Provancher (syn. by Habeck and Townes, 
1960. In Townes and Townes, p. 377). 

vancouveriensis (Provancher). Syn. of thoracica (Cresson). 

Genus NEOXORIDES Clement, p. 199 

borealis (Cresson). Hosts: Melanophila drummondi (Kby.), Laspeyresia to- 
reuta (Grt.). 

caryae (Harrington). Ont., Que. 

♦pilulus Townes. Yukon to Oreg., east to N.W.T., Que., N. B., Colo., also N. Y., 
N. C, N. J. Host: buprestid. 
Neoxorides pilulus Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216: 390. 9, $. 

vittifrons (Cresson). Transf. to *Podoschistus Townes. 

*Genus PODOSCHISTUS Townes 

Podoschistus Townes, 1957. Ent. Soc. Wash. Proc. 59(1) : 16 (key), 18. 
Type: Xorides vittifrons Cresson. Orig. desig. 

Place on p. 199 of Catalog after Neoxorides Clement. 

*vittifrons schlingeri Townes. Utah (Mantua). 

Podoschistus vittifrons schlingeri Townes, 1960. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 2) : 388. ?. 

vittifrons vittifrons (Cresson), changed status. Maine to Ga., west to Ont. and 
Tex. Host: buprestid. 
Neoxorides vittifrons (Cresson). Transf. from Neoxorides and reduced to 
subsp. rank {teste Townes and Townes, 1960, pp. 388, 389). 

Tribe RHYSSINI 

Genus RHYSSA Gravenhorst, p. 200, Sup. 1, p. 39 

alaskensis Ashmead. Wyo., Huds., Canad., and Transit. Zones. 

Rhyssa skinneri Viereck. Change bibliographic reference page from 86 
to 87. 

crevieri (Provancher). N. S., N. B. Que., Ont., Maine, N. H., N. Y., N. J., 
N. C. (Mts.), Mich., Minn. Host: Urocerus albicornis (F.). 
Rhyssa lineolata (Kirby) of Catalog, in part. Remove crevieri from syn. 
{teste Townes and Townes, 1960, p. 403). 

hoferi Rohwer. Ariz., N. Mex. 

*howdenorum Townes. Va., N. C, Nebr. 

Rhyssa howdenorum Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 396, 398. $, ?. 

lineolata (Kirby). Upper Austral. Zone. (Townes, 1960, p. 406 says "but 
there are no definite host records.") 
Rhyssa lineolata (Kirby). Remove crevieri (Provancher), a valid sp., 
from syn. {teste Townes and Townes, 1960, p. 403). 

persuasoria (Linnaeus). Host: Sirex areolatus (Cress.). 



78 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*ponderosae Townes. Calif. 

Rhyssa ponderosae Townes, 1960. In Townes and Townes, U. S. Natl, 
Mus. Bui. 216 (Pt. 2) : 397, 401. $, ?. 

Genus RHYSELLA Rohwer, p. 200 

humida (Say). Wis., Minn., Iowa. 

nitida (Cresson). Canad., Transit., Upper Austral. Zones. Que., Ont., Minn., 
Tenn., S. C, also B. C. 

*Genus EPIRHYSSA Gresson 

Epirhyssa Cresson, 1865. Ent. Soc. Phila. Proc. 4 : 39. 

Type: Epirhyssa speciosa Cresson. Desig. by Viereck, 1914. 

^Rhyssonota Kriechbaumer, 1890. Naturh. Hofmus. Wien. Ann. 5(3): 
489. Syn. by Townes and Townes, 1960, p. 413. 

Type: Rhyssonota tristis Kriechbaumer. By monotypy. 

Place before Megarhyssa Ashmead, p. 200. 

*dietrichi Townes. Ariz. (Mt. Lemmon). 

Epirhyssa dietrichi Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 414. $, 9. 

Genus MEGARHYSSA Ashmead, pp. 200-202 

Biology: Heatwole, Davis, Wenner, 1964. Amer. Midi. Nat. 71 (2) : 
374-381. 

atrata atrata (Fabricius), changed status. Conn., Ont. south to Fla. and Tex. 
Megarhyssa atrata (Fabricius). Reduced to subsp. rank {teste Porter, 
1957. Ent. News 68: 206). 

*atrata lineata Porter. N. S. to Ont. and south to Penna. Host: Tremex 
columba (L.). 
Megarhyssa atrata lineata Porter, 1957. Ent. News 68: 206, ?, 

*greenei floridana Townes. Fla. 

Megarhyssa greenei floridana Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 421, 424. 9. 

greenei greenei Viereck. Canad. to Upper Austral. Zone in east. No. Amer- 
ica. 
Megarhyssa greenei Viereck. Reduced to subsp. rank (teste Townes and 
Townes, 1960, p. 422). 

icterosticta Michener. A subsp. of macrums (Linnaeus). 

macrurus icterosticta Michener, changed status. Ariz. 

Megarhyssa icterosticta Michener. Reduced to subsp. rank {teste Townes 
and Townes, 1960, pp. 430, 435). 

macrurus lunator (Fabricius). Canad., Transit, and Upper Austral, zones 
east of 100°W. Host: Tremex columba (L.). 
Megarhyssa macrurus macrurus (Linnaeus) of Catalog, in part. Place 
lunator, and its syn. histrio, as valid subsp. of macrurus {teste 
Townes and Townes, 1960, p. 430). 

macrurus macrums (Linnaeus). Place lunator, and its syn. histrio, as valid 

subsp. of macrurus. 
*Ichneumon Georgicus Megerle, 1802. Appendix ad catalogum insecto- 

rum, quae mense Novembris 1802 Vienne Austriae auctionis lege 

vendita fuere, p. 16 (Syn. teste Townes and Townes, 1960, p. 

434). 
Megarhyssa macrurus phaeoptila Michener (syn. by Townes and Townes, 

1960, p. 434). 

macrurus phaeoptila Michener. A syn. of M. macrurus macrurus (Linnaeus). 
nortoni nortoni (Cresson). Ont., Mich., Vt. 
nortoni quebecensis (Provancher). P.E.I., Tenn. 



ICHNEUMONIDAE 79 

Tribe THERONIINI 

Genus PSEUDORHYSSA Merrill, p. 191 

Change type desig. to read as follows: Type: Pseudorhyssa sternata Mer- 
rill = ruficoxis (Kriechbaumer). Orig. desig. 

Transf. from Tribe Ephialtini, p. 191 {teste Townes and Townes, 1960, p. 
373). Place after heading Tribe Theroniini. 

*ruficoxis (Kriechbaumer). Host: Siricidae in conifers; ? also Coleoptera. 
"Rhyssa approximator (Fabricius)" of authors, not of Fabricius. 
Rhyssa approximator var. ruficoxis Kriechbaumer 1887. Ent. Nachr. 13: 

250. 9. 
Pseudorhyssa sternata Merrill (syn. by Townes and Townes, 1960, p. 
373). 
«/ernaf a Merrill. Syn. of ruficoxis (Kriechbaumer). 

Genus THERONIA Holmgren, pp. 202-203 

*Poecilopimpla Cameron, 1903. Straits Branch Roy. Asiatic. Soc. Jour. 
39: 140. Syn. by Townes and Townes, 1960, p. 351. 

Type: Poecilopimpla lucida Cameron. By monotypy. 
Liotheronia Enderlein, 1919. Transf. to syn. of Pimpla Fabricius. 

atalantae (Poda). Delete from Catalog. Place fulvescens (Cresson), and its 
syns. rufescens Taylor, fulvescens var. niellipennis Viereck and 
atalantae var. americana Krieger, as a subsp. of atalantae. 

atalantae fulvescens Cresson, changed status. T. atalantae atalantae (Poda) 
occurs in Europe. Host: Dasychira plagiata (Wlkr.). 
Theronia atalantae (Poda) of Catalog. Remove atalantae (Poda) from 
syn., and place fulvescens as valid subsp. of atalantae {teste 
Townes and Townes, 1960, p. 354). 

bicincta (Cresson). Delete from Catalog; occurs only in Cuba {teste Townes 
and Townes, 1960, p. 362). 

*bicincta floridana Townes. Fla., Ga. Host: Tromatobia nifopectus (Cress.). 
Another subsp., bicincta bicincta (Cresson), occurs in Cuba. 
Theronia bicincta floridana Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216: 361, 362. $, ?. 

hilaris (Say). Host: Heterocampa guttivitta (Wlkr.). 
septemtrionalis (Krieger). Fla., Ohio, Okla. 

Genus DELOMERISTA Foersier, p. 191 

Change desig. of type species to: Incl. and desig. by Schmiedeknecht, 
1888. 

Place in tribe Theroniini, p. 203. 

*borealis Walkley. Alaska, Yukon, N.W.T., Que., Colo. 

Delomerista borealis Walkley, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2): 365, 370. $. 

*geUda Walkley. Alaska, N.W.T., B. C. to Que., N. Y., Pa., Colo., Ariz., 
Calif. 
Delomerista gelida Walkley, 1960. In Townes and Townes, U. S. Natl 
Mus. Bui. 216 (Pt. 2) : 365, 366. $. 

*japonica diprionis Cushman, n. status. Alaska and B. C. to Calif. (Mts.) east 
to N. W. T., N. S. and N.Y., N C. (Mts.). D. japonica japonica 
occurs in Japan. Hosts: Diprion frutetorum (F.), Neodiprion spp. 

Taxonomy: Walkley, 1960, in Townes and Townes, pp. 367-368 (as syn. of 
japonica). 

Biology: Finlayson, 1960. Canad. Ent. 92(1) : 25-26 (as diprionis). 

*lepteees Walkley. Alaska, Que., Colo. 

Delomerista lepteces Walkley, 1960, In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 365, 368. 9. 



80 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

novita (Cresson). Hosts: Maeremphytus sp.; sawfly larva in elder stem; ? 
Mononychus vulpeculus (F.) ; Acrobasis rubrifasciella Pack.; ? 
Exartema olivaceanum Riley; ? Eublevima miniina (Guen.). 

*pfankuchi (Brauns). ? W. Va., Europe. 

[Pimpla] (Delomerista) Pfankuchi Brauns, 1905. Ztschr. f. Hym. 
Dipt. 5: 131. 9. 

texana (Cresson). Alaska, N.W.T., Que., Maine, N. H.; Europe. 

*Pimpla laevifrous (!) Thomson, 1877. Opuscula Ent. 8: 750. $, 9. 

Tribe LABENINI 

Genus LABENA Cresson, pp. 203-204, Sup. 1, p. 39 

*Microtritus Kriechbaumer, 1889. Ent. Nachr. 15 : 307. 

Type : Microtritus apicalis Kriechbaumer. By monotypy. 
*Neonotus Parrott, 1955 (1954). Linn. Soc. N. S. Wales, Proc. 79 (5-6) : 
230. 

Type: ISeonotus chadtcickii Parrot. By orig. desig. and mono- 
typy. 
(Syn. by Townes and Townes, 1960, p. 531.) 

*grallator balteata Townes. Tex. (nr. Presidio). Host: Tunnels in Lycium 
berlandieri. 
Labena grallator balteata Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 532, 536. $, $. 

grallator grallator (Say). Changed status. Ont. and Conn, to Fla., west to 
Kans., Tex., also Colo. Hosts: Chrysobothris tranqiiebarica Gmel., 
Cliion sp., Saperda puncticollis Say, Thrincopyge alacris Lee. 
Labena grallator (Say). (Reduced to subsp. rank by Townes and 
Townes, 1960, pp. 532-533.) 

*grallator lutea Townes. Calif. (Needles). 

Labena grallator lutea Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 532, 536. $. 

*graIlalor ochreata Townes. Ariz. 

Labena grallator ochreata Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 532, 535. $, ?. 

*grallator xutha Townes. N. Mex., Colo. 

Labena grallator xutha Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 532, 535. $, $. 

tinclipennis Rohwer. B. C. to Calif., Ariz. 

Genus GROTEA Cresson, p. 204 
Taxonomy: Townes and Townes, 1960, p. 537 (Tribe Labiini). 
anguina Cresson. Colo., Ariz. 
califomica Cresson. B. C, Wash., Idaho, Nev., Ariz. 

Tribe XORIDINI 

Genus XORIDES Latreille, pp. 204-206, Sup. 1, p. 39 

Gonophonus Foerster. Remove from syn. A valid subg. of Xorides 

(iesfe Townes and Townes, 1960, p. 494). Palearc. Oriental. 
Moerophora Foerster. Remove from syn. A valid subg. of Xorides {teste 

Townes and Townes, 1960, p. 495). Palearc. 
Perissocerus Smith. Trans, to syn. of subg. Periceros Schulz infra. 
Cyanoxorides Cameron. Remove from syn. A valid subg. of Xorides 

{teste Townes and Townes, 1960, p. 494). Oriental. 
Spiloxorides Cameron. Remove from syn. of Xorides. A syn. of subg. 

Cyanoxorides Cameron, supra. 
Macrosterotrichus Kokujev. Remove from syn. of Xorides. A syn. of 

subg. Moerophora Foerster, supra. 



ICHNEUMONIDAE 81 

Caenostoma Cameron. Remove from syn. of Xorides. A syn. of subg. 

Gonophonus Foerster, stipra. 
Periceros Schulz. Remove from syn. A valid subg. of Xorides (teste 

Townes and Townes, 1960, p. 501). Neotrop. 
Ahyhorhyssa Enderlein. Remove from syn. of Xorides. Syn. of subg. 

*Epixorides Smith {teste Townes and Townes, 1960, p. 494). 

Oriental. 
Lavaudenia Seyrig. Remove from syn. of Xorides. Syn. of subg. *Gonio- 

prymnus Cameron {teste Townes and Townes, 1960, p. 496). Eth- 

iop. 

♦XORIDES subgenus EXOMUS Townes 

Xorides subg. Exomus Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 491, 496. 

Type: Anonialon hunieralis Say. Orig. desig. 

Place on p. 204 as first subgenus under Xorides Latreille. 

*humeralis chiricensis Townes. Ariz. (Chiricahua Mts.). 

Xorides (Exomus) humeralis chiricensis Townes, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 497, 501. $. 

*humeralis excomptus Townes. Calif. 

Xorides (Exomus) humeralis excomptus Townes, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 497, 500. $, 9. 

humeralis humeralis (Say). Canad. Zone. Hosts •.Dicerca obscura (F.), Neo- 
clytus sp. in Ilex. 
IXylonomus australis Cresson. Syn. questioned by Townes and Townes, 
1960, pp. 498, 499. 

*humeralis niexicanus Townes. Tex., Mexico. 

Xorides (Exomus) humeralis niexicanus Townes, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 497. ?. 

humeralis piceatus Rohwer. 

*hunieraHs sierrae Townes. Calif. 

Xorides (Exomus) humeralis sierrae Townes, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 497, 500. $, 9. 

XORiDES subgenus XORIDES Latreille 

* Amphixylonomus Benoit, 1952. Ann. Ent. Soc. Belg. Bui. 88: 136. Syn. 
by Townes and Townes, 1960, p. 502. 

Type: Xylonomus (Amphixylonomus) konduensis Benoit. 
Orig. desig. 

albopiotus (Cresson). 

*atrox Townes. B.C. (Robson). 

Xorides (Xorides) atrox Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 492, 520. 9. 

*boharti Townes. Calif. 

Xorides (Xorides) boharti Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 493, 508. $, 9. 

calidus (Provancher). Misspelled caWidus in catalog. 

californicus (Cresson). Ariz., Colo. Host: Paranthrene robiniae perbicida 
(Busck). 

cinclicornis cincticornis (Cresson). Changed status. Alta., Mont. (Reduced 
to subsp. rank by Townes and Townes, 1960, p. 528). 

*cincticornis rufus Townes. Calif., Oreg. 

Xorides (Xorides) cincticornis rufus Townes, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 528, 530. $, 9. 

eastoni (Rohwer). Wis., B. C. 

frigidus (Cresson). Transcont. in Canad. Zone. 



82 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

harringtoni (Rohwer). liost : Dicerca divaricata (Say). 

insularis (Cresson). Alaska, Alta., Ariz., Que. Host: Scolytid in Pseudot- 
suga taxi folia. 

maudae (Davis). Wash. 

*medius Townes. Idaho, Calif. 

Xorides (Xorides) medius Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2): 493, 511. $, 9. 

neoclyti (Rohwer). N. Hex., Colo. 

*peniculus Townes. N. Y., Pa., N. J., S. C, Tex. 

Xorides (Xorides) peniculus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2): 494, 506. $, 9. 

*pictus Townes. Calif. 

Xorides (Xorides) pictus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 493, 505. 9. 

*planus Townes. Ariz., Colo. 

Xoriiles (Xorides) planus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 493, 511. $, 9. 

rileyi (Ashmead). Ga., Fla. Host: Phymatodes testaceus (L.). 

Xorides ruficoxis (Rohwer) (syn. by Townes and Townes, 1960, p. 508). 

*rudis Townes. Calif. (Lone Pine). 

Xorides (Xorides) rudis Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 493, 510. 9. 

ruficoxis (Rohwer). Syn. of rileyi (Ashmead). 

*semirufus Townes. Fla. (Archer). 

Xorides (Xorides) semirufus Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 494, 506. 9. 

stigmapterus floridanus (Ashmead). 111. 

stigmapterus stigmapterus (Say). Transcont. in Canad., Transit., and Upper 
Austral Zones. Host: ? Dryobius sexfasciatiis (Say). (Reduced 
to subsp. rank by Townes and Townes, 1960, p. 526.) 
*Xylonomus tartarus Morley, 1913. Fauna Brit, India . . . Hym., v. 3, 
pp. 77, 78. 9. (Syn. by Townes and Townes, 1960, p. 526.) 

Genus ODONTOCOLON Cushman, p. 206 

abdominale abdominale (Cresson), changed status. Idaho, Mont., Wyo., 
Utah, Colo., Ariz. Reduced to subsp. rank by Townes and Townes, 
1960, p. 463. 

*abdominaIe fergusoni Townes. Alta., B. C, Wash. 

OdontocoJon abdominale fergusoni Townes, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 463, 464. $,9. 

aethiops (Cresson). B. C, Mont. 

alaskensis (Rohwer), Alaska. A valid species (teste Townes and Townes, 
1960, pp. 450,453,460). 

albotibiale (Bradley). N. S., Ont., Minn,, Iowa, S. C, Fla. Remove question- 
marks before cit. of striatus Cushman and aciculatus Cushman. 

atripes (Rohwer). P. E. I., Ont., Alta., Ala, 
bicolor (Cresson), Ala,, S, C, Va„ Md. 
brevicaudus (Cushman), Wash., Idaho, 
canadense (Provancher) , N, S,, Ont,, Mich,, Minn. 

*ciHpes Townes, Ariz, (Chiricahua Mts,), 

Odontocolon cilipes Townes, 1960, In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 449, 452, 454. 9. 



ICHNEUMONIDAE 83 

*curtum Townes. Calif. 

Odontocolon curtum Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 450, 453, 470. $, 9. 

*depressum Townes. Colo. (Rocky Mt. N. P.) . 

Odontocolon depressuni Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 449, 453, 459. 9. 

dichrous (Rohwer). 

Taxonomy: Townes and Townes, 1960, pp. 450, 451, 453, 454, 475-478. ?, 

S. 

*dreisbachi Townes. Mich. (Isle Royale). 

Odontocolon dreisbachi Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 450, 453, 480. 9. 

*formicoides Townes. Ariz. (Humphreys Peak). 

Odontocolon formicoides Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 449, 452, 458. $. 

*ochropus Townes . Atlantic to Rocky Mts. in Transit, and U. Austral 
Zones, also B. C, Idaho. 
Odontocolon ochropus Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 451, 454, 482. $,9. 

*parvum Townes. B. C. to Oreg., Idaho, Wyo., Colo. 

Odontocolon parvum Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 450, 452, 454, 472. $, 9. 

polymorphum Cushman. B. C, Oreg. 

*pullum Townes. B. C, Calif. 

Odontocolon pullum Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 449, 452, 456. 9, $. 

punctatum (Cushman). B. C, Wash., Oreg., Calif. Wyo. Host: "Tremex" in 
A bies. 

punctulatum (Thomson). Delete from Catalog; European species. 

Odontomerus alaskensis Rohwer. Remove from syn., valid species. 

*sierrae Townes. Calif. 

Odontocolon sierrae Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 451, 454, 471. $, 9. 

strangaliae (Rohwer). N. B., Ont., Mich., Wis., Minn., N. C, Tenn., Ala. 
Hosts: "Leptura" larvae, Strangalina luteicornis (Fab.). 

vicinum (Cresson). Upper Austral Zone. Hosts: Buprestid in Betula nigra, 
Dicerca divaricata (Say). 

♦Genus ISCHNOCEROS Gravenhorst 

Ischnoceros Gravenhorst, 1829. Ichn. Europaea, v. 2, p. 949. 

Type: Ichneumon rusticus Fourcroy. Desig. by Viereck, 1914. 

Place before genus Aplomerus Provancher, p. 207. 

'clivulus Townes. Calif. 

Ischnoceros clivnlus Townes, 1957. Ent. Soc. Wash. Proc. 59: 21. 9. 

Genus APLOMERUS Provoncher, p. 207 

*arugosus Townes and G. Townes. Wyo. (Shoshone Canyon) . 

Aplomerus arugosus Townes and G. Townes, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 441, 446. 9. 

buprestivorus Rohwer. Calif., Ariz. 

*robustus Townes and G. Townes. B. C. to Calif. Host: Buprestid. 

Aplomerus robustus Townes and G. Townes, 1960. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 2) : 440, 441, 443. $, 9. 

tibialis (Provancher). Oreg. 



84 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Tribe ACAENITINI 

Taxonomy: Townes and Townes, 1960, pp. 4, 544 (Subfamily status). 
Genus COLEOCENTRUS Gravenhorst, pp. 207-208, Sup. 1, p. 39 
flavipes (Provancher) . N. S., N. B., Ont., Tenn. (Canad. Zone). 
harringtoni Cushman. N. B., Que., N. H., Mass., Pa., Minn. 

*nianni coloradensis Townes. Colo. (nr. Estes Park). 

Coleocentrus manni coloradensis Townes, 1960. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 2) : 555, 556. $. 

manni manni Cushman. Changed status. B. C, Wash., Oreg., Calif. Re- 
duced to subsp. rank by Townes and Townes, 1960, p. 555. 

occidentalis occidentalis Cresson, changed status. Mont. (Delete Calif, from 
dist.) Reduced to subsp. rank by Townes and Townes, 1960, p. 
548. 

occidentalis pulchripennis Cushman, changed status. Calif. Reduced to 
subsp. of occidentalis, teste Townes and Townes, 1960, p. 549. 

pettiti Cresson. N. S. to Ont. and Minn, south to Pa., also Colo. 

Coleocentrus Quebecensis Provancher. Remove from syn. A valid sp. 

quebecensis Provancher. Atlantic Ocean to Rocky Mts. in Canad. Zone. A 
valid species {teste Townes and Townes, 1960, pp. 547, 550). 

rufocoxatus Pratt. N. B., P.E.I,, N. Y., Pa., N.W.T. 

rufus Provancher. P. E. I., N. S., N. B., Ont., Iowa, N. Dak., Ga. 

Genus MESOCLISTUS Foerster, p. 208 

*cushmani Townes. Alaska (Mt. McKinley N. P.). 

Mesoclistus cushmani Townes, 1960. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 2) : 560. $, ?. 

Genus AROTES Gravenhorst, pp. 208-209, Sup. 1, p. 39 

*Retanisia Cameron, 1886. Biol. Centr.-Amer. Hym., v. 1, p. 299. Syn. 
by Townes and Townes, 1960, p. 561. 

Type: Retanisia fascialis Cameron. By monotypy. 

amoenus Cresson. Host : Melandrya striata Say. 

formosus Cresson. Transf. to Spilopteron Townes. 

maurus Rohwer. B. C. to Minn. 

melleus (Say). Md., D. C, N. C, Mich. 

occiputalis Cresson. Transf. to Spilopteron Townes. 

rupinsulensis (Cresson). Transf . to Yezoceryx Uchida. 

vicinus Cresson. Transf. to Spilopteron Townes. 

♦Genus SPILOPTERON Townes 

Spilopteron Townes, 1960. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 2) : 546, 568. 

Type: Spilopteron francleinonti Townes. Orig. desig. 

Place in Acaenitini after Arotes Gravenhorst, p. 209. 

*formosum australe Townes. Md., D. C, N. C, Ga. 

Spilopteron formosuni australe Townes, 1960. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 2) : 571, 573. $, 9. 

formosum formosum (Cresson), changed status. Que. to Man. south to N. J., 
Md., W. Va., Ohio and Minn. Transf. from Arotes Gravenhorst, 
and reduced to subsp. rank {teste Townes and Townes, 1960, pp. 
571,572). 



ICHNEUMONIDAE 85 



♦franclemonti Townes. N. B., Que. to Man. south to N. Y., Ohio, Minn. 

Spilopteron franclemonti Townes, 1960. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 569. $, 9. 

occiputale (Cresson). Ga. Transf. from Arotes Gravenhorst, p. 209. {teste 
Townes and Townes, 1960, pp. 569, 576). 

*vicinum melanderi Townes. Mich, to B. C., also N. S., Oreg. 

Spilopteron vicinum melanderi Townes, 1960. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 2) : 573, 575. 5, ?. 

vicinum vicinum (Cresson), changed status, P. E. I., N. B., Que. to Man., 
south to W. Va., Minn., also N. W. T., Colo., Idaho. Host: Ano- 
plodera nigrella (Say). Transf. from Arotes Gravenhorst, p. 209 
and reduced to subsp. rank {teste Townes, 1960, in Townes and 
Townes, p. 573). 

♦Genus YEZOCERYX Uchida 

Yezoceryx Uchida, 1928. Hokkaido Imp. Univ. Faculty Agr. Jour. 25: 
36. 

Type : Yezoceryx scutellaris Uchida. By orig. desig. 

Place in Acaenitini, p. 209 before subfamily Tryphoninae. 

rupinsulensis (Cresson). Transf. from Arotes Gravenhorst {teste Townes 
and Townes, 1960, p. 578). 

Subfamily LYCORININAE 
{^Tribe Lycorini, p. 309) 

Tribe Lycorinini transf. in Sup. 1, p. 39, from p. 309 to p. 209, following 
tribe Acaenitini. 

Genus TOXOPHOROIDES Cresson, p. 309 

This group needs revision. 

albomarginatus albomarginatus (Cresson). Colo. Host: Anarsia lineatella 
Zeller. 

*apicalis (Cresson). N. C, Fla., Mo.; Mexico. 

Lycorinal apicalis Cresson, 1874 (1873). Acad. Nat. Sci. Philadelphia 

Proc. 1873: 407. $. 
Toxophoroides xanthozonatus (Ashmead). N. syn. 

glaucomatus Cushman. Ohio. 

scitulus (Cresson). Host: Gelechia albisparcella {Chamh.) , Ancylis comptana 
fragariae (W. & R.). 

xanthozonatus (Ashmead). Syn. of apicalis (Cresson). 

Subfamily COLLYRilNAE 

This subfam. transf. in Sup. 1, p. 56, from p. 341 to p. 209, following 
Pimplinae. 

Genus COLLYRIA Schiedte, p. 341 
calcitrator (Gravenhorst). Md., Pa. 

Subfamily STILBOPINAE 
{zzStilbopini of Catalog) 

Place on p. 209, before Tryphoninae. An undescribed species represent- 
ing a new genus apparently belongs here. 



86 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus STILBOPS Foerster, p. 209 

Stilbops Foerster. After 1900, in type citation, delete period and add: 

(as veluta Gravenhorst) . 
Aphanoroptruui Foerster, Remove from syn. with its emendations; 

distinct genus {teste Perkins, 1962, p. 405). 

Genus APHANOROPTRUM Foerster 

Aphanoroptruui Foerster. Change type desig. to: 

Type: (Lissonota ruficornis Gravenhorst) = abdoniinale 

(Gravenhorst). Included by Thomson, 1877. 

Remove from syn. of Stilbops Foerster. Place in subfamily Stilbopinae, 
after Stilbops Foerster, p. 209. 

Subfamily ADELOGNATHINAE 

Genus CNEMISCHYS Foerster, p. 210 

Cnemischus Thomson. Add 'Emend.' after bibliogr. ref. Delete type ci- 
tation. 

Genus PAMMICRA Foerster, p. 210 

Pamiuicrns Thomson. Delete type citation. Add: Emend. 

Subfamily PHRUDINAE 
(^Phrudini of Catalog) 

Elevated from tribal rank {teste Perkins, 1959, Roy. Ent. Soc. London, 
Handbooks: 7 (Pt. 2(ai)): 8). 

Genus PHRUDUS Foerster, p. 210 

{ — Phrudus subgenus Phrudus Foerster, p. 210) 

Elevated to generic rank {teste Perkins, 1962, p. 445). 
Replace "Desig. by Viereck, 1914" of type citation, virith: Included by 
Dalla Torre, 1901. 

exarealatus Cushman. Transf. from subg. Mengersenia Schmiedeknecht, p. 
210. 

♦Genus ASTRENIS Foerster 

( = ? Phrudus subgenus Aiengersen/o Schmiedeknecht, p. 210) 

Astrenis Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25 : 148. 

Type: Hambergiella sinuata Roman. Included and desig. by 
Perkins, 1962. 
lOrona Cameron. N. syn. 
IMengersenia Schmiedeknecht. N. syn. Perkins considers this a valid 

genus. Type-species unknown to me. 
Hambergiella Roman (syn. teste Perkins, 1962, p. 408). 

?petiolaris (Cameron), n. comb. 

Subfamily TRYPHONINAE 
Tribe IDIOGRAMMATINI 

Genus IDIOGRAMMA Foerster, p. 218, Sup. 1, p. 40 

*titana Burdick. Calif. (Santa Clara Co.). Host: '!Xyela n. sip. 

Idiogramma titana Burdick, 1958. Pan-Pacific Ent. 34 (3) : 159. $,9. 



ICHNEUMONIDAE 87 

Tribe PHYTODIETINI 

Genus PHYTODIETUS Gravenhorst, p. 211 

*Euctenopu8 Ashmead, 1900 (Oct.). U. S. Natl. Mus. Proc. 23: 50 (sjm. 
teste Townes, Townes and Gupta, 1961, p. 90). 

Type: Euctenopus zealandicus Ashmead. Orig. desig. 
Euctenopus Ashmead, 1900 (Nov.). Place type-species name in parenth- 
eses and add : = zealandicus Ashmead. 
*Weisia Schmiedeknecht, 1907. Opusc. Ent. 3 (16) : 1257 (syn. teste 
Townes, Townes and Gupta, 1961, p. 91). 

Type : Weisia elegans Schmiedeknecht. By monotypy. 
*Neuchorus Uchida, 1931. Ins. Matsumurana 5 (3) : 143 (syn. teste 
Townes, 1957, Ent. Soc. Wash. Proc. 59 (3) : 113). 

Type: Neuchorus longicauda Uchida. By monotypy and orig. 
desig. 

This genus needs revision. 

vulgaris Cresson. Host: Dioryctria reniculella (Grt.). 

NETELIA subgenus PARABATES Foerster, p. 212 

Parahates Foerster. Considered valid genus by Perkins, 1962, p. 443. 
Parabatus Thomson. Emend. 

NETELIA subgenus *BESSOBATES Townes, Townes and Gupta 
(=:Nefefia subgenus Parabafus auctt, p. 216) 

Parabatus Thomson. Delete type citation and transf. to syn. of Parabates 

Foerster, p. 212, as emend. 
*Bessobates Townes, Townes and Gupta, 1961. Cat. and Classif. Indo- 
Austral. Ichn., pp. 93, 430. 

Type: Parabatus deceptor Morley. Desig. by Townes, Townes, 
and Gupta, 1961. 
Species listed under Parabatus Thomson, p. 216 all belong here. 

NETELIA subgenus NETELIA Gray, pp. 213-216 

*Scammatonotum Enderlein, 1914. Beitr. Kennt. Land-Sussw. 
Deut.-Sudwestafricas. tiym. IV, Ichn. p. 231 (syn. teste Townes, 
Townes and Gupta, 1961, p. 95), 

Type: Scammatonotum herero Enderlein. By monotypy and 
orig. desig. 
*Amebachia Uchida, 1928. Fac. Agr. Hokkaido Imp. Univ. Jour. 21: 186, 
198 (syn. teste Townes, Townes, and Gupta, p. 95). 
Type: Amebachia baibarana Uchida. Orig. desig. 

chloris (Olivier). 

Netelia (ISetelia) geminatus (Say). (Syn. teste Townes, 1961. Ent. Soc. 
Wash. Proc. 63(2) : 111.) 

geminatus (Say). A syn. of chloris (Olivier). 

sayi (Cush.). Host: Pseudaletia unipuncta (Haw.). 
Biology: Guppy, 1961. Canad. Ent. 93 (7) : 569. 

Tribe THYMARIDINI 

Genus THYMARIS Foerster, p. 217 

Thymaris Foerster. In type citation, delete "= [Hemiteles^ contamin- 
atus Gravenhorst" and substitute: = [Mesoleptus^ tener (Gra- 
venhorst) . 

Thymarus Thomson. Delete type citation. Add : Emend. 



88 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Tribe TRYPHONINI 

Genus CTENOCHIRA Foerster, pp. 221-222 

Ctenacmus Thomson. Add "Emend." after bibliogr. ref. Delete type ci- 
tation. 

Trichocalymmus Thomson. Delete type citation and transf. to syn. of 
Erromenus subg. Erromenus Holmgren, p. 222, as emendation of 
Trichocalymma Foerster. 

*Pauroctenus Cameron, 1909. Bombay Nat. Hist. Soc. Jour. 19: 726 
(syn. teste Townes, Townes, and Gupta, 1961, p. 105). 
Type: Pauroctonus paUipes Cameron. By monotypy. 

ERROMENUS subgenus ERROMENUS Holmgren, pp. 222-224 

Trichocalymmus Thomson. Emend. Transf. from syn. of Ctenochira Fo- 
erster, p. 221 (teste Perkins, 1962, p. 390). 

Genus MONOBLASTUS Hartig, pp. 224-225 

Otoblastus Foerster. Perkins (1962, p. 441) considers this a valid genus. 
Coeloconus Foerster. Change date in type citation to 1911. Perkins 

(1962, p. 414) considers this a valid genus. 
TSeleges Foerster. Perkins (1962, p. 438) considers this a valid genus. 

Genus DYSPETES Foerster 
( = D/spefu$ Thomson, p. 225) 

Dyspeies Foerster, 1868. Transf. from syn. of Horogenes Foerster, p. 
377. Replace type citation with: 

Type: Dyspetus praerogator (Gravenhorst) = arrogator 
Heinrich, 1949. Incl. by Thomson, 1883. 
Dyspetus Thomson, 1883. Emend. 

TRYPHON subgenus TRYPHON Fallen, pp. 225-226 
Psilosage Foerster. Perkins (1962, p. 448) considers this a valid genus. 

Tribe EXENTERIN! 

{zzzCfenisdni of Catalog) 

Taxonomy: Mason, 1962. Canad. Ent. 94 (12): 1273-1296 (generic li- 
mits). 

Genus ACROTOMUS Holmgren, p. 228 

Taxonomy: Mason, 1962, p. 1285. 

*succinctus (Gravenhorst). Ont. 

Tryphon succinctus Gravenhorst, 1829. Ichn. Europaea, v. 2, p. 166. $, 

9. 
Distribution : W. R. M. Mason, in litt. 

Genus EXENTERUS Hartig, pp. 228-230 

Biology: Finlayson, 1960. Canad. Ent. 92 (1, 12) : 26-30, 925-929 (larval 
descr. several spp.). — Finlayson, 1963. Canad. Ent. 95 (5): 
475-507 (hosts, larval keys, bibliography). 

amictorius (Panzer). N. H., Ohio, Wis. Hosts: Diprion hercyniae (Htg.), P. 
shjiilis (Htg.), Neodiprion nanulus nanulus Schedl, N. sertifer 
(Geoff.). 

canadensis Provancher. Hosts: Neodiprion abbotii (Leach), N. abietis com- 
plex, N. excitans (Roh.), N. nanulus nanulus Schedl, N. pratti 
pratti (Dyar). 
Biology: Finlayson, 1960. Canad. Ent. 92 (1): 26; 1963. Canad. Ent. 95 
(5) : 486, 488. 



ICHNEUMONIDAE 89 

claripennis Thomson. Remove from catalog; misidentification (see Exenterus 
confusus Kerrich) ; not in Nearctic region. 

*confusus Kerrich. N. B., Que.; Europe. Hosts: Diprion hercyniae (Htg.), 
Ncodiprion nanubis 7ianuJns Schedl. 
Exenterus confusus Kerrich, 1952. Brit. Mus. (Nat. Hist.) Ent. Bui. 2 
(6): 361, 366. 5, 2. Misdet. as E. claripennis Thomson in cata- 
log). 

tsugae Cushman. Host: Neodiprion sp. nr. scutellatus Roh. 

walleyi Cushman. Host: Neodiprion sertifer {Geoff .) . 

Genus SMICROPLECTRUS Thomson, p. 230, Sup. 1 , p. 43 
Excavarus Davis. Remove from syn. A valid genus. 
annulipes (Cresson). Transf. to Excavarus Davis. 
etrocaulus etrocaulus Mason. Transf. to Excavarus Davis. 
etrocaulus eurits Mason. Transf. to Excavarus Davis. 
velox Walley. Transf. to Excavarus Davis. 

Genus EXCAVARUS Davis 

A valid genus (teste Mason, 1962, p. 1293) ; not a syn. of Smicroplectrus 
Thomson. Place after Smicroplectrus, p. 230. 

annulipes (Cresson). Transf. from Smicroplectrus Thomson, p. 230 (teste 
Mason, 1962, p. 1293). 

etrocaulus etrocaulus (Mason). Transf. from Smicroplectrus Thomson, Sup. 
1, p. 43 (teste Mason, 1962, 1293). 

etrocaulus eurus (Mason). Transf. from Smicroplectrus Thomson, Sup. 1, p. 
43 (teste Mason, 1962, p. 1293). 

velox (Walley). Transf. from Smicroplectrus Thomson, p. 230 and Sup. 1, p. 
43 (teste Mason, 1962, p. 1293). 

Genus EXYSTON Schiodte, pp. 230-231 

Anecphysis Foerster. Remove from syn. A valid subg. 
Revision: Mason, 1959. Canad. Jour."Zool. 37: 1067-1115. 

cinctulus (Gravenhorst). Delete from Catalog (misdet.), not in Nearctic 
fauna; is Palearctic (teste Mason, 1959, p. 1110). 
Cteiiiscus clavatus Cresson. Remove from syn. A valid sp. in subg. Ex- 

yston Schiodte. 
Cteniscus abdominalis Cresson. Syn. of clavatus (Cresson). 
Mesoleptus macitlosus Provancher. Syn. of clavatus (Cresson). 
ExYston abdominalis Ashmead. Syn. of clavatus (Cresson). 
Rhimphalea erythrogaster Viereck. Syn. of clavatus (Cresson). 

contractus Davis. Not assignable to subg. or species (teste Mason, 1959, p. 
1112). 

excelsus (Cresson). Placed in subg. Exyston Schiodte. 

Exyston marginatum Provancher. A valid sp. in subg. Anecphysis Fo- 
erster. 

flavens Davis. Placed in subg. Exyston Schiodte. 

humeralis Davis. Placed in subg. Exyston Schiodte. 

nigreo Davis. Placed in subg. Anecphysis Foerster with its syn. as syn. of 
marginatus Provancher. 

nigroscutum Davis. Not assignable to subg. or species (teste Mason, 1959, p. 
1112). 

speciosus Davis. Placed in subg. Anecphysis Foerster. 

variatus Provancher. Placed in subg. Anecphysis Foerster. Misdet. in Cat. 
(teste Mason, 1959, p. 1082). 



90 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Exyston nigroscutum Davis. Remove from syn. Not assignable to subg. 

{teste Mason, 1959, p. 1112). 
Exyston horeotis Davis. Remove from syn. A valid sp. in subg. Anec- 
physis Foerster {teste Mason, 1959, p. 1082). 

venustus (Cresson). Placed in subg. Exyston Schiodte. 

Exyston politus Davis. Remove from syn. A valid species in subg. Exys- 
ton Schiodte. 

EXYSTON subgenus ANECPHYSIS Foerster 

Place on p. 230 before Exyston (Exyston) Schiodte (teste Mason, 
1959, p. 1067). Subgeneric placement of all species teste Mason, 
1959, pp. 1071-1097. 

*ater Mason. B. C, Wash., Oreg. 

Exyston (Anecphysis) ater Mason, 1959. Canad. Jour. Zool. 37: 1070, 
1095. $, ?. 

*austel!i Mason. Mass. to N.C. 

Exyston (Anecphysis) austelli Mason, 1959. Canad. Jour. Zool. 37: 1069, 
1078. $, ?. 

boreotis Davis. Mich, to Cont. Divide in Transit, and Canad. Zones. 

♦californicus Mason. Alaska, Oreg., Calif., Colo., Wyo., Mont., Idaho, Alta. 

Exyston (Anecphysis) calif ornicus Mason, 1959. Canad. Jour. Zool. 37: 
1070, 1093. 9, 5. 

*hadros Mason. Alta., Sask., Colo., Minn., Mich. 

Exyston (Anecphysis) hadros Mason, 1959. Canad. Jour. Zool. 37: 1069, 
1080. $, 9. 

*Iophotos Mason. N. W. T., Sask., Alta., B. C, Wash., Calif. 

Exyston (Anecphysis) lophotos Mason, 1959. Canad. Jour. Zool. 37: 

1070, 1091. $, ?. 

marginatus Provancher. Transcont. in Transit., Canad., and Huds. Zones. A 
valid species {teste Mason, 19.59, pp. 1071-1073). 
Anecphysis curvineura Davis (syn. teste Mason, 1959, p. 1071). 
Exyston nigreo Davis (syn. teste Mason, 1959, p. 1071). 

*reniformis Mason. Colo., Alta.. B. C, Oreg., Calif. 

Exyston (Anecphysis) reniformis Mason, 1959. Canad. Jour. Zool. 37: 

1070, 1089. $, 9. 

speciosus Davis. Alta., Idaho, Mont., Wyo. 

*spinuIosus Mason. Alaska, Yukon, N. W. T., Wash., Sask., Wyo., N. Dak., 
S. Dak., Minn., Iowa, Kans. (Misdet. as contractus in Catalog, 
teste Mason, 1959, p. 1086). 
Exyston (Anecphysis) spinulosus Mason, 1959. Canad. Jour. Zool. 37: 
1069, 1086. $, 9. 

*tectus Mason. N. B., P.E.I, to N. C. and Tenn., west to Colo., B. C, Wash., 
Oreg., also Calif., Ariz. Hosts: Pristiphora hivittata (Nort.), P. 
idiota (Nort.), P. sycophanta Walsh. 
Exyston (Anecphysis) tectus Mason, 1959. Canad. Jour. Zool. 37: 1069, 
1073. $, 9. 

variatus Provancher. Atlantic to Cont. Divide; B. C. Canad. Zone to U. 
Austr. Zone. 

EXYSTON subgenus EXYSTON Schiodte, pp. 230-231 

Subgeneric placement of all species, teste Mason, 1959, pp. 1097-1110. 

*chamaeleon Mason. Yukon, B. C, Wash., Oreg., Calif., Ariz., N. Mex., 
Utah, Colo., Alta., Sask., S. Dak., Ont. 
Exyston (Exyston) chamaeleon Mason, 1959. Canad. Jour. Zool. 37: 
1069, 1099. 9, $. 



ICHNEUMONIDAE 91 

clavatus (Cresson). Atlantic to Cont. Divide in Canad., Transit., U. Austr. 

Zones, B. C, Wash., Calif. Valid sp. teste Mason, 1959, pp. 1068, 

1110). 
Cteniscus abfloniinalis Cresson. 
Mesoleptus luacufosus Provancher. 
Exyston abdottiinalis Ashmead. 
Rhiinphalea erythrogaster Viereck. 

excelsus (Cresson). Colo. 

flavens Davis. N. J. 

humeralis Davis. Kans. 

*illinois Mason. Que., Ont., Mich., 111., Iowa. 

Exyston (Exyston) illinois Mason, 1959. Canad. Jour. Zool. 37: 1069, 

1103. ?, $. 

politus Davis. Calif., Wyo., Mont. A valid sp. {teste Mason, 1959, pp. 1069, 
1109). 

venustus (Cresson). B. C, Wash., Utah, Colo., S. Dak., Sask. 

Subfamily CRYPTINAE 
Tribe BRACHYCYRTINI 

Genus BRACHYCYRTUS Krtechbaumer, p. 203, Sup. 1, pp. 39, 43 

*Xanthocharops Morley, 1912. Linn. Soc. London, Trans. 15: 173 (syn. 
teste Townes and Townes, 1960, p. 542). 

Type: Xanthocharops primus Morley. By monotypy. 

pretiosus Cushman. Host: Chrysopa lateralis (Guer.). 

Tribe HEMITELINI 

Genus BATHYTHRIX Foerster, pp. 231-232 

Chaeretymma Foerster, 1868. Delete type desig. and transf. genus to 

syn. of Cubocephalus Ratzeburg, p. 253. 
*Stenoschema Foerster, 1868. Naturh. Ver. Rheinlande. Verh. 25: 220. 

Nom. Nud. (Syn. teste Townes, Townes, and Gupta, 1961, p. 119.) 
*Agenora Cameron, 1909. Bombay Nat. Hist. Soc. Jour. 19: 722 (syn. 
teste Townes, Townes, and Gupta, 1961, p. 119). 

Type: {Agenora hirticeps Cameron) = claviger Taschenberg. 
By monotypy. 

Genus MASTRUS Foerster, pp. 233-234 

*Perosis Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 169 (syn. 
teste Perkins, 1962, p. 443). 

Type: Echthrus armatus Gravenhorst. Desig. by Viereck, 
1914. 
*Daictes Foerster, 1868. Naturh. Ver, Rheinlande Verh. 25: 176 (syn. 
teste Townes, 1957. Ent. Soc. Wash. Proc. 59 (3) : 104). 

Type : Phygadeuon (Daictes) fukaii Viereck. First inch sp. 

argeae (Viereck). 

Biology: Finlayson, 1960. Canad. Ent, 92 (1) : 30 (larval). 

*Genus DICHROGASTER Doumerc 
( = Ofacus/es Foersfer, pp. 234-235) 

Dichrogaster Doumerc, 1855. Soc. Ent. France Ann. (Ser. 3) 3, Bui. 3: 

LXXXVIII-LXXXIX. 

Type: Microgaster perlae Doumerc. By monotypy. 
Otacuses Foerster (syn. teste Townes, Townes and Gupta, 1961, p. 121). 
*Microtorus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 174 

(syn. teste Townes, Townes and Gupta, 1961, p. 121). 



92 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Type: Microtorus kichijoi Uchida ( = Heniiteles (Rhadiurgus) 

chrysopae Uchida, preocc). Desig. by Townes, 1957. 

*Xenobrachys Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 179 

(syn. teste Perkins, 1962. Brit. Mus. (nat. Hist.) Ent. Bui. 11 

(8): 462;— Townes, Townes and Gupta, 1961, p. 121 [without 

type-species]. 

Type: Hemiteles longicaudatus Thomson. Desig. by Perkins, 
1962. 

bicolor (Cushman), n. comb. 

crassus chrysopae (Ashmead), n. comb. Hosts: Chrysopa sp., C. bimaculata 
McCl., C. cubana Hager, C. rufilabris Burm., C. sanchezi Navas. 

crassus patruelis (Cushman), n. comb. 

Genus ETHELURGUS Foerster, p. 235 

*lSuneches Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25 : 186. 

Type: Phygmleuon godalis Taschenberg. Incl. and desig. by 
Perkins, 1962. 

belfragi (Ashmead). Not congeneric with type-species syrphicola Ashm. Re- 
visionary work needed for placement. 

Genus ISDROMAS Foerster, p. 236 

Isdromas Foerster. In type citation change "Ashmead, 1900" to Ash- 
mead in Smith, 1900. 

lycaenae (Howard). Host: Apanteles congregatus (Say); delete Homadaula 
albizziae Clarke. 

Genus ALEGINA Foerster, pp. 236-237 

apantelis (Cushman). Host: Ocnerostoma strobivorum Free. 

*Iaricenae Mason. N. B., P.E.I., Ont. Host: Argyresthia laricella Kft. 
Alegina laricellae Mason, 1961. Canad. Ent. 93 (1): 4. $, $. 

niaesticolor (Roman). Transf. from Haplaspis Townes, p. 238 {teste Townes, 
1961. Ent. Soc. Wash. Proc. 63 (2) : 104). 

Genus CHIROTICA Foerster 

Chirotica Foerster. Transf. from syn. of Gelis Thunberg, p. 239. 

Type: (Hemiteles insignis Gravenhorst) = trifasciator (Thun- 
berg) . Included by Schimiedeknecht, 1897. 

*Spinolia Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 173. 
Preocc. (Syn. iesfe Perkins, 1962, p. 452.) 

Type: Hemiteles maculipennis Gravenhorst. Incl. by Schmie- 
deknecht, 1897. 
Diaglypta Foerster. Transf. from syn. of Phobetes Foerster, p. 237 (syn. 

teste Perkins 1962 p. 417). 
*Deuterospinolia Dalla Torre, 1902. Cat. Hym., v. 3 (pt. 2), p. 678. N. 
name. 

Place after Diaglyptidea Viereck, p. 237. 

conspicua (Cresson), n. comb. Transf. from Phobetes Foerster, p. 238. 

sheppardi (Walley), n. comb. Transf. from Phobetes Foerster, p. 238. 

Genus Phobetes Foerster, pp. 237-238 

Allocota Foerster. Transf. to syn. of Hymenosyneches Viereck. 
Syneches Foerster. Transf. to syn. of Hymenosyneches Viereck. 
Diaglypta Foerster. Transf. to syn. of *Chirotica Foerster. 
Phobetes Foerster, 1868. Delete type citation and transf. to Euryproctini, 

p. 338, as valid name for Ipoctoninus Hincks. 
Hymenosyneches Viereck. Remove from syn. A valid genus. 



ICHNEUMONIDAE 93 

con f ederatae (Ashmea.d) . Transf . to Hymenosyneches Viereck. 
conspicuua (Cresson). Transf. to *Chirotica Foerster. 
sheppardi (Walley). Transf. to *Chiroiica Foerster. 
thyridopteryx (Riley). Transf . to Hymenosyneches Viereck. 

Genus HYMENOSYNECHES Viereck 

Allocota Foerster, preocc. Transf. from syn. of Phobeles Foerster, p. 

237. N. syn. 

Syneches Foerster, preocc. Transf. from syn. of Phobetes Foerster, p. 

237. 
Hymenosyneches Viereck. Transf. from syn. of Phobetes Foerster, p. 

238. A valid genus. 

conf ederatae (Ashmead), n. comb. Transf. from Phobetes Foerster, p. 238. 
thyridopteryx (Riley). Transf. from Phobetes Foerster, p. 238. 

*Genus LYSIBIA Foerster 
i = Haplaspis Townes, p. 238) 

Lysibia Foerster 1868. Naturh. Ver. Rheinlande Verb. 25: 175. 

Type: (Tryphon fulvipes Gravenhorst) =nana Gravenhorst. 
Design, by Townes, Momoi, and Townes, 1965. An earlier 
desig. by Perkins, 1962, p. 436, of Tryphon nanus Graven- 
horst is invalid because that species was not one of the 
three (one through generic synonymy) included by 
Townes, Townes, and Gupta, 1961, p. 123, who made no 
type designation at that time. 
*Penion Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 174 (syn. 
teste Perkins, 1962, p. 443; also Townes, Townes, and Gupta 1961, 
but without type desig.) 

Type: Pemon proximum Perkins. Included and desig. by Per- 
kins, 1962. 
Haplaspis Townes (syn. teste Townes, Townes, and Gupta, 1961, p. 123). 

maesticolor (Roman). Transf. to Alegina Foerster, p. 236. 

mandibularis (Provancher) , n. comb. 

*Genus CHARITOPES Foerster 

Charitopes Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 181. 

Type: Hemiteles (Charitopes) chrysopae Brischke. First incl. 
sp. 

Place on p. 238 before Gelis Thunberg. 

gastricus (Holmgren). Transf. from Alphabetic List of Unplaced Species, p. 

408 (teste Townes, 1961. Ent. Soc. Wash. Proc. 63(2) : 106). 

plecticinus (Roman). Transf. from Alphabetic List of Unplaced Species, p. 

409 (teste Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 111). 

Genus GELIS Thunberg, pp. 238-243 

*Thaumatotypus Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 172 
(syn. teste Perkins, 1962, p. 457). 

Type: Thaumatotypus femoralis Brischke. Incl. by Brischke, 
1881. 
Chirotica Foerster. Delete type desig. and remove from syn. A valid 

genus. 
*Rhadiurgus Foerster, 1868. Naturh. Ver. Rheinlande 25: 177 (syn. 
teste Perkins, 1962, p. 449). 

Type: (Hemiteles bicolorinus Gravenhorst) =z *Gef is cinctus 
(Linnaeus). Incl. by Uchida, 1933. 
Ilapinastes Foerster. In type citation add "in Davidson" between Ash- 
mead and comma. 



94 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Urithreptis Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 180 (syn. 
teste Perkins, 1962, p. 462; Townes, Momoi, and Townes, 1965, p. 
133). 

Type: Not yet selected (Foerster spm., a Gelis $, not now 
recognized to species). 
*Thaumatogelis Kupka in Schmiedeknecht, 1933. Opusc. Ichn. Sup., fasc. 
17: 7-8 (syn. teste Perkins, 1962, pp. 392, 457). 

Type: Pezomachus aqnisgranensis Foerster. Desig. by Townes, 
Momoi, and Townes, 1965. 
*Cryptogelis Hellen, 1944. Notulae Ent. 24: 2. 

Type: Pezomachus aqnisgranensis Foerster. Desig. by 
Townes, Momoi, and Townes, 1965. 
*HolcogeUs Aubert, 1957. Miinchner Ent. Ges. Mitt. 57: 226. 
Type: Pezomachus corruptor Foerster. Orig. Desig. 

A revision will probably show this genus to be comprised of several 
segregates. 

obscurus (Cresson). Ont. Host: Ocnerostoma STp. 

tenellus (Say). Host: Coleophora laricella Hbn. 

urbanus (Brues). Que. 

Biology: Finlayson, 1960. Canad. Ent. 92(12) : 930 (larval). 

Genus GNYPETOMORPHA Foerster, p. 245 

*Daetora Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 175 (syn. 
teste Perkins, 1962, p. 416). 

Type: Hemiteles sotutus Thomson. Desig. by Perkins, 1962. 
*Opisthostenus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 175 
(syn. teste Townes, 1957. Proc. Ent. Soc. Wash. 59 (3) : 114). 

Type: Hemiteles (Opisthostenus) etorofuensis Uchida. Incl. 
by Uchida, 1936. 

*Genus XENOLYTUS Foerster 
( = Sfernocryptus Roman, p. 245) 

Xenolytus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 174. 

Type: * Xenolytus rufipes Cameron) = Xenolytus bitinctus 

(Gmelin). Included by Cameron, 1906 (Feb.). 
Sternocryptus Roman (syn. teste Perkins, 1962, p. 462). 

bitinctus (Gmelin). 

*Xenolytus rufipes Cameron, 1906 (Feb.). So. African Mus. Ann. 5: 154. 
9. 

liinatus (Cresson), n. comb. 

occidentalis (Harrington), n. comb. 

subspinosus (Provancher) , n. comb. 

ENDASYS subgenus ENDASYS Foerster, pp. 245-246 

*Scinacopus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 185 
(syn. teste Habermehl, 1912. Deut. Ent. Ztschr. 1912: 175; Per- 
kins, 1962, p. 451). 

TypeiCPhygadeuon cnemargus Gravenhorst = [Endasys] 
parviventris cnemargus (Gravenhorst). Included by 
Brischke, 1891. 

subclavatus (Say). Hosts: Neodiprion excitans Roh., N. lecontei (Fitch), N. 
namdus nanuhis Schedl. N. pratti pratti (Dyar), N. sertifer 
(GeoflF.), N. sivainei Midd., N. virginiana Roh., Zadiprion grandis 
(Roh.). Parasite: Euceros frigidvs Cress. 
Biology: Finlayson, 1960. Canad. Ent. 92 (1): 25, 32.— Tripp, 1961. 
Canad. Ent. 93 (1): 53.— Finlayson, 1963. Canad. Ent. 95 (5): 
476-485, 491.— Bobb, 1963. Jour. Econ. Ent. 56 (5) : 619, 620. 



ICHNEUMONIDAE 95 

*Genu$ THEROSCOPUS Foerster 

Theroscopus Foerster, 1850, Arch. f. Naturgesch, 16 (1) : 92. 

Type: (Pezomachus pedestris Gravenhorst) = Theroscopus 
pedestris (Fabricius). Desig. by Viereck, 1914. 

Place in Hemitelini in Catalog, p. 247, before Eriplanus Foerster. An 
undescribed species occurs in the midwest. 

Genus ERIPLANUS Foerster, p. 247 

Chamerpes Foerster. Perkins (1962, pp. 392, 413) considers this a syn. 

of Theroscopus Foerster. 
Phyrtus Foerster. Perkins (1962, pp. 392, 445) considers this a syn. of 

Theroscopus Foerster. 
*Thysiotorus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 181. 
Perkins (1962, pp. 392, 458) considers this a syn. of Theroscopus 
Foerster. 

Type: (Hemiteles (Physiotorus) [sic] brevipennis Brischke) 
= hemipterus (Fabricius) (teste Perkins, 1962, p. 458). 
Desig. by Viereck, 1914. 

I am not at all sure that the above genera, listed by Perkins as syno- 
nyms of Theroscopus Foerster, belong here. They are left here until a much 
needed revision of the Hemitelini places them elsewhere. 

Genus PHYGADEUON Gravenhorst, pp. 247-250, Sup. 1, p. 45 

*Stibeute8 Foerster, 1850. Arch. f. Naturgesch. 16 (1) : 76 (syn. teste 
Thomson, 1884. Opusc. Ent. f. 10, p. 948; Perkins, 1962, p. 453). 
Type: Stibeutes gravenhorstii Foerster. Desig. by Ashmead, 
1900. 
* Apterophygas Foerster, 1868. Naturh. Rheinlande Verh. 25: 172 
(syn. teste Perkins, 1962, p. 406). 

Type: Apterophrgus (Fst.) ? paradoxus Bridgman. Incl. by 
Schmiedeknecht, 1897. 
*Gunopaches Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 174 
(syn. teste Perkins, 1962, p. 426). 

Type: Gunopaches crassus Perkins. Incl. and desig. by Per- 
kins, 1962. 
*PantoUspa Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 178 
(syn. teste Perkins, 1962, p. 442). 

Type: Gunopaches crassus Perkins. Incl. and desig. by Per- 
kins, 1962. 
*Bathymetis Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 182 
(syn. teste Perkins, 1962, p. 410). 

Type: Phygadeuon (Bathymetis) cylindricus Brischke. Desig. 
by Viereck, 1914. 
Homelys Foerster. Remove "Included by Roman, 1909" from type cita- 
tion and replace with: Incl. and desig. by Viereck, 1914. (See 
Perkins, 1962, p. 428.) 
*Ernoctona Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 183 
(syn. teste Perkins, 1962, p. 423). 

Type: Phygadeuon rugulosus Gravenhorst. Incl. and desig. by 
Perkins, 1962. 
Plesignathus Foerster. Delete type citation and transf. genus to syn. of 

Polytribax Foerster, p. 257. 
Zaphleges Foerster. Misspelled in Index, p. 1284, as Zathleges. 
*? Odontoneura Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 185. 
Syn. (see Perkins, 1962, p. 440). 

Type: Phygadeuon annuUcornis Thomson. Desig. by Perkins, 
1962. 

victoriensis Townes. Calif. Host: Fannia canicularis (h.) . 

Centeterus canadensis Harrington. Add 1894 to bibliographic citation 
and change vol. no. to 26. 



96 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus ISELIX Foersier, p. 250 
Perkins, 1962, p. 432, considers this a syn. of Phygadeuon Gravenherst. 

*Genus CREMNODES Foerster 
( = Caenomer/s Foerster, p. 250) 

Cremnodes Foerster, 1850. Arch. f. Naturgesch. 16 (1) : 72. 

Type: Ichneumon atricapillus Gravenhorst. Orig. desig. 
Caenomeris Foerster (syn. teste Perkins, 1962, p. 411). 
*Stygera Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 176 (syn. 
teste Perkins, 1962, p. 397, 454). 

Type: Stygera rufipes Perkins. Incl. and desig. by Perkins, 
1962. 
*Cremnias Roman, 1939. Ent. Tidskr. 60 (3-4) : 186. N. name, unneces- 
sarily proposed. 

Genus STILPNUS Gravenhorst, pp. 250-251 
anthomyidiperda Viereck. Calif. Host: Fannia canicularis (h.) 

Genus MESOLEPTUS Gravenhorst, p. 251 

*Exolytus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 182. 
Preocc. 

Type: Ichneumon laevigatus Gravenhorst. Incl. and desig. by 
Foerster, 1871. 

Genus ATRACTODES Gravenhorst, pp. 251-253, Sup. 1, p. 45 
Asyncrita Foerster. Perkins 1962, p. 408, considers this a valid genus. 

Tribe HEMIGASTERINI 

[ziAp\es\n\ of Catalog) 

Revision: Townes and Gupta, 1962. Amer. Ent. Inst. Mem. 2, 305 pp., 
figs. 

♦Genus JAVRA Cameron 

*Cnemocryptus Cameron, 1903. Mem. and Proc. Manchester Lit. Phil. 
Soc. 47 (14) : 38. 

Type: {Cnemocryptus validicornis Cameron) = parviceps 
Cameron. By monotypy. 
*Javra Cameron, 1903. Mem. and Proc. Manchester Lit. Phil. Soc, 47 
(14) : 47. 

Type: Javra parviceps Cameron. Desig. by Viereck, 1914. 
*Monocryptus Hellen, 1956. Notulae Ent. 36: 135. New syn. 
Type : Cratocryptus opacus Thomson. By monotypy. 

Place as first genus in Hemigasterini. 

*glaesaria Townes. R. I., N. Y., Ont. 

Javra glaesaria Townes, 1962. Amer. Ent. Inst. Mem. 2: 12, 13. 9. 

*rugata Townes. Ohio, Va. 

Javra rugata Townes, 1962. Amer. Ent. Inst. Mem. 2: 12. 9. 

♦Genus PARMORTHA Townes 

Parmortha Townes 1962. In Townes and Gupta, Amer. Ent. Inst. Mem. 
2: 8, 14. 

Type: (Parmortha pleuralis atripes Townes) = pleuralis albo- 
maculata (Ashm.) n. comb., n. syn. (teste Momoi who 
examined types). 

Place after Javra Cameron in tribe Hemigasterini. 

circumcincta circumcincta (Provancher) . Canad. and Upper Austral Zones. 
Transf. from Cubocephalus Ratzeburg (teste Townes, in Townes 
and Gupta, 1962, p. 24). 



ICHNEUMONIDAE 97 



circumcincta flavipes (Harrington). Colo., Alta., Yukon. Transf. from Cu- 
bocephalus Ratzeburg, p. 254 (teste Townes, in Townes and 
Gupta, 1962, p. 23). 

*parvula (Gravenhorst) . Alaska, B. C. to N. S. south to Oreg. and N. J.; 
Europe. Hosts: Ametastegia glabrata (Fallen) ; sawfly in Ribes. 
Cryptus parvulus Gravenhorst, 1829. Ichn. Europaea 2: 459 "5", ? ("5" 

= Apsilops cincticornis F.). 
*Cryptus erythropus Gravenhorst, 1829. Ichn. Europaea 2: 469. $. 
Cuhocephalus erythropus velox (Cresson). (syn. by Townes, in Townes 
and Gupta, 1962, p. 16). 

*pleuralis alpina Townes. N. H. (Mt. Washington). P. pleuralis pleuralis 
Thomson occurs in Europe. 
Parmortha pleuralis alpina Townes, 1962. in Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 19, 22. 9. 

*pleuralis arctica Townes. Alaska, N. W. T., Que. 

Parmortha pleuralis arctica Townes, 1962, In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 19. $, 9. 

pleuralis signata (Provancher) , changed status. Alaska, B. C., Wash., Idaho, 
N. C. Host: Ametastegia glabrata (Fallen). Transf. from Cu- 
hocephalus Ratzeburg, p. 255 and reduced to subsp. rank {teste 
Townes, in Townes and Gupta, 1962. p. 20). 

Genus CRATOCRYPTUS Thomson 

Cratocryptus Thomson. Removed from synonymy of Cuhocephalus Ratze- 
burg, p. 253. Valid genus {teste Townes in Townes and Gupta, 
1962. Amer. Ent. Inst. Mem. 2: 9, 25). 

Place after *Parmortha Townes. 

*petilus Townes. Transcont. in Canad. Zone. Host: Cephaleia sp. 

Cratocryptus petilus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 26. $,9. 

Genus CUBOCEPHALUS Ratzeburg, pp. 253-255 

Chaeretymma Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 187. 
Transf. from syn. of Bathythrix Foerster, p. 231. Change type 
desig. to: 

Type: Cryptus anatorius Gravenhorst. Included (first sp.) 
and desig. by Schmiedeknecht, 1890 (p. 117). 
Cratocryptus Thomson, 1873. Remove from syn. Valid genus. 
*Planocryptus Heinrich, 1949. Munchner Ent. Gesell. Mitt. 35-39: 56. 

Type: *Planocryptus ntirabilis Heinrich. By monotypy and 
orig. desig. (Syn. teste Townes and Gupta, 1962, p. 92). 
Taxonomy: Perkins, 1962. Brit. Mus. (Nat. Hist.) Ent. Bui. 11 (8) : 413 
(Chaeretymma) . 

alacris alacris (Cresson), changed status. Ont., Mass., N. Y., Pa., N. J., S. 
C, Ind., Mich., Iowa, Kans. Reduced to subsp. rank (teste Townes 
and Gupta, 1962, p. 93). 

alacris canadensis (Provancher), changed status. Transcont. in Canad. Zone. 
Host: Hemitaxomis dubitatus (Nort.). Reduced to subsp. rank 
(teste Townes and Gupta, 1962, p. 89). 

alacris erythropygus (Provancher), changed status. Atlantic to Wis., in Can- 
ad. and Transit. Zones. Reduced to subsp. rank (teste Townes 
and Gupta, 1962, p. 92). 

alacris minutus (Cushman), changed status. B. C. to Calif. Hosts: Ametas- 
tegia glabrata (Fallen). Removed from syn. of canadensis Pro- 
vancher and placed as subsp. of alacris (teste Townes and Gupta, 
1962, p. 91). 

annectus Townes. Alaska, Yukon, B. C, Alta., Man., Que., N. S., south to 
Calif., N. Max., Mich., N. H. 



98 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Phaeogenes nigricornis Provancher (syn. teste Townes and Gupta, 1962, p. 

117). 
Cubocephalus atricornis Townes (syn. teste Townes and Gupta, 1962, p. 

117). 

annulatus (Cresson). Alta., Conn., Maine, Minn., N. C, Ont., Vt. Host: 
lAgrilus rubicola Abeille. 
Cryptus latus Provancher, Transf. to Ischnus Gravenhorst, p. 270, as 

valid species. 
Mesosienus nitidus Provancher. Remove from syn. ; a valid species. 
Cryptus riificoxus Provancher. Retain as syn. {teste Townes and Gupta, 

1962, p. 82). 
Mesosienus ruficoxus Provancher. Questionable syn. of Giraudia plana 

Provancher, p. 257. 
Cryptus occidentalis Provancher. Place as syn. of Cubocephalus nitidus 

Provancher. 
Phygadeuon inhabilis Provancher. Remove from syn.; a valid species. 
Phygadeuon nitidulus Provancher. Syn. of nitidus (Provancher). 
Mesostenus provancheri Dalla Torre. Questionable syn. of Giraudia 
plana Provancher, p. 257. 

^aquilonius Townes and Gupta. Alaska, Yukon, N. W. T., Labrador, B, C., 
Man., Colo. 
Cubocephalus aquilonius Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2: 30,37, 53. 3, ?• 

*ardens Townes. Alaska, Colo., Wis., Mich., Ont., N. H. 

Cubocephalus ardens Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 34, 41, 109. $, 9. 

ashmeadii (Harrington). Rocky Mts. to Pacific in Canad. and Transit. 
Zones. Removed from syn. of canadensis Provancher, p. 254 {teste 
Townes and Gupta, 1962, p. 96). 

atricornis Townes. Syn. of annectus Townes. 

*atriclunis Townes and Gupta. B. C, Man., Ont., Que., Maine, Vt., Mass., 
Conn., N. Y., Pa., N. J., Mich., Minn. 
Cubocephalus atriclunis Townes and Gupta, 1962. Amer. Ent. Inst. Mem. 
2: 31,37,56. $, 9. 

baldaufi (Dalla Torre). Maine, Mass., Ind., Minn. 

*brevicomis Taschenberg. Alaska, Yukon, Colo., Mich., Que.; Europe. 

Phygadeuon brevicornis Taschenberg, 1865. Ztschr. Gesam. Naturw. 

Halle 25: 48. 9. 
Taxonomy: Townes and Gupta, 1962, pp. 32, 38, 69. 

*caligneus Townes. N. C. (Mt. Mitchell). 

Cubocephalus caligneus Townes, 1962. In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 40, 109. 9. 

*callicerus Townes. N. S., Que., Ont., Mich., Ohio, N, Y. in Alleghanian 
Fauna. 
Cubocephalus callicerus Townes, 1962. In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 33, 41, 115. $, 9. 

canadensis (Provancher). A subsp. of alacris (Cresson). 

Chaeretynima Ashmeadii Harrington. Remove from syn.; a valid species. 
Chaeretymma minuta Cushman. Remove from syn.; a valid subsp. of 
alacris (Cresson). 

*camosus Townes. Que. and N. H. to Mich. 

Cubocephalus carnosus Townes, 1962. In Townes and Gupta. Amer. 
Ent. Inst. Mem. 2: 35, 41, 114. $, 9. 

cincticomis (Cresson). Mich., Pa., N. Y., Ont., Que. 

circumcinctus circumcinctus (Provancher). Transf. to Parmortha Townes. 

circumcinctus flavipes (Harrington). Transf. to Parmortha Townes, 



ICHNEUMONIDAE 99 



♦contractus Townes and Gupta. Transcont. in Huds. and Transit. Zones. 

Cubocephalus contractus Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2: 30, 35,46. $,9. 

*denticulatus denticulatus Townes. Mich., Que., Maine, Vt., N. Y. 

Cubocephalus denticulatus denticulatus Townes, 1962. In Townes and 
Gupta, Amer. Ent. Inst. Mem. 2: 30, 36, 43. $, $. 

*denticulatus obscuripes Townes. Alaska (Curry). 

Cubocephalus denticulatus obscuripes Townes, 1962. In Townes and 
Gupta, Amer. Ent. Inst. Mem. 2: 42. $. 

*dreisbachi Townes. Atlantic to Rocky Mts.: Alta. in Canad. and Transit. 
Zones. 
Cubocephalus dreisbachi Townes, 1962. In Townes and Gupta, Amer. 

Ent. Inst. Mem. 2: 33, 39, 94. $,9. 

erythropus velox (Cresson), Transf. to Parmortha Townes, as syn. of par- 

vula (Gravenhorst). 

erythropygus (Provancher) . A subsp. of alacris (Cresson). 

*euryops Townes. Minn., Mich., Ont., Vt. 

Cubocephalus euryops Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 31, 37, 64. 5, 5. 

eximius (Cresson). Syn. of semirufus (Brulle). 

*flavipes Townes. Mich., N. Y. 

Cubocephalus flavipes Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 32, 38, 72. $,9. 

*hebes Townes. Sask., Mont. 

Cubocephalus hebes Townes, 1962. In Townes and Gupta, 1962. Amer. 
Ent. Inst. Mem. 2: 32, 38, 66. $, 9. 

*hirtipes hirtipes Townes. Alta., Idaho, Wash., Calif., N. Mex,, Colo., Wyo., 
Mont. (In mts.). 
Cubocephalus hirtipes hirtipes Townes, 1962. In Townes and Gupta, 
Amer. Ent. Inst. Mem. 2: 34, 40, 121, 122. $,9. 

*hirtipes trachodes Townes. N. H. (Mt. Washington). 

Cubocephalus hirtipes trachodes Townes, 1962. In Townes and Gupta, 
Amer. Ent. Inst. Mem. 122. $, 9. 

ignotus (Provancher). Syn. of nitidus nitidus (Provancher). 

impressus (Provancher). Que. to Minn, south to Ohio and Ga.; Colo. In 
Alleghanian and Carolinian faunas. 

*incisus atriventris Townes and Gupta. Ariz. (Greer). 

Cubocephalus incisus atriventris Townes and Gupta, 1962. Amer. Ent. 
Inst. Mem. 2: 44. 9. 

*incisus incisus Townes and Gupta. Mich., N. Y. 

Cubocephalus incisus incisus Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2: 29,36,44,45. $, 9. 

incognitus (Provancher). Ont., Vt., Mass., Wyo. 

inhabilis inhabilis Provancher, changed status. Atlantic ocean to Rocky 
Mts.; Alta. Removed from syn. of annulatus (Cresson) and i-e- 
duced to subsp. rank (teste Townes in Townes and Gupta, 1962, p. 
85). 

*inhabilis umbonater Townes. Rocky Mts. to Pacific Ocean ; Sask. 

Cubocephalus inhabilis umbonater Townes, 1962. In Townes and Gupta, 
Amer. Ent. Inst. Mem. 2: 85, 87. 5, 9. 

laticeps (Cresson). N. B., N. S., Mass., N. Y., N. J., S. C, Mich., Minn., S. 
Dak., Calif. 



100 HYMENOPTERA OF AMERICA NORTH OF MEXICO 



*lissopIeuris Townes and Gupta. Maine, N. Y., Pa., N. C. 

Cubocephalus lissopleuris Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2: 30, 59. $. 

maurus (Cresson). Calif., B. C., Alta., N. W. T., Mich., Ont., Que. 

*miarus Townes and Gupta. Maine, N. H., N. Y., S. C., Mich, Wis. 

Cubocephalus miarus Townes and Gupta, 1962. Mem. Amer. Ent. Inst. 
2: 31,37, 54. $, 9. 

*inicans Townes. Que. and P. E. I. to R. I. west, to Minn. 

Cubocephalus micans Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 34, 40, 112. 8, 9. 

*inolaris cervinus Townes and Gupta. Calif., Oreg. 

Cubocephalus mofaris cervinus Townes and Gupta, 1962. Amer. Ent. 
Inst. Mem. 2: 50, 51. $, 9. 

*nioIaris leechi Townes and Gupta. B. C. (Steelhead), 

Cubocephalus molaris leechi Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2 : 50. $. 

*molaris molaris Townes and Gupta. Yukon, B. C, Wyo., Colo., N. Mex., 
Minn., N. Y., Ont., Que. 
Cubocephalus molaris molaris Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2: 29, 36, 50. $,9. 

♦nigriventris (Thomson). B. C, Wash., Oreg., Colo., Mich., N. B., Que., 
Maine, N. H., N. Y., N. C, Vt.; Europe. 
Stenocryptus nigriventris Thomson, 1874. Opusc. ent., fasc. 6, p. 604. $,9. 

nitidus nitidus (Provancher) , changed status. Alta. to N. S. south to Mich., 

Pa., and R. I. Reduced to subsp. rank {teste Townes, in Townes 

and Gupta, 1962, pp. 79, 80). 
Cryptus occidentalis Provancher, 1875. Transf. from syn. of annulatus 

(Cresson). 
Cubocephalus ignotus (Provancher) 1886. 
Phygadeuon tiitidulus Provancher. Transf. from syn. of annulatus 

(Cresson). (All syn. teste Townes in Townes and Gupta, 1962, p. 

80). 

*nitidus totus Townes. Alaska, B. C, Wash., Oreg., Calif., Wyo. 

Cubocephalus nitidus totus Townes, 1962. In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 79. $,9. 

pallidas pallidus (Cresson), changed status. Mich, and Kans. to Ont. and S. 
C. in Carolinian fauna. Reduced to subsp. rank {teste Townes 
and Gupta, 1962, pp. 100, 102). 
Mesoleptus vultus Cresson. (Syn. teste Townes, in Townes and Gupta, 
1962, p. 102.) 

pallidus spissicomis (Provancher), changed status. B. C, Wis. to Que. south 
to Mich, and N. C. in Alleghanian fauna. Reduced to subsp. rank 
{teste Townes and Gupta, 1962, pp. 100, 101). 

*personatus Townes. Wash., Oreg., Calif. 

Cubocephalus personatus Townes, 1962. In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 35, 41, 105. $, 9. 

*pictus Townes. Mich., S. C. 

Cubocephalus pictus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 34, 41, 111. $, 9. 

*prolixus phaeolepis Townes and Gupta. Calif., Oreg., Wash., B. C. 

Cubocephalus prolixus phaeolepis Townes and Gupta, 1962. Amer. Ent. 
Inst. Mem. 2: 48. $, 9. 

*prolixus prolixus Townes and Gupta. Man., S. Dak., Mich., Minn., Que., 
Maine, Vt., N. H., N. Y. 
Cubocephalus prolixus prolixus Townes and Gupta, 1962. Amer. Ent. 
Inst. Mem. 2: 30, 35, 48, 49. $,9. 



ICHNEUMONIDAE 101 

rectus (Provancher). Que. to N. Y. west to Minn. 

*rufibasis Townes and Gupta. Mich. 

Cubocephalus rifibasis Townes and Gupta, 1962. Amer. Ent. Inst. Mem. 
2: 36,63. 9. 

schmuckii (Dalla Torre). Que., Maine, Mass., Vt., Md., N. C., S. C., Ga., 
Wis., Minn. 
Cryptus longicaudus Provancher. Misspelled longicaudis in Catalog, p. 
255. 

*scorteus Townes. Que., Yukon. 

Cubocephalus scorteus To\\Ties, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 31, 38, 70. $,9. 

*8eniifulvus Townes. Wash., Alaska. 

Cubocephalus semifulvus Townes, 1962. In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 32, 38, 71. 5, ?• 

semirufus (Brulle). Maine to N. C., west to Mich. Transf. from Alphabetic 
List of Unplaced Species, p. 408 {teste Townes, 1961. Ent. Soc. 
Wash. Proc. 63 (2) : 105). 
Cubocephalus eximius (Cresson). (Syn. teste Townes and Gupta, 1962, 
p. 59). 

signatus (Provancher). Transf. to Pamiortha Townes as subsp. of pleuralis 

(Thomson). 

spissicornis (Provancher). Subsp. of pallidus (Cresson). 

*sternoloplius Townes. Alaska, Calif., N. Mex., Mich., Ohio, Penna., N. Y. 
Cubocephalus sternolophus Townes, 1962, In Townes and Gupta, Amer. 

Ent. Inst. Mem. 2: 32, 38, 74. $, 9. 

*subpolitus Townes. N. Y., Conn., Que., Minn, in Alleghanian Fauna. 

Cubocephalus subpolitus Townes, 1962. In Townes and Gupta, Amer, 
Ent. Inst. Mem. 2: 35, 40, 120. $, ?. 

*tincticoxis Townes. B. C, Oreg., Calif., Colo. 

Cubocephalus tincticoxis Townes, 1962. In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 31, 55. $. 

vultus (Cresson). Syn. of pallidus pallidus (Cresson). 

♦Genus PLEOLOPHUS Townes 

Pleolophus Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. Mem. 
2: 7, 9, 10, 223. 

Type: "Phygadeuon basizonus Gravenhorst, as interpreted in 
this publication." Orig. desig. 

Place after Cubocephalus Ratzeburg, p. 255. 

*annulosus Townes. Maine, N. Y., Mich. 

Pleolophus anmtlosus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 224, 226. $. 

*astrictus Townes. Maine, N. H., Conn., Mass., N. Y., Mich., Alaska. 

Pleolophus astrictus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 224, 226, 242. $, 9. 

basizonus (Gravenhorst). N. B., Ont., Mich. Host: Neodiprion nanulus 
Schedl. Transf. from Aptesis Foerster (teste Townes, 1962. In 
Townes and Gupta, p. 235). Mispelled basizonia in Catalog. 
Biology: Finlayson, 1960. Canad. Ent. 92 (1) : 33.— Griffiths, 1961. Can- 
ad. Ent. 93 (11) : 1005-1010. 

*borrori Townes. Maine (Lincoln Co.). 

Pleolophus borrori Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 225, 228. 9. 

*clypealis Townes. Alaska (Nome). 

Pleolophus clypealis Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 224, 242. $. 



102 HYMENOPTERA OF AMERICA NORTH OF MEXICO 



*contractus Townes. B. C, Wash., Oreg., Calif. Hosts: PHstiphora erichso- 
nii (Htg.), ILamhdina fiscellaria lugiihrosa (Hist.). 
Pleolophus contractus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 225, 237. $,9. 
*coriaceu8 Townes. Sask., Alta. 

Pleolophus coriaceus Townes, 1962. In Townes and Gupta, Amer, Ent. 
Inst. Mem. 2: 225, 229. $. 

*furvus Townes. Colo. (Rocky Mt. N. P.). 

Pleolophus furvus Townes, 1962. In Townes and Gupta, Amer. Ent, Inst. 
Mem. 2: 225, 228. $. 

*grossus Townes, N. Y., N. J., Pa., Md., D. C, N. C, Mich., Kans., N. Dak., 
B. C, Wash. Oreg. 
Pleolophus grossus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 225, 226, 229. $, 9. 

indistinctus (Provancher) . Sask., Man., Ont., Iowa, N. C. Hosts: Neodiprion 
ahhotii (Leach), A^. pratti banksianae Rohw., N. nanulus Schedl, 
N. sertifer (Geof.), N. stvainei Midd., N. virginianus Rohw., N. 
sp., Pristiphora geniculata (Htg.). Transf. from Aptesis Foerster, 
p. 255 (teste Townes, 1962. In Townes and Gupta, p. 238). 
Biology: Finlayson, 1960. Canad. Ent. 92 (1) : 32. 

*nigribasis Townes. Colo., N. Mex. 

Pleolophus nigribasis Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 225, 237. $. 

'^pilatus Townes. Labrador, Mich., Minn. 

Pleolophus pilatus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 224, 225, 227, $,9. 

rubrocinctus rubrocinctus (Provancher), Ont., N. H., Mass., Ohio. Transf. 
from Aptesis Foerster (teste Townes 1962. In Townes and Gupta, 
p. 231, 232). 

^rubrocinctus rufifemur Townes. Alaska, B. C, Alta., Wash., Oreg., Calif., 

Idaho, Mont., Wyo. 
Pleolophus rubrocinctus rufifemur Townes, 1962. In Townes and Gupta, 

Amer. Ent. Inst. Mem. 2: 231, 233. $,9. 
Aptesis rubrocincta n. subsp. Transf. from Aptesis Foerster, p. 255. 

♦Genus RHYTURA Townes 

Rhrtura Townes, 1962, In Townes and Gupta, Amer, Ent. Inst, Mem. 2: 
9, 10, 244, 

Type: Rhytura pendens Townes, Orig, desig, and by mono- 

typy. 

Place before Aptesis Foerster, p. 255. 

*pendens Townes. Que. and Ont. south to Ga., west to Minn. 

Rhytura pendens Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 244. $, 9. 

Genus APTESIS Foerster, p. 255 

*Oxytaenia Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 182. Syn. 
by Townes, 1962. In Townes and Gupta, p. 246. 

Type: (Oxytaenia rufolineata Cameron) = barda (Cresson). 
Included by Cameron, 1904. 

*aIbifrons Townes. N. Y. (Rome). 

Aptesis albifrons Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 248,267. $. 

*alpestris Townes. B. C. to Calif., Idaho, Colo., Ariz. 

Aptesis alpestris Townes, 1962. In Townes and Gupta, Amer, Ent. Inst. 
Mem. 2: 250, 255. $,9. 



ICHNEUMONIDAE 103 

'^'anaulus Townes. 

Aptesis anaulus Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 248, 272. 5- 

arridens (Gravenhorst). Transf. to Opidnus Foerster, p. 256 

*atrox Townes. Ohio (West Unity). 

Aptesis atrox Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 249, 269. $. 

*barda Carolina Townes. Que. to N. C. west to Mich., Iowa, Kans. A. barda 
barda Cresson occurs in Mexico. 
Aptesis barda Carolina Townes, 1962. In Townes and Gupta, Amer. Ent. 

Inst. Mem. 2: 270,271. 3, 9. 

ftasizonia [sic!] (Gravenhorst). Transf. to *Pleolophus Townes. 

*breviaria Townes. N. Y., Ohio, Mich., Minn. 

Aptesis breviaria Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 249,250,253. $,9. 

*catulus Townes. N. Y. to N. C., Kans. 

Aptesis catulus Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 249,250, 263. 5, 9. 

*fastigata Townes. Alta., Oreg., Calif., Wyo. 

Aptesis fasiigata Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 249,250, 267. $, ?. 

*fuscitibia Townes. Maine, Mass., R. I., N. Y., N. C., Mich. 

Aptesis fuscitibia Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 249, 251, 262. $, 9. 

^gracilis Townes. Alta., Mich. 

Aptesis gracilis Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 249, 251. $. 

*incompta Townes. S. C. (Cleveland) 

Aptesis incompta Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 249,251,265. $, ?. 

*inculta Townes. Que. to N. C. west to Mich. 

Aptesis inculta Townes, 1962. In Townes and Gupta, Amer. Ent, Inst. 
Mem. 2: 249, 250, 257. S, ?. 

indistincta (Provancher). Transf. to *Pleolophus Townes. 

iridescens (Cresson). Transf. to Schenkia Foerster, p. 257. 

*lissopIeuris Townes. N. C. (Mt. Pisgah). 

Aptesis lissopleuris Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 251, 257. 9. 

rubrocincta rubrocincta (Provancher). Transf. to *Pleolophu8 Townes. 

*scabra Townes. Md. (Glen Echo). 

Aptesis scabra Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 251,266. 9. 

*8egnis femorata Townes. Alta., Sask., Calif., Colo., Wyo. 

Aptesis segnis femorata Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 260. g, 9. 

segnis segnis (Provancher), changed status. Que. and Mich, to S. C. Reduced 
to subsp. rank {teste Townes and Gupta, 1962, pp. 260, 261). 
Aptesis tenthredinaruni (Rohwer). (Syn. teste Townes, 1962. In Townes 
and Gupta, 1962, p. 261.) 

*subguttata (Gravenhorst). Que., Europe. Host: Diprion her cyniae (Htg.). 
Crvptus subguttatus Gravenhorst, 1829. Ichn. Europaea, v. 2, p. 610. $. 
Biology: Finlayson, 1960. Canad. Ent. 92 (12) : 931, 932 (larva). 
Taxonomy: Townes in Townes and Gupta, 1962. Amer. Ent. Inst. Mem. 
pp. 247, 248. 



104 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

tenthredinarum (Rohwer). Syn. of segnis segnis (Provancher). 

*verrucata Townes. Maine, N. H., N. Y., Ohio, Mich. 

Aptesis verrucata Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 249,250, 252. $,9. 

*yosemite Townes. Calif., Wyo. 

Aptesis yosemite Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 250, 254. $. 

Genus MEGAPLECTES Foerster, pp. 255-256 

monticola blakei (Cresson). Yukon, N. W. T., and B. C. to Labrador and N. 
S. south to Oreg. and Pa., and Mts. of N. C. 

*moniicola dentatus Uchida. Alaska, Yukon (Dawson) ; Japan. 

Megaplectes monticola var. dentata Uchida, 1930. Hokkaido Imp. Univ. 
Faculty Agr. Jour, 25: 304. S, ?. 

Genus OPIDNUS Foersfer, p. 256 

*Ophiilnus Viereck, 1914. U. S. Natl. Mus. Bui. 83: 105. Misspelling. 
Syn. of Oresbius Marshall, 1867, teste Townes, Momoi, and Townes, 1965, 
p. 157. 

albicoxus albicoxus (Provancher), changed status. Host: Pristiphora erich- 
sonii (Htg.). Reduced to subsp. rank {teste Townes, 1962. In 
Townes and Gupta, p. 141). 

albicoxus schoberi Dalla Torre, changed status. Reduced to subsp. rank 
(teste Townes, 1962. hi Townes and Gupta, p. 141). 
*Cryptus koebelei Rohwer, 1917. Proc, U. S. Natl. Mus. 53: 158. N. 
name for ferrugineus Ashm. nee Smith; unnecessarily proposed. 
{Teste Townes, 1962. In Townes and Gupta, p. 141.) 

*alexanderi Townes. Que., Vt., Alta. 

Opidnus alexanderi Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 127, 131, 145. ^,9. 

*aniptus Townes. Alaska, Calif., Colo. 

Opidnus aniptus Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 127,131, 151. $, 9. 

arridens (Gravenhorst). N. H., Mass., N. Y., Ont., Mich. Transf. from Ap- 
tesis Foerster, p. 255 {teste Townes, 1962. In Townes and Gupta, 
p. 131). 

*ciliatus Townes. Que. (Payne Bay). 

Opidnus ciliatus Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 129, 164. ?. 

*cognatus Townes. B. C, Alta., Wash., Oreg., Calif., Colo., N.S., Que., N. H., 
N. Y. 
Opidnus cognatus Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 128, 131, 148. 5, ?. 

dorsator (Ashmead). Alaska (Nome). 

*fulvibasis Townes. Minn, to Que. south to N. C. 

Opidnus fulvibasis Townes, 1962. In Townes andd Gupta, Amer. Ent. 
Inst. Mem. 2: 127, 130, 144. 5, $• 

*Iindrothi Townes. Alaska (Umiat). 

Opidnus lindrothi Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 130, 158. ?. 

*major Townes. N. H., Que. 

Opidnus major Townes, 1962. In Townes and Gupta, Amer. Ent, Inst. 
Mem. 2: 129, 155. $. 

^mucronatus Townes. Maine, Mass., N. Y., Mich. 

Opidnus mucronatus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 127, 131, 150. $,9. 



ICHNEUMONIDAE 105 

nigriceps (Provancher) . Syn. of tegularis (Provaiicher). 

nivarius (Brues). Alaska (Point Barrow). 

Taxonomy: Townes, 1962. In Townes and Gupta, pp. 128, 130, 158 (first 
descr. $ ) . 

orbitalis (Provancher). Syn. of tegularis (Provancher). 

*orbitus Townes. N. H., Yukon. 

Opidnus orbitus Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 129, 130, 157. $, 9. 

*oresbius Townes. Wash., B. C. 

Opidnus oresbius Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 128, 136. $. 

*parallelus Townes. N. H., N. Y., Que., Man., Colo. 

Opidnus parallelus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 127, 130, 165. $, 9. 

*quarlriceps Townes. Alaska, N. W. T. 

Opidnus quadriceps Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 128, 129, 168. 5, ?• 

*rufinotuni Townes. Alaska. 

Opidnus rufinotum Townes, 1962. In Tovmes and Gupta, Amer. Ent. 
Inst. Mem. 2: 128, 161. $. 

schoberi (Dalla Torre) . Subsp. of albicoxus (Provancher). 

*sublaevis niiltomerus Townes. Wash. (Mt. Rainier). 

Opidnus sublaevis miltomerus Townes, 1962. In Townes and Gupta, 
Amer. Inst. Mem. 2: 133, 134, 135. $. 

*sublaevis sublaevis Townes. Colo., Wyo. 

Opidnus sublaevis sublaevis Townes, 1962. In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 126, 129, 133, 134. $, ?. 

*taeniatus Townes. N. H., Que., N. W. T., Yukon. 

Opidnus taeniatus Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 128, 130, 154. $, 9. 

tegularis (Provancher). Minn, to N. S. and Pa.; Wyo., Colo. Host: Pristi- 
phora erichsonii (Htg.). 
Ischnus placidus Provancher. Transf. from syn. Schenkia graminicola 

graminicola (Gravenhorst) . 
Phygadeuon orbitalis Provancher. 
Phygadeuon nigriceps Provancher. 

(All syn. teste, Townes and Gupta, 1962, p. 146.) 

*teres Townes. Alaska, Yukon. 

Opidnus teres Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 128, 129, 167. $, 9. 

*tibialis Townes. Alaska, Yukon, N. W. T., Que. 

Opidnus tibialis Townes, 1962. In Townes and Gupta, Amer. Ent. Inst, 
Mem. 2: 128, 130, 163. $, 9. 

trifasciatus (Ashmead). Yukon, N. W. T., Que. 

*tsugae infulatus Townes. N. C, Labrador. 

Opidnus tsugae infulatus Townes, 1962. In Townes and Gupta, Amer. 

Ent. Inst. Mem. 2: 137, 140. S, 9. 

*tsugae inferior Townes. Ariz., Colo., Wyo., Idaho, Alta. 

Opidnus tsugae interior Townes, 1962. In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 137, 139. $, 9. 

*tsugae ruficoxa Townes. Calif. Host: Cocoon on Abies concolor. 

Opidnus tsugae ruficoxa Townes, 1962. In Townes and Gupta, Amer. 
Ent. Inst. Mem. 2: 137, 138. $, 9. 

tsugae tsugae (Cushman), changed status. Host: Neodiprion sp. Reduced to 
subsp. rank {teste Townes, 1962. In Townes and Gupta, p. 137). 



106 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

♦tumulus Townes. N. W. T., Yukon, Alaska. 

Opidnus tumulus Townes, 1962. In Townes and Gupta, Amer. Ent. Inst. 
Mem. 2: 129, 160. $. 

*vockerothi Townes. Que. (Great Whale River). 

Opidnus vockerothi Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 166. $,9. 

*vultuosus Townes. Transcont. in Canadian zone. 

Opidnus vultuosus Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 127, 130, 152. 5, ?• 

Genus SCHENKIA Foerster, pp. 256-257 

*amphileuca Townes and Gupta. N. Y., Mich., Minn. 

Schenkia amphileuca Townes and Gupta, 1962. Amer. Ent. Inst. Mem. 
2: 200,201. $, 9. 

brevicauda brevicauda (Ashmead). Syn. of iridescens iridescens (Cresson). 

graminicola graminicola (Gravenhorst) . Yukon; Europe. 

*Cryptus brevicornis Gravenhorst, 1829. Ichn. Europaea, v. 2, p. 511. $. 

*Cryptus humilis Gravenhorst, 1829. Ichn. Europaea, v. 2, p. 604. $. 

Ischnus placidus Provancher. Transf. to syn. of Opidnus tegularis (Pro- 
vancher), p. 256. 

Phygadeuon rectus Provancher. Valid subspecies of graminicola. 

Phaeogenes tuberculifer Provancher, 1882. Syn. of graminicola recta 
(Provancher) . 

Cry plus pubescens Provancher, 1886. Syn. of graminicola recta (Pro- 
vancher). 

Cry plus hechfellneri Dalla Torre, 1902. Syn. of graminicola recta (Pro- 
vancher). 

*graminicola hesperus Townes and Gupta. B. C., Wash., Oreg. 

Schenkia graminicola hesperus Townes and Gupta, 1962. Amer. Ent. 
Inst. Mem. 2: 204, 207. $,9. 

graminicola recta (Provancher). changed status. B. C. to Labrador and 
Que.; south to Minn., Mich., N. Y. 

Phaeogenes tuberculifer Provancher (syn. teste Townes and Gupta, 1962, 
p. 205). 

Cryptus pubescens Provancher (syn. teste Townes and Gupta, 1962, p. 
205). 

Cryptus hechfellneri Dalla Torre (syn. teste Townes and Gupta, 1962, p. 
205). 
iridescens iridescens (Cresson), changed status. Ont. and N. S. to Ga. west to 
Mich, and Tenn. Transf. from Aptesis Foerster, p. 255 and re- 
duced to subsp. rank {teste Townes and Gupta, 1962, pp. 210, 
214). 

Cryptus brevicauda Ashmead (syn. by Townes and Gupta, 1962, p. 214). 

*iridescens melanops Townes and Gupta. B. C, Wash., Oreg., ?Idaho. 

Schenkia iridescens melanops Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2: 211,216. 3,5. 

*iridescens oreas Townes and Gupta. Alaska to Labrador south to Oreg., 
Colo., and Mich. 
Schenkia iridescens oreas Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2: 211,212. $, 9. 

♦iridescens pumila Townes and Gupta. Alaska to Wash., east to Que. and N. 
H., also Colo. 
Schenkia iridescens pumila Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2: 211. $, 9. 

*leucopus Townes and Gupta. Mich., Ohio, N. Y., N. J., S. C. 

Schenkia leucopus Townes and Gupta, 1962. Amer. Ent. Inst. Mem. 2: 
200,201,217. $, 9. 



ICHNEUMONIDAE 107 

*lonchura Townes and Gupta. Yukon. 

Sciienkitt loiicliura Townes and Gupta, 1962. Amer. Ent. Inst. Mem. 2: 
201, 217. $. 

*parallela evittata Townes and Gupta. Wash. 

Schenkia parallela evittata Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2: 208,209. . 

♦parallela parallela Townes and Gupta. Ont. to S. C, Sask. 

Schenkia parallela parallela Townes and Gupta, 1962, Amer. Ent. Inst, 
Mem, 2: 200,207,208, $,9. 

Genus GIRAUDIA Foerster, p. 257 

ferruginea (Cushman). Ont. to Md. and D. C, west to Mich, in Carolinian 
Fauna. Host: Tomostethus multicinctus Roh. Valid species. Re- 
moved from syn. of plana (Provancher {teste Townes and Gupta, 
1962, p. 186). 

*fulva fulva Townes and Gupta. Wash., Oreg., Calif. 

Giraudia fulva fulva Townes and Gupta, 1962. Amer. Ent. Inst, Mem. 2, 
pp. 184, 185, 189. $, ?. 

*fulva nigrithorax Townes and Gupta, B. C, Alta., Idaho, Oreg., Utah, Colo., 
N. Mex. 
Giraudia fulva nigrithorax Townes and Gupta, 1962. Am.er. Inst. Mem. 
2: 189, 190. $, 9. 

*leucopsis TowTies and Gupta. Calif. 

Giraudia leucopsis Townes and Gupta, 1962, Amer, Ent, Inst, Mem. 2: 
184, 196. $. 

♦minor Townes and Gupta. N. C, S. C. 

Giraudia minor Townes and Gupta, 1962. Amer. Ent. Inst. Mem. 2: 184, 
185, 196. $, 9. 

plana (Provancher). Ont., Maine, Pa., N. C, S. C, Ga., Ohio, Mich., Minn., 
Iowa, Delete host record. 
Aptesis ferruginea Cushman, 1935, Delete from syn., a valid species. 
IMesostenus ruficoxus Provancher. Transf. from syn. of Cubocephalus 

annulatus (Cresson) {teste Townes and Gupta, 1962, p. 192). 
IMesostenus provancheri Dalla Torre. Transf. from syn. of C. annulatus 
(Cresson) {teste Townes and Gupta, 1962, p. 192). 

*punctifera Townes and Gupta. Ont. and Que. to Ga., west to Mich, and Wis, 
Giraudia punctifera Townes and Gupta, 1962, Amer, Inst. Mem. 2: 184, 
185, 194, $, 9. 

rufa (Provancher), Que. to N. C. west to Ont., Minn, in Alleghanian Fauna. 

♦semiflava Townes and Gupta, Que., Maine, N. Y., Mich., N. Mex. 

Giraudia semiflava Townes and Gupta, 1962. Amer. Ent. Inst. Mem. 2: 
184,185, 191. $, 9. 
*subaequalis Townes and Gupta. N. Y., N. C, S. C, Ga. 

Giraudia subaequalis Townes and Gupta, 1962. Amer. Ent, Inst, Mem. 2: 
184, 185, 197, $, 9. 

Genus POLYTRIBAX Foerster, p. 257 

Plesignathus Foerster, 1868. Transf. from syn. of Phrgadeuon Graven- 
horst, p. 248 {teste Perkins, 1962, p. 446), Change type citation 
to: 

Type: Phygadeuon fulvescens Cresson, Included by Ashmead, 
in Smith, 1900 (June). 
*Neleophron Foerster, 1868. Naturh. Ver. Rhinelande Verh. 25: 184. No 
species. (Syn. teste Perkins, 1962, p. 438.) 

Type: Cryptus arrogans Gravenhorst. Included and desig, by 
Perkins, 1962. 



108 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Epiphohu8 Foerster, 1868. Naturh. Ver. Rheinlande 25: 183. No spe- 
cies. (Syn. teste Townes and Gupta, 1962, p. 170.) 

Type: Cry plus rufipes Gravenhorst. Included and desig. by 
Townes, 1962. 
Steriphocryptus Cameron, 1903. Transf. from syn. of Trachysphyrus Ha- 
liday, p. 259. (Syn. teste Townes, Townes and Gupta, 1961, p. 
137.) 
*Chasmocryptus Uchida, 1936. Ins. Mats. 11: 16. (Syn. teste Townes, 
1957, Ent. Soc. Wash. Proc. 59: 103.) 

Type: (Plectocryptus hokkaidensis JJ chida.) = penetrator 
(Smith). Orig. desig, 

contignus (Cresson). N. S., Ont., N. C, Ala., Minn., N. Dak., Nebr., Kans. 

crotchii crotchii (Cresson), changed status. Transcont. in Canad. and north. 
Transit. Zones. Reduced to subsp. rank {teste Townes and Gupta, 
1962, pp. 176, 177). 

*crotchii fumipennis Townes and Gupta. B. C, Calif., Ariz., Colo., N. Mex. 
(in Canadian Zone). 
Polytribax crotchii fumipennis Townes and Gupta, 1962. Amer. Ent. 
Inst. Mem. 2, pp. 176, 179. $, 

*crotchii hegpeinis Townes and Gupta. Calif., Oreg., B. C. 

Polytribav crotchii hesperus Townes and Gupta, 1962. Amer. Ent. Inst. 
Mem. 2, pp. 176, 180. $. 

fulvescens (Cresson). Wash., Oreg. 

pallescens (Viereck). S. C, Ga.; Mexico. Host: 1 Diacrisia virginica (F.). 

Genus RHEMBOBIUS Foerster, pp. 257-258 

*Vlothymus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 185. 
(Syn. teste Townes, Townes, and Gupta, 1961, p. 131; Perkins, 
1962, p. 462.) 

Type: Ichneumon perscrutator Thunberg. Included and de- 
signated by Perkins, 1962. 

abdominalis abdominalis (Provancher). Alta. Host: 1 Xylota. 

""abdominalis montanus Townes and Gupta. Alaska, Alta., Sask., Idaho, 
?Mont., S. Dak., Colo. 
Rhembobius abdominalis montanus Townes and Gupta, 1962. Amer. 

Ent. Inst. Mem. 2: 275. $, $. 

abdominalis pacificus (Harrington). Alaska, B. C, Oreg., Idaho. 

*parcus Townes. Sask., Man., Mich. 

Rhembobius parous Townes, 1962. In Townes and Gupta, Amer. Ent. 
Inst. Mem. 2: 273. $, . 

Genus DEMOPHELES Foerster, p. 258 

Change type citation to : 

Type: (Phygadeuon caliginosus Gravenhorst ?, not $)z=:cor- 
ruptor (Taschenberg). 

Townes places this genus in the Hemitelini ( = Gelini Townes) . 

corrupter maturus (Provancher), changed status. Maine and Que. to Ga., 

west to Ont., Mich. ? Alta. Upper Austral, to Canad. Zones. 
Reduced to subsp. rank (teste Townes, 1963. Ent. Soc. Wash. 
Proc. 65 (1): 48). 

*corruptor rufatus Townes. Wash., Colo., N. Mex. 

Demopheles corruptor rufatus Townes, 1963. Ent. Soc. Wash. Proc. 65 
(1) : 48,49. 9. 

Tribe SPHECOPHAGINI 

Taxonomy: Townes and Townes, 1962. U. S. Natl. Mus. Bui. 216 (Pt. 
3) : 16, 20, 522-523 (Sphecophagina). 



ICHNEUMONIDAE 109 

Genus SPHECOPHAGA Westwood, p. 258 

vesparum burra (Cresson), changed status, Transcont. in Canad. and Transit. 
Zones. S. vesparum vesparum occurs in Europe. Reduced to 
subsp. rank {teste Townes and Townes, 1962, pp. 525, 526). 

Tribe CRYPTINI 
( = Mesostenini of Catalog) 

Revision: Townes and Townes, 1962. U. S. Natl. Mus. Bui. 216 (Pt. 3) : 
I-VII, 1-602 (Mesostenini) 

♦Subtribe BARYCERATINA 
Place as first subtribe of Cryptini, p. 258. 

*Genus WHYMPERIA Cameron 

Whymperia Cameron, 1903. Entomologist 36: 122. 

Type: Whymperia carinifrons Cz.me\'ox\. By monotypy. 
*ProtocrYptus Schmiedeknecht, 1904. Opusc. Ichn., Band 2 (fasc. 6), p. 
414. Syn (teste Towiies and Townes, 1962, p. 22). _ 

Type: Protocryptus grandis Sclimiedeknecht. Desig. by Vier- 
eck 1914. 
*Zamansa Viereck' 1912. U. S. Natl. Mus. Proc. 42: 647. Syn. with 
Protocryptus by Townes, 1946, Bol. Ent. Venezolana 5 (2) : 33. 

Type : Cryptus aztecus Cresson. By monotypy and orig. desig. 

Place as first genus in subtribe Baryceratina, p. 258. 

♦tricolor Townes. Ariz. 

Whymperia tricolor Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 22. $. 

Genus ChristoUa Brulle, pp. 258-259 

Syn. of Baryceros Gravenhorst, infra. 

Lamprocryptidea Viereck. Delete from Catalog. A valid Neotrop. genus 

(teste Townes and Townes, 1962, p. 12). 
Haplomus Szepligeti. Syn. of Lamprocryptidea Viereck (teste Townes 

and Townes, 1962, p. 12). 
Aplomiana Strand. Syn. of Lamprocryptidea Viereck. 

*Genus BARYCEROS Gravenhorst 

(= ChristoUa Brulle, pp. 258-259) 

Baryceros Gravenhorst, 1829. Ichn. Europaea, v. 2, p. 777. 

Type : Baryceros guttatus Gravenhorst. By monotypy. 
ChristoUa Brulle. (Syn., teste Townes, 1961. Ent. Soc. Wash. Proc. 63 

(3): 167.) 
*Neochristolia Blanchard, 1936. Rev. Argentina de Ent. 1 (2) : 40. (Syn. 
by Townes, loc. cit. supra, p. 167.) 

Type: Neochristolia eucleides Blanchard. By monotypy and 
orig. desig. 

armatus (Provancher). Syn. of audax saundersi (Cresson). 

audax audax (Cresson), changed status. Mass., Conn., N. Y., Mich., Wis. 
south to S. C., Ala., Kans. Reduced to subsp. rank (teste Townes 
and Townes, 1962, pp. 27, 28). 
*Mesostenus exaptus Cresson, 1878. Canad. Ent. 10: 208. 9. (Syn. by 
Cushman, 1929. U. S. Natl. Mus. Proc. 74 (Art. 16) : 25.) 

audax saundersi (Cresson), changed status. Que., Ont., south to R I., N. Y., 
Ohio, Mich. Reduced to subsp. rank (teste Townes and Townes, 
1962, pp. 27, 29). 
ChristoUa armata (Provancher) (syn. teste Townes, and Townes, 1962, p. 
29). 



110 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

candidus (Cresson). 

ChristoUa dumetorum Townes (syn. teste Townes and Townes, 1962, p. 
25). 

dumetorum (Townes). Syn. of candidus (Cresson). 

fortis (Cresson). Pa., Ohio, Mich., W. Va., Ga. 

*halii Townes. Ariz. 

Baryceros halli Townes, 1962. In Townes and Townes, U. S. Natl. Mus, 
Bui. 216 (Pt. 3) : 24,30. 8. 

saundersi (Cresson). Subsp. of audax (Cresson). 

texanus (Ashmead). Va., N. C, Ind., Ky., Ark., Okla. ; Mexico. 

*Subtribe CRYPTINA 
(•=. tAesos^enlna Townes) 
Place as second subtribe of Cryptini, p. 259 

Genus CRYPTUS Fabricius, Sup. 1, p. 46 

( = Trochysphyrus of Catalog, pp. 259-263) 

Meringopus Foerster. Delete type citation and replace with: Cryptus 
recreator Fabricius. Included by Tschek, 1870 {teste Townes, 
Townes, and Gupta, 1961, p. 163). 
Itamoplex Foerster. Change "Desig. by Viereck, 1914" to: Included by 

Ashmead in Smith, 1900. 
*Costniocryptus Cameron, 1902. Amer. Ent. Soc. Trans. 28: 371. 

Type: Cosmiocryptus violaceipennis Cameron. By monotypy. 
Steriphocryptus Cameron, 1903. Transf. to syn. of Polytribax in (Apte- 

sini) =Hcinigasterini, p. 257. 
*Buathra Cameron, 1903. Ent. Soc. London, Trans. 1903 : 233. 
Type: Buathra rufiventris Cameron. By monotypy. 
*Plesiocryptus Cameron, 1903. Ztschr. Syst. Hym. Dipt. 3 : 299. 

Type : Plesiocryptus carinifrons Cameron. By monotypy. 
*Bathycrisis Cameron, 1905. Spolia Zeylanica 3 : 96. 

Type: Bathycrisis striaticollis Cameron. By monotypy. 
*Mesocryptus Szepligeti, 1916. Mus. Nat. Hungarici Ann. 14: 251. 
Preocc. 

Type: Mesocryptus pulcherrimus Szepligeti. By monotypy. 
*Pseudomesocrypius Strand, 1917. Ent. Ztschr. 10: 137. N. name for 

Mesocryptus. 
*Nippocryptus Uchida, 1936. Ins. Matsumurana 11: 3. 

Type: {Hemiteles suzukii Matsumura) = vittatorius (Jurine). 
By monotypy. 
*Neocryptopteryx Blanchard, 1947. Zool. Mus. Hist. Natl. Montevideo 
Comun. 2 (Pt. 42) : 1. 

Type : I\eocryptopteryx orientalis Blanchard. By orig. desig. 
Taxonomy: Townes, Townes and Gupta, 1961, pp. 163-164 (Generic 
syn.). 

albitarsis albitarsis (Cresson). Hosts: Heliothis zeae (Boddie) ; Macronoctua 
onusta Grt. 
Mesostenus tarsatus Provancher. Transf. to Mesostenus Gravenhorst, p. 
265, as questioned syn. of temporalis Townes. 

"^albitarsis argentifrons Cameron. Newfoundland to B. C. south to R. I., N. Y., 

Tex., and Calif.; Mexico, Costa Rica. 
*Cryptu8 bicolor Smith, 1879. Descr. of N. sp. of Hym. in the Coll. Brit. 

Mus., p. 231. Preocc. 
*Cryptus monticola Cameron, 1885. Biol. Cent.-Amer., Ins. Hym., v. 1, p. 

203. Preocc. 
Cryptus argentifrons Cameron, 1885. Biol. Cent.-Amer., Ins. Hym. 1: 

204. 
*Cryptus eggeri Dalla Torre, 1902. Cat. Hym., v. 3 (Pt. 2) p. 569. N. 

name for bicolor Smith. 



ICHNEUMONIDAE 111 

Trachysphyrus albitarsis coloradensis (Ashmead). 

Trachysphyrus inornatus (Pratt). 

(All syn. teste Townes and Townes, 1962, pp. 248-249). 

albitarsis coloradensis (Ashmead). Syn. of albitarsis argentifrons Cameron. 

albitarsis niutatus Pratt, changed status. 

Cry plus mutatus Pratt. Reduced to subsp. rank {teste Townes and 

Townes, 1962, pp. 244, 251). 
Cryptus curticaudus Pratt (syn. teste Townes and Townes, 1962, p. 251). 

albitarsis rufovinctus Pratt, changed status. B. C. Host: Trichoplusia sp. 

Cryptus rufovinctus Pratt. Reduced to subsp. rank (teste Townes and 
Townes, 1962, pp. 244, 253). 

altonii Dalla Torre. A. subsp. of laborator (Thunberg). 

arcticus Schiodte. N. W. T., Calif. (Arctic and Arctic-Apline). 

aridus aridus Pratt, changed status. B. C, Oreg., Calif., Colo. Reduced to 
subsp. rank (teste Townes and Townes, 1962, pp. 183, 229). 

aridus ocellaris Pratt, changed status. Oreg., Calif. Reduced to subsp. rank 
(teste Townes and Townes, 1962, pp. 229, 230). 

associatus Pratt. Syn. of persimilis Cresson. 

asymmetricus asymmetricus Pratt, changed status. B. C. and Sask. south to 
Mts. of Calif, and Ariz. Reduced to subsp. rank (teste Townes 
and Townes, 1962, p. 206). 

asymmetricus mirabilis Pratt, changed status. Reduced to subsp. rank (teste 
Townes and Townes, 1962, pp. 206, 207). 

bilobatus Pratt. A subsp. of symmetricus Pratt. 

calescens robustus Cresson, changed status. Alaska, Yukon, B. C. to Man. 
south to Calif, and N. Mex. in Canad. and Huds. Zones. Reduced 
to subsp. rank (teste Townes and Townes, 1962, p. 201). C. cales- 
cens calescevs Gravenhorst is European. 

crassipes Pratt. Transf. to Reptatrix Townes, infra. 

crassisculptus Pratt. A subsp. of rugosiscutum Pratt. 

crassulus Pratt. Mich. 

curticaudus Pratt. Syn. of albitarsis mutatus Pratt. 

dims Cresson. B. C. and Idaho south to Calif, and Ariz. 

Cryptus nitschei Dalla Torre. Remove from syn.; valid subsp. of rela- 
tivus Cresson. 

dorsicarinatus Pratt. Alaska, Alta., Mont., Colo. 

eleucopus Walkley. Syn. of relativus nitschei Dalla Torre. 
Cryptus eleucopus Walkley. Sup. 1, p. 46. 

*fasciatus (Townes), n. comb. Ariz. 

Trachysphyrus fasciatus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 180, 188. $, 9. 

flagellatus Pratt. A subsp. of rugosiscutum Pratt. 

genatus Pratt. Ariz., Utah. 

inornatus Pratt. Syn. of albitarsis argentifrons Cameron. 

iroquois Viereck. A subsp. of moschator (Fabricius). 

*krombeini (Townes), n. comb. W. Va. (Lost River State Park, Hardy Co.). 
Trachysphyrus krombeini Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 185, 243. 9. 

laborator altonii Dalla Torre, changed status. Host: Melanolophia imitata 
(Wlkr.). Reduced to subsp. rank (teste Townes and Townes, 1962, 
pp. 209,210). 



112 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

laborator fabric!! Schiodte. 

Taxonomy: Townes and Townes, 1962, pp. 214-215. 

laborator perplexus Cresson, changed status. Reduced to subsp. rank (teste 
Townes and Townes, 1962, pp. 209, 213). 

lat!genal!s Pratt. Alta., Mont., Ont. Host: Loxostcge sticticalis (L.). 

Biology: Ullyett, 1949. Canad. Ent. 81: 285-299; 82: 1-11 (as inor- 

natus). 
Taxonomy: Townes and Townes, 1962, pp. 184, 242-243. 

leucopus Ashmead. Syn. of relat!vus nitschei Dalla Torre. 

*Iochmaius (Townes) , n. comb. Ariz. 

Trachysphyrus lochmaius Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3): 183, 216. $, ?. 

luctuosus luctuosus Cresson, changed status. Reduced to subsp. rank (teste 
Townes and Townes, 1962, pp. 183, 221; also $ descr.). Other 
subspp. of luctuosus Cress, occur in Europe and Japan. 

*inent!gus nient!gus (Townes) , n. comb. Ariz. 

Trachysphyrus mentigits meniigus Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) ; 184, 234. $, 9. 

*inent!gus rhodomerus (Townes), n. comb. Alaska, Yukon, Wash, to Calif., 
Ariz., Utah, Colo., N. Mex. 
Trachysphyrus tnentigus rhodomerus Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 234, 235. $, ?. 

*ment!gus rufator (Townes), n. comb. Calif. (Blue Lake, Lassen Co.). 

Trachysphyrus tnentigus rufator Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 234, 235. ?. 

mimicus Pratt. A subsp. of tejonensis Cresson. 

in!n!mus Pratt. Sask., N. Mex., Nev. 

mirabilis Pratt. A subsp. of asymnietricus Pratt. 

moschator iroquo!s (Viereck), changed status. N. J., Ohio, Mich. Reduced to 
subsp. rank (teste Townes and Townes, 1962, pp. 223, 224). C. 
moschator moschator (Fabricius) is European. 

*murorum Tschek. Man., Calif. ; Europe. 

Cryptus murorum Tschek, 1872. Zool.-Bot. Gesell. Wien. Verhandl. 22: 

234. $, ?. 

mutatus Pratt. A subsp. of alb!tars!s (Cresson). 

ocellaris Pratt. A subsp. of ar!du8 Pratt. 

pacificus Cresson. B. C. to Calif, in Cand., Transit., U. Austral Zones. 

Cryptus punicus Cresson. Remove from syn. A valid subsp. of relat!vu8 

Cresson. 
Cryptus californicus Ashmead. Syn. of relat!vu9 punicus Cresson. 

perplexus Cresson. A subsp. of laborator (Thunberg). 

persimilis Cresson. 

Cryptus associatus Pratt (syn. teste Townes and Townes, 1962, p. 237). 

recurvatus Pratt. Calif., Tex. 

relativus nitscliei Dalla Torre, changed status. B. C, Wash., Calif., Alta., 
Mont., Nev., Utah, Ariz., Colo., N. Mex. Host: Aegeria tibialis 
(Harr.). Reduced to subsp. rank (teste Townes and Townes, 1962, 
pp. 193, 195). 

Cryptus violaceipennis Ashmead, preocc. 

Cryptus nitschei Dalla Torre, n. name. Removed from syn. of dirus Cres- 
son. 

Cryptus leucopus Ashmead (syn. teste Townes and Townes, 1962, p. 
195). 

Cryptus eleucopus Walkley (syn. teste Townes and Townes, 1962, p. 
195). 



ICHNEUMONIDAE 113 

relativus punicus Cresson, changed status. B. C. to Calif., Nev. Removed 
from syn. of pacifus Cresson and reduced to subsp. rank {teste 
Townes and Townes, 1962, pp. 193, 197). 
Cry plus calif ornicus Aslunead (syn. teste Townes and Townes, 1962, p. 
197). 

relativus relativus Cresson, changed status. Reduced to subsp. rank {teste 
Townes and Townes, 1962, p. 193). 

robustus Cresson. A subsp. of calescens Gravenhorst. 

rufopedibus Pratt. Syn. of tejonensis tejonensis Cresson. 

rufovinctus Pratt. A subsp. of albitarsis (Cresson). 

*rugifrons (Townes), n. comb. Ariz., Calif. 

Trachysphyrus rugifrons Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 184, 255. $, ?. 

rugosiscutum crassisculptus Pratt, changed status. Tex. Reduced to subsp. 
rank {teste Townes and Townes, 1962, pp. 226, 227). 

rugosiscutum flagellatus Pratt, changed status. Mexico. Reduced to subsp. 
rank {teste Townes and Townes, 1962, pp. 226, 227). 

rugosiscutum rugosiscutum Pratt, changed status. Reduced to subsp. rank 
{teste Townes and Townes, 1962, pp. 226, 228). 

rugulosus Pratt. Syn. of tejonensis mimicus Pratt. 

ruralis Pratt. B. C, Sask., Man., Idaho. 

*scapulatus (Townes) , n. comb. Calif. 

Trachysphyrus scapulatus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 184, 241. $, $. 

strigosus Pratt. Oreg. 

symmetricus bilobatus Pratt, changed status. Calif, in Canad. and Huds. 
Zones. Reduced to subsp. rank {teste Townes and Townes, 1962, 
pp. 204, 205). 

symmetricus symmetricus Pratt, changed status. Mont., Colo. Reduced to 
subsp. rank {teste Townes and Townes, 1962, p. 204). 

tejonensis mimicus Pratt, changed status. Alaska to Oreg. and Ariz., Alta., 
Utah, Colo., N. Mex. Reduced to subsp. rank {teste Townes and 
Townes, 1962, p. 190). 
Cryptus temporalis Pratt. (Syn. teste Townes and Townes, 1962, p. 190.) 
Cry plus rugulosus Pratt. (Syn. teste Townes and Townes, 1962, p. 190). 

tejonensis tejonensis Cresson, changed status. Calif., Ariz., Wash. Reduced 
to subsp. rank {teste Townes and Townes, 1962, pp. 190, 191). 
Cryptus vancouverensis Harrington. Remove from syn. A valid species. 
Cryptus rufopedibus Pratt. (Syn. teste Townes and Townes, 1962, p. 
191.) 

temporalis Pratt. Syn. of tejonensis mimicus Pratt. 

vancouverensis Harrington. B. C. to Calif. Host: Malacosoma sp. A valid 
sp. {teste Townes and Townes, 1962, pp. 181, 187). 

♦Genus LANUGO Townes 

Lanugo Townes, 1962. In Townes and Townes, U. S. Natl. Mus. Bui. 216 
(Pt. 3): 35,259,536. 

Type: Cryptus retentor Brulle. Orig. desig. (p. 536). 

Place on page 263 of Catalog after Cryptus Fabricius. 

*bicincta Townes. Kans. (Osborne). 

Lanugo bicincta Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3) : 261, 270. $. 



114 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*brunnipennis Townes. B. C. to Calif. 

Lanugo brunnipennis Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 260, 261, 275. 5, $. 

cestus (Say). Tenn. Transf. from Compsocrvplus Ashmead, p. 263 {teste 
Townes and Townes, 1962, pp. 261, 262). 

*deserii Townes. Calif., Ariz., Utah. Host: "Parasite of sawfly on Pinus." 

Lanugo deserti Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3): 260, 262, 268. $, 9. 

*excincta Townes. Kans., Colo., Utah, Ariz, N. Mex. 

Lanugo excincta Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3) : 261, 272. $, ?. 

*ferrugata Townes. Tex. 

Lanugo ferrugata Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 260, 262, 263. $, 9. 

*flavipennis Townes. Ariz. 

Lanugo flavipennis Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3): 260, 261, 271. $, 9. 

*longurius Townes. Ariz. (Sabino Canyon, Santa Catalina Mts.). 

Lanugo longurius Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 262, 269. 9. 

*picla Townes. Ariz., Tex.; Mexico. 

Lanugo picta Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3) : 260, 261, 266. $, 9. 

*poliia Townes. Ariz. 

Lanugo polka Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3) : 261, 271. 9. 

retentor (Brulle). Transf. from Compsocrrptus Ashmead, p. 264 (teste 
Townes and Townes, 1962, pp. 260, 261, 264). 

*6chlingeri Townes. Calif, (nr. Quincy). 

Lanugo schlingeri Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 261, 262. 9. 

sororia (Cresson). Calif., Ariz.; Mexico. Transf. from Compsocrvptus Ash- 
mead, p. 264 {teste Townes and Townes, 1962, pp. 261, 273). 

♦Genus REPTATRIX Townes 

Reptatrix To\vnes, 1962. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 3): 35,276. 

Type : Cryptus crassipes Pratt. Orig. desig. and by monotypy. 

Place in Cryptini on page 263 before Compsocryptus Ashmead. 

crassipes (Pratt). Wash, to Calif., Idaho, Utah, Wyo. Transf. from Cryptus 
Fabricius, p. 261 {teste Townes and Townes, 1962, p. 277). 

Genos COMPSOCRYPTUS Ashmead, pp. 263-264 

=^apicalis Townes. B. C, Oreg., Calif., Idaho, Nev., Utah, Colo., N. Mex. 

Compsocryptus apicalia Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 280, 282, 291. $, 9. 

*aridus Townes. Idaho, Wyo., Colo., south to Calif, and N. Mex. 

Compsocryptus ariilus Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 281, 282, 289. 5, 9. 

buccatus (Cresson). Mexico. 

calipterus brevicomis Cameron. Calif. Removed from syn. of calipterus calp- 

terus (Say). A valid subsp. {teste Townes and Townes, 1962, pp. 
286,287,288). 



ICHNEUMONIDAE 115 

calipterus calipterus (Say). Okla,, Tex, to Calif.; Mexico. 

*Cryptus fulvus Taschenberg, 1876. Ztschr. Gesam. Naturw. 48 (or N.F. 

14) : 6G. $. (Syn. teste Townes and Townes, 1962, p. 287.) 
*Ichneumon munerosus Cameron, 1885. Biol. Cent.-Amer. Ins., Hym. v. 

1, p. 159. $. (Syn. teste Townes and Townes, 1962, p. 287.) 
Campsocryptus (!) brevicornis Cameron, 1903. Remove from syn. A va- 
lid subsp. of calipterus (Say). 

cestus (Say). Transf. to Lanugo Townes. 

crotchii (Cresson). Calif. 

fasciipennis (Brulle). Fla. 

Taxonomy: Townes and Townes, 1962, pp. 279, 281, 282-283 (lectotype 
desig.). 

fletcheri (Provancher). B. C, Oreg., Calif., Idaho, Utah. 

*pallens Townes. Oreg., Calif., Idaho, Utah. 

Campsocryptus pollens Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 281, 282, 293. $, 9. 

purpuripennis (Cresson). B. C. 

resolutus resolutus (Cresson), changed status. Calif. Reduced to subsp. rank 
{teste Townes and Townes, 1962, pp. 303, 305). 

*resolutus subfasciatus Townes. Alta., south to Colo., west to Wash, and 
Calif. 
Campsocryptus resolutus subfasciatus Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 303. $, ?. 

retentor (Brulle). Transf. to Lanugo Townes. 

sororius (Cresson). Transf. to Lanugo Townes. 

*texensis Townes. Kans., Okla., Tex. 

Campsocryptus texensis Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 280, 281, 283. 5, $• 

*iricinctus Townes. Calif., ?Ga. 

Compsocrvptus tricinctus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 280, 281, 295. $, 9. 

turbatus (Cresson). Oreg., Idaho, Utah. 

*unicolor Townes. Calif. 

Compsocrvptus unicalor Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 280, 282, 297. $, $. 

Genus PYCNOCRYPTUS Thomson, p. 264 

*alexanderi Townes. Vt. (Lake Willoughby). 

Pycnocrrptus alexanderi Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 126, 128. e. 

director (Thunberg). Colo. 

*freemani Townes. N. B., Alta., Alaska. 

Pycnocryptus freemani Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 126, 127. $. 
*platyaspis Townes. Que., Man., Wyo. 

Pycnocryptus platyaspis Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 126, 127. 9. 

Genus MESOSTENUS Gravenhorst, pp. 264-265 
*Unlima Cameron, 1902. Ann. Mag. Nat. Hist., (Ser. 7) 9: 208. (Syn. 
teste Uchida, 1930, Hokkaido Imp. Univ. Fac. Agr. Jour. 25 (Pt. 
4): 317). 

Type: llnlima penetralis Cameron. By monotypy. 

albinotatus promptus Cresson, changed status. M. albinotatus albinotatus 
Gravenhorst is European. Reduced to subsp. rank (teste Tovraes 
and Townes, 1962, pp. 438, 439). 



116 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

angustus Townes. Carolinian fauna. 

=^clitellatus Townes. N. S., Que., to Va., west to Minn., Kans., and Tex., also 
Utah, B. C. Host: Pyrausta futilalis (Led.). 
Mesostenus clitellatus Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 436, 437, 447. $, ?. 

eisenii Ashmead. D. C, N. C, S. C, Ohio, Tex.; Mexico. 

gracilis Cresson. N. S., Que., Ont., B. C, Wash.; Baja California. 

*liogaster Townes. N. W. T., Sask., B. C, Oreg., Idaho, Wyo., Calif. ; Mexico, 
Mesostenus liogaster Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 435, 437, 440. $, ?. 

longicaudis Cresson. Transcont. in U. Austr. and L. Austr. Zones; Mexico. 

reliquus Cresson. Transf. to Chamula Townes. 

*sicarius Townes. N. C, Tex., Mont., Colo., Alta., Ariz., Calif.; Mexico. 

Mesostenus sicarius Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 436, 437, 448. $, ?. 

*temporalis Townes. Que., Maine, Ont., N. Y., Mich., Iowa, S. Dak. 

IMesostenus tarsatus Provancher, 1879. Transf. from syn. of Crypius 
albitarsis albitarsis (Cresson) {teste Towne sand Townes, 1962, p. 
441). 
Mesostenus temporalis Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 435, 436, 441. $, 9. 

ihoracicus Cresson. Transcont. Hosts: Acrobasis betulella Hulst; Macro- 
botys theseusalis (Wlkr.), Mineola sp. 

Genus POLYCYRTUS Spinola, p. 265 

*Cryptanuridimorpha Viereck. 1913. U. S. Natl. Mus. Proc. 46: 369. 

Type: Cryptanuridimorpha elegans Viereck. Orig. desig. 
*Cryptopterigimorpha Viereck, 1913. U. S. Natl. Mus. Proc. 46: 371. 

Type: Cryptopteriginiorpha tubulifera Viereck. Orig. desig. 
*Polycyrtimorpha Viereck, 1913. U. S. Natl. Mus. Proc. 46: 383. 

Type : Polycyrtimorpha amoena Viereck. Orig. desig. 
Taxonomy: Townes and Townes, 1962, pp. 14, 459. 

neglectus Cushman. Ohio. 

