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Full text of "Ichthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in .."

Woods Hole Oceanographic Institution 
ATLAS - GAZETTEER COLLECTION 



NOAA Technical Memorandum NMFS 




Sr ATES O* 



SEPTEMBER 1987 



ICHTHYOPLANKTON AND STATION DATA FOR 

CALIFORNIA COOPERATIVE OCEANIC FISHERIES 

INVESTIGATIONS SURVEY CRUISES IN 1955 

David A. Ambrose 

Richard L. Charter 

H. Geoffrey Moser 

Celeste R. Santos Methot 



PLEASE RETURN 

TO 

INSTITUTION DATA LIBRARY 

Mclean 



NOAA-TM-NMFS-SWFC- 83 






U.S. DEPARTMENT OF COMMERCE 

National Oceanic and Atmospheric Administration 

National Marine Fisheries Service 

Southwest Fisheries Center 



1265-A^ 
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NOAA Technical Memorandum NMFS 



The National Oceanic and Atmospheric Administration (NOAA), organized in 
1970, has evolved into an agency which establishes national policies and 
manages and conserves our oceanic, coastal, and atmospheric resources. An 
organizational element within NOAA, the Office of Fisheries is responsible for 
fisheries policy and the direction of the National Marine Fisheries Service 
(NMFS). 

In addition to its formal publications, the NMFS uses the NOAA Technical 
Memorandum series to issue informal scientific and technical publications when 
complete formal review and editorial processing are not appropriate or feasible. 
Documents within this series, however, reflect sound professional work and may 
be referenced in the formal sr.ipntific and technical literature. 



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Woods Hole Oceanographic Institution 
ATLAS - GAZETTEER COLLECTION 



NOAA Technical Memorandum NMFS 



This TM series is used lor documentation and timely communication ol preliminary results, imienm reports, of 
special purpose information, and have not received complete tormal review, editorial control, or detailed editing 



IZ65-A6 



SEPTEMBER 1987 



ICHTHYOPLANKTON AND STATION DATA FOR 
CALIFORNIA COOPERATIVE OCEANIC FISHERIES 
INVESTIGATIONS SURVEY CRUISES IN 1955 

David A. Ambrose 

Richard L. Charter 

H. Geoffrey Moser 

Celeste R. Santos Methot 

Southwest Fisheries Center 

National Marine Fisheries Service 

La Jolla, CA 92038 



NO AA-TM-NMFS-SWFC- 83 



U.S. DEPARTMENT OF COMMERCE 
C. William Verity, Jr., Secretary 

National Oceanic and Atmospheric Administration 

Anthony J. Calio, Administrator 

National Marine Fisheries Service 

William E. Evans, Assistant Administrator for Fisheries 



CONTENTS 

Page 

List of Figures iii 

List of Tables iv 

Abstract 1 

Introduction 1 

Sampling Area and Pattern 2 

Sampling Gear and Methods 3 

Laboratory Procedures 4 

Identification 5 

Computer Entry and Editing 10 

Species Summary 11 

Explanation of Tables 11 

Acknowledgments 12 

Literature Cited 14 

Figures 17 

Tables 28 

Index 182 



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LIST OF FIGURES 

Page 

Figure 1. Composite arrangement of diagrammatic 
charts showing areas sampled on each 

CalCOFI cruise during 1955 17 

Figure 2. Station pattern for CalCOFI Cruise 5501 

showing tracks for each vessel 18 

Figure 3. Station pattern for CalCOFI Cruise 5502 19 

Figure 4. Station pattern for CalCOFI Cruise 5503 20 

Figure 5. Station pattern for CalCOFI Cruise 5504 21 

Figure 6. Station pattern for CalCOFI Cruise 5505 22 

Figure 7. Station pattern for CalCOFI Cruise 5506 23 

Figure 8. Station pattern for CalCOFI Cruise 5507 24 

Figure 9. Station pattern for CalCOFI Cruise 5510 25 

Figure 10. Station pattern for CalCOFI Cruise 5512 26 

Figure 11. The basic station plan for CalCOFI 

cruises from 1950 to the present 27 



in 



LIST OF TABLES 



Page 



Table 1. Station and plankton tow data for CalCOFI 

cruises in 1955 28 

Table 2. Pooled occurrences of fish larvae taken 

during CalCOFI cruises in 1955 63 

Table 3 . Pooled numbers of fish larvae taken during 

CalCOFI cruises in 1955 66 

Table 4. Numbers of fish larvae taken on stations 

occupied during CalCOFI cruises in 1955 69 

Table 5. Summary of pooled occurrences of fish 
larvae taken on CalCOFI cruises from 1951- 
1960 177 

Table 6. List of stations with multiple occupancies 

in one month during 1955 181 



IV 



ABSTRACT 

This report provides ichthyoplankton and associated station 
and tow data from California Cooperative Oceanic Fisheries 
Investigations (CalCOFI) cruises conducted off California and 
Baja California in 1955. It is the fifth report in a series that 
presents these data for all biological-oceanographic CalCOFI 
surveys from 1951 to the present. A total of 13 08 stations was 
occupied during 9 monthly multivessel cruises over the quarter- 
million square mile survey area which extends from the 
California-Oregon border to Cape San Lucas, Mexico and seaward to 
several hundred miles. The data are listed in a series of 6 
tables; the background, methodology, and information necessary 
for interpretation and quantitative analysis of the data are 
presented in an accompanying text. All pertinent station and tow 
data, including volumes of water strained and standard haul 
factors are listed in the first table. Another key table lists, 
by station and month, standardized counts of each of the 124 
larval fish categories identified from survey samples. This and 
previous and subsequent reports make the CalCOFI ichthyoplankton 
and station data available to all investigators and serve as 
guides to the newly developed computer data base. 

INTRODUCTION 

This report, the fifth of a series, provides ichthyoplankton 
and associated station and tow data from California Cooperative 
Oceanic Fisheries Investigations (CalCOFI) joint biological- 
oceanographic survey cruises conducted in 1955. This program 
was initiated in 1949, under the sponsorship of the Marine 
Research Committee of the State of California, to study the 
population fluctuations of the Pacific sardine (Sardinops sagax) 
and the environmental factors that may play a role in such 
fluctuations. CalCOFI, known as the California Cooperative 
Sardine Research Program from 1949 to 1953, was made up of 
representatives of the South Pacific Fisheries Investigations 
(SPFI) of the U.S. Fish and Wildlife Service [now the La Jolla 
Laboratory, National Marine Fisheries Service (NMFS) ] , the 
Scripps Institution of Oceanography (SIO) , the California 
Department of Fish and Game (CDFG) , the California Academy of 
Sciences (CAS) and the Hopkins Marine Station of Stanford 
University. The first three of these agencies supplied ships and 
personnel to conduct the sea surveys. NMFS processed the 
plankton samples and analyzed the ichthyoplankton from them. SIO 
processed and analyzed the hydrographic samples and measurements 
and also analyzed invertebrate groups from the plankton samples. 

The boundaries, station placement, and sampling frequency 
for the CalCOFI survey area were based on the results of joint 
biological and oceanographic cruises conducted by NMFS and SIO 
during 1939-41. Those cruises were designed to collect sardine 
eggs and larvae and associated hydrographic data over the entire 
areal and seasonal spawning range of the species. On these 
survey cruises, plankton tows were made to 70 m, a depth which 



encompassed the vertical distribution of sardine eggs and larvae. 
Wide-ranging joint biological and oceanographic survey cruises 
were resumed in 1949 with sardine as the focus; however, an 
increasing interest in other biological components resulted in 
the deepening of standard tows to 140 m in 1951. This marked the 
beginning of truly quantitative ichthyoplankton sampling on 
CalCOFI surveys. 