Polycyrtus subgenus Polycyrtus Spinola, p. 265, Sup. 1, p. 46 

Delete from catalog and Supplement 1. 

Genus CRYPTANURA Brulle, p. 265 

banchifomiis (Megerle). N. Y". to Fla., west to Mich, and Tex. Transf. from 
Unplaced Species, p. 408 {teste Townes and Townes, 1962, p. 427). 
Mesostenus spinarius Brulle (syn. by Townes and Townes, 1962, p. 427). 
Biology: Townes and Townes, 1962, pp. 428-429. 

*compacta (Cresson). Tex.; Mexico, Honduras. 

Mesostenus (Mesostenus) compactus Cresson, 1873. [Phila.] Acad. Nat. 

Sci. Proc. 1873: 153. ?. 
*Cryptanura planiscutellata Cushman, 1945. U. S. Natl. Mus. Proc. 96: 
143, 152, 175. ?. (Syn. teste Townes, 1946, Bol. Ent. Venezolana 
5 (2): 35.) 

septentrionalis Cushman. N. Y., Pa., D. C, Va., N. C, S. C, Tenn., La. 

spinaria Brulle. Syn. of banchiformis (Megerle). 

♦Genus CHAMULA Townes 

Chamula Townes, 1962. In Townes and Townes, U. S. Natl. Mus. Bui. 
216 (Pt. 3) : 36, 430. 

Type : Mesostenus (Polycyrtus) reliquus Cresson. Orig. desig. 

Place in Cryptini after Cryptanura Brulle, p. 265. 

*reliqua (Cresson). Transf. from Mesostenus Gravenhorst {teste Townes and 
Townes, 1962, p. 431). 



ICHNEUMONIDAE 117 

♦Genus CAMERA Townes 

Camera Townes, 1962. In Townes and Townes, U. S. Natl. Mus. Bui. 216 
(Pt. 3) : 36, 432. 

Type : Mesostenus euryaspis Cameron. Orig. desig. 

Place in Cryptini on page 265 before Hidryta Foerster. 

*euryaspis (Cameron). Tex. 

Mesostenus euryaspis Cameron, 1885. Biol. Cent.-Amer., Ins., Hym. v. 1, 
p. 226. $. 

Genus HiDRYTA Foerster, pp. 265-266 

Hidryta Foerster. Change type citation to {Brachycryptus erythrocerus 

Thomson) = f rater (Cresson). 
Brachycryptus Thomson. Change type citation to {Brachycryptus eryth- 

rocerus Thomson) = fraier (Cresson). 
*Euthycryptus Jussei, 1907. Festsclir. zum 50 jahr. Bestande zugleich 44. 
Jahresb. iiber das Jahr 1906, Vorarlberger Landesmuseum 
Bregenz, p. 71 (syn. teste Townes and Townes, 1962, p. 318). 
Type: Euthycryptus scrobiculifer Jussei. By monotypy. 

f rater (Cresson). Transcont. in Canad. and Transit. Zones; Eurasia. 

* Brachycryptus erythrocerus Thomson, 1873. Opusc. Ent. fasc. 5, p. 488. 
2. (Syn. teste Townes and Townes, 1962, p. 318, 319.) 

nigrlcoxus (Provancher) . N. S., Ont., N. J., Pa., N. C, Mich., Minn. 

Genus TRYCHOSIS Foerster, p. 266 

Trychosis Foerster. Change type citation to: (Cryptus titillator Graven- 
h.ovs,t) z=Trychosis ambigua (Tschek). Desig. by Schmiedeknecht, 
1890 {teste Townes and Townes, 1962, p. 333; Perkins, 1962, pp. 
460-461). 

albicaligata (Walsh). Conn., N. Y., Md., D. C, Va., Ind., Kans., Tex. 

albisoleatus (Walsh). Transf. to Idiolispa Foerster. 

albitarsis albitarsis (Cresson), changed status. Newfoundland, Que., Ont., 
Maine, N. H., Vt., Mass., N. Y., Md., Va., Mich., Kans. Reduced 
to subsp. rank {teste Townes and Townes, 1962, p. 359). 

*albitarsis peratra Townes. B. C. to Calif., Idaho. 

Trychosis albitarsis peratra Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 358, 359. $, ?. 

*anagnius Townes. Transcont. in Transit, and Canad. Zones. 

Trychosis anagmus Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 335, 337, 376. S, ?• 

*apicalis Townes. E. I., N. Y., Pa., Md., Va., Fla., Tenn., Ohio, Mich., 111., 
Minn, 
Trychosis apicalis Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 336, 338, 347. $, ?. 

*atrorubens Townes. Colo., Ariz., Nev., Calif. 

Trychosis atrorubens Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3): 336, 337, 369. $, $. 

*coxaIis Townes. R. I. and N. Y. to S. C, west to Mich., Ala. 

Trychosis coxalis Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3) : 335, 337, 346. S, $. 

*cyperia Townes. Transcont. in Canad. and Huds. Zones. 

Trychosis cyperia Townes, 19S2. In Townes and Townes, U. S .Natl. 
Mu.s. Bui. 216 (Pt. 3) : 335, 336, 373. $, 9. 

*depilis Townes. B. C. to Calif., Idaho, Wyo., Utah. 

Trychosis depilis Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3) : 336, 338, 368. $, $. 

exulans (Cresson). Ont., Mass. to N, C, La., west to Mich., Kans., Tex. 



118 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*fuscata Townes. Labrador, Que., Maine. 

Trychosis fuscata Townes, 1962. In Townes and Townes, U. S. Natl, 
Mus. Bui. 216 (Pt. 3): 336, 337, 356. $, ?. 

fusifortnis (Provancher). Syn. of montivaga montivaga (Provancher). 

*kathrynae Townes. Que. to Sask., south to N. J., W. Va., Mich., Wis., Minn. 
Trychosis kathrynae Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 335, 337, 363. $, 9. 

'^'latidens Townes. Que. and N. Y. to Yukon, B. C, Wash., Oreg. ; also Colo. 

Trychosis latidens Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 336, 338, 355. $, 9. 

*montivaga austrina Townes. N. Mex. (Tajique). 

Trychosis montivaga austrina Townes, 1962. U. S. Natl. Mus. Bul. 216 
(Pt. 3): 336,338,341. $, 9. 

montivaga montivaga (Provancher). changed status. Transcont. in north 
U. S. and south. Canad. Reduced to subsp. rank (teste Townes and 
Townes, 1962, pp. 335, 338, 339). 
Trychosis fusijformis (Provancher) (syn. by Townes and Townes, 1962, p. 
339). 

*nlgripe8 Townes. Tex. 

Trychosis nigripes Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bul. 216 (Pt. 3) : 336, 338, 345. 5, 9. 

*reflexa Townes. B. C. to Calif., Idaho, Nev., Mont., Utah, S. Dak. 

Trychosis reflexa Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bul. 216 (Pt. 3) : 336, 337, 371. $, ?. 

*semirubra arizonica Townes. Ariz. (nr. Alpine), 

Trychosis semiruhra arizonica Townes, 1962. U. S. Natl. Mus. Bul. 216 
(Pt. 3): 351, 354. 3, 9. 

*semirubra pulla Townes. Wyo., Colo., Alta., Idaho, B. C, Wash., Calif. 
Host: egg sac drassid spider. 
Trychosis semirubra pulla Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bul. 216 (Pt. 3) : 351, 352. $, 9. 

semirubra semirubra Townes, changed status. Atlantic Ocean to Rocky Mts. 
in Transit, and Canad. Zones, Tex. Host: drassid spider egg 
cases. Reduced to subsp. rank (teste Townes and Townes, 1962, 
pp. 351, 353). 

^^slmilis badiarmus Townes. N. S. to Ont. south to Va., Ohio, Minn., also Tex. 
Trychosis similis badiarmus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bul. 216 (Pt. 3) : 342, 343. $, 9. 

similis similis (Cresson), changed status, N, H, to S. C. west to Wis., also 
Kans., Tex. Reduced to subsp. rank (teste Townes and Townes, 
1962, pp. 342,344). 

subgracilis (Cresson). Transit, zone. 

^sulcata Townes. Conn., N. Y. to N. C, west to Ont., Mo., Kans. 

Trychosis sulcata Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bul. 216 (Pt. 3) : 335, 337, 349. $,9. 

Genus IDIOLISPA Foerster, p. 266 

*Paracryptu8 Szepligeti, 1916. Mus. Nat. Hungarici, Ann. 14: 251. (Syn. 
teste Townes, Townes and Gupta, 1961, p. 166.) 

Type : Paracryptus hungaricus Szepligeti. By monotypy. 

^aestivalis Townes. Maine and Que. to Ga., west to Wis., Iowa, Kans., also 
Tex. 
Idiolispa aestivalis Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bul. 216 (Pt. 3) : 323, 331. $, 9. 

albisoleata (Walsh). Transcont. in Canad. and Transit. Zones. Transf. from 
Trychosis Foerster, p. 266 (teste Townes and Townes, 1962, p. 
323). 



ICHNEUMONIDAE 119 

analis analis (Gravenhorst), changed status. Transcont. in Canad. Zone; Eu- 
rope. Reduced to subsp. rank (teste Townes and Townes, 1962, 
pp.326). 

♦analis ignea Townes. So. Sask. to B. C, south to Calif., Ariz., N. Mex. 

Idiolispa analis ignea Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3): 326, 329. $, 9. 

analis limata (Cresson), changed status. Transcont. in Transit, and U. 
Austr. Zones. Reduced to subsp. rank (teste Townes and Townes, 
1962, pp. 326, 328). 

limata (Cresson). A subsp. of analis (Gravenhorst). 

*Genus HEMISPHRAGIA Seyrig 

*Micromavia Seyrig, 1952. Acad. Sci. Malgache, Mem. 19: 203, 204. 
(Syn. teste Townes and Townes, 1962, p. 378.) 

Type: Micromavia carinata Seyrig. Orig. desig. 
Hemisphragia Seyrig, 1952. Acad. Sci. Malgache Mem. 19: 204, 207. 

Type: Hemisphragia triangnlifera Seyrig. Orig. desig. 

Place in Cryptini on page 266 after Idiolispa Foerster. 

*debills Townes. Md., Va., Tenn., Ohio, Kans., La. 

Hemisphragia debilis Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 378. $, ?. 

Genus APSILOPS Foerster, p. 267 

*Heterotypus Foerster, 1868. Naturh. Ver. Rheinlande 25: 183. Syn. 
(teste Townes, Townes, and Gupta, 1961, p. 142). 

Type: Ichneumon cinctorius Fabricius. Desig. by Townes, 
Townes, and Gupta, 1961. 

bicolor (Cushman). Transcont. in Canad. Zone. 

Genus GAMBRUS Foerster, pp. 267-268 

Aritranis Foerster. Remove from syn. and place as valid genus on p. 268 
with Hoplocryptus Thomson as syn. 

ajfahilis (Provancher). Transf. to Aritranis, p. 269. 

apicatus (Provancher). N. S., Ont., Vt., Ohio, Mich. Hosts: lEuura gall; 

gall on Salix. 

bituminosus (Cushman). Calif. 

canadensis burkei (Viereck), changed status. Calif. Hosts: Malacosoma cali- 
fornicum (Pack.) ; H emerocampa vetusta (Bdvl.) ; H. oslari 
(Barnes). Removed from syn. of exareolatus (Viereck) (teste 
Townes and Townes, 1962, pp. 80, 84). 

canadensis canadensis (Provancher), changed status. Atlantic Ocean to 
Rocky Mts. in Canad. and Transit. Zones; Alta., B. C, Wash., 
Oreg. Hosts: Malacosoma sp. ; M. disstria Hbn., M. americaniim 
(F.) ; Cnrpocapsa pornonella (L.) ; tenthredinid. Reduced to 
subsp. rank (teste Townes and Townes, 1962, p. 80). 
ICryptus belangeri Provancher. Transf. to Aritranis Foerster as syn. of 
notata notata (Provancher). 

canadensis exareolatus (Viereck), changed status. Minn., Sask., Mont., Wyo., 
Colo., N. Mex., Alta., Idaho, also Wash. Hosts: Malacosoma sp., 
M. fragile (Stretch). Reduced to subsp. rank (teste Townes and 
Townes, 1962, p. 83). 

exareolatus (Viereck). A subsp. of canadensis (Provancher). In bibliog. ref. 
change p. 80 to 81. 
Cryptus (Gambrus) burkei Viereck. Remove from syn. Valid subsp. of 
canadensis (Provancher). 

extrematis (Cresson). Host: Hyalophora euryalus kasloensis (Ckll.). 



120 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

nuncius (Say). N. W, T., Mich., Minn. Hosts: Callosamia sp., C. angidifera 
(Wlkr.), ISamia cynthia (Dru.), lAntheraea polyphemus 
(Cramer), 

tunicula rubra (Fyles). 

Taxonomy: Townes and Townes, 1962, pp. 71, 75. (Lectotype desig.) 

ultimus (Cresson). Exyra semicrocea (Gn.), Hypera brunneipennis (Boh.), 
Hyalscotes pithopoera (Dyar), Loxostege sticticalis (L.), Symyra 
henrici (Grt.), Trichoplusia ni (Hbn.), Udea rubigalis (Gn.). 

*yukonensis Townes. Alaska, Yukon, N.W.T. 

Gambrus yukonensis Townes, 1962. In Townes and Townes, U.S. Natl. 
Mus. Bui. 216 (Pt. 3) : 71, 79. $, 9. 

Genus ARITRANIS Foerster 

( = Hop/ocrypfys Thomson of Catalog, pp. 268—269) 

Aritranis Foerster. Removed from syn. of Gambrus Foerster. A valid 

genus {teste Townes and Townes, 1962, pp. 70, 98). 
Hoplocryptus Thomson. Emend type citation to read {Hoplocryptus bin- 
otatulus Thomson) = *fugitiva (Gravenhorst) . 

affabilis (Provancher). N. C, Ind., Mich., Ont., Minn., Kans. Hosts: Phlyc- 
taenia tertialis (Guen.), Oberea tripunctata (Swed.), Oberea sp. 
Transf. from Gambrus Foerster {teste Townes and Townes, 1962, 
p. 100). 

albicoUaris albicollaris (Cresson), changed status. Que., Vt., N. Y., Va., N. C, 
Mich., Man., 111., Kans., Tex., Mont. Host: Phlyctaenia ter- 
tialis (Guen.). Reduced to subsp. rank {teste Townes and Townes, 
1962, pp. 100, 112). 

*albicollaris clypeata Townes. Calif., Wash. 

Aritanis albicoUaris clypeata Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 112, 114. $, 9. 

*barberi Townes. Md. (Plummei'S Is.). 

Aritranis barberi Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3) : 100, 111. $. 

*byrsina Townes. Que., Ont., south to Ga. and La., west to Wis., Iowa, Mo., 
Kans. Hosts: Achatodes zene (Harr.), Oberea sp., Papaipema sp. 
Aritranis byrsina Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3): 100, 102. $, 9. 

gracilis (Provancher). Alleghanian. 

imitator imitator (Provancher), changed status. N. B., Que., Ont. south to 
R. I., N. Y., Ohio, west to Minn., also Alta. Reduced to subsp. rank 
{teste Townes and Townes, 1962, pp. 99, 121, 123). 

*imitator monticola Townes. Alta., B. C, Idaho, Wash., Oreg. 

Aritranis imitator monticola Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 121, 124. $, ?. 

*imitator nigriventris Townes. Que., N. Y., Sask., Wyo., Colo., N. W. T., 
Yukon, B. C, Oreg. 
Aritranis imitator nigriventris Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 121. $, 9. 

*imifator ruficauda Townes .Calif. Host: Hoplitis hypocrita (Ckll.). 

Arcitransi imitator monticola Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 121, 125. $, 9. 

*iinnae linnae Townes. Que. to Alta., also N. Y., Mich., Minn. 

Aritranis linnae linnae Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 100, 105. $, 9. 

*linnae paoifica Townes. Calif. 

Aritranis linnae pacifica To^vnes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 105, 106. $, 9. 



ICHNEUMONIDAE 121 

notata newcomeri (Cushman), changed status. Mont, to B. C. south to Oreg., 
also N. J. Host: Ainetastegia glabroAa (Fall.), Allantus cinctus 
(L.). Removed from syn. of notata (Provancher) {teste Townes 
and Townes, 1962, pp. 117, 119). 

notata notata (Provancher), changed status. Atlantic to Rocky Mts. in Can- 
ad., Transit, zones. Hosts: "Odynerus" nest in Rhus stem, Ancis- 
troccrns brrenimaculatus (Sau?s.). 

Cryptus belangeri Provancher. Transf. from syn. of Gambrus canadensis 
canadensis (Provancher), p. 268 {teste Townes and Townes. 1962, 
pp. 80, 116). 

Caenocrrptus newcomeri Cushman. Remove from syn.; valid subsp. of 
notata (Provancher). 

*notata sierrae Townes. Calif. Hosts: Ancistrocerus spilogaster Cam.; Lep- 
tochilus tetradymia Park. ; Megachile gentilis Cress. 
Aritranis notata sierrae Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 116, 120. $,9. 

♦nubecula Townes. R. I., Fla., Mich., Tex. 

Aritranis nubecula Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 100, 109. S, ?. 

Genus AGROTHEREUTES Foerster, pp. 269-270, Sup. 1, p. 46 

*Dayro Cameron, 1902. Ann. and Mag. Nat. Hist. (Ser. 7) 9: 209. Syn. 
{te^te Townes and Townes, 1962, pp. 12, 42. An earlier reference 
to this synonomy not found). 

Type : Dayro pilosus Cameron. By monotypy. 

abbreviator iridescens (Cresson), changed status. A. abbreviator abbreviator 
is European. Minn., Wis., Ont. Host: Diprion frutetorum (F.). 
Reduced to subsp. rank {teste Townes and Townes, 1962, pp. 51, 
52,57). 

abbreviator rufopectus Cushman, changed status. B. C, Alta. to Calif, east 
to Wyo., Colo., N. Mex. Hosts: Neodiprion tsugne Midd., Pristi- 
phora erichsonii {Htg.), Laspeyresia vigricana (Steph.). Reduced 
to subsp. rank {teste Townes and Townes, 1962, pp. 51, 55). 

abbreviator similaris (Provancher), changed status. N. S., Ont., Mich., Wis., 
Minn., Man., Sask., Alta., N. W. T., Yukon, Alaska. Host: Neodi- 
prion lecontei (Fitch), N. abietis (Harr.), N. sertifer (Geoff.), 
Pristiphora erichsonii (Htg.). Reduced to subsp. rank {teste 
Townes and Townes, 1962, pp. 51, 52, 53). 
Biology: Finlayson, 1960, Canad. Ent. 92 (12) : 932-933 (larval). 

*alutarius Townes. Alaska, Yukon, N. W. T. 

Agrothereutes alutarius Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 44, 45, 47. $, 9. 

cimbicivorus (Cushman). Canad. Zone. 

*grandis Townes. Conn, south to S. C. 

Agrothereutes grandis Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 44, 45, 46. $, ?. 

iridescens (Cresson). A subsp. of abbreviator (Fabricius). 

lophyri lophyri (Norton), changed status. Reduced to subsp. rank {teste 
Townes and Townes, 1962, pp. 60, 62). 

*lophyri obscurus Townes. Newfoundland, N. B. Host: Neodiprion abietis 
(Harr.). 
Agrothereutes lophyri obscurus Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 60, 61. $, 9. 

*lophyri ontariensis Townes. N. B., Maine, Que., Ont., N. H., N. J., Mich., 
Wis., Minn., Man., B. C. Hosts: Neodiprion abietis (Harr.) ; N. 
lecontei (Fitch); N. nanulus nanulus Schedl; N. nigroscutum 
Midd.; N. pratti banksianae Rohw.; N. sp., Diprion similis 
(Htg.) ; "Spruce sawfly." 



122 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Agrothereutes lophyri ontariensis Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 60. S, 9. 

*mandator americanus Townes. Transcont. in Transit, and Canad. Zones. 
Hosts: Cimbex americana Leach, Trichiosoma triangulum Kby., 
T. sp. 
Agrothereutes mandator americanus Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 67, 68. $, 9. 

^montanus Townes. Colo. (Rocky Mt. Natl. Pk.). 

Agrothereutes montanus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 45, 48. 9. 

neodiprionis (Cushman). Canad. Zone. 

'^pallipennis Townes. Calif. (East Sonora Pass). 

Agrothereutes pallipennis Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3): 43, 45, 49. $, 9. 

rufopectus Cushman. A subsp. of abbreviator (Fabricius). 

similaris (Provancher). A subsp. of abbreviator (Fabricius). 

Genus JOPPIDIUM Cresson, p. 270 

*Isocryptu8 Brethes, 1927. Ent. Mitt. 16: 308. (Syn. teste Townes and 
Townes, 1962, p. 306.) 

Type: (Isocryptus azureus Brethes) = moerens (Perty). By 
monotypy. 

*arizonicuin Townes. Ariz. 

Joppidium arizonicum Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 307, 312. $. 

*brochum brochum Townes. Okla., Tex.; Mexico. 

Joppidium brochum brochum Townes, 1962. In To\vnes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 307, 308, 309. $, 9. 

*brocbum fattigi Townes. Ky., N. C, Ga. 

Joppidium brochum fattigi Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 308, 309. $, 9. 

*densum Townes. Ariz. ; Mexico. 

Joppidium densum Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 307, 312. $, 9. 

*discolor Townes. Ariz. 

Joppidium discolor Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 308, 314. $. 

*dubiosum Cresson. Va., S. C, Ga., Ala.; Mexico. 

Joppidium dubiosum Cresson, 1873. [Phila.] Acad. Nat. Sci. Proc. 1873: 

138. $, 9. 
*Opisosextu8 (!) ruficeps Sz^pligeti, 1916. Mus. Natl. Hungarici, Ann. 
14: 240. $. Preocc. (Syn. teste Townes and Townes, 1962, p. 317.) 

*fu9cipenne (Brulle). Ariz. (Huachuca Mts.) ; Mexico. 

Cryptus fuscipennis Brulle; 1846. In Lepeletier, Hist. Nat. Ins. Hym. v. 

4, p. 189. 9. 
*Joppidium donabilis Cresson, 1873. [Phila.] Acad. Nat. Sci. Proc. 

1873: 139. 5, 9. (Syn. teste Townes and Townes, 1962, p. 310.) 

* Joppidium yucatanense Cameron, 1885. Biol. Cent.-Amer. Ins., Hym., v. 1, 
p. 211. 9. (Syn. teste Townes and Townes, 1962, p. 310.) 

Genus ISCHNUS Gravenhorst, pp. 270-271 

*Aglaocryptus Cameron, 1903. Manchester [England] Lit. and Philos. 
Soc, Mem. and Proc. 47 (Pt. 14) : 31. (Syn. teste Townes, Townes 
and Gupta, 1961, pp. 133, 135.) 

*Type: Aglaocrrptug curvimaculatus Cameron. Desig. by 
Viereck, 1914. 



ICHNEUMONIDAE 123 

cinctipes (Walsh). Que., Ont., to Fla., west to Minn, and Tex. Hosts: An- 
cylis comptana (Froel.), A. divisana Wlkr., Polychrosis viteana 
(Clem.). Transf. from Alphabetic List of Unplaced Species, p. 
408 (teste Townes and Townes, 1962, pp. 133, 137). 
Ischnus fraterculus fraterculus (Provancher) Sup. 1, p. 46 (as 3-annu- 
latus fraterculus (Provancher) in Catalog, p. 270. (Syn. teste 
Townes and Townes, 1962, p. 137). 

*inquisItorius assimilis (Uchida), changed status. Alaska; Siberia, Japan. 
Reduced to subsp. rank (teste Townes and Townes, 1962, pp. 144, 
145, 146). 
Habrocrrptus assimilis Uchida, 1930. Hokkaido Imp. Univ., Fac. of Agr. 
Jour. 25: 313. 9. 

inquisitorius atriceps (Cresson). Colo., S. Dak. Hosts: ChoHstoneura fumi- 
ferana (Clem.) ; orange tortrix on Rliododendron sp. 

inquisitorius atricollaris (Walsh). B. C. Hosts: Choristoneura rosaceana 
(Harr.), Malacosoma disstria Hbn. 
Cry plus amblytelarius Provancher. Transf. to syn. of rhomboidalis 

(Walsh). 
Spilocryptus neomexicana Viereck. In bibliog. ref. change p. 80 to p. 81. 

^^inquisitorius pectoralis Townes. Alaska, Yukon, N. W. T., B. C. to Colo., 
N. Mex. east to Ont., Mich. Host: Choristoneura conflictana 
(Wlkr.). 
Ischnus inquisitorius pectoralis Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 144, 146. $, 9. 

*laevifrons Townes. Ariz. (Transit., Canad. Zones). 

Ischnus laevifrons Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 133, 156. $, ?. 

latus latus (Provancher), changed status. N. S., Maine, N. H., west to Alta. 

in Canad. Zone. Transf. from syn. of Cubocephalus annulatus 

Cresson, p. 253 and reduced to subsp. rank (teste Townes and 

Townes, 1962, p. 135). 
Ischnus triannulatus triannulatus (Provancher). Preocc. (Syn. teste 

Townes and Townes, 1962, p. 135.) 
Cryptus triannulatus Provancher (syn. teste Townes and Townes, 1962, 

p. 135). 
Ischnus fraterculus tricircularis Walkley (syn. teste Townes and Townes, 

1962, p. 135). 

latus pinguis (Provancher), changed status. Que., Ont. to N. C. west to Wis. 
and Kans. in Alleghanian fauna. Reduced to subsp. rank (teste 
Townes and Townes, 1962, pp. 135, 136). 

*laurae Townes. S. C, Fla. 

Ischnus laurae Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3): 133, 140. $, 9. 

""lautus Townes. B. C, Calif. Host: Grapholitha molesta (Busck). 

Ischnus lautus Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3): 133, 163. $, 9. 

*minor Townes. Ti'anscont. in Transit, and Canad. Zones. Hosts: Acleris 
variana (Fern.), Choristoneura fumiferana (Clem.), Polychrosis 
viteana (Clem.). 
Ischnus minor Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3): 133,159. $, ?. 

*politus Townes. Colo. 

Ischnus poliius Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3) : 133, 162. 9. 

rhomboidalis (Walsh). Transf. from Alphabetic List of Unplaced Species, p. 
408 (teste Townes and Townes, 1962, p. 157). 
Cryptus amblytelarius Provancher. Transf. from syn. of inquisitorius 
atricoUaris (Walsh) (teste Townes and Townes, 1962, p. 157). 



124 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*sparsus Townes. Que. to N. C. west to Minn. 

Ischnus sparsus Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3): 132, 141. $, ?. 

*velutinus Townes. Ariz. (Transit. Zone). 

Ischnus velutinus Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 133, 155. $, 9. 

Genus PSEUDISCHNUS Walkley, Sup. 1, p. 47 

Taxonomy: Townes and Townes, 1962, pp. 165-167. 

*oregonen8is coxalis (Townes) n. comb. S. Dak. (Deadwood). 

Habrocryptoides oregonensis coxalis Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 167. ?. 

oregonensis oregonensis (Cushman), changed status. B. C, Wash., Calif., 
Wyo. Reduced to subsp. rank (teste Townes and Townes, 1962, p. 
167). 

♦Genus HABROCRYPTOIDES Uchida 

Habrocryptoides Uchida, 1952. Insecta Matsumurana 18 : 19. 

Type : Habrocryptus shikokuensis Uchida. Orig. desig. 

Place after Pseudischnus [p. 271], Sup. 1, p. 47. 

*erasus Townes. Alta. (Lethbridge). 

Habrocryptoides erasus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 166, 175. ?. 

*inflexus Townes. Calif. (Alpine Co.). 

Habrocryptoides inflexus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (3) : 166, 177. 9. 

*Iunibarius Townes. Calif. (Lancaster). 

Habrocryptoides lumbar ius Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 166, 174. 9. 

*nitens Townes. Calif. (Kern Co.). 

Habrocryptoides nitens Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 166, 176. 9. 

*pictus Townes. Ariz. 

Habrocryptoides pictus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 166, 173. $. 

rufifrons (Walsh). N. Y. to Fla., west to Mich, and Tex. Transf. from 
Alphabetic List of Unplaced Species, p. 408 (teste Townes and 
Townes, 1962, pp. 166, 169). 

*virgeus ardalus Townes. Ariz., Colo. 

Habrocryptoides virgeus ardalus Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 171, 172. $, 9. 

*virgeus virgeus Townes. Mass. to S. C, Mich., Wis. Hosts: Harrisina amer- 
icana (G.-M.) ; ICarpocapsa pomonella (L.). 
Habrocryptoides virgeus virgeus Townes, 1962. In Townes and Townes 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 166, 170, 171. $, 9. 

Genus CHROMOCRYPTUS Ashmead, p. 271 

planosae planosae (Fitch), changed status. Reduced to subsp. rank (teste 
Townes and Townes, 1962, p. 257). 

*pIanosae vandykei Townes. Calif. ("Havilah", Fresno Co.). 

Chromocryptus planosae vandykei Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3): 257, 258. 9. 

Genus LISTROGNATHUS Tschek, p. 271 
♦LISTROGNATHUS subgenus FENESTULA Townes 

Listrognathus subg. Fenestula Townes, 1962. In Townes and Townes, 



ICHNEUMONIDAE 125 

U. S. Natl. Mus. Bui. 216 (Pt. 3) : 408. 

Type: Mesostenus paludatus Cresson. Orig. desig. and by 
monotypy. 

^paludatus californicus To\vnes. Calif. 

Listrognathus (Fenestula) paludata californica Townes, 1962. In Townes 
and Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 409, 410. $,9. 

*paludatus ocularis Townes. Ariz. (Chiricahua Mts.). 

Listrognathus (Fenestula) paludata ocularis Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 409, 411. ?. 

paludatus paludatus (Cresson), changed status. Transcont. in Transit, and 
Canad. Zones, Alaska, Tex. Hosts: Cerura cinerea Wlkr., C. sp. 
Reduced to subsp. rank {teste Townes and Townes, 1962, p. 409). 

LISTROGNATHUS subgenus LISTROGNATHUS Tschek 

*acheloma Townes. Md., Fla. 

Listrognathus (Listrognathus) acheloma Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 413, 425. 9. 

albomaculatus albomaculatus (Cresson), changed status. Ont., Ga., Mich., 
Kans., La., Tex. Host: Gretchena bolliana (Sling.). Reduced to 
subsp. rank (teste Townes and Townes, 1962, pp. 420, 421). 

albomaculatus var. multimaculatus Cushman. A valid sp. 

albomaculatus var. nubilipennis (Cresson). A valid species. 

albomaculatus sagax(Provancher) , changed status. 

Listrognathus albomaculatus var. «aga;«: ( Provancher) . Changed to subsp. 
rank {teste Townes and Townes, 1962, p. 420). 

*bicolor Townes. Ariz. (Oak Creek Canyon). 

Listrognathus (Listrognathus) bicolor Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 414. 9. 

*femoratus Townes. P. E. I., Que., Ont., Mich., Va., W. Va 

Listrognathus (Listrognathus) femorata Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 414, 415. $, 9. 

*glomeratus Townes. Pa. to Fla., W. Va., Ala., Tex. 

Listrognathus (Listrognathus) glomerala Townes, 1962. In Townes and 
Townes, U. S .Natl. Mus. Bui. 216 (Pt. 3) : 414, 424. $, 9. 

multimaculatus Cushman, changed status. 

Listrognathus albomaculatus var. multimaculatus Cushman. Elevated to 
sp. rank (teste Townes and Townes, 1962, pp. 414, 422). 

*nigrescens Townes. Alta., Utah, Wyo., N. Mex. 

Listrognathus (Listrognathus) nigrescens Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 414, 416. $, 9. 

nubilipennis (Cresson), changed status. 

Listrognathus albomaculatus var. nubilipennis (Cresson). Elevated to 
sp. rank (teste Townes and Townes, 1962, pp. 414, 416). 

paludatus (Cresson). Transf. to subg. Fenestula Townes. 

rufitibialis Cushman. Ind., Okla. 

*rufus Townes. N. Y., Kans. 

Listrognathus (Listrognathus) rufa Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 413, 419. $. 

victoriensis (Harrington). 

Taxonomy: Townes and Townes, 1962, pp. 417-419. 

Genus DIAPETIMORPHA Viereck, pp. 271-272 

acadia Cushman. D. C, Va., N. C, Tenn., Ala. 

alabama Cushman. 

Biology and Taxonomy: Townes and Townes, 1962, pp. 381, 383 384. 



126 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*brunnea Townes. Ga., Fla. 

Diapetimorpha brunnea Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 381, 382. $, 9. 

confederata Cushman. A subsp. of macula (Cameron). 

introita (Cresson). N. C, S. C, Fla., Miss. 

macula confederata Cushman, changed status. Va. Reduced to subsp. rank 
(teste Townes and Townes, 1962, p. 385). D. macula macula 
(Cameron) is Mexican. 

*macula ustulata Townes. Fla. 

Diapetimorpha macula ustulata Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 385, 386. 9. 

*picta Townes. Ga., Fla, 

Diapetimorpha picta Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 381, 387. $, 9. 

rufigaster Cushman. N. C, Tenn., Wis. 

*rugosa Townes. N. J. to S. C. west to Wis., Iowa, Kans., Ark., La. and Tex. 
Diapetimorpha rugosa Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 381, 382, 392. $,9. 

Genus MALLOCHIA Viereck, p. 272 

agenioides Viereck. Mich. 

♦frontalis Townes. N. J,, Md., N. C, Kans., Tex. 

Mallochia frontalis Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (3) : 93, 94, 97. $, ?. 

*laevis Townes. Ga., Fla. 

Mallochia laevis Townes, 1962. In Townes and Townes, U. S. Natl. Mus. 
Bui. 216 (Pt. 3) : 94,97. $, 9. 

strigosa (Cresson). ?Ariz. 

Taxonomy: Townes and Townes, 1962, pp. 94, 95. 

*Genus PACHYSOMOIDES Strand 
(=Pofist/phaga Cushman of Catalog, p. 272) 

*Pachysoma Szepligetl, 1916. Mus. Natl. Hungarici Ann. 14: 263, 290. 
Preocc. 

Type: (Pachysoma albopictum Szepligeti) = stupidus (Cres- 
son). By monotypy. 
Pachrsomoides Strand, 1917. Ent. Ztschr. 10 (24) : 137. N. name. 
Polis'tiphaga Cushman. Syn. {teste Townes and Townes, 1962, p. 398). 
Change type citation to read: {Mesostenus arvalis Cresson) = 
fulvus (Cresson). 

fulvus (Cresson). Cuba. 

stupidus (Cresson). Tenn. 

"Pachysoma albopictum Szepligeti, 1916. Mus. Natl. Hungarici Ann. 14.: 
290. 9. Syn. (fesfe Townes and Townes, 1962, p. 399). 

Genus Nasufocr/pfus Pratt, p. 272 

Syn. of Lymeon Foerster. 

nasutus Pratt. Transf. to Lymeon Foerster, infra. 

Genus LYMEON Foerster, pp. 272-273 

*Christolimorpha Viereck, 1913. U. S. Natl. Mus. Proc. 44: 564. (Syn. 
teste Townes and Townes, 1962, p. 393.) 

Type: Christolimorpha plesius Viereck. Orig. desig. 
*Zamastrus Viereck, 1913. U. S. Natl. Mus. Proc. 46: 385. (Syn. teste 
Townes and Townes, 1962, p. 393.) 

Type : Zamastrua photopsis Viereck. Orig. desig. 
*Neogoryphu8 Roman, 1936. Ent. Tidskr. 57: 3. (Syn. teste Townes and 



ICHNEUMONIDAE 127 

Townes, 1962, p. 393.) 

*Type: Ichneumon ariolator Linnaeus. 
Nasutocryptus Pratt ( syn. teste Townes and Townes, Op. cit. supra) . 

cinctiventris (Cushman). D. C. 

*leiponeuron Townes. Fla. 

Lymeon leiponeuron Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 393, 397. $,9. 

nasutum (Pratt). Transf. from Nasutocryptus Pratt (teste Townes and 
Townes, 1962, pp. 393, 398). 

orbum (Say). Host: Phereoeca walsinghami (Busck). 

Genus ACERASTES Cushman, p. 273 

pertinax Cresson. Cuba, Puerto Rico. 

Biology: Townes and Townes, 1962, pp. 404-405. 

Genus POLYCYRTIDEA Viereck, p. 273 

flavopicta (Ashmead). Remove from Catalog, p. 273, and Sup. 1, p. 47, not in 
Nearctic region. Remove limiiis Cushman from syn. {teste 
Townes and Townes, 1962, pp. 406-407). 

limitis Cushman. Tex.; Mexico, Costa Rica. Removed from syn. of flavopicta 
(Ashmead). A valid species {teste Townes and Townes, p. 406). 

*Subtribe NEMATOPODIINA 

Taxonomy: Townes and Townes, 1962. U. S. Natl. Mus. Bui. 216 (Pt. 
3) : 15, 20, 461. 

Insert as third Subtribe of Cryptini. 

Genus ACRORICNUS Ratzeburg, pp. 275-276, Sup. 1, pp. 47-48 

Xenodocon Foerster. Misspelled Xenocodon{l) in Catalog, pp. 275, 1283. 

Type species, Xenodocon ruficornis Foerster, is considered 
syn. of stylator (Thunberg) by Perkins, 1962, p. 462. 

stylator edwardsii (Cresson). Host: Sceliphron caementarium (Dru.). 

stylator excelsus (Cresson). Wyo., S. Dak. 

stylator junceus (Cresson). Host: Eu7nenes nest. 

Genus MESSATOPORUS Cushman, p. 276, Sup. 1 , p. 48 

compressicomis Cushman. Ohio, Tenn. Hosts: Ancistrocerus tuberculoce- 
phalus sutterianus (Sauss.) ; Trypoxylon collinum rubrocinctum 
Pack. 

discoidalis (Cresson). Miss. 

Mesostenus ferruniequinum Walsh and Riley. Preocc. 

Mesostenus jocosus Provancher, 1874. Nat. Canad. 6 (10) : 300. 5. 

Taxonomy: Townes and Townes, 1962, pp. 462-463, 465-467. 

ferrumequinujTi (Walsh and Riley) . Preocc. 

Mesostenus discoidalis Cresson. Removed from syn. as valid name for 

above. 

rufiventris Cushman. Transcont. Hosts: Auplopus nest; tPhanagenia hom- 
hycina nest; Trypoxylon (Trypoxylon) sp. 

♦Subtribe ECHTHRINA 

Taxonomy: Townes and Townes, 1962, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 
15, 20, 476. 

Place as fourth subtribe of Cryptini. 

Genus ECHTHRUS Gravenhorst, p. 273 

* Karaechthrus Uchida, 1929. Ins. Matsumurana 3 (4) : 176. (Syn. teste 
Townes and Townes, 1962, p. 486.) 



128 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Type: Karaechthrus tuberculatus Uchida. Orig. desig. and by 
monotypy. 
""Bioleter Meyer, 1930. Acad. Sci. USSR. Mus. Zool. Ann. 31: 169. (Syn. 
teste Townes and Townes, 1962, p. 486.) 

Type: (Bioleter eos Meyer) = tuberculatus (Uchida). Orig. 
desig. 

abdominalis abdominalis Cresson, changed status. Atlantic Ocean to Minn, 
in Canada. Zone. Reduced to subsp. rank (teste Townes and 
Townes, 1962, pp. 488, 489, 490). 

'''abdominalis diinidiatus Townes. Calif. 

Echthrus abdominalis dimidatus Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 488, 489, 491. $, ?. 

*abdominalis infuscus Townes. Wyo., Utah, Ariz., N. Mex. 

Echthrus abdominalis infuscus Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 488, 492. $, 9. 

abdominalis vancouverensis (Bradley), changed status. Alta. and B. C. to 
Wyo. and Calif. Reduced to subsp. rank {teste Townes and 
Townes, 1962, pp. 488, 489). 

adillae adillae Davis, changed status. B. C, Wash., Oreg. Reduced to subsp. 
rank {teste Townes and Townes, 1962, pp. 497, 498). 

*adillae brunneus Townes. N. Y., Va. 

Echthrus adillae brunneus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 497, 501. 2. 

*adillae nigriventris Townes. Labrador, Man., Mont., Colo., Alta., B. C, 
Alaska. Host: Paranthrene robiniae f. perlucida (Busck). 
Echthrus adillae nigriventris Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 497. $, 2. 

*adi!lae rubidus Townes. Oreg., Calif., Nev. 

Echthrus adillae rubidus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 496, 499. $, 5. 

^adillae semiruber Townes. Wash., Oreg. 

Echthrus adillae semiruber Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 2) : 497, 498. 3,9. 

*adillae tibialis Townes. Maine, Que., Ont. 

Echthrus adillae tibialis Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 497, 500. $,9. 

*niger emaculatus Townes. Maine, Que., Ont., Mich., Wis. 

Echthrus niger emaculatus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 493, 494. $,9. 

niger niger Cresson, changed status. Que., Ont., south to Va., Mich. Host: 
Saperda Candida F. Reduced to subsp. rank {teste Townes and 
Townes, 1962, pp. 494, 495). 

niger rufopedibus Harrington, changed status. Que. to B. C, Minn. Reduced 
to subsp. rank {teste Townes and Townes, 1962, pp. 493, 494). 

rufopedibus Harrington. A subsp. of niger Cresson. 

vancouverensis (Bradley). A subsp. of abdominalis Cresson. 

Genus XYLOPHRURUS Foersfer, pp. 273-274 

Xylophrurus Foerster. Change type citation to {Echthrus lancifer (Gra- 
venhorst)) = dispar (Thunberg). Included by Schmiedeknecht, 
1888. 

Nyxeophilus Foerster. Change type citation to: Cryptus bimaculatus 
Gravenhorst [as Thomson]. (Perkins, 1962, p. 439, considers 
this a valid genus.) 

Macrocryptus Thomson. Change type citation to (Echthrus lancifer Gra- 
venhorst) = dispar (Thunberg). By monotypy. 



ICHNEUMONIDAE 129 

*lSrxeophilus Thomson, 1885. Soc. Ent. France Ann. V (6) : 18. 
Type : Cryptus bimaculatus Gravenhorst. Orig. desig. 

agrili (Viereck). Mich. 

atr oruber Townes. Syn. of sitkensis (Ashmead). 

bicolor bicolor (Cushman), changed status. B. C, Idaho. Host: Saperda 
moesta LeC. Reduced to subsp. rank {teste Townes and Townes, 
1962, p. 482). 

*bicolor maurus Townes. Que. to Man. in Canad. Zone. 

Xlophrurus bicolor maurus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 482, 483. $,2. 

fasciatus fasciatus (Ashmead). Que., N. Y., Mich. 

*fasciatus hesperus Townes. Colo., Ariz., Calif. 

Xylophrurus fasciatus hesperus Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 480, 481. $, 9. 

luctuosus Provancher. A subsp. of nubilipennis Cresson. 

*nubilipennis abdominalis Townes. Sask., Utah, B. C. 

Xylophrurus nubilipennis abdominalis Townes, 1962. In Townes and 
Townes, U. S. Natl. Mus. Bui. 216 (Pt. 3) : 483, 484. $,9. 

nubilipennis luctuosus (Provancher), changed status. Que. and Ont. south to 
Fla. Host: Agnlus champlaini Frost; borer in Populus; Oberea 
praelonga Casey, Saperda Candida F. Reduced to subsp. rank 
(teste Townes and Townes, 1962, p. 483). 

nubilipennis nubilipennis (Cresson), changed status. Colo. Reduced to 
subsp. rank {teste Townes and Townes, 1962, pp. 483, 484). 

sitkensis (Ashmead). Alaska, B. C, Wash., Oreg., Calif. 

Xylophrurus atroruber Townes (syn. teste Townes and Townes, 1962, p. 
479). 

Genus AGONOCRYPTUS Cushman, p. 274 
discoidaloides (Viereck). Yiost: Psyrassa unicolor (Rand.). 
Genus CRYPTOHELCOSTIZUS Cushman, pp. 274-275 

alamedensis (Ashmead). B. C, Utah, Tex. Host: Borer in Agave stem. 

*caudatus Townes. Calif. (LakeTahoe). 

Cryptohelcostizus caudatus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 505, 511. ?. 

dichrous Viereck. Mo. 

*fuinipennis Townes. Ariz. (Sabino Canyon) . 

Cryptohelcostizus fumipennis Townes, 1962. In Townes and Townes, 
U. S. Natl. Mus. Bui. 216 (Pt. 3) : 505, 507. 9. 

*genalis Townes. Calif. (Capella, Mendocino Co.). 

Cryptohelcostizus genalis Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 504, 507. 9. 

*leionierus Townes. Calif. (Tanbark Flat, Los Angeles Co.). 

Cryptohelcostizus leiomerus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 505, 506. ?. 

*maculosu8 Townes. Calif. 

Cryptohelcostizus maculosus Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 505, 512. $. 

*nigricans Townes. Ariz. (Sierra Ancha) . 

Cryptohelcostizus nigricans Townes, 1962. In Townes and Townes, U. S. 
Natl. Mus. Bui. 216 (Pt. 3) : 505, 508. $, 9. 



130 HYMENOPTERA OF AMERICA NORTH OF MEXICO 



Genus HELCOSTIZUS Foerster, p. 275, Sup. 1, p. 47 

Brachycentrus Taschenberg. 

Helcostizus Foerster. 

Cyrtocryptus Marshall. 

Mesocryptus Thomson. 

Heterocryptus Woldstedt. 

Change type citation of each of above to (Cryptus brachycentrus Graven- 

horst) = albator ( Thunberg ) . 
Asternaulax Viereck. Change type citation to (Asternaulax fiskei Vier- 

eck) = albator canadensis (Provancher). 

albator canadensis (Provancher), changed status. Tenn., Mich. Reduced to 
subsp. rank {teste Townes and Townes, 1962, pp. 518, 521). H. 
albator albator (Thunberg) is European. 

albator rufiscutum Cushman, changed status. B. C, Oreg., Ariz. Host: 

borings of Plectura spinicauda Mannh. Reduced to subsp. rank 
(teste Townes and Townes, 1962, p. 518, 520). 

albator yukonensis (Ashmead), changed status. Alaska, N. W. T. south to 
Oreg. and Idaho. Reduced to subsp. rank {teste Townes and 
Townes, 1962, p. 519). 

annulicomis (Walsh). Md., Mich., Okla, Host: ? Elaphidionoides parallelus 
(Newn.). 
Helcostizus bicarinatus Cushman (syn. teste Townes and Townes, 1962, 
p. 515). 

bicarinatus CvLshman. Syn. of annulicornis (Walsh). 

canadensis (Provancher). A subsp. of albator (Thunberg). 

rufiscutum Cushman. A subsp. of albator (Thunberg). 

*subrectus Townes. Ariz. (nr. Alpine). 

Helcostizus subrectus Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 514, 517. 9. 

*tibialis Townes. Ont., N. Y., Mich., Wis. 

Helcostizus tibialis Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 514, 516. $, ?. 

*oxyura Townes. Ont., Mich. 

Helocostizus oxyura Townes, 1962. In Townes and Townes, U. S. Natl. 
Mus. Bui. 216 (Pt. 3) : 514, 517. ?. 

yukonensis (Ashmead). A subsp. of albator (Thunberg). 

Subfamily ICHNEUMONINAE 

Tribe PHAEOGENINI 

Genus PHAEOGENES Wesmael, pp. 276-278 

Phaeogenes Wesmael. After bibliog. ref. add: Proposed as subg. of "Ich- 
neumon Grav." 
*Tycherus Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 192 (syn. 
teste Perkins, 1962, p. 461). 

Type: Phaeogenes (Proscus) elongatus Thomson. Desig. by 
Perkins, 1962. 
*Proscus Holmgren, 1889. Ichn. Suecica 3: 420 (syn. teste Perkins, 1962, 
p. 461). 

Type: Phaeogenes cephalotes Wesmael. Desig. by Ashmead, 
1900. 

*Iaricellae Mason. N. B., Ont. Host: Argyresthia laricella Kft. 

Phaeogenes laricellae Mason, 1961. Canad. Ent. 93 (1): 3. $, 9. 

parvus (Provancher). Transf. to *Aethecerus Wesmael. 

pyriformis (Provancher). Transf. to *Jethsura Cameron. 



ICHNEUMONIDAE 131 

testaceicomis (Cameron). Transf. from Dicaelotus Wesmael, p. 280 (teste 
Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 104). 

vincibilis (Cresson). Transf. to *Takanona Uchida. 

walshiae walshiae Ashmead. Ohio. Host: Hendecaneura shawiana (Kft.). 

*Genus TAKANONA Uchida 

Takanona Uchida, 1926. Hokkaido Imp. Univ. Coll. Agr. Jour. 18: 158, 

163. 

Type: Takanona ishiyamanaV chida. Orig. desig. 
Syn. of Oronotus Wesmael, 1844, teste Townes, Momoi, and Townes, 1965, 

p. 420. 

Place after Phaeogenes Wesmael on p. 278. 

vincibilis (Cresson). Transf. from Phaeogenes Wesmael (teste Townes, 1957. 
Ent. Soc. Wash. Proc. 59 (3) : 119). 

*Genus JETHSURA Cameron 

Jethsura Cameron, 1902. Amer. Ent. Soc. Trans. 28: 373. 

Type: (Jethsura ferruginea Cameron) = pyrifortnis (Pro- 

vancher) . By monotypy. 

Place on p. 278, after Phaeogenes Wesmael and *Takanona Uchida. 

pyriformis (Provancher) . Transf. from Phaeogenes Wesmael, p. 277 (teste 
Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 109). 
* Jethsura ferruginea Cameron, 1902. Amer. Ent. Soc. Trans. 28: 374. §. 
(Syn. teste Townes, loc. cit. supra.) 

*Genus AETHECERUS Wesmael 

Aethecerus Wesmael, 1844. Acad. Sci. Bruxelles Nouv. Mem. 18: 166, 
203. Proposed as subg. of "Ichneumon Grav." 

Type: Aethecerus dispar Wesmael. Desig. by Ashmead, 1900. 

Place before Diadromus Wesmael on p. 278. 

parvus (Provancher). Transf. from Phaeogenes Wesmael (teste Mason, 1960. 
Canad. Ent. 92 (2): 145). 

*pinifoIii Mason. Calif., Alta. Hosts: Recurvaria milleri Busck, R. starki 
Freem. 
Aethecerus pinifolii Mason, 1960. Canad. Ent. 92 (2): 145. ?, $. 

*Genus THYRAEELLA Holmgren 

Thyraeella Holmgren, 1889. Ichn. Suecica, v. Ill, p. 402. 

Type: Ichneumon (Ischnus) collaris Gravenhorst. By mono- 
typy. 

Place before Diadromus Wesmael, on p. 278. 

erythrostomus (Cameron). Transf. from Diadromus Wesmael, p. 278 (teste 
Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 113). 

Genus DIADROMUS Wesmael, p. 278 
erythrostomus (Cameron). Transf. to *Thyraeella Holmgren. 

Genus DICAELOTUS Wesmael, pp. 279-280 

Leptoflemas Foerster. Considered syn. of Deloglyptus Foerster by Per- 
kins (first reviser), 1962, p. 435 (also in litt.). 
^Deloglyptus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 193. (A 
valid genus according to Perkins, 1962, p. 416.) 

Type: Deloglyptus punctiventris Thomson. Incl. by Thomson, 
1891. 
Deloglyptus and Leptodemas are left in synomymy here because a revi- 
sion of the species is not within the scope of this catalog. 

gelechiae (Ashmead). Ohio. 



132 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

missonriensis (Ashmead). 111., Va. 

testaceicomis (Cameron). Transf. to Phaeogenes Wesmael, p. 277. 

Tribe PLATYLABINI 
{ = Pr/it/'cerotm/ of Catalog) 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 697-704. 

Genus APAELETICUS Wesmael, p. 280 

americanus Cushman. 

Taxonomy: Heinrich, 1962, Pt. VI, pp. 792-793 ($ descr.). 

*brunnescens Heinrich. Maine. 

Apaeleticus brunnescens Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
793. $, ?. 

Genus CYCLOLABUS Heinrich, p. 280 

*albicinctus Heinrich. N. S. (Parrsboro). Host: A casis viridata Pack. 

Cyclolabus albicinctus Heinrich, 1962. Pt VI. Canad. Ent. Sup. 27: 756, 
757, 771. $, $. 

*arizonae Heinrich. Ariz. 

Cyclolabus arizonae Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 757, 
765. 5, ?. 

*carolmensis Heinrich. S. C. (nr. Tigerville) . 

Cyclolabus carolinensis Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
757, 764. 9. 

*dubiosus Perkins. (Misdet. as C. pactor (Wesmael) in Catalog, p. 280). 
Alta., B. C, Wash., Oreg., Colo., ?N. S., ?Newfoundland; Europe. 
Hosts: Eupithecia annulata Hist., E. filmata Pears, IE. luteata 
Pack., ?£■. palpata Pack. 
Cyclolabus dubiosus Perkins, 1953. Brit. Mus. (Nat. Hist.) Bui., Ent. 3 
(4): 156. $, 9. 

^gibsonatae Heinrich. Ont., B. C. Host: Eupithecia chagnoni Swett. 

Cyclolabus gibsonatae Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 757, 
764. ?. 

gracilicornis graciliconiis (Provancher) , changed status. Labrador, Que., 
Maine, N. H., N. Y., N. C. Transf. from Ectopius Wesmael, p. 281 
and reduced to subsp. rank (teste Heinrich, 1959, Ent. News 70 
(8): 216). 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 757, 758, 763 ($ descr.). 

gracilicornis subdentatus (Ashmead), changed status. Alaska to Calif., Colo. 
Host: Eupithecia edna. Hist. Transf. from Ectopius Wesmael, p. 
282 and reduced to subsp. rank (teste Heinrich, 1959. Ent. News 
70 (8) : 216. $, 9.). 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 757, 763-764 ($ descr.). 

impressus impressus (Provancher), changed status. Delete Wash, from 

distr., add: Maine, Ont. Transf. from Ectopius Wesmael, p. 281 

(teste Heinrich, 1959. Canad. Ent. 91 (12): 807). Reduced to 

subsp. rank (teste Heinrich, 1962, Pt. VI, pp. 757, 758, 759). 

Ichneumon erythropygus Provancher. Change sex sign to: "?" = 5- 

(Teste Heinrich, 1959, Canad. Ent. 91 (12) : 807.) 

*impressus infuscatus Heinrich. Alta., B. C, Wash. 

Cyclolabus impressus infuscatus Heinrich, 1962. Pt. VI. Canad. Ent. 

Sup. 27: 757, 758, 760. $, ?. 

*lobatus lobatus Heinrich. Maine, Que., N. Y. 

Cyclolabus lobatus lobatus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
757,758,761. $, ?. 



ICHNEUMONIDAE 133 



^lobatus occidentalis Heinrich. Alta. 

Cyclolabus lobatus occidentalis Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 

27: 756, 758, 761. $,9. 

pactor (Wesmael) . Remove from Catalog, p. 280; is European. 

^signatus (Provancher) . Maine, N. H. Transf. from Platylabus Wesmael, p. 
281 (teste Heinrich, 1959. Canad. Ent. 91 (12) : 807). 

Genus PLATYLABUS Wesmael, pp. 280-281 

IStenolabus Heinrich. Valid segregate; see *Asthenolabus Heinrich, n. 
name. 

*abbreviatus Heinrich. B.C. (Robson). 

Platylabus abbreviatus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 701, 
753. ?. 

*alaskae Heinrich. Alaska. 

Platylabus alaskae Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 700, 
721. ?. 

*albidorsus Heinrich. Maine, Que., Ont. 

Platylabus albidorsus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 702, 

740. 9. 

*arizonae Heinrich. Ariz., Calif. 

Platylabus arizonae Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 699, 

703, 715. $, ?. 

*benidi Heinrich, Maine, Ont. Hosts: Geometridae. 

Platylabus berndi Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 700, 703, 
731. $, 9. 

calif ornicus Cresson. Syn. of rufipes consors Cresson. 

canadensis Cresson. Transf. to Asthenolabus Heinrich. 

clarus (Cresson). Maine, Que., Ont., N. Y., Pa., N. C. Host: Abbotana cle- 
mataria A. & S. 
Platylabus metallicus Bradley. Remove from syn. A valid sp. 

*columbiae Heinrich. B. C, Calif. 

Platylabus columbiae Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 700, 
716. 9. 

consors Cresson. A subsp. of rufipes Provancher. 

*divisatae Heinrich. Newfoundland, Que., B. C. Host: Caripeta divisata 
Wlk. 
Platylabus divisatae Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 699, 

702,709. $,9. 

*duplificans Heinrich. Colo. 

Platylabus duplificans Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 700, 
703, 726. $,9. 

*erythrocoxa Heinrich. N. S., Maine, N. W. T., B. C, Calif. 

Platylabus erythrocoxa Heinrich, 1962. Pt. VI, Canad. Ent. Sup. 27: 

700, 703, 719. $, ?. 

*foxleei Heinrich. B. C, Alaska. 

Platylabus foxleei Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 700, 723. 
S, 9. 

gracllicomis (Viereck). Calif. Change bibliog. ref. p. 78 to 79. 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 702, 745-746. 

*hyperetis Heinrich. Maine, B. C. Host: Hyperetis amicaria. H.-S, 

Platylabus hyperetis Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 699, 

702,707. $,9. 

*imitans Heinrich. Maine, Que., B. C, Wash., Calif. Host: Enpithecia sp. 

Platylabus imitans Heinrich, 1962, Pt. VI. Canad. Ent. Sup. 27: 700, 
703, 725. $,9. 



134 HYMENOPTERA OF AMERICA NORTH OF MEXICO 



incabus Davis. B. C, Calif., Ariz. 

Taxonomy: Heinrich, 1962, Pt. VI, pp. 700, 703, 717-719 ($ descr.). 

lineolatus (Provancher) . Maine, Que., Ont., Mich., B. C. Host: Hydriomena 
renunciata columbiata Taylor. 
Phygadeuon rufipes Provancher. Remove from syn. A valid species. 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 701, 704, 734-736 {$ descr.). 

*lissosculptus Heinrich. Calif. (Leevining). 

Platylabus lissosculptus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 

704, 742. $. 

^^luteatae Heinrich. Que., Ont., Man., B. C, Oreg. Hosts: Eupithecia sp., E. 
luteata Pack. 
Platylabus luteatae Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 699, 

703, 715. $, 5. 

*melanocoxa Heinrich. B. C, Wash., Calif., Colo. Host: Eupithecia annulata 
Hist. 
Platylabus melanocoxa Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
700, 703, 727. $, 9. (P. pedatorius Fabricius in Catalog, mis- 
det.) 

metallicus Bradley. Maine, Que., N. H., Ont., Mich., Wash., B. C. A valid sp. 
{teste Heinrich, 1959. Ent. News 70 (8) : 215). 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 710-711. 

*micheneri Heinrich. Calif. (InyoMts.). 

Platylabus niicheneri Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 702, 
746. $. 

*monotonops Heinrich. N. Y. (Long Is.). 

Platylabus monotonops Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 

702, 744. ?. 

*monotonus Heinrich. N. H., N. Y., Ariz., Alta., B. C, Alaska. Hosts: Itame 
antaria Swett., I. qxiadrilinearia Pack., OpoHnia autumnata Gn. 
Platylabus monotonus Heinrich, 1962. Pt. VI. Canad. Ent, Sup. 27: 702, 

704, 741. $, ?. 

montanus Cresson. Mass., Mich. ; delete Wash. 

Platylabus vafcr (Cresson). Preocc. Change sex sign to: "?" = $. 
Platylabus perkinsi Walkley. (N. name for vafer (Cresson) Sup. 1, p. 
48.) Syn. teste Heinrich, 1959, Ent. News 70 (8) : 215. 

*opaculus americanus Heinrich. Transcont. in Transit. Zone. 

Platylabus opaculus americanus Heinrich, 1962. Pt. VI. Canad. Ent. 
Sup. 27: 701, 704, 730. $, . 

ornatus (Provancher). N. B., Maine, N. C, Ont., Sask., Alta., B. C. Hosts: 
Semiothisa granitata Gn. ; S. oweni Swett. 

pedatorius (Fabricius). Delete from Catalog; not in Nearctic fauna {teste 
Heinrich, 1962, Pt. VI, p. 727). 

perkinsi Walkley. Syn. of Platylabus montanus Cresson. 

*permodestus Heinrich. B. C, Wash., Calif., Colo. 

Platylabus permodestus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
700, 703, 720. $, 9. 

*polymelas Heinrich. Colo., Ariz., B. C. 

Platylabus polymelas Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 704, 
754. $. 

*pseudhistrio Heinrich. Maine (Dallas). 

Platylabus pseudhistrio Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
703,733. $. 

*rainieri Heinrich. Wash., B. C. 

Platylabus rainieri Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 700, 
702, 722, $,9. 



ICHNEUMONIDAE 135 

rubricapensis Provancher. Que., Ont., S. Dak., Idaho. 

*rubristernatus Heinrich. N. Y., N. C, Ohio, 111. 

Platylabus rubristernatus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
699, 713. 9. 

rufipes censors Cresson, changed status. Ariz., Calif., B. C, Alta., Alaska. 
Hosts: Hydriomena sp., H. renunciata Wlk. Reduced to subsp. 
rank (teste Heinrich, 1959. Ent. News 70 (8) : 215). 
Platylabus calif amicus Cresson (syn. teste Heinrich, 1959, op. cit. p. 
216). 

rufipes rufipes (Provancher), changed status. Que., Maine. A valid species; 
reduced to subsp. rank {teste Heinrich, 1959. Ent. News 70 (8): 
216). 

scutellatus (Provancher). Transf. to *AsthenoIabus Heinrich. 

Eurylabus agilis Cresson. Transf. to *Asthenolabus Heinrich as valid sp. 

*semiopacus Heinrich. Colo. (Phantom Valley). 

Platylabus semiopacus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 701, 

704, 730. $,9. 

*serratae Heinrich. N. B., Maine. Host: Euchlaena serrata Drury. 

Platylabus serraiae Heinrich, 1962, Pt. VI. Canad. Ent. Sup. 27: 702, 

704, 738. $, 9. 

*sexmaculatae Heinrich. Maine, Alta. Host: Semiothisa sexmaculata Pack. 
Platylabus sexmaculatae Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 

701, 749. 9. 

signatus (Provancher). Transf . to Cyclolabus Heinrich. 

*subrubricus Heinrich. Man., Sask., Alta. Host: Eupithecia ravocostiliata 
Pack. 
Platylabus subrubricus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
699,703, 714. $, ?. 

*tenuifonnis Heinrich. Ariz. (Sierra Ancha, Parker Creek). 

Platylabus tenuiformis Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 

702, 704,743. $, 9. 

tibialis Ashmead. N. Mex. 

Taxonomy: Heinrich, 1962, Pt. VI. pp. 700, 723 (relationships discussed). 

vafer (Cresson). Syn. of niontanus Cresson. 

*vaferops Heinrich. B. C, Wash., Calif. 

Platylabus vaf crops Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 701, 

704, 751. $, 9. 

♦Genus TROPICOLABUS Heinrich 

Tropicolabus Heinrich, 1959. Ent. News 70 (8) : 216. 

Type: Platylabus foxi Davis. By monotypy and orig. desig. 

Place in Platylabini after Platylabus Wesmael, p. 281. 

foxi (Davis). N. J. Transf. from Thaumatoteles Hopper and from syn. of T. 
ornatus (Cresson) p. 281 (teste Heinrich, 1959. Ent. News 70 
(8): 216). 

*Genus ASTHENOLABUS Heinrich 

Stenolabus Heinrich, 1935. Preocc. Transf. from syn. of Platylabus 

Wesmael (teste Heinrich, 1951. Bonn. Zool. Beitr. 2 (3/4) : 240). 
Asthenolabus Heinrich, 1951. Bonn. Zool. Beitr. 2 (3/4) : 240. N. name. 

Place on p. 281 of Catalog, after *Tropicolabus Heinrich. 

agilis (Cresson). N. S., P. E. I, Maine, Que., Mass., N. Y., N. W. T., S. Dak., 
Colo., Wyo., Oreg. Transf. from syn. of Platylabus scutellatus 
(Provancher) p. 281. A valid sp. (teste Heinrich, 1959. Ent. 
News 70 (8): 215). 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 778, 774-776. 



136 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

canadensis (Cresson). N. C. 

Taxonomy: Heinrich, 1962, Pt. VI, pp. 773, 776. $,9 ($ descr.). 

scutellatus (Provancher) . Que., Ont., N. Y. Transf. from Platylabus Wes- 
mael, p. 281 {teste Heinrich, 1959. Canad. Ent. 91 (12) : 807). 
Taxonomy: Heinrich, 1962. Pt. VI, pp. 773-774 (9 descr.). 

Genus PRISTiCEROS Gravenhorst, p. 281 

*Neopristiceros Heinrich, 1961 (1960). Pt. I, Canad. Ent. Sup. 15 [vol. 
92] : 13. (Syn. teste Heinrich, 1962, Pt. VI, pp. 783-784.) 

Type: Ichneumon lascivus Cresson. By monotypy and orig. 
desig. 

bakeri (Davis). Transf. to Pristicerops Heinrich. 

lascivus (Cresson). Maine, Que., Ont., Wash. 

Taxonomy: Heinrich, 1962, Pt. VI, pp. 783, 784, 785-786 {$ descr.; affini- 
ties). 

♦Genus PRISTICEROPS Heinrich 

Pristicerops Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 786. 
Type: Platylabus bakeri Davis. By orig. desig. 

Place on p. 281 of Catalog after genus Pristiceros Gravenhorst. 

bakeri bakeri (Davis), changed status. Maine, Que., N. H., Mich., Alta. 
Host: Deilinia sp. Reduced to subsp. rank and transf. from Pris- 
ticeros Gravenhorst (teste Heinrich, 1962, Pt. VI, pp. 786, 787, 
789). 

*bakeri townesi Heinrich. Wash. (Westport). 

Pristicerops bakeri townesi Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 

789. $, 9. 

Genus Thaumatofeles Hopper, p. 281 
Thauniatoteles Hopper. Syn. of Ambloplisus Heinrich. 

ornatus (Cresson). Transf . to Ambloplisus Heinrich. 

Platylabus foxi Davis. Transf. to Tropicolabus Heinrich as valid sp. 

*Genus AMBLOPLISUS Heinrich 

Ambloplisus Heinrich, 1930. Berlin Zool. Mus. Mitt. 15 (3/4) : 551. 

Type : Ambloplisus primus Heinrich. By monotypy. 
Thaumatoteles Hopper. (Syn. teste Heinrich, 1959. Ent. News 70 (8) : 
216.) 

ornatus (Cresson). N. C, Idaho. Transf. from Thaumatoteles Hopper (teste 
Heinrich, 1959. Ent. News 70 (8) : 216). Remove foxi Davis from 
syn., and transf. to Tropicolabus Heinrich as valid sp. 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 790-791 ($ descr.). 

Genus HYPOMECUS Wesmael, p. 281 
quadriannulatns (Gravenhorst). B. C. 

Genus Ecfopi'us Wesmael, pp. 281-282 
Delete from Catalog; no Nearctic spp. 
gracilicornis (Provancher). Transf. to Cyclolabus Heinrich, p. 280. 
impressus (Provancher). Transf. to Cyclolabus Heinrich, p. 280. 

subdentatus (Ashmead). Transf. to Cyclolabus Heinrich as subsp. of gracili- 
cornis (Provancher). 

thoracicus (Cresson). Transf. to Linycus Cameron as subsp. of exhortator 

(Fabricius). 



ICHNEUMONIDAE 137 

*Genus LINYCUS Cameron 

Linycus Cameron, 1903. Entomologist 36 : 234. 

*Type: Linycus rufipes Cameron. By monotypy. 

Place on p. 281 of Catalog, in place of genus Ectopias Wesmael (no 
Nearctic spp.). 

*exhortator montanus Heinrich. Colo. (Phantom Valley), L. exhortator ex- 
hortator (Fabricius) is European. 
Linycus exhortator montanus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 

27: 778,781. $. 

exhortator thoracicus (Cresson), changed status. Maine, Ont., Mich., Ind., 
Mo. Change $ sex sign after bibliog. ref. to: " $" = ?. Transf. 
from Ectopias Wesmael (teste Heinrich, 1962, Pt. VI, pp. 778, 
780). Reduced to subsp. rank (teste Heinrich, 1956. Canad. Ent. 
88 (11): 651). 

*nigriceps alaskae Heinrich. Alaska. 

Linycus nigriceps alaskae Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
778,782. $,9. 

nigriceps arizonensis Heinrich. Arizona (nr. Alpine). 

Linycus nigriceps arizonicus Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 
27: 778, 782. $. 

*nigriceps niger Heinrich. Alaska (Palmer). 

Linycus nigriceps niger Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
783. $. 

*nigriceps nigriceps Heinrich. B. C, Wash., Oreg. 

Linycus nigriceps nigriceps Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 

778, 781, 782. $, ?. 

Tribe LISTRODROMINI 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 687-688. 

Genus ANISOBAS Wesmael, p. 282 

Lycaenophilos Heinrich. Delete from syn. ; valid genus not in Nearctic 

fauna. 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 690-692 (key to spp.). 

bicolor Cushman. ?B. C. 

coloradensis (Cresson). S. Dak. 

Taxonomy: Heinrich, 1962, Pt. VI, pp. 691, 692, 693-694 ($ descr.). 

luzemensis (Bradley). Maine, Que., N. Y., Pa., Ont. 

Anisobas nearcticus Cushman (syn. teste Heinrich, 1962. Pt. VI, 691, 
692,695). 

texensis (Ashmead). Tex., Conn., ?Man. Hosts: llncisalia niphon Hbn., 
Strymon sp. 

Platylabus luzemensis Bradley. Remove from syn. Valid sp. (teste Hein- 
rich, 1962, Pt. VI, p. 695). 

Anisobas nearcticus Cushman. Transf. to syn. of luzemensis. 

Genus NEOTYPUS Foerster, p. 282 
nobilitator (Gravenhorst) . Alta. Host: Everes comyntas (Gdt.). 

Tribe iSCHNOJOPPINI 

Delete from Catalog (teste Heinrich, 1962, Pt. V, p. 661, in text under 
Trogomorpha Ashmead, which is placed in Ichneumonini). 



138 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Tribe ACANTHOJOPPINI 

Genus NARTHECURA Tov/nes, p. 284 

Transf. from Ichneumonini, p. 284 (teste Heinrich, 1962, Pt. VI, p. 681). 
Place after tribal heading Acanthojoppini, p. 283. 

*invadens Heinrich. Ariz., N. Hex. 

ISarthecura invadens Heinrich, 1962, Pt. VI. Canad. Ent. Sup. 27: 682. 

S, ?. 

Genus LOBAEGIS Townes, p. 284 

Transf. from Ichneumonini, p. 284 (teste Heinrich, 1962, Pt. VI, p. 683). 
Place before Pseudoplatylabus Smits van Burgst, p. 283. 

*8eptentrionalis Heinrich. N. Y., W. Va., S. C. 

Lobaegis septentrionalis Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 
684. $, ?. 

Genus PSEUDOPLATYLABUS Smits van Burgst, p. 283 

*towiiesi Heinrich. R. I., N. Y., Md. 

Pseudoplatylabus townesi Heinrich, 1962. Pt. VI. Canad. Ent. Sup. 27: 

686. $, 5. 

*violeiitus (Gravenhorst). Mich. ; Europe. Host: "Noctuid". 

Ichneumon violentus Gravenhorst, 1829. Iclin. Europaea, v. 1, p. 613. 
*Hoplismenus infaustus Wesmael, 1844. Acad. Sci. Bruxelles, Nouv. 
Mem. 18: 110. 

* Pseudoplatylabus caudatus Smits van Burgst, 1920. Ent. Ber. V (116) : 

282. ?. 
Taxonomy: Heinrich, 1962, Pt. VI, pp. 685-686. 

Tribe EURYLABINi 

Delete from Catalog; no Nearctic representatives. Transf. Probolus 
Wesmael to Ichneumonini, p. 283 (teste Heinrich, 1962, Pt. V, p. 518). 

Tribe ICHNEUMONINI 

[zzAmblytelini of Catalog) 

Genus TROGOMORPHA Ashmead, p. 282 
arrogans (Cresson). 

Taxonomy: Heinrich, 1962, Pt. V, pp. 662-663. 

Genus PROBOLUS Wesmael, p. 283 

detritus (Brulle). Transf. from syn. of Ctenichneuiiion svphax (Cresson), p. 
296; valid species (teste Townes, 1961. Ent. Soc. V/ash. Proc. 63 
(2): 107). 
Probolus illaetabilis (Cresson) (syn. teste Townes, 1961, loc. cit. supra). 

expunctus (Cresson). Maine, Man., B. C, ?Alaska. 

Probolus indistinctus (Provancher) (syn. teste Heinrich, 1962, Pt. V, p. 
520). 
illaetabilis (Cresson), Syn. of detritus (Brulle). 

indistinctus (Provancher). Syn, of expunctus (Cresson). 

*Genus PLATYLABOPS Heinrich 

Platylabops Heinrich, 1950. Mitt. Samml. Schmarotzerb. 27 : 4, 5. 
Type: Platylabops hinzi Heinrich. By monotypy. 

Place in lohneumonini before Hoplismenus Gravenhorst, p. 283. 

*apricus nearcticus Heinrich. Maine, N. Y. P. apricus apricus (Graven- 
horst) is European. 
Platylabops apricus nearcticus Heinrich, 1962. Pt. V. Canad. Ent. Sup. 
26: 641, 648. $,9. 



ICHNEUMONIDAE 139 

*eupitheciarum Heinrich. B. C, Alta. 

Platylabops eupitheciarum Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 
642, 648. $. 

*fraterculus Heinrich. Maine, Vt., N. Y. 

Platylabops fraterculus Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 641, 
642, 644. 5,9. 

*hoIervthrus Heinrich. B. C. Host: Eupithecia transcanndata Mack. 

Platylabops holerythrus Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 
641, 645. 9. 

leucopsis (Ashmead). Transf. from Aoplus Tischbein, p. 290 (teste Heinrich, 
1962, Pt. V, pp. 641,645). 
Taxonomy: Heinrich, 1962, Pt. V, pp. 641, 645-646 (9 descr.). 

*pecki novaescotiae Heinrich. N. S. 

Platylabops pecki novaescotiae Heinrich, 1962. Pt. V. Canad. Ent. Sup. 
26: 641, 643. 9. 

*pecki pecki Heinrich. Sask. 

Platylabops pecki pecki Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 
641, 642. 5, 9. 

Genus HOPLISMENUS Gravenhorst, pp. 283-284 

*Peritaenius Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 190. 
(Syn. teste Heinrich, 1949. Miinchen. Ent. Gesell. Mitt. 35-39: 
52.) 

Type: Peritaenius bavaricus Clement = Hoplismenus istrianus 
(Clement). Incl. and desig. by Clement, 1927. 
*Zanthojoppa Cameron, 1901. Ann. and Mag. Nat. Hist. (7) 7: 378. 
(Syn. teste Townes, Townes, and Gupta, 1961, p. 356.) 

Type: Zattthoioppa trilineata Cameron. By monotypy. 
*Xanthojoppa Cameron, 1903. Ann. and Mag. Nat. Hist. (7) 11: 182. 

Emend. 
*Taeniaspis Clement, 1927. Konowia 6 (1) : 67, 71. (Syn. teste Townes, 
Townes, and Gupta, 1961, p. 356.) 

Type: Taeniaspis flavitarsis Clement. By monotypy. 

morulus morulas (Say). Canad. Zone: N. W. T. 

morulus pacificus Cresson. Mont. 

*poIvleucos Heinrich. Ont. 

Hoplismenus polyleucos Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 
652, 659. $. 

praeruptus Swift. ?Ont. 

propitius (Cresson). Transf. to Ichneumon Linnaeus, p. 300. 

*rutilus borealis Heinrich. N. W. T., Man., Sask. 

Hoplismenus rutilus borealis Heinrich, 1962. Pt. V. Canad. Ent. Sup. 
26: 651, 652, 657. $, 9. 

rutilus rutilus (Cresson). P. E. I., Maine, Que., Ont. 

Hoplismenus scutellatus (Provancher) (syn. teste Heinrich, 1962, Pt. V, 
p. 656). 

rutilus tenuis Swift. Syn. of rutilus transversus Davis. 

rutilus transversus Davis, changed status. Oreg., Idaho, Wyo., Colo. Removed 
from syn. of scutellatus (Provancher) and reduced to subsp. rank 
(teste Heinrich, 1962, Pt. V, p. 656). 
Hoplismenus rutilus tenuis Swift (syn. teste Heinrich, 1962, Pt. V, p. 
656). 

scutellatus (Provancher). Syn. of rutilus rutilus (Cresson), p. 284. 
Hoplismenus transversus Davis. A valid subsp. of rutilus Cresson. 

feres Swift. Transf. to Hemihoplis Heinrich, infra. 



140 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Genus HEMIHOPLIS Hemrich 

Hemihoplis Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 15 [vol. 92]: 
15. 

Type: HopUsmenus teres Swift. By monotypy and orig. desig. 

Place on p. 284 after Hoplismenus Gravenhorst. 

teres (Swift). Transf. from Hoplismenus Gravenhorst, p. 384 (teste Hein- 
rich, 1961 (1960), Pt. I, p. 15; 1962, Pt. V, pp. 660, 661). 

*Genus HYBOPHORELLUS Schuiz 

*Hybophorus Tischbein, 1875. Stettin. Ent. Zeit. 36: 281. Preocc. 
Type: Ichneumon aulicus Gravenhorst. By monotypy. 
Hybophorellus Schuiz, 1911 (1909). Zool. Ann. 4 (1-2); 38. N. name. 

Place in Ichneumonini on p. 284 after Hoplismenus Gravenhorst. 

*baffinensis Heinrich. Baffin Is. 

Hybophorellus baffinensis Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 
562, 565. $. 

""injneundus nearcticus Heinrich, N. W. T. H. injucundus /injucundus/ (Wes- 
mael) is European. 
Hybophorellus injucundus nearcticus Heinrich, 1956. Canad. Ent. 88 
(2): 649,650. $, 9. 

*townesi Heinrich. Sask. 

Hybophorellus totcnesi Heinrich, 1956. Canad. Ent. 88 (2): 649. $, ?. 

Census LOBAEGIS Townes, p. 284 
Transf. to Acanthojoppini, p. 283. 

Census NARTHECURA Townes, p. 284 
Transf. to Acanthojoppini, p. 283. 

Genus PLACIOTRYPES Ashmead, p. 284 
concinnus (Say). N. Y. 

*Cenus RUBICUNDIELLA Heinrich 

Rubicundiella Heinrich, 1961 (1960). Pt. 1. Canad. Ent. Sup. 15 [vol. 
92] : 15. 

Type: Ichneumon tnucronatus Provancher. By monotypy and 
orig. desig. 

Place in Ichneumonini on p. 284 before Melanichneumon Thomson. 

annulicornis (Ashmead). Ga. Transf. from Melanichneumon Thomson, p. 
285 {teste Heinrich, 1962, Pt. V, pp. 568, 569). 

blanchardi (Davis). Maine, Que., N. H., Mass. Transf. from syn. of Melan- 
ichneumon rubicundus (Cresson), p. 286 {teste Heinrich, 1962, 
Pt. V, pp. 567, 568, 571). 
Taxonomy: Heinrich, 1962, Pt. V, p. 571 {$ descr.). 

*deuteromelas Heinrich. Maine, Yukon. 

Rubicundiella deuleromelas Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 
568, 573. $. 

mucronata (Provancher). Que., Md., Kans. 

Ichneumon mucronatus Provancher (erroneously syn. under Melanich- 
neumon rubicundus (Cresson), p. 286). Change sex symbol to: 
"9" = $. 

*perturbatrix Heinrich. Atlantic to 100° W. in Transit., U. and L. Austr. 

Zones, Calif.; Hawaii. 
Melanichneumon rubicundus (Cresson). Preocc. 
Melanichneumon mucronatus Auctt., not Provancher. 
Rubicundiella perturbatrix Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 

568, 577. N. name. 



ICHNEUMONIDAE 141 

*simpIicior Heinrich. Sask., B. C. Hosts: Fishia discors Grt. ; Ramosia bi- 
bioniformis (Bdvl.), IHeliothis ononis (Denis and Schiff.). 
Rubicundiella simplicior Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 

567,568,575. $, ?. 

Genus MELANICHNEUMON Thomson, pp. 284-287, Sup. 1, p. 49 

Bystra Cameron. Remove from Catalog. A valid, extralimital genus 
{teste Heinrich, 1937, Ann. and Mag. Nat. Hist. (Ser. 10) 20: 
261 ; Townes, Townes and Gupta, 1961, pp. 357, 464) . 

Gurfyia Cameron. Syn. of subg. Barichneiunon Thomson (teste Hein- 
rich, 1937, Ann. and Mag. Nat. Hist. (Ser. 10) 20: 265; Townes, 
Townes and Gupta, 1961, p. 363) . 

Harsaces Cameron. Syn. of subg. Barichneumon Thomson (teste Hein- 
rich, 1937, Ann. and Mag. Nat. Hist. (Ser. 10) 20: 265; Townes, 
Townes and Gupta, 1961, p. 363). 

Lissosculpta Heinrich (proposed as subg. of Melanichneunion Thomson). 
Raised to generic rank; extralimital (teste Heinrich, 1938. Mem. 
I'Acad. Malgache, f. 25: 23, 103; Townes, Townes, and Gupta, 
1961, pp. 378,466). 

Intermedichueumon Heinrich (proposed as subg. of Melanichneumon 
Thomson). Raised to generic rank; extralimital (teste Heinrich, 
1959. Ent. News 70 (8) : 211). 

Menkokia Heinrich (proposed as subg. of Melanichneumon Thomson). 
Removed from Catalog. Syn. of Bystra Cameron (teste Townes, 
Townes, and Gupta, 1961, p. 357) . 

absconditus (Provancher) . Placed in subg. Melanichneumon Thomson. 

albomarginatus (Cresson). Placed in subg. Vulgichneumon Heinrich. 

annulicornis (Ashmead). Transf. to Rubicundiella Heinrich, supra. 

blandii (Cresson), Placed in subg. Vulgichneumon Heinrich. 

brevicinctor (Say). Placed in subg. Vulgichneumon Heinrich. 

cordatus (Cresson). A subsp. of subcyaneus Cresson, in subg. Vulgichneu- 
mon Heinrich. 

disparilis (Cresson). Placed in subg. Melanichneumon Thomson. 

gestuosus (Cresson). Transf. to Ichneumon Linnaeus, p. 298. 

Cratichneumon pluto Viereck. Change page no. in bibliog, ref. to p. 79. 

Syn. of Melanichneumon nigripes (Provancher), p. 286. 
Amblyteles ohioensis DeGant. Syn. of nigripes (Provancher). 

heiligbrodtii (Cresson). Placed in subg. Melanichneumon Thomson. 

helvipes Cresson. Placed as syn. of anator (Fabricius) in subg. Barichneu- 
mon Thomson. 

honestus (Cresson). Placed in subg. Melanichneumon Thomson. 

leviculus (Cresson). Placed in subg. Melanichneumon Thomson. 

libens (Cresson). Placed in subg. Barichneumon Thomson. 

limbifrons (Cresson). Not subgenerically assigned. 

mimicus (Cresson). Placed in subg. Vulgichneumon Heinrich. 

mucronatus (Provancher), misdet. (Sup. 1, p. 49). Transf. to Rubicundiella 
Heinrich as syn. of perturbatrix Heinrich, n. name for rubicundus 
Cresson. 

nigripes (Provancher). Placed in subg. Melanichneumon Thomson. 

ornatipes (Cresson). Syn. of Melanichneumon (Rictichneumon) virginicus 

(Cresson) p. 287. 

residuus (Say). Placed in subg. Rictichneumon Heinrich. 



142 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

rubicundus (Cresson). See Sup. 1, p. 49. 

Ichneumon mucronatus (Provancher) . A valid sp. Transf. to Rubicun- 

diella Heinrich. 
Ichneumon blanchardi Davis. A valid sp. Transf. to Rubicundiella 

Heinrich. 

saevus (Cresson), Placed in subg. Vulgichneumon Heinrich. 

saundersU (Cresson), Transf. to Stenobarichneumon Heinrich. 

solitus (Cresson). Syn. of subcyaneus cordatus Cresson (subg. Vulgichneu- 
mon Heinrich) . 

soror (Cresson). Placed in subg. Barichneumon Thomson. 

subcyaneus (Cresson). Placed in subg. Vulgichneumon Heinrich. 

texanus (Cresson). Placed provisionally in subg. Melanichneumon Thomson. 

terminalis (Cresson). Placed provisionally in subg. Vulgichneumon Heinrich. 
Ichneumon finitimus Cresson. Syn. of subsp. terminalis (Cresson). 
Mesostenus apicalis Provancher. Placed in subg. Vulgichneumon Hein- 
rich as subsp. of terminalis (Cresson). 
Phygadeuon brevicaudus Provancher. Syn. of subsp. apicalis Provancher. 

virginicus (Cresson). Placed in subg. Rictichneumon Heinrich. 

vultus (Cresson). Not subgenerically assigned. 

zebratus (Cresson). Placed in subg. Vulgichneumon Heinrich. 

MELANICHNEUMON subgenus MELANICHNEUMON Thompson 

absconditus (Provancher). Maine, N. H., Alta., Wash., ?N. C. 

Taxonomy: Heinrich, 1962, Pt. V, pp. 584, 602-604 (2 descr.). 

belfragei (Cresson). Transf. from Pseudamblyteles Ashmead, p, 292 {teste 
Heinrich, 1959, Ent. News 70 (8) : 212). 

disparilis (Cresson). Que. 

Taxonomy: Heinrich, 1962, Pt. V, p. 587 (5 descr.). 

*dreisbachi Heinrich. Maine, Mich. 

Melanichneumon (Melanichneumon) dreisbachi Heinrich, 1962. Pt. V. 
Canad. Ent. Sup. 26: 584, 585. S, ?• 

*flavicarina Heinrich. Maine, Que., Ont., Man. Host: Protobarmia porcelaria 
Gn. 
Melanichneumon (Melanichneumon) flavicarina Heinrich, 1962. Pt. V. 
Canad. Ent. Sup. 26: 585, 594. $,9. 

*foxleei Heinrich. B.C. (Robson). 

Melanichneumon (Melanichneumon) foxleei Heinrich, 1962. Pt. V. 
Canad. Ent. Sup. 26: 584, 586. $. 

heiligbrodtii (Cresson). Md., N. C, S. C. 

Taxonomy: Heinrich, 1962, Pt. V. pp. 585, 592-594 (? descr.). 

honestus (Cresson). Md., N. C, S. C. 

Taxonomy: Heinrich, 1962, Pt. V. pp. 585, 592-594 (? descr.). 

*indecoratus Heinrich. P. E. I., Maine, Que., Ont., ? N. Y. 

Melanichneumon (Melanichneumon) indecoratus Heinrich, 1962. Pt. V. 
Canad. Ent. Sup. 26: 584, 589. $, 9. 

*leviculops Heinrich. Maine, Que., Ont. 

Melanichneumon (Melanichneumon) leviculops Heinrich, 1962. Pt. V. 
Canad. Ent. Sup. 26: 584, 600. 9. 

leviculus (Cresson). Que., Ont. 

Taxonomy: Heinrich, 1962, Pt. V, pp. 584, 599-600 (9 descr.). 

*lissorufus Heinrich. Labrador, Que., Maine, Ont., N. Y., Mich. 

Melanichneumon (Melanichneumon) lissorufus Heinrich, 1962. Pt. V. 
Canad. Ent. Sup. 26: 584, 591. $, 9. 

nigripes (Provancher). 

Cratichneumon pluto Viereck (syn. teste Heinrich, 1959, Ent. News 70 
(8): 212), 



ICHNEUMONIDAE 143 

Amblyteles ohioensis DeGant (syn. teste Heinrich, 1959, Ent. News 70 
(8): 212). 

* ?peranioenus Heinrich. Que., Ont., Md. 

Melanichneumon (?subgenus) peramoenus Heinrich, 1962. Pt. V. Canad. 
Ent. Sup 26: 631. $. 

?texanus (Cresson). Maine, Mich. 

Taxonomy: Heinrich, 1962, Pt. V, pp. 632-633 (9 needed for subg. as- 
signment) . 

*townesi Heinrich. N. Y., R. I. 

Melanichneumon (Melanichneumon) townesi Heinrich, 1962. Pt. V. 
Canad, Ent. Sup. 26: 585, 596. $, $. 

*MELAN!CHNEUMON subgenus VULGICHEUMON Heinrich 

Melanichneumon (Vulgichneumon) Heinrich, 1961 (1960). Pt. I. Canad. 
Ent. Sup. 15 [vol. 92]: 17. 

Type: Ichneumon brevicinctor Say. By monotypy and orig. 
desig. 

Place on page 287 of Catalog as second subg. of Melanichneumon Thom- 
son. Species placements teste Heinrich, 1962, Pt. V, pp. 605-619. 

albomarginatus (Cresson). Maine, N. J. 

Taxonomy: Heinrich, 1962, Pt. V, pp. 605, 611 (9 descr.). 

blantlii (Cresson). Ont. 

Taxonomy: Heinrich, 1962, Pt. V, pp. 605, 612-614 (9 descr.). 

brevicinctor (Say). Host: Papaipema nehris (Gn.). 

miniicus (Cresson). Newfoundland. 

Taxonomy: Heinrich, 1962, Pt. V, pp. 605, 614-616 (9 descr.). 

saevus (Cresson). Maine. 

subcyaneus cordaius (Cresson), changed status. Colo., Wash., B. C, ?N. Mex. 
Reduced to subsp. rank {teste Heinrich, 1959, Ent. News 70 (8): 
212). 
Ichneumon solitus Cresson (syn. teste Heinrich, loc. cit. supra). 

subcyaneus neutralis (Cresson), changed status. Reduced to subsp. rank 
(teste Heinrich, 1959, Ent. News 70 (8): 212). Transf. from 
(Pterocormus Foerster, Catalog pp. 299-300.) = Ichneumon Lin- 
naeus, Sup. I, p. 52 (teste Heinrich, loc. cit. supra). 

subcyaneus subcyaneus (Cresson), changed status. Eastern No. Amer. Host: 
Hyphantria cunea (Drury). Reduced to subsp. rank (teste Hein- 
rich, 1959, Ent. News 70 (8) : 212). 

?terminalis apicalis (Provancher) , changed status. Maine, Que., No. Ont. 
Removed from syn. and rdeuced to subsp. rank (teste Heinrich, 
1962, Pt. V, pp. 605, 619). 
Phygadeuon brevicaudus Provancher (syn. teste Heinrich, loc. cit. supra, 
p. 619). 

*?terminalis carolinensis Heinrich. N. C. 

Melanichneumon (Vulgichneumon) terminalis carolinensis Heinrich, 
1962. Pt. V. Canad. Ent. Sup. 26: 606, 619. 9. 

?terminalis terminalis (Cresson), changed status. So. Ont., Del., Md., 111., 
Iowa, Mo. Reduced to subsp. rank (teste Heinrich, 1962, Pt. V, 
pp. 606, 619). 
Taxonomy: Heinrich, 1962, Pt. V. pp. 605-606, 617-619 (generic place- 
ment). 

zebratus (Cresson). Transit. Zone in Mich., B. C. 

Taxonomy: Heinrich, 1962, Pt. V, pp. 605, 616-617 ($ descr,). 

MELANICHNEUMON subgenus BARICHNEUMON Thompson, Sup. 1, p. 49 
Place on p. 287 of Catalog as third subg. of Melanichneumon Thomson. 



144 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

anator (Fabricius). 

Ichneumon helvipes Cresson (syn. teste Heinrich, 1959. Ent. News 70 
(8) : 211; 1962 (Pt. V) : 621. 

*exceIsior Heinrich. Maine, Que., Ont. 

Melanichneumon (Bar ichneumon) excelsior Heinrich, 1962. Pt. V. Canad. 
Ent. Sup. 26: 620, 627. ?. 

libens (Cresson). Maine, N. C. 

Taxonomy: Heinrich, 1962, Pt. V., pp. 620, 626-627 (9 descr.). 

*sorex Heinrich. Maine, Que., Ont., Va. 

Melanichneumon (Bar ichneumon) sorex Heinrich, 1962. Pt. V. Canad. 

Ent. Sup. 26: 620,624. $, $. 

soror bimacularis (Dalla Torre), changed status. Maine, Que., Ont. Removed 
from syn. and reduced to subsp. rank {teste Heinrich, 1962, Pt. V, 
pp. 620, 624). 

soror soror (Cresson), changed status. Del. Reduced to subsp. rank {teste 
Heinrich, 1962, Pt. V, pp. 620, 623) . 
Ichneumon bimacularis Dalla Torre. A valid subsp. of soror {teste Hein- 
rich, 1962, Pt. V, p. 624). 

^MELANICHNEUMON subgenus RICTICHNEUMON Heinrich 

Melanichneumon (Rictichneumon) Heinrich, 1961 (1960). Pt. I. Canad. 
Ent. Sup. 15 [vol. 92] : 17. 

Type: Ichneumon residuus Say. By monotypy and orig. de- 
sig. 

Place on p. 287 of Catalog as fourth subgenus of Melanichneumon 
Thomson. 

residuus (Say) . 

Taxonomy: Heinrich, 1962, Pt. V, pp. 628-629. 

virginicus (Cresson). W. Va. 

Ichneumon ornatipes Cresson. Syn. {teste Heinrich, 1959, Ent. News 70 

(8): 212). 
Taxonomy: Heinrich, 1962, Pt. V, pp. 630-631. 

*Genus STENOBARICHNEUMON Heinrich 

Stenobarichneumon Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 15 
[vol. 92] : 16. 

Type: Ichneumon citator Thunberg. By monotypy and orig. 
desig. 

Place on p. 287 immediately following Melanichneumon Thomson. 

*agitator Heinrich. Maine, Que., Ont. 

Stenobarichneumon agitator Heinrich, 1962. Pt. V. Canad. Ent. Sup. 
26: 634, 639. ?, $. 

*duplicans Heinrich. Maine. 

Stenobarichneumon duplicans Heinrich, 1962. Pt. V. Canad. Ent. Sup. 
26: 634, 638. 5, ?• 

*pergracilis Heinrich. Maine, Que., Ont. 

Stenobarichneumon pergracilis Heinrich, 1962. Pt. V. Canad. Ent. Sup. 
26: 634, 635. $, $. 

*pygmaeops Heinrich. 

Stenobarichneumon pygmaeops Heinrich, 1962. Pt. V. Canad. Ent. Sup. 
26: 634,637. $, ?. 

saundersii (Cre?son). Maine, Que. Transf. from Melanichneumon Thomson, 
p. 286 {teste Heinrich, 1962, Pt. V, pp. 634, 636). 

Genus CRATICHNEUMON Thomson, pp. 287-290 

Revision: Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 18 [vol. 92]: 
98-173. 



ICHNEUMONIDAE 145 

altemans (Provancher) . 

Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 102, 153. 

*acronictae Heinrich. Que., Maine, N. Y., N. C. 

Cratichneumon acronictae Heinrich, 1961 (1960). Pt. II. Canad. Ent. 

Sup. 18 [vol. 92] : 101, 104, 125. $, 9. 

*anisotae Heinrich. Ont., Que., Maine, Conn., Mich. Host: Anisota i-uhicunda 
(F.). 
Cratichneumon anisotae Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 
18 [vol. 92]: 102, 105, 119. $, 9. 

annulatipes annulatipes (Provancher), changed status. Ont., Que., Maine, 
Mass., N. Y., Mich. Reduced to subsp. rank (teste Heinrich, 1961 
(1960), Pt. II, pp. 103, 107, 145, 146). 

*annulatipes facetops Heinrich. Mo. (Hannibal). 

Cratichneumon annulatipes facetops Heinrich, 1961 (1960). Pt. II. Canad. 

Ent. Sup. 18 [vol. 92]: 107, 147. $. 

annulatipes quintilis (Viereck), changed status. Conn, to N. C. Reduced to 
subsp. rank (teste Heinrich, 1961 (1960), Pt. II, pp. 103, 107, 
146). 

annulatus (Provancher). Maine. 

Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 101, 106, 140 (? descr.). 

arizonensis (Viereck). 

Taxonomy: Heinrich, 1961 (1960), Pt. II, p. 116 (under takomae) . 

ashmeadi (Schulz). N. H., Que., Colo. 

Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 99, 104, 118 (9 descr.). 

astutus (Holmgren). Transf. from Alphabetic List of Unplaced Species, p. 

408 (teste Townes, 1961. Ent. See. Wash. Proc. 63 (2) 106). 

*boreoalpinus Heinrich. Newfoundland, N. H. 

Cratichneumon boreoalpinus Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92]: 102, 168. 9. 

*boreovagans Heinrich. Maine, Que. 

Cratichneumon boreovagans Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 
29: 870. 9. 

brevipennis (Cresson). Maine, Ont., R. I., Fla. Host: Protoleucania albilinea 
(Hbn.). Transf. from Ichneumon (=Pterocormus in Cat., p. 297). 
Sup. 1, p. 51 (teste Heinrich, 1959, Ent. News 70 (8) : 214). 
Ischnus volens Cresson (syn. teste Heinrich, loc. cit. supra). 
Icheumon brevipennis var. obsoletus Riley (syn. teste Heinrich, Pt. II, p. 
170). 

citrinops kincaidi (Ashmead), changed status. Reduced to subsp. rank (teste 
Heinrich, 1959. Ent. News 70 (8) : 213). C. citrinops citrinops is 
European. 

confusus Ashmead. Transf. to *Aculichneumon subg. Crypteffigies Heinrich. 

duplicatus (Say). 

Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 100, 106, 149 (9 descr.). 

ericaceus Townes. Syn. of flavipectus Provancher. 

erythromelas (MacLachlan) . Transf . to Homotherus Foerster. 

*erythroscuta Heinrich. Mass. (Southbridge) . 

Cratichneumon erythroscuta Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92] : 101, 142. 9. 

facetus (Cresson). Mich. 

Taxonomy: Heinrich, 1961 (1960). Pt. II, pp. 103, 106, 159-160 (9 
descr.). 

*ferrugops Heinrich. Que., Ont., Mich. 

Cratichneumon ferrugops Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92] : 101, 124. 9. 



146 HYMENOPTERA OF AMERICA NORTH OP MEXICO 

flavipectus (Provancher). Ont., Que. to N. C. Deleted from syn. of scitulus 
(Cresson); a valid sp. {teste Heinrich, 1961 (1960), Pt. II, pp. 
103,107,162). 
Ichneumon pygmaeus Davis. Preocc. 

Cratichneumon ericaeus Townes, n. name (syn. teste Heinrich, 1961 
(1960), Pt. II, p. 162). 

*howdeni Heinrich. N. C. (Southern Pines). 

Cratichneumon howdeni Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 
18 [vol. 92] : 100, 172. ?. 

*insulae Heinrich. R. I., N. J. 

Cratichneumon insulae Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 
18 [vol. 92]: 100, 151. 9. 

kincaidi (Ashmead). A subsp. of citrinops Wesmael. 

nigritarius acerbus (Cresson). Que., Labrador, Huds. Zone. Host: Semioth- 
isa sexmaculata Pack. C. nigritarius nigritarius (Gravenhorst) is 
Eurasian. 

*paraparatus Heinrich. N. Y., N. J., Md., N. C, S. C. 

Cratichneumon paraparatus Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 
29: 869. $. 

paratus paratus (Say), changed status. Atlantic Ocean to 100° W in Transit. 
Zone. Reduced to subsp. rank {teste Heinrich, 1961 (1960), Pt. II, 
pp. 100, 105, 136). 
Ichneumon pusillus Cresson. Remove from syn. A valid subsp. of pa- 
ratus. 

paratus pusillus (Cresson), changed status. Del., Va. Reduced to subsp. rank 
{teste Heinrich, 1961 (1960), Pt. II, pp. 101 105, 137. 5, ?)• 

*paratus pseudovinnulus Heinrich. N. C, Fla. 

Cratichneumon paratus pseudovinnulus Heinrich, 1961 (1960). Pt. II. 
Canad. Ent. Sup. 18 [vol. 92] : 101, 105, 137. $, 9. 

?pectoralis (Say). Generic placement doubtful {teste Heinrich, 1961 (1960), 
Pt. II, p. 172). 

*pertenuis Heinrich. N. Y. (Ithaca). 

Cratichneumon pertenuis Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92] : 102, 169. $. 

*piceipes Heinrich. Maine, Que. 

Cratichneumon piceipes Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 
18 [vol. 92]: 101, 106, 130. $, ?. 

pilosulus (Pi'ovancher). 

Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 99, 104, 110 {$ descr.). 

popofensis Ashmead. A syn. of yakutatensis Ashmead. 

promptus (Cresson). 

Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 100, 104, 114-115. 

proximus (Cresson). 

Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 99, 104: 112-114. 

*pseudanisotae Heinrich. N. Y., N. J., Md. 

Cratichneumon pseudanisotae Heinrich, 1961 (1960). Pt. II. Canad. 
Ent. Sup. 18 [vol. 92] : 102, 133. 9. 

pteridis Townes. Ont., Man. Hosts: Protoboarmia porcelaria Gn., Semiothisa 
sp. 
Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 102, 107, 160-162 {$ 
descr.). 

*puncticoxa Heinrich. Que., Maine, Mass., N. Y. 

Cratichneumon puncticoxa Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92] : 101, 106, 133. $, 9. 



ICHNEUMONIDAE 147 

quintilis (Viereck). A subsp. of annulatipes (Provancher) . 
Ichneumon nanus Cresson. Preocc. 
Amblyteles nanodes Cushman. N, name. Syn. of C. scitulus (Cresson). 

*remanens Heinrich. Maine, Ont., N. Y., Md., N. C. 

Cratichneumon remanens Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92] : 107, 139, 157. $. 

*ritus Heinrich. Maine, Ont., N. C, Ind., Mich., Iowa. 

Cratichneumon ritus Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 18 
[vol. 92]: 100, 105, 143. $, ?. 

*rubricoides Heinrich. Que., N. Y., N. C, Mich. 

Cratichneumon rubricoides Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92] : 103, 106, 165. $, ?. 

*rubricops Heinrich. Que., Maine, Ont., N. Y. 

Cratichneumon rubricops Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92] : 103, 107, 167. $, 2. 

rubricus (Provancher). P. E. I., Maine, Mich. 

Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 103, 107, 164-165 {$ 
descr.). 

scitulus (Cresson). 

Ichneumon nanus Cresson (syn. teste Heinrich, 1961 (1960), Pt. II., p. 

154). 
ICryptus flavipectus Provancher. Remove from syn.; a valid sp. 
Amblyteles nanodes Cushman (syn. teste Heinrich, 1961 (1960), Pt. II, 

p. 154). 

signatipes (Cresson). Syn. of w-album w-album (Cresson) p. 289. 

Ichneumon lobatus Provancher. Remove from syn.; a valid subsp. of 
w-album (Cresson). 

suadus (Cresson). Maine, Ont. 

^subiilatus Heinrich. Que. to N. C. west to Mich. 

Cratichneumon subfilatus Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92]: 101, 106, 131. $, ?. 

sublatus (Cresson). Maine, N. H., Que., Ont. Host: Heterocampa biundata 
(Wlkr.). 

*takomae Heinrich. Md. (Takoma Park). 

Cratichneumon takomae Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 
18 [vol.92]: 104, 116. $. 

*tyloidifer Heinrich. Maine, Ont., N. Y., Pa., Md., N. C, Mich. 

Cratichneumon tyloidifer Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92]: 104, 115. $. 

unifasciatorius unifascialorius (Say), changed status. Eastern North Amer. 
Reduced to subsp. rank {teste Heinrich, 1961 (1960), Pt. II, p. 
109). 
Ichneumon vancouveriensis Provancher. A valid subsp. of unifasciato- 
rius (Say). 

unifasciatorius vancouveriensis (Provancher). B. C. Host: Arctiidae, IHali- 
siclota argentata (Pack.). Removed from syn. and reduced to 
subsp. rank {teste Heinrich, 1961 (1960), Pt. II, p. 109). 

*vaccinii Heinrich. Ont., Que. 

Cratichneumon vaccinii Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 
18 [vol.92]: 105, 141. $. 

variegatus (Provancher). Atlantic to 100° W in Transit, and U. Austr. 
Zones. Hosts: Anisota riibicunda (F.) ; A. sp. on beech; Hetero- 
campa guttivitta (Wlkr.). A valid species (teste Heinrich, 1959, 
Ent. News 70 (8): 214). 



148 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

vescus (Provancher). B.C. Host: Protobarmia porcelaria Gn. 

Taxonomy: Heinrich, 1961 (1960), Pt. 11, pp. 102, 105, 152-153 (? 
descr.). 

vinnulus (Cresson). Que., Ont., Maine to N. C. and Ky. 

*Cratichneumon gracilior Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup 18 [vol. 92]: 100, 105, 147. ? $, $. (Syn. (? only) teste 
Heinrich, 1962, Pt. VII, p. 868.) 

*vockerothi Heinrich. N. H. (Mt. Washington). 

Cratichneumon vockerothi Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92]: 102, 158. ?. 

volens (Cresson). Syn. of C. brevipennis (Cresson). 

w-album lobatus (Provancher), changed status. Que., Ont., New England, 
Mich. Transf. from syn. of signatipes Cresson and reduced to 
subsp. rank {teste Heinrich, 1961 (1960), Pt. II, pp. 101, 106, 
129). 

w-album w-album (Cresson), changed status. U. Austr. Zone north to South- 
ern N. England and Mich. Reduced to subsp. rank {teste Hein- 
rich, 1961 (1960), Pt. II, pp. 101, 106, 129). 

Cratichneumon signatipes (Cresson) (syn. teste Heinrich, 1961 (1960), 
Pt. II, pp. 127, 129). 

Ischnus variegaius Provancher. Remove from syn.; a valid sp. 

Ichneumon oryxicornis Viereck (syn. teste Heinrich, 1961 (1960), Pt. II, 
p. 129). 

Amblyteles duplicaliformis Viereck (syn. teste Heinrich, 1961 (1960), Pt. 
II, p. 129). 

yakutatensis Ashmead. 

Cratichneumon popofensis Ashmead. Syn. {teste Heinrich, 1959, Ent. 
News 70 (8): 214). 

*Genus HOMOTHERUS Foerster 

Homotherus Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25 : 185. 

Type: Ichneumon locutor Thunberg. IncL and desig. by 
Heinrich, 1961 (1960). 

Place in Catalog after Cratichneumon Thomson, p. 290. 

erythromelas (MacLachlan). Transf. from Cratichneumon Thomson, p. 288 
(teste Heinrich, 1962, Pt. VII, p. 871). 

*porceIariae Heinrich. Labrador, N. B., Maine, Ont., Man. Hosts: Protobar- 
mia porcelaria Gn., Eufidonia notatoria Wlkr., Semiothisa sp. 
Homotherus porcelariae Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 
18 [vol. 92]: 102, 105,177. $, 9. 

^semiaoplus Heinrich. Maine, Que., W. Va. 

Homotherus semiaoplus Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 
18 [vol. 92]: 103, 106, 173. $, ?. 

*townesi Heinrich. Maine, Que., Ont., N. Y., N. C, Mich. 

Homotherus townesi Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 18 
[vol.92]: 102, 106, 175. $, 9. 

*Genus ACULICHNEUMON Heinrich 
ACULICHNEUMON subgenus ACULICHNEUMON Heinrich 

Cratichneumon subg. Aculichneumon Heinrich, 1938. Acad. Malgache 
Mem. f. 25, p. 98. 

Type: Cratichneumon (Aculichneumon) longicauda Heinrich. 
By monotypy and orig. desig. 

This subgenus is not represented in the Nearctic Region. 

^ACULICHNEUMON subgenus CRYPTEFFIGIES Heinrich 
Aculichneumon subg. Crypteffigie* Heinrich, 1961 (1960). Pt. II. Canad. 



ICHNEUMONIDAE 149 

Ent, Sup. 15 [vol. 92] : 17. 

Type: Ichneumon (!) [Cratichneumon'] confusus Ashmead. 
By monotypy and orig. desig. 

Place in Catalog on p. 290 after Homotherus Foerster. 

confusus confusus (Ashmead), changed status. Alaska. Transf. from Cra- 
tichneumon Thomson and reduced to subsp. rank (teste Heinrich, 
1961 (1960), Pt. I, p. 17 (as Ichneumon confusus); Pt. II, p. 179). 

*confusus megalurus Heinrich. Ont. 

Aculichneumon (Crypteffigies) confusus megalurus Heinrich, 1961 
(1960). Pt. II. Canad. Ent. Sup. 18 [vol. 92]: 180. 9. 

*Genus EUPALAMUS Wesmael 

Eupalamus Wesmael, 1844. Acad. Sci. Bruxelles Nouv. Mem. 18: 13, 14. 

Type: Eupalamus oscillator Wesmael. Desig. by Ashmead, 
1900. 

Place in Ichneumonini, p. 290, after Cratichneumon. 

*andersoni Heinrich. B. C. (Stuarts Lake). 

Eupalamus andersoni Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 385. 
9. 

*melacneme Heinrich. Que., Ont. 

Eupalamus melacneme Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23; 
386. S, 9. 

*Genus PATROCLOIDES Heinrich 

Patrocloides Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 15 [vol. 
92]: 16. 

Type: Amblyteles perluctuosus Provancher. By monotypy and 
orig. desig. 

Place in Catalog on p. 290 before Aoplus Tischbein. 

montanus montanus (Cresson). Also no. Transit. Zone. Host: Syngrapha 
sp. Transf. from Patroclus Cresson, p. 291 {teste Heinrich, 1962, 
Pt. V, p. 517). 

'^montanus occidentalis Heinrich. Pacific to Cont. Divide in Canad. and 

Transit. Zones. 
Patrocloides montanus occidentalis Heinrich, 1962. Pt. V. Canad. Ent. 

Sup. 26: 517. 9. 
Patroclus montanus n. subsp. (of Catalog, p. 291). 

perluctuosus (Provancher). B. C. Hosts: Syngrapha epigaea (Grt.), S. 
selecta (Wlk.). Transf. from Patroclus Cresson. p. 291 (teste 
Heinrich, 1962, Pt. V, pp. 514, 516). 

walleyi (Heinrich). Que. Transf. from Patroclus Cresson, Sup. 1, p. 50 
(teste Heinrich, 1962, Pt. V, p. 513, 515). 

Genus AOPLUS Tischbein, pp. 290-291, Sup. 1, p. 49 

cestus (Cresson). Host: Melanolophia imitata Wlkr. 

Aoplus opacitas (Davis) (syn. teste Heinrich, 1959, Ent. News 70 (8) : 
211). 

confirmatus (Cresson). 

Taxonomy: Heinrich, 1962, Pt. V, pp. 535, 536, 537-539 ($ descr.). 

leucopsis (Ashmead). Transf. to Platylabops Heinrich (teste Heinrich, 1962, 
Pt. V, pp. 641, 645). 

*Iimbatae Heinrich. B. C. Host: Nematocavipa limbata Haw. 

Aoplus limbatae Heinrich. 1962. Pt. V. Canad. Ent. Sup. 26: 536, 542. 

9, $. 

*melanisticus Heinrich. Ont., Yukon, Canad. Zone. 

Aoplus melanisticus Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 535, 
547. 9. 



150 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*inoiIietti Heinrich. Alta. Host: Enypia moilietti Blkmre. 

Aoplus moilietti Heinrich, 1962. Pt. V. Canad. Ent. Sup. 62: 536, 544. 9. 

*inonotonus Heinrich. B. C, Wash., Oreg., Colo., Alta. 

Aoplus monotonus Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 536, 537, 
548. $, ?. 

*ochropis coloradensis Heinrich. Colo., ?Que. 

Aoplus ochropis coloradensis Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 
584, 536, 556. $, ?. 

ochropis ochropis (Gmelin), changed status. Reduced to subsp. rank (teste 
Heinrich, 1962, Pt. V, pp. 534, 536. 556). 

*permutabilis leucorbis Heinrich. Md., Mo. 

Aoplus premutahilis leucorbis Heinrich, 1962. Pt. V. Canad. Ent. Sup. 
26: 535, 542. 9. 

*perinutabilis permutabihs Heinrich. Maine, Que., Mass., Ont., Ind. 

Aoplus permutabilis perniutabilis Heinrich, 1962. Pt. V. Canad. Ent. 
Sup. 26: 535, 536, 541. S, 9. 

*pseudovelox Heinrich. Maine, Que., N. H., Ont., N. Y., Pa., Mich. Alta. 
Aoplus pseudovelox Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 535, 
537, 556. $, ?. 

*rarior Heinrich. Que., Ont. 

Aoplus rarior Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 535, 546. 9. 

ruficeps sitkensis (Ashmead). Not subsp. of vagans Provancher (teste 
Heinrich, 1959 (Oct.). Ent. News 70 (8) ; 211. A. ruficeps ruficeps 
is European. 

?Ichneutnon simulator Ashmead. 

Taxonomy: Heinrich, 1962, Pt. V, pp. 535, 536, 554. 

ruficeps vagans (Provancher). Maine, cooler parts of Transit. Zone. Host: 
Macaria sp. As here, not as vagans vagans Provancher (teste 
Heinrich, 1959 (Oct.). Ent. News 70 (8): 211). 
Taxonomy: Heinrich, 1959 (Dec), Canad. Ent. 91 (12): 807; 1962, Pt. 
V, pp. 535, 536, 553-554 (9 descr.). 

*thujarum Heinrich. Que., Ont . 

Aoplus thujarum Heinrich, 1962. Pt. V. Canad. Ent. Sup. 26: 536, 543. 9. 

*lorpidus (Wesmael). Maine, Que., Ont., Alta.; Europe. 

Ichneumon lorpidus Wesmael, 1857. Acad. Roy des Sci. Lettres, Beaux- 
arts Belgique Bui. (2)) 2: 385. 9. 

vagans sitkensis (Ashmead). Subsp. of ruficeps (Gravenhorst). 

vagans vagans (Provancher). Subsp. of ruficeps (Gravenhorst). 

velox oceidentalis (Harrington), changed status. B. C. Host: Lamhdina 
fiscellaria luqubrosa (Hist.). Reduced to subsp. rank (teste 
Heinrich, 1962, Pt. V, p. 551). 

velox velox (Cresson), changed status. Host: Ellopia athasaria (Guen.). 
Reduced to subsp. rank (teste Heinrich, 1962, Pt. V. p. 551). 
Ichneumon oceidentalis Harrington. Removed from syn. Valid subsp. of 
velox. 

Genus STENICHNEUMON Thomson, Sup. 1, pp. 49-50 

(Syn. of Aoplus Tischbein in Catalog, p. 290.) 
Change type citation to read: 

Type: (Ichneumon pistorius Gravenhorst) = militarius 
(Thunberg). 
*Myermo Cameron, 1901. Ann. Mag. Nat. Hist. (Ser. 7) 7: 523. (Syn. 
teste Townes, Townes, and Gupta, 1961: 345.) 

Type: (Myermo rufipes Cameron) = appropinquans 
(Cameron). By monotvpy. 
*Chiaglas Cameron, 1902. Ann. and Mag. Nat. Hist. (Ser. 7) 9: 152. 
(Syn. teste Townes, Townes, and Gupta, 1961: 345.) 



ICHNEUMONIDAE 151 

Type: Chiaglas nigripes Cameron. By monotypy. 
*Bolbomyschus Tosquinet, 1903. Soc. Ent. Belg. M6m. 10: 342. (Syn. 
teste Townes, Townes, and Gupta, 1961: 345.) 

Type: (Bolbomyschus albipictus Tosquinet) = nigripes 
(Cameron). By monotypy. 
*Stenichneumonoides Uchida, 1930. Insecta Matsumurana 9: 95. (Syn. 
teste Townes, Townes, and Gupta, 1961: 345). 

Type: Ichneumon posticalis Matsumura. Orig. desig. 
Taxonomy and Biology: Heinrich, 1962 (Pt. V): 522-524. 

culpator cinctioornis (Cresson). Transcont. in Canad. Zone. S. culpator 
cidpator (Schrank) is Eurasian. 
*Ichneuttton mathewi Cameron, 1907. Entomologist 40: 277. $. (Syn. 
teste Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 109.) 

^militarius leucopus Heinrich. Transcont. in Canad. Zone, N. B., Maine, Que. 
in North Transit. Zone. Host: Phisia sp. S. militarius militarius 
(Thunberg) is European. 
Stenichneumon militarius leucopus Heinrich, 1962. Pt. V. Canad. Ent. 
Sup. 26: 523, 524, 525. $, 9. 

pallidipennis (Viereck), changed status. Sask., N. W. T., B. C. Elevated to 
specific rank (teste Heinrich, 1962, Pt. V, pp. 523, 524, 531). 

salvus bioculaius (Cresson). N. B., P. E. I., Que., Man., Alta., B. C. 

Taxonomy: Heinrich, 1962, Pt. V, pp. 528, 531 (subsp. status doubtful). 

salvus pallidipennis (Viereck). A valid sp., not a subsp. 

salvus salvus (Cresson). 

Taxonomy: Heinrich, 1962, Pt. V, pp. 528-530 (specific status). 

Genus Patroclus Cresson, p. 291 

Remove from Catalog. Type-species is Neotropical. Species listed under 
this genus are transferred to *Palrocloides Heinrich. 

montanus n. subsp. Transf. to Patrocloides Heinrich as montanus occidentalis 

Heinrich {teste Heinrich, 1962, Pt. V, p. 517). 

montanus montanus (Cresson). Transf. to Patrocloides Heinrich. 
perluctuosus (Provancher) . Transf. to Patrocloides Heinrich. 
walleyi (Heinrich). Transf. to Patrocloides Heinrich. 

Genus ANISOPYGUS Kriechbaumer, p. 291 

*americanus Heinrich. Newfoundland, N. S., Maine, Mass., Ont., Wis., B. C. 
Host: Ichthyvra apicalis Wlkr. 
Anisopygus americanus Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 379. 

c?, 9. 

*pseudonyinus nearcticus Heinrich. Colo., Alta., B. C. Host: Notodontid 
pupa. A. pseudonymus pseudonymus (Wesmael) is European. 
Anisopygus pseudonymus nearcticus Heinrich, 1961. Pt. IV. Canad. Ent. 
Sup. 23: 378. $,9. 

Genus EXEPHANES Wesmael, 291 

*californicus Heinrich. Calif., Ariz., Utah, Colo. 

Exephanes californtcus Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 
476, 480. $,9. 

subfulvus (Cresson). Sask., ?Que. 

*terniinalis immaculatus Heinrich. Colo., ? N. W. T. 

Exephanes terminalis immaculatus Heinrich, 1961. Pt. IV. Canad. Ent. 
Sup. 23: 476, 479. $, 9. 

terminalis terminalis (Provancher), changed status. Transit. Zone, Maine, 
Que., Ont. Reduced to subsp. rank (teste Heinrich, 1961. Pt. IV, 
p. 479). 



152 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Spilichneumon pomilius Provancher (syn. teste Heinrich, 1959, Canad. 
Ent. 91 (12): 806). 

Genus ECTOPIMORPHA Viereck, pp. 291-292 

*boops Heinrich. Calif. (Crescent City). 

Ectopimorpha hoops Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 484, 
494. $. 

*caHfornira Heinrich. Calif. 

Ectopimorpha calif ornica Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 
484, 490. 3, ?. 

*hiulcops Heinrich. B. C, Sask. 

Ectopimorpha hiulcops Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 
484, 486. ?. 

indemnis (Cresson). Sask. 

*limerodops Heinrich. Calif. (Crane Flat). 

Ectopimorpha limerodops Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 

483, 487. 9. 

luperinae Cushman. 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 488-490 (disting. characters). 

*utahensis Heinrich. Utah (10 mi. north of Orderville). 

Ectopimorpha utahensis Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 

484, 493. $. 

*Genus PSEUDOAMBLYTELES Heinrich 

Pseudoamhlyieles Heinrich, 1926. Deut. Ent. Ztschr. 1926 (3) : 255. 

Type: Amblyteles homocerus Wesmael. By monotypy. 
^Ctenamhly teles Heinrich, 1928. Deut. Ent. Ztschr. 1928 (2) : 141. N. 
name. 

Place immediately after Ectopimorpha Viereck in Catalog, p. 292. 

*homoceru9 (Wesmael). Transcont. in Huds., Canad. Zones; Europe 
(Eurasia) ; Algeria; Tunisia. 
Atnblyteles homocerus Wesmael, 1854. Acad. Roy. des Sci. Lettres, 
Beaux -arts Belgique. Bui. Annexe: 121 $,9. 

Genus CHASMIAS Ashmead, p. 292 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 373-375. 

saucius (Cresson). Transf. to Ichneumon Linnaeus. 

scelestus (Cresson). Host: Papaipema cataphracta (Grote). 
Taxonomy: Heinrich, 1961, Pt. IV, pp. 374-375 ($ descr.). 

torvinus (Cresson). Syn. of saucius Cresson (transf. to Ichneumon, p. 301). 

*Genus NEAMBLYMORPHA Heinrich 

Neamblymorpha Heinrich, 1960. Pt. I. Canad. Ent. Sup. 15 [vol. 92]: 19. 
Type: Ichneumon milvus Cresson. By monotypy and orig. 
desig. 

Place on page 292 of Catalog after Chasmias Ashmead and before 
Eutanyacra Cameron. 

milva (Cresson). Transf. from Pseudamblyteles Ashmead, p. 293 {teste 
Heir.-.-ich, 1961, Pt. IV, pp. 376-377; $ descr.). 

Genus EUTANYACRA Cameron, Sup. 1, p. 50 

chillcotti Heinrich. N. W. T., Baffin Is., Que. 

consignata (Cresson). Ont. Transf. from Amblyteles Wesmael, p. 294 {teste 
Heinrich, 1961, Pt. IV, pp. 424, 425, 430; ? descr.). 



ICHNEUMONIDAE 153 

*hieinans Heinrich. Maine, Que., Ont., Pa. 

Eutanyacra hiemans Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 424, 
436. 9. 

*iinprovisa columbiana Heinrich. B. C. 

Eutanyacra improvisa columbiana Heinrich, 1961. Pt. IV. Canad. Ent. 
Sup. 23: 424, 428. $, 9. 

improvisa improvisa (Cresson), changed status. Maine, Ont., Sask. Hosts: 
? Ayiomogyna pergidvitata Morr. ("ex-A. pergui") ; Ceramica 
picta Harr. Reduced to subsp. rank {teste Heinrich, 1961, Pt. IV, 
pp. 424, 428). 

*miraoulosa Heinrich. Maine (Dryden). 

Eutanyacra miraculosa Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 
424, 434. $. 

munifica (Cresson). N. S., N. B., N. H., Sask. Transf. from Pseudamblyteles 
Ashmead, p. 293 {teste Heinrich, 1959, Ent. News 70 (8): 210). 
Taxonomy: Heinrich, 1961, Pt. IV, pp. 423, 424, 425, 426 444-446 (9 
tentative descr.). 

perannulata (Hopper). Transf. from Amblyteles Wesmael, p. 295 {teste 
Heinrich, 1959. Ent. News 70 (8): 210). 
Taxonomy: Heinrich, 1961, Pt. IV, pp. 424, 425, 429-430 (5 descr.). 

*pycnopus Heinrich. Ont., D. C, Ohio, Iowa, Ark. 

Eutanyacra pycnopus Heinrich 1961. Pt. IV. Canad. Ent. Sup. 23: 
424, 425, 435. $, 9. 

sag^uenaycnsis (Provancher) . Maine, Mich. Transf. from Pseudamblyteles 

Ashmead, p. 293 {teste Heinrich, 1961, Pt. IV, pp. 425, 426, 446- 
448; 9 descr.). 

*soIitaria Heinrich. Que., Ont., N. Y. 

Eutanyacra solitaria Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 425, 

442. S, 9. 

suturalis (Say). Labrador, N. S., Que., Ont., Man., Sask., N. W. T., B. C. 
Host: Actebia fennica Tausch. Transf. fi'om Pseudamblyteles 
Ashmead, p. 294 {teste Heinrich, 1959. Ent. News 70 (8): 210). 
Pseudamblyteles grotei (Cresson) (syn. teste Heinrich, loc. cit. supra). 

*trivittata Heinrich. Que. 

Eutanyacra trivittata Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 424, 
426, 448. $, 9. 

*valden!gra Heinrich. Que. (Pajme Bay). 

Eutanyacra valdenigra Heinrich, 1958. Canad. Ent. 90 (12): 740. $. 

*validiceps Heinrich. Maine, Que., Ont. 

Eutanyacra validiceps Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 425, 
426, 443. $, 9. 

*vilissima rubricosa Heinrich. Md., W. Va. 

Eutanyacra vilissima rubricosa Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 
23: 425, 441. $, 9. 

*vilissima vilissmia Heinrich. Maine, Que., Ont., Mass., N. Y., Mich. 

Eutanyacra vilissima vilissima Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 
23: 424, 426, 440. $, 9. 

Genus PSEUDAMBLYTELES Ashmead, pp. 292-294 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 397-401. 

*amoenipes Heinrich. Maine, Que. 

Pseudamblyteles amoenipes Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 
23: 400, 413. 9. 

animosus var. rubellus (Cresson). A valid species. 

belfragei (Cresson). Transf. to Melanichneumon Thomson, s. str. 



154 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

bizonatus (Cresson). Colo. 

laxonomy: Heinricn, 1961, Pt. IV, pp. 400, 401, 406 (possible subsp.). 

*carbo Hemrich. Maine, Que., Ont. 

Pseudamblyteles carbo Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 

399, 401, 419. $, $. 

citrinus (Provancher) . Transf. to Spilichneumon Thomson. 

♦distinctipes Heinrich. Maine, Que., Ont., N. Y., Md., Ohio, N. C. 

Pseudamblyteles distinctipes Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 
23: 400, 409. ?. 

*e£Sgialis Heinrich. N. B., Maine, Que., Ont. 

Pseudamblyteles effigialis Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 

400, 401, 407. $ 9. 

flebilis (Cresson). Maine, N. H., Que., Ont. 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 400, 401, 411 (? 5 descr.). 

grotei (Cresson). Transf. to Eutanyacra Cameron as syn. of E. suturalis 
(Say). 

*inlerdisli'nctus Heinrich. Maine, N. H., Que., Ont. 

Pseudamblyteles interdistinctus Heinrich, 1961. Pt. II. Canad. Ent. Sup. 
23: 400, 410. ?. 

koebelei (Swezey). Transf. to Spilichneumon Thomson as syn. of superbus 
(Provancher) . 

leuiisii (Cresson). Transf. to Ichneumon Linnaeus. 

micramoenus Heinrich. Maine. 

Pseudamblyteles micramoenus Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 
23: 399, 400, 420. ?. 

milvus (Cresson). Transf. to *Neamblymorpha Heinrich. 

mormonus (Cresson). Transf. with its syn. to *Thyrateles Perkins. 

munificus (Cresson). Transf. to Eutanyacra Cameron. 

*nigrotergops Heinrich. Que., Ont. 

Pseudamblyteles nigrotergops Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 
23: 399, 417. $. 

ormenus (Cresson). Transf. from Amblyleles Wesmael, p. 295 {teste Hein- 
rich, 1961, Pt. IV, pp. 399, 401, 414). 

*populorum Heinrich. Sask., Alta. Host: Brachylomia populi Strecker. 
Pseudamblyteles populorum Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 
23: 399, 401, 418. $, 9. 

provancheri (Cushman). Maine, Ont. Transf. from Amblyteles Wesmael, p. 
295 {teste Heinrich, 1961, Pt. IV, pp. 400,401, 404-406; $ descr.). 

robustus (Cresson). 

Taxonomy: Heinrich 1961, Pt. IV, pp. 400, 401 {$ descr.). 

rubellus (Cresson). Raised to specific rank {teste Heinrich, 1959, Ent. News 
70 (8): 211). 

saguenayensis (Provancher). Transf. to Eutanyacra Cameron. 

suturalis (Say). Transf. to Eutanyacra Cameron. 

♦walleyi Heinrich. Ont. (Smoky Falls). 

Pseudamblyteles walleyi Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 
399, 416. 9. 

*zebraticolor Heinrich. Que., Ont. 

Pseudamblyteles zebraticolor Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 
29: 882. $. 

Genus LIMERODOPS Hemrich, Sup. 1, p. 50 
Change type citation to read: Type: {Amblyteles elongatus Brischke) 



ICHNEUMONIDAE 155 

= "subsericans var. elongatus" (Brischke) {teste Heinrich, 1961, 
Pt. IV, pp. 380-382). 

belangeri (Cresson), changed status. Raised to sp. rank {teste Heinrich, 
1961, Pt. IV, p. 382). 

*niariannae Heinrich. N. S., Maine, Mass., Ont., Transit. Zone. 

Limerodops mariannae Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 383. 
S, ?. 

Genus AMBLYTELES Wesmael, pp. 294-295 

Delete from Catalog; contains no Nearctic species {teste Heinrich, 1961, 
Pt. IV, p. 398. 

aleatorius (Cresson). Transf. to Spilichneumon Thomson, p. 295, as syn. 
of subrufus (Cresson). 

belangeri Cresson. Transf. in Sup. 1 to Limerodops Heinrich. 

bronteus (Cresson). Transf. to Spilichneumon Thomson, p. 295. 

consignatus (Cresson). Transf. to Eutanyacra Cameron. 

crudosus (Cresson). Transf. tentatively to Spilichneumon Thomson, p. 295. 

fraternus Crefson. Transf. to Spilichneumon Thomson, p. 295, as syn. of 
nubivagus (Cresson). 

improvisus (Cresson). Transf. to Eutanyacra Cameron. 

inconstans (Cresson). Transf. to Spilichneumon Thomson, p. 295, as syn. of 
subrufus (Cresson). 

juxtus (Cresson). Transf. to Spilichneumon Thomson, p. 295. 

kodiakensis (Ashmead). Transf. to Spilichneumon Thomson, p. 295. 

nubivagus (Cresson). Transf. to Spilichneumon Thomson, p. 295. 

ormenus (Cresson). Transf. to Pseudamblyteles Ashmead, p. 293. 

perannulatus Hopper. Transf. to Eutanyacra Cameron. 

provancheri Cushman. Transf. to Pseudamblyteles Ashmead, p. 293. 

rufizonatus (Cresson). Transf. to Ctenichneumon Thomson as syn. of 
syphax ( Cresson ) . 
Amblyteles balteatus Hopper. Transf. to Ichneumon Linnaeus as syn. 
of volesus Cresson. 

subrufus (Cresson). Transf. to Spilichneumon, Thomson, p. 295. 

superbus Provancher. Transf. to Spilichneumon Thomson, p. 295. 

torontosus Davis. Transf. to Thyrateles Perkins as syn. of procax (Cresson). 

Genus SPILICHNEUMON Thomson, p. 295 

borealis (Provancher). Que., Ont. A valid species {teste Heinrich, 1961 
(1960), Pt. II, p. 185). 
Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 185-186 {$ descr.). 

bronteus (Cresson). Que., Ont., B. C. Transf. from Amblvteles Wesmael, p. 
294 {teste Heinrich, 1961 (1960) Pt. II, pp. 182, 183, 186). 
Taxonomy: Heinrich, 1961 (1960), Pt. II, pp. 187-188 (9 descr.). 

citrinus (Provancher). Transf. from Pseudamblyteles Ashmead, p. 292 {teste 
Heinrich, 1961 (1960), Pt. II, pp. 183, 189). 

?crudosus (Cresson). Transf. from Amblyteles, p. 294 (no spms. seen; descr. 
not definitive). 

juxtus (Cresson). Transf. from Amblyteles, p. 294 {teste Heinrich, 1959, 
Ent. News 70 (8) : 209). 

kodiakensis (Ashmead), n. comb. Transf. from Amblyteles, p. 294 {teste 
exam, of $ type spm.). 



156 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

uubivagus (Cresson). Labrador. Transf. from Amblyteles Wesmael, p. 294 
(teste Heinrich, 1959, Ent. News 70: 209). 
Amblyteles /raternus Cresson (syn. teste Heinrich, loc. cit. supra). 

*physcoteIoides Heinrich. Que., Maine, Mass. 

Spilichneumon phyacoteloides Heinrich, 1961 (1960). Pt. II. Canad. Ent. 
Sup. 18 Lvoi. 92]: 182, 183, 191. $ ?. 

pomilius (Provancher). Transf. to Exephanes Wesmael as syn. of terminalis 
tcrminalis (Provancher), p. 291. 
Amblyteles borealis Provancher. Remove from syn.; a valid species. 

subrufus (Cresson). Transcont. in Transit., U. Austr. zones, Newfoundland, 
N. W. T. Host: Crymodes devastator Brace. Transf. from 
Amblyteles Wesmael, p. 295 (teste Heinrich, 1959, Ent. News 
70 (8) : 209). 

Amblyteles aleatorius (Cresson) (syn. teste Heinrich, loc. cit. supra). 

Amblyteles inconstans (Cresson) (syn. teste Heinrich, loc. cit. supra). 

?superbus (Provancher). N. S. to B. C, N. W. T., Colo., Tex., Hawaii. Hosts: 

Euxoa flavicollis Sm., E. messoria Harr., E. ochrogaster Gn., E. 

scandens Riley, Feltia ducens Wlkr., Chorizagrotis auxiliaris Grt. 

Transf. from Amblvteles Wesmael, p. 295 (teste Heinrich, 1959. 

Ent. News 70 (8): 209). 
Pseudambly teles koebelei (Swezey) (syn. teste Heinrich, loc. cit. supra). 
Taxonomy: Heinrich, 1961 (1960). Pt. II. Canad. Ent. Sup. 18 [vol. 

92] : 182, 192-194. 

laos (Cresson). Que., Maine, Mass. 

*valfletypicus Heinrich. Man. (Riding Mt. Natl. Pk.). 

Spilichneumon valdetypicus Heinrich, 1961 (I960). Pt. II. Canad. Ent. 
Sup. 18 [vol. 92]: 182, 186. ?. 

*Genus OBTUSODONTA Heinrich 

Obtusodonta Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 872. 

Type: Spilichneumon obscuricolor Heinrich. Orig desig. 

Place in Catalog on p. 295 after Spilichneumon Thomson. 

*£oxleei Heinrich. B. C. 

Obtusodonta foxleei Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 873, 
874. 9. 

*manitobae Heinrich. Man., Alta. 

Obtusodonta manitobae Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 
873, 874. 9. 

*montana Heinrich. Que., Maine. 

Obtusodonta montana Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 
873, 875. S, ?. 

♦obscuricolor (Heinrich). Que., Maine, Conn., N. Y., Ont., Sask. 

(^?) Spilichneumon obscuricolor Heinrich, 1961 (1960). Pt. II. Canad. 

Ent. Sup. 18 [vol. 92]: 182, 183, 195. 1$, 9. 

Genus CTENICHNEUMON Thomson, pp. 295-296 

*caeruleop8 Heinrich. Sask., B. C. 

Ctenichneumon caeruleops Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 
452, 453, 461. $,9. 

"^columbianus Heinrich. B. C, Yukon, Alaska. 

Ctenichneumon columhianus Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 
23: 452, 453, 457. 5, ?• 

electus (Cresson). Questionable syn. of syphax (Cresson). 

cxcultus (Cresson). 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 452, 453, 454-457. 



ICHNEUMONIDAE 157 

•gracilior Heinrich. Maine, Que. 

Ctenichneumon gracilior Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 
452, 453, 470. $, ?. 

•heteropus Heinrich. B. C. 

Ctenichneiimon heteropus Heinrich, 1961. Pt. IV. Canad. Ent. Sup, 23: 

452, 453, 454. $,9. 

""holomelas Heinrich. Man. (Cedar Lake). 

Ctenichneumon holomelas Heinrich, 1961. Pt. VI. Canad. Ent. Sup. 23: 

451, 460. ?. 

♦minor Heinrich. N. S., N. B., Maine, Que., Ont., Conn., N. Y., W. Va., 
Man., Sask. 
Ctenichneumon minor Heinrich 1961. Pt. IV. Canad. Ent. Sup. 23: 451, 

453, 458. $, ?. 

*punotiscuta Heinrich. Man., B. C. 

Ctenichneumon punctiscuta Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 

452, 463. 9. 

*rufibasis Heinrich. Maine, Que. 

Ctenichneumon rufibasis Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 
452, 469. ?. 

ruidosensis ( Cockerel! ) . 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 453, 474-475. 

seinicaeriileus (Cresson). Man., Alta. 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 452, 453, 461 ($ descr.). 

syphax (Cresson). Atlantic to 100° W in U. Austr., Transit., Canad. Zones, 

Alta. 
?Ichneumon detritus Brulle. Transf. to Probolus Wesmael, p. 283, as 

valid sp. 
? Ctenichneumon electus (Cresson) (syn. teste Heinrich, 1961, Pt. IV, p. 

471). 
Ichneumon marianapolitanensis Provancher (syn. teste Heinrich, 1959, 

Ent. News 70 (8) : 210). 
?Ichneunton nitidus Provancher (syn. teste Heinrich, 1961, Pt. IV, p. 

471). 
Aniblyteles rufizonatus (Cresson) (syn. teste Heinrich, 1959, Ent. News 

70 (8): 210). 

*townesi Heinrich. Maine, N. Y., Mich. 

Ctenichneumon townesi Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 
452, 453. $, 9. 

ultus (Cresson). 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 452, 453, 464-467 (9 descr.). 

♦Genus PROTOPELMUS Heinrich 

Protopelmus Heinrich, 1959. Ent. Soc. Wash. Proc. 61 (5) : 200. 

Type: Trogus atrocoeruleus Cresson. By monotypy and orig. 
desig. 

Place on p. 296, before Tricholabus Thomson. 

atrocoeruleus (Cresson). Fla., Tex. Host: Ecpantheria deflorata (F.) ; E. 
muzina Oberth. Transf. from Trogus in Alphabetic List of Un- 
placed Species, p. 409 (teste Heinrich, 1959, Ent. Soc. Wash. 
Proc. 61 (5) : 200). 
Taxonomy: Heinrich, 1962, Pt. V, pp. 511-512. 

Genus TRICHOLABUS Thomson, p. 296, Sup. 1, p. 51 

Otohimea Uchida, 1926. Hokkaido Imp. Univ. Coll. Agr. Jour. 18: 146. 
Type: [Otohimea nigra uchida) = incanescens (Smith). 
Orig. desig. 

adventicus (Hopper). Ala., Fla. 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 389, 393-395 {$ descr.). 



158 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

citatus (Provancher) . 

Ichneumon merus Cresson. Remove from syn. and place as syn. of 

nortonii ( Cresson ) . 
Tricholobus tricolor (Davis) (syn. teste Heinrich, 1961, Pt. IV, p. 391). 

*foxIeei Heinrich. B. C, Calif. 

Tricholabus foxleei Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 389, 
396. S, ?. 

*milchelH Heinrich, Mass., N. Y. to Ga., Ark. 

Tricholabus mitchelli Heinrich, 1961. Pt. IV. Canad. Ent. Sup. 23: 389, 
395. $, 9. 

nortonii (Cresson). 

Ichneumon merus Cresson (syn. teste Heinrich, 1961, Pt. IV, pp. 389, 
390). 

pax (Dalla Torre). 

Taxonomy: Heinrich, 1961, Pt. IV, pp. 389, 397. 

tricolor (Davis). Syn. of citatus (Provancher). 

*Genus THYRATELES Perkins 

Thyrateles Perkins, 1953. Brit. Mus. (Nat. Hist.) Bui. Ent. 3 (4): 149. 
Type: Amblyteles camelinus Wesmael. By monotypy and 
orig. desig. 
Taxonomy: Heinrich, 1961. Pt. Ill, pp. 348-349. 

Place on page 296 of Catalog after Tricholabus Thomson. 

*amoenapex Heinrich. N. W. T. (Norman Wells). 

Thyrateles amoenapex Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 348, 357. 9. 

*caIiginops Heinrich. Maine, Que., Ont., Conn., N. Y., W. Va., Minn. Host: 
Vanessa atalanta (L.). 
Thyrateles caliginops Heinrich. 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 212, 215, 226, 348, 351. $, 9. 

*camelinops Heinrich. Alta., B. C. Host: Nymphalis milberti Godt; 
Polygonia sp. 
Thyrateles camelinops Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 212, 215, 348, 349. $, 9. 

instabilis (Cresson). Ho.^ts: Colias pJiilocUce Latr. ; place ? before Oeneis 
inelissa {¥.). Transf. from (Pterocormus Foerster of Catalog, 
p. 299) = Ichneumon Linnaeus, Sup. 1, p. 51 {teste Heinrich, 
1961, Pt. Ill, pp. 284 (text, 348, 352). 

lugubrator (Gravenhorst) . Transf. from Alphabetic List of Unplaced Spe- 
cies, p. 408 {teste Heinrich, 1961, Pt. Ill, p. 354). 

Pterocormus rufiventris (Brulle) (syn. teste Townes, 1961 (June). Ent. 
Soc. Proc. 63: 108). 

Pterocormus rufiventris var. incertus (Cresson) (syn. teste Heinrich, 
1961, Pt. Ill, p. 354). 

mormonus (Cresson). Ont. Host: Polygonia sp. Transf. (with its syn). 
from Pseudamblyteles Ashmead, p. 203 {teste Heinrich, 1959. Ent. 
News 70 (8) : 210). 

procax (Cresson). Host: Vanessa antiopa (L.). Transf. from {Pterocormus 
Foerster of Catalog, p. 300) = Ichneumon Linnaeus {teste 
Heinrich, 1959, Ent. News 70 (8): 210). 
Amblyteles torontosus Davis (syn. teste Heinrich, loc. cit. supra). 

Genus ICHNEUMON Linnaeus, Sup. 1, pp. 51-52 
( = P/erocormos Foerster of Catalog, pp. 296-301) 

Brachypterus Gravenhorst. Delete "Desig. by Westwood, 1839" and 
add: By monotypy. 



ICHNEUMONIDAE 159 

Pterocormus Foerster. N. name for Brachypterus Gravenhorst. Delete 
"Orig. desig." and add: By substitution of Pterocormus for 
Brachypterus. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 211-321; 1962, Pt. VII, pp. 877- 
881. 

ambulatorius Fabricius. Newfoundland, Que. in Canad. Zone; B. C. Hosts: 
Faronta diffusa Wlkr. ; Hydroecia immanis Gn. 
Taxonomy: Heinrich, 1961, Pt. III. pp. 218, 228, 229, 242-244 (var. 
mutator Cresson). 

*annulatorius Fabricius. Newfoundland. 

Pterocormus funestus (Cresson) (syn. teste Heinrich, 1961, Pt. Ill, pp. 

232, 246, 248). 
Taxonomy: Heinrich, 1961, Pt. Ill, pp. 223, 230, 233, 246-248. 

*anonymus Heinrich. Maine, Que., Ont., Mass., N. Y., Pa., W. Va. Md. 

Ichneumon anonymus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol: 
93]: 221, 222, 224, 255. $, 9. 

approximans Provancher. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 228, 343-344. 

artemis (Viereck). 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 213, 312-313. 

atrox Cresson: Labrador, Ont., Man., B. C. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 217, 226, 328-329 ($ descr.). 

bimembris Provancher. Ont. (Canad. Zone). Elevated to specific rank 
(teste Heinrich, 1961, Pt. Ill, pp. 336-337); reduced to subsp. of 
languidus Wesmael in Sup. 1, p. 51. 

brevipennis (Cresson). Transf. to Cratichneumon Thomson. 

*broweri Heinrich. Maine (Mt. Katahdin). 

Ichneumon broweri Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
931: 215, 337. 9. 

*browni Heinrich. Que. (Mt. Lyall, Gaspe). 

Ichneumon browni Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 93) : 

225, 277. 9. 

hurkei (Viereck). Transf. to Orgichncumon Heinrich as syn. of calcatorius 
(Thunberg). 

caliginosus Cresson. Transcont., N. W. T. Hosts: Delete Basilarchia 
archippus (Cram.) and Vanessa atalanta (L.) ; add Polygonia 
QTacilis Grote. 
Taxonomy: Heinrich, 1961, Pt. Ill, pp. 212, 215, 229, 332-334 ($ descr.; 
sp. restricted). 

calitergus Cresson. Que. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 216, 229, 326-327 ($ descr.). 

canadensis Cresson. Newfoundland, N. S., P. E, I., N. B., Maine, Que., Ont. 
Host: Pseudaletia unipuncta (Haw.). 
Amblyteles ctenuchae Cushman. Remove from syn. and place as syn. of 
trizonatus Provancher {teste Heinrich, 1961, Pt. Ill, p. 238; also 
Heinrich, 1959, Ent. News 70: 104 (desig. new lectotype)). 

*canadicola Heinrich. Newfoundland, Que., Maine, Ont., Mass. Transit, and 
Canad. Zones. 
Ichneumon canadicola Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93) : 219, 293. 9. 

centrator Say. B. C. 

?cervulus Provancher. Maine. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 228, 271. 

cessatorops kazubazuae Heinrich. Alta. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 217, 319-320 (5 descr.). 



160 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*chasmodops Heinrich. Maine, Que., Ont., N. H., N. Y., Minn., Man. 

Ichneumon chasmodops Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 223, 230, 259. 9. (S misdet. teste Heinrich, 1962, Pt. VII, 
p. 877). 

citrifrons Cresson. Ont. to N. J., Md., west to 111. and Mich. 

Ichneumon tumidifrons Cresson. Remove from syn. A valid species. 

*conscopa Heinrich. Maine (Dryden). 

Ichneumon conscopa Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 222, 250. ?. 

creperus Cresson. N. W. T., Alaska, B. C. Host: Actebia fennica Tausch. 
Taxonomy: Heinrich, 1961, Pt. Ill, pp. 222, 228, 237-238 (? descr.). 

curticrus (Provancher) . Man. 

deliratorius cinctitarsis Provancher. P. E. I., Ont. to B. C. Question host 

reference. See sup. 1., p. 51. 
Pterocormus stygicus (Cresson) (syn. teste Heinrich, 1961, Pt. Ill, p. 

331). 
Taxonomy: Heinrich, 1961. Pt. Ill, pp. 217, 226, 330-331. 

devinctor (Say). Ont., Alta., N. W. T. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 217, 227, 262-264. 

*diorj'ctriae Heinrich, emend. N. B., Maine, Que., Ont. Host: Dioryctria 
reniculella Grote. 
Ichneumon dioryctiae [sic!] Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 215, 226, 341. 5, ?• 

*eurypiis Heinrich. Que., Ont., N. W. T., Alta. 

Ichneumon eurypus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 216, 325. ?. 

feralis Cresson. Maine, Labrador. Host. IHepialus mustelimis Pack. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 216, 320-321; 1962, Pt. VII, pp. 
879-880 {$ descr.). 

*feriens Heinrich. N. B., Maine., Que., N. H., N. Y., Ont., Mich., Man., Alta., 
B. C, Alaska, Transit, and Canad. Zones. 
Ichneumon feriens Heinrich 1961. Pt. II. Canad. Ent. Sup. 21 [vol. 93] : 
216, 229, 231, 321. $, ?. 

*foxIeei Heinrich. Wyo., B. C, Yukon. 

Ichneumon foxleei Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 219, 309. 5. 

fuscifrons Cresson. 

Taxonomy: Heinrich, 1961. Pt. Ill, pp. 220, 229, 294-297. (5 descr.). 

gestuosus Cresson. Maine, N. H., Ont. Host: Choristoneura fumiferana 
(Clem.). Transf. from Melanichneumon Thomson, p. 285 {teste 
Heinrich, 1959, Ent. News 70 (8): 207). 

*gIaucopygos Heinrich. Maine, Que., Mass., N. Y. 

Ichneumon glaucopygos Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 224, 248. ?. 

*graciIicornops Heinrich. Labrador, Que., Maine, Ont., Man., Sask., Alta. 
Ichneumon gracilicornops Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 220, 278. 9. 

*grandisops Heinrich. Conn., N. Y., Ohio, N. C. 

Ichneumon grandisops Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 218, 305, 9. 

heterocanipae (Cushman). Que., Ont., Pa. 

*heterodon Heinrich. Ont., N. Y. 

Ichneumon heterodon Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 219, 306. 9. 



ICHNEUMONIDAE 161 

*hippisleyae Heinrich. Man. (Dauphin Lake). 

Ichneumon hippisleyae Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 225, 264. ?. 

'^homoioden Heinrich. N. J., Mich. 

Ichneumon homoioden Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 219, 311. ?. 

*homorus Heinrich. N. B., Maine, Que., Ont., Vt., N. Y. 

Ichneumon homorus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 216, 217, 322. ?. 

hospitus Cresson. Maine. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 224, 228, 271-272 (5 ?). 

*hypermelanos Heinrich. Ont. (Sudbury). 

Ichneumon hypermeianos Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 216, 259. 9. 

infidelis (Cresson). Transf. to Orgichneumon Heinrich as syn. of calcatorius 

(Thunberg). 

instabilis (Cresson). Transf. to Thyrateles Perkins. 

Amhlyteles (Pterocormus) winkleyi Viereck. Remove from syn.; a 
valid species in Ichneumon Linnaeus. 

inurbanus Cresson. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 219, 310. 

klagesi Viereck. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 227, 346 (type not found). 

lachrymans Provancher. Maine, N. Y. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 224, 229, 269-271 (9 descr.). 

lactus Brulle. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 222, 231-234 (table to sep. from 
annulatorius F.) . 

lariae labradoris Heinrich. ? N. H. 

lariae lariae Curtis. Greenland, N. W. T. (Boothia Peninsula), Baffin Is. 
Taxonomy: Heinrich, 1961, Pt. Ill, p. 254 (neotype selected). 

*leucocoxalis Heinrich. Maine, Pa. 

Ichneumon leucocoxalis Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 225, 266. 9. 

lewisii Cresson. Transf. from Pseudamblyteles Ashmead, p. 293 (teste Hein- 
rich, 1959. Ent. News 70 (8): 207). Question syn. of fulvopictus 
Ashmead {teste Heinrich, 1961, Pt. Ill, p. 300). 

lividulus (Provancher). Syn. of zelolypus Cresson. 

♦mainensis Heinrich. Maine (Dryden). 

Ichneumon mainensis Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 
880. 9. 

mains Cresson. N. B., Que., Mass., Ont. Host: Lexis bicolor Grote. 

Ichneumon versabilis Cresson. Remove from syn. A valid species. 
Ichneumon (Pterocormus) brittoni Viereck. Remove from syn. and 
transf. to syn. of versabilis Cresson. 

*nielanarcticus Heinrich. Que., Labrador. 

Ichneumon melanarcticus Heinrich, 1961. Pt. III. Canad. Ent. [vol. 931: 
215, 331. 9. 

mendax Cresson. 

Taxonomy: Heinrich, 1961. Pt. Ill, pp. 215, 226, 334-335 ($ descr.). 

♦microferiens Heinrich, Ont., Man., Alta., Alaska, Canad. Zone. 

Ichneumon microferiens Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 216, 323. 9. 



162 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*mystificans Heinrich. Maine, Labrador. 

Ichneumon mystificans Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 227, 342. $. 

*nearctivernus Heinrich. 

Ichneumon compufatorius nearctivernus Heinrich, 1956. Canad. Ent. 78 

(2) : 648. ?. 
Taxonomy: Heinrich, 1961, Pt. Ill, p. 339 (raised to sp. rank). 

nebularum Heinrich. ? N. H. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 217, 226, 317-318 {$ descr.). 

*neoinolitor Heinrich. Alta. (Banff). 

Ichneumon neomolitor Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 217, 329. ?. 

*neosarcitor Heinrich. Maine. 

Ichneumon neosarcitor Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 218, 230, 244. $, 9. 

*nereni emigrator Heinrich. P. E. I., Maine, Que., Ont. /. nereni nereni 
Thomson is European. 
Ichneumon nereni emigrator Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 220, 230, 289. $,9. 

neutralis (Cresson). Transf. Vfxih. its syn. to Melanichneumon subg. 
Vulgichneunion Heinrich as subsp. of subcyaneus Cresson {teste 
Heinrich, 1959. Ent. News 70 (8): 212; 1961, Pt. V, pp. 581, 
610-611). 

nigratorius (Fabricius). Preoccupied by Miiller and replaced by nigriculus n. 
name in Sup. 1, p. 52. 

nigriculus Walkley. Remove from Catalog and Sup. 1; sp. recorded in error 
from Nearctic region. 
Ichneumon nigratorius Fabricius (Catalog, p. 300), preocc. Syn. of 
Amblyjoppa proteus (Christ), a European species (teste Heinrich, 
1961, Pt. Ill, p. 346). 

nigrotergus Davis. 

Taxonomy: Heinrich, 1961, Pt. IV, p. 417 (in text). 

nigrovariegatus (Provancher) . Newfoundland. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 225, 229, 273-275 ($ descr.). 

*obIitus obiitus Heinrich. Maine, Que., Ont. 

Ichneumon obiitus obiitus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 223,228,234. $, ?. 

*oblitus polyglaucos Heinrich. N. Y., S. Dak. 

Ichneumon obiitus polyglaucos Heinrich, 1961. Pt. III. Canad. Ent. Sup. 
21 [vol. 93] : 223, 228, 236. $, ?. 

parvus Cresson. 

?Ichneumon xanthopsis Ashmead. (Syn. questioned by Heinrich, 1961, Pt. 
Ill, p. 345). 

pedalis Cresson. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 226, 313 (text of artemis Vier- 
eck),347. 

*perlugubris Heinrich. Que., Sask. 

Ichneumon perlugubris Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93] : 218, 306. ?. 

peroratus (Cameron). Transf. from Alphabetic List of Unplaced Species, p. 

409 (teste Townes, 1961, Ent. Soc. Wash. Proc. 63 (2) : 112). 

placidus Provancher. Maine, Ont., Utah. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 218, 227, 302 (? descr.). 



ICHNEUMONIDAE 163 

♦polargrus Heinrich. Labrador, Ont., Man., Canad. Zone. 

Ichneumon polargus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 216, 327. ?. 

procax (Cresson). Transf. to Thyrateles Perkins. 

propitius Cresson. Maine, Que., Ont. Transf. from Hoplismenus Graven- 
horst, p. 284 (teste Heinrich, 1959, Ent. News 70 (8) : 207). 

*pseudomaius Heinrich. Mass. (Cambridge). 

Ichneumon pseudomaius Heinrich, 1961. Pt, III. Canad. Ent. Sup. 21 
[vol. 93]: 220, 280. 9. 

*pseudovivax Heinrich. Maine (Dryden). 

Ichneumon pseudovivax Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 221, 286. 9. 

*pseudowinkIeyi Heinrich. Maine. Que., N. Y., Ont. Man., Alta. 

Ichneumon pseudotcinkleyi Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 221, 223, 252. $. 

*pumiliops Heinrich. Que., Ont., Mass., N. Y. 

Ichneumon pumiliops Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 
93]: 225, 276. 9. 

*pusillamoenus Heinrich. Que., Ont. 

Ichneumon pusillamoenus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93] : 224, 272. 9. 

putus Cresson. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 220, 290-291. 

*pycnocephalus Heinrich. B. C, Yukon. 

Ichneumon pycnocephalos (!) Heinrich, 1961. Pt. III. Canad. Ent. Sup. 
21 [vol. 93]: 223 [pycnocephalus] , 261. 9. 

quadrizonatus (Viereck). Syn. of subdolus Cresson. 

restrictus Cresson. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 228, 344-345 (9 prob. known but 
not associated). 

rufiventris (Brulle). Transf. to Thyrateles Perkins, p. 296, as syn. of lugu- 
brator ( Gravenhorst ) . 
Ichneumon semicoccineus Cresson. Remove from syn. A valid sp. 

rufiventris var. incertus (Cresson). Transf. to Thyrateles Perkins, as syn. of 
lugubrator ( Gravenhorst) . 

saucius Cresson. Que. Host: Paranthrene palmii Hy. Edw. Transf. from 
Chasmias Ashmead, p. 292 (teste Heinrich, 1961, Pt. Ill, pp. 212, 
335). 
Chasmias torvinus (Cresson( (syn. teste Heinrich, 1961, Pt. Ill, p. 335). 

semicoccineus Cresson. N. Y., Del., S. C, "Western Terr." Removed from 
syn. of rufiventris (Brulle). A valid sp. {teste Heinrich, 1959. 
Ent. News 70 (8): 207). 
Taxonomy: Heinrich, 1961, Pt. Ill, pp. 227, 310, 311-312. 

similaris Provancher. Transcont. in Canad. Zone. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 217, 226, 313-314 (9 descr.). 

*8tagniphi!os Heinrich. Ont. (Ottawa). 

Ichneumon stagniphilos Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 222, 224, 249. 9. 

stygicus Cresson. Syn. of deliratorius cinctitarsis Provancher. 

subdolus Cresson. N. S., P. E. I., Que., Ont., to N. C, Alta. 

Pterocormus quadrizonatus (Viereck) (syn. teste Heinrich, 1959, Ent. 
News 70 (8): 208). 

*substituens Heinrich. Mass. (Cambridge). 

Ichneumon substituens Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 225, 265. 9. 



164 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

♦suburbanus Heinrich. N. S., P. E. I., Que., Maine, Conn., Mass., Ont., Mich., 
Man., Alta. 
Ichneumon suburbanus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 219,227,307. $, 9. 

*subversatus Heinrich. Que., Maine, Ont, 

Ichneumon subversatus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 221, 283. 9. 

terminatus Provancher. Ont., Maine, Mich. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 224, 267, 268-269. 

*tritus Heinrich. Que., Ont., N. Y. 

Ichneumon tritus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 [vol. 93] : 
221, 224, 256. ?. 

trizonatus Provancher. 

Amblyteles ctenuchae Cushman (syn. teste Heinrich, 1961, Pt. Ill, pp. 
222,230,238 (text), 292). 

truculentus Cresson. Que. 

tumidifrons Cresson. 111. A valid species (teste Heinrich, 1961, Pt. Ill, p. 

224, 267-268). 

ultimus Cresson. Que., Maine, Ont., Md., W. Va., Canad. and Transit. Zones. 
Taxonomy: Heinrich, 1961, Pt. Ill, pp. 222, 229, 257 ($ descr.). 

uncinatus Cresson. Ont., Mich. 

*vaIdopacus Heinrich. Maine (Greenville). 

Ichneumon valtlopacus Heinrich, 1961. Pt. III. Canad. Ent. Sup. 21 
[vol. 93]: 220, 221, 286. 9. 

versabilis Cresson. Atlantic to Cont. Divide in Canad. Zone and cooler part 
Transit. Zone. Hosts: Euphydryas phaeton (Dru.) ; Polygonia 
f annus (Edw.) ; Lycaena hypophlaeas (Bdvl.). A valid species 
(teste Heinrich, 1959, Ent. News 70 (8) : 208). 
Amblyteles (Pterocormus) brittoni Viereck (syn. teste Heinrich, 1961, 
Pt. Ill, pp. 219, 221, 228, 230, 281-283, 284). 

vicinus Cresson. 

Taxonomy: Heinrich, 1961, Pt. Ill, pp. 224, 236-237 (generic placement). 

vivax Cresson. Ont. 

volesus Cresson. N. S., Maine, Que., Ont., Man. 

Amblyteles balteatus Hopper (syn. teste Heinrich, 1959, Ent. News 70 
(8) : 208). Transf. from syn. of Amblyteles rufizonatus (Cresson) 
(teste Heinrich, 1961, Pt. IV, p. 471). 

*walleyi Heinrich. Ont. (Smoky Falls). 

Ichneumon walleyi Heinrich, 1961. Pt. III. Canad. Ent. Sup 21 [vol. 
93]: 215,338. 9. 

winkleyi (Viereck). Que., Maine, Conn., Ont., ?N. B. A valid species (teste 
Heinrich, 1959, Ent. News 70 (8) : 208). 

zelotypus Cresson. Que., Maine, Ont. 

Pterocormus lividulus (Provancher) (syn. teste Heinrich, 1959, Ent. 
News 70 (8) : 209). 

*Genus ORGICHNEUMON Heinrich 

Orgichneumon Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 15 [vol. 
92] p. 15; Pt. II, Sup. 18, p. 95. 

Type: Ichneumon calcatorius Thunberg. By monotypy and 
orig. desig. 
Place in Ichneumonini on p. 301, immediately before (Ichneumonini) = 
Protichneunioni. 

calcatorius (Thunberg). P. E. I., N. S., N. B., Ont., S. C, B. C. Hosts: 
Dasychira plagiata (Wlkr.), Orgyia antiqua (L.), O. leucostigma 



ICHNEUMONIDAE 165 

A. & S. Transf. from Coelichneumon Thomson, Sup. 1, p. 52 

(teste Heinrich 1961 (1960), Pt. I, pp. 15, 95). 
Coelichneumon otiosus (Say) (syn. teste Heinrich, 1956, Canad. Ent. 87 

(11) : 647). 
Pterocormus infidelis (Cresson) (syn. teste Heinrich, 1959, Ent. News: 

211). 
Pterocormus burkei (Viereck) (syn. teste Heinrich, 1956, Canad. Ent. 

87 (11): 647). 

Tribe PROTiCHNEUMONINI 
( =: Ichneumonini of Catalog) 

I am not at all sure that this segregate deserves tribal rank. J. R. T. 
Short can find no larval differences, and the placement of some genera in 
either the Protichneumonini or Ichneumonini is more or less arbitrary. 

Revision: Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 15 [vol. 92]: 
20-87. 

Genus COELICHNEUMON Thomson, Sup. 1, p. 52 
( = /chneumon of Catalog, pp. 301—303) 

*albicoxa Heinrich. Ont., Mich., Wis., N. C. 

Coelichneumon albicoxa Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 
15 [vol. 92]: 34,81. $. 

ater (Cresson). Que., Maine, Vt., Ont., Ohio. 

Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 31, 35, 55-56. S, 9. 

azotus (Cresson). Maine, Que., Ont. 

barnstoni Morley. Ont., Pa., Md., Wis., Alta. In bibliog. ref. change: $, 2. 
to: (^ (^ misdet.). 
Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 29, 34, 54 {$ descr.). 

brunneri (Rohwer). Maine, Ont., Colo. Host: Dioryctria auranticella 
(Grote). 

calcatorius (Thunberg). Transf. to *Orgichneumon Heinrich. 

chalybeus (Cresson). Correct spelling of chalyhaeus in catalog, p. 303. 
Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 29, 50-51. 

citimus (Cresson). N. H. Removed from syn. of germanus Cresson; valid sp. 
{teste Heinrich, 1961 (1960), Pt. I, pp. 35, 81). 

*coIumbianus Heinrich. B.C. (Robson). 

Coelichneumon columbianus Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92]: 28, 38. ?. 

*eximiops Heinrich. Maine, 111., Wis., Iowa, Kans. Host: Achatodes zeae 
(Harr.). 
Coelichneumon eximiops Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 
15 [vol. 92]: 29,33,49. $, 9. 

eximius (Stephens). Hosts: Abagrotis placida Grote, "^Achatodes zeae 
(Harr.). 
Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 29, 33, 47-48 {$ descr.). 

*flagellator Heinrich. Colo. 

Coelichneumon flagellator Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92]: 32, 71. ?. 

*foxleei Heinrich. B. C. (Robson). 

Coelichneumon foxleei Heinrich, 1961 (1960). Pt. I, Canad. Ent. Sup. 
15 [vol. 92]: 29, 52. $. 

germanus Cresson. Syn. of histricus Cresson. 

Ichneumon citimus Cresson. Remove from syn.; a valid species. 

histricus (Cresson). P. E. I., Maine, Pa., Ont., Mich., S. Dak., B. C. 

Ichneumon germanus Cresson (syn. teste Heinrich, 1961 (1960), Pt. 1, 
pp. 31, 34, 66-67). 



166 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

jejunus (Cresson), Que., Ont., Fla., Mont., B. C. 

*leucographus Heinrich. Maine, Ont. 

Coelichneumon leucographus Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92]: 29, 33, 39. 9, $. 

*magniscopa Heinrich. Ont., Maine, Mass., N. Y., N. C, Fla. 

Coelichneumon magniscopa Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92] : 28, 33, 37. $, ?. 

*masoni Heinrich. B.C. (Squamish). 

Coelichneumon masoni Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 
15 [vol. 92] : 28, 43. ?. 

maurus (Cresson). Que., Ont., N. W. T., Alta., B. C, Calif. 

Taxonomy: Heinrich, 1961 (1960). Pt. I, pp. 30, 35, 68-70 {$ descr.). 

navus (Say). Que., Maine, Ont., Kans., Fla. 

*iieocitimus Heinrich. N. W. T., Yukon. 

Coelichneumon neocitimus Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92] : 30, 35, 79. $, 9. 

*neopretatiis Heinrich. N. Y., Maine, Que., Ont., Sask., ?Kans. Transit. Zone. 
Coelichneumon neocretatus Heinrich, 1961 (I960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92] : 29, 35, 51. $, ?. 

*neomexicanus Heinrich. N. W. (Manzano National Forest). 

Coelichneumon neomexicanus Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol.92]: 29,40. 9. 

nigraticolor (Viereck). Ont., Maine, Pa. 

Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 30, 34, 64-65 ($ descr.). 

nigrosignatus (Viereck). Transf. from Alphabetic List of Unplaced Species, 
p. 408 (teste Heinrich, 1961 (1960), Pt. I, p. 82). 

*nudus Heinrich. Ga. (Summerville). 

Coelichneumon nudus Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 
15, [vol. 92] : 29, 43. 9. 

*obscuratus Heinrich. B.C. (Robson). 

Coelichneumon obscuratus Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92]: 29, 45. 9. 

orpheus Cresson). Maine, Que. Host. "Notodontidae pupa". 

Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 30, 35, 61-63 {$ descr.). 

otiosus (Say). Transf. to *Orgichneumon Heinrich. 

pepticus (Cresson). Colo., Oreg., Wash. 

Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 32, 35, 63-64 (9 descr.). 

pequoitorunt (Viereck). Syn. of pervagus (Cresson). 

pervagus (Cresson). N. S., Maine, Labrador, B. C. Host: Scoliopteryx liba- 
trix L. 
Ichneumon pequoitorum (Viereck) (syn. teste Heinrich, 1959, Ent. 
News 70 (8): 207). 

*phaenomenon Heinrich. Maine, Alta., Alaska. 

Coelichneumon phaenomenon Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92] : 28, 32, 35. $, 9. 

*pomiHoaeneus Heinrich. Maine (Dryden). 

Coelichneumon pomilioaeneus Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92] : 29, 50, 87. 9 [spelled pomiloaeneus in key, p. 
29]. 

pulcher (Brulle). Maine, Ont. 

Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 28, 33, 46-47 (5 descr.). 

*puIcherior Heinrich. Ariz., Colo., B. C. 

Coelichneumon pulcherior Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92] : 28, 36. 9. 



ICHNEUMONIDAE 



167 



Ufens plicatus 
I. Canad. Ent. 

I. Canad. Ent. 



pumilionobilis Heinrich. Ont. 

*punctifer Heinrich. N. Y., Md., Va., Tex. 

Coelichneumon punciifer Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 
15 [vol. 92]: 31, 34, 77. $, 9. 

*quadraticeps Heinrich. Md., Va., W. Va., Mich. 

Coelichneumon quadraticeps Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92] : 31, 34, 78. $, 9. 

*rubricoxa Heinrich. Calif. 

Coelichneumon rubricoxa Heinrich, 1961 (1960). Pt. I. Canad. Ent. 

Sup. 15 [vol.92]: 32,72. $, 9. 

*rubroaeneus Heinrich. Calif. (Lake Almanor). Host: 
Grote. 
Coelichneumon rubroaeneus Heinrich, 1961 (1960). Pt. 
Sup. 15 [vol. 92]: 29, 43. $. 

*sassacoides Heinrich. B. C. 

Coelichneumon sassacoides Heinrich, 1961 (1960). Pt. 
Sup. 15 [vol.92]: 28,42. ?. 

sassacus (Viereck). Que., Maine, N. H., Ont., ?Calif. 

Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 28, 33, 40-42. 

seniilaevis (Cresson). Que., N. W. T., B. C, Calif. 

Taxonomy: Heinrich, 1961 (1960), Pt. I, pp. 32, 72-73 (5 descr.). 

*similior Heinrich. Ont., Maine. 

Coelichneumon similior Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 
15 [vol. 92]: 29,44. $. 

solutus occidentalis Roman. Transf. to *Coelichneumonops as a valid sp. 

tauma Heinrich. Ont., Que., N. Y., Minn. Hosts: Choristoneura fumiferana 
Clemson; Lexis bicolor Grote. 

*terebratus Heinrich. Colo. 

Coelichneumon terebratus Heinrich, 1961 (1960), 
Sup. 15 [vol. 92] : 30, 70. 9. 

*terebrifer Heinrich. Va. 

Coelichneumon terebrifer Heinrich, 1961 (1960). 
Sup. 15 [vol.92]: 36,71. 9. 

viola Cresson). Ont., Fla., Tex. 

vitalis (Cresson). Maine, N. H., Md., Va., N. C, Ind., Utah. 

*walleyi Heinrich. Ont., B. C. 

Coelichneumon walleyi Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 
15 [vol. 92]: 31, 34, 77. 5, 9. 

*Genus COELICHNEUMONOPS Heinrich 

Coelichneumonops Heinrich, 1958. Canad. Ent. 90 (12) : 739. 

Type: * (Coelichneumonops vockerothi Heinrich) = solutus 
(Holmgren). Orig. desig. 
Place in Catalog on page 303 after Coelichneumon Thomson. 

*cashmani Heinrich. Yukon (Firth R., 69° 30'N, 139° 31'W). 

Coelichneumonops cashmani Heinrich, 1958. Canad. Ent. 90 (12) : 740. 
9. 

occidentalis (Roman), changed status. Que., Baffin Is. Elevated to specific 
rank (teste Heinrich, 1961 (1960), Pt. I, pp. 83, 85. Transf. from 
Coelichneumon, Sup. 1, p. 52 (teste Heinrich, loc. cit. supra). 

*so!ulus (Holmgren). Que. 

Ichneumon solutus Holmgren, 1895. Ichn. Suecica I. r>. 145. 9. 
^Coelichneumonops vockerothi Heinrich. 1958. Canad. Ent. 90 C12) : 
739. 5, 9. Syn. (teste Heinrich, 1961 (1960). Pt. I. [vol. 921: 82. 
83). 



Pt. I. Canad. Ent. 



Pt. I. Canad. Ent. 



168 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*walkleyae Heinrich. B. C. (Vancouver) . 

Coelichneumonops walkleyae Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92] : 83, 85. 9. 

Genus PROTICHNEUMON Thomson, p. 304 

*effigies Heinrich. Newfoundland to B. C, Maine, Pa., Mich., Kans., S. Dak., 
Colo., Mont. 
Protichneumon effigies Heinrich, 1961 (1960). Pt. I. Canad. Ent. Sup. 
15 [vol. 92]: 22, 26. S, ?• 

grandis grandis (Brulle), changed status. Canad. and Transit. Zones. Re- 
duced to subsp. rank {teste Heinrich, 1961 (1960), Pt. I, pp. 22, 
25). 
Ichneumon regnatrix Cresson. Remove from syn. A valid subsp. 

grandis regnatrix (Cresson). Pa., Del., Ga., Fla., La., Kans. A valid subsp. 
(teste Heinrich, 1961 (1960), Pt. I, pp. 22, 25). 

^grandis victoriae Heinrich. B. C. 

Protichneumon grandis victoriae Heinrich, 1961 (1960). Pt. I. Canad. 

Ent. Sup. 15 [vol. 92] : 22, 25. $, 9. 

*polytropus Heinrich. N. C. (Boardman). 

Protichneumon polytropus Heinrich, 1961 (1960). Pt. I. Canad. Ent. 
Sup. 15 [vol. 92]: 22, 27. $ [misspelled polytropos in key). 

Genus AMBLYJOPPA Cameron, p. 340 

magna (Cresson). 

Taxonomy: Heinrich, 1961 (1960), Pt. I, p. 27. 

*Genus CATADELPHOPS Heinrich 

Catadelphops Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 863. 

Type: Tragus buccatus Cresson. By monotypy and orig. desig. 

Place on p. 304 of Catalog after Amblyjoppa Cameron. 

buccatus (Cresson). Transf. from Catadelphus Wesmael (teste Heinrich, 
1962, Pt. VII, p. 864). 

*Genus NEAMBLYJOPPA Heinrich 

Neamblyjoppa Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 865. 

Type: Neamblyjoppa nasuta Heinrich. By monotypy and orig. 
desig. 

Place on p. 304 of Catalog after * Catadelphops Heinrich. 

*na8uta Heinrich. Ariz. 

Neamblyjoppa nasuta Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 
866. $. 

Tribe TROGINI 
Revision: Heinrich, 1962. Pt. VII, Canad. Ent. Sup. 29: 807-860. 

*Subtribe CALLAJOPPINA 

Genus CATADELPHUS Wesmael, p. 304 

atrox (Cresson). Okla. Host: Proserpinus juanita Sikr. 

atrox var. marginipennis (Cresson). Elevated to specific rank. 

marginipennis (Cresson) , new status. Alta. 

Catadelphus atrox var. marginipennis (Cresson) of Catalog. 

*ochraceus Heinrich. Okla. 

Catadelphus ochraceus Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 
811,813. S. 

semiruber Hopper. B. C, Wyo. 



ICHNEUMONIDAE 169 

♦Genus CALLAJOPPA Cameron 

Callajoppa Cameron, 1903, Entomologist 36: 236. 

Type: (Callajoppa bilineata Cameron) = [Ichneumon'] cirro- 
gaster (Schrank). By monotypy. 

Place after Catadelphus Wesmael, p. 304. 

*cirrogaster (Schrank). ?Ont. ; Europe; Japan. 

Ichneumon cirrogaster Schrank, 1781. Enum. Ins. Austriae, p. 348. 
*Ichneumon lutorius Fabricius, 1787. Mant. Ins. I, p. 262. 
"^Callajoppa bilineata Cameron, 1903. Entomologist 36: 236, 9. 
Taxonomy: Heinrich, 1962, Pt. VII, pp. 814-815. 

Genus GNAMPTOPELTA Hopper, p. 304 
obsidianator obsidianator (Brulle). Que., Man. 

Genus TRICYPHUS Kriechbaumer, pp. 304-305 

aler Hopper. Fla. 

*elegans floridanus Heinrich. Fla. 

Tricyphus elegans floridanus Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 

29: 840,842. $. 

Genus TMETOGASTER Hopper 
( = Conoca/ama subgenus Tmefogasfer Hopper, p. 305) 
Raised to generic rank (teste Heinrich, 1962, Pt. VII, p. 836). 
nubilipennis (Haldemann). Maine. Host: lAmpelocea myron Cresson. 

Genus CONOCALAMA Hopper 

(r:Conoca/ama subgenus Conoca/ama Hopper, pp. 305-306) 

Raised to generic rank (teste Heinrich, 1962, Pt, VII, p. 815). 

nolteri (Cresson), changed status. Raised to sp. rank (teste Heinrich, 1962, 
Pt. VII, pp. 816, 818, 827; 9 discussed). 

brullei (Cresson). Hosts: Dolba hylaeus Drury, Paonias astylus Druiy, P. 
myops S. & A., Phlegethontius sexta Joh. 

canadensis (Provancher) . Atlantic to Cont. Divide in Transit., U. Austr. 

Zones. Hosts: Paonias myops S. &. A. Removed from syn. of C. 

copei (Cresson) ; a valid sp. (teste Heinrich, 1959, Ent. News 70 

(8): 206). 

Conocalama copei var. nigrocinctus (Hasey) (syn. teste Heinrich, 1962, 

Pt. VII, p. 822). 

*catalinarum Heinrich. Ariz. (Mt. Lemmon). 

Conocalama catalinarum Heinrich, 1957. Canad. Ent. 89 (7) : 334. $. 
Taxonomy: Heinrich, 1962, Pt. VII, pp. 817, 835-836. 

copei copei (Cresson), changed status. Reduced to subsp. rank (teste Hein- 
rich, 1962, Pt. VII, p. 821). 
Trogus canadensis Provancher. Remove from syn. Valid sp. 

*copei heteropteryx Heinrich. Maine, Que., Ont. 

Conocalama copei heteropteryx Heinrich, 1962. Pt. VII. Canad. Ent. 
Sup. 29: 816, 822. $, ?. 

copei var. nigrocinctus (Provancher). Syn. of canadensis (Provancher). 
fuscalata Hopper. Ont. 

*manitobae Heinrich. Man. (Aweme). 

Conocalama manitobae Heinrich, 1962. Pt. VII. Canad. Ent. Sup. 29: 
817,834. 5. 

mellosa mellosa (Cresson). N. Mex., Colo. Transf. from subsp. of occiden- 
taiis (Cresson). 
Taxonomy: Heinrich, 1962, Pt. VII, pp. 817, 818, 829, 830 (relationships 
discussed; 5 neallotype). 



170 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

mellosa rilevi (Cresson), changed status. Reduced to subsp. rank {teste 
Heinrich, 1962, Pt. VII, pp. 817, 818, 830; $ neallotype). 

occidentalis (Cresson), changed status. West. No. Amer. Host: Smerinthus 
cerisyi Kby. Raised to sp. rank {teste Heinrich, 1962, Pt. VII, p. 
823). 

fConocalama occidentalis occidentalis var. luteata Hopper (syn. teste 
Heinrich, 1962, Pt. VII, p. 823). 

Taxonomy: Heinrich, 1962, Pt. VII, pp. 823-825. 

occidentalis bolteri (Cresson). Given sp. rank. 

Tragus quebecensis Provancher. A valid species. 

occidentalis mellosa (Cresson). Now recognized as mellosa mellosa (Cres- 
son). 

quebecensis (Provancher). N. S., Maine, Que., Ont., Minn., Man., B. C. 
Host: lEnclcs imperialis Drury. Removed from syn. of occiden- 
talis bolteri (Cresson) ; a valid sp. {teste Heinrich, 1962, Pt. VII, 
pp. 817, 818,826). 

rileyi (Cresson). Subsp. of mellosa (Cresson). 

*Subtribe TROGINA 

Genus MACROJOPPA Kriechbaumer, p. 306 

Taxonomy: Heinrich, 1962, Pt. VII, pp. 845-846 (doubts of gen. syn. 
expressed). 

Genus TROGUS Panzer, p. 306 

Trogus Panzer. Under type citation change [Ichneumon'\ var. coerula- 

tor (Fabricius) to: [Ichneumon] lapidator coerulator (Fabri- 

cius). 
Dinotomus Foerster. Change type citation to: 

Type: Ichneumon lapidator Fabricius. Desig. by Ashmead in 

Smith, 1900. 
Psilomastax Tischbein. Remove from syn. A valid genus containing no 

Nearctic spp. {teste Heinrich, 1962, Pt. VII, pp. 844, 846, in text). 

edward«ii Cresson. 

Taxonomy: Heinrich, 1962, Pt. VII, pp. 848, 852-853 (prob. subsp.). 

flavipennis Cresson. Wyo. 

fulvipes Cresson. North. Transit. Zone. 

*lapidator coerulator (Fabricius). Yukon; Europe. T. lapidator lapidator 
(Fab.) is Palaearctic. Host: Papilio machaon L. 
Ichneumon coerulator Fabricius, 1804. Systema Piezatorum, p. 68, no. 

79. 
Taxonomy: Heinrich, 1962, Pt. VII, pp. 847, 848-849. 

pennator (Fabricius). Host: Papilio polyxenes asterias Putnam. 

*Dinotomus vulpinus var. nigrithorax Strand, 1912. Arch. Naturg. 

Jahrg. 78, Abt. A (6) : 74. $. (Syn. teste Townes, 1961. Ent. Soc. 

Wash. Proc. 63 (2) : 105.) 
Biology: Heinrich, 1962, Pt. VII, pp. 849, 850 (notes on hosts). 

Subfamily LISSONOTINAE 

Tribe GLYPTINI 
Genus GLYPTA Gravenhorst, pp. 306-309 

fumiferanae (Viereck). Utah, Colo., Ariz. Hosts: Dioryctria reniculella 
(Grt.), Sciaphila duplex (Wish.). 
Biology: Dodge, 1961. Canad. Ent. 93 (3) : 222-228. 



ICHNEUMONIDAE 171 

Tribe LISSONOTINI 

Genus ALLOPLASTA Foerster, p. 310 
( = Amers/b/a of Catalog, p. 310) 

Amersibia Auctt., nee Foerster (see Lampronola Curtis). 
*Trysicampe Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 207 
(syn. by Townes, Townes, and Gupta, 1961, p. 206; Perkins, 1962, 
p. 461). 

Type: Ichneumon piceator Thunberg. Desig. by Perkins, 
1962. 
*Trichopimpla Cameron, 1903. Ztschr. f. Hym.-Dipt. 3 (5) : 303 (syn. 
teste Morley, 1913, Fauna Brit. India, Hym. vol. 3, pt. 1, p. 222; 
Townes, Townes and Gupta, 1961, p. 205-206). 

Type: Trichophupla pilosa Cameron. By monotypy. 

bullata (Davis), n. comb. Transf. from Amersibia Foerster, p. 310; change 
vol. no. of bibliog. ref . from 21 to 22. 

superba (Provancher), n. comb. Transf. from Amersibia Foerster, p. 310. 

Genus Amersibia Foerster, p. 310 

Amersibia Foerster. Add pp. 214, 218 to bibliog. citation. Change type 
desig. to Ichneumon catenator Panzer. By substitution of Amersi- 
bia for Meniscus Schiodte. Amersibia was proposed by Foerster 
as a new name for Meniscus Schiodte which he considered preocc. 
by the plant name Meniscium (teste Foerster's handwritten MS; 
Schmiedeknecht, 1888, Zool. Jahrb. 3 (3) : 439). Syn. of Lampron- 
ola Cui'tis, p. 311. 

Alloplasta Foerster. Remove from syn. A valid genus (teste Perkins, 
1962, p. 403). 

bullata (Davis). Transf. to Alloplasta Foerster, p. 310. 

superba (Provancher). Transf. to Alloplasta Foerster, p. 310. 

Genus LAMPRONOTA Curtis, p. 31 1 

Meniscus Schiodte. Delete type citation. Add: Type: Ichneumon catena- 
tor Panzer. Desig. by Viereck, 1914. (Ichneumon setosus Holm- 
gren was not an orig. inch sp.). 

Amersibia Foerster. Proposed as n. name for Meniscus (erroneously be- 
lieved preocc). 

*Ctenopimpla Cameron, 1899. Manchester Lit. and Phil. Soc. Mem. and 
Proc. 43 (3) : 189. Preocc. (Syn. teste Townes, Townes, and 
Gupta, 1961, p. 204.) 

Type: Ctenopimpla alhomaculata Cameron. By monotypy. 

*Adelopimpla Schulz, 1906. Spolia Hym., p. 116. N. name for Cteno- 
pimpla. 

LISSONOTA subgenus LISSONOTA Gravenhorst, pp. 311-312 
bibrevicincla (Davis). Change vol. no. in bibliog. ref. from 21 to 22. 
occidentalis (Viereck). Change page no. in bibliog. ref. from 83 to 84. 

Genus PIMPLOPTERUS Ashmead, pp. 312-314 

argyreslhiae Walley. N. S., N. B., Que., Ont. Host: Argyresthia laricella 
Kft. 
Pimplopterus argyreslhiae Walley, 1961. Canad. Ent. 93 (1) : 1. 5, $• 

davisi Townes. 

Lampronota angusta Davis. Change vol. no. in bibliog. ref. from 17 to 

22. 

electra (Viereck) . Change page no. in bibliog. ref. from 83 to 84. 

fenellus (Viereck). 

Lissonota fenella Viereck. Change page no. in bibliog. ref. from 82 to 83. 



172 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Harrimaniella minor Viereck. Change page no. in bibliog. ref. from 86 to 
87. 

palloranus (Davis). Change vol. no. in bibliog. ref. from 21 to 22. 

parvus (Cresson). Host: Dioryctria reniculella (Grt.). 

sexcinctus (Ashmead). Does not belong in Pimplopterus; a revisionary study 
of Lissonotini necessary for placement. 

Genus CRYPTOPIMPLA Taschenberg, pp. 314-315 

Xenacis Foerster. Transf. from syn. of Cylloceria Schiodte, p. 342 (teste 
Perkins, 1962, p. 462). 

Genus Asphragis Foerster, p. 315 

Asphragis Foerster. Type-species misdet. Transf. to syn. of Cylloceria 

Schiodte, p. 342. 

michiganensis (Davis). Transf. to *Campociiieta Schmiedeknecht. 
mirabilis (Cresson). Transf. to *Campocineta Schmiedeknecht. 
subflavus (Davis). Transf. to *Campocinela Schmiedeknecht. 

*Genus CAMPOCINETA Schmiedeknecht 

Asphragis of authors not of type-species (misdet. by authors). 
Cainpocineta Schmiedeknecht, 1900. Zool. Jahrb. Abt. f. System. 13 (4) : 

301, 306, 309, 343. (See Townes, Townes, and Gupta, 1961, p. 

206). 
Type: Campocineta varicoruis Schmiedeknecht. By monotypy. 

Replaces Asphragis of authors, p. 314. 

michiganensis (Davis), n. comb. Transf. from Asphragis Foerster, p. 315. 

mirabilis (Cresson), n. comb. Transf. from Asphragis Foerster, p. 315. 

subflavus (Davis), n. comb. Transf. from Asphragis Foerster, p. 315. 

Genus SYZEUCTUS Foerster, p. 315 

Diceratops Foerster. In type citation after "Included" add, and desig- 
nated. 

Syzeuctus Foerster. In type citation after "Included" add, and designat- 
ed. 

Syzeucta Thomson. Delete type citation. Add : Emend. 

*L€ptoglyphis Brethes, 1927. Ent. Mitt. 16: 319 (syn. teste Townes, 
Townes, and Gupta, 961, p. 207). 

Type: Leptoglyphis minasensis Brethes. By monotypy. 

*Paratanera Rao, 1953. Indian Forest Rec. (n. s.) Ent. 8 (8) : 171 (syn. 
teste Townes, Townes, and Gupta, 1961, p. 207), 
Type: Paralanera indica Rao. Orig. desig. 

Tribe BANCHINI 

Genus EXETASTES Gravenhorst, pp. 316-319 

*Semnophrys Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 158 
(syn. teste Perkins, 1962, p. 451). 

Type: Exetastes notatus Holmgren. Desig. by Perkins, 1962. 

''Icyona Cameron, 1903. Ztschr. Syst. Hym. Dipt. 3: 340 (syn. teste 
Townes, Townes, and Gupta, p. 210). 

Type: (Icyona rufipes Cameron) = Exetastes cameroni 
Townes. By monotypy. 
*Rhynchexetastes Cameron, 1906. So. African Mus. Ann. 5: 102 (syn. 
teste Townes, Townes, and Gupta 1961, p. 210). 

Type: Rhynchexetastes violaceipennis Cameron. By mono- 
typy. 



ICHNEUMONIDAE 173 

Tegona Morley. Delete question-mark. Change type citation to: {Tegona 
rufipes Morley) = Exetastes canieroni Townes. 

pictus Cushman. Ariz, 

Subfamily CTENOPELMATINAE 
(zzScolobatinae of Catalog and A4e5o/e//nae of Sup. 1) 

Tribe EUCERATINI 

Genus EUCEROS Gravenhorst, pp. 321-322 

*Tautozeliis Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 212 
(syn. teste Perkins, 1962, p. 457). 

Type: (Euceros egregius Holmgren) = Euceros serricornis 

Haliday. Desig. by Perkins, 1962. 

frigidus Cresson. Que., N. H., N. Y. Hosts: Endasys subclavatus (Say), 
Laniaclms sp., Neodiprion lecontei (Fitch), TV. pratti banksianae 
Roh., Olesicamj)e lopliyri (Riley). 
Biology: Finlayson, 1960. Canad. Ent. 92 (1) : 34-36.— Tripp, 1961, Can- 
ad. Ent. 93 (1) : 40-58. 

thoracicus Cresson. B.C. Host: Melanolophiaimitata Wlkr. 

Tribe CATOGLYPTINI 
{zzPionini of Catalog) 

Genus TREMATOPYGUS Holmgren, p. 322 

* Ainorphognathon Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 
203 (syn. teste Perkins, 1962, p. 404). 

Type: Tryphon melanocerus Gravenhorst. Inc. by Kriech- 
baumer, 1897. 
Epachthes Foerster. Preocc. Remove from syn. (An isogenotypic syn. 

of Azelus Foerster with no Nearctic spp.) . 
*RhigeIus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 208 (syn. 
teste Perkins, 1962, p. 449). 

Type: (Trematopygus kriechbaumeri Thomson) = melanoc- 
erus (Gravenhorst). Desig. by Perkins, 1962. 
*Campoporus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25. 210 
(syn. teste Perkins, 1962, p. 412). 

Type: Ichneumon dictator Thunberg. Desig. by Perkins, 
1962. 
Epachtus Thomson. Emend. Remove from syn. 

Genus PION Sch!odte, pp. 323-324 

Sympherta Foerster. In type citation change "desig." to: Inch by Davis, 

1897. 
IStiphrosomus Foerster. Perkins, 1962, pp. 453, 458, considers this a 

syn. of Trapezocora Foerster. 
*1Trapezocora Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 208. 
Considered valid genus by Perkins, 1962, p. 458. 

Type: Mesoleptus antilope Gravenhorst. Desig. by Perkins, 
1962. 
*?Atrestes Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 209. Con- 
sidered a syn. of Trapezocora by Perkins, 1962, pp. 408, 458. 

Type: *Catoglyptus (Stiphrosomus) sulcatus Thomson. Desig. 
by Perkins, 1962. 
*?Campogenes Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 209. 

Type: (Tryphon antilope Gravenhorst) = Trapezocora anti- 
lope (Gravenhorst) . Desig. by Perkins. 1962. 



174 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

lEustiphrosomus Hincks. 

A revision will probably show several segregates here, or a single heter- 
ogeneous group. 

Tribe CTENOPELMATINI 

Genus CTENOPELMA Holmgren, pp. 324, 325 

Several undescribed Nearctic spp. belong here, 

XENOSCHESIS subgenus XENOSCHESIS Foerster, p. 325 

Polycinetus Thomson. Delete type citation. Transf. to syn. of subg. Po- 
lycinetis Foerster as emend. 

XENOSCHESIS subgenus POLYCINETIS Foerster, p. 325 

Polycinetus Thomson. Emend. Preocc. Transf. from syn. of subg. Xen- 
oschesis Foerster. 

Tribe MESOLEIINI 

Genus PERILISSUS Holmgren, pp. 326-327, Sup. 1, p. 55 

*Ichnaeops Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 197. 

Type: Perilissus lutescens Holmgren. Incl. and desig. by Per- 
kins, 1962. 
*Exacrodus Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 210 (syn. 
teste Townes, Townes, and Gupta, 1961, pp. 213-214). 

Type: (Exacrodus populans Morley) = cingulator (Morley). 
Incl. by Morley, 1913. 
*Pseudochorus Rao, 1953. Indian Forest Rec. (n.s., Ent.) 8: 195. (syn. 
teste Townes, Townes, and Gupta, 1961, pp. 213-214). 

Type: (Pseudochorus kuriani Rao) = cingulator Morley. 
Orig. desig. 

Genus LABROSSYTA Foerster, p. 328 

Labrossyta Foerster. Change type citation to: Ichneumon scotopterus 

Gravenhorst. Incl. by Thomson, 1894. 
*Labrossytus Thomson, 1894. Opusc. Ent., fasc. 18, p. 2001. Emend. 

Genus Laphyroscopus Foerster, p. 328 

Laphyroscopus Foerster. Delete type citation. Syn. of Lathrolestes Foer- 
ster, p. 328. 

capitatus (Cresson). Transf. to *Nanium Townes, n. gen. 

longipes (Davis). Transf. to *ZapIethocornia Schmiedeknecht. 

Genus LATHROLESTES Foerster, pp. 328-329 

Lathrolestes Foerster. Change type citation to: Tryphon clypeatus Zet- 
terstedt. Desig. by Viereck, 1914. (See Perkins, 1962, p. 435.) 

Laphyroscopus Foerster (syn. teste Townes, Townes, and Gupta, 1961, 
pp. 214-215; Perkins, 1962, p. 434). Change type citation to: 
{Tryphon gorskii Ratzeburg) = luteolator Gravenhorst. Desig. by 
Viereck, 1914. 

Lathrolestus Thomson. Delete type citation. Add : Emend. 

Luphyroscopus Thomson. Delete type citation. Add : Emend. 

♦Genus NANIUM Townes, n. gen. 
( = Laph/roscopus of authors) 

Type: Tryphon capitatus Cresson. Orig. desig. 
The genus is one proposed by Townes (MS) for Laphyroscopus capitatus 
(Cresson), invalid type desig. for Laphyroscopus Foex'ster. His description 
follows : 



ICHNEUMONIDAE 175 



Genus NANIUM 



Front wing 3.5 to 5.4 mm. long; clypeus separated from face by an 
impression, its profile weakly convex and apex broadly arcuate; mandibular 
teeth equal; junction of occipital and hypostomal carinae some distance 
above base of mandible; lateral ocellus separated from eye by about 2.5 its 
diameter; first flagellar segment about 1.55 as long as second; mesopleurum 
with a median longitudinal groove on only its hind 0.25 ± ; notaulus faint 
and rather short, or absent. Propodeum without carinae, its profile rather 
weakly convex; tarsal claws short, sparsely pectinate; areolet absent; me- 
diella strongly arched; nervellus vertical or inclivous, broken below the 
middle or sometimes not broken; first tergite rather short and broad, its 
dorsal profile weakly convex or almost flat, spiracle at the middle, its base 
without a dorsal pit, its dorso-lateral carina present to apex of glymma and 
sometimes present (but weak) between spiracle and apex; glymma rather 
short, deep, separated from glymma on opposite side by a translucent parti- 
tion; thyridium absent; sides of tergites 2 and 3 curved under, at least in 
female not clearly separated from their rather narrow epipleura; male sub- 
genital plate simple; tip of penis somewhat swollen, moderately decurved, its 
apex rounded; ovipositor sheath 0.35 to 0.6 as long as apical depth of abdo- 
men, moderately wide; ovipositor straight, moderately stout, with a strong 
subapical dorsal notch; cercus like the head of a mushroom, much wider than 
long. 

Type : Tryphon capitatus Cresson. 

This is an American genus, with the type-species in the eastern part of 
the Nearctic Region, and an undescribed species each in Mexico and in Chile. 

The generic name is from vaviov (puppet), in reference to the small 
size. 

capitatuni (Cresson), n. comb. Transf. from Laphyroscopus Foerster, p. 328. 

♦Genus ZAPLETHOCORNIA Schmiedeknecht 

Zaplethocornia Schmiedeknecht, 1912. Opusc. Ichn. fasc. 32, p. 2519. 

Type: Ichneumon procurator Gravenhorst. Incl. by Schmie- 
deknecht, 1913. 

Place on p. 329 before Synoecetes Foerster. 

longipes (Davis), n. comb. Transf. from Laphyroscopus Foerster, p. 328 
{teste Townes in litt.) . 

Genus SYNOECETES Foerster, p. 329 
Polyrhysius Thomson. Delete type citation. Add : Emend. 

♦Genus ESIGNA Perkins 
( = Gnes/a Foerster, p. 330) 

Esigna Perkins, 1962. British Mus. (Nat. Hist.) Bui. Ent. 11 (8) : 425. 
New name for Gnesia Foerster, 1868, preocc. by Doubleday, 1848. 
Type: Gnesia caliroae Rohwer. By substitution of Esigna for 
Gnesia, 

rotundata (Davis), n. comb. 

Genus PROTARCHUS Foerster, p. 330 

Otlophorus Foerster. Remove from syn. and from Catalog. A valid 
genus {teste Perkins, 1962, p. 441) with no descr. Nearctic spp. 

Neales Foerster. Isogenotypic syn. of preceding. Remove from Catalog. 

Protarchoides Cushman. A revision may prove this is a distinct but 
closely related genus. 

Genus HIMERTA Foerster, pp. 330-331 

Himerta Foerster, 1868. Change type citation to Euryproctus (Him- 

ertus) bisannulalus Thomson. 
Clepsiporthus Foerster. Change type citation to (Mesoleptusl ruhigitu 

osus Davis) = Himerta ruhiginosa (Cresson). 



176 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Dolioctonu8 Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25 : 212. 

Type: Ichneumon defectivus Gravenhorst. Incl. and desig. by 
Perkins, 1962. 
Himertus Thomson. Delete type citation and add: Emend. 
*Daisetsuzania Uchida, 1930. Hokkaido Univ., Fac. Agr., Jour. 25: 289 
(syn. teste Townes, 1957, Proc. Ent. Soc. Wash. 59 (3) : 104). 

Type : Daisetsuzania alhifrons Uchida. By monotypy and orig. 
desig. 

annulata (Davis). Change vol. no. in bibliog. ref. from 21 to 22. 

Genus MESOLEIUS Holmgren, pp. 331-334, Sup. 1, pp. 55-56 

Alexeter Foerster. Emend, type citation as follow^s: (Mesoleptus (Alex- 
eter) ruficornis Gravenhorst) = sectator (Thunberg), 1822. Per- 
kins, 1962, p. 403, considers this a valid genus. 
Lagarotis Foerster. Considered a valid genus by Perkins, 1962, p. 434. 
*Zemiophron Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 206 (syn. 
teste Townes, Townes, and Gupta, 1961, p. 215). Perkins, 1962, p. 
464, considers this a syn. of Alexeter Foerster. 

Type : Mesoleius laevissimus Strobl. Desig. by Perkins, 1962. 
Oneista Foerster. Perkins, 1962, p. 440, considers this a syn. of Lagarotis 

Foerster. 
*Dysantes Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 207 (syn. 
teste Townes, Townes, and Gupta, 1961, p. 215). Considered syn. 
of Lagarotis Foerster by Perkins, 1962, p. 419. 

Type: Ichneumon debitor Thunberg. Desig. by Perkins, 1962. 
Aeolometis Foerster. (Valid genus, teste Perkins, 1962, p. 402). 
Scopcsis Foerster. (Valid genus, teste Perkins, 1962, p. 451). Add to 

type citation: = bicolor (Gravenhorst). 
Tachyporthus Foerster. Considered syn. of Aeolometis Foerster by Per- 
kins, 1962, p. 456. 
*Scoparches Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 211 
(syn. teste Townes, Townes, and Gupta, p. 215). Considered syn. 
of Scopesis Foerster by Perkins, 1962, p. 451. 

Type : Tryphon bicolor Gravenhorst. Desig. by Perkins, 1962. 
*Allocritus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 211 (syn. 
teste Townes, Townes and Gupta, p. 215; Perkins, 1962, p. 403). _ 
Type: Mesoleius tenuiventris Holmgren. Desig. by Perkins, 
1962. 
Spudaea Foerster. Considered syn. of Rhinotorus Foerster by Pei'kins, 
1962, p. 452. Change type citation to: Tryphon leucostomus Gra- 
venhorst. Desig. by Viereck, 1914. {See Perkins, 1962, p. 452.) 
*Rhinotorus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 211 
(syn. teste Townes, Townes, and Gupta, p. 215). Considered valid 
genus by Perkins, 1962, p. 450. 

Type: Spudaea longicornis Schmiedeknecht. Desig. by Per- 
kins, 1962. 
*Campodoru8 Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 213 
(syn. teste Townse, Townes, and Gupta, 1961, p. 215; Perkins, 
1962, p. 412). 

Type: Mesoleius melangaster Holmgren. Desig. by Perkins, 
1962. 
Spudaeus Thomson. Delete type citation. Add: Emend. 
Lagarotus Thomson. Delete type desig. Add : Emend. 
Scopesus Thomson. Delete type desig. Add : Emend. 
Aelomeiis Thomson. Delete type desig. Add : Emend. 

A revision is necessary before the Nearctic species can be placed in the 
segregates comprising this genus. 

nigropictus (Davis). 

Lampronota nigropicta Davis. Change vol. no. in bibliog. ref. from 21 to 
22. 



ICHNEUMONIDAE 177 

tenthredinis Morley. Pa., Mich. 

Biology: Finlayson, 1960. Canad. Ent. 92 (12): 933-934 (larval).— 
Drooz, 1961. Canad. Ent. 93 (9): 804-807 (host immunity). 

Genus lAMACHUS Foerster, pp. 334-335 
angularius (Davis). Idaho. Host: Neodiprion sp. nr. scutellatus Roh. 
contortionis Davis. Ky. Host: Neodiprion lecontei (Fitch). 

Genus SAOTIS Foerster, p. 335 
Saotus Thomson. Add 'Emend.' after bibliog. ref. Delete type desig. 

Tribe EURYPROCTINI 

Genus SYNOMELIX Foerster, p. 335 

Zemiophora Foerster. In type citation change "Included by Dalla torre, 
1901" to: Included by Thomson, 1894. Perkins, 1962, p. 464, con- 
siders this a valid genus. 

Synomelix Foerster. Change type citation to: (Synomelix sieboldii 
Kriechbaumer) = Synomelix albipes (Gravenhorst) {teste per- 
kins, 1962, p. 456). 

Zemiophorus Thomson. Delete type citation. Add : Emend. 

Genus D/a/ges Foerster, pp. 335-336 

Dialges Foerster. Change type citation to: Tryphon vepretorum Graven- 
horst. Incl. by Kriechbaumer, 1897 (June) ; Davis' species was 
inch later. Syn. of Otlophorus Foerster {teste Perkins, 1962, pp. 

417,441). 

Delete from catalog. No Nearctic spp. Transf. Nearctic spp. to Pantor- 
haestes Foerster. 

Genus PANTORHAESTES Foerster 
( = Dia/ges of Catalog, p. 335) 

Trophoctonus Foerster. Transf. from syn. of Dialges Foerster. 
Pantorhaestes Foerster. Removed from syn. of Dialges Foerster as a 
valid genus (teste Perkins, 1962, pp. 443, 460). 

frontalis (Davis), n. comb. Transf. from Dialges Foerster, p. 336. 

rivalis (Davis), n. comb. Transf. from Dialges Foerster, p. 336. 

tricolor (Davis), n. comb. Transf. from Dialges Foerster, p. 336. 

Genus MESOLEPTIDEA Viereck, p. 336 
decens (Cresson). Wis. Host: A rc/z/ps sp. 

Genus POLYTERUS Foerster, p. 336 

Sychnoportus Foerster, 1868, p. 208 (as Sychnoporthus, p. 220). Change 
date in type citation to 1898 (1897). 

Genus SYNDIPNUS Foerster, p. 337 

Polypystis Foerster. Perkins, 1962, p. 447, considers this a valid genus. 
If so treated, most (all?) of the Nearctic spp. will go in Poly- 
pystis. 

Genus HYPAMBLYS Foerster, p. 337 

*Apystus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 212 (syn. 
teste Perkins. 1962, p. 406). 

Type: Tryphon albopictus Gravenhorst. Desig. by Perkins, 
1962. 
*Lathrophagus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 213 
(syn. teste Perkins, 1962, p. 435), 

Type : Mesoleius buccatus Holmgren. Desig. by Perkins, 1962. 



178 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus PHOBETES Foerster 
( = /pocfonmus Hincks, p. 338) 

Phobetes Foerster, 1868 (nee Auott.). 

Type: Tryphon fuscicornis Holmgren. Desig. by Viereck, 
1914, as type of Phobetus Thomson, emend. 
Ipoctonus Foerster. Preocc. (Syn. teste Townes, Momoi, and Townes, 

1965, p. 268.) 
Philotymma Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 209. 

Type: Ichneumon chrysostornus Gravenhorst. Desig. by 
Townes, Momoi, and Townes, 1965. An earlier desig. by 
Perkins, 1962 of Ichneumon leptocerus Gravenhorst is 
invalid because that sp. is not one of the four (two through 
generic syn.) included by Townes, Townes, and Gupta, 
1961, p. 217, who made no type desig. then. 
*Phohetus Thomson, 1889. Opusc. Ent., fasc. 13 p. 1430. Emend. 
*Griphodes Kriechbaumer, 1894. Termesz. Fiizetek 17: 57 (syn. of 
Philotymma Foerster teste Townes, Townes, and Gupta, 1961, p. 
217). 

Type: Griphodes caligatus Kriechbaumer. By monotypy. 

^Phobetellus Hincks, 1944. Proc. Roy. Ent. Soc. London (B) 13: 34. 

N. name for Phobetus. 
Ipoctoninus Hincks. N. name. 

Perkins, 1962, stated that Philotymma Foerster was a new synonym of 
Phobetes Foerster. As first reviser his choice is valid, even though Townes, 
Momoi, and Townes, 1965, use the name Philotymmn for the genus. 

egregia (Davis), n. comb. 

striata (Davis), n. comb. 

unicolor (Provancher) , n. comb. 

Genus HYPERALLUS Foerster, p. 338 

*Gastroporus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 206. 

Tvpe: Hyperallus caliroae Viereck. Incl. and desig. by Per- 
kins, 1962. 

Genus HADRODACTYLUS Foerster, pp. 338-339 

*Dizemon Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 199. 

Type: Mesoleptus typhae (Fourcroy). Incl. by Kriechbaumer, 
1891. 
Zemiodes Foerster. Change type citation to: Mesoleptus (Zemiodes) ery- 

thropus Kriechbaumer. Desig. by Perkins, 1962. 
*Narcopoea Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 204. 

Type: Ichneumon typhae Fourcroy. Incl. by Kriechbaumer, 
1891. 

Genus EURYPROCTUS Holmgren, pp. 339-340 

Euryproctus Holmgren. In type citation, replace "Desig. by Viereck, 
1912" with: Desig. by Thomson, 1894. 

bituminosus Davis. Mass., Minn., Idaho. 

Tribe OXYTORINI 
(zzCallidiotini of Catalog) 

OXYTORiNA was used by Kerrich in 1939 for this supergeneric category. 

Genus OXYTORUS Foerster 
( = Co//id(otes Foerster, p. 340) 

Delolytus Foerster. Syn. Change type citation to: {Atractodes varicornis 

Holmgren ) = luridator ( Gravenhorst ) . 
Callidiotes Foerster (syn. teste Meyer, 1936, Tabl. anal. Faune URSS 

(21) V. 5, p. 147). Change type citation to: {Mesoleptus coxator 



ICHNEUMONIDAE 179 

Gr avenhor st) =luridator (Gravenhorst) {teste Pfankuch, 1906, 
Ztschr. syst. Hym. Dipt. 6: 81; Perkins, 1962, p. 411). 
Oxytorus Foerster. Selected as valid name by first reviser (Meyer, 1936, 

Tabl. anal. Faune URSS (21) v. 5, p. 147). 
*Pantoporthus Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25 : 209. 

Type: Ichneumon luridator Gravenhorst. Desig. by Perkins, 
1962. 
Paracanidia Viereck. Syn. 
Prosmoridea Cushman. Syn. 

albopleuralis (Provancher) , n. comb. 

antennatus (Cresson). Va. Placed in Oxytorus by Dalla Torre, 1901. Cat. 
Hym., v. 3 (Pt. l),p. 352. 
Exolytus concamerus Davis, Transf. from Alphabetic List of Unplaced 
Species, p. 408. N. syn. 

elongata (Davis), n. comb. 

Subfamily ORTHOPELAAATINAE 

Genus ORTHOPELMA Toschenberg, p. 341 

calif ornicum Ashmead. B. C., Wash., Oreg., Idaho. Hosts: Diastrophus kin- 
caidii Gill. ; gall on thimbleberry. 

mediator (Thunberg). 111., N. Dak., Colo., Wash. Hosts: Diplolepis mayri 
(Schlecht.) ; D. viultispinosa (Gill.). 

♦Subfamily MICROCLEPTINAE 

{zzPlectiscinae of authors and Catalog, pp. 341-345, Sup. 1, p. 56) 

Townes selected the name Microleptinae, from Microleptes, the oldest 
generic name, for Plectiscinae of authors. Unfortunately, Microleptes is a 
lapsus calami for Microcleptes. In Gravenhorst's original description, the 
Greek word /jukpokXs-tv? was incorrectly spelled /j.iKpoXt'^TTTjs and ti-ansliter- 
ated from the latter, but the meaning given was for the former, "fureni 
rerum minutarum significante, quia hoc insectum minutum forsan non nisi 
larvas insectorum parvorum aggreditur." Paragraphs (a) (ii) and (c) of 
Article 32 of the International Rules apply here (see also Hopper, 1959. Ent. 
Soc. Wash. Proc. 61 (4) : 166). 

Genus CYLLOCERIA Schiedte, p. 342 

Asphragis Foerster. Transf. from Lissonotini (syn. teste Perkins, 1962. 

p. 407; misdet. type-species). 
Xenacis Foerster. Transf. to syn. of Cryptopimpla Taschenberg. 

Genus DIACRITUS Foersfer, p. 198 

Transf. from Neoxoridini (=Poememini of Catalog, p. 198) (teste Cush- 
man, 1917, Ent. Soc. Wash. Proc. 19 (1-4) : 162-165; Townes, 1957, Ent. Soc. 
Wash. Proc. 59 (1): 16), and place provisionally in *Microcleptinae after 
Cylloceria Schiodte on p. 342 (Townes, loc. cit. supra). Change type citation 
to: {Mesostenus rufipes Provancher) =mufe6ris (Cresson). Desig. by Vier- 
eck, 1914. (Davis, 1895, only doubtfully referred mulebris to Diacritus; spe- 
cies definitely incl. by Dalla Torre, 1902, teste Perkins, 1962, p. 417.) 

Genus SYMPLECIS Foerster, p. 343 

clipeator (Lundbeck). Transf. from Alphabetic List of Unplaced Species, p. 

408 {teste Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 106). 

Genus MEGASTYLUS Schiodte, p. 343, Sup. 1, p. 56 

Correct author name, misspelled "Schidte" in Sup. 1, p. 56. 
Brephoctonus Foerster. Transf. to syn. of Plectiscus Gravenhorst 
( -=Leipaulu8 Townes, p. 348). 



180 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus PROCLITUS Foersfer, p. 344 
Proclitus Foerster. Change type citation to: {Proclitus grandis Foerster) 
= praetor Haliclay {teste Perkins, 1962, p. 448). 

Genus MICROCLEPTES Gravenhorst, emend. 
( = A1/cro /epfes Gravenhorst, p. 345) 

See remarks under Microcleptinae. 

Subfamily ORTHOCENTRINAE 

{:=Plectiscinae of type-genus) 

Taxonomy: Townes, 1958. Ent. Soc. Wash. Proc. 60 (5): 221 (identity 
of type-species of Plectiscus Gravenhorst) . 

♦Genus PLECTISCUS Gravenhorst 
( = l.e/pau/us Townes, p. 348) 

Plectiscus Gravenhorst, 1829. Ichn. Europaea v. 2, p. 978. 

Type: Plectiscus impurator Gravenhorst. Desig. by West- 
wood, 1840. 
Brephoctonus Foerster. Syn. Transf. from syn. of Megastylus Schiodte, 

p. 343 {teste Townes, 1958. Ent. Soc. Wash. Proc. 60 (5) : 221). 
Leipaulus Townes (syn. teste Townes, loc. cit. supra). 

suffuscus (Davis). Transf. from Leipaulus Townes {teste Townes, loc. cit. 
supra) . 

Subfamily DIPLAZONTINAE 

( = D/p/azonmoe of Catalog) 

Revision: Dasch, 1964. Amer. Ent. Inst. Mem. 3: 1-273; 153 figs, (keys 
to genera and species) . 

Genus DIPLAZON Nees, pp. 348-349, Sup. 1, p. 57 

♦albotibialis Dasch. Que., Alaska. 

Diplazon albotibialis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 11, 13, 53. 

$, ?. 
*algidus Dasch. Greenland and Labrador to Alaska, south to north. Que., 
Man., Wyo., B. C. 
Diplazon algidus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 10, 13, 51. $, 

9. 

*angustus Dasch. P. E. I., N. W. T., Man., Yukon, Alaska, Calif., Huds. 
Zone. 
Diplazon angustus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 10, 13, 46. $, 

9. 
bizonarius (Gravenhorst). Transf. to Syrphophilus Dasch. 

*bradleyi Dasch. Que., Man., N. W. T., Alaska, B. C, Wash., N. Mex. 

Diplazon bradleyi Dasch, 1964. Amer. Ent. Inst. Mem. 3: 10, 12, 37. $, 

9. 

*cascadensi9 Dasch. Alta., Wash. 

Diplazon cascadensis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 12, 14, 49. 

$, ?. 

♦constrictus Dasch. Colo., Calif., Canad. Zone. 

Diplazon constrictus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 12, 14, 47. 

S, ?. 

'^deletus deletus (Thomson). Maine, Que. to N. W. T., Alta., Alaska south to 
N. C, Mich., Colo. Reduced to subsp. status {teste Dasch, 1964, p. 
41). 



ICHNEUMONIDAE 181 

*Ba8sus multicolor Holmgren, 1856. Svensk. Vet.-Akad. Handl. (n.s.) 1: 

Bassus deietus Thomson, 1890. Opusc. Ent. 14: 1471. $, ?. 

*deletus fuscifemoratus Dasch. Alaska, Yukon south to Oreg., Utah, Colo. 

Diplazon deietus fuscifemoratus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 
41,43. S, 9. 

*flavifrons Dasch. B. C, Wash., Calif. 

Diplazon flavifrons Dasch, 1964. Amer. Ent. Inst. Mem. 3: 10, 12, 39. $, 

?. 

*galenensis Dasch. Idaho (Galena Summit, Blaine Co.). 

Diplazon galenensis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 11, 14, 45. $, 

9. 

ichneumonoides {Proyancher) . Transf. to Syrphophilus Dasch. 

Bassus scapulaius Provancher. Transf. to syn. of Syrphophilus tricincto- 
rius tricmctorius (Thunberg). 

*insulcatus Dasch. Alta., Wash. 

Diplazon insulcatus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 10, 12, 58. $, 

9. 

laetatorius (Fabricius). 

Biology: Weems, 1954. Ohio Jour. Sci. 54: 45-54. — Butler and Dasch, 
1958. Ariz. Agr. Exp. Sta. Tech. Bui. 135: 5-7. 

orbitalis (Cresson). Transcont. in Huds., Canad. Transit, zones, but mostly 

west of 100° W. longitude. 
Bassus orbitalis Cresson. Deleted from syn. of tibiatorius (Thunberg) ; a 

valid species {teste Dasch, 1964, pp. 11, 13, 29). 
Bassus cinctulus Cresson (syn. teste Dasch, 1964, p. 29). 
Bassus orbitalis Ashmead (syn. teste Dasch, 1964, p. 29). 

pectoratorius (Thunberg). Transit. Zone into Mexico. 

rufiventris (Gravenhorst) . Transf. to Tymmophorus Schmiedeknecht. 

scutellaris (Cresson). Canad. Zone. 

*tetragonus rufigaster Dasch. Que., Mass., N. Y., Mich., N. W. T., Sask., 
Mont., Alta., Alaska, B. C, Wash. 
Diplazon tetragonus rufigaster Dasch, 1964. Amer. Ent. Inst. Mem. 3 
25,28. $, 9. 

tetragonus tetragonus (Thunberg), changed status. Reduced to subsp. rank 
(teste Dasch, 1964, p. 25). 
* Bassus albicornis Provancher, 1879. Nat. Canad. 11: 273 (lapsiis for 
albicoxus Provancher) . 

tibiatorius (Thunberg). Remove, in sensu strictu, from catalog. 
Bassus orbitalis Cresson. Remove from syn. ; a valid sp. 
Bassus cinctulus Cresson. Syn. of orbitalis (Cresson). 
Bassus orbitalis Ashmead. Syn. of orbitalis (Cresson). 

^tibiatorius occidentalis Dasch. Alta. to Yukon and B. C. south to Calif., 
Ariz., Colo. D. tibiatorius tibiatorius (Thunberg) occurs in Eu- 
rasia. 
Diplazon tibiatorius occidentalis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 
33, 34. S, 9. 

*wallevi Dasch. N. W. T., Yukon, Man. 

Diplazon walleyi Dasch, 1964. Amer. Ent. Inst. 3: 11, 14, 23. $,9. 

Genus SYRPHOPHILUS Dasch 

Zoolrephes of authors (in part) {=iZootrephus Thomson, emend). 
Syrphophilus Dasch, 1964. Amer. Ent. Inst. Mem. 3 : 8, 59. 

Type: Bassus bizonarius Gravenhorst. Orig. desig. 

Place in Diplazontinae after genus Diplazon Nees, p. 349, 



182 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*asperatus Dasch. Alaska. 

Syrphophilus asperatus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 60, 71. 

9. 

bizonarius (Gravenhorst) . Transf. from Diplazon Nees, Sup. 1, p. 57 (teste 
Dasch, 1964, pp. 60, 61). 

ichneumonoides (Provancher) , Transcont. in Canad. Zone. Transf. from Di- 
plazon Nees, Sup. 1, p. 57 {teste Dasch, 1964, pp. 61, 65). 

*tricinctorius niveus Dasch. Mass., Sask., Alta., Alaska south to S. C. and 
Mo. 
Syrphophilus tricinctorius niveus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 
67, 70. S, 9. 

tricinctorius tricinctorius (Thunberg), changed status. N. W. T. to Alaska, 
south to N. C, N. Mex., Calif, in Huds., Canad., Transit. Zones. 
Reduced to subsp. rank (teste Dasch, 1964, p. 68). 
Bassus scapulatus Provancher (syn. teste Dasch, 1964, p. 68). 

iricinctus (Ashmead). N. W. T. to Alaska south to Que., Colo., Wash., in 
Arctic and Huds. Zones. Transf. from syn. of Promethes elon- 
gatus (Provancher), p. 350 (teste Dasch, 1964, pp. 60, 61, 72). 

♦Genus TYMMOPHORUS Schmiedeknecht 

Zootrephes of authors (in part) (= Zootrephus Thomson, emend.). 
Tymmophorus Schmiedeknecht, 1913. Opusc. Ichn. 34: 2709, 2714. 

Type: Tymmophorus lacustris Schmiedeknecht = rufiventris 

(Gravenhorst). By monotypy. 

Place in Diplazontinae on p. 349, after *Syrphophilus Dasch. 

*fasciventris Dasch. Yukon, Alaska, Colo., Huds. Zone. 

Tymmophorus fasciventris Dasch, 1964. Amer. Ent. Inst. Mem. 3 : 75, 76, 

85. $, 9. 

*geHclus Dasch. N. W. T., Greenland, Arctic Zone. 

Tymmophorus gelidus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 75, 87. $, 
9. 

gillettii (Davis). Transf. from Syrphoctonus Foerster, p. 351 (teste Dasch, 
1964, pp. 75, 81). 

*graculus graculus (Gravenhorst). Que. to B. C, Alaska, Yukon, south to 
Colo., Wyo., Wash, in Huds. and Canad. Zones; Europe. 
Bassus graculus Gravenhorst, 1829. Ichn. Europaea 3: 339. "? 9". 
Taxonomy: Dasch, 1964. Amer. Ent. Inst. Mem. 3: 75, 76, 77. 

*graculus nigrofemoratus Dasch. N. W. T., Alta., B. C, Colo. 

Tymmophorus graculus nigrofemoratus Dasch, 1964. Amer. Ent. Inst. 
Mem. 3: 77, 79. $, 9. 

rufiventris (Gravenhorst). Transf. from Diplazon Nees, Sup. 1, p. 57 (teste 
Dasch, 1964, pp. 75, 76, 82). 

*thunbergi Dasch. Que. to Mass., west to B. C, Alaska. 

Tymmophorus thunbergi Dasch, 1964. Amer. Ent. Inst. Mem. 3: 75, 76, 
79. $, 9. 

Genus PROMETHES Foerster, pp. 349-350, Sup. 1, p. 57 

Sussaba Cameron. Remove from syn. ; a valid genus, infra. 

cultriformis (Ashmead). Transf. to Sussaba Cameron, p. 350, as subsp. of 
dorsalis (Holmgren). 

elongatus (Provancher). Transf . to Sussaba, p. 350. 

Bioblapsis tricincta Ashmead. Transf. to Syrphophilus Dasch, p. 349, as 

valid sp. 
Promethes rohweri Brues. Transf. to Sussaba Cameron, p. 350, as valid 
sp. 



ICHNEUMONIDAE 183 

*gravenhorsii Dasch. Newfoundland, N. B., Mass., N. Y., Mich., Wash., B. C, 
Alaska. 
Promethes gravenhorsti Dasch, 1964. Amer. Ent. Inst. Mem. 3: 231, 232, 
236. $, ?. 

*nominiiiguis Dasch. Que. (Nominingue). 

Promethes nomininguis Dasch, 1964. Amer. Ent. Inst. Mem. 3 : 231, 240. 

$, ?. 

*striatus Dasch. Pa., Ohio, Colo., Alta. 

Promethes striatus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 231, 233, 238. 

$, ?. 

*sulcator splendidus Dasch. Alaska to Oreg. 

Promethes sulcator splendidus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 
233, 236. $,9. 

sulcator sulcator (Gravenhorst) , changed status. Transcont. Huds., Canad. 
and Transit, zones. Hosts: Sphaerophoria robusta Curr., Melan- 
ostovia mellinum (L.). Reduced to subsp. rank {teste Dasch, 1964, 
pp. 232, 233). 

Genus SUSSABA Cameron 

A valid genus (teste Dasch, 1964, pp. 7, 230, 241). Transf. from syn. of 
Promethes Foerster. Place after Promethes Foerster, p. 350. 

*cognata faceta Dasch. N. S., Que., Ont. to N. Y., Mich., also N. C, Tenn., 
Alaska to Calif. Transit., Canad. and Arctic Zones. S. cognata 
cognata (Holmgren) occurs in Europe. 
Sussaha cognata faceta Dasch, 1964. Amer. Ent. Inst. Mem. 3: 264, 265. 

*coloradensis Dasch. Colo. (4 mi. west Cameron Pass). 

Sussaba coloradensis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 243, 270. 9. 

dorsalis cultriformis (Ashmead), changed status. Alta., B. C, Yukon, 
Alaska. Reduced to subsp. rank (teste Dasch, 1964, pp. 256, 257, 
259). 

*dorsalis dorsalis (Holmgren). Newfoundland, Labrador, N. W. T., Yukon, 
Alaska, south to Calif., Colo., Que., Huds., Canad., Transit. Zones. 
Bassus dorsalis Holmgren, 1856. Svenska Vetensk.-Akad. Handl. (n.s.) 1 
(2): 353,367. 9. 

elongata (Provancher) . Transf. from Promethes Foerster, p. 350 (teste 
Dasch, 1964, pp. 245, 250). 

*nigrithorax Dasch. Que., Baffin Is., N. W. T., Alaska, Colo. 

Sussaba nigrithorax Dasch, 1964. Amer. Ent. Inst. Mem. 3: 245, 249. $, 

9. 

*placita Dasch. B. C, Calif. 

Sussaba placita Dasch, 1964. Amer, Ent. Inst. Mem. 3: 242, 244, 269. $, 
9. 

*pulchella ooriacea Dasch. Que., Maine, Ont., south to N. C. west to Minn., 
Colo., also Alaska. 
Sussaba pulchella coriacea Dasch, 1964. Amer. Ent. Inst. Mem. 3: 246, 
247. $, 9. 

*pulchella pulchella (Holmgren). N. W. T., Que. Huds. Zone; Europe; No. 
India. 
Bassus pulchellus Holmgren, 1856. Svenska Vetensk.-Akad. Handl. (n.s.) 
1 (2): 353, 366. $, 9. 

*punctiventris (Thomson). N. S., Maine to Mass., west to Oreg., Wash., 
Alta., Yukon; also Ariz., Colo. Transit., Canad., Huds. zones: Eu- 
rope. 

Homoporus (!) punctiventris Thomson, 1890. Opusc. Ent. 14: 1490, 
1500. 9. 

Taxonomy: Dasch, 1964, pp. 242, 244, 267. 



184 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

rohweri (Brues). Alaska, Yukon, Sask. south to Calif., N. Mex., Canad., 
Transit Zones. Hosts: Endrosis sarcitrella (L.) and Metasyrphus 
pe7-plexus (Osborn). Transf. from syn. of elongatus (Pro- 
vancher), a valid species (teste Dasch, 1964, pp. 244, 245, 254). 

*rugipleuris ellesmerae Dasch. Ellesmere Is. 

Sussaba rugipleuris ellesmerae Dasch, 1964. Amer. Ent. Inst. Mem. 3: 
261, 262, 263. $, ?. 

*rugipleuris rugipleuris Dasch. N. W. T., Yukon, Alaska, Wyo. Arctic, 
Huds. Zones. 
Sussaba rugipleuris rugipleuris Dasch, 1964. Amer. Ent. Inst. Mem. 3: 
243,244,260,261, 262. $, ?. 

Genus HOMOTROPUS Foersfer 

Hotnotropus Foerster. Remove from syn. of Syrphoctonus Foerster, Sup. 
1, p. 57, a valid genus (teste Townes, Townes and Gupta, 1961, p. 
329). 

Place in Catalog before Syrphoctonus Foerster, on p. 350 (Sup. 1, p. 57). 

alaskensis (Ashmead). Yukon. Transf. from Syrphoctonus Foerster, p. 350 
(teste Dasch, 1964, p. 118). 

albopictus albopictus Davis, changed status. N. W. T., Yukon south to Mich., 
Idaho, B. C, also N. Mex. Reduced to subsp. rank (teste Dasch, 
1964, p. 101). 

*aIbopictus ithacae Dasch. N. S., N. W. T., Que., to N. C. west to Wye, 
Utah, Wash., Oreg. 
Homotropus albopictus ithacae Dasch, 1964. Amer. Ent. Inst. Mem. 3: 
101, 103. $, 9. 

*aquilonius Dasch. Greenland. 

Homotropus aquilonius Dasch, 1964. Amer. Ent. Inst. Mem. 3: 94, 135. 

?. 

*arizonensis Dasch. Colo., Ariz., Calif. 

Homotropus arizonensis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 90, 96, 

114. $, 9. 

'''badius Dasch. Calif. 

Homotropus badius Dasch, 1964. Amer. Ent. Inst. Mem. 3: 91, 99, 170. 

5, ?. 

belangerii (Provancher) . Ont. Transf. from Syrphoctonus Foerster, p. 350 
(teste Dasch, 1964, pp. 90, 96, 116. 5,9). 

*coIumbiensis Dasch. Alta., B. C. south to Calif., Utah, Colo. 

Homotropus colutnbiensis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 94, 99, 
123. S, ?. 

*compactus Dasch. Mich., N. W. T., Colo., Wash., Oreg. 

Homotropus compactus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 90, 96, 
119. $, 9. 

*confertus Dasch. Que., N. Mex., B. C. 

Homotropus confertus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 92, 97, 
142. 3, 9. 

cressonii (Davis). Transf. from Syrphoctonus Foerster, p. 350 (teste Dasch, 
1964, pp. 92, 178). 

cuhriformis (Davis). Alaska. Transf. from Syrphoctonus Foerster p. 350, 
Sup. 1, p. 57 (teste Dasch, 1964, pp. 91, 99, 179). 

decoratus (Cresson). Mex. Transf. from Syrphoctonus Foerster, p. 350 
(teste Dasch, 1964, pp. 93, 99, 127). 

*elegans (Gravenhorst) . Baffin Is., Que., Ont., Mich. 

Bassus elegans Gravenhorst, 1829. Ichn. Europaea 3: 313. $. 
*Bassus rufonotatus Holmgren, 1856. Svenska Vetensk.-Akad. Handl. (n. 
s.) 1: 369. $, 9. 



ICHNEUMONIDAE 185 

*fossatus aureopectus Dasch. N. W. T. to Alaska, south to Colo., Oreg. 

Homotropus fossatus aureopectus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 
158, 160. $, ?. 

*fossatus fossatus Dasch. Maine and Que., to N. Y. and Mass. west to B. C, 
N. W. T. 
Homotropus fossatus fossatus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 94, 
98, 157. $, ?. 

*fraudulentus fraudulentus Dasch. Maine, Que., Idaho, Alta., Yukon, B. C, 
Wash., Colo., Canad. Zone. 
Homotropus fraudulentus fraudulentus Dasch, 1964. Amer. Ent. Inst. 
Mem. 3: 93, 97, 149, 150, 151. $, $. 

*fraudulentus recurvatus Dasch. Calif., Wash. 

Homotropus fraudulentus recurvatus Dasch, 1964. Amer. Ent. Inst. 
Mem. 3: 150, 151, 152. $, 9. 

*iinperfectus Dasch. Labrador, Mich., B. C, Wash., Oreg. 

Homotropus imperfectus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 92, 97, 
155. $, $. 

*labradorensis Dasch. Labrador, Alta., Colo. 

Homotropus labradorensis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 93, 98, 
136. S, ?. 

laevis (Brues). Transf. from Syrphoctonus Foerster, p. 351 (teste Dasch, 
1964, pp. 92,97, 153). 

maculifrons (Cresson). Canad., Transit. Zones; Hawaii. Host: Metasyrphus 
perplexus (Osborn). Transf. from Syrphoctonus Foerster, p. 351 
(teste Townes, 1961, Ent. Soc. Wash. Proc. 63 (2) : 104). 
Taxonomy: Dasch, 1964, pp. 92, 97, 147. 

melanogaster (Holmgren). N. W. T., Que., Alaska, Wash. Arctic and Huds. 
Zones. Transf. from Syrphoctonus Foerster, p. 351 (teste Townes, 
1961. Ent. Soc. Wash. Proc. 63 (2) : 104). 

minimus (Cresson). Transf. from Syrphoctonus Foerster, p. 351 (teste 
Dasch, 1964, p. 171). 

*nigriclypealis Dasch. Que., Alta., Greenland. 

Homotropus nigriclypealis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 91, 
110. 9. 

nigritarsus fuscitarsus (Provancher) , changed status. Change last part of 
distr. to read: . . . also in Canad. Zone east to Atlantic. H. nigri- 
tarsus nigritarsus (Gravenhorst) (recorded in Catalog as elegans 
var. nigritarsus) occurs in Europe. 
Syrphoctonus fuscitarsus (Provancher), Sup. 1, p. 57. Reduced to subsp. 
rank (teste Dasch, 1964, p. 132). Transf. from Syrphoctonus Foer- 
ster, p. 351 (syn. of humeralis (Provancher)), and Sup. 1, p. 57 
(teste Dasch, 1964, p. 132). 

nigritarsus groenlandicus (Holmgren), changed status. Transf. from Syr- 
phoctonus Foerster, p. 351 (teste Townes, 1961. Ent. Soc. Wash. 
Proc. 63: 104). 
Syrphoctonus groenlandicus (Holmgren). Reduced to subsp. rank (teste 
Dasch, 1964, p. 132, 135). 

♦novitus Dasch. Alaska (Sable Pass). 

Homotropus noviius Dasch, 1964. Amer. Ent. Inst. Mem. 3: 98, 130. $. 

*nudus Dasch. Que., N. W. T. to Alaska south to Mich., Colo., Calif. Huds., 
Canad. Zones. 
Homotropus nudus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 90, 96, 111. 

5, ?. 

pacificus (Cresson). N. W. T., Canad. Zone; Mexico. Host: Bigonicheta sp. 
Transf. from Syrphoctonus Foerster, p. 351 (teste Townes, 1961, 
Ent. Soc. Wash. Proc. 63: 104). 
Taxonomy: Dasch, 1964, pp. 95, 99, 167. 



186 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

^pallipes (Gravenhorst). Transcont. in Huds., Canad., and Transit. Zones; 
Mexico, Europe. 
Bassus pallipes Gravenhorst, 1829. Ichn. Europaea 3: 325. "S" = ?. 
Syrphoctonus pallipennis (Provancher) . Transf. from Syrphoctonus 

Foerster (syn. and transf. teste Dasch, 1964, pp. 94, 99, 175). 
*Bassus pallidipes Bignell, 1898. Trans. Devon. Assoc. 30 : 500. Emend. 

pectoralis (Provancher). Transf. from Syrphoctonus Foerster, p. 351 (teste 
Dasch, 1964, pp. 94, 99, 121). 

pleuralis pleuralis (Cresson), changed status. East. No. Amer., centr. Tex. 
Transf. from Syrphoctonus Foerster, p. 351 (teste Dasch, 1964, 
pp. 92,97, 145). 
Syrphoctonus pleuralis (Cresson), p. 351. Reduced to subsp. rank (teste 
Dasch, 1964, p. 145). 

pleuralis syrphicola (Ashmead), changed status. Alta., B. C. south to Calif., 
Utah, Colo. Hosts: Scaeva pyrasti (L.) ; Metasyrphys perplexus 
(Osborn) . 
Bassus syrphicola Ashmead. Transf. from syn. of Syrphoctonus pleuralis 
(Cresson), p. 351, and reduced to subsp. rank (teste Dasch, 1964, 
pp. 145, 146). 

*quadrangularis Dasch. Colo., Alta, Yukon, Alaska, B. C, Idaho, also Mass., 
N.J. 
Homotropus quadrangularis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 91, 
96, 106. $, 9. 

*robsonensis Dasch. B.C. (Robson). 

Homotropus robsonensis Dasch, 1964. Amer. Ent. Inst. Mem. 3: 91, 96, 
183. $, ?. 

robustus (Davis). B. C, Mich. Transf. from Syrphoctonus Foerster, p. 351 
(teste Dasch, 1964, pp. 91, 96, 181). 

*rufulus Dasch. Maine, Ont. to S. C. vv^est to Wis., 111.; also La. 

Homotropus rufulus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 90, 96, 104. 

$, 9. 

*sicarius Dasch. Labrador, Que., N. W. T., Alaska in Huds. and Arctic 
Zones. Host: simuliid pupa. 
Homotropus sicarius Dasch, 1964. Amer. Ent. Inst. Mem. 3: 94, 98, 161. 

S, ?. 

'*'signatus (Gravenhorst). N. S., Que. to Md., Ohio, west to Colo., Oreg., B. C, 

Yukon, Alaska; Europe, Japan, Kuriles. Host: Sphaeroplwria 

cylindrica (Say). 
Bassus (Bassus) signatus Gravenhorst, 1829. Ichn. Europaea 3: 325. 9. 
Syrphoctonus compressiventris (Cresson) (syn. teste Dasch, 1964, p. 

164). 
Zootrephes montanus Davis. Transf. from Syrphoctonus Foerster, p. 350 

(teste Dasch, 1964, pp. 95, 100, 164). 

*tarsaiorius Panzer. Que., N. W. T., Ont., Alaska, B. C, Oreg., Idaho, Alta., 
Colo., Mich., N. Y. in Huds., Canad. Zones; Europe, Asia. 
Bassus tarsatorius Panzer, 1809. Faunae Ins. Gennanicae Jahrg. 9, 
H. 102, T. 19. 9. 

*teres Dasch. Colo. 

Homotropus teres Dasch, 1964. Amer. Ent. Inst. Mem. 3: 90, 95, 113. 5, 
9. 

*venustus Dasch. Colo., Utah, Idaho. 

Homotropus venustus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 89, 95, 138. 
$, 9. 

*vitreu8 Dasch, Labrador, N. W. T., Alaska, B. C, Alta., Wash., Oreg., 
Calif., Wyo., Mich. 
Homotropus vitreus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 94, 100, 173. 
$, 9. 



ICHNEUMONIDAE 187 

Genus SYRPHOCTONUS Foerster, pp. 350-351, Sup. 1, p. 57 

Homotropus Foerster. Remove from syn. A valid genus. 

agilis (Cresson). Syn. of flavolineatus (Gravenhorst) . 

alaskettsis (Ashmead). Transf. to Homotropus Thomson. 

albopictus (Davis). Transf. to Homotropus Thomson and reduced to subsp. 
rank. 

belangerii (Provancher). Transf. to Homotropus Thomson. 

*citropectoralis (Schmiedeknecht) . N. Y., Pa., Ohio, Mich., B. C, Alaska; 
Europe. 
Homocidus citropecioralis Schmiedeknecht, 1926. Opusc. Ichn. 5 (43) : 

3389,3412. $. 

compressiventris (Cresson). Transf. to Homotropus Thomson as syn. of sig- 
natus (Gravenhorst). 

cressonii Davis. Transf. to Homotropus Thomson. 

cultriformis (Davis). Transf. to Homotropus Thomson. 

elegans war. nigritarsus (Gravenhorst). Delete from Catalog. 

*flavolineatus (Gravenhorst). Ti-anscont. in Huds., Canad., Transit, and U. 

Austr. zones. Europe, Asia. Hosts: Baccha clavata (F.), Meta- 

syrphus lapponicus (Zett.). 
Bassus flavolineatus Gravenhorst, 1829. Ichn. Europeaea 3: 337. ?. 
Syrphoctonus agilis (Cresson) (syn. teste Dasch, 1964, p. 201). 

foutsi Cushman. Transf. to Campocraspedon Uchida. 

gillettii Davis. Transf. to Tymmophorus Schmiedeknecht. 

laevis (Brues). Transf. to Homotropus Thomson. 

maculifrons (Cresson). Transf. to Homotropus Thomson. 

melanogaster (Holmgren). Transf. to Homotropus Thomson. 

minimus (Cresson). Transf. to Homotropus Thomson. 

pacificus (Cresson). Transf. to Homotropus Thomson. 

pallipennis (Provancher). Transf. to Homotropus Thomson as syn. of pal- 
lipes (Gravenhorst). 

*patei Dasch. Colo., Alaska. 

Syrphoctonus patei Dasch, 1964. Amer. Ent. Inst. Mem. 3: 201, 207. ?. 

pectoralis (Provancher). Transf. to Homotropus Thomson. 

pleuralis (Cresson). Transf. to Homotropus Thomson. 

Bassus syrphicola Ashmead. Transf. to Homotropus Thomson as valid 
subsp. of pleuralis. 

robustus Davis. Transf. to Homotropus Thomson. 

vertebratus Cushman. Transf. to Xestopelta Dasch. 

*Genus XESTOPELTA Dasch 

Xestopelta Dasch, 1964. Amer. Ent. Inst. Mem. 3: 8, 208. 

Type: Syrphoctonus vertebratus Cushman. Orig. desig. 

Place on page 351 after Syrphoctonus Foerster. 

vertebrata (Cushman). Vt., N. W. T., B. C, Wash. Transf. from Syrphoc- 
tonus Foerster, p. 351 (teste Dasch, 1964, p. 209). 
Himertosoma quinquecinctum Ashmead Ms., in Slosson, 1902. Syn. 
wrongly placed at top of p. 58 in Sup. 1. Place after Syrphoc- 
tonus bilineatus Ashmead Ms, in Slosson, 1900. 

Genus PHTHORIMA Foerster, pp. 351-352, Sup. 1, p. 58 

Phthorima Foerster. In type citation, change "Desig. by Viereck" to: 
Inch by Thomson, 1890 [only one species]. 



188 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

bidens (Davis). Mass., Que., Ont., Ohio, Colo., Alaska. Host: Neocnemodon 
coxalis (Curr.). 

*compressa Desvignes. Alaska, Yukon, B. C, Alta., Colo. ; Europe. 
Bassus compressus Desvignes, 1856. Cat. Brit. Ichn., p. 91. ?. 

extensor Cushman. N. S., Que., Ont., Ohio, W. Va., Tenn., B. C. 

*rufipleurum Dasch. Ont., N. H., Mass., Conn., N. Y., Md., Ohio, Mich, 
N. W. T. 
Phthorima rufipleurum Dasch, 1964. Amer. Ent. Inst. Mem. 3: 217, 218, 
221. $, ?. 

*townesi Dasch. Ont., Mass., N. Y., Ohio, Mich., Iowa. 

Phthorima townesi Dasch, 1964. Amer. Ent. Inst. Mem. 3: 217, 218, 224. 

S, 9. 

*sulcigena Dasch. B.C. (Robson). 

Phthorima sulcigena Dasch, 1964. Amer. Ent. Inst. Mem. 3: 217, 228. $. 

*Genus CAMPOCRASPEDON Uchida 
Campocraspedon Uchida, 1957. Hokkaido Univ. Fac. Agr. Jour. 50: 226, 
237. 

Type: Campocraspedon satoi Uchida. Orig. desig. and by 
monotypy. 

Place on p. 352 after Phthorima Foerster. 
Taxonomy: Dasch, 1964, p. 211. 

foutsi (Cushman). Ont., N. C, Transf. from Syrphoctonus Foerster, p. 351 
(teste Dasch, 1964, pp. 211, 212). 

"'truncatus Dasch. Alaska, B. C, Idaho, Colo. Canad. Zone. 

Campocraspedon truncatus Dasch, 1964. Amer. Ent. Inst. Mem. 3: 211, 
212, 214. $, ?. 

Genus ENIZEMUM Foerster, p. 352, Sup. 1, p. 58 

*huronense Dasch. Mich. (Huron Mts., Marquette Co.). 

Enizemum huronense Dasch, 1964. Amer. Ent. Inst. Mem. 3: 186, 199. 
$. 

*imitatum Dasch. Ariz., Calif. 

Enizemum imitatum Dasch, 1964. Amer. Ent. Inst. Mem. 3: 187, 196. $. 

*insidiosum Dasch. Calif. (Carmel). 

Enizemum insidiosum Dasch, 1964. Amer. Ent. Inst. Mem. 3: 186, 197. 

9. 

*matanuskae Dasch. Alaska, Yukon. 

Enizemum matanuskae Dasch, 1964. Amer. Ent. Inst. Mem. 3: 186, 191. 
?. 

ornatum (Gravenhorst) . N. S., Colo., Yukon, B. C, Canad. Zone. 

petiolatum (Say). Canad. Zone. 

♦scorteum Dasch. Que., N. W. T., Colo., B. C. 

Enizemum scorteum Dasch, 1964. Amer. Ent. Inst. Mem. 3: 186, 194. $, 

9. 

*tridentatum Dasch. Colo., Alta., Alaska. 

Enizemum tridentatum Dasch, 1964. Amer. Ent. Inst. Mem. 3: 185, 187, 
189. a, 9. 

Subfamily METOPIINAE 

Revision: Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 1]: 
3-318. 

Genus METOPIUS Panzer, pp. 352-353 
METOPIUS subgenus METOPIUS Panzer, p. 352 

Delete statement in Catalog that this subgenus is not represented in the 
Nearctic Region. 



ICHNEUMONIDAE 189 

*galbaneus Townes and Townes. Calif., Oreg. 

Metopius (Metopius) galbaneus Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 63, 74. $, ?. 

*krombeini epixanthus Townes and Townes. Colo., Ariz., Tex. 

Metopius (Metopius) kromheini epixanthus Townes and Townes, 1959. 
U. S. Nat.l Mus. Bui. 216 [Pt. 1] : 69, 71. 5, 9. 

*kroinbeini krombeini Townes and Townes. N. Y. and N. J. south to Ga., 
west to Wis. and Ark. 
Metopius (Metopius) kromheini krombeini Townes and Townes, 1959. 
U. S. Natl. Mus. Bui. 216 [Pt. 1] : 63, 69, 70. $, ?. 

*miniicus Townes and Townes. Ariz. 

Metopius (Metopius) mimicus Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1]: 63, 68. $, ?. 

pulchellus montanus Cresson, changed status. Colo., Utah, Ariz. Transf. 
from subg. Clemontia Michener and reduced to subsp. rank (teste 
Townes and Townes, 1959, p. 72). 

pulchellus pulchellus Cresson, changed status. Colo., Wyo. Transf. from 
subg. Clemontia Michener and reduced to subsp. rank (teste 
Townes and Townes, 1959, pp. 72, 74). 

*pulchellus sonora Townes and Townes. Ariz., N. Hex. 

Metopius (Metopius) pulchellus sonora Townes and Townes, 1959. U. S. 
Natl. Mus. Bui. 216 [Pt. 1] : 72, 73. S, 9. 

robustus concinnus Cresson, changed status. Calif. Transf. from subg. Cle- 
montia Michener, and reduced to subsp. rank (teste Townes and 
Townes, 1959, p. 64). 

robustus niii-andus Cresson, changed status. Colo., Kans. Transf. from syn. 
of Metopius (Clemontia) montanus Cresson and reduced to subsp. 
rank (teste Townes and Townes, 1959, p. 65). 
Metopius (Clemontia) grandior Viereck (syn. by Townes and Townes, 
1959, p. 65). 

robustus robustus Cresson, changed status. N. Y. and N. J., to Ga., Ohio, 
Tenn., Mo. Transf. from subg. Clemontia Michener and reduced 
to subsp. rank (teste Townes and Townes, 1959, pp. 64, 66). 
Metopius (Clemontia) harbecki Skinner (syn. by Townes and Townes, 
1959, p. 66). 

*vittatus Townes and Townes. B. C, Oreg., Calif., Utah, Ariz. 

Metopius (Metopius) vittatus Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1]: 64, 76. $, 9. 

Metopius subgenus Clemontia Michener, pp. 352—353 

Delete from Catalog and place species in subgenera as indicated below. 
Type-species was misidentified; true Ichneumon micratorius Fabricius be- 
longs to Peltocarus Thomson of which there are no Nearctic species (teste 
Townes and Townes, 1959, pp. 61-62). 

basalts Cresson. Transf. to subg. Tylopius Townes and Townes. 

bellus Cresson. Transf. to subg. Tylopius Townes and Townes. 

birkmani Brues. Transf. to subg. Cultrarius Davis. 

comptus Cresson. Transf. to subg. Cultrarius Davis. 

concinnus Cresson. Transf. to subg. Metopius Panzer as subsp. of robustus 
Cresson. 

edwardsii Cresson. Transf. to subg. Tylopius Townes and Townes. 

Metopius nevadensis Cresson. Remove from syn. and transf. to subg. 
Tylopius Townes and Townes as valid subspecies of pollinctorius 
(Say). 

errantia Davis. Transf. to subg. Peltales Townes and Townes. 



190 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

grandior Viereck. Transf. to subg. Metopius Panzer as syn. of robustus mi- 
randus Cresson. 

harbecki Skinner. Transf. to subg. Metopius Panzer as syn. of robustus ro- 
bustus Cresson. 

medianus Morley. Transf. to subg. Tylopius Townes and Townes as question- 
able syn. of basalis basalis Cresson. 

montanus Cresson. Transf. to subg. Metopius Panzer as subsp. of pulchellus 
Cresson. 
Metopius mirandus Cresson. Delete from syn. and transf. to subg. Meto- 
pius Panzer as subsp. of robustus Cresson. 

pollinctorius (Say). Transf. to subg. Tylopius Townes and Townes. 

Metopius pinatorius Brulle. Remove from syn.; valid European sp., or 
subsp. of pollinctorius Say {teste Townes and Townes, 1959, p. 
98). 

pulchellus Cresson. Transf. to subg. Metopius Panzer. 

robustus Cresson. Transf. to subg. Metopius Panzer. 

rufipes Cresson. Transf. to subg. Cultrarius Davis. 

scitulus Cresson. Transf. to subg. Cultrarius Davis. 

uliimatus Davis. Transf. to subg. Cultrarius Davis. 

xanthostigma Ashmead. Transf. to subg. Cultrarius Davis. 

METOPIUS subgenus CULTRARIUS Davis, p. 353 

birkniani Brues. Calif., Ariz., Tex. Transf. from subg. Clemontia Michener. 
Change sex signs following bibliog. ref. to 5 (? misdet.) {teste 
Townes and Townes, 1959, pp. 78, 88). 

comptus Cresson. Ga., S. Dak. Transf. from subg. Clemontia Michener 
{teste Townes and Townes, 1959, p. 78). 

*consector Townes and Townes. Va., Minn., Mo., Nebr., Kans., S. Dak., Colo. 
Metopius (Cultrarius) consector Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 78, 83. $, 9. 

*pectoralis Townes and Townes. Mo., ?Tex. 

Metopius (Cultrarius) pectoralis Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 78, 85. $, ??. 

rileyi Marlatt. Nebr. 

rufipes Cresson. Nebr. Transf. from subg. Clemontia Michener {teste 
Townes and Townes, 1959, pp. 78, 84). 

*scapulatus Townes and Townes. Tex. (McDade). 

Metopius (Cultrarius) scapulaius Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 77, 81. $. 

scitulus Cresson. Transf. from subg. Clemontia Michener {teste Townes and 
Townes, 1959, pp. 78, 80). 

*secundus Townes and Townes. N. Mex. ("San Janacio"). 

Metopius (Cultrarius) secundus Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 78, 89. 9. 

ultimatus Davis. N. J., Ind., Mo., Kans., La., Miss. Transf. from subg. Cle- 
montia Michener {teste Townes and Townes, 1959, pp. 78, 86). 

xanthostigma Ashmead. Md. to S. C. west to N. Dak. Transf. from subg. 
Clemontia Michener {teste Townes and Townes, 1959, pp. 78, 82). 

*METOPIUS subgenus PELTALES Townes and Townes 

Metopius subg. Peltales Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 61,91. 

Type : Metopius errantius Davis. Orig. desig. 



ICHNEUMONIDAE 191 

Place under genus Metopius Panzer following subg. Cultrarius Davis, p. 
353. 

*errantius arizonicus Townes and Townes. Ariz. (Chiricahua Mts.). 

Metopius (Peltales) errantius arizonicus Townes and Townes, 1959. U. S. 
Natl. Mus. Bui. 216 [Pt. 1]: 93, 94. ?. 

*errantius californicus Townes and Townes. Calif., Oreg. 

Metopius (Peltales) errantius californicus Townes and Townes, 1959. 
U. S. Natl. Mus. Bui. 216 [Pt. 1]: 93, 95. $, 9. 

errantius errantius Davis, changed status. Minn., Ont., N. J., Va. Transf. 
from subg. Clemontia Michener and reduced to subsp. rank (teste 
Townes and Townes, 1959, pp. 92, 93). 

♦errantius floridanus Townes and Townes. Fla. (Alachua Co.) . 

Metopius (Peltales) errantius floridanus Townes and Townes, 1959. U. S. 
Natl. Mus. Bui. 216 [Pt. 1] : 93, 94. ?. 

*notatus Townes and Townes. Ala. (Pyriton). 

Metopius (Peltales) notatus Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1] : 92, 96. 9. 

♦METOPIUS subgenus TYLOPIUS Townes and Townes 

Metopius subg. Tylopius Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 61, 97. 

Type: Metopius pinatorius Brulle (= nticratorius of authors, 
not Fabricius). Orig. desig. 

Place under genus Metopius Panzer after subg. Peltales Townes and 
Townes, p. 353. 

basalis basalis Cresson, changed status. ?Ga. Transf. from subg. Clemontia 
Michener and reduced to subsp. rank {teste Townes and Townes, 
1959, pp. 104, 105). 
?Metopius medianus Morley (syn. by Townes and Townes, 1959, p. 105). 

*basalis heinrichi Townes and Townes. N. B., Maine, Ont., B. C. Host: 
Saturniidae. 
Metopius (Tylopius) basalis heinrichi Townes and Townes, 1959. U. S. 
Natl. Mus. Bui. 216 [Pt. 1]: 104. $, 9. 

bellus Cresson. Calif., Oreg. Transf. from subg. Clemontia Michener (teste 
Townes and Townes, 1959, pp. 98, 102). 

edwardsii Cresson. Alta. Transf. from subg. Clemontia Michener (teste 
Townes and Townes, 1959, pp. 97, 103). 

pollinctorius nevadensis Cresson, changed status. Nev., Calif., Wash. Host: 
Apatela sp. Transf. from syn. of Metopius subg. Clemontia Mi- 
chener edwardsii Cresson and reduced to subsp. rank (teste 
Townes and Townes, 1959, pp. 98, 100). 

pollinctorius pollinctorius Say, changed status. N. S. and Que. west to B. C. 
Hosts: Apetcla sp. ; A. sp. on Salix; Gbiphisia septentrionalis 
Wlkr. ; hairy caterpillar on chokecherry. Transf. from subg. Cle- 
montia Michener and reduced to subsp. rank (teste Townes and 
Townes, 1959, p. 98.) 

Genus PSEUDOMETOPIUS Davis, p. 353 

*Tylocomnoides Uchida, 1940. Sapporo Nat. Hist. Soc. Trans. 16 (3) : 178 
(syn. by Townes, 1957. Ent. Soc. Wash. Proc. 59 (3) : 119). 

Type: Tylocomnoides egatvai Uchida. By monotypy and orig. 
desig. 
*Odontotyloconinus Uchida, 1940. Sapporo Nat. Hist. Soc, Trans. 16 
(3): 179. Syn. by Townes, 1957. Wash. Ent. Soc. Proc. 59 (3): 
114). 

Type: Odontotylocomnus pilosus Uchida. Orig. desig. and by 
monotypy. 



192 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

hagenii (CreESon). Okla. 

Genus SPUDAEUS Gistel, pp. 353-354 

minoris (Davis). Trans, to *Bothromus Townes and Townes. 

scaber umbrosus (Davis), changed status. Mts. of Calif, and Ariz. Reduced 
to subsp. ranlc (teste Townes and Townes, 1959, p. 140). S. scaber 
scaber (Grav.) is European. 

♦Genus BOTHROMUS Townes and Townes 

Bothromus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 
1]: 8, 142. 

Type: Tylecomnus minoris Davis. Orig. desig. 

Insert following Spudaeus Gistel, p. 354. 

*gibbus Townes and Townes. Colo., Alta., Ont., Que. 

Bothromus gibbus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 143, 145. 9. 

*minoris cruralis Townes and Townes. Calif. (Camino). 

Bothromus minoris cruralis Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 143, 144. ?. 

minoris minoris (Davis), changed status. Transf. from Spudaeus Gistel and 
reduced to subsp. rank (teste Townes and Townes, 1959, p. 143). 

Genus CHORINAEUS Holmgren, p. 354 

♦aequalis Townes and Townes. Alta., Ont., Que., Mass., Conn., N. Y., Pa., 
Va., N.C. 
Chorinaeus aequalis Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 13, 14,21. $, ?. 

califomicus Ashmead. Alaska, B. C, Wash., Calif. After bibliog. ref. change 
$ to: "S" = 9 (teste Townes in litt.) . 

carinatus (Cresson). Subsp. of funebris (Grav.). 

cariniger Walsh. Syn. of funebris carinatus (Cresson). 

constrictus Davis. N. Y., Md., Va., Tenn., N. C, Ga. 

*emorsus Townes and Townes. N. H., N. S. 

Chorinaeus emorsus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 14,33. ?. 

excessorius Davis. Calif., Ont., Maine, Vt., N. Y., N. J., Pa., R. I.. Md., Va., 
Ga., Ala., Minn., Wis., Mich., Iowa, Mo., Kans., Utah. Hosts: An- 
cylis comptana (Frohlich), Choristoneura fumiferana (Clem.). 

funebris carinatus (Cresson), changed status. B. C. to N. S. south to Tex. 
and Ala. Hosts: Strepsicrates smithiana (Wlshm.), Tetralopha 
asperatella Clem., Acrobasis betulella Hulst, Anacampsis rhofruc- 
tella (Clem.), Argtjrotaenia velutinana (Wlkr.), ITrichotaphe sp., 
lAncylis comptana fragariae (Walsh & Riley). Reduced to subsp. 
rank (teste Townes and Townes, 1959, pp. 27, 28). C. funebris 
funebris (Grav.) is European. 
Chorinaeus cariniger (Walsh) (syn. by Townes and Townes, 1959, p. 28). 

*funebris clarus Townes and Townes. Ariz., Nev. 

Chorinaeus funebris clarus Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 27,30. ?. 

♦funebris divisus Townes and Townes. Idaho, Colo. Host: Grapholitha 
conversann Wlshm. 
Chorinaeus funebris divisus Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 27,31. $, 9. 

*Iabiosus Townes and Townes. N. C. (Mount Pisgah). 

Chorinaeus labiosus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 14,32. 9. 



ICHNEUMONIDAE 193 

*longicalcar flavicrus Townes and Townes. Oreg., "Wash. C. longicalcar lon- 

gicalcar Thomson is European. 
Chorinaeus longicalcar flavicrus Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 16, 19. $, ?. 

longicalcar pleturus Davis, changed status. B. C, Wash., Calif., Minn., Va., 
W. Va. Host: Choristoneura fumiferana (Clem.) ; geometrid. Re- 
duced to subsp. rank (iesfe Townes and Townes, 1959, pp. 15, 16). 

*IongicaIcar suralis Townes and Townes. Calif. (Berkeley). 

Chorinaeus longicalcar suralis Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 16, 20. $, 9. 

♦longicalcar talaris Townes and Townes. Ont., Que. Hosts: Choristoneura 
fumiferana (Clem.), Caripeta divisata Wlkr., ?C. angustiorata 
Wlkr., Semiothisa granitata (Guen.), Protoboarme porcelaria 
(Guen.) , Paraphia piniata Pack. 
Chorinaeus longicalcar talaris Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 16, 18. $, ?. 

opacitas Davis. Calif., Ariz. 

pleturus Davis. Subsp. of longicalcar Thomson. 

pusillus Davis. Syn. of Trieces coslatus (Davis). 

*recurvus Townes and Townes. N. Y., N. H. 

Chorinaeus recurvus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 14,31. $, 9. 

Genus TRIECES Townes, p. 354 

*aquilus Townes and Townes. N. C. (Highlands). 

Trieces aquilus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1] : 36, 50. $. 

*arcuatus Townes and Townes. N. Y., Pa., Md., Va., N. C, Man. 

Trieces arcuatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 35, 56. $, 9. 

*bradleyi Townes and Townes. Maine, N. Y., Md. 

Trieces bradlevi Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1] : 36, 52. ?. 

*calvatus Townes and Townes. Md., N. C, Tex. 

Trieces calvatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 34,54. $, 9. 

*ciliosus Townes and Townes. Md., Va. 

Trieces ciliosus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 36, 53. $, ?. 

costatus (Davis). B. C, Wash., Calif., Colo., Minn., Que., N. B., Maine, Vt., 
N. Y. Canad. Zone. 
Chorinaeus pusillus Davis (syn. by Townes and Townes, 1959, p. 38). 

*densus Townes and Townes. Mich. 

Trieces densus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 

1]: 35,42. $, 9. 

*dentatus Townes and Townes. Ont., R. I., N. Y., Md. Hosts: Geometridae. 
Trieces dentatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 34,55. $, 9. 

*diffidens Townes and Townes. Ont., Que., Maine, Mass., N. Y., N. J., Md., 
Colo. Hosts: Psilocorsis fletcherella Gbsn., Tetralopha sp. 
Trieces diffidens Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 35,57. $, 9. 

*ejectus Townes and Townes. Utah (Wasatch Co.). 

Trieces ejectus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 

1]: 36,46. 5. 

flavifrons ( Ashmead) . Conn., Mich., Minn., S. Dak., Kans., Colo. 



194 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*fusus Townes and Townes. Ont., Mass., N. Y., Pa., N. J., Minn, 

Trieces fusus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 
1]: 36, 49. $, $. 

♦integer Townes and Townes. Mass., N. Y., R. I., Va., N. C, Sask., Mich. 

Trieces integer Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 
1]: 36,48. $, 9. 

marlatti (Ashmead). R. I., N. C. 

'''masoni Townes and Townes. 