Data resulting from CalCOFI surveys in 1955 have been 
published in a number of forms. Hydrographic data (Reid et al., 
1962), zooplankton volumes (Staff, SPFI, 1956; Thrailkill, 1956; 
Smith, 1971) and ichthyoplankton data for selected species 
(Ahlstrom and Kramer, 19 57) were presented in standard formats. 
The latter lists counts for eggs and larvae of sardine and for 
larvae of northern anchovy {Engraulis mordax) , jack mackerel 
(Trachurus symmetricus) , Pacific mackerel {Scomber japonicus) , 
Pacific hake (Merluccius productus) , and rockfishes (Sebastes 
spp.). Also, length frequencies are listed for larvae of 
sardine, anchovy, jack mackerel and Pacific mackerel. 
Distribution maps of larvae of 5 of these taxa taken on CalCOFI 
surveys during 1955 are presented in the CalCOFI Atlas series 
(Kramer and Ahlstrom, 1968; Ahlstrom, 1969; Kramer, 1970; 
Ahlstrom et al., 1978). Other atlases provided distribution maps 
of 6 mesopelagic fish larvae (Ahlstrom, 1972) and 8 flatfish taxa 
(Ahlstrom and Moser, 1975) taken during 1955. 

A computer data base for eggs and larvae of sardine and 
anchovy and for larvae of hake, and the two mackerels was 
established in 1969. The development of a data base for other 
fish larvae is a complex undertaking because competency of 
identification has evolved steadily over the past 38 years. We 
began the task of producing a CalCOFI ichthyoplankton data base 
and associated data report series in 1983. All available 
original records for 1955 were subjected to an extensive 
verification and editing process to produce this report. This, 
and previous (Ambrose et al., 1987; Sandknop et al., 1987; 
Stevens et al., 1987; Sumida et al., 1987) and subsequent reports 
make the CalCOFI ichthyoplankton and station data available to 
all investigators and serve as guides to the computer data base. 
The data base will be modified when additional errors are 
discovered and when composite taxa from the earlier years are 
reidentif ied. These reports are the fundamental reference 
documents against which subsequent changes in the data base can 
be compared. 



SAMPLING AREA AND PATTERN 

In 1955, CalCOFI survey cruises were conducted at monthly 
intervals except during August, September and November. A total 
of 1308 stations included in this data base was occupied on 9 
cruises, with an average of 145 stations per cruise (range of 
105-196) . Coverage of the survey station pattern varied 
among cruises and the entire quarter-million square mile survey 
area was not covered on any single cruise (Figures 1-10; Table 



1) . The area off northern California (lines 40-57) was not 
surveyed. Central California (lines 60-77) waters were covered 
by stations occupied in May, June, July, and October. The area 
between Pt. Conception, California and Pt. San Juanico, Baja 
California (lines 80-137) was surveyed on all cruises. The area 
off southern Baja California (lines 140-157) was surveyed in 
January, February, March, and December. Coverage extended 
seaward to station 100 (approximately 3 00 miles offshore) on some 
lines in the main sampling area but typically did not extend 
beyond station 90 (approximately 160-250 miles offshore) . 

Six vessels were employed on these cruises: the Black 
Douglas, of NMFS, and the Crest, Horizon, Paolina T, Spencer F. 
Baird and Stranger of SIO. Two to three vessels participated on 
each cruise with three being the usual number. The Crest was 
used on all cruises except 5510 and 5512 and the Black Douglas on 
all but 5501, 5502, and 5506. The other four vessels 
participated on a total of 10 cruises (Reid et al., 1962). 



SAMPLING GEAR AND METHODS 

The standard CalCOFI net used from 1949 to 1969 had a 1-m 
diameter mouth opening (0.785 m area) and an overall length of 
about 5 m. The net was constructed of 30xxx gauze, a heavy duty 
grade of silk bolting cloth, with a mesh size of 0.55 mm after 
shrinkage. The last 40 cm of the cone and the cod end were 
constructed of 56xxx grit gauze which had a mesh size of 0.25 mm 
after shrinkage. The net ring was fastened to a short 3-lead 
bridle connected to several meters of line which attached to the 
towing cable by a clamp. A current meter was suspended in the 
center of the net mouth to measure volume of water filtered (see 
Kramer et al., 1972, for further details). 

The standard tow from 1951 through 1968 was an oblique haul 
to 140 m depth (to 15 m of the bottom in shallow areas) designed 
to filter a constant amount of water per depth interval (ca. 
3m 3 /m of depth) over the vertical range of most ichthyoplankters. 
Hauls were made at a ship speed of 1.5-2.0 knots and initiated by 



1 CalC0FI lines (Figure 11) are arranged perpendicular to the 
coastline and extend from the Canadian border (line 10) to below 
Cape San Lucas, Baja California (line 157) . Stations were 
established on the basis of a perpendicular to line 80 (off Pt. 
Conception) at a point designated as station 60. Stations were 
plotted seaward and shoreward from station 60 on each line. 
Cardinal CalCOFI lines (those ending in "0") are 120 miles apart 
and usually bracket two ordinal lines (ending in "3" or "7") , so 
that lines are 40 miles apart over most of the pattern. Cardinal 
stations are 40 miles apart and typically these are separated by 
a station number ending in "5" so that stations are 20 miles 
apart out to station 90 on most lines. Stations are placed at 
closer intervals near the coast and islands to accommodate these 
features (see Kramer et al., 1972 for further details). 



clamping the net line to the towing cable with the 4 5 kg terminal 
weight about 10-15 m below the surface. The net was lowered to 
140 m depth by paying out 200 m of wire over a 4 minute period 
(35 m of depth/min.). After fishing at depth for 30 seconds, the 
net was retrieved at 20 m/min. (14 m depth/min.). The angle of 
stray of the towing cable was recorded every 3 seconds and 
maintained at 45° (+3°) by adjusting the ship speed and course. 
After reaching the surface, the net was washed down and the 
samples preserved in 5% formalin buffered with sodium borate. 
Flowmeter readings were made at the beginning and end of each 
tow. Detailed descriptions of gear and methods are given by 
Ahlstrom (1953), Kramer et al. (1972), and Smith and Richardson 
(1977) . 



LABORATORY PROCEDURES 

Laboratory processing began with the determination of a 
displacement volume for each sample (methods described in Staff, 
SPFI, 1953 and Kramer et al . , 1972). Zooplankton volumes 
(including ichthyoplankton) of samples collected in 1955 are 
listed in Staff, SPFI (1956) and presented graphically in 
Thrailkill (1956) and Smith (1971). 

Sorting involved the removal of ichthyoplankton from the 
sample and identification and separation of eggs and larvae of 
selected species (see introduction) . Usually, each sample was 
sorted completely; however, some of the samples were fractioned 
into aliquots using a Folsom plankton splitter (McEwen et al., 
1954) prior to sorting. Several criteria were used to determine 
whether a sample was fractioned: samples containing an abundance 
of thaliacians and coelenterates and exceeding 150 ml in total 
plankton volume were fractioned (to 50%, 25%, 12.5%, or 6.25%) to 
approximate a reduced volume of 50 ml for sorting; samples with 
an excessive quantity of fish eggs and/or larvae were 
occasionally fractioned to expedite the sorting process in order 
to meet scheduled deadlines. If the identified fraction of an 
aliquot yielded rare or interesting species of fish larvae, the 
remaining fraction was frequently sorted and identified with the 
intent of findinq additional specimens. Aliquot percentaqes for 
fractioned samples from 1955 are listed in Table 1 under the 
"Percent Sorted" column. 

A "standard haul factor" (SHF) was calculated for each tow 
to make them comparable and allow estimations of areal abundance. 
This factor adjusts the number of eggs or larvae in a haul to the 
number in 10 m of water strained per meter of depth fished. If 
the vertical distribution of the species has been encompassed, 
then the adjusted value is equivalent to the number under 10 m 
of sea surface. The SHF is calculated for each haul by the 
formula: 



o . .... 