Trieces masoni Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 
1]: 35, 37. 9, $. 

onitis (Davis). Man. 

*sapineus litus Townes and Townes. B. C, Alta., Wash., Calif., Colo. 

Trieces sapineus litus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 43,44. 9. 

*8apineus sapineus Townes and Townes . Que., N. H., N. C, Mich., Minn. 

Trieces sapineus sapineus Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 35,43. 9. 

*8parsus Townes and Townes. Vt. to S. C. 

Trieces sparsus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 36,50. $, 9. 

*tegularis Townes and Townes. Calif. (Leevining). 

Trieces tegularis Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 35, 40. 9. 

*teres Townes and Townes. Calif., Colo., Man. 

Trieces teres Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 
1]: 35,41. $, 9. 

texanus (Cresson). N. J., Md., Fla., Kans. Change sex sign following bib- 
liog. ref . from 3 to " 5 " = 9 . 

*walleyi Townes and Townes. Ont., Que. Host: Herculia thymeUisalis 
(Wlkr.). 
Trieces walleyi Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 
1]: 36,47. $, 9. 

Genus PERIOPE Holiday, pp. 354-355 

*Monoplectrochus Heinrich, 1949. Munchner Ent. Gesell. Mitt. 35-39: 
109. (Syn. teste Townes, 1957, Ent. Soc. Wash. Proc. 59(3) : 111). 
Type: Monoplectrochus hoerhamnieri Heinrich. Orig. desig. 
and by monotypy. 

aethiops (Cresson). Alta. 

Genus LEURUS Townes, p. 355 

*caeruliventris borealis Townes and Townes. N. Y. and Mass. to Fla., Mich., 
La., Mo., Kans. Host: Desmia funeralis (HUbn.). 
Leurus caeruliventris borealis Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 148, 149. S, 9. 

caeruliventris (Cresson) of Catalog, in s. sir., excluded as Neotropical (teste 
Townes and Townes, 1959, p. 148). 

♦Genus SETICORNUTA Morley 

*Megatrema Cameron, 1907. Ztschr. System. Hym. Dipt. 7: 468. Preocc. 

Type: Megatrema albopilosa Cameron. By monotypy. 
Seticornuta Morley, 1913. Fauna British India, Hymenoptera, v. 3, Pt. 1, 
pp. 290, 310. 

Type: (Seticornuta albicalcar Morley) = albopilosa (Camer- 
on). Orig. desig. and by monotypy. 

Insert following Leurus Townes, p. 355. 



ICHNEUMONIDAE 195 

apicalis (Cresson). Hosts: Acrobasis betulella Hulst., A. indigenella (Zell.), 
A. sylviella Ely. Transf. from Exochus Gravenhorst (teste 
Townes and Townes, 1959, pp. 151, 153). Change sex sign after 
bibliog. ref. to "$" = 9. {Teste Townes and Townes, 1959, pp. 
153.) 

terminalis (Ashmead). Ont., Que., Pa., N. J., Va., W. Va., N. C, Ohio, ?Mo., 
Kans. Host: "tortricid". Transf. from Exochus Gravenhorst 
(teste Townes and Townes, 1959, p. 151). 

*Genus CARRIA Schmiedeknecht 

Carria Schmiedeknecht, 1924. Ent. Monthly Mag. 60: 112. 

Type : Carria paradoxa Schmiedeknecht. By monotypy. 

Insert before Colpotrochia Holmgren, p. 355. 

*dreisbachi californica Townes and Townes. Calif. 

Carria dreisbachi californica Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1] : 155, 157. $, 9. 

*dreisbachi dreisbachi Townes and Townes. Man to Que., south to 111. and 
N. J., N. C. 
Carria dreisbachi driesbachi Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 154, 155. $, 9. 

*dreisbachi montaiia Townes and Townes. Ariz., N. Mex., Colo. 

Carria dreisbachi montana Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 155, 156. $, 9. 

♦inculcata Townes and Townes. Md., N. J., N. C, S. C. 

Carria inculcata Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 154,158. $, 9. 

Genus COLPOTROCHIA Holmgren, p. 355 

Scallaina Cameron. Remove from syn. Valid subg. of Colpotrochia with 
no Nearctic species (teste Townes, 1957. Ent. Soc. Wash. Proc. 59 
(3): 103-104). _ 

Colpotrochoides Uchida. Place as syn. of subg. Scallama Cameron. 
Other generic synonymy in Catalog syn. of subg. Colpotrochia 
Holmgren (teste Townes and Townes, 1959, p. 128). 

COLPOTROCHIA subgenus COLPOTROCHIA Holmgren 

*Aithris Cameron, 1900. Proc. Manchester. Lit. Philos. Soc. Mem. 44: 
106 (syn. by Townes and Townes, 1959, p. 128). 

Type: Aithris coenutus Cameron. By monotypy. 

crassipes (Provancher) . 

Taxonomy and Biology: Townes and Townes, 1959, p. 131. 

*fuhoni Townes and Townes. Va., N. C, S. C. 

Colpotrochia (Colpotrochia) fultoni Townes and Townes, 1959. U. S. 
Natl. Mus. Bui. 216 [Pt. 1] : 129, 132. ?. 

lexana (Cresson). 

Taxonomy and Biology: Townes and Townes, 1959, p. 129. 

trifasciata (Cresson). Atlantic to 100° W. in Transit, and Upper Austr. 
Zones. 
Taxonomy and Biology: Townes and Townes, 1959, pp. 129, 133. 

Genus TRICLISTUS Foerster, p. 355 

*adustus Townes and Townes. Oreg., Calif., Colo. 

Triclistus adusius Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 109, 119. 9. 

brunnipes (Cresson). Yukon, B. C, Wash., Oreg., Calif., Ariz., Man., Sask., 
Mont., Utah, Wyo., N. Mex. 



196 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

♦chosis Townes and Townes. Ont., Mass., N. Y., Pa., N. J., Md., D. C, Va. 

Triclistus chosis Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Ft. 1]: 109, 121. 9. 

*crassus Townes and Townes. Alaska to Calif, east to P. E. I., Que. and 
N. Y. 
Triclistus crassus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
(Pt. 1) : 108, 109, 110. $, 9. 

emarginalus (Say). Transcont. in Transit., U. Austr. and Canadian Zones. 
Hosts: Archips argyrospilus (Wlkr.), Argyrotaenia vebitinana 
(Wlkr.), Anthophila pariana (Clerck), Croesia albicoinana Clem., 
Tetralopha rohustella Zell., Phlyctaenia extricalis Guen., Tholeria 
reversalis (Guen.), Bucculatrix sp. on Solidago; pyralid on Almis 
incana: Arundinaria sp. Transf. from Exochus Gravenhorst 
(teste Townes and Townes, 1959, p. 113). 
Exochus fulvipes (Cresson) (syn. by Townes and Townes, 1959, p. 113). 

*evexus Townes and Townes. N. H. to Md. and N. J., Minn., Iowa, Sask. 
Hosts: Archips cerasivorana Fitch, Acrobasis comptoniclla Froel. 
Triclistus evexus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 108, 109, 118. $, 9. 

fulvipes (Cresson). Syn. of emarginalus (Say). 

*meIanocephalus (Cameron). Ont. and Que. to N. C; Mich., Wis., S. Dak., 

Ariz. 
Exochus melanocephalus Cameron, 1886. Biol. Centr. Amer., Ins., Hym., 

V. 1, p. 280. "$" = 9. 
Triclistus thoracicus Ashmead (syn. by Townes and Townes, 1959, p. 

126). 

*occidentis Townes and Townes. Alta., Wash., Calif. 

Triclistus occidentis Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 108,109, 116. 9. 

*pallipes Holmgren. Alaska, Yukon south to Calif., Colo., Kans., Mich, to 
Maine south to Ala. and Ga.; Holarctic, Formosa. Hosts: Acleris 
variana (Fern.) , Paralobesia vitiana (Clem.). 

Triclistus pallipes Holmgren, 1873. Ofvers. Svenska Vetensk.-Akad. 
Forh. 30(4) : 59. 9. 

Exochus pygmaeus Cresson (syn. by Townes and Townes, 1959, p. 124). 

*podagricus (Gravenhorst). Alaska, Alta., Idaho, Sask., Ont., Que., N. S.; 
Europe. 
Exochus podagricus Gravenhorst, 1829. Ichn. Europeaea, v. 2, p. 336. ?9. 

propinquus (Cresson). Pa., Va., N. C, Ga., Ala., Ohio, Mich. 

pygmaeus (Cresson). Syn. of pallipes Holmgren. 

*rectus Townes and Townes. Ariz., Calif. 

Triclistus rectus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 108, 109, 117. 9. 

thoracicus Ashmead. Syn. of melanocephalus (Cameron). 

*Genus MACROMALON Townes and Townes 

Macromalon Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 
1]: 8,158. 

Type: Macromalon montanum Townes and Townes. Orig. 
desig. and by monotypy. 

Insert after Triclistus Foerster, p. 355. 

*montanum Townes and Townes. Colo. 

Macromalon montanum Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 159. $, 9. 

Genus HYPSICERA LafreMIe, p. 356 
*cuneata cuneata Townes and Townes. Ont., Maine, R. I., N. Y., Ohio. 



ICHNEUMONIDAE 197 

Hypsicera cuneata cuneata Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1] : 161, 165. 5, ?• 

*cuneata opaca Townes and Townes. Alaska, Alta., Calif., Ariz. 

Hypsicera cuneata opaca Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 165, 166. $, ?. 

*curvator Fabiicius. Que. to Pa., west to B. C. and Oreg. 

Ichneumon curvator Fabricius, 1793. Ent. System., v. 2, p. 177. 9. 
Ichneumon mansuetor Gravenhorst (syn. by Townes and Townes, 1959, 
p. 163). 

*fulviceps Townes and Townes. Va. (Galax). 

Hypsicera fulviceps Townes and Townes, 1959. U. S, Natl. Mus. Bui. 216 
[Pt. 1]: 161, 164. $. 

mansuetor (Gravenhorst). Syn. of curvator (Fabricius). 

♦Genus STETHONCUS Townes and Townes 

Stethoncus Townes and Townes, 1959. U. S, Natl. Mus. Bui. 216 [Pt. 1] : 
8, 167. 

Type: Stethoncus arcticus Townes and Townes. Orig. desig. 
and by monotypy. 

Insert following Hypsicera Latreille, p. 356. 

*arcticus Townes and Townes. Alaska, Labrador, Man., N. H. 

Stethoncus arcticus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 168. $,9. 

♦Genus SYNOSIS Townes and Townes 

Synosis Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 1] : 8, 
168. 

Type: Synosis clepsydra Townes and Townes. Orig. desig. 
and by monotypy. 

Insert before Exochus Gravenhorst, p. 356. 

*clep8ydra Townes and Townes. B. C., Calif., Wis., Mich., Ind., N. Y., Mass., 
Vt., Que. 
Synosis clepsydra Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 169. $, 9. 

Genus EXOCHUS Gravenhorst, pp. 356-357 

*Xanthexochus Morley, 1913. Fauna British India, Hymenoptera, v. 3, 
pp. 290, 292 (syn. by Townes and Townes, 1959, p. 170). 
Type: Xanthexochus scutellatus Morley. Orig. desig. 

albiceps Walsh. 

Taxonomy: Townes and Townes, 1959, p. 266 (sp. unknown). 

albifrons Cresson. Hosts: Choristoneura rosaceana (Harr.), Argyrotaenia 
velutinana (Wlkr.). 

alpinus Cushman. Syn. of flavifrontalis Davis. 

annulicrus Walsh. B. C. to N. B. south to Ariz., and N. C. 

Taxonomy: Townes and Townes, 1959, pp. 212-214 (? descr.). 

apicalis Cresson. Transf . to Seiicornuta Morley. 

*arniil?osus Townes and Townes. P. E. I., Mass., N. Y., N. J., Wis., Alta. 

Exochus armiUosus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 175,226. $, 9. 

atriceps atriceps Walsh, changed status. Ont. to Fla., also Wis., Kans., Tex. 
Reduced to subsp. rank {teste Townes and Townes, 1959, pp. 208, 
209). 



198 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*atriceps atricorpus Townes and Townes. N. B., Maine, Que., Man. 

Exochus atriceps atricorpus Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 208. $, ?. 

atrocoxalis Cresson. A subsp. of mitratus Gravenhorst. 

*brutus Townes and Townes. Ont., Que., N. Y., Pa., Md., N. C, Sask., Alta. 
Host: Phycitid on Tilia. 
Exochus brutus' Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 176, 228. $, 2. 

*bryaiiti Townes and Townes. N. S., Ont., Alta. 

Exochus bryanti Townes and Townes, 1959. U, S. Natl. Mus. Bui. 216 
[Pt. 1]: 173, 191. $, $. 

*canidens Townes and Townes. N. Y., Pa., Mich., Alta. 

Exochus canidens Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 178,250. $. 

*capnodes Townes and Townes. Calif. (Oakland). 

Exochus capnodes Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 177, 239. $. 

*cnemidotus Townes and Townes. Yukon, B. C. to Que., Mass., N. Y., R. I. 

Exochus cnemidotus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 176, 237. $, ?. 

*cuneatus Townes and Townes. Maine (Jimpond). Host: Heterarthrus ne- 
morahis (Fall.) . 
Exochus cuneatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 177,255. $. 

*decoratus hebes Townes and Townes. Alaska, B. C, Ariz., Colo. Host: 
Aclcris variana (Fern.). E. decoratus decoratus Holmgren is Eu- 
ropean. 
Exochus decoratus hebes Townes and Townes, 1959. U. S. Natl. Mus. 
Bui. 216 [Pt. 1]: 263,264. 9. 

decoratus scitulus Provancher, changed status. Ga., Mich., Minn., Ont., Que., 
N. S. Hosts: Acleris variana (Fern.), Argyrotaenia sp. on Abies; 
tortricid on A.bies. Reduced to subsp. rank {teste Townes and 
Townes, 1959, p. 265). 

*denotatus Townes and Townes. Ont., Que., Mass., N. Y., Pa., Md., N. J., 
N. C, Mich., Minn. Host: Zomaria interrupt olineana (Fern). 
Exochus denotatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 177,251. $, 9. 

*dentifrons Townes and Townes. Oreg., Calif., Ariz., N. Mex., Colo., Que. 

Exochus dentifrons Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 174,206. 5, $• 

dorsalis dorsalis Cresson, changed status. Reduced to subsp. rank {teste 
Townes and Townes, 1959, p. 248). 

dorsalis pictilis Walkley, changed status. Reduced to subsp, rank {teste 
Townes and Townes, 1959, p. 247). 

*elimatus Townes and Townes. Colo. (Rocky Mt. N. P.). 

Exochus elhnatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 173, 194. $, 9. 

emarginalus (Say). Transf . to Triclisius Foerster. 

*enodis Townes and Townes. Ariz., Colo., S. Dak. 

Exochus enodis Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 172, 189. $, 9. 

evetriae Rohwer. Wash. 

*externus Townes and Townes. Oreg. ("Summit Prairie"). 

Exochus externus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 175, 238. 9. 



ICHNEUMONIDAE 199 

*fastigatus Townes and Townes. Maine, Mass., R. I. Host: ? Ancylis comp- 
tana (Froel.). 
Exochus fastigatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 178, 249. $, 9. 

fernigineus (Ashmead). Mass., Pa., Md., Fla., Iowa. 

flavifrontalis Davis. Yukon, N. W. T., Wash., Oreg., Calif., Idaho, Utah, 
Colo., Minn., Mich., N .Y., N. B. Host: "tortricid." 

Exochus alpinus Cushman, 1922 (syn. by Townes and Townes, 1959, p. 
204). 

Exochus montanus Walkley, 1958. U. S. Dept. Agr. Monog. No. 2, Sup. 
1, p. 59. N. syn. 

*genualis Townes and Townes. N. J., Md., W. Va., Va., Ga., Ohio. 

Exochus geuualis Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 176,233. $. 

*gravipes (Gravenhorst) . N. S., Ont., Mich.; Europe. 

Ichneumon gravipes Gravenhorst, 1820. Accad. Sci. Torino, Mem. 24: 

384. $, 9. 
Taxonomy: Townes and Townes, 1959, p. 194. 

*liiiilcus Townes and Townes. B. C. (Rogers Pass). 

Exochus hiulcus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 175, 216. ?. 

*litus Townes and Townes. Alaska, Wash., Oreg., Calif., Nev., N. S. 

Exochus litus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 
1]: 178, 196. $, 9. 

mandibularis Cushman. Mich., N. H. 

*megadon Townes and Townes. R. I., N, Y., N. J., Mich., Colo. 

Exochus megadon Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 178,259. $. 

*mesodon Townes and Townes. R. I., Conn., N. Y. 

Exochus mesodon Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 178, 258. 9. 

*mesorufus Townes and Townes. N. B., N. Y., Del., Ala. 

Exochus mesorufus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 176, 232. S, 9. 

mitratus atrocoxalis Cresson, changed status. Man. to N. B. south to W. Va.; 
also "Dak. Terr.", Tex. Host: tortricid on Kalmia. E. mitratus 
mitratus Gravenhorst is European. 
Exochus atrocoxalis Cresson. Reduced to subsp. rank (teste Townes and 
Townes, 1959, p. 181). 

*mitratus orias Townes and Townes. B. C, Wash., Oreg., Calif., Alta., Wyo., 
Utah, Nev. 
Exochus mitratus orias Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 181, 182. $, 9. 

montanus Walkley. Syn. of flavifrontalis Davis. 

*nioniivagus Townes and Townes. Colo. (Rocky Mt. N. P.). 

Exochus montivagus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 174, 201. 9. 

*nigripalpis subobscurus Townes and Townes. Calif. Hosts: Acleris varie- 
gata (Schiff.), Archips argyrospiliis (Wlkr.), Argyrotaenia ci- 
trana. (Fern.), A. franciscana (Wlsm.), Cnephasia longana 
(Haw.), Pandemis pyinisana Kft. Platynota stidtana (Wlsm.)". E. 
nigripalpis nigripalpis Thomson is European. 
Exochus nigripalpis subobscurus Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 217, 219. $, 9. 

^^nigripalpis tectulum Townes and Townes. B. C. to N. S. south to Oreg., 
Colo., N. C. Hosts: Archips rosana (L.), Choristoneura fumifer- 
ana (Clem.). 



200 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Exochus nigripalpis tectulum Townes and Townes, 1959. U. S. Natl. 
Mus. Bui. 216 [Pt. 1] : 217, 218. $, 9. 

*ochreatus Townes and Townes. Ariz. (nr. Alpine). 

Exochus ochreatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 174,202. $. 

ostentatus Davis. Ont., N. Y., Pa., N. J., Md., Ala., Minn., Alta. 

Taxonomy: Townes and Townes, 1959, pp. 252-254 (? descr.). 

pallipes Cresson. Syn. of pleuralis Cresson. 

*peroniae Townes and Townes.2 B. C, Wash., Oreg., Colo., also Ont., Que., 
N. S. to N. Y. and Mich. Hosts: Acleris variana (Fern.), Clepsis 
melaleucana (Wlkr.), lArgyrotaenia sp., Archippus packardianus 
(Fern.). 
Exochus peroniae Townes and Townes, 1959. U. S. Natl. Mus. Bui, 216 
[Pt. 1]: 175,235. $, ?. 

pictilis Walkley. A subsp. of dorsalis Cresson. 

pictus xanthopsis Ashmead, changed status. B. C, Wash., Oreg., Calif., Colo., 
Ont., Que., N. S., Maine, N. H., N. Y., N. C. Host: Heterarthrus 
nemoratus (Fall.). Reduced to subsp. rank (teste Townes and 
Townes, 1959, p. 186). E. pictus pictus Holmgren is European. 

pleuralis Cresson. Mass. to Md., west to Mann., Utah, Ariz., Yukon. Hosts: 
Anacampsis agrimoniella (Clem.), Ancylis comptana (Froel.) ; 
"Peronea" on Viburnum. 
Exochus pallipes Cresson (syn. teste Townes and Townes, 1959, p. 256). 

*postfurcalis Townes and Townes. Ont., N. S. to N. C. west to Man., Colo. 
Hosts: Acleris oxycoccana (Pack.), Episimus argutanus (Clem). 
Exochus postfurcalis Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 177,242. $, 9. 

*pullatus Townes and Townes. Alaska, Man. 

Exochus pullatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 173,197. $, $. 

*quadradens Townes and Townes. Md. (Takoma Park). 

Exochus quadradens Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 177, 254. ?. 

*russeus Townes and Townes. Ariz., N. Mex., ? Pa. 

Exochus russeus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 172,188. $, 9. 

'^rutilatus Townes and Townes. B. C, Wis., Ont. and N. S. south to N. C. 

Exochus rutilatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 176, 225. $, 9. 

scitulus Provancher. A subsp. of decoratus Holmgren. 

semirufus Cresson. Sask. 

*signifer Townes and Townes. Alaska, B. C, Utah, Minn., Wis., Mich., Ohio, 
N. Y., Maine. 
Exochus signifer Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 176,240. ?. 

*silus Townes and Townes. Utah, Idaho, Calif. 

Exochus silus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 [Pt. 
1]: 175, 261. $, 9. 

*spilotus Townes and Townes. Wash., Ont., Maine. 

Exochus spilotus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 177, 245. 9. 

*spinalis Townes and Townes. Calif. 

Exochus spinalis Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 174,202. 9. 



ICHNEUMONIDAE 201 

*8teno8toma Townes and Townes. B. C. to Calif., Ariz. Host: Ephestia figu- 
lilella Greg. 
Exochus stenostoma Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 173, 179. $, 9. 

*8ulcatus Townes and Townes. N. J. ; ?Venezuela. 

Exochus sulcatus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 174, 211. $. 

*tenebrosus Townes and Townes. Wash., Wyo., Colo. 

Exochus tenebrosus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 216 
[Pt. 1]: 174,220. $, 9. 

terminalis (Ashmead). Transf. to Seticomuta Morley. 

*transversus Townes and Townes. Sask., Ont. 

Exochus transversus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 176, 241. ? $, 9. 

*turgidus Holmgren. Sask., Ont., Va., Fla., La., Calif.; Europe. Hosts: Dio- 
ryctria nmatella (Hist.), D. aiiranticella (Grt.), D. reniculella 
(Grt.), Petrova albicapitana Busck. 
Exochus turgidus Holmgren, 1856. Svenska Vetensk.-Akad. Handl. N. S., 
V. 1, n. 2, p. 312. 9. 

*veiitricosus Townes and Townes. R. I., N. Y. 

Exochus ventricosus Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1] : 178, 260. ?. 

virgatifrons Townes and Townes. Ont., Maine, N. H., Mass., Conn., N. Y., 
Md., 111., Wis., Iowa. Host: Canarsia ulmiarrosorella (Clem.). 
Exochus virgatifrons Townes and Townes, 1959. U. S. Natl. Mus. Bui. 
216 [Pt. 1]: 176,229. $, V. 

washingtonensis (Davis). Alaska, Alta., Wash., Idaho, S. Dak., Colo., N. M., 
Mich., Ont., Que., Maine, Vt., N. J., Pa. 

xanthopsis Ashmead. A subsp. of pictus Holmgren. 

Subfamily OPHIONINAE 
Tribe CAMPOPLEGINI 

Genus PYRACMON Holmgren, p. 358 

Rhimphoctona Foerster. Change type citation to: (Pyracmon fulvipes 

Holmgren) =z grandis (Boyer de Fonscolombe). 

Genus CYMODUSA Holmgren, p. 359 

laevissima (Viereck). Transf. from Campoplex Gravenhorst, Sup. 1, p. 59. 
(Suggested by Walley and verified by examination of type spm.) 

Genus CAMPOPLEX Gravenhorst, pp. 359-363 

Sinophorus Foerster. Considered a valid genus by Townes, Townes, and 
Gupta, 1961, pp. 219, 449, but syn. of Campoplex Gravenhorst by 
Perkins, 1962, p. 452. 
*Phaedroctonus Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25:153 
(syn. teste Townes, 1945. Amer. Ent. Soc. Mem. 11 (Pt. 2) : 593; 
Townes, Townes, and Gupta, 1961, p. 220). 

Type: {Phaedroctonus minutus Ashmead) = cupressi (Ash- 
mead). Included by Ashmead, 1902, 
Omorga Thomson. Delete type citation. Add: Emend. 

cupressi (Ashmead). ? N. H. 

frustranae Cushman. Md. "Host: Rhyacionia rioidana Fern. 

laevissima (Viereck). Transf. to Cymodusa Holmgren, p. 359. 

inellipe8(Provancher). Host: Eucordylea ducharmei Freeman. 



202 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

triannulatus (Cameron). Transf. from Alphabetic List of Unplaced Species, 

p. 408 {teste Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 106). 

validus (Cresson). Host: Dioryctria reniculella (Grt.). 

*Genus DEVORGILLA Cameron 

( = /dech>h/s Foerster, nee Huebner, p. 363) 

izzExidechthis Walkley, Sup. 1, p. 59) 

Devorgilla Cameron, 1907. Tijdschr. v. Ent. 50: 51 (syn. ieste Townes, 
Townes, and Gupta, 1961, p. 222). Perkins, 1962, p. 430, considers 
Exidechthis Walkley a valid genus. 

Type: Devorgilla dilatata Cameron. By monotypy. 

All spp. in Idechthis Foerster in catalog (except canescens) will be n. 
comb, in Devorgilla Cameron. 

canescens (Gravenhorst). 

*Omorga columbiana Johnson, 1899. U. S. Dept. Agr. Div. Ent. Bui. 20: 

67. Nom. nud. 
*Idechthis ephestiae Fletcher, 1903. Ent. Soc. Ontario, Ann. Rpt. 33 : 86. 
Nom. nud. 

Genus CASINARIA Holmgren, pp. 364-365 

Horogenes Foerster. Transf. from p. 376. (Syn. teste Townes, Townes, 
and Gupta, 1961, p. 224; Perkins, 1962, p. 428). 

Type: Campoplex tenuiventris Gravenhorst. Included by 
Brischke, 1880. 
Alcima Foerster. Delete type citation. Replace with: Campoplex orbi- 

talis Gravenhorst. Included by Brischke, 1880. 
*Rhythmonotus Foerster, 1886. Naturh. Ver. Rheinlande Verb. 25: 151, 
220 (as Rhytmonotus) (svn. teste Meyer, 1935. Tabl. anal. Faune 
URSS (16) V. 4, pp. 170-171; Perkins, 1962, p. 450). 

Type: Rhytmonotus (!) singularis Schmiedeknecht. Included 
by Schmiedeknecht, 1909. 
*Nothanomalon Szepligeti, 1905. Gen. Ins. 34: 39 (syn. teste Townes, 
Townes and Gupta, 1961, p. 224). 

Type: ISothanomalon novoguineensis Szepligeti. By mono- 

typy- 

Nothanomaloides Uchida, 1928. Fac. Agr. Hokkaido Imp. Univ. Jour. 
21: 273. Preocc. (Syn. teste Townes, Townes, and Gupta, 1961, p. 
225.) 

Type: Nothanomaloides matsuyamensis Uchida. Orig. desig. 
Deltops Seyrig, 1935. Paris Mus. d'Hist. Nat. Mem. 4: 84 (syn. teste 
Townes, Townes, and Gupta, 1961, p. 225). 

Type : Deltops granulicoxis Seyrig. Orig. desig. 
*Longicharops Uchida, 1940. Ins. Matsumurana 14: 131. N. name for 

JS othanomaloides. 
*Neonortoniella Blanchard, 1947. Zool. Mus. Hist. Nat. Montevideo 
Comun. 2 (42) : 4 (syn. teste Townes, Townes, and Gupta, 1961, p. 
225). 

Type: Neonortoniella plusiae Blanchard. By montotypy and 
orig. desig. 

^melanolophiae Walley. B. C. Host: Melanolophia imitata (Wlkr.). 

Casinaria m^elanolophiae Walley, 1959. Canad. Ent. 91 (9) : 565. $, 9. 

Genus CHAROPS Holmgren, p. 366 
annulipes Ashmead. Del. Host: Plathypena scabra (F.) . 

Genus BATHYPLECTES Foerster, p. 366 

*anura (Thomson). N. J., Pa. Host: Hypera postica (Gyll.). 

Canidia anura Thomson, 1887. Opusc. Ent., fasc. 11, p. 1113. $, 2. 

curculionis (Thomson). N. J., Va., N. C, Alta. 

Taxonomy: Aubert, 1960. Vie et Milieu 11 (3) : 490. 



ICHNEUMONIDAE 203 

Biology: Coles and Puttier, 1963. Jour. Econ. Ent. 56 (5) : 609-611.— 
Fisher, Schlinger, Van den Bosch, 1961. Jour. Econ. Ent. 54 (1) : 
196-197.— Puttier, Jones, Coles, 1961. Jour. Econ. Ent. 54 (5) : 
878-880.— Van den Bosch and Dietrick, 1959. Ent. Soc. Amer. 
Ann. 52 (5) : 609-616. 

Genus BIOLYSIA Schmiedeknecht, p. 366 

iristis (Gravenhorst). N. Y., Pa., N. J., Del. 

Biology: Puttier and Coles, 1962. Jour. Econ. Ent. 55 (6) : 831-833. 

Genus CAMPOLETIS Foerster, pp. 366-370 

argentifrons (Cresson). Host: Spodoptera frugiperda (J. E, Sm.). 

longiceps (Roman). Transf. from Ecophoropsis Ashmead, p. 370 (teste 
Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 105). 

Genus Ecphoropsis Ashmead, p. 370 
Delete from Catalog. No Nearc. spp. 
longiceps Roman. Transf. to Campoletis Foerster, p. 368. 

Genus DUSONA Cameron, pp. 370-375 

*ferruginea Walkley. Fla. (Dade Co.). 

Dusona ferruginea Walkley, 1963. Ent. News 74 (9) : 239. 9. 

pilosa (Walley). Host: Melanolophila imitata Wlkr. 

Genus NEPIERA Foerster, p. 375 

Nepiera Foerster. Change type citation to: (Limneria concinna Holm- 
gren) = Nepiera collector (Thunberg) {teste Perkins, 1962, p. 
438). 

marginata (Provancher) . N. B. Hosts: Actebia fennica (Tausch.) ; Graphi- 
phora smithi (Snell) ; Pseudaletia unipuncta (Haw.). 
Biology: Guppy, 1961. Canad. Ent. 93 (7) : 569. 

Genus PHOBOCAMPE Foerster, pp. 375-376 

Hypothereutes Foerster. Change spelling of genus in type citation to 

Hypotherentes (!) and change date to 1898 (1897). 
Phohocam pa Thomson. Delete type citation. Add: Emend. 

disparis (Viereck). N. Y. 

*Genus SYNETAERIS Foerster 

Synetaeris Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25: 155. 

Type: Synetaeris heteropus Thomson. Desig. by Viereck, 
1914. 

Place in Catalog on p. 376 after Phobocampe Foerster. 

*tenuifemur Walley. N. B., Ont. B. C. Host: Choristoneura fumiferana 
(Clem.). 
Synetaeris tenui femur Walley, 1963. Canad. Ent. 95 (1) : 23. $,9. 
Biology: Miller and Renault, 1963. Canad. Ent. 95 (1) : 24-28. 

Genus Horogenes Foerster, pp. 376—380 

Horogenes Foerster. Delete type citation; place as syn. of Casinaria 

Holmgi-en. 
Dyspeies Foerster. Delete type citation; transf. to Tryphonini as valid 

genus. 

Other generic synonyms listed are to be placed as syns. of (Horogenes of 
authors) =iDiadegma Foerster (teste Townes, Townes, and Gupta, 1961, p. 
234). Perkins, 1962, pp. 428-429 suggests Nythobia Foerster as the name to 
use for this concept. 

Stability in the Campoplegini will be possible only after a careful gener- 



204 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

ic revision of the whole tribe. Certainly Horogenes Auctt. has been a catch- 
all genus for many years. 

Transf. all species to Diadegma Foerster except as noted below. 

cacoeciae (Viereck). Transf. to *Tranosema Foerster as syn. of arenicola 

Thomson. 

Genus DIADEGMA Foerster 
( = Horogenes of Catalog and authors) 

*Areolina Enderlein, 1921. Stettin. Ent. Zeit. 82: 41 (syn. teste Townes, 
Townes, and Gupta, 1961, p. 234). 

Type: Areoluta imbecilla Enderlein. Orig. desig. 

Replaces Horogenes Foerster, p. 376. Species transf. from Horogenes in 
Catalog are n. comb, in Diadegma with the exception of a few spp. already 
cited in economic literature. 

*chrysostictuin (Gmelin), n. comb. Molarceic. Host: Ephestia sericarium 

Scott. 
Ichneumon chrysostictos Gmelin, 1790. In Linnaeus Syst. nat. Ed. 13, T. 

I, Pt. V, p. 2721, n. 407. 
Horogenes kiehtani (Viereck) (syn. teste Walley in Fisher, 1959. Canad. 

Jour. Zool. 37 (4): 429). 
Biology and Ecology: Fisher, 1959, loc. cit. supra, pp. 429-446; — 1961, 

Jour. Expt. Zool. 38 (2): 267-275;— 1962, Ecology 43 (2): 

314-316. 

eureka (Ashmead). Hosts: Acleris variepana (Schiff.), Croesia alhicomana 
(Clem.), lEpinotia crenana (Hbn.), Pandemis pyrusana Kft. 

hospitum (Holmgren). Transf. from Alphabetic List of Unplaced species, p. 

409 (teste Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 110). 

pterophorae (Ashmead). Hosts: Acleris variegana (Schiff.), Croesia alhico- 
mana (Clem.). 

stenosomus (Viereck). Que. Host: Eucordylea ducharmei Freeman. 

*Genus TRANOSEMA Foerster 

Tranosema Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 157. 

Type: Tranosema arenicola Thomson. Desig. by Viereck, 
1914. 

Place before Hyposoier Foerster, p. 380 of Cat. 

*arenicoIa Thomson. B. C, Europe. 

Tranosema arenicola Thomson, 1887. Opusc. Ent. fasc. 11, p. 1138. 
Horogenes cacoeciae (Viereck). Transf. from Horogenes Foerster, p. 

377. N. syn. {teste Walley, in litt.). 

Genus HYPOSOTER Foerster, pp. 380-383 

Hyposoter Foerster. In type citation change "Included by Viereck" to: 
Included and desig. by Viereck. 

Ischnoscopus Foerster. Under type citation change the date after Ash- 
mead to 1898 (1897). 

Anilastns Foerster. Remove question mark. 

Anilasta Thomson. Delete type citation and add: Emend. Remove ques- 
tion mark. 

annulipes (Cresson). Yiost : Pseudaletiaunipuncta (Haw.). 

exiguae (Viereck). Ariz. 

parorgyiae (Viereck). Host: Dasychira plagiata (Wlkr.). 

Genus OLESICAMPE Foerster, p.. 383-385 

Olesicampe Foerster. Change type citation to: Ichneumon longipes 
Mueller. Desig. by Viereck, 1912. 



ICHNEUMONIDAE 205 

Olesicampa Thomson. Delete type citation and add: Emend. 
?Holocremna Thomson. Delete type citation and add: Emend. (Perkins, 
1962, p. 427, considers Holocremnus Foerster a valid genus.) 

euurae (Ashmead). Idaho, ?Calif. 

Biology: Caltagirone, 1964. Ent. Soc. Amer. Ann. 57 (3): 287-289 (nr. 
or subsp.?). 

lophyri (Rilev). Minn. Hosts: Neodiprion lecontei (Fitch); N. swainei 
Midd. 

Tribe CREMASTINI 

Genus DIMOPHORA Foerster, p. 386 

Dimophora Foerster. Delete "Desig. by Viereck, 1914" from type cita- 
tion; add: Incl. by Brischke, 1894 (see Perkins, 1962, p. 418). 
Diniophorus Thomson. Change bibliog. ref. to: 1889. Opusc. Ent., fasc. 

13, pp. 1355, 1356. Emend. 
*Demophoru8 Thomson, 1890. Opusc. Ent. fasc. 14, pp. 1442, 1457. 

Type: (Demophorus arenicola Thomson) = Dimophora to- 
busta Brischke. First included species. 

Genus PRISTOMERUS Curtis, pp. 386-387 

euryptychiae Cushman. Hosts: Acrobasis caryae Grt. ; Gnorlmoschema gal- 
laesolidaginis (Riley), Polychrosis carduana Busck, Suleima lie- 
lianthana (Riley); Walshia amorphella (Clem.). 

Genus NEOCREMASTUS Cushman 

Neocreniastus Cushman. Removed from syn. of Zaleptopygus Viereck. A 
valid genus (teste Townes, 1958, Ins. of Micronesia v. 19, n. 2, p. 
66 [under Trathala Cameron]). 

mellipes (Provancher) . Transf. from Za/eplopygas Viereck. 

stigmaterus (Cresson). Transf. from Za/epfopygu* Viereck. 

Genus Za/epfop/gus Viereck, pp. 388—390 
(=:CREAiASTUS Gravenhorsf, nee Cremosfus of Catalog, oucff.) 

Neocremastus Cushman. Remove from syn.; a valid genus. 

aciculatus (Davis). Transf. to *TrathaIa Cameron. 

albipennis (Cresson). Tiansf. to Cremastus Gravenhorst. 

anomalus (Cushman). Transf. to Cremastus Gravenhorst. 

audax (Cresson). Transf. to Cremastus Gravenhorst as syn. of fuscipennis 
(Cresson). 

bilineatus (Cushman). Transf. to Cremastus Gravenhorst. 

brunneipennis (Walley). Transf. provisionally to Cremastus Gravenhorst. 

californicus (Provancher). Transf. provisionally to Cremastus Gravenhorst. 

champlaini (Cushman). Transf . to Cremastus Gravenhorst. 

cleridivorus (Cushman). Transf. to *Trathala Cameron. 

delicatus (Cresson). Transf. to *Trathala Cameron. 

dorcaschemae (Cushman). Transf. to Cremastus Gravenhorst. 

egregiacolor (Viereck). Transf. to Cremastus Gravenhorst. 

flavo-orbitalis (Cameron). Transf. to *Trathala Cameron. 

fuscipennis (Cresson). Transf. to Cremastus Gravenhorst. 

gallaecola (Cushman). Transf . to Cremastus Gravenhorst. 

granulatus (Davis). Transf. to *TrathaIa Cameron. 

hartii (Ashmead). Transf. to *Trathala Cameron. 



206 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

hyalinipennis (Cresson). Transf . to Cremastus Gravenhorst. 

incompletus (Provancher) . Transf . to Cremastus Gravenhorst. 

latithorax (Cushman). Transf. to *Trathala Cameron. 

mellipes (Provancher). Transf . to Neocremastus Cushman. 

mordellistenae (Cushman). Transf . to Cremastus Gravenhorst. 

nemoralis (Davis). Transf. to *Tratha!a Cameron. 

nigriclypealis (Cushman). Transf. to Cremastus Gravenhorst. 

obesus (Cushman). Transf. to *TrathaIa Cameron. 

orbitalis (Cresson). Transf. to Cremastus Gravenhorst. 

orfeus (Davis). Transf. to Cremastus Gravenhorst. 

plesius (Cushman). Transf. to *Trathala Cameron. 

quintilis (Viereck). Transf. provisionally to Cremastus Gravenhorst. 

retiniae (Cresson). Transf. to *Trathala Cameron. 

rohtveri (Cushman). Transf . to Cremastus Gravenhorst. 

rosae (Cushman). Transf. to *Trathala Cameron. 

rostratus (Cushman). Transf. to *Traihala Cameron. 

similis (Cushman). Transf. to *Trathala Cameron. 

snowi (Viereck), Transf. to Cremastus Gravenhorst, provisionally. 

stigmaterus (Cresson). Transf . to Neocremastus Cushman. 

tetralophae (Cushman). Transf. to *Trathala Cameron. 

vierecki (Cockerel!). Transf. to Cremastus Gravenhorst, provisionally. 

All these transfers, excepting the provisional ones are by Townes (see 
Insects of Micronesia 19 (2) : 65-66, 1958). 

Genus CREMASTUS Gravenhorst 
( = Zafepfop/gus Viereck, in part, pp. 388—390) 

The holotype of C. spectator Gravenhorst had been misidentified for 
many years {see Townes, 1958, Ins. of Micronesia, v, 19, n. 2, pp. 65-67). 

albipennis (Cresson). Transf . from Za/eptopygus Viereck. 

anomalus Cushman. Transf. from Zaleptopygus Viereck. 

bilineatus Cushman. Transf. from Zaleptopygus Viereck. 

?brunneipennis Walley. Transf. from Zaleptopygus Viereck. 

?californicus (Provancher). Transf. from Zaleptopygus Viereck. 

champlaini Cushman. Transf. from Zaleptopygus Viereck. 

dorcaschemae Cushman. Transf. from Zaleptopygus Viereck. 

egregiacolor (Viereck) . Transf. from Zaleptopygus Viereck. 

fuscipennis (Cresson). Transf . from Za/epiopygus Viereck. 

Zaleptopygus audax (Cresson). Syn. {teste Townes, 1958. Ins. of Mi- 
cronesia, V. 19, no. 2, p. 65). 

gallaecola Cushman. Mich. Host: Gnorimoschema sp. on aster. Transf. from 
Zaleptopygus Viereck. 

hyalinipennis (Cresson). Transf. from Za/eptopygiis Viereck. 

incompletus (Provancher). Transf. from Za/eptopygus Viereck. 

mordellistenae Cushman. Transf. from Zaleptopygus Viereck. 

nigriclypealis Cushman Transf. from Zaleptopygus Viereck. 

orbitalis (Cresson) . Ti-ansf . from Zaleptopygus Viereck. 



ICHNEUMONIDAE 207 

orbus (Davis). Transf. from Zaleptopygus Viereck. 
?quintilis Viereck. Transf. from Zaleptopygus Viereck. 
rohweri Cushman. Transf. from Zaleptopygus Viereck. 
?snowi (Viereck). Transf. from Za/ep/opvgus Viereck. 
?vierecki (Cockerell). Transf. from Zaleptopygus Viereck. 

*Genus TRATHALA Cameron 
( = Za/epfopygus Viereck, in part, pp. 388—390) 

See statement under Cremastus Gravenhorst. 

*Trathala Cameron, 1899. Mancliester Lit. Phil. Soc. Mem. Proc. 43: 122. 

Type : Trathala striata Cameron. By monotypy. 
*Epicremastus Szepligeti, 1905. Gen. Ins., v. 34, p. 51 (syn. teste 
Townes, Townes, and Gupta, 1961, p. 252). 

Type: Epicremastus concolor Szepligeti. By monotypy. 
Paurolexis Cameron, 1907. Syn. Transf. from syn. of Cremastus of Cat- 
alog {teste Townes, Townes, and Gupta, 1961, p. 252). 
*Haristaeus Cameron, 1910. Ent. Soc. London, Trans. 35: 442. (Syn. 
teste Townes, Townes, and Gupta, 1961, p. 252). 

Type: Haristaeus nigrifrons Cameron. By monotypy. 

aciculata (Davis). Transf. from Za/epropygits Viereck. 
cleridivora (Cushman). Transf. from Za/epfopygus Viereck. 
delicata (Cresson). Transf . from Za/epfopygus Viereck. 
flavoorbitalis (Cameron). Transf. from Za/epfopygus Viereck. 
granulata (Davis). Transf . from Zaleptopygus Viereck. 
hartii (Ashmead). Transf . from Zaleptopygus Viereck. 
latithorax (Cushman). Transf . from Za/ep/opygus Viereck. 
nenioralis (Davis). Transf . from Za/epfopygus Viereck. 
obesa (Cushman). Transf . from Za/epfopygus Viereck. 
plesia (Cushman). Transf. from Zaleptopygus Viereck. 
retiniae (Cresson). Transf. from Za/epfopygus Viereck. 
rosae Cushman. Transf. from Zaleptopygus Viereck. 
rostraia (Cushman). Transf. from Z«/eptopygus Viereck. 
similis Cushman. Transf. from Zaleptopygus Viereck. 
letralophae (Cushman). Transf. from Za/eptopygus Viereck. 

Genus Cremastus AucU. and Catalog, pp. 390-392, Sup. 1, 61 

Temelucha Foerster. Delete from syn., a valid name for species concept 

of Cremastus of auctt. and catalog. 
Neleothymus Foerster. A valid genus. 
Tarytia Cameron. Syn. of Temelucha Foerster. 
Cremastidea Viereck. Syn. of Temelucha Foerster. 
Paurolexis Cameron. Transf. to syn. of Trathala Cameron. 

Transf. all spp. listed on pages 390-392 under Cremastus, except atten- 
uatus Cushman, to Temelucha Foerster. Transf. attenuaius to Neleothymus 
Foerster. 

Genus TEMELUCHA Foerster 

(= Cremastus Auctt. and Catalog, pp. 390-392; Sup. 1, p. 61) 

Temelucha Foerster. Change type citation to: 

Type: (Porizon macer Cresson) = facilis (Cresson). Desig. 
by Perkins, 1962 (one of four spp. incl. by Ashmead in 
Smith, 1900). 
*Paracremastus Szepligeti, 1900. Termesz. Fuzetek 23: 28 (syn. teste 



208 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Townes, Townes and Gupta 1961, p. 256), 

Type: Paracremastus genalis Szepligeti. By monotypy. 
Tarytia Cameron. Syn. 

Cremastidea Viereck. Syn. Change type citation to: (Cremastidea chin- 
ensis Viereck) = biguttula Matsumui'a. 

brevicauda (Cvishman). Transf. from Crcnia«ms, p. 390. 

brevinervis (Cushman). Transf. from Cremastus, p. 391. 

brevipetiolata (Cushman). Transf. from Cremastus, p. 391. 

bruneiceps (Cushman). Transf. from Cremastus, p. 391. 

carpocapsae (Cushman). Transf. from Creniasius, p. 391. 

chilonis (Cushman). Transf. from Crema«fu«, p. 391. 

cookii (Weed). Transf. from Cremastus, p. 391. 

epagoges (Cushman). Transf. from Cremastus, p. 391. 

evetriae (Cushman). Transf. from CreinastHS, p. 391. 

facilis (Cresson). Transf. from Cremastus, p. 391. 

ferruginea (Davis). Transf. from Cremastus, p. 391. 

flaviceps (Cushman). Transf. from Cremastus, p. 391. 

forbesi (Weed). Transf. with its syn. (Sup. 1, p. 61) from Cremastus, p. 392, 
Sup. 1, p. 61. 

gracilipes (Cushman). Transf. from Cremastus, p. 392. 

gracilis (Cushman). Transf. from Cremastus, p. 392. 

grapholithae (Cushman). Transf. from Cremastus, p. 392. 

hamiltonensis (Viereck). Transf. from Cremastus, p. 392. 

interrupter (Gravenhorst) . Mich., Idaho. Transf . from Cremastus, p. 392. 

longigenalis (Cushman). Transf. from Cremastus, p. 392, Sup. 1, p. 61. 

minor (Cushman). Ga. Host: Ostrinia nubilalis (Hbn.). Transf. from Cre- 
mastus, p. 392. 

platynotae (Cushman). Also Yuma. Ariz. Transf. from Cremastus, p. 392. 

protracta (Cushman). Transf. from Cremastus, p. 392. 

pterophori (Cusliman). Transf. from Cremastus, p. 392. 

recta (Provancher). Transf. from Cremastus, p. 392. 

rhyacioniae (Cushman). Transf, from Cremastus, p. 392. 

ruficeps (Cushman). Transf . from Cremastus, p. 392. 

sinuata (Cushman), Transf. from Cremastus, p. 392. 

Genus NELEOTHYMUS Foerster 

Neleothymus Foerster. A valid genus {teste Townes, 1958. Ins. of Mi- 
cronesia V. 19, n. 2, p. 68 [under Temelucha] ) . 

attenuatus (Cushman). Transf. from Cremastus auctt. p. 390 (teste Townes, 
1958, loc. cit. supra) . 

Tribe TERSILOCHINI 

Genus BARYCNEMIS Foerster, p. 394 

Barycnemis Foerster. In type citation replace "Desig. by Viereck, 1914" 
with: Included by Brischke, 1880. 

*Genus ISCHNOBATIS Foerster 
l=:Luchatema Walkley, Sup. 1, p. 61) 
Ischnohatis Foerster, 1868. Naturh. Ver. Rheinlande Verh. 25 : 148. 



ICHNEUMONIDAE 209 

Type: Thersilochus (IchnobatisCl^ stramineipes Brischke. 
Inc. by Brischke, 1880. 
Luchatema Walkley (syn. teste Perkins, 1962, pp. 391, 431). 

baldufi (Walkley), n. comb. Transf. from Luchatema Walkley, Sup. I, p. 61. 

*Genus ALLOPHRYS Foerster 

Allophrys Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 147. 

Type: Thersilochus i I) oculatus Ashmead. Desig. by Viereck, 
1914. 

Place on p. 394, after Hetcrocola Foerster. There are several undes- 
cribed Nearctic spp. belonging here. 

Tribe ANOMALINI 

( = Sublribe ANOMALINA of Catalog) 

Dr. Short and Dr. Townes, in their published works, have included the 
tribes Anomalini, Gravenhorstiini, and Therionini in the subfamily Anoma- 
linae. Dr. Short also included the Ophionellini (Cushman, 1922) here. 

Genus ANOMALON Panzer, p. 395 

reticulatum (Cresson). Host: Conoderus vespertinus (F.). 
Biology: Robb, 1963. Ent. Soc. Amer. Ann. 56: 675. 

*Tribe GRAVENHORSTIINI 

( = Subtribe GRAVENHORSTIINA of Catalog) 

Genus APHANISTES Foerster, p. 396 

*Formosanomalon Uchida, 1928. Hokkaido Imp. Univ. Fac. Agr. Jour, 
21: 241 (syn. teste Townes, 1957. Ent. Soc. Wash. Proc. 59 (3): 
107). 

Type: (Formosanomalon baibarense Uchida) = elegans bai- 
barense Shestakov. Orig. desig. 
*MacTostemma Shestakov, 1923. Acad. Sci. Russ. Mus. zool. Ann. 24: 
46. (Syn. teste Townes, 1957. Ent. Soc. Wash. Proc. 59 (3) : 107.) 
Type: Macrostemma elegans Shestakov. By monotypy and 
orig. desig. 

Genus BARYLYPA Foerster, pp. 396-397 

Berylypa Foerster. Place type cited in parentheses and add: = delictor 

(Thunberg). 
*Laphyctes Foerster, 1868. Naturh. Ver. Rheinlande Verb. 25: 146. 
Preocc. 

Type: Laphyctes insidiator Foerster. Incl. by Foerster, 1878 
(syn. teste Perkins, 1962, p. 434). 
*1Magnibucca Morley, 1913. Revis. Ichn. British Mus., v. 2, pp. 50, 79 
(syn.? teste Townes, Townes and Gupta, 1961, p. 313). 
Type: Magnibucca testacea Morley. By monotypy. 

relicta (Fabricius). Transf. from Alphabetic List of Unplaced Species, p. 409 

{teste Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 111). 

Genus CAMPOSCOPUS Foerster 

Camposcopus Foerster, 1868. Removed from syn. of Gravenhorstia Boie, 

p. 397 of Catalog. 
Labrorychus Foerster (not of auctt. or of Catalog). 

Type: Anomalon nigricorne Wesmael. Included by Brischke, 
1880. N. syn. (Isogenotypic with Blaptocampus Thomson.) 
*Blaptocampus Thomson, 1892. Opusc. Ent., fasc. 16, p. 1769 (syn. teste 
Townes, Townes, and Gupta, 1961, p. 312). 

Type: Anomalon nigricorne Wesmael. Desig. by Viereck, 
1914. 



210 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Place in Gravenhorstiini, p. 397, before Gravenhorstia Boie. 

aclerivorus Rohwer. Transf. from Gravenhorstia Boie, p. 397 (teste Townes, 
Townes, and Gupta, p. 312) . 

Genus GRAVENHORSTIA Boie, pp. 397-398 

Gravenhorstia Boie. Not in Nearctic fauna. According to Perkins (in 
Hit.) the two characters that distinguish the genus are a cen- 
tral, preapical tooth on the clypeus and a strong, central tooth on 
face just below lower line of antennal insertions. I have seen no 
Nearctic species with both of these characters. 

Camposcopus Foerster. Remove from syn. A valid genus (teste Townes, 
Townes and Gupta, 1961, p. 312). 

Erigorgus Foerster. Delete type desig. and remove from syn. A valid 
genus (teste Perkins, 1962, pp. 412, 422). 

Sympratis Foerster. Syn. of Erigorgus Foerster. 

ddoniopsis Foerster. Isogenotypic syn. of Gravenhorstia Boie. 

aclerivorus (Rohwer). Transf. to Camposcopus Foerster. 
ambigua (Norton). Transf. to Erigorgus Foerster. 
cerinops (Gravenhorst) . Transf . to Erigorgus Foerster. 
curtus (Norton). Transf . to Erigorgus Foerster. 
edwardsii (Cresson). Transf . to Erigorgus Foerster. 
eureka (Ashmead). Transf . to Erigorgus Foerster. 
ferruginea (Norton). Transf . to Erigorgus Foerster. 
iuUginosa (Davis). Transf. to Habronyx Foerster. 
luteopecta (Norton). Transf . to Erigorgus Foerster. 
magniceps (Cresson). Transf. to Habronyx Foerster. 
propinqua (Cresson). Transf. to Erigorgus Foerster. 
rotunda (Davis). Transf . to Erigorgus Foerster. 
rufula (Provancher). Transf. to Erigorgus Foerster. 
semirufa (Norton). Transf. to Erigorgus Foerster. 
suhurhis (Davis). Transf . to Erigorgus Foerster. 
xanthopsis (Ashmead). Transf. to Erigorgus Foerster. 

Genus ERIGORGUS Foerster 

Erigorgus Foerster. From syn. of Gravenhorstia, Boie, p. 397. 

Type: Anomalon fibulator Gravenhorst, 1829. Desig. by Per- 
kins, 1962. (Anomalon carinatum Brischke was only 
doubtfully included and not available for type desig.). 
Sympratis Foerster. Transf. from syn. of Gravenhorstia Boie. In type 
citation change ^'■Erigorgus" to: Anomalon. 

ambiguus (Norton). Transf. from Gravenhorstia Boie, p. 397. 

cerinops (Gravenhorst). Transf. from Gravenhorstia Boie, p. 397. 

curtus (Norton). Iowa, Ark. Transf. from Gravenhorstia Boie, p. 397. 

edwardsii (Cresson). Transf. from Gravenhorstia Boie, p. 397. 

eureka (Ashmead). Transf. from Gravenhorstia Boie, p. 397. 

ferrugineus (Norton). Transf. from Gravenhorstia Boie, p. 397. 

fuliginosus (Davis). Transf. from Gravenhorstia Boie, p. 397. 

luteopectus (Norton). Transf. from Gravenhorstia Boie, p. 397. 

?neglectum (Morley). Transf. from Therion Curtis, p. 400. 

propinquus (Cresson). Transf. from Gravenhorstia Boie, p. 397. 



ICHNEUMONIDAE 21] 

rotundus (Davis). Transf. from Gravenhorstia Boie, p. 397. 
rufulus (Provancher). Transf. from Gravenhorstia Boie, p. 398. 
semirufus (Norton). Transf. from Gravenhorstia Boie, p. 398. 
suburbis (Davis). Transf. from Gravenhorstia Boie, p. 398. 
xanthopsis (Ashmead). Transf. from Gravenhorstia Boie, p. 398. 

Genus AGRYPON Foerster, Sup. 1, p. 61 
alaskensis (Ashmead). B.C. Host: Melanolophia irnitataWlkr . 

Genus TRICHIONOTUS Cameron 
( = (.abror/chus of Authors, p. 398) 

Labrorychus Foerster of Catalog, and authors, not of type (Labrorychus 

Foerster of type is syn. of Camposcopus Foerster). 
*Trichonotus Cameron, 1905 (July). Straits Branch Roy. Asiatic Soc. 
44: 124. Preocc. Syn. of Labrorychus authors (teste Townes, 
Townes, and Gupta, 1961, p. 316). 

Type: Trichonotits reticulatus Cameron. By monotypy. 
*Trichionotus Cameron, 1905 (Aug.). Ann. Mag. Nat. Hist. (ser. 7) 16: 
168 (syn. teste Townes, Townes, and Gupta, 1961, p. 316). 
Type : Trichionotus reticulatus Cameron. By monotypy. 
*Phaenolabrorychus Viereck, 1913. U. S. Natl. Mus. Proc. 46: 379 (syn. 
teste Townes, Townes, and Gupta, p. 316). 

Type: Phaenolabrorychus cuisiisi Viereck. By monotypy and 
orig. desig. 
*Paragrypon Uchida, 1941. Ins. Matsumurana 15 (4) : 159 (syn. teste 
Townes, 1957. Ent. Soc. Wash. Proc. 59 (3) : 115). 

Type: Gongropelma kikuchii Uchida. Orig. desig. 
All species are n. combs, in Trichionotus Cameron. 

prismaticus (Norton), n. comb. Maine, Que., N. H., Vt., N. B., Ohio, Ont., 
Minn. Hosts: Archips cerasivoranus (Fitch); Ostrinia nubilalis 
(Hbn.) ; Pyrausta penitalis Grt. 

Genus HABRONYX Foerster 

Habronyx Foerster, 1860. Naturh. Ver. Rheinlande Verh. 17: 145. 
Type: Habronyx gravenhorstii Foerster. By monotypy. 

Place on p. 398, before Atrometus Foerster. 

fuliginosus (Davis), n. comb. Transf. from Gravenhorstia, p. 397. 

magniceps (Cresson), n. comb. Transf. from Gravenhorstia Boie, p. 397. 

Genus ATROMETUS Foerster, p. 398 

*Parania Morley, 1913. Revis. Ichn. Brit. Mus. v. 2, pp. 50, 96 (syn. teste 
Townes, Townes, and Gupta, 1962, p. 316). 

Type: Parania nototrachoides Morley. By monotypy. 

clavipes (Davis). Host: Argyrotaenia velutinana (Wlkr.). 

♦Tribe THERIONINI 
( = SubtrIbe Tberionina of Catalog) 

Genus THERION Curtis, pp. 399-400 

fuscipenne (Norton). 

Therion verticale (Morley) (syn. teste Townes, 1961. Ent. Soc. Wash. 
Proc. 63(2): 106). 

neglectum (Morley). Townes places this sp. in Gravenhorstia Boie (1961, 
Ent. Soc. Wash. Proc. 63 (2): 106). Probably belongs in Eri- 
gorgus Foerster as treated in this supplement. 

sassacus Viereck. Host: Pseudaletia unipunctata (Haw.). 



212 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

tenuipes (Norton). 

*Exochilum diabolus Morley, 1913. Revis. Ichn. Brit. Mus., Pt. 2, pp. 71, 
73. ?. (Syn. teste Townes, 1961. Ent. Soc. Wash. Proc. 63 (2): 
106.) 

verticale (Morley). Syn. of fuscipenne (Norton). 

Tribe OPHIONINI 

Genus OPHION Fabricius, pp. 401-402 

bifoveolatus BruUe. Transf. to Enicospilus Stephens, p. 404. 

Ophion nigrovarius Provancher. Remove from syn. ; a valid sp. 

nigrovarius Provancher. (Misspelled nigrivarius in Catalog.) A valid sp. 
(teste Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 106). 
Ophion novascotiae (Morley) (syn. teste Townes, loc. cit. supra). 

novascotiae (Morley). Syn. of nigrovarius Provancher, p. 402. 

stramineus Taschenberg. Transf. to Enicospilus Stephens, p. 404 as syn. of 
bifoveolatus (BruUe). 

Genus CHILOPHION Cushman, p. 403 
abnormus (Felt). Wyo. 

Genus ENICOSPILUS Stephens, pp. 403-405 

Dicamptus Sz^pligeti, 1905. Delete from syn. Valid genus. 
AmesospUus Enderlein, 1918 (1914-1916). Change type citation to: 
Ophion unicallosus Vollenhoven. Orig. desig. 

americanus (Christ). 

*Ophion rugosus Brulle, 1846. Hist. Nat. Ins. Hym., v. 4, p. 138 (syn. 
teste Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 111). 

bifoveolatus (Brulle). Transf. from Ophion Fabricius, p. 402 (teste 
Townes, 1961. Ent. Soc. Wash. Proc. 63 (2) : 111). 
Ophion stramineus Taschenberg. Transf. from Ophion Fabricius, p. 402 
(syn. teste Townes, loc. cit. supra). 

cuhitalis (Morley), preocc. Syn. of glabratus (Say). 

excubitalis Walkley. Syn. of glabratus (Say). 

glabratus (Say). 

Enicospilus cuhitalis (Morley). Preocc. Syn. 

Enicospilus excubitalis Walkley. N. name. (Syn. teste Townes, 1961. 
Ent. Soc. Wash. Proc. 63 (2) : 103.) 

Subfamily MESOCHORINAE 

Genus ASTIPHROMMA Foerster, p. 405 
strenuum (Holmgren). Host (secondary ?) : Melanolophila imitataWYkr. 

Alphabetic List of Unplaced Species, pp. 407—409, Sup. 1, p. 62 

Cryptus cinctipes Walsh. Transf. to Ischnus Gravenhorst, p. 270, placing 
alphabetically. 

Cryptus rhomboidalis Walsh. Transf. to Ischnus Gravenhorst, p. 270, placing 
alphabetically. 

Cryptus rufifrons Walsh. Transf. to Habrocryptoides Uchida (to be placed on 
p. 271). 

Cryptus semirufus Brulle. Transf. to Cubocephalus Ratzeburg, p. 255, plac- 
ing alphabetically. 

Eriborus triannulatus Cameron. Transf. to Campoplex Gravenhorst, p. 362. 

Exolytus concamerus Davis. Transf. to Oxytorus Foerster (= Callidiotes 
Foerster) as syn. of antennatus (Cresson), p. 340. 



MYMARIDAE 213 

Hemiteles clipeator Lundbeck. Transf. to Symplecis Foerster, p. 343. 

Hemiteles gastricus Holmgren. Transf. to *Charitopes Foerster. 

Ichneumon astutus Holmgren. Transf. to Cratichneumon Thomson, p. 287. 

Ichneumon banchiformis Megerle. Transf. to Cryptanura Brulle, p. 265. 

Ichneumon lugubrator Gravenhorst. Transf. to *Thyratele8 Perkins (to be 
placed on p. 296) . 

Ichneumon nigrosignatus Viereck. Transf. to Coelichneumon Thomson, plac- 
ing alphabetically. 

Labrorychus relictus (Fabricius). Transf. to Barylypa Foerster, p. 396. 

Limnerium hospitum Holmgren. Transf. to Diadegma Foerster (= Horo' 
genes of Catalog, p. 378). 

Phygadeuon plectiscinus Roman. Transf. to *Charitope8 Foerster. 

Pseudambly teles peroratus Cameron. Transf. to Ichneumon Linnaeus ( = 
Pterocormus Foerster of Catalog), p. 300. 

Trogus atrocoeruleus Cresson. Transf. to *ProtopeImus Heinrich (to be 
placed on p. 296). 

Superfamily CHALCIDOIDEA 

Taxonomy: Boucek in Kratochvil, 1957. Klic Zvir. C. S. R., v. 2, pp. 
204-245, 269-288.— Ferriere and Kerrich, 1958. Handb. Ident. 
Brit. Ins. (Roy. Ent. Soc. London), v. 8, pt. 2a, 40 pp. — Nikols- 
kaja, 1952 (Eng. trans. 1963). Keys to Fauna U.S.S.R. 44, 593 
pp. (N. S. F.-Smiths. Inst, pub., OTS 61-31220, U. S. Dept. 
Comm.).— Peck, 1963. Canad. Ent. Sup. 30, 1092 pp. (catalog).— 
Burks, 1964 (1963). Canad. Ent. 95: 1254-1263 (Provancher 
spp.). 

Biology: Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, 151 pp. (biolo- 
gical control).— Keen, 1958. U. S. Dept. Agr. Tech. Bui. 1169, pp. 
93-112 (seed and cone ins. of forest trees). — Schaffner, 1959. U. S. 
Dept. Agr. Misc. Pub. 767, 97 pp. (lepidopterous hosts and 
par.). — Hedqvist, 1963. Stud. Forest. Suecica, no. 11, pp. 7-28 
(par. of beetles in bark and wood) . 

Family MYMARIDAE 

Taxonomy: Debauche, 1948. Mus. Roy. Hist. Nat. Belg. Mem. 108: 
1-248. — Debauche, 1949. Cong. Belg., Inst. Pares Nat., Expl. Pare 
Nat. Albert, Miss, de Witte, pt. 49, pp. 1-105. — Annecke and 
Doutt, 1961. So. Afr. Dept. Agr. Tech. Ser. Ent. Mem. 5: 1-71 
(generic classif.). 

Genus OOCTONUS Haliday, pp. 410-411 

americanus Girault. N. J., Ohio. Host: Eggs Philaenus leucophthalmus 
(Linn.). 

aphrophorae Milliron. Ohio. 

Genus COSMOCOMOIDEA Howard, p. 411 

Taxonomy: Debauche, 1948, loc. cit. supra, p. 114 (syn. of Ooctonus 
Haliday).— Ogloblin, 1959. Munch. Ent. Ges. Mitt. 49: 50 (subg. 
of Lymaenon Walker) . 

morrilli Howard. Ga., Tex.; Mexico, Cent. Amer. Host: Homalodisca coagu- 
lata (Say). 

Genus CAMPTOPTERA Foerster, p. 411 

*Stichothrix Foerster, 1856. Hym. Stud., v. 2, pp. 117-121. 
Type : Stichothrix cardui Foerster. By monotypy. 



214 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*CongoUa Ghesquiere, 1942. Rev. Zool. Bot Afr. 36: 320. 

Type : Congolia sycophila Ghesquiere. By monotypy. 
Taxonomy: Debauche, 1949, loc. cit. supra, pp. 4, 17. — Annecke and 
Doutt, 1961, loc. cit. supra, p. 15. 

Genus GONATOCERUS Nees, pp. 411-413 
( = Genus Lymaenon Haliday, Sup. 1, p. 63, cited in error) 

*Rachistus Foerster, 1847. Linn. Ent., v. 2, p. 203. 

Type: Ooctonus litoralis Haliday. Desig. by Gahan and Fagan, 
1923. 
*Oophilus Enock, 1909. Roy. Ent. Soc. London Trans. 1909, p. 458. 

Type : OophUus longicauda Enock. By monotypy. 
Taxonomy: Debauche, 1949, loc. cit. supra, pp. 4, 25 (Lymaenon). 

ashmeadi Girault. Ga., Miss., La., Tex. Hosts: Eggs Oncometopia orbona 
(Fab.), Homalodisca coagulata (Say), Cuerna costalis (Fab.). 

fasciatus Girault. Ga. Host: Homalodisca coagulata (Say). 

koebelei Perkins. Fla.; Mexico. 

marylandicus Girault. Ga. Host: Gypona S'p. 

novifasciatus Girault. Ga. Host: Graphocephala versuta (Say) . 

*triguttus Girault. Tex. ; West Indies. Host: Leaf hopper eggs. 

Gonatocerus triguttus Girault, 1916. Ent. Soc. Amer. Ann. 9: 297. 9, $. 

Genus ALAPTUS Haliday, p. 413 

*Parvnlinus Mercet, 1912. R. Soc. Esp. Hist. Nat. Bol. 12: 331. 

Type: Parvulinus auranti Mercet. By monotypy. 
Taxonomy: Girault, 1912. Ent. Soc. Wash. Proc. 14: 221. — Debauche, 
1949, loc. cit. supra, p. 9. 

*auranti (Mercet). La.; Europe. Hosts: Several spp. of Coccidae question- 
ably recorded ; true hosts prob. Psocoptera. 

Parvulinus auranti Mercet, 1912. R. Soc. Esp. Hist. Nat. Bol. 12: 333. 9. 

Metalaptus torquatus Malenotti (syn. by Mercet, 1917. Asoc. Esp. Prog. 
Ciencia, Cong, de Valladolid, sect. 4, p. 367). 

torquatus (Malenotti). Syn. of auranti (Mercet). 

Genus ERYTHMELUS Enoci<, pp. 413-414 

*Enaesius Enock, 1909. Roy. Ent. Soc. London Trans. 1909, p. 456. 

Type: Enaesius agilis Enock. Desig. by Gahan and Fagan, 
1923. 
Taxonomy: Annecke and Doutt, 1961, loc. cit. supra, p. 18. 

Genus PATASSON Walker, p. 414 

*Antoniella Soyka, 1950. Ent. Nachr. Burgdorf 3: 121. 

Type: Fulmekiella slubaiensis Soyka. Grig, desig. 
*Mariella Soyka, 1950. Ent. Nachr. Burgdorf 3 : 123. 

Type: Mariella superaddiia Soyka. Grig, desig. 
Taxonomy: Debauche, 1948, loc. cit. supra, p. 172 (Patasson a subg. of 
Anaphes Haliday). — Annecke and Doutt, 1961, loc. cit. supra, p. 
21 (syn.). 

conotracheli (Girault). Delete luna Girault from syn. 

luna (Girault). N. Y., N. J., Del., Md., Utah; Europe. Host: Hypera postica 
(Gyll.) eggs. 
Anaphoidea luna Girault, 1914. Erroneously placed as a syn. of conotra- 
cheli (Girault) in Catalog. Types re-examined. 
Biology: Clausen, 1956, loc. cit. supra, p. 116. 

Genus ANAGRUS Holiday, p. 415 

Pteratomus Packard, 1864. Transf. from Unplaced Genera, p. 593. 
*Packardiella Ashmead, 1904. Carnegie Mus. Mem. 1: 364. N. name, 



MYMARIDAE 215 

unnecessarily proposed, for Pteraiomus Packard. 
*Paranagrus Perkins, 1905. Hawaii. Sug. PI. Assn. BuL, Ent. Ser., 1: 
199. 

Type: Paranagrus optabilis Perkins. Desig. by Gahan and 
Fagan, 1923. 
Taxonomy: Annecke and Doutt, 1961, loc. cit. supra, p. 7. 

armatus nigriceps Girault. Calif. Host: Edwardsiana prunicola (Edw.). 
Biology: MuUa, 1956. Jour. Econ. Ent. 49: 438. 

epos Girault. Fla. Hosts: Edwardsiana prunicola (Edw.), Typhlocyba qu- 
erc2is (F.). 
Biology: Mulla, 1956. Jour. Econ. Ent. 49: 438. 

giraulti Crawford. Ga., Ala. Host: Homalodisca coagiilata (Say). 

putnamii (Packard). Transf. from Unplaced Species, Sup. 1, p. 84. 
Taxonomy: Annecke and Doutt, 1961, loc. cit. supra, p. 8. 

Genus ANAPHES Holiday 
(ZlAlymar Curtis of Catalog, pp. 415—416) 

*Ci;nomymar Kieffer, 1913. Soc. Agr. Alger. Bui. 4: 100. 

Type: Clinomymar peyeriinhoffi Kieffer. Orig. desig. 
*Stammeriella Soyka, 1950. Ent. Nachr. Burgdorf 3: 120. 

Type: Siammeriella wolfslhali Soyka. Orig. desig. 
Taxonomy: Debauche, 1949, loc. cit. supra, p. 63. — Annecke and Doutt, 

1961, loc. cit. supra, pp. 18-20. 
Application was made to Int. Comn. Zool. Nomencl. to preserve usage of 
Mymar Curtis in traditional sense. This requires substitution of Anaphes 
(Type: Ichneumon punctuni Shaw) for Mymar of Catalog, and Mymar for 
Myinarilla of Catalog. See Doutt and Annecke, 1963, Bui. Zool. Nomencl. 20: 
134-136. Opinion 729, Bui. Zool. Nomencl., 22: 82, 1965 authorizes this 
change, so traditional usages are restored in this Catalog for Anaphes Hali- 
day and Mymar Curtis. 

*alaskae Annecke and Doutt. Alaska. 

Anaphes alaskae Annecke and Doutt, 1961. So. Afr. Dept. Agr. Tech. 

Ser. Ent. Mem. 5: 47. 9, $. 

iole Girault. Calif. 

pratensis Foerster. 

Biology: Clausen, 1956, loc. cit. supra, p. 116. 

Genus ACMOPOLYNEMA Ogloblin, p. 416 

bifasciatipenne bifasciatipenne (Girault), corrected spelling of bifasciati- 
pennis bifasciatipennis of Catalog. 

bifasciatipenne varium (Girault), corrected spelling of bifasciatipennis »a- 
rium of Catalog. 

Genus BARYPOLYNEMA Ogloblin, p. 416 

Taxonomy: Annecke and Doutt, 1961, loc. cit. supra, p. 60. 

*niedicae Annecke and Doutt. D. C, Calif. 

Barypolynema medicae Annecke and Doutt, 1961. So. Afr. Dept. Agr. 
Tech. Ser. Ent. Mem. 5: 57. 9, $. 

Genus POLYNEMA Holiday, pp. 416-418 

*MaidlieUa Soyka, 1946. Zentbl. Ges. Ent. 1 : 178. 

Tvpe: Maidliella neofuscipes Sovka. Orig. desig. 
*I\ovickyeUa Soyka, 1946. Zentbl. Ges. Ent. 1: 179. 

Type : ISovickyella gracilior Soyka. Orig. desig. 
Delete Stephanodes Enock from syn. 

Taxonomy: Soyka, 1956. Zool.-Bot. Ges. Wien Abhandl. 19: 32.— An- 
necke and Doutt, 1961, loc. cit. supra, pp. 34-36, 60. 
Nomenclature: Peck, 1963, loc. cit. siipra, pp. 42-49 (sp. names changed 
to neuter case) . 



216 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

bimaculatipenne Girault, corrected spelling of bimaculatipennis of Catalog. 

boreum Girault, new status. Delete from syn. under striaticorne Girault, as 
var. boreum Girault, 1916, and ch. to valid sp., teste D. P. An- 
necke, in litt. 

caesariatipenne Girault, corrected spelling of caesariatipennis of Catalog. 

graculus Girault. [Name from L. subst., graculus, a jackdaw.] 

mymaripenne Dozier, corrected spelling of mymaripennis of Catalog. 

psecas Girault. Transf. to Stephanodes Enock. 

striaticorne Girault. Delete striaticorne var. boreum Girault from syn., and 
ch. to sp. 

Genus STEPHANODES Enock 

Delete from syn. of Polynema Haliday, teste Annecke and Doutt, 1961, 
loc. cit. supra, p. 34-35. Insert following Polynema Haliday on p. 418. 

psecas (Girault), n. comb. 111., Iowa. Trans, from Polynema Haliday; types 
examined. 

Genus CARAPHRACTUS Holiday, p. 418 

cinctus Haliday. N. W. T. (Southampton Isl.). Hosts: Eggs of Dytiscidae. 
Biology: Jackson, 1958. Roy. Ent. Soc. London Trans. 110: 533-566. 

Genus NEOMYMAR Crawford, p. 418 
vierecki vierecki Crawford. Fla., Iowa. 

Genus MYMAR Curtis 
(=:/Hymori//a of Cafalog, pp. 418-419) 

*Pterolinononyktera Malac, 1943. Ent. listy (Folia Ent.) Brno 6: 51. 
Type: Pterolinononyktera obenbergeri Malac. Orig. desig. 

Taxonomy: Annecke and Doutt, 1961, loc. cit. supra, pp. 26-29. 

Application was made to Int. Comn. Zool. Nomencl. to preserve tradi- 
tional usage of Mymar Curtis (Type: Mymar pulchellum Curtis). See Doutt 
and Annecke, 1963, loc. cit. supra, pp. 134-136. Opinion 729, Bui. Zool. No- 
mencl. 22: 82, 1965 takes favorable action on this application, so traditional 
usage for Mymar Curtis is restored in this Catalog. Mymarilla Westwood is 
a valid, but extralimital genus, different from Mymar, teste Annecke and 
Doutt, 1961, loc. cit. supra, pp. 31-32. 

Cincinnati Girault. 

Taxonomy: Annecke, 1961. So. Afr. Jour. Agr. Sci. 4: 551-552 (prob. 
syn. of Mymar regale Enock). 

^pulchellum Curtis. Pa.; Europe. 

Mymar pulchellus Curtis, 1832. Brit. Ent. 9: 411. 9, $. 
Mymar venustum Girault, 1911 (syn. by Annecke, 1961, loc. cit. supra, 
pp. 550-551). 

venustum Girault. Syn. of pulchellum Curtis. 

Genus ARESCON Haliday, p. 419 

*Xenomymar Crawford, 1913. U. S. Natl. Mus. Proc. 46: 349. 

Type: Xenomymar urichi Crawford. Orig. desig. 
*lSeurotes Enock, 1914. Roy. Ent. Soc, London, Proc. 1913, p. cxxxiv. 

Type : Neurotes iridescens Enock. By monotypy. 
Taxonomy: Annecke and Doutt, 1961, loc. cit. supra, p. 12. 

Family TR8CHOGRAMMATIDAE 

Genus TRICHOGRAMMA Westwood, pp. 419-421 

Taxonomy: Quednau, 1956. Ber. Hundertjahr. Deut. Ent. Ges. Berlin, 
pp. 87-92. — Quednau, 1956. Ztschr. Pflanzenkrank. und Pflan- 



TRICHOGRAMMATIDAE 217 

zensch. 63: 333-344.— Quednau, 1960. Biol. Bund, fur Land-und 

Forstwirtsch. Mitt. 100: 11-50. 

evanescens Westwood. Delete; valid but extralimital sp., teste Quednau, 
1960, loc. cit. supra, p. 31. 

*fasciatum (Perkins). Fla., La., South. Tex.; Mexico, West Indies, Cent, and 

S. America. 
Pentarthron fasciatum Perkins, 1912. Hawaii. Sugar Planters' Assn., 

Ent. Ser. Bui. 10, p. 19. 
Biology: Burrell and McCormick, 1962. Jour. Econ. Ent. 55: 880. 
Taxonomy : Quednau, 19G0, loc. cit. supra, p. 33. 

minutum Riley. South. Canada and U. S. north of Gulf Coast. Hosts: Eggs 
of over 160 spp., mostly Lepidoptera. 
Biology: Langston, 1957. Univ. Calif. Pubs., Ent. 14: 25-26.— Downey, 
1958 (1957). 111. State Acad. Sci. Trans. 50: 299.— Schaffner, 
1959, loc. cit. supra, p. 81. — Glass, 1963. Ent. Soc. Amer. Ann. 56: 
564. 
Taxonomy: Quednau, 1960, loc. cit. supra, p. 34. 

retorridum (Girault). Southwest. U. S. north to Iowa. 
Taxonomy: Quednau, 1960, loc. cit. supra, p. 31. 

*semifumaium (Perkins). South. Tex., South. Calif.; Mexico, Hawaii. 

Pentarthron semifumatum Perkins, 1910. Fauna Hawaii., vol. 2, pt. 6, p. 

659. 9. 
Biology: Hinckley, 1961. Jour. Econ. Ent. 54: 207. — Stern and Bowen, 

1963. Ent. Soc. Amer. Ann. 56: 358-372.— Poinar, 1964. Jour. 

Econ. Ent. 57: 381. 
Taxonomy: Quednau, 1960, loc. cit. supra, p. 33. 

semblidis (Aurivillius). Ont. and N. Y. west to Sask.; Europe. Hosts: Eggs 
of aquatic and subaquatic insects. 
Biology: Quednau, 1957. Deut. Ent. Ztschr. 4: 364. 
Taxonomy: Quednau, 1960, loc. cit. supra, p. 29. 

Genus APHELINOIDEA Girault, p. 421 

plutella Girault. 

Biology : Henderson, 1955. U. S. Dept. Agr. Cir. 967, pp. 1-16.— Flock et 
al, 1962. Jour. Econ. Ent. 55: 279. 

Genus LATHROMEROIDES Girault, pp. 421-422 
neomexicanus Girault. Iowa, Calif. 

Genus OLIGOSITA Haliday, pp. 422-423 
americana Girault. Ga. Host: Homalodiscainsolita {Wa-Da.) . 
sanguinea sanguinea (Girault). Calif. 

Genus UFENS Girault, p. 423 

niger (Ashmead). Ga., Utah, Wash., Calif. Hosts: CoUadonus geminatus 
(Van Duzee), Cuerna costalis (Fab.), Keonolla confluens (Uhl.). 

♦osborni Dozier. Fla.; West Indies. Hosts: Eggs of Pachnaeus litus (Germ.), 
Diaprepes abbreviatus (L.) ? secondary. 
Vfens osborni Dozier, 1932. Ent. Soc. Wash. Proc. 34: 36. 9, $. 
Biology: Tucker, 1936. Barbados Agr. Jour. 5: 9. — Baranowski, 1960. 
Fla. Ent. 43: 197. 

spiritus americanus Girault. Ga. Host: Homalodisca coagulata (Say). 

spiritus spiritus Girault. Ga., Fla. Hosts: Homalodisca coagulata (Say), On- 
cometopia orbona (Fab.). 

Genus HYDROPHYLITA Ghesqui^re, pp. 423-424 
In generic ref. change v. 34 to v. 39. 



218 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

aquivolans (Matheson and Crosby). Ohio, Kans. Host: Eggs of Ischnura 
verticalis (Say). 
Biology: Davis, 1962. Limnol. and Oceanog. 7: 390-392. 

Genus ABBELLA Girault, pp. 424-425 

acuminata (Ashmead). Ga., La., Minn., Tex., Calif. Hosts: Cuerna costalis 
(Fab.), DraeculacepJutla portola (Ball), Homalodisca insolita 
(Walk.). 

ceresarum (Ashmead). Calif. 

subflava Girault. 

Biology: Henderson, 1955. U. S. Dept. Agr. Cir. 967, pp. 1-16.— Flock et 
al, 1962. Jour. Econ. Ent. 55: 279. 

Family EULOPHIDAE 

Subfamily EULOPHINAE 

Genus PNIGALIO Schrank, pp. 425-426 

flavipes (Ashmead). Ohio, Minn. 

maculipes (Crawford). N. B., Que. Hosts: Phytagromyza populicola 
(Walk.), Bucculatrix ainsliella Murt., B. packardella Chamb. 
Biology: Schaffner, 1959, loc. cit. supra, p. 81. 

pallipes (Provancher). Ont. Transf. from Eulophus Olivier teste Burks, 
1964, loc. cit. supra, p. 1257. 

tischeriae (Ashmead). Ont. Host: Lithocolletis salicifoliella Clem. 

Genus SYMPIESIS Foerster, pp. 426-428 

*Teleogmus Foerster, 1856. Hym. Stud., v. 2, pp. 72, 74. 

Type : Teleogmus orbitalis Foerster. By monotypy. 
*Moroceras Erdos, 1954. Mus. Nat. Hung. Ann. Hist.-Nat., n. s., 5: 323. 

Type: Moroceras hiroi Erdos. Orig. desig. 
*Sympiesis subg. Cladosympiesis Graham, 1959. Soc. Brit. Ent. Trans. 
13: 182. 

Type: Eulophus gordius Walker. Orig. desig. 
Taxonomy: Boucek, 1959. Mus. Nat. Pragae Acta Ent. 33: 121 (subgen- 
era). — Delucchi, 1962. Awamia 5: 60 (disc, of Moroceras Erdos). 
—Graham, 1963. Soc. Brit. Ent. Trans. 15: 187 (syn.). 

ancylae Girault. Minn. 

Biology: Balduf, 1959. 111. Biol. Monog. 26, p. 146. 

bimaculatipennis (Girault). Ont., Wis., Minn., Oreg., Calif. 

compressicornis (Provancher). N. S., Maine and Ont. south to Va. west to 
Kans., Idaho, and Oreg. Hosts: Lithocolletis crataegella Clem., L. 
salicifoliella Chamb., L. caryaefoliella (Clem.), leaf miners on 
Amelanchier, maple, oak, black cherry. 
Sympiesis nigripes Ashmead, 1888 (syn. by Burks, 1964, loc. cit. supra, p. 
1257). 

conica (Provancher). Que. and Maine south to D. C, west to Idaho and Tex. 
Hosts: Listed under nigrifemora Ashmead in Catalog, p. 427. 
Sympiesis nigrifemora Ashmead, 1888 (syn. by Burks, 1964, loc. cit. 

supra, p. 1259). 
Biology: Schaffner, 1959, loc. cit. supra, p. 81 (nigrifeviora) . 

dolichogaster Ashmead. Holarctic. 

Taxonomy: Boucek, 1959. Mus. Nat. Pragae Acta Ent. 33: 130. 

guttatipennis Girault. Mass., N. C. Host: Exoteleia pinifoliella (Chamb.). 
Biology: Schaffner, 1959, loc. cit. supra, p. 81. 

lexingtonensis Girault. Syn. of miltoni Girault. 



EULOPHIDAE 219 

marylaiidica Girault. Wis., Minn. 

miltoni Girault. Maine south to W. Va., west to Wis. and Ky. Hosts: Litho- 
colletis spp. 
Sympiesis mil.oni Girault, May 1917. 
Sympiesis lexingtonensis Girault, Aug. 1917. N. syn. ; types examined. 

nigrifem ora Ashmead. Syn. of conica (Provancher) . 

nigripes Ashmead. Syn. of compressicornis (Provancher). 

siigmata Girault. Oreg. Host: Leaf miner on apple. 

tricladus (Provancher). Transf. from Eulophus Olivier teste Burks, 1964, 
loc. cit. supra, p. 1258. 

viridula (Thomson). Ont. and N. Y. west to Mo. and Wyo. 
Biology: Clausen, 1956, loc. cit. supra, pp. 129-130. 

Genus NECREMNUS Thomson, p. 428 

comptus Gahan. 

Taxonomy: Boucek, 1959. Mus. Nat. Pragae Acta Ent. 33: 150 (? syn. 
of artynes Walker) . 

duplicatus Gahan. Idaho. 

Biology: Walz, 1957. Ent. Soc. Amer. Ann. 50: 220. 

Taxonomy: Boucek, 1959, loc. cit. supra, p. 152 (? syn. of tidius 
Walker). 

Genus HEMIPTARSENUS Westwood, p. 428 

*Cleolophus Mercet, 1924. R. Soc. Esp. Hist. Nat. Bol. 24: 461-462. 

Type : Cleolophus autonomus Mercet. Orig. desig. 
Taxonomy: Boucek, 1959, Mus. Nat. Pragae Acta Ent. 33: 144 (syn.). — 
Hedqvist, 1963, loc. cit. supra, p. 142. 

*Genus DIGLYPHUS Walker 
( = So/enofus Foerster of Catalog, p. 429) 

Diglyphus Walker, 1844. Ann. Mag. Nat. Hist. 14 : 409. 

Type : Cirrospilus chabrias Walker. By monotypy. 

Taxonomy: Ferriere, 1952. Soc. Venez. Stor. Nat. Bol. 6: 176 (viridis 
Foerster, type sp. of Solenotus Foerster = Cirrospilus isaea 
Walk.).— Graham, 1959. Soc. Brit. Ent. Trans. 13: 178 (isaea 
Walk, transf. to Diglyphus Walk.). 

begini (Ashmead). Que. Host: Phytomyza atricornis Meigen, P. aquilegiae 
Hardy. 
Biology: Doutt, 1957. Jour. Econ. Ent. 50: 373-374. 

intermedius (Girault). Fla., 111. Host: Liriomyza properpusilla Yrosi. 

Genus EULOPHUS Olivier, Sup. 1, p. 64 
(=:Eu/ophus GeofFroy of Catalog, pp. 430—431) 

*Comedo subg. Onychocomedo Graham, 1959. Soc. Brit. Ent. Trans. 13: 

183. 

Type: Eulophus thespius Walker. Orig. desig. 
Taxonomy: Boucek, 1959, Mus. Nat. Pragae Acta Ent. 33: 160. — Gijs- 

wijt, 1964. Ent. Ber. 24: 31 {Eulophus Muller, 1764). 

anomocerus (Crawford). Que., Ont., N. Y., Va., Mich., N. Dak. 

magnisulcatus Girault. Maine south to W. Va. Host: Coleophora laricella 
Hbn. 
Biology: Schaffner, 1959, loc. cit. supra, p. 81. 

nebulosus (Provancher). Ont. and Mass. south to Fla., west to Wis. 
Taxonomy: Burks, 1964, loc. cit. supra, p. 1259. 



220 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

orgyiae (Fitch). Que., Miss., Wis., Wash., Oreg., Calif. Hosts: Acronicta 
grisea Walk., Alsophila povietaria Hbn., Gracillaria rhoifoliella 
Cham., Gluphisia septentrionis Walk., Hemerocampa plagiata 
Walk., Orthosia hibisci Gn. 

pallipes (Provancher). Transf. to Pnigalio Schrank; delete smerinthi Ash- 
mead from syn. 

smerinthi (Ashmead). Correct name for species listed as pallipes Provancher 
in Catalog teste Burks, 1964, loc. cit. supra, p. 1257. 

tricladus Provancher. Transf. to Symplesis Foerster. 

Genus DAHLBOMINUS Hincks, p. 431 

fuscipennis (Zetterstedt). Md., Va., 111., Mo., Minn.; Europe. Hosts: Neodi- 
prion lecontei (Fitch), N. nanulus Schedl, N. pinetiim (Nort.), N. 
pratti (Dyar), N. sertifier (Geof.), N. tsugae Midd., Pristiphora 
erichsoni (Htg.). 

Biology: Burnett, 1956. Ent. Soc. Amer. Ann. 49: 55. — Clausen, 1956, 
loc. cit. supra, p. 101-102. — Riordan, 1957. Canad. Jour. Zool. 35: 
603.— Burnett, 1958. Canad. Ent. 90: 279.— Finlayson and Finlay- 
son, 1958. Canad. Ent. 90: 558.— Finlayson, 1960. Canad. Ent. 
92: 38 ifuliginosus (Nees) [!]).— Szmidt, 1960. Anz. f. Scha- 
dlingsk. 33: 20.— Szmidt, 1961. Poznan. Wyzsza Skola Roln. Rocz. 
Dodatek 6: 11.— Bobb, 1962. Brooklyn Ent. Soc. Bui. 57: 102.— 
Finlayson, 1963. Canad. Ent. 95: 498. — Morris, Schroeder, and 
Bobb, 1963. Pine Sawfly in Va., Va. Div. For., pp. 15-16. 

Taxonomy: Finlayson, 1960, loc. cit. supra, pp. 37, 38, 44 (imm. stages). 

Genetics: Wilkes, 1964. Science 144: 305-307 Uuliginosus (Nees) [!]). 

Genus DIMMOCKIA Ashmead, p. 431 

incongrua (Ashmead). Ont. and Mass. south to Fla., Ohio, W. Va., 111. 
Hosts: Anacampsis innocuella (Zell.), Argyrotoxa semipurpu- 
rana (Kearf.), Episimus argutanus (Clem.) ; almost always a se- 
condary par. 
Biology : Schaffner, 1959, loc. cit. supra, p. 82. 

pallipes Muesebeck. 111., Wis. Hosts: Exartema ferriferanum (Walk.); 
Sparganothis pettitana (Rob.) ; prob. always secondary. 

Genus OICLADOCERUS Westwood, p. 431 

*Diglyphis Thomson, 1878. Hym. Scand., v. 5, p. 208. [Not Diglyphus 
Walker.] 

Type: Diglyphis aeniscapus Thomson. Desig. by Gahan and 
Fagan, 1923. 
Taxonomy: Graham, 1959. Soc. Brit. Ent. Trans. 13: 180. 

Subfamily APHELININAE 

Taxonomy: Flanders, 1955. Ent. Soc. Amer. Ann. 46: 84-93 (biol. crite- 
ria in tax.). — Compere and Annecke, 1961. Ent. Soc. S. Afr. Jour. 
24: 19-20 (fam. Aphelinidae). — Peck, 1963, loc. cit. supra, p. 239 
(fam. Aphelinidae). 

Genus ERETMOCERUS Haldeman, p. 432 

haldemani Howard. 111. Host: Trialeurodes ahutilonea (Hald.). 

*portoricensis Dozier. Fla.; West Indies. Host: Aleurothrixus floccosus 
(Mask.). 
Eretmocerus portoricensis Dozier, 1932. Ent. Soc. Wash. Proc. 34: 115. 

?. 

Genus EUDEROMPHALE Girault, p. 432 
(= Cardiogaster Motschulsky, Sup. 1, p. 64, cited in error) 

Taxonomy: Compere and Annecke, 1961. Ent. Soc. S. Afr. Jour. 24: 



EULOPHIDAE 221 

65-68 (Cardiogaster in Eulophidae). — Peck, 1963, loc. cit. supra, 
p. 238 {Cardiogaster in Entedontinae). Graham, 1963. Soc. Brit. 
Ent. Trans. 15: 273 {Euderonipliale in Entedontinae). 

"hyalina (Compere and Annecke), n. comb. Calif. Host: Aleurodes sp. 

Cardiogaster hyalina Compere and Annecke, 1961. Ent. Soc. S. Afr. 

Jour. 24: 68. ?. 
Biology: Poinar, 1964. Jour. Econ. Ent. 57: 382. 

Genus ARCHENOMUS Howard, p. 432 
{ — Pteroptrix Westwood of Sup. 1, p. 64) 

Taxonomy: Alam, 1956. Roy. Ent. Soc, London, Trans. 108: 374. — No- 
vitzky, 1961. Ent. Monthly Mag. 97: 193.— Peck, 1963, loc. cit. 
supra, pp. 241, 287. 

Genus MARIETTA Motschulsky, p. 433 

caniesi (Howard). D. C; Japan, China, India. 
Biology: Azim, 1961. Mushi 35: 107. 

*graminicoIa Timberlake. Md., Fla., Tex.; Hawaii. Hosts: Anagyrus anton- 
inae Timb., poss. other primary parasites of grass scales. 

Marietta graminicola Timberlake, 1925. Hawaii. Ent. Soc. Proc. 6: 180. 

9, 5. 
Biology: Dean and Schuster, 1958. Jour. Econ. Ent. 51: 364. 

mexicana (Howard). Md. Host: Thysanus merceti (Malen.). 

Biology: DeBach, Kennett, and Pence, 1958. Jour. Econ. Ent. 51: 115. 
pulchella (Howard). Sask. Hosts: Sec. parasite on Coccidae. 

Biology: Peterson, 1962. Canad. Ent. 94: 740. 

Genus MARLATTIELLA Howard, Sup. 1, p. 65 

prima Howard. Del.; Japan. 

Taxonomy: Tachikawa, 1962. Shikoku Ent. Soc. Trans. 7: 74. 

Genus APHYTIS Howard, pp. 433-434 

Biology: DeBach, 1959. Ent. Soc. Amer. Ann. 52: 354-359 (biol. con- 
trol). 

abnormis (Howard). 

Taxonomy: DeBach and Azim, 1962. Mushi 36: 1. 

chrysomphali (Mercet). 

Biology : Smith, 1957. Canad. Ent. 89: 219.— Muma, 1959. Jour. Econ. 
Ent. 52: 581.— DeBach and Sisojevic, 1960. Ecology 41: 153. 

citrinus Compere. Host: ? Aonidiella aurantii (Mask.). 

Taxonomy: DeBach, 1959, loc. cit. supra, pp. 360-361 (Japanese strain). 

*coheni DeBach. Calif.; Israel. Hosts: Aonidiella aurantii (Mask.), A. ci- 
trina (Coq.) , Chrysomphalus dictyospermi (Morg.). 
Aphytis coheni DeBach, 1960. Ent. Soc. Amer. Ann. 58: 705. $,5. 
Biology: Fleschner, 1963. Pan-Pacific Ent. 39: 116. 

*fishen DeBach. Calif.; Burma. Host: Aonidiella aurantii (Mask.). 
Aphytis fisheri DeBach, 1959. Ent. Soc. Amer. Ann. 52: 362. ?, $. 
Biology: DeBach and Landi, 1959. Calif. Agr. 13: 9. 

*hispanicus (Mercet). Fla., Tex., Calif.; Mexico, Europe, Orient. Hosts: 
Parlatoria pergandii Comst., ? Aonidiella aurantii (Mask.). 

Aphelinus maculicornis var. hispanica Mercet, 1912. Mus. Cien. Nat. Ma- 
drid Trab., ser. Zool., 10: 81-82. 

Taxonomy: Compere, 1955. Cahf. Univ. Pubs., Ent. 10: 289, 294 (sp.). — 
DeBach, 1959, loc. cit. supra, pp. 359-360 (Assam form). 

*holoxanthus DeBach. Fla., Tex.; China, Israel, Host: Chrysomphalus aoni- 
diim (L.). 
Aphytis holoxanthus DeBach, 1960. Ent. Soc. Amer. Ann. 53: 704. 9, $. 
Biology: Clancy et at., 1963. Jour. Econ. Ent. 56: 603-605. 



222 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

lepidosaphes Compere. Fla., Tex.; Mexico. 

Biology: Clausen, 1956, loc. cit. supra, p. 83 (sp. X). — DeBach, 1959, loc. 
cit. supra, p. 355. — Dean, 1961. Ent. Soc. Amer, Ann. 54: 918. — 
DeBach and Landi, 1961. Hilgardia 31: 461. 

lingnanensis Compere. 

Biology: Clausen, 1956, loc. cit. supra, p. 82 (sp. A). — DeBach and Si- 

sojevic, 1960, loc. cit. supra, p. 153. — DeBach and White, 1960. 

Calif. Agr. Expt. Sta. Bui. 770, p. 4. 
Taxonomy: DeBach, 1959, loc. cit. supra, p. 360. 

maculicornis (Masi). 

Biology: Clausen, 1956, loc. cit. supra, p. 82. — Huffaker et al., 1962. 

Hilgardia 32 : 569. 
Taxonomy: Doutt, 1954. Jour. Econ. Ent. 47: 39-40.— Hafez and Doutt, 

1954. Canad. Ent. 86: 90.— Sugonyaev, 1962. Akad. Nauk 

SSSR Izv., ser. Biol., 5: 755. 

*melinus DeBach. Calif.; India, Pakistan. Host: Aonidiella aurantii 
(Mask.). 
Aphytis melinus DeBach, 1959. Ent. Soc. Amer. Ann. 52: 361. 9, $. 
Biology: DeBach and Landi, 1959, loc. cit. supra, p. 9. 

mytilaspidis (LeBaron). 

Biology: Flanders, 1957. Jour. Econ. Ent. 50: 171 (Italian strain). — 
Flanders, 1960. Jour. Econ. Ent. 53: 757.— Tadic, 1961. Arh. za 
Poljopriv. Nauke 14: 118. — Tschumakova, 1961. Ent. Obozr. 40: 
317. 

Taxonomy: DeBach and Azim, 1962. Mushi 36: 1 (group char.). 

proclia (Walker). 

Taxonomy: DeBach, 1959, loc. cit. supra, p. 358 (diaspidis Howard). — 
Tschumakova, 1960. Leningrad. Vsesoi. Inst. Zasch. Rast. Trudy 
15: 57.— Benassy, 1961. Ann. Epiphyties, ser. C, 12: 115, 127.— 
Tschumakova, 1961, loc. cit. supra, p. 316. 

Genus APHELINUS Dalman, pp. 434-435 

mail (Haldeman). 

Biology: Evenhuis, 1958. Tijdschr. Plantenziekten 64: 3. — Lung, Wang, 
and Tang, 1960. Acta Ent. Sinica 10: 6.— Evenhuis, 1962. Ento- 
mophaga 7: 215. 

nigritus Howard. 111., Okla., Tex. (northern loc. in greenhouses). 
Biology: Wood, 1958. Jour. Econ. Ent. 51: 553. 

semiflavus Howard. [Prob. a composite sp.] Maine, Calif. Host: Ther- 

ioaphis maculatus (Buck.). 
Biology: van den Bosch, 1957. Jour. Econ. Ent. 50: 353. — Schlinger and 

Hall, 1959. Jour. Econ. Ent. 52: 154.— Barnes, 1960. Jour. Econ. 

Ent. 53: 1094.— Schaefers and Allen, 1962. Hilgardia 32: 426.— 

Force and Messenger, 1964. Ent. Soc. Amer. Ann. 57 : 405-413. 
Taxonomy: Janssen, 1961. Beitr, z. Ent. 11: 672 

Genus MESIDIA Foerster, p. 435 
Taxonomy: Ferriere, 1962. Schweiz. Ent. Ges. Mitt. 35: 139. 

Genus ABLERUS Howard, pp. 435-436 

clisiocanipae (Ashmead). 

Biology: Hughes, 1960. Fla. Ent. 43: 90. 

Genus AZOTUS Howard, p. 436 

Taxonomy: Alam, 1956. Roy. Ent. Soc. London Trans. 108: 358, 376. 

Genus TUMIDISCAPUS Giroult, p. 436 
cercopiphagns Milliron. Fla., Mich. 



EULOPHIDAE 223 

Genus PHYSCUS Howard, pp. 436-437 

*Encyrtophyscus Blanchard in DeSantis, 1948. La Plata Mus. Rev., n. s., 
Zool. 5: 192. 

Type: Physcus flaviflagellatus DeSantis. Desig. by DeSantis, 
1948. 
Taxonomy: Compere and Annecke, 1961. Ent. Soc. S. Afr. Jour. 24: 20. 

*flaviventris Howard. Tex.; Philippines. Host: Assoc, with Antonina gra- 
minis (Mask.). 
Physcus flaviventris Howard, 1910. Ent. News 21: 162. $. 

*fulvus Compere and Annecke. Calif.; Formosa. Host: Lepidosaphes beckii 
(Newon.). 
Physcus "B", Flanders, 1953. Lab. Agr. Gen. e Zool. Portici Bol. 33: 22. 
Physcus fulvus Compere and Annecke, 1961. Ent. Soc. S. Afr. Jour. 24: 

21. 9, $. 
Biology: Clausen, 1956, loc. cit. supra, p. 122 (sp. B). 

varicornis (Howard). Wis., Sask. Hosts: Phenacaspis pinifoliae (Fitch), 
Nuculaspis calif ormca (Colem.). 

Genus PROSPALTELLA Ashmead, p. 437-438 

Taxonomy: Alam, 1956. Roy. Ent, Soc. London Trans. 108: 359, 367. — 
Ferriere, 1961. Schweiz. Ent. Ges. Mitt. 34: 253. 

anrantii (Howard). 

Biology: Benassy, 1958. Entomophaga 3: 104. — Muma, 1959. Jour. 

Econ. Ent.'52: 579. 
Taxonomy: Ferriere, 1961, loc. cit. supra, pp. 256, 263. 

berlesei (Howard). N. C, Fla.; Europe, Japan. 

Biology: Gireau, 1956. Phytoma 82: 29. — Benassy, 1957. Entomophaga 

2: 283. — Benassy, 1958. Ann. Epiphyt., ser. C, 9: 459. — Benassy, 

1961. Ann. Epiphyt., ser. C, 12: 127. 
Taxonomy: Flanders, 1960. Jour. Econ. Ent. 53: 757.— Ferriere, 1961, 

loc. cit. supra, p. 256. 

diaspidicola Silvestri. Mass., D. C, HI.; West Indies, S. America, Europe, 
Africa. 
Taxonomy: Flanders, 1960, loc. cit. supra, p. 757 (? syn. of berlesei 
(Howard) ). 

fasciata Malenotti. D. C, Fla., Tex.; Europe. Host: Carulaspis visci (Schr.). 
Biology: Benassy and Bianchi, 1960. Entomophaga 5: 168. 
Taxonomy: Ferriere, 1961, loc. cit. supra, pp. 255, 259. 

fuscipennis Girault. Transf. to Coccophagoides. 

murtfeldtae (Howard). Ark. 

peltata (Cockerell), corrected spelling of peltatus of Catalog, teste Peck, 
1963, loc. cit. supra, p. 280. 

perniciosi Tower. 

Biology : Clausen, 1956, loc. cit. supra, p. 124. — Flanders, 1956. Lab. 
Zool. Agr. Portici Bol. 33: 23.— Chumakova, 1957. Zool. Zhur. 36: 
534. — Benassy and Bianchi, 1960. Entomophaga 5: 168. — 
Flanders, 1960. Jour. Econ. Ent. 53: 757. — Chumakova and Gor- 
junova, 1963. Ent. Obozr. 42 : 320. 

Genus ENCARSIA Foerster, p. 438 

*basicincta Gahan. Fla.; West Indies. Host: Aleurothrixus floccosus 
(Mask.). 
Encarsia basicincta Gahan, 1927. U. S. Natl. Mus. Proc. 71 (4) : 20. 9. 

coquilletti Howard. Idaho, Ariz., Wash. 

formosa Gahan. Calif. 

Biology: Burnett, 1958. Canad. Ent. 90: 179-191, 225-229.— Burnett, 



224 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

1962. Canad. Ent. 94: 673.— Timofeeva, 1963. Zashch. East. 
Vred. i Boleznei. 8: 44. 

luteola Howard. Calif. 

Biology: Poinar, 1964. Jour. Ent. 57: 382. 

meritoria Gahan. Ariz., Calif. 

pergandiella Howard. Host: Trialeurodes abutilonea (Hald.). 

quaintancei Howard. 111. Host: Trialeurodes abutilonea (Hald.). 

variegata Howard. Host: Aleurothrixus floccosus (Mask.). 

Genus ASPIDIOTIPHAGUS Howard, pp. 438-439 

Taxonomy: Alam, 1956. Roy. Ent. Soc. London Trans. 108: 359. 

citrinus citrinus (Craw). 

Biology: Benassy, 1958. Entomophaga 3: 104.— Hughes, 1960. Fla. Ent. 
43: 90.— Benassy, 1961. Ann. Epiphyt., ser. C, 12: 42. 

lounsburyi (Berlese and Paoli). 

Taxonomy: Muma, 1959. Jour. Econ. Ent. 52: 579. — Annecke, 1963. 
Ent. Soc. S. Afr. Jour. 26: 206. 

*Genus COCCOPHAGOIDES Girault 

Coccophagoides Girault, 1915. Queensland Mus. Mem. 4: 58. 

Type: Coccophagus abiiorniicornis Girault. Orig. desig. 
*Diaspiniphagus Silvestri, 1927. Lab. Zool. Gen. e Agr. Portici Bol. 20: 
35. 

Type: Prospalta similis Masi. Orig. desig. 
Taxonomy: Compere, 1936. Univ. Calif. Pubs., Ent. 6: 292. 

Place in Aphelininae following Aneristus Howard, p. 439. 

fuscipennis (Girault). D. C. Transf. from Prospaltella Howard, teste H. 
Compere, in litt. 

*kuwanai (Silvestri). Conn., N. Y., N. J., Ohio, Calif.; Japan. Host: Aspi- 

diotus perniciosus (Comst.). 
Diaspiniphagus kutvanae Silvestri, 1927. Lab. Zool. Portici Bol. 20: 39. 

9. 
Taxonomy: Compere, 1936, loc. cit. supra, p. 293. 
Biology: Flanders, 1960. Jour. Econ. Ent. 53: 757. 
Nomenclature: Peck, 1963, loc. cit. siipra, p. 294. 

'^similis (Masi). N. Amer.; Europe. Host: Aspidiotus perniciosus (Comst.). 
Prospalta similis Masi, 1908. Lab. Zool. Portici Bol. 3: 148. 9. 
* Prospaltella ilicis Mercet, 1921. R. Soc. Esp. Hist. Nat. 50: 304. 5. (Syn. 

by Silvestri, 1927, loc. cit. supra, p. 36.) 
Distribution : Flanders, 1960, loc. cit. supra, p. 757. 

Genus COCCOPHAGUS Wesfwood, pp. 439-441 

Taxonomy: Alam, 1956, loc. cit. supra, pp. 358, 363. — Tachikawa, 1957. 
Japan. Jour. Appl. Ent. and Zool. 1: 61. — Zinna, 1961. Lab. Ent. 
Agr. Portici Bol. 19: 303, 

albicoxa Howard. Hosts: Toumeyella pinicola Ferris, T. parvicornis (Ckll.). 

*basalis Compere. Calif. ; Brazil. 

Coccophagus basalis Compere, 1939. Calif. Univ. Pubs., Ent. 7: 86. ?. 
Biology: Flanders, Bartlett, Fisher, 1961. Ent. Soc. Amer. Ann. 54: 
227-236. 

capensis Compere. Calif.; S. Africa, Australia. 
Biology: Clausen, 1956, loc. cit. supra, p. 93. 

cowperi Girault. Tex., Calif.; Africa, Israel. Hosts: Saissetia oleae (Bern.), 
S. nigra (Niet.), Coccus hesperidum (L.). Incorrectly listed in 
Catalog, p. 441, as a syn. of lycimnia (Walker) . 



EULOPHIDAE 225 

Taxonomy: Compere, 1931. U. S. Natl. Mus. Proc. 78 (7) : 57, 60.— 
Flanders, 1953. Ent. Soc. Amer. Ann. 46: 90-91.— Alam, 1956, loc. 
cit. supra, pp. 363-364. 

Biology: Clausen, 1956, loc. cit. supra, p. 94. — Flanders, 1959. Jour. 
Econ. Ent. 52: 597. 

flavifrons Howard. Pa., Md., S. C, Ohio, Tex. Hosts: Phenacaspis pinifoliae 
(Fitch), ICoccus hesperidum (L.). 

gumeyi Compere. Calif. ; Australia, Africa. 

Biolog^y: Clausen, 1956, loc. cit. supra, p. 94. — Bartlett and Lloyd, 1958. 
Jour. Econ. Ent. 51: 92. 

lycimnia (Walker). Delete coipperi Girault from syn. 

Biology: Clausen, 1956, loc. cit. supra, pp. 94-95. — Peterson, 1962. Can- 
ad. Ent. 94: 740.— Sonina, 1962. Akad. Nauk. Uzbek. SSR, Biol. 
Zhur. 3: 62.— Sugonjaev, 1962. Akad. Nauk. SSSR Izv., Ser. Biol 
5: 755.— Phillips, 1963. Canad. Ent. 95: 402. 

*pulvinariae Compere. Calif.; Africa. Hosts: Coccus hesperidum (L.), Sais 

setia nigra (Niet.), S. oleae (Bern.). 
Coccophagus pulvhiariae Compere, Mar. 1931. U. S. Natl. Mus. Proc. 78 

(7): 53. 9, $. 
*Coccophagus eritroensis Compere, Apr. 1931. Univ. Calif. Pubs., Ent. 5: 

254. 9, $. (Syn. by Flanders, 1959. Jour. Econ. Ent. 52: 597.) 
Biology: Clausen, 1956, loc. cit. supra, pp. 94-95. — Flanders, 1959, loc. 

cit. supra, p. 597. 

rusti Compere. Calif.; Africa. 

Biology : Clausen, 1956, loc. cit. supra, p. 95. 

scutellaris (Dalman). Fla., Oreg. Host: Saissetia nigra (Niet.). 

Biology: Flanders, 1959. Jour. Econ. Ent. 52: 74.— Sonina, 1962. Akad. 
Nauk. Uzbek. SSR., Biol. Zhur. 3: 62. 

trifasciatus Compere. Transf . to Lounsburyia Compere and Annecke, infra. 

yoshidai Nakayama, corrected spelling of yoshidae Nakayama of Sup. 1, p. 
66, teste Peck, 1963, loc. cit. supra, p. 318. 
Biology: Sugonyaev and Peng, 1960. Ent. Obozr. 39: 702, 706. 

*Genus LOUNSBURYIA Compere and Annecke 

Lounsburyia Compere and Annecke, 1961. Ent. Soc. S. Afr. Jour. 24: 
28-29. 

Type: Coccophagus trifasciatus Compere. Orig. desig. 
Insert in Aphelininae after Coccophagus Westwood, p. 441. 

trifasciata (Compere). Transf. from Coccophagus Westwood, teste Compere 
and Annecke, 1961, loc. cit. supra, p. 29. 
Biology: Clausen, 1956, loc. cit. supra, p. 95. 

Subfamily TETRASTICHINAE 

Taxonomy: Ferriere, 1960. Acta Hym. (Japan) 1: 105-106 (key to gen- 
era) . 

Genus Thripocfenus Crawford, p. 442 
petiolata Ashmead. Ariz. 

Genus Thripocfenus Crawford, p. 442 

Transf. to syn. under Ceranisus Walker in the Entedontinae, p. 460; 
included spp. to be transf. to Ceranisus. 

Genus TETRASTICHUS Holiday 
(= Tefrosfichus Walker of Catalog, pp. 443-450) 

Paromphaloidomyiia Girault, 1917; Catalog, p. 452. N. syn.; tjT)es exa- 
mined. 



226 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

'!*Lygellus Giard, 1896. Soc. Biol. Paris C. R. 1896: 839. 

Type: Lygellus epilachnae Giard. By monotypy. 

*Myiomisa Rondani, 1877. Soc. Ent. Ital. Bui. 9: 189 (syn. by Boucek, 
1961. Min. Sel. Kruzy. Moldav. S.S.R. Trudy 7: 28). 

Type: Myiomisa microscopica Rondani. By monotypy. 

Taxonomy: Graham, 1961. Ent. Mo. Mag. 97: 34-64 (Aprostocetus). — 
Burks, 1962. Bui. Zool. Nomencl. 19: 306 (request for suppression 
of Tetrastichus Walker, 1842, and validation of Tetrastichus Hali- 
day, 1844). — Hedqvist, 1963, loc. cit. supra, pp. 146-147 
(Aprostocetus). — Int. Comn. Zool. Nomencl., 1965. Bui. Zool. No- 
mencl. 22: 26 (Opinion 720, suppression of Tetrastichus Walker, 
1842). 

argyrus Burks. Syn. of homeri (Girault). 

asparagi Crawford. Syn. of coeruleus (Nees). 

*auplopus Burks. Va. Host: Reared from cell of Auplopus sp. 

Tetrastichus auplopus Burks, 1963. Biol. Soc. Wash. Proc. 76: 55. ?. 

^barbarae Burks. Oreg. Host: Barbara colfaxiana (Kearf.), prob. secon- 
dary. 
Tetrastichus harharae Burks, 1963. Biol. Soc. Wash. Proc. 76: 52. 9, $. 

bruchivorus Gahan. Introd. from France and recovered in N. C., but prob. 
not est. in U.S. 

bruchophagi Gahan. 

Taxonomy: Butler and Hansen, 1958. Pan-Pacific Ent. 34: 224, 226. 

♦byersi Burks. Mich., Ind. Hosts: Pupae of Dolichopeza americana Need- 
ham, D. walleyi (Alex.). 
Tetrastichus byersi Burks, 1963. Biol. Soc. Wash. Proc. 76: 53. 9, $. 

carinatus Forbes. Syn. of charoba (Walker). 

*charoba (Walker). Generally dist. in U. S. and Canada; Europe, Asia. 

Cirrospilus charoba Walker, 1840. Ann. Nat. Hist. 4: 30. $. 

Tetrastichus carinatus Forbes, 1885 (syn. by Graham, loc. cit. supra, p. 
51 ^Aprostocetus^) . 

[Note: C. charoba Walk, selected by Graham from 13 other apparently, 
but not certainly, synonymous names, although it does not have priority. 
Article 23 (b) of Int. Code of Zool. Nomencl. would not preserve carinatus 
Forbes for this sp. if this selection is accepted.] 

chlamytis Ashmead. Mo., Ark., Kans., Tex., Utah. 

*clito (Walker). Ont.; Europe. Host: Cassida riibiginosa MnW. 
Cirrospilus clito Walker, 1840. Ann. Nat. Hist. 4: 30. $, 5. 
Taxonomy: Peck, 1963, loc. cit. supra, p. 128. 

coerulescens Ashmead. Mich., Wis., Minn., N. Mex., Ariz., Calif. Hosts; Cho- 
ristoneura fumiferana Clem., C. pinus Free., Dichomeris margin- 
ella (Fab.) ; prob. secondaries. 
Biology: Wester, 1956. Ent. Soc. Wash. Proc. 58: 284.— Schaffner, 1959, 
loc. cit. supra, p. 82. 

*coeruIeus (Nees). Wash.; Europe. 

Eulophus coeruleus Nees, 1834. Hym. Ichn. Affin. Monog., v. 2, p. 174. 

9. 
Tetrastichus asparagi Crawford, 1909 (syn. by Graham, 1961, loc. cit. 

supra, p. 39 [Aprostocetus]) . 
[Strict application of Article 23 (b) of Int. Code of Zool. Nomencl. 

would require retention of the name asparagi Crawford for this 

sp.] 

dolosus Gahan. Ariz. 

faustus Burks. Minn. 

Biology: Balduf, 1959. 111. Biol. Monog. 26, p. 146. 



EULOPHIDAE 227 

*garryana Burks. B. C, Oreg. Hosts: Besbicus mirahilis (Kinsey), several 
cynipid galls on Quercus garryana. 
Tetrastichus garryana Burks, 1963. Biol. Soc. Wash. Proc. 76: 49. 9, $. 
Biology: Evans, 1963. Canad. Ent. 95: 1002-1005. 

gibbon! (Girault). Okla. 

hagenowii (Ratzeburg). 

Biologv: Cameron, 1955. Bui. Ent. Res. 46: 137-147.— Cameron, 1957. 
Bui. Ent. Res. 48: 205-206.— Roth and Willis, 1960. Smithsn. 
Misc. Coll. 141: 249-253. 

homeri (Girault), n. comb. Va., S. C, Ga., Fla., Iowa. Host: Asteromyia sp. 
leaf gall on Solidago. Transf. from Paromphaloidomyiia Girault; 
types examined. 
Tetrastichus argyrus Burks, 1943, n. syn. ; types examined. 

*incertus (Ratzeburg). Mass., Pa., N. J., Md.; Europe. Host: Hypera pos- 

tica (Gyll.). 
Eulophus incertus Ratzebui'g. 1844, Ichn. der Forstins., v. 1, p. 168. 
Biologv: Chamberlin, 1925. Jour. Econ. Ent. 18: 597. — Coles and 

Puttier, 1963. Jour. Econ. Ent. 56: 610-611. 

johnsoni Ashmead. N. H., Md., Mo. Host: Prepupae of Dipogon sayi sayi 
Banks. 

longicorpus (Girault). Calif. 

malacosoinae Girault. 

Biology: Langston, 1957. Calif. Univ. Pubs. Ent. 14: 26-27. 

marcovitchi (Crawford). Ind., 111., Wis., N. Dak. 

Biology: Giese and Benjamin, 1959. For. Sci. 5: 205. 

marylandensis ( Girault) . 

Biology: Schalfner, 1959, loc. cit. supra, pp. 82-83. 

*megachilidi9 Burks. Fla., Ohio, Ariz. Hosts: Prepupae of Megachile 
(Litomegachile) gentilis Cress., M. (Melanosaurus) xylocopoides 
Smith, 

Tetrastichus megachilidis Burks, 1963. Biol. Soc. Wash. Proc. 76: 57. 9, 
$. 

Biology: Butler and Wargo, 1963. Pan-Pacific Ent. 39: 205. 

melanis Burks. N. H., Fla., Minn. [Prob. a syn. of the European coccinellae 
Kurdjumov.] 
Biology: Newcomer, 1958. Pan-Pacific Ent. 34: 89. 

*inilleri Burks. Calif. Host: Sec. -parasite ot Recurvaria milleri (Busck). 
Tetrastichus milleri Burks, 1963. Biol. Soc. Wash. Proc. 76: 56. $. 

modestus Howard. N. H. Host: Apanteles pyraustae Vier. 
Biology: Schaffner, 1959, loc. cit. supra, p. 83. 

*nordi Burks. Mich. Host: Agrilus sp. 

Tetrastichus nordi Burks, 1963. Biol. Soc. Wash. Proc. 76: 51. 5, $. 

oecanlhivorus compar Gahan. Md., Ark., Okla. 

paracholus Burks. Ont., N. C, Wis., Minn. 

*pausiris (Walker). Ont.; Europe. Host: Dasyneura leguminicola (Lint.). 
Cirrospilus pausiris Walker, 1839. Monog. Chalc, p. 327. 
Taxonomy : Peck, 1963, loc. cit. supra, p. 143. 

phegus Burks. Syn. of racemariae Ashmead. 

*phidippi Burks. Ariz. Host: Eggs of Phidippus opifex (McCook). 

Tetrastichus phidippi Burks, 1963. Biol. Soc. Wash. Proc. 76: 60. $, $. 

*poliii Burks. Va. Host: GaWs oi Xanthoteras politum (Bass.). 

Tetrastichus politi Burks, 1963. Biol. Soc. Wash. Proc. 76: 48. ^, $. 



228 HYMENOPTERA OF AMERICA NORTH OP MEXICO 

*pompilicola Graham. Newfoundland. Host: Anoplius nigerrimus (Scop.). 
Tetrastichus pompilicola Graham in Lindroth and Graham, 1960. Opusc. 
Ent. 25: 94. 9. 

productus Riley. Que., Minn. Host: Janus abbreviatus (Say); delete Phyto- 
phag a destructor (Say). 

*psyllaephagus Burks. Ariz. Host: Nymph of Trioza collaris Crawford. 

Tetrastichus psyllaephagus Burks, 1963. Biol. Soc. Wash. Proc. 76: 54. 

9 $ . 
Biology:' Cazier, 1964. Pan-Pacific Ent. 40: 71. 

racemariae Ashmead. Ont. south to Fla., west to Calif. 

Tetrastichus phegus Burks (syn. by Judd, 1963. Canad. Field-Nat. 77: 
63). 

rapo of Catalog. Syn. of sinope Walker. 

silvaticus Gahan. N. B. south to Conn., west to Wash, and Calif. 

*sinope (Walker). Cosmopolitan. Host: Apanteles glomeratus (L.). 
Cirrospilus sinope Walker, 1839. Monog. Chalc, v. 1, p. 295. 9. 
Tetrastichus rapo of Catalog and authors, long, misdet. teste Graham, 
1961, loc. cit. supra, p. 38. 

strobilus Burks. Host: (?) Barbara coif axiana (Kearf.). 
Biology : Keen, 1958, loc. cit. supra, pp. 95-96. 

triozae Burks. Hosts: Arytaina minuta Cwfd., Calophya calif ornica Schw., 
C. nigrella Jens., C. triozomima Schw., Euphalerus vermiculosus 
Cwfd., Pexopsylla cercocarpi Jens., Trioza albifrons Cwfd., T. 
b cayyieri. 
Biology: Jensen, 1957. Higardia 27: 79-80, 86-96. 

Irisulcatus Provancher. 

Taxonomy: Finlayson, 1963. Canad. Ent. 95: 498 (imm. stages). — 
Burks, 1964, loc. cit. supra, p. 1262 (types). 

lurionum (Hartig). Ont. 

Biology: Clausen, 1956, loc. cit. supra, pp. 130-131. — Juillet, 1959. Can- 
ad. Ent. 91: 716-718.— Schaffner, 1959, loc. cit. supra, p. 83.— Ad- 
lung, 1962. Ztschr. Angew. Ent. 50: 456. 

varicomis (Girault). Maine, Mass. Host: Epinotia nanana (Treit.). 
Biology : Schaffner, 1959, loc. cit. supra, p. 83. 

venustus Gahan. Que., Maine, Minn., Nebr. 

Taxonomy: Butler and Hansen. 1958. Pan-Pacific Ent. 34: 224-226. 

whitman! (Girault). 

Biology: Giese and Benjamin, 1959. For. Sci. 5: 205. 

xanthomelaenae (Rondani). 

Taxonomy; Boucek, 1957. Acta Ent. Mus. Nat. Pragae 31: 117 (galleru- 
cae Fonsc). — Peck, 1963, loc. cit. supra, pp. 132-133 (gallerucae 
Fonsc). 

xanthops (Ratzeburg). B. C. Host: Coleophora laricella Hbn. 
Biology: Schaffner, 1959, loc. cit. supra, p. 83. 

Genus APROSTOCETUS Westwood, pp. 450-451 

canadensis Ashmead. Syn. of daira (Walker). 

*daira (Walker). Que., Ont., Ohio, Nebr.; Europe. 

Cirrospilus daira Walker, 1839. Monog. Chalc, v. 1, p. 330. 9. 
Aprosiocetus canadensis Ashmead (syn. by Graham, 1961. Ent. Mo. Mag. 

97:47). 

Genus SYNTOMOSPHYRUM Foerster, pp. 451-452 

Tetrastichomyia Girault, 1916, n. syn. Transf. from syn. under Miotropis 

Thomson, p. 454 ; types examined. 
Taxonomy: Graham, 1961. Ent. Mo. Mag. 97: 36 (Aprostocetus). 



EULOPHIDAE 229 

blattae Burks. 

Biology: Roth and Willis, 1960, loc. cit. supra, p. 248. 

clisiocampae (Ashmead). Transf. from Miotropis Thomson; types examined. 

esurus (Riley). 

Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, pp. 91-92. 

ischnopterae (Girault). 

Biology: Roth and Willis, 1960, loc. cit. supra, p. 249. 

orgyiae (Girault), n. comb. Transf. from Miotropis Thomson; types exa- 
mined. 

philodromi (Gahan). Nebr. 

silvensis (Girault), n. comb. Transf. from Miotropis Thomson; types exa- 
mined. 

Genus Paromphaloidomyiia Girault, p. 452 
Syn. of Tetrastichus Haliday, supra. 
homeri Girault. Transf. to Tetrastichus Haliday. 

Genus MELITTOBIA Westwood, p. 452 

Delete Aceratoneuromyia Girault from syn.; a valid, but extralimital 
genus, teste Gradwell, 1959. Ent. Mo. Mag. 94: 277. 

chalybii Ashmead. Hosts: Attack almost any insect in lab. cultures or insec- 
taries, but in nature apparently attack only aculeate Hymenop- 
tera or their parasites. 
Biology: Medler, 1958. Ent. News 69: 21-23.— Medler, 1958. Ent. Soc. 
Amer. Ann. 51: 340-344.— Krombein, 1964. Biol. Soc. Wash. Proc 
77: 101. 

Subfamily ELACHERTINAE 

Genus EUPLECTRUS Westwood, pp. 452-453 

mellipes Provancher. Maine, N. H., Wis., Alaska. 
Biology: Guppy, 1959. Canad. Ent. 91: 426. 
Taxonomy: Burks, 1964, loc. cit. supra, p. 1258. 

Genus DIAULINOPSIS Crawford, pp. 453-454 

callichroma Crawford. Md. south to Fla., west to Nebr. and Calif.; Mexico, 
West Indies. 

Genus MIOTROPIS Thomson, p. 454 

Sienomesioideus Ashmead, n. syn. Transf. from Catalog, p. 460; types 

examined. 
Miotropis Thomson has long been misidentified in the N. Amer. lit. 

Delete Teirastichomyia Girault from syn. and transf. to syn. of 

Syntomosphyrum Foerster, p. 451. 
clisiocampae Ashmead. Transf. to Syntomosphyrum Foerster. 

mellea (Ashmead), n. comb. Transf. from Stenomesius Ashmead; types exa- 
mined. 

orgyiae (Girault). Transf. to Syntomosphyrum Foerster. 

silvensis (Girault). Transf . to Syntomosphyrum Foerster. 

Genus ELACHERTUS Spinola, pp. 455-456 

aeneoniger Girault. Wis., Minn., Oreg., B. C. Host: Choristoneura fumifer- 
ana (Clem.). 

glacialis Ashmead. Colo., Wash., Oreg. Hosts: Barbara colfaxiana (Kearf.). 
Mellisopus latiferreanus (Wlshm.). 
Biology : Keen, 1958, loc. cit. supra, p. 94. 



230 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

cacoeciae Howard. Oreg. Host: Choristoneura fumiferana (Clem.). 
Biology: Carolin and Coulter, 1959. Jour. Econ. Ent. 52: 552. 

coxalis Howard. Va. Host: Callisto ge7ninatella Pack. 

Biology: Hill and Hough, 1957. Va. Agr. Expt. Sta. Tech. Bui. 130, p. 
16. 

hyphantriae Crawford. Mass., Ark., Tex. 

proteoteratis Howard. Ont., Mich., Idaho, Ariz., Calif. 

Euplectrus viridaeueus Provancher, 1887 (syn. by Burks, loc. cit. supra, 
p. 1258). 

viridaeneus (Provancher). Syn. of proteoteratis Howard. 

Genus HYSSOPUS Giraolt, p. 456 

benefactor (Crawford). Md., Ohio. Host: Petrova virginiana (Busck), P. 
albicapitana (Busck) , P. metalhca (Busck). 

evetriae (Girault). Ohio. Host: Petrova wenzeli (Kearf.). 

Biology: Miller and Altmann, 1958. Ohio Jour. Sci. 58: 278-280.— Keen, 
1958, loc. cit. supra, p. 95. 

johannseni (Crawford). Va., Miss. Hosts: Laspeyresia spp. in pine cones. 

rhyacioniae Gahan. Ont. south to Fla. west to Mich., Ark., Tex. Hosts: 
Dioryctria abietella D. and S., D. amatella (Hulst), D. cambiicola 
(Dyar), D. zimmermani (Grote). 

thymus Girault. Ont. and N. Y. south to Fla. west to Idaho and Calif. 
Hosts: Dioryctria auranticella (Grote), D. zimmermani (Grote), 
Petrova comstockiana (Fern.), P. wenzeli (Kearf.), Rhyacionia 
rigidnna ( Fern. ) . 
Biology: Miller and Altman, 1958. Ohio Jour. Sci. 58: 278-280.— Keen, 
1958, loc. cit. supra, p. 95. — Watson and Arthur, 1959. Canad. 
Ent. 91: 480-484. — Schaffner, 1959, loc. cit. supra, p. 84.— Hart- 
man and Kulman, 1962. Jour. Econ. Ent. 55: 1007. 