Personal communication, James R. Thrailkill, National Marine 

Fisheries Service, Southwest Fisheries Center, La Jolla, CA. 



SHF = 10 D 



where D = depth of haul = cosine of the average angle 
of stray of the towing cable multiplied by 
cable length (m) 

V = total volume of water (m ) strained 

during the haul 

V = R • a • p 

where R = total number of revolutions of the current 
meter during the haul 

a = area (m ) of the mouth of the net 

p = length of column of water (m) needed to 
produce one revolution of the current 
meter. 

Tow depth, volume of water strained, and standard haul factor are 
listed in Table 1 for each tow taken during 1955. Detailed 
descriptions of factors involved in calculating these values are 
presented in Ahlstrom (1948), Kramer et al. (1972), and Smith and 
Richardson (1977) . 



IDENTIFICATION 

Identification of ichthyoplankton species beyond those 
separated during the sorting process was carried out by a 
separate group of specialists. Ontogenetic stages of fishes are 
inherently difficult to identify and this is further complicated 
by the large number and diversity of species which contribute to 
the ichthyoplankton of the California Current region. Most 
identifications were accomplished by establishing ontogenetic 
series on the basis of morphology, meristics, and pigmentation 
and then identifying these series by relating them to known 
metamorphic, juvenile, or adult stages with overlapping features 
(Powles and Markle, 1984) . A total of about 122 taxa was 
identified for 1955, with 72 taken to species, 22 to genus, 24 to 
family, and 4 to order. Some of the developmental series 
recognized originally could not be assigned scientific names, 
particularly in the Bathylagidae, Myctophidae, and 
Pleuronectiformes. These were given descriptive names, which 
later were changed to scientific names as they became known. 

The task of producing a reliable and equitable 
ichthyoplankton data base required extensive procedures to 
verify, correct, and edit the original identifications. The 
primary data source was the original identification sheets (see 
Kramer et al., 1972, for examples); however, a critical resource 
used in all phases of this process was the CalCOFI 
ichthyoplankton collection in which the samples are archived. 



Throughout the course of CalCOFI ichthyoplankton studies, samples 
have been identified to the lowest taxon possible. In reviewing 
these identifications for the data base, our approach has been 
conservative and we have preserved those identifications and 
counts which we could confirm, while correcting as many of the 
errors as possible. During the coding of the identification 
sheets, the "descriptive types" were assigned scientific names 
and reexamined, if necessary. After computer entry, taxonomic 
errors and inconsistencies in the data base were corrected and 
the most obvious identification errors were corrected. Our 
current knowledge of ichthyoplankton techniques coupled with a 
precise understanding of the development of identification 
competency in the program over the years allowed us to critically 
judge the historical records. Identifications were changed to 
different taxa, lumped to a higher taxonomic category, or given a 
more precise taxonomic name. In many cases, identifications of a 
taxon were inconsistent among cruises in a year, because of 
varying competency of identifiers. These records were made 
equitable by lumping to the higher taxonomic category to avoid 
biases that could result in quantitative misinterpretations. 

Next, statistical, seasonal, and geographic outliers were 
identified, employing a series of graphic summaries and listings. 
Examination of geographic outliers proved to be especially 
effective because of our accumulated knowledge of species 
distributions. In the course of examining samples for these 
outliers, other identification errors were discovered and 
eventually all taxa were scrutinized to some extent. Lastly, 
certain taxa were reexamined in all samples for the entire 
CalCOFI time series. These taxa were selected because of their 
commercial, ecological, phylogenetic, or zoogeographic importance 
or because taxonomic confusion was at the ordinal level. The 
following is a list of the taxa for 1955 which received special 
attention, with explanations and caveats intended to aid in 
quantitative interpretations: 

Anguilliformes - tentative and sporadic identifications to family 
or lower taxon lumped to order. 

Sardinops sagax - all specimens south of line 120 checked for 
misidentif ication of Opisthonema spp. 

Engraulidae - includes nearshore taxa (mostly Anchoa spp.) large 
enough to separate from Engraulis mordax . Some nearshore 
samples of small E. mordax may contain other anchovy genera, 
but could not be differentiated. 

Nansenia spp. - all specimens checked and identified as N. 

Candida or N. crassa ; all specimens of these species near 
their range boundaries checked. 

Bathylagus spp. - all specimens rechecked; residuals are small, 
poorly preserved specimens which possibly include 
Leuroglossus stilbius . 



Sternoptychidae - tentative and sporadic identifications of 
hatchet fishes to genus were lumped to family. 

Bathophilus spp. - all specimens checked. 

Tactostoma macropus - all specimens checked, and identifications 
changed to Bathophilus spp. 

Scopelarchidae - tentative and sporadic identifications to genus 
lumped to family. 

Lampanyctus spp. - tentative and sporadic identifications to 
species (mostly descriptive types) lumped to genus; 
identification of L. regalis and L. ritteri began in 1954. 

Lampanyctus regalis - underrepresented because of inability to 
differentiate small larvae (<5 mm) from those of other 
species of the genus; counts may include other species of 
the genus because of difficulty in identifying larvae of 
this large and complex genus. 

Lampanyctus ritteri - comment for L. regalis applies to this 
species. 

Stenobrachius leucopsarus - all specimens south of line 120 
checked. 

Diogenichthys atlanticus - all specimens at margins of range 
checked . 

Diogenichthys laternatus - all specimens at margins of range 
checked . 

Electrona rissoi - recognition of this species was inconsistent 
and others may be included in Protomyctophum crockeri or 
Myctophidae. 

Hygophum spp. - all specimens reidentified to species; residuals 
are small, poorly preserved specimens. 

Myctophum aurolaternatum - specimens checked; originally called 
"Astronesthidae" . 

Protomyctophum crockeri - some samples on northern lines may 
contain P. thompsoni , which was not identified at the 
time. 

Symbolophorus calif orniensis - all specimens south of line 120 
checked for confusion with Hygophum spp., stemming from 
descriptive names. 

Bregmaceros spp. - all gadiform types (see Index) , except 
Merluccius productus and Macrouridae, reexamined. 



Ophidiiformes - this category did not exist originally and 

ophidiiform larvae were included in Brosmophysis marginata , 
Carapidae, "Otophidium" , "Zoarcidae" , and "blenny"; identi- 
fications of B. marginata and Carapidae proved to be mostly 
correct and "Zoarcidae" to be a yet unidentified ophidiiform 
species; all "Otophidium" and "blenny" were reexamined and 
the former included Ophidion scrippsae , Chilara taylori and 
other ophidiiform taxa (moved to order) ; "blenny" contained 
0. scrippsae, C. taylori, and other ophidiiform taxa in 
addition to true blennioids. 

Ceratioidei - identifications of this group were inconsistent and 
specimens may be in the unidentified fish larva category. 

Trachipteridae - tentative and sporadic identifications to genus 
were lumped to family. 

Melamphaes spp. - all identifications ascribed to Melamphaidae 
were reexamined and assigned to genus (Melamphaes, 
Poromitra) or species (Scopelogadus bispinosus) . Larvae 
originally identified as Melamphaes spp. were not reexamined 
and this category may contain other melamphaid genera. 

Cottidae - some samples may include specimens of Scorpaenichthys 
marmoratus , hexagrammids (e.g., Oxylebius pictus, Zaniolepis 
spp.), and some blennioids (e.g., Hypsoblennius spp.). 

Zaniolepis spp. - all specimens checked. 

Sebastes spp. - in addition to other scorpaenid genera, 

category includes Prionotus spp. , serranids, scombrids, and 

other spiny-headed shorefishes, particularly in samples 
south of line 12 0. 

Sebastolobus spp. - absence of this category probably results 
from misidentif ication as Sebastes spp. 

Hypsoblennius spp. - some specimens remain in Cottidae. 

Clinidae - some specimens remain in Cottidae or unidentified fish 
larva category. 

Labridae - tentative and sporadic identifications to genus were 
lumped to family. 

Chromis punctipinnis - records south of about line 120 may 
include other pomacentrid taxa. 

Carangidae - all specimens checked; tentative and sporadic 
identifications to genus or species (except Trachurus 
symmetricus and Seriola lalandi) were lumped to family. 

Seriola lalandi - all specimens checked. 



8 



Gerreidae - larvae of this family and other shorefishes (e.g., 
Haemulidae, Girella nigricans, Mullidae, Priacanthidae) 
were not identified and may be in the unidentified fish 
larva category or may be misidentif ied as Sebastes spp., 
Cottidae, etc. 

Medialuna calif orniensis - all specimens checked. 

Caulolatilus princeps - all specimens checked. 

Sciaenidae - tentative and sporadic identifications to genus 
lumped to family. 

Serranidae - this family is underrepresented and some specimens 
may be in the unidentified fish larva category or may have 
been misidentif ied as Sebastes spp. 

Scombridae - all larvae identified to this family or constituent 
taxa (except Scomber japonicus) were reexamined and reas- 
signed; underrepresentation or absence of these taxa may be 
attributed to misidentif ication or they may be in the uni- 
dentified fish larva category. 

Nomeidae - absence of this family attributed to misidentif ication 
or placement in unidentified fish larva category. 

Pleuronectiformes - all available specimens of this category 

(originally called "flatfish") were examined and reidenti- 
fied; residuals are small, poorly preserved specimens. 

Bothidae - all specimens examined and reassigned; most were 

assigned to various paralichthyid genera or to Bothus spp. 

Citharichthys spp. - all larvae identified to genus or to a 

species of the genus from 1954 through 1960 were checked and 
identified to species; residuals are small, poorly preserved 
specimens or those with variable taxonomic characters. 

Etropus spp. - larvae of this taxon were originally lumped with 
Citharichthys spp. ; present records result from complete 
re identification of Citharichthys spp. 

Hippoglossina spp. - all specimens of this genus (originally 

called "pigmented bothid") were examined and assigned to 
H. stomata . 

Paralichthys spp. - all specimens of this genus were examined and 
most were assigned to P. californicus or Xystreurys 
liol epi s . 

Xystreurys liolepis - originally misidentif ied as Paralichthys 
californicus ; all specimens reidentif ied. 

Glyptocephalus zachirus - all specimens examined. 



Microstomas pacificus - all specimens examined. 

Pleuronichthys spp. - all larvae of this genus and constituent 

species were examined and assigned to species; residuals are 
small, poorly preserved specimens. 

Psettichthys melanostictus - absence of this species may be 

explained by misidentif ication with other flatfish species 
(e.g., Lyopsetta exilis) which we did not reexamine 
systematically. 



COMPUTER ENTRY AND EDITING 

Each taxon on the original identification sheets was given a 
3-digit code based on the list of codes in Haight et al. (1979). 
Taxon codes and counts from these sheets were keypunched by 
cruise and station, along with pertinent station and tow data and 
entered into the VAX 11/780 computer at the University of 
California, San Diego Computing Center. After entries were 
completed for an entire year, print-out listings of taxa and 
counts on each station were compared with the original data 
sheets to eliminate keypunch errors. Next, data in the file were 
cross-checked with data on an existing file which contained: 
station and tow data; numbers of eggs of sardine, anchovy, and 
saury (Cololabis saira) ; numbers of larvae of sardine, anchovy, 
hake, jack mackerel, and Pacific mackerel; total number of fish 
eggs; and total number of fish larvae. 

Discrepancies in ichthyoplankton data in these two files 
were corrected by inspecting original records from the sorting 
laboratory, the original ichthyoplankton identification sheets, 
and the samples themselves. Station and tow data discrepancies 
between the two files were corrected by reviewing ships' logs and 
deck tow sheets, original records from the sorting laboratory, 
cruise announcements, publications, header information on the 
ichthyoplankton identification sheets, and station plots 
generated for each cruise. Eventually all station and tow data 
were checked by comparing these sources. 

The corrected ichthyoplankton data base was then examined 
statistically and outliers were found and checked as above. 
Distributional plots were then prepared for each taxon and these 
were checked by reviewing the data sources mentioned above and by 
examining archived specimens. A listing of each taxon by station 
(Table 4) was produced, which became the primary document for 
subseguent checks. Misidentif ications found in geographic 
outlier checks and other misidentif ications and data problems 
discovered in the course of examining archived samples resulted 
in several iterations of Table 4. Finally, totals in Table 4 
were checked against annual summaries of incidence and abundance 
(Tables 2 and 3). Ecological analyses of the data (Moser et al., 
1987) were conducted concurrently with editing procedures and 
provided cross-checks that allowed correction of errors. 



10 



SPECIES SUMMARY 

Larvae of northern anchovy (Engraulis mordax) represented 
39% of all fish larvae taken on CalCOFI cruises during 1955 and 
numbered over twice as many as Pacific hake (Merluccius 
productus) , the next most abundant species (Tables 2, 3). 
Anchovy and hake ranked 2nd and 5th in incidence, respectively. 
Larvae of Sebastes spp., a composite of about 70 species of 
rockfish, were next most abundant (8% of total) and ranked 1st in 
occurrence. These 3 taxa accounted for about 2/3 of all fish 
larvae taken in 1955. Leuroglossus stilbius , a deepsea smelt, 
ranked 4th in abundance and 3rd in occurrence. Larvae of 
Sardinops sagax , Pacific sardine, ranked 5th in abundance but 
only 12th in occurrence, indicating relatively large sample 
sizes. Jack mackerel, Trachurus symmetricus , larvae ranked 6th 
in both abundance and incidence. The next most abundant were 
three midwater species, a lanternfish (Triphoturus mexicanus) , a 
lightfish (Vinci guerria lucetia) , and another lanternfish 
(Stenobrachius leucopsarus) , ranking 7th, 8th, and 9th, 
respectively. Triphoturus ranked 4th in occurrence, Vinciguerria 
was 7th, and Stenobrachius 13th. The 10th ranked taxon by 
abundance was the sanddab genus Citharichthys composed of 
specimens that could not be identified to species because of 
small size, poor condition, or variable taxonomic characters. 
These 10 top-ranking taxa contributed 87% of all larvae taken 
during 1955. The remaining 13% was represented by 112 taxa plus 
the unidentified and disintegrated categories. Of the 10 taxa, 4 
were midwater species, 3 were coastal demersal species or generic 
groupings, and 3 were coastal pelagic species. 



EXPLANATION OF TABLES 

Table 1 - This table lists by cruise the pertinent station and 
tow data for 1955, the volume of water filtered and 
standard haul factor for each tow, the percent of 
sample sorted, and the total numbers of fish eggs and 
larvae. CalCOFI cruises are designated by four digits; 
the first two indicate the year and the second two the 
month. Within each cruise the data are listed in order 
of increasing line and station number (southerly and 
seaward directions) ; the order of station occupancy is 
shown on the station charts (Figures 2-10) . Stations 
are designated by two groups of digits; the first set 
indicates the line and decimal fraction and the second 
set indicates the station on the line. Decimal 
fractions were not used in 1955. Time is listed as 
Pacific Standard Time at the start of each tow in 24- 
hour designation. Methods for determining tow depth, 
volume of water strained, standard haul factor, and 
percent sorted were described in the methods section. 
The values for total fish eggs and larvae represent raw 
counts (unadjusted for percent sorted or standard haul 
factor) . Ship codes are as follows: BD, Black 



11 



Douglas) CR, Crest; HO, Horizon; PT, Paolina T; SB, 
Spencer F. Baird; ST, Stranger . 