Genus APTEROLOPHUS Gahan, p. 457 
pulchricomis Gahan. S. C, Ind, Host: Taken sifting treehole litter. 

Genus CIRROSPILUS WesJwood, pp. 458-459 

*Plesiospnus Ferriere, 1953. Univ. Bologna 1st. Ent. Bol. 19: 398. 

Type: Eulophus unistriatus Foerster. Orig. desig. 
Taxonomy: Delucchi, 1958. Entomophaga 3: 253. 

cinctithorax (Girault). N. B., Que., Ont., Mass. Hosts: Arc/yresthia freyella 
Wlshm., Colcophora fiiscedinella Zeller, Lithccolletis salicif oliella 
Clem., L. tremuloiclella Braun, Ducculatrix ainsliella Murt. 
Biology: Schaffner, 1959, loc. cit. supra, p. 84. 

flavicinctus Riley. Host: Bucculatrix ainsliella Murt. 
Biology: Schaffner, 1959, loc. cit. supra, p. 84. 

flavoviridis Crawford. Ariz., Calif. Host: Liriomyza sp. 

pictus (Nees). 

Biology: Sundby, 1958 (1957). Norsk. Ent. Tidssk. Sup. 2, pp. 18-21. 

Genus ZAGRAMMOSOMA Ashmead, pp. 459-460 

Taxonomy: Boucek, 1959. Mus. Nat, Pragae Acta Ent. (Zagrammosoma 
a subg. of Cirrospilus Westwood). 

americanum Girault. Ariz., Calif. Host: Recurvaria milleri Busck. 

interlineatum Girault. Mass., Fla., Ohio. Hosts: Exoteleia pinif oliella 
(Chamb.), Lithocolletis cincinnatielln (Chamb.). 
Biology: Schaffner, 1959, loc. cit. supra, p. 84. 

mirum Girault. Hosts: Lithocolletis crataegella Clem., Liriomyza pictella 
(Thomson). 



EULOPHIDAE 231 



Genus Stenomesioideus Ashmead, p. 460 
Transf. to syn. of Miotropis Thomson, p. 454. 
melleus Ashmead. Transf. to Miotropis Thomson. 

*Genus MiRZAGRAMMOSOMA Girault 

Mirzagrammosoma Girault, 1915. Ent. Soc. Amer. Ann. 8: 279. 

Type: Mirzagrammosoma I'meaticeps Girault. By monotypy. 

Insert in Elachertinae after Zagrammosoma Ashmead, p. 460. 

*lineaticeps Girault. Fla. ; Cent. America, West Indies. Hosts: Leucoptera 
coffeella (Guerin), Liriomyza archboldi Frost. 
Mirzagrammosoma lineaticeps Girault, 1915. Ent. Soc. Amer. Ann. 8: 
279. 9. 

Subfamily ENTEDONTINAE 

♦Genus CERANISUS Walker 
l — Thripoctenus Crawford of Catalog, p. 442) 

Ceranisus Walker, 1842. Entomologist 1: expl. of PI. N, fig. 2. 

Type: Cirrospilus pacuvius Walker. By monotypy. 
Cryptomphale Girault, 1917. Spec. Gen. nova Euloph., p. 2. 

Type: T/iripocrenus nufci/ipcHJiis Williams. By monotypy. 
Taxonomy: Burks, 1943. U. S. Natl. Mus. Proc. 93: 509.— Bakkendorf, 
1957. Ent. Medd. 28: 1.— Graham, 1959. Soc. Brit. Ent. Trans. 
13: 203. — Peck, 1963, loc. cit. supra, p. 117 (n. comb's.). 

Insert in Entedontinae before Astichus Foerster, p. 460; transf. spp. in 
Catalog under Thripoctenus to Ceranisus. 

Genus OMPHALE Holiday, p. 460 

*Secodes Foerster, 1856. Hym. Stud., v. 2, pp. 78, 81. 

Type : Secodes fagi Foerster. By monotypy. 
Taxonomy: Graham, 1963. Soc. Brit. Ent. Trans. 15: 240. 

Genus CLOSTEROCERUS Westwood, p. 461 

cinctipennis Ashmead. Que. south to Fla. west to Wis., Iowa, Mo. Hosts: 
Brachys ovatus Web. (eggs and larvae), Neodiprion lecontei 
(Fitch), N. excitans Roh., N. virginianus complex (eggs). 
Biology: Benjamin, 1955. U. S. Dept. Agr. Tech. Bui. 1118, pp. 24-26.— 
Morris, Schroeder, and Bobb, 1963. Pine Sawfly in Va., Va. Div. 
For., p. 13. 

tricinctus (Ashmead). Host: Exoteleia pinifoUella (Chamb.). 
Biology: Schaffner, 1959, loc. cit. supra, p. 84. 

irifasciatus Westwood. Mass., Ariz. Host: Liriomyza pictella (Thomson), 
Biology: Sundby, 1957. Norsk Ent. Tidsskr., sup. 2, p. 41. 

Genus EUDERUS Haliday, p. 462 

*Secodelloidea Girault, 1917. Sp. Gen. nova Euloph., p. 2. 

Type: Secodella rugosus Crawford. By monotypy. 

Taxonomy: Erdos, 1956. Rov. Kozlem., n. s., 9: 2 (Euderini). — Graham, 
1959. Soc. Brit. Ent. Trans. 13: 172, 203 (Euderinae).— Peck, 
1963, loc. cit. supra, p. 172 {Secodelloidea Girault a syn.). — 
Boucek, 1963. Beitr. z. Ent. 13: 267 (Secodelloidea Girault dis- 
tinct, poss. a subgen.). 

argyresthiae (Crawford). N. Y. south to Fla., west to Wash., Oreg., Tex. 
Hosts: Recurvaria spp. 

crawfordi Peck. Ariz., Calif. Hosts: Galls of Dryocosmus coxii (Bass.). PZo- 
giotrochus suberi Weld. 



232 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

cushmani (Crawford). New England, N. Y., 111. Hosts: Coleophora laricella 
Hbn., Cydia packardii (Zell.), Dioryctria ziinmerinani (Grote), 
Proteoteras willingana (Kearf.). 
Biology: Balduf, 1959. 111. Biol. Monog. 26: 74.— Schaffner, 1959, loc. cit. 
supra, p. 85. 

rubriscapus (Girault), new status. Change from subsp. to sp. 

rugosus (Crawford), new status. N. Y. south to Fla. west to Tenn. and 
Kans. Change from subsp. to sp. 

rugosus rubriscapus (Girault). Change to sp. 

rugosus rugosus (Crawford). Change to sp. 

*Genus FLOCKIELLA Timberlake 

Flockiella Timberlake, 1957. Pan-Pacific Ent. 33: 109. 

Type: Flockiella eucalypti Timberlake. Orig. desig. 

Insert in Entedontinae following Euderus Haliday, p. 463. 

*eucalypti Timberlake. Calif. Host: Reared from seed capsules of Euca- 
lyptus uvibellata. 
Flockiella eucalypti Timberlake, 1951. Pan-Pacific Ent. 33: 110. 9. 
Biology: Flock, 1957. Pan-Pacific Ent. 33: 153-155. 

Genus BURKSIA Fullaway, Sup. 1, p. 68 
viridimaculata Fullaway. Calif. ; Cuba, Argentina. 

Genus CHRYSOCHARIS Foersfer, pp. 463-464 
ainsliei Crawford. S. C. Host: Liriomyza pictella (Thomson). 
compressicornis Ashmead. Syn. of viridis Provancher. 
*cubensis Gahan. Fla.; West Indies. 

Chrysocharis cubensis Gahan, 1932. Ent. Soc. Amer. Ann. 25: 748. 9, $. 
gemma (Walker). Transf. to Kratochviliana Mala5, infra. 

imbrasus (Walker). Transf. from Unplaced Species, p. 594, teste Graham, 
1963. Soc. Brit. Ent. Trans. 15: 234. 

laricinellae (Ratzeburg). Transf. to Kratochviliana Malac, infra. 

""majoriani (Girault). Fla.; West Indies, Brazil, introd. into Hawaii for biol. 
control. Hosts: Liriomyza sorosis (Will.), Phytobia sp., leaf- 
miners on Cassia and other plants. 
Chrysocharodes majoriani Girault, 1917. Descr. Hym. Chalc. var. cum 
Obs., p. 10. 9. 

viridis Provancher. Ont., Ohio. Host: Reared from gall on Lactuca. 

Chrysocharis compressicornis Ashmead, 1894 (syn. by Burks, 1964, loc. 
cit. supra, p. 1257). 

Genus KRATOCHVILIANA Malac 

Kratochviliana Malac, 1943. Ent. listy (Folia Ent.) Brno 6: 1. 

Type: Kratochviliana moravica Malac. Orig. desig. 

Epilampsis Delucchi, 1954. Neue Arten Subfam. Entedon., p. 3. 

Type : Chrysocharis boops Thomson. Orig. desig. 
Taxonomy: Boucek, 1961. Trudy Min. Sel. Kruzy. Moldav. S. S. R. 7: 28 
(syn.).— Graham, 1963. Soc. Brit. Ent. Trans. 15: 205 (both 
above gen. syn. of Chrysocharis Foerster) . 

Insert in Entedontinae following Chrysocharis Foerster, p. 464. 

gemma (Walker). B. C; Europe. Transf . from Chrysocharis Foerster. 

Taxonomy: Delucchi, 1954. Schweiz. Ent. Ges. Mitt. 27: 300-301 {Epi- 
lampsis Delucchi). 

laricinellae (Ratzeburg). N. B., N. S., Ont., Wis. Transf. from Chrysocharis 
Foerster. 



EULOPHIDAE 233 

Taxonomy: Bou2ek, 1961, loc. cit. supra, p. 28. — Peck, 1963, loc. cit. 

supra, pp. 205-207. 
Biology: Clausen, 1956, loc. cit. supra, p. 92.— Graham, 1958. Ent. Soc. 

Ont. Ann. Rpt. 1957, pp. 37-43.— Schaffner, 1959, loc. cit. supra, p. 

85. 

*Genus ENAYSMA Delucchi 
Insert in Entedontinae following Chrysocharis Foerster, p. 464. 

♦ENAYSMA subgenus ENAYSMA Delucchi 

Enaysma Delucchi, 1954. Neue Arten Subfam. Entedon., p. 1. 
Type: Enaysma zwiilferi Delucchi. Orig. desig. 

The typical subgenus does not occur in N. Amer. 

♦ENAYSMA subgenus PENTENAYSMA Graham 

Enaysma subg. Pentenaysma Graham, 1959. Soc. Brit. Ent. Trans. 13: 
196. 

Type: Eulophus latreillei Curtis. Orig. desig. 

Taxonomy: Miller, 1962. Canad. Ent. 94: 1039-1040. 

*arieiiascapus Miller. Que., Ont. Hosts: Lithocolletis ostryaefoliella Clem., 
Lithocolletis sp. on Viburmim. 
Enaysma arienascapus Miller, 1962. Canad. Ent. 94: 1050. 9, $. 

*carioca Miller. Que., Ont. Hosts: Lithocolletis sp. on Carya, leaf miner on 
oak. 
Enaysma carioca Miller, 1962. Canad. Ent. 94: 1043. 9, $. 

*clypeata Miller. Que. Host: Lithocolletis sp. on Acer. 

Enaysma clypeata Miller, 1962. Canad. Ent. 96: 1047, 1050. $. 

*gahani Miller. Que., Ont. Hosts: Lithocolletis lucetiella Clem., Lithocolletis 
spp. 
Enaysma gahani Miller, 1962. Canad. Ent. 94: 1041. ?, $. 

*hirtiscapus Miller. Ont. Host: Lithocolletis tilincella Chamb. 

Enaysma hirtiscapus Miller, 1962. Canad. Ent. 94: 1047. ?, $. 

*inyricae Miller. N. S. Host: Lithocolletis sp. on Myrica gale. 

Enaysma myricae Miller, 1962. Canad. Ent. 94: 1043-1047. 9, $. 

♦Genus MOSERINA Delucchi 

Moserina Delucchi, 1962. Kans. Ent. Soc. Jour. 35: 389. _ 
Type: Moserina niaculata Delucchi. Orig. desig. 

Insert in Entedontinae following Achrysocharoides Girault, p. 464. 

*maculata Delucchi. Ohio. Host: Reared from galls of Pachypsylla celtidis- 
mavwia (Riley), and P. celtidisvesicula Riley. 
Moserina maculaia Delucchi, 1962. Kans. Ent. Soc. Jour. 35: 392. 9, $. 

Genus PARACHRYSOCHARIS Girault, p. 465 

semiflava Girault. Wis. Host: Auplopus sp. 

Genus DEROSTENUS Westwood, pp. 466-467 

agromyzae Crawford. N. Y. south to Fla., west to N. Dak. and Ariz. Hosts: 
Delete Agromyza scutellata Fall and A. pusilla (Meig.) ; add 
Manturafloridana (Crotch) , Liriomyza sorosis (Will.). 
Biology: Peck, 1963, loc. cit. supra, p. 214. 

arizouensis Crawford. Hosts: Agromyza scutellata Fall, A. pusilla (Meig.), 
Liriomyza pictella (Thomson). 

fullawayi Crawford. Calif.; Hawaii; Mexico. 

punctiventris Crawford. Idaho, Calif. 



234 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Silvia (Girault). Maine south to Md. west to Wis., Calif. Hosts: Coleophora 
laricella (Hbn.), Exoteleia pinifoliella (Chamb.), Tetralopha ro- 
btistella Zell., Recurvaria milleri Busck, R. apicitripunctella 
(Clem.). 
Biology: SchaflFner, 1959, loc. cit. supra, p. 85. 

variipes Crawford. N. Y. south to Fla. west to La. and Tex, 

Genus HORISMENUS Walker, pp. 467-468 

*Pedlohioidea Girault, 1911. Canad. Ent. 43: 407. 

Type: Pediobioidea cyanea Girault. By monotypy. 
Taxonomy: Peck, 1963, loc. cit. supra, pp. 217, 220. 

lixivorus Crawford. Tex., Ariz, (delete Maine). Hosts: Lixus perforatus 
Lee. (delete Phyllocnistis populiella Chamb.). 

microgaster (Ashmead). Va., N. C. Hosts: CaUisto geminatella (Pack.), Te- 
tralopha robusiella Zell., Pyrausta futilalis (Led.), Choristoneura 
rosaceana (Har.), Exoteleia pinifoliella (Cham.), Coleophora lari- 
cella (Hbn.) ; secondary par. 
Biology: Hill and Hough, 1957. Va. Agr. Expt. Sta. Tech. Bui. 130, p. 
16. — Schaffner, 1959, loc. cit. supra, p. 85. 

violaceus (Ashmead). Mass., Va. Hosts: Lithocolletis cincinnatiella 
(Chamb.), L. crataegella Clem. 
Biology: Schaffner, 1959, loc. cit. supra, p. 85. 

Genus PEDIOBIUS Walker, Sup. 1, p. 68 
( = P/euro/rop/s Foerster of Catalog, pp. 468—470) 

*Me8tocharomyia Girault, 1915. Queensland Mus. Mem. 3: 188. N. syn. 

Type: Mestocharis lividus Girault. Orig. desig. 
*Eupleurotropis Girault, 1917. Spec. Gen. nova Euloph., p. 3. 

Type : Pleurotropis testaceipes Crawford. By monotypy. 
Taxonomy: Peck, 1963, loc. cit. supra, p. 228. 

albipes (Provancher) . Maine and Ont., south to Va., west to 111. Hosts: 
Apanteles congregatus (Say), ? Gelis tenellus (Say), Dibrachys 
cavus (Walk.), Euptcromalus hemipterus (Walk.), agromyzid on 
Helianthus, 1 Lithocolletis sp. 
Entedon albitarsis Ashmead, 1888 (syn. by Burks, 1964, loc. cit. supra, p. 
1259). 

albitarsis (Ashmead). Syn. of albipes (Provancher). 

benefica (Gahan). Syn. of nigritarsis (Thomson). 

bucculatricis (Gahan). Host: Xenochalepus dorsalis (Thunb.) ; may be secon- 
dary. 
Biology: Schaffner, 1959, loc. cit. supra, p. 85. 

metallicus ( Nees ) . 

Biology: Clausen, 1956, loc. cit. supra, p. 123. 

niger (Ashmead), corrected spelling of nigra of Catalog. 

*nigritarsis (Thomson). Ont. south to Va. west to Alta. and S. Dak.; Europe. 
Pleurotropis nigritarsis Thomson, 1878. Hym. Scand., v. 5, p. 251. 9, $. 
Pleurotropis benefica Gahan (syn. by V. Rosen, 1956. K. Lantbrukshog. 

Ann. 23: 19). 
Taxonomy: v. Rosen, 1959. Ent. Tidskr. 80: 163.— Peck, 1963, loc. cit. 

supra, p. 229. 

sexdentatus (Girault). Hosts: Sec. par. of Lepidoptera. 
Biology: Schaffner, 1959, loc. cit. supra, p. 86. 

splendens (Cook and Davis). Maine, Pa. Host: Eulophus anomocerus 
(Crawford). 

tarsalis (Ashmead). Hosts: Sec. par. of Lepidoptera. 
Biology: Schaffner, 1959, loc. cit. supra, p. 86. 

wilderi (Howard). Minn. Host: GeZis sp. in spider egg case. 



ELASMIDAE 235 

Genus ENTEDON Dolman, p. 470 

Taxonomy: Hedqvist, 1963, loc. cit. supra, p. 144. 

leucogramma (Ratzeburg). Conn., Pa., S. C; Europe. 

Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 145-146. — Graham, 1963. 
Soc. Brit. Ent. Trans. 15: 194-195 (possible syn.). 

Genus PARACRIAS Ashmead, pp. 470-471 

mirus (Girault). N. Y., Ohio. Hosts: Thysanocnemis spp. in ash seeds. 

*Genus CHRYSONOTOMYIA Ashmead 
Chrysonotomyia Ashmead, 1904. Carnegie Mus. Mem. 1: 344. 

Type : Eulophus auripunctatus Ashmead. Orig. desig. 

Insert in Entedontinae following Emersonopsis Girault, p. 471. 

'^'auripunctata (Ashmead). Fla. ; West Indies. Hosts: Ctenodactylomyia wat- 
soni Felt gall on Coccolobus uvifera; gall on Pithecolobium. 
Eulophus auripunctatus Ashmead, 1894. Linn. Soc. London Jour., Zool. 

25: 166. 9. 

Family ELASMIDAE 

This family is unnecessary: Elasmus itself is clearly eulophid, and the 
other genera now placed in the Elasmidae are closely related to Aphelinus 
Dalman. See Compere, 1947. Bui. Ent. Res. 38: 381. 

Genus ELASMUS Wesfwood, pp. 471-472 

apanteli Gahan. Wis. 

atratus Howard. N. H., Mass. 

Biology: Schaffner, 1959, loc. cit. supra, p. 86. 

*maculatus Howard. Fla.; West Indies. Hosts: Apanteles spp. 

Elasmus maculaius Howard, 1894. Linn. Soc. London Jour., Zool. 25: 
103. ?. 

setosiscutellatus Crawford. Hosts: Apanteles spp. on lepidopterous larvae. 

Family THYSANIDAE 

Genus THYSANUS Walker, pp. 472-474 

citrifolii (Ashmead). Delete entry; species cited in error for Siphonophora 
citrifolii Ashmead, an aphid. 

coquilletti (Ashmead). Fla. Hosts: Aleuroplatus coronatus (Quaint.), A. 

gelatinosus (Ckll.), Aleurodes sp. 

dactylopii (Ashmead). Tex., N. Mex. Host: Leucopis sp. pred. on mealybugs. 

elongatus (Girault). N. J., Md. 

Taxonomy: Ferriere, 1957. Opusc. Zool. 10: 6 (Signiphorina Nikols- 
kaja). 

fasciatus (Girault). 

Biology: Arnaud, 1959. Ent. News 70: 163 (primary par.). 

flavopalliatus (Ashmead). Hosts: ProspaZicWa spp. 

Biology: Muma, 1959. Jour. Econ. Ent. 52: 582 (sec. par.). — Hughes, 
1960. Fla. Ent. 43: 90 (sec. par.). 

louisianae Dozier. Fla. 

merceti (Malenotti). Calif. Hosts: Hemiberlesia lataniae (Sign.), H. rapax 
(Comst.) . 
Biology: DeBach, Kennett, and Pence, 1958. Jour. Econ. Ent. 51: 
114-115 (primary par.). — Valenzuela, 1962. Agronomia (Colom- 
bia) 1: 41 (primary par.) . 

iiiger (Ashmead). 

Taxonomy: De Santis, 1957. La Plata Univ. Nac, Mus. Notas 19: 113. 



236 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

occidentalis (Howard). Fla. 

pulcher (Girault). Ark. 

thoreauini (Girault). Hosts: Hemiberlesia lataniae (Sign.), H. rapax 
(Comst.). 
Biology: DeBach, Kennett, and Pence, 1958, loc. cit. supra, pp. 114-115 
(primary par.). 

Family TANAOSTIGMATIDAE 

Genus TANAOSTIGMODES Ashmead, p. 474 

albiclavus Girault, corrected spelling of albiclava Girault of Catalog. Host: 
Gall on Mimosa; prob. phytophagous. 
Nomenclature : Peck, 1963, loc. cit. supra, p. 331. 

Family ENCYRTIDAE 

Taxonomy: Ferriere, 1953. Schweiz. Ent. Ges. Mitt. 26: 1-45. — Erdos 
and Novicky, 1955. Beitr. z. Ent. 5: 165-202.— Hoffer, 1955. Na- 
rod. Mus. Praze Shorn. 11 (B) : 3-21.— Hoffer in Kratochvil, 1957. 
Cesk. Akad. Ved. Stud. Pram., Sek. Biol. 18: 245-269.— Hoffer, 
1960. Mus. Nat. Pragae Acta Faun. Ent. 6: 93-119. — Compere 
and Annecke, 1960. Ent. Soc. S. Afr. Jour. 23: 375-389.— Tachi- 
kawa, 1963. Ehime Univ. Mem. 9: 1-264 (host list, pp. 9-29).— De 
Santis, 1963. Com. Inv. Cient. Prov. B. A. An. 4: 9-398. 

Subfamily ENCYRTINAE 

Genus TINEOPHOCTONUS Ashmead, p. 475 
armatus (Ashmead). Fla.; Europe, S. Amer. Host: Anobimn punctatum 



(DeG.) 



*Genus ERICYDNUS Walker 



Ericydnus Walker, 1837. Ent. Mag. 4 : 363. 

Type: Ericydnus paludatus Walker. Desig. by Westwood, 
1840. 

Insert in Encyrtinae following Tineophoctonus Ashmead, p. 475. 

*sipylu8 (Walker). Calif.; Europe. 

Encyrtus sipylus Walker, 1837. Ent. Mag. 4: 445. 9. 

Genus LEPTOMASTIX Foerster, p. 475 

dactylopii Howard. 

Biology: Clausen, 1956, loc. cit. supra, p. 94. — Lloyd, 1958. Canad. Ent. 
90: 450-461. 

Genus ANAGYRUS Howard, pp. 475-476 

aiitoninae Timberlake. Mexico. 

Biology: Clausen, 1956, loc. cit. supra, p. 75. — Questel and Genung, 1957. 
Fla. Ent. 40: 123-125.— Dean and Schuster, 1958. Jour. Econ. 
Ent. 51: 363-365.— Dean, 1960. Jour. Econ. Ent. 53: 694. 

fusciventris (Girault). Calif.; Hawaii, West Indies. 
Biology: Clausen, 1956, loc. cit. supra, p. 75. 

"'graminicolens Dozier. Tex.; West Indies. Host: ? Antonina graminis 
(Mask.). 
Anagyrus graminicolens Dozier, 1937. Univ. P. R. Jour. Agr. 21: 123. 9. 
[This may be a synonym of Cerchysius pulchricornis Howard, 
1894, described from St. Vincent, W. I.]. 

pallidipes Ashmead. N. Y. south to Fla. west to Tex. 



ENCYRTIDAE 237 

*Genus NEODUSMETIA Kerrich 

Neodusmetia Kerrich, 1964. Entomophaga 9 : 76. 

Type: Dusmetia sangwani Subba Rao. Orig. desig. 

Taxonomy: Burks, 1958 (1957). Brooklyn Ent. Soc. Bui. 52: 124-125.— 
Richards, 1958. Roy. Ent. Soc. London Proc, ser B, 27: 61.— Ker- 
rich, 1964, loc. cit. supra, pp. 75-77. 

Insert in Encyrtinae following Anagyrus Howard, p. 476. 

*8angwani (Rao). Tex., Ariz.; Hawaii, Mexico, India, Pakistan. Host: An- 
tonina graminis (Mask.). 
Dusmetia sangtcani Rao, Dec. 1957. Indian Acad. Sci. Proc. 46: 385. ?, 

S- 
*Dusmetia indica Burks, Jan. 1958 (1957). Brooklyn Ent. Soc. Bui. 52: 

125. 9, $. (Syn. by Narayanan, Subba Rao, and Rao, 1960. Natl. 

Inst. Sci. India Proc. 26: 175.) 
Biology: Dean, Schuster, and Bailey, 1961. Jour. Econ. Ent. 54: 953. — 

Beardsley, 1961. Hawaii. Ent. Soc. Proc. 17: 327.— Kerrich, 1964, 

loc. cit. supra, p. 75. 

Genus COMPERIA Gomes, p. 476 

merceti (Compere). Calif. 

Biology: Roth and Willis, 1960. Smithsn. Misc. Coll. 141: 244, 255, 351. 

Genus APOANAGYRUS Compere, p. 476 

califomicus Compere. Ariz. 

Genus ECTROMATOPSIS Compere, p. 476 

americana (Howard). Tex., Ariz., Nev.; Mexico. 

*Genus HUNGARIELLA Erdos 
(^Tefrocnemus of Catalog, pp. 447—478) 
Hungariella Erdos, 1946. Mus. Natl. Hungarici Ann. Hist.-Nat. 39: 144. 

Type: Hungariella piceae Erdos. Orig. desig. 
Delete generic syn. in Catalog, p. 477, under Tetracnemus; Charitopus 
Foerster extralimital and valid. 

Taxonomy: Erdos, 1955. Acta Zool. [Budapest] 1: 218.— Hoffer, 1959. 
Mus. Nat. Pragae, Acta Ent. 33: 13-14.— Hoffer, 1960. Mus. Nat. 
Pragae, Acta Faun. Ent. 6: 101. 

peregrina (Compere). Calif.; S. Africa, S. America. Hosts: Pseudococcus 
gahani Green, P. marithnus (Ehrh.). 
Biology: Clausen, 1956, loc. cit. supra, p. 130. 

pretiosus (Timberlake). Calif.; Australia. Host: Pseudococcus maritimus 
(Ehrh.). 
Biology: Clausen, 1956, loc. cit. supra, p. 130. — Bartlett and Lloyd, 1958. 
Jour. Econ. Ent. 51: 92. 

*Genus TETRACNEMUS Wesfwood 
( = refrac/ad/a Howard of Catalog, p. 478) 

Paracalocerinus Girault, 1915; Catalog, p. 503. 

Masia Mercet, 1919 ; Catalog, p. 503. 

*AnusieUa Mercet, 1923. Real Soc. Esp. Hist. Nat. Bol. 23 : 286. 

Type: Anusiella heydeni Mercet. Orig. desig. 
*Placoceras Erdos, 1946. Frag. Faun. Hung. 9: 1. 

Type: Placoceras colocense Erdos. By monotypy. 
Taxonomy: Erdos, 1955, loc. cit. supra, p. 221-223. — Hoffer, 1960, loc. cit. 
supra, p. 101. 

americanus (Girault). Transf. from Paracalocerinus Girault, p. 503, teste 
Peck, 1963, loc. cit. supra, p. 347. 

marylandia (Girault). Transf. from Paracalocerinus Girault, p. 503, teste 
Peck, 1963, loc. cit. supra, p. 347. 



238 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

tertius (Girault). Transf. from Paracalocerinus Girault, p. 503 teste Peck, 
1963, loc. cit. supra, p. 347. 

Genus TETRACNEMOPSIS Ashmead, p. 478 

westwoodi (Cockerell). Que., Maine. 

Genus PENTACNEMUS Howard, p. 478 

bucculatricis Howard. Calif. Host: Argyresthia pilatella Braun. 

Genus LEPTOMASTIDEA Mercet, p. 479 

abiiormis (Girault). 

Biology: Clausen, 1956, loc. cit. supra, p. 109. 

Genus ENCYRTUS Latreille, pp. 479-480 

* Allorhopoideus Brethes, 1916. Buenos Aires Mus. Nac. Hist. Nat. Ann. 

27: 425. 

Type: Allorhopoideus mirabilis Brethes. Orig. desig. 
Taxonomy: Masi, 1921. [Genova]. Mus. Civ. di Stor. Nat. Ann., ser. 3, 

9: 239. 

f uscus ( Howard ) . 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1256. 

peckhaniii (Ashmead). Que. and Ont. south to N. C, west to Wis. and Tenn. 
Host: Eulecanium quercifex (Fitch). 

Genus PARALITOMASTIX Mercet, p. 481 

pyralidis (Ashmead). 

Biology: Clausen, 1956, loc. cit. supra, p. 22. 

Genus COPIDOSOMA Ratzeburg, pp. 481-483 

Taxonomy: De Santis, 1963. Com. Inv. Cient. Prov. B. A. An. 4: 
278-280, 284, 299-300. 

bakeri (Howard), new status. N. B. and Ont., south to Mass., west to Alta. 

and Calif. Hosts: 10 spp. of Noctuidae, listed in Catalog under 

subspp. of bakeri, p. 482. 
Copidosoma bakeri bakeri (Howard) of Catalog, raised to sp. by Peck, 

1963, loc. cit. supra, p. 361. 
Berecyntus bakeri var. arizonensis Girault, 1916 (syn. by Peck, 1963, loc. 

cit. supra, p. 361). 
Berecyntus bakeri var. gemma Girault, 1916 (syn. by Peck, 1963, loc. cit. 

supra, p. 361) . 
Berecyntus bakeri var. euxoae Strickland, 1916 (syn. by Peck, 1963, loc. 

cit. supra, p. 361). 

*deceptor Miller. Throughout Canada and north. U. S. Hosts: Epinotia nan- 
a7ia (Treit.), Acleris variana (Fern.), Duvita vittella Busck, £'?<- 
cordylia huntella Kief., Recurvaria apicitripunctella Clem., R. 
canusella Free., R. milleri Busck, R. moreonella Hein., R. nanella 
(Hbn.), R. starki Free., R. thujaella Kearf., Exoteleia dodecella 
(L.), E. pinifoliella (Cham.), Argyresthia aureoargentella 
B rower. 
Copidosoma deceptor Miller, 1958. Pan-Pacific Ent. 34: 58. 9, 5- 
Biology: Stark, 1961. Canad. Jour. Zool. 39: 895.— Telford, 1961. Jour. 
Econ. Ent. 54: 348.— Richards, 1963. Canad. Ent. 95: 519. 

gelechiae Howard, new status. 

Copidosoma gelechiae gelechiae (Howard) of Catalog, raised to sp. by 

Peck, 1963, loc. cit. supra, p. 363. 
Copidosoma gelechiae var. saga Girault, 1920 (syn. by Peck, 1963, loc. 

cit. supra, p. 364). 
Biology: Schaffner, 1959, loc. cit. supra, p. 86. — Miller, 1963. Ohio Jour. 

Sci. 63: 71. 

geniculatuin (Dalman). Ont.; Europe. Host: Rhyacionia frustrana 
(Comst.). 



ENCYRTIDAE 239 

Biology : SchaflFner, 1959, loc. cit. supra, p. 86. 
lymani Howard. Wis. Host: Rhyacionia buoliana (Schiflf.) 

Genus CERCHYSIUS Westwood, p. 483 
hubbardii Ashmead. Mont., Idaho, Ariz. 
marylandicus Girault. N. H., Ind. 

pallipes (Provancher). 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1257. 

Genus BAEOCHARIS Mayr, p. 483 

*pascuoruni Mayr. N. H.; Europe. Host: Eriopeltis festucae Fonsc. 

Baeocharis pascuorum Mayr, 1875. Zool. Bot. Ges. Wien. Verh. 25: 768. 

?, $. 

Genus CHALCASPIS Howard, p. 484 

pergandei Howard. Okla., Tex. 

Genus BOTHRIENCYRTUS Timberlake, p. 484 

planifonnis (Howard). Hosts: Hemerobius pacificus Banks; reared from co- 
coons of undet. Hemerobiidae. 

Genus PENTELICUS Howard, p. 485 

Taxonomy: Kerrich, 1963. Beitr. z. Ent. 13: 364. 

aldrichi Howard. N. H., Nebr. 

Genus HEMAENASIUS Ashmead, p. 485 

confusus Ashmead. Nebr. 

Genus TRICHOMASTHUS Thomson, p. 485 

Taxonomy: Alam, 1957. Roy. Ent. Soc. London Trans. 109: 446-447. 

^albimanus Thomson. N. H., Minn.; Europe. Host: Eriopeltis festucae 
Fonsc. 
Trichomasthus albimanus Thomson, 1875. Hym. Scand., v. 4, p. 144, 5 , 

$. 
Taxonomy: Alam, 1957, loc. cit. supra, p. 447. 

*c>aneus (Dalman). N. H., Mass.; Europe. Host: Eriopeltis festucae Fonsc. 
Encyrius cyaneus Dalman, 1820. Svenska. Vetensk. Akad. Handl. 41: 

160. 9. 
Taxonomy: Alam, 1957, loc. cit. supra, pp. 447, 450. 

cyanifrons (Dalman). N. H. 

Taxonomy: Alam, 1957, loc. cit. supra, p. 447. 

*Genus MIRA Schellenberg 

Mira Schellenberg, 1803. Gen. des Mouch., p. 68. 

Type : Mira macrocera Schellenberg. By monotypy. 

Insert in Encyrtinae following Trichomasthus Thomson, p. 485. 
*macrocera Schellenberg. Idaho; Europe. 

Mira macrocera Schellenberg, 1803. Gen. des Mouch., pp. 68-69. 

■•■Genus HAMBLETONIA Compere 

Hambletonia Compere, 1936. Hawaii. Ent. Soc. Proc. 9: 172. 

Type: Hambletonia pseudococcina Compere. By monotypy. 

Insert in Encyrtinae after Discodes Foerster, p. 485. 

*pseudococcina Compere. Fla., Tex., Calif; Hawaii, West Indies, Brazil. 
Host: Dysmicoccus brcvipes (Ckll.). 
Hambletonia pseudococcina Compere, 1936. Hawaii. Ent. Soc. Proc. 9: 
173. 9, $. 



240 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Biology: Carter, 1937. Jour. Econ. Ent. 30: 370.— Plank, 1938. P. R. 
Expt. Sta. Rpt. for 1937, pp. 98-99.— Bartlett, 1939. P. R. Univ. 
Jour. Agr. 23 : 67-71. — Clausen, 1956, loc. cit. supra, p. 106. 

Genus HOMALOTYLUS Mayr, p. 486 

*Lepidaphycus Blanchard, 1936. Soc. Ent. Arg. Rev. 8 : 13. 

Type: Lepidaphycus bosqi Blanchard. Orig. desig. 

Taxonomy: d'Araujo e Silva, 1945 (1944). Bol. Fitossanit. (Brazil) 1: 
29.— Hoflfer, 1959. Acta Ent. Mus. Nat. Pragae 33: 30.— De 
Santis, 1963. Com. Inv. Cient. Prov. B. A. An. 4: 162 (Echthro- 
plexis Foerster). 

Genus ANISOTYLUS Timberlake, p. 487 

pallentipes Timberlake. Mo. 

Genus PRIONOMITUS Mayr, p. 487 

mitratus (Dalman). B. C, Calif.; Europe. Hosts: Nymphs of Psyllidae. 
Biology: Jensen, 1957. Hilgardia 27: 72-78, 87-89, 92-95. 

Genus PSYLLAEPHAGUS Ashmead, pp. 487-488 

pachypsyllae (Howard). Calif. Hosts: Pachypsylla celtidisvesicula Riley, P. 
venusta (O. S.), Trioza beameri Tuthill. 
Biology: Jensen, 1957, loc. cit. supra, pp. 79, 95. 

trioziphagus (Howard). Ariz. Host: Kuwayama medicaginis (Cwfd.). 
Biology: Cazier, 1964. Pan-Pacific Ent. 40: 73. 

Genus MICROTERYS Thomson, pp. 488-489 

*Apentelicus Fullaway, 1913. Hawaii. Agr. Expt. Sta. Ann. Rpt. 1912, 
p. 26. 

Type: Apentelicus kotinskyi Fullaway. Orig. desig. 
Taxonomy: Timberlake, 1919. Hawaii. Ent. Soc. Proc. 4: 184. 

flavus (Howard). Cosmopolitan. 

Biology: Bartlett and Lagace, 1961. Ent. Soc. Amer. Ann. 54: 222-227. 
—Bartlett, 1964. Ent. Soc. Amer. Ann. 57: 344.— Bartlett and 
Ball, 1964. Ent. Soc. Amer. Ann. 57: 496-503. 
Taxonomy: Annecke, 1962. Ent. Soc. S. Afr. Jour. 25: 174, 175. 

Genus ISODROMUS Howard, p. 489 

iceryae Howard. Hosts: Chrysopa bimaculata McCL, C. cubana Hagan, C. 
sanchezi Navas. 

*montanus Slosson. N. H. Host: Chrysopa sp. 

Isodromus montanus Slosson, 1895. Ent. News 6: 278. [Type lost, but 
this may be lsodro')nus vinulus (Dalman), subsequently collected 
in P. E. I.] 
Taxonomy: Peck, 1963, loc. cit. supra, p. 395. 

montinus (Packard). Colo. 

puncticeps (Howard). N. J., Pa., Kans. 

*vinulus (Dalman). P. E. I.; Europe. 

Encyrtus vinulus Dalman, 1820. Svenska Vetensk. Akad. Handl. 41: 160. 
Taxonomy: Hellen, 1949. Notulae Ent. 29: 46 {Nobrimus Thomson). 

Genus COCCIDENCYRTUS Ashmead, p. 490 

*Coccidencyrtoides Blanchard, 1940. Soc. Cient. Arg. An. 130: 107. 

Type: Not desig. 
Taxonomy: Compere and Annecke, 1961. Ent. Soc. S. Afr. Jour. 24: 
59-61. 

ochraceipes Gahan. Calif.; Hawaii, Brazil. 

Genus PSEUDAPHYCUS Clausen, pp. 490-491 

Taxonomy: Hoifer, 1960. Mus. Nat. Pragae Acta Faun. Ent. 6: 94-96. 



ENCYRTIDAE 241 

malinus Gahan. Holarctic. 

Biology: Clausen, 1956, loc. cit. supra, p. 125. 

Taxonomy: Jasnoch, 1957. Ent. Obozr. 36: 720.— Hoffer, 1960, loc. cit. 

supra, pp. 95, 97. — Sugonyaev, 1962. Akad. Nauk S. S. S. R. Izv., 

ser. Biol., 5: 757. 

Genus ACEROPHAGUS E. A. Smith, p. 491 

notativentris (Girault). 

Biology: Flanders, 1959. Jour. Econ. Ent. 52: 72. 

Genus APHYCUS Mayr, pp. 492-495 

Aphycus subg. Aphycaspis Hoffer, 1954. Ochr. Prir. 9 : 170. N. syn. 

Type: Not desig. ; 2 spp. incl. 
Taxonomy: Compere, 1957. Portici Lab. Ent. Agr. Bol. 15: 221-230 

(Metaphycus.) — Compere and Annecke, 1960. Ent. Soc. S. Afr. 

Jour. 23: 375-389. 

alberti Howard. Calif.; Australia, Hawaii. 

Taxonomy: Compere, 1957, loc. cit. supra, pp. 222-224. 

apicalis (Dalman). [N. Amer. record almost certainly an error.] 

Taxonomy: Compere and Annecke, 1960, loc, cit. supra, p. 377-378. 

bifasciatus (Timberlake) . 

Taxonomy: Compere and Annecke, 1961. Ent. Soc. S. Afr. Jour. 24: 39, 

44 (W ater s tonia Mercet) . 

californicus Howard. 

Taxonomy: Compere and Annecke, 1961. Ent. Soc. S. Afr. Jour. 24: 34. 

eriococci Timberlake. Fla., Tex. Host: Coccus hesperidum {h.). 

eruptor Howard. Host : Ceroplastes floridensis Comstock. 

flavus Howard. Cosmopolitan. Hosts: Coccus hesperidum (L.), Saissetia 
oleae (Bern.). 
Taxonomy and Biology: Compere, 1957, loc. cit. supra, pp. 229-230. 

hederaceus (Westwood). 

Taxonomy: Compere, 1957, loc. cit. supra, p. 222. — Graham, 1959. Ent. 
Tidskr. 79 : 173. 

helvolus Compere. 

Taxonomy: Compere, 1957, loc. cit. supra, p. 223. 

Biology: Clausen, 1956, loc. cit. supra, pp. 80-81. — Flanders, 1959. Jour. 

Econ. Ent. 52: 74, 596-599.— Damaino, 1963. Rev. di Agr. Sub- 

trop. e Trop. 57: 452. 

lounsburyi Howard. Hawaii. Host: Saissetia hemispherica (Targ.). 

Biology: Clausen, 1956, loc. cit. supra, p. 81. — Beardsley, 1962. Hawaii. 
Ent. Soc. Proc. 18: 16. 

luteolus Timberlake. 

Taxonomy: Compere, 1957, loc. cit. supra, p. 229. 

Biology: Flanders, 1959, loc. cit. supra, p. 74. — Bartlett and Ball, 1964. 
Ent. Soc. Amer. Ann. 57: 496-503. 

maculipes Howard. Tex. 

mexicanus Howard. La. ; Mexico. Delete Pa. from range, 

pulvinariae Howard. 

Taxonomy: Compere and Annecke, 1961, loc. cit. supra, p. 34. 

Stanley! (Compere). 

Biology: Clausen, 1956, loc. cit. supra, pp. 81-82. 

Genus OOENCYRTUS, p. 496 

Taxonomy: Hoffer, 1963. Mus. Nat. Pragae Acta Ent. 35: 560-563 (div. 
of genus into subgenera). 



242 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

clisiocampae (Ashmead). Md., Ark., N. Hex., Idaho, Utah. Host: Malaco- 
soma fragile (Str.), prob. secondary par. 

kuwanai (Howard). N. Y.; Japan, Europe. 

Biology: Clausen, 1956, loc. cit. supra, pp. 116-118. — Templado, 1957. 
Inst. Biol. Barcelona, Apl. Pubs. 25: 119.— Hitchkock, 1959. Jour. 
Econ. Ent. 52: 764.— Tadic, Vasic, and Salatic, 1959. Zast. Bilja 
52-53: 47-52.— Dowden, 1961. Jour. Econ. Ent. 54: 876.— Bjegovic, 
1963. Zast. Bilja 14 : 544. 

*submetallicus (Howard). Fla.; West Indies, Brazil, Philippines. Host: Ne- 
zara viridula (L.). 
Encyrtus submetallicus Howard, 1896. Linn. Soc. London Jour., Zool. 26: 
151. 9. 

Genus SYRPHOPHAGUS Ashmead, p. 497 

quadrimaculati (Ashmead), corrected spelling of quadrimaculatae of Catalog, 
teste Peck, 1963, loc. cit. supra, p. 433. 

Genus PSEUDENCYRTUS Ashmead, p. 497 

cecidomyiae (Howard). Que., Ont., N. H., Mich. Host: Rhabdophaga strobi- 
loides (O. S.). 

Genus HOLCENCYRTUS Ashmead, p. 497 
physokermis Girault. Que., Maine., N. H. 

Genus CHEILONEURUS Westwood, pp. 497-498 

elegans (Dalman). [All forms currently under this name may not be 

conspecific] 
Biology: Ferriere, 1952. Ninth Int. Cong. Ent. Amsterdam Trans, v. 1, 

p. 594.— Ferriere, 1956. Lab. Zool. Gen. e Agr. Portici Bol. 33: 

354.— Sugonyaev, 1958. Ent. Obozr. 37 : 309. 
Taxonomy: Claridge, 1958, Ent. Mo. Mag. 94: 157. 

inimicus Compere. Fla., Tex. 

Genus CHRYSOPOPHAGUS Ashmead, p. 499 

americanus (Perkins). Calif.; Hawaii. 

amplicornis (Gahan). Ariz., Calif. Hosts: Dactylopius sp. on Opuntia, An- 
tonina graminis (Mask.), poss. sec. parasite. 

compressicornis Ashmead. Hosts: Hyperparasite of Chrysopa spp. 
Biology: Muma, 1959. Fla. Ent. 42: 151-152. 

Genus HABROLEPIS Foerster, pp. 499-500 

rouxi Compere. 

Biology: Clausen, 1956, loc. cit. supra, p. 106. — Flanders, 1956. Lab. 
Zool. Gen. e Agr. Portici Bol. 33: 15-16, 25.— Smith, 1957. Canad. 
Ent. 89: 219-228.— Doutt, 1959. Ann. Rev. Ent. 4: 170-171.— An- 
necke, 1963. Ent. Soc. S. Afr. Jour. 26: 207. 

Genus ADELENCYRTUS Ashmead, p. 500 

Taxonomy: Compere and Annecke, 1961, loc. cit. supra, pp. 47-49 (key to 
spp.). 

aulacaspidis (Brethes). Nebr. ; S. America, Europe. 

Genus ANICETUS Howard, p. 500 

annulatus Timberlake. Tex.; Mexico, Hawaii, Japan, China. Host: Coccus 
hesperidimi L. 

*toumeyellae Milliron. Del., Va. Host: Toumeyella liriodendri (Gmel.). 

Anicetus toumeyellae Milliron, 1959. Ent. Soc. Amer. Ann. 62: 29. 9, $. 



ENCYRTIDAE 243 

Genus ZARHOPALUS Ashmead, p. 501 

inquisitor (Howard). Md., Ariz.; Mexico, West Indies. Host: Ferrisiana vir- 
gata (Ckll.). 

Genus Coccob/us Ratzeburg, p. 501 

Delete; extralimital. 

diaspidis Ashmead. Syn. of Arrhenophagus chionaspidis Aurivillius, p. 507. 

Genus PSEUDHOMALOPODA Girault, p. 501 

prima Girault. 

Biology: Clancy et al, 1963. Jour. Econ. Ent. 56: 604. 

Genus EUSEMION Dahlbom, pp. 501-502 

*comiger (Walker). N. H.; Europe. Hosts: Lepidosaphes ulmi (h.) , Lecan- 
ium corni Bche., Pulvinaria vitis (L.), other spp. in European lit. 

Encyrtus corniger Walker, 1837. Ent. Mag. 5: 114. $. 

Taxonomy: Ferriere, 1949. Schweiz. Ent. Ges. Mitt. 22: 383. 

Biology: Schmutrerer, 1951. Ztschr. f. Angew. Ent. 33: 121. — Zak- 
Ogaza, 1961. Polskie Pismo Ent. 31: 365. 

*Genus COELOPENCYRTUS Timberlake 

Coelopencyrtus Timberlake, 1919. Ent. Soc. Hawaii Proc. 4: 218. 

Type : Coelopencyrtus odyneri Timberlake. Orig. Desig. 
*I\'esencyrtus Timberlake, 1919, loc. cit. supra, p. 223. 

Type: Adelencyrtus kaalae Ashmead. Orig. desig. 
Taxonomy: Burks, 1958. Wash. Acad. Sci. Jour. 48: 22. — Tryapitzin, 
1960. Ent. Obozr. 39: 697. 

Insert in Encyrtinae following Atropates Howard, p. 502. 

*hylaei Burks. Md., Nebr. Hosts: Hylaeiis cressoni (Ckll.), Hylaeus sp. 

Coelopencyrtus hylaei Burks, 1958. Wash. Acad. Sci. Jour. 48: 25. 9, $. 
Biology: Brandhorst, 1962. Ent. Soc. Amer. Ann. 55: 477. 

*hylaeoleler Burks. Ont., Idaho, Utah. Host: Hylaeus sp. 

Coelopencyrtus hylaeoleter Burks, 1958. Wash. Acad. Sci. Jour. 48: 24. 

Genus XANTHOENCYRTUS Ashmead, p. 502 

Delete Mirastymachus Girault from syn. and transf. to syn. under 
Rhopus Foerster, infra. 

Taxonomy: Ghesquiere, 1956. Lab. Zool. Gen. e Agr. Portici Bol. 33: 
683-689, 692-695. 

americanus (Girault). Transf. to Rhopus Foerster. 

nigroclavatus Ashmead. Md., Va., Nebr. 

♦Genus RHOPUS Foerster 

{ = Xanfhoencyrtus Ashmead of Catalog, p. 502, in Part) 

Rhopus Foerster, 1856. Hym. Stud., v. 2, pp. 34, 37, 151. 

Type: Encyrtus piso Walker. Orig. desig. 
Mirastymachus Girault, 1915 (transf. from syn. under Xanthoencyrtus 

Ashmead, teste Ghesquiere, 1956, loc. cit. supra, p. 689). 
Taxonomy : Ghesquiere, 1956, loc. cit. supra, pp. 683-692. 

Insert in Encyrtinae following Xanthoencyrtus Ashmead, p. 502. 

americanus (Girault). Del., Oreg. Transf. from Xanthoencyrtus Ashmead, 
teste Ghesquiere, 1956, loc. cit. supra, p. 698. 

Genus PAURIDIA Timberlake, p. 502 

peregrina Timberlake. Calif.; Hawaii, China. Host: Pseudococcus citri 
(Risso). 



244 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus PLAGIOMERUS Crawford, p. 502 
diaspidis Crawford. Fla., Tex. ; Mexico. Delete D. C. from range. 

*Genus TRECHNITES Thomson 
(= Psylledontus Crawford of Catalog, pp. 502-503) 

Trechnites Thomson, 1875. Hym. Scand., v. 4, p. 118. 

Type : Metallon fuscitarsis Thomson. By monotypy. 
Taxonomy: Hoffer, 1960. Mus. Nat. Pragae Acta Faun. Ent. 6: 109.— 
Ferriere, 1961. Entomophaga 6: 39-40. 

insidiosus (Crawford). Calif. 

Taxonomy: Ferriere, 1961, loc. cit. supra, p. 41. 

Biology: Madsen, Westigard, and Sisson, 1963. Canad. Ent. 95: 842-843. 

Genus AGROMYZAPHAGUS Gahan, p. 503 

detrimentosus Gahan. Wis., Iowa, Utah, Calif. 

Genus Paraca/ocerinus Girault, p. 503 

Transf. to syn. under Tetracnemus Westwood ( = Tetracladia Howard of 
Catalog, p. 478) ; transf. included spp. to Tetracnemus Westwood supra, not 
Tetracnemus of Catalog, p. 477. 

Genus IXODIPHAGUS Howard, p. 503 

texanus Howard. Iowa, Mont. 

Taxonomy : Quaraishi, 1958. Amer. Midland Nat. 59 : 489-504. 

Genus HUNTERELLUS Howard, pp. 503-504 

hookeri Howard. Cosmopolitan. 

Taxonomy: Quaraishi, 1958, loc. cit. supra, pp. 489-504. 
Biology: Klyushkina, 1958. Zool. Zhur. 37: 1561.— Ushakova, 1962. 
Akad. Nauk Kazakh. SSR Inst. Zool. Trudy 16: 183. 

Genus PARALEUROCERUS Girault, p. 504 

*Gibberella Miller, 1961. Canad. Ent. 93: 494. N. syn. 

Type: Gibberella scutellata Miller. Orig. desig. 

bicoloripes Girault. N. S., Ont., N. H., Mass., Conn., Mich., Tex. Hosts: 
Litliocolletis caryaefoliella (Clem.), L. lucetiella Clem., L. cincin- 
natiella (Chamb.), L. spp. 

*GibbereUa scutellata Miller, 1961. Canad. Ent. 93: 496. 9, $. N. syn.; 
types examined. 

Biology: Schaffner, 1959, loc. cit. supra, p. 87. 

Genus FORMICENCYRTUS Girault, p. 504 
thoreauini Girault. Ariz. 

Genus COMPERIELLA Howard, pp. 504-505 

bif asciata Howard. Fla., Calif. ; Japan. 

Biology: Clausen, 1956, loc. cit. supra, pp. 96-97. — Flanders, 1956. Por- 
tici Lab. Zool. Gen. e Agr. Bol. 33: 25-26.— Smith, 1957. Canad. 
Ent. 89: 219-228.— Teran and DeBach, 1963. Rev. Agron. No- 
roeste Arg. 4 : 5-23. 

Genus COCCIDOXENUS Crawford, p. 505 
mexicanus Girault, Ariz. ; Mexico, West Indies. 
nubilipennis (Girault). Va. 

Genus QUAYLEA Timberlake, p. 505 

whittieri (Girault). Calif .; Hawaii, Australia. 

Biologry: Clausen, 1956, loc. cit. supra, pp. 125-126. 



EUPELMIDAE 245 

Genus ANARHOPUS Timberlake, p. 505 

sydneyensis Timberlake. Calif. ; Australia. 

Biology: Clausen, 1956, loc. cit. supra, pp. 75, 130. — Flanders, 1959. 
Jour. Econ. Ent. 52: 75. 

Genus PSEUDORHOPUS Timberlake, pp. 505-506 

Taxonomy: Alam, 1957. Roy. Ent. Sec. London Trans. 109: 459-460. 
hartmani TimlDerlake. 

Taxonomy: Alam, 1957, loc. cit. supra, p. 460. 

Subfamily ARRHENOPHAGINAE 

Genus ARRHENOPHAGUS Aurivillius, pp. 506-507 

Taxonomy: Man Mohan, 1963. Ztschr. f. Parasitenk. 22: 394. 

chionaspidis Aurivillius. 

Coccobiits diaspidis Ashmead, 1900 (syn. by Ghesquiere, in litt.) . 

Taxonomy : Tachikawa, 1957. Jap. Jour. Appl. Ent. and Zool. 1 : 174. 

Biology: Benassy, 1958. Acad. Sci. (Paris) Compt. Rend. 246: 180.— Ta- 
chikawa, 1958. Ins. Mats. 21: 118.— Benassy, 1961. Ann. des fipi- 
phyt., ser. C, 12 : 42. 

Family EUPELMIDAE 

Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 131-132. 

Genus METAPELMA Westwood, p. 507 
schwarzi (Ashmead). Ark., Oreg. 

Genus EUSANDALUM Ratzeburg, pp. 507-508 

Taxonomy: Peck, 1963, loc. cit. supra, p. 472. 

acmaeoderae Rohwer, change to Girault. 

Eusandalutn acmaeoderae Girault, Mar. 1917. 
Eusandalutn acuiaeorerae Rohwer, June 1917. 

Genus CALOSOTA Curtis, p. 508 

Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 138-139. 
longiventris (Ashmead). Idaho. 

Genus EUPELMUS Ddman, pp. 508-511 

Taxonomy: Hedqvist, 1963, loc. cit. supra, p. 135. 

allynii (French). Hosts: Bruchophagus spp., Antistrophus pisum (Walsh) 
galls on Lygodesmia. 
Biology: Wester, 1956. Ent. Soc. Wash. Proc. 58: 285-286.— Neunzig 
and Gyrisco, 1959. Jour. Econ. Ent. 52: 900-901.— Allen, 1962. 
U. S. Dept. Agr. Tech. Bui. 1265, pp. 98-99.— Brandhorst, 1964. Ent. 
Soc. Amer. Ann. 57: 75-76. 

cyaniceps amicus Girault. Mass. 'Ro&t: Dioryctria disclusa "H-Qm. 

Biology: Schaffner, 1959, loc. cit. supra, p. 87. — Allen, 1962, loc. cit. 
supra, pp. 97-98. 

cyaniceps cyaniceps Ashmead. Calif. 

Biology: Allen, 1962, loc. cit. supra, pp. 96-97. 

Taxonomy: Finlayson, 1962. Canad. Ent. 94: 277, 278-279 (imm. stages). 

brevicauda (Crawford). Calif. 

inyoensis Girault. Host: Pimplopterus sp. 

Biology: Tilden, 1951. Microent. 16: 164, 173. 

limneriae Howard. 

Biology: Allen, 1962, loc. cit. supra, pp. 93-95. 



246 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

neococcidis Peck. Host: Tenodera angustipennis Sauss. 
Biology: Gurney, 1959. Ent. Soc. Wash. Proc. 61: 23. 

pini Taylor. N. H. 

popa Girault. Okla.; Hawaii, West Indies, Cent, and S. America, Orient, 
Australia. 
Biology: Passlow, 1958. Queensl. Jour. Agr. Sci. 15: 35.— Harris, 1961. 
Bui. Ent. Res. 52: 134. 

rhizophelus Brues. Syn. of Eupelmella dryorhizoxeni (Ashmead). 

spongipartus Foerster. N. B., Ont., Wis.; Europe. Hosts: Diprion similis 
(Htg.), Apanteles solitarius (Ratz.). 
Biology: Reeks and Smith, 1956. Canad. Ent. 88: 572.— Allen, 1962, loc. 

cit. supi'ii, pp. 95-96. 
Taxonomy: Finlayson, 1962. Canad. Ent. 94: 277, 279 (imm. stages). 

*urozoniis Dalman. N. J., Md. ; Europe. Host: ? Bruchidms ater (Marsh.). 
Eupehnus urozonus Dalman, 1820. Svenska Vetensk. Akad. Handl. 41: 

378. $. 
Biology: Leonide, 1960. Soc. Ent. France Bui. 65: 274, 279.— Rivosecchi, 

1960. Parassitol. 2: 289.— Feron, Benard, and Poitout, 1961. En- 

tomophaga 6 : 180. 
Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 135-137. 

Genus ARACHNOPHAGA Ashmead, pp. 511-512 

costalis Gahan. 

Biology: Allen, 1962, loc. cit. supra, p. 100. 

frontalis Gahan. 

Biology : Allen, 1962, loc. cit. supra, pp. 99-100. 

longicornis (Brues). 

Biology: Allen, 1962, loc. cit. supra, p. 100. 

picea (Riley). 

Biology: Cazier and Mortenson, 1962. So. Calif. Acad. Sci. Bui. 61: 74, 
83-84. 

Genus EUPELMELLA Masi, p. 512 

Nomenclature: Peck, 1963, loc. cit. supra, p. 492 (Macroneura). 
Taxonomy : Hedqvist, 1963, loc. cit. supra, p. 132. 

canadensis (Provancher) . Syn. of vesicularis (Retzius). 

dryorhizoxeni (Ashmead). Mass. south to Fla. west to Kans. 
Eupelmus rhizophelus Brues, n. syn. ; types examined. 

vesicularis (Retzius). Holarctic. 

Theocolax canadensis Provancher, 1883 (syn. by Burks, 1964, loc. cit. 

supra, p. 1262), 
Taxonomy: Finlayson, 1960. Canad. Ent. 92: 37, 38, 44 (imm. stages).— 

Hedqvist, 1963, loc. cit. supra, pp. 133-134. 
Biology: Allen, 1962, loc. cit. supra, pp. 100-101. 

Genus ANASTATUS Motschulsky, pp. 513-514 

blattidarum Ferriere. Syn. of tenuipes Bolivar y Pieltain. 

disparis Ruschka. 

Biology: Clausen, 1956, loc. cit. supra, p. 76. — Karnozitzky, 1957. Bulg. 
Akad. Nauk. Zool. Inst. Trudov 4: 37, 97.— Templado, 1957. Inst. 
Biol. Apl. Pubs. Barcelona 25: 126. — Vasic and Salatic, 1959. 
Zast. Bilja 52-53: 45. — Stefanov and Keremedziev, 1961. Viss. Le- 
sot. Inst. Sofia Nauch. Triidove 9: 162.— Tadic, 1961. Zast. Bilja 
13: 63-64.— Salatic, 1963. Zast. Bilja 14: 693-699. 

floridanus Roth and Willis. 

Biology: Roth and Willis, 1960. Smithsn. Misc. Coll. 141: 245-246. 

mantis (Ashmead). Ga., 111., Miss., Okla., Tex. 



EUCHARITIDAE 247 

microcentri Gahan. Kans., Tex., Ariz. 
semiflavidus Gahan. Calif. 

*tenuipe8 Bolivar y Pieltain. N. Hemisphere. Host: Egg case of Supella 

supellectilium ( Serv. ) . 
Anastatus tenuipes Bolivar y Pieltain, 1925. Soc. Roy. Ent. :figypte Bui. 

9: 43. 9. 
Anastatus blattidarum Ferriere, 1930 (syn. by Roth and Willis, 1960, loc. 

cit. supra, p. 246). 
Biology: Flock, 1941. Brooklyn Ent. Soc. Bui. 36: 1-4.— Roth and Willis, 

1960, loc. cit. supra, p. 246-247. 

Genus ENCYRTASPIS Ashmead, p. 514 

semirufa Gahan. Ariz. 

Biology: Allen, 1962, loc. cit. stipra, pp. 101-102. 

Genus LECANIOBIUS Ashmead, p. 514 

cockerelli Ashmead. Fla. ; West Indies, S. America. Host: Saissetia oleae 
(Bern.). 

utilis Compere. Calif. ; S. America. 

Biology: Clausen, 1956, loc. cit. supra, p. 108. 

*Genus PHLEBOPENES Perty 

Phlebopenes Perty, 1833. Delect, anim. artic, Brasil., p. 132. 

Type : Phlebopenes splendidus Perty. By monotypy. 
Revision: Roman, 1920. Ark. Zool. 12: 24-30. 

Insert in Eupelmidae following Zaischnopsis Ashmead, p. 514. 

*hetricki Burks. Md., Ga., Fla. 

Phlebopenes hetricki Burks, 1964. Ent. News 75: 1. 9. 

Family EUCHARITIDAE 

Genus PSEUDOCHALCURA Ashmead, pp. 514-515 

califomica (Ashmead). Oreg. 

gibbosa (Provancher) . N. H. 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1258. 

Genus PSEUDOMETAGEA Ashmead, p. 515 

Revision: Burks, 1961. Ent. News 72: 253-257. 

*bakeri Burks. Iowa, Wyo., Colo. 

Pseudometagea bakeri Burks, 1961. Ent. News 72: 256. 9, $. 

hillmeadia Girault. Syn. of schwarzii (Ashmead). 

schwarzii (Ashmead). Que. and Ont. south to Md. and D. C. west to Wis., 
Towa, Nebr. Host : Lasius neoniger Emery. 
Pseudometagea hilhneadia Girault, 1916 (syn. by Burks, loc. cit. supra, 

p. 255). 
Biology: Ayre, 1962. Canad. Jour. Zool. 40: 157-164. 

Genus ORASEMA Cameron, pp. 515-516 

*Semora Cameron, 1909. Amer. Ent. Soc. Trans. 35: 432-433. Preocc. 

Type : Semora xanthopus Cameron. Orig. desig. 
^Semorella Ghesquiere, 1946. Rev. Zool. Bot. Afr. 39: 368. N. name. 
*Semorata Strand, 1947. Folia Zool. Hydrobiol. 11: 393. N. name. 
Taxonomy: Kerrich, 1963. Bui. Ent. Res. 54: 366. 

*tolteca Mann. Ariz.; Mexico. Host: Pheidole sp. 

Orasema tolteca Mann, 1914. Psyche 21: 183-184. $, $. 



248 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Family PERILAMPIDAE 

Genus PERILAMPUS Latreille, pp. 517-519 

canadensis canadensis Crawford. Hosts: Ectemnius (Hypocrabro) stirpicola 
(Pack.), E. (H.) continuus (Fab.), E. ( H.) paucimaculatus 
(Pack.), Euplilis sp. 
Biology: Krombein, 1960. Ent. News 71: 32, 66.— Krombein, 1964. Biol. 
Soc. Wash. Proc. 77: 101-102. 

chrysopae Crawford. 

Taxonomy: Kerrich, 1958. Opusc. Ent. 23: 82. 

fulvicornis Ashmead, changed status. Raised from subsp. to sp. by Peck, 
1963, loc. cit. supra, p. 517. 
Biology: Schaffner, 1959, loc. cit. supra, p. 87. — Beique, 1960. Canad. 
Ent. 92: 859.— Bennett, 1961. Tenn. Acad. Sci. Jour. 36: 353.— 
Harman and Kulman, 1962. Jour. Econ. Ent. 55: 1007. 

fulvicornis prothoracicus Smulyan. Raised to sp. rank. 

hyalinus Say. [May represent a species complex.] 

Biology: Smith, 1958. Canad. Jour. Res. 36: 254-258.— Tripp, 1962. 

Canad. Ent. 94: 1250-1270.— Becker and Benjamin, 1964. Jour. 

Econ. Ent. 57: 356. 
Taxonomy: Finlayson, 1963. Canad. Ent. 95: 499-500, 501 (imm. 

stages). — Burks, 1964, loc. cit. supra, pp. 1259-1260 (aciculatus 

Provancher) . 

prothoracicus Smulyan, changed status. Raised from subsp. to sp. by Peck, 

1963, loc. cit. supra, p. 523. 

stygicus Provancher. Que. and Maine south to Va. west to Ohio, Kans. and 
La. 
Taxonomy: Burks, 1964, loc. cit. supra, p. 1260. 

Genus CHRYSOLAMPUS Spinola, p. 539 

Taxonomy: Boucek, 1956. Mus. Nat. Pragae Acta Faun. Ent. 1: 87-88. 

—Kerrich, 1958. Opusc. Ent. 33: 82. 
Transf . with component spp. from Pteromalidae, tribe Lamprotatini, and 
insert in Perilampidae following Perilampus Latreille, p. 519. 

Genus ELATUS Walker, p. 539 

*Parelatus Girault, 1916. Queensland Mus. Mem. 5: 226. 

Type : Elatus ater Girault. Orig. desig. 
Taxonomy: Boucek, 1956, loc. cit. supra, p. 88. 

Transf. with component spp. from Pteromalidae, tribe Lamprotatini, and 
insert in Perilampidae following Perilampus Latreille, p. 519. 

laevis (Provancher), n. comb. ? Ont. [may be foreign]. Transf. from Un- 
placed Species, p. 594, teste Burks, 1964, loc. cit. supra, p. 1260. 

Family AGAONIDAE 

Genus BLASTOPHAGA Gravenhorst 
( = B/asfophagus Gravenhorst of Catalog, p. 519) 

Nomenclature: China, 1960. Bui. Zool. Nomencl. 18: 69-72 (request for 
validation of Blastophaga Gravenhorst, 1829, and suppression of 
Blastophagxis Gravenhorst, 1827). — Peck, 1963, loc. cit. supra, p. 
526 (Blastophaga).— I.C.Z.'N., 1964. Opinion 694, Bui. Zool. No- 
mencl. 21: 31-32 (validation of Blastophaga Gravenhorst, type B. 
grossorum Grav.). 

psenes (Linnaeus). Cosmopolitan. 

Bibliography: Condit and Enderud, 1956. Hilgardia 25: 64-85, 616. 
Biology: Joseph, 1954. Agra Univ. Jour. Res. (Sci.) 3: 408. — Grandi, 



TORYMIDAE 249 

1955. Accad. Naz. dei Lincei Atti, Mem., CI. Sci., ser. 8, 5: 3. — 
Grandi, 1956. Bologna Univ. 1st. di Ent. Bol. 21: 111.— Mrak, 
1957. Food Tech. 11: 547.— Kuttamathiathu, 1958. Ann. Sci. 
Nat., Zool. et Biol. Anim., ser. 11, 20: 201. 
Nomenclature: I.C.Z.N., 1964. Opinion 694, Bui. Zool. Nomencl. 21: 32 
(validation of Cynips psenes Linnaeus, 1758). 

Family TORYMIDAE 

Subfamily IDARNINAE 

Genus COLYOSTICHUS Mayr, p. 519 

brevicaudis Mayr, corrected spelling of brevicauda of Catalog, teste Peck, 
1963, loc. cit. supra, p. 530. 

*Genus NEOSYCOPHILA Grandi 

( = Sycop/i//a of Catalog, p. 519) 

Neosycophila Grandi, 1923. Genova Mus. Civ. di Stor. Nat. Ann. 51: 106. 

Type : Neosycophila omeomorpha Grandi. Orig. desig. 
The genus Sycophila Walker is a member of the Eurytomidae, teste E. 
F. Riek, in litt. 

Subfamily TORYMINAE 

Genus ALLOTORYMUS Huber, p. 520 

splendens (Provancher) . 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1262. 

Genus TORYMUS Dalman, pp. 520-527 

baccharidis (Huber). Host: Rhopalomyia calif ornicaYeM. 

Biology: Doutt, 1961. Ent. Soc. Amer. Ann. 54: 51 (baccharicidis). 

bedeguaris (Linnaeus). Que., Ont., 111., Iowa, Nebr., Wash.; Europe. 
Biology: Judd, 1959. Canad. Ent. 91: 728-730. 

brevis Breland. Minn. 

califoniicus californicus ( Ashmead) . Delete Conn, and Fla. from dist. 

capillaceus (Huber). Tex., Ariz., Calif. Delete Conn, from dist. 

citripes (Huber). Host: Gall of Euaresta bullans (Wied.) on Atriplex. 

druparum Boheman. Syn. of varians (Walker). 

duplicatus (Huber). Que. and N. H., south to Fla. west to Nebr. 

fagopirum (Provancher), corrected spelling of fagopyrum of Catalog, teste 
Peck, 1963, loc. cit. supra, p. 541. 
Taxonomy: Burks, 1964, loc. cit. supra, p. 1256. 

flavicoxa (Osten Sacken). 

Cleonymus superbus Provancher, 1888 (syn. by Burks, 1964, loc. cit. 
supra, p. 1257). 

fullawayi (Huber). Host: Dryocosmus dubiosus (Full.). 

fulvus (Huber). Md., Iowa. 

koebelei (Huber). Host: Rhopalomyia calif oi^nica Felt. 
Biology: Doutt, 1961, loc. cit. supra, p. 51. 

longicauda (Provancher). 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1256. 

pachypsyllae (Ashmead). Ohio, Kans., Idaho. Host: Pachypsylla celtidis- 
mamma Riley. 



250 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

pilularidis (Huber). Host: Rhopalomyia calif ornica Felt. 

Biology: Doutt, 1961, loc. cit. supra, p. 51. — Burks, 1965. Ent. News 76: 
74-76. 

*rugglesi Milliron. Del., Md., Va. Host: Ilex opaca seeds. 

Torymus rugglesi Milliron, 1959. Ent. Soc. Amer. Ann. 52: 52. ?, $. 

scalaris (Huber). Kans. 

strobiloides (Huber). Ont. 

superbus (Provancher) . Syn. of flavicoxa (Osten Sacken). 

umbilicatus (Gahan). Md. south to Fla., west to Calif.; Mexico. 

*varians (Walker). N. C, Man.; Europe. 

Callimome various Walker, 1833. Ent. Mag. 1: 122. 9. 

Torymus druparum Boheman, 1834 (1833) (syn. by Olsson, 1957. 

Vaxtskyd. 21: 58). 
Taxonomy: Eady, 1958. Ent. Mo. Mag. 94: 260 (Syntomaspis Foerster). 

vesiculi Moser. Idaho. 

Genus DIOMORUS Walker, p. 527 

biorhizae Ashmead. Prob. a sp. of Torymus; type apparently lost. Transf. to 
Unplaced Species, p. 594. 

viridis (Provancher). 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1259. 

zabriskiei Cresson. Hosts: Crossocerus fergusoni Pate, Epicrossoceriis inso- 
lens (Fox), Ectemnius (Hypocrabro) spiniferus (Fox), E. (H.) 
paucimaculatus (Pack.) , Euplilis (Corynopus) rufigaster (Pack.), 
E. (C.) arapaho Pate. 
Biology: Krombein, 1960. Ent. News 71: 32, 67.— Krombein, 1964. Biol. 
Soc. Wash. Proc. 77: 102-103. 

Subfamily MONODONTOMERINAE 

Taxonomy: Szelenyi, 1957. Mus. Nat. Hung. Ann. Hist.-Nat., n.s., 8: 
381-388 (key to genera). — Szelenyi, 1960. Acad. Sci. Hung., Acta 
Zool. 5: 141-146 (rev. key to genera). 

Genus LIODONTOMERUS Gahan, p. 528 

insuetus Gahan. Nebr., Okla., Utah, Calif. Hosts: Bruchophagus kolobovae 
Fed., B. roddi (Guss.), B. platyptera (Walk.). 
Biology: Neunzig and Gyrisco, 1959. Jour. Econ. Ent. 52: 899-901. 

longfellowi (Girault). N. Y., Md., Va., W. Va., Ohio, Mich., Iowa, Utah. 
Hosts : Bruchophagus platyptera complex. 
Biology: Neunzig and Gyrisco, 1959, loc. cit. supra, pp. 899-901. 

perplexus Gahan. Holarctic. Hosts: Bruchophagus platyptera complex. 
Biology: Neunzig and Gyrisco, 1959, loc. cit. supra, pp. 899-901. 

Genus MONODONTOMERUS Westwood, pp. 528-529 

aereus Walker. Holarctic; India. 

Biology: Clausen, 1956, loc. cit. supra, pp. 114-116. — Wilbert, 1960. En- 
tomophaga 5: 203.— Blunck and Wilbert, 1962. Ztschr. f. Angew. 
Ent. 50: 203. 

dentipes (Dalman). Holarctic. 

Monodontomerus viridaeneus Provancher (syn. by Burks, 1964, loc. cit. 

supra, p. 1259). 
Biology: Finlayson and Finlayson, 1958. Canad. Ent. 90: 558-559. 
Taxonomy: Finlayson, 1960. Canad. Ent. 92: 37, 39, 44 (imm. stages). 

*mexicanus Gahan. Ind., Calif.; Mexico. Hosts: Trypoxylon mexicanum 
Sauss., Stenodynerus vagus (Sauss.). 



TORYMIDAE 251 

Monodontomerus mexicanus Gahan, 1941. U. S. Natl. Mus. Proc. 90: 
468. ?, $. 

*minor (Ratzeburg). Conn., N. Y. south to Va. west to Calif.; Europe. 

Hosts: Thyridopteryx ephemerae formis (Haw.), Phryganidia cali- 

fo7-nica Pack., Samia cyntliia (Drury) ; recorded as hyperparasite 

of Lepidoptera in European lit. 

Torymus minor Ratzeburg, 1848. Ichn. der Forstins., v. 2, p. 178. 9,5. 

Monodontomerus subobsoletus Gahan (syn. by Steffan, 1952. Paris Mus. 

d'Hist. Nat. Bui., ser. 2, 24: 293). 
Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, p. 102. 
montivagus Ashmead. Host: Megachile rotundata (Fab.). 

obscurus Westwood. Holarctic; Pa., Md., Ga., W. Va., Idaho. Host: Mega- 
chile rotundata (Fab.). 
Taxonomy: SteflFan, 1952, loc. cit. supra, pp. 289, 290.— Eady, 1958. Ent. 
Mo. Mag. 94: 269. 

subobsoletus Gahan. Syn. of minor (Ratzeburg). 

Genus ZAGLYPTONOTUS Crawford, pp. 529-530 
schwarzi Crawford. Idaho. 

Genus GLYPHOMERUS Foerster, p. 530 

Stigma (Fabricius). Nebr. Delete Monodontomerus viridaeneus Provancher 
from syn. and trans, to syn. under Monodontomerus dentipes 
(Dalman). 
Taxonomy: Burks, 1964, loc. cit. supra, p. 1259. 

Genus HYPERIMERU5 Girault, p. 537 

Transf. with included species from Pteromalidae, tribe Asaphini, and 
insert in Toryminae, following Glyphomerus Foerster, p. 530. 

corvus Girault, Hosts: Hemerobius sp., Sympherobius sp. 

Genus PSEUDOTORYMUS Masi, p. 530 

lazulellus (Ashmead) . N. B., Que., and Conn., west to Minn. 

Genus ANTISTROPHOPLEX Crawford, p. 531 

bicoloripes Crawford. Iowa. Host: Antistrophus pisum Walsh gall on Lygo- 

desmia. 
Biology: Brandhorst, 1964. Ent. Soc. Amer. Ann. 57: 74-77. 

Genus MICRODONTOMERUS Crawford, p. 531 

anthonomi (Crawford). Calif. Host: Leptoc hi lus S'p. 

fumipennis Crawford. Wyo. south to N. Mex., west to Wash., Oreg., Calif. 
Host: Malacosoma calif ornicum (Pack.). 
Biology: Langston, 1957. Univ. Calif. Pubs., Ent. 14: 29-30. 

Genus ROPTROCERUS Rafieburg, Sup. 1, pp. 73, 76 
(nPochyceros Ratzeburg of Catalog, p. 549) 

Taxonomy: Hedqvist, 1963, loc. cit. supra, p. 61. 

xylophagorum (Ratzeburg). Que. and N. H. south to N. C. and La., west to 

B. C, Wash., and Ariz. 
Biology: Reid, 1957. Canad. Ent. 89: 6-7. 
Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 61-62, 66-70 (eccoptogastri 

Ratzeburg a syn.). — Burks, 1964, loc. cit. supra, p. 1261 (rectus 

Provancher). 

Subfamily MEGASTIGMINAE 

Genus MEGASTIGMUS Dalman, pp. 531-533 
*Epimeg€istignius Girault, 1915, Queensland Mus, Mem. 4: 297, 307. 



252 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Type : Xanthosomoides fulvipes Girault. Orig. desig. 

Taxonomy: Milliron, 1949. Amer. Midland Nat. 41: 263. 

Biology: Lessmann, 1962. Ztschr. f. Angew. Ent. 49: 363-371 (imm. 

stages).— Lessmann, 1962. Ztschr. f. Angew. Ent. 50: 233-241 

(host plants). 

aculeatus aculeatus (Swederus). 111. 

Taxonomy and Biology: Balduf, 1957. Ent. Soc. Amer. Ann. 50: 

479-483.— Balduf, 1959. 111. Biol. Monog. 26, pp. 124-130. 
Nomenclature: Peck, 1963, loc. cit. supra, p. 578 (var.). 

aculeatus nigroflavus Hoflfmeyer. N. J., and Pa. south to Va., west to 111.; 
Japan, Russia. 
Nomenclature: Peck, 1963, loc. cit. supra, p. 579 (var.). 
Biology: Balduf, 1959, loc. cit. supra, pp. 124-126. 

albifrons Walker. 

Biology: Keen, 1958, loc. cit. supra, pp. 102-105. 

americanus Milliron. Cited as slossonae Crosby in Catalog in error; latter is 
a ywmen nudum, not validated through citation in synonymy. See 
Art. ll,d of Int. Code Zool. Nomencl. 

brevicaudis Ratzeburg, corrected spelling of brevicauda of Catalog. 

lasiocarpae Crosby. Alta., B. C, Wash. Delete Conn, from distb. 
Biology: Keen, 1958, loc. cit. supra, p. 105. 

milleri Milliron. 

Biology: Keen, 1958, loc. cit. supra, p. 106. 

nigrovariegatus Ashmead. 

Biology and Taxonomy: Balduf, 1957, loc. cit. supra, pp. 478-483. — Bal- 
duf, 1959, loc. cit. supra, pp. 130-145. 

picea montana Milliron. 

Biology: Keen, 1958, loc. cit. siipra, p. 106 (var.). 

picea picea Rohwer. 

Nomenclature: Peck, 1963, loc. cit. supra, p. 584 (var.). 
Biology: Keen, 1958, loc. cit. supra, p. 106. 

pinus pinus Parfitt. 

Nomenclature: Peck, 1963, loc. cit. supra, p. 585 (var.). 

Biology: Hussey and Klingler, 1954. Ent. Mo. Mag. 90: 198-201 {pinus 
Parfitt and var. crosbyi HofFmeyer). — Keen, 1958, loc. cit. supra, 
pp. 106-108.— Hussey, 1959. Ent. Mo. Mag. 95: 237. 

pinus marginatus Hoffmeyer. 

Nomenclature: Peck, 1963, loc. cit. supra, p. 586 (var.). 

rafni Hoffmeyer. 

Biology: Keen, 1958, loc. cit. supra, pp. 108-110. 

slossonae Crosby. Nomen nudum; see americanus Milliron. 

specularis Walley. 

Biology: Hedlin, 1957 (1956). Canad. Ent. 88: 691-697. 

spermotrophus nigrodorsatus Milliron. 

Biology: Keen, 1958, loc. cit. supra, p. Ill (var.). 

spemiotrophus spermotrophus Wachtl. 

Nomenclature: Peck, 1963, loc. cit. supra, p. 587 (var.). 
Biology: Hussey, 1955. Roy. Ent. Soc. London Trans. 106: 133-151. — 
Hedlin, 1958. Ent. Soc. B. C. Proc. 55: 37.— Keen, 1958, loc. cit. 
su2)ra, pp. 110-111. — Hussey, 1959. Ent. Mo. Mag. 95: 237. 

tsugae heterophyllae Milliron. 

Biology: Keen, 1958, loc. cit. supra, p. Ill (var.). 

tsugae tsugae Crosby. 

Nomenclature: Peck, 1963, loc. cit. supra, p. 588 (var.). 
Biology: Keen, 1958, loc. cit. supra, p. 111. 



ORMYRIDAE 253 

* Genus EUMEGASTIGMUS Hussey 

Eumegastigmus Hussey, 1956. Roy, Ent. Soc, London, Proc, ser. B, 25: 
157. 

Type: Eumegastigmus hypogea Hussey. Orig. desig. 

Insert in Megastigminae following Megastigmus Dalman, p. 533. 

*traiisvaalensis Hussey. Calif.; S. Africa, Canary Isls. Host: Seeds of 
Schinus molle. 
Eumegastigmus transvaalensis Hussey, 1956. Roy. Ent. Soc, London 

Proc, ser. B, 25: 161. 9, $. 
Biology: Harper and Lockwood, 1961. Calif. Dept. Agr. Bui. 50: 129. 

Subfamily PODAGRIONINAE 

Placed in Torymidae in Sup. 1, p. 82. 

Taxonomy: Ferriere, 1955. Soc. Roy. Ent. Belg. Mem. 27: 207 (Poda- 
grionidae). 

Genus PODAGRfON Spinola, p. 583 

*Cyanostola Saussure in Grandidier, 1890. Hist. Madagascar 20, Hym. 
1, pi. 15, fig. 30. 

Type : Cyanostola coerulea Saussure. By monotypy. 
*Podagrionella Girault, 1913. Arch. f. Naturgesch. 79 (A) pt. 6: 96. 

Type : Podagrionella bella Girault. By monotypy. 
*Pachytomoidella Girault, 1913. Wis. Nat. Hist. Soc. Bui. 11: 40. ^ 

Type : Pachytomoidella magniclavus Girault. Orig. desig. 
*Propodagrion Girault, 1915. Queensland Mus. Mem. 4: 287. 

Type: Propodagrion worcesteri Girault. Orig. desig. 
* Propachytomoides Girault, 1917. Ins. Ins. Mens. 5: 152. 

Type: Propachytomoides semialbiclavus Girault. Orig. desig. 
Taxonomy: Ferriere, 1955, loc. cit. supra, p. 216. 

*brasiliense Howard. Fla. ; West Indies. Host: Mantid egg mass. 

Podagrion brasiliensis Howard, 1894. Linn. Soc. London, Jour., Zool. 25: 

83 9 
Taxonomy: Gahan, 1951. Canad. Ent. 83: 176. 

crassiclava Gahan. Fla. ; Mexico. 

^mantidiphaguin Girault. Ariz.; Mexico, West Indies. Host: Egg mass of 
undet. mantid. 
Podagrion mantidiphagum Girault, 1917. Ent. 50: 37. 9, $. 

mantis Ashmead. Calif.; Mexico, Hawaii. 

Biology: Gurney, 1959. Ent. Soc. Wash. Proc. 61: 23. 

Family ORMYRIDAE 

Genus ORMYRUS Westwood, pp. 533-534 

*Wania Risbec, 1951. Inst. Franc. d'Afr. Noire M6m. 13: 294. 

Type: Wania ornata Risbec. By monotypy. 
Taxonomy: Peck, 1963, loc. cit. supra, p. 591. 

brunneipes Provancher, May 1887. Que. and Ont. south to Fla. west to Mich, 
and 111. Hosts: Listed under ventricosus Ashmead in Catalog, p. 
534. 
Ormyrus ventricosus Ashmead, Nov. 1887 (syn. by Burks, loc. cit. supra, 
p. 1259). 

vacciniicola Ashmead. 

Biology: Judd, 1959. Amer. Micro. Soc. Trans. 78: 218-214. 

ventricosus Ashmead. Syn. of bmnneipes Provancher. 



254 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Family PTEROMALIDAE 

Subfamily SPHEGIGASTERINAE 

Genus SPALANGIA Latreille, p. 535 
Revision: BouSek, 1963. Mus. Nat. Pragae Acta Ent. 35: 429-512. 

drosophilae Ashmead. N., Cent., and S. A-.merica, West Indies. Hosts: Mero- 
myza americana Fh., Oscinella frit (L.), O. minor (Ad.), O. car- 
bonaria (Loew), O. soror Macq., Hexacola sp., Loxotropa sp., 
Pseudogaurax anchora (Loew). 
Biology: Simmonds, 1952. Bui. Ent. Res. 43: 533. — Simmonds, 1953. 
Bui. Ent. Res. 44: 773.— Simmonds, 1954. Bui, Ent. Res. 45: 527. 
—Simmonds, 1956. Bui. Ent. Res. 47: 361-376. 

*endius Walker. Cosmopolitan. Hosts: Clirysoviyia spp., Fannia leucosticta 
(Meig.), F, scalaris (F.), Haematobia exigua Meig., H. irritans 
(L.), Musca domestica L., Stonioxys calcitrans (L.), Sarcophaga 
spp., Opiiyra anthrax Meig., Ceratitis capitata (Wied.). Dacus 
dorsalis Hend., D. ferrugineiis (F.). 

Spalangia endius Walker, 1859. Monog. Chalc, v. 2, p. 96. $. 

Spalangia muscidarutn var. stotiioxysiae Girault, 1916 (syn. by Boucek, 
19G3, loc. cit. supra, p. 458). 

*Spalangia philippinensis Fullaway, 1917. Hawaii Ent. Soc. Proc. 3: 
292. 9, $. (Syn. by Boucek, 1963, loc. cit. supra, p. 458.) 

Spalangia muscidaruin var. texensis Girault, 1920 (syn. by Boucek, 1963, 
loc. cit. supra, p. 458). 

*erylhromera brachyceps Boucek. ? Conn., ? Ont., ? Sask. ; Europe. [N. 
Amer. specimens ref. to this subsp. with hesitation.] 
Spalangia erythromera brachyceps Boucek, 1963. Mus. Nat. Pragae Acta 
Ent. 35: 471-472. 9, $. 

haematobiae Ashmead. Ont. and Alta. south to Va. and Tex. 

Taxonomy: Boucek, 1963, loc. cit. supra, pp. 475-476 {S. impuncta Ho- 
ward a possible syn.). 

muscae Howard. Syn. of nigra Latreille. 

muscarum Girault. Syn. of nigripes Curtis. 

muscidarum Richardson. Syn. of nigroaenea Curtis. 

*nigra Latreille. N. Amer.; Europe, West Indies, Hawaii. Hosts: Musca 
domestica L., Haematobia irritans (L.), Stomoxys calcitrans (L.), 
Phorbia antiqua (Meig.), Dacus cucurbitae Coq., Rhagoletis com- 
pleta Cress. 

Spalangia nigra Latreille, 1805. Hist. Nat. Crust. Ins., v. 13, p. 288. 

*Spalangia hirta Haliday, 1833. Ent. Mag. 1: 334. ?. (Syn. by Boucek, 
1963, loc. cit. supra, p. 443.) 

Spalangia rugosicollis Ashmead, 1894 (Syn. by Boucek, 1963, loc. cit. 
supra, p. 443). 

Spalangia muscae Howard, 1911 (syn. by Boucek, 1963, loc. cit. supra, p. 
443). 

*iiigripes Curtis. Holarctic: N. America, Europe, Asia. Hosts: Lucilia sp., 
Fannia leucosticta (Meig.), F. scalaris (F.), Musca domestica 
domestica L., M. d. vicina Macq.. undet. muscids. 

Spalangia nigripes Curtis, 1839. Brit. Ent., v. 16 [3], fol. 740 [p. 2]. 9. 

*Spalangia hyaloptera Foerster, 1850. Naturhist. Ver. der Preuss. 
Rheinl. Verhandl. 7: 509. 9, $. (Syn by Boucek, 1963, loc. cit. 
supra, p. 461.) 

*Spalangia formicaria Kieffer, 1905. Berlin Ent. Ztschr., 50: 1. 9. 
(Syn. by Boucek, 1963, loc. cit. supra, p. 461.) 

Spalangia muscarum Girault, 1920 (syn. by Boucek, 1963, loc. cit. supra, 
p. 461). 



PTEROMALIDAE 255 

*nigroaenea Curtis. Cosmopolitan. Hosts: Musca doviestica domestica L., M. 

d. vicina Macq., Musca spp., Haematobia exigua Meig., Stomoxys 

calcitrans (L.), SarcopJiaga spp., Chrysomyia spp., Lucilia spp., 

Paratheresia claripalpis (v. d. W.). 

Spalangia nigroaenea Curtis, 1839. Brit. Ent., V. 16 [3], fol. 740 [p. 2]. 

$. 
Spalangia hontalaspis Foerster, 1850. Naturhist. Ver. der Preuss. 

Rheinl. Verhandl 7: 505. 9, $, (Syn. by Boucek, 1963, loc. cit. 

supra, p. 448.) 
* Spalangia astuta Foerster, 1851, ibid. 8:1. 9. (Syn. by Boucke, 1963. 

loc. cit. stipra, p. 448.) 
Spalangia muscidaruw Richardson. 1913 (syn. by Boucek, 1963, loc. cit. 

supra, p. 448). 

rugosicollis Ashmead. Syn. of nigra Latreille. 
stomoxysiae Girault. Syn. of endius Walker. 

*Genus CEROCEPHALA Westwood 

Cerocephala Westwood, 1832. Mag. Zool. 2 : CI. ix, pi. 4. 

Type : Cerocephala cornigera Westwood. By monotypy. 
*Epimacrus Walker, 1833. Ent. Mag. 1: 368. 

Type : Epimacrus rufus Walker. By monotypy. 
*Sciatheras Ratzeburg, 1848. Ichn. der Forstins., v. 2, p. 209. 

Type : Sciatheras trichotus Ratzeburg. By monotypy. 
*Parasciatheras Masi, 1917. Nov. Zool. 24: 189. 

Type: Cerocephala caelebs Masi. By monotypy. 
*Proamotura Girault, 1920. Ins. Ins. Mens. 8: 143. 

Type: Proamotura aquila Girault. By monotypy. 
Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 127-128. 

Insert in tribe Cerocephalini following Choetospila Westwood, p. 535. 

*clubarae Wallace. Pa. Host: TProh. Hadrobregmus carinatus (Say). 

Cerocephala dubarae Wallace, 1959. Brooklyn Ent. Soc. Bui. 54: 84. 9, 

$• 

Genus ACEROCEPHALA Gahan, p. 536 

atroviolacea (Crawford). Colo., Calif. Host: ConophtJiorus flexilis Hopk. 
Biology: Keen, 1958, loc. cit. supra, pp. 49, 98. 

Genus ASAPHES Walker, p. 536 

All species of Asaphes are believed to be secondary parasites of Aphidi- 
dae. 

american MS Girault. Syn. of luceiis ( Provancher) . 

californicus Girault. Colo., Wash., Oreg. 

fletcheri (Crawford). Syn. of lucens (Provancher). 

lucens (Provancher). N. B., Que., Ont., Maine south to N. C, west to Iowa 
and Colo. Hosts: Aphidius testaceipes (Cress.), A. phorodontis 
Ashm., A. polygonaphis (Fh.), Diaretiella rapae (M'Int.), Praon 
aguti Smith, P. simulans (Prov.). 

Megorismus fletcheri Crawford (syn. by Burks, 1964, loc. cit. supra, p. 
1258). 

Asaphes americanus Girault (syn. by Burks, 1964, loc. cit. supra, p. 
1258). 

Biology: Sekhar, 1958. Ent. Soc. Amer. Ann. 51: 1-8 {fletcheri). 

Genus Hyperimerus Girault, p. 537 

Transf. with included sp. to Torymidae, subfamily Monodontomerinae, 
following Glyphomerus Foerster, p. 530. 

Genus DIPARA Walker, p. 537 
*Tricoryphus Foerster, 1856. Hym. Stud., v. 2, p. 46. 



256 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Type: Tricoryphus fasciatus Thomson. Included by Thomson, 
1876. 
*Hispanolelaps Mercet, 1927. Eos 3 : 60. 

Type : Hispanolelaps coxalis Mercet. Orig. desig. 
Taxonomy: Boucek, 1954. Mus. Nat. Pragae Acta Ent. 29: 52, 54. 
The female of this genus is wingless, commonly taken in Berlese 
samples; the male is winged. 

Genus APTEROLELAPS Ashmead, p. 537 

Taxonomy: Delucchi, 1958. Bol. Zool. Agr. Bachic, ser. 2, 1: 56 (syn. of 
Trichoryphus [!] Foerster). 

Genus LAMPROTATUS Westwood, pp. 537-538 

Delete Miscogaster Walker from syn. 

Taxonomy: Delucchi, 1955. Lunds Univ. Aarsskr., n. s., 50 (20) : 6-10. — • 
Peck, 1963, loc. cit. supra, pp. 606-608. 

abnormicolor (Girault). Transf. to Miscogaster Walker. 

auratus (Ashmead). Transf. to Miscogaster Walker. 

biguttus (Girault). Transf. to Miscogaster Walker. 

diaeus Walker. Maine, Minn. 

discoloripes (Girault). Transf . to Miscogaster Walker. 

flora (Girault). Transf . to Miscogaster Walker. 

keatsi (Girault). Transf. to Miscogaster Walker. 

marylandicus (Girault). Transf . to Miscogaster Walker. 

unguttus (Girault). Transf. to Miscogaster Walker. 

Genus MISCOGASTER Walker 
(= Lamprotafus Westwood of Catalog, in part, pp. 537—538) 

Delete from syn. under Lamprotatns Westwood and transf. with follow- 
ing spp. to p. 538, following Lamproiatus. 

Taxonomy: Delucchi, 1955, loc. cit. supra, pp. 6-7, 55-56. — Peck, 1963, 

loc. cit. supra, pp. 608-609 (transf. of spp. from Lamprotatus 

Westwood to Miscogaster Walker) . 

abnormicolor Girault. Transf. from Lamprotatus Westwood. 
auratus (Ashmead). Pa., Mich., Wis., Nebr. 
biguttata Girault. Transf. from Lamprotatus Westwood. 
discoloripes Girault. Transf. from Lamprotatus Westwood. 
flora Girault. Transf. from Lamprotatus Westwood. 
keatsi Girault. Transf. from Lamprotatus Westwood. 
marylandica Girault. Transf. from Lamprotatus Westwood. 
ungutta Girault. Transf. from Lamprotatus Westwood. 

Genus SYNTOMOPUS Walker, p. 538 
americanus Ashmead. N. Y. south to Fla., west to Oreg. and Calif. ; Mexico. 

Genus Chrysofampus Spinola, p. 539 

Transf. with included species to Perilampidae, following Perilampus La- 
treille, p. 519. 

Genus Elatus Walker, p. 539 

Transf. with included species to Perilampidae, following Perilampus La- 
treille, p. 519. 



PTEROMALIDAE 257 

Genus PACHYNEURON Walker, pp. 539-541 

*Serimus Brethes, 1913. Buenos Aires Mus. Nac. de Hist. Nat. Ann. 24: 
90. 

Type: Serimus argentinus Brethes. Orig. desig. 
Taxonomy: De Santis, 1957. Univ. Nac. de La Plata, Notas del Mus. 19: 

118. 
*Neopachyneuron Girault, 1917. Descr. Hym. Chalc. Var. cum Obs., pt. 
5, p. 9. 

Type: Pachyneuron eros Girault. By monotypy. 
Taxonomy: Peck, 1963, loc. cit. supra, p. 612. 

altiscutum Howard, ch. to Cook, teste Peck, 1963, loc. cit. supra, p. 613. N. B., 
D. C, S. C, Colo. Hosts: Cremifania nigricellulata Czerny, 
Leucopis sp., Leucopina americana Mall., Neoleucopis obscura 
(Hal.), N. pinicola Mall. 
Biology: Brown and Clark, 1957 (1956). Canad. Ent. 88: 679-682. 

anthomyiae Howard. Host: Leucopis sp. 

eros Girault. B. C. 

mucronatum Girault. Tex., Ariz.; Mexico. Host: Assoc, with coccids on 
Opuntia. 

virginicum Girault. Maine, 111. 

Genus CORUNA Walker, Sup. 1, p. 74 

clavata Walker. B. C, Alaska; Europe. 

Biology: Stary, 1962. Zool. Listy 11: 267. 

Genus PACHYCREPOIDEUS Ashmead, p. 541 

* Anisopteromalia Boucek, 1954. Mus. Nat. Pragae, Acta Ent. 29: 57-60. 

Type: Anisopteromalia crassinervis Boucek. Orig. desig. 
Taxonomy: Boucek, 1957 (1956). Casopis 53: 165. 

vindemiae (Rondani), corrected spelling of vindemmiae of Sup. 1, p. 75, teste 
Peck, 1963, loc. cit. supra, p. 620. Hosts: Fannia canicularis (L.), 
F. scalaris (Fab.), Calliphora vicina R.-D., Phorviia regina 
(Meig.). Lucilia sericata (Meig.), Anastrepha sp. 
Biology: Nostvik, 1954. Oikos 5: 195.— Steve, 1959. Jour. Econ. Ent. 52: 
530. 

Genus EUNEURA Walker, p. 541 

lachni (Ashmead). N. B. and Maine south to Fla., west to B. C. and Calif. 
Host: Paxiesia ponderosae (Mues.) on Cinara sp. 

Genus HALTICOPTERA Spinola, pp. 541-542 

*Pachylarthrus Westwood, 1832. Phil. Mag., ser. 3, 1 : 127. 

Type: Pachylarthrus insignia Westwood. By monotypy. 
Taxonomy: Nikolskaja, 1952, loc. cit. supra, p. 249. 

aenea (Walker). Syn. of patellana Dalman. 

brodiei Ashmead. Not Halticoptera Spinola; transf. to Unplaced Species, p. 

594. 

*patellana (Dalman). Holarctic. 

Diplolepis patellana Dalman, 1818. K. Svenska Vetensk-Akad. Handl. 

39: 80. 9. 
Dicyclus aenea Walker, 1833 (syn. by Ferriere, 1952. Soc. Ven. di Stor. 

Nat. Bol. 6: 172). 
Other syn., and hosts, under aenea in Catalog, p. 542. 

rosae Burks. Maine, Ont., Sask., Utah, Alaska. 

Biology: Balduf, 1959. HI. Biol. Monog. 26: 41-50, 170-173. 

Stella Girault. Ariz. Host: Aciurina maculata (Cole) . 



258 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Genus SPHEGIGASTER Spinola 
Iz^Trigonogastra Ashmead of Catalog, p. 542) 

Sphegigaster Spinola, 1811. Paris Mus. Hist. Nat. Ann. 17: 149. 

Type: Diplolepis pallicornis Spinola. Desig. by Ashmead, 
1904. 
Taxonomy: Delucchi, 1958. Bol. Zool. Agr. e Bach., ser. 2, 1: 56. 

*flavicomis (Walker). B.C.; Europe. Host: Phytomyza ilicis Curtis. 
Merismus flavicornis Walker, 1833. Ent. Mag. 1: 377. $. 
Biology: Cameron, 1939. Bui. Ent. Res. 30: 178-180, 192-197. 

Genus CYRTOGASTER Walker, p. 542 
glasgowi Crawford. Mo., Tex., Calif. 
vulgaris Walker. Wash. ; Europe. Host : Phytomyza nigra Meig. 

«Genus CAPELLIA Delucchi 
{zzEurYd'inota Foerster of Catalog, p. 543) 

Capellia Delucchi, 1958. Bol. Zool. Agr. e Bach., ser. 2, 1 : 59. 
Type: Eurydinota rufiventris Girault. Orig. Desig. 
Taxonomy: Delucchi, 1958, loc. cit. supra, pp. 57-60. 

lividicorpus (Girault). Mass., B. C. Host: Coleophora laricella Hbn. 
Biology: Schaffner, 1959, loc. cit. supra, p. 88. 

rufiventris (Girault). Miss., Minn., Calif. Host: Retinodiplosis sp. on pine. 

Genus HETEROSCHEMA Gahan, p. 543 

punctata (Ashmead). Miss., Mo.; West Indies. Host: Ophiomyia sp. in 
Lippa seed heads. 

rugosopunctata (Ashmead). Tex. 

Genus LOPHOCOMODIA Ashmead 

Transf. from Unplaced Genera, p. 593, to Sphegigasterini following He- 
teroschema Gahan, p. 543. 

americana Ashmead. Mich., Kans. Transf. from Unplaced Species, Sup. 1, p. 
84, teste G. E. Wallace in litt. 

Subfamily PTEROMALINAE 

Genus MORANILA Cameron, Sup. 1, p. 75 
(=:romocera Howard of Catalog, p. 543) 

calif ornica (Howard). Calif.; Mexico, Hawaii, South Sea Isls., Australia, 

Biology: Flanders, 1958. Jour. Econ. Ent. 51: 247. 

Genus EUNOTUS Walker, p. 544 

*crelaceus Walker. N. H., N. Y.; Europe. Host: Eriopeltis festucae Fonsc. 
Eunotus cretaceus Walker, 1834. Ent. Mag. 2: 298. $. 

Genus SCUTELLISTA Motschulsky, p. 544 

cyanea Motschulsky. Cosmopolitan. 

Biology: Clausen, 1956, loc. cit. supra, pp. 126, 128-129.— Flanders, 1958. 
Jour. Econ. Ent. 51: 247. — Flanders, 1959. Jour. Econ. Ent. 52: 
596-597.— Gexho, 1960. Tirana U. Shtet. Bui. Ser. Shken. Nat. 14: 
55.— Talhouk, 1961. Entomophaga 6 : 208. 

*Genus MACROGLENES Westwood 

Macroglenes Westwood, 1832. Phil. Mag., ser. 3, 1: 127. 

Type: Macroglenes oculatus Westwood. By monotypy. 
*Stenophru8 Foerster, 1841. Beitr. Monog. Pterom., p. 40. 

Type: Stenophrus compressus Foerster. By monotypy. 



PTEROMALIDAE 259 

Taxonomy: Gahan and Fagan, 1923. U. S. Natl. Mus. Bui. 124, p. 136. 

Insert in tribe Pirenini, before Pirene Haliday, p. 544. 

*penetrans (Kirby). N. J., Ont., Mich., Oreg. ; Europe. Host: Sitodiplosis 
mosellana (Gehin). 
Ichneumon penetrans Kirby, 1800. Linn. Soc. London Trans. 5: 109. 
*Macroglenes oculatus Westwood, 1832. Phil. Mag., ser. 3, 1: 127. 
*Stenophrus conipressus Foerster, 1841. Beitr. Monog. Pterom., p. 40. 
Distribution : Peck, 1963, loc. cit. stipra, p. 637. 

Genus PIRENE Holiday, p. 544 

marylandensis Girault. Transf . to Bairamlia Waterston, infra. 

*Genus BAIRAMLIA Waterston 

Bairamlia Waterston, 1929. Parasitol. 21 : 103. 

Type: Bairamlia fuscipes Waterston. Orig. desig. 

Insert in Tribe Pirenini following Pirene Haliday, p. 544. 

marylandensis (Girault). Transf. from Pirene Haliday, teste Waterston, 
1929, loc. cit. supra, p. 103. 

*Genus HERBERTIA Howard 

Herbertia Howard, 1894. Linn. Soc. London, Jour., Zool. 25 : 98. 
Type : Herbertia lucens Howard. By monotypy. 

Insert in tribe Pirenini, following Spathopus Ashmead, p. 544. 
Revision: Burks, 1959. Ent. Soc. Wash, Proc. 61: 249-255. 

*wallacei Burks. Pa., Md., Va., Ohio, Tex. Host: Phytomyza ilicicola Loew. 
Herbertia wallacei Burks, 1959. Ent. Soc. Wash. Proc. 61: 254. 9, $. 

Genus MORODORA Gahan, p. 544 
armata Gahan. B. C. 

Genus HEMADAS Crawford, p. 545 

nubilipennis (Ashmead). 

Biology: Judd, 1959. Amer. Micr. Soc. Trans. 78: 212-214. 

Genus Tr/gonoderos Westwood, p. 545 
Transf. with included spp, to tribe Cleonymini, p. 566. 

*Genus MACROMESUS Walker 

Macromesus Walker, 1848. List Hym. Brit. Mus., Chalc, pt. 2, pp. 106, 
101. 

Type: Macromesus amphiretus Walker. By monotypy. 
*Wesenbergia Kryger, 1943. Ent. Medd. 22: 360. 

Type: Wesenbergia occulata Kryger. Orig. desig. 
Taxonomy: Graham, 1959. Ent. Mo. Mag. 95: 73-78 (Macromesinae of 
Pteromalidae).— Hedqvist, 1960. Ent. Tidskr. 81: 140.— Boucek, 
1961. Mus. Nat. Pragae Acta Ent. 34: 57 (near Tridymini). — 
Hedqvist, 1963, loc. cit. supra, pp. 57-58 (Cheiropachinae) . 

Insert in tribe Tridymini after Elachertodomyia Girault, p. 546. 

*ainericanus Hedqvist. Ariz., Oreg., Calif. Hosts: Cylindrocopturus longulus 
(Lee), C. eatoni Buc, Pseudohylesinus nebulosus (Lee), Scolytus 
praeceps Lee, S. unispinosus Lee, S. ventralis Lee 
Macromesus americanus Hedqvist, 1960. Ent. Tidskr. 81: 140. 9, $. 

Genus TRIDYMUS Ratzeburg, p. 546 

aphidis Girault. Ark. 

gargantua Girault. N. H., Mich., Minn. Host: Reared from gall on willow. 

nigriclavus Girault, corrected spelling of nigriclava of Catalog, teste Peck, 
1963, loc. cit. supra, p. 643. 



260 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

oblongus (Provancher). Transf. from Unplaced Species, p. 594, teste Burks, 
1964, loc. cit. supra, p. 1261. 

Genus SYSTASIS Walker, p. 547 

diplosidis Eckel. Fla., Ohio, Wis., Miss. 

*encyrtoides Walker. Wis., Oreg., Alaska; Europe. Host: Contarinia sp. 
Systasis encyrtoides Walker, 1834. Ent. Mag. 2: 296. 9, $. 
Biology: Parnell, 1963. Roy. Ent. Soc, London Trans. 115: 271-272. 

Genus TEROBIA Foerster, p. 547 

Taxonomy: Delucchi, 1955. Soc. Roy. Ent. Belg. Mem. 27: 175 (Ormoc- 
erus Walker). — Peck, 1963, loc. cit. supra, p. 903 (N. Amer. spp. 
listed as unplaced) . 

Tribe MERISINI 

Taxonomy: Szelenyi, 1956. Mus. Nat. Hung., Ann. Hist.-Nat., n. s., 7: 
167-180. 

Genus MERISUS Walker, pp. 547-548 

Taxonomy: Szelenyi, 1956, loc. cit. supra, pp. 169-170. — Delucchi, 1956. 
Ztschr. f . Angew. Ent. 39 : 229. 

cupraeus (Provancher). Transf. to Caenacis Foerster in tribe Pteromalini, 

infra. 

destructor (Say). Holarctic. 

Taxonomy: Szelenyi, 1956, loc. cit. supra, p. 170 {Homoporus Thomson). 
— Delucchi, 1957. Ztschr. f. Angew. Ent. 40: 418-419 {Homoporus 
subg. Phaenacra Foerster) . 

febriculosus Girault. P. E. I. 

Taxonomy: Szelenyi, 1956, loc. cit. supra, p. 168 (? syn. of gibbiscuta 

Thomson; ref. to Homoporus Thomson). 
Biology: Cannon, 1963. Canad. Ent. 95: 34. 

mordellistenae Crawford. Iowa, Calif. 

*Genus HOMOPORUS Thomson 
( = A1er/soporus Mosi of Catalog, p. 548) 

Homoporus Thomson, 1878. Hym. Scand., v. 5, pp. 60, 64. 

Type: Pteromalus fulviventris Walker. Desig. by Ashmead, 
1904. 
[Not Homoporus Thomson, 1890, in Ichneumonidae, now considered a 

lapsus for Homotropus Thomson.] 
*Phaenacra Foerster, 1878. Naturhist. Ver. der Preuss. Rheinl. Ver- 
handl. 35: 51. 

Type: Phaenacra nubigera Foerster. Orig. desig. 
Taxonomy: Delucchi, 1957, loc. cit. supra, p. 402 (Homoporus subg. 
Phaenacra Foerster). — Erdos, 1953. Acad. Sci. Hung. Acta Biol. 
4: 239 (syn. of Phaenacra Foerster). — Peck, 1963, loc. cit. supra, 
p. 650 {Merisoporus Masi). 

chalcidiphagus (Walsh and Riley) . Holarctic. 

Taxonomy: Erdos,, 1953, loc. cit. supra, p. 239. — Szelenyi, 1956, loc. cit. 

supra, p. 168. — Delucchi, 1956, loc. cit. supra, pp. 417-418. 
Biology: Cannon, 1963, loc. cit. supra, p. 34. 

Genus CALLITULA Splnola, p. 548 

Taxonomy: Szelenyi, 1956, loc. cit. supra, pp. 167, 169-170. 

bicolor Spinola. Holarctic. 

Biology: Simmonds, 1952. Bui. Ent. Res. 43: 528-533.— Simmonds, 1953. 
Bui. Ent. Res. 44: 387-392. — Hemer, 1959. Ztschr. f. Angew. 
Ent. 44: 339. 



PTEROMALIDAE 261 

illinoensis (Ashmead). Md., Va., Fla., Tenn. 

Genus PSEUDOMICROMELUS Gahan and Fagan, p. 553 

deplanatus (Nees). Transf. to Cyclogastrella Bukovskij, infra. 

*Genus CYCLOGASTRELLA Bukovskij 

Cyclogastrella Bukovskij, 1938. Ent. Obozr. (Rev. Ent.) 27: 153. 

Type: Cyclogastrella quercina Bukovskij. By monotypy. 
Taxonomy: Graham, 1956. Ent. Mo. Mag. 92: 258. 

Insert in tribe Merisini following Callitula Spinola, p. 548. 

deplanata (Nees). Holarctic. Transf. from Pseudoniicromelus Gahan and 
Fagan, p. 553, teste Graham, 1956, loc. cit. supra, p. 259. 
Biology: Delucchi, 1962. Awamia 2: 123. 

Genus RHAPHITELUS Walker, pp. 548-549 

*Eucerchysius Brethes, 1913. Buenos Aires Mus. Nac. de Hist. Nat. 
Anal. 24: 103. 

Type: Eucerchysius scolyti Brethes. By monotypy. 
Taxonomy: De Santis, 1952. Soc. Ent. Argentina Rev. 15: 270 (generic 
syn.). — Hedqvist, 1963, loc. cit. supra, pp. 93-94. 

niaculatus Walker. Que., N. H., Pa., Wash., Oreg. ; Europe. 

Biology: Jamnicky, 1959. Biologia, Bratislava 14: 817. — Szczepanski, 

1960. Polskie Pismo Ent. 30: 409. 
Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 94-96. 

Genus METACOLUS Foerster, p. 549 

*Pterosema Foerster, 1878. Naturh. Ver. Preuss. Rheinl. Verh. 35 : 44. 

Type: Pterosema varicolor Foerster. Orig. desig. 
Taxonomy: Boucek, 1957. Mus. Nat. Prag. Acta Faun. Ent. 2: 76 (gen- 
eric syn.). — Hedqvist, 1963, loc. cit. supra, p. 97. 

fasciatus Girault. Idaho. Hosts: Assoc, with Scolytidae in Pinus spp. and 
Juniperus sp. 

Genus DINOTISCUS Ghesquiere, p. 549 
Taxonomy: Hedqvist, 1963, loc. cit. supra, p. 83. 
agrili (Rohwer). Transf. to Megairydynius Girault, p. 566. 

Genus TOMICOBIA Ashmead, p. 566 

Revision: Hedqvist, 1959. Opusc. Ent. 24: 177-184. 
Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 109-110. 

peratripes (Girault). 

Taxonomy: Hedqvist, 1959, loc. cit. supra, p. 184. 

tibialis Ashmead. Maine south to N. C., west to Idaho and Calif. Hosts: 
Adult Scolytidae. 
Taxonomy: Hedqvist, 1959, loc. cit. supra, pp. 178-181. 
Biology: Raid, 1957. Canad. Ent. 89: 7-8.— Ruckes, 1963. Pan-Pacific 
Ent. 39: 45. 

Genus RHOPALICUS Foerster, pp. 549-550 

Taxonomy: Hedqvist, 1963, loc. cit. supra, p. 71. 

pallipes Provancher. Transf. to Unplaced Species, p. 594. 

pulchripennis (Crawford). Que. and Maine south to Md. and Miss., west to 
Wash., Oreg., Calif. Hosts: Pissodes terminalis Hopping, P. ap- 
proximatus HoiDkins. 
Biology: Reid, 1957, loc. cit. supra, p. 7. — Keen, 1958, loc. cit. supra, p. 
100. 

tutela (Walker). Que., Mass., N. Y., Pa., Ont., Mich.; Europe. 

Biology: Nuorteva, 1956. Acta Ent. Fenn. 13: 64. — Chararas, 1959. Soc 



262 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Ent. France Bui. 64: 11. 
Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 71-79. 

Genus AMBLYMERUS Walker, p. 550 

Taxonomy: Graham, 1957. Ent. Mo. Mag. 93: 223 {Platymesopus West- 
wood).— von Rosen, 1958. Opusc. Ent. 23: 205 (Mesopolobus 
Westwood).— von Rosen, 1960. Ent. Tidskr. 82: 11-13 (Mesopo- 
lobus Westwood, in part) . 

bruchophagi (Gahan). Holarctic; N. Y. west to Man., Oreg., Calif. Hosts: 
Bruchophagus spp. 
Biology: Butler and Hansen, 1958. Pan-Pacific Ent. 34: 228. — Neunzig 
and Gyrisco, 1959. Jour. Econ. Ent. 52: 899. 

confusus (Ashmead). Idaho, Utah. Host: Tomicobia tibialis Ashm. par. on 
Ips oregonis (Eich.). 

flavipes (Walker). [Prob. not North American.] 

Taxonomy: von Rosen, 1958. Opusc. Ent. 23: 212.— Peck, 1963, loc. cit. 
supra, p. 660. 

mayetiolae ( Gahan ) . 

Taxonomy: von Rosen, 1958, loc. cit. supra, p. 207 (Mesopolobus). 

salicis (Girault). Mo. 

verditer (Norton). N. B., Que., and Maine south to N. J. and Pa., west to 
B. C. and Oreg. Hosts: Choristoncura pinus Free., Dioryctria 
disclusa Hein., Epinotia nanana (Treit.), Neocliprion lecontei 
(Fitch) ; may be pri. or sec. par. 

Biology: Keen, 1958, loc. cit. supra, pp. 98-99. — Schaffner, 1959, loc. cit. 
supra, p. 88. 

Taxonomy: von Rosen, 1958, loc. cit. supra, pp. 223-225 (Mesopolobus). 
— Finlayson, 1960. Canad. Ent. 92: 934-936, 939 (imm. stages). 

')'Genus AHLBERGiELLA von Rosen 

Ahlbergiella von Rosen, 1955. Ent. Tidskr. 76: 88. 

Type : Eutelus aequus Walker. Orig. desig. 

Insert in tribe Pteromalini following Amblymerus Walker, p. 550. 

*aequa (Walker). Nebr. [det. H. von Rosen]; Europe. Host: Delphacodes 
pelliicida (Fab.), eggs. 

Eutelus aequus Walker, 1834. Ent. Mag. 2: 364. 9. 

Biology and Taxonomy: von Rosen, 1955. Ent. Tidskr. 76: 88. — von Ro- 
sen, 1956. K. Lantbr. Ann. 23: 20-39.— von Rosen, 1958. Opusc. 
Ent. 23: 230-231 {Mesopolobus Westwood). 

Taxonomy: Graham, 1957. Ent. Mo. Mag. 93: 222. 

Biology: Raatikainen, 1962. Ann. Agr. Fenn. 1: 220 (Mesopolobus). 

Genus ZACALOCHLORA Crawford, pp. 550-551 

inilleri Crawford. Colo., Idaho, Oreg. Hosts: Laspeyresia piperana (Kearf.), 
Laspeyresia spp. in Pinus spp. cones; undet. Tachinidae assoc. w. 
pine cones. 
Biology: Keen, 1958, loc. cit. supra, pp. 100-102. 

Genus PLATYTERMA Walker, p. 551 

Taxonomy: von Rosen, 1958. Opusc. Ent. 23: 205 (syn. of Mesopolobus) . 

*nobiIis Walker. Minn., Wyo., Colo.; Europe. 

Platyterina nobile Walker, 1834. Ent. Mag. 2: 304. 9. 
Platyterma citripes Ashmead (syn. by von Rosen, 1959. Ent. Tidskr. 80: 
160 [Mesopolobus]) . 

Genus COELOPISTHIA Foerster, p. 551 

*Kranophoru8 Graham, 1956. Ent. Mo. Mag. 92 : 257. 

Type : Pteromalus extentus Walker. By monotypy. 



PTEROMALIDAE 263 

Taxonomy: Peck, 1963, loc. cit. supra, p. 665 (Kranophorus unnecessa- 
rily proposed). 

suborbicularis ( Prove ncher ) . 

Taxonomy : Burks, 1964, loc. cit. supra, p, 1262. 

Genus DIBRACHOIDES Kurdjumov, p. 551 

druso (Walker). Calif. 

Biology: Clausen, 1956, loc. cit. supra, p. 50. — Fisher et at., 1961. Jour. 

Econ. Ent. 54: 197.— Delucchi, 1962 . Awamia 2: 124. 
Taxonomy: Peck, 1963, loc. cit. supra, p. 667 (dynastes). 

Genus UROLEPIS Walker, p. 551 

i-ufipes (Ashmead). N. Y., Minn., Nebr., Kans., Tex., Utah, Oreg. 

Genus TRICHOMALUS Thomson, p. 552 

*perfectus (Walker). Idaho, B. C, Wash., Oreg., Calif.; Europe. Host: Ceu- 

torhynclms assimilis (Payk.). 
Pteromalus perfectus Walker, 1835. Ent. Mag. 2: 488. $. 
Trichoiitalus fasciatus of Catalog (misdet. teste Peck, 1963, loc. cit. 

supra, p. 669 ) . 
Biology: McLeod, 1953. Ent. Soc. B. C. Proc. 49: 12.— Walz, 1957. Ent. 

Soc. Amer. Ann. 50: 219.— Jourdheuil, 1960. Ann. Epiphyt. 11: 

462. 
Taxonomy: Delucchi and Graham, 1956. Beitr. z. Ent. 6: 571. 

Genus DIGLOCHIS Foerster, p. 552 

occidentalis (Ashmead). Wis., Man., Oreg. Hosts: Chrysops frigidus 0. S., 
C. furcatus Walk., C. nigripcs Zett., Hybomitra affinis (Kby.). 
Biology: James, 1952. Ent. Soc. Ont. Ann. 82: 70.— James, 1964 (1963). 
Canad. Ent. 95: 1229-1231 (imm. stages). 

Genus TRiTNEPTIS Girault, p. 552 

diprionis Gahan. 

Taxonomy: Finlayson, 1963. Canad. Ent. 95: 500. 

hemerocampae Girault. N. S., N. B., P. E. I., Tenn., N. Mex., Wash., Calif. 
Hosts: Platyprepia guttata Bdv. ; delete Diprion hercyniae 
(Htg.). 

Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, p. 102. 

klugii (Ratzeburg). Ont., Man., Sask. Host: Neodiprion sertifer (Geof.). 

Biology: Lejeune and Hildahl, 1954. Canad. Ent. 86: 341. — Graham, 
1956. For. Sci. 2: 152.— Griffiths, 1959. Canad. Ent. 91: 508.— 
Schneiderman and Horwitz, 1958. Jour. Expt. Biol. 35: 520. — 
Drooz, 1960. U. S. Dept. Agr. Tech. Bui. 1212, p. 33.— Ohnesorge, 
1962. Ztschr. f. Angew. Ent. 49: 142. 
Taxonomy: Finlayson, 1963. Canad. Ent. 95: 500. 

Genus DIBRACHYS Foerster, pp. 553-554 

oavus (Walker). Cosmopolitan. Hosts: Choristoneura pimis Free., Spilonota 
occllana (D. and S.), Ostrinia nuhilalis (Hbn.), Dioryctria dis- 
clusa Hein., Neodiprion excitans Roh., N. sertifer (Geof.), N. le- 
contei (Fitch), Apanteles glomeratus (L.) ; usually a sec. par., but 
may be primary. 
Biology: Reeks and Smith, 1956. Canad. Ent. 88: 572. — Lyons, 1957. 
Canad. Ent. 89: 75.— Baeva, 1959. Akad. Nauk Tadzh. SSR Inst. 
Zool. and Par. Trudy 119: 55, 69, 77.— Schaffner, 1959, loc. cit. 
supra, p. 89.— Richter, 1960. Ent. Ztschr. 70: 115.— Wylie, 1960. 
Entomophaga 5: 120. — Bendel-Janssen, 1962. Ztschr. f. Pflan- 
zenkrank. 9: 527.— Allen, 1962, loc. cit. supra, pp. 103-106.— Dixon 
and Benjamin, 1963. Jour. Econ. Ent. 56: 267. — Halstead, 1963. 
G. B. Pest Inf. Res. Bd. Rpt. 1962, pp. 5-6. 
Taxonomy: Finlayson, 1960. Canad. Ent. 92: 40, 44.— Finlayson, 1963. 
Canad. Ent. 95: 501 (imm. stages). 



264 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus EUPTEROMALUS Kurdjumov, pp. 554-555 

dubius (Ashmead). Minn. Host: Lydella sp., par. on Ostrinia nubilalis 
(Hbn.). 

hemipterus (Walker). Holarctic. Host: Lydella sp., par. on Ostrinia nubi- 
lalis (Hbn.), 
Biology: Clausen, 1956, loc. cit. supra, pp. 103-104. — Russo, 1959. Lab. 
Ent. Agr. Portici Bol. 17: 176.— Hemer, 1960. Ztschr. f. Angew. 
Ent. 46: 105.— Gersdorf, 1961. Ztschr. f. Angew. Ent. 47: 402. 

sarcophagae Gahan. Idaho, Ariz. 

subapterus (Riley). Holarctic. 

tachinae Gahan. Ohio, Iowa, Kans., Tex. Hosts: Ostrinia nubilalis (Hbn.), 
Hypoderma lineatum (DeV.). 

viridescens (Walsh). Ont., Minn., Wash. 

Biology: Harcourt, 1960. Canad. Ent. 92: 423. 

Genus LARIOPHAGUS Crawford, p. 555 

Taxonomy: Graham, 1957. Ent. Mo. Mag. 93: 217. 

distinguendus (Foerster). Cosmopolitan. 

Biology and Taxonomy: Kashef, 1954. Soc. Ent. France Bui. 59: 93. — 

Kashef, 1956 (1955). Soc. Ent. France Ann. 124: 58-88. 
Biology: Kaschef. 1959. Soc. Ent. Egypte Bui. 43: 171.— Kaschef. 1961. 

Ztschr. f. Parasitenk. 21 (1) : 65-70. 

dryorhizoxeni (Ashmead). Pa. 

texanus Crawford. S. C, Miss., Ariz., Calif. 

Biology: Hinckley, 1960. Hawaii. Ent. Soc. Proc. 17: 267. 

Genus SCHIZONOTUS Ratzeburg, p. 555 

Taxonomy: Boucek, 1958. Mus. Nat. Pragae Acta Ent. 32: 395. 

latus (Walker). N. Mex. Delete sieholdi (Ratzeburg) from syn. in Sup. 1, 
p. 77; N. Amer. records under that name in Catalog and Sup. 1 
refer to latus (Walker). 
Taxonomy: Boucek, 1958, loc. cit. supra, pp. 396-398. 

Genus METASTENUS Walker 
(=:Scymnophagus Ashmead of Catalog, p. 556) 

Taxonomy: Graham, 1956. Ent. Mo. Mag. 92: 256. 

lownsendi (Ashmead). Mont. 

Taxonomy: Delucchi, 1954. Bui. Ent. Res. 45: 265.— Graham, 1956, loc. 
cit. supra, p. 256. 

Genus XENOCREPIS Foerster, p. 556 

Taxonomy: Graham, 1957. Ent. Mo. Mag. 93: 235.— Boucek, 1958. Mus. 

Nat. Pragae Acta Ent. 32: 398.— von Rosen, 1960. Opusc. Ent. 

25: 8. 
Nomenclature: Peck, 1963, loc. cit. supra, p. 664. 

♦morys (Walker). Holarctic; Idaho, Wash., Oreg., Calif. Host: Ceutor- 

hynchus assimihs (Payk.). 
Pteromalus niorys Walker, 1848. List. Hym. Brit. Mus. Chalc, pt. 2, p. 

197. $. 
Xenocrepis pura Mayr, 1904 (syn. by Graham, 1957, loc. cit. supra, p. 

235). 
Biology: Walz, 1957. Ent. Soc. Amer. Ann. 50: 220.— von Rosen, 1958. 

Opusc. Ent. 23: 234.— Jourdheuil, 1960. Ann. des Epiphyt. 11: 

462. 

Genus MERAPORUS Walker, p. 556 
requisitus Tucker. S. C. Host: Taken in stored wheat. 



PTEROMALIDAE 265 

*Genus SPANIOPUS Walker 
( = Neopo//ce//s Hincks of Catalog, p. 556) 

Taxonomy: Graham, 1956. Ent. Mo. Mag. 92: 250.— Peck, 1963, loc. cit. 
supra, p. 695. 

modestus (Gahan). 111. 

Taxonomy: Graham, 1956, loc. cit. supra, p. 251 (? syn. of dissimilis 
Walker). 

websteri (Ashmead). Md. 

♦Genus ARTHROLYTUS Thomson 

Pteromaliis subg. Arthrolytus Thomson, 1878. Hyn. Scand., v. 5, pp. 147, 
158. 

Type: Arthrolytus punctatus Thomson. Desig. by Ashmead, 
1904. 

Place in tribe Pteromalini before Psychophagus Mayr, p. 557. 

fasciatus (Provancher) . Que., N. H., Mass., Pa. Transf. from Unplaced Spe- 
cies, p. 594, teste Burks, 1964, loc. cit. supra, p. 1262. 

Genus PSYCHOPHAGUS Mayr, p. 557 

omnivorus (Walker). Holarctic; Colo. Hosts: Neodiprion lecontei (Fitch), 

Cephalcia marginata Middk. ; often a sec. par. 
Taxonomy: Bakkendorf, 1955. Ent. Medd. 27: 137-143.— Boucek, 1961. 

Mus. Nat. Pragae Acta Ent. 34: 83.— Finlayson, 1963. Canad. 

Ent. 95: 499, 501-502 (imm. stages). 
Biology: Bogavac, 1958. Zash. Bilja 46: 80. — Keen, 1958, loc. cit. supra, 

p. 89. — Schaffner, 1959, loc. cit. supra, p. 89. 

tortricidis (Brues). Maine, Ont., Wis., Minn., Wyo. Host: Choristoneura 
pinus Free., C. conflictant (Walk.). 
Biology: Dixon and Benjamin, 1963. Jour. Econ. Ent. 56: 267. — McGu- 
gan and Blais, 1959. Canad. Ent. 91: 766, 775-776. 

Genus CECIDOSTIBA Thomson, p. 557 

Taxonomy: Delucchi, 1957. Entomophaga 2: 140. 

acuta (Provancher). N. S., Que., south to W. Va. west to Wash., Oreg., 
Calif. 
Taxonomy: Burks, 1964, loc. cit. sxipra, p. 1257. 

burkei Crawford. Colo., N. Mex., Idaho, Utah. Host: Dendroctonus obesus 
(Mann.). 

dendroctoni Ashmead. N. C, Calif. 

polygraphi Ashmead. Maine. 

thomsoni Crawford. Wyo., Wash., Oreg. 

Genus CATOLACCUS Thomson, pp. 557-558, Sup. 1, p. 78 

*Merisoides Masi, 1911. Lab. Zool. Gen. e Agr. Portici Bol. 5: 141. 

Type : Merisoides crassiceps Masi. By monotypy. 
Taxonomy: Delucchi, 1956. Ztschr. f. Angew. Ent. 39: 230. 

aeneoviridis (Girault). Throughout south. Canada, entire U. S.; Mexico, 
West Indies. Hosts: Apanteles medicaginis Mues., Hormius ba- 
salis (Prov.), Goniozus platynotae Ashm. 
Biology: Atkins et al., 1957. Ent. Soc. Amer. Ann. 50: 257. — Allen and 
Smith, 1958. Hilgardia 25: 29. 

cyanoideus Burks. Minn., Wyo. 

kansensis (Girault). Ariz., Calif . 

victoria Burks. Utah. 