Table 2 - This table lists pooled occurrences of all larval fish 
taxa taken during 1955 in ranked order. 

Table 3 - This table lists pooled counts of all larval fish taxa 
taken during 1955 in ranked order. Numbers are 
adjusted for percent sorted and standard haul factors. 

Table 4 - This table gives numbers of fish larvae for each taxon, 
listed by station and calendar month in which the tow 
was taken. Counts are adjusted for percent of sample 
sorted and standard haul factor. Average values are 
given for stations occupied more than once during a 
month. See Table 1 for station and tow data and Table 
6 for listing of stations with multiple occupancies 
during a month. Multiple occupancies occurred when a 
station was occupied more than once during a calendar 
month; in some cases multiple occupancies resulted from 
separate cruises. The orders are listed in 
"phylogenetic" sequence modified from Nelson (1984) . 
Subtaxa within each order are listed alphabetically. 
Page numbers for each taxon are given in the index at 
the end of the report. 

Table 5 - This table is a summary of pooled occurrences of all 
larval fish taxa taken on CalCOFI surveys from 1951 to 
1960. Taxa are listed in the same order as in Table 4. 

Table 6 - List of stations with multiple occupancies in one month 
during 1955. 



ACKNOWLEDGMENTS 

Elbert Ahlstrom, Robert Counts, Lois Hunter, and David 
Kramer originally identified larvae from CalCOFI cruises of 
1955. Ronald Whyte coded each larval fish taxon or type and 
Rita Ford entered them into the computer. Debby Snow efficiently 
assisted in all aspects of data editing and retrieval. Cindy 
Meyer, Larry Zins, and James Ryan provided programming 
assistance. Dorothy Roll designed the CalCOFI data acquisition 
system and provided data processing support. Ken Raymond, Roy 
Allen, and Henry Orr helped with graphics and production of the 
report. Lorraine Prescott and Diane Forsythe prepared the 
manuscript for printing. Paul Smith determined statistical 
outliers, provided assistance during geographical outlier checks 
and offered helpful suggestions throughout the project. Izadore 
Barrett, Director of the Southwest Fisheries Center and Reuben 
Lasker, Chief, Coastal Fisheries Resources Division, SWFC, 
provided the support critical to the completion of the project. 
James Thrailkill planned CalCOFI surveys and supervised cruises, 
data handling, and plankton sorting from 1949 to 1986 and is 
largely responsible for the high quality of these operations. 



12 



Without the vision and direction of Elbert Ahlstrom and Elton 
Sette and the dedicated efforts of the many people who collected, 
processed, and analyzed the samples, this data base would not 
exist. 



13 



LITERATURE CITED 



Ahlstrom, E. H. 1948. A record of pilchard eggs and larvae 
collected during surveys made in 1939 to 1941. U.S. Fish 
Wildl. Serv. SSRF 54, 82 p. 

Ahlstrom, E. H. 1953. Pilchard eggs and larvae and other fish 
larvae, pacific coast - 1951. U.S. Fish Wildl. Serv. SSRF 
102, 55 p. 

Ahlstrom, E. H. 1969. Distributional atlas of fish larvae in the 
California Current region: jack mackerel, Trachurus 
symmetricus , and Pacific hake, Merluccius productus , 1951 
through 1966. CalCOFI Atlas No. 11: xi + 187 p. 

Ahlstrom, E. H. 1972. Distributional atlas of fish larvae in 
the California Current region: six common mesoplagic fishes 
- Vinciguerria lucetia, Triphoturus mexicanus , Stenobrachius 
leucopsarus, Leuroglossus stilbius, Bathylagus wesethi , and 
Bathylagus ochotensis , 1955 through 1960. CalCOFI Atlas No. 
17: xv + 306 p. 

Ahlstrom, E. H. and D. Kramer. 1957. Sardine eggs and larvae 
and other fish larvae, Pacific Coast - 1955. U.S. Fish 
Wildl. Serv. SSRF 224, 90 p. 

Ahlstrom, E. H. and H. G. Moser. 1975. Distributional atlas 
of fish larvae in the California Current region: 
Flatfishes, 1955 through 1960. CalCOFI Atlas No. 23: xix + 
207 p. 

Ahlstrom, E. H. , H. G. Moser, and E. M. Sandknop. 1978. 
Distributional atlas of fish larvae in the California 
Current region: Rockfishes, Sebastes spp., 1950 through 
1975. CalCOFI Atlas No. 26: xxi + 178 p. 

Ambrose, D. A., R. L. Charter, H. G. Moser, and C. R. Santos 
Methot. 1987. Ichthyoplankton and station data for 
California Cooperative Oceanic Fisheries Investigations 
survey cruises in 1951. U.S. Dep. Commer., NOAA Tech. 
Memo., NMFS, SWFC, No. 79, 196 p. 

Haight, C. A., H. G. Moser, and P. E.' Smith. 1979. Data entry 
programs: CalCOFI. II. Fish eggs and larvae identification 
sheet. National Marine Fisheries Service, Southwest 
Fisheries Center, La Jolla, Admin. Rept. No. LJ-79-25. 

Kramer, D. 1970. Distributional atlas of fish eggs and 

larvae in the California current region: Pacific sardine, 

Sardinops caerulea (Girard) , 1951 through 1966. CalCOFI 
Atlas No. 12:vi + 277 p. 



14 



Kramer, D. and E. H. Ahlstrom. 1968. Distributional atlas of 
fish larvae in the California Current region: Northern 
anchovy, Engraulis mordax (Girard) , 1951 through 1965. 
CalCOFI Atlas No. 9: xi + 269 p. 

Kramer, D. , M. Kalin, E. G. Stevens, J. R. Thrailkill, and J. R. 
Zweifel. 1972. Collecting and processing data on fish eggs 
and larvae in the California Current Region. NOAA Tech. 
Rep. NMFS Circ. 370, 38 p. 

McEwen, G. F. , M. W. Johnson, and T. R. Folsom. 1954. A 
statistical analysis of the performance of the Folsom 
Plankton Sample Splitter, based on test observations. Arch. 
Meteor. Geophys. Bioklim. Ser. A, 7:502-527. 

Moser, H. G., P. E. Smith, and L. E. Eber. 1987. Larval fish 
assemblages in the California Current region during 1954- 
1960, a period of dynamic environmental change. CalCOFI 
Rep. 28:97-127. 



Nelson, J. S. 1984. 
N.Y. , 523 p. 



Fishes of the world. John Wiley and Sons, 



Powles, H. and D. F. Markle. 1984. Identification of larvae, p. 
31-3 3. In: Ontogeny and systematics of fishes. H. G, 
Moser, W. J. Richards, D. M. Cohen, M. P. Fahay, A. W. 
Kendall, Jr., and S. L. Richardson (eds.). Spec. Publ. 
No. 1. Amer. Soc. Ichthyol. Herpetol . , 760 p. 

Reid, J. L. , Jr., R. S. Arthur, and E. B. Bennett, (eds.), 
1962. Oceanic observations of the Pacific: 1955. Univ, 
Calif. Press, Berkeley, 477 p. 



Sandknop, E. M. , R. L. Charter, H. G. Moser, and J. D. Ryan. 
1987. Ichthyoplankton and station data for California 
Cooperative Oceanic Fisheries Investigations survey cruises 



in 1952. U.S. 
No. 80, 207 p. 



Dep. Commer., NOAA Tech. Memo., NMFS, SWFC, 



Smith, P. E. 1971. Distributional atlas of zooplankton volume 
in the California Current region, 1951 through 1966. 
CalCOFI Atlas No. 13: xvi + 144 p. 

Smith, P. E. and S. L. Richardson. 1977. Standard techniques 
for pelagic fish egg and larva surveys. FAO Fish. Tech. 
Pap. No. 175, 100 p. 