266 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus HETEROLACCUS Masi, Sup. 1, p. 78 
Taxonomy: Delucchi, 1958. Bol. Zool. Agr. e Bach., ser. 2, 1: 54. 
hunteri (Crawford). Ky. Host: Prionomerus calceatus Say. 

Genus ZATROPIS Crawford, pp. 558-559 

albiclavus (Girault). Mich., Wis., Idaho, Utah, Ariz. Host: Asphondylia sp. 
bud gall on Larrea sp.; Dasyneiira gleditschiae 0. S. 

bruchivorus (Ashmead). Fla., Tex., Ariz., Calif.; Mexico. 

catalpae Crawford. N. Y. west to 111. and Minn. Hosts: Dasyneura gledits- 
chiae O. S., Hyperdiplosis phlox Greene. 

incertus (Ashmead). Fla., Ariz. 

nigroaeneus (Ashmead). Fla., Mo., Nebr. 

perdubius (Girault). Mass., N. Y., N. J., D. C, Ohio, 111. Hosts: Anthon- 
omus musculus (Say), Smicronyx tychoides Lee. 

rosaecolis Burks. Mo., Nebr. 

Biology: Balduf, 1959. 111. Biol. Monog. 26, pp. 120-121. 

tortricidis Crawford. Fla., Wis. 

Genus CAENACIS Foersfer, p. 559 

cupraeus (Provancher) . Transf. from Merisus Walker, p. 547, teste Burks. 
1964, loc. cit. supra, p. 1261. 

Genus PSEUDOCATOLACCUS Masi, p. 559 
americanus Gahan. N. Mex., Utah. Host: Asphondylia sp. gall on Larrea. 

Genus MUSCIDIFURAX Girault and Sanders, p. 559 

raptor Girault and Sanders. Md., Nebr., Tex., Ariz., Calif.; Hawaii, West 
Indies, S. Amer., Africa. Hosts: Fannia caniciilaris (L.). Sto- 
moxys calcitrans (L.). 

Genus NASONIA Ashmead, pp. 559-560 

Taxonomy: Hedqvist, 1963, loc. cit. supra, p. 125. 

vitripeniiis (Walker). 

Biology: Schneiderman and Horwitz, 1958. Jour. Expt. Biol. 35: 520. — 
Wylie, 1958. Canad. Ent. 90: 597.— Ohgushi, 1959. Zool. Soc. of 
Japan, Zool. Mag. 68: 426. — Ohgushi, 1961. Jap. Jour. Appl. Ent, 
and Zool. 5: 154.— Saunders, 1962. Jour. Insect Physiol. 8: 309.— 
Nagel and Pimental, 1963. Canad. Ent. 95: 208.— Wylie, 1963. 
Canad. Ent. 95: 881-886.— King and Hopkins, 1963. Jour. Expt. 
Biol. 40: 751-761.— Beard, 1964. Jour. Insect Path. 6: 1-7.— Nagel 
and Pimental, 1964. Ecology 45: 658-660. 

Morphology: Cutler, 1955. Roy. Ent. Soc. London Proc. (A) 30: 73-81.^ 
King, 1962. Roy. Ent. Soc. London Proc. (A) 37: 73, 121. 

Genetics: Whiting, 1950. Genetics 35: 699.— Whiting, 1954. Evolution 
40: 135.— Whiting, 1955. Genetics 40: 602.— Whiting, 1958. In- 
ternatl. Cong. Ent., Proc. 10, vol. 2, p. 857.— Whiting, 1958. ASB 
Bui. 5: 16. 

Taxonomy: De Santis, 1957. Univ. Nac. de La Plata, Mus. Notas 19: 
169.— Tonapi, 1958. Roy. Ent. Soc, London Trans. 110: 550.— 
Saul, 1960. Rev. Suisse de Zool. 67: 270.— Hedqvist, 1963, loc. cit. 
supra, pp. 125-127. 

*Genus DINARMUS Thomson 

( = Bruchobius Ashmead of Catalog, p. 5601 

Dinarmus Thomson, 1878. Hym. Scand., v. 5, pp. 50, 56. 

Type: Dinarmus acutus Thomson. Desig. by Ashmead, 1904. 



PTEROMALIDAE 261 

Taxonomy: Delucchi, 1956. Ztschr. f. Angew. Ent. 39: 240. — Ferriere 
and Delucchi, 1957. Entomophaga 2: 121. 

*acutus (Thomson). N. J., Pa., N. C, Mich., Ky., Tenn., Minn., Nebr., Oreg., 
Europe. Hosts: Bnichus brachialis Fahr., Bruchidius ater (Mar- 
sham). 

Dimachus (Dinarmus) acutus Thomson, 1878. Hym. Scand., v. 5, p. 56. 
9. 

Sphaerakis mayri Masi (syn. by Delucchi, 1956, loc. cit. supra, p. 243). 

laticeps (Ashmead). 

Taxonomy: Delucchi, 1956, loc. cit. supra, pp. 248-246. 
Biology: Cheema and Misra, 1962. Cur. Sci. (India) 31: 21. 

mayri (Masi). Syn. of acutus (Thomson). 

Genus PTEROMALUS Swederus, pp. 560-561 

archippi Howard. N. H. south to Fla., west to Ark., Tex., and Calif. 

fuscipes (Provancher). 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1262. 

puparum (Linnaeus). Cosmopolitan. 

Biology: Clausen, 1956, loc. cit. supra, p. 49. — Blunck, 1957. Jour. Econ. 

Ent. 50: 836.— Wiltshire, 1958. Roy. Ent. Soc, London Trans. 

110: 241.— Blunck and Wilbert, 1959 (1957). Internatl. Cong. 

Crop Prot. Proc. 1 : 1017. 
Taxonomy: Burks, 1964, loc. cit. supra, p. 1261 {pieridis Provancher). 

vanessae Howard. Calif. 

*Genus SPINTHERUS Thomson 

Spintherus Thomson, 1878. Hym. Scand., v. 5, p. 88, 129. 

Type: Spintherus obscurus Thomson. By monotypy. 

Insert in tribe Pteromalini following Pteromalus Swederus, p. 561. 

orchestis (Ratzeburg), n. comb. N. Y., Ohio; Europe. Host: Rhynchaenus 
pallicornis (Say). 

Genus HABROCYTUS Thomson, pp. 561-563 

aculiventris Peck. 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1260 (acutus Provancher). 

analis (Ashmead). Nev. Hosts: Pemphredon giffardi (Roh.), Chrysidid, 
Omalus sp., par. on Pemphredon sp. 

bedeguaris (Thomson). Maine; Europe. 

Biology: Judd, 1959. Canad. Ent. 91: 728-730.— Gijswijt, 1962. Ent. 
Ber. 22: 250. 

capreae (Linnaeus). Delete; pp. is probably not a chalcid. N. Amer. records 
of a chalcid under this name cannot now be placed. 

giraulti Peck (= canadensis Girault, 1917, not Ashmead, 1887). Transf. to 
Lonchetron Graham, infra. 

medicaginis Gahan. N. Y., Mich.; Hawaii, Europe, Asia. Hosts: Brucho- 
phagus spp. 
Biology: Beardsley, 1956. Hawaii. Ent. Soc. Proc. 16: 11. — Butler and 
Hansen, 1958. Pan-Pacific Ent. 34: 228. — Carillo and Dickason, 
1963. Oreg. Agr. Expt. Sta. Tech. Bui. 68, p. 31. 

melanicrus (Provancher). 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1262. 

phycidis Ashmead. N. B., Mich. 

Biology: Dixon and Benjamin, 1963. Jour. Econ. Ent. 56: 267. 

thyridopterigis Howard. [This may be a composite sp.] Maine and Ont. 
south to N. C, west to Wis.. 111., and Mo. 



268 HYMENOPTERA OF AMERICA NORTH OP MEXICO 

Biology: Schaffner, 1959, loc. cit. supra, p. 89. — Harman and Kulman, 
1962. Jour. Econ. Ent. 55: 1007. 

*Genus LONCHETRON Graham 

Lonchetron Graham, 1956. Ann. Ent. Fenn. 22: 80. 

Type: Lonchetron fennicum Graham. Orig. desig. 

Insert in tribe Pteromalini after Habrocytus Thomson, p. 563. 

girauiti (Peck), n. comb. Transf. from Habrocytus Thomson; types exa- 
mined. 

Genus ANISOPTEROMALUS Ruschka, pp. 563-564 

calandrae (Howard). 

Biology: Chatterji, 1955. Zool. Soc. India Proc. 8: 11-23. — Ghani and 
Sweetman, 1955. Biologia (Lahore) 1: 115-139. — Cabal Concha, 
1956. Fac. Nac. Agron. Colombia Rev. 18: 61.— Calderon, 1962. 
Riv. di Parassitol. 23: 216. 
Morphology: Tonapi, 1958. Roy. Ent. Soc, London Trans. 110: 550. 

Genus TRIMEROMICRUS Gahan, p. 564 

maculatus Gahan. Mo., Okla., Tex., Idaho, Utah, Wash., Oreg. Hosts: Bm- 
chophagus spp. 
Biology: Walz, 1957. Ent. Soc. Amer. Ann. 50: 220. 

Genus SYSTELLOGASTER Gahan, p. 564 

ovivora Gahan. N. C, Ohio, Okla. 

Biology: Judd, 1955. Canad. Ent. 87: 98.— Roth and Willis, 1960. 
Smithsn. Misc. Coll. 141: 248. 

Genus HYPOPTEROMALUS Ashmead, pp. 564-565 

inimicus Muesebeck. N. B., Minn., Mo., N. Dak. 

percussor Girault. Nev., N. Mex., Oreg. Hosts: Campoplegini par. on Hem- 
erocampa pseudotsugata McD. and other Lepidoptera. 

Genus HEMITRICHUS Thomson, p. 565 

rufipes (Thomson). Syn. of seniculus (Nees). 

*seniculus (Nees). Que. south to D. C, west to Mich, and Kans.; Europe. 

Pteroma'us seniculus Nees, 1834. Hym. Ichn. Affin. Monog., v. 2, pp. 101, 

423. 9. 
Dimachus (Hemitrichus) rufipes Thomson (syn. by Delucchi, 1955. Soc. 

Roy. Ent. Belg. Mem. 27: 173). 

Genus NORBANUS Walker, p. 565 
perplexus (Ashmead). N. J., Del., Tenn., Kans. 

*Genus LAMPOTERMA Graham 
( = A1efosfenus of Catalog, pp. 565-566) 

Lampoterma Graham, 1956. Ent. Mo. Mag. 92: 256 (n. gen. for Metas- 

tenus Thomson, not Walker) . 

Type : Metastenus viridis Thomson. Orig. desig. 
Taxonomy: Peck, 1963, loc. cit. supra, pp. 741-742 (n. comb's.; ch. spp. 

name endings to neuter). 

faciale (Provancher) . 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1256. 

Genus NEOCATOLACCUS Ashmead, p. 566 

tylodcrmae (Ashmead). N. H. south to Fla. west to Mich., 111., Kans., Ariz. 
Host: Onychoharis subtonsn Lee. 
Biology: Wester, 1956. Ent. Soc. Wash. Proc. 58: 286. 



PTEROMALIDAE 269 

Genus MEGATRYDYMUS Girault, p. 566 

agrili (Rohwer). Transf. from Dinotiscus Ghesquiere, p. 549; type re-exa- 
mined. 

taciii Girault. N. J., Ark., Oreg., Calif. Host: Reared from cynipid galls, on 
oak. 

Genus HABRITYS Thomson, Sup. 1, p. 79 

latro Wallace, emend, for latrus. Va. Hosts: Ectemnius stirpicola (Pack.), 
Euplilis sp. 
Biology: Krombein, 1960. Ent. News 71: 32, 67.— Krombein, 1964. Biol. 
Soc. Wash. Proc. 77 : 103. 

Tribe CLEONYMINI 

Taxonomy: Kerrich and Graham, 1957. Soc. Brit. Ent. Trans. 12: 
265-311.— Boucek, 1959 (1958). Mus. Nat. Pragae Acta Ent. 32: 
353-386 (Cleonyminae). — Hedqvist, 1963, loc. cit. supra, p. 50. 

Genus HEYDENIA Foerster, p. 566 

Taxonomy: Hedqvist, 1957. Opusc. Ent. 22: 39.— Hedqvist, 1963, loc. cit. 
siipra, pp. 52-53. 

unica Cook and Davis. Ala., Tex., Ariz., Calif. 

Genus TRIGONODERUS Westwood, p. 545 

Taxonomy: Kerrich and Graham, 1957, loc. cit. supra, pp. 284-285. 
Transf. with included spp. from tribe Tridymini, p. 545, and insert in 
tribe Cleonymini following Heydenia Foerster, p. 566. 

algonquinia Girault. Transf. to Platygerrhus Thomson, infra. 

conicus Girault. Transf. to Gastracanthus Westwood, infra. 

unguttus Girault. Que., N. J., D. C, HI., Tex. Host: Euparius marmoreus 
Oliv. in shelf fungi. 
Biology: Graves, 1960. Ent. Soc. Amer. Ann. 53: 74. 

*Genus PLATYGERRHUS Thomson 

Platygerrhus Thomson, 1878. Hym. Scand., v. 5, pp. 4, 13. 

Type: Platygerrhus gracilis Thomson. By monotypy. 
Taxonomy: Kerrich and Graham, 1957, loc. cit. supra, pp. 300-303. 

Insert in tribe Cleonymini following Heydenia Foerster, p. 566. 

algonquinia (Girault). Transf. from Trigonoderus Westwood, teste Kerrich 
and Graham, 1957, loc. cit. supra, p. 303. 

*Genus GASTRACANTHUS Westwood 

Gastracanthus Westwood, 1833. Mag. Nat. Hist. 6: 121. 

Type: Gastracanthus pulcherrimus Westwood. By monotypy. 
*Hetroxys Westwood, 1833, loc. cit. supra, p. 495. N. name. 
*Etroxys Foerster, 1856. Hym. Stud., v. 2, pp. 66, 71. Emend. 
*Aetroxys Walker, 1872. Notes on Chalc, p. 110. Emend. 
*Photisntus Thomson, 1878. Hym. Scand., v. 5, pp. 66, 71. 

Tvpe: Photismus nubilosus Thomson. By monotypy. 
*Heteroxys Dalla Torre, 1898. Cat. Hym., v. 5, p. 101. Emend. 
Taxonomy: Kerrich and Graham, 1957, loc. cit. supra, pp. 281-284. 

Insert in tribe Cleonymini following Heydenia Foerster, p. 566. 

conicus (Girault). Que., Ont., N. Y., N. J., Md. Delete Ariz, from dist. 

Transf. from Trigonoderus Westwood, teste Kerrich and Graham, 1957, 
loc. cit. supra, p. 283. 



270 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Genus JANSSONIELLA Kerrich 

Janssoniella Kerrich, 1957. In Kerrich and Graham, Soc. Brit. Ent, 
Trans. 12 : 303. 

Type : Janssoniella caudata Kerrich. Orig. desig. 

Insert in tribe Cleonymini following Heydenia Foerster, p. 566. 

'*' caudata Kerrich. N. J., B. C, Calif.; Europe. Host: Reared from bracket 
fungi. 
Janssoniella caudata Kerrich, 1957, In Kerrich and Graham, Soc. Brit. 
Ent. Trans. 12: 304. ?. 

Genus EPISTENIA Westwood, p. 567 

*Idiobia Brethes, 1927. Ent. Mitt. 16: 330. 

Type : Idiobia schmidii Brethes. By monotypy. 
Taxonomy: Boucek, 1959, loc. cit. supra, p. 381. 

coeruleata Westwood. Va., Fla., Ariz., Calif. Hosts: Isodontia elegans 
(Smith), Leptochilus republicanus zendaloides (Robt.), Trypoxy- 
lon (Trypargilum) tridentatum Pack., T. (T.J sp. 
Biology: Krombein, 1959. Ent. News 70: 19. 

odyneri Ashmead. N. Mex., Ariz. 

osmiae (Ashmead). Mo., Tex. Host: Leptochilus republicanus zendaloides 
(Robt.). 

poliia (Say). D. C, N. C, Fla. 

Genus PTINOBIUS Ashmead, p. 567 

Taxonomy: Boucek, 1959, loc. cit. supra, p. 363 (prob. a syn. of Clean- 
ymus Latreille). 

Genus CHEIRO?ACHUS Westwood, pp. 567-568 

Taxonomy: Hedqvist, 1963, loc. cit, supra, pp. 102-103. 

colon (Linnaeus). Canada, north. U. S. and Pacific Coast; Europe. 
Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 103-107. 

*Genus COLOTRECHNUS Thomson 
Insert in tribe Cleonymini after Cheiropachus Westwood, p. 658. 

♦COLOTRECHNUS subgenus COLOTRECHNUS Thomson 

Colotrechnus Thomson, 1878. Hym. Scand., v. 5, p. 46. 

Type: Colotrechnus subcoeruleus Thomson. By monotypy. 

The typical subgenus does not occur in North America. 

♦COLOTRECHNUS subgenus ZANONIA Masi 

Zanonia Masi, 1921. Genova Mus. Civ. Stor. Nat. Ann., ser. 3, 9: 184. 
Type: Zanonia viridis Masi. By monotypy. 

Taxonomy: Masi, 1922. Soc. Ent. Ital, Bol. 54: 111 (closely rel. to Colo- 
trechnus Thomson). — Delucchi, 1956. Ztschr. f. Angew. Ent. 39: 
233 (syn. of Colotrechnus Thomson),— Burks, 1958. Fla. Ent. 41: 
13 (subg. of Colotrechnus Thomson). 

*ignotus Burks. N. C, Fla., Okla., Tex., Ariz., Calif. Host: Reared from 
heads of Bidens and other Compositae. 
Colotrechnus (Zanonia) ignotus Burks, 1958. Fla. Ent. 41: 14. 9, $. 

Family EURYTOMIDAE 

Genus HARMOLITA Motschulsky, pp. 568-573 

Taxonomy: Gahan, 1957. Brooklyn Ent. Soc. Bui. 52: 82-83.— Claridge, 
1958. Ent. Mo. Mag. 94: 81-85 (Tetramesa Walker) .—Claridge, 
1961. Roy. Ent. Soc. London Trans. 113: 175-216 {Tetramesa 



EURYTOMIDAE 271 

Walker) .—Peck, 1963, loc. cit. supra, pp. 751-783 {Tetramesa 
Walker). 

albomaculata (Ashmead). Holarctic. 

Taxonomy and Biology: Claridge, 1961, loc. cit. supra, p. 211. 

agropyrophila Phillips and Emery. Syn. of comuta (Walker). 

atlantica Phillips and Emery. Syn. of linearis (Walker). 

bambusae Phillips. Ala. 

*comuta (Walker). Atlantic Coast west to Kans.; Europe. Host: Agropyron 

repens. 
Isosoma cornutum Walker, 1832. Ent. Mag. 1: 19. $. 
Harniolita agropyrophila Phillips and Emery (syn. by Claridge, 1961, loc. 

cit. supra, p. 212). 

dactylicola Phillips and Emery. Syn. of longula (Dalman). 

feslucae Phillips and Emery. Poss. syn. of European H. giraudi (von 
Schlechtendal) teste Claridge, 1961, loc. cit. siipra, p. 194. 

hordei (Harris). P. E. I. 

Biology: Cannon, 1963. Canad. Ent. 95: 28. — Sterling and MacLaren, 
1964. Canad. Jour. Plant Sci. 44: 53-56. 

♦linearis (Walker). Holarctic; N. Y., Mich., Ind. Hosts: Agropyron spp. 
Isosoma lineare Walker, 1832. Ent. Mag. 1: 22. 9. 

Harmolita atlantica Phillips and Emery (syn. by Claridge, 1961, loc. cit. 
supra, p. 192). 

lolii Gahan. Oreg. 

*longicomis (Walker). N. Y., S. Dak. ; Europe. Host: Phalaris arundinacea. 
Isosoma longicorne Walker, 1832. Ent. Mag. 1: 17. 5- 
Harmolita phalaridis Phillips and Poos (syn. by Claridge, 1961, loc. cit. 
supra, p. 209). 

*Iongula (Dalman). Mass. and N. Y. south to Va., west to Mich., Tenn. and 

Utah; Europe. Host: Dactylis glomerata. 
Eurytoma longula Dalman, 1820. Svenska Vetensk. Akad. Handl. 41: 

173, pi. 7, fig. 15. $. 
Harmolita dactylicola Phillips and Emery (syn. by Claridge, 1961, loc. 

cit. supra, p. 210). 

maculata (Howard). England. Hosts: ^romus spp. 

ovata Phillips and Emery. Sec. homonym of Isosoma ovata Szepligeti, 1901; 
requires a new name, teste O. Peck in litt. 

opuntiae Muesebeck. Tex., N. Mex., Nev., Calif. 

phalaridis Phillips and Poos. Syn. of longicomis (Walker) . 

tritici (Fitch). 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1257 {basilaris Provancher). 

Genus GAHANIOLA Erdos, Sup. 1, p. 79 

phyllostachitis (Gahan). Ala.; China, Japan. 

Genus AXIMA V/alker, p. 573 

zabriskiei Howard. Que., Va., Ind., Tex., Calif. 

Biology: Krombein, 1960. Ent. News 71: 32, 68. 

Genus MACRGRILEYA Ashmead, p. 573 

oecanthi (Riley). Ariz. 

Genus RILEYA Ashmead, p. 573 

cecidomyiae Ashmead. Md., N. Mex., Tex., Idaho, Ariz. Hosts: Asphondylia 
spp. 
Rileya tegularis Gahan, 1918, n. syn. ; types re-examined. 



272 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Biology: Highland, 1964. Jour. Econ. Ent. 57: 83. 
mellea Ashmead. Host : Reared from leaf gall on Eugenia axillaris. 
piercei Crawford. Fla. 
tegularis Gahan. Syn. of cecidomyiae Ashmead. 

Genus CHRYSEIDA Spinola, p. 574 

*bennelti Burks. Tex.; Mexico, West Indies. Hosts: Bnichus sp. in Cajanus 
cajan, Merobmchus sp. in Havardia brevifolia. 
Chryseida bennetti Burks, 1956. Brooklyn Ent. Soc. Bui. 51: 115. 9. 

Genus TENUIPETIOLUS Bugbee, Sup. 1, p. 80 
medicaginis (Gahan). Tex., Calif. 

Genus EURYTOMA Illiger, pp. 574-579 

Delete Eurytomocharis from syn. ; valid genus, infra. 
Taxonomy: Hedqvist, 1963, loc. cit. supra, pp. 33-34. 

abnormis Ashmead. Syn. of conica Provancher. 

appendigaster (Swederus). 

Taxonomy: Claridge, 1959. Ent. Mo. Mag. 95: 4.— Claridge, 1960. Ent. 

Medd. 29: 248 (verticillata Fabricius) .— Peck, 1963, loc. cit. 

supra, p. 818 (verticillata) . 
Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, pp. 106-108. 

ashmeadi Peck (= minuta Ashmead, 1894, not Walker, 1832). Transf. to 
Eurytomocharis Ashmead, infra. 

auriceps Walsh, changed status. 

Eurytoma auriceps auriceps Walsh of Catalog. Raised to sp. rank (teste 

Bugbee, 1956. Ent. Soc. Amer. Ann. 49: 503). 
Eurytoma auriceps seminatrix Walsh (syn. by Bugbee, 1956, loc. cit. 

supra, p. 503). 
Eurytoma vagabuntla Ashmead (syn. by Bugbee, 1956, loc. cit. supra, p. 

503). 

auriceps seminatrix Walsh. Syn. of auriceps Walsh. 

bicolor Walsh. Md. Host: Reared from Auiaczdea gall on Lacittca. 

bigeloviae Ashmead. Utah, Calif. 

Eurytoma chalcidiformis Girault (syn. by Bugbee, 1956, loc. cit. supra, p. 
504). 

bolteri Riley, changed status. 

Eurytoma bolteri bolteri Riley of Catalog. Raised to sp. by Miller, 1963. 

Ohio Jour. Sci. 63: 72. 
Biology: Miller, 1963, loc. cit. supra, p. 72. 

*brevivena Bugbee. Ariz. Host: Disholcaspis quercusglobulus (Fitch) galls 
on Quercus gambelli. 
Eurytoma brevivena Bugbee, 1958. Ent. Soc. Amer. Ann. 51: 198. 9, $. 

*calycis Bugbee. Ont. Host: Phytophagous in shoots of Pinus banksiana. 
Eurytoma calycis Bugbee, 1961. Canad. Ent. 93: 33. 9, $. 
Biology: Thomas and Herdy, 1961. Canad. Ent. 93: 34-39. 

*celtigalla Bugbee. Tex. Host: ? Phytophaga celtiphylla Felt gall on Celtis 
spp. 
Eurytoma celtigalla Bugbee, 1957. Kans. Ent. Soc. Jour. 30: 49. 9, $. 

chalcidiformis Girault. Syn. of bigeloviae Ashmead. 

conica Provancher. Ont. south to N. C, west to Kans. and Tex. Hosts: 
Phloeosinus dentatus (Say), scolytids in elm. 
Eurytoma phloeosini Ashmead (syn. by Bugbee, 1956, loc. cit. supra, p. 
504). 



EURYTOMIDAE 273 

Eurytoma abnormis (Ashmead) (syn. by Bugbee, 1956, loc. cit. supra, p. 

504). 
Taxonomy: Burks, 1964, loc. cit. supra, p. 1259. 

crassineura Ashmead. Que. south to N. C. west to Idaho and Calif. Cited in 
error as sec. par. of Acrobasis comptoniella Hulst by Schaffner, 
1959, loc. cit. supra, p. 90; specimens re-examined by R. E. Bug- 
bee. 

eragrostidis (Howard). Transf. to Eurytomocharis Ashmead, iw/ra. 

*flavovultus Bugbee. Tex. Host: Pachypsylla venusta galls on Celtis spp. 
Eurytoma flavovultus Bugbee, 1957. Kans. Ent. Soc. Jour. 30: 45. 9, $. 

*furva Bugbee. Calif. Host: Disholcaspis washing tonensis (Gill.) gall on 
Quercus douglasii. 
Eurytoma furva Bugbee, 1958. Ent. Soc. Amer. Ann. 51: 198. 9, $. 

gigantea Walsh. Mont. 

Biology: Miller, 1959. Tenn. Acad. Sci. Jour. 34: 249-251. 

'''inornata Bugbee. Md. Hosts: Euplilis (Corynopus) rufigaster (Pack.), E. 
(C.) coarctata modcsta (Roh.), rarely Trypoxylon (Trypoxylon) 
sp. nesting in Hibiscus stems. 
Eurytoma inornaia Bugbee, 1962. Kans. Ent. Soc. Jour. 35: 347. 9, $. 
Biology: Krombein, 1964. Biol. Soc. Wash. Proc. 77: 103-104. 

juniperina juniperina Marcovitch. N. Dak. Host: Juniperus scopulorum ber- 
ries. 

*levivultus Bugbee. Ohio. Host : Pachypsylla gemma Riley galls on Celtis sp. 
Eurytoma levivultus Bugbee, 1957. Kans. Ent. Soc. Jour. 30: 48. 9, $. 

magdalidis Ashmead. N. C. Host: Oncideres cingulata (Say). 

minnesota Girault, corrected spelling of minnesotae of Catalog, teste Peck, 
1963, loc. cit. supra, p. 803. 

muhlenbergiae (Howard). Transf. to Eurjtomocharis Ashmead, infra. 

nigricoxa Provancher. 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1259. 

obtusiventris Gahan. 

Biology: Miller, 1959, loc. cit. supra, p. 249. 

orchidearum (Westwood). 

Eurytoma phoenix Girault (syn. by Bugbee, 1956, loc. cit. supra, p. 504). 

pachyneuron Girault. P. E. I. and N. Y., south to N. C, west to B. C. and 
Calif. Hosts: Harmolita spp. in Elymus spp. 
Eurytoma phoebus Girault (syn. by Bugbee, 1956, loc. cit. supra, p. 503). 
Eurytoma pater Girault (syn. by Bugbee, 1956, loc. cit. supra, p. 503). 

parva Phillips, new status. 

Eurytoma bolteri parva Phillips of Catalog. Raised to sp. by Bugbee, in 
litt. 

pater Girault. Syn. of pachyneuron Girault. 

phloesini Ashmead. Syn. of conica Provancher. 

phoebus Girault. Syn. of pachyneuron Girault. 

p/ioenijc Girault. Syn. of orchidearum (Westwood). 

*pini Bugbee. Que. south to Fla., west to B. C. and Wash. Hosts: Rhyacionia 

spp. in Pinus spp. 
Eurytoma pini Bugbee, 1958. Kans. Ent. Soc. Jour. 31: 197. 9, $. 
Biology: Watson and Arthur, 1959. Canad. Ent. 91: 480-484.— 

Schaffner, 1959, loc. cit. supra, p. 90 (tylodermatis) . — Arthur, 

1961. Canad. Ent. 93: 657.— Harman and Kulman, 1962. Jour. 

Econ. Ent. 55: 1007. 

punctiventris Walsh. Syn. of querciglobuli querciglobuli (Fitch). 



274 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

querciglobuli querciglobuli (Fitch), changed status. N. H. south to N. C, 

west to Minn, and Tex. Hosts: Disholcaspis mamma (Cress.), D. 

virens (Ashm.), D. quercus globulus (Fitch) galls on Quercus spp. 
Macroglenes querciglobuli Fitch (reduced to subsp. rank by Bugbee, 

1958. Ent. Soc. Amer. Ann. 51: 193-197). 
Eurytoma punctiventris Walsh (syn. by Bugbee, 1958, loc. cit. supra, p. 

193). 
Biology: Judd, 1957. Ent. News 68: 195. 

* querciglobuli rubra Bugbee. Fla., Miss. Hosts: Disholcaspis virens (Ashm.) 
and D. spongiosa galls on Quercus virginiana, D. sp. gall on Q. 
stellata. 
Eurytoma querciglobuli rubra Bugbee, 1958. Ent. Soc. Amer. Ann. 51: 
196. 9. 

^querciglobuli utahensis Bugbee. Utah. Host: Disholcaspis Colorado (Gill.) 
gall on Quercus utahensis. 
Eurytoma querciglobuli utahensis Bugbee, 1958. Ent. Soc. Amer. Ann. 
51: 197. ?. 

*sciromatis Bugbee. N. C, Ga., Fla., La. Host: Eeared from cankers on 
Pinus spp. of Cronartium fusiforme ; may be phytophagous. 
Eurytoma sciromatis Bugbee, 1962. Kans. Ent. Soc. Jour. 85: 346. 9, $. 

*semivenae Bugbee. N. Y. west to Kans. and Tex. Hosts: Pachypsylla vesi- 
cula Riley and P. mam7na Riley galls on Celtis spp. 
Eurytoma semivertae Bugbee, 1957. Kans. Ent. Soc. Jour. 30: 47. 9, $. 

solenozopheriae Ashmead. Fla. Name prob. cited in error as sec. par. of 
Bucculatrix ainsliella Murt. by Schaffner, 1959, loc. cit. supra, p. 
90. 

stigmi Ashmead. Nev., Calif. Hosts: Hoplitis producta gracilis (Mich.), 
Pemphredon giffardi (Roh.), Solierella similis (Bridw.), Trypoxy- 
lon sculleni Sandh. 

studiosa Say. Que. and Ont. south to Fla., west to Colo, and N. Mex. Host: 
Disholcaspis succinipes (Ashm.). 
Eurytoma succinipedis Ashmead (syn. by Bugbee, 1956, loc. cit. supra, p. 
505). 

succinipedis Ashmead. Syn. of studiosa Say. 

triodiae (Howard). Transf. to Eurytomocharis Ashmead, trr/ra. 

*tumoris Bugbee. Calif. Host: ? Phytophagous in stems and branches of 
Pinus sylvestris. 
Eurytoma tumoris Bugbee, 1962. Kans. Ent. Soc. Jour. 35: 345. 5, $. 
Biology: Stark and Koehler, 1964. Pan-Pacific Ent. 40: 41-46. 

tylodermatis Ashmead. Hosts: Curculionidae only; other published host re- 
cords should be disregarded, teste R. E. Bugbee in litt. Records as 
par. of Rhyacionia spp. in Schaffner, 1959, loc. cit. supra, p. 90, 
should be referred to pini Bugbee. 

vagabunda Ashmead. Syn. of auriceps Walsh. 

Genus EURYTOMOCHARIS Ashmead 

Delete from syn. under Eurytoma Illiger, p. 574, and insert as valid 
genus following Eurytoma, p. 579. 

Taxonomy: Bugbee, 1956, loc. cit. supra, pp. 505-506. — Peck, 1963, loc. 
cit. supra, pp. 831-832. 

ashmeadi (Peck), n. comb. Transf. from Eurytoma Illiger, teste Peck, 1963, 
loc. cit. supra, p. 831 (minuta Ashmead). 

eragrostidis Howard. Transf. from Eurytoma Illiger; types examined. 

muhlenbergiae (Howard). Transf. from Eurytoma Illiger, teste Bugbee, 
1956, loc. cit. supra, p. 505. 

triodiae Howard. Transf. from Eurytoma Illiger; types examined. 



EURYTOMIDAE 275 

Genus HEIMBRA Cameron, Sup. 1, p. 82 

opaca (Ashmead) l = opacas (1) of Sup. 1, p. 82]. Kans., N, Mex., Utah, 
Calif. 

Genus EUDECATOMA Ashmead, pp. 579-582 

Nomenclature: Peck, 1963, loc. cit. supra, pp. 833-844 (subspp. eh. to 
var.). 

amsterdamensis (Girault). Syn. of mellea (Curtis). 

florida (Girault), corrected spelling of floridae of Catalog, teste Peck, 1963, 
loc. cit. supra, p. 836. 

*mellea (Curtis). Que., N. Y., Oreg.; Holarctic. Hosts listed in Catalog 

under amsterdamensis (Girault), p. 579. 
Decafofna niel/ca Curtis, 1831. Brit. Ent. 8: 345. $ (= 9). 
Decatoma amsterdamensis Girault, 1917 (syn. by Claridge, 1959. Soc. 

Brit. Ent. Trans. 13: 152, 160). 

nubilistigma (Walsh). N. H. Host: Reared from gall of Euura sp. on Salix 
discolor. 

occidentalis occidentalis (Balduf). Utah, Oreg. Hosts: Disholcaspis elclora- 
densis (Beut.), Xantlioteras eburneum (Bass.), twig gall on Qu- 
ercus garryana. 

vacciniicola (Balduf). 

Biology: Judd, 1959. Amer. Micros. Soc. Trans. 78: 213. 

Genus SYSTOLE Walker, p. 582 

Taxonomy: Claridge, 1959. Ent. Mo. Mag. 95: 38. 

geniculata Foerster. 

Biology : Roberts, 1963. Jour. Econ. Ent. 56 : 541. 

Genus BRUCHOPHAGUS Ashmead, pp. 582-583 

Taxonomy: Boucek, 1954. Mus. Nat. Pragae Acta Ent. 30: 76 (subg. of 
Eurytoma) . — Fedoseeva, 1954. Mosk. Univ. Vestnik 9 (5): 
113-118.— Fedoseeva, 1958. Zool. Zhur. 37: 1345-1351. 

Biology: Burks, 1958 (1957). Ent. Soc. Wash. Proc. 59: 273-277 (hosts). 
—Fedoseeva, 1960. Nauch. Dok. Vyssh. Shkoly. Biol. 3: 14-18 
(ecology) . 

*kolobovae Fedoseeva. Cosmopolitan. Hosts: Seeds of Lofits spp, 

Bruchophagus kolobovae Fedoseeva, 1956. Akad. Nauk S.S.S.R. Dokl. 

Ill: 491. 2, $. 
Taxonomy: Fedoseeva, 1958, loc. cit. supra, p. 1349. — Perju, 1960. Acad. 
R. P. Romaine, Bucharest Comun. 10: 575. — Szelenyi, 1961. Buda- 
pest. Noven, Kut. Int. Evkon 8: 133, 136.— Strong, 1962. Ent. 
Soc. Amer. Ann. 55 : 1-4. 

mexicanus Ashmead. Idaho, Ariz., Calif. Host: Seeds of Astragalus mollis- 
simus. 

*platvptera (Walker). Cosmopolitan. Hosts: Seeds of clovers. 
Systole platyptera Walker, ISBi. Ent. Mag. 2: 153. 9. 
Eurytoma gibba Boheman, 1836 (syn. by Claridge, 1959. Ent. Mo. Mag. 

95: 39 [syn. questionably by Erdos, 1951. Folia Ent. Hung., n. s., 

4: 60 and Hellen, 1953. Not. Ent. 33: 18]). 
Biology: Carillo and Dickason, 1963. Oreg. State U. Agr. Expt. Sta. 

Tech. Bui. 68, p. 11 (gibbus). 
Taxonomy: Fedoseeva, 1954, loc. cit. supra, pp. 115, 117 (gibbus). — 

Strong, 1962, loc. cit. supra, pp. 1-4 (gibbus). 

*roddi (Gussakovsky). Cosmopolitan. Hosts: Seeds of MecZzca^fO spp. 

Euryfoma [Euryioma'\ roddi Gussakovsky in Rodd, Gussakovsky and An- 
tova, 1933. [Pests of dry-farming crops of Cent. Asia, Tashkent], 
pp. 122, 150. ?, $. 



276 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Bruchophagus gibbus medicaginis Kolobova, 1950. Ent. Obozr. 31: 70 

(syn. by Perju, loc, cit. supra, p. 575). 

Taxonomy: Nikolskaya, 1952. Acad. Sci. U.S.S.R., Keys to Fauna of 
U.S.S.R. 44: 173, 175.— Fedoseeva, 1954, loc. cit. supra, p. 116.— 
Fedoseeva, 1958, loc. cit. supra, p. 1349. — Strong, 1962, loc. cit. 
supra, pp. 1-4. 

Biology: Strong, 1962. Hilgardia 32: 229-248.— Carillo and Dickason, 
1963, loc. fit. supra, p. 25. — Strong, Bacon, and Russell, 1963. Hil- 
gardia 35: 1-12. 

sculptus (Ashmead), corrected spelling of sculpta of Sup. 1, p. 82, teste Peck, 
1963, loc. cit. supra, p. 831. 

Family CHALCEDECTIDAE 

Genus CHALCEDECTUS Walker, p. 584 

*Episystole Girault, 1927. S. Austr. Mus. Rec. 3: 317. 

Type : Episystole poeta Girault. Orig. desig. 
*I\eochalcodectus Masi, 1936. Soc. Ent. Ital. Bol. 68: 68. 

Type: Neochalcodectiis sinaiticus Masi. By monotypy. 
Taxonomy: Boucek, 1959. Mus. Nat. Pragae Acta Ent. 33: 568. 

Genus EUCHRYSIA Westwood, p. 584 

hyalinipennis Ashmead. Fla., N. Mex., Ariz. Host: Oncideres rhodosticta 
Bates. 

Family CHALCIDIDAE 

Subfamily HALTICHELLINAE 

Genus HALTICHELLA Spinola, p. 584 

longicornis Ashmead. 

Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, p. 110. 

rhyacioniae Gahan. Mass. south to S. C. and Miss., west to Nebr. and Tex. 
Host: Rhyacionia rigidana (Fern.). 
Biology: Miller and Neiswander, 1959. Ohio Agr. Expt. Sta. Res. Bui. 
840, p. 18. — Schaffner, 1959, loc. cit. supra, p. 91. 

xanticles (Walker). Iowa, Okla., Tex., Sask. Hosts: Grapholitha molesta 
(Busck), Coleophora laricella Hbn., Exoteleia pinifoliella 
(Chamb.). 
Biology: Schaffner, 1959, loc. cit. supra, p. 91. — Allen, 1962, loc. cit, 
supra, p. 109. 

Genus HOCKERIA Walker 
( = Sfomafoceras Kirby of Catalog, p. 585) 

Hockeria Walker, 1834. Ent. Mag. 2: 21, 34. 

Type : Chalcis bispinosa Fabricius. Desig. by Kirby, 1883. 

Taxonomy: Boucek, 1951. Mus. Nat. Pragae Acta Ent. 27, Sup., p. 61. — 
Steffan, 1953. Feuille Nat. (Cahiers Nat.), n.s., 8: 10.— Habu, 
1960. Nat. Inst. Agr. Sci. Japan Bui., ser. C, 11: 220. 

Use of this generic name is provisional, to conform with preponderant 
usage in current literature. Since the genotype of Stomatoceras is a misiden- 
tified species, the case should be referred to the Internatl. Comm. on Zool. 
Nomencl. 

americensis (Girault) , new status. 

Stomatoceras unipunctatipennis americensis Girault, 1918 (ch. from 
subsp. to sp.; types re-examined). 

eriensis (Wallace), new status. Del. south to N. C., west to N. Dak., Kans., 
Tex., Calif. 



CHALCIDroAE 277 

Siomatoceras rubra var. eriensis Wallace, 1942 (ch. to sp. by Steffan, 
1959. Vie et Milieu 10: 304). 

rubra (Ashmeacl), new status. Calif., Mexico. Host: Harrisina brillians B. & 
McD. Ch. from subsp. to sp. by Steffan, 1959, loc. cit. supra, p. 
304. 

tenuicomis (Girault). Calif. 

unipunctatipennis (Girault), new status. N. Y. south to Fla., west to Oreg., 
Calif. Hosts: Neocliprion spp. 
Stoniatoceras unipunctatipennis unipunctatipennis of Catalog (ch. from 
subsp. to sp.; types re-examined). 

Genus SCHWARZELLA Ashmead, p. 585 

The female of this genus has not been identified. 
arizonensis Ashmead. N. Mex., Calif. 

Subfamily BRACHYMERIINAE 
Taxonomy: Steffan, 1959. Mus. Nat. Pragae Acta Ent. 33: 287-290. 

*Tribe CRATOCENTRINI 

Genus ACANTHOCHALCIS Cameron, p. 585 

Revision: Steffan, 1959, loc, cit. supra, pp. 306-309. 

ACANTHOCHALCIS subgenus ACANTHOCHALCIS Cameron 

Changed from genus to subgenus, by Steffan, 1959, loc. cit. supra, p. 307. 

nigricans Cameron. Kans., Okla. ; Mexico. 

^ACANTHOCHALCIS subgenus BELOCHALCIS StefFan 

Acanthochalcis subg. Belochalcis Steffan, 1959. Mus. Nat. Pragae Acta 
Ent. 33: 308. 

Type: Acanthochalcis unispinosa Girault. By monotypy. 

unispinosa Girault. Ariz. 

♦Tribe PHASGONOPHORINI 

Genus PHASGONOPHORA Westwood, p. 585 
sulcata Westwood. N. H. 

Genus TRIGONURA Sichel, p. 586 

*Bactrochalcis Kieffer, 1912. Soc. Ent. de France Ann. 80 : 463. 

Type : Bactrochalcis reticulata Kieffer. By monotypy. 
Revision: Burks, 1959. Ent. Soc. Amer. Ann. 52: 75-81. 

*algerti Burks. Tex., Ariz., Calif. Hosts: Chrysobothris piuta Wickham, C. 
octocola Lee. 
Trigonura algerti Burks, 1959. Ent. Soc. Amer. Ann. 52: 77. $, $. 

californica Rohwer. Utah, Ariz., Oreg. Host: Chrysobothris femorata (Oli- 
vier). 

elegans (Provancher) . Conn., Ont., N. Y., Pa., W. Va., Ark. Hosts: Agrilus 
sp., Chrysobothris femorata (Oliv.), C. sexsignata (Say), Mag- 
dalis armicollis (Say), M. barbita (Say), M. olyra (Hbst.), ? Sco- 
lytus sp. 

Trigonura hicoriae Rohwer, 1919 (syn. by Burks, 1959, loc. cit. supra, p. 
78). 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1260 

hicoriae Rohwer. Syn. of elegans (Provancher). 



278 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*insularis (Cresson). Fla.; "West Indies, 

Phasgonophora iitsularis Cresson, 1865. Ent. Soc. Phila. Proc. 4: 102. 9, 
$. 
*pini Burks. B. C, Oreg., Calif. 

Trigonura pini Burks, 1959. Ent. Soc. Amer. Ann. 52: 80. ?, $. 

larsata (Dalla Torre). N. B., Vt., Mass., Ont., Pa., N. C, Ohio. Hosts: 
Magdalis armicollis (Say),M. barbita (Say). 

*iilmi Burks. Que., Ont., N. Y., N. J., Pa., N. C, 111., Iowa. Hosts: Magdalis 
armicollis (Say), Scolytus quadrispinosus Say., Scolytus sp. 
Trigonura ulnii Burks, 1959. Ent. Soc. Amer. Ann. 52: 80. $, $. 

*Tribe BRACHYMERIINI 

Genus BRACHYMERIA Westwood, p. 586 

Revision: Burks, 1960. Amer. Ent. Soc. Trans. 86: 225-273. 

BRACHYMERIA subgenus BRACHYMERIA Westwood 

Ch. from gen. to subg. by Habu, 1960. Nat. Inst. Agr. Sci. Japan Bui., 
Ser. C, 11, p. 144. 

*aeca Burks. N. S., Que., Maine, south to Fla., west to B. C. and N. Mex. 
Hosts: Chaetophlepsis sp., prob. other Tachinidae parasitic on 
Lepidoptera larvae, poss. also Tenthredinidae. 
Brachymeria (Brachymeria) aeca Burks, 1960. Amer. Ent. Soc. Trans. 
86: 242. 9, $. 

belfragei (Crawford). Tex.; Mexico, Honduras, Panama. 

coloradensis (Cresson). N. H. and Ont., south to Fla., west to B. C. and 

Calif.; Holarctic. Hosts: Sarcophaga kellyi Aid., S. hunteri 

Hough, par. on grasshoppers. 
Chalcis tachinae Howard, 1885 (syn. by Burks, 1960, loc. cit. supra, p. 

251). 
Biology: Smith, 1957. Canad. Jour. Res. 36: 258.— Spencer, 1958. 10th 

Internatl. Ent. Cong. Proc. 4 : 497. 

*comitator (Walker). Fla., Tex.; Mexico south to Colombia and Brazil. 
Hosts: Ceratnidia butleri (Mos.), C. viridis (Druce), C. sp., Alo/- 
baina argillacea (Hbn.), Pieris sp. 
Chalcis comitator Walker, 1862. Ent. Soc. London Trans., ser. 3, 1: 350. 

compsilurae (Crawford). Maine, N. H., and Ont., south to N. C. and Tenn.; 
Mich., 111., Wis., Minn., Man., Mont., south to Colo, and N. Mex., 
B. C, Oreg. Hosts: Rileymyia fulvipes (Bigot), other Tachinidae 
par. on Lepidoptera. 

hammari (Crawford). Ark., Kans., Tex. west to Oreg. and Calif.; Mexico. 
Hosts: Argyrotaenia citrana (Fern.), Carpocapsa pomonella (L.), 
Platynota stultana (Wlshm.), Psorosina hammondi (Riley), 
tCremastus cookii Weed. 
Biology: Allen, 1962. U. S. Dept. Agr. Tech. Bui. 1265, p. 110. 

fonscolombei (Dufour). Worldwide in warm-temperate zone. 

♦intermedia (Nees). Mass., Conn.; S. Europe, Near and Middle East. Hosts: 
Pupae of many Lepidoptera. 
Chalcis intermedia Nees, 1834. Hym. Ichn. affin. Monog., v. 2, p. 29. $. 

Distribution: Burks, 1960. Ent. News 71: 62. 

incerta (Cresson). Fla.; West Indies, Mexico. Hosts: Alabama argillacea 
(Hbn.), Celerio lineata (Fab.), Heliothis zea (Bod.), Laphygma 
frugiperda (Sm.), Lymire edwardsi (Grote), Megalopyge kr-ugii 
Dew., M. opercidaris (A. & S.), Neonympha sp., Olketicus abbotii 
Grote, Papilio cresphontes Cram., Mods latipes (Guen.). 

louisianae (Girault). Syn. of coluinbiana (Howard) in subg. Pseudobrachym- 
eria. 



CHALcmroAE 279 

*mo!estae Burks. Md. south to Fla., west to Ind., Tenn. and Miss. Hosts: 
Grapholitha molesta (Busck), Anchylopera platanana Clem., Aro- 
galea cristifasciella (Chamb.). 
Brachymeria (Brachymeria) molestae Burks, 1960. Amer. Ent. Soc. 

Trans. 86: 245. $, $. 

ovata (Say), changed status. Ont. ; U. S. south of Maine, N. H., and Mont. 

Hosts: Over 100 spp. of Lepidoptera, from Psychidae to Papilioni- 

dae. Raised from subsp. to sp. rank by Burks, 1960, loc. cit. 

supra, p. 258. 
Chalcis ovata var. ahiesae Girault, 1917 (syn. by Burks, 1960, loc. cit. 

supra, p. 259). 
Biology: Genung, 1959. Fla. Ent. 42: 40.— Schaffner, 1959, loc. cit. 

supra, p. 91. — Allen, 1962, loc. cit. supra, pp. 110-111. 

ovata ahiesae Girault. Syn. of ovata Say. 

ovata ovata (Say). Raised to sp. rank. 

robusta (Cresson), corrected spelling of rohustus of Catalog, teste Genung, 
1958. Fla. Ent. 41: 191. 
Biology: Genung, 1959. Fla. Ent. 42: 40. 

*russelli Burks. Miss., Tex., N. Mex., Ariz.; Mexico. Hosts: Coptocycla 
( Psalidonota) texana (Schaeffer), Chelymorpha cassidea (Fab.), 
Niizonia sp. 
Brachymeria (Brachymeria) russelli Burks, I960. Amer. Ent. Soc. 
Trans. 86: 240. 9, $. 

slossonae (Crawford.) Fla. Host: Ephyriades brunnea floridensis B. & C. 

tachinae (Howard). Syn. of coloradensis (Cresson). 

tegiilaris (Cresson). 111. and Tenn. west to Wash., Oreg., and Calif. Host: 
Hemithrixion oestriforme B. B. 
Biology: Smith, 1957. Canad. Jour. Zool. 36: 259.— Spencer, 1958. Int. 
Cong. Ent. 10 Proc, v. 4, p. 501. 

*truncata Burks. Syn. of truncatella Burks. 

*truncatella Burks, new name. Sask., Nebr., Tex., Colo., N. Hex., Ariz., 
Calif. 
*Brachymeria (Brachymeria) truncata Burks, 1960. Amer. Ent. Soc. 
Trans. 86: 234. ?, $. Preocc. by Brachymeria truncata 
(Schmitz), 1946. 

*weem8i Burks. Fla. 

Brachymeria (Brachymeria) tveemsi Burks, 1960. Amer. Ent. Soc. 
Trans. 86: 236. 9. 

*BRACHYMERIA subgenus GAHANULA Burks 

Brachymeria subg. Gahanula Burks, 1960. Amer. Ent. Soc. Trans. 86: 
261. 

Type: Brachymeria discreta Gahan. Orig. desig. 

discrela Gahan. Tex., Ariz., Calif., Hawaii; Mexico. Hosts: Tinea sp., 
? pyi-austid moth infesting Polistes instabilis Sauss. nest. 

discretoidea Gahan. Tex., Ariz.; Mexico, Panama. Host: Unident. moth in- 
festing Trig ana amalthea (Oliv.) nest. 

^BRACHYMERIA subgenus PSEUDOBRACHYMERIA Burks 

Brachymeria subg. Pseudobrachymeria Burks, 1960. Amer. Ent. Soc. 
Trans. 86: 263. 

Type: Pseudochalcis conica Ashmead. Orig. desig. 

Columbiana (Howard). D. C, Va., Fla., La. Host: ? Laetilia coccidivora 
Comst. 
Chalcis louisianae Girault (syn. by Burks, 1960, loc. cit. supra, p. 266). 

*conica (Ashmead). Tex.; south through Cent. Amer. to Brazil; Trinidad, 
W. I. 



280 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Pseudochalcis conica Ashmead, 1904. Carnegie Mus. Mem. 1: 407. 9. 

*flegiae Burks. Fla., Tex. Host: Palpita flegia (Cram.). 

Brachymeria ( Pseudobrachymeria) flegiae Burks, 1960. Amer. Ent. Soc. 

Trans. 86: 271. ?, $. 

*Iaetiliae Burks. N. Car., Okla., N. Hex., Ariz., Calif. Hosts: Laetilia cocci- 
divora Comst., L. zamacrella Dyar. 
Brachymeria (Pseudobrachymeria) laetiliae Burks, 1960. Amer. Ent. 
Soc. Trans. 86: 268. 9, S. 

pedalis (Cresson). Tex., Colo., N. Mex., Ariz., Calif. Hosts: Noctuelia elau- 
talis (Grote), Melitara prodenialis Walk., M. dentata Grote, Oza- 
mia clarefacta Dyar. 

*psyche Burks. Fla. Host: Oiketicus abbotii Grote. 

Brachymeria (Pseudobrachymeria) psyche Burks, 1960. Amer. Ent. Soc. 
Trans. 86: 267. $. 

Subfamily DIRHININAE 

Genus DIRHiNUS Dalman, p. 588 

*Dirrhinoidea Girault, 1912. Arch. Naturg. 78 (A) 9: 165. 

Type: Dirrhinoldea maculata Girault. Orig. desig. 
*Eniacella Girault, 1913. Wis. Nat. Hist. Soc. Bui. 11: 35. 

Type: Eniacella ruficornis Girault. Orig. desig. 
Taxonomy: Habu, 1960, loc. cit. supra, p. 309. 

lexanus (Ashmead). Ariz., Calif . 

Dirhinus paoli Burks, n. syn.; addl. mat. shows it to be in range of var. 
of texanus (Ashmead). 

paoli Burks. Syn. of texanus (Ashmead). 

Subfamily CHALCIDINAE 

Genus METADONTIA Ashmead, p. 588 

amoena (Say). Md. south to Fla. west to 111., Mo., Kans., Tex., N. Mex., 
Ariz., Calif.; Mexico, Cent, and S. America. 

Genus CHALCIS Fabricius, p. 588-589 

barbara (Cresson). Mich. In ref. to orig. descr. ch. $ to "$" = $. 

canadensis (Cresson). 

Biology: Judd, 1953. Amer. Midi. Nat. 49: 817. 

divisa (Walker). Calif.; Mexico. 

Morphology: Daly, 1963. Ent. Soc. Amer. Ann. 56: 303. 

lasia Burks. Ariz. 

megalomis Burks. Oreg. Host: Stratiomys sp. 

phoenicapoda Burks. 

Nomenclature: Peck, 1963, loc. cit. supra, p. 869 (rufipes Kirby). 

Genus SPILOCHALCIS Thomson, pp. 589-592 

albifrons (Walsh). 

Biology: Poinar and Gyrisco, 1963. Jour. Econ. Ent. 56: 534. 

dema Burks. Pa., Calif.; Mexico. 

mariae (Riley), corrected spelling of maria (Riley) of Catalog, teste Peck, 
1963, loc. cit. supra, p. 881. 

nortonii (Cresson). Conn. 

Biology: Schaflfner, 1959, loc. cit. supra, p. 91. 

odontotae Howard. Fla.; Mexico. 

phoenica Burks. Ariz., Calif. ; Mexico. 



LEUCOSPIDIDAE 281 



side (Walker). 

Biology: Arthur, 1958. Canad. Ent. 90: 590-595.— Schaflfner, 1959, loc. 
cit. supra, p. 91. — Allen, 1962, loc. cit. supra, p. 111. 

xanthostigma (Dalman). Delete; extralimital . Ont. specimen det. as this is 
albifrons (Walsh) ; specimen re-examined. 

Genus CERATOSMICRA Ashmoad, pp. 592-593 
debilis (Say). Md., Ariz., Calif.; Mexico, West Indies. 

Family LEUCOSPIDIDAE 

Genus LEUCOSPIS Fabricius, p. 593 

Taxonomy: Burks, 1961. Stud. Ent. Rio de Janeiro 4: 540.— Habu, 1962. 
Fauna Japonica, Chalc, Leucosp., and Podagr., pp. 165-170. 

affinis affinis Say. Hosts: Anthidium maculosum Cress., Ashmeadiella ari- 
dula Ckll., Megachile inermis Prov., M. relativa Cress., M. gentils 
Cress. Stelis sp., Hoplitis producta (Cress.), Osmia kincaidii Ckll. 

Biology: Medler, 1958. Ent. News 69: 21-24.— Medler and Koerber, 
1958. Ent. Soc. Amer. Ann. 51: 340-344.— Medler, 1958. Canad. 
Ent. 90: 326-327. 

Taxonomy: Michener, 1954. Univ. Kans. Sci. Bui. 35: 1743 (subsp. ch. to 
sp.). 

affinis floridana Cresson. Delete Mass. from dist.; spec, discolored by cyanide. 
Hosts: Megachile (Chelostomoides) georgica Cress., M. (Sayapis) 
sp. 
Nomenclature: Peck, 1963, loc. cit. supra, p. 894 (subsp. ch. to var.). 

birkmani Brues. Fla., Ariz., Calif; Mexico. 

robertsoni Crawford. Ala. 

slossonae Weld. Host: Anthidiellum sp. 

texana Cresson. Ala. 

Unplaced Genera, p. 593 

Lophocomodia Ashmead, 1888. Transf. to Sphegigasterini following Heteros- 
chema Gahan, p. 543. 

Ptertuomus Packard. Transf. to Mymaridae as syn. under Anagms Haliday, 
p. 415. 

Unplaced Species, p. 594 

Lophocomodia americana Ashmead. Transf. to Sphegigasterini, p. 543. 

Diomorus biorhizae Ashmead. Transf. from Diomorus Walker, p. 527. 
Type lost; species is apparently a Torymus. 

Halticoptera brodiei Ashmead. Transf. from Halticoptera Spinola, p. 542. 

Type lost, but original description clearly not of a Halticop- 
tera. 

Coccophagus brunneus Provancher. 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1257. 

Camptoptera clavata Provancher. 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1256. 

Semiotellus fasciatus Provancher. Transf. to Arthrolytus Thomson, p. 557. 

*Solenotus fletcheri Cockerell, 1889. Colo. Biol. Assn. Rpt. 6: [2]. 

Entedon imbrasus Walker. Transf. to Chrysocharis Foerster, p. 463. 

Perilampus laevis Provancher. Transf. to Elatus Walker, p. 519. 

Semiotellus minimus Provancher. 

Taxonomy: Burks, 1964, loc. cit. supra, p. 1262. 



282 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Meraporus monticola Ashmead, 1889. Colo. Biol. Assn. Rpt. 3 : [4]. 

Pteromalus nigricornis Provancher. 

Taxonomy : Burks, 1964, loc. cit. supra, p. 1260. 

Semiotellus oblongus Provancher. Transf. to Tridymus Ratzeburg, p. 546. 

Rhopalicus pallipes Provancher. Transf. from Rhopalicus Foerster teste 
Burks, 1964, loc. cit. supra, p. 1261. 

Eulophus ramosus Provancher. 

Taxonomy : Burks, 1964, loc. cit. supra, p 1258. 

Superfamily CYNIPOIDEA 
Family IBALIIDAE 

Genus IBALIA Latreille, pp. 594-595 

ensiger Norton. 

Biology: Wickman, 1964. Pan-Pacific Ent. 40: 19. 

Family CYNIPIDAE 

Taxonomy: Weld, 1957. Cynipid Galls of the Pacific Slope, 80 pp., Ann 
Arbor, Mich.— Weld, 1959. Cynipid Galls of the Eastern U. S., 
156 pp., Ann Arbor, Mich.— Weld, 1960. Cynipid Galls of the 
Southwest, 52 pp., Ann Arbor, Mich. (All privately printed.) 

TRYBLIOGRAPHA subgenus DIDICTYUM Riley 
IzzTrybliographa subg. Hexaplasfa Foerster of Catalog, p. 603) 

Taxonomy: Weld, 1962 (1961). Ent. Soc. Wash. Proc. 63(4) : 280. When 
Hexaplasta Foerster M^as made a synonym of Hexacola Foerster 
the name Didictyuvi was revived for Hexaplasta of authors and 
as a subgenus in Trybliographa. 

zigzag Riley. Fla. Host: Megaselia eletiae (Comstock). In reference to o.d., 
add pp. 214, 500, and Amer. Ent. 3: 52, 1880. 
Taxonomy : Weld, 1962, loc. cit. supra, p. 280. 

TRYBLIOGRAPHA subgenus TRYBLIOGRAPHA Foerster, pp. 603-604 

rapae (Westwood). Hosts: Hylemyia spp. 

Biology: Wishart and Monteith, 1954. Canad. Ent. 56: 145-154, 14 figs, 
(early stages). 

Genus SYNERGUS Hartig, pp. 610-613 

*inagnificus Weld. Calif. Reared from galls of Disholcaspis truckeensis 
(Ashmead). 
Synergus magnificus Weld, 1957. U. S. Natl. Mus. Proc. 107: 107. 9, $. 

Genus CEROPTRES Hartig, p. 614 

""montensis Weld. Calif. Reared from galls of Andricus reniformis McCrack- 
en and Egbert. 
Ceroptres montensis Weld, 1957. U. S. Natl. Mus. Proc. 107: 108. 9, $. 

Genus NEUROTERUS Hartig, pp. 620-624 

*floricomus Weld. Ariz. Gall on Quercus reticulata. 

ISeuroterus floricomus Weld, 1957. U. S. Natl. Mus. Proc. 107: 109. 9, 
5,0. 

^lamellae Weld. Ariz. Gall on Quercus subturbinella. 

ISeuroterus lamellae Weld, 1957. U. S. Natl. Mus. Proc. 107: 109-110. ?, 0. 

'^'florulentas Weld. Calif. Gall on Quercus dumosa and Q. lobata. 



CYNIProAE 283 

Neuroterus florulentus Weld, 1957. U. S. Natl. Mus. Proc. 107: 110. 9, 
5,0. 

Genus LIODORA Foersfer, p. 629 

*dumosae Weld. Calif. Gall on Quercus dumosa. 

Liodora dumosae Weld, 1957. U. S. Natl. Mus. Proc. 107: 111. 9, $, 0. 

Genus AMPHIBOLIPS Reinhard, p. 630-631 

*murata Weld. Fla. Gall on Quercus myrtifolia, Q. pumila, Q, laurifolia, and 

O CITICTCCI* 

Amphibolips murata Weld, 1957. U. S. Natl. Mus. Proc. 107: 111-112. 
9,0. 

Genus ANDRICUS Hartig, pp. 631-637 

crystallinus Bassett. Calif. 

Biology: Doutt, 1960. Pan-Pacific Ent. 36(5) : 167-170, 9, 5, (describes 
the alternating bisexual generation on the leaves of Quercus dou- 
glasii). 

foliaformis Gillette. Ariz. Gall on Quercus hicolor, Q. macrocarpa. 
Andricus foliosus Weld, 1926. N. syn. 

*nienclocinensis Weld. Calif. Gall on Quercus densiflora. 

Andricus mendocinensis Weld, 1957. U. S. Natl. Mus. Proc. 107: 112. 9, 0. 

Genus DISHOLCASPIS Dalla Torre and KiefFer, pp. 638-640 

*edura Weld. Ariz. Gall on Quercus oblongifolia and Q. arizonica. 

Disholcaspis edura Weld, 1957. U. S. Natl. Mus. Proc. 107: 113. 9, 0. 

*mamilIanaWeld. Calif. Gall on Quercus douglasii. 

Disholcaspis mamillana Weld, 1957. U. S. Natl. Mus. Proc. 107: 115. 9, 0. 

*mellifica Weld. Calif. Gall on Quercus garryana. 

Disholcaspis mellifica Weld, 1957. U. S. Natl. Mus. Proc. 107: 114. 9, 0. 

*prehensa Weld. Calif. Gall on Quercus dumosa. 

Disholcaspis prehensa Weld, 1957. U. S. Natl. Mus. Proc. 107: 113. 9, 0. 

*spissa Weld. Ariz. Gall on Quercus subturbinella, Q. arizonica, Q. harardi. 
Disholcaspis spissa Weld, 1957. U. S. Natl. Mus. Proc. 107: 114. 9, 0. 

Genus ANTRON Kinsey, pp. 640-641 

*magdalenae Weld. N. Mex. Gall on Quercus subturbinella. 

Antron magdalenae Weld, 1957. U. S. Natl. Mus. Proc. 107: 116. 9, 0. 

Genus LOXAULUS Mayr, p. 643 

*beuteiunuelleri Weld. N. J., Pa., 111. Gall on Quercus rubra, northern red 
oak. 
Loxaulus beutenmuelleri Weld, 1957. U. S. Natl. Mus. Proc. 107: 
116-117. 9. 

Genus DRYOCOSMUS Giraud, pp. 645-646 

dubiosus (Fullaway). Calif. Galls on Quercus agrifolia. 

Dryocosmus hicornis McCracken and Egbert (syn. by Doutt, 1959. Ent. 
Soc. Amer. Ann. 52(1): 69-74. 9, 0). D. hicornis is alternating 
agamic generation on the leaves. 

Genus HETEROECUS Kinsey, pp. 646-647 

pacificus (Ashmead). 

Biology: Lyon, 1963. Ent. Soc. Wash. Proc. 65(3): 250-254, figs. A, B, 
C, D. (Describes the alternating bisexual generation on the 
leaves. 9, 5,0.) 

Genus CALLIRHYTIS Foersfer, pp. 647-653 

agrifoliae (Ashmead). 



284 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Biology: Lyon, 1964. Ent. Soc. Wash. Proc. 66(3): 193. (Describes the 
alternating bisexual generation on Quercus agrifolia. ?, $, 0.) 

*balanella Weld. Ariz. Gall on Quercus emoryi. 

Callirhytis balanella Weld, 1957. U. S. Natl. Mus. Proc. 107: 117. ?, 0. 

*floripara Weld. Calif. Gall on Quercus agrifolia. 

Callirhytis floripara Weld, 1957. U. S. Natl. Mus. Proc. 107: 118. 9, $,0. 

*glomerosa Weld. Va. Gall on Quercus coccinea. 

Callirhytis glomerosa Weld, 1957. U. S. Natl. Mus. Proc. 107: 119. $, 0. 

*intersita Weld. Va. Taken ovipositing in internodes of new growth of Qu- 
erents alba. 
Callirhytis intersita Weld, 1957. U. S. Natl. Mus. Proc. 107: 119. 9. 

*medularis Weld. Pa., Va., Ohio. Gall on Quercus velutina and Q. coccinea. 
Callirhytis medularis Weld, 1957. U. S. Natl. Mus. Proc. 107: 121. 9,5,0. 

*perob8cura Weld. Va. Ovipositing in buds of Quercus velutina. 

Callirhytis perobscura Weld, 1957. U. S. Natl. Mus. Proc. 107: 122. 9. 

pomifomiis (Ashmead). 

Biology: Lyon, 1959. So. Calif. Acad, Sci. Bui. 58(1) : 33-37. (Describes 
the alternating bisexual generation on the leaves of Quercus agri- 
folia. 9, 5, 0.) 

Genus Undetermined, p. 653 

Oxylabis hifoveolata Brues, transf. from Diapriidae, p. 681 {teste L. Masner, 
who has examined the type in Amer. Mus. Nat. Hist. coll.). 

R/iorfifes /usi/ormans Cockerell, 1890. Entomologist 23 : 75. only. 

Rhodites globosus Cockerell, 1890. Entomologist 23 : 75. only. 

Rhodites spinosellus Cockerell. Catalog p. 654. Delete Norn. nud. 

♦Superfamily EVANIOIDEA 

Taxonomy: Short, 1959, Ent. Monthly Mag. 95 (1145) : 217-219 (larvae). 
— Crosskey, 1962. Roy. Ent. Soc. London, Trans. 114 (Pt. 12): 
399-400 (classification). 

Morphology, Biology, Taxonomy: Crosskey, 1951. Roy. Ent. Soc. Lon- 
don, Trans. 102 (5) : 247-301. 

Family EVANIIDAE 

Taxonomy and Biology: Crosskey, 1951. Roy. Ent. Soc. London, Trans. 
102 (5) : 285-286. 

Genus EVANIELLA Bradley, p. 656 

calif ornica (Ashmead). 

Taxonomy: Snelling, 1963. Pan-Pacific Ent. 39 (2): 107-108 (9 descr.). 

neomexicana (Ashmead). Calif. 

Family AULACIDAE 

{^Aviac'mae of Catalog) 

Taxonomy and Biology: Crosskey, 1951. Roy. Ent. Soc. London, Trans. 

102 (5) : 284-285. 
Morphology: Crosskey, 1951. Roy. Ent. Soc. London, Trans. 102 (5): 

247-281. 

Genus PRISTAULACUS Kieffer, Sup. 1, p. 88 
( = Ao/acos»e»/»us Philippi, pp. 657-659) 

ater (Westwood). Minn. 



GASTERUPTIIDAE 285 

rufitarsis (Cresson). Alaska. 

Genus AULACUS Jurine, pp. 659-660 
dispilus Townes. Host : Reared from Celtis laevigata. 
lovei (Ashmead). Host: Xiphydria tibialis Say. 

Family GASTERUPTIIDAE 

(^=zCasteruptiinae of Catalog) 

Revisions: Pasteels, 1956. Mus. Congo Beige, Ann., Sci. Zool., (Ser, 8) 
50: [l]-94 (No. Africa) .—Pasteels, 1957. Inst. Roy. des Sci. Nat. 
de Belg. Mem. (ser. 2) 56: [1]-125 (Australia).— Pasteels, 1957. 
Soc. Roy. d'Ent. Belg., Bui. et Ann. 93 (5-6) : 173-176 (New Zea- 
land) .—Pasteels, 1958. Op. cit. supra 94 (7-8) : 169-213. 

Taxonomy: Short, 1959. Ent. Monthly Mag. 95 (1145) : 219. 

Genus GASTERUPTION Latreille, p. 660 
assectator assectator (Linnaeus). Alaska. 
kirbii kirbii (Westwood) . Mich. 
kirbii russeum Townes. Host: Hoplitis sambuci Titus. 

Genus RHYDINOFOENUS Bradley, p. 661 

barnstoni bamstoni (Westwood). Alaska. 

floridanus bradleyi Townes. Hosts: Hylaeus cressoni (Ckll.) ; Trypoxylon 
frigidum Smith. 

pattersonae (Melander and Brues). Host: Dianthidium curvatum sayi Ckll. 

tarsatorius tarsatorius (Say). N. C. 

*Superfamily PELECINOIDEA 

Insert in Catalog before Proctotrupoidea, p. 655. 

Taxonomy: Handlirsch, 1933. Handbuch der Zoologie, v. 4 (2), Insecta 
2, p. 981. 

Family PELECINIDAE 

Remove from Proctotrupoidea, pp. 661-662 and place in superfamily 
Pelecinoidea before Proctotrupoidea, p. 655. 

Superfamily PROCTOTRUPOIDEA 

The families Evaniidae and Gasteruptiidae are removed to the superfa- 
mily *Evanioidea and the family Pelecinidae to the superfamily *PeIecinoi- 
dea. 

Morphology: Pschorn-Walcher, 1955. Schweiz. Ent. Gesell. Mitt. 28: 

214-219. 
Phylogeny: Bradley, 1955. Soc. Roy Ent. de Belg. Mem. 27: 129-131.— 

Bradley, 1958 (1956). Tenth Internatl. Congr. Ent. Proc. 1: 267. 

Family VANHORNIIDAE 

Genus VANHORNIA Crawford, p. 662 
eucnemidarum Crawford. Maine, N. H., Conn. 



286 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Family ROPRONIIDAE 

Genus ROPRONIA Provancher, p. 662 

Taxonomy: Yasumatsu, 1956. Insecta Matsumurana 19: 117-122. — Ya- 
sumatsu, 1958. Insecta Matsumurana 21: 112-114. — Heqvist, 
1959. Ent. Tidskr. 80: 137-139. 

Biology: Chao, 1957. Fukien Agr. Jour. 5: 73-76 {Ropronia recorded as 
parasitic on sawfly larvae — first record of host for the genus). 

Distribution: Yasumatsu, 1961. Mushi 35: 68-69. 

Family HELORIDAE 

Taxonomy: Riek, 1955. Austral. Jour. Zocl. 3: 258-265. — Pschorn- 
Walcher, 1955. Schweiz. Ent. Gesell. Mitt. 28: 233-250 (revision 
of Palearctic species). 

Morphology: Oeser, 1961. Berlin Zool. Mus. Mitt. 37: 72-73 (ovipositor). 

Genus HELORUS Latreille, p. 663 
paradoxus (Provancher). N. B. 

Family PROCTOTRUPIDAE 

Taxonomy: Riek, 1955. Austral. Jour. Zool. 3: 106-117 (revision of the 
Australian species). — Pschorn-Walcher, 1958. Beitr. z. Ent. 8: 
724-731.— Pschorn-Walcher, 1958. Schweiz. Ent. Gesell. Mitt. 31: 
57-64. — Masner, 1961. Pare Nat. Upemba Miss, de Witte, fasc. 
60: 37-47 (key to the genera of the world) . 

Morphology: Oeser, 1961. Berlin Zool. Mus. Mitt. 37: 71-72 (ovipositor). 

Genus PROCTOTRUPES Latreille, pp. 663-664 

caudatus Say. Pa., D. C. 

cockerelli (Brues), n. comb. Transf. from Phaenoserphus Kieffer, p. 664 
{teste L. Masner upon examination of type). 

pallidas (Say). Idaho. 

rufigaster Provancher. N. H. 

Genus PHAENOSERPHUS KiefFer, p. 664 
cockerelli (Brues). Transf. to Proctotrupes Latreille, pp. 663-664. 
obscuripes (Brues). Transf. to Codrus Panzer, pp. 665-666. 
placidus (Brues). Transf. to Codrus Panzer, pp. 665-666. 
serricornis (Brues). Transf. to Codrus Panzer, pp. 665-666. 
simplicior (Brues). Transf. to Codrus Panzer, pp. 665-666. 

Genus CRYPTOSERPHUS Kieffer, pp. 664-665 

^Brachyserphus Hellen, 1941. Notulae Ent. 21: 42. N. syn (teste L. 
Masner). 

Type : Codrus parvulus Nees. By orig. desig. 

abruptus (Say). N. H., N. J., Md., W. Va., S. C. 

flavipes (Provancher). N. B.; N. C. 

obsoletus (Say). Fla. 

Genus CODRUS Panzer, pp. 665-666 

longiceps (Ashmead). Ark. 

obscuripes (Brues), n. comb. Transf. from Phaenoserphus Kieffer, p. 664 
(teste L. Masner upon examination of type). 



CERAPHRONIDAE 287 

placidus (Brues), n. comb. Transf. from Phaenoserphus Kieffer, p. 664 {teste 
L. Masner upon examination of type). 

serricornis (Brues), n. comb. Transf. from Phaenoserphus Kieffer, p. 664 
(teste L. Masner upon examination of type) . 

simplicior (Brues), n. comb. Transf. from Phaenoserphus Kieifer, p. 664 
(teste L. Masner upon examination of type). 

Family CERAPHRONIDAE 

Taxonomy: Dessart, 1962. Soc. Roy, Ent. de Belg. Bui. et Ann. 98: 
305-309 (key to the genera of the world). 

Genus CERAPHRON Jurine, pp. 666-668 

Allomicrops Kieffer, 1914. Syn. by Dessart, 1963. Soc. Roy. Ent. de 

Belg. Bui. et Ann. 99: 524. 
*Eulagy nodes Girault, 1917. New Javanese Hymenoptera, p. 9 (Private 
pub.). Syn. by Masner, 1964. Acta Soc. Ent. cechosl. 61: 126. 

Type: Eulagynodes bicolor Girault. By monotypy and orig. 
desig. 

abnormis Perkins. Transf. from Allomicrops Kieffer, teste Dessart, 1963. 
Soc. Roy. Ent. de Belg. Bui. et Ann. 99: 523. 
*Eulagynodes bicolor Girault, 1917. New Javanese Hymenoptera, p. 9 
(private pub.). ?. Syn. by Masner, 1964. Acta Soc. Ent. Cechosl. 
61: 126. 

longicornis Ashmead. Transf. to Lagynodes Foerster, p. 669. 

salicicola Ashmead. Transf. to Aphanognius Thomson, p. 668. 

Genus APHANOGMUS Thomson, p. 668 

Taxonomy: Dessart, 1962. Soc. Roy. Ent. de Belg. Bui. et Ann. 98 
294-300.— Dessart, 1963. Soc. Roy. Ent. de Belg. Bui. et Ann. 99 
387-416.— Dessart, 1964. Soc. Roy. Ent. de Belg. Bui. et Ann. 100 
259-278. 

floridanus Ashmead. Ohio. Host: Mycodiplosis sp. 

salicicola (Ashmead), n. comb. Transf. from Ceraphron Jurine, p. 667 (teste 
L. Masner). 

Genus A/fomicrops Kieffer, p. 669 
Syn. of Ceraphron Jurine, supra. 
abnormis (Perkins). Transf. to Ceraphron Jurine, p. 666. 

Genus LAGYNODES Foerster, p. 669 

longicornis (Ashmead), n. comb. Transf. from Ceraphron Jurine, p. 667 
(teste L. Masner). 

Genus ATRITOMELLUS KiefFer, p. 669 

conwentziae (Gahan). Transf. from Dendrocerus Ratzeburg, p. 672 (teste 
Masner, 1964. Acta. Soc. Ent. Cechosl. 61: 126). 

conwentziae var. rufus (Gahan). Transf. from Dendrocerus Ratzeburg, p. 
672 (teste Masner, loc. cit. supra). Calif. 

Genus LYGOCERUS Foerster, pp. 669-670 

Biology: Muesebeck, 1959. Ent. News 57: 91. 

anomaliventris (Ashmead), n. comb. Transf. from Conostigmus Dahlbohm, p. 
670 (teste L. Masner). 

'''attentus Muesebeck. Maine. Hosts: Aphidiinae in Macrosiphum euphorbiae 
(Thomas). 
Lygocerus attentus Muesebeck, 1959. Ent. News 57: 94. 9, $. 



288 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

♦incompletus Muesebeck. Maine, Hosts: Aphidiinae in Macrosiphum euphor- 
biae (Thomas). 
Lygocerus incompletus Muesebeck, 1959. Ent. News 57: 96. 9, $. 

*latifrons Muesebeck. Calif. Host: Species of Hemerobiidae. 

Lygocerus latifrons Muesebeck, 1959. Ent. News 57: 93. ?, 5. 

*leucopidis Muesebeck. Calif. Host: Leucopis sp. 

Lygocerus latifrons Muesebeck, 1959. Ent. News 57: 93. $, $. 

*pinicola Muesebeck. Calif. Host: Leucopis atri fades Aldrich. 
Lygocerus pinicola Muesebeck, 1959. Ent. News 57: 92. ?, $. 

Genus CONOSTIGMUS Dahlbom, pp. 670-671 

Conostigmoides Dodd, 1914. Syn. by Masner, 1964. Acta Soc. Ent. 
Cechosl. 61: 124. 

anomaliventris (Ashmead). Transf. to Lygocerus Foerster, p. 669. 

eyrthrolhorax (Ashmead). Transf. from Conostigmoides Dodd, p. 672, teste 
L. Masner. 
Taxonomy: Masner, 1964. Acta Soc. Ent. Cechosl. 61: 124-125. 

Genus Conostigmoides Dodd, p. 672 
Syn. of Conostigmus Dahlbom, supra. 
erythrothorax (Ashmead). Transf. to Conostigmus Dahlbom, p. 670. 

Genus DENDROCERUS Ratzeburg, p. 672 

Delete; a valid genus but without known Nearctic species. Transf. the 
N. Amer. forms to Atritomellus Kieflfer, p. 669 {teste Masner, 
1964. Acta Soc. Ent. Cechosl. 61: 125). 

Family DIAPRIIDAE 

Taxonomy: Nixon, 1957. Roy. Ent. Soc, London Handbooks 8 (3) : 3. — 
Masner, 1961. Inst. Sci. de Madagascar Mem. 12: 289-292. 

Subfamily DIAPRIINAE 

Taxonomy: Risbec, 1950. Contribution a I'Etude des Proctotrupidae. Fr. 
W. Africa, Trav. du Lab. d'Ent. du Secteur Soudanais de Rech- 
erches Agron., pp. 224-225 (key to African genera). — Masner, 
1961. Inst. Sci. de Madagascar Mem. 12: 292. 

Genus BRUE50PRIA Wing, Sup. 1, p. 89 

americana (Brues). D. C. 

Genus AUXOPAEDEUTES Brues, p. 672 

Cracinopria Fouts. Syn. by Masner, 1964. Acta Ent. Soc. Cechosl. 61: 
135. All species listed under Cracinopria in Catalog are n. combs, 
in Auxopaedeutes. Endings of names requiring agreement in 
gender should be changed to masculine. 

Genus MYRMECOPRIA Ashmead, p. 673 

mellea (Ashmead). 

Taxonomy: Masner, 1964. Acta Soc. Ent. Cechosl. 61: 123. Redescrip- 
tion, with first description of female. 

Genus Adeliopra Ashmead, p. 673 

Syn. of Entoniacis Foerster, infra. 
longii Ashmead. Transf. to Entomacis Foerster, infra. 



DIAPRIIDAE 289 

Genus ENTOMACIS Foerster, p. 673 

Hemilexis Foerster, 1856. See 1st Sup. p. 90. 
Hemilexodes Ashmead, 1893. N. syn. ( teste L. Masner) , 
Adeliopria Ashmead, 1902. Syn. by Masner, 1964. Acta Soc. Ent. 
Cechosl. 61: 134. 

floridana (Ashmead), n. comb. Transf. with Hemilexodes, 1st Sup. p. 90 
{teste L. Masner). 

longii (Ashmead). Transf. from Adeliopria Ashmead, teste L. Masner. 
Taxonomy: Masner, 1964, loc. cit. supra, p. 134. 

Genus Aparamesius Kieffer, p. 673 

Syn. of Paramesius Westwood, infra. Transf. included spp. to that 
genus. 

Genus PARAMESIUS Westwood, p. 674 

*Lipoglyptus Crawford, 1910. U. S. Natl. Mus. Proc. 38: 123 (through 
synonymy with Aparamesius Kieffer, teste Baltazar, 1961. Philip- 
pine Jour. Sci. 90: 395). 

Type: Lipoglyptus primus Crawford. By monotypy and orig. 
desig. 
Aparamesius Kieffer, 1913. N. syn. (teste L. Masner). Cite analis (Ash- 
mead) and nigriclavis (Fonts) as n. combs, and remove paren- 
theses from author of clavipes. 

Genus ANEURHYNCHUS Westwood, pp. 674-675 

Glyptonota Foerster, 1856. Syn. by Masner and Sundholm, 1959. Acta 
Soc. Ent. cechosl. 56: 164. 

nigriclavatus (Ashmead). Transf . from G/ypionofa Foerster, p. 677. 

Taxonomy: Masner and Sundholm, 1959, loc. cit. supra, pp. 162-163. 

Genus SPiLOMICRUS Westwood, pp. 675-676 

Loxotropa Foerster, 1856. Syn. by Masner, 1964. Entomophaga 9 : 82. 

*Eriopria Kieffer, 1910. In Andre, Spec. Hym. Eur. Alg., v. 10, pp. 693, 
744. Syn. by Masner, 1964 (loc. cit. supra, p. 82). 

Type: Eriopria nigra Kieffer. Desig. by Kieffer, 1912. 
"Tritopria Kieffer, 1910. Loc. cit. supra, pp. 717, 748. Syn. by Masner, 
1964 {loc. cit. supra, p. 82). 

Type : Tritopria lusitanica Kieffer. By monotypy. 
*Cologlpytus Crawford, 1910. U. S. Natl. Mus. Proc. 38: 123. N. syn. 
{teste L. Masner). 

Type: Cologlyptus kiefferi Crawford. By monotypy and orig. 
desig. 
*Scutellipria Szabo, 1961. Mus. Nat. Hungarici, Ann. Hist.-Nat. 53: 
493. Syn. by Masner, 1964 {loc. cit. supra, p. 82). 

Type: ScutelUpria quinquepunctata Szabo. By monotypy and 
orig. desig. 

*foutsi Masner, n. name. Replaces kiefferi Fonts, preocc. 

kiefferi Fonts. Preocc. Replaced by foutsi Masner. 

*Genus PENTAPRIA Kieffer 

( = Xenopr/a Fouts, p. 676) 

Pentapria Kieffer, 1905. Mus. Civ. Stor. Nat. Genova, ser. 3, 2: 34. 

Type: Pentapria conjugens Kieffer. First included species. 
Xenopria Fouts, 1939. Syn. by Masner, 1964. Acta Soc. Ent. Cechosl. 
61: 134. 

Columbiana (Fouts). 

Taxonomy: Masner, 1964, loc. cit. supra, p. 134. 



290 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus Phaenopria Ashmead, pp. 676—677 
Syn. of Trichopria Ashmead, p. 678 ; transf . included spp. to that genus. 

Genus Clyptonota Foerster, p. 677 
Syn. of Aneurhynchus Westwood, p. 674. 
nigroclavata Ashmead. Transf. to Aneurhynchus Westwood, p. 674. 

Genus Acidopria KiefFer, p. 677 

Syn. of Basalys Westwood, p. 680. 

Genus Loxoiropa, Foerster, p. 677 

Syn. of Spilomicrus Westwood, p. 675; Loxotropa of Catalog = Basalys 
Westwood, p. 680. Transf. spp. included in Loxotropa in Catalog 
to Basalys. 

Genus TRICHOPRIA Ashmead, pp. 678-680 

Phaenopria Ashmead. Syn. by Sundholm, 1960. Opuscula Ent. 25: 220. 

All species listed in Catalog are transf. here as n. combs. 
Delete Ceratopria Ashmead from syn. and transf. to syn. of Basalys 

Westwood, p. 680. 
Taxonomy: Sundholm, loc. cit. supra, pp. 215-223. — Masner, 1964. Acta 

Soc. Ent. cechosl. 61: 136. 

bifoveolata (Ashmead). Return to Trichopria Ashmead from Ceratopria Ash- 
mead where it was transf. in Sup. 1, p. 90. 

Columbiana (Ashmead). Minn. 

floridana (Ashmead). Return to Trichopria Ashmead from Ceratopria Ash- 
mead where it was transf. in Sup. 1, p. 90. 

infuscaticeps (Ashmead). Return to Trichopria Ashmead from Ceratopria 
Ashmead where it was transf. in Sup. 1, p. 90. 

longiceps (Ashmead). Transf. to Basalys Westwood, p. 680. 

megaplasta (Ashmead). Return to Trichopria Ashmead from Ceratopria Ash- 
mead where it was transf. in Sup. 1, p. 90. 

popei (Muesebeck). N. Y., Minn. Hosts: Dictya s^p., Elgiva ruf a (Fanz.) . 
Biology: Knutson and Berg, 1963. Canad. Ent. 95: 724-726. 

pusilla (Ashmead). Transf. to Basalys Westwood, p. 680. 

simulans (Ashmead). Transf. to Diapria Latreille, p. 680. 

tabanivora Fonts. Misspelled tabanviora in Catalog. N. Y., Ala., Fla. Host: 
Tabanus liiieola F. 

virginica (Ashmead). Transf. with Phaenopria; preocc. in Trichopria; re- 
placed by virginiensis L. Masner, n. name. 

*virginiensis Masner, n. name. Replaces Phaenopria virginica Ashmead nee 
Diapria virginica Ashmead, both now in Trichopria. 

Genus Cracinopria Fouts, p. 680 

Syn. of Auxopaedeules Brues, p. 672; transf. all included species to that 
genus. 

Genus BASALYS Westwood, p. 680 
IzzLoxotropa Foerster of Catalog, p. 677) 

Ceratopria Ashmead, 1893. Remove from syn. under Trichopria Ash- 
mead. Syn. by Masner, 1964. Entomophaga 9 : 85. 
Acidopria Kielfer, 1913. Syn. by Masner, 1964 (loc. cit. supra, p. 85). 
*Tropidopsis Ashmead, 1893. U. S. Natl. Mus. Bui. 45, pp. 386, 402. N. 
syn. (teste L. Masner). 

Type: Tropidopsis clavata Ashmead. By monotypy and orig. 
desig. 



DIAPRIIDAE 291 

*Nesopria Muesebeck and Walkley, 1956. N. name for Tropidopsis Ash- 
mead, preocc. N. syn. (teste L. Masner). 
Taxonomy: Masner, 1964. Entomophaga 9: 85-86. 

All species listed in the Catalog under Acidopria KieflFer and Loxotropa 
Foerster, p. 677, are transf. to Basalys as n. combs, (teste L. Masner). 

longiceps (Ashmead). Transf. from Trichopria Ashmead, p. 679 (teste 
Masner, 1964. Entomophaga 9: 86). 

pusilla (Ashmead), n. comb. Transf. from Trichopria Ashmead, p. 679 (teste 
L. Masner). 

Genus DIAPRIA Latreille, p. 680 

Taxonomy: Sundholm, 1960. Opusc. Ent. 25: 215-223. 

simulans (Ashmead), n. comb. Transf. from Trichopria Ashmead, p. 679 
(teste L. Masner). 

Genus DOLIOPRIA KiefFer, p. 681 

americana Fouts. 

Taxonomy: Masner. 1964. Acta Soc. Ent. Cechosl. 61: 135. 

Genus Prosynacra KiefFer, p. 681 
Syn. of *Synacra Foerster ( = Paratelopsilus Whittaker) p. 690. 
gracilis Fouts. Transf. to Synacra Foerster. 

♦Subfamily AMBOSITRINAE 

Place in Diapriidae after Prosynacra Kieffer, p. 681. 

Taxonomy: Masner, 1961. Inst. Sci. de Madagascar Mem. 12: 289-295. 

Genus PROPSILOMMA Kieffer, p. 683 

Transf. from subfam. Cinetinae (replaced by subfam. Belytinae in Sup. 
1, p. 91) to subfam. Ambositrinae. 

♦Subfamily ISMARINAE 

Place in Diapriidae before Subfam. Cinetinae, p. 681 (replaced in Sup. 1 
by Belytinae). 

Taxonomy: Hellen, 1963. Fauna Fennica 14: 4. 

Genus ISMARUS Haliday, p. 681 

Transf. from subfam. Cinetinae (replaced by Belytinae in Sup. 1, p. 91) 
to subfam. Ismarinae. 

Biology: Chambers, 1955. Ent. Monthly Mag. 91: 114-115. 
Taxonomy: Nixon, 1957. Re 7. Ent. Soc, London, Handbooks 8 (3) : 11. 

Subfamily BELYTINAE 

Taxonomy: Nixon, 1957. Roy. Ent. Soc, London, Handbooks 8 (3), 107 
pp. (Keys to the British species). — Masner, 1961. Inst. Sci. de 
Madagascar Mem. 12 : 292. 

Genus OXYLABIS Foerster, p. 681 

Taxonomy: Nixon, 1957. Roy. Ent. Soc, London, Handbooks 8 (3) : 27. 

bifoveolata Brues. Remove from Oxylabis, p. 681, and from Proctotrupoidea; 
a species of Cynipoidea (teste L. Masner upon examination of 
type). 

fuscicornis (Ashmead), n. comb. Transf. from Aclista Foerster, p. 687 (teste 
L. Masner) . 



292 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus BELYTA Jurine, Sup. 1, p. 91 

*ParacU8ta Kieffer, 1909. In Andre, Spec. Hym. Eur. Alg., v. 10, p. 476. 
Syn. by Masner, 1965. Brit. Mus. Nat. Hist. BuL, Ent. Sup. 1: 
63. 
Type: Belyta brachyptera Thomson. Desig. by Kieffer, 1910. 

All species listed in Catalog under Cinetus Jurine, pp. 682-683, belong 
here (see Sup. 1, p. 91). Endings of names requiring agreement in gender 
should be changed to feminine. 

frontalis Ashmead. Transf. from Aclista Foerster, p. 687 (teste L. Masner). 

nasuta (Provancher), n. comb. Transf. from Cinetus Jurine, Sup. 1, p. 91 
(teste L. Masner). 

picipes (Ashmead, n. comb. Transf. from Aclista Foerster, p. 687 (teste L. 
Masner) . 

Genus ZYGOTA Foerster, pp. 683-684 

longicornis (Ashmead), n. comb. Transf. from Aclista Foerster, p. 688 (teste 
L. Masner). 

Genus MACROHYNNIS Foerster, p. 684 

*Cinelaptus Nixon, 1957. Roy. Ent. Soc, London, Handbooks 8 (3) : 7, 
91. N. syn. (teste L. Masner). 

Type: Cinelapius fragilis Nixon. By monotypy and orig desig. 

Genus MIOTA Foerster, pp. 684-685 

Synonymized under Cinetus Foerster by Muesebeck, 1958, Agr. Monogr. 
No. 2, Sup. 1, p. 91, and by Masner, 1964, Acta Soc. Ent. Cechosl. 61: 126, 
but now restored as a valid genus with Leptorhaptus authors, not Foerster, 
as synonym. Most of the species listed in the Catalog under Leptorhaptus 
are transferred here (teste L. Masner). 

*Leptonetus Masner. 1964. Acta Soc. Ent. Cechosl. 61: 128. N. syn. 
(teste L. Masner). 

Type : Cinetus politus Thomson. By orig. desig. 

alaskensis (Ashmead), n. comb. Transf. fi-om Aclista Foerster, p. 686 (teste 
L. Masner) . 

americana (Ashmead), n. comb. Transf. from Cinetus Jurine, Sup. 1, p. 91 
(teste L. Masner). 

ashmeadi (Kieffer), n. comb. Transf. from Leptorhaptus Foerster, p. 685 
(teste L. Masner). 

bakeri (Kieffer), n. comb. Transf. from Leptorhaptus Foerster, p. 685 (teste 
L. Masner). 

bisulcata (Kieffer), n. comb. Transf. from Leptorhaptus Foerster, p. 685 
(teste L. Masner). 

clarimontis (Kieffer), n. comb. Transf. from Leptorhaptus Foerster, p. 685 
(teste L. Masner). 

coloradensis (Ashmead). Transf. to Cinetus Jurine, Sup. 1, p. 91. 

filicornis (Keiffer), n. comb. Transf. from Leptorhaptus Foerster, p. 685 
(teste L. Masner). 

flavicomis Kieffer. Transf. from Leptorhaptus Foerster, p. 685 (teste L. 
Masner) . 

longinervis (Kieffer), n. comb. Transf. from Leptorhaptus Foerster, p. 685 
(teste L. Masner). 

pleuralis (Fouts). Transf. to Cinetus Jurine, Sup. 1, p. 91. 

rufa (Ashmead), n. comb. Transf. from Leptorhaptus Foerster, p. 685 (teste 
L. Masner) . 



DIAPRIIDAE 293 

vera (Fouts), n. comb. Transf. from Scorpioteleia Ashmead, p. 685 {teste L. 

Masner). 
Taxonomy: Masner, 1964. Acta Soc. Ent. Cechosl. 61:129. 

Genus CINETUS Jurine, Sup. 1, p. 91 
( ^ tepforhapf us Foerster, p. 685; Stylidon Ashmead, p. 685) 

Remove Miota Foerster from synonymy; a valid genus. Both Leptor- 
haptus Foerster and Stylidolon Ashmead are now considered synonyms of 
Cinetus Jurine {teste Masner, 1964. Acta Soc. Ent. Cechosl. 61: 126). How- 
ever, of the species listed in the Catalog under Leptorhaptus only conicus 
Ashmead and texanus (Ashmead) belong in Cinetus; the remainder are 
transf. to Miota Foerster, p. 684 {teste L. Masner). 

americanus (Ashmead). Transf. to Miota Foerster, p. 684. 

coloradensis (Ashmead), n. comb. Transf. from Miota Foerster, p. 684 {teste 
L. Masner). 

conicus (Ashmead). Transf. from Leptorhaptus Foerster, p. 685 {teste 
Masner, 1964. Acta Soc. Ent. cechosl. 61: 128). 

nasutus Provancher. Transf. to Belyta Jurine, Sup. 1, p. 91. 

pleuralis Fouts. Transf. from Miota Foerster, p. 685 {teste L. Masner). 

politus (Ashmead). Preocc. when transf. from S'ylidolon Ashmead to Cin- 
etus Jurine. Replaced by subpolitus Masner. 

ruficornis (Ashmead), n. comb. Transf. from Aclista Foerster, p. 688 {teste 
L. Masner). 

subpolitus Masner. Replaces Stylidolon politum Ashmead, preocc. when 
transf. to Cinetus, teste Masner, 1964. Acta Soc. Ent. Cechosl. 61 : 
128. 

texanus (Ashmead), n. comb. Transf. from Leptorhaptus Foerster, p. 685 
{teste L. Masner). 

Genus SCORPIOTELEIA Ashmead, p. 685 
Taxonomy: Masner, 1964. Acta Soc. Ent. Cechosl. 61: 130. 
vera Fouts. Transf. to Miota Foerster, p. 684. 

Genus ACLISTA Foerster, pp. 685-689 

Pantoclis Foerster. Remove from syn.; an independent genus {teste Nix- 
on, 1957. Roy. Ent. Soc, London, Handbooks 8 (3) : 11, 41). 

Zelotypa Foerster. Remove from syn. under Aclista and place as syn. of 
Panioclis Foerster. 

aculeata (Kieffer). Transf . to Pantoclis Foer.ster. 

alaskensis (Ashmead). Transf. to Miota Foerster, p. 684. 

analis (Ashmead). Transf. to Pantoclis Foerster. 

bakeri (Kieffer). Transf. to Pantoclis Foerster. 

bruesi (Kieffer). Transf . to Pantoclis Foerster. 

calif ornica (Ashmead). Transf . to Pantoclis Foerster. 

canadensis (Ashmead). Transf . to Pantoclis Foerster. 

clinoneura (Kieffer). Transf. to Pantoclis Foerster. 

coloradensis (Ashmead). Transf. to Pantoclis Foerster. 

consimilis Muesebeck. Transf. to Pantoclis Foerster. 

dubiosa (Kieffer). Transf . to Pantoclis Foerster. 

duplicata Muesebeck and Walkley. Transf. to Pantoclis Foerster. 

flavicrus (Kieffer). Transf. to Pantoclis Foerster. 

floridana (Ashmead). Transf. to Pantoclis Foerster. 



294 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

frontalis (Ashmead). Transf. to Belyta Jurine, Sup. 1, p. 91. 

fuscicornis (Ashmead). Transf. to Oxylabis Foerster, p. 681. 

georgica (Ashmead). Transf. to Pantoclis Foerster. 

integrifrons (Kieffer). Transf . to Pantoclis Foerster. 

klagesi (Kieffer). Transf. to Pantoclis Foerster. 

jeanneitensis Muesebeck and Walkley. Transf. to Pantoclis Foerster. 

longicornis (Ashmead). Transf. to Zygota Foerster, p. 683. 

monilicornis (Ashmead). Transf. to Pantoclis Foerster. 

montana (Ashmead). Transf. to Pantoclis Foerster. 

orthoneura (Kieffer). Transf . to Pantoclis Foerster. 

picipes (Ashmead). Transf. to Belyta Jurine, Sup. 1, p. 91. 

rufescens (Ashmead). Transf. to Pantoclis Foerster. 

ruficauda (Ashmead). Ti-ansf. to Pantoclis Foerster. 

ruficornis (Ashmead). Transf. to Cinetus Jurine, Sup. 1, p. 91. 

scutellata aptera (Kieffer). Raised to specific rank and transf. to Pantoclis 
Foerster. 

singularis (Kieffer). Transf. to Pantoclis Foerster. 

subfusca (Kieffer). Transf. to Pantoclis Foerster. 

subobtusa (Kieffer). Transf . to Pantoclis Foerster. 

Genus PANTOCLIS Foerster, p. 686 

Remove from syn. under Aclista Foerster and treat as valid genus, teste 
Nixon, 1957. Roy. Ent. Soc, London, Handbooks 81 (3) : 11, 41; place after 
Aclista, p. 689. 

Zelotypa Foerster; remove from syn. of Aclista Foerster. N. syn. {teste 
L. Masner) . 

aculeata Kieffer. Transf. from Aclista Foerster, p. 686 {teste L. Masner). 

analis Ashmead. Transf. from Aclista Foerster, p. 686 {teste L. Masner). 

aptera Kieffer, changed status. 

Pantoclis scutellata aptera Kieffer. Raised to specific rank and transf. 
from Aclista Foerster, p. 689 {teste L. Masner). 

bakeri Kieffer. Transf. from Aclista Foerster, p. 686 {teste L. Masner). 

bruesi Kieffer. Transf. from Aclista Foerster, p. 686 {teste L. Masner). 

calif ornica (Ashmead). Transf. from Aclista Foerster, p. 686 {teste L. 
Masner). 

canadensis Ashmead. Transf. from Aclista Foerster, p. 686 {teste L. 
Masner). 

clinoneura Kieffer. Transf. from Aclista Foerster, p. 686 {teste L. Masner). 

coloradensis (Ashmead), n. comb. Transf. from Aclista Foerster, p. 686 
{teste L. Masner). 

consimilis (Muesebeck), n. comb. Transf. from Aclista Foerster, Sup. 1, p. 
91, {teste L. Masner). 

dubiosa Kieffer. Transf. from Aclista Foerster, p. 687 {teste L. Masner). 

duplicata (Muesebeck and Walkley), n. comb. Transf. from Aclista Foerster, 
p. 686 {teste L. Masner). 

flavicnis Kieffer. Transf. from Aclista Foerster, p. 687 {teste L. Masner). 

floridana Ashmead. Transf. from Aclista Foerster, p. 687 {teste L. Masner). 

georgica (Ashmead). Transf. from Aclista Foerster, p. 687 {teste L. 
Masner). 



DIAPRiroAE 295 

integrifrons Kieffer. Transf. from Aclista Foerster, p. 687 {teste L. Masner). 

jeannettensis (Muesebeck and Walkley), n. comb. Transf. from Aclista 
Foerster, p. 688 (teste L. Masner). 

klagesi Kieffer. Transf. from Aclista Foerster, p. 687 (teste L. Masner). 

monilicornis (Ashmead), n. comb. Transf. from Aclista Foerster, p. 688 

(teste L. Masner). 
montana Ashmead. Transf. from Aclista Foerster, p. 688 (teste L. Masner). 
orthoneura Kieffer. Transf. from Aclista Foerster, p. 688 (teste L. Masner). 
rufescens Ashmead. Transf. from Aclista Foerster, p. 688 (teste L. Masner). 
ruficauda Ashmead. Transf. from Aclista Foerster, p. 688 (teste L. Masner). 
singularis Kieffer. Transf. from Aclista Foerster, p. 689 (teste L. Masner). 
subfusca Kieffer. Transf. from Aclista Foerster, p. 689 (teste L. Masner). 
subobtusa Kieffer. Transf. from Aclista Foerster, p. 689 (teste L. Masner). 

Genus ACROPIESTA Foerster, p. 689 

*Pantopies(a Maneval, 1939. Soc. Ent. France Bui. 44: 169, 170. Syn. by 
Nixon, 1957. Roy. Ent. Soc, London, Handbooks 8 (3) : 24. 

Type: Ciiteius flaviventris Thomson. By monotypy and orig. 
desig. 

Genus RHYNCHOPSILUS KiefFer, p. 689 

*Brunnicophilus Nixon, 1931. Ent. Rec. and Jour. Variation 43: 83. 
Syn. by Nixon, 1957. Roy. Ent., London, Handbooks 8 (3) : 13. 

Type: Brnnnicophilus donisthorpei Nixon. By monotypy and 
orig. desig. 

Genus POLYPEZA Foerster, Sup. 1, p. 89 
( = Afe/opsi7os Kieffer, pp. 689-690) 

Syn. by Masner, 1964. Acta Soc. Ent. Cechosl. 61: 131. Remove from 
Diapriinae to Belytinae where it replaces Atelopsilus Kieffer, p. 
689. 

Taxonomy: Masner and Sundholm, 1959. Acta Soc. Ent. Cechosl. 56: 
164. — Masner, 1964, loc. cit. supra, p. 131. 

alaskensis (Ashmead), n. comb. Transf. from Spilomicrus Westwood, p. 675 
(teste L. Masner). 

brunnea (Ashmead). 

Taxonomy: Masner, 1964, loc. cit. supra, p. 131. 

*Genus SYNACRA Foerster 

Synacra Foerster, 1856. Hym. Stud., v. 2, pp. 128, 130, 134. 

Type: Diapria brachialis Nees. Desig. by Ashmead, 1893. 
*Artibolu8 Haliday, 1857. Nat. Hist. Rev. 4: Proc. p. 173. 

Type: Diapria brachialis Nees, Desig. by Muesebeck and 
Walkley, 1956. Isogenotypic with Synacra Foerster. 
*Neuropria Kieffer, 1904. Soc. Hist. Nat. Metz Bui. 23: 53. Syn. by 
Pschorn-Walcher, 1957, Schweiz. Ent. Gesell. Mitt. 30 : 74. 
Type : Neuropria sociabilis Kieffer. By monotypy. 
Prosvnacra Kieffer, 1905. N. syn. (teste L. Masner). 

Para'telopsilus Whittaker, 1930. Syn. by Masner, 1964. Acta Soc. Ent. 
cechosl. 61: 131. 

Place at end of subfam. Belytinae. 

Taxonomy: Pschorn-Walcher, 1957. Schweiz. Ent. Gesell. Mitt. 30: 
74-76. 

canadensis (Whittaker) . 

Taxonomy: Masner, 1964. Acta Soc. Ent. Cechosl. 61: 131-132. 



296 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

gracilis (Fouts), n. comb. Transf. from Prosynacra Kieffer, p. 681 {teste L. 
Masner). 

Family SCELIONiDAE 

Subfamily TELENOMINAE 

Genus ARADOPHAGUS Ashmead, p. 690 
f asciatus Ashmead. Tex. ; Europe. 

Genus TIPHODYTES Bradley, p. 690 
gerriphagus (Marchal). Calif. 

Genus EUMICROSOMA Gahan, p. 690 

*T^ardo Nixon, 1938. Ann. Mag. Nat. Hist. (11), 1: 278. Syn. by 
Masner, 1965. Brit. Mus. Nat. Hist. BuL, Ent. Sup. 1: 108. 
Type: ISardo cumaeus Nixon. By orig. desig. 
*Szelenyiella Szabo, 1957. Opusc. Zool. 2: 49. Syn. by Masner, 1965, loc. 
cit. supra, p. 108. 

Type : Szelenyiella platythoracis Szabo. By monotypy and orig. 
desig. 

beneficum Gahan. U.S.S.R. 

bicolor (Ashmead). Mich. Host: Unknown eggs on Poa compressa. 

Genus PROTRIMORUS KiefFer, p. 690 

Transf. to Scelioninae as a syn. of Opisthacantha Ashmead {teste 
Masner, 1964. Acta Soc. Ent. Cechosl. 61: 140). 

TELENOMUS subgenus TELENOMUS Holiday, pp. 691-694 

alsophilae Viereck. Mass., Pa. Host: Ennomus sub si gnarius (Hhn.) . 

basalis Wallaston. Transf. to Trissolcus Ashmead, p. 694, 

bifidus Riley. Wis. Host: Dasychira plagiata (Wlkr.). 

californicus Ashmead. N. Mex., Nev., Oreg. 

chrysopae Ashmead. Host: Lomamyia flavicomis (Wlkr.). 

clisiocampae Riley. Colo., Wash. 

coloradensis Crawford. Idaho, Wash. Host: Hemerocampa pseudotsugata 
McD. 

cosmopeplae Gahan. Transf. to Trissolcus Ashmead, p. 694. 

dimmocki Ashmead. Ohio, Ga., Fla. Hosts: Cosmopepla bimaculata 
(Thom.), Podisus maculiventris (Say), Thyanta custator (F.). 

dolichocerus (Ashmead). Okla. Host: Emesaya brevipennis (Say). 

graptae Howard. 111., Colo., Ariz. Hosts: Lycaena thoe Guerin, Plebejus ica- 
rioides Bdv. 

heliothidis Ashmead. Fla. 

ichthyurae Ashmead. Maine, Okla. 

perplexus Girault. Hosts: Euschistus variolarius (Pal. de Beauv.), E. tris- 
tignius (Say). 

spilosomatis Ashmead. Ohio. 

utahenais Ashmead. Transf. to Trissolcus Ashmead, p. 694. 

Genus TRISSOLCUS Ashmead, p. 694 

*A8olcu8 Nakagawa, 1900. Spec. Rep. Imp. Agr. Sta. Japan 6, p. 17. 
Syn. by Masner, 1964. Acta Soc. Ent. Cechosl. 61 : 145. 



SCELIONIDAE 297 

Type : Asolcus nigripedius Nakagawa. By monotypy. 
*Aphanurus KieflFer, 1912. In Andre, Spec. Hym. Eur. Alg., v. 11, pp. 10, 
69. 

Type: Teleas semistriatus Nees. By orig. desig. 
*Microphanurus Kieffer, 1926. Das Tierreich, Lief. 48, pp. xv, 16, 91. 
Proposed for Aphanurus Kieffer, preocc. Syn. of Asolcus Naka- 
gawa, teste Watanabe, 1951. Shikoku Ent. Soc. Trans. 2: 21. 

basalis (Wollaston), n. comb. Transf. from Telenomus subg. Telenomus Ha- 

liday. Sup. 1, p. 92 (teste L. Masner). 

brochymenae (Ashmead). Ind., Tex. Host: A crostemum hilare (Say). 

cosinopeplae (Gahan), n. comb. Transf. from Telenomus subg. Telenomus Ha- 

liday, p. 692 (teste L. Masner). 

podisi Ashmead. W. Va., Ohio, Ind. Host: Banasa dimidiata (Say). 

thyantae Ashmead. Ga., Mo., Tex. 

utahensis (Ashmead), n. comb. Transf. from Telenomus subg. Telenomus 
Haliday, p. 694 (teste L. Masner). 

Genus Hadrophanurus KiefFer, p. 694 

Transf. to Scelioninae as a syn. of Gryon Haliday (teste Masner, 1961. 
Acta Soc. Ent. Cechosl. 58: 158). 

pennsylvanicus (Ashmead). Transf. to Gryon Haliday. 

Subfamily Bae/nae of Catalog 

Delete; include in Scelioninae. Place the contained genera at the begin- 
ning of the Scelioninae, p. 701, except Tetrabaeus Kieffer, which is transf. to 
Platygasteridae, p. 708. Transf. Acoloides xanthogaster (Ashmead) to 
* Gryon Haliday. 

Subfamily TELEASINAE 

Genus TRIMORUS Foerster, pp. 697-701 
( = rr(chos/us Provaneher, Sup. 1, p. 92) 

Trichasius Provaneher, 1887. N. syn. (teste L. Masner). 

Paragryon Kieffer, 1908. N. syn. (teste L. Masner). 

Transf. here all spp. in Catalog under Paragryon Kieffer, p. 696, and in 

Sup. 1 under Trichasius Provaneher, p. 92; all are n. combs, in 

Triniorus Foerster. 

Subfamily SCELIONINAE 

Genus CERATOBAEUS Ashmead, p. 695 
binotatus Ashmead. Md. 

Genus BAEUS Haliday, p. 695 

zabriskiei (Ashmead), n. comb. Transf. from Acoloides Howard, p. 696 
(teste L. Masner). 

Genus Acofoides Howard, pp. 695—696 

Syn. of Idris Foerster, infra. Remove Psilacolus Kieffer from syn. under 
Acoloides and place as syn. of Gryon Haliday. Transf. to Idris Foerster all 
species listed under Acoloides in Catalog except xanthogaster (Ashmead), 
which is transf. to Gryon Haliday, and zabriskiei (Ashmead), which is 
transf. to Baeus Haliday, p. 695. 

*Genus IDRIS Foerster 
( = Aco/o/des Howard, pp. 695—696) 

Idris Foerster, 1856. Hym. Stud. v. 2, pp. 102, 105. 



298 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Type: Idris flavicornis Foerster. By monotypy. 
Acoloides Howard, 1890. Insect Life 2: 269. Syn. by Masner, 1961. Acta 

Soc. Ent. Cechosl. 58: 163. 
*Megacolus Priesner, 1951. Inst. Fouad I du Desert Bui. 1: 121. 

Type: Megacolus desertoruin Priesner. By monotypy and 
orig. desig. 
*Philoplane8 Muesebeck and Walkley, 1956. U. S. Natl. Mus. Proc. 105: 

384. N. name for Megacolus Priesner, preocc. 
Taxonomy: Masner, 1961, loc. cit. supra, pp. 163-166. 

saitidis (Howard). Pa., Ariz. Host: Phidippus opifex (McCook). 

seminiger (Ashmead). Nebr. 

Genus THORON Haliday, p. 696 

*inetallicus Haliday. Europe; Mass. 

Thoron metallicus Haliday, 1830. Ent. Mag. 1: 272. ?. 

pallipes Ashmead. Transf . to Paridris Kieffer, p. 704. 

Genus SCELIO Latreille, pp. 702-703 

Taxonomy: Nixon, 1958. Roy. Ent. Soc, London Trans. 110: 303-318 

(revision of African species). 
Biology: Greathead, 1963. Roy. Ent. Soc, London Trans. 114: 439-442. 

bisulcus (Ashmead). Hosts: Melanoplus sip., M. differ entialis (Thomas). 

Scelio striaticollis Brues, 1906. N. syn. (teste C. F. W. Muesebeck on 

comparison of types) . 
Biology: Lanham and Evans, 1958. Pan-Pacific Ent. 34: 213-214.— Lan- 

ham and Evans, 1960. Brooklyn Ent. Soc. Bui. 55: 84-86 (pho- 

resy) . 

striaticollis Bvues. Syn. of bisulcus (Ashmead). 

Genus OPISTHACANTHA Ashmead, p. 703 

Protrimorus Kieffer, 1908. Transf. from Telenominae and made syn. of 
Opisthacantha by Masner, 1964. Acta Soc. Ent. Cechosl. 61: 140. 

americana (Ashmead). Transf. from Protrimorus Kieffer, p. 690, by Masner, 

1964. Acta Soc Ent. Cechosl. 61: 141. 

Genus PARIDRIS KiefFer, p. 704 

brevipennis Fouts. Calif. 

pallipes (Ashmead), n. comb. Transf. from Thoron Haliday, p. 696 (teste L. 
Masner). 

*Genus CALOTELEA Westwood 

( = Cerafofe/e/a Kieffer, p. 705) 

Calotelea Westwood, 1837. In Hope, Ent. Soc. London Trans. 2: 55. 

Type: Calotelea aurantea Hope. By monotypy. 
*Caloteleia Ashmead, 1893. U. S. Natl. Mus. Bui. 45: 209, 210, 212. 

Emendation. 
Ceratoteleia Kieffer, 1908. Syn. by Masner, 1965. Brit. Mus. Nat. Hist. 

Bui., Ent. Sup. 1: 69. 
Prosanteris Kieffer, 1908. Syn. with Ceratoteleia by Muesebeck, 1958, 

Sup. 1, p. 93. 
*l>leuroteleia Kieffer, 1910. Ent. Soc. France Bui. 1910: 293. Syn. by 

Masner, 1965, loc. cit. supra, p. 69. 

Type: Npuroteleia rujfa Kieffer. By orig. desig. 
*Lamproteleia Kieffer, 1910, loc. cit. supra, p. 293. Syn. by Masner, 

1965, loc. cit. supra, p. 69. 

Type: Lamproteleia fasciatipennis Kieffer. By orig. desig. 

Transf. here the species listed under Ceratoteleia Kieffer in the Catalog, 
p. 705, and in Sup. 1, p. 93. 



SCELIONIDAE 299 

*Genus GRYON Haliday 

Gryon Haliday, 1833. Ent. Mag, 1 : 271. 

Type: Gryon misellus Haliday. By monotypy. 
*Acolu8 Foerster, 1856. Hym. Stud., v. 2, p. 100. Syn. by Masner, 1961. 
Acta Soc. Ent. Cechosl. 58: 158. 

Type: Aeolus opacus Thomson. Desig. by Ashmead, 1903. 
Hadronotus Foerster, 1856. Syn. by Masner, 1961, loc. cit. supra, p. 158. 
*Plastogryon Kieffer, 1908. Soc. Sci. de Bruxelles Ann. 32: 119, 131. 
Syn. by Maneval, 1940. Faune de la France, v. 7, p. 113. 

Type: Plastogryon foersteri Kieffer. Desig. by Brues, 1908. 
Psilacolus Kieffer, 1908. N. syn. (teste L. Masner). 

*Holacolus Kieffer, 1912. In Andre, Spec. Hym. Eur. Alg., v. 11, pp. 89, 
94, 106. Syn. by Masner, 1961, loc. cit. supra, p. 158. 

Type: Aeolus opaeus Thomson. Desig. by Muesebeck and 
Walkley, 1956. 
Telenomoides Dodd, 1913. Syn. of Hadronotus Foerster. 
Hadronotellus Kieffer, 1917. Syn. of Hadronotus Foerster. 
Hadrophanurus Kieffer, 1926. Syn. by Masner, 1961, loc. cit. supra, p. 

158. 
*Heterogryon Kieffer, 1926. Das Tierreich, Lief, 48, pp. xxvi, 271, 446, 
448. Syn. by Masner, 1961, loc. cit. supra, p, 158, 

Type: Plastogryon sagax Kieffer, Desig, by Muesebeck and 
Walkley, 1956, 
Synteleia Fonts, 1927, N, syn, (teste L, Masner). 

Transf. to Gryon all species listed in the Catalog under Hadrophanurus 
Kieffer, p. 694, Hadronotus Foerster, pp. 704-705, and Synteleia Fonts, p. 706, 
and mark all, except pennsylvanicus (Ashmead), n, combs. Insert Gryon Ha- 
liday in place of Hadronotus Foerster, pp. 704-705, 

xanthogaster (Ashmead), n, comb, Transf, from Acoloides Howard, p, 696 
(teste L, Masner), 

Genus Pegofe/eio KiefFer, pp. 705—706 

Delete from Catalog; no North American species known. Transfer the 
species listed under Pegoteleia in the Catalog to *Probaryconus Kieffer, 
where all are n. combs, Chg. striata (Ashmead) to striatus. 

Genus Prosapegus Kieffer, p. 705 

Syn. of Macroteleia Westwood, infra. 
elongata (Ashmead). Transf . to Macroteleia Westwood, 

*Genus PROBARYCONUS KiefFer 
(=:Pegofe/e/o of Catalog, pp. 705-706) 

Probaryconus Kieffer, 1908. Soc. Sci. de Bruxelles Ann. 32: 118, 165, 
168. 

Type: Baryconus (Probaryeonus) spinosus Kieffer. By mono- 

typy. 

*Amblyconus Kieffer, 1913. In Andre, Spec. Hym, Eur. Alg,, v, 11 p. 221, 
Syn. by Masner, 1965. Brit, Mus, (Nat, Hist,) Bui,, Ent. Sup, 1: 
90, 

All species listed in the Catalog under Pegoteleia Kieffer, p, 706, are 
transf, here as n, combs,; striata (Ashmead) becomes striatus. 

Genus MACROTELEIA Westwood, p. 706 

*Baeoneura Foerster, 1856. H\Tn. Stud., v. 2, pp. 100, 102. Syn. by 
Muesebeck and Walkley, 1956. U. S. Natl. Mus. Proc. 105: 335. 

Type: Baeoneura floridana Ashmead. Desig. by Muesebeck 
and Walkley, 1956. 
Baeeeura Ashmead, 1887. Ent. Amer. 3: 99. Error for Baeoneura 

Foerster. 
Prosapegus Kieffer, 1908. Syn. by Masner, 1964. Acta Soc. Ent. Cechhosl. 
61: 137. 



300 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Stictoteleia Kieffer, 1926. Syn. by Masner, 1964, loc. cit. supra, p. 137. 
elongata (Ashmead), n. comb. Transf. with Prosapegus Kieffer, p. 705. 
virginiensis Ashmead. Transf. with Stictoteleia Kieffer, p. 706. 

Genus Sfkfofehia Kieffer, p. 706 

Syn. of Macroteleia Westwood, supra. 
virginiensis (Ashmead). Transf. to Macroteleia Westwood. 

Genus Synfelela Fouts, p. 706 

N. syn. of Gryon Haliday ( teste L. Masner) . The single species, coracina 
Fouts, is transf. to Grjon, and changed to coracinus (Fouts). 

Genus PSEUDANTERIS Foufs, p. 706 

Taxonomy: Masner, 1964. Acta Soc. Ent. Cechosl. 61:141. 

Family PLATYGASTERIDAE 

Taxonomy: Szabo, 1959. Mus. Nat. Hungarici Ann. Hist.-Nat. 51: 
389-390.— Masner, 1964. Acta Soc. Ent. cechosl. 61: 146-162. 

Subfamily INOSTEMMINAE 

Genus A1onocr//a Foerster, p. 707 
Syn. of Isostasius Foerster, p. 708. 
carinata Ashmead. Transf. to Metaclisis Foerster. 
nigripes Ashmead. Transf. to Platygaster Latreille. 

Genus METACLISIS Foerster, p. 707 
belonocnemae Ashmead. Transf. to Inostemnia Haliday, p. 708. 

carinata (Ashmead), n. comb. Transf. from Monocrita Foerster, p. 707 (teste 
L. Masner). 

erythropus Ashmead. Transf. to Platygaster Latreille, p. 710. 

floridana (Ashmead), n. comb. Transf. from Acerota Foerster, p. 707 {teste 
L. Masner). 

Genus Acerofa Foerster, p. 707 

Syn. of Inostemma Haliday; all included spp. are transf. to Inostemnia 
except floridana Ashmead, which is transf. to Metaclisis Foerster. 

Genus ALLOTROPA Foersfer, pp. 707-708 

burrelli Muesebeck. Established throughout range of host in U. S. 

Biology: Clausen, 1956. U. S. Dept. Agr. Tech. Bui. 1139, pp. 74-75. 

*citri Muesebeck. So. China; Calif. Host: Planococcus citri (Risso). 

AUotropa citri Muesebeck, 1954. Brooklyn Ent. Soc. Bui. 49: 18. $, 9. 
Release and recovery in U.S.: Bartlett and Lloyd, 1958. Jour. Econ. 
Ent. 51: 92-93. 

utilis Muesebeck. N. H., Vt. ; Japan. Host: Phenacoccus pergandei Ckll. 

Biology: Murakami, 1962. Kyushu Univ. Facul. Agr. Sci. Bui. 19: 
390-402. 

Genus Tridavus Brethes, p. 708 
Syn. of Fidiobia Ashmead, infra. 
drakei Ogloblin. Transf. to Fidiobia Ashmead. 

Genus FIDIOBIA Ashmead, p. 709 

Transf. from subfam. Platygasterinae to subfam. Inostemminae and 



PLATYGASTERIDAE 301 

place before Inostemma Haliday, p. 708 (teste Jansson, 1956. Ent. Tidskr. 
77: 87-90.— Masner, 1956. Acta Faun. Ent. Mus. Natl. Pragae 30: 114). 
Triclavus Brethes, 1916. N. syn. {teste L. Masner). 

drakei (Ogloblin), n. comb. Transf. with Triclavus Brethes, p. 708. 

Genus INOSTEMMA Haliday, p. 708 

Acerota Foerster. N. syn. (teste Masner, 1964. Acta Soc. Ent. cechosl. 
61: 146). All species listed in Catalog are transf. to Inostemma 
as n. combs, except floridana which is transf. to Metaclisis Foer- 
ster (teste L. Masner). 
*Brachiuostemina Kieffer, 1916. Zentbl. f. Bakt., Parasitenk. u. Infek- 
tionskrankeiten, Abt. 2, 46: 551. Syn. by Masner, 1964, loc. cit. 
supra, p. 146. 

Ty^e :Brachinostemma mediterraneum KiefPer. By monotypy 
and orig. desig. 
*Inocerota Szelenyi, 1939. Mus. Nat. Hungarici Ann. 32: 121. Syn. by 
Masner, 1964, loc. cit. supra, p. 146. 

Type: Inocerota discessus Szelenyi. By orig. desig. 

americanum (Ashmead), n. comb. Transf. from Acerota Foerster, p. 707. 
Taxonomy: Masner, 1964. Acta Soc. Ent. Cechosl. 61: 147. 

belonocnemae (Ashmead), n. comb. Transf. from Metaclisis Foerster, p. 707 
(teste L. Masner). 

cecidomyiae (Ashmead), n. comb. Transf. from Acerota Foerster, p. 707. 

leonardi (Fonts), n. comb. Transf. from Acerota Foerster, p. 707. 

Genus ISOSTASIUS Foerster, p. 708 

Monocrita Foerster. Syn. by Masner, 1965, Brit. Mus. Nat. Hist. Bui., 
Ent. Sup. 1: 130. 

Genus TETRABAEUS Kieffer, p. 695 

Transf. from Scelionidae to Platygasteridae, subfam. Inostemminae, and 
place after Isostasius Foerster, p. 708. 

*Crabroborus Muesebeck, 1963. Beitr. z. Ent. 13: 391. Syn. by Masner, 
1964. Acta Soc. Ent. cechosl. 61 : 148. 

Type: Crabroborus krombeini Muesebeck. By monotypy and 
orig. desig. 

americanus (Brues). Wis., Que., Md. Hosts: Ectemnius (Hypocrahro) pauci- 
viac2ilatus (Pack.), Euplilis (Corijnopus) rufigaster (Pack.), E. 
(C.) coarctata modesta (Roh.). 
*Crabroborus krombeini Muesebeck, 1963. Beitr. z. Ent. 13: 392. 9, $. 

Syn. by Masner, 1964. Acta Soc. Ent. cechosl. 61: 149. 
Biology: Krombein, 1964. Biol. Soc. Wash. Proc. 77: 104-108, fig. 11. 

Subfamily PLATYGASTERINAE 

Genus Fidiobia Ashmead, p. 709 
Transf. to Inostemminae, p. 708. 

Genus iSOCYBUS Foerster, p. 709 
canadensis (Provancher) . Ind. 

♦Genus METANOPEDIAS Brues 

Metanopedias Brues, 1910. Amer. Mus. Nat. Hist. Bui. 28: 79. 
Type : Metanopedias sicarius Brues. By monotypy. 
Insert in Platygasterinae before Trichacis Foerster, p. 709. 

rubicola (Ashmead), n. comb. Transf. from Trichacis Foerster, p. 709, with 
its synonyms (teste L. Masner). 



302 HYMENOPTEKA OF AMERICA NORTH OF MEXICO 

Genus TRICHACIS Foerster, pp. 709-710 
rubicola Ashmead. Transf. to Metanopedias Brues. 

Genus ERITRISSOMERUS Ashmead, p. 710 
noveboracensis Brues. Transf. to Platygaster Latreille, p. 710. 

Genus PLATYGASTER Latreille, pp. 710-715 

*Prosactogaster Kieffer, 1914. In Andre, Spec. Hym. Eur. Alg., v. 11, pp. 
352, 363. Syn. by Masner, 1965. Brit. Mus. Nat. Hist. Bui., Ent. 
Sup. 1: 135. 

Type: Platygaster lineatus Kieffer. Desig. by Muesebeck and 
Walkley, 1956. 
*Mi8ocyclops Kieffer, 1914, loc. cit. supra, pp. 353, 362. Syn. by Masner, 
1965, loc. cit. supra, p. 135. 

Type : Platygaster ornatus Kieffer. By monotypy. 
*Pareplmeces Kieffer, 1926. Das Tierreich, Lief. 48, pp. 563, 760. Syn. 
by Masner, 1965, loc. cit. supra, p. 135. 

Type: Epimeces ensifer Westwood. By substitution of Parepi- 
nieces for Epimeces Westwood, preocc. 

eryihropus (Ashmead), n. comb. Transf. from Metaclisis Foerster, p. 707 
{teste L. Masner). 

hyalinipennis (Ashmead). N. Y. Host: Caryomyia tubicola (O. S.) on hick- 
ory. 

nigripes (Ashmead), n. comb. Transf. from Monocrita Ashmead, p. 707 
(teste L. Masner). 

noveboracensis (Brues). Transf. from Eritrissomerus Ashmead, p. 710 {teste 
L. Masner). 

Genus EUXESTONOTUS Fouts, p. 715 

Remove from syn. under Xestonotidea Gahan. A valid genus containing 
all the species placed in Catalog under Xestonotidea, except andriciphila 
(Ashmead) which is transf. to Leptacis Foerster, of which Xestonotidea is a 
syn. 

Genus Soc/ogosfer Foerster, p. 715 

Syn. of Synopeas Foerster ( = Leptacis Foerster of Catalog in part) ; 
included spp. transf. to Synopeas. 

Genus SYNOPEAS Foerster, p. 716 

Remove from syn. under Leptacis Foerster, p. 716, and place as valid 
genus before Leptacis. 

Ectadius Foerster, 1856. Transf. here from syn. under Leptacis Foerster, 
p. 716. Syn. by Masner, 1964. Acta Soc. Ent. Cechosl. 61: 149. 

Polymecus Foerster, 1856. Transf. here from syn. under Leptacis Foer- 
ster, p. 716. Unnecessarily proposed as n. name for Ectadius 
Foerster. 

Sactogaster Foerster, 1856. N. syn, (teste L. Masner). All species listed 
in Catalog are transf. to Synopeas as n. combs.; where necessary 
ending is to be changed to neuter. 