Staff, South Pacific Fishery Investigations. 1953. Zooplankton 
volumes off the Pacific Coast, 1952. U.S. Fish Wildl. Serv. 
SSRF 100, 41 p. 



Staff, South Pacific Fishery Investigations. 

volumes off the Pacific Coast, 1955. U.S. 
SSRF 177, 31 p. 



1956. Zooplankton 
Fish Wildl. Serv. 



15 



Stevens, E. G., R. L. Charter, H. G. Moser, and M. S. Busby. 
1987. Ichthyoplankton and station data for California 
Cooperative Oceanic Fisheries Investigations survey cruises 
in 1953. U.S. Dep. Commer. , NOAA Tech. Memo., NMFS, SWFC, 
No. 81, 186 p. 

Sumida, B. Y. , R. L. Charter, H. G. Moser, and D. L. Snow. 
1987. Ichthyoplankton and station data for California 
Coopertive Oceanic Fisheries Investigations survey cruises 
in 1954. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC, 
No. 82, 207 p. 

Thrailkill, J. R. 1956. Relative areal zooplankton abundance 
off the Pacific coast. U.S. Fish Wildl. Serv. SSRF 188, 
85 p. 



16 




Figure 1. Composite arrangement of diagrammatic charts showing 
areas sampled on each CalCOFI cruise during 1955. 



17 



i i r 



CRUISE 5501 




PAOLINA-T 



J L 



FROM H.O. 0327 ft HO. 1006 IREO) 
I I II I I 



20° 



120" 



115° 



110° 



105° 



Figure 2. Station pattern for CalCOFI Cruise 5501 showing tracks 
for each vessel. Stations with plankton tows only are 
indicated by a dot; those with plankton tows and 
hydrographic measurements are shown by a dot and 
circle. Modified from charts in Reid et al . (1962) to 
include only those stations listed in Table 1 of this 
report. 



18 




SPENCER F. BAIRD 



120° 



J L 



115° 



J__J L 



FROM M-O-0327 6 HO, 1006 (RED) 

I 1 l__J I 



110° 



105° 



Figure 3. Station pattern for CalCOFI Cruise 5502 
Figure 2 . 



Symbols as in 



19 



105° 



CRUISE 5503 




HORIZON 



J L 



120° 



115° 



I ' L 



FROM H,0. 0527 8 H.O. 1006 (RED.) 

' I I ' i I 1 



110° 



105* 



Figure 4. Station pattern for CalCOFI Cruise 5503 
Figure 2. 



Symbols as in 



20 



40' 



35' 



50' 



25' 




CREST/ 
BLACK DOUGLAS 



BLACK DOUGLAS 



J L 



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110° 



Figure 5. Station pattern for CalCOFI Cruise 5504 
Figure 2 . 



Symbols as in 



21 



40' 



35 



30' 



25' 



CRUISE 5505 



BLACK DOUGLAS 




CREST 



FROM HG 0S2T 

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125° 



120* 



115° 



110° 



Figure 6. Station pattern for CalCOFI Cruise 5505, 
Figure 2 . 



Symbols as in 



22 



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30' 



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CRUISE 5506 




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Figure 7. Station pattern for CalCOFI Cruise 5506, 
Figure 2 . 



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Figure 8. Station pattern for CalCOFI Cruise 5507 
Figure 2 . 



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1 








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Figure 2. 



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CALCOFI 
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to the present. 



27 



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62 



TABLE 2. Pooled occurrences of fish larvae taken during CalCOFI 
cruises in 1955. 

Rank Taxon 



1 Sebastes spp. 

2 Engraulis mordax 

3 Leuroglossus stilbius 

4 Triphoturus mexicanus 

5 Merluccius productus 

6 Trachurus symmetricus 

7 Vinciguerria lucetia 

8 Protomyctophum crockeri 

9 Lampanyctus ritteri 

10 Bathylagus wesethi 

11 Diogenichthys laternatus 

12 Sardinops sagax 

13 Stenobrachius leucopsarus 

14 Citharichthys stigmaeus 

15 Cyclotbone spp. 

16 Melamphaes spp. 

17 Citharichthys xanthostigma 

18 Bathylagus ochotensis 

19 Citharichthys spp. 

20 Disintegrated fish larva 

21 Citharichthys fragilis 

22 Icichthys lockingtoni 

23 Tarletonbeania crenularis 

24 Symbolophorus calif orniens is 

25 Unidentified fish larva 

26 Scomber japonicus 

27 Diogenichthys atlanticus 

28 Stomias atriventer 

29 Diaphus spp. 

29 Argentina sialis 

31 Paralepididae 

32 Tetragonurus cuvieri 

33 Nansenia crassa 
33 Sciaenidae 

35 Myctophidae 

36 Ceratoscopelus townsendi 
36 Lampanyctus spp. 

38 Lyopsetta exilis 

38 Cottidae 

38 Labridae 

41 Gobiidae 

41 Citharichthys sordidus 

43 Cololabis saira 

44 Parophrys vetulus 

45 Chauliodus macouni 

46 Peprilus simillimus 

47 Hygophum atratum 

48 Synodus spp. 



Occurrences 


637 


569 


508 


475 


439 


369 


338 


312 


296 


286 


265 


255 


251 


206 


184 


166 


163 


162 


158 


124 


107 


105 


103 


102 


99 


93 


90 


87 


81 


81 


80 


65 


61 


61 


60 


58 


58 


57 


57 


57 


56 


56 


54 


50 


49 


47 


43 


41 



63 



TABLE 2. (cont. ) 



Rank Taxon Occurrences 

48 Sternoptychidae 41 

50 Gonichthys tenuiculus 37 

51 Scopelarchidae 34 

52 Ichthyococcus spp. 30 
52 Microstomus pacificus 30 
54 Pleuronichthys verticalis 26 
54 Ophidiif ormes 26 

56 Trichiuridae 25 

57 Chiasmodontidae 24 

58 Notoscopelus resplendens 23 
58 Pleuronichthys spp. 2 3 
60 Hippoglossina stoma ta 22 
60 Microstoma microstoma 22 
60 Paralichthys californicus 2 2 

63 Lampanyctus regalis 19 

64 Pleuronichthys coenosus 17 

65 Clinidae 15 

66 Chilara taylori 14 
66 Hypsoblennius spp. 14 
66 Loweina rara 14 
69 Hygophum reinhardtii 13 
69 Trachipteridae 13 
69 Pleuronectif ormes 13 
69 Myctophum nitidulum 13 

73 Symphurus spp. 11 

7 4 Lampadena urophaos 10 

74 Idiacanthus antrostomus 10 
74 Agonidae 10 

77 N an sen i a Candida 8 

78 Scopelogadus bispinosus 6 
78 Syngnathus spp. 6 
78 Diogenichthys spp. 6 
78 Hygophum spp. 6 
78 Brosmophycis marginata 6 
83 Medialuna californiensis 5 
83 Glyptocephalus zachirus 5 
83 Etrumeus acuminatus 5 
83 Cyclopteridae 5 

8 3 Bathophilus spp. 5 
88 Pleuronichthys decurrens • 4 
88 Anguilliformes 4 
88 Poromitra spp. 4 
88 Caulolatilus princeps 4 
88 Pleuronichthys ritteri 4 
88 Aristostomias scintillans 4 
88 Xystreurys liolepis 4 
88 Zaniolepis spp. 4 
88 Scorpaenichthys marmoratus 4 
88 Ophidion scrippsae 4 



64 



TABLE 2. (cont . ) 



Rank Taxon Occurrences 

88 Chromis punctipinnis 4 

99 Sphyraena argentea 3 

99 Bathylagus spp. 3 

99 Stomii formes 3 

99 Atherinidae 3 

99 Bregmaceros spp. 3 

99 Macrouridae 3 

105 Ophiodon elongatus 2 

105 Bathylagus pacificus 2 

105 Seriola lalandi 2 

105 Scombridae 2 

105 Macroramphosus gracilis 2 

105 Brama spp. 2 

111 Myctophum aurolaternatum 1 

111 Electrona rissoi 1 

111 Alepocephalidae 1 

111 Notolychnus valdiviae 1 

111 Engraulidae 1 

111 Bothus spp. 1 

111 Serranidae 1 

111 Diplophos taenia 1 

111 Evermannellidae 1 

111 Scopelosaurus spp. 1 

111 Gobiesocidae 1 

111 Bathylagus milleri 1 

111 Carapidae 1 

111 Tetraodontidae 1 



65 



TABLE 3. Pooled numbers of fish larvae taken during CalCOFI 

cruises in 1955. Counts are adjusted for percent of 
sample sorted and standard haul factor (see text). 

Rank Taxon Count 

1 Engraulis mordax 139090 

2 Merluccius productus 61516 

3 Sebastes spp. 27466 

4 Leuroglossus stilbius 14793 

5 Sardinops sagax 14189 

6 Trachurus symmetricus 13 269 

7 Triphoturus mexicanus 127 88 

8 Vinciguerria lucetia 12641 

9 Stenobrachius leucopsarus 7479 

10 Citharichthys spp. 7058 

11 Diogenichthys laternatus 4715 

12 Citharichthys xanthostigma 4280 

13 Citharichthys fragilis 3463 

14 Bathylagus wesethi 3231 

15 Citharichthys stigmaeus 2708 

16 Lampanyctus ritteri 2088 

17 Scomber japonicus 19 58 

18 Protomyctophum crockeri 1681 

19 Cyclothone spp. 152 3 

20 Icichthys lockingtoni 1372 

21 Bathylagus ochotensis 1295 

22 Sciaenidae 1211 
2 3 Diaphus spp. 9 36 

24 Peprilus simillimus 928 

25 Argentina sialis 846 

26 Tarletonbeania crenularis 842 

27 Melamphaes spp. 7 41 

28 Diogenichthys atlanticus 697 

29 Sy nodus spp. 643 

30 Symbolophorus calif orniensis 629 

31 Disintegrated fish larva 586 

32 Unidentified fish larva 553 

33 Labridae 526 

34 Lyopsetta exilis 514 

35 Tetragonurus cuvieri 486 

36 Ceratoscopelus townsendi 443 

37 Cottidae 428 

38 Stomias atriventer 418 

39 Lampanyctus spp. 388 

40 Cololabis saira 352 

41 Pleuronichthys verticalis 343 

42 Parophrys vetulus 342 

43 Paralepididae 317 

44 Citharichthys sordidus 312 

45 Hygophum atratum 301 

46 Nansenia crassa 277 

47 Pleuronichthys spp. 266 



66 



TABLE 3. (cont. ) 

Rank Taxon Count 



48 Gobiidae 234 

49 Myctophidae 228 

50 Paralichthys calif ornicus 217 

51 Chauliodus macouni 205 

52 Sternoptychidae 183 

53 Microstomus pacificus 111 

54 Gonichthys tenuiculus 144 

55 Pleuronectiformes 138 

56 Hippoglossina stomata 13 3 

57 Clinidae 122 

58 Scopelarchidae 113 

59 Ichthyococcus spp. 112 

60 Sphyraena argentea 109 

61 Trichiuridae 107 

62 Notoscopelus resplendens 101 

63 Ophidiiformes 99 

64 Microstoma microstoma 95 

65 Chiasmodontidae 93 

66 Lampanyctus regalis 92 

67 Pleuronichthys coenosus 91 

68 Pleuronichthys ritteri 87 

69 Trachipteridae 74 

70 Hypsoblennius spp. 71 

71 Symphurus spp. 66 

72 Etrumeus acuminatus 60 
72 Hygophum reinhardtii 60 

74 Xystreurys liolepis 52 

75 Myctophum nitidulum 51 

76 Caulolatilus princeps 50 

77 Chilara taylori 49 

78 Loweina rara 41 

79 Lampadena urophaos 39 

80 Bregmaceros spp. 36 
80 Agonidae 36 

82 Hygophum spp. 33 

83 Bathophilus spp. 32 

84 Nansenia Candida 31 

85 Idiacanthus antrostomus 30 

86 Glyptocephalus zachirus 29 

87 Cyclopteridae 27 

88 Brosmophycis marginata 24 

89 Syngnathus spp. 22 

90 Zaniolepis spp. 20 

91 Ophidion scrippsae 19 

92 Diogenichthys spp. 18 

93 Atherinidae 17 
93 Medialuna calif orniensis 17 
93 Scopelogadus bispinosus 17 



67 



TABLE 3. (cont. ) 

Rank Taxon Count 



96 Scorpaenichthys marmoratus 16 

97 Chromis punctipinnis 15 
97 Bathylagus pacificus 15 
97 Poromitra spp. 15 
97 Aristostomias scintillans 15 
101 Anguilliformes 14 
101 Stomii formes 14 
101 Pleuronichthys decurrens 14 
104 Bathylagus spp. 11 
104 Macrouridae 11 
106 Scombridae 8 
106 Seriola lalandi 8 
108 Brama spp. 6 
108 Macroramphosus gracilis 6 
108 Myctophum aurolaternatum 6 

111 Engraulidae 5 

112 Ophiodon elongatus 4 

113 Electrona rissoi 3 
113 Bathylagus milleri 3 
113 Serranidae 3 
113 Scopelosaurus spp. 3 
113 Evermannellidae 3 
113 Carapidae 3 
113 Tetraodontidae 3 
113 Diplophos taenia 3 
113 Alepocephalidae 3 
113 Notolychnus valdiviae 3 
123 Bothus spp. 2 
123 Gobiesocidae 2 

Total 357046 



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180 



TABLE 6. List of stations with multiple occupancies in one month 
during 1955. Stations were occupied twice in one month 
except those indicated by an asterisk, which were occu- 
pied three times. 



Station 


103.0 


35.0 


103.0 


40.0 


103.0 


50.0 


103.0 


60.0 


103.0 


70.0 


103.0 


80.0 


107.0 


80.0 


110.0 


60.0 


110.0 


80.0 


113.0 


60.0 


117.0 


60.0 


120.0 


35.0 


120.0 


45.0 


120.0 


50.0 


120.0 


55.0 


120.0 


60.0 


120.0 


70.0 


120.0 


80.0 


123.0 


50.0 


123.0 


60.0 


127.0 


50.0 


127.0 


60.0 


130.0 


50.0 


130.0 


60.0 


97.0 


30.0 


97.0 


32.0 


97.0 


35.0 


97.0 


40.0 


97.0 


45.0 


97.0 


50.0 


97.0 


55.0 


100.0 


29.0 


100.0 


30.0 


100.0 


35.0 


100.0 


40.0 


100.0 


45.0 


100.0 


50.0 


100.0 


55.0 


103.0 


30.0 


103.0 


35.0 


103.0 


40.0 


103.0 


45.0 


103.0 


50.0 


103.0 


55.0 


103.0 


60.0 


103.0 


65.0 



Month 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

4 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 

6 



Station 


Mon 


103.0 


70.0 


6 


103.0 


75.0 


6 


103.0 


80.0 


6 


103.0 


85.0 


6 


103.0 


90.0 


6 


107.0 


32.0 


6 


107.0 


35.0 


6 


107.0 


40.0 


6 



181 



INDEX 



This index lists taxa included in Table 4 with their page 
numbers. 