Dolichotrypes Crawford and Bradley, 1911. Transf. here from syn. 
under Leptacis Foerster, p. 716. Syn. by Masner, 1964. Acta Soc, 
Ent. cechosl. 61: 149. 

Taxonomy: Ghesquiere, 1948. Soc. Ent. de Belg. Bui. et Ann. 84:44. — 
Masner, 1960. Rev. de Zool. et de Bot. Africaines 62: 3. — Masner, 
1965. Brit. Mus. Nat. Hist. Bui., Ent. Sup. 1: 140. 

abdominator (Fouts), n. comb. Transf. from Leptacis Foerster, p. 716 (teste 
L. Masner) , 

aciculatum (Fouts), n. comb, Transf. from Leptacis Foerster, p, 716 (teste L, 
Masner) . 



PLATYGASTERIDAE 303 

angustulum (Fouts), n. comb. Transf. from Leptacis Foerster, p. 716 (teste 

L. Masner). 
anomalivenfre (Ashmead), n. comb. Transf. from Sactogaster Foerster, p. 

716 {teste L. Masner). 

ashmeadii Dalla Torre. Transf. from Leptacis Foerster, p. 716 {teste L. 
Masner) . 

auripes (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 716 {teste L. 
Masner) . 

bradleyi (Fouts), n. comb. Transf. from Leptacis Foerster, p. 716 {teste L. 
Masner) . 

breviventre (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 716 
{teste L. Masner). 

carinator (Fouts), n. comb. Transf. from Leptacis Foerster, p. 716 {teste L. 
Masner) . 

cynipsiphilum (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 716 
{teste L. Masner). 

dubiosum (Fouts), n. comb. Transf. from Leptacis Foerster, p. 717 (teste L. 
Masner). 

flavicorne (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 717 (teste 
L. Masner). 

floridanum (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 717 
(teste L. Masner). 

*foutsi Masner, n. name. Replaces longiventre (Fouts). preocc, upon transf. 
from Sactogaster Foerster, p. 716. 

globatum (Fouts), n. comb. Transf. from Leptacis Foerster, p. 717 (teste L. 
Masner) . 

hopkinsi (Crawford and Bradley), n. comb. N. C, Fla. Transf. from Lep- 
tacis Foerster, p. 717 (teste L. Masner). 

howardii (Ashmead), n. comb. Transf. from Sactogaster Foerster, p. 71 
(teste L. Masner). 

incertum (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 717 (teste 
L. Masner). 

longiventre (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 717 
(teste L. Masner). 

maculipes (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 717 (teste 
L. Masner). 

mucronatum (Fouts), n. comb. Transf. from Sactogaster Foerster, p. 716 
(teste L. Masner). 

nigripes Ashmead. Transf. from Leptacis Foerster, p. 717 (teste L. Masner). 

pennsylvanicum (Fouts), n. comb. Transf. from Leptacis Foerster, p. 717 
(teste L. Masner). 

pubescens (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 717 (teste 
L. Masner). 

punctatiun (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 717 
(teste L. Masner). 

rufiscapus Ashmead. Transf. from Leptacis Foerster, p. 717 (teste L. 
Masner) . 

rugosiceps (KieflFer), n. comb. Transf. from Leptacis Foerster, p. 717 (teste 
L. Masner). 

striatifrons (Ashmead), n. comb. Transf. from Leptacis Foerster, p. 718 
(teste L. Masner). 



304 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

texanum (Fouts), n. comb. Transf. from Leptacis Foerster, p. 718 (teste L. 
Masner) . 

varipes (Harrington), n. comb. Transf. from Sactogaster Foerster, p. 716 
(teste L. Masner). 

Genus LEPTACIS Foerster, pp. 716-718 

Ectadius Foerster, 1856. Remove from syn.; place as syn. of Synopeas 
Forester (teste Masner, 1964. Acta Soc. Ent. cechosl. 61: 149). 

Synopeas Foerster, 1856. Remove from syn.; a valid genus (teste 
Masner, 1960. Rev. de Zool. et de Bot. Africaines 62:3). 

Anopedias Foerster, 1856. Remove from syn.; a valid genus without 
known Nearctic species (teste L. Masner). 

Xestonotus Foerster, 1856. Syn. by Masner, 1964, loc. cit. supra, p. 151. 

Polymecus Foerster, 1856. Remove from syn.; syn. of Anopedias (unne- 
cessarily proposed replacement name). 

DoUchotrypes Crawford and Bradley, 1911. Remove from syn.; place as 
syn. of Synopeas Foerster (teste L. Masner, 1964, loc. cit. supra, 
p. 149.) 

Xestonotidea Gahan, 1919. Syn. by Masner, 1964, loc. cit. supra, p. 6, 
151. 

Axestonotus Kieffer, 1926. Syn. by Masner, 1964, loc. cit. supra, p. 151. 

*Froleptacis Kieffer, 1926. Das Tierreich, Lief. 48, pp. xxi, 562, 632. Syn. 
by Masner, 1960, loc. cit. supra, p. 3. 

Type : Leptacis foersteri Kieffer. By orig. desig. 

*Anacoryphe Debauche, 1947. Soc. Ent. de Belg. Bui. et Ann. 83: 247. 
Treated as subgenus by Masner, 1960, loc. cit. supra, p. 3. 

Type: Anacoryphe orchymonti Debauche. By monotypy and 
orig. desig. 

*Mandraka Risbec, 1953. Inst. Sci. de Madagascar Mem. 3: 343. Treated 
as subgenus by Masner, 1960, loc. cit. supra, p. 3. 

Type; Mandraka pauliani Risbec. By monotypy. 

Taxonomy: Masner, 1960, loc. cit. supra, pp. 1-34 (revision of the Afri- 
can species). 

abdontinator Fouts. Transf. to Synopeas Foerster. 

aciculata Fouts. Transf. to Synopeas Foerster. 

andriciphila (Ashmead). Transf. from Xestonotidea Gahan, p. 716 (teste 
Ghesquiere, 1948. Soc. Ent. de Belg. Bui. et Ann. 84: 42). 

angustula Fouts. Transf. to Synopeas Foerster. 

ashmeadii (Dalla Torre) . Transf . to Synopeas Foerster. 

auripes (Ashmead). Transf . to Synopeas Foerster. 

bradleyi Fouts. Transf. to Synopeas Foerster. 

breviventris Ashmead. Transf. to Synopeas Foerster. 

carinator Fouts. Transf. to Synopeas Foerster. 

cynipsiphila Ashmead. Transf. to Synopeas Foerster. 

dubiosa Fouts. Transf. to Synopeas Foerster. 

flavicornis Ashmead. Transf. to Synopeas Foerster. 

floridana Ashmead. Transf. to Synopeas Foerster. 

globata Fouts. Transf. to Synopeas Foerster. 

hopkinsi (Crawford and Bradley). Transf. to Synopeas Foerster. 

incerta (Ashmead). Transf . to Synopeas Foerster. 

longiventris Ashmead. Transf. to Synopeas Foerster. 

maculipes (Ashmead). Transf. to Synopeas Foerster. 

nigripes (Ashmead). Transf . to Synopeas Foerster. 

pennsylvanica Fouts. Transf. to Synopeas Foerster. 



CHRYSIDIDAE 305 

pubescens Ashmead. Transf . to Synopeas Foerster. 
punctata Ashmead. Transf. to Synopeas Foerster. 
rufiscapus (Ashmead). Transf. to Synopeas Foerster. 
rugosiceps Kieffer. Transf. to Synopeas Foerster. 
striatifrons Ashmead. Transf. to Synopeas Foerster. 
texana Fouts. Transf. to Synopeas Foerster. 

Superfamily BETHYLOIDEA 
Family CHRYSIDIDAE 

Biology: Krombein, 1958. Amer. Ent. Soc. Trans. 84: 141-168, 2 pis. 
(spp. of coastal N. C). 

Genus OMALUS Panzer, pp. 718-719 

*Philoctetes Abeille de Perrin, 1870. Soc. Linn. Lyon Ann. 26, sup.: 26. 

Type: Holopyga (Philoctetes) cicatrix Abeille de Perrin. De- 

sig. by Ashmead, 1902. 

Diplorrhos Aaron, 1885. Amer. Ent. Soc. Trans. 12: 216. 

Type: Diplorrhos plicatus Aaron. By monotypy. 

*EUampus subg. Dictenulus Semenov, 1932. Soc. Ent. Ross. Hor. 42: 6. 

Type: EUampus (Dictenulus) specularis Semenov. Orig desig. 

Revision: Bohart and Campos, 1960. Ent. Soc. Amer. Ann. 53: 235-250, 
30 figs. 

*aeneus (Fabricius). Canada and U. S. south to Ga., N. Mex. and Ariz., 
chiefly in Canadian and Transition Zones, occasionally Upper 
Austral; Europe. Hosts: Pemphredon sp. and Passaloecus sp. in 
U. S.; Stigmus, Passaloecus and Pemphredon in Europe. 
(Onialus) 

Chrysis aeneus Fabricius, 1787. Mant. Ins. 1: 284. 

Onialus laeviventris Cresson (syn. by Bohart and Campos, 1960, loc. cit. 
supra, p. 238). 

Omalus diversus Aaron (syn. by Bohart and Campos, 1960, loc. cit. 
supra, p. 238). 

*auratus (Linnaeus). N. Y. south to S. C, Ohio, Ind. Hosts: Pemphredon 
(Cemonus) I. lethifer (Shuck.) and Passaloecus turiomim Dahlb. 
in U. S.; in Europe Pemphredon (P.) lugubris F., Pemphredon 
(Cemonus) unicolor F., Trypoxylon figulum F. and T. attenuatum 
Sm. — the latter two records are very dubious. (Pseudomalus) 

Sphex aurata Linnaeus, 1758. Syst. Nat., ed. 10, p. 572. 

Onialus sinuosus (Say) (syn. by Bohart and Campos, 1960, loc. cit. 
supra, p. 241. Note that O. janus is removed from syn. and revali- 
dated with O. corruscans (Norton) and O. purpuratus (Pro- 
vancher) as syns.). 

Taxonomy: Krombein, 1959. Brooklyn Ent. Soc. Bui. 54: 95-96. 

Biology: Thomas, 1962. Ent. News 73: 217-218. 

*butleri Bohart and Campos. Ariz., N. Mex., Utah, Colo., B. C, Ind., N. Y. 
(Omalus) 
Omalus (Omalus) butleri Bohart and Campos, 1960. Ent. Soc. Amer. 
Ann. 53: 240, figs. 6, 17. $, $. 

cressoni (Aaron). Oreg., Calif., Ariz., Colo., Utah, Mont., Nebr. Hosts: Xy- 
locelia occidentalis (Fox), Pemphredon giffardi (Roh.), Stigmus 
inordinatus Fox. (Diplorrhos) 
Biology: Powell, 1964. Wasmann Jour. Biol. 21: 172-173, 

diversus Aaron. Syn. of aeneus (Fabricius). In orig. citation for diversus, 
change § to 5. 



306 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*downeyi Bohart and Campos. Oreg., Calif., Nev., chiefly in Canadian Zone. 
(Diplorrhos) 
Omalus (Diplorrhos) downeyi Bohart and Campos, 1960. Ent. Soc. 
Amer. Ann. 53: 246, figs. 5, 8. 5, ?• 

glomeratus (du Buysson). Calif, to Wash., east to Idaho, Colo., N. Hex. 
Host: Stigmus inordinatus Fox. (Omalus) 

*granti Bohart and Campos. Calif., Ariz., N. Hex., Nev., Colo. (Diplorrhos) 
Omalus (Diplorrhos) grand Bohart and Campos, 1960. Ent. Soc. Amer. 
Ann. 53: 247, fig. 19. $,9. 

intermedius (Aaron). Mont., Md. Host: Xylocelia virginiana Roh. Transf. 
from Elampus Spinola. (Diplorrhos) 

Biology: Krombein, 1963. Ent. Soc. Wash. Proc. 65: 264. 

iridescens (Norton). Throughout Canada and U. S., in Canadian, Transition 
and Austral Zones. Hosts: Stigmus inordinatus Fox, S. ameri- 
canus Pack., Diodontus trisculcus Fox. (Omalus) 

janus (Haldeman). Canada and U. S. south to Calif., Ariz., N. C, chiefly in 

Canadian and Transition Zones. Host: Pemphredon (P.) concolor 

Say, P. (P.) errans Roh., P. (Cemonus) giffardi (Roh.). {Pseudo' 

malus) 
Omalus sinuosus (Say) of Catalog, in part. O. janus not a syn. (teste 

Bohart and Campos, 1960, loc. cit. supra, p. 242) . 
Elampus coruscans Norton (syn. by Bohart and Campos, 1960, loc. cit. 

supra, p. 242). Note that coruscans was misspelled in orig. cit. 

under sinuosus in Catalog. 
Elampus purpuratus Provancher (syn. by Bohart and Campos, 1960, loc. 

cit. supra, p. 242) . 
Omalus semicircularis Aaron (syn. by Bohart and Campos, 1960, loc. cit. 

supra, p. 242). 

*krombeini Bohart and Campos. Ariz., southern Calif. (Diplorrhos) 

Omalus (Diplorrhos) krombeini Bohart and Campos, 1960. Ent. Soc. 
Amer. Ann. 53: 247, figs. 3, 13. $,9. 

laeviventris Cresson. Syn. of aeneus (Fabricius). 

*macswaini Bohart and Campos. Alaska, Canada and U. S. south to Calif., 
Ariz, and Ga. Hosts: Stigmus americanus Pack., Pemphredon 
(Cemonus) I. lethifer (Shuck.), ? P. (C.) harbecki Roh. (Pseudo- 
malus) 

Omalus (Pseudomalus) macstcaini Bohart and Campos, 1960. Ent. Soc. 
Amer. Ann. 53: 244, fig. 22. $,9. 

Biology: Krombein, 1960. Ent. News 71: 32-33. 

plicatus (Aaron). Alaska, Canada, western and northern U. S. south to 
Calif., N. Mex., Minn., Mich., N. Y. (Diplorrhos) 

semicircularis Aaron. Syn. of janus (Haldeman). 

seminudus (Aaron). Alaska, B. C. south to Calif., Idaho, Ont. Transf. from 
Elampus Spinola. (Diplorrhos) 

sinuosum (Say). Syn. of auratus (Linnaeus). N. B.: O. janus Haldeman is 
removed from syn. and revalidated, with O. corruscans Norton) 
and O. purpuratus (Provancher) as synonyms. (Pseudomalus) 

speculum (Say). Wash.. Oreg., Calif., Colo. (Diplorrhos) 

*telfordi Bohart and Campos. Tex. to southern Calif., Nev. (Diplorrhos) 

Omalus (Diplorrhos) telfordi Bohart and Campos, 1960. Ent. Soc. 
Amer. Ann. 53: 249, figs. 14, 27. $, 9. 

*irilobatus Bohart and Campos. B. C. and Idaho south to Calif., Ariz, and 
Colo., in Canadian, Transition and Upper Sonoran Zones. Host: 
Pemphredon (Cemonus) giffardi ("Roh.) . (Pseudomalus) 
Omalus (Pseudomalus) trilobatus Bohart and Campos, 1960. Ent. Soc. 
Amer. Ann. 53: 245, figs. 4, 15, 25, 28. $,9. 



CHRYSIDIDAE 307 

variatus (Aaron). Western N. Amer. in Canadian, Transition and Upper 
Sonoran Zones. Host: Stigmus inordinatus Fox. (Diplorrhos) 

Genus ELAMPUS Spinola, p. 719 

intermedius (Aaron). Transf. to Omalus, p. 718. 

seminudus (Aaron). Transf. to Omalus, p. 718. 

Genus Dip/orrhos Aaron, p. 719 

Transferred with component species to Omalus as a subgenus (teste 
Bohart and Campos, 1960, loc. cit. supra, p. 236). 

Genus HEDYCHRIDIUM Abeille de Perrin, p. 720 

amabile Cockerell. Ariz., Nev. 

fletcheri Bodenstein. Ala., B. C. 

polygoni Rohwer. Transf. to Hedychrum Latreille, p. 720. 

Genus HEDYCHRUM Latreille, pp. 720-721 

continuum Aaron. Fla. 

polygoni (Rohwer). Transferred from Hedychridium Abeille de Perrin, p. 720 
{teste K. V. Krombein who examined type). 

violaceum Brulle. Host: ? Cerceris halone Bks. In citation of tciltii Cresson, 
change $ to 5- 

Genus CHRYSURA Dahlbom 
( = Chrysis subg. Chryiura Dahlbom of Catalog, p. 721) 

Raised to generic rank (Krombein, 1963. Ent. News 74: 149). 

doriae Gribodo. Hosts: Trypoxylon (T.) bidentatum Fox, T. (T.) sculleni 
Sandh. Kans., Tex.; Mexico. 

*kyrae Krombein. Md., Va., Mo. (?), Ind. Host: Osmia (O.) I. lignaria Say. 
Chrysura kyrae Krombein, 1963. Ent. News 74: 150. $,9. 
Biology: Krombein, 1963, loc. cit. supra, p. 152. 

pacifica (Say). Delete host record. 

tola Aaron. B. C. 

CHRYSIS subgenus CHRYSOGONA Foerster, pp. 721-722 

densa Cresson. Delete Pseudodynerus quadrisectus (Say) as a host and add 
Pseudomasaris vespoides (Cr.), P. zonalis (Cr.) and P. edwardsii 
(Cr.). 
Biology: Hicks, 1927. Canad. Ent. 59: 78.— Hicks, 1929. Canad. Ent. 
61: 122. 

perpulchra Cresson. Host: Sphex aberti (Hald.). 
Biology: Hicks, 1932. Canad. Ent. 64: 145-151. 

verticalis Patton. Host: Trypoxylon backi Sandh. 

Biology: Thomas, 1962. Amer. Midland Nat, 67: 365-366.— Thomas, 
1963. Mich. Acad. Sci., Arts, Letters, Papers 48: 127-130. 

CHRYSIS subgenus DICHRYSIS Lichtenstein, p. 722 
annulipes (Mocsary). Syn. of bigeloviae Cockerell. 

bigeloviae Cockerell. Add $ after orig. descr. 

mesillae Cockerell. Tex., Okla.; Mexico (Zacatecas). Add 9 after orie. 
Chrysis (Dichrysis) annulipes (Mocsary) (listed in Sup. 1), n. syn., 

based on type examination by K. V. Krombein. 

descr. 



308 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

CHRYSIS subgenus CERATOCHRYSIS Cooper, Sup. 1, p. 95 

enhuycki Cooper. Host: Leptochilus republicanus zendaloides (Robt.). 
Biology: Krombein, 1959. Ent. News 70: 17-23. 

CHRYSIS subgenus TRICHRYSIS Lichtenstein, p. 722 

mucronata Brulle. Host: Trypoxylon {Trypargiluvi) t. tridentatum Pack. 
Biology: Parker, 1962. Pan-Pacific Ent. 38: 140. 

parvula Fabricius. Add Preocc. after citation for virens Cresson. 
Biology: Krombein, 1958. Amer. Ent. Soc. Trans. 84: 144. 

CHRYSIS subgenus CHRYSIS Linnaeus, pp. 722-725 

Taxonomy: Bohart, 1962. Acta Hym. 1: 361-375, 25 figs, (sexdentate 
forms) . 

aaroni Mocsary. 

Chrysis (Pyrin) nana Mocsary (syn. by Bohart, 1962. Acta Hym. 1: 375). 
Taxonomy: Krombein, 1958. Amer. Ent. Soc. Trans. 84: 146. 

alabamensis Mocsary. Transfer to *Neochry8is Linsenmaier as a syn. of pan- 
amensis (Cameron). 

*aridula Bohart. Ariz., southern Calif. 

Chrysis aridula Bohart, 1962. Acta Hym. 1: 364, figs. 1, 5, 17. $, 9. 

*arizonica Bohart. Ariz., Tex. Host: Stenodynerus (Parancistrocerus) tol- 

Chrysis arizonica Bohart, 1962. Acta Hym. 1: 366, figs. 2, 10, 18, 19. $, 9. 

*astralia Bohart. Calif., Nev., Utah, Ariz., N. Mex.; Mexico (Baja Califor- 
nia, Zacatecas) . 
Chrysis astralia Bohart, 1964. Biol. Soc. Wash. Proc. 77: 233, figs. 8, 13. 
$, 9. 

*bequaerii Bohart. D. C, Ga., Ark., Mo., Ind., S. Dak. to Tex., N. Mex. 

Chrysis hequaerti Bohart, 1962. Acta Hym. 1: 366, figs. 7, 9, 20, 21, 25. 

5, 9. 

*cembricola Krombein. Pa., Md., D. C, Va., W. Va., N. C. Host: Sijm- 
morphus canadensis (Sauss.). 
Chrysis (Chrysis) cembricola Krombein, 1958. Ent. Soc. Wash. Proc. 60: 

53, fig. 1. 9, S. 
Biology: Krombein, 1958, loc. cit. supra, pp. 57-58. 

clara Cresson. S. Dak., Colo., Wash. Transf. from Chrysis subg. Pyria Lepe- 
letier & Serville, p. 725. 
Taxonomy: Bohart, 1962. Acta Hym. 1: 367. 

*clypeata Mocsary. Syn. of lauta Cresson. 

coerulans Fabricius. Hosts: Ancistrocerus spilogaster Cam., A. tigris cy- 
tainus (Cam.), A. tuberculoceplialus siitterianus (Sauss.), Euo- 
dynerus leucomelas (Sauss.), E. f. foraminatus (Sauss.), E. f. 
scutellaris (Sauss.), Stenodynerus acarophonis Boh. 
Biology: Parker, 1962. Pan-Pacific Ent. 38: 140.— Medler, 1964. Ent. 
News 75: 26-27.— Medler, 1964. Ent. Soc. Wash. Proc. 66: 
209-215. 

cognata Gribodo. Syn. of intricata Brulle. 

*coloradica Bohart. Calif., Nev., Idaho. Host: Anthidium collectum Huard. 
Chrysis coloradica Bohart, 1964. Biol. Soc. Wash. Proc. 77: 224. N. 

name for pulcherrima Cresson, preocc, questionably placed in syn. 

of lauta Cresson in Catalog. 

*crotema Bohart. N. Mex., Ariz.; Mexico (Zacatecas). 

Chrysis crotema Bohart, 1964. Biol. Soc. Wash. Proc. 77: 228, figs. 5, 18. 
19. 5, 9. 

decepta Rohwer. Syn. of pattoni Aaron. 



CHRYSIDIDAE 309 

derivata du Buysson. Que., N. H., Mass., N. J., Pa., Md., N. C, 111., Minn., 
Mo. Hosts: Ancistrocerus catskill lialophilus Vier., Euodynerus 
foraminatus scutellaris (Sauss.). 
Taxonomy: Krombein, 1958, loc. cit. supra, p. 160, fig. 7. 

dorsalis Aaron. Delete Fla. from distribution. Host: Photopsis sp. 
Taxonomy: Krombein, 1958, loc. cit. supra, p. 146, fig. 5. 

equidens Viereck. N. C, Tex., Kans., Nebr. 

Taxonomy : Krombein, 1958, loc. cit. supra, pp. 145-146, fig. 6. 

fabricii Mocsary. Delete from North American fauna {teste Bohart, 1962. 
Acta Hym. 1: 375). 

fahifica Buysson. Syn. of lauta Cresson. 

fuscipennis Brulle. Calif. (Sacramento and Stanislaus Counties). Host: Sce- 
liphron caementarium (Dru.). 
Biology: Stage, 1960. Pan-Pacific Ent. 36: 191-195, 1 pi. 

inacquidens Dahlbom. Over most of U. S., N. Y. to Wash, and south into 
Mexico (Chihuahua). Hosts: Euodynerus megaera (Lep.), E. fo- 
raminatus scutellaris (Sauss.) and E. foraininatus apopkensis 
(Robt.). Transf. from Chrysis subg. Pyria Lepeletier & Serville, 
p. 725. 

Chrysls (Pyria) texana Gribodo (syn. by Bohart, 1962. Acta Hym. 1: 
368, fig. 14). 

Biology: Krombein, 1958, loc. cit. supra, pp. 161-163 (recorded as fabri- 
cii Mocs.).— Moore and Parker, 1962. Pan-Pacific Ent. 38: 14. 

inflata Aaron. Host: Ancistrocerus lineativentris Cam., A. tuberculiceps sut- 
terianus (Sauss.) 

intricata Brulle. Tex. (Brownsville) ; Mexico south to French Guiana. 
Transf. from Chrysis subg. Pyria Lepeletier & Serville, p. 725. 
*Chrysis sitiidti Dahlbom, 1845. Hym. Europaea 2: 317. 9. 
Chrysis (Pyria) cognata Gribodo (syn. by Bohart, 1962. Acta Hym. 1: 

368, fig. 16). 
""Chrysis proximo Cameron, 1888. Biol. Cent-Amer., Hym., v. 1, p. 465. 9. 
*Chrysis schulthessi Mocsary, 1889. Monog. Chrysid., p. 572. $. 
'^Chrysis aenescens Mocsary, 1889. Monog. Chrysid., p. 577. 9. 
*Chrysis aperta du Buysson, 1897. Ann. Soc. Ent. France 66: 559. $. 

kahli Viereck. Syn. of propria Aaron. 

laminifera BischofF. Ark. 

Taxonomy: Krombein, 1958, loc. cit. supra, p. 149, fig. 3. 

lauta Cresson. N. C, Ga., Miss., Tex., Kans., Nebr., Colo., N. Mex., Ariz.; 
Mexico (Morelos, Jalisco). Transfer pulcherrima Cresson to syn. 
of *coloradica Bohart. Host: Anthidium porterae Ckll. 
Chrysis (Chrysis) falsi fica Buysson (syn. by Bohart, 1964. Biol. Soc. 

Wash. Proc. 77: 224). 
*Chrysis (Tetrachrysis) clypeata Mocsary, 1889. Monog. Chrysid., p. 393. 

$. (Syn. by Bohart, 1964, loc. cit. supra, p. 224.) 
Biology: Custer, 1928. Ent. News 89: 123-134. 

tnontana Aaron. 

Taxonomy: Krombein, 1958, loc. cit. supra, p. 145. 

nana Mocsary. Syn. of aaroni Mocsdry, Transf. from Chrysis subg. Pyria 
Lepeletier & Serville. p. 725. 

nitidula Fabricius. Host: Ancistrocerus c. catskill (Sauss.), Euodynerus 
f. foraminatus (Sauss.). 
Biology: Medler, 1964. Ent. Soc. Wash. Proc. 66: 209-215. 

*oraria Bohart. Ariz., Tex. 

Chrysis oraria Bohart, 1962. Acta Hym. 1: 370. $, 9. 

pattonella Viereck. Syn. of propria Aaron. 



310 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

pattoni Aaron. Host: Ancistrocerus spilogaster Cam. 

Chrysis (Chrysis) decepta Rohwer (syn. by Krombein, 1957. Ent. News 
68: 191). 
pellucidula Aaron. N. Y., Pa., Calif. Host: Trypoxylon t. tridentatum Pack. 

Biology: Hicks, 1934. Univ. Colo. Studies 21: 267. 

*praestans du Buysson. Ky., Colo, to Tex., west to Wash, and Calif.; West 
Indies (Grenada). 
Chrysis praestans du Buysson, 1898. Rev. Ent. Caen 17: 144. 9. 
Taxonomy: Bohart, 1962. Acta Hym. 1: 370, fig. 8. 

*proIata Bohart. Kans., Wis. 

Chrysis prolata Bohart, 1964. Biol. Soc. Wash. Proc. 77: 229. ?. 

propria Aaron. 

Chrysis (Chrysis) kahli Viereck (syn. by Krombein, 1957, loc. cit. supra, 

p. 191). 
Chrysis (Chrysis) pattonella Viereck (syn. by Krombein, 1957, loc. cit. 

supra, p. 191). 

provancheri Schulz. S. Dak., B. C. 

*riva!is Bohart. Calif., Oreg., Idaho, Wyo., Utah, Colo., Nev. 

Chrysis rivalis Bohart, 1964. Biol. Soc. Wash. Proc. 77: 229, figs. 4, 22, 
23. S, 9. 

serrata Taylor. Wash., Calif., Nev., N. Hex., Utah, Mich. Transf. from 
Chrysis subg. Pvria Lepeletier & Serville, p. 725. 
Taxonomy: Bohart, 1962. Acta Hym. 1: 371, fig. 12. 

smaragdula Fabricius, Ariz. Host: Monobia quadridens (L.). Transf. from 
Chrysis subg. Pyria Lepeletier & Serville, p. 725. 
Biology: Krombein, 1958, loc. cit. supra, p. 163. 
Taxonomy: Bohart, 1962. Acta Hym. 1: 371, fig. 13. 

*8tenodytieri Krombein. N. Y. south to Fla., Ind., Mo., Kans. Hosts: Steno- 
dynerus ammonia histrionalis (Robt.), S. krombeini Boh., S. linca- 
tifrvns Boh. 

Chrysis (Chrysis) stenodyneri Krombein, 1958. Amer. Ent. Soc. Trans. 
84: 151, figs. 1, 2, 8. 9, $. 

Biology: Krombein, 1958, loc. cit. supra, pp. 155-160. 

*tensa Bohart. Calif., Ariz., N. Mex., Tex., Utah; Mexico (Sonora, Chihua- 
hua). 
Chrysis tensa Bohart, 1964. Biol. Soc. Wash. Proc. 77 : 230, figs. 2, 14, 15. 

$, 9. 

tenuicornis Taylor. Oreg., Calif., N. Mex.; Mexico (Baja California). 
Taxonomy: Bohart, 1962. Acta Hym. 1: 373, figs. 3, 15. 

texana Gribodo. Syn. of inaequidens Dahlbom. 

*tularensis Bohart. Calif. (Tulare Co.). 

Chrysis tularensis Bohart, 1962. Acta Hym. I: 373, figs. 4, 6. $,9. 

*vagabuiida Bohart. Calif., Oreg., Wash., Nev., Utah. 

Chrysis vagabutida Bohart, 1964. Biol. Soc. Wash. Proc. 77: 225, figs. 24, 

25. $, 2. 

*venustella Bohart. Calif., Nev., Wash. 

Chrysis venustella Bohart, 1964. Biol. Soc. Wash. Proc. 77: 235, figs. 9, 

12. $, 9. 

*vibex Bohart. Calif., Ariz., N. Mex., Tex.; Mexico (Chihuahua, Nuevo Leon, 
Jalisco). 
Chrysis vibex Bohart, 1964. Biol. Soc. Wash. Proc. 77: 231, figs. 1, 7, 11, 
16. $, 9. 

*wasbaueri Bohart. Generally distributed over U.S., Mass. to Wash., south to 
Mexico (Chihuahua, Nueva Leon). 
Chrysis wasbaueri Bohart, 1962. Acta Hym. 1 : 372, figs. 11, 22, 23, 24. 5, 9. 



CHRYSIDIDAE 311 

•Tcerophila Bohart. Calif. 

Chrysis xerophila Bohart, 1964. Biol. Soc. Wash. Proc. 77: 226, figs. 3, 
20, 21. S, ?. 

Chrysis subgenus Pyria Lepeletier and Serville, p. 725 

Delete this heading and transfer component species to Chrysis subg. 
Chrysis Linnaeus, pp. 722-725 (teste Bohart, 1962. Acta Hym. 1: 362). 

^Genus NEOCHRYSIS Linsenmaier 

Pleurocera subg. Neochrysis Linsenmaier, 1959. Schweiz. Ent. Ges. Mitt. 
32: 74. 

Type : Chrysis punctatissima Spinola. Orig. desig. 

Place after Chrysis subg. Chrysis Linnaeus, p. 725. Raised to generic 
rank (teste Bohart and Menke, 1963. Univ. Calif. Pubs. Ent. 30: 108). 

*panamensis (Cameron). Md. ; Panama. 

Chrysis panamensis Cameron, 1888. Biol. Cent.-Amer., Hym. 1, p. 464, pi. 

20, fig. 8. 
Chrysis (Chrysis) ahibamensis Mocsary (transf. from Chrysis and syn. by 

Bohart and Menke, 1963, loc. cit. supra, p. 108). 
Biology and Taxonomy: Krombein, 1958, loc. cit. supra, pp. 147-149, fig. 

4. 

Genus PARNOPES Latreille, pp. 725-726 

Revision: Telford, 1964. Univ. Calif. Pubs. Ent. 36: 1-42, 5 pis., 1 text 
fig. 

arizonensis Viereck. Syn. of concinnus Viereck. 

*borregoensis Telford. Southern Calif. 

Parnopes borregoensis Telford, 1964. Univ. Calif. Pubs. Ent. 36: 9, pi. 1, 

fig. 2. $, 9. 

*6oMt/ieryi Brethes. Syn. of edwardsii (Cresson). 

chrj soprasinus Smith. N. Y. south to Fla., west to Wis. and Tex. 

concinnus Viereck. Calif, to western Tex., Nev. ; Mexico (Baja California, 
Jalisco). 
Parnopes arizonensis Viereck (syn. by Telford, 1964, loc. cit. supra, p. 

11). 
*Parnopes digued du Buysson, 1904. Rev. d'Ent. Caen 23: 274. $. (Syn. 
by Telford, 1964, loc. cit. supra, p. 11.) 

diadema Viereck. Syn. of fulvicornis fulvicornis Cameron. 

*digueti du Buysson. Syn. of concinnus Viereck. 

edwardsii (Cresson). Change edwardsi in specific heading in Catalog to ed- 
wardsii. Alaska to California, east to the Rocky Mts. ; Mexico 
(Baja California). Host: Steniolia obliqua (Cr.). 
*Parnopes houtheryi Brethes, 1902. Mus. Nac. Buenos Aires An. 8 (3) : 

287. 9, $. (Syn. by Telford, 1964, loc. cit. supra, p. 6.) 
Parnopes hageni Viereck (syn. by Telford, 1964, loc. cit. supra, p. 6). 
Parnopes henshawi Viereck (syn. by Telford, 1964, loc. cit. supra, p. 6). 
Biology: Evans and Gillaspy, 1964. Amer. Midland Nat. 72: 277, fig. 14. 

excurvatus Viereck. Calif., Nev., Ariz. In orig. citation change 9 to "9" = $. 

festivus Cockerell. Ariz, to western Tex.; Mexico (Chihuahua). 

*fulvicornis atlanticus Krombein. Md., Va., N. C, ? Fla. Host: ? Microhem- 

bex monodonta (Say). 
Parnopes westcottii atlanticus Krombein, 1958. Amer. Ent. Soc. Trans. 

84: 164, 9, $. (Made a subsp. of fulvicornis by Telford, 1964. loc. 

cit. supra, p. 19.) 
Biology: Krombein, 1958, loc. cit. supra, pp. 166-167. 



312 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*fuIvicornis fulvicornis Cameron, changed status. Idaho and Iowa south to 

Calif, and Tex., and further routh to Guatemala. 
Parnopes fulvicornis Cameron, 1888. Biol. Cent.-Amer. Hym., v. 1, p. 

466, pi. 20, figs. 1, la. $. (Reduced to subsp. rank, teste Telford, 

1964, loc. cit. supra, p. 16.) 
Parnopes westcottii Melander and Brues (syn. by Telford, 1964, loc. cit. 

supra, p. 16), 
Parnopes diadema Viereck (syn. under typical westcottii by Krombein, 

1958. Amer. Ent. Soc. Trans. 84: 164). 
Parnopes taeniata Viereck (syn. by Telford, 1964, loc. cit. supra, p. 16), 

/lageni Viereck. Syn. of edwardsii (Cresson). 

/lens/iaici Viereck. Syn. of edwardsii (Cresson). 

taeniata Viereck. A syn. of fulvicornis fulvicornis Cameron, 

tcestcottii Melander and Brues. Syn. of fulvicornis fulvicornis Cameron, 

Genus CLEPTES Latreille, pp. 726 

purpuratus Cresson. Host: Neodiprion sp. 

Biology: Dahlsten, 1961. Canad. Ent. 93: 192-193. 

*seniiauratus (Linnaeus). New Jersey (Metuchen) ; introduced from Europe, 
Sphex semiaurata Linnaeus, 1761. Fauna Suec, Ed. 2, p. 413. $. 
*Cleptes pallipes Lepeletier, 1806. Mus. Hist. Nat. Ann. 7: 119, pi, 8, fig, 

1. ?, 

*Cleptes diana Mocsary, 1889. Monog. Chrysid., p, 46, 3. 

Genus MESITIOPTERUS Ashmead, p. 726 

*floridensis Krombein. Fla. (Gainesville). 

Mesitiopterus floridensis Krombein, 1960. Amer, Ent. Soc. Trans. 86: 28, 
fig. 6. $, 9. 

kahlii Ashmead, Ariz. (Portal), 

*Genus MICROSEGA Krombein 

Microsega Krombein, 1960, Amer. Ent, Soc. Trans. 86: 31. 

Type : Microsega bella Krombein. By monotypy and orig, desig. 

Place in Amiseginae (Sup. 1, p. 96) after Mesitiopterus. 

*bella Krombein. Tex. (Cameron Co.), 

Microsega bella Krombein, 1960, Amer, Ent. Soc. Trans, 86: 32, figs, 1, 

2, 5, 9, 

Genus ADELPHE Mocs6ry, Sup. 1, p. 96 

^anisomorphae Krombein, Ga., Fla,, La, Host: Anisom.orpha ferruginea 
(Beauv.), eggs. 
Adelphe anisomorphae Krombein, 1960, Amer, Ent. Soc. Trans, 86: 35, 
fig. 3. $, 9. 

Family BETHYLIDAE 

Taxonomy: Evans, 1964, Mus. Compar. Zool. Bui. 132: 1-222, 144 figs, 
(synopsis of American spp.), 

♦Subfamily EPYRINAE 
*Tribe CEPHALONOMIINI 

Genus CEPHALONOMIA Westwood, p. 727 

*Cephalonomia subg. Cephaloderma Hoffer, 1936, Festchr, Embrik 
Strand, vol, 1, p. 459, (Syn. by Evans, 1964, loc. cit. supra, p, 159,) 
Type : Cephalonomia (Cephaloderma) strandi Hoif er. By 
monotypy. 



BETHYLIDAE 313 

gallicola (Ashmead). 

*Cephalonomia (Cephaloderma) strandi Hoffer, 1936. Festschr. Embrik 
Strand, vol. 1, p. 460. ?. (Syn. by Evans, 1964, loc. cit. supra, p. 
159.) 

hyalinipennis Ashmead. Mass. Host: Conophthorus coniperda (Schw.). 

*perpusilla Evans. Oreg., Calif., Ariz.; Mexico (Baja California, Nayarit). 
Host: Ciid beetle larvae in fungi. 
Cephalonomia perpusiUa Evans, 1963- Psyche 70: 152. $,9. 

utahensis Brues. Hosts: Conophthorus radiatae Hopk. and Ernobius punctu- 
latus Fall. 
Biology: Schaefer, 1962. Ent. Soc. Amer. Ann. 55: 574-575, fig. 6. 

Genus PLASTANOXUS Kieffer 

Transferred from p. 731. 

chittendenii (Ashmead). Transcont. in U. S. and southern Canada. 

*incompletus Evans. Ariz. (Mesa). 

Plastauoxus incompleius Evans, 1964. Mus. Compar. Zool. Bui. 132: 

152. ?. 

laevis (Ashmead). Brazil (?). 

westwoodi (Kieffer). Widely distributed in U. S.; Africa; probably cosmopoli- 
tan. 

*Tribe SCLERODERMINI 

♦Genus NESEPYRIS Bridwell 

Nesepyris Bridwell, 1920. Hawaiian Ent. Soc. Proc. 4: 309. 
Type : I\esep)ris etva Bridwell. By monotypy. 

Place before Glenosema Kieffer, p. 727. 

*floridanus Evans. Fla.; Mexico. 

Nesepyris virginianus Evans, 1964. Mus. Compar. Zool. Bui. 132: 164, 
111, 112. 9. 

*virginianus Evans. Md., Va. 

Nesepyris virginianus Evans, 1964. Mus. Compar. Zool. Bui. 132: 164, 
fig. 110. $. 

Genus GLENOSEMA KiefFer, p. 727 

californicus (Bridwell). Transfer to syn. of Bethylus decipiens (Pro- 
vancher), p. 732. 

*crandalli Evans. Ariz. (Santa Catalina Mts.). 

Glenosema crandalli Evans, 1964. Mus. Compar. Zool. Bui. 132: 170, fig. 
L16. 9. 

Genus SCLERODERMA Latreille, p. 728 

*Neoscleroflerma Kieffer, 1905. Soc. Sci. Bruxelles Ann. 29: 106. (Syn. 
by Evans, 1964, loc. cit. supra, p. 174.) 

Type: Ateleopterus virginiensis Ashmead. Desig. by Kieffer, 
1914. 

*Tribe EPYRINI 

Genus LAELIUS Ashmead, p. 728 
utilis Cockerell. Ont. 

EPYRIS subgenus EPYRIS Westwood 
( = Genus Epyris Westwood, pp. 728-729) 
*Muellerella Saussure, 1892. In Grandidier, Hist. Nat. Madagascar 20: 



314 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

pi. 25, fig. 20. (Syn. by Evans, 1964, loc. cit. supra, p. 104.) 
Type: Muellerella amabilis Saussure. By monotypy. 
*Parepyris Kieffer, 1913 (not of Brethes, 1913). Lab. Zool. Gen. e Agr. 
Portici Bol. 7: 108. (Syn. by Evans, 1964, loc. cit. supra, p. 104). 
Type: Epyris interruptus Kieffer. Desig. by Kieffer, 1914. 
Psilepyris Kieffer, 1913. (Syn. by Evans, 1964, loc. cit. supra, p. 104.) 

indivisus Kieffer. Transf. from Psilepyris Kieffer, p. 730 {teste Evans, 1964, 
loc. cit. supra, p. 104). 

*vierecki Krombein. Md. (Plummers Island). 

Epyris vierecki Krombein, 1962. Biol. Soc. Wash. Proc. 75: 1. $,9. 

Nomen Nudum in Epyris Westwood 

*Epyris monticola Ashmead in Cockerell, 1889. Colo. Biol. Assn., 10th 
Kept., [p. 2] (probably published originally in Custer County 
Courant newspaper). 

=^Genus BAKERIELLA Kieffer 

Bakeriella Kieffer, 1910. Soc. Ent. France Ann. 78: 288. 

Type: Bakeriella flavicornis Kieffer. By monotypy. 

Place before Rhabdepyris Kieffer, p. 729. 

*floridana Evans. Fla. (Paradise Key). 

Bakeriella floridana Evans, 1964. Mus. Compar. Zool. Bui. 132: 134, fig. 
84. $, ?. 

Genus RHABDEPYRIS Kieffer, pp. 729-730 

Chlorepyris Kieffer, 1913. (Syn. by Evans, 1964, loc. cit. supra, p. 93.) 

aeneiceps (Ashmead). Transf. to Anisepyris Kieffer, p. 730. Delete Ga. from 

distribution. 
analis (Cresson). Transf. to Anisepyris Kieffer, p. 730. 

columbianus (Ashmead). Transf. to Anisepyris Kieffer, p. 730. 

fulgens (Brues). Transf. from Chlorepyris Kieffer, p. 730 (teste Evans, 1964, 
loc. cit. supra, p. 95). 

grandis (Ashmead). Transf. to Anisepyris Kieffer, p. 730. 

haemorrhoidalis Kieffer. Transf. to Holepyris Kieffer, p. 730. 

occidentalis (Ashmead). Transf. to Anisepyris Kieffer, p. 730. 

zeae Turner and Waterston. Transf. to syn. of Holepyris sylvanidis 
(Brethes), p. 730. 

Genus HOLEPYRIS Kieffer, p. 730 

*Holepyris subg. Rysepyris Kieffer, 1906. In Andre, Spec. Hym. Eur. 
Alg., V. 9, p. 341. (Syn. by Evans, 1964, loc. cit. supra, p. 140.) 
Type: Holepyris (Rysepyris) nutnidicus Kieffer. Desig. by 
Kieffer, 1914. 
*Misepyris Kieffer, 1913. Lab. Zool. Gen. e Agr. Portici Bol. 7: 108. 
(Syn. by Evans, 1964, loc. cit. supra, p. 140.) 

Type: Holepyris remotus Kieffer. Desig. by Kieffer, 1914. 
*Parepyris Brethes, 1913 (not of Kieffer, 1913). Buenos Aires Mus. Nac. 
de Hist. Nat. An. 24: 87. (Syn. by Evans, 1964, loc. cit. supra, p. 
140.) 

Type: Parepyris sylvanidis Brethes. By monotypy. 

haemorrhoidalis (Kieffer). Transf. from Anisepyris Kieffer, p. 729 (teste 
Evans, 1964, loc. cit. supra, p. 143). 

■^sylvanidis (Brethes). Widespread in U. S.; cosmopolitan. Host: Oryzae- 
philus surinamensis (L.). 
Parepyris sylvanidis Brethes, 1913. Buenos Aires Mus. Nac. de Hist. 
Nat. An. 24: 87. $,9. 



BETHYLIDAE 315 

Rhabdepyris zeae Turner and Waterston (syn. by Evans, 1964, loc. cit. 
supra, p. 143). 

Genus Psilepyrh Kieffer, p. 730 
Syn. of Epyris subg. Epyris Westwood, p. 728. 
indivisus (Kieffer). Transf. to Epyris Westwood, p. 729. 

Genus Chforepyris KiefFer, p. 730 

Transfer to synonymy of Rhabdepyris Kieffer, p. 729. 
fulgens (Brues). Transfer to Rhabdepyris Kieffer, p. 729. 

Genus Xanthepyris Kieffer, p. 730 
A synonym of Pseudisobrachium Kieffer, pp. 731-732. 

Genus Acrepyrh Kieffer, p. 730 

Transfer to Pristocera Klug, p. 731, as a valid subgenus {teste Evans, 
1963. Mus. Compar. Zool. BuL 129: 248). 

reticulatus (Kieffer). Transf. to Pristocera Klug, p. 731, as a syn. of armifera 
(Say). 

Genus ANISEPYRIS Kieffer, pp. 730-731 

*Rhabdepyris subg. Lophepyris Evans, 1959. Ent. Soc, Wash. Proc. 61: 
201. (Syn. by Evans, 1964, loc. cit. supra, p. 96.) 

Type: Rhabdepyris (Lophepyris) bridtvelli Evans. By orig. 
desig. 
Revision: Evans, 1959. Ent. Soc. Wash. Proc. 61: 97-120, 28 figs.— 
Evans, 1959. Ent. Soc. Wash. Proc. 61: 201-204. 

aeneiceps (Ashmead). Transf. from Rhabdepyris Kieffer, p. 729 (teste 
Evans, 1959, loc. cit. supra, p. 107). 

analis (Cresson). Fla. and Tex. to Tenn. and N. C. Transf. from Rhabde- 
pyris Kieffer, p. 729 {teste Evans, 1959, loc. cit. supra, p. 118). 

*arizonicus Evans. Ariz, (Santa Rita Mts.). 

Anisepyris arizonicus Evans, 1959. Ent. Soc. Wash. Proc. 61: 112, fig. 
15. 9. 

*bradleyi (Evans). Tex. (Wharton). 

Rhabdepyris (Lophepyris) bradleyi Evans, 1959. Ent. Soc. Wash. Proc. 
61: 204. ?. 

*bridwelli (Evans). Tex. (Brownsville, Laredo) ; Mexico. 

Rhabdepyris (Lophepyris) bridtvelli Evans, 1959. Ent. Soc. Wash. Proc. 
61: 202. $, $. 

eoluinbianus (Ashmead). Mass. west to Kans. south to Fla. and Tex. 
Transf. from Rhabdepyris Kieffer, p. 729 {teste Evans, 1959, loc. 
cit. supra, p. 104). 
Anisepyris pulchellus Fouts (syn. by Evans, 1959, loc. cit. supra, p. 104). 

*dietrichorum Evans. Ariz. (Arivaca in Pima Co.). 

Anisepyris dietrichorunt Evans, 1959. Ent. Soc. Wash. Proc. 61: 118, fig. 

21. ?. 

*gibbosifrons Evans. N. J., Fla. 

Anisepyris gibbosifrons Evans, 1959. Ent. Soc. Wash. Proc. 61: 116, figs. 
23,25,26. $, $. 

grandis (Ashmead). Transf. from Rhabdepyris Kieffer, p. 729 {teste Evans, 
1959, loc. cit. supra, p. 108). 

*laticeps Evans. Southeastern Ariz. 

Anisepyris laticeps Evans, 1959. Ent. Soc. Wash. Proc. 61: 113, figs. 18, 
19,24. 9, $. 



316 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

occidentalis (Ashmead). Ariz, and southern Calif, to Idaho and Wash. 
Transf. from Rhabdepyris Kieffer, p. 729 (teste Evans, 1959, loc. 
cit. supra, p. 109). 
Anisepyris punctaticeps Kieffer (syn. by Evans, 1959, loc. cit. supra, p. 
109). 

pulchellus Fonts. Syn. of columbianus (Ashmead). 

punctaticeps Kieffer. Syn. of occidentalis (Ashmead). 

subvioiaceus Kieffer. Fla., Tex. and N. Mex. north to Pa., Mich., Wyo., B. C. 

*williamsi Evans. Ariz, and Calif, north to Oreg., Idaho and Utah. 

Anisepyris williamsi Evans, 1959. Ent. Soc. Wash. Proc. 61: 106, fig. 9. 
?, 5. 

Genus Plasfanoxus KiefFer, p. 731 

Transfer with component species to *Tribe Cephalonomiini and place 
after Cephalonomia Westwood, p. 727 (teste Evans, 1964, loc. cit. supra, p. 
148). 

♦Subfamily PRISTOCERINAE 

Genus PRISTOCERA Klug, p. 731 

PRISTOCERA subgenus ACREPYRIS KiefFer 

Reduced to subg. of Prislocera Klug (teste Evans, 1963. loc. cit. infra 
p. 248). 

*Pristocera subg. Neopristocera Yasumatsu, 1955. Fac. Agr. Kyushu Univ. 
Jour. 10: 248. 

Type: Pristocera japonica Yasumatsu. Orig. desig. 
Revision: Evans, 1963. Mus. Compar. Zool. Bui. 129: 241-290, 60 figs. 

armifera (Say). N. H. and Vt. south to Fla., west to S. Dak., Colo, and Tex. 
Host: Melanotus sp. 
Scleroderma thoracica Westwood not a syn. of atra Klug but of armifera 

(teste Evans, 1963, loc. cit. supra, p. 256). 
Acrepyris reticulatus Kieffer (syn. by Evans, 1963, loc. cit. supra, p. 256). 

atra Klug. N. C. to Fla., west to N. Mex. Remove thoracica (Westwood) 
from syn. of atra and transfer it to syn. of armifera (Say) . 

*bridwelli Evans. Ark. (DoddCity). 

Pristocera (Acrepyris) bridtcelli Evans, 1963. Mus. Compar. Zool. Bui. 
129: 267. $, $. 

*caIifornIea Evans. Calif., Utah, Wyo. 

Pristocera (Acrepyris) cali/ornica Evans, 1963. Mus. Compar. Zool. Bui. 
129: 263. $. 

*chihuahua Evans. Ariz.; Mexico (Chihuahua, Zacatecas). 

Pristocera (Acrepyris) chihuahua Evans, 1963. Mus. Compar. Zool. Bui. 
129: 284. $. 

*cockerelIi Evans. Tex. to Ariz.; Mexico (Sonora, Oaxaca). 

Pristocera (Acrepyris) cockerelli Evans, 1963. Mus. Compar. Zool. Bui. 
129: 264. $, 9. 

*fraterna Evans. N. C. to Fla., Kans. 

Pristocera (Acrepyris) fraterna Evans, 1963, Mus. Compar. Zool. Bui. 
129: 261. $, 9 (?). 

hyalina Brues. La., Tex., N. Mex. 

*Genus APENESIA Wesfwood 

Apenesia Westwood, 1874. Thesaurus Ent. Oxon., p. 170. 

Type: Apenesia amazonica Westwood. Desig. by Westwood, 
1881. 
*Aeluroides Tullgren, 1904. Ark. Zool. 1: 428. 

Type : Aeluroides sjostedti Tullgren. By monotypy. 



BETHYLIDAE 317 

*Propristocera Kieffer, 1905. In Andre, Spec. Hym. Eur. Alg., v. 9, p. 
247. 

Type: Propristocera interrupta Kieffer, Desig. by Kieffer, 
1914. 
*Cleis!epyris Kieffer, 1910. Soc. Ent. France Ann. 79: 48. 

Type: Cleisiepyris punctatus Kieffer, 1910. Desig. by Kieffer, 
1914. 
*Dipristocera Kieffer, 1914. Das Tierreich, Lief. 41, p. 471. 

Type: Dipristocera microchela Kieffer. By montoypy. 
*Neopristocera Benoit, 1957. Explor. Pare Nat. Albert, Mission DeWitte, 
fasc. 88, p. 44. Preocc. 

Type: ISeopristocera triloba Benoit. Orig. desig. 
Revision: Evans, 1963. Mus. Compar. Zool. Bui. 130: 249-359, 138 figs. 

Place after Pristocera Klug, p. 731. 

*chiricahua Evans. Ariz.; Mexico (Mexico). 

Apenesio chiricahua Evans, 1963. Mus. Compar. Zool. Bui. 130: 306, figs. 

57.60, 69, SO, 84. $. 

*cochise Evans. Ariz. (Cochise Co.). 

Apenesia cochise Evans, 1963. Mus. Compar. Zool. Bui. 130: 296, figs. 
27,32. $. 

*dissomphaloides Evans. Ariz. (Pinal Co.), 

Apenesia dissomphaloides Evans, 1963. Mus. Compar. Zool. Bui. 130: 
297, figs. 37, 42. $. 

*exilis Evans. Ariz., Calif. 

Apenesia exilis Evans, 1963. Mus. Compar. Zool. Bui. 130: 293, figs. 28, 
30,33,34. $. 

*martiiii Evans. Fla. (Manatee Co.). 

Apenesia martini Evans, 1963. Mus. Compar. Zool. Bui. 130 : 294, fig. 29. 

$. 

*mohave Evans. Calif.; Mexico (Baja California). 

Apenesia mohave Evans, 1963. Mus. Compar. Zool. Bui. 130: 308, figs. 

58.61. $. 

*palliflula Evans. Ariz. (Cochise Co.). 

Apenesia pallidula Evans, 1963. Mus. Compar. Zool. Bui. 130: 300, figs. 
38, 43,46, 50. $. 

*parapolita Evans. Ark., La., Ala., and Ga., north to 111., W. Va. and N. Y. 

* Propristocera polita Evans, 1958. Ent. Soc. Wash. Proc. 59: 294, figs. 2, 

6. $. Preocc. in Apenesia. 
Apenesia parapolita Evans, 1963. Mus. Compar. Zool. Bui. 130: 322 {$), 

347 (9), figs. 81, 96, 127, 136. N. name. 

*pima Evans. Ariz. (Tucson). 

Apenesia pima Evans, 1963. Mus. Compar. Zool. Bui. 130: 292, figs. 26, 
31,35,36. $. 

♦Genus PARASCLERODERMA KiefFer 

Parascleroderma Kieffer, 1904. Mus. Civ. Stor. Nat. Genova Ann. 41; 
376. 

Type: Parascleroderma fulviceps Kieffer. Orig. desig. 

Place on p. 731 after *Apenesia Westwood. 

*insolita (Evans). Tex. (Brownsville). 

Apenesia insolita Evans, 1963. Mus. Compar. Zool. Bui. 130: 350, figs. 
130, 138. 9. 

Genus PSEUDISGBRACHIUM Kieffer, pp. 731-732 

*Monepyris Kieffer, 1905. Soc. Sci. Bruxelles Ann. 29: 101, 124. 

Type: Epyris halidayi Westwood (= suhcyaneum Haliday). 
By monotypy. 



318 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*Plutobethylu8 Kieffer, 1910. Soc. Ent. France Ann. 79: 51. 

Type: Plutobethylus distans Kieffer. Orig. desig. 
*Lyssepyris Kieffer, 1913. Portici Lab. Zool. Gen. e Agr. Bol. 7 : 108. 

Type: Holepyris flavicornis Kieffer. By monootypy. 
Xantepyris Kieffer, 1913. Portici Lab. Zool. Gen. e Agr. Bol. 7: 108. 

Type : Epyris flaviventris Kieffer. By monotypy. 
*Lissepyris (!) Kieffer, 1914. Das Tierreich 41: 236. Typ. error. 
Xa/if/iepyris Kieffer, 1914. Das Tierreich 41: 417. Emend. 
*Pari8obrachium Kieffer, 1914. Das Tierreich 41: 424. 

Type: Rhabdepyris (?) albipes Kieffer. By monotypy. 

Revision: Evans, 1961. Mus. Compar. Zool. Bui. 126: 211-318, 5 pis. 

Members of this genus have been taken in ant nests, but it is not certain 
whether they are parasitic on the ant larvae or on larvae of associated 
myrmecophilous beetles. 

agile Whittaker. Change orig. cit. to Roy. Ent. Soc. London Trans. 76: 386, 
pi. 13, fig. 4. $. Syn. of prolongatum (Provancher). 

*anomalum Evans. N. J., Va., Fla., 111. 

Pseudisobrachium anomalum Evans, 1961. Mus. Compar. Zool. Bui. 126: 
311, figs. 8, 54,64,67. $. 

*apache Evans. Ariz. (Santa Cruz, Gila and Pinal Counties). 

Pseudisobrachium apache Evans, 1961. Mus. Compar. Zool. Bui. 126: 
299, fig. 46. $. 

*arenarium Evans. N. J., Pa. (?), N. C, Ga., Fla. (?), Ala., Miss., Mo. (?), 
111. Associated with ponerine ants. 
Pseudisobrachium arenarium Evans, 1961. Mus. Compar. Zool. Bui. 126: 
263, figs. 20, 39, 57. 5, ? (?)• 

'^ashmeadi Evans. Mass., Ont., Mich, and Kans., south to Fla., Ala. and La. 
Associated with formicine and dolichoderine ants. 
Pseudisobrachium ashmeadi Evans, 1961. Mus. Compar. Zool. Bui. 126: 
275, figs. 1, 23, 38, 58, 63. $,9 (?). 

carbonarium (Ashmead), Md. south to Ga., Ala., W. Va., Ky. (?), 111., Kans,, 
N. Dak. (?). 
Taxonomy: Evans, 1961, loc. cit. supra, p. 271, figs. 23, 37. 5> ? (?)• 

*caroIinianuiii Evans. S. C, Fla. 

Pseudisobrachium carolinianum Evans, 1961. Mus. Compar. Zool. Bui. 
126: 254. $. 

*castaneum Evans. Calif. (San Diego). 

Pseudisobrachium castaneum Evans, 1961. Mus. Compar. Zool. Bui. 126: 
248, fig. 56. $. 

*conianche Evans. Ariz., Tex. 

Pseudisobrachium comanche Evans, 1961. Mus. Compar. Zool. Bui. 126: 
298, fig. 59. $. 

*crassum Evans. Tex. (Brownsville, San Antonio). 

Pseudisobrachium crassum Evans, 1961. Mus. Compar. Zool. Bui. 126: 
239, fig. 13. $. 

*emarginatuin Evans. Tex. (Kerrville). 

Pseudisobrachium emarginatum Evans, 1961. Mus. Compar. Zool. Bui. 
126: 303, fig. 49. $. 

flavicoxis Fouts. Syn. of rufiventre (Ashmead). 

flavinervis Fouts. Calif., Nev., Ariz., Tex. 

Taxonomy: Evans, 1961, loc. cit. supra, p. 305, figs, 6, 31, 38, 62. $, 9 (?). 

flaviventre (Kieffer). Del. to Fla., west to Tex., Kans., 111. Associated with 
myrmicine ants. 
Taxonomy: Evans, 1961, loc. cit. supra, p. 288, figs. 41, 60. $,9 (?). 



BETHYLIDAE 319 

*fout8i Evans. Tex., N. Mex. (?), Ariz. (?), Calif. (?) ; Mexico (Coahuila). 
Pseudisobrachium foutsi Evans, 1961. Mus. Compar. Zool. Bui. 126: 300, 
figs. 29,61. $. 

*gibbosum Evans. N. Mex. (Rodeo). 

Pseudisobrachium gibbosum Evans, 1961. Mus. Compar. Zool. Bui. 126: 

255. $. 

*krombeini Evans. N. Mex. (Albuquerque, White Sands). 

Pseudisobrachium krombeini Evans, 1961. Mus. Compar. Zool. Bui. 126: 
287, figs. 27, 50. $. 

*macrops Evans. Tex. (Cameron and Hidalgo Counties). 

Pseudisobrachium macrops Evans, 1961. Mus. Compar. Zool. Bui. 126: 
309. $. 

m,agnum (Ashmead). Syn. of prolongatum (Provancher). 

mandibulare (Ashmead). Syn. of prolongatum (Provancher). 

*inatthewsi Evans. Tex. (Van Horn, El Paso (?)). 

Pseudisobrachium matthewsi Evans, 1961. Mus. Compar. Zool. Bui. 126: 
249, fig. 36. $,9 (?). 

*minimuin Evans. N. Mex., Ariz: 

Pseudisobrachium minimum Evans, 1961. Mus. Compar. Zool. Bui. 126: 

280, fig. 24. $. 

*minutissimuni Evans. N. Mex., Ariz.; Mexico (Morelos, Baja California), 
Guatemala. 
Pseudisobrachium minutissimum, Evans, 1961. Mus. Compar. Zool. Bui. 
126, fig. 26. $. 

montanum (Ashmead). Syn. of prolongatum (Provancher). 

myrmecophilum (Ashmead). Syn. of prolongatum (Provancher). 

*navajo Evans. Ariz. (Coconino and Yavapai Counties). 

Pseudisobrachium navajo Evans, 1961. Mus. Compar. Zool. Bui. 126: 

283, fig. 25. $. 

*obscurum Evans. Tex., Ariz.; Mexico (Chihuahua, Baja California). 

Pseudisobrachium obscurum Evans, 1961. Mus. Compar. Zool. Bui. 126: 
258, figs. 16,36. 5, 9 (?)• 

*occidentale Evans. Calif., Ariz. Associated with dolichoderine ants. 

Pseudisobrachium occidentale Evans, 1961. Mus. Compar. Zool. Bui. 
126: 244, figs. 14, 55. $,9 (?). 

*otiosum Evans. Ariz. (Superior). 

Pseudisobrachium otiosum Evans, 1961. Mus. Compar. Zool. Bui. 126: 

256. $. 

*palli(lum Evans. Ariz. (Wellton). 

Pseudisobrachium pallidum Evans, 1961 Mus. Compar. Zool. Bui. 126: 

258, fig. 17. $. 

paucipunctatum Fonts. Utah, Calif. 

Taxonomy: Evans, 1961, loc. cit. supra, p. 316. 9. 

♦persimile Evans. Calif., B. C. (?). 

Pseudisobrachium persimile Evans, 1961. Mus. Compar. Zool. Bui. 126: 

284, fig. 2. $, 9 (?). 

prolongatum (Pi'ovancher) . Change 9 to "9" = 5 in orig. cit. N. B. and N. 

S. west to B. C., and Wash., south to N. C, Ky., 111. and Iowa. 

Associated with formicine and myrmicine ants. 
Pseudisobrachium magnum (Ashmead) (syn. by Evans, 1961, loc. cit. 

supra, p. 266, figs. 21, 22, 39, 57). 
Pseudisobrachium myrmecophilum (Ashmead), $ (9 misdet.) (syn. by 

Evans, 1961, loc. cit. supra, p. 266). 
Pseudisobrachium mandibulare (Ashmead), 9 ($ misdet.) (syn. by 

Evans, 1961, loc. cit. supra, p. 266), 



320 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Pseudisobrachium montanum (Ashmead) 9(5 misdet.) (sjm. by Evans, 

1D61, loc. cit. supra, p. 266). 
Pseudisobrachium rugosulum Fouts (syn. by Evans, 1961, loc. cit. supra, 

p. 266). 
Pseudisobrachium agile Whittaker (syn. by Evans, 1961, loc. cit. supra, 

p. 266). 

puncticeps Fouts. Syn. of rufiventre (Ashmead). 

*pusilluin Evans. La. (Shreveport) , Ark. (?). 

Pseudisobrachium pusillum Evans, 1961. Mus. Compar. Zool. Bui. 126: 
297. $, 9 (?). 

*rectangulatum Evans. Tex., Kans., Nebr.; Mexico (Nuevo Leon). 

Pseudisobrachium rectangulatum Evans, 1961. Mus. Compar. Zool. Bui. 
126: 304. $. 

rufiventre (Ashmead). Mass. and N. Y. south to Fla. and La. Associated 

with formicine ants. 
Pseudisobrachium flavicoxis Fouts (syn. by Evans, 1961, loc. cit. supra, 

p. 293, figs. 5, 40. $,9). 
Pseudisobrachium puncticeps Fouts (syn. by Evans, 1961, loc. cit. supra, 

p. 293). 

rugosulum Fouts. Syn. of prolongatum (Provancher). 

*texanuiti Evans. Tex. (Harlingen, Brownsville, Wharton, San Antonio). 

Pseudisobrachium texanum Evans, 1961. Mus. Compar. Zool. Bui. 126: 
240, figs. 48, 53. $. 

Genus DISSOMPHALUS Ashmead, p. 732 

*Thaumatepyris Kieffer, 1910. Soc. Ent. France Ann. 79: 47. 

Type: Thaumatepyris punctatus Kieffer. By monotypy. 
*Glenobethylus Kieffer, 1910. Soc. Ent. France Ann. 79: 50. 

Type: Glenobethylus montanus Kieffer. By monotypy. 
*Parecitopria Ogloblin, 1930. Soc. Ent. Arg. Rev. 3: 15. 

Type : Parecitopria azarai Ogloblin. By monotypy. 
Taxonomy: Evans, 1962. Ent. Soc. Wash. Proc. 64: 65-78, 4 figs, (re- 
vised key). 

altivolans Evans. Ala., 111., Tex., Ariz., Calif. 

Taxonomy: Evans, 1962, loc. cit. supra, p. 76. 9. 

apertus Kieffer. N. Y., Pa., 111. 

*arizonicus Evans. Ariz. (Portal). 

Dissomphalus arizonicus Evans, 1962. Ent. Soc. Wash. Proc. 64: 70, fig. 
3. S. 

barberi Evans. Maine, N. C. 

foveolatus (Brown and Cheng). Ga., Ark., 111. Delete Ariz, from distribu- 
tion. 

kansanus Evans. Pa., Fla., 111. 

nigrescens Evans. Ark. ? 

Taxonomy: Evans, 1962, loc. cit. supra, p. 75 (putative 9). 

xanthopus Ashmead. Ky., Miss., Ark. Ariz. 

♦Subfamily BETHYLINAE 

Genus BETHYLUS Lafreille, p. 732 

Digoniozus Kieffer (syn. by Evans, 1962, loc. cit. infra, p. 1). 
Revision: Evans, 1962. Breviora, No. 150, 12 pp., 3 figs., 1 map. 

amoenus Fouts. N. W. T. south to Alta., east through Wis. and 111. to N. Y., 

Maine, and N. S. 

brachypterus Whittaker. Syn, of decipiens (Provancher). 



DRYINIDAE 321 

decipiens (Provancher) . Alaska south to central Calif, eastward to N. Y., 

Maine, and N. S. 
Digoniozus oregonensis (Ashmead) (syn. by Evans, 1962, loc. cit. supra, 

p. 7). 
Glenoseina californicus (Bridwell) (syn. by Evans, 1962, loc. cit. supra, 

p. 7). 
Bethylus brachypterus Whittaker (sjTi. by Evans, 1962, loc. cit, supra, p. 

7). Change 9 5 to 9. 
Bethylus flavicornis Whittaker (syn. by Evans, 1962, loc. cit. supra, p. 
7). 
^aricorwis Whittaker. Syn. of decipiens (Provancher). 

Genus D/goniozus Kieffer, p. 732 

Place as a synonym of Bethylus Latreille, p. 732, and transfer oregon- 
ensis (Ashmead) to synonymy of decipiens (Provancher). 

Genus GONIOZUS Foerster, pp. 732-733 

*castaneicolor Evans. "United States." 

* Bethylus castaneus Kieffer, 1907. Berlin Ent. Ztschr. 51: 295. 9. 

Preocc. in Gonioziis. . 
Goniozus casianeicolor Evans, 1964. Mus. Compar. Zool. Bui. 132: 201. 
N. name. 

Genus Per/s/ero/a Kieffer, pp. 733-734 
Transfer to synonymy of Parasierola Cameron, p. 734. 

Genus PARASIEROLA Cameron, p. 734 

Perisierola Kieffer (syn. by Evans, 1964, loc. cit. supra, p. 195). 

Remove parens from names of authors of alutacea, gracilicornis, and 
punctaticeps, and place parens around names of authors of breviceps, 
cellularis, and emigrata. 

bicarinata Brues. Transfer to *Prosierola Kieffer which follows. 

"^distinguenda Kieffer. Calif.; Nicaragua. 

Parasierola distinguenda Kieffer, 1908. In Wytsman, Gen. Ins., fasc. 76, 
p. 14. 

*Genus PROSIEROLA Kieffer 

Prosierola Kieffer, 1905. In Andre, Spec. Hym. Eur. Alg., v. 9, p. 243. 
Type: Epyris (?) nasalis Westwood. By monotypy. 

Place after Parasierola Cameron, p. 734. 

bicarinata (Brues). Ga., Fla., Tex. Transf. from Parasierola Cameron, p. 
734 {teste Evans, 1964, loc. cit. supra, p. 185). 

Family DRYINIDAE 

Genus MESODRYINUS Kieffer, p. 1034 

*crawfordi Krombein. Md. (Plummers Island). 

Mesodryinus crancfordi Krombein, 1962. Biol. Soc. Wash. Proc. 75: 3. 9. 

Genus PSILODRYINUS Kieffer, p. 1035 

typhlooybae (Ashmead). Tex. Host: Metcalfa pridnosa (Say). 

Biology: Dean and Bailey, 1961. Jour. Econ. Ent. 54: 1104-1106, 2 figs. 

Genus EUCAMPTONYX Perkins, p. 1037 
secundus Fenton. Md., Kans. 

Genus GONATOPUS Ljungh, pp. 1037-1038 

*curriei Krombein. Md. (Plummers Island). 

Gonatopus curriei Krombein, 1962. Biol. Soc. Wash. Proc. 75: 4. 9. 



322 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus EPIGONATOPUS Perkins, p. 1039 

plesius Fenton. Ont. Host: Macrosteles fascifrons (Stal). 
Biology: George, 1959. Canad. Ent. 91: 266. 

Genus AMPULICOMORPHA Ashmead, p. 1043 

confusa Ashmead. Wis. Host: nymphs of Epiptera floridae (Wlkr.) on rotten 
pine and oak logs. 
Biology: Bridwell, 1958. Ent. Soc. Wash. Proc. 60: 23-26. 

Genus EMBOLEMUS Westwood, p. 1043 

Change citation for Embolemus to London and Edinb. Phil. Mag. and 
Jour. Sci. 2: 444. 

Family TRIGONALIDAE 

Genus LYCOGASTER Shuckard, p. 735 

pullata puUata Shuckard. Ga. Host: Telea polyphemus Crmr., probably sec- 
ondary. 

pullata nevadensis (Cresson). Host. Euodynerus foraminatum scutellaris 
(Sauss.). 

Genus BAREOGONALOS Schuiz, p. 735 

canadensis (Harrington). Calif. Host: Vespula (Dolichovespula) arenaria 
(F.). 
Biology: Stage and Slobodchikoff, 1962. Pan-Pacific Ent. 37(2): 97-98. 

Superfamily SCOLIOIDEA 

Family TIPHIIDAE 

Genus TIPHIA Fabricius, pp. 735-739 

Taxonomy: Allen, 1961. Amer. Ent. Soc. Trans. 87: 1-20 (key to de- 
scribed North American spp.). 

Tiphia femorata Fabricius and T. morio Fabricius were introduced from 
Europe and liberated in New York in 1956 and 1957 for control of 
Amphimallon majalis (Raz.), the European chafer. 

arida Malloch. Vt., Pa. 

atlantis Allen. 

Taxonomy: Allen, 1961, loc. cit. supra, p. 16. $. 

*bakeri Allen. Wash., Nev. 

Tiphia hakeri Allen, 1961. Amer. Ent. Soc. Trans. 87: 12. $. 
dentata Allen. Syn. of micropunctata Allen. 

fulvicauda Cameron. 

Taxonomy: Allen, 1962. Amer. Ent. Soc. Trans. 88: 24 (type redescrip- 
tion). 

hollowayi Allen. Syn. of infossata Allen. 

illinoensis Robertson. 

Tiphia voaldenii Viereck (syn. by Allen, 1961, loc. cit. supra, p. 18). 

incisurata Malloch. 

Taxonomy: Allen, 1961, loc. cit. supra, p. 20. 

*indistincta Allen. W. Va. (Kanawha Sta.). 

Tiphia indistincta Allen, 1961. Amer. Ent. Soc. Trans. 87: 14. 9. 

infossata Allen. 

Tiphia hollowayi Allen (syn. by Allen, 1961, loc. cit. supra, p. 19). 



TIPHIIDAE 323 

*irfla Allen. Fla. 

Tiphia irfla Allen, 1961. Amer. Ent. Trans. 87: 10. $,9. 

jaynesi Allen. Md. 

micropunctata Allen. 

Tiphia dentaia Allen (syn. by Allen, 1961, loc. cit. supra, p. 18). 

*inon»ana Allen. Mont. 

Tiphia montana Allen, 1961. Amer. Ent. Soc, Trans. 87: 13. $. 

nevadana Cameron. 

Taxonomy: Allen, 1962, loc. cit. supra, p. 22 (redescription of type). 

*pennsylvanica Allen. Pa., N. J., Del., Fla., La. 

Tiphia pennsylvanica Allen, 1961. Amer. Ent. Soc. Trans. 87: 15. 9. 

puncticeps Cameron. 

Taxonomy: Allen, 1962. Amer. Ent. Soc. Trans. 88: 23 (type redescrip- 
tion). 

relativa Viereck. 

Tiphia winnemanae Malloch (syn. by Allen, 1961, loc. cit. supra, p. 17). 

*tempeae Allen. Ariz. (Tempe)i 

Tiphia tempeae Allen, 1961. Amer. Ent. Soc. Trans. 87: 11. $. 

texensis Malloch. 

Taxonomy: Allen, 1961, loc. cit. supra, p. 19. 

vulgaris Robertson. 

Taxonomy: Allen, 1961, loc. cit. supra, p. 19. 

tcaldenii Viereck. Syn. of iilinoensis Robertson. 

tcinnemanae Malloch. Syn. of relativa Viereck. 

Genus NEOTIPHIA Malloch, p. 739 
( = Neo/ipfiia subg. Neofiphia Malloch, of Catalog) 

Revision: Allen and Krombein, 1964. Amer. Ent. Soc. Trans. 89: 241- 270. 

"'anima Allen and Krombein. N. Mex., Ariz. 

Neotiphia anima Allen and Krombein, 1964. Amer. Ent. Soc. Trans. 89: 
262. 9. 

*barbata Allen and Krombein. Ariz. (Continental, Tucson, Douglas). 

Neoiiphia barbata Allen and Krombein, 1964. Amer. Ent. Soc. Trans. 
89: 264. $. 

carinata Krombein. Syn. of novemexicana Allen. 

cockerelli Allen, chanjred status. Tex., N. Mex. 

Neotiphia (Neotiphia) crawfordi Allen of Catalog, in part. N. cockerelli 

Allen is a valid sp. {teste Allen and Krombein, 1964, loc. cit. supra, 
p. 255, fig. 50). 

conspicua Allen. Syn. of crawfordi Allen. 

crawfordi Allen. Tex. Remove cockerelli Allen, a valid sp., from synonymy. 
JSeotiphia (Neotiphia) conspicua Allen (syn. by Allen and Krombein, 
1964, loc. cit, supra, p. 257, figs. 28, 41, 42). 

cristata Allen. Transf. to Krombeinia Pate. 

novomexirana Allen. 

Neotiphia carinata Krombein (syn. by Allen and Krombein, 1964, loc. 
cit. supra, p. 251). 

*parma Allen and Krombein. Ariz., N. Mex.; Mexico (Mexico, Aguascalientes, 
Zacatecas). 
Neotiphia parma Allen and Krombein, 1964. Amer. Ent. Soc. Trans. 89: 
267, figs. 7, 54. $. 

pima Pate. Kans. and Colo, to Tex., N. Mex. and Ariz. 



324 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*rioverdci Allen and Krombein. Ariz. (Del Rio Verde). 

Neotiphia rioverflei Allen and Krombein, 1964. Amer. Ent. Soc. Trans. 
89: 263. 9. 

sulcata (Roberts). Mich., Iowa and S. Dak. south to Tex. and Ariz.; Mexico 
(Jalisco). 
Neotipbia iKronibeinia) sulcata (Roberts). Transf. from Kronibeinia 

Pate {teste Allen and Krombein, 1964, loc. cit. supra, p. 245). 
*l\eotiphia mexicana Allen (syn. by Allen and Krombein, 1964, loc. cit. 
supra, p. 245). 

waltoni Allen. Ariz.; Mexico (Chihuahua, Durango). 

Genus KROMBEINIA Pate, changed status, p. 739 
{zzNeotiphia subg. Krombeinia Pate, of Catalog) 

Revision: Allen and Krombein, 1964. Amer. Ent. Soc. Trans. 89: 214-241. 

cristata (Allen). Ariz., N. Mex. 

Neotiphia (ISeotiphia) cristata Allen. Transf. from Neotiphia Malloch 
(teste Allen and Krombein, 1964, loc. cit. supra, p. 238, figs. 3, 29). 

*jonesi Allen and Krombein. Ariz. (Douglas, Chiricahua, Huachuca Mts., 
Oracle). 
Kronibeinia jonesi Allen and Krombein, 1964. Amer. Ent. Soc. Trans. 
89: 239, figs. 6, 65. $, 9. 

sulcata (Roberts). Transf. to Neotiphia Malloch. 

Genus PARATIPHIA Sichel, pp. 739-740 

*aequalis atrata Allen. Ariz. The typical subspecies occurs in Baja Cali- 
fornia, Mexico. 
Paratiphia aequalis atrata Allen, 1963. Amer. Ent. Soc. Trans. 88: 228, 
figs. 5, 10, 15, 16. $, 9. 

algonquina Viereck. Pa. 

*asotinae Allen. Wash. (Asotin). 

Paratiphia asotinae Allen, 1963. Ent. News 74: 222. $, 9. 

bakeri Cameron. 

Taxonomy: Allen, 1962, loc. cit. supra, p. 28 (type redescription). 

*belfragei Allen. Tex. (Cypress Mill, Kerrville). 

Paratiphia belfragei Allen, 1963. Ent. Soc. Amer. Ann. 56: 575, figs. 

1, 2 c, d. $, 9. 

*bradleyi Allen. Tex. (Uvalde). 

Paratiphia bradleyi Allen, 1963. Amer. Ent. Soc. Trans. 88: 231, figs. 

2, 7, 12. $. 

*bridwelli Allen. Kans. (Dickinson Co.). 

Paratiphia bridwelli Allen, 1963. Amer. Ent. Soc. Trans. 88: 231, figs. 
1, 6, 11, 19. $. 

claripennis Cameron. 

Paratiphia ormshyensis Cameron (syn. by Allen, 1962, loc. cit. supra, p. 
29). 

*eph!ppiata Allen. Western Tex., N. Mex., Colo. 

Paratiphia ephippiata Allen, 1963. Ent. Soc. Amer. Ann. 56 : 574, figs. 1, 
2 a, b, f. S, 9. 

fuscinerva Cameron. 

Taxonomy: Allen, 1962, loc. cit. supra, p. 31 (type redescription). 

fortistriolata Cameron. Sjm. of robusta Cameron. 

insueta Allen. Tex. (Ft. Davis Mts.). 

Paratiphia insueta Allen, 1963. Ent. Soc. Amer. Ann. 56: 576, figs. 1, 
2 e. $. 



TIPHIIDAE 325 

intermedia Cameron, Wash. 

Taxonomy: Allen, 1962, loc. cit. supra, p. 32 (type redescription). — 
Allen, 1963, loc. cit. supra, p. 229, figs. 4, 9, 14, 17 (redescription 
S, 9). 

♦mitrhelli Allen. Tex., N. Mex. 

Paratiphia mitchelH Allen, 1963. Amer. Ent. Soc. Trans. 88: 232, figs. 3, 
8, 13, 18. $. 

neomexicana Cameron. 

Taxonomy: Allen, 1962, loc. cit. supra, p. 31 (type redescription). 

nevadensis Cameron. 

Paratiphia parvula Cameron (syn. by Allen, 1962, loc. cit. supra, p. 27). 

ormsbyensis Cameron. Syn. of claripennis Cameron. 

parvula Cameron. Syn. of nevadensis Cameron. 

robusta Cameron. Oreg., Wash., Idaho, Nev. 

Paratiphia fortistriolata Cameron (syn. by Allen, 1962, loc. cit. supra, p. 
33). 

texana Cameron. 

Taxonomy: Allen, 1962, loc. cit. supra, p. 25 (type redescription). 

%aripunetata Cameron. 

Taxonomy: Allen, 1962, loc. cit. supra, p. 30 (type redescription). 

*verna Allen. Calif., Nev. 

Paratiphia verna Allen, 1963. Ent. News 74: 219. $,9. 

Genus BRACHYCISTIS Fox, pp. 742-745 
aequalis Fox. Transf. to Acanthetropis Wasbauer. 
idiotes Cockerell. Transf. to Acanthetropis Wasbauer. 
noctivaga Bradley. Transf. to Acanthetropis Wasbauer. 
normalis Malloch. Transf. to Acanthetropis Wasbauer. 

♦Genus ACANTHETROPIS Wasbauer 

Acanthetropis Wasbauer, 1958. Pan-Pacific Ent. 34: 139. 

Type: Acanthetropis lamellatus Wasbauer. Orig. desig. 

Place after Colocistis Krombein, p. 745. 

aequalis (Fox). Transf. from Brachycistis, p. 742 (teste Wasbauer, 1958, 
loc. cit. supra, p. 142). 

idiotes (Cockerell). Transf. from Brachycistis, p. 743 (teste Wasbauer, 1958, 
loc. cit. supra, p. 142). 

noctivaga (Bradley). Transf. from Brachycistis, p. 744 (teste Wasbauer, 
1958, loc. cit. supra, p. 142). 

normalis (Malloch). Transf. from Brachycistis, p. 744 (teste Wasbauer, 
1958, loc. cit. supra, p. 142). 

♦Genus BRUESIELLA Mann 
{ = Eurycros Mickel and Krombein, p. 746) 

Bruesiella Mann, 1914. Psyche 21: 182. 

Type: Bruesiella formicaria Mann. By monotypy and orig. 
desig. 

This new synonymy is based on an examination of the type from Hidalgo, 
Mexico, in the Museum of Comparative Zoology. The genus was assigned 
erroneously by Mann to the Bethylidae. 

METHOCHA subgenus METHOCHA Latre'lle, p. 747 

califomica Westwood. Host: probably Cicindela senilis Horn, reared in lab. 
on Omus calif ortiicus Esch. 



326 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Biology: Burdick and Wasbauer, 1959. Wasmann Jour. Biol. 17: 75-88, 
6 figs. 

*impolita Krombein. W. Va., D. C, N. Y. 

Methocha (Methocha) impolita Krombein, 1958. Ent. Soc. Wash. Proc. 
60: 58. 9. 

MYRMOSA subgenus MYRMOSA Lafreille, pp. 747-748 

bradleyi Roberts. 

Biology: Linsley, 1960. Pan-Pacific Ent. 36: 36 (mating behavior). 

Genus MYRMOSULA Bradley, p. 748 

parvula (Fox). Host: Lasioglossum (Dialictus) inconspicuum (Sm.). 

Family SiEROLOMORPHIDAE 

Genus SIEROLOMORPHA Ashmead, pp. 748-749 

Taxonomy: Evans, 1961. Breviora, No. 140, 12 pp., 6 figs. 

ambigua (Ashmead). Syn. of canadensis (Provancher). 

*aparhe Evans. Ariz. 

Sierolomorpha apache Evans, 1961. Breviora, No. 140, p. 5, fig. 6. $. 

*bicolor Evans. Ariz. 

Sierolomorpha bicolor Evans, 1961. Breviora, No. 140, p. 4, fig. 5. 9, $. 

*brevicornis Evans. S. C. (Greenville). 

Sierolomorpha brevicornis Evans, 1961. Breviora, No. 140, p. 8, fig. 3. $. 

canadensis (Provancher). Alta., Ont. and Mass. south to Ariz., Tex. and 
Fla. 
Sierolomorpha ambigua (Ashmead) (syn. by Evans, 1961. Breviora, No. 
140, p. 11). 

*nigrescens Evans. Yukon, Alta. and Sask. south to Calif., Ariz, and Colo. 
Sierolomorpha nigrescens Evans, 1961. Breviora, No. 140, p. 9, fig. 1. 
$,9. 

*8imilis Evans. Conn, south to Ga., W. Va., Kans. 

Sierolomorpha similis Evans, 1961. Breviora, No. 140, p. 6, fig. 4. $, 9. 

Family MUTILLIDAE 

Subfamily APTEROGYNINAE 

Genus CHYPHOTES Blake, pp. 749-751 

^minimus Mickel, Ariz. (Organ Pipe Natl. Mon.). 

Chyphotes minimus Mickel, 1963. Pan-Pacific Ent. 39: 186. $. 

'^nitidus Mickel. Calif. (Winterhaven). 

Chyphotes nitidus Mickel, 1963. Pan-Pacific Ent. 39: 188. 9. 

Subfamily TYPHOCTINAE 

Taxonomy: Krombein and Schuster, 1957. Ent. Soc. Wash. Proc. 59: 209- 
232, 2 pis. 

Genus TYPHOCTES Ashmead, p. 751 

peculiaris mirabilis (Cockerell), changed status. Kans., Colo., Tex., N. Mex., 
Ariz.; Mexico (Chihuahua). 
Typhoctes peculiaris (Cresson) of Catalog, in part (mirabilis (Ckll.) is 
a subsp. of peculiaris, not a syn., teste Krombein and Schuster, 
1957, loc. cit. supra, p. 230, figs. 5, 6). 



MUTILLIDAE 327 

peculiaris pecuHaris (Cresson), changed status. Wash., Idaho, Utah, Ariz., 
Calif. 
Typhoctps peculiaris (Cresson) of Catalog, in part (peculiaris (Cr.) and 
difficilis (Mickel) constitute the subspecies p. peculiaris, teste 
Krombein and Schuster, 1957, loc. cit. supra, p. 227, fig. 3) 

*slrioIatus Krombein and Schuster. Ariz, to western Tex. 

Typhoctes striolatus Krombein and Schuster, 1957. Ent. Soc. Wash. Proc. 
59: 223, fig. 2. 9, $. 

*willianisi Krombein and Schuster. Southern Calif. 

Typhoctes tvilliamsi Krombein and Schuster, 1957. Ent. Soc. Wash. Proc. 
59: 220, figs. 1, 4. 3,9. 

Tribe SPHAEROPTHALMINI 

Taxonomy: Schuster, 1958. Ent. Amer. (n.s.) 37: 1-130, 7 pis. (Part 
II of a tribal revision of males containing keys to genera and 
subgenera and revisions of Acrophotopsis Schuster, Dilophotopsis 
Schuster and AcantJiophotopsis Schuster) . 

Insert after heading subfamily SPHAEROPTHALMINAE, p. 751. 

PHOTOMORPHUS subgenus PHOTOMORPHUS Viereck 
{ = Photomorphus Viereck of Catalog, pp. 751—752 

Reduced to subgeneric rank (teste Schuster, 1952. Brooklyn Ent. Soc. 
Bui. 47: 53). 

banks! (Bradley). Ala. 

johnsoni johnsoni Viereck, changed status. 

Photomorphus johnsoni Viereck. Reduced to subsp. rank (teste Schuster, 
1958, loc. cit. supra, p. 46). 

*johnsoni natehitoches Schuster. La. 

Photomorphus (Photomorphus) johnsoni natehitoches Schuster, 1958. 
Ent. Amer. (n.s.) 37: 46. $. 

PHOTOMORPHUS subgenus PHOTOMORPHINA Schuster 

Photomorphus subg. Photomorphina Schuster, 1958. Ent. Amer. (n.s.) 
37: 8 (in key), 39. Preocc. by Photomorphus subg. Photomorphina 
Schuster, 1952, and a syn. of it. 

Type: Photomorphus (Photomorphina') quadriangulata 
Schuster. Orig. desig. 

Place on p. 752 after Photomorphus subg. Photomorphus Viereck. 

*aurifera Schuster. Ariz. 

Photomorphus (Photomorphina) aurifera Schuster, 1958. Ent. Amer. 
(n.s.) 37: 44. $. Preocc. by and a syn. of Photomorphus 
(Photomorphina) aurifera Schuster, 1952. 

*auriventris Schuster. Type locality not cited. 

Photomorphus (Photomorphina) auriventris Schuster, 1958. Ent. Amer. 
(n.s.) 37: 40. $. 

bicolor (Blake). Transf. from Photopsis Blake, p. 753 (teste Schuster, 1958, 
loc. cit. supra, p. 42). 

*californicus Schuster. Calif. 

Photomorphus (Photomorphina) californica Schuster, 1958. Ent. Amer. 
(n.s.) 37: 44. $. 

clandestinus (Viereck). Transf. from Odontophotopsis Viereck, p. 756 (teste 
Schuster, 1958, loc. cit. supra, p. 45). 

*cobabi Schuster. Ariz.; Mexico. 

Photomorphus (Photomorphina) cobabi Schuster, 1958. Ent. Amer. 
(n.s.) 37: 41. $. 



328 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*cochisae Schuster. Ariz.; Mexico. 

Photomorphus (,Photomorphina) cochisae Schuster, 1958. Ent. Amer. 
(n.s.) 37: 43. $. 

*coloradcIIa Schuster. Colo. 

Photomorphus {Pholoinorphina} coloradella Schuster, 1958. Ent. Amer. 
(n.s.) 37: 44. $. 

*comobabi Schuster. Ariz. ; Mexico. 

Photomorphus {Photomorphina) comobabi Schuster, 1958. Ent. Amer. 
(n.s.) 37: 45. $. 

*dichrous Schuster. Ariz. 

Photomorphus (^Photomorphina) dichrous Schuster, 1958. Ent. Amer. 
(n.s.) 37: 41. $. 

*imperialoides Schuster. Ariz. 

Photomorphus (_Photomorphina) imperialoides Schuster, 1958. Ent. 
Amer. (n.s.) 37: 40. $. 

jason (Fox). Transf. from Photopsis Blake, p. 754 (teste Schuster, 1958, 
loc. cit. supra, p. 41). 

*juanita Schuster. Tex.; Mexico. 

Photomorphus (Photomorphina) juauita Schuster, 1958. Ent. Amer. 
(n.s.) 37: 43. $. Preocc. by and a syn. of Photomorphus 
(Photomorphina) juanita Schuster, 1952. 

*mininius Schuster. Ariz. (Wellton). 

Photomorphus (Photomorphina) minima Schuster, 1958. Ent. Amer. 
(n.s.) 37: 43. $. 

*nanullus Schuster. Ariz. 

Photomorphus (Photomorphina) nanulla Schuster, 1958. Ent. Amer. 
(n.s.) 37: 44. $ 

*obscurus Schuster. Calif. 

Photomorphus iPhotomorphina) obscura Schuster, 1958. Ent. Amer. 
(n.s.) 37: 45. $. 

*quadriangulatus Schuster. Ariz. 

Photomorphus (Photomorphina) quadriangulata Schuster, 1958. Ent. 
Amer. (n.s.) 37: 42. $. 

sarpedon (Fox). Transf. from Odontophotopsis Viereck, p. 757 (teste 
Schuster, 1958, loc. cit. supra, p. 43). 

spinci floridensis (Schuster). Transf. from Odontophotopsis Viereck, p. 757. 

spinci spinci (Bradley). Transf. from Odontophotopsis Viereck, p. 757 (teste 
Schuster, 1958, loc. cit. supra, p. 44). 

''spinciformis Schuster. Tex. 

Photomorphus (Photomorphina) spinciformis Schuster, 1958. Ent. Amer. 
(n.s.) 37: 44. $ 

subtenuis (Viereck). Transf. from Odontophotopsis Viereck, p. 757 (teste 
Schuster, 1958, loc. cit. supra, p. 43). 

*taeniatus parksi Schuster. Tex. 

Photomorphus (Photomorphina') taeniatus parksi Schuster, 1958. Ent. 
Amer. (n.s.) 37: 42. $. 

*taeniatus taeniatus Schuster. Tex. Ariz. 

Photomorphus (Photomorphina) taeniatus taeniatus Schuster, 1958. Ent. 
Amer. (n.s.) 37: 44. $. 

ihamyras (Fox). Transf. from Odontophotopsis Viereck, p. 758 (teste 
Schuster, 1958, loc. cit. supra, p. 43). 

trunculus (Viereck). Transf. from Odontophotopsis Viereck, p. 758 (teste 
Schuster, 1958, loc. cit. supra, p. 41). 



MUTILLIDAE 329 

*vegas Schuster. N. Mex. 

Photomorphus (Photomorphina) vegas Schuster, 1958. Ent. Amer. 
(n.s.) 37: 42. $. 

SPHAEROPTHALMA subgenus MICROMUTILLA Ashmead 
{ = Micromutilla Ashmead of Catalog, in part, p. 752) 

Micromntilla Ashmead is a weakly defined subgenus of Sphaeropthalma 
Blake, not a discrete genus (teste Schuster, 1958, loc. cit. supra, p. 9). 

acontia (Fox). Transf. from Photopsis Blake, p. 753 {teste Schuster, 1958, 
loc. cit. supra, p. 16). 

*arizonae Schuster. Ariz. 

Sphaeropthalma (Microtnutilla) arizonae Schuster, 1958. Ent. Amer. 
(n.s.) 37: 16. $. 

bellei-ophon (Fox). Transf. from Photopsis Blake, p. 753 {teste Schuster, 
1958, loc. cit. supra, p. 15). 

brachyptera (Schuster). Transf. from Photopsis Blake, p. 753 {teste Schuster, 
1958, loc. cit. supra, p. 18). 

*californiensis callforniensis Schuster. Calif. 

Sphaeropthalma {Micromntilla) californiense californiense Schuster, 
1958. Ent. Amer. (n.s.) 37: 18. $. 

*californiensis fuscatella Schuster. Calif. 

Sphaeropthalma {Micromutilla) californiense fuscatella Schuster, 1958. 
Ent. Amer. (n.s.) 37: 18. $ 

difficilis (Baker). Transf. from Photopsis Blake, p. 754 {teste Ferguson, 
1962. Univ. Calif. Pubs. Ent. 27: 82). Host: Auplopus architectus 
metallicus (Bks.). 

hyalina (Blake). Transf. from Photopsis Blake, p. 754 {teste Schuster, 1958, 
loc. cit. supra, p. 19). 

juxta (Blake). Kans. Transf. from Photopsis Blake, p. 754 {teste Schuster, 
1958, loc. cit. supra, p. 14). 

*maricopelIa castanea Schuster. Ariz. (Quijotoa). 

Sphaeropthalma (Micromutilla) maricopella castanea Schuster, 1958. 
Ent. Amer. (n.s.) 37: 17. $. 

*inaricopella maricopella Schuster. Calif. 

Sphaeropthalma {Micromutilla') maricopella maricopella Schuster, 1958. 
Ent. Amer. (n.s.) 37: 17. $. 

mesillensis (Cockerel]). Transf. from Photopsis Blake, p. 755 {teste Schuster, 
1958, loc. cit. supra, p. 17). 

pallida (Blake). Transf. from Photopsis Blake, p. 755 {teste Schuster, 1958, 
loc. cit. supra, p. 16). 

*pateli Schuster. Ariz. (Hereford). 

Sphaeropthalma (Micromutilla) pateli Schuster, 1958. Ent. Amer. 
(n.s.) 37: 15. $. 

*quijotoa quijotoa Schuster. Ariz. Another subspecies, quijotoa parrasia 
Schuster, occurs in Baja California. 
Sphaeropthalma (Micromutilla) quijotoa quijotoa Schuster, 1958. Ent. 
Amer. (n.s.) 37: 18. 5- 

*reducta Schuster. Ariz. 

Sphaeropthalma (Micromutilla) reducta Schuster, 1958. Ent. Amer. 
(n.s.) 37: 18. $. 

*sabino Schuster. Ariz. 

Sphaeropthalma (Micromutilla) sabino Schuster, 1958. Ent. Amer. 
(n.s.) 37: 19. $. 

*sonora Schuster. Ariz. (Tucson). 

Sphaeropthalma (Micromutilla) sonora Schuster, 1958. Ent. Amer. 
(n.s.) 37: 16. $. 



330 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*'sublobata Schuster. Idaho. 

Sphaeropthalma {Micromulilla) sublobata Schuster, 1958. Ent. Amer. 
(n.s.) 37: 16. $. 

*uvaldella Schuster. Tex. 

Sphaeropthalma (Micromutilla') uvaldella Schuster, 1958. Ent. Amer. 
(n.s.) 37: 19. $. 

*yavapai Schuster. Ariz. 

Sphaeropthalma (.Micromutilla) yavapai Schuster, 1958. Ent. Amer. 
(n.s.) 37: 19. $. 

*yuinaella Schuster. Ariz. 

Sphaeropthalma (Micromutilla) yumaella Schuster, 1958. Ent. Amer. 
(n.s.) 37: 19. $. 

♦SPHAEROPTHALMA subgenus PHYSETAPSIS Schuster 

Sphaeropthalma subg. Physetapsis Schuster, 1958. Ent. Amer. (n.s.) 37: 
9 (in key), 20. 

Type: Sphaeropthalma (Physetapsis) papaga Schuster. Orig. 
desig. 

Insert on p. 752 after Micromutilla Ashmead. 

*borealis Schuster. Southwestern Canada. 

Sphaeropthalma (Physetapsis) borealis Schuster, 1958. Ent. Amer. (n.S.) 
37: 20. $. 

*ecarinala Schuster. Calif. 

Sphaeropthalma (Physetapsis) ecarinata Schuster, 1958. Ent. Amer. 
(n.s.) 37: 20. $. 

*papaga ephysetos Schuster. Southwestern Ariz., southern Calif. 

Sphaeropthalma (Physetapsis) papaga ephysetos Schuster, 1958. Ent. 
Amer. (n.s.) 37: 20. $. 

*papaga papaga Schuster. Southeastern Ariz. 

Sphaeropthalma (Phvsetapsis) papaga papaga Schuster, 1958. Ent. Amer. 
(n.s.) 37: 20,'pl. 6, fig. 3. $. 

*subcarinata Schuster. N. Hex. 

Sphaeropthalma (Physetapsis) suhcarinata Schuster, 1958. Ent. Amer. 
(n.s.) 37: 20. 5. 

SPHAEROPTHALMA subgenus SPHAEROPTHALMA Blake 
( = Sphoeropfha/fno Blake of Catalog, p. 752) 

Reduced to subgeneric rank {teste Schuster, 1958, loc. cit supra, p. 10). 

albiplumosa Schuster. Reared from nest of solitary vespid in old clay cell 
of Scelipliron. 

*aiiripilis fasciventris Schuster. Type locality not cited. 

Sphaeropthalma (Sphaeropthalma) anripilis fasciventris Schuster, 1958. 
Ent. Amer. (n.s.) 37: 39. $. 

pennsylvanica pennsylvanica (Lepeletier). Host: Trypargilum clavatum 
(Say) prepupa in cocoon in rotten stump. 

SPHAEROPTHALMA subgenus PHOTOPSIS Blake 
izzPhotopsis Blake of Catalog, in part, pp. 753-756) 

Reduced to subgeneric rank {teste Schuster, 1958, loc. cit. supra, p. 10). 

Taxonomy and Biology: Ferguson, 1962. Univ. Calif. Pubs. Ent. 27: 
1-92, 7 pis., 2 text figs. 

abdominalis (Blake). Transf. to Sphaeropthalma subg. Photopsioides 

Schuster. 

abstrusa (Baker). Transf. to Sphaeropthalma subg. Photopsioides Schuster 
as a subsp. of amphion (Fox). 



MUTILLIDAE 331 

acontia (Fox). Transf. to Sphaeropthalma subg. Micromutilla Ashmead, p. 

752. 

albicincta albicincta (Fox). N. Mex. and northern Ariz, to Utah and Oreg. 
Photopsis albicincta (Fox). Reduced to subsp. rank (teste Schuster, 1958, 
loc. cit. supra, p. 36). 

♦albicincta flavipes Schuster. Southern Ariz. 

Sphaeropthalma (Photopsis) albicincta flavipes Schuster, 1958. Ent. 
Amer. (n.s.) 37: 36. $. 

♦albicincta piceoceps Schuster. Tex. 

Sphaeropthalma (Photopsis) albicincta piceoceps Schuster, 1958. Ent. 
Amer. (n.s.) 37: 35. $. 

albopilosa (Blake). Insert (Sphaeropthalma) after Mutilla in orig. cit. 

amphion (Fox). Transf. to Sphaeropthalma subg. Photopsioides Schuster. 

*angulifera Schuster. Type locality not cited. 

Sphaeropthalma (Photopsis) angulifera Schuster, 1958. Ent. Amer. 
(n.s.) 37: 32. $. 

anthophora Ashmead. Change spelling in specific heading and in orig cit. 
from anthophorae to anthophora. 

*arenicola Schuster. Type locality not cited. 

Sphaeropthalma (Photopsis) arenicola Schuster, 1958, Ent. Amer. (n.s.) 
37: 25. $. 

*baboquivari Schuster. Ariz. 

Sphaeropthalma (Photopsis) baboquivari Schuster, 1958. Ent. Amer. 
(n.s.) 37: 24, pi. 4, figs. 1-6. $. 

bellerophon (Fox). Transf. to Sphaeropthalma subg. Micromutilla Ashmead, 
p. 752. 

bicolor (Blake). Transf. to Photomorphus subg. Photomorphina Schuster. 

*bisetosa Schuster. N. Mex. 

Sphaeropthalma (Photopsis) bisetosa Schuster, 1958. Ent. Amer. (n.s.) 
37: 36. $. 

blakeii (Fox). Host: Diadasia vallicola Timb. 

Biology: Ferguson, 1962, loc. cit. supra, pp. 10-15, pis. 2, 3, 4, 5 and 7. 

brachyptera Schuster. Transf. to Sphaeropthalma subg. Micromutilla 

Ashmead, p. 752. 

ceyx (Fox). Syn. of blakeii (Fox). 

♦ceyxoides Schuster. Ariz. 

Sphaeropthalma (Photopsis) ceyxoides Schuster, 1958. Ent. Amer. (n.s.) 
37: 25. $. 

clara clara (Cresson), changed status. Colo, to Wyo. and Idaho. This subsp. 
is transitional to Sphaeropthalma subg. Physetapsis Schuster. 
Photopsis clara (Cresson). Reduced to subsp. rank {teste Schuster, 1958, 
loc. cit. supra, p. 27). 

*clara cleomella Schuster. Mont. This subsp. is transitional to Sphaer- 
opthalma subg. Physetapsis Schuster. 
Sphaeropthalma (Photopsis) clara cleomella Schuster, 1958. Ent. Amer. 
(n.s.) 37: 27. $. 

contractu (Blake). Transf. to Sphaeropthalma subg. Photopsioides Schuster. 

danaus (Blake). Kans., Tex.; Mexico. Transf. from Odontophotopsis 

Viereck, p. 756 (teste Schuster, 1958, loc. cit. supra, p. 24). 

*dentifera Schuster. S. Dak., Minn. 

Sphaeropthalma (Photopsis) dentifera Schuster, 1958. Ent. Amer. (n.s.) 

37: 24. $. 

difficilis Baker. Transfer to Sphaeropthalma subg. Micromutilla Ashmead. 



332 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

edwardsii edwardsii (Cresson), changed status. Oreg., Idaho. 

Photopsis edwardsii (Cresson). Reduced to subsp. rank {teste Schuster, 
1958, loc. cit. supra, p. 36). 

*edwardsii flammifera Schuster. Calif. 

Sphaeropthaltna (Photopsis) edwardsii flammifera Schuster, 1958. Ent. 
Amer. (n.s.) 37: 36. $. 

*facilis facilis (Cameron). Ariz.; Mexico. 

Sphaeropthaltna (!) facilis Cameron, 1896. Biol. Cent.-Amer., Hym. 2, 

p. 392. $. Reduced to subsp. rank {teste Schuster, 1958, loc. cit. 

supra, p. 26). 
Photopsis hubbardii (Fox). Syn. by Schuster, 1958, loc. cit. supra, p. 26. 

*facilis rubriventris Schuster. Tex. 

Sphaeropthaltna (Photopsis) facilis rubriventris Schuster, 1958. Ent. 
Amer. (n.s.) 37: 26. $. 

*ferrug:inopsis Schuster. Ariz., Calif. 

Sphaeropthaltna (Photopsis) ferruginopsis Schuster, 1958. Ent. Amer. 
(n.s.) 37: 29. $. 

*fuscipes Schuster. Ariz. This sp. is transitional to Sphaeropthalma subg. 
Physelapsis Schuster. 
Sphaeropthaltna (Photopsis) fuscipes Schuster, 1958. Ent. Amer. (n.s.) 
37: 21. $. 

*heIicaon diegrueno Schuster. Ariz. 

Sphaeropthaltna (Photopsis) helicaon diegueno Schuster, 1958. Ent. 
Amer. (n.s.) 37: 35. $. 

helicaon helicaon (Fox), changed status. 

Photopsis helicaon (Fox). Reduced to subsp. rank {teste Schuster, 1958, 
loc. cit. supra, p. 35). 

helicaon lingulatiis (Viereck), changed status. 

Photopsis lingulata Viereck. Reduced to subsp. rank {teste Schuster, 
1958, loc. cit. supra, p. 26). 

hubbardii (Fox). Syn. of facilis facilis (Cameron). 

hyalina (Blake). Transf. to Sphaeropthalma subg. Micromutilla Ashmead, 
p. 752. 

imperialiformis imperialiformis (Viereck), changed status. Tex. to Kans. 
and Colo. 
Photopsis iniperinlifortnis (Viereck). Reduced to subsp. rank {teste 
Schuster, 1958, loc. cit. supra, p. 34). 

*imperialiformis maricopae Schuster. Ariz, to Wash. 

Sphaeropthaltna (Photopsis) itnperialiforniis maricopae Schuster, 1958. 
Ent. Amer. (n.s.) 37: 34. $. 

indigens (Baker). Calif. 

ingenua (Baker). Syn. of orestes (Fox). 

jason (Fox). Transf. to Photomorphus subg. Photomorphina Schuster. 

juxta (Blake). Transf. to Sphaeropthalma subg. Micromutilla Ashmead, p. 
752. 

lingulata (Viereck). A subsp. of helicaon (Fox). 

*lui8eno Schuster. Calif. 

Sphaeropthalma (Photopsis) luiseno Schuster, 1958. Ent. Amer. (n.s.) 
37: 23. $. 

*megagmathos auriferus Schuster. Calif. 

Sphaeropthalma (Photopsis) itxegagnathos auriferus Schuster, 1958. Ent. 
Amer. (n.s.) 37: 36. $. 

""megaimathos megagnathos Schuster. Calif. 

Sphaeropthalma (Photopsis) megagnathos megagnathos Schuster, 1958. 
Ent. Amer. (n.s.) 37: 36. $ 



MUTILLIDAE 333 

melanderi (Baker). Transf. to Sphaeropthalma suhg. Photopsioides Schuster 
as a subsp. of uro (Blake). 

mesillensis Cockerell. Transf. to Sphaeropthalma subg. Micromutilla 

Ashmead, p. 752. 

*militaris Schuster. Ariz., Calif. 

Sphaeropthalma (Photopsis) militaris Schuster, 1958. Ent. Amer (n.S.) 
37: 27, pi. 5, fig. 2. $. 

*neoinoxicana Schuster. N. Mex. 

Sphaeropthalma (Photopsis) neomexicana Schuster, 1958. Ent. Amer. 
(n.S.) 37: 30. $. 

*noctivag;a infuscata Schuster. Ariz. 

Sphaeropthalma (Photopsis) noctivaga infuscata Schuster, 1958. Ent. 
Amer. (n.s.) 37: 32. $. 

noctivaga noctivaga (Melander), changed status. Tex. to Kans. 

Photopsis noctivaga (Melander). Reduced to subsp. rank {teste Schuster, 
1958, loc. cit. supra, p. 32). 

nokonii.«i nokoniis (Blake), changed status. Ariz. 

Photopsis nokomis (Blake). Reduced to subsp. rank {teste Schuster, 
1958, loc. cit. supra, p. 35). 

*nokoinis rufireps Schuster. Ariz. 

Sphaeropthalma (Photopsis) nokomis ruficeps Schuster, 1958. Ent. 
Amer. (n.s.) 37: 35. $. 

nudata (Baker). Transf. to Sphaeropthalma subg. Photopsioides Schuster 
as a syn. of amphion amphion (Fox). 

*ordae Schuster. Tex. 

Sphaeropthalma (Photopsis) ordae Schuster, 1958. Ent. Amer. (n.s.) 
37: 23. $. 

orestes (Fox). Wash, to Baja California. Hosts: Anthidium c. collectum 

Huard, Euodynerus hoscii auranum (Cam.), Tachysphex tenui- 

punctus Fox. 
Photopsis pattersonae Melander (syn. by Schuster, 1958, loc. cit. supra, 

p. 30). 
Photopsis uniformis Baker (syn. by Schuster, 1958, loc. cit. supra, p. 30). 
Photopsis pedata Baker (syn. by Schuster, 1958, loc. cit. supra, p. 30). 
Photopsis ingenua Baker (syn. by Schuster, 1958, loc. cit. supra, p. 30). 
Photopsis salmani Mickel (syn. by Ferguson, 1962, loc. cit. S7ipra, p. 10). 
?* Sphaeropthalma (Photopsis) salmani fresnoensis Schuster, 1958. Ent. 

Amer. (n.s.) 37: 30. $. Presumed a syn. because Ferguson placed 

typical salmani as a syn, of orestes. 
?* Sphaeropthalma (Photopsis) salmani oregana Schuster, 1958. Ent. 

Amer. (n.s.) 37: 31. $. Presumed a syn. because Ferguson placed 

typical salmani as a syn. of orestes. 
Biology: Ferguson, 19G2, loc. cit. supra, pp. 6-10, pis. 1, 6, tab. 1, 

palamedes (Fox), Syn. of pluto (Fox). 

pallida (Blake). Transf. to Sphaeropthalma subg. Micromutilla Ashmead, 
p. 752, 

*pallidipes gila Schuster. Type locality not cited — Ariz. (?). 

Sphaeropthalma (Photopsis) pallidipes gila Schuster, 1958. Ent. Amer. 
(n.s.) 37: 28. $. 

*pallidipes pallidipes Schuster. Type locality not cited. 

Sphaeropthalma (Photopsis) pallidipes pallidipes Schuster, 1958. Ent. 

Amer. (n.s.) 37: 27. $. 

*parkeri Schuster. Ariz. 

Sphaeropthalma (Photopsis) parkcri Schuster, 1958. Ent. Amer, (n.s.) 
37: 28. $. 

pattersonae (Melander). Syn. of orestes (Fox). 



334 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

pedata (Baker). Syn. of orestes (Fox). 

*piceogaster Schuster. Type locality not cited. 

Sphaeropthalma (Photopsis) piceogaster Schuster, 1958. Ent. Amer. 
(n.s.) 37: 27. $. 

*pinalea pinalea Schuster. Ariz. 

Sphaeropthalma (Photopsis) pinalea pinalea Schuster, 1958. Ent. Amer. 
(n.s.) 37: 32. $. 

'''pinalea texanella Schuster. Tex. to N. Dak. 

Sphaeropthalma (Photopsis) pinalea texanella Schuster, 1957. Ent. 
Amer. (n.s.) 37: 32. $. 

*pinales Schuster. Ariz. 

Sphaeropthalma (Photopsis) pinales Schuster, 1958. Ent. Amer. (n.s.) 
37: 36. $. 

pluto (Fox). 

Photopsis palamedes (Fox) (syn. by Schuster, 1958, loe. cit. supra, p. 31). 

*rustica occUaria Schuster. Calif. 

Sphaeropthalma (Photopsis) rustica ocellaria Schuster, 1958. Ent. Amer. 
(n.s.) 37: 32. $. 

rustica rustica (Blake), changed status. 

Photopsis rustica (Blake). Reduced to subsp. rank (teste Schuster, 1958, 
loc. cit. supra, p. 32). 

salmani (Mickel). Syn. of orestes (Fox). 

*salmani fresnoensis Schuster. Syn. (?) of orestes (Fox). 

*salmani oregana Schuster. Syn. (?) of orestes (Fox). 

*8cudderi Schuster. Ariz. 

Sphaeropthalma (Photopsis) scudderi Schuster, 1958. Ent. Amer. (n.s.) 
37; 23. $. 

*similis Schuster. Type locality not cited. 

Sphaeropthalma (Photopsis) similis Schuster, 1958. Ent. Amer. (n.s.) 
37: 30. $. 

*spinifera Schuster. Kans. 

Sphaeropthalma (Photopsis) spinifera Schuster, 1958. Ent. Amer. (n.S.) 
37: 23. $. 

*tetracuspis Schuster. Type locality not cited. 

Sphaeropthalma (Photopsis) tetracuspis Schuster, 1958. Ent. Amer. 
(n.s.) 37: 31. $. 

*tuberculifera Schuster. Type locality not cited. 

Sphaeropthalma (Photopsis) tuherculifera Schuster, 1958. Ent. Amer. 
(n.s.) 37: 25. $. 

unicolor (Cresson). U. S. west of Rocky Mts.; Mexico (Baja California 
del Norte). Hosts: Anthidium c. collectum Huard, Callanthidium 
illustre (Cr.), Ashmeadiella c. calif omica (Ashm.),_ Anthophora 
abruptella Ckll., A. bomboides neomexicana Ckll., A. linsleyi Timb., 
?A. montana Cr., A. occidentalis Cr., Diadasia bituberculata (Cr.), 
Melissodes robustior Ckll., Xeromelecta calif omica (Ashm.). 
Biology: Ferguson, 1962, loc. cit. supra, pp. 15-22, pis. 3, 5, and 7. 

uniformis (Baker). Syn. of orestes (Fox). 

itro (Blake). Transf. to Sphaeropthalma subg. Photopsioides Schuster. 

♦SPHAEROPTHALMA subgenus PHOTOPSIOIDES Schuster 

Sphaeropthalma subg. Photopsioides Schuster, 1958. Ent. Amer. (n.s.) 
37: 10 (in key), 36. 

Type: Agama uro Blake. Orig desig. 

Place after Sphaeropthalma subg. Photopsis Blake, p. 756. 



MUTILLIDAE 335 

abdominalis (Blake). Transf. from Photopsis Blake, p, 753 (teste Schuster, 
1958. Ent. Amer. (n.s.) 37: 37). 

amphion abstrusa (Baker). Transf. from Photopsis Blake, p. 753 and re- 
duced to subsp. rank (teste Schuster, 1958, loc. cit. supra, p. 38). 

amphion amphion (Fox), changed status. Transf. from Photopsis Blake, p. 
753 and reduced to subsp. rank (teste Schuster, 1958, loc. cit. 
supra, p. 38). 
Photopsis nudata Baker (syn. by Schuster, 1958, loc. cit. supra, p. 38). 

contracta (Blake). Transf. from Photopsis Blake, p. 754 (teste Schuster, 
1958, loc. cit. supra, p. 37). 

uro melanderi (Baker), changed status. Transf. from Photopsis Blake, p. 
755 and reduced to subsp. rank (teste Schuster, 1958, loc. cit. 
supra, p. 38). 

*uro stenognatha Schuster. N. Mex., Ariz. 

Sphaeropthalma (Photopsioides) uro stenognatha Schuster, 1958. Ent. 
Amer. (n.s.) 37: 38. $. 

uro uro (Blake), changed status. Transf. from Photopsis Blake, p. 756 and 
reduced to subsp. rank (teste Schuster, 1958, loc. cit. supra, p. 38). 
Hosts : Dianthidium curvatum sayi Ckll., Trypargilum tridentatum 
(Pack.). 

ODONTOPHOTOPSIS subgenus ODONTOPHOTOPSIS Viereck 
i^Odontophotopsis Viereck of Catalog, pp. 756—758) 

Tetraphotopsis Ashmead, p. 758. Syn. by Schuster, 1958, loc. cit. supra, 
p. 54. 

Reduced to subgeneric rank (teste Schuster, 1958, loc. cit. supra, pp. 8 
(in key), 46). 

acmaea Viereck. A subsp. of adonis (Fox). 

adonis acmaea Viereck, changed status. 

Odontophotopsis acmaea Viereck. Reduced to subsp. rank (teste Schuster, 
1958, loc. cit. supra, p. 54). 

adonis adonis (Fox), changed status. 

Odontophotopsis adonis (Fox). Reduced to subsp. rank (teste Schuster, 
1958, loc. cit. supra, p. 54). 

alamonis Viereck. Transf. to Dilophotopsis Schuster as a syn. of concolor 
concolor (Cresson). 

alemon (Fox). 

Odontophotopsis trita (Melander) (syn. by Schuster, 1958, loc. cit. supra, 

p. 56). 
Odontophotopsis crucis Viereck (syn. by Schuster, 1958, loc. cit. supra, p. 

56). 

""anomala Schuster. Tex. 

Odontophotopsis (Odontophotopsis) anomala Schuster, 1958. Ent. Amer. 
(n.s.) 37: 60. $. 

*apacheorum apacheorum Schuster. Tex. 

Odontophotopsis (Odontophotopsis) appacheoruni (!) appacheoruni (!) 

Schuster, 1958. Ent. Amer. (n.s.) 37: 55. $. 

*apacheorum rostrata Schuster. Ariz. 

Odontophotopsis (Odontophotopsis) appacheoruni (!) rostratus Schuster, 
1958. Ent. Amer. (n.s.) 37: 55. 5- 

*armata Schuster. Type locality not cited. 

Odontophotopis (Odontophotopsis) armata Schuster, 1958. Ent. Amer. 
(n.s.) 37: 60. $. 

atripes Mickel. Syn. of pudica (Melander). 

aulus (Blake). A subsp. of tapajos (Blake). 



336 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*bequaertiJ Schuster. Tex. 

Odontophotopsis (Odontophotopsis) bequaertii Schuster, 1958. Ent. 
Amer. (n.s.) 37: 52. 5. 

*brarcata Schuster. Type locality not cited. 

Odontophotopsis (Odontophotopsis) braccatus Schuster, Ent. Amer. 
(n.s.) 37: 54. S. 

brevicornis (Fox). Syn. of melioausa (Blake). 

clandestina Viereck. Transf. to Photoiuorphus subg. Photomorphina Schuster. 

*clypeata Schuster. Ariz. 

Odontophotopsis (Odontophotopsis) clypeatus Schuster, 1958. Ent. 
Amer. (n.s.) 37: 59. $. 

concolor (Cresson). Transf. to Dilophotopsis Schuster. 

*conifera Schuster. Ariz. 

Odontophotopsis (Odontophotopsis) coniferus Schuster, 1958. Ent. 
Amer. (n.s.) 37: 48. $. 

cookii Baker. 

Biology: Ferguson, 1963. Pan-Pacific Ent. 39: 65-66. 

crucis Viereck. Syn. of alenion (Fox). 

danaus (Blake). Transf. to Sphaeroplhalma subg. Photopsis Blake, p. 754. 

*edentaia distans Schuster. Type locality not cited. 

Odontophotopsis (Odontophotopsis) edentata distans Schuster, 1958. 
Ent. Amer. (n.s.) 37: 53. $. 

*edentata edentata Schuster. Type locality not cited. 

Odontophotopsis (Odontophotopsis) edentata edentata Schuster, 1958. 
Ent. Amer. (n.s.) 37: 53. $. 

fallax Viereck. 

Odontophotopsis ocellata Baker, n. syn. Schuster, 1958, loc. cit. supra, 
p. 57, placed fallax as a syn. of ocellata, but the former antedates 
the latter. 

*grandiceps Schuster. Type locality not cited. 

Odontophotopsis (Odontophotopsis) grandiceps Schuster, 1958. Ent. 
Amer. (n.s.) 37: 53. $. 

*hexadonta Schuster. Type locality not cited. 

Odontophotopsis (Odontophotopsis) hexadontus Schuster, 1958. Ent. 
Amer. (n.s.) 37: 51. $. 

hubbardi Ashmead. Syn. of venusta (Blake). 

*inconspicua brunnea Schuster. Type locality not cited. 

Odontophotopsis (Odontophotopsis) inconspicua brunnea Schuster, 1958. 
Ent. Amer. (n.s.) 37: 54. $. 

inconspicua inconspicua (Blake), changed status. 

Odontophotopsis inconspicua (Blake). Reduced to subsp. rank {teste 
Schuster, 1958, loc. cit. supra, p. 54). 

indotata Viereck. Syn. of melicausa westcottii (Melander). 

*lamellifera Schuster. Ariz. 

Odontophotopsis (Odontophotopsis) laniellifera Schuster, 1958. Ent. 
Amer. (n.s.) 37: 56. $. 

melicausa melicausa (Blake), changed status. 

Odontophotopsis melicausa Blake. Reduced to subsp. rank (teste 

Schuster, 1958, loc. cit. supra, p. 58). 
Odontophotopsis brevicornis (Fox) (syn. by Schuster, 1958, loc. cit. 

supra, p. 58). 
Odontophotopsis mellicornis Baker (syn. by Schuster, 1958, loc. cit. 

supra, p. 58). 
Odontophotopsis obliqua Viereck (syn. by Schuster, 1958, loc. cit. supra, 

p. 58). 



MUTILLIDAE 337 

*inelicausa piceipes Schuster. Mont. 

Odontophotopsis (Odontophotopsis) mellicausa (!) piceipes Schuster, 
1958. Ent. Amer. (n.s.) 37: 58. $. 

melieausa westcottii (Melander), changed status. 

Odontophotopsis westcottii (Melander). Reduced to subsp. rank {teste 

Schuster, 1958, loc. cit. supra, p. 59). 
Odontophotopsis indotata Viereck (syn. by Schuster, 1958, loc. cit. supra, 

p. 59). 

mellicornis Baker. Syn. of melieausa (Blake). 

ohliqua Viereck. Syn. of melieausa (Blake). 

ocellata Baker. Syn. of fallax Viereck. 

*parksiana Schuster. Tex. 

Odontophotopsis (Odontophotopsis) parksiana Schuster, 1958. Ent. 
Amer. (n.s.) 37: 57. $. 

*parva Schuster. Type locality not cited. 

Odontophotopsis (Odontophotopsis) parva Schuster, 1958. Ent. Amer. 
(n.s.) 37: 55. $. 

pudiea (Melander). 

Odontophotopsis atripes Mickel (syn. by Schuster, 1958, loc. cit. supra, 
p. 57). 

♦quadridentata Schuster. Type locality not cited. 

Odontophotopsis (Odontophotopsis) quadridentata Schuster, 1958. Ent. 
Amer. (n.s.) 37: 51. $. 

sarpedon (Fox). Transf. to Photomorphus subg. Photomorphina Schuster. 

spinci floridensis (Schuster). Transf. to Photomorphus subg. Photomorphina 

Schuster. 

spinci spinci (Bradley). Transf. to Photomorphus subg. Photomorphina 

Schuster. 

suhtenuis Viereck. Transf. to Photomorphus subg. Photomorphina Schuster. 

tapaios aulus (Blake), changed status. 

Odontophotopsis aulas (Blake). Reduced to subsp. rank {teste Schuster, 
1958, loc. cit. supra, p. 53). 

tapajos tapajos (Blake), changed status. 

Odontophotopsis tapajos (Blake). Reduced to subsp. rank {teste Schuster, 
1958, loc. cit. supra, p. 53). 

*territa obscura Schuster. Ariz. 

Odontophotopsis (Odontophotopsis) territa obscura Schuster, 1958. Ent. 
Amer. (n.s.) 37: 59. $. 

territa territa (Cockerell), changed status. 

Odontophotopsis territa (Cockerell). Reduced to subsp. rank {teste 
Schuster, 1958, loc. cit. supra, p. 59). 

thamyras (Fox). Transf. to Photomorphus subg. Photomorphina Schuster. 

trita (Melander). Syn. of alemon (Fox). 

truncula Viereck. Transf. to Photomorphus subg. Photomorphina Schuster. 

*unicornis Schuster. Ariz. 

Odontophotopsis (Odontophotopsis) unicornis Schuster, 1958. Ent. Amer. 
(n.s.) 37: 52. $. 

venusta (Blake). 

Tetraphotopsis hubbardi Ashmead (syn. by Schuster, 1958, loc. cit. 
supra, p. 54). 

tvestcottii (Melander). A subsp. of melieausa (Blake). 



338 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*ODONTOPHOTOPSIS subgenus PERIPHOTOPSIS Schuster 

Odontophotopsis subg. Periphotopsis Schuster, 1958. Ent. Amer. (n.s.) 
37: 8 (in key), 60. 

Type: Odontophotopsis (Periphotopsis) mamatus Schuster. 
By monotypy and orig. desig. 

Place after Odontophotopsis Viereck, p. 758. 

^mamata Schuster. Ariz. 

Odontophotopsis (Periphotopsis) mamatus Schuster, 1958. Ent. Amer. 
(n.s.) 37: 60. $. 

Genus Tefraphotopih Ashmead, p. 758 
A synonym of Odontophotopsis subg. Odontophotopsis Viereck. 

*Genus ACROPHOTOPSIS Schuster 

Acrophotopsis Schuster, 1958. Ent. Amer. (n.s.) 37: 4 (in key), 61. 
Type: Acrophotopsis eurygnathus Schuster. Orig. desig. 

Insert on p. 758 after Morsyma Fox. 

*eurygnathus Schuster. Southern Ariz. 

Acrophotopsis eurygnathus Schuster, 1958. Ent. Amer. (n.s.) 37: 10 (in 
key), 65, pi. 1, figs. 1, 2. $. 

♦Genus DILOPHOTOPSIS Schuster 

Dilophotopsis Schuster, 1958. Ent. Amer. (n.s.) 37: 5 (in key), 71. 
Type: Mutilla concolor Cresson. Orig. desig. 

Insert on p. 758 after Acorphotopsis Schuster. 

concolor concolor (Cresson), changed status. Kans. to Colo, and Wyo., Tex., 

N. Mex., eastern Ariz.; Mexico (Zacatecas, Durango, Jalisco). 
Odontophotopsis concolor (Cresson), Transf. from Odontophotopsis 

Viereck, p. 756 and reduced to subsp. rank (teste Schuster, 1958, 

loc. cit. supra, p. 84). 
Odontophotopsis alamonis Viereck. Syn. by Schuster, 1958, loc. cit. supra, 

p. 84. 

*concolor laredo Schuster. Tex. (Winterhaven, Laredo, Cotulla) . 

Dilophotopsis concolor laredo Schuster, 1958. Ent. Amer. (n.s.) 37: 84 
(in key), 86. $. 

*concolor sonorensis Schuster. Southern Ariz, and Calif.; Mexico (Baja 
California). 
Dilophotopsis concolor sonorensis Schuster, 1958. Ent. Amer. (n.s.) 37: 
84 (in key), 88. $. 

*concolor utahensis Schuster. Mont., Utah, western Colo., Nev., northeastern 
Calif. 
Dilophotopsis concolor utahensis Schuster, 1958. Ent. Amer. (n.s.) 37: 
84 (in key), 87. $. 

*stenognatha Schuster. Ariz. (Tucson). 

Dilophotopsis stenognatha Schuster, 1958. Ent. Amer. (n.s.) 37: 11 (in 

key), 74. $. 
Taxonomy: Mickel, 1963. Pan-Pacific Ent. 39: 184 (5). 

♦Genus ACANTHOPHOTOPSIS Schuster 

Acanthophotopsis Schuster, 1958. Ent. Amer. (n.s.) 37: 5 (in key), 88. 
Type: Acanthophotopsis falciformis Schuster. Orig desig. 

Insert on p. 758 after Dilophotopsis Schuster. 

*bequaertii Schuster. Ariz. (Hereford). 

Acanthophotopsis bequaertii Schuster, 1958. Ent. Amer. (n.s.) 37: 12 
(in key), 101. $. 



MUTILLIDAE 339 

*bifurca Schuster. Tex. (Val Verde Co., Winterhaven). 

Acanthophotopsis bifurca Schuster, 1958. Ent. Amer. (n.s.) 37: 13 (in 
key), 98, pi. 3, fig. 3. $. 

*dorophora Schuster. Ariz. (Tucson, Yuma). 

Acanthophotopsis dorophora Schuster, 1958. Ent. Amer. (n.s.) 37: 13 
(in key), 104, pi. 2, fig. 1, pi. 3, fig. 2. 5. 

*evansii Schuster. Tex. (Alpine and Chisos Mts.) ; Mexico (Durango). 

Acanthophotopsis evansii Schuster, 1958. Ent. Amer. (n.s.) 37: 12 (in 
key), 93, pi. 3, fig. 1. S. 

*falciformis falciformis Schuster. Calif. (Palm Springs); Mexico 
(Zacatecas). 
Acanthophotopsis falciformis falciformis Schuster, 1958. Ent. Amer. 
(n.s.) 37: 13 (in key), 108, pi. 2, figs. 2, 3. $. 

*falciformis furcislerna Schuster. Ariz. (Tucson, Phoenix, Ajo). 

Acanthophotopsis falciformis furcisterna Schuster, 1958. Ent. Amer. 
(n.s.) 37: 14 (in key). 111. $. 

Genus PSEUDOMETHOCA Ashmead, pp. 758-761 

contiunax (Cresson). Mexico. 

*Mutilla microphthalma Gerstaecker, 1874. Arch. Naturges. 40: 64. $. 
Taxonomy: Mickel, 1935. Amer. Ent. Soc. Trans. 61: 393. 

frigida frigida (Smith). Probable hosts: Lasioglossmn (Dialictus) incon- 
spicmim (Sm.), L. {D.) zephyrum (Sm.),L. {D.) rohweri (Ellis), 
L. {D.) versatum (Robt.) and Augochlorella striata (Prov.). 
Biology: Lin, N., 1964. Ins. Sociaux 11: 187-192. 

Genus DASYMUTILLA Ashmead, pp. 761-770 

asteria Mickel. Host: Microbembex nigrifrons (Prov.) (teste J. D. Haddock, 
in. litt.). 

biguttata (Cockerell). Ark. 

creon (Blake). Ark. 

dngesii (Cockerell and Casad). Host: Anthophora occidentalis Cr. 

gloriosa (Saussure). Host: Microbembex nigrifrons (Prov.) (teste J. D. 
Haddock, in litt.). 

leda (Blake). N. Mex. 

macilenta (Blake). Delete Ala. from distribution. 

nigripes (Fabricius). B. C. Host: Cerceris fiavofasciata H. S. Smith. 
Biology: Krombein, 1959. Ent. Soc. Wash. Proc. 61: 198. 

pyrrhus (Fox). Host: Bembicine sp. 

Biology: Krombein, 1964. Amer. Mus. Novitates 2201: 2-4, fig. 1. 

vesta vesta (Cresson). B. C. 

Genus LOMACHAETA Mickel, p. 770 

variegata Mickel. Hosts: Solierella plenoculoides similis (Brid.), S. blaisdelli 
(Brid.), nesting in stems of Eriogonum elatum. 
Biology: Parker, 1962. Pan-Pacific Ent. 38: 116. 

*Genus SMICROMUTILLA Mickel 

Smicromutilla Mickel, 1964. Pan-Pacific Ent. 40: 108. 

Type: Smicromutilla potvelli Mickel. Orig. desig. and by 
monotypy. 

Insert on p. 770 after Lomachaeta Mickel. 

♦powelli Mickel. Calif. (San Luis Obispo Co.). Host: ? Xylocelia occidentalis 
Fox. 
Smicromutilla potvelli Mickel 1964. Pan-Pacific Ent. 40; 108, 1 fig. 5, ?. 



340 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

TIMULLA subgenus TIMULLA Ashmead, pp. 770-772 

grotei (Blake). Mexico (Jalisco, Guanajuato). 

*Mutilla fulviventris Gerstaecker, 1874. Archiv. Naturges. 40: 323. $. 
Taxonomy: Mickel, 1937. Ent. Amer. 17: 78. 

oajaca (Blake). Mexico (Morelos, Jalisco, Colima). 

*Mutilla aniens Gerstaecker, 1874. Arch. Naturges. 40: 323. $. 
*Mutilla mazatlanae Cameron, 1894. Biol. Cent.-Amer. Hym., v. 2, p. 294. 

S' 
Taxonomy: Mickel, 1938. Roy. Ent. Soc. London Trans. 87: 641. 
Biology: Linsley, 1960. Pan-Pacific Ent. 36: 36 (mating behavior). 

ocellaria rufidorsa Mickel. Ark. 

EPHUTA subgenus EPHUTA Say, pp. 772-773 

Taxonomy: Schuster, 1957. N. Y. Ent. Soc. Jour. 64: 7-84 (Part II of 
revis. treating Species Group Grisea). 

This revision reduces Ephuta of the Catalog to subgeneric rank. 

albiceps Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 75. 9. 

argcnticeps Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 16. $,9. 

auricapitis Schuster. Possibly the opposite sex of ecarinata ecarinata Schuster. 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 76. ?. 

babocfuivari Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 79, ?. 

cephalotes Schuster. The opposite sex is possibly minuta Schuster. 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 68. $. 

coloradella Schuster. Possibly the opposite sex of grisea Bradley, 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 73. 9. 

conchate Mickel. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 21. 9, 5- 

ecarinata ecarinata Schuster. The opposite sex is possibly sudatrix (Melander) 
or auricapitis Schuster. 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 49. $. 

ecarinata neomexicana Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 52. $. 

ecarinata pima Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 51. $. 

floridana dietrichi Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 40. 9. 

floridana floridana Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 34. 5, 9. 

grisea fuscosericea Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 14. $. 

grisea grisea Bradley. The opposite sex is possibly coloradella Schuster. 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 13. $. 

margueritae margueritae Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 56. 5, 9. 

margueritae xanthocephala Schuster. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 63. 5, 9. 

minuta Schuster. Possibly opposite sex of cephalotes Schuster. 

pauxilla pauxilla Bradley. Host: Dipogon s. sayi Bks. {teste H. E. Evans, 
in litt.) . 



SCOLIDAE 341 

pscphenophila Schuster. The opposite sex is possibly tentativa Schuster. 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 44. $. 

rufisquamis (Andre). The opposite sex is possibly tumacacori Schuster. 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 65. $. 

scrupea Say. 

Taxonomy: Schuster, 1957, loc. cit. supra, p. 26. ?, $. 

sudatrix (Melander). Possibly the opposite sex of ecarinata ecarinata 

Schuster. 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 54. ?. 

tegulicia Bradley. Possibly a syn. or subsp. of idiasta (Cameron) of Mexico. 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 47. $. 

tentativa Schuster. Possibly the opposite sex of pscphenophila Schuster. 
Taxonomy: Schuster, 1957, loc. cit, supra, p. 42. $. 

tumacacori Schuster. Possibly the opposite sex of rufisquamis (Andre) . 
Taxonomy: Schuster, 1957, loc. cit. supra, p. 82. ?. 

*EPHUTA subgenus XENOCHILE Schuster 

Ephuta subg-. Xenochile Schiister, 1957. N. Y. Ent. Soc. Jour. 64: 8. 

Type: Ephuta (Xenochile) kronibeini Schuster. Monotypic 
and by orig. desig. 

Place after subgenus Ephuta Say, p. 773. 

*krombeini Schuster. Ariz. (Ramsey Canyon in Huachuca Mts.). 

Ephula (Xenochile) kronibeini Schuster, 1957. N. Y. Ent. Soc. Jour. 64: 
9. $. 

Family SCOLIiDAE 

SCOLIA subgenus SCOLIA Fabricius 

The typical subgenus is restricted to the Palaearctic Region (teste 
Betrem and Bradley, 1964. Zool. Meded, 40: 90). 

SCOLIA subgenus DISCOLIA Saussure 
(=:Sco//a subgenus Sco/i'a Fabricius of Catalog, pp. 774—775) 

Transfer citations for Scolia, Lacosi, and Lisoca to the typical sub- 
genus above. Change citation for Discolia as follows: 

Scolia subg. Discolia Saussure, 1863. Soc. Ent. France Ann. (4) 3: 18. 
Type: Scolia (Discolia) nohilitata Fabricius. Desig. by 
Betrem and Bradley, 1964. 

Add the following to the subgeneric synonymy: 

Scolia subg. Scolia sect. Scolioides Guiglia and Capra, 1934. Soc. Ent. 
Ital. Boll. 66: 115. 

Type: Scolia hirta Schrank. Orig. desig, 

dubia monticola Cameron. Syn. of mexicana Saussure. 

*mexicana Saussure. 

Scolia (Lacosi) mexicana Saussure, 1958. Soc. Ent. France Ann. (3) 

6: 219. 9. 
Scolia (Scolia) dubia monticola Cameron (syn. by Bradley, 1964. In 

Betrem and Bradley, Zool. Meded. 40: 96. 

nohilitata otomita Saussure, changed status. 

Scolia (Scolia) otomita Saussure. Reduced to subsp. rank (teste Bradley, 
1964. In Bertem and Bradley, Zool. Meded. 40: 96). 

otomita Saussure. Subsp. of nohilitata Fabricius. 

Genus TRISCOLIA Saussure 
(=:Sco/ia subgenus Tr/sco/ia Saussure and Sichel of Catalog p. 775) 

Raised to generic rank (teste Betrem and Bradley, 1964. Zool. Meded. 



342 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

39: 435). Change bibliographic citation and genotype designation as follows: 
Scolia subg. Triscolia Saussure, 1863. Soc. Ent. France Ann. (4) : 3. 17. 
Type: Scolia (Triscolia) badia Saussure. By monotypy. 

Genus CAMPSOMERIS Guerin, pp. 775-776 

*CaTnpsomeris subg. Xanthocanipsotneris Bradley, 1957. Amer. Ent. Soc. 
Trans. 83: 70. 

Type: Tiphia tricincta Fabricius. Orig. desig. 
*Campsomeris subg. Pygodasis Bradley, 1957. Amer. Ent. Soc. Trans. 

83: 72. 

Type: Scolia quadrinotata Fabricius. Orig. desig. 
Taxonomy: Bradley, 1957. Amer. Ent. Soc. Trans. 83: 65-77 (subgeneric 
reclassification Amer. spp.), — Betrem, 1962. Ent. News 73: 207 
(type of Dielis). 

Delete heading, Subgenus CAMPSOMERIS Guerin, in Catalog, and refer- 
ence to Colpa; and change type designation for Elis subg. Dielis to {Scolia 
radula Fabricius =) Sphex plumipes Drury. 

*completa Rohwer. Ariz. (Chiricahua and Santa Rita Mts.) ; Mexico to El 
Salvador. (Xanthocatnpsomeris) 
Campsomeris (Campsomeris) cotnpleta Rohwer, 1927. Wash. Acad. Sci. 
Jour. 17: 151. 9. 

*druryi Cockerell. Syn. of quadrimaculata (Fabricius). 

ephippium ephippium (Say). (Pygodasis) 

fulvohirta (Cresson). (Xanthocatnpsomeris) 

limosa (Burmeister). (Xanthocainpsomeris) 

*maculata (Drury). Syn. of quadrimaculata (Fabricius). 

pilipes (Saussure). (Subgenus?) 

plumipes confluenta (Say). On flowers of Rubus, Arabis, Melilotus, 
Solidago, and Cephalanthus. 1 Host: Cotalpa lanigera (L.) 
(Dielis). 
Biology: Kurczewski, 1963. Ent. News 74: 21-24. 

plumipes fossulana (Fabricius). (Dielis) 

*Campsomeris szetschwanensis Forma D Betrem, 1932. Ent. Ber. 9: 414. 
$ (Syn. by Betrem, 1964. In Bradley, Spolia Zool. Mus. Hau- 
niensis 21: 35.) 

plumipes plumipes (Drury). (Dielis) 

* quadrimaculata (Fabricius). (Pygodasis) 

*Vespa maculata Drury, 1773. Illustr. Nat. Hist. 2: 74, pi. 39, fig. 2. 

5. Preocc. 
Scolia quadrimaculata Fabricius, 1775. Syst. ent., p. 355. 9. 
*Campsomeris druryi Cockerell, 1907. Entom. 40: 50. N. name. 
Campsomeris (Campsomeris) quadrinotata (Fabricius) (syn. by Bradley, 

1964. Ent. News 75: 105). 

quadrinotata (Fabricius). Syn. of quadrimaculata (Fabricius). 

tolteca (Saussure). (Dielis) 

*trifasciata nassauensis Bradley. Fla. (Key Vaca) ; Bahamas. (Dielis) 

Campsomeris (Campsomeris) trifasciata nassauensis Bradley, 1928. Acad. 
Nat. Sci. Phila. Proc. 80: 322. ?, $. 

trifasciata trifasciata (Fabricius). (Dielis) 

Genus TRIELIS Saussure 
( = Campsosco/io Betrem of Catalog, p. 776) 

Trielis Saussure, 1863. Soc. Ent. France Ann. (4) 3: 18. 

Type: Elis xantiana Saussure. By monotypy. 
Taxonomy: Betrem, 1962. Ent. News 73: 146. 



SAPYGIDAE 343 

Family SAPYGIDAE 

Genus SAPYGA Latreille, pp. 777-77B 

aculeata Cresson. Minn. 

louisi Krombein, N. J. (Princeton), Fla. (Orlando). 

pumila Cresson. Hosts: Ashmeadiella (A.) aridula Ckll., Megachile 
(Eutricharaea) rohindata (F.), 

Family FORMICIDAE 

Morphology: Wilson, 1954. Insectes Sociaux 1: 75-80, figs. 1-2 (poly- 
morphism). — Eisner and Brown, 1958 (1956). 10th. Int. Cong. 
Ent. Proc. 2: 503-508, figs. 1-9 (evolution and social significance 
of proventriculus) . — Eisner, 1957. Harvard Univ., Mus. Compar. 
Zool. Bui. 116: 439-490, pis. 1-25 (studies of the proventriculus).— 
G. C. and J. Wheeler, 1960. Psyche 67: 87-94 (techniques for 
study of ant larvae). — Nachtweg, 1961. Insectes Sociaux 8: 
369-381.— Nachtweg, 1963. Insectes Sociaux 10: 43-57 (sound 
organs).— Nachtweg, 1964 (1963). Insectes Sociaux 10: 359-378 

Taxonomy: Van Pelt, 1948. Fla. Ent. 30: 57-67, figs. 1-24 (key to 
species of Alachua County, Florida). — G. C. and J. Wheeler, 1963. 
The Ants of North Dakota. Univ. N. Dakota Press, Grand 
Forks, 326 pp., 56 figs., 20 illus., 81 maps.— Gregg, 1963. The 
Ants of Colorado: Their Taxonomy, Ecology and Geographic 
Distribution. Univ. of Colorado Press, Boulder. 792 pp., 26 tables, 
24 pis., 16 distribution maps, 163 altitudinal charts, 1 colored 
vegetation map. 

Economics: Smith and Weiss, 1942. U. S. Dept. Agr., Tech. Bui. 798: 
1-44 (relation of insects to spread azalea flower spot). — Smith, 
1950. In Pest Control Technology. Natl. Pest Control Assoc. 
Inc., Garrard Press, Champaign, 111., pp. 261-300, figs. 104-119 
(house-infecting ants). — Simeone, 1954. State Univ. N. Y., Col- 
lege of Forestry, Syracuse, Bui. 34: 1-19 (carpenter ants and 
their control).— Pimentel, 1955. Jour. Econ. Ent. 48: 28-30.— 
Reid, 1962. Ga. Agr. Expt. Sta. and Univ. Ga. Handbook, 71 pp., 
17 pis., 154 figs, (ants as intermediate hosts of chicken and turkey 
tapeworms).— Orlob, 1964 (1963). Ent. Expt. and Appl. 6: 95- 
106 (plant virus trans.). 

Embryology: Ledoux, 1954. Ins. Sociaux 1: 149-175, figs. 1-55, pi. 1 
{Oecophylla longinoda Latreille). 

Evolution: Brown and Wilson, 1960 (1959). Quart. Rev. Biol. 34: 278- 
294, figs. 1-33 (dacetine ants). 

Ecology: Van Pelt, 1956. Amer. Midland Nat. 56: 358-387, figs. 1-8, 
tables 1-3 (ants of Welaka Reserve, Fla.). — Kannowski, 1956. 
Amer. Midland Nat. 56: 168-185, table 1 (ants of Ramsey County, 
N. Dak.).— Van Pelt, 1958. Amer. Midland Nat. 59: i-57 (an- 
notated list of ants of Welaka Reserve, Fla.). — Carter, 1962, 
Elisha Mitchell Sci. Soc. Jour. 78: 150-204, fig. 1, table 1 (dis- 
tribution N. C. ants). — ibidem, 78: 1-18, tables 1-2 (distribution 
ants in N. C. Piedmont) .—Van Pelt, 1963. Amer. Midland Nat. 
69: 205-223, figs. 1-2, tables 1-2 (altitudinal distribution of ants 
in southern Blue Ridge Mts.) 

Physiology: Schneirla, 1953. Insect Physiology. John Wiley and Sons, 
Inc., N. Y., ch. 25, pp. 656-685 (basic problems in insect behavior), 
ibidem, ch. 26, pp. 685-723 (insect behavior in relation to its set- 
ting), ibidem, ch. 27, pp. 723-748 (modifiability in insect behavior), 
ibidem, ch. 28, pp. 748-780 (collective activities and social patterns 
among insects). — Glockner, 1957. Insectes Sociaux 4: 83-90, figs. 
1-3 (effect of hormones on metamorphosis) . — Eisner and Wilson 
1958 (1956). 10th. Int. Cong. Ent. Proc. 2: 509-513, figs. 1-3 
(radioactive studies, food transmission). — Wilson, 1959. Anat. 



344 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Rec. 134: 653 (pheromones in the organization of ant societies). — 
Eisner and Happ, 1962. Psyche 69: 107-116, text fig. 1; figs. 1-3, 
pi. 6 (infrabuccal pocket of a formicine ant, a social filtration de- 
vice).— Roth and Eisner, 1962. Ann. Rev. Ent. 7: 107-136, tables 
1-2 (chemical defenses of arthropods). — Wilson, 1963. Sci. Amer. 
208: 100-106, illus. (pheromones).— Wilson and Bossert, 1963. 
Recent Progress 19: 673-716, figs. 1-10, table 1 (chemical com- 
munication in animals). Academic Press, Inc., N. Y. 
Biology: Lafleur, 1941. N. Y. Ent. Soc. Jour. 49: 227-231 (civil 
disturbances in ant communities), ibidem, pp. 199-204 (communal 
disaffection).— Flanders, 1945. Sci. 101 (2619): 245-246 (Are 
castes determined by rate of egg deposition?). — Brackbill, 1948. 
Auk 65: 66-77 (anting by four species of birds). — Nixon, 1951. 
The Association of Ants with Aphids and Coccids. Commonwealth 
Inst. Ent. (London) pp. 1-35. — Flanders, 1951. Canad. Ent. 83: 
93-98 (role of ants in biological control of homopterous insects). — 
Flanders, 1952. Jour. Econ. Ent. 45: 38-39 (ovisorption as the 
mechanism causing worker development in ants). — Flanders, 1953. 
Sci. Monthly 76: 142-148, fig. 1 (caste determination in the social 
Hymenoptera). — Talbot, 1953. Mich. Univ., Contrib. Lab. Vertebr. 
Biol. 63: 1-13 (fauna and populations). — Wilson, 1953. Psyche 
60: 15-20 (caste determination). — Gosswald and Bier, 1954. In- 
sectes Sociaux 1: 229-246, figs. 1-5, tables 1-5 (caste determina- 
tion in genus Formica). — Chapman, 1954. Pan-Pacific Ent. 30: 
93-102, fig. 1, tables 1-2 (swarming of ants on summits of U. S. 
western mountains). — Bier, 1954. Insectes Sociaux 1: 1-19, figs. 
1-2 table 1.— Gosswald, 1955. Rev. Suisse de Zool. 62: 372-386 
(caste determination in ants). — Wilson and Eisner, 1957. In- 
sectes Sociaux 4: 157-166, figs. 1-6 (liquid transmission of food). — 
Talbot, 1957. Insectes Sociaux 4: 375-384 (populations). — Brian, 
1957. Ann. Rev. Ent. 2: 107-120 (caste determination).— Goss- 
wald and Bier, 1957. Insectes Sociaux 4: 335-348, tables 1-9 
(caste determination in genus Formica). — Weber, 1958 (1956). 
10th. Int. Cong. Ent. Proc. 2: 459-473, figs. 1-12 (evolution of 
ants and feeding habits). — Wenner, 1959. Amer. Midland Nat. 
62: 174-182 (ants of Bidwell Park, Chico, Calif.).— Kannowski, 
1959. Insectes Sociaux 6: 115-162, fig. 1, tables 1-4 (flight ac- 
tivities and colony founding of bog ants in southeastern Mich.). — 
Downey, 1962. Ent. News 73: 57-66 (ants associated with larvae 
of a lycaenid butterfly).— Ayre, 1962. N. Y. Ent. Soc. Jour. 70: 
159-167 (problems in using Lincoln Index for estimating size ant 
colonies). — Kincaid, 1963. Amer. Micros. Soc. Trans. 82: 101-105, 
figs. 1-5 (pollination of plants by ants). — Wilson, 1963. Ann. Rev. 
Ent. 8: 345-368, tables 1-2 (social biology) .—Wilson, 1963. Evolu- 
tion 17: 249-253, table 1 (social modification). — Ayre, 1963. Canad. 
Ent. 95: 712-715 (feeding habits). 

Subfamily DORYLINAE 

Taxonomy: Borgmeier, 1958. Studia Ent. 1: 197 (key for identifica- 
tion of females to genera, in tribe Ecitonini) . 

Biology: Schneirla, 1953. Insectes Sociaux 1: 29-41 (army ant queen). — 
Tafuri, 1955. N. Y. Ent. Soc. Jour. 63: 21-41, pis. 1-3, table 1 
(growth and polymorphism of larva of Eciton hamatum (F.)). — 
Schneirla, 1956 (1955). Smithsn. Inst. Ann. Rpt., pp. 379-406, 
figs. 1-4, pis. 1-2 (army ants). — Schneirla, 1957. Insectes Sociaux 
4: 259-298, figs. 1-2, tables 1-3 (comparison of functional patterns 
in army ants). — Schneirla, 1957. Amer. Phil. Soc. Proc. 101: 
106-133, figs. 1-8, tables 1-4 (theoretical consideration of cyclic 
processes). — Schneirla, 1958. Insectes Sociaux 5; 215-255, figs. 1- 
6, tables 1-2 (behavior and biology of Neivamyrmex nigrescens 
(Cress.) and iV. opacithorax (Emery)). — Brown, 1960. Psyche 66: 
25-27 (alarm and attack behavior). — Schneirla, 1960. Smith- 
sonian Treasury of Science. Simon and Schuster, N. Y., pp. 664- 



FORMICIDAE 345 

696 (the army ants). — Schneirla, 1961. Ztschr. f, Tierpsychol. 
18: 1-32, figs. 1-5, tables 1-4 (sexual broods and colony division 
of Neivamyrmex nigrescens (Cress.)). — Rettenmeyer, 1963. Univ, 
Kans. Bui. 44(9): 287-465, figs. 1-23, tables 1-4 (behavioral 
studies). 
Morphology: Hagan, 1954. Amer. Mus. Nat. Hist., Amer. Mus. Novitates 
1663: 1-12, figs. 1-2 (anatomy of queen of Eciton). — Ibidem 1664: 
1-17, figs. 1-18 (reproductive system of queen of Eciton). — Ibidem, 
1665: 1-20, figs. 1-21 (the oocyte cycle). — Lappano, 1958. In- 
sectes Sociaux 5: 31-66, figs, 1-25, tables 1-2 (morphological study 
of larval development in all worker broods of Eciton burchelli 
(Westw.)).— Whelden, 1963. N. Y. Ent. Soc. Jour. 71: 158-178, 
figs, 10 (A-D4), 11 (A-G), 12 (A-1) (reproductive system of 
vi^orker and female of Eciton burchelli (Westw.) and E. hamatum 
(F.)).— Whelden, 1963. Ibidem, pp. 246-261, figs. 1 ((A-K, 14 (A- 
E), 15 (A-D), 16 (A-E) (antennae and legs, E. hamatum and 
E. burchelli) ). 

Genus NOMAMYRMEX Borgmeier, Sup. 1, p. 108 

Morphology: Borgmeier, 1957. Rev. Brasil. Biol. 17: 390. 
Taxonomy: Borgmeier, 1958,. Studia Ent. 1: 201. 

Genus LABIDUS Jurine, Sup. 1, p. 108 

Morphology: Borgmeier, 1957. Rev. Brasil. Biol. 17: 389. 

coecus (Latreille). Ark. 

Biology: Kempf, 1961. Studia Ent. 4: 551-552 (as a cave ant). — Retten- 
meyer, 1963. Univ. Kans. Sci. Bui. 44 (9) : 418-424. 
Morphology: Borgmeier, 1957, loc. cit. supra, p. 390. 

Genus NEIVAMYRMEX Borgmeier, Sup. 1, pp. 108-110 

Morphology: Borgmeier, 1957, loc. cit. supra, pp. 392-393. 

ealifornicus (Mayr). Nev. 

carolinensis (Emery). Ohio, Va. 

fuscipennis (W. M. Wheeler). Kans.; Mexico. 

harrisii (Haldeman). 

Taxonomy: Borgmeier, 1958. Studia Ent. 1: 206-207. 

melanocephalus (Emery). Honduras. 

nigrescens (Cresson). Nebr., Colo. 

Biology: Rettenmeyer, 1963. Univ. Kans. Sci. Bui. 44 (9): 446-452, fig. 

23. 
Morphology: Borgmeier, 1957, loc. cit. supra, p. 393. 
Ecology: Gregg, 1963. The Ants of Colorado, pp. 286-288, map. 

opacithorax (Emery). Va., Ariz. 

Biology: Rettenmeyer, 1963. Univ. Kans. Sci. Bui. 44 (9): 437-446, 

fig. 22. 
Morphology: Borgmeier, 1957, loc. cit. supra, p. 393. 

pauxillus (W. M. Wheeler). La. 

pilosus mandibularis (M. R. Smith). N. Max. 

wheeleri (Emery). Ariz. 

Subfamily CERAPACHYINAE 
Biology: Wilson, 1958. Insectes Sociaux 5: 129-140 (behavior). 

ACANTHOSTICHUS subgenus CTENOPYGA Ashmead, p. 782 

texanus Forel. 

Taxonomy: M. R. Smith, 1955. Brooklyn Ent. Soc. Bui. 50: 48-50 (type 
lost). 



346 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Subfamily PONERINAE 

Revision: Brown, 1960. Harvard Univ., Mus. Comp. Zool. Bui. 122: 145- 

230, figs. 1-48 (tribe Amblyoponini) . 
Biology: Wilson, 1958. Evolution 12: 24-31 (nomadic and predatory 

behavior) . 

*Genus AMBLYOPONE Erichson 
( = Sfigmafomma Roger of Catalog, pp. 782-783) 

Amblyopone Erichson, 1842. Arch. f. Naturgesch. 8: 260. 

Type: Amblyopone australis Erichson, By monotypy. 
Taxonomy: Brown, 1960. loc. cit. supra, pp. 155-169. 
Morphology: Eisner, 1957. Harvard Univ., Mus. Compar. Zool. Bui. 

116: 476, pi. 22, fig. 91 (proventriculus). 
Biology: Haskins, C. P. and E. F., 1951. Amer. Midland Nat. 45: 432- 

445, fig. 1 (colony founding of A. australis Erichson). 
Amblyopone replaces Stigmatomma, which is a synonym (teste Brown, 

1960, loc. cit. supra). 

oregonense (W. M. Wheeler), changed status. 

Stigmatomma pallipes oregonense W. M. Wheeler. Raised to sp. rank 
{teste Brown, 1960, loc. cit. supra, p. 169). 

pallipes (Haldeman), changed status. Ariz., Colo., Okla. 

Stigmatomma pallipes pallipes (Haldeman). Raised to sp. rank {teste 

Brown, 1960, loc. cit. supra, p. 169). 
Stigmatomma pallipes montigena Creighton (syn. by Brown, 1960, loc. cit. 

supra) . 
Stigmatomma pallipes subterranea Creighton (syn. by Brown, 1960, loc. 

cit. supra). 
Morphology: Whelden, 1957. N. Y. Ent. Soc. Jour. 65: 1-21, pi. 1, figs. 

1-6; pi. 2, figs. 1-7. 
Ecology: Gregg, 1963. The Ants of Colorado, pp. 279-280, 282, map. 

pallipes montigena Creighton. Syn. of pallipes (Haldeman). 

pallipes oregonense W. M. Wheeler. Raised to specific rank. 

pallipes pallipes (Haldeman). Raised to specific rank. 

pallipes subterranea Creighton. Syn. of pallipes (Haldeman). 

Genus PLATYTHYREA Roger, p. 783 

Taxonomy: Brown, 1952. Breviora 6: 4 (in key). 

punctata (F. Smith). 

Biology: Brown, 1957. Harvard Univ., Mus. Comp. Zool. Bui. 116: 229. 

♦Genus PRIONOPELTA Mayr 

Prionopelta Mayr, 1866. Akad. der Wiss. Wien, Math.-Nat. Kl. Sitzber. 
53: 503. 

Type: Prionopelta punctulata Mayr. By monotypy. 
Taxonomy: Brown, 1960, loc. cit. supra, pp. 173-178. 

Place after Platythyrea, p. 783. 

*antillana Forel. Fla. (Juniper Springs in Marion County). 

Prionopelta punctulata antillana Forel, 1909. Deut. Ent. Ztschr., p. 239. 
Biology: Kempf, 1961. Studia Ent. 4: 489-490. 

Genus ECTATOMMA F. Smith 
( = Ecfafomma subgenus Ectafomma F. Smith of Catalog, p. 783) 

Revision: Brown, 1958. Harvard Univ., Mus. Comp. Zool. Bui. 118: 206- 

211. 
Ectatomma raised to generic rank {teste Brown, 1958, loc. cit. supra). 

tuberculatum (Olivier). 

Taxonomy: Brown, 1958, loc. cit. supra, pp. 209, 211, 298, 299. — Kempf, 
1962. Studia Ent. 5: 2-3. 



FORMICIDAE 347 

*Genus GNAMPTOGENYS Roger 
l^Ecfatomma subgenus Parecfafomma Emery of Catalog, p. 783) 

Gnamptogenys Roger, 1863. Berlin. Ent. Ztschr. 7: 174. 

Type: Ponera tornata Roger. Desig. by Emery, 1911. 
Ectatoma subg. Parectatoma Emery (syn. by Brown, 1958, loc. cit. supra, 
p. 212.) 

hartmani (W. M. Wheeler). La. (Lucky); Honduras. Emendation in spell- 
ing of hartmanni ! of catalog. 
Taxonomy: Brown, 1958, loc. cit. supra, pp. 228, 230, 234, 302. 
Historical: Brown, 1961. Psyche 68: 69. 
Biology: Echols, 1964. Ent. Soc. Amer. Ann. 57: 137. 

Genus PROCERATIUM Roger, p. 784 

Sysphincta Roger (syn. by Brown, 1958, loc. cit. supra, p. 241). 
Biology: Brown, 1958 (1957). Psyche 64: 115. 

croceum (Roger). 

Biology: Brown, 1958, loc. cit. supra, pp. 246, 247. 

melinum (Roger). 

Taxonomy: Brown, 1958, loc. cit. supra, pp. 243, 246-248 (also biology). 
Brown, op. cit., p. 334 considers melinum a European species 
which very doubtfully occurs in North America. 

pergandei (Emery). Fla. 

Taxonomy: Brown, 1958, loc. cit. supra, pp. 243, 246, 247-248 (also 
biology) . 

silaceum Roger. Okla. 

Taxonomy: Brown, 1958, loc. cit. supra, pp. 241, 245-248 (also biology). 

Genus Sysph'mcfa Roger, p. 784 

Transfer to synonymy of Proceratium, p. 784. 

Genus DISCOTHYREA Roger, pp. 784-785 

Taxonomy: Borgmeier, 1957. Acad. Brasil. de Cien. An. 29: 124-125, 

figs. 36-39 ( $ ) .—Brown, 1958, loc. cit. supra, pp. 248-253. 
Biology: Brown, 1958 (1957). Psyche 64: 115. 

testacea Roger. Okla., S. C. 

Taxonomy: M. R. Smith, 1955. Brooklyn Ent. Soc. Bui. 50: 98 (probable 
type locality, collector) . — Brown, 1958, loc. cit. supra, pp. 253, 
841-342. 

PACHYCONDYLA subgenus PACHYCONDYLA F. Smith, p. 785 

Taxonomy: Kempf, 1961. Rev. Brasil Ent. 10: 189-204, figs. 1-13 
(Brazilian species). 

harpax (Fabricius). Jamaica, Trinidad. 

Taxonomy: Kempf, 1961, loc. cit. supra, p. 194. 
Biology: Kempf, 1961. Studia Ent. 4: 493. 

Genus BRACHYPONERA Emery 
(=:Euponera subgenus Brac/iyponera Emery of Catalog, p. 785) 

Raised to generic rank by Wilson, 1958. Harvard Univ., Mus. Compar. 
Zool. Bui. 119: 346. 

Genus TRACHYMESOPUS Emery 

[ziEuponera subgenus Trachymesopus Emery of Catalog, p. 786) 

Raised to generic rank by Wilson, 1958, loc. cit. supra, p. 352. 

gilva (Roger). Transf. to Cryplopone Emery, infra. 

stigma (Fabricius). 

* Ponera quadridentata Smith, 1859. Linn. Soc. London Jour., Zool. 3: 



348 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

143. (Syn. by Wilson, 1958, loc. cit. supra, p. 355.) 
Biology: Kempf, 1960. Studia Ent. 3: 427-428. 

♦Genus CRYPTOPONE Emery 

Cryptopone Emery, 1892. Soc. Ent. de France Bui. 61:CCLXXV. 

Type: ? Amblyopone testacea Motschulsky. By monotypy. 

Place after Trachymesopus, p. 786. 

gilva (Roger). Ky., N. C. Transf. from Trachymesopusy p. 786 {teste Brown, 
1963. Breviora No. 190, p. 6). 

Genus PONERA Latreille, pp. 786-787 

coarctata pennsylvanica Buckley. Ariz., Ark., Okla. 

Biology: G. C. and J. Wheeler, 1963. The Ants of North Dakota, pp. 

91-92, map. 
Ecology: Gregg, 1963. The Ants of Colorado, pp. 281-282, 284, map. 

♦exotica M. R. Smith. N. C. (Piedmont and Coastal Regions; apparently 
introduced) . 
Ponera exotica M. R. Smith, 1962. Acta Hym. 1 (4) : 377-382, fig. 1, 
A, B, 2, 9. 

inexorata W. M. Wheeler. Ariz. (Cochise County, 5400 ft.) 

opacireps Mayr. Uruguay. 

Taxonomy: Kempf, 1962. Studia Ent. 5: 7-9, figs. 8, 9. 

Biology: Kempf, 1960. Studia Ent. 3: 391. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 283-284, map. 

Irigona opacior Forel. Oreg. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 284-286, map. 

Genus ODONTOMACHUS Latreille, pp. 787-788 

Morphology: Eisner, 1957. Harvard Univ., Mus. Compar. Zool. Bui. 116: 
475-476, pi. 22, fig. 90 (proventriculus). 

haematoda insularis Guerin. 

Biology: Taylor and Wilson, 1961. Psyche 68: 142. 

Subfamily PSEUDOMYRMECINAE 

Taxonomy: G. C. and J. Wheeler, 1956. Ent. Soc. Amer. Ann. 49: 376- 
379, pi. 3, figs. 1-6 (larva). 

Genus PSEUDOMYRMEX Lund, Sup. 1, p. 112 

Taxonomy: Kempf, 1958. Studia Ent. 1: 434 (gracilis (Fabricius) 

group). 
Biology: Brown, 1960. Ecology 41: 590. 
Morphology: Eisner, 1957, loc. cit. supra, pp. 452-453, pi. 1, fig. 5 

(proventriculus). 

apache Creighton. 

Biology: Creighton, 1963. Amer. Mus. Novitates 2156: 1-4, table 1. 

elongatus (Mayr). 

Taxonomy: Creighton, 1955. N. Y. Ent. Soc. Jour. 63: 17-20 (also key 
U. S. species of Pseudomyrmex) . 

gracilis mexicanus (Roger). Fla. (Dade County). Probably introduced. 
Taxonomy: G. C. and J. Wheeler, 1956, loc. cit. supra, pp. 385-386, pi. 1, 
figs. 18-21; pi. 2, fig. 3 (larva). — Brown, 1957. Harvard Univ., 
Mus. Comp. Zool. Bui. 116: 235. 

paUidus (F. Smith). 

Physiology: Blum and Callahan, 1963. Psyche 70: 69-74 (morphology 
and physiology of poison glands and venom). 



FORMICIDAE 349 

Subfamily MYRMICINAE 

Taxonomy: G. C. and J. Wheeler, 1960. Ent. Soc. Amer. Ann. 53: 98- 
110, pi. 1; figs. 1-3 (ant larvae). — G. C. and J. Wheeler, 1960. 
Ent. Soc. Wash. Proc. 62 : 1-32, pi. 1-2 (ant larvae) . 

MYRMICA subgenus MYRMICA Latreille, pp. 789-791 

Taxonomy: Ccllingvirood, 1958. Roy. Ent. Soc, London Proc, Ser. A.33: 
65-75, figs. 1-2 (Myrmica of Britian).— G. C. and J. Wheeler, 1960, 
loc. cit. supra, pp. 98-110, figs. 1-3 (larvae).— G. C. and J. Wheeler, 
1963. The Ants of North Dakota, pp. 94-108, figs. 

Biology: Brian, 1957. Insectes Sociaux 4: 177-190, fig. 1, table 1 (growth 
and development of colonies). — Weir, 1958. Insectes Sociaux 5: 
315-339, figs. 1-7, tables 1-9 (polyethism in workers) .—Weir, 
1958. Jour. Ins. Physiol. 1: 352-360, figs. 1-5 (effect of tempera- 
ture variation on queen ovisposition and colony formation). — 
Kannowski, 1959. Insectes Sociaux 6: 143-144 (flight compari- 
son).— Weir, 1959. Insectes Sociaux 6: 271-290, figs. 1-5 (in- 
fluence of worker age on trophogenic larval dormancy). — Ibidem, 
pp. 167-201, fig. 1-tables 1-9 (egg masses and early larval growth). 
—Brian and Hibble, 1963. Insectes Sociaux 10: 71-82, figs. 1-2, 
tables 1-5.— Weir, 1959. Physiol. Zool. 32: 63-77, figs. 1-8 (in- 
terrelation of queen and worker oviposition). — Carr, 1962. In- 
sectes Sociaux 9: 177-211. — Brian and Hibble, 1963. Insectes 
Sociaux 10: 71-82. 

Physiology: Weir, 1958 (1957). Quart. Jour. Micros. Sci. 98: 499-506, 
figs. 1-2 (functional anatomy of mid-gut). 

brevinodis brevinodis Emery. 

Biology: Kannowski, 1959, loc. cit. supra, p. 121 (p. 155, pleometrosis) . — 

G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 99-101, map. 
Ecology: Gregg, 1963. The Ants of Colorado, pp. 291-292, 294, map. 

brevinodis brevispinosa W. M. Wheeler. Alta., Wash. 

Taxonomy: Kannowski, 1956. Amer. Midland Nat. 56: 176, $ (also 

brevinodis discontinua S). — Gregg, 1961. N. Y. Ent. Soc. Jour. 

69: 211 5. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 296-297, 299, map. 

brevinodis discontinua Weber. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 298-300, map. 

brevinodis sulcinodoides Emery. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 293-295, 297, map. 

laevinotlis Nylander. Syn. of rubra (Linnaeus). 

lobicornis fracticornis Emery. 

Biology: Kannowski, 1957. Psyche 64: 1-5 (host of Leptothorax 
provancheri Emery). — Kannowski, 1959, loc. cit. supra, pp. 121- 
124, 155-156.— G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 104- 
106, map. 

Ecology: Gregg, 1963, loc. cit. supra pp. 300, 302-303, map. 

punctiventris punctivenlris Roger. Ark., Mass. 

Taxonomy: Gregg, 1961. N. Y. Ent. Soc. Jour. 69: 211. 5. 
Biology: Kannowski, 1959, loc. cit. supra, p. 124. 

punctiventris pinetorum W. M. Wheeler. N. Y. 

*rubra (Linnaeus). 

Formica rubra Linnaeus, 1758. Syst. Nat., Ed. 10, v. 1, p. 580. 

Myrmka (Myrmica) lacvinodis Nylander (syn. by Yarrow, 1955. Roy. 
Ent. Soc, London, Proc, Ser. B, 24: 113-115). 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, pp. 4-5 (larva). 

Biology: Brian, 1951. Experientia 7: 182 (caste determination). — Brian, 
1951. Physiol. Compar. and Oecol. 2: 248-262, figs. 1-2, tables 
1-7.— Brian, 1953. Ibidem 3: 25-36, tables 1-2 (oviposHion by 
workers). — Brian, 1954. Insectes Sociaux 1: 101-122. fig. 1-3, 



350 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

tables 1-6 (caste differentiation, larval dormancy, winter size, 
vernalization). — Brian, 1955. Ibidem 2: 1-34, figs. 1-5, tables 1-12 
(caste differentiation, growth of workers and intercastes) . — Brian, 
1955. Ibidem 2: 85-114, figs. 1-5, tables 1-8 (caste differentiation, 
larval dormancy, winter size, and vernalization). — Brian M. V. 
and A. D., 1955. Evolution 9: 280-290, figs. 1-2, tables 1-3 
(macrogynes and microgynes). — Brian, 1956. Insectes Sociaux 3: 
869-394, figs. 1-3, tables 1-8 (caste differentiation, controlled larval 
nutrition) .—Brian, 1956. Jour. Anim. Ecol. 25: 319-337, fig. 1, 
tables 1-2 (segregation of Myrtnica spp.). — Brian, 1957. Insectes 
Sociaux 4: 191-210, fig. 1, tables 1-7 (serial organization of brood). 
— Brian, 1957. Physiol. Compar. and Oecolog. 4: 329-345, figs. 
1-3, tables 1-8 (food distribution and larval size in cultures). — 
Weir, 1958. Insectes Sociaux 5: 315-339, figs. 1-6, tables 1-9 
(polyethism in workers). — Brian 1962. Ibidem 9: 295-310, illus. 
(social conditions affecting early larval differentiation). — Brian, 
1963. Ibidem 10: 91-102 (caste differentiation). 

Morphology: Allen, 1957. Ent. Monthly Mag. 93: 136-139 (intercastes). 

Physiology: Weir, 1959. Insectes Sociaux 6: 375-386, figs. 1-6, table 
1 (retro-cerebral endocrine glands and their relation to larval 
growth and development). 

sabuleti americana Weber. N. C, Ark. 

Taxonomy: Kannowski, 1956. Amer. Midland Nat. 56: 175. $. 
Biology: P. B. and P. M. Kannowski, 1957. Ohio Jour. Sci. 57: 371-374 
(mating activities). — G. C. and J. Wheeler, 1963, loc. cit. supra, 
pp. 96-99, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 308-310, 312, map. 

sabuleti hamulata Weber. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 300-301, map. 

scabrinodis mexicana W. M. Wheeler. 

Taxonomy: Gregg, 1961. N. Y. Ent. Soc. Jour. 69: 211. 5. 

schencki emeryana Forel. 

Biology: Kannowski, 1959. Insectes Sociaux 6: 121. — Medler, 1958. Ent. 

Soc. Wash. Proc. 60: 258 (swarming). — G. C. and J. Wheeler, 

1963, loc. cit. supra, pp. 103-104, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 311-312, 314, map. 

schencki spatulata M. R. Smith. ? N. Y. 

striolagaster Cole. Colo. 

Taxonomy: Cole 1957. Tenn. Acad. Sci. Jour. 32: 208-209 (S).— Gregg, 

1961, loc. cit. supra, p. 211. 5 . 
Ecology: Gregg, 1963, loc. cit. supra, pp. 313-314, map. 

*Genus PARAMYRMICA Cole 

Paramyrmica Cole, 1957. Tenn. Acad. Sci. Jour. 32: 37-42. 

Type: Paramyrmica colax Cole. By monotypy. 
Taxonomy: G. C. and J. Wheeler, 1959. Tenn. Acad. Sci. Jour. 34: 219, 

fig. 1, A-G (larva). — Gregg, 1961, loc. cit. supra, pp. 209-220, 

fig. 1. 
Place after Myrmica subg. Myrmica, p. 791. 

*coIax Cole. Tex. (Limpia Canyon in Davis Mts., Jeff Davis Co.). Inquiline 
in nest of Myrmica (Myrmica) striolagaster Cole. 
Paramyrmica colax Cole, 1957. Tenn. Acad. Sci. Jour. 32: 37-41, fig. 1, 

a, b, c. 5 9. 
Taxonomy: G. C. and J. Wheeler, 1959, loc. cit. supra, pp. 219-220, fig. 
1, A-G (larva).— Gregg, 1961, loc. cit. supra, pp. 213-215. 5 2. 

rugiventris (M. R. Smith). Ariz., Colo. Transf. from Tetramorium, p. 824 
(teste Gregg, 1961, loc. cit. supra, p. 215). 
Taxonomy: Gregg, 1961, loc. cit. supra, pp. 210, 215-220 fig. 1. 5 $ 5- 
Historical: Brown, 1957. Breviora No. 72, pp. 5-7. 



FORMICIDAE 351 

Genus MANICA Jurine, p. 791 

Taxonomy: Cole, 1957. Tenn. Acad. Sci. Jour. 32: 213 (key to males). — 
G. C. and J. Wheeler, 1960. Ent. Soc. Wash. Proc. 62: 5 (larva).— 
G. C. and J. Wheeler, 1963, loc. cit, supra, pp. 108-110, figs. 

bradleyi (W. M. Wheeler). 

Taxonomy: Cole, 1957, loc. cit. supra, pp. 210-212, fig. la (? $). — G. C. 
and J. Wheeler, 1960, loc. cit. supra, p. 6 (larva). 

hunteri (W. M. Wheeler). 

Taxonomy: Cole, 1957, loc. cit. supra, pp. 212-213, fig. 1, b ($). 

mutica (Emery). 

Taxonomy: Cole, 1957, loc. cit. supra, p. 213, fig. 1, c {$). — G. C. and J. 

Wheeler, 1960, loc. cit. supra, pp. 5-6 (larva). 
Ecology: Gregg, 1963, loc. cit. supra, pp. 314-317, map. 
Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 109-110, map. 

POGONOMYRMEX subgenus POGONOMYRMEX Mayr, pp. 791-794 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit, supra, p. 2 (larva). 
Morphology: Eisner, 1957. Harvard Univ., Mus. Compar. Zool. Bui. 116: 
477-478, pi. 23, figs. 95-96 (proventriculus). 

apache W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 318-319, map. 

badius (Latreille). La. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 2 (larva). 
Physiology: Wilson, Durlach, and Roth, 1959 (1958). Psyche 65: 108- 
114 (chemical releasers of necrophoric behavior). — Wilson, 1959 
(1958). Psyche 65: 41-51, fig. 1, table 1 (chemical releasers of 
alarm and digging). 

barbatus barbatus (F. Smith). Ark. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 2 (larva). 

Biology: Young and Howell, 1954. Okla. Acad. Sci. Proc. 35: 60-62 (mat- 
ing sw^arms). 

Ecology: Gregg, 1963, loc. cit. supra, pp. 319-321, 323, map. 

Economics: Gordon, 1943. Jour. Econ. Ent. 36: 354 (damage to air- 
plane runways). — Young and Gonzalez, 1957. Mexico. Secretaria 
de agricultura y ganaderia, fol. tec. 23: 1-20, figs. 1-6, tables 1-5. 

barbatus fuscatus Emery. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 322-324, map. 

barbatus rugosus Emery. Colo., Tex. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 325, 327, map. 

maricopa barnesi M. R. Smith. Utah. 

maricopa maricopa W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 326-328, map. 

occidentalis comanche W. M. Wheeler. La. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 332-334, map. 

occidentalis occidentalis (Cresson). 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 118-120, figs., 

map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 328-333, map. 
Physiology: Weber, 1959. Ent. News 70: 85-90 (stings, also population). 

subdentatus Mayr. Nev., Oreg. 

POGONOMYRMEX subgenus EPHEBOMYRMEX W. M. Wheeler, p. 794 

Biology: Creighton, 1956. Psyche 63: 54-66, fig. 1. 

imberbiculus W. M. Wheeler. Colo. 

Taxonomy: Creighton, 1956. Psyche 63: 63-64, fig. 1 ($). 
Ecology: Gregg, 1963, loc. cit. supra, pp. 334-336, map. 



352 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Genus STENAMMA Westwood, pp. 794-795 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 120-122, maps 
lU-11. 

brevicorne (Mayr). ? Colo. 

Biology: Kannowski, 1958. Ent. News 69: 231-233 (swarming of males). 
Ecology: Gregg, 1963, loc. cit. supra, pp. 346-348, 350, map. 

diecki Emery. Colo. 

Biology: G. C. and J, Wheeler, 1963, loc. cit. supra, pp. 121-122, map. 
Ecology: Gregg, 1963, loc. cit. s:upra, pp. 349-350, map. 

huachucanum M. R. Smith. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 350-351, 353, map. 

impar Forel. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, p. 122, map. 

meridionale M. R. Smith. N. C, Okla. 

occidentale M. R. Smith. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 352-354, map. 

APHAENOGASTER subgenus ATTOMYRMA Emery, pp. 795-798 

ashmearli (Emery). N. C. 

flemingi M. R. Smith. Ala., Ky., La., N. C. 

Aphaenogaster (Altotnyrma) niacrospina M. R. Smith (syn. by M. R. 
Smith, 1957. Brooklyn Ent. Soc. Bui. 52: 113). 

floridana M. R. Smith. N. C, S. C. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 7 (larva). 

fulva Roger. Colo., Fla. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 336-337, 339, map. 

huachucana huachucana Creighton. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 338-340, map. 

niacrospina M. R. Smith. Syn. of flemingi M. R. Smith. 

mariae Forel. Ky. 

*megoinmatus M. R. Smith. Nev., Ariz., Calif., Oreg. 

Aphaenogaster (Attomyrnia) megonimatns M. R. Smith, 1963. N. Y. Ent. 
Soc. Jour. 71: 244-246, fig. 1, A. B. 2. 

rudis picea Emery. 

Biology: Haskins, 1960. N. Y. Ent. Soc. Jour. 68: 66-67 (longevity of 
fertile females). 

rudis rudis Emery. Colo., Wyo. 

Biology: Brown 1959 (1958). Psyche 65: 39-40 (nest raided by Formica 

subintegra Emery.) 
Ecology: Gregg, 1963, loc. cit. supra, pp. 340-342, map. 

subterranea valida W. M. Wheeler. Rocky Mt. Region from Colo, to B. C. 

and west to Utah. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 342-344, map. 

uinta W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 344-345, 347, map. 

Genus NOVOMESSOR Emery, pp. 798-799 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 8 (larva). 

albisetosus ( Mayr) . 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, pp. 8, 10, pi. 2, 
fig. 1 (larva). 

cockerelli (Andre). 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 10 (larva). 



FORMICIDAE 353 

Genus VEROMESSOR Forel, p. 799 

Taxonomy: G. C. and J. Wheeler, 1956. Psyche 63: 142-143 (larva). 

anrlrci (Mayr). 

Biology: McCluskey, 1958. Science 128: 536-537, fig. 1 (daily rhythm 
of male). 

lariversi M. R. Smith. 

Taxonomy: Cole, 1963. Ent. Soc. Amer. Ann. 56: 680, 682, figs. 1, B; 
fig. 3, B; fig. 4, B (5 $ $). 

lobognathus (Andrews). N. Dak. 

Taxonomy: G. C. and J. Wheeler, 1956. Psyche 63: 143-145, fig. 2, A-J 

(larva). — Cole, 1963, loc. cit. supra, pp. 680-682, fig. 1, C; fig. 2, 

C; fig. 3, C; fig. 4, C (5 9 5). 
Biology: G. C. and J. Wheeler 1956. Psyche 63: U0-U2.— Ibidem, 1959. 

Ent. Soc. Amer. Ann. 52: 176-179, figs. 1-5 (behavior, nest 

structure). — Ibidem, 1963, loc. cit. supra, pp. 123-125, figs., map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 354-356 map. 

pergandei (Mayr). 

Taxonomy: Cole, 1963, loc. cit. supra, p. 681 (5). 

*smithi Cole. Nev. (Mercury in Nye County). 

yeromessor smithi Cole, 1963. Ent. Soc. Amer. Ann. 56: 678-680, fig. 1, 
A; fig. 2, A; fig. 3, A; fig. 4, A. 5 9 $. 

PHEIDOLE subgenus PHEIDOLE Westwood, pp. 799-806 

Taxonomy: Creighton, 1957. N. Y. Ent. Soc. Jour. 65: 203-212 (vasliti 
complex).— Gregg, 1958. N. Y. Ent. Soc. Jour. 66: 7-48 (key 
to species of the U. S.). — G. C. and J. Wheeler, 1963, loc. cit. 
supra, pp. 125-131, maps 13, 14. 

Biology: Weber, 1948. Ent. News 59: 31-35 (food of larvae and adults). 

bahai (Forel). 

Taxonomy: M. R. Smith, 1955. Brooklyn Ent. Soc. Bui. 50: 99 (opinions 
of authors concerning correct taxonomic placement) . 

bicarinata bicarinata Mayr. 

Pheidole (Pheidole) vinelandica buccalis W. M. Wheeler (syn. by Cole, 

1956. Tenn. Acad. Sci. Jour. 31: 114). 
Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 126-127, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 408-409, map. 

bicarinata longula Emery. N. Dak. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, p. 127, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 408, 410, map. 

*bicarinata paiute Gregg. {In subarmata group.) Nev. (Goldfield). 

Pheidole bicarinata paiute Gregg, 1958. N. Y. Ent. Soc. Jour. QQ: 17-18. 
2 U. 
bicarinata vinelandica Forel. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 127-128, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 410, 412-413, map. 
Economics: Reid and Nugara, 1961. Jour. Parasitol. 47: 885-889, pi. 
1, figs. 1-16. 

calif ornica niicula W. M. Wheeler. Raised to sp. rank. 

californica oregonica Emery. 

Pheidole (Pheidole) californica shoshoni Cole. (syn. by Gregg, 1958, 
loc. cit. supra, p. 33). 

californica shoshoni Cole. Syn. of californica oregonica Emery. 

ceres W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 413-415, map. 

cockerelli W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 415-417, map. 



354 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

constipate W. M. Wheeler. Subsp. of fluridana Emery. 

creightoni Gregg. Calif. 

davisi W. M. Wheeler. N. C. 

dentata Mayr. 

Biology: Van Pelt, 1950. Ent. News 61: 161-163, fig. 1, A, B, C 
(parasitism by a chalcid). 

dentigula M. R. Smith. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. siipra, p. 12 (larva). 

desertorum W. M. Wheeler. Okla., N. Mex. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 417-419, map. 

flavens sculptior Forel. Delete from Catalog; a misdetermination {teste 
Gregg, 1958, loc. cit. supra, p. 45). 

floridana constipata W. M. Wheeler. 

Pheidole (Pheiilole) constipata W. M. Wheeler. Reduced to subsp. rank 
{teste Gregg, 1958, loc. cit. supra, p. 35). 

floridana floridana Emery, changed status, N. C. 

Pheidole (Pheiflole) floridana lauta W. M. Wheeler (syn. by Gregg, 

1958, loc. cit. supra, p. 35). 
Pheidole (Pheidole) floridana Emery. Reduced to subsp. rank {teste 
Gregg, 1958, loc. cit. supra, p. 35). 

floridana lauta W. M. Wheeler. Syn. of floridana floridana Emery. 

grallipes W. M. Wheeler. 

Pheidole grallipes var. vistana Forel (syn. by Gregg, 1958, loc. cit. supra, 
p. 22). 

grallipes var. vistana Forel. Syn. of grallipes W. M. Wheeler. 

hyatti hyatti Emery. Okla., Tex. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 419-420, map. 

lamia W. M. Wheeler. 

Biology: Gregg, 1956. Ent. News 67: 37-39 (also distribution). 

macclendoni W. M. Wheeler. 

Taxonomy: Gregg, 1958, loc. cit. supra, pp. 46-47. 24!.— Cole, 1958 (1957). 
N. Y. Ent. Soc. Jour. 65: 130-131 (5, especially 24:). 

megaccphala ( Fabricius ) . 

Biology: Broekhuysen, 1948. Union So. Africa Dept. Agr. Bui. 266, pp. 
1-40, figs. 1-22, tables 1-10.— -Brown, 1958. Acta Hym. 1: 47.— 
Weber, 1960. Ent. Soc. Wash. Proc. 62: 232.— Taylor and Wilson, 
1961. Psyche 68: 143.— Kempf, 1962. Studia Ent. 5: 18-19. 

metalleseens metallescens Emery. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 12 (larva). 

micula W. M. Wheeler, changed status. 

Pheidole (Pheidole) californica micula W. M. Wheeler. Raised to sp. 
rank {teste Gregg, 1958, loc. cit. supra, p. 33). 

militicida W. M. Wheeler. 

Biology: Creighton, 1960 (1959). Psyche 66: 1-12, fig. 1 (major worker). 

♦moerens W. M. Wheeler. Ala. (Mobile). Apparently introduced. Originally 
described from Puerto Rico. 
Pheidole moerens W. M. Wheeler, 1908. Amer. Mus. Nat. Hist. Bui. 24: 
136-138. 5 n. 

morrisii morrisii Forel. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 12 (larva). 

*obtusospinosa Pergande. Syn. of *8ubdentaia Pergande. 

pilifera coloradensis Emery. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 128-130, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 419, 421-422, map. 



FORMICIDAE 355 

pilifera pilifera (Roger). N, Dak. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 130-131, map. 

*rugulosa Gregg. {In subarmata group.) Ariz. (Harding's Ranch near 
Tucson.) 
Pheidole rugulosa Gregg, 1958. N.Y. Ent. Soc. Jour. 66: 26-29. 5 U 

? $. 

senex Gregg. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 422-424, map. 

sitarches campestris W. M. Wheeler, changed status. Tex. and Colo, to Mo. 
and Miss. 

Pheidole (Pheidole) soritis W. M. Wheeler, in part of Sup. 1. Remove 
sitarches rufesrens var. campestris W. M. Wheeler and its 
synonym, rufescens from synonymy of soritis and place as a 
subspecies of sitarches W. M. Wheeler {teste Gregg, 1958, loc. cit. 
supra, pp. 30, 38). 

Ecology: Gregg, 1963, loc. cit. supra, pp. 424-425, map. 

sitarches sitarches W. M. Wheeler. Tex.; Mexico. 

Biology: Wilson, 1957. Psyche 64: 46-50 (nuptial flights). 

sitarches soritis W. M. Wheeler, changed status. Ariz., N. Mex., Tex., Utah; 

Mexico. 
Pheidole (Pheidole) soritis W. M. Wheeler. A subsp. of sitarches not a 

species {teste Gregg, 1958, loc. cit. supra, pp. 30, 38). 
Pheidole (Pheidole) tepicana cavigenis W. M. Wheeler (syn. by Gregg, 

1958, loc. cit. supra, p. 48). 

soritis W. M. Wheeler, of Sup. I. A subsp. of sitarches W. M. Wheeler. N.B., 
sitarches rufescens var. campestris W. M. Wheeler and its synonjmi 
sitarches rufescens W. M. Wheeler, is another subsp. of sitarches. 

*subdentata Pergande. Ariz.; Mexico. 

Pheidole subdentata Pergande, 1895. Calif. Acad. Sci. Proc. 5: 888. $. 
Pheidole (Pheidole) vasliti arizonica Santschi (syn. by Creighton, 1957. 

N. Y. Ent. Soc. Jour. 65: 211). 
*Pheidole obtusospinosa Pergande, 1895. Calif. Acad. Sci. Proc. 5: 889. 

H. (Syn. by Creighton, 1957, loc. cit. supra, p. 211.) 

tepicana cavigenis W. M. Wheeler. Syn. of sitarches soritis W. M. Wheeler. 
vasliti arizonica Santschi. Syn. of subdentata Pergande. 
vinelandica buccalis W. M. Wheeler. Syn. of bicarinata bicarinata Mayr. 
PHEIDOLE subgenus CERATOPHEiDOLE Pergande, p. 806 
Taxonomy: Gregg, 1958, loc. cit. supra, pp. 9, 16 (key to species of U. S.). 

grundmanni M. R. Smith. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 424, 426-427, map. 

Genus EPIPHEIDOLE W. M. Wheeler, p. 806 

inquilina W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 427-429, map. 

Genus SYMPHEIDOLE W. M. Wheeler, p. 807 

elecebra W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit supra, pp. 429-431, map. 

Genus CARDIOCONDYLA Emery, p. 807 

Taxonomy: Weber, 1952. Amer. Mus. Novitates No. 1548: 5. 
Biology: Wilson, 1960 (1959). Psyche %Q: 29-34 (tandem running in 
genus Cardiocondyla) . 

emeryi Forel. 

Taxonomy: Weber, 1952, loc. cit. supra, p. 5. 
nuda minutior Forel. Calif., Tex., La. An introduced form. Probably a na- 
tive of the Oriental Region. 



356 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

CREMATOGASTER subgenus ORTHOCREMA Santschi, pp. 807-808 

minutissima minutissima Mayr. N. C. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 14, pi. 2, fiers. 
11-13 (larva). 

minutissima missouriensis Emery. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 363-364, map. 

CREMATOGASTER subgenus CREMATOGASTER Lund, pp. 808-810 

Taxonomy: Buren, 1958. N. Y. Ent. Soc. Jour. 66: 119-134 (also biology). 
Biology: Soulie, 1962. Insectes Sociaux 9: 181-195 (colony foundation 
and development). 

cerasi (Fitch). 

Biology: Kannowski, 1959. Insectes Sociaux 6: 125. 

lineolata emeryana Creighton. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 358-360, map. 

lineolata lineolata (Say). 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 132-133, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 357, 359, map. 

punctulata Emery. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 361-362, 364, map. 

vermiculata Emery. N. C. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 13, pi. 2, figs. 
14-15 (larva). 

MONOMORIUM subgenus MONOMORIUM Mayr, pp. 810-811 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 133-136, fig., 
map 16. 

floricola (Jerdon). 

Biology: Brown, 1964. Ent. News 75: 15. 

*metoecus Brown and Wilson. Ala. (Tuscaloosa). Parasite of Monomorium 

mininrnvi (Buckley). 
Monomorium metoecus Brown and Wilson, 1957. Ent. News 68: 239-244, 

fig. 1, a and b, ergatogyne. 
Taxonomy: Wilson and Brown, 1958. Ent. News 69, 33-38, fig. 1, A and 

B ( 2 also biology). 

minimum (Buckley). Change heading in Sup. 1, p. 128 from minimum 
minimum to minimum. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 133-135, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 366-368, map. 

pharaonis (Linnaeus). 

Biology: Peacock, Hall, Smith and Goodfellow, 1950. Scot. Dept. Agr. 
Mi?c. Pub. No. 17: 1-50, figs. 1-5, tables 1-3.— Sudd, 1953. Adv. 
of Sci. (Brit.) 10: 17-18 (colony formation). — Peacock, Water- 
house and Baxter, 1955. Ent. Monthly Mag. 91: 37-42 (viability 
in regard to temperature and moisture). — Brown, 1958. Acta 
Hym. 1: 36. — G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 135- 
136, map.— Brown, 1964. Ent. News 75: 14-15.— Eichler, 1963. 
Deut. Ent. Ztschr. 10: 207-215. 

Morphology: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 16 (larva). 

Embryology: Smith and Peacock, 1957. Roy. Soc. Edinbh. Proc, Section 
B, 66: 235-261, figs. 1-4, table 1 (cytology). 

viridum peninsulatum Gregg. Fla. and Gulf Coast States to Ariz., including 
southern Colo. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 368-370, map. 

viridum viridum Brown. N. Y., Long Island (Coram, Selden). 



FORMICIDAE 357 

MONOMORIUM subgenus PARHOLCOMYRMEX Emery, p. 811 

destructor (Jerdon). 

Economics: Kempf, 1960. Studia Ent. 3: 506-507. 

Genus XENOMYRMEX Forel, p. 811 

Taxonomy: G. C. and J. Wheeler, 1960. Ent. Soc. Wash. Proc. 62: 19- 
20 (larva). — Creighton, 1957. Amer. Mus. Novitates 1843: 1-14 
(including key to species). 

florldanus floridanus Emery, changed status. Cuba, Bahamas. Another sub- 
species, /. skwarrae W. M. Wheeler, occurs in Mexico. 
Xenomyrmex stolli floridanus Emery. A subsp. of floridanus not a 
subsp. of stolli Emery (teste Creighton, 1957, loc. cit. supra, p. 6). 

SOLENOPSIS subgenus SOLENOPSIS Westwood, pp. 812-813 

Taxonomy: Snelling, 1963. Calif. Dept. Agr., Bur. Ent., Occas. Papers 

No. 3: 1-10, figs, 1-12 (keys to all castes). 
Physiology: Blum, Walker, Callahan and Novak, 1958. Science 128: 

306-307 (chemical, insecticidal and antibiotic properties of fire 

ant venom). 

aurea W. M. Wheeler, changed status. 

Soleuopsis (Solenopsis) aurea amblychila W. M. Wheeler (syn. by Snell- 
ing, 1963, loc. cit. supra, p. 7). 

aurea amblychila W. M. Wheeler. Syn. of aurea W. M. Wheeler. 

aurea aurea W. M. Wheeler. Raised to sp. rank. 

geminata geminata (Fabricius). 

Biology: Kempf, 1961. Studia Ent. 4: 507. 

saevissima richteri Forel. 

Taxonomy: Wilson, 1953. Evolution 7: 262-263, fig. 1 (origin of varia- 
tion) .—Brown, 1957. Quart. Rev. Biol. 32: 258-261, figs. 4-7.— 
Wilson and Brown, 1958. Evolution 12: 211-218, figs. 1-3, table 
1 (morphological changes in introduced population). 

Biology: Wilson, 1958. Sci. Amer. 198: 36-41, maps, illus.— Green, 1962. 
Jour. Econ. Ent. 55: 1003-1004, table 1. 

Morphology: Callahan, Blum and Walker, 1959. Ent. Soc. Amer. Ann. 
52: 573-590, figs. 1-19 (also histology of poison glands and sting). 

Physiology: Caro, Derbes and Jung, 1957. Amer. Med. Assoc. Arch. 
Derm. 75: 475-478, figs. 1-7.— Favorite, 1958. U. S. Pub. Health 
Serv., Pub. Health Rpts. 73: 445-448.— Wilson, Durlach and Roth 
1959 (1958). Psyche 65: 108-114 (chemical releases of necrophoric 
behavior). — Wilson, 1959. Science 129: 643-644 (nature of odor 
trail).— Wilson, 1962. Anim. Behavior 10: 134-147, figs. 1-8, 
tables 1-4 (chemical communication among workers-mass forag- 
ing) .—Blum and Callahan, 1960. XI Int. Kongr. F. Entom. Wien, 
Verb. B. Ill: 290-293 (chemical and biological properties of 
venom).— Wilson, 1962. Anim. Behavior 10: 148-158, figs. 1-5, 
tables 1-2 (chemical communication among workers — analysis of 
the odour trail). — Wilson, 1962. Ibidem, 159-164, table 1 (chemi- 
cal communication among workers — induction of social responses) . 

xyloni McCook. Colo. 

Biology: Wray, 1962. Jour. Econ. Ent. 55: 145 (apparent introduction 

into N. C.) 
Ecology: Gregg, 1963, loc. cit. supra, pp. 370-371, 373, map. 
Physiology: Blum, Roberts and Novak, 1961. Psyche 68: 73-74 (chemical 

and biological characters of venom). 

SOLENOPSIS subgenus EUOPHTHALMA Creighton, p. 813 

globularia littoralis Creighton. 

Taxonomy: G. C. and J. Wheeler 1960, loc. cit. supra, p. 22 (larva). 



358 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

SOLENOPSIS subgenus DIPLORHOPTRUM Mayr, pp. 813-814 

molesta molesta (Say). Colo. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 136-138, map. — 

Macnamara, 1945. Canad. Ent. 77: 40. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 372-373, 375, map. 

molesta validiuscula Emery. Pacific Coast States east to Idaho, Colo., 
N. Mex. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 374-375, 377, map. 

pergandei Forel. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, pp. 21-22 (larva). 

picta Emery. N. C. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 22 (larva). 

salina W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 376-378, map. 

tennesseensis M. R. Smith. Kans. (Kearny County), Calif, (near Azusa). 

texana texana Emery. N. C. 

texana truncorum Forel. Colo., Ariz., Calif. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 378-380, map. 

Genus ANERGATES Forel, p. 815 

atratulus (Schenck). Md. (Baltimore), N. Y. (Suffolk County, Long Island). 
Morphology: Meyer, 1955. Insectes Sociaux 2: 163-170, figs. 1-4. 

Genus EREBOMYRMA W. M. Wheeler, p. 815 

*Spelaeomyrmex W. M. Wheeler, 1922. Amer. Mus. Novitates No. 45: 
9. (Syn. by Wilson, 1962. Psyche 69: 63.) 

Type: Spelaeomyrmex urichi W. M. Wheeler. By monotypy. 

Biology: Wilson, 1962, loc. cit. supra, pp. 62-72, fig. 1. 

longii W. M. Wheeler. Okla. (Fort Sill). 

Genus MYRMECINA Curtis, pp. 815-816 

americana Emery. Colo., N. Mex. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 364-365. 367, map. 

LEPTOTHORAX subgenus NESOMYRMEX W. M. Wheeler, pp. 816-817 

Revision: Kempf, 1959. Studia Ent. 2: 391-432, figs. 1-31. 

wilda M. R. Smith. Tex.; Mexico. 

Taxonomy: Kempf, 1959, loc. cit. supra, 421-423, figs. 8, 28 (5 $). 

LEPTOTHORAX subgenus MYRAFANT M. R. Smith, pp. 817-819 

Taxonomy: Cole, 1958. Ent. Soc. Amer. Ann. 51: 535-538 (nitens- 
carinatus complex) . — G. C. and J. Wheeler, 1960. Ent. Soc. Wash. 
Proc. 62: 23 (larva). 

andrei Emery. Nev., Ariz., N. Mex. 

Taxonomy: Cole, 1958, loc. cit. supra, pp. 537-538 (5, ?, 5)- 

ambiguus ambiguus Emery. Slave of L. duloticus Wesson. 

Biology: Kannowski, 1959. Insectes Sociaux 6: 124 (150-151, parasitism; 
155 pleometrosis). — G. C. and J. Wheeler, 1963, loc. cit. supra, 
pp. 140-141, map. 

*carinatus Cole. Tex. (Limpia Canyon, Davis Mts., 5,400 ft.), Ariz. 

Leptothorax (Leptothorax) carinatus Cols, 1957, Tenn. Acad. Sci. Jour. 

32: 213-215, fig. 1. $ 9. 
Taxonomy: Cole, 1958, loc. cit. supra, pp. 537-538 (5, ?). 

curvispinosus Mayr. Okla., Ariz. 

Biology: Talbot, 1957. Ecology 38: 449-456 (populations). — Kannowski, 
1959, loc. cit. supra, p. 124. 



FORMICIDAE 359 

foveata M. R. Smith. Va. 

furunculus W. M. Wheeler. Wyo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 381, 382, map. 

*hispidus Cole. Tex. (Limpia Canyon, Davis Mts., 5,400 ft.) ; Mexico. 

Leptothorax (Leptothorax) hispidus Cole, 1957. Tenn. Acad. Sci. Jour. 
32: 42-45, fig. 1, a, b, c. 5 $. 

longispinosus Roger. 

Biology: Kannowski, 1959, loc. cit. supra, p. 125. 

mariposa W. M. Wheeler, changed status. 

Leptothorax (Myrafant) nitens nitens Emery, in part of Sup. 1. Re- 
move mariposa W. M. Wheeler from syn. and raise to sp. rank 
{teste. Cole, 1958, loc. cit. supra, p. 536). 

nitens Emery, changed status. Wash., Oreg., Nev. 

Lepothorax (Myrafant) nitens nitens Emery. Raised to sp. rank {teste 

Cole, 1958, loc. cit. supra, p. 536). 
Leptothorax (Myrafant) nitens heathii W. M. Wheeler. (Syn. by Cole, 

1958, loc. cit. supra, p. 536). 
Ecology: Gregg, 1963, loc. cit. supra, pp. 382-384, map. 

nitens heathii W. M. Wheeler. Syn. of nitens Emery. 

nitens nitens Emery of Sup. I. Raised to sp. rank; remove mariposa W. M. 

Wheeler from syn. and also raise it to sp. rank. 

obliquicanthus Cole. 

Taxonomy: Gregg. 1953. Breviora No. 22, pp. 1-3, fig. 1. 5. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 384-385, 387, map. 

rugatulus brunnescens W. M. Wheeler. Colo, and Utah to Mont, and the 
Dakotas. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 388-389, map. 

rugatulus rugatulus Emery. 

Taxonomy: Kannowski, 1956. Amer. Midland Nat. 56: 178-179 (5 ; also 

rugatulus brunnescens W. M. Wheeler) . 
Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 141-143, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 386-388, map. 

schaumi Roger. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 23 (larva). 

schmittii W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 388, 390-391, map. 

lexanus texanus W. M. Wheeler. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 24 (larva). 

tricarinatus neomexicanus W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit supra, pp. 393-394, map. 

tricarinatus tricarinatus Emery. Ariz., Idaho, N. Dak. 

Biology: G. C. and J. Wheeler, 1963, loc cit. supra, p. 143, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 391-393, map. 

tuscaloosae Wilson. N. C. 

LEPTOTHORAX subgenus DICHOTHORAX Emery, p. 820 

pergandei pergandei Emery. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, p. 23 (larva). 

LEPTOTHORAX subgenus LEPTOTHORAX Mayr, pp. 820-822 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 143-148, figs., 
maps 21-23. 

crassipilis W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 401-402, 405, map. 

duloticus L. G. Wesson. Enslaves Leptothorax ambiguus ambiguus Emery. 



360 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Biology: Talbot, 1957. Ecology 38: 449-456 (populations).— Kannowski, 
1959. Insectes Sociaux 6: 124-125 (150-151, parasitism). 

hirticornis Emery. Calif. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 145-146, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 403, 405, map. 

muscorum (Nylander). 

Biology: Kannowski 1959, loc. cit. supra, p. 125. — G. C. and J. Wheeler, 

1963, loc. cit. supra, pp. 143-145, map. 
Ecology: Brown, 1955. Ent. News 66: 47-50. — Gregg, 1963, loc. cit. supra, 
pp. 393-400, map. 

provanclieri Emery. Colo. Host: Myrmica lohicomis fracticomis Emery. 

Biology: Kannowski, 1957. Psyche 64: 1-5.— G. C. and J. Wheeler, 1963, 

loc. cit. supra, pp. 146-148, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 404-407, map. 

Genus HARPAGOXENUS Forel, pp. 822-823 
americanus (Emery). N. C, Ont. 

Genus TRIGLYPHOTHRIX Forel, p. 823 

striatidens (Emery). 

Biology: Taylor and Wilson, 1961. Psyche 68: 142. 

Genus TETRAMORIUM Mayr, pp. 823-824 

caespitum (Linnaeus). 

Biology: Brown, 1964. Ent. News 75: 15. 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, pp. 24-25 (larva). 

Historical: Brown, 1957. Breviora 72: 1-4. 

guineense (Fabricius). 

Biology: Taylor and Wilson, 1961. Psyche 68: 138.— Brown, 1964. Ent. 

News 75: 14-15. 
Ecology: Brown, 1958. Acta Hym. 1: 28. 

rugiventris M. R. Smith. Transf. to Paramyrmica Cole, p. 791. 

simillimum (F. Smith). 

Biology: Taylor and Wilson, 1961. Psyche 68: 142-143. 

Genus XIPHOMYRMEX Forel, p. 824 
Taxonomy: Cole, 1957. Tenn. Acad. Sci. Jour. 32: 209-210 ($). 

PARACRYPTOCERUS subgenus HARNEDIA M. R. Smith, p. 825 
Revision: Kempf, 1958. Studia Ent. 1: 65-144, pis. 1-8. 

rohweri (W. M. Wheeler). 

Taxonomy: Kempf, 1958, loc. cit. supra, pp. 129-132, pi. 2, fig. 8; pi. 4, 
fig. 2; pi. 6, fig. 12. 

texanus (Santschi). 

Taxonomy: Kempf, 1958, loc. cit. supra, pp. 123-127, pi. 2, fig. 5; pi. 4, 

fig. 4; pi. 6, fig. 7. 
Biology: Creighton, 1963. Psyche 70: 133-143, figs. 1-3. 

PARACRYPTOCERUS subgenus CYATHOMYRMEX Creighton, p. 825 

Revision: Kempf, 1958. Studia Ent. 1: 144-158, pi. 8. 

varians (F. Smith). 

Taxonomy: Kempf, 1958, loc. cit. supra, pp. 155-158, pi. 8. figs. 7, 8. 

STRUMIGENYS subgenus STRUMIGENYS F. Smith, pp. 825-826 

Taxonomy: G. C. and J. Wheeler, 1960, loc. cit. supra, pp. 25-26 (larva). 
—Brown, 1960 (1959). Psyche 66: 37-52, figs. 1-9 {qundlachi 
group).— Brovm, 1962. Psyche 69: 238-267, figs. 1-30 (Neotropi- 
cal species and key). 



FORMICIDAE 361 

Biology: Brown and Wilson, 1960 (1959). Quart. Rev. Biol. 34: 281. 

*eggersi Emery. Fla. (Miami, Fisher's Isl., Archbold Biol. Station). Prob- 
ably introduced. 

Strumigenys eggcrsi Emery, 1890. Soc. Ent. Ital. Bui. 22: 68, pi. 7, fig. 
9. 2 9. 

Taxonomy: Brown, 1960 (1959). Psyche 66: 46-47, figs. 2, 6 (5, $).— 
Brown, 1962. Psyche 69: 249, 257, 264. 

Biology: Weber 1952. Amer. Mus. Novitates No. 1554: 5. — Kempf, 1958, 
loc. cit. supra, p. 554. — Kempf, 1961. Studia Ent. 4: 515. 

*gundlachi Emery. Fla. (Royal Palm Ranger Station, Everglades Natl. Park; 

Key Largo.) Probably introduced. 
Strumigenys gundlachi Roger, 1862. Berlin Ent. Ztschr. 6: 253, pi. 1, 

fig. 18 a 5 not 9. 
Taxonomy: Brown, 1960 (1959). Psyche 66: 37-45, figs. 1, 5 ( 5 9).— 

Brown, 1962. Psyche 69: 249, 257, 264. 
Biology: Weber, 1952, loc. cit. supra, pp. 4-5. 

louisianae Roger. Ariz., Okla. 

Taxonomy: Brown, 1953. Psyche 60: 1-2.— G. C. and J. Wheeler, 1954. 
Psyche 61: 136-137, pi. 10, figs. 1-6 (larva) .—Brown, 1957. Quart. 
Rev. Biol. 32: 271.— Brown, 1961. Psyche 68: 64-67, 68 (varia- 
tion and synonymy).— Brown, 1962. Psyche 69: 246-247, 257, 263. 

Biology: Wilson, 1953. Ent. Soc. Amer. Ann. 46: 481-483.— Kempf, 
1958. Studia Ent. 1: 555. 

*silvestrii Emery. La. (Paradise). Apparently introduced. 

Strumigenys silvestrii Emery, 1905. Soc. Ent. ItaL Bui. 37: 168, fig. 27. 

5 9. 
Taxonomy: Brown. 1959. Studia Ent. (n.s.) 2: 25-28, fig. 1 (2 9).— 
Brown, 1962. Psyche 69: 246, 257, 259. 

SMITHISTRUMA subgenus WESSONISTRUMA Brown, p. 826 

Biology: Brown and Wilson, 1960 (1959). Quart. Rev. Biol. 34: 282.— 
Brown, 1964. Amer. Ent. Soc. Trans. 89: 190-197. 

angulata (M. R. Smith). 111. (Giant City State Park). 

pergandei (Emery). N. C., Mo., Kans. 

Taxonomy: G. C. and J. Wheeler, 1954. Psyche 61: 144-145 (larva). 
Note: W. L. Brown, 1964, loc. cit. supra, p. 197 remarks that this species 
was taken at Billerica, Mass. "the northernmost locality from 
which any dacetine ant has been taken in the New World." 

SMITHISTRUMA subgenus SMITHISTRUMA Brown, pp. 826-828 

Biology: Brown, 1964, loc. cit. supra, pp. 185-186, 189-191, 193-200. 

abdita (L. G. Jr., and R. G. Wesson). N. C. (Great Smoky Mt. Natl. Park, 
2600 ft.), 111., Iowa and Va. 
Taxonomy: Brown, 1964, loc. cit. supra, pp. 199-200 (abdita-missouriensis 
complex). 

bimarginata (L. G. Jr., and R. G. Wesson). 111. (Red Hills State Park). 

brevisetosa (M. R. Smith). Syn. of pilinasis (Forel). 

bunki W. L. Brown. Fla. (Duvall Co). 

*carolinensis W. L. Brown. S. C. (U. S. Hwy. 17, 1 mile east Ravenel), N. C. 
(Marshville in Union Co.). 
Smithistrnwa carolinensis W. L. Brown, 1964. Amer. Ent. Soc. Trans. 
89: 185-186, pi. 16, fig. 1, 2. 

cloydi Pfitzer. 

Taxonomy: Brown, 1964, loc. cit. supra, p. 191. 

clypeaia (Roger). Ark., Ala., N. C. (Great Smoky Mts. Nat. Park at 2,000 
ft.), N. J. 
Taxonomy: G. C. and J. Wheeler, 1954. Psyche 61: 143 (larva). 
Biology: Wilson, 1953. Ent. Soc. Amer. Ann. 46: 486. 



362 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

creightoni (M. R. Smith). N. C, Ga., Tenn. 

dietrichi (M. R. Smith). N. C. 

Taxonomy: G. C. and J. Wheeler, 1954, Psyche 61: 143 (larva). 
Biology: Wilson, 1953, loc. cit. supra, p. 487. 

filirrhina W. L. Brown. Mo. 

filitalpa W. L. Brown. Ind. (Brown Co. State Park). 

laevinasis (M. R. Smith). N. C, Ala. 

Taxonomy: Brown, 1964, loc. cit. supra, p. 199 {pilinasis-laevinasis com- 
plex). 

missouriensis (M. R. Smith). Iowa, 111., N. C. 

Taxonomy: G. C. and J. Wheeler, 1954. Psyche 61: 143 (larva). 
Biology: Wilson, 1953, loc. cit. supra, pp. 485-486. 

ohioensis (Kennedy and Schramm). Ga., Ind., N. J. 

ornata (Mayr). Okla., Mich., Mo., Tenn., S. C, La., Ark., Tex., Del. 

pilinasis (Forel). Mo., Ark., La., Tenn., N. C. 

Smithistruma (Smithistruma) brevisetosa (M. R. Smith) (syn. by Brown, 

1964, loc. cit. supra, p. 197). 
Taxonomy: G. C. and J. Wheeler, 1954. Psyche 61: 143 (larva). — Brown, 

1964, loc. cit. supra, p. 199 (pilinasis-laevinasis complex). 
Biology: Wilson, 1953, loc. cit. sup^'a, p. 485. 

pulchella (Emery). Remove Iowa from range of states. 

reflexa (L. G. Jr., and R. G. Wesson). Md. 

rostrata (Emery). 

Taxonomy: G. C. and J. Wheeler, 1954. Psyche 61: 144 (larva). 
Biology: Wilson, 1953, loc. cit. supra, p. 485. 

lalpa (Weber). 111. (Red Hills State Park). 

Biology: Wilson, 1953, loc. cit. supra, p. 486. 

Genus TRICHOSCAPA Emery, p. 829 

Biology: Brown and Wilson, 1960 (1959). Quart. Rev. Biol. 34: 283. 

tnembranifera (Emery). Calif, (near Stockton). Apparently introduced. 
Taxonomy: G. C. and J. Wheeler, 1954. Psyche 61: 145 (larva). 
Biology: Weber, 1952. Amer. Mus. Novitates No. 1554: 7. — Wilson, 

1953, loc. cit, supra, pp. 483-485. — Brown, 1954. Harvard Univ., 

Mus. Comp. Zool. Bui. 112: 6. 

Genus QUADRISTRUMA Brown, p. 829 

Biology: Brown, 1954, loc. cit. supra, p. 6. — Brown and Wilson, 1960 
(1959), loc. cit. supra, p. 281. 

emmae (Emery). 

Biology: Kempf, 1961. Studia Ent. 4: 516. 

*Genus EURHOPALOTHRIX Brown and Kempf 

Eurhopalothrix Brown and Kempf, 1960. Studia Ent., Petropolis (n.s.), 
3: 202-203. 

Type: Rhopalothrix Bolaui Mayr. Desig. by Brown and 
Kempf, 1961. 
Taxonomy: Brown and Kempf, 1961 (1960): Psyche 67: 44. 

Place after genus Quadristruma Brown, p. 829. 

*floridana Brown and Kempf. Fla. (Highlands Hammock, Highlands 
County). 

Eurhopalothrix floridana Brown and Kempf, 1960. Studia Ent., Petropolis 

(n.s.) 3: 207-208, fig. 34 (in key, p. 246). 5. 



FORMICIDAE 363 

CYPHOMYRMEX subgenus CYPHOMYRMEX Mayr, pp. 829-830 

Biology: Weber, 1957. Ecology 38: 480-494, figs. 1-5 (cultivation of 
fungus). 

rimosus (Spinola). Miss, (near Gulfport). 

Taxonomy: Weber, 1958. Ent. Soc. Wash. Proc. 60: 259-260.— G. C. and 
J. Wheeler, 1960. Ent. Soc. Wash. Proc. 62: 28 (larva). 

CYPHOMYRMEX subgenus MYCETOSORITIS W. M. Wheeler, p. 830 

hartmani (W. M, Wheeler), corrected spelling of hartmanni (!) of catalog, 
p. 830. 

Genus TRACHYMYRMEX Forel, pp. 830-831 

Biology: Weber, 1960. Ent. News 71: 1-6 (comparison of worker and 
female behavior) . 

septentrionalis (McCook), changed status. 

Trachymyrinex septentrionalis obscurior (W. M. Wheeler) (syn, by 

Weber, 1958. Ent. News 69: 52). 
Trachymyrmex septentrionalis seminole (W. M. Wheeler) (syn. by Weber, 

1958, loc. cit. supra, p. 53). 

septentrionalis obscurior (W.M.Wheeler). Syn. of septentrionalis (McCook). 
septentrionalis seminole (W. M. Wheeler). Syn. of septentrionalis (McCook). 
septentrionalis septentrionalis (McCook). Raised to sp. rank. 

Genus ATTA Fabricius, pp. 832-833 

Revision: Borgmeier, 1959. Studia Ent. 2 (fasc. 1-4): 321-390, figs. 

1-29. 

Biology: Weber, 1962. Natural History (N. Y.) 61: 45-51, illus.— Smith, 

1963. Ent. Soc. Wash. Proc. 65: 299-302. 

mexicana (F. Smith). Extreme south. Ariz.; Mexico, Honduras, Guatemala, 

El Salvador. 
Taxonomy: Borgmeier, 1959, Studia Ent. 2 (fasc. 1-4): 351-353, figs. 

16, 28 (5, 9, $). 
Biology: Smith, 1963, loc. cit. supra, pp. 299-302. 

texana (Buckley). Thirteen parishes in western La., much of the eastern half 
of Tex.; States of Tamaulipas and Vera Cruz, Mexico. 
Taxonomy: Borgmeier, 1959. Studia Ent. 2 (fasc. 1-4) : 356-358, fig. 29 

Biology: Bennett, 1958. U. S. Dept. of Agr., For. Pest Leaflet 23, pp. 
1-4, figs. 1-4. — Warter, Moser and Blum, 1962. La. Acad. Sci. 
Proc. 25: 42-46 (correlation between night hawk feeding and 
nuptial flights of A. texana and Iridomyrmex pniinosus (Rog.)). 
— Moser, 1962. Forests and People 12: 40-41, illus. (laboratory 
and field studies of nesting habits). — Moser, 1963. Ent. Soc. Amer. 
Ann. 56: 286-291, figs. 1-6, table 1 (contents and structure of nest 
in summer). — Smith, 1963, loc. cit. supra, pp. 299-302. 

Physiology: Moser and Blum, 1963. Science 140: 1228 (trail marking 
substance, source and potency). 

Subfamily DOLICHODERINAE 

Morphology: Pavan, 1955. Riv. Sci. Nat., "Natura" 46: 135-145, figs. 
1-6, table 1. 

Physiology: Pavan and Trave, 1958. Insectes Sociaux 5: 299-308 (venom 
of Tapinoma nigerrimum Nyl.). — Torossian, 1959. Ibidem, 6: 369- 
374, pi. 1 (tropholactic and proctodeal exchange in Dolichoderus 
guadripunctatus (L.)). — Wilson and Pavan, 1960 (1959). Psyche 
66: 70-76, tables 1-2 (glandular sources of chemical releasers of 
social behavior) . 



364 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

DOLICHODERUS subgenus HYPOCLINEA Mayr, pp. 833-835 

Taxonomy: G. C. and J. Wheeler, 3963, loc. cit. supra, pp. 149-154, figs., 
maps 24, 25. 

Biology: Kannowski, 1959. Insectes Sociaux 6: 144-145 (flights com- 
parisons of species). — Torossian, 1960. Insectes Sociaux 7: 383- 
393, figs. 1-3 (biology of Dolichoderus quadrip2inctatus (L.)). 

Morphology: Eisner, 1957. Harvard Univ., Mus. Compar. Zool. Bui. 116: 
453-454, pi. 2, fig. 7 (proventriculus). 

tuariae Forel. 

Biology: Talbot, 1956. Psyche 63: 134-139 (flight activities). — Kan- 
nowski, 1959, loc. cit. sup7-a, 129-133. 

plagiatus (Mayr). 

Biology: Kannowski, 1959, loc. cit. supra, p. 133. — G. C. and J. Wheeler, 
19G3, loc. cit. supra, pp. 149-151, map. 

pustulatus Mayr. 

Biology: Kannowski, 1959, loc. cit. supra, pp. 133-134 (155, pleometrosis) . 

taschenbergi (Mayr). 

Biology: Kannowski, 1959. Ent. Soc. Amer. Ann. 52: 755-760, figs. 1-3 
(flight activities). — G. C. and J. Wheeler, 1963, loc. cit. supra., pp. 
151-154, fig., map. 

Genus LIOMETOPUM Mayr, p. 835 

Morphology: Eisner, 1957, loc. cit. supra, p. 455, pi. 5, fig. 22 (pro- 
ventriculus) . 

apiculatum Mayr. 

Biology: Gregg, 1963. Univ. Colo. Studies, Series in Biol. No. 11, pp. 

1-6, figs. 1, 2. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 438-441, map. 

occidentale luctuosum W. I\I. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 441-443, map. 

Genus IRIDOMYRMEX Mayr, pp. 835-836 

Morphology: Eisner, 1957, loc. cit. supra, pp. 459-460, pi. 7, fig. 32; pi. 
8, fig. 34 (proventriculus). 

humilis (Mayr). 

Biology: Flanders, 1943. Calif. Citrograph 28: 117, 128, 137, illus. 
(Argentine ant versus parasites of black scale). — McCluskey, 
1958. Science 128: 536-537, fig. 1 (daily rhythm in male Argentine 
ant). 

Morphology: Pavan, 1955. Soc. Ital. di Sci. Nat. Atti 94: 379-477, figs. 
1-32; pis., tables 1-8, illus. — Nachtwey, 1963. Insectes Sociaux 
10: 50-53, figs. 3, 4; photograph 8. 

Physiology: Pavan, 1950 (1948). 8th Internatl. Cong. Ent. Proc, Stock- 
holm, pp. 863-865, illus.— Weber, 1961. Ent. Soc. Wash. Proc. 
63: 218 (use of poison). — Torossian, 1961. Insectes Sociaux 8: 
189-191. 

pruinosus analis (Andre). Calif, to Tex., Okla., and Kans., north to Idaho, 
and N. Dak. 
Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 154-155, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 436-438, map. 

pruinosus pruinosus (Roger). N. Y. to Fla. west to Wise, Tex., and N. Mex. 
Biology: Warter, Moser and Blum, 1962. La. Acad. Sci. Proc. 25: 42-46 

(correlation between night hawk feeding and nuptial flights of 

pruinosus pruinosus.) 
Physiology: Blum, Warter, Monroe and Chidester, 1963. Jour. Ins. 

Physiol. 9: 881-885, fig. 1 (chemical releasers of social behavior- 

methyl-n-amyl ketone). 



FORMICIDAE 365 

Genus FORELIUS Emery, p. 836 

foetidus foetidus (Buckley). Kans., Okla. and Tex. to Colo, and Calif., south 
to Mexico. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 434-436, map. 

DORYMYRMEX subgenus CONOMYRMA Forel, p. 837 

Morphology: Eisner, 1957, loc. cit. supra, pp. 463-464, pi. 7, fig. 33; pi. 
8, fig. 37; pi. 10, fig. 46 (proventriculus). 

bicolor W. M. Wheeler, changed status. 

Doryinyrniex (Conoinyriua) pyramicus bicolor W. M. Wheeler. Raised 
to sp. rank (teste Cole, 1958 (1957). N. Y. Ent. Soc. Jour. 65: 
130). 
Taxonomy: Wilson, 1958 (1957). Psyche 64: 76. 

pyramicus bicolor W. M. Wheeler. Raised to sp. rank. 

pyramicus pyramicus (Roger). Entire U. S. with the possible exception of 

New England States north of Conn., and Wash. 
Taxonomy: Wilson, 1958 (1957). Psyche 64: 76. 
Biology: Wilson 1958 (1957). Psyche 64: 48.— G. C. and J. Wheeler, 

1963, loc. cit. supra, pp. 155-157, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 432-434, map. 

Genus TAPINOMA Foerster, pp. 837-833 

Morphology: Eisner, 1957, loc. cit. supra, pp. 456-457, pi. 6, fig. 26 

(proventriculus) . 
Physiology: Toropsian, 1960. Insectes Sociaux 7: 171-174 (tropholactic 

and proctodeal in Tapinoma erraticum (Latreille) ). — Weber, 1961. 

Ent. Soc. Wash. Proc. 63: 217-218 (use of poison). 

melanocephalum (Fabricius). 

Biology: King, 1948. Iowa Acad. Sci. Proc. 55: 395.— Kempf, 1961. 

Studia Ent. 4: 520.— Brown, 1964. Ent. News 75: 14-15. 
Physiology: Weber, 1961, loc. cit. supra, p. 218 (employment of venom by 

ant). 

sessile (Say). 

Biology: Kannowski, 1959. Insectes Sociaux 6: 126-129 (155, pleo- 

metrosis). — G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 157-160, 

map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 443-447, map. 

Subfamily FORMICINAE 

Biology: Pontin, 1960. Insectes Sociaux 7: 227-230 (experiments in 
colony founding in Lasius niger (L.) and Lasius flamis (F.)). — 
Kannowski, 1963. Pavia Univ., Atti IV Cong. U.I.E.I.S., v. 12, 
pp. 74-102, tables 1-7 (flight activities). 

Morphology: Eisner and Wilson, 1952. Psyche 59: 47-60, fig. 1, pis. 
5, A, B, C, D; pi. 6, A, B, (proventriculus). 

*PLAGiOLEPIS subgenus PLAGiOLEPIS Mayr 

Plagiolepis subg. Plagiolepis Mayr, 1861. Europ. Formicid., p. 42. 
Type: Formica pygmaea Latreille. By monotypy. 

Insert before Brachymyrmex Mayr, p. 838. 

*alluaudi Emery. Calif. (Catalina Isl.). Introduced. An Old World species 
that has been widely disseminated by commerce. 

Taxonomy: M. R. Smith, 1957. N. Y. Ent. Soc. Jour. 65: 195-198 (also 
biology). 

Biology: Taylor and Wilson, 1961. Psyche 68: 143. 

BRACHYMYRMEX subgenus BRACHYMYRMEX Mayr, pp. 838-839 

Biology: Arnaud and Quate. 1951. Pan-Pacific Ent. 27: 171 (note on 
swarming) . 



366 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

depilis Emery. N. S. and B. C. south to Fla. and Calif. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 161-163, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 447-449, map. 

heeri var. obscurior Forel. Raised to sp. rank. 

obscurior Forel, changed status. 

Brachyntyrmex ( Brachyinyrmex) heeri var. obscurior Forel. Raised to 

sp. rank (teste Brown, 1957. Harvard Univ., Mus. Compar. Zool. 

Bui. 116: 237). 
Morphology: Eisner, 1957. Ibidem 116(8), pi. 20, fig. 84 (cuticular 

framework of proventriculus of worker) . 

CAMPONOTUS subgenus CAMPONOTUS Mayr, pp. 839-842 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 163-170, figs., 

maps 30-32. 
Biology: Riordan, 1960. Insectes Sociaux 7: 353-355 (location of nests 

of carpenter ants, Camponotus spp. by a radioactive isotope). 
Morphology: Eisner, 1957, loc. cit. supra, pp. 470-473, pi. 15, fig. 67; pi. 

16, fig. 68; pi. 17, figs. 70-74 (proventriculus). 

americanus Mayr. 

Morphology: Eisner and Wilson, 1952. Psyche 59: 47-60, fig. 1; pi. 5, 
A, B, C, D; pi. 6, A, B, C (proventriculus). 

herculeanus herculeanus (Linnaeus). 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 164-165, map. 
Ecology: Brown, 1955. Ent. News 66: 47-48. — Gregg, 1963, loc. cit. supra, 

pp. 656-660, map. 
Physiology: Ayre 1964 (1963). Ent. Exp. and Appl. 6: 165-170. 

herculeanus niodoc W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 665-669, map. 

laevigatus (F. Smith). 

Ecology: Gregg, 1963, loc. cit. supra, pp. 660-663, map. 

noveboracensis (Fitch). 

Biology: Kannowski, 1959. Insectes Sociaux 6: 134-135. — G. C. and J. 

Wheeler, 1963, loc. cit. supra, pp. 165-168, figs., map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 663-665, map. 

pennsylvanicus ( DeGeer ) . 

Morphology: Forbes and McFarlane, 1961. N. Y. Ent. Soc. Jour. 69: 

92-103, figs. 1-5 (comparative anatomy of digestive glands of 

female and male). — Keister, 1963. Ent. Soc. Amer. Ann. 56: 336- 

340, figs. 1-3 (anatomy of tracheal system). 
Biology: Van Pelt, 1958. Tenn. Acad. Sci. Jour. 33: 120-122, fig. 1 

(parasitism by a fungus). — G. C. and J. Wheeler, 1963, loc. cit. 

supra, pp. 168-170, map. 

CAMPONOTUS subgenus TANAEMYRMEX Ashmead, p. 843 

sansabeanus sansabeanus (Buckley). 

Ecology: Gregg, 1963, loc. cit. supra, pp. 669-671, map. 

sansabeanus torrefactus W. M. Wheeler. Colo., Nev. 

vicinus Mayr. Alta., N. Dak. and Okla. to Calif, and B. C. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 170-171, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 671-675, map. 

CAMPONOTUS subgenus MYRMOTHRIX Forel, pp. 843-844 

abdominalis floridanus (Buckley). Dauphine and Horn Islands in the Gulf 
of Mexico. 

CAMPONOTUS subgenus MYRMENTOMA Forel, pp. 844-845 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 171-174, figs., 
maps 34, 35. 



FORMICIDAE 367 

caryae discolor (Buckley). N. Dak. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 172-173, map. 

nearcticus Emery. 

Biology: Kannowski, 1959. Insectes Sociaux 6: 134. — G. C. and J. 

Wheeler, 1963, loc. cit, supra, pp. 173-174, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 675, 677, map. 

rasilis W. M. Wheeler. N. C., Calif. (Imperial County). 
Ecology: Gregg, 1963, loc. cit. supra, pp. 677-679, map. 

subbarbatus Emery. N. C, Md., N. Y. 

CAMPONOTUS subgenus COLOBOPSIS Mayr, p. 846 
pylartes fraxinicola M. R. Smith. N. C. 
mississippiensis M. R. Smith. N. C, Md. (Plummers Island). 

PARATRECHINA subgenus PARATRECHINA Motschoulsky, pp. 847-848 

iongicornis (Latreille). 

Biology: Brown, 1964. Ent. News 75: 14-15. 

Economics: Fox and Garcia-Moll, 1961. Jour. Econ. Ent. 54: 1065-1066, 
figs. 1-2. 

PARATRECHINA subgenus NYLANDERIA Emery, pp. 848-849 

parvula (Mayr). Ariz., N. Dak., ? Utah. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 175-176, map. 

Genus PRENOLEPIS Mayr, pp. 849-850 

imparis coloradensis W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 496-498, map. 

LASIUS subgenus LASIUS Fabricius, pp. 850-851 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 177-188, maps 
37-40. — Gregg, 1963, loc. cit. supra, pp. 449-453. 

Biology: Kannowski, 1959. Insectes Sociaux 6: 145-146 (flight com- 
parisons) . 

alienus (Foerster). 

Biology: Kannowski, 1959. Insects Sociaux 6: 135-136. — Judd and 
Benjamin, 1958. Canad. Ent. 90: 419.— G. C. and J. Wheeler, 
1963, loc. cit. supra, pp. 178-179, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 454-457, map. 

crypticus Wilson. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 179-182, map. 

neoniger Emery. 

Biology: Ayre, 1962. Canad. Jour. Zool. 40: 157-164, figs. 1-5 (para- 
sitism). — G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 182-185, 
fig., map. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 457-461, map. 

sitkaensis Pergande. 

Biology: Medler, 1958. Ent. Soc. Wash. Proc. 60: 258 (emergence of 

males, winged females for flights). — G. C. and J. Wheeler, 1963, 

loc. cit. supra, pp. 185-188, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 461-465, map. 

LASIUS subgenus CAUTOLASIUS Wilson, Sup. 1, pp. 148-149 

Taxonomy: Gregg, 1963, loc. cit. supra, pp. 449-453. 

flavus (Fabricius). N. Dak. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 189-191, map. — 
Waloff, 1957. Insectes Sociaux 4: 391-408, tables 1-11 (effect of 
number of queens on their survival and the development of first 
brood) . 



368 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

Ecology: Gregg, 1963, loc. cit. supra, pp. 465-469, map. 

nearcticus W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 469-471, map. 

LASIUS subgenus CHTHONOLASIUS Ruzsky, pp. 851-852 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 191-194, maps 

42, 43. — Gregg, 1963, loc. cit. supra, pp. 449-453. 
Biology: Kannowski 1959. Insectes Sociaux 6: 145-146. 

*atopus Cole. Calif. (3 miles south of Leggett, Mendocino County). 

Lasius (Chthonolasius) atnpus Cole, 1958. Tenn. Acad. Sci. Jour. 33: 
75-77, fig. 1, A, B. 5 . 

hiunilis W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 473-475, map. 

minutus Emery. 

Biology: Kannowski, 1959. Insectes Sociaux 6: 136-138 (151-153, colony 

founding; possible parasitism on Lasius alienus (Foerster) and 

Lasius sitkaensis (Pergande). 
Physiology: Kannowski, 1959. Ecology 40: 162-165 (radioactive 

phosphorus in study of colony distribution). 

speculiventris Emery. Host ant, Lasius minutus Emery. 

Biology: Kannowski, 1959, loc. cit. supra, pp. 138-141 (153-154, para- 
sitism on minutus and possibly other Lasius spp). 

subumbratus Viereck. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 191-192, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 475-478 map. 

umbratus (Nylander). N. Dak. 

Biology: Kannowski, 1959, loc. cit. supra, p. 141 (154-155, possible para- 
sitism of other Lasius spp.). — G. C. and J. Wheeler, 1963, loc. cit. 
supra, pp. 192-194, map. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 478-481, map. 

Genus ACANTHOMYOPS Mayr, Sup. 1, pp. 150-151 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 194-204, figs., 
maps 45-49. — Gregg, 1963, loc. cit. supra, pp. 449-453. 

claviger claviger (Roger). 

Biology: Talbot, 1963. Ecology 44: 549-557, tables 1-3, (local distribu- 
tion, flight activities). — G. C. and J. Wheeler, 1963, loc. cit. supra, 
pp. 195-197, map. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 481-482, map. 

Physiology: Chadha, Eisner, Monro and Meinwald, 1962. Jour. Ins. 
Physiol. 8: 175-179, figs. 1-4 (citronellal and citral in mandibular 
gland secretions). 

claviger coloradensis (W. M. Wheeler). 

Ecology: Gregg, 1963, loc. cit. supra, pp. 481, 483-484, map. 

inter jectus interjectus (Mayr). 

Biology: Talbot, 1963. Ecology 44: 549-557, tables 1-3 (local distribution, 
flight activities). — G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 
197-200, figs,, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 484-486, map. 

latipes (Walsh). 

Biology: Talbot, 1963. Ecology 44: 549-557, tables 1-3 (local distribu- 
tion flight activities). — G. C. and J. Wheeler, 1963, loc. cit. supra, 
pp. 200-202, fig., map. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 486-488, map. 

murphy i ( Forel ) . 

Biology: Talbot, 1963. Ecology 44: 549-557, tables 1-3 (local distribu- 
tion, flight activities). 
Ecology: Gregg, 1963, loc. cit. supra, pp. 489-490, map. 



FORMicroAE 369 

occidentalis (W. M. Wheeler). N. Dak., ? Wis. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 490-493, map. 

parvulus (M. R. Smith). 111., N. Dak. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 202-203, map. 

pogonogynus (Buren). Idaho. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 493-494, map. 

subglaber (Emery). Wis., Tenn. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 203-204, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 493. 495-496, map. 

Genus MYRMECOCYSTUS Wesmael, pp. 854-857 

comatus W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 643-645, map. 

lugubris W. M. Wheeler. Nev., Ariz. 

mendax W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 645-648, map. 

mexicanus hortideonim McCook. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 648-651, map. 

navajo W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit. stipra, pp. 651-653, map. 

pyramicus M. R. Smith. Calif., Idaho. 

Taxonomy: Cole, 1958 (1957). N. Y. Ent. Sec. Jour. 65: 129-130 (9, $). 

semirufus Emery. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 653-655, map. 

FORMICA subgenus PROFORMICA Ruzsky, p. 857 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 206-214, figs., 
maps 50-53. 

bradleyi W. M. Wheeler. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 206-209, fig., 

map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 602-606, map. 

lasioides Emery. Coast to coast in south. Canada and north. U. S. with ex- 
tensions in Rocky Mts. to N. Mex. and Appalachians. 

Biology-: Kincaid, 1963. Amer. Micros. Soc. Trans. 82: 101-105, figs. 1-5, 
(as pollinators of plants). — G. C. and J. Wheeler, 1963, loc. cit. 
supra, pp. 209-211, map. 

Ecology: Gregg, 1963, loc. cit. siipra, pp. 498-500, 502, map. 

Ilmata W. M. Wheeler. Utah. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 211-212, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 502, 504 map. 

manni W. M. Wheeler. 

Formica (Formica) manni W. M. Wheeler. Transf. by Wilson and Brown, 

1955. Psyche 62: 126. 
Taxonomy: Wilson and Brown, 1955, loc. cit. supra, pp. 127-128. 

neogagatcs Emery. Coast to coast in south. Canada and north. U. S. with 
extensions in the Appalachians to N. C. and in the Rocky Mts. 
to N. Mex. Slave of Formica wheeleri Creighton. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 212-214, fig., map. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 503-505, map. 

obtusopilosa Emery. 

Formica (Formica) obtusopilosa Emery. Transf. by Wilson and Brown, 

1955. Psyche 62: 128. 
Taxonomy: Wilson and Brown, 1955, loc. cit. supra, p. 128. 
Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 217-219, fg., map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 608-611 map. 



370 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

perpilosa W. M. Wheeler. Wyo. 

Formica (Formica) perpilosa W. M. Wheeler. Transf. by Wilson and 

Brown, 1955. Psyche 62: 128. 
Taxonomy: Wilson and Brown, 1955, loc. cit. supra, p. 128. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 611-613, map. 

FORMICA subgenus NEOFORMICA W. M. Wheeler, pp. 858-860 

Biology: Kannowski, 1959. Insectes Sociaux 6: 147 (flight comparisons). 

pallidefulva pallidefulva Latreille. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 625-628, map. 

pallidefulva nitidiventris Emery. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 628-630, map. 

schaufussi dolosa W. M. Wheeler. Va. to Fla. west to Iowa, Mo., Tex., and 
Colo. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 630-632. 

FORMICA subgenus FORMICA Linnaeus, pp. 860-873 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 214-228, figs., 
maps 54-60 (sanguinea group) ; pp. 229-252, figs., maps 61-71 
(rufa group) ; pp. 252-253, map 72 (microgyna group) ; pp. 254- 
258, figs., map. 73 (exsecta group) ; pp. 258-274, figs., maps 74- 
79 {fusca group). 

Biology: Gosswald, 1957. Insectes Sociaux 4: 335-348, tables 1-9 (caste 
determination in the genus ForTuica). — Kannowski, 1959. Insectes 
Sociaux 6: 147. 

Morphology: Eisner, 1957, loc. cit. supra, pp. 473-474, pi. 25, figs. 100-103 
(proventriculus) . 

Physiology: Osman and Kloft, 1961. Insectes Sociaux 8: 383-395 
(venom). 

altipetens W. M. Wheeler. N. Dak., Mont., and B. C. to Mex. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 260-261, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 507-509, map. 

aterrima Cresson. 

Taxonomy: Gregg, 1963, loc. cit. supra, pp. 544, 546. 

ciliata Mayr. N. Dak. Temporary parasite of Formica cinerea lepida W. M. 
Wheeler. 
Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 233-234, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 545-547, map. 

cinerea var. canadensis Santschi. Syn. of cinerea lepida W. M. Wheeler. 

cinerea lepida W. M. Wheeler. Sask., Utah, Tex. Host of temporary parasites, 
Formica ciliata Mayr, F. dakotensis Emery and F. criniventris 
W. M. Wheeler. 

Formica (Formica) cinerea var. canadensis Santschi (syn. by M. R. 
Smith, 1960. Ent. Soc. Wash. Proc. 62: 251-252). 

Biology: G. C. and J. Wheleer, 1963, loc. cit. supra, pp. 261-263, map. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 509-512, 514, map. 

cinerea montana Emery. Ohio to Dakotas, Nebr., Colo., and N. Mex. 

Biology: G. C. and J. Wheeler, 1963, loc cit. supra, pp. 263-264, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 513-514, map. 

comata W. M. Wheeler. N. Dak. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 547-548, 550, map. 

criniventris W. M. Wheeler. Probably a temporary parasite of F. cinerea 
lepida W. M. Wheeler. 
Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 234-236, fig., map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 549-551, map. 

cariosa Creighton. 

Taxonomy: Wilson and Brown. 1955. Psyche 62: 123-125, fig. 2. 



FORMICIDAE 371 

dakotensis Emery. Alaska, Nev., N. Dak., Ohio. 

Formica {Formica) dakotensis var. montigena W. M. Wheeler (syn. by 

Brown, 1957. Ent. News 68: 167). 
Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 236-238, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 551-554, map. 

dakotensis var. montigena W. M. Wheeler. Syn of dakotensis Emery. 

densiventris Viereck. Utah. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 589-593, map. 

exsectoides Forel. Miss. (Holly Springs). 

Ecology: Gregg, 1963, loc. cit. supra, pp. 597-599, map. 
Physiology: Eisner and Happ, 1964. Sci. 134: 329-331, figs. 1-4 (haemor- 
rhage of a coccinellid beetle and its repellent effect on ants). 

fossaceps Buren. N. Dak. 

Biology: King, 1949. Iowa Acad. Sci. Proc. 56: 367-370 (mixed colony 
with F. obscuriventris clivia Creighton). — King, 1951. Op. cit., 
pp. 487-489.— King, 1952. Op. cit. pp. 469-474, figs. 1, 2 (macrop- 
seudogynes, or ? pterergates). — King and Sallee, 1957. Op. cit. 
64: 667-669, table 1 (mixed colony with F. obscuriventris clivia). 
—King and Sallee, 1959. Op. cit. 66: 472-473 (slave of F. rubi- 
cunda Emery).— King and Sallee, 1962. Op. cit. 69: 531-539, figs. 
1-12, table 1 (mixed colony with F. obscuriventris clivia). — G. C. 
and J. Wheeler, 1963, loc. cit. supra, pp. 238-239, map. 

fusca Linnaeus. Holarctic. Labrador and Alaska south through Canada. 
Occurs in every state in U. S., but absent from certain unfavorable 
habitats, such as deserts. 

Biology: King, 1949. Iowa Acad. Sci. Proc. 56: 367-370 (in mixed colony 
with Formica reflexa Buren). — King, 1951. Op. cit. pp. 487-489 
(ibidem). — Kannowski, 1959. Insectes Sociaux 6: 141-142. — 
Wallis, 1960. Op. cit. 7: 187-190. figs. 1-8, tables 1-2 (spinning 
movements of larvae). — G. C. and J. Wheeler, 1963, loc. cit. supra, 
pp. 264-267, map. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 514-525, map. 

hewitti W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 526-527, map. 

Integra haemorrhoidalis Emery. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 854-857, map. 

integroides coloradensis W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 557-560, map. 

integroides planipilis Creighton. Utah, Colo., Wyo., N. Dak., Wash. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 560-561, map. 

integroides propinqua W. M. Wheeler. Colo., Utah. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 562-564, map. 

laeviceps Creighton. N. Dak., Colo. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 239-240, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 564-565, map. 

manni W. M. Wheeler. Transf. to Formica subg. Proformica Ruzsky, p. 857. 

marcida W. M. Wheeler. N. Dak., S. Dak. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 267-268, map. 

mucescens W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 566-567, map. 

microgyna W. M. Wheeler. Wyo., Utah, N. Mex. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 584-587, map. 

neoclara Emery. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 268-271, fig., map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 526, 528-530, map. 



372 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

neorufibarbis gelida W. M. Wheeler. Calif, to Alaska to Sask. and south 
through Rocky Mts. to N. Mex. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 533-536, map. 

neorufibarbis neorufibarbis Emery. Nebr. 

Biology: Kannowski, 1959. Insectes Sociaux 6: 142. — G. C. and J. 

Wheeler, 1963, loc. cit. supra, pp. 271-273, map. 
Ecology: Brown, 1955. Ent. News 66: 47-50. — Gregg, 1963, loc. cit. supra, 

pp. 530-533, map. 

obscuripes obscuripes Forel. South. Canada and north. U. S. from Ind. and 
Mich, to Calif, and B. C, also the Rocky Mt. states. 

Biology: King and Sallee, 1957. Iowa Acad. Sci. Proc. 63: 721-723, tables 
1-3.— Talbot, 1959. Amer. Midland Nat. 61: 124-128 (flight ac- 
tivities). — G. C. and J, Wheeler, 1963, loc. cit. supra, pp. 240-244, 
fig., map. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 566, 568-572, map. 

obscuripes ravida W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 579-581, map. 

obscuriventris clivia Creighton. Colo., N. Mex. 

Biology: King, 1949. Iowa Acad. Sci. Proc. 56: 367-370 (mixed colony 
with Formica fossaceps Buren). — King and Sallee, 1951. Op. cit. 
58: 487-489 (ibidem) .—King, 1955. Op. cit. 62: 509-513, figs. 1, 
2 (winged workers).— King and Sallee, 1959. Op. cit. 66: 472-473 
(slave of Formica rubicunda Emery). — King and Sallee, 1962. 
Op. cit. 69: 531-539, figs. 1-12, table 1 (mixed colonies with 
Formica fossaceps Buren). 

Ecology: Gregg, 1963, loc. cit. supra, pp. 574-577, map. 

obscuriventris obscuriventris Mayr. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 244-246, map. 
Ecology: Gregg, 1963, loc. cit. supra, 572-574, map. 

obtusopilosa Emery. Transf. to Formica subg. Proformica Ruzsky, p. 858. 

opaciventris Emery. N. Dak. 

Biology: Scherba, 1961. N. Y. Ent. Soc. Jour. 69: 71-87, tables 1-6 (re- 
production, nest structure). — Scherba, 1963. Op. cit. 71: 219-231, 
tables 1-6, figs. 1-3. — G. C. and J. Wheeler, 1963, loc. cit. supra, 
pp. 254-256, map. 

oreas comptula W. M. Wheeler. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 246-248, map. 

oreas oreas W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 577-579, map. 
oregonensis Cole. 

Biology: Kincaid, 1963. Amer. Micros. Soc. Trans. 82: 101-105, figs. 
1-5 (pollinates a plant). 

Taxonomy: Cole, 1956. Tenn. Acad. Sci. Jour. 31: 212-214. 

parcipappa Cole. 

Taxonomy: Wilson and Brown, 1955. Psyche 62: 125. 

perpilosa W. M. Wheeler. Transf. to Formica subg. Proformica Ruzsky, p. 858. 
postoculata Kennedy and Dennis. Pa. (Mt. Morris). 

pruinosa W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 537-539, map. 

puberula Emery. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 220-221, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 613-616 map. 

rasilis W. M. Wheeler. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 587-589, map. 

reflexa Buren. N. Dak. 

Biology: King, 1949. Iowa Acad. Sci. Proc. 56: 367-379 (mixed colony 



FORMICIDAE 373 

with fusco L.).— King, 1951. Op. cit. 487-489 {ibidem). — G. C. 
and J. Wlieeler, 1963, loc. cit. supra, pp. 249-250, map. 

rubicunda Emery. 

Biology: King and Sallee, 1959. Iowa Acad. Sci. Proc. 66: 472-473 
(slaves — Formica fossaceps Buren and F. obscuriventris clivia 
Creighton). — G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 221- 
223, map. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 616-618, map. 

rufibarbis gnava Buckley. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 539-541, map. 

spatulata Buren. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, p. 253, map. 

subintcgra Emery. 

Biology: Brown, 1959 (1958). Psyche 65: 39-40.— G. C. and J. Wheeler, 
1963, loc. cit. supra, pp. 225-227, map. 

sublucida W. M. Wheeler. New England states to S. Dak., Colo. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 622-623, map. 

subnitens Creighton. N. Dak., Ol-eg., Colo. 

Taxonomy: Miller, 1957. Insectes Sociaux 4: 253-257, figs. 1-4 (2, ?, $). 
Biology: Ayre, 1957. Insectes Sociaux 4: 173-176 (ecological notes). — 

Ayre, 1958. Op. cit. 5: 1-7.— Ayre, 1958,. Op. cit. 5: 147-157, fig. 

1 (meterological factors affecting foraging). — Ayre, 1959. Op. 

cit. 6: 105-114, figs. 1-3 (food habits). 
Ecology: Gregg, 1963, loc. cit. supra, pp. 581-584, map. — G. C. and J. 

Wheeler, 1963, loc. cit. supra, pp. 250-252, map. 

subnuda Emery. 

Biology: G. C. and J. Wheeler 1963, loc. cit. supra, pp. 223-225, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 618-621, map. 

subpolita ficticia W. M. Wheeler. 

Ecology: Gregg 1963, loc. cit. supra, pp. 541-542, map. 

ulkei Emery. Iowa, Ind., Wyo. 

Biology: Scherba, 1958. Insectes Sociaux 5: 201-213, figs. 1-5 (reproduc- 
tion, mounds).— Kannowski, 1959. Op. cit. 6: 142-143.— Talbot, 
1959. Amer. Midland Nat. 61: 128-132 (flight activities).— 
Scherba, 1959, Op. cit. 61: 499-508, figs. 1-2 (moisture regulations 
in mounds).— Talbot, 1961. Ecology 42: 202-205 (mounds).— 
Scherba, 1962. Amer. Midland Nat. 67: 373-385, figs. 1-4 (mound 
temperatures). — G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 
256-258, fig., map. 

wheeleri Creighton. N. Dak. Slave: Formica neogagates Emery. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 227-228, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 623-625, map. 

whymperi alpina W. M. Wheeler. 

Taxonomy and Ecology: Gregg, 1963, loc. cit. supra, pp. 593-596 (9). 

Genus POLYERGUS Lafreille, pp. 873-875 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 274-277, figs., 
maps 80, 81. 

Biology: Dobrzanska and Dobrzanska, 1960. Insectes Sociaux 7: 1-8 (new 
observations on Polyergus rufesccns Latr.). 

Morphology: Forbes and Brassel, 1962. N. Y. Ent. Soc. Jour. 70: 79-87, 
pi. 1, figs. 1-5; pi. 2, figs. 8-19 (male genitalia and terminal seg- 
ments of Polyergus riifescens bicolor Wasmann, P. r. breviceps 
Emery, P. r. umbratus W. M. Wheeler, and P. lucidus Mayr). 
lucidus longicornis M. R. Smith. Ga., N. C. 

lucidus lucidus Mayr. N. Dak., Ark.. Colo. 

Biology: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 274-275, map. 
Ecology: Gregg, 1963, loc. cit. supra, pp. 633-635, map. 



374 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

rufescens bicolor Wasmann. Colo. 

Taxonomy: G- C. and J. Wheeler, 1963, loc. cit. supra, pp. 275-277, map 

(as rufescens Latr.). 
Ecology: Gregg, 1963, loc. cit. supra, pp. 635-638, map. 

rufescens breviceps Emery. 

Taxonomy: G. C. and J. Wheeler, 1963, loc. cit. supra, pp. 275-277, map 

(as rufescens Latr.). 
Ecology: Gregg, 1963, loc. cit. supra, pp. 638-641, map. 

rufescens umbratus W. M. Wheeler. Colo. 

Ecology: Gregg, 1963, loc. cit. supra, pp. 641-643, map. 

Superfamily VESPOIDEA 
Family VESPIDAE 

Taxonomy: Bohart and Bechtel, 1957. Calif. Ins. Survey Bui. 4: 73-101, 
34 text figs., 7 maps, 2 pis. (social Vespidae of Calif.). — Richards, 
1962. A Revisional Study of the Masarid Wasps, 294 pp., 241 figs. 

Biology: Krombein, 1961. Wash. Acad. Sci. Jour. 51: 89-93, 6 figs, 
(symbiosis between saproglyphid mites and solitary vespid 
wasps). 

Richards (1962) proposed dividing the Vespidae into three families. 
In North America the Vespidae would include the social wasps (Vespinae, 
Polistinae and Polybiinae of the Catalog), and the solitary wasps would be 
assigned to the Eumenidae (Zethinae and Eumeninae of the Catalog) and 
the Masaridae (Masarinae and Euparagiinae of the Catalog). 

Genus VESPA Linnaeus, p. 875 
crabro germana Christ. W. Va., Ohio, Ind., Ala. 

Genus VESPULA Thomson, pp. 875-877 

Revision: Miller, 1961. Canad. Ent., Supp. 22, Vol. 93, 52 pp. 

VESPULA subgenus VESPULA Thomson, pp. 875-877 

acadica (Sladen), changed status. 

Vespula (Vespiila) rufa acadica (Sladen). Restored to sp. rank (teste 

Miller, 1961, loc. cit. supra, p. 15). 
Vespula (Vespula) rufa sladeni Bequaert (syn. by Miller, 1961, loc. cit. 

supra, p. 15). 

atropilosa (Sladen), changed status. 

Vespula (Vespula) rufa atropilosa (Sladen). Restored to sp. rank (teste 
Miller, 1961, loc. cit. supra, p. 17). 

consobrina (Saussure), changed status. 

Vespula (Vespula) rufa consobrina (Saussure). Restored to sp. rank 
(teste Bohart and Bechtel, 1957, loc. cit. supra, p. 79). 

intermedia (Buysson), changed status. 

Vespula (Vespula) rufa intermedia (Buysson). Restored to sp. rank 
(teste Miller, 1961, loc. cit. supra, p. 23). 

maculifrons (Buysson). 

Biology: Haviland, 1962. Ent. Soc. Wash. Proc. 64: 181-183 (colony 
size). 

pensylvanica (Saussure). 

Biology: Chapman, 1963. Ecology 44: 766-767 (predation on winged 
ants). 

rufa acadica (Sladen). Raised to sp. rank. 

rufa atropilosa (Sladen). Raised to sp. rank. 

rufa consobrina (Saussure). Raised to sp. rank. 



VESPIDAE 375 

rufa intermedia (Buysson). Raised to sp. rank. 

rufa sladeni Bequaert. Syn. of acadica (Sladen). 

rufa vidua (Saussure). Raised to sp. rank. 

vidua (Saussure), changed status. 

Vespula (Vespulu) rufa vidua (Saussure). Restored to sp. rank {teste 
Miller, 1961, loc. cit. supra, p. 26). 

VESPULA subgenus DOLICHOVESPULA Rohwer, p. 877 

adulterina adulterina (Buysson). Delete; the color variant in western North 
America thought to be typical adulterina from Asia is just a varia- 
tion of arctica Rohwer. 

adulterina arctica Rohwer. Raised to sp. rank. 

albida (Sladen), changed status. 

Vespula (Dolichovespula) norwegica albida (Sladen). Restored to sp. 
rank {teste Miller, 1961, loc, cit. supra, p. 28). 

arctica Rohwer, changed status. 

Vespula (Dolichovespula) adulterina arctica Rohwer. Restored to sp. 
rank {teste Miller, 1961, loc. cit. supra, p. 30). 

arenaria (Fabricius), changed status. 

Vespula (Dolichovespula) arenaria arenaria (Fabricius). Raised to sp. 

rank {teste Bohart and Bechtel, 1957, loc. cit. supra, p. 88). 
Vespula (Dolichovespula) arenaria fernaldi (Lewis) (syn, by Bohart 
and Bechtel, 1957, loc. cit. supra, p. 88). 

arenaria fernaldi (Lewis). Syn. of arenaria (Fabricius). 

norvegicoides (Sladen), changed status. 

Vespula (Dolichovespula) norvegicoides (Sladen). Restored to sp. rank 
{teste Miller, 1961, loc. cit. supra, p. 36). 

norwegica albida (Sladen). Raised to sp. rank. 

norwegica norvegicoides (Sladen). Raised to sp. rank. 

Genus POLISTES Latreille, pp. 878-879 

annularis (Linnaeus). 

Biology: Rabb, 1960. Ent. Soc. Amer. Ann. 53: 111-121.— Balduf, 1961. 
Ent. News 72: 259-260. 

exclamans exclamans Viereck. Md., N. J. 

Biology: Hodapp and Bickley, 1959. Ent. Soc. Wash. Proc. 61: 73.— 
Rabb, 1960, loc. cit. supra, pp. 111-121. 

fuscatus fuscatus (Fabricius). 

Biology: Rabb, 1960, loc. cit. supra, pp. 111-121. 

Genus PSEUDOMASARIS Ashmead, pp. 882-883 

Taxonomy: Richards, 1962. A Revisional Study of the Masarid Wasps, 
pp. 276-281, figs. 236-241 (revised key and places the subgenera 
Toryna Bradley and Holopticus Bradley in synonymy). 

Revision: Richards, 1963. Univ. Calif. Pubs. Ent. 27: 283-310, 3 pis., 
26 text figs. 

bariscipus Bradley. Syn. of basirufus Rohwer. 

basirufus Rohwer. Ariz., Calif. 

Pseudomasaris (Holopticus) maculifrons (Fox), of Catalog in part. Re- 
move zonalis basirufus Rohwer from syn. {teste Richards, 1962, 
loc. cit. supra, p. 281). 

Pseudomasaris (Holopticus) bariscipus Bradley (syn. by Richards, 1962, 
loc. cit. supra, p. 281). 

coquilletti Rohwer. Ariz. 

edwardsii (Cresson). Mexico (Baja California). 



376 HYMENOPTERA OF AMERICA NORTH OF MEXICO 

*macneilli Bohart. Calif., Utah. 

Pseudoinasaris macneilli Bohart, in Richards, 1963. Univ. Calif. Pubs. 
Ent. 27: 291. $, 5. 

*inaoswaini Bohart. Southern Calif. 

Pseudoinasaris macswaini Bohart, in Richards, 1963. Univ. Calif. Pubs. 
Ent. 27: 301. $, $. 

maculifrons (Fox). Utah, N. Mex.; Mexico (Sonora, Baja California). Re- 
move basirufus Rohwer, a valid sp., from syn. 

inarginalis (Cresson). Nev., Wash., B. C. 

Pseudoinasaris (Holopticus) rohweri Bradley (syn. by Richards, 1962, 
loc. cit. supra, p. 280). 
*inicheiicri Bohart. Calif. (Inyo Co.). 

Pseudoinasaris micheneri Bohart, in Richards, 1963. Univ. Calif. Pubs. 
Ent. 27: 298, figs. 1, 14, 21. $, 9. 

occidentalis (Cresson). Kans., N. Mex. 

rohweri Bradley. Syn. of maculifrons (Fox). 

texanus (Cresson). Ariz. 

vespoides (Cresson). Wash., Mont.; Mexico (Baja California). 

zonaiis (Cresson). Mont., Nebr. 

Pseudoinasaris (Preudomasaris) sonalis alhopictus Bohart (syn. by 
Richards, 1962, loc. cit. supra, p. 280). 

zonaiis alhopictus Bohart. Syn. of zonaiis (Cresson). 

Subfamily EUPARAGIINAE 

Taxonomy: Richards, 1962. A Revisional Study of the Masarid Wasps, 
pp. 49-51, figs. 47-54. 

Genus EUMENES Latreille, pp. 884-885 

aureus Isel