Page 

Anguilliformes 69 

Clupei formes 

Clupeidae 

Etrumeus acuminatus 69 

Sardinops sagax 69 

Engraulidae 7 2 

Engraulis mordax 7 2 

Salmoni formes 

Alepocephalidae 75 

Argentinidae 

Argentina sialis 75 

Microstoma microstoma 76 

Nansenia Candida 77 

Nansenia crassa 77 

Bathylagidae 

Bathylagus spp 78 

Bathylagus milleri 78 

Bathylagus ochotensis 78 

Bathylagus pacificus 80 

Bathylagus wesethi 81 

Leuroglossus stilbius 83 

Stomiiformes 87 

Gonostomatidae 

Cycl othone spp 87 

Diplophos taenia 89 

Ichthyococcus spp 89 

Vinciguerria lucetia 90 

Sternoptychidae 92 

Stomiatoidea 

Chaul iodont idae 

Chauliodus macouni 93 

Idiacanthidae 

Idiacanthus antrostomus 94 

Malacosteidae 

Aristostomias scintillans 95 

Melanostomiidae 

Bathophilus spp 95 

Stomiidae 

Stomias atriventer 95 

Myctophi formes 
Alepisauroidei 

Evermannellidae 96 

Paralepididae 97 

Chlorophthalmoidei 

Notosudidae 

Scopelosaurus spp 98 

Scopelarchidae 98 



182 



Page 

Myctophoidei 

Myctophidae 99 

Lampanyctinae 

Ceratoscopel us townsendi 100 

Di aph us spp 101 

Lampadena urophaos 10 3 

Lampanyctus spp 103 

Lampanyctus regalis 104 

Lampanyctus ritteri 104 

Notolychnus valdiviae 108 

Notoscopelus resplendens 108 

Stenobrachius leucopsarus 108 

Triphoturus mexicanus 110 

Myctophinae 

Di ogeni chthys spp 114 

Diogenichthys atlanticus 114 

Diogenichthys laternatus 115 

Electrona rissoi 117 

Gonichthys tenuiculus 118 

Hygophum spp 118 

Hygophum atratum 119 

Hygophum reinhardtii 119 

Loweina rara 120 

Myctophum aurol aternatum 120 

Myctophum nitidul urn 120 

Protomyctophum crockeri 121 

Symbolophorus californiensis 124 

Tarletonbeania crenularis 12 5 

Synodontoidei 

Synodontidae 

Synodus spp 127 

Gadiformes 

Bregmacerotidae 

Bregmaceros spp 128 

Merlucciidae 

Merl ucci us productus 128 

Macrouridae 131 

Ophidiiformes 132 

Bythitidae 

Brosmophysis marginata 132 

Carapidae 132 

Ophidiidae 

Chilara taylori 133 

Ophidion scrippsae 133 

Gobiesociformes 

Gobiesocidae 133 

Bel on i formes 

Scomberesocidae 

Cololabis saira 133 

Ather in i formes 

Atherinidae 134 

Lampri formes 

Trachipteridae 13 5 

183 



Page 

Beryci formes 

Melamphaidae 

Melamphaes spp 135 

Poromi tra spp 137 

Scopelogadus bispinosus 137 

Syngnathi formes 

Macroramphos idae 

Macroramphosus gracilis 138 

Syngnathidae 

Syngnathus spp 138 

Scorpaeni formes 
Cottoidei 

Agonidae 138 

Cottidae 138 

Scorpaeni chthys marmoratus 139 

Cyclopteridae 139 

Hexagrammidae 

Ophiodon elongatus 139 

Zani ol epi s spp 140 

Scorpaenoidei 

Scorpaenidae 

Sebastes spp 140 

Percif ormes 
Blennioidei 

Blenniidae 

Hypsobl enni us spp 144 

Clinidae 144 

Gobioidei 

Gobiidae 144 

Labroidei 

Labridae 14 5 

Pomacentridae 

Chromis punctipinnis 147 

Percoidei 

Bramidae 

Brama spp 147 

Carangidae 

Seriola lalandi 147 

Trachurus symmetricus 147 

Kyphosidae 

Medialuna calif orniensis 151 

Malacanthidae 

Caulolatilus princeps 151 

Sciaenidae 151 

Serranidae 152 

Scombroidei 

Scombridae 152 

Scomber japonicus 152 

Trichiuridae 154 

Sphyraenoidei 

Sphyraenidae 

Sphyraena argentea 154 



184 



Page 

Stromateoidei 

Centrolophidae 

Icichthys lockingtoni 155 

Stromateidae 

Peprilus simillimus 156 

Tetragonuridae 

Tetragonurus cuvieri 157 

Trachinoidei 

Chiasmodontidae 158 

Pleuronecti formes 158 

Pleuronectoidei 
Bothidae 

Bothus spp 159 

Paralichthyidae 

Citharichthys spp 159 

Citharichthys fragilis 161 

Citharichthys sordidus 162 

Citharichthys stigmaeus 163 

Citharichthys xanthostigma 165 

Hippoglossina stomata 167 

Paralichthys californicus 167 

Xystreurys liolepis 168 

Pleuronectidae 

Glyptocephalus zachirus 168 

Lyopsetta exilis 168 

Microstomus pacificus 169 

Parophrys vetul us 170 

Pleuronichthys spp 171 

Pleuronichthys coenosus 171 

Pleuronichthys decurrens 172 

PI euroni chthys ri tteri 172 

Pleuronichthys verticalis 172 

Soleoidei 

Cynoglossidae 

Symphurus spp 172 

Tetraodonti formes 

Tetraodontidae 173 

Disintegrated fish larva 173 

Unidentified fish larva 175 



185 



RECENT TECHNICAL MEMORANDUMS 

Copies of this and other NOAA Technical Memorandums are available from the National Technical 
Information Service, 5285 Port Royal Road, Springfield, VA 22167. Paper copies vary in price. 
Microfiche copies cost $4.50. Recent issues of NOAA Technical Memorandums from the NMFS 
Southwest Fisheries Center are listed below: 

NOAA-TM-NMFS-SWFC- 73 United States North Pacific Albacore Fishery 1961-1980. 

A.P. MAJORS 
(April 1987) 

74 Abundance of zooplankton species in California coastal waters 
during April 1981, February 1982 and March 1985. 

A. ALVARINO and C.A. KIMBRELL 
(June 1987) 

75 Data report on the vertical distribution of the eggs and larvae of 
northern anchovy, Engraulis mordax, at two stations in the 
Southern California Bight, March-April 1980. 

T. POMMERANZand H.G. MOSER 
(July 1987) 

76 Report of a marine mammal survey of the Eastern Tropical Pacific 
aboard the research vessel David Starr Jordan, July 29-December 
5, 1986. 

R.S. HOLT and S.N. SEXTON 
(August 1987) 

77 Report of a marine mammal survey of the Eastern Tropical Pacific 
aboard the research vessel McArthur, July 29-December 6, 1986. 
R.S. HOLT and A. JACKSON 

(August 1987) 

78 Results of the Bay Area Sportfish Economic Study (BASES) 
C.J. THOMSON and D.D. HUPPERT 

(August 1987) 

79 Ichthyoplankton and station data for California Cooperative 
Oceanic Fisheries Investigations survey cruises in 1951 . 

D.A. AMBROSE, R.L CHARTER, H.G. MOSER and OR. SANTOS 

METHOT 

(September 1987) 

80 Ichthyoplankton and station data for California Cooperative 
Oceanic Fisheries Investigations survey cruises in 1952. 

E.M. SANDKNOP, R.L. CHARTER, H.G. MOSER, and J.D. RYAN 
(September 1987) 

81 Ichthyoplankton and station data for California Cooperative 
Oceanic Fisheries Investigations survey cruises in 1953. 

E.G. STEVENS, R.L. CHARTER, H.G. MOSER, and M.S. BUSBY 
(September 1987) 

82 Ichthyoplankton and station data for California Cooperative 
Oceanic Fisheries Investigations survey cruises in 1954. 
B.Y. SUMIDA, R.L. CHARTER, H.G. MOSER, and D.L SNOW 
(September 1987)